Por Hin co)ivctiio realizado eni 7 de dezembro de 1954,

entre o Estado do Pará e o Instituto Nacional de Pesíjnisas

da Amazônia, foi a direção cie7itifica e a administração do

Museu Paraense Emílio Goeldi entregue a esse órgão a par-

tir de IP de Janeiro de 1955, pelo espaço de 20 ai7os.

O Museu Paraense Emílio Goeldi continuará a perten-

cer ao Estado do Pará, recebendo o Instituto Nacional de

Pesquisas da Amazónia todo o seu acewo, (jue administrará

com ainpla e total autononúa, sem, entretanto, poder alienar

qualquer parcela de seu património . Gonstruções e benfeito-

rias que forem feitas nos terrenos c prédios do Museu, nesse

período, passarão a constituir seu patrianónio.

Todo o pessoal do Museu Paraense Emílio Goeldi f>assa

a ser de livre escolha do Diretor do Instituto Nacional de

Pesquisas da Amazómia, obedecido o disposto nos decretos

federais -nP 31.672, de 29 de outubro de 1952 e

35.1 >3, de 1.^’ de março de 165 -/, ijue, respectivamente, cria-

ram e regulammitaram o funcionamoito do Instituto. Os

servidores do Museu que não f<n-em aproveitados em sua

nova organizaçãio, ficarão à disposição do Governo do

Estado .

Deverá o Instituto Nacional de Pesíjui.sas da /ímazónia

promover o reeíjuipamento do Museu Paraoise Emílio Goel-

di, restaurar suas instalações, atualizar sua biblioteca e dotar

o mesmo coni um tiuadro de especialistas nacionais ou es-

trangeiros à altura da elevada mis.são de contitiuar a obra

com tanto brilho realizada por .seus aniece.ssores .

Dois anos a)ites de findo o prazo do conxiénio, poderá

éste ser denunciado por (inabjuer das partes, em caso con-

trário sendo considerado auton\àticamente prorrogado por

períodos suce.ssivos de cinco anos, .sempre que não .seja de-

nunciado com dois anos de antecedência .

SciELO

10 11 12 13 14 15 16

Mus. Goki.di

Tomo XI (I) — Setembro 1955

Denunciado o coiwênio e fhido o prazo de sna vigência,

passarão para o (iovêrno do Estado do Pará a responsabi-

lidade da adniiyiistraçãio do Mnseyi Paraense Emílio Goeldi,

bem como os respectivos encargos relativos a pessoal e ma-

terial, inclusive os resultantes de contratos, acordos ou con-

vênios assinados pela administração do Instituto Nacional de

Pesquisas da Amazônia.

O Museu Paraense Emílio Goeldi continuará, através

de seus técnicos e cientistas de outras instituições a contri-

buir para o estudo das ciências naturais na região Amazô-

nica, ynatitendo, a.ssim, a sua tradição e área de ação.

O Boletim será mantido sob o formato anterior e pu-

blicaréi trabalhos também etn líjiguas estrangeiras qyumdo

o âmbito ou natureza purarnente técnica do trabalho assiyn

o exigir.

REVISÃO DO COMPLEXO CYRTORHINUS FIERER -

MECOMMA EIEBER (IlEMIPTERÁ-HETEROPTERA,

MÍRIDAE)

I’or

[osí; C. M. Carvalho

Museu Paraense Kniílio Goeldi, líeiéin, Pará, Hrasil

'r. R. E. SouiHwooi)

r)c|>artanicnto dc Entomologia, EstaçHo Experimental de Rotiiamsted, Ilarpenden,

Inglaterra

INTRODUÇÃO

Os percevejos de plantas on miridios incluídos no complexo aci-

ma apresentam grande interesse econômico e taxonômico . Algumas

espécies são utilizadas no controle biológico de cigarrinlias (liomóf)-

leros) nas Ilhas do Pacifico c encontram-se, no momento, em estado

de confusão taxonômica .

Atualmente estão incluídas no complexo espécies pertencentes

a duas subfamilias diferentes, Orthotylinae e Phylinae, fàcilrnente

separáveis pela presença ou ausência de verdadeiros arólios entre

as unhas.

No.<!sos estudos mostram que Eieber estava certo quando esta-

beleceu, em 1864, o género Tylllitts para abranger duas espécies sem

arólios, os quais .ie acham substituídos por pêlos entre as unhas.

Assim sendo, tódas as espécies do complexo com êste e outros carac-

teres dos Phylinae deverão passar ao gênero Tyttiuis, embora auto-

res recentes, não reconhecendo a importância das unhas, hajam co-

locado as mesmas em (^yiloiliintis (Orthortylinae). .Ilgumas espé-

cies dêste último gênero deverão .ser transferidas para Mccoiiiina,

que continua sendo mantido como género independente . Um gê-

nero noxio, Eicl)crocaj)stis, c proposto para a espécie ílaveohis Reuter.

Essa confusão existente entre entomólogos experimentados re-

sulta do fato de serem as espécies dêsse complexo, embora perten-

centes a subfamilias diferentes, muito .semelhantes entre si c com os

mesmos hábitos e halritttts, constituindo, assim, um caso de acentua-

da e curiosa convergência .

SciELO

10 11 12 13 14 15 16

CONTENTS

Introdiiction

raxonomic history of lhe Cyrtorhinus — complex

Mnjor charactcrcs separating tlic Orthotylinac and 1’liylinae

Index to speties which inay l)e referred to the Cyrtorhinus — Mccommn

complex

Kcy lo genera

Tytthus Fiebcr

FieherorajKsus nov. gen

C\rlorliiuus Fieber

Mfcoinnia Fiebcr

IMology

l’lciygopolymorpbisni

Feeding babits

immatnie stages

Habitat

Zoogeogiapby

C.lictk list

Snnnnary

AtknotvIedgemeiUs

Referentes

SciELO

10 11 12 13 14 15 16

INTRODUCTION

The Miricl biigs in tliis coinplcx are of both cconomic and

taxonomic intcrcst; for not only are tbey iniportant in Ijiological

control of ieaf boppers, esj)ecially in tbe Pacific Islancls, but

taxonomically tbey are in a confnscd State.

At jnesent tbe coiuplex inchides species belonging to two

distinct subfaniilies, tbe Ortbotylinac and Pbylinae, wbicb, liave

rcspectively, flap-like and bristie-like arolia.

Otir stndies bave sliown tliat Fiel)cr was corrcct tvben in 1801

lie crectcd tbe gemis Tytllnis for two .sjtecies witb Itristie like arolia.

Ilcnce all specics witb tbis and otber pbyline cbaracters (see pj).

l.H-M) sbonbl now be included in Tyttlnis; altboiigb later autbors,

not recognising tbe iniportance of tbe claws, bave placed all spccies

incUidcd in Tyttlius under Cyrtorhimts, wbicb is in tbe Ortbotylinae.

Tbose s|)ecies bitberto in Cyrlorliinus wbicb bave Ortbotyline

cbaracters niostly reniain in tbis gentis, biit a few sboiild be

transferred to tbe closely related Mccomma, wbicb is retainetl as a

genus. A new gemis, riehitroatpsus, is raised for flnxieoliis Reuter.

'íbis confnsion bas arisen ainongst expericnced entomologists

becaiise tbese species, belonging to two diffcretu snb-fainilies are

so inncb alike in ajipearance and bave similar biibits and babitat;

tbns beitig a striking case of convergente.

TAXONOMIC MISTORY OK TIIK CVRTORHINUS

COMTLEX

'I be gemis Cyrlorliiuus was describcd by Fieber (Wien. Knt.

Monat . 1858) to incbide a single species, Capsits clcgtnilulus

Meyer-Dih, 1815. Fieber (Fiir. llein.: 0'.), 1800) separateil it, in

a key, froin otber Kuropean getiera and redescribeil tbe type (F.iir.

Iletn.: 285, 1800).

riie getms Tytlliu.s, geneially legarded by aiitbors as a synonym

of f.’)'i7or/(init.v, was descí ibed by Fielter (Wien. làit. Monat. 8: 82,

1801) to incbide C<il>sus grniiniis Flor and bVi/rvics l)ygiiiacus

Zetterstedt. I be ty]H; of tbe gemis was latei' fixed by Kirkaldy

(('Fraiis. .\iiier. F.nt. .Soe. 52:128, 1000). Tlie lirst anilior lo incbide

Tyttlius in tlie synonyiny of Cyrtoiiiiiius was Reiiter (bib. K. .Sv.

Vet. Akad. For. 5 (1): 51, 1875).

IJoi.. Mus. Goki.di — Toino XI (I) — Setembro 1955

riiomson (Opuse. Enl. 4: I.'!?, 1871) cletected thc synoiiymy

of Capsus carieis Fallen, 1807 anel Capsus eleganlulus Mcycr-l)ür,

1843.

Reuter (Rev. Crit. Gaps. 1: 91, 2: 12, 1875, and Hih. K. Sv.

Vct. Akad. Mandl. 3 (1): 31, 1875) considered Cyrtorhimis as a

subgemis of Chlamydatus Clurtis listing four species witb synonyniy.

l)escri{)tion of tbe gentis and species Avas given latcr (Hem. Gymn.

Eur. 3: 379, 545, 1883).

Uhler (Proc. Zool. Soc. London: 711, 1893) described Cylloceps

peilicin, new genus and sjjecies from St. Vicent II., Avbich was later

found by China to be a synonym of Cyrtorhinus pawiceps Reuter

(Ann. Mag. Nat. Hist. (9) 14: 444, 1924).

Breddin (I)eut. Ent. Zcit.: 106, 1896) describetl Periscopus

mundulus, new genus and species from Java, thc gcncric name

being preorcu{)ied by /Vrúfo/Jtw Eitzinger, 1843 (Rcj)tilia). Kirkaldy

(VVien. Ent. Zeit. 22: 13, 1903) establisbed lireddiniessa a n. nov.

for Periscopus llrcddin.

Distam (Faun. llrit. Ind. Rliync. 2: 476, 1903) rcdcscribcd

thc gentis CyrlorliÍ7ius Ficber, listing its synonymy and giving a

figtire of C. lividipCTitiis Reuter thcn found in Ceylon, Hurma and

Great Nicobar.

Knight (Conn. Nat. llist. Surv. lUil. 34: 509, 511, 1923)

keyed tbe genus and desci ibed (iyrlorliiuiis carieis var. vagus, hc

mentions thc typical carieis from Colorado and says that hc had

coni|jarcd it with a Finnish spccimcn named by Reuter. A femalc

of tliis species from Colorado and a series from Wrattgel, Alaska

are to be found in thc U.S. National Museuni. Most aincrican

rccords are howevcr Tyttlius vagus (Knight, 192.3), a species of

Phylini .

Hueber (Syti. lUindw. 2: 106, 1908) gives a key and dcscriptions

of thc Gcrman species.

Poppius (Acta .Soc. Sei. Fenn. 11 (3): (U), 70, 1911) redcscribcs

tlie genus from África, listing with dcscriptions, C . pandeeps and

. lucgalogps (error pro iticlauops Reuter).

üsiiiger (Proc. llaw. Ent. .Soc. 10 (2): 271, 19.39) gives host,

distribiition, notes and a key for the Pacific species, and is tlic

first recent author to draw attetition to lhe faci th;ii some species

had only brisile likc arolia while olheis had tiue convergem arolia.

Latcr (.Soc. Sei. Fenn. Cominem. Hiol . 12 (8): I, 1951) thc same

aulhor in a revisioii of lhe Pacific species proposed lhe subgemis

liculericssa for lhe species with Oiiliotylini claws and arolia, kceping

lhe ('.yrtorliiiiu.s s. sir. lor lhe Phylini species. Ilis tvork was

baseei on spccimens misnamed by F. . P. Van Du/ce in tlic Califórnia

Acacleniy ol Sciences, who named a Phylini species (which was

Carvalho S; Southwood — Cyrtorliinus-Meconinia Complex

actually llie sainc as Kniglu’s CyrtorJiinxis carieis var. xiagtis) as thc

true Eiiropcaii Capsus carieis Fallcn. In tlie saine pajjcr a new

sjiecics, C. vitiensis was clcscribcd Irom Fiji.

lllatchlcy (Hcin. Hct. F.N. Amer.: 845, 853, 1926) mentions

carieis vagas Kniglit aiul pygmacas Irom thc United States. Here

again the aiithor was misled, since pygmacas Flor does not occiir

in America. Specimens wliicli were named as siieh by Van Dtizec

are conspecilic with Cylloeeps pellicia Ubier, acttially Tytthas

paiineeps (Reiiter).

Zimmcrman (Ins. Hawaii, 3 Het.: 205, 1918) gives the history

of Cyrloriiimts in Hawaii, biological control and notes for mundalas

and f alvas Knight, with good ilhistrations. Notes on species from

Guam are to be found in Usinger (Ins. Guam, II: 79, 1916).

Knight (Ins. Samoa II, Meni. 5, 1935) described C. fulvas from

Samoa, lhe same atithor (111. Nat. Hist. Siirv. Bul. 22 (1): 82,

95, 1941) mention the presence of carieis in Minnesota.

Carvalho (An. Acad. Brasil. Ci. 24 (1): 76, 1952) includes

in the .synonymy of Cyrtorhhms the gentis Arislobalas Distant (Ann.

Mag. Nat. Hist. (8) 5: 16, 1910) and Nycticapsus Poppius (Acta

Soc. Sei. Fcnn. 44 (3): 74, 1914). 'Fhc authors inchide them as

synonyms of Meeotnma Fieber in thc present pajter.

Wagner (ricrw. Deut. 41, Blindw.: 110, 127, 1952) deals

with thc gentis Cyrlorliitias in Gcrmany in which hc includes earieis,

flaveohis, pygmacas and geminas with ilhistrations.

Kiritchenko (Hem. Ftir. URSS: 175, 1951) gives keys for the

four sjiecies mentioned above (in Ritssian).

Cat;ilogtie rcfcrcnces on the geiuis are to bc fotind in Atkinson

(1890), Oshanin (1906, 1912), Van Du/ce (1917), .Stichcl (193.3),

China (1943), Carvalho (1952) and Carvalho S: Leston (1952).

M.VJOR chakacteres separa einc; orehoty-

LINAE AND PIIYl.lNAE

riic Orthotylinae and Phylinac are most easily sciiaratcd on

three characters:

1 . Prrlaxsal struetare: thc claws of thc Orthotylinae have

sniall pads, the pseudarolia, on their umlcrsidcs, whilst bctwccn the

claws are a p:tir of coiivergcnt membr.ineons arolia (Fig. I F);

jjseiuIoaroli;t are aiso present in the Phylinac, biit thc arolia are

thin and hairlike (Fig. 1 G).

2. hemale gctiilalia: .Slater (1950) found lhat in typical

Orihotylinac the scicrotiseti rings on the tlorsal wall of thc bursa

copulatrix have their later;d margiii strongly folded dorsoines.id:

SciELO i'o

Carvalho & Southwood — Cyrtorliinus-Meconinia Coinplex

l'vvo claspcrs or parameres are situatccl on either side oí the

ápices of the subgenital jtlate. They are asymmetrical and in general

in the Orthotylinae difler niarkedly Irom one spccies to another.

In the Phylinae, the niost striking snjjerlicial featiire is the

posterolaterally directed sclerotiscd sheatli (Fig. 1 E). According

to Kidlenberg this consists of a fusion of the subgenital plate and

theca. lUit it sceins that owing to the niechanics of the aedaegus

during copnlation snch a fusion is impossible and this sheath cannot

be homologised with the tlicca of the Orthotylinae. It corresponds

only with the subgenital plate or aedeagal support and titis is

confirnted by the atlachment of the stapes (Fig. 1 D) to its margin.

This structure sharply distinguishes the typical Phyline froin the

Ortliotyline genitalia and is refcrred to as the aedeagal sheath. The

Phylinae differ further in the lack of sclerotised vesical appendages

and in tlie modification of the aedeagus into a strap like structure.

riie claspers of relatcd spccies of Prylinae are oftcn vcry similar

in form, more so than in the Orthotylinae.

INDEX TO .SPECIES VVHICH MAY 15E REFERRED TO THE

CYR TORIIINUS-MECOMMA COMPLEX

cYirroniiiNus

Original gcncric

assígntncnt

Present gcncric

assignmcnt

alboonmtus Kiiighl. 1931

C;ynorliimi5

Tytlliiis

* aiiiiulirollis I’()p])ius, 1915

Cyrlorhimis

'rytlluis

chiiinisiu (.Slal, 1859)

Ixilli Kiiight, 1931

Cyrtorliinus

Tyllhus

cnricis lallcn, 1807

Capsiis

Cyilorhimis

rollfiris Malsuiiuira, 1911

C.lilainydatus

chiucrisis Slal, 1859

Oapsiis

Tyuhus

• chloroplrrus Hcrrich-Sflicfícr, 1853

Cajisus

Ciyriorhimis

= carieis (l■'alU■n, 1807)

nimberi tVoodwarcl, 1950

Cyrlorhimis

Cyrloihimis

• rlc/^aiiltilus Mcytr, 1813

Capsus

Cyriurhimis

— carieis (l-allcn, 1807)

• clnui^alus goppiiis, 1915

Ciyrlorhimis

■rytihiis

— chinensis (Slal, 1859)

• filius Dislaiit, 1910

Arisloholus

Mccoiiiina

— ninicus (Diilaiit. 1909)

ilavcnlus Kriilfr, 1870

lyllliiis

hicherorapsus

lulvus Kiiíglit. 1955

Cyrlorhimis

f^rmitius Mor, 1800

Capstis

■ryllhiis

— sYiioiiyin

Hoi,. Mus. Gokldi — lomo XI (I) — Setembro 1955

CVUrORlIlNUS

Original gcncric

assignmcnt

I*rcscni gcncric

assigninciu

* insignis Douglas ft Scott, 1866

'Fytthus

= pyginaeus (Zcttcrstcdt, 1810)

iiisf/eralus Knight, 1925

Cyrtorhinns

Tytlhus

lividipennis Reuter, 1884

Cyrtorhinus

marginatus Uhlcr, 1895

Cyrtorhinns

Orthotylus

melatiocephalus Poppius, 1914

Nycticapsus

.Mcconnna

rnelanops Reuter, 1905

Cyrtorhinus

muridulus Brcildiu, 1896

Pcriscopus

'Fytthus

iieotropicalis Carvalho, 1954

Cyrtorhinus costae

Carvalho ncc Stal

Tytthus

parviceps Rcutcr, 1890

Cyrtorhinus

'Fytthus

• pellida Uhler, 1893

Cylloccps

'Fytthus

— parvicejis (Rcutcr, 1890)

* pelluce7is Holicman, 1852

Capsus

'Fytthus

= (Zcttcrstcclt, 1840)

“pubescens Knight, 1931

Cyrtorhinus

'Fytthus

= geminus (Flor, 1860)

frygtnaus Zctterstcdt, 1840

Capsus

Tytthus

* riveti Chccsnian, 1927

Cyrtorhinus

'Fytthus

= chinetisis (Stal, 1859)

1 'agus Knight, 1923

Cyrtorhinus

Tytthus

‘viliensis Usingcr, 1951

Cyrtorhinus

= lividipennis Rcutcr, 1884

swaluwenburgi Usingcr, 1944

Cyrtorhinus

'Fytthus

MECOMMA

arnbulans Fallcn, 1807

Capsus

Mccomma

amicus Distant, 1909

Antiphilus

.Mccomma

nntennnia Van Du/cc, 1917

.Mccomma

Mccomma

• dnbius Zctterstcdt, 1840

= ainbulans (Fallcn, 1807)

Capsus

Mccomma

chinensis Rcutcr, 1906

Mcioinnia

Mccomma

gilvipes .Stal, 1858

I.cptonicrocoris

iMccoinina

• luctuosHS 1’rovanchcr, 1887

= gilvipes (Stal, 1858)

Chlainydattis

Mccomma

tnadagasrariensis Rcutcr, 1892

Mctoinina

Mccomma

* nigrilulus Zctlerslcdt, 1840

= ambninns (Fallcn, 1807)

Capsus

Mcconnna

• ocUripes Curtis, 1838

Chiainydatus

.Mccomma

= arnbulans (Fallcn, 1807)

n «ynonyni

Carvalho & Solthwooi) — Cyitorhiiuis-Mcconima Coniplex

KEY rO GENERA

1. Prctarsus with a pair of hristle-likc arolia (Eig. 1 G), inale

pygophorc witli a strongly sclcrotisccl tubular shcat íor the

actlacgus (Eig. I D) (Phylinac) Tytthus Ficber

Prctarsus witb a pair of flap-likc arolia (Eig. 1 F), malc

jjygophorc uot tlcvclopctl to form siich a projcctiug shcath

(Orthotyliuac) 2

2. Prouotum straiglu sitletl (Eig. 10 G), its lower margiu slightly

coucavc, ctiueus witlcr thau loug (Eig. 16 C), (pubcsccuce

short: 0.08-0.12 uiiu. without auy black coloratiou)

Ficherocapstis ii. gcu.

Prouottmi bell-sbapctl or caiupautililorm (Eig. 15 A), its lower

margiu straiglu (fretjtieiuly partly or eutirciy black) 3

3. Ctiueus ouly slightly louger thau witle (Eig. 16 A); slightly

rotiutlctl iu outliue; pubcsccuce short; 0.15 iiuu. or Icss, rostrum

uot rcachiug posterior coxa (uever comjilctcly brachyptcrous)

Cyrloritimis Ficbcr

Ctiueus cousitlerably louger thau wiilc (ofteu twicc as uuich)

(Eig. 16 1$); males almost ])arallcl sitletl; pubcsccuce loug:

mostly tiver 0.15 mm.; rostrum reat:hiug bcytuitl base ol

posteritir coxa (females ofteu completely brachyptcrous (Eig.

22 A, 11) Mecomma Eieder

c;enijs tytthus fieber

Tytthus Eiehcr, Wieu. Eut. Mouat. 8:82, 1864.

• C.yUoceps llhier, Proc. Zool. .Soc. Eoutlou : 711, 181)3 (uov. syu.).

• ITriscopus llretltliii, Deut. Eut. Zeit. . 106, 1806 (uoem. pretic.

hy Periscopus Eit/iuger, 1813, Reptilia).

• lircdditiirssa Kirkaltly, Wieu. Eut. Zeit. 22:13, 1003 (uom. uov.

ftir Periscopus llretitliu, 1806) (u. .syu.).

Eype species: Ctipstis gcmiuus Elor, 18()0, lixetl by Kirkaltly, Iraus.

Amer. Eut. Soc. 32:128, 1006.

Small bugs (2. 2-3. 6 mm.); heatl rtiuutletl auteriorlf]. face semi

vertical; prouotum camjiauuliform, calli slightly marketl; tipeuiug

of otliferoiis glantl raisetl auti well marketl, |)ret;usus with bristie

like artilia; male pygophorc with a projcctiug aetleagal shcat; lemale

Inirsa copuhitrix with symmetrit;il A struetures. Coveretl by simple

erect pubesceuce (0.100.10 mm. in leugth); rostrum rcachiug the

hiutl coxae; colour patteru geuerally black aiitl [lale greeu.

_ 2

]5ol. Mus. Goeudi — loino XI (1) — Setembro 1955

KEY rO THE SPECIES OF I HE GENUS TYTTIIUS

1 . Colour pale yellorvish testaceous

zwahmeiiburgi (Usinger)

C^olour ií pale then jjionotuni, head and scutellum black or

Avith daik brown to black areas 2

2. Firts antennal seginent pale, darkened only at extreme apex

or extreme base 5

First antennal segment black, pale only at extreme base and

apex b

3. Hemelytra white at least on basal thlrd 4

llemelytra darkened basally or unicolorous but not as above

4. Hemelytra witb a wide dark brown to black faseia, the basal

thiixl and cuncus whitish albooiiiiitus (Kniglit)

Hemelytra with only basal tliird wbitisb . . . mmitamis n. sp.

5. Hemelytra noticeably inínscate longitudinally along clavns

and endocorium, pronotum totally black

muntlulus (llreddin)

Hemelytra pale green, if infuscate, tbe ])ronotnm pale at basal

angles gctniinis (Flor)

G. First antennal segment entirelly black, tibiae black to dark

brown; |)ronotum sligbtly constricted at middle 7

First antennal segment jtale at least on extreme apex or base,

tibiae pale to tuscous, not black; pronotum not constricted

at midtlle 8

7. Pronotum brownisli black witb a transverse itiilky wbite faseia

in front of calli, ciineus pale iii.sl)ci(itus (Kniglit)

Pronotum orange yellow on anterior margin of disc, cuncus

witb a darkened apex bnlli (Kniglit)

8. First antennal segment witb a black ring at middle Icaviug

apical and basal third wbite ncoIrDpicíilis (Garvalbo)

First antennal segment mostly black, witb only extiemc apex

or base pale 9

9. Pronotum usually pale anteriorly (iii front, betwcen or over

calli) 10

Pionottim usually totally black or dark brown II

10. 1'ibiae eiitirely yellow; piouotum varying front pale witb only

posterior angles black to black rvitb a pale tentral are.i

anteriorly fiygniavus (Zetterstedt)

Tibiae witb extreme base dark; pronotum black, only pale

anteriorly futnficeps (Reuter)

Carvai.ho &: SouTiiwooD — Cyrtoihimis-Mecomnia Complcx

1 1 . l ibiac eiuircly pale, Iicinclyira largcly infuscate, palcr along

anterior niargin (large sj)ecies, about 2.8 iinn long)

vagiis (Knight)

Tibiae witb extreme base or basal ])ortioii brovvnish to black;

hemelytra mostly pale green 12

12. Pale spots on vertex obsolete; iiiulersicle rufescent

panarnensis n. sp.

Pale spots on vertex ilistinct; niulerside dark brown to black

; 13

13. Larger sjjecies, tbe spots on vertex larger

paniceps var. tlioracicus (Morvatb)

Sinaller species, tbe sj)ots on vertex ininnte . . cliinensis (Stal)

TYTTIIUS ZWALUWEN nu R('.l (II.SINCIPR) nov. coinb.

Cyrtorliinus zuialimenbitrgi Usinger, Proc. Hawaii. Ent. Soc. 12

(1): 148, fig. 1, 1944.

(Fig. in Usinger, 1944)

Cliaracterized by its entirely jtale ycllowisb testaccoiis colour anil

dimensions.

Male: lengtli 2.3 mm., widtli 0.8 mm.

Colom pale ycllowisb testaceoiis exce|)t for dark brown cyes and a

vague brown area at middie of bead.

Rostrmn reatbing nearly to ajjices of middie coxac.

Dislrilnitioii: Cantou Is.

Specirnens studicd: I $, ryjje, Cantou Is., Ilawaii .Sngar

Plantcr’s Exp. .Station.

'í bis speiics arcording lo Usinger (1944) is very do.se to riveti

Clieesman (= rliincnsis .Stal) in size and stniclnie bm is sirikingly

diffeient from lifcti and all oilier described species in colom pattei n.

It will nin to riveti in Usinger’s key (Ilawaii. Ent. .Soc. Proc. 10:

271, 1939), bnt riveti lias sliorier sccond antennal segment (less tban

tbree times as long as lirst, 12: í/), a longcr rostrmn wbicb siirpas.ses

apex of middie loxa and differeiu colom .

It was takcn on Rocriiaavia associaled wiili tbe cicadellid,

Nesaloha cantonis Oman, and is possibly a predator of tbis .s|)ecies.

TYTTHUS CH/NENS/S (.S I AU) nov. coinb.

Capstis cliiiieitsi.t Sui\,, Ercg. E.ng. Resa, liem.: 2.'í8, 1859.

Cyrtorhiuwi cliiuensis Rcuicr, ()(v . F. \'et. Soc. Forb. 45 (10): 22,

1903.

20 ]5()i,. Mus. (ioF.!,!)! — 'l'oi7i() XI (I) — Setembro 1955

* CyrlorhÍ7ius elongalus I’o])|)iiis, Areli. Naturges. 80 A (8): 65,

1914 (n. syn.)

* Cyrlorhinns anniilicollis l’oj)j)iiis, Areli. Naturges. 80 A (8); 66,

1914 (?í. syn.)

* Cyrtorhinus riveli Chccsmun, Anu. Mag. Nat. Ilist. 19: 94, 1927

(n. syn.); Usiiiger; Soc. Sei. Feiiii. Comment. Hiol. 12 (8); 4,

1951.

(Figs. 2 A-E)

Characteri/ed by its colour aiul male genitalia.

Malc; length 2. 1-2.5 mm., widtli 0.8 mm.; hcad, width 0.6 mm.,

verlex 0.31 mm.; aiueiiiiae, segment 1 length 0.23 mm.; II

0.7 mm.; III 0.45 mm.; IV 0.28 mm.; proiiotum, length 0.3

mm., width 0.75 mm.; rostrnm legtb 0.70 mm.

llead blaek a|)art Irom two areas adjacent to tbe eyes, which

are pale; antennae brown-black with extreme apex ol lirst segment

pale; pronotnm aiul scntellnni blaek; bemelytra j)ale green, mem-

Í)rane and nervnres pale; Icgs yellow green to pale Inlvoiis exeept

for extreme base of tibi;i wbieb is blaek; rostrnm pale, tip tlark;

nnderside of tborax dark, abdomeii dark exeept for extreme venter

of segmems 2-8 wbieb is greeii-yellow.

Pnbescenee of fine pale adjjiessed bairs, some of tbose on tbe

baek of tbe liead anti anterior of tbe pronotnm are sligbtly longer

and more ercct. Macro])terons.

(ienitnlia: aedeagns (fig. 2 11) of pbyline type; left elasper

(lig. 2 (i) witb basal proeess sligblly sborler tban in panriceps;

riglit elasper (fig. 2 D) simple.

1'cniale; similar to male in eolonr and dimensions, sligbtly more

robnst .

Dislrihnlion; Ciliina, Formosa, Uonin lls. ((Ibiehi jima),

Marianas lls. (Cbiam, .Saipan, Rola, Finian), Caroline lls. (1‘alan

lls., Ngnin, Faraniep, Yap, Koror, 1’nlo Anna), New llebrides,

Samoa, l abiti, Fiji.

Specimctis sludicd; 1 á (lloloty|)e of rlongatus), Anping,

Formosa, Santer 1911 (Dentsdies Fntomologisehes Institnt); 1 <5

(lloloiype of nnnulir(dlis), Fainan, Formosa, 11. .Santer (Denisdics

Kntomologisehes Instilnt); I 9 ( Fy|)e of riveli), 1’apeete, l abiti,

Mareh-April 1925, L. E. (iheesman (líritisb Mnsenm); I <1,2 9,

Amabii, Fntnila, Samoa, 9.6.23, Swe/ey and Wilder; I <J , I ç,

Cibekiang Prov., Clbina, jnly 1927, Dora Wrigbt; 2 i , d|)oln, .Sonioa,

9.12-23 (Ib innida grass), Swe/ey and Wilder; I á , I 9 . Erromanga,

New llebrides, jnly 1930. L. E. (ibeesman; 26 <í , 29 9, MARl.V

Carvamio &: SouTiiwoon — Cyrtoiiiinus-Mccomma Coniplcx

NAS ISL., Guam: l’t. Oca, 5.12.15, J. L. Grcssit (collectcd at

light): laloloío, Aug. 1949, N. L. II. Krauss; Agana aíiport,

15.8.15, 11. S. Dybas; Saipan-. 1-2 m. ol' Tanapag, 10.1.45, 11. S.

Dybas; Titiian: [uly 194G, H. K. 1'ownes; Rola: ]unc 1930, T.

Esaki; CAROUNE ILS., Fulo Ami: 13.9.52, N. L. II. Krauss;

Koror Is: 30.11.47, II. S. Dybas; Ngw/» Atoll: Nguiu Ils, 3.10.52,

N. L. II. Krauss; Faraulej) Atoll: Faralep Is. 21.9.52, N. L. II.

Krauss; Yap group: E. Malrid, Kaiiif Yal, Colouia Yafrid, [uly 1950,

R. J. Goss; HONIN ILS.: CAiiclii Jiuia Relto: Aug. 19.34, ‘ll. Ikcda.

Usiugcr (1939) louud this spccics suckiiig eggs of Sogata ochrias

Kirkaldy ou Sporolobus virginicus aud ol Nilapawata lugois (Stal)

ou rirc.

'Eliis is thc smallcst spccics of Tytthus aud is distiugiushcd

by its black prouotuiu aud scutclium, tlic dark bases of thc tibia

aud thc suiall size.

1). R. Malaisc of thc Riksmtiscuiu, Stockholm, has kindly

iufonucd us titat thc typc of Capsus cliiuciisis imist bc cotisidcrcd

lost. \VY' coiisidcr thai this spccics is referred to iu StaTs dcscriptiou

{Frrg. Eug. Resa: 258, 1859): -

"Nigricaiis, uiaculis duabtis basalibus cajritis albidis; hctuclytris

fusccsccnic albidis: itcdibus abdomiuistpic disco pailide tcstacco-

flavis.

Eciualc. Loug. 3, lat. 1 1/3 luilliiu. 1’atiia: Ghiua (llougkoug).

C. elegnntulo affiuis et similis, capite aiigustiorc, thoracc

posterius latiorc, antciiuis, |)racscrtiui articulo basali, brevioribus

differt.”

lhe Dircctor of thc Dcutsdies Eutomologischcs Iiistitut has

kindly sent Poppius’ typcs for study. 'Ehat of elougalus rcprcscuts

a tcncral lualc spcciuicu of 7’. citiurnsis; thc uicasurciuciits agree

cxccpt for thc third aud fotnth antcuual scgiucnts whidi are some-

what shrivcllcd, whilst thc hc:id (.iptii t Iroiu thc spots ou thc vertex),

thc pronoum aiitl scutclium are light brovvn iiistcail of black. 'lhe

typc of autiiilicollis is a typital malc of T. cliinciisis.

TYTrUUS FARVICEFS (REI J TER) nov. comb.

Cyrtorhinus paiviccps Rculcr, Rcv. d'Eut. 9: 258, 1890; Poj)pius,

Acta Soe. Sei. Ecuu. 44 (3): 70, 1914.

• Cylloccps pcllicia Uhlcr, Pioc. Zool. .Soe. I.ondon: 712, 1893

(syn. by Ghiua, Auu. Mag. Nat. llist. (9) 14: 441, 1924).

(Eigs. 3 1-M)

Charattcri/cil by its colour aud gciiitalia.

Malf. Icugth 2.4-2. 7 mm.: width 0.85-0. 95 mm.; hcad width 0.58

mm.; vcitcx 0.3 mm.; autcutiac, segemeut 1 Icugth 0.3 mm.,

SciELO

10 11 12 13 14 15

Hol. Mus. (íoki.di — l onio XI (1) — Setembro 1955

II ().8'1 mm., III 0.5 mm., IV 0.4 mm.; jiroiiotum, length

0.29 mm., widlh 0.75 mm.; rostrum lengtli 0.75mm.

Meacl blatk wilb two j)ale aieas adjacenl to tbe eyes; antemiae

blaek except apex ol basal joiiit wbich is pale; [jronotum blaek with

two paler (usually yellowgreen) areas at anterior angles; sciitellnm

blaek; bemelytra pale yellow-green, memlnane and nervnres pale;

legs pale yellow-green with extieme base ol tibia dark; rostrnm pale,

its tip dark; mulerside ol tliorax and pygophone blaek, remainder

of abdômen green .

Pid)e.scenee of fine |)ale adpressed bairs, longer tban in pygmaeus,

es])eeially on tbe back of ibe bead. Matroplerous.

('tciiildlia'. aedeagns of pbyline ly|)e (fig. ‘I C), lelt elasper

(fig. ‘i I, |, K) with terminal proeess blnnt and slightly enrved

inwards at a|)ex, rigbt (las|)er (fig. ‘5 M) sim|)le.

Femnlc. lenglb 2. 5-5.0 mm.; widtb 0.95-1.1 mm.; bead widib 0.().‘!

mm., veriex 0..‘55 mm.; antennae, segment 1 length 0.28 mm.,

II 0.78 mm., 111 0.5 mm., IV 0.4 mm.; pronotmn, length

0.5() mm., widtb 0.92 mm.; rostinm length 0.75 mm.

Similar lo male in coloiir. Abdômen with mulerside pale green-

yellow, lateral margins dark, dorsmn pale. Dorsal wall of bnrsa

eopnlatrix very simple, as in halli and ixigns.

Disirihulioit: Kgypt, Si . Vieent, .Seyclielles, Rodrigne/. I., l’a-

ragnay. Florida, F.ast and West Afiica, Morocco, Gigepio Is. Italy,

S. /Mrica (Cape Provinee), St Helena.

Sprriíiiciis studied'. 9 ( Fyiie) Gairo, May 188(1, F. ,\ntian

(P>.M.); I 9 (Fyjieof G. peilieia). St. ViceiK, May, II. II. Smith

(B. M.); I (5 5 9 Rodrigues II., Ang.-Sept. 1918, 11. |. Snell

& II. P. riiomassel; 5 <5 I 9 l.akeland, Idorida (at ligbt), Oct.

1918, May 1952, R. I'. llussey; I 9 Mossel Bay, Gape Piovinee,

May 1952, R. F. 'I nrner; I i beaten from native Gomposite Tree,

Piapiel Post, St. Helena 27/2/.5(i, 11. F. 1). Bartlett; I 9 Waldia,

Abyssinia, l-2()/2/5(), (. \V'. G. Maclde; 1 9 Libreville, Gabon,

J. Primol; I 9 Serpent Fake, e. 9,000 ft., \Vomand)onlcbi,

Abyssinia 5/10/2(1, ). Omer Go<»per; I 9 Hora Keloli, Abyssinia,

Dec. 192(1, Omer Gooper; Gnba, Uhier (ol.; Pto. Obiddia,

Gode Prov., Panamá, Blanion tol.; Managna, Niiaragna, Baker eol.

Fhis spet ies is dose lo pygtmunts (Zetterstedt) bnt dilfers

in tbe nairower bead, in llu; blatk ring at extreme bases ol tibiae

and in tbe stiiulnre of tbe male genilalia; ilie pronotmn is geneially

daikei . l he lorm lliorariíiis was deseribed from tbe (ianaiy Ils. by

Horvalb (Ann Mns. Nat. Hnng. 8; 289, 1909) and is diaraitei ised

by baving tbe pionolmii enliiely blaek. 'I bis foiin bas subse(|nently

Carvalho & Southwood — Cyrtorhinus-Meconinm Coinjilcx

bccn rcfouiid in thc saine arca by Lindbcrg (1936, 1953), biit it

does iiot aijpcar to have bccn íound clscvs’hcrc . Howevcr thc amount

ol black colouration on thc jnonotum is very variable witliin one

popidation oi T. pyginaciis ami this probably applies to other

spccícs .

Since thc conimcnts above wcre writen thc senior aiithor has

sccn sjteciincns oi tliis sjtccics in thc U.S. National Musenm in

wbich thc variation mentioned above is to bc sccn. lhe iollowing

localities shoidd be added: Charlottc Anialic, St. Ihonias, Virgin Is.

VI, 917, M. Morrison col.; Macoris River, San Domingo, Vil, 917,

n. Morrison col.; Cuba (labcllcd by Uhlcr as Cylloceps pellicia);

Gnanajibo, Pnerto Rico, VIII, 935, H. L. Dozier col.; Lake Placid,

Florida, Bcaincr col.; íYdlsmorc, Florida ■ (naincd by liarber as

pellicia), in action oi ovijjositing in cgg oi Saccliarosydre sachaiyvora

(Wcstwood); Cuapara, Carabobo, Venezuela, IX, 938, C. II. Bailou

(on cotton).

TYTTIIUS PYGMAEUS (ZET TERSTED F)

Capsus pygtnaeus Zetterstedt, Ins. Lapp.: 279, 1810.

Tytthus pygniaeus Fieber, Wien Ent. Monat. 8: 82, 1861.

* Capstii pelhicois Bohcinan, Oiv. Sv. Vct. Akad. Forh.: 76, 1852

(syn. by Reuter, Not. Sallsk. F. Fl. Fenn. Forh. 14: 16, 1873).

Cyrtorliinus pygmaeiis Reuter, llein. Gymn. Eur. 3: 381, 551, pl.

2, lig. 4, 1883: .Saunders, Ilem. llet. Brit. Is.: 283, pl. 26,

iig. 6, 1892; AVagner, 'Fierw. Deut. 41, Blindw.: 128, 1952.

• Tylthus iusignis Douglas íc Scott, Ent. mon. Mag. 2: 247, 1866

(syn. by Saunders, Ent. inon. Mag. 13:113, 1876).

(Figs. 3A-G)

Characterised by its coloination and genitalia.

Male: length 2.85 nnn.; width 1.0 mm.; bead, width 0.7 mm.,

vertex 0.32., anicnnac, segment I length 0.32 mm., II 1.03

mm., III 0.76 mm., IV 0.76 mm.; pronotum, length 0.34 nnn.,

witlth 0. 80 nnn.; rostrum length 1.15 mm.

Head black with two pale aieas adjacent lo thc eyes; antennac

dark exce|)t Íor apex oi basal and base oi setond .segments pale;

pronotnm varying Írom pale with only thc exiieme posterior angles

dark to black with a pale cenlial arca anieriorly; scutelhim black;

hemclytra pale grey-green, nervures and membrane itale grey: legs

yellow; rostrum pale, its ti|) dark; underside oi thorax and pygophorc

black. resi oi abdômen pale green suiiuseil with iuscous towards

pygoi)hore.

1 SciELO

24 lk)L. Mus. Goei.di — lomo XI (I) — Setembro 1955

Piibescence of fine pale adpressetl liairs. Macropteroiis.

Genilalia: aeileagiis (fig. C) of pliyline type, lefi rlasper

(fig. 3 E, H) witli comparatively sbort process, riglit elasper (fig.

3 F) siniple.

Fcmale-, length 3.2 mm., widtb I .22 mm.; head, widtb 0.78 mm.,

vertex 0.38 mm.; anteimae, I 0.32 mm., II 0.87 mm. 111 0.70

mm,; IV 0.77 mm.; ])ronotum length 0.37 mm., widtb 0.87

mm., rostium length 1.2 mm.

• Colonr and pidrescencc as in inale. Maciopterons.

Distributiou: England, Wales, Netberlands, N. Erance,

Sweden,Lappland, Finland, N. Rnssia, N. Germany.

Specimeyis stxidied'. (i $ 0 ç Ilonndslow Ileatb, Middlescx,

II. K., '2.0/1 jb?), G. E. Woodrofle; 1 ç Marpenden, llerts, U. K.

11/8/54, 'E. R. E. Sontlnvood; 1 9 Elatford, Suffolk, II. K.,

20/8/54, 'E. R. E. Sontlnvood; I 9 1’ergas, Finland, Dr. Eger; 1

9 whieh has been designated tbe leelotype, labelled ‘‘Esber” aiul

“'Eyttbns Fieb, insignis Scott f . nova s|jec.” from tbe Scott eollection,

now in Power eollection (H. M.).

Ehe degree of pigmentation of tbe pronotnm of this s])ctics

varies greatly, even within one |)o|)nIation; biit in general it is largely

|>ale at tbe anterior and this togetber witb tbe entirely yellow legs

and tbe mostiy dark basal antennal segment distingnisbes it. Wagner

(1952) records this species from marsby jilaces aronml tbe bases of

rnslies and grasses; it is liowever occasionally taken by swecping and

ocenrs as an adidt in jnly and Angnst (earlier than gcmixnis wbose

range is similar). 4 bc winter is passed in tbe egg stage.

TYTTÍIUS VAGUS (KNIGIEE) nov. comb.

Gyr/«r///;!i/,v fííWr/s rwgi/s Knigbt, Gonn. Nat. llist. Snrv. Ifnl. 34:

511, 1923.

(Fig. 4 A-E)

Gbaractcrised by its colonr and male genitalia.

Mnle: lengtb 2.8 mm., widtb 1.2 mm.; bead, lengtb 0.2 mm.,

widtb 0.0 mm., verlex 0.32 mm., aniennae, segment I, lengtb

0.2 mm., II 0.8 mm., III ... mm., IV ... mm., jironotnm,

lengtb 0.3 mm., widtb at base 0.8 mm., rostrnm, lenglli 0.7 mm.

Ilead black, sbining, pale spois on vertex starcely apparent.

Antennae witb firsl segment black, apex pale, remaining segments

black; pronotnm and scntellnm black; bemelytra nniformly fnscons,

Carvalho & Sournwooi) — Cyrtoi hiiuis-Mcconinia Coniplcx

the latter slightly ])alcr along cniboliiiin; cuneus and incmbranc pale

fiiscous; legs pale, coxac, bintl fcMiiora exccpt base and more narrowly

at ajiex, íuscons.

Morpbological characters as given for genns.

Gcuilaliii: aedeagns of the Phyline type. Lcft tiasper (fig. 4 E)

with both branebes strongiy pointed. Right clasjrer (fig. 4 D) sbort

and comjxict.

Female: similar to male in colonr and dimensions. Srlerotized ring

and dorsal wall of biirsa copidatrix as in figures 4 15, C.

Distribiítion: New York, Massaclnissets, New Jersey, Virginia,

N. Carolina, Colorado, U.S.A.

Specimens studied: 1 á and 1 9 paratypes, New Jersey,

Lakebnrst (Knigbt’s collection); 12 specimens, Piney Point, Md.,

\'lll ÍIK). A. 1. .Sailer col.; 1 sp]). Hoston, Mass. (named by

Knigbt as C. carieis vaj^us Knigbt.).

'riiis species shows some convergente witb Cyrtorliinus carieis -

(Ortbotylini) bnt is a typical Pbylini in the strnctnre of tlaws and

genitalia. It is closest to pannceps, neolropicalis, pygmaeus and

chinensis, bnt distinguisbctl in the roloiir of antennae and prf)notnm,

as well as in the stnictnre of claspers. .So far mí>st American

specimens seen by tbe sênior antbor and named as carieis atict. are

to be referretl to tbis species.

TYTTHUS NKOTROPICALIS (CARVALHO) nov. comb.

Cyrlorliiniis costae Carvalho nec .Stal, Rev. Hrasil. Hiol. 5 (1): 310,

figs. 1, 2, .3, 1915.

Cyrlorliitius ucoiropicalis C:\r\:\\ho, Au . Acad. Hrasil. Ci. 2(‘) (3-4):

125, 1954.

(Figs. in Carvalho, 1945)

Characlerised by its colonr, dimensions and male genitalia.

Male: length 2.9 mm., width 0.8. mm.; head, length 0.2 mm.,

wiilth 0.5 mm., vertex 0.28 mm., antennae. segment 1, Icngtb

0.2 mm.. 11 0.8 mm., III 0.0 mm.; IV 0.3 mm., ])ronotnm,

lengtb 0.4 mm., width at base 0.7 mm., rostrnm, lengtb

1 .1 mm .

Colonr fnscons to black on head, pronotnm and scntelhnn, the

hemelytra and legs i)ale; antennae blatk (except apex and base of

first segment): hemelytra infnscate on the clavns; pale s])ots ol vertex

obsolete. Rostrnm leaching the middie toxae.

26 ]}oL. Mus. CoELDi — lomo XI (I) — .Sctciiiliro 1955

Genitalia: left claspcr bifuicaic witii scvcral dorsal setae.

Riglit claspcr falciform.

Feinale: similar to male in coloiir and dimcnsions.

Distributio7i: Km. 47 Estrada Rio S. Paulo, Rio de Janeiro,

Viçosa, Minas Gerais, Goias, Hrazil.

Specimens studicd: 3 5 5 9 D. Federal, Rio de Janeiro,

Brazil .

riiis spccies was dcscrihed and ligurcd by tbe sênior autlior in

1945 as Cyrlorliinus costae (Slal, 1860). Alter studyng types in

Stockliolm it was fonnd tliat costae (Stal) is a spccies of Falconia

Distam and not a Cyrtorliiiius as stated by Berguotb (1922). Ihc

name tieotropicalis was preposed for tbe sjíccies An. Acad. Brasil.

Ci. 26 (3-4): 425, 1954. It can bc distinguisbed from otlier spccies

by tbe blac ring oí tbe lirst antcnnal segmcnt, being closct to

lYjgi/s Knight which lias tbe lirst antcnnal segmcnt largerly black aiul

pale spots on vcrtex well marked.

TYTTHUS MUXDULUS (BREDDIN) nov. comb.

Periscopus mimdulus Breddin, Dcnt Ent. Zcit. . 106, 1896.

Cyrlorliinus muudulus Reuter, Ofv. E. Vet. .Soc. Eorli. 14: 178,

1902, Zimmerman, Ins. llawaii, 3, Ilet.: 206, fig. 88, 1948;

Gsingcr, Soc. .Sei. Ecmi. Gominent . Biol. 12 (8): 4, fig. 1951.

(Fig. in Zimmerman, 1918)

Cbaractcrised by its size, colonr and genitalia.

Male: Icngtb 3.5. mm., width 1. 0 mm.; bead, widtb 0.7 mm.,

vcrtex 0.36 mm.; antennae, segmcnt 1, lenglli 0.35 mm., 11

1.10 mm., 111 0.62 mm., IV 0.5 mm.; pronotmn, lengtb 0.38

mm., widtb 0,85 mm., rostrmn lengtb 1.1 mm.

llead black-brown cxce|)t for two ])ale arcas adjacent to tbe cycs,

orcasionally entirely dark; basal segmcnt of antennae jiale fulvous

witii tbe apex sometimes darker, other segments nsnally fnsemis;

pronotmn and sculellnm blaik-brown; liemelytra pale grecn suflnscd

svitli brown towards lhe sntmc, ccll nervnres brown; legs ycllow-

grccn to pale fulvous; rosirum pale grecn tip daik; uuderside

brown blat k .

Pubcscence of jiale line bairs espccially long at tbe posterior of

bead and anterior of tborax; m:icro|)tcrous.

Carvalho & Solthwooi) — Cyrtoihinus-Mecomnia Coniplex

(ieuilalia: aecleagus of j)hyliiic typc (íig. 5 E), leít claspcr

(fig. 5 C) wilh tenninal jjrocess stouter tliaii in germinus, right

claspcr (fig. 5 G) siinple.

Female: length 3.25 iiim.; width 1.2 inm.; licacl, wiclth 0.75 mm.,

vertex 0. 38 mm.; anlcmiac, 1 0.35 mm., 11 1.0 mm., III 0.50

mm., IV 0.35 mm.; j)ionotiim, length 0.‘I5 mm., width 0.00

mm.; rostrum lengtli 1.0 mm.

Coloiir and pnbcsccncc as in male; macropteroiis.

Distrihuliotr. Java, Eiji, Philippinc Ils. Qiieensland (Anstra-

lia), (Introdmctl intto Ilawaii).

Sjjeciinens xhtdied: 5 á -1 $ Natoiia, Eiji, April 1910, R.

Vcitch; 2 $ Lahaina Mani, Ilawaii, .Swc/cy col. Dec. 1028; 2 9

Ilalifax, Qnecnsland, i\nstralia, April 1020, E. iMnir; 1^19 Los

Banos, Philippinc Ils., Williams col.

The largcst sjtccics of Tytllius, distingtiishcd by its palc basal

antennal scgmcnt and general dark. colonration.

Aceording to Zinnnerman (1018) “this species was introdiiccd in

Ilawaii in 1020, from Qnecnsland and Eiji, to aid in thc control of

thc sngarcanc leafhop|)er, PcrkiiisielUi .sncliaririda Kirkaidy. It

herame estahiished and conslitutcs one of the ontstanding rccords

in the history of hiologiral control. It has saved thc llawaiian sngar

indnstry the 'Eeiiitory millions of ilollars— its trne worth can hardly

be estimatctl”.

The pretiatory hahits of thc species wcre discovercd by Dr.

Mnir in Qnecnsland (1020) (.Swe/ey, 1930).

It may also fcetl on thc eggs of Peyegriinis iiKiidis (Ashmcad) on

corn :ind Mcgamchis Inoscrpitui Kiikaldy, on taro.

ryrrnus aiaíoornatus (RNiíiiee) nov. comh.

C.yrtorhiuus nlbomnntus Knight, Bnl. Brook. Ent. .Soc. 20 (1); 172,

1031.

(Eigs. 0 A-Ci, 7 A)

Charticteriired hy tlic dark colonr witli basal two fifths of hemelytra

pale whitish, (imens likewisc palc and male genitalia.

Male, inacr(>l)leroiis: length 2.3 mm., width 0.8 mm., hcad,

Idiglit 0.1 mm., witith 0.5 mm., vertex 0.30 mm., antennac,

scgmcnt 1, lenght 0.3 mm., II 1 .0 mm.. III 0.7 mm.. IV 0.7

mm., pronotnm. length 0.3 mm., width at base 0.7 mm.,

rostiinn, length 0.0 mm.

Bol. Mus. Goei.di — roiiio XI (1) — .Setembro 1955

Colour dark brown to black, basal two liftbs of liemelytra atui

entire cuneiis jrale whitisli; distai hall ol coxae and bases oi femora

pale to wbitisb; tibiae and tarsi pale Inscoiis, spines black; inembrane

and veins ratlier imilonnly pale luscons.

brachypterous: lenglit 1.7 inni., widtb 0.7 inm., hcad, length

0.2 inni., widtb 0.0 nnn., vertex 0.50 nnn., antennae, segment I,

length 0.2 mni., II 0.7 nnn., III 0.2 nnn., IV 0.2 nnn., pronotinn,

length 0.3 nnn., widtb at base 0.0 nnn., rostrinn, length 0.7 mm.

Colour black; basal two thirds antl apical one sixth of

hemielytra, white; first antennal segment (excei)t extreme base),

coxae, extreme bases and ápices of iemora, the tibiae towards the

apex and rostrnm, sordid yellow.

I lemelytra withont indication of cnneiis or inembrane; rostrimi

reaching the posterior coxae; posterior femora mnch longcr and

stonter than others; pubescence short and semiadpressed .

Genilalia'. no macropterons fornis were available for dissection.

In the brachypterous forni the aedeagns is of the IMiyline type (fig.

0 !•■). Left clasper (fig. 0 (i, 11) as seen in ilhistration, with tlie

arm somewhat laminate and nuicli less sclerotized, the left arm end-

ing by a sjnnelike ])oint. Right clasper very small. Pygojihore

(fig. 0 B, D) as shown in illnstrations; the sheath is noticeably

pointed (fig. 0 E).

Femole: similar to inale in colour, slightly more robust.

Distrihutioii: Florida, New York, IJ.S.A.

Specimtnis sliidicd: I <} holoty|)e, Jacksonville, Florida

(K.night’s collection); 2 á brachy|)terons, l itnsville, Floriíla, -1 .2.52,

II. C. Cihapman.

riiis sjiecies can be easily recognized by lhe whiiish base ol

liemelytra and ciinens, which seeiii to be contant aiso in the brachyp-

terons form .

TYTTIIUS GEMINUS (FLOR)

Cdpsus goninus Flor, Rhync. l.ivi. I; (iOti, IHtiO.

Tytllius grmitwis Fieber, VVien. Fnt . Monat . 8 (3); 83, 18(il.

Cyrlorhiiuis gemiinis Renier, liem. Cymn. Fnr. 3: 382, 551, pi. 2,

fig. 5, 1883; Wagner, Fierw. Dent. 11, Bliiidw.: 129, 1952.

Clilotnydahis (C.yrlorhiuiis) gemiiius Heuter, Rev. Crií. Caps. 2:

I2(i, 1875.

* C.yrlorliitnis puhcscni.s Knight, Bnl. Brook. Fnt. .Soc. 2() (1):

172, 1931 ( 11 . syn..).

(Figs. 8 A-C)

SciELO

10 11 12 13 14 15

Carvalho & Southwooi) — Cyrtorhinus-Meconinia Complex

Charactcriscd l)y its colour, the long black hairs and inale genitalia.

Male: length 2.(i‘nnn., width 1.0 inin.; hcad, width 0.65 mm.,

vertex 0.'ò"ò mm.; anteimae, scgment I, length 0.38 mm., II

1.0 mm., III 0.79 mm., IV 0.65 mm.; pronotnm, length 0.33

mm., width 0.76 mm.; rostriim length 1.15 mm.

Head hlack with two pale areas adjacent to the eyes; antcnnae

dark excejit for hasal joint which is only dark at extreme hase;

pronotnm antcriorly dark fading to pale hrown at posterior,

scutellmn hlack or hrown, darker in the median arca; hcmelytra

jiale grccn-hrown, meinhranc pale; legs cntircly jialc yellow-green,

rostrinn yellow-grecn, dark at apex; plenrites and sternites of tliorax,

dorsnm and lateral margin of ahdomen and pygophorc dark hrown,

extreme vcntral arca of ahdomen pale grccn-yellow.

Piihcscencc, long (0.19 mm.) dark ercct hairs on posterior of

hcad antl anterior of pronotnm (fig. 8 15), short dark and mostly

adprcsscd hairs on the legs, othcrwisc pidiescencc of short adpresscd

mixed pale aiuI dark fine hairs; macropterous.

Ciciiilalid: aedeagns (fig. 8 E) simplc.

Uemale: length 2.5-3.25 mm., width 1. 05-1. 25 mm.; hcad, width

0.75 nnn., vertex 0.35 nnn.; antcnnae, 1 0.10 mm., II 0.90

mm., 111 0.75 mm., IV 0.7 nnn.; pronotnm, length 0.'12

nnn., width 0.85 mm.; rostrum length 1.18 nnn.

Colonr as in the male, hnt pronotnm sometimes cntircly dark

an with the dorsnm and lateral areas of the ahdomen dark, the

entire venter pale yellow-green.

Pnhescence as in the male. .Semi-hratliy|)tcrons, hcmelytra rcach-

ing to the h:ise of the 7th abdominal scgment, mcmhranc rcdnccd.

Distributioii: England, N . Rnssia, Finlaiul, Sihcria, Livonia,

N. Cermany, Dcnmark, .Swcden.

Sl)C(iiiH-iis sltiditul-. 1 9 Wickcn Ecn. Camhs., 19/9/31, II. .St.

J. Donisthorpe; I <í 2 9 Madcley, Stalfs., 22/9/31 11., \V. Daltry;

2 9 Einlaiul, R. l.innavnori; I á Porgos, Einland, |. Sahlhcrg.;

1 á .“l 9 .\nchoragc, Alaska, Vlll, 918, R. I. Sailer col., 1 9

Coloratlo, Uhici col.. holotypc (piihesccus), Wray, Colorado,

Knighl’s collcclion.

riiis .sj>ccics is casily distingnishcd hy the long hlack hairs, the

cntircly ycllow legs, the pionotnm nsnally |)alcr |)ostcriorly and the

hasal scgment of ilie antcnnae ahnost cntiicly pale. .Semi-hrachyp-

teions males and inacroplerotis feniales are known (Wagner 1952),

SciELO

10 11 12 13 14 15

30 15ol. Mus. Goeu)I — Tomo XI (I) — Setembro 1955

but none wcre available for study. It is associated with varioiis

species of Carex and is found as an adidt iii September and October

(Butler, 1923).

Tbc anionnt of dark colour on head and jjronotuni varies

within tlie species.

TYTriIUS BALIA (KNlGin ) nov. comb.

Cyrtorliimis balli Kniglit, I5ul. Brook. Ent. Soc. 26 (1): 171, 1931.

(Fig. 5 A, D F)

Cliaractcrized by its tolonr, sliort antennal segincnts and bi-coloured

aspcct of tbe beinclytra.

Male: lengtb 2.7 inm., width 0.7 mm., head, Icngth 0.1 mm.,

width 0.6 mm., verlex 0.32 mm.; antennae, segment I, Icngth

0.2 mm., II 0.() mm., III 0.7 mm., IV 0.5 mm.; pronotum,

lengtb 0.2 mm., width at base 0.6 mm.; rostrnm, length

1 .0 mm .

lleatl Idack, shining, a jtale spot cach side of vertex Itordering

cyc; antennae black, a trace of palc at ti]) of sccond segment;

pronotum i)ale fuscous to blackish, anterior margin of disk orange

ycllow; mcsosctitnm atui scutelium dark fuscous, witii orange tint

in hypodermis; hemclytra pale and fuscous, inner half of clavus

l)ordcring scutelium, ajtical half of corium, apical half of embolium

except tij), and ajtical balf of cuncus fuscous to blakislij membranc

rather unílormily pale fuscous, a shade darker on areoles atui veins.

Underside of body brownisb to black, abdômen pale beneath, genital

segment black; legs i)alc to orange coloured, tibiae and tarsi blackish.

(ic)iilali(i: aedeagus of the Phyline type. Left clasj)er (fig. 5 D)

with ápices of branches short and pointed. Right clasper (fig. 5 F)

elongate with a jtointed outgrowth near apex.

Lainale: similai to male in colour and dimensions.

Disirihiilinn: Florida, 'Fexas, U.S.A.

Sfirriínnis studicd: I í 1 9 paratypes, |a( ksonville. Florida

(Knight's ( ollection); I J , .México (an roses), intercepted llrown-

sville, 'Fexas.

I his speties is wcll distinguished amongst the others in the

genus by ibe anterioily narrowed |>ronotum and (olour pattern of

the body. It is nearest to C. insprratus Knight, especially in antennal

lengtb and shape of pronotum, but dilleis in tolour and si/e.

Carvalho & Southwood — Cyrtoiliimis-Mcconima Complex 31

TYrriIUS INSPERATUS (KNIGHT) nov. cob.

Cyrtolihinus insperatus Knight, Bul. Brook. Ent. Soc. 30: 43, 1925.

(Fig. 9)

Cliaracterized by its coloiir and male genitalia.

Male: length 3.0 mm., widlh 0.8 mm.; hcad, lengtb 0.2 mm.,

■width O.G mm., vcrtcx 0.28 mm.; antcimac, scgment 1, length

0.3 mm., 11 1.5 mm., 111 1.0 mm., IV 0.5 mm.; pronotum

length 0.3 mm., width at base 0.7 mm.; rostrum, length

1 . 0 mm .

Head black with a small pale ocellate spot cach side of vertex

above eye; rostrum reddish yellow, basal segment greenish, a])ical

segment black; antennae black, lincly pale to dusky pidjescent;

pionotiim brownish black, becoming fulvous basally, anterior margin

pale, scutelhnn and mesoscutum fulvous, the mesoscutum broadly

exposed and tinged with fuscous; hemelytra semitranslucent, fumate,

basal area of corium aiul narrow margin of clavus pale transluccnt,

claviis fuscous, cxcept aloug claval suture, the scutellar margin and

slcnderly along commissure fulvous; membnme and veins uniformly

fumate, anal area darker; steruum fulvous, sides, pleura and ostiolar

|)eritreme becoming fuscous; legs ])ale fulvous, tibiae and tarsi black,

femora with fuscous linc along dorsal margins, also a reddish to

fuscous longitudinal linc on anterior face, this line being more

sidtvcntral on fiont femora; abdômen greenish with a mctallic luster,

genital segment black.

(Irnitaliir. no male specimens werc availablc for disscction.

Fcmalc: more robust lhan male but similar in colouration.

Distyiliulioii: Arizona.

Spccivifus stiulifíl’. 1 á iH'd 1 9 paialypes, 1 ueson, Arizona,

A. .\. Nichol (Knight's collection); 1 <{ . I 9. Calexico. Calif.

Bali col., 1 <5, Buckcye, .\rizona, [ohnsou col.

'I his species is easily recognized by its very long second antcnnal

antennal segment and the white tiansverse area on anteiior portion

of pronotum. It shows also a prolrudiug clypeus and a short ncck.

(haraclcis which set it apart from the remaming species and probably

from the genus. Once specimens are availablc for disscction, it will

bc possiblc to place it with (Citainty.

Boi.. Mus. Gokldi — lonio Xi (1) — Seteniliro 1955

TYTTHUS MONTANUS n. sp.

Characteri/,cd by its coloiir, shapc of pronotuni aiul inale gciiitalia.

Male: lengtb 2.7 inni. widtli 0.8 mm. llead'. Icngth 0.2 mm.,

width (i.O mm., vcrle.\ 0.33 mm. Anteiifuie: scgment I, lengtli

0.2 mm.; II, 0.9 mm.; 111, IV, brokcn. Pronolum: Icngth

0.1 mm., width at hasc 0.7 mm.

Castancons to dark hrown; liead, pronotnm and sctUelltnn

(cxccpt jiale arcas on vcrtc.x) dark hrown; antcnnac ])alc ycllow,

sccond and two last joints ítiscons towards apcx; licmclytra grcyish

hrown with hasal third wliitish to palc ycllow, mcmhranc palcr

towards apcx; nndcrsidc of hody castancons, Icgs iialc ycllow cxccpt

hascs of coxac which arc rcddish and apical jiortion of fcmora

(csj)ccially hind pair) which arc castancons to dark hrown, tihiae

sligluly darkcr towards hasc.

Rostnim rcaching thc hasc of hind coxac. llcad ronndcd in

front, clypcns not sccn from ahovc, cycs placcd ncar middle of hcad,

distant from pronotnm hy a s|jacc ctpial to thickncss of first antcnnal

scgmcnt, pronotnm noticcahle constrictcil hchind middle, disc

convex, smooth, calli largcr hnt not marked, lateral margins hroadly

ronndcd and converging antcriorly, posterior margin s innatc

intcrnally, hnmcral anglcs ronndcd; pnhcsccncc scanty and short,

mcsoscntnm hroadly exposed.

Geuitnlia: aedeagns of thc connnon Bhylinc typc. Lcft claspcr

(fig. 7 C, F) as sccn in fignres, thc longcr arm with many hairs on

externai margin. Right clasiicr (fig. 7 B, D) somcwhat laminatc, as

sliown in fignres.

Pemale: nnknown.

í!ololyl>e: inale, Drnnimoiul. Montana, Oman col., VI 1. 935,

in thc (oilcction of thc U..S. National Mnscnin.

‘I his spccics shows a peculiar ])ronottmi hnt slill can hc mantai-

ned in thc genns Tyttlius Ficher, since lhe posiiion ol lhe cycs

and shapc of pronotnm varies within thc genns. It differs from thc

othcrs kiiown so lar in thc colonr and thc ahovc mcniioncd

{ haracters .

ryrrnus paxami-nsis n. sp.

Charactci i/cd hy its si/.c, (olonr and inale genitalia.

Mair: Icngth 2.1 mm., width ().(> mm. Hcnd: Icngth 0.2 mm.,

width 0. 1 mm., vcricx 0.23 mm. Anleiiuar: scgmcnt I, 0.2

mni.; II, 0.9 mm.; III and l\', hrokcn. 1’ionotnm: Icngth

0.2 mm., width at hasc 0.5 nnn.

Carvalho &: Soutiiwood — Cyrtorhinus-lMccomnia Complcx

IlcacI, proiiotum and first antennal scgmeiit (cxccpt extreme

apex) black lo dark brown; reinaiiiing aiiteniial segments, ápices

of íeniora and bases of tibiac castaiieoiis to íulvons; hemelytra pale

to greenish yellow, niembrane translncent; rostrum, coxae (except

for reddisb tinge on first pair), bases of femora and ápices of tibiae,

pale yellow; nnderside of body (except nicdian portion of abdômen)

dark brown to castaneous; tlie ligbt spots on liead obsolete.

Rostrum reaching base of hind coxae, body elongate, hemelytra

parallel sided.

Genitalia: aedeagus of the common Phyline typc. Left clasjier

(fig. 7 G) as seen in figure. Right clasper (fig. 7 E) with a long

setae on truncate apex.

Fernale: unknown.

Holotypc. inale, Corozal, C. Z. Panama, A. lUicsk col. (at

ligbt).

Paratypes: 3 males, same data as the type, in the collection

of the U.S. National Musenm and of the author.

'I his species is near pnrviceps in colonr bnt with more elongate

body, the spots on vertcx obsolete and a typical left clasper. From

balli and alboornntus it ditfers in colonr and sirnctnre of claspers.

GENUS EIEIiEROCAPSUS nov. gcii.

1 lead rounded anteriorly, face short, distance from base of

eye to ajiex of tyhis half height of eye: antennae with the first atui

secoiul segments at letisl half as broad again as the third and fourth;

pronotnm trapezeform, straight sided antl only slighily wider

posteriorly than anteriorly (Fig. 10), its lower margin nsnally

slightly concave, calli only slightly raiscd; hemelytra with the

cunens broader than long; covercd with pale semi-crect hairs,

longer (average 0.12 mm.) on head and anterior of pronotnm

atui shorter on hemelytra atui legs (average 0.08 mm.), occasiotial

spinose hairs oti hind tibia, rostrum reaching to apex hind coxa.

Genitalia comparatively large, the left clasper simple and cnrved,

the right roniuletl with a niedian row of the tecth; the vesica ol the

aedaegus has a pair of aj)|)endages arising from near its apex, as

well as the more basal vcsical appendage; in the fernale the K

stiticlure is large and its postei ior median concave. Hrachypterism

otcurs in both se.xes.

Fype species: Fyltlius jUweolus Renter 1870.

Althongh snperficially similar to Cyrtorhiuus aiul Mi-romnin,

this gentis is sharply tlisiingnesheil from iheni on genitalia, in the

stiiutnre of lhese it shows alfinities with Cylleroris llahn and

Dtyophilocoris Renter .

— 3 _

31 Boi- Mus. Goi i.di — 'lomo XI (1) — Setembro 1955

FIEBEROCAPSUS ELAVEOLUS (RliUlER) nov. comb.

Tyttlius flaveolus Reuter, Not. Sallsk. F. Fl. Fenn. II: 323, pl. 1,

fig. 6, 1870.

* Tytthus iusiguis Reuter iiec Douglas &: Seott, Rev. Crit. Caps.

2: 126, 1875 (syn. by Reuter, Ent. mon. Mag. 1 1: 131, 1877).

Cyrlorhinus flaveolus Reuter, Hem. Gymn. Eur. 3: 380, 55-1, 1883;

■Sauiulers, Hem. Het. íbit. Is.: 281, 1892; Wagner, "Fierw.

Deut. ‘11, Blindw.: 128, 1952.

(Fig. 10 A-H)

Cbaracterized by its sbape, tolour and genitalia.

Male: lirachypterous: lengh 2.8 mm., widtb 1.50 mm.; lieail,

width 0.91 mm., vertex 0.‘10 mm.; antennae, segment 1, lengtli

0.‘M mm., II 1.2 mm.. Hl 0.75 mm., IV 0.5 mm.; pronotnm,

length 0.52 mm., width 0.97 mm.; rostrnm 1.3 mm.

Entirely pale yellow except for antennae, tbc first and second

segments of which are brown with ])ale backs, wliilst tbe tbird and

fourtb segments are pale grey-brown; tbe tylus is dark brow as are

two areas on tbe frons (lo wbirb tbe leeding pump muscles are

attacbed); lateral region of pronotnm and meso— and meta— tboracic

jiletirites snffnsed witii brown, tip of rostrnm dark brown.

Bubescence of ])alc fine adpressed Iiairs. Bracbypterous,

bemelytra reaching just bcyond posterior of eigbtb segment,

incinbranc and ttineal snture absent.

Genitalia: aedeagus (fig. 10 B) of orthotylinc lype with

straplike tootii vesieal ajipendages; left dasper (fig. 10 I), E) narrow,

its apex prodiued and grooveil; rigbt elaspers (fig. 10 ;\) broatl

with a teniral tootbed sbelf-like projedion.

Macropterous: length 3.5 mm., widtb 1.17 mm.; jironotum,

length 0.52 mm., widtb I.I5 mm.; otberwise as in bracliyp-

terons inale.

Eeinalc: brachyptcrous: length 2.97 mm., widtb 1.75 mm.; bead,

widtli 0.91 mm., vertex 0.‘I5 mm.; antennae, I 0.13 mm., 11

I.I5 mm., 111 0.72 mm.. I\' 0.5 mm.; pronotum, length 0.5

nnn., widtb 1.12 mm.; rostiiim length 1.28 mm.

Golour :ind pubeseeiue as in m;de.

Genitalia: K strudures sliaped in a «urved 1.. tpiile distinct

from any oiber s|>eeies (fig. 10 I').

Maí^ropterous: length 3.75 nnn., widtb 1.75 nnn.; pronotum,

length 0.62 nnn., width 1 .35 nnn.; oilierwise as in braebytemus

lemale.

Carvalho &; Solithwooi) — Cyitoiliimis-Mccomina Complcx

Englaiul, .Svvcdcii, Finlancl, Dcnniark, N. Rússia,

Houclslow Hcath, Middlesex,

S, Finland, Aug. 1947, R.

Distribiitioii:

Gci niany .

Specimens stiidied: 6^09

U.K, 20/7/53, G. F. WoodroLíe, I

Linnavuori; 1 9 Finland.

'l liis spccies is casily recogniscd by its eniircdy ])ale coloiation.

'Fhc adult, wliicli is íoiind íroin July to Scpleinbcr (lUitler, 1923),

is said to bc phytojjbagoiis (Wagner, 1952) (see bowcvcr p. Gl).

It is loiind at tbe bases oi reeds and 1 iislies; lhe braeliypterous ionn

is tbe connnonest.

C.ENUS CYRTOR/ÍINUS FIEBER

Cyrtorhiniis Fieber, Wien. Fnt. Monat. 2: 313, 1858; Reiiter, Ilem.

Gyinn. Ftir. 3: 379, 1883; Distant, Faiin. Hrit. Ind. Rhynç.

2: 47G, 1904; Hueber, .Syn. Hlindw. 2: 10(i, 1908; Popjtius, Acta

Soc. Sei. Fenn. 44 (.3): 70, 1914; Wagner, 'Fieivv. Deut .41,

Blindw.: 127, 1952.

* Cyriorrhima Reuter, Acta .Sf)c. .Sei. Fenn. 13: 379, 1884 (emenda-

tion).

* Reuleriessd Usinger, .Soc. .Sei. Fenn. Connnent. Biol. 12 (8): 3,

1951 (7WV. syu.).

'Fype spccies: Capsus elegaiiluliis Meyei, 1843, a synonyin oi

Cyrtorftititm carieis (Fallen, 1807) — iTionobasic.

Sntall to inedinni si/.ed biigs (2. 5-4. 5 inin), liaed rounded

antciiorly, lace senii-vcrtical; antennae aiising adjacent to anterior

inargin oi eyes; pronotinn cainpanuliioi in witii calli sligbtly raised,

its lower inargin straiglU; posterior oi pronotuin sligbtly wider tban

liead; opening oi oiliieroiis gland raised; pretarsus with ilaplikc

arolia; rostrtnn reacbing base oi inid-ioxa; inale jiygopbore with

siibgenital plate or adeagal su|)port only sligbtly projecting; bursa

copiibitrix of 1'einalc witli rounded or senii-tpiadrale K striu lures.

Gosered with siin|)lc seini-ered pubesctauc (0 . 08() . 1 5 nnn. in

lengtb;) tolour p.ittern generally black and pale green. I'einale

soinetiines partially brarhypterous.

KFV 10 1 IIF SBFGIF.S OF CVlCrORII/NUS

1 . Settind antennal segment aliout twice as long as pronotal widtli

at liase (1.5 : 0.8) 2

Seiond antennal segment noticeably less tban twite as long as

pronotal widtli at base (1.5 : 1.0 to 0.8 ; 0.7) 3

SciELO

10 11 12 13 14 15

3G l}oi>. Mus. CioKi.Di — loiiio XI (1) — Seicinljio 1955

2. Coloiir tliiclly fulvoiis, apcx ol lirst aiitcnnal scgnicnt j^ale ..

julvits Kniglit

Coloiir palc ycllow giccii, aj)cx ol lirst antcnnal scgiiicnt clark

ctunberi Woodwarcl

• 3. Sccoiul antcnnal scgmcnt liall as long again as width ol

pronotnin at liasc (1.5 : 1.0 nnn.), riglit claspcr ol inale not

bilid, basal scgmcnt ol antcnnac cntiicly black; pronotinn and

scutcllinn cntirely black, bcad black with only two sniall pale

arcas adjaccnt to tbc cycs carieis (Fallen)

.Sccond antcnnal scgmcnt only sligbtly longcr tban tbc pronotuin

is wide at its base (0.8 : 0.7 nnn. or 1.18 ; 0.93 nnn.), right

claspcr ol inalc bilid, basal .scgmcnt ol antcnnac palc at apex;

(pronotnin and scntcllmn oltcn partially jialc; bcad gcncrally

with wide palc arcas adjaccnt and posterior to tbc cycs) .... 4

4. Small spccics (lindcr 3.0 nnn.), tibiac cntirely ycllow

lividipennis Renter

Larger spccics (over 3.5 mm.), tibiac gcncrally inluscate

cspccially at base melauops Renter

CVIiTORIlINUS CUMliEIU WOODWARD

Cyrtorltinus ciirnberi VVoodward, Rcc. Anck. Inst. Mus. 4 (1):

9-23, 1950.

(Figs. II A-G)

Cliaractcrizcd by its long sccond antcnnal scgmcnt, palc colourcd

jironotum ol thc inale and gcnitalia.

Malc: Icngth 3.85 nnn., width 1.0 nnn.; hcad, width 0.75 mm.,

vertex 0.1 mm.; antcnnac, scgmcnt 1, Icngth 0.5 mm., II

1.5 mm., III 1.34 nnn., IV 0.55 mm.; pronotinn, Icngth

0.18 nnn., width 0.84 mm.; lostrmn Icngth 1.0 mm.

I’alc ycllow grccn, cxccpt antcnnac (apart Irom extreme hasc

ol lirst scgmcnt), anterior and lateral regions ol hcad, lateral arca

ol thorax and apices ol tarsi and rostrmn which are hlack.

1’nbc.sccticc jialc and linc; macropterous.

(tcnitalin: aedeagns lig. II l.) ol orlhotylinc typr, thc thcca

with a dorsal projettion and thc vcsital ap|>cndagc trnncatc ;u its

apex; light dasper (lig. 1 1 G) with a ciirvcd jaotess which is tcMithcil

at its apex, Iclt tla.s|>ci (lig. II D) with a long simplc curved

|)I ()( css.

Carvalho &: Southvvood — Cyrtorhinus-Mccomma Complcx

Female, liracfiypleroiis: Icngth 3.84 inin., wiclth 1.25 mm.; hcacl,

width 0.78 mm., vertex 0.40 mm. Antciinac, I 0.5 mm., 11

1.33 mm., III 1.07 mm., IV 0.55 mm.; jnonotiim, length

0.5 mm., width 0.85 mm.; rostnim length 1.1 mm.

Head red with anterior and lateral regions hiack and two areas

oir the vertex, median to the eyes, yellow; antennae with hasal joint

red, second red-brown hecoming darker at apex; pronotnm yellow-

gieen suHiised with red aiul anterior and lateral regions hiack; legs

and scutellum yellow green; hemelytra jxde green; nnderside yellow-

green; tip of rostrnm tlark.

Pnbescence as in male; brachypterons, hemelytra reaching to

end of seventh abdominal segment, membrane mnch rednced.

Genitnlia: K striictiires (fig. 11 C) semi tjiiadrate.

Macropterons not seen, based on Woodwartl (1950); length 3.0 mm.,

width 1.3 mm. As brachypterons female Itnt hemelytra reach-

ing, bnt not entirely covering the ninth tergite.

Distribution: New Zealand (N. Island).

Specirneiis sludied: 1 á , Paiaka, Manawatn, New Zealand,

2/2/51, T. K. Wootlward (1$. M.); 4 $, tlitto, New Zealand,

4/1/50, T. E. Woodward (15. M.).

This sitecies is very similar to jnlvus Knight in jtroportions,

bnt is distingnished by ils coloration and genitalia. It was fonnd

below and in tidts of riishes and grasses, where Deljíhacids occurred

abnndantiy (Woodward, 1950).

cYitroRiiiNUs lui.vvs KNicirr

C.yrtorliiniis ftilvus Knighl, Ins ..Samoa, II, Ilem. 5: 205, 1935.

Cyrlor)iinus fulvus Zimmerman, Ins. Ilawaii, 3, lleterojjt.; 205, lig.

8, 1948.

(Eig. 12 A-E, (Ml)

Cliaracteri/ed by its colonr, si/e, length of second antennal segment

and genitalia.

Mnir: lenglli 3.4 mm., width 1.0 mm.; head, length 0.2 mm.,

width 0.7 mm., vertex 0.34 nnn.,; antennae, .segtnenl 1, letigth

0.4 mm., II 1.5 mm., lH 1.3 tmn., IV 0.0 mm.; [jionotitm,

length 0.4 mtn., width at base 0.8 mm.; rostritm, length

1 . 1 mm .

SciELO

10 11 12 13 14 15

38 J5()i,. Mus. CioKi.Di — Tomo XI (I) — Sctcml)ro 1955

General coloiir lulvous, antennae except ajrex of segment I,

liead exce])t on verlex and genac and loia, pronotinn except niedian

ray on basal lialf ol disc, mesonotuin more or less, and median line

ol sentellnm, blac.k; legs lulvons, luscous on knees; membrane pale to

dusky, cnbitns tnscons.

Rostriim readiing lhe middie coxae. Gcnitalia witb aedeagns

sbowing a typical s|)ic'nlinn (lig. Ili 11). Lelt dasper (lig. 12 D)

branebed at middie, ibe lower arm cnrved, tbe lipper lobe witb

dorsal setae. Rigbt dasfier (íig. 12 G, 11) also branebed, one lobe

witb setae, tbe otber ended by a serrate margin. Pygopbore (lig.

12 G) as .seen in lignre.

Uemalc: lengtb 3.8 mm., widtb 1.3 mm., sligbtly more robiist

tban male biit very similar in striictnre and colonration . K

strnetnre as seen in lignre 12 K.

DislribxUioiv. Micronesia, Garoline Is. (Palan, Ilabeltbnap,

Vap, Koi'or), .Samoa, Java, Pbilippines, Idji, New Gninea (Introdueed

into Ilawaii).

Sf)ccimeiis sludicd-, 1 á (Ilolotype) Savaii, Samoa, Lower

lorest (l()()()-2()()() It.), P. II. llryan (Ibit. Mns.); 3 <} (Paraty[)es)

Savaii, Samoa, K. II. Ilryan (Ibit. Mns.); 2 $ (Paratyjjes)

iMalololedei, dpolii, Samoa 1/5/21, P. .\. Ibixton and G. 11.

llopkins (Ibit. Mns.); 2 $ Snva, Idji, 7/5/13, R. Lever; 3 é

New Gninea, Pemberton col.; I <5 java, P. Mnir eol.; 1 5 Los

Ibinos, Pbilijipine lis. IL Mnir eol.; I i llonolnln, Ilawaii; 9 ^

■11 9, CAROI.INP ll.S., hilatr. Ibdidlbuap Is., 7/12/52, j. L.

Giessit; Yaj) Croiip: ^'apid, Goloni;i, Ronil, S. \’apid, Pomil Dist.,

jnly 1950. R. J. (brss. Koror; 30/11/17, 11. .S. Dybas; 15/3/18

K. L. .Maekler; 20/1/53, L. \V. Ibiirdsley; IUil/cltliU(iiil>: Kaisban,

Ang. 1939. P. Pisaki.

CYR TO R lííN US I.ll d IHriüVXIS R P. 1 1 ’ PP R

Cyrlorhiiixis lividipniiiis Renler, Pnt. l idskr. 5: 199, 1881; Distant,

laiin. Ibit. Ind. Rbyne. 2: -no, lig. 308, 1901.

* Cyrloiliiiiiis xnliciisis Lsinger, Soe. Sei. Penn. Gonnn. lliol. 12

(8); 3, ligs. I. 2. 1951 (nov. .syn.).

(Pigs. 12 P, 13 A-íi)

Cbaraeleri/cd by ils si/e, lengtb ol second antennal segment ;nid

geniialia .

Mair: lengtb 2.50-2.78 mm., widtb 0.95 mm.; be;id, widtb 0.0

mm., vcilex 0.27 mm.; anieimae, segment I. lengtb 0.3 mm.,

II 0.80 mm., III 0.78 mm., IV 0.52 mm.; |)ron()inm, lengtb

0.33 mm., widtb 0.78 mm.; loslinm lengtb 0.75 mm.

CIarvai.iio &: Soiniiwooi) — Cyrtoiiiimis-Mccoinma Comijlcx

Ilcail, pronotiim and scntclliim pale ycllow witii a variablc

amoiiiu ()[ black or dark hiown niarkings, ranging from ainiost

ciuirely dark to entirely liglit; antennae dark cxcejit for apex ot

Itasal aiul base of segnient; lieinclytra pale green, menibranc pale

grey with ncrvnres grey-green; legs pale ycllow; rostruin pale green,

tip dark; nndersiile of thorax, and all abdômen pale yellow-grcen .

Pnbescence of sliort pale adprcssed hairs. Macroitterons.

(ieniíalia: aedeagns (fig. 12 K, 13 C) of orthotylinc tyite, the

single vesical ajtjtendage sharply widened on one side and then

narrowing again beforc the tij); left clasper (fig. 13 F) with an

elongated enrved arm with tliree teeth on th eventral aspect, as in

fulvus; the right elasiter (fig. 13 H, G) with a sliort ctirved process.

Femalc: Icngth 2.75-3.0 nnn., width 1.1 mm.; head, width 0.03

mm., vertex 0.31 mm.; antennae, 1 0.26 mm., II 0.78 mm.,

III 0.71 mm., 1\' 0.58 mm.; jironotnm, length 0.33 mm.,

width 0.85 mm.; rostrnm Icngth 0.7() mm.

Colonr and |mbescencc as in male. Macropterotis.

GenitoUif. K strnctnres (fig. 13 D) ctirvetl.

Dislrihiítiow. S. Iiulia, Ccylon, lUirma, Gt. Nicobar, Ghina,

FoVmosa, Jap.in, IMiilipjiine lls., Java, Snmatra, New Gninea,

Marianas lls. (Gnam, Rota), Caroline lls. (Yap, Babeltlmap, Koror,

Vajiid), Samoa, New llcbrides (Malcknia), llornco.

Sfx-cimeus sludicd: 1 $ (lloiotypc of xiitiaisis Usinger)

Dobnilevn, Fiji (swecping yonng rice), 15/0/18, H. A. 0’Conncr

(llrit. Mnsenm); 2 <í Dobnilevn, Fiji (sweeping yonng rice),

15/0/18,1$. A. 0’G()nnei; 10 á.9 9 Nadnrolonlon, Fiji, (sweejiing

yonng rice), 10/10/50, 1$. A. ()'Gonner; I <5 2 9 Mokassan,

(iliekiang, Ghina, Dora F. Wright; several á, 9 San joe, Mindoro,

IMiilippinc lls., March 1915, F. S. Ross; Fos Hanos, Philipjiine

lls., October 1915, G. 1$. Vi;tdo: 1 <J New Gninea, Pemberton col.;

I á Malcknia, Malna l$ay, New llebrides May 1929, L. F.

íiheesman; 25 $ 13 9 MARl.VNAS IFS., Guam: Pt. Oca, Agana

May Jnne and Jnly 1915, G. F. l$oh;irt and J. F. Gressit; Rota:

22/0/52 V. Koiido; GAROFINF IFS., Va/) Grouj): Yap Is., Oct .

1952, N. F. II. Kranss; Ilill behind ^'aptown, 29/11/52, J. F.

Giessiti, Rahdlliunf): IMiiming. 25/12/17 II. S. Dybas, 23/5/53 Oller;

yafid: Golonia, Jnly 19.50 R. |. (hiss; Koior: jnly 1953, j. \V.

Beaiilsley, Sandakan. l$oi iico, Pcmbei ton toF; Fly River, New

Gtiinea, Pemberton tol.

I bis species is near to julvus Knight bnt tlillcrs in the shoitcr

secoiul tintennal segment, less tnmid calli, color;ition, smaller sizo

;md in the strnctnre of the male genitalia, whilsl the two lattct

tharacters distingnish it Irom mc/mio/w Rcnter.

•10

lloi.. Mus. Goei.di — loino XI (I) — Setembro 1955

Usinger (19.^9) founcl this species associated with Peregrinus

maidis (Aslimeacl) on roni, the eggs of tlie íulgoricl appareiitly being

its preíened food. It was also common on rice where it preyed iipon

the eggs of Nilaparvatn liigeus (.Stal). Aceording to Usinger (1946)

lividipeiuiis Reuter was introduced into the Ilawaiian Islands

recemly in an effoit to eontrol the com leafhopper biit did not

become establisbed. Ziminennan (1918) says tluu the species was

introduced from Guam into Ilawaii by the Board of Agriculture

and Forestry, in 1939, and confirms the fact that it did not become

establisbed .

CYRTORUINUS MELANOPS REUTER

Cyrtorhinus melanops Reuter, Ofv. E. Vet. Soe. Eorh. 47 (2): 6,

1 905 .

* Cyrlorliiiius tiiegalops Poppius, Acta Soc. Sei. Ecnn. 44 (3): 71,

1914 (error pro melanops Reuter).

(Figs. 14 A-j)

Characteri/ed by the wide pale area adjacent to the eyes in the malc

and the genitalia.

Male: length 3.8 mm., width 1.2 mm.; head, width 0.73 mm.,

vertex 0.37 mm.; antennae, I 0.3 mm., II 1.18 mm., 111 1.0

mm., IV, broken; pronotum, length 0.45 mm., width 0.93 mm.;

rostium length 1.15 mm.

Type: ^'ellow-green apar from anterior of head, basal segmenl

of antennae (exte|)t extreme base and apex) black-brown, second

antennal .segment light brown, paler in the centre, hind tibia with

pale brown mark a third length from the apex, a|)ical segment ol

tarsus and ciaws ligth brown.

Other specimnis: Head black with two wide pale areas

adjacent to the eyes, stretching Irom just posterior to the base ol

lhe antennae to the posterior margin of the vertex; antennae black-

brown with the extreme base and apex of lhe basal segment pale;

l)ronotum and .scutelium entirely black-brown; hemelytra yellow-

green, claval suture slightly iniuscale, membrane pale, nervures, p;de

green; legs pale green, with extreme base of the tibia brown, the

hind tibia sometines enliiely pale brown, with the base darker, apex

ol tarsus and ciaws brown; umierside and abdômen brown-black

rosirum |)ale green, tip dark.

Pubesccnce pale and fine, macroplerous.

SciELO

10 11 12 13 14 15

Carvai.iio & SoirniwooD — Cyrtoihinus-Mecoinnia Coinj)lex

Getiitalia: accleagiis with a single vesical aijpendagc (fig. 14 }),

Iclt claspcr bifid, lowcr arni curved (figs. 14 F, G), riglu clasper

also bifid, its lower arni with inwardly curved teeth apically (figs.

14 H, I).

Female: length 3.88 min., width 1.21 nnn.; head, width 0.77 mni.,

vertex 0.38 mni.; antennae, segment I, length 0.32 mm., ff

1.1!) mm., III IV broken; pronotnm, length 0.5 mm., width

0.98 mm., rostrum length 1.17 mm.

Generally similar in coloration to male, biit pale areas adjacent

to eyes sometimes very small and claviis infuscate; abdômen pale

green with dorsinn brown or entirely fuscous.

Distributiori: “Caffrorca” [Natal], Abyssinia.

Spccimcns sludied: i ype, I $ , ‘‘Caffrorca” (Riksmnseum,

-Stockholm), 3 á I 9 Mawash River, \V. of Mount Zaquala (c. 6,000

ft.), Abyssinia, 28/11/26, J. Onier Coo|)er (15. M.); 1 á -Serpeilt

Lake, Wonramboidchi, (c. 9,000 ft.) Abyssinia, 5/10/26, [. Omer

Cooper (15. M.); 1 9 nr. Addis Aliem (c. 8,000 ft.), Abyssinia,

19/9/26, J. Omer Cooper (15. M.); I 9 Natal, 15rown col.

Although the other specimens are imich darker in colonration

than the lype, examination of lhe genitalia left no doidn that lhey

were the same spccies. On genitalia and other characters it is closest

to lividipennis Reuter, but is distinguished from ihis species (jn si/.e,

and from carieis (Fallcn) l)y the pale areas on its head, the pale

ajjcx of lhe basal antennal segment and genitalia.

CYliTüRIllNUS CAlilCIS (FALI.EN)

Capsus carieis Fallen, Mon. Cimic. .Succ.; 102, 1807.

carieis Vollenhoven, liem. llei. Neerl.: 228, j)!. 16, fig. 4,

1878.

Cyllecoris carieis llahn, Wan/, Ins. 2: 100, fig. 184, 1834.

Cyrtorhinus carieis Reuter, liem. Gymn. Fur. 3: 383, 555, pl. I,

fig. 10, jjl. 2, fig. 3, 1883: .Saunders, liem. llei. 15rit. Is.: 283,

pl. 26, fig. .5, 1892: Stichel, lllus. 15est. Deut. Wan/. 8: 227,

figs. 590, 591, 19.33: Wagner, 'Fit-rw. Deut. 41, 151indw.: 127,

1952.

• Capsus clcfrantulus Meyer-Dür, Ver/. .Schw. Rhync.: 86, pl. 5,

fig. 2, 1843 (syn. by Fhomson, ()j)usc. Fnt. 4: 437, 1871).

Sphyraccplialus clegautulus Douglas Ik .Siolt, I5rit. Mcm.: 351, 1865.

• Capsus cliloropicrus Umich-ScUiicUci, Ins. Ver/.: 34, 1853

(syn. by Reuter, liem. Gymn. Fur. 3: 383. 1883).

(Figs. 15A-II)

■12

Hoi,. Mus. CioKiDi — l oiiio XI (1) — Setembro 1955

Cliaracleri/.ecI Ijy the entirely Ijlack antcmiac aiul the gcnitalia.

Male: Icngth 'l.l mm., wickli 1.25 mm.; head, widtli 0.85 mm.,

vertex 0.35 mm.; antcnnae, segmciu I, Icnglh 0.'15 mm., II

1.50 mm., III 1.20 mm., IV'^ 0.53 mm.; pronotmn, Icngth

0.5 mm., widtli 1.0 mm.; rostnim Icngth 1.2 mm.

Head hlack, exce])t for tvvo triangidar arcas on the vertex

adjacent lo tlic eyes, pale green; antcnnae entirely hlack; pronotinn

and sditellnm hlack; hcmelytra pale green with clavns and median

arca of corinm snffiiscd with dark hrown, nervnrcs hrown; Icgs ])ale

green with apex ol tarsi dark; rostnim pale ti|) dark; nmlerside of

head and thorax hlack; ahdomcn pale green .

Pnhcsccnce pale and fine: all specimens seen macropterons.

C.enitalia: aedeagns (Fig. 15 II) of orthotyline type, with lhe

single vcsical ajiiiendagc cnrved at its apex; left clasiier (I''ig. 15 D,

K) with process simply cnrved; right clasper (Fig. F) sinijile

tnincatc slightly cnrved, Iceth of dorsal arca not raised on a process.

Feiijdlc: Icngth -1.0 mm., widtli I .-15 mm.; head,.width 0.85 mm.,

vertex 0.38 mm.; antcnnae, I 0.15 mm., II 1.22 mm., 111

1.0 mm., IV^ 0.53 mm.; pronotnm, Icngth 0.5 mm., width

1.0 mm.; rosiriim Icngth 1.1 mm.

Coloration and piihcscencc as in inale. Macropterons.

(icnitnlia: 'Flie K stnictiircs are small semiijiiadrate with the

inner posterior corner elongated (Fig. 15 G).

Disirihnlioii: Ireland, Scotlaiul, Wkaies, Fngland, Nclliei laiuls,

France, Swii/erland, llnngary, Denmark, Norway, Sweden, Finland.

Fhiropean Rnssia (iiid. Gamasns), Siheria, Fnrkestan.

Sperimens sludicd: I á Sannders coll.; 2 í I 9 1 loundslow

Ilcaih, .Middiesex, U. K. 20/7/53, G. F. Woodroffe, 2 9

Wimhledon (iommon, Siiriey, F.ngland, 8/10/51, 'F. R. F.

Soiithwood; 2 9 Aviemore, Scotlaiul, Angust 1938, A. M . Massee:

I (J 1’iesling, I.ower .\nstria: I 9 Colorado. Uhler col.; Several .J

and 9, Wrangcl, Alaska, 15. Malkin, \'ll. 951.

’I liis large sjiecies is similar in appearance to inrl(iii(>j)s Rcnier,

hiit its entirely hlack tintemiae, sinall pale arcas on the head and

gcnitalia distingiiish it. It is foiind at the hases of rnslies (/i/nrii.v)

and sedges {('.arrx, Sri}l)iis), as an adnlt hetween jnne and Octoher.

'lhe winter is passed in the egg State and tlie yonng nymphs liatih

tlic lollowing spring (hiitler 1923, Knllenherg 1910). l he eggs,

wliicli aie sansage-shaped with theit micropylat end strongly ciiived,

arc laid in the Icavcs or stems of vations Scirl>us spp. (Knllenherg

1913, 1910) (Fig. 22).

SciELO

10 11 12 13 14 15

•13

Carva i-iio &: Southwood — Cyrtorhimis-Mccoinnia Comjilcx

Most oí tlie American records for tliis spccies are to be relerred

to Tyttlius xmgus Kiiight {Cyrtorliinus carieis imgus Knight, 1923),

a species of 1’liylini. Several specimens labellcd as carieis in American

collections and seen by the sênior autlior do not ])ossess the

convergem arolia of tbe Orthotylini. C. carieis (Fallen) is apparently

rare in the United States where it seems to be restricted to tlie

Rocky Monntain range.

GENLJS MECOMMA FIE15ER

Mecomma Fielter, VVien. Ent. Monat. 2: 313, 1808; Reiiter, llerm.

Gymn. Eiir. 3: 383, õlã, 1883; Ilneber, Syn. lilidw. 2: Kib,

1-13, 1908; Fojjpiiis, Acia Soe. Sei. Feim. 11 (3): (iO, 72, 1911;

Wagner, Fierw. Dent. 1, lllindvv.: 110, 129, 1952.

* Spltyracej)halus Douglas & Scott, Ibit. Hem.: 318, 1805.

* Sphyrops Douglas & , Scott, Ent. Mon. Mag. 3: 10, 1800 (nom.

nov. for Spliyraccplialiis Douglas 8: Scott, 1805).

* Antiphihts Distam, Ann. Mag. Nat. Ilist. (8) 1: 521, 1909 (syn.

by Uarvalbo, An. ,\cad. brasil. Ci., 21 (1); 78, 1952).

* Aristobiiliis Distam, Ann, Mag. Nat. Ilist. (8) 5: 10, 1910

(n. .syn.).

* Nyclicapsus Pojtpius, Acta Soc. Sei. Fenn. ll (3); 71, 1911

(n. syn.).

* Aristoholiis CarviiUxo, \n. Acad. brasil. Ci. 21 (1): 79 (error pro

Aristoholus Distam).

'1’ype species: Capsus ambulans Fallen, 1807 — monobasic.

Males ahvays fnlly winged medinnt si/ed bngs (1-5 mm,), females

nsnally brat liypierons (2.0-3. 5 mm.), occttsionally fnlly uingend. lleail

ronndde anteriorly; lace vertical, deplli from base oí eyes to a|)ex of

tyliis aboni hall lhe vertical iliameter of tbe eyes, antennae arising

aiijacent to lower median conter (tf the eyes, basal and second anten-

nal segments maikedly thitker ihan ihiiil and fomih; pronotnm

camptimilifoiin wilh nairow anteiior collar and in the tttale strottgly

t;iised posteriorly; itt llte tttales heald widih iitdiidiitg e)e.s ttearly

twice :ts tvide tis protiotal collar attd .snbe(|nal to the wiiíth of the

b;tse of the ptottolimt; o])ettiitg of otloriferoiis s;ic tttised; |)reiarsits

wilh flaplike arolia: tostrnttt reachittg beyottd base of hittd coxa;

tttale pygophore wilh stibgeniial plale or aedeagal snpport oitly

slighlly projet iiiig: Ifll claspei sirongly cm ved, l iglit < lasper wilh

a ridge ol leelh, aedeagtis wilh single vesital appendage and vesica

ctirved tioisally jnsl before goiiopore; bmsa (opnlairix of fenialc

with loiindetl to conical K stnicttires; coveied wilh siniple .senii-crect

I5oi.. Miis. Goei.di — loino XI (1) — Setembro 1955

jjubescence (0. 10-0. 18 mm.), usually loiigcr in tlie malc; general

coloiir of males light brown and black and in íemalcs brown and

black or almost entircly brown.

Mecomma is closest to Cyrlorhinus in general faseies and in tlic

strnctnrc of the genitalia of botb sexes. It differs liosvcver in thc

following cbaractcrs: the female is nsnally braebyptcrons, the outlinc

of the male is almost parallel sided, the rostrnm reaclies beyond thc

base of thc hind coxa, thc pidje.scencc is long (nsnally over 0.15 mm.),

thc enneus is long (in thc malc twicc as long as wide), the left

claspcr is strongly curved and nnbranched, thc right claspcr has an

apical row of spines and a sjrinelcss vcntral process, thc vcsica is

cnrvcd n])wards just before the gonoporc and thc K strnctnrc of

the bursa copidatrix has its posterior margin jrrodnccd almost to

a point.

.Spccics of Mecomma ocenr amongst grasses (gcnerally damp)

in or aronnd temperate forests. Ilencc in tropical regions (c.g.

Índia) Mecomma sp. arc |)rescnt only in mountainons districts,

wherc these conditions arc foiind; this rcstdts in gcographical isola-

tion, followcd by sid)s|K‘ciation . Snbspcciation has also orenred

in the Ncarctic region. It seems that thc isolating effcct of the

gcographical barriers arc strcngtbcned by thc brachyptcrism common

in thc fcmalc. One spccies is |K)lymorphic.

l he striking convcrgencc betwcen thc allied Orthotylinc genns

Cyrtorhinus and the Phylínc genns Tyttlius has alrcaily bcen noted;

a similar convcrgencc in both sexes occiirs bctwecn Mecomma and

the phylinc genns Orlhoiiotas which in habits similar sitnations.

A further stndy of Mecomma with more material aiul ccological

data woidd bc of great interest, intrinsically and from thc more

general aspect of spcciation.

ki;y ro tiiil si*i;cii:,s of mecomma fifiifr

Eemales: (at ])rescnt known)

1 . .Sccond antennal segment distinctiy clavatc, clytra reaching only

lo sccond abdominal segment mimelica n. sp.

.Sccond aniennal segment if incrassale towards apex, nol clavatc,

clytra reaching posterior margin of third abdominal segment

or beyond 2

2. Flytra entircly black 3

Flytra pale oi il infuscatc, ycllowish at base or margins . . 7

.3. 1'ronolnm noticcably broader posteriorly than anteriorly (Figs.

22 B. A), segment two of antcnnae enlirely black •!

1’ronolnni only slighlly broader posteriorly than anteriorly

SciELO

10 11 12 13 14 15

Carvalho S: Souihwood — Cyitorhiiius-Mecoinma Complex 45

(Figs. 17 A, B; 20 A) scgment two of antennae at least

partially palc 6

4. Antennae entirely black aminis (Distant)

Antennae partially pale 5

5. Basal segnient of antenna pale cfiijtoisis Renter

Basal segnient of antennae black, the third partially pale . .

ainbiilans (Fallen)

6. Secoiul antennal segnient dark at apex, npper surface of

pronotinn and hemelytra shining

orientalis orieutalis n. sp. forni a

Second antennal segnient entirely yellow, npper snrlace of

pronotnni and lienielytra rngose i

orientalis orientalis n. sp. form b

7. Antennae entirely black, larger species grandis n. sp.

Antennae partially palc or brown 8

8. First antennal segnient black or fnlvons 9

First antennal segnient jiale 10

10. Antennae nsnally castancons to fnlvons; K strnctnre as in fig.

20 I antennata Van Dnzee

Antennae black; K strnctnre as in fig. 20 F .... gilvipes (Stal)

11. Pronotnni and scntcllnni nnicolorons .. luetnosa (1’rovanclier)

Pronoinni witii a longitiulinal faseia and apex of scntcllnni

pale 7nelanoceplialus (Poppins)

Males', (known at iircscnt)

1 .

. 8 .

(■).

Pronotnni and first antennal segnient castancons to fnlvons

antennata Van Dnzee

Pronoinnt black, first antennal .segnient black or palc 2

riiird antennal segnient partially jialc 3

'Fliird antennal segnient black 4

Basal antennal segnient palc, cnnens long (0.70-0.81 nini), licad

nairow (0.r)8-0.71 nnn) orientalis n. sp. orientalis for a

Basal antennal .segnient dark or palc, cnnens sliort (0.()0().(i2

nnn). Iiead ivide (0.73-0.78 nnn)

orientalis liynialaycnsis n. snbsp.

Antctinac black

Antennae ivitli first segnient pale 8

Aiiices of corinni aiul cnnens pale to ocliraccons b

Ápices of corinni atui cnnens snffnsed witii black /

F.ndo,.,iinni black nnulagascariensis Renter

Fndocorinni traiislncent. palc ..luetnosa luetnosa (Provanclicr)

•lü

Mus. Goki.di — Tomo XI (I) — Setembro 11)55

7. Larger, liead width 0.83-0.87, riglit claspcr as in lig. 1*1 E

spicidum as iii íig. 21 E nmbulans (Fallcn)

Smaller, hcad widtli 0.79-0.83 mm, riglit claspcr as iii fig. 19 li,

C spiculmn as in íig. 19 1) (unicus (Distant)

8. Genitalia as in íigs. 17 G, F orientalis oricnlnlis form b

Genitalia as in íigs. 21 A, I .... luctuosa pacifica n. subsp.

MECOMMA ORIENTALIS n. sp.

(Figs. 17 A-H)

Cliaractcri/cd by tlie pronotmn of tlie femalc only sliglitly wider

postcriorly tlian antciiorly, tlie partially palc coloured antennac, thc

general jrroportions and genitalia.

[Note: forms a and b of tbe femalc may not correspond with

tlic rcspectivc forms in tbe inale and hcnce an allotype bas not

bccn designated].

Subspccies orientalis nov. sp. Form a (typc form)

Male, Ilololype: Icngth -1.25 mm., width 1.25 mm.; hcad width

0.08 mm., vertex 0.32 mm., antennal .segment I, Icngth 0.38

mm., II 1.48 mm., III I.I2 mm., IV 0.40 mm.; pronotum

Icngth 0.47 mm., wiilth 0.92 mm.; rostrum Icngth 1.39 mm.;

cimeus Icngth 0.80 mm.

Ilcad black, antennac with thc first segment ycllow brown, thc

sccond black, the third palc ycllow with its apical half black-brown,

thc foiirlh black-brown; pronotmn and scntcllnm black; hcmclytra

light brown with clavns, iimcr maigin and angle of (orinm darker,

onter anglc of (orinm and apex of ennens dark brown; Icgs ycllow-

red, with base of roxa black, and ajiex of tarsi darker; abdômen and

niulcrsidc entirely black.

Fnbcscencc of fine scmi-ercct hairs. Macropterons.

Genitalia: aedeagns of orthotyline typc with a single vesical

aiipendage (Fig. 17 F), this is not bifid at the apex; right claspcr

with vcntral jiroiess cnived postcriorly (Figs. 17 G, K); left claspcr

strongly (iirvcd (Fig. 17 D, I) slightly more so than in ainiens and

with dorsal cornei of bas;il region slightly prodiufd.

Otlicr spcciincns: lenglh 1.0 1.4 mm.; width 1. 201. 29 mm.

(for other measnrements sec Tablc I).

Goloration only diflering in that of third antennal segment

which may be palc with only extreme b.ise and a|)ex daik or varia-

tions between this and the (ondition in lhe I lolotype, bnt aiways at

Icast hall palc.

SciELO

10 11 12 13

C^ARVAi.iio is: Soiniiwooi) — Cyrloi Iiiiius-Mccomma Coniplcx

•17

Specitncm stiidied: 1 <} (Holotypc) South Iiulia, V.

Oaiiiphcll (Hiitish Muscuin); 17 <5 (1’aiatypcs) Soutli liulia, T. V.

Caiiipbcll (Ilritish Muscuni): 3 á Kodai-Kaiial, S. Iiitlia, '1’. V.

Campbell; 1 $ Chikballapur, Mysorc, Jan. 1915, T. V. Campbell.

Fomí b

Male: leiigth 1.05 mm., widtli 1.15 mm.; head widtb 0.72 mm.,

vertex 0.31 mm.; anteiinae, segment 1, lengtb 0.37 mm., H

1.23 mm., III 0.90 mm., IV 0.18 mm.; pronotum, leiigth

0.50 mm., widtb 0.85 mm., rostrum lengtb 1.33 mm.; cuneus

lengtb 0.01 mm.

Similar to foim a in general coloration, but differing in baving

tbe tbird antennal segment entircly black and ibc ápices oí the

corium and cuneus oltcn paler, :dso in tbe sborter cuneus and

antcnnae and tbe narrowcr base ol tbe pronotum. Ibe genitalia

are virtually ideniical witb tbose of lorin a.

Specimens sludicd: 3 <} .South índia, V. Campbell; 1 $

Kodai-Kanal, S. Índia (7 000 It.) 21/3/30; 7 á Nilgiri Mills., S.

Índia, r. V. Camjjbell.

Fonn n, l'emale: lengtb 2.10 mm., widtb 1.1 mm.; bead witlth

0.70 mm., vertex 0.33 mm.; antennae, segment 1, lengtb 0.31

mm., II 1.20 mm., 111 0.83 mm., IV 0.33 mm.; pronotum,

lengtb 0.12 mm., wititb 0.71 mm.; rostiinu lengtb 1.12 mm.;

bemelytra witltb 0.72 mm.

Ileail black witb two arcas ailjacent to tbe eyes sligbtly paler;

antennae, basal segment tlark or jtale, second segment witb basal

two tbirds yellow, apex black, tbird segment basal tbird pale, apex

and fourtb segment tlark brown; pronotum and scutcllum black;

bemelytra black witb extreme bumeral angle sligbtly jtaler; legs

yellow— lidvous witb apical segment ol tarsus tlark antl ctixa, at

íeast tbeir base black; abtiomen bbick witb extteme margin ol

connexivum pale; rtistrum fulvttus, its basal antl apical segments

tlark brown t>r blatk.

Mpper surface smootb antl sbining, especially on bemelytra

wbete tbe only maikings are tbose t)l tbe alveoli, lrt)ni wbicb ari.se

pale semifiett bairs ol metlium lengtb. Ilratbypterous.

(ieiiildlifi: K strutture ol bursa topulatrix as in Fig. 17 II.

Spdriíneiis sliidied: 2 Ç l.oveilale, S. Intlia, 1 . \'. (.ani|)ell.

2 9 Nilgiri Mills, S. Intlia, T. Campbell,

Farm h. Fnmilr. lengtb 2.18 mm.; widtb 1.38 min.; beatl widtb

0.75 mm., vertex 0.32 mm., antennae, segment 1, lengtb 0.29

SciELO i'o

Carvalho & .Southwoo» — Cyrtorhinus-Meconima Conijjlex

female: unknown.

Speciinens sludied: 1 $ (Holotyjse), Kiirseong, E. Himalayas

(5,000 ft.), 7/7/08 (IJrit. Mus.); 2 á Gaiigtok, Sikkim, (6,000 ft.),

29/4/28, F. M. liailey.

Distribiilion of M . orientnlis Jiimalayensis: Eastern Himalayas

(Sikkim and N. Bcngal).

Notes on M . orientalis: this species is sharply distinguished

from M. aniictis by thc ionn ot' the vcsical ajjpcndagc of the male

genitalia, in Aí. orientnlis thc apcx of this stiucturc is single,

rescmhling Cyrtorliinus in this respcct. Aí. orientalis difíers forin

Aí. aniicns in coloration and pro])ortions, and can, with expericnce,

he distinguished Avith the naked cye hy the slightly nanower and

less convcx form of the anterior pronotuin, the anterior collar heing

well marked. lhe two suhspccics orientalis and himalayensis are

clearly distinguished hy the form of the right genital claspcr and

l)y thc slightly diffcrcnt proportions, especially thc widcr hcad and

short cuncus of himalayensis . l he two suhspecies whosc hahitats

are montaine grassland associated with damp, hut temperate

woodlands, are .separated from one anothcr hy the Deccan plateau

and the Canges valley.

Forms a and h of Aí. o. orientalis are distinguished in the male

hy thc partially pale third antennal segment, the longcr cuncus

and antennae and widcr pronotum ol form a and in thc samc form

of femalc hy the partially dark sccoiul antennal .segment and the

shining hcmclytra. 1'hcre are no differcnccs of any magnitude in

thc male genitalia and those in the femalc are only slight. It will

he scen from Figs. 23-25 that there is some overlap hetween the

range of variation of these species even with thc most diagnostic

measurements. l he majority of lhe material availahle was collected

hy Mr. 1'. V. Camjjhell; vvho, it is understood, mounted his captures,

generally on thc .same day as capture. As forms a and h (of hoth

•sexes), wcre often mounted on lhe same card, suggcsting that they

werc collected togcther, it scems uniikely that they could Í)c diffcrent

scasonal lorms of thc same í|)ecies or different ecotypes, although

the latter jrossihility cannoi he eliminated. Until further material

atui ecological informalion is availahle Aí. o. orientalis should he

regarded as polymorphic.

MECOMMA AMÍCUS (DhSl AN 1)

Antipliiliis amiens l^isVãul, Anu. Mag. Nat. llist. (8) 4: 521, 1909;

Distant, Fauna. Hrit. Ind. Rhynt. 5: 272, fig. 117, 1910.

— 4 _

SciELO

10 11 12 13 14 15 16

50 Hol. Mus. Goki.di — 'loiiio XI (I) — Setembro 1955

Mecomrnn arnicus Carvallio, An. Acad. brasil. Ci . 2-1 (I): 78, 1952.

* Aristobiilus filitis Distant, Ann. Mag. Nat. Hist. (8) 5: 17, 1910

n. syn.) Distant, Faun. Brit. Incl. Rliync. 5: 28(5, fig. 157,

1910.

* Cyrtorhinus filius Carvalho, An. Acatl. Brasil. Ci. 24 (1): 79,

1952.

(Figs. 19 A-G)

Characterized by its entirely black antennae, size and genitalia.

Male: length 4. 0-4. 5 nnn., width 1.20-1.28 mm.; head, width

0.80 mm., vertex 0.38 mm.; antennae, segment I, length

0.45 mm., II 1 ,35 mm., III 1 . 15 mm., IV 0.43 mm.; pronotnm,

length 0.50 mm., width 0.95 mm.; rostnim length 1.33 mm.;

ciineus length 0.72mm.

Head black, soinetimes two slightly paler arcas adjacent to the

cyes; antennae black; pronotnm and scntclhim black; hcmelytra

light brown with clavtis, inner and oiiter angles oí corium and apex

of ciineiis tlark brown; mendjranc light grey, ncrviires brown; legs

yellow-rcd, ápices of tarsi slightly d.irker; lindersidc and whole ot

abdômen black.

Pubcsccncc of fine semi-erect hairs. Macropterons.

Genitalia: aedeagns of orthotyline type, the vesical appendage

with dorsal projection and apically bifid (Fig, 19 D), leít clasircr

strongly curved (Fig. 19 E). 'Fhc right clasper with the ventral

process curved antcriorly (Figs. 19 B, C).

I'euuile,brachyf)ter()us: length 2. 1-2.4 mm., width 1.4 mm.; head,

width 0.83 mm., vertex 0.38 mm.; antennae, I 0.30., II 0.87

mm., III 0.74 mm., IV 0.37 mm.; pronotnm, length 0.41 nnn.,

width 0.80 mm.; rostnim length l.I mm.

Entirely black apart from two sm.dl areas median and adjacent

lo the cyes which are pale yellow-green and the legs which arc

tcstaccons, with only the base of the coxa black.

1’nbesccnce of fine pale adpressed hairs, some longcr ones on

the anterior of the thorax, ihose on the legs dark. Brachypterons,

hemclytra reaching to posterior of segment six, membrane absem

and cnneal sntnre absent.

Genitalia: K strnctnre (Fig. 19 F) with an clongated enrve

jirocess .

Macropterons: length 3.75 mm., withh 1.35 mm.; hcati, witlth

0.90 mm., vertex 0.30 mm.; proiumiin, length 0.53 mm., width

Carvalho & Southwood — Cyrtorhimis-Mcconinia Coniplcx

1 .10 mm.; otheivvi.se as in brachypterous form. Hemelytra dark

brovvn to black, membrane tliisky vvitb nerviires tlarker, otber-

wise as in brachypterous form.

Distributioiv. Sikkim, N. llengal, and Manipur (Assam).

Sspecirnens sludied: 1 $ (Holotype of mniciis) “Darjiling

(6,000 ft.), 25/9/08, Brunetti (Hrit. Mus.); 1 $ (Holotype oí

filius), “Darjiling” (6,000 ft.), 23/9/08, lírunetti (llrit. Mus.); 2

$ “Darjiling” (6,000 ft. — sweeping grass and low berbage), 25/9/08,

Brunetti; 2á Ukbral, Manipur (6,400 ft.), Aug. 1908, Pettigrew;

1 $ Kurscong, E. Himalayas (5,000 ft.), 1 jl 23 9 Darjceling

(7,000 ft.) 11-20/3/24, R. W. G. llingston; 9^69 (2 macropt.),

Gangtok, Sikkim (6,000 ft.), 24 April— 2 May 1928, E. M. llailey.

An examination of tbe types of A7itipltililus amicus Distant and

Aristobulus filius Distant for tbeir coloration and size, in relation

to otber Indian specimens, shows tbat ibese are tbe male and female

of tbe samc specics. 'Ebis conclusion is supportcd by tbe similarity

in tbe locality data. AI. nrnicus is distinguisbcd by tbe entircly

black antcnnae in botb sexes and tbe genitalia; tbe bifid apcx of

tbe vcsical appendage and tbe longer basal antennal segment clearly

sejiarates it from Áf. orieulalis. It is dosest to Al. auibulans,

unfortunately bovvevcr most vvork on tbis latter spccies bad to be

done on material from England; it is bigbly desirable tbat larger

samples from otber jxirts of its range sbould lie compared vvitb Al.

amicus, for even vvitbin tbe Englisb material one specimen (out ot

several) of AI. ambulnns from Ilarjx.'mlen bad mcasurements just

vvitbin tbe range of variation of Al. amicus (Eigs. 23, 24).

Measurements of .several specimens of eacb species are given in

'1'able 1 aiul 11 and tbe diagnostic diaratters |)lotted on Eigs. 23, 25;

it vvill be sccn tbat Al. amicus differs from Al. auibulans in being

smaller in botb sexes. It also tliffers in baving tbe antcnnae entircly

black in tbe female, vvbilst tbey are pariially jjale in tbis scx of Al.

ambulnns and tbe male genitalia of tbe two specics, altbougb dose

are dislinct.

llitbcrio Al. amicus only .bas been recoitled from Sikkim, N.

Bengal and Manipur, Assam, but it pioliably occurs ibrougbout tbe

Eastern llimalayau and Assam regious beivveen 5,000 and 7,000 ft.

vvbcre tbe cold tcmperate forest occurs.

AIECOAIAIA CIUNKNSIS (REU TER)

Alecommn chiuctisis Reuter, Annu. Mus. Zool. Acad. St. 1’etcrsb.

10: 63, 1905.

Cbaracleri/ed by its black coloration,. tbe loug antcnnae vvitb tbe

basal segment jialc.

]5()i,. Mus. (;()i;i,i)i — 'loino XI (I) — Scteinljio 1955

Male: unknown.

Fcmale: Rciiter’s dcscrij)tioii: —

Fcmiiiíi late ovata, nigia, (apite al)(loiiiine(|iie iiitidissiimis

hoc subaenco-niicante, ])ronoto, scutello liemielytris(|iic o])aculis cuni

dorso abdoniinis longe cinereo-pultescentibus; verticc iitrinuiiie

guttula minuta ol)soleta testacea; rostro, artieido primo nigricante

cxcepto, antennis arlicido |)rimo tolo seciindotjne basi, nec non

pedibiis llaventibus; antennis articulo setnndo margine basali pronoti

(formae bracbypterae) saltem dimidio longiore versus apicem in

ciavam elongatiim incrassato, tertio secundo fere breviore, dimidio

basali albo. Long 9 ^ mm.

.Ad llinuen Scimbagu d. 8 anguste 189,3, uniemn spccimcn.

j\í . ainlnilinili (Fali.) simillima, antennis longioribns, aliter

ronstrnclis, articulo jtrimo flavo-lestaceo divergens. Ciapiit (9

bracby|)t.) basi |)ronoti ac<pie lalimi, valae nitens, ab antico visnm

latitiidine verticis ociilitpie linici aetpie longnm, vertice ociilo circiter

duplo lutiore. Rosirnm coxas postiças snbattingens, apicc nigro.

Antcnnae ( 9 ) corjxjiis longitudine, articulo secutulo .sat longe

tidpressim piloso, primo lere .3-'- longiore, a medio lortuis incrassato,

ultinuts scmicrecte pilosis. Pronotum (9 bracbypt.) basi longitudini

lere du])lo latins apice (piam basi vix tnagis (piam — augustius disco

bori/ontali, anguli.santitts rotundatis, latcribus mox atUc ângulos

postiços leviter situiatis, tnaigine basali late situiato. Iletnielytra tota

nigia, médium abdomiuis |)aull() supeiantia, (lavo et cuneo batid

discielis, margine apicali latissime lottmdato, membratia attgttstissi-

ma. 1 ibi.ie tenuiler concoloriter spinulosae. Farsi articujo ultimo

apice lusco.

Distriliiition: Sdmbagu, Cliitia (otily knowti Irom tbc typc

local ity) .

•Specimetis of titis species were iiot available for study, bttt ít

woiild appeat lo be ( losesi lo atnhuldii.s lútil. on si/e, differing in

tlic antennae svliidi are lotigct willi llie basal .segment palc.

MECOMMA AMIiül.ANS (FAI.I.FN)

C(//ui/.ç tnnhtilatis Fallen, Moit. Citnic ..Sttec.: 101, 1807: Ileiiidi

.Sdtaefrei, Watt/. Ins. .3: 109, figs. .3.35;$,37, I8.3(i; Reuier, liem.

(ivmn. Kui . .3: .381, 555 ,pl . 2, ligs. 1-2, 1883: .Saunders, Ilem.

Ilel. lirit. Is.: 280, pl . 20. lig. .31. 1892: .Siidiel. llltts. Ilest.

Detil. Watt/. 8 : 220, ligs. 588-89, 19.33; Wagner, 1 ierw. Deut.

II, lilindw.: 129, lig. 19:72.

Carvaí.ho ÍL- Soirniwoon — Cyrtorhimis-Mccoiinna Complcx

* Capsus dubitis Zctlcrstcíh, Iiis. Lajij).: 279, 1810 (syn. by l’liom-

soii, Opuse. Ent. -1: 437, 1871).

* Capsxis ochripes Curtis, llrit. Ent. 15, jtl . 093, 1838 (syu. by

Douglas &; Scott, lirit. liem.: 319, 1805).

* Capsus nigritiiliis Zettcrstetlt, lus. Eapp.: 279, 1810 (syn. by

'Ihonison, Ojjusc. Ent. 4: 437, 1871).

(Fig. 22 11)

Cliaractcri/cd by its size, pr()|)ortious, geuitalia and thc ícnialc witli

cntircly black bemclytra, but paitially pale antennac.

Male: lengtli 4. 4-4.0 mm., width 1.38-1.42 mm.; licad, width

0.80 mm., veitcx 0.30 mm.; antennac, segment 1, leugtb 0.40

mm., 11 1.50 mm., 111 -.38 mm., IV 0.52 mm.; inonotum

lengtli 0.50 mm., widtli 1.07 mm.; lostrum lengtli 1.30 mm.;

cuneus lengtli 0.90 mm.

llead black with tvvo ])alc arcas adjarent to thc cyes; antennac

cntircly dark; jrronotum and sctitcllum black: bemclytra liglit

oebrous brown with thc clavus, apices of corium and cunctis, and

inner anglc ol corium sulltiscd with black, and thc remaining

margins ol thc cuncus and outer margin ol thc corium a dceper

ycllow-brown; membrane pale, nervures margined with black: Icgs

ycllow-brown, thc apex ol thc tarsi daikcr and base of roxa black;

rostrum ycllow-brown, its tip darker; ‘ sides and venter ol thorax

black; abdômen cntircly black.

1’ubcsccncc ol linc scmi-crccl hairs of medium Iciigth (0. 15 mm.)

their alveoli being wcll maikcd on tlic hcmclyira. Macropterous.

('.niilalio: aedeagus (Fig. 21 F) ol orlholylinc typc with bilid

tootli vcsical appendage, k ll t laspcr stiongly (urvcd (higs. 20 C, D),

light (laspcr (Fig. 21 E) with ;m tipital toothcd i idgc aml a thin

median projettion. .Singlit lh uthis (1925) ligurc ol thc aedeagus ol

this spccics is (pnte iiKorrctt.

Icmale: l{nirliyptc'r<>iis: leugtb 2. 4-3.0 mm., width 1.58-1.70 mm.;

autcnnac, 1 0.48 mm., II 1 .25 mm., III I . 12 mm., IV 0.53 mm.;

pronotum, Icngth 0.5 mm., width 0.98 mm.; rostrum Icngth

1 .30 mm .

llead bhick, two slightly palcr arcas adjatent to thc cyes, anten-

uac, lirst and setond scgmciits bl.u k, thc third pale loi basal thiid

ol its Iciigts, thc remaiuder ol thc third and thc louith ■■egment

biown; pronotum, st titcllum and hcim4ytra black; Icgs light ycllow-

Itiown with b;iscs and a|)iccs ol tarsi daikcr; lostrum ycllow biowit,

hs apex daik; cntiic undcisidc black, extieme maigins ol connexi-

vum biown.

SciELO

10 11 12 13 14 15

15oi,. Mus. Cioi.i.Di — 'Toino XI (I) — Setembro 1955

Piibcsceiice of scmi-ercct Iiairs of inetiiiiin length (0.13 mm.),

tlicy are particailarly deiise on tlie second antcnnal segmeut and

paler and sijarser on ilie lliird and íoiirtli. lírachypterons, Iicmelytra

rearliing posterior oí tliird abdominal segment; membrane ;md cnneal

stitnres lost .

Gcnilalia: thc K strncttirc (lig- 20 I) is tapered and cnrved.

Macroplerous: length 4. 25 mm.; vvidth 1.50 mm.; pronotum,

length 5.00 mm., width 1.20 mm.

Otherwisc measitrements, colonration and pubesccnce as in

hrachypterons female, with membrane dusky, thc nervurcs black-

grey ontlined with black.

Dislribulioiv. Entope, Caticasns, Algeria, N. Asia, Alaska attd

Canada .

Specimcns sludicd: 1 5 New lorest, Ilants., U. K.

'E. R. E. Sotithwood; 1 $ E. Peckham, Kcnt, U. K., 10/8/48,

E. R. E. Stjiithwood; 5 á 2 9 I larpeiuleti, Ileits., U. K.

14-25/8/53, E. R. E. Sonthwood; 5 S 0 9 (1 macro]). ) S. Lakc

District, U. K., f. E. Satchell & 'E. R. E. Sonthwood n-'ÒOjllM;

I 9 .Mofiat, Scotland, U. K. Eryercol.; 3 5 9 Camhridgeshirc,

U. K. Fryer col.; 1 3 , I 9 Ihadore bay, Qncbec.

Elte ahnost entiiely black female, thc black antennac, its si/e

and genitalia distingnish this spetics Irom amicus Dislatit thc closcst

Old \Vorld species and from gihnpcs Stal and lucluosus 1’rovanchet

in thc New World. Ktiight says that all lecords oí amhnlans for

Anerica are incorrect and that males of this species and gilvipcs can

be separated only by the strnctnre of the genitalia.

Ehis assertion was correcled later in a pajter by Walley (Can.

Ent. 01; 152, 1932) in which a series of 37 males and 21 fetnales

from Ihadore Ihiy were foniul by Knight to be (iiubulans (Fallen)

alter (omparison with Enropean .sj)ecimetis. A malc atid female ol

this sj)ecies were stndied by lhe sênior anthor at the Califoiiiia

Academy of Sciences.

Ehe egg of this species, descrihed by Ktdleid)erg (1913) is laid

in lhe stems of v.irions Jiinots s|)p. and grasses (Knllenberg, 1910)

(Eig. 22) dnring late snmmer ()nly and Angnst) and hatch the

following spring, mostiy in May. Ehc nymphs, which are giecnish-

red, matine from late jnne onwards (Ifntlcr, 1923, .Sonthwood

nnpnhlished) .

M/:c:OMALI MADAGASCAHIENSIS REU EER 1892

Mrcotiirtia madagasairirnsis Renler, Ent, mon. Mag.

1’oppins, Acl;i Soc . Sei. Eenn. 41 (3): 75, 1914.

28: 185, 1892;

SciELO

10 11 12 13 14 15

Carvai.iio & SouTuwooi) — Cyrtorhimis-Mcconuna Comjilcx 55

Male: Rcutcr’s dcscription: —

Elongatus, nigcr, niticliis, longius palliclo-j)iibescens: gutta

iitrinquc vcrticis rostro j)eclibii.s(pic llavo-ochiaccis; hcniielytris divi-

dis, clavo conimissura apicccjiie late, corio intravenam cubitalem,

angulo anteriorc, cunci niembranatpie ciiin venis brachiali et cubitali

nigricantibus, corio extra venain cubitalem, cuneo, angulo interno

exce]Jto, areola meinbranae minore cum vena connectante limbocjue

laterali externo pallidis, angulis ajjicalibus corii exteriore et cunei

concoloribus. Long. 1 — mm.

Specimens of tbis species were not available for study.

MECO MM A GRANDIS nov. sp.

Cliaracterized by its size, entircly black antennae and partially palc

hemelytra .

(Fig. 20 A)

Male: unknown.

Eemale: length 3.25 mm., width 1 .08 mm.; head, widtb 1 .00 mm.,

vertex 0.18 mm.; antennae, segment 1, length 0.'15 mm.; II

l.lí) mm., III 1.01 mm., IV 0.48 mm.; pronotum length

0.55 mm., width 1.05 mm.; rostrum length 1.50 mm.;

hemelytra rvidth 0.87 mm.

Head black, wilh two slightly paler arcas on the vertex adjaccnt

lo lhe eyes aniennae entircly black; pronotnm and scutelhnn black;

hemelytra black wilh a lateral and apical band of about j thcir

withh yellow-testaceons; legs tcstaccons with extreme ba,scs of coxac

mui a])iccs of tarsi darker; abdômen black with margins of connexi-

vum retlish; underside black; rostrum yellow-testaccous with basal

•'uul aj)ical .segments fuscous.

Clovered wilh pale short adpressed hairs and longcr line crect

hairs; hemelytra anil pronotum strongly rugose; brachypterous, but

trace of membrane remaining.

Oislrihutiou: l)jem-I)jem Forest, 15 miles west of Abbdis

Ababa, Flhiopia.

Spfcimnu studied: I 9 (Holotype), Fdge of Djem-Djcm

Forest (c. 0,000 ft.), Abyssinia, 1/10/20, II. Scott (Hrit. Mus.).

I his species is distinguished from M . ainhulaiis and M . amicus

liy its partially yellow coloured hemelytra, rcsembling in ihis lespect

lurtuDsus nm\ M . gilxiipcs, and from lhesc species and M. t/iela-

by its large size.

Sf) Hoi,. Mus. (;oi, 1 . 1)1

Toiuo XI (I) — Sctcinhro

lhe only known spccimcn of this bug was rolleeted by Dr .

lingb Scott at tbc cdgc of tbc Djein-Djem (or Jeni-Jcni) Forest in

Oetober. D. Scott bas kinclly given us tlic loliowing inloniiation

about tbc arca: tlic forest itsclf is coniferous witb giaiit Jmiipers,

()() ft. or more in bcigbt and giant Podocarpus . TIic forest is

reeking wet during tbe Great Rains wbicb end in September or

later, and probably rcniains dani]) for inost of tbc year; tbe general

cliniatc is c.ool tcmperatc. Froni liis Journal Dr. Scott finds tbat on

October ■Itb, l‘J2(), bc collected imicli material from tbc grassiand

and isolated cliimps of troes at tbc edge of tbe forest. An accoimt

of tbis arca is given by Sxott (1950).

Ibus tlic habitat of Aí. gratidis agrees rioscly witb tbat of

otber members of tlie genus, viz. damp grassiands in or around

tbc margins of cold tcmperatc forests.

MEC.OMMA ANTENNATA (VAN DIIZFF)

Mecomtnn aiitetninla Van Dtizee, Proc. Cal. Acad. Sei. 7: 275 ,1917.

Cliaracterized by its coloin aiul genitalia.

Male: Icngtb •1.5 mm., widtb 1..S mm.; bead, Icnglb 0.2 mm.,

widtb ().8 mm., vertex ().,'{5 mm.; antennac, segment I, lengtb

0.3 mm., II 1.5 mm., III 1.1 mm., IV 0.3 mm.; pronotmn,

lengtli 0.5 mm., widtli at base 1.0 mm.; rostrtmi, lengtb

1 .1 mm .

Colotir dark brown to castaneous (antennac, bead, pronotmn

and scutellmn); two last antennal segments darker (base of tliird

segment sliglitly jialer); two faint spots on vertex ncar eyes, liemelytra

(except inftiscate arca of claviis aiul coriínn along commisstire),

Icgs and rostnnn |)ale yellow transbiceiu to ocbraceons; apex of

ctinetis and membrane infmnatc, tbe first and veins tlarker; nnderside

of body castaneous.

Càineus twice as long as wide at base, vertex witii a row of

bristies.

(icnildlitr. aedeagus witb a spidilum (big. 21 II) less dirved

apicalbj as in gilviptw. l.clt dasper very similar to tbat ol nilvipn.

Kigbt claspcr (Fig. 21 b) cliaraderistic, as .seen in figure.

Enntile-, lengtb 2. 7 mm., second antennal segment I .1 mm., tbird

segment 0.9 mm.; pronotmn, lengtb 0.5 mm., widib at base

0.9 mm.

Simibu in tolour lo inale bui br;u bypterous, wiibout cuneus,

membiane and (lav;il sulure, tbe second anlemnd segmeni mo-

deialely inciassate calli ol pionotum noi noticeidily piomineni.

SciELO

10 11 12 13 14 15

Fig. 13

A — Cyrtorhinus lividipcnuis, hcad and pronotuin, spccinicn from índia.

II — Idcm, spccitncn from Fiji.

C — Idcm, acdcagus.

D — Idcm, K «tructurc of bursa copnlatríx.

E — Idcm, pygophore.

F — Idcm, Icft claspcr.

G, II — Idcm, right claspcr.

Fig. H

A — Cyrtoriuniis mclanops, hca<l and prnnoUiin of nialc.

H — Idcni, prctarHiis.

C — Idcni, K stnicturc of biirso ('o|)idatrix .

1> — Idcni, pygophorc, dornal vicw ,

E — Idciii, pygopliorr, Inlcral vicw.

F — Idcin, Icft cla.vpcr, vcntral view.

G — Icft, cla!i|>cr, inirrnal lateral vicw.

H — Idcni, riglit rlav|H-r, dorsal vicw,

I — Idcni, riglit cia.sper, internai lateral view.

f — Idetn, aedragiis.

Fig. 15

A — Cyrtorliinus cariris, licad niul iironotum c>f frmalr.

II — l(U-ni, pygopliorc .

C — Idcni, prclarsu».

I), E — Idcin, Icft claspcr, internai and dorsal views.

F — Idem, right claspcr.

G — Idein, K striicturc of bursa ropnlatrix,

II — Idem, aedeagns.

SciELO

cm 1

Fig. 15

SciELO i'o

SciELO

cm 1

Fig. 17

SciELO i'o

SciELO

cm 1

Fig. 18

Fig. 19

A — Mccomma amicus, inacrupicruus fcnialc.

B, C — Idcm, right cla^pcr, dorsal and vcntral viesvi.

I) — Idcm, a|>cx of vcsical appcndagc.

E — Idcm, Icft clasper, dorso lateral vicw.

F — Idcm, K strueture of bursa copulatrix.

G — Idcm, preCarius.

Hr. 2(1

A — Nfccomnin graiidin. Icnialc.

U, E — Icft «lasper: II — M. gilvipca.

n — M. limbiilaii.s.

E — M. lurtiiosa.

E, J — K slriicturr of buiu ropiilatrix.

F — M. gilvipc».

(> — M. liKliiosí parifica.

11 — M. luctuosa lurtiio.ta.

I — M. anibiilaiM.

j — M. aiKciinata .

Fík 21

Fig. 21 A, F^ — Kiglit cla.sj>cr of M('a>iiiui:i

A — Uictuosa pacifica.

1$ — antc-niiata.

C — gilvipcs.

I) — lurtiiosa Itictiiosa.

E — anibiilnns.

F, J — .Spiciili of acclragti.s of Mccciiiiiiia .

F — ambiilans.

(i — gilvipcs.

II — antciiiiata.

I — lucliiosa parifica.

J — luctliosa liictuosa.

Fír, 22

Fig. 22 A — Mccomma amiciis, brachyptcroii.s fciiiaU!.

B — Mccomma ambulans. brarliy|)trrnii<i fcmalr.

C, E — Cainciiv.

C; — M. amiciis.

I) — M. oricnlalis (orm b.

E — M. oricntalis fomí a.

F, I - Eggs.

F — T. mundtiliis.

G — C. carieis.

II — M. ambulans.

I — M. oricntalis forni b (ovaríaii cgg) (f aflcr AVilliaiiH, lOSIj (i, II

affrr Kiillrnlicrg, 1912).

Fig. 23

Fig. 23-25 — Mccomma, scallcr diagrani.i u( «liagnoslir niraMirrtncnU: Fig, 23

— nialrü; Fig. 2í — malc»; Fig. 2.5 — fciiialM. AII mra<urrmrnt%

in min., I>a.<ird on tal>ch I and II.

0 ambiilan.i

H ainiciis

Q oricntali.<i nrientalis fnrm a

X nrientalis oricnialis fnrm h

□ nrientalis liimalayensís

CIarvai.iic) & Soirmwooi) — C^yrtoi liimis-Mccoiiinia (.'omplex

Genitalin: K structurc (Fig. 21 H) sliglitly curvccl, tapering to

ajjcx .

Dislril/tilioti: Calilornia, U.S.A.

SpcciincDs sltidied: 5 (} .‘5 9 iMuir \\5)ods, Marin, Califórnia,

I5/7/I7, F. Miiir and \V. M. GilTard, 1^59 San Francisco,

Calilornia, 21/7/17, \V. iM. Giflonl.

Diffcrs froni M. gihipes in tlic colour of body which is

noticcahiy brown tcnding lo castancoiis, witb a dark brown first

antcnnal scgincnt aiul in tbe structurc of thc rigbt clasjtcr and

sj)iculnin of acdcagiis.

MECOMMA MIMETICA n. sp.

(Fig. 18 A)

(iliaractcrizcd by its colour, strongly convcx calli, vcry sliort clytra

and noticeably clavatc sccond antcnnal scgincnt.

Icmalc: Icngtli ‘5.2 nnn., widtli 1.5 niin.; bcad, Icngth 0.5 min.,

widib 0.8 inni., vertcx 0.37 mm.; antcnnac, scgmcnt 1, Icngth

0.3 nnn., 11 I .-1 inin., 111 0.8 nnn., IV 0.3 nnn.; pronotuin,

Icngth 0.5 nnn., widtb at base 0.0 nnn.; rostruni, Icngth

I . I nnn .

Colour black wilh rcddish or castancous tinge, strongly shining;

antcnnac (cxccpt clavatc aj)cx of .sccond scgmcnt), tvvo sjiots on

vertex ncar cycs, rostruni and Icgs i)alc fulvous, thc b:iscs of fcinora,

trot luintcrs and apices of coxac |)alc white, hind tibiac towards

b;isc and b:isc of c()x:ic fuscous; clytra palc transluccnt with a

transversal dark brown faseia; conncxivinn witb rcddish to palc

tucas 011 imicr niargin.

I Icad strongly roundcd, with a short ncck, pronotuni slighily

widci' at b;isc than long, r:dli vcry strongly raiscd with ;i wide

lurrow bctwccn thcni. scíond :intcnnal scgmcnt strongly ( lavalc on

iipical h;dl, clytr;i vcry short rcadiing only to third abdominal

scgmcnt .

unknown.

Ilolotypc: Icimdc, .\spcn Giovc, Nicola, Hritish Columbia.

-1/8/32. G. I. S|)cn(cr, l’rof. R. L. llsingcr’.s colicction; parnlype:

Icmalc, same dat;i ;is typc.

■|'his spcíics dilicrs from olheis in thc genus (Icimdcs) in lhe

vcry shoit clvira. < lavalc setond antcnnal scgmcnt, colour of lhe

Boi.. Mus. Gokldi — Tomo XI (I) — Setembro 1955

elytra aiul legs, tlie sliape of lieatl and very promineiue jironotal

calli. It was taken togetiier witii tlie icimeiimconidae {(ielis sp.,

Hernitelwi, Cryptiiiae) wliich it strongly mimies.

MECOMMA MELANÜCKPIIALUS (POFPIUS)

Nycticapsxís xnelanocephalus Pop])iiis, Acta. Soe. Sei. PTnn. -14 (5):

74, 1914.

Feiiiales Popjiiiis description (translatcd);

Mead, pronotum, sciiteliim, pro— and mesostcnnnn black; collar

anteilorly, ealli, a longitudinal faseia at middle of disk, apex of

scutellum widely, hemelytra, mesosteriunn posteriorly, metastennim,

nnderside of body, rostnnn, first antennal segment and legs yellow;

apex of clavus, coritnn internally and middle of metastenium

lateraly, dark; membrane yellowisb brown, strongly iridescent,

extreme apex of rostnim, second antennal segment and a])ex of

clyjiens black; two last antennal segments dark-brown .

Lengtli 2.8 mm., widtb 0.8 mm. Nyassa Lakc.

Po])|)ius jirobably stndied a macropterous female of tbis species

and nntil furtbcr sjjecimens are fonnd and stmlied (none Avere

available to tlie antbors) \ve piefer to inclnde it in tlie geiuis

Mecomnui Fiebcr and not in Cyrtorliinus as previonsly considered

by Carvalho (1952).

MECOMMA GIEVIPES (,S1'AL)

Lrptomcrocoris ^iloipes Sf,\\ . Stett. Knt. Zeit. 19: 187, 1858.

Mecnmma Renter, liem. Gymn. Eur. .8: 380, .555, pl. 2,

fig. 0; 1883.

CliliDnydiiliis ('ihiipes Renter, Ofv. E. Vet. .Soe. Forli. 21: 57, 1879.

Cliaracteri/ed by its colonr and genilalia.

Malc: Icngtb 4.7 mm., width 1.3 mm.; Ilead, length 0.3 mm.,

Avidtli 0.7 mm., vertex 0.35 mm. Antcnnae, segment 1. length

0.4 mm., II 1.4 mm., III 1.2 mm., IV 0.5 mm.: Pronotnm,

length 0.4 mm., Avidth at base 1.0 mm.; Cuneus, length

0.928 mm.

Antcnna. hcad (except two ohsolele spois near the eyes),

pronotnm, stntcllnm, black to dark brown: clavns inínscatc, black

along (ommissiirc, coriimi tratishicid (extept on (imimissinc);

nnderside of body black, legs pale.

SciELO

10 11 12 13 14 15

Carvai. ho &; South \v(){)i) — Cyrtorliimis-Meconuna Coniplex

Genil/iliti: acdeagiis uith tipical spiculinn (Fig. 21 G) branchcd

aj)ically, tlic largcr bianth provided with a lairly large prong.

Lcft clasper (Fig. 20 H) as scen in figure, the apical portiou tapering

and not swollen as in otlier sjjecies. Riglit clasjier (Fig. 21 C) as

seen in fignrc.

Fetnale: length 2.7 nnn., width l.fi nim.; head, length 0.4 mm.,

width 0.8 mm., vertex 0..S9 mm.; antennae, segment I, length

0.3 mm., II 1 .0 mm., 111 0.9 mm., IV 0.3 mm.; jjionotnm,

length 0.3 mm., width at base 1. 0 mm.

Black exccpt the base of third antennal segment, two spots

ncar the eycs, hemclytra, rostrnm and legs which are translncent,

base of coxae dark.

Genitnlin: K strnctnre (Fig. 20 F) with a hnmp on tlie externai

margin, tlie apex somewhat acnte.

Distribtilion: Dcscribcd originally from Sitka, y\laska.

Spcciineris studicd: several males and femalcs, Anchorage,

27/7/918; Falmer, Vlll, 918; Valde/, VII, 918, Alaska, R. I. Sailer

col.; 3 femalcs, Ketchican, Falis Creck, Alaska, IX, 951, B. Malkin

col.; male and female, Willow, Alaska, VII, 918, F. S. Blanton col.,

several males and femalcs, Pojjoff Is., VII, 1899, Ilarriman Ex])cdi-

tion; I m;de, Wrangel, Alaska, VIII, 1951, B. Malkin col.

Fhis species differs from M. lucluosa (Provancher) in the

complctely black first antennal segment of femalcs and in the typical

male genitalia; from M . ambulans (Fallcn) it differs in the

Iranslnceiu hemclytra of femalcs and in the male genitalia.

With cxception of Alaskan records for this species, all rcfcrnccs

to ('ilvipes (Stal) concern cither to luetuosa lucluosa (Provancher)

or luetuosa j>acifica n. snbsp.

MECOMMA I.UCTUOSA LUCTUOSA (PROVANCHER)

Clilamydatus lucluostis Provancher, Pet. Fann. Ent. Can. 3: 137,

1887.

Mccointua f^iluijjrs Knight, Comi. Nat. Ilist. Snrv. Bnl. 34: 510,

1923; Blatchley, lict. E. N. Amcr.: 852, fig. I7f), 1920; Slater,

lowa St. Coll. jonr. Sei. 25 (1): 52, pl. 0, fig. 19, 1950.

(Fig. in Bhitdilcy, I92())

Malr: Ictigth 1.5 mm., nidth 1.2 mm.; head, length 0.3 mm.,

width 0.7 mm., veiicx 0.21 mm.; antennae. segment I, length

(iO

Hoi.. Mus. CioKi.Di — 'lomo XI (I) — Sctcinljio 1955

0.4 inin., II 1.5 inni., III I. .4 mm., IV 0.4 mm.; pronotiim,

Icngth 0.4 mm., wicith at base 1.0 mm.; ciineiis, length

0.88 mm.

(loloiir hlack to dark brown, cxcept sjtots iicar the cyes, rostrum,

Iicmc4ytra aiul Icgs whicli are [tale lo traiisliicent, rlaviis and ape.K

of ciinetis inliiscate; lindcrsidc of boily black.

GeniUilia'. sjjicidum of aedeagiis (I''ig. 21 J) ciirvcd apically

in almost a straiglu angle. Left clasiier (Fig. 20 E) as seen in

figure. Riglu clas]jer (Fig. 21 D) as seen in figure.

Feinale: length 2.7 mm., width 1.‘5 mm.; bead, length 0..4 mm.,

width 0.9 mm., vertex 0.50 mm.; tuUennae, segment I, length

0..‘5 mm., II 1.1 mm., 111 1.0 mm., IV ()..4 mm.; jnonotum,

length 0.5 mm., width at base 0.9 mm.

Illatk except for basal half or more of second tintennal segment,

lirst antennal segment, two s])ols near the eyes, hemelytra, legs

and rostrtim.

Geniinlia: K strueture (Fig. 20 II) somewhat similar to that

of M . luetnosa j)iicificn n. subsp.

Specimens sludied: 7 males and 5 females. Machias, Mc.

Janson col.; 2 males and 2 females, (flen llouse, N. H., Parshley

col.; 5 males and I female, Mt. Washington, 1’arshley col.; 2 tnales

and I female, East Fort Me., Farshiey col:; I inale and I

female, Fulton (io., N. Y., Farshiey col.; I inale, I.ake Flacid,

N. 5'., Farshiey col.; several males and lemales, Monimor (ianada,

Llhier col.; Indian I.ake, N. Y., 11. Cl. llarber col.; 2 males,

Illinois (brooklin Museum); several males and females, .Smith River,

(lalifornia, .Mdrich tol.; females, Liberty (lo.. Florida, V, 924, F. II.

I lubbcl col .

Flie s|)ecies was originally described from (lap Rouge, (lanada.

'Fhe series from .Smith River, (lalifornia heie referreil to this

species ((jiild not be sepaiated from the typical lurluo.sit (Frovan-

(hcr). Ii may actually have a (ontinuous disiribution atross the

northciti 1'niied States of .America. ,\ll retords for gt/ctpe.v (Sl;d)

knotvn up to date Irom the eastern United States and (lanada should

be relerred to tyjiit al luí liiosii .

'I his stibspecies iliflers from hirlu<i.sit pntilini n. subsp. in llic

black liist antennal segment ol the males, in the imich longer cuneiis

and in lhe shape of lhe spiciilum of lhe aedeagus. 1 he species can

10 11 12 13 14 15

Carvalho & Soiniiwooi) — Cyitorhimis-Mecoinina Coniplex

l)c reaclily separatcd IVom the otlicrs iii the genus by the pale

íirst antennal scgment of fcinalcs and iii the structure of the inale

genitalia.

MECO MM A LU cru OS A PACA EI CA n. snbsp.

Cdiaractci i/ed by its coloin and length of cnnens on males.

Mdle: length '1.0 nnn., width l.;5 nnn.; head, length 0.2 nnn.,

width 0.7 nnn., vertex 0.‘f‘l nnn.; antennae, segment I, length

0.1 nnn., II 1.5 nnn., III 1.5 nnn., IV^ 0.5 nnn.: pronotinn,

length 0.1 nnn., width at ba.se 0.0 nnn.; rostrinn, length 1.5

nnn.; cnnens, length 0.()07 nnn.

Cooinr daik biown to black; first antennal .scgment, base of

tliird segment, two spots near the eyes, hemclytra, rostnnn and Icggs-,

pale to dnll yellowish tianslncent, tinged with fuscons along claval

and roti;il coinmisinc; veins of membnine and third segtnent of

tai si bfown to castaneons; nndersidc of botly dark brown .

Oeniíalia: aedcagns (Fig. 21 I) with a spicnlnm broatlly cnrved

apically. Left clas])er (Fig. 20 lí) strongly cnrved and providcd

with a sivollen ajiical arcas covercd by minute teeth. Right clasper

(F'ig. 21 A) as seen in lignre.

Eeiiidlc: similar to m;de in colonr, brachyjiterons, the elytr;i as in

diiteinidln. Length 2. 7 mm., width 1. 5 nnn.; head, length

0. 5 mm., width 0.0 nnn., vertex 0.5() mm.; antenmie, segment

1, length 0.5 nnn., II l.l nnn., III 1.0 nnn., 1\' 0.5 mm.;

pronotnm, length 0.1 mm., width ;it b:ise 0.8 nnn.; lostrnm,

length 1.5 mm.

Ceuildlid-. K strnctnre (Fig. 20 O) broail basally, tajicring to

apex, only slightly t iirved.

H<)l(>lyj)c: male, Iftukley, \\':ishington, \'II, 055, Oimin col.

in lhe collettion of the U.SN.M; dllolyfic: lemale, same dat:i ;is tyjie:

l)drdtyf)CS'. 22 males and feimdes, Whangel, Heanderc, Dncan Canal,

Alaska, \’lll, 1051, b. Malkin (ol.; 6 males and 5 females. Forks,

Clallan Co.. Washington. \'II. 1020, lí. P. Van l)n/ee col.; I

lemale, ,S;isni(h Dist. 15. C., VIII, 1018, \V. Downes col.

Fhis species dillers Irom liirliiosd hiiliiosd (Provancher) in the

jiale first iiniennal segment ol males, lhe mnch shorier tniiens ;md

in lhe slriutnie ol male genitalia, espetiidly in lhe shape ol the

spitnhnn .

G2 Bol. Mus. Goeldi — Tomo XI (I) — Setembro 1955

TAIJLE I

Measiirements of males o£ Mecomma ambiilafis, M. amicus, M.

oriefiíalis and Aí. o. hitnalayeusis (all measiirements in mm.)

SPKCIES-LOCALITY

HEAD

PKONOTUM

ANTENNAIÍ LENOTU

Cuncu B

iength

Widil

Vertex

width

Widtl

I/cngtl

‘

iii

Ámbulan*

Kendal, Wcfltm

0.87

1.15

0.60

0.49

1.59

1.40

0.50

1.01

Cambridg^hirc

0.80

0.30

I.03

0 52

0.47

1.52

1.53

0.51

0.97

Ilarijondon, Ilcrts

0.80

3.30

1.I0

0.50

0.47

1.43

1.38

0.52

0.84

** ”

0.83

0.35

1.01

0.50

0.45

1.33

1.23

0.48

0.7S

M .f

0,80

0.37

1.07

0.50

0.45

1.51

1.39

0-53

0.87

M M

0.87

0.37

0.98

0.40

0.47

1.13

1.38

0.52

0.86

Amicui

Kuraeong, Himalayas

0.80

0.37

1.09

0.52

0.45

I.4G

1.20

0.48

0.74

Sikkim

0.79

0.37

0.99

0.50

0.40

1.28

1.10

0.39

0.70

0.82

0.37

0.91

0.53

0.44

—

0.09

0.80

0.37

0.90

0.49

0.41

1.25

1.08

0.37

0.69

0.83

038

0.94

0.61

0.45

1.32

1.15

0.40

0.72

Ukhrjl, Himalayas

0.81

0.37

0.95

0.48

0.40

1.29

1.12

0.76

Darjpeling

0.82

0.39

0.92

0.52

0.40

1.50

1.25

0.45

0.73

0.80

0.30

1.08

0.51

0.43

IJO

1.20

0.73

Orifntalii crievtfjlit

Form 0

S. Ind*a

0.08

0.32

0.92

0.47

0JJ8

1.48

1.12

0.40

OJjO

••

0.70

0J16

0.95

O-IÜ

0.38

1.40

I.1Ü

0.40

0.79

0.70

0.31

0.91

0.47

0.38

1.43

1.14

0.40

0.79

0.09

0.33

0.92

0.48

0.39

1.42

1.10

0.41

0.78

0.71

0.34

0.97

0.49

0.3S

1.47

1.12

0.44

0.78

Mysorc

0.72

0.35

0-97

0.48

0.39

1.52

1.13

0.00

0.72

fl. índia

0.71

OJÍI

0.05

0.47

0.30

1.31

1.03

—

0.70

KodniKanal, fe. India

0.75

nj7

0.05

OJiO

0.37

1.45

1.12

0.75

Form b

8. índia

0.72

0.32

0.85

0.4J

0.38

tJ7

i.no

0.44

0.05

0.71

OJI

0.82

0.50

0.31

1.27

0.08

0.45

0.0 1

0.70

0h3I

0.7(1

0.60

0.30

1.15

0.93

0.49

0.03

KiKlaiKanat, 8. InJia

0.70

0.30

0.87

0.51

0J7

0.02

0.72

Nilgiri Jlills

0.70

0.31

0.80

0.49

0.37

1.20

0.90

0.52

0.04

Orientoiti himnlyrntii

Sikkim

0.74

0J2

OJiS

0.50

0.37

Ü.OO

0.78

035

0.87

0,50

037

0.02

Kunif^ong

0.73

0.33

0.88

0.4S

0.39

1.29

0.Í8

0.40

0.02

Carvalho &: Southwood — Cyrtorliinus-Meconinui Coniplex

TAIiLE II

Mcasurcmcnts of^nialcs of Mecomma ambiãans, M. atnicus, M.

amicus and M. orienlalis (all nicasurcnicnts in inm.)

SPECIES-LOCALITY

HEAD

PRO.VOTUM

ANTENNAE LENGTH

Width

Vertex

widtli

VVidtl

Lrngtli

s t

tS-

Xmòu/ori«

Ilarpcndon

0.SS

0.42

0.96

0.48

0.40

0.10

1.10

0.50

0.73

Knst IVckham

Ü.PO

0.42

0.90

0.60

0.38

1.11

1.10

0.51

0.70

(íranKc, Lanes

Ü.S8

0.40

0.89

0.47

0 37

1.15

1.12

0.51

0.75

Nowby Bridíte

0.9Ü

0.41

0.92

0.48

0.39

1.20

1.12

0.54

0.75

Kcndal, Westm

0.92

0.40

0.92

0.51

0.40

1.24

1.11

0.52

0.75

Amicxi»

Darjcellng

0.84

0.37

0.85

o.to

0-36

0.90

—

0.70

0.S6

0.30

0.87

0.42

0.36

—

...

0.67

0.S4

0.39

0.S6

0.40

0.33

0.80

0.75

0.40

0.68 -

0.83

0.38

0.84

0.40

0.34

0.85

0.70

0.32

0.64

0.83

037

0,86

0.41

0.36

0.87

0.76

0.35

0.66

0.S4

0.38

0.85

0.41

0.35

0.91

0.75

0.32

0-64

Sikkim

0.S5

0.39

0.89

0.40

0.36

—

0.65

Orirntalu oríentait

Farm o

0.S5

0.38

0.86

0.41

C.35

0.67

Nilgiri Hilla

0.70

0.31

0.72

0.43

0.30

0.S7

0.74

—

0.68

* *

0.09

0.30

0.73

0.4A

0.31

1.17

0.87

0.30

0.70

S. índia

0.70

0.32

0.70

0.41

0.30

1.18

0.79

0-36

0.73

'■

0.73

0.32

0.73

0.44

0.32

1.26

0.90

—

0.73

Farm b

8. índia

0.75

0.32

0.75

0.45

0.31

0.92

0.74

0.36

0.75

0.74

0.32

0.74

0.44

0.32

0.96

0.75

'1 4i.

0.73

0.75

0.33

0.75

0.45

0.30

0.87

0.76

0-34

0.75

0.76

0.32

0.75

0.45

0.30

0.96

0.75

—

0.73

0,74

0.32

0.75

0.49

0.32

l.OO

0 76

0.74

ni()LX)GY

Ptei-ygo-polymorphism

In niaiiy llctcroptcra varyiiig tlcgrccs of wiiig rcduction arc

known. riiis pliciioincnon, firsi revicwcd by Pcncaii (1905), lias

subscxjucntiy bccn slndicd by maiiy autliors; tbc most rcccnt general

stiulíes arc by Larscn (1950) and I*oisson (1951).

All stages of wing retliution, froin tlie fiilly developed lo tbc

ajitcrous condiiion arc rarely fomul in one speeics (c.g. Gerris

lacustris 1,. — 1’oisson. 1951). Witbin tbc Cyrtorhiuus-Mccomma

(oniplcx (old .sensc) tbrec grades ean be fixed:

(a) fully developed lieinclytra manopterous (Eig. 1 A)

(b) rednred lienielytra, incinbrane present scmi-bmchypleroiis

(Fig. 8 A)

(il

IJoL. Mus. (ioia.i)! — "ronio XI (1) — .Setembro 1955

7 A)

A variety of intermecliate coiuiitioiis exist betwecn tbese grades

(.Stehlik, 1952), biit tbese are generally imieb rarer.

It is iiot known to wliat extern tliis reductioii is enviroiimcntal

oí gcnctical I)ut in Mecomvin it is related to sex, lor tlie males are

always fully develojied and tbc females are generally bracbypterous.

Occasioiiidly macropterons íemalcs are loiind and even more rarcly

semi-brac liypteroüs individuais and otber intcrmeiliates . 'I be prodiic-

tion oí macrojjterons forms in lemalc M. amhitltiiis scems to Ite

associated witb citbcr nortiicrn latitude or monntainons regions

(Stehlik, 1952, in Czechoslovakia, Soiithwood, nn|)blisbed, in Gt.

llritain). Sncb a pbenomenon could bc a direct environmental cllect

or tine to a selection lactor more lavorable to tbe macroj)terons.

forni in tbese localities. Stehlik says tbat in general monntain-macrop-

terism is loiind in species tbat overwinter in tbe egg condition,

wbilst tbe reverse (a tendency towartls bracbypterism) is fonnd in

sjjeries tbat overwinter as adiilts.

'I lie sex in wing rediu tion oeenrs and its degree are cbaractcristic

for eadi gemis now recognised in tbe Cyrlorhinus-iMcrotnma

comjjlex: —

Cyrlorhinus

in female only

Meco mina

all species, otber conditions very rarc

(Af. mimelica female micropteroiis)

Fielierocapsiis — in botb sexes, bracbypterous and more rarely

inaci ojitcrons condition .

Tytthus — complete bracby|)terism is found in tbe male of nlhoornn-

lus, a semi brocliypterism in tbe females of jiiibesccns anil botb

sexes of gci/j/ni/.s.

Fccding liabits

Mnir (1920) foiind tbat T. miniiliiliis in Qneensland lived

excliisively on tbe eggs of tbe sngar-cane leafboppcr, PnhinsicUn

sccharicida Kirkaidy; wlien tbis miiid was introdnced into llawaii

it brongbt abont tbe lontrol of tbe lealbop|)ei' (Swe/ey, líI.St)).

Snbseipient workers, notably Usinger (I9.‘i9), bave lecoiilcd similai

babits for 7'. cliinrn.sis, C. fuhma and C. liindi jieiniis (tbe details

are givcn nnder eaib species. l'.iiro|)ean antbors (e.g. Knllenberg

1910, Wbigner 1952) bowever bave generally considered tbeir species

to be pbytopbagons. Knllenberg obseited iiiiicis feeding on

varimis Caiex and Sciijni.s species, biit Usinger points ont tbat even

Carvalho & Soüthwood — Cyitorhimis-Meconiina Coiiiplex 65

when thc bug apj>earcd to be íccding on an unbrokeii plant surface

thcre was always a Dclphacid egg prescnt, oíten laid from the other

sitie of the leaf or steni. Soüthwood (unpublished) lias noted that

C. carieis and T. pygmaeus are always fonnd togethcr with large

nunibers of Delphacids, especially Conomelus limbatus Fab. and

Massee (1954) records T . pygmaeus feeding on the early instar larva

of a leafhopper. Kidlcnberg considered Mecomma ambulans to be

phytophagous, but littlc is known of the biology of this gentis.

Immature stages

The eggs of 7'. mundulus are laitl in the leaves of the sugar

cane, fretpiently in an old leafhopper egg slit (Williams, 1931);

they are of typieal iniritl forni and the opcrcnluin aiitl microjiylar

region just projects above the Icaf surface. Tliose of C. carieis and

M . ambulans which have been described hy Kullcnberg (1913) are

laid in the steni of varioiis species of Scirpus and grasses rcspectively

(Kullcnberg, 1916) and the ovarian egg of M. orietitalis is figured

lielow (Fig. 22 I). The eggs of these Orthotyline spccies ajipear

to be niuch more strongly curved than thosc of T. mundulus.

lhe young nynijihs of T . mundulus are bright red in colour

and also feed on leaf hopper eggs (Williams, 1931); Butler (1923)

dcscribes the first instar of C. carieis as orange, though later instais

of this species and of T. pygmaeus and /•'. flaveolus resemble thc

adiilts in general coloration (Butler, Soüthwood unpub.).

Habitat

In temperaie regions the species of Cyrlorbinus Fieberocap.ius

and 'Fytthus are found around the bases of tufts of various rushes

(Juncu.s), sedges (Carex) and grasses growing in very damp or water

logged sitiiations. This is closcly correlaled with thc tlistrihution of

Dclphacid eggs, which as indicatetl ahove are probably their majoi

food. In troiiical regions thc habitats are analagous; for e.xample

bamhoo grass, young rice, sugar cane; it seems tliat here they aro

more rcadily taken by swceping than in temjierate areas.

Mccotnma ambulans is iiormally found amongst tlaiiiji grass,

especially in or around lemperate woodlands. From what is known

of lhe ilistributioii of M. amictis and M. orientalis logclher wilh

lhe collectors note that lailer was aken "sweeping gra.ss”, it would

seem that they are confined lo similar situatioii, which in the liidian

suh'Contiiicnl are only found over 5,000 ft. It is noleworthy that

the solitary Atrican sjiecimen was taken ;it 9,000 fl. at lhe edge ol

the cold temperate l)jem-l)jem Forest, Abyssinia. lhe Ncarctic

tommon neartic spccies accoiding Blalchley occuis upon rank

herbage iti moist shaded locations.

— 3 _

()() Hoi,. Mus. Gokldi — Tomo XI (I) — Setembro 1955

Thus it woukl secm tliat thc genus Mecomma is always associatecl

with tlamp grassy arcas in temperatc, mostly broadlcaved, woodiands.

The males of M . ambulans are very active and fly readily; whcn

alive they have a marked superficial rcsemblance to parasitic Hyioc-

noptera; tliis probably appiics to the wliole genus. M. miineticn

mimies tlie Ichnemnonid, Gclis sp. and was found witli it.

ZOOGEOGRAPHY

Tytthus is thc most witlcly distribnted of the foiir gcncra revised

in this paper, it oceurs in all tlie major zoogcograpliical regions.

T. pnn>iceps is jtarticularly notcworthy witli its wide distril)ution

from Floricia Central America, Venezuela and Paraguay in thc west,

to thc Rodriguez II. and the Seychclles in thc cast, whilst northwards

it has bccn recorded from Giglio I., Italy, by Mancini (1952). As

T. pamiceps oceurs on so many islands, espccially isolatcd ones

like St. Helena, it can be assumed that it has obtained this wide

distribution in comj)arativeIy recent times. In thc Oceania two

spccics oceur, chinensis and itmndxilus; on |)rcscnt knowledgc the

ranges of thc spccics appear to be distinct, rniindulus oceuring in

Melanesia and chinensis in Micronesia. T. gerninns and T. pyg-

inacus have ovcriapping ranges in thc Palcarctic, whilst T. pubes-

cens and T. mgns are jjrobably an analogous pair of non-ailied

spccics in the Ncarctic.

Cyrtorhinus is absent from thc Américas but is present in both

tropical and temperatc regions. C. curnberi, in many ways the most

primitive spccics, oceurs in New Zcaland; whilst C. fnlvns, which is

similar lo it in many respects, is lound from Java to Samoa .

Ovcriapping with C. fnlxnis, but extending much íurther west into

China, llurma and índia is C. lividipennis, a spccics having

affinitics with both C. fnlims and the Klliiopian C. tnelanops. Set

somcwhat apart from thc othcr spccics structurally is the Ilolarctic

C . carieis. l he distribution of thc .spccics of Cyrlorhinns and its

correlalion with their structural rclationships shows that it is an

old genus; this is further supjtorted by the way thc sjiccies are

clearly separated.

In contrast to Cyrtorhinus, Mecorntna is a genus of closely

rclaied spccics, often very similar in strueture and confined to the

broadlcaved or mixed forest of temperatc region. Whcn piescnt

in (•(jualorial regions, they oceur only on tlie mountains (e.g.

Nilgiri llills, ü. índia) where this type of forest oceurs. Of the .'I

ncarctic spccics known before, existing records show that two are

restrictctl to the Pacific coast, antennala being known only from

lhe San Francisco llay arca of Galífornia and gilvipes from the Coastal

Carvalho &: Sournwooi) — Cyrtorhinus-Mecomma C.oinplex

region and sontheastcrn Alaska west to tlic treelcss Aleutian Islands.

The third spccies, Aí. luctiiosa occurs from coast to coast across

soiithern Canada and northcrn United States. Most of its range is

oceupied by the typical subspccies which is known froni such widely

distributed localities as northern Califórnia, Michigan and Florida.

The other subspecies appears to be restricted to the Coastal region

from southeastern Alaska soutli to Washington. In índia, Aí.

orientalis, a distinct subspecies occurs in the E. Himalayas, separated

from the typical subspecies by the Deccan plateau and Canges

Valley.

Fieberocapsus is represented by one sjjecies, E . flaveolus whose

range is confined lo northern Europe. Structural evidences shows

that vvhilts Mecovirna is very dose related to Cyrlorliinus, Eiebero-

capsus belongs lo another branch of lhe Orthotylinac.

CHECK LIS r

(Stíb-family Phylinnc)

Tyuliiis lüclKr 18G1

Cylloccps Ulilcr 1893 (nov. syn.)

rcriscopiis Brcdditi 1890

Brctldinicssa Kirkaidy 1903 (nov. syn.)

zwahiwcnl>urgi Usinger 19'l'l (nov. comb.)

chinensis Stal 1859 (nov. comb.)

clongatus Toppins BJH (nov. cyn.)

annulicollis 1’oppius 1911 (nov. syn.)

riveti Cheesman 1927 (nov. syn.)

parviceps (Rculcr) (nov. comb.)

pclicia Ulilcr

pygmaciis /cuersiedt 1840

pclluccns llolicman 1852

insignis Dciiiglas ic Scott 1800

vagiis Knigbt 1923 (nov. comb.)

ncotropicalis Carvalbo 1951

costae Carvalho 1915 ncc Stal

munduins llrcddin 1890 (nov. comb.)

panamensís n. sp.

allxxirnatns Knigbt 1931 (nov. comb.)

montanus n. sp.

gcminns Mor 18(K)

piibcsccns (Knigbt) (nov. syn.)

balli Knigbt 1931 (nov. comb.)

lns[)cratus Knigbt 1925 (nov. comb.)

lU)i.. Mus. Goki.i)! — Toino XI (I) — Sctcinbro 195.*)

t)8

(Suh-fatJiily Orthotylinae)

Fiel)Ci()(:a|)siis nov. gen.

flaveoliis Rciiter 1870 (nov. comb.)

Cyrtorhiniis Fiebcr 1858

Cyrlorrhiniis Rciitcr 1884 (emenda liou)

Rciiteriessa Usinger 1951 (nov. syn.)

cuml)eri Woordward 1950

fulvus Knigbt 1935

lividipennis Reuter 1884

vitiensis Usinger 1951 (nov. syn.)

niclanops Reuter 1905

mcgalops 1’oppius 1914 error ])ro melanops

raritis Fallcn 1807

cirgantulus Meycr-Diir 1843

chloropterus Ucrritk-.Schacffer 1853

Mcconiina Fieber 1858

Spliyraceplialus Douglas 8c Scott 1865

.Spbyrops Douglas & Scott 1866

.Vntiphilus Distam 1909

,\ristobulus Distant 1910 (nov. syn.)

Nycticapsus Poppius 1914 (nov. syn.)

Aristolxrlus Carvalho 19.52 error (rro Aristobulus (nov. syn.)

Orirntniis nov. sp.

sub. sp. orientalis nov. sub. sp.

sub. sp. himalayensis nov. sub sj).

aniictis Distant 1909

filius Distant 1910 (nov. syn.)

rliinensis Reuter 1905

atnbiilans Fallen 1807

dubius Zcltcrstcdt 1810

ocliii|)es Curtis 1838

nigrilulus /etlcrstcdt 1840

tnadagascatiensis Reuter 1892

melanoccpbaltis 1’oppius 1914

grandis nov. sp.

luctuosa liictuosa 1’rovamlicr 1887

gilvipcs auett. nec Stal 1858

luctuosa pacifica n. siibsp,

gilvijrcs Stal 1858

antennata Van Durce 1917

inlmelica nov. sji.

Spreirs iiicerlae seilit

Chiamydatus collaris Matsumuia 1911

Carvalho & .Souiiiwood — Cyrtoihinus-Mccoiimia Coin)jk‘x

SUMÁRIO

O presente trabalho é uma revisão do complexo Cyitorliimis

— Mecomma (Ilemiptera, Miridae). Êsses pequenos percevejos pos-

suem grande importância económica e larga distribuição geográfica.

Algumas espécies são usadas no combate biológico das cigarrinhas.

Segundo Zimmerman (I94S), a espécie Tytthus mundulus foi intro-

duzida em Ilaiuaii em 1920, proveniente de (.hieensland e Fiji, para

auxiliar o combate à cigamnha de cana de açúcar. A espécie esta-

beleceu-se e constitue um dos marcos na .história do controle por

meios biológicos. Ela contribuiu para a economia da indústria açu-

careira de Ilatvaii com milhões de dólares — o seu valor real podendo

dificilme7íte ser estimado.

As espécies aqui tratadas se acluwam em estado confuso taxa- '

nóirnicamente , existindo tanibé7n dúvidas quanto à sua área

geográfica.

Iniciamos o trabalho com uma introdução e um histórico dos

estudos prévios sôbre espécies do grupo. A seguir, damos os carac-

teres usados na separação das subfamilias Orthotylinae c Phylinae.

Foi incluído um índice das espécies que já foram descritas no com-

plexo com sua posição genérica atual, bem como uma chave para

separação dos géneros nêle envolvidos . Cada género é tratado sepa-

radamente, as espécies descritas e ilustradas com chaves apropriadas

para sua separação. As espécies novas estão descritas. Comentário.-,

sóbre a biologia e distribuição geográfica das espécies foram feitos

e uma lista da bibliografia mais manuseada acha-se incluída no fim

do trabalho.

SUMMARV

A rcvisiuii of llic spccifs hillicito incliulcd iii Cyrtorhinus liclior aiul

Mfcornma LípIkt haj shuwn thal tlic speties witii bristlc-likc arolia aiul ollicr

riiyliiif characters slunild ite placed in tlic genus Tyttlnwi l''icl)cr williin lho

jub-family 1’liylinac'. Tlic Ortliotylinc jpccics arc allotcd to Cyrtorhinus and

to tlic floscly allied Mriotninti, whilst flaivolus Rputer is placcd in a ncw

gcmis Heberocafmv.

Keys arc given to gcncra and .«pccics, wbidi aip rcdcsciibcil and figincd or

in a fcw laitcs wlicrp inaiciial was not availablc for stiidy ibc original dcsiriiitioni

arc given. 'riircc ncu' spcric» of Mrcoinma arc d<"icril)cd. onc of ibem with two

Miljspcricj, onc of tbem wbicb is |M)lyinorphic in lK>th sexes. A ncw snbspccics

of Af. luctuosui l’rov. is drseribed.

Notes arc given on: tbc coinparativc morphology of thc genitalia of tbc

PliyUiiac and tlic Oitbotylinac; on tlic biology of thc spccics, intinding ptcrygsi

polyinorphisin and their nsc in thc biological tontrol of lcafbt>ppcis; and on

their roügcograpby.

70 IJoL. Mus. Goeldi — loino XI (1) — Setembro 1955

ACKNOWLEDGEMEN rs

This Work was carriccl oiii whilst thc scnior aiitlior was worUing in the

U. S. National Museiim iinclcr a Giiggenheim grani and the junior

anthor was in rcccipt ot a grant from thc Agricnltnral Research Conncil; wc

wish to exprcss onr indebtcdncss, lo thcsc anthoritics and to those of thc

following institntions who have pcrniittcd iis to examine thcir collcction:

British Muscnin (Nat. Hist.), U. S. National Mnscum, llcrnicc 1’. llishop

Miiseum and Californian Acadcniy of Sciences,. It is a plcasiirc to rccord our

gratitude to the following colleagnes who have hclpcd tis with advicc on

spccific points or hy thc loan of thcir spccinicns or those in thcir chargc: Dr.

W. E. China, Mr. II. VV. Dallry, Drs. C. G. Joliirson and St. von Kclcr,

Prof. H. 11. Knight, Mr. D. Lcston, Drs. R. Malaisc and A. M. Masscc,

Prof. H. Sachtichcn, Drs. R. I. Sailcr and Jl. .Scott, Prof. R. L. Usingcr and

Mr. G. E. Woodroffe.

REFERENCES

(Only general rcfcrcnces inclnded, pnrciy taxonomic refcrcnccs are givcn in

lhe appropriale placc in lhe Icxt)

ATKINSON, E. 1800. Catalogue of lhe Insecta No. 2. Order Rhynnchota,

sulxjrder Hcmiptera-Hetcroptcra. Family Capsidac. /. Asiat. Sor.

lininal fS (2): snppi. 1: 25-200.

RONH.AG, P. F., \\’ICK, T. R., 1953. The fnnctional anatomy of thc malc

and fcmalc reproduetive systems of lhe milkwecd bng, Oncopellus fasciatus

fDalIas) (llctcroptcra: I.ygacidac). J. Morph,, Philadelphia 93: 177-230.

lUJTLER, E. A., 1923. A Iliology of lhe Rritish Ilcmiptcra-Hclcroptcra.

London .

CARVALHO, J. C. M. 19.52. On lhe major classification of thc Miridac

(Hcmiplcra)^ (With keys lo siihfamilics and trilres and a catalogue of

lhe world genera). Anu. Arad. fírasil. Ci. 21 (1): 3I-11I.

CARVALHO. |. C. M. Xc LLSFON, 1)., 19.52. The classification of lhe lUitish

.Miridac (Hem.), with keys to thc genera. Ent. mon. Mag. 88: 231-251.

CHINA, VV. F.., 1913. Thc Gcncric names of llrilish Insccis. 8. Thc gcncric

naines of lhe llrilish I lemiplera-Helcroptera, with a chcck Ilst of thc

llrilish s|>cries, London.

DUZEE, E. P. van, 1917. ('atalognc of lhe Hemiptera of America Nort ol

México excepting lhe Aphididac, Coccldae and Aleiirodidac. Univ. Cali-

fórnia l’uhs. Ertt., 2, xlv, — 902 pp.

HORVATII, G., 1909. Hcinipicrcs rcciicillis par M. Th. Ilcckcr anx Ilcs

Canaries. Ann. Mus. Nat, Hung. 7: 289-301.

Carvai.uo &: SouTHWoou — Cyi torhimis-Mccoirnna Complex

KNIGHT, H. II., 1921. Sciciuitic resuits of tlic Katmaí cxpeclitions of the

National Gcographic Sonety. XIV. Ilcmiptcra of lhe family Miridae.

Oliio ]. Sei. 21 (3): 107 112.

KIILLKNIIKRG, 11., 1!)13. l)ie Kier der schwcdisthen. capsiden (Rhynchota)

II. Ark. Zool., Slockiwlm 34 A (15): 1-8.

1910. Studien üher die Riologie der Capsiden. Z. 00 I. Kidrag.,

Uppsata 23 (snppl.) 522 pp.

1947. Uher morphologie und fiinktion des kopnlationsapparals der

Capsiden nnd Nabiden. Z.ool. Ilidrag., Uppsala 24: 217-419.

L.\R.SEN, O.. 1950. Die Veranderungen im llait der Ileleropteren liei der

Rednktion des l''Iuga])parates. Opus. enl., l.und., 15: 17-51.

l.E.SrON, I)., 1953. “IMiloeidac” Dallas: syslematics and inorpliology, with

reinarks 011 lhe phylogeny of ''Pentalonioidca” Leach and npon lhe position

of ''Serhana" Dislani (1 Icniiplcra). liev. llrasil. lliol. 13 (2): 121-110.

I.INDllKRG, II., 1930. Die Ileleropteren der Kanarischen Inscin. Comm.

lliol. 6 (7). 43 pp.

GINDIIERG, II. 1953. Ileniiptera Insniaruin Canairiensium (Systeinatik, Okologie

nnd Vcrhrcitnng der Kanorischen Ileleropteren nnd Cicadinen). Comm.

lliol. Soc. .Sei. Fcnn. 14 (1); 1-304.

MANGINI, G., 1952. Matcriali per ona fauna delli arcipelargo Toseano XXII

(I) Eniittcri dell isola dei Giglio. Atiti. Mus. civ. Sloria Nat., Genoua

66: 1-32.

MASSEE, A. M., 1951.

MIIIR, !■'. 1920. Repoit of Enloniological work in Anslralia 1919-1920. llawaii

1’lant. Ilrc. 23: 125-130.

O.SIIAMN, U., 1900. Vcrreiihnis der 1’alaearklischcn llemipieren init heson-

derer herücksichtigung ihrcn verieilung iin Riissischcn Reich I (1 & 2).

Anu. Mus. zool. Acnd. hnp. Sei. II: 1087 p|).

■ 1910. dilto, III. ihid 15, 218 jip.

1912. Katalog der 1’alacarliseheu, Si. 1’ctershiirg 8c Ilerlin, XVI

-(-187 pp.

I’ENEAU, M. ]., 1905. 1’reniicrcs notes sur Ic polyinorphisine des llcmiptcres

dans Touest de la Erancc. C. 11. Ass. franc. Av. Sei. 1905: 502-509.

1’<)1.S.S()N, R., 1951. Ordere des Ilcleropleres In Gra.ssc, 1’. 1‘. ed., Traiu de

Z.oologie, 10 (2): 1057-1803, l’aris.

SCOl T, II., 1950, in JcanncI, R., llaiites Montagnes d'Afriipic. Paris. p. 155-157.

■SHARP, 1890. On lhe strnctnrc of lhe terminal segnicnts in some inale lleniip-

icra. Trans. fiil. Soc. l.oud. 1890 (3): 390-427 (3 pis.).

SINíilI PRU Tlll, II.. 1925. The morphology of lhe inalc genilalia in Rhynchota.

Trans, enl, Soc, l.ond. 1925: 127-207.

S1..\TER, J. A., 1950. An investigalion of the fcinale genilalia as taxonomic

rharacters in lhe Mirid.ic (Ilcmiptcra). lowa St. Coll. J. Sei. 25: 1-81.

15oi.. Mus. (ÍOKI.I)!

loiiu) XI (I) — Setembro 11)55

TÀ

.SOUTHVVOOD, T. R. E., 1953. The inoipliology and taxonoiny of thc

geniis Orthotylus Fieber (Hem., Miridae), wilh spedal rcfercnce to the

Itritish spccies. Traus. H. ent. Soc. 104 (11); 415-149.

.SIEHLIK, J., 1952. The fauna of Heteroptera of the inountain High jesenik.

Acta Mus, Moraw., 37: 131-248. [In C./edi., with English and Riissian

siiinmartesj.

SriCHEL, \V., 1925-38. llluslricrte liestimmungslabelUu der Deutschen Wanzeti

{Ilemiptera Iletcrojitera), Bcrlin.

,SWE/.EY, O., 1930. Iliological control of the sugar canc I.cafhoppcr in Ilawaii.

Buli. Haxvaii Sug. Ass. ent. Ser. 21: 57-101.

USINGER, R. L., 1939. Distribution and host relationshi|)s of Cyrtorhinus

(Hemiptera; Miridae). Proc. llaw. ent. Soc. 10 (2): 271-273.

1940. Insects of Guam II, Ilemiptera, Heteroptera of Guam. llull.

II. P. liishop Mus. 189: 9-103.

WAGNER, E., 1952. Blindwanzcn order Miridcn in Pierurlt Oeutschlands 41,

218 pp., jena.

WILLIAMS, F. X., 1951. The insetts and othcr inverlebrates of Hawaiian

sugar cane ficids, Honolulu. 100 pp.

WOODWARI), T. E., 1950. New rccords of Miridae (Heteroptera) from New

/.caland, with descriptions of a ncw gemis and four ncw spccies. Pec.

Auck. Inst. Mus. 1: 9-23.

ZIMMERMAN, E. C. 1948. Heteroptera, Insccts of Hawaii 3, 255 pp., Ho-

nolulu.

fík. I

A — Lateral viow of at-deagii.s ot C. carieis, wilh siiiiport and part of

floor of pygophorc.

II — LnngitiKlinal hall of inale pygopliore of L. earicis (diagranialie)

to Show striictiire of aedeagal snpport or siihgenital plale.

C — l’ostero-ventral view of pygophorc of C. carieis, clas|)ers removed.

I) — yVedeagiis wilh aedcagiis shealh of T. pygmacus, lateral view.

E — Anterior view of pygophorc of 'P. pygmacus, witii clas[)crs and

aedcagiis rcinovcd, showing tubular striicturc of aedeagal sheath.

F — Lateral view of prctarsiis of C. carieis (Orlholylinac) .

(í — Lateral view of pretarsus of T. pygmacus (1’hylinae).

1 — lateral arin of basal platc; p — promotor apodeme of lhe

phallobasc; ph — parainerc (or claspcr) htiles; s — aedeagal sheath

( = aedeagal support or siiirgenital plale); I — theca; v — vesica;

va — vesical ap|>cndagc or spiculuni; bp — ba.s;il platc.

Fig. 2

A — Tytlhus chincnsis, iiialc.

It — Idcni, acdcagus.

C — Idciii, Icft claspcr.

1) — Idcni, right claspor.

E — Idciii, pygophorc .

ríR. 3

A — Tyttluis pygmacus, hcad aml thorax of inalc.

B — Idcni, pygopliore.

C — Idciii, acdcagus.

D — Idcm, apcx of pygophorc with claspcra and acdragus romoved,

E, H — Idcin, Icft claspcr.

F — Idcin, riglit claspcr,

í; — Ideni, prctarsus.

I, J, K — Tytlhiis parviccps, Icft clasi>cr.

L — Idcm, pygopliorc.

M — Idcm, riglit claspcr.

Fír. 8

A — Tytthus pubesccns (Knight), fcnialc hololyi>c.

n — Tytthus gcminiis, hrad and pronotiim nf inalc.

C — Idcnt, acdcagus.

D, F — Idcm, Icft clas|Kr,

E — Idcm, right clas|)cr.

G — Idcm, pygophorc dorsal vicw.

FIr. II

A — Cyrtorhiniis cuinbcri, iiialc.

It — Idciii, pygophorc.

C — Iilcin, rigth claspcr.

I) — Idcm, Icft claspcr, inicnial lateral vicw.

L — Idcm^cdcagui.

F — Idnii, prctirsuü.

(• — Idcni, K itnirtiirr of frinalc l>ur»i cupulalrix.

SciELOjlo

lioL. Mi;s. Goki.di — M oino XI (II) — Dezcmlno 1955

contrur no uso desLas chaves e, por isso, deseja chamar a alC7ição

para os segjiintes pontos:

Torna-se absolutamente impossível estabelecer chaves que pos-

sam ser usadas satisjatòriamentc para todas as espécies de todos os

géneros conhecidos até o presente, tia fauna mundial. Sabemos tam-

bém que a evolttção pode estar agindo ativamente em muitas e.spé-

cies ou gêneros, de tal forma que os extremos se completam. Chaves

extremamente complexas apareceriam, se tal trabalho fósse tentado,

requerendo o estudo de todos os tipos conhecidos para milhares de

espécies existentes nos museus de vários países, tornando-se, assim,

o trabalho muito oneroso e exigindo tempo considerável .

As chaves aqui apresentadas são baseadas, principalmente, tio

estudo das espécies típicas de cada gênero e, sempre que po.ssivel, no

estudo das demais espécies que compõetti êsses géneros. O ditnor-

fismo sexual contribui também para tornar as chaves mais difíceis,

sobretudo nos casos cm que somente o macho ou a fêttiea são conheci-

dos, e são numerosas as espécies desta categoria. As chaves foram

feitas, setnpre que possível, para abranger os dois .sexos.

(hiando surgiretn dúvidas sobre se se deve seguir êste ou aquêle

ramo da chave, torna-se aconselhável, apéts atingir o fim do ramo em

questão, rever a descrição original do gênero para uma cotifirmação

mais satisfatória, tios casos duvidosos. Parece ao autor ser êste o meio

mais eficiente e certo de verificar a exatidão do trabalho. As nume-

rosas ilustrações incluídas no texto sentirão para tornar o trabalho

mais fácil e em muitos casos, elas séiziiihas indicarão a posição correta

de um gênero entre outros afins.

As dnwes foram elaboradas com o auxilio de um tnicro.scópio

binocular, usando um aumento bastante elevado (cêrea de 7(>x)\

È muito importante o conhecimento dêste fato, uma vez que certos

caracteres mencionados como forte, fundo, grosseiro etc. podem .ser

interpretados de outra maneira com o uso de pequeno aumento.

A mesma consideração deverá ser dada à iluminação, que deverá

ser forte c incidente sobre o campo estudado, finando estão sendo

estudadas estruturas delicadas como arólios, pseiidarólios, cerdas etc.

devemos experimentar fundos diferentes por baixo do inseto e deve

ser tentado qualquer meio ou mesmo instrumento que permita livre

movimentaçãio .

O uso de medidas é absolutameiite essencial ao emprêgo das

chaves. A vi.uio apenas é muitas vézes enganadora, .sendo necessário

o uso de um micrômetro ocular. As medidas são obtidas com mais

êxito, quando o inseto está sóbre fundo branco. A luz é a.ssim re-

fletida, permitindo uma definição mais clara de margem, extremi-

dade etc. (hiando determinada parte do inseto é mencionada como

mais comprida ou mais curta que outra (v.g. segundo segmento

SciELO

10 11 12 13 14 15 16

Carvai.ho — Keys Gcnera Miridae of the World

da antena, mais cojnprido que a largura da cabeça), significa que uma

variação ate 50 micra não deve ser tomada como conclusiva. Nestas

chaves, tôdas as medidas com diferença acima de 50 micra foram

consideradas conclusivas . Assim, se o segundo segmento da antena

è mais de 50 tnicra mais longo que a largura da cabeça, êle é con-

siderado 7nais longo que a largura desta última (no binocular usado,

cada divisão da ocular micrométrica media 15.5 micra).

O autor procurou usar o mais possível caracteres que são mais

comumente preservados em exemplares de museu e que possam ser

vistos exlernamente, sobretudo os encontrados na cabeça de pronoto.

Em muitos casos, todavia, isso não foi possível e caracteres como ros-

tro, segmentos da antena, pubescência etc. tiveram que ser consi-

derados.

Jletalhes estruturais superficiais, como pontuação, rugosidade e

pubescência fo7'am considerados com o inseto sob luz incidente. É

de lamentar que em certos grupos, v.g. Phylini, a pilosidade tenha

que vir a ser forçosamente considerada. São coinuns os exemplares

onde ela foi totalmente perdida e transtornada. Nesses casos, somente

um especialista ou c7itomólogo bem t7-eÍ7wdo será capaz de colocar o

gênero co7'reta/ne7ite . Pessoas com pouca expe7'iê77cia devem co7i-

S7iltar o especialista, em vez de se arriscarem a um mero palpite.

(hiando se 7nencio7ia pilosidade ou pêlos comuns, significa que

são êles os co7numcnte encontrados, seja7/i ei-etos ou recumbe7ites

(deitados) po7'ém se/npre cilhidricos, direitos e afilados para a exíre-

midnde apical. Pêlos sedosos ou lanosos são os geral/nente enrolados

ou ondulados, deitados e brilhantes sob luz mcide7ite. Pêlos esca-

7nifor/nes ou achatados são os arredo7idados ou em foiina de escama

de peixe, coiniunente tim potico alongados ou deprimidos, possuindo

côr prateada sob luz incidente. Pêlos idjos e geralmente fortes, alo7i-

gados, recebe7n o 7wmc de cerdas ou pêlos setiformes.

Nestas chaves estão incluídos lodos os gê7ieros co7ihecidos desde

I75S até P)5 l . Fora7n ta7nbêm adicionados os gêrieros descritos em

1955, seja por comunicação dos autores ou por co7isulta de trabalhos

que pudera7/i chegar às 7nãios do atitor.

cm i

SciELO

10 11 12 13 14 15 16

CONFKNrS

Pdgs.

Kcys lo llic goncra of iniridae of lhe world 11

Key lo llie suhfainilics of Miridae Ilalin, 1831 13

Key to lhe trihcs of Mirinae M

Key lo llic trihes of Orthotyiiune 15

Key lo llic tvibes of Biyocoriiiae 15

Key lo lhe Irihcs of Phyliuae Ki

Key to lhe trihes of Dnacocoiiiine Ki

Key lo lhe trihes of ('.yl/ijiiiiae 17

Key lo lhe genera of llollnioiiiiihii 17

Key lo lhe genera of Piilxnini 18

Key lo lhe genera of Cyliijiiiii 20

Key lo lhe genera of 'l'i’niialol>lii!itii 22

Key lo lhe genera of CIwiticnnni 23

Key lo lhe genera of I lyttliodini 21

Key lo lhe genera of Drrncororitii 20

Key lo lhe genera of Ihyoroiiin 20

Key lo lhe genera of MonaUmiitii 38

Key lo lhe genera of Odonirilitii 40

Key lo lhe gener.i of Pliyliiii 13

Key lo lhe genera of Dityjiliiiii ^ 58

Key to lhe genera of llidlodnliiiii 00

Key lo lhe genera of ll(dli(iiii 05

Key to lhe genera of Orlliolyliiii 08

Key lo lhe genera of Piloldioiiiii 70

Key lo lhe genera of Miriiii 82

Key lo lhe siihgenera of /.ygiis Ilalin 101

Key lo lhe genera of Mti islnrctini 102

Key lo lhe genera of Pillinnini 102

Key lo lhe genera of llyalojnldini 100

Key lo lhe genera of Ufslliniini 107

Key lo lhe geiieia of lltrdiniiini 100

Aiknowledgeinenis

Kxplanalion of figures I'’'

i\ni:\ K) (;i;\i'.KA "7

cm 1

SciELO

10 11 12 13 14 15 16

KEYS lO riIE GENERA OE jMIRIDAE OF THE WORLD

(HEMIPTEILA)

José C. M. Carvalho

^ íllscu Paraense Emílio Goeldi, Belém, Pará, Brasil

(Wilh ;;G3 figures iii lhe text)

Wlicii thc aiithor began bis studies on lhe Miridac lie was faced

witb two major difíicidties. 'Fhe first was thc doubt coiicerning

thc torrcct sytematie positions oí ncarly 150 gciiera, most of which

coidd iiot hc placcd in their jiroper trihcs of even siibíamilies be-

causc of thc inacciirate ■ dcscriptions thcn availahlc. Iii Reiiter’s

gcncric catalogue (Acta Soe. .S<i. Fciin. .S7 (.S), 1910) ncarly 100

gciicra wcre regarded as of uneertain position, and scveral othcrs

allottcd positions on thc hasis of their dcscriptions only.

Unforlunatcly this master laxonomist was not ahlc to examine

lhe types of tlicse genera, scatlercd as tliey wcre among thc muscums

of Eiiro|)c and America.

Tliis major difficully scems to he removed now with thc jju-

hlication hy lhe aulhor of lhe paper cntitled; “On thc Major

Classification of thc Miridac etc.” (An. Acad. llrasil. Ci. 24 (l):

•11-110, 1952) whicli ha.sed on the stndy of types in seycral muscums

of Europe and America, lists ail genera nnder tiicir correct suh-

familics and trilres, tinis allowing entomologisls to work up sa-

lisfactorily tlicse two lirst steps in tlie taxonomy helow thc family

levei .

'I iie sccond major difficidty was lhe lack of apjrropriate keys,

witii illustration of criticai characters, to thc genera or even groups

of genera which wonld emhrace thc w'orld latina. lhe identific-

ai ion of a cerlain gentis was usually dclayed considcrahly hy having

lo (onsuli a great immher ol pajrers, and check the litcratuic ex-

hausiively. On lhe other liand lhe study of regional keys only was

not salislactory siiice in niany cases it was douhtlul whether a

geinis had heen introdnced recentiy or not into thc region.

The preseni paper has heen" wrillen in order lo remove ihis

second difficulty. The author is aware of the dilliculties which his

(olleagues niay enconiiter in nsing lhe acconipanying keys, and

aileiilion is called to the lollowing iroints.

cm 1

SciELO

10 11 12 13 14 15 16

12 Mus. Goki.di — loiiio XI (II) — l)c/cmbro 1955

It .scems almost iinjx)ssiblc to cstablish kcys to work satisfacto-

rily for all .spccics of all gciicra kiiowii at thc prcscnt tinic tbroug-

hoiit the world. Wc kiiow tliat thcrc arc niurh gradation in naturc,

and that cvoliition may bc on thc inarch in ccrtain spccics or gcncra

.so that thcy niay incrgc into onc anothcr so that thc extremes come

in contact. A very compicx kcy would rcsidt if such a work wcre

aUeinjjted, and thc time recjuired to stiidy all the types roncerned

in mnseums in many conntries woidd be almost probibitive.

riic prcscnt kcys wcre baseei mainly on the types spccics of

thc gcncra and, as miich as possiblc on thc study of othcr s])ccics

inclndcd in thc gcncra. .Sexual dimorphism tends aiso to render

tlic kcys more difficnit, csjKxially as oídy thc fcmale or malc sex

is known for a great nnmbcr of s|)ccics. l he kcys wcre made when-

ever possiblc to inchidc both sexes. Whcn a doubt ariscs whcthcr

to follow this or that branch of thc kcy, it is adviscd that after

rcaching thc end of thc branch conccrncd a chcck shonld bc made

with lhe original dcscription of thc gemis in order to ascertain

if onc is right or not. 'I his .seems to bc lhe safest way to do such

work. lhe mnncrous ilhistrations anncxcd to thc text will render

lhe work mnch casicr and in many ca.scs thcy alonc will point out

a ccrtain gentis among their rclativcs.

'l he kcys wcre construeted with thc hclp of a binocular micros-

copc, using a fairly high magnification (about 70 x). 'í bis is very

importam since ccrtain characters mentioned as strong, dccp, coarse

etc., may bc interpreted othcrwisc if low magnification is used.

'lhe same consideration ajiplics to thc illumination, which nuist

bc strotig and incident ovei thc inscct. Whcn dclicatc struetures

arc to bc sccn, sucb as thc arolia, jiscudarolia, sctac etc., diffcrctit

backgrounds shonld bc tricd under thc inscct, and any system per-

mittíng thc lattcr to bc turncd around or u|j and down is cu-

cou ragccl.

Measurements arc absolutcly csscntial to dcal with thc kcys.

'l he cyc alonc is sometimes mislcading and a micrometrie cycpiccc

niust bc used. Metisurements arc better takcn witb a white back-

groiind rcflcctitig thc light which will pcrniit a better cicfinition

of margiti or cxlrcmitics. Wbcn a ccriaiti part of tbc inscct is said

to be longcr or shorter lhan anothcr, such as thc second antcnnal

segment longcr or shorter thati thc hcad it means that a variatioti

uj) to 50 microns mnst not be regarded as conchisive. In thc prcscnt

kcys all meastirctncnts above 50 microtis wcre considered as con-

clusivc, ibus if tbc .second tintennal segment is more than 50 microns

as long as thc width of hc';ul, it is considered longcr than thc heaci

(in thc binocular microscoj)c used thc micrometrie cyepiece mea-

siired 15.5 microns for each division).

cm i

SciELO

10 11 12 13 14 15

Cíarvai.iio — Kcys Gciicra Miridac ol llic World

'riic aiithor has tricd as íar as jjossiblc to use eharacters wliich

are to be íouiul iii imiseimi sjtecinieiis and are visiltle externally

siieli as titose ol the head pronotiim. In inany cases however this

was not jtossilde and eharacters snch as rostrinn, antennal segnients,

jtnljescence etc., had to be considered.

vStrnctnral details ol the npper snrlace, snch as pnnctnration,

rngosities and the pnbescence were delined with the insect nnder

incidem light. It is nnlortnnate that in certain gronjts, snch as the

IMiylini, the ptibescence ninst be cotisidered, since in tnany cases

these hairs aie easily dislodged ;tnd lost. In stich cases only tlie

specialist ot a well trained entoinologist will be able to ]ilace the

genns coirectly. lhe inexjterienced jterson shonltl consnlt a spe-

ciitlist rather tlian risk a ntere gness.

When coinnion pnhescence or hairs ttre nientioned it tneatis

that they ;ire tlie nsnal ones lonnd, either erect or rectnnbeiit (atl-

])ressed), Itttt always cylindrical, straight, atid ta])ering towards tlie

ajticttl extreinity. .Silky or woolly hairs are soinewhat cnrled, nsnally

rectmibent atui brilliant with incident light. .Scale like or llattenetl

hairs are llattcned or ronnded with typical silvery colonr nnder

incident light. .Still and nsnally stroiig, long hairs are called bristles

or setilonii htlirs.

So lar as the anthor is awaie these keys inclnde all genera

described niilil the end ol 1951 and also a lew described in 1955.

KKY ']'() riIK .SUllFAMIl.lE.S OF MllUDAE IlAllN, 1831

Arolia ])resent, large and Iree, arising between the claws

(ligs. 19, 20) 2

Arolia absent, snbstitnted hy a [tair ol straight hairs (ligs. 2,

7, l<>) •■•••• ^

Arolia distinctly divergem toward lheir ápices (lig. 20) nsnally

dilated; |)i()notal collar always present and well sejtarated Irom

])ronotnin by a Itirrow (ligs. 31, 38)

M/n/iVAE llahn, 1831 pg.

.Arolia parallel or (onvergent towtntl their ápices (lig. 19),

nsnally slender; pronotal collar il ])resent, ol the tlepiessed

lype (ligs. 32, 31, 30), not se])arated Irom pronotnm by a

liirrow onriIüTYLINAE Van l)n/ee, 1910 pg. 15

r.scndaiolia ])re.sem, Iree or connected with the claw, sonietimes

ininnie and dillicnlt to see (ligs. 510) in this case the pro-

notal (ollar absent (lig. 33)

Psendasolia absent (ligs. I, 2, 3, 1): i)sonot;il collar ))resem

or in case not. the claws very long, smooth and slemlcT

(IÍMS. 1, 2) ^

lioi,. Mus. (loKi.Di — 'rf)ni() XI (II) — Dcvcmbro 1955

Psciularolia arising from tlie base or iiiiier niargin oí claw

(ligs. 5-I()); incinbranc with two cells (íig. 22); tarsi linear

(fig. 29) PHYLINAE Douglas & Seott, 191)5 pg. Ki

— Pseiidarolia arising írom tbe veiUral suurlace ol elaw (figs. 17,

18); ineinbrane witli one cell (fig. 21); tarsi tnickenecl toward

ápices (fig. 28) lillYCOR/NAK llaerensprung, 10r)O, pg. 15

Clavvs toothed or tickened at base (figs. 3, 4)

DERAEOCORINAE Douglas k .Scott, 10(iÔ, pg. Ib

— Claws sniooth at base, long and slender (figs. 1,2)

CYLAPINAE Kirkaldy, 1903 pg. 17

KLY TO THE d RI15ES OF MIRINAE

1 . Idrst seginent of bind tarsi as long as or longer tlian secontl

and tbird togetiier (fig. 23) or when tliis is not tlie case, pro-

notal collar incoinplcte or pronoluin with a lateial riilge at

least anteriorly 2

— F'irst seginent of nind tarsi not as long as second and tliiicl to-

getlier of if so, jnonotinn witiiont a lateral ridge and jnonotal

collar distinct, se|5arated froin disc by a furrow (fig. 35) ... 1

2. F'irst segnient of antennae as long as head aiul pronotum toge-

ther (fig. 44); pronotal collar distinct and complete; legs and

antennae very long MECASTOSC.EEIN/ Renter, 1910 pg. 102

— F'irst segnient of antetniae sliorter tbati lictid and pronoluin to-

gctlier; pronotal collar if [iresent usually inroni])lete; legs and

antennae not noticeably long 3

3. Myrniecoinopliic species, iisutdly with elytra not divided, the

ctineus anil ineinbrane vestigial ar absent

PITIÍANINI Douglas St -Scott, 1905 pg. 102

— Species not inyrniecoinor|)hic, the heinielytra diviiled into co-

riuin, < lavus and emboliuin, the cuneus and ineinbrane present

(TrENODEMfNI China, 19.34 pg. 103

4. Myrniecoinor|)hic speiies with the abdômen constricted at base

(fig. 39); collar usually represented by a depressed line

IIERDONftNI Distant, 1901 jig. 109

— Species not myrincconiorphic, the abilomcti not constricted at

base; collar distinct, separated from jironotuni by a funow 5

5. Ostiolar jieiiticme small (fig. 25), its dorsal niaigin scariely

extenditig dorsal as far as ventral margin of iiic.scpimeiini; pro-

notal collar very wide, with mesal Icnght usually as great as

width of lalli (fig. 47); ilull black species with reddish, luteous

or yellow maiks RIÍSTIIENINI Reuter, 1905 pg. 107

SciELO

10 11 12 13 14 15 16

Carvalho — Kcys (icncra Miriclae of thc World

— Ostiolar pcritrcmcp roniinent (fig. 24), its dorsal margin cx-

tcnding wcll above vcntral margin of nicscpimeron; ]n-onotal

collar (fig. 45) not as broad as width of calli; specics if dark,

nsually shining (>

(). 1 Icinic-lytra glassy and transparcnt allowing thc abdômen and

mcmbranons wings to be seen froni above

HYALOPEPUNI Cífi valho, 1952 pg. lOC.

— 1 lemfelyira not glassy and transjiarent, the abdômen an mem-

branons wings not seen from above MIRINI Hahn, 1831 pg. 82

ki:y ro ihe iribes oe orthotylinae

1 . Small, nsually dark compact species with saltatorial femora, the

genae very high (fig. 43), equal to or more than height of onc

eye; vertex very wide, eycs i)rominent: third antennal segment

nsually nnich more slender than second; body frequently with

scale like i)nbescence; brachypterons fornis very common ....

HALriCINI Kirkaldy, 1902 jtg. ()5

— Species of médium size, nsually greenish of light coloured,

without saltatorial femora; the genae low (fig. 40) equal to

or less than the height of one eye; vertex if wide then eyes not

prominent; third antennal segment frequently equal to thick-

ness of second: if body with scale like pubescence then both

sexes macropterous 2

2. Myrmecomorjihic species with abdômen constricted at base ...

PILOPHORINI Renter, 1883 pg. 79

— Species not myrmecomrphic, the abdômen not constricted at

base ORTHOTYLINI Van Duzee, 1910 pg. 68

KEY ro rilE I RIBES OE RRYOCORINAE

1 . Eirst antennal .segment incrassate, equal in lenght to half lhe

width of vertex (fig. 27) about as long as wide; species nsually

of large size, with coarsely, punctate pronotum and strongly

iullated sculelium, if first antennae longer than half lhe width

— of vertex and scutellum no cystiform then thc membranc with

auxiliary veins or head with three poinied tuberclcs anterioily

ODONIELl.INI Renter, 1910 pg. 40

Eirts aulcunal segment if not longer than half thc width ol

vertex, then distinclly narrowcr than long, thc pronotum smoolh

and shining: sculelium never inflated or cystiform 2

2. Earge, long aml slender species with smoolh and shining boily;

j)ronoinm strongly constricted anterioily, thc head with a dis-

SciELO

10 11 12 13 14 15 16

Hoi,. Mus. CioK.i.Di — 'loino XI (II) — De/cnihio 195.5

tinct iieck (lig. 30); rostnini rcacluiig apcx ol anterior co.xac

oi' so; secoiul antcnnal segineiu about llirec limes (ir more as

loiig as lirts MONAl.ONÍINI Reuter, 1892 pg. 38

— Meilimii si/.e to small species; pronotiim usiially ])imetiiretl or

if smooth, not coiistricted anteriorly or lhe rostrum longer,

reaching beyoiul the apex ol anterior coxae; head withont a

distinet neck ol il present, the second antennal segment less

than three times as long as lirst

ISRYOCORINI IJaerenspning, 1880 pg. 29

KF.Y C)F rilE I RIllKS OI' RHYLINAE

1. 1’ronotmn withont an a|)ical collar (ligs. 33-37)

RIIYJJNI Douglas 8: Seott, 1885 pg. 13

— 1’ronotum with a well marked aiiieal collar or when this is not

the case, species with anl-like a])peatance 2

2. Myrmecomorphic species, with ahdomen constricted at hase;

pronotum nearly tritingnlar or elotigtite, witli a more or less

llattetied tipical colhir separated Ironi disc hy a sligltt Inriow;

claws ustially long and sletider, not henl at hase; hemielytra in

niost cases with a wliite or yellow tioss hatui or with pale areas

IIAl.LODARINI Van Dnzee, 19l(i j)g. (iO

— .Species withont myrmecomorjjhic appeatance or il so, then ah-

domen tiot constricted at hase, protiotimi with a well inaiked

collar (lig. 12); claws il long and sletider notiteahiy hent at

hase; hemielytra withont a wnitisn or yellow cross hand or |)ale

aieas I)/CYI‘U/NI Reuter, 1883 pg. 58

KEY IO 11 lE I RIISE.S OE DERAEOCORINAE

1. I lead elongate, pointed, slightly shoi ter than pronotum, lions

hoti/.ontal or neaily so; eyes tistially large; ;intetniae very shoit

(lig. '18); species ol small si/e

TERMATOrUYEINI Reuter, 1881 pg. 22

— I lead vertical or sttotigly declivotts, intit h shorter than jtiono-

tnm; aitlennae not tioticeahly shoil; species ol inedínm or laige

si/e • 2

2. Ihonotnm with an impic.ssed line iimning liom tinteio latetal

(otiier to postei ior margin ol calli (lig. 28)

CEIEINEMINI Retiter, 1875 pg. 23

— Pronotum withotit the line mentioned ahove 3

cm i

SciELO

10 11 12 13 14 15

Carvai.ho — Kcys Gcnera Miriclae of thc World

llciiiiclytra liyaline, transpareiu and glassy, cmboliar margin oC

corium grcatly cnlarged

í lYALIODINI Carvallio & Drakc, 1913 pg. 24

Meniicdytra not hyalinc, glassy or transparcnt 4

Pronotum constrictcd antcriorly, thc calli large and fuscd; eyes

scnii-stylatc; nicinbrane with one cell

SArURNlOMIRINI Carvalho, 1952 pg. 29

1’ronotnm not constrictcd antcriorly; calli not prominente and

liiscd, ncithcr arc thc cycs scini-stylate; membrane nsually with

tvvo cclls . . DERAEOCORINJ Douglas 8: Scott, 1865 pg. 26

KKY rO 4 HE I RlllES OE CYLAVINAE

Hcad long and ijointcd, gida long, frons horizontal or nearly

so, clipens distinctly, curved, its ajjcx nsually vcntral of its base;

calli very large, confluent, occupying thc anterior two thirds

ol' pronotum (ligs. 41, 51, 53) EULVIINI Uhlcr, 1886 pg. 18

Ilcad short and roundcd, gula short, 1'ronis vertical or strongly

tlcclivous (lig. 48), clyjxms in thc same plane as frons; calli i£

large not occupying the two anterior thirds of pronotum

(‘ig- 18) 2

Jíody strongly siiining and coarsely punctate, thc size of the

puncturc about etpial thickness of first antcnnal segment at

l)ase; osliolar peritreme with an ocelloid shining tuberclc; mem-

brane distinctly pilose

JiOTIIRIÜMlR/N/ Kinkaidy, 1906 pg. 17

Ilody more finely j)unctatc; ostiolar ])eritreme without an

ocelloid shining tid)ercle; membrane glabrous or if pilose, very

minutely so CYLARINI Kirkaidy, 1903 pg. 20

KEV ro rilE CENERA OE ROTIIRIOM/RIN/

Pronotum with tubercular shining sweilings; scutellum with

two latertd high lobes sej)arated by a deep sidcus (Sumatra) . .

LEEROCAESUS Poppius, 1914

Pionotum without tubercidar sweilings; .scutellum not as

above 2

Second aniennal segment four times an long as the first, strongly

thickened, with short hairs anti two long, erect bristies; rostrum

readiing thi' middie coxae (Índia)

DASHYMENEi Po])pius, 1910

18 IloL. Mus. Goei.di — loiiu) Xl (II) — Dezembro 1955

— Secoiul antennal segmciit aboiu twice as long as tlic lirst, not

noticeably tiiickenecl; rostium reaching tbc anterior eoxac or

slightly beyoncl it .8

3. Scutelhmi with a meclian, apical and smooth lobe (fava) ....

DASIIYMENIELLA Poppins, 1914

— Ssutellum witliout a median ajiical lobe 4

4. First antennal segment as long as or longer than width ol vertex;

second segment twice as long as tbe first (índia, Malay, llorneo,

Formosa) BOTTIRJOMIRIS Kirkaldy, 1902

— First antennal segment shorter than width of vertex; second

segment 2-1/3 as long as tbe first (Prilippines)

BAKER! O LA Ilergrotb, 1920

KEY 40 FIIE GENERA OF EULVIINI

1 . Pronotal collar absent or obsenred by tbe calli 2

— Pronotal collar present 3

2. Tarsi two segmented; body shagrecned, oval or rounded in

outline (fig. 232) (África, Australia, índia, Américas)

PERlTROriS llbler, 1891

— 'Farsi tbree segmented: body not sbagreened, elongate in ont-

line (Pbilippines) EULVIDIUH Popj)ins, 1909

3. Pnbescence on eyes longer than diameter of ommadium; cuneus

very narrow, not as broad as tbe embolinm or absent .... 4

— Pnbescence on eyes absent or if present not longer than dia-

meter of ommatidinm; cuneus if jjresent, wider than embo-

lium 5

4. Lateral margins of proiiotum strongly carinate; cuneus absent

(Australia) LYGAEOSCYTUS Reuter. 1893

— Lateral margins of |)ronotum not carinate; cuneus present, very

narrow (África)

IIEMlüPIirilALMOCORIS Poppins, 1912

5. Eyes reaching tbe gula below in lateral view (fig. 53) .... 0

— Eyes not reaching the gida below in lateral view (fig. 51) . . 13

0. llody oval; ))osterior angles of pronotum not prodiued and tbe

lateral margins straiglu or convex; head and iHonotum greenisb

metailie, puiictured (New Guinea)

BIRONfELI.A Poi)pius, 1909

— ISody elongate, sometimes slightly widened hiteially; posterior

angles of pronotum produted, the later:d margins slightly

emarginate 7

CaRVAI.IK)

Kcys Cíciicra Miridae of tlie W^orlcl

7. Ilciniclytra witlioiit clistinct cunciis and cniljoliiim, tlie coiium

dividcd into ccto, incso and endocoriinn (fig. 75) (Central &

Soiitli America) XENOCYLAFUS liergToth, 1922

— Hemielytra witli a distinct cunens and emboliinn 8

8. Einbolinm strongly widened after basal third; antennae in-

serted lar 1'rom tlie anterior margin ol thc eyes 9

— Einboliinn ol' abont the saine width thronghont, not noticeably

widened alter thc basal tliird; antennae inserted contiguous

to tlie eyes 10

9. Rostrnm reaching the middle coxae; emboliinn narrowed at

llie apex (Ceylon) LEFIDOhULVIUS Poppins, 1913

— Rostnnn reacbing tiie base ol abdômen; cmboliiim not nar-

rowed at the apex (Mcntawei L)

EUCEIILOEULVIUS Poppins, 1909

10. Rostnnn reacbing lhe middle coxae (México)

ORASUS Distant, 1883

— Rostnnn reacliing beyond tbc posterior coxae H

1 1 . Eirst antennal segment reaching beyond ajiex ol bead (Cosmo-

politan) EULVIUS Stal, 1802

— First antennal .segment not reaching beyond apex ol head 12

12. Rostnnn very long, reaching ajicx ol abdômen (Atrica) ....

MICROEULVIUS Poppins, 1912

— Rostrnm reaching only thc middle ol abdômen (Colombia,

Panama) EERITROPOJ DES Carvalho, 1955

13. Pxuly smooth or shagrecned, not pnnctnred H

— boily ;ibove distinctly pnnctnred 10

11. First segment ol rostrnm reaching thc lirst coxae; antennae

very long; cnnens indistinct l'^

— First segment ol rostrnm not reaching beyond thc base ol head;

thc antennae not very long; cnnens nsnally distinct (at least

on m;u hopteions lorms)

15. Antenna very long; cnnens indistinct; spccics ol large size K)

— .\nlenna not' very long; cnnens distinct; spccics ol .small size

(IJR.\/IL) PAIIAFUÍA lUS Carvalho, I9 j4

lli. Ileinielytra with spar.se, tnbercnlar swellings; rostrnm reaching

lhe apex ol abdômen (Malay, Philippines)

RII/NOMIRIS kirkaldy, 1902

— Ileinielytra withont tnbercnlar swellings; rostrnm not reaching

beyond middle ol abilonien (AIrica) ’ i/wU

JilIlNOMIRIDIUS Po|)j)ins, 1909

Bol. Mus. Goki.di — 'lomo XI (II) — Dc/.cniljio 11)55

11 ).

20 .

17. First antennal scgmcnt linear; first rostral segincnl rcadiing

only thc middie of llie cycs towards thc apex (África)

lUIINOlULVIUS Reuter, l'J02

— First antennal seginent incrassate; first rostral segincnt as long

as thc hcad 18

18. Species with aspect of beetle; brachypterous; calli and scntellum

strongly raised (Jamaica)

ISRACIIYFULVJUS Carvalho, 11)55

— Spccics without aspcct of bcctlc; niacroptcrons; calli and scutcl-

hnn not strongly raised (Australia)

CERATOFULVIUS Rcntcr, 11)02

Frons depressed, striolatcd and pnnetate; thc body strongly

punctatc, tunens absent (fava, Snmatra)

TERATOFULVIUS Poppius, 11)11

Frons smooih or snleate, withont punctures 20

Ilcmiclytra with tnbcrcular swcllings, vertex protrnding np-

wartls with two convex tubcrclcs (Phili|)pincs)

LIINDHLADIOLLA n.gcn.

typc; Psicolrranphus albomaculnlxis .Stiil.

— Ilcinielytra without tubcrcular swcllings; vertex not as

abo ve 21

21 . Cuneus absent; first antennal segnient shorter than width of

liead, with two or three long setae; small, compact species

NEw Gninea) CYI.AFOFUIA íUS Poppins, 11)01)

— Gnnens present; first antennal segment as long as or longer

than width of hcad, withont setae; median size species ... 22

22. Ilody glabrons; cnnens shorter than broad at base; lostrnm

reaching the middie of abdômen (llorneo)

RUINOCYLARUS Poppins, 11)01)

— Body with sparse, yellow, adjjressed pid)escence on hemielytia;

cnnens abonl as long as wide at base; rosinnn leaching the

airex of the abdômen or nearly so (Foiinosa)

RUlNOCYl.AriDIUS Poppins, 11)15

KFV FO I IIF, GFNF.RA OF CYl.AFINI

1 . Body above smooth, rngose or shagreened

— Body above, at least on ])ir)notnm, distinctiy pnnctnred

.‘ 1 .

Body with very shoit, ad|)ressed pnbesceiue

Body with erect or semierect |)nbescencc . .

frons with a pointed piocess

SciELO

10 11 12 13 14 15

Carvai.iio — Keys Gcncia Miridac of the World

10 .

II.

Froii.s without a poiiitcd process (If fions is prodiiccd then

bhmt or sulcate) 5

Sditellum Hat; clypcus coinpressed; anterior íemora incrassate

(Bornco) RHINOPHRUS Hsiao, 19M

.Sciitclluni witli a mcdiiim tubercle; clypcus not compressed;

anterior femora not incrassate (Ceylon)

CINNAMUS Distant, 1909

Anterior tibiae strongly compressed, íoliaceus (Ceylon) ....

PHYLLOCYLAFUS Poppius, 1913

Anterior tibiae not comjrrcssed and íoliaceus ü

First antennal segnicnt very short and thick, about as long

as balí the widtli oí vertex; hcad vertical (Philippines, Palau)

CYLAPOMORPHA Poppius, 1911

F'irst antennal segment long slcndcr, about as long as or longer

tban width oí vertex; liead inclined (Philippines, Koror) . .

MYCA:T()CA’LAPUS Poppius, 19M

Frons protuding in front, dccply sulcate (Hg. 244) 8

Frons if jrrotruding as above not sulcate 9

First antennal segment as long as the bead, ibe second three

times longer tban tbe íirst (íig. 244) (Mailagascar, Américas,

New Cuinea) VANNÍUS Distant, 1883

First antennal .segment as long as tbe bead and pronotum

togetber, tbe second .segment only 1-1/3 as long as tbe íirst

(New llebridcs, Daupbin, Esp. .Santo)

VANNIOPS/S Poppiu.s, 1909

Posterior femora noticeably cnlarged towards tbe base; second

antennal segment about 7 times as long as íirst segment (íig.

24.')) (Madagascar) PARACYLAPUS Carvalho, 19.')2

Posterior femora not noticeably enlarged towards tbe base;

second antennal .segment less tban 4 times as long as íirst

segment 10

Hody witb semierect sbort |)ubescence; ícmales bracbypterous

(Iba/.il) CORCOVADOCOLA Carvalbo, 1948

Hoily witb long anil erecí pidrescence; botb sexes macro-

pterous 11

Frons sulcate, eyes very large and sbortly pednnculatc; bead

as wide as pronotum at base; rostrum rcacbing bind coxae

(15,a/il) CALAPOIDKS Carvalbo, 19.’)2

Frons smootb, eyes not pedunculate; bead narrowcr tban pro-

noium at base; rostrum longer (Philippines, New Cuinea) . .

TRICIIOPULPIVS Poppius, 1909

SciELO

10 11 12 13 14 15

Mus. CioKi.Di — Toino XI (II) — Dc/cmhio 1955

12. Anterior Icmora strongly enlarged; tiinciis alrseiit or long as

wide as base (índia) PliOAMBLlA Ilergroth, 1910

— Anterior íeniora not noticcably enlarged l.'i

13. Eycs rising a considerable distante above dorsimi of head wbicb

is very deeply snleatc (Américas) CYLAPUS .Say, 1832

— Eyes not rising a consideraljle distance above dorsimi ot head

wliich is not deejjly sidcate M

H. 1’ubesccnce erect and very fine; rostrnm reaehing genital seg-

nient; cmboliinn wide, laniinate, tlaval, eorial and eml)oIial

veins witii a row' of punctures (Amazônia)

CYLAPOCOlilS Carvallio, 1951

— 1’ubescence not as above or if so tlien emljolium narrow, eorial,

tlaval and embolial veins witliont a row of punctures (New

Caledónia) PALISCUS DistanI, 1901

KEV IO EIIE CENERA OE TPAiMATOPllYLINl

I . Eirst antennal segment reaehing apex of head; seeond segment

strongly enlarged, foliaeeus (fig. 76) (Nortli America)

IIESPEROPUYI.UM Reuter & Poppius, 1912

— Eirst antennal segmnet reaehing to or beyond apex of head;

seeond segment if inerassate, not foliaeeus or flaitened (fig. 71) 2

2. llemielytra trans])arent; eycs distant from pronotum, witb long

pubeseenee (fig. 21) (Central & South America)

TEliMATüPHYLlDEA Reuter & Poppius, 1912

— llemielytra not transjjarcnt; eyes eontiguous witb pronotum or

nearly so, not or only sliortly pubescent 3

3. Ilead dislinetiy longer tlian wide •!

— I lead wider than long or as wide as long 5

'l. llody witb seale like pubeseenee in rows; eyes glabrous (Egy|)t)

A l{(iY ROPELAENUS Reuter & Poppius, 1919

— P.ody witbout .seale like pube.seenee; eyes pubeseeut (Nortb

America) CONOCEPI lAEOCORLS Kuigbt, 1927

5. Ilead almost twiee as wide as long; meinbrane (oriaeeus (Cen-

tral ílc .Sotitb America)

TERMATOPIIYLELEA Carvalho, 19.55

— Head as long as wide oi neat ly so; membratie not (oriaeeus 6

6. Pronotum witb a row of punctures on sideus bebind and

between (alli; pidxseeiue very long and etect; rostrum readiing

apex ol anteiioi (oxae (Cential Ameiiea)

TERMATOPUYEOIDES Carvalho, 19.55

Carvai.iio — Reys Gcncra Miridae o£ tlic World

— Pronotimi without the row of punctiircs as above; pubescence

not noticcably long; lostrum reaching ihe middle coxae or

bc-yond (fig. 46) (África, Índia, Malay, liorneo)

TERMATOPHYLUM Reuter & Poppius, 1912

KEY lü THE GENERA OF CLIVINEMINI

1 . Large sj)ccies with sevcral short spurious veins arising from

large cell (fig. 67) (Central America)

MAGAMIIUS Hsiao, 1947

— Mcinbranc withoiit spnrious veins on membrane 2

— Anterior inargin of pronotnin if cystfiorm, not hooded or pro-

jccting over the head (fig. 62) 3

— .Vntei ior inaigin of jnonotnm if cystiforin, not hooded or pro-

jccting over the head (figs. 60, 63) 5

3. llody with short, adpresscd pubescence; cnneus twice or more

as long as witle at base (Central & South America)

OFELLUS Distant, 1883

— lloily with erect or semicrect pubescence; cnneus less thau

twice as long as wide at base 4

4. Hairs of body strongly curled somewhat flattened at middle,

very dense; frons itointcd (Central S; North America)

CU V IN EM A Reuter, 1875

— Hairs normal, not curled; frons not jiroduced (Central & South

America) AMIUUCIUS Stal, 1860

5. Prouotum distinctly carinate between the calli aml also in the

middle of collar (Central America) . . . ZOILUS Distant, 1884

— Pronotum not carinate between the calli and the middle of

collar 6

6. Lateral margins of pronotum distinctly carinate 7

— Lateral margins of pronotum not carinate 8

7. Setoiul antennal segment stout and clavate (North America)

LAliC.IDEA Van üu/.ee, 1912

— Second antennal segment linear (Panama)

8. llody smooth, 'shining and glal)rous (Jamaica)

LAMPllOSCYTVS Reuter, 1907

— llody pilose, pronotum distinctly punctate 9

9. Mcmlnane distinctly pilose (Europe, Asia Ib North America)

liüTlIYNOTUS Fiebcr, 1864

Memljiane glabrons . . . .

24 Kol. iMus. Goei.i)i — 4'oino XI (II) — Dezembro 1955

10. Seconcl antennal segmcnt threc times longer than first; body

punctatc only on pronotum, thc rest smooth and shining (íigs.

00, 63) (Central & South America)

GUANABAREA Carvalho, 1918

— Second antennal segment approximately five times as long as

first; body rugously pnnctate (West Indies)

HEMICEROCORIS Lethierry, 1881

KEY OF THE CENERA OF HYALIODINI

1 . Scutellum with a median stout, suberect sj)ine-like projection

(lig. 25) (Sonth America)

KNIGHTONÍA Carvalho & Drake, 1944

— Scutellum smooth, without a spine-likc projection 2

2. First and sccond antennal seginents very wide, laminate or

foliaceus (fig. 58) (Central & South America)

AUCflUS Distant, 1893

— First and second antennal segments cylindrical; if incrassate,

never foliaceus 3

3. Pronotum strongly constricted on ajjical half, (figs. 56, 61)

this j)ortion being much narrover and as long as or longer

than the head; embolium slightly wider than length of first

antennal segment 4

— Pronotum not as abovc or of constricted anteriorly then end^o-

lium less wide than the length of the first antennal segment 5

4. Rostrum reaching apex of anterior coxae; eyes not contiguous

with anterior margin of pronotum (fig. 61) (Central America)

TRYGO Distant, 1884

— Rostrum reaching to or beyond middle coxae; eyes contiguous

with anterior margin of pronotum (fig. 56) (South America)

CARIJOANUS Carvalho, 1955

5. Eyes distinctly separaled from jnonotal collar this distance

being about 1/3 or more length of one eye (figs. 65, 68, 257) 6

— Eyes contiguous with jnonotal collar or nearly so (figs. 57,

59, 74) 10

6 . Eyes large, occujjying most of the sides of the head as seen from

dorsal asjiect (fig. 65), the distance between eye and collar

etjual to 1 /3 or less the length of eye 7

— Eyes ^lot noticeably large,occuj)ying only anterior jjortion of

head (fig. 68), the distance between eye and collar etjual about

length of eye 9

Carvalho — Kcys Gencra Aíiridae o£ thc World

7. Sccond aiitennal seginent incrassate towards thc apex; pro-

notuni strongly constrictcd anteriorly, long and erectly pilosc;

incsoscutuin broadly exposed (Haiti)

FENNAHIELLA Carvalho, 1955

— Sccond antcnnal scgmcnt not incrassate towards apex or if so

then jjronotnm not strongly constrictcd anteriorly and body

ahnost glabrous 8

8. Hcad about 2.5 times wider than long; íirst antcnnal seginent

1 .5 times or more longcr than length ol' hcad; species usually

over 4 mm. long (fig. 65) (South America)

HYALIODOCORIS Knight, 1943

— Head only two times wider than long or less; first antcnnal

.scgmcnt usually less than 1.5 times longer as head; species

usually less than 4 mm. long (West Indics & C. America)

PARACARNUS Distant, 1884

9. Pronotum very coarsely j)unctate, with tubcrcular shining

sivellings; first antcnnal scgmcnt slightly longcr than head;

rostrum reaching thc posterior coxae (Ecuador)

LYDE Distant, 1893

— Pronotum without shining tubcrcular swellings (excepting ca-

lina); first antennal scgmcnt usually distinctly longer than

hcad; rostrum reaching the middlc coxae (North, Central &

S. America) HY ALIO DES Reuter, 1876

10. Head strongly pointed in front (fig. 253); first antcnnal seg-

ment as long as width of onc cye seen from above (Central

America) I' USCUS Distant, 1884

— Head rounded in front; first antcnnal segment longer than

width of cye seen from above 1 1

1 1 . First antennal segment three times as long as length of head;

sccond antcnnal segment as long as íirst; rostrum reaching thc

posterior coxae (fig. 52) (Central & South America)

« ANNONA Distant, 1884

— First antennal segment less than three times as long as lengtli

of hcad; sccond antennal segment longer than first segment 12

12. Pronotum strongly convex and declivous towards the head and

margins; first antennal scgmcnt as long as width of vertex;

clavus usually brack with an ocellatc white spot (fig. 74)

(Central &: South America) FLORUS Distant, 1884

— Pronotum not noticeably convex of if so, then first antennal

scgmcnt longer than width of vertex and clavus not as above 1 3

13. First antennal segment incrassate, usually distinctly longer than

hcad; species usually over 4.5 mm. long (figs. 59, 64) (Central

& Soutli America) PSEUDOCARNUS Distant, 1884

I$<)L. ^íu.s. CioKi.Di — loino XI (II) — Dc/.cnibio 1955

— First aiitcniial scgincnt slcmlcr and aboiit as long as head;

si>ccies usually Icss than -1.5 min. long (lig. 57) (Central Sc

Sauth America) ANTiAS Distant, 1884

K1:Y to 1 HF CiKNERA OF DERAEOCOIUNI

1 . .Sccond antennal segincnt broad aiul distinctly ílattened

(í'g- 76) ;.■■■■;„

— Second antennal seginent linear or clavate (fig. 07) .... 2

2. Calli strongly raiscd and pointed at antero lateral anglc; cyes

set at iniddle ol head; einbolium very wide (Pern, 'Frinidad)

EKRUANOCOIilS Carvalho, 195.‘5

— Calli ií convex not as above; eyes not set at iniddlc of head

bnt if so then einbolinm not noticeably wide

.'l. First antennal segment very short and thick, strongly narrowed

basally, aboiit as long as widtli of vertex; lirst tarsal seginen

innch tliicker than tlie others with a tnft of hairs inferiorly;

antennal peduncle very large (fig. 243) (Madagascar)

PAUL! AN AN A Carvallio, 1952

— First antennal segment not as ahove; first .segment of tarsi if

tliick, then withont the tnlf of liairs; antennal j)eduncle not

very large 4

4. Frons pnnetate (Fgypt) .... CRANOCAPSUS Wagner, 1951

— Frons sniooth 5

5. Frons transverscly striate, the vertex snleate (fig. 54) .... 6

— Frons more or less polislied, scarcely striate, vertex not sidcatc 7

0. .Second antennal segment clavate; tliird antl fonrth shoit and

thick, fnsiform (Nortli ík .Sontii America)

DIPÍ.O/.ONA Van l)n/ee, 1915

— Antcnnae linear oi neai ly etpial thickness throngliont; second

joint scarcely eidarged at apex, tliird and loiirth linear (Norili

)k- .South America) EUSTÍCEUS Renter, 1909

7. Ilcmielytra smooth, somewhat tianshicent, cnneiis strongly in-

(lined; body ttstially shining, clavo (orial and endtolio corial

(ommissiire at base with a low ol pnnt tines 8

— Ilemielytra i)imtlate 9

8. IJody glabrons, ciineal frattnre not notiieably witle anil ileep;

hemielytra translncent (Cenltal Ameiiia)

CARMELUS Distant, 1881

— ISody ])nbescent, cnneal fraetnre wide and dcep; hemelytia

more or less opaipie (Iha/il) . . . .LVNHÍIA.A Carvalho, 1951

cm 1

SciELO

10 11 12 13 14 15

Carvalho — Kc-ys Gcncra Miridac of tlie World

!). Cly])cus i)r()jcctinj^ hcyond apcx of first aiitennac; cmbolium

vcry wide and thiii ,fig- 55) (North &; Sontli America) ....

KUllYClllLOPTERELLA RciUcr, 1909

— Clypeiis iioi |)r<>jecting beyond apex of first antcimal segment;

cmbolium not as above 10

10. Rosinim reacbing to or sligbtly beyond anterior coxae; cyes

removed from pronotnm by a ilistance ecpial to at Icast one-

balf of its lengtb; small glabroiis siiccies with strongly produeed

Iicad and pronotnm (fig. 00) 11

— Rostrnm reacbing middie or jjosterior coxae; eycs contiguous

to pronotnm or so; species if small, with bead and pronotnm

not as above 12

1 1 . llody fincly piinctatc; second antennal segment incrassate

towards apex, as tbick as the first (Hra/.il)

ANNIESSA Kirkaldy, 1903

— Ikidy coarsely and dccply pnnetate; second antennal segment

linear, mneb more slender than first (fig. (iO) (África, Anstra-

lia, Cbina, IMtilijjpincs) EÍNGULUS Distant, 1901

12. 1’ionotnm carinatc on lateral margins 13

— Pronotnm not carinatc on lateral margins M

13. Ilemielytra setose; eyes tbick and erectly pilose (Anstralia, Ín-

dia) CÍM1CAJ’SUS Poppins, 1915

— Ilemielytra not setose; eyes glabions (New Zealand)

ROMNA Kirkaldy, 1900

11. Second antennal .segment distinclly clavate apically .... 15

— Second antennal segment not clavate apically 17

15. Cnnens strongly inclitied; ba.sal joint of biiul tarsi tliickcned;

mendtrane nniareolate, claval sntnre with a row of pnnctnres;

small species abont ■! mm lotig (Nortb America)

KEOPICORÍS Van l)n/(;e, 1915

— Cnnens not stiongly inclined or if so tbcn corial sntnre witliont

a low of pnnctnres: sjtecies over •! mm long 15

10. Deep black, polisbed species abont 8 mm long; general aspect

of Dcrncocoi is or (iaj)sus; claws not distinctiy tootlied at base

(North .\merica) PERAROCAESUS Knigbt, 1920

— Species not cleej) black, nsmdly less than 8 mm long; claws

distinctiy tootlied at base (New («ninea)

IRASEUS Distant, 1901

17. Scntellnm byaline, vitieons; bocly glabrons; eyes removed from

pionotnm by an espace ccjiial to abont twic.e tbe thickncss of

second antennal segment (lig. 210) (Alric:a)

YERONIA Carvalho, 1951

cm i

SciELO

LO 11 12 13 14 15 16

Hoi,. Mus. Goki.di — 'loiiio XI (II) — Dezembro 1955

— .Scutclhim not liyaline or vitreous; if so, tlieii cycs toucliiiifí

pronotum or body pubescciu 18

18. Collar covcrccl witli a whitish <lust-likc powder; antciinac witli

loug sctac and short hairs; head alniost horizontal and pro-

dnced bctwccn thc bases of antennae (Europe, Asia)

ALLÜEOTOMUS Eiebcr, 1885

— Collar witboiii ilie whitish dnst ahove; antennae with single

type of ])uhescence: head not noticeahly horizontal or elearly

prodiued hetween bases of antennae 19

19. 1’ronotnin very strongly ]ninctate, vertex distinctly carinate 29

— Pronotum if strongly punctatc, then vertex not carinate . . 21

20. body long and erectly j)ilose: second antennal segment with

a few hairs and long erect .setae ( rasinania)

PSEUDOCAMrrOIiROClIIS Popinus, 1911

— P.ody glahrons; second antennal segment without long erect

■setae (Australia) EUItYIillOCIlIS Kirkaldy, 1902

21 . Rostrnm reaching the ai)ex of mesosternum or middlc

coxae 22

— Rostrnm reaching the posterior coxae 2.^

22. .Second antennal segment a little incrassate near apex; clavus

with a series of piinctiires following claval commissure (Java)

LAMPROCRANUM Reuter,1891

— .Secoml antennal segment linear; clavus without a series of

punetnres following claval comissure (índia)

DüRTUS Distant. 1910

2.S. First segment of rostrnm reaching for heyond base of head;

second segment of hind tarsi slightly longer than first (China)

CY PI IODEM] DEA Reuter. 1901

— First segment of rostrnm reaching ha.se of head; second segment

of hind tarsi usually shorter or as long as first (Cosniopolitan)

DERAEOCORIS Kir.sc hhamn, 18.55

21. First antennal segment very thick. thc internai margin slightly

roíindcil, the externai sinnatc: vertex sulcatc longitudinally:

second antennal segment longer than head and pronotum to-

gether; cmholium wide (Ihazil)

PíESrorOMUS Ilergroth, 1922

— First antennal segment not noticeahly thick, more or less

straight; vertex smooth; second antennal segment shorter than

head and pronotum togethei ; cmholium narrow (lha/il) . . .

KiASriarUS lleigrotli, 1922

Note: '1 he genus Rnidn bmh. \Vhite. 1878. (Fnt. Mo. .Mag.

15: I.S2) froin New /.ealand is no| imluded in this Key, due to its

incomplete des< ription .

cm 1

SciELO

10 11 12 13 14 15

(Iarvai.iio — Kcys Ciciicra Miiidac ol lhe VV^orlcl

ki:y ro riiii gknera of saturniomiiuni

1 . Calli witli a V-shaped depression between tliein and a very

distinct occlloid structuic laterally; pronotum punctatc (New

Ciuinca) SATUIINIOMIIUH Kirkaldy, 1902

— Calli witliout a tlepression between thein; pronotum sinooth 2

2. Calli witb tbree laiy^e lossae or depressions behind tliem; pro-

notinn carinate laterally behind thc ealli (lig. 70) (Anstralia)

TRILACCUS Horvath, 1902

— Calli with a continuons snlcns behind; pronotum not carinate

laterally; an ocelloid shallow fossa laterally to calli present (New

Cbiinca) IMOGEN Kirkaldy, 1905

KKY ro I’HE CiENERA OF liRYOCORINI

- 1 .

llemielytra withont membranc on both sexes or the latter

only vestigeal; clavns conflnent with corinm (lig. 137).. 2

llemielytra Avith a distinct membranc at least on malc; clavus

and corinm iisiially ilistinct (fig. 131) 1

Rostiinn rcaching thc middlc coxac; .sccond antcnnal segment

as thi(k as first segment; outer margin of corinm roimded;

aspect of roleoptera (fig. 137) (Hrazil)

COl.EOrrEROMIRIS Carvalho, 1910

Rostnim rcaching lhe posterior coxae 3

Eyes pednncnlatc; sccond antcnnal segment more slender than

first segment; inner margin of corinm straight (fig. 231)

(México) AZTECARIELLA Carvalho, 1951

Eyes not |)eilimcnlatc: sccond antennal segment as thick as

first segment; inner margin of corinm stiongly sinnate till

lhe apical third (Cuba, San Domingos, Hra/il)

HEMISriIAERODEEEA Renter, 1908

Eyes on a snberect styhis which is at least as long as tbc widih

of onc eye, thc cnnens at mosi twice as long as wide

(lifí- 17») ^

Eyes sessil or at most bni siibstylate, in ihis casc, (iinens more

than twice as long as wide ‘

Rosliiim veiy shoit. not rcaching lhe apex of anterior coxae;

body with shorl, adpiesseti pnbescem e, lhe hemielylia iii-

gonscly pmutatc (Mentawei) SIEORÍA 1’oppins, 1915

Roslinm leathing the poslciior coxae or beyond il; body

withont adpressed pnbescence; hemielylia pnnctaie or smoolh

bni nevei nigonse *’

cm i

SciELO

10 11 12 13 14 15 16

‘iO Hói.. Mus. (ioKi.Di — romo XI (II) — Dc/ciiihro I95.5

(i. Rostruin rcacliing bcyoiul tlic posterior coxac; pygopliorc wilh

a S|)iiiil()i in projection bciit clown; bemiclytra ilistinctly puiu-

liired; first antemial scgiuciit about as thick as sccoiul (Norib

& CIciitral America) ÍIESPEROLAliüPS Kirkaicly, líIOli

— Rostrum rcacbing tlie posterior coxae only; pygojjborc witboiit

a s|)iiiirorin projection bent dowii; bemiclytra smooth or so;

first antcnnal .segment Iwicc as tbick as secoiul (Clentral Ame-

rica) NEOLEUCON Distant, 1881

7. Eycs substylatc with a sliort j)cdnnclc or if iiot, tbeii cimciis

very long and narrow, abom tbrec to four times as long as

wide at base, witb inner margin bent following tbe cnrvatnre

ol onter margin of membranc (figs. 127, Ml, MD) 8

— Eyes not sidrstylatc, ennens not as long and narrow as above,

nsnally witb straight margins 17

8. Apex of ennens reaching ibe distai or apical margin ol mem-

branc (cxcept femalcs of NcouecUa in tbis case, tbe rostrnm

leacbing tbe base of abdômen) (figs. Ml, Mb) í)

— Airex of ennens not reacbing lhe rlisial or a])ical margin of

membranc: ibe rostrnm never reacbing tbe base of abilomen

(fig. Mí>) lli

9. Rostrnm reacbing at most to a|)ex ol middle coxae; s|)ecies

ncv'cr leddisb or Inteons 10

— Rostrnm ccacbing tbe posterior coxae or beyond it; species

witb reddish or Inteons colonr 11

10. Rostrnm reacbing tbe middle of mesosternmn: scntellnm

smootli: eycs not strongly rccnrved (Clentral ík- Sontii ,\meri-

cas) SPARTACUS Distant, 1881

— Rostsnm reacbing a|>ex of middle coxae; scntellnm distinctly

[mnctnred; eyes strongly lecnrved ([ava)

MíCROliRVOCORÍS Eopirins, líH I

1 1 . Ecmalcs witb ennens not reacbing apex of mcmbiane; ennens

of malc very broad (fig. M(i) (.Sontli America)

NEONEEI.I.A Costa I.ima, 11)12

— Ecmalcs witb ennens reacbing ajrex of membrane: ennens of

niale very narrow — (India, Malay, iMiilippincs)

TlíAUMASrOMlRlS Kirkaicly, 11)02

12. .Seconcl antctinal segment longer tlian first segment; rostimn

very sliort, reacbing apex of first ccixac; ennens sbort (fig. 2.S7)

(África) AÍ/iVf/AY;f/,l Carvalbo, ID.ál

— .Seccmcl antcnnal segment longer tban lirst segment: rostrnm

reacbing beyond a|H'x of first coxae I.'l

I.H. Calli very large witb a deep meclian tiiangnlar depression; an-

terior area of pronolnm smootb; eyes Inrned backwaicis reacb-

ing beyond |Misleiior maigin of bead by a clislance ccpial to

(Iarvai.iio — Kcys Clciicra Miridac of thc World

•11

about l/‘l oI Icngth of cyc; sidcs of licniielytia parallcl (fig.

125) (África) STKNOPTEROCORIS China, HMl

— Calli willioiit a incdian triangular dcprcssion betvvcen tbcm;

collar j)unctatc: cycs not as abovc; sidcs of bcmiclytra sligbtly

widcncd at iniddlc 14

11. Kycs scssil; first antcinial scgincnt slcndcr only at extreme base;

vertex convex 15

— Kycs strongly prominent, sidtstylated; first antcnnal segment

mnch slender on basal half or third; vertex concave and sligbtly

sulcate 1()

15. Collar as wide as thc cyc; thc latter emarginate postero-inter-

nally, recnrvetl over tbe collar (Java)

' MYOC APUS Poppins, 1!)11

Collar not as wide as tbe cyc; tbe latter ont rcctn vcd backwardly

over tbe collar (Central America)

NEOSII.IA Distam, 1881

IC). Cnnens very narrow on apical balf or tbronghont, somcwbat

enrved; cye pednnclc abont as higb as widtb of onc cyc (fig. 1 15))

(C. k S. America, índia, Malay) .... SlNERl'US Sial, 1860

— Cnnens not noticeably narrow, more or Icss triangular; cyc

pednnde not as higb as widtb of onc cye (África, Índia)

PRODROMUS Distant, 15)01

Rostrnm reacbing tbe posterior coxac or bcyoiul it or wbcn

tbis is not tbe case, color reddish or Intcons and black .... 18

Rostrnm not extending bcyond tbe ajtcx of middlc coxac .3.8

.Small species, nsnally with orange or Intcons and blnish or

black color; males with a characteristic prong on Icft dorsal

side of pygopbore 15)

— .Speties with otber color pattern or if so tben males witbont

tbe prong on genital segment 20

15). Calli siongly convex (fig. 113) (Ameiicas)

CAUI.OTOPS Bergiotli, 185)1

— Calli not strongly convex (fig. 126) (Nortb America)

IIALTICÜTOMA Renter, 1913

20. Rostrnm reacbing fonitii abdominal segment or bcyond it 21

— Rostinm not reacbing thc lonrtb abdominal segment .... 22

21. Collar stiongly depressed, tbe apical corneis of pronotnm tn-

bcrcniate; second antennal segment incrassate at apex, sborter

tban tbird; claval corial and tostai veins with pnnctmes and

a low of brigbt bairs; cnneal fraetnre oblitpie, tbe tnnens

sboit, ronnded apitally, oblitpie, with same wititb tinongbont

(Vene/nela) PRISEOS EM RA Renter, 185)2

17,

18.

cm i

SciELO

10 11 12 13 14 15

•52

Mi s. (íoi i di — 'I oiiio XI (II) — Dt/cmliro 19.5.5

— Collai nat sti()n>>ly dcjircsscil and a])ic:al (()riicr,s t)[ pronolinn

not tuliciTulatc; vcins of liemiclytra not as abovc; cuiicus slcn-

dcr at apex (Hra/il) .... STICTOLOP! WS HcrRiotli, 1922

Íi2. Pionotum sinootli or riigoiisc 23

— 1’roiiotmn distinctly puiictate, soinctiincs vcry fincly so (fig.

23. Rüstrum surpassing tlie apcx of jiostcrior coxac: Icinoia in-

ciassati;: collar alxnu as long as íirst aiitcnnal .scginciu tliick,

liead triangidar (IJomco) NOTIDIUS Usiao, 1914

— Rostnnii not surpassing api-x of posterior eoxae; Icinora not

iiurassatc; eollar twice as long as first antennal scgnient tltick;

licad roíinded 21

21. Rostrnin not rcaeliing bcyond liind eoxae (Central America)

PACHVroihí Carvalho & Cliina, 1951

— Rosinnn reaeliing lhe ihird abdominal segment (I-iva)

MAUliOCOniS Poppins, 1911

25. Pronoinm witb anterior portion strongly convex and .somewliat

liooded over tbe bead, distinelly loveolate in tbe tentei of

(onstrit tion: seulellnm partially tovered by trnneate |K)sterior

margin of pronoinm (Ceylon, Pliilippines)

i:iiNEST/i\US Distam, 1911

— Pionotum not as above; sentcllum ex|Kjsed 2(')

2(i. F.yes sligbtly stylate or prominent, tiirved posteriorly, toiiili-

ing apex of pronotnm or protrnding baekward over lateral

sides of tbe latter (fig. 171) 27

— Fyes sessil, not prominent or (iirvcd |M>steriorly 30

27. Color orange red and bluisli metallit; bemielytra notiteably

longer tban abdômen, areola triangular its apex reailiing muth

beyoiid a|K'x of lunetis, veins densely pilose; setond autennal

segment witii long, erett selae (Malay, Java)

Ml.kril.A Distam, 1901

— (.olor orange led or luieoiis, ii bluisb tinge present tlien liemi-

elytia not notiiably longer tban abdômen, areola not as above,

veins glabrous or nearly so; setond antennal segment svitiiout

long, erett setae 28

28. I.arge speties over 5 mm witii a basal extavation on siutelium,

|K)sterior margin ol pronotnm snbtruntatc (Cential .Soutb

Amerita) SI.El.l.A Reuter. 1909

— .Small sjhtícs not over I mm long witii convex stutelium, |kis-

terior maigin ol pionotum noticeably einaiginate (Pbilippi*

lies) KMC.IITIOEA llsiao, 1911

29 llemielytia parallel: s|h ( ies witiioni traces oí Intcons or leddisli

colonr (fig. 1.50) 30

— llemielytia distinctly or sligbtly wicleneci at miticlle; speties

Carvai.ik) — Keys (kiicra Miritlac of tlic \V<)ilcl

30.

31 .

31».

33.

31 .

30.

liitcou.s, oiangc or rcddish with black or bluish inctallic tinge

(fig. 131) 31

.Sdiiellnm vonvex and sinootb ((Ibile)

PACHYNEURRHI MENUS Rcuter, 1909

.SdUelbim ilai triangidarly cònipresscd iii iniddle, the impres-

sioii angu late (New Ciuinea)

ELATYPELTOCORIS Poppius, 1912

l''irst antennal .segincnt about a.s long a.s lialf lhe width oI

vertex; pronotal collar soinewbat boodcd over the hcad; hemi-

elylra very broad ])ostcriorly'(lig. 132) (Clentral ík .South .\me-

rica) MECOLAEMUS Ilsiao, 1917

First antennal seginent longcr lhan half the width of vertex;

pronotal collar not hooded over the heatl; hemielytra not no-

ticeably broad jwstcriorly 32

lleinielylra finely ])unctulate, without bluish inctallic colour

(Ilidia, .Malay, Philippines, Australia)

IIROMEIJAEMÍRIS .Schuinacher, 1919

lleinielytra sinooth, with bluish inetallic tinge (Central &

.South America) TENTIIECORIS .Scoti, 188(5

Rostruin short and usually very thick, reaching the anterior

toxae 31

Kosiruin usually sleiidcr, reaching to iniddle of incsostcrnuin

or lo iniddle coxae, sonictiincs extending to its apex or so 39

Fiist antennal segnient shorler lhan width ol vertex; species

with black heniielytia and luteous or reddish heail amí pro-

noiuni (lig. 12.3) .35

First antennal segiiieiu as longa as or longer than width of

vertex; siiiall speties without traces ol luleouS or reddish .3(5

.Shining blatk .s|)ecie.s with head slrongly producetl inleriorly;

pionotuiii iioliceably narrowed anteriorly, glabrous, sditelhnn

pundate; leinales niat loplerous, cuneus dedivous, cuneal fra-

duie deep and wide (Fig. 153) (Central Jk- .South .\incrica)

ROrUROPHOREA.EA Reuler, 1907

Color not as above, head not pioduted inleriorly; pronotuin

not iiolíteably naiiowed anteriorly, pilose, sditelhnn oiily rn-

gose; l('iiialcs usually brachypleious, diiieus hori/ontal, diiieal

liaduie shallow (lig. I 12) (F.ui(>|K‘, .\sia)

RRYOCORIS F.illeii. 1829

I lead not noiiceably piodiued belweeii lhe anieiiiiae below;

Itoiis siiKHith, eyes siiiall, not diived |H)slei ioi ly; (ollai wide

(lig. 135) (.South .Vnieiií.i)

iSPlDono riIRUS Keuin, 1907

— 3

34 MtJS. Goki.di — r<)ino XI (II) — Dc/cnibro 1955

— IleacI strongly produccd bctwccii thc antcnnae bclow; fioiis

striatc; cycs and vcrtcx soincwhat cmvcd postcriorly; collav

narrow (figs. 123, 134. 13G) !... 37

37. Pronotum glabrous, strongly shining; general color blnisb ine-

tallic (fig. 130) (Central & South America)

líURYCIPITíA Renter, 1905

— Pronotum distinctly jiilose; general color not blnisb me-

tallic 38

38. Small sjjccies, nsnally 4.5 mm long or less, with boily strongly

ovoid (fig. 123) (Central America)

DICRIIOÜCORIS Renter, 1909

— Species nsnally over 4.5 mm long, with body clongate (fig.

134) (Central & South America) .... SYSÍNAS Distant, 1883

39. Pronotum smooth, nsnally shining, sometimes rngose, bni never

punctate 40

— Pronotum punctate, sometimes finely but distinctly so . . 42

40. Pronotum raiscd in middle behind calli and produccd ever the

lattcr; hind femora curved with a decp exeavation ou thc upper

surfacc (Cuba) NOTüLOIiUS Rcuter, 1909

— Pronotum not as above; hind femora without an exeavation 41

41 . Pronotum not strongly narrowed in front; head withont a net k;

short, ovoid, dark and small sjrecies (Madagascar)

MüNALOCüRüRSlS Poppius, 1914

— Pronotum strongly narrowed in front; hcail with a distinct

ncck, distinctly longcr than wide; claval veins usual ly with a

row of punctures (fig. 164) (Pacific Is. Madagascar)

1‘KI.ISACUS lYisV.un, 1901

42. Small reddish or black and red spccies less than 4.5 mm long;

sccond antennal scgmeiit about as thick as first, with unusually

short pubescence; third ;md fourth segments very slendcr (fig.

196) (.South America)

RACIIYMRROCI-RUS Renter, 1909

— Spccies with other color or if so, thc .secítnd antennal segment

not as tliick as first or beset with long, eicci pubescence and

larger than 4.5 mm long 43

43. Kyes producctl out and hackwards, their innei margin levei

with externai margin of collar; the latter distim tly set oíf fiom

pronotum by a postei ior furross' as wide as thitkest portion

of first antennal segment (about as long as hall the lenghi ol

one cyc or 0.12 mm) (índia, (ieylon)

DIOCI lAiUS Distant, 1910

- liyes and collar not as above or íl so, cithersvise colonrcd II

C^ARVAi.iU) — Kcys (ic-ncra Miridae oí tlic World

‘M. Proiiotal rollar ditinct iiot widcr than width of sccond antcnnal

scgmciit, distinctly dcliinitcd íroin disc of pronotuin, dark small

spccics usually witii light cmboliiiin, nieinbraiic fincly densely

pubcsccnt (fig. 172) 45

— Tronotal collar indistinct or if distiiict tbcii widcr than width

of sccond antcnnal scgincnt, usually not distinctly dcliinitcd

from disc of ])ronotuin, scldom small, dark pilosc spccics if

so incinbranc always glabrous (fig. 133) 47

15. First antcnnal scgincnt longcr than width of vcrtex, distinctly

narrowcr on basal balf; clongatc, convcx spccics with hcmi-

clytra parallcl sidcd (Malay, Burma)

IIEKISTA Kirkaldy, 1902

— First antcnnal scgnicnt narrowcr oídy on basal third or extreme

base: sborter or c<pial to width of vcrtex; ovoid spccics with

bcmiclytra ililatcd at midtlle (fig. 172) 46

16. (aincal fracturc dccp and very witle; cuncus curved cxtcrnally;

bcmiclytra flat (Mcxico) .... CYCI.lDOí.ON Reuter, 1909

— Ouncal fracturc sbalow and narrow; cuncus straight cxtcrnally;

bcmiclytra more or Icss convcx (Cosmopolitan)

MONALOCORIS Dablborn, 1851

17. .Small spccics mostly bbuk witb bcad porrcct, apically acute,

mote or Icss triatigular; first antcntial segment about as long

as or sborter iban widtb of vcrtex; |)ronotum slrongly piceous,

usually inilatcd and mneb bigbcr tban tbc bcmiclytra (figs.

133, 118) 48

— Spccics witbout tbc altove combination of cbaractcrs .... 50

18. Ilcmiclytra covcrctl by silvery silky or woolly pubcscence; outer

margin of cycs about levei witb anterior margins ofp ronotum

(fig. 133) ((ictitral & .Soutb America)

CYRTOCAPSUS Reuter, 1875

— Mcmiclytra witbout silky or woolly pubcsccticc; outer margin

ol cycs ptodiucd bcyond anterior margin of |)ronoluin by at

Icast balf tbc widtb of onc cyc (fig. 118) 19

19. Fmboliuin narrow and iiuras.salc; bcmiclytra witb latbcr long,

scinicrcc t pubcstcncc, witbout silvery sjtots or arcas; pionotum

|)ostci iorly niodcratcly inilatctl (Nortb, (Icntral ík S. America)

SlXEONOrUS Reuter, 1875

— F.mbolium broadly expanded and flat; bcmiclytra witb very

linc, sbort and crcct pubcsccncc and silvery spots or arcas; pro-

nottim |M)stciiorly gicatly inflatcd (Noitb, Clcntial & .Soutb

amcrica) PYCiVODERl.S Clucrin ík Meu. 18.56

5(1. Fmbolium vciy wblc and tonspit iions or swollcn at mitldlc

always witb a pit likc dcpicssion or wbcn tbis is not tbc case

(fcm.de of //. ililtitiitíi\) body very stiongly sbining, bluisb oi

3() Uoi.. Mus. (ioKi.Di — Tomo XI (II) — Divcmbio l!)r)5

j>rccnisli mctallic (figs. 150, 105) 51

— Kniholiiun il motlificd, ncvcr willi a pit likc clcprcssion 52

51 . Hody witii mctallic bhiisli or grccnish colour; ciiiiciis with a

pit likc dcprcssioii, cmholiar pit conspicuoiis, opcn towards

tlic oiitsidc (lig. 105), (Cuba)

IIETEROCORIS Gucriii & Mcn., 1850

— Hody witbout mctallic bluish or grcciiish colour; cuiicus witli-

out a pit likc clcprcssion, cmboliar pit dccp, round or oval, not

opcn towards tlic outsidc (fig. 150) (Central America)

METAEUR/US Carvalho & China, lí)51

52. Kmbolium distinctly laminatc about, as wide as or wider than

balí tlic width oí vertex; spccics with otlicr colour than black

(íigs. 158, 108) 5.3

— r.mbolium not laminatc, ccpially wide throughout, usually

incrassatc, realy narrowing toward aj)cx, ncvcr as wide as balí

tbc width of vertex or if so, small and black sj)ccics 55

53. Kmbolium strongly dilatcd on bascd tbird .so ibat tbc basal

|)arl of costa fornis an obtuse anglc with tbc apical jiart of

costa (fig. 108) (Central &: .South America)

EMIiüUüCORIS Carvalho & China, 1951

— r.mbolium laminatc tbrougbout, not noticcably widened on

basal tbird, narrowing gradually toward apex 55

51. l-.mbolium strongly arcuatc cxtcrnally; sculcllym with a tumid

basal lobc projccting backwards and a Hat, pointed a])ical

lourtli; hody very long, fine and crcctly pubcsccnt cycs not

rccurvcd (New Càilcdonia) .... (iUS’HAI)lA Distant, 1920

— Kmbolium not strongly arcuatc cxtcrnally; scutcllum not as

above, convex; hody with sbort, crcct pubestente; cycs strongly

rccurvcd, collar sonicsvbat boodcd over lhe vertex (fig. 158)

(Ilrairil) y.lKANlOEA Carvalho, 1910

55. Kmbolium wide at base, narrowcd lowartl apex, aficr tbc mid-

dic; coslal vein with a row oí punctures; hody strongly lound-

cd (New (iuinca) ... HEM/SEIIAEROCORIS Koppius, 1912

— Kmbolium if wider at base, ncvcr wiih a row of punctures over

(osial vein; botly not noticcably rouiulcd 50

50. Ihonotum vciy toaiscly and dccply punclalc, glabrous, lhe .si/.c

of thc pum lures cipial lo lhe width of second ;inlcnnal seg-

incnt (fig. 17.3) (Central .Amciiia)

XO TO TREMA EES Carvalho !k China. 1951

— 1’ionotum not nolitcably toarscly pumiale, pilosc, si/c ol thc

pundmcs sm.dici than widih ol sciond anicim.il scgmcnl 57

Carvai.ho — Kcys Ciciicra Miridac of tlie World

57.

58.

5'.).

I Icad in dor.sal vicw apically pointed; proiiotuin finely piibcs-

ccnt and punctuicd; hind tibiac linear (íig. 110) (Central

America) KNlGHTOCOIilS Carvalho k China, 1951

llead in dor.sal view a[)ically roíind; pronotum distinctly pu-

bescent; hind tibiac usually tbickcned toward ape.v 58

Idytra witli vcry long, crcct, fine pubesccncc; inale with lirst

antennal segmcnt tootiicd (fig. 150) (Central America)

• ODONTOCKROCORIS Carvalho &: China, 1951

Elytra with short adprcssed pubesccncc, if crcct or scinierect

tlicn witlioiit a tooth in last antennal scgnicnt 59

Klytra jxirallel sided (fig. 150) 00

Klytra wider across iniddlc than at base or apcx (fig. 122) 01

(iO. llead ilistinctly exserted, with a neck as long as the eyes which

are jilaced aboiit its iniddle (North America)

TYí.OCAPSUS Van l)u/ce, 192.3*

— llearl not exserted, if a short neck is present, the eyes Icss distant

from pronoinm 01

(il. Scntelhnn totally covered by pronotnm (Central &; .South ,\mc-

íica) PSEUDOIiRYOCORIS Distant, 1881

— Scntelhnn not covercd by pronotnm 02

02. Sinall, elongatc dark .siiecics; pronotnm with two distinct cons-

trictions, collar and calli together abont as long as disc; males

with a wide and dcep snlcns on frons, the vertex bifoveolate

(Índia, .Ceylon, IJnrma) HARPEDONA Distant, 1901

— Species with a distinct color pattern; anterior iiortion of pro-

notmn nsnally shortcr than disc; males withont the snlcns

mentioncd above 03

03. Pronotnm Hat, the cnnens very narrow and pointed (Philip-

pines) EüPURIVS Poppins, 1915

— Pronotnm more or less inllated convex; cnnens long and wide

at base (Central America) NEOEURÍUS Distant, 1881

01. I.lytra vciy oval and Hat, with dense, shoit adprcssed pnbes-

cence (fig. 122) 05

— I.lytia veiy slighlly widened in iniddle, with rather sparse

erett or scmierect pubesccncc 0<'>

05. Setond antennal segmcnt more than twitc as long as the líisi;

division beiween loiinm and cnnens not distinct; species nsnally

laiger than (i mm long (Central .Vmciica)

MA/.A Distant. 1881

* I lii< RriiiM M.M sliiilird irtriill)' liy llic anilioi .iti<l lnuiid Ki Ik- :i Synniipii

"I Mni iitlitlilnn l irhrr ittii >|ddiii).

38 Iloi.. MtJs. (ioKi.Di — 'lomo XI (II) — Dc/iniliro 1955

— Sccoiul antcuruil scgmcnt Icss than twicc as loiig as firts; clivi-

sion bctwccii coriuin aiuI cuiieiis distinct: spccics usually Icss

lliaii 5 nim long (Central & South America)

PARAFURIUS Carvalho & China, 1951

fifi. CoIIar area somcwhat projecting over hase ol head; second

segnicnt of antcnnac sliortcr than wiilih o! head; scntelhnn

punctatc (fig. 218) (México)

EURYCAULEIA.A Reuter, 1909

— Collar arca not projecting over hase of head; second antennal

segniciu longer than widlh of heail; scutelium not pnnclate

(Central & South America, New Guinea)

ECC.RITOTARSUS Stal, 1800

Note: riie following genera are not iiuhuled iu tlie key, since

thc typcs were not seen and the descriptions are incom])lcte:

Cobalorhynclius Reuter, 19()(’) (Anu. Mus. 7.ooI . St. Petersh.

10: I): China.

Eopidolon Poppius, 1911 (Ofv F. \Y’t. So( . Forh. 5.SA (2):

7): índia.

Pcrissobdsi.s Reuter, 1892 (Ann. Soc Ent. Fr. (il: 397); Venc-

7ucla.

KEV IO FIIE (iENERA OF MONAI.ONIINI

1 . Femora with twoswcllings at apex, thc ilistal onc large, halloon-

like; scutelium smooth and flat (fava)

iRTinUTlCUS Hergroth, 1923

— Femora not .swollen at a|K'x as ahove or if so, scutelium armed

with one or more spines or processes 2

2. Scutelium flat or convex, sometimes hiaddei like or tystiform

with tuherdes, hut never true spines or pnxesses 5

— .Ssiitelium armed witii one or more spines or processes (figs.

120, 121, 159) 3

3. .Scuteilutn with a single, very long and slender spine ending

in a hution like knoh; aittcnnae very long :ind linear; apex

of etnium witliout a shield shtiped elcvation (fig. 159) (África,

índia, .'\nstr;ili;i) UEI.OPEI.TIS Signoret, 1858

— Scutelium witliout a very long spine ending hy a holton-like

knoh; antennac with segments incrassate apically oi at middie;

a|K'X of roíium with a shield slia|K'd elcvation ■!

1. .Scutelium with a thick, high, mushroon-like piocess; |>osteiior

angles ol pionotum not piiKluced into a Hat spine; aniennae

with ápices not h.dlcKm-like inilated (lig. 121) (.Mrica) ....

PUrSOPUOROP PERA Poppius, 1910

Carvai.iu) — Kcys Gcncra Miiiclac of the World

Sciitflliim witii a nicdiiim prorcss braiichcd apically iiito a pair

o[ sliorl, poiiUed s|)ms: jwstcrior aiiglcs of pronotuin produced

iiito a flat, jjoinfcd proccss; antennac with apiccs of joints

iidiatcd iiito balloon-likc knobs (fig. 120) (África)

rilYSüPlK) liOrTKRELLA Poppius, 1914

Kinbolio (orial and clavo corial suture rvithout piiucturcs;

prouotiuu and beniiclytra glabrous 6

Euil)olio corial and clavo corial sutures witb a row of dose

set punctures (bigh luagnificatiou); pronotiun or hemielytra

pubescent (fig. 139) 7

Secoud autcnual segincut little louger than first; apical angle

of areolac or veiu of lueiubraue aciitc and jOTiuted (Australia,

Fiji) EUCEROCÜRIS Westwood, 1837

Secoud autcnual scgnieiit about six times as loug as first; apical

aiigle of areole or veiu of membrane rouuded (fig. 138) (Cen-

tral & South America)

MONAI.ONION Herricb Schaeffer, 1850

Piouottuu strougly writikled; rostrum rcacliing the middle of

mesosteruum (África) .... PARARCULANUS Poppius, 1912

Prouotum smooth and shining; rostrum reachiug the apex of

anterior coxac 8

Fious strougly swolleu, ;is seeu from above produced well iu

fioiit of eyes above base of clypeus; first .lutemial segmeut

strougly thickcued but glabrous: membrane witb a siturious

veiu; calli promiueut as two rouiid, erect protubcranccs (Con-

go, .Madagascar) IRCULANUS Distaut, 1901

Frous uot swolleu ueitlicr irroduced iu frout of the eyes above

base of clyjKuis, tiuterioi" margiu seeu from above almost

straigbt between and levei with anterior margiu of eyes; first

aiiteimal .segmeut uot iuordiuate thickeued with erect hairs;

mcmbrauc wilhout spurious veiu; calli uot as erect protube-

rauce 9

Fmbolium as wide as thickuess of first autcnual segmeut: pro-

uotum (i)veied by loug setiloim hairs (fig. .30) (.Afiica) ....

POPPIUSEi China, 1914

Fmbolium uot as wide as tliicuess of fiisi aiiteuual segmeut:

if so. frous with three ttdteides: piouotum smooth or with

(ouuuou hairs oídy

Piouotum completely glabrous

1’iouotum pubescent **

Fioiis with tliiee tubeides beaiiiig loug liaiis; body above and

lhe auteuuae loug and eiecily pilose; ciiueus shoii; aieola with

iuuer apical aiigle almost straighi (Foimosa, Malay)

EVPACAIYPEEPIS Poppius, 1915

40 Bol. Mus. Goei.di — J onio XI (II) — Dc/cnil)ro 15)55

— Frons without tubcrclcs; Iwxly ahovc and aiitcnnae nol noti-

ccably Jong and crcctly pilose; cuncus twicc as long as wide;

arcolac with inncr apical angle slrongly acntc (Índia, Malay,

Pacific Is.) PACHYPKLTIS .Signoict, 1858

12. Hciniclytra with short,. adprcsscd jjubcsccncc (crect only on

clavus); scutcllinn covered by pronotinn at base (Indochina)

MANSONIELLA Poi)pius, 11)15

— Heniyclytra glabrous; scntelhnn not covered by pronotnin at

base (Anstralia) PACAIYPELTOPSIS Poppius, 15)12

KFY TO THF GENERA OF ODONíEEEINí

1. Mcndnane ot heiniclytra with a miinber of aiixiliary veins or

vein-likc iniprcssions extending Iroin basal cell to apex ot

meinbranc, thc cnbital vein distinct, arising froin the l)asal

angle of the ccll and extending along anal niargin (fig. 121) 2

— Meinbranc of heinielytra without such anxiliary veins, soinc-

tinics with a spnrious vein arising froin the ajiical angle of

hasal ccll 5

2. Frons distinclly swollen and jirodnced anteriorly bctween bases

of first antennal segnicnls, dcliinited froin verte.x by a sinuate

iinprcssion, first antennal seginent short and thick, shorter ihan

Icngth ot head with neck, twice as long as wide, tliird seginenl

very strongly thickened in iniddie, ninch thickcr lhan first

•scgnient (fig. 124) (.'\frica)

PANTIEÍOMORPUA Sclunnacher, 15)17

— FYons feebly swollen and not produced anteriorly hctween bases

of antennae, not deliinited from vertex by a sinuate iinpression,

first antennal seginent less thickened, slighlly longer than

length of head with neck four times as long as wide, ihird

segment not inuch thicker in middie than a|)ex of second seg-

nicnt (África) .... EYCÍDOCORIS Rcuter ík Poppius, 15)11

‘i. Frotis in front with threc strong anteriorly or upwardiy ili-

rected spines; first antennal segment densely beset with long,

crect scalc likc hairs 1

— Frotis without three of such s])itics; sometimes with two or threc

ttibercular process in which case, the first antennal segment

without scalc likc hairs 0

4. Last threc antennal segments with short hairs; lateral spines oí

frons distinctly bent outwards (África)

CIIAMOPSÍS Rcuter Ik- Pop|)ius, IDI I

— I.ast three atUetinal segtnents with long hairs; lateral spines of

frons distinctly hent u|)wai(ls (fig. 2.S1)) 5

cm i

SciELO

10 11 12 13 14 15

Cíakvai.iio — Kcys Genera Miriclac oi thc World *11

Frontal sj)iiies vcry Iiigh, as long as thc depth oi onc cye sccn

froin sidc; pronotiim with two lateral discai lohes; scale likc

hairs oi iirst antcnnal seginent vcry large (Airica)

PARACIJAMUS! .Schouteden, lO-lG

Frontal sjtincs short, sccn froin sidc not as long as thc depth

oi onc cye; jtronotiini without lobes; scale likc hairs oi iisst

antcnnal scgnicnt slcndcr (íig. 239) (Airica)

CflAMUS Distant, 190-1

Frons, ahove base oi clypeiis bctwccn antcnnac, with a pair

oi distinct conical jnotnberances, these rarcly niinntc or iused

into onc in which casc apex oi sccond and third and iourth

antcnnal scgincnts strongly swollcn; soinctiincs tnbcrcles minute

and setigerous and rathcr indistinct in which casc pronotinn,

posterior latcrally, strongly dilatcd, its niargins serrate antl

))ronotal collar anncd with íour tiibcrcular inoccsses CÜíís.

128, 129) ^7

Frons, above oi clypctis bctwccn antcnnac without a pair oi

conical protubcranccs or setigerous tidtcrcles, thc írons soinc-

times prominent bctwccn antcnnac in whidi casc apex oi sc-

cond antcnnal scgnicnt not or only slightly thickcncd .... M

Pronotal collar with four tubcrtular processes, thc inner ]3air

clongatc; suriacc oi pronotum wiih ten crcct conical processes

in two rows, thc two centre ones oi posterior row oi six, much

longcr and biggcr than othcrs; jrosterior lateral niargins oí

pronotum dilatcd and serrate; .seutelhun sjilit uj) into six lobes

(iig. 11)9) (Airica) YANGAM HIA , Schouteden, 1912

Pronotal collar without crcct tubcrcidar processes; suriacc oi

pronotum without crcct conical processes, thc posterior margin

oi pronotum not serrate and scutcllum not nmltilobatc ... 8

Puncluration oi jnonottim dccp and more or le.ss regular, thc

suriacc wiihotit small, shining, tuberctilar .swellings (iig. 129) 9

Puncturation oi pronotum Icss dcc]), rugouscly coniused, sur-

iace with small irrcgularly jilaced tubcrcidar, shining swellings

(fig. 128) 12

Sctitcllum strongly inilatcd, cystiiorm (iig. 129) 10

Scutcllum not noticcably inilatcd or cystiiosm (Philipjiincs)

VOLKELlOrSlS Poppius, 1915

Connexivum oi abdômen hirgely exposed; .scutcllum not covci-

itig thc clavus on sides; irontal tuberclcs longcr than wide

at base (iig. 128) (Spanish Guinca, Fernando Pó)

lUlYOCOROPSIS .Schumacher, 1917

Gonncxivum oi abdotncn covered by thc hemiclytra or only

slightly exjjoscd; sditclhim ptodmcd over thc clavus hitcrally,

l>or,. Miis. Goki.di — 'romo XI (II) — Dczenihio 1955

covcring it almost cntirely; frontal tubcrcies shortcr than wide

at base 11

11. llcad scen from above with two distinct tid)crclcs on anterior

inaiffins, tbc clyjtcns clistinctly visible betwccn thcin; scntellinn

bcinispberical, widest in middlc, tbe basal niargin ovcrlying

pronotnm arcuatc (fig. 16G) ('riiibet)

RIIOrAUCESCIIATUS Rcnter, 1893

— llcad scen froin above with tbe anterior tidrcrclcs fused to

forni an anteriorly trnncatc jiroccss wbich bides tbe clypens;

scntellinn shield sbaped, widest at base, tbe basal niargin overly-

ing tbc jironotnin, straigbt not arenate (fig. 129) (New Bri-

tai) PSEUDÜDONIELLA Cliina & Carvalho, 1951

12. .Scntellnin abont as higli as tbc pronotnm, pointed apically;

eoiinexiviim of abdômen nsnally covered by tbc bcmiclytra or

so; form elongatc 13

— .Scntellnm ronnd, mneb higbcr than inonotnm; conncxivnm of

abdômen largely exposed; form ovoid (fig. 128) (New Gni-

nea, New Jiritain)

EARAIiRYOCOROrSlS China &: Carvalho, 1951

13. Hind tibiac clistinctly nodnlonsely swollcn; eycs small, only

one cpiarter tbc widtb of vertex; acetabnla of anterior Icgs lar-

ge, visible from above on cach side of anterior collar (fig. M7)

(África) DlSTANTfEEEA China, 1914

— Ilind tibiae sim|jlc, not nodnlonsely swollcn; eycs large, abont

one Iialf tbe widtb of vertex scen from above; acetabnla of

from Icgs small, not visible from above (fig. 145) (África)

SAIU.liERGEEl.A llaglnnd, 1895

14. Rostrnm extending to tbe posterior coxae 15

— Rostrnm extending to tbe anterior or iniddle coxae .... IG

15. Ilcmielytra with minute scalelike bairs giving a sbagreencd

a|ipcarcn(e; connexivinn not exposed; bead pointed in front

(África) noXíA China, 1943

— Ilemielytra witiiont minnte scales; connexivnm largely ex-

posed; bead ronnded in front (Malay)

REATYXaOàíIRIS Kirkaidy, 1902

l(). Rostrnm reacliing to middle coxae; second antennal segment

stiongly incrassate at a|»ex; davns ])nnctate (tig. 130) (África)

IlEE/ERSICORIS Delattre, 1950

— Rostrnm reacbing to anterior coxae or sligbtl beyond; secotid

antennal segment not imrassate at apex; clavns not pnn-

ctate 17

17. Ibird and lonitb antennal segments clistinctly cbdisbaped;

scnicdlnm convex, not bigber than pKinotnm wben seett liom

C^ARVAi.iio — Kcys Gcncra Miriclae of thc World

side; hcniiclytra aml scutc-llum densely pllosc, moderately shin-

ing (Australia) VOLKELIUS Distant, 1901

— I liird and lourth antcnnal segincnts inorc or less linear; scntel-

linu strongly inllatcd, a littie liigher tlian pronotum when

seen froni the sídc; hemiclytra and scutellnm glabrons or

sligluly pilose, strongly shining (África)

OnONIELLA Haghlund, 1895

KI:Y to 4'HE GENERA of PHYLINl

1 . lUack spccies, brachyjjcrous; elytra without incnibrane, the

coriínn, clavns and cuneus fused 2

— Species with otlier color or i£ black and brachypterous then

the heinielytra with ineinhrane or coriínn, clavns and cuneus

not fused 3

2. Small species ahout l.G nnn long; second antennal segment

very thick, as wide as length of first segment (fig. 73) (Rra-

zil) TAPUliUYUNUS Carvalho, 191G

— Species ahout 4 inni long; second antennal segment cylindrical,

not incrassate (Caucasus)

IIOMALANEli Kiritshenko, 1951

3. Second antennal segment of ionsj)icuous shape (fig. 93),

strongly hent at middle like a letter U with a shorter arm;

legs vesy long (l)utch Guiana)

ANOMALOCORNUS Garv. & Wygodz. 1945

— Second antennal segment linear or incrassate hut never hent

or U sliaped; legs not noticeahly long 4

4. Third antennal segment glohose, heset with long, llattened

hairs (fig. 00); puhescence of hody erect, inlermixed with

silvery, flat hairs; species of small size and light color (North

and Central America) . . 1 1 AMULE/PON lOLA Carvalho, 1954

— Third antennal segment cylindrical, with common puhes-

cence ^

5. 1’ronotum ahove distinctly punclurcd (fig. 74) G

— 1’ronotum ahove smoolh or very line and iiulistinctly pun-

ctured

(). Pronotum with lateral margins explanate, straight, hroadly

and strongly rellexed; disc iriegidarly rugose; second antcnnal

segment strongly rellexed; disc irregnlarly lugose; second an-

tennal segment stiongly clavate (North America)

PRONOTOCREPIS Rnigiit, 1929

— Pronotum with lateral margins not rellexed or explanate; disc

not rugose; second antennae not davate /

cm i

SciELO

10 11 12 13 14 15

lloi,. Mus. Goki.di — 'J onio XI (II) — I)cv.cinl)ro 1955

'l’arsi oí posterior tiliiae very long, aboiit as long as hall' tlie

Icngth of thc lalter; cycs very large ( Turkestan)

liOOPIDOCORIS Reuter, 1879

'larsi imich sliorter; eyes not very large 8

Proiioturn coarsely i)unctinecl; clorsuin brilliaiit metallic; two

last antennal joiiits linear (África)

LAMPROSTHENARUS Poppius, 19M

Pronotum finely ]Hiiicturetl: dorsum not metallic; two last

joints of antennae fiisiform (St. Helena)

AGRAMETRA Biic. Wbite, 1878

Eyes snbstylate, distant from j)ronotnm; heinielytra transpa-

rent; first and second antennal segments incrassate (Brazil)

(fig. 78) CRASSÍCORNUS Carvalho, 1945

Eyes not snbstylate, contiguons with pronotum; heinielytra not

transparent 10

Ilody beset with scale-like hairs or llattened silvery hairs in-

termixed with common pnbescence (in thc lattcr case iisnally

tibitd spines with black sjiots at ba.se, eyes noticcably grann-

lose, second antennal segmetn lotiger than width of hcad,

rostrnm rcaching thc posterior coxac or littlc bcyond, vertex

not carinatc II

llody withotit scalc-Iikc or flattcned silvery hairs intermixed

with common pnbesícncc (if silky or wooly hairs are present

amongst othcr hairs, thcn withont thc set of characters pointed

above) 30

Fairly large, dark species with hcad much wider than long,

eyes snbstylate; vertex strongly carinatc; body beset with dense,

elongate whitish scalc-Iikc hairs; jiscndoralia apprcsscd to thc

claw, tdmost rcaching its apex (África)

EASIOLAíiOPS Pop[)ins, 1914

IJsnall small species; if large, then thc cycs sessile; p.sendarolia

not as above 12

Hcad iransver.se, frons vertical, not protrnding in front of

antennal bases as seen fiom dorsal a.s])ect (fig. 77) .... 13

llead prodiiced in front of antennal bases, if not distinctiy

.so then thc scale-like ]mbesccnce blatk (fig. 82) 15

Espace betwccn bnccula and cye not greater than thickncss

of first antennal segmeni cxcejn in females where distance

may exceed width of last antennal .segment bnt does not etpial

its leiigth (Américas) RillNACLOA Rcnter, 1870

Espace bctwecn bnccnla and eye greater than thickncss of

fitst antennal segment, nsindly stdiecpial to Icngth of seg-

ment 14

Carvalho — Kcys Ciciicni Miridac ol tlic World

■15

M. Second aiUcnnal scgniciu five times Icngtli ol íirst; liind tibiae

wiih light sjjincs without daik spots at base ( Transcaspia)

STHENAROPSIS Popjjius, 1912

— Second anteiinal segmeiu not over tbree times length of íirst;

bind tibiae rvitb black s])ines baving dark spots at base (North

America) LEFIDOPSALLUS Knight, 1923

15. Clypeiis sharjily ])iodiiced, apex irointed (íig. 99) 16

— Clypeus not ])rodnced, vertical, tbe a])ex blnnt (fig. 82) 17

16. I5oth sexes witb second antennal segment strongly incrassatc,

abont twice as long as íirst segment (Enrope, North yVmerica)

EXCENTRICOJilS Carvalho, 1955

— Only tbe male witb second antennal segment incrassate ( abont

íonr or more times as long as íirst (Kuro|)e, Asia, North Ame-

rica) CAUOCORIS Fieber, 1858

17. Second antennal segment strongly thickened, mnch broader

tlian tbe íirst (íig. 82) (Cosmopolitan)

ATRACTOTOMUS Fieber, 1858

— Secoiul antennal segment not strongly thickened, itsntilly more

slcnder tlian the íirst segmctit 18

18. Secotid antennal segment sligbtly compressed, tliickcst at mid-

dle wcre it is tbicker than íirst, coveretl witb very dense, íairly

long, semierect bbick ])nbescence (Aírica)

LEP/DOCAPSUS l’o])pins, 1911

— Second antennal segment linear, not tbi(ker tban and not

pnbescent as above 19

19. l.engtli oí second antennal segment or ecjnal to width oí head

across eyes

— Length oí second antennal segment greater than width oí

liead across eyes -1

20. liead inclined, ])i()dticetl in íront ol antennal bases: jnibes-

cence rvitb erect ;tnd silvery decidnotis bairs; íirst antenn;il

as long as lortnn (North Americ;i)

MEdAEOPSAEEUS Knight, 1927

— . liead loimded in íront; ])nbescence mostly oí sericeoits deci-

dnons bairs: íirst antenntie longer than loinm (North .\meri-

ra) EUROPIEEEA Renter, 1909

21. Scale-like pnhescetne black; a pseiido-pronotal collar present

(Kiii-ope) EXAERETUS Fieber, 1861

— Scale-like laibescence silvery; a psendo-i)ronotal collar

absent

22. llind tibiae witb light spines baving dark spots at base . . 23

— .Spines ol bind tibiae witb otber colonr ií light, tben witbotit

datk s|)ots at base

cm i

SciELO

10 11 12 13 14 15

■l(i

23.

I5()i,. Mus. GoKi.nt — roíiio XI (II) — Dezembro 1955

24.

25.

26.

27.

28.

29.

30.

]5o(ly legs and antennae witli minute fuscous or reddish spots;

scales on cuneus bfack (Guatemala)

CAPELLANUS Distant, 1904

Ilody legs and antennae without minute íuseous or reddish

spots; scales on cuneus not black (Ceylon)

DEMOPLESIA Poppius, 1913

Hiud tibiae with dark spiues without dark spots at base 25

Hind tibiae with dark spiues having dark si)ots at base or

with light spines without dark spots at base 27

Pseudarolia attached oídy at base of claw, tijj íree and extend-

ing to middle of claw (Américas)

REUTEIiOSCOPUS Kikraldy

Pseudarolia United with claw 26

.Setiform hairs black, strong and erect, espetially on vertex

and anterior margin of jtronotum (Algeria)

CUlirsOCUNODES Reuter, 1901

•Setiform hairs yellow or whitish, long and fine (Europe) . . .

PIIYEIDEA Reuter. 1899

Rostrum uot reaching the aj)ex of hind coxae or beyond it;

head seen frf)m above as long as pronotum (North America)

HOPEOMACHIDEA Reuter, 1909

Rostrum reaching the a])ex of hind coxae or beyond it; head

seen from above shorter than pronotum 28

Clyjjeus prominent, distinctly visible from the side; antennae

long, segment II linear, in male somewhat thicker ;ij)ically,

length ecpial to or greater than basal width of pronotum

(Cosmopolitan) PSALEUS Fieber, 1858

Clypeus nearly flat, scarcely visible from the side; antennal

segment II rather short, length not over onedialf or two-thinls

the basal width of pronotum 29

Vertex carinate or marginate; third segment of hüul tarsus

longer than second (Cosmopolitan) ^/'ACrVAnm

Vertex smooth, not marginate; third segment of hind tarsus

shorter than secoml (África)

STENOCAPSUS Bergroth, 1926

P.seudarolia arisiug from base of claw, free and convergent at

ápices 31

Pseudarolia if free and arising from base of claw, never cou-

verging at aj)ice.s 32

Head vertical, tibiae with black spots (África)

SCJIROEDERIEEl.A Poppius, 1914

Head uot veitical; tibiae without black sjtots (África, Madei-

ra, Madagascar) CEPUAEOCAPSUS Popiuus, 1911

SciELO

10 11 12 13 14 15

CIarvai.ik) — Kcys Cíciiora ^íil■i(lae of tlie World

‘17

32. Lcngth ol sccoiul anteiiiial segnicnt lesse tlian widtli oí hcad

across cyes; in species in wliicli tlie two are alinost equal, hind

rciiiora liglit with dark spots (fig. 97) 33

— Leiigth ot secoiul aiitennal segment greater tlian width ol

head acro.ss eyes; in species in which the two are ahnost eqnal,

hind feniora not light with dark spots 51

33. Light coloured sjjccies with fcmora yellow and beset with

conspicnons black spots ((íig. 90) 31

— Dark colonrcd species or if light, femora black to dark brown

or of 1'einora light, then without dark spots (sometiines with

light fiicons jx)ints or clondings) 39

31. Pnbcscence distinctly silky or woolly, adpresscd; black spots

of feniora only on the externai margins, not very conspicnons

(África) liRACHYCRANELLA Reuter, 1905

— Pubesccncc with sctiforni or fine hairs, but not silky or woolly

or adprcssend, if so then black spots of femora large and irre-

gnlarly jilaced 35

35. Rostrinn reaching slightly bcyond anterior coxae; pseudarolia

free and paralell (Enroiie, North África)

MOISSONIA Reuter, 1891

— Rostrinn reaching the middlc coxae or bcyond it; pseudarolia

connected with claw or not visible 36

36. Pubescence erect, bristlc-like; tibiae strongly spinose, length

of spines about twicc the diameter of tibiae (North America)

(fig. 191) PHYLLOFIDEA Knight

— Pubescence simple, not bristle-likc; tibial spines shorter 37

37. Distancc from lower apex of eye to buccula, seen from sidc,

etpial or less than lialf the hcight of eye: arolia visible (Cos-

mopolitan) CAMPYLOMMA Reuter, 1878

— Distance from apex of eye to buccula equal or about equal

the heiglit of one eye; arolia not visible (fig. 180) .... 38

38. Llypcus not extending backwards to a point beneath front

margin of eye; as seen from sidc clypeus and juga narrow

(Asia, Lurope, North America)

ATOMOSCELIS Reuter, 1878

— (ilypeus extending backwards to a point beneath front maigin

of eye; as seen from sidc, the clypeus and jugum very broad

(Algeria) APHAENOPHYES Reuter, 1899

39. Male antenuae with first aiul secoiul joint greatly thickenetl,

hemielytra black with a jjale mark ou clavus; femalcs some-

times brachypterous (fig. 97) (Américas) •• •

SPANOaON/CUS Berg, 1883

cm i

SciELO

10 11 12 13 14 15

IU)L. Mus. Goi.i.di — 'Joiiio XI (II) — Dc/cnibro I!)r).5

— Ilcmiclytra witlunit a l)alc niark ou clavus or if so, nialc an-

tennae siciuler, scarcciy thicker tlian iii ícmalc; thc latter ina-

(Toptcroiis 40

40. l*iibc.scciicc setiform, stiff and black 41

— Pnbcsccncc vcry íine, crect or sciniadprcssed, not setiform 42

41 . Pscndarolia alinost rcaching thc aiJC.x of daw; first rostral

segment rcaching anterior coxae; dorsuni and hcad not

strongly hirsute (Enropc) LITOXENUS Rcuter, 188!3

— Pscndarolia rcaching at niost thc middle of claw, first rostral

segment scarcciy surpassing thc base of hcad; dorsnm and

itead stronglv hirsntc (North America)

' PIIYLLOPIDEA Knight, 1919

42. Pnbcsccncc vcry short, almost glabrons sjjccics with body more

or Icss dnll; tibial s|)incs short and jjlaccd hcyond thc middle

of tibiac 43

— Ilotiy distinctiy jjnbcsccnt, nsnally shining; tibial spines placcd

thronghont thc tibiac, rclativcly long 44

43. Sides of inonotiim cmarginatc; hcad hlnntly ronndcd in front;

third segment of hind tarsi longcr than sccond (fig. 37) (En-

ro])c, Asia, North America) . . CONOSTETUUS Eicbcr, 1858

— .Sides of pronotnm straight; hcad distinctiy pointed in front:

third segment of hind tarsi as long as .sccond (fig. 100) (En-

ropc) STENOPAREl Eicbcr, 1870

41. Hcad vcry wide, lhe posterior margin scmicircniar, thc cycs

jnomincnt rcadiing backwards to thc middle of pronotnm

(Egypt) EURYCIIANEEEA Renter, 1901

— Ilead if wide, not with a scmicircniar jarsterior margin, thc

cycs never ícaching h:ickwat(ls to thc middle of pronotnm 45

45. Hind femora palc wilhont black or fnscons spots 40

— Iliiul femora black or fnstons or palc bnt in lhe latter case

with some blatk or fnscons s|)ots 47

40. Tibial spines palc; rostinin leaching apex of inesostcrnnm;

■setond anlennal segment slighily incrassatc toward thc apex

(África) J.EProXANTIlUS Reiltcr, 1905

— 'Tibial s])ine,s blaik: rostinm leadnng apex of mithllc coxae

or beyond; setond antennal segment linear (Entope, Asia,

Madeira) MAURODACTYEUS Renter, 1878

17. Hcad sccn lioni ;ibove strongly piodiited in fiont bctwccn

lhe aniennac (lig. 88) -18

Hcad sccn iioin ;d)ove Kninded in fioni, not or only slighily

pKxhitcd bciwecn lhe aniennac 49

10 11 12 13 14 15

Carvai.hc) — Kcys Cicncra Miiiclac of the \V'orkl

48. lk)(Iy vvitii silvcry silky jnibesceiicc intcnnixcd with linc hairs;

libial spincs witii clark spots at base (Forniosa)

CEPIIALOCAPSIDEA Poppius, 1915

— llocly vvitb a single ty})e ol jjiibescence; tibial sjnnes without

tlaik sjjots at Ijase (lig. 88, 235) (Central America)

RANZOVIUS Dhvant, 189.3

49. 1 iltial spincs yellow; sinall species with apex oí' scutclluin,

a sjjüt on corinin and base oi cnneus wbite (África)

TÜRMA Cbina, 1927

— l ibial sj)ines liark, with or withont dark spots at base; apex

oi sciitelhnn and base oi cnneus not white 50

.50. Clypeus with a distinct suture at base; hcad narrow; apex oi

cuueus white (Noith yVmerica)

STROPIIOPODA Van Du/ee, 1921 *

— (dyj)eus without a suture at l)ase, coniluent with irons; head

wide; ape.\ oi cuneus eoneolorous . (Cosmopolitan)

CHEAMYDATUS Curtis, 1833

.51 . Pseudarolia large, reaching to or projecting slightly beyond

ápices oi claws, connccted with tliem or not; disc oí proster-

nal xy[)hus depressed and with elevated margins (ügs.

9, 10) 52

— Pseudarolia minute or not visiljlc, never reaching tips oi

claws; dist: oi prosternal xyphus convex; margins not elevated

(figs. 5, 8) 57

52. Rostrum reaching the middlc coxae 53

— Rostrum reaching the hind coxae or beyond 54

.53. Pubescence oi body yellowish; hemielytra jxde (North Ame-

rica) NICHOIAA Knight, 1929

— Pube.scence oi body black; hemielytra with dark points (Tur-

kestan) SCEODAMIA Poppius, 1912

54. Pseutlaroliii not reaching beyond .ápices oi claws; rostrum

surjxissing the hind coxae 55

~ Pseudarolia reaching beyond the ápices oi claws; rostrum not

surpassingt he posterior coxae (figs. 9, 10) 5G

55. Ilead not strongly j)ioduced aiiterioily; anterior niargin oi

prímotum straight (Rússia) . . . E PllELAS PIA Reuter, i87(i

— lleatl strongly j)roducetl anteriorly; anterior margin oi prono-

tum concave (Kurope, N. ^Miica, iN . y\merica)

AMHEYTYEUS Fieber, 1858

* 1 liis was Iiicnlly l(Min(l lo hc a ,s\iiiin)in ol Chhimydatus Curlis

(aiillioi).

— i —

50 ]5oi,. Mus. Gokudi — J oiiio XI (II) — Dc/,cinl)io 1955

50. Claw clistiiictly toothcd at base; psciuiarolia free, liiscd to

claw only at ba.sc (íig. 10); clypcus strongly piotiiidiiig (Ku-

rope^ Asia, Nortb África) MACRO TYLUS Fieber, 1858

— Claw not tootiicd at base; pseudarolia fuscd to claw iii its

greater or wholc cxtciision (fig. 9); clypcus not protniding

(Europc, Asia, África, N. America) .... I.OPUS llabn, 18.1S

57. Margin of cyc well separated from aiitcnnal fossa, miniimim

sjjacc betwecn tbc two usually more tlian one tliird as great

as diameter of aiitennal fossa; margin of cyc near antennal

fossa alniost straight (fig. 95) 58

— Margin of eyc almost or (piitc tonebing antennal fossa, mini-

mum spacc bctween tlie two not more tlian one ciglitb as great

as diameter of antennal fossa; margin of eye more or less emar-

ginate near antennal fossa (fig. 85) 71

58. Lateral margin of jnonotum widely reflected, sinnate before

the posterior angles; legs very short; witb black pnbescencc

(Turkestan) PLEUROXONOTUS Reuler, 1901

— Lateral margins of pronotinn not reflexed; legs not as

above 59

59. Ilody witb black setiform liairs only (sometimes easily rubbed

off) 1)0

— Hody witb golden or yellow fine pubescence otily or witb seti-

form bairs intermixed witb silky liairs 05

00. Rostrum reaching the mesosternum; pubescence very short;

frons tumid, striolate (Algeria) EUOEROX 1’uton, 1888

— Rostrum reaching the ajx^x ot middie coxae or beyond; pn-

bcsccnce not notir.eably short; frons not striolate 01

01. Rostrum reaching the apex of middie coxae; xypinis ol pros-

terniim imjiresscd at middie, obtnsely marginate (.Sibéria) . . .

IliíAR/S llorvat !k Reuter, 1900

— Rostrum reaching the posterior coxae or beyond; xyphus ol

prosiernum not imirginatc, if so, tlie rostrum re;iching the

5th abdominal segment . .• 02

02. Rostrum reaching the posterior (oxae; xyphus of piosteinnm

with two jiarallel im|)ressed lines at apex (Asia Minor)

ÜTOPNIA Reuter, 1881

— Rostrum reaching tlie Vth abdominal segment or beyond; xy-

pims of prosterimm wilhotit lhe two impiessed lines meniion-

ed above ti.i

03. Rosirmn reacliing ihc genital segment; xyphus ol piosieinnm

(onvex, wiihotit imtassate margins (l.m()|K', .\sia, Norlli

África) PACAIYXYPÍIVS Fiebei, 18.58

Oarvai.ho — Kcys Ciciicra Miridac of llic World

(il,

(i5 .

(ili .

(i7.

(i8.

(ü).

VO.

Rostrum rcacliing at iiiost thc Vth abdominal scgmcnt; xyphus

of prostcrmim plane with margins incrassatc 64

Ilead distinctly transverse, first antennal segnient not rcach-

ing or extending bcyond apex of clypeus, inserted dose to

aj)cx of eye (Asia Minor)

OFISTHÜTAENIA Rcuter, 1901

llead as long as wide; first antennal segment reaching quite

beyond apex of dypens, inserted not dose to the apex of the

eye (Einoi)c, Asia, North África)

THERMOCOlUS Puton, 1875

Rostrum reaching thc apex of anterior coxae or very slightly

beyond: .second antennal scgmcnt shorter than third (fig. 89)

( (F.urope, Asia, Nortli África)

HARFOCERA Curtis, 1838

Rostrum reaching beyond thc apex of first coxae; second an-

tennal segment longer tlian third 66

Antennac with the second joint incrassatc toward thc apex;

genae witliout long liairs (Entope)

CREMNORRUÍNUS Rcuter, 1880

Antennae slendcr or iiuiassate, the second joint narrowed

towaid the apex, linear or so; genae witli long hairs or

setae ^7

Ilody with a single type of pubescence, ])allid, yellowish or

goklen, semiadpressed; first antennal segment dark (Europe)

MOXOSYNAMMA Scot, 1861

Ilody with silky, bright itubescence intermixetl with setiform

haii.s; lirst antennal segment light with one or more blatk

setae

Eairly large hlatk species; rostrum reaching the posterior co-

xae; tibiae blatk (.Sibéria) NYCIIOEA Rcuter, 1901

Species of small or médium si/e, not Itlack; the tibiae light 69

Rostium leadiing distinctly beyond the posterior coxae; head

stiongly pointed in litint

Rostium not smpassing the posterior toxae; head not strongly

pointed in lioiu

l ibial spines with blatk spots at base; secontl segment if hmtl

taisus shortei than thinl (Europe)

U.EOEOEAJiSUS Rcuter, 1885

•l ibial spines without lihu k spots at base; second segment ol

hinti taisus longei than thiiil (Europe. Asia. North Alrita)

MEdAEOCOEEUS Rcuter, 1890

r>()i.. Mus. Cior, 1 , 1)1 — 'loino XI (H) — De/uinliro I !).').')

71. Roslrum rcachiiig tlic posterior coxae; xy[)luis ot prostcrmim

convcx, iiot tarinatc; liiiul tibiac with (lark spincs williout

(lark sjrots at base 12

— Roslrum not rcaching bcyoiul tbc iniddlc eoxac; siilcs oi

xy[)luis distiiictly tarinatc; Iiiiid tiliiac witii dark sjiincs liaving

dark sjiots at base 73

12. Xypluis oi prostcrnuiii convcx and smooth; pubcsccncc not

noticcalily long and dense (Eiirope, Nortb Airica)

UADROPllYES 1’uton, 187i

— Xyj)luis oi jirostcrnum with a incdian carina; pubcsccncc very

long and dense (Algeria) .... DAÜYCAPSUS Popj)ius,1911i

73. .Sniall spccics witii long, slcndcr sccond antcnnal scgincnt; tbc

silky pubcsccncc long and irregular; bcmiclytra with a dark

spot at apex oi coriuni (Europc, Asia)

CAMPTOTYLUS Fieber, 1800

— Usually large spccics; sccond antcnnal scgincnt usually ibick;

silvery pubcsccncc scanty and short; hcniiclira witliout a

blatk spot at apex oi toriuín (Europc, Asia, Nortb America)

ONCOTYJMS Eicbcr, 1858

71. Rostrum ratbing tbc posterior coxae or bcyond ii; ii sborlcr,

tibiac witb blatk spines aml dark spots at base 75

— Rostrum not or surprassing sligbtly apex oi mcsoslcrnum

(cxceptionally rcatbing middlc coxae) 105

75. Hody beset witb silky or woolly pubcscence or ibis type plus

sctiiorm bairs or Irue liairs only 70

— liody beset witb black sctiiorm bairs oídy, witboui silky or

woolly pubcscence 103

70. Vertex distinctly and strongly tarinatc iMislcriorly .... 77

— Vertex not tarinatc j>osterioriy or only very lincly so ... 7'J

77. l’oslcrior iemora witliout rigid ercct sclac ou anterior

margin 78

— Posterior iemora witb several rigid and ercct setae on anterior

margin (EurojK-, Asia and N. Airica)

fSAlA.or.SlS Reuter, 1001

78. Roslrum suipassing sligbtly tbc bind coxae; eyes not reatbing

bcyond anterior maigins oi pronotum; pubesicnce not luiti-

ceably long and still (.\sia Minor and N. Airica)

rAR.lMIXlA Reuter, lOOO

— Rostrum rcatbing tbc apical tbird ol abdômen; eyes reacb-

iiig beyond anterior margins ol pronotum; pubesceme very

long and still ((iuam Is.) RSAl.l.ORS Usinger, 1010

70. Roslium very long, exiending lo tbc 8lb ol Olb abdmnin.il

segment; c.dli well devcloped 80

CIarvai.ik) — Kcys Gcncra i\[iriclac ol the World

5.S

80.

82.

83.

81.

85.

8r>.

'/ .

Rostrum ii surj)as.siiig thc posterior coxac not rcaching the

8tli abdominal segment; calli small, indistinct 82

First antcnnal .segment as long as width of vertex; rostrinn

reaching the last abdominal segment (Eiiroj)e, N . África, N.

America) riNICEPHALUS rieber, 1858

First antcnnal .segment .shorter than width of vertex; rostrnm

not reaching thc last abdominal segment 81

Femora and tibiac with black or colonrcd spots; anxiliary

veins present in membrane (Europc, Asia)

SOLENOXYPHUS Renter, 1875

Femora and tibiac nnicolonr; mejnbranc withont anxiliary

veins (Asia Minor) VOlWCIIlELLA Popin.s, 1912

Tibiac withont distint spines; very small spccies, chocolate

brown with whitish marks at base of pronotum, hcmiclytra,

apex of scntcllnm, corinm and ciinens (Rodriguez Is.) ....

TREVESSA C.hina, 1921

Fibiac with distinct spines; spccies nsnally over 2 mm. long;

(olonr not as above

1’nbe.scence very short, sparse and atlpressed; pronotum, hcad

and sciitellnm ahnost glabrous; hemielytra with a riddish

transver.se faseia on a]»ex of corinm (Euro])e, N. África) ....

MEGAEODACryi.US Fieber, 1858

Rubescence usually longcr and .semierect, if short then reddish

iransverse faseia on apex of corinm abseni 81

Ridtescence of hemielytra composed of briglit silky or woolly

ailpressed hairs intermixed witli black, setiform ones .... 85

Rnbesccncc of hemielytr;i yellowish or dark, bnt never with

two types of hairs 8()

Eegs |)ale withont dark spots or ])oints (Enrope, N. America)

ASC/ODEMA Renter, 1878

l.egs dark or fttstons, if pale with dark spots or points (Noith

America) MEGAl.OPSAl.IAJS Knight, 1927

Ilind tibiac with black spines, thesc s|)ines withont datk spots

at base (sometimes only a laint tlotul, bnt not a delinite

spot)

Ilind tibiac nsnally with ligbt, ycllow oi colottrless spiitcs, il

daik sijines, then with distinct black spots at their bases 87

l’.de giecnish s|)ecies; tibiac with black spines havitig d.nk

spots at base; females brac hypteions, male with sccond anten-

nal segment imtassate (f.niope)

MAEACGTES Renter, 1878

Uoi.. Mus. (ioiiDi — roíiio XI (II) — l)e/cml)i() 1955

— .S|)ccics willi otlicr coloiir tliaii palc ^rccnisli, l)Ut if so, tlicn

fcinalcs inaciopterous and males witii secoiul antcnnal scg-

ment linear and sicndcr 88

88. Ilind tibiae with dark or black .sj)incs, nsually witb dark sj)()ts

at base 87

— Ilind til)iac with ycllow or colourless spincs 90

89. \'ertc.\ finely carinate; niale witli a [jiong on genital seginent;

calli not distiiut (África)

PLAGIOC.N ATI I IDEA Poi)i)ius. 1911

— Vertcx not carinate; inales withont a prong on genital seginent

(Cosmoix)litan) PLAGlOGXATIíUS Ficber, 1858

90. l ibial s|)ines with fuscons spots at base 91

— l ibial spincs withont dark or fuscons spots at base .... 92

91. First rostral segmcnt not reaching beyond base of hcad ((ihi-

na) LEUCODELLUS Rcutcr, 190()

— First rostral segmcnt ícaching beyond base of hcad (Algeria)

COMPSONANNUS Reuter, 1902

92. Femalcs nsually brachyptcrons; sities of pronotum emarginatc

at middle, the calli large and tnmid (Fnropc)

ORTIIÜNOTUS .Stcphens, 1829

— Femalcs always macro[)tcrotis: sides of pronotum not cmargi*

nate at middle; calli small and flat 9.1

93. .Second antennal segmcnt at least in malc strongly incrassatc;

rostrnm not reaching beyond posterior (oxae (Furope, .Asia)

liRAClIYAirrUIWM Ficber, 1858

— , Second antennal segment slender in both sexes; rostrnm reacli-

ing slightly beyond ihc posterior (oxae 91

91. 'I hird segment of hind tarsns as long as lirst and second toge-

ther (Fnrope, N. Alrica) .... PI.ESfODEMA Renter. 1875

— rhird segment of hind tarsns sliorter than firsi and second

togetiier 95

95. First rostral segment not reaching xyphns ol piosteinnm; head

fairly oblicpie (Furope, Asia, N. .\frica)

PUYI.VS llahn, 1831

— First rostral segment reaching abont the b;isal thiid ol xyphns

of piosteiinnn; head fairly vertical 91)

90. Ilannis ol areola and connecling vein cliveiging dose to the

base (Fnrkestan) VORVGUIA Renter. 1879

— Il.nnns oí ;neola and connecling vein not cliveiging dose lo

lhe base 97

97. Head veilical; second and thitcl segmenis of hind laisns snl)-

ecpial in leiigth (diina) ... EI CIIARICORIS Renter. 1900

cm 1

SciELO

10 11 12 13 14 15

Carvai.ik) — Kcys Gcncia Miridae of thc World 55

— llcad subvcrtical; tliird scginent of hind tarsus shorter thaii

sccond (Kiiiopc) ICO DEM A Rcuter, 1875

98. General eolour greenish-ycllow or greenish-white ininute fus-

coiis spois on corium 99

— C.eneral eolour tlark. red or brown to black; if grccnisli then

tlie corium without fuscous spots; pubesceiice not strongly

.silky and uiieven 100

99. Pidjescencc distinrtly silky and imcven; rostrum reaching thc

apex of iniddlc coxae (Indiaii, Tibet)

TIBETOCOlUS liutchiiison, 1931

— Pubesceiice not distinctly silky or uneven; rostrum reaching

a littlc beyond jiosterior coxae (Rússia, 'rurkcstan)

MALTHACOSOMA Rcuter, 1879

1(1(1. General eolour grecnisb yellow 101

— General eolour dark red, brown or black 102

101 . Kyes sliglitly removed from ])ronotum; clypcus strongly pro-

minent (fig. 182) (Europe, Asia)

PAREDRüCOlUS Rcuter, 1878

— Eyes contiguous witli |)ronotum; clypcus vertical, not notice-

ably prominent (fig. 9()) (Europe, Asia, N, África, N. Ame-

rica) TURONIA Rcuter, 1875

102. .Sccond antennal segment slightly swollen at apex, so as to

become as wide as first segment; general eolour dark red

(North America) lUI/NOCARSUS IJhler, 1890

— Sccond antennal segment linear, not so wide as first segment;

rarcly wilb reddish (North America)

M/CRÜPIIYLELLUS Rcuter, 1909

103. Xyplius of prosternum convex, sides not carinatc (Europe,

Asia) PEACÜCIIIEUS Eicber, 1858

— Xyplius of prosternum Hat, the margins obtusely subincras-

sate iOl

KM, Rostrum rcathing beyond the hind coxae; first .segment reaih-

ing beyond the middie of xyplius of prosternum; posterior

tibiae without dark s|M)ts (Gaiicasiis)

DAMIOSCEA Rcuter, 1881

— Rostrum re;iching the posterior coxiie, thc lirst segment not

iiMching the middie ol xyplius of prosternum; ])osterior tibiae

with dark sixits (Einope, Asia, .Mrica)

IIOPEAMACllUS Eicber, 1858

10.5. Pionotinn strongly deilivoiis with two dark spots beliin thc

tali; fairly laige speties (\ . .Miiia)

HOVDAIREA Piiton & Rcuter, 188()

Pionotum not as ;d)ovc '***’

Sfi lioi.. Mus. CioKi.Di — Toino XI (II) — Dezembro 195.*)

106. Vertex very stroiigly rariiiatc, lhe eariíia arniatc siiniatc aiite-

riorly: Iiead traiisvcrsc and vertical, lhe anlennae very tliick

(North America) MYOCÍIROOCORIS Reuter, 1909

— Vertex not carinatc as above; antcniiae slender, or if tbick,

tben pronotnm carinate Ictcrally 107

107. First antennal segnient very tbick; pronotnm carinate latcrally;

rostrnm rcacbing only apex of anterior coxac (fig. 87) (Kn-

rope, Asia, N. África) NASOCORIS Renter, 1879

— First antennal segment not noticeably tbick; pronotnm not

carinate latcrally; rostrnm reacbing l)eyoml apex of anterior

coxae 108

108. .Sides of pronotnm dislinctiy emarginate; eycs sligbtly removed

fiom anterior niargin of j)ronotinn (fig. 98) (Enropc, Asia,

N. África) EURYCOl.PUS Reuter, 1879.

— Sides of pronotnm straigbt or sinuate; eyes contignons wiib

anterior niargin of pronotnm 109

109. Small, strongly shining dark brown species wiili a wbilisli

arca on middle of bemielylra (clavns, corinm, sometimes sen-

tellnm): pnbescence very scanty, snbglabrons; coxac and mid-

dlc of mesosternum wbiiisli (Fnrope, Asia, N. África) ....

AUCIÍENOCRERIS Fieber, 1858

— .Species if dark brown, not strongly sbining and snbglabrons;

wliitisb arca of bemielylra absent 110

110. Fnbescencc black, setiform, intermixed witli fine, whitisli

liairs (sometimes tbe .setiform bairs are present only on sides

of jironoinm and apex of corinm); bemielylra witiiont minute

dark or fuscons spots 111

— 1’nbesccncc fine, wliitisli, no black setiform bairs present 116

111 . Femora with black or fnscons spots I lli

— Femora withont black or fnscons spots 111

112. black spots of femora only on inferior niargin; .scconcl an-

tcnnal segment mncli longer lhan wiclth of bead (.Mongolia)

IdRArroCORlS Remer, 191

— black spots of femora on botii sides; .scconcl antennal segment

sligbtly longer tlian widtb of licacl ll.'!

1 1‘5. Vertex margineci postei ioily, as well as, tbe sides ol pronolnm.

bemielylra very finely |)nnclnrecl (,\sia, Inclia)

RARARAC.MVS 1‘oppins, 1911

— Vertex and pronolnm not margineci; bemielylra wilb dark

and golclen bairs (India) RAC^MVS Distani, 1910

IM. bocly wilb a íew sc-tiform bairs; lostrnm leacbing tbe apex

ol middle coxae; lirsi segment ol bind taisns inmb sboiier

CIakvai.ik) — Kcys (icncra Miridac of thc World

í>i

tlian sccoiul (Asia, Europe)

LEUCOPTERUM Rciiter, 187!)

— Uody piibcsccMice very clcnsc; body witb black hairs (nuiiicr-

ous)o rostnmi rcaching tbc niiddlc coxac (base); clytra willi

sniall dots '1^

115. Heniiclytra with dark or brown .spots; strongly shining; pscu-

darolia rcarbing tbc iniddlc of claw (Atrica)

' PARASCIODEMA Poppius, 1911

— Mcinielytra tvitbout dark or brown spot.s, not strongly sbining;

j)scudarolia ininute (Eiirope, N. Alrica)

TRAGISCOCORIS Fieber, 1801

110. First antcnnal scgmcnt vcry sliort, as long as thc length ol

( lypcus; rcddisb spccics with wbitish arca on iniddlc of claviis

and base of (iincns; licad black ( Fnrkcstan)

EPHIPPÍOCORIS Poppins, 191Ü

— If thc first antcnnal scgmcnt vcry short, colonr not rcdtlish

as above, hcad not black .' 117

117. Externai incisnre of cuncal fraetnre dccp; sccond antcnnal

scgmcnt of malc incrassatc toward a])cx (Clhina)

ECTENEUAIS Reuter, 1900

— Externai incisnre of cnneal fraetnre not dccp: .sccond an-

icnnal scgmcnt of malc linear 118

118. Heniiclytra black or brownish black, base of ennens wiili two

whitish sinits; sccond anlcnnal scgmcnt as thick as or thickcr

than first scgmcnt (Ccylon) SEJANUS Distant, 1190

— llcmiclytra with scvcral miiuitc or dark or orange spots; sc-

cond antcnnal scgmcnt more slcndcr tlian first scgmcnt 11!)

119. First rostral .scgmcnt rcaching only abont iniddlc of cyc; hc-

niiclytra with some mcdiiim si/.cd s])ots (Egypt)

ECyPAGKI.A Schmidt, 1959

— First rostral scgmcnt rcaching xypluis of ])rostcrmini; hemi-

clytra with scvcral miniitc ilark spots

120. llcad proditccd bctwccn bases of antcnnac; cycs not noticc-

ably large; tibiac withont fiiscons sjiots at base ol spines (Eii-

ropc, Asia .Minor, N. África .. PASTOCORIS Rciitcr, 1879

— llcad not proihiccd bctwccn thc bases of antcnnac; cycs vcry

l irgc- tibiac iisnally with fiiscons spots :tt thc base ol sjmics

(Asia’ Minor, North Alrica) . . ATOMOPUORA Rcntci, 1879

I hc following gcncra are not inchidcd in thc kcy:

ncamn.t Poppins, 1915 (Ardi. f. Natnrgcs. 80 A (8): 75);

Ihii iiiosa .

SciELO

10 11 12 13 14 15

58 Boi,. Mus. Goi:i.I)I — 'l oiiio XI (II) — Du/cmbro 1955

Oligobliclla Rcutcr, 1885 (Kiit. Mo. Mag. 21: 201 ; St. Helena.

P/ioe7iicocal)siis Reutcr, 1871) (Pct. Nouv. Kiit. II: 51); Europc.

Tocniopliorus Liniiavuori, 1952 (Ann. Ent. Eciiii. 18 (1): 36);

"rurrnienia (near Icodema Reuter).

Psallomimus Wagner, 1951 (Bnl. Soc. Eouacl I Ent. 35: 119);

Egypt.

KEV IO IIIE GENERA OE DICYPÍIINI

1 . Bocly above sniooth, raiely very finely or snjieificially piiiu ta-

te, more or less slcnder 2

— Body above coiirsely and deeply inmctate, tliick aiul ronnded

(fig- 7-1) Ití

2. Sinall cell oí ineinbrane wilb a distinctly inaiked blaek S|)ot

(África) IIÁEMÁTÜCAPSUS 1’oppius, 19M

— Sinall reli of niendirane williont a dark spot 3

3. Eyes contiguous witli anterior niargin of pronotnni (fig.

157) 1

— Eyes more or less removed froin anterior maigin of jironotnm

(figs. 161, 162) 7

'1. 1’ronotnm constricted antcriorly, tlie anterior lobe jiroduced

forward covóring tlic collar; liead stdrate on vertex; arolia

aiising from tlic claw bnt tonverging ai :ipices (fig. 160) (.Sa-

moa) ÜNCONOTPl.l.US Knigbt, 1935

— 1’ionotinn if (onstiirted ;interiorIy, svitliont a lobe as ;d)ove;

liead not snlcate and arolia not (onveiging at iipices .... 5

5. Rosinim readiing tbe a|)ex of posterior coxae oi beyond (Afl i-

ta) D/CYPIIOPSÍS Popjiins, 1911

— Rostrnni leatbing tlie apex oí middie coxae 6

6. .Seiond antennal segment inciassate tmvard tbe a|K'x; large

tell ol membr.ine almost rcctangniar (Enro|H', .Mrita)

CAMPYI.OiSPl/nOPSIS Poppins, 1911

— .Sttond :inienna segment linear; large cell ol membrane lound-

ed apically (fig. 157) (;\lrica)

CAMPYl.ONEVHA Eieber, 1860

7. líütli sexes biadiypterotis; body lovered witii long and sigiti

seiae; pionottim as long as wide; abdômen eiitiiely ex|)osed

(Ansirali;i) SETOCOHIS Gliina Sc Carvalho, 1951

Botb sexes iii.n io|)terons oi if one sex biadiypteions, ilien thc

.ibdomen paitially (oveied 8

ClARVAi.iio - Kcys Gciiera Miriclac of tluí \VorId

H. Ncck stroiigly coiistrictccl tovvards lhe prouotum, bchind tlic

eycs (África) ORTIIOTILIDEA Poppius, 1914

— Neik noi iioticcal)ly constricted bchind tbe eycs 9

9. Lyes situatcd distinctly in front of tbe iniddle of tbe licad;

tlie ncck very long (África)

MACRÜLOPHIDEA Poppius, 1914

— l'.yes situatcd bchind tbe iniddle of tbe bead or occujjying a

nicdian jiosition 10

19. Cloilar and pronotum very narrow, tbe sides of pronotum ca-

rinate (África) IIYALOSOMELLA Poppius, 1914

— Ciollar not very narrow; sides of jnonotuin not carinate 1 1

1 1 . P.yes renioved froin pronotum by a distance about equal to tbe

tbi(kne,ss of sccond antcnnal segment (fig. 107) (Cosinopoli-

tan) CYRTüPELriS Ficber, 1800

— Kyes renioved from pronotum by ;i distance mucb greater tban

tbe tbickness of sccond antennal segment (fig. 101) .... 12

Kyes very small, .separated from pronotum by more tban Icngtb

of eye scen from above; bead about as long as wide 1.4

— Kyes relatively large, se|)arated from pronotum by a distance

c(|U,d to or Icss tban lengtb of eye seen from aliovc; bcatl

sligbtly wider tban long 11

14. (àmeus more titan tbree times as long as wide; frons not pro-

duced betwecn tbe antennae; first antcnnal segment sborter

tban widtb of bead (fig. 104) (Central America)

Cn/US Distam, 1884

~ Cuneus about twice as long as wide; Irons produced betwecn

tlie antennae: first antennal segment as long as widtb ol bead

(lig. 101) (Kiiroiie, .\mericas, África)

MACROEOPUUS Kieber, 1858

l'l- (iiineus long and narrow, iiboul tbree times longer tban wide

:u base: jironotum not (onstricted at mitldle (Índia)

UiUlAEUS Distam, 1909

Cuneus sligbtly longer tlian wide at base; pronotum (onstiided

ai iniddle

15. \'eitex sligbtly (arinatc at tbe sides; posterior angles ol pro-

l•olum not produced; scutellum witbout subus (lig. 12) ....

(( iosmo|Militan) DICAPIIVS Idcbei, 1858

^eIl(■x (ompleicly smootb; basal maigin ol pronotum stmngly

emaiginate, tbe |M)steiior angles piotluced; siutclium witb .i

longitudinal subus at base (.\lrica)

nUCOIflA Poppms, 1911

(iO

Ifi.

18 .

1 !).

20 .

I k)!.. Mu.s. Gohi.1)i - l oiiio XI (11) - Dezembro 1955

Ncck strongly nanowed behind the eyes; incision of cuneal

iiactuie very deep and wide when the wing is in horizontal

position (índia) ANGERIANUS Distant, 190!

Neck not strongly nanowed behind eyes incision of ciineal

frature not as ahove 17

IHonotuiu with slilning tuhcrcular swellings; scutelluin wiih a

short tubercnlar process (Euro]je, Asia)

STETIIOCÜNUS Flor, 1801

Pronotiim withoiit shining tnbercnar swellings; scntelhnn

withont a tid)ercidar process 18

Eyes distant írom anterior margin of pronotinn, very small 19

Eyes contignoiis with anterior margin of pronotnm 20

Hemielytra and scntelhnn ]ninctate, the latter flat (Mada-

gascar) crCAIROCAPSUS Popi)ius, 1914

Hemielytra and sciitelhim smooth, the latter strongly convex

(fig. 162) (Itorneo) APOLLODÜTIDEA FIsiao, 1914

Rostrnm reaching the apex of posterior coxae; scutclhim

si7iooth (Madagascar)

JIlLDEBRAirriEI.LA Poppins, 1944

— Rostrnm reaching ajjex of middle coxae; scntelhmi punctate

(New Caledónia) TERATÜCARSUS Poppins, 1911

N() I’E: 'Ehe genns Isoproha Osborn ík Drakc, 1915 (Ohio

Nat. 15) is not inchided in this key becausc it coidd not be ]3laced

from its original ilescrijition and no sijecimens werc examined.

Uíibrocoris Wagner, 1951 rnns in this key with Dicypinis Eicber, and

liiicohia Popjjins.

KEY I O THE GENERA OE IlALLODAPIN I

1. Psendarolia large reaching or not tpiitc reaching apex of claw;

sijccies of large size, elongatc (fig. 11) 2

Psendarolia minnte, not as above (figs. 14, 15, 10) 4

2. Second antennal .segment strongly clavate apically, its thick-

ness at a|)ex more than twice that of base; rostrnm reaching

almost to hind coxae; hemielytra fnlly develojied in both sexes

(North Ameiica) rEEEOPIUNUS Uhier, 1890

— .Second antennal segment linear or slightly incrassate at apex;

rostrnm reaching midille coxtie; females brachy])terous or

winglcss .8

.‘5. Second antennal segment linear; psendaiajolia attached at base

ol claw, fiee aj)i(aily; females wingless (North America) ....

COOUIEEETTH Uhlei, 1870

cm i

SciELO

10 11 12 13 14 15

Carvalho — Kcys Gcncni Miriílac ol tlie World

1 1 .

.Scco.ul amennal scgmeiu sligluly incrassate at apex; pscudaro-

lia coinpletely joiiitcd to daw; tcnialcs hrachypteroiis (Nortli

America) ORECTüDERUS Uhler, 187()

I Icmiclytra with sharjily delimited whitish or ycllowish white

tiansverse íasciac or sjiots or elongatc markings; seldom im-

ilormily pale biit then long and erectly pilose <)

I Icmiclytra withoiit sharply delimited whitish or yellowish

white transverse fasciae or spots or elongatc markings; on the

t laviis and on the eorium there is seldom a very small ycllow

transverse spot, this same spot however on the clavus lies fav

in Iront ol that on the coriíim and is postcriorly bordered by a

ileep velvety black elongatc spot 5

llody above with long, ereet and black bristles; gronnd colour

dirty greysh ycllow (África)

TRICEIORHOREELA Reuter, 1905

botiy above usnally with short, ercct hairs or il these are dark

then the dorsiim black 6

Dorsimi with short, ereet dark bristles (yVfrica)

lUIiUN DIELEA Po|)pius, 19M

Dorsiim withoiit short, ereet dark bristles 7

Dorsinn totally black, clavus and coriíim with a small ycllow

transverse s|X)t (África) SYNGONUS llergroth, 192()

Dorsitm not totally black 8

Dorsiim with short, adjiressed, yellowish piibescence; pronotum

not longitudinally sulcate at middle ol base (Índia)

A/JZUS Distant, 1910

Dorsnm with short, ereet, whitish bristles: pronotum with a

longittidinal sulctis at middle ol base (North yVlrica)

AEOEOCORES Reuter, 1901

1’rottotum distitictly ptinctate (lig. 71) 10

Pionotum smooth or very finely rugouse 1 ‘5

Utiderside long and erectly pilose; tibiiie with lotig bristles 11

l/tiderside tiot lotig and arecty jiilose; tibiae withoitt bris-

tles 12

Piody above lotig aml erectly pilose (llra/il)

IMA/.OiXOCOR/S Carvalho, 1952

Piody above tvith short atui ailpiessetl pidiesceme (lloliviti)

' EUC.EREEEA Poppius, 1921

Twc) last antemial segmetits thinncr th;in secotid; pionotum

widei than long, the lateial margins straight; scutelhun with-

otit a basal lobe (Alrica) EYEOPEETIS Retiter, 1901

Two htst antemial segmeiits abotit as ihiiuk as seioiid; jiro-

iiotum aliout as long as witle; the lateial margins rouiuled;

cm i

SciELO

10 11 12 13 14 15

02 lioL. Mus. CioEi.Di — loino XI (II) — Dc/cinbro 1955

sciitelluin with a basal blimt lobo alinost as high as pronotiiin

(Sonialiland, logo) GLOSSOPELTIS Rcutcr, 1901

l.B. Sciitelluiii of male with a hlgb, crect spinc likc projcction, as

high as Of highcr than pronotimi (fig. 213) M

— .Scutcllmn of nialc unarnicd, sonictimes strongly convcx of

tuniid, if so, ncvcr as high as disc of proiiotum 17

H. Anterior portion of pronotuin with two crect sinnelike jno-

cesses (fig. 81) (índia, Malay, Horneo)

NICOSTRATUS Distaut, 1901

— Anterior portion of jjronotnin without the above inentioned

processes 15

15. .Scntellar s])inc directed forwards; lostnnn reaching apex ol

iniddle coxae; antcnnae inserted near the eye (North America)

HFJDEMANNIELLA Poppins, 1911

— Scntellar s])inc directed backwards; antcnnae inserted lar froni

the eye (fig. 229) 10

10. Rostrnm reaching apex of anterior coxae; .second antcnnal seg-

ment not strongly incrassate; scntellar spinc short ('rinkes-

tan, África) ASE/DACANTílUS Renter, 1901

— Rostiinn reaching posterior coxae: secoiul antennal seginent

strongly incrassate towards apex; .scntellar spinc long (fig. 229)

(África) MYOMBEA China & Carvalho, 1951

17. Eycs distam from the pronotiim by a s|)ace abont e([iial the

width of vertex (fig. 81) 18

— Eyes contiguous with the pronotnm or renioveti froin it by a

distance abont etpial to half the width of vertex (brachyptcr-

ons femalcs) or mnch less (inales) (fig. 80) 19

18. Eyes pilose; disc of pronotnm strongly convcx posteriorly; he-

mielytra long, reaching distinctly beyond apex of abdômen

(fig. 81) (índia) SOUENUS Distant, 1910

— Eyes glabrons; disc of pronotnm not noticeably convcx poste-

riorly; hcmieytra only covering apex of abdômen (Entope,

Asia, África) AÍ.EOEOMIMUS Renter, 1910

19. Antennae very densc, short and adpres.sed pnbesccnt with a few

intermixed bristlcs; sccoiul segment incrassate towaitis the

apex were it is thicker than first; body with short, stont, creci,

black bristlcs (fogo)

CIIAETOGAESUS Pojipins, 1911

— Antennae with a single tyjie of inibescence; body withont short,

stont, crect black bristlcs 20

20. Pronotnm strongly constricled and narrowed on anterior half;

posterior leiimia incrassate apitally, posterior libiac cnrved in-

' Carvalho — KL‘ys Ciciiera Miridac of thc World

()3

ternally (fig. 211) (Madagascar)

MALGACHEOCORIS Carvalho, 1952

— Pronotuni not as above, femora not noticcably incrassate at

apcx 21

21 . Froiis projccting in írom into a tidjcrcular conical process co-

vcring part of tbc laminatc (strongly coinpressed) clypeus;

Icngth of projcctioH aboiu cqual to tliickncss of first antennal

scginein (fig. 2.86) (África)

ACRORRHINIUM Noualhicr, 1895

— Frons roundcd, flat or convex, witboiit a tubercular conical

jtrojcction 22

22. Antennae Inscrlcd aboiit thc iniddlc of cyes (África)

P AN GANI A Poppius, 1914

— Antennae in.serted bellow or about the inferior third or levei

with apex of eye 28

28. .Scutellnni sbort and narrow, with a j)it shapjted depression at

base, bebind the iinpression soniewhat swollen; feniales bra-

cbypteroiis with elytra sharp and transverscly flattened at base;

heatl as wide as ba.se of ])ronotinn, both didl, the hemiclytra

strongly shining (Madagascar)

MYRM/COPSELEA Poppius, 1911

— Scutelhim without a })it like de|)rcssion at base 21

24. Genae higher than width of one eye; bead distinctly ant like,

with a raised carina in front of apex of eye (Eiiropc)

MIRMJCOMÍMUS Reuter, 1881

— Genae inuch ess high than width of one eye; ape.x of the latter

not contiiuied by a carina, bead not noticcably inodificd 25

25. lleinielytra witb longitudinal oblique white lines or faseia;

large s|)ecies about 5 inni long or more 26

— lleinielytra without longitudinal oblitpic whitish lines or

faseia; .sjiecies nuuh sinaller in si/.c 27

26. .Second antennal seginent linear; juonotum strongly narrowcd

in front (North America)

CLOSTEROCORIS Uhlcr, 1890

— .Secoiul antennal segment incrassate towards the ajtex; prono-

tum not notiteably narrowcd in front (Etirope)

CREMNOCEPIIALUS Fieber, 1861

Vertex maiginate or tarinatc, sometimes angulate iiostcriorly,

pronotum strongly tonvex ou disc and declivous 28

\5'itex not marginate or carinate; jironotum not noticcably tle-

divous 28

27 .

1)01,. Mus. Cioi.i.i)! — 'l‘om() XI (II) — Dc/cnibro lí)55

28. Ilcail vcry largo, as vvido as with ol jironotum at base; oycs

Occaipying tlio wliolc sidos ol lioad, roaohiiig llio gula liolow

(Pomba Is.) liüOrJDELLA Roíilor, 1907

— lload not vory largo, iiarrovver lhan widih ol proiiotiim al baso;

oyos not roatliing tlio gula bolow 2t)

29. Collar vory wiilo, otpially lovclod and arcliod with pronotinn,

vory sliglitly dojnossod, soparato írom disc by a laint linc; pro-

notum long, narrovv, tho sidos strongly roíindod (Aírica) ....

DIOCORIS Kirkaldy, 1902

— Collar distinctly .sot oll and doprossod, sidos ol pronotum not

noticoably ronndod ilO

30. líody with vory long oroct pulrosconcc intonnixod with short,

adprossod silvory wooly hairs; liind tibiao íairly thick, with

vory long and slondor spinos (Alrica)

TRIClIOriITlIALMOCAPSUS Poppius, 1911

— P>ody with a singlo typo ol pnhosconco; hind tibiao il thick

ihon lho spinos not vory long and slondor 31

31. lloatl soon Iroin Iront distinctly longor than wido, gula long,

horizontal upjior lip largo, coll ol moinbrano with righi or

straight apical anglo (Alrica)

SYSTIÍLLONOTIDKA Popius, 1911

— lload soon liom Iront not or slightly longor, usually shorlor

than wido; gula short, il long not horizontal, uppor lip sinall.

apical anglo ol moinbrano roundotl 32

32. l ibiao with spinos at loast apically 33

— l ibiao without spines (North Alrica)

C.LARIIYROCORIS Routor, 1903

33. Kyos largo, olongato, distinctly produtod boyond sidos ol collar;

hoacl strongly inclinod, Hat antorioi ly (Kuropo, Not th zMrica)

MI MOCO RIS .Scott, 1871Í

— l'.yos small, roundod, not noticoably |M()tlutod outwards; hoacl

not strongly inclinod, Irons roundod 31

2'1 . .Scutolhun strongly tumid (maio) or with a spino liko ])rocoss

(lomalo); pronotum vory strongly doclivous 3ã

— .Sciitcllum Hat or convox on both soxos; pronotum not strongly

doclivous (Kiiiopo, Alrica, Índia)

IIAI.LODARVS l•■iobor, IHtiO

3.ã. I hird and lourth antonnal .sogmoni ihickor than .socond; seu-

tolluin with a spinilorm jnotuboiaiuo (.Alrica)

IIYROMIMUS l.indborg, 1910

- rhiid and louilh antonnal sogmonls moio slondor than socond;

sculoll.u pi otidioranc c' bhmi (l.nropo, ,\sia. .Mrica, )ava) ...

I.AEMOCORIS Routor, 1879

Carvai.iio — Kcys Gcncia Miridac of thc World

Í56. Dorsum jndjostciu 37

— Dorsum glahrous or only very scaiity pubcsccnt 39

37. Pubescciicc vcry short, dcnsc and atlprcsscd (North África) . . .

FORMICOPSELLA Poppius, 1914

— Pubesccnce crcct, scMiiiercct, not noticcably short 38

38. Eycs contiguous with j)ronotum (África)

SYSTELLONOTOPSIS Poppius, 1914

— Eycs rcinovcd from prouotuui (Europc, Asia, África)

SYSTELLONOTUS Eieber, 1858

.39. Cdypeus proiuincnt, visible from above; hcad as wide as width

of ba.sc of pronotum (Europc, Asia, África)

OMPffAEONOTUS Reutcr, 1876

— Clypcus not noticcably proiuincnt and visible from above; hcad

not as wide as base of jnonotum (Madagascar)

LISSOCAPSUS licrgroth, 1903

NO TE: riic genus Trachelnnotus Reutcr, 1901 (Ann. Mus.

7ooI. St. Petersb. 9:8) from Persia is not includcd in lhe kcy. Mimo-

W^agner (Egy|)t) runs in this kcy with Glapliywcoris Reutcr

.jtnd Mimocori .1 Scott.

KEY TO EllE GENERA OE IIALTICANI

1 . Erons vcry proiuincnt, roundcd and convex; posterior femur

vcry thick, its largcst diameter about ccpial to width of onc

clytra; botii sexes brachyptcrous; small spccics with rcddish

marks (llawaii) NES/DIORCflESTim Kirkaldy, 1902

— Erons not noticcably prominent; femora not as thick as width

of clytra: if bolh .sexes brachyptcrous, rcddish marks absent

fig. 210) 2

M. Anlcmiac vcry long and sicndcr, setond segment four or more

times as long as first segment; brachyptcrous forms conunon,

with oval body, strongly convex (fig. 210) 3

— Antcunac shorter, setond .segment littlc more than three times

as long as first segment or much shorter 4

.3. Eycs removed from anterior margin of pronotum; vertex

siraight postcriorly, hcad sccn from from jicntagonal (Ghina)

ECTOME/IOPTERUS Reutcr, 1906

— Eycs (oniiguous tvith pronotum; vertex somcwhat arcuatc ])os-

tci iorly; hcad sccn Irom from not pentagonal (fig. 210) (Cos-

mopoliian) ILILTIC.VS llahn, 18.32

1. Eycs disliiu lly pedmu ulalc. thc vertex vcry wide (fig. 170) 5

— Eycs noi pctíum ulatc, sometimes substylate 6

— 5

Of)

Boi.. Mus. Goei.di — 'roino XI (II) — De/cnihro 1955

5. Pronotum riigulosc piinctatc, vcrtcx carinatc, tlic carina ar-

cuatc; rostnini reaching base of middle coxac (North America)

LABOPELLÁ Knight, 1929

— Pronotum not piinctate; vertex witliout an arcuate carina; ros-

trum reaching apex of middle coxae or beyond (fig. 170) (Eu-

rope, Asia, North America) LAliOPS Burmeister, 18,‘{5

f). Body glabrous, distinctly punctured; both sexes brachypterous,

first antennal segment with stout black bristles and sbort pu-

bescence (Europe, Asia) .... EURYOPIC.ORIS Reuter, 1875

— Body not glabrous and pn neta te, if so, tbcn one sex macro-

pterous and first antennal segment with a single type of jni-

bescence 7

7. Body, especially bead and pronotum, with long stiff black bris-

tles; lengtb of liairs of third antennal .segment iisually more

tban twice as great as thickness of segment 8

— Body without long stiff black bristles; antennae with mucb

shorter pubescence 17

8. Both sexes brachypterous; elytra with whitish flattened or scale-

like pubescence amongst the bristles 9

— Males macropterous, females brachypterous; hemielytra with-

out flattened or scalo-like whitish puliescence or if so, no bris-

tles present but only adpressed pube.scence 12

9. First antennal segment with very short bristles and scale-like

pubescence; femora with many thick and stout hristles (Euro-

pe, Asia)w AN APUS Stal, 18.58

— First antennal segment with hristles and common pubescence;

femora with only a few or without stout :ind thick bristles 10

10. Hemielytra not reaching beyond the second abdominal seg-

ment; hind femora with numerous fine bristles 11

— Hemielytra reaching beyond second abdomimd segment; hind

femora with a single seta (Asia)

PEATYPORUS Reuter, 1900

1 1 . Hind femora with only one type of long and fine bristles; body

with dense whitish flattened |)ubescence and bristles (Fur-

kestan) SCI RTETEELUS Reuter, 1891

— Hind lemora with a lew long bristles mixed with very short

ones; body with very sianty silky pidiesience and (ommon bris-

tles (Spain) nOMOEOCORIS Fuente, 1918

12. Pubescence of body coniposed oI Hat whitish scale-like hairs

mixed with hairs or bristles; seiond antennal segment about

as thick as first 15

cm i

SciELO

10 11 12 13 14 15

(Lvrvai.ik) — Kcys Gcncni Miriclac ot thcv Workl

— Pubesccncc of bocly composed of onc typc only, or silky or

woolly inixcd with cominou haiis; sccond antennal segmcnt

slender tlian first 14

]‘5. Amennac with vcry short inibesccncc; vertcx somewhat ciirvcd

jiostcriorly; rostnim rcaching middlc of abdômen (North Áfri-

ca) ORANIELLA Rcntcr, 1894

— Antcnnac witb Inistles and bairs; vertex straiglu posteriorly;

rostrimi shorter (Enropc, Asia, N. África, U.S.A.)

ORTIIOCEPHALUS Fieber, 1858

M. Mcmielytra witli single type of semi-adpressed hairy jnibescen-

ce; bead as wide as base of pronotnm (Europe, Asia, N. África)

PAC.HYTOMEELA Reuter, 1890

— llemielytra witii silky bairs mixed with common ones; head

narrower tban base of jnonotnm 15

15. Males witb ciineiis more tban three times as long as wide;

memlnane and areolae elongate; eyes snbstylate (Europe, Asia,

África) DIMORPFIOCORIS Reuter, 1891

— Males with cuneus aboui twice as long as wide; eyes uot sub-

stylate 16

16. .Small black species; rostrum reaching apex of posterior coxae

(male.s) (Europe) SCHOENOCORÍS Reuter, 1891

— Médium size greeuisli species; rostrum uot reaching bcyond

apex of middle coxae (males) (Europe, Asia, N. África) ....

PEAGIOTYEUS .Scott, 1874

17. Male witb vertex strougly carinate and depressed, frons separa-

te Irom clypeus by a semicircular ridge; rostrum uot rcacliing

middle of mesosternum; fcmales brachy|)terous, piceous bluisb,

punctate (fig. 8,S) )Europa, yVsia)

PIEZOCRANUM Horvatb, 1877

— Male witb vertex and fions uot as above; rostrum longer, reacli-

ing miildle (oxae or beyotid; females macropterous 18

18. Hody distiiKtly punctate; head uot jiroducetl in middle of

antennal bases; cuneal fracture uot deep and wide 19

llody smootb; head proilmed iu miildle of antennal bases;

small species witb detiduous pubescence; (imeal fracture wide

and deep (India) STUENARIDEA Reuter, 1884

Embolium wideued towaril tbe apex; body with erect jmbe.s-

<ence; seioud auleimal segment about twice as long as head;

females with hemielvtra eularged apically (Índia)

.' ICRATIIEUS Distam, 1910

Emholium narrowed toward the apex; body glabrous or with

semi adpressed pubescence; sccond antennal segmcnt shorter or

slighily longer than head; females with hemielytra uot eidarged

19 .

f)8

Jk)L. Mus. Goei.di — 'loino XI (II) — l)c/cinbro 1955

apically (Europc, Asia, N. África, and N. America) ....

STRONGYLOCORIS lilanchard,

1840

NO'rE: The genus DASYSCYTUS Fieber, 1861 (Wien. Ent.

Monat. 8:84) íroin .Spain, is not included in Key, as well as 11 AL-

riCÍDEA Reiiter, 1901 (Oív. F. Vet. Soc. Forh. 45:172), Rússia;

and SARONA Kirkaldy, 1902 (Fauna Havv. 7(2):142), Hawaii.

KEY J'0 THE GENERA OF ORTllOTYLlNl

1 . .Second, third and fourth segments of antennae incrassatc, about

cípial in thickness (fig. 102) 2

— Second segment of antennae ihicker tban third or fourth

(fig. 10.5) 3

2. PronotuiTi anterior to iniddle nearly cylindrical, ratlier

abruptly flaring behind middle, basal half of disc strongly

convex; eniboliar inargins stdcate on basal half (North y\me-

rica) PAMILÍA Uhler, 1887

— Pronotuni rcgularly narrowcd anteriorly, its sides tiot cons-

tricted at niiddle; eniboliar niargins not sulcate (North aiul

.South America) C.ERATOCAPSUS Reuter, 1875

3. Scutellum elevated and swollen, curving cystiformiy forward

over the disc of iironottun, biconstricted, with a small erect

dorsal spine in front of the anterior constriction (fig. 257)

(Australia) CYSTEORRACllA Kirkaldy, 1907

— .Scutellum not as above 4

4. Pronotum dislinctly ptinctale, if jitinctures are obscured by rti-

gosities then hemielytra jninctate (fig. 74) 5

— Pronotum smooth or rtigose 14

5. Hemielytra witbout an ajijiarent cuneus; simill cell of mem-

1)1 ane faintly delineate or not visible (Hawaii)

SULAMITA Kirkaldy, 1902

— Hemielytra with a distinct ctineus aiul small cell of mem-

brane 6

f). Small species with head as wide as j)ronotum at ba.se; frons

striolate finely i)uncttired; antennae very short; margins of

pionotum carinate, a pseudocollar present (Australia)

COR/DROMÍUS Signoret, 1862

— Frons not punctate or striolate; i)ron<)tinn not carimtie

laterally 7

7. Glypeus strongly |)i()minent: eyes seen Itom Ironi Ihittened and

somewlKit pedunculate; scutellum strongly |)rominenl, raised

al middle; body wilh short and deiise adpiessed pubescence

(Ibiwaii) KAEAiVEl Kirkaldy, 1901

cm i

SciELO

10 11 12 13 14 15

Carvai.iio — Keys Gcnera Miridac of thc World

— Clypeiis iiot noticcably proininent; eycs sccn Croin front not pc-

dimcidatc; srutclluni Hat or convcx, not raiscd at middlc; body

pubcsccnce not noticcably sliort, dense and adprcssed .... 8

8. Dorsuin tbickly clothcd with scmidecnmbcnt pnbcsccnce; ver-

tcx strongly dcclivous antcriorly, carinatc; colonr back (North

America) LO Pl DELL A Knigbt, 1925

— Dorsnm with crcct |nibc.sccncc or alinost glabrons; vertex not

as above 9

9. Hemiclytra sinootli, transparent; calli with two dec|) fossac

bchind ihcni; body with fine, long and crect |)nbcsccncc

(brazil) SOLANOCORIS Carvalho, 1915

— lleniielylra ])imctatc; calli withont two fossac bchind thern;

body withont long, fine aml crcct pid)csccncc 10

10. First and second antcnnal segments very thick, thc sccond seg-

incnt flattcncd, third and foiirth segments very short and slcn- ■

der (África) MILLERIMIIUS Carvalho, 1951

— F'lrst and second antcnnal segments linear and not noticcably

thick 11

11. Dorsal snrface cvcnly pnnctnrcd, a |)nbcscent hair arising

from eadi punetnre 12

— Dorsal snrface punctured biit withotit a pnbesccnt hair arising

from each pnnctnre 13

12. Cnncns veiy small, wider at base than long; secoml antcnnal

segment incrassatc towanls tbc apex; rostrnm reaching the

postei ior coxae (Samoa)

PSEUDONEORORUS Knigbt, 1935

— Ciinens abont twice as lotig as wide at base; secoml antcnnal

segtnent linear; rosirnm reaching thc middlc coxae (África)

liUNSUA Carvalho, 1951

13. Hemiclytra distinctly ronnded latcrally; eycs ronnded, some-

what removed from pronotnm (Venezuela)

EALCA)NrODES Rentcr, 1905

— 1 lemielytra more or less straight latcrally; eycs straight pos-

teriorly, (ontignous with pronotnm (fig. 108) (Central &

Sonth America) EALCONIA Distant, 1881

M. First aml sccond antcnnal segments conspienons, thc sccond

strongly enlarged, compressed or foliacens; thiid and fonrth

very sitort and slender; black species (figs. III, 230) .... 1.5

— First and second antcnnal segments not noticcably inoilified, or

if so, then second not foliacens 18

15. First and second antcnnal segments with flattcncd hairs; frons

prodnced between :intenn;ie (Fnrope)

I IEEEROEOMA l.epeletier .Servillc, 1825

cm i

SciELO

10 11 12 13 14 15

70 lloi.. Mus. CloKi.Di — 'l oino XI (11) — Dcztnibro 1955

— First and sccond antennal scgnicnts withont llattened hairs;

IVons iiot produccd (fig. 230) 10

10. Mend)iane apically acutely pointcd (l'miisia)

ACRODERRUIS liergioth, 1911

— Mcinbranc not acutcly pointcd at ajjcx 17

17. Vcrtcx carinatc; roslrum rcaching hasc ol iniddlc coxac (Áfri-

ca, Ccylon) DRUTIIMARUS Distam, 1909

— Vcrtcx not carinatc; rostrimi rcaching base of abdômen (fig.

Ilt) (Fiiroj)c, Asia, .South America)

EXCENTRICUS Reuter, 1878

18. 1’ronotuin with plcural arca sc]jaratctl froin dorsal part by a

distinct suture, jrronotal disc raiscd jrostcriorly and jtrojccting

above scutclliun, clothcd with dense, bristiy pubcsccncc (fig.

100) (North America) SEMI UM Reuter, 1875

— Pronotum witiiout a distinct lateral suture antl not projecting

postcriorly over scutclluin 19

19. IJody with distinct scalc-likc or llattened pubescence interiuixed

with hairs or bristlcs or dcnscly covered with whitish flattcnerl

somewhat scale-like hairs specially on uuderside 20

— liody clothcd with a single type of pubescence or someliines

interinixed with silky hairs 32

20. llead without a wcll defined posterior niargin 21

— llead with a well defined posterior niargin 23

21. llead rounded in front, sccond antennal seginent incrassale

towards the apex; pronotum constricted anteriorly with raiseil

calli; dark species with jiale arcas; antminiic (Kuro]ie, .í\sia,

North America) .. (UJ)lilCACRS I.e 1’elletier & .Serville, 1825

— llead uoi rounded in front, sccond antennal .seginent cylin-

drical; pronotum if constricted anteriorly then calli not raiscd;

sjiecies usually with grccnish color or if dark then not

autinimic 22

22. llead noticeably produccd in front; ciy])eus very large and

proininent; species usually over -1 nnn. long with norinal le-

niora (North America) .. ARfiVROCOR IS Van l)u/ec, 1912

— llead uoi noticeably iiroduced in front; cly|)eus not as above;

species usually less thaii 1 mm. long, with enlarged |)osterioi

femora (North America) RARTIIENICUS Reuter, 187ii

23. Tibiae with hhuk spots at base of s|)iues; body (overed by

very dense whitish llattened hairs on the uuderside .... 21

— I ibiae without black s|)ots at base of spines; uuderside of body

with (omnion pubescence 25

Carvalho — Kcys Gciicra Miriclae oí the World

2-1. Femora with black spots or if not tlicn color greenish; eyes

romuled (Nortli America)

PSEUDOPSALLUS Van Duzee, 1916

— l'enu)ra without sjjots: general color black; eyes strongly com-

pressecl (Europe) IIYPSELOECUS Reuter, 1891

25. Secoiul antennal segmeiu thickened at apex or if not, then

very black species (fig. 104) (Enrope, Asia, Nortli America)

IIETEROCORDYLUS Fieber, 1858

— Second antennal segment linear or so; never very black

spocies 26

26. Ciineus rounded externally, cnneal incisiirc deep; pronotinn

carinale laterally; redilish species (.Scychelles Is.)

MARALAUDA Distant, 1913

— Cnnens not as above (except brachyj)teroiis Ibrins); pronotum

not carinate 27

27. Claws deejjly cleft witli inner half wider; head inclined and

distinctly prodnced before bases of antennac (North America)

RIFIDUNCiULUS Knight, 19.30

— Claws not divided, head not noticeably prodnced in front 28

28. Vertex very wide; íirsi antennal segment about etpial to half

widtli of vertex: bristle like jjubescence very fine and erect;

females brachypterous or almost so (North America)

LAIIOPIDEA Uhler, 1877

— Vertex not noticeably witle; first antennal segment distinctly

longer than half the widlh of vertex; bristle like pnbescence

short; females macroj)terous 29

29. Rostnnn short, .scarcely atlaining hind margin of mesosternum;

large, elongale fnscoiis species 5 to 6 mm. long (North Ame-

rica) NOCTUOCORÍS Knight, 1923

— Rostmni reaching the miildle coxae or beyond; species nsutilly

greenish in color 30

30. First antennal .segment shorter than width of vertex; scalc-like

pnbescence usually silvery (Euro|x;, North America)

MEEANOTR/CAJUS Renter, 1875

— First antennal segment as long as or longer than witdth ol

vertex; scale-like pnbescence mostly biit not necessarily

black 31

31 . Rostrnm reaching lar beyond ápices of hind coxae: cly|)eus very

laige, nsnally, wider than ihiíkness of first antennal .segment

(North America) MACROTYEOIDES Van Dnzee, 1916

— Rostnnn not reaching beyond hind coxae; clypens not large,

nsnally as wide as or narrower than first antennal segment 32

72 líoL. Mus. Goki.di — 'l oino XI (II) — Dc/cnibro 1955

32. líristlcs blnck witii or silvcry scalcs bctwccn; proiiotuni wilh-

out blac scalc-likc sjiots; first antcnnal scginciu iiuub longcr

tbaii wicltb o[ vcitex (Nortb America)

ILNACORELIA Kniglit, I92&

— llristlcs liglit witb black scalcs bctwccn; pronotiim vvitb black

scalc-likc s|)ots; first antcnnal scgincnt altoiu as long as width

oí vertex (Nortb ík Central America)

ILNACOIIA Renter, 187(>

35. First antcnnal segment very tbick, aboiit as long as hcatl anil

pronotum togetber; ciincns abont fonr times as long as wide

at base, somcwliat ciirvcd ontwards cxtcrnally (fig. 228) (Áfri-

ca) UELKANA Carvalho, 1951

— First antcnnal segment if long, not noticcably tbick; enneus

not as above 31

31. Eycs roíindcd bebind anil set in front, at or ncar middlc of

bcad, nsnally wcll removed from anterior margin of pronotimi

by a spacc cqnal at Icast to tbickncss of first antcnnal segment,

long cnnens (figs. 114, 155) 35

— Eyes rclaiively straigbt bebind, contigiions or .set very dose to

anterior margin of pronotum (figs. 117, 118) 54

35. Veitcx depressed at middlc; areolae of membrane scleiotizeil

as cnnens, ap])arcntly witb a .sinijjlc cell; male witb :i conspi-

cnoiis antenna (first and sccond segments witb iwo long s|)incs,

tbe sccond aiso witb a medinm fossa snrrounded by small dark

spines) as in fig. 00 (Nortb, Central & Soutb America)

ÍIYAÍ.OCIILORIA Renter. 1907

— Vertex convex; aieolae not cbitini/ed or if so witb two distinct

cells; male antennae not as above 3(5

30. Eirst antcnnal segment sborter or abont as long as widib of

vertex; if not, tben cnnens 2 times or more as long as wide at

base 37

— First antennal segment dislinctiy longcr tban widtb of vertex,

nsnally as long as or longcr tban widtb of bead witb eycs 40

37. Cialli witb two deep fnrrows bebind tbem containing ])nnt.tnres;

bead witb a sbort neck; eyes removed from pronoinm by a

distance eipial to more tban balf tbe Icgtb of one eye O'*'

niaica) MESOEROEIS Renter, 1907

— Calli witbont two deep fnrrows witb pnnctnres bebind tbem;

bead vvitbontb a sbort neck 38

Ilead very flat, wider or abont ;is wide as ])ronotnm at b;ise,

clotbed witb silky silvcry pnbescence; pionotnm rei langnlar

(lig. 107) 39

Ilead not very Hat, it wider tban pronotum at base, tbeti witb-

ont silky pnbcscetice 10

38 .

Carvalho — Kcys Cíciicra Miiiclac of tlie World

- 10 .

•II

39. Internai niargin of eye straight; rostrum rcaching apex of inid-

dle coxac (Australia) COMPSOSCYrUS Reuter, 1009

— Internai niargin of eye strongly divergent; rostrum reaching

sligluly beyond anterior coxae (fig. 107) (Eurojie, Nortli

Afrira) PLATYCRANUS Fieber, 1870

Head vvitb a sliort neck; eyes sitnateil more towtirds anterior

end ol head; body fairly long and ereetly pilose (fig. 155) -11

Head without a sliort neck; eyes sitnated more towards base of

head; body not long and ereetly j)ilo.se (fig. 110) -12

Clypens seen from above; eyes very small, head elongate; lirst

antennal .segment with a black faseia, inicriorly (Euro]ie, North

Afriea, North America) MALACOCOIUS l-icber, 1858

— Clypens not seen from above; eyes not very small, head loimedd

(fig. 155) (Índia, Formosa, Ceylon)

/.ANCIIIUS Ditsant, lOOT

-12. First segment of antennae with a longitudinal black line on

eilher side, these lines coimected on ventral side near apex

(Kurope, North America) lUiVTERI A Puton, 1875

— First segment of antennae not marked with longitudinal black

lines as above -13

Flongate sleiuler s|)ecies; cuneus twice or more as long as wide

at base -11

Species if elongate, cuneus shorter -15

F.yes small, placed at middie of head, usually distam Irom

lironotum by more than diameter of lirst anienna; il this dis-

tance ecpials diameter of segment (male) then hemielytra reach-

ing lar beyond tip of abdômen (Américas)

PA RA PROBA Distam, 1883

— F.yes large, placed on posterior iiortion of head, usually not

more removetl from ])ronotum then thickness of forst antenna;

if this distance etpials diameter of segment (male) then henie-

lytia not as above (fig. 203) (North America)

D! AP! IN 11)1 A Uhler, 1895

•15. .Setond antennal segmem about as thicks as the lirst; membra-

ne (clls membranous (Central & .South .\merica, Jamaica) . .

.SAI LERIA llsiao, 1915

— .Setond antennal segment dislincily more slemler than the lirst;

memhrane tells toiiateus ((.entrai N Noilh Ameiita)

PI.ATY.SCYTU.S Reuter, 1907

•1(). Rosii um leaihing the middie ol the abilomen oi beyond .. -17

— Rositum not leathing beyond lhe posterior toxae 50

-13.

■ 11 .

cm 1

SciELO

10 11 12 13 14 15

7‘1 Hoi.. iMus. Goki.di — 'l oiiio XI (II) — De/cnibio 1955

47. Pronotuin stroiigly coiistrictcd antcriorly; clavii.s with a row

(>f puncturc.s; ciincus alinost four tiincs as long as wide at base

(Madagascar) MADAGASCARIELLA Carvalho, 1955

— 1’ronotuin iiot strongly constricted aiUeriorly; clavus without

a row ol jnuictiires; ciiiieiis shorter 48

48. Vertex witli a straight carina posteariorly, the eyes blutnly mar-

gined behiiul; body rvitli esparse, long, erect and fine pubcs-

cence (fig. 115) (Central & Sonth America)

JOIiERTUS Distam, 1884

— Vertex without a straiglit carina bcliind or if so, eyes not

inargined posteriorly; body not noticeably long pilose (fig.

110) 49

49. Pronoluni bisinuate posteriorly; eyes distant froin pronotum

by an esjxice ctpial to thickness of first anteniial seginent (fig.

IIO) (Ihazil) RRAS/I.IOMIRIS Carvalho, 1940

— 1'ronotuin straigth posteriorly; eyes sitnated in front of the

iniddie of head (fig. 114) (Ihazil)

ITACORÍS Carvalho, 1947

50. 1’ronotinn strongly constricted at niiddle; claval vcin witli a row

of piinctures; species noticeably elongate and slender with

tiansparent heinielytra (África)

rElJSACODKS Ikrgroth, 1920

— 1’ronotnm not strongly constricted; claval vein without j)nn-

ctnres 51

51 . lJp()cr lip very thick, inllaled, dense and shortly pidrcscent;

firsi antennal scgment abont twice as long as the head (Ca-

nary Is.) AETORRl UNEEl.A Nonalhier, 1893

— Uppcr lip noi inflattcd or pnbescent; fiist antennal e<]nal or

slighlly longcr than the head 52

52. First scgment of antennae longer than head .seen from above;

rostrnm reaching slightly bcyond ]K)sterior (oxae; eyes small

((icntral America) RARACAIIUS Distant, 1881

— First antennal scgmen t as long as head seen Irom above; ros-

Irinn reaching the jrosterior (oxae; eyes large 53

53. Head strongly narrowed behind the eyes; pronotum with two

snblateral furrows leathing posteiior margin of c;dli (Formo-

sa) y.ONODOROrsIS Poppius, 1915

— Head not strongly nairowed behind the eyes; pronotum with-

oiiti the two liiriows above ()amaica)

MEEANOSriCTES Reuter, 1907

51. A wi ll delined oblitpie sntme on gena extending Irom antennal

lossa to bene:ith eye, this .sntnre (lig. 231) fre<inently ontlined

5(i.

l)/ .

r.8.

5!) .

(iO.

Carvalho — Keys Gcnera Miridac of thc World

by a dark stripc; rcd orange and black spccies (Nort and Cen-

tral America) LOPIDEA Uhler, 1872

Genal sntnre absent or extending directly from antennal fossae

eyes or present, bnt vagne and not ontlined by a dark

sti

ipe

Pronotnm strongly declivons, clypeus oblicjne, situated beneath

liead; eyes very large; body with silky piibescence; rostrnm

reacliing sligluly beyond anterior coxae (lig. 212) (Argentina)

HYPORRHINOCORIS Reuter, 1909

Pronotnm not strongly declivons, clypens vertical, situated in

Iront of head; eyes not very large, body with other tyjre of

l)nbescence or if silky, then rostrnm longer 56

Species mormorate (as in Plty tocaris)', rostrnm reaching middle

of abdômen; ])osterior femora flattened (also as in Phytocoris);

abdômen long an.l erectly pilose (Hawaii)

KAMEHAMEHA Kirkaldy, 1902

Species not marmorate or if so, rostrnm sborter and jjosterior

femora not flattened; abdômen not noticeably long jrilose 57

1 leatl strongly prodnced in front of eyes, tbis sjxicc being aboiit

twice as long as length of eyes; antennac inserted for from the

eye, the space between tbem eqnal to or sligluly over the

tbickness of firtl segment; head somewhat hori/ontal, body

glabrons (Hawaii) PSEUDOC.LERADA Kirkaldy, 1902

Head not as above, antennae inserted mnch closcr to the eye;

head vertical or strongly inclined 58

Small pale species with both sexes nsnally bracbypterons; mem-

brane and cnnens absent; pronotnm tra|)e/,ilorm (Enrojje) . . .

EIEIiEROCAPSUS Carvalho & Sontbwood, 1955

Species macropterons or if one sex bracby])tcrons, then cnnens

or membrane present; pronotnm not as above 59

Vertex willi a distinct raised carina at posterior margin (fig.

112) 60

Veilex williont a distinct raiseil (arina at posterior margin

lig. 56) 77

Carina extemling from eye lo eye and bearing erect bristles

(fig. 259) 61

Calina if extending from eye lo eye witbont stonl black bristles

or ered setae (bristles may be present on vertex) 65

fiist antennae aboiil as long as widtb of Itaed indiiding eyes;

greenisb (< lotired .sitecies (Ktirope, Asia, Norlb Aliica)

lU.EPIEÍRIDOPTERUS Kolenati, 1915

l'iist aiilennae sltorier llian oi as long as witltli of vertex 62

SciELO

10 11 12 13 14 15

70 Hor.. Mus. Goei.di — Tomo XI (II) — Dczcmljro 1955

02. Pronotiim clistinctly narrowccl and constrictcd in Iront, calli

promincnt; spccics witii antmimic coloiiration (Euiopc) ....

DRYOPini.OCORIS Reutcr, 1875

— Pronotum not noticcably constrictcd in Iront; spccics witiioiit

antniintic colouration 0.5

0. 5. Rcddisli or ycllow bnt rcddisli inarkcd spccics; body noticc-

ably long ant crectly pilose; hcmiclytra dnll, not trans-

I)arcnt Od

— Otbcrwise colorcd, somctinics with traces of brick rcddisli, if

long, fine and crcctly pilose, thcn thc bcniiclytra trans-

parent 05

01 . Prons soincwliat protriiding bctwccn bases ot antcnnac, wbich

lias bristies and some crcct sctac (Eiiro])c, Asia, Índia)

RSEUnOI.üXüRS Kirkaidy, 1905

— Erons not protrnting bctwccn bases of antcnnac wbich has

bristies only (Ccylon) TIIERMUS Disttmt, 1909

05. Ilcmiclytia glassy, transparent, lotig, esparse and crectly setose;

pronotnm strongly cnrvcd postcriorly ,fig. 109) (.South Ame-

rica) PLINÍELEA Hcrgroth, 1922

— Hcmiclytra not glassy or tratispaictU; jironotnm not strongly

cnrvcd jiostcriorly 00

00. Chccn to ycllowish spccics; jironotnm not carinatc latcrally

(North Atncrica) LAIiOPÍDEA Uhicr, 1877

— Dark sjiccics, sometimes with traces of reddish; jironotiim ca-

rinatc latcrally 07

Íi7. Pronotnm flattcncd latcrally with tlic sides strotigly carinatc;

bristly jnibcsccncc verv long (Mcxico)

.'.... IIADRONEMIDEA Rcntcr, 1909

— Pronotnm not fhittcncd l.itcially only slightly caritiatc; bristly

jitibcsccncc fairly short (fig. 112) (North Atncrica)

líADRONEMA Uhicr, 1871

08. Vertex with a shallow longitudinal snhtis; hcad hot i/.ont;il;

an .S-.sha|)cd sinooth ridgc bent forward arising from thc jicri-

trctnc jircsctit (Formosa) .. ZONODOREEÍAJS Po|)|)ins, 1915

— Vertex not longitndinally sithatc; an .Sshajicd ridgc arising frotn

jicrilrcntc absent 09

09. Rostumi rcaching thc ajicx of antetior coxac; (tinctts ahont

as long as wide at hasc 70

— Kostititn icitíliing thc middlc of mcsostci imtit ot hcyond;

(imcits tisttally longci than wide 71

Ilcmiclytia glahroiis, shining; pionotinn smooth (Gciiitiil ík-

.Siitith America) JORXAXDES Distant, 1881

70 .

(jARVAi.Ho — Kcys Gcncra Mirídac of tiie World

— llcinielytra pubcsccnt; pronotum sonicwliat nigosc (Hawaii)

KOANÜA Kirkaldy, lOOÍi

71 . Frons and vcitcx strongly dcclivous, thc latter soinewhat dc-

jjicsscd, carina Iiigh with sliarp cdgc; body sbagrinc, ahnost

glal)r()us; nicinbranc noüccably long (Mcxico)

FICANUS Distant, 1893

— Frons and vcrtcx not as abovc or it so, thc carina low and

blnnt; body not sbagrinc and mcinbranc not noticcaldy

long 72

72. Vcrtcx with a l)Iack. fossa cach sidc ncxt to tlic cyc; with

striatcd black niarks on frons (Enro]JC, North África)

UYOIDEA Rcutcr, 1876

— Vcrtcx and frons withont thc abovc inarks 73

73. Ilcad strongly dcclivous, pointcd bclow; cycs contiguous with

])ronotai anglcs; first antcnnal scgincnt abotit as long as hall

thc width of vcrtcx 71

— Ilcad not strongly dcclivous; cycs slightly rcmovcd froni pro-

notal anglcs; first antcnnal segmcnt abont as long as vcrtcx 75

71. .Arolia large, broadnctl toward thc apex which is trtincatc (fig.

69); si)ccics olivaccons in color (Malay)

PARASTIIENAIUDEA Millcr, 19.37

— y\rolia sinall, tapering loward thc apex which is pointcd; spccies

nsnally black to ridcscciu (Micronesia, Philippines, .S. Ame-

rica, Pnerto Rico) ORTUO EYLELLUS Knight, 1935

75. Carina of vcrtcx arenate postcriorly; spccics over 7 min. long,

with rcscnd)lancc to Cyllccoris and CAohiceps (Korca)

CAMRYLOrROPÍS Renter, 1901

— Carina of vcrtcx not arenate postcriorly; .sjhxícs Icss than V

nnn . long, withont icscinblancc to thc gcncra abovc nicn-

tioned 76

76. .Sccond atitcnn;il scgnicm abont three times as long as third;

genital segmcnt of males with a median projcction dirccted

backwaidly (Canary Is.) .. CANA R/()C()RIS Lindbcrg, 1951

— .Second antennal .segmcnt abont twicc or less as long as third;

genital segmcnt of males withont a vctitral |)roje(tion dirccted

badwaiils (Ftirope, Noith America, África)

ORTIIOTYI.US Fichei, 1858

77. Rostiinn reaching apex of anterior coxae or slightly bcyond 78

Rostnnn reathing bcyond middie ol mcsosternmn /9

Pionotnm not noticeably narrowed anteriorly; .sccond segmcnt

ol antenna linear; slender, elongate greenish spccies (Fnro|)e)

nRACJIYNOrOCORIS Renter, 1890

78 .

78 • lk)i,. Mus. (ioKi.Di — l oino XI (11) — Oc/cinbio 1955

— Proiiotum narrowcd aiitcriorly; sccond aiUcnnal scgmcnt

sliglitly iiicrassatc; spccics black and rufescciu (Central Ame-

rica) RUINOCAPSIDKA Renler, 1908

79. Clypeu.s .sirongly eoinpres.sed and proininent, ronndish in

front, secn froin above distinctly protlnced betvveen antennac;

l)()dy witii long erect and e.spar.se brisllc.s (íig. 154) (Sontli

America) CYRTOTYIÁIS (krgroth, 1922

— Clypeu.s not as above or if so, tbe body rvitbont long erect and

esparse bristies 80

80. Ilody witb silvery silky pnbescence intermixed vvitb line erect

liairs; tibiae witb long siiines baving dark spots at base (AIrica,

Cientral ík .Sontli America) RLLENIA Renter, 19-10

— Hody witb a single type ol pnbescence 81

81 . First antcnnal segmeni shorter tban widtli of vertex; eyes con-

tignons witb anterior margin ol pronoinm 82

— First antennal segmem longer tban widtb of vertex or if tiot

tlieti eyes somewbat removed from atiterior margiti of pio-

tiotinn 8.‘[

82. 1 lead poitited in front; body fairly jjilose: veitex smootb; se-

cottd anteimal segtnent less tban 5 times lottger tban first seg-

ment (Cirecce) AMIX/A Renter, 1885

— Head tiot jointed in frotit; body coveied witb sbort liairs;

vertex longitnditially snlcatc; secotul titUennal segtnetit 5 titnes

lottger tbati first segtnetit (índia) .. ASERYMl/S Distant, 1901

8.5. 1’rotiotnm tiotieeably tiairowed iti frotit, cotistricted, calli pio-

minenl; species witb atitmitnic colonratioti; female antenn:il

scgmetit clavate (Knrope, Asia)

CALLECORÍS Ilalni, 18:M

— Pronotnm tiot noticeably cotistricted and tiairowed in front;

ctlli not protnitietit; sjiecics witbont anttniniic colonratioti 81

81. Fyes contigiiotis witb atiterior margin of pronotnm; rostritm

leacbitig apex of mesosterninn (Kttrope)

RACAlYl.ors Fieber, 1858

— Fyes sligbtly retnoved frotn piotiotittn; rostnmi readiitig middie

ol posteiiot eoxae 85

85. Rostnini reacbitig tbe middie (oxae; protiottnn witb a sletider

atiterior (olbir (F.hiojm-, Asia, Nortb .Mrica, Nortb .Atnetita)

MECOMMA Fieber, 18.58

— Kostrntn leadiitig tbe posterior eoxae; pronotntn witbont an

tinterior (ollar (Costnopolitan)

CYRTORII/NUS Fieber, 1858

CIarvai.ik) — Kcys Gcnera Miridac oí tlic VVorkl

KEY ix) riiE c;enera oe pilopuorini

] . Frons witli a sj)iniforni |)rojction bent downvvards ovcr base oí

clypcus wíiich is coniprcssccl; first rostral seginent not leacbing

base oí Iiead; proiiotuni and abdômen very strongiy constricted;

species cxtreniely inynnicoinorjjliic (Ceylon)

LUTIIERIELLA Poppius, 1913

— Frons witbont a spinilorm jnojection 2

2. Scutelhmi with an erect sjjiniform process (fig. 213) .... 3

— Sciitelliim flat or conve.x, without a spiniform process .... 5

3. Pronolum dense and finely puncturcd (África)

ÜPYSTOCYCLUS Poppius, 19M

— Pronolum not |mnctured 4

4. .Sccond antennal segmeiu abouth as thc thick as third (North

America) CYRTOPEETOCORIS Reiiter, 187.3

— .Sccond antennal scgment distinctly thicker tlian third (Ar-

gentine) MYRMECOZELOTES Bcrg, 1884

,3. Hcad much higbcr than long or wiile; eyes substylate rising

distinctly abovc levei oí vertex; antcnnac inscrted far írom

eyc, tbis dislancc being about etiual the eight oí cyc; botli scxes

Itrachypterous (Euroj)e, Asia)

MYRMECOPIIYES Fieber, 1870

— 1 lead not as above; eyes not substylate; antcnnac inscrted

nearer to the eyes (1

(). Anterior (oxac with a slrong tuberclc at hase; pronotum

strongiy constricted and narrowed on anterior third, thc sides

carinate anteiiorly (Madagascar)

EUCOMPSEl.LA Poppius, 1914

— yVnterior coxae without a strong tuberclc at base; pronotum if

c:onstrictetl not carinate anteriorly 7

7. Chda strongiy carinate; upper lip as long and wide as lirsl

rostral segmenl; head (vertex and frons) very depressed, suiik

bellow eye levei (fig. 80) (Chile)

DOEICAIOSTEN/A Poppius, 1921

— (hda not strongiy carinate; upper lip smaller, not as wide as

first rostral segmenl; head convex oi Hat, not uotice*d)ly tle-

pressed on vertex ^

Pronotum stiong and transver.sely rugose; rostrum rcaching

apex ol middie coxae (Austrtdia)

‘ KIRKAEDIEEIA Poppius, 1921

Pionotum not rugose or only slighily so 9

SO 1U)I.. Mus. (iOKI.DI

roíiu) XI (11) — l)c7cmbro 1955

9. Sccoiul antciiniil scgiiiciu as tliick as tliird, or louitli; fenialcs

usually bracliyptcrous or with niodiliccl hciniclytra (short

mciubranc) (lig. 102) 10

— Scioiid aiitcnnaí scgineiu tliickcr tlian tliiril and íourtb; tcnia-

Ics usually iiiacroptcrous (lig. 103) 19

10. Vertex strougly caiiiiatc, tlic carina as liigli as tliiclkuess of

sctoiid aiitcnnal scgincut at base; scutclluin strougly tuinid;

frons striolate; bciuielytra wilh .soincwhat rugose surfacc (Aíri-

ca) NICIIOMACIIUS Distant, 1901

— Vcrtcx carinate iiot as abovc; scutcllum not noticcably luniid;

írons iiot striolate 11

1 1 . Ileiuiclytra with .scalc likc liairs or transversal silvery scalc likc

Ijubescent baiuls 12

— 1 lemielytra witbout seale like bairs or silvery baiuls 14

12. llemielyira with long erect bristlcs aiul srale likc bairs; pro-

notuin not strougly narrowed in iront 13

— lleinielylra with .seniiadpre.ssed pubeseence plus scale likc bairs;

jironotuni strougly narrowed in Iront (Bolivia)

LEPinOTAENIA Poppius, 1921

13. Pronotuin eovering inesoscutinn aiul part ot seutelbnn (Cen-

tral Nortb America) liENODAEUS Distant, 1893

— Pronotuin not eovering niesoscutinn (Nortb America)

Pn.OPllORüPSIH Poppius, 1914

14. Pronotum noticcably constricteti at middle; tiliiae usually some-

wbat curved; only females known 15

— Pronotum not noticcably constricteti at middle; tiliiac straiglit;

males known 20

15. Anterior portion ol jironotum witb two or tinee ttibercies,

erect and pointed (Vene/uela)

ZANCIUSMK Kirkaidy, 1901

— Anterior portion ol pronotum witbout tidierclcs 10

10, Pronotum not deeply (onstricled at middle, linely punctate,

sbining; bemielytra eovering tbe abdômen (Intlia)

y.AHATVS Distant. 1909

— Pronotum deeply (onstricted at middle, smoolb or il puiutate,

tbe bemielytra not reacbing apex ol abdômen 17

17. llemielytia distiiutly puiutate at base, reatbing beyond tbe

posterior loxae (.Mrita) l.ASIOMI MVS Poppius, 1911

— llemielytia smootb, uot leatbing beyond tbe posterior

toxae 18

Postei ior lobe ol pronotum (ouically piodiued upwards; eyes

elongate aud obliipie on bead (Nortb .Mrita)

lORMlCOCOniS I.indberg, 1910

18 .

Carvalho — Keys Cencra Miriilae of the World

lí).

. 20 .

2 \ .

22 .

Z‘5.

1 ' 1 ,

Posterior lobc ol proiiotuin not eonically produced upwards;

cyes roimded (Aiistralia) . . MYRMECORIDEA Poi)Jmus, 1921

Vertcx not coinprcssed posteriorly, neither overlapping ante-

rior edge ol jrronotinn 20

Verlex noticeably coinjnessed and carinate posteriorly, iisually

overlapping slightly anterior edge ol j)ronotinn (lig. 261) 22

Length ol lirst antennal segment shorter than width ol head;

pronotum not eonstrieted anteriorly (Anstralia)

LEUCOPÍJORÜETERA Poppius, 1921

Length ol lirst antennal segment abont etpial the width ol

head; pronotum distinetly eonstrieted anteriorly 21

Seiitelhim strongly tninid, elcvated; cyes distant Irom prono-

tuni by an esj)ace greater than the thiekncss ol lirst antennal

segment (North America) .. CYEllOPELTA Van Duzec, 1910

Seiitelhim eonvex not noticeably prominent; eyes contiguous

with anterior margin ol pronotum or so (lig. 262) (North

America) PSEUDOXENETUS Renter, 1909

Memielytra eonstrieted and recurved ventrad, bcaring white

or silvei y ])ubeseent htinds '. 23

Memielytra withont white or silvery j)iibescent baiuls .... 21

Vertex and Irons very Hat, laintly snleate longitnilinally; vertcx

straight jrosteriorly; inembrane long; s])eeics noticeably elon-

gate (Chile It Argentina) .... TUXENEEEA Carvalho, 1952

Vertex and Irons not snlctite longitiidinally, roímilctl or not

noticeably dej)ressed; veitex eurved jjosterioily towards the

pionotinn; meinbrane short; speties not noticeably elongatc

(l'.moj)e, Asia, Alrica, N. .Vmerica)

PIl.OPIIORUS Westwood, 1876

llody ahnost glabroits; lemalcs noticeably dimorjthic with head

strongly roimded in Iront :ind j)ronotum (onstritted anteriorly;

small species .3 mm long or less (Paragiiay)

JIAELODAPOIDES Carvalbo, 1951

llody distinetly itnbescent; lemales not strongly ilimorjthic;

s|)ecies nsnally over 3 mm long 25

.Setond antennal segment .starcelly ihickeneil towards lhe ajtcx;

width ol head e(|nal or larger than base ol pronotum (North

Ameri(a) ALEPIDIA Renter, 1909

.Seconil antennal segment incrasstite towards the apex; width ol

head le.ss than width ol posterior margin ol pronotum (North

Ameti(a) Al.EPlDIELLA Poppius, 1911

Pionolimi and síiitellmn linely piindate (.\igeniina)

I.AEMOCORIDEA Po])pius, 1921

. fl .

cm i

SciELO

10 11 12 13 14 15

82 Hoi,. Mus. Goki.di — l onio XI (II) — Dezembro 19.55

— Pronotum anel sciUellum not punctate (Américas)

SEIIICOPHANES Reuter, ISTC)

NOTE: l he following genera wcre not inclndecl in tlie key:

Antliropopítagioles Kirkakly, 1908 (roc. Linn. .Soc. N. S. Wkiles

3.^:378) from Fiji.

KEY EO 1 IIE GENERA OE MIIIINI

1 . U])per wing withont tnnens and membrane, lhe divisions into

coriínn, clavus and emboliiim not distinct; .secoiul antennal

segment long and clavate at a]jcx; bracbypterons (Eiiroj)e,

zVsia) APIÍANOSOMA Costa, 1811

— Upper wing with (iineus and membrane, tbe divisions into

corinm, clavus and embolium distinct 2

2. First antennal segment very tliick, its diameter being about

cqnal to that of one eye, consti icted ai miildle; secoinl segment

strongly incrassate on apical balf; third and lourtli very sbort,

togetlier snbeepial to second; clypens strongly |)rominent, licme-

lytra witb palcbes ol silvery pnbescence (Central Atncrica)

(íig. 202) EUROTAS Distant, 1881

— y\ntennae not tis above 3

3. Alltel ior tibiac strongly enlarged and ílaltened, Ibliacens or

■so; ciineus about ;is long as wide at base (Cential & .Soiitli

America) 1 lENICOCNEM/S ,Stal, 1800

— Anterior tibiae cylindiical, not as above; if enlarged apically

tlien (iineiis longer tban wide 1

'1 . Hody above smooth, sbagrine or rngose, raicly laintiy |)nnc-

tulate, is tbis case, tbe lirst segment ol liiiul tarsi very long.

about as long as tbe two last ones togetlier or tbe lorae strongly

carinate (fig. 199) 5

— líody above distinctiy pnmtaie (pronotum). tbe pnnctnres

rarely 1'aint, in tbis case, tbe snriaie deeply rngose or witb

esparse siale-like bairs or silky and silvery pnbesieme, tbe

bead pointed and longly ptodnted 77

5. l.orae very strongly develo|)ed and (arinate 0

— l.orae il developed, never carinate 7

0. liody long, large and elongate; ciine.d incistire small, sballow

(AIrica) l.liWOCEROCORIS Karseb. 1892

llody lairly small, roimded; cnneal iiuisnre deep (.Miica.

Eernando Po) EAM PROCA PSIDEA Poppins, 1912

Carvai.iio — Kcys (icncra Miritlac of tlic World

7. Posterior femora very long, extending iiuicli bcyoiid tip of

altdonicn and flattcncd, broadest belorc middlc and tbcn ta-

})cring gradually lo a])cx or if not flattcncd, tbcn cnrved on

itppcr snrfacc 8

— Posterior femora sborter, not or scarccly extending bcyond

tip of abdômen, not noticcably broad at base II

8. Pronoinm sid)marginate latcrally at apex; Itead more or Icss

borizontal and |)ointed: claval vein distinctiy raiscd (Enrojjc,

Asia, North Afriea) MIR/DIUS Fieber, 1858

— Pronotnm not marginate; licad vertical not pointed; claval

vein not raiscd 9

!). Ilind femora cnrved on npper snríace, witb a fevv spines on

|)osterior side; pronotnm strongly dcclivons (fava)

EUPUYTOCORIS Poppins, 1914

— Ilind femora not cnrved on npper margin, witiiont spines;,

jnonotnm not stiongly dcclivons 10

10. Pemora not notiteably narrowed towards ajiex; calli rcacliing

sides of pronotnm: female bracbyplcrons (Asia)

PIIYTOCORIDEA Renter, 1905

— Femora llaltencd and noticcably narrowed towards tbc aj)cx;

(alli not leaching sides of pronotnm: lemalcs macropterons

(Cosmopolitan) PÍIYTOCORIS Fallcn, 1811

I 1 . Fiist antennal segment tbickcned, cloted tvith nnmcrons ílat-

tened baiis (lig. 1138) (Nortb & (lential /Vmerica)

NEUROCOLPVS Renter, 1875

— First antennal segmeni devoid of llattencd liairs 12

12. Piononlm witii two snbext iivated, dnll bbick spots sitntited

bebind tlie c .illosilies (lig. 207) 13

— Pronoinm wiihoni ilie spots above or witb only superficial

ones above lhe <lisc 15

13. Fiist antennal segment large, strongly (ompressed laterally

(foliacens) (lig. 21) (Américas)

LAMPETUVSA Distant, 1881

First antennal segment (ylindrical, not noticcably ati-

lei ged bl

11. Fiist aniennal segment ilotlied witb long black Iiairs; vertex

(on\i'x, Iroiis snioolh; rosirnm reailiing lhe lasl coxae (y\me-

'EA EDI A Distant, 1883

l iisi antennal segment (lolhed with short liairs and two or

ilnee selae; veitex depressial, fiotis siriolaie: rostriim readt-

ing lhe 8th alidoniiiial segment (.Mrici)

OXACiaORIS Renter, 1905

Mus. Goki.di — 'ronio XI (II) — Dc/cnihro 1955

15. líody sliagrinc and marblcd; heiniclytra witii soincwliat scale-

likc liairs; anterior tibia with an apical tnft oi bairs internally

(fig. 199) (South America) . . GUIANELLA Carvalho, 19‘1()

— llody not sbagrine and inarbled; il scale-like bairs present

tlien anterior tibia witliout an apical tiift oí bairs IG

IG. .Second antennal seginent at least on males noticeably incras-

sate towards tbe apex, somewbat spindle sbaj)ed (tig. 197) 17

— .Second antennal segment linear or only very sligbtly incrassate

at or towards tbe apex (lig. 219) 27

17. Head borizontal, clyjjens abnost reacbing a|)ex ol first anten-

nal segment; jngum distinctly set oü Irom lonnn, Irons jjoint-

ed; first rostral segment reacbing only levei of middle of cye,

wbicb is removed írom anterior margin of pronotiim; first

tibiae enlarged and flattened, body witb j)atcbes of silvery

silky bairs (África) .... TRACIIELUCHUS Bergrotb, 192G

— Head not as above or if so, tben first tibiae not enlarged and

body witbont patebes of silvery silky bairs 18

18. Body witb adpressed silvery pnbescence; first segment of bind

tarsi nuicli longer tban second (Australia)

PSEUDEUliYSTYLUS Po])pius, 1915

— Body witbont adpressed silvery ])id)escence or if so, tben a

second type of bair or bristle present; first segment of biml

tarsi not longer tban second 19

19. Claviis witb two rows of piinctnres; eyes set at middlle of

bead; antennae and legs very long (Madagascar)

INOSlIiEA Carvalbo, 1953

— Clavns witbont two rows of jtnnctures; eyes closer to collat

20. 1’ronottnn witb black setiform bristles intermixed witii golden

or silvery pnbescence; second antennal segment gradnally in-

crassate towards apex 21

— 1’ronotnm witb a single tyjte of jtnbescente or il not, tbe

second antennal segment very slender at base and incrassate

oídy ajtical balf or so 23

21. First antennal segment large, compressed, witb two types of

pidtestence (Asia, África, Kast Indies, Pat ific Is.)

EURYSTYEUS .Stiil, 1870

— First antennal segment cylindrical, witb one lype of pnbes-

22. .Senteibim stiongly laised ilistally; (iinens abont as Iniig as

wide at base; lirst antennal segment abont as tbick as a|)e.\

ol second segtnenl (Noitb .Vmeiita)

RYCNOCOItlS V’an l)n/ee, 1911

Carvai.iio — Kcys Gcncni Miridac ol tlic World

— Sciitclluin modcrately convcx; cimeus loiiger than wide at

base: first anlciinal scgmcnt not as trick as aj>cx of second

segincnt (Américas) NOTHOLOPUS IJcrgroth, 1922

23. Hody strongly sliining; pronotum crcctly and hemelytra ad-

presscd pilosc; rostnim rcaching middlc of mcsostcrnum (Ma-

dagascar) SCHOUrKDENIELLA Poppius, 1912

— llody not noticcably sliining; pronotum and hemelytra with

samc typc of pubescence 24

24. Pronotum finely nigonse, almost glabrons, body totally black

above (North America) .... ECTOPIOCEliUS Uhler, 1890

— Pronotum not nigonse, pnbesccnt, body not totall black

above 25

25. Verlex sulcate in middle; rostrnm reacliing tbe middle coxae;

second antennal scgmcnt tlnckest at apcx 26

— Vertcx smooth; .second antennal scgmcnt tliickcst on apical

tliird; rostrnm rcaching thc posterior coxae (fig. 179) (Amé-

ricas) GARGANUS Stal, 1858

26. Pronotum constrictcd antcriorly; thc anterior portion much

lower than posterior; jugnm very large and iiromincnt ('l au-

ria) EPIMECEÍ.Í.US Rentcr, 1894

— Pronotum not constricled antcriorly; jugnm not noticcably

large and prominent ('rasmania)

PSEUDOPANTILIUS Rentcr, 1905

27. First scgmcnt of hind tarsi distinctly longer tban tbird (fig.

23) 28

— First scgmcnt of t*'»'*' ''«t longer than third (fig. 29) 31

28. Large spccics with cuneus long and pointed, about threc

times as long as wide; head and jironotum covered by black

setiform hairs, thc latter faintly rugosc; lornm very small

gena prodiiccd aiiically soincwhat tubcrcular, touching apcx

of jugao (lUirma)

CllEILOCAPSUS Kirkaldy, 1902

— .Small or médium si/.c s])ecics with shorter cuneus; hody with-

out setiform hairs or if so, then gena not as aíiove .... 29

29. Ilead much longer than wiile, distinctly pointed in front,

somewhal hori/ontal (índia) .. ZAí.MUNNA Distant, 1909

— Ilead about as long as wiilc, not pointed in front, vertical 30

30. Fyes slightly removed from |)ronotum; second antennal seg-

ment thinner than first (Lurope, Asia, Alrica, Índia, North

.America) SPENO PlhS (akovlcv, 1877

— F.yes contiguous with pronotum; second antennal scgmcnt as

thick as lirst (Alrica) LYGOPSIS Poppius, 1912

80 líoi,. Mus. CioKi.Di

ronio XI (II) — Dc/cinhro 11)55

31. Dorsal siirfacc distinctly jjubcscciit, diill 32

— Dorsal siirlacc glabrous, liigldy j)olislictl, somctiiiics nigosc or

roiigli witb esparsc, ícw and sliort liairs 71

32. Hcinclytra covercd by dark rccuinbcnt hairs plus scalc-likc

liairs; cunciis aboiit as wide as long, mcinbraiic very short,

liemclytra rounded cxlernally (New Zcalaiul)

CIUNAMfJilS VVoodward, 1951

— llcinelytra witb a single tyjie ol hairs or il scale-like liairs

present tlien cnnens longer tliaii wide, ineiiibranc long and

lieinelytra not ronndcd e-\lernally 33

33. First antennal seginent as long as or longer tliaii liead and

jironotnni togetlier; Iroiis horizontal sliglitly protrnding over

liase of clyiieiis; rostrnm reatliing tlie loiirtli visible abdominal

scgnient (Nortli America) .. PALLACOCOIllS Renter, 1870

— Fhrst antennal segmeiu not as long as lieail and jironotinn

togetlier or ií so tlien rastrinn sliorter and Irons not as

above 31

31. Collar very wide, abont, 1-1/2 times as trick as lirst antennal

segment; eyes renioved Iroin pronotmn, piomineni; ilark sjie-

ties witb liiteons markings, tlie emboliinn iisnally llavoiis or

yellowisli (Fin()])e, Asia, N. África)

CAPSODES Dahibom, 1851

— Collar not noticeably wide: if tlie eyes renioved Ironi jirono-

tnni tlien tlie licad witb a sliori neck 35

35. liody covered witb still black .setifoini brislles; bead witb a

sbort neck; eyes sligbtly renioved froin anterior maigin ol

])ionotnm; legs and aiitennae witb many brislles; roslrinn

reacbing tbe niiddle coxae (aspect ol Dicylflitis) (lig. 188)

PAMKIiíni-A Renter. 1900

— body witb otber type ol piibescence or il setiform bairs pre-

sent, tbe eyes (ontignons to pronotmn and bead witliout a

sbort neck 30

30. 1’ronotnni <;ninate at sides; vertex snbate (lig. 192) . . 37

— 1’ronolmii not larinate lateially 38

37. IJody covered by sbort setilorm black brislles; rostrnm reacb-

ing beyond niiddle coxae; vertex slndlowly snicate (.\sia,

Nortb America) ALLüllIlINOCOniS Renter, 1870 *'

— body (overed by sbort setilorm black and adpressed brislles

plns woolly silvery bairs; rostrnm not leadiing tbe midille

(oxae; veitex deeply snbate (lig. 192) (lúirope, Asi;i) ....

PAiWTlíAUS Cnriiss, 1833

* l he ficiim A Imiililius Kblliliciilo, liljl runs lo Ihis toii]>lct of tlic key.

Carvamio — Kcys Gcncra Miridae of thc World

38.

39.

40 .

41.

42.

43.

41.

45 .

40.

47 .

Hcmelytra witli black sctiform liairs and silvcry silky pubes-

ccnsc (thc bristlcs soinetinies seen only on exocorium) . .39

Hcniclytra with a single typc of pubescence 43

Rosirum not reaching bcyond thc apex of hind coxae . . 40

Rostrimi reaching bcyond thc apex of hind coxae 42

Vertex sulcate longitndinally; rostrinn reaching the apex of

hind coxae (Asia) PARAPANTILIUS Rcuter, 1904

Vertex sinooth; rostrnm reaching thc apex of middle

coxae 41

First and sccond antennal segnicnts linear or so; tibiae com-

j)ressed and carinate on njjper niargln with strong black

spines, the posterior tibiae distinctly ciirved (lindia)

LUCITANUS Distant, 1904

First and sccond antennal segments fairly thick; tibiae not

as above (África) VOLUMNUS Stál, 1865

Roslrinn reaching thc 8th abominai segment; first antennal

segnicnt withont long, dense and rigid sctac; tibiae withont

long s])ines (North America) .... KCAüiTOIilA Uhler, 1909

Rostrnm not reaching the 8th ahdominal segmcnl; first ati-

tennal segment with long, dense and rigid setae; tibiae with

long spines (África) TIl/C.HOliASlS Renter, 1904

First antennal segment longcr than width of head (fig.

198) 44

First antennal segment shorter or etpial to widtli of head 48

Vertex snleate longitndinally; frons striolate; third antennal

segment equal in Icngth to first; sccond slightly longer tlian

tliiril; long p;nallel sided species (fig. 198) (Gosmopolitan)

CAIEONTIADES Distant, 1883

Vertex not snleate, frons not striolate 45

F.yes removed Iroin anterior margin of jtronotinn by an espace

;d)ont ecinal to length of one eye; claval vein distinctly raised;

pnbescence black and setilorm (Fmope, Asia)

' onONTOPLATYS Ficber, 1858

Fyes contignons with anterior margin of pronotnm or so;

tlav;d vein mjt laised 46

Posterior lemor;i comi)ressed, fairly mnrowed towards thc

anex- membiane marmorate (as in Pliylocoris) (North Afri-

EREMOIUEI.EUS Renter ,1895

Posterior lemoni not comi)ressed and noiiceably narrowed

tow;nds ;ipcx; membiane not maimoiate 47

Fyes somewhat

(New (hiinea)

.emoveil Irom anterior margin of iironotnm

lUSI ROPEPLUS Poppius, 1915

88 15()i,. Mus. G()ei.i)I — '1'onio XI (11) — Dezembro 19 j5

— Eyes conliguous with anterior niargiii of pronotnin; species of

large size; body not densely ])ubescent; cuneiis longer tban

wide (Asia, índia) .. TIIICHOPIIORONCUS Reuter, 189(5

-IS. Eiongate sjiccies with bracliyptcrons feinales; males with

cunens two and balf times as long as wide at base; tibiae with

long spines; pronotnin sitnate laterally, strongly narrowed

towards apex; feinales witbont distinet cunens and mem-

brane 19

— If bracbyjiterism orrnrs in feinales, tben cunens less tlian two

and balf times as long as witie at base; tibiae witbont noticc-

able long siiines; pronotnm not strongly narrowed in front;

feinales with cuneiis and membrane üO

’ 49. Pronotnin simiate laterally: clypeiis flattened, scarcely conve.x;

eyes inclined forward; ciinens absent on feinales (Asia, Euro-

pe) ALLOEONOTUS Eieber, 1858

— Pronotum not simiate laterally; clypens arcuate-convex; eyes

vertical in jiosition; cnnens distinet on feinales (North Ame-

rica) srrrrocAPSUS Knight, 1912

50. Rostnim reacliing sliglitly beyond apex of anterior coxae:

body long and ereclly jiilose (Asia, Eiirope, North África)

liRACllYCOLEUS Eieber, 1858

— Rostrum reacliing ilie middie coxae or beyond 51

51 . lllack species witii jnginn .separated froni lornni by a deep

suture; first rostral segment iiinch tliicker tliaii otliers; body

above nigose, often willi pnnctale as|)ect (Norili America)

IRlilSlA Rcntcr, 1879

— .Species if black, witbont tlie combination of diaracters

above 12

52. Scutellnm strongly tnniid and stee]) |)osterioi ly, feebly decli-

vons towards apex; body covered by adpressed, silvery pnbes-

cence (New Cbiinca) .... PEl/El DOEICEEUS Poppins, 1912

— Scntellnni not strongly tnniid or if so, llie body witiioul sil-

very adpressed piibescence i 5!$

5.H. Eyes renioved froni pronotnm by a distance abont etpial to or

more tliaii tliickncss of second antennal .segment; collar wide,

talli large and conllnent, reacliing tlie lateral niargins ol pro-

notinn. tlie latter soniewliat tlepresscd beliind talli (ligs. I8(i,

187) 51

— Eyes contignons with pronotnm or if not tlien talli not large,

not toniincnt tir rtiatliing lateral niargins of |>ronolnni (lig.

2(».'5) 57

8í)

Carvalho — Kcys Cíciicra iMiiiclac ol thc World

51.

SC).

58.

5í).

(iO.

Collar very wide with inesal lengtli about eíjual to half the

wiilth of one cye; pubescencc long, 1'inc and crect (Europc)

DIONCONOTUS Rcuter, 1891

Collar abont as thick as íirst aiitennal scgmcnt; pidjesccnce

sliort ainl adprcsscd, easily rubbcd oíf 55

Calli vennícidatc scidptnrated; posterior lobe oí pronotuin

carinatc at middie (Corsica)

RHABDOSCYTUS Ilorvath, 1921

Calli not as above; pronotiim witliont a tarina on posterior

lobe 56

Froiis strongly .swollen in front (fig. 187); pronotmn dis-

tinctly iiarrowed and soinewliat coiistriclcd in Iront (Eiirope,

Asia) GRYPOCORIS Douglas &; Scott, 1868

lòons not noticeably swollen in front; pronotinn not cons-

liicted in front (fig. 186) (Enrope)

líADRODEMUS Fieber, 1858

First antcnnal scginent alinost as thick as diameter of eye

(fig. 2().'5), slender at base, the othcrs very slender; posterior

feniora very thick, tibiae with long spines; pronotmn and

ennens strongly declivous; hcinclyria adpressed pilose, ])rono-

tinn with setiform and silky ])nbescence (.South America) ....

PORAS Distam, 189.8

First aniennal .segmem not as above; posterioi- femora not no-

ticcably thick; pubcscence not as above 58

llody above (overcd with short black setiform bristles, cs])e-

cially visible on sides of pronotmn and exocorimn .... 59

Hody above covercd with hairy i)iibescence only, withont seti-

form bristles 61

Corium with very prominent veins, the cid)ital branched at

apex; first segment of himl tarsi thicker than .second, dcci)ly

excised at apex, longer than .second; pronotmn with lateral

margins acnte anteiiorly; females identical to males, cnnens

more than twice as long as wide (it)

Corinm withont iirominenl veins; pronotnm with ronndetl

lateral margins; females with very short membrane, the < nnens

as long as wide at base (Enrope)

no RV ATI II A Renter, 1881

llody oval; head seeti fiom above as long as witle, cyes conti-

;nons with pronottim; rostrnm reaching slightly beyond mid-

ile coxae; Íirst rostrai segment reaching middie of xyjdms

iCTINONOrUS Renter, 1896

Ifody std)elongate; head seen hom abo\e iniuh longtt than

vide; eyes sotnewhat removid fiom pionotnm; rostinm readi-

90 Mi;.s. CioKU)!

lonu) XI (II) — Dc/cinbro 1955

ing the base of heacl (Eiiropc, North America)

MI RIS 1-abricius, 1791

GI . Vertex longitiulinally sulcate; botiy cscarcely j)iibescent, al-

most glabrous or with very long, erect and esparse jjubes-

ceiice 02

— Vertex iiot sulcate longitudiiially; body distinctly pidies-

cent 04

02. 15ody with sliort golden and black piibescence (North África)

REUTEIUSTÁ Kirkaldy, 1904

— body almost glabrous or witlt long, erect pubcscence .... 03

03. body almost glabrous: eyes not very large; antennae not notice-

ably long; bemelytra not shagrine (Kurope, Asia, África, Aus-

trália) MEGACOEEUM Fieber, 1858

— body than long. erect pidjescence; eyes very large; antennae

longer than tbe body; bemelytra finely shagrine (Ceylon)

RIIARYEEUS Distant, 1904

01. Rostrum reacbing beyond apex of posterior coxae; head broad

eyes proctically in contact with jironotal angics, hind margin

of eyes somevvíiat flattened and forming an arcuate line with

base of vertex 05

— Rostrum not reacbing beyond apex of posterior coxae; head

not unusually broad, eyes convex behiiul and well lemoved

from pronotau angles 00

05. Vertex distinctly ctirinate; lieatl very sltort, vertical, eyes com-

pressed; pubcscence not silky (Índia)

SARINNIUS Distant, 1909

— Vertex not tarinate; beatl not noticeably sliort and vertical:

eyes not strotigiy {omjirc.ssed; pubcscence not noticeably silky

(lúiroj)e, Noitb iAmerica) . . DIEI I ROOSEVEUS Eieber, 1858

00. .Second antennal .segment about as tbick as or tliicker tliati

lirst; eyes large, margined posteriorly (lig. 201); clavus witlt

duli tomentose dust (,\frica, 1’acilic Is.)

SI DMA Retiter, 1905

— .Second antennal segtneiU sleniler than first; eyes not margined

posteiioily; clavus witiiottt totnentose dtist 07

07. Head stiongly pointed in liont; veitex (arinate; up|)er suilace

ol body lugose; jnonotutn with a sliglit median imjiression

(Aflita) CIXACORIS 1’oppiits, 1912

— Head not stiongly pointeil; body smootli; jironotum convex

or Hat, not imptessed 08

08. riiitkness of fointli antennal segment almost etpial to that

ol base ol secoiul segment; mesal lengtb ol tollar subetpial to

Carvalho — Kcys Gencra Miriclac of thc World

tliickncss of íouitli aiUcnnal segmciu (fig. 219) (Cosmopo-

litaii) ADELPHOCORIS Rcuter, 189G

— Fourth aiitcnnal scgmciu distinctly thinncr thaii base of

sccoiul scgiiicin; incsal Iciigtli of tollar distinctly greater than

tliickncss of foiirtii scginciu 69

69. Proiiotimi glabroiis, sliiiiiiig; pubcsccncc of hciiielytra very

sliort (North yVmcrica) .... GANOCAFSUS Van Duzec, 1912

— rronotiiin pubcsccnt; jjubcsccncc of licinelytra not noticcably

shoit 70

70. Vcrtcx fincly carinatc; collar vcry slcnder (Java)

CALOCOliOPSIS Popitis, 1911

— Vertc.K iiot carinatc; collar not noticcably slcnder (fig. 38)

(Cosinopolitan) CALOCülilS Fiebcr, 1858

71. Rostrinn sliort, cscarccly surpassing anterior coxac or rcach-

ing niiddlc of incsostcrntim, in a fcw spccics rcaching anterior

niargin of middlc coxae (Índia)

LtOCAPSUS Poppitis, 1915

— Rostnnn longcr, ícacliing at Icast posterior niargin of niiddlc

coxac 72

Vertex distinctly snleate (Central Aincrica)

JACCJl/NUS Distam, 1873

Vertex not distinctly sulcatc or only vcry fincly so 73

Kycs vcry large, coniprcsscd, sniooth bcliiml, loiicliing tlic an-

terior anglcs of jironotinn and giila bclotv, strongly rcnilorni

in front; vertex tliickly niargincd; first antciiiíal scgiuciil

sliortcr or ctinal to rvitli of vertex (África)

JACJDOLON Remei, 1907

— Kycs not (onipres.sed or toncliing anterior anglcs of pronottim

and gnla lielow; vertex not niarginate; first antennal segment

longcr tlian widili of vertex 71

71. Large spccics over 10 mm long; .sciitelltnn rngose, iiniire.ssed

longitiidinally: dypetis strongly proininem (New Ctiinea) . .

MACIiOPP.Pl.US Pop])ins, 1912

— .Sinaller spccics witii snioolli .scntelliim, not iinpiessed longi-

tndinally; (ly]K'tis not noticcably proniinent 75

75. Kycs coinpressed and large; body oiiaipie; iironolinii strongly

dedivons (Madagascar)

IDEPPIlüCOniDEA Poppius, 1912

— I■'.yes not compressed; body shining; pronoiinn not strongly

dedivons

76. Posterior tibiae witli spines tlirongliont; lortmt prominem,

soniewli;it taiitiate (,\ineii(as) .... lIOJiCAAS Distam, 1881

72.

73.

cm i

SciELO

10 11 12 13 14 15

92 Hoi- Mus. Goki.di — 'lonio XI (II) — Dezembro 1955

— Posterior libiac vvitli oiily a fcw apical spines; loruni not

iioticcably proniinciu (Java) ... GIANELLA Poppius, 1911

77. Heacl horizontal, vertex longitudinally siilcatc and striolatc;

jnguin vcry proininent, set off as two [jointed tubercles at

tbe sities of cly])eiis: third and fonrtli joints ot antennae ini-

mite; rostrinn reathing tbe posterior coxae (Central zVmerica)

MÍNYTUS Distant, 1883

— Jngnin and vertex not as above; if tbird and [onrth joints of

antennae ininnte, tbe rostrnm reacbing tbe iniddle coxae 78

78. llody covered by vcry long, fine and crect itnbescence, cspe-

cially on exocoriíim and scntelhnn; lieinclytra with silvery

areas or adpresscd silky or wooly pidresccnce aniongst tbe crect

bairs 79

— llody sbortly jrilose or if long pilose, tbe hcinclytra withont

silvery spots or silky or woolly pidjcsccncc intcrinixed with

long liairs 83

79. Pnbesccnce of heinielytra with silky silvery adj)resscd bairs

amongst tbe crect pnbesccnce; or silvery toinentosc arcas

|)rescnt (Ceylon) DIOGNETUS Distant, 1901

— Pnbcscencc of bcmclytra witb silvery tonicntosc spots on bc-

niielytia, siinctiines aiso a few sbort silvery bairs 80

80. Pronotnm strongly convex at iniddie of disc, witb a tnbcrcnlar

clevation (an crect lobe) (fig. 210) (Formo.sa)

riNGINOTOPSIS Poppins, 1915

— Pronotnin not as above 81

81 . Kinbolinin witbont transverse datk and byalinc fasciae; ])n-

bescence of body not noticeably long (Ceylon)

ARGENIS Distant, 1901

— KinboliniTi witb transverse dark anil byalinc fasciae; body pn-

bescence vcry long 82

82. Rostrnm reacbing ajn-x of mesosternnm; embolio-corial tom-

missiire at base witb pnncinres; bead somcwb.it i)roiln(ed

betwcen antennae (Fiji, Samoii)

NESODAPIINE Kirkaidy, 1908

— Rostrnm reacbing tbe |»osteiior coxae or beyond; embolio-

(oriid connnissnrc witbont pniutines; bead not prodnccil an-

teriorly (Índia, África, Java, Pbilippines, etc.)

'. .. riMGINOTUM Kirkaidy. 1902

First antennal segment notiteably enlarged, lompressed or fo-

liaceons or witb two fiist segment noti(e;d)ly tbick (fig.

200) 81

l'iits antennal segment if enlaigeti, cylindric.d, or lirst two

segments not notiteably tbiik (iig. 201) 85

83 .

CIakvai.ik) — Kcys Ciencra Miridac ol tlic \V'oi'lcl

í).‘5

81.

87.

88 .

8 !).

;»().

í»2.

llcad more or Icss horizontal, acutely pointcd in íront; pro-

notum .strongly narrowcd and declivons; 1'irst antcnnal seg-

inciu withoni l)ristlcs and scalc-Iike hairs (fig. 200) (Ceylon)

CLAPMARIUS Distam, 1901

llcad not horizontal: inonotnm not strongly narrowed in

Iront; 1'irst antcnnal scgmcnt vvith hristlcs and scale-likc or

woolly hairs (Formosa)

EURYSrYLOMORFHA Poppins, 1915

Sccond antcnnal scgmcnt distinctly clavatc 86

.Sccond antcnnal scgmcnt linear or so 91

Upper snrlacc with goldcn ycllowish scale-likc jndrcscencc;

hcmclytra rngosc, vertex carinate (Peru)

ACANTUOREPLUS Poppins, 1912

Upper snrlacc without scalc-likc ))ubcsccncc 87

liody (cxccpt hcmclytra) very long and crcctly ])ilosc (Aírica)

TRICnOCAPSUS Poppins, 1912

Hody without noticcahly long jnibesccncc 88

Pronotnm crcctly and hcmclytra shortly adjncsscd pilosc;

first antcnnal scgmcnt ver short and thick (North África)

niSTRlOCORIDEA Poppins, 1912

Hody pidrcsccncc not as above; first antcnnal scgmcnt not

noticcably thick 89

Ilcmclytra covcrctl by silky adprcsscd pnbcsccncc; scntcllnm

transvcrsally rngosc; rostrnm snrj)a.ssing slightly thc apex of

anterior (oxac ( Fonkin) P/IAN/A Poppins, 1915

Hcmclytra withont silky adincsscd pidjcsccncc; scntcllnm not

transvcrsally rngosc; rostrnm longcr <)()

Vertex very witlc and with a median shallow depression; jn-

gnm strongly tnmid; frons .smooth, pronotnm pnnetate (fig.

197) (F.nropc, Asia, North America)

CAPSUS Fabricins, 180:!

V'crtcx not noticcably wide, withont median depression; jn-

gnm not strongly tnmid; frons striolatc; iwonotnm rngosc

(África) ]>SEUn()RTII(>TYLUS Poppins, 1911

Pronotnm strongly declivons; .scntcllnm strongly prominent;

hcmclytr;i shíigrinc covered by small sralc-likc dchisctmt haiis,

ennens strongly inclincd (fig. I«5) (Central & .Sonth Ameri-

; DEROPirriLUMA llcrg, I88;i

Pionotnm if strongly declivons. lhe .scnicllum not as above

or if so. thcn lhe hcmclytra wilhonl adprcsscd scalc-likc hans

giving shagrccncd a|)pcaiancc *-

llcad shoit, vertical, wide. frons siriolalc and snlcatc; cycs

large, compicsscd. ocdipying thc sides of thc licad; pronoinm

91 lloi.. Mus. r.oEi.Di — lomo XI (11) — Dc/cinhro 1955

stroiigly pimctatc, Iicinclytra l)csct willi goldcn adpicsstcl pu-

bcsccncc or tliis typc pliis roninioii liairs (lig. 195) 95

— Ilcad not as abovc, if írons sliiolatc tboii bcmclytra with

otlicr typc of pubcsccncc or pronotimi not strongly piiii-

ctatc 95

95. Aiitcnnae with sbort liairs and long, crcct linc .sctae; lieincly-

tra witb crcct hairs intcnnixcd with adprcsscd oncs (lig. 195)

(Central &: .Sonth America) .. CALOCOIUSCA Disiant, 1881

— Antcnnae with a single oí pubcsccncc; bcmclytra witb a single

typc of pubcsccncc 91

91. llody witb densc goldcn atljircsscd pubcsccncc; membrane

very sbort and dcnscly pnbcscciit (Central íb .Sonth America)

KUCIIIÍ.OCORIS Renter, 1907

— IJody not noticcably dcnscly pilosc; pronotnm and sditcllnm

strongly pniutatc; membrane glabroiis (Vcnc/ncla)

CURYSODASIA Renter, 1892

95. I Icmclytra clotbcd with distiiu t silvery or woolly or silky

pubcsccncc, single or mixed witb linc, crcct hairs, sometimes

casily nibbcd oH 9()

— I Icmclytra glabrons or clotbcd with oiily a single typc of

]mbcsccncc, sometimes scmiadpressed bnt nevei i.idy silky

or woolly lOti

90. First antcnnal segment very ihick, as wide as widih ol one

cyc; pronotnm with three collonsed lines anteriorly on the

sides (Anstralia) 1)1 R I lORA I.IA Renter, 1905

— 1'irsi antcnnal segment not as ihick as width ol eye; proncinm

withont the lines abovc " 97

97. 1’ronotnm tvith a median subexcavaled dnll spot behind calli;

scnielhnn strongly tnmid; imbesceme ol black siilf brisiles

and silvery hairs (índia)

lAUiYSrVl.OrsiS Ooppins, 1911

— Pronotnm withoni the spot abovc; scntelhnn not noticcably

ttmiid or íl so, tben body pnbesccme not as .ibove 98

98. Pronotnm with iwo dnll depressed black spois (as in Tnrdiu),

covereci by black brisllcs only (.\lrica)

ri.tMOC irsUS Bergroih, 1920

— Pronoimn withont two dnll black spois as abovc or il so.

lhen pronoimn with woolly pubesceme 99

99. body covereci by silky pnbescencc iniei mixed with crcct and

linc hairs; |)ronolnm coarsely and cleeply pnnclale; scnielhim

Hat ()ava) CORXA Poppins, 1911 '

• lhe neiiii' Síi/igiuo Kellciii, (Ndilli Aineiii.i) usiches abii lliis (ini|)lel

ol lhe Key.

Cakvai.iu) — Kcys Clencra Miridac of tlic World

!)5

— Hody covcrcd hy aiiprcsscd ])id)cscencc oídy; pronotiiin linely

punctatc or if coarscly punctatc then sciitellimi tuinid lOO

100. llcad stroiigly poiiitctl in front; daws loothcd at base; ostiolar

])critreiiic vcry large (África, índia, Madagascar, Java, etc.)

PROliOSC/DOCOJlIS Reuter, 1894

— Ilead not strongly poiiued in Iroin or if so, then claws not

tootlicd at base or ostiolar pcritrcnie not vcry large .... 101

101. .Sciitellinn strongly tnniid 102

— .Scutellnni not strongly tnniid 103

102. Pronotnni coarsely punctatc, glabrons (índia)

PELTIDOLYC.IJS Poppius, 191.5

— Pronotmn finely nigoiisc ])iinctate, dorsinn clothed witb sil-

very woolly ailpressed |)ubesccnce (Nortb yXincrica)

CA LYP PODERÁ Van l)ii/ee, 1923

103. Pronotnin soinewbat rngose punctatc; rostrimi rcacbing thc

VI to VII abdominal scgincnt; clongatc spccics with tbc aspcct

of Lygii.s bnt witb silvery bairs amongst tbc pubcsccncc of

hcinclytra (Índia, Ccylon) El/EIÍEMUS Distant, 1909

— Pronotmn not rngose; rostrnm shorter, ])id)csccncc distinctly

silky: nsnally roniulisb daik spccics 104

101. Pronolnm csparscly |)nnctatc, stiongly shining, scantily pnbcs-

cent; hcmiclytra somcwbat rngose punctatc (Asia, índia) . . .

EfOCORIDEA Rcntcr, 1903

— Pronolnm distinctly pnbcstcnt; licmcivira not i ngoscly pnne-

tatc '. 105

105. First and setond scgmciits of bind tarsi ctpial in Icngth; collar

tviili mcsal Icngtb largcr tlian ibickncss of first antcnnal seg-

ment (Fnropc, y\sia, África)

CIlARACOCHir.VS Ficbcr, 1858

— First segment of bind tarsi distinctly sborter tban sccond;

colbn witb mcsal Icngtb ctpial to tbickncss of first antcnnal

scgmcnl ((losmopolitan) .... POiAMERUS Wcstwood, 1839

lOft. Pronolnm stiongly tonvex above and lonndcd latcrally, biil-

liant, (alli not visiblc, (ollar vciy linc and dcpicsscd covcicd

by tbc vcitcx wbidi is sbarply (arinatc and continnons witb

postcrioi maigin of cycs; scntcibmi stiongly tiimid raising

mndi above tbc sinfaíc bcmicdytia, c lavai (omissnre sboi tcr

tban s( nlclbim ((Icylon, Java) lU.R ESA Kirktildy, 1901

— Pionolnm ;ind (ollar not as aliovc, il so tbcn scntcllnm not

noti(c;ibly Inmid or cycs diliciont 107

107. llcad bmgci tban wiilc, logctbcr witb (ollar abont as long

as |)tonotnm, sballowly snlcatc at middie, somcwbat bori-

/ontal. dypcns vciy laigc; cycs distant liom inonotnm by an

Boi,. Mus. (íoki.di — 1'oimo XI (II) — Dc/cinbro 1955

ÍK)

csjiacc cijiial to cliainctcr of íirst antennal scgincnt; jaonotum

strongly narrowecl iii front; first antennal segnient alinost as

long as liead and pionotinn together (lig. 221) (África) . . .

^ KIAMIiURA China, 193G

— Ilcad and íirst antennal seginent not as above 108

108. Pronotum distinctly punctate hetween calli and posterior to

collar (fig. 190) 109

— Pronotum impiinctatc or nigose hetween calli and posterior

to collar 117

109. .Sinal I highly polished tleeply and coarsely punctate alniost

hairless species; vertex thickly inargined; pronotum strongly

declivous; corium jmnetate only uear claval commissurc and

on emholio-corial commissure (fig. 201) (Samoa)

PLESIOLYGUS Knight, 1935

— Body suríace not as ahove or if so, then vertex smooth, prono-

tum not strongly tieclivous, corium otherwise piictate . . 110

110. Lateral margins oí pronotum not carinate; form more elong-

ate and suhparallel 111

— Lateral margins of pronotum carinate or at least with a cal-

loused liiie; form Ovoid or body very strongly punctate, jiro-

notiim with the espace hetween jiiinctures forming here and

there tubercular sweilings Ilfi

111 . Rostrum reaching the 5th abdominal segment 112

— Rostrum uot reaching the 5lh abilomiiial .segment .... 113

112. Anteuuae iuserted distinctly below apex of eye; ostiolar peri-

treme iudistiuct; hemelytra rugosely punctate (Suécia) ....

y.YG/MUS iMcber, 1870

— Antennae iuserted above apex of eye; ostiolar peritreme dis-

tinct; hemelytra punctate only (Kurope, Asia)

CAMrroZYGUM Reuter, 1890

113. Membrane linely and ilensely jiilo.se; head smooth, roslrum

reaching ihe middie toxae (.\kiita)

YXGVEEl.lA Popiiius, 1912

— Membrane smooth, head punctate or striolate II 1

H l. First antennal segtnent as long as width ol head; species of

liitge si/e, Irous striolate (fig. 195) (Central Xc .South America)

EíASUS Distam, 1883

— First atitcnmd segment shottei than width ol vertex: speiies

of médium si/e 115

115. Irons stiiolate; setond anteuual segment five times as long

as Íirst segment (Not th .America)

XEOIIOHEI.I.A Ktiight, 1925

SciELO i'o

98 ]ioi.. Mus. Goeldi — l oino XI (II) — Dezembro 1955

125. Vertex with median loiifritudinal sulcus; lonim prominent

and carinatc; memhranc short and densely pilosc; pronotinn

strongly shining (fig. 183) (Central &: South America) ....

CALONDAS Distant, 1883

— Vertex not sulcate or if so, lomm not imominent or carinatc;

membranc glabrous 12()

126. First and sccond antcnnal segments thickly clotbed with beavy

black pubescence; large red spccies (North America)

COCCOn APUES Uhler, 1878

— First Antcnnal segment more esparsely clotbed with pale pu-

besccncc or if black, colour not red 127

127. Pronotum with lateral margins sharply angulatc, carinatc near

jiostcrior angle; body red or brick red and black; clavus with-

out punctures (Américas) NEOCAPSUS Distant, 1881

— Pronotum with lateral margins rounded or angulate, not ca-

rinate, but if so, then clavus with two rows of punctures 128

128. Pronotum slightly carinatc ou lateral anterior margin; clavus

beset with onc row of |)unctures ou each side of claval vein;

large sjrccics with pronotum strongly punctatc, ahnost gla-

brous, frons striolatc (Sumatra)

rOI.ONGíA Poppius, 1915

— Pronotum not carinatc; clavus without punctures .... 129

129. Pronotum distinctly and coarsely rugosc tratisvcrsally, with

punctures mostly obscured by rugosities 130

— Protiotum punctatc but not rugosc, sometimes only very fin-

cly so 135

130. Pubescence very short atid csj)arsc; rostiinn reaching apex

of iniddlc coxae 131

— Pubescence not as above; lostrum naching ihe hiiul (oxae

or bcyond 132

131. Calli large, confluent, reaching sides of pronotum; width of

vertex greater thati width of an eyc (Europc, North África,

Asia) PI. ES! OCO RIS Fichei , 1861

— Calli small, not cotillueiU our readiing sides of pronotum;

witlth of vertex less tlian width of an cye (North .America)

NEOIiOROPS Uhler, 1895

132. .Antcnnae very long, first segment as long as head and collai

together, thc sccotid segment etpial in leiigth to third; pro-

notum distinctly narrowed anteriorly (África)

inJTTNERIEI.I.A Poppius. 1912

— Antetinae shortei, not as above, pionotum not notiteably nat-

rowed anteriorly 13.1

CLvkvai.ik)

Kcys Gcncra Miriclae of the Woikl

133. Hcacl vcry wiclc, vcrtex twicc tlie dorsal width of an eye, cari-

natc; cycs cxtciuliiig bcyoiid anterior anglcs of pronotum;

small species not over b.b nini long (North America)

liOLTERlA Llhler, 1877

— llead and vertex not as above 134

131. Hody almost glabrons, the hcmielytra sinooth; rostrnm reach-

ing tbe (ith abdominal segment (New Gninea)

IRISTOPEPLUS Poppins, 1912

— Hody long, fine and erectly pilose; bemielytra rngosely pnnct-

ate; lostnnn not reacbing tbe btb abdominal segment (Áfri-

ca) IIORVATIIIEELA Poppins, 1912

135. Pronotum pisceons, slrongly sbinnig; bemielytra black, opatpie;

arca between calli distinctly rngose (Enrope)

SAUNDERSIELLA Renter, 1890

— Pionotnm not as above; tirea betwenn calli smooth .... 13(i

13(i. .Setond antennal segment sliorter or abont as long as width

of liead ticross eyes (Etirope, North America)

AGNüCORIS Rcuter, 1875

— .Secoml antennal segment distinctly longer than width of head

acros eyes 137

137. Eions tnmid, somewhat snlcate, ilistinctly stiiolatc; clongate

species with pionotnm strongly declivons 138

— Frons if tnmid not striolate or snlcate 139

138. Rostrnm reaching the Ith or 5th abdominal segment; prono-

tum pnbescent (África)

MACEDANUS Hergroth, 1920

— Rostrnm reaching the hind coxae or nearly so; pronotum

almost glabrons (Índia, Ghina)

PniLOSTEPflANUS Distant,, 1909

1.39. Large clongate species itarallel sided (males), oval or as wide

at base, with apiiroximale same width throughout; body al-

most glabrons (North AIrica, Asia Minor)

ISCAINOSC.EIJCORIS Rcuter, 188(i

— .Sixaies if large or clongate then cnnens mnch shorter, body

pnbescence distinct

110. llead pointeti between antennae, somewhat hori/ontal; se-

cond antennal segment abotit as thick as fiist, thitd tiiitl loiuth

very slmrt; rostinni reathing the /th abdominal segtncMit,

bodv imbestence very short and .scanty (Centi.il Ametica)

_ ' ; PAPPUS Distant, 1883

llead not noiiteably pointed between antennae, vertical; se-

lond antennal segment as well as thiid ;ind lontlh not .rs

above or il so then lostinm shorter I’!*

100 15()i,. Miis. CioKi.Di — Tonio XI (H) — l)c/cinl)ro 1955

Hl. Scutellmn stiongly tiiniiil, imith highcr tlian jjroiiotimi; ros-

triim not rcaching inickllc coxae; bocly with loiig, crcct hairs

(fig. 189) (Chile) CHILEAIÁ Carvalho.

— Sculclluin il timiid not liigher than jnonotum Hli

142. 1’ronotuni distinctly and dcnscly punctate, hairs nsually ad-

pressed or suhadprcssed 1 l.‘i

— 1’ronotuni shallowly and csparsely puetate, hairs nsiially

crcct, íinc and long 148

ll.H. Ilind tihiac hlack or witli a hiack spot or ring 144

— Iliiul tihiac nnicolorons, palc 145

144. Rostrum rcaching thc iniddie coxae; hciniclytra not rugosc

pimctalc; sccond antenna sliort and incrassatc (África) ....

líISTRIOCORÍS Renter, 1905

— Rostiinn rcaching thc hind toxac; hciniclytni rngosc pnnetate;

sccond antenna long and incrassatc only at apex (holartic)

EXOLYGUS Wagner

145. riiird scgincnt of hind tarsus longcr than sccond; hind Icnioia

imich stouter than othcrs (Kuropc, North África)

CYPIIODEMA Ficher, 1858

— 'Fhird scgincnt of hind tarsiis as long as or shorter than .sc-

cotid; hind fcniora not nnich stouter than othcrs 14(5

14(5. Pronottnn rngosc pnnetate; s])ccics over 5.5 nnn long (Asia,

North America) LYGIDEA Renter. 1875

Pronottnn jtnnctatc hnt not rngosc; spccics Ic.ss than 5.5 nnn

long 147

147. First and sccond antcnnal scgincnts iinrassatc, dianictcr of

scconil scgincnt ctpial to tliamctcr of forc tihia (Formosa) . . 1

EOEYGUS Po])pins, 1915

— First aml setond antcnnal segment not incrassatc oi vciy

slightly so, greatest diamctci of setond antcnnal segment not

ctpial to tliamctcr t)f lorc tihiac (F.nro|)c. North >\Íiica. Asia,

North America) ORTIIOPS Ficher, 18.58

148. Frons with ftitir to live pnnctnrcs ahove anicimal lossa; (spc-

cics of small si/c (Intlia) SA liAGTUS Distam, 1910

— Frtms wilhoni thc pnnctnrs mcniionctl aliovc 119

149. l.cngth t)l lirst antcnnal segment shorter than cyc height. if

this ralio etpial then vci tex notiteahly sinnatc at postei ior

m.ngin 150

— I.ength oi fiist antennal segment longer than eyc height. il

this ratio eipial then vcitex stiaight at postetioi maigin 15">

150. .Setond atitennal segment slioit. intiassate. thiisely pilosc;

hotiy laiily long. eiettly pilosc (lig. 191) ((iential aml .Sotith

Ameiita) EUllATAS Distant. 1881

Carvai.ik) — Kcys GcMicra \íiiidac of lhe World

101

— Sccoiul aiucnnal scgincnt if short or incrassatc not densely

pilosc : 151

151. Vertex straiglu i)ostcriorly: sccond aiitcmial segnicnt four

times longer than first segment; rostnim leaehiiig tlie middle

ol abdômen (Asia, North America)

PINALITUS Kelton, 1955

— Vertex simiate a posterior margiii; secoiul antemial segment

iisually less than ■! times longer than first segment; rostnnn

nsnally reaching the hind coxae (may reach heyond) . . 152

152. l o this conplet come the genera Dagberius Distant, 1901 and

Taylorilygus Leslon. 1952. .Since their seiraration hased on

externai characters is dilficnli and the latter may prove to

he a synonym of (hilrida Kirkaldy, 1902, further stiidics

shonld he lindertaken on the snhject.

155. Rostiimt reachitig the middle or hittd coxae; first atttennal

segtnent shorler than width of head (Knrope, Asia, North

Americti) LYdUS llahn, 18.55

— Rostnnn rcachitig the middle of ahdomcti; first atttennal seg-

ment longer than width o fhead 151

151. Ilaed seeti frotn ahove mote thati twice as wide as long, lacial

atigle acnte (Vene/nela) . . . NEOSTENOTUS Reiiter, 1905

— 1 lead seen Irom ahove twice as wide as long, seen Irom sido

as lotig as high, facial atigle siraight (15ra/il) .

A EDA Renter, 1909

KKY TO SlIUGhNKR.V Ol' lA GUS IIAIIN

1 . d ratisverse caritia of vertex ])re.setit hnt incompletc at middle,

evident only near eyes LlírUS (lA(iUS) Ilalin

— 'riansverse catina of head (otnplete -

2. Tihial spittes hlatk l.YCtUS {,-\POIAGVS) (diina

— 'l ihial spines pale or hrown . . . LYdUS (^NEOL) (AIS) Knighl

Tlir jollotving ^ciicrn of Miiiiii nre not includcd iu the kcys

Aratitlioniiiirllii l’()|i|)iin, llll l (Atla .Soe. .S(i. Icnii. 1 1 (.1): lll), iicar Itopido-

filioirlld Kciitcr, /an/il)ar.

(]hiiia, Itt.tl (.\iiii. /.(><>I. Ja|). 15: 20.')). Japai).

4uUii>((,l)sui Kiikaldy, liUlI (^.ntom. .51: 110), nlliccl lo Uyidoprjdus, Aiistialia.

Diplotutliirlln l’o|>i)iiii, ItH.') (Ann. Mus. Iluiin. 15; 0.')), Índia.

I.hlix Kiikaldy, l'.KI2 (Trans. r.nl. .Soe. I.ondon, 250), allinl lo (.'«psio, India.

(■iilmiti Kiikaldy, 1!H)2 (Kniom. .55: 581), CaOooii.

102 lloi,. Mus. CioKi.i)! — loiiio XI (11) — Dczcinlíio 11)55

Kmigra KirkaUly, 1902 (Trans. Kiit. Soc. I.oiulon, 257), allicil lo Jlyalopeplus,

Ilidia.

lAocapsidea Poppius, 1915 (Ami. Mus. Hung. 13: 16), iicar Liocapsus, índia.

Macgregorius Kirkaldy, 1903 (Wicn. Ent. Zcil. 22: 11), Quccnsland.

Mennilolccerns Rculcr, 1907 (Ann. Mus. Zool. St. Pctcrb. '189), Asia.

Nesosylplias Kirkaldy, 1908 (Proc. Linn. Soc. N.S. Walcs, 33: 379), Fiji.

Ninslanin Rculcr, 1901 (Ofv. E. Vct. Soc. Forh. 17 (5): II), 'Fasmania.

Ocierocapsus Poppius, 1915 (Ann. Mus. Ilung. 13:17), Atistralia.

Olympiocapsus Kirkaldy, 1902 ('Frans. Ent. Soc. London, 255), China.

Pachyptcrna Ficljcr, 1858 (Wicn. Ent. Monat. II: 301), lúiropc.

Poccilonotus Rculcr, 1897 (Ofv. F. Vct. Soc. Forh. 38: 167), Asia Minor.

Ruspolielta Poppius, 1921 (Ent. Mitt. 10 (3): 82), ncar Lamprocapsidea, África.

Tropidopliorella Rcutcr, 1907 (Ofv. F. Vct. Forh. 49 (7). 15). África.

Megacoelopsis Poppius, 1912 (Acta Soc. Sei. Fcnn. 41 (3): 40), África.

Ommatodrmn Poppius, 1911 (Ofv. 1’. \'ct. .Soc. Forh. 53 ,\ (3): 4), Fasmania.

KKY rc) riIE GENER.V OE MECISTOCEIJNI

1 . Rostriiin rcacliiiig bcyoiul tlic posterior toxae; caiiiciis a litllc

loiiger tliaii broatl; hemiclytra glassy transparciit (fig. 11) (Ín-

dia)

MYSTII.US Dislaiu, l'J()l

— Rostriiiii rcatliing tlic niitldlc toxae; timeus nuith longer tliaii

broatl; beinielytra opatjue (Iiitlia, Java)

MEClsrOCEl.lS Reiilcr, 18í)l

KEY K) riIE GENER.Y OE P! EHANINI

1. .Spet ies noticeably inyrmecomorpliif wiili alitloincii strongly

tDiistrit teti at base aiitl not eoveretl liy tbe very sbort liemiely-

— .S|)etie.s witli a cerlain ant-like appearente but not noticeably

inyrinecoinorphic, tlie abtionien eoveretl at least pralitally by

tlie beinielytra 2

2. I.engtli of first antennal .scgnient less tlian wititli of vertex;

pionotinn not extentling batk lo liasal angles ol lieinielyira

(Etirope, Asia, N. África, N. .\nierita)

PI TII.LWS Eieber, 1858

— I.tngib ol lirst antennal segment greater iban wiiltb ol veriex;

pionolinn extentling b.ick to basal angles of lieinielytra (Norili

Ainei ica)

iMlMOCEPS Uliler, 181)0

Carvauio — Kcys Genera Miiidac of thc World

103

KKY rO I HE GENERA OF STENODEMINI

1. líead strongly cxscrted with cyes placed near nriddle, thus far

rcinovcd írom anterior inargin of pronotuni, the distance

l)ctwecn collar and eye cqnal to thc width of one eye secn

írom above (fig. 174) 2

— Hcad not or only slightly cxscrted, the cyes in contact with

l)ronotinn or nearly so, the distance betwecn collar and eye less

than the width of one eye scen from above (figs. 175, 178) 5

1'irst antennal segment about as long as head and pronotum

togethcr; Irons strong and conically jjrodnced (Asia)

CriOROSOMELLA Horvath, 1906

— First antennal scgincnt shorter than head pronotnin togethcr;

frons not strong and conically produced 3,

3. Clypens horizontal; pronotnin carinatc latcrally (África)

NARIDOMllUS Poppius, 1914 '

— Clypens vertical; pronotnin not carinate laterally 4

4. Eyes soinewhat pediinculate; body glabrous; pronotum smooth

(Anstralia) EURYMIRIS Kirkaldy, 1902

— Eyes sessile; body pubcscent; jironotum jninctate (fig. 174)

(Américas) GOLEARIA Provancher, 1872

5. llemielytra smooth or rngosc, .sometime very finely but never

distinctly jninctured (3

— llemielytra distinctly and dccply punctured (fig. 176) 24

6. Pronotum coarsely and dceply pnnetate (fig. 176) (Cosmopo-

lilan) STENODEMA Laporte, 1832

— Pronotum itnpunctatc or only very finely or obscnrely so 7

7. First antennal segmetit covercd by long, crect pubcscencc, the

hairs at least as long as half the width of the .segment (fig.

175) 8

— l irst antennal segment (overeil by veiy short jnibescencc, thc

hairs shorter than half the svidth of the segtncnt (fig. 178) 17

8. Frons rounded atitciiorly, dcclivoiis, at most swollcn or with

:i faint l idge; vei tex with a median shallow dcinession (no true

siihus piivsetit); eyes slightly removed from pronotum (fig.

175) 9

IVoiis with a iiromincnt tnbercle or aiiteriorly; vertex with a

distimt longitudinal snleus; eyes bordering pronotum or very

near so (fig. 178) 13

llemielytia complctely glabrous and soft 10

llemielytia pubcscetit, haid and well (hitini/ed 12

9 .

104 Bor.. Mus. Gokidi — Toino XI (II) — Dc/cnibro lí)55

10. Sccoiul aiueniial scgiiieiit distinctly incrassatc towaid tlic a])cx

(Finland) ICTITOCORIS Rcutcr, 1880

— Sccoiul antcniial scgiiicnt linear II

11. Bronotuin constiicted oii anterior tliird, ilie anterior lobe

rounded lalerally (Anstralia)

AUSTROMIRIS Kirkaldy, 1902

— Pronotiim not constrieted on anterior tbird, the lateral niargins

straigbt (Índia) EliUTíUS Distant. 1909

12. Body eovered by long, erect pnbescence; ])ronolnin witbont a

niedian eonstriction dividing it into an anterior lower and

narrower porlion aiul a posterior eonvex disk; lateral inargin

of ])ronotinn distinetiy earinate; no vestige ol pionotal eollar

(lig. I?.*)) (Euro|)e, Asia, N. America)

LErrOPTERNA Fieber, I8.ã8

— Body coveied witb seini-erect, stifl bairs; pronotntn rvitli an

anterior eonstriction dividing it into an anterior lower and

narrower portion and a eonvex posterior disk; lateral inargins

ol pronotnin rounded; a narrow pronotal eollar |)resent (Ila-

waii) N ESI ÜM ÍRIS Kirkaldy, 1902

I.H. Frons protmding anteiiorly, eovering ibe base ol or tbe wbole

elypeus wben seen froin above 14

— Fions not protinding anteriorly so as to eover tbe base or tbe

wbole elyiteus wben seen froni above (lig. 178) Ki

14. First segnient ol tbe bind tarsi dislinetiy sborter tban tbe seeond

and tbird togetber (Anstralia) .. DASYMIRIS 1’oppins, 1911

— First .segment ol tbe bind tarsi as long as or aboui as long as

tbe seeond and tbird togetber 13 /

13. Body witb line, long and ereet pnbesíenee; lirsl antennal seg-

inent abont bali as long as tbe be.ad and pronotnin togetber

(índia) LASIOMIRIS Renter.. 1891

— Body abnost glabrons; lirst antennal segnient abont as long as

tbe bead and pronottnn togetber (Fnrope, Asia, Alrica, -\ns-

tralia) , NOTOSEIRA Fieber, 1838

Ki. Kyes small, ronnded. sligbtly reinoved Iroin pronoinm; bemi-

elytra rngose: rostrmn reaebing beyond apex ol bind (oxae

(java) NOTOSriROPS Popiiins, 1914

F.yes ol niedinm si/e, elongate, bordering |)ronottnn; beiniely-

tia smootb; rostrnm not readiing base ol jiosterior loxae (Fn-

rope, Aineiiea, Alrita, ;\sia)

DOI.ICIIOMIRIS Renter, 1882

Ile.id sbori ;ind llatlened, Irons siarctly protmding beyond

bases ol anlennae; tbe lirst antennal segment slender and tniv-

17 .

105

Cakvai.ik) — Kcys Cieiicra Miridac oI tlic World

ed, ihickc.st ncar base llien ta]jcring to a]K‘x wlicrc oii it cnlargcs

agaiii (fig. 181) (Kuropc, Asia, N. America)

TERATOCOlilS Fieber, 1858

— llcad long or sltort biit pointed, frons projccting sliarply

l)eyoiul Irascs ol íirst antcniial segment, which is not as

abo ve 18

18. Pronotum wiih onc central and two lateral strongly dcvcloi)cd

carinae: bead witb a median tleinession, nO longitudinal snlciis

(Enrope, Asia, Nortb America, África)

ACETROriS Fieber, 1858

— Pronotum without a developcd cential carina; bead witb a

dislincl longitudinal sidcus 11)

1!). First segment of bind tarsi shorter or etpial to third; body

witb leddisb areas (Índia) ZANESSA Kirkaldy, 1902

— First segment of bind tarsi longer tlian tliird; body witbont

reddish areas 20

20. Rostrnm e.xtending to base of abdometi; first antennal .segmctit

as lotig ;is bead and pronotum togetber 21

— Uostrnm not reacliitig beyond midille coxae; first atitcnnal seg-

mctit as lotig as bead 22

21 . First tintetnial segment as long ;is bead and ])rf)notnm togetber;

frons ])i()rln(ed iU base of clypens (Closmopolitan)

MEC.ALOC.ERAEA Fieber, 1858

— First antennal segment shorter tban bead, fions not prodnced

at base of chpens (.Vfrica) NYMANNl/S Distant, 1901

22. Protiotnm distinclly carinate laterally 28

— Pronotum tiot caiinate laterally: bons piodnced into a pointed

tnbercle ( Fastnatiia) PJ{()'E()M / RIS Po|)j)ins, 1911

28. Ilitid tibiae witb long erect i)nbescence: frons smootb, flat;

first antennal .segmetit almost glabrons (fig. 288) (África) . . .

SCIIOUTEDENOMIRÍS (larvalbo, 1951

— Ilind tibiae witb sbort j>nbescence: frons witb a point or |)ro-

minence; lirst antennal segment distiiutly pnbescent (figs. 105,

178) ((:()smo|)olitan) rRIC.ONOTY/Ars Fieber, 18,58

21. Posterior tibia very long and strongly pilose; bracby|)terons,

tbe bctnelytta witbottt divisiotis (Inaii Fernatule/,)

KVSCUEI.IANA Carvalbo, 19,52

— l’osterioi tibiae not as above; maci optei ons 2;>

25 Rostrnm leac bing to or beyotid tbe postei ioi (oxae (.Xnictitas)

OrirniAI.MOMIRIS lUrg, 1888

— Kosirnm not (|nite reacbing middle (oxae (Atneritas)

' roRROMIRIS berg, 1881

106 líoi.. Mus. Gof.i.i)I — J onio XI (II) — Dc/cmbio 19.55

Ki: Y ro riiE gknera oe iiyalopeplini

1. Proiiotiini distinctly and coarsely pnnctatc (íig. 218) .... 2

— 1’ionotum iin])unctaie, somctimcs nigosc or only lincly jjimctu-

latc (fig. 225) 5

2. Collar punctatc with mcsal Icngth ccpial to half ihc width of

onc cyc (fig. 218) 3

— Gollar not pnnctatc and not as wide as above 4

3. Scntcllnni sniooth above and stiongly clevalcd (fig. 218) (Ilor-

nco) MACltOLONIDEA Hsiao, 1914

— Scntcllnni pnnctatc (Malay) .... MAC.ROI.ONIUS Slal, 1870

4. Elead strongly vertical; rostnnn rcaching tbc 7tb or 8th obdo-

ininal seginent; eycs very large, occnjrying tbc wliolc sides o£

liead; lorae strongly ])ronnnent (Malasia)

KOSMÍOMIRIS Kirkaldy, 1902

— Hcad not strongly vertical; rostnnn not extending beyond a])ex

of bind coxae; eycs not occiipying tbc wliole sides of hcad and

lorae not noticeablc proininent (Malay)

CAIRYSORRUANIS Kirkaldy, 1902

5. 1’ronotnin coarsely nigose transvcrsally (fig. 225) 12

— Pronotnni smootb or veiy fincly pnctnl.ite 6

6. 1’ronotnin beset with innnerons sliort bristies; first antcnnal

segnient incrassated towards base and ajtex 7

— Pronotnni witbont sboit bristies; first antcnnal segincnt

linear 8

7. A sinall tidtercnlar flat |)rocess between inferior niargin of

antcnnal .socket and eye; genae not caiinatc; antcnnae in-

crassated towards apex; large, elongate species (Ilritisb Cbiia-

na) IRIDüPEPEUS Ilergrolb, 1910

— ^\'itb tlie small tnbeicnlar protess above; genae carinatc on

iipjjer niargin; antennae incrassated towards tbc base; ine-

dinin si/ed species (Manritins)

CORI/.inoEON Rcnier, 1905

8. Hody polisbed glabrons; pronotnm more or less (arinated late-

rally (África) PI.EVROCIIII.OPflORVS Renter, 1905

líody pnbesceiit; pronoinni not carinateil laterally 9

9. First anttmnal segnient inncb sborter tban widtb of bead. tlie

latter strongly vertical ;ind transverse; tbe eyes very large, oc-

(iipying tbe wbole sides of bead, (oniignons witb pronotnm

(New (biinea) MOROCA Poppins, 1912

— l iist antennal segnient longer tban widtb ol bead; eyes not

otdipying tbe wbole sides of bead. renioved from pro-

notnm 10

C^ARVAi.iio — Kcys Gciiera Miridae of thc World

107

10. Pronotiiin vcry lincly {)imctidate; first antcnnal scgment alniost

twicc as long as width of hcad (Philippines, Siiniatra)

RA MB EA Poppius, 1912

— Proiiotmn siiiooth; first antennal scgnicnt slightly longer than

width of liead 11

1 1 . Pronotum strongly constrictcd anteriorly; body glabrous

(lUinna) ONOMAUS Distant, 1904

— Pronotum not constrictcd anteriorly; body j)nbescent (Índia,

Philippines) GUIANERIUS Distant, 1903

12. Gorinm without veins 13

— Coriínn with veins (Índia, Malasia)

ISABEL Kirkaldy, 1902

13. Emboliinn and cnnens distinctly pilose (New Gninea, Philip-

pines) IIYALOPEPLOIDES Poppius, 1912

— Endroliinn and cnnens glabrous M

14. Clavns distinctly pnbescent; first antennal scgment longer than

width of head (Índia, Samoa)

GUISARDUS Distant, 1904

— Clavns glahrons; first antcnnal scgment shorter or abont as

long as withh of hcad lõ

15. First antcnnal .scgment thickcr at hasc; hcad vertical (fig. 225)

(Mallaca) EUIIYALOPEPLVS llsiao, 1944

— First antcnnal scgment iiuiassatcd towards tlic apex; hcad not

noticcably vertical (África, índia, Malay, Pacific Is.)

IIYALOPEPEUS .SUil, 1870

KEV K) EME (lENERA OE RESTUENINÍ

I . .Scntellnm strongly convex, witl» a longitudinal basal snlcns

or impression: til)iac as thick as tlic femora, compressed, snicatc

on both sides or infcrioily (üg- -IIH) (South America)

RESTUENIA .Sjnnola, 1837

— .Scntclhnn not stiongly (onvcx, ncithcr snleate no impicsscd

at l)asc: tihiae cylimhical, not snleate 2

2. Pronotal {ollar not rcaching thc sides of pionotnni. lhe pio-

notnm strongly carinatc antl prodnccd aiiieiioily bcyond sides

ol (ollar so as to cnclosc thc latlcr (lig. 251) ((^cnti.d K: .South

America) MIMONCOPEI.TUS Kirkaldy, 190()

— Pronotal collar ícaching thc sides of pronotnin, thc lattcr not

OI mmh Icss strongly carinatc ;is above (lig. 220) 1

body oval, vcry wide; hcmiclytra widened latcrally, distinctly

wider than iironotnm at base (fig. 210) 4

3 .

108 lioi,. Mus. CioKi.Di — loino XI (II) — Dc/cnibio 1955

— liody clongatc or obloiig, parallcl-sidccl; hcndclytra larcly

widcncd at iniddlc, usiially j)arallcl or ncaicjly so, not or only

sliglitly wider than |)ronotiiin at base 5

‘1 . Sccoiid aiiteniial segmeiit tliieker tban lirst; tibiac rvitliout

spines (.South America) .. EURYSCYTOVIIORÁ Retiter, 1909

— .Second anteimal segment mote slentler than íirst; tibiae very

siiort (Chile) EURYLOMATA Reuter, 1909

5. Pionoium emarginate latcrally and postcriorly (>

— Pronotiim laterally nearly straight or slightly rounded, some-

times laintly simiate behind collar but never at middle . . 7

0. Pid)esceiu'e very short and scanty; anterior coxal clelt seen

írom above; slightly antmimic (Hra/il)

KAMAÍURANA Carvallio, 1952

— Pidte.scence distinct and altiindaiu; anteiior coxal clelt seen

Irom above; not antmimic (Argentina)

IIETEROSCYTUS Reuter, 1909

7. Ilead including eyes ccpial or only scarcely wider than collar;

pronotnm distinctly carinate lateially; second antenmd segincnt

ccpial in thickness to the lirst segment, linear (lig. 220) (South

America) CIIÍEOXIONOTUS Reuter, 1909

— Ilead including lhe eyes distinctly wider than colhir; pronolum

ir carinate hitcrally, only at iinterior portion and in this case

with second antennal .segment more slentler than lirst segment

or incrassate towards the a])ex (ligs. 217, 227) 8

8. Uody with hrillianl metidlic s|)ots or aie:is; tibi;ie strongly in-

crassate towiirds the ápices with densely subadprcssed pubes-

ceni (.South America) I.AMESOEI ÍORVS Reuter. 1909

— Uody without biilliauí metailie spots or areas; tibiae not

strongly incrassate towards their apites, the pubesteiue more

or less erect 9

9. Pronotnm distinctly setose (true setae) (lig. 219) 10

— Pionotum glabrous, linely pubestent or shoitly pilose, but

never setose (figs. 217, 227) II

10. Antennae and legs witii unilorm short pubesteiue, large species

over 12 mm. long (fig. 219) (.South Aineric:i)

CAEEICIIII.EI.LA Carvalho, 1955

— • Antennae and legs with numeroiis long set;ie in adilition to

the short pubcsccnte; snialler speties, less than 10 mm. long

lig. 209) (lha/il) MAIIELIA Kiikalily, I90.S

11. Pioiioliim punilate or toarscly rugose; botly mirrow, siib-

glabroiis; iibi:ie with spines; (iineiis ol mat it>|)ierous loims

more than iwite as long as wide at base (Chile)

S EEXOrAREDRA Reuter, 1909

C^ARVAi.iio — Kcys Cicncia Miridac ot thc World

10!)

— Pronotum smootli, body iisiially not narrow, W so thcn thc

tihiac pilosc: cuiiciis ncvcr inorc tlian twice as long as wide

at base 12

12. First aiitcniial scgment sbortcr tbaii widtb ol vcrtcx; .sccond

scginciit more tban thrcc times as long as lirst (lig. 211) (Nortli

Central America) ONCEROMETOPUS Reiiter, 1875

— First tintennal segment longer tban widtb of vertex; second

segment abont twice as long as lirst l.i

1‘5. Pronotum distinctly caiinate laterally bcbind thc collar, on

onter side ot calli; tlie sccond antennal segment nsnally in-

crassate, as tbitk or thickcr tban first segment; species of large

si/e, iistially ovcr 10 mm. long (fig. 227) (Central & South

America) PEATYTYEUS Fieber, 1858

— Pionotiim not ilistinctly carinate laterally, as above; sccond

antennal segment more slendcr tban the first, if incrassatc, tbcn

the base tbinncr tban first segment; specics ot metliuni or small

si/e, nstially less tban 10 mm. long (fig. 217) (Américas) ...

PREPOPS Renter, 1!)05

KFY FO FllF C;F.NFR.\ OF IIERDONIINI

1. Scntellmn wilb an erect sj)ine-like projection (fig. 213) .. 2

— Scntellmn smootli, flat or convex, withont a spinc-likc pro-

jedion 8

2. Pionotnm stiongly constritted at midtllc, the anterior portion

Hat and bori/ontal (figs. 25F,252) (Central America)

ZACYNTIJUS Distant, 1881

— Pronotum not strongly constricted at middie 3

3. Ilead witb a sbort neck, narrowcd basally; eyes sejtarated íroni

pronotum by a distance abont eipial to tbe Icngth ot onc eye

(fig. 21.3) ■!

— Ilead witiiotit a sbort neck, not narrowcd ba.sally; eyes bordei-

ing anteiioi maigin ol pionotnm (lig. ;>

■1. Cnnens impertepiibly merged witb membrane; rostrnm reatb-

ing middie toxae (lig. -13) (.Sotitb Americ;i)

UERDOMUS Sliil, I8(i0

Cnnens absent; rostrnm reacbing anterior co.xae (fig. 3!))

(.Sontb .Vmerita) (iLIARA\E'í C.aivalbo Sj Cbin.i, l!)al

5. Prcnoial (oll;n distimt (lig. 2-2)

— Pionolal (ollar absent oi indistinct 7

(i. I egs witb long, ereit, wbite biislles; tbe bind libi.ie sii.iigbt

(Sonih ,\meii(a) ILIARUPIOEA 1 oj)|)ins, l!)_l

110 Koi.. Mus. Goei.I)! — lomo XI (11) — Dc/cinbio lí)L.5

— Lcgs without loiig, crcct wliite brislles; liiiul til)iac fairly curvctl

(South America) HAAltUPIA P<)j)|)ius, 1921

7. Vertex sulcatc longitudinally; calli not distinct; bind tibiao

fairly curvcd (fig. 177) (South America)

FIEBIUGIELLA Poppius, 1921

— Vertex not sidcatc lougitudiually; calli distinct posteiiorly;

hind tibiae not curved (Américas)

HAllIiERIEELA Poppius, 1911

8. Both sexes brachy|)terous; eyes distaut from pronotum by a

spacc etpial or more thau leugth of eyc (fig. 255) (Europe

Asia) CAMPONOTIDEA Reuter, 1879

— At least oue macropterous; eyes mucli closer to pronotum or

contiguous with it 9

9. llemiclytra glal)rous or with short, adpressed pubesceuce only,

never wiili long erect bristles 10

— llemielylia witii long, erect bristles, sometimes intei luixeil

with semi-erect pubestence 15

10. Pronotum raisetl posterioily into a spiiie like, erect projectiou

(North America) DACERl.A (lignoiet, 1887

— Pronotum not as above ; II

11. Posterior femora with long, erect bristles; posterior tibiae with

long spines and small, dark tuberdes (fig. 220) (Américas) ..

PARAXENETUS Reuter, 1907

— Posterior femora witiiout long, erect biistles; postei ior tibiae

with only a short puhescence or spines 12

12. Hody glabrous; head with a short netk (Holivia)

ACEdlMA Pojipius, 1921

— Hody pubescent; head without a short netk 15

I. 5. Eyes removed from anterior margiu of pronotum; thc latter

strongly constricted in mitldle (Afrita)

SPUíNCrOTIIORAX Stiil, 185.5

— Eyes (ontiguous with pronotum, the latter not strongly tons-

tricted in mitldle II

II. First antennal segment starccly leaching the ajn-x ol head

(North África) EAURIMA Reuter. 1881

— First antennal segment reathing tlistiiutly bevonti the apex

of head (África) XEN E.POMORPU A Poppius, 1912

15. Stulelium strongly ttmvex, with a pitmiineni blunt meilian

elevation :intl long. eret t setae 10

— .Stulelium, ii tonvex, without a metiian blunt eleviilion 17

Ki. Rostrum stanely surpassing mitldle ol mesosteinum (.Stmih

Amerita) \/./A)MMA PI S Reuter, 19117

111

(Iarvauk) — Kcys Gcncni Miriclae of the World

— Rostrimi reaching the base of posterior coxac (Central Ameri-

ca) ZOSIPPUS Distant, 1883

17. Pronotal collar iiot visible, indistinct (Américas)

XENETUS Distant, 1883

— Pionotal collar distinct (South America)

LEPIDOXENETUS Poppins, 1921

ACKNOLEDGEMENTS

riiis paper is in many ways complementary to that jtnblished

in the “Anais da Academia brasileira de Ciências 24 (1):31-11(),

1952”, and the acknolcdgemcnts mado there in scill aj)ply. In

atldition to the iriends and colleagncs mentioned in that paper

the anthor wishes now to express his gratitiide also to Mr. T.R.E.

SotiTiiwooi), Rothamsted Exjx.'rinient Station; L. KiaaoN, Depart-

ment of Agricnltnre, Ottawa; E. Wagni.r, Ilandmrg and D. Lf.stün,

l,ondon, who have more recently helj)cd with iiscfid criticism of

parts of the j)resent paper.

riie illnslrations at the end of the paper werc tlrawn by Mr.

Antonio Vir.GAs Piigas nnder the anthor’s snjjervision .

cm 12 3

10 11 12 13 14 15

I•lalc I

Tarstl (laws: I — l'ulvius hrunncus. 2 — Cylapiis (cniiiiitornis. S

Dcracix ()rÍ5 riihcr. ‘í — Kury(hil()|>(crclla liiridiila. 5 — 1’sallus

aiKorifcT. I> — Monosyiiamma boliciiiaiiiii . 7 — Khinocapsus vaii-

(luzcci. 8 — Rcutcrosíopiis «rrialiis. 9 — l,<)pus (Ictdlor. 10 — Ma-

(Totyliis sfxgiiltadis. II — Cocpiillclia niinictica. 12 — Dityplius

(li^rcpaiis. 13 — Diiyjiliiis famclicus. 1-1 — Syslclloiiolus triguttatus.

15 — llall(>(la|>iiii s]». 10 — IlalUidapiiü corizoidcs. 17 — l‘yaiodcrcs

dilataliis. 18 — .Spartaius alliatiis. 19 — Siroiigyloforis stygiciis.

20 — I.ygtis vandiizcei. (1-13 and 19-20, aficr Kiiiglit).

IMatc I

21 — llnitfllular inrinbninc of llryocorinae . 22 — llitrlltilar iiicin-

biaiic of Mirini. 2,1 — Tarsiis of Stcnodcina . 21 — Ostiolar pcri-

Iri-iiic of Kc.sllirniiii (Prcpops) . 2(> — Ilrad and pronoliini of (li,

viiu-mini (Ambrncius) . 27 — ilcad and ()ronolinn of OdonicIIini

(l’arabryocoro])sis) . 28 — Tarsus of IlrymoriíiL' (Ncclla). 29 —

Tursns of Mirinar (IIor(ia.f), 10 — Ilcad and pronotinn of Mona-

loniini (1’oppiiisía) .

riatc III

31 — l’r(inotal (ollar <ií Miritiac (C.alocoris) . — 32 — l)t'|iicssf(I («illar

<>( Orthotyliiiac (Ortiiotylus) . 33 — llcad aiul pronoliiin of l*Iiylini

(IMagiogiiatluis) . 31 — Pronolal collar ol Orthotyruiac (Cyllccoris) .

35 — Pronotum of .Stciiodciiiiiii (I.qdopicriia), sliowing llic jiro-

iiiiiicnt latrral rigdc cliararicriütic of llic Sirnodrniiiii (aflcr

Kiiight). 36 — l’rniio(al collar of Ortiiolylinac (Cyllccoris). 37 —

l’roiio(uin of Phyliiii ((ionostcllnis) . 33 — llcad and pronoliiiii of

.Mirini (Calocoris). 3!) — lailcral viccv of ilcrdoniini (Cuarania).

1’liitc IV

•(() — llcad of Oithotylini (Orlliolylus). 11 — rroiioliiin aiul lirad

of 1'iilviini (Fiilvius), aflcr Kiii^lit. '12 — llca<l aiul proiioUini «f

«f Dirypliiiii (l)icyi>hiis) . 4,'I — Ilt-ad of Ilaltiiini (IlaUicus). -ll

— Ilocly of iMctisloiscclini (Mystilus). 45 — l’roiioHiin and lirad of

Miriíii (floreias). 4(> — I’roiioluiii and hcad of Trrmatojdiylini

( I crniatopliylmn) . 47 — I’roiiotnm and hcad of Kcslhcnini (l’rc-

pojis). 4K — Hcad of Cylapini (Cylapus).

rialc V

4!) — Oylapus iciiiiiiiconiis Say, frontal vicw (Afctcr Kniglit). 50

— First aiiteiiiial scgniciit of OdoiHoccrocoris ornantUis (I)istaiit).

51 — Ilcatl of Rliiiioiniris cainclus Poppitis, sidc virw. 52 — llrad,

|>roiiotiiin and aiitrniiao of Aniioiia l)iina(ulatn (I)istaiit). 53 —

Ilcatl of Fiilvitis liisbislillatus Stal, sidc vicw. 54 — llcad of Fiis-

lictiis Kciitcr. 55 — llcad of Eiiryrhiloptcrclla liirida Kciitcr. 50

— llcad and pronotnin of Carijoatuis ruhcrfascialus Carvallio. 57 —

Ilcatl and pronotnin of Antia.s cliilcnsis Carvalho. 58 — llcad and

antcnna of Aucluis hrasilicnsis Kiiight & Carvalho. 5!) — llcad and

first aiitcnna of Pscutlocarnus fraudans (Stal). 00 — llcad and pro-

notnni of Guanaharca angrcnsis Carvalho. 01 — Ilcatl and prono-

tnni of '1'rygo iinitatíonis Distant. 02 — llcad and pronotiim of

Amhracius diifonri Stal, sitie vicw. 03 — Ilcatl anil pronotnin of

(iuanaharca angrcnsis Clarvallio, sitie vicw. Iil — llcad and first

antcnna of P.scntiocarniis Distant.

SciELO i'o

rlatc IX

120 — IMiysoplioroplficlla tlcnlitollis I’<)|i|>iiis, sidc vicw. 121 —

IMiysoplioroptcra iiiiraliilis Poppiiis, .sulf vicw. 122 — raiafiiiiiis

(liscifcr (.Slal), .socii rrnin abovc. I2.i — llcad of DichuxKoris inalai-

sei Carvalho. 121 — Panlilioniorfa iinprcs.sopuiKiala .Sclniinatlirr.

12.'} — llcad and proiiotiiiii of .Sicnopicrocoris Ialiceps China. 121»

llcad and pronoliiin of llallirolonia valida UciKcr. 127 — llcad

and pronolinn of .Sincrviis hyalipcdcs Carvalho. 128 — 1’arabryoco-

ropsis lypictis China & Cairvalho, secn froni ahove. 12!) — llcad

ainl pronotinn of Pscudodoniclla pacifica China & Carvalho. I.'l()

— llcad and anienna of Villicrsicoris liolasi Dclaltrc. 131 — llc-

iniclylra of Tcnlhccoris orchidcarnin (Rculcr). 1.32 — llcad anil

pronoinin of Mccolaciniis carvallioi (Caista Lima). 1.33 — llcad and

pronolinn of Cyitcxapsiis caligincns (.Slal). 1.31 — llcad of Sysinas

paliidl|>cs (Sliil). 13.3 — llcad of .VspidolKilhrus flavicosla Carva-

lho. 1.31» — llcad of Knrycipitia splcndcns (Dislanl). 137 — Colco

ptcrciiniris siniilaris Càirvalho, sccn froin ahove.

IMalc X

138 — Ilciul and pronotiini of Monalonion xanthophiliiiu Walkcr.

139 — IlcinU-lylra of 1’oppiusia conibrctonim China. MO — Ilcad

of KnightíHoris r.arvallio íC- China. Ml — Iloniiolylron of Spartacus

alhalus ni.staiit. M2 — ÍU'a<l an<l pronoinin of llryoioris ploridi.s

(Fallcii). M3 — llrad and pronoltini of Caiilotops piinrticollis Kcr-

grolli. Ml — Ilcad and pronntuni «f .Stictolopluis hicolor Cana-

lho. lia — Tihia of .Sahlhcrgclla singularis llaglnnd. M(i — IIc-

niiclytron of Nconcclla niilzac Ciarvalho. M7 — Tihia of Distaii-

ticlla thcohroma (Oi.slani). M8 — Ilcad and pronoliim of l’ycno-

deres 4-inatulatus (Cuèiiii & iMcncvilIc). 119 — Hcniiclytioii ol

.Siiicrvils harenspriingi Stal. 150 — llciniclylra of Neofnriiis Distam.

151 — Ilcad and pronoliim of I.ahops hirtus Knighl. 152 — He-

miclylron of Prodroimts thaliae China. 1.53 — Ilcmiclylron of

llothrophorcila nigra (.Slal). 151 — Ilcad of Cyrtolylus nihricaUis

llcrgrolh, side vicw. 155 — Ilcad and prnnotiiin of /.anthiiis fra-

gi|is IJsingcr (Aficr li.singcr). 1.50 — Ilcmiclylron of Mclariirins

Canalho 8.* China.

in? — :in(l pronoliim ol (:ain|))'l<)n('iir;i viigiila (llcrrUli- .Scliacl

frr). 158 — /jkaniola cicgans C.arvaIho sccii fioin almvi'. 15!) — Stiilcl-

liiiii of I Iclopcllis .SigiiorcI, .Hidi- vicw. KiO — llcad and pronotiiiii

of Oiuonolclliij huxioni Kniglil. Ilil — llcaul and proiiotiiiii ol

Ma( rolopliiis niibiliis (ll<'iTÍ('h-.S(lia('rr('r). Kill — llcad and |>rn'

notnni of Apolkxiotidra y signala llviao. I(>;{ — (:niu'n<i and nicni

branr of C.liíus niaculatiiü DivIanI. Kil — llrad ainl pronotnin of

l■■l■lisacn^ «nliraícns mingcr, I(i5 — llcinclylron of lIcUTotoiis

dilalatiis ((>iK'rin-Mriu'villr) . Ililí — llcad of Kbopalisccdiains Kcii-

ler. Ili7 — lload and pronotnin of ( '\i lopcilis inodcNta (l)ixlant).

1(>8 — llcniiclylron of l•',lnb(llio(l)|•is ('.^irvalbo &.• ( liliia. Ili!) — lli-ad

and (ollar of Vatngainbia inaiarangac ((Ibina). 170 — llcad ol

llcspcrolabops kiikaldy. 171 — llcad and pronotnin of Nccila

IntcMcnr (.Stal). 1711 — llcad and pronoinin of Monalocoris filieis

(l.innaciis) . 17.1 — llcad and proiioinni of Noioliciiialcs ('.arvalba

K- (liina).

l*lii(c XM

171 — Ilucd ;iikI proiioliim of Colhiria scctiica (Stal). 175 — lli'a<l

and prniiotuiii ot Lcplotcrna dídabrala (Liniiac-iis). 17(i — llcad

and prnnotiim of Stcnodoina insuavis (Stal). 177 — Tilda «f l'ic-

brigiclla luimcralis 1’oppiiis. 178 — Iload and pronotiiin of Trigo

notylus riifitornis (Cícoffroy) . 17!) — llcad and anicnna of Ciargamis

gracilcntiis (Sliil). 180 — llcad of Atomosccliíi OTiustiis Ficbcr, sidc

vicw. 181 — Tibia of Kllenia cuncata (Stal). 182 — llcad of 1’arc-

drocoris pcitoralis Kciitcr. fS.') — llcad of Calondas fasciatus Div

lant. 184 — llcad and pronotnni of Tcralocoris saiindcrsi. 185 —

Dcroplitlialnia minúscula C!arvaliio, .sidc vicw of bcad and pioiio-

tuin. 180 — llcad of lladrodcmus inflavum (Goc/c). 187 — llcad

of (>ry|>ucoris coslai Uciilcr. 188 — llcad and pronotiim of 1’amcri-

fica inarlollii Uciilcr. 18!) — .Sciilcllnni of Chilcaia ('.arvalho. 1!)0

— l*ronolum of Ncoborus glabcr Kniglit. l!)l — llcad and proiio-

liini of 1’byllopidca picla (llbicr). I!)2 — llcad of rantilius luni'

ralus (Fabriciiis) . I!),i — llcad of Caloforisca villosa Distant. l!)l

— llcad of Kubatas chirifpiinus Distant,

rialr \lll

l!)5 — llciul «f Piasu.H ( ribricoilis (Slal). HMi — Aiiictina of l’a-

thymcrcKCTiis erylhronotus (Ilrrg). 197 — IIi'a<l and aiitrnna of

('.a|)sus ater (Liniiaeiis) . MW — llead and anienna of Crtonliadfs

Di^lant. 199 — (dlianclla inarniorata CUirvallio, M-rn froin alH>v«‘.

20(1 — lli-ad and first aniennal srgnu-nt of Clapniariini Disianl .

201 — llcarl, pionoinni and anIenna of Sidnia kcllcrsi Kniglit

202 — Ifead and antcnnai' of l'.iiro(aH nodosiiH I)i>iant. 20.'l — Ilead

and first anienna of 1’oeas reiiteri Distam. 201 — Ilead and jiro-

noliiin of 1’lesiülygiis jiiinetatiis Kniglit. 20.5 — Uead and pronoinni

of Monaloiorisea ((inspiirrata DistanI. 200 — Ilead, pronoinni and

anienna of I.ygidea nicndax Keiiter. 207 — Ilead and pronotiini

of laedia neotropiralis lleigrolli. 20S — Ilead, pronotnni and an-

ienna of l-ygns (Neolygus) loinniiinis kniglil. 209 — Ilead, eollar

and firsl antenna of Malielia piileherriina Kiikaldy. 210 — I’ro-

notnni <d linginotop.sis 1’oppiii.s. 211 — Ilead and first antennal

srgnient of I.ainpethusa anatina Dislan. 212 — Ilead of llyporrliino-

eoris tonienlosiis Keiiler, side vicw.

riiiu- XIV

213 — llciid anil iironotiiiii <>f llcriloníiis annatus Stal, sidc vitw.

211 — Ilcad and first aiilriinal srgniciit of OiucrnniclopiiH inipíctiis

Kniglit. 215 — Ilcinirlylroii of Angcrianiis rradii!) Distaiit. 210 —

l■.llry!l(y(o|)ll()^a lalorallis KcuUt, mtii froin abovo, 217 — Ilcad,

liroiiottiiii and antcnnar of IMatytyllus Uciilrr. 218 — l’i'ono(nin

and bcad of Macrolonidca (yancKCcns Ilsiao. 21!) — Itcad ainl an-

tcnna of Adcipliocoris lincolatus ((íoc/c). 220 — Ilind fcninr of

raraxcnctiis gnlliilalus (Uldcr). 221 — Ili-ad and pionolinn ol

Kiainbura (offcac China. 222 — Ilra<f an<l inonoliiin of llaanipía

.xpinoHa I’op|>iii>i. 22,3 — Ilcad, pronoluni ainl first (ihia of llcni

cotiicmis alIxKirnatii.s ,Slal. 221 — 1’ionoliini of Darcrla inflata

(llldrr). 225 — Pronotnni of ladiyalopopliis piilchclliis Ilsiao. 220

— Ilcad and pinnoliini of C.hiloxionotus nigrosc ulcllatiis Carva-

lho. 227 — l’ronoti:ni, lirad and anirnnac of 1’lalylylliis 1'ichcr.

riato XV

22K — llcad and pionntnin oF IJc-lcaiia liiiigiiaiiicala Carvalho.

22!) — llrail and pronoinin ot Myoinbca C.hina &: Carvalho. 2.‘)0

— Aniinna of DriUliinarus Di.slant. 231 — Ilcad and pionntnin of

Aztccariclla (áirvalho. 232 — Uppi-r sidc of 1’critropis saldarforinis

Clilcr. 233 — Ilcad of SchoiilcdcnoinirÍ!i íáirvalho, sidc vicw. 231

— Ilcad of I-ojiidca IJhIcr, sidc vicw (aficr Knighl). 235 — Ilcail

and pronotinn of Kanrovius Distant. 230 — Ilcad of Atroirhiniinn

Nnnalliicr, sidc vicw. 237 — Ilcad and pronoinni of Kunungiia

('.arvalho. 23H — Ilcad and antrnna of Nciirocolpus tiliac Knighl.

2.3!) — Ilcad of Chainus tiihcrciilatns (l)islant). 210 — llcad, pro-

iioinni and anicnna of Ilalticiis bractcallis (.Say). 211 — Ilcad anil

pronninni of Malgacbcocoris ('.arvalho. 212 — Ilcad and pronoinin

of .SctiKoris China & Cairvalho. 2'I3 — Ilcad and pronotiiin of

ralilíaiiaiia Cairvalho. 211 — Ilcad of Vannilis Distant.

riatc XVI

'Zi!i — llrad and proniiliiiii of I’ara(y1a|)us Carvalho. 2'U) — Ilcad

and |ir«no(nni of Ycbonia (jirvallio. 2i7 — l’ronoluni of Zacynthlis

Diviant. 2111 — Ilcad of lúirythilclla Kcntcr. 21!) — l’ronoliiin of

C.allichila Kciitcr. 2.50 — .Srutclium of Knightoiiia, lateral vicw.

251 — Ilcad and |)ronotuni of Miinoncopclttiv Kirkaidy. 252 —

Ilcad and |)rnnotnni of /.acynthus (after llistant), 25.5 — Ilcad of

l■■uscln Distant. 2,51 — Ilcad and pronotiiin of Myriiiccoris Corski.

2.55 — Ilcad and pronoluni of Cainponotidca Kciitcr (after Ilohcr-

landt). 2.50 — Ilcad and pronotinn of Ilalludapoidcv Carvalho.

257 — Ilcad ainl pronotiiin of Cystcorratlia Kirkaidy. 25H — llrad

ainl pronotnni of PcruantKoris òirvalho. 259 — Ilcad of I ladro-

ncina l/hirr (after Knight). 200 — Ilcad, pronotiiin and M-ntrIhini

of Kcvtliciiia .Spinola. 201 — llrad and pronoinni of l'il()|>honi»

WrstwtMMl. 202 — Ilcad and pronotinn of I’sciido\cncltis Kcntcr.

205 — Ilcad and pronotiiin of Diaphnidia IMdrr.

Ahibnliis

. . 59

INDEX TO GENERA

Aiistopephis 99

C.umpyloiieura . . .

. . 58

Acaiilhoriannlld . .

.. 101

Artliriticus

.. .38

Ca mjfylon euro p.sis

. . 58

Acantlioj>cj>his . . .

. . 9.1

Asciodema

. . 53

Campylolropis . . . .

. . 77

Arcgimn

.. 110

Aseiymiis

. . 78

Canariocoris

. . 77

Arclroliis

. . 105

Asjiidacaiilhus . .

.. 02

Capdanus

. 40

Acrnihcus

. . 07

Asjiidobolhrus . . .

. . 33

Capsodfs

. . 80

Acrodcrrhis

. . 70

Alomopliora ....

. . 57

Acronhiiiiitiii ....

. . 03

Atomosrclis

.. 17

Carmdus

. 20

Aftiuimotus

. . 03

Atrarlotomus ...

. . -15

Carijoanus

. 21

Aclilocoris

. . 101

Aurhciincrel>is ...

. . 50

Caulolops

. .31

A(lelj>)iocori(lcn ..

. 91

Auchus

. . 21

Cephalocapsidfti

. 49

Adeljiliocoris ....

. 91

Aushorolisus ....

.. 101

Ccphalocapsus . . .

. 40

Adinrius

. 23

Auslromiris

.. 101

Ccratocapsus

. 08

Atolocoris

. 01

AiisWiijirfdus ....

. . 87

Ct-ialnfulviu.s

.' 20

Arthorhiurlln ....

. 71

/lííztu

.. 01

Chiutorapsus

. 02

.Ípi.v/ic/uí

. 28

Aztecariella

.. 29

Chaiiitipsis

. IO

Aí^iwroris

. 99

Hakrriola

.. 18

Chain us

. 41

Agmtticirn

. 11

Itarhrriclla

.. 110

Charag^ochilus ....

. 95

AgrnlHoioris

. 50

Ilertsa

. . 95

Chdlocapsiis

. 85

A Ida

. 101

Ilihuiidiclla

. . 01

Chileaia

. 100

Atcliidin

. 81

Ililiduiijtulus . . . .

.. 71

Chiloxiimoliis

. 108

Alijiididla

. 81

HironicUa

. . 18

Chinamiris

. 80

Allorix hnts

. 97

Illcidiaridopinus

.. 75

Chius

. 59

Alloríiiiihiii.s

. 02

líoUcria

. . 99

Chhimydatus

. 49

Alloctiiioliis

. 88

Iltttipiddla

. . 01

Cliorosoindla

. 10.3

AUorolarsus

. 51

Iloopidocoris

. . ■! 1

C.hrysodiiiodc.s . . . .

. 40

AUocoIoiiims

. 28

Ilolhritimiris

, . 18

Chiysndasia

. 91

AlUíimiinlas

. 110

Ilolhropliordia . . .

. . 33

Chiystinliaiiis

. 100

Allíirliiiitirons

. 80

lIlltliMUlIUS

. 23

Cimiiapsus

. 27

A niítztnutcoris

. 01

Iliixia

.. 12

Ciniiaiiius

. 21

Aiitl)l\l\liis

. 19

Ilradiyarllirum ..

. . 51

Cixatiiiis

. 90

Am hl tuins

. 23

llradiyroints

. 88

Clapniaiius

. 93

Amixia

. 78

Ilradiyrrniirlla . . .

. -17

Clixáiifiiia

. 23

Amiihit alisiis

. 101

ltradi\luh’ÍH.s ....

Clo.sltrocoris

. 03

. 38

Aiialms

Jtradiyiioloroii.s . .

. 77

Ctihaloihyiidius . . .

Aiif^nianiis

. 00

lira.siliomiris

. 71

Cocnihaplirs

Aiiiiirssii

. 27

Ilriimrlianiiiris . . .

. .33

('iilrtiplnoiniiis . . .

. i;9

Aimuiia

. 25

Iliyiiriiris

. 3.3

Collaiia

103

Aiiiimaliiiiinius

. 13

Ilryocoriipsis

. 11

Ciiiiipsoiiaimus ....

. 51

Aiiiisihra

. 81

Iliirohia

. 59

Cumpsiiscylu.s

Aiilhiiilitilduinitilfs

. 82

Iltiiisiia

. 09

Ciiiiiirrphahiriiris . .

A 11 lias

. 20

Ilulliiifiirlla

. 98

Ciiiiiisirllius

A/iaiili/iiis

('.allifliilrlla

C.iiijiiilldlia

A lihanioldixn

Caliirori.s

. 91

CoiTOfadiinila

Afihauosama

Valorurisra

. 91

Coridromius

A liidliidiitidfa

C.alormopsis

. 91

CoihidoUiii

100

Ajiiilypis

101

Caitmdas

. 98

Cranocapsus

Arasfiiis

Cahplodria

. ■ 95

Crassiciiniiis

Iini/aniis

Campiiiiiitidra

. 110

Cirmiiocrplialus ...

o,i

Aiffniis

Ciimplolylus

. 52

Ciriiiiliadr.s

AiKyrmiiiis

C.amplir.yiium

. 90

Ciriiinoirhiiius ....

•15

■filisiidrlanius ....

Camiiylomma

Ciiocoiis

SciELO

10 11 12 13 14 15

1'tH ]k )i.. Mus. (ioKiDi — 1'onio XI (II) — I)ezcinl)ro 1955

CychrocafKxux . . .

Cyclxdoloxx

... .35

Cylapoidcx

. .. 21

('•yhxpocorix

... 22

('•yUipoinorpha . .

... 21

('.ylíipojxxhmix . . .

.. . 20

Cylnpux

(',\llecorix

... 78

(•yphodenin

('•yphodfxnidcü . .

(•yphopclla

... 81

Cvrtocnpxiix

Cyrtopellix

. . . 59

Cyrlopcllocorix . .

... 79

Cyrtorliinxix

... 78

Cyrtniylxix

... 78

('yxtcorraclia . . . .

Daccrla

Dagbertxix

Datnioxcea

Daxhyrtietiin

... 17

Dnxhyiixexiielln . .

... 18

tiaxycapxxix

. . . 52

Dnsyxcylxtx

. . . 08

Daxymirix

. .. 101

Drcoinia

. . . 57

Demoplexxa

. . . 40

Prrarorapxux ....

... 27

Orracocorix

. . . 28

Drropbthaixxxa ...

. . . 93

Diaphiiidia

. . . 7.3

Dirhroocnrix

, . . 31

Dichxooxcytux . . . ,

, . . 90

líicypbopxix

,.. 58

nicyphxix

. . 59

Ilimorplirnxix ....

. . 07

Dxoderxix

Dinrnrxx

Diogiirtux

Dioiiroiiolxix

. . 89

Diploliicliirlltx . . , .

.. 101

Diplozdxin

. . 20

Dirlitipalin

. . 91

IHxInxilxrlIa

Dolbhotiiirix

.. 101

I><)lirh<ixlrtii(x

.. 79

Dorlxix

. . 28

llriillixiKinix

Drynphilonirix ...

. . 70

Eblix

l-.bxítxxxx

En riloinrxxix

Errriobia

.. 87

Ertagrln

Etinirllxts

. . 57

Ectmetopterus (>5

Ectopiocerus 85

Ellcnia 78

Etthemus 95

EviboUocoris ,8(i

Eofurius 37

Eolyjrus 100

Ejihippiororis 57

Eliiinerelhis 85

Ereniobictlus 87

Enies/iniis

Ethclaslia 19

Eubalas 100

EnrcrcUa 01

E.urrrocoris 39

Exirharicoris 54

Etichilocoris 91

Euchilofulvius 19

Eucomjisrla 79

E.wlcron 50

Eubynlopcpliis 107

Eujmcbyprllis .39

Enphylocons 83

Europiella 15

Eurotíis 8U

Eurybrochis 28

EitrychilFlIn 38

Eutychitoplcirlla ... 27

ExirycipUia 31

Exixycolpxxs 50

ExtrycxaxxcUa 18

Extiyloxntxia 108

Euxytxiixis 103

Exixyupirorxs 00

Exixjscytophíxx/i 108

Exiryxtyloixiorpha . . . 93

E.xxrystylopiis 91

E.unslylxxs 81

Exislxrtxis 20

EMtrrrIxi.x 15

Exrnilrinxris -15

Exrrxitxií X 1 .X 70

Exnlyi^xis |00

Ealrtxxxia 09

Etilnxxiiodr.x 09

l alixx XIX 22’

Erlixitrtxilrs 7I

Irlixixixix 31

Irxiiiixhxrllti 25

liiixixix 77

Eirbrxixrapxux 75

ExrbxxKXflIíX 110

Eixif^xilxis 27

lloxxix 25

Itxxxtxkxxcnxii 80

Eoixxxicopsrllíi 05

Etihndiii.x 18

ExxMxis 19

Exisnis 25

Caxxorapsus 91

('•<11 frmxiis 85

('•iatiella 92

('•bipliyrocorxx 04

(•lobiceps 70

('•lossopclli.x 02

Conia 94

('•ly pocoriò 89

(iimiiabuxca 24

('•xumxxxin |09

('•xiiaiirlln 84

('•xiiixiicxius 107

Ciiixiirdus 107

('•xixxhadya 30

(iiilxida 101

Unarupia 110

I Inax iipiobx 109

llabrocoii.x 00

I ladiodriiixii 89

IladrotiriiKx 70

! hidxoxxenxxdea 70

Uadropíxyrs 52

llixniuxlocapxus .... 58

IhiUodapoidcx 81

llallodapxts 01

lldltxcidFn 08

ll(xllirotoiiin 31

llallxriis 05

/.'ainblcloiiiold 43

I larprdoixn 37

llniporrxn 51

llridrmíiiiiiirllri .... 02

llritixin .35

lleloprttis .38

llrniirrrociixix 24

llniiiopblhaltixocorxx 18

/Iriixxxpbiirriinixxs .. .30

llfiiiisplKintidrIln .. 29

I Irtiinuririiiix 82

I Irxilíiiiixix 109

lírspnobibopx 30

llrxpniipliyliitti .... 21!

Ilrlrrtifiixilylxix .... 71

I Irirroroxix 30

UrIrKXxcylux 108

llrtnolnmix 09

IlitdrbxiiiKilirItri ... 00

Uxxlrxiinxxxdrn ..... 9.3

llxxlxiororit 1(8)

Htutxnbxnrr 43

lloiiiiiriicoiii (Wl

(Iakvai.ik) — Kcys Cicncra Miriclac of thc World

149

I lolHoninrliiden . . . ■

llophniachus

Hordas

IfoiTutliin

I toiTnthieHn

Ilyalioílcs

I haliodororis

Ilyalorlitoria

lhalojiejdoides

I lyaíojiejdtis

llyalosomcUn

Ilyoidra

Ilyjioiniiiius

llyjmrrlihiororis

lí\l>srlorriis

Ihiaiis

Irndeiiin

Hoacoru

Iltiororella

Inwgni

hhisiíi

/)í(/()/)r/)/ií,ç

Isíilirl

íschtiosrflitoris

Isojnoba

Itnroris

Jatrhiiius

lohrrlus

loriKiiiilis

Kahttia

Kr.maiiirniia

Kaiiiiluiiiirlia

Kiainlnira

Kititaldirlla

Klojdroris

fyllifrlltioli,

/y iiif^lilororis

Allífr/i/oílál

Koaiioa

/yosntiomiiis

lyiiiiuiiniiri

I\usrhrlia>ia

I aholirlla

I iihol)idra

l.aholi\

I arinororidra

/ arinorijiis

/ aiii/>rrliii\a

/ íim/)Mir(i/nií/rn ....

I attijiroí iiiiiHiii

/ v/m

I 'itiiiooslhriianis .

I niiilitojilioi iis

l ai f^idrii

■1() I.asiolabops . . .

r)5 J .asiominus . . .

!)1 I.usiomiris ....

89 l.nurinia

99 l.fl>idoca{>sus .

25 l.cjndofuhniis .

25 l.cj)idol>snlliis .

72 l.cjndotaenia . .

107 I.cjddoxenetus .

107 /.ejirocafisus . .

59 l.eplojdcnia . .

77 l.eploxniithus .

01 l.ntcodcUus ...

75 l.cucoplioroptcrí

71 l.rucoptnuin .

50 l.iisiniwlus . . .

55 l.hiocerororis .

72 Liocapsidca . . .

72 l.iocajisus

29 l.iocoridca ....

88 Lissocafnus . . .

10(i l.itoxenus ....

107 l.opidca

99 l.oliiddla

00 l.ol>idoloii ....

71 l.ojius

91 l.iidiniuis ....

71 l.tnidhiadiola

70 f.iindirlla

08 l.iitlifrirlla . ■ ■

108 l.yddororis ...

75 lydr

102 l.\f;;nrosrylus ..

90 l.ynidra

79 l.yjridolon ....

27 l,\j!;ol>sis

.51! /A'í!i<.í

.57 Mahrlin

21 Marrdniiiis ...

77 Marnirf^oiiii.s . .

100 Marioloiiidin .

.50 Mairidoiiiiis ...

105 Miirudojdiidra

00 Manotoldius ..

71 Manoprlilus ..

00 Manotytoidrs .

81 Marrolyliis ...

01 Madaiiiisraiidla

85 Mala

82 Malaroroiis ...

28 Malaioirs ....

25 Maluat lirororis

1 1 Mailharosoma

108 .Mansoiiirlla ...

25 Maralaudti ....

•14

104

110

111

104

102

100

101

108

102

100

Maurocoris

Maurodactylus

Mcdsloscelis

Mrcolacinus

Mecomma

Megaroelopsis

Megacoeltini

Megaluceraea

Megulocolcns

Mcgalodartylus . ■ . .

Megalopsallus

Megamiris

Melanoslictus

Melanoiriclius

Mi-nnitolf(rnis

Merlila

Mcsolropis ■

Mrlafurius

Microbryororis

Microjnhnus

Mirrnpliylrllus

Millniiiiiris

Miinorapsus

Mimoccps

Mimocoris

Miinniicopdlus

Minytus

Miridius

Miris

yfoissouia

Monatocoris

Mouaktrorisca

Moiialornropsis

Moiialotiioti

Moiiosyiiaiiiina

Moroai

102

105

102

107

100

150 lioi.. Mus.

CioKI.DI — ToilU) XI

(11) -

- Divciiiliro 1055

Neolygtts

. 101

PíuhytomvUa

. . . 07

Neoticellit

. .IO

Parhvxybhus ....

. . . 50

Neosilin

]‘idlíi(di<)ii.i

. . . 80

Pbdycratius

. 71

Meostenolits

. 101

Pudicridca

. . . 80

Platylyy^Hs

. 07

Nvsiditirchcslfs ....

. (>:>

I'ainilin

. . . 08

1'lntyugdiiiiris

■12

Nrsiomiiis

. 101

1’iinf^idiiu

. . . 01

PlatyjirltdCdiis ....

. 11

Nesüdajfhue

Pídililidiiid) Idin . .

. . -10

Plidylidi lis

. 00

Nesosyljihas

. lOU

Paidilhts

. . 80

Xnirocoljm.s

' H.l

I’aj>l>iis

. . Oi)

Platytylus

. 100

\iaiilutiia

lo;!

1‘ntabryocoropiis .

.. 12

PlrsidCdjisus

. 01

íWirholia

•li)

Paracarutts

. . 27

Plcsiocoris

. 08

Nicliomurhiís

Pnrachíimus

. . 11

r,.t

Nirnstratns

(iU

1’ariichiits

. . 71

Plfsiolyptis

, 00

Xoclitorori.i

l’/iraiyliisi(s

, . . 21

Pleurorliildjihoi ns , .

. 100

Sothololms

H.''.

Pmnfuhdus

. . 10

Ptcuroxdudlus

Xolidius

Pnraltiiius

Pliiiielld

Notniohus

.11

Paramixia

. . 52

Xidoslirn

101

P{ir{ij}fintilius . . . .

. . 87

117

Xiil()slir<)l>s

101

Parnpruba

.. 71

Pdccilutwltis

102

Xulolremalcs

P(i>iir<minus

. . 70

Pdlymcnis

Xycliden

.'il

Pdrnrnilniius

. . 10

Pdjilnusiii

Xytiin/itnis

lor.

Piira.sriddeiiut

. . ,77

Pdijioiiiiris

107

Orhterocnl>sus

lOi»

Piirastiiniuridca ■ .

. . 77

Pii.sldtinira

Oddiiirlln

■11

Paraxriictus

.. 110

PirjfojKs

100

Odoiitíxrrorori.i . . . .

Pnrcdrocoris

. . 5,7

Pidiimbliíi

Odoitloldalys

Pnriltniints

(}frl{jt%

'.li

Olii^nliirlla

.58

Píiidiamuia

. . 20

Olytnjdorniisu.i

lOU

Pítundygiis

. . 07

Ptdtidldcrrpis

■11

Oinniiilodniia

lOi»

Prilididyxus

. . 05

IO'

Onijdinloiiohis

0.7

/V//iV/f;/;r/)lií! ....

. . 88

Psfdidiiiiiiitis

.78

OncriDinrloliti.s

1 0!)

Prtissdba.iis

. . 18

Piídlojii

OiK oiiDlrllus

.78

Prrilrdjds

. . 18

Psattdlms

Oiirolyliií

Pnilropdidr.s

. . 10

Psattus

■10

OlIDIllllUS

107

I'i'runndtdtis

P^nitlfuiyslyliis . . . .

(>/dil/inhiioiiiiiií ....

10.7

l’liiinUits

. . 00

PiciidobrydCdiis . . . .

()/d.^l/i(diiniiti

.71

Phildslrphiiints ...

. . 00

I‘5riidd(itntjddbr()< his

O/dsl/inryrhií

7!)

/'/uiriiíforn/utu

. . .78

Psniddrniiiiis

Onniirllii

I*Íi\li(Íru

. . 10

P\niddrln(iilii

Ortiíus

li)

Pliylloiyliilitif ....

. . 21

Psniildddtiirlbi

■12

<)rrrlodrru.i

Phylldjiiilra

PsnididdXdps

Orlhorrpludus

Phylluí

, . 51

P\niddiirdbd>u\ ....

(htlidiiidii^

.71

Pltysdldioidlilntí .

, , 18

P\niitdilhdlylii\ ....

(Irtlidjis

100

Physdlilididplnrlbi

. 10

Psr\tddjm>ililun ....

8.7

Oilhidylrlliií

Phytdídridnt

Psruddjt\idt\i\

Orllidlylidni

.70

Pliylondif

. 81

l'\nido\r)irtiiy

Oillidlytu^

Pinxiií

. i))i

Pydididiiy

()\iitirdri%

Piiyldldinii\

. 28

Pyi iiddnrs

1.7

Vorh\lnft\

tnnmn

(17

I'iii liytnrrorfnif ....

Pitdpbdrdjnis

. 80

iintnhro

107

I’<irh\>irunhimrniif .

PUdjdidun

. Hl

Itnuidxdin

I’iirh\liflli^

Piiiidiliif

. 101

i{rtnuiants

I’iii liylirlldpsii

Pilldlllllf

. 102

Hrxtliriiiii

107

I'iicli\j)ddn

1'.1

Plardt liiliis

. 77

Urxxdn

I'iêchyf)lniiii

102

l’l'iKÍ<>K""lliidni . . .

. 71

Ilfiiiniii

Cakvai.ho — Kcys Gcncia Miriclae ot thc World

151

Hrulirista 90

lieuterosiofms 10

Uhabdoscytus 89

JUiasis 97

Rhiiiacloa 41

Hhinocapsidea 78

J{hi>ior(il>sus 55

Ilhiiiocylajiidius .... 20

Hhiuorylajms 20

lUiiiiofnhdiis 20

Ilhiiioiniridius 19

Hhinomiris 19

Rhinojdirns 21

Rhodolygxts 97

Rliolmlisccscluitiis . . 12

Romua 27

Roudairca 55

Riisjiolielln 102

Snhnclus 100

Snlilhrrgrila 42

Sailfún y.i

Saligiitis 91

'iíiroiia (íh

'iiihitiiioiiiiiíx 29

''iiiiiidersiclln 99

Srroilaiitia 19

Schortiocotis 07

''('hoiitrdniirlln .... 85

''I hoiitrxlrtiomitis . . 105

'irhrocdrrirlln 10

'xrirtrlrlliis 00

''rjiiniix fi7

''niiiiim 70

VfnVo/i/Kiiirj H2

'•rttxiiiis ")H

Sitliiia 90

'•innfiis .SI

''ilmiin 29

'xÍMonoiux S5

''ohniiix 02

''"laiionii ix 09

ux ■17

■'•1'iiiliu ii \ SO

Sphinclothorax

Sletiocapxux . .

Stciiodftiia . . .

Stenoparcdra

Stctiojmria . . .

Stciioptcrocoris

Sletioliix

SIflhocoiiiix . .

Slhcuaridea . .

Sthexiaropsis .

Stiftolophux . .

Stiltonipxux . .

SIroitgytocorix

Slrophopoda .

SiiUunitn

Syiigoiiits ....

Xyxitinx

Syxicllotiolidca

SysIíIRmolopsix

S\slcll(»iohtx .

Tardia

Tarniophornx

'Tapuruyjitiux

'Taylorilygux .

Tricot Itititix .

Triillirroiix . .

Trraiorapxttx

Tnaiororis . . .

Trralofuhàux

Tcrmalopityiriia

Trniiatopliyiidra

Trrtiialopliyioidrx

Trriiiatopliyiuin

Tliatiia

Thaamaxtotaitix

Thrnaororis

TItrriaus ....

l iiirlortirix . .

'l'iiigiiiolopsix

'1'illgillOlltllt

Tiiiit rpliaiitx .

’l'oioiigia ....

Toiiiia

Trarlirionoltix

Tiat liriíicliitx

110

Trairiscororis

Trevrsxa

10.4

Tricliobasix

108

Triciiocapxus

Tricliofuiviux

Tricliopborcila

Tricbophoroiicux . . .

Trichophthaimocapsus

Trigotiotyiux

105

'15

Triiarcux

Trojtidaphoreiia . . .

102

Tropidoxieptrx

Trygo

4í)

Tupotiia

Tuxcnriia

Txlocapstts

Tyiopcitix

i^ricana

Vtopnia

()5

Vanniopxix

Oi)

i’aniiiax

8,3

Tiiiirrxicorix

Voikrliopxix

Toikriiax

101

Toiuiiiiiiis

Tonirbia

l'oiiicliirila

Xriirloiiiorpba

110

105

Sriirlii.x

111

Xriioitorttx

Xrtiocyiapux

yaiigaiiibia

Yriioitia

Yiigorrila

/.ar\iillitix

109

/.aiiiiutitia

'/.atirhixmr

/.aiirliinx

/.aiicxxa

105

'/.aratux

/.ikaiiioia

/.oiiiix

/.onodotcllus

■19

/.ottodoropsix

(ir»

/oxippiis

) 1 1

/xgimiis

9(i

SciELO

10 11 12 13 14 15