ISSCA

International Society for the Study

and Conservation of Amphibians

(International Society of Batrachology)

SEAT

Laboratoire des Reptiles et Amphibiens, Muséum national d'Histoire naturelle, 25 rue Cuvier, 75005

Paris, France. — Tel.: (33).(0)1.40.79.34.87. — Fax: (33).(0)1.40.79.34.88. — E-mail: ohler@mnhn.fr.

BOARD

President: C. Kenneth Don», Jr. (Gainesville, USA).

General Secretary: Annemarie OHLER (Paris, France).

Deputy Secretary: Alain PAGANO (Angers, France).

Treasurer: Stéphane GROSIEAN (Paris, France).

Deputy Treasurer: Rafael De SA (Richmond, USA).

Councillors: Lauren E. BROWN (Normal, USA); Alain Dusois (Paris, France): JIANG Jianping (Chengdu,

China); Esteban O. LavILLA (San Miguel de Tucumän, Argentina); Thierry LODÉ (Angers, France):

Miguel VENCES (Amsterdam, The Netherlands).

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Bibliothèque Centrale Muséum

ALvTEeS IN

INTERNATIONAL JOURNAL OF BATRACHOLOGY

July 2005 Volume 23, N° 1-2

Alytes, 2005, 23 (1-2): 1-24.

Amphibia Mundi. 1.1. *

An ergotaxonomy of recent amphibians

Alain DuBoIs

Vertébrés: Reptiles et Amphibiens,

USM 0602 Taxonomie and Collections

Département de Systématique and Evolution,

Muséum national d'Histoire naturelle,

25 rue Cuvier, 75005 Paris, France

<adubois@mnhn.fr>

The new publication series Amphibia Mundi was presented by Dugors (2004a). This will be a series

of taxonomic catalogues and regular lists of taxonomic novelties concerning the AmPHiBlA, that will

allow users of taxonomic data (biologists, conservationists, administrators, etc.) to find updated

information on the state of the art. Contributors to this ambitious endeavour are welcome and should

contact our editorial board, either to write some contributions, or to provide information, or to correct

some of the mistakes or omissions that our catalogues will unavoidably contain. This first issue of the

series presents a list of taxonomic novelties in recent amphibians, i.e., basically a list of new nomina

Œugois, 2000) recently proposed for amphibians. Information on these novelties must be provided

within the frame of a given taxonomy, and respecting strictly the Rules of the /nternational Code of

Zoological Nomenclature (“the Code”; ANONYMOUS, 1999). These rules, often designated as “Linnaean”,

have force of law for all zoologists worldwide except those who expressly state that they are following

other rules, but then the nomenclature adopted is incompatible with a “Linnaean”? one (for details, see

Dusois, 2005a). The taxonomy used as a framework for Amphibia Mundi deserves a few comments.

In most zoological groups, and especially in those like the amphibians, which are currently the

matter of numerous phylogenetic works often followed by drastic reappraisals of relationships, any

given taxonomy is bound to be provisional. This is by no means problematic, as long as one

understands the “heuristic value” of taxonomy (e.g. MAyR, 1981). Taxonomies are not only “results”?

of phylogenetic and taxonomic research, but may serve as starting points for further research, as they

provide hypotheses on relationships that can be tested. For each z0ological group, until we have

reached its “final taxonomy”, a goal that is legitimate but which will probably remain out of reach for

many decades yet, any taxonomic frame must be viewed as a “working taxonomy” (Duois, 1999) or

more shortly an ergoraxonomy (DuBoIs, 2005a).

Taxonomy under the “Linnaean” system consists in wo different aspects (e.g., DUBOIS, 2005a):

establishment or use of raxa, and allocation of ranks to these taxa. These two aspects are independent

and widely different. Establishment or use of taxa is a scientific work that relies on a philosophy of

taxonomy: it requires a decision regarding which information is believed to be important or crucial to

be carried by taxa and their nomina. Many authors consider that taxonomy should be “phylogenetic”,

ie., that taxa should, as far as possible (but see DELORME et al., 2004), be “monophyletic sensu

Source : MNHN, Paris

2 ALYTES 23 (1-2)

Hennig” or holophyletic. Taxa are hierarchically nested within one another, some being more inclusive

than others, and there is a single hierarchy of all living beings. À distinct matter is the ranks that are

given to these more or less inclusive taxa. Despite several atrempts in this respect, there is at present no

homogenizing principle that would allow equivalence of taxa at a given rank in different groups: a

family ofbirds is by no criterion equivalent to a family of frogs (Dunots, 1988). Ranks are arbitrary and

subjective, as are the nomina of taxa. However, just like for the latter, this does not mean that they are

useless or harmfül and that they should be abandoned. Ranks provide a useful, if not indispensable,

system of hierarchisation and indexation of taxonomic information (for more details, discussion and

references, see Duois, 2005a). À careful use of ranks allows them to play an important rôle in the

robustness of ergotaxonomies. Such robustness is an appropriate goal for ergotaxonomies, as they are

meant to be useful not only to phylogeneticists and taxonomists but also to all other users of zoological

nomina.

To avoid unnecessarily frequent changes in ergotaxonomies, especially back and forth movements

between two related taxonomic schemes, any ergotaxonomy chosen for a group should be largely

conservative. To attain this goal it should preferably afford primary key ranks (e.g., ordo or familia;

Dusois, 2005b), to taxa that are widely recognized as valid, i.e., that are considered by most authors, on

the basis of apparently reliable data, as corresponding to well-supported clades. Such taxa and their

nomina are likely to remain unchanged for long periods, which will be appreciated by non-taxonomist

users. In contrast, taxa that are more controversial, being less robustly supported by the current set of

data, should be afforded secondary key ranks (e.g., legio or phalanx) or even subsidiary ranks (e.g.

superfamilia or subfamilia) (for more details, see DuBois, 20054). This philosophy was followed for the

choice of the ergotaxonomy used in this issue of Amphibia Mundi.

In the ergotaxonomy presented below, subfamilies and tribes are recognized only when supported

by published phylogenetic hypotheses, even when provisional: some of these taxa are likely to change,

but this will not affect very much the overall familial scheme. At higher levels, although hypotheses

about the relationships between the provisional families as recognized here do exist (for recent data, see

e.g.: Haas, 2003; HorGG et al., 2004; RoELANTS & Bossuyr, 2005; San MAURO et al., 2005), they are not

yet consensual and are still likely to be modified in the coming years. Until a robust cladistic hypothesis

is widely accepted, it seems better to refrain from recognizing taxa of rank suborder between family and

order, especially as this would raise various nomenclatural problems concerning their best designation

(for more details, see Dupois, 20046, 2006). At any rate, in the future, the two nomina ARCHAFOBA-

rracia Reig, 1958 and Nrosarracura Reig, 1958 must be definitively abandoned, for two distinct but

complementary reasons: (1) these nomina are junior homonyms of the nomina ARCHAFOBATRACHI

Sarasin & Sarasin, 1890 and Neogarracui Sarasin & Sarasin, 1890; (2) the nomen ARCHAEOBATRACHIA

Reig, 1958 was proposed for a taxon that is clearly paraphyletic (references above). The nomen

Nrorarracut Sarasin & Sarasin, 1890 is the valid nomen of the subclass of recent amphibians, that has

sometimes been called LissamPmBia Haeckel, 1866. The latter nomen must also be abandoned, being

an invalid junior synonym of Baracuia Brongniart, 1800. More details on nomenclature of higher taxa

(above superfamily) of Amrmia were provided by Durois (20045).

Rather than recognizing suborders, a better solution for the time being is to recognize higher ranks

in the family-series, i.e., superfamilies (ending in -omr4) and epifamilies (ending in om), as

redefined by Dumois (20056): these taxa do not require the use of other nomina than those of families

and may be easily abandoned or modified whenever changes are brought to the clado-taxonomic

scheme. The cladistic scheme of San Mauro et al. (2005), which largely agrees with other recent studies

(Haas, 2003; RoELANTS & Bossuvr, 2005) was used as the basic framework for recognition of these

higher family-series taxa. For fossil groups, SaNCHiz (1998) and S. E. Evans et al. (2005) were largely

followed. As explained by DuBois (2004b, 20058), any higher taxon that only includes one taxon of next

lower rank, a situation that is sometimes made necessary for taxonomic balance and homogeneï

bears the same nomen as this lower taxon: e.g., epifamily PaLonaromDiA and superfamily PELOBATOIDEA,

Source : MNHN, Paris

Dugois £)

or superorder + ALLOCAUDATA and order + ALLOCAUDATA. In such cases, the two ranks are redundant,

which causes no nomenclatural problem as long as this does not require formation or recognition

of a distinct nomen for each rank (for more details on this controversial question, see DuBOIs,

20056).

The general ergotaxonomic frame used here is conservative at the family level: most of the families

have been used for more than a century, and correspond to clades that are consensually recognized by

most current authors. In most groups, the ergotaxonomy retained is similar to that given in HUTCHINS

et al. (2003), but there are a few important differences, largely related to the inclusion of extinct taxa,

but also in a few cases to the acknowlegement of recent cladistic data. Unlike the recent lists of FROST

(1985), DurLLMAN (1993) and GLAw et al. (1998), Amphibia Mundi provides information on fossil taxa.

This has some bearing on the taxonomic frame retained. Because many fossil taxa are known only from

partial skeletons, their inclusion in the taxonomy of recent amphibians results in some uncertainties. To

limit the impact of this problem, a conservative familial ergotaxonomy was adopted here whenever

phylogenetic relationships between taxa are still unsolved or controversial. These taxa were maintained

in comprehensive families, which may be split later whenever relationships berween them are better

understood. Not doing so would result in having many genera unallocated to families, which would

simply have to be listed as “incertae sedis” at family level. Given these premises, the taxonomic-

nomenclatural scheme adopted here for some families needs a short discussion:

(1) For the “discoglossoid” frogs, the taxonomic scheme of SancHiz (1998: 18) was here followed,

with a single family including the ALvriAr, BomminaroriNAr, DIscoGLossiNAE and + GOBIATINAE, as the

relationships between these four groups are still controversial (Haas, 2003; HoEGG et al., 2004;

RoELanrs & Bossuyr, 2005; San MAURO et al., 2005). Additionally, a few problematic fossil genera are

simply referred to the family without subfamilial allocation. The family as here recognized will most

probably have to be dismantled when the relationships between all its genera are better understood. In

the meanwhile, the valid nomen for this family is BomBiNarorIDAr, as pointed out long ago (DUBOIS,

1984). As the International Commission of Zoological Nomenclature decided not to use its plenary

powers to protect the nomen DiscocLossipar (see DuBois, 19874), the Code’s Rule of Priority must be

followed. The nomen Bomsaroribar has been used repeatedly in the recent years (e.g.: FORD &

CANNATELLA, 1993; Bu & BossuyT, 2003; MAGLIA, 2003; CANNATELLA & HILLIS, 2004; ROELANTS &

Bossuyr, 2005), so it cannot be rejected as a nomen oblitum.

€) The case of the “pelobatoid” frogs is similar. Recent discussions have not yet led to a

consensual hypothesis for relationships among groups (GarCia-PaRis et al., 2003; HAAS, 2003; CaNNa-

rer & Hizuis, 2004; HorGG et al., 2004; ROELANTS & Bossuvr, 2005; San MAuRO et al., 2005). À

provisional conservative scheme with a single family Pecorariar was adopted here. This family

includes four subfamilies (Macormevinar, PeLonaTINAE, PELODYTINAE, ScapatoPoDINAE) and several fossil

genera that cannot be allocated to subfamilies in the present state of knowledge, especially because of

apparent convergences between PrLonariNar and Scarsroropinar in their fossorial adaptations.

(3) Two subfamilies are recognized here within the family Praz following B. J. Evans et al. (2004,

200$). Priority requires that the subfamily including Xenopus and Silurana be called DACTYLETHRINAE, as

already pointed out repeatedly (DuBois, 1983, 1984, 1985, 1987b-c).

(4) The epifamily Ranorbta as recognized here corresponds to the “NroBaracuta” of several

recent authors. This clade is robustly supported by most recent analyses (e.g., HOEGG et al., 2004; Van

DER MEIDEN et al., 2005). It includes two well-supported large clades, recognized here as the

superfamilies HyLomdrA and Raxom£A, and two smaller groups of uncertain afinities (HOEGG et al.,

2004), recognized here as the superfamilies HeLrorHRYNOIDEA and S006LOSSOIDEA.

(5) No subfamilies are currently recognized in the family Buronipar, although this huge assem-

blage clearly consists of several subclades, some of which have a limited geographical range whereas

others have a much larger distribution. In case future works support the formal recognition of

Source : MNHN, Paris

4 ALYTES 23 (1-2)

subfamilies, several family-series nomina are already available and should be used to nominate them

rather than coining new nomina (DuBois, 1984: 34-35, 1987a: 24-29).

(6) The results of DARST & CANNATELLA (2004) and HoEGG et al. (2004) suggest that the subfamily

PseuniNar Fitzinger, 1843 should be considered a synonym of Hyz1var Rafinesque, 1815, not a distinct

subfamily.

(7) The recent finding (DARST & CANNATELLA, 2004) that the genus Brachycephalus Fitzinger, 1826

(including Psyllophryne Izecksohn, 1971, according to KAPLAN, 2002) is phylogenetically nested within

eleutherodactylines, as usually understood, suggests that the taxon including these genera should be

called BracuvorrxauiNar Günther, 1858 instead of Ezsurmeronacrvuinar Lutz, 1954 (as used e.g. by

DuELLMAN, 2003). The genus Craugastor Cope, 1862 was recently recognized for a large part of the

species usually placed in Eleutherodactylus Duméril & Bibron, 1841 (CrawrorD & SmrrH, 2005), and

the latter genus might have to be further split.

(8) The family RAMIDAE as understood here is a very conservative group which corresponds to the

epifamily RANOIDAE as recognized by VENCES & GLAW (2001) and VAN DER MEIJDEN et al. (2005). This

huge assemblage includes a number of taxa whose relationships are not yet clarified and most of which

are here provisionally treated as subfamilies, following Dusors (2003) but adding the MANTELLINAE and

RHAcOPHORINAE. Treating the latter as families makes the RANIDAE paraphyletic (VENCES & GLaAw, 2001;

Van Der MEIDEN et al., 2004, 2005). This family will probably have to be split in several families, but

these may correspond only in part to the subfamilies as recognized below, so this move appears

premature. Changes are here brought to the following taxa:

(a) Van per MEUDEN et al. (2005) recently pointed to the well-supported existence of a

previously undetected radiation in African ranid frogs that includes all genera placed by Durots

(2003) in the Cacosrenivar Noble, 1931 but also the genera Afrana, Natalobatrachus and Pyxicepha-

lus. This finding is ackowledged here in placing all these genera, as well as the clearly related Amietia

and Aubria, in the same subfamily, for which the nomen PyxIc£PHALINAE Bonaparte, 1850 has pri-

ority.

() The data of Van DER METDEN et al. (2005) also suggest that the genus Occidozyga is a member

of the DicroGLossiNAE (as already proposed by DuBois, 1987a, 1992), and therefore the subfamily

OccipozyGiNar in Duois (2003) is here downgraded to the rank of a tribe of the later.

genera are not members of the DickOGLOSSINAE and that the tribe CerarosarracHinI of DuBois (2003)

should be provisionally treated as a subfamily of its own. The valid nomen for this subfamily is

Crsarosarracminar Boulenger, 1884, not PLarymanrivar Laurent, 1986, as suggested by VAN DER

MEMDEN et al. (2005). The genus Batrachylodes, placed by DuBois (1987a, 1992, 2003) in the RAnIN4E

without robust evidence, is here tentatively referred 10 this subfamily mostly on the grounds of

reproductive mode (direct development) and biogeography.

() In the tribe Lmnonecrint of the DicroGLossinar, the genus Liurana Dubois, 1987 is here

considered a strict synonym of Taylorana Dubois, 1987 (Dusois & OHLER, in preparation). Priority of

Taylorana over Liurana was fixed by the first-reviser action of DuBois (1999: 91).

(e) Cladistic relationships within the subfamily Ranrxar as recognized by Dumois (2003) remain

very poorly known and will require additional data. This will not be not an easy task because, as already

pointed out (Dugois, 1981, 1987a, 1992, 2003), such a revision to be meaningful cannot be limited 10

analysis of a subsample of the subfamily, chosen e.g. on geographical grounds (e.g., HiLLIS & WILCOX,

200$), but must include representative of at least all groups and subgroups defined by DuBois (1992),

and probably more. The tribe Amozomni Yang, 1991, recognized by Dumois (2003), is not adopted

here, as its relationships and contents are not yet fully understood. The genus Odorrana Fei, Ye &

Huang, 1991 should probably be separated from Rana (Dugots, 2003), but its relationships with several

Source : MNHN, Paris

Dugois 5

other groups (e.g.: Bamburana Fei, Ye & Huang, 2005; Chalcorana Dubois, 1992; Eburana Dubois, 1992;

Nasirana Dubois, 1992) should first be clarified. Except for the genera Afrana, Amieria and Strongylo-

pus, here placed in the Pyxicrræaunar for reasons explained above, the genus Rana is here kept as a

wide and probably polyphyletic assemblage (Durois, 1992) to avoid the creation of paraphyletic

genera, which would certainly be the case if e.g. Amnirana Dubois, 1992, Hylarana Tschudi, 1838,

Pelophylax Fitzinger, 1843, Pseudorana Fei, Ye & Huang, 1991 or Rugosa Fei, Ye & Huang, 1991 were

raised 10 generic rank, as proposed by some recent authors (e.g., FE et al., 1991, 2005; VAN DER

MEUDEN et al., 2005).

( On the other hand, within the RAMINAF, a new tribe Srauromnt (type-genus Staurois Cope, 1865)

is here erected for the genus Sraurois alone. The diagnosis of this tribe is as follows: its members share

two important characters with the RHAcoPHoRINAr, ie. completely closed ventral cells on digital disks

and glandular belly skin (Dugois, 1992: 321, 334; Bossuyr & Dugois, 2001: 4), and show other

synapomorphies for the RAMIN4E, such as special courtship display (HARDING, 1982) and unusual

keratodont formulae in tadpoles (ALTIG & MCDiarMID, 1999: 331). BossuyT & DuBois (2001: 4) wrote

about the genus Staurois: “In fact, but for the absence of intercalary cartilages on digits, there seems to

be little reason for not assigning the genus Staurois to the Rhacophorinae”. According to recent

molecular data (ROELANTS et al., 2004), this genus appears as the sister-group of all other RANINAE.

(@) The relationships of the other groups of RANIDAE (as here understood) are not yet clarified (e.g.,

Van DER MEIDEN et al., 2005), so these groups are here provisionally maintained as the subfamilies

CONRAUINAE, LANKANECTINAE, MICRIXALINAE, NYCTIBATRACHINAE, PTYCHADENINAE, RANIXALINAE, PETROPEDE-

TiNAE and ParyNosarracHinar, The latter two groups were included by Dupois (2003) in a single

subfamily PerrorsperiNar, but this taxon appears paraphyletic according to the data of VAN DER

MEIJDEN et al. (200$).

(8) The recent molecular data of Darsr & CANNATELLA (2004) and Van DER MEIJDEN et al. (2004,

2005) support the opinion of LauRENT (1980, 1986) and Dumois (1981, 1987a, 1992) that the

ARTHROLEPTINAr, AsrYLosrERNINAE and Hyrerozunar belong in the same clade, recognized by VENCES &

GLaw (2001) and Van DER MEUDEN et al. (2005) as the epifamily ArrakoLsprompas. But these data also

suggest that two other groups, traditionally recognized as the Micronvuinar BRevicirrriNa and the

Hemisoripar, are also members of this clade. To account for these findings, these five groups are here

recognized as subfamilies of a single, purely African, family, which must bear the nomen BkevictrrrIDAE.

This is not because “the oldest available genus name in this clade is Breviceps Merrem, 1920°* (DARsr

& CannarELLa, 2004: 468), as priority among family-series nomina is determined by the dates of the

latter nomina, not by those of the nomina of their included general The valid nomen in this case is

Brsvicrrmina Bonaparte, 1850, which has priority over Hemsiae Cope, 1867 and ARTHkOLEPrINA

Mivart, 1869. In Linnaean nomenclature, Brevicrmina, Brevicrninar, Brevicirripar or Brevicterror-

DEA are simply different aponyms of the same nomen (see DuBois, 2000), which have the same author

and date but “simply” different ranks: it is thus incorrect to write that there “seems to be no available

superfamily name” for this taxon (DaRsT & CANNATELLA, 2004: 468). Besides the five subfamilies listed

above, a sixth subfamily is here recognized in this family for the Lerrorsunar, which according to

EMERSON et al. (2000) represent a subclade distinct from the HyPEROLUNAE.

() In the urodelan family PzerHoponrIDAr, the traditional taxonomy (WaKkE, 2003) has been

challenged by recent findings. The molecular phylogenetic data recently provided by CHIPPINDALE et

al. (2004) suggest the existence of two major lineages, for which the nomina HemDAcrYLuNAE and

PLerHoDoNTINAE are available. The first lineage seems to include three subclades, which can be

provisionally recognized as tribes, under the nomina BozrroGLossni, Hemibacryuunr and SrezerPint. In

this group, the genus Eurycea Rafinesque, 1822 is here understood as including the taxa traditionally

known as the genera Haideorrion Carr, 1939, Tiphlomolge Stejneger, 1896 and Tphlorriton Stejneger,

1893, as well as other taxa more recently recognized (HiLLis et al., 2001). The second lineage of

Source : MNHN, Paris

6 ALYTES 23 (1-2)

PLETHODONTIDAE seems to include two subclades, for which the nomina DESMOGNATHINI and PLETHO-

Doxrir are available. The new data obtained by MUELLER et al. (2004), and by MIN et al. (2005) on the

occasion of the discovery of Karsenia koreana, furthermore suggest that the genus Hydromantes s.].

(including Speleomantes) must be placed in the DesmoGnaTHINI, rather than in the BozrrocLossinr. The

phylogeny and taxonomy of this family are still under intense study and will probably have to be

modified in the near future.

(0) In the family SALAMANDRIDAE, on the basis of “molecular studies in progress” (“estudios

moleculares en curso”), Garcia-Paris et al. (2004) recently split the genus Trirurus in four genera,

recognizing the genera Lissorriton, Mesotrion and Ommatorriton. No subfamilies are currently recog-

nized in this family, although LAURENT (1986) had recognized three subfamilies, including respectively

the genera Pleurodeles, Salamandra and Triturus (and other genera in each). If, following ongoing works,

this or a similar arrangement had to be adopted, the valid nomina for these three subfamilies would be,

respectively, PLeuropeuNA8 TscHunt, 1838, SaLamanprinar Goldfuss, 1820 and Mozcivar Gray, 1850

(see Dugoïs, 1985).

(11) Beside the three traditional orders AnurA, URODELA and GymNorxronA (for their valid

nomina see DuBois, 2004), an order + ALLocAUDATA is here tentatively recognized for the family +

ALBERPETONTIDAE in order to account for the results of McGowan & Evans (1995).

The ergotaxonomy used in this first issue of Amphibia Mundi will certainly have to be modified in

subsequent issues. The list below only mentions the nomina of taxa currently considered valid on the

basis of published evidence, except in a few cases mentioned above. Hierarchy of taxa is shown by

indentation from margin, and ranks of class-series and family-series taxa (DUBOIS, 2005a-b) are written

in full. Taxa of same rank subordinate to the same taxon are listed by alphabetical order. Synonyms,

subgenera and other infrageneric supraspecific taxa, species and subspecies are not listed. Nomina of

entirely fossil taxa are preceded by the sign +.

Classis Ammimia De Blainville, 1816

Subclassis NeoraracHn Sarasin & Sarasin, 1890

Superordo + ALLocauDaTA Fox & Naylor, 1982

Ordo + ALLocauDaTA Fox & Naylor, 1982

Epifamilia + ALsansrreronrontA Estes & Hofistetter, 1976

Superfamilia + AzsanersronronnsA Estes & Hoffstetter, 1976

Familia + Azsansreeronripar Estes & Hoffstetter, 1976

+ Albanerpeton Estes & Hoffstetter, 1976

+ Anoualerperon Gardner, Evans & Sigogneau-Russell, 2003

+ Celtedens MeGowan & Evans, 1995

+ Nukusurus Nessov, 1981

Superordo Barracmia Brongniart, 1800

Ordo Axura Duméril, 1806

Incertae sedis

+ Aralobatrachus Nessov, 1981

+ Avitabatrachus Bâez, Trueb & Calvo, 2000

+ Batrachulina Kuhn, 1962

+ Comobatrachus Hecht & Estes, 1960

+ Czatkobarrachus Evans & Borsuk-Bialynicka, 1998

+ Bobatrachus Marsh, 1887

+ Eorubera Hecht, 1960

+ Estesina Roëek & Nessov, 1993

+ Gobiatoides Roëek & Nessov, 1993

Source : MNHN, Paris

Dugois 7

+ Hatzegobatrachus Venczel & Csiki, 2003

+ Jtemirella Nessov, 1981

+ Liventsovkia Ratnikov, 1993

+ Luteriobatrachus Wattke, 1988

+ Mesophryne Gao & Wang, 2001

+ Negatchevkia Ratnikov, 1993

+ Nezpercius Blob, Carrano, Rogers, Forster & Espinoza, 2001

+ Novooskolia Ratnikov, 1993

+ Probarrachus Peters, 1878

+ Procerobatrachus Roëek & Nessov, 1993

+ Protophrynus Pomel, 1953

+ Saevesoederberghia Roëek & Nessov, 1993

+ Sunnybatrachus Evans & MeGowan, 2002

+ Thaumastosaurus De Stefano, 1903

+ Theatonius Fox, 1976

+ Yichoubatrachus Gao & Chen, 2004

Familia + Prosauibar Shubin & Jenkins, 1995

+ Prosalirus Kuhn, 1964

Familia + Raxavinar Fcjérväry, 1920

+ Ranavus Portis, 1885

Familia + Tkecosarracuiar Holman, 1974

+ Tregobatrachus Holman, 1974

Familia + Vieraezunar Kubn, 1964

+ Vieraella Reig, 1961

Epifamilia Bomsvarorornia Gray, 1825

Superfamilia BompinarororpeA Gray, 1825

Familia BomiNarTorIDAr Gray, 1825

Incertae sedis

+ Alranulia Gubin, 1993

+ Callobatrachus Wang & Gao, 1997

+ Enneabatrachus Evans & Milner, 1993

+ Latoglossus Hossini, 2000

+ Montsechobatrachus Fejérväry, 1921

+ Opisthocoelellus Kuhn, 1941

+ Pelophilus Tschudi, 1838

+ Scotiophryme Estes, 1969

Subfamilia Azyrnar Fitzinger, 1843

Alyres Wagler, 1829

+ Kizylkuma Nessov, 1981

Subfamilia Bomanarorivar Gray, 1825

Barbourula Taylor & Noble, 1924

Bombina Oken, 1816

Subfamilia DiscocLossivae Günther, 1858

Discoglossus Otth, 1837

+ Eodiscoglossus Villalta, 1956

+ Latonia Meyer, 1843

+ Paradiscoglossus Estes & Sanchiz, 1982

+ Paralatonia Venczel & Csiki, 2003

+ Wéaldenbatrachus Fey, 1988

Source : MNHN, Paris

ALYTES 23 (1-2)

Subfamilia + Gomarinar Roëek & Nessov, 1993

+ Crerasalia Gubin, 1999

+ Gobiates Spinar & Tatarinov, 1986

Epifamilia Lerorezmaromnta Mivart, 1869

Superfamilia LerorszmaromrA Mivart, 1869

Familia Ascarminar Fejérväry, 1923

Ascaphus Stejneger, 1899

Familia Lerorezmaripas Mivart, 1869

Subfamilia LeorezmariNar Mivart, 1869

Leiopelma Fitznger, 1861

Subfamilia + Norosarracminar Reig, 1957

+ Norobatrachus Reig, 1956

Epifamilia PrLosarorDiA Bonaparte, 1850

Superfamilia PeLoraromnza Bonaparte, 1850

Familia PeLogATIDAE Bonaparte, 1850

Incertae sedis

+ Liaobatrachus Ji Shu’an & Ji Quang, 1998

+ Macropelobates Noble, 1924

+ Uldzinia Gubin, 1996

Subfamilia MeGop#ryINAE Bonaparte, 1850

Tribus Leprosracauni Dubois, 1983

Leptobrachella Smith, 1925

Leprobrachium Tschudi, 1838

Leptolalax Dubois, 1980

Orcolalax Myers & Leviton, 1962

Scutiger Theobald, 1868

Tribus Mecorarvinr Bonaparte, 1850

Brachyrarsophrys Tian & Hu, 1983

Megophrys Kuhl & Van Hasselt, 1822

Ophryophryne Boulenger, 1903

Xenophrys Günther, 1864

Subfamilia PeLosaninar Bonaparte, 1850

+ Eopelobates Parker, 1929

Pelobates Wagler, 1830

Subfamilia PeLonyriNae Bonaparte, 1850

+ Miopelodyres Taylor, 1941

Pelodyres Bonaparte, 1838

+ Rphrodyres Henrici, 1994

Subfamilia Scasioronivar Cope, 1865

Scaphiopus Holbrook, 1836

Spea Cope, 1866

Epifamilia Prroupta Gray, 1825

Superfamilia Prromp£a Gray, 1825

Incertae sedis

+ Aygroua Jones, Evans & Sigogneau-Russell, 2003

À Thoraciliacus Nevo, 1968

Familia + PaL4roBATRAcHIDAE Cope, 1865

+ Albionbatrachus Meszoely, Spinar & Ford, 1984

+ Messelobatrachus Wauttke, 1988

Source : MNHN, Paris

Dugois

+ Palacobatrachus Tschudi, 1838

+ Pliobatrachus Fejérväry, 1917

Familia Prripar Gray, 1825

Incertae sedis

+ Cordicephalus Nevo, 1968

+ Eoxenopoides Haughton, 1931

+ Llankibatrachus Bâez & Pugener, 2003

+ Shomronella Estes, Spinar & Nevo, 1978

+ Thoraciliacus Nevo, 1968

Subfamilia Dacrvzerurinas Hogg, 1838

+ Pachycentrata Bâez & Rage, 2004

+ Saltenia Reig, 1959

+ Shelania Casamiquela, 1960

Silurana Gray, 1864

Xenopus Wagler, 1827

Subfamilia Prnar Gray, 1825

Hymenochirus Boulenger, 1896

Pipa Laurenti, 1768

Pseudhymenochirus Chabanaud, 1920

Familia RuNorrynipar Günther, 1858

+ Chelomophrynus Henrici, 1991

+ Eorhinophrynus Hecht, 1959

+ Rhadinosteus Henrici, 1998

Rhinophrynus Duméril & Bibron, 1841

Epifamilia Raxorpu Rafinesque-Schmaltz, 1814

Superfamilia HeLrorurynomra Noble, 1931

Familia Hezroraynipae Noble, 1931

Heleophryne Sclater, 1899

Superfamilia HyLomrA Rafinesque, 1815

Familia AzLorarvmpar Goin, Goin & Zug, 1978

Allophryne Gaige, 1926

Familia Buronipar Gray, 1825

Adenomus Cope, 1860

Aliphrynoides Dubois, 1987

Andinophryne Hoogmoed, 1985

Ansonia Stoliczka, 1870

Atelophryniscus MeCranie, Wilson & Williams, 1989

Atelopus Duméril & Bibron, 1841

Bufo Laurenti, 1768

Bufoides Pillai & Yazdani, 1973

Capensibufo Grandison, 1980

Churamiti Channing & Stanley, 2002

Crepidophryne Cope, 1889

Dendrophryniscus Jiménez de la Espada, 1871

Didynamipus Andersson, 1903

Frostius Cannatella, 1986

Laurentophryne Tihen, 1960

Leptophryne Fitzinger, 1843

Melanophryniscus Gallardo, 1961

Mertensophryne Tihen, 1960

Source : MNHN, Paris

ALYTES 23 (1-2)

Metaphryniscus Señaris, Ayarzagüena & Gorzula, 1994

Necrophryne Buchholz & Peters, 1875

Nectophrynoides Noble, 1926

Nimbaphrynoides Dubois, 1987

Oreophrynella Boulenger, 1895

Osornophryne Ruiz-Carranza & Hernändez-Camacho, 1976

Parapelophryne Fei, Ye & Jiang, 2003

Pedostibes Günther, 1876

Pelophryne Barbour, 1938

Phrynoidis Fitzinger, 1843

Peudobufo Tschudi, 1838

Rhamphophryne Trueb, 1971

Schismaderma Smith, 1849

Spinophrynoides Dubois, 1987

Stephopaedes Channing, 1978

Truebella Graybeal & Cannatella, 1995

Wérneria Poche, 1903

Wôlterstorffina Mertens, 1939

Familia CexrroLeNIDAr Taylor, 1951

Centrolene Jiménez de la Espada, 1872

Cochranella Taylor, 1951

Hyalinobatrachium Ruiz-Carranza & Lynch, 1991

Familia Dexproraripar Cope, 1865 (1850)

Allobates Zimmermann & Zimmermann, 1988

Aromobates Myers, Paolillo & Daly, 1991

Colostethus Cope, 1866

Cryprophyllobares Lôtters, Jungfer & Widmer, 2000

Dendrobares Wagler, 1830

Epipedobates Myers, 1987

Mannophryne LaMarca, 1992

Nephelobates La Marca, 1994

Phollobates Duméril & Bibron, 1841

Familia Hyipar Rafinesque, 1815

Subfamilia Hewraracrinaz Peters, 1862

Cryptobatrachus Ruthven, 1916

Flectonotus Miranda-Ribeiro, 1920

Gastrotheca Fitzinger, 1843

Hemiphractus Wagler, 1828

Srefania Rivero, 1968

Subfamilia Hyzia Rafinesque, 1815

Acris Duméril & Bibron, 1841

Anotheca Smith, 1939

Aparasphenodon Miranda-Ribeiro, 1920

Aplastodiscus Lutz, 1950

Argenteohyla Trueb, 1970

Corythomantis Boulenger, 1896

Ducllmanohyla Campbell & Smith, 1992

Hyla Laurenti, 1768

Lysapsus Cope, 1862

Nyctimantis Boulenger, 1882

Source : MNHN, Paris

Dusois 11

Osteocephalus Steindachner, 1862

Osteopilus Fitzinger, 1843

Phrynohyas Fitznger, 1843

Phyllodytes Wagler, 1830

Plectrohyla Brocchi, 1877

+ Proacris Holman, 1961

Pseudacris Fitzinger, 1843

Preudis Wagler, 1830

Prrnohyla Boulenger, 1882

Prychohyla Taylor, 1944

Scarthyla Duellman & De Sä, 1988

Scinax Wagler, 1830

Smilisca Cope, 1865

Sphaenorhynchus Tschudi, 1838

Tépuihyla Ayarazagüena, Señaris & Gorzula, 1992

Trachycephalus Tschudi, 1838

Triprion Cope, 1866

Xenohyla Izecksohn, 1998

Subfamilia PeLoprYanivar Günther, 1858

Cyclorana Steindachner, 1867

+ Emabatrachus Hochnull, 2003

Litoria Tschudi, 1838

Nycrimystes Stejneger, 1916

Pelodryas Günther, 1859

Subfamilia PavzLomrusiar Günther, 1858

Agalychnis Cope, 1864

Hylomantis Peters, 1872

Pachymedusa Duellman, 1968

Phasmahyla Cruz, 1991

Phrynomedusa Miranda-Ribeiro, 1923

Phyllomedusa Wagler, 1830

Familia Lerropacrvuipar Werner, 1896 (1838)

Incertae sedis

+ Estesiella Bâez, 1995

Subfamilia Beacavcernazixar Günther, 1858

Adelophryne Hoogmoed & Lescure, 1984

Aropophrynus Lynch & Ruiz-Carranza, 1982

Barycholos Heyer, 1969

Brachycephalus Fitzinger, 1826

Craugastor Cope, 1862

Dischidodactylus Lynch, 1979

Eleutherodactylus Duméril & Bibron, 1841

Euparkerella Griffiths, 1959

Geobatrachus Ruthven, 1915

Holoaden Miranda-Ribeiro, 1920

Ischnocnema Reinhardt & Lütken, 1862

Phrynopus Peters, 1874

Phyllonastes Heyer, 1977

Phyzelaphryne Heyer, 1977

Source : MNHN, Paris

ALYTES 23 (1-2)

Subfamilia Cerarorr viva Tschudi, 1838

+ Baurubatrachus Bâez & Peri, 1990

Ceratophrys Wied-Neuvwied, 1824

Chacophrys Reig & Limeses, 1963

Lepidobatrachus Budget, 1899

+ Wävelia Casamiquela, 1959

Subfamilia CrcLoramrmNar Bonaparte, 1850

Crossodacrylodes Cochran, 1938

Cycloramphus Tschudi, 1838

Paratelmatobius Lutz & Carvalho, 1958

Rupirana Heyer, 1999

Sesthrophrys Lynch, 1971

Thoropa Cope, 1865

Zachaenus Cope, 1866

Subfamilia HyLoninar Günther, 1858

Crossodactylus Duméril & Bibron, 1841

Hylodes Fitzinger, 1826

Megaelosia Miranda-Ribeiro, 1923

Subfamilia Lerropacryuinae Werner, 1896 (1838)

Adenomera Steindachner, 1867

Edalorhina Jimênez de la Espada, 1870

Hydrolaerare Gallardo, 1963

Leptodactylus Fitzinger, 1826

Limnomedusa Fitzinger, 1843

Lithodytes Fitzinger, 1843

Physalaemus Fitzinger, 1826

Pleurodema Tschudi, 1838

Pseudopaludicola Miranda-Ribeiro, 1926

Vanzolinius Heyer, 1974

Subfamilia Opowroraryaar Lynch, 1969

Macrogenioglottus Carvalho, 1946

Odontophrynus Reinhardt & Lütken, 1862

Proceratophrys Miranda-Ribeiro, 1920

Subfamilia Tkzmarogunas Fitzinger, 1843

Alkodes Bell, 1843

Atelognathus Lynch, 1978

Barrachophrynus Peters, 1873

Barrachyla Bell, 1843

Caudiverbera Laurenti, 1768

Eupsophus Fitzinger, 1843

Hylorina Bell, 1843

Insuetophrynus Barrio, 1970

+ Neoprocoela Schacffer, 1949

Somuneuria Lynch, 1978

Télmatobius Wiegmann, 1835

Telmatobufo Schmidt, 1952

Familia Mvosarracminar Schlegel, 1850

Incertae sedis

+ Indobatrachus Noble, 1930

Source : MNHN, Paris

Dugois 13

Subfamilia Lumvonyasrinae Lynch, 1969

Adelorus Ogilby, 1907

Heleioporus Gray, 1841

Kyarranus Moore, 1958

Lechriodus Boulenger, 1882

Limnodynastes Fitzinger, 1843

Mixophyes Günther, 1864

Neobatrachus Peters, 1863

Notaden Günther, 1873

Philoria Spencer, 1901

Subfamilia Mrorsrracænar Schlegel, 1850

Arenophryne Tyler, 1976

Assa Tyler, 1972

Bryobatrachus Rounsevell, Ziegeler, Brown, Davies & Littlejohn, 1994

Crinia Tschudi, 1838

Geocrinia Blake, 1973

Meracrinia Parker, 1940

Mobatrachus Schlegel, 1850

Paracrinia Heyer & Liem, 1976

Pseudophryne Fitznger, 1843

Rheobatrachus Liem, 1973

Spicospina Roberts, Horwitz, Wardell-Johnson, Maxson & Mahony, 1997

Taudactylus Straughan & Lee, 1966

Uperoleia Gray, 1841

Familia RHNopERMATIDAE Bonaparte, 1850

Rhinoderma Duméril & Bibron, 1841

Superfamilia RaomprA Rafinesque-Schmaltz, 1814

Incertae sedis

+ Ranomorphus Ratnikov, 1993

Familia Brericrripar Bonaparte, 1850

Subfamilia Arrarozrrrivar Mivart, 1869

Artholeptis Smith, 1849

Cardioglossa Boulenger, 1900

Subfamilia AsryLosTERNINAE Noble, 1927

Assylosternus Werner, 1898

Leptodactylodon Andersson, 1903

Nyctibates Boulenger, 1904

Scorobleps Boulenger, 1900

Trichobatrachus Boulenger, 1900

Subfamilia Bezricrerninar Bonaparte, 1850

Balebreviceps Largen & Drewes, 1989

Breviceps Merrem, 1820

Calldina Nieden, 1910

Probreviceps Parker, 1931

Spelacophryne AB, 1924

Subfamilia Hewsorinar Cope, 1867

Hemisus Günther, 1859

Subfamilia HyP£roLINAE Laurent, 1943

Tribus HyreroLuNI Laurent, 1943

Acanthixalus Laurent, 1944

Source : MNHN, Paris

ALYTES 23 (1-2)

Afrixalus Laurent, 1944

Alexteroon Perret, 1988

Arlequinus Perret, 1988

Callixalus Laurent, 1950

Chlorolius Perret, 1988

Chrysobatrachus Laurent, 1951

Cryprothylax Laurent & Combaz, 1950

Heterixalus Laurent, 1944

Hyperolius Rapp, 1842

Kassinula Laurent, 1940

Tachyenemis Fitzinger, 1843

Tribus Kassivr Laurent, 1972

Kassina Girard, 1853

Opisthothylax Perret, 1966

Paracassina Peracca, 1907

Phlyctimantis Laurent & Combaz, 1950

Semnodactylus Hoffman, 1939

Subfamilia Leprorsuivae Laurent, 1972

Lepiopelis Günther, 1859/1869

Familia Microuvuipar Günther, 1858 (1843)

Subfamilia Asrrrorarvinar Günther, 1858

Incertae sedis

+ Australobatrachus Tyler, 1976

Tribus Asrerormevinr Günther, 1858

Asterophrys Tschudi, 1838

Hylophorbus Macleay, 1878

Mantophryne Boulenger, 1897

Pherohaspis Zweifel, 1972

Tribus Barvozwvin Burton, 1986

Barygenys Parker, 1936

Tribus Cazzuzorr Dubois, 1988

Callulops Boulenger, 1888

Tribus Xexorvivr Mivart, 1869

Xenobatrachus Peters & Doria, 1878

Xenorhina Peters, 1863

Subfamilia Cazzurz.unar Fei, Ye & Jiang, 2005

Calluella Stoliczka, 1872

Subfamilia Coruvuixar Cope, 1889

Anodonthyla Müller, 1892

Cophyla Boettger, 1880

Madecassophryne Guibé, 1974

Plarypelis Boulenger, 1882

Plethodontohyla Boulenger, 1882

Rhombophryne Boettger, 1880

Stumpffia Boettger, 1881

Subfamilia Dyscormnar Boulenger, 1882

Dyscophus Grandidier, 1872

Subfamilia Gexvorueyxixar Boulenger, 1890

Alberieus Burton & Zweifel, 1995

Source : MNHN, Paris

Dugois 15

Aphantophryne Fry, 1917

Austrochaperina Fry, 1912

Choerophryne Van Kampen, 1915

Cophixalus Boettger, 1892

Copiula Méhelÿ, 1901

Genyophryne Boulenger, 1890

Liophryne Boulenger, 1897

Orcophryne Boettger, 1895

Oxydactyla Van Kampen, 1913

Sphenophryne Peters & Doria, 1878

Subfamilia Horzoruryninaz Noble, 1931

Hoplophryne Barbour & Loveridge, 1928

Parhoplophryne Barbour & Loveridge, 1928

Subfamilia Microavziar Günther, 1858 (1843)

Tribus Gasrroraevnin Fitzinger, 1843

Adelastes Zweifel, 1986

Alrigius Wild, 1995

Arcovomer Carvalho, 1954

Chiasmocleis Méhelÿ, 1904

Ctenophryne Mocquard, 1904

Dasypops Miranda-Ribeiro, 1924

Dermatonotus Méhelÿ, 1904

Elachistocleis Parker, 1927

Gastrophryne Fitznger, 1843

Hamptophryne Carvalho, 1954

Hyophryne Carvalho, 1954

Hypopachus Keferstein, 1867

Myersiella Carvalho, 1954

Nelsonophryne Frost, 1987

Orophryne Boulenger, 1900

Stereocyclops Cope, 1870

Synapturanus Carvalho, 1954

Syncope Walker, 1973

Tribus Micronvyzini Günther, 1858 (1843)

Chaperina Mocquard, 1892

Gastrophrynoides Noble, 1926

Glyphoglossus Günther, 1868

Kalophrymus Tschudi, 1838

Kaloula Gray, 1831

Melanobatrachus Beddome, 1878

Meraphrynella Parker, 1934

Microhyla Tschudi, 1838

Micrylerra Dubois, 1987

Phrynella Boulenger, 1887

Ramanella Rao & Ramanna, 1925

Uperodon Duméril & Bibron, 1841

Subfamilia PHRYNoMERINAE Noble, 1931

Phrynomantis Peters, 1867

Subfamilia Scarmioraev var Laurent, 1946

Paradoxophyla Blommers-Schlôsser & Blanc, 1991

Scaphiophryne Boulenger, 1882

Source : MNHN, Paris

ALYTES 23 (1-2)

Familia Ranipar Rafinesque-Schmaltz, 1814

Subfamilia CerarosarracHinar Boulenger, 1884

Batrachylodes Boulenger, 1887

Ceratobatrachus Boulenger, 1884

Discodeles Boulenger, 1918

Ingerana Dubois, 1987

Palmatorappia Ahl, 1927

Platymantis Günther, 1859

Subfamilia Conrauvar Dubois, 1992

Conraua Nieden, 1908

Subfamilia DicroossiNar Anderson, 1871

Tribus DickoGLossinI Anderson, 1871

Euphlyetis Fitzinger, 1843

Fejervarya Bolkay, 1915

Hoplobatrachus Peters, 1863

Minervarya Dubois, Ohler & Biju, 2001

Nannophrys Günther, 1869

Sphaerotheca Günther, 1859

Tribus Luvonscriv Dubois, 1992

Annandia Dubois, 1992

Elachyglossa Andersson, 1916

Limnonectes Fivinger, 1843

Taylorana Dubois, 1987

Tribus Occrnozvomt Fei, Ye & Huang, 1991

Occidozyga Kuhl &Van Hasselt, 1822

Phrynoglossus Peters, 1867

Tribus Pur Dubois, 1992

Chaparana Bourret, 1939

Nanorana Günther, 1896

Quasipaa Dubois, 1992

Subfamilia Lankaxscrinas Dubois & Ohler, 2001

Lankanectes Dubois & Ohler, 2001

Subfamilia Manrezunar Laurent, 1946

Tribus Boopmnr Vences & Glaw, 2001

Boophis Tschudi, 1838

Tribus LazrosromunI Vences & Glaw, 2001

Aglyprodactylus Boulenger, 1919

Laliostoma Glaw, Vences & Bôhme, 1998

Tribus Manreuunr Laurent, 1946

Mantella Boulenger, 1882

Mantidacrylus Boulenger, 1895

Subfamilia Micrixaivar Dubois, Ohler & Biju, 2001

Micrixalus Boulenger, 1888

Subfamilia NrcrsarracmiNar Blommers-Schlôsser, 1993

Nyctibatrachus Boulenger, 1882

Subfamilia Perrorsperinae Noble, 1931

Arthroleptides Nieden, 1910

Petropedetes Reichenow, 1874

Subfamilia ParvvonarracHinar Laurent, 1941

Dimorphognathus Boulenger, 1906

Source : MNHN, Paris

Dugois L7

Ericabatrachus Largen, 1991

Phrynobatrachus Günther, 1862

Phrynodon Parker, 1935

Subfamilia Prrcxapewinar Dubois, 1987

Hildebrandtia Nieden, 1907

Lanzarana Clarke, 1983

Prychadena Boulenger, 1917

Subfamilia Prxiceraazinae Bonaparte, 1850

Afrana Dubois, 1992

Amietia Dubois, 1987

Anhydrophryne Hewitt, 1919

Aubria Boulenger, 1917

Arihroleptella Hewitt, 1926

Cacosternum Boulenger, 1887

Microbatrachella Hewitt, 1926

Natalobatrachus Hewitt & Methuen, 1913

Nothophryne Poynton, 1963

Poyntonia Channing & Boycott, 1989

Pyxicephalus Tschudi, 1838

Strongylopus Tschudi, 1838

Tomopterna Duméril & Bibron, 1841

Subfamilia Raxnar Rafinesque-Schmaltz, 1814

‘Tribus Ravvr Rafinesque-Schmaltz, 1814

Amolops Cope, 1865

Pseudoamolops Fei, Ye & Jiang, 2000

Rana Linnaeus, 1758

Tribus Srauromr nov.

Staurois Cope, 1865

Subfamilia RanIXALINAE Dubois, 1987

Indirana Laurent, 1986

Subfamilia Raacornorinar Hoffman, 1932 (1858)

Incertae sedis

Dendrobatorana Ahl, 1927

Tribus Busroerunt Channing, 1989

Buergeria Tschudi, 1838

Tribus PrLaurivr Dubois, 1981

Aquixalus Delorme, Dubois, Grosjean & Ohler, 2005

Kurixalus Fei, Ye & Dubois, 1999

Nyctixalus Boulenger, 1882

Philautus Gistel, 1848

Theloderma Tschudi, 1838

Tribus RaacorxoriNI Hoffman, 1932 (1858)

Chirixalus Boulenger, 1893

Chiromantis Peters, 1855

Polypedares Tschudi, 1838

Rhacophorus Kuhl & Van Hasselt, 1822

Superfamilia Sooczossownra Noble, 1931

Familia NasiKABATRACHIDAE Biju & Bossuyt, 2003

Nasikabatrachus Biju & Bossuyt, 2003

Source : MNHN, Paris

18 ALYTES 23 (1-2)

Familia Sooczossiar Noble, 1931

Nesomantis Boulenger, 1909

Sooglossus Boulenger, 1906

Epifamilia + Tkzpoarracaomia Kuhn, 1962

Superfamilia + TraposarracHoreA Kuhn, 1962

Familia + TRzaposarracminar Kuhn, 1962

+ Triadobarrachus Kuhn, 1962

Ordo UropeLa Duméril, 1806

Incertae sedis

+ Apricosiren Evans & MeGowan, 2002

+ Bishara Nessov, 1997

+ Bissekria Nessov, 1981

+ Comonecturoides Hecht & Estes, 1960

+ Galverpeton Estes & Sanchiz, 1982

+ Hylaeobatrachus Dollo, 1884

+ Jeholotriton Wang, 2000

+ Kivatriton Averianov & Voronkevich, 2002

+ Laccotriton Gao et al., 1998

+ Marmorerperon Evans, Milner & Mussett, 1988

+ Ramonellus Nevo & Estes, 1969

+ Sinerpeton Gao & Shubin, 2001

Familia + Barracosaurordipar Auffenberg, 1958

+ Batrachosauroides Taylor & Hesse, 1943

+ Mynbulakia Nessov, 1981

+ Opisthotriton Auffenberg, 1961

+ Palacoproteus Herre, 1935

+ Parrisia Denton & O’Neill, 1998

+ Peratosauroides Naylor, 1981

+ Prodesmodon Estes, 1964

Familia + Prosievipar Estes, 1969

+ Prosiren Goin & Auffenberg, 1958

Familia + ScaraerpzronriDar Auffenberg & Goin, 1959

+ Eoscapherpeton Nessov, 1981

+ Horezmia Nessov, 1981

+ Lisserpeton Estes, 1965

+ Piceverperon Meszoely, 1967

+ Scapherperon Cope, 1877

Epifamilia CryprosrancHomntA Fitzinger, 1826

Superfamilia CryrrosrancnomrA Fitzinger, 1826

Familia Cryrrosrancuiar Fitzinger, 1826

Andrias Tschudi, 1837

+ Aviturus Gubin, 1991

+ Chumerperon Gao & Shubin, 2003

Cryprobranchus Leuckart, 1821

+ Ulanurus Gubin, 1991

Familia Hyoanar Cope, 1859 (1856)

Subfamilia Hyvoguxar Cope, 1859 (1856)

Batrachuperus Boulenger, 1878

Hynobius Tschudi, 1838

+ Liaoxitriton Dong & Wang, 1998

Source : MNHN, Paris

Dugois 19

Liua Zhao & Hu, 1983

Onychodactylus Tschudi, 1838

Pachyhynobius Fei, Qu & Wu, 1983

+ Parahynobius Venczel, 1999

Pseudohynobius Fei & Ye, 1983

Ranodon Kessler, 1866

Salamandrella Dybowski, 1870

Subfamilia Proronvnounar Fei & Ye, 2000

Protohynobius Fei &Ye, 2000

Epifamilia + Karauromia Ivachnenko, 1978

Superfamilia + KarauromeA Ivachnenko, 1978

Familia + KarauRIDAE Ivachnenko, 1978

+ Karaurus Ivachnenko, 1978

+ Kokartus Nessov, 1981

Epifamilia Saamanprouta Goldfuss, 1820

Incertae sedis

+ Jridorriton Evans, Lally, Chure, Elder & Maisano, 2005

+ Valdotriton Evans & Milner, 1996

Superfamilia Amaysromaromnes Gray, 1850

Familia Ausysromaripar Gray, 1850

Ambystoma Tschudi, 1838

+ Amphirriton Rogers, 1976

Familia Dicamrroponrinar Tihen, 1958

+ Ambystomichnus Peabody, 1954

+ Bargmannia Herre, 1955

+ Chrysorriton Estes, 1981

Dicamptodon Strauch, 1870

+ Geyeriella Herre, 1950

+ Wülrerstorffella Herre, 1950

Superfamilia Amemrumorpea Gray, 1825

Familia Aursumnar Gray, 1825

Amphiuma Garden, 1821

+ Paleoamphiuma Rieppel & Grande, 1998

+ Proamphiuma Estes, 1969

Familia PLrraoonripar Gray, 1850

Subfamilia Hewoacrvzunae Hallowell, 1856 (1850)

Tribus BourroaLossir Hallowell, 1856

Batrachoseps Bonaparte, 1841

Bolitoglossa Duméril, Bibron & Duméril, 1854

Bradytriton Wake & Elias, 1983

Chiropterotriton Taylor, 1944

Cryprorriton Garcia-Paris & Wake, 2000

Dendrorriron Wake & Elias, 1983

Ixalotriton Wake & Johnson, 1989

Lineatriton Tanner, 1950

Nororriton Wake & Elias, 1983

Nycranolis Elias & Wake, 1983

Oedipina Keferstein, 1868

Parvimolge Taylor, 1944

Source : MNHN, Paris

ALYTES 23 (1-2)

Pseudoeurycea Taylor, 1944

Thorius Cope, 1869

Tribus Hembacrvzunr Hallowell, 1856 (1850)

Hemidactylium Tschudi, 1838

Tribus Srzzerrint Cope, 1859

Eurycea Rafinesque, 1822

Gyrinophilus Cope, 1869

Pseudotriton Tschudi, 1838

Stereochilus Cope, 1869

Subfamilia PLertoponrinar Gray, 1850

Tribus Desmooxarinr Gray, 1850

Aneides Baird, 1849

Desmognathus Baird, 1850

Ensarina Gray, 1850

Hydromantes Gistel, 1848

Karsenia Min, Yang, Bonett, Vicites, Brandon & Wake, 2005

Leurognathus Moore, 1899

Phaeognathus Highton, 1961

Tribus PLeraoDonrINI Gray, 1850

Plethodon Tschudi, 1838

Superfamilia ProrzomrA Gray, 1825

Familia Proripar Gray, 1825

+ Mioproteus Estes & Darevsky, 1978

Necturus Rafinesque, 1819

+ Orchophyia Meyer, 1845

Proteus Laurenti, 1768

Superfamilia RavacorrrronomneA Tihen, 1958

Familia Ravacorrrroninar Tihen, 1958

Rhyacorriton Dunn, 1920

Superfamilia SazamanDromDzA Goldfuss, 1820

Familia SaLamanpeipar Goldfuss, 1820

+ Archacotriton Meyer, 1860

+ Brachycornus Meyer, 1860

+ Chelotriton Pomel, 1853

Chioglossa Bocage, 1864

Cynops Tschudi, 1838

Echinotriton Nussbaum & Brodie, 1982

Euproctus Gené, 1838

+ Koalliella Herre, 1950

Lissotriton Bell, 1839

Lyciasalamandra Veith & Steinfartz, 2004

+ Megalorriton Zittel, 1888

Mertensiella Wolterstorff, 1925

Mesorriton Bolkay, 1927

Neurergus Cope, 1862

Notophthalmus Rafinesque, 1820

+ Oligosemia Naväs, 1922

Ommatorriton Gray, 1850

Pachytriton Boulenger, 1878

+ Palaeopleurodeles Herre, 1941

Source : MNHN, Paris

Dugois 21

Paramesotriton Chang, 1935

Pleurodeles Michahelles, 1830

+ Procynops Young, 1965

Salamandra Laurenti, 1768

Salamandrina Fitzinger, 1826

Taricha Gray, 1850

Triturus Rafinesque, 1815

TBlototriton Anderson, 1871

Epifamilia SenoIDIA Gray, 1825

Superfamilia SiRENOIDEA Gray, 1825

Familia SIRENIDAE Gray, 1825

+ Habrosaurus Gilmore, 1928

+ Kababisha Evans, Milner & Werner, 1996

+ Norerperon Rage, Marshall & Gayet, 1993

Pseudobranchus Gray, 1825

Siren Osterdam, 1766

Superordo Gymnorioxa Rafinesque-Schmaltz, 1814

Ordo GymnorHiona Rafinesque-Schmaltz, 1814

Incertae sedis

Ÿ Rubricacaecilia Evans & Sigogneau-Russell, 2001

Epifamilia Cazcurombi4 Rafinesque-Schmaltz, 1814

Superfamilia CarciLrorpeA Rafinesque-Schmaltz, 1814

Familia Carcuupar Rafinesque-Schmaltz, 1814

+ Apodops Estes & Wake, 1972

Boulengerula Tornier, 1897

Brasiloryphlus Taylor, 1968

Caecilia Linnaeus, 1758

Dermophis Peters, 1879

Gegeneophis Peters, 1879

Geotrypetes Peters, 1880

Grandisonia Taylor, 1968

Gymmopis Peters, 1874

Herpele Peters, 1879

Hypogeophis Peters, 1879

Idiocranium Parker, 1936

Indotyphlus Taylor, 1960

Luetkenoryphlus Taylor, 1968

Microcaecilia Taylor, 1968

Mimosiphonops Taylor, 1968

Oscaecilia Taylor, 1968

Parvicaccilia Taylor, 1968

Praslinia Boulenger, 1909

Schistometopum Parker, 1941

Siphonops Wagler, 1830

Sylvacaccilia Wake, 1987

Familia IcaravoranpAr Taylor, 1968 (1843)

Caudacaecilia Taylor, 1968

Ichthyophis Fitzinger, 1826

Familia ScozscomorPaipar Taylor, 1969

Crotaphatrema Nussbaum, 1985

Scolecomorphus Boulenger, 1883

Source : MNHN, Paris

22 ALYTES 23 (1-2)

Familia TieuLonecripas Taylor, 1968

Atretochoana Nussbaum & Wilkinson, 1995

Chthonerpeton Peters, 1879

Nectocaecilia Taylor, 1968

Poromoryphlus Taylor, 1968

Tÿphlonectes Peters, 1879

Familia Urasorvrauinar Nussbaum, 1979

Uraeoryphlus Peters, 1879

Superfamilia RmnarremarorneA Nussbaum, 1977

Familia Renarremarinar Nussbaum, 1977

Epicrionops Boulenger, 1883

Rhinatrema Duméril & Bibron, 1841

Epifamilia + EocarciLromnia Jenkins & Walsh, 1993

Superfamilia + Eocaseuropea Jenkins & Walsh, 1993

Familia + Eocasciupar Jenkins & Walsh, 1993

+ Eocacailia Jenkins & Walsh, 1993

ACKNOWLEDGMENTS

For helpful comments, suggestions and information that allowed to improve this paper, I am

grateful to Lauren E. Brown, Ronald I. Crombie, Frank Glaw, Jean-Claude Rage, Borja Sanchiz,

Miguel Vences, Nicolas Vidal and David B. Wake.

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à zoologique de France, 104:

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Alytes, 2005, 23 (1-2): 25-69. 25

Amphibia Mundi. 1.2.

Recent amphibians: generic

and infrageneric taxonomic additions

(1981-2002)

Alain Dugois*, Ronald I. CROMBIE** & Frank GLAW***

* Vertébrés: Reptiles et Amphibil

USM 0602 Taxonomie and Collections,

Département de Systématique and Evolution,

Muséum national d'Histoire naturelle,

25 rue Cuvier, 75005 Paris, France

<adubois@mnhn.fr>

#* California Academy of Sciences,

875 Howard Street, San Francisco, CA 94103-3098, USA

<rcrombie@calacademy.org>

*#* Zoologische Staatssammlung,

81247 München, Germany

<Frank.Glew@zsm.mwn.de>

The present list concerns additions in the taxonomy of NeosaTacuI (i.e., recent amphibians, taxa

represented by at least one species in the currently living fauna of our planet: see DuBors, 2004), for taxa

at rank genus and below, published before 2003 after the three lists of recent amphibians taxa of FROST

(1985), DuELLMAN (1993) and GLaw et al. (1998) and the two lists of fossil taxa of this group of ESTES

(1981) and SancHiz (1998), or absent from these five lists. The period covered by these additions starts

in 1981 for taxa of fossil gymnophiones and urodeles, in 1993 for taxa of recent amphibians, and in 1998

for taxa of fossil anurans. It ends on 31 December 2002 for all these groups. We tried to include all new

nomina that had been overlooked in the lists cited above, or for which we identified errors in these lists.

However, nomina of lower recent taxa anterior to 1993 not considered in Frosr's (1985) and

DuELLMaN’s (1993) checklists (i.e., most synonyms and most nomina of valid subgenera and subspe-

cies) were not included, as their inclusion would have increased the present list by hundreds, if not

thousands, of nomina. Most of these nomina anterior to 1° January 1970 are to be found in GORHAM

(1974), but a gap exists for the period 1970-1992.

Only new nomina are listed, and taxonomic or nomenclatural changes other than additions (e.g.,

synonymisation or revalidation of nomen, change of rank or higher taxonomic allocation of taxon,

subsequent type designation, first-reviser action, orthographic emendation) are not considered here.

The new nomina are listed below by alphabetical order under families and subfamilies according to the

general taxonomic frame of Dusois (2005a).

New nomina of the species-series (ie. species and subspecies; Dunois, 2000, 2005-c) are printed

in lower case Jralics, followed by the country of the sype-locality of the taxon. Although many species are

described on the basis of specimens collected in a single locality or a few neighbouring localities (see

Dubois, 2004: 24), this is not a general rule, and at any rate mention of the country of the type-locality

is not to be contrued as the known or inferred geographical distribution of the taxon. In most cases,

only the name of the country is given, but for a few countries that are either very large (Australia, Brazil,

Canada, China, India, USA) or that cover several important land masses (Indonesia, Malaysia), the

first-level administrative division (province, state, etc.) is also given. Names of countries are given in

English, but those of first-level administrative divisions are in the language of the country, even when

a common English translation exists.

Source : MNHN, Paris

26 ALYTES 23 (1-2)

New nomina of the genus-series (i.e., genus or subgenus; DuBois, 2000, 2005b-c) are printed in

lower case bold Italies, followed by the nomina of their sype-species and the country of the type-locality

of the latter (not the known or inferred geographical distribution of the taxon).

The nomina of fossil taxa are preceded by the sign +. For such taxa, beside the country of the

type-locality, the sratigraphical level of the latter (i.e., not the known or inferred stratigraphical

distribution of the taxon) is provided.

Nomenclaturally unavailable nomina (i.e., nomina nuda and other kinds of anoplonyms, as

defined by Dugois, 2000) are presented below “between quotation marks”.

In z0ological taxonomy, among two synonymous nomina, the valid one is the first published (rule

of priority). For this reason, knowing the actual date of public distribution (publication date) of a book

or paper is an important information. Below, publication dates as indicated on the works themselves are

accepted as true in the absence of contradictory information. Whenever more precise information is

available, the actual date of publication, either exact [day] or rough [month], is added between square

brackets after the year at the beginning of reference. The source of this information is given, also

between square brackets, at the end of the reference. When only incomplete information is available

(e.g., evidence that a work was published later than its printed date, but no precise publication date),

this information is also given between square brackets at the end of the reference. Publication dates as

given on the covers of some journals or reprints are not accepted as genuine evidence of exact

publication date, because the exact date at which a publication was actually distributed can be known

with certainty only afterwards (e.g., a planned date of distribution can have to be modified because of

unexpected delays, strike, etc.).

We did our best to provide the complete titles of publications and the names of administrative

divisions of countries in their original languages, as well as the names of authors, with proper accents

and other diacritic marks, e.g. using “6” and not “oe” or “o?”, “ñ” and not “n”, or “Z” and not “?”.

Such a respect for persons and languages other than English is becoming rare in many journals and

databases, even major ones. We would appreciate receiving corrections from readers if mistakes in this

respect remain in the present document. Titles ofworks were presented under their English translation

only when the title in the original language was not written in Roman alphabet.

This list only provides information on new nomina published in the period 1981-2002 for

amphibian taxa then considered new. Since their original description, some of these nomina have

already been synonymized, or have had a change of nomenclatural rank (subspecies raised to species

rank, or the reverse, etc.), or have been shown to apply to taxa that were wrongly allocated to higher

taxa: e.g., Scuriger mokokchungensis Das & Chanda, 2000 (PeLosaripAr, MeGorHRYINAE), which was later

shown to be a member ofthe Rap4r DicroGLossiNar. In the list below, only the original combination

(including its original mispellings if present) is given, not the subsequent synonymisation or change of

status. However, as nomina are listed under family-series taxa, a few taxa, for which wrong taxonomic

allocation is certain, were listed under their proper taxon, albeit in their original combination, in order

not to create any new combination in the present work. Reference to evidence supporting the

taxonomic transfer, when relevant, is provided as a “Comment” after the nomen. In a few cases, no

previous published statement is known to exist, so the responsibility of the transfer is acknowledged

between square brackets (e.g., for Rana charlesdartoini).

A few nomina were initially published with “incorrect original spellings” as defined by Art. 32.4 of

the Code. For such nomina, we provide the “correct original spelling” according to Art. 32.5 of the

Code, with reference to the first-user of this spelling if the latter has already been corrected. This applies

to species-series nomina mispelled because of wrong agreement in gender with the generic nomen

(Art. 31.2), but not to nomina unusually formed from personal names (Art. 31.1), for reasons explained

in detail by Crocuer & Dugois (244: 496). À few other nomina were initially published with “multiple

original spellings” as defined by Art. 19.3 of the Code: in such cases, we provide information of the

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 27

“correct original spelling” chosen by the first-reviser (Art. 24.2.3 and 32.2.1), and, if no such

first-reviser action has yet been taken, we provide it.

The present list will be regularly followed by updates published in Amphibia Mundi. Although we

did our best to collect all the available information on amphibian taxonomic novelties published from

1981 to 2002 that had not been provided in the five lists mentioned above, no doubt we overlooked a

number of publications. This is unavoidable, as even the Zoological Record, with an experience of one

and a half century, still overlooks a large proportion of publications and new nomina (BOUCHET &

Rockoï, 1992, 1993). Readers and users of this first list are therefore encouraged to send us corrections

and additions. This new information will be included in our subsequent lists. Furthermore, in order to

avoid our overlooking their forthcoming works, all amphibian taxonomists worldwide are strongly

encouraged to send spontaneously a copy of each of their publications (books and reprints) having

taxonomic, nomenclatural or distributional contents to the coordinator of Amphibia Mundi, Alain

Duois (Reptiles & Amphibiens, Muséum national d'Histoire naturelle, 25 rue Cuvier, 75005 Paris,

France). The works so received will be deposited in the public herpetological library of the Paris

Museum, where they will be freely available to all visitors and library users.

Classis Aura De Blainville, 1816

Subclassis NroBarracun Sarasin & Sarasin, 1890

Superordo + ALLocauDarA Fox & Naylor, 1982

Ordo + ArLocaupara Fox & Naylor, 1982

Epifamilia + ALBANERPETONTOIDIA Estes & Hoffstetter, 1976

Superfamilia + ALBANERPETONTOIDEA Estes & Hofistetter, 1976

Familia + Azsanerreronrinar Estes & Hoffstetter, 1976

+ Albanerpeton arthridion Fox & Naylor, 1982. - USA (Texas). Cretaceous.

+ Albanerperon cifellit Gardner, 1999. - USA (Utah). Creraceous.

+ Albanerpeton galaktion Fox & Naylor, 1982. - Canada (Alberta). Cretaceous.

+ Albanerperon gracilis Gardner, 2000. - Canada (Alberta). Cretaceous.

+ Celtedens MeGowan & Evans, 1995. - Type-species, by original designation: + Triton megacephalus

Costa, 1864. - Italy. Cretaceous.

+ Celtedens iberius McGowan & Evans, 1995. - Spain. Cretaceous.

+ Nukusurus Nessov, 1981. - Type-species, by original designation: + Nukusurus insuetus Nessov,

1981. - Uzbekistan. Cretaceous.

+ Nukusurus insuetus Nessov, 1981. - Uzbekistan. Cretaceous.

+ Nukusurus sodalis Nessov, 1997. - Uzbekistan. Cretaceous.

Superordo BaraGia Brongniart, 1800

Ordo Axura Duméril, 1806

Incertae sedis

+ Avitabatrachus Bäez, Trueb & Calvo, 2000. - Type-species, by original designation: + Aviraba-

trachus uliana Bâez, Trueb & Calvo, 2000. - Argentina. Cretaceous.

Source : MNHN, Paris

28 ALYTES 23 (1-2)

Ÿ Avitabatrachus uliana Bâez, Trueb & Calvo, 2000. - Argentina. Cretaceous.

+ Mesophryne Gao &Wang, 2001.- Type-species, by original designation: + Mesophryne beipiaoensis

Gao & Wang, 2001. - China (Liaoning). Mesozoic.

+ Mesophryne beipiaoensis Gao & Wang, 2001. - China (Liaoning). Mesozoic.

+ Nezpercius Blob, Carrano, Rogers, Forster & Espinoza, 2001. - Type-species, by original

designation: + Nezpercius dodsoni Blob, Carrano, Rogers, Forster & Espinoza, 2001. - USA

(Montana). Cretaceous.

+ Nezpercius dodsoni Blob, Carrano, Rogers, Forster & Espinoza, 2001.- USA (Montana). Cretaceous.

+ Sunnybatrachus Evans & McGowan, 2002. - Type-species, by original designation: + Sumyba-

trachus purbeckensis Evans & McGowan, 2002. - England. Cretaceous.

+ Sunnybarrachus purbeckensis Evans & McGowan, 2002. - England. Cretaceous.

+ Thaumastosaurus wardi Holman & Harrison, 2002. - England. Eocene.

Epifamilia BomevaroromnA Gray, 1825

Superfamilia Bomsivarororpe Gray, 1825

Familia Bommnaroriar Gray, 1825

Incertae sedis

+ Callobatrachus Wang & Gao, 1997. - Type-species, by original designation: + Callobatrachus

sanvanensis Wang & Gao, 1997. - China (Liaoning). Jurassic-Cretaceous boundary.

+ Callobarrachus sanvanensis Wang & Gao, 1997. - China (Liaoning). Jurassic-Cretaceous boundary.

+ Latoglossus Hossini, 2000. - Type-species, by original designation: + Laroglossus zraus Hossini,

2000. - Morocco. Miocene.

+ Laroglossus zraus Hossini, 2000. - Morocco. Miocene.

Subfamilia AzyriNar Fitzinger, 1843

Alytes obstetricans pertinax Garcia-Paris & Martinez-Solano, 2001. - Spain.

Subfamilia Bommvarorinar Gray, 1825

Bombina lichuanensis Ye & Fei in Ye, Fe1 & YU, 1993. — China (Hubei). - Comment: Species

redescribed as new by YE & FE1 (1994a), with the same nomen and authors (see VE, Fet & Hu,

1993: 364), but nomen is available as from VE, Fet & Hu (1993: 113). Not being mentioned in the

nal publication of the nomen, the “holotype” designated by YE & FE1 (1994a: 22, 25) is in fact

the lectotype of this nominal species.

Subfamilia + GomariNAar Roëek & Nessov, 1993

+ Cretasalia Gubin, 1999. - Type-species, by original designation: Cretasalia 1sybini Gubin, 1999. -

Mongolia. Cretaceous.

+ Crerasalia 1sybini Gubin, 1999. - Mongolia. Cretaceous.

+ “Gobiates” Spinar, 1983. - Mongolia. Cretaceous. - Comment: Nomenclaturally unavailable

genus-series nomen, as published without designation of a type-species. Nomen made nomencla-

turally available in SpNAR & TATARINOV (1986).

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 29

Epifamilia Lerormaromta Mivart, 1869

Superfamilia Lerorszmaronnra Mivart, 1869

Familia Lerorzzmarinar Mivart, 1869

Subfamilia Lerorgzmarinar Mivart, 1869

Leiopelma pakeka Bell, Daugherty & Hay, 1998. - New Zealand.

Epifamilia Peosaroipla Bonaparte, 1850

Superfamilia PaLoraromea Bonaparte, 1850

Familia PeLogarpas Bonaparte, 1850

Incertae sedis

+ Liaobatrachus Ji Shu’an & Ji Qiang, 1998. - Type-species, by original designation: + Liaobatrachus

grabaui Ji Shu’an & Ji Quang, 1998. — China (Liaoning). Mesozoic.

+ Liaobatrachus grabaui Ji Shu’an & Ji Qiang, 1998. - China (Liaoning). Mesozoic.

Subfamilia MrcoPsryiN4r Noble, 1931 (1850)

Tribus Leprogracaunr Dubois, 1983

Leptobrachium banae Lathrop, Murphy, Orlov & Ho, 1998a. - Vietnam.

Leptobrachium hainanensis Ye & Fei in Ye, FEI & Hu, 1993. - China (Hainan).

Leptobrachium smithi Matsui, Nabhitabhata & Panha, 1999. - Thailand.

Leptobrachium xanthospilum Lathrop, Murphy, Orlov & Ho, 1998a. - Vietnam.

Leptobrachium (Vibrissaphora) echinatum Dubois & Ohler, 1998. Vietnam.

Leptolalax alpinus Fei, Ye & Li in FE1, YE & HuANG, 1991. - China (Yunnan). - Comment: Species

redescribed as new, with the same nomen, authors and holotype, in Fer, YE & Li (1992), but

nomen is available as from FE1, YE & HUANG (1991).

Leprolalax liui Fei & Ye in F1, YE & HUANG, 1991. - China (Fujian).

Leptolalax nahangensis Lathrop, Murphy, Orlov & Ho, 1998b. - Vietnam.

Leptolalax pluvialis Ohler, Marquis, Swan & Grosjean, 2000. — Vietnam.

Leprolalax sungi Lathrop, Murphy, Orlov & Ho, 1998b. - Vietnam.

Leptolalax tuberosus Inger, Orlov & Darevsky, 1999. - Vietnam.

Leptolalax ventripuncratus Fei, Ye & Li in FEI, YE & HUANG, 1991. - China (Yunnan). - Comment:

Species redescribed as new, with the same nomen, authors and holotype, in Frt, YE & Li (1992),

but nomen is available as from FEI, YE & HUANG (1991).

Oreolalax granulosus Fei, Ye & Chen in Fe1, YE & HUANG, 1991. - China (Yunnan). - Comment:

Species redescribed as new, with the same nomen, authors and holotype, in FEI, YE & CHEN

(1992), but nomen is available as from FEI, YE & HUANG (1991).

Oreolalax nanjiangensis Fei & Ye in FEt, YE & Li, 1999. - China (Yunnan).

Scutiger (Aelurophryne) bhuanensis Delorme & Dubois, 2001. - Bhutan.

Source : MNHN, Paris

30 ALYTES 23 (1-2)

Scutiger (Aelurophryne) jiulongensis Fei, Ye & Jiang in Fe1, JANG, Ye & CHEN, 1996. - China (Sichuan).

Scutiger (Oreolalax) xiangchengensis degenicus Yang & He, 1990. - China (Yunnan).

Tribus MrGoruryin Noble, 1931 (1850)

Brachytarsophrys chuannanensis Fei, Ye & Yang in Fer & YE, 2001a. - China (Sichuan).

Brachyrarsophrys platyparietus Rao &Yang, 19976. - China (Yunnan).

Megophrys caudoprocta Shen, 1994. - China (Hunan).

Megophrys daweimontis Rao &Yang, 1997a. - China (Yunnan).

Megophrys glandulosa Fei, Ye & Huang, 1991. - China (Yunnan). - Comment: Species redescribed as

new, with the same nomen, authors and holotype, in FEI, YE & HUANG (1992), but nomen is

available as from FEI, YE & HUANG (1991).

Megophrys mangshanensis Fei &Ye in FE1, YE & HuANG, 1991. China (Hunan). - Comment: Species

redescribed as new, with the same nomen, authors and holotype, by Fet, YE & HUaNG (1992), but

nomen is available as from FEI, YE & HUANG (1991).

Megophrys minor binchuanensis Ye & Fei, 1995. - China (Yunnan).

Megophrys shuichengensis Tian, Gu & Sun, 2000. - China (Guizhou).

Megophrys watouensis Fei, Jiang & Zheng in Fer & Ye, 2001a. - China (Sichuan). - Comment: Species

redescribed in detail by JIANG, FEI, ZHENG, YE, XIE & CHEN (2002).

Megophrys wuliangshanensis Ye & Fei, 1995. - China (Yunnan).

Megophrys wushanensis Ye & Fei, 1995. - China (Sichuan).

Megophrys zhangi Ye & Fei, 1993. - China (Xizang).

Megophrys (Xenophrys) auralensis Ohler, Swan & Daltry, 2002. - Cambodia.

Panophrys Rao & Yang, 1997. - Type-species, by original designation: Megophrys omeimontis Liu,

1950. - China (Sichuan).

Subfamilia Popyrinae Bonaparte, 1850

Pelodytes ibericus Sänchez-Herräiz, Barbadillo, Machordom & Sanchiz, 2000. - Spain.

Epifamilia Prporpia Gray, 1825

Superfamilia Prornea Gray, 1825

Familia + Pazarorarrammpar Cope, 1865

Ÿ Palaeobatrachus robustus Hossini & Rage, 2000. - France. Miocene.

Familia Priar Gray, 1825

Subfamilia Dacrvzeraenar Hogg, 1838

+ Pachybatrachus Bâez & Rage, 1998. — Type-species, by original designation: + Pachybatrachus

raqueti Bâez & Rage, 1998. - Niger. Cretaceous.

+ Pachybatrachus raqueri Bâez & Rage, 1998. - Niger. Cretaceous.

Source : MNHN, Paris:

Dugois, CROMBIE & GLAW 31

+ Shelania laurenti Bâez & Pugener, 1998. - Argentina. Palacogene.

+ Xenopus arabiensis Henrici & Bâez, 2001. - Yemen. Oligocene.

Familia RHINOPHRYNIDAE Günther, 1858

+ Rhadinosteus Henrici, 1998. — Type-species, by original designation: + Rhadinosteus parous

Henrici, 1998. - USA (Utah). Jurassic.

+ Rhadinosteus parous Henrici, 1998. - USA (Utah). Jurassic.

Epifamilia Ravornra Rafinesque-Schmaltz, 1814

Superfamilia HyLoprA Rafinesque, 1815

Familia Buronipar Gray, 1825

Adenomus dasi Manamendra-Arachchi & Pethiyagoda, 1998. - Sri Lanka.

Ansonia anotis Inger, Tan &Yambun, 2001. - Malaysia (Sabah).

Ansonia inthanon Matsui, Nabhitabhata & Panha, 1998. - Thailand.

Ansonia kamblei Ravichandran & Pillai, 1996. — India (Maharashtra).

Auelopus angeliro Ardila-Robayo & Ruiz-Carranza, 1998. - Colombia.

Atelopus guanujo Coloma, 2002. - Ecuador.

Atelopus guitarraensis Osorno-Muñoz, Ardila-Robayo & Ruiz-Carranza, 2001. - Colombia.

Atelopus lozanoi Osorno-Muñoz, Ardila-Robayo & Ruiz-Carranza, 2001. - Colombia.

Atelopus mandingues Osorno-Muñoz, Ardila-Robayo & Ruiz-Carranza, 2001. - Colombia.

Atelopus mono-hernandezit Ardila-Robayo, Osorno-Muñoz & Ruiz-Carranza, 2002. - Colombia. -

Comment: The original spelling of the epithet of this new species is incorrect and should be

emended into monohernandezi according to Art. 32.5.2.3 of the Code.

Atelopus nanay Coloma, 2002. - Ecuador.

Atelopus petriruizii Ardila-Robayo, 1999. - Colombia.

Atelopus reticulatus Lôtters, Haas, Schick & Bôhme, 2002. - Peru.

Atelopus siranus Lôtrers & Henzl, 2000. - Peru.

Arelopus sonsonensis Vélez-Rodriguez & Ruiz-Carranza, 1997. - Colombia.

Bufo amieti Tandy & Perret, 2000. — Ivory Coast.

Bufo chavin Lehr, Kôhler, Aguilar & Ponce, 2001. - Peru.

Bufo cristinae Vélez-Rodriguez & Ruiz-Carranza, 2002. - Colombia.

Bufo danatensis taxhorensis Fei, Ye & Huang in FEL, YE, HUANG & CHEN, 1999. - China (Xi

Bufo jimi Stevaux, 2002. - Brazil (Bahia).

Bufo kumquat Das & Lim, 2001. - Malaysia (West Malaysia).

Bufo leucomyos MeCranie & Wilson, 2000. - Honduras.

Bufo melanostictus hazarensis Khan, 2001. - Pakistan.

Bufo noellerti Manamendra-Arachchi & Pethiyagoda, 1998. - Sri Lanka.

Bufo pseudoraddei baturae Stôck, Schmid, Steinlein & Grosse, 1999. - Pakistan.

Bufo sclerocephalus Mijares-Urrutia & Arends, 2001. - Venezuela.

Bufo stanlai Lôtrers & Kôhler, 2000. - Bolivia.

Bufo taiensis Rôdel & Ernst, 2000. — Ivory Coast.

Bufo zamdaensis Feï, Ye & Huang in FEI, YE, HUANG & CHEN, 1999. - China (izang).

Churamiti Channing & Stanley, 2002. - Type-species, by original designation: Churamiti maridadi

Channing & Stanley, 2002. - Tanzania.

Churamiti maridadi Channing & Stanley, 2002. - Tanzania.

Melanophryniscus klappenbachi Prigioni & Langone, 2000. - Argentina.

Source : MNHN, Paris

32 ALYTES 23 (1-2)

Melanophryniscus simplex Caramaschi & Cruz, 2002. - Brazil (Santa Catarina).

Melanophryniscus spectabilis Caramaschi & Cruz, 2002. - Brazil (Santa Catarina).

Necrophrynoides asperginis Poynton, Howell, Clarke & Lovett, 1999. - Tanzania.

Rhamphophryne ruizi Grant, 2000. - Colombia.

Stephopaedes howelli Poynton & Clarke, 1999. - Tanzania.

Stephopaedes usambarae Poynton & Clarke, 1999. - Tanzania.

“Torrentophryne” Rao &Yang, 1994. - China (Yunnan).- Comment: Nomenclaturally unavailable

genus-series nomen, as published without designation of a type-species.

Torrentophryne Yang in YANG, Liu & RAo, 1996. - Type-species, by original designation: Torren-

tophryne aspinia Yang & Rao, 1996. - China (Yunnan).

Torrentophryne aspinia Rao & Yang, 1994. - China (Yunnan). - Comments: (1) Although published

combined with a nomenclaturally unavailable genus-series nomen, this specific nomen is available

as the Code expressly states that the generic nomen with which a new specific nomen must be

combined “need not be valid or even available” (ANONYMOUS, 1999: Art. 11.9.3.1). (2) Species

redescribed as new by YANG & RAO in YANG, Liu & Rao (1996), with the same nomen but with a

different order of names of authors, but nomen is available as from Ra & YANG (1994). Not being

mentioned in the original publication of the nomen, the “holotype” designated by YANG & RAO in

YANG, Liu & RAO (1996) is in fact the lectotype of this nominal species.

“T{orrentophryne] tuberculous” Rao & Yang, 1994. - Nomen nudum.

Torrentophryne tuberospinia Yang & Liu in YANG, Liu & RA0, 1996. - China (Yunnan). - Comment:

Specific nomen misspelled ruberospina in GLAW et al. (1998).

Wolterstorffina chirioi Boistel & Amiet, 2001. - Cameroon.

Familia CENTROLENIDAE Taylor, 1951

Centrolene papillahallicum Noonan & Harvey, 2000. - Guyana.

Cochranella rosada Ruiz-Carranza & Lynch, 1997. - Colombia.

Cochranella spilota Ruiz-Carranza & Lynch, 1997. - Colombia.

Hyalinobatrachium crurifasciatum Myers & Donnelly, 1997. - Venezuela.

Hyalinobatrachium eccentricum Myers & Donnelly, 2001. - Venezuela.

Hyalinobatrachium esmeralda Ruiz-Carranza & Lynch, 1998. - Colombia.

Hyalinobatrachium guairarepanensis Señaris, 2001. - Venezuela.

Hyalinobatrachium ibama Ruiz-Carranza & Lynch, 1998. - Colombia.

Hyalinobatrachium mondolfii Señaris & Ayarzagüena, 2002a. - Venezuela.

Hyalinobatrachium nouraguensis Lescure & Marty, 2000. - French Guyana.

Hyalinobatrachium ruedai Ruiz-Carranza & Lynch, 1998. - Colombia.

Familia DENDRoBATIDAE Cope, 1865 (1850)

Colostethus alessandroi Grant & Rodriguez, 2001. - Peru.

Colostethus atopoglossus Grant, Humphrey & Myers, 1997. - Colombia.

Colostethus avarzaguenai La Marca, 1996. - Venezuela.

Colostethus borjai Rivero & Serna, 1995. - Colombia.

Colostethus cacerensis Rivero & Serna, 1995. - Colombia.

Colostethus caeruleodactylus Lima & Caldwell, 2001. - Brazil (Amazonas).

Colostethus cepedai Morales, 2002. - Colombia.

Colostethus conspicuus Morales, 2002. - Peru.

Colostethus crombiei Morales, 2002. - Brazil (Parä).

Colostethus dysprosium Rivero & Serna, 1995. - Colombia.

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 23;

Colostethus erasmios Rivero & Serna, 1995. - Colombia.

Colostethus excisus Rivero & Serna, 1995. - Colombia.

Colostethus fascianigrus Grant & Castro, 1998. - Colombia.

Colostethus fratisenescus Morales, 2002. - Ecuador.

Colostethus fuscellus Morales, 2002. - Brazil (Amazonas).

Colostethus gasconi Morales, 2002. - Brazil (Amazonas).

Colostethus guanayensis La Marca, 1996. - Venezuela.

Colostethus insperatus Morales, 2002. - Ecuador.

Colostethus larandinus Yüstiz, 1991. - Venezuela.

Colostethus Iynchi Grant, 1998. - Colombia.

Colostethus masniger Morales, 2002. - Brazil (Paré).

Colostethus melanolaemus Grant & Rodriguez, 2001. - Peru.

Colostethus murisipanensis La Marca, 1996. - Venezuela.

Colostethus ornatus Morales, 2002. - Peru.

Colostethus parimae La Marca, 1996. - Venezuela.

Colostethus picachos Ardila-Robayo, Acosta-Galvis & Coloma, 2000. - Colombia.

Colostethus praderioi La Marca, 1996. - Venezuela.

Colostethus pseudopalmatus Rivero & Serna, 1995. - Colombia.

Colostethus ramirezi Rivero & Serna, 1995. - Colombia.

Colostethus roraima La Marca, 1996. - Venezuela.

Colostethus saltuarius Grant & Ardila-Robayo, 2002. - Colombia.

Colostethus sumtuosus Morales, 2002. - Brazil (Par).

Colostethus tamacuarensis Myers & Donnelly, 1997. - Venezuela.

Colostethus tepuyensis La Marca, 1996. - Venezuela.

Colostethus undulatus Myers & Donnelly, 2001. - Venezuela.

Colostethus vanzolinius Morales, 2002. — Brazil (Amazonas).

Colostethus soayuu Acosta, Cuentas & Coloma, 2000. - Colombia.

Cryptophyllobates Lôtters, Jungfer & Widmer, 2000. — Type-species, by original designation:

Phyllobates azureiventris Kneller & Henle, 1985. - Peru.

Dendrobates amazonicus Schulte, 1999. - Peru.

Dendrobates claudiae Jungfer, Lôtters & Jôrgens, 2000. - Panama.

Dendrobates duelmani Schulte, 1999. - Peru.

Dendrobates flavovitratus Schulte, 1999. - Peru.

Dendrobates imitator intermedius Schulte, 1999. - Peru.

Dendrobates imirator yurimaguensis Schulte, 1999. - Peru.

Dendrobates rubrocephalus Schulte, 1999. - Peru.

Epipedobates planipaleae Morales & Velazco, 1998. - Peru.

Epipedobates pongoensis Schulte, 1999. - Peru.

Epipedobates simulans Myers, Rodriguez & Icochea, 1998. - Peru.

Mannophryne caquetio Mäjares-Urrutia & Arends R.; 19996. - Venezuela.

Mannophryne lamarcai Müjares-Urrutia & Arends R., 1999a. - Venezuela.

Familia Hyzipar Rafinesque, 1815

Subfamilia HemipHRacTiNAr Peters, 1862

Gastrorheca sticropleura Duellman, Lehr & Aguilar, 2001. - Peru.

Hemiphractus helioi Sheil & Mendelson, 2001. - Peru.

Stefania ackatwaio MacCulloch & Lathrop, 2002. - Guyana.

Source : MNHN, Paris

34 ALYTES 23 (1-2)

Stefania ayangannae MacCulloch & Lathrop, 2002. - Guyana.

Stefania coxi MacCulloch & Lathrop, 2002. - Guyana.

Stefania oculosa Señaris, Ayarzagüena & Gorzula, 1996. - Venezuela.

Stefania percristata Señaris, Ayarzagüena & Gorzula, 1996. - Venezuela.

Stefania riveroi Señaris, Ayarzagüena & Gorzula, 1996. - Venezuela.

Stefania satelles Señaris, Ayarzagüena & Gorzula, 1996. - Venezuela.

Stefania schuberti Señaris, Ayarzagüena & Gorzula, 1996. - Venezuela.

Stefania ramacuarina Myers & Donnelly, 1997. - Venezuela.

Subfamilia Hyuar Rafinesque, 1815

Hyla abdivita Campbell & Duellman, 2000. - Mexico.

Hyla ameibothalame Canseco-Märquez, Mendelson & Gutiérrez-Mayén, 2002. - Mexico.

Hyla amicorum Mijares-Urrutia, 1998. -Venezuela.

Hyla annectans chuanxiensis Ye & Fei in Y£, Fet, Li & Li, 2000. - China (Sichuan).

Hyla annectans jingdongensis Ye & Fei in Ye, Fer, Li & Li, 2000. - China (Yunnan). - Comment:

his

new nomen appears under two different spellings in the original publication: jingdongensis (once in

p.88, twice in p. 89, twice in p. 91, twice in p. 93) and jindongensis (once in p. 89). These spellings

are “multiple original spellings” according to the Code. Acting as first revisers, we hereby choose

the spelling jingdongensis as “correct original spelling” of this nomen.

Hyla annectans tengchongensis Ye, Fei & Li in Ye, Fet, Li & Li, 2000. - China (Yunnan). - Comment:

This new nomen appears under three different spellings in the original publication: tengchongensis

(once in p. 88, twice in p. 89, once in p. 90, once in p. 91, once in p. 93), rangchongensis (once in

p.91) and renchongensis (once in p. 93). These spellings are “multiple original spellings” according

to the Code. Acting as first revisers, we hereby choose the spelling rengchongensis as “correct original

spelling” of this nomen.

Hyla annectans wulingensis Shen, 1997. - China (Hunan).

Hyla araguaya Napoli & Caramaschi, 1998. — Brazil (Mato Grosso).

Hyla buriti Caramaschi & Cruz, 1999. - Brazil (Minas Gerais).

Hyla cachimbo Napoli & Caramaschi, 1999b. - Brazil (Par).

Hyla calthula Ustach, Mendelson, McDiarmid & Campbell, 2000. - Mexico.

Hyla cerradensis Napoli & Caramaschi, 1998. - Brazil (Mato Grosso do Sul).

Hyla cruzi Pombal & Bastos, 1998. - Brazil (Goiâs).

Hyla cyelada Campbell & Duellman, 2000. - Mexico.

Hÿla delarivai Kôhler & Lôtwers, 20016. - Bolivia.

Hyla dendrophasma Campbell, Smith & Acevedo, 2000. - Guatemala.

Hyla elianeae Napoli & Caramaschi, 2000. - Brazil (Mato Grosso do Sul).

Hyla ericae Caramaschi & Cruz, 2000. - Brazil (Goiäs).

Hyla gaucheri Lescure & Marty, 2000. - French Guyana.

Hyla jimi Napoli & Caramaschi, 19994. - Brazil (Säo Paulo).

Hyla joannae Kôhler & Lôtters, 2001a. - Bolivia.

Hyla nephila Mendelson & Campbell, 1999. - Mexico.

Hyla palaestes Duellman, De la Riva & Wild, 1997. - Peru.

Hyla phaeopleura Caramaschi & Cruz, 2000. - Brazil (Goiäs).

Hyla psarosema Campbell & Duellman, 2000. - Mexico.

Ha pseudomeridiana Cruz, Caramaschi & Dias, 2000. - Brazil (Rio de Janeiro).

Ha ravida Caramaschi, Napoli & Bernardes, 2001. - Brazil (Minas Gerais).

Hyla rhea Napoli & Caramaschi 199%. - Brazil (So Paulo)

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 35

Hyla rhythmicus Señaris & Ayarzagüena, 20026. - Venezuela. - Comment: The original spelling of the

epithet of this new species is incorrect and should be emended into rhyhmica according to

Art. 31.2 of the Code.

Hyla simplex hainanensis Fei &Ye, 2000b. - China (Hainan).

Hyla stenocephala Caramaschi & Cruz, 1999. - Brazil (Minas Gerais).

Hyla yaracuyana Mijares-Urrutia & Rivero, 2000. - Venezuela.

Osteocephalus ayarzaguenai Gorsula & Señaris, 2000. - Venezuela.

Osteocephalus deridens Jungfer, Ron, Seipp & Almendäriz, 2000. - Ecuador.

Osteocephalus exophthalmus Smith & Noonan, 2001. - Guyana.

Osteocephalus fuscifacies Jangfer, Ron, Seipp & Almendäriz, 2000. - Ecuador.

Osteocephalus heyeri Lynch, 2002. - Colombia.

Osteocephalus leoniae Jangfer & Lehr, 2001. - Peru.

Osteocephalus mutabor Jangfer & Hôdl, 2002. - Ecuador.

Osteocephalus yasuni Ron & Pramuk, 2000. - Ecuador.

Plectrohyla exquisita MeCranie & Wilson, 1998. - Honduras.

Plectrohyla psiloderma MeCranie & Wilson, 1999a. - Honduras.

Pseudis cardosoi Kwet, 2000. - Brazil (Rio Grande do Sul).

Pseudis tocantins Caramaschi & Cruz, 1998. — Brazil (Tocantins).

Prychohyla acrochorda Campbell & Duellman, 2000. - Mexico.

Pychohyla zophodes Campbell & Duellman, 2000. - Mexico.

Scinax arduous Peixoto, 2002. - Brazil (Espirito Santo).

Scinax jolyi Lescure & Marty, 2000. - French Guyana.

Tépuihyla Ayarzagüena, Señaris & Gorzula, 1993. - Type-species, by original designation: Hyla

rodriguezi Rivero, 1968. - Venezuela.

Rpuihyla celsae Mijares-Urrutia, Manzanilla-Puppo & La Marca, 2000. - Venezuela.

Xenohyla Izecksohn, 1998. - Type-species, by original designation: Hyla wuncata Izecksohn,

1959. — Brazil (Rio de Janeiro).

Xenohyla eugenioi Caramaschi, 1998. - Brazil (Bahia).

Subfamilia PELODRYADIN,

E Günther, 1858

Litoria andiürrmalin McDonald, 1997. - Australia (Queensland).

Litoria daviesae Mahony, Knowles, Foster & Donnellan, 2001. - Australia (New South Wales).

Litoria elkeae Günther & Richards, 2000. - Indonesia (Irian Jaya).

Litoria macki Richards, 2001. - Indonesia (Irian Jaya).

Litoria wapogaensis Richards & Iskandar, 2001. — Indonesia (Irian Jaya).

Subfamilia PayzLomepusiNAr Günther, 1858

Phyllomedusa camba De la Riva, 2000. - Bolivia.

Phyllomedusa oreades Brandäo, 2002. - Brazil (Goiäs).

Familia Lepropacrvuipar Werner, 1896 (1838)

Incertae sedis

+ Estesiella Bäez, 1995. - Bolivia. Paleocene. - Comment: Nomen novum pro Estesius Bâez, 1995

Inec Estesius Wallach, 1984].

Source : MNHN, Paris

36 ALYTES 23 (1-2)

Subfamilia BracaycerHALINAE Günther, 1858

Brachycephalus pernix Pombal, Wistuba & Bornschein, 1998. - Brazil (Parané).

Brachycephalus vertebralis Pombal, 2001. - Brazil (Rio de Janeiro).

Eleutherodactylus actinolaimus Lynch & Rueda-Almonacid, 1998. - Colombia.

Eleutherodactylus amniscola Campbell & Savage, 2000. - Guatemala.

Eleutherodactylus anemerus Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus angustilineatus Lynch, 1998a. - Colombia.

Eleutherodactylus anthrax Lynch, 20016. - Colombia.

Eleutherodactylus araiodactylus Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus ardalonychus Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus ashkapara Kôhler, 2000b. - Bolivia.

Eleutherodactylus atrabracus Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus avicuporum Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus avius Myers & Donnelly, 1997. - Venezuela.

Eleutherodactylus baiotis Lynch, 1998a. - Colombia.

Eleutherodactylus blairhedgesi Estrada, Diaz & Rodriguez, 1998. - Cuba.

Eleutherodactylus capitonis Lynch, 1998a. - Colombia.

Eleutherodactylus catalinae Campbell & Savage, 2000. - Costa Rica.

Eleutherodactylus cavernibardus Myers & Donnelly, 1997. - Venezuela.

Eleutherodactylus charadra Campbell & Savage, 2000. - Guatemala.

Eleutherodactylus coffeus MeCranie & Kôhler, 19996. - Honduras.

Eleutherodactylus cuneirostris Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus duende Lynch, 20014. - Colombia.

Eleutherodacrylus dundeei Heyer & Muñoz, 1999. - Brazil (Mato Grosso).

Eleutherodactylus epacrus Lynch & Suärez-Mayorga, 2000. - Colombia.

Eleutherodactylus exoristus Duellman & Pramuk, 1999. - Ecuador.

Eleutherodactylus factiosus Lynch & Rueda-Almonacid, 1998a, - Colombia.

Eleutherodactylus fallax Lynch & Rueda-Almonacid, 1999. - Colombia.

Eleutherodactylus ferosus Lynch & Rueda-Almonacid, 1998a. - Colombia.

Eleutherodactylus glamyrus Estrada & Hedges, 1997e. - Cuba.

Eleutherodactylus helvolus Lynch & Rueda-Almonacid, 1998b, - Colombia.

Eleutherodactylus ibischi Reichle, Lôtters & De La Riva, 2001. - Bolivia.

Eleutherodactylus inachus Campbell & Savage, 2000. - Guatemala.

Elutherodactylus infraguttatus Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus jaumei Estrada & Alonso, 1997. - Cuba.

Eleutherodactylus kelephus Lynch, 1998a. - Colombia.

Eleutherodacrylus lemur Lynch & Rueda-Almonacid, 1998b. - Colombia.

Eleutherodactylus llojsintura Kôhler & Lôtters, 1999. - Bolivia.

Eleutherodactylus melanogaster Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus memorans Myers & Donnelly, 1997. - Venezuela.

Eleutherodactylus merabates Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus mmionaeres Lynch, 1998b, - Colombia.

Ekutherodactylus muscosus Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus myllomyllon Savage, 2000. - Guatemala.

Eleutherodactylus myops Lynch, 1998a. - Colombia.

Eleutherodactylus nephophilus Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus olanchano MeCranie & Wilson, 19996, - Honduras.

Eleutherodactylus olivaceus Kôhler, Morales, Lôtters, Reichle & Aparicio, 1998. - Bolivia.

Source : MNHN, Paris

DuBois, CROMBIE & GLAW

Eleutherodactylus operosus Savage, McCranie & Wilson, 1999. - Honduras.

Eleutherodactylus opimus Savage & Myers, 2002. - Colombia.

Eleutherodactylus paisa Lynch & Ardila-Robayo, 1999, - Colombia.

Eleutherodactylus palenque Campbell & Savage, 2000. - Mexico.

Eleutherodactylus paranaensis Langone & Segalla, 1996. - Brazil (Paranä).

Eleutherodactylus parectatus Lynch & Rueda-Almonacid, 19986, - Colombia.

Eleutherodactylus pataikos Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus pechorum MeCranie & Wilson, 1999b, - Honduras.

Eleutherodacrylus pelorus Campbell & Savage, 2000. - Mexico.

Eleutherodactylus penelopus Lynch & Ruedas-Almonacid, 1999, - Colombia.

Eleutherodactylus perenopterus Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus phalarus Lynch, 1998a. - Colombia.

Eleutherodactylus pinguis Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus principalis Estrada & Hedges, 1997a. - Cuba.

Eleutherodactylus prochus Lynch, 1998a. - Colombia.

Eleutherodactylus quantus Lynch, 1998a. - Colombia.

Eleutherodactylus quidditus Lynch, 20016. - Colombia.

Eleutherodactylus renjiforum Lynch, 2000. - Colombia.

Eleutherodactylus rhodostichus Duellman & Pramuk, 1999. — Peru.

Eleutherodactylus rhyacobatrachus Campbell & Savage, 2000. - Costa Rica.

Eleutherodactylus riparius Estrada & Hedges, 1998. - Cuba.

Eleutherodactylus rivularis Diaz, Estrada & Hedges, 2001. - Cuba.

Eleutherodactylus rivulus Campbell & Savage, 2000. - Guatemala.

Eleutherodactylus rufioculis Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus rupinius Campbell & Savage, 2000. - Guatemala.

Eleutherodactylus sabrinus Campbell & Savage, 2000. - Guatemala.

Eleutherodactylus sambaqui Mendes Castanho & Haddad, 2000. — Brazil (Paranä).

Eleutherodactylus sanguineus Lynch, 1998. - Colombia.

Eleutherodactylus serendipitus Duellman & Pramuk, 1999. - Peru.

Eleutherodactylus simulans Diaz & Fong, 2001. - Cuba.

Eleutherodactylus suetus Lynch & Rueda-Almonacid, 19986. - Colombia.

Eleutherodactylus tinker Lynch, 2001, - Colombia.

Eleutherodactylus tonvi Estrada & Hedges, 19978, - Cuba.

Eleutherodactylus torrenticola Lynch & Rueda-Almonacid, 1998a. - Colombia.

Eleutherodactylus turpinorum Hardy, 2001. - Trinidad & Tobago.

Eleutherodactylus viejas Lynch & Ruedas-Almonacid, 1999. - Colombia.

Eleutherodactylus xeniolum Lynch, 2001a. - Colombia.

Eleutherodactylus zophus Lynch & Ardila-Robayo, 1999. - Colombia.

Phrynopus adenopleurus Aguayo, Rodrigo & Harvey, 2001. - Bolivia.

Phrynopus barthlenae Lehr & Aguilar, 2002. - Peru.

Phrynopus carpish Lehr, Rodriguez & Cérdova, 2002. - Peru.

Phrynopus dagmarae Lehr, Aguilar & Kôhler, 2002. - Peru.

Phrynopus fallaciosus Duellman, 2000. - Peru.

Phrynopus heimorum Lehr, 2001. - Peru.

Phrynopus horstpauli Lehr, Kôhler & Ponce, 2000. - Peru.

Phrynopus iatamasi Aguayo, Rodrigo & Harvey, 2001. - Bolivia.

Phrynopus kauneorum Lehr, Aguilar & Kôhler, 2002. - Peru.

Phrynopus pinguis Harvey & Ergueta, 1998. - Bol

Phrynopus spectabilis Duellman, 2000. - Peru.

Phrynopus thompsoni Duellman, 2000. - Peru.

Source : MNHN, Paris

38 ALYTES 23 (1-2)

Phyillonastes carrascoicola De la Riva & Kôhler, 1998. - Bolivia.

Phyllonastes ritarasquinae Kôhler, 2000a. - Bolivia.

Psyillophryne hermogenesi Giaretta & Sawaya, 1998. - Brazil (Säo Paulo).

Subfamilia CerarorHRyINAE Tschudi, 1838

+ Ceratophrys ameghinorum Fernicola, 2001. - Argentina. Neogene.

Subfamilia CycLoramPHINAE Bonaparte, 1850

Paratelmatobius cardosoi Pombal & Haddad, 1999. - Brazil (Säo Paulo).

Paratelmatobius mantiqueira Pombal & Haddad, 1999. - Brazil (Säo Paulo).

Rupirana Heyer, 1999. - Type-species, by original designation: Rupirana cardosoi Heyer, 1999.

Brazil (Bahia).

Rupirana cardosoi Heyer, 1999. - Brazil (Bahia).

Subfamilia HyLoDN4E Günther, 1858

Hylodes amnicola Pombal, Feio & Haddad, 2002. - Brazil (Minas Gerais).

Hylodes dacrylocinus Pavan, Narvaes & Rodrigues, 2001. - Brazil (Säo Paulo).

Hylodes uai Nascimento, Pombal & Haddad, 2001. - Brazil (Minas Gerais).

Megaelosia boticariana Giaretta & Aguiar, 1998. - Brazil (Säo Paulo).

Subfamilia LepropacryLi4r Werner, 1896 (1838)

Adenomera araucaria Kwet, 2003. Brazil (Rio Grande do Sul).

Physalaemus maximus Feio, Pombal & Caramaschi, 1999. — Brazil (Minas Gerais).

Pseudopaludicola mirandae Mercadal de Barrio & Barrio, 1994. - Argentina.

Pseudopaludicola riopiedadensis Mercadal de Barrio & Barrio, 1994. - Brazil (Säo Paulo).

Subfamilia OnonroP#RYNINAE Lynch, 1969

Odontophrynus cordobae Martino & Sinsch, 2002. — Argentina.

Procerarophrys brauni Kwet & Faivovich, 2001. - Brazil (Rio Grande do Sul).

Procerarophrys concavirympanum Giaretta, Bernarde & Kokubum, 2000. - Brazil (Rondônia).

Proceratophrys cururu Eterovick & Sazima, 1998. — Brazil (Minas Gerais).

Subfamilia TëLmaroBuNAE Fitzinger, 1843

Alsodes australis Formas, Übeda, Cuevas & Nuñez, 1998. - Chile.

Alsodes hugoi Cuevas & Formas, 2001. - Chile.

Alsodes kaweshkari Formas, Cuevas & Nuñez, 1998. - Chile.

Alsodes valdiviensis Formas, Cuevas & Brieva, 2002. - Chile.

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 39

Atelognathus cei Basso, 1998. - Chile.

Télmatobius dankoi Formas, Northland, Capetillo, Nuñez, Cuevas & Brieva, 1999. - Chile.

Télmatobius fronteriensis Benavides, Ortiz & Formas, 2002. - Chile.

Télmatobius huayra Lavilla & Erguera, 1995. - Bolivia.

Télmatobius ifornoi Lavilla & Ergueta Sandoval, 1999. - Bolivia.

Télmatobius philippii Cuevas, 2002. - Chile.

Familia Mvosarracmar Schlegel, 1850

Subfamilia Lumonyxasrinar Lynch, 1969

Neobatrachus albipes Roberts, Mahony, Kendrick & Majors, 1991. - Australia (Western Australia).

Superfamilia Raoipra Rafinesque-Schmaltz, 1814

Familia Brevicirrribar Bonaparte, 1850

Subfamilia Asryzosrarninar Noble, 1927

Leptodactylodon blanei Ohler, 1999. - Gabon.

Leptodacrylodon æildi Amiet & Dowsett-Lemaire, 2000. - Cameroon.

Subfamilia Hemsonnar Cope, 1867

Hemisus barotseensis Channing & Broadley, 2002. - Zambia.

Subfamilia Hyrskozunar Laurent, 1943

Tribus Hyrsrouuni Laurent, 1943

Alexteroon hypsiphonus Amiet, 2000 - Cameroon.

Alexteroon jynx Amiet, 2000. - Cameroon.

Heterixalus carbonei Vences, Glaw, Jesu & Schimmenti, 2000. - Madagascar.

Hyperolius kihangensis Schiotz & Westergaard in Scmiorz, 1999. - Tanzania. - Comment: Species

redescribed in detail by Scxi@Tz & WESTERGAARD (2000).

Hyperolius nienokouensis Rôdel, 1999. — Ivory Coast.

Hyperolius pseudargus Schiotz & Westergaard in Scmiorz, 1999. - Tanzania. - Comment: Species

redescribed in detail by ScHiorz & WESTERGAARD (2000).

Tribus Kassiini Laurent, 1972

Kassina schioetzi Rôdel, Grafe, Rudolf & Ernst, 2002. - Ivory Coast.

Subfamilia LePropeLiNAE Laurent, 1972

Leptopelis sebra Amiet, 2001. - Cameroon

Source : MNHN, Paris

40 ALYTES 23 (1-2)

Familia Micronvziar Günther, 1858 (1843)

Subfamilia Asrsrorarvinar Günther, 1858

Tribus Asrsrormryivr Günther, 1858

Hylophorbus nigrinus Günther, 2001. - Indonesia (Irian Jaya).

Hylophorbus picoides Günther, 2001. - Indonesia (Irian Jaya).

Hylophorbus richardsi Günther, 2001. - Papua New Guinea.

Hylophorbus sextus Günther, 2001. - Indonesia (Irian Jaya).

Hylophorbus tetraphonus Günther, 2001. — Indonesia (Irian Jaya).

Hylophorbus wondixsoi Günther, 2001. - Indonesia (Irian Jaya).

Tribus XENORHININI Mivart, 1869

Xenobatrachus zweifeli Kraus & Allison, 2002. - Papua New Guinea.

Xenorhina arboricola Allison & Kraus, 2000. - Papua New Guinea.

Subfamilia CoryuiNar Cope, 1889

Stumpffia helenae Vallan, 2000. - Madagascar.

Subfamilia GenvorurYNINAE Boulenger, 1890

Albericus brunhildae Menzies, 1999. - Papua New Guinea.

Albericus fafniri Menzies, 1999. - Papua New Guinea.

Albericus gudrunae Menzies, 1999. - Papua New Guinea.

Albericus gunnari Menzies, 1999. - Papua New Guinea.

Albericus laurini Günther, 2000. - Indonesia (Irian Jaya).

Atbericus rhenaurum Menzies, 1999. - Papua New Guinea.

Albericus siegfriedi Menzies, 1999. - Papua New Guinea.

Albericus swanhildae Menzies, 1999. - Papua New Guinea.

Albericus valkuriarum Menvies, 1999. - Papua New Guinea.

Austrochaperina adamantina Zweïfel, 2000. - Papua New Guinea.

Austrochaperina aquilonia Zweïfel, 2000. - Papua New Guinea.

Austrochaperina archboldi Zweifel, 2000. - Papua New Guinea.

Austrochaperina blumi Zweifel, 2000. Indonesia (Irian Jaya).

Austrochaperina derongo Zweifel, 2000. - Papua New Guinea.

Austrochaperina guttara Zweifel, 2000. - Papua New Guinea.

Austrochaperina kosarek Zweifel, 2000. - Indonesia (Irian Jaya).

Austrochaperina novaebritanniae Zweïfel, 2000. - Papua New Guinea.

Austrochaperina parkeri Zweifel, 2000. - Papua New Guinea.

Austrochaperina rivularis, Zweifel, 2000. - Papua New Guinea.

Austrochaperina velaensis Zweifel, 2000. - Papua New Guinea.

Choerophryne longirostris Kraus & Allison, 2001. - Papua New Guinea.

Cophixalus bewaniensis Kraus & Allison, 2000. - Papua New Guinea.

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 41

Cophixalus pulchellus Kraus & Allison, 2000. - Papua New Guinea.

Cophixalus variegatus Richards, Johnston & Burton, 1992. - Papua New Guinea.

Cophixalus zweifeli Davies & McDonald, 1998. - Australia (Queensland).

Copiula expectata Günther, 20026. - Indonesia (Irian Jaya).

Copiula major Günther, 2002a. - Indonesia (Irian Jaya).

Copiula obsti Günther, 2002a. — Indonesia (Irian Jaya).

Liophryne allisoni Zweifel, 2000. - Papua New Guinea.

Liophryne rubra Zweifel, 2000. - Papua New Guinea.

Liophryne similis Zweifel, 2000. - Papua New Guinea.

Oreophryne atrigularis Günther, Richards & Iskandar, 2001. - Indonesia (Irian Jaya).

Oreophryne minuta Richards & Iskandar, 2000. — Indonesia (Irian Jaya).

Oreophryne wapoga Günther, Richards & Iskandar, 2001. - Indonesia (Irian Jaya).

Oxydactyla alpestris Zweifel, 2000. - Papua New Guinea.

Oxydactyla coggeri Zweïfel, 2000. - Papua New Guinea.

Oxydactyla stenodactyla Zweifel, 2000. - Papua New Guinea.

Subfamilia MicronyLiNar Günther, 1858 (1843)

Tribus Gasrroraryninr Fitzinger, 1843

Chiasmocleis alagoanus Cruz, Caramaschi & Freire, 1999. - Brazil (Alagoas).

Chiasmocleis jimi Caramaschi & Cruz, 2001. - Brazil (Amazonas).

Elachisocleis erythrogaster Kwet & Di-Bernardo, 1998. - Brazil (Rio Grande do Sul).

Orophryne pyburni Campbell & Clarke, 1998. - Colombia.

Tribus Micronyzini Günther, 1858 (1843)

Kalophrynus orangensis Dutta, Ahmed & Das, 2000. - India (Assam).

Kaloula walteri Diesmos, Brown & Alcala, 2002. - Philippines (Luzon).

Microhyla sholigari D'utta & Ray, 2000. — India (Karnataka).

Ramanella nagaoi Manamendra-Arachchi & Pethiyagoda, 20014. - Sri Lanka.

Familia RanDar Rafinesque-Schmaltz, 1814

Subfamilia CERATOBATRACHINAE Boulenger, 1884

Platymantis banahao Brown, Alcala, Diesmos & Alcala, 1997. - Philippines (Luzon).

Platymantis bimaculata Günther, 1999. - Indonesia (Irian Jaya).

Platymantis browni Allison & Kraus, 2001. - Papua New Guinea.

Plarymantis cagayanensis Brown, Alcala & Diesmos, 1999. - Philippines (Luzon).

Platymantis cryproris Günther, 1999. - Indonesia (Irian Jaya).

Platymantis indeprensus Brown, Alcala & Diesmos, 1999. - Philippines (Luzon).

Plarymantis isarog Brown, Brown, Alcala & Frost, 1997, - Philippines (Luzon). - Comment: Nomen

novum pro Plarymantis reticulatus Brown, Brown & Alcala, 1997 [nec Platymantis reticulatus Zhao

& Li, 1984].

Platymantis luzonensis Brown, Alcala, Diesmos & Alcala, 1997. - Philippines (Luzon).

+ Plawymantis megabotoniviti Worthy, 2001. - Fiji. Quaternary.

Source : MNHN, Paris

42 ALYTES 23 (1-2)

Plarymantis naomi Alcala, Brown & Diesmos, 1998. - Philippines (Luzon). - Comment: Although this

species was dedicated to a woman (Naomi Alcala), according to Art. 31.1.1 of the Code its nomen

does not have to be emended for reasons explained by CROCHET & DuBors (2004: 496).

Platymantis negrosensis Brown, Alcala, Diesmos & Alcala, 1997. - Philippines (Negros).

Platymantis pseudodorsalis Brown, Alcala & Diesmos, 1999. — Philippines (Luzon).

Platymantis pygmaeus Alcala, Brown & Diesmos, 1998. - Philippines (Luzon).

Platymantis rabori Brown, Alcala, Diesmos & Alcala, 1997. - Philippines (Bohol).

Platymantis sierramadrensis Brown, Alcala, Ong & Diesmos, 1999. - Philippines (Luzon).

Platymantis taylori Brown, Alcala & Diesmos, 1999. — Philippines (Luzon).

Subfamilia DicroGLOSSINAE Anderson, 1871

Tribus DicRoGLOSsINI Anderson, 1871

Féjervarya iskandari Veith, Kosuch, Ohler & Dubois, 2001. - Indonesia (Java).

Minervarya Dubois, Ohler & Biju, 2001. - Type-species, by original designation: Minervarya

sahyadris Dubois, Ohler & Biju, 2001. — India (Karnataka).

Minervarya sahyadris Dubois, Ohler & Biju, 2001. - India (Karnataka).

Tigrina Fei, Ye & Huang, 1991 [nec Tigrina Grevé, 1894]. - Type-species, by original designation:

Rana tigerina Daudin, 1802. - India (West Bengal).

Tomopterna maskevi Schleich & Anders, 1998. - Nepal. - Comment: The nomen Tomopterna is now

applied only to an African genus referred to the Pyx1c#PHALINAr (DuBois, 2003, 2005a); the Asian

species formerly placed in this genus are now referred to the genus Sphaerotheca, which is related

with Feyervarya (VENCES et al., 2000) and therefore considered a member of the DICROGLOSSINI

(Dusois, 2003).

Tribus LimNonecrINI Dubois, 1992

Limnonectes fujianensis Ye & Fei in Ye, Fe1 & Hu, 1993. - China (Fujian). - Comment: Species

redescribed as new by YE & FEI (1994b), with the same nomen and authors (see YE, FEI & HU,

1993: 370), but nomen is available as from YE, FEI & Hu (1993: 113). Not being mentioned in the

original publication of the nomen, the “holotype” designated by YE & Fer (1994b: 494, 4995) is in

fact the lectotype of this nominal species.

Liurana alpinus Huang & Ye, 1997. - China (Xizang).

Liurana medogensis Fei, Ye & Huang, 1997. - China (Kizang).

Rana charlesdarwini Das, 1998a. - India (Andamans & Nicobars). - Comment: This species with

forked omosternum is clearly not a member of the genus Rana, let alone of the RANINAF; pending

examination of specimens, it is here referred to the LimNONECTINI, without generic allocation.

[Alain Dugois].

Tribus Par Dubois, 1992

Paa (Feirana) taihangnicus Chen & Jiang, 2002. - China (Henan). - Comment: The original spelling

of the epithet of this new species is incorrect and should be emended into raïhangnica according to

Art. 31.2 of the Code, a justified emendation which was first used by JIANG et al. (2005).

Paa (Feirana) yei Chen, Qu & Jiang, 2002. - China (Henan). - Comments: (1) Species redescribed as

new, with the same nomen, authors and holotype, in CHEN, JIANG & QU (2004), but nomen is

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 43

available as from CHEN, Qu & JIanG (2002). (2) Although this species was dedicated to a woman

(Ye Changyuan), according to Art. 31.1.1 of the Code its nomen does not have to be emended for

reasons explained by CrocHET & Duois (2004: 496).

Paa (Paa) medogensis Fei & Ye, 20016. — China (Xizang).

Paa (Paa) rarica Dubois, Matsui & Ohler, 2001. - Nepal. - Comment: Nomen novum pro Rana

(Paa) rara Dubois & Matsui, 1983 [nec Rana danubina var. rara Fraas, 1903].

Rana robertingeri Wa & Zhao, 1995. - China (Sichuan). - Comment: Referred 10 the genus Paa by

JranG & ZHou (2005) and to the genus Quasipaa by JIANG et al. (2005).

Scutiger mokokchungensis Das & Chanda, 2000. - India (Nagaland). - Comment: Referred to the

genus Paa by DuBois (2002).

Unculuana Fei, Ye & Huang, 1991. - Type-species, by original designation: Rana unculuana Liu, Hu

& Yang, 1960. - China (Yunnan). - Comment: Created as a subgenus of Paa Dubois, 1975.

Subfamilia LaNKANECTINAE Dubois & Ohler, 2001

Lankanectes Dubois & Ohler, 2001. - Type-species, by original designation: Rana corrugata

Peters, 1863. - Sri Lanka.

Subfamilia MANTELLUINAE Laurent, 1946

Tribus Boop Vences & Glaw, 2001

Boophis bortae Vences & Glaw, 2002. - Madagascar.

Boophis feonnvala Glaw, Vences, Andreone & Vallan, 2001. - Madagascar.

Boophis haematopus Glaw, Vences, Andreone & Vallan, 2001. - Madagascar.

Boophis lichenoïdes Vallan, Glaw, Andreone & Cadle 1998. - Madagascar.

Boophis picturatus Glaw, Vences, Andreone & Vallan, 2001. - Madagascar.

Boophis pyrrhus Glaw, Vences, Andreone & Vallan, 2001. - Madagascar.

Boophis schuboeae Glaw & Vences, 20028. - Madagascar.

Boophis tasymena Vences & Glaw, 2002. - Madagascar.

Boophis vitratus Glaw, Vences, Andreone & Vallan, 2001. - Madagascar.

Tribus Larosrommni Vences & Glaw, 2001

Aglyptodactylus laticeps Glaw, Vences & Bôhme, 1998. - Madagascar.

Aglyptodactylus securifer Glaw, Vences & Bôhme, 1998. - Madagascar.

Laliostoma Glaw, Vences & Bôhme, 1998. - Type-species, by original designation: Tomopterna

labrosa Cope, 1868. - Madagascar. - Comment: Created as a subgenus of Jomoprerna Duméril &

Bibron, 1841.

Tribus ManreLuni Laurent, 1946

Chonomantis Glaw & Vences, 1994. - Type-species, by original designation: Rana albofrenata

Müller, 1892. - Madagascar. - Comment: Created as a subgenus of Mantidactylus Boulenger,

1895.

Mantella aurantiaca milorympanum Staniszewski, 1996. - Madagascar.

Source : MNHN, Paris

44 ALYTES 23 (1-2)

Mantella aurantiaca rubra Staniszewski, 1996. - Madagascar.

Mantella manery Vences, Glaw & Bôhme, 1999. - Madagascar.

Mantidactylus ambohitra Vences & Glaw, 20016. - Madagascar.

Mantidactylus brunae Andreone, Glaw, Vences & Vallan, 1998. — Madagascar.

Mantidactylus enki Glaw & Vences, 2002e. - Madagascar.

Mantidactylus kathrinae Glaw, Vences & Gossmann, 2000. - Madagascar.

Mantidactylus madinika Vences, Andreone, Glaw & Mattioli, 2002. - Madagascar.

Mantidactylus moseri Glaw & Vences, 20024. - Madagascar.

Mantidactylus sarotra Glaw & Vences, 2002a. - Madagascar.

Mantidactylus schilfi Glaw & Vences, 2000. - Madagascar.

Mantidactylus striatus Vences, Glaw, Andreone, Jesu & Schimmenti, 2002. - Madagascar.

Mantidactylus randroka Glaw & Vences, 2001. - Madagascar.

Mantidactylus 1schenki Glaw & Vences, 2001. - Madagascar.

Ochthomantis Glaw & Vences, 1994. - Type-species, by original designation: Rana femoralis

Boulenger, 1882. - Madagascar. - Comment: Created as a subgenus of Manridacrylus Boulenger,

1895.

Pandanusicola Glaw & Vences, 1994. - Type-species, by original designation: Rhacophorus bicalca-

ratus Boettger, 1913. - Madagascar. - Comment: Created as a subgenus of Mantidacrylus Bou-

lenger, 1895.

Phylacomantis Glaw &Vences, 1994. — Type-species, by original designation: Mantidactylus corous

Glaw & Vences, 1994. — Madagascar. - Comment: Created as a subgenus of Manridacrylus

Boulenger, 1895.

Subfamilia MicrixaLiN4E Dubois, Ohler & Biju, 2001

Micrixalus gadgili Pillai & Pattabiraman, 1990. - India (Kerala).

Subfamilia NycriBATRACHINAE Blommers-Schlôsser, 1993

Nyctibatrachus hussaini Krishnamurthy, Reddy & Gururaja, 2001. - India (Karnataka).

Subfamilia PerrorrpeTiNAE Noble, 1931

Arthroleptides vakusini Channing, Moyer & Howell, 2002. - Tanzania.

Subfamilia PHRYNOBATRACHINAE Laurent, 1941

Phrynobatrachus inexpectatus Largen, 2001. - Ethiopia.

Phrynobatrachus irangi Drewes & Perret, 2000. - Kenya.

Phrynobatrachus phyllophilus Rôdel & Ernst, 2002. - Ivory Coast.

Subfamilia PrvcHapeniNar Dubois, 1987

Prychadena fikuoha Largen, 1997. - Ethiopia.

Piychadena harenna Largen, 1997. - Ethiopia.

Prychadena wadei Largen, 2000. - Ethiopia.

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 45

Subfamilia Pyx1cEPHALINAE Bonaparte, 1850

Arthroleptella drewvesit Channing, Hendricks & Dawood, 1994. - South Africa.

Arthroleprella landdrosia Dawood & Channing, 2000. - South Africa.

Cacosternum karooicum Boycott, de Villiers & Scott, 2002. - South Africa.

Strongylopus kitumbeine Channing & Davenport, 2002. - Tanzania.

Tomopterna damarensis Dawood & Channing, 2002. - Namibia.

Subfamilia RanNAE Rafinesque-Schmaltz, 1814

Tribus Raninr Rafinesque-Schmaltz, 1814

Amolops bellulus Liu, Yang, Ferraris & Matsui, 2000. - China (Yunnan).

Amolops chakrataensis Ray, 1999. — India (Uttar Pradesh).

Amolops cremmobarus Inger & Kottelat, 1998. — Laos.

Amolops jaunsari Ray, 1999. - India (Uttar Pradesh).

Amolops mengyangensis Wa & Tian, 1995. - China (Yunnan).

Amolops spinapectoralis Inger, Orlov & Darevsky, 1999. - Vietnam.

Amolops tuberodepressus Liu & Yang, 2000. — China (Yunnan).

Amolops (Huia) modiglianit Doria, Salvidio & Tavano, 2001. - Indonesia (Sumatra).

Odorrana exiliversabilis Fei, Ye & Li, 20016. - China (Fujian).

Odorrana hainanensis Fei, Ye & Li, 2001a. - China (Hainan).

Odorrana jingdongensis Fei, Ye & Li, 2001a. - China (Yunnan).

Odorrana junlianensis Huang, Fei & Ye in Fe1 & YE, 2001a. - China (Sichuan).

Odorrana nasuta ei, Ye & Li, 20018. - China (Hainan).

“Pseudoamolops” Jiang, Fei, Ye, Zeng, Zhen, Xie & Chen, 1997. - Taiwan. - Comments: (1)

Created as a subgenus of Amolops Cope, 1865. (2) Nomenclaturally unavailable genus-series

nomen, as published without designation of a type-species.

Pseudoamolops Fei, Ye & Jiang, 2000. - Type-species, by original designation: Rana sauteri Bou-

lenger, 1909. - Taiwan.

Rana auigua Inger, Orlov & Darevsky, 1999. - Vietnam.

Rana balcanica Schneider & Sinsch, 1992 [nec Rana balcanica Schneider, Sinsch & Sofanidou, 1993].

— Greece. - Comment: See DuBois & OHLER (1995).

Rana bannanica Rao & Yang, 1997c. - China (Yunnan).

Rana chitwanensis Das, 19986. - Nepal.

Rana dhakuriensis Ray, 1997. - India (Uttar Pradesh)

Rana epeirotica Schneider, Sofianidou & Kyriakopoulou-Sklavounou, 1984. - Greece.

Rana huanrenensis Liu, Zhang & Liu, 1993. - China (Liaoning). - Comment: As noted by DUELLMAN

(1993: 262), the nomen of this species was made available by its publication in a key in FE1, YE &

HuaxG (1991: 131). The nomen Rana huanrenensis introduced by Liu, ZHANG & Liu (1993) in their

formal description of the species can be regarded either as a brand new nomen or as an unjustified

emendation of Rana huanrensis Feï, Ye & Huang, 1991 (see below). In both cases it is a distinct

available nomen and an invalid junior synonym of the latter nomen.

Rana huanrensis Fei, Ye & Huang, 1991. - China (Liaoning). - Comment: The original spelling

huanrensis appears three times in the original publication (FE1, YE & HUANG, 1991: 131, 298, 347),

so it cannot be considered an “inadvertent error”, and it does not have to be corrected because of

so-called “incorrect latinization” as the latter is not a case of “incorrect original spelling”

according to the Code (ANONYMOUS, 1999: Art, 32.5).

Source : MNHN, Paris

46 ALYTES 23 (1-2)

Rana kunyuensis Lu & Li, 2002. - China (Shandong).

Rana lessonae bergeri Günther, 1985. - Italy.

Rana lini Chou, 1999. - China (Yunnan).

Rana mangyanum Brown & Guttman, 2002. — Philippines (Mindoro).

Rana multidenticulata Chou & Lin, 1997. - Taiwan. - Comments: (1) Nomen mispelled Rana

multidentata in GLaw et al. (1998: xxii). (2) Species referred to the genus Pseudoamolops by FEI, YE

& JraNG (2000).

Rana omeimontis Ye & Fei in Ye, FE & Hu, 1993. - China (Sichuan).

Rana osca Paolucci, Fuhn & Bruno, 1993. - Italy.

Rana ridibunda caralitana Arikan, 1988.- Turkey.

Rana tipanan Brown, McGuire & Diesmos, 2000. - Philippines (Luzon).

Rana zhengi Zhao, 1999. - China (Sichuan).

Rana zhenhaiensis Ye, Fei & Matsui, 1995. - China (Zhejiang).

Rana (Sylvirana) faber Ohler, Swan & Daltry, 2002. - Cambodia.

Tenuirana Fci, Ye & Huang, 1991. - Type-species, by original designation: Rana taipehensis

Van Denburgh, 1909. -Taiwan. - Comment: Created as a subgenus of Hylarana Tschudi, 1838.

Subfamilia RaAacopaorINAE Hoffman, 1932 (1858)

Tribus PrzaurINI Dubois, 1981

Kurixalus Fei, Ye & Dubois in Fr1, 1999. - Type-species, by original designation: Rana eiffingeri

Boettger, 1895. - Japan.

Philautus abditus Inger, Orlov & Darevsky, 1999. - Vietnam.

Philautus cardamonus Ohler, Swan & Daltry, 2002. - Cambodia.

Philautus erythrophthalmus Stuebing & Wong, 2000. - Malaysia (Sabah).

Philautus griet Bossuyt, 2002. — India (Kerala).

Philautus odontotarsus Ye & Fei in Ye, Fet & Hu, 1993. - China (Yunnan).

Philautus terebrans Das & Chanda, 1998. - India (Andhra Pradesh).

Tribus Raacorsorini Hoffman, 1932 (1858)

Chirixalus dudhtoaensis Ray, 1999. India (Uttar Pradesh).

Polypedates fastigo Manamendra-Arachchi & Pethiyagoda, 20016. — Sri Lanka.

Polypedates pingbianensis Kou, Hu & Gao, 2001. - China (Yunnan).

Polypedates pseudocruciger Das & Ravichandran, 1998. - India (Tamil Nadu).

Polypedates puerensis He, 1999. - China (Yunnan).

Rhacophorus achantharrhena Harvey, Pemberton & Smith, 2002. - Indonesia (Sumatra).

Rhacophorus baliogaster Inger, Orlov & Darevsky, 1999. - Vietnam.

Rhacophorus barisant Harvey, Pemberton & Smith, 2002. - Indonesia (Sumatra).

Rhacophorus catamitus Harvey, Pemberton & Smith, 2002. - Indonesia (Sumatra).

Rhacophorus eyanopuncratus Manthey & Steiof, 1998. - Thailand.

Rhacophorus duboisi Ohler, Marquis, Swan & Grosjean, 2000. - Vietnam.

Rhacophorus exechopygus Inger, Orlov & Darevsky, 1999. - Vietnam.

Rhacophorus hoanglienensis Orlov, Lathrop, Murphy & Cuc, 2001. - Vietnam.

Rhacophorus orlovi Ziegler & Kôhler, 2001. - Vietnam.

Rhacophorus pseudomalabaricus Vasudevan & Dutta, 2000. - India (Tamil Nadu).

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 47

Superfamilia Sooczossores Noble, 1931

Familia Sooczossibar Noble, 1931

Sooglossus pipilodryas Gerlach & Willi, 2002. — Seychelles.

Ordo UropeLa Duméril, 1806

Incertae sedis

+ Apricosiren Evans & MeGowan, 2002. - Type-species, by original designation: + Apricosiren

ensomi Evans & McGowan, 2002, - England. Cretaceous.

+ Apricosiren ensomi Evans & McGowan, 2002. - England. Cretaceous.

+ Bishara Nessov, 1997. - Type-species, by original designation: Bishara backa Nessov, 1997. —

Kazakhstan. Cretaceous.

+ Bishara backa Nessov, 1997, - Kazakhstan. Creraceous.

+ Galverpeton Estes & Sanchiz, 1982. — Type-species, by original designation: + Galverperon

ibericum Estes & Sanchiz, 1982. - Spain. Cretaceous.

+ Galverpeton ibericum Estes & Sanchiz, 1982. - Spain. Cretaceous.

+ Jeholotriton Wang, 2000. — Type-species, by original designation: + Jeholorriton paradoxus

Wang, 2000. - China (Nei Mongol). Cretaceous.

+ Jeholorriron paradoxus Wang, 2000. - China (Nei Mongol). Cretaceous.

+ Kivatriton Averianov & Voronkevich, 2002. - Type-species, by original designation: + Kivarriton

leshchinskiyi Averianov & Voronkevich, 2002. - Russia. Cretaceous.

+ Kivarriton leshchinskivi Averianov & Voronkevich, 2002. - Russia, Cretaceous.

+ Laccotriton Gao, Cheng & Xu, 1998. — Type-species, by original designation: + Laccorriron

subsolanus Gao et al., 1998. - China (Hebei). Mesozoic.

+ Laccotriron subsolanus Gao, Cheng & Xu, 1998. - China (Hebei). Mesozoic.

+ Sinerpeton Gao & Shubin, 2001. - Type-species, by original designation: Ÿ Sinerperon fengshanensis

Gao & Shubin, 2001. - China (Hebei). Jurassic.

+ Sinerperon fengshanensis Gao & Shubin, 2001. — China (Hebei). Jurassic.

Familia BarracHosAURoIDIDAE Auffenberg, 1958

+ Mynbulakia Nessov, 1981.- Type-species, by original designation: + Mynbulakia surgayi Nessov,

1981. — Uzbekistan. Cretaceous.

+ Mynbulakia nongratis Nessov, 1981. - Uzbekistan. Cretaceous.

+ Mynbulakia surgayi Nessov, 1981. - Uzbekistan. Cretaceous.

+ Parrisia Denton & O’Neill, 1998. - Type-species, by original designation: + Parrisia neocesariensis

Denton & O'Neill, 1998. - USA (New Jersey). Cretaceous.

+ Parrisia neocesariensis Denton & O'Neill, 1998. - USA (New Jersey). Cretaceous.

+ Peratosauroides Naylor, 1983. - Type-species, by original designation: + Perarosauroides proble-

matica Naylor, 1983. - USA (California). Miocene.

+ Peratosauroides problematica Naylor, 1983. - USA (California). Miocene.

Familia + Scartereronripar Auffenberg & Goin, 1959

+Eoscapherpeton Nessov, 1981. - Type-species, by original designation: + Foscapherpeton asiaticumr

Nessov, 1981. - Uzbekistan. Cretaceous.

Source

: MNHN, Paris

48 ALYTES 23 (1-2)

+ Eoscapherpeton asiaticum Nessov, 1981. - Uzbekistan. Cretaccous.

+ Eoscapherpeton superum Nessov, 1997. - Tajikistan, Cretaceous.

+Horezmia Nessov, 1981. - Type-species, by original designation: + Horezmia gracile Nessov, 1981.

— Uzbekistan. Cretaceous.

+ Horezmia gracile Nessov, 1981. - Uzbekistan. Cretaceous.

Epifamilia CryPrograNcHoipiA Fitzinger, 1826

Superfamilia CrvrrosraxcnomrA Fitzinger, 1826

Familia Crvrroskancæinar Fitzinger, 1826

+ Andrias karelcapeki Ckhikvadze, 1982. - Kazakhstan. Miocene.

+ Aviturus Gubin, 1991. - Type-species, by original designation: + Avirurus exsecrarus Gubin,

1991. - Mongolia. Paleocene.

+ Aviturus exsecratus Gubin, 1991. - Mongolia. Paleocene.

+ Ulanurus Gubin, 1991. - Type-species, by original designation: + Ulanurus fractus Gubin,

1991. - Mongolia. Paleocene.

+ Ulanurus fractus Gubin, 1991. - Mongolia. Paleocene.

Familia Hyvonpar Cope, 1859 (1856)

Subfamilia Hyvopnvar Cope, 1859 (1856)

Batrachuperus taibaiensis Song, Zeng, Wu, Liu & Fu, 2001. - China (Shaanxi).

Hynobius amjiensis Gu, 1992. - China (Zhejiang).

Hynobius yunanicus Chen, Qu & Niu, 2001. - China (Henan).

+ Liaoxitriton Dong & Wang, 1998. - Type-species, by original designation: Ÿ Liaoxitriton zhongjiant

Dong & Wang, 1998. - China (Liaoning). Cretaceous.

+ Liaoxitriton zhongjiani Dong & Wang, 1998. - China (Liaoning). Cretaceous.

+ Parahynobius Venczel, 1999. - Type-species, by original designation: + Parahynobius betfianus

Venczel, 1999. - Romania. Pleistocene.

+ Parahynobius betfianus Venczel, 1999. - Romania. Pleistocene.

+ Parahynobius kordosi Venczel, 1999. - Hungary. Miocene.

Pseudohynobius shuichengensis Tian, Gu, Sun & Li, 1998. - China (Guizhou). - Comment: This

new nomen appears under three different spellings in the original publication: shuichengensis

(wice in p. 7, twice in p. 12, once in p. 13), xuichengensis (once in p. 11) and suichenensis (once in

p. 12). These spellings are “multiple original spellings”” according to the Code. Acting as first

revisers, we hereby choose the spelling shuichengensis as “correct original spelling” of this nomen.

Subfamilia Proromynopunar Fei & Ye, 2000

Protohynobius Fei & Ye, 2000a. - Type-species, by original designation: Prorohynobius puxiongensis

Fei & Ye, 2000. - China (Sichuan).

Protohynobius puxiongensis Fei & Ye, 2000, - China (Sichuan).

Source : MNHN, Paris

Dugois, CROMBIE & GLAW

Epifamilia + Karaurorpra Ivachnenko, 1978

Superfamilia + KarauroiprA Ivachnenko, 1978

Familia + Karauriar Ivachnenko, 1978

+ Kokartus Nessov, 1988. — Type-species, by original designation: + Kokartus honorarius Nessov,

1988. — Kirgiztan. Jurassic.

+ Kokartus honorarius Nessov, 1988. - Kirgiztan. Jurassic.

Epifamilia SaamaNDroipiA4 Goldfuss, 1820

Incertae sedis

Haldotriton Evans & Milner, 1996. - Type-species, by original designation: + Valdorriton gracilis

Evans & Milner, 1996. - Spain. Cretaceous.

+ Valdorriton gracilis Evans & Milner, 1996. - Spain. Cretaceous.

Superfamilia Amaysromaromnes Gray, 1850

Familia Auavsromarpar Gray, 1850

Familia Dicampropowrinar Tihen, 1958

+ Dicamptodon antiquus Naylor & Fox, 1993. - Canada (Alberta). Paleocene.

Superfamilia Aupsrumornra Gray, 1825

Familia Aupmrumpar Gray, 1825

+ Paleoamphiuma Rieppel & Grande, 1998. - Type-species, by original designation: + Paleoam-

Phiuma retradacrylum Rieppel & Grande, 1998, - USA (Wyoming). Eocene.

+ Paleoamphiuma tetradacrylum Rieppel & Grande, 1998. - USA (Wyoming). Eocene.

Familia PLETHODONTIDAE Gray, 1850

Subfamily HemibacryLuNar Hallowell, 1856 (1850)

Tribus BozrToGLo:

Hallowell, 1856

Batrachoseps diabolicus Jockusch, Wake & Yanev, 1998. - USA (California).

Batrachoseps gavilanensis Jockusch, Yanev & Wake, 2001. - USA (California).

Batrachoseps gregarius Jockusch, Wake & Yanev, 1998. - USA (California).

Batrachoseps incognitus Jockusch, Yanev & Wake, 2001. - USA (California).

Batrachoseps kawia Jockusch, Wake & Yanev, 1998. - USA (California).

Batrachoseps luciae Jockusch, Yanev & Wake, 2001. - USA (California).

Batrachoseps minor Jockusch, Yanev & Wake, 2001. - USA (California).

Source : MNHN, Paris

50 ALYTES 23 (1-2)

Barrachoseps regius Jockusch, Wake & Yanev, 1998. - USA (California).

Batrachoseps robustus Wake, Yanev & Hansen, 2002. - USA (California).

Bolitoglossa anthracina Brame, Savage, Wake & Hanken, 2001. - Panama.

Bolitoglossa decora MeCranie & Wilson, 1997. - Honduras.

Bolitoglossa guaramacalensis Schargel, Garcia-Pérez & Smith, 2002. - Venezuela.

Bolitoglossa hiemalis Lynch, 2001a. - Colombia.

Bolitoglossa lozanoi Acosta-Galvis & Restrepo, 2001. - Colombia.

Bolitoglossa mombachoensis Kôhler & McCranie, 1999. - Nicaragua.

Bolitoglossa oaxacensis Parra-Olea, Garcia-Paris & Wake, 2002. - Mexico.

Bolitoglossa spongai Barrio Amorôs & Fuentes Ramos, 2001, - Venezuela.

Bolitoglossa synoria McCranie & Kôhler, 1999a. - Honduras.

Boliroglossa zaporeca Parra-Olea, Garcia-Paris & Wake, 2002. - Mexico.

Cryptotriton Garcia-Paris & Wake, 2000. - Type-species, by original designation: Oedipus nasalis

Dunn, 1924. - Honduras.

Lineatriton orchileucos Brodie, Mendelson & Campbell, 2002. - Mexico.

Lineatriton orchimelas Brodie, Mendelson & Campbell, 2002. - Mexico.

Nototriton brodiei Campbell & Smith, 1998. - Guatemala.

Nototriton gamezi Garcia-Paris & Wake, 2000. - Costa Rica.

Nototriton lignicola MeCranie & Wilson, 1997. - Honduras.

Nototriton limnospectator MeCranie, Wilson & Polisar, 1998. - Honduras.

Nototriton monzoni Campbell & Smith, 1998. - Guatemala.

Nototriton saslaya Kôhler, 2002. - Nicaragua.

Nototriton stuarti Wake & Campbell, 2000. - Guatemala.

Notorriton wakei Campbell & Smith, 1998. - Guatemala.

Oedipina maritima Garcia-Paris & Wake, 2000. - Panama.

Oedipina savagei Garcia-Paris & Wake, 2000. - Costa Rica.

“Peudoeuryeea amuzga” Perez-Ramos & Saldana de la Riva, 2000. - Mexico. - Comment: Nomen-

claturally unavailable nomen, as having not been published on a “permanent support”, Nomen

made nomenclaturally available in PEREz-RAMOS & SALDANA DE LA Riva (2003).

Pseudoeurycea aquatica Wake & Campbell, 2001. - Mexico.

Pseudoeurycea Iynchi Parra-Olea, Papenfuss & Wake, 2001. - Mexico.

Pseudoeuryeea naucampatepeil Parra-Olea, Papenfuss & Wake, 2001. - Mexico.

Thorius grandis Hanken, Wake & Freeman, 1999, - Mexico.

Thorius infernalis Hanken, Wake & Freeman, 1999. - Mexico.

Thorius lunaris Hanken & Wake 1998. — Mexico.

Thorius magnipes Hanken & Wake 1998. - Mexico.

Thorius minydemus Hanken & Wake 1998. - Mexico.

Thorius munificus Hanken & Wake 1998. - Mexico.

Thorius omilteni Hanken, Wake & Freeman, 1999, - Mexico.

Thorius papaloae Hanken & Wake, 2001. - Mexico.

Thorius spilogaster Hanken & Wake 1998. - Mexico.

Tribus Srzserivr Cope, 1859

Blepsimolge Hillis, Chamberlain, Wilcox & Chippindale, 2001. - Type-species, by original designa-

tion: Eurycea nana Bishop, 1941. - USA (Texas). - Comment: Created as a subgenus of Eurycea

Gray, 1850.

Eurycea chisholmensis Chippindale, Price, Wiens & Hillis, 2000. - USA (Texas).

Eurycea naufragia Chippindale, Price, Wiens & Hillis, 2000. - USA (Texas).

Eurycea tonkatoae Chippindale, Price, Wiens & Hillis, 2000. - USA (Texas).

Source : MNHN, Paris

DuBois, CROMBIE & GLAW 51

Eurycea waterlooensis Hillis, Chamberlain, Wilcox & Chippindale, 2001. - USA (Texas).

Notiomolge Hillis, Chamberlain, Wilcox & Chippindale, 2001. - Type-species, by original designa-

tion: Eurycea neotenes Bishop & Wright, 1937. - USA (Texas). - Comment: Created as a

“division” of Eurycea Gray, 1850.

Paedomolge Hillis, Chamberlain, Wilcox & Chippindale, 2001. - Type-species, by original designa-

tion: Eurycea tonkatae Chippindale, Price, Wiens & Hillis, 2000. - USA (Texas). - Comment:

Created as a “section”? of Eurycea Gray, 1850.

Septentriomolge Hillis, Chamberlain, Wilcox & Chippindale, 2001. - Type-species, by original

designation: Eurycea chisholmensis Chippindale, Price, Wiens & Hillis, 2000. - USA (Texas).

Comment: Created as a subgenus of Eurycea Gray, 1850.

Subfamilia PLETHODONTINAE Gray, 1850

Tribus DESMOGNATHINI Gray, 1850

Aneïdes vagrans Wake & Jackman in JackMaN, 1998. - USA (California).

Desmognathus folkerisi Camp, Tilley, Austin & Marshall, 2002. - USA (Georgia).

Speleomantes imperialis sarrabusensis Lanza, Leo, Forti, Cimmaruta, Caputo & Nascetti, 2001. — Italy.

Tribus PLETHODONTINI Gray, 1850

Plethodon ainstorthi Lazell, 1998. - USA (Mississippi).

Plethodon amplus Highton & Peabody, 2000. - USA (North Carolina).

Plethodon cheoah Highton & Peabody, 2000. - USA (North Carolina).

Plerhodon electromorphus Highton, 1999. - USA (West Virginia).

Plethodon meridianus Highton & Peabody, 2000. - USA (North Carolina).

Plethodon montanus Highton & Peabody, 2000. - USA (Virginia).

Plethodon virginia Highton, 1999. - USA (Virginia).

Superfamilia Prorrorra Gray, 1825

Familia Prorsipar Gray, 1825

+ Mioproteus wezei Miynarski, Szyndlar, Estes & Sanchiz, 1984. - Poland. Pliocene.

Superfamilia S, NDROIDEA Goldfuss, 1820

Familia SaLAMANDRIDAE Goldfuss, 1820

+ Chelotriton pliocenicus Bailon, 1989. — France. Pliocene.

“Chioglossa lusitanica brevidigitata” Ferrand de Almeida, Ferrand de Almeida, Gonçalves, Sequeira,

Teixeira & Ferrand de Almeida, 2001. - Portugal. - Comment: Nomenclaturally unavailable

nomen, as having been published without type-specimen designation and without explicit

statement of the intention to establish a new taxon (Crocer & DuBois, 2004: 496).

Paramesotriton laoensis Stuart & Papenfuss, 2002. - Laos.

Trivurus karelinit arntzeni Litvinchuk, Borkin, Dzukié & Kalezié in LrrviNcHuk, BorkiN, DZUKté,

KaLezié, KHALTURIN & Rosanov, 1999. — Serbia.

Source : MNHN, Paris:

52 ALYTES 23 (1-2)

Tlototriton asperrimus wenxianensis Feï, Ye & Yang, 1984. - China (Gansu).

Tlototriton hainanensis Fei, Ye & Yang, 1984. - China (Hainan). - Comment: Authorship of nomen

wrongly credited to “Fei & Yang” by FROST (1985: 617) and DUELLMAN (1993: 310).

Epifamilia Sirevornia Gray, 1825

Superfamilia Sreevorra Gray, 1825

Familia Srenipar Gray, 1825

+ Kababisha Evans, Milner & Werner, 1996. - Type-species, by original designation: + Kababisha

humarensis Evans, Milner & Werner, 1996. - Sudan. Cretaceous.

+ Kababisha humarensis Evans, Milner & Werner, 1996. - Sudan. Cretaceous.

+ Kababisha sudanensis Evans, Milner & Werner, 1996. - Sudan. Cretaceous.

+ Noterpeton Rage, Marshall & Gayet, 1993. - Type-species, by original designation: + Norerperon

bolivianum Rage, Marshall & Gayet, 1993. - Bolivia. Cretaceous.

+ Norerpeton bolivianum Rage, Marshall & Gayet, 1993. - Bolivia. Cretaccous.

Superordo Gymnormowa Rafinesque-Schmaltz, 1814

Ordo Grmnormioxa Rafinesque-Schmaltz, 1814

Incertae sedis

+ Rubricacaecilia Evans & Sigogneau-Russell, 2001. - Type-species, by original designation:

+ Rubricacaccilia monbaroni Evans & Sigogneau-Russell, 2001. - Morocco. Cretaceous.

+ Rubricacaecilia monbaroni Evans & Sigogneau-Russell, 2001. - Morocco. Cretaceous.

Epifamilia Carosromma Rafinesque-Schmaltz, 1814

Superfamilia CarcirromA Rafinesque-Schmaltz, 1814

Familia Carcunoar Rafinesque-Schmaltz, 1814

Boulengerula fischeri Nussbaum & Hinkel in FiscHer & Hinkez, 1994 (nec Boulengerula fischeri

Nussbaum & Hinkel, 1994]. - Rwanda. - Comment: See LôTTERS (2003).

Gegeneophis krishni Pillai & Ravichandran, 1999. — India (Karnataka).

Familia Icuravoraupas Taylor, 1968

Ichthyophis garoensis Pillai & Ravichandran, 1999. - India (Meghalaya).

Ichthyophis husaini Pillai & Ravichandran, 1999. — India (Meghalaya).

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 53

Familia Scozrcomorrmupar Taylor, 1969

Crotaphatrema tchabalmbaboensis Lawson, 2000. - Cameroon.

Familia TÿPHLONECTIDAE Taylor, 1968

Atretochoana Nussbaum & Wilkinson, 1995. - Type-species, by original designation: Tiphlonectes

iselti Taylor, 1968. - “South America”.

Pseudotyphlonectes Lescure, Renous & Gasc, 1986. — Type-species, by original designation:

Caecilia natans Fischer, 1879. - Colombia.

Familia UraroryPaLipaE Nussbaum, 1979

Uraeoryphlus interruptus Pillai & Ravichandran, 1999. - India (Kerala).

Epifamilia Eocarcuzaoma Jenkins & Walsh, 1993

Superfamilia + Eocarcaomnea Jenkins & Walsh, 1993

Familia + Bocascuzupar Jenkins & Walsh, 1993

+ Eocaecilia Jenkins & Walsh, 1993. - Type-species, by original designation: + Focaecilia micropodia

Jenkins & Walsh, 1993. - USA (Arizona). Jurassic.

+ Eocaccilia micropodia Jenkins & Walsh, 1993. - USA (Arizona). Jurassic.

ACKNOWLEDGMENTS

For helpful comments, suggestions and information that allowed 10 improve this paper, we are

grateful to Lauren E. Brown, Jean-Claude Rage, Borja Sanchiz, Miguel Vences and David B. Wake.

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Doria, G., SALviDIo, S. & TavaNo, M. L., 2001. - Description of Amolops (Huia) modiglianii, new species from

Sumatra (Amphibia, Anura, Ranidac). Doriana, 1999”, 7 (317): 1-9. [Printing finished in February 2001

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Drewess, R. C & PERRET, J.-L., 2000. — À new species of giant, montane Phrynobarrachus (Anura: Ranidac) from the

central mountains of Kenya. Proceedings of the California Academy of Sciences, 32 (5): 55-64.

Dusois, A.; 2000 [13 July].- Synonymies and related lists in z00logy: general proposals, with examples in herpetology.

Dumerilia; 4 (2): 33-98. [Exact publication date obtained directly from publisher].

2002 (24 January]. - The taxonomic status of Seutiger mokokchungensis Das & Chanda, 2000 (Amphibia, Anura).

Amphibia-Reprilia, “2001”, 22 (4): 499-506. (Exact publication date: Amphibia-Reptilia, 2002, 23 (1): inside

front cover].

— 2003. - True frogs (Ranidae). In: M. HurcHns, W. E. DUELLMAN & N. ScHLAGER (ed.), Grzimek's animal life

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— 2004. Developmental pathway, speciation and supraspecific taxonomy in amphibians. 1.Why are there so many

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— 20056. - Proposed Rules for the incorporation of nomina of higher-ranked zoological taxa in the International

Code of Zoological Nomenclature. 1. Some general questions, concepts and terms of biological nomenclature.

Zoosystema, 1 (2): 365-426.

—— 200%. - Proposed Rules for the incorporation of nomina of higher-ranked zoological taxa in the International

Code of Zoological Nomenclature. 2. The proposed Rules and their rationale. Zoosystema, in press.

Dusors, À.; Marsut, M. & Our, À, 2001 [31 May]. À replacement name for Rana (Paa) rara Dubois & Matsui,

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Dusors, À. & OLer, À., 1995. — Frogs of the subgenus Pelophylax (Amphibia, Anura, genus Rana): a catalogue of

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—— 1998 [7 October]. - À new species of Leprobrachium (Vibrissaphora) from northern Vietnam, with a review of the

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_— 1999. — Asian and Oriental toads of the Bufo melanostictus, Bufo scaber and Bufo stejnegeri groups (Amphibia,

Anura): a list of available and valid names and redescription of some name-bearing types. Journal of South

Asian natural History, 4 (2): 133-180.

_— 2001 [20 December]. — A new genus for an aquatic ranid (Amphibia, Anura) from Sri Lanka. Alyres, 19 (2-4):

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Dusors, A., OHLER, À. & Buju, S. D., 2001 [20 December]. - A new genus and species of Ranidae (Amphibia, Anura)

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DueLLMaN, W. E., 1993. - Amphibian species of the world: additions and corrections. University of Kansas Museum of

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DusLLMan, W. E., 2000 [8 September]. — Leptodactylid frogs of the genus Phrynopus in northern Peru with

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DueLmaN, W. E., Leur, E. & AGUILAR, C.; 2001. - À new species of marsupial frog (Anura: Hylidae: Gasrrotheca)

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Durra, S. K. & Ray, P. 2000. - Microhyla sholigari, a new species of microhylid frog (Anura: Microhylidae) from

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Esrrapa, À. R. & ALONSO, R., 1997. - Nueva especie del grupo lmbatus (Leptodactylidae: Eleutherodacrylus) de la

region oriental de Cuba. Caribbean Journal of Science, 33 (1-2): 41-44.

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58 ALYTES 23 (1-2)

Esrrapa, À. R. & HEDGES, S. B., 1997a. - Nueva especie de Eleutherodactylus (Anura: Leptodactylidae) del macizo

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Evans, S. E. & SiGoGnrau-RussELL, D., 2001. - A stem-group caecilian (Lissamphibia: Gymnophiona) from the

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Ft, L., JIaNG, J., Ye, C. & CHENG, S., 1996. - Electrophoresis analyses of crystalline [sic] lens protein of twenty-one

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Far, L. & Ye, C, 20004. - À new hynobiid subfamily with a new genus and new species of Hynobiidae from west China

(Amphibia: Caudata). Culrum herperologica sinica, 8: 64-70. [In Chinese].

—— 20008. - À new subspecies of Hyla simplex Boettger from China (Amphibia: Hylidae). Culrun herpetologica sinica,

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— 2001 a. - The colour handbook of the amphibians of Sichuan. Chengdu, Chengdu Institute of Biology: [i-xv] + 1-263.

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20016. - A new species of the genus Pa from southeast Xizang, China (Amphibia: Ranidae). Acta zoologica sinica,

47 (2): 476-478. [In Chinese].

Fu, L,, YE, C. & CHEN, H., 1992. - A new pelobatid toad of genus Orolalax (Amphibia, Anura) from Yunnan-

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13 (1): 5-12. [In Chinese].

—— 1997. -Taxonomic studies on the genus Liurana of China including discriprions [sic] of a new species (Amphibia:

Ranidae). Cultun herpetologica sinica, 6-7: 75-80. [In Chinese].

Fu, L., YE, C., HUANG, Y. & CHEN, X., 1999. - Taxonomic studies on Bufo viridis from west China (Amphibia:

Anura). Zoological Research, 20 (4): 294-300. [In Chinese].

Fe, L, Ye, C. & JiaNG, J., 2000. - À new genus of subfamily Amolopinae — Pseudoamolops, and its relationship to

related genera. Acra 2oologica simica, 46 (1): 19-26. [in Chinese]. [Received on 16 May 2000 in the Paris

Museum Central Library]

VE, C., ANG, J. & CHEN, S., 1999. - Description of a new Scuriger from Mt. Gongga in Sichuan Province of

China. Zoological Research, 20 (3): 172-177. [In Chinese].

Fi, L., YE, C. & Li, C., 1999. - Description of a new pelobatid toad of genus Orcolalax - Oreolalax nanjiangensis from

Daba mountain region of China. Acta zootaxonomica sinica, 24 (1): 107-113. [In Chinese].

2001a.- Descriptions of two new species of the genus Odorrana in China (Anura: Ranidae). Acta s00raxonomica

simicas 26 (1): 108-114. [In Chinese].

20016. - Taxonomic studies of Odorrana versabilis in China. II. Descriptions of wo new species (Amphibia:

Ranidae). Acta zootaxonomica simica, 26 (4): 601-607. [In Chinese].

Fe, L, YE, C. &Li,S., 1992. -Two new species of pelobatid toads of genus Leprolalax from Yunnan province of China

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Fm, L, VE, C. & YANG, R 1984. - À new species and a new subspecies of the genus Tÿlororriton (Caudara:

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FeRNicoLA, J. C., 2001. - Una nueva especie de Ceratophrys (Anura, Leptodactylidae) en el Neogeneo de la provincial

de Buenos Aires, Argentina. Ameghiniana, 38 (4): 385-391.

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ForMas, J. R., Cuzvas, C. C. & Briva, L. M., 2002. À new species of Alsodes (Anura: Leptodactylidae) from Cerro

Mirador, Cordillera Pelada, southern Chile. Proceedings of the biological Society of Washington, 115 (4): 708-719.

Formas, J. R,, Cuevas, C. & NUREz, J., 1998. - À new species of Alkodes (Amphibia: Anura: Leptodactylidae) from

southern Chile. Proceedings of the biological Sociery of Washington, 1 (3): 521-530.

Formas, J. R., NorrHLaNn, L., CaperiLLo, J., NUNE7, J. J., Cuevas, C. C. & BRIEVA, L. M. 1999. - Télmatobius dankoi,

una nueva especie de rana acuatica del norte de Chile (Leptodactylidae). Revista chilena de Historia natural, T2

G): 427-445.

Formas, J. R., Uepa, C., Curvas, C. & NUNEz, J., 1998. — Alsodes ausrralis, a new species of leptodactylid frog from

the temperate Nothofagus forest of southern Chile and Argentina. Studies on neotropical Fauna and Environment,

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Fox, R. C. & NavLoR, B. G., 1982. - A reconsideration of the relationships of the fossil amphibian Albanerperon.

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Frosr, D. R., (ed.), 1985. - Amphibian species of the world. Lawrence, Allen Press & Assoc. Syst. Coll.

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Gao, K.-Q,., CHENG, Z.-W. & XU, X., 1998. - First report of a Mesozoic urodele from China. Chinese Geology, 1 (248):

40-41. [In Chinese].

Gao, K.-Q. & SuuBiN, N. H., 2001. - Late Jurassic salamanders from northern China. Nature, 410 (6828): 574-577.

Gao, K.-Q. & WanG,Y., 2001. - Mesozoic anurans from Liaoning Province, China, and phylogenetic relationships of

archaeobatrachian anuran clades. Journal of vertébrate Paleomology, 21 (3): 460-476.

Garcia-Paris, M. & Marrinez-SoLANO, L., 2001. — Nuevo estatus taxonômico para las poblaciones ibero-

mediterräneas de Alyres obsterricans (Anura: Discoglossidae). Revista española de Herpetologia, 15: 99-113.

Garcia-Paris, M. & Wake, D. B., 2000. - Molecular phylogenetic analysis of relationships of the tropical salamander

genera Ocdipina and Nororriton, with descriptions of a new genus and three species. Copeia, 2000 (1): 42-70.

GarKER, J. D., 1999. - New albanerpentotid amphibians from the Albian to Coniacion of Utah, USA - bridging the

gap. Journal of Vértebrate Paleontology, 19 (4): 632-638.

—— 2000. = Albanerpetontid amphibians from the upper Cretaceous (Campanian and Maastrichtian) of North

America. Geodiversitas, 22 (3): 349-388.

GrrtaGH,J. & WiLLI, J., 2002. — À new species of frog, genus Sooglossus (Anura, Sooglossidae) from Silhouette island,

Seychelles. Amphibia-Reprilia, 23 (4): 445-458.

Grarerra, À. À. & AGuIAR, O. Jr., 1998. — À new species of Megaelosia from Mantiqueira range, southeastern Brazil.

Journal of Herpetology, 32 (1): 80-83.

GrarerTa, À. A. BERNARDE P. S. & KokuBuM M. N. DE C., 2000. — A new species of Proceratophrys (Anura:

Leptodactylidae) from the Amazon rain forest. Journal of Herperology, 34 (2): 173-178.

GrarerTa, À. À. & Sawaya, R. J.; 1998. - Second species of Psyllophryne (Anura: Brachycephalidae). Copeia, 1998 (4):

985-087.

GLaw, F, KôHLer, J., LôrreRs, S. & VENCEs, M., 1998. - Vorläufige Liste und Bibliographie neubeschriebener

Amphibienarten und -unterarten von 1993 bis 1997. Elaphe, 6 (1): i-xxiv.

Graw, F. & VENCES, M., 1994 [Mid December]. - À fieldguide 10 the amphibians and repiles of Madagascar. Second

edition, including mammals and freshroater fish. Kôin, Vences & Glaw: 1-480, 48 pl. [Publication date according

10 F. Glaw].

2000. À new species of Mantidactylus from northeastern Madagascar with resurrection of Manridactylus blanci

(Guibé, 1974) (Amphibia, Anura, Ranidae). Spixiana, 23 (1): 71-83.

2001. - To new sibling species of Manridactylus cornutus from Madagascar (Amphibia, Anura, Ranidac).

Spixiana, 24 (2): 177-190.

2002a [30 April]. - À new sibling species of the anuran subgenus Blommersia from Madagascar (Amphibia:

Mantellidae: Mantidacrylus) and its molecular phylogenetic relationships. The herperological Journal, 12 (1):

11-20. [Exact publication date: The herpetological Journal, 2002, 12 (2): back cover]

20028. - À new cryptic treefrog species of the Boophis luteus group from Madagascar: bioacoustic and genetic

evidence (Amphibia, Anura, Mantellidae). Spixiana, 25 (2): 173-181.

2002 [15 October]. - À new cryptic frog species of the Mantidactylus boulengeri group with a divergent vocal sac

structure. Amphibia-Reprilia, 23 (3): 293-304, [Exact publication date: Amphibia-Reprlia, 2002, 23 (4): inside

front cover].

20024. - À new species of Mantidactylus (Anura: Mantellidae) from Andasibe in eastern Madagascar. Journal of

Herperology, 36 (3): 372-378.

GLaw, F., VENCES, M., ANDREONE, F. & VALLAN, D., 2001. - Revision of the Boophis majoré group (Amphibia:

Mantellidae) from Madagascar, with descriptions of five new species. Zoological Journal of the Linnean Society,

133 (4): 495-529

GLaw, F., VeNcEs, M. & Bôume, W., 1998. - Systematic revision of the genus Aglyprodacrylus Boulenger, 1919

(Anura: Ranidae) and analysis of its phylogenetic relationships with other ranoid genera from Madagascar

(Tomopterna, Boophis, Mantidacrylus and Mantella). Journal of soological Systematies and evolurionary Research,

36 (1-2): 17-37

GLaw, F., VeNCEs, M. & GossManx, V., 2000. — À new species of Mantidacrylus (subgenus Guibemantis) from

Madagascar, with a comparative survey of internal femoral gland structure in the genus (Amphibia: Ranidae:

Mantellinae). Journal of natural History, 34 (1): 1135-1154.

GornaM, S. W., 1974. - Checklisr of seorld amphibians up t0 January 1, 1970. Saint-John, The New Brunswick Museum:

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60 ALYTES 23 (1-2)

GorzuLa, S. & Searts, C.; 2000. - Una nueve especie del género Osteocephalus (Anura: Hylidae) de la Gran Sabana,

Venezuela. Acta biologica venezuelica, “1996”, 16 (4): 19-22.

Grant, T,, 1998. -Una nueva especie de Colosterhus del grupo eduardsi de Colombia. Revista de la Academia colombiana

de Ciencias exactas, fisicas y naturales, 22 (84): 423-428.

——--- 2000. - Una nueva especie de Rhamphophryme (Anura: Bufonidae) de la Cordillera Central de Colombia. Revista

de la Academia colombiana de Ciencias exactas fisicas y naturales, “1999”, 23 (Suplemento especial): 287-292.

{Printing finished in February 2000 according to last page of fascicle].

Gran, T. & ARDILA-Rosavo, M. C.; 2002 [23 May]. - À new species of Colosterhus (Anura: Dendrobatidae) from the

eastern slopes of the Cordillera Oriental of Colombia. Herpetologica, 58 (2): 252-260. [Exact publication date:

Herpetologica, 2002, 58 (3): 393].

Grant, T. & Casrro, F., 1998. - The cloud forest Colosterhus (Anura, Dendrobatidae) of a region of the Cordillera

Occidental of Colombia. Journal of Herpetology, 32 (3): 378-392.

Granr, T., Humpnrey, E. C. & Mvers, C. W., 1997. - The median lingual process of frogs: a bizarre character of Old

World ranoids discovered in South American dendrobatids. American Museum Novitates, 3212: 1-40.

Grant, T. & RoDRiGUEz, L. O., 2001. - Tiwo new species of frogs of the genus Colostethus (Dendrobatidae) from Peru

and a redescription of C. rrilineatus (Boulenger, 1883). American Museum Novitates, 3355: 1-24.

Gv, H., 1992. - À new species of Hynobius - Hynobius amjiensis. In:Y. QuaN, E. ZHAO & K. ZHAO. (ed:), Animal science

research, À volume issued 10 celebrate the 90” birthday of Professor Mang-ven L.Y Chang, Beïjing, China Forestry

Press, “1991”: 39-43. [In Chinese]. [Publication date: ZHaO & ADLER, 1993: 372].

Gui, Y. M., 1991. - Paleocene salamanders from southern Mongolia. Paleontologicheskit Zhurnal, 191 (1): 96-106.

-— 1999. - Gobiatids (Anura) from the Upper Cretaceous locality Khermeen-Tsav (Gobi Desert, Mongolia).

Paleontologicheskit Zhurnal, 1991 (1): 76-87.

Günruer, R, 1985. - Ordnung Anura, Froschlurche. In:W.-E. ENGELMANN, J. FITZSCHE, R. GÜNTHER & F. J. Onsr,

Lurche und Kriechiere Europas, Leipzig & Radebeul, Neumann: 113-184.

—— 1999. - Morphological and bioacoustic characteristics of frogs of the genus Plarymantis (Amphibia, Ranidae) in

Irian Jaya, with descriptions of new species. Mitteilungen aus dem Museum für Naturkunde in Berlin, Zoologische

Reihe, 75 (2): 317-335.

— 2000. - Albericus laurini species nova, the first record of the genus Albericus (Anura, Microhylidae) from the west

of New Guinea. Mireilungen aus dem Museum für Naturkunde in Berlin, Zoologische Reine, 16 (2): 167-174.

—- 2001. - The Papuan frog genus Hylophorbus (Anura: Microhylidae) is not monospecific: description of six new

species. Russian Journal of Herpetology, 8 (2): 81-104.

—— 20024. - Westernmost records of the Papuan frog genus Copiula with descriptions of two new species (Amphibia:

Anura: Microhylidae). Faunistische Abhandlungen, Staatliches Museum für Tierkunde Dresden, 23 (2): 35-58.

— 2002. — Beschreibung einer neuen Copiula-Art (Amphibia, Anura, Microhylidae) von der Insel Yapen im

Nordwesten von Papua, Indonesien. Zoologische Abhandlungen, Staatliches Museum für Tierkunde Dresden, 52:

77-86.

Günruer, R. & RICHARDS, S., 2000. - A new species from the Livoria gracilenta group from Irian Jaya (Anura: Hylidac).

Herperozoa, 13 (1-2): 21-43.

Güxruer, R. RICHARDS, S. & Iskanpar, D., 2001. - Two new species of the genus Orcophryne from Irian Jaya,

Indonesia (Amphibia, Anura, Microhylidae). Spixiana, 24 (3): 257-214,

Hanken, J. & Wake, D. B., 1998. — Biology of tiny animals: systematics of the minute salamanders (Thorius:

Plethodontidae) from Veracruz and Puebla, México, with descriptions of five new species. Copeia, 1998 (2):

312-345.

2001 [10 December]. — À seventh species of minute salamander (Thorius: Plethodontidae) from the Sierra de

Jurez, México. Herpetologica, 57 (4): 515-523. [Exact publication date: Herpetologica, 2002, 58 (1): 138].

Hanken, J, Wake, D. B. & FREEMAN, H. L., 1999. - Three new species of minute salamanders (Thorius:

Plethodontidae) from Guerrero, México, including the report of a novel dental polymorphism in Urodeles.

Copeia, 1999 (4): 917-931.

Haroy,J. D.Jr., 2001. - À new frog of the genus Elewherodactylus from the island of Tobago, West Indies. Bulletin of the

“Maryland herpetological Society, 37 (1): 25-31.

Harvey, M. B. & P. ERGUETA S. (1998. - À new speci

Bolivian Andes. Journal of Herpetology, 32 (4): 536-540

Harvey, M. B., PEMBERTON, À. J. & SMITH, E. N., 2002. - New and poorly known parachuting frogs (Rhacophoridae:

Rhacophorus) from Sumatra and Java. Herperological Monographs, 16: 46-92.

He, X., 1999. - À new species of the family Rhacophoridae: Polypedates puerensis. Sichuan Journal of Zoology, 18 (3):

99-100.

HenriGi, A. C., 1998. — À new pipoid anuran from the Late Jurassic Morrison Formation at Dinosaur National

Monument, Utah, Journal of vertebrate Paleontology, 18 (2): 321-332.

Henrici, A. C. & Bâez, A. M, 2001. - First occurrence of Xenopus (Aura: Pipidae) on the Arabian peninsula: a new

species from the upper Oligocene of Yemen. Journal of Paleontology, 75 (4): 870-882.

Hver, W. R., 1999. - À new genus and species of frog from Bahia, Brazil (Amphibia: Anura: Leptodactylidae) with

comments on the z0ogeography of the Brazilian campos rupestres. Proceedings of the biological Sociery of

Washingron, 12 (1): 19-39.

Hever, W. R. & Muoz, À. M. 1999. - Validation of Eleutherodacrylus crepitans Bokermann, 1965, notes on the types

and type locality of Télarrema heterodacrylum Miranda-Ribeiro, 1937, and description of a new species Of

Eleutherodactylus from Mato Grosso, Brazil (Amphibia: Anura: Leptodactylidae). Proceedings of the biological

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Source : MNHN, Paris

62 ALYTES 23 (1-2)

KônLer, G. & MCCRANIE, J. R, 1999. À new species of salamander from Volcän Mombacho, Nicaragua, formerly

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—— 20008. - New species of Eleutherodacrylus (Anura: Leptodactylidae) from cloud forest of Bolivia. Copeia, 2000 (2):

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KôuLer, J. & Lôrrens, S., 1999. - New species of the Æleutherodacrylus unistrigatus group (Amphibia: Anura:

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—— 20016. - Description of a small tree frog, genus Hyla (Anura: Hylidae), from humid Andean slopes of Bolivia.

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Lararor, À, Muray, R. W., ORLOv, N. L. & Ho, C. T., 19986 [12 August]. -Twvo new species of Leprolalax (Anura:

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Leur, E., KôuLER, G., AGUILAR, C. & PONCE, E., 2001. - New species of Bufo (Anura: Bufonidae) from central Peru,

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Leur, E., KôHLER, G. & PONCE, E., 2000. - À new species of Phrynopus from Peru (Amphibia, Anura, Leptodacty-

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Leur, E., RoDRiGuEz, D. & CRpova, J. H., 2002. - À new species of Phrynopus (Amphibia, Anura, Leptodactylidac)

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Lrrvincnux, S. N., BorkiN, L. J., Dzukié, G., Kaze216, M. L., KHaLTuRIN, M. D. & Rosanov, J. M., 1999. —

Taxonomic status of Triturus karelinii on the Balkans, with some comments about other crested new taxa.

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Liv, W. & YANG, D., 2000 [1 June]. - À new species of Amolops (Anura: Ranidae) from Yunnan, China, with a

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Liv, W., YanG, D., FerraRts, C. & MaTsut, M.; 2000. - Amolops bellulus: a new species of stream-breading frog from

western Yunnan, China (Anura: Ranidae). Copeia, 2000 (2): 536-541.

Lôrrers, S., 2003. - Boulengerula fischeri Nussbaum and Hinkel, 1994 is an objective junior synonym and a homonym

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Lôrrers, S. & HENzL, M. J,, 2000. - À new species of Atelopus (Anura: Bufonidae) from the Serrania de Sira,

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Lôrrens, $., JUNGrER, K.-H. & WIDMER, À., 2000. - À new genus of aposematic poison frog (Amphibia: Anura:

Dendrobatidae) from the upper Amazon Basin, with notes on its reproductive biology and tadpole morphol-

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Lôrrers, S. & KôHLER, J., 2000. - À new toad of the Bufo ryphonius complex from humid montane forests of Bolivia

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Lu, Y.-Y. & Li, P-P, 2002. - À new wood-frog of the genus Rana in Mt. Kunyu, Shandong Province, China

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Lynn, J. D. 1998a. - New species of Eleutherodactylus from the Cordillera Occidental of western Colombia with a

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—— 1998. — À new frog (genus Eleutherodactylus) from cloud forests of southern Boyacä. Revista de la Academia

colombiana de Ciencias exactas, fisicas y naturales, 22 (84): 429-432.

— 2000. - À new species of frog, genus Eleutherodacrylus (Leptodactylidae), from the Sabana de Bogotä. Revista de

la Academia colombiana de Ciencias exactas, fisicas y naturales, 24 (92): 435-439.

-— 20014. À small amphibian fauna from a previously unexplored paramo of the Cordillera Occidental in western

Colombia. Journal of Herpetology, 35 (2): 226-231.

— 20016. - Three new rainfrogs of the Eleutherodactylus diastema group from Colombia and Panama. Revista de la

Academia colombiana de Ciencias exactas, fisicas y naturales, 25 (95): 287-297.

-— 2002. À new species of the genus Osreocephalus (Hylidae: Anura) from the western Amazon. Revista de la

Academia colombiana de Ciencias exactas, fisicas y naturales, 26 (98): 289-292.

Lynen, J. D. & ARDILA Rorayo, M. C., 1999. -The Eleutherodactylus of the taeniatus complex in western Colombia:

taxonomy and distribution. Revista de la Academia colombiana de Ciencias exactas, fisicas y naturales, 23 (89):

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Lyncn, J. D. & RUEDA-ALMONACID, J.V,, 1998. - Additional new species of frogs (genus Eleutherodactylus) from cloud

forests of eastern Departamento de Caldas, Colombia. Revista de la Academia colombiana de Ciencias exactas,

Jisicas y naturales, 22 (83): 287-298.

— 19986, — New frogs of the genus Eleutherodactylus from the eastern flank of the northern Cordillera Central of

Colombia. Revista de la Academia colombiana de Ciencias exactas, fisicas y naturales, 22 (85): 561-570.

= 1999, - New species of frogs from low and moderate elevations from the Caldas transect of the eastern flank of

the Cordillera Central. Revista de la Academia colombiana de Ciencias exactas,fisicas y naturales, 23 (ST): 307-314.

Lyxen, J. D. & Suarez-MayorGa, À. M., 2000. - À new frog (Eleutherodacrylus: Leptodactylidae) from the southern

part of the Cordillera Oriental of Colombia. Revista de la Academia colombiana de Ciencias exactas, fisicas ÿ

naturales, 24 (91): 289-293.

MacCuLLocu, R. D. & Laruror À., 2002 [6 August}. - Exceptional diversity of Srefania (Anura: Hylidae) on mount

Ayanganna, Guyana: three new species and new distributional records. Herperologica, 58 (3): 321-346. (Exact

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Source : MNHN, Paris:

64 ALYTES 23 (1-2)

Manoxy, M., KNOWLES, R., FOSTER, R. & DONKELLAN, $., 2001. - Systematics of the Litoria citropa (Anura: Hylidae)

complex in northern New South Wales and southern Queensland, Australia, with the description of a new

species. Records of the Australian Museum, 53: 31-48.

MANAMENDRA-ARACHCHI, K. & PETHIYAGODA, R., 1998. A synopsis ofthe Sri Lankan Bufonidae (Amphibia: Anura),

with description of two new species. Journal of South Asian natural History, 3 (2): 213-248.

—- 20014. - Ramanella nagaoï, a new tree-hole frog (Microhylidae) from southern Sri Lanka. Journal of South Asian

natural History, 5 (2): 121-133.

——— 20016. — Polypedates fastigo, a new tree frog (Ranidae: Rhacophorinac) from Sri Lanka. Journal of South Asian

natural History, 5 (2): 191-199.

ManrHEy, U. & ST#1OF, C., 1998. Rhacophorus cyanopunctatus sp. n. (Anura: Rhacophoridae), ein neuer Flugfrosch

von der Malaïschen Halbinsel, Sumatra und Borneo. Sauria, 20 (3): 37-42.

MarTiNo, A. L. & SscH, U., 2002. - Speciation by polyploidy in Odontophrynus americanus. Journal of Zoology,

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Marsut, M., NaBHITABHATA, J. & PANHA, $., 1998 [29 December]. - À new Ansonia from northern Thailand (Anura:

Bufonidae). Herpetologica, 54 (4): 448-454. [Exact publication date: Herperologica, 1999, 58 (1): 129].

1999. - On Leprobrachium from Thailand with a description of a new species (Anura: Pelobatidae). Japanese

Journal of Herpetology, 18 (1): 19-29.

MGCRANIE, J. R. & KôHLER, G., 1999a. - À new species of salamander of the Bolitoglossa dunni group from Cerro El

Pital, Honduras and El Salvador (Amphibia, Caudata, Plethodontidae). Senckenbergiana biologica, 18 (1-2):

225-229.

— 19996. A new species ofrainfrog ofthe Eleutherodactylus gollmeri group from western Honduras. Senckenbergiana

biologica;, 79 (1): 83-87.

MGCRANIE, J. R. & WILSON, L. D., 1997. Two new species ofsalamanders (Caudata: Plethodontidae) of the genera

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1998. Specific status of the Honduran frogs formerly referred to Plectrohyla teuchestes. Journal of Herpetology, 32

(1): 96-101.

—— 1999. - Description of a new species of Pkctrohyla from Cerro Celaque, Honduras, formerly referred to

Plectrohyla glandulosa (Amphibia, Anura, Hylidae). Senchenbergiana biologica, 78 (1-2): 231-236.

——— 19996. "Two new species of the Eleutherodacrylus rugulosus group (Amphibia: Anura: Leptodactylidae) from

Honduras. Proceedings of the biological Society of Washington, 112 (3): 515-522.

-— 2000. - À new species of high-crested toad of the Bufo valiceps group from north-central Honduras. Journal of

Herperology, 34 (1): 21-31.

MCCRANIE, J. R., WiLsON, L. D. & PoLIsaR, J., 1998 [29 December]. - Another new montane salamander (Amphibia:

Caudata: Plethodontidae) from Parque Nacional Santa Barbara, Honduras. Herpetologica, 54 (4): 455-461.

[Exact publication date: Herpetologica, 1999, 58 (1): 129].

McDonarp, K. R 1997. - À new stream-dwelling Litoria from the Melville Range, Australia. Memoirs of the

Queensland Museum, 42 (1): 307-309.

McGowan, G. & Evans, S.E., 1995. - Albanerpetontid amphibians from the Cretaceous of Spain. Nature, 373 (6510):

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MExDELsON, J. R., III & CamPBLz, J. A, 1999 [9 March]. - The taxonomic status of populations referred to Hyla

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Mexpes CasraNHo, L. & HappaD, C. F. B., 2000 [4 August]. - New species of Eleutherodactylus (Amphibia:

Leptodactylidae) from Guaraqueçaba, Atlantic Forest of Brazil. Copeia, 2000 (3): 777-181. (Publication date

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Mexziss, J.L., 1999. — À study of Albericus (Anura: Microhylidae) of New Guinea. Australian Journal of Zoology, 47 (4):

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MERCADAL DE BaRklO, L.T. & A. BARRIO (1994. — Reconsideraciôn del género Pseudopaludicola de Argentina y

descripcién de dos nuevas especies P mirandae ÿ P riopiedadensis (Amphibia, Anura). Revista del Museo

argentino de Ciencias naturales “Bernardino Rivadavia”, 16 (6): 65-80.

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357-368. [Exact publication date: Amphibia-Reprilia, 20 (1): inside front cover].

Vasupevax, K. & Durra, S. K., 2000. - À new species of Rhacophorus (Anura: Rhacophoridae) from the Western

Ghats, India. Hamadryad, 25 (1): 21-28.

Vers, M., Kosucn, ]., OuLer, À. & DuBois,À.,2001 [31 May]. Systematics of Fojervarya limmocharis (Gravenhorst,

1829) (Amphibia, Anura, Ranidae) and related species. 2. Morphological and molecular variation in frogs from

the Greater Sunda Islands (Sumatra, Java, Borneo) with the definition of two species. Alyres, 19 (1): 5

{Exact publication date obtained directly from publisher].

VéLez-Ronrquez, C. M. & Rurz-CarRANZA, P. M; 1997. - Una nueva especie de Arelopus (Amphibia: Anura:

Bufonidae) de la Cordillera Central, Colombia. Revista de la Academia colombiana de Ciencias exactas, fisicas y

naturales, 21 (81): 555-563.

2002 [24 October]. - À new species of Bufo (Anura: Bufonidae) from Colombia. Herperologica, 58 (4): 453-462.

[Exact publication date: Herperologica, 2003, 59 (1): 143]

Vexces, M. ANDREONE, F., GLAW, F. & MaTr1OL1, F, 2002. - New dwarf species of Manridactylus from northwestern

Madagascar (Anura: Mantellidae). Copeia, 2002 (4): 1057-1062.

Vaxces, M. & GLAW, F., 2001a. - When molecules claim for taxonomic changes: new proposals on the classification

of Old World treefrogs. Spixiana, 24 (1): 85-92

20016 [20 December]. - Systematic review and molecular phylogenetic relationships of the direct developing

Malagasy anurans of the Mantidacrylus asper group (Amphibia, Mantellidae). Alyes, 19 (2-4): 107-139. [Exact

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2002. - Tivo new treefrogs of the Boophis rappiodes group from eastern Madagascar (Amphibia Mantellidac).

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Vences, M., GLaW F, ANDREONE, F., JESU, R. & SCHIMMENTI, G., 2002. - Systematic re

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Vexces, M., GLaw, F. & BôHME, W, 1999 [4 November]. - À review of the genus

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ion of the enigmatic

position within the

antella (Anura, Ranidae,

72. [Exact

Source : MNHN, Paris

Dugois, CROMBIE & GLAW 69

Veces, M., GLaw, F, Kosucn, J., Das, IL. & Verrs, M., 2000. - Polyphyly of Tomopterna (Amphibia: Ranidae) based

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Wake, D. B, & CAMPBELL, J. À; 2000. - À new species of diminutive salamander (Amphibia: Caudata: Plethodontidae:

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— 2001 [10 December]. - An aquatic plethodontid salamander from Oaxaca, Mexico. Herpetologica, 57 (4): 509-514.

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Wake, D. B., YANEv, K. P. & HANSEN, R. W,, 2002. - New species ofslender salamander, genus Batrachoseps, from the

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WaxG, Y., 2000. - À new salamander (Amphibia: Caudata) from the early Cretaceous Jehol biota. Jértebrata

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Wan, Ÿ. & Gao, K., 1997. - Earliest Asian discoglossid frog from western Liaoning. Chinese Science Bulletin, 44 (7):

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Wu, G. & Tian, W, 1995. - À new Amolops species from southern Yunnan. Sichuan Journal of Zoology, 14 (suppl.):

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——— 1994b. - À new species of family Ranidae - Limmonectes fujianensis from Fujian, China (Amphibia: Anura). Acta

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— 1995. - Taxonomie studies on the small type Megophrys in China including descriptions of the new species

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Ë Zua0, CHANG & ZHaO, 2000: 190].

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Ye, C., FE, L. & Marsut, M., 1995. - Taxonomic studies of Chinese Rana japonica Guenther. Acta herpetologica sinica,

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Yüsnz, E., 1991. - Une nuevo Colosterhus (Amphibia: Dendrobatidae) de la Sierra de Barbacoas, Estado Lara,

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Zuao, E.-M., 1999. - Diagnoses of à new frog and a new snake from China. Sichuan Journal of Zoology, 18 (3): inside

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Zicuer, T. & KômLER, J,, 2001. - Rhacophorus orlovi sp. n. ein neuer Ruderfrosch aus Vietnam (Amphibia: Anura:

Rhacophoridae). Sauria, 23 (3): 31-46.

Zwrez, R. G., 2000. - Partition of the Australopapuan microhylid frog genus Sphenophryne with descriptions of new

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Source : MNHN, Paris

Alytes, 2005, 23 (1-2): 70-80.

Amphibia Mundi. 1.3.

Recent amphibians:

suprageneric taxonomic additions

(967-2002)

Alain DuBois

Vertébrés: Reptiles et Amphibiens,

USM 0602 Taxonomie and Collections,

Département de Systématique and Evolution,

Muséum national d'Histoire naturelle,

25 rue Cuvier, 75005 Paris, France

<adubois@mnhn.fr>

The current International Code of Zoological Nomenclature (ANONYMOUS, 1999) only regulates some

of the nomina of zoological taxa, belonging to three “groups of names” or better “nominal-series”

(Dusoïs, 2000): the species-series, the genus-series and the family-series. It is currently not concerned

with the nomenclature of lower-ranked taxa, ie. of the “variety-series” (DuBois, 200$c-d), or of

higher-ranked taxa, i.e. of the “class-series” (Dumors, 2000, 2005c-d). As a result, the nomenclature of

such taxa, supposedly regulated by “usage” and “consensus” among specialists, is in fact arbitrary and

chaotic, which causes problem for communication among taxonomists and especially between the

latter and all non-specialist users of zoological nomina. For this reason, Duois (2005c-d) recently

proposed a set of rules for the nomenclature of class-series taxa. For the time being, only brief

summaries of these proposed rules have been published (Durois, 2004, 2005a), and their discussion by

the international community of zoologists, before their possible incorporation in the Code, may take

time. Regarding the Neosarracum (i.e., recent amphibians, taxa represented by at least one species in

the currently living fauna of our planet: see DuBois, 2004), in the series Amphibia Mundi, for reasons

explained in Dusois (20056), such nomina are currently not used, but this may change in the future,

when more robust hypotheses on the relationships among amphibian fossil and recent groups are

available and widely accepted. It will then be useful to have a list of available class-series nomina, some

of which may have then to be considered as valid. As changes are also likely to occur at family level and

below, a similar list for family-series nomina will also be useful.

The present list presents additions in the taxonomy of Neogarracut for taxa above rank genus,

published until 2003 after the two lists of such taxa of KUHN (1967) and Duois (1984), or ignored in

these two lists. The period covered by these additions starts in 1984 for family-series taxa of living

anurans, and in 1967 for all other taxa and nomina. It ends on 31 December 2002 for all these groups.

New nomina of the family-series (i.e., families, subfamilies, tribes and subtribes; DUBOIS, 2000,

2005c-d) are printed in sazz carrraz Irauics, followed by the nomina of their rype-genera, and by the

country of the rype-localiy of the type-species of the latter (not the currently known or inferred

geographical distribution of the taxon, that may be much larger).

New nomina of the class-series (i.e., orders, classes, etc.; DuBois, 2000, 2005a, c-d) are printed in

BOLD SMALL CAPITALS. Às class-series nomina below the rank order are not recognized in the

Source : MNHN, Paris

Dugois 71

ergotaxonomy used here (DuBois, 20056), any new nomen of this nominal-series is simply listed below

the nomen of the least inclusive class-series taxon including all its originally included genera or

conucleogenera (see Dugois, 20054), followed between square brackets by the rank afforded to this

nomen in the publication where it was created.

Only new nomina are listed, and taxonomic or nomenclatural changes other than additions (e.g.,

synonymisation or revalidation of nomen, change of rank or or higher taxonomic allocation of taxon,

first-reviser action, orthographic emendation) are not considered here. The new nomina are listed

below by alphabetical order under taxa according to the conservative general taxonomic frame of

Durois (20056). The nomina of all-fossil taxa are preceded by the sign +. Nomenclaturally unavailable

nomina (i.e., nomina nuda and other kinds of anoplonyms, as defined by DuBois, 2000) are presented

below “between quotation marks”.

Classis Ampmimta De Blainville, 1816

Subclassis Nroarracnt Sarasin & Sarasin, 1890

Superordo + ALLocauDATA Fox & Naylor, 1982

Ordo + Azzocaupara Fox & Naylor, 1982

+ Auzocaupara Fox & Naylor, 1982 [ordo].

Superordo Barracuia Brongniart, 1800

Ordo Anura Duméril, 1806

ArGHarosALIENTIA Roëek, 1981 [ordo].

BomBiNanurA Ford & Cannatella, 1993 [“taxon”’].

DiscocLossanurA Ford & Cannatella, 1993 [“taxon”].

LeioPELMATANURA Ford & Cannatella, 1993 [“taxon”].

Néocaupara Cannatella & Hillis, 1993 [no rank given].

Nrosauenria Roëek, 1981 [ordo].

Pararombia Gardiner, 1982 [superordo]. - Comment: Nomen misspelled PararomEA by MILNER

(1988).

PipanuRra Ford & Cannatella, 1993 [“taxon”].

Pirimorpna Ford & Cannatella, 1993 [“taxon”?].

Procera Feller & Hedges, 1998 [superordo].

Incertae sedis

+ Prosaureinar Shubin & Jenkins, 1995. - Type-genus, by original designation: + Prosalirus Kuhn,

1964. - USA (Arizona). Jurassic.

osarracmnae Holman, 1974. - Type-genus, by original designation: + Tregobatrachus Holman,

1964. - USA (Kansas). Miocene.

Source : MNHN, Paris

72 ALYTES 23 (1-2)

Epifamilia Bomavaroromnia Gray, 1825

Superfamilia BomeiaroromdrA Gray, 1825

Familia Bomexaroriar Gray, 1825

Subfamilia + Gomanvar Roëek & Nessov, 1993

+ Gorrarimar Roëek & Nessov, 1993. - Mongolia. Cretaceous.

Epifamilia PeLoaromA Bonaparte, 1850

Superfamilia PsLoraromdea Bonaparte, 1850

Familia PaLorarmar Bonaparte, 1850

Subfamilia Mrcoraevinar Noble, 1931 (1850)

Tribus Lerrosracaunr Dubois, 1983

“ Lrprosracann®” Dubois, 1980. - Type-genus, by implicit etymological designation: Leprobrachium

Tschudi, 1838. — Indonesia (Java). - Comment: Nomenclaturally unavailable nomen, as

published conditionally (Art. 15.1).

Larrogracuunar Dubois, 1983. - Type-genus, by implicit etymological designation: Leprobrachium

Tschudi, 1838. — Indonesia (Java).

OrsozaLaxiNar Tian & Hu, 1985. - Type-genus, by original designation: Oreolalax Myers & Leviton,

1962. - China (Sichuan). - Comment: The original spelling of this nomen is incorrect and should

be emended into ORoLALAGINAE, a justified emendation which was first used by Duois (19876).

Subfamilia P#Logarinar Bonaparte, 1850

+ EorgzoranNae Spinar, 1972. - Type-genus, by original designation: + Eopelobates Parker, 1929. —

Germany. Oligo-Miocene boundary.

Epifamilia Prromia Gray, 1825

Superfamilia Proipea Gray, 1825

Familia Prrib4r Gray, 1825

Subfamilia Dacrvzeruinar Hogg, 1838

Sizvraninar Cannatella & Trueb, 1988. - Type-genus, by implicit etymological designation: Silurana

Gray, 1864. - Nigeria.

Source : MNHN, Paris

Dugois 73

Epifamilia Ravoroza Rafinesque-Schmaltz, 1814

Superfamilia HyorrA Rafinesque, 1815

Familia Buroxipar Gray, 1825

Srersoraemint Dubois, 1987a. — Type-genus, by original designation: Stephopaedes Channing,

1978. - Zimbabwe.

Superfamilia RanorprA Rafinesque-Schmaltz, 1814

Familia Micronvunar Günther, 1858 (1843)

Subfamilia Asrrrorarvivar Günther, 1858

Tribus Barvcexvinr Burton, 1986

Barvéenyinr Burton, 1986. - Type-genus, by original désignation: Barygenys Parker, 1936. —

Papua New Guinea.

Tribus Cazzuzorvr Dubois, 1988

Cazzuzorinr Dubois, 1988. - Type-genus, by original designation: Callulops Boulenger, 1888. —

Papua New Guinea.

Subfamilia Micronvzinar Günther, 1858 (1843)

Ororneywnar Wassersug & Pyburn, 1987.- Type-genus, by original designation: Orophryne Boulen-

ger, 1900. - Guyana.

Subfamilia Parynomrivar Noble, 1931

PHRYNOMANTINI Burton, 1986. - Type-genus, by original designation: Phrynomantis Peters, 1867. —

South Africa.

Familia Brevicierrioar Bonaparte, 1850

Subfamilia Brevicmnar Bonaparte, 1850

Tomorrerinr Dubois, 1987a. - Type-genus, by original designation: Tomoprerna Duméril & Bibron,

1841. - South Africa.

Source : MNHN, Paris

74 ALYTES 23 (1-2)

Familia Ran1DAE Rafinesque-Schmaltz, 1814

Subfamilia CoNrauIN4r Dubois, 1992

Conan Dubois, 1992. - Type-genus, by original designation: Conraua Nieden, 1908. - Came-

roon.

Subfamilia DicroGLOSSINAE Anderson, 1871

Tribus LimnonecrINI Dubois, 1992

Lumnonscrnr Dubois, 1992. - Type-genus, by original designation: Limmonectes Fitzinger, 1843. -

Indonesia (Java).

Tribus Occmnozron Fei, Ye & Huang, 1991

Occunozvoinar Fei, Ye & Huang, 1991. - Type-genus, by original designation: Occidozyga Kuhl &Van

Hasselt, 1822. — Indonesia (Java).

Tribus Par Dubois, 1992

Painr Dubois, 1992. - Type-genus, by original designation: Paa Dubois, 1975. - Nepal.

Subfamilia LANKANE

1x4 Dubois & Ohler, 2001

Lankaxecrinar Dubois & Ohler, 2001. - Type-genus, by original designation: Lankanectes Dubois &

Ohler, 2001. - Sri Lanka.

Subfamilia Manrzuuinar Laurent, 1946

Tribus Boon Vences & Glaw, 2001

Boomnar Vences & Glaw, 2001. - Type-genus, by original designation: Boophis Tschudi, 1838. -

Madagascar.

Tribus LazrosromniVences & Glaw, 2001

LauosronarVences & Glaw, 2001. - Type-genus, by original designation: Laliostoma Glaw, Vences &

Bôhme, 1998. - Madagascar. - Comment: This family-series nomen was ill-formed as the stem of

the nomen Laliostoma is Laliostomat-. However, according to Art. 29.4 of the current version of the

Code (ANONYMOUS, 1999), in such cases the original spelling “must be maintained as the correct

Source : MNHN, Paris

Dugois 75

original spelling”, artificially considering that “its stem is formed from the name of the type genus

as though it were an arbitrary combination of letters”.

Subfamilia Micrixauiar Dubois, Ohler & Biju, 2001

MicrixauNar Dubois, Ohler & Biju, 2001. - Type-genus, by original designation: Micrixalus Boulen-

ger, 1888. - “Southern India”.

Subfamilia NycrsarracHiNar Blommers-Schlôsser, 1993

NycnsarracuNar Blommers-Schlôsser, 1993. - Type-genus, by original designation: Nycribatrachus

Boulenger, 1882. — India (Kerala).

Subfamilia Prrcxapenvar Dubois, 1987

Prycuapexinr Dubois, 1987a.- Type-genus, by original designation: Prychadena Boulenger, 1917.-La

Réunion, Mascarene Islands.

Subfamilia Ranrxar Rafinesque-Schmaltz, 1814

Tribus Ranir Rafinesque-Schmaltz, 1814

< AnoLonnar * Yang, 1989. - Type-genus, by original designation: Amolops Cope, 1865. - “Afghanis-

tan”. - Comment: nomen nudum.

AmoLorsiNar Yang, 1991. - Type-genus, by original designation: Amolops Cope, 1865. “Afghanistan”.

— Comment: The original spelling of this nomen is incorrect and should be emended into

AMOLOPINAE, à justified emendation which was first used by FEI, YE & HUANG (1991).

Subfamilia RaNIXALINAE Dubois, 1987

Ranixazinr Dubois, 1987a. - Type-genus, by original designation: Ranixalus Dubois, 1986. — India

(Karnataka).

Ixpraniar Blommers-Schlôsser, 1993. - Type-genus, by original designation: Indirana Laurent,

1986. - India (Kerala).

Subfamilia Raacorsoinar Hoffman, 1932 (1858)

Tribus Burerrunr Channing, 1989

Busrcrrunar Channing, 1989. - Type-genus, by original designation: Buergeria Tschudi, 1838. —

Japan.

Source : MNHN, Paris

76 ALYTES 23 (1-2)

Ordo UronELa Duméril, 1806

Incertae sedis

Familia + ProsIRENIDAE Estes, 1969

+ ProsmenDar Estes, 1969. - Type-genus by original designation: + Prosiren Goin & Auffenberg,

1958. - USA (Texas). Cretaceous.

Familia + Scarnerreronripar Auffenberg & Goin, 1959

+ Eoscaraerprronnar Nessov, 1981. - Type-genus by original designation: + Foscapherpeton Nessov,

1981. - Uzbekistan. Cretaceous.

Epifamilia CrvrrosancnomtA Fitzinger, 1826

Superfamilia Crvprosrancnonea Fitzinger, 1826

Familia Crverosrancanar Fitzinger, 1826

+ Avrrurinas Gubin, 1991. - Type-genus, by original designation: + Aviturus Gubin, 1991. - Mongolia.

Palacocene.

Familia Hyxogupar Cope, 1859 (1856)

Subfamilia Proronvwosunar Bei & Ye, 2000

Prorouvnosunar Fei & Ye, 2000. - Type-genus, by original designation: Prorohynobius Fei & Ye,

2000. - China (Sichuan).

Epifamilia + Karaurompra Ivachnenko, 1978

Superfamilia + KarauroipeA Ivachnenko, 1978

Familia + KaraURIDAE Ivachnenko, 1978

+ Karauribar Ivachnenko, 1978. - Type-genus, by original designation: + Karaurus Ivachnenko,

1978. - Kazakhstan. Jurassic.

Epifamilia Sireort4 Gray, 1825

Superfamilia Sikevorra Gray, 1825

Familia Sexroaë Gray, 1825

+ Norsersronnoar Rage, Marshall & Gayet, 1993. - Type-genus, by original designation: + Norerpeton

Rage, Marshall & Gayet, 1993. - Bolivia. Cretaceous.

Source : MNHN, Paris

Dugois 77

Superordo Gymnormona Rafinesque-Schmaltz, 1814

Ordo Grmnormoxa Rafinesque-Schmaltz, 1814

Ericruer Lescure, Renous & Gasc, 1986 [infraordo].

Rmvarremaromt Lescure, Renous & Gasc, 1986 [subordo].

Srænonoribet Lescure, Renous & Gasc, 1986 [subordo].

Familia Casciznpar Rafinesque-Schmaltz, 1814

Arrocarcirni Lescure, Renous & Gasc, 1986. - Type-genus, by original designation: Afrocaccilia

Taylor, 1968. - Kenya.

Brasorvrau Lescure, Renous & Gasc, 1986. - Type-genus, by original designation: Brasiloryphlus

Taylor, 1968. - Brazil (Amazonas).

Dermornmnar Taylor, 1969. - Type-genus, by original designation: Dermophis Peters, 1879. - Mexico.

Grorevrenmar Lescure, Renous & Gasc, 1986. - Type-genus, by original designation: Georrypetes

Peters, 1880. - Gabon.

Granisonuzar Lescure, Renous & Gasc, 1986. - Type-genus, by original designation: Grandisonia

Taylor, 1968. - Seychelles.

Gruorntar Lescure, Renous & Gasc, 1986. - Type-genus, by original designation: Gymmopis Peters,

1874. - Panama.

Herreunar Lescure, Renous & Gasc, 1986. - Type-genus, by original designation: Herpele Peters,

1879. - Gabon.

Ixporvemrini Lescure, Renous & Gasc, 1986. - Type-genus, by original designation: {ndoryphlus Taylor,

1960. — India (Maharashtra).

Oscarcunpar Lescure, Renous & Gase, 1986. - Type-genus, by original designation: Oscacailia Taylor,

1968. - Panama.

Psrunosnonorrni Lescure, Renous & Gasc, 1986. - Type-genus, by original designation: Pseudosi-

phonops Taylor, 1968. - Brazil.

Familia Jcravormupas Taylor, 1968 (1843)

Icrrnvormmpar Taylor, 1968. - Type-genus, by original désignation: Jchthyophis Taylor, 1968. — Sri

Lanka.

Familia ScozrcomorPHibAr Taylor, 1969

Scozrcomorrmpar Taylor, 1969. - Type-genus, by original designation: Scolecomorphus Boulenger,

1883. - Tanzania.

Familia TrxLoNecrIDAE Taylor, 1968

Poramoryrazinar Lescure, Renous & Gasc, 1986. - Type-genus, by original designation: Poromotyphlus

“Taylor, 1968. - Venezuela. - Comment: The original nomen of this family is incorrect and should

be emended into PoromorvræLinar, according to Art. 35.4.1 of the Code.

Psrunoryenronecrinr Lescure, Renous & Gasc, 1986. - Type-genus, by original designation: Pseudo-

Lyphlonectes Lescure, Renous & Gasc, 1986. - Colombia.

Source

: MNHN, Paris

78 ALYTES 23 (1-2)

Trrmonecnnae Taylor, 1968. — Type-genus, by original designation: Tiphlonectes Peters, 1879. —

French Guyana.

Familia UraroryPaLib4E Nussbaum, 1979

UrarorvraLinar Nussbaum, 1979. - Type-genus, by original designation: Uracoryphlus Peters, 1979. —

India (Kerala).

Superfamilia RavarremaromneA Nussbaum, 1977

Familia Rannarremarmoar Nussbaum, 1977

RHINATREMATIDAE Nussbaum, 1977. - Type-genus, by original designation: Rhinatrema Taylor, 1968. —

Venezuela.

Epifamilia + EocarcrLromia Jenkins & Walsh, 1993

Superfamilia + Eocaseiroipea Jenkins & Walsh, 1993

Familia + Eocascinpar Jenkins & Walsh, 1993

+ Eocarcntamnar Jenkins & Walsh, 1993. - Type-genus, by original designation: + Eocaccilia Jenkins &

Walsh, 1993. - USA (Arizona). Jurassic. - Comment: The original nomen of this family is

incorrect and should be emended into Eocarcizupar, a justified emendation first used by DuBoIs

(2005b). The original spelling was clearly derived from that of the familial nomen CarciLtAIDAE.

The latter spelling was once adopted by ICZN (ANONYMOUS, 1987) to avoid homonymy with a

familial nomen of Insects, but this decision was later modified by ICZN (ANONYMOUS, 1996) to

return to the well-known spelling Carcizupar. In 1993, the Code in force was the so-called third

edition (ANONYMOUS, 1985) according to which an incorrect original familial nomen must be

corrected. This rule was changed in the so-called fourth edition (ANONYMOUS, 1999; see above

under LaL1osrromNas) but it applies to any familial nomen published before 31 December 1999.

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B8L DU

MUSÉ

PARIS

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AIVTES

International Journal of Batrachology

published by ISSCA.

EDITORIAL BOARD

Chief Editor: Alain Dumors (Laboratoire des Reptiles et Amphibiens, Muséum national d'Histoire naturelle,

25 rue Cuvier, 7500$ Paris, France; <adubois@mnhn.fr>).

Deputy Editor: Thierry LODÉ (Laboratoire d’Ecologie animale, Université d'Angers, 2 boulevard Lavoisier,

49045 Angers Cedex, France; <thierry.lode@univ-angers.fr>).

Conservation Editor: Stephen J. RICHARDS (Vertebrates Department, South Australian Museum, North

Terrace, Adelaide, S.A. 5000, Australia; <Richards.Steve@saugov.sa.gov.au>).

Alytes Editorial Board: Franco ANDREONE (Torino, Italy); Lauren E. Brown (Normal, USA); Janalee P.

CALDWELL (Norman, USA); Ulisses CARAMAsCHI (Rio de Janeiro, Brazil); Günter GOLLMANN (Wien,

Austria); Heinz GriLLIrsCH (Wien, Austria); Tim HALLIDAY (Milton Keynes, United Kingdom); W. Ronald

Heer (Washington, USA); Esteban O. LaviLLA (Tucumän, Argentina), Masafumi MATSUI (Kyoto,

Japan); Donald F. MCALPINE (Saint John, Canada); Alain PAGANO (Angers, France); John C. POYNTON

(London, England); Miguel Vences (Konstanz, Germany).

Amphibia Mundi Editorial Board: Alain Dunois, Chef Editor (Paris, France); Ronald IL. CRoMBIE (San

Francisco, USA); Stéphane GROSIEAN (Paris, France); W. Ronald HEYER (Washington, USA); JIANG

Jianping (Chengdu, China), Esteban O. LAVILLA (Tucumän, Argentina); Jean-Claude RAGE (Paris,

France); David B. Wake (Berkeley, USA).

ee He Team (Paris, France): Alain Durois (texts); Roger BouR (tables); Annemarie OHLER

igures).

Book Review Editor: Annemarie OHLER (Paris, France).

SHORT GUIDE FOR AUTHORS

(for more detailed Instructions to Authors, see Alytes, 1997, 14: 175-200)

Alytes publishes original papers in English, French or Spanish, in any discipline dealing with amphibians.

Beside articles and notes reporting results of original research, consideration is given for publication to synthetic

review articles, book reviews, comments and replies, and to papers based upon original high quality illustrations

(such as colour or black and white photographs), showing beautiful or rare species, interesting behaviours, etc.

The title should be followed by the name(s) and address(es) of the author(s). The text should be typewritten

or printed double-spaced on one side of the paper. The manuscript should be organized as follows: English

abstract, introduction, material and methods, results, discussion, conclusion, French or Spanish abstract,

acknowledgements, literature cited, appendix.

Figures and tables should be mentioned in the text as follows: fig. 4 or tab. 4. Figures should not exceed 16 X

24 em. The size of the lettering should ensure its legibility after reduction. The legends of figures and tables

should be assembled on a separate sheet. Each figure should be numbered using a pencil.

References in the text are to be written in capital letters (BOURRET, 1942; GRAF & POLLS PELAZ, 1989; INGER

et al., 1974). References in the Literature Cited section should be presented as follows:

BouRRET, R., 1942. — Les batraciens de l'Indochine. Hanoi, Institut Océanographique de l’Indochine: i-x + 1-547,

1. 1-4.

GRAF, J.-D. & PoLLs PELAZ, M. 1989. - Evolutionary genetics of the Rana esculenta complex. In: R. M. DAWLEY

& JP Bogarr (ed.), Évolution and ecology of unisexual vertebrates, Albany, The New York State Museum:

289-302.

INGER, R. E, Voris, H. K. & Vois, H. H., 1974. - Genetic variation and population ecology of some Southeast

Asian frogs of the genera Bufo and Rana. Biochem. Genet., 12: 121-145.

Manuscripts should be submitted in triplicate either to Alain Duois (address above) if dealing with

amphibian morphology, anatomy, systematies, biogeography, evolution, genetics, anomalies or developmental

biology, or to Thierry LODÉ (address above) if dealing with amphibian population genetics, ecology, ethology or

life history, or to Stephen J. RICHARDS (address above) if dealing with conservation biology, including declining

amphibian populations or pathology. Acceptance for publication will be decided by the editors following review

by at least two referees.

If possible, after acceptance, a copy of the final manuscript on a floppy disk (3 % or 5 4) should be sent to

the Chief Editor. We welcome the following formats of text processing: (1) preferably, MS Word (1.1 to 6.0, DOS

or Windows), WordPerfect (4.1 to 5.1, DOS or Windows) or WordStar (3.3 to 7.0); (2) less preferably, formated

DOS (ASCII) or DOS-formated MS Word for the Macintosh (on a 3 % high density 1.44 Mo floppy disk only).

Page charges are requested only from authors having institutional support for this purpose. The publication

of colour photographs is charged. For each published paper, 25 free reprints are offered by ISSCA to the

author(s). Additional reprints may be purchased.

Published with the support of AALRAM

(Association des Amis du Laboratoire des Reptiles et Amphibiens

du Muséum National d'Histoire Naturelle, Paris, France).

Directeur de la Publication: Alain DuBOr.

Numéro de Commission Paritaire: 64851.

Source : MNHN, Paris: