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BRITISH MUSEUM (NATURAL HISTCRY)
INSECTS OF SAMOA
AND OTHER SAMOAN TERRESTRIAL
ARTHROPODA
PART II. HEMIPTERA
FASC. 2. Pp. 47-80
CERCOPIDAE
By V. LALLEMAND, M.D.
CICADIDAE
By J. G. MYERS, Sc.D.
AQUATIC AND SEMI-AQUATIC HETEROPTERA
By PROF. TEISO ESAKI
WITH THIRTY-EIGHT TEXT-FIGURES
LONDON :
_ PRINTED BY ORDER OF THE TRUSTEES OF THE BRITISH MUSEUM
SOLD AT
Tur Brivish Museum (Naturan Hisvory), CRomwrLL Roan, S.W.7
AND BY
B. Quaritcu, Lrp,; Dotav & Co., Lrp.; Tax Oxrorp University Press; anp
Wuetpon & Wustry, Lrp,, Lonpon; atso py Oxiver & Boyp, EpinsurcH
1928
L3sued 23rd June, 1928.] : [Price Tro Shillings and Sixpence.
INSECTS OF SAMOA
AND OTHER SAMOAN TERRESTRIAL
ARTHROPODA
Although a monograph, or series of papers, dealing comprehensively with —
the land arthropod fauna of any group of islands in the South Pacific may be
expected to yield valuable results, in connection with distribution, modification
_ due ito isolation, and other problems, no such work is at present in existence. oe
In order in some measure to remedy this deficiency, and in view of benefits
directly accruing to the National Collections, the Trustees of the British
Museum have undertaken the publication of an account of the Insects and other
Terrestrial. Arthropoda collected in the Samoan Islands, in 1924-1925, by
Messrs. P. A. Buxton and G. H. E. Hopkins, during the Expedition of the
London School of Hygiene and Tropical Medicine to the South Pacific.
Advaritage has been taken of the opportunity thus afforded, to make the studies © 4
as complete as possible by including in them all Samoan material of the groups
concerned in both the British Museum (Natural History) and (by courtesy . 7
the authorities of that institution) the Bishop Museum, Honolulu. |
le ic not wtended Ghet contutore te une ce shall i contac to de :
Museum Staff or to any one nation, but, so far as possible, the assistance of the
leading authorities on all groups to be dealt with has been obtained.
The work will be divided into igh’ ‘Parts ” {see p. 3 of wrapper), which :
will be subdivided ito “Fascicles.”’ Each of the latter, which will appear as
ready in any order, will consist of one or more contributions. On the
completion of the work it is intended to issue a general survey, summarising
the whole and drawing from it such conclusions as may be warranted.
A list of Fascicles already issued will be found on the back of this cee
| E. E. AUSTEN,
Keeper of Entomology. Aa
British Museum (Naturat History),
Cromwe.t Roap, S.W.7.
INSECTS OF SAMOA |
AND OTHER. SAMOAN TERRESTRIAL
ARTHROPODA |
Although a monograph, or series of Sapele desing comprehensively with :
the land arthropod fauna of any group of islands in the South Pacific may be -
expected to yield valuable results, i in connection with distribution, modification ©
_ due to isolation, and other problems, ne such work is at present in “existence. :
_In order in some measure to remedy this deficiency, and in view of benefits
directly accrumg to the National Collections, the Trustees of the British —
Museum have undertaken the publication of an account of the Insects and other —
‘Terrestrial. Arthropoda collected in the Samoan Islands, in 1924- 1925, by :
Messrs. P. A. Buxton and ey H. E. Hopkins, during the Expedition of the —
London School of Hygiene and Tropical Medicine to the South Pacific.
Advaritage has been taken of the opportunity thus afforded, to make the studies
as complete as possible: by including in them all Samoan material of the. groups.
concerned in both the British Museum (Natural History) and (by conti 6 e ‘
the authorities of that institution) the Bishop Museum, Honolulu. :
It is not intended that contributors to. the text shall Me eonkined t to ie
Museum Staff or to any one nation, but, so far as possible, the assistance of the
leading authorities on all groups: to be dealt with has ao obtained.
The work will He ede cight “Parts (see p. 3 of ween aiiich
will be subdivided into “‘Fascicles.” Each of the latter, which will appear as
ready in any order, will consist of one or more contributions. On the
completion of the work it is intended to issue a general survey, ee ;
the whole and drawing from it such conclusions as may be warranted. :
A list of Fascicles already issued will be found on the Sade of this wrapper.
EE AUSTEN,
Keeper of pap
British Museum (Natura Hisrory), thas
CromwEL.L Roap, S.W.7.
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INSHCPS OF SAMOA
Parr II, Fasc, 2
HEMIPTERA
CERCOPIDAE
Par V. LALLEMAND, M.D., Uccle, Belgique
(Avec 10 Texte-figures)
I. INTRODUCTION
Le genre Clovia est répandu en Afrique, en Asie et en Océanie a peu prés
entre les latitudes 30° N. et 8.; les espéces sont nombreuses en Océanie.
Malheureusement les homoptéres et spécialement certains groupes, dont les
Cercopides, ont été peu recherchés dans plusieurs de ces iles et les rapports
entr’elles des différentes espéces de ces régions sont encore assez difficiles a
établir. Une de celles ci-dessous décrites C. guddi est voisine de C. insignis
Distant, recueillie sur Vile Elisabeth (ou Henderson), de C. oceanica Jacobi,
trouvée dans les iles Tonga, et également de deux autres (C. rapana de Vile Rapa
et C. soceabilis de Vile Tahiti) * ; ces quatre Clovia constituent un groupe bien
distinct, ce sont probablement quatre formes locales d’ une seule et méme souche.
Le genre Perinoia est propre a VOcéanie, la P. caput-ranae (= birarensis
Kirkaldy) a été trouvée en Nouvelle-Guinée, Nouvelle-Bretagne et enfin dans les
iles Samoa.
Le genre Aufidus est également spécial & l’Océanie ; la plupart des espéces
décrites proviennent de la Nouvelle-Guinée, qui parait étre le centre de disper-
sion ; quelques unes ont été recueillies dans les iles environnantes 4 Aroé, Birara
(Nouvelle-Bretagne), une sur l’ile Gilolo (Moluques), et enfin deux autres, pro-
venant du Queensland, ont été décrites par Kirkaldy ; pour ces derniéres, il
* Ann. Mag. Nat. Hist. (10) I, p. 634, 1928.
II, 2 1
an
48 INSECTS OF SAMOA.
avait eréé les genres Aufidellus et Aufiterna (Rep. Exp. Stat. Haw. Sug. Plant.
Assoc., Vol. 9, pp. 381 et 382, 1906). Elles ont été rapportées au genre Aufidus
par Distant (Rhynchota malayana, Vol. 1, part 1, p. 132, 1908).
Il. Les ESPECES RECUEILLES DANS LES ILES SAMOA.
APHROPHORINAE.
1. Clovia swezeyi, sp. n. Fig. 1.
29. Noire ou brun-noir; sur les cotés du front, des stries transversales
jaunatres ; sur le vertex de chaque cété, une ligne partant de l’angle postérieur
de la partie frontale, longeant l’oeil et s’étendant jusqu’au bord postérieur et
derriére elle, sur le pronotum, une petite
tache, jaune-brunatre ; tiers basal du
clavus, ainsi que sur le corium, une
petite tache longitudinale prés de la
suture et sur le méme plan au bord
externe une autre en forme de trapéze
(ces trois taches formant une bande
transversale), enfin a la partie apicale,
partant du bord externe, une tache
allongée s’étendant jusqu’ entre les
branches du radius, jaune-clair; ex-
trémité du clavus et partie adjacente
du corium d’un jaune légérement brun ;
la partie apicale des élytres est plus
Texte-Fic. 1.—Clovia swezeyi, sp. n. claire et plus ou moins transparente ;
alles enfumées, a nervures noires; sur
le pro- et mesosternum, partant de l’oeil, une bande jaune; rostre, pattes,
hanches, jaune-brunatre ; tibias postérieurs plus foncés; extrémité du rostre,
des tarses et des épines, noire; la couleur noire du vertex peut se muer en un
brun plus ou moins foneé.
Vertex en angle aigu, a extrémité arrondie, aussi long que large entre les
yeux ; ocelles a peu prés a égale distance l’un de l'autre et des yeux ; pronotum
et écusson assez densément ponctués en stries transversales, sur le premier,
une trace de caréne longitudinale ; deux épines sur les tibias postérieurs. Cette
espéece, comme les suivantes, a les nervures saillantes spécialement les apicales ;
le réseau apical est composé de cing cellules.
CERCOPIDAE. 49
Longueur—10-5 mm.
Habitat—lle Savaii, Safune, mai (Swezey) ; [le Tutuila, Leone, juillet(Bryan).
Je dédie cette espéce a Mr. Swezey qui l’a récoltée dans lile Savaii.
2. Clovia oceanica Jacobi.
Arch. f. Naturges. Jahrg. 87, Heft 12, p. 14, 1921.
Tonga (type) ; Samoa (Jacobi).
3. Clovia juddi, sp. n. Fig. 2.
35,492. Noire; vertex (sauf une bande postérieure), écusson, une bande
longitudinale s’élargissant légérement en arriére le long du bord externe et
extrémité du clavus, jaunes; sont d’un_ blanc-
jaunatre, transparentes, les taches suivantes: sur
le corium, une prés de l’extrémité du clavus, en
face de celle-ci, partant du bord externe une
deuxiéme de forme allongée, se dirigeant en arriére
et en dedans, et se terminant en pointe, enfin prés
du bord apical deux autres plus petites (sur cer-
tains exemplaires ces derniéres, ainsi que celles de
Vextrémité du clavus peuvent disparaitre) ; front
jaune, ayant 6 a 7 stries transversales latérales et = Texrx-ric. 2.—Clovia juddi,
en dessous de celles-ci les cdtés bruns. Espéce trés shea:
voisine de C. insignis Dist., dont elle se différencie par le dessin du pronotum et
des élytres, et de C. oceanica Jacobi, dont elle se distingue principalement par
l’écusson jaune.
Longueur—g, 65 mm.; 97 a7-5 mm.
Habitat—lles Manua ; Ofu et Tau, février et septembre (Judd, Swezey et
Wilder).
Je dédie cette espéce & Mr. Judd, un de ceux qui en ont récolté des
exemplaires.
4. Clovia biformis, sp. n. Fig. 3.
g. Téte, écusson, sternum, abdomen, jaunes; élytres jaunes a la base,
plus pales ensuite et plus ou moins transparents, recouverts d’une villosité
blanche, finement bordés de brun au bord apical, deux ou trois petites nervures,
situées prés de celui-ci plus ou moins brunes ; sont noirs, le pronotum, 6 lignes
50
INSECTS OF SAMOA.
transversales interrompues & la partie médiane du front, l’extrémité du rostre,
des tarses et des épines ; yeux bruns tachés de noir.
? différe par le pronotum jaune, le bord apical et les nervures @ peine bruns.
Vertex en angle obtus, sa longeur égale environ les trois quarts de sa largeur
au bord postérieur ; ocelles égérement plus prés l’un de Vautre que des yeux ;
TEXTE-FIG 3.—Clovia biformis, sp. n.: A, male; B, femelle.
pronotum ponctué en stries plus ou moins transversales; derriére son bord
antérieur, une série de fossettes formant une ligne paralléle a celui-ci; deux
épines sur les tibias postérieurs.
Longqueur—9 rom.
Habitat—9 3 et 6 9, recueillis entre 100 et 400 m. d’altitude durant les mois
TEXTE-FIG. 4.—Clovia bryant, sp. n.
de janvier, avril, juin, juillet, aout et septembre,
sur les herbes dans Vile Tutuila: a Pago Pago
et dans le centre; et Vile Upolu: a Malolo-
lelei.
5. Clovia bryani, sp. n. Fig. 4.
2 3. Orange; yeux gris-brun fortement
tachés de noir; bords latéro-antérieurs du
pronotum noirs ; petites nervures apicales plus
ou moins brunes; alles enfumées a nervures
noires ; face supérieure de l’abdomen et bords
de sa face inférieure, extrémité du rostre, des
tarses et des épines, brun-noir; surface supérieure du corps recouverte d’une
villosité argentée.
CERCOPIDAE. 51
Front légérement aplati, 4 stries latérales assez légéres; vertex a bord
antérieur en angle obtus, arrondi, & surface déclive en avant, sa longeur égale &
peu prés les trois-quarts de sa largeur au bord postérieur ; deux épines sur les
tibias postérieurs.
Longueur—7-5 mam.
Habitat—Ile Sava: Salailua, mai (Bryan).
Je dédie cette espéce & celui qui l’a récoltée, Mr. K. H. Bryan.
6. Clovia armstrongi, sp. n. Fig. 5.
2. Jaune-orange, recouverte d’un duvet blanchatre ; élytres oranges et
opaques sur le clavus et le long de la moitié antérieure du bord externe entre
celui-ci et le radius, blanc-jaunatre et trans-
parents sur le restant; six stries latérales,
transversales sur le front, une fine bande
longitudinale sur le radius depuis la base
jusqu’un peu au dela du tiers basal, bord
externe, des nervures apicales, bord apical,
bord interne trés finement, extrémité des
rostre, des épines et des tarses, brun-noir.
Ces différentes taches a |’exception des lignes
du front peuvent plus ou moins seffacer.
Abdomen brun-clair ; sur le prosternum, au
niveau de la base des ailes, une tache brun- [yyep-pe. 5.—Clovia armstrongt, sp. n.
foncé.
Vertex long, en angle aigu, 4 extrémité arrondie ; sur le pronotum une série
de petites fossettes allongées, formant une ligne paralléle au bord antérieur ;
deux épines sur les tibias postérieurs ; front relativement assez aplati.
Longueur—8 mm.
Habitat—Ile Upolu: Apia; Siumu ; recueillis en aoait et novembre.
Je dédie cette espéce a Dr. J. 8. Armstrong, qui l’a capturé.
7. Clovia buxtoni, sp. n. Fig. 6.
13. Blanc-jaunatre, & téte égérement plus jaune ; huit stries transversales,
latérales du front, yeux et sur les élytres, une large bande transversale, médiane
ainsi qu’un petit triangle prés de la base entre la suture et le radius, noirs ;
extrémité du rostre et des tarses, bande longitudinale médiane du vertex, nervures
52 INSECTS OF SAMOA.
apicales et bord apical jusque la pote du clavus, abdomen, d’un brun plus ou
moins foncé ; partie apicale des élytres transparente ; ailes hyalines, a nervures
brunes. Partie médiane du front lisse, sur les cétés, des stries transversales ;
rostre s étendant entre les hanchés médianes ; yeux assez gros, saillants ; ocelles
petits, légérement plus prés un de l'autre que des yeux; vertex aplati, a peu
prés aussi long que large au bord postérieur entre les yeux ; pronotum ponctué
. en stries transversales, 4 faible sillon médian, longitudinal ; deux épines sur les
tibias postérieurs.
Longueur—6-5 ; élytres, longueur: 5 mm., largeur: 1-5 mm.
Habitat—Ile Upolu: Vailima, recueilli en décembre.
. Je dédie cette espéce a celui qui la récoltée et a rassemblé la collection
présente des homopteéres des iles Samoa.
TrexteE-FIc. 6.—Clovia buxtoni, sp. n. TEXTE-FIG. 7.—Clovia navigans Jacobi.
Led
8. Clovia navigans Jacobi. Fig. 7.
Arch. f. Naturges. Jahrg. 87, Heft 12, p. 15, 1921.
Habitat—lle Upolu: Apia; Mulifanua; Vailutai; Falelatai; Lalomanu.
Tle Savaii: Tuasivi; Salailua; Fagamalo; Safune. Ile Tutuila: Pago Pago ;
Amauli.
Hspéce répandue de préférence, semble-t-il, non loin des cotes.
9. Perinoia caput-ranae Le Guillon.
Rev. Zool., Vol. 4, p. 262, 1841 (Aphrophora caput-ranae).
Samoa; Nouvelle-Guinée ; Nouvelle-Bretagne (Jacobi).
CERCOPIDAE od
TOMASPIDINAE.
10. Aufidus hopkinsi, sp. n. Fig. 8.
1 3. Partie supérieure du corps noir-brun, brillante, recouverte d’un duvet
brun ; partie antérieure du vertex brune; ailes brun-noir, plus foncées vers la
partie apicale; front, clypeus, rostre, pattes, jaunes; extrémité des tarses et
des épines, noire; abdomen brun; élytres, a base légérement plus brune, a
extrémité brun-clair, transparente, montrant en arriére du milieu une grande
tache blanche, hyaline, s’étendant du bord externe au médian.
Ocelles trés proches l'un de l’autre; sur le pronotum, ponctué en stries
transversales, une fine caréne médiane ; deux épines sur les tibias postérieurs.
Longueur—s8 mm.
Habitat—lle Upolu: Malololelei; recueilli en juin par P. A. Buxton et
G. H. E. Hopkins.
Je dédie cette espéce a ce dernier.
TEXTE-FIG. 8.—Aufidus hopkinst, sp. n. TEXTE-FIG. 9.—Aufidus samoanus, sp. D.,
2
forme typique.
11. Aufidus samoanus, sp. n. Figs. 9 et 10.
26 g et 47 9. Orange ou quelquefois brun-orange ; abdomen plus foncé ;
élytres plus clairs, transparents spécialement dans la partie apicale; ocelles,
yeux, extrémité des tarses, épines et sur les élytres une bande et une tache,
noir-brun ; la bande des élytres transversale, située & peu prés au milieu, part
du bord externe et s’étend plus ou moins loin vers l’intérieur, elle peut méme
sur certains exemplaires atteindre le bord interne et alors est ondulée, sur d’autres
elle est trés courte ; la tache part de l’extrémité du bord externe et s’étend plus
54
INSECTS OF SAMOA.
ou moins loin dans la direction de la pointe du clavus ; sur certains spécimens
bien marqués, la pointe du clavus est brune, la tache apicale peut s’allonger vers
celle-ci et méme l’atteindre ; de méme, les bords postérieurs du vertex et du
pronotum peuvent étre bruns aussi; partie supérieure du corps recouverte d'une
villosité rousse.
TEXTE-FIG. 10.—Aufidus samoanus
var. wmmaculata (A) ;
obscura (B).
Longueur—6-5 & 7 mm.
Sillon frontal large et relativement peu profond, commencant
non loin du bord antérieur de la téte, celui-ci
est en angle obtus; ocelles trés proches lun de
Vautre ; milieu de l’écusson creusé en fossette
longitudinale ; une épine sur les tibias postérieurs.
Var. unmaculata : aucune tache a la face
supérieure du corps.
Var. obscura: Vertex, pronotum, moitié
basale et tache apicale des élytres brun-noir ; le
bord postérieur de la partie brune n’est pas droit
mais légérement ondulé comme la bande sur la
forme type ; la coloration orange primitive peut
apparaitre par places, a la partie antérieure du
vertex et du pronotum, ainsi qua la base des
élytres.
Habitat—Ue Savai: Tuasivi; Salailua; Safune. Ile Upolu: Malololelei ;
Vaea; Apia; Vailima; Mulifanua; Falelatai; Siumu; Aleipata. [le Tutuila:
Amauli; Afono; Fagasa; Leone; Pago Pago, et le centre de Iile.
Var. ammaculata: Ile Savaii: Safune. Ile Tutuila: Pago Pago.
Var. obscura: Ile Upolu: Siumu; Malololelei; Vaea. Ile Savai: Safune.
Recueillis durant toute année, sauf pendant les mois de janvier et d’octobre.
Texte-fig.
HE oo be
or D
pa
oo
LISTE DES FIGURES.
Clovia swezeyt, sp. n.
Clovia juddi, sp. n.
. Clovia biformis, sp. n.: A, male, B, femelle.
Clovia bryant, sp. n.
Clovia armstrongi, sp. n.
Clovia buxtoni, sp. n.
Clovia navigans Jacobi.
Aufidus hopkinsi, sp. n.
9. dufidus samoanus, sp. n., forme typique.
. Aufidus samoanus var. immaculata (A) ; var. obscura (B).
CICADIDAE
By J. G. Myers, Sc.D.
(With 22 Text-figures.)
THE cicadas seem to be represented in the Samoan Islands by one abundant
and widespread species, and by two others which, judging from the collections
examined, are very rare. Baeturia exhausta Guér. is essentially Austro-Malayan
in distribution, and Samoa would appear to be its eastern limit, which it has
reached by a route north of Fiji, in which group it does not occur. Tibicen
kuruduadua Dist. occurs elsewhere only in Fiji, while the third species belongs
to a remarkable new endemic genus—the first peculiarly Polynesian genus of
cicadas.* This latter has Austro-Malayan affinities, but is strongly isolated.
It is thus apparent that the evidence of the cicadas, so far as it goes, corroborates
the general conclusion drawn by Muir from a study of the Fulgoroids, “ that the
Samoan Islands form an outpost of the Polynesian plateau.”
I am deeply indebted to Mr. A. J. E. Terzi for the beautiful drawing of the
_ venation which illustrates the new genus; and to Mr. W. E. China for many
courtesies at the British Museum.
Subfamily PLATYPLEURINAE Handl.
Genus Trpicen Latr. 1825
Type Cicada plebera Scop.
1. Tibicen kuruduadua Dist.
Cicada kuruduadua Distant, Trans. Ent. Soc. London (1881), p. 645, 1881; Syn. Cat. Hom.
Crcad., p. 39, 1906.
Upolu: Malololelei, 1 9, 30.viii.1925 (Armstrong).
In the absence of a male, this seems indistinguishable, save by paler colour,
from 7’. kuruduadua, hitherto known only from Fiji. As a matter of fact this
* Dasypsaltria Haupt. is synonymous with Platypleura, and is not Samoan. See below.
5D
56 INSECTS OF SAMOA.
female looks more truly congeneric with the type than do the females placed
with the latter in the British Museum.
Structurally, except in the greater length of the teemina, this species
resembles 7. (Cicada) plebeia Scop., with which it may be considered strictly
congeneric. It certainly does not fall into any of the segregates—Diceroprocta
Stal, Chremistica Stal, Rihana Distant—which it has been proposed to separate
from the genus of which C. plebeia Scop. is the type. With regard to this genus
I cannot accept Horvath’s (Ann. Hist. Nat. Mus. Hung., 23, pp. 93-98, 1926)
new name Lyristes. The name Trbicen was first proposed in 1825, by Latreille
(Fam. nat. Regne Anim., p. 426) with C. plebeia as the only species mentioned.
* Tibicen ” is not a French word, as Horvath claims, nor is there any excuse for
considering it, as Horvath does, a nomen nudum, since C. plebera, given as its
example, is still and was then, the best-known cicada in the world. I therefore
follow Van Duzee in considering C. plebeva Scop. as the type of the genus Tibicen
Latr. (= Cicada auctt. nec Linn.*). This course creates, in the subfamily
nomenclature, a difficulty which Van Duzee (Cat. Hemipt. North Amer., pp. 488,
498, 1917) has not met by calling the subfamily based on Trbicen Latr. Tibi-
ceninae, and the other, founded on Tibicina Amyot, Tibicininae. Formed
correctly from Trbicen and from Tibicina respectively, by adding -inae to the
stem, the two subfamily names would be identical. I have therefore followed
Handlirsch (a Schroder, Handb. d. Entom., Bd. 3, pp. 1115-1117, 1925) in calling
the former subfamily Platypleurinae, after another well-known, old and repre-
sentative genus.
Moana,t ecm.
Head, including eyes, much narrower than base of mesonotum, head pro-
duced conically half its length in front of eyes. Frons elongate, considerably
swollen basally, smooth and shining, the cross striae indistinct. Antennal scrobes
prominent. Ocelli almost contiguous, the median ocellus directed straight
forward. Pronotum apically the widest part of body, basally greatly con-
stricted, so that the margins, which are slightly expanded at the apical angles,
run very obliquely to the head. Sulci of vertex and pronotum deeply marked.
* Since C. plebeia Scop. was not among the originally included species of Cicada Linn. 1758,
it obviously cannot be the genotype. The genotype now generally recognised is C. orm Linn.
+ Moana—Samoan name for the ocean—thus appropriate for a cicadid genus, endemic in
Oceania.
CICADIDAE. 57
Mesonotum twice the length of pronotum; basal three-quarters domed and
swollen; cruciform elevation relatively depressed, little developed, much con-
stricted laterally. Abdomen small, conical, four-thirds length of mesonotum,
than which it is almost wholly narrower, giving whole head and trunk a general
fusiform shape. Tymbal covers complete, consisting of a somewhat narrow
tongue reaching antero-laterally from each side of 2nd segment. Legs long
and slender. Meracanthus long and slender, and hollowed ventrally.
Tegmina unusually expansive, much longer than body, with wings also,
cross-ribbed at wide intervals, especially in the huge cell Cu;. Stem of M
extraordinarily thin and weak, leaving basal cell practically contiguous with
TExt-FIic. 1.—Moana expansa g. et sp. n., tegmen. A. J. EH. Terzi del.
that of Cu;. Cu, very stout, rib-like and compressed almost at right angles to
the plane of the tegmen, strongly arched towards costal margin. A, very
stout, practically contiguous with anal margin, which is itself much thickened,
and bent outwards, so that clavus is wider than usual. Nodal line barely
discernible on membrane. Hind wings normal, with six apical cells. Pygophor
with no median dorsal spine, but two lateral projections directed caudad, as in
Sawda, Fatima, Cosmopsaltria and related genera, but shorter. (Rest of the
pygophor missing in the case of the type.) Curiously enough in the last nymph
of same sex the pygophor bears a typical median dorsal spine.
The position of this genus, partly on account of the poor condition of the
unique type, is obscure. The venation, however, shows it to be an extraordinarily
aberrant form, which, with some hesitation, I place nearer to Sawda Dist. than
to any other known genus. The tymbal covers closely resemble those of Sawda,
though the opercula are much shorter than is usual in that genus. The narrow
58 INSECTS OF SAMOA.
head is another common feature, as-are also the close proximity of the ocelli
and the sudden dip of the crown, so that the median ocellus looks directly
forward. The meracanthus resembles closely that of Sawda froggatti Dist. It
may be noticed that Sawda is an Austro-Malayan genus, with one species (known
only from the female) also in Fiji. The present form could perhaps have been
derived from a Sawda-like immigrant in Samoa.
2. Moana expansa, sp. n. Figs. 1-3, 22.
Male: dull olivaceous brown (type probably faded). Frons, sides of
abdomen, and legs somewhat paler. A little scattered pale decumbent
pubescence on thorax and abdomen, but no hairs. Ocelli edged with blackish.
An indistinct paler median longitudinal stripe on pronotum. Mesonotum
shining, with two narrow black longitudinal stripes on each side of disc, the
Text-ric. 2.—Moana expansa, last nymphal Trext-Fic. 3.—Moana expansa, abdominal
instar, fore femur. extremity of last nymphal instar
of male.
inner pair impressed and about two-thirds length of outer pair, which nearly
reach cruciform elevation. Tergites II-VII smudged with blackish centrally,
and II—-VIII laterally also. Tegmina hyaline, the veins brownish, darker
distally.
Crown (including that part of frons visible dorsally) a little longer than
basal width between eyes, and equal in length to dise of pronotum without
CICADIDAE. 59
posterior depressed margin. Eyes very large. Posterior margin about one-
fourth total length of pronotum. Opercula rounded apically, barely longer
than meracanthus, widely visible in dorsal view. Fore femora weakly armed.
Hind tibiae with five non-apical spines. (Hypandrium missing in type.)
Lateral projections of pygophor short, blunt, emarginate dorso-apically.
(Rostrum and most of legs missing in case of type.)
Length to tip of VIIIth tergite 25-3 mm., tegmen length, 43-3 mm., greatest
width of tegmen (about halfway), 15-6 mm.
Male nymph, last instar: Eyes very large, antennae with 8 segments, of
which I-IV have the usual few very large setae, while the rest are densely and
evenly clothed with numerous small bristles. The tegmen pads show distinctly
the strong and much curved Cu, and the weak M stem alongside it. Fossorial
fore-legs with femora as figured. Pygophor ends in a typical median tergal
spine, which is, however, delimited by a furrow shown dotted in the figure.
Length 24-0 mm. (Length of head and thorax added to that of the abdomen,
to avoid curve.)
Upolu: Lanutoo, 1 3, more or less fragmentary, taken from a spider’s
web ; skilfully mended by Mr. China; no date (Hopkins) ; Malololelei, exuviae
of one last instar g nymph collected in rain forest at 2,000 feet, 26.iv.1924
(Bryan).
I have felt very chary of erecting a new genus on such poor material, but
when one considers how many new Orders and lesser groups are founded on far
less complete fossil insects, the objection ceases to have weight. The genus is
well characterised, and the species should be recognisable from the above
description.
The nymph, which is in perfect condition, is as aberrant as the imago.
In most cicada nymphs known to me (e.g. Platypleura, Henicopsaltria, Dicero-
procta, Odopoea, Magicicada, Melampsalta, Baeturia, Tettigarcta), the antennal
segments, even to the apical one, are supplied with a relatively small number
(2 or 3 up to about 6 per segment) of very large setae, sometimes as long as the
segment itself, whereas in the present species these are replaced on segments
V-VIII by numerous small bristles.
60 INSECTS OF SAMOA.
Subfamily TIBICININAE Dist.
Genus Barturta Distant, 1892.
Type Cicada conviva Stal.
3. Baeturia exhausta Guér. Figs. 5-9, 11-17, 19-21.
Distant, Syn. Cat. Hom. Cicad., p. 157, 1906.
Cicada exhausta Guér., Voy. “ Coquille,” Zool., 11, p. 181 (1838), pl. x, fig. 6 (1831).
C. hastipennis Walk., List Hom. Brit. Mus., Suppl., p. 30, 1858.
Dundubia parabola Walk., Ins. Saund. Hom., p. 6, 1858.
Savai: Safune, up to 4,000 feet in rain forest, 4 gg, 1 9, 2 exuviae, v. and
xi. (Buxton, Hopkins, Bryan).
Upolu: Apia, Malololelei, Aleipata, Vaea, up to 2,000 feet, 14 gg, 25 99,
3 exuviae, every month except 1. and x. (various collectors).
Tutuila: Pago Pago, Fagatoga, Amauli, 8 §g, 11 99, i.., UL, ix. (Steffany,
Judd, Swezey, Wilder).
Manua: Ofu, Tau, 3 3g, u. (Judd).
Samoa (island not specified): 2 9g, 5 99, ui., iv. (Tate, O'Connor, and
collector unknown—Brit. Mus.).
Total Samoan material: 31 34, 42 99, 5 exuviae (last instar).
There are two abundant and widespread Austro-Malayan species of Baeturva,
which differ from the other species known to me in the blackish speckling of
their pale ground colour. These two, B. exhausta Guér. and B. conviva Stal,
have sometimes been considered only doubtfully distinct (Kirkaldy, Haw. Sugar
Planters’ Exp. Sta. Entom. Bull., 12, p. 8, 1913). The characters on which
they are usually separated are the greater average size of B. exhausta, and the
extension of its tegminal cell R; (3rd apical cell) basad of cell Ry (1st apical
cell), whereas in B. conviva the bases of these two cells are opposite. On these
two characters the Samoan material all agrees with B. exhausta.
Save for the Samoan extension of B. exhausta, the range of these two forms
seems practically coincident. Judging from the above macroscopic criteria there
are in the British Museum examples of both from Buru, Ceram, Amboina and
New Guinea. There are specimens of B. conviva only from Doré, Timor, Mysol,
Batchian and Duke of York I. (Bismarck Arch. ?) ; and of B. exhausta only from
Gilolo, Torres Strait and Samoa. Size is at best an untrustworthy character,
and in examples from several Austro-Malayan islands, notably Buru, the vena-
tional distinction between the two forms also breaks down. In the British
Museum there are two males from the same locality in Buru, taken at the same
CICADIDAE. 61
time, and almost identical in other respects, but with the base of cell R; basad
of cell Ro in the one, and distad in the other. The apical venation in Cicadidae
is not very generally constant in details, although in the 73 Samoan examples
this particular character holds. Unfortunately the type of neither species is
available for study. The male external genitalia show certain variations, both
in the aedeagus and in the copulatory claspers of the Xth segment, but at least
in the two males from Buru already mentioned (figs. 6, 7, 12, 13) these are not
ee
=
re
.
oe sie
Text-Fies. 4-9.—Fig. 4, Baeturia conviva Stal, Ceram, aedeagus, lateral view; fig. 5, B. ex-
hausta Guér., type of Dundubia parabola Walk., aedeagus, lateral view ; fig. 6, B. exhausta
Guér., (example from W. Buru with venation of B. conviva), aedeagus, lateral view ; fig. 7,
B. exhausta Guér., (example from W. Buru with normal venation), aedeagus, lateral view ;
fig. 8, B. exhausta Guér., Samoan example, aedeagus, lateral view; fig. 9, B. exhausta Guér.,
(another specimen from Samoa), aedeagus, lateral view.
correlated with the venational differences. On characters presented by the
genitalia these two males belong to B. exhausta. Figs. 4 and 10 show the corre-
sponding parts in what we must provisionally regard as a typical B. conviva,
to be contrasted with figs. 5 and 11, drawn from Walker’s type of Dundubia
parabola, which, with Distant, I consider as synonymous with B. exhausta.
The other figs. (8, 9, 14-17) illustrate the variation among the Samoan material.
If the specimen of B. conviva dissected by me is really typical, then this species
may perhaps be regarded as distinct owing to the longer aedeagus and accom-
62 INSECTS OF SAMOA.
panying minor differences in this organ and in the shape of the claspers. On the
2
ZB
=
=!
Text-Fies. 10-17.—Fig. 10, B. conviva Stal,
Ceram, left clasper (Xth segment) ; fig. 11,
B. exhausta Guér., type of D. parabola
Walk., left clasper (Xth segment) ; fig. 12,
B. exhausta Guér., (example from W. Buru
with venation of B. conviva), left clasper
(Xth segment) ; fig. 13, B. exhausta Gueér.,
(example from W. Buru with normal
venation), left clasper (Xth segment) ;
fig. 14, B. exhausta Guér., from Samoa, left
clasper (Xth segment) ; fig. 15, B. exhausta
Guér., (another specimen from Samoa,
Apia), left clasper (Xth segment) ; fig.
16, B. exhausta Guér., (still another speci-
men from Samoa, Apia), left clasper
(Xth segment) ; fig. 17, B. exhausta Guér.,
Samoan example (Savaii), left clasper
(Xth segment).
basis of macroscopic and microscopic
characters it is evident that we are deal-
ing with one polymorphic form, com-
parable with Metrosideros villosa auctt.
among the Pacific Myrtacae. To settle
the question it will be necessary firstly,
of course, to examine the respective
types, and secondly to study series
from the different islands comparable
with the extensive one from Samoa
that I have been privileged to ex-
It seems to me very likely
that material from more numerous
intermediate stations would yield still
more annectent examples than we have
at present. In the meantime B. conviva
is kept doubtfully distinct, and the
Samoan forms are regarded as belonging
to B. exhausta.
mental variation in genitalia in the
genus, the aedeagus of an undoubtedly
distinct species, B. famulus Dist., is
amine.
To show more funda-
represented in fig. 18.
The type of Cicada hastipennis
Walk. is a female which agrees specifi-
cally with D. parabola, and thus pre-
sumably with B. exhausta.*
The nymph of Baeturia exhausta.
—As pointed out on other occasions,
the last nymphal exuviae of cicadas
make beautiful objects of study. In
the present species the antennae have
9 segments, including a very minute
* In view of the above discussion, the Baeturia species of Buru are of the greatest interest.
Schmidt (Tvreubia, 7, p. 222, 1926) records from the island in question B. conviva, B. exhausta,
CICADIDAE. 63
apical one. These segments are supplied, as usual, with a few setae, up to half
a dozen on a segment, and sometimes as long as the segment itself. Between
these large setae the surface is bare of ordinary hairs. The fore femur and the
extremity of the abdomen in the last instar male nymph are shown in figs. 19, 20.
Length (length of head and thorax added to that of abdomen) 18-8 mm.
.
Trxt-Fic. 18.—B. famulus Text-Fic. 19-22.—Fig. 19, B. exhausta Gueér., last
Dist., aedeagus, lateral nymphal instar, fore femur; fig. 20, B. exhausta
“view. Guér., abdominal extremity of last nymphal instar
of male; fig. 21, B. exhausta Guér., apical antennal
segments of last nymphal instar of male; fig. 22,
Moana expansa, apical antennal segments of last
nymphal instar of male.
One male specimen of B. exhausta from Apia (Buxton and Hopkins) has a
mass of hardened adhesive matter clogging its face and mouth-parts. It bears
the highly interesting explanatory label, “‘ The value of latex to plants! This
cicada tried to suck Carica papaya.”
B. bicolorata Dist. and B. schulzi, sp. nov., but, from his notes based on badly-preserved material
I doubt whether the third species is correctly recorded. His B. schulzi is said to be related to
B. viridicata Dist., of New Guinea, but judging from the description it is evidently much more
typical. The type of B. viridicata has almost opaque tegmina with the ambient vein almost
coincident with the margin, and greatly reduced opercula, which reach only half-way along the
meracanthus. In these three respects B. schulzi would seem more like B. exhausta and B. conviva.
Schmidt's new genus, Toxopeusella (I.c., p. 224) is apparently well-founded, although the geno-type,
Cicada stigma Walk., is represented in the British Museum only by the original female. The
common stem from the basal cell of the tegmen is not, however, M plus Cu as stated by Schmidt,
but M plus Cu, only. The statement that fore and hind wings are in 7’. stigma “ quergerippt ”’
and in Baeturia not, is incorrect, as an examination of B. exhausta and B. conviva will show, though
this character is certainly more marked in 7’. stagma.
TW. 2 2
64 INSECTS OF SAMOA.
A female example (Apia, 11.1924, Buxton and Hopkins) has a large piece
bitten out of the base of the abdomen, and was taken from a large introduced
wasp, Polistes macaensis which had captured it. Vespids are also known as
enemies of cicadas in North America (Davis).
“ Dasypsaltria maera”’ Haupt = Platypleura dwisa (Germ._).
In 1917 Haupt (Stett. Ent. Zeit., Bd. 18, p. 303, fig. 1) erected under the
name Dasypsaltria, a new monotypic genus to receive D. maera, a supposed
new species believed to have been collected in Samoa. On 7th June, 1926,
Herr Haupt was so good as to write to me regarding this species as follows :—
“Tn meiner Dasypsalirva maera mochte ich bemerken, dass ich da getiiuscht
worden bin. Sie gehért zu der stark varnerenden Platypl. divisa Germ., was ich
aber wegen des etwas abnormen Pronotums leider zu spit erkannte. Getiaiuscht
wurde ich auch durch die Herkunfts-Angabe. Einer unserer Kolonial Beamten
hatte das Tier in Stid-Afrika erbeutet, reiste von dort nach Samoa und sandte
es von dort zur Bestimmung nach Halle.” Dasypsaltria is thus a synonym of
Platypleura.
In the meantime, unfortunately, Kato (Trans. Nat. Hist. Soc. Formosa, 17,
p- 210, 1927) has described a new Formosan cicada as Dasypsaltria formosana.
CICADIDAE. 65
LIST OF TEXT-FIGURES.
(Corresponding parts shown at similar magnification ; hairs and setae omitted.)
I.
2.
3.
4.
5
for)
“J
21.
22.
Moana expansa g. et sp. n., tegmen, A. J. E. Terzi del.
Moana expansa, last nymphal instar, fore femur.
Moana expansa, abdominal extremity of last nymphal instar of male.
Baeturia conviva Stal, Ceram, aedeagus, lateral view.
. B. exhausta Guér., type of Dundubia parabola, Walk., aedeagus, lateral view.
. echausta Guér., Bara, W. Buru (example with venation of B. conviva), aedeagus,
lateral view. -
. exhausta Guér., Bara, W. Buru (example with normal venation), aedeagus, lateral
view.
. echausta Guér., Apia, Samoa, aedeagus, lateral view.
. echausta, Guér., Apia, Samoa (another specimen), aedeagus, lateral view.
. conviva Stal, Ceram, left clasper (Xth segment).
. echausta Guér., Bara, W. Buru (example with venation of B. conviva), left clasper
(Xth segment).
. exhausta Guér., Bara, W. Buru (example with normal venation), left clasper
(Xth segment).
. echausta Guér., Apia, Samoa, left clasper (Xth segment).
. echausta Guér., Apia, Samoa (another specimen), left clasper (Xth segment).
B
B
B
B
B
. B. exhausta Guér., type of D. parabola, Walk., left clasper (Xth segment).
B
B
B
B
. B. exhausta Guér., Apia, Samoa (still another specimen), left clasper (Xth segment).
. B. exhausta Guér., Savaii, Samoa, left clasper (Xth segment).
. B. famulus, Dist., aedeagus, lateral view.
. B. exhausta Guér., last nymphal instar, fore femur.
. B. exhausta Guér., abdominal extremity of last nymphal instar of male.
B. exhausta Guér., apical antennal segments of last nymphal instar of male.
Moana expansa, apical antennal segments of last nymphal instar of male.
- AQUATIC AND SEMIAQUATIC HETEROPTERA.
By Tetso Hsaxt.
(With 6 Text-figures.)
ALTHOUGH a few representatives of this group of insects have been described
from material collected in Fiji, the Marquesas Is., New Caledonia and the Society
Is., none have yet been recorded from Samoa.
In the present paper ten species, representing four families, are mentioned
or described. All belong to genera typical of the South Pacific region, but they
include five new species, some of which may prove to be endemic.
Unless otherwise stated, all the specimens were collected by Messrs. P. A.
Buxton and G. H. Hopkins.
VELIIDAE.
Microvelia Stal.
Besides those occurring in Australia, three species of this genus, Mvcrovelia
oceanca Distant (New Caledonia), M. pacifica Kirkaldy (Fiji), and M. prompta
Cheesman (Tahiti), are known to exist in the South Pacific Islands. Another
new species is now added to the fauna.
1. Microvelia samoana, sp. n. ‘Text-fig. 1.
6 2. Black, covered with grayish pubescence. Head black, lateral portions
of vertex along the eyes brown, pubescence whitish along the median longi-
tudinal line. Eyes black, shining. Antennae dark brown, apical segment
darker. Rostrum dark brown, with the extreme apex black. Pronotum black,
with a conspicuous brown band along the anterior margin where the pubescence
is pale brown. Dorsal and ventral surfaces of meso- and metathorax, and of
abdomen totally black, connexivum sometimes slightly brownish. Legs brown,
with the apical one-third to two-fifths of femora, tibiae and tarsi much darker
than the rest of femora, trochanters and coxae.
67
68 INSECTS OF SAMOA.
Size, small. Body elongate, lateral margins roughly parallel, covered with
coarse pubescence and punctures. Vertex conically produced anteriorly,
rounded at the apex. Three or four pairs of sensory pits along the lateral
submarginal areas; the apex with long setae. Eyes much rounded laterally,
agglomerate. Antennae slender, first
segment stout, slightly curved out-
wardly, slightly thickened at the
middle ; second, three-fourths the
length of first, thinner than the latter,
a little thickened towards the apex ;
third very slender, nearly as long as
first ; fourth much the longest, twice
as long as second, one and a _ half
times as long as first, thickened im
the middle. Ratio of the antennal
seoments—12: 9:11:18. Rostrum
reaching the anterior coxae, second
segment (in reality this may be the
third) much the longest. Pronotum
transverse, both anterior and posterior
margins shallowly sinuate posteriorly,
lateral margins slightly rounded, con-
vergent anteriorly ; a more or less dis-
tinct submarginal depression along the
posterior margin; meso- and meta-
notum together about half as long as
pronotum. Legs moderately slender,
covered with setose hairs, femora moderately thickened in the middle; anterior
tibia about three-fourths of femur, tarsus long, a little longer than half the
length of tibia; intermediate femur and tibia nearly equal in length, tarsus
about half as long as tibia, two segments of equal length ; posterior tibia longer
than femur, slender, tarsus short, two segments of equal length. First four
dorsal abdominal segments subequal in length, fifth a little longer than fourth,
sixth nearly as long as fourth and fifth taken together. Connexivum obliquely
erect in male, turning over the dorsal surface of abdomen in female, in which the
dorsal surface of the last four segments is scarcely visible.
Y|
Text-Fie. 1.—Microvelia samoana Esaki,
sp. By, 6:
AQUATIC HETEROPTERA. 69
Length of body: ¢ 1-6 mm., 9 1:7 mm.; breadth of body: ¢ 0:6 mm.,
2 0-7 mm.
Upolu: Mulifanua, type 3, type 9, 2 ¢ paratypes, x.1925, in marsh.
This species is smaller than Microvelia prompta Cheesman and larger than
M. oceanica Distant.* From the former M. samoana Esaki differs also in the
stouter shape of the body and the much narrower connexivum. From the
latter it differs in the ratio of the antennal segments.
GERRIDAE.
Limnogonus Stal.
Three closely allied species of this genus occur in the Pacific Islands. These
are :
Lamnogonus fossarum (Fabricius), type locality, E. India.
Lamnogonus discolor (Stal), type locality, Tahiti.
Lamnogonus luctuosus (Montrouzier), type locality, New Caledonia.
Lamnogonus nymphae (Ksaki), of Formosa, can be synonymised with L.
fossarum (Fabricius), and L. lineatus (Carpenter), of Murray Island, may be
synonymous with L. discolor (Stal). The type (¢) of L. lineatus (Carpenter)
is in the British Museum ; with that exception I have not yet had the oppor-
* Hale (Records S. Austr. Mus., Adelaide, iii, p. 208, 1926) identified an Australian Microvelva
as M. oceanica, but his species is quite distinct from WM. oceanica Distant, and is apparently new.
It may be characterised as follows :
Microvelia halei, sp. n.
$2. Differing from Microvelia oceanica Distant in the following characters :
Larger size: Length of body in apterous form ‘‘ 1-66 mm. to 2 mm.,” while in the apterous
form of M. oceanica, the body measures 1-51 mm. in the 3, and 1-65 mm. in the 9. These measure-
ments were made from the cotypes of WM. oceanica Distant in the British Museum. Distant’s
original description (Sarasin and Roux, Nova Caledonia, Zool., i, pt. 4, no. 10, p. 385, 1914) was
based only on the winged form, which measures “2 mm.,” though the author gave figures of
both forms.
Different ratio of the antennal segments: Hale describes the antennae of M. oceanica Hale,
nec Distant, as follows: “ first segment nearly one-fourth longer than second, subequal in length
to third and about three-fifths as long as fourth, which is more than twice as long as the second.”
In the true M. oceanica the ratio is 10:8:10:21 and 11:9:11:23 in g and Q respectively.
Thus in the latter species the last segment is considerably longer, in comparison with the other
segments, than in M. halei, in which it is much less than twice the length of the third segment.
Reference should be made to the full description and figures given by Hale (loc. cit.).
70 INSECTS OF SAMOA.
tunity of studying the types of these species. It is therefore impossible to
identify the species correctly, since the original descriptions are all insufficient
to distinguish one from another. For the time being, therefore, I have identified
them with three distinct species occurring respectively in the type localities.
A common Indian species, which is also widely distributed over the Malay
Archipelago, the Philippine Is. and Formosa, has been selected as probably
identical with ZL. fossarum (Fabricius); the only Tahitian species of which
there are examples in the British Museum has been regarded as representing
L. discolor (Stal); and a New Caledonian species has been identified as L.
SAO
Text-Fic. 2.—Ventral view of the genital segments of Limnogonus-species, semi-diagrammatic.
a, Limnogonus luctuosus (Montrouzier), ¢; 6, 9. 6 Limnogonus pacificus Esaki, sp. n., 3;
d, 2. e, Limnogonus fossarum (Fabricius), 3; jf; &.
luctuosus (Montrouzier). When the types of the species in question are
examined it is possible that some alterations will have to be made. Of these
three species, one occurs in Samoa.
2. Limnogonus luctuosus (Montrouzier). Text-fig. 2, a, b.
Gerris luctuosa Montrouzier, Ann. Soc. Linn. Lyon, xi, p. 242, 1864; Distant, Sarasin and Roux,
Nova Caledonia, Zool., i, pt. 4, no. 10, p. 384, 1914.
Upolu: Laulu, 1 9, 21.1.1925.
This species was hitherto only known to occur in New Caledonia.
AQUATIC HETEROPTERA. 71
3. Limnogonus pacificus, sp. n. Text-fig. 2, c, d.
3 2. Very closely allied to Limnogonus fossarum (Fabricius) and its allies,
differing only in the structure of the genital segments of both sexes, except that
the female is proportionally much stouter in shape than the male.
The differences in the genital segments of the two species (better shown in
text-fig. 2, ¢, d, e, f) are indicated in the subjoined tabular statement.
L. pacificus, sp. n. L. fossarum Fabricius.
S$. Posterior ventral margin of sixth G. Posterior ventral margin of sixth
abdominal segment sinuate on each side of abdominal segment deeply excavated in middle
middle line (see text-fig. 2, c). _ line, thus forming a parabola (text-fig. 2, ¢).
[The figure is made from a specimen from
Bor Ghat, India. The shape of this sinuation
seems variable to some extent, but it is never
as in L. pacificus.|
Posterior ventral margin of first genital Posterior ventral margin of first genital
segment distinctly pointed in the middle. segment slightly sinuate, not at all pointed in
the middle. Second ventral genital segment
broader, lateral margins convergent posteriorly.
9. Abdomen and connexiva much broader ; 9. Last abdominal segment: less reflexed
sternite of last abdominal segment much more dorsally, forming three equal teeth seen from
reflexed dorsally. venter.
All the specimens of L. pacificus in the collection are apterous. Pronotum
protruded posteriorly, covering a part of mesonotum, as in the apterous form
of the allied species. Second and third abdominal tergites in female with
bluish-gray pubescence. Antennae similar to those of L. fossarum, the first
segment much the longest, the remaining three segments subequal in length :
the exact ratio shows a slight variation to some extent.
Length of body: 3 8-0-8-5 mm., 9 8-5-9 mm. ; breadth of body: ¢2mm.,
° 3 mm.
Upolu: Malololelei, 2,000 feet, type gy 1., v.1924; Apia, 2 99 (including
type 9), 9.1v.1924, 1 3 paratype, 25.1v.1924; Mulifanua, 1 9, 17.v1.1925.
4. Limnogonus buxtoni, sp. n. Text-fig. 3.
The apterous form is the only one represented in the collection. Males
much smaller than females.
3g. Fusiform. General markings similar to those of the other species of
the genus, but the insect is more or less thickly covered with minute golden
72 INSECTS OF SAMOA.
pubescence, partially shining on pronotum and dorsal surface of abdomen.
Head black above, with a pale yellowish brown U-shaped marking and a small
spot just in front of each eye, which is confluent with the pale yellowish brown
under side of head. Eyes dark brown, shining. Antennae dark brown.
Rostrum pale yellowish brown, except extreme apex, which is black. Pronotum
black, lateral and postero-lateral margins, two spots on the anterior area, and
a stripe along the median longitudinal line except the anterior part pale yellowish
Cc d
Text-ric. 3.—Limnogonus buatoni Esaki, sp. n. a, 3, dorsal view; 6, antenna; c, dg, genital
segments, ventral view, shown upside down; d, the same of 9.
brown. Dorsal surface of meso- and metanotum, and of abdomen black, with
a more or less distinct stripe along the median line, pale yellowish brown ; in
© lateral portions of last three or four dorsal segments faintly brown. Legs
dark brown, anterior femur paler. Sides and under side of body pale yellowish
brown, with the lateral area of prothorax, a small spot on pro-acetabulum,
lateral stripes on mesothorax, a very small spot at the end of meso-acetabular
suture, and a large spot on mego- and meta-acetabulum, black. Sides of abdomen
dark brown, or a dark brown spot on each segment.
yes much rounded exteriorly, distinctly but not much emarginate interiorly.
Three pairs of very long sensory setae along the submarginal line on vertex
(easily visible in profile). Antennae very long and slender, first segment the
AQUATIC HETEROPTERA. 73
stoutest and slightly curved near base; second and third segments equal in
length ; fourth a little longer than first or subequal to it. Hind margin of last
(sixth) abdominal sternite roundly sinuate, forming an arc, genital segments
much rounded at the apex (see text-fig. 3).
2. Similar to male, but much larger. Fourth segment of antennae some-
times slightly shorter than, generally subequal to first segment in length. Last
(sixth) abdominal sternite truncate at apex, without tooth-like process.
a
C
Text-Fie. 4.—Limnogonus hopkinsi Esaki, sp. n. @, antenna: 6b, ¢ genital segments, ventral
view ; ¢, the same of 9.
Length of body: ¢4mm., 26mm. ; breadth of body: $13 mm., 2? 2mm.
Upolu: Apia, type g, type 2, 3 5 paratypes and 1 @ paratype, 5.v1.1924 ;
Vailima, 1 gf, 1 9, 20.11.1925.
This interesting species is unlike any other known member of the genus
Limnogonus owing to the striking difference in size between the sexes, the shape
of the last abdominal and genital segments, and the considerable length of the
last antennal segment, which is the longest of all in the male. With the excep-
tion of some oceanic forms, the sexual difference in size is not so striking in other
members of the Gerris-group. Loamnogonus hopkinsi, sp. n., described below—a
much larger species—exhibits a similar sexual dimorphism in the matter of size.
5. Limnogonus hopkinsi, sp. n. Text-fig. 4.
6. Apparently very similar to the foregoing species, except in size. The
description of the colour and markings of L. buxtoni Esaki will serve also for
the present species. The 3 of this species is about as large as the 9 of L. buxtons,
74 INSECTS OF SAMOA.
while the 2 of the former is considerably larger than the male. Among the
material collected, there is a slight variation in size and coloration. The
specimens taken in Savai are much darker than those obtained in Upolu. In
the latter material the brown margins of the pronotum are sometimes much
more widened at the apex.
First segment of antennae much the longest, second and fourth subequal,
or the former a little shorter than the latter ; third, the shortest ; fourth, unlike
that in L. buxtoni, always much shorter than the first.
Genital segments of both sexes very similar to those of LZ. buxtont as shown
in the figures.
Length of body: ¢ 5-6 mm., 2 9-10 mm.; breadth of body: ¢2mm.,
O38 mm;
Upolu: Malololelei, 2,000 feet, type g, type 2, 3.g and 2 2 paratypes,
4.v.1924; 1 g, l.v.1924; 6 dg, 6 OY, 14-30.v1.1924. Savaii: Safune, rain
forest, 2,000-4,000 feet, 4 fg, 3 29, 5.viii.1924 (Bryan).
It appears probable that this species is confined to the mountain streams ;
L. buxtom occurs at lower levels.
6. Halobates princeps B. White.
Halobates princeps B. White, Rept. Voy. “‘ Challenger,” Zool., vii, pt. 19, pp. 39, 44, pl. 1, fig. 3, 1883.
Halobates alluaudi Bergroth, Rev. dEnt., xii, p. 204, 1893; Kirkaldy, Ann. Soc. Ent. France,
Ixvii, p. 103, 1899 ; Esaki, Ann. Mus. Nat. Hung., xxiii, p. 133, 1926 (Syn. nov.).
Halobates sericeus Matsumura (nec Eschscholtz), Thous. Ins. Japan, Addit., i, p. 97, pl. xi, fig. 8,
91: .
ae matsumurat Esaki, Psyche, xxxi, p. 117, pl. v, fig. D, 1924 (Syn. nov.).
Upolu: Apia, 1 3, 1 9, 2.11.1924, on surface of sea.
Bergroth, who described Halobates alluaudc from material from the
Seychelles, wrote (loc. cit., p. 205): “ H. princip: B. White affinis, sed tarsis
omnibus, praesertim posticis, multo brevioribus et trochanteribus mediis iner-
mibus bene distinctus.” Recently I have examined the type of H. princeps
B. White (a single female, in the British Museum), and have found an important
particular in which B. White’s statement and figure are incorrect. He writes
(loc. cit., p. 45): “ [the hind] tarsus about five-eighths of the length of tibia,”
but, in reality, the measurements afforded by the unique type are :—hind tibia :
tarsus = 43: 138, 2.e. roughly 3:1. This agrees quite well with the statement
of Bergroth (loc. cit., p. 205), in the description of H. alluaudi : “ [tibia postica]
tarso saltem triplo longiore,” which is the only difference between H. princeps
AQUATIC HETEROPTERA. 15
and H. alluaudi, except the armature of the middle trochanters. B. White
(loc. cit., p. 45), writes: “[the middle] trochanter armed with spines,” but
these “ spines” are not real spines, but stout setae, which are sometimes torn
off even during the life of the insect. I have also examined the type material
of H. alluaudi Bergroth in the Muséum National d’ Histoire Naturelle, Paris, and
am now quite sure that this supposed species is identical with H. princeps B.
White. Halobates matsumurar Ksaki is also synonymous with this species.
H. princeps is now known to be widely distributed over the Indian and
Pacific Oceans, as far as the Seychelles in the west, and the Pacific coast. of
Southern Japan in the north.
b)
GELASTOCORIDAE.
7. Peltopterus macrothorax (Montrouzier).
Galqulus macrothorax Montrouzier, Ann. Sc. Phys. Nat., Lyon, (2), vii, pt. 1, p. 110, 1855.
Scylaecus macrothorax Stal, Of. Vet.-Akad. Forh., xviii, p. 201, 1861; Enum. Hemip, v (Svensk.
Vet.-Akad. Handl., xi, no. 4), p. 139, 1876. é
Peltopterus macrothorax Stal, Berlin, Hint. Zeit., vii, p. 408, 1863; Ofv. Vet.-Akad. Férh., xxvii,
p-. 706, 1870; Montandon, Bul. Soc. Sev. Bucuresci, viii, p. 779, 1899.
Tutuila: Leone Road, a single J, 8-5 mm. in length, 9.1x.1926 (Judd).
This species was previously known to occur in Woodlark I., Fiji, the
Philippine Is., N. Borneo, Aru, the Solomon Is., and Marianne or Ladrone Is.
In the British Museum collection there are, in addition to those from Woodlark I.,
Fiji, the Marianne and Philippine Is., specimens from Ke Dulan (in the Ké [= Kei]
Islands, 1 g, 1 9, 25.1x.1874, “‘ Challenger ” collection), Formosa (1 3, without
further data), and ‘‘ Madgico-sima, Corea ” * (3 29).
* These specimens from “ Corea” were collected by Arthur Adams during the voyage of
H.M.S. Samarang under Captain Sir EH. Belcher, C.B., R.N., and deposited in the British Museum
in 1845 with many other specimens. The three specimens are labelled “ Corea,” and only one
of them has a second label, ‘‘ Dry sand, Madgico-sima.”’ There is no doubt that these three were
collected at the same time. I believe that “‘ Madgico-sima ”’ is not an island near Corea, because
at the time of this voyage no Japanese name was officially used for the Corean geographical places
(“‘ sima’’ or shima means “island”’ in Japanese). On the other hand, the Yayeyama Islands in
the Loo Choo Group, lying between Okinawa and Formosa, have sometimes been called as “ Majico-
sima”’ or “‘ Mayico-sima.”’ It may be fairly safe to assume that this ‘‘ Madgico-sima ”’ is identical
with “ Mayico-sima.” If my assumption is correct, it is very interesting to find this typical
oceanic species on these islands (the chief islands are Ishigaki-jima and Iriomote-jima), which
are, together with Kotdsho (= Botol Tobago), zoogeographically much more closely related to
the Philippine Islands than to Formosa, in spite of their closer proximity to the latter than to the
76 INSECTS OF SAMOA.
NOTONECTIDAE.
Anisops Spinola.
This large genus, which is represented by three species in the collection,
has now become taxonomically one of the most difficult groups of water-bugs.
As Bergroth * remarked, “ Kirkaldy has published revisions of the different
genera + of this family, but his attempt to make them better known was a
failure. These insects offer good specific characters, but Kirkaldy was unable
to express the differences in words, and his descriptions are so vague and con-
fusing, and in so many cases positively wrong (especially in the comparative
measurements of the tibiae and tarsal joints) that they are practically almost
useless.”
The species mentioned below have been carefully determined by the study
of many types and of more recent literature of the group.
8. Anisops fieber1 Kirkaldy.
Anisops niveus Fieber (nec Fabricius), Abh. bihm. Gesel. Wiss., Prague, (5), vii, p. 484, 1852 ;
Matsumura, Trans. Sapporo Nat. Hist. Soc., i, p. 28, 1906 ; Esaki, Ent. Mag., Kyoto, i, p. 31,
1915.
Anisops fiebert Kirkaldy,{ Entomologist, zexiv, ip: 5; 190); Ween: Rant: Zet., xxi, ps 16a 1904
Distant, Faun. Brit. Ind., Rhynch., iii, p. 46, 1906 ; Hale, Record. S. Austral. Mus., 1, p. 400,
fig. 363, 1923; Arkw Zool., Stockholm, xvii, A, no. 20, p. 17, 1925; Esaki, Ann. Mus.
Nation. Hungar., xxiv, p. 188, 1926.
Anisops kuroiwae Matsumura, Ent. Mag., Kyoto, 1, p. 109, pl. in, fig. 38, 1915 (Syn. nov.).
former. It may also be pointed out here that a Malayan cicadid, Cryptotympana aquila (Walker)
was originally described by Walker (List Homop., i, p. 84, 1850) from a specimen from “ Corea.”
The type is still preserved in the British Museum, and the label refers the specimen to the same
source as the specimens of Peltopterus macrothorax from “* Corea’ mentioned above. It is, there-
fore, very reasonable to suppose that this specimen was also collected on “ Madgico-sima”’ or
Yayeyama Islands instead of Corea, though there are no further data on the label, since, judging
by the distributive range of this species (known to include the Malay Peninsula, Borneo, Sumatra
and Java), it is almost impossible to believe that the specimen was actually collected in Corea, as
has already been doubted by several authors (e.g. Moulton, Journ. Feder. Malay Stat. Mus., xi,
pp- 138 and 169, 1923). This species also has recently been recorded from Formosa, though I
have not seen the actual specimen (cf. Kato, Trans. Nat. Hist. Soc. Formosa, xvii, p. 23, 1927).
* Meddel. Goteborgs Mus. Zool. Afdel. 4 (Géoteborgs Kungl. Vetenskaps- och Vitterhetssamhdlles
Handlingar, xvi, no. 2), p. 15, 1914.
+ For Anisops, see Wien. Ent. Zeit., xxiii, pp. 111-119, 1904.—T.K.
+ Kirkaldy bestowed this new name upon 4. niveus Fieber, on the assumption that this
species was not identical with A. niveus (Fabricius) (Syst. Lnt., p.690, 1775). In the opinion of the
author in question, Notonecla ciliata Fabricius (Ent. Syst., Suppl., p. 52, 1798) and Anisops hyalinus
AQUATIC HETEROPTERA. 77
Upolu: Mulifanua, 2 3g, 3 99, 9.x1.1925.
Previously known to occur in India, Ceylon, Celebes, New Guinea, N.
Australia, Formosa, and Loo Choo.
: Anisops kurowwae Matsumura is identical with A. fiebert. Matsumura says
that A. kurowae is distinctly larger (“ deutlich grosser ’’) than A. fieber?, but the
former is “6-6-5 mm.” in length, whereas A. fiebert measures 4:9-7-2 mm.,
generally 6-5-7 mm. (Fieber’s measurements, “34” lines for $ and 9,
practically = 7 mm.). As a matter of fact I have seen many specimens from
Formosa and the Loo Choo Is., in both of which localities individuals forming
part of Matsumura’s typical series of A. kurovwae were obtained, but no differences
were observed between them and examples collected elsewhere.
9. Anisops cleopatra Distant. Text-fig. 5.
Anisops cleopatra Distant, Sarasin and Roux, Nova Caledonia, Zool., i, pt. 4, no. 10, p. 386, pl. xi,
fig. 8, 1914.
Upolu: Laulii, 5 specimens, 21.1.1925; Apia, 16 specimens, 1.1924, 3
specimens, 19.vi.1924; Malololelei, 4 specimens, 9.vi.1924; Mulifanua, 2
specimens, 9.x1.1925. Also 1 specimen, “ Samoa,’ 1920 (O’Connor).
‘This species was hitherto known only from specimens from New Caledonia.
The description by Distant, though accompanied by a beautiful drawing, is
very insufficient. The type is not to be found in the British Museum, which
however possesses the types of some other species described in the same paper.
But there are two specimens * from New Caledonia (J. J. Walker) in the
Fieber (Abh. béhm. Gesel. Wiss., Prague, (5), vil, p. 482, 1852) are identical with Anisops niveus
(Fabricius) (cf. Kirkaldy, Ann. Soc. Hnt. France, Ixviii, p. 106, 1899). All these theories, however,
were based on the “type” of Notonecta nivea Fabricius, which was said to be preserved in the British
Museum (Kirkaldy, Wren., Hnt. Zeit., xxii, p. 119, 1904). But although a Fabrician specimen,
which is identical with A. niveus auct., and was determined by Fabricius as “ nivea”’ is contained
in the British Museum (Banks’s Collection), it is not the type of A. niveus, because the species
was described from specimens in the collection of D. Koenig. The real types (two specimens)
are now kept in the Zoological Museum of the University of Kiel, and seem to represent a different
species from A. niveus auct. Even A. niveus auct., which is known to be widely distributed
throughout Southern Asia and Central and Southern Africa, seems to be a composite species, as
has already been suggested by Hutchinson (Ann. Mag. Nat. Hist., (10), i, p. 164, 1928). If the
Fabrician types and Fieber’s types in the Zoological Museum of the University of Berlin could be
carefully studied, some changes in the nomenclature of this genus would undoubtedly become
necessary.
* These specimens were identified by H. M. Hale as “ Anisops doris Kirkaldy,” but A. doris
is much larger than this species (“* g 83-9 mm., 9 8-9} mm.’—Kirkaldy, Wien. Ent. Zeit., xxiii,
78 INSECTS OF SAMOA.
collection, one of which at least agrees very well with Distant’s description and
figure, while the other is not quite identical in the colour of the scutellum. The
latter character is very variable in many species of the genus.
Comparing the specimens from Samoa with those from New Caledonia I have
identified them .as Anisops cleopatra Distant. I give below some average.
measurements of the specimens
to facilitate the identification of
the species. (30 units = 1 mm.)
Length of body: 35-5 mm., 9 6 mm.
Breadth of vertex at the anterior
margin viewed from above: ¢ 6
units, 2 8 units.
Breadth of synthlipsis: ¢ 3 units,
© 4 units.
Anterior tibia: ¢ 31 units, 2 30 units.
Anterior tarsus: ¢g 20 units, 9 14+9
units.
Intermediate tibia: g 33 units, 2 33
units.
Intermediate tarsus: g 15+11 units,
© 16+12 units.
Text-Fia. 5.—Amsops cleopatra Distant. a, anterior posterior tibia : 3.50 units, 253 units.
tibia and tarsus of ¢; 0b, the same of 9. Posterior tarsus: ¢ 19-121 units, 2
20-+24 units.
De
47
10. Anisops leucothea, sp. n. Text-fig. 6.
3g. Shining, pale sordid yellow or pale sordid brown like many other
species of Anisops. Eyes pale brown. Base of scutellum more or less obscurely
dark. Under side of abdomen black.
Head including eyes slightly broader than pronotum in ¢ and slightly
narrower in 2, with a short median groove in middle of vertex.* Vertex narrowed
posteriorly, more divergent anteriorly in 2 than in g. Breadth of vertex at the
anterior margin seen from above: synthlipsis = 7:5 in g, 9:5 in 9. Eyes
more prominent in g than in 9. Pronotum much longer than head seen above,
anterior margin more convexly sinuate than posterior margin. Scutellum
p. 112, 1904), and the eyes in the g, contrary to what is the case in the present species, almost
touch each other at the base (“am Grunde fast zusammenstossend,” Kirkaldy, loc. cit.).
* The term “ vertex” (Kirkaldy, 1904) is equivalent to “ notocephalon ”’ (Kirkaldy, 1897).
See Kirkaldy, Wien. Ent. Zeit., xxiii, p. 93, 1904.
AQUATIC HETEROPTERA. 7
broader than long, about one and a third times as long as pronotum in both
sexes. Shape of the anterior tibia and tarsus in both sexes is as shown in the
text-fig. 6. Average dimensions: (30 units = 1 mm.).
Length of body: § 8 mm., 28 mm.
Breadth of body: g 1-9 mm., 9 2-1 mm.
Breadth of vertex at anterior margin, seen
from above: ¢ 7 units, 9 9 units.
Breadth of synthlipsis: $5 units, 9 5 units.
Length of head: ¢ 25 units, 2 20 units.
Length of pronotum: ¢ 33 units, 2 31 units.
Anterior femur: 4 35 units, 2 34 units.
Anterior tibia: ¢ 44 units, 2 44 units.
Anterior tarsus: $27 units, 917-++10 units.
Intermediate femur: 9 48 units, 9 47 units.
Intermediate tibia: ¢ 45 units, 9 44 units.
Intermediate tarsus: 3 20-+14 units, 2 20
+15 units.
Posterior femur: ¢ 75 units, 2 73 units.
Posterior tibia: ¢ 63 units, 9 63 units.
Posterior tarsus: ¢ 20+25 units, 9 22
25 units.
TExt-FIG. 6.—Amisops leucothea Ksaki, sp. n.
a, anterior tibia and tarsus of g; 6, the
same of Q.
‘Upolu: Mulifanua, type 3, type 2, 6 33, 8 2°, paratypes, 9.xi.1925.
In general appearance, this species is very similar to Anisops assumilis
B. White, of New Zealand, but differs in having a shorter pronotum. The
following measurements are taken from the types of Anisops assiemilis B. White,
which are now preserved in the Perthshire Natural History and Antiquarian
Museum, Perth. (30 units = 1 mm.)
Length of body: $65 mm., 9 8 mm.
Breadth of vertex at the anterior margin seen above: ¢ 11 units, 2 9 units.
Breadth of synthlipsis: ¢7 units, 2 5 units.
Length of head: ¢ 25 units, 9 17 units.
Length of pronotum: 3 42 units, 2 33 units.
Anterior tibia: g¢ 40 units, 2 30 units.
Anterior tarsus: g 45 units, 2 19-++10 units.
In A. assimalis, the head and pronotum are comparatively much wider
in the § than in the 9, but there is apparently no distinct sexual difference in
the ratio between the breadth of the anterior margin of the vertex and the
synthlipsis, as in A. leucothea : the ratios are as follows :
A. assimilis : 11:7, 99:5.' A..leucothea: 67:5, 29:5.
Tis 2 3
80 INSECTS OF SAMOA.
LIST OF TEXT-FIGURES.
Text-fig. 1. Microvelia samoana Esaki, sp. n., 3.
» 2. Ventral view of the genital segments of Limnogonus species, shown upside down,
semi-diagrammatic.
a, Limnogonus luctuosus (Montrouzier), 3.
b, oe) > Q.
c, Lamnogonus pacificus Esaki, sp. n., 3.
d, 99 99 2.
e, Limnogonus fossarum (Fabricius), 3.
Ff; ” 9 eh :
. Limnogonus buxtont Esaki, sp. n. a, g, dorsal view; 6, antenna; c, 4, genital
segments, ventral view, shown upside down; d, the same of .
» 4. Limnogonus hopkinsi Esaki, sp.n. a, antenna; 6, ¢ genital segments, ventral view,
shown upside down; c, the same of 9.
» 5. Anisops cleopatra Distant. a, anterior tibia and tarsus of J; b, the same of 9.
» 6. Anisops leucothea Esaki, sp. n. «a, anterior tibia and tarsus of ¢; 6, the same of 9.
.
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PRINTED IN GREAT BRITAIN BY WILLIAM CLOWES AND SONS, LIMITED, LONDON AND BEOCLES.
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INSECTS OF SAMOA
AND OTHER SAMOAN TERRESTRIAL
ARTHROPODA
LIST OF PARTS AND SYSTEM ‘OF PUBLICATION a
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Orthoptera and Dermaptera.
Hemiptera.
. Lepidoptera.
. Coleoptera.
Hymenoptera.
Diptera.
Other Orders of Insects.
Terrestrial Arthropoda other than Insects.
ist “The Wark will be pabtichcd at intervals’ in the foun of numbered fascicles.
i an individual fascicles may contain contributions by more than one
author, each fascicle will be so arranged as to form an iilearsl portion of one or
ae other oi the ae specified above.
INSECTS OF SAMOA
AND OTHER SAMOAN TERRESTRIAL
ARTHROPODA
“List of Fascicles issued to 23rd June, 1928 —
: Date Issued.
Insects of Samoa and other Samoan Terrestrial Arthropoda. ve I and eae
2 (in envelope). 1927, 4to. 6d. ee 26th February, 1927.
Part II. Hemiptera. : ee i Se
Fi ae 1. Tule By F. Mae: 25 text-figures. Paylide ( (Chermid). By
j rawford. 4 text-figures. C idide a t re
By F. Laing, MA, BSc. 3 cexthgares, Pp: 15) (927 eo Gd 8 25th June Waa
Fasc. 2. Cercopide. By V. Lallemand, M.D. 10 text-figures. Cicadide. By. ine
yers, Sc... 22 text-figures. Aquatic and Semi-aquatic ee sini hi Faso Be
sy Prof. Teiso Esaki. 6 text-figures. Pp. 47-80. 1928, 4to. a 6d. ? hinges 23rd June, 1928. —
Part III. LEepipoPTera. sash eee
Fase. |. Butterflies of S nd some neighbouring Island-groups. B GH py) ee Wa a
Hoskie MAU EE. Sol icaehoare and 4 Ae Pp. 1 en a} to, 5s. 9th April, 1927.
Fasc. 2. Micro-Lepidoptera. By Edward Meyrick, B.A., FRS. Pp. 65-116.
1927, 4to. 2s. 6d. 28th May, 1927.
Foie 3. Gomeride. (By Lone Be P t. FES, 2 textfgur dt plate. 5, Oa A era
‘Pp. 117-168. 1928, 40, Tae ule Es Dai March dee
Part IV. CoLEopTEra. — es
Fasc. 1. Carabide. By H. E. Andrews. 9 | Dytiscide. By A.
Zimmermann. 2 text-figures. Staphylinidee. y meron, M.B. 2 text- SAE a an
gures. Hydrophilide. By A. d Orchymont. | text-figure. Clavicornia and bre UE Ben’
-Lamellicronia. “By G. J. rows 13 text-figures. Pp. 1-66. 1927. 4to. 3s. 19th December, 1927.
Fasc. 2. _Heteromera, Bostrychoidea, Meecode a and Buprestide. ByK.G. Tier amese vie
Blair, B.Sc. 14 ee Elaterid, a H. van Zealuvenbere. 10.
text-figures. Melee z ee z). By ay Fleutiaux.. Cerambycide. By
hr. Aurivilli ae late. Brent idee. By R. Kleime. 4° text-figures. oF ara
Anthribide. By Kark Jordan, Ph.D. 11 eee Proterhinide.. ey, pS a) BP aaraaitint )
R.C. L. Perlans, D.Sc, F-R-S. Pp. 67-174. 1928. nee 3s. 25th February, 1928.
Part V. Hymenoptera.
Fasc. 1. Apoidea, Sphecoidea, and Vonact By R. IG; L. Perkin, D.Sc. ss
F.RS., and_L. Evelyn Cheesman, F-E.S., F.Z.S. 12 text-hgures. Larride. BY (ieee ant ae
By Francis X. Williams. 12 text-Agures, oe os Dr. F. Santschi. SH ss! vt
9 text-figures. Pp. 1-58. 1928, Ato. 5s. ae Dth February, 1928.
Part VI. Duprera. : eye
Fasc. 1 Streblide and N abi ‘By L. Fal 7 text-f Hy aye been
heed: By GF. Fave cide, By fe ae 1927, figures, eee 23rd July, 1927.
F 2. Nematocera. By F.W.Edwards, M.A. 20text-fisures. Cecid %
By ALF Bames BA, PRD. 4temhgures, Pp. 25-108 ee 23rd June, 1928.
Part VII. Orner OrbeErRS OF ace . Peaks :
F a a ae Hels ee oe eae era Gull, af Sent and Gyiicaiee 5 h
t t Lt-Col. F. 3 text- 4 en ian
Ped. 198 do, 2s 6 oe a | 28th May, 1927. kes
Fasc. 2. ance By St 13 Tillyard, Sc.D. (Cantab,), FR. Se pet i i Raw
By Rehards Baul EROE ELS. te ‘ ne AR 1928, to, ae
t xt T te thang
bd yeaa "23rd June, 1928,
Part VIII. TerrestRidL ARTHROPODA OTHER THAN INSECTS.
Fasc. 1. Isopoda Terrestria. By Harold G. Jackson, D.Sc. 2 plates. Scor-
ines ea. ee Eat: oy oe Parnes yee raster at ut
text-figur rine t rst. 2. text- = x AS
ry Ms ema aaa Rp TaN M IS G1 hy 23rd July, 1927.
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