List of Fascicles issued to 26th July, 1930 (continued) :—

Part IV. CoLeopTerRA.

Fasc. 1. Carabide. By H. E. pads: HR text-figures. Dytiscide. By A. Date Issued.

ane Lk 4 tin feutes ide yd 2. By y M. pany ce 2 ce

r ir t. text- mi

[erneliiconsies 1B Vee eT Aci. iaicreheures: | “Pp ee one "35. 19th December, 1927.

Fase. 2. ate. Bostrychoidea, pe ecodenaas = Buses: ByK.G

Blair, B.Sc. text-figures. Elateride. . van Zwaluwenberg. 10

text-figures. Melande te, Bred By aR Fleutiaux. Cerambycide. By

Chr. Fears late Bet By R. Kleine. 4 text-figures.

ae Kare ordan, Ph.D. 11 text-figures. Proterhinide. By

CL Pate D'Sc., F-R:S. Be 67-174. 1928, 4to. 5s. 25th February, 1928.

Face 3) Theeade By Ko Go Blin BSe- Veexehee 16 fae .

"By S. Maulik, MLA, {8 text-figures. Pp. 175-215. 1929, 4to. 2s. 6d. 23rd February, 1929.

Guc A Pilg neudes and Seclvidie By C.F Cl Been De 13 Le ive

“Feares. Pp 217-248, 1929, Ato, 2s, 6d. Chae GrW nome ye

Part V. HyMENOPTERA.

Fasc. 1. Apoidea, Sphecoidea, and Vespoiden: By R. C. L. Perkins, D.Sc.,

F.R.S., and_L. Evelyn Cheesman, F.E.S., F.Z.S. 12 text-figures. Larride.

By Francis X:"Williams. 12 text-figures. Formicide. By Dr. F. Santschi. |

tat igurer’ Po a8) WOM den Se ts Bane a ABE

PART VI. Duprera.

F; 1. Streblide and Nycteribiide. By L.Falcoz. 7 text-figuri Hi

erade: By GE Foe Biese betes Pp. 21. 1927, 4to. nes eae 23rd July, 1927.

Fasc. 2. NEG ea By F.W.Edwards, M.A. 20text-figures. Cecidomyiine. ig

By HF. Barnes, BA., Ph.D. 4 text-figures.. Pp. 23-108. 1928, 4to. 5s. 23rd June, 1928.

eee Bs Spatenonde! Tabanide and Asilide. By Gataide Ricardo. 6 text-

figures. Larve o ames z. P. A: Buxton, M.A. 2 tet hele:

Se.

Dolicho a By C 8 text-figures. Sarcophagide.

P. A. , MA. aren: M B R- Malloch,

Pp. 109-175, "1929, dto, 5s, gure, Musride. "By J, R./Malloe 11th May, 1929.

Fasc. 4. Empididae and Pipunculidae. By J. Collin. 7 text-figures.

Pee eae By fh Millet es ‘Gust Be Tres eteat oS.

ae eprom ae y alloc ext-higures. Fp H 27th Joly, 1929,

Fasc. 5. Ortalidac, By J’. R. Malloch. 6 text-f Calliphoridae. B

ee Melee Bo Hea Od ee tee a Oat Wiaree ean

Part VII. Orner Orpers oF He

webs le eects: Fomily Ter ne By GecaldtE Etil ES. ee and eh

t ta Lt-C F. * text- :

a i i ae ie ins eer 28th May, 1927.

Fasc. 2. Plectoptera. By R. 12: Tillyard, Sc.D. (Cantab.), F.R.S., and J. A.

By RehardS BoE RGR ELS. Ben es ee |

Denes ee eee "23rd June, 1928.

ee zx alongs By J Hea DSc. 2 text-figures. Anoplura. By g

Buxton, richoptera. y Martin ae Mosely. gure. (aa

Neier By P. Esben-Peter | text-figure and 2 plates. Apterygota. Sikes,

By George H. Carpenter, D. oe We gtevemeares! PS ate ic. : 1928, 4to. 2s. 6d. 28th July,§1928. ry

Part VIII. TerRrestRIAL ARTHROPODA OTHER THAN INSECTS.

Fasc. 1. Isopoda Terrestria. By Harold G. Jackson, aie 2 plates. Scor-

pionoidea. paae ie Recep at 7 na

Pak ae carina. By Stan ey Hirst text-figures. Pp. I-. 23rd July, 1927.

Fasc, 2. Myriopoden (M da). By C. Att 4 text-

st. 2: Mutiopoden (Myrigpoda), By C. Attems. 4 textures, Araignées 5) hone 1900,

A : Oe

je” BRITISH MUSEUM (NATURAL HISTORY)

INSECTS OF SAMOA

AND OTHER SAMOAN TERRESTRIAL

ARTHROPODA

PART IL HEMIP'TERA

FASC. 4. Pp. 163-494

CICADELLIDAE (JASSIDAE)

By HERBERT OSBORN, Ohio State University,

Columbus, Ohio

WITH FIFTEEN TEXT-FIGURES

LONDON:

PRINTED BY ORDER OF THE TRUSTEES OF THE BRITISH MUSEUM

SOLD AT

Tue Britiss Museum (Naturat History), CromwEeLL Roan, $.W.7

AND BY

B. Quaniten, Lrp.; Dunavu & Co., Lrp.; anp Tur Oxrorp University PREss.

: 1934

Issued 27th January, 1934.] [Price Two Shillings.

INSECTS OF SAMOA

AND OTHER SAMOAN TERRESTRIAL

| ARTHROPODA

Although a monograph, or series of papers, dealing comprehensively with

the land arthropod fauna of any group of islands in the South Pacific may be

expected to yield valuable results, in connection with distribution, modification

_ due to isolation, and other problems, no such work i is at Piescni in existence. .

In order in some measure to remedy this deficiency, and in view of benefits:

directly accruing to the National Collections, the Trustees of the British

Museum have undertaken the publication of an account of the Insects and other

Terrestrial Arthropoda collected m the Samoan Islands, in 1924-1925, by

Messrs. P. A. Buxton and G. H. E. Hopkins, during the Expedition of the ;

London School of Hygiene and Tropical Medicine to the South Pacific.

Advantage has been taken of the opportunity thus afforded, to make the studies

as complete as possible by including in them all Samoan material of the groups |

concerned in both the British Museum (Natural History) and (by ee) of

the authorities of that institution) the Bishop Museum, Honolulu.

It is not intended that Coe ae to the text shall be confined to the

Museum Staff or to any one nation, but, so far as possible, the assistance of the

leading authorities on all groups to be dealt with has been obtained.

The work is divided into nine | “Darts'e: (see Ds 3 of wrapper), of oe

the first eight are subdivided into “ Fascicles.” Each of the latter, hich

appear as ready in any order, consists of one or more contributions. On the

completion of the systematic portion of the work it is intended to issue Ais

general survey (Part IX), summarising the whole and drawing from it such |

conclusions as may be warranted. a

A list of Fascicles already issued will be found on pp. 3 and 4 of this wrapper.

N. D. RILEY, —

) Keeper of Entomology.

British Museum (INarurat History). .

CromweLt Roap, S.W.7.

INSECTS OF SAMOA

Part II. Fasc. 4

HEMIPTERA—CICADELLIDAE (JASSIDAE)

By Herpert Oszorn, Ohio State University, Columbus, Ohio

(With 15 Text-figures)

THE collection of Cicadellidae received includes collections made by Dr. P. A.

Buxton and Mr. G. H. E. Hopkins in 1924-25, together with collections by

O. H. Swezey and G. P. Wilder and EH. H. Bryan. It represents the largest

series known from this group of islands and is of special interest in comparison

with the Fiji, Society and Marquesas groups and their possible affinities with

Oriental or Malayan species, and also with the Hawaiian Islands to the north.

All these groups of islands have had a long isolation, with evidently abundant

time and opportunity, so far as food plants and other ecological factors are

concerned, for the formation of species or varieties from what must have been

early introductions from more fully populated regions. While it would be

intensely interesting to attempt the tracing of affinities and lines of migration

it is plain that this would involve much pure speculation and, with the material

at present available, so little of certainty that this had best be left for future

students. To the extent that we have species occurring in adjacent territory

or for which there may be evident affinities, we can make suggestions in con-

nection with particular species. In a general way we can be assured that the

Samoan Cicadellid fauna has its affinities with other islands of the South Pacific,

but we have a number of forms not known from other regions and for them we

may postulate an endemic evolution.

The literature for Cicadellidae of Samoa is very meagre and it would seem

that little attention to this group has been given in the past. Only scattered

records for individual species have come to my notice and, while I have not

i 4

164 INSECTS OF SAMOA.

had opportunity to consult all possible sources, I have carefully searched through

all the publications that would be expected to include such records. The works

of Walker, Stal, Signoret, Melichar, Distant, and Kirkaldy have been the main

sources for descriptions of species from the broader Oriental faunal region,

including India, Ceylon, Malaya, Hast Indies, and Philippine Islands.

The figures illustrating the paper have been drawn by Mrs. Celeste Taft.

Types collected by Buxton and Hopkins are placed in the British Museum

(Natural History). Those collected by Bryan, and by Swezey and Wilder, are

in the collection of the Bishop Museum, Honolulu. Duplicate or paratype

specimens when available are in the author’s collection at the Ohio State

University.

Agalliopsis Kirkaldy.

Agalliopsis Kirkaldy, Bull. No. 3, Dw. Ent. Exp. Sta. H.S.P.A., p. 31 (1907). (Based on

Div. II of Osborn & Ball.)

Agallia (group II) Osborn & Ball, Proc. Dav. Acad. Sci., VII, p. 46 (1897).

1. Agalliopsis fasciatus, n. sp. (Text-fig. 1).

Small, head decidedly broader than pronotum, eyes prominent; vertex

i angulately arched, very short, mid-length

less than length at eye; hind border

sinuate; ocelli well below border, near

together; front narrowing at apex;

clypeus longer than broad, sides parallel.

Pronotum produced anteriorly much

beyond line of eyes, more than half as

long as broad, hind border concave.

Hlytra narrowing apically to rounded tip,

venation obscure. Female, last ventral

segment truncate, faintly sinuate.

Colour: fuscous above, pale below ;

vertex basally, except next the eye,

ocellar dots, pronotum, except central

BO”

Text-ric. 1.—Agalliopsis fasciatus, n. sp.: and lateral fasciae, scutellum in basal

a, dorsal view; 6, face; ¢, female angles and elytra, except costa and

ponies. claval stripes, dark fuscous ; face, except

angular projection from vertex and the ocellar dots, a median narrow and wider

lateral stripes on pronotum, disc of scutellum, broad costal border and two.

HEMIPTERA—CICADELLIDAE (JASSIDAE). 165

stripes on clavus and the claval suture, light yellowish brown. Beneath pale,

disc of venter infuscate.

Length, female, 5-5 mm.

Described from one female (holotype) Malololelei, Upolu, 2,000 ft.,

14-30.vu.1924, Buxton and Hopkins.

This species has the pronotum more decidedly produced than in any species

known to me and in this respect resembles Tartessus, but the ocelli are distinctly

below the margin of vertex and in size and general facies the species falls in the

Agallia group and may be placed in Agalliopsis as now constituted. The

pronotum is even more produced anteriorly than in the genotype species novella,

which is a common North American species. In general facies the species

resembles Agallia atrovenosa Melichar, from Ceylon, but in details of picture, as

well as the pronounced difference in form of head and pronotum, it is quite

distinct.

Genus Bythoscopus Germar.

Bythoscopus Germar in Silberman, Rev. Ent., I, p. 180 (1833).

Batracomorphus Lewis, Trans. Ent. Soc. London, I, p. 51 (1834).

Macropsis Amyot & Serville, (nec Lewis), Hist. Nat. Ins. Hémip., p. 585 (1843).

Strangania Stal, Rio Jan. Hem., I, p. 49 (1858).

Pachyopsis Uhler, Bull. U.S. Geol. & Geog. Survey, UI, p. 466 (1877).

Gargaropsis Fowler, Biol. Cent. Am. Rhynchota Homop., I, p. 167 (1896). Genotype B. lanio (L.)

Body depressed, face retracted, vertex short, ocelli on face below margin.

This genus is world-wide in distribution, and many of the species in widely

scattered localities have many features in common so that separation of species

is difficult. The species of the Pacific island groups are no exception, and it is

quite possible that some of those given names here may be considered identical

with species already described from other regions. However, if we bear in mind

the isolation they must represent it is less likely to result in confusion to give

them distinctive names than to refer them to species known in other regions.

2. Bythoscopus chlorophana (Melichar).

Pachyopsis chlorophana Melichar, Hom. Faun. Ceylon, p. 153 (1903).

Bythoscopus chlorophana Distant, Faun. Brit. Ind. Rhynchota, IV, Pt. 1, p. 191 (1907).

This is a pale green species sometimes faintly suffused with reddish on the

head and pronotum but without markings, except a small blackish spot at tip

of clavus.

Length: female, 5 mm.; male, 4-5 mm.

166 INSECTS OF SAMOA. ~

Five specimens: one female, Upolu, 2,000 ft., vii.1924; one male, Upolu

Island, Vailima, 12.x11.1925, Buxton and Hopkins; one male, Upolu, Tuaefu,

9-16-1925, Swezey and Wilder; one male, Malololelei, 2,000 ft., 1.1.1925 ; and

one female, Upolu, Apia, 2.111.1924, Buxton and Hopkins.

3. Bythoscopus angustatus, n. sp. (‘Text-fig. 2).

Small, narrow ; head scarcely as wide as pronotum ; vertex short, evenly

rounded ; front tapering rapidly to a narrow clypeus; clypeus somewhat

inflated on basal one-half, sides

parallel; lorae distant from mar-

gin of cheek ; cheek margin evenly

rounded ; pronotum three times as

long as vertex, minutely striate ;

elytra hyaline, veins margined with

minute punctures and hairs very

minute or wanting.

Male, last ventral segment

elongate, somewhat produced at

the middle, pygofers short, plates

hidden.

Length: 4 mm.

Described from two males:

one (holotype) Neiafu, Vavau,

Tonga, 1.11.1925, Hopkins; and

TEXT-FIG. 2.— Bythoscopus angustatus, D. sp. one (paratype) Tutuila, Pago Pago,

a, dorsal view ; b, face; c, male genitalia. 9-24-1923, Swezey aad Wilder:

This species is smaller, the colour paler, and the elytra more hyaline than

in chlorophana.

4. Bythoscopus tutuilanus, n. sp. (‘Text-fig. 3).

Broad, robust, head scarcely as wide as pronotum, very short; vertex

with parallel borders, rounded to the front; front narrowed abruptly to short

clypeus ; lorae small; cheek margins sinuate ; pronotum about five times as

long as vertex ; elytra opaque, minutely and sparsely hairy.

Female, last ventral segment a little longer than preceding, truncate or

slightly concave.

HEMIPTERA—CICADELLIDAE (JASSIDAE). 167

Male, last ventral segment long, rounded behind, as long as pygofers ;

pygofers short, ventral margins closely appressed.

Colour dull stramineous, probably green in life, without markings except

minute blackish point at tip of clavus.

Length: female, 5mm.; male, 4-25 mm.

Described from four adults: one female (holotype) Tutuila, 1,070 it.,

eastern end of island, Keilers, 6-21-1918 ; and two males (allotype and paratype)

TExtT-FIG. 3.—Bythoscopus tutuwilanus, n. sp.: a, dorsal view; 6, face ;

c, female, and d, male genitalia.

Tutuila, 760-900 ft., Kellers, iv.1918; one male (paratype) Tutuila, Samoa,

1,200 ft., Kellers, xii.1918. Three nymphs referred to this species with the

same data as the last specimen, have the vertex slightly produced, about one-

third as long as pronotum, the wing pads extending to first abdominal segment ;

the body with short appressed hairs, probably about the third instar.

This species is more robust than chlorophana, with the head decidedly

shorter, and the stramineous colour more intense.

168 INSECTS OF SAMOA.

5. Bythoscopus buxtoni, n. sp.

Body elongate ; head as wide as pronotum ; vertex considerably arched,

borders parallel; front flattened, abruptly narrowed from the antennae ;

clypeus narrowed toward the apex with polished elevation basally ; lorae small ;

margin of cheek sinuate below the eye, subangular at the sides ; pronotum three

times as long as vertex ; elytra shiny, sparsely hairy and punctate.

Female, last ventral segment fully twice as long as preceding, slightly

emarginate and very broadly notched at the middle. Pygofers broad, sparsely

ciliate.

Colour: dull greenish ; ocelli red ; elytra hyaline; pygofer with a broad

fulvous patch either side.

Female: length, 6-25 mm.

Described from two specimens: female (holotype) Upolu, Malololelei,

2,000 ft., 25.x1.1924, Buxton and Hopkins; one with same locality, 23.x1.1924,

abdomen wanting ; and one female Tutuila, Afono Trail, 9-25-1923, Swezey and

Wilder, referred here, but the female pygofer lacks the coloured patch at the

side.

6. Bythoscopus laticeps, n. sp. (Text-fig. 4).

Broad ; head narrower than

pronotum ; vertex very short ;

front much broader than long ;

antennae under distinct ledges ;

\\ clypeus half longer than wide ;

lorae distinct from margin of

cheek ; cheek broad, the mar-

gin scarcely sinuate below eye ;

pronotum more than twice as

wide as length at centre, hind .

margin truncate ; elytra slightly

longer than abdomen, smooth,

minutely punctate. Female,

last ventral segment at apex

concave; pygofer short,

Text-ria. 4.—Bythoscopus laticeps, n. sp.: a, dorsal view ; scarcely exceeded by the ovi-

b, face ; c, female genitalia. positor.

DON OE

I = )

HEMIPTERA—CICADELLIDAE (JASSIDAE).

169

Pale green, stramineous, unmarked. Length, 5 mm.; width of pronotum

at base, 1°75 mm.

One specimen: female (holotype) from Tutuila, 900-1,200 ft., centre of

island, 6-30-1918, Kellers.

This species is proportionately broader than related forms and the broad

face and comparatively short elytra and lack of colour markings serve as

differential characters.

7. Bythoscopus hyalinus, n. sp. (Text-fig. 5).

Head distinctly narrower than pronotum, rather prominently rounded,

vertex with parallel borders ;

face broad ; clypeus slightly

narrowing to rounded tip ;

lorae broad, approaching

border of cheek; pronotum

nearly four times as long as

vertex, over twice as broad

as long, concave behind ;

elytra but little longer than

abdomen, costa strongly

curved. Female, last ventral

segment quite deeply con-

cave, as long as preceding

segment; pygofer long,

scarcely extending to tip of

ovipositor.

Pale green without

marking. Length, 6-5 mm.

One specimen, female

(holotype) from Tutuila,

1,200 ft., 7-21-1918, Kellers.

Text-Fic. 5.—Bythoscopus hyalinus, n. sp.: a, dorsal view ;

b, face ; c, female genitalia.

The larger size, pale translucent colour and the strongly curved elytra

appear sufficient to separate this from chlorophana, to which it seems related.

170 INSECTS OF SAMOA.

Genus Idiocerus Lewis.

Idiocerus Lewis, Trans. Ent. Soc. Lond., I, p. 47, 1834. Genotype I. adustus H.S.

8. Idiocerus tutuilanus, n. sp. (Text-fig. 6).

Head distinctly wider than pronotum ; eyes large; vertex short, scarcely

as long as width between eyes, sinuately narrowing from antennae to clypeus ;

clypeus half longer than width

at base, expanded apically ; apex

broadly rounded ; lorae fusiform ;

margin of cheek nearly straight ;

pronotum nearly three times as

wide as length at middle; hind

border slightly concave; elytra

with obscure venation; four

apical cells and broad appendix.

Male valve rounded behind, plates

elongate.

Pale yellow, with pronotum

except two oval spots, scutellum

except central stripe and apex and

Text-Fic. 6.—Idiocerus tutwilanus, N. 8p.: 4, dorsal inner claval border and irregular

Weis), dee ormale gente stripes toward apex, black or dark

fuscous. Beneath yellowish, legs yellowish, hind femur and tarsal claws blackish.

Length, 4 mm.

Three specimens: one male (holotype) Upolu, Vailima, 1.1925, 1,200 ft. ;

one male (paratype) Malololelei, iv.1925, Buxton and Hopkins; and one male

(paratype) Tutuila, xii.1918, Kellers.

This species will fall into a subgenus in which the male antennae do not

have the flat discs of typical Jdcocerus.

Xestocephalus Van Duzee.

Xestocephalus Van Duzee, Trans. Am. Ent. Soc., XIX, p. 298, 1892 ; Bull. Buffalo Soc. Nat. Sci.,

V, pp. 197, 214 (1894). Genotype pulicarius Van. D.

“ Head tumid and rounded before ; ocelli placed on the rounded anterior

edge superiorly, distant from the eyes ; vertex convex, closely evenly punctured,

without longitudinal rugae.” (Van Duzee.)

HEMIPTERA—CICADELLIDAE (JASSIDAE). 171

Species included in the genus have now been recognised from Eastern

Palaearctic, Ethiopian and Oriental Regions, as well as the Nearctic and

Neotropic.

9. Xestocephalus pardalinus Distant.

Xestocephalus pardalinus Distant, Fauna Brit. Ind. Rhynchota, IV, Pt. 2, p. 348, 1908.

In this species, as defined by Distant, there is a distinct colour pattern and

a well-marked dark line above and below the ocelli, including a pale area within

which the ocelli are located. Except for the more distinct colour marking and

a slight difference indicated in size (which may be a variable character) there is

little to separate the species from apicalis of Melichar. However, specimens

agreeing very perfectly with Distant’s description (and type in British Museum)

of specimens from Ceylon, Maskeliya, and bearing labels “ Upolu Is., Malololelei,

2,000 ft., x11.1925, and vi.1924, P. A. Buxton and G. H. Hopkins,” and one

“ Savai, Saitalus, v-17-24, Rain Forest, 2,000-4,000 ft., E. H. Bryan collector,”

are among the specimens referred to me and I am therefore including the record

under this name. If future studies prove the species to be identical Melichar’s

name has priority.

10. Xestocephalus apicalis Melichar.

Xestocephalus apicalis Melichar, Hom. Fauna, Ceylon, p. 207 (1903).

Xestocephalus apicalis Distant, Fauna Brit. Ind. Rhynchota, IV, Pt. 2, p. 349 (1908).

The Samoan specimens agree well with Melichar’s description with only

slight variations in extent or intensity of colour pattern. Four from Malololelei,

Upolu, 2,000 ft., vi.1924, Buxton and Hopkins ; one Malololelei, Upolu, 2,000 ft.,

vi.1924; one Tutuila, 760-900 ft., Kellers, iv.1918 ; one Tutuila, Afona Trail

9-25-1923, Swezey ; one Savail, Salailua, v-17-1924, Rain Forest, 2,000—4,000 ft.,

Bryan.

Considering the apparent wide distribution now accorded the species and

the habitat indicated, the variation noted does not seem unusual. The specimens

listed above merge almost into the paler form of what I have taken to be the

pardalinus of Distant. See ante.

172 INSECTS OF SAMOA.

11. Xestocephalus tutuilanus, n. sp. (Text-fig. 7).

Head narrow, scarcely as wide as pronotum. Vertex as long at middle as

at eye, subangulate, rounded to front ; front broad, narrowed abruptly to base

of short clypeus. Clypeus about as broad

as long, sides parallel, apex truncate.

Pronotum a little longer than vertex, trun-

cate behind; elytra with distinct veins.

- Female last ventral segment long, twice as

long as penultimate, faintly simuate and

with shallow notch at middle of hind border.

Male valve hidden, plates elongate tri-

angular, tips acute.

Colour pattern similar to pulicarius and

pardalinus. Vertex pale with two dark

spots on disc and dark borders to lighter

TEXT-FIG. 7.—Xestocephalus tutwlanus, spot enclosing ocelli. Pronotum brown with

n. sp.: a, dorsal view; 0 face; c,

female, and d, male genitalia. rounded spots anteriorly and two larger

ones on the disc near the hind margin and a

broken median line, yellowish. Scutellum yellow with darker areas in basal

angles. Hlytra yellowish subhyaline with distinct brown or fuscous spots as

shown in the figure. Beneath light brown.

Length: female, 2-5 mm.; male, 2:25 mm.

Two specimens: female (holotype and paratype) Tutuila, Pago Pago, one

9-9-1923 and one 9-24-1923, Swezey and Wilder ; one male (allotype) Lau, Fiji,

Namuka, 8-13-1924, Bryan ; one male (paratype) Tutuila, 760-900 ft., Kellers,

iv.1918 ; one female (paratype) Pago Pago, Tutuila, 0-300 ft., Kellers, iv.1918.

12. Xestocephalus piceatus, n. sp.

Head slightly narrower than pronotum, rounded before, vertex broader

than long, scarcely longer at middle than at eye. Pronotum twice as wide as

long, hind margin slightly convex; elytral veins obscure. Male valve very

short, almost hidden under preceding segment. Plates long, tapering to blunt

upturned apices, clothed with long cilia.

Dark piceous above except two whitish subhyaline spots on costa near

HEMIPTERA—CICADELLIDAE (JASSIDAE). 173

apex and two faint whitish areas on inner margin, one at tip of clavus and the

other half-way on appendix and invading inner apical cell. Wing veins dusky,

beneath piceous ; margins of ventral segment, basal half of plates and the legs

pale.

Length: male, 3 mm.

One male (holotype) Apia, Upolu, 1.x.1925, Buxton and Hopkins.

This species seems well marked by the rounded border of vertex and the

dense piceous colour.

13. ‘xestocephalus reflexus, n. sp.

Head narrower than pronotum, broadly subangulate. Vertex half longer

at middle than at eye, ocelli close to anterior margin; front broad, narrowed

rather abruptly to clypeus ; cheek convex ; pronotum half longer than vertex ;

elytral veins mostly obscure. Female, last ventral segment long, twice as long

as preceding, truncate behind, faintly notched at middle, pygofer nearly as long

as ovipositor, ciliate on apical half.

Pale brown; the scutellum and elytra infuscate or piceous ; two strongly

reflexed pale spots on costa. Beneath pale testaceous with some piceous

suffusion.

Length, 3-5 mm.

One specimen, female (holotype), Upolu, Malololelei, 24.11.1924, Buxton and

Hopkins.

This might seem possibly to be the female of prceatus, but aside from the .

difference in locality there is a distinct difference in the colour pattern and it

seems wiser to consider them distinct. If additional material or the finding of

the sexes together proves an identity one may be placed in synonomy without

serious consequences, a result that would seem preferable to confusing two

species under one name.

Genus Hecaloidella, new genus.

Allied to Hecalus and Parabolocratus but with different venation. Body

depressed, head thin, margin of vertex acutely angular, almost foliaceous,

rounded in front. Elytral venation distinct, two claval veins and a cross-vein

from outer claval to suture. Three anteapical, four apical cells, the outer one

very broad and its base resting on the nodal outer anteapical. Genotype

Hl, mitida, nu. sp.

174 INSECTS OF SAMOA. -

14. Hecaloidella nitida, n. sp. (Text-fig. 8).

Female: head slightly wider than pronotum; vertex large, scarcely as

long as wide at base, rounded, in female faintly subangulate, margin thin ;

ocelli close to eyes; front broad, sinuate next antennae, tapering sharply to

clypeus ; clypeus longer than broad, widening apically ; lorae broad, almost

touching margin of cheek ; cheek broad, margin

ve faintly smuate. Pronotum scarcely longer than

ee vertex, strongly arched before, lateral margin

| very short, hind border slightly emarginate ;

aid

scutellum wider than long, with a curved in-

dented line behind the half-way point. Elytra

polished, veins distinct, nodal areole shorter than

outer anteapical, outer apical broad, resting on

nodal, outer anteapical and part of middle ante-

i ca apical. Second apical narrow at base, much

Text-ric. 8.—Hecaloidella nitida, Wider at apex. Female, last ventral segment

n. sp.: @, dorsal view, head, about twice longer than preceding, truncate ;

pronotum and scutellum; 6, 3 :

Genie adecmaleneentana pygofer scarcely exceeded by ovipositor and

d, elytron. not reaching tip of elytra.

Colour: vertex, pronotum and face pale

yellowish, translucent. Margin of vertex, scutellum, elytral veins tinged with

fuscous or pale brown. LElytra hyaline with fuscous patches on tips of claval

veins, the discal cross-veins and the veins at base of outer apical areole and

margin of apical cells.

Length, 5 mm

Male. What I am confident is the male of this species, having same locality

of capture, has the vertex rounded and not at all subangulate and the most of

the pronotum and all of scutellum, elytra, and all beneath except the face are

deeply suffused with piceous. The male valve is very small, the hind border

rounded, the plates broad at base, narrowed and acutely pointed at apex.

Length, 4-5 mm

The female (holotype) and male (allotype) are both from Malololelei, Upolu,

2,000 ft., vii.1924, Buxton and Hopkins.

This species seems to bear some resemblance to Parabolocratus wallengrent

Stal, but must, I believe, have different generic position.

HEMIPTERA—CICADELLIDAE (JASSIDAE). 175

Genus Nephotettix Matsumuri.

Nephotettix Matsumuri, Termés ftizetek, XXV, pp. 356 and 378 (1902). Genotype N. apicalis

Motsch.

Related to Thamnotettix.

15. Nephotettix plebetus Kirkaldy (Text-fig. 9).

Nephotettia plebevus Kirkaldy, Bul. No.1, pt.9, Div. Ent. Exp. Sta. H.S.P.A., p. 331 (1906).

This species was described by Kirkaldy from specimens taken in Australia

at various localities and is recognised by the grey colour, dark transverse bar

Trext-ric. 9.—Nephotettix plebeius Kirkaldy: a, dorsal view ;

b, face; ¢, female, and d, male genitalia.

on vertex just behind ocelli and extending the distance between them with a

trace of angular projection at centre. Length: female, 5-5-5-75 mm.; male,

4°75-5:25 mm.

The collection from Samoa includes a large number of specimens from

different localities and indicates a quite general distribution on different islands

176 INSECTS OF SAMOA.

of the group. It is probably a grass feeder, as records include Sporobolus as a

host plant. The locality records are as follows :—

A large series of specimens of both sexes and many nymphs from Tutuila,

collected in March, Sept., Oct., and Dec.

Eleven from Tau, Manua, 17-27.ix.1923; three from Savaii, Safune,

11.v.1924, Bryan; two Upolu, Mulifanua, 16.vu.1925, Wilder; two Apia,

13.1x.1923, Sporobolus, Swezey and Wilder; four Malololelei, xii and 11.1924 ;

four Apia, i, iv.1924, Buxton and Hopkins.

Also there are two smaller males, otherwise apparently identical, from

Neiafu, Vavau, Tonga, 5.11.1925, Hopkins.

This would seem to be one species that has had favourable opportunities

for migration, and it may be that its association with a grass that was scattered

by human agencies was a distinct factor in its distribution.

Drabescus Stal.

Drabescus Stal, Ofv. Vet.-Akad. Férh., XXVII, p. 738 (1870). Genotype D. remotus Walk.

Distribution apparently limited to Oriental Region, with records from

India, Malaya, and the Philippines.

16. Drabescus samoanus, n. sp. (Text-fig. 10).

Head distinctly wider than pronotum ; eyes large ; vertex broad, depressed,

angulate, one-fourth longer at middle than at eye; margin elevated, ocelli

minute on extreme edge half-way from eye to middle ; front depressed at base,

scarcely longer than broad, sides nearly parallel from antennae to abruptly

contracted apex; clypeus long, narrow, widening to apex and with strong

median keel; lorae large, broad, nearly touching border of cheek; cheek

depressed below antennae ; lower part flattened, margin deeply sinuate below

the eye; pronotum twice as long as vertex, hind border concave, disc trans-

versely minutely striate, scutellum minutely granulate, elytral veins punctate.

except in apical cells; claval and discal areoles rugose. Female, last ventral

segment as long as preceding, slightly sinuate and with a shallow notch at

middle of hind border. Male valve small, plates small, triangular, tips acute.

Dark brown, infuscate in typical example, vertex fuscous, pronotum

yellowish on anterior border ranging in extreme forms of males to entire pronotum

and scutellum; scutellum pale yellowish; elytra brownish subhyaline, the

HEMIPTERA—CICADELLIDAE (JASSIDAE). 177

veins dotted with yellow and a paler transverse band before apex of clavus.

Base of front and eyes fuscous; a yellowish band crossing lower part of front

and cheeks below eyes. Clypeus and lorae fuscous. Beneath dark brown, a

yellow or whitish spot on propleura.

Length: female, 9 mm.; male, 7 mm.

Nineteen specimens. Four females (holotype and paratypes) and two

males (allotype and paratype) Tutuila, centre of island, 900-1,200 ft., Kellers ;

TEext-FiG. 10.—Drabescus samoanus, n. sp.: a, dorsal view ;

b, face; c, female, and d, male genitalia.

one male (paratype) Tutuila, Leone Rd., 1x-7-1923, Swezey and Wilder; one

female (paratype) Upolu, Tuaefu, 1x-16-1923, Swezey and Wilder; one female

Savail, Safune, v-3-1924, Rain Forest, 2,000-4,000 ft., Bryan; four Apia,

Upolu; one female (paratype) Pago Pago, 4.x1.1925, Tutuila Island; one

female (paratype) Vailima, Upolu; two females, one male (paratypes) Malolo-

lelei, Upolu, 2,000 ft., 25.vi.1924, Buxton and Hopkins ; two nymphs.

Two nymphs, evidently associated with this species, having date and

locality same as holotype, have the head produced into a narrow snout as long

as eye. The head broader than the pronotum and the abdomen long and

tapering to a very narrow tip.

178 INSECTS OF SAMOA.

Eutettix Van Duzee

Eutetiiz Van Duzee, Psyche, V1, p. 307 (1892). Genotype L. luridus Van Duzee.

17. Eutettix ornatella, n. sp. (Text-fig. 11).

Head as broad as pronotum, short, vertex broadly rounded, slightly longer

at middle than at eye. ‘Transverse line scarcely impressed ; rounded to front ;

front broad at base, narrowed from below antennae ; clypeus rather long, sides

nearly parallel, narrowing

slightly to apex. Lorae broad,

approaching margin of cheek ;

cheek broad, margin slightly

sinuate; pronotum twice as

long as vertex, lateral margins

very short, hind border emar-

ginate ; scutellum small, elytra

with distinct veins and numerous

cross-veinlets. Apical cells four,

apex rounded. Female ventral

segment elongate, sinuate at

side, produced medially, hind

border at centre rounded. Male

valve small, obtusely angulate

Text-rie. 11.—Eutettia ornatella, n. sp.: a, dorsal behind; plates long, narrowed

view ; 6, face ; c, female, and d, male genitalia. behind middle to acute tips :

densely hairy.

Colour pale grey, vertex anteriorly yellow, base tinged with orange, two

dark dots near tip and a series of triangular points along base. Face yellowish-

srey mottled with light fuscous, pronotum irrorate with brown or pale fuscous.

Klytra grey-white with veins and coarse veinlets mostly brown or fuscous,

about four post-nodal oblique fuscous bars. In some specimens a wavy white

patch in distal part of costal areole and outer discal areole. This area otherwise

paler than discal area. Beneath grey, mottled with fuscous.

Length: female, 4mm.; male, 3-75 mm.

Sixteen specimens: nine females (holotype and paratypes) and six males

(allotype and paratypes), one abdomen missing. Upolu Island, Malololelei,

2,000 ft., x11.1925, Buxton and Hopkins.

HEMIPTERA—CICADELLIDAE (JASSIDAE), 179

This species is distinctly more slender than discigutta Walker, with which

it has some characters in common.

Stirellus Osborn and Ball.

Stirellus (subgenus) Osborn & Ball, Ohio Nat., I, p. 250 (1902).

Stirellus Sleesman, Ent. Am., X, p. 124 (1930). Genotype S. bicolor Van D.

In this genus as here recognised the vertex is long and angular, though not

always as long at middle as between the eyes, the elytra with three anteapical

cells, one cross-nervure from inner sector to claval. They are insects of small

size and similar to Deltocephalus in general facies.

18. Stirellus pacificus, n. sp. (Text-fig. 12).

Head wider than pronotum, vertex three-fourths as long as width between

eyes, obtusely angulate ; front long narrowing uniformly to base of clypeus ;

clypeus about twice as long as width at base

and much narrowed at apex; lorae long,

rather narrow, nearly reaching margin of cheek ;

cheek broad, broadly subangulate ; pronotum

arched before, lateral margins short, hind

border concave. LElytra longer than abdomen,

outer sector twice forked, inner apical areole

large, stylate at tip, appendix narrow, extend-

ing to tip. Female last ventral segment long,

concave behind ; male valve very short, plates

broad at base narrowing abruptly from middle

to acuminate tips.

light grey, vertex with two transverse T=*t Ie. 12.—Stirellus pacificus,

n. sp.: @, dorsal view; b, face ;

c, female, and d, male genitalia.

discal spots and faimt oblique dots near apex ;

front with faint lateral arcs. Tergum black

margined with yellow, venter black, last ventral segment pale. Pygofer black

with whitish setae.

Length: female, 3 mm.; male, 2°75 mm.

Twenty-one specimens: six females (holotype and paratypes), four males

(allotype and paratypes) Tutuila, Pago Pago, 9-9-1923, Swezey and Wilder ;

ten specimens (paratypes), one female and five males, Malololelei, Upolu, 2,000 ft.,

gt 2

180 INSECTS OF SAMOA..

vi.1924, Buxton and Hopkins; four females and one male (paratypes) Upolu,

Mulifanua, 7-16-1923, Wilder.

In general appearance this resembles Stirellus (Phrynomorphus) hospes

Kirkaldy, but the pattern on the vertex is different.

19. Stirellus samoanus, n. sp.

Head slightly wider than pronotum; vertex about as long at middle as

width at base, distinctly angulate, nearly twice as long at middle as next to

eye; ocelli close to the eye; front slightly convex, sloping evenly to base of

clypeus ; clypeus narrowing slightly toward apex, half longer than wide ; lorae

large ; margin of cheek sinuate ; pronotum as long as vertex, scarcely concave

behind ; elytral veins distinct, claval veins united by a single cross-vein, one

cross-vein in disc, central anteapical cell narrowed at the middle, four apical

cells.

Male, last ventral segment short, valves small, plates short (apparently with

broken tips) not reaching tip of pygofer. Tip of pygofer with stiff bristles

darkened at the tip.

Pale straw-coloured, the apex of vertex and a line each side of the central

impressed line and margin at eye, figure on anterior part of pronotum and elytral

veins pale yellow, front pale brownish with faint yellowish-white arcs. Abdomen

beneath black, with lateral margins of plates and pygofer pale yellow.

Length, 2-6 mm.

Described from one specimen, male (holotype) Savai, Safune, 4-v-1924,

Bryan.

This species approaches in size Stcrellus (Phrynomorphus) hospes Kairk.,

but is slightly larger, and the picture on the vertex is distinctly different.

20. Surellus affnis, n. sp.

Head wider than pronotum; vertex sharply angulate, length at middle

equal to width between the eyes; ocelli close to the eyes; front narrowed

slightly, curved from antennae to the clypeus; clypeus broad, narrowed slightly

toward the apex, half longer than width at base ; lorae narrowed below ; cheek

margin distinctly sinuate ; pronotum scarcely as long as vertex, slightly concave

behind ; claval veins without cross-vein, one cross-vein on the disc.

Male, last ventral segment scarcely as long as preceding; valves small,

HEMIPTERA—CICADELLIDAE (JASSIDAE) 18]

rounded behind ; plates broad at base, constricted to acute tips, margins ciliate,

slightly exceeded by the pygofer which is quite distinctly ciliate. Pale straw

colour, markings of vertex and pronotum obscure ; elytra hyaline with traces

of fuscous margins in the cells and a distinct fuscous spot at base of middle

anteapical cell. Beneath pale, abdominal segments somewhat suffused, especially

at the sides, with fuscous.

Length, 2-5 mm.

Described from one male (holotype) Tau, Manua, 9-27-1923, Swezey.

Collected from grass.

This species comes very close to Stirellus (Phrynomorphus) hospes Kirk.,

but differs distinctly in the picture of the vertex and the distinct frontal arcs :

it agrees with it in the fuscous spot at base of middle anteapical cell and differs

but slightly in the male genitalia.

I have compared it with one of Kirkaldy’s specimens, probably a co-type,

and I am satisfied that it cannot be considered identical with that species.

21. Stirellus tauensis, n. sp.

Head wider than pronotum; vertex subangulate, wider at base than

length at middle; length at middle one-third longer than at eye, rather

distinctly rounded to front; front distinctly convex, narrowing rather sharply

to the base of clypeus; clypeus broad, narrowing slightly toward the apex ;

cheek margin sinuate ; pronotum as long as vertex, slightly concave behind ;

elytral veins rather weak with one cross-vein on disc.

Female, last ventral segment longer than preceding ; hind border truncate,

the lateral angles slightly produced ; pygofer broad, scarcely exceeded by the

ovipositor.

Straw-coloured ; vertex with a pair of oblique dashes at the apex, a faint

dot near the base; pronotum with impressed fuscous markings anteriorly ;

elytra with whitish veins, the areoles in part faintly margined with fuscous ;

front with faint lateral arcs ; pygofer at base somewhat suffused with fuscous.

Length, 3 mm.

Described from one female (holotype) “Tau, Manua, 9-27-1923, O. H.

Swezey collector, grass.”

This specimen was mounted with the one described as affinis, evidently

collected at the same time but certainly a different species. It has the vertex

182 INSECTS OF SAMOA.’

shorter, much less angulate, and the picture would, I think, preclude its being

considered the female of any other species referred to this genus.

22. Jassus fijianus, n. sp.

Head narrower than pronotum ; eyes large ; vertex narrow, widening from

base to front, scarcely longer than width of the eyes and but little longer than

at base; front narrow, contracted at clypeus; clypeus broadly carinate, long,

twice as long as width, broadening toward the apex ; pronotum short, scarcely

longer than vertex, distinctly granulate, hind border concave; scutellum

sparsely granulate on the basal part, apical part elevated; elytra with the

veins not granulate, and faintly marginate with punctures.

Female, last ventral segment elongate, slightly produced and rounded on

the hind border. Colour dark brown ; front border of vertex, ocelli, basal part

of frons, a few transverse arcs and some other indistinct maculae on the elytra,

fuscous ; disc of vertex, pronotal and scutellar granules and marginal dots and

dashes on elytral veins, yellowish. Beneath yellowish, legs tinged with brown.

Length, 7 mm.

One specimen (holotype) Fiji, Lau, Kambara, 23.viii.1924, Bryan.

This species can hardly be considered a member of the Samoan fauna and

there does not appear to have been any member of this genus recorded from

Samoa.

Genus Jassoidula, new genus.

Nearly related to Jassus and having considerable resemblance to Neocoelidia

but with produced head, which is extended distinctly beyond the margin of the

eyes, large eyes and very large, conspicuous ocelli located half-way between eyes

and tip of vertex. These are fairly distinct differences from either Jassus or

Neocoelidia.

The male plates seem to offer a distinctive character as they are very much

expanded posteriorly and appear in position very much like the expanded

pygofer, but close examination indicates that the oedeagus is a slender structure

within these plates or outer claspers and the pygofer is a much reduced structure

hidden from below by the expanded plates. Valve apparently wanting or

covered. Elytra with five apical cells. Antennae quite long. Genotype:

Jassoidula straminea.

HEMIPTERA—CICADELLIDAE (JASSIDAE). 183

23. Jassoidula straminea, n. sp. (Text-fig. 13).

Head narrow ; vertex produced, subangulate, widening before eyes ; front

long, narrow, tapering to base of clypeus ; clypeus elongate, carinate ; pronotum

short, arched before, concave behind; elytra with Jassine venation, veins

distinct, apical cells four.

Female, last ventral segment long, hind border sinuate, produced medially ;

ovipositor extending one-fourth its length beyond tip of pygofer. Male valve

hidden. Plates narrow, elongate, tapering to acute tips.

Text-Fic. 13.—Jassoidula straminea, n. sp.: a, dorsal view ; 6, vertex, pronotum

and scutellum ; c¢, face; d, female, and e, male genitalia.

Colour, dull stramineous to ochreous, without evident markings except a

smoky band and spot on elytra beyond tip of clavus.

Length: female, 6 mm.; male, 5-25 mm.

¥rom Tutuila: seven female specimens (holotype and paratypes), 760—

900 {t.; five females, 900-1,200 ft. ; one male, 1,070 ft.; two males (allotype

and paratypes) Tutuila, Pago Pago, 30.ix.1923, Swezey and Wilder.

24, Jassoidula niuensis, n. sp.

Similar to straminea but with more distinct fuscous markings on the elytra

and two distinct fuscous points on the scutellum. Vertex with sides nearly

parallel, widened slightly in front of eyes, subangulate; ocelli large, con-

184. INSECTS OF SAMOA. ©

colorous with front; front narrow, tapering moderately to clypeus, broadly

carinate, widening distinctly toward the tip; lorae narrow; cheeks narrow ;

pronotum scarcely as long as vertex, minutely rugose; scutellum smooth,

deeply indented behind the middle ; elytral veins distinct.

Female, last ventral segment elongate, more or less compressed and carinate,

hind border produced, rounded. Male, plates long, very narrow at the base,

broadly expanded, foliaceous, apex subangulate, with an apical seta; with the

plates, outer surface densely minutely hairy; pygofer short and hind margin

ciliate.

Colour, dark stramineous. Vertex with a more or less obsolete reddish

stripe on either side; the pronotum slightly infuscate on the disc; scutellum

with two distinct fuscous dots; elytra with the apical part of clavus and two

bands across the apex fuscous, veins partly infuscate, the central part of the

clavus and a broad costal area subhyaline. Beneath paler.

Length: female, 5-5 mm.; male, 5 mm.

Eight females (holotype and paratypes), twelve males (allotype and para-

types), from Savage Island, Niue, 6.vii.1918, Kellers.

This species is evidently very close to straminea, but it is uniformly smaller

and the females are distinctly marked with a fuscous pattern that is different

from that of straminea, which, with the difference in location, seems to justify

a separate description.

25. Jassoidula ochracea, n. sp.

Vertex a little longer than width at base, rounded, scarcely subangulate in

front; ocelli very large, filling a large part of the space between eyes and tip

of vertex ; front slightly elevated above antennae, narrowing gradually to base

of clypeus; clypeus narrowed at base, expanded to tip; lorae rather broad,

distant from margin of cheeks ; margin of cheeks scarcely smuate ; pronotum

short, scarcely as long as vertex, hind border slightly concave ; scutellum rather

small.

Male, valve wanting ; genital plates long, much expanded apically, densely

hairy, the margins, especially on the outer part, deeply fuscous.

Colour: vertex black marked with yellowish at the base and with two

large yellowish spots including the ocelli; front brown, bordered with fuscous ;

clypeus yellowish with a fuscous carina; pronotum yellow; elytra ochreous,

HEMIPTERA—CICADELLIDAE (JASSIDAE). 185

apical border somewhat infuscate, beneath dull ochreous ; abdominal segments

infuscate with yellowish margins.

Length, 6 mm.

One specimen (holotype), male, Upolu, Malololelei, 2,000 ft., xu.1925,

Buxton and Hopkins.

26. Jassoidula infuscata, n. sp.

Similar to straminea but with a distinct infuscate pattern covering most of

the pronotum, scutellum and central part of the elytra.

Vertex longer at middle than width at base, distinctly produced in front of

eyes ; ocelli very large ; antennae reaching second coxae ; front narrow, sloping

to clypeus ; clypeus narrow at base, expanded and emarginate at tip; vertex

minutely granulate, at base finely shagreened, above minutely rugose and

carinate ; elytral veins prominent.

Male, plates rather thick at base, extremely flat and oeended apically,

with long dense hairs.

Basal part of vertex, pronotum, and base of scutellum and a broad middle

irregular stripe on the elytra, infuscate ; apex of vertex, tip of scutellum, a

ragged inner border and a broad costal margin, whitish or subhyaline ; under-

neath light brown ; the face somewhat yellowish.

Length, 5-5 mm.

One male (holotype) Tutuila, 900-1,200 ft., centre of island, 30.vi.1918,

Kellers.

27. Jassoidula metallica, n. sp.

Head slightly narrower than pronotum; vertex as wide as the eyes,

widening slightly toward the apex; margins elevated, rounded anteriorly ;

front narrow, slightly contracted below the eyes; clypeus half longer than

broad, sides nearly parallel, slightly expanded toward apex ; lorae small, distant

from cheek margin ; cheek margin scarcely sinuate ; pronotum rather minutely

granulate as well as vertex, sinuate behind ; scutellum slightly granulate on the

basal part ; elytra with strong veins, five apical cells.

Female, last ventral segment twice as long as preceding, produced and

distinctly carinate ; pygofer short, slightly exceeded by the ovipositor, which

extends nearly to tip of elytra. Colour dark fuscous, and pale fulvous and

186 INSECTS OF SAMOA. °

yellow. Vertex yellow with curved fuscous stripes at side; pronotum dark

fuscous ; lateral portions yellowish ; elytra metallic, bronzy, somewhat irides-

cent, purplish toward the tip ; base of clavus and patch on base of corium dark

greenish, apical cells smoky, costa with a hyaline patch at centre and in the

first apical cell; beneath the front fulvous, cheeks yellowish, legs yellowish

becoming fulvous before tips of femora and tibiae, abdomen black above, pale

brown below, base of last ventral segment and sides of pygofer fuscous.

Length, 5-5 mm.

One specimen: female (holotype) Nukualofa, Tonga, 16.11.1925, Hopkins.

This interesting little species seems to be somewhat intermediate between

Jassus and Jassoidula, but in most characters it seems to fit in with the latter

genus.

28. Jassoidula limbata, n. sp.

Head narrower than pronotum ; vertex moderately produced before the

eyes, widening gradually from the base, a little longer than width at base ;

antennae long, reaching hind coxae; front narrow, narrowed below the eyes,

tapering to clypeus; clypeus long, twice as long as width at base, weakly

carinate ; lorae narrow; cheeks scarcely sinuate below the eyes; pronotum

nearly as long as vertex, beneath carinate, hind border sinuate ; elytral margins

nearly parallel, apex rounded.

Female, last ventral segment half longer than preceding, distinctly pro-

duced, a broad median lobe with a small notch at the tip ; ovipositor half longer

than pygofer, reaching tip of elytra. Male, plates rather thick at base, much

expanded, flat on the apical part and densely covered with long whitish hairs.

Colour, light brown; the eyes, apex of vertex, sides of pronotum, most of

the elytra, fuscous; the disc of vertex, pronotum, and scutellum brown: sides

of vertex, the inner border of clavus and the costa and an apical margin of

elytra, pale brown or yellowish; beneath pale yellowish, the front fuscous at

base, lower part and clypeus fulvous; veins and tergum blackish; ovipositor,

margin of pygofer black, tip of ovipositor brown. In the male the abdomen

is dark beneath as well as above with the margins of the segments pale.

Length, 8 mm.

One female specimen (holotype) Malololelei, Upolu, 2,000 ft., vi.1924,

Buxton and Hopkins; one male (allotype) Savaii, Salailua, v.1923-1924,

Bryan.

HEMIPTERA—CICADELLIDAE (JASSIDAK). 187

The male allotype associated with this species agrees so perfectly in colour

pattern that I am confident it should be placed here in spite of the different

locality record.

29. Jassoidula cuprescens, n. sp.

Head distinctly narrower than pronotum, vertex broadening from the

base, elongate, fairly subangulate before; front narrow, somewhat sinuate

between antennae, tapering to narrow clypeus; clypeus narrow at base,

expanding to truncate tip with a distinct keel on basal half; lorae rather large,

with angular projections toward antennae; cheek margins slightly sinuate ;

pronotum short, two-thirds as long as vertex, distinctly granulate, hind border

scarcely sinuate; scutellum minutely granulate or shagreened; elytra with

distinct venation.

Male, plates rather thick at base, narrow, broadening to moderately expanded

apical portion, with rather short fine hairs.

Colour dark fuscous to blackish, the vertex with a median narrow pale line,

margins of eyes pale, hind border of pronotum narrowly brownish; apex of

scutellum bordered with light brown ; elytra metallic green at base, clavus and

most of corium hyaline, the patch on the costa and a round spot in the outer

apical cell pale yellowish. Beneath entirely blackish, except the sutures and

antennae; beak and legs yellow; abdomen black, the segments with pale

narrow margins, base of plates pale. Leneth, 5 mm.

One specimen, male (holotype), Neiafu, Vavau, Tonga, 5.11.1925, Hopkins.

This species has the male plates somewhat less expanded than in the typical

forms of the genus, but it agrees better here than with Jassus. The sinuated

frons seems to approach the character given in the genus Guliga Dist., but in

that genus the sinuation is very pronounced and apparently the frons widens

toward the apex, which is not the case in this species.

30. Jassoidula pallida, n. sp.

Head narrower than pronotum ; vertex longer at middle than at base and

base concave, apex rounded ; ocelli conspicuous, half-way between eyes and tip ;

antennae long, reaching hind coxae, front with sides nearly parallel to below

the middle, narrowing sharply to clypeus ; clypeus one-half longer than broad,

188 INSECTS OF SAMOA.

widening a little to the tip; lorae rather small, distant from cheek margin,

margin of cheek sinuate below the eye; pronotum about two-thirds as long as

vertex, weakly granulate ; hind border sinuate ; scutellum with a deep impres-

sion at the middle, smooth ; elytra with five apical cells, veins distinct.

Female, last ventral segment one-half longer than the preceding, hind

border sinuate, median portion slightly produced, a very shallow notch at

middle, ovipositor exceeding tip of pygofer, reaching nearly to tip of elytra.

Colour pale stramineous without markings, except a fulvous stripe at the

sides of the vertex. LHlytra hyaline. Length, 6 mm.

One specimen, female (holotype), Haapai, Tonga, 11-26-1925, Hopkins.

This species appears to fit into the genus although there is no male for

comparison with other species of the genus.

Genus Nesosteles Kirkaldy.

Nesosteles Kirkaldy, Bul. No. 1, Div. Ent. Exp. Sta. H.S.P.A., p. 343 (1906).

Very similar to Hugnathodus Baker, possessing a vestige of nervure forming

an areolet in basal part of costal areole, scarcely evident in the Nearctic forms.

Genotype: Nesosteles hebe Kirkaldy.

31. Nesosteles hebe Kirkaldy.

Nesosteles hebe Kirkaldy, Bul. No. 1, Pt. 9, Div. Ent. Exp. Sta. H.S.P.A., p. 343 (1906).

Head as wide as pronotum; vertex short, as long at middle as at eye.

Body slender, sides subparallel, abdomen black above.

Length: female, 4mm.; male, 3 mm.

Specimens from Savaii, Safune, v-4-1924, Lower Forest, Bryan, and Savaii,

Sailailua, v-16-1924, lowland to 1,000 ft., Bryan.

Originally described from Fiji. This species seems to be a common, often

very abundant, insect in many of the Polynesian islands.

32. Nesosteles tutuilana, n. sp. (Text-fig. 14).

Head wider than pronotum, distinctly arched or curved. Vertex short, as

long at middle as at eye. Front a little longer than broad, sides curving to

narrow apex; clypeus rather long, narrowing slightly to apex; lorae close to

HEMIPTERA—CICADELLIDAE (JASSIDAE). 189

margin of cheek; cheek margin faintly sinuate; pronotum nearly four times

as long as vertex, lateral margins short, hind border truncate ; elytra with costa

convex. Female, last ventral segment

as long as preceding, slightly indented

to form broad median lobe, the indenta-

tions accentuated by black projection.

Male valve large, triangular; plates

short, narrowed to broadly rounded tips,

scantily setose.

Pale gray marking. LElytra sub- °

hyaline; veins concolorous. Beneath

pale except the black marking on hind

border of female last ventral segment.

‘Length : female, 3 mm.; male, 3 mm.

Tutuila, Leone Rd., 2-19-1924,

Bryan ; Pago Pago, 9-4-1923, Swezey Text-ric. 14.—Nesosteles tutuilana, n. sp. :

and Wilder; and Savaii, Safune, v-11- GaadowsAl wicw>, Urtacesy oc, female, and

d, male genitalia.

1924, Lower forest, 1,000-2,000 ft., Bryan.

This species is smaller than hebe, more yellowish in tint, and lacks the

longitudinal stripes usually evident in that species, and the genitalia are

distinct.

Genus Homa Distant.

Fauna Brit. India, Rhynchota, IV, p. 400, 1908.

Type H. insignis, Dist. Distribution Ceylon.

“ Vertex subquadrate, longer than breadth between eyes at base, rounded

but very slightly produced in front of eyes, which are almost as long as the

vertex and do not project over the anterior angles of the pronotum ; face long,

broad, narrowing to clypeus which is short and broad ; pronotum shorter than

vertex, transverse, a little rounded in front, the lateral margins nearly straight ;

scutellum broad, subtriangular ; legs slender, posterior tibiae strongly spinulose

and slightly curved; tegmina long, much longer than abdomen, apical cells

three, large, and subequal in length.” (Distant.)

190 INSECTS OF SAMOA.

33. Homa upoluana, n. sp. (Text-fig. 15).

This species appears to be nearly related to Homa insignis Distant, but

aside from a different colour pattern the vertex is slightly subangulate and

apparently not quite so much longer than the pronotum. Vertex is longer than

pronotum, faintly subangulate, about one-

third longer at middle than next the eye;

front narrow tapering gradually to clypeus ;

clypeus short, apex rounded, lorae elongate,

reaching close to margin of cheek. Cheek

depressed below the eye; outer margin

shghtly sinuate ; pronotum broadly rounded

in front, scarcely concave behind; elytra with

three large apical cells, the outer one sessile

on transverse veins, wing with submarginal

vein and one apical cell.

Female, last ventral segment short, trun-

cate ; ovipositor extending beyond the pygofer

Text-ric. 15.—Homa ra n.sp.: about one-fourth its length. Male, valve

a, dorsal view ; 6, face; ¢, female, hidden ; plates elongate, triangular, narrowed

and d, male genitalia. . .

to acute points extending beyond the pygofer.

Colour ivory white, face and lateral margins of pronotum in some specimens

and the clavus and corium of elytra and tergum brilliant red. Beneath white

or lvory white, abdomen tinged with yellow.

Length: female, 2 mm.; male, 1-75 mm.

Described from sixteen specimens (holotype, allotype, and paratypes) :

fourteen from Apia, 15.ix.1923, Swezey and Wilder; and two specimens (para-

types), Savaii, Safune, 2.v.1924, Rain Forest, 2,000-4,000 ft., Bryan.

This species evidently belongs to Distant’s genus Homa and his division of

Empoascaria, the wing having a definite single cell. He was uncertain of its

position and says, “I have included Homa in this division, but have very

imperfectly seen the wing venation in the unique typical specimen.” This

record evidently gives a considerable additional range to the genus.

HEMIPTERA—CICADELLIDAE (JASSIDAE). 191

Genus Erythroneura Fitch.

Erythroneura Fitch, Trans. N.Y. State Cab., p. 62 (1851).

34. Erythroneura samoana, nN. sp.

Head narrower than pronotum ; vertex obtusely angulate, three-fourths as

long as width at base ; front margins nearly parallel to near the apex, narrowing

abruptly to clypeus ; cheek margins slightly sinuate ; pronotum a little longer

than the vertex, the hind margins slightly concave.

Female, last ventral segment twice as long as preceding, rounded behind,

the strongly curved ovipositor extending distinctly beyond the tip of pygofer.

Male, last ventral segment about as long as preceding; somewhat tubular,

valve wanting or hidden; plates or outer claspers narrow and strongly curved

and faintly ciliate.

Colour ivory white tinged with vellowish and in some specimens distinctly

tinged with fulvous or orange on the pronotum ; the elytra hyaline suffused with

yellow to nearly the cross-veins, an elongate ivory plaque on the costa; apex

hyaline and wings milky hyaline. Beneath pale, tips of tarsi black.

Length: female, 3 mm.; male, 2-5 mm.

Described from a series of specimens: three female (holotype and para-

types), Savaii, Salailua, 23-v-1924, Bryan; four male (allotype and paratypes),

Savaii, Safune, 13-v-1924, Bryan; and also specimens referred to this species

from Tutuila, Fagasa, 9-9-1923, Swezey and Wilder.

192 INSECTS OF SAMOA. |

PRINCIPAL REFERENCE WORKS.

Distant, W. L. Fauna of British India, Rhyncota, IV, pp. 157-418, 1907-1908.

Kirkaupy, G. W. Div. Ent. Bulletin No.1, pt. 9, Hawarian Sugar Planters’ Association, pp. 271-

479, 1906, and No. 3, pp. 1-186, 1907.

Meuicuar, B. Homop. Faun. Ceylon, 1903.

Stenoret, V. Essai sur les Jassides, Ann. Entom. Soc. de France, Tome 1853.

Sau, 0. Ofv. Vet.-Akad. Férh., p. 737, 1870.

Waker, F. List of Homoptera, British Museum, 1851-5 and Supplement, 1858.

LIST OF TEXT-FIGURES.

Fig. 1. Agalliopsis fasciatus, n. sp.: a, dorsal view ; b. face ; c, female genitalia.

Fig. 2. Bythoscopus angustatus, n. sp.: a, dorsal view ; b, face; c, female genitalia.

Fig. 3. Bythoscopus tutuilanus, n. sp.: a, dorsal view ; 6, face ; c, female, and d, male genitalia.

Fig. 4. Bythoscopus laticeps, n. sp.: a, dorsal view ; 6, face ; c, female genitalia.

Fig. 5. Bythoscopus hyalinus, un. sp.: a, dorsal view ; b, face; c, female genitalia.

Fig. 6. Idiocerus tutuilanus, n. sp.: a, dorsal view ; 6, face; c, male genitalia.

Fig. 7. Xestocephalus tutuilanus, n. sp.: a, dorsal view ; 6, face ; c, female, and d, male genitalia.

Fig. 8. Hecaloidella nitida, n. sp.: a, dorsal view, head, pronotum and scutellum ; b, female, and

¢, male genitalia ; d, elytron.

Fig. 9. Nephotettix plebeius Kirkaldy : a, dorsal view ; 6, face; c, female, and d, male genitalia.

Fig. 10. Drabescus samoanus, n. sp.: a, dorsal view ; b, face; c, female, and d, male genitalia.

Fig. 11. Hutettix ornatella, nu. sp.: a, dorsal view ; 6, face; c, female, and d, male genitalia.

Fig. 12. Storellus pacificus, n. sp.: a, dorsal view ; b, face; c, female, and d, male genitalia.

Fig. 13. Jassoidula straminea, n. sp.: a, dorsal view ; 6, vertex, pronotum and scutellum ; c¢, face ;

d, female, and e, male genitalia.

Fig. 14. Nesosteles tutuilana, n. sp.: a, dorsal view ; b, face; c, female, and d, male genitalia.

Fig. 15. Homa upoluana, n. sp.: a, dorsal view ; 6, face ; c, female, and d, male genitalia.

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PRINTED IN GREAT BRITAIN BY WILLIAM CLOWES AND SONS, LIMITED, LONDON AND BECCLES.

INSECTS OF SAMOA

AND OTHER SAMOAN TERRESTRIAL —

ARTHROPODA

List OF PARTS AND SYSTEM OF PUBLICATION :—

Part). 6 21.

ce ols

mudora As

Be ges NI

eee VT,

Reon WL

oar VILL.

| ae XG

The work

Orthoptera and Dermaptera (Complete.)

Hemiptera. : a

Lepidoptera.

Coleoptera. (Complete.)

Hymenoptera. (Complete.)

Diptera. i

Other Orders of Insects. (Complete.) _ |

Terrestrial Arthropoda other than Insects. (Complete.)

Summary and Index. |

is published at intervals in the form of numbered fascicles.

be | Although’ individual fascicles may contain contributions by more than one

author, each fascicle is so arranged as to form an integral portion of one or

other of the Parts specified above. |

Bier oe eeeicles aesncd a) 2th Lae 1934 - ates

" Insects of Somoa and other Samoan Terrestrial Aahrapoda Maps 1 and Date heey

* >) 2iGm envelope). 1927,'4to. “6d. | oi ye Oe February, 1927.

ye rf: ‘ORTHOPTERA AND DERMAPTERA. Be Sat Gates

_ Fase. 1. Dermaptera. By Dr. Alfredo Borelli. Pp. 1-8. 1928, 4to. Is. 28th Jal 1928.

Fase. 2 Orthoptera. By Dr. L. Chopard. 51 text- figures. Pp. 9-58. 1929, Ato. 5s, 26th January, 1929.

Delay I]. HeEmiprera.

ror.

Fasc. 1. Fulgoroidea. By F. Muir. 25 pete’ Psyllide cl panama) By

male Crawford. 4 text-figures. Coccide, Aphididae and Aleyrodide. Fees ers Sa

By F. Laing, M.A., BSc. 3 text-figures. Pp. 1-45. 1927, 4to. 2s.6d.. 2th June, 1927.

Fase. Ds -Cercopide. By V. Lallemand, M.D. 10 text-figures. Cicadide. By

ee Myers, Sc.D. 22 text-figures. Aquatic and Semi-aquatic Heteroptera. real des IDM a

y Prof. Teiso Esaki. 6 text-figures. Pp.47-80. 1928, 4to. 2s. 6d. 23rd June, 1928.

Fa ‘asc. 3. Heteroptera. By W. E. China, B.A. ‘(Cantab)). 28 text Aoures, (

Pp. 81-162.

opkins,

ee Ato.

19300 ee) eu 26th July, 1930.

‘ : Fase. 4. Cieadellid (Jassid By Herbert Osborn, Ohio State DS by aie

“Columbus, Ohi: *tetehewre Bp. 163- 194. 1934, 4to. 2s. 2th January, 1934,

oar Hi. Lainie

ace ABunerthes of Sat ae ge By GoHubal

MA. FE. pn and ame nl reese 0 “Br dio. 5s, 9th April, 1927.

Fasc. 2. ee ee By Edward Meyrick, BA., FR S. Pp. 65-116.

- 28th May, 1927.

Fase acu de, By” Lege B! Pronts beEs.) 2 tentfire and 1 plate.

Pp. T7168

1928, hae 2s. 6d. ce ey 24th March, 1928.

List of Fascelés issued to 27th January, 1934 (continued) —

Part. W. COLEOPTERA.

Fasc. 1. Carabide. By H. E. Andrewes. 9 text-figures. Be) By. a

yA ey text-figures. nae linide. By M. Cameron, MLB.) 2 text. ia

Lamellicornia. By G. J. Arrow. 13 text-figures. Pp. 1-66. 1927, 4to. 3s.

Fasc. 2. _Heteromera, Bostrychoidea, Malacodermata ae Buprestide. By K. ie

Blair, BSc. 14. text-figures. Elateride. By R. H. van Zwaluwenberg. 10 —

text-fgures. Melaside Eucnemide). By E. Fleutiaux. Cerambycide. By

hr. Aurivilltus. 1 plate. Brenthide. By R. eme. 4 text-figures.

Anthribide. By Karl Jordan: Ph.D. 11 text-figures. Proterhinide. By

R.C.L. Perkins, D.'Sc., F.R.S. Pp. 67-174. 1928, 4to. 5s

Fase. 3. Throscide. By KUCE BB Sect MO es —

By S. Maulik, M.A. 18 text-figures. Pp. 175-215. 1929, 4to

tae ey 217-248. 1929, Ato. 2s. 6d.

Parr v. toe)

_ Fasc. 1. Apoidea, Sphecoidea, and Vespoidea. By R. oe a ee DSc. fe :

FE RS,, and L. Evelyn Cheesman, F.E.S., F.Z.S. 12 text-figures. Larride.

By Francis X. Williams. 12 text-figures. Formicide. By Dr. 1B, cy i 25th Bein y ary, 1928. a

Pier hautest Pp. 1-58. 1928, tio. bse

pint VL Diptera.

19th December, 197.

DBrd F ebruary, 1929

Fasc. 4. Platypodide: and Scolytide. By O HG; oe DSc. 2B text- “Dad Jum 1909,

Date Masued. L ee oe

25th F cbruary, | 928.

Fasc. |, Streblide and Nycteribiide, By L. Fee, 7 Coe SP se

boscide. By G. F. Ferris. 6 text-figures. Pp. 1-21. 1927, 4to. 2s. 6d.

| oe 2. Nematocera. By F..W. Edwards, M.A. 20 text-figures. eae

. F. Barnes, B.A., Ph.D. 4 text-figures. Pp. 23-108. 1928, 4to

Fasc. 3. Stratiomyude, eG) and Ae By Gertrude Bea 6 text-

fisures. Larve o tratiomyiide. By P. A. Buxton, M.A. 2 ae

Dol ichopodide. By C. G. Lamb, Sc.D. 8 text-figures. Sacorpaeey

uxton, ae 3 text-figures. Muscidae. By R Malloch. } ‘

Pp. 109-175. 1929, 4to. 5s.

Fase. 4. echoes and Pipuncutidae! By ie Calin. 7 Pau eel

yrpnidae. By eae M. Hull. 2 text-figures. ee (Heteroneuridae

4to.

Fal 5 One Bee Roliod 46 text figures. Calliphoridae. as

j.R. Malloch. Pp. 215-237. 1930, 4to. 2s.

Part VII. Otrner Orpers oF Inseers.

Fasc. |. Isoptera: Family Termitide. By Gerald F. Hil 14 asa sidan

1 plate. Odonata. By Lt.-Col. F. C. Fraser, ILM'S., FES. 5 text-figures. 81h vee 1927.

Pp. 1-44. 1927, 4to. 2s. 6d.

Fasc. 2. Bei By ee J Tillyar,, Sep: chu E RS., and J. A.

Lestage. 2 text-figures. Si aptera. By P. A. Buxton, M.A. Thysanoptera.

By Hees Bagnall, F Rel ae ee

Buxton, M.A. Trichoptera. By Martin E. Mosely. | figure.

Neuroptera. By P. Esben-Petersen. 1 text-figure and 2 plates. Apterygota

By George H. Carpenter, D.Sc. 32 text-figures. Pp. 77-\16. 1928, Ato. 2. 6d.

Part VIII. TrrrestriaL ARTHROPODA OTHER THAN Insecrs..

Fasc. |. Isopoda Terrestria. By. Harold G. Jackéouy DSc. 2 plates. Score ‘

pionoidea. By P. A. Buxton, M.A. Pseudo-scorpiones. By A. Kastner. 11

4to. 2s. 6d. 7

Fase. 2. Myriopoden (Myriopoda). By C. Att 4 ecient Waenkes

(Araridida). By DrinieenBeviaad to tree Pp. 29-78, 1929, 4to. 25. 6d.

text-hgures. Acarina. By Stanley Hirst. 2 text-figures. oe 1-27. 1927,

and oe By. R- Malloch. 6 ‘text-figures. Pp. ATF2N3. 1929, |

‘ Pp 76) Fore es

Tangs cae figures. Pp. Me O34 Tune: 192

Fi 3. Mallophaga. By J. Waterston, D.Sc. 2 text-figures. Anoplura. By) eel 3 Peay

23rd July, 1927.

22nd June, | 929. >) os