FORSCHUNGS
museum
KOENIG
Bonn zoological Bulletin
Supplementum Vol. 61 (2014)
formerly: Bonner zoologische Monographien
A monograph of the Sericini of Sri Lanka
(Coleoptera: Scarabaeidae)
Silvia Fabrizi & Dirk Ahrens
An open access journal of organismal zoology, published by
Zoologisches Forschungsmuseum Alexander Koenig - Leibniz-lnstitut fur Biodiversitat der Tiere, Bonn
BONN ZOOLOGICAL BULLETIN - SUPPLEMENTUM (BzB-S),
formerly “Bonner zoologische Monographien”,
ss as above, tel. +49 228-9122-425,
;e 1971. It is published by the Zoological Research
;inal zoology papers too long for inclusion in our in
rs of this issue: Silvia Fabrizi & D
:r. 129, 53113 Bonn, Gen
nt of the map of the e>
I bzm_61_ahrens_BZB-eml 22.04.14 12:41 Seite 1
A monograph of the
Sericini of Sri Lanka
(Coleoptera: Scarabaeidae)
Silvia Fabrizi & Dirk Ahrens
Bonn zoological Bulletin - Supplen
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_ahrens_
Contents Volume 61
Abstract
Key words
Introduction
Material and methods
Depositories
Systematic part
Key to genera of Sri Lanka
Periserica Brenske, 1898
Selaserica Brenske, 1 897
Neoserica (s.l.) Brenske, 1894
" Maladera"
Maladera lugubris group
Maladera (sbg. Cephaloserica)
Maladera fistulosa group
Maladera madurensis group
Maladera setosa group
Maladera detersa group
Maladera (sbg. Hemiserica)
“Serica"
Species incertae sedis
Zoogeography
Acknowledgements
References
Tables
Appendix. Gazetter of localities
Index
13
34
39
39
47
58
90
91
91
92
97
97
98
100
101
122
124
Bonn zoological Bulletin - Supplementum 61: 1-124
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April 2014
A monograph of the Sericini of Sri Lanka
(Coleoptera: Scarabaeidae)
INTRODUCTION
sive study of this fauna was conducted by Walker (1859)
eration of the island followed later (Moser 1916; Frey
1972, 1974).
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(Figs 2A-C, 16A)
spec ” (BMNH).
Description. Length: 7.2 mm, length of elytra: 4.3 mm,
iiiililiiiiiil
12
serrated line; posterior ventral margin almost straight,
weakly widened in apical half, neither serrated ventrally
nor dorsally, glabrous. Metatibia moderately wide and
short, widest at middle, weakly narrowed apically, ratio
width/ length: 1/ 3.6, longitudinally convex dorsally, with
two groups of spines, basal one at one quarter, apical one
at three quarters of metatibial length; lateral face longi-
tudinally convex, smooth, laterally with very sparse and
fine punctures; ventral margin with four strong equidis-
tant spines; medial face smooth, apex interiorly near tarsal
articulation moderately truncated. Tarsomeres impunctate
dorsally, with dense, fine setae ventrally; metatarsomeres
glabrous ventrally, with a strongly serrated carina, subven-
trally with a second, smooth longitudinal carina, first
metatarsomere a little shorter than following two tar-
someres combined and one third of its length longer than
the dorsal tibial spur. Protibia moderately long, tridentate,
basal tooth small. All claws symmetrical, feebly curved
and long, with normally developed basal tooth.
Aedeagus: Fig. IF-H. Habitus: Fig. 14C.
Variation. Female: Antennal club composed of three an-
tennomeres, slightly shorter than remaining antennomeres
combined.
Remarks. The abdomen and genitalia of the holotype of
Periserica nigripennis are lost. However, it is virtually
identically in all other external features with the lectotype
of Periserica fulvostriata except for the colour of the ely-
tra. Although the elytra of P. nigripennis are darker, the
pattern of spots is recognizable and virtually identical to
that of P. fulvostriata.
Selaserica Brenske, 1897
Selaserica Brenske, 1 897: 355 (type species by monotypy:
Selaserica nitida (Candeze, 1861)).
Key to species of Selaserica (SS):
1 Hypomeron without a transverse carina basiventrally.
Metafemur without a serrated line adjacent to anteri-
or margin. Antenna with nine antennomeres
{Selaserica nitida group) 2
r Hypomeron with a transverse carina basiventrally.
Metafemur with a serrated line adjacent to anterior
margin. Antenna with ten antennomeres
{Selaserica splendiflca group) 13
2 Antennal club in male with 4 antennomeres 3
2’ Antennal club in male with 3 antennomeres 4
3 Antennal club almost as long as remaining anten-
nomeres combined Se. nitida (Candeze)
3’ Antennal club distinctly longer than remaining anten-
nomeres combined Se. knucklensis sp. n.
4 Frons at least in anterior two thirds shiny 5
4’ Frons extensively dull, shiny only narrowly behind
frontoclypeal suture 7
5 Pronotum shiny. Large species, body larger than
11 mm 6
5’ Pronotum dull. Small species, body smaller than 6.5
mm. Frons completely shiny. Elytra glabrous, except
the lateral setae Se. kanneliyana sp. n.
6 Dorsal surface dark or dark reddish brown. Elytra dull,
intervals deeply impressed. Metatibia wide and strong-
ly flattened, dorsal edge sharp. ...Se. lucidicollis sp. n.
6’ Dorsal surface light reddish brovm. Elytra complete-
ly shiny, intervals weakly impressed. Metatibia slen-
der and moderately flattened, dorsal edge almost lon-
gitudinally convex Se. scutellaris Avcosn ($ !)
7 Metatibia wide and strongly flattened, dorsal margin
sharp 8
7’ Metatibia slender and moderately flattened, dorsal
margin almost longitudinally convex 12
8 Apex of phallobasis (in dorsal view) strongly asym-
metrical Se. maculicauda (Arrow)
8’ Apex of phallobasis (in dorsal view) almost symmet-
rical 9
9 Antennal club 1.5 times as long as remaining anten-
nomeres combined Se. implicata (Brenske)
9’ Antennal club as long as or slightly longer than re-
maining antennomeres combined 10
10 Basal lobe of left paramere narrow and long
Se. padukkana sp. n.
10’ Basal lobe of left paramere large and lobiform, indis-
tinctly defined 11
1 1 Eeft paramere evenly pointed at apex. Right paramere,
relative to overall size of aedeagus shorter
Se. impexa sp. n.
1 1 ’ Eeft paramere bluntly truncate at apex. Right para-
mere, relative to overall size of aedeagus longer
Se. convexiuscula sp. n.
12 Antennal club as long as remaining antennomeres
combined Se. confirmata (Walker)
12’ Antennal club more than twice as long as remaining
antennomeres combined. ..Se. distincticomis (Brenske)
13 Antennal club composed of three antennomeres.
Small species, body smaller than 5.5 mm. Frons pos-
teriorly narrowly dull 14
13’ Antennal club composed of four antennomeres. Larg-
er species, body larger than 6 mm 15
14 Dorsal surface nearly glabrous.
Se. wilpattuensis sp. n.
14’ Dorsal surface densely setose. ..Se. meridionalis sp. n.
15 Metatibia laterally with a very short serrated line on-
ly. Body larger than 9 mm 16
15’ Metatibia laterally with a serrated line as long as one
third of metatibia... 17
16 Dorsal surface in parts dull Se. ser icea Axxosn
16’ Dorsal surface completely shiny Se. nuwarana sp. n.
illHlill
i im i «i 1 1 1 IIP 1 1 pi I i !i I illl 1 1 nil 1 i i i II 1 1 HI If Hi i | i
l!liiPiif!i|liiil!iii!iliifilW lii I
18
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20
22
24
tioii. Length: 9.6 mm, length of elytra: 7.3 mm,
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30
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1 ’ Body size smaller, total length <5.4 mm. Body surface
yellowish brown and shiny N. kalaarensis sp. n.
2 Antennal club in both sexes with four antennomeres..
N. bombycina (Karsch)
2’ Antennal club in male with six antennomeres
N. sexfoliata Moser
Remarks. Neoserica resulted polyphyletic in a first mo-
lecular phylogeny of Sericini (Ahrens & Vogler 2008),
however, a revision of the genus concept based on a com-
bined molecular and morphological analysis needs a sig-
nificantly wider and possibly complete taxon sampling.
Therefore, following Ahrens (2004), we consider all
species that were so far assigned to Neoserica and that do
not belong to the monophyletic Neoserica (sensu stricto)
group (Ahrens 2003b) preliminary as Neoserica sensu la-
to.
Neoserica bombycina (Karsch, 1882)
(Figs 5M-0, 141, 17A)
Serica bombycina Karsch, 1882: 387.
Neoserica bombycina. Brenske 1898: 281; Arrow: 1916:
431.
Serica quadriflabellata Brenske, 1896: 154, syn. n.
Neoserica quadriflabellata. Brenske 1898: 253.
Neoserica sericata Brenske, 1898: 280, syn. n.
Type material examined. Syntypes {bombycina)-. 1 $
“61918/ ^/bombycina Karsch* Ceylon. Mus. Colombo”
(ZMHB), 1 $ [without labels but the same type of pin -
therefore it is to assume that thios specimen is a syntype
as well. Consequently, the authors have added a copy of
the labels of the other syntype:] “61918/ $/ A. bombyci-
na Ceylon, Mus. Colombo” (ZMHB). Syntypes (quadri-
flabellata)-. 3 1 $ “Konbir/ 4-jlabellata type Brsk.”
(ISNB), 1 $ “Konbir [subsequently labelled by corator]/
4-jlabellata type Brsk.” (ISNB), 1 6' “Konbir/ A-flabel-
lata! coll. Brenske” (ZMHB), 1 6' “Konbir/ Coll.
Brenske/4-flabellata” (ZMHB). Syntype {sericataf. 1 $
“Madras/ sericata type Brsk./ Type/ Museum Paris Coll.
Generale” (MNHN), 1 $ “Madras” (MNHN).
Additional material examined. S. India: 1 ex. ($) “Coll.
R.I.Sc.N.B. Inde Barway P. Cardon/ Type/ det. Brenske
1898 Berk Ent. Zeit: 252/ Serica 191. Brsk.” (ISNB), 3
ex. “India: Pondicherry Karalkal, IX. 1988 T.R.S. Nathan”
(CMNC), 1 6', 1 9 “Madras” (offensichtlich keine Typen
von quadroflabellata, da das Mannchen Brenske unbekan-
nt blieb, ZMHB), 1 $ “Mysore S. Indien III.53” (CF), 1
$ “Bengal Mandar/ etwas abweichend [hand written
Brsk.]/ coll. Brenske” (ZMHB), 1 ex. “India: Anamalai
Hills, Cinchona 700‘ [m] X.1959 / P. S. Nathan Bishop
Museum” (BPBM), 1 ex. “S. India: Madras State, Ana-
malai Hills, Cinchona 1050 m 1.67/ Collector P. S. Nathan”
(BPBM), 2 ex. “South India Pondicherry State, Karikal
1. 1963/ Collector P. S. Nathan” (BPBM), 1 ex. “India (S):
Kerala State: Walayar Forests, 213 m, X.1959/ P. S.
Nathan Bishop Museum” (BPBM), 1 ex. “Ramnad Hin-
doustan” (MNHN), 1 ex. “India Kistna Dt. Avanigadda
VII.53 Rimondi” (MSNM), 1 ex. “Cote de Malabar M.
Maindron/ Mahe Chass. inidgenes Aout 1901/ Museum
Paris Cote de Malabar Mahe M. Maindron” (MNHN), 1
ex. “S. India Coimbatore PS. Nathan” (MNHN), 1 ex. “In-
dia Coc[r?]onada/ Simson/Fry Coll. 1905.100.” (BMNH),
1 ex. “W.S. Millard 3rd Nov. 1910 Madras/ Bombay Nat.
Hist. Soc. B.M. 1914^11” (BMNH), 4 ex. “Mysore H.K.
Slater 1901-182.” (BMNH), 1 ex. “India: ex Mus. Cal-
cutta. B.M. 1930-1./ Ind. Mus. Jubbulpore district Cen-
tral prov. 1914. Purchased” (BMNH), 2 ex. “India: Oris-
sa Bhubaneshwar Feb. 13, 1975, 500’ Mary L. Ripley”
(UNSM), 1 ex. “South India Pandioberry State, Karaikal
X.1990 leg. Theresa Raj abai Selva Nathan” (CARL). Sri
Lanka: 1 S “Sri Lanka Ambalangoda 12.8.1985, Madl”
(NHMW), 1 $ “Slid Ceylon Mai 1889 H. Fruhstorfer/
Coll. Brenske” (ZMHB), 1 $ “Ceylon 2de trim 1889 (H.
Fruhstorfer)” (ZMHB), 13 ex. “Ceylon, V.1969 50 kmN
Colombo leg. P. Schmitz” (CF), 1 S “Inde Anglaise Shi-
moga” (MNHN), 1 ex. “Colombo 10.09/ Colombo Cey-
lon \9\6. 169! Neoserica bombycina Karsch Determined
from Description G. J.A.” (BMNH), 2 ex. “Colombo 9.05/
Ceylon 1910-20.” (BMNH), 1 ex. “Bentota 21.7.1973/
Brancucci” (NHMB), 1 ex. “Ceylan M. Maindron/ Co-
lombo 12-20 mai 1901/ Museum Paris Ceylan M. Main-
dron” (MNHN), 3 ex. “Colombo Ceylon, India/
14-VIir44 DE Hardy” (UNSM), 1 ex. “Ceylon, W. Prov.
Yakkala, 18 mis NE Colombo, 16-28.11.62 At light Loc.
10/ Lund University Ceylon Expedition Brinck-Andersen-
Cederholm” (MZLU), 7 ex. “Sri Lanka: Put. Dist.
Wilpettu, National Park, Kali Villu 12-14 June 1975/ Col-
lectors: D. H. Messersmith G. L. Williams & P. B.
Karunaratne” (USNM), 1 ex. “Ceylon: Ham. Dist. Yala,
Palatupana 8-10 March 1972 K. V. Krombein & P. B.
Karunaratne” (USNM), 1 ex. “Sri Lanka: Put Dist. Wilpat-
tu, Maddikulam 15 June 1975/ Collectors: D. H. Messer-
smith G. L. Williams & P. B. Karunaratne” (USNM), 1
ex. “Ceylon: Ham. Dist. Bambowa, 1 . 5mi NW Palatu-
pana 23-X-1970, 50’ O. S. Flint, Jr.“ (USNM), 1 ex.
“Ceylon: Jaf Dist. Elephant Pass 7-XI-1970, 5’ O. S.
Flint, Jr.“ (USNM), 1 ex. “Sri Lanka: Ham. Dist. Palatu-
pana Tank 3-A February 1975/ K. V. Krombein P. B. Kara-
naratne P. Fernando E. G. Dabrera” (USNM). Burma: 1
S “Victoria Point Tenasserim E.T. Atkinson/ Atkinson
Coll. 92-3/ 8.87” (BMNH).
Redescription of male syntype of quadriflabellata.
Length: 7.0 mm, length of elytra: 4.6 mm, width: 3.9 mm.
Body oval, yellowish brown, antenna yellowish, head
shiny, dorsal surface dull, nearly glabrous.
Labroclypeus wide and trapezoidal, widest at base, lat-
eral margins straight and strongly convergent anteriorly.
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Silvia]
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40
ly serrated longitud]
rina beside it, first i
Variation. Length: 7.7-8.4 mm, length of elytra: 5.6-6.4
ly shorter than remaining antenneomeres combined; py-
[hand written G. Frey]” (CF). Paraty]
Bur. det G. Frey, 1972” (CF), 1 6' “Ceylon:
Frey, 1972” (CF), 1 “Ceylon: Padukka43km
0 28.IL-2.IIL1971/ J. & M. Sedlacek Bishop
Museum” (BPBM), 1 6' “Colombo, Ceylon 1.IL58 leg.
G. Frey/ cum Type comparatum/ Autoserica proxima Bur.
det G. Frey, 1972” (CF), 1 6', 5 $$ “Ceylon: Kan Dist.
Hasalaka, 500’ 22-25-XI-1970 O.S. Flint, Jr.” (USNM),
1 6 ', 2 $$ “Ceylon: Kan. Dist. Laksapana, 1200’
23-29-IX-1970 O. S. Flint, Jr.“ (USNM), 1 6 ', 1 $ “Cey-
lon: Kan. Dist. Kandy 1-15 March 1971 Piyadase &
Somapala” (USNM), 1 $ “Sri Lanka: Kan. Dist. Per-
adeniya Hantana Hill 29-III-1973, 2000 ft. Baumann &
Cross” (USNM), 1 $ “Sri Lanka: Rat. Dist. Kalu Ganga
Induruwa Jungle 23-III-1973, 1000 ft. Baumann &
Cross” (USNM), 1 $ “Ceylon: Rat. Dist. Uggakaltota 350
ft. Irrigation Bungalow 31 Jan.-8 Feb. 1970 Davis &
ma Reservoir 7 Aug. 1975 Peyton & Huang” (USNM), 1
$ “Sri Lanka: N. E. Dist. Hakgala Botanic Garden, 23-25
Feb 1977/ K. V. Krombein P. B. Karunaratne P. Fernan-
do D. W. Balasooriya” (USNM), 1 6' “Sri Lanka: N. E.
Dist Hakgala Botanic Garden, 23-25 Febl977 blacklight.
: Tri. Dist. 1
8-11-X-1977/ K. V. Krombein P. B. Karunaratne P. Fer-
nando M. Jayaweera” (USNM), 1 ? “Sri Eanka: Bad. Dist.
light” (USNM), 1 $ “Sri Eanka: Pol. Dist. Polounaruwa
6 May 1974 Gans & Prasanna” (USNM), 1 $ “Sri Ean-
ka: Mate. Dist. Elkaduwa, Hunas Falls 5-IV-1973, 3000
t” (USNM), 1 $ “Sri
I 61: 1-124
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48
It Loc. 10/ Lund
inck-Andersen-C
(MZLU), 1 ex. “Ceylon, W. Prov. Yakkala, 18
Colombo, 1.-14.IIL62 At light Loc. 10/ Lund Ui
Ceylon
(MZLU),
27-VII-1975/Y.-M.
D.W. Balasooriya” (USNM),
Kandy Tour Inn 31-III-1978 M. D.
i.L. Peyton, S.
lex. “SriLan
(MZLU), 4 ex. “Ceylon, W.
5-13.1.62 Loc. 3 At light / Li
(USNM), 1 ex. “Ceylon: Kan. Dist. Nanu Oya, nr Per-
adeniya, 1500’ 22-IX-1970 O. S. Flint, Jr.“ (USNM),1 ex.
“Ceylon: Kan. Dist. Haslaka, Dam Spillway, 300’
23-XI-1970 O. S. Flint, Jr.“ (USNM), 1 ex. “Ceylon: Kan.
y 1971 F' ■
), 2 ex. “Ceylon: Kan. E
a” (USNM), 3 ex. “SriLan-
ht 13-14' '
collected by G.F. Hevel F.E. Dietz IV S. Karunaratne D.W.
Balasooriya” (USNM), 1 ex. “Sri Lanka: Col. Dist. Gam- blackl
paha Botanical Garden 24 May 1975/ Collectros D.H. maV.
Messermilch G.L.
(USNM), 4 ex. “Ce:
Padaviya 180 ft. 27 Feb.-9 Mar. 1970 Davis &
fiiya 1-15
ex. “Cey-
Dist. Kandy 1-15 March 1971 Piyadasa &
(USNM), 7 ex. “Ceylon: Kan. Dist. Kandy
■ 1971 P. P. Spangler/ Collect at black light”
Davies, S. (USNM), 1 ex. “Sri Lanka: Kan. Dist. Kandy, Udawat-
1-3-X-1973 at black light/ K.V. Krombein PB.
2 ex. “Sri Lanka: Kan.
(USNM), 2 ex.
15-17 Sept ;
1977/Collected by KV
^ri Lanka: Mate. Dist. Sigiriya,
collected by D.M. Davies, S.
-unaratne, D. W. Balasooriya” (USNM), 1 (
Mate. Dist. Deniyaya, black light trap 9-
lected by D. M. Davies, S. Karunaratne
Tiya” (USNM), 2 ex. “Sri Lanka: Kal. D
a: Pol. Dist. Polonnamwa 6 May 1974 Gans&Pra-
a” (USNM), 2 ex. “Sri Lanka: Rat. Dist. Udawalawa
June 1975/ Collecters: D. H. Messersmith G. L.
ams & P. B. Karunaratne” (USNM), 1 ex. “Sri Lan-
ka: Amp. Dist. Ekgal Am Tank, 100 m 19-23 Febmary
1977/ K. V. Krombein P. B. Karunaratne P. Fernando D.
W. Balasooriya” (USNM), 1 ex. “Sri Lanka: Col. Dist.
Y. Huang E.
(USNM), 1
Sit:-
MaaB” (NME), 2 ex. “Ceylon E. Prov. Pottuvil
l-12/vii-1983 Ole Mehl leg.” (CDKC).
liiiliii
50
i61: 1-124
_BZB-eml 22.04.14 12:41 Seite 51
Sericini of Sri Lanka 5 1
ver 1906-7 E. Bugnion” (MNHN), 9 ex. “S. Ceylon Ha-
baraduwa 20.1.^.2.1983 T. Osten leg.” (SMNS), 1 ex.
“Ceylon Nuwara Eliya 17.III.78 Renner” (SMNS), 3 ex.
“Ceylon N.W.P. Rajakadaluwa 23.VIII.53 F. Reiser”
(NHMB), 6 ex. “Sri Eanka (Ceylon) Southern Province
Galle, Koggela Lake 14.-28.12.1992 leg. Gerd Kune”
(NME), 8 ex. Sri Lanka Vilpattu 8.X.1982 leg. G. Med-
vedev” (ZIN), 2 ex. “Beruwela Ceylam 22.IV.75 C.G. Mi-
net” (CF), 2 ex. “Beruwela Ceylam 21. IV. 75 C.G. Minet”
(CF), 1 ex. “Beruwela Ceylam 23.IV.75 C.G. Minet” (CF),
1 ex. “Ceylon Bentota 21.7.1973/ Brancucci” (NHMB),
3 ex. “Coll. R.I.Sc.N.B. Ceylan/ J.J. Gillet del, vendit.:”
(ISNB), 3 ex. “Batticaloa, Ceylon. June 1915. C.F.S. Ba-
ker. 1915-338” (BMNH), 1 ex. “Trincomalee, Ceylon.
1915. C.F.S. Baker. 1915-338” (BMNH), 1 ex. “Trinco-
malee, Ceylon. C.F.S. Baker. 1914-448” (BMNH), 4 ex.
“Nilaveli. Ceylon. C.F.S. Baker. 1914^48” (BMNH), 2
ex. “Trincomalee Ceylon, XI-06/ 1989/ Ceylon, E.E.
Green 1910-19” (BMNH), 1 ex. “Batticaloa Ceylon,
5-06/ 1988/ Ceylon, E.E. Green 1910-19” (BMNH), 1 ex.
“Ceylon. Trincomali. 16. VI. 07 Oopah Fst. Bainbrigge-
Fletcher/ Ceylon T.B. Fletcher. 1909-80” (BMNH), 1 ex.
“7.VII.07 Ceylon. Trincomali. / Ceylon T.B. Fletcher.
1909-80” (BMNH), 1 ex. “Colombo, Ceylon/ C.F.S. Ba-
ker/April 1915 1915-199.” (BMNH), 4 ex. “Colombo
Ceylon, India/ 14-VIIF44 DE Hardy” (UNSM), 14 ex. Sri
Lanka Southern Province Galle Distric, Habaraduwa
20.8.^.9.1982 H.J. Bremer leg.” (ZSM), 9 ex. “Ceylon,
W. Prov. Yakkala, 18 mis NE Colombo, 14-31.1.62 At
light Loc. 10/ Lund University Ceylon Expedition
Brinck-Andersen-Cederholm” (MZLU), 6 ex. “Ceylon,
W. Prov. Yakkala, 18 mis NE Colombo, 1-14.IIL62 At
light Loc. 1 0/ Lund University Ceylon Expedition Brinck-
Andersen-Cederholm” (MZLU), 1 ex. “Ceylon, NW. Prov.
Chilaw, 31.1.62 At light Loc. 38/ Lund University Cey-
lon Expedition Brinck-Andersen-Cederholm” (MZLU),
6 ex. “Ceylon, S. Prov. Hikkaduwa, 11 mis NW Galle,
25.1.62 Loc. 22/ Lund University Ceylon Expedition
Brinck-Andersen-Cederholm” (MZLU), 2 ex. “Sri Lan-
ka: Put. Dist. 9 mi E Puttalam 19 June 1975 S. L. Wood
& J. L. Petty/ Collected in black light trap” (USNM), 10
ex. “Sri Lanka: Man. Dist. Cashew Corp., Ma Villu, 17-21
Feb. 1979, blacklight/ Collected by K. V. Krombein T. Wi-
jesinhe S. Siriwaradne T. Gunawaradne” (USNM), 1 ex.
“Sri Lanka: Put. Dist. Wilpattu Park, 100ft. Talawila,
9-1 0-IV-73 Baumann & Cross At black lighf ’ (USNM),
2 ex. “Sri Lanka: Man. Dist. Olaithoduvai, lOmi NW of
Mannar, 0-50 feet black light, 4-5 November 1976/ Col-
lected by G. F. Hevel R. E. Dietz IV S. Karunaratne D.
W. Balasooriya” (USNM), 2 ex. “Sri Lanka: Man. Dist.
8mi SE of Mannar, black light, 15 feet 6 November 1976/
Collected by G. F. Hevel R. E. Dietz IV S. Karunaratne
D. W. Balasooriya” (USNM), 6 ex. “Sri Lanka: Col. Dist.
Hanwella Resthouse 200ft, black light, 2 October 1976/
Collected by G. F. Hevel R. E. Dietz IV S. Karunaratne
D. W. Balasooriya” (USNM), 3 ex. “Ceylon: Kan. Dist.
Nanu Oya, nr. Peradeniya, 1500’ 22-IX-1970 O. S. Flint,
Jr.“ (USNM), 4 ex. “Ceylon: Kan. Dist. Kandy, 1800Ft.
Peak View Motel 7-14 Jan. 1970 Davis & Rowe”
(USNM), 2 ex. “Ceylon: Kan. Dist. Kandy, 1800 ft. Peak
View Motel 15-24 Jan. 1970 Davis & Rowe” (USNM),
1 ex. “Sri Lanka: Kan. Dist. Kandy, 23 November 1974/
Collectors: C. Gans P. Fernando S. Farock” (USNM^ 1
ex. “Sri Lanka: Man. Dist. Kondachchi, Ma Villu
19- IX-1979 In Malaise trap/ T. Wijesinhe L. Jayawick-
rema R. Subasinhe” (USNM), 2 ex. “Sri Lanka: Man. Dist.
Kondachchi, Ma Villu 1 9-IX-l 979 In Malaise trap/ T. Wi-
jesinhe L. Jayawickrema R. Subasinhe” (USNM), 4 ex.
“Sri Lanka: Man. Dist. Kondachchi, Ma Villu
20- IX-1979/ T. Wijesinhe L. Jayawickrema R. Subasin-
he” (USNM), 1 ex. “Sri Lanka: Man. Dist. Kondachchi,
17-19 Feb 1979Malaise trap/ Collectors: K.V Krombein
T. Wijesinhe S. Siriwaradne T. Gunawaradne” (USNM),
1 ex. “Sri Lanka: Man. Dist. Kondachchi, 22-28 January
1978/ PB. Karunaratne T. Wijesinhe M. Jayaweera G. Rat-
navira” (USNM), 1 ex. “Ceylon: Kan. Dist. Hantane Hill
10 December 1970 Piyadasa & Sompala” (USNM), 1 ex.
“Sri Lanka: Galle Dist. Hiniduma 22 Oct. 1973 M. & R.
Robinson” (USNM), 4 ex. “Sri Lanka: Put. Dist. Wilpat-
tu National Park, Kali Villu 12-14 June 1975/ Collectors:
D. H. Messersmith G. L. Williams P. B. Karunaratne”
(USNM), 1 ex. “Sri Lanka: Col. Dist. Beyagama, sea lev-
el 28-29 VIII 1973 Ginter Ekis” (USNM), 2 ex. “Sri Lan-
ka Bentota 23.5.-15.6. 1990 leg. U. Schmidt/ Coll. A. Ska-
le Hof/ Saale” (NME), 2 ex. “Ceylon N.P Mannar 3 1 . 1 .54
F. Reiser/ a. Licht” (NHMB).
Redescription. Length: 7.7 mm, length of elytra: 5.5 mm,
width: 4.5 mm. Body oblong-oval, light reddish brown,
elytra and antenna yellowish brown, labroclypeus shiny,
dorsal surface dull, glabrous except for a few small setae
on head and lateral margins of pronotum and elytra.
Labroclypeus trapezoidal, distinctly wider than long,
widest at base, lateral margins weakly convex and
strongly convergent to moderately rounded anterior an-
gles, lateral border and ocular canthus producing an in-
distinct blunt angle, margins moderately reflexed, anteri-
or margin distinctly and narrowly sinuate medially; sur-
face slightly carinate along the middle in anterior half,
shiny, moderately coarsely and densely punctate, distance
between punctures subequal their diameter, partly fusing
with each other, behind anterior margin with a few erect
setae; frontoclypeal suture feebly incised and weakly
curved; smooth area anterior to eye approximately three
times as wide as long; ocular canthus moderately long and
moderately wide, finely and densely punctate, with two
long terminal setae. Frons dull, anterior quarter shiny, with
fine, dense punctures, glabrous except for two to three sin-
gle setae beside eyes, punctures partly with minute setae.
Eyes moderately large, ratio of diameter/ interocular
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58
widest just before apical third, ratio width/ length: 1/ 2.9,
Key to species
60
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num/ metacoxa: 1/ 2.05. Median apophysis of metacoxa
with very dense and evenly short, white, scale-like setae.
Ion Expedition 1962
(MZLU). Paratypes: 5
(USNM), 1 $ “Sri La
Kandy 29-III-1973,
(USNM).
ly
ly
64
68
iiiiaiiiiiiii isiiWiilSIl ili-iill!
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74
Silvia]
;st just behind middle, st
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76
78
(Figs 9P-R, 18D)
Loc. 167:1/ Lund L
h: 6.2 mm, length of elytra: 4.6 mm,
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ill iiii
i 111 i
If filial II
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86
s, 1 .2 times (6') as long as re-
iiiiliiiiiiiii iili
88
90
92
sal margin impunctate, basally and at middle with a few
single spines beside the serrated line; lateral face longi-
tudinally convex, shiny, with sparse and fine punctures
basally and on ventral portion, with short setae in punc-
tures; ventral margin coarsely serrate, with three robust
and long spines; medial face impunctate and glabrous,
apex interiorly near tarsal articulation sharply and deeply
truncate. Tarsomeres smooth dorsally, with short, sparse
setae ventrally; metatarsomeres with a strongly serrated
ridge ventrally, beside the ridge with a fine longitudinal
Carina, not carinate laterally; first metatarsomere as long
as following two tarsomeres combined and slightly less
than twice as long as the dorsal tibial spur. Protibia mod-
erately long, bidentate, lateral teeth distinctly displaced
apically, slightly but not bluntly widened at middle of the
external margin; protarsal claws symmetrical, basal tooth
of inner claw simply pointed.
Aedeagus: Fig. 13A-C. Habitus: Fig. 15J.
Serica lurida Brenske, 1898
(Figs 13D-F, 20B)
Serica lurida BxQmkQ, 1898: 261.
Serica maculifera Brenske, 1898 (nee Brenske, 1894):
260; Ahrens 1996: 31, syn. n.
Serica rubescens 1916: 436, syn. n.
Type material examined. Syntypes {lurida)'. 1 S “Cey-
lon./ Nietner 860/ lurida Type Brsk./ Typus” (NHMW),
1 S “Ceylon./ Nietner 860/ lurida Type Brsk./ Typus”
(NHMW), 1 $ “Ceylon./ Nietner 860/ Type Brsk./
Typus” (NHMW), 1 $ “Ceylon/ lurida m./ coll Brenske”
(ZMHB), 1 $ “277/ Haputate Tea estate at 5000 ft./ Mus.
Colombo Ceylon/ Serica lurida m./ Museum Leiden Ser-
ica lurida Det. Brenske/Cat. No. 1” (RMNH). Syntypes:
{maculifera Brsk., 1898 (nee Brsk. 1894); see Ahrens
1996): 1 6 ', 2 $$ “Ceylon Nietner” (ZMHB), 1 $ of these
additionally with ^^maculifera Brsk.” (ZMHB), 1 S “Cey-
lon Nietner/ 38970” (ZMHB), 1 6' “38970” (ZMHB), 1
6' “Ceylon/ 24943/ Brsk.” (ZMHB), 1 6', 1 ?
“24943” (ZMHB), 4 6'6', 3 $$ “Ceylon/ coll. Brenske”
(ZMHB), 1 6' of these additionally with "'maculifera
Typ.Brsk.” (ZMHB), 1 6 ', 1 $ “Ceylon/ zwaczz/z/era Brsk.”
(ZMHB). Syntypes {rubescens)'. 1 S “Bogawantalawa
4.900-5.200 ft. 21.III.^.IV.82/ Ceylon G. Lewis 1910-
320/ Serica rubescens type Arrow/ =S. maculifera Brsk.,
1 898 (nee Brsk. 1 894) det. Ahrens 1 995” (BMNH), 4 $ $
“Bogawantalawa 4.900-5.200 ft. 2LIIL^.IV.82/Ceylon
G. Lewis 1910-320” (BMNH). Syntype {nano)'. 1 S
“Homaloplia nana Ceyl. Dohrn/ nana Brsk. type/ Muse-
um Paris ex coll. R. Oberthur” (MNHN).
Additional material examined. 8 ex. “Dikaya
3.000^.200 ft. 21. L-7.II.82/ Ceylon G Lewis 1910-320”
(BMNH), 1 ex. “Dikaya 3.000-4.200 ft. 21.L-7.II.82/
Ceylon G Lewis 1910-320/ Determined from description
G. J. A. Serica maculifera Brsk.” (BMNH), 1 ex. “Hak-
gala Ceylon 26-VIII-29/ Ceylon B.M. 1929-370”
(BMNH), 3 ex. “Ohiya Ceylon IV-29” (BMNH), 1 ex.
“Ohiya Ceylon 6500‘ [feet] 30-IV-28/ Ceylon B.M.
1929-370” (BMNH), 7 ex. “S Sri Lanka 17.-19.III.1994
Polhera - on light 3 km W of Matara Z. Kejval Igt.” (CP),
1 ex. ($)”Belihul-Oya Ceylon 2^^ trin 89 1. Z. Kannegie-
ter” (ZMHB), 4 ex. “Ceylon Nietn./ Zool. Inst. Mus.
Greifswald Coll. Gerstaecker” (EMAU), 1 ex. ($) ’’Cey-
lon, Centr. Prov. Horton Plains, Alt. 7000 ft. 12 mis SSE
Nuwara-Eliya 19.111.62 Eoc. 163/ Eund University Cey-
lon Expedition 1962 Brinck-Andersson-Cederholm ”
(MZEU), 2 ex. “Ceylon N. E. Dist. Kanda-ela Reservoir
5. 6mi SWNuwara Eliya, 6200 ft. 10-21 Feb. 1970 Da-
vis & Rowe” (USNM), 1 ex. “Sri Eanka: N. E. Dist. Kan-
de-ela 2 June 1975 S. E. Wood & J. E. Petty” (USNM),
1 S “Nuwara-Eliya E. Simon 1892/ Slg. R. Oberthur
(Coll. C. Martin) Eing. Nr. 4, 1956” (ZFMK), 1 $ “Cey-
lon/ variolosa Ceylan/ Coll. Kraatz/ Serica maculifera
Brsk. J. Moser det. 1917.” (DEIC), 1 $ “Ceylon Cande-
ze/ Coll. Haag/ Coll. Metzler/ Serica maculifera Brs./
Moser det.” (DEIC), 1 $ “Ceylon/ Coll. Metzler/ Moser
det.” (DEIC).
Redescription of syntype 1 {lurida). Eength: 5.5 mm,
length of elytra: 3.8 mm, width: 3.2 mm. Body moderate-
ly oblong, suboval, including antenna and legs reddish
brown, head and pronotum with strong green metallic
shine, elytra with weak green metallic shine, without dark
spots, dorsal surface including entire head dull, nearly
glabrous.
Eabroclypeus moderately wide, widest at base, lateral
margins convexly convergent toward broadly rounded an-
terior angles, margins moderately reflexed, anterior mar-
gin weakly sinuate medially; surface slightly elevated me-
dially and dull, finely and very densely punctate, with a
few long, sparsely scattered, erect setae; frontoclypeal su-
ture slightly elevate and weakly curved; smooth area an-
terior to eye 2.5 times as wide as long, flat; ocular can-
thus short but slender (ca. 1/4 of ocular diameter), smooth,
without a terminal seta. Frons dull, with fine, dense punc-
tures, with minute setae in punctures, with a few setae be-
side interior margin of eyes and at middle. Eyes moder-
ately large, ratio of diameter/ interocular width: 0.6. An-
tenna with ten antennomeres; antennal club in both sex-
es with three antennomeres, in male slightly longer than
remaining antennomeres combined. Mentum convexly el-
evated anteriorly.
Pronotum moderately transverse, widest at base, later-
al margins evenly convex, moderately convergent anteri-
orly, behind anterior angles convex; anterior angles pro-
duced and moderately acute, posterior angles almost
right-angled, moderately rounded at the tip; anterior mar-
gin convexly produced medially, without a fine marginal
line, basal margin without marginal line; surface with
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Fig. 1. (A-B) Periserica picta Brenske (lectotype), (C-E) P. subsignata (Walker) (syntype of P. picta, ZMHB), (F-H) P. ful-
vostriata Brenske (lectotype), (I, J) Periserica interrupta (Walker, 1859) (lectotypeX (K-M) P triflahellata sp. n. (holotype), (N-P)
P. densipunctata sp. n. (holotype). (A, C, F, I, K, N) Aedeagus, left side lateral view; (B, E, H, M, P) aedeagus, left side lateral
view; (D, G, J, F, O) parameres, dorsal view. Scale bar: 1 mm.
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Fig. 3. (A-C) Selaserica distincticornis Brenske (syntype), (D-F) Se. implicata (^renske) (syntype), (G-I) Se. convexiuscula
sp. n., (J-L) Se. impexa sp. n., (M-0) Se. padukkana sp. n., (P-R) Se. kanneliyana sp. n. (A, D, G, J, M, P) Aedeagus, left side
lateral view; (C, F, I, L, O, R) aedeagus, left side lateral view; (B, E, H, K, N, Q) parameres, dorsal view Scale bar: 1 mm.
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12:41 Seite104
Fig. 4. (A-C) Selaserica pusilla Arrow (syntype), (D-F) splendifica (Brenske) (syntype), (G-I) sericea Arrow (holo-
type), (J-L) Se. nuwarana sp. n., (M-0) Se. praetexta sp. n., (P-R) Se. sinharajana sp. n. (A, D, G, J, M, P) Aedeagus, left side
lateral view; (C, F, I, L, O, R) aedeagus, left side lateral view; (B, E, H, K, N, Q) parameres, dorsal view Scale bar: 1 mm.
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Sericini of Sri Lanka
105
Fig. 5. (A-C) Selaserica wilpattuensis sp. n., (D-F) Se. meridionalis sp. n., (G-I) Neoserica sexfoliata Moser (syntype), (J-L)
N. kalaarensis sp. n., (M-0) N. bombycina (Karsch) (syntype Neoserica quadrijlabellata), (P-R) Maladera calcarata (Brenske)
(Lectotype). (A, D, G, J, M, P) Aedeagus, left side lateral view; (C, F, 1, L, O, R) aedeagus, left side lateral view; (B, E, H, K, N,
Q) parameres, dorsal view. Scale bar: 1 mm.
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Fig. 6. (A-C) Maladera breviata (Brenske) (syntype), (D-F) M. breviatella sp. n. (holotype), (G-1) M. rotundata (Walker) (holo-
type Autoserica srilanka Frey), (J-L) M. iuga sp. n. (A, D, G, J) Aedeagus, left side lateral view; (C, F, 1, L) aedeagus, left side
lateral view; (B, E, H, K) parameres, dorsal view. Scale bar: 1 mm.
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107
Fig. 7. (A-C) Maladera mollis (Walker) (Sri Lanka: Marawila), (D-F) M. pseudomollis sp. n. (holotype), (G-I) M carinifrons
(Brenske) (syntype), (J-L) M. westermanni (Brenske), (M-0) M. krombeini sp. n. (holotype), (P-R) M. ekisi sp. n. (holotype),
(S-U) M cinnaberina (Brenske) (syntype), (V-X) M rufocuprea (Blanchard) (S. India Coimbatore). (A, D, G, J, M, P, S, V) Aedea-
gus, left side lateral view; (C, F, I, L, O, R, U, X) aedeagus, left side lateral view; (B, E, H, K, N, Q, T, W) parameres, dorsal view.
Scale bar: 1 mm.
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Dirk Ahrens
Fig. 8. (A-C) Maladera fistulosa (Brenske) (lectotype), (D-F) M dubia (Arrow) (holotype), (G-I) M. weligamana (Brenske)
(syntypQ Autoserica fistulosa, Ceylon), (J-L) M coxalis (Moser) (syntype). (A, D, G, J) Aedeagus, left side lateral view; (C, F, I,
L) aedeagus, left side lateral view; (B, E, H, K) parameres, dorsal view. Scale bar: 1 mm.
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109
Sericini of Sri Lanka
Fig. 11. (A-C) Maladera kandyensis sp. n. (holotype), (D-F) M hastata sp. n. (holotype), (G-I) M lindulana sp.
(J-L) M. tricuspidata sp. n. (holotype), (M-0) M. hortonensis sp. n. (holotype), (P-R) M hadullana sp. n. (holoty
n. (holotype),
pe), (S-U) M
view; (C, F, I,
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Fig. 12. (A-C) Maladera bandarwelana sp. n. (holotype), (D-F) M. setosa (Brenske) (syntype, Konbir), (G-1) M. strata (Brenske)
(syntype Autoserica apogonoides Brenske), (J-L) M. brevis (syntype). (A, D, G, J) Aedeagus, left side lateral view; (C, F, 1, L)
aedeagus, left side lateral view; (B, E, H, K) parameres, dorsal view. Scale bar: 1 mm.
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113
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Fig. 14. Habitus. (A) Periserica picta Brenske, (B) R interrupta (Walker), (C) R fulvostriata Brenske, (D) Selaserica lucidicol-
lis sp. n., (E) Se. pusilla Arrow, (F) Se. nitida (Candeze), (G) Se. impexa sp. n., (H) Se. meridionalis sp. n., (I) Neoserica bombyci-
na (not to scale).
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Sericini of Sri Lanka
group and M. madurensis group (in part D).
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Fig. 18. Distribution of the Sericini species of Sri Lanka (as indicated by the legend): (A-C) Maladera subgenus Cephaloserica
and (D) Maladera fistulosa group (part).
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119
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Fig. 20. Distribution of the Sericini species of Sri Lanka (as indicated by the legend): (A) Maladera setosa group and (B) Seri-
ca (sensu lato). (C) Climatic zones (see text), their number of species (D) and their faunistic similarity based on the Jaccard Index
(agglomeration method: unweighted pair-group average) in Sericini (E).
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1 1 a iiiiiiiiiMiiiiii iiiiiiii I ill iiiiiii ill iiir 3.
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anderssoni sp. n. (Maladera) 59, 88, 110,
apogonoides Brenske, 1 898; syn
carinirostris BTQnskQ, 1896; syn
atrata Burmeister, 1855 (nec Reiche 1847); syn
atratula Dalla Torre, 1917; syn
badullana sp. n. {Maladera) 59, 69,
bandarwelana sp. n. {Maladera) 89,
bisornata sp. n. {Maladera) 59, 74,
bombycina (Karsch, 1882) {Neoserica) ..35, 97, 105,
breviata (Brenske, 1898) {Maladera) 39, 41, 44,
breviatella sp. n. {Maladera) 39, 43,
(Blanchard, USO) {Hemiserica) 91,
brincki sp. n. {Maladera) 59, 79,
buruensis Brenske, 1899, syn
calcarata (Brenske, 1898) {Maladera) ...39, 90, 105,
carinifrons (Brenske, 1896) {Maladera) 47, 49,
chalybaeaBxQxs&ks, 1898; syn
cinerea Brenske, 1898, syn
cinnaberina (Brenske, 1 898) {Maladera)
confirmata (Walker, 1859) {Selaserica ) ...
convexiuscula sp. n. {Selaserica)
coxalis (Moser, 1915) {Maladei
densipunctata sp. n. {Perisericc
distincticornis (Brenske, 1898) {Selaserica) ...13, 25,
diyalumana sp. n. {Maladera) 59, 77,
dubia (Arrow, 1916) {Maladera) 59, 66, 69, T.
ekisi sp. n. {Maladera) A1 , 57, 9^
Brenske, 1898; syn
fistulosa (Brenske, 1898) {Maladera) .58, 59, 62, 97,
flinti sp. n. {Maladera) 59, 86,
fulvostriata Brenske, 1898 {Periserica)
... Al, 54
....13, 17
....13, 24
....59, 60, 62, 64
ica)
fusa Brenske, 1898 {Serica)
hastata sp. n. {Maladera)
heveli sp. n. {Maladera)
hortonensis sp. n. {Maladera) .
zmmMw/to Brenske, 1898; syn. .
impexa sp. n. {Selaserica)
implicata (Brenske, 1 898) {Sel
interrupta (Walker, 1859) {Pen
iuga sp. n. {Maladera)
kalaarensis sp. n. {Neoserica) .
kandyensis sp. n. {Maladera) ..
kanneliyana sp. n. {Selaserica)
kitulgalana sp. n. {Serica)
knucklensis sp. n. {Selaserica)
krombeini sp. n. {Maladera) ....
kuruwitana sp. n. {Maladera) .
Imdulana sp. n. {Maladera) ....
lucidicollis sp. n. {Selaserica) .
92, 97, 113
5, 9
59, 83, 85
59, 78, 80
....58, 67, 70,
13,23, 103,
serica) ....\3, 19, 23,
erica) 5, 11, 101,
39, 46,
35, 38,
59, 84, 86,
13,21,
92, 96,
13, 16,21,
47, 56, 58
..59, 62, 63, 66,
59, 71,
13,20,25, 702
urida Brenske, 1898 {Serica) 92, 94, 96, 97, 113, 120
naculicauda Arrow, 1916 {Serica) 13, 18, 98, 102, 116
naculifera Brenske, 1898 (nec Brenske, 1894); syn 94, 97
navilluensis sp. n. {Maladera) 59, 85, 87, 770, 119
teridionalis sp. n. {Selaserica).... \3, 32, 33, 97, 105, 114, 116
nollis (Walker, 1859) {Maladera) 47, 50, 54, 57, 707, 118
^ana Brenske, 1898 {Serica) 92, 94, 95, 97, 113, 115, i:
7a(Candeze, \^6\) {Selaserica) ...
nilaveliensis sp. n. {Malader
padukkana sp. n. {Selasericc
pavieana Brenske, 1 899; syn.
...13, 14, 16,
28, 702, 114,
59, 80, 770,
14,27, 104,
...13, 22,
17 ,
7 poyagana sp. n. {Maladera) .
9 praetexta sp. n. {Selaserica) .
8 pseudomollis sp. n {Maladera)
;2 pubescens (Arrow, 1916) {Maladei
1
8 pusilla Arrow, 1916 {Selaserica) . .
i) 13, 22, 24, 103, 116
91
5, 6, 7, 8, 9,
10, 707, 114, 116
59, 70, 109
14, 29,31,704, 116
47, 52, 707, 118
59, 72, 75, 76,
84, 770, 119
14,27,28, 30,
32, 104, 114, 116
quadriflabellata {BrmskQ, 1896); syn 35, 36, 705
romanoi sp. n. {Maladera) 59, 87, 88, 89, 110, 119
rotundata (Walker, 1859) {Maladera) 39, 44, 47, 106, 117
rubescens Arrow, 1916; syn 94
rufocuprea (Blanchard, 1850) {Maladera) 47, 52, 97,
707, 115, 118
schintlmeisteri sp. n. {Maladera) 59, 64, 66, 109, 119
scutellaris Arrow, 1916 {Selaserica) 1 3, 15
sericata Brenske, 1898; syn. 35
sericea Arrow, 1916 {Selaserica)
(Brenske, n96) {Maladera) ...
ta Mose
1915 {Neose
’ca)
14, 33, 104, IK
39, 73, 90, 97
772, 115, 12(
35, 36, 705, 111
.49, 50, 5:
14, 30, 704, IK
13, 14, 26,
srilanka Frey, 1974; syn
straba (Brenske, 1898) {Maladera)
stupida Brenske, 1900; syn. .
subsignata (Walker, 1859) {Periserica) 5, 6, 8, 9, 7
sumatrensis Bronsko, 1898; syn
tricuspidata sp. n. {Maladera) 59, 75, 77, 81, 7
truncatus Mittal, 1976; syn. .
uggalkaltotaensis sp. n. {Maladera) 59, 76, 7
vflno7o.sfl Motschulsky, 1863 {Serica)
ventralis WdXkor, 1859; syn. ..
;nske, 1900) {Maladera) .59, 60, 61, 108,
termanni (Brenske, 1898) {Maladera) 47, 5
attuensis sp. n. {Selaserica) 13, 32, 705,
di sp. n. {Maladera) 59, 65, 109,
lensis sp. n {Maladera) 59, 81, 110,
Bonn zoological Bulletin - Supplementum 61: 1-124
©ZFMK
Bonn zoological Bulletin - Supplementum Vol. 61 (2014)
Managing Editor: Thomas Wesener
Zoologisches Forschungsmuseum Alexander Koenig -
Leibniz-lnstitut fur Biodiversitat der Tiere (ZFMK)
Adenaueralllee 160, D-53113 Bonn, Germany
ISBN-13: 978-3-925382-65-9
ISSN: 0302-671X
Back-Cover illustration:
Male genitalia of Maladera tricuspidata sp. n.: newly developed processes of the aedeagus replace
functionally the reduced parameres.
Cover illustration:
Habitus of Selaserica impexa sp. n. in front of the map of the examined material from Sri Lanka.
Leibniz-Gemeinschaft
Bundesministerium
SorsSung Wissenschaft und Forschung
Ministerium fiir Innovation,
des Landes Nordrhein-Westfalen
Bonn zoological Bulletin - Supplementum Volume 61 (2014)