AMERICAN NATURALIST,
An Hllusteate Hlagazine
OF
NATURAL HISTORY.
EDITED BY
EDWARD D. COPE anD J. S. KINGSLEY,
ASSISTED BY
' Dr. C. O. WuiTMan, Dr. C. E. Bessey, THOMAS WILSON, ProF. C. M.
WEED, ProF. W. S. BAYLEY, Pror. E. A. ANDREWS.
VOLUME XXVII. jen -Ju
Mo. Bot. Garden,
1895.
PHILADELPHIA, U. S$. A.
THE EDWARDS & DOCKER Co.,
518 anp 520 MINOR STREET.
1893.
CONTENTS.
LEADING ARTICLES:
Gases in Living Plants, J. C. ARTHU
1
Certain Shell Heaps of the St. John’s River, Florida (Illustrated), CLARENCE
B. Moore 8, 113, 605,
14
Legends of the Sumiro Accadians of Chaldea, ALICE BODINGTON
The Flight of Birds (Illustrated). J. LANCASTER
Joint Formations Among the Invertebrata (Illustrated). BENJAMIN SHARP...
The Ancyclopoda, Chalicotherium and Artionyx (Illustrated) H. F. OSBORN..
The Quantity of Human Life, J. Lawron WILLIAMS
The Titanotherium Beds (Illustrated).
An _—_ Produced oe without Charictatiasice of the Mother (Illus-
é H. Bo
‘On the ee of p POES RSE R. W. SHUFELDT.
The Genealogy of Man (Illustrated). E. D. COPE
The Probable Physiognomy of the Cretaceous Plant Population, C. MACMIL-
LAN...
An Extreme Case.of Parasitism, ROBERT HESSLER
'On the TUNE and Distribution of the North Aaron Unionidae, with
s on the West Coast aries C. T. Sm
Recent uaa of. Catnivorous Plants; J.-G. Saabr its. ois cists wsiesseiesn’: dhevens
A New Theory of the Mechanical rt x the Metapodial Keels of Di-
plarthra (Illustrated). J. L. WORTMA
Among the Cliff-Dwellers (Illustrated). C. c WEBSTER
New Discoveries of Fossil Mammalia of Southern Patagonia, F. AMEGHINO..
Recent Researches upon the Succession of the Teeth in Mammals, H. F. Os-
BO:
Symbiosis and Mutualism in Lichens, ROSCOE POUND.....-.sssseee cesses sesseseee
Evolution and Dichromatism in the Genus Megascops, E. M. HASBROUCK ...521,
The Cinnamon Harvest-Spider and its Variations (Illustrated). C. M. WEED.
Some —— in etic and Phylogeny in Brachiopoda (Illustrated),
C. E.B
ssssesse
Notes on nog Tarea of Europe ue BasHFoRD DEAN. 625,
The Spore-Forming Species of the Genus Saccharomyces, J. C: BAY...........
The Philosophy of Flower Seasons, H. L. CLARKE
The Morphology of Root ebeiles of Leguminosae (Illustrated). ALBERT
On oe — of the Carapace in the Devonian Crustacean Rhinocaris ; and
e Relation of the Genus to Mesothyra and the Phyllocardia (Ilustra
p J M. CLARKE
708
10
118
599
697
685
General Physiology and its Relation to Morphology. C. O. WHITMAN........
rece in its General AEN H. L. RUssELI
847,
f Lateroversion of the Ophidian Heart (Illustrated). P. A. FISH.
Woes as an Acquired ak erpai ALPHEUS HYATT......
a Eggs of Pityophis melanoleucus (Illustrated). J. P. MOORE
he Genera of the Dipnoi Dipneumones, HOWARD AYRES
Wied Intelligence, JAS. WEI
Notes on a Collection of A from North Western Louisiana, and Harri-
son County, Texas, L. W. VAUGHN
Trenton and ceg mme Gravel Specimens Compared with the SA "9
Refuse in America and Europe rapsi C.M
Notes on E ppap insect, TD; AG
The Color Variations of the Milk Snake (Illustrated). E. D. Cope.............
EDITORIALS.— The Antivivisectionists in Pennsylvania,-26; The Coming
Peary Expedition, 26; Dates of Issue of the NATURALIST for 1892,
27; The Law of Priority in Nomenclature, 184; The Meeting of the
Societies at Princeton, 135; The American Table at Naples, 136; ae
Geological Survey of Inoi, 238; Reform in Orthography, 238;
tality of the English Sparrow, 239; The American aa So.
rnal o
ogy, 360; A Preserve for New Zealand, 360; The Geological Survey
of Georgia, 450; The Exhibition of Monstrosities, 450; Game Protec-
tion in New York, 451; The ee of the American Philo-
sophical Society, 542; A Museum for Chicago, 886; Duplication of
Names for Postoffices, 886 ; Biography of American Botany, 887;
he Zoologists and the Am n Association, maipi Specimens in the
Mails, 979; The Glacialists? Macs ne, 980; The Bureau of Ethnol-
gy ; The Pronunciation of Arkansas, ie, College and Univer-
sity Periodicals, 1072; Appropriation for the Gypsy Moth Commission,
1072; Dates of Issues of the NATURALIST for 1893, 1073.
RECENT BOOKS AND PAMPHLETS.—28, 137, 240, 361, 452, 544, 650, 724, 808,
889, 982,
iv The American Naturalist. ; [Vol. XXVII,
802
1050
RECENT LITERATURE.—The Apodi idae, 31; Darwin and After Darwin, 31;
The Diseases of Personality, 33; Two Text-Books of Human Embry-
ology, 140 ; Geological Survey of Texas, 1891, 142; Mineral Resour-
ces of the United States, 1889 and 1890, 143 ; Memojrs of the PRS
Academy of Sciences, Vol. V, 243; Brooks and Bruce on the E bry-
ology of the Macrura, 243; Campbell’s Biology, "e laia
Papers of the U. S. Geol. Surv., Neocene, 246; Cary on the Evolution
of the Minnesota Valley, 365; Ridgwa way, on the Anatomy of the Hum-
ming-Birds and Swifts—A Rejoinder, 367; Beddard’s Animal Colora-
Vols. I and II, 455; The Earth’s Hi story, 456; Wright’s Light, 496;
Evolution of the Colors of North American Land Birds, 547 ; Wright’s
Man and the Glacial Period, 550 ; Some Recent Books on B Bacteriology,
554; Report on the Fish and Fisheries of the United States for 1888,
1893] s Contents. v
556; A Popular Botany, 653; Two Text-Books of Physiology, 654;
Caldenwood on Mental Evolution,654; Hawks and Owls of the United
States, 727; Fresh Water Algae and the Desmidieae of the United
States, 727; Gasteropoda and Cephalopoda of the New Jersey Creta-
ceous _— 7285 smears # oe Veuetants vers ali; Report of the
Year,1891,811;
The ae Paleolgy aa the Liane itatacado, 811; Weider-
sheim’s Comparative Anatomy, 892; Mill’s Diatomaceae, 892; Ken-
nel’s Zoology, ae Iowa Geological Survey, First Annual Report for
2,985; Correlation Papers—The Newark System, 987; Spalding’s
Guide to the Study of Common Plants, 988; Annual Report of the
United States Geological Survey, 1889-90, 990; The Report of the
Death Valley Expedition, 990 ; Piersol’s Histology, 1077; First Annual
ol for sping of the Iowa ie Rept niki Survey, 1078; The Pale-
ozoic p of afi 1078; The Mesozoic Echinodermata of the
de seein. 1079; The Flora a the Dakota Group, 1079; Fritsch’s
Fauna of the faiths of Bohemia 1079
GENERAL NOTES.— Geography and Travels.—Africa, 253, 458, 730 ; America,
255; Asia, 257, 461; Europe, 261, 730; Polar Regions, 268, 468;
ee 260, 466 ; The Iowa State sees ere to the West
dies and Florida Keys, set General Not 464
cae sald Paleontology.—On e T SPAT y ie 34; Geology of
Northwestern Alabama, nates ; The Mesosauria of South Africa, 35; Kan
sas Pterodactyls, 36; dies Formations of Southern ennen
44; The Genus Hybodus, 145 ; On some Dicynodont and Other Rep-
tilian Remains from the Elgin Sandstone, peed Cenozoic Insects, 146:
Uplifts in the Sierras of Califo ornia, 147; w Pliocene Ruminant,
147; A Remarkable Artiodactyle from a el River Epoch, 147 ;
The White Clays of the Ohio Region, 148 ; Geology of Eastern Siberia,
257 ; Geological Features of Arabia Petraea and Palestine, 267; Ver-
tebrate Fauna of the Ordovician of Colorado, 268; The Loess in South-
ern Russia, 269; Sources of the Texas Drift, 269; Currents of the
North Atlantic, 375; Fins of Paleaspis americana, 375; New Re eptiles
from the Elgin Sandstone, 376; Fossil Reptiles from the Parana, visas
On the Systematic Position of the Genus Protogonodon, ash, Bro
Coal and Lignite of Texas; Extinct Fauna of Mauritius, 379; The
‘Western oman of Ecuador, 469; Devonian Fossils fm Manitoba,
469; Jura and Trias in Taylorville, California, 470; The Post-Lara-
mie Beds of Middle Park, Colorado, 471; Marine Pliocene Beds of the
aea 471; The Oneonta and Chemung Formations in Eastern
Cen New York, 558; Tertiary Insects from Colorado and Utah,
e A Supposed New Order of Gigantic Fossils from Nebraska, 559 ;
Mammalia from the “ Pits of Gargas,” 560; The Norian nag of Can-
ada, 656; The Caudal Fin of Ichthyosaurian Reptiles
sil Fishes from the Upper Lias, 657 ; Affinities of sins Se 657;
Cretaceous Formations of Mexico, 657; On a New Musteline from the
John Day Miocene, 658 ; The Mammals of the Deep River Beds, 659;
- Conditions of Eraio Beneath Deep Glaciers, 662 ; The Moon’s Face,
The American Naturalist. [Vol. XXVII,
734; North America During Cambrian Time, 734; Lower Silurian
Brachiopoda of Minnesota, 735; Recent Volcanic Eruptions in Cali-
fornia, 818; Continental Problems, 816; Mineral Resources of the
United States for 1891, 816; Notes on Upper Devonian Fishes from
Canada, 817; The Diatomaceae of the Triassic (?) Sandstone of New
Jersey, 817; Do Glaciers Excavate? 818; Plistocene Deposits of Rus-
sia, 819; A New Plistocene Sabre-Tooth, 896; The Laurentian of the
Ottawa District, 996 ; Relations of the Laurentian and Huronian Rocks
North of Lake Huron, 996; The Carboniferous Glaciers of Central
France, 997 ; Quicksilver Ore Deposits, 998; The Discovery of Mio-
cene Amphisbaenians, 988; On Symmorium and the Position of the
Cladodont Sharks, 999; Mud Avalanches in the Mustagh Mountains,
82; Cladodont Sharks of the Cleveland Shale, 1083; The Neocene
Sierra aoe 1084; Geological News, 37, 149, 270, 471, 560, 662,
Minera S Petrography.—The Origin and ae of Igneus Rocks,
; The Novaculites of Arkansas, 42; ew Occurrence of Prtiolite,
43; Mineral Syntheses, 45, 277, 567; Lande Blowpipe Analysis
46; The Rocks of the Thalhorn, 272; The New Jersey Eleolite- ag
nite, 272 ; Mica a from kai ms; Rhyolites in Maryland
and Penn., 273; The Nepheline and Lencite Rocks of Brazil, 274;
Cecialinerschic aoe of Diopsides, 275; Hore from Hebron,
Maine, 276; Methods and Instruments, 278; An Appendix to ‘* Gems
of North America,” 278; Description of the New Rock Type, Mal-
chite, 880; Petrography of Hokkaido, Japan, 880; Two Peculiar
Rocks from Siberia, 381; An Ottrelite Bearing Conglomerate in Ver-
mont, 382; Lithophysae in the Rocche-Rosse, 382; The Composition
of Dune Sands of the Netherlands, 882; Quartz-Gabbro in Mary-
d, 383; Minerals from the Diamond Fields of Brazil, 383 ; ta
Resins 385; Isomorphism, 386; Etched Figures 6;
chemical Reaction. 387; Miscellaneous, 387 ; The Besisgrapky at ad
anada, 564; The Melibocus “Massiv” and its Dyke Rocks,
; The Granites of Argentina, S. A., 565; New Edition of Rosen-
ater Volume of Minerals, 567 ; eae Teal 567, 1093; Rock Sep-
arations, 568; Anorthorites and Diabases from the Minnesota Shore of
Lake Superior, 898; Volcanic Rocks of the Andes, 898; Basalts and
Trachytes from Gough’s Island, 899; Origin of the Gneisses of Heidel-
rg, 899; American Minerals, 901; Trachytes and Andesites of the
Siebengebirge, 1003; Variolite Dyke in Ireland, 1003; Chemical Na-
ture of Eruptive Rocks, 1003; Norites in the Eastern United States,
1004; Ottrelite Conglomerate in Vermont, 1004; Chalcedony and
other Silicious Spherulites, 1005; Danalite from Redruth, Cornwall
1005; Mirabilite Changed to Theradite, 1006 ; The Schists of South-
ern Berkshire, Massachusetts, 1087; The Phonolytes of the Hegan,
1087 ; The Ricks of a-New Island off Pantelleria, 1088; Analy:es
of Aalan Minerals, 1090; North American Minerals, 1091; Phy-
Contents.
cal Properties of Minerals, 1092; Petrographical News, 42, 274,
yan 1008 ; Mineralogical News, 43, 276, 384, 903, 1006; New Min-
erals, 45, 566, 900; Miscellaneous, 387, 903
Botany. —A New Edition Wolle’s Desmids, 47 ; Botanical Definitions, og
ngler
Timely Words o the Nomenclature Question, 48;
Prantl’s Waiuitichea Pa anil. 49; An International Sisk
Congress, 279; N orth American shai Century, , 474; New
York Fungi, 474; Seymour and Earle’s Economic Fungi, 475; H
sted’s Weed. Seeds, 475 ; Morong’s earki e he Canadian Mosses
and Lichens, 476; Caeoma nitens, 569; Our Naids, 569; Hough’s
American Woods, 570; Allen’s Characeae of Pai 570; The
Plants of the Bahamas, Jamaica, and Grand Cayman, 664; “The e Sapro-
legniaceae of the United States, 665; The Coming Botanical Meetings
at Madison, 738 ; ae Algae, 739; Check-List of New North
American ta, 821; Shall we Decapitalize Specific Names? 821;
The Use of Personal imi in Designating Species, 822; Botany at
the Madison Meetings, 823; Kuntze’s Revisio Generum Plantarum,
0 ws
n
Zoology.—Locomotion of Limpets, 51; Tunicate Studies, 51; Skeleton and
Teeth of the Australian Dugong, 51; On the Cephalohumeral Muscle
and the So-Called Clavicle of mers 52; A New Synaptomys from
New Jersey, 53; A New Evotomys from Southern New Jersey, 54;
The Cercaria Stage of Aiia, 151; Fecinidioi of the Eggs of
phiuma, 154; On a New Spade-Foot from Texas, 155; The Pedal
Skeleton ofthe Dorking Fowl,156; Allen’s Faunal hi of North Amer-
ica, 282; The Madagascar Fauna, 282; The Nephridia of Amphioxus,
283; The Position of the Marsipobranchs, 284; Degeneration of the
Clitoris, 284; Sed German Zoological Society, 388; Eyes of sont
tes, 389; Arachnida, 389; The Fishes of the Pacific Coast of Am
North of Cerros ae 390 ; On the Mechanical Genesis of the Sais
of Fishes, 391; Systematic Position of the Kiwi, 392; Mammals from
the Galapagos Islands, 394; The New England Species of Balanoglos-
sus, 477 ; Marsipobranchs, 477; Lateral Line of Siluroids, 477; Pro-
dromus of a New System of the Non-venomous Snakes, 477; A Medusa
from Lake Tanganyika, 571 ; The Air-Bladder and Weberian Ossicles
in the Siluroid Fishes, 571; Age Modifications of the Mucous Lining
of the Stomach of Ruminants, 572; A Deformity Inherited, 667; Pre-
liminary Note on the ae of the Species Usually United under
the Generic Name Sebastodes, 668 ; Batrachians of British India, 671;
Washington and British soar Ornithology, 671; Notes on the
Classification of the Cryptodira, 672; Two New Species of North
American Testudinata, 675; Zoology of the Lower Saskatchewan River,
741; Classification of the Actiniae, 829; Maioid Crabs in the National
Museum, 830; A New Lancelet 830; Descriptions of Four New Ro-
dents from California, 831; How Young Flickers are Fed, 1014; For-
syth Major and Rése on the Theory of Dental Evolution, 1014; Effects
Vil
1008
viii The American Naturalist. [Vol. XXVII,
of Temperature on the aape of Lepidoptera, 1016; Fish Acclimati-
zation on the Pacific Coast, 1017; The Molluscs of the Water Mains of
Embryolo — trulation of Aurelia, 57; Cleavage n Aeguoria Jorskila:
58; Experimental Embryology, 158 ; Studies in n Insect Embryology,
i erm- Epi
ade pee 397; Experiments mi “Cleav e, 388; Germ-Layers of
Vertebrates, 485 ; The Mantle of Ascidians 486; The Sea-Urchin Egg,
743 ; A Contribution to Insect Embryology, 745 ; Frog Eggs Under
Pressure, 1097; Em PTR i Chiton, 1098; Lithium Monsters,
1099; Mechanics of Embr
Entomology. —The Pupa of Areyramoti oedipus, 60; Horn-F ly in Cana ie
and Texas, 63; The Wheat Frit-Fly, 64; On a Small Collection of
c nent from the High pink B of British Columbia, 164 ;
Peculiar Seed Like Case-Worm from t an n, 166, 402; The
Pear-Tree ; Notes on Ohi other Phalangiidae, 294;
Gall-Producing Insects, 296 ; Recent Publications, 29 a ; Termi-
tophilous Insects, 400; Notes on the Mou s and Thorax of In-
sects and Chilopods ; The Woolly Alder Aphis, 402; The upar
i (0) pùaripeza, 402; No eric ormetida 74;
om 680 ;
S
e
5
& a
[e]
as
=
an
*&
o
CO
pi
T; $ i
n Atriplex, 1104; Honey Adulterations, 1105; North Ameri-
can italien: 1106; Entomological News, 64, 169, 578
AES Fa Solution of the Dust oe m in Microscopy ee ed),
ing P ne, 407; A Meth he fey sori the Blood-Vessels
i n Ova
Psychology.—Notes on Habits of European may e; A Nest-Building F ie,
67 ; Horse ‘‘Human Nature,” 68; Vision in a Young Girl Six Years
of Age, Operated upon for Double Congenital Cataract, 171; The
i e Snakeman
Archeology and Ethnology.—Lege ndary Evolution of the Navajo Indians, 69
ea and Population of European Co untries, 173; A Measure of Civil-
ization, 174; International Congress of Americanists, 300, 579, 755
838; Language Versus Ana’ atomy in Determining Human Races, 581 :
The cents of New: Zew Zealand, 582; The Exposicion Historico.
. Americano, 838, 907; The Explorations in the Delaware Valley, 1023;
The Tatera sl Congress of ee Archeology and Anthro-
at prer
RECORD oF AMERICA j van beiesi
PROCEEDINGS OF Scimvrivic SOCIETIES, 80; 176, 306, 409, 488, 585, 682,
762, 918, 1
ScIENTIFIC News, a 191, 410, 489, 592, 683, 764, 844, 917, 1088... ..000000
1096
1100
298
Ne
1893.]
Index. ix
INDEX.
CANTHODES semistriatus.... 149
Accipiter nisus 65
oe A poe rast 36 166
red Character 191
Acquired Pohibdetertetic Dic uevieesvebeed 865
Acr eu perinus 662
Actiniae, “Clastcaton of PRE EE ST 829
Acuto ME A Eni 669
Adelonycters Pho vise heverenes UO
Aisculus hi stanum Lesecssssen 589
Aine of Ichthyormisi.s. sos 657
African Travel, notes of........s.00 254
Apa Scientific Society of Oregon 683
Agathaumas, plaster cast of........ - 593
Agelacrinitidaë... siseses. sesse -soem 87
Age Fy ator! of the mucous
ing of the Stomach of
gues I ts 572
Agrimonia mollis 1112
nia striata 1112
a falcata.......+. se reer 153
ASi e ERN 1
riage ay and Weberian Ossicles
n the Siluroid Fishes........... 7
P ee a al Survey. 192
Algae (Fr h Water) 73
Algae abe Deamids me Water)
of the U..S., Stokes’..2..6<.2...
pheus saulcyi 243
Amara remotestriata De iiias
Ameghino, F,, New Discoveries
of re en in South-
ern 439
Soka F AERE ERPE Society... 289
American g pigain Society, Fifth
_ Annu tin 17,
American Aes ra iE ee 901
American Morphological Society. 307
American Psychological Associa
GO a e ciseeeee 186
ican Society of Naturalists... . 306
erican Woods, Houghs.......... 570
Amnicola lutetiana . 1094
Amone God Sera ame Cok.
4
Am Peis lips leni Harr...... 289
Amphibolips ilicifoliaé Bass......... é
Amphibolips i me Ses ics, 289
Amphisbaenians, Miocéne.......... 998
Amphistomum subclavatum. ........ 151
rr ala POCORN oss Bs cccese 296
Anal 007
Analysis of American Minerals.... 1090
Aoii OF PIE Sa e cssss nah 008
Anchitherium equinum 661
ee .&B 670
Ancyk 118
Ancylopoda Chalicotherium and
Artionyx, H. F. Osborn........ 118
Andricus sep si Rw: 289
pe bie M Seana iia ore 1018
al Colo loration, Beddard’s......
Animal Intelligence, Jas. Weis Jr. 933
736
a S A dae Wes bwl E E NE 1093
Anorthosites of Cana 563
ET and “se
he Mapaana Shore of Lake
3 perio
rcti c Con
Antivivisectionists i - n Pennsylvania 26
omus mu
Authon 64
Aphis persicoe- eae E Seuveceus 578
Aphodius pi obey Mani. ics 166
Aphodius granarius Linn............. 165
Aphodius ursinus a Ma ts. 165
Apodidae, Bernard’s 31
Appendicularia mossii 51
Pran proe ate Fauna of
Wyoming and Montana........ 1096
Arabia Petraea 267
Arachinda...... 389
ralia eocenica 971
Archeology 69, 173, 300, 579, 755
838, 907, 1023
Archean News 149 270
Penge ge MS auses
Area a A pues of European
Cot 173
teens ipus Fabs usri. 60
Arkansaw or Arkans
Arthur, J. j D in n Living
Plants
baat hal from the White “ages
Epoch, description of, E.
118
x The American Naturalist.
Tn gaudryi
Ascidia
Asia Minor, ve tbe co peepee
Aspidium cris
a a ee Anato-
mi
Astr. wis MRO
Fesinsoe ron lucayanum...
Atalaph a bra chyoti Ss
Atriplex canescens 1021
pugon separ ~ the Advance- i
Adain, A OTA 466
Avian fauna of yeki beds
OF Qacenlahd assein
joai Pocket i in certain Chame-
Ayes H., On the Genera of thé
Dipnoi Dipneumones............ 91
Azurite 276
ACTERIA and their Products. 556
E T a a PR ‘554
Bacteriology in its General
Relations, A, L. Russell
847 1050
Baden, Geolagical History of...... 1089
Badde ley
Bailey, L H, What is an Acquired
NEED 91
Delano hs kowalevskii........... 77
Balanoglossus, the New England
species of
Banks, N., Notes on the Mouth-
parts ‘and akg of Insects
and. Chilopodt i ise Feri 400
Baram LT and Mi, Dulit, Bor-
oie phon Of. ccs 1 FES
Basaltsa and” Trachytes from Gough’s
8
Batrachians 9 a Tegea Indias. vise 671
Potra hiii News, nei a sso 28
Batrachia fi from "Tex
mar, = Dasit of Testudi-
675
nen, G; bee Hantal of Mio-
cene Amphisbænians............ 998
ga G, Notes on haiak Box
77
ur, G., Rep tes on the Clas:
tion of the Cry PR cs <07
y, J-C., Spore-forming
of the genus Saccharomyces... 685
Bay of Fundy ‘Coast during t the
Beecher, es Sn ey eal Correlations
in the ee and Phylo-
geny of the Brachiopoda...... 599
[XXVII
Belonostomus sweetii 150
nse aia insertum Mots. siess eee 164
384
Bessey, E E., Daia Nomen-
E 47
ons e E., Botany at the Mad-
ison Meeting 823
Bessey, C. E., Allen’s Characeæ
of America 570
Bessey, C. E., Review of Dana’s
How to know Wild Flowers.. 653
man Ba Hough’s American
70
S r jN Review of Humph-
rey’s Saprolegniaceæ of the
UMEO AMES sioneer parate 664
Bessey, C. E., on s Naiads... 569
Bessey, C. E., Review of MacMil-
lan’s gat of the Min-
NESDIS VANE V cis centinsoses saene 5
Bessey, C. E., Review of Spald-
ines Gilde X the Study of
Com Plants 988
Bessey, C. E, Richard’s Develop-
ment of Caeoma TT NOORA 9
Dai, C. E., Shall we decapi-
talize specific names 21
B G E of
ames in desiginating species. 822
Bibliography of American Py 887
ig -» Review of Hae
sky s Cleavage i in Aequoria ra
cannes 58
Biological, aaas of the Iow
e University to the West
Indies and the Florida Keys... 894
Biological Station at Plon............ 17
Biology, Camp 245
Bird Bones (Neocene)........0. sses: 737
Birds, Mexican 742
Birds of Michig 096
Birds of Paradise and the Bower-
Birds ~- BA
Bismuthinite 903
ege ae hydrographical explor-
n of. 262
itean ias ni odias 573
tga fa antilopinus APITAR 662
Bles 844
Blowpipe pecan Landauer’s.... 46
at “a on, A., Legends of the
Sumiro-Accadians of Chaldea
14,1 410
Boliyina pe TREE 561
Book Publication i in “America. stessa 859
Borneo, travels i
1893,]
T pocer Natural History, 81
, 313, — = 585, 682, 1113
ienis Club, eedin pes of,
Madison, pele 18 913
Botanical Definitions 7
AE hee at Madison...... 738
Botanical 279
Bot ms Bd 2 474, 569, 664, 738, 821
Botan e Madison Meetings,
eel a 823
erence of silicious me init Retire 44
Boveri, The Organism produced
exua aed await aiarar
f the 222
Box Tortoises, PARTEA of, G,
Packscb i aeae ches Sorte anar 677
Brachiopoda of Minnesota Lower
Silu 735
i 45, 1007
ect C. L. Restoration of Osmic
Acid Solutions 175
Brook, Ja 917
Brooks and Bruce on ia: Embryol-
ogy of the Macroura....... sii 243
Brown Coal and Li lais of Texas,
Dumble’s..: 3
Bufo compactilis 156
Bufe debitis 156
Bufo punctatus 156
Bulima trigona..........066 seo 561
B s lauta I 165
Bureau of Ethnology 980
a rma = eee TOA TOM. cissi 65
of Rodents (fossil)......... 559
tng eyelophorus Kisbycnacna 16:
Byrrhus kirbyi I 135
ACHE, Long Bluff..ss.e..-......…- 616
Cacoma nitens 56
Calcium carbonate crystal..... 276
Call, Prof. R. E 411
Calyptrate Muscidæ 578
Canis ANCEDS icc ss cece: S TR
Canis lupt 5i
Cape Town University 817
Carabus oregonensi p becrreand 1
Carapace in ~ Devonian Crusta-
cean Rhino J. M.C _ 793
Cocboatinness Eustis of Centra
France 997
Carnivorous AFU 413
Carrington PERN 4
Case- worm from the Grand Cation;
a 1
Cauc i Rs s erat 261
Caudal "Fin of ran Rep- i
til es 65
Cecidomyia atriplicis............+.-.- 1021
Cecidomyia caryocola O. S........... 273
Indez.
xi
Cecidomyia holotricha O. S.. 73
Cecidomyia alae a Q. S.. iaie - 278
Cecidomyia tubicola O. S............ 273
Cecro se a cocoons 184
Cenozoic In 146
Genote news, 38 150, 271, 473,
561, 663, 737, 100 2, 1085
Cephalo-humeral muscles and the
so-called Clavicle of Carniv-
ora 52
Cephalophus spadix 56
Cercaria Stage of Amphistomum.... 151
Cerussite 44
Chabasite 884
io crue and other Silicious
Sp RRB ET anv bins: O RR 1095
E bálicaiké ME TERE OREO 118, 128
Chameleon isabellinus 286
Chameleon vulga 1095
Ch mpia parvu ula 979
C treme of America, Allen’s...., 570
CUO Mi a E ues, 1038
C baek- List of New North Ameri-
ant 821
Chelonia ERS iirde foriros 673
a ao e E E O 673
TNIV OVONCER. core sesesicssess scevcsaes 673
ara: pe ft a pati A 830
on marmoratus 1098
oie squamosus 1098
Christianite 884
Cinnamon Harvest Spider and its
Varieties. e M. Weed......... 534
Cladod ont SHARES nini ear 1083
Cladodus tepeleri 1083
Clarke, te t : Philosophy of
Flower Sea 769
Clarke, o n the structure of
the Carapace in tbe’ Devonian
Crustacean Rhinocaris ;
Relation of the genus to
Moothgma and the Phyllocar-
San s s Mesozois Echinodermata of
the United States.................
oe? wet Aequor forskalea,
58
Cams k’s ne on... 398
Clidastes velox......... sisereses 1985
Clidas aaa somsis OS
Chi Dwellers seess iisas sedeesses 435
CupE spiatteliaties thdoewrnebaee be 56
Clymnella elongata...........ssseeseee 1096
C te ahoga: sssi aeus 560
Coccinella transversoguttata a Fabr.. +- 105
occus cacti 1041
Cockerell, T. D. A. N on the
ochineal Insec 1041
Coelocerus grandis
FOOTE ee come ee seewee
xii
Cole, A. H., Solution of the Dust
copy
The American Naturalist.
Problem in Microscopy.....+++- 405
Coleoptera from High Mountains of
British Columbia 164
Coloceras globatum 872
Color Variations of the Milk Snake
E. D. ope
a "Anatomy, Wieder-
sheim 892
Cone-in-Cone Structure 0
` Congress oerna ry of the World’
Columbian Exposition of 1893. 1028
pos a at Teius Bey i. 1085
a a $ came of pre-
tag M olluscan Ova........... 1026
Conquest nt es Vegetable ¥ World,
ETS 811
E Poblet 16
Conway, W. M., expedited of..... 465
Cooke, C 170
Cookeite ......... 901
Cope, E. 038
Cope, E. D., Color Variations of
the Milk Snake 6
Cope, E. D., pare of Spea
laticeps Co 155
Cope, E. D., Fossil Reptiles from
iha Danan 76
e E. D., The Genealogy of
Man 21
Cope, E. D., A New Plistocene
Sa ae h 896
Cope. E. D., morium, and
the Position of the Cladodont
999
Cope, E. D., Prodomus of a new
system of the ek ra
Snakes var
Cope, E. D,, Review of Beddard’s
Animal Coloration 871
Cope, D.. Review of Calder-
wood’s Mental aran cutee 654
Cope, E. D., Review of Cary
Study 48
Cope, E. D., Review, ul The
Death o oo EER
i ‘ ew of Earle s
Species of albert -- 250
Cope, E. D., Review of Fritsch’
Gaskhole of Bohemia............. 1079
Cope, E. D., Review of- Forsyth
Major and Röse on the Theory
of Dental Evolution............. 1014
Cope, E. D., Review of Keeler’s
Evolution of the North Amer-
ican Land 47
Cope, E. D., Review a Wrights
Man and the Glacial Period... 550
Cope E. D., Remarkable Artio-
dactyle from the White River
Cope, E D., Vertebrate nies
ogy of the Llano Esta
Corning, Dr. H.
Correlation — a = Lt RS
Geol.
Correlations ol On + i lien d Phy-
n the Besthiopida, C,
ne ae
Corvus monedula
S Earle’s species
Co: ene, ao American ......
Coultero pre E
Crassidulus
sce
Cre apis EE of Mexico..
Cretac
atins strata on Staten Island..
Crommelin, Dr., Jan Pieter van
Wickevoor
Plants
Crustaceans from the Indian Arch-
ipelago
Cryptodira, classification of, G.
Baur
Crystalline Schis
rys roma l experiments on..
er
O
Cuprocassiterite ohian lioes
Currents of ain North Atlantic......
Cyclus scottii
G
Cc
ycl
yclopidius i es Sopeapvessertes
ynodesmus thooide
AHOMEY, coquest of.........+++
Daimon aire
Danais archippus. ......:....se0-
ney shee Redauth, Cornwall..
Dar after Darwin, Ro-
Dates sof is issue of Naturalist for 1892
Datholit
Davii, 1. W
an, B., N n the ga rine
Biological LAG atori of
Europe 625
Death Valley Expedition............
Deca preh from the Indian Archi-
DeCandoile, , Alphonse Louis Pierre
et tira a Se N SERS
Degeneration of es LSAtons E E
Dendrohyrax vali
Dendromys nigttrons. Sebscenes
or Ai oa.
Dermatemydidae........crsssvece psss
PE MOCheIyidne, sinesi nirondrerti
[XXVII, .
a ia EEA i or ee
1893.] Index xiii
“> pis crenidens 660 descriptions of new Fishes
Detection of Minerals 387 from the Northwes me | |
De opinen of the vertebrae and Eigenmann, C a ac. H. Bee-
he my ic coel in An- son, Pre ry note e
guis and Tropidnotus............ 285 prone 4 of the Species
Devonian Fossils from Manitoba... 469 peo A United under the Gen-
Devonian Plants 084 c Name Sebastodes
Diamond in Meteoric Iron.......... 1009 Eighth “International Congress of
Diapheromera femorata.............. 1113 American PONS 755
Diatomaceae, Mill’s............000. 00 892 | Eggs of Pityophis melanoleucus, J.
iatomaceae of the yea (?) P 878
Sandstone of New Jersey...... 817 ta 1115
eas maae EEEE 6 la aps fulvus 286
vinula 170 | Electricity and Cleava 396
Dieyorhabdus T priscus 268 ephas meridionalis.......0.sessssess 87
Dicynodont Remains from the EBlginia: mirabilige. kine: 876
lgin Sandstone......... asveves 145 prs es 57, 158, 288, 395,
Dinichthys intermedius 270 743 1097
Dinobastis 897 PUNY bonnie Of: Chilton R AnA; 1098
Dinornis queetetendise MEG 1002 | Embryology, Hertwig’s a aay
iopsides 275 Srog of the Macrura... EUR 243
Diplodu us problematicus.............. 149 675
Diptera brasiligne. s cninn 578 ra Aoo bleekeri 737
Disastrophus cocan O. S.. 239 | Engadine Lakes, origin Of..sssss...…. 1086
Disastrop vie sne us O., S......... 289 | Entomology 164, 293, 400,
Disclose ti UET 736 74, 679, 750 rases ... 904, 1104
eart of Ati rt Ribot’s. 33 Entomological SOMO 6 caves vedas 1106
rei m curtains StS yee < 1094 Entomological Notes 64......... 99 681
Diviessaie parador soi... 1004 ntomological Study 170
i gegia tumida 1094 | Eolite Pene from New Jersey.... 272
Ennes. 69 | Epigenes 289
Dugong, skeleton: ait ner of an Equus mi 812
51 | Equus sivalensi 39
Wie eri 1006 racia brevico 1096
oes ‘Sends in the Netherlands, Eriptychu ericant 268
382 Erosion beneath Dees Glaciers..... 662
Tichieation of Names of Aione 886 | Eruptive Ro cks, chemical nature of 1003
Dust ie em in Microscopy, A. eprinhe graminis 474
H. 405 eae: i BF
Etehed FIPS.. i iseen svete ke
Enoka @ 69, 173, 300, 579, 755,
ARLE, C., On the Pega 838 907
position of Pro otogonodon...... 77 | Euclase 384
Earth’s History, abate Sask 456 | Eudamns tityrus 1018
Echi ic EAE E 395 | Eulalie lobułata 1096
Echinoderms of the Eocene......... 473 nno: pire ME Jccbieses 1016
Echinus microtuberculatus 226, 228 Eunotosaurus africanus............. i
743 oe gen pholyphylls pasruar wwe
Stine hip pest Lowlands ete 467 Dorin SUCHE oss: fac ccs.
Ecuadorian Andes, Travels in...... 255 reie and Dichromatism of the
Binasi 26, 134, 238, 359, 450, Genus Megascops, E. M. Has-
54l, 1072 638
Edwards, A. M; tomaceae of Rilition of Foot Structure, Cary’s 248
the Triassic 0) § idiot of Evolution d the North American
New jersey. meg eeose ocsssss 547
cee ai Temperature o on the Col- Evolution of T Mammalia
of Lepidoptera...... “sa 1016 in its bearing aN the Problem
ichisenie crassi pes 1113 of Phylogeny............+0 586
Eigenmann, C. & R., Prelimi Evotomys gapperi rho ell isc 54
Xiv
co Aaah in the Delaware Val-
02
Explorations in the Amazon Basin 1115
Exposicion Historico Americano;
P Madrid Spain
Experimental Embryology........... 158
Extreme Case of Parisitiem, R.
Hessler.. 346
AN dentition 321
TOOS. rras of North Amer-
82
Fauna pe aos of Manuritius........ 379
— as sae! o the Eggs of Chinus
151
Fe lis ck ae eidy 897
Fertilization of the a ee. 285
Finger Lakes of New i sabaw . 1085
Fins of Palaeaspis americana....... 75
Fish Acclimatization on he Pacific
Coast 1017
Fishes (dwarfed) 741
Fishes lossil from the Upper
657
raae ce the Pacific Coast of
ca north of the Cerros
390
fois
ha per, of the Permian of
France 560
Fish a Fisheries of the U. S. for
Ta P A of " Laterover-
sion of the ‘Ophidian Heart... 860
1014
20
Flickers.
Kupat of Dads aa Lancaster........
Fol gerite
566
Forest Flora of New South Wales. 1039
* 1014
Fossil Booki, Hetin age of. 14!
Fossil Plan m Kootanie......... 1085
Fossil Pants fi from Texas.es. reocseaee 1085
Fourchite 275
Foxes in git Talend 1108
Triedelite 276
Trit-fly
fritsch’s Fauna of the Gaskohle of
Bohemia
Frog’s Eggs under Pressure...... ss. 1097
Fuchsite 090
—— < (Eeonomic) Seymour and 4
75
Fungi o NoT 474
Fungi (North P aia E voopees 474
The American Naturalist
[XXVII
ADUS tomcodus
Gall ona ene Bush, de-
588
scriprion o 906
Gall (Lepidopteros on Bigelovia. 680
Gall o ar Grand Cañon,
de: aenn o 05
epe on rahe ee fremontii, descrip-
E a TA T E cei seis A 904
Gall- veer ing Insects 296
Game Protection in New York..... 451
Canai lite 217
Gases in aside rises, J. C.
Arthu 1
Cikingniat, Frare ims a 253
"S and Cenia opoda of
New Jer er Cretaceous
Mans Whitfiel 728
Gastrulat mata Aurelia 70
eb elginens pote aSocciscteccssens 356
Geikielit 576
Geis oe pi 286
ere of North America, appendix
278
Gha of Man, E. D. Cope. 321
Genera ai Dipnoi Dipneumones,
H. Ayres 919
Genth, F. "A 314
Geography, a Chair of.........00+++0 314
Geography and Travels 253, 458,
730 894
Geological Congress........s-s0000++ 1031
eoon = ri cal of Arabia Pet- a
Geological geimi of ai 1040
Geological Survey of Illinois......... 238
Geological Survey of Georgia....... 450
Geological News 37, 149, i a J;
2, 73 1 1084
Geology of Eastern Siberia..........- 267
Geology of asasina — 34
Geology of Northern Afric: oot aE
Geology and Paleontology $ 34, 144,
267, 375 , 656, fo
813, 96 1082
Geology of Pennsylvania, —
description of. 455
Germa 1 Society.......... 388
Germ-layers of Amphioxus, Lwoff's 288
Germ-layers of Vertebrates........... 485
Glacialists’ ERE spiris eerseveshas 980
laciers e 817
Glaciers = ‘New ‘South Zealand.... 260
Glacial sandera ibo 37
Gla aieea 1009
marr a Te re; origin of.. 889
pede sadist wasa 108
sabes. basen be vunneneae’ bait SOME
ba et chir ragra.. 243
eevee eeeweee
1893.] Index XV
Gordonia duffiana 376 at Arrochar, Staten Island..... 1110
Gordonia huxleyana 376 | Homfray, 917
Gordonia juddiana 876 Honey. AE tio 1105
Go ia traquairii 376 | Hoplocepalus suboccipiialis RATUNE 56
Grahamite in Texa 561 | Horn-fly in Canada and ANES 63
Granite, formation of........-...ssse0. 70 | Horse, benevolence m TEL Mh, oni Oban cove 68
Granites of Argentina, a = a 565 | Human Embryolocy, Mino ath 140
472 | Hum nming Birds and Swifts, R. W.
Gr raphit 901 Shufeld 367
Grephai’ mien heap seseseeeeeess O78 | Humphrey, 412
rapte: Ichra 675 | Hyaena brevirostris 561
poe Richard Alt | Hykena recite Esie aan 560
Grossularit 4 Hyaena macrostoma........ssscesseeee 39
Ground Sii of the Mississippi stenus caribbeus 30
alley ö | Hyatt, A., Phylogeny of an Ac-
on of the Rattle of the Rattle- quired vean SKEER G 865
yd Hybodti basanu 1
Gu W A deed & 1038 | Hylocopa prorstsys 1022
wig A Moth Commission, C. M. i ymenocaris vermicauda,........... 99
dedavntees 128
Gypsy Moti, extermination of....... 1072
BEA, travels in 458
oo of certain European Ice-age 561
65 Igneou ocks, origin and
ae Serra a ck 63 _ Classification Of.ssssseressssesee 40
a ae H. 170 hering, Dr. H. von 844
Ha | Incolaria securiformis .........sses0s 662
Haue erium schinzii 51 | Indiana Academy of Scienta., 187
H lus innocuus Lec............... 164 ee ss i si 663
Hasbrouck, E. M., Evolution and oceramus problematicus............ 658
ere the nsect paraa logy, Wheeler S. 745
a A O 38 nsec mbr ryology, Henking’s S
Hatcher, ST aes , Titanotherium Beds 204 sale 160
Hauchecornite........cecsessssscssseaes 566 | Insects, Geneon 146
Hau von 918 Insects (injurious) 681
Hauyne in pum 275 | Insects injurious to Čranberries.. 578
peice vei Owls of the U. S., Insects sia us) of Kansas........ 297
is 727 | Insects (injurious) in Okisthoutn.: 65
Heat Conductivity of Uniaxial Insects of Sou thern Alaska.......... 294
Crystal, 1093 nstrum 7 567
Heleopelta 150 anhaa Botanical Congress... 279
Helvit 1006 | International Congress Ameri-
ematite 1092 Canister ne SS 3 579
ye a in Tertiary formations 561 rep onal Congress of Archeol-
RIN E E OE Sess A 671 ogy and Ethn nology, sketches
Hemimorphic ‘Compounds........... 1092 Of SOD, S02 Aisen +8038 579
412 International Congress of Prehis-
a E E E E o 68 to rcheology and Paleon-
Herderite from Maine kas 6 tol at Ehh meeting c.i
perei pide ‘An Extreme Case of International Congress of Anthr o-
3 pology
Hippodaaii à ai ie CURS 165 | International Congress of Zoology
Hippodamia 5-si Kirby...:..... 164 312 764
Hippopotamus, E serisi 318 pha engl fossils of the Trinity
Fippiscti FSR aeeai
Holland, W: J., Nest Building Invertebrate Paleontology of the
Hollick, A., Notes on the Geology Towa Geological Survey.. sorses 985 1078
xyi
eases
Lee, F.S., Review 6 Rowan
The American Naturalist.
[XXVII
Ipomoea carletonii..........0.0.00. ees LEO Darwin and after Darwin...... 1
Ischyodus egertonii 472 | Legendary Evolution 2 the age
Isomorphism 38 Indians n Vle 69
Iva frutescens 1089 | Lepidodendron sasea 1001
Lepidodiscus mil 38
E in Burmah 56 Lepidolite 90 1091
Fa Joess sel, J. G 192 Lepidoptera i in the ards coll
Joes sel, JG 411 tion of Insects, list of............ 297
ma Hopkins Marine Se oe Lepidosiren paradoxa 920
Laborat 491 Lepidoste n 473
Johnston- Evik M Lepteces ornatus ores irn 830
Joint F ormation among the Inver- Leptolepis affinis: Sauvage........... 657
tebrata, B. Sharp 89 jpe iiiaio dorais Sau- as
1 i 1
Journ ot es Geol vey: $60 ae Flora of the Dakota
Lie and Trias i in Taylorville, Cal- Liit eTe ht of Tesi. phic
Lome 470 eucite-Tephrite of Hussak.........
eukart < Festschrift 7.005. hiaai 411
Libytherium maurusium............... 147
Ko wee and the Irawadi.. 462 facheeuhaneks G. W.... ii 491
17 | Light, Wrehtstav na a 456
ia ee T ek of TERS zE i ee locomotion SE 0000+ 51
Inderhook. Faun Liobunum hemisphericum........... 389
Kingsley, J. A., Recor of North laaa ventricosu ood....... 541
_ American Zoolog toners esesssese 8 Liobunum ventricosum iyena
Kinoste idae...... TA 673 525
inosternum ees sessesosssesooe 676 | List of Plants rarely found in
Kinosternum penns 677 Staten: x eee >- 589
Knop’ Pe ae History of B Baden 1089 | Lithiu nsters 1099
Koksha TR 315 iomplio Rmestoné? RENNIN 900
Korsche 845 io a in the Tauk Rosse.. 382
Ku es Terbis Generum Plan- Litton a lores 1112
Tarn ete Ea r 1010 | j re 681
Loess in Southern Russia............. 269
ae decagonale.... 39 | Loew, Prof 1115
561 Longipennes, een of, R.
Sen T, p3 Tign of Birds 20 W. Shufel 233
Lancelet “et the Bahamas........ 830 | Lorsen, Prof. K. A 593
Language versus Anatomy in deter- xonema winnipegense Perc Sevens 662
mining Human Races............ 579 | Lungula monesii............scc00. sees. 736
weer —— seteeneeer eesees 66 ake ators orin a Atriplex, C. H. vin
of Batrac 154
Lateral Line of Silusoids ecg 477 | Lydekker, R. 1088
Lateroversion of the Ophidian
He - 860
one and Huronian Rocks ACACUS tolosanus..............
die of Lake ens relations Muacdouyal, D. To cia. 17
996 MacMillan, C., The Proba-
Mee of the DE District.. 996 le Physiognomy of the
Lavas of a Terti ONE iiei 275. Nt eous Plan T
marie æ Ne] Bibliography of, 317
Leaia 271 | Macleay, “Sir. ‘William, ‘scientific
Tei a the Paris Museum...... 592 memorial volume in honor of. 1039
ium hyalin »- 484 | Macrochelys lacerti 742
Leechs o iae Madagascar fauna 982
Lee, S: F., Review of Ribot’s Dis- Magothy TE T 662
of Personality... seeeecees 33 | Maioid Crabs ji in the National Mu-
=
ET «4 ine
Mites by
1893. ] Index xvii
Malchite, description Of...ssesse s.. Mesosaurus pleurogaster...... 85
Mammalia (fossil) in Southern M id 35
Patagonta.....crcccscrsesen eomeers 439 | Mesothyra oceana..........c0+ sesersees 796
separate" from the Galapagos Is- Mesozoic Echinodermata of "the
United States.......ccisssscsveesie 079
k, from the Pits of Gargas. Mesozoic News 38, 150, 271, 472,
Mammalia News 72 560, 662, 7 1 1084
Mammals of ihe Deep River Beds, Metacoceras da a N 867
hide B. Scott etasperma of the Minnesota V
Mammals, Mexican 742 ley, pea BEO 865
Man wand ig Glacial Period, Metat atiformis Hyatt...... 866
550 | Method of injectin the Blood ves-
Man bes righ 321 sels of Birds, W. S. Miller..... 582
Mantle of Ascidians 486 | Mica Perodite............c..ssecosese ve: BE
Maori dentition,....00 ¢--.+sers+ssees 321 -chemical . 903
Marine’ Biological al Laboratories of Micro-chemical Reactions............ 387
Eu 697 | Mi LT SOD anteoesens: 1026
Marine Biological a Laboratory = Miller, Prof. a
Wood’s 841 | Miller, W. S., Method of injecting
Marine ne Laboratories of aac B the Blood ben of Birds...... 582
625 illerite ssov Ef
limes position of......... 283 mee ee to 276... = 1006
ite 44 1092 and rography
Mastodon borsonii 561 272, 3 380, 562, 398. Akis ba ‘1008 1087
Mastodon oligobunis.......+.00 onsbie ee Mineral Resources of the United
MoGee, Wf scsiiseises secetisccanent LODO States, 1889 i ARRAN
Measur iization...........++++ 174 | Mineral ral Resources of the United
Mechanical Evolution ne oe ome 1891 816
apodial Keels of Diplarthra, J. erpin er SEEC 45 567
L. Wortman...s+e..sssse0eeereee- 421 | Minerals from the Diamond felds
ical Genesis of the Scales of Brazi 383
Fishes. 1 | Minerals, prear peye of..... 1092
Medals psn ar: by the Columbian Miocloenus pentacus.,......-.ss.s0-se0 377
Exposition at Madrid....... 316 | Miocloenu: 377
pee ee rs depths of....... 262 | Mirabilite changed t to Thenardite... 1016
rom Lake Tanganyika... 571 | Möller, Dr. H 1115
are madagascaricnsis...... 1002| Molluscan News...
] eeeceeseccesceseseecacesscnees 638 Molluscan Ova, ITAR poomi
Melanostibian. ..............se.2222-.... 901 ing 1026
Menten „Massiv ” andi its Dyke Mollusks, cretaceous....... desservie: aw
De B pen Mollusks of Lou socteenscsamhid
elilite lusks of Water Mains of
Membranes of the Sea Urchin Egg. 397 _ Paris ii 1094
Memoirs of the National Academy MOR wise tidecieidivieeecses SOR
Sciences, Meee regen e ria parisiensis...s..s..0-. 272
M e Eran ues ae oe oiimhih sssr
Pi Monocladodus pinnatus.........s00e00
Mercer, H. C., Trenton and Som- onstrosities, exhibition of. ETTET 450
rel Specimens com- Moon’s Face. ‘ 734
red with Ancient Quarry Moore, T. J va Se
America and Eu- — pe É 708
a C. B., Shell H of the
K. S. von 317 Moore, “
T E Sic PAT TEIR, 270 | ii so John's River 8, 113...600: 708
se wa, tyme peo melanoleucus 878, Errata..... 1115
——- chel Sr GGL :
intermedius.......-.--0= => | Moraines in i
el ama
XVili
pE i at Root Tubercles of
osae, s Schn ae 782
a pig pued j Caa RNE 7
T ate and Thor x of Insects
E Ehifopa ds, N. Te
ETTEN 1082
Mummied Child. from the Cliff
lers 437
Mus aeie
4
56
Museum for Chicago 886
Musteline from the John Day Mio- `
cene
658
Nigar SEAE, E R 475
Naiads, Regie ON SS eee, ck 69
uta 87
eter S "Pola ti 263
TEA Murine Biological Labora-
136
Naples Zoological Station 491
National Academy of 'Science...:... 585
atrolite 903 1007
Naturlichen Pflanzenfamilien, Eng-'
er a itl’s 49
Nebalia bipes 799
ebalia geoff oyi -800
N ebraska Aca ademy of Sciences... 80 |
ebri a Piset: 64
ctonema ag VARA
Neocene i Hra Nevada.. a! 1084
Neotoma ences Fanit cee 574
Apor ane! and Leucite Rocks of
Brazil 274
ba tie of Amphioxus,............. 283
Nephrite of api Zealand: 582
Nest Building F a OT
Neusiedler I kes Rensi 730
WeustiqosauUtign: si i 0
Newark System 987
Newberry, Je -Sovsiivwey scerna 83
Newberry Library of Geok sitis 164
in the sil
Mianais.: of Southern Patago
nia, F.. Ameghnio........00.0.608 439
New G , travels in 466
New Mexico Society for the Ad-
vancement of Science............ 310
Naw Minerale
New York Academy of Sciences
80, 182, 308, 409... vereeeeee 483 586
Nickle ‘arsenide à
Nile, sources 731
Ninth “tntermationa Congress of >
Rakes: sense sean 838
Noctuidee, N North American......... - 1106
Noll, F. C 411
Norian Rocks of Canada............. 656
The American Naturalist
885 | Orth possi
[XXVII
Norites in the Eastern United
Stat es.
hore America during Cambrian
ee
Notes on. the -Cochineal Insect, T.
D. Cockerell -041
Notes on the, Mollusks from North-
western. Louisiana and Harri-
son Count Texas, T. W.
Vau ghn 944
rope albzolus 152
tropis het terolepis 152
Makro jordanii 592
N tropis reticulatis 152
otropis scopiferus 153
otulinia noctula 53
Novaculites of Arkan 42
oxious — aia Tala lk
tin of
Sinikari JODESIANANT. iigiin
Nutting, C. C.,. Biological "Expedi-
Oe in West Green-
]
468
Odontophorus consobrinus.... 887,
Officers of B aa pa heme.
Brit. Ass. Adv. 593
Olivine : ` 1008
Om: s phillipsi 1001
Oolite, mee ation’ of.
Oneonta sare ciie mung Fornali
tid comin New York. 558
Picea 833
| Ophibolus doste Ti in 1066
Opossums on eeg Telanid ae .. 1108
Optical An 385
pre: aeria sexually with-
characteristics bad the
Th. Boveri ;
CO. sesso ee aee
ipewe Insects ‘of the Gala-
_ pagos Islan ae
Peewee eee ene e nese
1021
— H., Note on Trichodactylus |
xylocopæ
H. F., The Ancylopoda,
ohi and Artionyx.
Osborn, H. F., Recent Researches
ession of the
eee eee ewes eres
Otocoris alpestris Linn
Otaria jubata
— ite pressing Conglomerate ros
Bagg Wickens
ALAEASPIS americana.........
eee Beene od OF dssisyinn
Paleolith
Palæortyx grivensis. Eessbddsssssesoisos
Palæortyx maxim
apeta inta
Palæotermes ellisii. ee Keh diiva
Paleontology 34, 144, 267, 375,
380, 469, 558 558, 650, 734, —
Paleozoic
Pekcan Formations " —
Min
oa of Geor
Paleo iy
Pidesi pant 37, 149, 70, 479,
-s eet awweewwee AU
pd 5 eae Books on
rio
Pantosteus ini Aeae TEAS ga ”
Pantosteus discobolus
Bocsvescddcccssecd
enon eeeeeeeae
~ Seen ee ene eeeeee
RERAN
Peary Expedition of y f 1808 eens 6
Index
oy
—
ea
2 1003
83
Pedal RON of the Dorking
1
PEET 56
Peik- Tu: E Sian, A visit Oleana tiss 461
Pelias 299
elope carer pee wenencdas 279
45
Sasori = Cecussatus.. cesa sissie 741
rogna athus merriamii 573
Peronospora echinospermi.........., 474
Pers 465
Petrograption! News, 42, 274, nih 3
1005 1088
Saki eA 40, 272, 380:.....: 562 898.
Pet mcg aphy of the Abukuma,
Japan 562
Petrography of ans Japan... 380
fitzner, Pro 411
Phalangiidae, pore üa, C. oM:
294
Phallogaster saccatu 280
aneropleuron te Whiteaves, 817
Phone lomys mitche 88
ascolom ‘Paths. sissors 38
Philosophical Congress............... 1028
earns © of. Flower Seasons, H:
rke 769
T arsam ogre sestese 150
ae Of, the FI on 1087
n 586
Piieeiy, of a an n Acquired Charac-
ristic ya 865
Physiology, foakite’ oresoresbeotanes es 654
Piersol’s Histology. 1077
Pinicola enucleator Linn............. 587
Pityophis melanolencus..........0. + 878
iaul awsonii 150
Planorbis contortus 151
ee. ° Bahamas, Jamaica
Capriati idsccciwceses
Pans, fossil gine aA Beds. ine 0
PUA ster nine gs cos ssi nevorseds irisa 674
Pate hee att EN OT E 674
Paid ayes nan Linn.. ssesessre 7
Plaat ussia...... 819
iocene ‘(Mari soy Beds of the
Carolina 471
Podabrus iae Alii isisa i105
Poikilite 900
Polar Regions, explorations in...... 468
Polar Regions, travels in............. 263
Pol rieten aot ien = CSF
P ybasite r 43
Polychaetes, ey yes 388
Po moa POET CR ODM icin voce cs «xs 281
Poly, onum, pringlei serrer sceesns... 281
Pontine Marshes, scheme for re- ;
Popenoe, Prof. 170
; 653
The American Naturalist
xx [XXVII
P yry breccia 1005 Pyroxenite 1088
Portage Stage... 001 1084
Pangean Niak. lens Spe- 845 Qoi Wi of Human Life, J.
x x Williams r3
AP nne Modint Midd. Dak 471 Qnarterly Bulletin of the Uni-
aed ieee f ideas versity of Minnesota........ 412
wpe ie ee ne Ma 383
ig "acne m Plantarum, Quercus brittonii Davis 587
P Err 1010 Quercus ilicifolia TN PINEA 587
ound, d aana and ee: 587
8 Lichensicccsccscoececcooe. 509 | Quercus nigra a L.. TER ARRE ;
Pani, t, Prof, R 593 Quicksilver Ore Deposits ADAS oo BBs
Pratt, Miss Anna.........+..-- eteasens 917
Preserve for] Rae. Zemani 3 ABBIT’S Feet, igi of. 672
<E 317 k Radiolaria archean............. 149
Pri neeton Meeting of Societies....... 135 Radiolaria, plcocsin bint 149
Prim -& B 669 | Radiolites davidsonii..... sess 1088
Priority in Nom WOE DENEAN 134 | Rampheoleon spectrum. ........... . 109%
rize for best method of destroying Rampheoleon herstenii 1095
pernicious son ts se rem For- Ranunculus r 280
sts of western Prussia......... 15 | Recent Books or on gr ane r e
mua hie Physiognomy e the Cret- Pam
Plant Population, C Recent Beaks: a Pamphlets 28,
MacMillan 336 137, 240, 361, 455, 444, 650,
Proceedings of Scientfic Societies TOE: BOG D.. arrika 074
8), 176, 306, 409, 488, 585, Recent Tiea 31, 140, 243,
G82, 702, DAB sissi -1028 1108 364, 455, 547, 653, 727, 811,
— of a new es of the
venomous S. » me. D, Recent Researches upon the Suc-
cue 477 on of the Teeth in Mam-
eS SCE te 493
Prosopon etheridgei. severeececssesees 271 | Recent Smdies in "Carnivorous
ocaris marshii.........0000+ pdboase s, J. G. Smit 443
Protoceras arisi Miriio oiai 137 Tiea oe in Ppr
149 a TOENE paapne 1006
Protegonodon, preemie position of pas an Zoology....... 85
fC. E 377 | Reithrodontomys pallidus 835
iba, sarange at 812 | Relationships and Distribution of
Protolemus 560 the North American Unionidae
rotolemus paradoxoides............ 560 with notes on the West Coast
Protozoan News 484 Species, C. T. Simpson......... 303
P seneeesececeorsesescssesenes Relationship of the species usually
ee Science nee Congress.......+. 1029 the generic name
171 298 Sebastodes, C. H. Eigenman
Paylla TA RE A AEE 293 and C. H. PE E EERE ae
a 20 maculata Say. PER 165 ; Report of the Minnesota Geologi-
OE eemrenesenehshenersess se Gk 1085 pa a ERKE History Sur
Prrodacty from Kansas.........37 1085 91 811
E. & i 670 alkar r pe U. S. Geological
essatork classification of......... 87 Survey, 1889-90, Part I....... 999
io onda riparius Dej........ ... 164 | Reptiles and Batrachians of Wis- a
43 consin 7
Parent -Argyramoeba oedipus Fab., Reptiles from the Elgin Sandstone. 376
cription o: 60 mepa sarg from the Parana, E.
bip eaae d Blepharipeza, descrip- age DE
204 > sea Seer e eee ene nerene eee 255
"a Daa ae Rentiles from Texas 672
eames > 1006 | Review of CER: Animal Col-
TIERS schon a’ soniethoe costes erin 277 oration, E. D. Cope......0.20-0. 371
Ee aE SS R
FETS ES ER ee re an eee pe ed Ne ey ee eee eae eee
egies eas
OS erie are PT eee ree Sree se eer
1893.]
Review of an iy in Foot
Structure, E. D. Cope.......... 248
Review of The qi Valley Ex-
990
ias 250
Review of E Fritsch’s Fauna of the
le of Bohemia, E. D.
1079
seeeee
Review of F. orsyth-Major
Röse’s eens: of De
and
ntal Evo-
lution, E. 1014
Review of Hacker Cleavage in
Aequoria forskalea.............. 58
Review of Humphrey's Saproleg-
niaceae of the United States,
co: . Bessey
Review of Keeler’s Evolution of
the Pa h American Lan
Birds, E. D.:Cope..........000.. 547
Review of MacMillian’s Metasper-
the Minnesota Valley,
Glacial oy E. D. Cope...
. 988
550
286
R iba... 286
Rhinocaris columbina.,........... 795 797
Rhoads, S. N., Descriptions of 4
Rodents from California....... 831
Rhoads, S. N., Description of Sy-
naptomys stonei..,............... 53
Rhyolite in Maryland and Pennsyl- al
Rhytina gigas Linn...... Pie Resa . 490
enter iatea Š
Rocks i at Cingolina. binding
Bails aidie La Si PEREA .- 1089
Rocks of a New Island oft Pantel-
leria... 1088
Rocks of Odenwald. aoe ENAP AES pe
Rocks from Siberia...... 381
Rocks from Southern Borneo....... 899
Rocks of the Thalhorn..,............. > 2
Rocks of the Valley of Mifior,
+a from Californi
Rosenbusch’s Volume on Minerals. 567
Rothrock, J. T... 83
Retcllite 1091
Index xxi
General Relations........ ---847 1050
ABACON ama EURE TAA N 575
Saccharomyces..........0.:sss000 685
Sacken, monet AARNE RA
lt Crystals...... PE NR SEE PAANS EA
Saprolegniaceae of the United
ates 664
in, M. M 593
arcolestes leedsii 1002
Scaumenacia curta 817
Schaaffhansen, Prof. H.............. 412
Schists of Berkshire, Eng-
land 1087
mre wake. od derstand of Root
of Leguminosae..... 782
Scientific eon Be: Bem ee a
592,
3 1115
Scientific Publications................ 072
Siwas mndalates. 6.0 cc6 cscs ocese sis
Scott, W. B., sere nels of the
659
Scott, B.A A ener Musteline
from the John Day Miocene... 658
E.F., n E TER 316
Sea-urchin 743
Sebastichthys Gill 669
Sebastodes .... 668
Sebastodes 670
Sebastosomus Gill 670
i 149
Seitz, Dr. A 844
Selénia illustraria. «..........00.0...00+ 1016
RES cr ni oar me cath awenti teat 1091
Semper, Prof. Carl......... ...0-++00 845
Serropalpus barbatus Schall.......... 166
Sesquicentennial of - ae
Philosophical ROTE
iiep; = ses ama
Shasta-Chico series. ..........
Shell Sae, ti ats the § St. John’s Rive River
Florida, C. B. Moore 8, 113,
605 708
Shimek, Pro 317
Shufelde, R. Ww. Classification of
nes, 2338
sr rt: S i
5
Sitomys heranii..........5.0.0ccersescee
xxii
Sitom mys
befe Ga R
Sei Plants o rere
Shaken; (CCIE Ol ies 50st scr secon
Soda lite-Syeriite from Montana......
een oes COMBE, Billings reves
Solidag
Sollas EDE g p e a
Spade- foot, new from Texas.........
Spaere echinus Con 227.. ei
Spaldin gs s on vr o the Study
om 55
parrow, English, orali of. otic.
Spea laticeps Cope
noe Names, decapitalized..
Specific N.
sete eee
, Pateosote Group ca
rl
tk
Sahihi a
Spessartite
sessessososss
Cer er
Mexico ` a
6
Spore-forming species ni Dsi gents
Bay R
The American Naturalist.
831
185
Saccharomyces, aX sooege , 685
MALON, Fig Laras oa icy SO et mo 814
tar-fish ] BETO Soicc tosis cvuns n Soe
Staten Island Natural Science Asso-
ciation 182, 488.............. 586 1108
St PICRO rs.) seas AA eae ie oe 2.67;
wnat ME el TRE DONE 474
Steneofiber ntanus HIY 660
Ste: siop s hi pieta 90 4243
Stereosternum tumidum ip, BD
Stibarus obtusilol ; 148
Stichodactyina 8
ae MA /dtictiption of roion,
ad eo a a 5
ri ‘Alexa AUA ee ER E O 038
rig ocephelus zone in Mani 471
. 66
Strix ianeriaiee on
umiro Actedians of Chaldea
Depends o are Bodington 14,
105, 410
Sundtite
Swifts and Humming Bids. EAR 573
Sylvia hort 65
ieia stonei, description of. 53
Syneta albida Le 166
; sees and Maina in Lich-
Symmorium and the Position of the
t Sharks, E, D. Cope,
999
AE E HA
[XXVII,
Synthesis of the Members of the
Sodalit Group
Sycteinatic position of the Kiwi..... 892
AENIOPTERIS missuriensis. 662
uas dentition 321
902
“apir 428.
poing crassus 867
e it 43
ZA 471
'errapene aa LE a SRD E E 677
‘errapene major Ag 677
Perr T mexicana os ne ae 677
ferrapene ornata 678.
etrapene ag aa a S POR SA 678
Termitophilu 400:
ertiary 1 tases froni Gikaa and
558
Pei 672:
Testudinidae 675-
Testudinoidea 675
Tetrabelodon serridens 812:
Tetrabelodon shepardii Leidy 473,
12
Texas Geological foes hf 1801.. Tae
Textularia manly 560
Textularia 560
hecla canines 18.
Thomas, M. B., The Androchonia :
vi ‘Lepidoptera E E seine 1018
Thomomys..... 337
Thonvortr a Tk i E AE 1096
Thomomys’ au 837
Thomomys fossor 837
Thomomys monticolus 837
amna toltecus 837
Tibet, exploration of. 465
M aieea Beds, J. 5. Hatcher 204
otanus majori T.
Tourmaline
Townsend; Co PT 2 170
Townsend, C. H. T., Description
of a Case- -worm from Grand l
Canon .... 166
Townsend, CH. T., Description `
of Cecidomyiá atriplic | RARE 1021
| Townsend, C. H: T., Description
of the Popa of Argyramoeba
oedipus Fab 60
Townsend; w e T, aapi po ecw
of Puparium of Jurinia. 576
Townsend H.T., Hym menop-
terous Gall on a Credo te
7 Bush 905
| Townsend, C. Lepdopte- `
Tous gall ¢ on pahe 680
1893.]
To C..H. T., Lycaenid
larva on ae ETE TEUA E
Townsend, C. T., Puparium
; ceeded
o
Townsend, C.. H. T., Spiders col-
lected in New Mexico and
Ari io
Towns, Cc H. T., Tineid Canes
n Po opu ulas fremontii
Townsend. C..H..T., Wooly Leaf
sit on Oak near ee calles on.
ndesites of the
rge
Trenton ye Somme Gravel Speci-
mens compared with nanan
Quarry Refuse in America and
et bes Mater: DO
‘Trichob ta
Trichoeactylas BESRA in Cali-
fornia
Trilobite pher “ Skiddaw Slates.
Trilobite, 1
eee cristatus
Trit palmatus f
Tropidoclonium 1 lineatüm,. soss..
Tunicate Stu
Sa cevveccsvcesscescessces
Ne
t
=
“ao
ri
=|
=!
a §
=
n
‘Typhlotriton speloeus
CA gti one , exploration of.
ropoeus L
ween ene
Unionidae, North A
Uplifts i = the California ‘Shere. Ee
Uranopbait E E eens
Urous SECT i. 025 oira vies eee 5
Ursus speloeus. PS EE E iin
PYRUS spettim.. sisse
Vanellus ¢ eng cbeavevercceess
Van Vleet, T Legendary
GEET of the ane Indi-
Variation | ~ the snakes of North
eiit itie Dyke i in one among
Vaughn, T. W., n Mol-
lusks from nm L
Index
1104
402
53
65
S
1003
ay growths as Sidat of
or impurity of Water...
Ver la
Vertebrate dw oe or the Ordovician
mn
news
Vert 1
araa Daae aa, of the
Aci Estacado, Texas, E. D.
xxiii
viisiobiis 811
Vaals 44
Vetter, 412
Viallanes, Henry 844
ine, G. R..... 917
irchow, Prof.
Vision in. a girl 6 years of age,
operat aie cd w double Con-
oe Sr A TE A 171
Volcanic Dus Te 663
Volcanic “seater in i Coline. 813
Volcanic Rocks of the Andes ...... 898
I A. S., expedition
64
i DE T B iiias 844
sae n An nthropological So-
ert 81, 187, 410, 488....682 737
Washington Biological Society 81,
187, 410, 488, 591 ......... 682 1114
Washington Geological Society 591
udar ai ah 1 Among the Cite
Dweller 435
Weed, 170
Weed, € M, An American spe-
cies of Sabacon 574
Weed, C. M., Cinnamon . Harvest
Spider and its "Variati Ce EEan 534
Weed, C. M., The Gipsy Moth
maiin 750
Wecd, C. M., North American
Coimeldae. sie. siise 74
Weed, C. M., Notes on Ohio and
other Phalangii IABE, siiis
Weed-seeds, Halsted’s............. 475
Weir, Jas., cog Intelligence... 933
estw > 192
hoo cen R. von 317
PEA E G DEA ss scewede ss. < OO
Wheatland, Dr. Henry, vk. onc. 410
| ia Clays of a “Ohio Region... 148
Wickham, H, F., escriptian ns of
Daie ra from the th-
west 164
ee Lda; the sprees of
Hotties Life. cocci: 193
Wolle’s Desmids 47
Woman =o 321
Wood-Mas: 845
seamen ys of es Soin walk, Lum- in
XXIV
The American Naturalist.
[XXVII
Woodward, A. Sm ATE | Lapan ea erica rieres coissrsosivsss 672
ba. Alder er iphis. sseseseessee 400 402 | Zeuzera oesculi Linn 586
World’s Congress Auxiliary of me uglodon CaUcasiCus...... ssesss sses 663
wun Catenbints Expositio; Zoological News 56, i 285, 484, e.
of 1893 762 TE, 741 837
Wortman, J. L., A New Theory of
the Mechanical Evolution of
the Metapodial Keels of Dip-
larthra
421
ENOTIME 901
Xantusia henshawii............ 837
AMITES montanensis............ 1085
= Diras ELLELE ETETET
Zoological Congress......+..:ssssseeee
mae eea and the parenie ae Asso-
on 667, FAR EPER ENA ...829 1014
ogy
eres of ste Lower Saskatche-
wan Riv
Zunyi
Zuyder Zee, reclamation of,,.......+
AMERICAN
NATURALIST
A MONTHLY JOURNAL
DEVOTED TO THE NATURAL SCIENCES
IN THEIR WIDEST SENSE.
MANAGING EDIT
Prors. E. D. COPE ann J. S pE
j ee a EDITORS: ;
Dr. C. O. WHITMAN, . C.-E, BESSEY, _ THOMAS WILSON, ;
Pror. C. M. WEED, Pror. W. S. BAYLEY, Pror. E.-A. ANDREWS. —
J ANUARY, 1893.
CONTENTS.
PAG
Gases IN Livinc Prants. (To be continued.)
Arthur.
i HEAPS. oF THE ST: Tii Kiven,
mic nae ee OF CHALD
ued.) parsen
FLIGHT OF erh (Mlustrated. = 3 Lanéaster,
— as ai San in Penn
Com Pea Expedition ;—
Senet ofi issue of M Naturalist for 1892.
Embryology.—Gastrulation p ure
Darwin, $ im Aequoria cise alea å
Frit-FI ao rh
and P Pelentogy On 1 the Formation of $ iere al — Notes on eS
ortheastern Alabama—The pean Birds—A Nest Buldi ing Frog Hore
a of South Africa— eens as Ptrerodactyls— kaer
eral— onl
jal News, General- aleozoic Mesozoic— a : 7 ae at ar E thn ogy Leg endary Evo
| on of che Nayajo Indians. .
aey a nd Patregrap iy. y — The Origin and ` PROCEEDINGS OF santas Societies,
Petr eapi ical Ne ws—A New Occur- | Sctentiric New
ot Pe seat News—Mineral ~ RECORD OF Noita Maske ZooLocy
PHILADELPHIA, yU. S A.
BINDER & KELLY,
Oe ano 520 MINOR STREET. >
Ch. Marchand’s
= CH. MARCHAND’S
_ PEROXIDE OF HYDROGEN
(MEDICINAL) H202 (ABSOLUTELY HARMLESS.)
MOST POWERFUL BACTERICIDE AND PUS DESTROYER.
ENDORSED BY THE MEDICAL PROFESSION,
UNIFORM IN STRENGTH, PURITY AND STABILITY.
RETAINS GERMICIDAL POWER ANY LENGTH OF TIME.
DR. E. R. SQUIBB, of Brooklyn; N. Y., DR. ROBT. T. MORRIS, of WN. i
DR. EGBERT H. GRANDIN, of N. Y, DR. JOHN AULDE, of Phila., Pa, DR
W. F. WAUGH, of Phila., Pa., DR, GEO, B. HOPE, of N. Y., DR. E. CHAREST,
of St. Cloud; Minn., DR, G. F. ADAMS, of Pulaski, N. Y, DR. H. F. WIGGIN,
of N. Y., DR. J. A. LARRABEE, of Louisville, Ky., DR. R. M. CHASE, of Bethel, Vts
DR. ROBT. T. WILSON, of Baltimore, Md., DR. J. H. SHERMAN, of Boston, Mass.
DR. C. M. WOODWARD, of Tecumseh, Mich, DR? GEO. H. PIERCE, of Brey
NY and many others
NOTE,_Avota substitutes—tn ee ef the commercial tea gai soat
orthless to use as a medicin
Marchand’s Pevesinne of Hydrogen or nee jane aeg ja only in 40%)
S-0z., a 16-0z. bottles, bearing a blue la os Sai © letters, red and gold
border, y ih his s signature, Never: sold in b
_ GLYCOZON Ea
TS ee hice as ene IN THE a
OST POWER LING PURPOSES.
DYSPEPSIA. GASTRITIS, “ULOER OF “THE STOMACH, HEART-BU
ase _ Glyeozone is sold only in 4-02.,
8-0z,, and 16-0z, bottles, Never sold in balk.
___ PHYSICIANS WILLING TO PAY EXPRESS CHARGES WILL RECEIVE
Boru OP THE Above REMEDIES
"ARE PREPARED ONLY RY
me F Mention this publication,
and Graduate Of the “ eola Centrals des Arts et Manufactures de Paris (France).
a —_ paucaisrs. + Laboratory, 28 Prince St., New Yi
AMERICAN NATURALIST
VOL XXVILI. January, 1893. 313
THE GASES IN LIVING PLANTS:
By J. C. ARTHUR?
The present state of knowledge regarding the kinds, sources
and movement of gases in plants does not constitute a com-
pleted volume. There is much yet to be learned, old views
are to be corrected, and alleged facts are to be more firmly
established. The subject is thoroughly modern, the first
writer to give any connected and intelligent account of the
behavior of gases in connection with living plants being De
Saussure in his brilliant and epoch-making work describing his
chemical researches upon vegetation, published in 1804.
THE COURSE or Discovery UP To 1865.
The various life functions of plants have been slowly estab-
lished by first assuming them to be individually the same as
those of animals, and from this basis gradually evolving their
true nature. The early naturalists saw nothing in plants that
suggested lungs or the movement of air, and it was not till the
time of Malpighi, 1671, that breathing was supposed to have
any part in the plant economy. He saw in the wood vessels,
known then and long afterward as spiral vessels, an analogous
1 Read before the Biological Section of the Amer. Assoc. Adv. Science, in Wash-
ington, August, 1891.
? D, Sc. Professor of Vegetable Physiology and Pathology, in Purdue University,
Indiana.
T
2 The American Naturalist. [January,
set of organs to the trachee of insects, and therefore believed
them to have the same office. His views were accepted by the
Englishmen, Grew and Ray, who wrote about the same time,
but found no supporters in Germany or France. The views
of Malpighi, who in many respects was far in advance of the
other botanists of his age, fell into obscurity, insomuch that
even the existence of ducts was finally denied.
The subject was not revived until 1715, when Nieuwentyt
demonstrated the presence of air in plants by placing sections
of stems in water under an air pump. The demonstration
was better performed by Christian Wolff, who was a philo-
-= sophical naturalist of much insight. He placed leaves, wood,
and other parts of plants, in water, freed from air, under the
air pump, and after seeing the bubbles rise from the tissues as
the air was exhausted from the receiver, he allowed the air to
re-enter the receiver, and found that the tissues were at once
filled with water, and that some kinds were thereby made so
heavy as to sink.
In England, a few years afterward (1727), Stephen Hales,
the real founder of experimental vegetable physiology,
repeated and improved upon the air pump experiments, but
used his knowledge to explain the life processes in a different
manner from his predecessors. He combined the fact that
gases are recovered from plants by dry distillation and fer-
mentation to support a well arranged theory of the use of
gases in forming the solid parts of plants. The use of gases
by plants was, therefore, according to Hales, a part of the sub-
ject of the nutrition of plants.
But this small advance led to no further developments, and
was again lost sight of for many years. After nearly half a
century, in 1771, Priestley hit upon a discovery which, coming
as it did only three years before the discovery of oxygen, and
only shortly before the re-organization of chemical theories by
Lavoisier and others, proved very fruitful.
Priestley’s discovery was simple enough, amounting only to
the fact that plants give off oxygen. He tells of his discovery
in an interesting way, and I, therefore, quote a few paragraphs
LEA Ti Sy Dido ae
PW gt) NRO olen Rese Pier a
ee?
oe eee et eS LER è 3 ae aS
sha AN E E EA I OENE N A E O ts May ETN
Nr ii ep ta NRA
1893.] The Gases in Living Plants. 3
from his communication to the Royal Society, announcing the
matter:
“The quantity of air which even a small flame requires to
keep it burning is prodigous. It is generally said that an
ordinary candle consumes, as it is called, about a gallon in a
minute. Considering this amazing consumption of air, by
fires of all kinds, volcanoes, ete., it becomes a great object of
philosophical inquiry, to ascertain what change is made in
the constitution of the air by flame, and to discover what pro-
vision there is in Nature for remedying the injury which the
atmosphere receives by this means.
“ I flatter myself that I have accidentally hit upon a method
of restoring air which has been injured by the burning of
candles, and that I have discovered at least one of the restora-
tives which Nature employs for this purpose. It is vegetation.
* * * One might have imagined that, since common air is
necessary to vegetable as well as to animal life, both plants
and animals had affected it in the same manner, and I own I
had that expectation when I first put a sprig of mint in a
glass jar standing inverted in a vessel of water; but when it
had continued growing there for some months, I found that
the air would neither extinguish a candle nor was it at all
inconvenient to a mouse which I put into it. * * * Finding
that candles burn very well in air in which plants had grown
a long time, and having had some reason to think that there
was something attending vegetation which restored air that
had been injured by respiration, I thought it was possible that
the same process might also restore the air that had been
injured by the burning of candles. Accordingly, on August
17, 1771, I put a sprig of mint into a quantity of air, in which
a wax candle had burned out, and found that on August 27
another candle burned perfectly well in it. This experiment
I repeated, without the least variation in the event, not less
than eight or ten times in the remainder of the summer.”
Had Priestley had the good fortune to have set his jar con-
taining green sprigs into direct sunlight, he would have made
an additional discovery of almost equal importance. But the
world did not wait long till Ingenhousz went over the ground
4 The American Naturalist. (January,
and discovered (1779) that light was an essential factor in
restoring air, and that by the aid of sunlight he could per-
form in a few hours the experiments which took Priestley five
or six days.
In 1800 Senebier added the discovery that plants obtain all —
their carbon from carbon dioxide, but fell into the error of
supposing that part, at least, of this gas was taken up by the
plant through its roots, an error that has proved extremely
tenacious, existing in our text books to the present day,
although repeatedly and fully disproven. :
. e have now arrived at the time of De Saussure (1804),
who, with his superior chemical knowledge, placed the whole
subject in excellent shape. He distinguished between carbon
assimilation and true respiration. He dealt with the subject :
quantitatively, and showed that there was a definite relation |
between the carbon dioxide taken up by the plant and the
oxygen evolved by the action of light.. He clearly pointed |
out that the presence of oxygen was as essential to the growth 2
of plants as to animals, the most active parts, such as green l
leaves, opening flowers, ete., requiring the most, and that this
requirement had no relation to the presence of light. .
De Saussure also pointed out that while plants receive their |
supply of carbon dioxide for assimilation, and oxygen for res-
piration directly from the atmosphere, yet the nitrogen, which _
is an essential constituent of their organization and by far the
most abundant gas in the atmosphere is not utilized by plants |
in the gaseous form.
Having now established that plants contain gases, that these
gases are the same as those of the atmosphere surrounding the ©
plant, that oxygen and carbon dioxide are made use of in _
their gaseous forms in the life processes of plants, while nitro- `
gen as a gas is not actively connected with the life of plants, —
and having established these facts with a wealth of accurate —
experiment and logical deduction that permitted no doubt of
the truth, it was left to De Saussure’s successors to elaborate
the structure which he had so ably built, without being called
upon to again readjust the foundations.
1893.] The Gases in Living Plants. 5
It was over thirty years before a work of importance in this
line again appeared. In 1837 Dutrochet published his ana-
tomical and physiological memoirs, in which he carefully
studied the structure of vegetable organs as well as their
functions. He was the first to rightly point out the relations
of the cavities in plants to the movement of gases in respira-
tion, that only cells with chlorophyll are able to decompose
carbon dioxide, and to distinguish sharply between respiration
and assimilation. But although he recognized the essentially
different character of the two processes, respiration and assim-
ilation, yet he used the erroneous and absurd nomenclature of
the time, and called them nocturnal and diurnal respiration.
The weight of his example did much to fix the terms in popu-
lar usage, where they still persist, in spite of the protests of
every ableinvestigator and writer upon the subject since; and
even though the matter was set right by Garreau in 1851, who
did admirable work upon plant respiration.
In 1865 appeared the handbook of experimental physiology
of plants by Julius Sachs, the founder of the modern school
of vegetable physiologists. The work was comprehensive,
well balanced, and replete with clear ideas of the theoretical
bearing and logical association of the facts.
The author’s laboratory at Wiirzburg, where he shortly
became established, has been the school from which most of
the great plant physiologists of the present have received
instruction and from which all have drawn inspiration. It is
sad that its doors should now be darkened by the mists that
have gathered over the intellect of its honored director.
In the handbook of Sachs a chapter is devoted to each of `
the three branches of the subject; aeration, or the movement
of gases in the plant; respiration, or the action of atmos-
pheric oxygen; and the effect of light upon vegetation, chiefly
in assimilation. Each of the os is treated in a clear and
masterly manner.
NATIONALITY OF DiscOVERERS AND WRITERS.
Having traced the growth of knowledge regarding the vital
relations of gases to plants up to the time when it was possible
6 The American Naturalist. [January,
to present the subject in a reasonably complete and well bal-
anced manner, it will be more satisfactory to drop the chrono-
logical method of treatment, and to outline the salient features
of the subject as they are understood at the present time.
Before doing so, however, it will not be unprofitable to glance
for a few moments at the parts which the several nations have
played in this growth of knowledge, and at the reciprocal
influence which has been exerted upon the teachers of those
countries.
Science in its highest aspects has always been, as at the
present time, the property of the whole world, knowing no
political restraint or nationality. The barrier of language,
however, has had much to do with retarding the diffusion of
knowledge from the original sources of discovery into the text
books, which serve as the means of enlightenment for the
mass of learners.
The great discoverers and thinkers in our present subject,
since the days of Malpighi, an Italian, have been either
French, German or English. The Germans, before 1865,
made no discoveries of commanding importance, and even
their text books barely gave a true account of the subject as
known at the time. Link, in 1807, ignored the all-important
discoveries of Senebier and De Saussure, the more readily,
doubtless, because they were Frenchmen. ‘Twenty-five years
afterward De Candolle’s general treatise was translated from
` the French and became one of the most influential text books
in Germany. $
The chief activity among Englishmen occurred before 1800,
and brought forward the names of Hales, Priestley and Ingen-
housz. The advanced work was taken up by Frenchmen after
1800, among whom Senebier, De Saussure, Dutrochet and
Boussingault are the most conspicuous investi gators.
It is chiefly the French botanists, particularly De Candolle ` :
and Dutrochet, who have had the most potent and lasting
influence upon the popular conceptions of the English regard-
ing vegetable physiology. To them we can also trace a num- :
ber of errors and omissions which figure in our school text
books at the present time. De Candolle was the author of the
ee ae eta ES N S ORE
Sa AEN, EE E E E A EN E oes
i VESES gg E CDAS
ra Stee ae et ee
1893.] The Gases in Living Plants. 7
imaginary “spongioles” upon the root tips, which still have a
sort of backwoods existence in the minds of some persons,
although practically eradicated from the text books. He sub-
scribed to the dual respiration of plants by which they gave
off oxygen in daylight and carbon dioxide in-darkness, which
is still taught by certain American text books. In other
American text books, which are still standard, a reaction is
shown by the suppression of any suitable account of respira-
tion proper, this important subject being referred to only inci-
dentally in a line or two in connection with a short account
of the use plants make of stored food material. Thus, in the
latest revised edition of Gray’s Lessons in Botany, as in the
preceding edition, barely three lines are devoted to respiration,
while two pages are given to assimilation. This work also
teaches the incorrect doctrine’ that carbon dioxide beside
reaching the plant through the surface of the leaves, “is
absorbed by the roots of plants, either as dissolved in the
water they imbibe, or in the form of gas in the interstices of
the soil.” In Bessey’s Botany, first issued in 1880, respiration
is treated in essentially the same brief manner, and it is
curious to note that the unusually complete index to the work
does not contain the words gas, breathing, or respiration.
The modern phase of plant physiology may be said to have
been introduced to English speaking students by the transla-
tion of Sach’s text book in 1875, and reinforced by the appear-
ance of Goodale’s work in 1885, on this side of the Atlantic,
and of Vines’ work the following yearin England. In these
works the balance between respiration, assimilation and the
physical movement of gases is fairly well maintained.
Another work in English, less pretentious, but equally accu-
rate and discriminating with the last mentioned, and antedat-
ing them, should be spoken of here, that of Johnson’s How
Crops Feed, published in 1870. The work was deservedly
popular, and is still a source of exact information.
5In the discussion which followed the reading of this paper, Prof. Geo. L. Goodale
gave Dr. Gray’s reasons for retaining his early views. It was Dr. Gray’s belief that
his statement would prove, upon more extended investigations, to be essentially cor-
rect. Prof. W. H. Brewer spoke in further support of the conservative views of
Dr. Gray.
(To be continued.)
Ps
8 The American Naturalist. (January,
CERTAIN SHELL HEAPS OF THE ST. JOHN'S RIVER, ©
FLORIDA, HITHERTO UNEXPLORED. A
By CLARENCE B. Moore.
_ (Continued from November Number, 1892.)
(Second Paper.)
SHELL HEAP THREE MILES NORTH OF PALATKA.
This shell heap on the west bank of the St. John’s has been |
largely washed away by the river, and in addition great quan-
tities of its shells have been “lightered” to Palatka for use
upon the streets. It was visited by Wyman and by Le Baron?
who probably made no excavations, or at all events, did not
put them on record. Implements found by them both at this
point can be seen at the Peabody Museum, Cambridge. |
This shell heap is peculiarly rich in relics of stone and of
ment; one flat fragment of bone lined along the entire length,
W. W. Calkins (Proceedings of the Davenport Academy of Natural Sciences,
Vol. 2, pages 226-227) explored a river mound “north of Palatka,” which may be
dentica with the one under consideration, -
' Smithsonian Report, 1882, p. 771 et seq.
1393.] Shell Heaps of Florida. 9
with a hole in the center extending almost through; a small
piece of greenstone cut into the form of a pyramid, and six
arrowheads, one of quite unusual pattern for the St. John’s
River, though found in some of the Western States‘ (Fig. 1).
Fic. ]. Size, +.
Captain Rossignol, formerly in charge of lighters carrying
shell to Palatka, presented the writer with a collection of
Fic. 2. Size, t+.
implements found in situ, at this place, by him, comprising a
number of rude arrow-heads and an implement of chert very
‘Charles C. Jones, Jr., in “ Antiquities of the Southern Indians,” describes imple-
ments of this character as being broken arrow-heads turned into scrapers. Fig. II,
Plate xiv, represents a specimen from Georgia.
10 The American Naturalist. [Jonna
roughly wrought on one side, the other being left flat and
smooth, recalling the implements of the Moustier Cavern,
Dordogne, France (Fig. 2). This shell heap is probably one of
the earlier class. A careful search along the entire section of
pottery, and none was met with in digging. Two pieces upon
the surface were probably from later Indians. Investigations
at this spot, however, were not based upon enough excavations
to give a final judgment upon the subject.
._TWO SHELL HEAPS ON SALT RUN.
Salt Run makes into Lake George about two miles south-
west of where the St. John’s leaves the lake. On the right
bank, going up, at a distance of about half a mile from the
mouth, is a shell deposit some two hunderd yards in length —
and one hundred yards in breadth, with a height of from four —
to five feet on the water’s edge, increasing to a maximum of —
ten feet somewhat beyond the middle toward the land. On
the same side of Salt Run, about half a mile farther up, is a-
deposit of shell presenting no irregularities of surface, though
varying in depth at different points, owing to unevenness of
ground upon which the deposit was made.
EXCAVATION I.
53x5x3% feet deep; after surface loam no pottery was met
with. Fragment of bone awl at a depth of 34 feet. About |
three feet down was found, within half a foot of the bottom of
Fic. 3. Size, }.
1893.] Shell Heaps of Florida. 11
the shell deposit, a lance-head of graceful pattern, perfect in
every respect; the only lance-head, as far as the writer has
been able to learn, ever found at a considerable depth from
the surface in any of the shell heaps of the St. John’s (Fig. 3).
Other excavations yielded nothing of marked interest.
HITCHEN’S CREEK.
At the point where the St. John’s River enters Lake George
is Volusia Bar. About half a mile south, Hitchen’s Creek
joins the St. John’s on the east side of the river. A short dis-
tance above, on the left hand side, going up the creek, are
shell heaps and fields under cultivation; in all, about seven
acres. A number of excavations yielded the usual bones of
edible animals, and showed traces of numerous fire-places at
varying depths. In the rear of the dwelling the shell deposit,
considerably higher than elsewhere, is closely packed, the
shells being crushed to a marked extent and having a large
admixture of sandy loam—a “kjökkenmödding.” In this
deposit, below two feet, no pottery was met with, and the Pal-
udinæ were of small size, in comparison to those of some of
the shell heaps. Scattered on the surface were Paludinæ
georgianæ of large size, mingled with a Paludina previously
unknown, Paludina georgiana, variety altior, Pilsbry The
portion of the shell deposit, toward the swamp, is composed
of unbroken shells, mostly of the two varieties of Paludina,
of unusually large size and unmixed with sand or loam.
Plain pottery is found in great abundance throughout. Water
is reached at a depth of two feet. This deposit was probably
made by the aborigines living upon the shell ridge adjacent,
after the ridge had attained considerable size, since but few of
the new variety of Paludina were found in the ridge below
the surface and at comparatively little depth.
SWAMP SHELL RIDGE NEAR MORRISON'S CREEK.
About three miles south of Volusia Bar, Morrison’s Creek, a
“ cut-off,’ divided from the St. John’s by an island, enters the
’The Nautilus, April, 1892, p. 142, et seq.
12 The American Naturalist. (January,
river. Below this point, in the swamp, entirely surrounded
by water when the river is high, is a ridge of shell running
north and south, 350 feet in length, with a maximum breadth
of 180 feet. The southern extremity, the lowest portion of
the ridge, is from 4} to 6 feet in height; while the northern
end attains a maximum elevation of 11 feet, 10 inches, above
the level of the swamp when dry. Two excavations, 8x5 feet,
4 in.x9} feet deep and 7}x4x6 feet deep, were made. Two
or three bits of rude, plain pottery were met with, but none at
a greater depth than two feet from the surface. Just below 4
the surface a human humerus was found, and a human verte-
bra at adepth of one foot. A fire-place was at the same depth
but at a distance from the vertebra, which showed no marks of
fire. At a depth of four feet was found a triangular imple-
ment of shell; while 4 feet, 8 inches, down, immediately upon
a fire-place, were two rude arrow-heads, one with ashes upon
it. Animal bones, disconnected, mainly of the alligator, the
turtle and the deer, were encountered throughout.
MT. TAYLOR.
This great swamp shell ridge,’ the highest fresh-water shell
deposit on the St. John’s River, lies on the east bank, 200
yards (paced) from the water’s edge. It is about one mile
south of Volusia, and in dry seasons can be reached from the
river by wading through the swamp; though access from dry
land in the rear, about forty yards distant, is advisable.
Under any circumstances, the services of a guide are a neces-
sity. This shell heap is not referred to by Le Baron, nor is it |
particularly mentioned by Wyman, who could not have ©
failed to describe so remarkable a heap, had it been accorded
a visit. On page 44 of his memoir, Fresh Water Shell a
Mounds of the St. John’s River, Florida, “two mounds, right
bank, between Lake Dexter and Volusia,” are included in the —
6The thanks of the writer are due to Mr. William Edgar Bird, of Brooklyn, for
much information and valuable assistance, and for most cordial permission to prose-
cute investigations on every portion of his 5000 acres lying between Lake Dexter —
and Volusia; including the great shell heaps and sand mound of Bluffton, in addition | 2
to Mt. Taylor and other shell heaps in the swamp.
1893.] ° Shell Heaps of Florida. 13
list of shell heaps. This somewhat indefinite description
would seem to indicate that his knowledge of the existence
and location of Mt. Taylor was based upon information
derived from others and not personally verified.
Mt. Taylor, standing alone in the swamp, which ¿at high
water is covered to a depth of 13 feet. rises abruptly on every
side, the ascent of one portion being at an angle of 40°. The
maximum height of the ridge is 27 feet, 2 inches; its length
at base 500 feet, with a maximum breadth of base of 175 feet.
An almost level plateau on the summit has a length of 266
feet, with a maximum breadth of 80 feet. The mound is
overgrown with palmettoes, palmetto scrub, live oaks and
cedars. It lies longitudinally east and west, and is composed
almost exclusively of Paludine of a smaller size than those
of many of the later shell heaps. With the exception of a
few fragments on the surface, no pottery was found in any por-
tion of the heap, while implements of any description were of
infrequent occurrence in the various excavations. As a rule,
it may be said, the older the shell heap, the fewer relics are met
with, though weapons of stone exist at all depths, even in
mounds which contain no pottery, and in others below the
level at which fragments of pottery are found.
(1o be continued)
14 The American Naturalist. [January,
LEGENDS OF THE SUMIRO-ACCADIANS OF
CHALDEA.
By ALICE BODINGTON.
In THE American Naturatist for August, 1892, Mr. Wilson —
puts in a strong plea for the study of prehistoric anthropology,
nor can the claims of this science be overrated. But of equal —
interest in its own line is the study of that earliest civilization a
of Western Asia, which a few years ago was itself prehistoric, —
and which has only emerged into the light of day since the
deciphering of the Cuneiform inscriptions of Assyria and —
Chaldea. | a
Some 5000 years B. C., wandering Turanian tribes! settled —
in the fertile alluvial plains at the mouths of the great rivers,
Tigris and Euphrates, round the head of the Persian Gulf.
Materials for building, it might be thought, did not exist, save |
for the giant reeds, fourteen to fifteen feet high, with which
the Arabs of that marshy region still construct their huts.
But the Sumiro-Accadians, as these Turanian tribes were |
named, had the faculty possessed by their relations,
Chinese, of taking the first steps in inventions. Out of the
mud and clay of their new home they made bricks, at first-
mere cakes of sund-dried clay; then these cakes were found
to gain consistency by mixture with finely chopped straw;
finally the clay was kiln-dried and gained a hardness ani
consistency equal to the best bricks produced now. The kiln-
dried bricks were highly valued and were stamped with the
name and titles of the king for whose palaces and templ
they were to be used. Some bearing the name and title ¢
-Gudéa, the patesi or priest-king of Sirgulla, have inscriptions 0
Chaldea. Story of the Natives. Z. A. Ragozin.
_ I must disclaim all responsibility for the spelling of proper names, since
every oct I have consulted spells the names differently, and no
standard to have been arrived at. For nase se the spelling is some-
times Shnmiro-Akkadian, sometimes Sumiro-Accadian
B
1393.] Legends of the Sumiro-Accadians of Chaldea. — 15
a highly archeic character. A statue of unique interest was
found at Sirgulla;* the head is strikingly Turanian in form
and feature and bears a turbaned cap such as may still be seen
in Mongolia. No type can be more strikingly unlike that of
the Semitic Assyrians who were to be in later times the rulers
of Chaldea. This statue, and the bricks with their archeic
inscriptions found with it, are considered to be as old as
between 4000 and 3000 B. ©. A successor to this oldest of
known monarchs was Ur-ea, king of Ur, whose date can be
approximately arrived at,‘ and whose reign was over before
the Elamite Conquest of Chaldea ; when Chedorlaomer (Kha-
dar-Lagamar), of Genesis; chapter xiv, marched an army
across the desert to attack the rich and populous valleys of
Jordan, and carried off Lot, the brother of Abraham, among
his captives.
In the materials for holding their bricks together there was
also progressive improvement; in the oldest buildings discov-
ered, a sticky red clay or loam was used; then bitumen was
substituted, which, being applied hot, adheres so strongly to
the bricks that pieces of these are broken off when an attempt
is made to take a fragment of the cement. Finally, in the
latest Babylonian period, a beautiful white cement made of
calcareous earth was used, which has never been surpassed
for lightness and strength.
The whole country of Chaldea was absolutely flat; no ves-
tige of natural hillock occurred throughout its whole extent.
But the Accadians, whose very name shows their origin as
mountaineers, were determined to raise their most impor-
tant buildings above the inundations, and the wild beasts and
- noxious insects of the marshy plains. They erected artificial
mounds of a size almost incredible. The great mound of
Koyun-jik, which represents the palaces of Nineveh’ itself,
covers an area of one hundred acres, and reaches an elevation
of ninety-five feet at its highest point. To “heap up such a
3Modern Tell-Loh.
*Chaldea, p. 214-19.
ough an oo city, Nineveh was built on the Chaldean plan, on a
tell” or moun
r
16 The American Naturalist. [January,
pile of brick and earth would require the united exertions of 2
10,000 men for twelve years.” Then only could the construc-
tion of the palaces begin! The mound of Nebbi-Yuma,
which has not yet been excavated covers an area of forty acres
and is loftier and steeper than its neighbor. The platform of ©
the principal mound of Mugheir (the “ Ur of the Chaldees y
from which Abraham went forth) is faced with a wall ten feet
thick, of red, kiln-dried bricks cemented with bitumen.
The sub-structure of these mounds wes made up of rough
bricks and rubbish, hence the inherent weakness of the whole
structure. The heavy semi-tropical rains falling for weeks at
a time soaked through the casing of fine bricks, and the foun-
dation became a mass of yielding mud. The mighty palaces
and temples upon which the Assyrian and Chaldean kings
lavished all the resources of wealth, all the treasures of art,
sank into sand-choked, shapeless heaps. But the treacherous
clay could preserve, hidden from the prowling Arabs who ~
roam over this land of once mighty empires, priceless treas-
ures of art and literature. Exquisite alabaster slabs, richly
engraved, beautiful enamelled tiles forming colored friezes;
the great human-headed bulls whose very discovery made the
name of Layard famous; the life-like groups of lions and
lionesses ; and incomparably more precious than all, the royal
libraries formed by the great kings, have been preserved for. —
centuries beneath these unsightly mounds. For the one avail- :
able substance, clay, formed the almost imperishable material
of which the Chaldean and Assyrian “ books” were made.
In the great mound of Koyun-jik (Nineveh) Layard found
the remains of two sumptuous palaces, the residences of Sen-
nacherib and of his grandson Asshurbanipal, who lived some: —
650 years B. C., two of the mightiest sovereigns and conquer-
ors of the Eastern World. In Asshurbanipal’s palace the —
explorer found two small chambers, containing a layer, more
than a foot in height, of baked clay tablets, covered on both
sides with cuneiform writings. Some were still entire, others
ae ee a Se eS
yokes
teks
whe Be
in fragments. Layard filled many cases with the tablets, 2
broken and unbroken; they were sent to England, and lay _
Five Monarchies. Rawlinson, Vol. 1, pp. 317-18.
t
1893.] Legends of the Sumiro-Accadians of Chaldea. 17
for years in the British Museum untouched and unnoticed.
George Smith, a young archeologist whose devotion to science
and untiring industry and patience enabled him to undertake
—and to succeed in—an apparently impossible task, deter-
mined not only to arrange and engrave the cuneiform texts '
on the tablets, but to read them, and this he succeeded in
doing. The result was something astonishing. A series of
twelve tablets was brought to light containing an epic poem
of the highest antiquity and interest, the one alluded to fur-
ther on, containing the earliest versions of the great Sun,
Moon and Earth myths, of the Deluge, of Bel and the Dragon
and of the Creation of the world. Fragments, of course, were
missing, and to seek these George Smith was sent (by the gener-
osity of the owners of The London Daily Telegraph) to search
the Archive Chambers at Koyun-jik, and by inconceivable
good fortune, found many of the missing pieces. On his sec-
ond visit to Chaldea he fell a victim to plague. His last legi-
ble words were worthy of a martyr to science. “Not so well.
If doctor present I should recover, but he has not come; if
fatal, farewell to . . . My work has been entirely for the
science I study. ‘There isa large field for study in my collec-
tion. I intended to work it out, but desire now that my
antiquities and notes may be thrown open to all students. I
have done my duty thoroughly. I do not fear the change,
but desire to live for my family.”
Besides the tablets containing the epic poem, two hundred
tablets divided into three books were found at Nineveh, fifty
of which have been deciphered. The contents of these also
are supremely interesting; one book, the oldest, reveals the
Shamanitie stage of the Sumiro-Accadian religion; a stage in
which many Turanian tribes still remain. It treats of “evil
spirits” with which earth, sky and the “abyss” under the
earth were conceived to be filled; of sorcerers who could
employ the power of the evil spirits for the destruction of
mankind, and of magicians who understood incantations and
spells capable of driving away these malignant powers,
answering to the “ black” and “white” magic of the Middle
"La Magie et la Divination chez les Chaldeérs. François Lenormant.
2
18 The American Naturalist. [January,
Ages. The second book treats of diseases, for which no cure
was known but exorcisms, since diseases were conceived to be
personal demons. Even so late as three or four hundred years
B. C., Greek travelers visiting Babylon beheld sick persons
brought out into the streets, where any passer by could enquire
as to their malady and suggest a remedy! Even this strange
plan was not resorted to till all known forms of incantation
had been gone through and proved vain. The third book
shows a great advance from this religion of pure terror.
Beneficent spirits, gods in fact, were appealed to, especially
Una, the Heaven-god; Ea, the great deity of the Earth and
Waters; Im, the Storm Wind; Ud, the Sun, and Gibil, Fire.
Ea, above all, was beloved by the Sumiro-Accadians for his
goodness and trusted for his wisdom. His very name was a
terror to evil spirits. But beneficent as he was, Ea was con-
sidered too great a deity to be lightly invoked, and in his son
Meridug, they found a spirit whose sole office was to mediate
between his father and suffering mankind. A whole tablet is
devoted to a description of one such intercession, where the
“ Disease of the Head (insanity) has issued from the Abyss,
from the dwelling of the Lord of the Abyss,” and has attacked
a human being. Then “Meridug has looked on his misery:
He has entered the abode of his father, Ea, and has spoken
unto him: ‘My father, the Disease of the Head has issued
from the Abyss. What he must do against it the man knows
not. How shall he find healing?” Ea replies, “ My son, how
dost thou not know? What should I teach thee? What I-
know, thou also knowest. But come hither, my son Meridug
.” Here follow directions for the cure of the sufferer
that the “ Disease of the Head may vanish like a phantom of
the night.”
The conception of conscience was also carried to a high
degree among the Sumiro-Accadians. With such insistence _
and authority did it speak that it was believed to be the voice
of an indwelling guardian spirit. Some most beautiful —
prayers took their origin from this belief; they have been —
called the Penitential Psalms, from their striking likeness to
those psalms in which King David confesses his iniquities and
1893.] Legends of the Sumiro-Accadians of Chaldea. 19
humbles himself before the Lord. I have space but for a few
verses of the Sumiro-Accadian psalm called “The complaint
of the repentant heart.”
“O my God, my transgressions are very great; very great
are my sins. I transgress and know it not. I wander in
wrong paths and know it not. The Lord in the wrath of His
heart has overwhelmed me with confusion. I lie on the
ground and none reaches a hand to me. I cry out and there
is none hears me. . . . My God, who knowest the unknown
be merciful. . . . How long, O my God? . . . Lord
Thou wilt not repulse Thy servant. In the midst of the
stormy waters, come to my help, take me by the hand.”
Since the key to the cuneiform inscriptions has been discov-
ered, it has been evident that many legends of Genesis are
varients of Sumiro-Accadian originals, and that from this
source too was drawn the Jewish belief in magic, witchcraft,
dreams, supernatural serpents, sacred trees, etc.; whilst the
pure Monotheism of later times was fighting hard to establish
itself in the hearts and minds of a people, who came from a
cradleland of many gods. For in those palmy days of the
Yellow Race, when it was at the head of human progress, the
Semites are seen as nomad tribes dwelling amongst the Acca-
dians, and in one most noted instance wandering from Ur of
the Chaldees, till they finally reached Egypt and the Nile. And
the legends, the superstitions, the forms of prayer of Accad
are faithfully reflected in the earliest traditions of Israel.
(To be continued.)
20 The American Naturalist. (January,
THE FLIGHT OF BIRDS.
By I. LANCASTER.
I have been asked what effect the application of soaring
methods has upon active wing flight. If soaring goes on 80
easily where gravity is the motive power, why do not all birds
soar? o
The soaring activity is not understood when such a question
is put, obscurity arising, doubtless, from misconception.
The usual statement of soaring held by everybody, and
especially by mechanical authority, is diagrammed in 1,
where g represents weight, w, horizontal air resistance, and p,
Pate Sree E
én A? (1)
normal pressure. When a horizontal force, f, is applied, of
sufficient magnitude to cause p to equal both g and w, the —
plane moves on the horizontal path of soaring flight. Soar-
ing, or indeed any sort of bird flight, would be impossible —
thus stated, which may be called the formula method. |
The soaring statement presented by nature, is seen in 2, 8
where the horizontal force, f, and the horizontal resistance, W, a
are stricken out. Vertical g is replaced by gn and gp, while p- 4
remains the same in direction but not in magnitude. The —
pressure plane of air throws g out of vertical, making gn and —
gp out of it; p is equal to gn; fis equal to gp, and the plane ‘
soars; f pushes gp up as fast as gn pushes p down, producing —
a horizontal resultant. a
Compare both statements with 3. In either case a prism of
air as wide as the distance across the extended wings of the —
|
y SW
|
|
|
|
|
1893.] The Flight of Birds. 21
bird, as thick as the perpendicular distance between the front
and rear edges of the wings, and say fifty feet long, is driven
bie a FOP
tor Np A
out of the way in one second of time. This is done, as above
stated, in both cases, but the way of doing it is the vital -
matter. It is driven out by gnand not by f. Suppose f should
stop acting. Motion of bird would then be downward on the
normal line, in which motion gn would do the same work in
one second as before. Motion both ways goes on simultan-
eously, neither having the slightest effect upon the other, for
the reason that they are 90° apart, and rectangular forces do
not affect each other, the bird moving under each force as if.
the other were not acting. The wav in which the air is driven
out is determined by f, but gn does all the work. The law of
meer ee eee ere
om>
fluid reactions throws all the air resistance to the bird’s motion
around, normal to the plane of its wings; gn then forces it
down, while f keeps the bird level.
There is yet a very important point to be understood. In
methods sanctioned by mechanical authority weight and air
resistance are added in pressure. Both are assumed to be
resistance to the soaring force. They should be subtracted,
22 The American Naturalist. [January,
not added. pis a reaction against gn and hot a force equalto
gn, acting with it against the soaring force. Atmosphere
resistance to the bird must be overcome, but weight overcomes
it, and is itself used up in the exertion.
I once saw a parody on Jack, the Giant Killer. Jack was ~
set upon simultaneously by two giants, either of which could i
have demolished him at once. He adroitly set them to fight- __
ing each other and then cut off their heads. Nature met with
much the same problem. She desired a soaring bird. Two
antagonists confronted her, air and weight. She so fashioned
a bird as to take advantage of the law of fluid pressure, which
set weight upon air resistance, in which contest they were both :
destroyed. She outdid Hercules in details of destruction. l
Pressure first cut weight into two unequal parts, then fell upon :
the greater and transformed it into a stream of escaping air
condensation, while the smaller still offered resistance. Then |
from these condensations, equal to the total normal part, she 3
obtained enough force through wing details to destroy the ¥
parallel part, and still have a large surplus. And all this was
accomplished by a bird’s wing. It seems a pity that such a
magnificent piece of work should belong only to fishhawks,
carrion crows, and the like.
Further, a single matter must be noticed. By referring to
3, it will be seen that there is a region behind and above the
lower surface of the bird, and beneath the front edge, of tri-
angular shape, that is not affected by the air. If the bird had
but one motion on the upward slant, any thickness of parts,
either of wing or body, would resist. But normal motion
confines air collision to the under side, leaving a confused
mass of eddies and reacting currents at the upper side. This
region of eddies may be filled with solid materials and still
not destroy the effect of a mathematical plane devoid of
thickness. The front edge must be sharp but the rear edge
may be overreached, as in 4, without further resistance. Su
shape will move farther in the same time unit normally than a
flat shape, while parallel motion is not changed in the slightest
degree. The least projection above a b increases resistance.
1893.] The Flight of Birds. 23
I have called this region the “neutral zone,” for want of a
better name. The bodies of all birds are almost entirely con-
fined to this zone.
The original question is now in order. Why do not all
birds soar? For many reasons, among which are the follow-
ing:
1. A bird may be of such shape that to throw it over to an
inclination that gives a parallel factor small enough to be
neutralized by the forward thrust would get part of the ani-
mal out of the neutral zone. The gray pelican of the Gulf
coasts is a case in point; especially when its gullet is full of
fish. It must flap to get thrust for its large parallel factor. If
its wings were one foot longer on each side, it could doubtless
soar continuously, even if its body did encroach on resisting
air to some extent.
2. Weight may be too great for surface. In this case con-
densations escape too readily to be utilized. A wild turkey,
prairie-chicken, or pheasant, are examples.
3. Weight may be too little for surface. Here condensation
is too weak to give thrust. A species of sea-gull found on the
lower Florida peninsula is a good example. In May and
June, when food is scarce, they flap continually. In Novem-
ber and December, when food is plenty, they rarely move a
wing. They will put into an empty craw their weight in food. |
4. Life habits may prevent suitable feather construction. I
presume this reason may apply to many small birds.
In the case of bats, the small factor may be entirely neu-
tralized by flapping, as there seems to be at least no surface
provision for utilizing the escaping condensations.
Flying squirrels do not sustain themselves.
24 The American Naturalist. [J anuary,
Ba
But in no case of bird flight that I am acquainted with, can
it be said that the normal factor is opposed by flapping. That
is cancelled by its own work on air. Flapping goes on for the
sole purpose of producing parallel motion by either increasing
the energy of the condensations, or by a backward push
against the air.
It must be borne in mind that the nearer the bird’s wings
approach horizontal, the less will be the obstructive gravity
factor, but on the other hand, the more contracted will be the
neutral zone. The moment this zone is encroached upon,
more is lost in resistance of air, than is gained by lessening
the small factor. >
|
Sie a
In experimenting, I never pay the least attention to what I
have called the “soaring force,” meaning thereby the force —
required to push the plane to the resultant, after the small
factor is neutralized. I have never used scales delicate
enough to measure this force. The moment the small factor
is out of the way, the plane runs to the front to the limit of ,
its freedom. :
There is the narrowest possible margin between active and |
fixed wing flight. The only group of white pelicans I ever —
met with, eight in number, moved through the air in alternate
flapping and gliding motion. Once only, I found them facing
a southwester on fixed wings. They rested in the gale as
firmly as if fastened to a rigid support. I had been observing
them daily for five months and was rewarded by this very |
unusual exhibition. I shot one of them and found its gullet mM |
and intestines full of fish, and it could only spread one square — |
foot of surface to each five pounds of weight, the greatest con-
trast of surface to weight I ever found. :
The entire subject of bird flight has been persistently mis-
conceived. It must be recast in toto to rescue it from the
mass of delusion that involves it. To speak of erroneous
details is labor lost. It is all erroneous. There is no stress in
the entire activity, either in direction or magnitude, where
stresses are supposed to be. When it is seen that from eleven-
twelfths to seventeen-eighteenths, approximately, of total
weight is employed in overcoming total air resistance to the
1893.] The Flight of Birds. 25
bird’s motion, it is easily conceivable that the animal may
overcome the remaining resistance without the necessity of
estimating the muscular exertion of a creature weighing eight
ounces in terms of horse power.
26 The American Naturalist.
EDITORIALS.
EDITORS, E. D. COPE, AND J. 8. KINGSLEY.
—Ar the October meeting of the American Humane Association, —
held in Philadelphia, a resolution was passed of a very dangerous —
nature. It urged upon the Legislature of every State in the Union —
the enactment of laws which shall prohibit, under severe penalty, the a
repetition of painful experiments upon animals for the purpose fa
teaching or demonstrating well-known and accepted facts. The danger
lies not in the intent, but in the fact that incompetent persons will feel —
it their duty to say that this experiment is painful, that another is 4
unnecessary ; and further, it strikes a deadly blow at all future increase —
of knowledge. An investigator in physiology can only be trained
by the laboratory method. He cannot read the accounts of previous =
work and, with no further preparation, proceed at once to the solution
of new problems. He must, rather, test his powers of experimenta-
tion by this very repetition which the proposed law prohibits. He —
must demonstrate for himself “well-known and accepted facts,” and
until he is able to bring his results into full accord with those facts
he is incompetent to enter untrodden fields where he is without checks —
upon the accuracy of his results.
That the proposed legislation is not so vicious as some, which, for- —
tunately, has been rare in the United States, is a matter for which We —
should be thankful; but, on the other hand, it would place a danger
ous tool in the hands of such fanatics and unqualified persons as com- —
monly occupy important positions in connection with the societies for
the prevention of cruelty to animals. The writer has a somewhat —
extensive acquaintance with the physiological investigators of both
Europe and America, and he knows them to be as a class humane per-
sons unwilling to inflict unnecessary suffering upon any animal, and
at the same time fully as competent to judge of the necessity of any
experiment as the persons whom the proposed legislation would put m
the position of prosecutors and judges.
— LIEUTENANT PEARY has obtained leave of absence from the Sec-
retary of the Navy for the purpose of further prosecution of Arcti¢
explorations. He proposes to establish himself at a point on the
northern coast of Greenland already visited by him as a base for —
4
al
1893.] Editorials. 27
explorations northward. He expects to travel over the ice which
covers the ocean, toward, and if circumstances permit, to the pole.
It will be a fortunate circumstance, and one conducive to the success
of the expedition, if land shall be found to the north of Greenland.
This, Lieutenant Peary suspects, may be the fact. Transportation will
be thus more easy and less dangerous, and much of interest to science
may be expected to result. The determination of the geologic and
paleontologic features of the region is of first-class importance to
world-history, and much important evidence will be contributed
toward the solution of some at present obscure problems.
—Tuer numbers of Toe AMERICAN Natura.isr for 1892 were
issued at the following dates: January, March 26; February, March
31; March, April 25; April, April 29; May, May 1; and all subse-
quent numbers on the first of the month named on the cover and
pages.
28 The American Naturalist. (January,
RECENT BOOKS AND PAMPHLETS.
ALLEN, J. A.—On Colaptes auratus and C. cafer. Ext. Proceeds. Am. Mus. Nat.
Hist., 1892. From the author q
Barley, L. H.—The Hortieulturalis’s Rule-Book, completed to the beginning of —
the year 1892. From the aut
BANKS, N.—The Spider a of the Upper Cayuga Lake Basin. Ext. Proceeds. —
Acad, Nat. Sci. Phila., 1892. From the author. - n
Baur, G.—Ein Besuch der Galapagos Inseln. München, 1892. From theauthor. —
EAN, T. H.—Description of a new species of Star-gazer (Cathetostoma albigutta) —
from the Gulf of Mexico. Ext. Proceeds. Natl. Mus., Vol. xv. From the author.
Bulletins No. 16 and 18 Agri. Exp. Station of the Rhode Island College of Agri-
culture and Mechanic Arts.
Bulletin No. 18 Iowa Agri. Exp. Station. From Herbert Osborn.
Bulletin No. 43 Mass. State Agri. Exp. Station, Aug., 1892.
CLAYPOLE, E. W.—Traces of the Ice Age in the Flora of the Cuyahoga Valley.
Tract No. 84 Western Reserve Hist. Soc. From the author.
DALL, W. H.—Tertiary Mollusks of Florida, Part 2. Trans. Wagner Inst. Science.
From the mither. ;
RET for Collecting Mollusks and other Useful Hints for the Concholo-
gist. Part G of Bull. U. S. Natl. Mus. No. 39. From the Smithsonian Institution.
Day, D. alist Resources of the United States. Washington, 1892. From
the author.
Dotto, L.—Sur le Lepidodteus suessoniensis ——Sur la Morphologie de la Colonne
vertébrale.
——Sur la Morphologie des Côtes. Exts.du Tome xxiv, Bull. Sci. de la France
et de la Belgique.
—Nouvelle Note sur le Ch lien adapté à la vie fluvia-
tile. Ext. Bull. Soc. Belge de Geol. , Paleont., et Hydrol. Bruxelles, Tome v, 1891
From the author. ;
DumBLE, E. T.—Third Annual Report of the Geological Survey of Texas for
1891. Austin, 1892. 4
——Notes on the Geology of the Valley of the Middle Rio Grande. Ext. Bull,
Geol. eer Am., Vol. iii, pp. 216-230. From the Society. a
Dyar, H. G.—Descriptions of Some Butterfly Larvæ from Yosemite. Ext. Can:
ES Oct., 1891. From the author.
Evolution in Science, Philosophy, and Art. Popular Lectures and Discourses
Before the Brooklyn Ethical Association. New York, 1891. From D. Appleton &
Co., Seen
GAER’
TNER, F.—Concerning the Differentiation of Black Pigment in the Liver, |
Spleen, and Shean from Coal-dust Deposits. Reprint Microscope, July, 1889.
From the author,
©- —Vivisection. Ext. AM. NAT., Oct., 1891. From the author.
ILL, TH.— Notes on the Tetraodontoidea. Ext. Proceeds, Natl. Mus., Vol. xi%
From the author.
1893.] Recent Books and Pamphlets. 29
GRAFF, L. von.—Bibliothek des Professors der Zoologie und Vergl. Anatomie.
Leipzig, 1892. From the author
Hasets, M.—Extrait du rapport sur l'Industrie des Sardages à l’Exposition Uni-
verselle de 1889. From th
HARLE, E.—Une Mandibule de Singe du Repaire de Hyenes de Montsaunes ,
Ext. Compte-rendu fèv., 1892. From the author.
Hay n the Breeding Habits, Eggs, slik Young of Certain Snakes,
S0 the agg og of Blood from the Eyes of Horned Toads. Exts. Proceeds.
U. S. Natl. Mus., . xv. From the Smithsonian Institution.
Hayes, C. ie Expedition Through the Yukon District Ext. Natl, Geog.
Mag., Vol. iv, pp. 117-162. From the author
HERNDON, J. H.—Plea for the Life of the Geological Survey of Texas.——
Review of the charges preferred against Prof. E. T. Dumble, State Coble: for
“ Incompetency, Plagiarism, and Maladministration in Office,” and the sham trial
thereof. Austin, 189]. From the author
Hircucock, R.—The Ancient Pit. Dwellers of Yezo, Japan. Ext. Reprint Natl.
Mus., 1808. PP- 417-427.
Descriptions of Two New Forms of Basileuterus rufiforms from Mexico.
Ext. Proceeds. Natl. Mus., Vol. xv. From the author
aye of Life. A Sym püsik on the Contig Man. Edited by Wallace
$ D. New York, 1892. From E. B. Treat, Publisher.
een D. S.—Evolution. Syllabus of Lectures, Alameda, 1892, From the
author.
KEpzik, J. H.—Speculations. Solar Heat, Gravitation, Sun Spots. Chicago, 1891.
From the author.
Kirsy, W. F.—Elementary Text-book of ee Second Edition. London
and New York, 1892. From Macmillan & Co.,
LERCH, O.—A Preliminary Report on the eae z pe North of the Vicks-
~ Shreveport and Z Railroad. From the author i
U, J.—The Geolo KaR Map of the United States the United States
Gelgie sale: oe 892.
Mason, O. T.—The oh ak or Womans Knife, of the Eskimo. Ext. Rept. Natl.
Mus. tor 1006 From the Muse
MERRILL, G. P.—Handbook K the Department of paed in the U. S. National
Museum. Ext. Rep. Natl. Mus. for 1890. From the Mus
OSBORN, HERBERT.— Lice Affecting Domestic Animals. Ea. Bull. No. 16, Iowa
Agri. Exp. Station
Report of the Cio of Fish and Fisheries Respecting the Establishment
of Fish Cultural Stations in the Rocky Mountain Region and Gulf States. Wash-
ington, 1892.
Report of the Conduct of the British Museum for 1891. London, 1892.
Ripcway, R.—The Humming-Birds. Ext. Rep. Natl. Mus., 1890, pp. 253-383.
tion.
Riley, C. V.—Directions for Collecting and cramer de Insects. Part F sid Bull.
U. S. Natl. Mus., No. 39. From the Smithsonian Insti
Rust, Dr. D. —Contributions to Canadian Sitio: Paacoatciogy, Part 4, with an
introduction by J. B. Tyrrell. Ottawa, 1892. From the Can. Geol. and Nat. Hist.
Survey.
30 The American Naturalist. [Janui
Science Teaching in the Schools. Report of the Committee of the American
Society of Naturalists, 1891. J
SuuFELDT, R. W.—The Myology of the Raven. A Guide to the Study of the
Muscular System in Birds. London and New York, 1890. From Macmillan & Co.,
Publishers.
STEJNEGER, L.—Two Additions to the Japanese Avifauna, including Description —
of a New Species. Ext. Proceeds. U. S. Natl. Mus., Vol. xv, pp. 371-373.
the Smithsonian Instituti ;
ARR, R. S.—A Hint pur Respect to the Origin of Terraces in Glaciated Regions.
Ext. Am, Jour. Science, Vol. xliv, 1892.
——Reconnaissance of the BPA Mountains. Bull. No. 3, Dept. Agri.
Insur. Statis. and Hist. Geol. Survey Texas, 1892. 2
The Relation of Secular Decay of Rocks to the Formation of Sediments. —
Ext. Am. Geol., a y, 1892. From the author. E
TOWNSEND, C. .—Notes on North lian Tachinide sens str. with
ee of ee Genera and aie Ext. Canadian Entomologist, March, —
A Parasite of the Fall Web-Worm.——A New Simulium from Southern New —
Mexico.—Description of a Sarcophaga Bred from Helix. PEPA Pryche, 1892.
——A Tachinid Parasite of Crysophanus dione. Exorista chrysophani n. sp.——
Description of a Muscid Bred from Swine Dung, with Notes on Two Muscid Gen-
era. Exts. Entomol. News, 1891. i
Notes on North American ear with Descriptions of New Genera and —
Species. pr 1892. From the autho
Wurre, I. C.—The Mannington Oil ida and the History of Its Development.
ossil Plants from the Permian Beds of Texas. Exts. Bull. Geol. Soc. Am.
Vol. iii iii, pp. 187-218. From the Society.
1893.] Recent Literature. 31
RECENT LITERATURE.
The Apodidez.'—This, one of the latest of the “ Nature Series,”
is not up to its predecessors either in accuracy of statement or sug-
gestiveness of matter. The author, starting off with the intention of
working up the comparative anatomy of this family of phyllopod crus-
tacea, has been led to regard them as all-important in phylogenetic
speculations, but, unfortunately, the good points of his volume are not
original, while the original portions cannot be praised. Thus the cen-
tral position of the Phyllopods in the Crustacean branch was recog-
nized long ago, while the comparison of the foot of the Apus with the
parapodum of a Polychzte worm was made long before Bernard entered
the field of zoology. On the other hand the special studies of Mr.
Bernard have led him to regard the differences between the annelids
and Apus as of extremely minor importance. All you have to do,
says he in effect, is to bend the anterior end of a carnivorous annelid
back upon itself to produce this portion of Apus, and the thing is
done. Resemblances are magnified and differences are minimized or
ignored, and presto! Apus is the all-important arthropod. The name
of Macmillan & Co. is so uniformly associated with only first-class
works that we were surprised to see it on the present volume.
Darwin, and After Darwin; I, The Darwinian Theory,
by Grorce Jonn Romanes.’—Romanes has devoted the best years
of his’ life to the defence of the evolutionary faith and to making
himself acquainted with, advocating, and extending Darwin’s ideas.
The present work consists of two volumes, viz., “The Darwinian
Theory,” and “ Post-Darwinian Questions.” The latter, soon to be
issued, is to treat of heredity, utility, isolation, etc., which have
become prominent since the death of Darwin. The former is a
systematic exposition of the Darwinism of Darwin. It gives a
résumé of the evidence, as it is known at present from class-
ification, Morphology, Embryology, Paleontology, and Geographical
Distribution, and includes a full discussion of the Theories of Natural
and of Sexual Selection. It is a neat volume of 460 pages, fully
1The Apodide. A morphological study, by Henry Meyners Bernard. London
and New York. Macmillan & Co., 1892.
2Chicago. The Open Court Publishing Company, 1892.
32 The American Naturalist. [January,
illustrated with new figures that largely increase its value. It is the
best single volume on the general subject that has appeared since —
Darwin’s time, and it is doubtless destined to be for years to come the —
The principle of continuity makes antecedently probable the theory
of organic evolution. The probability is strengthened by the fact that ;
a natural classification of organie beings seems with the advance of
knowledge more and more evident. The evidence from Morphology —
in the present volume is confined largely to a discussion of rudimen- —
tary structures, and especially such as are found in the human body. —
Muscles of the external ear, panniculus carnosus, feet, hands, tail, ver-
miform appendix of the cxcum, ear, hair, teeth, perforation of the i
humerus and flattening of the tibia are all treated. In this connection — |
Dr. Louis Robinson’s recent interesting observations on the grasping — :
power of the infant’s hand are reported. In discussing Embryology, —
considerable space is devoted to the phenomena of fertilization and
karyokinesis, since the author believes the great similarity in these
highly complex and specialized processes, shown throughout the animal —
and vegetable kingdoms, to be indicative of organic continuity, and
hence evidence of the highest importance. The testimony afforded by i
connecting links, which has accumulated mainly since Darwin first
published, forms an interesting section, made more interesting by good i.
figures. Geographical Distribution is mainly a summary of Wallaces
observations. E
' The evidence for and against the theory of Natural Selection is fully
and fairly given. Romanes believes this principle to be not the sole,
but the most important, factor of organic evolution. The main gen- —
eral arguments in favor of the theory are three, viz., its inherent
necessity, the facts of heredity, variation, and struggle for existence
being excepted ; the fact that among all the millions of structures and
instincts, each is developed for the benefit of its own species, and m
not a single case for the exclusive benefit of another species; also the
facts of domestication. Protective coloring, warning coloring and
Ne o aE Sepp sagas E es OS Cg EE E SE E E re Ea nm E AS ATER
similar organs in widely different groups (Mivart’s instance of the p
of the cuttlefish and of vertebrates), and the preservation of the ft
beginnings of structures (the Duke of Argyll’s “Prophetic Germs "h |
1893.] Recent Literature. 33
where the principle of correlation must play so important a part. The
electric organ in the tail of the skate is a formidable case, which our
present knowledge is not able satisfactorily to dispose of. In the last
chapter Mr. Wallace’s objections to Darwin’s theory of Sexual Selec-
tion are replied to. The relations of the Darwinian doctrine to adap-
tation and beauty in organic nature are discussed in brief, and finally
its relations to the fundamentals of religion —F. 8. Ler. »
The Diseases of Personality, by Tu. Rısor.’—The new Psy-
chology is under a great debt to Ribot for his studies of nervous dis-
eases. In this last volume he bases personality as the highest form of
psychic individuality upon the organic sense. All the bodily organs
are constantly sending into the central nervous system impulses that
give rise to sensations. These organic sensations are relatively more
prominent in the lower animals, because there they are not, as they are
higher in the scale, covered up by desires, passions, perceptions and
ideas. Everywhere, however, they form the physical basis of person-
ality. The author analyzes the organic, emotional and intellectual
conditions and disorders of personality. The discussions include the
meaning of “individual” in various forms of animal life; the person-
ality of twins and double monsters; the rôle of memory ; transforma-
tions brought about by hallucinations and by ideas; the phenomena
of the dissolution of personality in cases of progressive dementia. A
convenient, if not entirely comprehensive, classification of the diseases
mentioned is that its three categories, viz., alienation (where the
changed person is either entirely ignorant of his former self or regards
it objectively), alternation (ordinary cases of double consciousness),
and substitution (where the individual takes on a new character, yet is
conscious of his former one, as where he now believes himself a king,
although he remembers that he was formerly a poor man).
It is to be hoped that the same publishers will issue in the same neat
form the author’s works on the diseases of memory and of the will.
F. 5. Lee.
SAuthorized translation, Chicago. The Open Court Publishing Co. pp. 157.
34 The American Naturalist. [January,
General Notes.
GEOLOGY AND PALEONTOLOGY.
On the Formation of Oolite.—Dr. A. Rothpletz has proposed
a theory of the formation of odlite, which is as interesting as it is
novel. He noticed on the shores of Great Salt Lake, Utah, snow-white
and silver-gray calcareous corpuscles in great numbers. They form @
large part of the beach sand, and where they lie in the water they are
partly covered with a bluish-green alga-mass. On examination the
algoid bodies proved to be colonies of cells of the lime secreting alge,
Gleocapsa and Gleothece. By a series of experiments Dr. Rothpletz
satisfied himself that the calcareous bodies secreted by the plants and
the calcareous bodies which compose the beach sand are identical.
Pursuing his researches, the author investigated the odlites from the
strand of the Red Sea, and found that although slightly differing m.
structure, these odlites originate similarly to those of Salt Lake; that
is, from lime secreting algæ.
In studying fossil forms Dr. Rothpletz has found in a gray limestone
from the Lias of the Vilser Alps, and in the great oölite structure of
gated by Wethered, and more recently by Bleicher (May, 1892),
closely resembles that of the Red Sea odlite. In view of these
Dr. Rothpletz is inclined to believe that at least the majority of the
marine cal¢areous odlites with regular zonal and radial structures are
of plant origin; the product of microscopically small alge of very
low rank, capable of secreting lime—From Botanisches Centralbla
Translated by F. W. Cragin for the American Geologist, Nov., 1892
intended as a basis for the economic geology of that region, and 1
therefore, general rather than special and detailed. ‘The rocks of the
*Report on Geology of Northeastern Alabama and adjacent portions of Georgis
d Tennessee. Bulletin No. 4, Geological Survey of Alabama. By C. Wil
Hayes, Assistant Geologist U. S. Geol. Surv., 1892.
1893.] Geology and Paleontology. 35
area under consideration are all Paleozoic, and include representatives
of all the larger subdivisions of that system. A columnar section of
the strata exposed east of Browntown Valley gives the Cambrian rocks
an average thickness of 7550 feet; the Silurian, 5935 feet; the Devon-
ian, 180 feet; and the Carboniferous, 2175 feet. The formation names
are all new, being purely geographic and local. It is questionable if
the making of new names is necessary in regions contiguous to those
with similar formations whose names have been generally adopted.
The reason advanced, “ to avoid all remote correlations,” does not seem
sufficient to warrant such innovations.
The report is accompanied by an excellent Geological map which
shows a structure section through Northeastern Alabama.
The Mesosauria of South Africa.—Paleontologists are indebted
to Prof. H. G. Seeley for a detailed description of the Mesosauria of
South Africa, and an exact statement of the relations of Mesosaurus
with Stereosternum.
For many years the genus Mesosaurus has been evidenced by a
single fossil from Griqualand, South Africa, described by Gervais in
1865 under the name Mesosaurus tenuidens. In 1878 four specimens
from the shale at the margin of the Kimberley Diamond field were
obtained by Mr. G. H. Lee, and deposited in the British Museum. They
show that Mesosaurus was a long-tailed reptile, with hind limbs well
developed. Mr. Seeley found it impossible to refer the Kimberley speci-
mens to M. tenuidens, and described them under the name M. pleuro-
gaster. The well-developed abdominal ribs, formed of flattened
plates, give this species its most distinctive feature.
A second specimen of M. tenuidens found in the district of Albania
by David Arnold, preserved in the Cape Town Museum, shows the
ventral aspect of the anterior part of the skeleton. There are some
differences between this specimen and the type, but Mr. Seeley does
not consider them important enough to prove specific distinction.
Another specimen of the same genus, from near Burghersdorf, is in
the Albany Museum at Grahamstown. It shows the dorsal aspect of
dorsal vertebre and ribs. It indicates a new species, but there is no
character available for its definition except that of relatively stout
ribs.
In discussing the relations of Mesosaurus with Stereosternum,
Mr. Seeley refers to Stereosternum tumidum from Brazil, described
by Cope in 1886, and figures the shoulder girdle of that species
to show a pair of wide, thin, crescentic bones in advance of the
36 The American Naturalist. [January,
shoulder girdle. In regard to these bones Mr. Seeley says: “ If
the transverse expansion seen in the Paris type of Mesosaurus is the
‘same bone, its form is imperfect, but it is in the same position as the
lateral crescentic bone of Stereosternum. There is nothing in the Cape
Town Mesosaurus which corresponds in form with these bones in
Stereosternum, and the shoulder girdle in the two types seems to be
unlike, because the coracoids in the Brazilian genus met (as shown by
the thickened margin) in the median line, while in Mesosaurus there
seems to have been a squamous overlap as in Monotreme mammals,
and as the coracoid cartilages overlap in Triton and Salamandra.
This condition, so far as I am aware, is not otherwise suggested by
remains of fossil reptiles. There is also a possible resemblance to Sal-
amanders in the fact that the scapula and coracoid are not separable,
though the Cape Town Mesosaurus appears to indicate a suture. °
These African Sauromorpha closely resemble some genera from the
Trias of Europe in general form and characters of the humerus. This
leads Mr. Seeley to present, for the present, the following classification
of a small group to which he gives the name “ Mesosauria: ”
MeESOSAURIA.
General Characters.—Palate closed in the median line, teeth slen-
der, prehensile; cervical ribs with a single articulation, dorsal ribs
articulated to the anterior face of the neural arch. The shoulder-
girdle formed of scapular and clavicular arches. Humerus expanded
distally with an ent-epicondylar foramen. Digits terminating in claws.
Division I. Proganosauria.
Articular faces of centrum conically cupped, coracoid and scapula
anchylosed, a larg’: clavicle (or separate episcapule), a sacrum of four
vertebrz, a foramen in the pubis, five bones in the distal row of the
tarsus, neck short, tail long. South Africa, South America.
Division IT. Neusticosauria.
Articular faces of centrum flat, coracoid and scapula separate, clav-
icles relatively small (no separate episcapula), sacram unknown, @
notch instead of a foramen in the pubis, neck long, tail short. Europe.
This order is an important diagnostic type of its horizon, and all
additional knowledge respecting it is welcome.
Quarterly Jour. Geol. Soc., Nov., 1892.
E ee ae ee a tes s
PS -EN it Se
E EA ee SEa VNE
e T E ae ae nine eee
See Te EE EOE
1893.] Geology and Paleontology. 37
Kansas Pterodactyls.—The wealth of material in the museum
of the Kansas University affords Prof. Williston the opportunity to
compare the genera Nyctodactylus, Pterodactylus and Pteranodon,
with the following result:
“ It seems very probable that the genus Nyctodactylus has no teeth
in its jaws; it agrees in every other respect with the genus Pterodacty-
lus, so far as known. Now, in not a few species of Pterodactylus the
teeth are confined to the anterior end of the jaws, and their entire
absence, unaccompanied by other structural differences, will hardly
constitute an order, or even a family.
“ Pteranodon differs from Pterodactylus, so far as that genus is
known, in the united coracoscapule and pubes, both of which charac-
ters are found in Rhamphorhynchus.
“The sole family characters remaining then, for Pteranodon, are
absence of teeth, a supra-occipital crest, and the articulation of the
upper end of the scapula.”
Prof. Williston, therefore, proposes the following classification :
Order Pterosauria.
Family Pterodactylide; sub-families, Pteranodontine, Pterodactyl-
ine.
Family Rhamphorhynchide.
Family Ornithocheiride.—Kansas University Quarterly, July, 1892.
Geological News, General.—Prof. S. W. Williston considers the
practice of American text-books in Geology in introducing generic
names of characteristic fossils as the names of the geological horizons
whence they come as very reprehensible. Leconte’s Elements contains
a long list of such names that have long been out of use by paleon-
tologists.—Kansas Univ. Quart., July, 1892. According to T. Mel-
lard Reade, marine sands are rounded and highly polished, while
non-marine but purely glacial sands are invariably angular.— Geolog-
ical Magazine, Oct., 1892.——M. Boule calls attention to some well-
preserved remains of Elephas meridionalis found in the volcanic
terranes of Senéze (Haute-Loire). They resemble E. meridionalis of
the English Crag. This fossil confirms M. Boule’s previous statement
that while some of the volcanoes of the valley of the Allier (Coupet
and Chilhas) were active during the middle Pliocene, others, like
Senéze, are contemporary with E. meridionalis, and are therefore more
recenit.—Revue Scientifique, Nov., 1892.
Paleozoic.—A new form of the rare group Agelacrinitide has _
been found in the lower carboniferous limestone of Cumberland,
38 The American Naturalist. [January, .
England. The fossil is described and figured in the Quart. Jour.
Geol. Soc. May, 1892, under the name Lepidodiscus milleri, by
G. Sharman and E. T. Newton. Mr. H. G. Seeley describes a new
reptile from Welte Vreden, Cape Colony, Eunotosawrus africanus.
The dorsal vertebræ in form and number suggest the Chelonian type,
but the specimen affords no proof that the whole of the dorsal verte-
bre are preserved. Every character preserved differs from those of
South African fossils hitherto known, with the exception of the pubis,
which suggests that the specimen is referable to the Mesosauria in a
division distinct from the Proganosauria—Quart. Jour. Geol. Soc.,
Nov., 1892.——Eleven new species of Lower Silurian Ostracoda refer-
able to the two genera Leperditia and Schmidtella are described and
figured by E. O. Ulrich in the Amer. Geol., Nov., 1892.
Mesozoic.—Mr. Arthur Hollick calls attention to some fossil mol-
lusesfound at Tottenville and Arrochar, Long Island. Prof. Whitman
has determined them to be marine cretaceous species. These, in con-
nection with cretaceous plants found in the same locality, establishes
the cretaceous strata which have hitherto been assumed to extend
along the southern and western shores of Long Island.—Trans. New
York Acad. Sci., 1892, p. 96. A new Mosasaur, Clidastes westii, is
described by Mr. Williston. The fossil was found in the uppermost of
the Niobrara beds, and consists of a complete lower jaw, quadrate,
portions of the skull, the larger part of the vertebral column, and the
incomplete hind and fore paddle. It is estimated that the animal in
life measured seventeen and one-half feet.—Kansas Univ. Quart., July,
1892. Mr. Ubler’s observations show three structural units in the
Tuscaloosa formation instead of the one insisted upon by McGee and
Darton. His division is as follows: 1 The Potomac formation prope?
laid down on the broken border of the crystalline rocks and capped —
RIM
Y
ai
i
3
He
3
K
aE
LEd
a
a
by the variegated clay. 2 "The Albirupean formation, which includes
the series of clays, sands and cobble-stone deposits resting between the
variegated clay and the base of the Severn formation. 3 Alternate
clay sands resting upon the irregular and eroded surface of the white
1892.
Cenozoic.—The collection of fossil marsupials at Queensland
includes a fine series of mandibles of Phascolomys mitchellii, which sup” ,
_ port Mr. De Vis in making this a distinct species from P. platyrhinvs-
—Proceeds. Linn. Soc. N. S. W., 1891——Two species, Laganum
Ma
clay or sand of the preceding group.—Trans. Maryland Acad. Sci, a
1893.) Geology and Paleontology. 39
decagonale and Cassidulus florescens, have been added by Mr. Gregory
to the Australian fossil Echinoidea. The papers recently published
by Cotteau, Tate, Bittner, and Gregory on Australian Cenozoic Ech-
inoids, show that the fauna in question is Eocene and Oligocene instead
of Miocene, and that it is remarkably rich and varied in genera —
Geol. Mag., Oct., 1892.——A collection of mammalian bones from
Mongolia reported on by Lydekker are of interest since they carry the
Chinese mammalian fauna to a more northern district than has hith-
erto been known, and they indicate two Indian Siwalik species not
previously recorded from Chinese territory, viz., Hyena macrostoma
and Equus sivalensis.— Records Geol. Surv. India, Vol. xxiv.
40 The American Naturalist. (January, q
MINERALOGY AND PETROGRAPHY:'
The Origin and Classification of Igneous Rocks.—Mr.
Iddings’ has recently published at length the data upon which are
based his conclusions concerning the causes of the different structures
exhibited by the igneous rocks of Electric Peak and Sepulchre Moun-
tain and of their varied mineral composition. The main results
reached by this study have already been noticed in these pages? It
may be well again to call attention to the fact that in this region the —
different conditions attending the final consolidation of the ejected and —
of the intruded magmas affected not only their crystalline structure, 4
but also their essential mineral composition; consequently, the mole-
cules in a chemically homogenous fluid magma combine in various
ways and form quite different associations of silicate minerals, produc-
ing mineralogically different rocks. For instance, biotite is an essen-
tial constituent of even the most basic of the intrusive rocks, while in
the effusive phases it is rarely found in rocks containing less than 61%
of SiO, Again, quartz is common in the coarser grained varieties of —
the former and is absent from those of the latter. Therefore, it is more
proper to consider intrusive and effusive rocks that havea like chemical —
composition as corresponding or equivalent rocks, than those forms of
the two series that have similar mineral compositions. The classifica-
tion of igneous rocks should recognize the close dependence of struc- n
ture and mineralogical composition upon geological relations. But,
since the structure is the best exponent of these relations, structure ;
should form the basis of this classification. Though giving most 0
his attention to the general subject of the relation existing between the —
structure and the geological position of the rocks of the area described, —
the author devotes a portion of his article to illustrating the inter —
growths of hypersthene, pyroxene and hornblende that occur so plen- —
tifully developed in the rocks of the region.——In a second paper the
same author‘ attacks the great problem of the origin of igneous rocks. :
He introduces the subject by outlining the growth of the theory first
enunciated by Scrope, that the varieties of igneous rocks are the result —
‘Edited by Dr. W. S. Bayley, Colby University, Waterville, Maine.
*Twelfth Ann. Rep. Di U.S.
3Cf. AMERICAN NATURALIST, April, 1890,
360. ie
‘The Origin of Igneous Rocks. Bull. Philos. Soc. Wash., xii, 1892, p- 89. se
Geol. Survey, Washington, 1892, p- 569-
Be nO ny ae ae yo eee ee eee
ig a ea
1893,] Mineralogy and Petrography. 41
of the differentiation of a homogeneous magma. Scrope’s notion was
a crude one, but it has been built upon little by little until it has, in
the hands of Mr. Iddings, been placed upon a footing secure enough
to warrant its being thoroughly tested by observation and experimen-
tation. The author points out the evidences of the close relationships
exhibited by the rocks emanating from a volcanic center and their
differences from similar groups from other centers, and then takes up
the question of the differentiation of molten magmas. He brings for-
ward geological and chemical evidences of the fact of differentiation,
and explains the act upon Soret’s principle that in a solution whose
parts are at different temperatures there will be a concentration of the
salt in the colder parts. Lagorio has shown that rock magmas are
solutions, and Iddings believes they are solutions of the chemical ele-
ments or of their oxides. Consequently, after differentiation has taken
place and cooling sets in, different mi ] fi d ing to laws
that depend upon the proportions of the oxides occurring in the differ-
entiated portions. This is apparently contradictory to the view of
Rosenbusch,* who regards rocks as having originated in the differen-
tiation of a magma, but of a magma which is a solution of silicate salts
in a silicate solvent. As a result of the condition of affairs suggested
by Iddings the first eruption from a volcanic center would naturally
possess a composition intermediate between those of succeeding erup-
tions. As a fact the author states that the sequence is usually a rock
of intermediate composition, followed by less siliceous and more silic-
eous ones, to those very basic and very acid. The last eruptions are
of very exceptional character. These will occur in small quantity
only, and will be first eroded from the surface. Consequently these
forms will be found principally in dykes. They are the forms to which
Rosenbusch has given the group name “Ganggesteine.” These rocks,
according to Iddings, have their equivalents among volcanic flows, but
the association of minerals in them is different. It is simply their
structure, therefore, that characterizes the dyke rocks. They have
originated in the same manner as have other eruptives, and conse-
quently are not essentially different from them. The author’s views
are developed carefully and at considerable length. They will
undoubtedly serve to turn the attention of petrographers to a subject
that has lain neglected long enough—the comparative study of rocks
of single geological provinces. The paper will well repay very careful
reading by all petrographers and theoretical geologists, who should be
SAMERICAN Natura.ist, Nov., 1890, p. 1071.
42 The American Naturalist. asia :
glad to know that it is on sale by the Philosophical Society of
Washington, from whose secretary it may be purchased for $1.
The Novaculites of Arkansas.—In his excellent discussion of
the novaculites of Arkansas, Griswold* describes most of these rocks a8
consisting of very tiny irregular grains of quartz with occasional —
specks of carbonaceous matter. Originally the rock contained also
wel) crystallized rhombohedra of calcite, traces of which are sometimes q
seen in the sections. Generally, however, the calcite has entirely
disappeared, and its place is now occupied by a rhombic cavity, around
which the quartz grains are packed as though they had been shoved —
about by the crystallizing carbonate. The good cutting qualities of
Arkansas whetstones are thought to be due to the presence of these —
cavities. The purity of the Hot Springs novaculite is shown by an
analysis that yielded :
ent ars sie &
Bias Sond ae MGA > oe Mah ae A ges N
SiO, ALO, aiit be MgO K,O Na,O Loss Total
99.45 2 tr. 19 64 e .06—100.62
According to the author the rocks were first deposited as a mud or —
ooze, in which calcite crystallized. They were then consolidated by —
simple pressure, and finally, after upturning and erosion, they weré :
supplied with a small quantity of secondary silica. i
Petrographical News.—Osann’ has discovered that the mineral —
heretofore regarded as sodalite in the Montreal eleolite-syenite is
nosean, as it contains 5-6% of SO,, and very little calcium. It is quite 4
abundant in the rock, and is Kaea as idiomorphic grains in its :
garnets. A microscopical test proposed by the author for distinguish- a
ing between nosean and sodalite is as follows: Moisten slide with .
dilute acetic acid to which a little barium-chloride has been added, and ig
allow to stand in an atmosphere of the acid. Sodalite remains trans- 2
parent, while nosean is covered with an opaque coating of barium —
sulphate. a
The coloring matter of the black limestone of the Pyrenees is show? —
by Jannetaz* to be carbon, probably in the form of anthracite.
The new catalogue of geology and petrography issued by Ward's 2
Natural Science Establishment, of Rochester, N. Y., deserves mention —
*Ann. Rep. Geol. Survey of Ark. for 1890, Vol. iii, pp. 122-168.
‘Neues Jahrb. f. Min., etc., 1892, i, p. 222.
*Bull. Soc. Franç d. Min., xv, 1892, p.'101.
1893.] Mineralogy and Petrography. 43
in these notes because of the full list of rock names contained in it.
The principal rock types are defined, and under each are given the
technical names of all its varieties. It is further interesting as an
indication of the growing importance of lithology in this country,
since it is quite evident that Prof. Ward would not find it advisable to
keep in stock such a large quantity of rock material were the demands
for it rare. The catalogue may well serve the geologist as a table of
petrographical synonyms.
A New Occurrence of Ptilolite.—A new occurrence of ptilo-
lite has been discovered by Cross and Eakins’ in Custer County, Col.,
about three miles southeast of Silver Creek, in the vesicles of a dull
‘green devitrified pitchstone. The mineral is in very slender needles
that are optically negative. An analysis made on very carefully
selected material gave:
SiO, AlĻO, CaO K,O NaO H,O Total
67.83 11.44 3.30 .64 263 13.44—99.28,
which is equivalent to R, Al, Si, O,,+6% H,O, a formula identical with
that determined for mordenite by Pirsson."” Clarke" regards a part
of the water in each mineral as basic, and believes that mordenite, the
ptilolite from Silver Creek and the original ptilolite (which is poor in
Na,O) are mixtures of the salts. Al, (Si,O,); Ca H, 3Aq, Al, (Si,0,),
Ca H, 6Aq, Al, (Si,O,), Na, H, 6Aq and Al, (Si,O,), K, H, 6Aq.
Mineralogical News.—Polybasite and tennantite are reported by
Penfield and Pearce” from the Mollie Gibson Mine in Aspen, Col.
The former is the ore of the mine. It occurs massive, often associa-
ted with barite and siderite. It is of a grayish-black color, and has
disseminated through it patches of the lighter tennantite. Analyses,
corrected for impurities, follow:
As b Ag Pee Ca Za Be
Polybasite.... 18.13 7.01 .30 56.90 14.85 2.81
Tennantite... 25.04 17.18 .13 13.65 .86 35.72 6.90 .42
Crystals of both minerals are known to occur in several of the Colorado
mines, though they have not yet been described.
®Amer. Jour. Sci., August, 1892, p. 96-
Cf. AMERICAN NATURALIST, 1891, p- 372.
Amer. Jour. Sci., August, 1892, p. 101.
2 Amer. Jour. Sci., July, 1892, p. 15.
44 The American Naturalist. [
The cerussite from Pacaudiére, near Roanne, Loire, France, is stat
by Gonnard” to be associated with copper, silver and lead compounds
pyrite, limonite, quartz and calcite. Its simple crystals present ng
variety of planes. Twinned crystals are common, and trillings an
known. A description of the several types is given by the autho
For sixty years past the same mineral has been known to occur at th
argentiferous galena mines of Pontgibaud Puy-de-Dôme, but the
has not been noted in the treatises on Systematic Mineralogy.
crystals seem to have been formed at the expense of galena and b
nonite by the action of CO, from the neighboring volcanic vents.
habit of its crystals is well described by Gonnard".
Morenosite [(Ni Mg) SO,+7H,0] in green stalactites from the
of the Breithorn in Zermatt, yielded the same mineralogist” the fig
SO, = 28.7; NiO = 185; MgO = 6.5; H,O = 46.5. As
fragment of an ochre-yellow mineral from New Caledonia is a si
of nickel, magnesium and iron:
SiO, FeO, AlO, NiO MgO H,O Total
83.0 185 15 26.3 80 140 = 1019
Frossard” substantiates the statement of Mallard that the b
garnet pyreneite is a grossularite and not a melanite as repor fs
Raymond. Its density varies between 3.375 and 3.53.
Vesuvianite is reported by Pisani" from Settino in the Rhetian
Its analysis gave:
SiO, AlO, FeO CaO MgO MnO Loss Total —
39.0 143 18 3874 67 tr. .9=1001
The supposed martite crystals in the rock of Cuzeau, Mont -
are tabular hematites cemented into octahedra by magnesio-fer
determined by Lacroix.”
In the basic clays of Condorcet near Nyons, Drôme, France, are b
ders of siliceous limestone, with cavities whose walls are lined with bi-p
amidal quartz crystals, transparent celestite, dolomite and calcite. `
quartz and celestite both contain rare planes beautifully develo
“Bull. Soe. Franç d. Min., xv, 1892, p- 35.
“Ib., xv, p. 41.
1893.] Mineralogy and Petrography. 45
Mineral Syntheses.—Bourgeois and Traube” having failed to
produce carbonates of the magnesium group of elements by the reaction
of urea, water and metallic chlorides on each other at 130° in sealed
tubes, have made another attempt at their synthesis by substituting
potassium cyanate for the urea. The attempt proved successful,
needles of aragonite and rhombohedra of dolomite and magnesite
having been produced under the conditions mentioned, when the
chloride used was a mixture of the magnesium and calcium salts in
molecular proportions.
By the slow action of dilute solutions of copper chloride upon freshly
precipitated lead hydroxide at ordinary temperatures there is produced
a blue powder consisting of octahedra and, cubes of percylite, with
which are associated quadruple twins of a colorless mineral supposed
by C. Friedel” to be phosgenite.
Crocoite has been obtained by Ludeking” upon allowing a strong
solution of caustic potash to stand for some time in contact with lead
chromate in the presence of a little potassium chromate. By using a
large excess of very strong caustic potash phenicochroite forms. The
crystallization of the latter substance is due to the abstraction of the
solvent by the carbon-dioxide of the air, and of the former by a fur-
ther reaction between the caustic potash and chromic acid.
New Minerals.—Penjieldite—This mineral, discovered by Prof.
Genth® on the slags from Laurion, Greece, is evidently produced by
the action of sea water upon the materials of the slag. It is usually
in the form of hexagonal prisms with basal planes, or in prisms tapered
by pyramids. The color is white and the lustre vitreous to greasy.
An analysis of the tapering crystals gave: Cl = 18.55, Pb = 78.25,
O = , indicating the formula Pb ©. 2Pb Cl,
Brazilite is a new tantalo-niobate from the iron mine Jacupiranga,
in S. São Paulo, Brazil. Hussak* describes it as occurring in the
magnetite-pyroxene rock called by Derby jacupirangite. It was ap
rated by washing the decomposed residue of this rock in a miner’s
pan, and has heretofore been taken for orthite. Its crystallization is
monoclinic with a: b : e = .9859 : 1 :.5109. A= 98° 457. The
forms observed in its crystals are oP, œP, «P2', sP œ, OP, Po,
*Ib., xv, 1892, p- 13.
“1Tb., xv, 1892, p. 96.
"Amer. Jour. Sci., July: 1892; p. 57.
Amer. Jour. Sci., 18
“Neues. Jahrb. f. Min., ae 1892, ii, p. 141.
46 The Amervean Naturalist. (January, |
2P œ`, P and —P. The crystals are tabular parallel to the orthopin-
acuid and are nearly always twinned, frequently yielding very compli-
cated groupings. The color of the larger crystals varies from sulphur
yellow to black. Their hardness is 6.5 and density 5.006. The pl
of their optical axes is parallel to the clinopinacoid, and the double
refraction is negative. The extinction is 8°-15° in obtuse £, and the
pleochroism varies between dark-brown and oil-green. The minerals
associated with brazilite are apatite, magnetite, perofskite, ilmenite,
and a spinel. An analysis of the new minerals is promised shortly.
Landauer’s Blowpipe Analysis.—This little book” will be
cordially welcomed by English and American teachers in colleges
which the use of a large manual of blowpipe analysis is undesirable.
It is as suitable for classes in mineralogy as in chemistry, since it will
enable the student to determine the composition of a mineral as rapidl
as will the use of the great majority of Determinative Mineralogies
upon the market. Moreover, it possesses one desirable advantage over
compounds are made to serve as distinctive tests for them, in that it
compels the experimenter to study the chemical nature of the sub-
stance with which he is working. A mineral is a definite chemical
substance. A student of mineralogy who is unfamiliar with the com-
position of bodies with which he is working, though he may know
considerable about their physical properties, is neglecting the founda-
tion upon which his knowledge of minerals must rest. The little book
before us is an excellent introduction to the larger works like those
of Brush and Plattner. It is, besides, complete enough for most of
the purposes to which such a book is usually put. Beginning with &
good description of the apparatus and reagents necessary to blowpi
manipulation, it follows with an account of the operations employe
describes Bunsens flame reactions, mentions the distinctive tests for the
various chemical elements, gives Landauer’s and Egleston’s schemes
for the systematic examination of inorganic substances, and closes
with tables exhibiting the reactions of the various metallic oxides,
and in a condensed form the results of the different operations
described in the text. The book must find a place in many labor-
atories.
“Blowpipe Analysis, by J. Landauer. Authorized English Edition by
James Taylor. Second Edition. Macmillan & Co., 1892, pp. 14 and 173.
1893.] Botany. 47
BOTANY.
A New Edition of Wolle’s Desmids.—Botanists who failed
to secure a copy of the first edition of Wolle’s “ Desmids of the United
States,” and who were meditating whether or not to pay the extortionate
prices charged by the antiquarian booksellers, will be glad to know
that the author has brought out a new edition with considerable addi-
tions, which he is offering at $6.50. The book was well worthy of
this new edition, which will doubtless find a ready sale.
Botanical Definitions.—It is the misfortune of every science
that it has a mass of technical words or of words with technical mean-
ings, which must be defined before they can be understood by the
general reader. Indeed the number of these terms is so great in some
sciences, notably Botany, that even the professional botanist is obliged
to turn to some handy volume for the meaning of a strange word. So
we must have glossaries and dictionaries of scientific terms. The latest
one to appear is Crozier’s “ Dictionary of Botanical Terms,” a pretty
volume of about 200 pages, upon which the publishers (Holt & Co.)
have done well their share of the work. Turning to the substance of
the volume we find it disappointing. While it catalogues about 6000
words and omits few words of importance, and while its definitions are
generally not false, they are in very many cases so meagre as to leave
the user of the book little wiser after than before consultation. The
author has failed to distinguish between a true statement regarding a
word, and a definition of the word. Many of the definitions in this
book do not define. As examples, see Accessory Gonidia (“gonidial
formations in some species of Mucorini in addition to the typical
kind”), Apical Cell (“the generating cell of a growing point”),
Archegonium (“the female organ in the higher cryptogams”), Basid-
iospore (“a spore borne on a basidium, as those of mushrooms”’),
Linnzan System (“the system of classification devised by Linnzus,
founded upon the number and arrangement of the stamens and pis-
tils; sexual system”), Sexual System (“see Linnean system”). On
the other hand, some of the definitions of the new terms are well
drawn. The older terms fare pretty well, and are as well defined as
they usually are. It is the new terms which often fare badly. Yet such
a book is not wholly useless. When one needs to confirm his impress-
ion as to the meaning of a word it will be helpful, for, as indicated
48 The American Naturalist.
above, the statements are generally true. The general reader, howey
and the beginner in botany who meets a word for the first time a
who seeks a definition which will give him a clear notion of its
ing, will often turn away disappointed.—Cuarwes E. Bessey.
Timely Words as to the Nomenclature Question.—
this time, when there is not a little of ferment and effervescence 6
the rules which should govern in the nomenclature of plants, it wil
be well for us all to read the following remarks made by Al
De Candolle in the introduction to the “ Paris Code” of 1867. T
convey very well the ideas of the “ moderns ” of to-day. f
“ The system of nomenclature of organized beings, founded by Linn
zeus, was looked upon until the middle of this century as extremely
ingenious, and has been thought, by some authors, a most admirable
one. It was quoted in philosophical lectures and found superior t
that of chemical nomenclature, on account of its adapting itself mo
readily to changes necessitated by the progress of discovery. Bo
ists professed for it the greatest veneration. They boasted of hav
developed a better nomenclature than zoologists, which is not surp i
‘ing, as the most illustrious botanists, thirty or forty years ago, gay
infinitely more attention to this subject than zoologists. í
Nevertheless, of late years, a change has been perceptible; op
is wavering, enthusiasm abated. Here and there, in different countries,
doubts have arisen and complaints have been made regarding the 8y%
tem of botanical nomenclature.” * * *
“Tt follows that it is useful—every ieenty years, for instance
revise the ensemble of received rules.”
“Without going far back it is easy to see om since the end of
eighteenth century botanists have endeavored to free themselves os
many useless shackles put on by Linnzeus and tightened by his disc
ples, above all with relation to the choice of generic names. Jan
dolle [the elder, in Théorie Elémentaire] was ruled by the idea
having the law of priority properly roes a law which, fifty y
ago, was often unscrupulously infringed.” * * *
“ The time must however come, when actually existing vegetable forms
having all been described, herbaria containing undoubted types ®
them, botanists having made, unmade, or oftentimes remade, elevated
or lowered, and above all modified, some hundred thousand groups
from orders downward to simple varieties of species, the num
synonyms tivis, become infinitely greater than that of admi
groups—it will become necessary to effect some great revolution in!
1893.] Botany. 49
formation of science. This nomenclature that we are striving to
improve will have the appearance of an old scaffolding, made up of
parts laboriously renewed one by one, and surrounded by a heap of
more or less embarrassing rubbish, arising from the accumulation of
ieces successively rejected. The edifice of Science will have been
constructed, but it will not be sufficiently clear of all that has served
to raise it. Perhaps there will then come to light something very
different from the Linnzan nomenclature—something will have been
devised for giving definite names to definite groups.” * * *
“Tn the meanwhile, let us improve the system of binomial nomencla-
ture introduced by Linnzus. Let us endeavor to accommodate it to
the continued and necessary alterations that take place in science, and
for this purpose let us diffuse as well as we can the principles of the
method ; let us attack slight abuses, slight negligence, and let us come,
if possible, to an understanding on debated points. We shall thus
have prepared, for some years to come, the way for better carrying out
works on systematic botany.”
Engler and Prantl’s “ Naturlichen Pflanzenfamilien.’’—
This great work is making such headway that another year will nearly
complete it. During 1893 we are promised the Fungi, Hepatice,
Musci and the Pteridophytes. The Gymnosperms and Monocotylodons,
are already completed, while but few families of the Dicotylodons
remain to be worked. Recent numbers treat of the Composite (74),
Oleaceæ, Salvadoracese and Loganiacee (75), and Myxogasters and
Fungi (76), the last by the well-known Mycologist Schröter. His tab-
ular view of the system of classification of the Fungi which he adopts
is instructive. See page 50.
Mo. Bot. Garden,
1895.
50 The American Naturalist. [January,
Fg Hemisporangieæ ........ PAESE
% | Sporangieæ Ancylisti
a | È Eusporangiee.........+++ Monoblepharidinea.
s | Saprolegninee.
2 © | Conidieæ A aape
El Peronosporinee.
g| 3
a= ol § 4
Bi ES Te 4 Mucorinee.
A Conidieæ è
è al { Entomophthorinee.
Hemiascee .......... | Fretomyretina
scoidinee.
r Protoascese Sacchar omycetinee.
2 Endomycetinee.
gi ( § (Gymnocarpee............. Taphrine
S g A Helvellineæ
g1 2438 | $4 Hemikleistocarpee....... Pezizine
S SI ET E| Rieistoe Phacidiineæ
4 £a £ 4 2 ATPCH......eeeeees 4 Tuberinee.
a Plectascese mn
= | Elaphomycetinee
| Pyrenoasceæ Perisp bi r
ine
g Hysteriinee.
$ Pa ae
>} Ustilaginee.
I r Tilletiineæ.
Si Sg oe ese a... | Uredinee.
idi à Auricularinee.
; Schizobasidiee............ Tremellinee.
3 pe Dacryomycetinee.
S . ‘ Exobosidiinee.
g j.s Thel. ephori COB»
2 4 = r = Gymnocarpeæ.. cesetini sbss Clavariineæ
E E 451% Hydneinee
EIERE cae Les
AS 2) Hemtenet oletinece.
E É arma aiaaoasit a \4 aricinee
= a Phallinee.
Anr ymenogastrneæ.
; | OIOCArPeæ..... eessen | jeop erdineæ.
L | Plectobasidieæ. Nidulariinee.
4 Scleroderminee-
ST ee Rah es ee ie
1893.] Zoology. 51
ZOOLOGY.
Locomotion of Limpets.—Herdman records’ several facts
which seem to militate against the view that limpets do leave their
resting place and return to it again. It has been shown that they can
leave and travel some distance, but he found a specimen of Patella
vulgata which was sticking to a cylindrical bar of iron and which had
the shell molded to fit the surface. Now as the bar was short and free
to move about, the probabilities are that if it once left the support it
would never be able to return to it. In other cases he found limpets
at the bottom of deep pits, from which it would be very difficult, if
possible at all, for them to extricate themselves.
Tunicate Studies.—Herdman publishes’ some notes on the
structure of the Appendicularian, Œkopleura. This form was studied
by serial sections, and the results, most interesting, are: The condition
of the endostyle as a diverticulum to a great extent shut off from the
branchial sac; the presence of a genital duct; the distribution of the
enlarged ectoderm cells and the cuticular test; the exact course of the
nerve cord through the posterior part of the body; and the shapes and
positions of the alimentary and reproductive viscera.
In the same publication’ Garstang points out that Appendicularia
mossit (Mossia dolioides) is to be regarded as a member of the genus
Kowalevskia of Fol, and that it in reality has not that importance
from the phylogenetic standpoint which was attributed to it by Herd-
man in his “ Challenger” report.
The Skeleton and Teeth of the Australian Dugong.—
Zoologists are indebted to Prof. G. B. Howes and Mr. J. Harrison fora
valuable paper on the skeleton and teeth of the Australian Dugong,
of which the following is an abstract :
“The authors showed that the vertebral epiphyses are more fully
developed than Albrecht has suspected, and that they appear late and
rapidly ankylose with the centra, a feature of especial interest, in view
of Lefévre’s alleged discovery of fully developed epiphyses in Hali-
therium schinzii and Metaxytherium. On comparison with the Cetacea
‘Trans. Liverpool Biol. Soe., vi, 22, 1892.
*Trans. Liverpool Biol. Soc., vi, 40, 1892.
3L. C., p- 57, 1892.
52 The American Naturalist. [January A
they sought to associate the reduction of the epiphyses with adaptation
to an aquatic existence.
“In dealing with the limb-skeleton they described a longitudinal
cleavage of the phalanges, akin to that recorded by Kükenthal for the
Cetacea. The only structures observed which were at all comparable
to sup y phalanges derivations of the terminal (ungual)
ones, arising proximally; and the observations lend no support to
Kiikenthal’s view that supernumerary phalanges are epiphysial in
origin. :
“ The first upper incisor and the four lower ones of either side were
shown to have milk predecessors, which are early absorbed. Five
teeth were shown to be present on either side of the symphysial region
of each mandibular ramus of Manatus, the fifth one being claimed as
a canine; and in this animal the authors described milk predecessors
to the two anterior pairs of mandibular cheek teeth.”
On the Cephalo-humeral Muscle and the So-Called Clav-
icle of Carnivora.—At a meeting of the Philadelphia Academy,
Dr. Harrison Allen spoke of the peculiarities of the cephalo-humeral
muscle in mammals and invited especial attention to the presence of a
small fibro-cartilaginous disc in the junction of the cephalo-humeral
with the muscles which are inserted in the bones at the region of the:
shoulder. This is well defined in Felis and is identified as a rudimen-
tal clavicle. Dr. Allen has detected this structure in Herpestes, Taz-
idea, Cercoleptes, Bassaris, and Procyon.
The cartilage is either in the form of a flat dise or a minute scythe
shaped rod, and is constant in lying directly over the greatest convexity
formed by the round of the shoulder. It seems to give strength to the
center of a muscle system of which the cephalic, cervical, pectoral and
latissimal sheets are parts. The identification of such a plate or rod
with a true clavicle is doubtful, since in Balantiopteryx (a genus of
bats) the structure above described is remarkably developed, while the
clavicle is as well formed as in any other animal. ‘Phe long rod-like
body is continuous with a fascicle of fibres arising from the pectoralis
and receives the insertion of the occipito-pollicalis. The anterior end
of the rod lies in the upper border of the wing membrane and is con-
tinuous with the fibrous thread which represents the tendon of the
occipito-pollicalis as this muscle is defined in the bats generally. From
both the proximal and distal divisions of this muscle delicate fascicles 4
pass toward the elbow and the entire plan appears to be associated
with the rudiment of the characteristic skin sac. Slight modification —
_ of this arrangement is met with in the allied genus Rhynchonycteris.
Re A
ane
ai
aa
ay
a $
S
1893.] Zoology. 53
Comparison of this arrangement with that seen in the common
brown bat (Adelonycteris fiuscus), the noctula bat (Noctulinia noctula),
and the false vampire ( Vampyrus spectrum) showed by the part taken
by the rod in Balantiopteryx is the tendon of a pectoral muscle-fasci-
cle which is inserted into the occipito-pollical muscle as it crosses the
shoulder, while in the group of the Molossi the muscle-fascicle is fleshy
throughout its entire extent, but on the whole preserving the same
relations, Thus the fibro-cartilage of Balantiopteryx is. represented
by fibrous tissue in Adelonycteris and both these in turn by muscle in
the Molossi. Dr. Allen believed that it was inexact to speak of a
clavicle and of this rod as things which were equal. The clavicle acts
with the scapula in supporting the head of the humerus but in no wise
limiting or determining its movements, while the rod is always over
the outer aspect of the shaft of the humerus below its head and acts
as a check to abduction of this bone—Proceeds. Phila. Acad., Pt. 2,
1892.
A New Synaptomys from New Jersey.—While trapping
for a type series of the new race of Evotomys described by Mr. Stone
in the present number of Tor Naturatist, I had the fortune to
secure a specimen of this long-looked-for genus, which is, I believe,
the first taken in flesh east of the Alleghany Mountains.
It had previously been detected by the U. S. Department of Agricul-
ture in the rejects of a barn-owl living in the tower of the Smithsonian
Institution.
_ A comparison of the New Jersey specimen with two Synaptomys
cooperii from Ohio, courteously loaned by Mr. J. A. Allen, of the
American Museum of Natural History, N. Y., shows such marked
specific differences that it will be unnecessary to more than briefly
allude to them.
SYNAPTOMYS. STONEI.—Sp. nov. Type No. 567. ad. 9. coll. S. N.
Rhoads, May’s Landing, N. J. Dec. 2, 1892.
Special Characters—Outward appearance and proportions as in
S. cooperti. Above blackish-brown, with black hairs more predomi-
nant over the shorter brown hairs than in cooperii. The same color
reaching around sides of belly instead of being confined to dorsal area
as in cooperit. Hoary gray belly and neck of cooperii replaced by dark
plumbeous gray. Feet, including soles, plumbeous, without brown
shade. Two middle toes of fore-feet and four inner toes of hind feet,
including nails, white. Tail unicolor plumbeous gray. Lips encircled
with narrow white edgings.
54 The American Naturalist. [ January,
Skull narrower, shallower, and, viewed from above, less angular than —
that of cooperii, but of same length. Lower jaws viewed from below,
ditto. Incisors shorter, broader, and less cylindrical, with sulcation
of upper pair much more distinct. Zygomatic foramen longer and
narrower. Sagittal suture and parietals relatively much longer;
interparietal transversely narrower, longitudinally longer. Supra- —
occipital in cooperii twice as wide as deep, in stonei thrice as wide as
p. eo
Molars one-third wider and one-eighth longer in stonei. In cooperii :
the length of the symphysis mandibuli just equals the distance from
its posterior end to the angle formed by the antero-inferior border
of the masseteric fossa ; in stonei the symphysis i is one-third longer.
Posterior face of angle of lower j jaw in stonei very stout, abruptly
rounded, and recurved outward; in cooperii it is slender, spatulate, .
elongated posteriorly in a nearly vertical plane, and the margin below
the condyle not thickened as in the former species. :
Measurements in millimeters of the New Jersey specimen in the
flesh, with averages of six alcoholic specimens of cooperii, made by —
Dr. Coues, are given :
Full length. Tail. Foot. Ear. ;
Synaptomys cooperii.......... 105 18 18 ba
Synaptomys stonei 117 18 18 A
The age of specimens on which the above cranial and color charac —
ters are based is evidently about the same. In other respects they
may be safely considered normal adult representatives of the species 2
the different localities where they were taken. The new species may
Curator of Birds in the Philadelphia Academy of Natural Sciences. —
.SamuEL N. Rwoaps. 4
-A New Evotomys from Southern New Jersey.—On October
25, 1892, while collecting small mammals near May’s Landing, N
Jersey, in company with Mr. S. N. Rhoads, I captured a specimen of
Evotomys, a genus which has not previously been reported from souti
of Massachusetts and the Adirondacks, except in the higher mountains
of North Carolina, The next day three more specimens were secu
and subsequently (December 2) Mr. Rhoads collected four others 5
the same locality. A comparison of these specimens with a series 0
Evotomys gapperii from Northern New York, which is apparently the
most closely related form, shows them to be subspecifically distinct,
put
1893.] Zoology. 55
and I therefore propose for the New Jersey animal the name Evotomys
gapperi rhoadsii in honor of my friend, Mr. Samuel N. Rhoads.
The comparison of a series of skulls of E. gapperti and E. g. rhoadsii
fails to show any constant differential characters, though the immature
specimens of the new race are peculiar in the structure of the last
upper molars. In these teeth the first reentrant angle on the inside is
opposite the second salient angle on the outside instead of the first
reentrant angle as is the casein the adults of both forms. One young
specimen of E. gapperii shows a tendency to this structure, but in all
the other specimens that I have examined the reentrant angles meet,
and the outer one is deflected posteriorly.
In proportions the New Jersey race seems to average rather smaller
than E. gupperii from the Adirondacks, while the tail is shorter and
the feet slightly longer than in that species.
As regards coloration E. g. rhoadsii is everywhere darker than E.
gapperii, and has a plumbeous cast on the sides and flanks, while it
lacks almost entirely the buff suffusion generally seen on the sides and
under surface of the latter species.
Above the color is decidedly darker than in E. gapperii, and there
are a great many more black hairs scattered over the back. The red-
dish area is not so well defined and the color is darker—more of a
mahogany shade. The tail is distinctly bicolor, but the upper surface
is darker than in E. gapperii, and the feet have a decidedly gray suf-
fusion, contrasting strongly with the pure white of the latter species.
Some immature specimens of E. gapperii approach adult E. g. rhoadsii
in general coloration, but the young of the latter race with which
they should properly be compared have scarcely a trace of the reddish
dorsal area, the middle of the back r Being brownish and the sides gray.
The table on next page, will show the ts of the
two forms, the specimens of E. gapperii being selected from a series
kindly loaned me by Mr. G. S. Miller.
Dr. C. Hart Merriam, of the Department of Agriculture, Washing-
ton, D. C., has kindly examined my New Jersey material and compared
it with Evotomys carolinensis and other species to which I had not
access, but its closest relationship appears to be with E. gapperti. All
the specimens of this new mouse so far secured were taken in a cran-
berry bog on the Egg Harbor River, about a mile above the town of
May’s Landing, N. J. The unexpected occurrence of this boreal genus
well within the Carolinian Fauna may probably be accounted for by the
theory already advanced by Dr. Merriam that in these damp bogs,
where the temperature is much lower than in the surrounding dry
56 The American Naturalist. (January,
Evotomys gapperii.
Tail Hind
No. Sex. Locality. Date. Length. Vertebra. Foot
444% 3 Peterboro, N. Y. July 17,1892 151mm. 41mm. 17
THE Q x ` July 19, 1892 165 51 18.4
THE 9 A S Aug. 1,1892 160 46 19.6
4344 3 Keene Valley, N. Y. Mar. 17,1892 155 45 20
Average 158 46 19
Evotomys gapperii rhoadsii.
Tail Hind
No. Sex. Locality. Date. Length. Vertebre. Foot.
-160 ¢ Type, Coll. of W. Stone. Dec. 2,1892. 142 40
161 9 s a “ “ 130 37 2%
570 9 Coll. ofS. N. Rhoads. “ « 123 34 21
571 3 7 ss “ 130 36 20
Average “ii 87 20
areas, the conditions of life are quite suited to more boreal species,
especially animals of nocturnal habits. The presence of various Eri-
caceous and other boreal types of plant life in these locations also
supports this hypothesis.—Wrrmer Srone, Academy of N atural
Sciences, Philadelphia.
- Zoological News.—Verrepratra—Some new reptiles and fishes
from Australia are described by J. Douglas Ogilby. The list com-
prises Typhlops curtus from the Gulf of Carpentaria, Hoplocepalus
suboccipitalis from Morel, and Clupea sprattelloides from rivers flowing
into Port Jackson and Botany Bay. The latter species has until now
been supposed to be the young of C. nove-hollandia.—Records Austr.
Mus., Vol. ii, No. 2——F. W. True reports that the collection of
African mammals presented to the National Museum by Dr. Abbott
contun several species apparently new: Dendrohyrax validus, Mus
aquilus, Dendromys nigrifrons, Seiurus undulatus, Cephalophus spadia
The known range of several species is considerably extended by Dr.
Abbott’s labors. The mammalian fauna of the Kilima-Njaro region —
as indicated by this collection includes seventy-one to seventy-three —
species.— Proceeds. U. S. Natl. Mus., Vol. xv, pp. 445-480.
1893.] Embryology. 57
EMBRYOLOGY.
Gastrulation of Aurelia.*—Frank Smith has entered into the
controversy between Claus and Goette concerning the origin of the
entoblast of Aurelia. The results obtained from his first sections led
him to think that the conclusions reached by Goette for Aurelia aurita
would be confirmed in the case of A. flavidula. Better staining,
thinner sections and more accurate orientation have, however, made it
certain that the gastrulation in this species is much more in accord
with the description given by Claus and that the process really is one
of invagination. ‘The result of cleavage is a one-layered blastosphere
as in A. aurita. The cells of the blastosphere are usually somewhat
shorter at one pole than elsewhere, and it is from this region that the
entoblast is formed. It develops as a single continuous layer of cells
surrounding a small cavity, the coelenteron. From the beginning
there is a narrow blastopore. Only a small portion of the wall of the
blastosphere is concerned in the invagination, and to that extent it is
not typical. The celenteron is, however, at all stages, an open sac-
like cavity, and therefore noticeably different from that of A. aurita as
described by Claus. The colenteron enlarges until the cleavage cavity
is entirely obliterated and the entoblast everywhere comes into contact
with the ectoblast. The entoblast, at first thin, thickens after the
completion of gastrulation.
While the entoblast is formed by invagination, ingression of cells
from the wall of the blastosphere into the cleavage cavity does occur,
although only in a minority of cases. It may happen any time after
the blastosphere contains about 100 cells, sometimes before invagina-
tion. When this phenomenon takes place it is similar to that repre-
sented by Goette (Figs. 1-5) for the earlier stages of the blastula in
A. aurita, and consists of the migration into the cleavage cavity of
one or two, rarely more than three, of the cells of the blastospheric
wall. Soon after invagination the nucleus of the cell disappears and
the cell breaks down, or, less frequently, persists until gastrulation is
complete. In the latter case it becomes forced through the entoblastic
wall into the cavity of the ccelenteron. The cause or purpose of this
immigration does not appear.
ea department is edited by Dr. E. A. Andrews, Johns Hopkins Uni-
versity.
2Bull. Mus. Comp. Zool., Harvard, xxii.
_ now much smaller than it was and has no nucleolus, while in the
58 The American Naturalist. [January,
The author thinks that the difference in opinion between Claus and —
Goette is partially due to there being two kinds of cells that find t
way into the cleavage cavity. Besides the large cells just deseri
he found in a much smaller number of cases one or two very sm
cells that look precisely like the small cells that appear in the deep
part of the ectoblast at about the time gastrulation begins.
There is no evidence that the immigrating cells have anything to
with the formation of the entoblast, and Goette’s case is further weak-
ened by the fact that all the conditions shown in his figures (6-9)
easily be reproduced from sections of invaginating gastrule of
single stage of development.
Cleavage in Aequoria Forskalea.—Dr. V. Hecker’ contri
utes an interesting series of observations on this subject. He fim
that when the specimens are in good condition the time relatio
between the successive periods of activity are remarkably precise.
If ripe, the eggs are laid between 7 and 7.30 A. M. The first polar
body is extruded at 9 A. M. The entrance of the spermatozoan and
the division to form the second polar nucleus takes place at 9.30.
10 A. M. the dyaster stage of the first cleavage nucleus occurs,
with it the first indication of the division of the cell body. 4
daughter nuclei are undergoing metakinesis at 11, and at 12 the four
nuclei of the next set are in the dyaster stage. The nuclear divisions
continue to take place an hour apart at least as far as the sixty-four —
cell stage, and this seems to show that the nucleus is not affected
this period by the amount of cell protoplasm that it controls. 4
mally nuclear division takes place at the same time throughout i
egg, and the blastomeres are of equal size up to the sixty-four-cell sta
When eggs are laid after the specimen has been kept in an aquari
several days, irregularities generally occur in the time of nuclear
ion and in the size of the blastomeres. At the same time a path
ical form of nuclear division, the triaster, appears, and the mas
cells loses its spherical shape.
A remarkable feature of the cleavage of the egg of Aequoria
presence of a body for which the author proposes the name Me
eleolus. In the older ovarian eggs and in eggs just laid there 18 ®
large nucleus containing a very fine network of chromatin and àla!
~ spherical or renifurm nucleolus. About half an hour after the egg
_ laid this nucleolus appears to have been extruded, for the nucleus
"Archiv. fur Mikro. Anat., 40 Bd., 2 Heft.
1893.] ` Embryology. 59
outside of the nucleus but close to it there is a body resembling the
former nucleolus in every particular, except position. This body, the
metanucleolus, never divides, there is never any radial arrangement
of the protoplasm about it, and it may be found in one of the blasto-
meres until a later stage in the cleavage. From a review of the work
of Metschnikoff, Boveri, and others, the author thinks that homologous
structures have been seen, although wrongly interpreted, in the Lepto-
A ntl d apparently also in Mytilus
and Sagitta. He has also examined Weismann and Ischikawa’s prep-
arations of the winter eggs of Daphnids, with the result that he
regards the paracopulation cells as not cells at all, but as in all proba-
bility structures homologous with the metanucleoli of the meduse.
Another point of interest in this paper is the numerical relation
between the chromosomes of the second polar spindle and of the first
eleavage spindle, there being six in the former and twelve in the latter.
Boveri had pointed out that in the forms in which this point had been
studied, while the number varied in different species, the number of
chromosomes in the cleavage spindle was always just double the number
in the last polar spindle, and he had also noticed that the number of
the latter in certain species could be arranged in a geometrical series
in which the numbers are forms of two (2, 4, 8, 16,32). Hecker
reviews the literature of the subject and shows that there may be two
other series besides this one. There is a series of the forms of three
(3, 9, 27, etc.), of which, however, there is but one example, Echinus
with nine chromosomes, and then there is a mixed double and triple
` system (6, 12, 18, 24, 36, 48, etc.), to which Aequoria belongs, as well
as the greater part of the insects and the vertebrates. He concludes
that all cases so far known may. be arranged in three systems in such
a way that in general nearly related forms belong to the same system.
R.
meduse,
IGELOW.
(1) Zeit. f. Wiss. Zool., 49, 1890, pp. 503-580, plates 24-26.
(2) Zeit. f. Wiss. Zool., 51, 1891, pp. 685-730, plates 35-37.
(3) Zeit. f. Wiss. Zool., 54, 1892, pp. 1-249, plates 1-12.
little wider than head, the neck being somewhat constricted; &
60 The American Naturalist.
ENTOMOLOGY."
The Pupa of Argyramceba cedipus Fab.—The descripti
given below is drawn from a pupal skin sent to me with the fly by
Prof. C. P. Gillette, who bred the latter from a nest of Odynerus
at Fort Collins, Colo. In a paper which will be published in Psyche,
I have described the pupa of Toxophora virgata O. S., and also made
some mention of the pupæ of Bombyliide which have so far been
described. The pupa of the present species differs quite markedly
detail from that of the Toxophora above mentioned.
Pupa of Argyrameba edipus: General color of empty pupal
very pale straw colored; the cephalic horns black, reddish-brown
basally, anal horns black, slightly reddish-brown at base; dorsal rows
of ridges reddish-brown, the terminal spinous processes blackish; pro
thoracic spiracles reddish-brown, other spiracles but little darker than
rest of integument, slightly brownish. Head conforming to shape
head of adult fiy, more or less sub-spherical in form. Eight ceph
horns or teeth arranged in four pairs, the three anterior pairs joi $
in a common base, the posterior pair removed from others; anterior
pair longest, their rufous brown joined hasal portion distinctly shorter
than their black free terminal portion, moderately slender, nearly :
straight, directed forward, gradually tapering to tips; second and third
pairs more closely approximated one pair to the other than are !
first and second pairs, but the two horns of each pair widely remo
from each other, both pairs directed nearly forward but at a slightly
more downward angle than first pair; third pair much shorter than
‘second, more curved or claw-like ; posterior pair closely approxi mash
at base, nearly as large as third pair, but straight and directed infer-
iorly, situated nearly in middle of ventral surface of head segment; :
pair of quite widely separated divergent bristles on outer dorsal sur-
face of anterior cephalic horns; a more approximated nearly para
pair just posterior to these but arising from integument of head-
anterior dorsal edge, being situated just behind the suture at base
cephalic horns; a divergent pair situated just anterior to base of tas
posterior of fourth pair of cephalic horns; a small bristle on each side
at hind margin of ventral surface of cephalic segment. Thorax
.
‘This department is edited by Clarence M. Weed, Hanover, N. H.
Pe ie Seer eee ey eee ee eS yey ee
7 pa z ji = ee ey e
1893.] Entomology. 61
closely approximated short bristles arising from the same papilla on
lateral dorsal surface of thorax a little anterior to middle; a bristle
below these on pleural surface, another still below and a little anterior
to this one. Wing cases reaching about to base of third abdominal
segment, leg cases a little longer. Scutellar segment about as wide as
thorax, with a transverse row of ten or eleven long, more or less
curled hairs on each side of dorsum approximated to anterior margin,
there being a bare space on median portion of segment between the
inner ends of the rows; about ten somewhat longer similar hairs on
extreme Jateral portion of segment on each side, arranged in a more
or less complete semicircle, the open portion of the semicircle being
toward the posterior end of body; the lateral hairs are longer and
slightly stouter than those of dorsum, being nearly as long as trans-
verse width of segment. Abdominal segments one to four, about same
width as scutellar segment, each armed on dorsum with a transverse
row of short longitudinal parallel chitinous ridges or very narrow
plates, there being thirteen in a row on first and second segments,
twelve on third segment, and nine on fourth, the rows a little approxi-
mated to posterior margin of segment, especially in middle; these
ridges are about two-sevenths as long as length of segment, those in
middle of rows being the largest and heaviest, the outer ones shorter
and diminishing in size, those on fourth segment less heavy than those
of first to third segments, and each ridge is produced at its ends into
a spinous or hook-like process, the ridges in profile presenting a cres-
-centic appearance with the concavity uppermost. The other abdomi-
nal segments without these rows of ridges, fifth segment nearly as wide
as preceding, sixth segment hardly narrower than fifth, the fifth and
sixth segments each with a transverse continuous row of hairs on
dorsum arising from a transverse ridge, extending down to lateral
ventral edge of segment and continued on sides of venter, these rows
somewhat approximated to posterior margin of segment. Segments
(abdominal) one to four with a thin transverse row of shorter weak
hairs on each side of dorsum, arising in posterior edge of rows of ridges,
extending down on each side to lateral margin, no hairs on median
dorsal portion ; same segments with a more or less complete lateral
semicircle of longer hairs as on scutellar segment, but somewhat
weaker and shorter than those on that segment, the hairs on fourth
segment extending beneath on sides of venter. Seventh abdominal
segment much narrowed, rapidly and evenly narrowiug from
posterior margin, its width on posterior margin hardly more than one-
third its width anteriorly, its mean width about one-half that of sixth
62 The American Naturalist. [January,
segment, with a transverse row of several hairs on each side of dorsum
extending below on edge of venter, discontinued in middle on dorsum,
slightly approximated to posterior margin of segment. Eighth or
anal segment narrow, same width as posterior margin of seventh, nearly
as long (to base of horns) as wide, terminated by three pairs of anal
horns; anterior or upper pair short, small, situated at base dorsally of
middle pair; middle or second pair long, curved slightly upward ter-
minally, nearly as long as length of segment, widened inwardly on
basal half so that the bases are closely approximated, inner outline
hollowed out on apical portion, longitudinally corrugated at base
above, with a dorsal longitudinal groove widening to hollowed portion
and then extending narrowly to tip, moderately sharp at tips; third
or inferior pair short, small, hardly as large as anterior pair and not so
stout at base, directed more downward than middle pair, situated on
outer base ventrally of middle pair ; just anterior to first pair on dor-
sum there is a median very small spinous tubercle, apparently a rudi-
ment (or herald) of a fourth pair of anal horns. Prothoracic spiracle
situated on lateral front border of thorax (prothorax) just anterior to
wing bases, mesothoracic spiracle not apparent, metathoracic spiracle
situated anteriorly on lateral edge of dorsum of scutellar segment;
first to sixth abdominal pairs of spiracles situated on anterior edge
laterally of dorsum of first to sixth abdominal segments ; seventh pair
situated one on each side of dorsum of seventh segment immediately
behind the transverse row of hairs. It is interesting to note that a
quite long section of the trachez is left attached in most cases to the -
spiracles on inside of the pupal skin, especially to the thoracic pairs.
The fly escaped by the pupal skin splitting along the dorsal median
line of the head and thorax, the slit extending slightly into the scutel-
lar segment; also splitting laterally backward on each side of head
from a little above the base of anterior cephalic horns along what
would nearly correspond to the frontal fissure in Muscide, the break
curving shortly and obliquely upward to thoracic suture, and allowing
the nearly triangular posterior dorsal or upper section of the integu-
ment of the head to become loosened laterally below from its junction —
with the thorax, and hanging like a flap by its median dorsal junction.
Length, 93 mm.; width of basal abdominal segments, 2} mm,
The anal extremity of this pupal skin is distended with a dirty
colored hardened fluid ventrally, just below and anterior to anal
horns, into a large round tubercle with a subcentral deep pit or orifice-
like depression which is approximated to posterior margin, the anterior
portion of the tubercle being greatly bulged and distended. The
1893.] | Entomology. ` 63
diameter of this false tubercle is as great as the posterior width of the —
sixth abdominal segment. The fluid which distended it is perhaps
homologous with the meconium of butterflies.
The description of the manner in which the pupal skin splits to
allow the escape of the fly was omitted in the description of the pupa
of Tdxophora virgata in the article above referred to. It is accom-
plished in the same way as just described for the present species, except
that the dorsal median split does not reach posterior margin of thorax,
and the dorsal pieces of head are notso much detached from their lat-
eral thoracic fastenings, and are left more quadrangular in shape by
the oblique lateral breaks of head. It may also be mentioned that a
section of the trachez is left attached to inside of prothoracic spira-
cles.
. My reasons for calling the first abdominal segment of other authors
the scutellar segment, are stated in the article on Toxophora.
C. H. TYLER TOWNSEND.
The Horn-Flyin Canada and Texas.—Mr. James Fletcher,
Entomologist to the Canadian Department of Agriculture, announces?
that the Horn-fiy (Hæmatobia serrata) has appeared in enormous
numbers in the Provinces of Ontario and Quebec, causing considerable
anxiety to stock-owners. It was first definitely heard from at Oshawa,
Ont., July 30, 1892. An excellent résumé of the life-history of the
pest and of the means of' preventing its injuries is given.
That this insect is also spreading rapidly in the southwest is shown
by the following note from Dr. Mark Francis, of the Texas Agricul-
tural College, who wrote me under date of Oct. 18, 1892, from College
Station: “The horn-fly seems to be spreading westward. I saw it at
Stillwater, Oklahoma, two weeks ago. It has not reached here yet,
but I saw great numbers of them at Hempstead, Texas (forty miles ,
southeast of here) last Friday. I think there can be no doubt as to
its identity, as I have compared them with type specimens from Prof.
H. Garman, of Kentucky.”
Two days later Dr. Francis again wrote that the horn-fly was
observed at College Station, Oct. 19, for the first time.
In Southern New Hampshire this insect has been very numerous
the past season, and it has been gradually spreading northward through
New England. Buta hopeful report comes from New Jersey, where
the insect was first observed. Prof. J. B. Smith states that it now
causes little trouble there, and is seldom noticed as specially abundant.
2Central Exper. Farm, Ottawa, Bull. No. 14.
64 The American Naturalist. [January,
The Wheat Frit-Fly.—Dr. Otto Lugger reports’ extensive
damage to wheat in the northwest by a larva supposed to belong to one
of the frit flies. The stem is injured about three inches above the
ground, the larve occurring immediately above a node. The insect so
weakens the plant at this point that the stalk falls over some time
before harvest, the grains do not fill out, and reaper passes over the
fallen stem. ‘The puparia resemble the “ flaxseed” state of the
Hessian fly, and are found within the culm. It is supposed that the
insect hibernates with the puparia. Burning and plowing under the
stubble are the remedial measures recommended. ‘This is apparently
the worst frit-fly attack on wheat yet recorded in America. Dr.
Lugger says that in many places fully one-fourth of the entire crop of
wheat has been destroyed and in a great many more the losses amount
to at least one-tenth,
Entomological Notes.—That excellent periodical, Entomologi-
cal News, has instituted a department of economic entomology, with
Prof. J. B. Smith in charge. This magazine will prove very useful to
amateur as well as professional entomologists, and deserves cordial
support. There has lately been a tendency to insert only very short
articles, or to continue a single article through several issues (some-
what after the fashion of Entomologica Americana), which is unsatis-
factory to all concerned.
Mr. M. H. Beckwith discusses‘ the injuries’of the strawberry weevil
(Anthonomus musculus) in Delaware, and reports finding the larve
feeding upon the ovaries of strawberry blossoms. He surmises that
there may be two or possibly three broods each year, but has been
unable to trace the life-history of the insect during the latter summer
months.
Concerning the recent bestowal by the University of Heidelberg of
the honorary degree of Doctor of Natural Science upon Baron C. R.
von Osten Sacken, Prof. S. W. Williston writes: “Baron Osten
Sacken’s work has been chiefly related to American Dipterology, but —
the ripe fruits of his wide experience and broad grasp of principles
have epriched all dipterology, and, I believe, all entomology. Others
there are and have been who have won enviable honors in systematic
dipterology ; others who have written more extensively than be, but
no one has written more that will be appreciated in the future than
has Baron Osten Sacken.
*Minn. Exp. Station, Bull. No. 23.
‘Delaware College Exp. Pinin, Bull. No. xviii.
*Payehe, Vol.6, p. 346.
1893.] Entomology. 65
Dr. J. C. Neal discusses’ a number of injurious insects that have
appeared in Oklahoma. He includes Preris rape, Plusia brassice,
Heliothis armigera, Diabrotica vittata, Lytta cinerea, Oncideres cingu-
lata, and Blissus leucopterus
The recent biennial report of Prof. S. A. Forbes as Director of the
Ilinois State Laboratory of Natural History, shows that entomologi-
cal studies are being vigorously prosecuted in that favored State.
Fully 20,000 specimens have been added to the pinned collections, and
2700 bottles and vials to the biological series. We are glad to note
the announcement to two important papers soon to appear in the Bul-
letin of the laboratory, the first by Mr. John Marten, containing
descriptions of new species of Illinois gall gnats, and the second by
Mr. C. A. Hart, a descriptive list of the aculeate Hymenoptera of
Illinois.
PSYCHOLOGY.
Notes on Habits of Certain European Birds.—M. Ch. von
Kempen has recently published some observations on birds from which.
the following extract is quoted to show the voracity of the ordinary
sparrow-hawk (Aceipiter nisus):
“ For several years I lived in the country, and was accustomed to
write during the summer near an open window. The apartment had
from one side a view of the garden ; from the other one looked out
over the fields. Suddenly I saw a sparrow-hawk dart through the
room; he flew with such violence that he broke the glass of the win-
dow, against which he dashed in his impetuous flight. I soon had an
explanation of the circumstance. A linnet (Sylvia hortensis) perched
near me was eyidently the attraction. The warbler had flown into the
room to escape the hawk, which in headlong pursuit, had gone through
the room like an arrow from a bow.
“In February, 1889, I had in my town garden a certain number of
lapwings ( Vanellus cristatus); each evening, when I would go to shut
them up in a cage, I would find one less than I had counted in the
morning ; I attributed this loss to a cat belonging in the neighborhood.
The third day on missing another of my pets, I resolved to discover
the thief, and concealed myself for that purpose. In the morning I
saw a sparrow-hawk coming straight to my garden from the old tower
Oklahoma Agri. Exp. Station, Bull. No. 3.
5
66 The American Naturalist. [January,
of Saint Bertin. In spite of my presence he tried to carry off his
daily meal, but I struck him with my hands and made him drop his
prey. I then put my lapwings in their cage, as I was expecting to go
out after dinner. What was my astonishment on going to see my
birds in the evening to find the sparrow-hawk keeping them company
in the cage. He had forced himself in through the bars but could
not get out in the same way, and so was a prisoner with the lapwings,
which he had not, as yet, dared to touch. The hawk was a young
male, and now forms a part of my natural history collection.”
Two other citations show that birds can familiarize themselves with
objects which ordinarily terrify them.
“The jackdaws (Corvus monedula) and barn-owls (Strix flammea)
are very numerous in all the towers of Saint-Omer; they are so accus-
tomed to the noise of the clocks that they build their nests against the
clappers.
“Last year I saw a nest of a titmouse (Parus major) built in a
little mill that children played with in a garden. This noisy scare-
crow, turning with every wind, did not frighten the saucy birds,
and they reared their young with comfort.
“ I now give two observations of another sort that prove beyond a
doubt that birds possess a memory :
“Thad in the country two domestic peafowls (Pavo domesticus) ;
they were accustomed to come every evening to get their slice of bread
cut in small bits before perching themselves on the roof; and if they
were forgotten they would wait nearly all night before abandoning all
hope of the treat. They were so tame that the male, as well as the —
female, would eat from our hands. After I had gone to the city, in
order to keep the peafowls out of the garden, where, it is well known,
they cause great havoc among the vegetables, the berries, and the cur- _
rants, they were given corn in abundance in a place quite remote, but —
they declined these overtures and returned constantly to the place —
where I had fed them; I found them there on my return the following —
year. During the summer the female laid ten eggs, a less number —
than usual; nine young chicks were born, which, following the exam- —
ple of their parents, came every evening to look for their repast. i
_ “During the winter a storm, accompanied by a fall of snow, burst _
upon us during the night, and the unfortunate peafowls were thrown —
from their perch on the roof; some wandering dogs strangled them, —
and we found their remains scattered over the field. :
“T have at this time two laughing gulls (Larus ridibundus) living.
I give them twice a day, at regular hours, bits of meat. Some jack-
4893.] Psychology. 67
daws (Corvus monedula) come every day, at the exact hour, never too
soon, never too late, from the towers of the Chapel of the Lyceum, an
old church of the Jesuits, to snatch from me, or from any one who
takes my place, the bits of meat that we give to the gulls.
“Last year a dwarf hen which belonged to me chased from its nest
a female pigeon which had been setting for two days, broke up the
eggs, and laid one of its own in the nest. The pair of pigeons contin-
ued to care for the egg of the hen, and, at the end of twenty-one days
(which was really twenty-three for the pigeons) the chick came out of
the shell. To see the efforts of the parents to feed it was curious. The
second day, seeing that their efforts were in vain, I gave it some moist-
ened bread, then I put it under the pigeon; so matters went on for
three days, but the chicken wished to run about and I was obliged to
take it from its adopted parents.”—Bull. Soc. Zool., No. 4, 1892.
A Nest Building Frog.—In your issue for May, 1889, page
383, you published a paper in reference to certain batrachian nests
discovered by me at Nikko in Japan. This summer I was shown by
Dr. Guenther, at the British Museum, a couple of similar structures,
though very much smaller in size, preserved in alcohol, and which had
been received by the Museum from Japan. One of them had been
taken from a shrub growing in the mouth of a well. Dr. Guenther
told me that this nest is referable to a species of Polypedates. Day
before yesterday I received a letter from my friend, Dr. A. C. Good,
who is at present conducting a series of explorations in German West
Africa. I take the liberty of transcribing a portion of the letter as
follows:
“ I desire to write you of something I saw on my last trip. As we
brushed against the bush, that frequently overhung our path, I several
times noticed, now on my shoe, now on my knee, a white froth. I
thought it belonged to some insect, but for a long time I only noticed
the white foam-like substance when I had gotten past the bush from
which I had brushed it.
“ At length, however, I brushed off a large bunch of substance, and
when I tried to brush it from my clothes I uncovered some small crea-
tures which wiggled about in it and evidently made this froth-like
matter their home. On closer examination I discovered, very much
to my surprise, that they were tadpoles. ;
« Later on I found on the underside of a leaf, a mass of this white
substance that had not been disturbed since it had been placed there
by the mother frog. I take it for granted that these tadpoles produce
68 The American Naturalist. [January,
tree-frogs. The nest was about three inches long, by two or two and
one-half inches wide, and nearly an inch in depth. In the inside and
at the edges the frothy mass was quite white, and in consistency resem-
bled the white of an egg after being thoroughly beaten. The lower
face of the nest had taken on a yellowish tinge from long exposure.
In this I found eggs, or semi-transparent jelly-like bodies as large as a
small pea, which had already some power of motion, and on a few of
which the tail was just beginning to take form. In another nest I
found similar eggs just developing and also well-developed tadpoles.
These last were about one-eighth of an inch in length and had tails
one-half of an inch long. They seemed to move with difficulty
through the mass.
“I wonder whether this froth is at once home and food to them, but
am unable to say. These nests are frequent everywhere except near
the coast. I saw none nearer than ten miles from the beach.
“I remember your saying something about finding tadpoles in trees
in Japan, and I have the impression that you published something
on the subject. If so this will be of interest to you.”
Whether the creatures, the young of which Dr. Good found in this
frothy mass were the tadpoles of Chiromantis guineénsis, to which a
similar habit is ascribed by Bucholtz, of course I cannot say.—W. J.
Hortan, Chancellor Western University of Pennsylvania.
Horse “ Human Nature.’’—My son writes from a Wyoming
ranch that a blind bay mare is ostentatiously protected by a black —
mare, the two having been raised together.
The blind horse would suffer greatly for feed, water, and from herd —
interference if the black were not constantly on guard. The latter
watches the bay and grazes in a circle about her, keeping other horses 2
at a distance by kicks and bites if necessary, selecting good grazing =
spots and watering places.
The guardian is rewarded with occasional kicks and other human-
like evidences of gratitude —S. V. CLEVENGER.
1893.] Archeology and Ethnology. 69
ARCH ZXZOLOGY AND ETHNOLOGY.
* Legendary Evolution of the Navajo Indians.'—The Nav-
ajos once lived in a world below this earth. The tribe had twelve
chiefs, and the chief had four wives. This head chief arose early in
the morning and commanded his people to go to work. One morning
he failed to arise. The third morning he failed to arise. The fourth
morning he made no appearance. On the fifth morning the Navajos
became uneasy, and went to find their chief. The other eleven chiefs
wondered what became of him, and when they found him they learned
that. his oldest squaw had left him, and had married another man.
The old chief grieved very much and refused to be comforted.
In a short time the squaw came in and said, “I have left you
because I have ceased to love you. I can make my own living, and
you can make yours.” So they had a row.
This woman was chief of the women of the tribe.
Then the squaw called all of the women to council and said, “ Let
us part from the men!” So the women said, “ Take all the men, boys,
and male babies and cross the large river... Burn logs out to make
the canoes, and stay over there four years.” ;
They gave the male children into the hands of some hermaphro-
dites, who raised them on the brains of wild sheep and deer.
The men sailed across the large river, planted large fields of corn,
and raised immense crops. The first year the women raised a fair
crop; the second year they raised less ; the third year they had hardly
enough to eat; and the fourth year was a complete failure.
The women became discontented, and were in a starving condition.
Some of them ran into the river and were drowned.
1These legends were collected by the writer while employed in the Indian service at
the Navajo Indian agency. They were related by Tsé-di-dhl-ha’-in-be-géh, or
Rocking Sun, the great Lightning Medicine-Man of the Navajos. These legends
were carefully interpreted, and are given word for word as they were related.
2The “ Happy Hunting-grounds” of the Navajo Indians are represented as a land
full of forests and lakes which abound in various kinds of game and fish. Flowing
through the center of this land is a huge river which separates the braves and pretty
maidens from the inferior members of the tribe and the old women.
70 The American Naturalist. [Junuary,
So the women begged the men to come back to them. The woman-
chief admitted that she had done wrong, that the women could not
make their living, and that the men could not make their living.
Then the young chiefs held a council and said, “ Let us go back to
the women in four days, or they will die.” Soin four days they went
back to the women, and had a feast of deer meat and love-making.
While they were having a good time the Coyote picked a young
whale out of the water and hid it under his blanket. On the fourth
morning after this, when they awoke, they saw a large blue wave of
something coming fromthe east. The old chief sent an Indian to see
what it was. The Indian returned and said that it was water. They
looked to the north and saw a big white wave coming. The chief
sent an Indian to see what it was. The Indian returned and said, “ It
is water.” They looked to the west and saw a black wave coming.
The chief sent an Indian to see what it was. The Indian returned
and said, “It is water.” Then they looked to the south and saw a
green wave coming. The chief sent an Indian to see what it was.
The Indian returned and said, “It is water.’ Then the chief called
the tribe to council and said, “Something is wrong, we all will be
drowned.”
At that time the Navajos were animals and had squirrel blood in
them. So the White Squirrel planted a pine tree; the Gray Squirrel
planted a rattoon tree; and the Turkey planted a pipe-stem reed.
The Navajos all ran into this reed and began climbing up on the
inside. The reed grew very fast, even faster than they climbed. The
water began rising higher and higher, and followed close to their heels.
The Coyote was among them. The Badger went up first, making way
for the rest with his paws. The Badger consoled the rest by saying,
“Iam very near the top of the earth.” In getting his feet muddy
his legs and feet have remained black from that time. The Turkey
came up last, and the foam of the water touching its tail caused its
tail-feathers to be tipped with white from that time. :
Finally they came up through a lake, and they knew they had
reached the top of the earth.
The Badger looked out and said, “ I see a big Water Animal and some
Big Men who are very mean.” Then they sent the Locust out to see
what kind of an earth this was. A big White Bird came from the
north, met the Locust and said, “ Things like you are not to be seen
here!” Then the Locust replied, “ We will see about this.” A Yellow
Bird came from the west, a Black Bird came from the south, and a Blue
Bird came from the east, and they all said, ,“ Things like you are not
1893,] Archeology and Ethnology: 71
to be seen here.” But the Locust said, “ We will see about that.
If you will do as I do you may have this land; but if you do not do
as I do I will beat you.” The Locust had two arrows. He stuck one
of them up through his body and the other one down his mouth.
Then he took the two arrows and crossed them through his heart.
He next threw the arrows at the White Bird and said, “ If you do not
do as I have done I will beat you.” The White Bird took the arrows
and pretended to do these things, but he only ran the arrows through
his feathers.
There was so much water that the Locust could not bring his com-
panions up out of the reed. So he took a mountain-sheep’s horn and
broke the land to the north and to the south and to the west and to the
east, and the water all ran off. The Locust then went back and
brought his companions out of the hole which the Badger had made.
But the water still followed them up through this hole.
Then the chief said, “ Some one has been playing a trick.” He said
to the Coyote, “ You are always up to some meanness! What have you
under your blanket?” The Coyote opened his blanket, showed the
young whale to the chief, and then dropped it down this hole. The
water immediately went back down the reed into the river. They all
came out, but could not walk because of so much mud. Then the
chief prayed to the wind, and the wind dried the mud.
The Navajos were now changed to people, but they did not know
what to plant. The Turkey flew up, and the first time he dropped some
yellow corn ; the second time he dropped some red corn ; the third time
he dropped blue corn; and the fourth time he dropped all kinds of
corn.
The Navajos then made hé’-gins (houses), and the women and
children played in them while the men worked. Some of them made
houses in the rocks.
The chief then said, “ We will see if there will be any deaths up in
this world. I will throw a big log into the water, and if it sinks, we
will each one have to die; but if the log floats, we will never die.”
Then the Coyote tied a string to a rock and said, “I will throw
this into the water, and if it sinks we will each have to die, but if it
comes up and floats, we will never die.”
The chief then said to the Coyote, “ You are always doing some mean
trick!”
But the Coyote said, “I cannot help it. If the Navajos never die,
we will always be the same; but if the Navajos die, we will all be
72 *The American Naturalist. [January,
different. We all have children, and if none ever die, this earth will |
not hold us.” a 4
On the fourth morning, one of their number died. They all looked |
for this one, but they could not find him. Then they looked down this
hole which they came out of a few days before, and they saw this man :
down there combing his hair. This man looked up and said: a
“ I am happy down here. In time, you all will be down here where
Iam.” Then there was a famine, and about half of them died.
:
|
Tue MYSTERIOUS MAIDEN.
There was a little girl found at daylight one morning. The woman
who found her claimed that she was the “ Mysterious Maiden,” andso
another woman took her and raised her. This child soon grew to
womanhood.
This maiden conceived from a piece of petrified wood and borea —
Giant. She conceived from a feather and bore a Large Bird. Then she i
conceived from a horn and bore a Large Animal (something like a but —
falo), which ate the Navajos. She next conceived from a berry and
bore a Bear. She then rubbed herself against a rock. Behind this —
rock wasa patch of berries. When the Navajos went to gather the
berries, the rocks would crash together and kill them. The sides of _
the rocks were covered with blood. The maiden then conceived from
a reed and bore a patch of reeds. If a Navajo went into these reeds, —
he never was seen again. She next conceived from a battle-axe, and
bore an Old Hag who lived among the rocks, This Old Hag would moan —
and cry for some one to come and kill her. When a Navajo went to —
kill her, she would blow on the battle-axe and the axe would kill the A
Navajo. She then conceived from a hair, and bore an Animal Whose
Hair Grew Fast tothe Rocks. This Animal stood on the brow ofa preci- _
pice. Over in a corner of his den were some beautiful arrows. This
Animal would tell the Navajos to come and get those arrows, but when
they went there he would kick them over this precipice, and his chil-
dren, who lived down below him, would devour them. Next, this
maiden conceived from the sand and bore a pair of Big Eyes. Atnight, —
those Eyes shone like a big fire, and they would hollow for people to
come over there. Then the Eyes would pierce their hearts and kill
them. She next conceived from an antelope-hoof and bore Twelve
Antelopes, who used to destroy Navajos. Lastly, she conceived and
bore two sons. The oldest boy was conceived from the sun, and the
younger one from the water. These boys were going to kill all of
1893.] Archeology and Ethnology. 73
these animals which destroyed the Navajos, but the Navajos were
nearly all killed before this time. These boys grew up to be very
large, had bows and arrows, and they used to run off. One day these
boys asked their mother who their father was. She replied. “ The
cactus and the water.” But the oldest boy said, “I do not believe
this.” Then the mother said, “ The sun is your father, but he lives a
long way off.”
ORIGIN oF THE YAY-Br-Cuys.
The father of the Red Yaybichy was the sun. The father of the
White Yaybichy was the water. The Mysterious Maiden‘ conceived
from the sun and bore the Red Yaybichy. She conceived from the
water and bore the White Yaybichy.
This Mysterious Maiden was out picking up wood, and was going to
put it on her back, when the sun came up to her, dressed in turquois,
beads, feathers, and fine skins. He told this maiden to be by herself
that night, and he would come to her.
The Mysterious Maiden went home and told her father what the sun
had said. The sun came and talked with her, but she did not know
it; but she heard a noise going out from the hogan (house) where she
stayed. She saw this man (the sun) four days afterward, and told her
father that this was the same man she saw while picking up the wood.
She saw the sun abusing himself at daylight, and this made fleas and
mosquitos.
In four days, these two sons were born to the Mysterious Maiden,
and in four days more, these sons went up to visit their father.
The younger son had a cedar bow, and the older son had a piñon
bow. They started toward the east to see their father.
The Black Yaybichy met them there and told them to go back. He
told them that there were oceans and cañons and deserts and cactus fields
and great fires and great wolves and great snakes and great bears that
would destroy them, and said, “ Your father lives a long way off.”
3For a description of the Yaybichy Dance of the Navajo Indians see pages 435-436
of the Annual Report of the Bureau of Ethnology for 1883-84, by Dr. Washington
Mathews, U. 5. A., under the direction of Major J. W»? Powell, director of U. S.
Geological Survey.
_ The Yaybichy medicine-men are the leading medicine-men of the Navajo tribe,
and play an important part in all their religious ceremonies and fetichistic mysteries.
4The same maiden referred to in the Legend of the Mysterious Maiden.
74 The American Naturalist. [January,
These boys’ (the Red and White Yaybichys) went by a large ocean,
and looked down into the valley, and saw the smoke coming out of the
ground. Here lived an old Woman-spider. When the boys came
up the Spider said, “ Hallo, grand-children, where did you come from?
People of your class never come here! This place is not for you!”
“Our mother told us to go to see our father, the sun,” said the boys.
“But your father lives a long way off, and he is not a good man.
He will kill you with sweat-houses and red-hot irons.”
Then this old Woman-spider gave them each a white feather, and
told them it would be a Spirit to guide and defend them. Then she
said, “Stop here to-night with me.”
The boys said, “ We can not get through that hole in the ground.”
Then the old Woman-spider blew into the hole and it became larger.
She then vomited, and gave them (the boys) their suppers.
The sun was now straight over their heads. The boys told the old
Woman-spider that they wanted to get as far as they could before sun-
down.
* The old Woman-spider was a spirit ; so she pulled the sun down with
a net and then told the boys that it was now sundown. The boys
stayed all night, and they grew to manhood during that night.
The Black Yaybichy met them again, and told them that they would
reach their destination about noon that day, and that their father would
come to them at that place at night. `
At noon that day they saw a big house and started to goin, Two
big bears met them and snarled, but the boys said, “ We are going to —
to see our father.” Then the bears lay down and the boys passed over
them. They next met two large, vicious snakes. The snakes rattled
and hissed, but the boys said, “ We are going to see our father.” Then
the snakes lay down and the boys passed over them. They next met
two big lightnings and thunders. These stopped the boys, but the
_ boys said, “ We are going to see our father.” Then the lightning lay
down and the boys passed over it. They next met a number of little
snakes of various kinds. The boys said, “We are going to see our —
father.” Then the snakes lay down and the boys passed over them.
They next met the sun’s young wife. The boys said to her, “ We are
going to see our father.” The young wife replied, “ What are you
doing here? Boys like you and people of your class are not allowed
aoe here.” But the boys replied, “ The sun is our father.” Then the young
7 -~ wife wrapped them up in a white cloud that pointed toward the north.
__ She also made a black cloud that pointed toward the east; a yellow
~ “The boys referred to in the Legend of the Mysterious Maiden.
1893.] Archeology and Ethnology. 75
cloud that pointed toward the south, and a red cloud that pointed to-
ward the west. It was now pretty near night (sundown).
The sun had two children by this young wife, a girl and a boy.
The boy spoke and said, “ I hear my father coming home, for I hear
the white gourd rattle.”
The girl spoke and said, “ My father is coming, for I hear the blue
gourd rattle.” Then the boy said, “ My father is coming home, for I
hear the ivory gourd rattle.” The girl spoke and said, “ My father is
coming close, for I hear the turquois gourd rattle.”
At that moment the father came, making a fearful noise rattling the
irons on his body.
“The sun demanded of his young wife who those two young
men were that he saw come into the house, but did not see go out
again.
The wife replied, “You think you are pretty cunning. You told
me that you had no wife but me. These young men claim to be your
sons.” .
At this the sun became angry and rattled his gourds, and the earth
began to tremble, the lightning flashed, the bears roared, and the
snakes rattled and hissed. Thesun then demanded where the two
young men were, but the wife made no answer. He demanded this
again, but no answer. He then demanded the fourth time, but still
the wife refused to answer. Then the sun went to the cloud in the
east, and knocked that down; but no one fell out of it. He went to
the cloud in the west, and knocked that down; but no one fell out of
it. He went to the cloud in the south, and knocked that down; but
no one fell out of it. He went to the cloud in the north and knocked
that down, and the boys fell out of it and stood before him.
All at once four sharp irons, corresponding to the four clouds,
pointed toward the boys. There was a white iron from the north, a
black iron from the east, a yellow iron from the south, and a red one
from the west. The sun threw the boys violently against these irons,
the north one first, then the east one, then the south one, and then the
west one, but this white feather which the old Woman-spider gaye them
would let them down easy; so they remained unhurt.
The sun became angry and said, “I will find out if you are my
children. If you withstand my test you are my children.” Then a
spirit descended and stood on each of their ears, and told the boys how
to answer the sun’s questions. It said, “ Tell him he is your father.”
Then the sun took a huge turquois hammer and tried to mash the boys,
but the feathers made the turquois hammer come down easy. The sun
76 The American Naturalist. [January,
then made the boys smoke some poison in first, a turquois pipe, and
second, in an ivory pipe. He did this the second time, apd still the
boys were unhurt.
The sun said to his servants “Make a sweat-house and put four
irons in it, one of the irons shall be white, one blue, one yellow, and
one black, and make the house boiling hot.”
Just as the boys started to go into the sweat-house a Gopher came
up through the ground and told the boys to crawl into his hole. The
hole was inside of the sweat-house.®
The Gopher said “If you stay in there the sun will throw water on
the irons and the irons will break and kill you.” So the boys went
into the Gopher’s hole. The Gopher then said, “If your father asks
you if you are warm you go out of the hole and say, yes. You can
thus fool him. He will throw the water, but you will be safe in this
hole. He will then be through with you.”
The sun placed a blanket over the mouth of the sweat-house ‘sil
did as the Gopher had said. When the sun looked in he saw the boys
sitting there unhurt. Then he kissed the boys and told them that
they were his sons, and that they had gone through with all of the
forms that could kill them. The sun then took the boys home with
him and made his other son and daughter shake hands with them.
The young wife was then in a good humor, and dressed up her step-
sons. One of them she painted red, with white streaks down his back,
representing the lightning; the other one she painted white.
The sun then asked the boys what they wanted as a gift. The Spirits
on their ears said, “do not answer him until he asks you another
question.” The father took them through a large iron gate to the east
and showed them fine horses of all colors. The father said, “ Boys,
do you want these?” The Spirit said, “Tell him no.” Then he
opened a large iron gate to the north and showed them some fine
sheep, and said, “Do you want these?” The Spirit said, “ Tell him
no.” He next opened a large iron gate to the west and showed them
some fine goats, and said, “Do you want these?” ‘The Spirit said,
“Tell him no.” He then opened a large iron gate to the south and —
‘The Navajo Indians have sweat-houses at the present day. The house is made in —
a hemispherical form. Its first roof consists of poles, the second one is stones, and
the third one is dirt. A hole is left in one side for ingress and egress. The house
is usually located in close proximity to some stream or pond, and is used for medical
_ purposes. Rocks are heated and thrown into the sweat-house and water is throws
on n the heated rocks, causing steam to fill the apartment. The patient now goes into —
covers the door with a blanket. After a time he comes out and i
a pga into the adjoining lake or river. This process is often repeated in winter.
1893. ] Archeology and Ethnology. 77
showed them deer, buffalo, antelope and all kinds of game, and said,
“Do you want these?” The Spirit said, “Tell him no.” The father
then brought the boys home, and said, “ My children, what can I do
for you?”
The boys looked and saw four lightning arrows and a huge bow hang-
ing on the wall. The spirit said, “Tell him that you want these; that
some animals, a Huge Giant, Twelve Antelope, a Huge Bird, an Animal
Whose Hair Grew Fast to the Rock, are eating all the Navajos. Those
arrows will kill them.”
The sun replied, “ The Giant’ is my son and your brother, but if he
is eating the Navajos you have my permission to kill him. He has
no right to live. I will take you and go to-morrow morning.”
On the following morning they started, and came to Sanmateo
Mountain® about noon. The sun said, “ Boys, where did you start
from?” The Spirit said, “ Tell him that it was from here ; that here is
where the Big Giant was.” Then the sun let the lightning down, and
the boys climbed down the lightning to a big spring at the foot of the
mountains.
The Giant drank the water from the spring, and then lay down on a
rock to rest. He did this the second time and the third time and the
fourth time. As he turned his face toward the north the boys saw him
lying on the rock. The Giant turned his face toward the east, and
they could see his shoulders. He turned his face to the south, and
they could see his waist. He turned his face to the west, and they
could see his whole body.
The Giant now saw the boys and said, “ Will not they make fine
eating?” The Spirit said, “ Tell him that he will make fine eating.”
The giant then flew in a rage and threw an iron boomerang at the
head of the older boy. The Spirit said, “Stoop low, for he is throwing
at your head.” The Giant then threw one at his middle; the Spirit
said, “Jump to the right or he will hit you.” He next threw one
very low, and the Spirit said, “ Jump high, for he is going to throw it
low.” .
The sun then appeared and said, “ He is my son and I will have the
first chance at him.” Then the sun struck him with the lightning.
The Giant fell to the earth and grew weak because he lost his blood.
Navajo mythology is so replete. Dr. Mathews refers to it several times in his
« Mountain Chant,” found in the annual report of the Bureau of Ethnology, 1883-
84, J. W. Powell, Director.
78 The American Naturalist. [January,
The black rocks (igneous rocks) are his blood, and the petrified
wood is his bones. The Spirit then said, “Do not let his blood run
together, or he will get up again.” His blood ran down the hill.
The boys then shot him with the four arrows which their father had
given them, and killed him. The younger boy picked up the iron
boomerangs and kept them. The younger boy was given the turquois
gourd from his father.
They then saw these animals—the Buffalo, the Twelve Antelope, the
Large Bird, and the Animal Whose Hair Grew Fast to the Rocks over
in a little valley. They shook this turquois gourd at these animals
and the animals all died.
The boys then went to the Cafion de Shelley’ and went into one of
the cliff-dwellings, known as the “White House,” and disappeared, for-
ever to remain as Yaybichys.
THE MISSION OF THE YAYBICHYS.
A man was once struck by lightning and knocked all to pieces.
The Yaybichys came and sang over him and brought him to life again.
The White Yaybichy was the first one who came. This one went over
his body from east to west, from west to east, from north to south, and
from south to north, and had four songs. This one picked up his
meat.
The Black Yaybichy did the same as the white one. ;
The Red Yaybichy came, and when the man came partly to life he
came from the east, and had a gourd in his hand, and made a noise —
like lightning. He came from the south and made another queer noise.
Then he came from the west, and then from the north, and shook the
gourd” over the dead man’s head.
The gourd represents the noise of the lightning when it strikes a
person. ;
The White Yabichy took the man home, after he got alive, and —
showed him all these medicine things, and how he worshiped them.
*Pronounced de Shay. It is a beautiful little cañon situated about fifty miles from
Fort Defiance, Arizona Territory. It contains many cliff dwelli ngs, among which is
2 the one known as the “ White House,” (because of its whitened walls) which is vis-
ited by numerous adventurers every summer. Most of the walls remain at the pres-
ent day. There are now twenty-six Yaybichys in the Navajo tribe, including the
_ $un and these two boys. :
~ “The turquois gourd referred to in the Origin of the Yaybichys, The medicine-
-men of to-day seem to have unlimited faith in the turquois gourd,
1893.] Archeoloyg and Ethnology. 79
This Yaybichy took him to Sanmateo Mountain and told him these
things. He told him that a thousand years from this time the people
would follow the teachings of the Yaybichys, that his son and his son’s
son through a period of a thousand years would be able to bring peo-
ple back to life.
Then the Red Yaybichy took this man to Sanmateo Mountain and
shook the gourd over him and told him how to wave the gourd over
the dead man, from east to west (from sunrise to sunset), and from
north to south. Then this man came back into the Navajo tribe and
showed them how to use the medicine things,” and he was a great
_ medicine-man.
But when this man came back and the Navajos broke their arms
and legs, then they used these same medicine things, and they got well.
So that made the Navajos have confidence in the medicine-man and
the medicine things.
When any one gets sick we rattle the gourd over him and he gets
well.
These two Yaybichys, the red one, called Ya-na’ Yä-zän, and the
white one called T6-wizh-zhiis-chi-ni, were made on the top of Sanma-
teo Mountain.
There were some animals that ate the Navajos at that time, viz., a
Bear, a Large Bird, a Huge Giant, and a Fierce Animal Whose Hair
grew Fast to the Rocks, and which coaxed the people to pass that way,
when he would kick them down over the rocks and then go down and
eat them up.
There was another fierce animal which chased the Navajos, killed
them, and devoured them.
The Red Yaybichy killed all these animals” off.
' There was a sister who had twelve brothers. This sister became a
bear and killed the twelve brothers. Then the White Yaybichy killed
this sister.
The father of the Red Yaybichy was the sun. The father of the
White Yaybichy was the water—T. STANTON Van VLEET.
The “ medicine things ” referred to are the things which are used by the Yay-
bichy medicine-men at the present day. They consist of five pieces, each one of
which performs a specific duty in restoring the health of the patient. The medicine-
men claim that these “ medicine things ” have been handed down from generation to
generation since the origin of the Yaybichys.
These are the animals to which the “ Mysterious Maiden” gave birth
seem to have a significant place in capes mythology, and find their way into a large
number of their legends.
80 The American Naturalist. [January,
PROCEEDINGS OF SCIENTIFIC SOCIETIES.
New York Academy of Sciences.—At the meeting of the
Biological Section, Nov. 14, Prof. H. F. Osborn was elected chairman,
and Bashford Dean secretary. The papers of the evening were:
_ Arthur Hollick, On Additions tothe Palzeobotany of the Cretaceous
of Staten Island. These include about forty species not previously
recorded from eastern North America, although in part described as -
occurring in the cretaceous of Greenland and in the Laramie. About
fifteen new species were recorded, representing Populus, Platanus,
Myrica, Kalmia, Acer and Williamsonia. The fossils were in the main
taken from fire-brick clay. H. F. Osborn, Report Upon a Collection —
of Mammals from the Cretaceous (Laramie). The multituberculates
Meniscoéssus and Ptilodus were assigned to the Plagiaulacidz, the for- _
mer a probable ancestor of Polymastodon. The relations of these —
mammals were shown to be closer to Puerco than to upper Jurassic
i Sake
Soe sgt SE
foe; a Sao a
forms. Arthur Willey, On the Significance of the Pituitary Body, —
suggesting from studies on Ascidians and Amphioxus a primitive mon- —
orhinic condition in vertebrates. The nasal sac of Petromyzon is of —
secondary nature, as shown by development (Dohrn) and nerve sup- —
ply, but the nose in the monorhinic ancestor of vertebrates was the —
pituitary body of existing forms, this being represented in Ascidia, a8 4
shown by Julin, by the sub-neural gland and its duct, and in Amphi- —
oxus by the so-called olfactory pit. The pituitary body is to the
lateral nares what the pineal body is to lateral eyes. a
Bashford Dean exhibited an entire Ctadodus, a unique specimen
recently collected in the Cleveland shales. The tail is for the first
time shown, and indicates historically the origin of the ray parts of
this organ in modern elasmobranchs. ; :
re
jot A. H. Van Vleet, of Peru, Secretary and Treasurer. The fok
lowing papers were read : i
Psychology a Science, Dr. D. R. Dungan; Evidences of two Pre :
morainie Glacial Movements, Prof. G. D. Sweezey ; Evolution of the
Loup Rivers, Dr. L. E. Hicks; Some Notes on the Fringillide
Nebraska, D. A. Haggard; The Myriapoda of Nebraska, F.C. Ken-
_ yon; The Canyon Flora of Northwest Nebraska, A. F. Woods; Notes
1893.] Proceedings of Scientific Societies. 81
on the Flora of the Black Hills of South Dakota, P. A. Rydberg;
Notes on Nebraska Phosphates, H. E. Fulmer; Some Notes on Min-
eral Water from Odell, Nebraska, Rosa Bouton ; Systems of Notation
in Numbers, Dr. H. E. Hitchcock ; The Flora of Long Pine Canyon,
Julius Conklin; The Flora of the Sand Hills, Roscoe Pound; A New
Miocene Rodent, Prof. E. H. Barbour; The Fishes of Nebraska, M. E.
O’Brien ; Descriptions of Some New Nebraska Orthoptera, L. Bruner ;
Catalogue of the Orthoptera of Nebraska, L. Bruner ; Notes on the
Composition of the Lincoln City Gas Supply, Prof. H. H. Nicholson ;
The Relationship of the Nebraska Flora to That of the Regions Fur-
ther West, H. S. Clason; The Erysiphee of Crete, W. H. Skinner ;
The Fresh-Water Algæ of Kearney County, Nebraska, Dr. H. Hape-
man; Some Mexican Lichens, Prof. T. A. Williams.
Boston Society of Natural History.—November 16.—The
following paper was read: The Origin of Drumlins, Mr. Warren
Upham ; Profs. Shaler and Davis also spoke on the Origin of Drum-
lins.
December 7.—The following papers were read : Some Indian Quar-
ries in Arkansas, Mr. Leon S. Griswold; Notes on a New Order of
Schizomycetes (Bacteria). Specimens were shown with both papers.
SAMUEL HENSHAW, Secretary.
The Biological Society of Washington.—November 19.—
The following communications were read: On Certain Minute (para-
sitie?) Bodies Within the Red Blood Corpuscles, Dr. Theobald Smith ;
The Topographical Relations of the Excretory Canals of Cestodes,
Dr. C. W. Stiles; A Walchia from New Mexico, Mr. David White ;
Some Entomological Factors in the Problem of Country Fences, Mr.
F. M. Webster ; Comparative Value of Plants in Determining Floral
Zones, Mr. F. V. Coville.
December 3.—The following communications were read: The
Cruise of the U.S. Fish Commission Steamer Albatross in Alaskan
Waters in 1892, Prof. B. W. Evermann; Some New Grasses, Dr.
George Vasey; On the Rediscovery of Certain Rare Plants, Mr. J.N.
Rose; Exhibition of a Complete Series of the Large American Ground
Squirrels of the Subgenus Otospermophilus, Dr. C. Hart Merriam ;
The Mathematics of Forest Growth, Dr. B. E. Fernow.
Freperic A. Lucas, Secretary.
- Anthropological Society of Washington.—November 15.—
The following papers were read: Singular Copper Objects from
6
82 The American Naturalist. [January,
Ancient Mounds in Ohio, Mr. Warren K. Moorhead; Geographic
Nomenclature of the District and Vicinity,a Symposium, Mr. James
Mooney, Prof. Lester F. Ward, Mr. W. H. Holmes, Mr. W. Hallet
Phillips, Mr. W. H. Babcock, and Dr. Frank Baker.
Weston FLINT, Secretary.
SCIENTIFIC NEWS.
Prof. John S. Newberry, Professor of Geology in Columbia Col-
lege, New York, died at New Haven, December7. He wasborn at Wind-
sor, Conn., in 1822, and was the descendant of an old and distinguished
Puritan family. He was graduated from Western Reserve College in
1846, and from Cleveland Medical College in 1848. After two years’
travel and study in Europe he established himself as a physician in
Cleveland. He returned to his scientific studies, which had long been
in abeyance, in 1855, when he accepted an appointment as acting
assistant surgeon in the army, and accompanied, as surgeon and geolo-
gist, the expedition under Lieutenant R. S. Williamson, U. S. A.
which explored the territory lying between San Francisco and the
Columbia River. In 1857-58 he was attached, in the same capacity,
to the expedition under Lieutenant J. C. Ives, U. S. A., which made
the first exploration of the Colorado River, one of the most important
of the western territory surveys. Dr. Newberry, in 1859, participated
in the exploration of the country bordering the upper Colorado an
San Juan Rivers. During the war of the rebellion Dr. Newberry was-
a member of the United States Sanitary Commission, and directed its
operations in the Valley of the Mississippi. In 1866 he was appointed
professor of geology in the School of Mines, Columbia College. In
1869 he was appointed head of the reorganized Ohio Geological Sur-
vey, and under his direction the work was vigorously pushed to com-
pletion.
Prof. Newberry had probably seen more of the United States from
a professional point of view than any other of our geologists. He
will be best known from his work on fossil plants and fossil fishes.
- He was especially conscientious in his comparisons of American with
European forms of extinct life that came under his observation. Per- _ :
sonally he was of a rather impetuous- temperament, whose strong
friendships were offset by a spice of irascibility without malice. He
will be greatly missed from his place in the scientific life of America.
ea
1893.] Scientific News. 83
Sir Richard Owen died December 18. He was born at Lancaster
on July 20,1804. He received his early education in his native town,
and at the age of 20 he began a medical course in the University of
Edinburgh. He completed his studies in London and Paris medical
schools. |
When 30 years old he was appointed to the chair of comparative
anatomy at St. Bartholomew’s Hospital, and two years later he suc-
ceeded Sir Charles Bell as Professor of Anatomy and Physiology in
the College of Surgeons. He held the latter place for twenty years,
leaving it only to take charge of the Department of Natural History
in the British Museum.
He had the Cross of the Legion of Honor, was a Chevalier of the
Prussian Order of Merit, and was one of the eight foreign associates
of the French Institute. He was created a commander of the Bath
in 1873, and subsequently was made K.C. B. Sir Richard Owen was
the successor of Cuvier as the leader in the progressive advance of the
science of comparative anatomy. He filled up many of the gaps
unavoidably left by the great Frenchman, which he was enabled to do
by the opening up of many parts of the world by British commercial
and colonial enterprise. His contributions to paleontology are even
more important, his researches having covered regions that Cuvier
could not in his day reach. , Antarctic paleontology was founded by
him, for South America, South Africa and Australia yielded their
treasures to him first of all. Besides being an accurate observer and
describer he was a good systematist, many of the current terms of
zoology having originated with him. In generalizations of a higher
grade he was not active; the doctrine of evolution having arrived
rather late to get that attention from him which its earlier advent
would have secured.
Owen was a tall and stalwart man of spare habit. He was charac-
terized by a mental and physical tenacity, which was exhibited in his
psychic structure in the great difficulty he experienced in changing
an opinion he had once formed. He was fond of diplomacy, and
could dissect an adversary crosswise of the grain in the most bland and
sympathetic manner imaginable.
Prof. J. T. Rothrock has resigned from the faculty of the Univer-
sity of Pennsylvania, and will occupy the position of Secretary of the
Forestry Commission of Pennsylvania.
What is an Acquired Character ?—A Correction.—In the
December number of THe Natura.ist, page 1010, occurs about as
84 The American Naturalist. [January,
unfortunate an error in proof-reading as could have been made. The
sentence “ I must confess my inability to see why this variation is not
qualitative as well,” is put in quotation marks, as if it were credited to
Weismann. The sentence is my own, and would doubtless be indig-
nantly repudiated by the great apostle of Neo-Darwinism, as it is flatly
Neo-Lamarckian in fact.
In explanation I will simply add that I did not read the proof.
The original manuscript in my possession is correct, although I am.
not positive that the typewritten copy forwarded to Tue NATURALIST
is not at fault—C. C. NUTTING.
1893.] . Record of North American Zoology. 85
RECORD OF NORTH AMERICAN ZOOLOGY.
(Continued from Vol. XX VI, p. 798.)
Grote, A. R.—[Validity of ] Halisidota trigona. Can. Ent., xxiii,
109, 1891.
Grote, A. R.—Note on Graphiphora Hubn. Can. Ent., xxiii, 101,
1891.
Grorg, A, R.—The male genitalia and the subdivisions of Agrotis.
Can. Ent., xxiii, 147, 1891.
GROTE, A. R.—Halisidota trigona. Can. Ent., xxiii, 201, 1891.—
Validity of.
Grote, A. R.—On Catocala flebilis and C. fratercula. Can. Ent.
xxiii, 281, 1891.
GROTE, A. R.—Agrotis subgothica. Can. Ent., xxiii, 202, 1891.—
Reply to Tutt.
GROTE, A. R.—An explanation. Can. Ent., xxiv, 17, 1892.—Reply
to certain criticisms of J. B. Smith, as to arrangement of Heterocera.
Grore, A. R.—Remarks on Prof. John B. Smith’s revision of the
genus Agrotis. Can. Ent., xxiii 45, 1891.
Howarp, L. O.—The larger corn stalk borer. Insect Life, iv, 95,
1891.— Diatrea saccharalis.
Hupson, G. H.—A new species of Cerura. Can. Ent., xxiii, 197,
1891.
Hutst, G. D.—Prof. J. B. Smith’s List of Lepidoptera. Can. Ent.,
xxiv, 74, 1892.
Hurst, G. D.—New species of Pyaalide. Can. Ent., xxiv, 59,
1892.—15 sp., Ocala and Chipeta, nn. gg.
Ke.uicort, D. 8.—Notes on two borers injurious to the Mountain
Ash.—Podoseiia syringe, Zeuzophora semifuneralis.
Kewuicort, D. 8.—Notes on the Ægeridæ of Central Ohio. Can.
Ent., xxiv, 42, 1892.
Keuuicorr, D. 8.—[Tortricid feeding on Silphium perfoliatum].
Can. Ent., xxiii, 218, 1891.
LINTNER, J. A—([An onion pest, Agrotis ypsilon]. Can. Ent.,
xxiii, 220, 1891.
LINTNER, J. A.—On the eye-spotted bud-moth ( Tmetocera ocellaria),
in Western New York. Can. Ent., xxiii, 231, 1891.
#
86 The American Naturalist. [January,
LINTNER, J. A.—On some of our Orgyias. Can. Ent., xxiii, 232
1891.
Lyman, H. H.—Pamphila manitoba (Scud.) and its varieties. 22
Rep. Ent. Socy. Ontario, 1891, 27. Can. Ent. xxiv, 57, 1892.
Morratt, J. A—A microscopical examination of an unexpanded
wing of Callosamia promethea. 22 An. Rep. Entom. Socy. Canada,
32, 1891
Morrart, J. A——Additions to the Canadian list of Microlepidop-
tera. Can. Ent., xxiii, 167, 1891.
Morratt, J. A.— Melitta phaeton [in Ontario]. Can. Ent., xxiv,
18, 1892.
Morratt, J. H.—Petrophora silaciata. Can. Ent., xxiv, 18, 1892.
—Distribution and varieties.
NeEuMOGEN, B.—About Pseudohazis and its variations. Can. Ent.,
xxiii, 145, 1891.
Osporn, H. and Gossarp, H. A.—The clover-seed caterpillar
(Grapholitha interstinctana Clem.). 22 Rep. Ent. Soc. Ontario, 74,
1891. Insect Life, iv, 56, 1891.
Ossory, H.—Asopia farinalis as a clover pest. Can. Ent., xxiii,
283, 1891.
PACKARD, A. S.—Notes on some points in the external structure and
phylogemy of Lepidopterous larve. Proc. B. S. N. H., xxv, 82, 1891.
P . H.—Aphidivorous habits of Teniseca tarquinius (Fabr.)
Grote. Can. Ent., xxviii, 66, 1892
Ritey, C. V.—A new herbarium pest. Insect Life, iv, 108, 1891.—
| Carphozera ptelearia n. g. and sp. Geometride.
Situ, J. B.—Contributions toward a monograph of the Noctuide
of Temperate North America.—Revision of Homohadena, Grote.
Proc. U. S. Nat. Mus., xiii, 397, 1891.—H. deserta nov.
> Sumra, J. B.—Contributions, ete., Revision of the species of Hadena
referable to Xylophasia and Lapai t c., p- 407, 1891.—X. cogitata,
alticola, nigrior, antennata, centralis, are new.
Smıra, J. B.—Remarks on the classification of the Lepidoptera.
Can. Ent., xxiii, 245, 1891.
Smiru, J. B.—Notes on blackberry borers and gall makers. Insect
Life, i lv, 27, 1891. 22 Rep. Ent. Soe. Ontario, 52, 1891. —Bembecia.
SMITE, J. B.—Limenitis arthemis, etc. Can. Ent., xxiii, 104, 1891.
Smita, J. B—The squash borer, Mellitia ‘cucurbite, and remedies
mae 22 Rep. Ent. Soc. Ontario, 55, 1891. Insect Life, iv, 30,
1893.] Record of North American Zoology. 87
SMITH, J. B.—Halisidota trigona Grt. Can. Ent., xxiii, 158, 1891.
Sairu, J. B.—Prof. J. B. Smith’s List of Lepidoptera. Can. Ent.,
xxiv, 103, 1892—Reply to H. G. Dyar.
TownsEnD, C. H. T.— Chilo saccharalis in New Mexico. Insect
Life, iv, 24,1891. 22 Rep. Ent. Soc. Ontario, 50, 1891.
Turr, J. W.—Agrotis subgothica Harr. Can. Ent., xxiii, 159, 1891.
Van Duzer, E. P.—List of the Macro-Lepidoptera of Buffalo and
vicinity. Bull. Buff. Soc. N. H., v. 105, 1891.
WEBSTER, F. M.—Some studies of the clover hay worm, Asopia
costalis. Insect Life, iv, 121, 1891.
‘Winn, A. F.—Some rare Lepidoptera taken near Montreal. Can.
Ent., xxiii, 96, 1891.
Wricut, W.G.—Getting butterfly eggs. Can. Ent., xxiv, 70, 1892.
DIPTERA.
Atwoop, W. B.—A note on remedies for the horn fly. 22 Rep. Ent.
Socy. Ontario, 83, 1891. Insect Life, iv, 68, 1891.
Curtis, C._—Oxwarble in the United States. Jour. Comp. Med.
and Veterin. Archives, June, 1891.
CoquitteTt, D. W.—Revision of the Bombylid genus Aphcebantus.
W. A. Scientist, vii, 254, 1891.
CoquittettT, D. W.—New Bombylide of the group Paracosomus.
W. A. Scientist, vii, 219.
Coquittett, D. W.—Revision of the Bombylid genus Epacmus
(Leptochilus). Can. Ent., xxiv, 9, 1892.—2 n. sp.
CoQUILLETT, D. W.—New Bombylid® from California. W. A.
Scientist, vii, 197, 1891.
GARMAN, H.—An undescribed larva from Mammoth Cave. Bull.
Essex Inst., xxiii, 136, 1891.—Near Sciara.
Ke .icort, D. 8.—Note on the horn fly in Ohio. Insect Life, iv,
35, 1891. 22 Rep. Ent. Soc. Ontario, 59, 1891.
LINTNER, J. A.—The pear midge (Diplosis pyrivora) in New York.
Can. Ent., xxiii, 223, 1891.
Smrra, J. B.—Notes on blackberry borers and gall makers. Insect
Life, iv, 27, 1891.—Cecidomyid.
Smita, J. B.—Habits of Volucella fasciator. Can. Ent., xxiii, 242,
1891.
Snow, W. A.—The moose-fly—a new Hematobia. 22 Rep. Ent.
Soc. Ontario, 96, 1891.
88 ; The American Naturalist. . (January.
TownsEnD, C. H. T.—Notes on North American Tachinide with
descriptions of new genera and species—-Paper V. Can. Ent., xxiv,p.
64, 1592.—8 n. sp.
Townsend, C. H. T.—A Tachinid bred from a chrysalis. Can. —
Ent., xxiii, 206, 1891.— Meigenia websterii nov.
TownsEnp, C. H. T.—Notes on North American Tachinide, with
descriptions of new genera and species—Paper V. Can. Entom. xxiv,
p. 76, 1892.—Clistomorpha, n. g
TownseEnD, C. H. A PATA of a Muscid bred from swine
dung, with notes on two Muscid ee Can. Ent., xxiii, 152, pk .
—Lists of Cordylura and Clei
Weep, H. E.—The natural Gi of the screw-worm. Can. Ent,
xxiii, 243, 1891.
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CONTENTS.
White River Epoch gorse le White Clays
Reg ecologie ws, General—
j 89 i of the Obio ion—G
a Apion E come m a
_ AMONG THE INVERTEBRATA,
, Benjamin Sharp
ANTS. C EA 98
ENEO O or CH. Zoology.—The Cercaria Stage of Amihai 1
perpen 105. —Fecundation’ of the Eggs of Clinus argentatus— —
EST. garir S RIVER, Preliminary Desëriptions New Fishes from the —
eae Fee BD OR RED (Llus-., | Neg ple oT Larynx of Batrachia—The ete
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HE Asc i kinta P ag Caste J- The Pedal Skeleton of the Dorking” ak
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ie a: fit a 34 ( bgalome arienk S wali Cothectben of Calsopters’ :
Sa A ae h Mountains of British Columbia—A
from the "Hig
~ 137 “liar Seed-Like Gase-Worm from the Gran
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TOES: of the. Ua! ited gaa Age, Operated Upon for Double Congenital Cataract
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AMERICAN NATURALIST
VoL. XX VIL. February, 1893. 314
JOINT FORMATION AMONG THE INVERTEBRATA.
By BENJAMIN SHARP.
z The following observations were prompted by a discussion
= on the subject of joints, at a meeting of the Academy of
Natural Sciences of Philadelphia held’ on November 92nd,
1892, when Prof. Cope considered some facts relating to his
theory of joint formation as seen among the vertebrata.
i As his theory has been known for some time, and an
extended consideration of it has recently been published’ I
merely quote as a thesis, a sentence published in 1878’;
“change of structure is seen to take place in accordance with
"the mechanical effect of three forms of motion, viz, by friction,
Pressure and strain.”
: It occurred to me that if his theory had a general application,
= Some additional proofs could be shown to exist among the
-= Invertebrates, where we have the action of muscular force
upon hard and resisting parts of the skeleton. Those which
“present the best study for this purpose, appeat to be the
a crustaceans, where we find an immense variety of articulations
_ im the body and in the limbs; highly complicated locked
joints, others allowing motion in but one plane, as well as
: __ ‘The Mechanical Causes of the development ‘of the Hard Parts of the
_ Mammalia, by E. D. Cope. Jour. of Morph. Vol. 1I, 1889, p- 137-290.
*The relation of animal motion to animal evolution, by E. D. Cope, Amer-
“e ican Naturalist, Vol. XII. 1878, p- 44-
i
90 ` The American Naturalist. [February,
loose joints, where the hard parts scarcely come in contact with
one another, and cases of degeneration of the hard parts, lead-
ing to total disappearance of a previously existing joint.
In the annelides, from which, there is no doubt, the arthro-
pod branch sprang, we find no deposit of inorganic salts in
the epidermis. (Ido not regard the hard, calcareous, and
other secretions of tubicolous annelides as entering into this
question, as they are merely protective coverings, not acted
upon by the museles of the body, nor is there even any mus-
cular connection with it.) The outer layer of the body is
generally of a horn-like character, adhering closely to the —
secretive cells of the epidermis, very flexible and thrown into —
folds by the vermicular motion of its possessor. In the —
leeches, the body consists of a flexible cylinder, made up of
two sets of muscles, an outer longitudinal cylinder and an —
inner cylinder of circular fibers, the contraction of which, —
causes the animal to increase in length, while shortening is.
effected by the contraction of the longitudinal layer. The —
external surface of the medicinal leech, for example (see fig. 1) a
- is thrown into a regular series 0
very fine folds, extending across the
longitudinal axis of the body
These folds do not correspond in-
OE are a S
to one somite. When the ne
shortens its length, these folds
closely eh eg when extension
takes place, the folds are flatten
spread open, although not wh lly
disappearing, as they are a fix
quantity so to speak. I be
1893.) Joint Formation Among the Invertebrata. 91
valleys between the folds, this being aided by some of the cir-
cular fibers which pass through the longitudinal sheath, and
find their attachment to the bases of the valleys.
Starting from this point, and supposing the regularity
of the folds to have become established from pre-existing
irregular folds by the regularity and stress of muscular action,
we can conceive that when deposits of calcareous matter took
place, rings similarly formed by a folding of a soft skin
would receive that deposit at the most prominent portion of
this fold, the convex face and not in the protected valleys, as
there would be more friction or pressure from external
causes, and no deposits would take place in the valleys them-
selves because they would not be subject to external friction,
and their continual flexion would prevent any such deposits.
Should such a deposit take place in the valleys, there would be
a stiffening of the whole surface which would defeat motion.
In fact, in the leech, the cuticle is already much thicker on
the crests of the folds than in the valleys.
In the more primitive crustacea, we find the animal made up
of ‘rings extending over the whole length of the body, similar
to the rings of the leech, save (fig. 2) that there is but one ring
i ‘to one somite, and instead of a per-
pendicular valley between the folds,
this valley has an inward and a
forward direction, allowing the ante-
s rior edge of a caudad ring to fit into
the posterior edge of a cephalad
ring.
In the higher crustacea, several of
the anterior rings have coalesced,
and form a solid shield which is
known as the carapace. This has
no doubt arisen by the lessening of
the action between the anterior rings
oe T . : became the more active propelling
kepama becca aed organ. As the action ceased forward
showing the action of the mus- the valleys came to rest, and became
cles. ‘exposed to friction and pressure, and
_ ular ball and socket joint, but with a flexible part of th
92 The American Naturalist. [February, 1
consequently a deposit of calcareous matter took place produc-
ing the stiffening aboye hinted at. 4
The formation of jointed appendages from parapodic pad-
dles of the annelides can be followed out in the same manner,
since the manner of mutual relation of the segments is the
same as in the case of the body segments. =
It has been stated that in the leech the folds do not corre-
spond in number to the somites of the body, while they do in :
the crustacea. Allannelides do not move by means of a mus- 2
cular system built upon the plan found in the leech. In ~ 4
many the circular layer has to a large extent disappeared, |
for the longitudino-circular plan is undoubtedly ante-annelit
dan. The movement of the free medusoid forms, and of the —
Ctenophora, is the result of a modified arrangement of this 4
lan. i
With the disappearance of the circular layer, we, find a 4
peculiar modification of the longitudinal layer. This layer
becomes broken up and the fibers act in moving the setæ,
which answer to limbs. In a segment of a setiferous i
annelid, we may observe that the longitudinal muscles of the i
somite in section at the position of the seta are arranged like
the letter “ V,” in the fork of which the seta lies, the fibers to
the left (anterior) pull the seta externally backward, those on —
the right (posterior) pull the seta forward. The introduction —
of the sete, the origin of which I do not here attempt to —
explain has no doubt been, together with the establishment of —
external segmentation, a strong factor in causing the breaking ©
up of the muscular tube into sections (myotomes), which by —
use and consequent increase have extended each arm of the
“ V” into the segment on each side, while the insertion of the
end of the seta has caused a break in the muscle by the for-
mation of an aponurosis. This gives us the peculiar disposition
of a myotome to extend arcoss the union of two somites.
If we examine the segments of the so-called abdomen of the
macrurous crustacea, as the lobster, we will find that the
anterior face of one abdominal ring is pulled into the posterior
orifice of the ring lying anterior to it, forming a kind of eer :
ET ey TE oe Y
Fehon
te eo gee ee Ta eet Ohare
integument with no calcareous deposit, folded upon itself,
1893.] Joint Formation Among the Invertebrata. 93
and acting physiologically as a tubular ligamentum teres. On
examining the different joints, we will find that commencing
at a fixed point, as at the base of the thorax, the movable
ring of the first abdominal somite is pulled into the fixed part.
Then the first abdominal somite becomes the fixed point for
the movable ring posterior to it, and so on, so that we find
that the rings proceed away from the thorax, each is pulled
into the opening of the one in advance. This is true of all
those forms where the abdomen is well formed, strong and an
active organ in the economy of the animal; when this organ,
the abdomen, ceases to be an active organ of motion ‘as in the
burrowing forms, as Callianassa, Gebia, some of the Squillide,
etc., or where it is folded upon the sternum of the thoracic
region, the muscles becoming weaker through disuse, the
rings are not subject to the powerful muscular strain, and they
as a rule overlap but little if at all, but lie so that the edge of
one ring rests upon the edge of another. In those forms
where degeneration of the abdomen has proceeded so far as
not to have even the usual deposit of calcareous matter, as in
the hermit crabs, there are simply indications of rings on the
abdomen, and this organ is but little more than a fleshy sae
containing some of the viscera, and supplied with a few mus-
cles which act together, with the form of the organ, to keep
the abdomen curled so that it may hold as a hook, the animal
within the molluscan shell which it habitually occupies.
In forms as highly developed as the cray-fish and the lob-
ster where the calcareous deposits are great, there have crept
in many modifications. One of the main features in the
articulations, of the rings of the abdomen is the “lock” or
hinge which allows no lateral movement in the tail whatever;
the lateral movement-is also prevented by the overlap of the
pleura. This lock consists of a rounded prominence on each
of the rings projecting forward from a posterior segment into a
socket in the anterior one. The movement in any two seg-
ments so formed takes place in the plane, the axis of which
passes through these two points so that the are described by
the movement is only in a vertical plane. The terga of the
segment is much arched in the lobster, and the upward
motion is limited after a certain point is reached by the dome
>
Which follows it so that it may be covered by the posterior
_and the surface of the ring.
to hold good viz.: that the movable part is pulled into the
_Ibacus, the first pair, and in fact all of the thoracic limbs end -
gestion of a chela. In Crangon, (fig. 3) on the other hand
OS me “hand? of & form of Cre
-o a “eda eof ie Increase of the muscular po
94 The American Naturalist. | [Februray,
of the segment itself. In some of the Scyllaridw, there isa
transverse groove on the terga of the somites merely indicated — ;
in the lobster, into which the posterior face of the terga ahead _
of it fits when the abdomen is fully extended, and which pre-
vents any further upward movement. It will be noticed
further, in those forms that the sternum of each abdominal |
segment is very narrow, and that they are connected by a _
broad uncalcified membranous area which allows plenty of 1
room for flexion downward. ;
The first abdominal segment is narrow, and in the vast |
majority of cases it is considerably narrower than the one —
ce
edge of the carapace and the anterior edge of the second
abdominal ring. In the Homaridea and Loricata there is an a
interesting provision to prevent the strain of the powerful
abdominal muscles from drawing the smaller segment too far
into the thorax. On each of the lateral faces of the ring there
is a broad tooth or calcified lobe, which overlaps the posterior —
edge of the carapace allowing the latter to play between it
When the limbs are examined, the same rule will be found
fixed part. A modification of this is well illustrated in the
evolution of the large chelæ. In some forms, take for example
in a sharp pointed segment, there being not the slightest sug-
the ternimal segment is pulled
against the broad face of thé
penultimate one thus making 4
shift for a chela. In the Stom-
~K ~ ` and forms with ita very effective
6 figs ~ grasping organ. The continual
| use of the terminal segment, the
1893.] Joint Formation Among the Invertebrata. 95
will tend to draw this terminal segment backward (into) on
the penultimate which enlarges with the increase of bulk of
muscle, so that a well developed chelæ, as in the lobster, is
found where the ultimate segment is pulled backward to
about the middle of the penultimate segment.
A true ball and socket joint rarely if ever obtains in the
crustacea, and for very obvious reasons. The nature of the
impassive connection of the joints would not allow of the
motion and the active element, the muscle acts upon the
inside of the joint or through its center, thus making it almost
impossible for muscles to act so as to give the free movement
of a ball and socket joint. The same end is obtained by a
series of ginglymoid joints working in different planes so that
in a series of three or four joints the end of the limb, for
example, can be made to describe a circle, and I am inclined
to believe that in order to obtain this free motion, the first two
or three joints of crustacean limbs are as a rule very short |
and work in different planes.
Several objections may be easily found to the statement
that posterior segments are always drawn into the anterior
ones. Take for example, the shrimps, where the second
abdominal ring overlaps the first so that the ring ahead of it
as well as the others behind it is drawn within its anterior
opening. I am inclined to regard this exception as more or
less proving the rule, and to consider the second segment the
fixed point into which the other two rings are drawn. We
see a foreshadowing of this plan in the lobster group, where,
as in Astacus, to use the words of Huxley, “the plure of the
second somite are much larger than any of the others, and,
their front edges overlap the small pleure of the first abdom-
inal somite, and when the abdomen is much flexed these
plure even ride over the posterior edges of the branchioste-
gites. ”* |
- An exception much more difficult to explain is that found
in some of the anomurous forms as Galathea, where the fourth, —
fifth, sixth and seventh segments follow the rule, and the first
The Crayfish, an introduction to the study of zoology, by T. H. Huxley,
New York and London, 1880, p. 98 and 99.
96 The American Naturalist. (February, — a
in regard to its anterior face, but the second overlaps the :
first, the third, the second, and the fourth. In other words,
the fourth has pulled both ways as the second has in the —
shrimps, but in the latter the second segment is obviously the
largest of the series, while the abdominal segments of Galathea
are all of about the same size and strength. A
We may, however, provisionally accept as a natural conse-
quence and general rule, that the motion of a movable part
naturally hollows out for itself a cavity in which to move —
within the solid and fixed basis. This may be further illus-
trated in the hard parts of the vertebrata.
It is a well known fact that even when the muscles are said
to be at rest, there is more or less of a strain, or pull exerted.
When the human hand hangs loosely down, it rests half way `
between extreme flexion and extreme extension ; the same may
be said of the fingers, although in some persons, as in sailors,
where the flexor muscles are very strong the fingers hang as s0
many “ hooks, ” owing to the pull of the more powerful flexor
muscles. This is well shown in cases of the paralysis of onesetof
muscles ; the parts are drawn to the side of the healthy muscles :
even when no voluntary action is exerted by the normal mus-
cles. This is too well known to medical men to require —
further comment, and is especially illustrated in cases of facial _
paralysis where the face is drawn strongly to one side by the |
normal muscles. 1
Taking the action of the normal muscles when at so-called
rest, it is not surprising to find that in the joints the lighter
or more mobile part, being continually pulled against the
fixed part and moved upon it, that a cavity should be
formed in the fixed parts by the continual action and pres-
sure of the mobile part. ‘Take for example, the humerus in
_ man, the head is hemispherical and fits into a concave portion —
of the scapula, a true ball and socket joint, and it is formed, if |
there be any truth in the idea, by the humerus being pulled `
into the scapula by the powerful muscles of the shoulder.
And as this is probably the first joint which was formed in the
=~ evolution of the limb which then acted as a fin, the rotatory
_ motion aided and finally established the ball and socket form
1893.] Joint Formation Among the Invertebrata. 97
of joint. Other cases might be mentioned as that of the lower
jaw and the tarso-tibial joint.
In the above remarks, I have simply made an attempt to
apply the theory of mechanical genesis to a much neglected
group of the invertebrata, and if I, in this, have only been able
to direct attention to the field of carcinology, I will feel that
this short paper will have served its purpose.
98 The American Naturalist. [February,
THE GASES IN LIVING PLANTS.
(Continued from Vol. XXVII, page 7.)
By J. C. ARTHUR.
Kinps oF GASES AND RELATIVE AMOUNTS.
It is.now our duty to give a brief statement of the kinds, `
the source and the movements of gases in plants. E
Plants are permeated by the same gases that make up the |
atmosphere surrounding them: oxygen, carbon dioxide and
nitrogen. Nitrogen in the form of a gas is neither used nor
generated by any part of plants, unless we except the tuber-
cles of certain roots, and so it occurs in about the same per- —
centage inside the plant as outside of it. On the other hand,
both oxygen and carbon dioxide enter into combination with,
and. are liberated from, the plant tissues in varying amounts
at different times. The percentage of these two gases in the —
cavities of the plant vary through a considerable range. Ina
series of determinations made by Lawes, Gilbert and Pugh, in ~
England, the oxygen ranged from 3 to 10 per cent., and the —
_ carbon dioxide from 14 to 21 per cent in plants which had —
been for some time in the dark, while plants which had been ©
standing in sunlight reversed these figures, and gave 24 to 27 _
per cent. of oxygen and 3 to 6 per cent. of carbon dioxide.
The two gases, therefore, bear a somewhat reciprocal relation,
their sum usually being about 25 to 30 per cent. of the total —
gas in the plant. K
a a eee a, ee et eee Spe
VARIATION IN AMOUNT DUE TO ASSIMILATION.
1893.] The Gases in Living Plants. 99
rapidity in healthy cells, but is entirely checked upon the
withdrawal of light, or when it reaches a certain low intensity.
Of course it never takes place in roots, flowers, the central
portion of large stems, or other parts which are not green, nor
in any fungi or other plants not possessed of green coloring
matter.
VARIATION IN AMOUNT DUE TO RESPIRATION.
The other great cause of disturbance in the relation of oxy-
gen and carbon dioxide in the plant, is the process of respira-
tion.
Respiration in plants is essentially the same as in animals,
and consists in the fixation of oxygen and the liberation of
carbon dioxide. It takes place in every living cell, whatever
the kind of plant, whatever the part of the plant, and what-
ever the conditions of active existence. The rate of respira-
tion varies with the temperature, the age of the cell, and the
nature of the chemical transformations. In normal respira-
tion the amount of oxygen absorbed is approximately the
same as the amount of carbon dioxide evolved. There are,
however, certain modified forms of respiration in which this
does not hold true.
If living plants be placed in a vacuum, or in an atmosphere
deprived of oxygen, it is found that they can still carry on
life processes for some time, accompanied with an evolution
of carbon dioxide. The oxygen necessary for this process is
obtained from the breaking up of compounds in the cells, and
it is therefore called intramolecular breathing. >
The germination of seeds, which contain a large amount of
oil, is somewhat the opposite of this last process. In order
to convert the fat into a more directly servicable food material
for the plant, a large amount of oxygen enters into the new
combination, for which there is no equivalent amount of gas
liberated. It consequently comes about that oily seeds in ger-
minating absorb a far larger amount of oxygen than they
liberate of carbon dioxide. This is known as vincular
breathing. -
100 The American Naturalist. t [Febina
Another variation from normal respiration is known as
insolar breathing, and which, with still some other modifica-
tions, I need not stop to explain. To this brief statement of
plant respiration must be added that much yet remains to be
discovered regarding the details of the processes.
Assimilation and respiration are the two great causes which
disturb the relative volume of the two variable gases in plants.
MOVEMENT OF THE SAME TWO GASES.
_ We shall now turn to the movement of the same two gases,
oxygen and carbon dioxide. There has never been a disposi-
tion, as in the case of many other plant phenomena, to explain
the movement of gases upon any other than purely physical
principles. We have therefore to do simply with the question
of the aids and hindrances to the establishment of an equilib-
rium between the gases inside and outside the plant, irrespect
ive of whether the cells are alive or dead. |
OUTER AND INNER PRESSURE OF GASES.
It has already been stated that the relative amounts
oxygen and carbon dioxide inside the plamt are usually very
different, and that within a few hours the relation of the two
_ may be completely reversed. To this may be added that the
pressure of the gases inside the plant is sometimes more, 80
times less than that of the atmosphere outside the plant
almost never the same. Hales observed ‘in his early wort
that a mercury guage connected with the inside of the trunk
of a tree showed an internal pressure when the hot rays ©
sun warmed the trunk. This was largely due, undoubted,
the expansion of the gasesin the trunk, by the heat. Suc
excess of pressure in water plants is very common, althoug
undoubtedly many have noticed in gathering water lilies,
other water plants.
an On the other hand, the pressure of the gas inside the
‘may be less than on the outside. This has long been
1893.] The Gases in Living Plants. 101
nized, but was best demonstrated by von Héhnel in 1879, to
whom it occurred to cut off stems under mercury. In doing so
the mercury rose to a considerable height in the vessels of the
stem, and as mercury is without capillarity, this can only be
ascribed to the greater pressure of the outside air, or in other
words, to a partial vacuum in the plant.
An observation was made by Hales, whom I have mentioned
so often, which we may use to illustrate how such a negative
pressure, as it has been called, can be brought about. He cut
off a branch, fastened an empty tube to the cut end, and
plunged the other end of the tube into aliquid. He found that
as evaporation of moisture from the leaves took place, the
liquid was drawn up into the empty tube. This phenomenon
can now be explained more satisfactorily than could be done
at that early day. By evaporation the liquid water inside the
plant escapes in the form of vapor, and the space it occupied is
filled by the gases, thus rarifying them. This rarifaction
may go on in uninjured plants until the internal pressure is
greatly reduced. But in the experiment, the pressure is equal-
ized by the rise of the liquid in the tube. A later modifica-
tion of Hales’ experiment is to use a forked branch, place the
cut end in water to give a continuous supply of moisture for
transpiration, and attach the empty tube to one of the side
forks of the stem, cut away for that purpose.
PERMEABILITY OF TISSUES.
It is self-evident that such condensation and rarifaction of
the gases in the plant could not take place if the cell walls
were readily permeable to gases. Thus it comes about that
one of the most important topics in connection with the move-
ment of gases in the plant, is the permeability of tissue walls
of various kinds, and especially those constituting the surface
covering of plants.
I shail not attempt to conduct you through the tangle of.
supposition and fact, errors in experiments, correct and incor-
rect conclusions, and the general confusion which has come
from the labors of physicists, chemists and botanists for the
last twenty-five years, during which the subject has received
104 = _ The American Naturalist.
coefficient of absorption and the density of the gas. Cuticular —
and corky formations also permit the passage of gases when.
dry. Thus we see, that gases may be forced through the
stomata, or breathing pores, by varying pressure, but can only
pass through the epidermis and bark of plants by diffusion.
We therefore arrive at the conclusion that the gases inside and
outside of the plant are brought to an equilibrium by direct
interchange through the stomata and intercellular spaces,
aided by the comparatively slow process of diffusion through — :
the whole surface of the plant, both above and below ground.
In CONCLUSION.
After so long a discussion of the subject of the origin, kind
and movement of gases in plants, I trust I have not only
brought out the main facts regarding our present knowledge,
but also made evident some of the numerous directions in —
which further experiment and research is needed. It would
be most appropriate to transfer the activity in this subject to
the laboratories of this country, and let the Americans take up
the line of discoveries carried on so far by the English, French,
and lastly, the German scientists:
1893.] Legends of the Sumiro-Accadians of Chaldea. 105
LEGENDS OF THE SUMIRO-ACCADIANS OF
CHALDEA.
By Atice BODINGTON.
(Continued from Vol. XXVII, p. 19.)
In Genesis, as is well known, two distinct accounts exist of
the creation of the world and of Man, of which variants are
found in the cuneiform inscriptions. One of these variants,
lately found on a tablet of baked clay, with a parallel Semitic
translation, is of great interest. It begins at an earlier period
of cosmic history than either the Biblical or the hitherto known
Chaldean account! Whilst the version of the first chapter of
Genesis’ begins with a description of chaos and the old Semitic
Babylonian version with the time “ when the heavens were not
proclaimed and the earth recorded not a name,” the Sumiro-
Accadian account begins with a description of the time when
the “ glorious house of the gods (apparently the sky,) had not
been made, a plant had not been brought forth nor a tree
created; when a brick had not been laid, a beam not shaped,
a house not built, a city not constructed and a glorious foun-
dation or dwelling of men had not been made.” But when
“within the sea there was a stream,” then “ the glorious city
of the gods, the divine Eridhu,” was built, of which Babylon,
the earthly Eridhu, was a faint copy. Then the tablet men-
tions the creation of living beings, not men as yet, but gods
and the spirits “Anunnaki” ; and the supreme deity proclaims
the existence of the “ glorious city, the seat of the joy of their
hearts.” Meridug, son of Ea, now “ made a foundation before
the waters”; made dust and poured it out with the blood,
and in one single line “he made mankind.” The female
principle, the goddess Araru, (the Bohu of Chaldean legend
the Bohu of Genesis 1.2) “ made the seed of mankind with him.”
1That deci ith.
*New Vasa a ope ence Paper read by Mr. T. G. Pinches,
of the British Museum, at the International Oriental Congress.
8
planets conceived of as resplendent animals of benef
106 The American Naturalist. [February,
Then he made the beasts of the field and the living creatures
of the desert; the Tigris and the Euphrates, and “ proclaimed
their name well,” in fact said that creation was good, as in
Genesis. Then Herodach created grass, the plants of the
marshes and the forests; and with the plains and forests in
which they were to dwell, he made oxen and other large cattle
and sheep. This deeply interesting account has been neces-
sarily much abbreviated.
The Accadian idea of the structure of the world must be
understood before the various legends as to the creation and
the early history of mankind become “thinkable” to the mod-
ern mind. In our mind’s eye we see the world as a compara-
tively insignificant planet whirling at headlong speed round a
central sun; we know the blue sky to be only the effect of
distant air; it is not “thinkable” to us that the sun should
stand still at the bidding of the leader of a small tribe of half
savage Nomads, nor that a tower should be built to attain a
solid sky, above which lived gods, alarmed and jealous at so
daring a proceeding. Nor can we picture to ourselves a great
reservoir or “Heavenly Ocean” above a solid sky, which
could be “ opened ” to let the waters submerge a sinful world.
But all these stories are in harmony with Accadian cosmogony- —
The Chaldeans imagined the earth as an inverted boat or bowl,
the thickness of which would represent what we call the crust
of the earth; while in the hollow beneath this crust lay the
_ “abyss,” the abode of many powers, and answering to |
Elysium and Tartarus of classic mythology. Here dwelt
Allat, the remorseless Queen of the Dead; from the Abyss
issued the terrible Maskim, the seven evil spirits, who knew
neither mercy nor pity. Here too was Eridhu, the “ glorio
abode,” within which was the tower which reached the skies. —
Above the convex surface of the earth spread the sky, (ana)
divided into two regions, the highest heaven or firmament,
which with the fixed stars immovably attached to it, revolved
_ as upon an axis, round an immensely high mountain, which
joined it to the earth as a pillar. In the lower heaven th
` 3Chaldea, pp. 153-157.
1893.] Legends of the Sumiro-Accadians of Chaldea. 107
nature, wandered forever on their appointed paths. The
earthly ocean, a counterpart of the heavenly ocean, was imag-
ined as-a broad river, or watery rim, flowing all round the
edge of the inverted bowl, precisely as Herodotus described it.
And so late as the Middle Ages, we find Dante’s conception of
the structure of the world to be substantially thatof the Accad-
ians; the “ purgatorio ” is an immensely high mountain join-
ing earth and sky, through whose circles pass the souls in a
state of probation, till at the top they step into the lower
heaven.
In the fine Accadian epic of which the account of the Deluge
furnishes the eleventh book, the ark is represented as resting
on this mountain. It relatés* that Tzdubar, the national hero,
was beloved by the goddess Tshtar. She promised him a “ char-
iot of gold and precious stones; that kings and princes should
bow before him, and kiss his feet; that his flocks and herds
should multiply two fold, and his mules and oxen be peerless
of their kind.” But Tzdubar laughed her love to scorn, and
the enraged goddess implored her father, Anu, to take ven-
geance on him. A monstrous bull was sent against his city of
Erech, but it was slain by Tzdubar’s friend, Eâbani, and laid
before the altar of the god Shamash, whilst the people spent
the night in feasting and rejoicing. But the vengeance of
Tshtar was not to be so easily foiled. With the help of her
mother she smote EAbani with sudden death and Tzdubar
with a dire disease, which made life a burden. He determined
in his anguish to seek relief from his great ancestor, Hasisadra,
who dwelt, immortal, in the Blessed Land, at the “ mouth of
the rivers” beyond the Waters of Death. Long and weary
was the journey, and on his way Tzdubar passed the giant
warders of the sun—half men and half scorpions—who kept
watch over his rising and his setting. On the shore of the
Waters of Death the hero met the ferryman Urubél, and for a
month and fifteen days they journeyed together over that
dreary sea, till they reached the Chaldean “ Valley of Avilon,”
and Tzdubar met his ancestor face to face. Here Hasisadra
related to his descendant the story of the Deluge, and his own
‘Chaldea, pp. 301-17.
108 The American Naturalist. [February,
share in that great event. Tzdubar was purified and healed
by laving in the Waters of Death ; he returned safely to Erech,
and there offered up a touching prayer to the beneficent god,
Ea, that his lost friend, Eabani, might be restored to life.
Ea, through his son, Meridug, brings Eâbani from the world
of Shades to the Land of the Blessed, there to live forever
among the heroes of old, and so all ends happily as a fairy
tale.
The Chaldean legend of the Deluge is, as is well known, an
extremely close variant of the account found in Genesis, even
to the rainbow asa sign of the repentance of the God Anu,
for the havoc he had wrought. Only it is Tshtar who “ spreads
out the great bows of her father Anu,” and who says “ I shall
be mindful of these days; never shall I lose the memory of
them,” and the assurance is given that though pestilence and
wild beasts may be sent as a punishment for the wickedness
of man, never shall a universal flood again overwhelm the
earth. In the Chaldean account it is the ever beneficent Ea
who warns Hasisadra of the coming flood, and bids him
prepare the ark for himself and his family, and who himself
sends the cattle and the wild beasts of the field to their
haven of safety; he who reproaches the other gods with the
wanton destruction they have wrought, and brought Bel,
hasty but quickly repentant to his senses. Bel himself took
Hasisadra by the hand and led him out of the ark, after
the sacrifice had been offered up, “ when the gods smelled a
sweet savour.” See Genesis viii, 21.
The great solar and catholic myths are found in their earliest
forms amongst the Sumiro-Accadians, and from them they
passed through the Pheenicians to the Greeks and Romans.
Tshtar, the great goddess of Nature, personifying the life-
producing earth, loves the young Sun-God, Dumuzi. The
tablet of the national Epic, which describes the manner of
his death, has unfortunately but one fragment left, which
speaks of the “ black pine of Eridhu, marking the centre of
the earth, in the. dark forest, into the heart whereof man
has not penetrated; within it Dumuzi. . . .” A month was
set apart, (June-July), both in Chaldea and Assyria, as
1893.] Legends of the Sumiro-Accadians of Chaldea. 109
mourning for the death of Dumuzi, however it occurred ;
the mourners wept and wailed, and tore their hair for the
first six days, and at the close of the sixth day the wildest
and most extravagant rejoicing marked his resurrection and
restoration to Tshtar. Six hundred years B. C., Ezekiel, in
captivity at Babylon, speaks of the “ women weeping for Tam-
muz,” (Dumuzi). We have, fortunately, the tablet, nearly
complete, which relates Tshtar’s descent to “the land whence
there is no return, towards the dwelling that has an entrance
but no exit, towards the hall from which the light of day is
shut out, where the shades of the dead dwell in the dark.”
Tshtar haughtily orders the warder to open the gate, “ If thou
_openest not, I will demolish the threshold. . . . I will
let loose the dead to return to earth. . . . I will make the
risen dead more monstrous than the living.” The gatekeeper
humbly answers the angry goddess: “ Be appeased, O Lady,
let me go and report thy name to Allat, the Queen.” And
Tshtar declares that she comes only to “weep over the heroes
who have lost their wives; over the wives who have been
taken from their husbands’ arms. I wish to weep over the
Only Son, (a name of Dumuzi,) who has been taken away be-
fore his time.” Allat, full of evil delight that a hated rival
has come within her power, orders the keeper to open the
seven-fold gates for Tshtar, with the stipulation that at every
gate she should strip off some of her attire. The warder of
Arallu takes from the goddess her earrings, her necklace, her
jewelled girdle, the bracelets on her arms, and the bangles at
her ankles, and lastly her long flowing garment, and with
these her divine power departed, and she stood powerless
before the spiteful Queen of the Dead. Allat orders her chief
minister, Namtar, the Pestilence, to lead Tshtar away, and to
afflict her with sixty dire diseases, in the deepest darkness of
the abyss. Meanwhile, as when Demeter mourned incon-
solably for her lost Persephone, all went ill in the upper world.
Life and love had gone out of it; there were no marriages
and no births, and the gods held council as to the release of
Tshtar. The beneficent Ea conceived a plan. He created a
phantom, Uddusunamir. “Go,” he said, “ to the Land whence
110 The American Naturalist. [February,
there is no return, and the seven gates of Arallu will open
before thee. . . . Conjure Allat with the name of the great
gods; stiffen thy neck and keep thy mind on the Spring of
Life. Let the Lady (Tshtar) gain access to the Spring of
Life, and drink of its waters.” Allat in her fury beat her
breast and bit her fingers with rage. She ordered Namtar to
let Tshtar drink of the Spring of Life, and bear her from her
sight. Namtar took the goddess through the seven enclosures,
restoring at each the article of her attire that had been taken
from her. At the last gate he said: “ Thou hast paid no ran-
som to Allat for thy deliverance; so now return to Dumuzi,
the lover of thy youth; sprinkle over him the sacred waters,
clothe him in splendid garments, adorn him with gems.”
The last lines of the poem are mutilated, but it is evident
that they bear on the reunion of Tshtar with her young lover.
Not only does this myth remind one of the legends of Deme-
ter and Persephone, of Adonis, of Balder, the beautiful, and of
Osiris, but what—if it were possible—seems like a variant of
the same myth, is the legend amongst the Tee-Wahn’ Indians
of Nah-chu-ru-chu (the Bluish Light of Dawn) and his lost
wife, the Moon-maiden. Nah-chu-ru-chu held the well-being
of all his people in his hands for life and death. When the
jealous Corn-maidens had thrown his wife down a deep well
where none could find her, Nah-chu-ru-chu sat for days,
neither speaking nor moving, his head bowed upon his hands.
Then no rain fell and the crops died, and thirsty animals wan-
dered, crying along the dry rivers. The coyote, the badger
and the eagle went to seek the lost Moon, and when at last she
was found, the choked earth drank and was glad and green,
the dead crops came to life, and for four days the people danced
and sang in the public square.
In innumerable ways have the Accadians been the priests
and schoolmasters of mankind in Europe and Asia, for through i
the Bak tribes of Elam, in Southern Chaldea, they are consid-
ered to have been the founders of the ancient civilization of
China; and through the Semitic ‘peoples they have conferred
upon us gifts, good and bad; the art of writing, the signs of
’Tee-Wahn Folk Stories. St. Nicholas, March, 1892.
1893.] Legends of the Sumiro-Accadians of Chaldea. 111
the zodiac, the measurement of the year, the early legends of
Genesis, the keeping sacred of the seventh day, the belief in
magic and witchcraft, and the medieval devil. An Assyrian
calendar mentions a day called Sabattu, “a day for completion
of work, of rest for the soul.” On that day it was not lawful
to cook food, to change one’s dress, to offer a sacrifice; the
king was forbidden to speak in public, to ride in a chariot,
or perform any civil or military duty”; a strictness equal to
the Sabbatarianism of the most orthodox Jew.
On their way to a pure monotheism, the early Jews seem to
have combined in their “ Elohim,” and their “ Yahveh,” the
characteristics of the principal gods of the Sumiro-Accadians.
We recognize the features, sometimes of the calm and benefi-
` cent Ea, and of his son Meridug, Intercessor for men; some-
times of the hasty, vindictive Sun-god, Bel, who “ consumes in
a moment” the victims of his anger, till the features of all
these Nature gods fade away in the moral effulgence of the
sublime God of Isaiah. |
No object is commoner in Chaldean and Assyrian pictorial
representations than the Sacred Tree, the Tree of Life, with
and without the serpent. To the old Chaldeans the Sacred
Tree was intimately connected with an original ancestral
abode, an earthly Paradise, watered by springs which became
great-rivers.
The “ serpent of Gatien we roçognize as Mummu-Tiamat,”
the “ Dragon,” the “ Great Serpent,” who was the sworn enemy
of the gods and their creation ; the principle of opposition and
destruction. The gods determine to fight the great Serpent ;
Anu, the heaven God, prepares the sickle-shaped sword, and
the beautifully bent bow, whilst Bel goes forth in his match-
less war-chariot, sending the lightning before him, and scatter-
ing his arrows around. Tiamat comes forth to meet him,
attended by evil demons, and bearing death and destruction
in her train. But it is Meridug, the intercessor for mankind,
who “ bruises the serpent’s head” ; who binds her and puts an
end to her works, while her followers fly terror-stricken.
I have not space to give a tithe of the profoundly interesting
and suggestive facts revealed by the deciphering of the cune-
112 | The American Naturalist. [February,
iform inscriptions. We find ourselves looking at the cradle-
land of our religion, our sciences and our literature; we find
_ the most touching and passionate appeals to the Divinity in
the very spirit of the finest Psalms of David, in a land looked
upon as the seat of the grossest idolatry ; we find a civiliza-
tion old enough to have been the parent stock of the civiliza-
tion of China; we look upon an early world, which knew
nothing of the proud white race which now girdles the earth
with empire.
Amidst the surging waves which have carried away so
many beliefs, we find two anchors linking us to the Unseen,
which become the firmer the more we know of evolutionary
processes. One is the enormous strength of the religious in-
stinct, even at the earliest stages of the civilization of man-
kind. And if the study of evolution teaches us one thing
more than another, it is that no instinct exists in vain. The
other foreign to my present subject is the inscrutable nature
of the “noumenon,” which lies behind all phenomena; that
Energy, which, whether we think of it as “gods” with the old
Accadians, as the “Supreme Being,” or by whatever name we
strive to approach the Unknown Reality, it remains equally
Unknown, (though perhaps not forever Unknowable,) and
equally the eternal object of search and worship.
1893.] Shell Heaps of Florida. 113
CERTAIN SHELL HEAPS OF THE ST. JOHN’S RIVER
FLORIDA, HITHERTO UNEXPLORED.
By CLARENCE B. MOORE.
(Continued from January Number, 1893.)
(Second Paper.)
West end of ridge on summit, 7x4x7} feet deep. Ata
depth of 1 foot 6 inches, an arrow-head with bones of lower
edible animals, was found. Two feet down were fragments of
turtle shell, and a half a foot lower, a fragment of a bone awl
was met with. At 5 feet, 6 inches, from the surface was a well
defined fire-place, with bones of the turtle. The shells are
almost exclusively Paludine, with occasional Unionide# and few
Ampullariz. .
EXCAVATION II.
North-east side of mound about ten feet vertically from the
base. Besides the usual fire-places and bones of edible animals,
a fragment of a Fulgur was found at a depth of four feet.
EXCAVATION III.
On summit near center of ridge, 9x9x11 feet deep. No
clearly marked strata were met with, the excavation being ~
carried through shell with a considerable percentage of broken
shell and sand. The following objects were found ;—flint flake
at a depth of 6 inches; fragment of bone awl 1 foot down;
small pendant ornament of shell 1} feet from surface, (Fig. 4)
Fic. 4. Size, }.
114 The American Naturalist. [February,
with bone implement; shell gouge (broken) 2 feet down; rude
arrow-head on fire-place, 44 feet down; flint flake 5 feet down ;
arrow-head lying on fire-place, at a depth of 6} feet from sur-
face ; portion of arrow-head on fire-place, bearing marks of fire
at a depth of 8 feet; fragment of bone awl 9 feet 9 inches
down ; rude bone awl 10 feet down; bones of edible animals,
mainly of alligator, deer and turtle, at all depths.
EXCAVATION IV.
At margin of base north-east portion of mound. Trench
113x10x6 feet deep at end. A piece of coquina, smooth
on one side, evidently used for polishing, at a depth of 2} feet,
and a fragment of arrow-head with the usual bones of edible
animals, were the only objects of interest met with.
As before stated, absolutely no pottery was seen in the exca-
vation, and this, with the 600 feet of swamp between the
mound and the river, would argue for Mt. Taylor a place
among the earliest shell heaps of the St. John’s. It was the
habit of the builders of the shell heaps where solid ground was
to be found, to locate their refuse heaps upon it, and it seems
unlikely that a spot so far distant from the river’s edge as is
Mt. Taylor to-day, should have been selected. It is a much
more likely hypothesis that the swamp is the result of a
change of channel in the river, and has formed since the
abandonment of the mound. Professor Wyman has graph-
ically described in his memoir (page 83 et. seq.) the manner of
formation of similar swamps. The time required to trans-
, form a portion of the river into swamp-land, the writer of this -
paper believes to be somewhat less than is generally supposed,
owing to the great abundance of floating vegetation upon the
river. At Little Orange mound, opposite Lake Dexter, is a
- steam boat wharf in use within eight years, according to’ the
owner of the place. At present, masses of vegetation extend-
ing some distance into the river have made access impossible _
for a boat of any draught, and are speedily forming the
nucleus of a future swamp.
_ Assuming this hypothesis of the formation of the swamp
after the completion of the mound to be correct, an indication
1893.] Shell Heaps of Florida. 115
of minimum age may be obtained in the case of Mt. Taylor
- through the size of trees growing between it and the water’s
edge. Between it and the river grow two cypresses, respec-
tively 14 feet in circumference four feet from the ground, and
19 feet, 3 inches, 5 feet from the ground ; while a little to the
north-west of the mound in the swamp grows a cypress 23}
feet in circumference at a height of 5 feet from the ground.
The cypress is a tree of slow growth, by some being placed
next to the live oak in this respect.
Another point illustrated by Mt. Taylor is that neighboring
shell heaps are by no means of necessity contemporary. In close
proximity to it in the river, is Bird’s Island,’ covered with a
shell deposit between five and six feet in thickness. Through-
out this deposit to the very bottom unornamented pottery is
found in great abundance. North of Bird’s Island, on the
western bank of the river, is the shell bluff described by Pro-
fessor Wyman (Note A.). This shell bluff, by the absence of
pottery and the small size of its Paludinæ, differs widely from
Bird’s Island, and is probably contemporary with Mt. Taylor.
Just what impelled the makers of Mt. Taylor to pile the
debris of their meals to such a height and at such an angle it
is impossible to surmise. No ridges of shell led to it, while no
shell fields surround it, and it is evident that great care was
taken looking to its isolation and to give to it its height and.
shape. In comparison with the shell deposits of Huntoon |
Island, of Bluffton, of Silver Spring (Lake George), of Tick
Island and other localities where the shell deposit, though
large, is spread over great areas and not piled at a single spot,
the contrast is sistema striking.
SHELL Deposit Orrostre BLUFFTON.
On the west bank of the river, opposite Bluffton, is a level
deposit of shell bordering the water. About fifty yards
inland is a small symmetrical mound, not over three feet in
The Paludinz on Bird’s Island, it may be remarked, are far larger than
any found at Mt. Taylor; well illustrating a point stated in the preceding
paper that shell fish of the St. John’s had a middle period of increased
development, and a subsequent relapse to their earlier condition.
116 The American Naturalist. [February,
height, composed of the same material as the shell deposit,
namely shell and powdered shell. An excavation was made
in the center of the mound, and within a foot of the surface
were found various parts of a skeleton, apparently of a woman,
The jaws and all other facial bones were missing, as were one
femur, one humerus, both tibiæ and all bones of the feet.
Many of the bones had fractures not caused during the excava-
tion, the area of the fracture being of the same color as the re-
mainder of the bone. With the remains were several bits of
pottery which seemed to be wanting in the mound, save in
association with the bones, and this fact would indicate a burial,
as would the symmetrical shape of the mound. The fractures
ean readily be explained by the proximity to the surface, where
the tramping of feet could without difficulty separate and
fracture the bones. On the other hand, the absence of so
many important bones—for which very careful search was
made—would seem to indicate that the entire body was not
placed within the mound. The various conflicting facts con-
nected with this case offer one of those questions, unfortunately
without solution, which from time to time confront the
explorer of the shell heaps.
Near this point Mr. Dease, of Volusia, found a small earthen-
ware bowl clinging to the roots of an overturned palmetto,
and presented it to the writer.
NOTE A.
As To A NATURAL ORIGIN OF ANY OF THE FRESH-WATER
SHELL HEAPS OF THE RIVER.
It will be remembered that at the time when Professor Wy-
man wrote, the agency of man in connection with the shell
heaps of the St. John’s was nowhere admitted. It is therefore
quite natural that taking the initiative in this matter he
should have been averse to making any sweeping assertions.
There is, however, little room to doubt that all the shell heaps
are the work of man. Of the shell bluff near Astor, (Fort
Butler) he says? (page 38) “taken by itself, in view of the
2 Op. cit.
1893.] Shell Heaps of Florida. 117
absence of human implements we should hardly feel jus-
tified in saying that it was made by man; though in its
general features it resembles his work. ” At the base of this
bluff the writer picked up a chisel of shell, and, what is more
conclusive (since objects not in place prove but little) a clearly
defined fire-place was discovered at a depth of four feet from
the surface. It is perfectly safe to say that no fresh-water
shell heap can be found on the St. John’s River where a care-
ful search would not reveal unmistakable evidences of human
agency in its origin.
118 The American Naturalist. [February,
THE ANCYLOPODA, CHALICOTHERIUM AND
ARTIONYX.'
By Henry FAIRFIELD OSBORN.
Chalicotherium was the most unique mammal of the Mio-
cene period. So far as known at present, it combined cheek
teeth of a distinctively ungulate type, ankle and wrist joints
resembling those of the Perissodactyla, with highly modified
phalanges, terminating in large cleft ungues which were
undoubtedly covered with strong pointed claws. It was
widely distributed over Europe, Asia and North America from
the oligocene Phosphorites of France (C. modicum) to the early
Pliocene of the Siwalik Hills (C. sivalense).
The recent discoveries of Filhol and Depéret in France
have fully revealed the structure of the skull and limbs, but
with the exception of the axis, little or nothing is known of
either the vertebree or scapular and pelvic arches. As we
learn from these authors the Chalicotherium magnum of Sansan
(Middle Miocene) was an animal taller than the grizzly bear.
The head was about nineteen inches long and raised five feet
above the ground. The limbs were rather slender, and a strik-
ing peculiarity of this species is that the fore limb is nearly
twice as long as the hind limb (as indicated by the proportions
of the radius and tibia). This was not the case in its upper
Miocene ally (Ancyllotheriwm pentelici) from Pikermi, in which
the limbs were heavy and of the same length. There were
three toes upon each foot, but in both manus and pes, the
fourth or outer digit was the largest instead of the third or
middle digit as in the Perissodactyla. M. Filhol has repre-
sented C. magnum as fully plantigrade, but this appears to mẹ
an error; the perissodactyl displacement of the carpals and
1 Under the title “ The acne of Chalicotherium,” this article was written some
months ago. While it was in press, Artionyx, a new type of the order was dis-
covered, and at the saio suggestion, the article was held back to include a descrip-
tion of it. o
1893.] The Ancylopoda, Chalicotherinm and Artionyz. 119
tarsals, the structure of the ankle and wrist joints, and the
terminal facets of the metapodials all indicate that the feet
were sub-digitigrade. Displacement is very rare in planti-
grade types. The proximal phalanges are peculiar in having
the metapodial facets directed obliquely upward; both these-
and the median phalanges are short; the distal phalanges are
` very peculiar, they present a deep terminal cleft and were evi-
dently drawn up or retracted, as in the cats, instead of being
drawn down or flexed, as in the Edentates. Thus, it is proba-
ble that Chalicotherium walked more like a clumsy-footed cat
than like the bear. The soles of the feet were slightly turned
inward and the fore-limb was adapted to digging or partial
prehension. :
Restoration. The axial skeleton in the accompanying draw-
ing is mainly based upon Filhol’s restoration, but the scapula
and pelvis are independently restored upon the perissodactyl
model; the sloping trunk leads us to expect a rather broad
` pelvic basin. All the above parts are purely conjectural.
The proportions of the fore and hind limbs with each other
and with the skull are based upon the figures of Depéret. In
Filhol’s drawing the skull is proportionately too small. The
side views of the feet are after Filhol’s excellent drawings, but
the articular positions are drawn differently, for M. Filhol
makes the animal fully plantigrade, while the metapodial and
phalangeal and higher articulations prove that it was digiti-
grade. s
What is the explanation of this remarkable assemblage of
characters? Where shall we place Chalicotherium, with ref-
erence to the two great orders between which it seems to stand
midway. If, observing the teeth and the carpus and tarsus,
we relate it to Perissodactyla, we find ourselves placing a
clawed type among hoofed types. Or is it to be considered an
Edentate because of its claws and edentulous premaxillaries
as Filhol has suggested? Or is it representative of a distinct
order midway between the Ungulata and Unguiculata? It is
too early to attempt a final solution of these questions. We
must wait for the discovery of the middle and upper Eocene
Chalicotheriide. It is evident from Gaudry’s discovery of the
120 The American Naturalist. [February,
feet in the Phosphorites, C. (Schizotherium) priscum, that the
finding of other Eocene types is not far off. In the meantime
I have tentatively offered the suggestion that Chalicotherium is
KOEI LIOR APR
CATTERY
N bat a PAA }
ae Fey if Lae k
en & ray i Wa 4 oa
Fig. 1. Restoration of Chalicotherium. Modified from Filhol and Depéret.
distantly related to Meniscotherium of the lower Eocene (Wa-
satch), and that this may give us a clue to its zoological posi-
tion. I will now expand and discuss this idea which may be
expressed as follows:
That the group to which Chalicotherium
belongs was derived from the Condylarthra
of the lowest Eocene, with affinities to the
Meniscotheriidae and primitive Perissodac-
tyla. It represents a distinct order, the
Ancylopoda (Cope). The likeness tothe Ungui-
culates and especially to the Edentata is dué
to secondary adaptations and contains no
proof of real affinity.
The history of the discovery of this animal is as unique as its
structure, and illustrates the uncertain path of the vertebrate
paleontologist. We are indebted for a full résumé, to Dr. Ch.
Depéret, who has fully discussed the bibliography, structure
and relations of this genus in his admirable memoir. Until
1893.] The Ancylopoda, Chalicotherium and Artionyz. 121
recently the skull and limb bones, wherever found, were placed
not only in different genera but in distinct orders. The first
remains discovered were the terminal phalanges, found in the
upper Miocene of Eppelsheim in 1825; they were sent to
Cuvier, who, noting the deep terminal cleft, named them Pan-
golin gigantesque. In 1833 Kaup found in the same beds the
isolated upper molars which he naturally attributed to an
ungulate, and named Chalicotherium goldfussi (syn. anti-
quum). Thus early began the confusion due to the wholly
diverse affinities suggested by the phalanges and the teeth.
In 1837 Lartet found the feet (in somewhat earlier Miocene
beds of Sansan) which he supposed belonged to a huge eden-
tate and termed Macrotherium giganteum. In 1853 Lartet also
discovered a skull and teeth at Sansan; he was unaware of
Kaup’s priority and first called the skull Anoplotherium, but
later separated it as Anisodon; Gervais, later, pointed out the
priority of Kaup’s term. In the upper Miocene of Pikermi
Gaudry found feet and limbs quite different from the Macro-
therium type; these he termed Ancylotherium. From the
Phosphorites, in 1875, he described the teeth of a new species,
C. modicum, and the feet of a supposed new genus, Schizo-
therium priscum. The skull and teeth were also found by
Falconer in the Siwaliks of India and termed C. sivalense.
Considering this exceptional mingling in so many horizons,
of one genus represented exclusively by skulls or teeth and
another by feet and limbs, Filhol, in 1888, first advanced the
conjecture that the two might really be one, and was happily
able to confirm this by his own discoveries in Sansan. Forsyth
Major independently arrived at the same concl usion from his
explorations in the Pikermi beds. Finally, in his memoir,
Depéret describes parts of a skeleton and skull found together
at Grive St. Alban, in beds nearly contemporaneous with San-
san. In the meantime it is probable that a similar confusion
has arisen with us. In 1877 Marsh announced the discovery
of phalanges of a large edentate in the middle and upper
Miocene of Oregon and Nebraska, and compared them with
those of Ancylotherium rather than with any true edentates;
he distinguished them by the coalescence of the first and sec-
9
122 The American Naturalist. [February,
ond phalanges as Moropus (species distans, senex and elatus).
The first undoubted remains of Chalicotherium in this country
were found by Garman in the Loup Fork (upper Miocene) and
described by Scott and Osborn. Cope has also described a
lower Miocene species (C. bilobatum) from the White River —
beds (Swift Current Creek) of Canada. It is not improbable
that Moropus belongs to the same types.
Nomenclature and Synonomy. Depéret advocates an impor-
tant modification -in our nomenclature. He believes that the
Eppelsheim type, to which the name Chalicotherium was origi-
nally applied, can be identified by the form of its phalanges
- with the Pikermi type (Ancylotherium Gaudry), and is, there-
fore, distinct from the Sansan type. For the latter he proposes
to retain the name Macrotherium Lartet. If this proves to be
correct these types should be distinguished as follows:
Middle and Lower Miocene.
Upper Miocene, Lower Pliocene.
(Sansan, Grive St. Alban, ? Phos- eens
phorites.
(Eppelsheim, Pikermi.)
es.)
Macrotherium. — Chalicotherium.
Syn.: Anisodon, ? Schizotherium, Syn.: Ancylotherium.
! Moropas. Quadrupedal. Heavy skeleton.
Semi-arboreal and fossorial. Light
skeleton. Fore-limb greatly elonga-
ted. Radius (= .70), tibia (= .29).
Radius with large external, and
small internal, fossa for humerus.
Fore- and hind-limbs nearly equal.
Radius and tibia of nearly same —
size; of heavy proportions. Radius
with subequal external and internal —
fosse of humerus. Ulna coalesced
Ulna distinct or very slightly coal- with radius.
esced with radius distally.
While many striking differences separated these two type —
especially in the proportions of the limbs, the teeth were sub- —
stantially similar. A strong family relationship is also exhib- —
ited in a peculiarity of the feet pointed out by Gaudry, namely; —
the proximal phalanges were retracted and not the distal, while - ’
in the cats the distal phalanges are retracted and in the edentates —
the phalanges are not retracted at all but are flexed. In the a
edentates the foot is thrown upon its outer side; in the Chali- —
cotheriide this was only partly the case. i a 2
1893.] The Ancylopoda, Chalicotherium and Artionyz. 123
Ungulate affinities of chalicotherium. M. Filhol’s view is
that this genus belongs to a group which bridges the gap
between the Edentates and Ungulates. When he showed me
his newly discovered skeleton in 1889, I was especially struck
by the perissodactyl character of the carpus and tarsus, and
considered it best to leave the genus in the Perissodactyla as “ an
aberrant form, with nearest affinities to Palaeosyops and gen-
era of that line.” It has also been doubtfully placed near the
Perissodactyla by Flower and Lydekker.
Depéret has now advanced sufficient evidence to exclude the
edentate idea entirely and shows clearly that the resemblances
which Chalicotherium bears to the sloths are purely superficial.
As shown above the adaptations of the phalanges for prehension
or digging involve an entirely different set of muscles from those
employed in either the Cats or the Edentates. This genus has
attained a somewhat similar functional result by a different
route—a case of analogy but not of homology. So with the
elongation and curvature of the fore-arm, the ultimate coales-
cence of the ulna and the radius, the backward direction of
the olecranon, the cleft ungual phalanges. All these are inde-
pendent and “ parallel” or “homoplastic” adaptations. Chal-
icotherium is still more positively separated from the Edentates
by the numerous ungulate characters which it displays.
Depéret is very much impressed by these ungulate structures
and places this genus definitely among the Perissodactyla. -
Let us, therefore, examine the ungulate hypothesis. Upon the
affirmative side is the structure of the molar teeth; they cer-
tainly bear a distant resemblance to those of Anoplotherium
and a more striking likeness to those of Palæosyops and Titan-
otherium. The reduction of the upper cutting teeth may be
regarded as a secondary adaptation which does not affect the
question of ungulate affinity either way. There are really
several questions involved; shall we waive the structure of
the terminal phalanges and call this genus an Ungulate?
Shall we then direct our attention upon the teeth and carpals
and tarsals and call it a Perissodactyl,or shall we follow Cope
and remove it entirely to the Unguiculata? Let us sum up
some of its leading characters:
124
The American Naturalist.
[February,
CHARACTERS OF CHALICOTHERIUM.
Shared by Ungulates.
Radius transverse
proximally, with two
fossæ for humerus.
Tibia with two fossze,
_ spine and crest.
um with
broad sustentaculum.
iplarthry in carpus
and tarsus.
Shared by Perissodactyla.
Tridactyl manus and
pes.
Astragalus and
carpal and tarsal facets
diplarthrous.
` Not found in Perisso-
dactyla.
Not mesaxonie.
fourth digit.
Secondary unquicu-
ate modifications of
limbs and phalanges.
Third trochanter ab-
sent.
' Large curved tym-
Buno-selenodont
Jaws with elevated molar type.
condyle and broad an-
panic bull. i
Reduction of upper
gle. incisors.
Molar teeth of sexi-
tubercular origin. :
This table shows that there are many characters, beside the
form of the phalanges, which excludes this family from the
Perissodactyla proper. The detailed comparisons of the skull
which Depéret institutes with Palxotherium and Palsosyops
relate to structures which are shared by all primitive Ungu-
lates. The only structures if which there is apparently con-
clusive evidence of a remote relationship to this great division
of the Ungulates is first in the teeth and second in the carpus
and tarsus. It remains, therefore, to be seen whether these
- also are products of homoplasy or “parallel adaptation,” oF
whether they really evince affinity. 3
Now let us turn to the broader question. Is Chalicotherium
to be classed as an ungulate or as an unguiculate in the Lin-
nean sense? We should, of course, compare it first with the
primitive ungulata which exhibit many unguiculate characters.
We are again guided partly by the excellent figures and descrip-
tions given by Depéret of C. magnum (race Rhodanicum). The
skull resembles that of the Ungulata Condylarthra in many —
features. The occiput is very low and broad and widely over-
hangs the condyles. There is a well arched sagittal crest and
a rather small cranium. The lower jaw has an elevated con-
dyle and a deeply extended angle. The external auditory
meatus is widely open below and there is a large mastoid pro-
1893.] The Ancylopoda, Chalicotherium and Artionyz. 125
cess. The maxillary bones are high. All these characters are
very similar to those in the skulls of the primitive Condy-
larthra, such as Periptychus, Phenacodus and Meniscotheriwm.
We should also mention many cranial features which we
believe belong to the category of secondary adaptive modifi-
cations; among such are the following: (1) the large curved
cylindrical tympanic bulle, (2) the feeble character of the
nasals, premaxillaries and anterior portion of the mandibular
rami. The latter modifications are associated with the reduc-
tion of the cutting teeth. Filhol has shown that the young
individuals of Sansan have a full set of incisors, while in the
adult the upper incisors are rudimentary or wanting. `
There are other secondary features of note. The condylar
facets are prolonged forward upon the basi-occipitals. The
posterior face of the centrum of the axis is directed upward ;
this and the preceding features point to peculiar flexibility of
the neck. The centrum of the axis is long and slender, the
odontoid process is round, the spine is unusually long and
high.
Looking again at the limbs we observe that the radius
covers the entire front face of the ulna and presents two dis-
tinct fossee for the condyles of the humerus. The tibia closely
resembles that of some of the short-limbed ungulates in its
cnemial crest and two fosse for the femur, in its spine and dis-
tal grooves for the double trochlea of the astragalus which
is like that in the short-footed Aphelops. This likeness to the
ungulates, and especially to the perissodactyls, in the form of
the upper foot bones and of their articulations with the meta-
podials is most striking. The calcaneum has a long tuber
and very broad sustentaculum. The astragalus rests widely
upon the cuboid, its tibial trochlea is very similar to that in
the short-footed rhinoceroses. The cuboid, navicular and
cuneiforms are, however, extremely flattened and specialized.
The diplarthrous character of the carpus is precisely like that
of Perissodactyla, the lunar rests upon the cuneiform and the
scaphoid upon the magnum, while the metapodials abut later-
ally against the unciform and magnum.
126 The American Naturalist. [February,
Upon the whole we find that primitive and second-
ary ungulate characters decidedly predomi-
nate in the skull, teeth and skeleton. The
secondary characters, especially in the ankle and wrist joints,
are parallel with those in the perissodactyls, and are not
mingled with unguiculate adaptations until we reach the
phalanges.
Relation to the Unguiculata. Cope was the first to advocate
the more radical view that this genus represents a distinct
order. He wrote: “It has little relation to the family of Per-
issodactyla to which it has given the name (Paleosyops).
Unlike the serial manus and pes of the edentata the carpus
and tarsus are here diplarthrous in structure or displaced upon
each other. While the Condylarthra are ungulate with an
unguiculate carpus and tarsus, this order Ancylopoda presents
the antithesis of including unguiculates with an ungulate car-
pus and tarsus.” This antithesis he assigned as the main — A
character of the new order? He still considers the form of
the terminal phalanges as of fundamental importance and
believes (in a letter of May 14, 1892) that Chalicotherium must
have been derived from some primitive unguiculate.
Lhe ancestry of Chalicotherium. Without any knowledge of
the ancestors of Chalicotheriwm we thus reach a dead lock.
Shall the unguiculate structure of the phalanges outweigh the — A
ungulate structures in other parts of the skeleton? Is it pos-
sible to derive such a skull and skeleton from any unguiculate
or such terminal phalanges from any ungulate? In looking
about for relatives of this genus we must now entirely discard
the Palæosyops group, and hunt among the lowest Eocene
types. The lowest Eocene unguiculates are wholly dissimilar.
The only form at all similar is the genus Meniscotherium.
Meniscotherium was a little plantigrade slightly larger than
_. Hyrax, and of very similar proportions. It is found in the — a
beds between the Wahsatch and Puerco and was even older than _ -
Coryphodon. The teeth are of the ungulate type known aS
7Gil’s term “Chalicotheroidea” was applied to these forms, considered
as a super-family, equivalent to the Camelide, Giraffidæ, etc., and not as 4 n
suborder. Arr. Fam. of Mamm., 1872, p. 71-77.
1893.] The Ancylopoda, Chalicotherium and Artionyz. 127
“buno-selenodont.” The feet are in the border region between -
the unguiculates and ungulates. This genus also occupies an
isolated position; although placed by Cope in the family
Meniscotheriide among the Condylarthra it is not even
remotely related to any of the contemporary ungulates. Its
molar evolution was precocious in both jaws, far more so than
in Phenacodus. I was led to the idea that this might possibly
be the long sought ancestor of Chalicotherium by the struct-
ure of the molar teeth. In 1886 Wortman regarded Menisco-
therium as an ancestor of the Hyracoidea. Later Schlosser
recognized the striking likeness of its molars to those of Chal-
icotherium, and considered it with Macrauchenia as a repre-
sentative of Perissodactyla which had retained a primitive
foot structure.
The comparison of the entire dentition of Meniscotherium
and Chalicotherium led me in 1891 to the discovery of many
very significant details of resemblance. In both, the anterior
portion (or cutting teeth) of the dental series is reduced and —
the posterior or cheek teeth are enlarged. In both, the true
molars have identically the same pattern in detail in both
jaws, including the absence of the third lobe upon the last
lower molar, which separates Meniscotherium from all early
Condylarthra and Perissodactyla. I was especially struck
also by the presence of a short posterior crest formed of the
hypocone and hypoconule in the upper molars, and by the
reduplication of the antero-internal tubercle (metaconid) in
the lower molars. At the tinie only the upper tarsals of Men-
iscotherium were known; these possess the unguiculate char-
acters which are exhibited in Phenacodus and all early ungu-
lates but are distinguished by a fibular facet upon the calca-
neum. It appeared to me altogether probable that the Wahsatch
form was related to the ancestors of Chalicotherium, and that
the question would be decided by the discovery of its feet.
This discovery was reported sooner than was expected in
Marsh’s recent paper upon M. (Hyracops) sociale in which the
fore and hind feet are fully figured. —
‘The feet of Meniscotherium present only one feature which
definitely points to those of Chalicotherium, namely, they are
128 The American Naturalist. (February,
functionally tridactyl, the outer toes being much shorter than
the three median toes. Marsh describes the terminal pha-
langes as intermediate between hoofs and claws, the extremi-
ties are thin, slightly expanded, and apparently covered by
thin nails. In other respects the feet are very primitive and
exhibit all the unguiculate features we now invariably expect
to find among the primitive ungulates, namely, (1) the “ cen-
trale ” and possibly (2) the “ tibiale,” (3) the fibular facet upon
2
_ Fig 2. Fore and hind Feet of Meniscotherium (Hyracops) sociale after Marsh.
the caleaneum, (4) the “astragalar foramen,” (5) the serial oF
undisplaced facets in both tarsus and carpus. As this was the
parent ungulate type there is no insuperable difficulty in sup-
1893.] The Ancylopoda, Chalicotherium and Artionyz. 129
posing that the ungulate carpus and tarsus of Chalicotherium
were derived from those of Meniscotherium.
The terminal phalanges present a greater obstacle ; is it pos-
sible that having progressed only so far toward hoofs as to be
called “intermediate,” they have subsequently retrogressed
into curved claw-bearing ungues? T consider this possible but
not in itself probable. We must not forget the manus of the
subungulate Lemurs which bears three or four terminal nails
and one large claw; or the phalanges. of the unguiculate
Mesonyx which are cleft, but also flattened and very ungulate .
in appearance. The line of division between claws, nails and
incipient hoofs was not very sharply drawn. Another diffi-
culty is seen in the fact that the median toe (III) of Menisco-
therium is enlarged, it is mesaxonic; while the lateral toe (IV)
of Chalicotherium is enlarged.
Fig 3. Fore and hind Feet of Chalicotherium after Gervais, (from Sansan.).
To offset these difficulties we find the striking similarities in
the molar teeth already noted, and in the dental series as a
whole. Meniscotherium is very exceptional in the absence of
~
130 ‘The American Naturalist. [rebrand
a third lobe to the last lower molar. The resemblance between
the skulls of these two types is also very marked in the whole -
region behind the infra-orbital foramen. We see the same
form of cranium, occiput, and auditory region.
I conclude, therefore, by giving a table of the résemblances
and differences between the two types. Both are numerous,
The question is, which are most fundamental ?
MENISCOTHERIUM. CHALICOTHERIUM.
1° Cutting teeth somewhat reduced............ Reduced or rudimentary.
s per molars, buno-selenodont e same ooo
A short posterior crest. Protocon- The same. Protoconule re-
ule large. duced.
Lower molars \opho-selenodont The same.
Metaconulid reduplicate. No 3rd
lobe on Mè. ` The same.
2° Five digits. Functionally tri- Three digits. Structurally —
dactyl. Mesaxonic. tridactyl. Not mesaxonie,
Carpus and tarsus, serial, (un-
guiculate type)........ Displaced (ungulate type).
3° Centrale, tibiale, 3rd trochan-
ter, entepicondylar foramen,
Fibulo-calcaneal facet All wanting.
4° Terminal phalanges sub-ungu-............... Unguiculate.
ate.
5° Plantigrade. Sub-digitigrade.
Many of these differences are such as separate higher from
lower forms, especially those under 3° and 5°. We witness
the loss of the primitive characters under 3° in many distinct
phyla, also the modification of the characters under 2° and 5°.
I have now endeavored to clearly state the pros and cons
of this question. The evidence for the original ungu-
late affinities of the Ancylopoda seems to me much stronger
than that for its purely unguiculate origin. Supporting this
view is the strong likeness of the skull and teeth of Chalice
therium to those of Meniscotherium, and finally the analogies
which it presents to the perissodactyls in the modernization of
the feet, wrist and ankle. My conception of its origin and
zoological relations can be expressed in the following diagra™-
aay ogee
Pan
Reh as
eS EE
1893.] The Ancylopoda, Chalicotherium and Artionyz. 131
Ancylopoda Perissodactyla
Meniscotheriide : Phenacodontidæ
i Condylarthra
Protungulata
(unguiculate)
The purpose of this paper is not; however, to express a final
opinion but to suggest inquiry. While agreeing with Cope
that the Ancylopoda represent a distinct division of the mam-
malia, we must admit that the broad zoological relations of
this division are yet to be determined.
c. External.
a. Internal.
Fig. 4. Right hind foot of Artionyx gaudryi.
POSTSCRIPT. — THE DISCOVERY OF ARTIONYX.
The hind foot of Artionyz gaudryi was found last summer
by the American Museum party, under Dr. Wortman, in the
White River Miocene of South Dakota. ;
The discovery of this foot was one of those complete surprises
which render paleontological research so fascinating. As indi-
-cated in the above article, we were daily expecting to find
remains of Chalicotherium in the lower Miocene of America,
but no one could have anticipated finding a type related, and
yet so entirely different.
132 The American Naturalist. [February,
As Chalicotherium may be broadly termed an unguiculate
perissodactyl,so Artionyx may be called an unguiculate artio-
dactyl. This, in fact, sums up the main difference between
these types when we add that the terminal phalanges in the
former are cleft, and in the latter are uncleft.
If this foot of Artionyx had the metatarsals cut off half way
down, no one would hesitate to call it truly artiodactyl. The
tibia, fibula, astragalus and calcaneum, the navicular with its
posterior hook for the great-flexor tendon, the conjoined
ecto- and mesocuneiforms, the nearly even pairs of toes on
either side of the middle line—all these structures bear a
marked artiodactyl stamp. The entocuneiform is missing,
but there is no doubt that it supported a first digit, of which
the proximal phalanx is fortunately preserved.
The distal ends of the metatarsals and the phalanges exhibit
the same sudden transition to an unguiculate type which we
observe: in Chalicotherium, in fact they bear a very marked
resemblance to the corresponding parts in the feet of the
carnivora, until we come to the terminal phalanges, which are
short, deep and laterally compressed, but not hooked or retrac-
tile. As observed above, they are not cleft.
The foot differs as widely from that of Chalicotherium as
that of the pig does from that of the tapir. It is somewhat
hazardous to make a deduction from the foot alone, but wè
have ventured to divide the Ancylopoda into two subdivisions.
TEREE. SUB-ORDERS.
Aneylopoda. Ungulate, with A. Perissonychia (odd clawed),
unguiculate terminal phalanges. with perissodactyl tarsus and
mesaxial reduction. Ungues cl
B. Artionychia (even clawed),
with artiodactyl tarsus and parax
jal reduction. Ungues uncleft.
Subsequent discoveries may show that these are merely twO
families—the Chalicotheride and Artionychide. This discov-
ery rather strengthens the idea of the relationship of Menis-
cotherium to the Ancylopoda—for we observe in Artiony* the
fibulo-calcaneal facet of the older genus, also the depression
upon the inner side of the astragalus.
1893.] The Ancylopoda, Chalicotherium and Artionyx. 133
RECENT BIBLIOGRAPHY.
1. Cope: “The Vertebrata of the Swift Current River, II.”
AMERICAN NaturRAuist, March, 1889, p. 151 (species
C. bilobatum, order Ancylopoda).
2. Depéret: “La Faune de Mammifères Miocénes de la Grive-
St. Alban.” Arch. Mus. d’Hist. Nat. d’Lyon, 1892.
3. Filhol: Assoc. Franç. Congrès Toulouse, 1888, p. 265.
“ Mammifères de Sansan.” Ann. Se. Géol., 1891, pp.
294-300, Plates 43—46.
4. Gaudry: Journ. de Zool., 1875, Plate 18 (Schizotheriwm
priscum).
“Les Enchainments du Monde Animal.” 1878, pp.
194-198.
“Fauna Attica.”
5. Osborn: “ Chalicotherium and Macrotherium.” AMERICAN
NATURALIST, 1889, p. 729.
“Meniscotheriide and Chalicotherioidea.” AMERI-
CAN NATURALIST, Oct., 1891, p. 911.
_ “Ts Meniscotherium a Member of the Chalicotheri-
oidea.” AMERICAN NATURALIST, June, 1892, p. 507.
“ Artionyx, a new member of the Ancylopoda.”
Bull. Am. Mus. Nat. Hist., February, 1893. (Osborn
and Wortman.)
134 The American Naturalist. [February,
EDITORIALS.
EDITORS, E. D. COPE, AND J. S. KINGSLEY.
—Boranists are waking up to the fact that they have too long
neglected to conform to the law of priority in nomenclature. The
return to older names of genera, species, etc., produces temporary
inconvenience, but it is in the interest of stability, which is the funda-
mental condition of convenience in this matter. Persons who are
accustomed to use certain names find it difficult to change them, and
endeavor to make exceptions to the general rule, as some leading bot-
anists are now proposing. But the sooner a clearing up of the whole
matter is made the shorter will be the diseomfort, for the law of pri-
ority is fundamental and must be supported under all circumstances
that fall within the scope of nomenclature.
It is evident, however, that in cases which violate other laws of —
nomenclature, the law of priority has no application. Such are typo-
_ graphic and orthographic errors ; also terms which cannot be given a
Latin or Greek form: Such rules are the conditions of ordinary schol-
arship. There is another rule which belongs to scientific scholarship
which has been hitherto respected, but which a small number of
American zoologists are endeavoring ‘to set aside. This is, that no
name or term is available for scientific use which has not been accom-
panied by a definition prior to the publication of any other name for
the same thing published with a description. So self-evident is the
necessity of this rule that it has been adopted without exception by all
scientific societies and committees in all countries which have proposed
or revised laws of nomenclature. The primary element in all exact
language is the definition of terms, and to omit this condition from
scientific nomenclature is to abandon science and to go into literature,
if mere word-making can be dignified by:such a title. When we
remember what a passion this word-making becomes in some persons,
it is clear that science must have some protection from it. The only
_ way that this protection may be had is to demand a raison detre for
every name that is proposed. The student then knows with what he
has to deal, and the proposer places himself on record in an available
form. The abuses of the neglect of this rule are self-evident, since it
lays the sciences open to charlatanism and to pretence of every i
tion, and makes scientific literature of the catalogues of the showman z
and the salesman.
1893.] Editorials. ; 135 .
The only body which has encouraged this abuse is the American
Ornithologists Union, which in its rules straddles the question. One
result of this position is that members of that body are not a unit in
their opinions on this point. A concrete case is the lists of names
published a half century ago by Fitzinger. In his early days this
naturalist made a creditable beginning of scientific work, but he soon
lapsed into a cataloguer and list-publisher. He projected comprehen-
sive systems which have little or no correspondence with nature, and
filled them out with names without definitions, many of which are not
applicable to anything which can be defined. So well known has this
- been in Europe that this work of his has been long since relegated to
the waste heap, where it should by all means be permitted to remain.
But the love of change has induced a few Americans at this late day
to resuscitate this rubbish. No better motive can be ascribed for this
resurrection, for it is not in accordance with existing rules; it is very
inconvenient; and it opens the door to continued inconvenience of the
same kind. It is a blow at scientific scholarship, and a distinct
encouragement to dishonest work. i
It is hardly likely that the botanists will split upon this rock.
—THE late meetings of the American Society of Naturalists and
the allied societies of morphologists, anatomists and physiologists, held
at Princeton, N. J., brought together a representative body of the
most active workers in the natural sciences of the eastern part of the
country. The west was ably represented by members from Cincinnati,
_ Denison and Chicago Universities. These meetings add to the pleas-
ures of the holidays, supplementing family reunions by professional
reunions of men bound together by similarity of tastes and modes of
thought. The advantages of these meetings are various, but one most
obvious one is the stimulus to care and precision which results from
free and friendly criticism. After living in an atmosphere of this
kind, the mind more fully appreciates the contrast between the more
or less careless, not to say disingenuous habits of thought that are so
common, and the clear, rational, and truthful attitude of the mind
which is so conspicuous in the workers in the fields of science. The
welcome given by President Patton was as clear and honest as the
minds of the men he faced, and it was a welcome to truth, let it come
from what quarter it may. :
The ceded were ok from an assembly which should include
suck a body of energetic workers as now constitute the Geological
Society of America. In view of possible further segregation, a com-
- 136 The American Naturalist. [February,
mittee was appuinted by the parent Society of Naturalists to suggest to
the society some measures looking toward a combined organization,
so that future cooperation might be assured. Such a combination
would hold a winter meeting corresponding to the summer meeting of
the American Association for the Advancement of Science, with the
difference that it would consist of professional naturalists only.
—Tue Smithsonian Institution has been requested by a petition
signed by a large number of American naturalists and by a resolution
of the American Society of Naturalists, to pay for the support of a
table at the Naples Marine Biological laboratory to be occupied by .
American students. It would seem that such an appropriation of
money is within the objects of the Institution as defined by the will of
Smithson. As a school for original investigators, the Naples Labora-
tory has been, and doubtless will continue to be, of the greatest utility. ,
PLATE I.
Protoceras celer Marsh.
1893.] Recent Books and Pamphlets. 137
RECENT BOOKS AND PAMPHLETS.
BEAL, W. J., AND C. F. WHEELER.—Michigan Flora. From the States Agri. Coll.
Mich.
BENEDICT, J. E.—Preliminary Descriptions of Thirty-seven New Species of Her-
mit Crabs of the Genus Eupagurus in the U. S. National Museum. Ext. Proceeds
Natl. Mus., Vol. xv, pp. 1-26. From the Smithsonian Institution.
BOETTGER, O.—Herpetologische Mitteilungen. Aus dem 29, 80,31 und 32 Berichte
des Oppenbacher Vereins fiir Naturkunde pag. 61, bis 164. From the author
— Katalog der Batrachier-Sammlungen des ‘asia der Senckenbergischen
Naturforschenden Gesellschaft in Frankfurt am Main, 1892. From the author.
Bulletin No. 7, Iowa Agricultural Experiment Station, May, 1892
CALL, R. E—The Tertiary Silicified Woods.——The Fishes of the Des Moines
Basin. pis 6% Abnormal Hyoid Bone in the Human Subject. Artesian Wells
in Iowa. Exts. Resend Iowa Acad. Science, Vol. i, Pt. 2,1890-’91. From the “aren
Cops, N. A.—Arabian Nematodes. Ext. Vol. v, Proceeds. Linn. Soc. N. S. W.
From the author
Corr, E. D.—On Some New and Little Known Paleozoic Vertebrates. Reprint
Am. Philos. Soc., May, 1892
Darton, N. H.—Fossils in the — Rocks of Central Piedmont Virginia.
Am. Jour. Sci., July, 1892. From the a
Day, D. T.—Mineral Resources -i < “United States, 1889, 1890. Seventh
Span of the Series. From the Smithsonian ——
DERBY, O. A.—Nepheline Rocks in Brazil. t. Quar. Jour. Geol. Soc., May,
1891. From the author
DEVIS, C. W Repuki on Post-Tertiary Phascolomyidæ.—[In Confirmation of
the Genus Owenia so-called.——The Incisors of Sceparnodon. Exts. Proceeds.
Linn. Soc, N. S. W., 1891, From the author.
Dona.pson, H. H.—The Extent of the Visual Cortex in Man as Deduced from
the Study of Laura Bridgman's Head. Reprint Am. Jour. Psychol., Aug., 1892.
From the author.
ETHERIDGE, R —On the Occurrence of the Genus Belonostomus in the Rolling
Downs Formation (Cntr of Central Queens!and. Ext. Trans. Roy. Soc. Vic-
toria, 1891. From the a
Garman, S.—On oe Collected by Dr. George Baur near Guyaquil, Ecuador.
——On Cophias and Bachia. Extr. Bull. Ess. ae t 1892.——Dr. D. H. Storer’s
Work on the Fishes. Ext. Proceeds. Boston Soc. Nat. Hist., Dec., 1891. From
the author.
GAUDRY, A.—Excursion dans l’ Amérique du aaa Ext. Bull. Soc. Geol. de
France, 8d série, t. xix, pp. 936-1024. From the au
GILL, go wie g yi Genus Hiatula of pea or Fantoga of Mitchell.
Ext. Proceeds. U. S. Nat. Mus., Vol. xiv. From the author.
_ HALE, H.—Language as a Test of Mental Capacity, Being an Attempt to Demon-
` Strate the True Basis of Anthropology. Ext. Trans. Roy. Soc. Can., Vol. ix, E
, From the author,
10
138 The American Naturalist. [Febra
HASSALL, A., AND C. W. StiLes.—Strongylus rubidus, a New Species of Nema-
tode Parasitic in Pigs. Reprint. Jour. Comp. Med. and Vetern. Arch., 1892. From
the authors.
Hircucock, R.—The Ainos of Yezo, Japan. Ext. Rep. Nat. Mus., 1890, pp.
429-502. From the Smithsonian Institution.
HOoLLICK, A vous Paleontology of the Cretaceous Formation on Staten Island.
Ext. Trans. N. Y. Acad. Sci., Vol. xi, No. 5. From the author.
Hom, TH he on the Flowers of Anthoxanthum odoratum L. Ext.
ceeds. U, S. Nat. Mus., Vol. xv, pp. 399-403. From the sctdoshad Institution.
Houcu, W.—The Methods of Fire-Making. Ext. Rep. Nat.-Mus., 1890, pp- `
395~409. From the Smithsonian Institution.
Howarp, L. O.—Insects of the Subfamily Encyrtine with Branched Antenne.
Ext. Proceeds. U. S. Nat. Mus., Vol. xv, pp. 361-369. From the Smithsonian
Institution.
James, J. F.—Of the Age of the Point Pleasant, Ohio, Beds. On Problematic
Organisms and the Preservation of Algz as Fossils. From the author.
KOEHLER, S. R.—White-line Engraving in the Fifteenth and Sixteenth Centuries. —
n DE Nat. Mus., 1890, pp. 385-394. From the Smithsonian Institution. i
VERETT, F.—On the Shediiewice of the White Clays of the Ohio Region.
a o Geol., Vol. x, 1892. From the author
MATTHEWS, W.—The Catlin Collection of Indian Aog Ext. Rep. Nat.
Mus., 1890, pp. 593-610. From the Smithsonian Institutio
PoucHET, G.—Contribution à l’histoire des Cyames, Eat Jour. Anat. et Physiol.
de Homme et des Animaux, 1892. From the author.
Report Regarding the Proposed Establishment of a National Aquarium in Sydney.
1892.
Roser, C.—Ueber die.v. Koch’sche Versteinerungsmethode. Aus Anat. Anz,
1892, Nos. 16 and 17. From the author.
SMITH, F.—The Gastrulation of Aurelia favidula Per. and Les, Ext. Bull. Mus.
Comp. Zool., Harvard Coll., Vol. xxii, No. 2. From the author. ;
Srarhpacinen, F.—Ueber neue und seltene Lacertiden aus den H Herpetologischen
Sammlungen des k. k. Naturhistorischen hofmuseums. Separat Abdruck aus Band
vi, Heft. 3und 4; Ann. des k. k. Piai sionen Hofmuseums. From the
STORER, H. R.—The Medals of Natural Scientists, Part 1. Ext. Proceeds. New-
port Nat. Hist. Soc., 1888-1891. a
- Third Annual Report of the Geological Survey of Texas, 1891. From the
rvey.
“‘Townsesm, C. H. T—A Remarkable New Hippoboscid from Mexico. Ext —
Entomol. News, 1891, pp. 105-106, From the author
_ Unter, P. R.—Observations on Some Remarkable Heteroptera of North Amet-
ica. Ext. Maryland Acad. Sci., 1892, p. 179. From the aut Si
WADSWORTH, M. E.—The Doña Inez and the Llano del the Meteorites from
, Chile, with EPEE A criptions. Ext. Proceeds. Rochester Acad,
Sci., Vol. 1. From the au
WATKINS, J. E.—The. Log » the Savannah. Ext. Rep. Nat. Mus. for 1890, pP:
611-639. From the Smithsonian Institution. _ B
TUER M.—Zoologische Ergebnisse einer Reise in Niederländisch Ost-Indien
Zweiter Band. Erstes Heft. Leiden, 1891. From the author. i
1893.] Recent Books and Pamphlets. 139
WILLISTON, S. W.—Higher Professional Requirements. From the author.
WILSON, TH.—Anthropology at the Paris Exposition in 1889. Ext. Rep. Nat.
Mus. for 1890. From the Museu
WOODWARD, A. S.—On a Ms ibik Tooth from the Wealden Formation of
Hastings. Ext. Proceeds. London Zool. Soc., 1891.
Supplementary Observations on oe non Fishes of the English Lower
Oolites. tena Proceeds. Geol. Asso., Vol.
keleton of a Chimeroid Fish (lschyodua) from pees ieee Clay of
Christian Maiford, Wiltshire. Ext. Ann. Mag. Nat. Hist., July, 1
holidophorus ae gee an Addition to the Fish Fauna i Z Upper Lias
of Whitby. Ext. Geol. Mag., Dec., 1891.
On the Lower sh Fish Fauna of Campbellton, N. B. Ibid., Jan., 1892.
The Hybodont and Cestraciont Sharks of the a es ricco Ext. Pro-
ae iseset Geol. and Polytechnic Soc., rom the fi
—A Review of Artesian Horizons in Southern aie Jersey. Ext.
“as en git Geol. New Jersey for 1891. From the a
ZIEGLER, F.—Modelle zur Demonstration der aon des Zähne des Mens-
` chen. From the author.
140 The American Naturalist. [ February,
RECENT LITERATURE.
Two Text-Books of Human Embryology.—Only a few
years ago the physician who wished to know anything about the
development of the human body found his whole supply of infor-
mation in one or two chapters of his physiological text-book. In
some respects the statements to be found there were reliable, but, at
least in most American physiologies, they were strange misconceptions,
as they could scarcely help being, for they were abstracts, at sixth or
seventh hand of the wonderful researches of Bischoff, of Rathke and
of von Baer, with never a bit of autoptic knowledge on the part of
the compiler. i
In the last two decades our knowledge of the embryology of man
and of the other mammals has been greatly increased, so that the
summary given by Balfour scarcely ten years ago is in many respects
behind the times. So a new résumé in English is very acceptable.
“Tt never rains but it pours.” Thesame week brings to our table two
works with essentially the same scope—that is the embryology of man
and the mammals, with side lights on the lower vertebrates—but of |
greatly different treatment.
The first to be mentioned, Minot’s Human Embryology, 18 A
large octavo volume, especially designed for the physician. That
is, it seems admirably adapted to lead the medical student or prac
titioner into a clearer knowledge of the history of the human body,
a knowledge of which the average physician has but the veriest
smattering. Minot gives first an introduction in which he describes
the uterus and outlines the stages of human development. Then the
main portion of the book is divided into five parts. In the first W°
Ot eee
have an account of the history of the genital products and the theory
of sex, together with a brief discussion of heredity. In Part Two the
origin and meaning of the germ layers are discussed. This portion
will prove more or less familiar to our readers, as several of its chap-
ters have already been given in our pages. In the Third Part the
chapters treat of the building up of the embryo, those of the Fourth E
the fætal appendages, while the Fifth Part takes up the fotus. AS
an appendix given is an exceedingly condensed list of references t0
the authorities quoted in the text. Such a table of contents as that =
‘Human Embryology by Charles Sedgwick Minot. N. Y., William Wood :
& Co., 1892, pp. xxiii + 815.
1893.] Recent Literature. 141
given above must fail to give an adequate idea of this book, and yet
it is about as much of detail as our space will allow. In general we
may say that the book pleases us; that while we find a few statements
here and there which we might question, we feel deeply indebted to
Prof. Minot not only for the compilation of such a valuable résumé of
the work of other embryologists, but for the numbers of new contri-
butions which we find in his pages. One feature which is of especial
_ value, and which cannot but strike the reader, is the richness of cita-
tion of the works of other students. The illustrations also are good,
and while most of them are process cuts, the majority have a freshness
which is very pleasing.
The other work upon our list, Prof. Mark’s Translation of Hertwig’s
Embryology,’ does not need very extended mention, as the original
has already been noticed in this journal. Its plan is greatly different
from that of Minot in that after the introductory chapters of Part
One the history of development is followed for the organs of the differ-
ent germ-layers: Entoderm, mesoderm, ectoderm and mesenchym.
The translation is done in a very satisfactory manner, and a careful
reading of a considerable portion of the translation fails to reveal
many of the “awkward renderings and German idioms” to which
Dr. Mark refers in the preface. One feature of the translation
deserves mention. For anlage the term fundament is used. Had this
translation been introduced several years ago it might hope for general
acceptation, but now that anlage is being bodily carried over into
English (Minot, for instance, uses it throughout), we hardly think
that the translation will prevail, especially in the face of the other and
not obsolete meaning of the word. The printing of the text is good,
but some of the cuts of the translation (clichés from the German cuts)
are muddy and lack the clearness and delicacy of the German edition.
It would have been difficult to find two men better adapted for their
respective tasks than these. For ten years Dr. Minot has been accumu-
lating the materials for his portly volume, and he has had excep-
tional facilities in working over the preparations of some of the
European masters. Dr. Mark, on the other hand, brings to the posi-
tion of translator an accurate and detailed knowledge of the subject
matter, and an almost extreme care that every word in his translation
shall express, so far as the genius of our language permits, exactly the
*Text Book of the Embryology of Man and Mammals, by Dr. Oscar
Hertwig, translated from the third German edition by Edward L. Mark.
London, Swan Sonnenschein; New York, Macmillan & Co. 1892, pp. xvi
+ 670. -
142 The American Naturalist. [February,
idea of his author. A comparison of the two works is difficult. For
the student who has already a good foundation in the broader features
of general embryology, possibly Mark’s Hertwig will prove the more
useful, for it gives more the broader features of ontogeny without pay-
ing great attention to minor and, for him, unimportant detail. Minot’s
work, on the other hand, from its method of treatment, may be better
adapted for the student of medicine, for it takes up the subject morein
accordance with the ideas which he receives from the rest of his special
training, and in those points which may prove of value in questions of
medical jurisprudence it gives a fulness of detail which is foreign to
the purpose of the other work. Yet the average practitioner will
probably have little to do with either volume. All that he cares to
know concerning the questions discussed, would occupy very few
pages, and the 670 pages of Mark’s Hertwig, and the over 800 in
Minot’s volume, will, we fear, scare away the very men who most need
the information they contain. Yet it is difficult to see how the matter
could be much more condensed. ‘The fault is not, in either case, with
the author, but rather with the extent of our knowledge. This, how-
ever, will prove no drawback for the student of morphology, and, we
doubt not, these will be two of the most consulted books in his library,
for they are the two most important additions in our language to the
literature of biology for the year 1892.
To conclude, we would say that both books are valuable, and should
be owned by all morphologists. Minot’s volume is possibly the better _ f
adapted from its arrangement for the physician, and the greatest
objection which we can see to its use by medical men is its size.
Geological Survey of Texas, 1891.’—This volume constitutes
the Third Annual Report of the Geological Survey of Texas. It com —
sists of the usual introductory statements of the State Geologist, E. T.
Dumble, followed by the several reports of his assistants. The accom —
panying papers are reports on the geology and paleontology Eo
important regions by specialists in those departments. Mr. Kennedy a
contributes the result of investigations which carried him across the .
post-cretaceous deposits, from Terrell, in Kaufman County, in a south- =
easterly direction to the mouth of the Sabine River. Mr. Cummins —
gives a detailed report on the geology of the Llafio Estacado. He r
traced the Carboniferous formation to its farthest outcrop in Central ,
SS he Sea
eS So
Texas, determined the northern extension of the Cretaceous along the
"Third Annual Report of the Geological Survey of Texas, 1891. E.T
Dumble, State Geologist, Austin, 1892.
1893.] Recent Literature. 143
eastern escarpment of the Staked Plains, and determined the extent
of the Dockum and Blanco Canyon Beds and their relation to the
underlying strata. The Stratigraphy of the Triassic formation in
northwest Texas was assigned to N. F. Drake, who embodies his obser-
vations in a short paper of twenty-one pages. Mr. Steernwitz is still
at work on the Trans-Pecos region. In his paper he states that rocks
of Carboniferous age have been traced over a large area of this region,
gives the age and relations of certain conglomerates, and the strati-
graphic relationship between the schists and the red grit. Prof. E. D.
Cope, who had in charge the fossil vertebrata, reports on collections
from the Fayette formations, the Blanco Canyon bed, and the Triassic
or Dockum beds. Dr. Sterki furnishes notes on shells found in a dry
salt lake near Eddy, New Mexico.
The illustrations are numerous and compare favorably with those of
previous publications of the survey.
Mineral Resources of the United States, 1889 and 1890.'—
` This volume is the seventh of the series of Mineral Resources of the
United States, and carries the statistical data to Dec. 31, 1890. An
idea of the amount and value of the various useful mineral products
is given in a brief summary which forms the opening chapter. The
remaining 530 pages comprise carefully prepared statistical papers
containing much valuable information.
According to the report the year 1890 was a period of unexampled
activity in mining, particularly so in iron, silver, copper, coal and
petroleum. The total value of the product is put at $656,604,698, an
increase far beyond any previous year.
In addition to its individual index, this volume contains the general
index to the entire series from 1882 to 1890 inclusive.
“Mineral Resources of the United States, 1889 and 1890. David T. Day,
Chief of the Division of Mining Statistics and Technology, Washington,
1892.
144 The American Naturalist. [February,
i General Notes.
GEOLOGY AND PALEONTOLOGY.
Paleozoic Formations of Southern Minnesota.—A sys
tematic account of the Paleozoic formations of southeastern Minnesota,
by C. W. Hall and F. W. Sardeson, is given in the Bulletin of the
Geological Society of America for 1892. The rocks described occupy
an area some 13,200 square miles in extent. They stretch eastward
from a straight line between Mankato and Hinckley to the State of
Wisconsin, and from Chengwatona southward to Iowa. These periods
of geologic time are represented by three formations: Upper Cam-
brian, Ordovician, and Devonian. These Paleozoic rocks are under-
lain by the Archean and Algonkian, and lie beneath patches of Cre-
taceous and a covering of quaternary débris save in the extreme south-
eastern corner included in Chamberlin’s “ driftless area.”
The Upper Cambrian is represented in this region by the Potsdam
sandstone and the Magnesian series ; the Ordovician by the St. Peter, —
the Trenton, and the Cincinnati beds. The Devonian is present in so
thin a layer in which so few fossils occur that it cannot be assigned to
any division of that group, but is supposed to belong near the middle.
The writers give the following summary of thickness and leading
lithological characters based chiefly upon their own measurements and
determinations : :
FEET.
Devonian. Not subdivided. php be and shale 10-15
( Cincin- f Wyko Limeston 20 70
nati. | Maquoketa. Shale at limestone 20
Maclurea “sins re
j Lingulasina limes
= Camar nas shaly risen »
: | Orthisina calcareous shale
Ordovician } Trenton 4 Zyeorpire shale Be
0
Stictopora shal 30
L Speco can ey bees Tamer añd shale 10
Blue rosi .12 =
Bu Paroa 15) “u
_ Saint Peter, not subdivided, iiio 15-11 an
t Shakopie dolomite 10-5 |
evator B (Ric hmond) medics
Magnesian, piin Sapa dolomite
Jordan sand
Caelio. 1 Saint Law kka me Malisacties and shales
Raden not subdivided, sandstones and shales
Total thickness of Paleozoic strata in Minnesota........++++
<
ined
S
1893.] Geology and Paleontology. 145
` The Genus Hybodus.—<According to Mr. A. S. Woodward, the
genus Hybodus, which was determined solely from the evidence of the
teeth and associated ribbed dorsal fin-spines, is now definitely deter-
mined by the other essential regions revealed by new specimens.
The fossils in question comprise numerous skulls and two portions of
the trunk of the typical Hybodus basanus from the Wealden of the
Sussex coast.
The characters which afford information are (1) the presence of
hooked cephalic spines, and (2) the occurrence of a vacant space
between the neural and hemal arches of the endoskeleton, which
must have been occupied by a persistent notochord. Mr. Woodward
is inclined to think that the presence or absence of cephalic spines will
eventually prove to be a sexual character—Proceeds. Yorkshire Geol.
and Polytechnic Soc., Vol. xii, 1891.
On Some Dicynodont and Other Reptilian Remains from
the Elgin Sandstone.—At the Aberdeen meeting of British Asso-
ciation in 1835, Dr. Traquair called attention to the skull of a Diey-
nodont which had been discovered in the Elgin sandstone of Cutlies
Hillock (= New Spynie). Since that time several other specimens
have been obtained from the same place, some of which are the prop-
erty of the Elgin Museum, while others belong to the Geological
Survey of the United Kingdom. These specimens are now being
worked out by Mr. E. T. Newton, and the following is a preliminary
note on the interesting results which have been obtained.
All the reptile remains obtained from Cutlies’ Hillock are in the
condition of hollow casts, the bones themselves having been dissolved
away; this, it will be remembered, was the case with some of the
examples of Stagonolepis from the Elgin sandstone, described by Prof.
Huxley, and the method of taking casts from the hollow cavities
which was adopted in that case, has been found of great advantage in
the present instance. The blocks when brought from the quarry were
more or less split open, exposing portions of the specimens. In some
cases these cavities were traced out and developed with the chisel,
while in others they were further split open, thus allowing casts to be
taken. In many cases these casts had to be made in several parts and
afterwards fitted together. The time and labor involved in this task
have been repaid by the restoration of the skulls and parts of skele-
tons of several Dicynodonts, and one or two other equally remarkable
forms of reptiles.
146 The American Naturalist. [February;
In most of these specimens, including that noticed by Dr. Traquair,
the skulls are similar in form, although differing in minor details, and
have a general resemblance to the South African Dicynodon and
Oudenodon, some of them having small tusks in the maxillary bones.
With most of these skulls parts of the skeleton have been found.
Two or three show the position of the vertebral column and ribs, but
up to the present no definite centra have been traced; besides this
there is evidence of scapula, clavicle, humerus, radius, and ulna, the
humerus having the characteristic anomodont expansion of the two
extremities. In two specimens the ilia are preserved. These forms
appear to be distinct from Dicynodon, and probably repro at least
two or three species.
Another skull presents most of the characters of Lystrosaurus,
but has a short muzzle and no teeth. The last, and by far the most
remarkable skull of this series, is about six inches in length, and has
the outer surface completely covered in by bony plates, the nostrils,
eyes, and pineal fossa being the only apertures. The chief feature of
this skull is the extreme development of horns upon the face and
cheeks, there being about thirty of these formidable defences varying
from about a fourth of an inch to nearly three inches in length,
besides some smaller bones. The dentition is pleurodont, and resem-
bles very closely that of the living Iguana; the palate is lacertilian,
but with the pterygoids united in front of the pterygoid vacuity. This
skull reminds one very strongly of the living Moloch and Phrynosoma, —
but it probably finds its nearest ally in the Pareiasawrus from the
South African Karoo bed. The detailed description of these speci-
mens is nearly completed and will, it is hoped, be shortly published—
Geol. Mag., Nov., 1892.
Cenozoic Insects.—In discussing the Rhynchophora of North
America, Mr. S. H. Scudder refers to the variety and abundance of
insect life during the early Cenozoic period as follows :
“In the earlier Tertiaries we not only possess in profusion represen-
tatives of every one of the orders of insects, but every dominating
family type which exists to-day has been recognized in the rocks;
even many of the families which have but a meager representation
to-day have also been discovered, and though many extinct genera
have been recognized, no higher groups, with a single exception oF
two, have been founded upon extinct forms. This is one of the most
striking facts which confronts the student of fossil insects. It is the
more striking from the delicacy, the tenuity, and miniatures of many
1893.] Geology and Paleontology. 147
of the forms which are here concerned; and the statement can be
enforced by the further fact that the parasitic groups—those which are
entomophagous—are represented, as well as many of those which in the,
present time show peculiar modes of life; thus we have representatives
of such microscopie parasitic insects as Myrmar, strepsipterons have
been discovered, the viviparity of the ancient Aphides has been shown
probable, the special sexual forms of ants and white ants were as
clearly marked as to-day, and the triungulin larva of Meloe has been
found enclosed in amber, showing that the phenomenon of hypermeta-
morphism had already been developed.—Proceeds. Boston Soc. Nat.
Hist., Jan., 1892.
Uplifts in the Sierras of California.—A paper of Mills in
the Bull. G. S. A., Vol. 3, contains the following account of the oro-
graphic movements from which result northern half of the Sierra
Nevada range.
Along the axes of the greatest uplifting of the present range a pre-
Mesozoic range arose carrying up both crystalline and metamorphosed
sedimentary rocks, which partially disappeared through erosion and
subsidence; then a mesozoic range arose and its strata became uptilted,
and it in turn was reduced by erosion and subsidence to very small
proportions, and then in Tertiary and Quaternary time has arisen the
present range, which is now undergoing its erosion.”
A New Pliocene Ruminant.—The right mandible of a large
animal found in the lowest Pliocene formation in the vicinity of Oran
has beer referred to the order of Artiodactyla by M. Pomel under the
name Libytherium maurusium. In dimensions it rivals Helladothe-
rium, with which it has some affinities, but from which it differs unmis-
takably. The fossil measures 30 centimetres from the posterior border
of the last molar to the posterior part of symphysis.—Revue Scien-
tifique, July, 1892.
A Remarkable Artiodactyle from the White River
Epoch.—In a late number of the Bulletin of the American Museum
of Natural History, Drs. Osborn and Wortman describe a species of
Artiodactyle which differs much from any form previously known.
A considerable part of the skeleton was obtained, together with crania
of both male and female. , The superior premolars (except the last)
are simple, thus excluding the genus from the Bodidea, and the cuboid
and navicular bones are distinct, a character which separates 1 from
148 The American Naturalist. [February,
the Tragulide. The cervical vertebrx are undescribed, but the char-
acters in general refer it to the Oreodontide rather than to any group
of the camel series. The metapodial bones are distinct, and there are
four in front and two posteriorly. Both sexes have a pair of low
horn-cores or tuberosities on the parietal bones, but the male has other
tuberosities and processes as follows: The superciliary borders are
produced posteriorly into a prominent lobe, and a low tuberosity rises
from the frontal at its anterior border, just in front of the orbit,
Each maxillary develops an anteriorly projecting process above the
last premolar tooth, and then rises into a huge elevated compressed
plate with thickened apex at the side of the narrow nasal fissure. The
latter opens far posteriorly, as the nasal bones are very short and wide.
The molar teeth are Oreodontoid, and the superior canines are worn
posteriorly, showing that there was a large inferior first premolar as in
Oreodon. ‘I'he canine has also the triangular section seen in that
family. The inferior canine is recumbent, and resembles an incisor.
No superior incisors. ;
The absence of superior incisors is a character which is uncommon
in the Oreodontidæ, but is found in the genus Merycopater, and prob-
ably in Pithecistes. The discovery of a horned member of the group
is interesting. The female of the species was first described by Marsh
under the name of Protoceras celer. It is probable, however, that it
is nearly allied to, if not actually pertaining to the genus Stibarus
Cope, described in 1873. The latter was founded on a fragment of
lower jaw with premolar teeth, a part which has not been found in
the species described by Drs. Osborn and Wortman. The latter was
found by Dr. Wortman in the upper White River beds; the Stibarus
obtusilobus is a smaller species, and comes from the lower beds of the
same series. I give figures of parts of the skeleton from clichés kindly
furnished by Dr. Osborn (Plates I and II).—E. D. Corr.
The White Clays of the Ohio Region.—In southeastern
Indiana and southwestern Ohio occur a series of clays overlying the
glacial beds and differing from them in color and structure. There
are similar clay deposits in southern Ohio outside the glacial boundary.
Mr. Frank Leverett, in a paper published in the Amer. Geol., July,
1892, cites evidence to show that these deposits are synchronous, that
they have a common origin, that that origin is independent of both
ice-dams and organic agencies, but is best explained by a fluvio-lacu®
trine hypothesis. Furthermore, they furnish important evidence
a the sequence of events in this region during the glacial
period.
1893.] Geology and Paleontology. 149
Geological News, General.—Mr. R. S. Tarr has published a
paper in the Am. Geol., July, 1892, calling attention to the relation of
secular decay of rocks to the formation of sediments. The essay is in
the main a summary of the present knowledge of secular disintegra-
tion. The rocks of the Guadalupe Mountains are referred by
R.S. Tarr to the Upper Coal Measures instead of the Permian as
determined by Shumard. Bull. No. 3, Geol. Surv. Texas——A Geo-
logical and Agricultural Survey of the State of Louisiana has been
undertaken in the interest of agriculture, the first publication of which
has just been issued. It consists of a preliminary report on the north-
ern part of the State, by Dr. Otto Lerch, covering the Topography,
General Geology, Water Supply, Distribution of Soils and Useful
Minerals. M. Adolph Carnot states that the relative age of fossil
bones can be ascertained by estimating the amount of fluoride of cal-
cium entering into their percentage composition. His method was
applied recently to determine whether a human tibia was contempora-
neous with the bones of animals with which it was associated. The
chemical analysis proved that the tibia contained only the same per-
centage of fluoride as a recent human bone, while the associated animal
remains vielded seven to nine times as much fluoride as their recent
analogues. Hence the tibia in question belongs to a later period than
the other fossils with which it was found, a result in which M. Riviére —
acquiesces on other grounds.—Comptes- Rendus, Vol. exv, pp. 337-339.
Archean.—Dr. Charles Barrois announces the discovery of Ra-
diolaria in the Archean rocks of Brittany. The fossils occur ina fine
siliceous matrix (phtanite) associated with fragments of graphite, in
the neighborhood of Lamballe, Cotes-du-Nord. M. Cayeux, to whom
the fossils were submitted, regards them as simple Radiolaria, of the
family Monospheridee—Natural Science, Oct., 1892. -
Paleozoic.—According to Dr. Riist, Radiolarian fossils are abund-
ant in Paleozoic rocks, and nearly all of the Paleozoic Radiolaria can
be referred to the genera recognized by Haeckel in the seas of to-day.
—Palxontographica, Vol. xxxviii, pp. 107-200.
Three me ai of fish—Protodus jexii, Diplodus problematicus,
and Acanthodes semistriatus—have been added to the list of Lower
Devonian Fish-fauna by Mr. A. S. Woodward. The fossils were col-
lected by Mr. Jex at Campbellton, New Brunswick, and are now In
the British Museum.—Geol. Mag., Jan., 1392.
150 The American Naturalist. | [February,
Mr. J. C. White reports a small collection of fossil plants from the
Wichita beds of Texas. They are essentially the same with the flora
from southwestern Pennsylvania described by Prof. Fontaine. This
makes an equivalency probable between the uppermost beds of the
Carboniferous system in West Virginia, southwestern Pennsylvania
and southern Ohio (the Dunkard Creek series), and those of the
Wichita beds of Texas—Bull. Geol. Soc. Am., Vol. iii.
Mesozoic.—A Termite, discovered in the English Lias by Mr.
Montagu Brown, exhibits even the patches of pigment on the wings.
It is described by Dr. Henry Woodward in the May number of the
Geol. Mag. under the name Paleotermes ellisii ——Mr. A. S. Wood-
ward reports Pholidophorus germanicus from the Upper Lias of
Whitby, Yorkshire. This Lepidosteoid fish was observed by Quensted
in the Lias of Wiirtemburg and named in 1858.—Geol. Mag., Dec.,
1891.—A mammalian tooth discovered by Mr. Charles Dawson has
been given the provisional name Plagiaulaz dawsonii by Mr. A. S. Wood-
ward, who describes and figures it in the Proceeds. London Zool. Sot.
Nov., 1891. This fossil is the first evidence of a European Cretaceous
mammal, A fine specimen of Belonostomus from the Rolling
Downs formation (Cretaceous) of Central Queensland is described and
figured by Mr. Etheridge in the Trans. Roy. Soc. Victoria, 1891, under
the name Belonostomus sweetii. The specimen is supplemented by four
other fragments so that all the more important features in the skeleton
of the fish are made known.
Cenozoic.—A proof of the identity of the diatomaceous beds of
New Jersey, Delaware and the outerops on the western shore of Ches-
apeake Bay is afforded by the presence of a single specific form of
diatom, Heliopelta, which occurs at the base only of this bed, and has
not been found elsewhere in the world, either fossil or recent.—Ann.
Rep. State Geol. New Jersey, 1891. Mr. Harlé reports the finding
of the right mandible of a monkey in a cave in the stone quarry °
Montsaunés. The fossil in question is that of a Macaque and as it
appears to Mr. Harlé to be a distinct species, he has given the name
Macacus tolosanus.—Comptes-rendus, fev. 1892.——A list of Birds from
the so-called Post-Pliocene drifts of Queensland, by Mr. De Vis, is 9°
interesting contribution to avian literature. The list comprises twenty-
eight species referred to twenty-four genera. The whole of the twenty-
eight species and seven or perhaps eight of the genera are extinct.
This change is very much the same as that observed in the case of the
marsupials——Proceeds Linn. Soc. N. S. W., Vol. vi.
1893.] -~ Zoology. 151
ZOOLOGY.
The Cercaria Stage of Amphistomum,.—Dr. Albert Lang
has completed' the life-history of Amphistomum subclavatum. The
adult lives in the frog, the young in the small Planorbis contortus of
Germany. He finds that the young become encysted upon the skin of
the frog, and as this is eaten at the time of molting the young obtain
access to the alimentary canal in that way. Lang also describes some
points in the anatomy of the Cercaria. Especially interesting is the
description of the excretory apparatus which cannot well be repro-
duced without a figure.
Fecundation of the Eggs of Clinus argentatus.—M. Fred-
eric Gintel gives the following account of the habits of a fish known
as Clinus argentatus during the deposition of the eggs and their fecun-
dation by the male:
“The extrusion of the eggs occupies about half an hour, and they
are deposited among the branches of a clump of seaweed, after which
the female goes away and the male comes in his turn to the alge to
fertilize the eggs, of which he then becomes the guardian until they
are hatched, driving away all the males and females that come near to
the nest. If the latter, however, are gravid, he invites them to lay
their eggs near those which he is already guarding. M. Gintel has
seen a male fecundate successively, and to care for seven separate
depositions by as many different females, at intervals of several days.
—Revue Scientifique, Août, 1892.
Preliminary Descriptions of New Fishes from the North-
west.—In the November number of Tae Naturatisr we described
a few of the new species collected in western Canada. We shall here
describe in a brief way the remainder of the new species. Full descrip-
tions as well as a general report on the collections und a comparison
between the fish faunas of the Red River of the North, the Saskatch-
ewan, the Missouri, the Columbia, and the Fraser River will appear
later, the MS. of the entire report being ready for the press.
1. Pantosteus columbianus E. & E.
Three specimens, 92-100 mm. Boise River, Caldwell, Oregon.
Related to P. generosus, the eye larger, the caudal longer. Head
"Berichte Naturf. Gesell. Freiburg vi, 81, 1892. `
152 The American Naturalist. [February,
42-43; D. II, 114 or 123; A I, 83; scales, 16-19-80-100-15 ; eye, 13-
2 in. snout, 11 to 12in interorbital, 3} to little more than 4in head. All
the fins pointed, the caudal lobes considerably longer than the head.
Light brown, with indistinct clouds of darker.
2. Notropis albeolus E. & E.
One specimen 73 mm. long. Medicine Hat, Assiniboia.
Related to N. maculatus and N. heterodon. D.94; <A. 84; scales,
4~35~—4 ; 15 scales before dorsal; teeth 4-4, 1, 2.
More slender than heterolepis ; fins all small; origin of dorsal over
ventral, equidistant from base of middle caudal rays and nares ; scales
closely imbricated, the exposed edges little higher than long. Lateral
line decurved, the tubes developed on fewer than ten scales; ventral
surface entirely white ; a plumbeous lateral band overlaid with silvery ;
a dark vertebral line from occiput to caudal; sides with a few dark
specks, dorsal surface more densely specked, the margins of the scales
darker.
3. Notropis heterolepis E. & E.
One specimen 35 mm.long. Fort Quappelle.
Related to N. heterodon and N. anogenus. D. 9}; A. 93; scales
5-35-4; 15 scales in front of the dorsal; teeth 4-4; dorsal inserted
equidistant between base of upper caudal rays and anterior margin of
eye, behind the last ray of the dorsal; scales loosely imbricated, almost
imbedded in front of the dorsal; scales along the median line with a
deep notch near the middle of the posterior margin; the line nearly
straight; a few black specks along the base of the anal, a dark line
from anal to caudal; a dark band from tip of snout along the sides to
the caudal; a conspicuous black curved line at the base of each scale
of the lateral line; all the scales above the lateral band dotted with
black; a narrow vertebral line from occiput to dorsal, a broad dusky
band on the back between the dorsal and caudal, between this and the
lateral band a lighter band; scales of back with dark markings;
series of minute black dots along each ray of the dorsal, anal an
anterior portion of pectoral ; dorsal and caudal quite dark.
4. Notropis reticulatus E. & E.
Twenty-four specimens from Fort Quappelle. Thirteen specimens,
the largest 76 mm. from Brandon.
Related ' to N. spectrunculus, fretensis, nitidus and topeka. It
approaches nearest to N. fretensis and topeka. From the former it
differs chiefly in the larger scales in front of the dorsal and from the
latter in the naked breast.
PLATE II.
Protoceras celer Marsh.
1893.] Zoology. 153
Head 4, depth 4-43; D. 9# or 103 (I or II, 83); A. 94 (II, 74);
scales 4 or 5-34-3 or 4; scales in front of dorsal, 12 or 14; teeth 44,
hooked, with evident grinding surface; head pointed, broad above,
and slightly convex ; snout decurved, pointed, the lower jaw included ;
mouth oblique, the premaxillary on a level with the lower margin of
the pupil or somewhat lower; maxillary reaching front of orbit; eye
large, considerably longer than snout, 3 in head, greater than interor-
bital; origin of dorsal over ventrals, equidistant from tip of snout
and base of upper caudal rays; longest ray scarcely extending beyond
tip of last when depressed ; anal low, the longest ray not extending
beyond tip of last when fin is depressed, equal to snout and eye; ven-
trals reaching vent, slightly longer than the highest anal ray; pecto-
rals little longer than head, less opercle; scales closely imbricated, the
exposed edges considerably deeper than long in the largest specimens ;
lateral line decurved, complete; breast naked.
Dark streak from anal to caudal, lower parts otherwise plain; a
dark vertebral line, a plumbeous band along the side: a faint spot at
the base of the caudal about as large as the pupil; a series of spots
along each side of the lateral line; upper parts of sides and the back
profusely spotted, the edges of the scales are black, giving the whole
part a reticulated appearance. The specimens from Quappelle are
darker than those from Brandon.
5. Notropis scopiferus E. & E.
Many specimens from Winnipeg, Brandon, Fort Quappelle,
Medicine Hat. ae
This species is evidently closely related to W. luciodus, from which it
differs in having a conspicuous jet black spot about as large as the
pupil at the base of the caudal fin.
Head 4-43; depth 44; D. 93; A. 104 (II, 83); scales 6-36 to 42-4 ;
14-18 scales in front of the dorsal; teeth 2, 4-4, 2 grinding surface
very narrow, on two teeth only; head heavy, compressed, flat above;
snout blunt, much decurved ; mouth small, little oblique; maxillary
extending to eye; eye, 3 in head, little less than interorbital ; lateral
line complete, evenly and gently decurved to above origin of anal Toon
conspicuous black spot at the base of the middle caudal rays; a sil-
` very lateral band, its dorsal margin distinct, color otherwise variable.
6. Agosia faleata E. & E.
Many specimens. Boise River, talene
Scales larger and fins much longer than in A. nubila.
Head 34 po 4}-5}; D. 11}; A. 9}; scales 53-60, about ten
above the lateral line; dorsal usually inserted directly over the origin
11 í
154 The American Naturalist. [February,
of the ventrals, the fin large, its anterior rays prolonged ; origin of
dorsal equidistant from base of middle caudal rays and nares; caudal
deeply forked, the lobes acute, 34 to 3% in the length ; anal rays 43-47
in the length; ventrals always more posterior in position than in
nubila, their tips extending to or past middle of base of anal; pector-
als not reaching ventrals.
A dark band forward from eye ; dark lateral band scarcely evident;
silvery below ; sides and back with numerous irregular, well-defined —
dark blotches; anal and sometimes ventrals with a dusky spot near
base in front; dorsal and caudal faintly mottled; crimson spots on
mandible, axil of ventrals and along base of anal.
T. Agosia shuswap E. & E.
Four specimens. Shuswap Lake, near mouth of Eagle River.
These specimens differ from A. falcata in a few characters; inter
gradations may be found, but none exist in our specimens.
Dorsal equidistant from base of middle caudal rays and posterior
half of eye, inserted directly over origin of ventrals; lateral band
well marked, otherwise as in A. falcata.
C. H. anp R. S. EIGENMANN.
The Larynx of Batrachia.—Dr. H. H. Wilder states that
Amphibia typically possess two pairs of laryngeal cartilages, an ante
rior pair of arytenoids, while the posterior, containing tracheal rings
as well as the future cricoids, are best called tracheal elements. In
Proteus and Necturus but one -pair is. present, while in many Anun
the tracheal elements fuse in the median line, forming a sort of cricoid.
Wilder suggests, arguing from position, innervation, and musculature,
that the arytenoids are the fifth branchial arches and are further
homologous with the so-called inferior pharyngeal bones of Teleosts.
The tracheal elements, on the other hand, are new formations.
The Kidney of Amphiuma.—Dr. H. H. Field has studied the
kidney anlage in young Amphiuma and finds’ that here as in the ge)
cilians as described by Semon, there is a true metamerism. In
material (which has already served for other studies published in these
pages‘) the kidney reached back to the cloacal region, and it was poe
sible to follow the different stages of formation of the canals. In the
caudal region in the middle of each somite there was a wide, sharply
*Anat. Anzeiger, vii, 570, 1892.
*Verh. Deutschen Zool. Gesellschaft, 1892, p. 113.
‘Hay, Am. Nar.. xxii, p. 315, 1888; Kingsley 1. c. xxvi, p. 671, 1892-
i
1893.] Zoology. 155
bent tube which communicated on the one hand by a nephrostome
with the body cavity, and on the other empties into the segmental duct.
Immediately above the nephrostome was the malpighian body. Going
forward the looping of the duct becomes more extensive, and extends
from wall to wall of the somite, but no anastomoses occur and the
nephrostome is always in the middle of the somite. The ontogeny of
this organ will doubtless prove of extreme interest, and this point goes
to support Cope’s thesis of the relationships of Amphiuma to the
Gymnophiona.
On a New Spade-Foot from Texas.—Spea laticeps sp. nov.
—Frontoparietal bones without fontanelle, smooth. Form of head
and body short and wide. Width of head at tympana entering length
of head and body two and one-quarter times. Front wide and per-
fectly flat, its width entering the length of the tibia two and one-quarter
times, and considerably exceeding the width of an eyelid. Muzzle
Spea laticeps Cope, nat. size.
projecting little beyond lip; nostrils nearly terminal ; canthus ros-
tralis not marked ; tympanum concealed; vomerine fascicles small,
posterior to a line connecting the choane. The latter exceed in diam-
eter the well-developed ostia pharyngea of the eustachian tubes.
The heel of the extended hind leg reaches the position of the tym-
panum. Its digits are half-webbed, three and one-half phalanges of
the fourth digit remaining free. On closing the fingers the second a
little exceeds the third in length, and the third exceeds the fifth. The
integument of the superior and lateral surfaces extends to or beyond
the middle of the femur. It is obscurely tubercular, and the inferior
surfaces are smooth. There are no distinct glandular aggregations of
156 The American Naturalist. ` [February,
crypts. The length of the head and body is 38.5 mm. ; length of head
(axial), including tympanic region, 14.5 mm.; width of interorbital
space, 6 mm. ; of head at tympana, 18 mm. ; length of fore leg from
axilla, 19 mm.; length of hind leg from urostyle, 41 mm.; length of
tibia, 12.5 mm.; of foot, 20 mm.
Color of superior surfaces brown, varied with large obscure spots of
‘dark brown.
There are no pale longitudinal stripes, but the borders of the sacral
region are pale and contract posteriorly, enclosing a narrow dark band
above the urostyle. This pale outline is bordered externally by brown,
and from this dark spots or bands extend transversely. There is a
dark spot on the canthus rostralis ; one on each side of the end of the
muzzle, one below the eye, and one posterior to the eye. Gular region
light yellow, rest of lower surface dirty white. Sheaths of hallux black
bordered.
The single specimen of this species before me is smaller than the
average S. hammondii proper, and is equal to the larger individuals of —
the 8. h. bombifrons. Its color and the position of its vomerine teeth
are more like those of the Scaphiopus couchii, but the proportions and
the perfectly smooth cranial bones with free integument, distinguish it
readily.
This interesting species approximates more closely the genus Scaph-
iopus than the known species of Spea. It was found by Mr. Wm. L.
Black, of Prof. W. F. Cummins’ party of the geological survey of
Texas, on the journey between Seymour, in northwest Texas south
the Red River, and Austin. Mr. Black obtained on the same expedi-
tion three species of Bufo, viz., B. compactilis, B. debilis and B. pune
tatus. The range of the last named is thus much extended eastward.
A specimen of the snake Tropidoclonium lineatum was also obtained,
which extends its range much westward, Dallas having been the pre
vious limit in this direction —E. D. Corr.
The Pedal Skeleton of the Dorking Fowl.—Mr. G. p. ;
Howes has been studying the pedal skeleton of two Dorking chicks,
reaching conclusions somewhat different from those of Mr. Cowper,
who has recently published studies of the adult structure. As a result
of his anatomical researches Mr. Howes advances the theory that the
two innermost digits of the Dorking fowl’s pes are the product of the
cleavage of the hallux, and adduces the recent discovery by Kiikenth
of an undoubted occasional splitting of the Cetacean pollex asa SUP
port of this argument. Mr. Howes states also that the types of Pr
1893.] Zoology. 157
hexadactyle limb met with in the Amniota, as well as the pentadactyle
pes of the Dorking fowl, is a monstrous formation, and that arguments
deduced from their study in support of hexa- and heptadactyle limbs
are premature. The author calls attention to the fact that inasmuch
as no observed amphibian either living or extinct is known to have
borne more than two phalanges upon its second digit, neither the sau-
-ropsidian nor the mammalian types can be derived from that of the
Amphibia except by a process of intercalation such as there is no evi-
dence to support. As involving the Mammalia, the force of the argu-
ment against a belief in the amphibian origin of the class it is second
only to that of the discovery of the mammalian condition of the
limbs in the Theromorous Reptiles—Jour. Anat. and Phys., Vol.
XXvi.
158 The American Naturalist. [February,
EMBRYOLOGY.’
Experimental Embryology.—Under this title may be included
two most interesting recent contributions to our knowledge of the sig-
nificance of the cleavage phenomena in animal eggs. Both deal with
. the cleavage phenomena of eggs influenced by experimental interfer-
ence; the one in the case of a Vertebrate, the other in an Echino-
derm.
Prof. E. B. Wilson,’ of Columbia College, finds that the cleavage
cells of an Amphioxus egg when shaken apart are yet capable of
developing into gastrulas or even into complete larve of perfectly
normal structure. Yet the size of the gastrule or larve is determined
by the size of the cleaving cell; thus if one of the first two cells is
taken the resulting gastrula is half the normal size, if one of the first
four is taken, the gastrula is one-fourth the normal bulk. Gastrulas
of one-eighth the normal size occurred but were not products of isolated
cells in the eight-cell stage; in fact, the experiments go to show that
one of these first eight cells is incapable of forming a gastrula. There
is thus an early limit set upon the potentialities of the cleavage cells.
This, moreover, seems to be a qualitative and not a quantitative limit.
The homogeneous, undifferentiated character of the cells is lost at the
eight-celled stage.
When the cells are incompletely separated, double, triple, and even
quadruple embryos are formed.
Comparing the development of the isolated cells of Amphioxus with
what is known of similar phenomena in-other Vertebrates as known
from the work of Roux upon the frog, or with the facts established for
Echinoderms by Dreisch and others, we find an important and fun
mental difference in the time at which individuation and symmetry be
come apparent. While in the Echinoderm the partial development pro-
ceeds at first as if all the cells were present, and only later what maf,
improperly, be called a p fregeneration, forms that which waslack =
ing to the completion of a normal symmetrical embryo, in Amphioxus =
as the author shows by di ivi te ol tions than
had hitherto been made upon the exact mode of cleaving,—“the isolated
bss department is edited by Dr. E. A. Andrews, Johns Hopkins Unr
"On Multiple and Partial ioxus. Anat. Anz, IAA i
Nos. 23-24. Pp. ; ial Development in Amphioxus. Ana , a
1893.] Embryology. 159
blastomere develops as a unit, not asa half-unit ; and the cells to which
it gives rise cannot be individually identified with those of a normal
embryo-half. The development is transformed from the beginning.”
The second paper, by Hans Dreisch, of Zürich,’ contains many
important facts not presented in the preliminary article noticed in the
December NATURALIST.
When the eggs of certain sea-urchins are exposed to high tempera-
tures an acceleration in the rapidity of cleavage processes is not the
only result—but at a temperature of 31° C. certain marked changes
both in the size and position of cells, the entire absence of micromeres
and other such fundamental effects are produced. Nevertheless eggs
that cleave in these abnormal ways may develop into perfectly normal
larval forms, plutei. Long continued exposure to such temperatures
acts like the mechanical force applied in shaking eggs in a test-tube
and results in the separation of cleaving eggs into two, rarely more
than two portions, each of which may then develop by itself. As
some of these portions form normal larve, high temperature may be
one of the factors concerned in the formation of multiple embryos.
Returning to the cleavage of cells separated by shaking echinoderm
eggs, a series of figures are given to illustrate the fact that the isolated
cells continue to cleave, for a time, as if still in combination with their
fellow cells. Thus three cells taken from a four-celled stage do not at
once fill up the gap left by the removal of the fourth and this fourth
cell divides by itself just as it would have done if remaining in contact
with the three others. It is only at a later stage that the products of
isolated cells arrange themselves so as to form a complete, symmetrical
individual.
A portion of the paper is taken up with certain abnormal methods
of cleavage, beginning with the simultaneous formation of four instead
of two cells, at the first cleavage. In these cases the cleavage contin-
ues to be double, a sixteen-cell stage having two sets of each eight cells
comparable to the entire eight-cell stage of a normal egg. No larvæ
could be reared from such eggs, yet special interest attaches to them
upon the assumption that they owe their dual nature to a double fertili-
zation. The only evidence that these eggs were fertilized by two
sperms is the assumption by Fol and others that double fertilization
causes the formation of four in place of the normal two first cleavage
A most important addition to the methods of experimentation has
resulted in unexpected results. It consists in subjecting echinoderm
*Entwicklungsmechanische Studien. Zeit. f. Wiss. Zool., 55, 1892. .
160 The American Naturalist. [February,
eggs to mechanical pressure by the very simple expedient of allowing
a cover-glass to rest upon them with more or less force as controlled by
` a hair inserted under the glass.
Eggs flattened out to several times their original diameter do not
Jose their power to develop, even when the egg membrane is ruptured.
If the eggs are fertilized and undergo cleavage under pressure the
resulting cells are not arranged in the form of a sphere but form a flat
isk. This arrangement of the cells results from the arrangement of
the nuclear spindles. While in a normal egg the spindles of the first
four cells would stand vertically, as it were, and produce a new set of
cells by a horizontal cleavage, under pressure the spindles are found
all lying at right angles to the compressing force; that is, parallel to
the cover glass or horizontally, and hence give rise to cells all lying in
one plane. When the pressure is removed the cells remain at first in
a flat disk, though each cell becomes rounded. Subsequent divisions
of these cells give rise to a spherical mass capable of forming a com-
plete normal embryo.
With the aid of figures the author makes it clear that in such cases
material which would normally have taken part in the formation of
only one pole or side of the embryo must now contribute to opposite
poles or sides. In the sea-urchin there is thus no early specialization
or differentiation of the material found in the cleavage cells; the mix-
ing up of the blastomeres would not prevent the formation of a com-
plete normal animal: the author concludes they could be mixed in all
_ possible ways without destroying the normal symmetry of the adult.
In the philosophical discussion that takes up the final portion of the
paper, the author considers the methods of morphological research and
upholds the experimental method, which tends to promise the redut-
tion of Biology to a scientific basis sooner than the descriptive, histor-
ical or mechanical lines of inquiry possibly can.
Studies in Insect Embryology.—Dr. H. Henking* having pře-
viously published extensive researches upon the formation, maturation
and union of the sexual cells in Pieris brassica and Pyrrhocoris apterus
concludes this comparative study with additional facts upon the eB o
of the latter insect as well as those of Agelastica ulni, Donacia lam-
pyris, Tenebrio, Lasius, Rhodites, Bombyx, Musca, and other represen-
tatives of nearly all the chief groups of insects and then summarizes
the whole with the aid of most interesting comparisons and sugges- |
.
tions.
* Zeit. f. Wiss. Zool., 54, 1892; 51, 1891; 49, 1890.
1893.] Embryology. 161
Along with the observation of normal processes, attempts were also
made in some cases to influence these processes by increasing or dimin-
ishing the atmospheric pressure, by placing the egg in horizontal and
vertical positions, by surrounding them with CO, and with O; but the
results are not definite enough to afford more than suggestions for
future work.
Regarding the maturation of the egg the author holds views that
depend upon the interpretation of peculiar appearances seen in insect
eggs, where, as is well-known, the technique is a most difficult factor.
The first polar body receives one-half of the chromosomes, one-half
the normal number of the ordinary cell, and also the first thelyid.
This thelyid or “ acromatic polar body,” is a rather vague mass of
acromatic substance including the connecting filaments and adjacent
substances between the two separating sets of chromosomes.
The second polar body receives the same number of chromosomes as
the first, these being formed by a division of each, so that a half of
each remains to form the egg nucleus; in some cases there is a second
thelyid, but this remains inside the ovum.
The two polar bodies may separate completely from the egg, but
more often they remain either in protoplasmic processes of its surface
or entirely beneath its surface; the first polar body may divide into two.
Even when the polar bodies are extruded they may be later taken
into the egg again. Their nuclei may undergo changes similar to that
of the egg nucleus, either retaining the separate chromosomes or form-
ing a chromatic network. The first and second or the product of the
division of the first and second may fuse with one another inside the
egg, much as the sexual pronuclei do. :
Meanwhile numerous sperms may enter an egg; polyspermy mr
insects has no apparent injurious results. The sperm passes into the
yolk in a bent attitude, with the union of head and tail preceding. A
clear area about the sperm, the arrhenoid, seems to give rise to radia-
ting striæ in the protoplasm. This peculiar body is regarded as formed
from the acromatic connecting filaments given to the sperm at the sec-
ond division of its mother cell; is thus comparable, in a way, to the
thelyid. Increase in size and change in the sperms take place simulta-
neously in all that penetrate the yolk; only one meets and unites with
the female pronucleus. i :
The number of chromosomes in the nuclei of ordinary somatic cells
and in the cleaving egg is 28, 24, 20, 18-20, 24-30, in five different
insects studied ; the number in the first and also in second polar bodies
in these same species, 14, 12, 10, 9, about 12. As far as studied the
162: . The American Naturalist. [February,
spermatocytes have also this same half-number after their first and
second divisions; with some interesting exceptions the sperm and the
polar body and the female pronucleus have the same number, in any
species, and this is half the number of other cells of that species.
With the aid of a diagram the author emphasizes the close parallel
between the formation of sperms and polar bodies. Starting from a
mother cell with, say 24 chromosomes, a reducing division forms two
sperm mother cells, or in the female an egg and a polar body, in each
of which are only 12 chromosomes, and these are half of the original
24, unchanged or undivided individually. A second division of each
cell results in the formation of four sperm mother cells or in the
female of three polar bodies, and the remaining egg, in each of which
are 12 chromosomes. These, however, are formed by an equal divis- .
ion, that is, from the previous 12, 24 arise by a division of each—not
by the separation of the whole set into two sets as in division with
reduction. Later the 12 chromosomes of a sperm may be added to the
12 of the egg nucleus to make the normal 24 of the cleavage nucleus,
whence, by equal division, all subsequent cells have 24 also. Likewise,
in the insects, one polar body may unite its 12 with the 12 of another
polar body.
Utilizing the recent researches of Guignard, the author draws &
most interesting comparison between the processes taking place in
insects andin plants. Without the aid of diagrams this can scarcely
be rendered intelligible ; the most novel part, however, is an attempt
to explain the polar bodies of animal eggs as, in a sense, “ larval
organs,” somewhat comparable to rudimentary structures within the
embryo sac of a flowering plant.
The fact that in insects the polar bodies may remain within the egg
and that they, or their descendants, may unite with one another, seems
to form a parallel to what takes place in the flowering plant. As 1$
well-known, the embryo-sac cell forms two sets of each 4 nuclei within
its substance, 4 lower antipodal cells and 4 upper cells, of which one
is the ovum.
The formation of these is directly comparable to the formation of
polar bodies in the animal—supposing a second series of divisions to
supervene and convert the 4, ovum and three polar bodies, into eight.
In the embryo-sac a remarkable fusion takes place between one of the
antipodal nuclei and the sister cell of the ovum; this union forms the
endosperm. Compared with the insect, this union is like the fusion of
polar bodies, only in a later generation. In the insect nothing appa
See this Journal, May, 1892.
1893.] Embryology. 163
ently results from the fusion of polar bodies; they are rudimentary
larval organs.
The fact that we here compare union of later generations in the
plant with that of earlier ones in the insect or animal is perhaps made
less objectionable by considering the state of things in the infusoria,
where, as Maupas has shown, the final fusion takes place between
nuclei resulting from more divisions than are employed in the
formation of sperm and female pronucleus.
The observations of Henking upon the number and division of the
chromosomes in insects conflict in an important particular with the
facts necessary to support Weismann’s theoretical explanation of the
use of polar bodies. In his “ Amphimixis” Weismann holds that
reduction takes place in the formation of both polar bodies, and that
there was previously a doubling in the number of chromosomes or
idanten to allow of greater complexity of combination of the iden or
hypothetical hereditary units found in the chromosomes. Hinking
denies that the second polar body is formed by reduction ; it is formed
by an equal division. Moreover, there is no previous doubling of
chromosomes; the apparent doubling is only a more or less pronounced
division into parts still remaining subservient to the whole, not acquir-
ing individuality. -
Weismann’s views are moreover limited in that they depend upon the
occurrence of two polar bodies, two divisions in the formation of sex-
ual cells; whereas in plants and in infusoria, nuclei that need to copu-
late result from more numerous divisions.
Parthenogenesis again, as seen in hymenoptera and lepidoptera, may
occur where two polar bodies with a smaller number of chromosomes
are present. > l ;
Though recognizing the strength of the evidence that heredity is
closely associated with the chromosomes, the author thinks they are
not unchangeable, and that at times they may be more intimately com-
bined with the other parts of the nucleus. If the centrosomes are to
be regarded in heredity as of like value with the chromosomes then
we must seek in them a process of reduction before copulation.
All present theories of fertilization and of inheritance have only a
provisional character.
164 The American Naturalist. [February,
ENTOMOLOGY.
On a Small Collection of Coleoptera from the High
Mountains of British Columbia.—A small series of Coleoptera
furnished the writer by Messrs A. G. Smith and F. Russell is of inter-
est sufficient to deserve notice; the collection was made near Donald,
B. C., on the top of the first range of mountains to the northeast and
above the base of the snow line. Some of the beetles are circumpolar
species, others are melanistic forms of those found in more favored sit-
uations. A list of them with remarks is appended :
Carabus oregonensis Lec.
One specimen of the insect was taken ; it is darker than those found
on the coast and somewhat smaller.,
Nebria sahlbergii Fisch.
Numerous specimens of small size and with rather dark legs. This
seems to have been the most common beetle on the mountain, and is
characteristic of boreal or mountainous districts; it extends from the
hite Mountains of New Hampshire to Alaska, and Dr. Hamilton
writes that he has it from the Coast Range of the south of California.
Bembidium incertum Mots. i
One specimen ; this is reported from Sitka, Alaska.
Pterostichus riparius Dej.
Four specimens; these are a little smaller than those from the coast
of Southern Alaska, but about the same as a number from the Stikine
River of British Columbia. It has been reported from Gray’s Peak,
Colo., at an elevation of 12,000 feet. Dejean described it from our
northwest coast in 1828 (Species General des Coleopteres, iii, 333).
Amara remotestriata Dej.
Three specimens were brought from the mountain top. It occurs in
mountainous and northern regions of our continent.
Harpalus innocuus Lec.
- Two specimens of this species are very black, but present no other
difference.
Hippodamia 5-signata Kirby.
_A series of five specimens show great variation in size and color
the smallest one being also the darkest. One of them is remarkable
‘This department is edited by Clarence M. Weed, Hanover, N. H;
1893.] Entomology. 165
in that the thorax has a large, perfectly regular fovea on each side
near the middle.
Hippodamia convergens Guer.
One specimen showing nothing of interest.
Coccinella transversoguttata Fabr.
One specimen of an ordinary type. Widely distributed in North
America, and occurs also in Northern Europe and Asia.
Psyllobora 20-maculuta Say.
A specimen of this species is the most thoroughly melanized of any
seen. All of the elytral spots, excepting the sub-apical, are more or
less confluent, forming two irregular vitte—one near the suture, the
other discal. This would go under the variety tedata Lec., but is
larger than typical 20-maculata instead of being smaller, as is usual
with that variety.
Byrrhus kirbyi Lee.
One specimen.
Byrrhus cyclophorus Kirby. i
Two specimens. This and the preceding species are northern or
montane in habit.
Buprestis lanta Lec.
One specimen.
Podabrus xanthoderus Lec.
Four specimens of a Podabrus which agree with Dr. Leconte’sshort
description of his variety a (Trans. Amer. Ento. Soc., ix, p. 48), are
referred here. The mandibles and sides of the head are yellow, as are
also the antennal tubercles and the first two joints of the antennæ.
One specimen (possibly a distinct species) has the thorax entirely black,
except a very narrow margin on the sides in front of the middle ; it is
also a little less in size, but seems to agree in other characters.
Aphodius ursinus Mots. __ :
Two specimens of this fine insect are in the lot. It is a black variety
of aleutus, according to the latest researches and is known from Col-
orado (Dr. Horn, Trans. Amer. Ento. Soc., xiv, p. 13) and from
Kamtschatka. Compared with examples of aleutus from Southern
Alaska these latter are only about half as large.
Aphodius granarius Linn.
Four rather large specimens. This species is found over nearly all
of North America and the rest of the civilized world.
166 The American Naturalist. [February,
Aphodius congregatus Mann.
One specimen of the variety with testaceous elytra.
Acmæops proteus Kirby.
A specimen of this variable species is almost exactly like one from
Ottawa, Canada. ;
Syneta albida Lec.
One specimen. Thesuture is narrowly dark, the thorax and head
also quite dark. The species of Syneta vary in color and it is quite
unnecessary to erect new species on such characters until the extent
of this variation is more definitely known.
Gonioctena arctica Mann.
Four specimens of this interesting species were taken, two of them
nearly typical, differing from Mannerheim’s description (Bulletin de
la Societe Imp. des Naturalistes des Moscow, 1853, p. 166) only in the
absence of- the pale thoracic borders and dorsal line. The remaining
two show both of these markings very well, but differ from each other
in minor particulars. , As the species is rare in American collections I
Fig. i:
have prepared the accompanying figure, which will give some idea of
the color variations exhibited even by a small set. This species *
known from Northern Alaska, the Nelson and Churchill Rivers and
from Arctic Siberia southward to the Amur, according to Dr. Hamil-
p. 88
ton, from whose valuable paper (Trans. Amer. Ento. Soc., xvi,
et seq.) much of the distribution herein quoted is derived.
Serropalpus barbatus Schall.
One specimen occurs “ from Maine to Alaska, Central and Western
Europe, Western and Eastern Siberia.” (Hamilton.)
H. F. Wickam, Iowa City, low’
A Peculiar Seed-Like Case-Worm from the Grand oer
yon.—While collecting in the Grand Canyon, July 8 to 11, 1892,
members of our party frequently found themselves covered with num-
1393] Entomology. 167
bers of very small, thin, seed-like bodies, which became attached to
their arms, hands, clothing, and faces. Examination proved these to
be the cases of a very small larva, which lived within. They were
afterward observed to be very common on the narrow-leafed willows
(Salix sp.), a round-leafed species of leguminous tree, and on many,
other plants. They were usually found on the edges of leaves, the
Fig. 2.—Larva, dorsal view—much enlarged. (Hair line shows nat-
ural si
case being attached thereto by a few extremely fine strands of silk.
They occurred especially on the Salix, and were found at 2500 feet and
more below the rim.
An examination of this larva proves it to be lepidopterous. It has
not been bred, but the description of the larva and case given below,
together with the figures, will serve to identify it. It is peculiar as
exhibiting no trace of legs or prolegs, yet it is unmistakably lepidop-
‘terous.
Description of Larva.—Body 13-jointed, whitish, very sparsely
clothed with weak hairs; head and second segment with anal hooks,
somewhat rufous. Head narrower than second segment, & little more
than one-half as wide, subcircular in outline from above, widely and
deeply notched behind; two small approximated simple a A
extreme outer anterior edge of lateral dorsal plates of head just ac
of antennæ, the anterior one slightly larger than the —
Antenne small, short, 3-jointed ; first joint stout, a little longer t
wide, with terminal bristles on outer edge; second joint very sm
and short ; third joint a little narrower and longer than second, or
to a point at tip. Mandibles stout, narrowed and twice-
168 The American Naturalist. : February,
(3-toothed?) apically. First pair of maxillee 3-jointed, joints about
equal in length, basal joint stoutest, second and third joints decreasing
in width ; two pairs of palpi springing from inner side of basal joint,
these palpi apparently also 3-jointed, joints of equal length, basal
stoutest, second and third successively narrowed. Second pair of max-
illæ with one pair of palpi, these palpi apparently 2-jointed, first joint
moderately narrow, terminal joint elongate, bristle-like, tapering to a
Fig. 3.—a, case—natural size; b, case—enlarged ; c, portion of case—
enlarged ; d, portion of case—greatly enlarged.
point ; spinneret 3-jointed, joints sub-equal in length, basal joint stout,
terminal joint pointed. Second segment corneous, with a transverse
row of about eight roundish markings near posterior margin, and 8.
median pair just anterior to row. Other segments fleshy, the integu
ment of all segments except head microscopically warted, in the pres-
ent alcoholic specimen much wrinkled, especially on third segment ;
third segment somewhat wider than second, fourth a little narrower,
segments 5 to 10 about same width, 11 to 12 nearly as wide; segmen
2 to 12 subequal in length. Thirteenth or anal segment narrowed,
shortened, longitudinally bifid on the median line posteriorly, fur-
1893.] Entomology. 169
nished with a pair of curved claw-like corneous hooks springing from
sides of ventral surface, their tips closing on each other posteriorly.
(Ninth and tenth segments each with a black circular or oblong con-
tained body.) Spiracles showing on eleventh and twelfth segments as
small nearly circular openings about the middle of segment near lateral
margin, not apparent on other segments. No indication of true thor-
acic legs or of prolegs.
Length of larva (slightly curved as in drawing), 23 mm.; width of
third segment (second from head), about $ mm.
Description of Case.—Case is small, seed-like in shape, very flat and
thin, a little longer than wide, pale brownish in color, composed of
two thin layers sub-oval in outline, glued together on edges, narrowly
separated or free at one end, the larva living between the layers. Under
the microscope a single layer presents a reticulated structure, appear-
ing as if uniformly made up of very small irregular pieces of nearly
equal size. It is apparently composed of vegetable tissue, probably |
very minute fragments of leaves.
Length of case, about 33 mm.; greatest width 2 mm., or hardly
more.
C. H. TYLER TOWNSEND.
Entomological News.—At the meeting of the American Asso-
ciation of Agricultural Colleges and Experiment Stations held in New
Orleans, Nov. 15 to 18, Chairman Lawrence Bruner, of Nebraska, and
Secretary F. M. Webster, of Ohio, of the Entomological Section, were
reelected for another year. But few entomologists were in attendance
at the meeting, most of the Station entomologists seeming to prefer the
Association of Economic Entomologists for the presentation of papers
in reference to their work. The report of the Chairman of the Sec-
tion of Entomology, Prof. Lawrence Bruner, was read by Prof. Osborn.
The report largely consisted in a statement of the equipment of the
Station entomologists as a whole, to which was added a statement in
detail of the entomological equipment of each station having an ento-
mologist. The report showed that while some of the stations were,
well equipped with laboratory, insectary, and other requirements for
work, yet there were many stations which had not given entomolo-
gists proper equipment. Some of the Station entomologists lack -e
for laboratory, others library, and many, microscopes. Valuable resu
of a practical as well as a scientific nature are rarely attained without
proper apparatus, and the Station entomologists need something more
than jack-knives with which to carry on their work.
12
170 The American Naturalist. ' [February,
It is gratifying to learn of the advancement being made by many of
the entomologists of the experiment stations. Prof. H. A. Morgan, of
Louisiana, and Prof. Popenoe, of Kansas, who were formerly entomol-
ogists and horticulturists of their respective stations, have, upon
their urgent request, been made entomologists of their stations only.
‘This is a step in the right direction.
Prof. John B. Smith will have charge of the entomological part of
the biological exhibit of the office of experiment stations at the
World’s Columbian Exposition.
Howard Evarts Weed, of the Mississippi Agricultural College is
making a special study of North American Myriopods, and would be
glad to exchange insects of any order for Myriopods of any locality,
especially the west. His address is Agricultural College, Mississippi.
Prof. C. H. T. Townsend is preparing a monograph of North Amer
ican Tachinidz, and desires material in this family from any locality,
especially bred specimens. His address is Las Cruces, New Mexico.
A recent issue of Psyche says: “ Entomologists everywhere will
regret to hear that the serious illness which has, for the past two years,
incapacitated Dr. H. A. Hagen, renders it improbable that he will be
able to do any further work. Dr. Hagen has had charge of the col-
lections of insects in the Museum of Comparative Zoology at Harvard
University since Oct. 12, 1867, and during this long period of twenty-
five years has applied himself with entire devotion to the interests of
the department. The scientific value and present excellent condition
of the collections are the result of his faithful and disinterested work. i
Recently the department has been placed in charge of Mr. Samuel
Henshaw.”
Dr. M. C. Cooke, the well-known English mycologist, has brought
together an account of the fungi parasitic upon insects in a volume
recently published by the Society for Promoting Christian Knowledge
(London, 1892).
We learn from The Entomologists Monthly Magazine that Mr. Oswald
- Latter reported at a recent meeting of the London Entomologics
Society that the moth Dicranura vinula produces, probably from
mouth, a solution of caustic potash for the purpose of softening the
cocoon.
An interesting summary of the value of entomological study bas
been sent out by Mr. James Fletcher in the report of his Evidot
mons.
before the Committee on Agriculture of the Canadian House s “
1893.] Psychology. 171
PSYCHOLOGY.
Vision in a Young Girl Six Years of Age, Operated Upon
for Double Congenital Cataract.—M. Grafé has published in the
Revue Scientifique July 16, 1892, an interesting account of an opera-
tion for double cataract, performed by M. Bribosia upon a blind girl
5 years of age, and discusses, in the same paper, some questions of
sight psychology. This reminds me of a similar operation performed
by myself several years ago upon a young girl 6 years old, born with
a double white cataract. Some notes of that operation, which I have
kept, will perhaps be of use to M. A. Grafé in the researches he has
undertaken in a new field.
What are the sensory impressions and emotions of a person born
blind when the veil which has hidden from him the marvellous world
is suddently rent? Would he, at first sight, see, as has been taught,
objects reversed, in consequence of the crossing of the rays of light
` from the object before their contact with the retina? Would he have
intuitively an idea of distance and of the third dimension of matter,
or would this idea be gained by observation and experience?
In order to succeed in solving the problems of optical physics and -
visual psychology, M. A. Grafé arrived a little late after the operation
by M. Bribosia. I had opportunities in the case of the child 6 years
of age, more advantageous for studying the curious phenomena which
took place, not only during the operation, but also during the dressing
of the wound and the first exercise of the newly acquired sense.
The following are the notes of the operation :
Julie D. had a double white congenital cataract at the age of 6 years.
She perceived less and less from infancy the light of day, even after
the local use of belladonna. Being intelligent, she had acquired a
variety of knowledge from conversation. Touch and hearing had
become very sensitive, and her memory was excellent.
I operated upon both eyes at the same time, as was my custom with
double cataract in old people as well as young. But, in order to avoid
a sudden flood of light, I operate always in a slightly darkened room ;
and before attending to the second eye, I hermetically seal the first
with a simple bandage. The dressing of the wound for the first few
days is done in the same manner in a similarly darkened room.
When I allowed Julie D. for the first time to remove the bandage
from her eyes she had a slight sensation of dizziness, which made her
172 The American Naturalist. [February,
put her hands to her face. Then she saw at a little distance objects
which she recognized by their shape; a coffee-mill, my hands, knives,
forks, a plate, a cup, a watch, a handkerchief, etc., the forms of which
she had become familiar with through touch during her long blindness,
But she could not distinguish color. She would make hap-hazard
guesses at red, green, blue, and yellow, but could sometimes recognize
black and white. I was anxious to know her first optical impression
as to the position of objects, whether they appeared right or reversed.
With this idea in mind I showed her, first of all, a coffee-mill, held at
the distance of a meter from her, and said, “ What is this?” “My
coffee-mill,” said she without hesitation. ‘How do I hold it, with the
drawer on top or on thebottom?” “It is below, as it ought to be.”
Thinking that the child was answering in accordance with her precon-
ceived notions contrary to her first visual sensation, I showed her a
knife fixed vertically by two threads to a sheet of white paper, aid
asked her to show me with her hand where was the handle of the knife,
at the top or the bottom of the paper. She pointed immediately with
her right hand to the handle, saying: “It is there, at the top.” This
was correct.
I did not indulge my scientific curiosity further at that time. At
the second and third interviews which took place the sixth and seventh
. day after the operation, Julie was no longer deceived as to colors, but
she still imagined as she had at first, that everything she saw was
within reach. Space, height and volume did not as yet exist for her.
However, she appreciated perfectly the distance of sounds. When
objects were held for her at less than a meter’s length, she would -
instinctively draw back, feeling, as she said, that they would strike her
eyes. As to everything she saw that was at more than arm’s length,
they all seemed ranged on the same line, that is, all seemed equally
distant. But this illusion did not last dong. The sense of touch had
- given her such just ideas of the form, the size, and the different dimen-
sions of buildings, persons and the objects in the midst of which she
had lived, that ideas of perspective and appreciation of distances were
speedily acquired. os
At the end of fifteen days she said she had almost forgotten that she
had ever been blind.
Had she, in the presence of what we call the wonders of nature,
the flowers, the trees, the monuments, the country, the storms, the sky
with its myriad stars, the sun and the moon, that awakening of ~ n
imagination roused by a spectacle as magnificent, as unexpected, whic
expresses itself in an ecstatic admiration? She was astonished at all, A
1893.] Psychology. 173
but she admired nothing. Had she dreamed, perhaps, during that
long night of six years more and better things than the realities which
unfolded themselves before her eyes?
It is a fact that the persons who have lost their sight, and recovered
it through an operation, have shown more pleasure at seeing again
their relations, their friends, and their coffee-mills than did Julie D.
at seeing for the first time all the objects of which she could have had
only a feeble and imperfect idea gained either through the sense of
touch or from descriptions given her by others—H. Boéns, Revue
Scientifique, Oct. 29, 1892.
ARCH ZOLOGY AND ETHNOLOGY.
Area and Population of European Countries.—M. E. Lav-
asseur, in a communication to the Academie des Sciences on March
21, 1892, calls attention to the diversity usually to be found in statis-
tics of area and population in standard works of reference. Consider-
ing only such high class publications as “ Die Bevölkerung der Erde,”
the “Almanach de Gotha,” and the “Statesman’s Year-Book,” he
points out that diversity does not necessarily convict any of error, as
the approximation to the exact figures may be arrived at in different
ways; that, in fact, absolute agreement in statistics is a sign that they
have been copied by one writer from another. In some cases the areas
of countries are officially derived from cadastral surveys, which, as a
matter of fact, often do not include the whole land and water area of
a country, geographers not being agreed as to what water areas should
__ be reckoned along with the land. In other cases official or semi-offi-
cial measurements are made on large-scale topographical maps, and the
degree of approximation must vary with the scale, and with the tech-
nical skill of the computor. Some countries are content with meas-
urements or estimates made by individuals, such as those of Strel-
bitsky and of Perthes’ Institute. The calculation of population is
still more uncertain, being dependent on census returns (themselves
imperfect), calculated to any given intermediate date by eeguesemnaes
derived from registers of births, deaths, immigration and emigration.
In a table brought down to the end of 1890 M. Levasseur gives a con-
Spectus of the area, population, and density of population of the
countries of Europe, classed in four groups, as follows :
174 The American Naturalist. [February,
Density
Inhabitants. Area, sq. m. per sq. m.
Western Europe......... 87,109,000 352,300 247
Central Europe........... 93,609,000 464,400 200
Southern Europe......... 71,826,000 557,800 130
rn Europe........+. 98,000,000 2,106,500 47
Northern Europe......... 9,100,000 378,000 23
All Europe 359,635,000 3,859,200 93.1
Notes as to the various estimates are appended ; those relating to the
German Empire may be cited as an example. The “ Bevölkerung der
Erde” in 1872 gave the area as 207,816 square miles ; in 1874 as m
935; in 1876 as 207,943; in 1878, as in 1880 as 207,883;
1882 as 207,899; in 1891 as 207,861; while Strelbitsky’s measure-
ment is 208,008, and that adopted in the tables from the “ Statistisches a
Jahrbuch fiir deutsches Reich” for 1891 is 207,929 square miles— —
Proceeds. Roy. Geog. Soc., July, 1892. i
t
A Measure of Civilization.—Dr. Lamborn suggests that the
proportionate length the artist gives the face or head to the body
might be considered a measure of civilization. He finds that the Jais
Spanish artists give 103 faces to the height. Dürer 10 to 1.
Spanish artists 9 to 1. The measurements of a number of wood cuts
of the Fifteenth Century run 5} to 1 and 6} to 1. The Mexican
images are 4 to 1, 3} to 1, and 3 to 1. The gold figures of Peru masè
the head larger than the body. (Lamborn’s History of Mexican Art)
1893,] Microscopy. ` 175
MICROSCOPY
On the Restoration of Osmic Acid Solutions.—The
osmic acid so valuable to the histologist as a fixing reagent, is
known to the chemist as osmium tetroxide, Os O,, and is one of the
series of osmium compounds with oxygen. It is readily reduced in
the presence of an organic substance, giving up two atoms of oxygen
and forming the deutoxide, Os O,. This is the well-known black pow-
der so familiar in specimens killed in any of the fluids into which osmice
acid enters. Osmium tetroxide affects fatty substances first, and these
are the substances that most readily undergo oxidation. Prolonged
exposure to the action of the tetroxide blackens the tissue so that it is
useless for histological purposes.
In preparing stock solutions of osium tetroxide the greatest care
must be taken to exclude organic matter, even acetic acid in the quan-
tity called for in Flemming’s stronger fluid, is liable to cause the
reduction of the tetroxide. A trace of impurity in the distilled water
sooner or later produces the fatal blackening, while a solution made
with pure water will keep indefinitely in the light. Let but a trace
of dust fall into the bottle and reduction will take place to some
degree.
During the summer of 1892, at the Marine Biological Laboratory
at Woods Holl, Mass., a large number of solutions were treated with
peroxide of hydrogen, H, O,, and different investigators tested them
with uniformly good results. The restored solutions acted in the same
manner as the fresh solutions, and produced the same results ; the only
noticeable difference was a gradual weakening of the solution, as
would naturally be expected. Specimens overblackened by the action
of the peroxide were rendered colorless without perceptibly changing
the character of the tissue, and a few experiments were made to deter-
mine whether the bleaching with peroxide could be so controlled as to
leave certain tissues blackened while others were cleared. This quan-
titative bleaching with peroxide yielded no very good results in the
few experiments made, but enough was accomplished to warrant further
attem
cacy of the restoration and the bleaching further demon-
strates that no injurious factor enters into the result as is shown by the
following equations :
This department is edited by C. O. Whitman, University of Chicago.
176 The American Naturalist. [February,
Os O, + organi¢ substance = Os O, + oxidized organic substance.
Os O, + 2H,0, = Os O, + 2H,0. .
That is, in bleaching a tissue, the Os O, is converted into Os O, and
water is formed, with which the tissue is already saturated.
To restore 100 cc. of a 1% solution of osmium tetroxide requires
from 10 to 20 drops of fresh peroxide of hydrogen, and clearing goes
on better in full sunlight. To bleach tissue with peroxide requires
about the same amount in 100 ce. of water, and the bleaching takes
place better in the sunlight also. The amount needed cannot be stated
with any more precision because the peroxide loses its strength some-
what with age. The peroxide made by the Oakland Chemical Co., 54
South Avenue, New York, is more stable and keeps better than many
of the preparations in the market.—C. L. BRISTOL.
PROCEEDINGS OF SCIENTIFIC SOCIETIES.
The Geological Society of America.—The Fifth Annual
Meeting was held at Ottawa, Canada, on December 28, 1892, and fol-
lowing days. The following papers were read: Notes on the Early
Palæozoic Sponges, Sir J. William Dawson, Montreal, Can.; Noteson
the Devonian Formation of Manitoba and the N. W. Territories, J.F.
Whiteaves, Ottawa, Ont. The object of this paper was to show the
relationship of Devonian formation of Manitoba and the N. W. Ter-
ritories with that of Northern Europe. In Manitoba the Stringoceph-
alus zone is remarkably well developed and holds a rich fauna, whereas
in the Mackenzie River district, most of the fossils so far collected
seem to be from the Cuboides zone. On the Coals and Petroleums of ‘a
the Crows’s Nest. Pass, Rocky Mountains, A. R. C. Selwyn, Ottawa,
Ont.; On the Geology of Natural Gas and Petroleum in Ontario, H.
P. Brumell, Ottawa, Ont. A brief résumé of work done in gas and
oil explorations in Ontario, and dealing with the geological evidence 5
adduced from record of wells sunk. Including also a short account of
the geology of Southwestern Ontario. Some Features of the Phos
phate Bearing Rocks of Ottawa County, P. Q., Elfrie Drew Ingall, :
Assoc. Royal School of Mines, Eng. (Introduced by J. B. Tyrrell.)
This was a preliminary paper, based upon field observations, discussing
_ the phenomena exhibited by the Apatite deposits and their associated —
rocks in their bearing upon the question of the probable mode of for
mation of these deposits. Some Maryland Granites and Their Origm,
Charles Rollin Keyes, Des Moines, Ia. A brief sketch of the granite
1893.] Proceedings of Scientific Societies. 177
rocks of Marylend, with a summary of the reasons for regarding them
as eruptive in origin. Epidote as a Primary Component in Granites,
Charles Rollin Keyes,!Des Moines, Ia. Occurrences of the mineral in
certain granites regarded as eruptive described, the evidences of its
original nature explained and its associations with the closely related
Allanite considered. On the Laurentian of the Ottawa District, Rob-
ert W. Ells, Ottawa, Ont. The paper briefly reviewed the work of
the late Sir Wm. Logan in this district, forty years ago, and discussed
the structure of the calcareous portion of the system, more partic-
ularly as regards the relationship of the crystalline limestones to the
underlying gneiss; with a brief statement of the several kinds of
intrusive rocks which penetrate both the gneiss and limestone, and
their apparent influence upon the occurrence of the economic minerals,
such as graphite, apatite and mica. The Contact of the Laurentian
and Huronian North of Lake Huron, Robert Bell, Ottawa, Ont. The
line of contact between the Great Huronian Belt to the northward
and the Laurentian area to the southward was traced from near Killar-
ney on Lake Huron to its intersection with the Ottawa River. The
two sets of rocks in this region were described with regard to their
structure and their relations to one another. The nature of their con-
tact here and in other instances in Canada was described in connection
with the question of the conformability or otherwise of the two sys-
tems. Reference was made to tl t linary develop tof quartzit
in the LaCloche region, to certain forms resembling fossils lately dis-
covered by the writer in the Huronian rocks of the vicinity, to the
greenstone masses and dykes, to the nature of the granites and to other
points in connection with the geology of the district. The marked
dependence of the physical geography upon the phenomena of erys-
talline rocks of the region was likewise pointed out. The Archean
Rocks West of Lake Superior, W. H. C. Smith, Ottawa, Ont. (Intro-
duced by R. W. Ells.) This paper gave a brief description of the
rocks and their distribution between the Lake of the Woods and Lake
Superior north of the International boundary and referred to some
of the theories of origin and structure of the various members of the
Archean system and to the iron orés and gold-bearing rocks of the
region. On the Archean of Sudbury Mining District, Alfred E.
Barlow, M. A., Geological Survey Department, Ottawa, Ont. This
paper related in detail the various phenomena in connection with
the contact between the Laurentian and Huronian to the northeast of
Lake Huron, with a discussion of the origin and relative age of each,
based on these observations. Notes on Cambrian Fossils from the
178 The American Naturalist. [February,
Selkirks and Rocky Mountain Region of Canada, Henry M. Ami,
Ottawa, Ont. This paper was based mainly upon a collection of Lower __
and Middle Cambrian fossils made by the author in the summer of
1891. It contains notes on some eight species of Lower Cambrian
(Olenellus zone) fossils from the gray, glossy and calcareous schists
and limestones of the entrance to the Selkirks, some two miles west of
Donald, British Columbia. The latter part of the paper dealt with
the forms met with in the Middle Cambrian of Mount Stephen in the
Rocky Mountains, near Field, B. C., where the terrane is highly fossil- _
iferous. Upwards of twenty species have been recorded from this
locality, many of which are very interesting and well preserved. On |
the Potsdam and Calciferous Terranes of the Ottawa Paleozoic Basin,
Henry M. Ami, Ottawa, Ont. The stratigraphical, lithological and
paleontological relations of the Potsdam and Calciferous terranes, as 3
seen and known in the Ottawa palæozoic basin and elsewhere weredis- |
cussed in this paper ; also the reference of these two terranes to the
Cambro-‘Silurian or Ordovician Epoch instead of to the Cambrian
Epoch, inferred from the internal evidence. The Criteria for the Rec-
ognition of Separate Ice Epochs, R. D. Salisbury, Chicago, Ill.; Note
on the Geology of Middleton Island, Alaska, George M. Dawson, a
Ottawa, Ont. This short paper was devoted principally to the descrip- a
tion of a boulder-clay or till from Middleton Island, which is foundto — n
contain some marine fossils. Eskers Near Rochester, N. Y., Warren
Upham, Somerville, Mass. The Pinnacle Hills, a very remarkable
esker series in the southeastern suburbs of Rochester, were described |
and attributed to deposition by a glacial river flowing between wals
of ice and open above to the sky. Their material, which is shown "
have been englacial, is chiefly gravel and sand, but also comprises m
some parts very abundant and large boulders, some of which are lified
200 feet or more above their sources within a few miles on a nearly —
plain country. Similar explanations are also applicable to other oo
eskers in Pittsford, several miles farther southeast. Comparison of a
Plistocene and Present Ice-Sheets, Warren Upham, Somerville, Mas. a
The Plistocene ice sheets of North America and Europe were compare? a
with the now existing Malaspina, Greenland, and Antarctic ice-sheet :
as to their areas, surface slopes and probable thickness, rates of erosioP
and ablation, subglacial and englacial drifi, and manner of deposition a
of various drift formations. The Malaspina glacier or ice-sheet, COW
`. ered on its wasting borders by much drift and growing forests, we
believed to afford explanations of forest beds between deposits fe
till, and of the peculiar drift accumulations named drumlins, both
1893.] Proceedings of Scientific Societies. 179
being attributable to stages in the general recession of the North
American ice-sheet when increased snowfal] and onflow of the ice slack-
ened its retreat or caused it temporarily to re-advance. Under this
view, the Ice-Age seems probably to have comprised only one great
epoch of glaciation, with moderate oscillations of the ice-front, and to
have been geologically brief. Plistocene Phenomena in the Region
Southeast and East of Lake Athabaska, Canada, J. B. Tyrrell, Ottawa,
Can. The paper was the result of an exploration conducted by the
writer during the past summer in the hitherto unexplored region lying
southeast of Athabaska Lake and north of Churchill River. The
region has some strongly marked glacial features. The writer dis-
discussed the striation and character of the rock surfaces, the occurrence
of till, drumlins, kames, and other glacial phenomena, Some high-
level beaches, terraces, and other post-glacial deposits were also noted.
Notes on the Glacial Geology of the Northeast Territories, A. P. Low,
Ottawa, Ont. A short account of the geography and surface geology
of the western water-shed of Labrador Peninsula, derived from explor-
ations on the Rupert; East Main, Big, Great Whale and Clearwater
Rivers, and along the east coast of Hudson Bay. Notes on the Gold
Range in British Columbia, James McEvoy, B. A. Se., Ottawa, Ont.
(Introduced by H. M. Ami.) A short description of the topography
of the Gold Range and part of the adjoining Interior Plateau country,
with notes on the glacial geology of the same. The Glacial Gravels
of Glacier Bay, Alaska, Harry Fielding Reid, Cleveland, Ohio (Intro-
duced by J. S. Diller). The Post-glacial Outlet of the Great Lakes
Through Lake Nippissing and the Mattawa River, G. Frederick
- Wright, Oberlin, O.; The Height of the Bay of Fundy Coast in the
Glacial Period Relative to Sea Level, as Evidenced by Marine Fossils
in the Boulder Clay at Saint John, New Brunswick, Robert Chalmers,
Ottawa, Ont. Description of locality, area and thickness of boulder-
clay. Divergent strie on underlying rocks. The materials of the
boulder-clay from the north. Intercalated stratified portions and their
fossils deposited in the sea. Sections at Fern Ledges and Negrotown
Point given. The conclusions are that the climate was Arctic or sub-
arctic; that the boulder clay here was formed by a number of suc-
cessive accretions in a zone of oscillation of the ice-front, and that its
upper portion at least was thrown down when the lands stood atom
200 feet below its present level. On Certain Features in the Distribu-
tion of the Columbia Formation on the Middle Atlantic Slope, N. H.
Darton, Washington, D. C. This paper was a description of rela-
tions indicating an interval of erosion between the depositions of the
180 The American Naturalist. [February,
high-level and low-level portions of the formation, beginning in South-
ern Maryland and gradually increasing northward to New Jersey. A
Geological Reconnoissance in the Central Part of the State of Wash-
ington, Israel C. Russell, Ann Arbor, Mich. The region traversed
embraces about 10,000 square miles in the arid region east of the cas-
cades and drained by the Columbia. The reconnoisance was made for
the purpose of learning how far the conditions there existing favor the
project of obtaining artesian water for irrigation. The following is
a syllabus of the paper.
Geography of the region explored.
Geological formations.
Crystalline rocks. |
Columbin toe } Tertiary. Glacial records, ) Pleistocene.
ake Lewis,
Geological structure. Existing lakes. Artesian water supply.
The Abandoned Strands of Lake Warren, Andrew C. Lawson,
' Berkeley, Cal. The strands of Lake Warren, on the north side of
Lake Superior, up to an elevation of 1200 feet above sea-level are
post-glacial. It was not an ice-dammed lake. There was an outlet
northward corresponding to one of its higher stages. A post-glacial
depression of Central Canada whereby the James Bay slope was covered
with marine sediments to a present altitude of 450 feet above tide and
only 150 miles distant from Lake Superior is correlated with the max-
imum fullness of Lake Warren, and the subsequent uplift is correlated
with its subsidence. The strand lines show no evidence of deforma-
tion. In the absence of ice dams and of a gorge of permanent drain-
age, the Level of Lake Warren could only have been lowered by
epeirogenic depression along its southeastern margin, i. e., in the region
south of Lakes Huron and Michigan, which depression is thus coeval —
with the post-glacial uplift of central Canada. High terraces and
beaches are known to extend along the north side of Lake Huron from
Sault St. Marie eastward and they are reported on the high lands of
the Peninsula of Ontario; so that Lake Warren must have been the
greatest of the known Quaternary Lakes. Its area is roughly estima-
ted at 150,000 square miles. The Importance of Photography in
Illustrating Geological Structure, R. W. Ells, Ottawa, Ont. In con-
nection with this paper was shown a series of large photographs by
Topley, taken under the direction of the writer, showing more clearly
the mode of occurrence of the apatite in the deposit of the Bucking- |
ham and Lievre district, as well as the relations of the intrusive
1893.] Proceedings of Scientific Societies, 181
apatite-bearing rocks to the surrounding gneiss. Two Neocene Rivers
of California, Waldemar Lindgren, Washington, D. C. It wasthe pur-
pose of this paper to briefly indicate the direction and grade of the
principal forks of the Neocene equivalents of the present Yuba and
American Rivers. These streams now drain an area in which many
important Neocene gravel deposits are located, and to trace the old
water courses is of the highest importance in order to successfully open
and mine the auriferous channels. The results recorded here have
been reached during the course of the investigation of the Gold Belt
carried out under the direction of Dr. G. F. Becker, by the U. 8.
Geological Survey.
The geological events related to the accumulation and covering up
of the Neocene channels were briefly sketched and the different systems
of channels described. The fact was accentuated that the Neocene
rivers closely correspond to those of the present day in the extent of
their watersheds, although by no means always in the direction of
their principal forks. A short statement was given of the work pre-
viously done in the same field by other investigators.
The course of the Neocene Yuba and American Rivers was des-
cribed, by the aid of a map, on a small scale. In places where
differing results have been obtained from those of other observers, or
where new facts have been brought out, a more detailed statement was
given. Practical results : The greater part of these Neocene channels is
now covered by heavy masses of volcanic material and many of them,
though auriferous, have not yet been opened for mining operations.
The vontinuity of certain channels under the lava cap can be asserted
and their approximate positions indicated ; some of them, most likely
to prove remunerative, but which have not yet been opened up, were
mentioned; among them are the Blue Tent Channel, the Long
Canyon Channel, and the Webber Creek Channel.
Boston Society of Natural History.—Dec. 21.—The follow-
ing papers were read: Some New Experiments on the Absorption of
Liquids by Aerial Parts of Plants, Mr. W. F. Ganong; The Abdom-
inal Pouch of Butterflies of the Genus Parnassius, Mr. 8. H. Seudder;
Columnar Structure in Stratified Rock, Prof. W. H. Niles.
- January 4, 1893.—The following papers were read : Account of Some
of the Botanical Establishments of Europe (photographs were shown),
Prof. W. G. Farlow ; Application of the Microscope to the Study of
Rocks (colored stereopticon views were shown), Dr. J. Eliot Wolff.
SAMUEL HENSHAW, Secretary.
182 The American Naturalist. [February,
New York Academy of Sciences, Biological Section.—
Meeting of Dec. 12.—The following is a synopsis of the papers: On
the Miocene Deposits of the White River, by Dr. J. L. Wortman;
These deposits were arranged in three groups, lower or Menodus beds,
middle or Oreodon beds, and upper or Protoceras beds. The Protoceras
beds were regarded as in part contemporary with the John Day beds
of Oregon. On the ileo-colic junction of Procyon lotor and allied
Arctoids, by G. S. Huntington. The absence of cecum in Procyon
was noted as repeating the condition found in Hyzena and the Urside.
The provision for preventing return of contents of large intestine
appears to consist in a series of constrictions in the terminal part 0
the ileum together with increase in the circular muscular fibres in
these situations as well as at the ileo-colic junction itself. There is a
complete absence of an ileo-colic valve. On the Origin of West Indian
Bird Life, by F. M. Chapman. Conclusions from study of bird and
mammal life were (1) distinctness geologically of Lesser from Greater
Antilles ; (2) independence of islands from mainland since the appear- —
ance of the present fauna; (3) original connection of West Indies to
Central America by way of Jamaica ; Central America at this time an
archipelago created by passage leading from Pacific to Carribean Sea;
(4) the older faunal forms of the West Indies represent survivors of the
insular Tertiary species ; (5) the newer forms are immigrants and become
differentiated under new conditions of living. H. F. Osborn reported
the discovery in the Miocene of South Dakota of a horned artiodac-
tyle represented by male and female skulls and complete fore and hind
fect. The female skull is comparatively hornless and proves to be
identical with Protoceras celer Marsh. The male skull exhibits no less
than five protuberances upon each side or ten altogether. Two of these
upon the frontals and sides of the maxillaries are very small ; the
parietal, supraorbital and maxillary protuberances are very promi-
nent and had apparently a dermal covering as
are four toes in front and two behind as in the early
types were found by Dr. J. L. Wortman, and are in
tions of the Am. Mus. Nat. Hist.
Basurorp DEAN, See. Biol. Section.
Tragulide. The
the recent collec:
Natural Science Association of Staten Isla
1892.—The election of officers for the ensuing year resulted as follows:
President, Walter C. Kerr ; treasurer, Thos. Craig ; recording secretary»
Joseph C. Thompson ; corresponding secretary, Arthur Hollick. A
paper on Our Historic Landmarks, by Mr. Ira K. Morris, was by
in the giraffe. There
EA E,
A E RERE ANT EE E E PEN
a ea ee Su REY
nd.—Nov. 1%
1893.] Proceedings of Scientific Societies, 183
title, and will be published as a“ special” later on, Dr. N. L. Britton
presented specimens of Aspidium cristatum (L.) Sw., collected by Mrs.
N. L. Britton, near Oakwood. This fern is exceedingly rare on Staten
Island, having been previously reported from but two localities, in one
of which, the Clove Lake Swamp, it has been since exterminated.
Mr. Arthur Hollick read the following paper Upon Additions to the
Cretaceous Flora of Staten Island.
At the meeting of January 2, 1892, I gave a preliminary account
of the Cretaceous fauna and flora of the island so far as studied up to
that date. At the meeting of March 12, the rediscovery of fossil
leaves in the Kreischerville clays was recorded, and also the discovery
of a large fossil leaf near Richmond Valley. At the meeting of
September 10 the discovery by Mr. Heinrich Ries of fossil léaves in
the Cretaceous clay at Green Ridge was recorded and since that time
Mr. Wm. T. Davis has turned over to me a number of other specimens
found by him in the clay at Kreischerville.
All this new material and such of the old as had not been critically
examined has been the subject of careful study by me during the past
few months with the result that there has been added at least thirty-
eight species to our Cretaceous flora.
Most of them were known previously from the clays of Woodbridge
or Amboy, N. J., but others have not before been reported from East-
ern North America and we here have their first recorded occurrence in
this region. Of these fourteen represent species new to science.
The specimens have all been shown from time to time at our meet-
ings, and from these careful drawings have been made. The entire
subject will be presented before the New York Academy of Sciences
shortly, with full descriptions and figures of all the new species, which
will be published in the Transactions in due time.
We have forty-seven species representing the flora of the Cretaceous
formation on Staten Island: All the specimens from Tottenville,
Richmond Valley, Princes Bay and Arrochar are in ferruginous rock
of a concretionary character and are well preserved. Those from the
clays of Green Ridge and Kreischerville soon began to disintegrate
upon exposure to the air and are now mostly useless for purposes of
identification. <
Dec. 17, 1892.—A specimen of European “Gorse” or “ Furze,” sent
by Dr. F. Hollick, was shown and the following memorandum read :
The accompanying specimen was obtained Nov. 10, from a wren
bush, about a foot in height, of European furze (Ulex mop ),
Which is growing wild on Ward’s Hill, Tompkinsville. I first discov-
184 The American Naturalist. (February, a
ered it during the early autumn. To all appearances the plant is
self-seeded, and is remote from any garden or cultivated ground. The
only other vegetation in the vicinity is short stunted grass, so that the
bush in spite of its small size, is a conspicuous object and has probably
only escaped destruction by reason of its sharp prickly foliage. The
locality is strikingly similar to the “commons” or “downs” where it
grows in Great Britain. How it came here is a mystery, as the only
other place where it is known to occur in America, so far as I am
aware, is on the Island of Nantucket, where it was introduced about :
1860, and has since spread to some extent. If protected and encour- 7
aged it would doubtless become established here and spread as it has
done in the latter locality.
Mr. William T. Davis exhibited Cecropia cocoons which had been
partly destroyed by woodpeckers, and read the following paper on
woodpeckers and cecropia cocoons.
The caterpillars of the Ceeropia moth spin their cocoons in a variety
of places; occasionally on the flat side of a board fence, sometimes at :
the base of bushes such as the elder, and sometimes at the ends of sway- a
ing branches when the food plant happens to be a tree. The cocoons
spun near the ground are often devoured by mice that gnaw through
the silken coats to the edible pupa within. Specimens,which had been
thus destroyed were shown to this Association by Mr. Joseph C.
Thompson on Dec. 12, 1889. Cocoons that are placed on tree branches
are more safe from the attacks of mice, but are liable to be eaten by
woodpeckers.
On January 14, 1888, I saw a downy woodpecker investigating a
Cecropia cocoon in a white maple, the woodpecker thrusting its bill i
and pulling it out of the cocoon quite frequently. After a while it
flew to another cocoon a few feet away, but it being on such a small
branch it was unable to successfully pick it open, as the branch swayed —
up and down. It was then plain what a great protection it was to the —
insects to place their cocoons near the branch ends, though no doubt |
they are sometimes killed by the swaying of these branches during &
storm. ; L
_ When the woodpecker was gone I cut the cocoon off and found a —
small hole on its side quite near the branch, where it was easiest to $
drill because the silken fabric gave way the least to the strokes of the
bird. Cutting open the other side of the cocoon, I found that the pup#
shell was sucked nearly dry of its contents. A
1893.] Proceedings of Scientific Societies. 185
_ The Cecropia cocoons, occurring commonly on white maples, are
generally placed near the ends of the long drooping branches, and it
will be seen from the foregoing that it is probably the safest situation
afforded by the tree. Near my home there isa small white maple that
has eight Cecropia cocoons on its branches, but only two of them, from
their position, can be injured by a woodpecker.
If a woodpecker is successful in making a hole into a cocoon it is
nevertheless sometimes disappointed at its contents. I have found a
cocoon that contained the tough case of an Ichneumon fly pupa
(Ophion), which had been drilled in the side by a woodpecker, and
then abandoned, leaving the parasite unharmed.
Mr. Davis also submitted the following note:
The tawny thrush or Veery (Turdus fuscescens), has not been-
reported as nesting on the Island, but during the last spring and sum-
mer it was not uncommon at Watchogue. Several would often be
heard singing at the same time, or be seen walking, hopping or run-
ning, for they do all three, along the wood paths and stopping by the
way to turn over a countless number of dead leaves for the insects to
be found beneath them. In every instance they frequented the edge
of the close timber near to some open or half cleared ground, whither
they often flew.
On June 26 I noticed a Veery carrying food in her bill, and was
thus enabled, after a time, to discover a young bird perched on a log
in a thick growth on the edge of a’ swamp. On July 10, with Mr.
Walter Granger, of the American Museum, I heard many Veeries
singing, but we were unsuccessful in finding a nest of the second brood.
It seems probable that this thrush has only been plentiful at Watch-
ogue during the past summer, for just as close attention has been given
to the district for a number of years, and yet only once before, namely
in May, 1891, was a Veery heard singing.
Mr. Arthur Hollick read Notes on Staten Island Clays collected for
the World’s Fair. Le
Mr. L. P. Gratacap exhibited berries of Smilox rotundifolia and
submitted a memorandum in connection with them. an
Mr. Gratacap also exhibited drawings from microscopic examination
of rocks to which seaweed had been attached, and read the following
note:
The great quantity of rock used in constructing the roadbed of the
Rapid Transit Railroad along our north shore has introduced upon the
island a very representative collection of N. Y. Island rock, including
mica schists, gneiss and granite. It serves the double purpose afford-
15
186 The American Naturalist. [February,
ing the local mineralogists an opportunity to study and collect feldspar,
' quartz and mica, with associated garnets and tourmalines and is a
suggestion as well of the underlying reefs of archæan metamorphic
masses upon which our own island is built, and our geological relation-
ship with New York from whose ledges the rocks have been trans-
ported. Wherever these fragments have rolled down to the waters’
edge or are so placed as to be inundated by the high tides, a very vig-
orous growth of the green filamentous seaweed (Calothrix?) has
appeared. I am not sure that it is more marked than the similar
growth over the traps, sandstones and shales which were the previous
occupants of the shores of the Island, and which have been somewhat
displaced from their intimacy with the waves and currents of the Kill
van Kull by these later arrivals, but it does seem to me as if it had
developed with interesting rapidity. The schistose flaky character of
many or most of these fragments may have conduced to facilitate this
luxuriant development; and where, as in the case of granite veins, the
feldspar is coarsely brecciated, the interlacing lines of angular defini-
tion between the crystals offer a crevice for attachment. As a fact the a
lamellar fragments are more quickly invaded. a
Placing under a microscope flakes of mica, feldspar and quartz, we :
can see the delicate filaments of the alga penetrating the films of the
mica or occupying pits and irregular surfaces of the feldspar with dis-
tributed spots of green granules. Sometimes lacunæ or minute cavi-
ties, situated a little way within the edge of mica flakes, will be filled
with the green spore-like groups. A few rude drawings show approx-
imately some of the phases in this process of investment of the rock
by the seaweed. say
Cumulative effects have of late been much emphasized, and it is not cs
unlikely that the alternating drying and swelling, from exposure to the —
sun at low and submergence at high water, of this alga may sensibly
determine the duration, as a mass, of the more schistose or friable
rocks, when the tiny extensions of this subtle enemy have effected an
entrance within their substance. : ge
The American Psychological Association—Met at Phila-
delphia, University of Pennsylvania, Tuesday and Wednesday, Dee. .
27 and 28, 1892. On Tuesday, Dec. 27 there was a business meeting
at which the nature of a permanent organization and other important -
questions will be discussed. The following papers were read : :
Errors of Observation in Physics and Psychology; Prof. J. McK. —
Cattell, Columbia College; Experiments Upon Pain, Dr. Herbert —
1893.] Proceedings of Scientific Societies. 187
Nichols, Harvard University ; Tactile Estimates of Thickness, Prof.
Edward Pace, Catholic University, Washington; Some Experiments
Upon the Æsthetics of Visual Form, Prof. Lightmer Witmer, Univer-
sity of Pennsylvania ; Investigations of Reaction-times of Various
Classes of Persons, Prof. Lightmer Witmer, University of Pennsyl-
vania ; History and Prospects of Experimental Psychology in America,
President G. Stanley Hall, Clark University ; Experimental Psychol-
ogy at the World’s Fair, Prof. Joseph Jastrow, University of
Wisconsin.
Wednesday, Dee. 28,—Certain Phenomena of Rotation, Dr. Herbert
Nichols, Harvard University ; Note Upon the Controversy Regarding
the Relation of the Intensity of the Stimulus to the Reaction Time,
Prof. W. M. Bryan, University of Indiana ; Minor Studies at the
Psychological Laboratory of Clark University, Dr. E. ©. Sanford,
Clark University ; Preliminary Notes Upon Psychological Tests in the
Schools. of Springfield, Mass., Prof. W. M. Bryan, University of
Indiana.
Anthropological Society of Washington.—Dec. 20, 1892.—
The program of exercises was as follows: Is Simplified Spelling Feas-
ible? (Symposium.) Discussion by Prof. F. A. March, of Lafayette
College, Hon. A. R. Spofford, Hon. Wm. T. Harris, Hon. Edwin
Willits.
January 3, 1893.—The program of exercises was the following: Is
Simplified Spelling Feasible as Proposed by the English and American
Philological Societies? (Close of the Symposium. ) Discussion by
Prof. Lester F. Ward, Prof. Wm. B. Powell, Mr. James C. Pilling,
Prof. Benj. E. Smith, Editor Century Dictionary ; Dr. Charles R. G.
Scott, Editor Worcester Dictionary; Mr. E. T. Peters, Major John W.
Powell and Weston Flint. The discussion was closed by Hon. A. R.
Spofford and Dr. Wm. T. Harris—W Eston FLINT, Secretary.
The BiologicalSociety of W ashington.—Dec.17,1 892.-The prin-
cipal topic of the evening was What Should be the Seope and Object of a
Biological Society ? Introduced by Mr. B. E. Fernow. The following
communications were read: Frost Freaks of the Dittany, Prof. Lester
F. Ward ; Notes on Peach Rosette, Dr. Erwin F. Smith ; Destruction
‘of Lichens on Pear Trees, Mr. M. B- Waite; Notes on Apple and Pear
Fusicladii, Mr. D. G. Papen Se 1 toi Rovio
Indiana Academy of Science.—The eighth annual meeting
was held in the Capitol Building at Indianapolis Dee. 28 and 29.
188 The American Naturalist. [February,
There was a large attendance of members and visitors. President J.
L. Campbell, Wabash College, Crawfordsville, Ind., presided. The
large number of papers entered made imperative the meeting of the
Academy in three sections in the afternoon of the first day. These
were so constituted that papers on mathematics, physics, chemistry
and geology were read in one section; on botany in another; on
zoology and anthropology in a third. On the evening of the first —
day the President deliyered his address on The Independence of Lib-
eral Pursuits. The following officers were elected for the next year:
President, J. C. Arthur, Purdue University, Lafayette, Ind.; vice
president, W. A. Noyes, Rose Polytechnic Institute, Terre Haute, Ind.,
secretary, A. W. Butler, Brookville, Ind.; assistant secretary, Stanley
Coulter, Purdue University, Lafayette, Ind. ; treasurer, ©. A. Waldo
De Pauw University, Greencastle, Ind. ; auditors, Thomas Gray, Rose —
Polytechnic Institute, Terre Haute, Ind.; W. S. Blatchley, High
School, Terre Haute, Ind. L. M. Underwood, De Pauw University,
Greencastle, Ind.; and W. A. Noyes, Rose Polytechnic Institute, Terre
Haute, Ind., were appointed program committee. The Academy —
decided to hold its spring meeting the third week in May in Parks —
County, Ind., closing with an evening meeting Friday at Terre
Haute, Ind.
The following papers were presented: Notes on the Reproduction —
and Development of Grinnellia americana Harv., M. A. Brannon; —
tion of High Temperatures, W. A. Noyes; Tests of the Torsional
Strength of a Steel Shaft, Thos. Gray ; An Extreme Case of Parasit- i
ism, Robert Hessler; Exhibition and Explanation of a Geologi ay
Chart, Elwood P. Cubberly; Local Variations, C. H. Eigenmann;
Botanical Field Work in Western Idaho, D. T. MacDougal; The -
Quaternion Treatment of the Motion of Two or More Bodies U ae i
the Law of Gravitation, A. S. Hathaway; The Electrical Oxidation
of Glycerin, W. E. Stone and H. N. McCoy ; Notes Concerning Testsof
the Purdue Experimental Locomotive, W.F. M.Goss ; The Electrostati 7
Theoryof Cohesionand Vander Waal’s Equation, Reginald A. Fessenden
On Sulphon-pthaleins, Walter Jones’; Quartz Suspensions, Benj. W
Snow; Observations on Glacial and Pre-glacial Erosion at Richm¢
Indiana, Joseph Moore; A Modification of Grandeau’s Method | :
TE linear eg 2 ot ft dae ak et ge Fa eee ~~
j ARE A E eee A TI T o ee EE See A
.
1893.] Proceedings of Scientific Societies. 189
Determination of Humus in Soils, H. A. Huston and F. W. McBride ;
Experiments with and Phenomena of Vacuum Tubes, R. A. Fessen-
den; The Extraction of Xylan from Straw in the Manufacture of
Paper, W. E. Stone and W. H. Test; The Electro-magnetic Inertia of
a Large Magnet, Thos. Gray; The Determination of Chlorine in Nat-
ural Waters, W. A. Noyes; Some New Electrical Apparatus, R. A.
Fessenden ; Thiofurfurol and its Condensation Products, W. E. Stone
and Clinton Dickson ; On the Construction and Use of a Bolometer,
B. W. Snow; On the Determination of Valence, P. S. Baker; An
Application of Mathematics in Botany, Katharine E. Golden; On
the Fertilization and Development of the Embryo in Senecio aureus,
D. M. Mottier; Distribution of the North American Cactacee (by
title), John M. Coulter; Marchantia polymorpha, not a Typical or
Representative Liver-wort, L. M. Underwood; Notes Concerning
Certain Plants of the Southwestern Counties of Indiana, John S.
Wright; Spines and Epidermis of the Cactaceæ (by title), E. B.
Uline ; Preliminary Notes on the Genus Cactus, E. M. Fisher; An
Auxanometer for the Registration of Growth of Stems in Thickness,
Katharine E. Golden; The Apical Growth of the Thallus of Fucus
vesiculosus, D. M. Mottier ; Symbiosis of Orchidaceæ, M. B. Thomas;
Notes on Pediastrum, W. L. Bray; The Genus Corallorhiza, M. B.
Thomas; Notes on Root Tubercles of Indigenous and Exogenous
Legumes in Virgin Soil of the Northwest (by title), H. L. Bolley ;
Notes on Archeology in Mexico, J. T. Scovell ; Notes on the loss of
the Vomerine Teeth with Age in the Males of the Salamander,
Desmognathus fusca (by title), F. C. Test: Modern Geographical Dis-
‘tribution of Insects in Indiana (by title), F. M. Webster; New Species
of Indiana Hymenoptera, reared at LaFayette, Ind. (by title), F. M.
Webster; Description and Elevation of Mount Orizaba, J. T. Scovell:
The Climate and Glaciers of Mounts Orizaba and Popocatapetl, J.
T. Scovell; A Mite, Probably Hypoderas columbe, Parasitic in the
Pigeon, W. W. Norman; The Locustide of Indiana with Description
of New Species, W. S. Blatchley ; Early Stages in, the Development
of Cymatogaster, C. H. Eigenmann ; Some Remarks Regarding the
Embryology of Amphiuma, O. P. Hay ; Some Structural Peculiarities
of Pacific Slope Fishes (by title), A. B. Ulrey ; Peculiar Death of an
Oriole (by title), T. B. Redding ; The Range of the American Cross-
bill (Loxia curvirostra minor) in the Ohio Valley, with Notes on its
Unusual Occurrence in Summer, A. W. Butler; A Note on Loria
curvirostra, W. S. Blatchley ; Notice of a Terrapin to be Restored to
the Fauna of Indiana, O. P. Hay; A Migration of Birds and one of
Insects, T. B. Redding: The South American Catfishes Belonging to
190 The American Naturalist. [February,
Cornell University (by title), E. M. Kindle; Notes on the Genus Lytta,
W. P. Shannon; The Ichthyologic Features of the Black Hills Region,
B. W. Evermann; Explorations in Western Canada, C. H. Eigen-
mann; Ancient Earthworks Near Anderson, Indiana, Francis A.
Walker ; The Work of the U. S. Fish Commission Steamer Albatross
in the North Pacific and Bering Sea in 1892, B. W. Evermann; A
Thermo-regulator for rooms Heated by Steam, J. C. Arthur; Archæ-
ology of Tippecanoe County, O. J. Craig; Some Indian Camping Sites
near Brookville, A. W. Butler; Relation of Kings County Traps to
those of Cumberland County, N. S., V. F. Marsters; The Traps of
Red Head, N. B., V. F. Marsters; On Birds in Western Texas and
Southern New Mexico (by title), A. W. Butler; An Account of Veg-
etable and Mineral Substances that Fell in a Snow Storm in LaPorte
County, Jan. 8-9, ’92 (by title), A. N. Somers; How a Tendril Coils,
D. T. MacDougal; Remarkable Prehistoric Relic, E. Pleas; The
Brun’s Group of Mounds, H. M. Stoops ; Some Points in the Geology of
Mount Orizaba (by title), J.T.Scovell ; Two-ocean Pass (by title), B. W.
Evermann ; The Blattide and Phasmide of Indiana, W. S. Blatchley;
Forestry Exhibit of Indiana at the Columbian Exposition, Stanley
Coulter; The Yolk Nucleus, J. W. Hubbard; Some Causes Acting
` Physiologically Toward the Destruction of Trees in Cities, J. ©.
Arthur ; British Columbia Glaciers, C. H. Eigenmann ; A State Bio-
logical Survey—a Suggestion for Our Spring Meeting, L. M. Under-
wood ; The Mounds of Brookville Township, Franklin County, Ind, S
H. M. Stoops; How the Colleges could aid the Public Schools in
Teaching Biological Subjects, W. W. Norman ; Notes on the Flora of
the Chilhowee and Great Smoky Mountains, Stanley Coulter; The”
Need of a Large Library of Reference in Cryptogamic Botany in
Indiana—W hat the Colleges are Doing to Supply the Deficiency, L. ae
M. Underwood ; Exhibition of a Series of Grouse and Ptarmigan from
Alaska, B. W. Evermann ; Botanical Assemblies in the United States
Announced for the Year 1893, J. C. Arthur; Development of Ovule
in Aster and Solidago (by title), G. W. Martin; Remarks on Arche
ological Map Making (by title), A. W. Butler ; The Lilly Herbarium
and its Work, John S. Wright; Additional Facts Regarding Forest —
Distribution in Indiana, Stanley Coulter; Rotary Blowers, John T.
Wilken ; Some Effects of Mutilation on the Forms of Leaf and Sex —
of Morus alba and Morus nigra (by title), A. N. Somers; The Craw
ford Mound (by title), H. M. Stoops.
1893.] Scientific News. 191
SCIENTIFIC NEWS.
What Is an Acquired Character ?—I suppose that I am the
person to whom Prof. Nutting refers (December, p. 1009) as “ the
member ” who, at the Rochester meeting, “ had the temerity to confess
that he was not sure that he knew the exact nature of an ‘acquired
character. ” I have been strongly impressed with the feeling that
argument against the position of Weismann and his followers is useless,
from the method of these writers in explaining away whatever proofs of
the hereditability of acquired characters one may bring forward. The
definitions of an acquired character are themselves sufficiently vague
and various, but the treatment of individual instances by the Neo-
Darwinians seems to preclude the possibility of argument. Prof.
Ward has well said that Weismann’s “ treatment of this point (hered-
itability of acquired characters) often borders on the dogmatic.” The
practice seems to be to use the definition as a stalking-horse to shield
the philosopher from the onslaughts of his opponents. If he is driven
into a corner it is easy enough to say that the character in question
is not acquired, but that there existed a predisposition to it; and while
he cannot afford any proof of the assertion and may not be able to
bring furward any reason for it beyond its convenience as a method of
escape, his opponent cannot dislodge him from his position. It seems
to me, as I said at Rochester, that much of this argument is but a war
of words; and Prof. Nutting has precisely indicated my own difficul-
ties concerning it.
I have been growing plants for a single generation in soils of
different chemical composition, the seeds having been taken from the
same fruit. These plants show characteristic differences, and when
the seeds from the different lots are sown in one soil the resulting
plants show that some of the novel characters become hereditary for a
generation or more. If I were to bring these experiments forward as
proof of the transmission of acquired characters, my opponents would
simply say, “Oh, well, these are not acquired characters; there was a
predisposition to them.”
I may remark here that an inexhaustible field for study of contem-
- poraneous evolution is afforded by common cultivated “plants ; and if
we are to consider acquired characters in the Darwinian sense, I fancy |
that no one could long be a horticulturist without becoming a con-
firmed believer in their transmission.—L. H. Barry, Cornell Univer-
sity. j
os pag ` ‘The American Naturalist. [February,
The Geological Survey of Alabama.—The Legislature four
years ago reduced the tax rate, unwisely, as now appears, for it has
caused the money in the treasury to run now about $200,000 short of
what is needed to carry on the affairs of the State. When the present
Legislature met and was confronted with this deficiency in the treas-
ury, the first thought was to reduce the expenses of the State within
the present income by retrenching in every direction. Accordingly a
number of bills were introduced aiming to cut off everything that we
could possibly do without. Among these bills was one prohibiting the
payment of any money to the survey for the next two years, saving
$15,000 on this item. Many of the members both of the House and
of the Senate were disposed to support this wholesale retrenchment if
by it the raising of the tax rate could be avoided; for the raising of
the rate would be an unpopular measure as a matter of course, and
would be made the most of by the enemies of the party in power.
Upon closer examination it has become perfectly evident to every one -
that the tax rate must be raised, for all the retrenchment proposed and
in any way possible will fall far short of remedying the trouble. But
some of the members are afraid to vote for raising the rate without, as-
an offset, voting to cut down every expense in the State to its minimum. —
While it is not likely that the extreme measures at first proposed will
pass, there may have to be some compromise that will cut off part of
the sum usually coming to the survey. There is no one fighting the
survey, for the sentiment is very favorable to us, but they may haveto
show that they have economized.— EUGENE A. SMITH.
Recent Deaths.—John Obadiah Westwood, M. A., F. L. S., hon-
orary president of the British Entomological Society, died January Lsa
Prof. Westwood was born in Sheffield in 1805. He was educated at
Litchfield and was appointed in 1861 to the professorship of zoology
founded at Oxford by the munificence of the late Rev. F. W. Hope. —
In 1855 the Royal Society awarded bim one of the royal medals
for his scientific works, and in 1860 he was elected to fill the place
of the illustrious Humboldt as corresponding member of the Ento-
mological Society at Paris. He wrote “Introduction to the Modern
lassification of Insects,” “ Entomologists’ Text Book,” published in ef
.1838 ; “British Butterflies and Their Transformations” in 1841, and
a number of other works of a similar nature. The number of his”
memoirs on special groups of insects is large, and he introduced many —
important forms to the scientific record. 2
Prof. J.G. Joessel, of the University of Strasburg, is dead. He was
3 born April 27, 1839, and was ordinary professor of anatomy.
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CONTENTS:
PAGE. P
Mineralogy and Pelrography-—The- Rocks of.
| Lawton Williams. 193 | the Thathorn- The New Je nate bog aay Seike Mies
NOTHERIUM. BEDS. “(Clstrated.) eres from Kenti fe vs
atcher, 204 d Penn—The Nepheline and Tepe o!
DUCED- SEXUA ALLY WITHOUT Brazi — Petrographical Ne ews — Crys aa a
R IONS OF THE MOTHER. {(Ilus- opsides—Herderite Ange n Hebron, Maine
. Boveri. 222 Notes—New i
CEASSIFICATION OF THE LONGIPENNES.
Shufeldt: 233
The Se sg Survey of Mlinois—
n Ortho SA _Tutersiional Botanical Ga
ical Society. -238 Erna Not :
Zoology. pE a s Faunal Areasof North America—
Memoirs of the ‘National en Madagascar Fauna—The Nephridia of A
o ise — Brooks and s—The Position pr the Marsipobranchs—]
ogy Macrura— SETAA of the Human sce T oe — New
Biology Correlation rien of the | —Repti
Ses as oe e
soe eas on the
si , 243 | Composition.
Entomology. —The Pear-Three Psylla.
and Travels — Africa — Ametica—
‘Sia—Europe—The oe caer
gress o on taat he ists.
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Vor. XXVII. Match, Igy: 315
THE QUANTITY OF HUMAN LIFE.
By J. Lawron WILLIAMS.
Compared with periods of terrestrial history the duration of
a human life even at the very longest is exceedingly brief. In
our own time only a very few people ever live to see their
hundredth birthday, and the average age is now calculated on
the basis of elaborate statistics to fall between thirty-three
and thirty-four years.
It seems to bea prevalent opinion among unrefiecting peo-
ple, that the quantity of an individual life is expressed by the
number of days and years recorded on his or her tombstone.
That such an opinion is erroneous will be rendered evident by
a little consideration. First of all, in estimating the quantity
of human life, we must possess some well defined conception
of what we mean by the term life. This point settled, we can
proceed with some degree of certainty in our calculation. Who-
ever has devoted a little thought to the subject, cannot fail
to be deeply impressed with the multitude and diversity of
the factors which enter into the experiences of a life-time.
There are the ever changing phenomena of the external world
awakening and controlling the states of consciousness. There
are the equally changeful individual activities perpetually
giving rise to new internal changes. In other words, there is
the physiological life utilizing the forces of nature, and mak-
ing “ai subservient to the wants of the individual economy ;
4
194 The American Naturalist. [Mareh,
and, by its side, generating and directing its activities is its
ever present companion the psychological life. Here then, we
have a duplicate existence, extending over a finite period of
years, incessantly changing from the time of its beginning,
throughout the periods of growth, maturity and decay, and
ultimately resolved, so far as human observation goes, into other
existences of a lower order.
Life, then is an existence. It is an existence characterized
by incessant changes, and those changes are the manifestations
of force. Hence, disregarding all ontological questions, the
quantity of life may be measured like any other force, by the
product of its intensity into the time of its duration. Let us
‘now consider the factors which enter into and control the
intensities of human life. Subjectively considered, life is ob-
viously made up of a series of conscious experiences ; objectively
considered, it is made of a succession of molecular and molar
motions. To avoid too great complications we will consider it
only subjectively, since the objective life has much in common
with the motions of dead matter.
Hobbes has well said “it is almost all one for a man to be
sensible of one and the same thing, and not to be sensible at
all of anything.” For instance the ticking of a clock contin-
ued unintermittently through days, months, and years, forms —
a rhythm in the conscious life which is equivalent to silence: 2
Not until it ceases are we aroused to a consciousness of its
former existence. The same is true of all familiar and oft ¢
repeated experiences. No matter how intense and vivid may —
have been the first appeal of an experience, sooner OF later its
reiterated occurrence establishes itself in the rounded and
frictionless ruts of an indifferent consciousness. It might at
first be inferred from this law, that for the conscious life to
attain to its maximum intensity, it should experience the
rarest stimuli; and those stimuli should be as intense as pos-
sible. There is a limit however, beyond which nove ;
to be impressive, and the over stimulated and flagging COD-
sciousness relapses into apathy and indifference. But as &
general rule we may be guided by Hobbes’ statement, and
from its eonverse deduce the proposition, that the intensity of
= Se
IE ee
1893.] The Quantity of Human Life. 195
conscious life varies as the number and rarity of its experi-
ences. But the number and rarity of its experiences depend
upon a great variety of circumstances.
They depend upon the diversity of external conditions.
They also depend upon the extent and acuteness of surfaces
of contact, and the rapidity with which those surfaces are
transported into new environments. Finally they will depend
upon the stock of vitality of the individual. All these factors
must be considered in estimating the quantity of the conscious
life. The diversities of external surroundings are very great.
Even in primitive communities, and amid the habitual isola-
tions incident to a nomadic life, the differences in the attitudes
and groupings of the objects of inanimate nature, and of the
lower orders of animate nature, are ever stimulating the con-
scious life to new and intense experiences. Surprises, alarms,
friendship, apathies, pleasures and pain are ever appealing in
new and varied relations to the would-be slumbering sensibil-
ities. But when we pass from the primitive condition of
solitude to the later condition of society where the wide play
of forces concerned in human interests is brought to bear in
a multiplicity of combinations upon the individual life, the
number of conscious experiences increases with the members
of the community not in an arithmetical, but in a geometri-
cal ratio. This holds up toa certain point when the maxi-
mum of conscious experiences during a given time is reached.
The limit is a subjective and not an objective one. It is
determined by the capacity for reception of the individual.
Amid the intricacies of modern civilized life that limit 1s
speedily reached. Even the most capacious and versatile
minds soon weary of the endless solicitations of the senses and
the trains of ratiocination to which they give rise. The keen
activity of such minds is rapidly dulled by the continuous
impact of stimuli, and soon falls back upon the restful reitera-
tion of former experiences. But even the repose of such
reminiscences is disturbed by the jars and turmoils of inevita-
ble environments, and the drowsy sensibilities are spurred on
to renewed activities day after day so long as life continues.
Thus it happens that the average life of the social individual
196 The American Naturalist. [March, e
:
is much more intense than that of the inhabitant of solitudes.,
The reason is, that the sociological stimuli affect us more than
stimuli of equal or even greater intensity proceeding from ~
inanimate nature, or the lower orders of animate nature. So-
ciety also engenders wide differences in the experience of its
various members on account of financial, political and other —
differences. The humdrum life of the common laborer is
much less varied, and, therefore much less intense than the
life of the gentleman of affluence and leisure who diversifies a
his experiences by travel, letters, society and recreative F
sports. l
The lavish distribution of cheap literature within the last —
quarter of a century has immensely augmented the differences
in the quantities of living among people of the same social —
standing. It is not uncommon to find two men engaged in —
the same occupation one of whom cannot read or write, while —
the other is conversant with the widest questions of political —
and social progress. Literature thus tends to obliterate the —
quantitative differences of life arising from certain social —
accidents of birth, and places all men, so far as the intellectual —
consciousness is concerned, on pretty much the same level as
quantitative experiences. Much more might be said as to the ©
diversities of the external conditions, but this must suffice.
We now pass on to consider the surfaces of contact. Und
this designation are included all the external integuments
the body as the skin, eyes, hair, and so on. For the sake
of mathematical simplicity we will assume at present that-
equal surfaces of contact possess equal degrees of sensibility 1m
two individuals during equal periods of time. Then if onè —
person present twice as much surface of contact as another, a
that person will, other things equal, receive twice as mam
peripheral stimulations as the other. (We here assume |
an equal number of stimulations come from every direction
against the unit of the surface.) Now the surfaces of contae
vary as the square of some lineal unit. Suppose the su l
of contact be 4 and 8 respectively ; then the linear units are 2
and 2.8+. But the masses and weights of the bodies w99
surfaces are 4 and 8 vary as the cube of the linear units, oF @
fo $e We ae 5
CR te re Oe ee eae TM US
Dees d
“Haas A asa 9G a oa A NEL aa E
1893.] The Quantity of Human Life. 197
2° = 8 and (2-84 F = 22-44. Now in order that the individ-
ual with double the surface of contact may encounter twice as
many stimuli as the other, when both are moving through
their environment, it must move the same distance as the
other. In moving such a distance however, it will perform
not twice but nearly three times the amount of work that the
other performs since it will transport nearly three times the
weight. Thus, if the expenditure of muscular energy detracts
from the stock of vitality, and hence dulls the keenness of the
sensibilities, even though the surfaces of contact of the two
individuals possess initially the same degree of sensibility
by reason of that impairment, the larger animal would expe-
rience less than twice as much consciousness as the smaller.
Viewed from another standpoint, the smaller animal could
execute more than twice the velocity of the larger with equal
expenditure of strength, ,(disregarding the resistance of the
medium), and hence it would encounter more than an equiva-
lent number of experiences.
It was here assumed that equal surfaces of contact conveyed
equal quantities of sensibility in the two individuals, but it
is well known to physiological psychologists that there are
immense differences in the relative acuteness of the different
sense organs in the same individual, and in the same sense
organs in different individuals. The optical, auditory and
tactile powers are quite variable. Probably the experiential
products of all the other senses combined do not equal those
which come through the medium of vision. It will thus be
readily inferred that the extent of surfaces of contact is less
important in relation to the experiences of the conscious life
than their quality; and the movements of the individual
tending to multiply experiences are less important than the
movements of the environment. For example, an observer
standing on a street corner in a great city, or riding in a rail-
way coach at forty miles an hour, will gather more experiences
than he would were he to run atthe top of his speed in a
deserted field. Doubtless he might experience a high degree
of exhilaration at first, yet experience of effort would prevail.
Such experiences like all other somatic feelings have a place
198 The American Naturalist. [Mareh,
in consciousness, but they certainly do not have ‘that vivid
and definite character which distinguish peripheral sensations.
Moreover they must detract from the general vitality, and
lower the tone of the higher orders of experience.
There can be no doubt that the stock of vitality, and the
natural endowments of body and mind immensely transcend
all the other differences hitherto considered. Not only do
ER E cro 3. ei gE eat ee
they give rise to differences in the intensity, but also in the — :
duration of life. The vigorous, alert, and impressionable nature
has a livelier consciousness of all that it experiences, responds
more keenly to all its enjoyments, and resists more stubbornly
the wracking and dissolution of conflict and disease. Hence
such an individual lives longer and lives more. There are
still other differences of a psychological character which give
rise to still more important variations in the quantity of the
conscious life. Owing to the absence of any standards in the
higher fields of psychometric research, it is hard to discrimin-
ate between quantitative and qualitative differences. Yet
viewed from a rational standpoint it cannot be doubted that
different kinds of consciousness are quantitatively different,
though there is no perceptible disparity in their relative
intensities. That the untutored savage feels as intense grati-
fication in the pursuit of his game, as the astronomer in the
pursuit of undiscovered asteroids, cannot reasonably be doubted.
But how totally different the quality of the feeling, and the
psychological grounds upon which it is based! Where the
thoughts and emotions are different in kind, this disparity 18
even greater. There are certain thoughts and emotions access
ible to the few which are totally beyond and above the range
of the average mind. Such thoughts as those of Laplace and
Kant, and such emotions as those of Schiller, are of this order.
While it may not be easy to defend the position, yet We feel
forced to believe that there are quantitative differences between
such thoughts and emotions, and the stolid apathy, and or ,
materialistic conceptions of the mediocre mind.
Up to this point we have considered the conditions W
modify the quantity of the conscious life in a generat
But the quantity of the life of every human being is subj
hich
] way: 7
ect to Ta a
aa
1893.) The Quantity of Human Life. 199
innumerable vicissitudes both accidental and periodic. While
there is an endless influx of disturbing forces which augment
or deplete the quantity of life at all times, yet there are cer-
tain periodic causes which promote recurrence of cycles of
consciousness. Such are the alterations of day and night, the
recurrence of the seasons, and the widely extended observance
of days and customs. How much the quantity of life is tem-
porarily augmented by the convivialities of Christmas and
other commemoration days! Then there is a diurnal rise and
fall in the conscious life. The morning with its vivid and
buoyant sensations; the rising of the energies and their tem-
porary lull at midday, and their slow descent with the
descending of the sun. Then the twilight comes on with its
calm reflections and feelings, succeeded by the serenity of
sleep. But even here the pulse of consciousness rises and falls,
summoned into being by the vague apparitions and muffled
ounds of dreams. At length these increase in vividness
and frequency, the pulse of consciousness grows more intense
until we are again ushered into the hazy realities of another
morning. Then there are the periodicities arising from the
changing seasons.
To one who dwells during a period of years in the same
place, the events which are repeated from year to year are
much in excess of those which are variable. Hence the
annual rhythms of consciousness are quite well marked. The
conscious life throughout the time of its existence is subject to
quite well defined periods of change. There are several strik-
ing analogies between the consciousness of infancy and that
of extreme old age. Both are oblivious of details, apprehend-
ing only to the salient outlines and strong qualities of things.
Both repeat and re-repeat simple things, the babe its lullaby,
the old man his favorite story. The consciousness of old age
is reminiscent because the powers are feeble, and reminiscences
are paths of least resistance. Quite otherwise is the consciousness
of youth and the prime of life. It is vigilant and aggressive,
and ever seeking new combinations of experience. Not only
is the quantity of the conscious life extremely varied in its
different periods of development, but also in its quality. In
200 The American Naturalist. [March,
early years and even far into the period of maturity the phy-
siological activities prevail. Childhood and early youth are
characterized by the aimlessmovements of spontaneity and
the prodigal expenditure of physical force ; later on the psycho-
logical life rises in importance and attains its maximum
when the physiological life is on the wane. This continues
until the decay of old age sets in.
A grander series of changes remains yet to be considered.
This is suggested by the query, how does the quantity of
human life existing on the globe at the present time compare
with the quantity of the antecedent times? Whoever reviews
the history of the race during the nineteenth century as related —
to its history in previous times will observe a tremendous —
acceleration in the rate of living, and a vast augmentation in
the intensity of human life. Notable among the causes which
have secondarily conspired to this end, may be mentioned the —
centralization of populations. Within a century, according to —
the London Journal of Statistics the relative populations of the
country and city of England have changed sides. Up to
about 1840 the rural population exceeded the urban. About —
that year they were equal, and ever since, the cities have had —
a constantly increasing majority. The same is true in general —
of other civilized countries. Facts like these tend to show —
that the society of modern life is becoming more consolidated,
and hence the aggregate of human experiences is becoming
increasingly intense from year to year. Among the primary —
causes which have brought about this consolidation, are the i
improved facilities of travel, commerce, and communication
between mind and mind. Not only have they done this,
they are the indispensible conditions of an intricate co-opera-
tive existence. But these are only a few of the many
which enter the problem of this augmentation of life. 48
differentiation of pursuits and the widely diversified pr?
1893. ] The Quantity of Human Life. 201
lead a more varied life than primitive men, probably more
varied than our contemporary the Fuegian, whose life is a
monotonous ordeal of physical suffering, or the Esquimau
whose days are passed in the unbroken solitudes of Arctic
America. In view of these unprecedented changes, and the
steady increase of population, it is but a step to the conclusion
that the quantity of human life on the globe to-day is greater
than ever before. In all probability the aggregate quantity
of conscious life which has been experienced during the nine-
teenth century is |far greater than the life experienced in any
antecedent period of equal duration. Never was there a
recorded time when the drain upon natural resources was so
great as it is to-day. These resources are all turned either
directly or indirectly toward the furtherance of human experi-
ence. Notonly do these agencies of civilization add to the
multiplicity of human experiences, but they also add to their
pleasurableness. Thus undaunted by the clamors of pessimism
we may firmly believe that the sum of human happiness is
greater to-day than ever before.
The marked feature with which we are impressed in review-
ing the evolution of human life, is its growing complexity.
But by the very terms of a previous proposition, this signifies
a growth in the quantity of conscious experience. All primi-
tive life was simple and plain. Monotony was stamped upon
its music, language, gesture, and the rounds of domestic, social
and political life. The frequent repetitions of barbarian
speech, and the sing-song tones of their music are in pitiable
contrast to the sweeping climax and anti-climax of the civil-
ized orator or opera singer.
The principle is of general application. All orders of expe-
rience are more varied in the civilized than in the savage
state. Hence the quantity of life for each individual is greater,
and the totality of human experiences immensely greater with
the diffusion of civilization. We have now considered some of
the factors which determine the quantity of human life. The
final inquiry remains, how are we to live most? and how is the
race to live most? Judging from the foregoing conclusions
we would say in the case of the individual, that this end is
202 The American Naturalist. [Mareh, .
insured by multiplying the number, intensity, and variety of
his experiences. Spinoza had fora motto, “to live is to think.’
Surely the quantity and quality of our thinking has much to _
do with the quantity and quality of our living, but from con-
siderations already discussed, this utterance seems to be want-
ing in comprehensiveness. If this statement were true without
qualifications, then he who thinks most lives most, but we
have already seen that other experiences than those of thought
have perhaps an equal degree of intensity, and involve an
equal amount ofliving. Any conclusion which can be reached —
on this question is necessarily colored by personal prejudices,
and cannot be set up as a general standard. To me, at least, `
he seems to live most who enjoys the greatest possible range
of conscious experiences of the highest order, during the long- —
est period of time. And the quantity of life seems to be truly
greatest which embodies the fullest and highest expression of |
the attributes of the intellect, the emotions and the will. :
Not where the experiences are confined to one of these
alone, but where all three have equal sovereignty in the
dominion of the mind. He seems to me. to live less whose
life embodies one or all of these in a less degree, or who has
sullied their original purity with baser purposes and ends.
Perhaps this may seem too ethical for a scientific statement
but until that same ethical spirit shall more fully actuate the
promoters of science, its highest beauty and usefulness cannot —
be completely realized. Even the best of us lead much more —
diminutive lives than our circumstances require.
but remember that it is as easy to think great thoughts as w
think little thoughts, and feel great emotions instead of base
and belittling ones, we might all enter into a largeness í
life which far exceeds its present dimensions. But we must
pass on to consider the question how can the race live most?
One thing is certain, while all are permitted to think and fi
and will, all cannot be brain workers. There must be doers
as well. Society is so organized that no small part of
members must perform physical labor. This being the case,
to realize the maximum of conscious life, there must be 1
greatest possible number of mental laborers, and the least po
1893,] The Quantity of Human Life. 203
sible number of physical laborers ; and in each case the maxi-
mum of consciousness must be sought. Speculative as these
views may appear yet they are substantiated by historical fact.
To-day when the quantity of human life is greatest, the phy-
sical laborers effect most with the least effort, and the mental
laborers are constantly diminishing that effort. So that were
a given status of living to continue, the physical laborers
would decrease, and the mental laborers would increase until
finally equiblibrium should be attained.
Such equilibrium however, is indefinitely postponed by the
constantly ascending standard of living. But as the standard
of living ascends, all live more, and, since the physical laborers
diminish and the mental laborers increase, the quantity of life
is still further augmented. This would happen were the pop-
ulation to remain constant. But as the standard of life rises,
the possibilities of existence increase, and the death rate
diminishes. Consequently the population increases as the
quantity of the individual life increases, and hence the total
quantity of human life is augmented by both of these recip-
rocating factors. How great the quantity of human life may
some day be is a question which the developments of the
future alone can decide.
If the present rate of its growth shall continue unabated,
sooner or later the time will come when it can increase no
more. All the resources of the earth will be utilized and
strained to the utmost to augment that life. But they cannot.
Gradually the potential energies of nature which support life
will be dissipated as energy of motion. And as those energies
dwindle and disappear, the quantity of human life will fall
away contemporaneously. The time must come when that
quantity will be zero.
204 The American Naturalist. (March,
THE TITANOTHERIUM BEDS.
By J. B. HATCHER.
In 1857 Meek and Hayden gave the name Titanotherium bed
to the lowest member of the fresh water, Miocene, lake depos-
its of the White river, Bad Lands of Dakota and Nebraska.’
They named these beds from the genus Titanotherium, estab-
lished by Leidy in 1852? This genus embraces the largest,
most abundant, and most characteristic vertebrate fossils found
in these beds. In 1870 Professor Marsh discovered, in north-
eastern Colorado, an exposure of these beds, farther south than
they had then been reported? He then referred them to “The |
true Titanotherium beds.” In 1877 Professor Marsh proposed
the name Brontotherium beds for this same series of strata, from —
his proposed genus Brontotherium.' Since there can be no doubt
that the Titanotheriwm bed M. and H., and the Brontotherium beds
Marsh, are identical, and since the former has a priority of 3
at least twenty years, it should be retained.” In this paper —
the plural will be used, as expressing more nearly the true —
nature of the deposits.
GEOGRAPHICAL DISTRIBUTION OF THE TITANOTHERIUM BEDS. ©
The Titanotherium beds are known to have a surface expo
sure in various portions of the western interior plains region.
That exposure from which they were first named and descri n
and from which Hayden first made and published a section of
them in 1863, is of far greater extent than any other. +”
Titanotherium beds of this exposure are known to occupy è
1Proc. Phila. Acad. Nat. Sci., 1857, p. 120.
2Ancient Fauna of Nebr., Leidy, 1852, p. 72.
*Am. Jour. Sci., Sept., 1870, p. 292.
1Am. Jour. Sci., 3d series, Vol. xiv, p. 354.
‘Titanotherium, Leidy, 1852, is antedated by Menodus Pomel, 1849 5
Menodus is essentially preoccupied by Menodon von Meyer, 1838.
Proc. Am. Phil. Soc., 1863, Vol. xii, p. 31.
3
1893.] The Titanotherium Beds. 205
considerable portion of that country embraced between the
Cheyenne, Missouri, and White rivers in S. Dakota, with a
western continuation, extending along the eastern and south-
ern slopes of the Black Hills, and westward to near the town
of Douglas in Wyoming. While this exposure is seen to have
a considerable extent from east to west, yet from north to south
it is of very limited extent. This is due to avery slight dip
of these beds to the southeast, where they soon pass under the
overlying Oreodon and Loup Fork beds. That range of bluffs
in southwestern S. Dakota, northwestern Nebraska and central
Wyoming, known as Pine Ridge, may be considered as the
southern limit of this exposure of the Titanotherium beds. The
beds of this region occupy a considerable portion of south-
western S. Dakota, extreme northwestern Nebraska, and a very
narrow strip in central and eastern Wyoming. Besides this
exposure of the beds, there are other isolated exposures at
Long Pine, Nebraska; in northeastern Colorado; in Wyoming,
along the Rattlesnake Range, north of the Sweetwater; and
Professor Cope has described remains of Titanotheriide from
Canada.’
Although the surface exposures of the Titanotherium beds
are of comparatively small extent, yet from the distribution
of these.exposures the beds are known to extend over a con-
siderable portion of S. Dakota, Nebraska, Colorado, Wyoming,
and perhaps portions of Montana and N. Dakota.
DESCRIPTION OF THE DEPOSITS.
The Titanotherium beds may be briefly characterized as a
series of usually light colored, sometimes variegated clays,
alternating with less extensive deposits of sandstones and con-
glomerates, situated at the base of the Miocene and containing
among other fossils the remains of Titanotherium and allied
forms.
The typical locality from which Meek and Hayden first
named the Titanotherium beds and from which the latter first
made and published a section of them, is also that locality in
7Contributions to Can. Pal., Vol. iii, p. 8-
206 The American Naturalist. [March,
the Bad Lands of S. Dakota, which has been more thoroughly
explored for vertebrate fossils than any other locality in these
beds. It is located in that portion of Washington county
embraced between the White and Cheyenne rivers, at that
point where they approach nearest to each other. This par-
ticular region has not only proved especially rich in vertebrate
fossils, but the Titanotherium beds and the entire Miocene
series, up to the Loup Fork, has attained a development here
unsurpassed, if not unequaled, elsewhere. In no other region
has erosion done more to aid the investigator than here;
whether his aim be to study the country from a purely geo-
logical standpoint, to collect material for the paleontological
laboratory, or to study the laws and observe the rate and
effects of erosion, under the many and varied circumstances
which this region presents. Since the materials composing
the Titanotherium beds are very similar throughout the entire
distribution of the beds, a description of the deposits at the
typical locality may be considered as representing fairly well
the character of the deposits at any other locality.
The section published by Hayden in 1863 agrees in all
important points with those published since by other authori-
ties. As stated by Hayden, the beds vary in thickness at
different localities. At no locality has he given a thickness of
more than 100 feet. The writer has found, by actual meas- —
urement, that they attain a thickness in some places at least,
of 180 feet. As stated before, they are composed of clays,
sandstones, and conglomerates. The clays greatly predomi- —
nate, consist of very fine particles, and are quite compact. | Ine
places they are composed almost entirely of pure kaolin, but
they often contain a considerable portion of sand. Near the
bottom of the beds the color is often red or variegated, due to
the presence in them of small quantities of red oxide of iron; —
but the prevailing color is a very characteristic and delicate
greenish white. The clays usually contain little or no cement-
ing substance, and are held together almost entirely by adhe-
sion. Occasionally, however, there are quite persistent layat
of clay nodules, or concretions in which the clays are firmly
cemented by carbonate of lime. Owing to the extreme min
1893.] The Titanotherium Beds. 207
uteness of the particles forming the clays and the absence of
sufficient cementing material in them, in most places they
readily yield to the action of water and are quite rapidly
eroded. The clays of the Titanotherium beds were probably
derived from two sources, viz., from the Cretaceous clays and
shales, and from the kaolinization of granitic feldspars.
The sandstones are never entirely continuous, and never
more than a few feet thick. They present every degree of
compactness, from loose beds of sand to the most solid sand-
stones. They are composed of quartz, feldspar and mica, and
are evidently of granitic origin. When solidified the cement-
ing substance is carbonate of lime.
The conglomerates, like the sandstones, are not constant, are
of very limited vertical extent, never more than a few feet
thick. They are usually quite hard, being firmly held
together by carbonate of lime. A section of the beds taken
at any point and showing the relative position and thickness
of the sandstones, clays, and conglomerates is of little value,
since these vary much at different and quite adjacent locali-
ties. ‘ ;
The varying hardness of the different strata of the Titano-
therium and overlying later Miocene beds, by offering unlike
degrees of resistance to erosion, have succeeded in producing
in this particular region an exhibition of characteristic Bad
Lands scenery, unsurpassed elsewhere in the Miocene. The
surface of the country in this region is scarred by numerous
deeply eroded canyons. The intervening ridges are rugged
and barren, often terminating for miles in sharp, serrated
crests. The walls of these canyons present a series of terraces
or projecting ledges from top to bottom, due to different
degrees of hardness in the different strata. Sometimes these
ledges are but a few feet wide, or just wide enough to allow a
person on foot to pass along on them. At other times when
the harder layers of which they are composed are of greater
thickness, they may be several hundred feet wide, and their
surface strewn with the fossil bones of Titanotherium, Elo-
therium, Hyracodon, Hyzenodon, Oreodon, turtles, ete., washed
208 The American Naturalist. [March,
out of the overlying beds, by the slow process of denudation,
presents a remarkable, not to say, sepulchral appearance.
Toward the interior of the Bad Lands, midway between the
White and Cheyenne rivers, side branches of the main can-
yons have cut their way entirely through the dividing ridges
and have produced particularly picturesque effects. Charac-
teristic among these are the Devil’s Tower, at the south end of
Sheep Mountain, in the eastern portion of the Bad Lands;
Chimney Rock, near their center, and the Tabled Rocks along
_ their western border.
The location of such especially characteristic Bad Land
scenery, in this particular region, is doubtless due to the near
approach of the White and Cheyenne rivers to each other at
this place. Since the Miocene deposits of this region are
essentially horizontal, and form the summit of the divide
between these two streams; this divide would be of essentially
the same altitude, in reference to the beds of these streams,
whether the latter were near together or far removed from each
other. Therefore these streams upon approaching nearer to
each other, as they do in this region, where they are only
about 20 miles apart, would greatly reduce the length of their
respective, intervening tributaries, without decreasing the
height of the divide which separates them. This would nec-
essarily increase the average rate of fall of the tributaries, and
correspondingly the rapidity of the flow of their waters,
therefore the erosive power of the latter. ,
In various portions of the Titanotherium beds there are
numerous vertical veins of chaleedony running through the :
beds in every direction. These veins vary in thickness from ri
that of a sheet of paper to about two inches. On first thought
the writer was inclined to attribute their origin to mud cracks,
any particular region where they now occur having been wr :
short periods, during seasons of low water, above the water
level and subjected to the action of the atmosphere and the
heat of the sun became baked and cracked ; just as we now 80
1893.] The Titanotherium Beds. 209
with the same materials as the overlying beds; for when the
waters again covered this region, the mud cracks would be
immediately filled by sedimentation and with the same mate-
rials that now compose the overlying beds. It has sinee
occurred to the writer, that these cracks were not made while
the particular strata in which they now appear occupied the
immediate bottom of the lake, but after the overlying beds
were deposited. The extreme fineness of the particles form-
ing the clays of the Titanotherium beds in those places where
these veins occur is evidence that the clays were deposited by
a slow process of sedimentation in still waters. The bottom
of a lake where such materials were being laid down would
consist for several feet of a very thin mud or ooze. This would
gradually become firmer toward the bottom as deposition con-
tinued, but would still mechanically retain a considerable per
cent. of water. Later, when the entire overlying series of
strata were deposited and the region brought permanently
above the water level this imprisoned water would gradually
disappear, by filtration or otherwise, aided perhaps by the
pressure of the superincumbent beds. This loss of moisture
in the clays would diminish their volume and bring about a
readjustment of the particles composing them. The decrease
in volume would be taken up in two ways: First, as in the
case of mud cracks, the particles would tend to collect about
certain centers in the beds, and these centers of adhesion
would increase laterally by the attraction of adjacent parti-
eles until cracks of varying thickness would form between
the peripherys of adjacent centers of adhesion. The pressure
of the overlying beds would determine the vertical direc-
tion of these cracks, and would also afford the means for
the second way, by which the decrease in the volume of Ne
clays would be takem up, viZ., by a decrease in the verti
thickness of the beds. These cracks, thus formed far beneath
the surface, were afterward filled by chaleedony —
out of the overlying beds by heated waters perco EA
through them. Occasionally other minerals, as ordinary cal-
cite and its less common, variety known as Iceland spal ars
found in small cavities in these vains.. Localities containing
15
210 The American Naturalist. [Mareh,
these peculiar veins of chalcedony resemble both in appear-
ance and structure a huge septaria, except that the veins in the
former are much smaller in comparison with the area of the <3
intervening spaces than in septarie. A. singular fact in
connection with the position of areas exhibiting this peculiar
structure is also thought to afford in itself evidence in favor
of this view of the origin of the veins. These veins occur
only in certain localities of limited area. Any single locality
is never more than afew square milesin extent. These small
and isolated areas, wherever it has been possible to determine
their relation to the underlying beds, have been found to have —
been deposited in an eroded valley or depression of the latter.
Such depressions would afford a natural basin for the waters, —
held mechanically by the clays, and would prevent a rapid —
drainage during their deposition.
STRATIGRAPHICAL POSITION OF THE TITANOTHERIUM BEDS.
Tur UNDERLYING BEDS.
The Titanotherium beds are underlaid by various older for-
mations, from the Laramie to early Paleozoic or perhaps
Archean. In most places they rest unconformably upon the
eroded surface of some member of tħe Cretaceous, most fre-
quently the Fort Pierre shales. At the mouth of Beaver creek,
a tributary of White river, near the Nebraska and Dakota
State line, the writer in 1886 observed the Titanotherium beds-
resting upon the eroded surface of the chalk of Cretaceous No. |
3, so well developed in central and western Kansas. This is
believed to be a more northwestern extension of these beds
than had then been reported. ;
THE TITANOTHERIUM “BEDS.
The Titanotherium beds may be divided into Lower, Middle,
and Upper beds, each characterized by different forms % —
the Titanotheriide. Accepting a total thickness of 180 feetas
the maximum thickness of these beds, 50 fect of this would
belong to the Lower, 100 feet to the Middle, and 30 feet to the
Upper beds. The different forms of Titanotheriide, especia
1893.] The Titanotherium Beds. 211
characteristic of these three divisions will be pointed out and
figured later in this article.
THE OvERLYING BEDS.
The Titanotherium beds are everywhere overlaid by the
Oreodon beds, except where the latter have been removed by
erosion. So far as noticed there are in the Bad Lands of §.
Dakota no evidence of any unconformity between the Oreodon
and Titanotherium beds, unless the sudden change from one
fauna to another should be regarded as evidence of an uncon-
formity or at least as indicating a considerable break in sedi-
mentation between the two beds. ‘
Southwest of the Dakota Bad Lands in the extreme north-
western portion of Nebraska there seems to be some evidence
of an unconformity between the Titanotherium and Oreodon
beds, or at least that for a considerable period immediately
following the deposition of the Titanotherium beds, this region
became dry land and that the Oreodon beds were subsequently
laid down upon the eroded surface of the Titanotherium beds.
At the extreme head of Cottonwood creek, a small tributary of
White river, from the northwest, in the northwestern part of
Dawes county, there is an isolated butte composed entirely of
the Titanotherium beds and containing many Titanotherium
bones. The different strata composing this butte are horizon-
tal. About one mile south of this isolated butte is a range of
bluffs of the characteristic Oreodon beds, containing numer-
ous bones of Oreodon and other associated animals.
In the first mentioned butte Titanotherium bones are found
at the same altitude at which Oreodon bones occur in the
bluffs to the south. Whether this occurrence of Titanothe-
rium and Oreodon bones at the same altitude in quite adja-
cent beds is due to the Oreodon beds having been deposited
upon the eroded surface of the Titanotherium beds, or to
differences in the level of the bottom of the lake, at the begin-
ning of the Miocene period, could not be determined. The
horizontal position of the strata composing the Titanotherium
butte would seem to favor the first conclusion, since if these
212 The American Naturalist. [March,
strata had been laid down upon a sloping bottom they would,
in their lower members at least, partake of the inclination of
the bottom of the lake.
About six miles northeast of the town of Chadron, in Dawes
county, Nebraska, on a small branch of the west fork of Dry —
creek, the writer, in 1886, while collecting missing portions of ©
the skeleton of Professor Marsh’s type of Titanotherium (Brontops)
robustum, discovered by H. C. Clifford in 1875, observed several
feet above the horizon in which this remarkably complete
skeleton was found, and certainly not far from the top of the
Titanotherium beds, positive evidence of a break in sedimen-
tation at this place at least. This evidence consists in the occut-
rence at this place of a very dark colored layer of vegetable
mould about two feet thick, accumulated no doubt during 4
period of elevation above water level. x
Owing to the fact that the Titanotherium and Oreodon beds
of S. Dakota and Nebraska are approximately horizontal, itis
almost impossible to determine there, just what their strati-
graphic relations are. Farther west, near the head of Lance
and Harney creeks, in Wyoming, the Titanotherium beds, at
least, dip to the southward at a considerable angle. The writer
has been unable to make any but the most superficial obser-
vations in this region, which, if carefully examined, would
doubtless furnish a solution of the actual stratigraphic rela-
tions of these beds. au
Professor W. P. Jenney, of the U. S. Geol. Survey,
reported to the writer an account of the discovery of & sM
Miocene lake in the Black Hills. As a description of th
singular and interesting deposit a portion of Professor Je
letter is quoted. He says: “ About 1879-’80 I was into?
that fossil bones . . . . had been found in a prospe
shaft near Lead City. I visited the locality and obtained
session of three nearly perfect skulls a: These mE
I sent to Professor Marsh at New Haven, accompanied by
letter giving a sketch and full description of the occur
and received a reply stating that they were all bape
bertsonii (this specific name is from memory but E
correct). iM
1893.] The Titanotherium Beds. 213
“ This Miocene deposit is situated about half a mile south-
easterly from Lead City, South Dakota, and nearly five miles
south of the city of Deadwood, at an elevation of about 5,200
feet above the ocean. . . . . Of the extent of this tertiary
deposit very little is known. It does not outcrop at the surface
and appears to occupy a small, deep channel or basin eroded
in the Archean slates which outcrop on all sides within a
radius of 1000 to 3000 feet. . . . . This occurrence is
peculiar not only in forming an outlier to the great Miocene
deposits and in occupying a basin eroded in a range of steep
Archean hills where lakes are now unknown, but also in the
nature of the deposit itself, which so closely resembles, in
appearance at least, the deposits of the same age on the plains.
The clays of the latter were probably derived from Cretaceous
shales, but the sediments filling this lake would naturally be
expected to be very different, the rocks in the vicinity being
Archean slates capped by Potsdam sandstone and shales, and
locally overlaid by masses of eruptive post-Cretaceous por-
phyry
That this ennai lake was, during Miocene times, entirely iso-
lated from the great body of water to the eastward can hardly
be questioned. The similarity just referred to by Professor
Jenney as existing between the deposits of the small lake and
those of the larger one is probably due to the materials com-
posing both having been derived largely from a common
source. The prevalence everywhere throughout the Miocene
of large quantities of sands and conglomerates, composed of
coarse grains or pebbles of quartz, feldspars, and mica, is suffi-
cient evidence that the materials composing the beds of the
Miocene were derived largely from sources other than the
Cretaceous shales, which contain little sand and are always
very fine grained. The coarser sands and conglomerates were
doubtless derived from Jurassic, Paleozoic, and Archean rocks,
and a considerable portion of the Miocene clays probably owe
their origin to the decomposition of the slates of these older
_ formations and the kaolinization of granitic feldspars.
214 The American Naturalist. [March,
DIVISION OF THE TITANOTHERIUM Bens Intro Lower, MIDDLE
AND UPPER BEDS.
During the seasons of 1886, 1887, and 1888, the writer spent `
fifteen months in the White River Miocene beds of S. Dakota —
and Nebraska, collecting material for Professor Marsh’s Mon-
ograph on the Titanotheride (Brontotheriidx). Among the
material collected was 105 nearly complete Titanotherium
skulls, and many portions of skeletons and disarticulated
bones; besides the remains of many other associated animals.
Early in the season of 1886 it became apparent that certain
forms of skulls were characteristic of certain horizons in the
beds. This fact showed the importance of keeping, so far as
possible, an exact record of the horizon from which each skull
or skeleton wastaken. From actual measurement the vertical —
range of the Titanotheriide was found to be about 180 feet.
For convenience in keeping a record of horizons the beds were
divided into three divisions of 60 feet each, and each of these
three divisions was subdivided into three divisions of 20 feet
each. The different skulls and skeletons, when dug out, were —
each given a separate letter or number, and this letter or num —
ber was placed in that subdivision of the beds from which the
skull or skeleton was taken. :
At present about 60 of these skulls and several more or less
complete skeletons have been freed from their matrix. When
studied in connection with the horizons from which they were
taken, these remains show that a regular and systematic devel-
opment took place in these animals from the base to the top
of the beds. The most noticeable change which took place 10
the Titanotheriidæ was a gradual and decided increase in their
size from the lowest to the uppermost beds, as is shown by the —
increase in the size of the skulls, fore and hind limbs,
other portions of the skeleton. Individuals found near
bottom of the beds are little, if any, larger than the liv
rhinoceros. From this they gradually increase in size 88 ew
go up until at the top we find a type described by Professor
Marsh as Titanops, rivaling the modern elephant in size.
This increase in size from the base to the summit of the beds
1893.] The Titanotherium Beds, 215
was attended by a very marked development in certain por-
tions of the skeleton, noticeable among which are the follow-
ing: A variation in shape and an increase in the size and
length of the horncores as compared with the size of the skulls,
attended, near the summit of the beds at least, by a decided
shortening of the nasals.
There were also changes taking place in the dentition of
these animals, especially in the number of incisors and in the
structure of the last, upper, true molar. The number of inci-
sors, though probably never constant, even in the same spe-
cies, shows a tendency to decrease in skulls found near the
summit of the beds. At the base of the beds the number of
incisors is from one to three on aside, while at the top there
‘are never more than two on a side, often only one, sometimes
none. In skulls from the very lowest beds the incisors have
already become so rudimentary as to be no longer functional.
As would be expected, the number of incisors decreased after
they became of no functional value. In the matter of incisors
the Titanotheriidse were, at the time of their extermination, in
a fair way to accomplish just what the somewhat related but
more persistent Rhinocerotide have very nearly succeeded in
doing, viz., the elimination of the incisor dentition. The
number of incisors in the Titanotheriide varies with age in
the same individual. These teeth were but loosely set in the
comparatively thin alveolar border between the canines, and
showed a decided tendency to drop out in old age, their alve-
oles afterward becoming entirely closed. This is well shown
in several instances where on one side of a jaw there may be
1, 2, or 3 incisors, while on the other side of the same jaw
there will be 0, 1, or 2 and no alveole for the missing tooth.
The number of incisors can hardly be considered as of either
generic or specific importance in the Titanotheride where
they are no longer functional and vary with individuals in
the same species and with age in the same individual. The
same may be said of the presence or absence of the first pre-
molar.
In the structure of the last upper true molar we find a
change taking place in the development of a posterior, inner
216 ? The American Naturalist. [March,
cone. Individuals from the base of the beds show scarcely
any indications of this cone, but as we proceed upward in the
beds, there is a marked increase in the development of the inner
basal ridge on the inner posterior angle of the last upper true
molar. This development frequently succeeds near the middle
of the beds in producing a perfect posterior inner cone.
Other variations are noticeable in the dentition and in the
character of the nasals, but they are common to individuals
from any horizon whatsoever, and are thought to be of sexual
importance only. Among variations of this nature may be —
mentioned the following: Differences in the size of the canines —
as compared with the molars and premolars; presence, absence,
or want of continuity of an inner basal ridge on the superior
premolars; comparative strength and rugosity of the nasals.
Since slender and delicate nasals, small canines and upper
premolars with only a very slight or no basal ridge on the
inner border, are associated in the same skulls; these skulls
are considered to have belonged to females, while those skulls
in the same horizons, with stronger and more rugose nasals,
larger canines and a well developed inner basal ridge on the
upper premolars represent their male companions.
There were also changes taking place in other portions of
the skeletons of these animals, important among which may
be mentioned the development of a third trochanter. Femora
from the bottom of the beds were quite small and with only a
rudimentary third trochanter. Toward the middle of the beds
they increase in size and show a well marked third trochanter,
while at the top of the beds this process has become quite well
developed and is of moderate size. :
The most important change that took place in the Titano-
theriide, and the only one perhaps of generic importance, 8”
be found in the fore foot. Early in the season of 1886 the
writer discovered near the base of the Titanotherium beds a
portion of the skeleton of a very small individual with both
fore feet and limbs in position. The carpus of this individual
was found to possess an additional bone (the trapezium) hitherto
uhknown in the Titanotheriide. This fortunate discovery
a skeleton, at the base of the deposits, with a distinct trapezium
1893.] The Titanotherium Beds. 217
showing a perfect articulation with the trapezoid, may be
found to be of considerable importance as establishing a closer
relation of Titanotherium to Diplacodon from the upper
Eocene than has yet been pointed out, especially if this latter
form should prove to possess a trapezium, as it is now almost
certain it did. The trapezium, though present in the earliest
forms of the Titanotheridz, was quite small and soon disap-
peared entirely, leaving no vestige of a first digit. Among
many more or less complete fore feet found in the upper part
of the Lower, in the Middle, and Upper beds, no trapezium has
yet been found, while several have been found at the base of the
beds. :
It is quite probable that those forms of Titanotheriidw with
three incisors described by Professor Marsh under the generic
name of Teleodus will be found to possess a trapezium.’ If
such should prove to be the case, the genus Teleodus, Marsh can
then be considered as distinguished from that of Titanotherium,
Leidy by a definite and constant character, viz., the presence
of a trapezium. Individuals with three incisors collected by
the writer have invariably been found at the bottom of the
beds and the small size of other individuals with three inci-
sors, discovered by other collectors, indicate that they also
were taken from the same horizon. There would appear,
therefore, to be little doubt that the presence of a trapezium
and three incisors are associated in the same individuals. That
the latter character was not constant is shown by the presence
of three incisors on one side and two on the other in the same
jaw, with no alveole for the missing tooth.
Variations occur also in the number of sacral vertebra.
But since these are clearly dependent upon the age of the
individual they are not considered here. All fully adult speci-
mens show four sacrals, younger ones only two or three.
In figure 3 the side view of a skull from the base of the
‘Titanotherium beds is represented, showing the small round
horncores and long nasals which are characteristic of skulls
from this horizon. In figure 2 the side view of a skull from
the Middle beds is represented, showing horncores and nasals
of medium length. Figure 1 represents the side view of a
8m. Jour. Sci, June, 1890, p: 524.
-~
to
_—
io 2)
The American Naturalist. [March,
Oreodon Beds.
Reddish brown clays, with occasional concretionary layers,
Remains of Oreodon, Hyracodon, Hyzenodon, Elotherium, etc.
About 500 feet thick.
Upper Beds, 30 ft.
_ Characterized by Titanotheriidz of large size. Horns 10-
18 in. long, elliptical to sub-ovate in cross-section. Nasals
very short and pointed. Incisors never more than 2. Inter-
nal cingulum on upper premolars not strongly marked in
either sex. Posterior inner cone on last upper molar. Third
trochanter present. Trapezium absent. General form of
skull shown in fig. 1.
Middle Beds, 100 ft.
Characterized by Titanotheriidz of medium size. Horns
4-10 in. long, circular to sub-triangular in cross-section. —
Nasals of moderate length, with broad or pointed extremi-
ties. Incisors never more than 2, Sfrong internal cingulum
on upper premolars of males only. Posterior inner cone on
last upper molar. Third trochanter present. Trapezium
absent. General form of skull represented in fig. 2.
TITANOTHERIUM BEDS, 180 FEET THICK.
_ Lower Beds, 50 ft.
Characterized by Titanotheriide of small size. Horns
rudimentary or from 1-4 in. long, circular in cross-section
Nasals long and pointed. Incisors occasionally as many aS
3. Strong internal cingulum on upper premolars in males
only. No posterior inner cone on last upper molar. Third
trochanter somewhat rudimentary. Trapezium present in
earliest forms. General form of skull shown in fig. 3.
Underlying
Represented by various formations from Laramie to Archean.
Section of Titanotherium Beds showing relative thickness of Upper, Middle, and
Lower Beds, with brief descriptions of forms of Titanotheriidee common to
1893.) The Titanotherium Beds. 219
skull from the Upper beds with the'characteristic long horn-
cores and short nasals. The nasals are not -sọ short in this
Fic. 1.
S nea
Titanotherium acre, py Cope.
Fic. 2.
Titanotherium ingens Marsh; yy (After Marsh).
Fic. 3.
Titanotherium heloceras Cope ; tz (After Marsh).
skull as in many others. The figures are drawn to the same
scale, one-twelfth natural size, and represent well the increase
220 The American Naturalist. (March,
in the size of the skulls from the base to the summit of the
beds. They are arranged on the opposite page in the order
in which they occur in the beds commencing at the top.
Intermediate forms in the middle of the beds have a much
greater vertical range than their smaller ancestors in the
underlying beds or their larger survivors in the overlying
eds.
Between the skulls here figured there are many intermediate
forms, showing every stage of development from a very small
skull found at the very bottom of the beds, which measures
only 22 inches from occipital condyles to the extremity of
nasals, with very rudimentary horncores only 14 inches high
to forms similar to that represented at the top, which measure
more than 36 inches from occipital condyles to extremity of
nasals, with horncores over a foot long and having an expanse
of two feet or more.
In the section of the Titanotherium beds accompanying
this paper the relative thickness of the Lower, Middle, and
Upper beds are shown, with brief descriptions of forms of
Titanotheride characteristic of each. a
Much additional information bearing upon the develop-
ment which took place in the Titanotheriide will doubtless be
brought to light, when the large collections either made or
purchased by the writer for use in the preparation of Professor
Marsh’s monograph and now numbering nearly 200 complete
skulls and many more or less complete skeletons shall have
been worked up. It issafe to say that there is scarcely a single
foot of sediment within the entire vertical range of the Titano- :
theridæ that is not represented by material in this collection.
Only those facts bearing upon the evolution of the Titano
theriidæ which have presented themselves to the writer m
connection with his field work are set forth in this paper-
is to be hoped that the few results here recorded may suggest
to other collectors the importance of keeping, as nearly as po%
sible, accurate data of horizons; since a familiarity with the
These facts were communicated by the writer to Professor Marsh in 168.
were referred to by the latterin the Am, Jour. Sci., Feb., 1889, p. 163, and the Nun
Ann. Report U. S; Geol. S., p. 114. fe
1893.) The Tilenstheriam Beda. 221
different horizons as characterized by different and closely
related forms is clearly indispensible in making an intelligent
classification of any group of fossils, more especially when
they are as highly specialized and as susceptible to the influ-
ence of constantly changing environments as were the Titano-
theriide.
Yale Museum, Jan. 16, 1893.
229 The American. Naturalist. [March,
AN ORGANISM PRODUCED SEXUALLY WITHOUT
CHARACTERISTICS OF THE MOTHER.
By Tu. BOVERI.
[In offering a translation of Dr. Th. Boveri’s paper, entitled
Ein geschlechtlich erzeugter Organismus ohne miitterliche
Eigenschaften, I have been guided by two motives. First, to
make this paper, which will certainly become a classic in
Biological literature, accessible to American students, since the
journal in which it appeared “Sitz. d. Gesellschaft für Mor-
phologie und Physiologie zur München ” X Sitzung am 16 Juli,
1889, has a very limited circulation in this country.
In the second place, to point out the new avenues of research
that such work opens. Results of this kind are of the utmost
importance, inasmuch as they touch the very heart of the
question of Heredity. Each advance in our knowledge gained
by experimental work of this sort, carries forward rapidly our
understanding of the most vital phenomena of life.
Results of this importance must be verified over and over
again, until all chances of error (by no means small) are elimi-
nated. Dr. Boveri writes that during the present Winter it is
his hope to carry forward this work at Naples.
Three small wood cuts were given in the original paper.
Dr. Boveri has most generously placed at my disposal the orig-
“inal drawings as well as five additional figures. These are
added to the present account. The original figures gave side
views of the Echinus larva, the Spherechinus larva, and the
side view of the bastard larva (Echinus 4, Spherechinus @ ).
Three of the new figures give posterior views of the same larve
and two of the Dwarf Larve.
Weismann, referring to Boveri’s work, in his essay on Amphi-
mixis affirms that two spermatozoa enter the enucleated egg to form
the segmentation nucleus. I wish to call attention to the foot
note in Boveri’s paper, where it is positively stated that only @
single spermatozoon enters the enucleated egg.
T. H. Moraay. |
1893.] Organism without Characteristics of Mother. 223
Although the law, that the substances giving the definite
and hereditary characters to the cell are entirely contained in
the nucleus, is at times spoken of as a very probable hypothe-
sis, but again even as a fact, yet it may be easily shown that
this can be known to us neither in the well known phenome-
non of fertilization of the egg, nor through the researches
already carried out, concerning the rôle of the nucleus in the
protozoa.
Simple reflection shows moreover that the determination
whether or not this Theory of Inheritance (Vererbungs-Theorie)
is true, can be settled in one way alone, viz., to take two differ-
ent sorts of cells, utilizing the nucleus of one and the proto-
plasm of the other, to form a new cell. If the nucleus and
protoplasm are so constituted that they can exist together, then
will the properties arising from this cell, made artificially,
answer our question. For then either the exclusive qualities
of that cell will develop which had held the nucleus, or those
will arise that come from the protoplasm, or lastly, from a
mixture of both; showing whether the nucleus alone or the
protoplasm alone (or both of the two constituents) is able to
transfer to the other the properties it possesses on account of
its origin, so as to bring about a reverse result in the other.
The idea of inheritance in the narrower sense corresponds to
such a choice of the two cells, that the cell produced arti-
ficially out of the nucleus of the one and the protoplasm of
another forms a cell capable of development.
Moreover in the case of the egg, as in no other, an opportu-
nity is given to solve the question; because for no other cell do
We possess SO accurate a method of judging of its qualities as is
given in the egg by means of the adult organism that comes
from the egg.
Rauber' has already isidi on a wodburel of this sort. He
describes it as follows. “The nucleus of the first segmentation
sphere of a frog’s egg was drawn out an hour after fertilization,
by means of a pipette stuck into it. The same process was
carried out in a fertilized toad’s egg. These two pipettes were
exchanged, and the nucleus of the toad’s egg placed in
1Rauber Zoologischer Auzeiger 15 March, 1886, pp. 17-.
224 The American Naturalist. [March,
the frog’s egg, and the nucleus of the frog’s egg into the toad’s
egg. Now if the nucleus alone carries the hereditary functions
then there must develop out of the frog’s egg a toad, and out of
the toad’s egg a frog.” As was to be anticipated, the eggs did
not develop further, and at first sight there seems to be no
reason to expect that such an experiment should succeed. For
if we were in a position to take out, without further injury, the
nucleus from cells, still we could scarcely introduce by artificial
means a new nucleus, without causing such fundamental alter-
ations of the one or the other part, as to make further life
impossible.
But just here Nature herself offers the solution by means of
which we can accomplish our purpose, because it makes the
second and more difficult part of the experiment an entirely
` normal process—the entrance of the spermatozoon into the egg
may be utilized.
The basis for my research is founded on the discovery of the
Hertwig brothers? When these investigators shook for a long
time the eggs of the sea-urchin in a test tube, containing a
small amount of water, in order to alter them mechanically,
they found that in consequence of the shaking some of the
eggs fell into pieces, and while some of the pieces contained
nuclei, others did not. It was further shown that these enu-
cleated (non-nucleated) fragments, as well as the nucleated
ones could be fertilized, and that an active segmentative pro-
cess took place in them. What followed in these cell masses
was not determined by the Hertwigs. I have myself verified
this discovery during my last visit to the Zoological station
of Naples, and found that the enucleated and fertilized fragments
of eggs developed as far as and formed as complete larve as did
the perfect nucleated egg. From the enucleated fragments which
I isolated I reared in a series of dishes larvæ’ (about half of
20. and R. Hertwig. Ueber den Befruchtungs und Theilungs-Vorgang des.
Thierischen Eies unter dem Einfluss ausserer Agentien Jena 1887.
` 3In a more extended account of my experiments I will describe in detail the meth- -
ods which must be used in order to be certain that we are dealing with pieces with- =
out nuclei, and on the other hand the best method te. give the more delicate pieces
favorable conditions of development. Here I may remark that in order to obtain
positive results, a sufficient size of the fragment is ‘necessary, also that they have @
nearly spherical form and sufficient amount of water for their development, and that
MONOSPHERIC FERTILIZATION TAKES PLACE,
1893.] Organism without Characteristics of Mother. 225
them developing normally) that formed dwarf-larve. These
were in some cases only one-fourth as large as the normal larve,
but otherwise agreed perfectly with these, and even lived for
the same length of time, i. e., about seven days.
This result is in itself certainly full of interest. It shows
that the sperm-nucleus itself possesses all of the properties
necessary to function as the first segmentation nucleus, and
the result, which shows the prevailing opinion concerning the
nature of fertilization to be erroneous, gives an important sup-
port to my view, published in several places, concerning the
interpretation of the process. I desist from considering the
point further here, in order to turn to the interpretation of the
experiments dealing with the questions raised concerning
inheritance.
With the possibility of fertilizing enucleated egg-pieces and
of bringing these to a normal dévelopment, we have fulfilled
all of the conditions in order to reach the end which Rauber
failed to attain. We have an enucleated egg—for the frag-
ment has the value of an entire egg, and we are able by means
of a process of fertilization to introduce another nucleus into
the egg, the egg still possessing the power to develop. And
now instead of using a spermatozoon of the same species, we
introduce the spermatozoon of another in order to produce the
hypothetical case of “the toad’s egg and the frog’s nucleus.”
It is even possible to bastardise the egg fragments (obtained
by shaking) of one species of sea-urchin with the sperm of
another species, and to rear them far enough to determine
whether the developing organism shows the qualities of both
species or only of the one species. Forsuch an experiment the
sea-urehin presents favorable conditions. It is possible,
although the results are somewhat variable, to cross-fertilize
two species standing quite far apart. And in the second place
the larva of the sea-urchin shows in a very few days, prin-
cipally through the development of the calcareous skeleton, a
very well defined shape. This is constant for each species and
so well defined for each species, that at the age of four days
we can distinguish the two species of larve just as easily and
surely as in the adult condition.
16
a
226 The American Naturalist. [Marech,
The two species that I have used in my research are Echinus
microtuberculatus and Sphzrechinus granularis. I am certain
that amore favorable combination than this can be found, for
although the two species are well adapted inasmuch as their
larvee differ very considerably from one another, yet there is to
Fig. 1. Echinus microtuberculatus larva; pure type.
be taken into account, the fact that cross-fertilization between
the two is so difficult that in my experiment out of 1000 picked
eggs (Sphzrechinus 9, Echinus ¢) nota single fertilized egg
reattiod Of the four species found at Naples in great pba
.
1893.]
Fig. 2. + i é le re typ Bra -
ance, these two alone are useful, and lend themselves to the
solution of our problem.
228 The American Naturalist. [March,
Since the differences throughout are dependent on the dis-
tinctive peculiarities of the larve used, it is necesssry first to
make clear the difference in structure of the larval forms that
we are considering. The two accompanying figures will ren-
der a long account needless. :
Fig. 1, A and B, represents an Echinus larva, and fig. 2, A and
B, that of Spherechinus. The two figures oriented alike, show
the circumference of the body and the calcareous skeleton seen
in profile [and in front]. The differences in structure of the
Fig. 3.
Larva of bastard of Echinus microtuberculatus 3 and Spharechinus granularis Q -
body, particularly in the sheltered structures, are brought in
these figures in the position best suited to the observer. The
arrangement is sufficiently well shown in the figures to obviate
any further description The enucleated egg-fragments of
Spherechinus were brought together with the sperm of Echinus.
`The generic name will suffice, from this on, in speaking of these two species.
PLAE S eres
1893.] Organism without Characteristics of Mother. 229
If the nuclear substance alone is the bearer of the parental
qualities, then larva of the pure Echinus type will be produced
(Fig. 1). It is clear that in the experiment this conclusion can
only be proved, if the bastards produced from fertilized eggs
(we will call these the genuine Bastard larve) give a middle-
form standing between the larve of the two parental species,
For it is conceivable that the particular structures of the two
Fig. 3.
Larva of bastard of Echinus alee 8 and Sphaerechinus granularis Q .
species might not be able to commingle, so that the larvæ
would, according to their surroundings, develop either the
paternal or maternal qualities alone (the qualities of the
other parent in each case remaining latent). In this case there
would be no true bastard.
I have reared such larvæ in great numbers and from many
different individuals, and have in all certainly seen more than
230 The American Naturalist. [March,
a thousand of them. The result was always the same. All
genuine bastards, with not a single exception, represent both in
the shape of the body as well as in the skeleton an almost
exact middle form standing between the two parents. This is
a new well defined form, and is always as such to be recog-
nized and cannot be confused with either of the two parent
larval forms. Fig. 3, A and B, which shows such a genuine
Bastard larva, illustrates, as compared with figs. 1 and 2,
this law better than many words could possibly do.
The result is otherwise if we shake into pieces the Sphere-
chinus egg before the entrance of the Echinus-spermatozoon.
Certain of the eggs remain intact after the process, and these do
give the genuine bastard form (Fig. 3). These -are those
formed from nucleated pieces, but another portion of the crossed
H i
Fig. 4. Dwarf bastard larva of Echinus g and Spherechinus Q (fragment) of pure
Echinus type.
larvæ agree entirely with the simple larva derived from
the egg and spermatozoon of Echinus microtuberculatus (Fig.1)-
These must have been produced chiefly from enucleated frag-
ments. (Fig.4 A and B.)
That this is the correct interpretation is more clearly demon-
strated by rearing in isolation the enucleated pieces. * My
1893.] Organism without Characteristics of Mother. 231
results in this direction are on the whole not successful, and
the explanation for this is to be found in the circumstance cited
above, i. e., that the cross-fertilization between Echinus microtu-
berculatus 3 and Sphxrechinus granularis 2 were scarcely ever
successful out of 1000 cases. This small percentage of the
whole number fertilized, where many millions of eggs or egg-
fragments may be used, lends great improbability to the chance
of getting successful results by isolating a few chosen examples
of eggs. It is such a long and tiresome procedure to pick out
and isolate the enucleated fragments from the shaken mass
that in the limited time of my stay at the sea-shore I did
not have the good fortune to see them develop in a single
example.
I hold as sufficiently strong proof, in absence of direct
knowledge, the result of those experiments in bastardization
where enucleated were mingled with nucleated fragments, and
with entire eggs. For after I have been able to reach the con-
clusion on the one hand by observing eggs and spermatozoa of
the same species that enucleated pieces still develop, and on
the other hand after it has been shown that the fertilization of
nucleated eggs of Spherechinus with Echinus sperm larve
occasionally arise standing between the parent forms, there
remains no other interpretation as to the origin of the
dwarf-larva of pure Echinus except out of the enucleated frag-
ments found there. But there remains a more definite result
pointing to the correctness of such a conclusion. One can dis-
tinguish in a preserved and colored larva whether or not it has
originated from a nucleated or enucleated egg, and this from
the size of its nuclei. If the fertilized enucleated egg-fragment
was the first segmentation nucleus which is formed entirely from
A SPERM ATOZOON NUCLEUS, the nucleus is half as large asa normal
first segmentation nucleus, and this difference in size is handed
down to all of the descendants of the egg cell, even to the lar-
Arg b Ree If now we B larvæ of the. same age derived
ego-fi tilized by Echinus
sperm, we have mixed together with the Bastard-type the true
Echinus type, and the latter (true Echinus type) shows propor-
tionately considerably smaller nuclei than the preceding one.
232 The American Naturalist. (March,
All doubt is thus removed, and it may be regarded as con-
clusive that by bastard fertilization of whole eggs or protoplas-
mic pieces having nuclei, larve are formed that stand midway
between the larval forms of the parent species; but the larvae
arising from the enucleated fragments of eggs have entirely
the characteristics of the parent (male) species. Herewith is
demonstrated the law that the nucleus alone is the bearer of
hereditary qualities. With the removal of the maternal nucleus
are removed at the same time the maternal hereditary ten-
dencies of the egg. And the maternal protoplasm, although
in this case furnishing a large share of the material for the
development of the new organism, is without influence on the
form of the organism.
The phenomenon of growth of the enucleated protoplasmic
mass is entirely governed by the introduced spermatozoon, and
indeed to conclude by analogy, by the nucleus of the sperm.
Whether the spermatic nucleus penetrates the egg protoplasm
of its own species or of another species, the same result appears,
namely, the larva is that of the male species. I might say
that the nucleus possesses an assimilating power toward the
protoplasm, since it produces the same sort of organism from
material that appears to be different, according to our stand-
ards of truth. That the egg-protoplasm is similar in the two
cases in every respect pertaining to its chemical constitution,
I cannot admit, and do not even hold to be probable. Still
such differences may, if they were present, be very easily
explained, as due to the previous action of that nucleus, to
which the enucleated protoplasm was at one time united, and
any difference would thus not be incompatible with the law
that all changes of form of the protoplasm are dependent on the
constitution of the nucleus.
1893.] Classification of the Longipennes. 233
ON THE CLASSIFICATION OF THE LONGIPENNES.
By R. W. SHUFELDT.
Few groups of birds there are that have received a greater
amount of attention at the hands of comparative morpholo-
gists than the one I here consider as the Longipennes.
Considered apart from related ones, the suborder is fairly
well circumscribed, and structurally, very surely a most homo-
geneous collection of bird-forms. Itis abundantly represented
in the United States avifauna, and here, as in nearly all other
parts of the world, consists chiefly of the Gulls and Terns, the
Skuas and Jaegers, and the Black Skimmer of the family
Rhynchopidæ. We have the Skuas represented by but one
species, Megdlestris skua; and there are three Jaegers of the
genus Stercorarius. There are upward of thirty Gulls referred
to the genera Gavia, Rissa, Larus (20 species), Rhodostethia,
and Xema, and about seventeen Terns, which are referred to
the four genera Gelochelidon, Sterna (13 species), Hydrochelidon,
and Anous.
Among the earlier classifiers of the Laridæ, we find P. Bruch
and C. L. Bonaparte, the former being the author, of two
celebrated papers', and the latter of another which appeared
between the times of their publication’. The confusing and
1Bruch, P., Monographische Uebersicht der Gattung Larus Lin. J. f. O., i, 18538,
Ibid. Revie oe Gattung Larus Lin. J. f. O., iii, 1855, pp. 273-293, pll. IV, V.
2Ronaparte, C. L., Notes sur les Larides. Naumannia, IV, 1854, pp. 209-219.
Of these two exploits Coues says in his Ornithological Bibliography, “It may be
remarked, in fine, of this article [ Bonaparte’s], that it is worse than worthless, being
pernicious. It is ostensibly a review of Bruch’s paper of 1853 [supra]; this being
itself a very incompetent performance, confusion is here worse confounded by Bona-
parte’s criticisms and ‘‘rectifications.”” It seems to have had, among other undesira-
ble results, the effect of setting Bruch at the babiaees nen, as cobra age » zy garis
paper of 1855, [above cited]. The two auth
as can be found in ornithology; woe to the confiding student who trusts either of
themn—Crede — ! Bruch and Bonaparte are “ Scylla and Charybdis of Gull
literature.” (p. 1001.)
234 The American Naturalist. [Mareh,
misleading statments made in these three memoirs were
subsequently to a large extent, eradicated by the excellent
labors of Coues which appeared a few years after them,—
although Coues, as he himself admits, suffered from the
reliance he placed in their observations.’
But we are more interested in those works wherein the
taxonomy of the Longipennes is based largely upon the struc-
ture of the various species, and this Dr. Coues gave us later in
his “ Birds of the Northwest,” a very solid contribution to the
subject. Here a very thorough study was made of the
Gulls, Terns, Skimmers and others. His final classification,
however, appeared ten years later, or in 1884° and stood thus:
SUB FAMILIES,
[arin
ORDER. SUB-ORDER. FAMILY. } La
Longipennes. Gavie. Laride. | Sternin
[R Rhynichopitee.
Other Laridists have also ably treated the group and we
can here but refer the student to the works of Nitzsch, Huxley,
Schlegel, Blasius, Saunders, and Sclater and Salvin, bringing
us dows to the year 1880. Of these Huxley’s work is deserving
of especial mention f and he upon certain structural characters
placed the Gulls and their allies in his group “ Cecomorphe,”
remarking that “ This group contains, the Laridæ (Longipennes
Nitzsch), the Procellariidæ, the Colymbide, and the Alcidx.”
(p. 458). As a knowledge of the morphology of a number of
these families became extended, this classification has been
very much modified and altered. Forinstance, Alfred Newton
in the 9th edition of the Encyclopedia Britannica under the
article GuLL (v. x1, p. 274) says, “The Family Laride is
composed of two chief groups, Larinæ and Sternine——the Gulls
Coues, E., Revision of the Gulls (Larine) of North America; based upon
specimens in the Mus of the Smithsonian Institution. Proc. Acad. Nat. Sct.
Phila., XIV, 1862, pp. “291-812 Also a review of the sates ncaa of North
America. Proc. Acad. Nat. Sci. Phila. XIV, 1862, pp. 535-559.
*Coues, E., Monograph of the North American Laridz, 1874, pp. 587-715.
*Coues, E., Key to North American Birds, 1884, p. 732.
®Huxley, Thomas H., On the Classification of Birds; and on the Taxonomic
Value of the Modifications of certain of the Cranial Bones observable in that Class-
P. Z. S. 1867, pp. 415-472.
1893.] Classification of the Longipennes. 235
and Terns, though two other sub-families are frequently
counted, the Skuas (Stercorariing), and that formed by the
single genus Rhynchops, the Skimmers; but there seems no
strong reason why the former should not be referred to the
Larine and the latter to the Sternine.” Without being
especially a classificatory work, W. K. Parker has given us some
valuable researches on the development of the skull of Gavia
ridibunda that has materially furthered our knowledge of the
structure of the Gulls.
In the Check List of the American Orinthologists’ Union,
(1886, p. 84) we find the Longipennes divided into the three
families Stercorariidx, (2) Laride and the (3) Rhyncopidx,—
the second group being again sub-divided into the two sub-
“families, (1) the Larinæ to contain the Gulls, and (2) the
Sternine to contain the Terns.
Finally, I have examined the classifications of Stejneger,
Cope, and more particularly the one proposed by F iirbringer,
and that distinguished ornithologist presents us with the
following arrangement’ :—(p. 1566.)
> G. S. lat. j G. s. sir. F. Charadriidæ.
i] Laro-Limicolæ. | Charadrii. | F. s. str. Glareolidz.
= F. s. str. Dromadidz
& S.O. Char- F. Chionididæ.
2 adrii- | F. Laride.
S formes. F. Alcidæ.
a G. Parrze. F. Thinocoride.
5 Parridæ.
S J G. Otides. b potice
His order Charadriornithes, is also made to include the Im.
S. O. Gruiformes and the Im. S. O. Ralliformes, but we are
not especially concerned with them here. Among the most
recent classification we find that of Doctor Sharpe; and the
Longipennes correspond to his Lariformes*, thus :—
kami
> 3 SUB-ORDER. SUB FAMILIES.
P FAMILIES, inæ
bE Lari. 1. Stercorariidze. l Sterninæ
TA J2 Laridæ. Rhynchopinæ
O ~
` TFürbringer, Max, Untersuchungen Zur Morphologie und Systematik der Vogel,
Amsterdam, 1888.
Sharpe, R. Bowdler, A review of Recent Attempts to Classify Birds. (An
address delivered before the 2d Intern. Ornith. Cong., on the 18th of May, 1891.)
Buda 1891. A most valuable contribution to scientific ornithology, and of grea
aid to all workers.
236 The American Naturalist. [March,
and he places his Lariformes between his Alciformes and
Charadriiformes. (p. 72.)
My own cabinet contains numerous skeletons of Longipen-
nine birds, and, thanks to the U. S. National Museum, I have
had the opportunity of examining very many more.
Recently I have gone thoroughly over this material and
written out full descriptions of the characters presented on :
the part of our American Longipennes, and chiefly from
osteological data it would seem to me that the following classi-
fications can be sustained as being probably the most natural
one.
SUB-ORDER. FAMILIES. SUB-FAMILIES.
f LARIDÆ. / Leine.
Sterninæ
LONGIPENNES. 1 STERCORARIIDA.
i RHYNCHOPIDÆ.
So far as our American Laridæ are concerned, and it is fair
to presume that the fact obtains elsewhere in the world, we
observe that certain genera among the Gulls almost insensibly
approach in their structure and merge into certain genera of
Terns. Xema sabinii is closely related to such a species as
Sterna paradisea, while Gelochelidon comes near some of the
heavier built Larinz.
Iam of the opinion that the Skuas and Jaegers (Stercora-
riidx, stand between the Laridæ and the Rhynchopidz, being
more nearly related to the first named than they are to the
Skimmers. Indeed it would seem that the family Rhynchop-
idx is more remotely related to either of the other two
families of the Longipennes, than has heretofore generally
been supposed. Rhynchops I have shown in a memoir that I
have sent to the Journal of Anatomy (Lond.) for publication,
is, in some of its osteological characters notably in the skull,
the vetebral chain, and pelvis, not very unlike the fossil
cretaceous bird Ichthyornis, and this is a very remarkable
fact.
Through the Laride of the present SUB ORDER, the Longi-
pennes are found to be not so very distantly connected with the
Alex, the genus Uria among the Auks standing between
certain Gulls and the more ancient types, such as Plautus and
1893,] Classification of the Longipennes. 237
Alca. Huxley has already shown that in another direction
“the Gulls grade insensibly into the Procellariide ” (P. Z.S.
1867, p. 445), and it is now generally admitted that through
an easy gradation, the osteological characters of a Gull or
Tern, pass into those of the Plovers and their allies among the
Limicole.
_ Beyond the Plovers, we are led on the one hand to the Rails
through the Otididz, the Cranes, the Rhinochetidx, and the
Psophiide, while on the other, through other branches we
find the Plovers linked by Hemipodius with the Galline.
238 The American Naturalist. [March,
EDITORIALS.
EDITORS, E. D. COPE, AND J. S. KINGSLEY.
—In the States of New York, New Jersey, Pennsylvania, Alabama,
and some others, the governments have been alive to the importance
of retaining in the office of State Geologist a tried and competent
expert. In this way they have secured the best results with the least
expenditure, and have escaped the inevitable loss which results from
changes in the personnel of an expert office. This loss is serious in
consequence of the continuous nature of the work of a geological sur-
vey. Collections are in process of being formed, and reports of being
written, and at no time can a change be made without a loss of work
already done, and a loss due to the lack of continuity of work already
in hand. Some States have pursued a different course, notably Indiana
and Michigan, where competent men were replaced by incompetent
men, and the surveys have been of little value since those changes were
made. Good geologists competent for both scientific and administra-
tive work are rare, and their excellence is in no way related to their
political affiliations.
It is proposed, we regret to learn, to remove from his place the pres-
ent very competent State Geologist of Illinois, Professor Lindahl.
Not only has this officer had an excellent training for the place, but
he has already done a vast amount of unpretentious work of the great-
est value to the State of Illinois. He not only secured to the State
great collections of its minerals and fossils which would have otherwise
passed into private hands, but he has reduced them to order, so that
they are available to the student and business men of the common-
wealth. He is now engaged in making sections across the State with
the view of preparing a full and final geological map. The accom-
plishment of this enterprise can be safely entrusted to the hands of
Dr. Lindahl, and he should be supported by larger appropriations
than those he has been receiving. He has done much with the limited
funds at his disposal. _
—M. Gréard has proposed to the French Academy that it authorize
a number of changes in orthography. Among these are two which
especially commend themselves to writers on scientific subjects, since
they relate to words mostly derived from Greek roots. He proposes to
abolish the vowel Y, and the diphthong PH, using I in place of the
1893.] Editorials. 239
former, and F in place of the latter. These reforms have been before
the American public for many years, and their utility is self-evident.
If the French Academy of Letters gives them the weight of its
authority, we may hope to see them generally adopted. In anticipation -
of such approval, the Revue Scientifique initiates the movement by
writing “ fisiologie,” “ fisiognomie,” “ psichique,” “ilia,” ete. (Feb.,
1893, p. 175). The American Naturalist will follow the example of
the Revue Scientifique so far as it can do so at present consistently with
the liberty of action of contributors and other persons, whose preju-
dices in favor of the old orthography it is necessary to respect.
—It is to be hoped that the present severe Winter has materially
reduced the number of English sparrows in this country. Some beney-
olent persons have been feeding them, and no doubt have thus reduced
the mortality to some extent. Suchy that their
benevolence is misplaced, since they are sustaining the most active
enemy of the farmer known among the feathered tribes.. It has been
thoroughly proven that the English sparrow is not an insectivorous
bird, but that it destroys great quantities of grain and fruit. It also
drives away the insectivorous birds, thus doing double injury. It is
not even ornamental, and the loss of the beautiful or melodious native
species from our parks, which it has occasioned, is greatly to be regret-
ted.
` —THE Amertcan ENTOMOLOGICAL SOCIETY occupies a room in
the hall of the Academy of Natural Sciencesof Philadelphia. Anxiety
has been expressed by some of. the members of the former lest they be
compelled to vacate their quarters in the Academy Building. On the
other hand some of the members of the Academy have expressed some
fear lest the Entomological Soeiety. vacate voluntarily. Under the
circumstances it would seem that both parties have the same object in
view. This being the case, cooperation and mutual admiration must be
the result, and the satisfaction of both parties be guaranteed.
240 The American Naturalist. [Mareh,
RECENT BOOKS AND PAMPHLETS.
Aarsberetning, Bengens Museum for 1890. From the Director of the Museum,
Aarsberetning ORE Museum for 1891. From the trustees of the Museum.
Administration Report of the pereas Central Museum at oe 1891-'92.
ALLEN, J. A ENPE escription of a New Species of Perognathus from Southeastern
x On a Small Collection a% Musial from the Galapagos sacs, collected
by G. Baur. Por Bull. Am. Mus. Nat. Hist., Vol. iv, 1892. From the author.
BEECHER, C.—Development of the Brachiopoda, Pt. II. iniia of the
— of Growth and Decline. Ext. Am. Jour. Sci., Aug., 1892. From the author.
E, C.—Life Histories of North ae Birds, with special reference to
their ce ae Habits and Eggs. Special Bull. No. 1, Smithsonian Institution. From
us.
BEUTENMULLER, W.—On i Earlier Stages of sams ce of North American
Moths, Art. viii. Am. Mu s. Nat. Hist. From the a
BRA aea ’s Klassen und Ordnungen des Thier-Reichs. Ccelenterata,
Tenio, Würmer, Gliederfüssler, Vögel, Säugethiere.
Bulletin No. 42, Massachusetts State da E eeN Station, 1892.
BURCKHARDT, R.—Das Gehirn Agathaum abell om us Marsh. Separat
abdruck aus dem Neuen Jahrbuch fiir Mineo < tee m the author.
CADY, A. C.—The American Continent an ts Takaia sion Columbus, Pts.
14, v 16. From Gebbie & Co., Publishers.
Cort, E. D.—A Synopsis of the Tiid genus Cnemidophorus. Trans. Amer.
Tip Sic, Vol. n PRG:
CORTAZAR, D. DeE.—Descripción Fisica y Geologica de la Provincia de Segovi
Extr. Bol. de la k del Mapa geológico. Madrid, 1891. From the wae
DAREWIN, G. S.—Synopsis of the Lives of V. C. Woodhull and T. Claflin. From
the on
DE T, M., AND M. LoHEsT ee TaS du Mammouth à Spy
Wik de Namur (Belgique). From the ;
EERS ns and Regulations of the West Laurel Hill Cemetery. 1891.
ALFRED.—Ave nueva de Mexico Dendroica dugesii, H. K. Coale. La
eee 2d Series Tomo 2. From the au
Exposition Rétrospective du Travail et des Scie Anthropologiques. Sec. 1,
888.
Anthropologie-Ethnographie, Pari
GARMAN, S.—The Discoboli, Crop, Liparopside, and Liparidide. Mem.
Mus. Comp. Zool., Harvard Coll., V: v, No. 2, 1892. From thea
uthor.
GEGENBAUR, C.—Die Epiglottis. beein Anatomische Studie. FE 1892.
From the avthor.
1893.] Recent Books and Pamphlets. 241
Hunt, T. S.—Sy-tematic Mineralogy Based on a Natural Classification. New
York, 1891. From the author
KINGSLEY, J. S.—The Head of an Embryo Amphiuma. Ext. AM. NAT., 1892,
. 671-6
KRASNOPOLSKY, A.—Carte Géologique génerale de la Russe. Feuille 126.
Mémoires du Comité Geol., Vol. xi, No. 1, 1891. Perm-Sohkamsk. From the
ence
— Beiträge zur Anatomie des Myrmecobius facsiatus.—Zur tre se
ur SEn AAS Ext. Biol. Foren. Förhandl. Verhand]. Biologio. Ver. Stoc
holm, 1891. From the author.
Macoun, J., AND N. C. Kioii .—Catalogue of Chnadiaa Plants, Part vi.
——Musci, 1892. From the author.
Map of the Madras Presidency. From the Government Central Museum, Madras,
1892.
Monthly Review of the Iowa Weather and Crop Service, — iii, Nos. 2 and 3,
1892. From J. R. Sage.
MourLon, M.—Sur une réforme à oe a dans les Bibliothiques Scientifiques.
Ext. des Ann. des Tray. Pub. de Belg., T. xli
Légende de la Carte Geologique de Kitali: Groupe Quartenaire. From
the author.
NEGRI, A.—Trionici Eocenici ed Oligocenici del Veneto. Mem. estr: dal Tom.
viii, Serie 3a, No. 7, della Soc. Ital. Sei
Ninth Annual Mapai of the Board of Control of the State Agricultural Experi-
ment Station at a Mass., 18
Osporn, H. F., L. WORTMAN. E r of the Wasatch and
Wind River Beds, aaa of 1891. Ext. . Nat. Hist., Sept. 1892.
From the authors
PoHLic, H oi and ae des Elephas antiquus Falc. mit Beiträgen
über Elephas primigenius Blum. und Elephas meridionalis Nestl. Extr. Nova
Acta der Kgl. Leop.-Carol. Deutschen Akademie der Naturforscher, Bd. lvii, No.
5. Halle, 1891.—— Ueber neue Ausgrabungen von Taubach bie Weimar. Ueber
Petersburger fossile Säugethierreste.—— Ueber Americanische Proboscidierreste.
—— Ueber Glacial geschiebe von Leipzig. Separat-Abdruck aus den Sitzungs
Re site der Niederrh. Gesellschaft in Bonn. Feb., 1891. From the author.
PILLING, J. C.—Bibliography of the Algonquian Languages. Wishincin, 1891.
From me Suites Institution
PRESCOTT, A. gags ress before the Am. Asso. Adv. Silanes, Aug., 1892. Ext.
Proceeds. A. A. A. S.,
Prospekt Allgemeine pr Leipzig und Wein. From
Rust, Dr.—Beitrage zur Kenntwiss der fossilen Radiolarien aus Gesteinen der
Trias und der palzozoichen Schichten, ipri? Beiträge zur Naturge-
schichte der Vorzeit, herausgegehen von K. A. Zittel. Acht und dreissigster Band.
Dritte bis sechste Lieferung. Stuttgart, 1892. From Karl A. Zittel.
RUTIMEYER, L.—Die Eocāne Säugethier-Welt von Egerkingen. Abhandl. der
eeu. palicntologischen Gesellschaft, Vol. xviii, 1891. Zürich, 1892.
From the author.
242 The American Naturalist. [March,
SAITZEW, A.—Geologische untersuchungen im Nikolai-Pawdinschen Kreise und
lan sae im Gebiete des Central-Ural und dessen östlichen Abhinge. Mém
Comité Geol., Vol. xiii, No. 1, 1892. From the author
Scanian Leon. U. v.—Das Mikr apes Wien, Leipzig, 1892. From
HARMAN, G., AND E. T, NEwron.—On a new form of Agelacrinites (Lepidodis-
cus milleri n. sp.) from the Lower Carboniferou Limestones of Cumberland. Extr.
Quart. Journ. Geol. Soc., May, 1892. From the author
SHUFELDT, R. W.—Notes on the Blowing Viper. ba Great Divide, July, 1892.
From the author.
SIEBENROCK, F.—Zur Kenntwiss des Kopfskelettes der Scincoiden, Anguiden und
Gerrhosauriden. Separat aes aus Band vii, Heft. 83,1892. From the author.
imen Map from W. . K. Johnston’s School Atlas of Physical Geog-
raphy, 1892. From W. and d 3 eats
hird Annual Report of the Missouri Botanical Garden, 1892. From the Board
TROUESSART, E.—Les Primates tertiares et Phomme fossile sud américain. Extr.
L’ Anthropologie Mais e 1892, No. 3. From the author.
TUCKERMAN, F.—On the Terminations of the Nerves in the Lingual Papillæ of
the Chelonia. Aus der ieni Monatsschrift f. Anat. u. Phys., 1892. Bd.
ix, Heft 1.
The Gustatory Organs of Ateles ater. Extr. Journ. Anat. and Phys., Vol.
xxvi. From the author.
1893.] Recent Literature. 243
RECENT LITERATURE.
Memoirs of the National Academy of Sciences, Vol. V.—
The memoirs of 1891, five in number, are published in the usual
quarto form under the following titles: Energy and Vision, by S. P.
Langley ; Contributions to Meteorology, by Elias Loomis; Report of
Studies of Atmospheric Electricity, by T. C. Mendenhall ; The Embry-
ology and Metamorphosis of the Macroura, by W. K. Brooks and F.
H. Herrick ; On the Application of Interference Methods to Astro-
nomical Measurements, by A. A. Nicholson. The illustrations are
numerous and good.
Brooks and Bruce on the Embryology of the Macroura.!
—The two hundred and fifty quarto pages of this memoir are not
easily summarized in a page or so of THE Natura.ist, for they
include a wide range of subjects. After an introduction by Professor
Brooks comes an account of the life-history of Stenopus hispidus by
Professor Herrick ; next the habits and metamorphosis of Gonodacty- .
lus chiragra by Professor Brooks ; fourth the Metamorphosis of Alpheus
sauleyi by Brook [sic] und Herrick, while the bulk of the volume is
taken up by Dr. Herrick’s paper: Alpheus, a study in the development
of the Crustacea, which extends from page 371 to the end. ;
The most interesting facts connected with the reproduction of the
almost cosmopolitan Stenopus are the hatching of the larva as a pro-
tozoxa with enormous mandibles and its later metamorphosis into a
larva with an enormously developed fifth pereiopod, the use of which
as a swimming organ has doubtless played an important part in the
wide distribution of the species.
Much more important are the observations on Gonodactylus, for
every fact concerning the early embryology of the Stomatopods is a
positive addition to knowledge. Gonodactylus, like the others of its
tribe, deposits its eggs at the bottom of its burrows, where they are
aérated by the currents produced by the pleopoda of the parents.
From these eggs the young hatches in an advanced condition, five
abdominal segments with their appendages being outlined before the
young escapes from the chorion. Professor Brooks was enabled by this
material to conclusively show that the larval Stomatopod to which he
The Embryology and Metamorphosis of the Macroura, by W. K. Brooks
and F. H. Herrick. Memoirs Nat. Acad. Sci., Vol. v, pp- 321-576, 57 pls.
Washington, 1892.
244 The American Naturalist. [Mareh,
had previously given the name Gonerichthus is the young of Gono-
dactylus.
The account of the various species of Alpheus is most detailed and
one of the most striking facts brought out is that the same species in
different localities may have an almost totally different development.
Thus, in the Bahamas, Alpheus heterochelis? hatches from the egg with
all its appendages functional as far back as the third pair of maxilli-
peds, while the pereiopoda 1, 2, and 5 are bud-like rudiments, and the
joint between abdominal segment 6 and the telson has not appeared
and no pleopoda are outlined. At Beaufort, N. C., the species is much
more advanced before leaving the egg, but the stage at which it hatches
is not directly comparable with any stage in the life history of the
Bahaman form. At hatching all the appendages of the adult are
present, and all become functional after the first molt. In the Bahamas
there is a long series of larval stages, while at Beaufort there isa great
acceleration, and even this is not all; the Beaufort stages are so mod-
ified that at no time can exact parallels be drawn between them and
the more southern form. In the Bahamas there are three, then four,
then five, and then seven schizopodal feet with functional exopodites,
while at Beaufort there are never more than three. Yet these differ-
ent types of metamorphosis result in the production of adults which
are almost exactly alike. It must be noted that Packard has described
still another type of development for what he regards as the same spe-
cies at Key West. For the details of the development of the different
species of Alpheus studied we have but little room; those interested
must seek the memoir itself. The segmentation in A. sauleyi and A.
heterocheelis is typical centrolecithal, with the formation of yolk pyra-
mids ; in A. minus it is irregular and has no yolk pyramids, but the
statement (pp. 427 and 457) that it is amitotic deserves further inves-
tigation. All of the nuclei resulting from segmentation migrate to
the surface and there by delamination they produce “ wandering cells”
which pass into the yolk and give rise to both mesodermal and ento-
dermal structures. The subject of degeneration of certain nuclei is
also very interesting. Dr. Herrick has carefully followed the increase
of nuclei in the various parts of the embryo, and has plotted curves
illustrating the distribution of primary yolk nuclei, of wandering cells,
*There isa great diversity in this memoir as to the spelling of various
scientific terms. Thus we have usually heterochelis regardless of the fact
that Say, who described the species, wrote heterochelis; Alpheus minus
appears as minus, minos and minor; centrolecithal sometimes as centroly-
-cethal; Lucifer.as Leucifer.
1893.] Recent Literature. 245
and of all other cells, from which the reader, can almost reconstruct the
. eggs for himself.
Only a few points more can þe noted in the later history. The
mouth invagination occurs on a line between the bases ofthe antennular
buds; the history of the eye is followed, the author agreeing well in
most points with Parker, but affording little support to Watase’s theo-
retical views. The antennal gland (green gland) is regarded as meso-
dermal, but its opening was not found even in the larval stages; the
alimentary canal proper is almost wholly made up from stomodeal and
proctodeal invaginations, the true entoderm, which arises by the migra-
tion of yolk cells to the posterior end of the yolk being chiefly con-
fined to the hepato-pancreatic diverticula and their ducts.
The greatest fault which one can find with the paper is that which
is due to its composite nature, the result being an apparent lack of
arrangement, so that it is difficult to follow in detail certain structures.
This possibly was unavoidable where two authors were each contribut-
ing their parts and also where the composition of the text was done at
different times. The volume is filled with valuable facts and cannot
be ignored by the student of Crustacean ontogeny. It is by far the most
valuable zoological memoir yet published by the National Academy.
Campbell’s Biology.*—It is rarely that such a veritable hodge
podge as this comes to our table. It is an example of absorption with-
out assimilation on the part of the author. The plan of the work is
fairly good but it is a misfortune for any student to have it as a guide
in his studies. It is worse than the notorious works by the late Dr.
Steele, for their faults were largely negative; they taught absolutely
nothing good or bad, but this is “filled with lots of things that are not
so.” The work intends to be a companion to the laboratory work, and
gives much space to protoplasm, the cell and the like, and then takes
up without any apparent order the structures and classification of
animals and plants. A few passages out of over a hundred which we
have marked will illustrate the chief shortcomings of the work.
P. 137. The lungs “ develop as an outgrowth of the alimentary canal.
This outgrowth becomes completely separated off from the oesophagus,
and at its lower end divides into two or more tubes, which communi-
cate with the pharynx by a single tube, the trachea.”—Pp. 145-6. The
statement is made without a single qualification that the ureter of ver-
3Text book of Elementary Biology, by H. J. Campbell, M. D., Lond.
London, Swan Sonnenschein, New York, Macmillan & Co., 1893. 12 O., pp.
xii -+ 254.
246 The American Naturalist. [March,
tebrates is developed from the ectoderm and in the development of the
nephridia “as a rule the ciliated funnel which was present during
development, becomes completely closed.” P. 85. Between the ecto-
derm and the entoderm of the sponges “is a gelatinous layer, the mes-
oglea, amongst the cells of which crystals of lime salts occur,” which
we suppose to be the spicules, silicious as well as calcareous, of these
organisms. P. 83. The animal kingdom is divided into Protozoa, and
Invertebrate and Vertebrate Mesozoa. The Invertebrates are defined
as follows: “ They possess no backbone, the nerve cord or nerve cords
are never dorsal, . . . and the heart is always placed in the dorsal
region.” The Invertebrata are subdivided into Ceelenterata and Cœ-
lomata, but never a word that the Plathelminthes are not Ceelomata
and that the vertebrates are. But enough. The work is well illustra-
ted, mostly by cuts from Claus, Sachs, Prantl and other recent text-
books.
Correlation Papers of the U. S. Geological Survey
Neocene.*—This memoir is the fifth of a series, having been pre-
ceded by essays on the Carboniferous and Devonian by Mr. Williams,
on the Cambrian by Mr. Walcott, on the Cretaceous by Mr. White,
and on the Eocene by Mr. Clark. To an excellent summary of pub-
lished material on the subject discussed the authors have added
important original matter based on personal investigations by Mr. Dall
in the field and laboratory. The following is an outline of the memoir
as given in the introduction:
“ This paper, after discussing general principles connected with the
study and description of the Tertiary or Cenozoic rocks and fossils
contained in them, takes up the Neocene deposits of the United States
in particular.
“ A chapter is devoted to a summary of what is known in regard to
the Neocene of the eastern coast of the United States, each State in
geographical order being separately considered, beginning at the north.
The State of Florida, in regard to which much unpublished informa-
tion was available, being entirely composed of Cenozoic rocks, and
therefore as a type of such structure peculiarly interesting, is treated
of in greater detail and at more length than in other cases. The part
of this essay relating to the State of Florida is really a preliminary
geological report on that State, of which the structure has hitherto
been very little known. The important fact that until the Pliocene
‘Bulletin of the United States Geological Survey No. 84. Correlation
Papers—Neocene, By W. H. Dall and G. D. Harris. Washington, 1892.
ae. Sic ES gall Ae ee So UN aa Sarai ks = is hepa eA tee. =
oe om BOs oid peta
1893.] Recent Literature. 247
period, Florida, so far as it was elevated above the sea, was an island
separated from the mainland by a wide strait, is here first demonstra-
ted. It is also shown that the strata are probably gently folded
lengthwise of the peninsula, and that in the trough now occupied by
the “lake region ” of Florida in Pliocene time a large lake probably
existed, to which the name of De Soto has been applied. The age of the
remains of fossil vertebrate animals, which in south Florida are asso-
ciated with the so-called “ pebble phosphates,” is here definitely deter-
mined.” i
After discussing by States the character and distribution of the
Atlantic Neocene, a chapter is devoted to the consideration of the
general geological movements and fluctuations of land, sea, currents,
and water temperatures which appear to have been concerned in pro-
ducing the characteristics described.
“Tn like manner the Neocene geology of the Pacific coast has been
treated, and in addition to that of California, Oregon, and Washing-
ton, a synopsis of data relating to British Columbia has been included,
together with a summary of what is known in relation to Alaska
during this epoch. The latter discussion contains a large amount of
material extracted from unpublished notes covering some fifteen years’
study and exploration by W. H. Dall in the Alaskan region, and
therefore adds materially to the sum of our knowledge in regard to
that part of the United States.
“The Great Interior region of the west is then taken up, and a sum-
mary of our knowledge in regard to its Neocene geology is brought
together for the first time. While this is necessarily far from perfect,
the very fact that such gaps exist will stimulate the collection of infor-
mation to supply the missing links.
“The essay closes with a list of names proposed for geological beds,
groups, and formations in the American Cenozoic strata, and a descrip-
tion of the data upon which the coloration of the general map is
b. d 9
The work while an eminently important and useful one, is very
unsymmetrical, as the authors themselves recognize. They are very full
in describing the formations that they have seen, those of Florida, for
instance, and deficient in those which they have not seen, as the lacus-
trine formations of the interior. The latter yet remain to receive ade-
quate treatment from the U. S. Survey, since to do this requires the
aid of a competent paleontologist of the vertebrata.
The geological map of Florida contained in this volume expresses
clearly the latest discoveries in that state. The coloration will surprise
248 ` The American Naturalist. [March,
geologists who supposed that Major Powell had abandoned his extraor-
dinary position on the question of coloration of geological maps. We
seem to see in Florida a good representation of the Archean, Paleozoic
and Mesozoic beds, as well as the Cenozoic. For this Mr. Dall is in
no way responsible. It is a pity that expense should be incurred in
printing such maps, since they will have to be republished with the
customary colors.
Cary on the Evolution of Foot Structure.’—We have in this
paper a study of the fore foot of Palseosyops, from a specimen in
the museum of Princeton College, conducted with a view of ascertaining
the mechanical relations of the parts when in action. The ultimate
object is to determine whether the structures presented (facets, etc), can
have been produced by direct mechanical impacts, strains, ete., as is
alleged by the Neolamarckian school of evolutionists. The study is
conducted with care, so far as it goes, but it is not always easy to under-
stand the drift of the author’s argument. He reaches but one definite
conclusion, viz.; that the trapezoid is too small to express properly a
result of direct mechanical causes. This fact, the author says is
incompatible with the Lamarckian principle. He informs us that in
reaching this result he has applied geometrical methods. “ First, the
volume of the bones was got at. Next the area of the bearing surfaces
and their inclination to the digits were measured. Then giving to the
thrust of each metacarpal a value proportional to its volume, the distri-
bution of that thrust can by resolution and composition of forces, be
traced through the foot, and the pressure on each surface and bone
approximately obtained.” Further than this the author does not
explain how he reached the result that the trapezoid is too small. It
is quite essential that this demonstration should be given if we are
expected to accept his conclusion. An essential part of the problem
is, however, unnoticed by Mr. Cary; and that is the condition of the
trapezoid in the reptilian ancestors of the Mammalia. The phylogeny
of an element must be known, since it furnishes the “ physical basis”
of the problem.
. Cary then proceeds to criticize the explanations offered by
Professor Osborn and myself, in accounting for the origin of certain
structures. He finds our explanations to be self-contradictory, and
that we also contradict each other. Osborn has supposed that the
conules of the molars are produced by friction of the molars of opposite
šA study in Foot Structure; by Austin Cary. American Journal of Morphology
Dec. 1892, p. 305.
1893.] Recent Literature. 249
series on each other. I have expressed the opinion that the shear of
the sectorial teeth of Carnivora was produced by lateral friction
during vertical movement of the lower tooth on the upper. I have
also asserted that the forms of facets of limb articulations are due to
pressure. Mr. Cary sees here the attempt to explain the origin of
totally different structures through identical mechanical processes, and
believes that the attempt is a failure. Were the conditions of the
problems alike, as Mr. Cary thinks them to be, he would have good
reason for this opinion. But the conditions in the three cases are
entirely different, and our author’s conclusion is due to neglect of
the elementary facts of the proposition.
The development of conules at the points indicated by Professor
Osborn, has been supposed by him to be due to friction between exist-
ing ridges of enamel which cross each other when in action, at the
points in question. In the case of the development of the sectorial
shear, the faces between which the shearing motion takes place are
smooth, and without ridges or crests. ence the entire surface
receives a homogeneous friction. In the third case, that of the foot
articulations, there is no friction, but there is pressure which when
abruptly applied in movement becomes impact. There is really no
parity between the three cases.
The author of this paper also thinks that the explanation of the
elongation of bones through use of different kinds is not a permissible
hypothesis. He cites my attempt to account for the elongation of the
leg bones of higher mammals through impact-stimulus ; and of other
limb bones of other mammals through stretching.. But he does not
prove that similar results may not flow from mechanical stresses
applied in different ways. I suppose that any mechanical stress which
determines nutritive processes to a part, will increase its size,-
caeteris paribus ; and the stretch as well as the impact has this effect.
se is a term which is too indefinite for purposes of exact demon-
stration, and I have endeavored to reduce it to precision so far as
regards the skeleton, by defining it as “friction, pressure and strain.”
Precisely how these processes affect nutrition is not yet clear. We refer
the production of various animal fluids to “secretion ”, knowing that
the products of secretion are most various, as bile, gastric juice, saliva,
etc. The exact cause of the diversity remains unknown. So with the
effect of stimuli on bone nutrition, we see the cause and the effect, but
the ultimate process, as in all nutrition, has as yet eluded our view.
In concluding, Mr. Cary admits one of the two contentions of the
Neolamarckians in his two closing propositions. He says “ Plasticity
250 The American Naturalist. [March,
of bone, using the word plasticity not in a physical sense merely, but to
include absorption under pressure, will probably account for much
structure in the foot and elsewhere, especially the connection with the
joints, and in the fields of variation and correlation.” In the second
proposition he says that facts have been adduced by him which are
inconsistent with the theory that the size of bones has been increased
by the stimulus they receive, and with the theory that regions of
growth are determined by regions of pressure and strain. “ The testi-
mony of the literature as to the latter point he says is conflicting.” I
have shown that the supposed conflict is due to a misunderstanding on
the part of the author of this paper. The proposition that pressure
does not affect growth is in contradiction to the admission made by the
author in his first proposition, where he admits that pressure determines
structure; for in such change of structure there is always growth.
inally Mr. Cary remarks “ That race changes follow those produced
in the individual life, or that they are directly caused by their mechan-
ical surroundings, I do not think it has been satisfactorily shown. ”
The fact that the characters of bone structure admitted by-Mr. Cary to
have had a mechanical origin appear in the young before birth, is
evidence that race characters are produced, and that they are produced
by mechanical surroundings.
Such criticisms as are contemplated by the author of the paper
reviewed above, are important and are what the subject needs. It is
along the line followed by him that the ultimate demonstration of the
problems involved will be made. We trust that we shall hear from
him again in this field, and that in his labors he will be well supplied
with the phylogenetic details as a foundation.
E. D. COPE.
Earle on the Species of Coryphodontide.’—In preparing
this paper Mr. Earle had the advantage of the use of the material in
the collections of the New York American Museum of Natural His-
tory, and the private collection of Professor E. D. Cope. He presents
us with a brief résumé of the results of his comparisons, and adds con-
siderably to our knowledge of the characters of the skeleton and
dentition of some of the species. He gives a list of the described
species, which number twenty-one, and which were referred by Cope
to five genera. He concludes that these should be reduced to ten
“Revision of the species of Coryphodon, Art. xii, Bull.Am. Mus. Nat-
History New York, iv, pp. 149-66; Oct. 18, 1892.
1893.] Recent Laterature. 251
species and three genera, viz., Coryphodon Owen, Ectacodon Cope, and
Manteodon Cope.
Mr. Earle’s conclusion that the supposed genus Bathmodon is not
distinct from Coryphodon may be well founded, as the material at his
disposal is better than mine. The difference in the forms of the astragali
of the two types is, however, greater than is usual in a single genus, and
is seen in material from all localities. The character on which the
genus Metalophodon rests is a strong one, provided it be constant.
Mr. Earle says it is not constant, and if his material demonstrates this
to be the case the genus must be abandoned. I do not, however, think
that he demonstrates his case in the paper under review.
Let us now see the evidence on which he reduces my reputed twenty-
one species to ten. In the first place he fails to state that I had
already reduced two of the names to the rank of synonyms, leaving
nineteen species to my credit ; that is to say, nine specific names remain
which are alleged to be superfluous. One of these is, however, admitted
to be good by Mr. Earle. He refers C. simus and C. latidens to
C. elephantopus, but says also that the former two are “quite radically
distinct.” Both cannot, therefore, be synonyms of the same species
unless “ things not equal to one another are equal to the same thing.”
I described the lower molars which probably belong to the C. elephan-
topus, and they are totally different from those of the C. latidens. The
superior last molar of C. simus is different from that of the C. elephan-
topus. The three species are in my opinion well distinguished. This
reduces the supposed superfluous names to seven.
Mr. Earle does not admit the Metalophodon armatus for reasons
which are insufficient. As I took the greater part of the dental series
from une decayed skull, and an almost equally large series from a sec-
ond skull, and as the two series confirm each other, I believe the spe-
cies to be one of the most distinct of the family. This reduces the
supposed excess to six. As to the C. cuspidatus, the last inferior molar
teeth of three individuals are now known, and they confirm each other
not only by their characters but by their inferior size. Mr. Earle
admits this species with doubt. The C. marginatus is rejected by him
as probably founded on a milk tooth of C.anaz. But it is not a milk
tooth,® but an unworn permanent tooth of a species of hardly half the
bulk of the C.anax. Coryphodon did not possess milk teeth of this
form. The surplus is now five names. There are three forms of
approximately similar and smaller size, viz., C. latipes, C. molestus and
6] have represented the milk dentition of Coryphodon on Plate liv, fig. 3-
of the U. 8. G. G. Survey, Report Capt. Wheeler, iv, 1875.
252 The American Naturalist. [Mareh,
C. simus, which Mr. Earle refers to the C. elephantopus, but of which
the second and third do not exhibit the form of the last superior molar
which he regards with me as characteristic of the latter species. It is
not unlikely that C. simus and C. molestus are one and the same, but
the evidence is not yet in favor of their being identical with the C.
marginatus. It is probable that these specimens represent at least one
distinct species of rather small size. ;
In conclusion [ think that Mr. Earle has been hasty in his whole-
sale reductions, and that instead of ten species in the American
Wasatch beds there are at least fifteen recognizably described. It is
evident that more material and more research are necessary before a
larger number than this can be demonstrated and before those which
are admitted can be fully defined. A considerable part of Mr. Earle’s
conclusions may be due to the fact that, as he says, he has “ labored
under the disadvantage of not being able to study any of the types of
Coryphodon from New Mexico which have been described by Professor
The Coryphodontidse were the predominant type of the Wasatch
(Suessonian) Eocene, and they were probably numerous in species and
varied in character. If the bones and teeth of the existing African
antelopes were mixed up and discovered piecemeal, they would puzzle
naturalists, who would at first be incredulous as to their representing
over forty species.—E, D. Corr.
1893.] Geography and Travels. 253
General Notes.
- GEOGRAPHY AND TRAVELS:
Africa.—GaRENGANZE.—Garenganze, the land ruled by Mshidi,
between Lakes Mwero and Tanganyika, has recently been visited both
from the east and from the west. Mr. Alfred Sharpe, proceeding from
Lake Tanganyika, discovered west of that lake and east of Mwero
another Mwero into which run the rivers Mkabe, Mwambezi, the
Choma from the north, and the Chisela from the north-east. This lake
is now partially desiccated, and consists of an extensive salt-marsh with
several pools of water, including a large central one about fifteen miles
long by ten wide. The former outlet of this lake was by the Movu
river-bed to the Kalongwizi River, a feeder of Lake Mwero. Kazembe,
the powerful chief who resides on the Luapula, above Lake Mwero, |
though very polite to the traveller, would not permit him to cross
because of his enmity to Mshidi, so that, finding all attempts useless,
Mr. Sharpe was compelled to retrace his steps to Abdallah’s near the
salt-marsh, and from thence to make his way westward round the north
of Lake Mwero. Abdallah is an agent of the famous Tippo Tib, and
bears sway eastward as far as Kabunda, near Tanganyika. The high-
est point between Chipenbiri, on the level of the swamp, ( 3050 feet )
and Lake Mwero, is at an elevation of 3850 feet. The éastern water-
shed of the northern part of Mwero is not more than six to eight miles
from the shore, and that lake is 2900 feet above the sea. The streams
Luao and Luchinda flow into the lake from the north, and Mpweto’s
town, east of the Luabula, is subject to Abdallah. The Luapula, at
its point of exit from the lake, is of less volume than above. Beyond
the lake rises one of those plateaux so frequent in Africa, and the
country beyond this, after passing Chuako (subject to Mpweto ),
belongs to Mshidi. The Luvule River falls into the Luapula not far
from its exit, and the next river reached after crossing the watershed
is the Luvula, an affluent of the Lufira, into which it falls a few miles
above the junction of the Lufira with the Likulwe, from its left bank.
The Lufira itself like the Lapula, is a tributary of the Congo. The
soil of Garenganze is rich, butthe country shadeless. Mshidi is now an
old man; originally he was a trader from the Wa-Nyamyezi, but
became chief. He has many wives. On his return by the same route
1This department is edited by W. W. Lockington, Rugby, England.
254 The American Naturaust. {March,
that he had entered, Mr. Sharpe explored the salt-swamp, and found it
rich in large game. Lake Bangweolo is placed at 4260 feet above sea-
level; thus, as the sources of the Chambezi are only 4400 feet, the fall
of that river must be very slight. Our traveller states that the prac-
tice of maiming the person is rife among the natives southwest of Tan-
ganyika, among whom the Wa-Wemba are perhaps thé worst in this
respect.
The visitor to Mshidi from the west, was Lieutenant Paul Le Marinel,
the service of the Congo State. The route necessitated the crossing of
the Sankuru, Lomami, and Lualaba basins. The first is situated
further to the east than has before been supposed, and the travellers
discovered the Luembe, an affluent of the Lubilash, itself a tributary
ofthe Sankuru. The source of the Lomami was found in 8° 45’ B. and
24° 55 E. long; the river occupies a long narrow basin, running
almost due north and south for 750 miles. During the whole of this
course it does not stray more than a degree from a straight course, and
it receives only two important tributaries, the Lurimbi and the
~ Lukassi. South of the plateau in which the Lomami rises, is a district
named Samba, with a most European aspect. Marinel gives Bunkeia
as the name of Mshidi’s capital. East of the Lualaba the country is
mountainous.
AFRICAN Norrs.—One of the chief discoveries of Dr. Stuhlmann,
who last year accompanied Emin Pasha into his old Equatorial prov-
ince, was the river Kifu, which is stated to have a course of 250 miles
before it falls into Lake Albert-Edward, and isthus the most southerly
source of the Nile yet known. Dr. Stuhlmann started from Kafure, on
the Muta, a tributary of the Kangere, on the west shore of Victoria
Nyanza, descended to the Kagere, then crossed the Mpororo Moun-
tains to the Ruchuru and to Lake Albert-Edward ( Mwutanzige ) at
Vichumbi (0° 44’ S., 2850 feet). The country to the south was a vast
savannah with mountains in the distance, among which Kisigali, 13,000
feet, seems to be the highest. Beyond this is the active voleano of
Virungo Viagongo. Along the west shore of the lake the mountains
approach almost to the edge. From Karevia he attempted to ascend
the snowy mountain (probably a peak of Ruwenzori), and reached
_ 12,500 feet. He found bamboo and grasses from 3850 to 5350 feet ;
‘colocasia beans, grass, and the upper settlements up to 6700; then
deciduous forests with Erica and bamboo to 8530 feet, followed by a
belt of Erica and Vaccinium to 11,800, above which, to the snow line
at 13,000 feet, the vegetation was reduced to mosses, lichens and
1893.] Geography and Travel. 255
Senecio. The plateau west of Lake Albert is known as Lendu. He
reports a considerable shrinkage of the waters of the Albert Nyanza
since it was first discovered, so that islands have become peninsulas.
M. Crampel and his party was set upon by Snoussi and his fanatical
Arabs, who killed all the whites, asserting that they had no business in
the country, but did not hurt the Senegalis who composed his escort.
M. Nebout, who was coming up, carried back the sad news, and the
fate of Crampel was avenged by M. Dybovski, who entered the country
with a considerable force.
On the Mobangi, Captain Van Gele has “forged the last link ”
between that river and the Welle, which is thus definitely included
in the Congo basin.
The Kong Mountains of the maps have been found to have no actual
existence, proving to be but an elevated watershed.
The Italians are endeavoring to explore Somaliland, but some of
their I h t been st Captain B. de Vesme,
however, passed from Berbera by Harrar, over the waterless plains
of Milmil to the rivers Kishen and Amaden, and then to the upper
course of the Webbe.
p + oases
America.—Tar ECUADORIAN AnpEs.—Dr. Wolf, who has devo-
ted much time to the study of the Andes of Ecuador, describes them
as consisting of two ranges, connected at intervals by cross chains, and
thus enclosing basins. The eastern cordillera is as a whole more prom-
inent, geologically older, and of greater average height than the west-
ern, the rocks of which are much more heterogeneous, while its elevation
exhibit greater diversity and irregularity. A porphyritic cross ridge
at the mountain knot of Acayana and Guagrauina has much gold-
bearing quartz, and isthe richest mining district of the country. From
this the eastern cordillera continues onward, broad and lofty, into the-
province of Cuenca, with a wide bend, and a second cross ridge of por-
phyry forms the knots of Portete and Tinajillas. The lofty basin of
‘Cuenca is one of the most beautiful in the Andes, and the chief town
is situated at an altitude of 8460 feet. North of there is the irregular
mountain mass of Azuay, which sends spurs in every direction, two of
which reach the western cordillera. Beyond this the eastern cordillera
becomes of greater extent, and rises into the gigantic volcanoes of
Sangay (17,880 feet), the most active volcano in the world, Altar, or
Collares (17,710 feet), and Tunguragua, (16,696 feet). Opposite to
these, on the western cordillera, rises the still loftier Chimborazo
(20,660 feet). Here commences a third cordillera, parallel with and
256 The American Naturalist. [March,
geologically resembling the western range. The waters of the Laca-
tunga basin flow south and burst through the eastern range. The
drainage of the basins of Quito and Ibarra goes to the Pacific. The
western cordillera, by the Quito basin, is quite low, not more than
10,000 feet, but is raised by superimposed voleanic rock into the lofty
volcanoes of Corazon, (15,804), Atacoza, (14,390) and Pichincha. The
eastern cordillera is here more complicated, and bears the giant volca-
noes of Cotopaxi (19,480), Sincholagua (16,360), Antisana (18,885)
and Cayambe (19,450). Northward the eastern range is continued
into Columbia, increasing in height and breadth, but without any vol-
cano north of Cayambe. In the province of Quito the western range
rises into the volcanoes of Cotocachi (16,295) and Yanaurai (14,000.)
Both Dr. Wolf and Mr. Whymper describe travelling in the Ecua-
dorian Andes as depressing in the extreme. The lower regions, up to
ten thousand feet, are thickly covered with a forest, within which it
rains forever, the home of fever and dysentery. The higher regions
are relatively healthy, but past expression dreary, all the more or less
rounded heights being covered with coarse brown olive paramo grass,
and presenting quite a contrast to the picturesque ruggedness of the
Alps. These bleak paramos, as the tracts 10,000 feet or more above
the sea are called, are poorly populated and badly tilled, while the huts
are windowless and the people in rags. Here and there are fertile val-
leys with forest in sheltered spots. The temperature has little
variety, ranging from 39° to 46°.
Mr. Whymper describes the climate of the paramos as a perpetually
wet afternoon, and speaks of Chimborazo as a long extinct volcano,
and states that the existing centres of volcanic activity are Cotopaxi
and Sangai. The whole of the Andes of Ecuador are, according to
Dr. Wolf and Mr, Whymper, situated more to the east than they are
shown on Humboldt’s and other maps.
THe Ucayaur—The Ucayali, generally considered the leading
affluent of the Amazon, has been found to be navigable for 1040 miles. i
The junction of the Perene with the Ene forms the Tambo, and that
- of the Tambo with the Urubamba forms the Ucayali. The Perene,
which curves westward toward the Andes, can be navigated ten miles
above its junction with the Ene, while the Huallaga, a large tributary
` tothe north, can be entered by large steamers to 100 miles from its
mouth. {daiis on the Amazon, on the frontiers of Peru and Brazil,
is 350 feet above sea level, the mouth of the Ucayali 370, that of the
` Tambo 800, and that of the Ene 1000 feet. ‘The native population of
1893.] Geography and Travel. 257
the Ucayali Valley, according to Col. Church, is not more than forty
thousand, and this is as many as can be supported with their modes of
life. Life is not plentiful ; fish are scarce in the waters, and in correl-
ation with this, birds are not abundant. The natives cannot procure
iron. The eastern slopes of the Peruvian Andes are clothed with thick
forests, among which cinchona trees abound.
Asia.—Tue Pamirs.—During the last few years much has been
written about the Pamir, a region the very name of which was unknown
until recently. Many travellers have found their way across this now
celebrated plateau, or rather congeries of plateaux, which may be con-
sidered to form the western and narrower end of the great Tibetan
table-land, the knot, in fact, from which the bounding ranges of that
plateau take their origin. Among its late visitors have been the
Frenchmen Bonvalot, (in company with the Prince of Orleans ), and
Sauvergne, the Russian Grombchevysky, the Englishmen Younghusband
and St. George Littledale, accompanied by his wife. The main object
of the latter seems to have been the destruction of Ovis poli, yet they
could scarcely avoid adding to our knowledge of this not readily acces-
sible tract.
The Pamir, or rather Pamirs, for there are several, the Little Pamir,
the Great Pamir, that of Tagh-dum-bash, Alichur Pamir, and Sarikol,
consist of a series of flat valleys surrounded by mountains, the really
lofty crests of which do not rise greatly above the level of the valley
ms.
‘The Pamir is situated at the junction of three great empires, and the
ownership of the comparatively barren region may lead to war. The
lake on the Great Pamir has been named Victoria Lake, but is called
by the Kirghis,Gaz Kul, a name also given to two or three other lakes.
Victoria Lake is 13,980 feet above sea level, while the Khargash Pass
to the north is 14,500 feet, and the Andenin Pass, between it and the
Little Pamir, 15,500 feet. The Pamir River from this lake flows west
into the Wakhan, which seems also to bear the name of Kala-i-Panj,
and which, after its junction with the Murghab, becomes the Amu-
Daria or Oxus. The Little Pamir Lake is said to be 13,850 feet above
sea level, and its outlet is by the Aksu, which, after running north-
east until it has rounded the Great Pamir, unites with the Aik Bailul
to form the Murghab affluent of the Oxus, flowing through Roshan
Valley. Between the westerly courses of the Wakhan River and the
Murghab flows the Ghund-Dara, the chief sources of which seem to be
Lake Yashil-Kuhl and the Alichur River—the valley of the Ghund-
18
258 The American Naturalist. [March,
Dara is known as Shignan. Littledale’s encampment on the Alichur
Pamir, was at a height of 13,625 feet. Lake Kara Kul, north of the
Akru-Murghab, is 12,400 feet above the sea.
East of the Great and Little Pamirs are Tagh-kum-bash and Sari-
kol, the streams from which, among them the Markhan-su, flow into
the Kashgar and Yarkand Rivers, and thus geographically fall to
China, which has,in fact, raised her banner on the Sarikol plateau.
The two maps which have been published in the Proc. Roy. Geog.
Soc. during the past year exhibit differences in the course of these
atiuents.
The important range of the Hindu Kush separates the Chitral and
Hunza district from the Pamir and more northern valleys. This range
is crossed by the extraordinary depression, two to three miles wide,
known as the Baroghil Pass (12,480 feet). Through Chitral Valley
flows the Yarkhun River, rising in a small lake bearing the title of
Gaz Kul, and flowing somewhat south of west toward the Indus. Close
to this Gaz Kul, even if not occasionally united with it, is another
small lake, from which issues the Karambar River, which with the
Gilghit and Hunza Rivers, join the higher course of the Indus, where it
flows toward the west, before bursting through the Himalayas. The
western part of this Indus-draining region is known as Chitral, and is
situated at an average elevation of 5200 feet on the southern slopes of
the Hindu Kush, amid spurs from fourteen to twenty-five thousand
feet above sea level. On the south it is bounded by the petty States
of Asmar and Dir. The valley is reputed to be very fertile, and is
said to have a population of a hundred and fifty to two hundred thous-
and, without reckoning the Bushgali Kafirs. The Mehtar or Bad-
shah of Chitral, is said to be able to bring into the field 6000 fighting
men, all careful marksmen. Chitral, about 150 miles from the town
of Gilghit, is an aggregation of six large villages situated along the
river, which in the Geographical Journal is called the Kashkar. The
names Yarkhun and Kashkar seem to be identical with the Yarkana
and Kashgar of Chinese Turkistan.
Mr. and Mrs. Littledale reached the Pamir region via the Russian
‘Trans-Caspian railway. The route from Samarkand to Marghilan, the
capital of Kokhand, is described as an alternation of barrenness with
gardens of Eden. Continuing southward, they followed the Gulcha
River and crossed the Alai plateau over the Little Alai range.
‘Through the passes of Taldik (11,600), Shart (12,800), Terek (over
12,000), Kalin Art (10,800) and Kizil Art (14,200), they reached
‘Kara Kul Lake, and then, after encamping within sight of Lake
` 1893.) Geography and Travels. 259
Yashil Kul, crossed the Great and Little Pamirs, and proceeded through
Baroghil Pass to Chitral, Darkot and Yasin, whence they made their
way to Gilghit. From the Great Pamir they enjoyed a view of magni-
ficent mountain peaks over 20,000 feet high. They describe the Pamirs
as flat valleys, with lakes having low shelving banks; barren areas
with little grass. In Chitral the climate was much better, the warmth
and the abundance of apricots contrasting sharply with the barren
coldness of the Pamirs.
D. W. Freshfield thus speaks of the Pamirs, after stating that the
different strips of table-land form a district 280 miles long, from 120
to 140 wide, and twelve thousand feet high, “tent-shaped glacier-
covered mountains divided by broad easy gaps, bare heights naked of
verdure and shorn of forests by bitter winds and frosts ; desolate lakes,
a region which for the most part has neither fuel nor food; an Enga-
dine of Asia, with nine months of winter and three months of cold
weather; the home of wild sheep and that of a few wandering shep-
herds; nomads’ land if not no man’s land.” The Chinese name for the
eaid Woi the “half-way house to heaven,” while the word
“Pamir” appears to be a Turki term for a plateau. Grombchevsky
gives a more favorable account of the district. Wakhan, Shignan and
Roshan are at present claimed by the Amir of Afghanistan. Mr.
Younghusband found his way to Hunza in 1889 and to the Pamirs in
1890. Between Leh and Hunza, or Kanjut, four passes, from 17,500
to 18,500 feet high, must be crossed, amid glaciers and the grandest
scenery. The home of the robber Kanjuts, or Hunza, (in which latter
name some think to find the origin of the Huns), is eastward of Chi-
tral. The villages of the Hunza are stone-walled forts, the entrance
into which would be difficult if resisted. The sovereign, who made his
way to power in good old Oriental fashion, by the murder of his rela-
tives, owes allegiance to the Maharaja of Kashmir. The Taga-dum-
bash Pamir descends as low as to 800 feet.
Another recent traveller in this region is the Frenchman Dauvergne,
who entered by Srinagar and Leh, over the Karakoram Pass to Sanju
Kurgan, took a new route to the sources of the Oxus, crossed Baroghil
Pass into Chitral, and then through unexplored passes found his way
into Karambar Valley, and via Gilghit back to Srinagar. Beyond
Kilian Pass (17,450), at Namelong, Dauvergne turned west and reached
Kugiar Valley by the following passes, Namelong (12,140), Saraghar
(13,250), Tuslar Dawan, (14,500), Tupa Dawan (15,400), (a great
opening in a chain parallel to the Kuen Lun, not yet marked on the
maps, with peaks sixteen to nineteen thousand feet in height); Sanich
260 The American Naturalist. [March,
Dawan (16,170), and Kichokin Dawan, a double pass, 15,300 feet
above sea level. The Tisnaf Valley, Egisarak Kurgan, where copper
is found, the village of Langar on the Zarafshan, an affluent of the
Yarkand, are other points in his journey. He followed the Zarafshan
to its junction with the Tung, and then turned S. 8. W. to the plateau
region, where he reached 16,350 feet. According to him, the real
sources of the Oxus are those of the Panjah. The difficult Karambar
Pass was crossed with ten horses.
Australasia.—Tue GLACIERS OF Sourn New ZeaLrann—The
glacier region of the southern island of New Zealand has been to a great
extent explored during the last thirty years. The smaller glaciers,
which lie north of the larger, were first visited, the earliest visit to the
latter being that of Sir J. von Haast in 1862. After his exploration
of the headwaters of the Godley and Tasman rivers, came the visit of
E. P. Seely between 1867 and 1870. Little more was done until 1882,
when the Rev. W. S. Green almost ascended Mt. Cook, the loftiest peak
of the region (12,349 feet). Dr. Lendenfeldt followed next year, and
ascended Hochstetter Dome (9,258 feet). Since that year many
ascents have been made by G. E. Mannering, Dixon, Johnson, Inglis,
Brodrick and A. P. Harper, the contributor of a notice to the Royal
Geographical Society. From the account and accompanying map it
may be gathered that there is a great difference between the conditions,
character and scenery of the eastern and western parts of the range.
The glaciers of the east- slope descend gradually from elevations of
5,300 to 8,600 feet to their terminations, 2,354 to 2,882 feet above the
sea; the comparatively flat surface of their ice is hummocky, and for
about a quarter of the length of the glaciersis covered with a consider-
able quantity of rough moraine; the old lateral moraines of these
eastern glaciers are also distinct, especially that of the Hooker glacier,
where in one part there are no less than five. On the Mackenzie
plains terminal and lateral moraines have been followed for forty miles.
The glaciers of the east slope have been tolerably well explored; the
principal ice sheets are Tasman, 18 miles long, 1.25 to 2.14 wide, and
covering 13,664 acres; Murchison, 5,800 acres, and eleven miles in
length; Müller, 3,200 acres, and eight miles in length ; and Hooker,
2416 acres, and seven and a quarter miles long. North of these lie the
_ Classen glacier of 1,707 acres, 4.70 miles long, and Godley glacier, cov-
ering 5,312 acres, extending over eight lineal miles. The Aletsch
glacier of Switzerland is fifteen miles by about a mile, and thus is
inferior in dimensions to the Tasman glacier.
1893.] Geography and Travels. 261
The western slopes have not yet been thoroughly explored, but it is
known that they are subject to a very large rainfall, amounting to
about 120 inches against 25 or 30 on the eastern slope. Though Mt.
Cook is nearer to the equator than Switzerland, the glaciers descend
some 3,000 feet lower than in that country. The Fox glacier, indeed,
on the western slope, descends to 700 feet, and one or two others to less
than 1,300 feet. These western glaciers are steep ice-falls to a short
distance from their terminations, and their surfaces are almost free
from moraine stuff—in consequence of the steep dip and smooth surface
presented by the rock strata on this face, as compared with the jagged
edges on the east slope. The lower parts of the Fox glacier, and of
some others on the west, is overhung by tree-ferns and bushes growing
almost on the moraine, and near to the terminal face of the Fox issues
a hot spring with a temperature of 100°. A phenomenon of the same
kind, on a larger scale, occurs at Mount Ruapehu (9,100 feet) in the
northern island. This mountain is covered with snow and ice to the
summit, where is situated a boiling lake, into which the ice melts.
These glaciers seem on the whole to be receding, especially those at
the head of the Rangitata River, but last year the Miiller advanced so
as to dam up the Hooker River, and the Ball glacier is certainly
advancing. The daily rate of flow varies ; on the Ball it is from ten
to eighteen inches, on the Murchison from 2.6 to 8, on the Hooker very
slight, on the Miiller from 3 to 12.
In 1890 Mannering and Dixon discovered that the loftiest peak,
Mt. Cook, was not situated on the main range, but on its eastern side,
so that it sends no water to the west coast. The next most lofty peaks
are Dampier (11,823), Tasman (11,475), and a peak which bears no
title on the map, though it reaches the altitude of 11,844 feet. There
are several other summits that attain above ten thousand feet. The
writer states that he has not seen vegetation at above 6,200 feet, and
that 6,500 may be considered the highest limit. The flora is not so
varied as that of the Swiss Alps.
Europe.—Txe Cavucasus.—The Caucasus still continues to be a
favorite region for Alpine exploration by English and other clubs,
Mr. D. W. Freshfield, himself an ardent climber, from time to time
contributes a note to the Royal Geographical Society. In one of these
it is mentioned that new maps show in the central Caucasus one peak
over 15,000 feet, nine over 14,000, and five others over 13,000, also that
in Suanetia four elevations of more than sixteen thousand feet are
known, two over 15,900, three more over 15,000, and three over 14,000,
262 - The American Naturalist. [March,
The Ossete villages, built upon summits, are stated to abound in inter-
esting antiquities.
Tue Brack Sea.—A hydrographical exploration of the Black Sea,
recently conducted by the Russian Government, has proved that the :
waters of that large area exist under anomalous conditions. The 100
fathom line follows the coast outline closely save in the north-west
where it crosses with a slight northern inflexion, from Varna to the
Crimea. From this the bottom descends at the tolerably steep gradient
of twelve degrees to a depth of 500 fathoms, and from thence with a
slighter inclination to 1,200 fathoms. The 1,200 fathom line encloses
an oval in the center, with 1,219 to 1,227 fathoms as its greatest
depths. :
The upper stratum of twenty-five fathoms changes in temperature
with the seasons; between this and 100 fathoms the coldest water, as
low as 44° is found, while below 100 fathoms the constant temperature
is 48°. In August the upper surface of the cold layer sinks to from
thirty-five to fifty fathoms. The vertical circulation of the water is
limited to the upper one hundred fathoms, and there is no trace of
organic life below this level. The lower layers contain much sulphur-
etted hydrogen, besides sulphates of alkaline and earthy metals, yet
the principal reason of the absence of life is probably the deficiency of
oxygen in a dissolved state. The bottom from ten or twenty to a hun-
dred fathoms is covered with a light gray mud; from 100 to 800 fath-
oms the mud is dark gray, and at greater depths it assumes a bluish
tint through the prevalence of carbonate of lime. Remains of brackish
water mussels, of which species some still live in the Caspian, while
others inhabit the rivers flowing into the Black Sea, are found at
depths of from one to six hundred fathoms. It has been assumed that
at the beginning of the Quaternary and end of the Pliocene periods,
the Black Sea was a brackish lake, without any communication with
the Mediterranean, and that at the end of the glacial period the very
salt waters of the Mediterranean bursting in through the newly opened
Bosphorus, accumulated at the bottom, prevented circulation, and
annihilated the brackish water fauna. ` '
THE MEDITERRANEAN.—The depths of the eastern Mediterranean
have been explored by the Austrian expedition in the Pola. A depth
of 4,400 metres was found in 35° 44’ 20” N., and 21° 44’ 50” E. long.,
at a distance of only about fifty knots from Cape Matapan. A few
miles farther eastward 4,080 metres was found—these are the greatest
1893.] Geography and Travels. 263-
known depthsin the Mediterranean. Between Candia and Alexandria
3,310 metres was found twenty miles south-east of Grandes Bay, and
this depth gradually diminished to the east, to 2,120 metres near Alex-
andria. :
The Polar Regions.—Dr. Naxsex’s Voyace.—The opening
article of the Geographical Journal for this month is a reprint of the
address read before the Royal Geographical Society on November 14,
1892, by Dr. Fridtjof Nansen, the adventurous traverser of Greenland,
and the daring aspirant to the honor of reaching the North Pole by
drifting with the currents.
Of the existence of these currents ample proofs were adduced i in the
course of Dr. Nansen’s speech, but of their sufficiency, their regular-
ity, as well as of the character of the region to be traversed, whether
largely open ocean, or whether cut up into intricate channels by a
maze of islands, the bold argonaut could not say anything convincing.
As proofs of the general course of the currents, Dr. Nansen stated
that ships turned back by floe ice drifting southward were carried
between Greenland and Spitzbergen, through which passage he estima-
ted that in every twenty-four hours a startlingly enormous quantity of
water passed southward; also that other southward currents ran
through Smith, Jones and Lancaster Sounds; and that undoubted
relics of the unfortunate Jeannette were picked from a floe at Juliane-
haab three years after she sank near the New Siberian Islands. He
believes the regions around the pole to act like an enormous pump, suck-
ing in the water from Bering’s Strait and East Siberia, and returning it
by the Greenland Seas. Among other facts, he mentioned that several
years ago, a throwing-stick of a peculiar form used only by the natives
of Port Clarence, Norton Sound, Alaska, was found near Goathaab on
the west coast of Greenland, also that the driftwood which reaches every
year the shores of Greenland and Spitzbergen, is the timber of Ameri-
can and Siberian species. At the NewSiberian Islands the ice is thin,
while on the east coast of Greenland it is thick, and the speaker main-
tained that it grew in bulk as it drifted across the pole.
Specimens of mud, collected by Nansen from floes between Iceland
and Greenland, had been examined by Dr. Tornebohm, of Stockholm,
who had come to the conclusion that it consisted of mud from the great
Siberian rivers. Dr. Cleve, of Upsala, had also examined the dust from
the snow of these floes, and had identified sixteen species of diatoms,
all of which were known to be found at Cape Wankarena, near Behring
Strait, and twelve of which were only known from there. The pumice
"264 The American Naturalist. [March,
abundantly strewn on some parts of the shores of Norway, Spitzbergen
and Greenland, is stated by Bäckström to be andesitic, and must
therefore have been derived from some andesitic volcano near Bering
Sea—this proves a post glacial communication. A great deal of this
pumice is now, probably on account of the rising of the land, at from
thirty to seventy feet above sea-level.
In order to guard, as far as possible, against the danger of being
crushed in the solid ice, a ship of small size and great strength, yclept
the Fram, bas already been built. She displaces about 600 tons, is
large enough to accommodate twelve men with food and fuel for five or
six years, and has engines of 160 horse power, enabling her to steam
some six knots hourly. The sides slope from bulwarks to keel, so that
the pressure of the ice may tend to lift her out of the water (as occurred
in the case of the Tegethoff); she is broad in proportion to width,
pointed at both ends, flat-bottomed near the keel, which is almost cov-
ered by the boarding, and has a considerably curved stem. A special
point is the arrangement for raising the screw clear out of the water
—two reserve screws are to be carried. The sides are planked with
pitch-pine, oak, and greenheart to a total thickness of over twenty'eight
inches. The length at water-line is 113 feet, and the sailing speed is
reckoned at from 8 to 9 knots. The Fram, or Forward, was launched
at Laurvik, October 28, 1892, and will start on her hazardous cruise
in the Spring. She has ten boats of various kinds, including two
specially large and strong, intended for the homeward voyage in case
the ship should be crushed; a dynamo will also be taken.
The reading of this paper was followed by a long discussion, in
which various objections were brought forward by distinguished Arctic
navigators. Some of these were successfully answered by Dr. Nansen,
who, however, seemed inclined to follow the advice of Captain Wiggins,
and to start by way of the Kara Sea, rather than from the New Sibe-
rian Islands. Sir George Nares pointed out the danger of disregarding
the usual maxim to keep close to shore, and said that several years
would be required for the drifting, during the whole of which time the
vessel would be in danger. A vessel frozen-in became a solid block
with the ice. He accepted the homing current as proven, but doubted
` the existence of the out-going one, or at least its power to overcome the
force of the winds. The largest fresh-water rivers lost all influence at
a distance of 200 miles from their mouth. The Fram, he said, could
not at best expect to reach more than sixty miles north of 76° 30’
before meeting with the ice-pack, and would thus be frozen-in 730
miles from the Pole and 600 from the home current. He believed that
1893.] Geography and ` Travels. 265
the wind would be more likely to drift the vessel to the west than to
the east,
Sir A. Young said that the greatest danger arose from the land.
The remains of the Jeannette had probably found their way through
narrow channels, in the course of years. He considered the fact that
a tame reindeer had once found its way from Siberia to Upernavik,
in Greenland, proved the existence of land. The advice of Captain
Wiggins was not to wait for a squeeze, but to endeavor to run the ves-
sel on to the top of the ice. 2
To all these Dr. Nansen answered by a declaration that, as he had
abandoned his ships to cross Greenland, so he meant to abandon the
coast to reach the Pole; he believed that the winds acted with and
caused the currents.
Admiral Sir G. H. Richards, in a written communication states his
belief that the enormous ice-cap of the pole, and the weight of snow,
cause an outflow in all directions. Solid ice north of 78° is his concep-
tion of the Arctic.
Finally Sir J. D. Hooker writes a word of earnest warning. The
lines of a ship may be of use in an open pack, but not when forced
against land, amongst floes and bergs, or when the vessel is on her
beam ends. He then enlarges upon the depressed spirits produced by
the long confinement in close quarters, ever in prey to .a haunting
uncertainty, to say nothing of the risk of scurvy.
General Notes—The Geographical Journal, the first number of
which appeared in January, can scarcely be called a new magazine,
since it is but the well-known “ Proceedings of the Royal Geographical
Society ” in a new dress, a dress which will, it is hoped, tend to
popularize the study of Geography, and thus gain for itself a wider
circle of subscribers than was reached by the “ Proceedings.” One of ©
the most noticeable new features is the more extended reviews of
recently issued books, printed in the same type with the principal
articles.
Before the outbreak of the war with Chili, a Geographical Society
had been founded at Lima, by D. Manuel Pardo. This was broken
up by the war, but was again started on April 15th, 1891. The first
volume of the “ Proceedings ” contains much of interest; among other
things an article on the phenomenon known as the “ Callao Painter,”
by S. Raimondi, and others on the River Purus, the geology of
Huanta, and on the subsidence of Lake Titicaca. The president is-
266 The American Naturalist. [March,.
D. Louis Carranza, a physician and a close observer, the secretary of
the philologist, G. P. Zegarra.
Among well-known geographers who have passed away during 1892
may be mentioned Bates, the “ Naturalist on the Amazon”; Grant,
Speke’s companion to the sources of the Nile; Lord Arthur Russel),
Professor Moseley, and Sir Lewis Pelly, who, while stationed at
Bushire, adventurously penetrated in British uniform to the stronghold
of the fanatical Wahabis. This was in 1865. At Riyadh he met the
- blind Saiyid Amir, and was allowed, in appearance, to depart in safety,
but on their homeward march the party found that their water-bottles
had been poisoned, and suffered agonies of thirst, the only refreshment
they could gain being to pour the water over their wrists.
In an address delivered before the Berlin Geographical Society,
Professor J. Walther compares the desert regions of America with
those of Africa, and finds the similarity greater than is generally sup-
posed. Both are characterized by four distinct types of denudation,
gravel beds, sand-dunes, loam regions and salt deposits. In both the
mountains rise out of the plains like so many islands, without any inter-
vening debris, and in both the “amphitheatre ” formation is common.
Both exhibit the denudating powers of heat and dryness, the first split-
ting the rocks into fragments, while the dry winds whirl the dust
into heaps. No doubt water, even here, is the chief agent, but it has
not more than sixty days in which to accomplish its work. The chief
difference between the American arid lands and those of North Africa
is the steppe vegetation of the former.
1893.] Geology and Paleontology. 267
GEOLOGY AND PALEONTOLOGY.
Geology of Eastern Siberia.—The Jnvestia of the East Siberian
Geographical Society (Vol. xxiii, 3) contains an account of M. Obrut-
cheff’s further researches in the Olekma and Vitim highlands. In
the northeastern part of this region the author found a further contin-
uation of the “ Patom plateau ”—that is, a swelling from 3500 to 4000
ft. high, devoid of trees, with ridges and mountains rising over it to
heights of from 5000 to 5600 feet. They consist of granite and crys-
talline schists, probably of Laurentian age, covered with younger,
probably Huronian, gneisses and schists. The other parts of the
highlands consist of Cambrian and Ordovician deposits, while
Silurian limestones and Devonian Red sandstones are met with in
the Valley of the Lena. We thus have a further confirmation of
the hypothesis, according to which the great plateau of northeastern
Asia is a remnant of an old continent which has not been submerged
since the Devonian epoch. Further traces of mighty glaciation have
been found in the southeastern part of the region. As to the gold-
bearing deposits, they are pre-glacial in the south and post- glacial in
the north. The high terraces in the valleys are indicative of a con-
siderable post-pliocene accumulation of alluvial deposits, and of a sub-
sequent denudation on a large scale. (Nature, Jan. 12, 1893).
Geological Features of Arabia Petræa and Palestine.—
At a recent meeting of the London Geological Society, the following
communication was read by Professor Edward Hull on the Geology of
Arabia Petra and Palestine :
The most ancient rocks (Archean) are found in the southern por-
tion of the region ; they consist of gneissose and schistose masses and
are penetrated by numerous intrusive igneous rocks. They are suc-
ceeded by the Lower Carboniferous beds of the Sinaitic peninsula and
Moabite tableland consisting of bluish limestone with fossils, which
have their counterparts chiefly in the Carboniferous limestone of Bel-
gium, and of a purple and reddish sandstone (called by the author
“the Desert Sandstone,” to distinguish it from the Nubian Sandstone
of Cretaceous age), lying below the limestone. The Nubian Sand-
stone, separated from the Carboniferous by an enormous hiatus in the
succession of the formations, is probably of Neocomian or Cenomanian
age, and is succeeded by white and gray marls, and limestones with
268 The American Naturalist. [March,
flint, with fossils of Huronian and Devonian ages. The Middle Eocene
(Nummulitic Limestone) beds appear to follow on those of Cretaceous
age without a discordance, but there is a real hiatus notwithstanding
the apparent conformity, as shown by the complete change of fauna.
In Philistia a calcareous sandstone in which no fossils have been dis-
covered is referred to the Upper Eocene; for the Miocene period was
a continental one, when faulting and sliding were taking place, and the
main physical features were developed—e. g., the formation of the
Jordan-Arabah depression is referable to this period. i
In Pliocene times a general depression of land took place to about
200-300 feet below the present. sea-level, and littoral deposits were
formed on the coasts and in the valleys. To this period belong the
higher terraces of the Jordan-Arabah valley. The Pliocene deposits
consist of shelly gravels. Later terraces were formed at the epoch of
the glaciation of the Lebanon Mountains, when the rainfall was exces-
sive in Palestine and Arabia.
The volcanoes of the Jaulân, Hauran, and Arabian Desert are con-
sidered to have been in active operation during the Miocene, Pliocene,
and Plistocene periods, but the date of their final extinction has not
been satisfactorily determined. (Geol. Mag., Jan., 1893).
The Vertebrate Fauna of the Ordovician of Colorado.—
Mr. Charles D. Walcott has recently published a paper on the verte-
brate fauna in strata of Ordovician age near Canyon City, Colorado,
already noticed in Tae Narurauist. The fossils consist of what
appear to be the plates and scales of fishes and the ossified chordal
sheath of a fish allied to the recent Chimera. The remains occur in
a sandstone which is correlated with the lower Trenton and the lower
Bala of Wales. Microscopic sections of a dermal plate belonging to
an Asterolepis-like form, examined by Dr. Otto Jaekel, show (1) the
dentine tubules that are characteristic of vertebrates ; (2) the occurrence
of true osteoblasts, which exclude the forms from the Elasmobranchii,
and relegates them to other low divisions of the fishes ; (3) the absence
of enamel and the distinct concentric lamination of the dentine tubules
which indicate a low stage of development.
In view of the objections that can be made to -a classification based
entirely upon the characters of the dermal plates and scales Mr. Wal-
cott has made his classification tentative and has only outlined the
characters of the fragmentary remains. For the present, then, the
fossils will be known respectively as Dictyorhabdus priseus, a supposed
chimaeroid, and Astraspis desideratum and Eriptyehus americanus,
1893.] Geology and Paleontology. 269
rhipidopterygians. It is, however, extremely unlikely that these forms
are fishes, but they are more likely Agnatha.—Bull. Geol. Soc. Am.,
Vol. 3, pp. 153-172. C
The Loess in Southern Russia.—In his notes on Russian
Geology Mr. W. F. Hume gives an interesting account of the Russian
Loess, its character and distribution, together with a theory of its
origin
The Russian Loess is a yellowish-brown sandy clay, often rich in
grains of quartz and mica, and in many places rich in carbonate of
lime and humus. In lies unconformably on all the principal forma-
tions. To the west of the Dnieper it conceals the broken and contorted
gneisses and granites of the Archean axis in S. Ekaterinoslav and the
Don Cossack country it covers the shales and sandstones of the Carbon-
iferous, whilst in the more central governments of Kursk, Karkoff,
and Tchernigov it overlies the Cretaceous and the whole Tertiary
series. Also along a definite line running to the north of these gov-
ernments it rests upon the Boulder-clays and sands of the Glacial
period. From its general appearance Mr. Hume considers the Loess
of Russia to be more or less coeval with that of Central and Western
Europe, and the paleontological evidence seems to confirm his view.
In discussing the origin of the Loess Mr. Hume calls attention to
the relation, pointed out by Professor Suess between the Loess and the
Glacial Drift, and gives in detail the theories of Professor Armache-
vsky and Baron von Richthofen, neither of which is sufficient in itself
to account for the distribution of the Loess
In conclusion the author gives the following statement of the proba-
cle sequence of events:
1. The Loess particles may be originally derived from the finely
ground material resulting from the wearing of the subjacent beds by
the ice-sheet.
II. The same have been deposited in tundra-like depressions under
the influence of slowly moving waters or by the action of rivers in
flood.
` IIL. This deposit under more temperate conditions dried up, and
was then suitable material for the redistributive action of the wind.—
(Geol. Mag., Dec., 1892).
Sources of the Texas Drift.—Mr. Dumble divides the Texas
drift area into four districts. First the Trans-Pecos Texas, the valley
of the Rio Grande, and the Rio Grande Divide. The origin of this
270 ‘The American Naturalist. [March,
drift is traceable to the mountainous region of Trans-Pecos Texas,
where nearly every variety of pebble can be found in its original loca-
tion. Second, the country between the Nueces and the Brazos. This
region is covered with pebbles, gravel and sand derived from the rocks
found in the Central mineral region. Third, from the Brazos to the
Sabine, where the gravel is largely made up of ferruginous material
which had its origin in the iron-capped hills which cover so large a
portion of Eastern Texas. Fourth, the area known as Northwest
‘Texas according to the survey division. The drift of this region came
from the hills enclosing the waters under which they were deposited,
viz., Wichita Mountains and the mountains of New Mexico.—Trans.
‘Texas Acad. Sci., Vol. i, 1892.
Archean.—According to Mr. H. V. Winchell, Minnesota possesses
the greatest iron district known in the world to-day. It lies in the
Mesabi range and extends from the Canadian boundary line in a direc-
tion a little south of west, beyond the Mississippi River, a distance of
140 miles. The ore on the eastern end is hard, black and magnetite,
owing to the heat of the gabbro overflow. In the central and western
portions of the range the ore is soft hematite, limonite and geothite.
As to the source of the iron, it is believed by Mr. Winchell to be
largely the result of oceanic deposition, both chemical and mechan-
ical, and to have been concentrated in its present situations. (Twentieth
Ann. Rept. Minn. Geol. Surv., 1892).—Mr. T. R. Struthers advances
the theory that the primitive rocks, i. e., granite, were formed by the
cooling of the exterior of the globe under the primeval deep. The
pressure of the sea at the depth of two miles would be sufficient to
account for the structure of granite. This theory explains the fea-
tures presented by the bedded or stratified granite of the British
islands and many other parts of the world. (Geol. Mag., Dec., 1892.)
Paleozoic.—A skull of Dinichthys intermedius recently examined
by Professor E. W. Claypole supplies details previously unknown
regarding the plates of which it is composed. The especial points of
interest are the forms of some of the plates and the over and underlap
- which has not been represented, and sore additions to the structure of
the upper jaw. (Am. Geol., Oct., 1892.)—Mr. Herbert Bolton reports
a triolobite from the Skiddaw slates of the Isle of Man. The specimen
belongs to one of two genera, Asaphus or Æyglina, both of which are
Ordovician forms. This in connection with the occurrence of two
specimens of Palzeochorda (an Arenig form) is strong evidence that
1893.] Geology and Paleontology. 271
the Skiddaw slates are of Llandeilo age. (Geol. Mag., Jan., 1895).—
A species of Cyclus from the Coal Measures of Lancashire, England,
is described and figured in the Geol. Mag., Jan., 1893, by Dr. Wood-
ward. The fossil is about as large as a shilling and resembles C. agno-
tus H. von Meyer. Mr. Woodward considers it a new species, and
names it Cyclus scottii.
Mesozoic.—A new crustacean, Prosopon etheridgei, from the Cre-
taceous beds of Queensland is described by Dr. Henry Woodward.
This crustacean is closely related to P. verrucosum Reuss, and P. tubero-
sum, von Meyer, two Neocomian species from the Cretaceous of Boucher-
aus, Dept. Jura. It differs from both, however, in several important
points. (Proceeds. Linn. Soc. N. S. W., Vol. vii, 1892).—Mr. R.
Etheridge reports a new fossil Phyllopod from the Upper Coal Meas-
ures of the Newcastle District, N.S. W., belonging to the genus Leaia.
This is the second genus of that family known from the whole of
Australia. Mr. Etheridge has named the species Leaia mitchellii in
honor of its discoverer, Mr. Mitchell. (Proceeds. Linn. Soc. N. S
Wales, Vol. vii, 1892).
Cenozoic.—Mr. J. H. Cooke reports finding the jaw of an Arctic
bear, Ursus arctosin pleistocene strata of Malta. The fossil, consist-
ing of an entire ramus with its canine and molar teeth, was found in
a cavern together with bones of elephants, hippopotami, a stag, and a
large dog. (Knowledge, Dec. 1, 1892).—A ccording to Baron DeGeer
the Pleistocene changes of level in North America as well as in
Europe, are closely connected with the local structure of the earth’s
crust and with the local extension of the glaciations, and that these
changes cannot be accounted for by changes in the level of the sea.
(Amer. Geol., Jan., 1893).—M. Bureau has described two fossil plants
from the Caleaire grossier parisien. The first, called by the author
_Aralia eocenica, is represented by an impression of a leaf remarkable
` for the slender, long petiole. The second, Monochoria parisiensis,
resembles strongly certain species of living Monochoria found in India,
Ceylon, Malay, Chinaand Japan. (Revue Scientifique, Jan., 1893).
bo
~J
by
The American Naturalist. [March,
MINERALOGY AND PETROGRAPHY:'
The Rocks of the Thalhorn.—In the Thalhorn of the Upper
Amariner Thal are found a porphyritic granite, between conglomerates
composed of gabbro pebbles in a schistose matrix, and also serpen-
tines, massive gabbro, schists, and various contact rocks. Linck? gives
a good petrographical description of all these, and geological notes of
their occurrence. e main granite mass is a portion of the well-
known Kamm granite. It is found in dykes and flows, and it varies
in its composition and structure from a typical granitite containing
two feldspars, through porphyritic granite and syenite to lamprophyric
minettes. The unaltered sediments near the eruptive are graywackes.
On the contact with the granite the clastics are altered to knotty schists
that are predominantly biotite schists flecked with light spots, consist-
ing mainly of quartz and feldspar in micropegmatitic intergrowths,
surrounded by biotite. Extreme alteration gives rise to hornstones, of
which the writer recognizes several varieties. In these biotite, feld-
spar, hornblende and micropegmatite are so orientated as to resemble
the poicilitic structure of many diabases and other basic rocks. Horn-
blende is abundant in them as needles scattered through the ground-
mass and as large phenocrysts. The conglomerates occupy the greater
share of the writer’s attention. In one group acid pebbles occur in 4
sandy or clayey matrix of basic detritus, in which biotite, feldspar and
hornblende are new products of alteration. A second group includes
rocks made up partly of gabbro material. Here the author again
recognizes two groups, in one of which diallage and other gabbro con-
stituents are occasionally present in the groundmass, and a second in
which gabbro material forms a very large portion, either of the matrix
or of the pebbly portion of the rock. In either case the rock is much
altered, with the resulting formation of plagioclase and hornblende.
The serpentine of the region was originally an olivine-enstatite rock
and not a gabbro as has been supposed.
The New Jersey Eleolite-Syenite.—The New Jersey Eleolite-
syenite dyke described by Emerson’ is again studied by Kemp,’ who
' 1Edited by Dr. W. S. Bayley, Colby University, Waterville, Me.
?Mitth. d. geol. Landesanst v. Elsass-Loth., iv, 1892.
5Amer. Jour. Science, iii, xxiii, p. 302.
*Trans. N. Y. Acad. Sci., Vol. xi, p. 60.
PLATE IIl.
1. Cecidomyia holotricha O.S. | 3. Cecidomyia persicoides O. Ss
4. Cecidomyia tubicola O. S.
{ų 2. Cecidomyia caryæcola O. S.
ee e TOET OR
gi,
33.84
1893.] Mineralogy and Petrography. 273
declares that the earlier description applies only to that phase of the
rock occurring in the northern and the southern portions of its extent.
The pyroxene throughout the dyke is aegerine. Cancrinite and soda-
lite are both fairly abundant in it. An analysis of specimens collected
from about the point visited by Emerson gave:
SiO, AlO, FeO; MnO CaO MgO K,O Na,O Loss Total
50.36 19.34 6.94 .41 3.43 notdet. 7.17 7.64 3.51=99.80
Eleolite porphyries with a tinguaitic groundmass are closely asso-
ciated with the more abundant syenite, and along the eastern side of
the great dyke are smaller ones of ouachitite and fourchite. The basic
material of these small dykes, when first’ studied, was regarded as
porphyrite. Contact effects produced by the intrusion of the syenite
through the surrounding shales are noticed on the east side of the
dyke, where the sedimentaries have been changed to biotitic hornfels.
Mica Peridotite from Kentucky.—A mica peridotite’ from a
dyke in Crittenden Co., Ky., is composed essentially of biotite, serpen-
tine, and perofskite, with smaller proportions of apatite, muscovite,
magnetite, chlorite, calcite, and other secondary products. The bio-
tite and serpentine constitute about 75% of the entire rock. The mica
is in large plates in which are scattered the grains and shreds of Ser-
pentine. The composition of the rock follows:
TiO, Al,O, Fe,0, FeO CaO MgO K,O NaO H,O P,O;
‘3.78 588 7.04 516 946 22.96 2.04 38 7.50 .89
and small quantities of Cr,O,, MnO, NiO, CoO, BaO and CI. The rock
represents a new type of peridotite in which biotite takes the part of
an amphiboloid in the more usual types.
Rhyolites in Maryland and Penn.—G. H. Williams’ has iden-
tified an extensive series of old volcanic rocks in the South Mountain
region of Pennsylvania and Maryland. The rocks have hitherto been
considered sedimentaries, but to the writer they exhibit all the pecu-
liarities of eruptives, though some of the beds are fragmental tufas
and breccias. The two principal types are rhyolite and basalt. The
former possesses all the features of recent eruptives, such as flowage
5Amer. Jour. Sci., III, xxxviii, p. 130.
6J, S. Diller, Amer. Jour. Sci., xliv, 1892, p. 286.
TIb., xliv, 1892, p. 482.
19
274 The American Naturalist. [March
lines, spherulites, lithophysae and amygdaloidal cavities. Quartz and
an alkaline feldspar are the prevailing phenocrysts, while the ground-
mass is a quartz-feldspar mosaic. The basalts are much altered, but
their structure is clearly that of an eruptive. A detailed account of
the rocks is promised later.
The Nepheline and Leucite Rocks of Brazil.—A more
careful study of a few of the Brazilian nepheline and leucite rocks
undertaken by Hussak* has resulted in the discovery of leucite in some
of the phonolites, and in the detection of leucite-tephrites containing
pseudo-crystals. The leucitophyres consist of phenocrysts of sanidine,
augite, nepheline and pseudo-leucites in a groundmass of small zeoli-
tized leucites, augite, magnetite and nepheline. The leucite-tephrites
are all characterized by the possession of the pseudo-leucites. In many
eases these are nothing but spherical masses of the rock material sur-
rounded by biotite plates. In other cases the biotite surrounds anal-
cite or mixtures of analcite and calcite. The structure of several of
these rocks is the diabasic. With these the author would place a rock
described by Eigel’ from the Cape Verde Islands, and the augite-por-
phyrite described by Kemp" from Deckertown, N. J.,in both of which
traces of leucite are thought to have been discovered. Hussak has
also found a leucitite dyke in phonolite near Pocos de Caldas, and a
leucitite tufa composed of fragments of basalt, isolated crystals of
leucite changed to analcite, pieces of augite and crystals of magnetite.
The author concludes his paper with remarks on ‘ pseudo-erystals’
combating the view of Derby that they are true leucite erystals filled
with inclusions of the rock’s groundmass.
The last named writer" has examined the Peak of Tingua with some
care, finding eleolite-syenite, phonolite and dykes of basic rocks. The
syenite and phonolite are thought to be phases of the same magma, as
they apparently grade into one another. The phonolitic phase occurs
both in dykes and in flows associated with phonolite tufas. The origin
of the pseudo-crystals is discussed briefly.
| Petrographical News.—Brauns” has discovered hauyne in the
pumice sandstone near Marburg, a mineral hitherto unobserved in the
ŝNeues, Jahrb. f. Min., etc., 1892, II, p. 141.
PAMERICAN Naturatst, Feb., 1892, p. 165.
VSee above under ‘The New Jersey Eleolite-Syenite.’
"Quart. Jour. Geol. Soc., May, 1891, p. 251.
*Zeits. d. deutsch. geol. Gesell., xliv, 1892, p 149."
1893.] Mineralogy and Petrography. 275
rock because of the loss of its characteristic blue color through alter-
ation. The list of minerals common to this rock and to those of the
Laacher See is now complete, so that the belief in a common origin
for them is rendered almost a certainty.
C. W. Hall” gives a few notes on rocks collected from Central Wis-
consin, describing very briefly hypersthene and quartz gabbros in
which there is much secondary hornblende, and quartz diorites and
gneisses regarded as squeezed gabbros.
A fourchite boulder in which are large arfvedsonite phenocrysts is
mentioned by Kemp” as occurring at Aurora, Cayuga Co., N. Y. The
same author mentions the existence of rhyolite, hypersthene, andesite
and andalusite-hornstone from near Gold Hill, Teole Co., Utah.
Spherulites® of andalusite occur in the carboniferous clastic schists
of Beaujeu, France. The schists are composed of black and white
mica fragments in a paste of sericite and hematite.
Turner” makes brief mention of basaltic, andesitic and rhyolitic
lavas, whose source was the late Tertiary cone Mt. Ingalls, in Califor-
nia.
Crystallographic Study of Diopsides.—Some very careful
crystallographic observations have been made by A. Schmidt" upon the
diopsides of the Alathal, of Achmatowsk, of Nordmark, of the Ziller-
thal and the Arany-Berg. Many crystals from each -of these famous
localities were examined, and much new data was obtained concerning
the mineral. The following new planes were discovered: 4P2 and
5P§ on the white diopside from Achmatowsk; Pœ on the green
variety from the same place; œ P6' in the Nordmark species; 2 P10,
oo P4 and oo Pz on the nearly colorless small crystals from Schwarzen-
stein in the Zillerthal, and œ P7 and P4 on the black Arany-Berg
mineral. The form P4 appears in Goldschmidt’s ‘ Index,’ but no ref-
erence to it could be found by the author in the original memoirs.
The axial ratios of the different varieties are :
Alathal 1.0895 : 1: 5894 8 = 74°15'47”
Achmatowsk (white)......... ........ 1.0909 : 1 : 5899 6 = 74°10'42”
Achmatowsk (green)...............00: 1.0951 : 1 : 5985 2 = 73°31'8”
Minn. Ac. Nat. Science, III, No. 2, p. 251.
“Trans. N. Y. Acad. Sci., xi, p. 92.
Lévy. Bull. Soc. Franç d. Min., xv, 1892, p. 121.
16Amer. Jour. Sci., Dec., 1892, p. 455.
MZeits. f. Kryst., xxi, 1892, p. 1.
276 . The American Naturalist. {March,
Nordmark 1.0915: 1: .5848 7 = 74°38'59”
Zillerthal (colorless) 1.0922 : 1: 5887 8 = 74°16/28"
Arany-Berg (yellow) 1.0945 : 1: .5918 6 = 74°19'38”
Arany-Berg (black)........... ..... anO Ts 6876. P = TA° 4687
The optical angle for the Nordmark crystals is 2Vna = 60°44’, and
C/\e¢ = 45°21’. For the dark Zillerthal diopside 2Vna = 58°56’
and C/\c = 34°4.
Herderite from Hebron, Maine.—A single specimen of Her-
derite from Hebron, Maine, is described by Wells and Penfield” as a
few yellowish white crystals on albite. The crystals have a tabular
habit, with oP, æ P, 3P and £P the only forms observed. The density
is 2.975 and composition :
P,O, BeO CaO (by diff) H,O F Insol. Total
40.81-15.32 32.54 5.83 .40 5.27 = 100.17
Corresponding to Ca Be (OH) PO, or aherderite in which nearly all
of the fluorine is replaced by hydroxy].
Mineralogical Notes—A calcium carbonate of secondary origin.
from the Marble Mountains of Wolmsdorf in Glatz has been analyzed ,
by Kosmann” with the following astounding result: Ca CO, = 4.32;
chemically combined H,O = 1.54; mechanically combined H,O =
94.13. The author believes the mineral to be a hydrated carbonate
CaCO, + 2H,0 capable of absorbing a large quantity of water, simi-
lar to the ‘ Mountain Milk’ of Rose.
The friedelite of the Manganese mine of Sjogrube, Orebro, Sweden,
occurs in large quantity in clefts, veins, etc., that are partially filled |
with calcite. An analysis yielded Igelström :”
SiO, Cl MnO FeO CaO MgO NaO H,O Total
34.36 3.00 45.88 1.35 150 150 2.79 9,00 = 99.38
On the Azurite from the Laurion Mts., Greece, Zimanyi” has found
28 forms, three of which (?Pa, $Pæ and $Pæ ) are new. The crys-
tals have the usual habit of the mineral, and they compare favorably
in beauty with those from Chessy, Arizona and Utah.
Amer, Jour. Sci., xliv, 1892, p. 114. |
WZeits. d. deutsch. geol. Ges., xliv, 1892, p. 155.
*Zeits. f. Kryst., xxi, p. 92.
“Ib. xxi, p- 86.
ear aana e e aa oe eae eae
a sas cia ee i
ee
1893,] Mineralogy and Petrography. 277
Tremolite” pseudomorphs after sahlite from the limestone of Canaan,
‘ Ct., have the composition :
SiO, AlO, FeO, FeO CaO MgO K,O NaO
60.98 “10 12 “19 14649362 18 21
The controversy over the nature of Melanophlogite is not yet ended.
Bombicci” has recently defended himself against the attack of Friedel,
and in his defense he accuses his opponent with misquoting him.
New Minerals.—Ganophyllite, from Harstige, near Pajsberg,
Sweden, is a manganese zeolite™ that is associated with barite, lead,
and rhodonite. It occurs in large brown monoclinic, prismatic crys-
tals, in which æ P is combined with the base and theclinodome. a: b
: è = .413 : 1 : 1.831. / = 86°39’. On cleavage plates parallel to oP
a percussion figure may be produced, one of whose rays is parallel to a
- and the other two inclined at 60° to this. Plane of optical axes is
perpendicular to 0 Pæ , with c the first bisectrix.. 2E (air) = 41°53’
for sodium light, and 2V = 23°52’. The pleochroism.is strong ¢ = A
= yellow-brown; a = B and 6 = C = colorless. The density is
2.84 and hardness = 4. A mean of two analyses gave Hamberg a
result that may be represented by 8Si0,'Al,0, 7MnO + 6H,O/
SiO, AlO, FeO, MnO CaO uso PbO?) K,O NaO H,O Total
39.67 7.95 .90 35.15 111 2 20. 270 218 919 = 99.85
Pyrophanite, described by the same author as occurring in the same
mine, is a manganese titanium compound isomorphous with ilmenite.
_ An analysis gave:
SiO, TiO, MnO Fe,O, Sb,O, Total
1.58 50.49 46.92 1.16 .48= 100.63
It is found as brilliant, deep red, transparent tables, associated with
ganophyllite. a:e¢— 1: 1.369. The double refraction is strong, and
the indices of refraction for sodium light are w = 2.481, € = 2.21.
Density is 4.537.
Synthesis of the Members of the Sodalite Group.—The
minerals of the sodalite group have been manufactured by Morozie-
2W.H. Hobbs. Amer. Geol., July, 1892, p. 44.
Bull. d. 1. Soc. Franç d. Min., xv, p. 144.
“Ref. Neues. Jahrb. f. Min., etc., 1892, II, p. 234.
278 The American Naturalist. [March,
wies” as microscopic crystals, A mixture of 65 parts SiO, + 3Aq,
44 parts Al,O, + 3Aq, and 33 parts gypsum, heated in a platinum
crucible with an excess of Glauber’s salt, yielded tiny cubes and dode-
cahedra of hauyne or sodalite. When heated with an excess of Na,
SO, + Na Cl a substance was obtained that is supposed to be an iso-
morphous mixture of the two minerals above mentioned, and in addi-
tion some sodalite crystals were produced. When heated with Na Cl
alone sodalite only resulted.
Methods and Instruments.—A simple method for determining
the value of the optical angle in thin sections of minerals is described
by Lane.” It consists essentially of the measurement of the angular
distance between the hyperbolas of the biaxial interference figure by
means of the sub-staye mirror.
A cheap form of crystal refractometer constructed on the same prin-
ciples as the larger Zeiss instrument has been made by Czapske.” The
height of the complete instrument is only 25 cm. It is suitable for
all ordinary refraction work.
` An Appendix to the ‘‘Gems of North America.’’—Mr.
Kunz has issued an appendix to his valuable ‘Gems and Precious
Stones of North America’™ that brings the volume up to date. Most
of the material in the appended chapter has appeared in the journals,
but some of the information it contains is new. The author states that
the sapphire gravels of Ruby Bar, Montana, and the turquoise mines
of New Mexico are now being worked by companies that expect their
outlay of capital justified by a goodly yield of gem material. The
turquoise company has already taken from their diggings about a
hundred thousand dollars worth of gems.
Neues. Jahrb. f. Min., etc., 1892, II, p. 139.
6Science, Dec. 23, 1892, p. 354.
2™Neues. Jahrb, f. Min., etc., 1892, I, p. 209.
**Cf., AMER. NATURALIST, Dec,, 1891, p. 1119.
1893.] Botany. 279
BOTANY.
An International Botanical Congress.—After careful cousid-
eration of all the conditions it has been thought advisable to take
steps toward securing an International Congress of Botanists in con-
nection with the meeting of the American Association for the Advance-
ment of Science in Madison next August. Upon the return of
Professor Underwood from Genoa with his report of what was done
there, as well as of what was left undone, such a Congress seemed a
necessity, especially when it was learned that the delegates to the
Genoa Congress expected one to be held in America this year in order
to complete the work left by them. Moreover, the Columbian Expo-
sition will doubtless bring many foreign botanists to this country
during the year, and most of these will attend our scientific meet-
ings whenever it is possible to do so. It seems wise, therefore, to
take advantage of these favorable conditions and to arrange for a
formal Congress.
After a good deal of consultation on the part of those who could
more readily do so, it was suggested that the Chairman of the Sec-
tion of Botany of the American Association for the Advancement of
Science and the President of the Botanical Club (Dr. Wilson) should
appoint a committee to take the matter in hand. In accordance with
this suggestion notices were sent on Dec. 9 to the following gentle-
men with the request that they serve on such committee: J. ©. Arthur,
L. H. Bailey, N. L. Britton, D. H. Campbell, J. M. Coulter, B. T.
Galloway, Conway MacMillan, B. L. Robinson, William Trelease, L.
M. Underwood, George Vasey.
It is to be hoped that the heartiest support will be given to the com-
mittee in their effort to bring to a successful issue the purpose for
which they were appointed.—CHARLES E. Bessey, Chairman of Sec-
tion G (Botany), A. A. A. 8.
Botanical Notes.—Mr. B. M. Davis, in the December number of
the “ Annals of Botany” describes and figures all the stages of devel-
opment from the carpospore to the young growing plant of Champia
parvula, one of the common red seaweeds of both the Atlantic and the
Pacific coasts. It is a valuable contribution to the embryology of the
Floridex.—A. P. Morgan describes in the “ Journal of the Cincinnati
Society of Natural History (October) a singular new fungus of the
280 The American Naturalist. {Mareh,
family Phallaceæ, for which he proposes the generic name Phallogaster. a
It appears to connect the Phallaceæ with the Lycoperdacez ; in fact it 1
is difficult to say why it may not be placed near the Puff-Balls rather a
than near the Stink-Horns. The single species is P. saccatus. Speci-
mens have been collected in Ohio, New York and Connecticut, show- —
ing that it is not local in its distribution. —In the “ Contributions from —
the U.S. National Herbarium,” issued in December, 1892, J. M. Holz- —
inger publishes lists of the plants collected by C. S. Sheldon and M. A.
Carlton in the Indian. Territory in 1891. The novelties are, a variety
( fasciculata) of Solidago missouriensis; a species of trailing morning
glory (Ipomea carletoni) with narrowly lanceolate leaves and large
flowers (2 to 2) inches long) which are solitary or occasionally in twos
- or threes; and a new Euphorbia (E. strictior), with very narrow leaves.
- Dr. Engelmann’s E. polyphylla, the description of which has hitherto
not been published, is here characterized and distinguished from £.
wrightii, E. strictior and E. discoidalis—In the same publication Mr.
. Carleton publishes some useful “Observations on the Native Plants of
. Oklahoma Territory and Adjacent Districts.” His observations upon
the native grasses are especially valuable.—Mr. B. B. Smyth, of Topeka,
| Kansas, has published a useful “Check-list of the Plants of Kansas,”
with especial reference to his proposed distribution of botanical speci-
-~ mens.—The Contributions to American Botany from the Herbarium
of Harvard University which the writings. of Gray and Watson have
: made familiar to botanists throughout the world, have been resumed
- by Dr. B. L. Robinson, the Curator of the Gray Herbarium. His
. Jatest.contribution consists of Descriptions of New Plants Collected in
. Mexico by C..C. Pringle in 1890 and 1891, with. notes upon afew other
. species.. Among the more notable things is a new genus of Umbel-
liferæ to which he gives the name Coulterophytum, which suggests that
the author has taken the hint given by Otto Kuntze in regard to the —
_ manufacture of names! .. Geissolepis, a new. genus of Composite is —
- represented by a single species of prostrate plants from San Luis
Potosi—Dr. Britton, in the Transactions of the N. Y. Academy of
Sciences (Nov., 1892), discusses “ Ranunculus repens and its Eastern
North American Allies.” He recognizes six species as follows: 1. B.
repens L., sparingly introduced from Europe; 2. R. macounit Hook,
Canada and in the Rocky Mountains of U.S.; 3. R, hispidus Michx.,
Ontario to Georgia and west to Michigan, Northwest Territory and
, apparently to Texas; 4. R. fascicularis Muhl., widely distributed ; 5.
R. septentrionalis Poir., eastern Canada to Minnesota, south to Pa. and
_ Ky; 6. R. palmatus Ell., South Carolina, Ga: to Fla.—In a “ Prelim-
1893.] Botany. 281
inary List of American Species of Polygonum” in the Torrey Bulletin _
for December Mr. John K. Small enumerates seventy-nine species.
Some changes are made in the nomenclature, and two new species (P.
mexicanum and P. pringlei are described from San Luis Potosi, Mex-
ico.)—* Amherst Trees,” by Professor J. E. Humphrey, and “The
“Woody Plants of Manhattan in their Winter condition,” by Professor
A. S. Hitchcock, are two pamphlets: which indicate the increasing
interest in forest trees as constituents of the flora of a locality. The
first named is the more popular and treats of many New England
trees from the standpoint of the tree lover and the landscape gardener ;
the second is quite scientific, and is intended to aid the people of the
plains (Kansas) to identify trees in their winter state-——A. W. Bennett
has published in the “St. Thomas Hospital Reports ” (London) a use-
ful paper entitled “ Vegetable Growths as Evidence of the Purity or
Impurity of Water.” He discusses the subject under four heads, as
follows: I. Flowering Plants; IJ. Fungi; IIL Alge ; and IV, Char-
aces, The presence of the first is “a sign of comparative purity of
the water;” of the second of the impurity of the water. The blue-
green algæ (Cyanophycew) “should be regarded as rendering it (the
water) unfit for domestic purposes.” . The chlorophyll-green algæ
( Chlorophycee) are probably innoxious, in spite of the prevalent opin-
ion to the contrary. The Characee are regarded as noxious “since
when decaying they give off a strong fetid odor, accompanied by evo-
lution of sulphuretted hydrogen gas.”
282 The American Naturalist. [Mareh,
ZOOLOGY.
Allen’s Faunal Areas of North America.—In a paper on
the Distribution of North American Mammals, Mr. J. A. Allen gives
the following tabular synopsis of the faunal areas of North America:
Realms {xo North E Papiiblen
American Tropical.
North American. < North Temperate Realm.
re ge | Cenal A | American.) _ american Tropical Realm.
Subregions { Wand Na \ ==North American Region.
Provinces { ey } ==Warm Temperate Region.
Appalachian. : :
; Austroriparian. l —Humid Province.
Subprovinces Ci yi
| Sonoran. 5 } =aria Province.
Great Plains.
| Grea Basin. } —campestian Subprovince,
Pacific Coast
( Barren Ground, X
Alaskan-Arctic. \ Arctic.
Districts
Canadian | Cold Temperate.
Faune 4 Sitkan.
hanian.
Carolinian, } Humid Warm Temperate
Louisianian,
Floridian, :
| Tamaulipan. } Tropical,
(Bull. Am. Mus. Nat. Hist., Dec., 1892.)
The Madagascar Fauna.—At a recent meeting of the Royal
Geographical Society, Canon Tristram made the following remarks oD
the Fauna of Madagascar :
“Madagascar has an extraordinary natural history. One would
suppose, from its position, that this would be African, but it, like its
people, is thoroughly un-African. The monkeys and lemurs of Mada-
gasear are not to be found in Africa, while all the great African ani-
mals of prey are absent. Among the lemurs is one known as the
ayeaye, the formation of whose digits is unique. The botany is almost 4
as peculiar.
1893.] Zoology. 283
“ We saw, at the last meeting of the Zoological Society, a specimen
of the egg of an extinct bird of Madagascar, which is fifteen times the
bulk of an ostrich egg, and yet the bird itself does not appear to have
been larger than—as far as we can judge from remains—the New Zea-
land moa, an extinct bird, to which it had an affinity. This same
peculiarity runs through all the birds of Madagascar. Of course, the
water-birds and sea-fowl are the same as those of Africa, but there are
one or two extraordinary exceptions. There is the snake-bird, a long-
necked bird of very great beauty and grace, allied to the cormorant,
which it resembles in its habits, and of which there are four species in
the world—the Madagascar one is certainly Indian. Then, again,
another puzzling bird to naturalists is the Mesites, a water-hen peculiar
to Madagascar. These birds are usually distinguished by a small tail
and a short tarsus, whereas, the Madagascar, which is related to
the others, has a long tail and tarsus, and no one, until M. Audebert,
thought the bird was allied to the rails. There is a group of cuckoos
entirely peculiar to Madagascar—the coua—of which there are nine or
ten species, which have no relations at all in Africa or India. Then,
in another group, we have a bird allied to the thrushes, but not Afri-
can, although allied to a species in the Mauritius and al] the Mascarene
Islands—the Hypsipetes. Altogether, we cannot explain the Madagas-
car Fauna, but it shows that Madagascar must have been separated
from Africa for an infinity of ages; and its natural history affinities
are certainly rather with India than Africa, and yet they are entirely
distinct and peculiar. No doubt there is a great deal more to be found
out than we have yet obtained. The most peculiar specimens seem to
come from the northwest part, which, I believe, has been but slightly
explored. We know less of it than of any other part, and that leads
one to hope that we may still have further specimens, and that we may
get something which will throw light generally on the Mad
fauna, which is represented also in the Seychelles Islands, in the Rod-
rigues, and in Réunion, also in the Mauritius.” (Proceeds. Roy. Geog.
Soc., Nov., 1892.)
The Nephridia of Amphioxus.—Boveri, in an article’ which
deserves more space than we can give it, describes the nephridia of
Amphioxus, and, in conclusion, summarises his results in the follow-
ing words: There are present in Amphioxus all the elements of the
nephridial system of the Craniata, part with the same function (pro-
ME
É gi oe
Zool. Jahrbücher. Abth. f. Anat. u. Ontog. V., 1892.
284 The American Naturalist. [March,
chamber—pronephric duct), part in wholly other associations (genital
_ chamber=mesonephric tubules). As in the whole of the rest of its
organization, Amphioxus, in its urogenital system, shows in contrast
. with the Craniata, a condition of simplicity and indifference, which is
recapitulated by the latter in their ontogeny.
These facts show that we may recognize the conditions of the excre-
. tory and sexual apparatus of Amphioxus as primitive from which the
_ relations found in the Craniata have probably developed. Amphioxus
_is therefore to be taken from its former isolated position, and it shows
itself to be, as in all its other organs, so with reference to its urogenital
system actually as the primitive type of the vertebrates, as the true
. primitive vertebrate.
The Position of the Marsipobranchs.—Prof. G. B. Howes
__ has reviewed’ the various conflicting views as to the systematic position
and affinities of the lampreys and hag-fishes, and reconsiders the various
_ structural points of value in that connection. He points out that these
forms must be regarded as aberrant gnathostomata; that their uro-
genital apparatus with that of the Teleosts is the least modified survival
„of an hermaphroditic apparatus possessed by the ancestors of the
_ vertebrates; that the sucking mouth of these forms has been second-
arily acquired, and-is not genitically connected with that of the batra-
_chian larva. The arguments from the hypophysis are also considered
„and assigned great weight, and the rasping tongue is given a greater
‘value. in uniting the lampreys and myxinoids than is the sucking
‚mouth. As a result, dismissing, as shown above the term Agnatha for
these forms, Howes divides the Vertebrata proper into Epicraniata and
. Hypocraniata, basing the division upon the position of the hypophysis;
. the Epicraniata containing only the Marsipobranchs. He has also a
. secondary division into Euthorehidic and Nephrorchidie series—the
_lampreys, Teleosts and Dipnoi belonging to the former; all other verte-
_ brates (except, possibly, some ganoids) belonging to the latter series.
He thinks that Haeckel’s famous aphorism that the Marsipobranct
“are further removed from the fishes than the fishes are from man,”
fails to express the enormity of the gap between these forms and the
“ higher vertebrates.
-Degeneration of the Clitoris.—In a paper read before the
American Association of Obstetricians and Gynecologists at the St.
- Louis, N. W., meeting in 1892, Dr. Robert T. Morris stated that about
_ 80 per cent. of all Aryan American women have adhesions which bind — 4
"Trans. Biol. Socy. Liverpool, VI, 1892.
1893.] Zoology. : 285
together the glans of the clitoris and its prepuce. These adhesions
may bind down the prepuce so closely that not a particle of the glans
clitoridis is in sight. They may involve half of the glans, or they may
form only a small band. Adhesions which involve the whole, or a
large part of the glans clitoridis cause profound disturbances in the’
physical and mental health of the individual, and probably form the
most common single factor in invalidism in young women.
In compiling statistics upon the subject, Dr. Morris found that pre-
putial adhesions are rare among negresses, and seem to occur only in
those possessing a large admixture of white blood.
The author considers the degenerate clitoris a chatrieteiiitic of the
civilized white race. (Am. Journ. of Obstetrics, Vol. xxvi, 1892.)
Zoological News—-Reptiles.—Professor O. P. Hay has a
valuable paper’ on the breeding habits, eggs, and young of certain
snakes, to which referencé must be made by all who wish information
on this subject. The same author also notes‘ the ejection of blood from
the eyes of the horned toad. Thesame habit on the part of Phrynosoma
has been noted by other observers, but Professor Hay has settled, by
microscopic examination, the fact that it is really blood which- is
squirted out from the outer canthus of the eye.
Dr. Oppel, of Freiburg, i.B., deals* with the fertilization of iis Bei-
tilian Egg. His observations were made upon Anguis fragilis, Tropid-
notus natrix and Lacerta viridis. The article deals with the behavior.
of the male and female pronuclei and the accessory sperm nucleus, the
questions a to the latter being still left open.
Dr. H. K. Corning, of Prague, deals with some points in n the develop-
ment of the vertebree and the myotomic ceelom in Anguis and Tropido-
notus®, The myotomic celom persists until after the formation of the
neural arches of the vertebræ, hence it is easy to see that the segmenta-
tion of the vertebre results from the formation of inter-vertebral split-
tings which correspond in position to the divisions between the primi-
tive myotomes. The whole question of resegmentation of the vertebral.
column is not, says Corning, so simple as has been thought.
The subject of Variation in the snakes of North America, treated
of by Cope in a late paper,’ is taken up by Hay, in his Presi-
i a U. S. Nat. Mus. XV., 385, 1892; cf. Proc. Ind. Acad. Sci., 1891, p. 109,
“L.c., p 875.
5Archiv. f. mikr. Anat. xxxix., 215., 1892.
ê Morph. Jahrbuch., xvii., p. 611, 1
1Proc. U. S. Nat. Mus., xiv., 589, 1892.
286 The American Naturalist. (March,
dential address before the Indiana Academy of. Science,’ where he
finds in four species of snakes that in Eutaenia sirtalis the variation
from the average number of body vertebræ amounts to 14 per cent., in
Bascanion constrictor to 6 per cent., in Cyclophis vernalis to 4.5 per
cent., and in Diadophis punctatus to 13 per cent. In the caudal verte-
bre the variations amount to 35, 20, 23, and 23.5 per cent. respect-
ively, while in proportion of tail to body the per cents are 9.4, 28, 25,
and 35. Hay states that were breeders interested, they could very
soon produce breeds of snakes with long bodies and short tails, and
short bodies and long tails, or any other combinations that might be
desired. The same author has also some interesting notes’ upon the
systematic names and the habits of the species of Malaclemys.
A. J. Bigney notes” the occurrence of Elaps fulvus in Ripley
County (south eastern) Indiana.
Some observations on the growth of the rattle of the rattlesnake are
given by Dr. Feokistow, who studied specimens sent him from America.
He finds" that the rattle is frequently shed, and (his snakes were kept
in a very warm room) in three or four months two rattles were present,
and that their formation has nothing to do with ecdysis. The snakes
were made to register the vibrations of the rattle on smoked paper, and
it was found that the vibration was a compound one, consisting of the
vibration of the tail as a whole, and of the rattle independently of the
tail vibrations. The approximate figures of vibrations are given for
the tail seventy-five, of the rattle one hundred and ten per “minute.”
Mr. W. E. Taylor has published a paper on the Snakes of Nebraska,
giving descriptions of both the adult and the young of every species
found in Nebraska, together with remarks upon their habits and pecu-
liarities. (Rept. State Board Agric., 1891.)
At the November meeting of the London Zoological Society, Dr.
Gunther read a paper descriptive of a collection of reptiles and Batrach-
ians from Nyassa land, transmitted by Mr. Johnston, containing
examples of several remarkable new species, amongst which were three
new Chameleons, proposed to be called Chameleon isabellinus, Rham-
pholeon platyceps, and R. brachyurus. (Nature, Nov. 17, 1892.)
8Proc. Ind. Acad., 1891, p. 37.
*Proc. U. S. Nat. Mus. xv., 1892. Cf. Proc. Ind. Acad. Sci., 1891.
Proc, Ind. Acad. Sci., 1891, p. 151, 1892.
Mélanges Biolog. Acad. Imp. St. Petersburg, xiii. Translated in Annals and
Mag. Nat. Hist. vi, xi, 54, 1893.
1893.] Zoology. 287
Batrachia—Mr. Stejneger reports a blind Salamander from Rock
House Cave, Missouri. This is one of the most interesting herpetological
events of recent years, since it is the first and only blind form among the
true salamanders. Mr. Stejneger considers it a new genus of the family
Desmognathide, and gives a preliminary description of it under the
name Typhlotrition speleus, in the Proceeds. U.S. Natl. Mus., Vol. XV.
288 © The American Naturalist. [March,
EMBRYOLOGY.
The Star-fish Larva.—Dr. G. W. Field: has published a fully
illustrated study of the development of our common star-fish obtained
at the U. S. Fish Commission Laboratory, at Woods Holl, Mass. Spe-
cial attention was directed to the mesenchyme, mesodermal pouches,
ciliated bands and watervascular system of the larva with a view to,
solving the meaning of the echinoderm larva.
The two symmetrically placed outgrowths of the. archenteron
acquire connection with the exterior by uniting with two dorsal
ectodermal invaginations. These invaginations form two water
pores, which are both open for awhile. The one on the right closes-
up. This double condition isnot, as has been maintained, an abnormal
condition, but as Professor W. K. Brooks showed, a normal and
significant fact.
In discussing the application of these and other facts to the phyllo-
geny of the Echinoderms, the author holds that—‘the Echinoderm
IFNA was ery a free-swimming animal, in general characters
fthe Turbellarians ; a creature witha.
well-differentiated digestive tract, ciliary locomotor apparatus, excretory
system, respiratory surface not localized; coenogenetrically modified’
by the acquirement of transparency, long arms and particularly by
modification of the external form, by changes in the direction of the
ciliated bands, as pointed out by Johannes Miiller, into the forms.
characteristic for the various Echinoderm groups.”
Germ-layers of Amphioxus.’—Basilius Lwoff has recently
published a short paper in which he gives the results of his observa-
tions on the early stages of Amphioxus so far as they differ from those
arrived at by Hatschek.
He finds that there is no period of rest at the end of the cleavage,
at least as far as the smaller ectoblastic cells are concerned, for they
continue to divide frequently. This causes a change in the relative
position of the cells at the boundary between the micromeres and
macromeres, and results in the passive invagination of the entoblast,.
ectoblastic cells being the active agents,
1Q. J. Mic. Sci. 1892.
*Biol. Centralblatt, 12, 1892, pp. 729-744, 8 figs.
PLATE IV.
Sa i a Daal A PEER te al
1. Diastrophus cuseuteformis O. B.
4. Amphibolips confluentus HARB.
5. Amphibolips inanis O.S.
3. Andricus cornigerus O. S. | 6. Amphibolips ilicifoliw BASS.
2. Diastrophus nebulosus O. $.
1893,] Embryology. 289
This multiplication of the entoblastic cells is not equal on all sides,
however, but is most frequent on the dorsal side of the invagination ;
so that the ectoblast becomes invaginated itself at this point and push-
ing the entoblast before it finally forms the roof of the cavity, while, the
sides and floor are lined by entoblast. At the same time the margin
grows backward and the gastrula mouth gradually closes. In this
invagination, then we have two distinct processes: first, the invagina-
tion of the entoblastic cells from which the gut is formed, (it is a palin-
genetic process—the gastrulation); second, the invagination of the
dorsal ectoblastic cells ; this is to be regarded as a coenogenetic process,
that has nothing to go with gastrulation, but is preparatory to the
formation of the chorda and the mesoblast.
Lwoff was unable to find the pole cells of the mesoblast described by
Hatschek, and it is interesting to note that more recent observations
by E. B. Wilson have had the same result. Lwoff points out, more-
over, that even if these cells did exist they could have no connection
with the mesoblastic bands; the latter are upon the opposite side from
the position assigned to the former.
The multiplication of the cells in the medullary plate causes it to
fold inward along the median, and it pushes before it the dorsal wall of
the archenteron, leaving a mesoblastic fold on each side. The mesoblast
is purely passive in its evagination. The inner half of each fold is
composed of ectoblast cells from the roof of the gastrula cavity, and
the outer half of entoblast from the side.
The lumen disappears in each mesoblastic segment after it has been
constricted off from the general fold. Afterward the true mesoblastic
cavity, which is to become the body-cavity, is formed by the separation
of the cells in the process of their growth. The body-cavity, therefore,
is not a true enterocoel.
The chorda is formed from what is left of the ectoblast in the gas-
trula cavity, aided, perhaps, by the entoblast at the anterior end. The
chordal plate becomes folded outward, and the two sides of the fold
are pushed together by the entoblastic cells that at this stage are
multiplying rapidly to form the dorsal wall of the gut.
P. B.
Epigenesis.—lIn an interesting review of the history of Evolution
versus Epigenesis, Prof. C. Hertwig’ contributes a few experiments
upon the eggs of Triton, to those of Chabry, Fiedler, Driesch and
others all tending to overthrow the position occupied by Roux and
35Entwichlungs-Theorieen, Berlin, 1892.
290 The American Naturalist. [March,
Watase regarding the pre-formation and early localization of embryonic .
- organs. While Roux held that the frog’s egg is a mosaic in which
definite regions must become certain organs, the following experi-
ment of O. Hertwig seems to show that this is unlikely in the related
form, the Triton.
When the egg of Triton palmatus and T. cristatus taken in May and
June 1892 was dividing into two cells, a delicate silk thread was
passed around it and drawn together so as to gently squeeze the two
first cells somewhat apart. This made the egg somewhat dumb-bell
shaped.
Each cell divided and finally an embryo with chorda, somites and
nerve tube was formed. As the embryo was not formed so as to lie
with its left on one of the hemispheroid parts of the egg and its right
upon the other, we may conclude that the first cleavage did not divide
the Triton egg so as to separate its right-forming from its left-forming
material. The right and left halves are not separated by the first
cleavage. In fact in one case the thread separated the head from the
tail region.
It is only, the author thinks, by understanding the multiplication of
the egg as an organism and the gradual interaction of the numerous
cells of any stage that we can arrive at a true conception of the
epigenesis-like formation of an embryo.
Form and Chemical Composition.—Curt Hebst* of Zurich
has published a series of experiments made at Naples and at Triest in
the endeavor to determine if the form of organic structures is dependent
upon their chemical composition. To this end the eggs of sea urchins,
(three species were tried), were reared in sea water to which definite,
small amounts of certain salts were added. The salts used were LI
Cl, Li Br, Lil, Li NO,, Li, SO, Na Br, NaI, Na, SO, Na NO, K
Cl, KBr, KI, K NO,, K,, SO, RhCl, CsCl, Mg SO, and Ca Cl, ; the
results obtained were certain peculiar forms of larvæ, and the expla-
nation adopted for the results was that the salts acted osmotically, not
iby altering the chemical constitution of the eggs.
Before speaking of the character of the larve reared under these
abnormal conditions we will first note a few incidental results sometimes
seen. One is that in a number of eggs, two blastule were seen inside
the egg membrane so that separate twins had been formed from one egg-
Again it was sometimes observed that only part of the cleavage cells
formed the blastula, the rest remaining as an irregular mass within the —
‘Zeit. f. wiss. zool. 55, Dec., 1892.
1893.] Embryology. 291
same egg membrane. ‘These facts favor the views of Driesch as to the
equality of the cleavage cells in the echinoderm. Another peculiar
result sometimes followed the addition of salts, namely, the production
of twin gastrulse or plutei (without processes) or even of multiple larvæ
all to be regarded in these cases as due to fusion. The author enter-
tains no doubt, that by some change in the character of the ectoderm cells
the larve first adhere and then fuse till there are formed complete twins
with two separate mouths, ani and (ingomplete) skeletal systems,
though but a single body space.
Coming to the main results of the experiments, we find that two
peculiar larval forms, the potassium larve and the lithium larve may
be formed instead of the normal larvæ when salts of potassium or of
lithium are added to sea water.
The potassium larva is simply a pluteus with its normal digestive
tract and cilia but without the characteristic pluteus processes or arms
and: with little or no skeleton. As an example of the amount of
material necessary to effect this result may be cited one experiment in
which eggs fertilized in normal sea water were put into 860 ccm. sea-
water diluted by 140 cm. of 3.7% KNO, solution. The larve lived
for fourteen days but had scarcely any or no skeleton and no arms.
The absence of the skeleton is regarded as the chief thing determin-
ing the absence of the pluteus arms; these, it is believed, naturally
growing as they are constantly stimulated by the growing skeleton. The
cells to form the skeleton may be properly arranged, but do not secrete
the lime salts to form the skeleton.
This kind of larva may be formed by other salts than those contain -
ing potassium. The lithium larva, however, is formed only by
salts containing lithium.
This latter form may be described as two vesicles attached to one
another by a hollow stalk. One vesicle has a thicker wall and finer
cilia on its outside, the other a thin wall and fewer, longer cilia on its
outside. This larva is actually formed by the elongation of a normal
blastula followed by a partial abstriction into two vesicular portions.
It is regarded, however, as having the morphological value of a gas-
trula which has grown in an evaginated form so that the thicker walled
vesicle represents the entoderm and the other the ectoderm.
One experiment taken at random will serve to illustrate the
amount of salts used; to 1950 ccm. sea-water 50cm. 3.7% LI Cl
solution were added; the characteristic double vesicle larva were
formed, but all died on the eighth day.
292 The American Naturalist. [Mareh,
Most interesting differences obtain amongst the various salts of
lithium in the strength of their action. If the eggs of the same sea
urchin are treated simultaneously with the various salts we find that
at a given time the larve were not all equally far advanced, equal
amounts of the some salts acting sooner than others in producing the
lithium larva, and larger amounts of some salts being necessary to
produce the same results as smaller amounts of other salts of lithium.
From a table of such experiment the author concludes that Li Cl,
Li NO,, Li Br, and Lil are less and less active in this order which is
also the order of increasing molecular weights. Thus in these experi-
ments where the same per centage of salts was always used the heavy
molecules were less numerous and less active ; the action of these salts
in producing the lithium larva diminishes with the number of mole-
cules used. This rule, however, finds an exception in Li, SO,.
In NaCl, Na NO,, Na Br, and NaI as well asin K Cl, K NO,, K
Br and KI we find again the same rule; the larger number of mole-
cules being most efficient in forming the so-called potassium larve, and
so on down to the heaviest. The results hold only for salts of the same
metal.
Now since it is known from the work of H. De Vries and others
that osmotic pressure is associated with the number of molecules in a
given volume, increasing with diminishing molecular weight we find so
close a similarity between the effects of salt upon larve and their
osmotic action, that we may conclude, the author thinks, that these
effects are due to their osmotic action.
Thus the potassium larva is to be regarded as the result of dis-
turbing those chemical processes which would have normally formed a
lime skeleton, and this disturbance is by the removal of water osmoti-
cally. Again the lithium larva may be regarded as due to some
peculiar impermeability of sea urchin larval cells toward salt of
lithium ; this produces strong osmotic pressure. The pressure is not
regarded as working in a gross mechanical way, but rather as a stimu-
lus that causes the larval cells to grow in an abnormal way. 7
Though this explanation leaves no room for chemical changes as a
source of change of form in these echinoderm experiments, yet the
author is inclined to think that in some cases, as in the formation of
galls, chemical changes of the protoplasm may cause the changed form
that results. ae
The application of this study in experimental embryology is that-
the normal course of ontogeny is dependent upon the conditions of —
osmotic pressure within and without the body.
1593.] Entomology. 293
ENTOMOLOGY.
The Pear-Tree Psylla.—During the last two or three seasons
many pear orchards in Connecticut, New York and other States have
suffered severely by the attacks of a small jumping plant louse of the
family Psyllide. This insect is supposed to have been imported from
Europe early in the present century, having been first noticed in Con-
necticut. It now occurs throughout many of the Northeastern United
States, and as far west as the Mississippi Valley.
* Ina recent discussion `of this insect in Bulletin 44 of the Cornell
University Experiment Station, Mr. Mark V. Slingerland records
one of the most notable pieces of entomological work done since the
establishment of the stations. After an introductory paragraph indi-
cating the recent losses due to the pest the author considers its past
history in America ; its place in zoological classification ; the indica-
tions of its presence; the appearance of the immature and mature
insect; its life history in detail, and the methods of preventing its
ravages, all of which is followed by a series of technical descriptions
and a full bibliography. One of the interesting points brought out is
that the species is dimorphic, the summer form having been described
as one species, and the winter form as another. Concerning this the
author writes:
“ This difference between the summer and the winter adults is common
among the Psyllide, and has before led to their being described as
different species. It seems not to have been suspected that these insects
were truly dimorphic or appeared in two distinct forms during the
year. The general impression seems to have been that the adults
appearing in the fall were at first the same as the summer form; and
that as winter approached these adults gradually assumed the charac-
teristics of the hibernating form. However, frequent observations
upon Psylla pyricola in the field during August and September, 1892.
have shown that from eggs lain about August 20th by typical summer
adults, there hatched nymphs which showed no variations from the
typical summer nymphs and from these nymphs there emerged about
September 25th the distinct hibernating form simulans. The hiberna-
ting forms feed until the leaves fall and then seek their hiding places
in which to pass the winter. None have been seen to copulate in the
fall. But very few summer forms were seen after September 20th.
1 Edited by Clarence M. Weed, Hanover, N. H.
994 The American Naturalist. [Mareh,
Thus in our Pear-tree Psylla we have a case of true dimorphism. The
summer form is the typical Psylla pyricola, and may be designated
when necessary to refer to this form alone as Psylla pyricola pyricola;
while the hibernating form should be known as Psylla pyricola simu-
lans.”
Experiments showed that the immature stages of the Psylla were
easily destroyed by spraying with kerosene emulsion. Weare indebted
to Mr. Slingerland for the use of the engravings on the accompanying
plate, showing the stages and structure of the insect. Fig. 1 repre-
sents the adult insect; fig. 2, its head and antenna greatly magnified ;
3, the abdomen of the male; 4, the abdomen of the female ; 5, the
wings; 6, the full-grown nymph, and 7, the egg.
Insects of Southern Alaska.—An important contribution to our
knowledge of the coleopterous fauna of Southern Alaska has recently
been made by Mr. H. F. Wickham,’ who has published some of the
results of a collecting trip made during the summer of 1891. The
points visited were Fort Wrangel, mainland near Wrangel Island,
Yes Bay, Loring; Hunter’s Bay and Port Chester in Alaska, and the
Stikine River Valley and Glenora in British Columbia. From his
studies Mr. Wickham concludes (1) “That the fauna of Southern
Alaska is less closely related to our alpine, northern inland or north-
east coast faune than is that of the Stikine Canyon or of Glenora.
(2) That the Stikine Canyon fauna is more closely allied to that of
the north and east than is that of the coast, and about the same as is
that of Glenora. (3) That the chief relations of all three are in the
direction of Lake Superior. With larger lists this affinity might turn
to the Rocky Mountains, especially in the case of Glenora.”
Notes on Ohio and Other Phalangiidz.— A recent study of a
large collection of harvest-spiders (Phulangiide) from all parts of
Ohio shows that the State is unusually rich in these interesting Arach-
nids. The following species occurred in the collection :
1. Liobunum vittatum (Say) Weed.
2. Liobunum vittatum dorsatum (Say) Weed.
3. Liobunum nigripalpis (Wood) Weed.
4. Iiobunum nigripes Weed.
5. Liobunum politum Weed.
6. Liobunum longipes Weed.
2Entomology of Southern Alaska, Bull. Nat. Hist. Labrt. Iowa State Univ., V. ii,
pp- 202-233.
1893.) Entomology. ; 295
. Liobunum ventricosum (Wood) Weed.
. Liobunum bicolor (Wood) Weed.
_ 9. Liobonum-(?) calcar (Wood) Weed.
10. Liobunwm maculosum (Wood) Weed.
11. Liobunum grande (Say) Weed.
12. Liobunum grande var. simile Weed.
13. Mitopus pictus (Wood) Weed.
14. Mitopus ohioensis Weed.
15. Phalangium cinereum Wood.
An examination of more than fifty specimens of a harvest-spider in
various stages of development taken along the banks of the Maumee
River in Henry County, leads to the conclusion that the form from
Illinois described some years ago as Liobunum elegans is an immature
stage of the male of L. bicolor. A reexamination of the type speci-
men of Mitopus ohioensis after it has been in alcohol more than four
years shows that it was apparently just ready to moult when captured.
This gives rise to the suspicion that this is an immature form of M.
pictus, the pink coloring possibly being due to the peculiar conditions
of the moulting period.
The study and measurement of a considerable number of specimens
of Liobunum ventricosum from many States shows that this species
increases in size to the southward in a way similar to that of L. vitta-
tum. The southern form is evidently sufficiently distinct for a subspe-
cific name, and as the form now standing as Forbesium hyemale is
pretty certainly an immature stage of it, the subspecies may well take
its name and be known as L. ventricosum hyemale.
Illustrated papers on both these subjects are ready for the printer,
and will appear in the near future.
Professor C. H. Tyler Townsend, of the New Mexico Agricultural
College, recently sent me specimens of an undescribed species of Lio-
bunum taken at Las Cruces. It may be called L. townsendii. Its
description is as follows:
Male.—Body 5 mm. long, 3.7 mm. wide; palpi, 5 mm. long. Legs,
first, 43 mm.; second, 80 mm.; third, 45 mm.; fourth, 59 mm. Gen-
eral color of dorsum brown, approaching raw umber, with indistinct
darker blotches, but no central marking. Ventrum light grayish
brown. Palpi similar in color to ventrum, with dorsal surface of
patella and of tip of femur darker brown. General color of legs raw
umber, with whitish rings near articulations and blackish ones at
articulations. Dorsum minutely tuberculate; articulation of the three
posterior segments very distinct. Eye eminence rather high, nearly
co oo =]
‘296 The American Naturalist.
square as seen from above ; canaliculate; smooth except for twosparse
rows of spinose hairs. Palpi rather long, slender, with no projecting
angles ; clothed with rather short hairs and afew minute spinosetu
cles; claw slightly pectinate near base. Mandibles normal; light
brown with tips of claws black. Femora angular. Second legs much
more slender than others.
Female.—Body 7 mm. long, 5 mm. wide; palpi 4.5 mm. long,
Legs, first, 35 mm. ; second, 61 mm. ; third, 37 mm. ; fourth, 46 mm.
Differs from the male chiefly by its larger body and shorter legs.
Described from three specimens (1 ¢ 2 9 ).—CLAReENcE M. WEED.
Gall-Producing Insects.—Mr. Wm. Beutenmiiller, of the Amer- —
ican Museum of Natural History, publishes’ a useful Catalogue of © :
Gall-producing Insects found within fifty miles of New York City
Eighty-eight species are enumerated, the family distribution of which —
is: Cynipide, 40; Tenthredinide, 2; Cecidomyide, 32; Tripetide, 2;
Psyllidæ, 5; Aphidide, 6; Acaroidea,1. The next to the last family —
is called Aphide instead of the more correct Aphidide. “The vege —
table deformations called galls,” writes Mr. Beutenmiiller, are pro- —
duced by insects. Generally an egg is inserted in a bud, a leaf, a root, —
or some other part of the plant, and the presence of this foreign body —
among the vegetable cells causes an abnormal growth of a definite —
shape. The variety of galls in respect to texture and substance is very —
great. Every species of gall-producing insects attacks its own partic:
ular plant, and a particular part of that plant, and produces a gall of |
a definite and uniform structure.” The two plates accompanying (due
to the courtesy of the author) show a variety of common galls. Plate
(IIL) represents those made by the minute two-winged flies of the genus
Cadomyia ; and (IV) those made by the four-winged flies of the family
Cynipidæ.
Recent "Publications.—Mr. Henry G. Hubbard publishes’ an
extended description of thelarva of Amphizoa lecontei, illustrated by
an admirable plate. He extends Schiödte’s table of the larval charac-
ters of the principal families of adephagous coleoptera. :
Through the cooperation of the Massachusetts Society for Promoting
Agriculture, the Hatch Experiment Station of the Agricultural Col-
lege has issued an edition of 45,000 copies of a bulletin concerning
canker-worms, tent caterpillars, fall web-worms and tussock moths
*Bull. Am. Mus. Nat. Hist., V. iv, Art. xv.
*Proc. Ent. Soc, Wash., 1i, pp. 341-346.
1893.) Entomology. | 297
written by Professor C. H. Fernald. A special attempt is being made
to induce the people of the State to suppress these pests. The author
states that “There has been such culpable negligence on the part of
many of our people with regard to the tent caterpillar that there can
be no doubt that some legislation is needed to compel the negligent to
destroy this pest on all the trees on their own land, and thus prevent
it from extending to the trees in the surrounding orchards. Provision
should be made for the destruction of tent caterpillars on all public
lands as well as in the forests, and village improvement societies should
urge such action in town meetings as shall make it the duty of the
superintendent of roads to destroy all tent caterpillars on the trees and
shrubs along the sides of the roads.”
Mr. S. H. Seudder’s Monograph of the Orthopteran Genus Hippis-
cus which has been running through Psyche for some time has been
issued as a reprint. The thirty-eight species are grouped under the
subgenera Hippiscus, Sticthippus, and Xanthippus.
Two interesting papers on the Butterflies and Crickets of Indiana
have been published by Mr. W. S. Blatchley, of the Terre Haute High
School. The former is extracted from the 17th Report on the Geology
and Natural History of Indiana, and the latter from the Proceedings
of the Indiana Academy of Science, 1891.
Mr. Wm. Beutenmiiller records in the Bulletin of the American
Museum of Natural History (v. IV, Art. XIII) an important List of
Types of Lepidoptera in the Edwards Collection of Insects. This
collection (made by the late Henry Edwards) “ consists of about 250,-
000 specimens and about 25,000 species, representing all the orders
and gathered in various parts of the globe. It is especially rich in
Australian species and in North American species from the Pacific
Coast. The present list enumerates 465 types of species.
The department of entomology of the University of Kansas has
recently published a bulletin of 126 pages concerning “Common Inju-
rious Insects of Kansas,” prepared by Vernon L. Kellogg. The paper
is well illustrated and will prove valuable to Kansas farmers.
298 The American Naturalist. [Mareh,.
PSYCHOLOGY.
The Sense of Taste in a Sea-Anemone.—It is a well known
fact that sea-anemones possess a faculty which enables them to recog-
nize food. This sense has been recently studied by M. Nagel of the
Zoological Station at Naples. The following is one of his experiments.
A small piece of sardine was gently pushed toward the tentacles of a |
sea-anemone ; the morsel was first touched, then seized, then swallowed.
A small ball of blotting-paper, similar in appearance, saturated with
sea-water was next offered, but it was not taken. The paper was then
saturated with the juice of the fish, when it was seized with the same
avidity as the bit of sardine, but often it was rejected after the lapse of
a few moments.
When the blotting-paper is impregnated with quinine the tentacles
recoil, The quinine does not affect the external surface of the body,
except the part situated between the tentacles and the mouth. If food
is placed in the mouth, or near the open mouth, the animal does not
notice it, but will only take it when the tentacles have touched it. The
sense of taste, then, is localized in the tentacles, which serve as organs
of touch ; and also appreciate changes of temperature. (Revue Scien-
tifique, Dec. 1892.)
M. Vaillant on the Feeding of Snakes.—In a paper read
before the Academie des Sciences de Paris, M. Léon Vaillant made the
following interesting remarks concerning the alimentation of Snakes.
These remarks were the result of a series of observations of a large
Anaconda from South America, Eunectes murinus, one of the Boidae,
about 6 meters long, which, contrary to the usual habits of this species
accepted food very soon after its arrival at the menagerie of the Rep-
tiles of the Museum, and has continued to eat with regularity up to the
present time. In fact, since its entrance into the Jardin des Plantes,
August 8, 1885, until the end of the year 1891, this serpent has eaten
34 times, or about 5 times a year. Its food has generally been small
or young goats; three times it took a hare and once a goose. The
intervals between the times of feeding have varied from 23 days to 204
days; this last interval occurred but once. The snake decided for
itself the time for feeding, manifesting its desire by increased activity, 2
and by other signs. i
1893.] Psychology. 299
As to quantity of food, in order to avoid all accidents which might
result from indigestion, the Anaconda was given animals of moderate
size; the largest it has swallowed is a kid of about 12 kilograms
weight ; which represents one sixth the weight of the snake. Thereis
no doubt, however, that in a wild state, a snake of its size could swal-
low animals three or four times as large. ;
M. Vaillant adds, in this connection, the following fact which shows
the stretching capacity of certain snakes. In the menagerie of the
museum, a viper from France, (Pelias berus) had to be put in the
same cage with a horned viper, ( Cerastes.) As the individuals,
although they belonged to different species, were about the same size,
it was supposed that they would live peaceably together. It was a
mistake. During the following night the Cerastes swallowed the Pelias
berus, and in order to accommodate himself to his huge prey, his body
was distended so that the scales which touch each other laterally, and
even lap in its normal state were now so spread apart that between the
longitudinal rows, a bare space equal in size to the scales was left.
Digestion went on regularly, however, and the Cerastes did not appear
to suffer.
The author also remarked that snakes in general do not accept
indifferently all sorts of food, but appear to exercise a choice. It is
often difficult to induce a snake to take food for the first time, but
once this is accomplished, it accepts more readily succeeding proffers.
A Pelophilus madagascariensis has been known to refuse a variety of
food for 22 months, when it atea starling, then a few other small birds
and finally some rats, which are still fed to him without any difficulty,
although previously they had been offered to him in vain.
M. Vaillant also remarks that the residue of digestion are evacuated
at a single time after each feeding, at the end of a certain number of
days. However, it often happens that the fæces contain the residue of
a former meal. (Revue Scientific, aont, 1892.)
300 The American Naturalist.
ARCHEOLOGY AND ETHNOLOGY.
International Congress of Americanists.—It is proposedin
this department to make a series of sketches of some of the Interna- —
tional Congresses held in Europe. Two of these Congresses, to wit, that
of “Criminal Anthropology,” of Brussels, and that of the “ American- —
ists,” at Huelva, voted to hold a special meeting in the United States
during the Summer of 1893; both, presumably, to be at Chicago. —
The increased interest thus to be engendered justifies this publication.
Ba ogee
THE FIRST InrERNATIONAL CONGRESS OF AMERICANISTS.—On the
25th of August, 1874, the Société Geographique Francaise held a meet- |
ing in the City of Paris, and formulated a Constitution, which should
serve as a foundation for a new Society, to be called the Congrès —
Internationale des Americanistes. A
The object was to contribute to the progress of Ethnographic,
Linguistic, Historic studies relative to North and South America, and —
especially those of antiquity. The Historic or Proto-Historic portion —
would naturally be directed to the time of the discovery by Christopher
Columbus. 7
Committees were appointed, officers and their duties provided, requi-
sites of membership laid down, all in the shortest and simplest manner
possible, and it finally provided that the First Congress should be held
= the City of Nancy, on the 19th of July, 1875, and to continue four
ys. r
` The First Congress was entirely successful, and demonstrated the
good judgment of its founders. =
Twenty-eight countries were represented. Professor Henry, Secre-
tary of the Smithsonian Institution, and Hon. Robert T. Winthrop,
General Clary and Professor Henry W. Haynes, of Boston, were dele- — |
gates from the United States; although it does not appear that the two
former were present. ` :
The report of the Treasurer showed 1572 subscriptions at 12 francs
each, with total receipts of 23,106 franes. The meetings were held in —
the Ducal Palace, at Nancy, under Government patronage and protec” ;
tion; the city was put in gala costume, the palace and streets were
draped with the flags of all nations, and there were the usual exeur-
sions, receptions, fétes, concluding with the official banquet. :
‘This department is edited by Dr. Thomas Wilson, of the U. S. National Museum.
ea ae Pa D AT RIA E eA PE e pony Ee See ee MORO can an ae Se eee ce ee eS a ee
1893. ] Archeology and Ethnology. 301
There was an Exposition of American Antiquities, principally from
the collections of the Archeologists of two countries; the most exten-
sive being that of a most esteemed friend, M. E. Boban, of Paris and
Mexico.
The papers read and the subjects discussed are here given to show
the scope of the organization.
Discovery of America before the time of Columbus, by M. Benedict
Grondals, 7 pages; Discovery of the New World, by M. E. Beauvois,
52 pages; The Pheniciens in America, by Gaffarel, 48 pages; Bud-
dhism in America, by M. Foucaux and M. De Rosny, with discussion,
12 pages; Fou-Sang, by Lucien Adam, 18 pages; Atlantis, by M.
Chill y Naranjo, 3 pages; The Dighton Rock, by M. G. Gravier, 26
pages; A Dream of Christopher Columbus, by M. Castaigne, 23 pages ;
Inscription on the Grave Creek Tablet, by M. Levy Bing, 17 pages;
America and the Portuguese, by M. Luciano Cordiero, with discussion
by Professor Haynes, 97 pages; The Arctic and Antarctic Regious, by
M. Daa; The Esquimaux, by R. P. Petitot; The Indians of the United
States, by M. De Semallé, 9 pages; The Ancient Races. of Peru, by
M. John Campbell, 19 pages; Columbian Skulls, by M. Paul Broca,
16 pages; The Aborigines of Haiti, by Madiou; The Tradition of the
White Man, by Madier De Montjau; An Iroquois Manuscript, by
Leon De Rosny ; The Mound Builders, by M. Joly; The Indians of
French Guiana, by M. Dupont; The Caribs, by M. Ballet; The Ori-
gin of America, by Baron de Bretton; The Indians of Peru, by
M. Ber; Le Maguey, by M. Gordron; The Guano of Peru, by M.
Ridel; Language of the Cheyenne Indians and the Quichua, by
Lucien Adam,7 pages ; Les Déné-Dindjies, by R. T. Petitot, 24 pages,
and An Iroquois Manuscript ; The Relation of Words, by Lucien Adam,
6 pages; Comparison between the Basque Language and Indian Lan-
guages of America, by M. Julien Vinson, 14 pages; Deciphering the
Maya Language, by Leon De Rosny,5 pages; Central America, by M.
Blaise, 2 pages ; The Society of Quakers, by M. Magnin ; The Crees and
the Chippeways by Lucien Adam, 59 pages; Anthropology in the
Antilles, by M. Cornilliac, 22 pages; Traditions of the Greenlanders, by
M. Rink and Valdemar Schmidt, 9 pages; The falsity of the tirai
inscription found on a stone at Newark, Ohio, by M. Henry Harrisse, 7
ges; America in Antiquity, by Francis A. Allen, 47 pages; Asiatic
Emigration to the New America, by R. P. Petitot, 11 pages; Ancient
Mexican Stirrups, by Eugene Boban, 3 pages; Archeologie Analogies,
by M. Morey, 7 pages; Prehistoric Canada, by M. Le Metayer-
Masselin, 8 pages; The Museum of St. Petersburg, by M. Schoebel, 3
302 The American Naturalist. [Man 4
:
pages; Ancient American Music, by Oscar Comettant, 27 pages; —
Alphabet of one of the Ancient Languages in the interior of South —
America, by M. Pacheco-Zegarra, 28 pages; Memoire on the numera-
tion of the Maya Language, Leon De Rosny, 20 pages. es
THE SECOND INTERNATIONAL CONGRESS OF AMERICANISTS Was —
held at Luxembourg, from the 10th to the 14th of September, 1877. —
There were 1047 subscriptions, and the total receipts, including con- —
tributions, amounted to 18,338 francs. Twenty-seven countries were
represented. a
The titles of the papers presented and addresses delivered are as H
follows :— 4
The Ancient Pueblos, by Edwin A. Barber, 17 pages; The Mound
Builders, by Mr. Robert S. Robertson, 26 pages; The Chinese in Cal
ifornia, by M. Emile Guimet, 9 pages ; Osteologic Evidence furnished
by the Ancient Mounds of Michigan, by Mr. Henry Gillman, 15 pages;
The Origin of Civilization in the New World, by Mr. F. A. Allen, 20 3
pages, with the Protestation, by M. L’abbe Hengesh, 4 pages; The
Mound Builders, by Mr. S. D. Peet, 17 pages; To What Race Belon a
the Mound Builders? by Judge M. F. Force, 25 pages; The Origin of
the Language, Mythology and Civilization of America, found in the Old |
World, by Mr. Hyde Clarke, 14 pages; The History of America and
its Discovery, by Mr. De Hellwald, 4 pages; The European Colonies —
in Markland and l’Escociland, in the XVI and XVII Centuries, by 1
M. E. Beauvois, 61 pages; A Comparison between the Civil Legisla- 4
tion of the Mexicans under the Aztec Emperor, and of the Peruvians —
during the epoch of the Incas, by Sr. J. F. Nodal; The Route of the a
Mississippi, by M. Gabriel Gravier, 75 pages; Points or Marks in the —
Chronology of the History of the Mound Builders, by M. Stronck, 18
pages ; The Migration of the Nahuas, by J. H. Becker, 26 pages; —
Address of Mr. Da Silva Paranhos and his criticism of the work of —
M. le Doctor Couto deMacalhaes, entitled “ O Selvagem ” or the Sav-
age, being an investigation into the Antiquity of Man in South Amer a
ica, and especially in Brazil; Americus Vespucci, by Dr. Schoetter, 8
pages; Theories on Ka-kwaks, and their destruction by the Senecas,by
Abbe Schmidt; Conquest of the Ancient Chilleans by the Peruvians
in the times of the Incas, by M. H. Savary, 2 pages ; Pay-Tuma, by Abbe —
Schmidt, 6 pages; Prehistoric Synchronism, by M. Anatole Bamps, 29
pages; The Discovery of Brazil by the French, by M. P. Gaffarel, st
pages; Memoirs of Brazil, by M. Burtin ; the voyage of Verrazzano, by —
M. Desimoni; The Ethnographic distribution of Nations and Lan- —
1893.] - Archeology and Ethnology. 303
guages in Mexico, by M. V. A. Malte-Brun, 36 pages; The Language
Atacameiia, by M. Moore, 22 pages ; The Manuscript of M. Platzmann ;
The Indian Languages of America compared with the Ural-altaischen
Languages, by Forchhammer, 19 pages; Is the Quichua an Aryian lan-
guage? Being a critical examination of the works of Don V.F. Lopez;
The Aryenne Races in Peru, by M. V. Henry, 83 pages; An inscribed
tablet, by M. J. Gass, 2 pages; The Engraved tablet of Rockford, Illi-
nois, and its evident fraudulent character, by N. Moody ; A grammat-
ical examination and comparison of 16 American Languages, by M.
Lucien Adam, 83 pages; Principles of the Cree Language, by M. R.
P. Remas, 10 pages; The Age of Stone, at the exposition of Philadel-
phia, 5 pages, by Emile Guimet ; The National Library of Rio Janeiro,
by Ferdinand Denis, 8 pages; American Antiquities in the Royal
Netherland Museum at Leyden, by M. Leemans, 20 pages; A chapter
in American Archeology, 16 pages, by C. Schoebel ; A rock-shelter in
Pennsylvania, by Mr.8. S. Haldemann, 8 pages; A primitive habita-
tion of the Esquimaux, by M. H. Rink, 14 pages; The antiquity of
Man in America attested by the Silex. The evidence upon which this
paper relies to prove its proposition, is the similarity of the flint imple-
ments of America with those of Europe, by Jean Engling ; The collec-
tion of M. Emile de Ville, Belgium Consul at Quito, and its proposed
deposition or gift to the city of Brussels and its deposition in that
Museum; Antiquities of Greenland, by M. Valdemar Schmidt, 3 pages.
Tur THIRD INTERNATIONAL CONGRESS OF AMERICANISTS was held
at Brussels, from the 23d to the 28th of September, 1879.
The number of Nations represented was about the same at the two
previous Congresses ; while the number in attendance, and the papers
read and addresses made were greater. The proceedings were reported
in two large volumes containing together 825 pages, with an atlas of
lates,
The following papers were read and addresses delivered :—
Pre-Columbian Historic Documents from Mexico and Anahuac, by
M. Andre de Bellecombe, 13 pages; The Calpullis of Mexico, their
administration, origin and communistie principles, by Mr. Ad. F.
Bandelier, 3 pages; The Norambégue, with the proof of its Scandina-
vian origin furnished by the language, institution and belief of the
Aborigines of Acadia, by M. Eugene Beauvois, 38 pages; The explo-
ration of the Amazon by the Franciscans of Peru, by P. Servais
Dirks, 31 pages; Progrese of American Cartographie during the XVI
Century, by Rev. F. B. De Costa, 8 pages; The same (Continued ) by
304 The American Naturalist. -[Mareh,
M. Gabriel Gravier and by Lieut. Col. Adan, 89 pages; An unknown
chart, the first one made by Louis Joliet in 1674, after his exploration _
of the Mississippi with Father Marquette in 1673, by M. Gabriel Gra-
vieri with explanations by M. Lucien Adam, 38 pages; Observations
on the earlier letters of Americus Vespucius, by Judge M. F. Force, 36
pages; On the influence of orography on the march of civilization in
America and Europe, by Dr. Charles Barrois, 4 pages ; Printing and
Bookmaking in Spanish America, from the XVI to the XVII Cen-
tury, by Vicente G. Quesada, 68 pages; The antiquity of different
Canadian States, by M. Burtin, 3 pages; Verraxxano, discoverer of
certain regions in North America, by Cornelio Desimoni, of Genoa, 67
pages; The Carib Language, and the differences between it as spoken
by men and women, L. Adam; Peruvian Ceramics in the Society of
Americanists at Nancy, by M. Jules Renauld, 20 pages; Christianity
and the White Man in America before its discovery by Christopher
- Columbus, by Abbe Schmitz, 15 pages; The White Man and the Cross
in America, by M. Peterkin, 20 pages ; The White Man and the Cross in
Peru before the Discovery by Columbus, by D. Marcos Jiminez de la
Espada, 28 pages; Collection of M. Guesde, of Port-au-Prince, spec
ially the Carib stone hatchet, 3 pages; American Antiquities recently
acquired by the Royal Netherland Museum of Antiquities at Leyden,
by Dr. Leemans, 3 pages.
Antiquities of Greenland, by Valdemar Schmidt, 6 pages; The last
discoveries in the Mounds of America, by Rev. J. Gass, with a letter —
from M. Francis A. Allen, 26 pages; Effigy Mounds in America com- ;
pared with those of the same kind in Europe and Asia, by Dr. Phéne,
7 pages; The probable destination of the Inca-Chunca, by Jean Van
Volxem; Antiquities from the Equator at the Royal Museum at Brus
sels, by Anatole Bamps, 96 pages; The antiquities from the valley of
San Augustine, United States of Columbia, by M. Jose Maria Gutier-
rez de Alba, 5 pages ; The discourse of Dr. Virchow ; The influence of —
American surroundings upon the White Race, by M. Grattan, 8 page> í
Existence of Man in North America during the Glacial Period, by M,
Sidney Skertchly, 4 pages; The reefs of St. Paul and the question of
Atlantis, by Abbe Renard, 22 pages; The last of the Mexican Races in
the United States; by M. Edwin A. Barber, 6 pages; The highest i
antiquity of Man in America, by M. Florentine Ameghino. This :
paper has reference to the man during the tertiary period in Brazil, 52 a
pages ; Cosmogonye Algique, by Count de Charencey, 28 pages; Orig
of the Primitive Indians of South America; by Mme. Marcella J. Wik
kins, 8 pages; A critical examination and comparison of fourteen
PLATE V.
The Pear Tree Psylla.
1893.) Archeology and Ethnology. 305
American Languages, by M. Lucien Adam, 61 pages; Grammar of
the Moskito Language, by Edward Grunewald, and Dictionary of the
same, Moskito and German ; Inscribed Stones purporting to be in
Hebrew, from Licking County, Ohio, by M. Charles Whittlesey ; A
comparative grammar of three languages of Greenland, by Victor
Henry; A comparison between the Aymara of the Quichua and the
dialect of Quito, by Jose Fernandez Nodal; Principles of Language
and Othomie, by Professor de Harlez, 45 pages; The Affiliation of
American languages, by M. Jobn Campbell; A Communication on the
Idol of Guaqui, by M. Marcos Jiminez de la Espada; Philologic
Notes, by M. P. Vegreville, 39 pages; The Mexican Calendar, by Man-
uel Orozcoy Berra; Ancient inscriptions in the Argentine Republic,
by Florentine Ameghino, 27 pages ; Prehistoric Clocks in South Amer-
ica; The language Maskoki and its dialects, by M. Albert 8. Gatschet,
16 pages; The deciphering of certain Maya characters, by the Count
de Charency.
21
306 The American Naturalist. — [March,
PROCEEDINGS OF SCIENTIFIC SOCIETIES.
American Society of Naturalists.—This body met at Prince-
ton, N. J., in J. C. Green Science Hall of Princeton College, on Decem-
ber 28-9, 1892. Professor Osborn, of Columbia College, N. Y., was
President, and Professor T. H. Morgan, of Bryn Mawr College, Pa., was
Secretary. On the afternoon of Dec. 28th, President Patton, of Prince-
ton College, gave an address of welcome. A committee was appointed
to confer with the executive committees of the affiliated societies to
bring about a union between the societies of Morphologists and Anato-
mists. Professor Sedgwick read a report by Professor S. F. Clark, on
Science Teaching in Schools. Reports on Marine Biological Laborator-
ies were presented by Dr. E. A. Andrews, of Johns Hopkins University
on the Marine Station in Jamaica; Dr. D. Bashford Dean read a
paper on the Marine Laboratories of Europe, which was handsomely
illustrated ; Professor C. O. Whitman read a paper on the Outlook of
the Marine Laboratory at Woods Holl, Mass. Dr. Stiles read a report
of progress in obtaining support for an American table at the Marine
Laboratory at Naples. The Society adopted resolutions favoring the
establishment and continuous support of a table at that Laboratory.
In the evening the Annual Dinner took place in University Hall
After the address by the President, Professor Osborn, remarks were
made by Professor Brewer, of Yale, Professor Cope, of Pennsylvania,
Dr. Heitzman, of New York, Professor Minot, of Harvard, and others.
On Dee. 29th, the following members were elected: Howard Ayers, —
J. H. Bamhart, G. W. Calkins, T. M. Chapman, D. Bashford Dean,
D. G. Elliot, B. D. Halsted, Ida A. Keller, Edwin Linton, T. B. Lotsy,
J. I. Peck, H. 8. Pratt, J. E. Reighard, Wm. E. Ritter, J. P. Smith,
O. S$. Strong, R. Thaxter, H. H. Wilder, Arthur Willey, J. L. Wort- —
man.
Professor Libbey offered the following resolution, which was carried :
That a committee be appointed to bring about an affiliation of the
Physiological, Te am des Anatomical and Geological Societies
with this body.
New Haven was adopted as the next place of meeting. The follow-
ing officers were elected for the ensuing year: President, Professor R.
H. Chittendon, New Haven; Vice-Presidents, Professor G. Baur, Chi- —
cago, and Dr. Wm. H. Dall, Washington, and Professor Wm. Libbey, :
Princeton; Secretary, Professor T. H. Morgan, Bryn Mawr, Pa.;
1893.] Proceedings of Scientific Societies. 307
Treasurer, Professor W. T. Sedgwick, Boston ; Executive Committee,
Professor W. G. Farlow, Cambridge; Professor J. A. Ryder, Philadel-
phia.
Special reports on the work of the preceding season were made by
Professor W. Libbey, Jr., on the summer work of the U.S. Fish Com-
mission schooner, Grampus ; by Dr. J. L. Wortman on the Exhibitions
of the American Museum of Natural History in New Mexico,
Wyoming and Dakota; and by Professor E. D. Cope on the Paleonto-
logical Exploration of the Staked Plains of Texas by the Geological
Survey of Texas.
On Dec. 29th the Annual Discussion took place. The question was,
What were the former Areas and Relations of the American Continent
as determined by Faunal and Floral Distributions? The discussion
was opened by a paper from Professor W. B. Scott, of Princeton, on
“ Evidence from past and present Distribution of Mammalia.” Pro-
fessor J. A. Allen read a paper on the Evidence from the Distribution
of Birds; and Dr. N. L. Britton on Evidence from the Distribution of
lants.
American Morphological Society.—The annual meeting was
held at Princeton, N. J., Dec. 28-29, 1892. The following papers
were read: Dr. E. B. Wilson, Columbia College, The cleavage of the
ovum and the teloblasts of Amphioxus. Dr. C. W.Stiles, Department
of Agriculture, Washington, The topographical anatomy in the family
Tæniadæ. Dr. E. O. Jordan, University of Chicago, The maturation
and Fertilization of the egg of the Newt. Professor E. D. Cope, Phil-
adelphia, False Elbow Joints in Man and the Horse. Mr. Arthur
Willey, Columbia College, On Acinetæ parasitic in the buccal tube of
Diplosoma. Dr. C. B. Davenport, Harvard College, The development
of the cerata of Eolis. Dr. H. B. Ward, University of Michigan, The
host of Nectonema. Dr. C. O. Whitman, University of Chicago, The
metamerism of Clepsine. Dr. W. B. Scott, Princeton College, The
evolution of the premolars. Dr. H. A. Ayers, of the Lake Laboratory,
Milwaukee, The ending of the auditory nerves in the hair-cells. Dr.
E. A. Andrews, Johns Hopkins University, Notice of a new sort of
Amphioxus. Professor A. E. Verrill, Yale College, Some New forms
of Nemerteans. Dr. T. H. Morgan, Bryn Mawr College, Preliminary
note on Balanoglossus. Professor B. Sharp, Academy of Natural
Sciences, Philadelphia, Joint formation among the Invertebrata. Pro-
fessor W. A. Locy, Lake Forest University, The formation of the
308 The American Naturalist. [March,
medullary grooves, and some other features of embryonic development
in the Elasmobranchs.
The following gentlemen were elected officers for the current year:
President, Dr. C. O. Whitman, University of Chicago ; Vice-President,
Dr. E. B. Wilson; Columbia College; Sec’y-Treas., Dr. J. Playfair
McMurrich, University of Cincinnati. Members of the Executive
Committee elected from the Society-at-large: Dr. T. H. Morgan, Bryn
Mawr College; Dr. C. B. Davenport, Harvard College.
Association of American Anatomists.—Fifth annual session,
Tuesday to Thursday, December 27 to 29, 1892, at Princeton, N. J.
The officers for the year 1891-1892 were—President, Harrison Allen,
M. D., Philadelphia; First Vice-President, Charles Heitzman, M. D.,
New York City; Second Vice-President, Theodore N. Gill, M. D.,
Washington, D. C.; Secretary and Treasurer, D. S. Lamb, M. D.,
Washington, D. C. Executive Committee: Thomas Dwight, M. D.,
Harvard University ; E. C. Spitzka, M. D., New York City; Dr. F. H.
Gerrish, Portland, Me., the President and Secretary, ex-officio. Com-
mittee on Anatomical Nomenclature: Harrison Allen, M. D., Phila-
delphia; Frank Baker, M..D., Washington ; Thomas Dwight, M. D..
Harvard University; Thomas B. Stowell, Ph. D., Potsdam, N. Y.;
Burt G. Wilder, M. D., Cornell University, Secretary. Delegate to
American Congress of Physicians and Surgeons, F. J. Shepherd, M. D.,
Montreal, Canada. Alternate, R. W.Shufeldt, M. D., Washington, D. C.
Tuesday, December 27, the following business was transacted: 1,
Opening of the session by the President; 2, Report of the Executive
Committee ; 3, Report of the Secretary and Treasurer ; 4, Election of new
members ; 5, Report of the Committee on Anatomical Nomenclature;
6, amendment to Constitution, abolishing dues and substituting assess-
ments; 7, Miscellaneous business.
Then followed the address of the President, Dr. Harrison Allen.
The following papers were read: 1. History of the development of
bone tissue. Illustrated by microscopic slides. Dr. Carl Heitzmann,
New York City; discussed by Professors Macloskie and Libbey. 2.
Crania of the Cetacea (15 minutes), with specimens, Dr. Harrison
Allen, University of Pennsylvania. 3, The human lower jaw (10
minutes), with specimens, Dr. Allen, The second and third discussed
by Dr. Herrick and Professor Macloskie. 4, An anomalous human
sternum, Dr.C. S. Lamb, Washington, D. ©. Discussed by Dr.
Dwight. 5. Observations on the psoas parvus and pyramidalis. A
study on variation, Dr. Thomas Dwight, Harvard University. 6.
1393.] Proceedings of Scientifie Societies. 309
Significance of percentages in reversions in human anatomy, Professor
H. F. Osborn, Columbia College, New York. Discussed by Professor
Cope and Drs. Dwight and Lamb. 7. Series of thirty-five natural-
size photographs of sections of human brain, with brief remarks,
Dr. I. S. Haynes, University of New York. 8. Histogenesis in the
brain, and its bearings on development and decline, Professor C. L.
Herrick, University of Chicago. Discussed by Drs. Heitzmann and
Piersol. 9. The metapore or foramen of Magendie, with photographs,
Professor G. B. Wilder, Cornell University. Read by Mr. Clark.
10. Neuromerism and the cranial nerves of Ophidia, Professor Herrick,
11. The insula of the pig, with specimens, Tracy E. Clark, B. S., Clin-
ton Liberal Institute, Fort Plain, N. Y. Discussed by Drs. Allen,
Dwight and Spitzka. 12. The posterior surface of the liver ; described
by Vesalius; Dr. F. H. Gerrish, Bowdoin College, Maine. Discussed
by Drs. Dwight, Heitzmann and Allen. 13. Embryos of bats, (with
specimens), Dr. Allen. Discussed by Professors Cope and Minot. 14.
Meckel’s diverticulum, Dr. D. S. Lamb, Army Medical Museum, Wash-
ington. Discussed by Drs. Minot and Dwight. 15. Delimitation of
abdominal regions, Dr. E. A. Balloch, Howard University, Washing-
ton. Read by the Secretary. 16. The need of agreement in the
limits of the abdominal regions, Dr. Gerrish. Discussion of the last
two papers together, by Dwight, Piersol, Kemp, Heitzmann and Lamb.
17. Physical characters of the Kootenay Indians, Mr. Alex. F. Chamber-
lain, Clark University, Worcester, Mass. 18. Discovery of an ossified
thyroid cartilage, and a supposed rudimentary clavicle in an artiodac-
tyl, with the specimen; Professor W. B. Scott, Princton, N. J. Dis-
cussed by Professor Cope and Dr. Allen. 19. Notes on diagrams of the
spinal cord, Dr. J. T. Duncan, Toronto, Can. 20. Duration of motion
of human spermatozoa, Dr. Geo. Piersol, University of Penna. Dis-
cussed by Drs. Spitzka and Heitzmann. 21. The innervation of the
organ of Corti, Dr. Howard Ayers, Lake Laboratory, Milwaukee, Wis.
Microscopical slides, with remarks
New York Academy of Sciences.—Biological Section, Jan.
9.—The following papers were presented:—A. A. Julien, Suggestions
in Microscopical Technique, including (@) a carrier of cover impres-
sions (mycoderm blood) utilizing as clampsa coil of brass wire mounted
in a phial. The same device with a platinum coil serves as a conveni-
ent staining phial for cover glass preparations; (b) a suggested
medium for mounting delicately contractile protoplasmic objects; (¢)
310 The American Naturalist. [Mareh,
devices for avoiding inclusion of air bubbles in mounts; (d) balsam-
paraffin as a ring varnish.
O. S. Strong: On the components of cranial nerves of Amphibia.
In the seventh a dorsal root was shown to pass off into branches rep-
resenting ophthalmicus superficialis, facialis and buccalis of fishes, and
innervating the lateral sense organs of the head. In vagus, a root of
similar internal origin passes into the R. laterales, innervating the
lateral sense organs of the body. Another component of the facialisis
the fasciculus communis of fishes. This passes off into the palatinus
and mandibularis internus, innervating the mucous epithelium of the
oral cavity ; while in the glosso-pharyngeus and vagus, similar com-
ponents divided from this fasciculus innervate in like manner portions
of the alimentary canal and its appendages. The relation of the results
to segmentation of head was discussed.
N. L. Britton: A Review of the N. A. species of Lespedeza, with
comments on the eleven native species, shown to be divisible into two
groups, (a) those producing both petalous and apetalous flowers, and
(b) those in which no petalous flowers are developed. Of the two
naturalized species, one in S. E. part of U. S., L. striata (Thunb.) H
and A, is a native of E. Asia, appearing (about 1845) in Georgia.
Basurorp Dean, Ree. Secy. of Section.
Proceedings of the New Mexico Society for the Advance-
ment of Science.—Meeting of November 2, 1892, at La Cruces.—
(No. 1.) Professor Townsend read a paper entitled “A partial comparison
of the insect fauna of the Grand Cafion with that of the San Francisco
Mountain in Arizona.” The paper discussed briefly the conditions
which exert an influence on the vertical range of species, and then gave
some exact data on the topographical features of the above region.
Dr. Merriam’s life-zones of the San Francisco Mountains were re-stated,
and the following provisional zones were offered for the Grand Cafion
(from top of south rim at Hance’s, 7500 ft., to Colorado River, 2500 ft.) :
Canadian or balsam fir zone (only on north or northeast exposure).
Neutral or pine zone (only on north or northeast exposure).
Piñon or cedar gst Bot te to 7500 ft. (top of south or southwest
exposure).
Sub-desert zone—4500 to 6000 ft.
Gila zone—2500 to 4500 ft.
The insect fauna was considered in three sections—that peculiar to
the cafion, that peculiar to the mountain, and that common to both.
divisions were compared with each other, and also with the fauna
1893.] Proceedings of Scientific Societies. 311
of the surrounding region. Specimens of insects were exhibited, belong-
ing to three divisions of the fauna, and many photographs and
views of the cañon region were shown.
(No 2.) “ Exhaustion and renewal of soils,’ by Professor Arthur
Goss. He first described soils chemically, and told where they obtain
their various constituents. He next divided the productive existence
of a soil into three periods—1. The pioneer period. 2. The period
when soiling crops are necessary. 3. The fertilizer period. The
desired thing in fertilizers is the element or compound which has been
taken from the soil. The best way to find this is not by chemical
analysis of the soil, but by judicious experimentation,
(No. 3.) Professor C. T. Hagerty read a paper entitled “ Computa-
tion of the Comparative strength of Insects and the Higher Animals.”
He proved conclusively that the higher animals are much stronger
than insects in proportion to their size, and readily accounted for the
apparent superior strength of the latter. In the course of his remarks
he referred to a computation on the comparative strength of the honey-
bee and horse, made by Miall and Denny in a work entitled “ The
Cockroach ” (page 82, edition of 1886), and pointed out an error in
their computation. They state that the relative muscular force of the
horse is more than fourteen times as great in comparison with that of
the bee as it would be if the muscles of both animals were similar in
kind and the proportions of the two similar in all respects, and he
showed that according to their own method of solution, it would be 3.08
instead of 14. Quite a number of specimens of the insect fauna of
Grand Cañon and the San Francisco Mountains of Arizona were
exhibited by Professor Townsend, besides some photographs taken
during the trip. ;
Dec. 1, 1892.—(No. 1.) Professor Townsend read a paper entitled
“ Notes on the occurrence of the puma (Felis concolor L.) in Southern
New Mexico.” The animal was reported on good authority as toler-
ably common in the Organ Mountains, and still more so farther east on
the Pefiasco River in Lincoln County. It has killed colts in Soledad
Cañon (Organ Mts.). Measurements of some large skins are given,
one of more than eleven feet being vouched for on good authority.
(No. 2.) Notes on the Ferns of the Organ Mountains, by Professor
E. O. Wooton, following the distribution zones of ferns as advanced
by Dr. Underwood in “Our Native Ferns and Their Allies” (pages
61-65, edition of 1888), he enumerated the various species and genera
which had been collected on several trips to the mountains without any
_ idea of such paper in view. Whole number of Genera collected, 7;
312 The American Naturalist. [March,
whole number of Species collected, 16. He then noticed the general
characteristics to be: rigid stems, thick leaves, deeply-seated veins,
fronds either very hairy, chappy, granulated, or with very thick epi-
dermis. All of which characteristics were taken as being simply differ-.
ent vaat of protecting themselves from the intense dryness of the
aris 3) “ Notes on the Mound Builders,” by Professor J. P. Owen,
was a paper concerning what is known of that ancient race. The pro-
fessor showed, in several different ways, the difference between this race
and the North American Indian found in possession of this continent
at its discovery. He told of the migrations of this pre-historic race
from the south and west to the central:and northern states, and their
probable return to the sections from whence they came. He led us to
believe that these ancient peoples might have been connected with our
Pueblo Indians, and were connected with the Aztec tribes of Mexico. '
International Congress of Zoology.—The second meeting was
held at Moscow, August, 1892. The fellowing papers were read :—
General Questions Concerning Biology: Reply to questions proposed
by Professor L. Cosmovici: 1. On a definite division of the animal
kingdom into “ phyla.” 2. Natural basis of a system of the type of
worms. 3. Uniformity in the terminology of the secretory organs of '
worms, J. von Kennel.—Observations on some points in Zoological
Nomenclature, Ch. Girard.—On the importation and hybridization of :
reptiles and amphibians, J. de Bedriaga.
Special questions Concerning Biology :—Note on Parapaguris pilo-
simanus, a Pagurid from the abyss of the Atlantic Ocean, A. Milne
Edwards and E. L. Bouvier—Note on the fauna of the Black Sea, P.
N. Bontchinsky.—On a fresh-water Thuricola, F. Vejdovsk y.—Essay
on the Classification of Animal variations according to their causes, A.
Brandt.—The fauna of the eastern part of the Baltic Sea, and the
problems for the next investigations in that fauna, Gr. Kojevnikov.—
Geographical Distribution of the carnivora, Ch. Grévée.—The
European and Circum-Mediterranean vipers, J. de Bedriaga.— Chal-
cides symonyi Steind., and Molge luschiani Steind., J. de Bedriaga.—
Notes on the nests of insects made of clay, H. von Ihéring—The
Cetaceans of the Black Sea, living and fossil, J. van Beneden.—On
the geographical distribution of the Cladocera, T. Richard—On the
Monodontophrya, a new species of the Opalinide, Fr. Vejdovsky.
Histology and Embryology :—Contributions to the theory of the
pgs
mesoderm and of metamerism, N. Cholodovski—Note on the forma- . —
_ -1893.] Proceedings of Scientific Societies. 313
tion of the germ of the peripheral nervous system, A. Plitzine-—Note
on the development of the endothelium of the heart in the amphibians,
V. Roudnev.—On the development of Chrysopa perla, Mme. O. Tic-
homirova.—On the segmentation of the egg, and the formation of the
blastoderm of the Pseudoscorpionidex, Fr. Vejdovsky.—On an embry-
onic organ of the Pseudoscorpionide, Fr. Vejdovsky.—Contribution to
the history of the parasites of the Hymenoptera, N.|Koulaguine—The
use of embryological researches for classification, A. Tichomiroff.
Physiology and Physiological Chemistry :—On the phosphorescence
of animals in the Black Sea, P. Khvorostansky.
Morphology and Comparative Anatomy :—What is meant by “the
water vascular system, the segmental organs, the excretory organs, and
the nephridia, L. Cosmovici—Presence or absence of an excretory appa-
ratus in the genital organs of the Metazoa, H. von Ihering.—Note on
metameric signification of the cranial nerves, P. Mitrophanovy.—Posi-
tion of the Strepsipteride in the system according to the data of post-
embryonic development and of anatomy, N. Nassonov.—On the excre-
tory organs of terrestrial arthropods, A. O. Kovalevsky.—On the
origin and ot of the Arthropods, particularly the tracheated
arthropods, N raf.
Boston Society of Natural History.—January 18, 1893.—
The following paper was read: Report on a study of Glacial sand-
plains in Eastern Massachusetts, illustrated by lantern slides, by Pro-
fessor W. M. Davis and students in geological field-work in Harvard
University. SAMUEL HENSHAW, Secretary.
314 The American Naturalist. [Mareh,
SCIENTIFIC NEWS.
Recent Deaths.—Henry Tispors Srarnron.— Word comes from
England of the death of this eminent Lepidopterist on December 2d,
1892, in his 71st year. For the past fifty years he has been one of the
most active British entomologists. During this period he has been
president of the Entomological Society of London, secretary and vice-
president of the Linnean Society, and secretary of the Ray Society.
One of the founders of the Etomological Monthly Magazine, he con-
tinued on the editorial staff from its beginning until his death. From
this magazine we learn that he has published more than twenty-five
volumes on natural history, besides frequent contributions to entomo-
logical periodicals. He studied chiefly the Micro-Lepidoptera, and
was best known as a student of the Tineina. Two of his associates on
the Monthly Magazine, Messrs J. W. Douglas and R. McLachlan,
write: “ Naturally diffident and unobstrusive in society, he yet pursued
the objects that interested him, with ardor and perseverance, and his
liberality in the cause of the advancement of entomological studies in
Britain, which was always dear to him, and his unstinted aid in the
identification of species, are too well known to require eulogy. From
the first he restricted his researches to Lepidoptera, but he had sym-
pathy with the students of all orders of insects, and of natural history
generally. Possessed of an ample fortune, he used his means freely to
assist any cause or person that he deemed to be deserving . . . In
1871 he was instrumental in founding the Zoological Record Anoia
tion, for the purpose of continuing the Zoological Record, which had
been relinquished by Er. Van Voorst, and largely through his liberal-
ity this indispensable publication appeared regularly until 1886, when
it was taken up by the Zoological Society of London.”
FREDERICK Avueustus Gentu.—The death of Frederick Augustus
Genth occurred at his residence, No. 3937 Locust street, Philadelphia,
and ended, at the age of seventy-three, the career of a chemist and
mineralogist whose reputation was not confined to one hemisphere, but.
was co-extensive with the world of scientific investigation. Professor
Genth was born in Waechtersbach, Hesse-Cassel, on May 17, 1820.
After attending the Gymiiasium, in Hanau, he studied at the University
of Heidelberg, under Liebig; at Giessen, and finally under Bunsen, at
1893.] Scientific News. 315
Marburg, where he received the degree of Ph. D. in 1846. For three
years he acted as assistant to Professor Bunsen, and soon afterward came
to the United States, where he has since resided
In 1872 he was called to the chair of chemistry and mineralogy in
the University of Pennsylvania, which place he held up to within a few
years ago. He had also held the office of chemist to the Geological
Survey of Pennsylvania, and also to the Board of Agriculture of this
State. Professor Genth was a member of many scientific societies in
the United States, and was elected in 1872 to membership to the
National Academy of Sciences. He had no superior in this country
as an analytical chemist, and he greatly enriched the literature of
chemistry with his very many and careful analyses of minerals. His
name is associated with the ammonia cobalt bases, which he discovered
in 1846, and, in joint authorship with Dr. Wolcott Gibbs, he contributed
to the “Smithsonian Contributions to Knowledge,” a monograph on
“ Researches on the Ammonia Cobalt Bases” (Washington, 1856).
Professor Genth was the author of nearly one hundred separate
papers on subjects in chemistry and mineralogy, and published Tabel-
larische ubersicht der Uichtigsten Reactionen welche Basen in Salzen
zeigen (Marburg, 1845) ; also the same in relation to “ Acids” (1845) ;
his “ Minerals of North Carolina,” being appendix “ C ” of the “ Report
on the Geology of North Carolina” (Raleigh, 1875). He also was the
author of “ First and Second Preliminary Reports on the Mineralogy
of Pennsylvania” (Harrisburg, 1875-6) and “‘ Minerals and Mineral
Localities of North Carolina” (Raleigh, 1881).
Nrxouat Ivanovircn Koxsnarorr, the well-known Russian miner-
alogist, died at St. Petersburg, January 2, 1893. During his early
years he lectured on geology and physical geography, but later on
devoted himself to the description of Russian minerals of which he dis-
covered, and described many new ones. His chief works are embod-
ied ineleven large quarto volumes of Beiträge zur Mineralogie Russlands,
illustrated with numerous plates. The twelfth volume was in type
when he died. He was a member of the St. Petersburg Academy of
_ Sciences, and many of the scientific bodies of Western Europe elected
him corresponding or honorary member.
—The address delivered by Professor Virchow on his assumption of
the office of Rector of the University of Berlin, has been issued by the
ne
A
316 The American Naturalist. [March,
German publisher, August Hirschwald, of that city, under the title
“ Lernen und Forschen.” i
Unofficial information has been received by Professor G. Brown
Goode, of the National Museum, and member of the United States
Commission to the Madrid Exposition, that awards have been made to
the following exhibitors from this country :—
Gold medals — Smithsonian Institution, Bureau of Ethnology,
National Museum, University of Pennsylvania, William Ellery Curtis, 2
of Washington ; the Hemenway Expedition, Dr. J. Walter Fewkes, of —
Cambridge.
Silver Medals—United States Geological Survey, the Numismatic
and Antiquarian Society of Philadelphia, the Philadelphia Academy
of Natural Sciences, the Peabody Museum of Archeology, Cambridge;
The United States Mint, the United States Navy, The Army Medical
Museum, Dr. D. G. Brinton and Mr. Stewart Culin, of Philadelphia; —
Professor O. T. Mason, Professor Thomas Wilson, Mr. W. H. Holmes
and Mr. Walter Hough, of Washington; Mrs. Zelia Nuttall, of Cam-
bridge. |
Bronze Medals—Fish Commission, Department of Ayriculture;
Indian School at Carlisle, Pa.; Sons of the American Revolution, Dr.
James ©. Welling and Mrs. Tillie E. Stevenson, of Washington; Dr, —
Cyrus Adler, of New York ; Mr. Warren K. Moorehead, of Xenia, O.
The gold medals, it is understood, are very handsome, with an intrin-
sic value of $150 apiece. Final action by the jury is not known, and —
others may be honored. i
At the annual meeting of Bowdoin College Alumni held at Boston, —
Jan. 25th, Professor Robinson of the chemical department of —
Bowdoin, announced that Mr. Edward F. Searles, whose wife, the
widow of Mark Hopkins, left him millions, had promised to erect the
finest and best equipped building for the study of the sciences that
could be had in this country. He stated that Mr. Searles had placed —
no limit on the cost. oo
"
bi;
7
ie
4
i
Dr. Osaun has been appointed Geologist on the Geological Survey , —
of Texas, and will take charge at once of the petrographic work of the _
Survey. Dr. Osaun is well qualified for the position, having been for
many years first assistant to Professor Rosenbusch, in Heidelberg, and) _
later, extraordinary professor of mineralogy and petrography in that —
University. 2
1893.] Scientific News. 317
Dr. Karl Spruner von Merz, the author of “ Historisch Geograph-
ischer Hand Atlas,” died August 24th, 1892, at the age of 89. His
great work, commenced in 1837, was not completed until 1852. Spru-
ner was also the author of two school atlases of historical geography.
The University of Cape Town has been admitted to affiliation with
the University of Cambridge. This is the second of the Colonial
Universities thus affiliated, the other being that of New Zealand.
According to the Anatomischer Anzeiger, the Biological station at
Plön, under the direction of Dr. Otto Zaccharias, has accomplished
successful work during the past Summer. Of the species of animals
occurring in the Lake of Plön, there have been determined 20 fishes, 40
crustacea, 69 worms, 14 mollusca, and 74 protozoa. Of greatest inter-
est is the discovery in fresh water of many organisms that have hitherto
been supposed to occur only in the sea; and among these more espec-
ially may be noted certain genera of diatoms, rhizopods and worms.
An illustrated report on results is soon to be published,
—Dr. R. von Wetlstein has been appointed Professor of Botany in
the University of Prague, Bohemia.
—Mr. E. E. Prince, M. A., Professor of Zoology in St. Munigo’s
College, Glasgow, has been appointed Commissioner and General
Inspector of Fisheries in Canada.
The Indiana Academy of Sciences has issued its Proceedings for
1891, in the shape of a pamphlet of 176 pages. It contains not only
the proceedings of that year, but also a list of all papers read before
the Academy since its organization in 1885. The Academy has 121
active, 11 non-resident and 1 honorary members. Copies of the Pro-
ceedings may be had of the secretary, Amos W. Butler, Brookville,
Indiana.
Professor Bohumil Shimek has gone to Nicaragua to collect in the
interests of the State University of Iowa. He will pay especial atten-
tion to the invertebrata and the cryptogamous plants. `
Bulletin No. 40, of the U. S. National Museum, contains the fourth
of the bibliographies of American Naturalists, that of George N. Law-
rence by L. S. Foster, 121 titles are enumerated. A good steel por-
trait accompanies the Bulletin.
318 The American Naturalist. [Mareh,
New York, Feb. 3.—A broad smile illumined the folio face of
Caliph, the hippopotamus at the Zoo this afternoon. In the tank
adjoining his a baby had arrived at exactly 12.15 P. M., and Miss
Murphy, his mate, was its happy mother. Director Smith and his
keepers knew Tuesday that something was in the air, for Miss Murphy
was restless and excited. She is a nervous animal at all times, but
Thursday she paced her cage all day, and Thursday night did not lie
down once. The warning came just in,time, and a thick straw bedding
was given her, and the partition separating her from Fatima was put
up. The latter has the empty tank and the mother has the landing.
After the youngster came, Mr. Smith said the first thing it did was
to walk about shaking its tiny ears and giving little grunts. Murphy
followed wherever it went, and would, allow no one to approach within —
the railing, and so he didn’t attempt it. Caliph meanwhile found a
peek-hole in the boards, and viewed his progeny with a great deal of
interest. He refused to go into his tank, and last night slept within
touch of the mother and baby. i
The baby weighs about thirty pounds, being very small. Its color
is pink—a salmon hue—and its legs are so short as to be almost imper-
ceptible at first sight. It is lively and runs about a great deal, much
to the mother’s discomfort. It is a peculiarly shaped animal. It %
might be said that it has no shape at all. Itis about eighteen inches
long, has no tail, ears that can hardly be seen and mouth no larger
than a pint cup. The mother guards it with a great deal of care. It
is her third born. Four years ago one was born which weighed sixty- —
‘three pounds. It died. Fatima came next, having been born October
4, 1890. Murphy is 8 years old and cost $5000. She weighs between —
3000 and 4000 pounds. Caliph was brought from the Cincinnati Zoo —
and cost $5000. He weighs 7000 pounds. To-night Murphy and her —
baby were doing nicely.—Late paper. -
Table of Contents of the North American Review for
February, 1893.—How to Revise the Tariff, by the Hon. Wm. M. —
Springer, Chairman of the Ways and Means Committee; Recollections
of the Panama Canal Congress, Rear-Admiral Ammen, U. S. N.;
Changes in the Church of England, The Dean of St. Paul’s; Criminal
Law in France, Madame Adam; Boons and Banes of Free Coinage,
I. “In the Interest of Shylock, ” by the Hon. R. P. Bland, Chairman
of the Committee on Free Coinage, Etc., II. A Warning to Savings
Bank Depositors, by John Harsen Rhoades, President of the oa
1893.] Scientific News. 319
wich Savings Bank, N. Y., III. A Depositor’s Point of View, by a
Depositor in a Savings Bank ; Wild Stag Hunting in Devon and Som-
erset, The Countess of Malmesbury ; Government Aid to the Nicaragua
Canal, Senator John T. Morgan; Shall Our Laws be Codified ? Fred-
. eric R. Coudert; Needed Reforms in the Army, General John Gibbon,
U.S. A; Why Immigration Should Not be Suspended, Senator H. C.
Hansbrough ; The Hope of a Home, Erastus Wiman ; Europe at the
World’s Fair, I. The British Section, by Sir Henry Trueman Wood,
Secretary to the British Commission, IJ. The French Section, by
Theodore Stanton, Commissioner Resident in Paris; Notes and Com-
ments, Mistakes—but Not of Moses, Charles W. Trickett ; Science and
the Woman’s Question, Lydia Lvovna Pimenoff; From Renan’s Point
of View, Arthur Reed Kimball; The American Common Schools,
Rev. James A. King.
320 The American Naturalist. [Mareh,
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‘ Vol. XXVII. APRIL, 1893. ` No. 316-
CONTENTS.
THE RELATIONSHIPS ND DISTRIBUTION OF THE —Isomo
NORT ERICAN UNI Mouar WITH NOTES eenitab praia nn
> <ON nie wie COAST SPECIE
= PAGE. PAGE.
' THE GENEALOGY OF MAN. E DA New Rock Type, A stray aaa Petrography of —
z D. Cope. 32 ey ae = pan—Two Peculiar Rocks from Siberia ~
THE PROBABLE Eo OF me CRETA- | —An Ottrelite Basie parc AETS in Vermont—
_ CEOUS PLANT POPULATI | Lithapbiyes in the Rocche-Rosse—The Composi- ~~
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Ch E Da Macmillan’s rae re sre ager Tm Eggs El lectricity ond
rmz of the Minnesota ereiaro on eavage—Membranes of the Sea Urchin ae
omy of the Humming-Birds and. Experiment on Clea iy :
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sn O ERTS, wie - 364 | the Mouth Parts and Thorax of Insects and Chilo-
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PLATE IX.
t
U
iq
i) WSs
\
Bis ce
1-2. Man and Woman of Spy. 3. Maori. 4. Tahitian.
7. Esquimaux,
THE `
AMERICAN NATURALIST
Vor. XXVII. April, 1893. a 316
THE GENEALOGY OF MAN.
By E. D; Cops.
The ancestry of man is a question to be solved by paleon-
tology. Within the last twenty years, important progress has
been made in our knowledge of the phylogeny of the mam-
malia, and some points have been gained which throw consid-
erable light on the more immediate predecessors of man.
These additions to knowledge have been made in the fields of
both simian and human paleontology, and they have naturally
received attention from archeologists as well as paleontologists.
The question must be approached from the side of anatomy
primarily, although aid from any other source is essential.
Philology can, of course, give us no assistance, since the first
man can have left no trace of his language. It is a fact that
in some quarters, archeologists who are not anatomists, seem
to underrate the value of anatomy in the premises, and are
inclined to dispute the existence of men of very primitive or
| simian physical characteristics. But archeology, apart from
anatomy, is a poor guide in the field of human ancestry.
oN othing is better known than that some races of men present
a larger proportion of simian characteristics than others. Thus
these approximations are fewest in the Indo-European race,
and are rather more evident in the Mongolian. In the Poly-
nesian, Negro, and Bushmen, they are most numerous, and in
about equal b t differently distributed proporti Confining
22
322 The American Naturalist. (April,
our attention to the skeleton, it is known that almost every
part of it is the seat of variations of greater or less significance.
Much importance attaches to the dentition. Many years
ago, Owen’ called attention to the fact that in the dark races
the roots of the last superior molar are distinct from each
other, while in the Indo-Europeans they are known to be more
or less fused together. These now well-known characteristics
of human dentition constitute one of the examples of transi-
tion from a simian to a human type. I have pointed outa
corresponding modifigation in the structure of the crown of!
the superior true molars, viz: the transition from a quadri- ~
tubercular to a tritubercular structure in passing from the
lower to the higher races. As this point has some interesting
implications in the earlier phylogeny of man, and as its value |
has been disputed, I give it a little attention. |
The facts of the case are as follows: I have demonstrated’
the fact that all forms of dentition exhibited by the Eutherian |
Mammals have been derived from a primitive tritubereular :
type. Professor Osborn says that he expects to be able to do
the same for the Multituberculate (? Prototherian) dentition. |
I have also shown that man exhibits a tendency to revert
from his primitive quadritubercular molar to this tritubercular 3
type.’ These facts are now common knowledge among paleon-
|
tologists, although Dr. Brinton states in one of his late works
that the latter proposition has been “refuted ” by Dr. Harrison
Allen and Professor Virchow. As to the significance of these
facts, I have expressed the view that this acquisition of a tri-
tubercular molar is a reversion to the lemurine type. This
conclusion is necessary because the lemurs are the last of
the families in the line of the ancestry of man which pre-
sent this dentition. The monkeys and anthropoid apes are all
quadritubercular, except a few limited collateral branches of
the former, which still retain the lemurine type. There are i
also a few collateral types of lemurs which have acquired one
* Odontography, 1840-5, p. 454.
Amer. Philosoph. Soc., Dec, 1883; Origin of the Fittest, 1887, PP-
2 Proceeds.
245, 347, 359.
3 Amer, Journal of Morphology, II, 1888, p: 7.
1893.] The Genealogy of Man. 323
or more quadritubercular molars, but they are not typical. In
many tritubercular mammals, a precocious form or two can
be found, which has acquired the fourth tubercle. But the
further back we go in time, the fewer they become, until,
in the Puerco fauna, of eighty-two species of Eutherian Mam-
mals, but four have true quadritubercular superior molars.
I take this opportunity of saying, however, that reversion
is not necessarily a result of heredity. It may be simply a
retrogression on a line of advance already laid down. What
influence lemurine heredity may have had in the case of
man, it is not easy to know. But it must be borne in mind
that various forms of degeneracy of molar teeth are possible
other than the resumption of the tritubercular type, yet the
normal reduction generally presented, is just this lemurine
and primitive Eutherian condition. The simplicity of the
elements involved, has something, but not everything, to do
with this reversion.
Dr. Paul Topinard has made an investigation‘ of the char-
acters of the crowns of the molars in man, and has reached
general conclusions identical with my own. He remarks (p.
665), “ It-is demonstrated, in conclusion, that the teeth of man
are, at present, in process of transformation, and that in some
future which is remote, the inferior molars shall certainly be
quadricuspid, and the superior molars tricuspid. It will
be curious to have the statistics as to prehistoric man;
unfortunately, their crania are rare, and their molars gewil í
much worn.” In the details of his examination, there are some
divergencies from my results. Thus he finds the quadrituber-
cular second and third superior molar relatively of more fre-
quent occurrence in Europeans than I did. But the absence
of Europeo-Americans. from his tables reduces the percentage
of tritubercularsin the Indo-Europeans. He makes no report
of Esquimaux. Had he observed this type, he would have
found a higher per cent. of tritubercular upper molars than in
any race that he has recorded. He confirms my conclusion as
to the high percentage of quadritubercular superior molars in
the Malays, Polynesians and Melanesians.
* L’Anthropologie, 1892, p. 641 (Nov., Dec.).
`~ 324 The American Naturalist. [April,
The relation of this fact to phylogeny is to confirm Heeckel’s
hypothesis of the lemurine ancestry of man. I have advanced
the further hypothesis that the Anthropomorpha (which
include man and the anthropoid apes) have been derived
directly from the lemurs, without passing through the
monkeys proper. This close association of man with the apes,
is based on various considerations. One of them is that the
skeleton of the anthropoid apes more nearly resembles that of
man in the most important respects than it does that of the
monkeys. This is especially true of the vertebral column,
where the anapophyses are wanting in the Anthropomorpha
(insignificant rudiments remaining on one or two vertebre,
as pointed out by Mivart), while they are well developed in
the monkeys and lemurs. The frequent presence of the tri-
tubercular molar in man disposes of the superior claim of the
monkeys over the lemurs to the position of ancestor. Another
significant fact pointing in the same direction is the existence
of large-brained lemurs with a very anthropoid dentition
(Anaptomorphidae) in our Eocene beds. This resemblance is
very remarkable, much exceeding that lately observed by
Ameghino in certain extinct forms of monkeys in Patagonia,
which appear to be ancestors of the existing South American
monkeys (Cebidae), and possibly of the Old World monkeys
also. The superior molars of Anaptomorphus are trituber-
cular, while the premolars, canines, and incisors are essentially
: ` anthropomorphous, and rather human than simian. Anapto-
morphus is probably at the same time the ancestor of the
Malaysian lemurine genus Tarsius, and M. Topinard remarks
that Tarsius has as good claims to be regarded as ancestral to
Homo as Anaptomorphus. But M. Topinard must be aware
that in the existing genus the character of the canine and
incisive dentition is very unlike that of the Anaptomorphus
and Homo. It is specialized in a different direction. The
dentition of Anaptomorphus being so generalized as compared
with Tarsius, I suspect that its skeleton will be found to pre-
sent corresponding characters. Of course, if it be found here-
after to have the foot structure of Tarsius (which I do not
1893.] The Genealogy of Man. ‘B25
anticipate), it must be removed from the line of ancestry of
the Anthropomorpha.
It must be further observed that the ancestral line of the
Anthropomorpha cannot be traced through any existing type
of Lemuridae, but through the extinct forms of the Eocene
period.’ This is on account of the peculiar specialization of
the inferior canines, which are incisor-like, and because of the
peculiar character of the incisors themselves, in the modern
lemurs in the restricted sense. But we have numerous lemur-
ine types of the Eocene of both America and Europe which
satisfy the conditions exactly, so far as the dentition is con-
cerned. These are mostly referable to the family Adapidae..
Unfortunately, we do not know the entire skeletons of
these Eocene lemurs, but as far as we have them (genera Tom-
itherium and Adapis), they are monkey-like. But we have
what is almost as useful, the skeleton of their Eocene and
Puerco ancestors, the Condylarthra. I long since pointed out
that the latter order (not the genus Phenacodus, as Lydekker
has represented me as saying), must be the ancestors of the
lemurs, basing my views expressly on the general structure of
Phenacodus, Periptychus and Meniscotherium. The structure
of the ungual phalanges of Periptychus is very significant, and
even more so is that in Meniscotherium, as recently shown by
Marsh, who adopts (without credit) my hypothesis of lemurine
affinities of the Condylarthra (which he renames the Meso- .
dactyla). From Condylarthra back to Creodonta is an easy
transition, and I have always assumed that the Creodonta
were derived from generalized Polyprotodont Marsupialia.
This view has been entirely confirmed by the recent discover-
ies of Ameghino in Patagonia, where he has found forms
whose remains may be referred with equal propriety to the one
group or the other. M. Topinard has been rather hasty in
reaching the marsupial ancestry in supposing that Phenacodus
belongs to that order. All the evidence shows that Phenacodus
is a generalized ungulate placental. :
To return to the more immediate ancestry of man. M.
Topinard believes that I hold the opinion that the Hominidae
5 On the Primitive Types of the Orders of the Mammalia Educabilia, 1873, p. 8.
326 The American Naturalist. [April,
descended directly from the lemurs without the intervention
of the Simiidae. I have, on the conirary, expressed’, and now
maintain as a working hypothesis, that the Anthropomorpha
were descended from the Eocene Lemuroids. In my system!
the Anthropomorpha includes the two families Hominidae
and Simiidae. The sole difference between these families is
seen in the structure of the posterior foot ; the Simiidae hav-
ing the hallux opposable, while in the Hominidae the hallux
is not opposable. This is not a strong character, since it —
depends on a slight difference in the form of the entocunel- }
form bone. In some vertebrates, as the tree frogs, the same
and similar characters (genus Phyllomedusa) are notregarded =
as of family value. It is then highly probable that Homo is
|
|
:
descended from some form of the Anthropomorpha now extinct,
and probably unknown at present, although we do not yet
know all the characters of some extinct supposed Simiidae, of
which fragments only remain to us. It cannot now be deter-
mined whether man and the Simiidae were both descended
from a genus with opposable hallux, or without opposable
hallux, or whether from a genus presenting an intermediate
character in this respect. This genus was, in any case, distinct
from either of the two existing genera of Simiidae, Simia :
and Hylobates, since these present varied combinations of a
anthropoid resemblances and differences, of generic and speci-
fic value.
Professor Virchow has recently* thrown down the guage to
the evolutionary anthropologists by asserting that “ scientific
anthropology begins with living races,” adding “that the first
step in the construction of the doctrine of transformism willbe
the explanation of the way the human races have been ©
formed,” etc. These two assertions are inconsistent, since the
only way of solving the latter problem will be by the discovery
of the ancestral races, which are extinct. The ad captandum
remarks of the learned professor as to deriving an Aryan from
ê American Naturalist, 1885, p. 467.
- 7 Origin of the Fittest, 1887, p. 346, from pres Naturalist, 1885, p. 346, wh
the classification of the Taxeopoda should be in a foot-note; loc. cit., 1889, cull
* Popular Science Monthly, Jan. 1893, p. 373, translated.
1893.] The Genealogy of Man. 327
a Negro, etc., remind one of the criticisms directed at the doc-
trine of evolution when it was first presented to the public, as
to a horse never producing a cow, etc. It is well known to
Professor Virchow that human races present greater or less
approximations to the simian type in various respects. Such
are the flat codssified nasal bones of the Bushmen ; the quadri-
tubercular molars of the Polynesians; the flat ilia and prog-
nathous jaws of the Negro; the flat shin-bones of various
races; the divergent hallux of some aborigines of farther India,
etc. But that no nearer approximation to the simian type
than these has been discovered in any race will surprise most
students of the subject. Professor Virchow does not satisfac-
torily explain the significance of the Cannstadt and Neander-
thal skulls, and in his address he ignores the important and
well authenticated discovery of the man and woman of Spy.
To these ancient people I will now devote some space, describ-
ing more especially the dentition so far as it has been pre-
served.
What had been long suspected is now established, through
the discovery and descriptions of MM. Lohest and Fraipont of
Fic. 1—SKULL OF THE MAN OF Spy. From Professor G. F. Wright’s Man
and the Glacial Period. From a photograph.
328 The American Naturalist. [April,
Liége, viz.: that there dwelt in Europe during paleolithic times,
a race of men which possessed a greater number of simioid
characteristics than any which has been discovered elsewhere.
The important discovery in the grotto of Spy, of two skeletons,
almost complete, served to unify knowledge of this race, which
had previously rested on isolated fragments only. These skel-
etons proved what had been previously only surmised, that the —
;
7
skeleton of Neanderthal, the lower jaw of Naulette, and the
crania of Cannstadt, belong to one and the same race. The
simian characters of these parts of the skeleton are well known.
These are the enormous superciliary ridges and glabella; the
retreating frontal region; the thickness of the cranial wall;
e massive mandibular ramus with rudimentary chin,and the —
large size of the posterior molars. MM. Fraipont and Lohest —
have added other very important characters to these, viz: the —
tibia shorter than in any other known human race; the sig-
moid flexure of the femur; the divergent curvature of the
bones of the fore-arm, and a very peculiar form of the posterior —
face of the mandibular symphysis. Of these characters Ma
-mm mm m
- —
~-
RS eae a ett
i EE Se cores 5
.
s
2 o ae.
oo weet
Fic. 2.—Outlines of calvaria of the Neanderthal man in solid line; of the Spy man *
no. Lin dots; and of the Spy man No. 2 in broken line. From Fraipont and Lohest.
oe and Lohest write as follows: :
I.—The prominent superciliary crests, which are character-
istic of the Cannstadt race. No existing race presents such 4
’ Archives Belges de Biologie, VII, 1886, p. 731, Gand.
1893.] - The Genealogy of Man. 329
development, neither the Papuans, Australians, nor Negroes of
any race. But we find the superciliary crests and underlying
sinuses identical in adult female orangs and chimpanzees and
young male gorillas. In the female chimpanzees the crests are
almost inferior in size to those of the man of Spy.
I1.—The retreating forehead of the two crania of Spy, is not
found in any existing human race, while it is typical of that of
Cannstadt. It is characteristic of female orangs and gorillas
and of the young males of both species, and of adult males and
female chimpanzees. It appears in existing men in rare and
isolated cases.
III. —The prominent transverse superior semicircular crest
of the occipital bone is found in existing races among the
Fellahs of Africa and the Nigritos. It is characteristic of the
Cannstadt skulls, and presents exactly the same characters as
the young male and female orang and gorilla, and young male
and adult female of the chimpanzee.
IV.—No human race presents the characters of the lower jaw
exhibited by those of Spy, Naulette and Shipka. In this part
of their osteology the anthropoids depart widely from man, the
most conspicuous point in the latter being the presence of a
chin. Accordingly, the angle formed by the anterior face of
Fie. 3. ; Fic. 4.
Vertical sections of symphysis mandibuli of gorilla (Fig. 3), and orang (Fig. 4).
From Fraipont and Lohest.
330 The American Naturalist. [ April,
the symphysis with the inferior border of the horizontal ramus,
is less than a right angle in man, and much more than a right
angle in the anthropoids. According to Topinard, this angle in
fifteen Parisians is 71°.4; in fifteen African Negroes, 82°.2; in
fifteen Neocaledonians, $3°.9; in the jaw of Naulette, 94°. In
the best preserved jaw of Spy, the angle-is 107°, if measured
from the inferior symphysial border, or 90° if measured from
the inferior border of the ramus. There is no chin in the jaw
of the Cannstadt race, and the large angle approaches without
nearly equaling that of the anthropoids. But the posterior
face of the symphysis presents the most remarkable peculiarity.
In the symphysis of the apes (Figs. 3, 4,5) the posterior border
is a continuous slope from the alveolar border to the inferior
K
Fic. 5. Fic. 6. Fic. 7.
Vertical sections of symphysis mandibuli of chimpanzee (Fig. 5) ; of Spy man No.
1 (Fig. 6), and Spy man No. 2. ( Fig. 7). From Fraipont and Lohest.
margin, interrupted by a slight concavity below the middle.
In the human jaw this line slopes backward to near the middle,
where are situated the small tuberosities for the insertion of the
genio-glossal muscles. (B in the accompanying figures.) The
surface then slopes rapidly forward to pass into the narrow
inferior border of the chin (Figs. 8, 9). In the jaws Naulette
and Spy, the structure is exactly intermediate between the two,
and quite different from both. It commences above with a 2
posterior slope similar to that of the apes, exhibiting what is
called by Topinard, internal prognathism, as it appears in the
lower human races. The surface then descends abruptly, form-
1893.] The Genealogy of Man. 331
ing a vertical concavity, which is bounded a considerable
distance below by another protuberance, the insertion of the
genio-glossal muscles. This concavity is not present in the
human symphysis, while it is less developed in the simian.
The surface then slopes forward, as in the human symphysis,
but this portion is shorter than in human jaws generally. It
is represented by a convex face in the simian jaw. This
character taken in connection with the others cited, goes a
long way toward justifying the separation of the Cannstadt
race as a distant species, as has been done by some author
under the name of Homo neanderthalensis (Figs. 6, 7).
Fic. 8. Fic. 9
Sections of symphysis mandibuli of modern Liegois (Fig. 8), and of an ancient Par-
isian (Fig. 9). From Fraipont and Lohest.
V.—The curvature of the ulna and radius, which produces a
wide interosseous space, is not found in any human race, but is
common to the apes. On the contrary, the shortness of these
bones is entirely human.
VI.—The anterior convexity of the femur, with its round
section, is only found among living races among the Nigritos
of the Philippine Islands. It isseen in a less degree in femora
of neolithic men, and occasional instances are seen among exist-
ing Europeans. It is the normal condition in the apes.
VII.—The tibia is shorter in its relation to the femur than
in any human race, and is more robust than in most of them.
This character, with the oval section, while not identical with
what is seen in the apes, forms an approximation to it.
VIII.—The condyles of the femur are not only wider than
in human races generally, but they are more produced posteri-
orly. The portion habitually resting on the head of the tibia
332 The American Naturalist. [April,
is so far posterior as to throw the femur backward when in a
place. This requires that in order to maintain equilibrium,
the head and body must have been thrown forward. This
relation of the femur and tibia is what we find in the apes,
and it is highly probable that the man of Spy presented a
somewhat similar figure when walking. That is, the knees
were bent and the body thrown forward. In the best preserved
leg, the knee was also slightly bent inward. The femur had,
however, the superior length generally characteristic of man, A
though this, it is well known, is a variable character. l
MM. Fraipont and Lohest have pointed out the genera
characters of the dentition of the man of Spy. They show that
the molars increase in size posteriorly to the same extent that
they do in the ‘apes, which is the reverse of what is usual in
man, where they diminish posteriorly, or, in a few lower races
(Australians, ete.), remain equal. They show that the superior —
molars are all quadritubercular, and that the internal root is
distinct in all of them. Through the kindness of M. Lohest,I
received casts and photographs of these teeth, and I give here
figures the former (Plate IX), which are more satisfactory
than those in the memoir from which I have already quoted
so fully, where, indeed, the grinding faces are not represented
at all. I also make some remarks which they suggest, on some
details not noticed by MM. Fraipont and Lohest. ve
The casts of the teeth of the man are those of P.m. 2 and M. |
= of one side, and of M. *** of the other side. Those of the —
woman are alveoli of I. * and C. +, and the entire teeth of P.
m. ` and M. =+ all of one, the right, side. The crowns of
the man’s teeth are moderately worn, so that their composition
can be studied, while those of the woman are more worn, 80
that the composition is somewhat obscure. Comprehensively,
it may be stated that the true molars are fully quadrituber
cular, as in the lowest known races of men, and that the
hypocone is in the anterior molars, as large as the protocone,
while it is always smaller in man when present. In the female
the last superior molar has a well-developed internal root, which —
diverges widely from the posterior external, even more widely
than in a chimpanzee in my possession. This arrangement!
1893.] The Genealogy of Man. 335
even more simian than that seen in Australian and Maori denti-
tions in my collection. The last molar of the male is stated
to have similar structure, but the roots cannot be clearly seen in
the plaster cast. These two characters then place the man of
Spy i in the lowest category, just as would be naturally expected
in the presence of the other simian characters already referred
to. The dentition is inferior to that of the neolithic man of
France,” where the last molar at least is tritubercular.
Some other peculiarities of these dental series may be now
referred to. In the female dentition, where alone the canine
alveoli remain, the size of the latter tooth and that of the
premolar is large, relatively, to that of the true molars. The
canine is not remarkably large, but exceeded by a little, that
commonly seen in Polynesians and Australians, where it is
largest among human races. The premolars exceed a little
the proportions seen in the same races, approximating but not
quite equaling the relative dimensions seen in the chimpanzee.
The roots are relatively more robust in the transverse direction
than in human dentitions generally. The first and second
molars of the female are of equal transverse diameters, while
the third is a little wider transversely and quite as wide
anteroposteriorly. This large dimension for the third molar
exceeds anything I have observed in human dentitions except
in the case of a single Australian in my collection, surpassing
a little that of the chimpanzee, and reminding one of some of
the gibbons (Hylobates), where, however, the crown is subcir-
cular in outline. In the male the transverse diameter of the
third molar is a little smaller than that of the other molars,
but the tooth is larger than is usual in man, and equals in its
dimensions that of the chimpanzee.
A peculiarity which is pronounced in the male, and observ-
able in the first molar only of the female, is the prolonged pos-
terior convexity of the grinding surface. This is unlike any-
ng I have seen in human molars, and resembles the char-
acter of the orang" more than that of any other quadrumane
known to me. This appearance is increased by a contrac-
“1 Cope loc. ĉit., p. 17.
‘N Owen Odontography, p. 444.
334 The American Naturalist. (April,
tion of the grinding surface posteriorly, by the convergence
of the internal and external borders, a peculiarity apparently
unique to the man of Spy. The forward overlap of inferior
molars on the superior is much greater than usual in the male.
This contraction of the posterior part of the external border is.
equally marked in the female, but the posterior border is not so
convex. It results from the peculiar form of the external borders
of the crowns of thetrue molars in both sexes, that they form an
echelon arrangement. This does not seem to be an adaptation
to an oblique outline of the maxillary arch, which is broadly
rounded as in the Australians and Polynesians.
Summarizing the above observations, I would say that in
the quadritubercular molars, with distinct and divergent roots,
the man of Spy takes the lowest position among the subspecies
of man. Second, that in the relative proportions of the pre-
molars and molars with each other and among themselves, he
possesses some characters lower than any of the known races of
man, and approximates near to the apes. Third, that in the
echelon arrangement of the external wall of the molars, he
presents a peculiarity more strongly pronounced than either
man or apes, and whose true significance can not now be deter-
mined. It occurs in the abnormal V-shaped maxillary arch of
existing men, but the arch in the man of Spy has not this
character, but is fully rounded, and as it occurs in both sexes,
it is not probably an abnormality.
In conclusion, it may be observed that we have in the Homo
neanderthalensis a greater number of simian characteristics
than exist in any of the known races of the Homo sapiens,
although, so far as known, he belongs to the genus Homo. The
posterior foot, so far as preserved, indicates this to be the case.
The foot-character, which distinguishes the genera Homo and
and Simia still remains. There is still, to use the language of
Fraipont and Lohest, “an abyss” between the man of Spy and
the highest ape; though, from a zoological point of view, it is o
not a wide one. |
The flints which were discovered in the stratum of cave
deposit containing the human remains, are of the paleolithic
type known as Mousterien in France, which are of later origin
1393.] The Genealogy of Man. 335
than the Chelléen or older paleolithic. The older paleolithic
man isnot yet known. It is interesting to observe that these
flints (Mousterien) are of the same form as the obsidian imple-
ments which I collected at Fossil Lake, in Oregon, with the
bones of extinct llamas, horses, elephants, sloth, etc. The ani-
mals which accompanied the man of Spy are, Cælodonta anti-
quitatis (wooly rhinoceros), Equus caballus, Cervus elaphus, Cervus
tarandus, Bos primigenius, Elephas primigenius, Ursus spelius,
Meles taxus, Hyaena spelæa ; five extinct and four existing
species.
As the evidence now stands, the most primitive and simian
of human races inhabited Europe. No trace of the Homo
neanderthalensis has been found in any other region of the
earth. As, however, paleolithic implements are found in all
continents, we may anticipate that this or some similar species
of man will be discovered there also. An equally probable
conclusion is that, taking into consideration the characters of
the Neolithic man, the Europeans originated in Europe, and
that some of us are the direct descendants of the Homo nean-
derthalensis.
EXPLANATION OF PLATE.
Dentition of the man and woman of Spy, natural size; taken
from casts.
Fic. 1. Superior molars of the man from below.
Fic. 2. Do. of the woman from below; 2a do. last posterior
molar from behind.
Fic. 3. Maori, last superior molar from behind.
Fie. 4. Do. of Tahitian from below ; a do. from behind, show-
ing fused roots.
Fia. 5. Superior dentition of a Fan from West Africa, from
the collection of the University of Pennsylvania; a last super-
ior molar.
Fig. 6. Inferior dentition of Fan of Fig. 5.
Fig. 7. Esquimaux, superior molars; from cranium in Army
Med. Museum, Washington, D. C.
336 The American Naturalist. [April, >
THE PROBABLE PHYSIOGNOMY OF THE CRETA-
CEOUS PLANT POPULATION.
By Conway MAcMILLAN.
The sudden appearance of metaspermic plants in the rocks
of the Cretaceous period has occasioned much discussion, and
has always proved to be a crux to the writers on evolutional
philosophy. Even in the latest work of authority on general
evolutionary topies—Darwin, and after Darwin, by Romanes, the
„old and vague statements—concerning “probable breaks in
the records,” and possible remarkable accelerations of what
are usually very slow processes—may be found again, and
have the air of being all that can yet be said with much show
of reason from the evolutional point of view. The various
opponents of Darwinism, and of evolution in its broader sense,
have ever appealed to the sudden emergence of Cretaceous
metaspermic plants as an unanswerable argument against the
different development-hypotheses. There is not space here —
even to cite the numerous works in which the unexplained
and sudden appearance of Cretaceous metaspermic plants has
been adduced in condemnation of the various evolutionary
hypotheses. All students who have kept pace with the con-
troversial biological literature of the last thirty years, will
readily recall the important part that the Cretaceous plants
have played in the anti-evolution polemic.
In view of what has been stated above, the intent of this
paper will be recognized as a sufficiently ambitious one, when
it is stated to be an effort to show two facts, as developed by
the study of modern plant-formations and the evidence of the
fossils ; first, that the appearance of Cretaceous metaspermic
plants is proved, by the fossils, not to have been sudden, but
ual, and consequently, in Cretaceous time, the general
preponderance of plant-population was strongly coniferous,
fern and cycadean; and second, that the conditions of Creta~
ann eee ee
ee E aes ae = Mem E EAEE tomate
I Cee cre: ae Oe eT rs
ceous time were such that the new and scattered metaspermi¢ a
plants were placed under circumstances similar to those in
NS an ERRE ane De
ee ar eee ee a a =
oe pipes
1893.) Cretaceous Plant Population. 337
which to-day variation is most rapid and plasticity is greatest
for each species and even for every individual.
The attempt to show that the high preponderance of metas-
permic remains in the Cretaceous rocks is a sufficient proof of
the high preponderance of pines, cycads, ferns and club-mosses
in the Cretaceous forests is paradoxical enough to excite grave
suspicions of the method by which such a result is reached,
but the writer hopes to show that the absurdity of the state-
ment is solely a superficial one, and that it is really capable of
a defence both from experiment, observation and legitimate
deduction. I have already presented in my recent work on
the plants of the Minnesota valley the skeleton of the argu-
ment, but desire here to amplify and extend the discussion,
somewhat, to more fully cover the ground. If the conclusions
that I have drawn are legitimate, and my knowledge of the
literature of the subject sufficient to uphold my belief that
certain new laws are determined, both of plant-distribution
to-day and paleontological interpretation, it will be clear that
the argument is an important one, and should be examined for
what it may prove to be worth.
In the first place let us examine some of the observed facts
of plant-distribution to-day. It is apparent that while species
themselves are generally limited in some more or less definite
manner and occupy a more or less considerable area of the
earth’s crust, there are several degrees of aggregation in which
species play asubordinate part. We note the coniferous forest
as distinct from the hardwood forest, the prairie knoll from
the prairie swale, the mountain-side herbage and shrubbery
from the herbage and shrubbery of the valley. The presence
of these different zonal groups at different attitudes and lati-
tudes was first pointed out in the modern way, by von Hum-
boldt2 Since his time, the various groups of species, condi-
tioned in some definite manner and, under climatic influences,
acquiring characteristic habits or habitats, have been studied
1Metaspermz of the Minnesota Valley, p. 602 (1892).
- 2Essai sur la Geographie des Plantes ,suivi d'un Tableau Physique des Regions
De Distributione Geographica Plantarum, secundum cceli Temperium et altitudi-
nem montium, Prolegomena
23
*
338 The American Naturalist. [April,
somewhat in detail. The great works of DeCandolle®, Grise- l
bach‘, Engler’, Drude’, and the numerous special brochures of
these and of many other writers on geographical botany, have
pointed out with much distinctness the existence of plant-for-
mations. These formations are of different degrees. In the
United States there are, in the eastern portion, two very distinct
formations that belong to the highest division—namely, the
forest and the prairie. These two great formations, in the
broadest sense, are scarcely anywhere in contact with each
other, but are separated by what may be a very narrow strip
of plants imperfectly developed in either forest or prairie.
The region occupied by such a transitional strip of plants I
have named in the work cited above, a tension-line. Wherever
two formations of different sorts, such as hardwood forest and
coniferous forest, meadow and hillside-grassland, or any other
two, come together, there is such a tension-line developed along
the line of contact.
The character of the tension-line between formations has
received as yet very little study, and it is not possible to speak
of it as exactly as one would wish. My own observations are
confined to the tension-lines between forest and prairie in
the Minnesota valley, a region of special study for the past two’
years. Certain interesting facts have been deduced. These
may be noted as follows: .
The plants of the tension-line consist of such species as are `
sparsely developed in the more central areas of the two adja-
cent formations, or in some cases of species very poorly devel-
oped in the solid parts of the two formations,
The number of individuals of the first group is generally
greater over a given area along the tension-line than in the
solid portions of the formations.
*Géographie Botanique Raisonnée, on exposition des Faits Principaux et des Lois
papers la Distribution Geographique des Plantes de Pepoque actuelle. Tom. I,
‘La Vegetation du Globe, d’apres sa Disposition suivant les Climats, esquisse d’une
Geographie Comparée des Plantes, trad. par P. de Tchihatchef. Tom. I, II. (1877).
1893.] Cretaceous Plant Population. 339
There is a tendency for plants of a higher phylogenetic type
to accumulate in the tension-line area than characterizes
either of the adjacent formations. Thus in the border of the
wood and prairie one finds Asters, Helianthi, Solidagos, pre-
dominant over the lower Archichlamydeae or Monocotyledones.
There is a transition from typically prairie groups of grasses
to the typical tension-line groups of species, and similarly from
the typically forest groups of trees, in the hardwood belt.
The tension line is characterized by a greater comparative
richness of species and luxuriance of individuals, in the mat-
ter of number, than the average of the adjacent formations.
The tension-line is not a fixed, but a moving line, and
usually one formation or the other is in a condition of
encroachment while its adjacent formation is in a condition
of retreat.
The movement of the tension-line is felt some distance from
the contact line of the two formations, so that, if forest is
encroaching upon prairie, the characteristic composites of the
tension-line are thrust out farther into the prairie.
From the interior regions of both adjacent formations plants
“work out” into the tension-line area. Such plants are gen-
erally the imperfectly established.
The last point lends itself very easily to direct experiment.
If a field of some well established perennial plant, as for -
instance, blue-grass (Poa pratensis) be selected, and several
sunflower seeds, or thistle seeds, be planted in the center of it,
it will happen that one or two plants of the composite species
may reach maturity, and will then distribute seed outward in
every direction. In three years time the composites will prob-
ably be ejected to the periphery of the field, and will have `
gained there a considerable foothold, while in the center of the
field they will have gained but a slight foothold, and perhaps
will not have retained what they had given them at first. A
strong formation thus ejects to its periphery the plants that
not in harmony with it, still strive to establish themselves. In
the work cited above, I have termed this “the law of the ejec-
tion of the weaker.” It may be seen operating in any wood or
field, and is, I believe, one of the most important factors in the
340 The American Naturalist. [April,
origin and modification of metaspermic species—although not
generally recognized. It will be observed that while this ejec-
tion is purely biological, it nevertheless bears comparison with
certain physical phenomena as when, for example, the water
contained between the pores of adjacent solids is squeezed to
the surface of the mass, and is thus ejected from its original,
more central position. :
It appears from the preceding analysis of conditions certainly
existing and verifiable, that two adjacent formations do not
actually come in contact, but are separated by the thicker or
thinner band of tension-line species. It further appears that
the tension-line group is made up of, and constantly replen-
ished by plants ejected from the interiors of both formations,
and reaching the tension-line as it were, to use a metaphor
from chemical science, “in the nascent condition.” This
explains, or at least suggests the explanation of what other-
wise would be difficult, viz., the large comparative number of
species in the tension-line area, and the fact that higher types
are aggregated there rather than in the solid portions of adjacent
formations. For not only is the tension-area populated from the
interiors of the adjacent formations, and thus comes to have a
diversified group of inhabitants injected into it, but these
inhabitants are likely to be the new forms, not in perfect har-
mony with the general mass of the formation in which they
originated—and therefore they are ejected. Furthermore
plants reach the tension-area “on the move ”—their variations
already begun and accentuated, it may be, by the forced jour-
ney to the periphery of the parent formation. Therefore both
the actual numbers of species in the tension-area being great,
and these species being derived from widely different habitat
and environment conditions, together with the high degree of
plasticity acquired in the ejection and after establishment in
the tension-area, it becomes evident that it is precisely at such
a contact-region between two formations that conditions are
highly favorable for specific and generic and eyen ordinal
variations to set in. Thus the increased number of species in
the tension-area is seen to be in part the cause, and in part the
;
i
1893.] Cretaceous Plant Population. 341
result, of the high variability that is experimentally noted in
such an area.
From the above, the explanation of the various peculiarities
of the tension-line will readily be derived. Under the law of
the struggle for existence, it will be seen that this struggle may
be between individual and individual, individual and forma-
tion, and formation and formation. All three phases are
developed at the tension-line and since the causes of evolu-
tion are working at their highest along this area, it happens
that the results are numerically most apparent. The three
phases of the struggle are not elsewhere so vividly brought to
one’s notice, for at least the third will be wanting or felt but
indirectly. It appears, as a result of the examination, that the
character of the tension-line, as made out by observation, is
capable of illumination by experiment and of explanation by
the general and well-known laws of evolution.
It is now my intention to show that the metaspermic plant-
population of the Cretaceous, and especially of the Upper
Cretaceous was a tension-line population. If this be possible,
we shall expect that population to manifest tension-line char-
acteristics.
In the first place it is necessary to gain some comprehension
of the Cretaceous geography. I shall concern myself here only
with the Upper Cretaceous of America. At the time of the
deposition of the Cretaceous clays and sandstones, the North
American continent was much more restricted than now. It
extended little south of Illinois, and was cut by a great Med-
iterranean Sea into an eastern, larger and shorter, and a west-
tern, smaller and longer continent. The Mediterranean Sea
extended from the south to the extreme north, and was doubt-
less highly saline in character. The coasts of this sea were
generally low—at least this is derived from the evidence of the
flora, according to Lesquereaux—but, here and there, doubt-
less varied by some rocky promontory or headland. The
shores were marked by frequent bays and estuaries, and the
rivers which flowed into the Cretaceous oceans were probably
tide-streams to a marked degree. The general character of
the country was low or morassic, but this was certainly broken
342 The American Naturalist. [April,
by high uplifts, usually far inland. The coast-line of the
ocean was therefore, so far as may be determined up to the
present time, a generally low area, where vast stretches of sands
or mud-flats were shifted about by the surf and by the inflow-
ing waters of tributary streams. It is these sandy beaches
_and mud-flats that persist to-day as the Cretaceous rocks, bear-
ing their countless imprints of archaic plants.
Considering next, for a moment, the character of the plant
remains from the Cretaceous, we find that in the Lower Creta-
ceous, as studied by Fontaine, there are numerous ancient fern ©
and cycadean types. These become less prominent in the Upper
Cretaceous, so that in 1874, when, under the researches of
Heer and Lesquereaux, already 130 species were described
from the Dakota group, of these only five are ferns or crypto-
gamous plants, one doubtfully cycadean, six coniferous, two
monocotyledonous, and the rest Dicotyledons—principally in
the Archichlamydez, with only a very few ( Bicornes, Legu-
minosæ, Ebenacex, ete.) referable’io the Metachlamydexe. The
presence of the older types of cycads, conifers and ferns is
important to be noted, for it is possible that the sole reason for
their not appearing in countless numbers is the fact that they
were cut off from the sandy-beaches and mud-flats by a thin
but well-nigh continuous tension-line group of new and strug-
gling Metasperme.
The study of shore-lines of modern seas is essential to the
further working out of the hypothesis in process of develop-
ment in this paper. The writer is not aware that observations
have been made with the considerations in view that would
make them useful either for support or objection. His own
studies of sea-shore areas are confined to the observation of
Cape Cod, Massachusetts, during a single month. While there
he found that the older plants are more numerous in the inner
regions, and the newer more numerous nearer the coast. The
_ coast-line then seems to be a tension-area. While there are
many pines on the Cape, these, so far as I could discover, were
generally distributed inland, reaching the border of the sea
only on high promontories, and on low lands being generally
separated from it by strips of herbage, tov shrubbery and hard-
pS oe se
1893.] Cretaceous Plant Population. 343
wood timber. Further study of the sea-shore is needed along
this line. But, apart from the testimony of the limited obser-
vation, it appears demonstrable that the coast-line, simply
because it was peripheral, thus terminating ejections, must have
developed a tension-line group of plants in Cretaceous times.
That. this tension-line was a relatively narrow one is proved
by the great preponderance of metaspermic remains. For if it
had been a broad one, the competition between older ferns and
cycads, and newer metaspermic plants would doubtless have
been sufficient to have brought down to the coast in larger
numbers the more inland types of plants.
The picture of the Cretaceous plant physiognomy which
these considerations brings before our eyes is of a great solid
inland forest of conifers, cycads, ferns, and Calamites (?), bounded
by a thin peripheral line of metaspermic trees, shrubs and
herbs. This peripheral line extended up both sides of all
estuaries, streams and rivers, and thus shut off the older, luxu-
riant and dominant flora from proper representation in the
rocks of the period, as they have come down to us. An occa-
sional inland plant was carried down some river and buried in
the sands or shales at 1ts mouth, and sometimes when the shores
were high the inland types came close to the sea, and in the
Sequoias, Cunninghamites, A bietites, Zamieites and other genera
of the Cretaceous rocks, we derive a hint of the inland and
general character of the plant population. The very fact,
therefore, of the high metaspermic preponderance in the fos-
sils—since it shows metaspermic plants to have been uniformly
the coast population, and hence the tension-line population, is
an argument in favor of supposing that metaspermic plants
were actually very much less predominant in the general flora
then than they were in Tertiary or Recent time.
Together with this disposition of the oft-repeated declaration
that the Cretaceous Metasperme appeared with inexplicable
suddenness, it will be seen that the other difficulty of their
rapid and inordinate multiplication of species and genera is
solyed—so far as such notes constitute a solution. The tension-
line position, with its tremendous increase of urgency to mod-
ify, and its concomitant increase of individual plasticity,
344 The American Naturalist. [April,
explains the matter, so far as general explanations are of
value. As the Metasperme began to fight their way back into
the solid coniferous fern and cycadean forest, doubtless the same
phenomena might have been noticed that are apparent to-day.
The examination of a region where hardwood forest is encroach-
ing upon coniferous is especially suggestive. The forerunners
of the hardwood trees, various herbs and shrubs, insinuate
themselves up the rivers and tributaries, closely followed by
the trees themselves. In this way the coniferous plants and
their numerous allies, the various heaths, cornels, club-mosses,
etc., are isolated into phalanxes of smaller and smaller size.
At last they become entirely surrounded by the invading
types, and at the first opportunity are exterminated altogether.
Without question similar scenes might have been noticed in
the ancient Cretaceous forest when the tension-line group of
plants waxed strong and began the backward march carried
on through Tertiary time and resulting in the domination to-
day of archichlamydeous trees in the extra-tropical forest
regions, while metachlamydeous plants are in their turn
characteristic of the tension-lines.
To summarize, in conclusion, what is at best a disconnected
and imperfect attempt to contribute somewhat toward the
analysis of a very difficult group of problems. A study of
modern plant formations reveals the presence of tension-line
areas in which higher types of plants may become aggregated,
and where the general tendencies are toward a high degree of
individual and specific variability. Such tension-lines are
peripheral in position, and may be of different degrees of
organization as they lie around formations of greater or less
degree of integration. Newer types of plants are particularly
likely to be forced into the tension-line habitat, under the law
of the ejection of the weaker. Both observation and experi-
ment show this. Such tension-lines doubtless characterized
the Cretaceous formations as well as those of to-day. In them
the weaker, because newer, types were aggregated, and in this
way the littoral regions of the Cretaceous oceans came in all
cases to be densely metaspermic in the character of their
plant-population. The generally low elevation of the coasts
:
ir
K
¥
;
2
:
T
Se cere E E
1893,] Cretaceous Plant Population. 345
contributed to this condition. The older types of plants,
making up the bulk of the forest, were isolated from the shores
and from the banks of streams, except very far inland. Con-
sequently their remains are preserved only under exceptional
conditions. That the metaspermic plants occupied the
tension-line position indicates the preponderance of older types
in the great inland stretches of country. The rapid appear-
ance of new metaspermic types during the Cretaceous—and
probably too in Jurassic time—was a result of the tension-line
position into which they had been forced. The high develop-
ment of the arboreal habit may be referred to similar causes.
The picture thus brought before the eyes differs much from
the usual reconstructions of Cretaceous landscapes, but may be,
nevertheless, a closer approximation to the truth. A final
word may be added in regard to the conditions of the Triassic
and even of earlier time. In those ages we may imagine the
metaspermic types developing in the ancient forests of cycads
—or even of Lepidodendrons and Annularias—and beginning
their forced marches to the peripheral regions of the solid,
archaic formations of their birth. Thus for century upon
century they may have existed, and fail to appear in the fos-
sils, because of their internal position in the formations. But
when, after their long struggle for food and light and soil for
support, they emerge upon the periphery of their parent for-
mations we find them already arborescent or arboreal, and soon
almost universal upon littoral areas. They then first come
into a position where preservation in the rocks is possible.
346 The American Naturalist. [ April,
AN EXTREME CASE OF PARASITISM:
By Ropert Hessier, A. B., M. D.
There is a small family of minute animals belonging to one
of the sub-divisions of the arthropods which is of especial inter-
est on account of the parasitic habits of many of its members.
One of these, Sarcoptes scabiei De G., is parasitic on the genus
Homo.
Historicat.—It seems that the Jewish physicians dwelling
among the Moors of Spain, in the 12th century, knew some-
thing about these animals and their causative relation to the
affliction known as the itch. The discovery of the microscope
early in the 17th century (1619) led to many investigations,
some of which had reference to the itch mites. The first good
description of the mite bears the date of 1687, and was written
‘by Bonomo to Redi, at Florence. After this, various observers
saw the insect, but frequently it was confounded with other
mites, especially the cheese mite. Linneus in his Systema
Nature, 1734, had a very imperfect idea of it.’
In the century following the first appearance of the Systema
Nature, the mites, as causes of the itch, were frequently the
‘subject of heated discussions among medical men. The iden-
tity of the human itch mite was better known to naturalists
than to physicians.’ For instance, in a large treatise on skin
diseases by P. Rayer, originally published in Paris in 1835,
the author, in describing the causes of the itch, says: “ Causes
—Scabies is one of the most universally disseminated conta-
gious diseases known; the most momentary contact of the
fluid secreted by its vesicles is enough to communicate the
infection ; it is observed in every climate during every season
of the year, attacking all ages and persons in every rank of
life, without discrimination. It is most commonly observed,
however, among the poor and wretched,” etc. Further on he
1 Read before the Indiana Academy of Science, December 28, 1892.
2 Hebra, “ Diseases of the Skin,” 1868. P. 168-180
3 Hebra, “ Diseases of the Skin,” 1868. P. 180-181.
1893.] An Extreme Case of Parasitism. 347
says: “Several authors of modern times have reported that
they have discovered an apterous insect, almost invisible to the
naked eye, within the vesicles of scabies, which they have
described under the name of acarus scabiei,” ete. The author
does not state whether he believes this.’
The labors of many observers in 1834 and 1836 firmly estab-
lished the true nature of the itch or scabies. In 1845 and
1846 the minute anatomy of the insect was thoroughly studied
and described.’ The mite Sarcoptes scabiei, as the cause of the
itch, was established beyond all doubt.
In recent medical works, I find there is little uniformity in
regard to the systematic classification of this animal and its
generic and specific names. The latter are variously given,
the most popular seems to be Acarus scabiei. Frequently, all
the different names are given. A recent treatise has it thus:
“The acarus scabiei, sarcoptes scabiei, or sarcoptes hominis,
commonly called the itch mite.” Authorities for names are
never given. The accounts of the animal are often very brief
and unsatisfactory. The size of the mites is variously given.
According to some it is “ very minute, almost microscopic; ”
according to others, “ the size of a pin head.” The pictures or
figures of the mite differ greatly. In one work consulted—a
standard text-book by the way—an entirely different mite is
figured.’ From the great numbers of mites which I had for
comparison, I may say, without going into invidious distinc-
tions, that the best figures I have seen are those in Taylor's
“Atlas of Skin Diseases.” Dr. Piffard’s atlas contains an excel-
lent photo-micrograph of the female acarus.
The itch mite is now universally called by entomologists
Sarcoptes scabiei De Geer. The names Acarus scabiei and
Sarcoptes hominis, given in many medical works should be
dropped. Acarus and acari as synonyms for mite and mites
are, of course, allowable.
The best description of the mite I have seen is Hebra’s. It
‘runs: “ Body oval, tortoise-shaped, with indentations in the
t Reyer, “ Diseases of the Skin” (John Bell), 1845. P. 135-136.
` 5 Hebra, “ Diseases of the Skin.” P. 187-489.
6 Shoemaker, “ Diseases of the Skin,” 1892. P. 700.
1 Frey, “ Microscope and Microscopical Technology,” Cutter, 1880. P. 562.
348 The American Naturalist. [April,
lateral margins. Skin provided with shallow, undulating,
transverse furrows. Dorsal surface covered with numerous
small and large appendages of a clavate or conical form, or
resembling scales, and with ‘spines resting on papillary eleya-
tions. Head apparently distinct from the trunk, with four
pairs of jaws, and two strong, three-jointed palpi, placed near
the jaws, and of the same length. Legs eight in number, five
jointed ; the first and second pairs provided with pedunculated
suckers—the peduncles of the same length as the legs them-
selves, and having no joints; the third and fourth pairs in the
female terminating in long bristles. In the male, the first,
second, and fourth pairs of legs are all provided with suckers,
the third pair alone haying bristles; the epimera of the first
pair united. Larva presenting six legs—the first and second
pair with suckers, the third pair terminating in long bristles.”
In regard to size, Hebra says: “The adult acarus varies in
size. On an average, its length is 0.45 mm.; its breadth, at
the fourth ring of the thorax, 0.35 mm. The male is consid-
erably smaller than thefemale. In length it measures 0.23 mm..;
in breadth, 0.19 mm. The eggs of the itch mite are of an ovid
form, and are 0.16 mm. long, 0.11 mm. broad.” It is, perhaps,
unnecessary to say that the size of the animal is entirely out
of proportion to the commotion it occasions when its presence
becomes known.
The presence of the mite on the human body is character-
ized by certain appearances and symptoms known as ‘scabies
or ‘itch’. Scabies comes from scabere, to scratch. “The
name of ‘scabies’ is, at the present day, used to designate the
numerous appearances produced on the skin by the presence
of acari which dwell within it, and which, in obtaining food
for themselves, and in propagating their species, give rise to a
continual irritation of its component tissues. The morbid
appearances themselves resemble, in many respects, those
caused by other irritants, but, taken as a whole, they never-
theless present certain peculiarities, which justify our making
of them a Seance form of disease g ' Hebra.
“Though is the essential cause of scabies,
it must be remembered that EES almost, if not wholly.
? Hebra, “ Diseases of the Skin.” P. 192.
1893.] An Extreme Case of Parasitism. 349
identical, are produced by other species of the Sarcoptes
derived from horses, dogs, cats, camels, sheep and rabbits.”
Such cases are, however, rare.
It is, perhaps, unnecessary to add that the mite is never
found in the body, only upon it, never in the organs or in the
blood.
The human itch mite is perhaps coeval with man. It has
certainly always existed as far back as written history goes.
Hebra, for good reasons, believes that many of the afflictions
of the Israelites, as mentioned in the Old Testament, were
nothing other than the itch.” Aggravated forms no doubt
frequently occurred. Geologically speaking, the mites, as a
group, are of recent appearance (Garman)."
The itch is, at the present time, becoming one of the rarer
afflictions of mankind—like all filth diseases, it is gradually
disappearing.
(In his Atlas of Skin Diseases (p. 319), Doctor R. W. Taylor
says: “ Until a few years ago, scabies was a rather uncommon
disease in this country, but it has been noticed that in the
various large cities and in some small communities, there has
been of late, a remarkable increase in its prevalence.” Italics
mine.
In a recent letter to the writer, he expresses the belief that
this is mainly due to the influx of Russian and Polish immi-
grants.
Cases are seldom seen outside of the slums of large cities ;
but it sometimes happens that if a case occur, say in a small
town where none had appeared for years, perhaps half of the
population becomes affected before its true nature is known,
and appropriate remedies are applied. Vigorous medical treat-
ment soon blots out an epidemic.
In an ordinary, acute or epidemic case of itch, the number
of mites is quite small, probably not exceeding one hundred
adult animals.
® Taylor, ‘Atlas of Skin Diseases.” P. 327.
10 Hebra, “ Diseases of the Skin.” P. 164-166.
1 «Insect Life.” Vol. IV., P. 182.
350 The American Naturalist. (April,
Recently I met with an extreme and rare case of the itch,”
brought about by the presence in the skin of an immense
number of itch mites. It shows to what extent parasitism will
go under favorable conditions. The first detailed description
of this rare- form of itch was made by Hebra, of Vienna, in
1852, and who gave it the name of Scabies norvegica, or Nor-
way itch, because the case was from Norway. Since then,
other cases have been seen at various times and in different
places, still, it is so rare that modern treatises on skin diseases
do not describe it. Taylor’s large “Atlas of Skin Diseases,”
devotes just two lines to it, as follows: “ European writers,
particularly of previous decades, have described a severe form
of encrusted eczematous scabies, which they call Scabies nor-
vegica, or Norway itch. Itis not seen in this country.””
DESCRIPTION OF THE CASE.
The host of the mites was a middle-aged white man; he was
partly paralyzed, and hence comparatively helpless. At the
time of his admission to the City Hospital, his entire body
was covered with thick, yellowish-white, leathery scales, the
largest measuring over one inch in diameter, and over one-
tenth inch in thickness. He was literally covered with scales
like a fish:
On rubbing the body slightly, a large quantity of these
scales fell off; rubbing more briskly removed an additional
quantity, but with the production of small bleeding places
(these scales were not crusts or scabs, they were epithelial pro-
liferations). Ata few places on the body crusts were found,
mainly where the skin was bent, as at the joints. On account
of the bleeding, no vigorous mechanical treatment was used at.
_ anytime; the scales were gently rubbed off. A handful could
be gathered daily. Despite this great alteration of the skin, the
general health remained good. Taken in its entirety, the case
was a puzzling one.
In a search for the cause of this eruption, and as an aid to
the determination of the disease, some of the scales were sec-
“ The case was described from a medical standpoint in the Jndiana Medical Jour-
nal for November, 1892, by Dr. Brayton and myself,
13 Taylor, “Atlas of Skin Diseases.” P, 327.
*
.
1893.] An Extreme Case of Parasitism. 351
tioned. Itch mites were found in abundance, and the nature
of the disease was soon established.
In order to study the case more thoroughly, a number of the
scales were prepared by usual histological method. The scales
were fixed and hardened by immersion in alcohol, stained in
picrocarmine, finally imbedded in paraffin, sectioned, affixed to
slides with clove-oil collodion, and mounted in series. The
shrinkage in passing through this process was very slight. The
differential staining in this case was peculiar: the epithelium
stained red, the mites yellow; the eggs remained unstained,
that is, white; the fæces appeared dark brown or black. In
staining sections of normal skin with picrocarmine, the outer
or horny layer stains a bright yellow, the underlying proto-
plasmic layers stain more or less deeply red. In the present
case the epithelium is peculiar in that well-stained sections
show very little yellow; this indicates that the component
cells are still protoplasmic, and have not cornified.
The activity going on in the columnar layer to replace this
constant and enormous loss of cells must have been great.
Apparently a vicious circle was established; the presence of
the mites irritating the skin—the skin, to protect or rid itself
from the parasites, pushed out a large number of new cells.
These succulent cells, however, furnished a good nidus for the
mites. The mites increased, causing increased irritation of the
skin, and this caused an increased proliferation of cells. Thus
the case went on, until it assumed this extreme form.
I may here say, that under appropriate anti-parasitic treat-
ment at the hospital, the mites were soon exterminated. The
cause of the irritation once removed, the skin gradually
regained its normal character.
NUMBER or Mires.—In®*order to obtain an idea of the num-
ber of mites present on the host, that is, at the time the mites
were present in greatest numbers, several scales were cut to
measure one-half by one-eighth inch, sectioned and mounted
serially. It will be readily understood that in such a pro-
cedure the mites were cut in various planes and directions;
sometimes the head would be seen in one section, the body in
one or two other sections, and the tail in still another. Many
of the smaller embryos and eggs appeared entire.
352 The American Naturalist. [April,
Anyone who has attempted to get numerical results from
serial sections, will understand the difficulty of obtaining
approximately correct figures. In the present case, two counts
were made in each of three series of sections, and the mean of
these counts was taken as an approximately correct number.
Here are the figures:
Eggs and egg cases, 380.
Mites in all stages of development, 109.
Assuming that the above figures are approximately correct,
we can make a little calculation: Multiplying the above
numbers by 16, gives the number for one square inch ; again
multiplying by 144, gives the number for one square foot.
_ The average amount of epithelial covering on an adult
human being, is said to be about 16 square feet. To be on the
safe side, we will say that only half of the body was covered
with scales, therefore, multiplying our last figures by eight,
gives the number of eggs and egg-cases and mites on the host
at the time of greatest abundance. Here are the figures in
round numbers:
Egg cases and eggs, 7,000,000.
Mites, 2,000,000.
Since the reading of this paper, I tried to determine the proportion of eggs which
are empty, that is, hatched. It seems that from one-half to three-fourths are empty
shells.
As to the number of ving mites in the shed scales. This is a difficult question. I
am inclined to believe that only a small Proportion were alive at the time the scales
became detached.
Indianapolis Ind.
1893.] Distribution of the North American Unionide. 35
v3
ON THE RELATIONSHIPS AND DISTRIBUTION OF
THE NORTH AMERICAN UNIONIDAE, WITH
NOTES ON THE WEST COAST SPECIES.
By Cuas. T. Simpson.
For some years past, it has been generally acknowledged by
students of the Unionidæ that a more natural system of classi-
fication was needed for that family than any that has hitherto
been offered.
The genera or sub-genera of Rafinesque, Swainson, Agassiz
and Conrad are of little value, for while certain typical species
may be fairly referred to their divisions, in many others the
characters fade out, or so blend together that it is impossible
to place them, and sub-generic lines cannot be drawn with any
degree of accuracy. Lea’s arrangement, in which he divided
the genera into two great groups, characterized by the presence
or absence of a dorsal wing on the shell, and then into smaller
divisions based on sculpture and form, was, like the Linnæan
classification in botany, almost entirely artificial, consequently
in many cases it brought together side by side forms not at all
nearly related, and in others it widely separated those having
close affinities, or even sometimes individuals of the same
species.
I believe that the best and simplest system that can be,
adopted at the present, at least, is to place those species, which
by the characters of the shell, by what we know of those of the
animal, and by the facts of geographical distribution are most
nearly related, in groups, each founded on and taking its name
from some common, characteristic, and widely distributed
form. Thus Unio gibbosus, which is nearly everywhere found
throughout the Mississippi drainage area, is well known to
every collector who has only a few species of Unios, and, with
its elliptic oval and sometimes arcuate outline, thick epider-
mis, heavy cardinal and solid club-shaped lateral teeth, fairly
typifies an assemblage of forms found from Canada to Central
America, and from the Missouri river to Florida. Unio liga-
24
354 The American Naturalist. [April,
mentinus represents another section, Unio complanatus another,
| mph 2 margaritifera a fourth, Anodonta ovata a fifth, and
so on.’
This plan of grouping by relationships is not a new one,
having been adopted by R. Ellsworth Call, Wm. A. Marsh, and
other students of the Unionide in the arrangement of their
collections, and even Mr. Lea acknowledged some such affini-
ties, though he did not put his ideas into practice.
It is found that a common assemblage of Unionide inhabits
the entire Mississippi drainage basin, and that a considerable
number of the species have a distribution covering the- greater
part of this area, as well as the whole of Texas, and even parts
of eastern Mexico.
Those streams which fall into the Atlantic are peopled by
an entirely different set of forms, the Appalachian chain seem-
ing to act as a sharp barrier between the two regions. Many
of the Mississippi Valley species have spread into Michigan, a
few into Canada, into streams in New York that flow into the
Atlantic, and two or three of these are met with in the Red
River of the North, and the waters about Hudson Bay; while
below the southern end of the mountain range, there is another
mingling of groups.”
There are some forms which, apparently, may be with equal
propriety assigned to either of two or three groups, but, in
such cases, the location of the species with regard to drainage
often gives us a key as to its relationship. There is a group
of oval, rather smooth, compressed Unios, which may be fairly
typified by U.favosus, which is found in the rivers from North
Carolina to Alabama, several members of which very strongly
1 With such an arrangement, I believe that the North American Unios would fall
into something like forty quite natural assemblages, whose group characters agree
fairly well with their distribution so far as drain areas are concerned. There
would be some half dozen sections of Margaritana, and ten or a dozen of Anodonta.
*In many cases, the Unionidz seem to have had no difficulty in migrating across
the country from river to river; an example of this being the Mississippi Valley species
which now inhabit all the rivers of Texas, and some of those of Eastern Mexico;
„in some cases, may be carried by aquatic birds in the manner else-
where mentioned in this paper; in others, they probably migrate across overflowed
regions near the sea, in time of floods.
I
or aed
1893.] Distribution of the North American Unionide. 355
resemble some of those of the Clavus Group, but the species of
the latter assemblage are confined, with one or two exceptions,
to the streams of the Mississippi Basin, while those of the
former belong entirely in waters falling into the Atlantic or
the Gulf. Another small group of rounded or oval inflated
forms typified by U. irrasus likewise approaches in a few of its
members some of the species of the clavus Group, but it is con-
fined, so far as my knowledge goes, to the waters of the Gulf
drainage; the Etowah, Coosa, Chattahoochee, and a few neigh-
boring streams, and nota specimen has, I believe, ever come
from Tennessee or the rivers emptying into the Atlantic,
though several of them rise near the Chattahoochee.
In Mexico and Central America, a totally different Unio fauna
is found. Some of the Anodons extend into South America,
and others belong to groups that have their metropolis on that
Continent, while at the North there is a greater or less ming-
ling of the species with those of the Mississippi Valley.
I have been greatly puzzled to account for the origin and
relations of the few species found on the Pacific slope of North
America. ‘Two Unios, one Margaritana, and some half dozen
Anodons, are all that have been hitherto credited to this
immense region. Of the latter, Anodonta wahlametensis, A.
nuttalliana, A. oregonensis, and A. californiensis, are believed by
Dr. Stearns’ and other competent conchologists to be merely
variations of one species, although they often differ greatly in
forms and general appearance. A. wahlametensis is provided
witha wide dorsal wing, which gradually fades out through
the other forms in the order I have given them, to A. califor-
niensis, which is nearly or quite destitute of this appendage.
This very abundant species is found from the Rocky Mount-
ains on the east—in all the waters draining into the Pacific—
to the coast, and from British Columbia on the north, well
into Mexico on the south. Six shells in the collection of the
United States National Museum (Mus. No. 117951), which
were identified by Dr. Stearns as Anodonia californiensis, were
collected by Mr. Duges in the Province of Oajacay.
3 } Jistory and Distribution of the Fresh Water Mussels, and
rotate ae fikies by R. E. C. Stearns, Proc. Cal. Acad. Sci., Nov.
20th, 1882.
356. The American Naturalist. [April,
To the northward, this seems to be replaced by Anodonta
kennerlyi, a species closely resembling some of the varieties of
A. cygnea of Europe, and in Alaska another nearly related
form, A. youkonensis of Lea, is met with. The latter is an
absolute synonym of A. herculea Midd., of which specimens `
are in our collection, obtained in Kamschatka by Dr. Dall. 1
had noticed the resemblance of all these forms to that of the
well known, abundant, variable, and widely distributed Ano-
donta cygnea and its variety anatina, and on hunting up the
distribution of that species, I have been able to trace it through
Northern Asia into Eastern Siberia. Any one who will care-
fully examine an extensive series of Anodonta cygnea, anatina,
herculea, youkonensis, kennerlyi, wahlametensis, nuttalliana, oregon-
ensis, and californiensis, cannot, I think, fail to notice their
remarkable resemblance in many essential points: the pecu-
liar and usually exact rounding of the anterior end of the
shell, the flattening of the umbonal region, which, in good
specimens, is generally overlaid with hair-like, greenish, radi-
ating, wavy lines, and the singular cutaway in the posterior
slope, all indicate close relationship, and the fact that they
have sprung from a common origin.
I should unhesitatingly place these forms in a natural
group, typified by Anodonta cygnea. Dr. Stearns, in the paper
to which I have already alluded, believes that Anodonta wahl-
ametensis, nuttalliana, oregonensis, and californiensis, are but —
variations of the European A. cygnea, and that A. fluviatilis of
the Atlantic drainage is probably but another form of this
protean species.
While I was formerly inclined to this view, a more careful
and thorough examination of large quantities of material has
_ induced me to somewhat modify my opinions. Though the
nuttalliana and its variations are very close to cygnea, I do
not, at present, feel justified in uniting them specifically, and
I regard the A. fluviatilis as the type of a rather nearly related
group, having a distribution from Canada to Florida, in the
streams which fall into the Atlantic. The shells of this section
are generally more inflated than those which belong with
eygnea ; they are not flattened at the umbonal region, and atè
more full on the dorsal slope.
1893.] Distribution of the North American Unionidæ. 357
IfI am right in this view of the case, it is probable that the
group of Anodonta cygnea is confined in North America to the
Pacific slope, and that its connection with the rest of the
species in the Old World was made over a land passage
between Asia and North America. Dr. Theodore Gill informs
me that the boreal fresh-water fishes of the Old World and
those of the western slope of our continent have a similar dis-
tribution to the cygnea group of Anodons as I have outlined it.
One of the most remarkable of the Unionidæ, Anodonta
angulata, is an inhabitant of the waters of Oregon, Washing-
ton, and California. The anterior end of this species in ordi-
nary specimens is much narrowed ; the posterior is wide and
inflated, and, running from the beaks to the posterior ventral
regions is an extraordinarily developed sharp-edged ridge or
keel. Nothing in the least resembling this peculiar form has
hitherto been found. But the National Museum possesses
specimens in which the anterior end is wider, and the keel is
far less developed, and others in which it is almost entirely
wanting, and that indefatigable collector, Mr. Henry Hemp-
hill, has sent me a photograph of a specimen in his collection
which has no vestige of a ridge, but is furnished with a slight
groove running down the posterior slope. Recently, Messrs
Mearns and Holzner, of the International Boundary Commis-
sion, have sent to the National Museum a large number of
specimens of what is apparently a new species of Anodonta,
from San Bernardino Ranch, Arizona, which, externally, very
much resembles some of the varieties of Unio complanatus,
but has a very different texture, and, on comparing these
shells with Mr. Hemphill’s photograph, and the more com-
pressed specimens of A. angulata, I was convinced at once that
they group together. They have the same outline, the same
beak sculpture, and, when viewed from the dorsal or ventral
region, have a much greater diameter through the posterior
area than elsewhere, and nearly all exhibit the slight groove
I have mentioned.
The type of Unio oregonensis was presented to Mr. Lea by
Mr. C. M. Wheatley, and the former mentions that the latter
reserved a specimen for his own collection. Lea’s shell is said
to come from the Columbia River, and is in rather bad condi-
358 The American Naturalist. [April,
tion, as it is dead and somewhat eroded. I have very carefully
examined it, and after comparing it with everything that I
could think might be related to it, I have come to the conclu-
sion that it is a form of the protean Unio luteolus.
This is the most abundant, widely distributed, and variable
Unio in the world, being found almost everywhere throughout
the 1,200,000 square miles of the Mississippi drainage basin,
through Texas, and probably into Mexico, in New York in
some of the streams that flow into the Atlantic, and northward
in the vicinity of Hudson Bay, and probably throughout the
British possessions east of the Rocky Mountains. I have
traced it up the Missouri River to near its source, and when it
is taken into consideration that the Marias, a tributary of this
stream, heads within a few miles of Flathead Lake on Clarkes
River, a branch of the Columbia, and that the embryos of
Unios are provided with hooks by which they can attach
themselves to the feet or feathers of aquatic birds, it is very
easy to see how this species might have been carried from the
waters of one drainage system to those of another. Margari-
tana margaritifera, which inhabits Europe, Northern Asia, and
the Pacific slope of North America, is also found in the upper
Missouri. Two specimens of Unio luteolus in the Museum col-
lection, from the North Shore of Lake Superior, are almost
exactly like Lea’s type of U. oregonensis.
So far as I am aware, nothing is stated of Unio famelicus,
the other Pacific slope species, save that Gould received his
type from Dr. Pickering, and that it is said to have come from
Walla Walla, Oregon. The type is in the National Museum,
and is undoubtedly a young Unio ellipticus Spix, from Brazil, a
species belonging to an extensive South American group,
characterized by having oblong, compressed, sulcate shells,
which are often granulated on the ridges. Any one familiar
with Unios would know at once on carefully examining the
type, that it came either from South America, or some part of
the Southern Hemisphere of the Old World, a fact which is
made evident by its form, its peculiar greenish olive epider-
mis, the strong growth lines, and, most of all, the compressed,
parallel, cardinal teeth of the right valve. It is perfectly safe
to say that the locality given is an error, and that this species
was never found in the United States.
1893.] Editorials. 359
EDITORIALS.
EDITORS, E. D. COPE, AND J. S. KINGSLEY.
—Sratistics as to the publication of original books by the four
leading nations of readers have recently appeared. America and
Great Britain have published during the last decade between four
and five thousand books each. In America the proportion of fic-
tion was considerably larger than in Great Britain. In Germany
the number of books published in the same time was over ten
thousand, with a smaller absolute number of works of fiction than in
Great Britain. The figures for France fall between those of Great
Britain and Germany.
In view of the fact that America exceeds in population each of the
countries named, and includes in this population a greater number of
readers, these statistics are not pleasing to our national pride. Great
Britain, with a much smaller population, is our superior in the quality
of books produced, and yet falls far behind Germany.
There are several reasons for this state of affairs. One is theabsence
of compulsory education, which exists in an effective condition in
Germany. Another, and probably more efficient cause, is the fact
that our publishers think it necessary to bring out books of value in
an expensive style, so that they are out of the reach of students gen-
erally. The same cause probably conduces to the depression in the
publishing business in Great Britain. If American students could
purchase scientific and other serious books at as low rates as can be
done in Germany, the general standard of intelligence would be greatly
elevated. If American publishers would get out cheap editions of
such literature they would greatly profit by it. The extravagant style
in which popular scientific books are published in this country is an
anachronism. There is no nobility here whose attention it is neces-
sary to gain for material reasons, nor is social position the object for
which the scientist is supposed to be striving. Hence the absurdity of
clothing science in a garb which can attract a small class only, while
it makes her inaccessible to the mass of mankind. The supposition
that people will not buy solid literature is negatived by the experience |
of Germany. If American publishers will not publish cheap works
of this class we shall become dependent on foreign countries, whose
books will soon become accessible to us by the removal of the oppress-
ive tax on intelligence, known as the protective tariff.
360 The American Naturalist. [April,
— W2 have received the first number of the New Journal of Geology
published by the geological faculty of the University of Chicago, and
printed by the press of the University. It is edited by six members
of the United States Geological Survey, together with twelve associate
editors, of whom five are Americans, and one resides in each of the
following countries, viz.: Canada, Great Britain, Germany, France,
Austria, Norway, and Sweden. The articles of the number are by
Messrs Geikie, Holmes, Williams, Chamberlin and Salisbury, and are
all of much interest. Editorials, Reviews, Abstracts, and Acknow-
ledgments follow. The beginning of this new journal is auspicious,
but it remains with the future to determine whether it can be contin-
ued on its present basis. On the face of it the proposition is broad,
but the basis is narrow. The names of the associate editors indicate
cosmopolitan aspirations, but “the immediate editorship,” we are told
in the announcement, “will rest with the geological faculty of the
University of Chicago,” who are all members or ex-members of the
U.S. Geological Survey. The leading article is by a foreigner, but
this is appropriate in a city which has legal holidays in honor of
William of Germany and Saint Patrick. We will observe with
interest whether such a constitution can stand the test of time, or
whether it will meet the difficulties which ‘befel the image of the
prophetic vision, in which the geological order was reversed, since the
head was of archean gold, but whose feet were of glacial clay. Not
but that the precious metal may be found in the superficial stratum in
this instance; but placer deposits in glacial drift are not as permanent
as those of greater geologic age. We have an excellent geological
magazine, the American Geologist, which is not identified with any
institution or institutions, and whose editorial corps is not interested in
supporting geological literature in other countries. While, therefore,
we welcome all additions to scientific periodical literature, it is always
with a reservation which requires that they shall be viable infants,
endowed with a good constitution. i
—A movement is on foot in New Zealand to secure an island not far
from the coast as a preserve for the native animals and plants of that
country. In view of the extreme importance to biological science
of the preservation of the remarkable types of this ancient flora and
fauna, and in view of the other fact that many of its representatives
have become extinct within recent times, and others are disappearing,
we sincerely hope that this effort will be successful. If there is any
doubt about it, a concerted effort should be made by the scientific men
of all countries to secure favorable action from the colonial legislature.
a eras ahs Sateen eee
1893.] Recent Books and Pamphlets. 361
RECENT BOOKS AND PAMPHLETS.
ALLEN, H.—On a New Subfamily of Phyllostome Bats.—— Description of a New
Genus of Phyllostome Bats.——On Temminck’s Bat, Scotophilus temminckii. Extr.
Proceeds. U. S. Natl. Mus., Vol. xv. From the author
Biennial Report of the Director of the Minois State Laboratory of Natural His-
tory, 1891-1892.
OLLMAN, C. H. E ay G Fontaria pulchella T Strawberry Plains,
Tenn. Extr. Proceeds. U. S. Natl . 1888. From the author.
BONAPARTE, R.—La Novège et k geni résumé de a dua Extr. Le
Globe Journ. Geog., T. 28,1889. From the a
BRIGHAM, A. P.—A Chapter in Glacial History with ‘Illustrative Notes from Cen-
tral New York. Extr. Trans. Oneida Hist. Soc., 1892.
—— The Geology of Oneida County, seh York, Extr. Trans. Oneida Hist. Soc.,
1887-88.
Rivers and the Evolution of Geographic Forms, Extr. Bull. Am. Geog. Soc.,
1892. From the
Bulletin of the THlinois State Laboratory of Natural History, Vol. iii, 1892.
BURMEISTER, G.—Anales del Museo Nacional de Buenos Aires para dar á conocer
los objectos de Historia Natural Muevos 6 Poco Conocidos conservados en este
establecimiento. Entrega Decimaoctava, 1891. From the au
Crook, A. R.—Ueber einige fossile Knochenfische aus der aiai Kreide von
Kansas. Separat-Abdruck aus Palaeontographica, xxxix, Band, Stuttgart, 1892.
Cross, WHITMAN.—The Post-Laramie Beds of Middle Park, Colo. Extr. Pro-
ceeds. Col. Sci. Soc., 1892. From the author.
Dorsey, J. O. —The Cegiha Language. Contributions to san American Eth-
nology, Vol. v i, Washington , 1890. From the Dept. of the Interi
EARLE, C.—A Memoir deren the Genus Palzosyops Leidy and ie Allies. Reprint
Journ. H Acad., Vol. ix, Oct., 1892.
—— Revision of the Species of Coryphodon. Extr. Bull. Am. Mus. Nat. Hist.,
Oct., 1892. Bee Ar
The Variability of Specific Characters as Exhibited by the Extinct Genus
ee: Extr. Science, July 1, 1892. From the ai >
Fos. P. M.—The Fertility of Hybrids. Extr. Medical News, 1892. From
the Sakae
GARMAN, H.—A Synopsis of the Reptiles and Amphibians of Illinois. Bull. Ill,
oy — Vol. iii, 1892. From the author.
GARNIER, Dr.—The Reptiles of Canada, Extr. The Land we Live In. From
the author.
GEINITz, H. B.—Bericht über die neue Aufstellung in dem kénigl. Mineralo-
gischen Museum zu Dresden. Sitzung der Gesellschaft Isis in Dresden, April, 1892.
From the author.
GILL, TH.—On the jan Generic Name of the Tunny and Albicore. On the
Psychrolutidæ of Guenther. Extr. U. S. Natl. Mus., 1888. From the aut
Haves, C. W.—Report on the Geology of Northeastern Alabama and pE
Portions óf Georgia and Tennessee, 1892. From the author.
362 The American Naturalist. [Apri},
Horm, TH.—The Flora of the Dakota Group. Extr. Botanical Gazette, Vol.
xvii. From the author.
Howes, G. B.—On the Affinities, Interrelationships, and Systematic Position of
the Marsipobranchii. Extr. Trans. Biol. Soc. Liv erpool, Vol. vi.
——On the Pedal Extremity of the Dorking Fowl, with Remarks on Hexadac tyl-
ism and Phalangeal Variation in the Amniota. Notes Upon the aoi Pris
of Certain Dicynodontid Reptiles. Extrs. Jour. Anat. and Physiol., Vol. x
——On the Arrangement of the Living Fishes, as based upon the study p ‘their
Reproductive System.—On the Customary Methods of Describing the Gills of
Fishes, Read at the Cardiff Meeting of the British Assoc., 1891.
OWES, G. B., AND J. HARRISON.—On the Skeleton ae Teeth of the Australian
ass Read at the Edinburgh Meeting of the British Assoc., 1892. From the
author
JORDAN, D. S.—List of Fishes collected by Alphonse Forrer about Mazatlan,
with konka of two new species, Heros beanii and Peæcilia butlerii. Extr. U.S.
Natl. Mus., 1888. From the author.
Keyes, C. R.—Iowa’s Natural Resources, Extr. Iowa State Register, July 12,
1892. From the author
LYDEKKER KER, R.—On a a Collection of Mammalian Bones from Mongolia. Extr..
Records Geol. Surv. India, Vol. xxiv, 1891. From the author
Macoun, J.—Catalogue of Canadian Plants. Part VI, Wiad. From the author.
East Runton, near Cromer. Extr. Trans. Zool. Soc. London, Vol. xiii, Part IV,
—— The Vertebrata of the Pliocene Deposits of Britain. Extr. Mem. Geol.
Surv. of the koer) Kingdom, 1891. From the author
NIBLACK, A. Bi Ethaolesy of the Coast Indian Tribes of Alaska. Extr. Pro-
ceeds. U. S. Natl. Mus., 1888. From the author.
Pocta, PH.—Ueber Spongien aus der oberen Kreide py in dem k.
Mineralogischen Museum in Dresden, mit Vorwort von Dr..H. B. einitz. From
pie T, M. G.—Nouvelles observations sur la Sardine océanique. Extr. Comptes
ais pia, 1889.
Quarterly Bulletin of the University of Min :
Report of the Commissioner of Education, 1806-86, Vol. i. From the Dept. of
the Interior.
Rouon, J. V.—Die Obersilurischen Fische von Oesel. Mém de l'Acad. Sci. de
St. Petersbourg, vii E Serie, T. xxxviii, No. 18, 1892.
——Ueber Pterichthys. Separat-Abdruck a. d. Verh. d. Russ. Kais. Miner.
Ges. zu St. Petb., Bd. xxviii, 1891. From the author.
RN, L.—Nos Fleurs. From Armand, Collin & Cie, Editeurs, piene 1892
SCHELLWEIN, E.—Die Fauna des Kinis Fusulinenkalks, A s Palzonto- .
graphica, Neununddreissigster Band. Erste Lie eferung, Stuttgart satay
ie 2 logical and Min neralogical Notes, No 5. pe Bull. Essex
Inst., 1892. From the author.
SımPsoN, C. T.—Notes on the Unionidæ of Florida and the Southeastern States.
Extr. Proceeds. U. S. Natl. Mus., Vol. xv. From the Smithsonian Institution.
1893,] Recent Books and Pamphlets. 363
Report U. S. National Museum, Part II, 1886; Part II,1887. From
the M
Top J. E —Notes on the Geology of Northwestern Iowa. Extr. Proceeds. Iowa
Acad. of Sciences, 1891.
——Striation of Rocks by River Ice. Extr. Am. Geol., Vol. ix, 1892. From the
author.
TRUE, F. W.—An Annotated Catalogue of the Mammals collected by Dr. W. L.
Abbott in Ps Kilimanjaro Region, East Africa. Extr. Proceeds. U. S. Natl. Mus.,
Vol. xv. he Smithsonian Institution. .
United Sar Commission of Fish and Fisheries, Part xvi.
missioner for 1
EAVES
Basin.
Report of the Com-
ES, F —The Orthoceratidz of the Trenton Limestone of the Winni-
Extr. Trans. Roy. Soc. Canada, Vol. ix, 1891. From the author
364 : The American Naturalist. [April,
RECENT LITERATURE.
The Woodland of the Southwest, by Chas. F. Lummis.!
—For those readers who have read but a few books of travel on the
Southwest, this snug little volume will be quite a revelation. The
contents of the twenty-two chapters scarcely contain anything that has
been written or sketched before, except a few pages on the Moqui
Snake dance and Indian superstitions. The thoroughness of his famil-
iarity with Pueblo customs and folklore is only equalled by the graphic
qualities of his style. In looking about the “strange corners” which
the author describes, we are first attracted by a prairie-dog hunt, with
the Navajo Indians put in scene to fill their larder. White people of
the Southwest never think of killing this rodent for food, because it is
so difficult to attain with a rifle ball; but these natives utilize abund-
ant downpours of rain to conduct the floods into their tunnels and
afterward haul up their dead bodies for a feast. To get rid of the
prairie-dog plague, people have proposed to kill them with poisoned
apple-quarters. The belief in witchcraft is as potent among the whites
and Indians of New Mexico as it ever was during the middle ages.
Manslaughter is committed for any act arousing even the suspicion of
witchery, and the fact that one-half of the Isleta people are wizards.
and witches speaks loud enough. The “Finishing an Indian Boy”
shows principles of education in full force now, which our Northern
Indians began to drop as early as one century ago. In the chapter
“The American Sahara” the wide waste is delineated in colors none
too sharp or cruel. Lieut. Wheeler is mentioned by mistake as its
earliest explorer instead of Lieut. Whipple. The marvelous wealth of
objects presented in Lummis’s volume will attract ever and again the
class of readers and tourists which seeks instruction rather than pleas-
ure in books of travel, and they will hold it dear as a publication of
really scientific value, standing far above most of the productions of
our present sensation-loving period of literature.
“The Wanderings of Cochiti” is another very interesting sketch
from our “ Wonderland” on the upper Rio Grande. It is printed in
the “Century Magazine,” January, 1893, and describes and also pic-
tures in photographic reproductions the people, customs, history and
scenery of Cochiti, one of the Quéres Pueblos of Northern New Mex-
e strange corners of our country, the Woodland of the Southwest, p- 270. p
mo., richly illustrated. New York, The Century Co., 1892.
GS NCE irs So ae oa ie Sac AN
NERO so Ro Sat Seen ee Lear a. Sc ah ee eae a tah to? Oa 7 Bays CP ae ee eam oR Soh a
1893.] Recent Literature. 365
ico, and the celebrated gorge of Tyii-on-yi with its rock carvings in the
vicinity of that pueblo. The scene of Bandelier’s arch:eologic novel,
“ The Delight-Makers,” is placed in that locality.
Macmillan’s Metasperma of the Minnesota Valley.’—
In this thick, handsomely printed book, in which excellent paper and
an equally excellent selection of type are manifest, we have a notable
departure from the usual “ report” issued from a State printing office.
So unlike the State or Government reports is this volume that one has
to turn to the imprint to satisfy himself that it is actually the work of
the State printer. If this model is followed in the future by other
State printers then indeed is this book a noble pioneer in a much
needed reform in State book making.
Turning to the more important part of the book, namely its con-
tents, we find that the author also has broken away from the tradi-
tional form and style of State reports, and has given to the botanical
world a useful book. He first of all chose an area with natural
boundaries, instead of one arbitrarily set off for political and not sci-
entific purposes. In his choice of the Minnesota Valley the author
seems to have been fortunate in having to deal with an area of more
than usual interest, but it is likely that if in other States natural areas
were studied, similar interesting features would be discovered.
The list of plants includes 1174 species and varieties, a large num-
ber when one remembers that the area covers but 16,000 square miles,
and the only plants enumerated are the Metaspermz (Angiospermz).
These species are found in prairies and woodlands, in “ meadows,
marshes, swamps and bogs.” ‘There is, however, a considerable pre-
ponderance of the rolling prairies. The soils are not much varied,
and the difference in altitude is little more than 1300 feet from the
lowest point at Fort Snelling (688 ft.), to the highest hills in the east-
ern part of North Dakota (2000 ft.).
The author aims to trace the history of the flora of this valley. To
him it is not enough to tell what plants occupy the territory ; he seeks
to show how they came to do so, from whence they came, and why
they came. Accordingly we have a series of discussions such as we
rarely (if ever) have had in books dealing with local floras. Thus
one finds such headings as these: “ The Dynamic Relations of Plants,”
2Geological and Natural History Survey of Minnesota. Conway Macmillan, State
Botanist. Zhe Metasperme of the Minnesota Valley, a list of the higher seed-pro-
ducing plants indigenous to the Drainage Basin of the Minnesota River, by Conway
Macmillan. Reports of the Survey, Botanical Series, i, Dec. 29, 1892, Minneapo-
lis, Minnesota, pp. xiii, 826. Large octavo.
366 The American Naturalist. [April,
“General Features of Plant Distribution,” “The General Factors in
Floral Differences,” “ Pressures and Tensions,” “ Outlines of Meta-
spermic History in the Northern Hemisphere,” etc.
Enough has been said to indicate that this work is an unusual one.
It is impossible to give a summary of its many interesting chapters.
Perhaps the conclusion which the author reaches that while the region
is geographically central in the continent, the flora is “ preponderantly
southeastern,” sums up the results of the work as well as any which
could be given.
It remains to speak somewhat more as to the list itself. In the first
place it is ultra-modern in nomenclature and arrangement. Engler &
Prantl’s work, “ Die Natiirlichen Pflanzenfamilien,” is followed in the
sequence of the families, so that one finds the simple Monocotyledons
first (not the grasses, and sedges, however), and following the well-
known succession of families, reaching the culmination of the Dicoty-
ledons in the Composite. The manuscript of the book went to the
printer four months before the Rochester meeting of botanists last
year, hence it does not conform in all respects to the present rules as
to nomenclature. Thus the initial date adopted for the genera is 1737,
instead of 1753, and now and then one finds a specific name identical
with that of the genus to which it belongs. In the preface, which
bears date of Dec. 24, 1892, the author records his hearty acquies-
cence with the action of the botanists at Rochester, and suggests the
changes which must be made in order to bring the nomenclature in
the book into harmony with the present code.
A very valuable feature of the work is the fulness of citations of
synonyms, with dates. Thus one finds:
Lacrnraria Hill, Syst. Veg. iv, 49 (1762).
Psilosanthus Neck. Elem. (1790).
Liatris Schreb. Gen. Pl. 572(1791).
Calostelma Don, Sweet. Brit. Fl. Gard., 2, 184 Seegi
The following may serve to show the triatinent of s
LACINIARIA sQUARROSA (Linn.) Hill. Syst. Veg. iv, 49 (1762).
Serratula squarrosa Linn. Spee. 88 (1753).
Pteronia caroliniana Walt. F1. Gar, 292 (1788).
Liatris squarrosa Willd. Spec. IIT, 1065 (1802).
Then follow many references to descriptions or lists giving locali-
ties, followed by statements as to its general and particular distribu-
tion, ete.
The book will be read with interest by all advanced hoticnints: and —
it is to be hoped that it will serve as a model for future plant lists.
CHARLES E. Bessey.
1893.] Recent Literature. 367
Ridgway on the Anatomy of the Humming-Birds and
Swifts.—A Rejoinder.—What I say here has reference to Mr.
Ridgway’s reply to my review of his “ popular monograph of the
Humming-Birds,” which review appeared in the October NATURALIST,
and his reply in the December following issue of that journal (1892,
p. 1040).
In that reply Mr. Ridgway remarks that his description of the
humming-bird’s tongue “is substantially a condensation of MacGil-
livray’s”* (in Audubon’s Birds of America, Vol. iv, pp. 197, 198),
and that his “knowledge of the subject is based chiefly upon it.”
Now one of the errors I pointed out in Mr. Ridgway’s “ Humming-
Birds” was that he claimed the tongue in those birds was “ hollow,”
and yet he now states that his account is a condensation of Macgil-
livray’s lucid description. Let me contrast the statement of the two
authors, thus:
MACGILLIVRAY.
In the tongue of the humming-
bird.“‘there are, it is true, two cy-
lindrical tubes, but they gradually
taper away toward the points, and
instead of being pervious, form two
sheaths for the two terminal parts
or shafts of the glosso-hyal portion
of the tongue, which run nearly to
the tip.” (Aud. Birds of Amer.,
Vol. iv, p. 198).
RIDGWAY.
the humming-birds “The
‘hai is slender and very exten-
se like that of the woodpeckers.
* * * * Instead, however, of
its being as in the woodpeckers,
solid and tipped with a barbed,
horny point, it is hollow.” (The
Humming-Birds, p. 290).
These sentences, in either case, are completed by their respective
authors in their calling attention to the bifurcated condition of the
extremity of the tongue, and, as that is not the point in question, I
purposely omitted it.
arther on in his reply Mr. Ridgway remarks that his knowledge
of the structure of the humming-birds is to some extent based upon
the “later dissections of thirteen species,” made by Mr. F. A. Lucas,
“instead of one, as in the case of Dr. Shufeldt’s ‘extensive dissec-
tions.” If my friend will again allow me to invite his attention to
the literature of the subject, I would call it to my memoir published in
If Mr. Ridgway will permit one who has long been familiar with the “ee of
“the Scotch anatomist,” I would “kindly invite” his attention to the fact that
that careful dissector of birds spelled his name Macgillivray, and of jaiii
as Mr. Ridgway always writes it.—R. W. S.
368 The American Naturalist. [April,
the Linnean Societys Journal, Vol. xx (London), pp. 303, 304, where
he will find that I had the following material for my “extensive dis-
sections,” namely, spirit specimens of Celigena clemencie, Trochilus
colubris, T. alexandri, T. anna, T. platycercus, T. rufus, T. calliope and
Jache latirostris—in all 3 genera, 8 species, and 62 specimens. When
this material was presented by me to the collections of the U.S.
National Museum, Mr. Lucas remarked that it was the largest and
best collection of spirit specimens of North American humming-birds
ever donated to the institution by one individual. It naturally seems
to me that the further Mr. Ridgway goes into this subject the greater
is the display of his ignorance of the literature of it. Now when I
charge a writer with being ignorant of the literature of a subject I
take the pains, as I have done above, to point out wherein his ignor-
ance lies, and do not resort to the remarkable method adopted by Mr.
Ridgway—thus, and alluding in bis reply to myself, he continues:
“ Apparently he is not familiar with the literature of the subject, for,
if he were posted, he would know that leading authorities on avian
comparative anatomy are overwhelmingly if not unanimously against
his side of the question.” Rather remarkable logic—especially to be
immediately followed up in the very next sentence by, “ I would there-
fore suggest that he consult Fürbringer, Parker, Garrod and Gadow,
and thus learn regarding the matter which he handles with so much
urance. Even a careful perusal of Huxley (whom, by some strange
hallucination, he imagines his abettor), may also prove instructive to
him (p. 1040).” To this I would simply reply that in so far as Pro-
fessor Fürbringer is concerned, I can say that, thanks to his generosity,
I am the fortunate possessor, not only of many of his works upon
comparative avian anatomy, but also his two massive and superb vol-
umes so familiar to us all, and while I can thank him for his numer-
ous and courteous references to my memoirs in that great work, I
must dissent from his views in the matter of the taxonomy, of the
Cypseli and Trochili. Apparently Professor Fiirbringer had not seen
my Linnean memoir, which was defended for me by Professor Par-
ker before the London Linnean Society a number of years ago. There
is a reason for that, however, as both works appeared early in 1888.‘
*‘Shufeldt, R. W., Studies of the Macrochirem morphological and otherwise, vih jsi
view of indicating their relationships and d
Linn. Soc. Jour.—Zool., Vol. xx, pp. 299-394, Pls. Meira (Communicated by wW.
K. Parker, F. R. S., F. L. S., 19th Jan., 1888). When I spoke of my extensive dis-
sections this was the work I had reference to, fern not to the brief note in Forest and
Stream, as Mr. Ridgway seems to think. It stands to-day as the most extensive
paper on the entire structure of the Humming-birds ever published by a single indi-
vidual.
1893.] Recent Literature. 369
As for Professor Parker, I can say that I had the honor of being
one of his constant correspondents for a number of years, and I
have from him several hundred letters. A year or so before his
lamented death it was his intention to supplement my work upon ‘the
“Macrochires,” and I had sent him some material to that end. I
have letters to show from him that he practically agreed with me in
the taxonomy of the swifts, swallows and humming-birds, and he pro-
posed to figure one of the latter “as big as a Cochin-China cock!” It
is to the loss of the science of systematic ornithology that he never
lived to accomplish it. Next as to Garrod. My library contains a
copy of his “ Collected Scientific Papers,” a work I have had occasion
to refer to almost daily since its appearance. He was disposed to
classify birds upon altogether too few characters; there is no evidence
whatever in his work that he ever critically compared the entire strue-
ture of a humming-bird with that of a swift; he notes simply the fact
that “ the tensor patagii brevis (in the Cypselide and Trochilide) and
the pterylosis are characteristic, as is the sternum,” but he failed to
point out how widely they differed morphologically (p. 222). I dis-
sent altogether from his views not only upon his taxonomy of the
Cypseli and Trochili, but also from much else that he has left us upon
the classification of birds. I must ask Mr. Ridgway for the title of
any work from the pen of Professor Gadow wherein I may find a crit-
ical comparison of the entire structure of a swift and a humming-bird.
I know of none, though I do know, as do many others interested in
the classification of birds, that at the present time Dr. Gadow is hard
at work upon probably what will prove one of the best practical
schemes for the taxonomy of the class, and it will shortly appear.
He has already done me the great honor in inviting me to submit my
own views to him on the subject, and in a valuable letter I have just
received from that able taxonomer I find that he is prepared to admit
that the goatsuckers no longer should be retained in the same group
with the forms they have usually been associated with heretofore, and
he says, “I shall reconsider the position of the Caprimulgide and ele-
vate them perhaps on account of Steatornis as Caprimulgi, thus
making the difference from Cypselide and Trochilidæ more marked,
although I shall not go so far as Fiirbringer has done.” Mr. William
Brewster, of Cambridge, the distinguished American ornithologist, and
one of the Committee responsible for the “ Check-List ” of the Amer-
ican Ornithologist’s Union, and the classification therein set forth,
having read my contribution to The Ibis (January, 1893) on Swifts
and Humming-birds, writes me under date of February 13, 1893: “It
370 The American Naturalist. [April
is comforting to find that there are now, among scientific men of dis-
tinction, champions of the belief of my boyhood days that a swift is
merely a peculiar kind of swallow.”
Now as to my “strange hallucination” in the matter of Professor
Huxley’s views upon the classification of the Swifts, Swallows, and
Humming-birds. I have a well-worn copy of his famous P. Z. 8.
memoir of 1867—one of the most remarkable contributions ever
written upon the classification of birds—and, I may add, a perfect
monumental beacon, warning comparative avian anatomists for all time
in their attempts to classify birds, against trusting to any single set of
characters. In his “Cypselomorphe” Huxley placed only the Swifts,
Humming-birds and Goatsuckers (pp. 468, 469). It was done upon
only too few characters, and mainly based upon the osteological ones
seen at the base of the skull. But Huxley believed the vomer of a
Humming-bird was “ truncated at the anterior end,” an error which
both Parker and myself independently pointed out for him. But a
quarter of a century is a long time in comparative anatomy (1867-
1892), and if Professor Huxley has kept up with the literature of the
subject he may hold entirely different views at the present writing.
This would appear to be the more probable, as his misgivings were
sufficiently strong as to have him write in his 1867 memoir that “In
their cranial characters the Swifts are far more closely allied with the
Swallows than with any of the Desmognathous birds, the Swift pre-
senting but a very slight modification of the true Passerine type exhib-
ited by the Swallow (Joe. cit. p. 456). May I ask Mr. Ridgway what
he thinks Professor Huxley meant when he wrote that sentence?
And does he believe that to-day Professor Huxley would retain the
Caprimulgi with the Humming-birds and Swifts together in one group
and consider it to be a natural one?
One word more and I have done for the present. In the case of
doubtful affinity among birds, as with all other forms, there is but one
true way of getting at a solution, and that is to critically weigh and
compare everything that is known about the one group with evergthing
that is known about the other, before finally deciding. Now I chal-
lenge Mr. Ridgway to take any species of North American Humming- _
bird and any species of North American Swift, and arrange in two
columns in a comparative way all that is known about either species,
paleontologically, biologically, morphologically, or otherwise, and then
upon summing up give any real reason why the Trochili and the Cypseli
should be considered to constitute by themselves a natural group of
birds. I have very serious doubts as to his ability to do this. If he
1893.] Recent Literature. 371
ever undertake such a task, he must bear one thing well in mind,
and that he must not do not do it with the view of “exploding” any
one’s “ peculiar notions,” but rather with the view of getting at the truth
of the matter. Then, too, do not condemn another’s views simply
because “ leading authorities on avian anatomy are overwhelmingly if
not unanimously against his side of the question (p. 1040).” That has
been the case with a great many anatomists in the world’s history, and
even I can remember when the “ leading authorities” overwhelmingly
if not unanimously believed that Chamea was a Wren; but the present
writer in his published account’ based upon his “ extensive dissections ”
claimed it was most nearly related to the Bush-Tits (of the genus
Psaltriparus), to which opinion the “leading authorities,” who have
reviewed my work, are now rapidly coming. But no doubt Mr,
Ridgway remembers something of this; he most assuredly does if he is
familiar with the literature we have upon the subject.
R. W. SHUFELDT.
Animal Coloration.'—This volume has grown out of materials
collected by Mr. Beddard for the Davis lectures delivered at the Zoo-
logical Garden of London in 1890. It contains the latest information
as to the phenomena of coloration exhibited by animals, together with
the most important theories upon the subject, including those of Dr.
Eisig and M. Stolzmann, which had not previously found their way
into works of a popular character. Mr. Beddard groups his facts under
the following heads: Coloration Affected by Environment ; Protective
Coloration ; Warning Coloration ; Protective Mimicry ; Sexual Color-
ation.
In summarizing the facts concerning Protective Mimicry, the author
concludes that the theory of Natural Selection alone does not satisfac-
torily explain the remarkable resemblances included under that
caption. seit
The chapter on Sexual Coloration calls attention to some objections
to Darwin’s theory of sexual selection, and gives briefly a sketch of
the views of Mr. Stolzmann and those of Mr. Wallace.
Mr. Beddard’s objections to the use to which the theory of Natural
Selection has been put by various naturalists, and especially by Messrs
5« On the Position of Chamza in the System.” The Jour. of Morph., Vol.: iii,
maiaa a a eal of the Principal Facts and Theories Relating to
the Colors and Markings of Animals, by Frank E. Beddard, M. A. Oxon., F. R.
S. E., with four colored plates and wood cuts in the text. London, Swan, Sonn-
enschein & Co.; New York, Macmillan & Co., 1892. j i
872 The American Naturalist. [April,
Wallace and Poulton, are well taken. While he is far from denying
the efficacy of that process, he does not trace to it the origin of char-
acters, and he presents various instances where the latter seem to have
een the consequence of the direct action of physical causes. His
investigations are especially valuable because they cover a field which
has been especially cultivated by the advocates of the exclusive effi-
ciency of natural selection, and he thus throws new light on the class
of facts where the evidence for natural selection is strongest. For
this reason the work will well repay perusal, as it serves as a corrective
to thought which has, in the opinion of many, gone to an extreme in
one direction.
Ñ
Fic, 1. The Stoat in winter pelage, or Ermine.
The author cites the following remarkable example of the direct
effect of physical causes in producing changes of coloration :
“That the yellow color of canaries can be altered to an orange red
by mixing cayenne pepper with their food has been known for a long
ime. This curious fact was first discovered in England, as was also
the fact that the different races of canaries vary in their susceptibility
to the action of the pepper; some kinds are more, others are less
affected, while one race is absolutely without any power of having its
coloration altered by these means. The color change is produced by
feeding the newly hatched young with the pepper conveyed in their
food or the old birds while sitting upon the nest are furnished with
food containing the cayenne, which they in turn feed their offspring.
The color change can, in fact, be only brought about in very young
1893.] Recent Literature. 373
birds whose feathers are not completely matured ; it is quite impossi-
ble to produce any alteration upon the full grown canary. Clearly,
therefore, here is an instance of the direct effect of food upon color.
An interesting paper upon the subject, which has also furnished me
with the facts already mentioned,’ and it will be of interest to give
some account of. the author’s (Dr. Sauermann’s) experiments for
reasons that will appear. Cayenne pepper, of course, is a composite
substance, from which a number of distinct chemical substances can
be extracted ; the red color is caused by a pigment termed capsicin,
which can be separated from the pepper; and it might easily be sup-
posed that the change from yellow to red in the feathers of the canary
was simply caused by a transference of the pigment, as in the cases
mentioned on p. 127; but Dr. Sauermann shows that it is not so.
Yellow colored canaries were not in the very slightest degree affected
by the pigment alone; but, curiously enough, parti-colored birds did
react—the brown parts of the feathers became distinetly lighter in hue.
It is a fatty substance (triolein) which appears to convey the pigment
and produce thus a changing of the color from yellow to red; and
further experiments were made with other birds, showing that it is not
only canaries which are influenced by their food in this way. Some
white fowls, belonging to a special breed, showed traces of yellow
among the feathers after feeding with cayenne; but in this case these
were not racial, but individual differences in susceptibility, for all the
specimens of birds experimented with did not react to the stimulus.
“ A similar series of experiments was made with some other colors;
it was found with carmine that the yellow color was destroyed and the
birds became white. This unexpected effect is explained by the fact
that a mixture of violet and yellow produces white. The fact that the
fatty constituent, triolein, plays the chief part in the coloring of the
feathers may perhaps help to explain the very singular fact that the
Amazon parrots change from green to yellow when fed upon the fat of
certain fishes.
“With regard to the white fowls referred to, the experiments made
by Dr. Sauermann were particularly interesting. The interest lies in
the fact that the pigment was not absorbed equally by all the feathers ;
only special tracts were affected; the breast feathers, for instance,
became red, while the head remained white. It is therefore quite cred-
ible that in a state of nature partial alteration of color may be pro-
duced by a change of diet.”
*Archiv. Anatomie und Physiol., 1889. Physiol., Abtheil., 543.
374 The American Naturalist. [April
In a chapter relating to protective resemblances will be found an
account of several examples of animals which have apparently
acquired a resemblance to their surroundings by the transference of
pigment to their bodies in their food.
The list of illustrations comprises four very attractive colored plates
and thirty-six wood cuts, each of which repay study.
We reproduce three of these; one that represents color harmony
Fic. 2.
The Polytmus humming bird;
lower figure the male; the upper
the female.
Fics’ $
The male argus pheasant.
with the environment in the winter pelage of the stoats; and two
showing similar feather developments in the tails of the Polytmus
humming bird and of the Argus pheasant, in America and Asia
respectively.
1893.} Geology and Paleontology. 375
General Notes.
GEOLOGY AND PALEONTOLOGY.
Currents of the North Atlantic.—Prince Albert, of Monaco,
has recently published a chart of the currents of the North Atlantic,
entirely based upon the results of a series of experiments he has been
conducting since 1885. In that year he launched 160 floats at inter-
vals, along a line of 170 miles, in the direction of N. 14° W. from a
point 110 miles northwest from Cowo, one of the Azores. The follow-
ing year 510 floats were launched, nearly in the meridian 17° 40° west
of Greenwich, between 42° 34’ and 50° N. latitude. In 1887, 931
floats were launched along a line extending from the Azores to the
Grand Banks, of Newfoundland. During the same year, a group of
floats was launched between two other points in the North Atlantic,
between the Azores and Newfoundland.
Of the floats thus launched, 227 were sent back to the experimenter
with detailed information as to the place and date of finding. These
several histories gave indications of the direction and speed of the cur-
rents which floated them. The internal edge of the great vortex per-
forms a revolution with a very short radius round a point south-west
of the Azores, and not very distant from them. The cycle described
by objects drawn into the vortex is renewed indefinitely, except in the
case where they escape by an offshoot into the Arctic regions.
The mean velocity for the region comprised between the Azores,
Ireland, and Norway is 3.97 nautical miles in 24 hours; between the
Azores, France, Portugal, and the Canaries it is 5.18 nautical miles in
24 hours. From the Canaries to the West Indies, the Bahamas, and
even to Bermuda, it attains 10.11 nautical miles in 24 hours. In the
eastern portion of the arc, which extends from Bermuda to the Azores,
it falls again to 6.42 nautical miles in 24 hours. The mean velocity is
4.48 nautical miles in 24 hours. ( Proceeds. Roy. Geog. Soc., Sept.,
1892. )
Fins of Paleaspis Americana.— Mr. Claypole has recently
published the description of certain objects which he believes to be
the fins of Paleaspis americana. This fossil Mr. Claypole at first sup-
posed to be a spine, but upon examination it was found to be mar-
376 The American Naturalist. [April,
gined with a flat fringe around the pointed end and the rounded conical
parts were filled with matrix ; this central spine-like portion is covered
with a layer of the same tissue as that of which the shield is composed ;
the fringe around the conical central portion is distinctly striate in a
direction approximately parallel to its axis, These points of resem-
blance to a Crossopterygian fin, together with the fact that one of the
specimens was found so close to the side of the dorsal shield as to irre-
sistibly suggest a connection, have decided Mr. Claypole to add the
character of a fin to his amended definition of Paleaspis americana.
(Quart. Journ. Geol. Sur. 1892.)
New Reptiles from the Elgin Sandstone.—We have already
noticed the reptilian remains from the Elgin Sandstone, which are in
the condition of hollow moulds, the bones themselves having entirely
disappeared. Casts of these moulds have been made of gutta-percha,
which upon examination prove to be models of eight distinct skeletons,
of seven species of remarkable reptiles new to science. Professor E.
T. Newton, to whom the casts were sent for identification, refers 5 of
them to the genus Gordonia; G. traquairi (2), G. hualeyana, G.
dufiana and G. juddiana. In Gordonia, as in Dicynodon, the maxilla
is directed downward and forward to end in a small tusk. The for-
mer is distinguished from the latter by the presence of two post-tempo-
ral fossæ on each side of the occiput, by the small size of the maxillary
tusk, and by the slight ossification of the vertebral centra. A second
generic form, Geikia elginensis is nearly allied to Lystrosaurus Cope,
but is distinguished by its shorter muzzle and the entire absence of
teeth. The maxilla is produced into a tooth-like prominence, which
occupies a similar position to thetusks of Gordonia. For the remain-
ing species, Mr. Newton proposes the name Elginia mirabilis. The
exterior of the skull is covered in by bony plates, the only apertures
being the pair of nostrils, the orbits, and the pineal fossa. Horns and
spines, varying from } inch to 3 inches in length are found upon nearly
every bone of the exterior. This feature reminds one of the living
lizards Moloch and Phrynosoma. With the exception of a smaller
number of teeth, the dentition is a repetition on a large scale of Iguana.
This peculiar skull seems to show affinities with both Stegocephali
and Lacertilians, and is unlike any living or fossil form; its nearest,
though distant ally apparently being the Pariasaurus from the Karoo
beds of South Africa. (Nature, Dec., 1892. )
Fossil Reptiles from the Parana.—JIn the Boletin of the
Academia Nacional de Ciencias of Cordova (Argentina), Mr. Juan B.
1893.] Geology and Paleontology. 377
Ambrosetti describes three new species of fossil reptiles from the Oligo-
cene of the Parana. He also deserbies more fully three other species
already named by Bravard, Burmeister, and Scalabrini. The entire
number are referred to five genera, of which four are reported to be
‘new. The descriptions of species are quite full, but the new genera are
not described at all, so that they have, so far, no scientific standing.
One of them is said to be near to Tupinambis ( “ Podinema” ), and
the others are crocodiles. Three of them are regarded as near to the
gavials, and are an interesting addition to the South American fauna,
if this statement is confirmed. But the author is not yet fully compe-
tent for the task he has undertaken. He writes of the“ Ameivide,”
meaning Teidæ, and imagines that Rhamphostoma was proposed by
Burmeister in the Annals of the Museum of Buenos Ayres! His
knowledge of classic composition is indicated by his construction of
‘the abortion Oxysdonsaurus, of which an American grammar school
boy would not have been guilty. He should have written Oxyodonto-
saurus, a very awkward name at best. As the genus is not defined,
however, nobody is under obligation to adopt it.—E. D. C.
_ On the Systematic Position of the genus Protogonodon.
—The collection of fossil mammalia in the American Museum from
the Puerco formation contains among the numerous specimens a few
upper teeth, and a number of lower jaws belonging to the genus Proto-
gonodon, Scott! (— Mioclenus Cope, in part). I consider the material
referable to the genus Protogonodon as the most important in this col-
lection, as it enables me to probably settle the question as to its phylo-
gentic position in the system.
Professor Cope’s type of Protogonodon is a lower jaw with teeth. In
his description’ of this specimen, he considered it as probably closely
related to the Condylarthra, but as he was not familiar with the upper
molars, he placed it in the rather indefinite genus Mioclenus. Later —
Professor Scott? in his valuable paper on the Creodonta removed the
M. pentacus from Mioclenus, and established the genus Protogonodon
for its reception. Professor Scott places this genus among the Condyl-
arthra, and probably in the family Phenacodontide.
The series of upper molars in the collection which I refer to Proto-
gonodon were not found associated with lower teeth, so that their refer-
ence to this genus is not, absolutely certain. However, as the lower
1Proc. Philad. Academy, 1892, p. 322.
*Synopsis of the Vertebrate fauna of the Puerco series, 1888.
Op. cit.
378 The American Naturalist. April,
true molars of Protogonodon are of the sextubercular type, I believe
they should be associated with a superior molar which is tritubercular.
These upper molars consist of three principal cones with well devel-
oped intermediate tubercles; the external cones differ considerably in
form from those of the Creodonta ( Sarcothraustes ), but the general
form of the upper true molars in Protogonodon closely resembles that .
of the Creodonts. _
Another character of the teeth of Protogonodon which differentiates
it from the Perissodactyle line is the character of the inferior premo-
lars. The last tooth of this series is much simpler in structure than in
Protogonia, and closely resembles that of Pantolestes. It consists of a
simple cone with a slightly enlarged heel. In Protogonia puercensis
on the other hand, a well marked deuterocone is present, and this tooth
is nearly as complex as it is in Phenacodus.
_ I believe from these characters of the teeth, and also from the fact
that the lower jaw is elongated and slender, as in the genus Pantolestes,
that Protogonodon should be placed near the latter genus, and may
stand in ancestral relation to it.
Unfortunately the foot structure of Protogonodon is totally unknown,
so until that is discovered we cannot decide its affinity with certainty.
It is of great interest to note, if my supposition be correct, that in
the genus Protogonodon we are dealing with an ancestral Artiodactyle,
and that already as far down as the lowermost Eocene ( Puerco ) the
main types of the Ungulata were differentiated. It will not then be
necessary to resort as Schlosser has done to derive the Artiodactyla -
from any of the known Periptychide, but that the Perissodactyla were
represented by Protogonia,‘ the Artiodactyla by at least Prutogonodon
and perhaps other genera, as suggested by Professor Scott. Lastly a8
Professor Cope has shown the Amblypoda were already evolved and
represented by the genus Pantolambda.
Whether or not these Ungulate stems were in all cases distinct from
the Condylarthra, remains for future research to prove; but at least
one of them, the Amblypoda, was differentiated. In the case of Proto-
gonodon the structure of its teeth shows that it was fairly well on the
Artiodactyle line, but the discovery of its foot structure will demon-
strate whether or not it had passed the Condylarthrous stage.
I believe that among the Puerco Condylarthra that Protogonia was
a persistent type, and nearly all of the Periptychids with the possible -
‘Professor Cope informs me by letter that the name Protogonia is preoccupied,
and he proposes as a substitute the name Euprotogonia.
1893.] Geology and Paleontology. 379
exception of Mioclenus turgidus died out and left no descendants.—
Cuas. EARLE.
_ American Museum of Natural History, New York.
. The Brown Coal and Lignite of Texas.°—Mr. E.T. Dumble’s
comprehensive report on the fossil fuel supply of Texas has been issued
as a separate publication by the Geological Survey of that State. In
the course of his investigations, Mr. Dumble found it necessary to visit
the mines and factories of Austria and Germany, where by personal
observation he ascertained the character, use, and value of lignites,
together with the best methods of utilizing this fossil wealth. His
report is complete, both froma geological and an economic standpoint.
Geologically, he describes the origin, formation, physical and chemical
character, and the occurrence of the brown coal. Incidentally a clas-
sification of the Texas brown coals is given. As an economist, Mr.
Dumble suggests the particular uses to which the varieties of brown
coal may be put, discusses the valuable products that may be obtained
from it, and gives in detail the most approved methods of making it
into artificial fuel by brignetting. In a word, Mr. Dumble has demon-
strated that in her brown coals and lignite, Texas has an abundant
and cheap fuel supply.
The text is illustrated with a number of cuts of machinery, and an
excellent map.
Extinct Fauna of Mauritius.—At the meeting of the Zoological
Society, of London, Nov. 1, 1892, a communication was read from Sir
Edward Newton and Dr. Gadow, describing a collection of bones of
the dodo and other extinct birds of Mauritius, recovered from the
Mare aux Songes in that island by the exertions of Mr. Theodore
Sauzier. The collection contained examples of the atlas, metacarpals,
prepelvic vertebre and complete pubic bones of the dodo, which have
heretofore been wanting, as well as additional bones of Lophopsittacus,
Aphanapteryx and other forms previously known to have inhabited
that island. Besides these there were bones of other birds, the exist-
ence of which was not before suspected, and these were described as
belonging to the genera Strix, Astur, Butorides, Plotus, Sarcidiornis,
and Anas.
5Report of the Brown Coal and Lignite of Texas. Character, Formation, Occur-
rence and Fuel Uses. By E. T. Dumble, F. G. S. A., State Geologist.
380 The American Naturalist. [April,
MINERALOGY AND PETROGRAPHY:'
Description of the New Rock Type, Malchite.—The new
rock, malchite, referred’ to a few months ago as the granitic dyke form
of diorite, is now described in some detail by its discoverer, Osann
It forms dykes cutting granite in the Odenwald, Germany. Ina dense
groundmass are rare phenocrysts of dark mica, pale green plagioclase
and quartz. The mica is biotite and the plagioclase labradorite. In
addition to these the microscope reveals the presence of idiomorphic
green hornblende, allanite and sphene. The groundmass in which
these lie strongly resembles that of some tinguaites, with hornblende
and quartz in place of aegerine and nepheline. It consists of a fine
granular aggregate of feldspar and quartz, the latter with occasional -
idiomorphie contours, and prisms of hornblende imbedded in the
aggregate, the prisms often arranged in flowage lines. An analysis of
a fresh specimen of the rock yielded:
_ SiO, AI,O, Fe,O, FeO MgO CaO Na,O K,O.H,0 SO, P,O, Total
63.18 17.03 .24 637 92 417 4.44 2.91 .52 .19 .283—100.20
The Petrography of Hokkaido, Japan.—In a general geo-
logical sketch of Hokkaido, (Jezo or Yesso), Japan, Jimbo‘ declares
that the island consists largely of paleozoic beds, probably underlain
by amphibolites and various other schists, and cut by granite, diorite,
gabbro, peridotite, and serpentine. In the lower portion of
the paleozoic the beds consist largely of pyroxenites, with traces of rad-
iolarian remains, phyllites, quartz-schists, limestone, and serpentine.
The pyroxenites are aggregates of light colored augite, quartz and
feldspar, in which the augite is often more or less changed to epidote
and glaucophane. Where the granite cuts the clastics the clay slate
is changed by contact action to a biotitic clay slate, to hornfels and to
mica schist, with the latter nearest the eruptive. Tourmaline occurs in
the schist and cordierite in this rock and in the mica slate. An
amphibolite in the contact belt is supposed to be an altered tufa.
Schistose granites, diorites and gabbro are phases of the corresponding
- Edited by Dr. W. S. Bayley, Colby University, Waterville, Me.
s CAN NATURALIST, May, 1892, p. 422.
> . 380.
‘General Geological sketch of Hokkaido, with special refer€nce to the petrography.
Satporo, Hokkaido, Japan, 1892.
1893.] Mineralogy and Petrography. 381
massive rocks associated with the contact products. Diabases occur as
sheets in the unaltered paleozoic beds, and serpentines derived from
gabbros and from dunites are met with cutting these at various locali-
ties. In addition to paleozoic there are also tertiary rocks on the
island, and these are cut by their own systems of dykes and bosses, and
are interbedded with characteristic sheets of lava, and layers of tufas.
The tertiary voleanic rocks are pyroxene and hornblende andesites,
propylites and rhyolites. The pyroxene andesites contain both ortho-
rhombic and monoclinic pyroxenes and occasionally some olivine.
They have also a glassy base which sometimes becomes so abundant as
to 'resemble pumice. The hornblende andesite is strongly porphy-
ritic with large phenocrysts of hornblende. The rhyolites are both
compact and glassy, in which latter case they are vesicular.
Two Peculiar Rocks from Siberia.—Two very remarkable
rocks are described by von Chrustschoff* from Taimyr-Land, Siberia.
One is an ophitic aggregate of anorthoclase and nosean, containing as
accessories sanidine, plagioclase, amphibole, biotite, melanite, magne-
tite, sphene, zircon and glass. The anorthoclase is in long, narrow
crystals of the following composition :
SiO, AlO, FeO, CaO MgO K,O NaO Total
64.59 19.84 2.24 1.26 .63 3.3 7.88=99.97
Corresponding to Or, Ab, An, The feldspar is usually idiomorphic
with respect to the nosean, whose period of formation was between
that of the biotite and that of the hornblende. The nosean is in very
large quantity. Its density is 2.266 and composition :
SiO, AlO, FeO, NaO K,O CaO H,O Cl SO, Total
37.83 2659 .38 2240 1.63 .54 .87 1.66 8.68 — 99.98
The zircon is of the trachytic type, and is the only accessory of any
importance. The author calls the rock taimyrite. The second rock
is composed of anorthoclase, sanidine, biotite, and amphibole as essen-
tial components, and the other minerals mentioned above in connec-
tion with taimyrite as accessories, except that sodalite here replaces
nosean. The zircon is of the granitic type, and the rock possesses the
granitic texture.
5Bull. d. Acad. Imp.des Sciences St. Petersb. Mél. Geol. et Paleont., i, p. 153.
382 The American Naturalist. [April,
An Ottrelite Bearing Conglomerate in Vermont.—It is not
uncommon to find ottrelite forming 25% of the schistose groundmass
of the conglomerate? at the base of the Lower Cambrian, near Rut-
land, Vermont. The same mineral occurs along shear planes in a
blue quartzite and constitutes 40% of a massive bed of the conglom-
erate. In the last named rock the ottrelite is in rudely circular areas,
lying in a dark colored quartz. The areas consist usually of radiating
plates of the mineral, disposed in a single plane. Its commonest
inclusions are quartz and feldspar, while sericite often forms the cen-
ters of the radiating bundles. In the latter case the ottrelite is
oriented in parallel position with the mica. Other inclusions within
the ottrelite besides those above mentioned are crystals of zircon and
rutile, flakes of graphite and plates of ilmenite. In other cases the
ottrelite is in plates including large areas of the groundmass of the
rock, which is a granulated mixture of quartz and albite (?) in about
equal proportions, a large quantity of sericite, and some biotite. In
this groundmass associated with rutile are crystals and plates of ana-
tase. No traces of its original clastic structure remain in the rock,
though its conglomeratic character is beyond dispute.
Lithophysz in the Rocche-Rosse.—In parts of the Rocche-
Rosse lava stream of Monte Pelato, Lipari, are spherulites with litho-
physal characteristics. In some specimens examined by Cole and
Butler’ the spherulitic growth originated about the walls of steam
vesicles, and progressed outward into the rock; in other cases they
grew inward until they have completely filled the space that was for-
merly vacant. The importance of the paper lies in the fact that it
acknowledges the correctness of many of Idding’s views with respect
to the formation of lithophysz, and contradicts the view that regards
all hollow lithophysz as the result of the decomposition of spherulites.
The Composition of the Dune Sands of the Netherlands.
—A very elaborate paper by Retgers® on the constitution of sand
composing the dunes on the west coast of Holland at Sheveningen,
near the Hague, contains a large amount of information concerning
the character of sands and the method of determining the nature of
their constituents. The author carefully fractioned large quantities o of
the dune sand by the ordinary methods of fractional precipitation in
SC. L. Whittle, Amer. Jour. Sci., Oct., 1892, p. 270.
"Quart. Jour. Geol. Soc., xlviii, 1892, p. 438.
"Recueil des Travaux Chimiques des Pays-Bas., xi, 1892, p. 169.
1893, ] Mineralogy and Petrography. 383
the usual heavy liquids and by means of the molten substances sug- '
gested by himself’ for this purpose a few years ago, thus obtaining
mixtures of mineral grains of about the same density. These then
were studied carefully by comparison of their indices of refraction, by
immersing them in liquids of known optical densities, until one was
found in which the grains became almost invisible. The index of
refraction of these is nearly that of the liquid, consequently their
nature is thus approximately determined. Microchemical tests and
the ease with which cleavage laminze were produced, served to distin-
guish accurately between minerals having nearly the same refractive
index. The principal minerals identified by the author are ortho-
clase, quartz, microline, plagioclase, cordierite, calcite, apatite,.amphi-
bole, tourmaline, pyroxene, epidote, sphene, sillimanite, olivine, gar-
net, staurolite, disthene, corundum, spinel, rutile, zircon, magnetite and
ilmenite. The surprising discoveries are those of cordierite, calcite
and olivine, and of glaucophane among the amphiboles. The propor-
tions of the various minerals present according to specific gravity was
2.5% between 2.5 and 2.6; 85% between 2.6 and 2.7; 7.5% between
2.7 and 3. ; 1.5% between 3 and 3.3; 1% between 3.3 and 3.6; 2.4%
between 3.6 and 4.2; .1% between 4.2 and 5.2. The sands are sup-
posed to have come mainly from the rocks of archean terraces.
Quartz-Gabbro in Maryland.—In the Baltimore gabbro area,
according to Grant,” are quartz. gabbros consisting of bytownnite,
quartz, hypersthene, secondary hornblende, and a few accessories.
The quartz is limpid, and is almost free from inclusions, except for
lines of small liquid cavities that traverse the grains, as is usual in
granitic quartz. Diallage, which is so common in the normal gabbro
` of the region, is entirely absent from the quartz-bearing phases, which
thus becomes a quartz norite.
Minerals from the Diamond Fields of Brazil.—Hussak"
describes the characteristics of crystals of brookite, cassiterite and
-Xenotime from the diamond region of Dattas, Minas Geraes, Brazil.
On brookite from the sands of Diamantina was found the new pyramid,
4P%. The cassiterite is from Manquinho, near Sao Paulo. It occurs
in a rubellite-bearing lepidolite granite. The zxenotime accompanies
the brookite in the sands of Dattas. On one doubly terminated crys-
9Cf., AMER, NATURALIST, 1890, p. 175.
Johns Hopkins Univ. Circ. No. 103.
"Min. u. Petrog., Mitth. xii, p. 455.
384 The American Naturalist. [April,
` tal were found the two new pyramids 4P and iP. The axial ratio of
these crystals is 1: .61775.. The author has also made a crystallo-
graphic examination of the monazite occurring so abundantly in the
Brazilian granites and gneisses. The crystals of this substance are
always tabular parallel to « Pæ. They contain the same forms as
do the Ilmengebirge crystals, but are never twinned. Upon washing
a portion of sand from Bohia a 3 mm. long crystal of euclase was
obtained whose density is 3.1. It is very rich in planes, being pos-
sessed of not less than three prisms, six clinodomes and three negative
pyramids, beside the clinopinacoid.
Mineralogical Notes.—Christianite crystals are reported by
Gonnard” as lining geode cavities in the basalts of dykes at Quey-
riéres and Fay le Froid, Haute Loire, France. In the latter case the
christianite groups enclose many crystals of augite. The trachyte of
Montcharet, occurring as a dyke in granite, is cut’ by fissures whose
walls are lined by chabasite.* The cubic faces of galena crystals
implanted in druses of quartz at Pontgibaud are roughened by little
cavities whose walls have the positions of octahedral planes. The
phenomenon is regarded by Gonnard" as the result of corrosion. The
same author mentions the existence of large crystals of beryl in the
granites of Droiturier, near La Palisse, Allier, psilomelane in mam-
millary forms at Croix Moraud, Mt. Doré, and cubic pseudomorphs of
artz after some unknown mineral, probably fluorite, in the vicinity
of d’Aubenas, Ardéche.
Three specimens of melilite from Mt. Somma, with densities of 2.917,
2.932 and 2.945 respectively, were powdered, purified, and analyzed by
Bodlander” with this result :
SiO, AlO, FeO, CaO MgO K,O NaO H,O Total
41.34 10.37 429 33.84 5.79 113 3.45 .08 — 100.29
The author combats the view of Vogt that melilite is an isomor
phous mixture of the gehlenite and akermanite molecules. He thinks
that the negative variety is an admixture of R”SiO, and the alumin-
ate R”,0,R”, while the positive variety is a compound of the same
silicate with the aluminate R,’”O,R,”. Intermediate varieties are iso-
morphous mixtures of these.
“Bull. Soc. Franç d. Min., 1892, xv, p. 28.
“by p. 31.
MTb., p. 34.
Neues. Jahrb. f. Min., ete., 1893, i, p. 15.
See
Sy
1893.] Mineralogy and Petrography. 385
Moses" records the analysis of a granular nickel arsenide associated
with native silver and siderite in a mine 18 miles west of Silver City,
N. Mex. The silver is imbedded in arborescent forms in the brittle
gray nickel ore, and this in turn is in a gangue of siderite. The
analysis made on impure substance gave:
je Pb Ag As Ni Co. Fe _ Total
tr. 8.383 67.37 11.12 5.13 2.64=— 99.20
Regarding the SiO, and Ag as impurities the iat aig takes a form
that may be represented by RAs, in which R == 4 Ni 3 Co and } Fe,
corresponding to a nickel skutterudite.
On crystals of topaz from the Province of Omi and from the tin
mines of Yenagari Mino, Japan, Matthew" finds four pyramids, seven
prisms, one of which, æ Py, is new, the three pinacoids, three brachy-
domes and two macrodomes.
Optical Anomalies. — After an exceedingly careful examination
of many sections of appophyllite crystals and a comparison of the phe-
nomena they present with those presented by combinations of thin
biaxial plates placed one upon the other, Klein concludes that the
mineral in its geometrically tetragonal crystals is an intimate mixture
of optically positive and optically negative triclinic lamelle. The
positive constituent seems to differ from the negative element in con-
taining no crystal water, since upon heating the positive component
appears to increase in quantity. Negative appophyllite becomes posi-
tive upon loss of 43 molecules of crystal water. The investigation is
a beautiful piece of accurate optical work.
In a reply to Mallard’s” remarks on the black -garnet pyrenaite
Brauns” states that the structure described by the first mentioned
anthor is exactly what should be expected of a dodecahedral substance
under strain, and that the peculiarities of this garnet’s optical proper-
ties may be easily explained on the Klein-Brauns theory of strain.
Upon soaking in oil sections of zeolites that have been rendered
cloudy by loss of water, they again become Snoen transparent for
the study of their optical properties. Rinne” has taken advantage of
School of Mines Quart., xiv, No. i, p. 49.
"Iib, xiv, No. 1, p
18Neues. Jahrb. f. Min., etc. oe II, p. 165.
19AMERICAN NATURALIST, Oct., 1892, p. 849.
_ Neues. Jahrb. f. Min., w, sens E pe 287.
™Ref. ar Jahrb. f. Min., etc., 1892, II, p. 237.
386 The American Naturalist. [April,
this phenomenon and has carefully examined a number of the mem-
bers of the group with a view to learning something of the changes
effected in them by the loss of water. Natrolite appears monoclinic
under these conditions, scolecite orthorhombic, stilbite orthorhombic,
and each of the other zeolites affords a corresponding meta-zeolite.
The optical anomalies often observed in these minerals is thought to be
undoubtedly due to partial loss of water.
Isomorphism.—<After a long mathematical discussion of the
theory of the structure of isomorphous mixtures and upon compari-
son of the results of investigations upon the optical properties of
mixed crystals, Poeckel™ concludes that we have not yet sufficient
data to decide as to whether Mallard’s lamellz theory of the constitu-
tion of these bodies is correct or not.
By the use of the method” in which colored and colorless crystals
of supposed isomorphous substances are allowed to form under the
microscope Retgers™ has proven that the alkaline ferrates are isomor-
phous with the corresponding sulphates, selenates, molybdates and
tungstates, that the potassium tellurates and osmiates are isodimor-
phous, and that the rutheniate of this metal is isomorphous with its
uranate.
The arguments for and against the view as to the isomorphism of
calcite and dolomite are given respectively by Brauns and Retgers”
in a recent letter to the Neues, Jahrbuch. The discussion is too
involved to warrant an intelligible abstract in these notes.
Etched Figures.—The matrix of the African diamonds is capable
of resorbing” diamonds, producing on their faces irregular, long, and
hemispherical hollows, associated with which are little spheres and
grains of black carbonaceous substance, supposed to be a compound of
iron and carbon.
Hofer” describes corrosian forms on the calcites of Steierdorf, Banat,
and of Rauris and Salzburg, and ascribes the hexoctahedral faces
*#02 on the fluorite of Sarnthal, Tyrol, to corrosive processes.
™Neues. Jahrb. f. Min., etc.,-B. B., viii, 1892, p. 117.
AMERICAN NATURALIST, June, 1892, p. 517.
*Zeits. f. Physik. Chem., x, 5, 1892, p. 529.
Neues. Jahrb. f. Min., etc., 1892, II, p. 210.
%Ber. deutsch. chem. Ges., 1892, p. 2470.
"Min. u. Petrog., Mitth. xii, p. 487.
1893.] Mineralogy and Petrography. 387
Microchemical Reactions.—The methods of testing for traces
of ammonia under the microscope, and ri prempteres metals with
HS are described in a few words by Stre
Directions for the detection of the ERA minerals in small par-
ticles are given by Lemberg:” Scapolite, hauyne, sodalite, eudialite,
lazurite, sulphur, okoia celestite and melilite.
Mincellasiwedea- iai the title “Rapid Qualitative Examina-
tion of Mineral Substances,” Moses and Wells” publish a scheme for
the detection of minerals. The blowpipe method is used with the
metallic minerals, but in the silicate group a mixture of the dry and
wet methods is made use of. From a hasty reading of the scheme it
seems to be a practicable and convenient one.
Neues. Jahrb., 1893, I, p. 49.
*Zeits. d. deutsch. geol. Ges., 1892, p. 224.
School of Mines Quart., Nov., 1892, p. 25.
388 The American Naturalist. ‘[April,
ZOOLOGY.
= The German Zoological Society.—The annual report of this
society for 1892 is out, and contains among others the following
The opening address by the President, Professor F. E.
Schultze, described the zoological department of the University of
Berlin, where the meeting was held, giving also a sketch of its history.
Professor H. E. Ziegler spoke upon the embryonal anlage of the blood
in the vertebrates. At first the blood is a clear serum in which no
corpuscles appear. Later they are present. Various ideas have been
advanced as to their origin. Ziegler vigorously discusses the view
which assigns them to the entoderm and decides that they are without —
doubt mesodermal in origin. Dr. L. Plate described the anatomy and
affinities of the Onchidiide. These forms belong to the Asiatic region,
and are amphibious, living equally well in or out of the salt water.
To separate the genera the author uses the relative width of the foot
and of that suprapedal region to which he gives the name of hypono-
tum. The group is of great interest from the fact that though usually
regarded as Opisthobranchs they must be considered as near the
ancestral Pulmonate. Plate, however, thinks that they have crossed
the line and are true Pulmonates, but that their Nudibranch charac-
ters are such as cannot be explained except upon the hypothesis of
descent from that group. Dr. Thiele, in discussing the phylogeny of
the byssus apparatus of the Lamellibranchs, thinks that it must be
traced back to some such type as the adhesive glands of Haliotis, and
does not agree with Pelseneer in regarding the byssal glands of the
Nuculide as the most primitive. The fact that either in young or in
adult the byssus occurs in most Acephals, is proof to Thiele that the
gland must be traced back to the ancestral mollusc. Von Nathusius
discussed the form and color of the hair as a basis of classification of
the Equidæ. Dr. Korschelt presented a paper on the differentiation
of the germ-layers in the Cephalopods with especial reference to the
formation of the alimentary canal and the nervous system. Accord-
ing to Bobretzky both the midgut and the nervous system are of meso-
dermal origin, while several other authors think that the whole
alimentary canal is formed by ectodermal inpushings (stomo- and proc-
todeum). Korschelt finds' that at an early date an epithelial layer
arises just above the yolk in the anal region. It is distinctly separa
1The full paper with illustrations appears in the Leuckart Festschrift.
E
5s Se
SERTE
x
E T ENN Hai ss 5i
Pe
Sn Re eS eg ee te a eh a ty) he et
1893.] ; Zoology. 389
from the ectoderm by the mesoderm. Later it becomes elevated
from the yolk, forming a small cavity open to the yolk, the future
mesenteron. This layer is regarded by Korschelt as entoderm, and has
been regarded by several authors as the proctodeal inpushing. He also
finds the nervous system to be, as might have been expected, ectoder-
mal. Dr. Burckhardt described the central nervous system of the
Dipnoi. He has subjected the brain to sections, and by study of these
has been able to elucidate points not before understood. He concludes
that in its brain Protopterus is not an Amphibian, but that it is in this
respect a problematic organism between the fishes (Elasmobranchs)
and Amphibia, whose exact position will first be solved when the
ontogeny is studied. Professor R. Hertwig had a review. of recent
work upon canjugation and fertilization, and Dr. H. H. Field a paper
on the Metamerism of the Kidney of Amphiuma, already noticed in
our pages. The last paper was a’ view of the idea of instinct, by Pro-
fessor H. E. Ziegler. |
Eyes of Polychætes.—Dr. E. A. Andrews has an extended
article? on this subject. He has studied a large series of genera, some
both as young and adult, and among the conclusions at which he
arrives are the following: The eye is an epidermal organ consisting of
a pigmented cup containing a refracting mass, which at the orifice of
the cup comes in contact with the cuticle. The wall of the cup con-
sists of a single layer of cells forming the retina, each retinal element
containing pigment, and being produced internally into a nerve pro-
cess, externally into a clear rod. This, as will be seen, is a different
conception of the annelid eye from either that of Carriére or that of
Graber, but one which approaches most nearly to that of the former
author.
Arachnida.—Mr. B. I. Pocock discusses’ the external anatomy of
the members of the group Arachnida (s. s.), and concludes that the
members must be arranged in two sub-classes, Ctenophora, with the
single order Scorpiones, and Lipoctena, containing the superorders
Caulogastra (Pedipalpi and Aranez) Mycetophora (Solifuge), and
Holostomata (Pseudoscorpiones, Opiliones, Acari).
F. Purcell, after studying the anatomy and development of the eyes
of Liobunum hemisphericum, concludes‘ that contrary to the com-
2Jour. Morph., vii, 169, 1892.
3Ann. and Mag. Nat. Hist., VI, xi, p- 1, 1893.
4Zool. Anz., xv, 1892.
390 The American Naturalist. [April,
monly accepted ideas, these organs in the Phalangids are to be regarded
as compound, and further that they are inverse eyes homologous with
the anterior middle eyes of spiders and the middle eyes of the scor-
pions.
The Fishes of the Pacific Coast of America North of
Cerros Islands.—An important paper recently published is an
enumeration of the fishes occurring on the Pacific coast of America
north of Cerros Island, and to a depth of 150 fathoms by C. H. and
R. S. Eigenmann. The authors have placed the dividing line between
the littoral and the bathybial faunas of this region at 150 fathoms,
because all of the genera so far recorded from this depth have repre-
sentatives in the shallower water. The few littoral genera which have
representatives in deeper water are added as footnotes. Cerros Island
is a natural boundary to this region, since south of it few of the char-
acteristic genera of this region are found. The author’s observations
show that the old division, at Point Conception, of the northern and
southern California fauna is incorrect, and that, in reality, no definite
boundaries can be assigned for a northern and southern fauna.
The California fauna is distinct from that of Alaska, although four
of the species found at San Diego are found in Alaska. The Califor-
nia Fauna is characterized by the abundance of species of Sebastodes,
of Cottidæ, and of Embiotocidæ. The last are entirely absent from
Alaska, and only a few species of Sebastodes are found there. The
boundary between these two regions lies somewhere between Sitka and
Puget Sound.
The relative number of species at the principal localities is as fol-
lows :
Species.
The whole of Alaska 108
Fuget Sound: «acs cnssastccroveaatreassciiy eee a be 107
Han: PERE OIIOD ioanen a a 156
Monterey 148
Santa Barbara 117
San Pedro 82
San Diego, including Cortes Banks , 168
There are known from the entire region 382 species, belonging to
228 genera. Of these 116 genera, or more than half, are also found
in the Atlantic Ocean, and 32 species are found both in. the Atlantic
and in the Pacific.
- 1893.] Zoology. 391
The genera having species in both oceans practically all belong to
one of three classes: 1st, Tropical genera; 2d, Arctic genera; 8d,
Pelagic and other genera (Ann. N. Y. Acad. Sci., vi, 1892).
On the Mechanical Genesis of the Scales of Fishes.—
Mr. J. A. Ryder has advanced a theory that the genesis and arrange-
ment of the scales of fishes is due to the movements of the fishes in
swimming. The scales originatein a continuous subepidermal matrix
which may be regarded as a basement membrane. Such a matrix is
found to actually exist in some forms at an early stage, just beneath
the epidermis; in other forms it exists in the larval stages, and in the
young of many types of fishes it is coextensive with the whole epider-
mal layer just at the time when the scales are forming.
In the Clupeoids and Cyprinoids it is found that the myocommata,
or sheets of connective tissue intervening between successive somites
are attached firmly to the deeper layers of the skin. This construc-
tion, together with the peculiar arrangement of the muscle plates at
the time the scales begin to form, conditions the further growth of the.
scale matrix. The ordinary movements of the fish in swimming
throw the whole integument into definitely circumscribed areolæ, of
which the central portions are in a quiescent state, while the margins
are wrinkled or folded as a result of the current action of the lateral
muscles of the body.
` In each of the circumscribed areolz a scale develops; the continuity
of its development with its fellows across the margins of the areola is
prevented by the flexures to which the dermis is there subjected.
The author shows that the imbrication as well as the arrangement of
the scales is determined by the action of the segmentally arranged
muscles of the body.
Special types of squamation, as those of the eel and the sturgeon,
confirm the author’s views. In cases where the scales are very fine,
that is, several oblique rows corresponding to each somite, the principle
holds good, since the rows still conform to the lines of tetision of the
linear attachment of the myocommata to the integuments.
From the hypothesis and the evidence presented, Mr. Ryder draws
the following important conclusion :
1. “ The scales of fishes bear a segmental relation to the remaining
hard and soft parts, and are either repeated consecutively and in
oblique rows corresponding to the number of segments, or they may
be repeated in rows as multiples of the somites, or segmental reduction
may oceur, which may affect the arrangement of the scales so as to
392 The American Naturalist. ‘[April,
reduce the number of rows below the number of somites indicated by
the other soft and hard parts.
2. “ The peculiar manner of interdigitation of the muscular somites
as indicated by the sigmoid outline of the myocommata, as seen from
their outer faces, and the oblique direction of the membranes separa-
ting the muscular cones, has developed a mode of insertion of the
myocommata upon the corium which has thrown the integument into
rhombic areolæ during muscular contraction. These areole are in
line in three directions, and the folds separating them, particularly at
their posterior borders, are inflected in such a manner by muscular
tensions, due to the arrangement of muscular cones, as to induce the
condition of imbrication so characteristic of the squamation of many
fishes.” —( Proceeds. Phila. Acad., Part II, 1892.)
The Systematic Position of the Kiwi.—In a paper on the
History of the Kiwi, Professor T. Jeffery Parker gives the following
conclusion as to the systematic position of this species :
“On the whole it wil be seen that the study of the development of
the kiwi tends to lessen the gulf between it and ordinary birds, and
to show that its ancestors probably possessed many of the more impor-
tant and distinctive features which characterize the Carinats of to-day.
The facts clearly indicate that the founder of the Apterygian house
had interrupted plumage, functional wings, an ordinary avian tail, a
keeled sternum, a double-headed quadrate, lateral optic lobes, and a
pecten in the eye; in other words, that the ancestors of the genus were
typical flying birds and not bird-like reptiles. a
“Of the eight characters enumerated above as separating the Ra
titze from the Carinatæ it will be noticed that the first five are directly —
connected with the power of flight. We should expect to find such
adaptive characters in purely cursorial birds whether they arose from
a common stock or sprang separately from early flying birds, and’ as
a matter of fact they occur to a greater or less extent in such flightless
birds as the Dodo, Weka, Notornis, etc., which we know have no gen-
etic connection with one another, but have independently acquired the
characteristics of flightlessness. I think, therefore, that the possession
of the characters referred to by the whole of the Ratite is no argu-
ment for their common origin.
“The peculiarity of the quadrate has been shown to be a secondary
matter, and we have left only the characters of the base of the skull.
These certainly form an excellent diagnostic character by which the —
whole of the Ratitæ are separated from the majority of the Carinate,
t
Ee
1893.] Zoology. 393
but even here the distinction is not absolute, for the Tinamous approach
in many respects more nearly to the Ratite than to the rest of the
Carinatæ. Still it seems probable that the various genera of Ratite
must have diverged from the main line of descent at a comparatively
early period, though perhaps not earlier than some of the existing
orders of Carinatz. The Penguins, for instance, are far more reptilian
in their vertebral column and less typical in the structure of their
wings than the Ratite. The Ostrich, however, shows the unique and
very reptilian character of two claws on the wing, and the very gen-
eral presence of wing-claws in the group is a distinctly primitive
character.
“ Leaving the skull, in which the whole group shows primitive char-
acters, and the wing and related parts, in which the resemblances
between the genera are largely adaptive, we find the range of variation
in the Ratitæ to be very great indeed. Two genera (Apteryx and
Dinornis) have a normal four-toed foot; in three others (Cassowary,
Emu and Rhea) the hind toe or hallux has disappeared ; while in
another (Ostrich) only two toes are left. The pelvis of the kiwi and
moa is of the simplest avian type, both pubis and ischia being free;
in the cassowary and emu the ischium unites with the ilium ; in the
rhea the ischiæ unite with one another above the intestines—a unique
arrangement ; in the ostrich the pubes unite to form a symphysis as
in most of the higher vertebrates. The feathers have an aftershaft in
the emu, cassowary, and moa, none in the ostrich, rhea, or kiwi. In
no order of carinate birds do we find such a wide range of variation
as this, and when we add to the characters enumerated the extraordi-
narily aberrant skull and the structure of the egg shell of the Apteryx,
the total atrophy of the wings in Dinornis, and even of the shoulder-
girdle in some species of the genus, and the striking differences between
the sterna, the shoulder-girdle and the wings of the various genera, we
are forced to the conclusion that the existing or lately extinct cursorial
birds now known to us are divisible into five well-marked orders, each
the equivalent of an entire order of Carinate. Of these one order
contains the ostrich alone, another the rheas, a third the emu and the
cassowaries, a fourth the moas, and a fifth the kiwis. ;
“« As to the relation of the kiwi to the other genera it has been shown
to be most nearly allied, as far as its skeleton is concerned, to the moa,
differing from it, however, in many important respects. It must cer-
tainly have been isolated at a very distant period, and as far as we can
sée, some of its more striking peculiarities are distinctly correlated to
its method of feeding. Most nocturnal animals have large eyes suited
394 The American Naturalist. [ April,
for taking the utmost advantage of the semi-darkness, but the kiwi, `
finding its prey by scent alone, has developed an extraordinarily per-
fect olfactory sense, while at the same time, having no need to keep
watch against beasts of prey, its eyes have diminished in size and
efficiency to a degree elsewhere unknown in the bird class.”—(New
Zealand Jour. Sci., March, 1891.)
Mammals from the Galapagos Islands.—The mammals col-
lected by Dr. Baur on the Salisbury Expedition to the Galapagos
Islands were sent to Mr. J. A. Allen for identification. The collection
numbers 12 specimens, representing 4 species, of which 2 prove to be
_ introduced species of Mus. The other species are both new, one a bat,
Atalapha brachyotis, the other a mouse, Oryzomys baurii, closely related
to O. galapagéénsis. The only other species of mammals thus far
reported from the Galapagos Islands are two species of Eared Seals,
Otaria jubata (Forst.) and Arctocephalus australis (Zimm.), which
were obtained by the Hassler Expedition. The list of known indige-
nous species thus numbers only five—(Bull. Am. Mus. Nat. pki
Vol. iv.)
1893.] Embryology. 395
EMBRYOLOGY.’
Echinoderm Eggs.—Karl Fiedler’ endeavored to corroborate the
work of Roux by experiments made at Naples upon the eggs of three
species of sea urchin. At first the method used was that of Roux,
piercing the egg with a needle. It was found necessary, however, to
use a lancet shaped needle made by hammering out and sharpening the
tip of a common needle. Later the method of shaking employed by
the Hertwigs and by Boveri was, for the first time, adopted in separating
the cleavage cells.
The use of the needle under the microscope was attended with a
great many difficulties, increased by the large mortality amongst the
few individuals that could be successfully operated upon. After the
membrane is quickly cut the cells may be individually punctured
and destroyed. Such embryos can be kept only in large vessels with
algee.
By this means it was possible to pierce a cell so that some of the
contents ran out without destroying its power of cleaving, providing
the nucleus remained ; but in other cases where the nucleus escaped,
even with but little protoplasm, the cell died. In the cases where
cleavage continued the diminished cell gave rise to small progeny ; thus
in the four cell stage two cells were much smaller than the others; in
the sixteen cell stage two of the four polar micromeres were plainly
smaller than the others; in the blastula one half was less convex than
the other; but later the difference seems to have been equalized.
When cells are separated by shaking, the remaining ones may change
their shape, becoming more spherical where no longer in contact with
other cells, but they retain their normal position much as if the other
cells had not been removed. Thus when one of the first two cleavage
cells was destroyed there resulted an eight celled stage that was half
the normal sixteen celled stage, having half the normal number, four,
of micromeres at one pole and the other cells likewise arranged as if
the sixteen celled stage had been cut into two. The same was true of
the half-twenty-eight cell stage. Such half embryos can be formed
either by destruction of one of the first two cells or by destruction of
two of the first four. The same result is produced by destroying any
two of the first four cells, whether they are sister cells or first cousins.
1This department is edited by E. A. Andrews, Johns Hopkins University.
2Entwicklungsmechanische Studien. Festschrift fiir Nägeli, Zurich, 1891.
396 The American Naturalist. [April,
The first four cells are thus all alike in their possibilities as well as
in appearance ; not so the first eight cells, though they all look alike.
When the eight cells are separated into groups of four these give
rise to quite different sets, some with eight large, others six large and
two little cells, others four large and four small cells. Thus the devel-
opment of the eight cell stage is largely a process. of “self differentia-
tion ” of these cells and not due to the mere interaction of the cells of
the group.
By destroying one of the first two cleavage cells half-blastulas even,
perhaps, half-gastrulas were reared in a few cases from the single
remaining cleavage cell. The half-blastulas were obtained both after
piercing and removing one cell and by killing it by shaking, which
left it still inside the intact membrane.
The half-blastulas showed a tendency to close in as spheres, but died
first.
One interesting case suggesting Roux’s “ postgeneration ” was seen,
but lost. It consisted of a half blastula lying against a second solid
hemisphere covered over by a layer of cells continuous with the half-
blastula.
Electricity and Cleavage.—Wilhelm Roux,’ seeking to deter-
mine if electrical phenomena are involved in the process of karyoki-
netic cell division, subjected frogs’ eggs to the action of a current from
three Bunsen cells. In these experiments, made in 1885‘, the eggs
were placed in a glass tube surrounded by the coiled wire conveying
the current. :
The result was negative. In the present paper the author describes
the results obtained by the use of an alternating current of 100 volts
or less, used for lighting the Anatomical Institute at Innsbruck. Here
again, the result as far as any connection of cleavage or cell division
and the electric current is concerned, was entirelyfnegative. When
the current was not strong enough to kill the eggs they divided in the
glass tube without any reference to the presence and direction of the
current. The same is found to be true of the maximum continuous as
well as of the alternating current.
The alternating current is also found to have no directive effect upon
the entering sperm or the fusing pronuclei, factors which Roux regards
as determining the first cleavage plane. The electric current has,
` however, a marked effect upon the egg, visible as a change in color at
5Bresl. ärzt. Zeitschrift, 1885, No. 6.
‘Sitzb. Akad. Wiss. Wien., Jan., 1892.
1893.] Embryology. 397
each pole as contrasted with the equator, where such a contraction
takes place that at first one might regard the equatorial belt as a
cleavage furrow. These changes involve the death of the egg, and are
merely a “ morphological ” polarization produced by the passing cur-
rent and localized by its direction irrespective of any axial or polar
differentiation within the egg itself.
However interesting and valuable the long series of experiments
recorded in the two hundred pages of this memoir may be to the phy-
. Sicist and to the biologist, they have for the embryologist of the present
day too little direct bearing to make it worth while reviewing them at
length.
It should be mentioned, however, that the most diverse objects, frogs’
eggs, gall bladders, embryos, hearts, hydras, tritons, lizard and fish
embryos, chick and mammal embryos, as well as inorganic substances,
such as mercury, copper, lead, ete., exhibit visible differentiation of
polar and equatorial areas dependent upon the direction of passing
currents. Yet this is not true of even all organic bodies experimented
with.
Membranes of the Sea Urchin Egg.—Curt Herbst’ repeated
the experiment of the Hertwigs, and used in addition to chloroform,
clove oil, creosote, xylol, toluol and benzole.
Eggs of the sea urchin shaken in water that had been mixed with
small quantities of any one of these substances form an artificial egg-
membrane just like that normally formed after the entrance of a
sperm in fertilization.
This membrane, the author holds, is made by the hardening of the
preexistent hyalin outer layer of the egg.
The subsequent separation of the membrane from the surface of the
egg is probably due to the secretion of some jelly-like substance. The
egg does not shrink away from the membrane at all.
The sperm has no direct part in the formation of the membrane,
but merely acts as a stimulus to the egg. If the membrane is
removed from a fertilized egg (by shaking) the presence of more
sperms does not cause the formation of a new membrane. If, however,
some of the above substances are used, a second membrane is formed.
Two membranes, one inside the other, may be formed from eggs having
one membrane, or even from those having the first membrane removed.
The cause of this membrane formation is to be sought for in the egg
itself.
5Biol. Centralblatt, Jan., 1893.
398 The American Naturalist. [April,
Experiments on Cleavage.’—Dr. Jacques Loeb, of the Uni-
versity of Chicago, has madea most valuable addition to our know-
ledge of the cleaving ovum. His experiments were simply the expo-
sure of the eggs of sea urchins (Arbacia) to water containing more or
less than the normal amount of sodium chloride.
The general result is stated as follows: “If we reduce the irritabil-
ity of the protoplasm of the egg by reducing the amount of water
contained in it, the nucleus can segment without segmentation of the
protoplasm. If we increase again later the amount of water, and -
consequently the irritability of such an egg, the protoplasm at once
divides into about as many cleavage cells as there are nuclei pre-formed.
The segmentation of the protoplasm in the egg, and probably in every
cell, is only the effect of a stimulus exercised as a rule by the nuclei.”
- The following illustration of the character of the experiment is
presented by the author: Eggs taken a few minutes after impregna-
tion were divided into four lots, one put into normal sea water, one
into that concentrated by adding 2 g. Na Cl per 100 ccm., the other
two into sea water concentrated by addition of 1.3 g. NaC and 1 g.
Na Cl per 100 cem. in each case.
When the eggs in normal sea water had arrived at the two celled
stage none of the others had as yet begun to cleave. In the least con-
centrated solution the cleavage soon followed and in the more concen-
trated solution it followed about an hour later, but in the most con-
centrated solution no cleavage took place. Concentrations greater than
2 g. per 100 ccm. produced plasmolysis. The form of the cells indi-
cates the amount of water and the intracellular pressure; thus in
normal water the first two cells are nearly hemispheres, but in concen-,
trated solutions the cells approach more and more toward a spherical
shape.
Other experiments bring out the interesting point that the effect of
salt is not to destroy but to suspend the cleavage phenomena. When
the eggs are put back into normal water after staying some time, but
not as long as twelve hours, in concentrated water, the suspended
cleavage begins and goes on much as in a normally situated egg. The
longer the eggs have been in the concentrated water the more numerous
are the cleavage cells formed all at once when the egg is returned to
normal water. An interval of about twenty minutes in the normal
water must elapse before the sudden appearance of the retarded
cleavage cells occurs.
®Journal of Morphology, vii, 1892.
1893.] Embryology. 399
The behavior of the nuclei of the eggs in concentrated sea water
was observed somewhat in live eggs and in certain stained eggs which
Dr. Conklin prepared for the author. It seems that the nuclei increase
in numbers in the salted sea water when there are no cleavage furrows
visible on the outside of the egg, but this increase is not alwàys accom-
panied by a normal separation.
In the light of the conceptions of Fol and O. and R. Hertwig
regarding the effects of polyspermy in producing a cleavage into many
simultaneously formed cells it might have been urged that Dr. Loeb’s
results were due to polyspermy.
Granting, however, that the increase in nuclei takes place while the
eggs are in the salted sea water such facts as the effect of this water in
paralyzing the spermatozoa, as observed by the author, show that the
spermatozoa cannot be connected with these peculiar cleavage phe-
nomena. i
Regarding the method of action of the salt used in these experi-
ments we must premise that the author in previous work upon hydroids
came to the conclusion that growth and regeneration in animals and
plants is a function of the amount of water contained in the cells.
The application of this to the present case is in the idea that increas-
ing the concentration of the liquid about a cell decreases its irritabil-
ity by removing water from it ; the effect is the same quantitatively
and qualitatively as would be produced by lowering the temperature.
- The normal source of the stimulus which the abstraction of water
is supposed tọ render no longer efficient to produce cleavage is con-
sidered to be the nucleus. The nature of this stimulus is unknown,
but some facts lead toward the sumption that it may be a chemical
one
On the other hand the protoplasm has some influence upon the
nucleus ; possibly the intracellular pressure determining the form of
the cell also fixes the direction of the nucleus, which will then be less
defined in a mass without cell walls.
400 The American Naturalist. [ April,
ENTOMOLOGY:
Termitophilous Insects.—E. Wasman has lately described’ a
number of new termitophilous insects, erecting for part of them the
genus Termitobia and has summarized our present knowledge of the
guests of termites. He says that very little is known concerning the
guests of the termites as compared with those of ants, largely because
the former are chiefly confined to the tropical and sub-tropical regions
and also because the investigation of their nests is more difficult. But
in spite of this more than fifty species of termitophilous animals are
known. Of these forty belong tothe Coleoptera, two to Orthoptera,
one to Heteroptera, one to Lepidoptera, two or three to Thysanura,
one to the mites (Acaroidea), one to the Arachnida, and one is a Nem-
atode. Of the beetles one is a species of Glyptus with its larva, and
thirty-nine are Staphylinidæ.. The author refers to the investigations
of Mr. E. A. Schwarz in North America.’
Notes on the Mouth Parts and Thorax of Insects and
Chilopods.—The difference between the mouth parts of true insects
and Chilopods are usually mentioned in text-books. But when one
believes that insects must have come from some form similar to living
Chilopods, it becomes a matter of interest to see what is the homology
between the various parts of the mouth. As it is usually stated, Chilo-
pods have one pair of mandibles, two pairs of maxilla, and a pair of
maxillipeds ; insects a pair of mandibles, one pair of maxillæ, and a
labium. The mandibles are of similar structure in both groups. In
Chilopods the first pair of maxillz are two-lobed; the second pair of
maxillz, or first pair of legs (as it is sometimes called), is, in struc-
ture, like a pair of legs; the maxillipeds are a modified pair of legs,
large and powerful, used for seizing the prey. The dorsal scuta of the
segment to which the maxillipeds belong is frequently not wholly
united to the head, but is plainly visible (Lithobius).
We here see what is well-known among the higher Crustacea; the
tendency to the cephalization of thoracic segments in order that their
appendages may function. as mouth parts. If such is the case, what
‘Edited by Clarence M. Weed, Hanover, N. H.
*Neues Termitophilen, mit einer Uebersicht über den Termitengiste. Verhandl.
Zoologisch botanish Gesellsch. in Wien, xli, 647.
Proc. Ent. Soc. Wash., i, 161.
PLATE X.
The Woolly Alder Aphis.
1893.] Entomology. 401
has become of the second maxille and the maxillipeds? The maxilla
of insects is composed of three portions, the palpus, the galea, and the
lacinia with the basal joints. If now we look at a Perlid larva (the
lowest winged insect) the secret lies revealed; the basal joints of the
maxillipeds have united to form the sub-mentum, the remaining joints
by their pressure have united to the basal joints of the maxilla. Yet
in the Perlid larva the union is not complete and one can plainly see
that the palpiger is not articulated to the stipes, but has basal joints
which are concealed by the stipes and the two parts of the cardo. So
that the maxill of insects is equal to the first maxille of Chilopods
plus the maxillipeds. The labium of insects appears to be the same
as the second maxille of Chilopods, the mentum being the united
basal’ joints. ;
I would also call attention to the steady tendency observable in
Chilopods toward the union of certain thoracic segments. In the lower
Chilopods (Geophilus) the segments are all of about equal size, and
each one bears a spiracle. A little higher, in Scolopocryptops, we see
that there are two kinds of segments, the large with spiracles, the small
without them. Segments 2, 4 and 6 are small, the 7th and 8th both
large, after that every other one is small. In Lithobius the process is
continued still further, the small being smaller, the large larger. In
Scutigera (the highest Chilopod) the dorsal scutæ of the small segments
become united to the larger ones, so that seen from above, the first
scuta (prothorax) covers one pair of legs, the second scuta (mesothorax)
covers two pairs, the third scuta (metathorax) covers two pairs, and
the fourth scuta covering three pairs of legs. Therefore I conceive
that the mesothorax and metathorax of insects are each composed of
two segments. This would appear quite probable if one but look at a
grasshopper; but there exists greater proof. Machilis (a common
Thysanuran) has on each abdominal segment a pair of small append-
ages, which have been recognized as representing legs, but the meso-
and metathorax also bear these appendages beside the normal legs ;
hence if these appendages represent legs the meso- and metathorax of
Machilis must be compound segments. It is possible that each abdom-
inal segment is compound, but I hardly think this probable, since the
first abdominal segment of insects, which is very small, appears to
represent the small sixth segment of Scutigera. If this theory of the
insect thorax be true, the thorax consists of five segments, segments
one, three, and five bearing legs, segments two and four bearing wings.
Narman Banks, Sea Cliff, N. J.
27
402 The American Naturalist. [ April,
The Woolly Alder Aphis.—In the current report of the New
Hampshire Experiment Station observations are recorded by Clarence
M. Weed which show that in autumn large numbers of the young of
this species (Schizoneura tessellata Fitch) are born by the partheno-
genetic females. These young descend the alder shrubs to the surface
of the ground, where they congregate, under various sorts of shelter, in
enormous numbers. They remain there throughout the winter, and in
spring those which have not been washed away or otherwise destroyed
climb up the trunk to the limbs, where they fix themselves on the
lower surface of the bark and grow rapidly into other parthenogenetic
females. No males, females, or eggs have been found. This insect
has been very abundant in New England during recent years; the
appearance of an infested branch is shown in the accompanying plate.
Further Note onthe Tineid Case-worm from the Grand Can
yon—lIn my description of the seed-like lepidopterous case-worm from
the Grand Canyon in the February, 1893, number of Tar NATURALIST,
pp. 166-169, figs. 2 and 3, it should be mentioned that the hair line in
fig. 2 shows twice the natural size of the larva. Fig. 2 was intended to
be reduced one-half, and was so indicated on the margin of the original
drawing. The hair line was made twice the length of the larva, so
that when the whole was reduced one-half it would represent the nat-
ural size, stated in the description as 2¢ mm.
It seems very certain that this larva is a tineid. It is well-known
that these small larvz often construct cases in which they live; and,
as Dr. Packard states (Guide, p. 342), the larve of some of the
smaller tineid genera (Antispila, Tinagma, etc.) are absolutely footless.
The narrow-leafed willow referred to in the description is Salix
longifolia Muhl., while the round-leafed leguminous tree is Cercis occi-
dentalis Torr. The elevation of that part of the canyon referred to as
2500 ft. below the rim is 5000 ft. above sea level.
C. H. TYLER TOWNSEND.
The Puparium of Blepharipeza.—Blepharipeza i is a genus of
Tachinidæ s. str., belonging in the subfamily Hystriciine, and thus
closely allied to Dejeania and the other Tachinidæ that are character-
ized by the presence of dense and spiny macrochaetæ on the scutellum
and abdomen. Blepharipeza may at once be recognized in this group
by its hind tibiæ being pma ciliate on their outer oe) with more
or less flattened brist
Ses
SS SS Se ete eee
1893.] Entomology. 403
The following description is drawn from a puparium of Blepharipeza
adusta Lev., from which emerged a 9? specimen of the fly, both of
which were sent to me by Mr. Harrison G. Dyar, who bred the fly
from a larva of Halisidota argentata. It will serve to indicate the
distinctive features of the puparium in this interesting genus. I do
not know that any description has ever been published of the pupa-
rium in this group, except the very brief one of Belvosia bifasciata by
Dr. Riley (Bull. 3, U. S. Ent. Com., p. 42). The latter indicates the
puparium of Belvosia to be very different from that of Blepharipeea
in the structure and situation of the anal stigmata or spiracles. This
may serve to emphasize the distinctness of the two genera, which some
authors have been prone to combine under the name Belvosia.
Description of Puparium.—Length, 10 mm.; greatest width and
thickness, 5 mm. Color dark brown. Almost cylindrical, perfectly
rounded at both ends, the anterior end a little less in diameter than
the posterior, the transverse and dorso-ventral diameters of each seg-
ment equal. There appear to be 12 segments, including the capital
and anal plates, but the segments are very ill-apparent on the posterior
end, very plain anteriorly. A belt or zone of circular (longitudinal
_ to the belt, transverse to the puparium) minute corrugations alternate
with one having a rugose surface, there being 10 of the corrugated or
fluted belts, and 10 of the rugose or punctured belts. The rugose belts
are about one-half the width of the fluted belts, and there is appar-
ently one of each to each segment of the puparium, not including the
capital and anal plates. The anterior one-third of each segment is
` occupied by the rugose belt, and the posterior two-thirds by the fluted
belt, these belts more or less clearly defining the segments. Mouth
parts of larva showing on capital plate as a wrinkled protuberance
consisting of 4 principal eminences, separated from each other by a
wrinkle or fold of the integument. Each eminence is more or less
minutely wrinkled, the wrinkles as well as the large folds radiating
from the center of the protuberance. Anal stigmata showing an anal
plate at center of posterior extremity of puparium, as a pair of organs,
each marked by 3 short raised ridges, the two upper ones more or less
parallel with each other, inclined outward above and at an angle of
about 45 degrees with the lower one, which is not quite parallel with
the transverse diameter of the puparium. These ridges are usually
more or less slightly curved ; when curved, it is always with the slight
convexity outward from the center of the organ or stigma. The two
stigmata occupy the same transverse plane of the puparium. What
appears like an anal vent is situated exactly ventrally of the pair of
404 The American Naturalist. [April,
anal stigmata, forming with them the corners of an equilateral trian-
gle, the vent representing the lower or exactly inferiorly directed
angle. This vent consists of a circular raised portion of the integu-
ment, rugose in appearance, of about the same diameter as either
stigma, and with a little circular pit-like depression on top. Exactly
ventrad of this vent is a smaller, more irregularly shaped tubercle,
apparently situated on the tenth segment. It is hardly more than
three times as much removed from the anal vent as that is from either
stigma, measuring from center to center.
_ The puparium is furnished inside with a fine, white, silken, membra-
nous lining —C. H. TyLer Townsenp, Las Cruces, New Mexico.
a
1893.) Microscopy. 405
MICROSCOPY.
The Solution of the Dust Problem in Microscopy.—The
statement of the dust problem is this: Given a stock of cleaned micro-
slips and cover-glasses, to keep them clean and ready for use at any
moment, without the necessity of brushing or wiping them. The fol-
lowing solution is the result of a laboratory study of the problem, and
is now announced after having received the approval of leading micros-
copists,
The objects to be accomplished are :—
1st, To secure a dust-proof magazine for storing the cleaned micro-
slips and cover-glasses in separate compartments for the different sizes
of squares, circles, and oblon
2d, To provide simple mechanical appliances for removing a single
slip or cover without exposing those remaining in the case
3d, To provide an automatic device for warning the operator of the
approaching exhaustion of his stock of any of the shapes of covers and
of slips, thus avoiding the necessity of opening the case except to replen-
ish stock.
4th, To provide against the breaki li g t of the covers
iu case of the nocideutal ‘overturning ye the case
5th, Incidentally to provide a mounting table with guides for center-
ing the objects and cover-glasses, the whole apparatus being so con-
structed that the glasses are not touched by the fingers, and only once
by the forceps until the slide is completed and labeled.
The Dust-proof Slip and Cover-glass Case” fully meets these require-
ments.
Plate VI is a front view of the case which is 4} inches square on
the base and 6 inches high. The slips and mounting table are con-
tained in the lower half and the cover-glasses in the upper portion. The
mounting table is shown in position for work, and the concentric lines
for guidance in properly centering the objects and covers are seen
on the celluloid slip. Four dust-excluders are hinged to the front
of the grooved table, and two of them are shown opened so as to expose
the slots through which the cover-glasses pass out upon the grooved
table. The milled head of the roller which pushes the cover-glasses out
of the slots is shown at the side of the case. A screw stop is inserted in
the front of the case above each slqt.
1Ed. by C. O. Whitman, University of Chicago.
2Patent applied for.
406 The American Naturalist. [April,
The case is fitted for 3 inch by 1 inch glass slips of any thickness,
and has a capacity of six to twelve dozen. It has four compartments
for the cover-glasses of the sizes 4 inch, $ inch, ł inch, $ inch, or 12mm,
to 24mm., either circles or squares or oblongs of the above widths, and
any lengths up to 2} inches. The screw stops regulate the depth of
the slots for either No. 1, 2 or 3 thickness of cover-glasses.
Plate VII is a vertical section from front to back, showing the plan
of the case. The section of the lower half is made in the median plane,
that of the upper half in the plane of a screw stop. Through the
base (b ) projects the stop (s ) into a narrow groove in the mounting
table ( mt ) which slides in and out between the base and the floor (f)
of the stock compartments (sc’). The service compartment (sc) is
deepened by a shallow cavity cut into the upper surface of the mount-
ing table, and in the bottom of this cavity is fitted a sheet of celluloid
bearing a series of concentric lines. The slips ( ss ) are stacked in the
service compartment, the lowest slip filling the shallow cavity in the
mounting table as shown at (ss’). A warning slip of paper ( ws ) is
folded around the front edge of the top slip. The loose partitions ( p )
running in grooves in the sides of the case can be removed after slid-
ing back the cover (c). In the base (f’) of the cover-glass case is the
groove (1 ) for the roller (r ) which is made of brass and covered with
chamois. To the front of the base is hinged the brass dust-excluder
(de) whose lining of silk plush protects the grooved table (t) and the
slot (sl) from dust. The dust-excluder also prevents the delivery of
a cover glass by the roller except when the slot is opened. Into the
slot projects the point of the serew-stop (st ) which is so adjusted as to
permit the passage into the slot and out upon the grooved table of only
one cover-glass at a time. Above the roller are stacked the cover-
glasses (cg). Near the top of the stack is placed the warning disk
(w) and upon the top cover-glass rests the loosely-fitting follower
(cf) which prevents displacement and breaking of the covers, if the
case is accidentally overturned. The adjustable partition ( ap) may
be moved backward or forward to fit any length of cover-glass from
} inch to 2? inches. The cover (c) is so constructed as to exclude
ust.
Plate VIII is a front view of a form of the case as made for one size
cover-glass, either circles or squares, and of either ŝin., łin., or žin.
diameter. The open dust excluder is hinged above the slot at the sides
of the case. The engraving does not show the groove in the table in
front of the slot. A slip is shown in position in the mounting table.
One of the lateral grooves in the mounting table is not represented.
1893. ] Microscopy. 407
The plan in general, is the same as in the complete form illustrated in
Plates VI and VII.
Method of operation.
Thoroughly dust the inside of the case and partly, or completely,
fill the compartments with clean slips and cover-glasses of the proper
sizes and place the warnings in position. The apparatus is then ready
for immediate use, or at any time during successive weeks or months until
the stock of slips and covers is exhausted.
The slips are withdrawn from the case singly by pulling forward
the mounting table, and from this they are removed, either before or
after an object is mounted, by inserting a finger into the notches cut
into the sides of the table. The mounting table is automatically
reloaded on being pushed back into the case. The warning slip having
been withdrawn, the service compartment is again filled by removing
the sliding cover which carries with it the cover-glass case, withdraw-
ing a loose partition and sliding the cleaned slips from a stock com-
partment into the service compartment.
Whenever a cover-glass is needed, the dust-excluder in front of the
proper slot is opened by a touch of the finger, the milled wheel is
- rotated and the chamois covered roller pushes the bottom cover-glass
through the slot and out upon the grooved table, where it is readily
grasped by the forceps. This action is positive because the friction
between chamois and glass is greater than between two clean glass
surfaces. The table being grooved, only the extreme margins of the
cover-glass touch it. A touch of the finger closes the dust-excluder.
It will be noted (a) that the slips and covers are doubly protected
from atmospheric impurities by the dust-proof case and the constant
contact of clean glass surfaces with each other; and (b) that in the
process of removal for use they do not come in contact with any sur-
face from which they receive dust. They come out as clean as when,
weeks or months before, they were put into the case. Thus the dust
problem is solved—A. H. Cons, A. M., University of Chicago.
Cooling Paraffine.—To those who have experienced the incon-
venience and difficulty connected with the preparation of good par-
affine sections in the Tropics or even in the Tropical Summers of other
climates, the following mee device of Professor C. Gracomini' may
be a welcome suggestion
1Arch. Ital. de Biolegie, IX, 1888, p. 380.
408 The American Naturalist. [April,
A small rubber tube, a meter or so long is connected with a vessel of
ice-water partly stopped at the other end so that a slow stream passes
through it. This tube is wrapped around the paraffine block upon the
microtome and cools it off so that thin sections can be cut.
If in winter the paraffine is too hard, the same device with warm
water may be resorted to.
PLATE VI.
PLATE VII.
P
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PLATE VIIL
1893.] : Proceedings of Scientific Societies. 409
PROCEEDINGS OF SCIENTIFIC SOCIETIES.
Boston Society of Natural History, February 1.—The fol-
lowing papers were read: Mr. H. L. Harris, A new instance of the
capture of streams; Professor W. T. Sedgwick, The natural history of
disease.
February 15.—The following papers were read: Professor Henry
W. Haynes, More evidence of cannibalism among the Indians of New
England; Dr. R. T. Jackson, Notes on the development of palms;
Dr. 8. J. Mixter, A Massachusetts beaver dam.
March 1.—The following papers were read: Prof. E. S. Morse, A
curious Aino toy; Dr. C. Willard Hayes and Mr. M. R. Campbell,
The structural features (geomorphology) of the southern Appalach-
ians. Maps and diagrams were shown and a discussion followed, Pro-
fessor Davis, Mr. Upham and others participating.
SAMUEL HENSHAW, Secretary.
New York Academy of Sciences, Meeting of Biological
Section, Feb. 13, 1893. A paper on the Functions of the Internal
Ear was presented by Dr. F. S. Lee, based upon a study of dog-fish.
The results of ease: were given showing that the semicircular
canals are sensory organs for dynamical (rotational) equilibrium,
otolithic rat for statical (resting) equilibrium. Each canal
appreciates movement in its own plane and by a definite functional
combination of canals all possible rotational movements are radiated.
This theory explains compensating movements of eyes, fins and trunk.
The method of experiment was that of sectioning the branches of the
acoustic nerve and stimulation (by rotational movements) of the
living swimming fish.
In a paper by Bashford Dean on the Marine Laboratories of Europe
a series of views was shown of the stations of Naples, Banyuls,
Roscoff, Plymouth, Arcachon, the Helder, and St. Andrews
H. F. Osborn described the foot of Artionyx, the new member of the
order Ancylopoda Cope. It is distinguished from Chalicotherium by
the artiodactyle character of ankle and pes.
Basurorp Dray, Ree. Sec.
410 The American Naturalist. . [April,
The Biological Society of Washington.—The following
- communications were made: Mr. M. B. Waite, the Destruction of
Lichens on Pear Trees; Mr. C. H. Townsend, The Propagation of the
Atlantic Coast Oyster on the Pacific Coast; Mr. Charles Hallock, The
' Geographical Distribution of the Musk-Ox; Dr. C. Hart Merriam,
The Four-toed Kangaroo Rats (with exhibition of specimens).
February 25.—Communications: Mr. Sheldon Jackson, The Intro-
duction of Reindeer in Alaska; Mr. M. B. Waite, Variation in the
Fruit of the Pear due to Difference of Pollen ; Mr. E. M. Hasbrouck,
On the Development of the Appendages of the Cedar Waxwing; Mr.
F. A. Lucas, The Food of Humming-birds.
FREDERIC A. Lucas, Secretary.
Anthropological Society of Washington.—Special Meeting,
February 14.—The program consisted of a lecture, A Primitive Priest-
hood, by an Initiate. Illustrated by lantern slides by Frank Hamil-
ton Cushing —Weston FLINT, Secretary.
SCIENTIFIC NEWS.
Errata in Mrs. Bodington’s article “ Legends of the Sumiro-Acca-
dians of Chaldea” in THE American Naturauist for February,
1893, p.105.
Page 105, Line 2 from bottom, for Bohu read Bahu. (N. B.
Bohu in the last line is correct.)
Page 106, Line 4; for Herodach read Merodach.
Page 107, Lines 13, 18, 21, 25, 30, and 36; for Tzdubar, read Izdu-
bar.
Page 107, Line 14; for Tshtar read Ishtar.
Page 108, Line 1; for Tzdubar read Izdubar.
Page 108, Lines 12 and 31; for Tshtar read Ishtar.
Page 209, Lines 5, 8, 12, 18, 24, 31 and 37; for Tshtar read Ishtar.
Page 110, Lines 4, 7 and 15; for Tshar read Ishtar.
In the January Number, 1893.
Page 14, Note 1; for Story of the Natives read Story of the
Nations.
Dr. Henry Wheatland, of Salem, Mass., died February 27, 1893
aged 81 years. Dr. Wheatland graduated at Harvard in 1834, and
1893.] Scientific News. | 411
later at Harvard Medical College. While a student at college he
received instruction in zoological studies from Nuttall, the ornitholo-
gist and botanist. Later he turned to anatomical work and then. to
the study of marine zoology. Under his instruction the late William
Stimpson did his first dredging in Massachusetts Bay. For many
years Dr. Wheatland has been President of the Essex Institute at
Salem, Mass., and he has been trustee of both the Peabody Academy
of Science at Salem, and the Peabody Museum of Archeology and
Ethnology at Cambridge since their foundation. In later years his
energies have been devoted to the study of local history. Personally,
Dr. Wheatland was a delightful companion, and his stories of the
older days of American Science were especially interesting.
Mr. A. Smith Woodward, of the British Museum, has gone to the
Lebanon to explore the deposit of fossil fishes.
Professor R. Ellsworth Call is now instructor in Physics and Chem-
istry in the Manual Training High School, Louisville, Ky.
The Tacoma Academy of Sciences is endeavoring to have the name
Mt. Tacoma adopted as the official title of what was formerly known
as Mt. Ranier. If the Academy could have the name of the State of
Washington changed to Tacoma it would be a great convenience. We
have too many Washingtons already. x
The Leuckart “ Festschrift,” the volume of the contributions of the
former students of the celebrated Leipzig zoologist, brought together
at his seventieth birthday, makes a splendid quarto volume illustrated
by forty plates. Thirty-five pupils contributed to the memorial.
Richard Greeff, Professor of Zoology in Marburg, died August 30,
in his sixty-fifth year.
J. G. Joessel, Professor of Anatomy in the University of Strasburg,
died Dec. 4, 1892, in his 55th year.
Professor W. Pfitzner, of Strassburg, has been appointed to the
position left vacant by the death of Professor Joessel. His assistants
will be Drs. Mechnert and Heinrich Hoyer (of Würzburg).
Professor F. C. Noll, for many years editor of the “ Zoologiscenh
Garten,” died in Frankfurt a. M. Jan. 14, 1892.
Dr. Jan Pieter van Wickevoort Crommelin, an ornithologist, died
in Haarlem, Holland, Oct. 20, 1892.
412 The American Naturalist. [April,
Thomas John Moore, Curator of the Liverpool Museum, died Oct.
31, 1892
Dr. Benjamin Vetter, Professor of Zoology in the Royal Polytech-
nicum of Dresden, and well-known as the translator of many works,
died in the night between Jan. 1 and 2, 1893.
Professor H. Schaaffhausen, the anthropologist of Bonn, is dead in
his 77th year.
Professor James Ellis Humphrey sailed Feb. 11th on a botanical
collecting expedition to Jamaica, where he wishes especially to study
the algae of the Caribbean Sea.
Mr. Samuel Henshaw is in charge of the entomological collection of
the Museum of Comparative Zoology at Harvard.
The University of Minnesota is publishing a“ Quarterly Bulletin.”
From the last issue (Jan. 30, 93) we gain the following: The. herba-
rium of the University and of the State survey are merged together
In the past three years 350 species of Minnesota fungi have been
determined. In 1892 over 20,000 species of flowering plants were
collected in the limits of the State, and the collections now contain
about 900 species from the region. In the line of geology considerable
work has been done in collecting lower Silurian fossils near the Uni-
versity; while the collections have received many additions from other
States. In zoology considerable work is being done. The Museum ‘
has been fitted with cases, but the curator reports that the collections :
still leave much to be desired. Not half the birds of the State are a
represented, while the collections of fishes, reptiles, amphibia, etc., are
largely in prospect. The labors of the zoological survey during the
past summer were mostly in the neighborhood of Mille Lac, which
was pretty thoroughly explored, and work is being carried on, so that
a few years hence there will be one of the best local faunal collections
for reference and comparison in the country. The University also
proposes to establish a Biological station on the upper Mississippi to be
open without fees to all who may wish to come. The location will be
e decided later. A laboratory for histology and embryology in the
College of Medicine has been fitted up with equipment for elementary
and advanced work.
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Vol. XXVII. MAY, 1893.
CONTENTS.
PAGE,
RECENT STUDIES OF CARNIVOROUS PLANTS. asia—Polar Regions. .
Jared G. Smith. 413 Geology and Pai. The Western Te
land of Ecuador—Devonian Fossils from Manitoba
—Jura and Trias at Taylorville, reecaye ates
A NEw THEORY OF THE MECHANICAL EVOLUTION
OF THE METAPODIAL KEELS oF DIPLARTHRA,
š (Mustrated:) . a o, a J Li Wortman.
a CLIrF-DWELLERS. (IUustrated.)
: Clement L. Webster.
Marine Pliocene Beds of the Carolinas—Geological
News—General—Paleozoic — Mesozoic —Cenozoic, | 4
New DISCOVERIES OF Fossit MAMMALIA OF Dan T yoan rk Ea Pe .
: -SOUTHERN PATAGONIA. Florentino Ameghino.
EDITORIALS.— The Geological Survey of Georgia—
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AMERICAN NATURALIST
Vou. XXVII.
May, 1893. 317
RECENT STUDIES OF CARNIVOROUS PLANTS:
By JARED G. SMITH.
N. Tischutkin published an article in 1889 in the Berichte der
Deutschen Botanischen Gesellschaft on the cause of the digestion
of albumen by the leaves of Pinguicula vulgaris L. in which he
endeavors to show that the process of digestion is the result of
the action of Bacteria. This is in opposition to the theory
of Darwin and other authors that the digestion is analogous to
the digestion by means of pepsin in the animal kingdom.
In an article in volume XII of the Acta Horti Petropolitani
he further discusses the subject and concludes that the pepsin
of the leaves of insectivorous plants is not a secretion of the
plants themselves but is a by-product formed by the numerous
species of Bacteria found in the digestive fluid.
In his former study Tischutkin performed over one
hundred and fifty experiments, using extracts from the leaves
of Pinguicula and small amounts of the secretions of the insect
catching leaves. Into these extracts and secretions he put
small cubes of coagulated egg albumen. In no case did there
follow any peptonization of the egg albumen. These results
impelled him to take up the further study of thé relation of
micro-organisms to the digestion of albuminous compounds by
insectivorous plants, using other species and genera.
1$haw School of Botany, St. Louis. Read before the Botanical Seminar of the
University i Nebraska, Feb. 18, 1893.
414 The American Naturalist. [May,
Certain authors who have investigated the digestive process
(among them Darwin, Gorup, Besanz, Reess and Will) have
concluded that it is the result of pepsin secreted by the plants.
Others, as C. Morren, see only a process of rotting and decay.
Tischutkin in his former paper has reviewed the methods by
which these various authors have arrived at their results.
Morren placed on the leaves of Drosera binata Labill. different
insects and cubes of coagulated egg albumen and observed
that a few hours afterward the albumen became transparent,
the angles were rounded off, and after a day or two the pieces
entirely disappeared. Microscopic examination of the trans-
parent fragments of albumen revealed the presence of bacteria,
monads, and filaments of a mycelium which resembled the
conjugating stage of some species of Mucorineæ. He concludes
that while this dissolving of the albumen by some absorbent
principle is possible, the intervention of pepsin is entirely
problematic.
Schimper in the Botanische Zeitung for 1882 tells of some
experiments which he made with Sarracenia purpurea. These
are of great value because made on plants growing wild in
Massachusetts. A part of the observations were made on
insects caught by the plants, and a part by placing pieces of
meat on the leaves. He found that on stopping up the
mouths of the leaves at their time of opening and thus pre-
venting the free entrance of bacteria the digestion took place
no more rapidly than when the bits of meat were placed in
pure water, and from this fact he concluded that pepsin was
not present. He did not test the liquid by chemical means.
He found great numbers of worms in many of the leaves and
thinks that they may have something to do in rendering
organic matter absorbable.
Hildebrandt in the Botanische Zeitung for 1870 tells of the
experiments made on the leaves of Dionxa muscipula Ell. an
on the cups of Sarracenia, Cephalotus and Nepenthes. He
doubts very much whether the insects caught are of value to
the plants. “My observations on Dionza muscipula showed
me that the long legged spiders which are caught in the
leaves exert an injurious influence upon them. A slimy fluid
1393.] Recent Studies of Carnivorous Plants. 415
would be secreted, but after a short time the leaves would
commence to die, if I had not removed the spider in time. ”
Aschman in 1875-6 in his work on insectivorous plants
throws doubt on the presence of pepsin in the secretions of
Nepenthes and Sarracenia, and believes that the digestive
process is simply putrefaction.
Thus we see that many authors have expressed the opinion
that the digestion of nitrogenous substances by the secretions
of the leaves of insectivorous plants is due to disintegration
through the action of the lower organisms rather than by
active ferments produced by the plant itself.
Following this preliminary statement of the published
data of the subject, Tischutkin takes up his own experiments
and gives the results which he has obtained.
The objects which he kept in view while making these
experiments were, first, to see whether the presence of lower
organisms in the secretions was a constant occurrence ; second,
if uniformly present, could these organisms dissolve
albumen; and, third, to compare the results obtained with
other insectivorous plants with Pinguicula vulgaris. The
plants used were Pinguicula vulgaris, Drosera longifolia, D.
rotundifolia, Dionxa muscipula and Nepenthes mastersii. All of
these plants, the last excepted, were cultivated under bell-jars
to prevent the visits of insects. To procure the secretion he
excited the glands by means of cubes of sterilized egg albumen.
The leaves of the first three species responded once to the irri-
tation, those of Dionga muscipula, three times. The acid sap
thus obtained was examined microscopically every 24 hours
for five days. In every case great multitudes of bacteria were
discovered, and frequently there were various moulds as well.
The author considers this discovery of much importance,
because Darwin had stated that the secretions possess antisep-
tic properties which prevent the rapid appearance of micro-
organisms, and compared their action with that of the gastric
juice in the higher animals, which, as is well known, destroys
injurious and decay-producing lower organisms.
Gelatine cultures were made in the usual manner, using a
drop of the secretion from Dionza and Drosera 24 hours
416 The American Naturalist. [May,
after the commencement of the experiment. After the devel-
opment of from nine to ten generations on Fleisch-Peptone-
Gelatine, the different species of bacteria were transferred to
test tubes which contained dilute acid peptone-free meat
bouillon. Control experiments were made with test tubes
containing sterilized water acidulated with dilute hydrochlo-
ric acid. Into each test tube was placed a small cube of
sterilized egg albumen. In all of those test tubes in which
the bacteria developed there followed a rapid solution of the
egg albumen commencing at the surface of the cubes and
invariably continuing to their centers until solution or pep-
tonization was complete.
By these experiments the author proved that bacteria
which possess the power of dissolving egg albumen are —
always present in the secretions of insectivorous plants.
The number of species that had this power were: 7
In Pinguicula vulgaris, 4 species.
In Drosera longifolia, 2 species. ; l
In Drosera rotundifolia, 1 species. 7
In Dionæa muscipula, 2 species.
In Nepenthes mastersii, 2 species.
His next experiments were made using the leaves of Nepen-
thes, where comparatively large quantities of the fluid could
be procured without special irritation. He examined fully -
developed pitchers of Nepenthes mastersii and N. kennedyana.
The reaction of the secretion was neutral. The secretion con-
tained neither peptone nor insects, and yet there were vast
numbers of micro-organisms. The liquid from a leaf of each 4
species was poured into two test tubes, one acidulated with
dilute hydrochloric acid, the other without addition of acid.
A small cube of coagulated egg albumen was placed in each
test tube, and they were then placed in a thermostat at a tempert-
ature of 37.5° C. At the end of sixteen hours the egg albumen
in the tubes to which hydrochloric acid had been added, had
commenced to dissolve, but in the other two was unchang
This experiment was made to satisfy himself concerning the
truth of observations made by Gorup, Besanz and Will that _
the secretion of unirritated pitchers possessed the property
1893.] Recent Studies of Carnivorous Plants. 417
of peptonizing egg albumen only after it had passed from
neutral to acid reaction. He further says that it is extremely
improbable that this peptonizing principle is secreted by the
glands of the Nepenthes pitchers, but that it is produced by
certain micro-organisms present in the solution. Hence the
digestion that takes place is not analogous to digestion of
albuminoids in the stomach of man, because in the latter case
the secretion of gastric juice follows as a result of direct
irritation.
To demonstrate this more fully he repeated his experiments
in a modified form, using a neutral secretion which contained
no bacteria. For this purpose he selected two unopened cups
of Nepenthes coccinea and Nepenthes—sp. ?——which were so far
developed that the liquid was already present in considerable
quantity. The leaves were separated from the plant and the
petioles were attached to a ring stand so that the pitcher of
the leaf was vertical. The outside of each pitcher was disin-
fected with a freshly prepared 1:1000 sublimate solution in
water. After the water of the sublimate solution had par-
tially evaporated, a small hole 3 mm. in diameter was cut in
the wall of the pitcher with a pair of sterilized scissors. The
secretion from each was then transferred by means of a steril-
ized pipette into two test tubes each containing two cubic
centimeters of distilled water and a small cube of coagulated
egg albumen. One of the test tubes was slightly acidulated
with hydrochloric acid, the other remained neutral. The four
test tubes were then placed in a thermostat at 37.5° C.
At the end of 48 hours there wasno solution of the egg
albumen, though a control experiment with a test tube con-
taining pepsin gave positive results. Microscopic examination
and gelatine cultures gave negative results. These experi-
ments were repeated, using the remainder of the secretion
diluted with a little pure glycerin, but the result was the
same.
In these experiments the only question that could be raised
was whether the pitchers from which the secretion was taken
were not too young to contain pepsin. But if one remembers
the experiments of Wunschmann in 1872, and also considers
418 The American Naturalist. [May,
that the cups were more than half filled, there can be little
doubt on this ground. :
To obviate this objection the experiment was repeated in
another way. Two cups were taken, the one from Nepenthes
distillatoria and the other from N. hirsuta, which were still
closed but appeared to be just ready to open. Considerable
secretion was present. The cups were not separated from the
parent plants. The wall of the cup was sterilized with a sub-
limate solution which was washed away with sterilized water.
Then a small V shaped opening was cut in the wall above the
surface of the liquid. Through this opening in the wall of the
cup a piece of sterilized egg albumen one centimeter long and ©
one millimeter thick was introduced into the cup of N. hirsuta
and a piece } cm. long and 1 mm. thick, into that of N. distil-
latoria. The openings in the walls were then covered with
pieces of court plaster, and these for protection from the
moisture of the culture house were covered with varnish.
After four days the cups opened. The pieces of egg albumen
were unchanged, not even the corners being rounded off.
Pepsin was not present in the secretion and bacteria only in
very small numbers.
The secretion from each cup was then poured out into ster-
ilized test tubes containing small pieces of white of egg, and
they were set aside at a temperature of 20-22° C. The white
of egg in the secretion from N. distillatoria was dissolved in
four days, that in the secretion of N. hirsuta in five. At the
same time enormous quantities of bacteria were developed.
After finishing these experiments the author noticed an
article in the Comptes Rendus for 1890, by Dubois, who had
experimented with the secretions of various Nepenthes spe-
cies taken before the cups had opened. Dubois’s results agree
with those of Tischutkin. The secretion taken from the cups
just as they were ready to open did not contain pepsin and
did not affect the cubes of albumen. But after having been
exposed to the air for some time, putrefaction commenced and
the resultant liquid contained traces of pepsin. Dubois draws
the following conclusions from his experiments. :
1893.) Recent Studies of Carnivorous Plants. 419
Ist. Thatthe secretion of Nepenthes does not contain any
pepsin and that Nepenthes is not an insectivorous plant.
2d. That the phenomenon of disintegration, called “ diges-
tion” by Hooker, was without doubt due to the presence of
micro-organisms and not to the secretions of the plant itself.
The author finds that the results of his various experiments
show that all the disintegration of nitrogenous substances by
the leaves of insectivorous plants is directly dependent upon the
presence of certain lower organisms, principally bacteria, in
the secretions of the plants.
The secretion contains bacteria of many different species,
micrococei and rod-shapes, and sometimes the mycelia of
moulds or other fungi. It is evident that most of these orga-
nisms are carried into the cups by the air, though some may
be washed in by rain drops or carried on the bodies of insects.
The solution of albuminous compounds commences simulta-
neously with the development of great numbers of micro-
organisms in the secretion.
The secretion of the leaves of insectivorous plants is not
analogous to the gastric juice of the higher animals. It is
only a medium in which bacteria may live and develop
while they are breaking up nitrogenous organic insoluble
compounds and preparing them for assimilation by the plant.
It will not be uninteresting to present a table showing the
chemical composition of the secretion of various Nepenthes
species. The calcium and magnesium are in composition
with malic and citric acids.
Malic acid and a trace of citric acid 38.61%
Potassium chloride (Chlorkalium) 50.42
Sodium carbonate 6.36
Calcium 2.59
Magnesium 2.59
Organic matter—a trace
100.57.
There are no free acids in the liquid, though Dr. Turner in
1828 reported a trace of oxalic acid.
420 The American Naturalist. [May,
Without doubt these genera constitute a group entirely
unique. They furnish great natural bacteriological laborato-
ries in which the cultivation of microbes which are beneficial
to the plant is carried on. It isin a certain sense a symbiosis
between the lowest and the most highly organized plants.
The higher organisms furnish a food supply for the use of the
microbes in order that they may in turn be benefited by food
stuffs which the lower organisms manufacture and furnish in
convenient form. That the nitrogenous compounds set free
by the dissolution of the egg albumen are really absorbed is
shown by Darwin in his experiments.
On the ground of his experiments the author offers his con-
clusions as follows:
Ist. The disintegration of albuminous compounds by the
secretions of carnivorous plants is due to the growth of micro-
organisms, principally bacteria.
2. Micro-organisms possessing the power of dissolving
albuminous compounds always vegetate in the secretions of
completely developed carnivorous plants.
3. The disintegration of the albumen does not commence
at the moment of the secretion of the fluid, but only after
micro-organisms have developed in sufficient numbers in the
secretion.
4. The micro-organisms found on the leaves of carnivorous
plants come principally from the air, though they may be
derived from other sources.
The name “ carnivorous” plants is to be understood in
the sense that the plants only assimilate the products which
the lower organisms have set free.
6. The rôle of the plant itself is only to furnish a medium
in which certain micro-organisms may live and develop.
1893.] Evolution of Metapodial Keels of Diplarthra. 421
A NEW THEORY OF THE MECHANICAL EVOLUTION
OF THE METAPODIAL KEELS OF DIPLARTHRA.
By J. L. Wortman, M. D.
In the last number of the Journal of Morphology, Mr. Aus-
tin Carey gives the results of his studies in the foot structure
of the Ungulates. A fair statement of his main conclusions,
or rather perhaps the pith and substance of his argument is to
be found in the concluding sentence of his article which reads
as follows: “ That the lines of evolution have progressed with
but few useless side variations seems to be the uniform testi-
mony of paleontologists; but that race changes follow those
produced in individual life, or that they are directly caused
by their mechanical surroundings, I do not think has been
satisfactorily shown.”
It is not my intention to enter here into a criticism or gen-
eral discussion of the points this author has raised in his
argument, nor will I undertake to discuss at this time the
broad question of the transmission of acquired characters. I
do, however, wish to say a few words upon the question of the
production of crests and keels upon the distal extremities of
the metapodial bones. Upon this point Mr. Carey says, (p.
341) “The crests and grooves on the lower metacarpal ends
in some forms, produced apparently in relation to the sesamoid
bones, is one of the most marked examples of probable
mechanical evolution.” “But before such structures can be
said to prove the inheritance of acquired characters, the ques-
tion should be tested whether they are not produced some-
where in the history of each individual by the necessary
interaction of parts.” He further adds in a footnote that
“The crests in certain highly specialized forms, like the horse
and deer, reach round to the anterior face of the bone and
apparently cannot thus be interpreted or assigned to any
mechanical origin that is obvious.”
422 The American Naturalist. [Máy
With reference to Mr. Carey’s first proposition that these
metapodial crests are produced during the life of each individ-
ual by the necessary interaction of parts, it appears to me to
be a very simple one indeed. If they are produced, by pres-
sure during the lifetime of each individual, and are not
inherited, then clearly we should find the crests absent in new
born animals that had never walked, and in which the meta-
podials had not been subjected to any impact or pressure
whatever. I have taken the trouble to examine a number of
such examples in which the distal ends of the bones were entirely
cartilaginous, and I find that the keels and grooves are as
well developed as they are in the adult animal. I will cite
one case in particular in which I happen to know the history
completely. During the past winter, a young hippotamus was
born in the Zoological Gardens in Central Park, and it was
stated to have been a premature birth; the animal lived but-
twenty-four hours, and I was informed by the keeper that it
never stood upon its feet. An examination of the feet shows
that the distal ends of the metapodials are entirely cartilagi-
nous, and in them the keels are as well prefigured in cartilage
as they are performed in bone in the adult animal. Ihave
also found the same to be true of newborn rabbits and guinea-
pigs. In another case of a young buffalo calf preserved in the
American Museum Collection, the distal keels of the metapod-
ials are complete notwithstanding the fact that the epiphyses
of all the bones are very imperfectly ossified. This evidence,
it appears to me, effectually disposes of the question of the
production of these structures during the lifetime of the indi-
vidual. They are as truly inherited as is the number of digits
or any other important organ in the animal economy.
Mr. Carey further states in his concluding remarks that
“ Plasticity of bone, using the word plasticity not in a physical
sense merely, but to include absorption under pressure, will
‘probably account for much structure in the foot and elsewhere,
especially in connection with the joints and in the field of
variation and correlation.” Now just what Mr. Carey means
by “absorption under pressure” is not clear to my mind. If
he means a process similar to that of the absorption of the roots _
1893.] Evolution of Metapodial Keels of Diplarthra. 423
of the temporary or milk teeth, then the term “ under pres-
sure ” is especially inappropriate, since Tomes has shown’ that
this process is entirely independent of pressure of any kind
whatever, but is in some way connected with the presence of
“myeloid” or “giant” cells. The only other process of
“ absorption under pressure ” with which I am acquainted, is a
pathological one. In this case the pressure is sufficiently
severe to cut off entirely, or seriously affect the supply of
nutrition, thereby causing the death of the part and its subse-
quent removal by sloughing. This however may be very
gradual, so gradual in fact, as to resemble a perfectly normal
change, as is often seen in the pressure of an aneurismal sac
upon a neigboring bone. It may be true that a process of this
nature, having its origin in a strictly pathological cause, may
have become normal. Indeed, Huxley has remarked’ that it
is not always easy to draw the line between pathological and
normal changes. Upon the whole, however, it seems to me
that before we can admit this principle of “absorption under
pressure ” as having played any important part in the modifi-
cation of the skeleton, we must require further proof of its
existence and its method of operation.
That living bone tissue is plastic, I should say highly plastic
and that too in the ordinary physical sense, is so abundantly
demonstrated upon every hand, that no further argument is
necessary to establish the truth of this proposition. It is rec-
ognized and acted upon in everyday surgical practice, and the
number of cases wherein entirely new joints have been formed
in old dislocations, leaves nothing to be desired to complete
the proof. The changes thus produced however, have all
occurred within the lifetime of the individual and it yet
remains to be proven that they are capable of transmission to
succeeding generations. _
With reference to the production of the-metapodial keels or
crests, Cope has explained it in the didactyle foot at follows’ :
“ A similar cause produces a similar result in the development
1Dental Anatomy, p. 197.
2Address to the British Medical Association, London Lancet.
3The Artiodactyla, American Naturalist, March 1889, p. 115.
424 The American Naturalist. [May,
of the tongue and groove articulation between the metapodials
and first phalanges. In alighting on a didactyle foot, the toes
are naturally spread, the result being to throw both the first
phalanges away from the median line, or axis of impact, in
divergent directions. The result of this impact is to produce
upon each metapodial condyle as in the case of the humerus,
an external roller of smaller diameter than the rest of the
condyle, and separated from it by an abrupt crest. In both
humerus and metapodial bones these crests are accentuated by
a pinching process during flexion and extension. This is
produced by the longitudinal torsion which results in all
limbs in motion from the arrest of the outward rotation of the
foot by the earth on alighting. The pinching of a keel by
its groove takes place at all points in the length of the former,
reached by the opposite sides of the extremities of the latter
during flexion and extension. This keel becomes acute and
prominent in the Bodidea and extends to the anterior face of
the condyle. This development has been apparently
especially due to the presence of two sesamoid bones, embed-
ded in the flexor tendons, one on each side of the middle line
of the posterior side of the metapodial condyle. The fissure
between these two bones has permitted the moulding of the |
surface into a keel to fit it. That this has been the case is
further indicated by the fact that a median trochlear surface
exists at the distal extremity of the first phalanx in all mam-
mals. But asingle flexor tendon crosses this articulation, and
it contains but one sesamoid bone, to which the trochlear sur-
face is moulded in a concave surface, as is the case of the
patella and the rotular groove of the femur. ”
In support of Cope’s proposition, that the development of
crests or keels upon the plantar or palmar aspect of the distal
end of the metapodials, is due primarily to the presence of
two sesamoids imbedded in the flexor tendons, I can add
that in every case where these sesamoids are present, and
pressure is exerted by the flexure of the phalanges upon the
metapodials, the keels are developed, and conversely, that
where the sesamoids are absent or are present without pres-
sure, the keels are absent. The human foot and hand are
1893,] Evolution of Metapodial Keels of Diplarthra. 425
excellent examples of this. In the foot there are two sesa-
moids developed in the tendons of the flexor brevis hallucis as
they pass over the end of the metatarsal of the great toe, to be
attached to the phalanx. Now in the act of walking the
greater part of the weight of the body falls upon this digit and
as the heel is raised and the footis brought into a more or less
vertical position, these tendons are put upon the stretch and
pressure results; in this metapodial, therefore, wé find the
keel present. In the other digits the sesamoids are absent
and there is no keel developed. In the upper extremity there
are sesamoids developed in the tendons of the flexor brevis pol-
licis at the metacarpo- phalangeal articulation, but owing to
lack of pressure the keel is absent. Again in the Spider
Monkey there are two sesamoids present in the tendons of the
short flexors of the great toe and the keel is developed, while
in the other digits there are no sesamoids and no keels. In
other species of monkeys on the other hand such as the Mac-
aques, the sesamoids are present and the keels are developed
upon all the metapodials. These monkeys are saitl to be less
arboreal in their habits, which would explain the difference in
the matter of sesamoids and keels. Among the Marsupials
there is no patella in the Vulpine Phalanger, and here we find
that the rotular groove of the femur is very little developed.
= Hmilut is without the sesamoids and there is no keel,
he fact that there are both sesamoids and keels
in the other metapdids These cases could be multiplied
indefinitely showing the same results. I havein fact failed to
find a single example of a metapodial, in which sesamoids are
present and pressure is exerted, which does not show the presence
of the primary keel.
I will now consider Mr. Carey’s other proposition viz: that
the crests or keels in certain highly specialized forms reach
around to the anterior face of the bone, and are not explain-
able by, nor cannot be assigned to any mechanical reason that
is obvious. I must say that when I met with this problem
several years ago, I was somewhat at a loss to understand how
any mechanical explanation could possibly be applied to its
Professor Cope’s explanation of the extension of these keels in
=
426 The American Naturalist. [May,
the didactyle foot may possibly be the correct one, but I am
inclined to doubt it seriously. It certainly cannot apply to
the monodactyle foot since it is inconceivable that any
“pinching” could occur in flexion and extension of the
phalanges upon the metapodials; for the reason that there is
no spreading such as he describes in the didactyle foot. It is
moreover manifestly impossible that the sesamoids could have
come into play to form the keel, because the amount of exten-
sion necessary to bring them into the proper position would
be so great as to cause complete dislocation of the phalanx.
How then can this extension of the keels be explained upon
the basis of mechanics?
FIGURE 1.—Coryphodon, Median digit of the Manus, posterior view. (Coll. Am.
ist
In the first place it is necessary to call attention to the fact
that at least three positions have been assumed in the various
stages of the evolution of the ungulate foot, viz: a planti-
grade or subplantigrade position, in which not only
the phalanges but the metapodials are applied to the ground
in walking, as in the hind feet of Coryphodon and all the earlier
forms; second, a digitigrade position, in which only the
phalanges are applied to the ground in walking, as in such
forms as the tapir, rhinoceros and the earlier representatives
of the horse series ; and finally the unguligrade position
in which the weight of the body is supported entirely upon the
terminal or ungual phalanges. This position of the foot
1393.] Evolution of Metapodial Keels of Diplarthra. 427
belongs to the higher forms of both the odd and the even-toed
ungulates. Now if we attempt to trace the history of the
metapodial keels in the evolution of such a form as the horse,
no one will deny, it seems to me, that we have here an
unusually complete phylum represented, which clearly indi-
cates the different stages in their development. In the earlier
members of this series, such for example as Hyracotherium, the
keels are confined to the plantar and palmar surfaces of the `
metapodials, and the animal was subdigitigrade. In such a
form as Mesohippus, on the other hand, we pass from the digi-
tigrade position of the foot to that of the unguligrade, wherein
the weight of the body was supported upon the terminal
phalanx or coffin bone. Just what induced this change is
unknown, but it is more than probable that it was in some
way connected with the reduction of the lateral toes.
We do not yet know with certainty the ancestors of the
horse series beyond Hyracotherium, but there can be little doubt
that the feet were pentadactyle and plantigrade. So far at
least as the metapodials are concerned, we have such a condi-
tion in Coryphodon, in which the keels are but faintly indica-
ted, and the two lateral grooves of which the primary keel
forms the common median wall, are in the first stages of
development. There is every reason to believe that this ani-
mal was plantigrade or subplantigrade in gait, and that the
failure to develop the keels was due to the more equal distri-
bution of the strain upon all the digits. This istrue of the
foot of the elephant in which we observe another structure
which has certainly had much to do with the retardation of
keel development viz: a very thick plantar pad. The posses-
sion of this pad not only tends to distribute the strain and
equalize the pressure but at the same time limits the amount
of flexion and extension of the phalanges. It is a fact worthy
of especial note, and one which I will make use of later on,
that in all these forms wherein the lateral grooves
andkeelsare verylittledeveloped,the plantar
borderofthearcticularsurface ofthe first pha-
lanx is not at all notched. It is also to be noted
that the feet are short and spreading.
428 The American Naturalist. [May,
With the assumption of the digitigrade position however, —
the plantar pad was reduced, the foot became more elongated,
and the strain became more localized. In this position of the _
foot the weight of the body falls upon the phalanges, which
are now bent at a considerable angle upon the metapodials,
and an especial strain is produced by the flexor tendons where a
they pass over the distal ends of the metapodials. In a foot of —
this type therefore, do we find a considerable advance in the
development of the median keel and lateral grooves of the
metapodials? Those of the third digit are most advanced for —
the reason that the strain is more concentrated and localized —
FIGURE 2.—Modern Tapir. Posterior or plantar surface of the hind foot.
at this point. In none of the metapodials however, do the
lateral grooves and keel extend more than half-way around
the articular ends, nor does the keel rise but little above the
lateral boundaries of the grooves. This is well shown in such
forms as the tapir, rhinoceros, Hyracotherium and nearly
the Eocene Perissodactyla. While the foot remained in the
digitigrade position, the median keel of the metapodials made
little or no impression upon the lower edge of the articular
surface of the proximal phalanx, for the reason that the main
flexure of the foot was between the first phalanges and the
1893.] Evolution of Metapodial Keels of Diplarthra. 429
metapodials, which would throw the keels so far back as to
seldom come in contact with the proximal phalanx. In the
middle toe or third digit, of the digitigrade foot however, the
main flexure of this toe is momentarily transferred from the
articulation between the metapodial and first phalanx to that
between the second and third phalanges in the act of walking,
and this straightening out of the first two phalanges, so as to
bring them in a line with the long axis of the metapodial, has
caused the keel to impinge upon the lower edge of the articu-
lar surface of the first phalanx which has become distinctly
notched.
FIGURE 3.—Mesohifpus. Outer view of the left hind foot. B, Distal surface of
median metatarsal, C, Proximal surface of median first phalanx. (Collection
Amer. Mus. Nat. Hist.)
It follows that in such forms as the tapir, rhinoceros, Hyra- —
cotherium, and in fact in all of the truly digitigrade Perisso-
dactyla, the first phalanx of the median digit is always
ae
430 The American Naturalist. [May,
slightly notched, while the first phalanges of the lateral digits
are without the notch. This is an important fact, and in
connection with what has just been said of Coryphodon, I think
that it may be accepted as demonstrated, that the form a-
tion of this notch is dueto the impact of the
keel upon the lower border of the phalanx. Up to this stage
therefore, we have the following conditions, viz: a compara-
tively low primary keel bounded upon either side by well
marked lateral grooves whose outer borders are as much
elevated as the keel. We also note that the grooves and keel
extend but half-way around the end of the metapodial, and
that the lower edge of the first phalanx in the middle toe is
distinctly notched. This may be termed the digitigrade
stage in the development of the keel.
In the next or unguligrade stage of this development,
beginning with Mesohippus,' the foot changed from the digiti-
grade to the unguligrade position in which the main flexure
of the foot was transferred to the articulation between the two
last phalanges, and the two proximal phalanges came to
occupy a position in line with the long axis of the metapodial.
It is in this genus therefore, that we find the first distinctive
evidence of an advance of the metapodial keel around the
lower surface toward the dorsal aspect of the bone. While
this advance or extension of the keel is slight, it is interesting
to note that the notch in the first phalanx, which we see just
beginning in the tapir, has now been transformed into a
groove and extended nearly acrossthe articular
face of the bone. Weare thus able to demonstrate that,
in the further extension of the keel of the metapodial beyond
that of the digitigrade stage, the groove was formed in the
phalanz first. It is to be further noted that the keel has
become prominent and that the lateral grooves of the meta-
podials are almost entirely obliterated, at least their outside
walls have disappeared, leaving two nearly plain articular
surfaces separated by a now prominent, or secondary
. median keel. Again, if the bones be placed in position,
*It is highly probable that the most ponies stages of this process are to be traced 2
to forms somewhat older than this genus. l
1893.] Evolution of Metapodial Keels of Diplarthra. 431
that is, if the first phalanx be articulated with the metapodial,
it will been seen that when the phalanx is placed in a line
with the metapodial or very nearly so, there is equal bearing
upon all parts of the two articular surfaces of the bones ; but
if the phalanx is flexed even to a slight extent, then the bear-
ing falls largely upon the keel for the metapodial surface and the
shallow groove for the phalangeal surface. When I say flexed
I mean bent in the direction of the plantar side of the foot.
FIGURE 4.—Modern horse. Distal view of median metatarsal or cannon bone,
showing complete keel.
How now can we apply these facts to the mechanical
explanation of the further extension of the keel toward the
dorsal side of the metapodials? When the foot assumed the
ungaligrade position permanently, we can understand, and
havé in fact already seen how the keel impinges upon and
produces the notch in the phalanx. Any flexure of the pha-
lanx upon the metapodial, under pressure would bring the
keel to a more advanced position with reference to the phalan-
geal articulation, and would cause an extension of the notch
in the phalanx so as to form a groove. ` That the phalanx is
so flexed is evident to anyone who has ever studied carefully
the movements of the foot of a horse in the act of pulling a
heavy load up an inclined surface. Here the phalanx is seen
to be greatly flexed when the foot strikes the ground and is
therefore under great strain and pressure. I speak of it par-
ticularly in a horse pulling a load, because it is more noticea-
ble in this case than in the unencumbered animal, but what
is true of one is also true of the other. Now in an animal in
which the keels and grooves were little or not at all advanced,
such as was the case with theancestor of the horse when he
emerged from the digitigrade and assumed the unguligrade
position of the foot, the flexure of the phalanx brought the
keel forward and produced a groove in the phalanx. As the-
432 The American Naturalist. [May,
phalanx went back to its more normal position, that is, in line
with the long axis of the metapodial, there would then be a
_ plane surface of bone lying immediately in front of the keel
which would be opposed to a groove in the phalanx. An
extension of the keel would necessarily follow as a result of
moulding of this metapodial surface to fit the groove. Flex-
ion, therefore, of the phalanx upon the metapodial under
pressure, bringing the keel already formed to a more and more
anterior position with reference to the articular surface of the
phalanx, thus causing it to become grooved, and the subse-
quent moulding of the keel to fit this groove, I conceive to be
the complete mechanical explanation of the production of this
structure. The various stages in the forward extension of the
keel is to be found in such series as Hyracotherium, Epihippus,
Mesohippus, Anchitherium, Hipparion and Equus.
FicurE 5.—Modern Tapir. Middle toe inside view, showing position of the
phalanges upon the ground.
In the preceding discussion, only the possibility of the meta-
podial keels having been produced and extended forward by
mechanical means, has been shown. Iwill now turn to the
matter of the application of this reasoning to the somewhat
broader question of the transmission of acquired characters,
and see whether or not any argument can be adduced in sup-
port of the proposition that the development of these structures
has been due solely to mechanical causes, and that they have
in time been transmitted to succeeding generations. Upon
this hypothesis it must be assumed that the changes took
place first within the lifetime of the individual and that they
finally came to be inherited. It must likewise be shown that
wherever a groove or a keel has occurred, there must have
been pressure, impact or strain, exerted at that particular
point sufficient to accomplish the result. The only other pos-
1893.] Evolution of Metapodial Keels of Diplarthra. 433
sible explanation of these structures is upon the hypothesis of
the accumulation of a number of spontaneous variations in
this direction, entirely independently of any mechanical
results acquired during the lifetime of the individual. Accord-
ing to this theory, all mechanical influence must of necessity,
be rigidly excluded, for the reason that it is possible for it to
have been exerted, or come into play only during the time -
that the foot was in use, and therefore, during the lifetime of
the individual. If mechanical influence is admitted even to
the slightest extent, then the whole proposition of the trans-
mission of acquired characters is at once proyen, and who can
say how much is due to mechanics and how much to natural
selection ?
In the production of the completed keel and phalangeal
groove, it must be shown by those who hold the theory of
origin purely by natural selection that they are congenit-
ally correlated structures, otherwise they must, according to the
very nature of the case, have proven injurious to their
possessor. In order for a keel to be useful it must be accom-
panied by a groove into which it is received. In other words
it must be shown that any variation in the one must have
affected the other. I shall now attempt to show that the
keels and grooves are not congenitally cor-
related. It has been shown in the preceding pages, that
the phalangeal notch which is the earliest and incipient stage
of the phalangeal groove is not correlated with the keel. The
evidence for this is to be seen in Coryphodon, Elephas, Metamy-
nodon, Titanotherium and others in which the keel exists
without any vestige of the phalangeal notch.
Again, in the tapir and rhinoceros, the lateral metapodials
are provided with distinct keels but the notch is absent.
In the middle toe of many of the lower Perissodactyla, the
keel is associated with the notch, but it was not until the foot
assumed a position whereby it was possible for the keel to
impinge upon the lower border of the phalanx that the notch
appeared. It would indeed require a great stretch of the
imagination to believe that the keel and notch were congenit-
ally correlated structures in one toe and not in the others of
434 The American Naturalist.
the same foot! In like manner it can be shown that neither
the sesamoids, keels nor grooves of the metapodials are corre-
lated structures, for we have seen that in the human thumb —
there are sesamoids present but no keels nor lateral grooves. _
If upon the other hand we look at the problem from the —
standpoint here advocated of the mechanical explanation, or —
kinetogenesis, and we admit that the bone is plastic, then we
are forced to conclude, it seems to me, that the lateral grooves _
and keels of the metapodials are the direct and unavoidable —
results of pressure exerted by the sesamoids. This is proven —
by the fact that wherever there are sesamoids present, and
es
ieee
pleted keel. Up to this point the lateral grooves and keel had
a distinct function, viz: to serve as guides for the sesamoids,
but when the change was made in the position of the foot to
that of the unguligrade, the lateral grooves began to disappear
and the keel, which we must regard in the light of a necessary
accompaniment of these grooves, began to perform a new
function. The excavation of the phalangeal notch, its exten-
sion into a groove, and the subsequent moulding of the me
podial surface into a secondary keel to fit the groove were j
as much a mechanical necessity as was the original or prim
formation of the lateral grooves and keel. The proof of
is to be found in the fact that in the further extension of
keel beyond the digitigrade stage, the phalangeal groove
the way. In the mechanical explanation therefore, we
every condition satisfied, while upon the theory of natw
selection of favorable variations the explanation is vag
unsatisfactory and not in accord’ with the facts.
; American Museum of Natural History, March 29th, 1893.
1893.) Among the Cliff-Dwellers. 435
AMONG THE CLIFF-DW ELLERS.
By CLEMENT L.. WEBSTER.
In the August number of the Naturalist, for 1891, the writer
in an article entitled “ Preliminary Notes on the Archeology of
New Mexico,” alluded to, for the first time, the
most important discovery of a Cliff-dweller Mummy, made
by him in a Cliff-dwelling of that region; and promised to,
later on, give a more detailed account of the discovery.
The present sketch then may be considered as a partial
redemption of the promise then made.
The Mogollon Mountains of southwestern New Mexico, are
one of the most rugged and broken of the great Rocky Moun-
tain system.
This region abounds in deep and vast cafions, sometimes so
profound as to almost exclude the light of day. This region
is also deeply cut and scored by the channels of the head-
waters of the Rio Gila, known as the East, West and Middle
branches.
At no other place in this great system of mountains, is there
presented so abundant and diversified scenes of wild and
romantic beauty.
A strange charm surrounds this region; and there is here
presented a remarkably rich field for the study and investiga-
tions of the Archeol
For almost everywhere here in the natural caves, in, and at
the base of the almost innumerable cliffs, are found the for-
mer abodes of the Cliff-dwellers. Some of these dwellings are
solitary, and only large enough to accommodate one or two per-
sons; while others occupy a series of large connected caves,
where are sometimes found as many as twenty-five or thirty
beautifully preserved rooms or dwellings. The writer spent
some time in this region making explorations in these ancient
ruins, sketching and drawing; and here obtained a large and
exceedingly valuable collection ” specimens.’
See report alluded to in the A f this journal, for 1891.
iy _ y a “5 F
436 The American Naturalist. reed
Extending out from the West branch of the Gila, about three
_and one half miles above its confluence with the main Gila, is
one of the most beautiful, wild and romantic canyons possible
to be found.
. The average width of this canyon is perhaps thirty-five or
forty feet, while towering cliffs rise perpendicularly on either
side to a height of from two hundred to three hundred feet K
estimate.
Through this wild glen there rushes, during the rainy sea-
son, a small stream of pure water, clear as crystal. The lower —
nearly one-half of these great cliffs is composed of a hard,
dark colored basalt, upon which the elements have made but
slight impression; and overlying them is a vast stratum of —
buff colored conglomerate, such as occupies such extensive
areas in southwestern New Mexico, portions of Arizona and
California.
Here the lower portion of this vast stratum of conglomerate
has been worn away by the slow action of the elements, leav-
ing a narrow projecting shelf of basalt extending along
horizontally for some distance. i
In the base of this conglomerate, nature has formed numer- —
ous large connected caves.
These caves the ancient Cliff-dwellers had taken possessi
of, and in them reared their strange dwellings.
Here may be seen upward of twenty-eight rooms, of di
ent sizes, shapes and designs; which taken together form the
most interesting group of these dwelling which the writer has
ever seen.
As the object of this paper is mainly to deal with the
covery of the mummy already alluded to, a detailed descrip-
tion of these and other Cliff-dwellings, ete., will be deferred to
a future report.
= One of the questions regarding the Cliff-dwellers, is
: en to the dispositions made of the remains of th
: reek very few facts have been obtained thus far, w
shed much ae upon this question
vel jes made, however, which demo
PLATE XI.
1893.] Among the Cliff-Dwellers. aat
strate that in the matter of disposing of the dead, departures
from the general custom, whatever that may have been, were
occasionally practiced. One of these exceptional cases was
brought to light in the discovery of the mummy, which forms
the subject of this paper. E
This mummy was discovered in one of the rooms of the large
collection of Cliff-dwellings just described; and had been care- .
fully placed in an excavation made in the floor, on the side
and slightly under the partition wall. The wall here, as well
as elsewhere, rested upon the debris which had previously
collected in the cave. Every portion of the body, including
the nails, hair and teeth, were perfectly preserved.
The preservation of this body was due to the great dryness
of the atmosphere of the country, and the chemical elements
of the soil, etc., in which it was entombed. The mummy was
small, being that of a child apparently about seven years of
age; although inferior in size to that of a child of the same age
with us.
It had been carefully and completely wrapped in two large
pieces of coarsely woven cloth of different textures, made from
the fibers of the “Spanish dagger”—which was used so
extensively by the Cliff-dwellers for this and other purposes
—and then again as carefully wrapped in a large and nicely
woven mat of bear grass.
After this, it was bound with cords, onto a small and >
curiously shaped board of cottonwood ; the cords crossing the
body and passing through small circular holes made in each
corner of the board.
The board had apparently been fashioned with a stone ax,
and afterward finished by being rubbed with some instru-
ment. |
The hair on the head of the mummy was of a beautiful
dark brown color, and of a soft and silky texture. The arms
were drawn up near to the sides of the head, and hands
clinched ; and the legs also were somewhat drawn up.
This description is better illustrated by the plate Figs. 1, and
2, (copied from a photograph) which represent the back and
front views of this mummy.
438 The American Naturalist. [May,
At the feet is shown a portion of the mat which formed the
outer covering, and the board to which it was bound.
There existed on the wall of one of the rooms, located a few
feet from where the mummy was discovered, many strange
and interesting characters, or picture-writings, of these ancient
people, painted in the provincial red color. |
Close by where the mummy was found, was also discovered
an ancient weaving-room, of the greatest interest. In this were
found portions of the ancient loom, woven fabrics, etc. ;
Near the mummy were discovered long braids of human
hair identical in all respects with that of the mummy, save
that it was much longer and of a coarser texture. In other
rooms of this Pueblo, were found great quantities of corn-cobs,
husks, and kernels of corn ; squash or pumpkin rinds, seeds
and stems; gourds, sandals, wearing apparel, basket and
other woven work, pottery, stone mills, weapons, utensils, ete,
ete. Such amass of evidence was secured, in connection with
the discovery of the mummy, as to leave no doubt as to the
genuineness of this relic. So far as the writer is aware, this is
the only specimen of its kind of the Cliff-dwellers, ev
discovered. a
1893.] Fossil Mammalia of Southern Patagonia, 439
NEW DISCOVERIES OF FOSSIL MAMMALIA OF
SOUTHERN PATAGONIA.
By FLORENTINO AMEGHINO.'
I.
At this time, the most important discovery in South Amer-
ican paleontology is the presence of true monkeys in the
Eocene of Patagonia. Certain paleontologists have expressed
doubts as to the age of the remains. I believe that these
doubts have no foundation from whatever point of view one
approaches the subject.
From the geological standpoint, the latest researches of my
brother, Carlos Ameghino, show that the marine formation,
which I have named Subpatagonian, and the formation of
sub-aerial or terrestrial origin that I have designated Santa-
cruzian (the formation which contains the mammals in ques-
tion), are conformable without a break. The characteristic
fossil of Subpatagonian formation is Ostrea bowrgeoisii (R. de
C.). In the east, toward the Atlantic, the lower part of the
Santacruzian formation, is found to be intersected with beds of
Ostrea bourgeoisii, representing the upper part of the Subpata-
gonian with all the beds of the two formations resting upon
each other conformably. In the west, toward the Andes, in
the lower reaches of the Rio Sehuen, and in the vicinity of
the lakes Viedrua and Saint-Martin, the marine beds of the
Subpatagonian formation rest on the Dinosaurian beds of
terrestrial origin, contemporary with the North American
Laramie (Pehuenche formation) ; the transition from one for-
mation to the other is almost imperceptible. Throughout this
region no mammalian remains have been found in any forma-
tion more recent than the Santacruzian. It is then certain
that the fossil monkeys belong to this formation. The Ho-
munculus specimens have always been found in place, and in the
lower Santacruzian beds.; It is beyond a doubt that the fos-
sil monkeys belong to the Santacruzian fauna.
1 From the Revue Scientifique, Jan. 7, 1893.
440 The American Naturalist. [May,
It remains now to ascertain if the Santacruzian fauna is
really Eocene, or if it belongs to a more recent epoch. The
affinities of the lower part of this formation with the Dinosaur
beds, the presence of Creodonts and Plagiaulacidae, the
absence of the modern groups of Ungulates, seem to me to
demonstrate plainly that we have to do here with a fauna
which dates at least from the commencement of the Tertiary
epoch, and which represents the development upon the spot of
an autochthonic fauna dating from an anterior geological epoch.
Another proof of its antiquity is that a large number of Lara-
mie mammals from the United States described by Marsh,
have their nearest allies in the Eocene of Patagonia.
On the other hand, some of the forms characteristic of the
Puerco fauna of North America are found in beds much more
recent (Lower Oligocene and Upper Eocene) of the Argentine
Republic in the neighborhood of Parana (Periptychus Cope).
It must also be said that all the authors who have studied
the conchological fauna of the rocks of Parana (d’Orbigny,
Darwin, Bravard, Doering, Philippi, etc.), have referred
this fauna to the Upper Eocene or to the Lower Oligocene.
But, between the Santacruzian fauna and the fossil fauna of
the Parana, there is, when we consider the development of
forms, a considerable hiatus, almost an abyss, that can only be
filled by the aid of a half-dozen intermediate faunas yet to be
discovered. Moreover, the greater part of the types of the
Santacruzian fauna, considered from the point of view of an
evolutionist, represent forms less advanced than do the totality
of similar forms found in other continents.
Thus all the evidences—geological, paleontological and evo-
lutional—are in favor of the great age of the Santacruzian
fauna. I place this fauna at the beginning of the Tertiary,
and consider it to be contemporary with the fauna of the ©
Puerco of North America. I believe that the more we seek to
place this fauna at a later date, the harder it will be to explain
the affinities it presents with the faunas of other continents.
Returning now to the fossil monkeys of this formation, I can
pics that my brother, in his last exploration, found some neW
Specimens, among them, a skeleton, almost complete, of Ho-
ee
1893.] Fossil Mammalia of Southern Patagonia. 441
munculus patagonicus, in the lowest bed of the Santacruzian
formation. This skeleton is imbedded in a large block of
stone, and, as yet, only a few bones have been exposed. The
lower jaw has been taken out almost intact, with its entire
dentition. This new specimen, the preservation of which
leaves nothing to be desired, shows that the jaw is not so nar-
row nor so compressed as I have figured recently in drawing
from a specimen whose symphysis was evidently distorted by
pressure. In the new specimen now at hand, the distance
between the internal posterior border and the second true
molar on each side is 10 mm., 5. It thus results that the two
dental series are more widely separated at the back, and that
the whole jaw is much more elevated.
The exact dental formula is I 3, C+, P M.§,M%. The first
two true inferior molars are very nearly of the same size, but
the third is smaller. The internal incisors are weaker than
the external ones. The canine is a little more developed than
in the first specimen, and is separated from the premolar by
a small diastema. It is probable that the two specimens are
respectively male and female. Unfortunately, the individual
represented by the new specimen, was very old, so that one
can no longer detect the details of the surface of the crown of
the molars. The dental series measure, in a straight line, 31
millimeters. The femur is 11 centimeters, and the radius 95
millimeters long. The humerus has a large epitrochlear per-
foration, but it lacks the intercondylar foramen. Each of
these bones, in form, are those of a man in miniature.
Among the other forms belonging to the Santacruzian for-
mation, and which ought to be better known, I will select only
the most important, beginning with the Ungulates which are
exceedingly interesting.
IL
I have been able to reconstruct the foot of the Mesorhinidæ
(the genera Mesorhinus, Theosodon, Pseudocoelosoma), which
are the predecessors of the Macraucheniidæ. The genus Theoso-
don had five toes in front as well as behind ; the three in the
middle were well-developed, very similar to those of Macrau-
442 The American Naturalist. (May,
chenia, but the internal and external on each foot were very
small,
The Homalodontheride also had five toes on each foot, but
with them all the toes are well-developed. The feet are strong
and robust, and it is worthy of note that the hoof-bearing
phalanges are split up, and have the same shape as those of
the Chalicotherium of Europe. The carpal and tarsal bones
have the alternate arrangement. The legs are equally robust,
and the humerus hasan epitrochlear foramen. The bones of the
feet and of the legs bear a singular resemblance to those of the
Edentates, but there are also more characters of specialization
altogether peculiar, without which they might be taken for
the direct ancestors of the Edentates. The Homalodontheride
are the ancestors of the more modern Chalicotheride of the
Northern Hemisphere. It is a mistake to look -for the stock of
the Chalicotheride in the Meniscotherid. The latter represent
a type allied to the Proterotheridx, with which they ought to
have a common ancestor as yet unknown.
-~ The Proterotheride present us with peculiarities still more
surprising. The discovery of the leg bones of the Thoatherium
prove that it was a monodactyl like the horse. Moreover, in
the Thoatherium minusculum, the reduction of the parts was still
more advanced than in the horses; the posterior feet are very
slender, provided with a single toe, the third, while the second
and the fourth are represented ouly by vestiges of the meta-
tarsals, much more atrophied than are the same bones in the
horse. One sees, in this case, an example of parallel evolution
very remarkable. A complete reduction of the bones of the
foot among the ungulates has taken place in two different fam-
ilies, at two different epochs, a fact perhaps without precedent:
in the Vertebrate series,
It is, however, certain that there exists a certain ancestral
relation between the Proterotheridse and the Equide, for the
latter have descended from a form approaching the Protere-
therium, but with complete dentition. I have elsewhere stated
that in Europe as in North America, one may be able to trace
the genealogy of the horse beyond Anchitherium (Mesohippus ’
included), and that one ought to separate definitely from the:
1393.] Fossil Mammalia of Southern Patagonia. 443
ancestral line of the horses, the genera Palaeotherium and
Hyracotherium, as well as the different types of the sub-order of
Condylarthra. In my opinion, the horses originate from a
group of Litopterna, from which sprang, at the same time, the
Paleotheriidæ and the Hyracotheriidæ, and this ancestral type
would be closely related to the Proterotheriidæ.
It would seem that I am right, for the splendid monograph
of the genus Mesohippus, recently published by M. Scott, shows
that the calcaneum of this genus still possessed a very small
articular facet for the fibula, the last vestige of the litoptern
organization, however, the shape of the astragalus, as a whole,
is already like that of a Perissodactyl.
We now have evidence to warrant the statement that the
horses are descended from a form allied to the Proterotheride,
and which ought to differ from it above all by its orbits open
behind, and its complete dentition, in a continuous series.
This ancestral is, perhaps, the Notohippus or a kindred genus,
but, be that as it may, it is undoubtedly in the Northern Hem-
isphere that the group of horses has acquired the characters
which distinguish it at the present time.
The genus Astrapotherium is now known, not only by a skull
almost entire, but by many of the bones of the skeleton. The
skull, in its anterior part, agrees in a number of points with
those of the Proboscidians, and could have carried a proboscis
as fully developed as that of the elephants. The bones of the
limbs also greatly resemble those of the elephants, and there is
no doubt that this type represents the nearest relative of the
Proboscidians discovered up to this time in the older forma-
tions. The genus Astrapotherium is certainly not the direct
ancestor of the Proboscidians, but only a collateral branch of
the trunk from which the latter have sprung. However, some
of the other genera of the family of the Astrapotheride, Astra-
podon for example, may well be considered the ancestor of the
living Proboscidians.
In any case, the latter have acquired their characters upon
some other continent, as it is certain that during the epoch of
the formation of the fossiliferous beds of the Parana, the Pro-
boscidians had no representatives in our country. These ani;
444 The American Naturalist.
mals did not make their appearance in the Argentine Repub-
lic until the end of the Miocene. ;
The new specimens of fossil rodents taken from the Santa-
cruzian formation afford unexpected opportunities for com-
parison. The fossil Cercolabinæ of Patagonia are the ancestors
of all the hystricomorphous rodents. The Stiromys have,
while young, five superior molars on each side, but the next to
myinæ. The Eocardiinæ, in their turn, have given rise to the
Caviidæ and to the Dasyproctinæ. The Octodontidæ, which
now seem to be isolated, have come from certain forms of
true origin of the Octodontidæ. There are still some inter-
mediate forms wanting, but, no doubt, they will be found soon
in the fossiliferous formation of the Paranà.
Acaremys and Sciamys, from the lower Eocene of Patagonia.
The Myomorphs are not derived from the Acaramyinæ, they
having lost their premolars and undergone various modifica-
tions in the conformation of the skull. If fossil rats have not
yet been found in the Parana beds, it is on account of their
fragile bones, but, in all probability, they will be found there
some day. : -
I now know a part of the dentition of the animal that I ha’
named Tidaeus—this name being preoccupied, I substitute :
it, Mannodon. The Mannodon trisulcatus is the first gen
Plagiaulacidæ of Patagonia which has the inferior molars
structed on the same type as those of the Multituberculata. It
1s a genus closely related to Neoplagiaulax and Ptilodus.
_ Many of the types of the Plagiaulacidw present a featu
that is strange to be found among mammals. The Epant
thidee, for example, have the inferior incisors beveled on t
1893.] Fossil Mammalia of Southern Patagonia. 445
internal border, and not on their upper face as is the case
among rodents and all other known mammals which have
chisel-like incisors the. mandibular branches of the same
animals have no symphyseal suture, the surface of the bone,
at this point, being nearly as smooth as the rest of the jaw.
It thus results that the two mandibular rami of the Epanorthide
were susceptible of lateral movements, and the singular
incisors could perform, up to a certain point, the function of
horizontal pincers.
Also, although I am quite convinced that the Plagiaulaci-
dæ and the allied fossil forms have been derived from the
Didelphian Diprotodontia, the relation with the living
Diprotodontia is not so close as I had thought. The discov-
ery I made showing that the large furrowed tooth of the man-
dible of the Plagiaulacide is not the last premolar, as had
been believed, but is the first true molar, separates the latter
from the living Diprotodontide. This character justifies a
sub-order in the Diprotodontia, to which can be given the
name Plagiaulacoidea, a name which is as appropriate for the
northern forms known as Multituberculata as it is for those of
the Eocene of Patagonia.
The recent studies that I have made of the Microbiotheriide
have confirmed me in my opinion that they represent the
direct ancestors of the living Didelphyide. The latter have
certainly originated in South America, and although their
remains have not yet been found in the Parana formations,
they will be found there some day.
One of the fossil groups of Patagonia which presents the
greatest interest is that of the primitive Carnivora, allied both
to the Dasyuride and to the Creodonts. It seems to me impos-
sible to find a line of demarcation between them. Instudying
the material in my possession, I arrive at the following conclu-
sions: The Dasyuride are descended from the Microbiotheriidz
and are then modified into Creodonts; these, in their turn, are
transformed into the more modern Carnivores. The dental
formule of these primitive carnivora are varied—take, for
example, the superior incisors, of which the Microbiotheride
have five pairs like the Didelphyde ; the Dasyuride have
30
+
446 The American Naturalist.
only four pairs, and the same number is found among ot
forms allied at once to the Dasyuride and to the Creodo;
The larger number of the Creodonts have only three pa
and, in some genera, but two, and even a single pair. Th
are some specimens in which one can trace the evolution fi
the primitive formula. Thus, in the genus Amphiproviv
the internal pair of superior incisors are entirely atrophied 4
useless, while the other three pairs are well-developed.
proves that it is the internal pair of upper incisors that is
in the passage from the Dasyurian form to the creodont
carnivorous forms.
Perhaps the most remarkable specimen discovered during
the last exploration is a skull almost entire of Borhyena tu
ata, an animal which has hitherto been known only by so
insignificant fragments. It is certainly one of the most sin
lar carnivores that is known. It must have been rather le
since its skull is 205 millimeters long, and nearly 15 cer
meters in transverse diameter at the level of the zygomé
arches. The upper arch of the skull is in an almost horizor
plane with the cerebral cavity excessively reduced, but
sagittal crest is very long and well-developed; the fron
have no postorbital apophyses, and the nasals are wide
behind in an extraordinary manner; the anterior nasal op
ing has the characteristic form of placental Carnivora. *
lachrymal canals open entirely inside of the orbits. Them
bone is prolonged behind to the glenoid cavity, of wh
border it forms a part of the boundary as in the Marsupials
The palate has no palatine foramina. The upper incisors |
reduced to two pairs, of which the internal one is atrophied.
Between the canine and the external incisor on each side, 1
large groove intended to accommodate the lower canine; t
groove replaces the deep fossa which is found in the
place in Dasyurus and Amphiproviverra. These groove
the contraction of the anterior part of the intermaxillary
followed by the reduction in the number of the incisors,
the anterior part of the skull the appearance of a rodent.
canines, premolars, and molars, of each side, form a conti
series. There are seven molars, of which three are pr
1393.] Fossil Mammalia of Southern Patagonia. 447
and four are back molars. The first have but a single cusp,
but they have a posterior heel, well-developed in the last.
The first three true molars increase regularly in size from the
first to the third, and have the anterior internal cusp atrophied,
which gives them a cutting form, especially noticeable in
in the third. The fourth is much reduced, and occupies the
same position as in the genera Thylacynus, Prothylacynus and
Amphiproviverra. Perhaps Dynamictis, of which only the
lower jaw is known, is a synonym for Borhyæna. In any case,
the latter animal was a formidable Carnivore, as dangerous as
our tigers, and which it is hard to classify. It may belong
with the placental Carnivores, or among the marsupial flesh-
eaters (Dasyures), or with the ambiguous forms known as
Creodonts.
It would take too long to speak of the numerous fossil Eden-
tates of the Santacruzian formation and of the many peculiari-
ties they present. However, I cannot resist mentioning one of
the greatest novelties of the new family of the Metopotheriide,
which comprises the genera Metopotherium, Pelecyodon, and
Zamicrus. In this group, each of the horizontal branches of
the mandibula is formed of two distinct bony parts, one in
front, the other behind. These two pieces are united by a
suture which commences at the alveolar border, very nearly
in the middle of the dental series, and is directed obliquely
back and down, terminating in the lower border, almost just
below the last tooth. This is evidently a character inherited
from the reptiles. The same suture can be seen, but not
quite so plainly, in the mandible of some of the Orthotheriide.
I remarked, in the beginning of this article, that I had
found some forms characteristic of the Puerco of North Amer-
ica in the beds much more recent than those of the Santa-
cruzian epoch. In fact, I have received from the Parana, a
fragment of a upper maxillary, which shows several tooth
_ sockets and one tooth in place. The latter corresponds to a
third premolar. It is impossible to separate this specimen
generically from Periptychus (Cope). The tooth resembles
perfectly the third upper premolar of Periptychus rhabdodon
and shows the same grooves or wrinkles in the enamel which
448 The American Naturalist. [May,
distinguishes the North American species. However, the
South American species (Periptychus argentinus Amegh.) is —
easily distinguished by its size—almost a third smaller—an
by the insertion of the molar teeth which are closer together
forming a continuous series. This specimen was found in th
Lower Oligocene or the Upper Eocene in the neighborhood of
Parana.
While I am speaking of the fossils of that region, I must —
add that the doubts which I had concerning the real nature —
of Ribodon are now set at rest. The new material at han
one of the family of Halitheriide.
TG
Argentine Republic, and particularly to the assertion of Pro-
fessor Steinmann, who maintains that the beds I have desig- —
nated Miocene and Pliocene are only Plistocene (or Quater- |
nary). *
I have not the complete work of Professor Steinmann, and
therefore will refer only to the extracts that he has published.
I am more and more convinced that the Pampean forma
tion is tertiary, and not quaternary. The Araucanian, of whic!
Professor Steinmann speaks, is entirely distinct from the Pam
pean formation. The vast difference which exists between the
Parana fauna and that of Monte-Hermoso is easily explai
by taking into account the fact that the first is Lower Oli
cene or Upper Eocene of the old geologists, while Monte-Her-
moso is Upper Miocene. Some day, when the Lower Miocem
of Catamarca (an Araucanian bed) is explored, and the mam:
malian fauna of that formation is satisfactorily known, the
great break that is now held to exist between the fauna of tf
Parana and that of Monte-Hermoso will largely disappear.
I am astonished that any one can still speak of a glac
epoch and of traces of glacial action in connection with |
Pampean, and still more so in regard to the pre-Pam
formations. The Araucanian bed is formed by the accum
‘American Naturalist, 1891, p. 855. :
-< 1893.] Fossil Mammalia of Southern Patagonia. 449
lations of volcanic detritus. The Pehuelche is of fluviatile
origin. Is it necessary to repeat for the hundredth time, that
there is not the least vestige of glacial action in the Pampean
formation ?
It is impossible to name single fossil species from the Pam-
pean formation which is characteristic of a colder climate
than now prevails in the same country. On the contrary, not
only does the mammalian fauna indicate a warmer climate
` than at present, but the fresh water conchological fauna also
furnishes similar evidence. The many fossil plants of the
` Pampean formation are the same that are seen to-day in the
province of Buenos-Ayres, or in the northern part of the
Argentine Republic—not a single form belonging to a cold cli-
mate. I am almost certain that an examination of the marine
_ shells of the same formation would lead to the same result. I
am about to make a collection of them to send to M. von
Jhering, whom I will request to study them from this point of
view.
450 The American Naturalist. [May, |
EDITORIALS.
EDITORS, E. D. COPE AND J. 8. KINGSLEY.
—WE have had occasion to refer in previous numbers of this journal
to the difficulties under which the Geological Survey of Georgia has
labored, by reason of the attempts of a political board to assume its direc-
tion. This board appears to consist of men ignorant of Geology, and
who use the survey as an opportunity for giving places to their friends.
The director of the Survey, Dr. J. W. Spencer, has not been permitted —
to direct, nor even to select his subordinates, so that chaos hasbeen the _
natural result. Ignoring the fact that this state of affairs has been due
to their own incompetency, the board has asked for the resignation
of the director of the survey. This Dr. Spencer has very properly —
refused to furnish. In a letter to the Governor he presents a strong
. defense, which is an arraignment of the board of a convincing character. —
It is evident that the board should resign, rather than the director of —
the Geological Survey. If they will not resign, the board should be —
abolished. It appears that their misgovernment has not been confined ES
to the Geological Survey, but that many of the educational interests of —
the State have suffered at their hands. If Governor Northen desires
the acquisition of accurate information as to the material resources of —
Georgia, he can not do better than retain Dr. Spencer in his present
position. If he regards the fair fame of Georgia, he will use what
power he has to abolish the present board, and when a competent one
is selected, it will permit the State Geol
—A RIDICULOUS bill has been introduced into the Legislature of
Pennsylvania with the following title and provisions, by Representative
1893.) Editorials. 451
Jno. H. Fow, of Philadelphia, “ (No. 163) An Act to prohibit public
exhibitions of physical or mental deformities in certain public places.”
“ Srcrion 1. Be it enacted by the Senate and House of Represen-
tatives of the Comn.onwealth of Pennsylvania, in General Assembly
met, and it is hereby enacted by the authority of the same, that on and
after the passage of this act, it shall be unlawful for any person to
exhibit in any public hall, museum, theater or any other building,
tent, booth, or public place, for a pecuniary consideration or reward,
any insane, idiotic or deformed person or any imbecile, and whoever
shall exhibit such mental or physical deformity, shall be guilty of a
misdemeanor, and, upon conviction thereof, shall be sentenced to pay
a fine not exceeding one thousand dollars, or suffer imprisonment not
exceeding six months or both, or either at the discretion of the Court.”
If this bill becomes a law, it will require the closing of scientific
exhibitions of abnormalities to students where a fee is charged, as well
as the more popular exhibitions of a miscellaneous character. It will
thus deprive the public of a real source of instruction, which is important
from various points of view. It will deprive scientific men of the
opportunity of examining many curious pathological specimens which
the industry of the exhibitors brings together, and which would be
otherwise generally inaccessible to them. The Alexis St. Martins,
Siamese Twins, double monsters and other such specimens, which are
most important for physiologic research, would no longer come to
Philadelphia. It will also deprive many unfortunate persons of
making a'very good livelihood, who would otherwise become charge-
able to their relatives or to the State. What good is to be attained by
the passage of such a bill it is impossible to imagine.
—Tue New York game protector, Mr. Willett Kidd, and the Forest
and Stream periodical, are to be congratulated on the successful result
of their endeavors to enforce the game protection law of the State
against the restaurateur Delmonico, of New York City. For having
in his possession eighteen woodcock killed out of season, Delmonico
has been compelled to pay a fine of $25 apiece—total, $450.00. This
result was only attained after persevering efforts, prolonged through
many months. The delay was due to the culpable neglect of Prose-
cuting-Attorney Nicholl, of New York, who would have permitted the
culprit to escape, had it not been for the interest of Judge McCarthy,
who took up the case himself. We trust that the game and fish laws
of all the States may be enforced as effectually, and a great deal more
promptly.
452 The American Naturalist.
RECENT BOOKS AND PAMPHLETS.
LLEN, J. A.—On a Collection of Birds from Chapada, Matto Grosso, Brazil, Brazil,
op Mr. H.-H. Smith. Part II, Tyrannidae. Extr. Bull. Am. Mus. Nat. Hist,
iv, 1892. From the author
Annual Report of the Poatiaster- General of the United States for the Fiscal
‘ending June 30, 1892.
=~ ARANZADI, D. T. De.—Fauna Americana. Ateneo de Madrid. Leida eld
; de Abril, de 1891. From the author.
BEECHER, C. E.—Development of the Brachiopoda. Extr. Am. Journ. S
xli, 1891. From the author.
Bissins, A.—On the Distribution of te in the Chesapeake E
and the Character of its Habitat. Extr. Trans. Maryland Acad. Sci., Vol. i,
From the author.
BOETTGER, O.—Drei i: colubriforme Schlangen, Separat-Abdrucks
Zool, Anz., No. 405, 1892
—— Katalog der Herr so af im Museum der Senckenbergischen
forschenden Gesellschaft in Frankfurt a. M. From the author.
Bouter, M.—Notes sur le Remplissage des Cavernes, Extr. de L’Anthre
1892.
——Decouverte d'un squelette d’ Elephas meridionalis dans les cendres t
are du volcan de Senéze (Haute-Loire). Extr, Comptes Rendus, 1892. F
ct D. G.—Address delivered on Columbus Day, Oct. 21, 1892,
Library and Museum Building of the University of Pennsylvania, Phila. Fre
author,
Bulletin Agri. Exp. Station of the Rhode Island College of Agric
Arts, No. 19, 1892.
Mechanic ;
Bulletin Government oe Experiment Station for North Dakota.
1892.
Bulletin Massachusetts State Agricultural Experiment Station, No. 45, 1892.
. Bulletin North Carolina Agri. Exper. Station, No. 87, Sept. 15, 1892.
1393,] Recent Books and Pamphlets. 453
FABRINI, E.—Su Alcuin Feliné del Pliocene ergg Estratto Rendiconti della
R. Accad. dei e Vol. i, 2° Sem, Serie 5a, fasc. 7.. From the author
FAIRCHILD, H. L. Paie of the Fourth Annual Meeting, hid at Columbus,
Ohio, Dec. 29, 30 and 31, 1891. Extr. Bull. Geol. Soc. Am., Vol. iii, 1892. Fro:
the Society.
GARMAN, I.—The Reptiles of the Galapagos Islands..——0On Reptiles rai by
G. Baur near Guayaquil, Eucador. —-On Cophias and Bachia——On n Rep-
tiles. Extr ll. Essex Inst., Nos. 4, 5,6. From the a
Horer, W. aT anatomische Studien über die Nerven des Armes und
- der Kas bei den Affen und dem Menschen. From the author.
HOLEROOK, M. L.— Microscopical Researches of the eed Elements of
- HOLLAND, -—On the Occurrence of Riebeckite in India. tea and
Physical Hotes s on Rocks from the Salt ae Punjab, Extr. Records Geol. Surv.
India, Vol. xxv, pt. 3, 1892. From the
owEs, G. B.—On the Pedal me ” the Dorking Fowl, with Remarks on
Dama and Phalangeal Variation in the Amniota, .- Extr. Journ. Anat. and
Physiol., Vol. xxvi. From the author.
Hoyt, J. W.—Memorial in regard to a National University, From the author.
Hutton, F. W.—On the ag of the Struthious Birds of Australasia. Extr.
Proceeds. Australasian Ass. Adv. Sci., 1892. From the author.
ress Pieces at Méchante’ ee Boston, on Public Benefits of
the Work of the Ags Association, From the
KELLocG, V. L.—Common Injurious Insects of a From the author.
Kuprrer, C. V.—Ueber die Entwickelung von Milz und Pankreas. Extr. Mün-
chener medicinsche Abhaudlungen. 36 Heft., V. iii, Reihe., 4 Heft., 1892. From
the author
. Letter from the Secretary of the Interior transmitting in response to Senate resolu-
tion of July 16, 1890, information relative to the Geological Survey
MARCOU, J.—A Little More Light on the U. S. Geological Survey. From the
author,
ASON, O. T.—The Birth of Invention. Extr. Proceeds. From the author.
Norris, H. W.—Studies on the page of the Ear of Amblystoma. Extr.
Journ. Morph., Vol. vii, 1892. From the
pera C. V.— Directions nat Colle an oe he serving Insects. Part F, Bull. U
S. Natl. Mus., No. 39. From the Smithsonian Institution
Sainz, L. DE H., AND T. DE PARERE Avance à fe Autopologia de Espafia,
Madrid, 1892.
SEELEY, H. G.—The Mesosauria of South Africa. Extr. Quart. Journ. Geol. Soc.,
Nov. 1892. stags: the author.
SLINGERLAND, M. V.—The Pear-Tree Psylla. Bull. 44, Cornell University Agri.
Exper. Station, ee 1892. From the author.
SMITH, a A.—Sketch of the Geology of Aiea From the author.
E. S.—Statistics of Constitutional and Developmental EES of the
Jaws and Teeth of Normal, Idiotic, Deaf and Dumb, Blind and Insane Persons.
Extr. Denta meathouroany July, 1889.—— The Jaws and Teeth of a Party of Cave and
Cliff-Dwellers. ification of Typical Irregularities of the Maxillee and Teeth.
454 The American Naturalist. r] i
Reprint NR hpne, 1889.—— The Differentiation of Anterior Protrusions of
pper Max d Teeth. Read before Internat]. Med. Congress, Aug. 4, 1
From the rere
TOWNSEND, c. H. J.—Notes on North American bine sens, str., with
descriptions of New Genera and Species. Extr. Trans. Am. Ent. Soc., Vol. xix, 1892.
——On a Leaf-Miner of Populus fremonti, Se on Two Mexican Species of
Ceroplastes, with a Record of Parasites reared from one.
——New Jamaica Tachinide.——New North American Tachisi Extrs. Entomo.
News, 1892.——North American Genera of Calyptrate Muscidae. Extr. -Ta
Ent. Soc., Vol. xix, 1892.——A Sarcophagial Parasite of Cimbex a. Extr.
Can. Ent., 1893.——An Aporia bred from Limacodes Sp. Extr. peek June, ag
From the author.
ANDERVEER, A.—Hystero-Epilepsy, with report of cases. Extr. Trans. Med.
Soc. N. Y., 1892. From the author.
WILLIAMS, G. H.—The Volcanic Ss aki of South Mountain in Pennsylvania and
Maryland. Extr. Am. Journ. Sci., Vol. xliv., Dec., 1892. From the author.
ia D, H, C.—The Relation of Neuropathic Insanity to Crime. Reprint Internatl.
nds July, 1892. From the author,
"Wan T, G. F.—Unity of the Glacial Epoch. Extr. Am. Journ. Sci., Vol. xliv,
Nov. 1992. From the author.
1893.] Recent Literature. 455
RECENT LITERATURE.
A Summary Description of the Geology of Pennsyl-
vania. Vols. I and II.'—The results of the Geological Survey of
Pennsylvania from its beginning, in June, 1874, to the close of its field
work, June 1, 1890, have been summarized by Professor J. P. Lesley, in
three octavo volumes of 700 to 800 pages each, of which Vols. I and
II have just been issued, while Vol. III is promised in June, 1893.
This summary is compiled from the reports published by the Board
since 1875, with the corrections, and the additions that the accumula-
tion of fresh facts make necessary. Vol. I describes the most ancient
deposits: The Laurentian granites, gneisses and mica schists; the
Cambrian gneisses, schists and limestones ; the lower Silurian lime-
stones and iron ores, roofing slates and mountain sandrocks ; Forma-
tions II, III and IV, in the order of advancing time and superposition.
The second volume describes the Upper Silurian red shales, limestones,
and fossil iron-ore beds; the glass sand-quarry rocks; the black and
gray shales, and the gray and red sandstones, with fish beds; Forma-
tions V, VI, VII, VIII and IX reaching to the top of the Catskill
Mountain rocks, the upper limit of the Devonian system.
Each formation is discussed in detail. Its geographical distribution,
thickness, lithological characters and fossil contents are carefully
described, together with its relation to similar formations of neighbor-
ing states.
The question of sedimentary Archean rocks receives attention, and
while discussing Archean rocks in general, Professor Lesley calls atten-
tion to the fact that the Huronian rocks of Canada are nowhere recog-
nized along the Highland Belt from the Hudson to the Schuylkill. In
the author’s opinion, the term Huronian, “ must be used simply as a
proper and private name for a series of rocks exposed along that
part of the northern boundary of the United States.” In the paleozoic
series, Professor Lesley does not clearly discriminate between the Cam-
brian and Ordovician limestones, including them all under the one
head of No. II, as was done by the first geological survey. He leans
toward the explanation given by Dr. Fraser of the structure of the
Chester Valley, rather than those of Hall and Rogers.
The illustrations are numerous, many of them being photo-electro-
1 A Summary Description of the Geology of Pennsylvania. Final Report ordered
by Legislature, 1891. Vols. I and II. By J. P. Lesley, Director, Harrisburg, 1892.
456 The American Naturalist.
type reductions of the drawings made by the assistant geologists.
plates of fossils accompany the chapters of the several format
These are half-sized reproductions of the figures of fossils given
Report P. 4, Dictionary of the Fossils of Pennsylvania and the
rounding States, published in 1889-1890. ;
The present report has been evidently written with an eye tothe
general rather than the scientific public. As a popular synopsis of
the geology of Pennsylvania, it will have much utility. Asad
sion of the more abstruse problems presented by the structure of
formations of the State, especially of the eastern region, it has
value. a
The Earth’s History,’ by R. D. Roberts, is an attempt “to
nish a sketch of the methods and chief results of geological eng
such as a reader interested in the subject for its own sake would ¢
- to obtain. It is not intended to be a text-book of Geology.” I
_ ning with a brief history of geological thought, it takes up the
history and shows how geologists are enabled to trace the succe
steps in the development of the earth’s physiography, by reference
the records of the rocks as interpreted by means of the results of
tigations into the processes at present active on and beneath its sur
The central idea of the author is “ to reconstruct, from ancient
mentary remains, the old conditions that characterized the succe
stages in the evolution of the land areas; to make out the life
- of the earth,” and this idea has determined the method of
the subject matter. The book is not crowded with geological fae
1s not a dry abridgment of some popular manual of geology, but
exactly what its author intended it should be—an exposition
logical logic. Its style is simple and clear, its statements are
and its various parts are well proportioned. In short, it is an ©
volume for collateral reading for college classes in geology, and
Introduction to the geological methods for general students.—W.
_ Wright’s Light,’ although mainly a discussion’ of light
nomena that may be projected upon the screen by means of a
: with the lime light, is nevertheless, an excellent volume for §
___*The Earth’s History, an Introduction to Modern Geology, by R- D.
A Pri 4 Apren fan ~ York: Scribner's Sons, 1893, pp-
ae ‘Light, nt, a course of Experimental Optics, chiefly with the Lantern.
_ Wright; 2d Ed, Macmillan & Co., pp. 301, PI. 9, Figs. 207. Price, $3:
1893.) : Recent Literature. 457
the students in the theory of light—natural and polarized. The value
of the work to the student of polarized light lies in the fact that many
experiments are described that aid one to gain a clear notion of double-
refraction, of interference, and of the resolution of vibrations into two
sets at right angles to each other, without compelling him to accept the
conclusions of mathematical processes. Scattered between chapters
dealing with the experimental data are others, in which these data are
explained, and a few in which the theory of light vibrations is dis-
cussed, the whole forming a logical exposition of the modern theory of
light.
Unfortunately for the reader, there are many clumsily-constructed
sentences in nearly every chapter of the book, which need much close
study before the meaning of their author can be discerned. This fact
necessarily detracts from the delight with which the volume would
otherwise be perused, but not so much so as would be the case were it
not so compactly written. An immense amount of material is collected
within its covers, and very little of this matter could well be spared by
the student. A little more confidence might have been felt in the
author’s statements, as representative of the most modern views of
light specialists, had he given some evidence of his acquaintance with
the wonderful grating of Rutherford and Rowland, instead of mention-
ing only Noblet’s gratings and Barton’s buttons in his discussion in
diffraction. The imperfections, however, are slight as compared with
the abundant good qualities of the book.
The volume is well printed and splendidly illustrated with a
wealth of cuts and nine lithographic plates, of which four are beauti-
fully colored.
The student of light phenomena will find many of his difficulties
in understanding the theory of his subject cleared away if he will care-
fully examine the experiments described by Mr. Wright, and thor- -
oughly digest the accompanying explanations—W. T. B.
Co, and, as this name is both short and pretty, it may be hoped tl
458 The American Naturalist.
General Notes.
GEOGRAPHY AND TRAVELS.
Africa.—Isra, on Brivisn East Arrica.—Although the con
nent of Africa is now gay with the colors which distinguish the sphen
of influence of the various European countries, and though it has no
been crossed and re-crossed in almost every conceivable directio
there are still considerable areas about which very little is kno
This is especially true of the regions adjoining and intervening betwe
the great lakes of the Equator; those vast fresh-water areas ¢
known to the ancients, but entirely lost to the moderns until the
ent generation. The politico-religious troubles in Uganda, which
endeavored to convert into reasons for the abandonment of that
try by the British, have directed considerable attention toward th
region, and Captain Lugard, whose name has been prominent in ti
recent troubles, has not been slow to furnish information, The Briti
sphere of influence in this quarter is separated on the north from
Italian by the river Juba, while to the southwest a line drawn
the center of the east shore of Victoria Nyanza, to a point on the ¢
near the island of Penba marks the edge of the German sphere.
line is however, bent northward around Kilima njaro, so that the’
of that snow-clad elevation is within the German sphere. At
toria Nyanza the Anglo-German boundary is deflected
Upon the west, the British protectorate is bounded by the Congo®
The territory of Uganda stretches along the north and nortl
shores of the great lake, extending inland as far as Unyoro,
skirts to the southeast shores of the Albert Nyanza; east
Uganda toward the Albert-Edward Nyanza, lies the district of
The whole of the British sphere in northeast Africa has
name of “ Ibea,” formed from the initials of Imp. British East
u will persist when the company itself is a thing of the past.
Captain Lugard’s route to Uganda was from Mombasa to t
_ Tiver Sabaki, tothe south of Mount Kenia, which is entirely
British sphere. According to him, the source from which
Victoria Nyanza derives ite supply of water is among
1893.] Geography and Travels. "459
geographical puzzles of Africa. All the rivers flowing into the lake
are very small, andthe broad Somerset Nile which flows out of its
northern coast certainly carries away more than is received by their
combined streams.
The greater part of the waters which fall upon the Uganda region
seem to make their way either to Lake Rudolf or Lake Baringo, and
we must therefore assume that Victoria Nyanza is fed by springs.
The western shore of Lake Albert is faced by a lofty plateau, the
waters of which flow, not to the Nile basin, but to the Ituri, a tributary
of the Congo. The Senliki River, which, under various names, unites
the Albert-Edward or Mwutanzige Lake with the Albert Nyanza, is
relatively small where it issues from the former, and its considerable
volume at its entrance into the latter is due to the affluents which flow
from the western extremity of the snow-capped range of Ruwenzori.
Though the eastern portion of this range drains directly into the
Mwutanzige, the waters flowing from the western part are sufficient to
convert the lower portion of the Senliki into a deep and rapid river.
Adjoining the Victoria Nyanza on the northeast is the fertile district
of Kavironda, where much grain is grown and where the natives, a
good-natured and merry set, wear no clothing of any kind unless the
tusks of wart-hog, horns and wings with which the warriors decorate
their heads can be considered as clothing. Passing west from Kavi-
rondo to Usoga, a district tributary to the Waganda, the character of
the agriculture suddenly changes, the grain giving place to bananas,
potatoes and cassava, though no marked difference of soil or aspect
can be detected in the surface of the country. Moreover, the transit
is from nudity to clothing, for every native of Usoga, like the Waganda,
is dressed in the mbugu, an ample garment manufactured from fig-
bark. The dress of both sexes is the same, but while the mbugu of
Usoga is black, that of Uganda is brick-red. East of Kavirondo is an
equatorial plateau seven to eight thousand feet above the sea, appar-
ently well-suited for the residence of Europeans, and gradually sloping
to the Kavirondo lowlands. In various parts of this plateau are still
to be found remnants of the once powerful nation of the Gallas whose
_ forces have been exhausted in the struggle with the Masai to the south
and the Somals to the north. A curious feature of the districts of
Uganda and Unyoro, according to Captain Lugard, is the scarcity of
running water, and the occurrence of swamps in every depression.
The average elevation of Uganda is 3,700 feet, while Unyoro, though
not greatly dissimilar in general aspect is higher, and has loftier hills.
The Waganda possess great natural aptitude, are clever carpenters
i A The American Naturalist. [May,
and smiths, and can be readily taught European methods of working,
They are fond of music, have many rude musical instruments, and pos- —
sess a currency of cowries. Mentally, they are certainly superior to
the surrounding races. Uganda is divided into ten provinces, each of:
which has its chief, while over all is set a vizir. The system of land-
tenure is very complex, and chieftainships are complicated. The »
northern part of Ankoli is thickly peopled, but the land has suffered
terribly from the cattle-plague, which has reduced all the pastoral ~
tribes of this region to a state bordering on starvation. Not only have
millions of domestic cattle fallen before this dread disease, but the
buffalo has been practically exterminated, while the eland and the
smaller kinds of antelopes have suffered severely.
The Albert-Edward or Mwutanzige is really a double lake, for the
long northeastern arm known as Ruvango is only connected with the ©
larger body by a river not more than 500 yards across. This lake is
abundantly supplied; the Mpanga alone brings down more water
from Ruwenzori than is taken out by the Senliki. y
During his circuit from the Victoria by the Albert-Edward to the
Albert Nyanza and Wadelai, Captain Lugard picked up the Sudanese
refugees left by Emin Pasha, and upon his return was accompanied by _
some 9000 people. He concluded his address before the Royal Geo-
graphical Society with a graphic picture of the present disordered state of
the countries west and north of the Uganda, smitten by the cattle-plague,
oppressed by a usurper named Kabarega, and only prevented from
freely accepting protection, from the fear that all protectors, like Stan-
ley, will depart and thus leave them a helpless prey to the vengeance
of their oppressors. ‘sf
Uganda is a region of rounded grass-clad knolls, while Unyoro is more
rugged, often with fantastic piles of granite, and abounds in caves-
The rivers of Uganda are large papyrus swamps with no current, and
little open water. The hills are of red marl or shaly gravel. In past
times Uganda had broad roads, often with culverts across water, 40° —
even now it would be easy to make a good road to the Albert Nyanza.
More grain is grown in Unyoro than in Unganda. A survey for |
projected railway from the coast to Victoria Nyanza has already been
made for 350 miles.
There seem to be several small lakes in Buddu, a province í
Uganda west of the Victoria Nyanza; in Koki, a tributary state as
of Buddu, and in Ankoli, the eastern part of which has loftier hills
and poorer soil than Buddu. One of these lakes, between Anko’
and Koki, is named Kashera. Iron seems to be abundant in the
a
1893.] Geography and Travels. 461
districts. The natives of Ankoli are Wahuma, a fine race apparently
with affinities to the Galla and Somali, but mingled with Bantu.
The Tana, a large river south of the Juba, has been navigated by
Captain E. G. Dundas in the stern-wheel steamer Kenia, to within a
short distance of the mountain of that name, which was afterward
ascended to a height of 8,700 feet. The Tana has two mouths, the
northern one, the Ozi, is not now important, but the entire district
between it and the Tana mouth is alluvial. The lower portion of the
river is almost meridianal, but close to the equator a broad westerly
curve conducts to the congeries of affluents which descend from the
forest: covered slopes west of Mt. Kenia. The steamer could proceed
as far as Hameyé, near the equator, at the start of the great bend.
The peoples met with in the ascent were the Kalindi at the mouth, the
Ngao, Omoina, Ndera, Guano, Ndura, Sabaki, Malululu, Malakote,
Wasania, Korokoro, Wandorotu, Murdoi, Wathaka, Mumomi, Mbe,
Kikoya and}Muea near Mt. Kenia.
The smaller Sabaki river to the south of the Tana, also has its head-
waters in the forest belt S. W. of Kenia. The existence of such a river
as the Tana, navigable through more than three degrees of latitude,
was until recently pratically unknown. A considerable fall about
189 south of thejequator, and sixty feet high, terminates the navigable
part of the stream.
It may be remembered that Mt. Kenia has been ascended by Count
Teleki to a height of 15,350 feet, yet he estimated that there was still
above him 3,500 feet to the highest summit. The forest reaches to
3,500 feet, bamboos to 10,500, after which the vegetation as far as the
snow-line at 15,000 feet consists of mosses and the curious tree Senecio
johnstonii. The slope toward the west is very gentle, but southward
the mountain presents a serrated ridge four miles in length.
An expedition has started for the Juba River commanded by Lieu-
tenant C. R. Villiers, and accompanied by J. W. Gregory of the
British Museum.
Asia.—A Vistr to Prerk-Tu-SHan.—Mr. Carles has recently paid
a visit to the singular white-topped and flat-topped mountain in
Mantchuria, from which issue the sources of the three great rivers
Tumen, Yalu and Sungari. This mountain, which has upon its sum-
mit a lake some twelve miles in circuit, at an elevation of 7,500 feet,-
had been previously been visited by Mr. Younghusband and his party,
but Mr. Carles was moved to undertake the journey from Seul, the
capital of Korea, by reading an account of it in a Chinese work.
31
ee
462 The American Naturalist.
This account, exaggerated though it was, giving to the mounta
height of five miles, and to the lake a proportionately expanded
cumference, showed that there really existed a remarkable eleva
Chan-y-Peik, the “ Ever White Mountain,” is well-wooded to the
and thus presents a contrast to the isolated Peik-tu-Shan or
White Mountain the summit of which is bare, and white |
pumice. It is the latter which encloses the lake.
The account of the journey, in the March issue of the Proceedi
Royal Geological Society is full of interesting matter concerning
ways of the large-hatted Koreans, their threshing-floors of beaten ¢
the pickled cabbages and turnips so dear to their palates, their ¢
and domestic animals, among which the black and hairy pig is í
spicuous. The untidiness of a Korean village offers a marked
trast to the neatness of a Japanese one, but Mr. Carles gives
Korean rustics credit for a more thorough appreciation of nat
beauty than is possessed by any other people. A Korean who has
climbed a summit stops, not to complain of the toil, but to admire
landscape. Ham-Heung, the chief town of N. E. Korea, is a wa
town with twenty-five to thirty thousand inhabitants.
Tot KACHINS AND THE IRAwADIL—The recent expedition of
Indian Government among the hills of the rebellious Kachins, Chi
paws, or Singphos, who occupy much of northern Burma, toward
Chinese frontier, though they have added to our geographical r
ledge of the upper waters of the Irawadi, have not solved the my:
of the Salwin. The upper Irawadi divides into two principal st
the Mali Kha to the west, and the more easterly Nmai Kha.
course of the former has been mapped as a result of the Kachin
ditions, but little is really known of the latter, save that at its
with the Mali Kha it is somewhat the larger.
The Kachins seem to be descended from the Karengs, and hi
ages been subject to hereditary chiefs or Sawbwas; but of late
many villages have rebelled against these, driven them out or
them, and these villages are now only governed by ‘headmet
little authority. This, together with the fact that the peaceful
villages of the Shans and Chinese are each under the pre
The district is not one of lofty mountains, but of rugged
three thousand feet in height, intersected by well-watered
1393.] Geography and Travels. 463
good deal of opium is grown, but not enough to fully supply this
opium-consuming race. :No reliable information can be obtained
respecting the head-waters of the Nmai Kha. It is thought that the
Phung Mai, east of Kanti and the Nmai Kha, may be the latter,
while the Lu is the Salwin. All the water that falls upon the land
up to within a few miles of the Lu drains to the Irawadi, which thus
rapidly develops into a noble river, though it is doubtful whether any
of its sources are farther north than 28° 30’.
East Asta Minor.—In these days of exploration of the unknown,
some of the districts that have been longest known, and which fill large
pages of the world’s history, seem to be least visited. It is this which
lends additional interest to D. G. Hogarth’s account of his travels in
Asia Minor—that land of many peoples and of many ruins which has
for centuries been the stronghold of the Turkish power. Mr. Hogarth
not only examined many ruined cities, but penetrated through several
little known passes of the Taurus and Anti-Taurus. In the district
between Lakes Egerdir and Beysheher there are no passes, but the
Eurymedon cuts its way through a great gorge, and the population
interests from its primitive and apparently indigenous character.
The Pisidian city of Adada, now Kara Bavlo, perched high on its hill-
top, is a most perfect specimen of an Anatolian city of Roman times.
Not very distant are the ruins of Lystra and Derbe.
In a small island upon Lake Egerdir are the remnants of a small
colony of Greeks, who in the twelfth century deliberately preferred
Mohammedan to Byzantine rulers. The monastery of Koja Kalessi is
a remarkable ruin, containing a church of the fourth or fifth century,
restored in the reign of Justinian, and evidencing the power still
possessed by heathen traditions in the entire absence of Christian sym-
bolism from its sculpture. ‘Ihe city of Coropissus is a veritable
Pompeii, the church probably of the seventh or eighth century, most
of the ruins alittle later. The east Taurus is richly wooded, and is
traversed by the tremendous canyons of Samanti, Saros and Jihan, not
passable even on foot. Several important passes occur farther east
than the famous Cilician Gates. This district was the last refuge of
the independent Armenians of Cilicia, and their robber towns, Hadjin
and Zeitun, still exist. The half-troglodyte Kurds, nominally Mussul-
mans, but with pre-Mussulman customs, have penetrated to these parts.
The Anti-Taurus, deserted since the eleventh century, is now inhabited
only by the nomad Ayshar and some Kurds, who found their way there
about fifty years ago. The great Roman road to the Euphrates can
i
464 The American Naturalist. Eo
still be easily traced in the valley of the Saros, and has many mile-
stones. Tne fortified tower of Maidan dates from the first or second
century, when the Cilicians were lords of the Mediterranean. Among
so-called Hittite monuments, one of the most important is the obelisk
at Albistan. This the inhabitants refused to sell to some Germans,
the Government heard of it, and took possession without payment. It
has sixty-seven lines of inscription on all four sides.
NORTHEAST SIBERIA.—The region of the Verkhoyansk and Stano-
voi Mounts, in N. E. Asia, with the upper basins of the Kolyma,
Indigirka and Lena, have been recently explored by M. Cherski.
Proceeding up the Chandyga River, Cherski reached the heart of the
Verkhoyansk, whose summits are below the snow-line. Then by way
of the river Dyba, a tributary of the Tyra, he reached the valley of
the Omekon, which stream the natives regard as the true head-waters —
of the Indigirka. The Uchagei-urach does not, as shown on the maps,
empty into the Omekon. The Verkhoyansk was then crossed at its
junction with the Stanovoi range, where the latter runs eastward.
The region between the Indigirka and the Kolyma, covered by the
bend of Stanovoi, is split into several subdivides, of which the expedi-
tion crossed three, and found the third to be the water parting between
the Indigirka and Kolyma basins, the Moma flowing from one side,
the Syrjanka, an affluent of the Kolyma, from the other. The valleys-
of this district are broad with gentle slopes, and without terraces,
which are destroyed by the shifting courses of the rivers, the tendency
of the latter being rather to fill up the valleys with rubbish than to
deepen them. On the southern slopes magnificent poplars and willows
flourish, and the meadows have a rich flora. Notwithstanding the
high latitude, the temperature in June and July rose to 113 in t
sun, though for fourteen days in August it sank below 32. The gr
longitudinal valleys seem to have possessed immense glaciers in
istocene period; while the mountains are formed of Silurian
Jurassic folds.
Astatic Norrs.—The recent expedition of Lieutenant H. S. Wal-
ker, from Napeh in Upper Burma to Arakan, possesses considerable
interest, were it only for the visit paid to Myohaung, the ancient cap-
ital of Arakan, now peopled only by some 2500 souls. The ancient
magnificence of the place is attested by the great ruined walls, fort
three squares one within the other, and by two gigantic pagodas,
1893.] Geography and Travels. 465
with eighty, and the other with ninety thousand images. This city
was known to Ptolemy, who speaks of it as Triglyphon.
The outward route was through the An Pass, while the return route
was eighty miles to the north. The former was found far the better
for a railway, though beyond Dalet it was cut up by creeks, many of
which, however, could be missed by hugging Myaintaung Hill.
Lieutenant Walker speaks of the Arakanese as indolent and cowardly
beyond measure.
W. M. Conway, with a party, started in February of the past year
to explore the glacier regions of Baltistan and Karakoram, southeast
of Pamir. News of his expedition has several times been received,
but no full account has yet been given. He has been the first
European to cross the Nashik pass; has explored the Hispar Glacier,
a vast level sea of snow of 300 square miles, surrounded by a ring of
giant peaks—the largest glacier outside of the polar circle; has
ascended many lofty peaks, among which is one which he has named
Pioneer Peak (over 23,000 feet) because it was found to precede the
Golden Throne, some 2000 feet higher still.
A new map of Persia, compiled from a great number of general and
divisional maps, may be found in the issue of the Proceedings Royal
Geographical Society for February, 1892. It is on a scale of 600 miles
to the inch, and includes Afghanistan and Beluchistan. The bounda-
ries of the famous Persian desert, the Dasht-i-Kavir, with its two large
salt-swamps, and the Kuh-i-Gugird, are clearly shown. The desert is
shaped like a dumb-bell, and the largest salt-swamp is in the centre of
the western portion.
Captain Bower and Dr. Thorold left Leh in June last year, crossed
the widest part of Tibet, and passed a chain of salt-lakes, one of
which, Bor-Ba-Tu, is probably the highest lake in the world, since it is
situated 17,930 feet above sea-level. Their course lay from Ladak to
Chiamdo, about 150 miles to the north of the route of Pundit Nain
Singh. Though ordered back by the Tibetan authorities, they made
their way through Tibet to west China.
Burma and Assam seem temptingly close upon the map, and are
only separated by about a hundred miles of hill country, yet a satis-
factory road from one to the other has not yet been found. The high-
est hills or mountains of the district range from but four to eight
J
466 The American Naturalist. [May, '
thousand feet, but as they are clothed with dense forest and impenetra- —
ble jungle, with few inhabited or cultivated spots, the passage is very —
difficult. The political officer of Assam, Mr. Needham, after having
had experience of one route, tried that by which in past times the
Burmese armies used to find their way to conquer Assam. This leads
across the Patkoi, among hills below 3000 feet in height, to Nongyong —
‘Lake, and then via the Loglai to the Singpho village of Ham Yung, —
thence to Ningbu, three short marches from Maingkhwan, ;
Australasia.—AusTRALIAN ExPpLoration.—The news of Austra- —
lian exploration is always tame and little varied, like the aspect of —
that “ scrubby ” interior. Dr. Lindsay and party crossed the space
between the courses of Forrest and of Giles. They found the Queen —
Victoria spring on the latter course nearly dry, and the country seemed —
to have had no rain for two years, Various bushes were found, alsoa —
gum-tree forest which extended into 8. Australia. From the roots of
certain mallee trees, which the natives know by experience, they pro- —
cure a quantity of pure water. ‘
In South Australia the country between lakes Eyre and Amadeus has
been explored, and seems to be rather more varied than most of the —
interior, as several ranges of hills, the Musgrave, Everard, Mann, ete, —
have been found, rising four to five thousand feet above the sea. The ;
Musgrave hills are of red granite, which appears to predominate inthe —
other ranges. Spinifex, a few pines, stunted gums and mulga, are the —
prevalent trees, but between the Musgrave and Mann ranges is 4
growth of large casuarinas,
In West Australia the country between Northam and Eucla has
White gum and gimlet-wood were the most conspicuous trees.
land seems rich in minerals. The natives seen were of much
physique than those which frequent the towns.
New Gurnea.—Sir W, Macgregor continues his work of vis
the various tribes of British New Guinea with the aim of bringing the
to acquiesce in the government and laws of Great Britain. G
progress has already been made in this matter, so that already
1893.] Geography and Travel. 467
villages which were built upon piles for security are now removed to
dry land. The want of recognized hereditary chiefs among the natives
resent causes some difficulty, but the coast tribes have to a great
extent accepted the change, and the inland tribes, who were for the
most part driven inland by the coast residents, are commencing to
come in. The area of the British portion is 86,000 square miles, and
the population is probably 350,000. On their part the Germans are
doing similar work. Between Pouro and Milne bays there is a ridge
of lava and coral limestone, rising to 850 feet. The close relationship
often supposed to exist between the flora of New Guinea and that of
Australia seems scarcely borne out by the facts. Though Proteaceæ
and Myrtacee abound in the savannahs of Fly River, yet the palms
are numerous in species, and at least fifty indigenous plants are
known.
Sir W. Macgregor has been specially engaged among the D’Entre-
casteaux and Trobriand Islands. The latter are a little known group
of small islets, with one language, and a population of some 15,000,
The most interesting discovery has been that of a number of atolls
which have, since their formation, been elevated by a horizontal uplift.
Kitava or Nowan has a surface of 5.6 square miles, and is now girt
by a fringing reef: almost its whole circuit is surrounded by a low
sloping margin about a quarter of a mile wide, covered with trees.
This abuts against a steep coral wall three to four hundred feet in
height, also covered with forest. Within this wall the land dips gently
to a plateau fifty to a hundred feet below the edge of the wall, plainly
the ancient lagoon. The soil in this interior plain is of a rich choco-
late tint, and very fertile. All the natives reside in this protected
area, which is drained by filtration through the porous coral rock.
Kwaiwata is similar, but much smaller. Gawa is a still more perfect
specimen, having a coral wall four hundred feet high, and so steep
that it must beclimbed by ladders, while the interior is a 100 feet
lower. On approaching these islands from the sea they seem to be
uninhabited, since the natives live in the saucer-like hollow. In Iwa
the raised border has been worn away. A great portion of the south
of Fergusson Island, one of the D’Entrecasteaux group, is occupied by
the mountain mass of Edagwaba, 4-5000 feet high, composed of mica
schist, and in the N. W. corner Kubioia rises to 3-4000 feet. The
islets of Namu and Bagiagia are of coral, low and uninhabited. On
each of the small Trobriand Islands there is a single village.
468 The American Naturalist. 0
Polar Regions.—Tue Danisu Exprprrion.—Lieutenant R
of the Danish navy, contrived in spite of the floating ice whic
extended from three hundred to three hundred and fifty miles f
Greenland, to enter Scoresby Sound last year, and to discover that:
was really an extensive fjord with several branches. Hurry’s Inlet,
branch toward the north, proved to be a fjord 28 miles long, wit
gneissic cliffs 3000 feet in height on the east, and westward the crag
of Jameson’s Land, 2500 feet high, seemingly composed largely |
glacial moraine. Many Jurassic and Tertiary fossils were found í
Jameson’s Land, while at Cape Brewster, where the cliffs rose
from 300 to 500 feet, fossils of older date were found.
The southern shore of Scoresby Sound presents a lofty and unbroke
granite wall sixty miles in length, the ground rising in the interior
3000 feet. The widest inlet is named Halls Inlet, and trends n
west. On its shores are roches moutonnées and strie in abundane
but noice. The inland ice is not met with until a distance of 1
miles from the sea is reached, it is found at the heads of the sm
bays and fjords, and all the fjords seem to reach it. The gneiss
to 500 feet in the southwest, to 3000 in Milne’s Land (west of Halls
Inlet) and to 6000 feet in the northern region explored—on the westit
is covered with basalt. Animal life proved to be rich, especially
Jameson’s Land, where reindeer occurred in wonderful num
while the musk-ox was found on Hurry’s Inlet. No less than }
species of flowering plants were gathered. No inhabitants were
with, but winter houses and graves were seen.
PoLAR Nores.—Several whalers have left Dundee for the An
tic, in the hope of combining discovery with a good catch.
Dr. Drygalski, in connection with the Geographical Society of Bert
has erected an observatory between the Great and Little Karajak gl
iers, on Umanak Fjord, West Greenland.
Coal has been discovered in Spitzbergen by L. Cremer.
w
1893.] Geology and Paleontology. 469
GEOLOGY AND PALEONTOLOGY.
The Western Lowland of Ecuador.—A paper by Dr. Th.
Wolf, read at the Berlin Geographical Society, Dec. 3, 1892, gives the
following account of the Western Lowland of Ecuador :—
Its superficial area is 25,950 square miles. As late as the end of
the Neocene period the waves of the ocean washed the foot of the
Western Cordillera of Ecuador. At the commencement of the
Plistocene period a part of the northern half of the lowland appeared
above the ocean while the southern half was covered over by an
immense sandstone formation. The latter, containing here and there
the bones of mastodons and horses, was, at a very recent period, sub-
jected to upheaval, which gave to the country very nearly its present
configuration.
e great low-lying plain east and west of the Gulf of Guayaquil as
well as the smaller alluvial plains, are quite recent formations, true
deltas. The highest points of the coast ridge attain an elevation of
2,300 feet in a chalk range between Guayaquil and the sea, and in the
Cordillera of Chongon and Colonche. In the Neocene region there are
summits of from 650 to 1000 feet. The Plistocene marine strata form
a gently undulating country rising from 65 to 260 feet above sea-level,
while the extensive plain of alluvium is quite level. The rivers which
issue from the Western Cordillera, do not flow due west ; but, in con-
sequence of the chalk range which runs north and south, form two
extensive river systems, Rio Guayas emptying into the Gulf of Guaya-
quil, and Rio Esmeralda, flowing without a delta into the sea at 1°
N. latitude. (Geog. Journ. Feb., 1893).
Devonian Fossils from Manitoba.—During the summer sea-
son of 1888 and 1889, Mr. J. B. Tyrrell made a collection of the fossils
of the Devonian rocks of the islands, shores and immediate vicinity of
Lakes Manitoba and Winnipegosis. This collection forms the subject
of a report by Mr. J. F. Whiteaves, published by the Geological Sur-
vey of Canada. According to the author, this collection is one of the
most important that have been brought back by the Survey collectors
for many years. The species represented are of unusual interest, not
only on account of the number of new forms, but also as showing the
close relations that exist, in so many respects, between the fauna of
these rocks and that of the Devonian rocks of Europe.
470 The American Naturalist. [Me
Mr. Whiteaves’ report comprises 105 pages of text illustrated by 15 _
full page plates. The classification followed, with few exceptions, is
that adopted by Dr. Karl Zittel in his Handbuch der Paleontologie.
(Contributions to Canadian Paleontology. Vol. I, Pt. IV, 1892).
Jura and Trias in Taylorville, California.—Professor Hyatt
gives the following preliminary results of an examination of four
different collections of intervebrate fossils from the Jura and Trias of
the Taylorville region in California.
A general comparison of the Trias of Taylorville with that of Idaho
and of the Star Peak range in the Humboldt region, Nevada, shows
that the Idaho Trias hasa well marked Triassic fauna, with fi
cephalopods recognized in Europe by Mojsisovics, Steinmann and
Karpinsky, as belonging to the lower part of the Triassic system.
This fauna appears to be more nearly the equivalent of that of the
‘Werferner beds of the middle Buntersandstein of the German Trias
than of any other.
The Trias of the Star Peak range contains an unmistakably younger
fauna. The species show a parallelism with the Muschelkalk instead a
of the Saint Cassian stage, as has been supposed.
The Trias of Taylorville is quite as interesting as that of the other
two localities, and it is suggestive that its age, as indicated by the fos- 2
sils, is that of the Noric and Karnic series in the upper Trias. :
The Lower, Middle and Upper Jura, are represented by character-
istic fossils which can be closely compared with representative
European species. No remarkable or entirely new types occur, such
as have been found among the vertebrata of this continent. A
On comparing the Taylorville with that of Aurora, Wyoming, and
of the Black Hills, it is found that the latter were deposited in
same basin, the species being largely identical, and that they can te
spoken of together as having characteristice of the fauna of |
pavia or Oxfordian in the upper Jura of Europe.
The supposed Callovian of Mount Jura has no species in commod
with those of Aurora or the Black Hills. This accords with the 4
_ tinct fauna of the Bicknell sandstone and the Hinchman Tuff,
would go to show that there was no direct connection between
upper Jurassic faunas of the Rocky Mountain region.
The Oolite of the Rocky Mountain region is too little known to Pi
mit comparison. Dr. White has described a few fossils found by
Peale near the lower cafion of the Yellowstone in Montana.
1893. J Geology and Paleontology. 471
The lower Lias, having certain forms of undeniably European
facies, occurs in western and southwestern Nevada, and perhaps in
California east of the crests of the Sierra. These faunas are older than
those found at Mount Jura.
It is obvious that the Jura occurs in widely separated patches, and
that, so far as now known, Mount Jura exhibits a larger number of frag-
ments of the series of the Jurassic system than any other locality in
the United States. (Bull. Geol. Soc. Am., Vol. 3, 1892.)
The Post-Laramie Beds of Middle Park, Colorado.—Mr.
Whitman Cross, in a paper read before the Colorado Scientific
Society in October, 1892, points out the unconformities that exist
between the Middle Park Beds of Colorado and the underlying rec-
ognized Laramie formations, and calls attention to the difference in
lithological characters. A study of specimens collected at different
points show that a large series of andesite rocks strongly resembling
those that constitute the Denver conglomerates take part in this for-
mation. The Lignitic beds also resemble those of the Denver beds.
In regard to the fossil plants Mr. Cross finds that 12 out of 21 species
from the Middle Park beds occur in the Denver beds. These facts,
the author believes, justifies him in correlating the Middle Park beds
with the Denver, and he agrees with Mr. R. T. Hill in assigning them
to the Post-Laramie group.
It has been suggested by Mr. J. B. Hatcher,' that in view of the fact
that at least two species of Agathaumas have been described from the
Denver beds, that it would be well to take into account the evidence
from the vertebrate fauna before settling definitely the age of the beds
under discussion. (Proceeds Col. Sci. Soc. 1892).
Marine Pliocene Beds of the Carolinas.—The tables of the
Molluscan fauna of the Waccamaw and Croatan Beds of the Carolinas
recently published by Dr. Dall, establish the presence of genuine
Pliocene beds in both the Carolinas. The Pliocene of Tuomey and
Holmes is shown to be based on a confusion of species belonging to at
least two horizons, and hence the classifications based upon the
supposed characters of this non-existent fauna may now be consigned
to oblivion. (Trans. Wagner Free Inst. 1892.)
Geological News.—General.—Mr. Ralph S. Tarr has called
attention to the resemblance between the terraces now forming on the
1Amer. Journal Sci., Feb., 1893.
472 The American Naturalist.
Colorada River in Texas, and the older ones of the Connecticut
He suggests that the modes of formation may be identical. In
case the floods and sentiment supply arise chiefly from the c
accident of desiccation, while in the other they are the result of
accident. (Am. Journ. Sci. July, 1892.)
M. E. Ficheur has added an important fact to the knowledge of
geology of northern Africa. He finds that on the northern flan
the Atlas Mountains in Blidah (Algeria), and throughout the ext
of the region occupied by the Chiffa schists, there is an anteclinal
affecting the whole sedimentary series, a fold long drawn out
turned toward the north, This phenomenon appears to be the re
of a lateral thrust from the north against the resisting mass of
Chiffa schists which formed, to all appearance, an island in the C
aceous Sea. This folding took place after the Carté¢irenne period,
very probably after the Helvétien stage, for these beds are seen to
disturbed at the end of this range on the slopes of the spur on the
side of the Harrach. M. Ficheur is of the opinion that the foldi
limited to the Blidah range. (Revue Scientifique, Feb. 1893.)
Paleozoic.—In his notes on the Devonian formation of Mar
Mr. J. F. Whiteaves remarks that in Manitoba the Stringoceph
zone is remarkably clearly developed, and holds a rich fauna, whe
in the Mackenzie River district, most of the fossils so far coll
seem to be from the Cuboides zone. (Geol. Mag., Feb., 1893.)—
Rollin Keyes calls attention to a well defined Kinderhook fauna i
ealated in the Burlington limestone in northeastern Missouri.
fossils, chiefly molluscan, are found in a heavily bedded white e
limestone with a peculiar white chert in nodules and i
bands. The forms are all species which characterize the Kin¢
of Burlington, Iowa. This seems to be an illustration of Ba
celebrated “ doctrine of colonies, ” (Am. Journ. Sci., Dec., 1892)
Mesozoic.—According to Mr. H. W. Fairbanks the granitic
of southern California are not Archean. They are present as
sions squeezed into rocks varying in age from the Jurassic to F
inclusive at the close of the Jurassic. (Geol. Mag., Feb., 1893.
Mr. A. Smith Woodward reports the skeleton of a fish from the
Clay of Wiltshire, England, which he “provisionally quotes
immature example of Ischyodus egertonii.” The fossil tends
firm the reference of Ischyodus-like fishes to the existing
1893.] Geology and Paleontology. 473
Chimeride. (Ann. Mag. Nat. Hist., Jan., 1892.) ——The fossil
fragments of jaws and teeth described by Gervais under the name
Lepidosteus suessoniensis have been referred by M. V. Lemoine to
the genus Champsosaurus Cope. M. Dollo has recently published a
critical comparison of Gervais’ Lepidosteus with Champsosaurus to
show that there is no point of resemblance between them, but that they
present numerous and radical differences, and that Gervais’ reference
is correct. (Bull. Scien. de la France et de la Belgique, 1892.)
Cenozoic.—Sejior Nogués, who has been exploring the Andesian
volcanoes of Chillan, confirms the supposition of Pissis (Geografia
fiscia de la Republica de Chili, 1875, Paris) concerning the existence
of moraines of two distinct epochs, and therefore of glacial periods, one
of them being anterior to the present volcanoes. This explains the
orography of the region. (Geographical Journal, Feb. 1893.) Mr.
G. Cotteau, who is studying the fossil echinoderms of the Eocene of
France, has recently described a number of new species belonging to
the genera Coptosoma (Desor), Liosoma (Cotteau), Micropeltis (Pomel)
Circopeltis (Pomel), and Gagaria (Duncan). The latter genus is
represented in France by six new species belonging to the middle
Eocene. (Revue Scien., Jan., 1893.) Professor Cope recently
exhibited to the Academy of Philadelphia a nearly entire mandible of
the Tetrabelodon shepardii (Mastodon shepardii Leidy) from the
Blanco (Pliocene) of Texas. He showed that it differs from the known
species in the decurvation of the mandibular symphysis, which
equals that of the corresponding part of the jaw of Dinotherium, but
lacks the incurvature of that genus. He shows that the Mexican
species formly referred to that species is distinct, and he proposed for
it the name of Mastodon oligobunis.
474 The American Naturalist.
BOTANY. 4
North American Fungi, Century KXIX.—Subscribers to _
Ellis & Everhart’s North American Fungi were gratified recently by s
the receipt of the 29th volume of this great collection, with its neatly —
mounted and labelled specimens. It is one of the “ miscellaneous” —
volumes, nearly all groups of the fungi being represented. Thus we
find five Myxomycetes, among which is the new Stemonitis virginiensis
Rex; the recently described Peronospora echinospermi Swingle; a —
dozen or so Uredinez, some but recently described ; Erysiphe graminis
in frait ; many Pyrenomyceteæ, some Hymenomycetes, and a good many
“ imperfect fungi,” the latter well represented by species of Phyllostica —
and Septoria—Cuarves E. BESSEY. :
New York Fungi.—A new and interesting distribution of speci :
mens of fungi has been undertaken by Mr. C. L. Spear, of Alcove, N. —
Y., under the title of “ New York Fungi.” It is designed particularly
to illustrate the work of the well-known mycologist, Charles H. Peck, i
who, in the capacity of State botanist, has for so many years been —
engaged in the study of the fungi of New York. Century I contains —
the following species: Amanita cesaria Scop., A. phalloides Fr., A.
musearia L., var. alba Pk., A. vaginata Bull., Tricholoma album Schell,
Collybia radicata Relh., Mycena galericulata Scop., M. leaiana Berks —
Omphalia campanella Batsch., Pleurotus sapidus Kalchb., Clitopilus
cæspitosus Pk., Naucoria pruinatipes Pk., Hypholoma aggregata Pk
H. sublateritium Bchæff., H. candolleanum, Fr., Coprinus picaceus ta
C. micaceus (Bull.) Fr., C. plicatilis (Curt.) Fr., Lactarius deceptivus —
Pk., Cantharellus floceosus Schw., Marasmius oreades (Bolt.) Fr, W —
rotula (Scop.) Fr., Panus stipticus (Bull.) Fr., Schizophyllum commune
Fr., Lenzites betulina (L.) Fr., Boletinus pictus Pk., B. porosus (Berk.) a
Pk., var. opacus Pk., Boletus subluteus Pk., Polyporus perennis (E)
Fr., P. sulphureus (Bull.) Fr., P. fumosus (Pers.) Fr, P. adustus
(Willd.) Fr., P. pubescens (Schum.) Fr., P. applanatus (Pers.) Ft»
P. fomentarius (L.) Fr., P. conchatus (Pers.) Fr., P. hirsutus (Wulf)
Fr., P. pergamenus Fr., P. versicolor (L.) Fr., Dedalea confragost —
(Bolt.) Fr., D. unicolor (Bull.) Fr., Glæoporus conchoides Mont., Hyd-
num zonatum Batsch., H. graveolens Delast., H. coralloides Scop., Irpa
paradoxus (Schrad.) Fr., Phlebia zonata B. & C., P. radiata FT.
Craterellus cornucopioides (L.) Fr., Thelephora sehweinitzia P
1393.] Botany. 475
Stereum rugosum Pers., S. acerinum (Pers.) Fr., Hymenochete corrugata
(Fr.) Lev., Corticium salicinum Fr., Cuphella tilie (Pk.) Cke., C. fulva
B. & Rav., Clavaria botrytis Pers., C. formosa Pers., C. stricta Pers., C.
fusiformis Sow., Spathularia flavida Pers., Guepinia spathularia (S.) Fr.,
Lycoperdon gemmatum Batsch., L. pyriforme Schæff., Scleroderma vul-
gare Fr., Phragmidium subcorticium (Schrank.) Wint., Puccinia pecki-
ana Howe, P. graminis Pers., P. circææ Pers., P. pimpinelle (Strauss)
Lk., P. sorghi Schw., P. malvacearum Mont., P. mariæ-wilsoni Clint.,
P. fusca (Relh.) Wint., P. podophylli Schw., Uromyces junci (Desm.)
Tul., U. polygoni (Pers.) Fekl., U. hyperici (Schw.) M. A. Curt., Ræs-
telia lacerata (Sow.) Fr., Æcidium clematidis DC., Ustilago caricis
(Pers.) Fkl., U. avenæ (Pers.) Jensen, U. erythronii Clint., U. utriculosa
(Nees) Tul., Sphacelotheca hydropiperis (Schum.) Schrot., Urocystis
waldsteinie Pk., Microsphera alni (DC.) Wint., Hypocrea richardsoni
B. &. M., Hypomyces lactifluorum Schw., Diaporthe acerina (Pk.) Sace.,
Hypoxylon fuscum Fr., H. blakei B. & C., Pezicula carpinea (Pers.)
Tul., Cenangium cerasi (Pers.) Fr., Patellaria rhabarbarina Berk.,
Aphia tigillaris B. & Br., Monilia fructigena Pers., Coniosporium
rhizophilum (Pr.) Sacc., Trijara americanum Thüm., Septoria
enothere West.—CHARLES E. BEssEY.
Seymour and Earle’s Economic Fungi.—The fifth fascicle
of this set was distributed February 1st, 1893. It includes numbers
201 to 250, represented chiefly by Uredineæ parasitic on woody plants,
many numbers (226 to 250) being devoted to the gymnosporangia,
with their aecidial stages. Accompanying the set is a systematic list
of hosts, from which one learns that the 50 species of fungi are parasitic
upon 35 hosts, representing 13 families of flowing plants.
Of the usefulness of this set to students of the injurious — little
need be said. It is simply indispensable.—CHARLES E. Bess
Halsted’s Weed - Seeds.—One of the most useful sets of speci-
mens recently distributed is the “One Hundred Species of American
Weed-Seeds,” issued by Dr. B. D. Halsted, of New Brunswick, N. J.
It consists of 100 small screw-capped vials, each containing a good _
quantity of cleaned seeds or achenes, in the condition in which they
usually occur as impurities in horticultural and agricultural seeds. It
supplements most admirably the two centuries of “American Weeds,”
issued by the same author.—CHARLES E. BEssEY.
Morong’s Naiadacez.—Students of the Naiadacee will be
greatly helped by the sets of named specimens which Dr. Thomas
476 The American Naturalist.
Morong, of Columbia College, is now distributing to subscribers,
set recently received contains 83 North American and Eu
species and varieties, representing quite satisfactorily this differ
group. When the author's “ Revision of the North American
ace,” now in the press, is received, there will be little reason
dents and collectors neglecting these interesting acquatics.
E. Bessey.
Canadian Mosses and Lichens.—Several years ago, Dr.
Macoun began the distribution of sets of “Canadian Musei;
material secured in his extensive collecting expeditions in the
ests of the north and west. During the winter the fifth century
to hand, including numbers 501 to 600, some of which are inte
new species.
With this century of mosses came Century I of “ Canadian Li
which promises to be fully as interesting a distribution as the
The specimens are ample, and the labels are of the same neat
those of the mosses——CHArtes E. Brssry.
1893.] Zoology. ; 477
ZOOLOGY.
The New England species of Balanoglossus.—In 1873,
Alex. Agassiz described a Tornaria from New England which he
ascribed to the common Balanoglossus of the region. Bateson later,
studying in Virginia and North Carolina, showed that in B. kowalevskii
there is a direct development. The question at once arose, what form
did Agassiz have? Either his Tornaria did not belong to B. kowalev-
skii or the same species has a different development in different regions.
Now Prof. T. H. Morgan announces’ that he has found Bateson’s larvee
with the Balanoglossus kowalevskii on the southern coast of New Eng-
land. This clears up one problem but it still leaves the origin of the
Tornaria questionable.
Marsipobranchs.—Beard reports’ that in young Myxines in which
the pronephros was functional, the teeth were present in several rows
upon the roof of the mouth. Apparently there is no Ammocete stage,
but there is a metamorphosis. He further finds that at least in one
specimen of a male Petromyzon in about one section in forty of the
testis there was a welljmarked ovum, occupying a follicle for itself.
The Lateral Line of Siluroids.—Pollard has been tracing the
cephalic divisions of the lateral line system in several Siluroids’ and
while his work is largely descriptive and admits of no abstract, one is
struck with the support he brings to the old view of Huxley that the
Arthrodira are near relatives of the Siluroids and especially to the
close affinities of Clarias and Coccosteus. The parallels in the canals
of the two forms are very exact. Oñ the other hand Pollard points out
a close resemblance between the African genera Clarias and Auchen-
aspis and the South}American, Chetostomus. The facts remains, that
while careful studies of the lateral line like these of Pollard may in
future throw much light upon the phylogeny of various groups of fish-
like forms, we have as yet not data enough for much generalization.
Prodromus of a new System of the non-venomous Snakes.
—The unsatisfactory character of the existing classifications of the
1 Zool. Anz. XV, 456, 1892.
? Anat. Auz. VIII, p. 59, 1892.
z eya arrea ita
478 The American Naturalist.
non-venomous snakes is well known. That authors are not ag
to the principles on which these animals should be classified
learned by comparison of the systems published by Duméril and
in 1853, Giinther in 1858, myself in 1886 (Proceeds. Amer.
Society), and Boulenger in 1892 (Reptilia of Zoology of British
It had appeared evident to me that a further examination of the a j
omy of these reptiles is necessary before a correct account ot i
mutual relations can be given, and that the organs of the reproducti
system especially, were likely to yield important results. W rt
object of obtaining light on this question, I have made an exami a
of the hemipenis, and have obtained valuable indications of relationsh
which have been hitherto unknown. I have had the opport
examining material from the collections of the Academy of Phila
of the U. S. National Museum, and of my own cabinet. I present
a general synopsis of the results, reserving for a future occas
publication of a fuller illustrated memoir on the subject. The
classification is more in harmony with the systematic indi
obtained from the study of other vertebrata, than those hitherto a
which is itself an indication of its greater approximation to
Some points remain obscure, and many details are omitted fi
present prodromus.
The hemipenis of the Ophidia is traversed by a groove which
the muscular investment to the internal integument (or external
ument when the organ is retracted), which commences at t
internally, and soon turns to the external side of the organ al
tinues to its extremity. This is the sulcus spermaticus. This
always bifurcated in venomous snakes, and I find it to be equi
cated in many harmless snakes. The investing tissues may
correspond with this bifurcation. Thus the hemipenis may
nay bifureate. we states that it is bifurcate in venomous
ut it is not so in Hydrophis hardwickii, B us semifasciatus,
cephalus coronatus, ete., while it is bifureate in many masm
Next to the bifurcation of the sulcus in importance, is the nature
surface of the external investment (internal when retracted). 4
most perfect types both venomous and non-venomous, this
a suctorial function. Their borders are generally
i s0 deeply divided into papille as to lose their orig
These papillæ may be the seat of osseous s
bristles or spines, which become larger toward the middie oft
and lose their mutual membranous connections. These isol
1893.] Zoology. 479
may extend to the apex, but they rarely extend to the base. The sur-
face may, however, be laminate and not recticulate, and the laminæ may
be longitudinal or transverse. In either of these cases they may not
be spiniferous. The apex or apices of the organ may be furnished
with a rigid papilla or awn. The entire surface of the organ
when protruded, is designed for the maintainance of its position in
the oviduct of the female, from which it cannot be withdrawn, except-
ing by invagination.
In the Tortricina and Peropoda, the hemipenis is not spinous, and
the sulcus is bifurcate, and in the Boidae the hemipenis is bifurcate also,
although in some genera (Xiphosoma, Ungualia), the branches are very
short. The external integument is never reticulate, but is always
laminate with elongate papill at the extremities, in Epicrates, Xipho-
soma, and Ungualia. The laminae are pinnate from the sulcus as an
axis, in Morelia, Enygrus, Lichanura and Eryx, and are transverse
in Charina. In Ilysia they are pinnate, with a few longitudinal plicæ
below.
The general definitions of the families exclusive of the Peropoda are
as follows.
No spines; surface longitudinally plicate ; Calamariide.
The surface of the hemipenis is flounced more or less transversely; Lycodontide.
F ace is more or less reticulate, and the sulcus spermaticus is undivided ;
kei eae anterior ; Colubride.
The surface is reticulate or JongimdinaHy plicate, and the sulcus is divided ; hypa-
pophyses anterior Xenodontide.
The surface is Dras reticulate nor flounced, and the Peson when present are dis-
connected; hypapophyses continued to caudal verteb Natricide
The Calamariidæ approach in the oon of the hemipenis to
such Peropoda as Eryx. The character assigned to the Lycodontide
is more or less distinctly present in the typical or Solenoglyph venomous
snakes; while the Najide (exclusive of Elapide) on the other hand,
cannot be distinguished from the Xenodontide, by any general char-
acter. The Lycodontide are Old World with a single genus in America;
a distribution resembling that of the Pythonide. The Colubride
inhabit the Old World and North America, a few genera entering
uth America. The distribution is like that of the firmisternial
By tia. The Xenodontidsx are of South America and Madagascar, a
few genera entering Africa and North America, a distribution nearly
like that of the Iguanian lizards. The Natricide are distributed in
the Northern Continents, very few types occurring in Africa and none
in South America
480 The American Naturalist.
It will be observed that all of the families include burrowing typ
and that the Colubride and Xenodontide include also arboreal type
This result is in accordance with the general rule that adaptations
present environment are not indications of the deepest affinities.
the families include glyphodont genera. The genera included in
families are the following : |
CALAMARIIDÆ.
maria gervaisii the plice are well developed.
formerly referred to Simotes belong here. In Holarchus* ((
ancorus, the sulcus is undivided, and the distal part of the exi
side is membranous. In the H. trinotatus the sulcus and hemip
. shortly bifurcate, for which reason I refer it to a distant genus
the name of Dicraulax. Ihave had access to very few types of
group, and consider the present arrangement provisional.
LYCODONTID&.
I. Aglyphodont; sulcus spermaticus undivided.
No palatine or pterygoid teeth. (In Ovigodon subguadratus the only
the genus examined, there are two robust papilla on the extremity of the he
ni
Hemipenis simple; palatine and pterygoid teeth ; Lye
II. Aglyphodont ; sulcus spermaticus, and generally hemipenis, a,
III. Glyphodont ; sulcus and hemipenis bifurcate ;
IV. Glyphodont; sulcus simple;
To the Lycodontine belong Lycodon and Anoplophallus (=
ops Hallow.) which has a long loreal and no preocular gene ;
spines on the hemipenis. To the Boddontine belong Boddo
COLUBRID.
The subfamilies are as follows:
gaer S. Natl. Museum, 82, 1887, p. 54.
1893.] Zoology. 481
Fusiform. Chilomeniscus; Fici imia; Geagras; Cemophora; Rhinochilus; Con-
opsis (has papillæ at the apex in addition to a few cups). ; Stylosoma (spinous
nearly to apex.)
Colubriform. Contia; Hypsiglena (pocketed below apex) ; Proterodon; Dianodon ;
Coronella; Ophibolus; Rhinechis ; ie awa (four rows of spines dividing
calyculate apex); Salvadora ; T e ; Epiglot aed pansies Spilotes ;
Coluber; Bascanium; Drymobius ; Zam seals Ptyas ; Herpetodryas ; itn wet
Liopeltis; Cynophis (has a ki papilla which is pro! pan ed into an awn
Crossanthera, (g. n.) established for Deudrophidium melanotropis Cope, on
account of the total division of the walls of the small cups into papillæ
Attenuate, Dendrophis; Leptophis; Bucephalus; Dasypeltis (papillæ spinous to
apex.)
In the DrrsaprxÆæ the same gradation appears.
Fusiform. Tantilla; Scolecophis; Ogmius (spines to apex).
Dipsadiform; (generally pocketed below apex). Sibon ; Trimorphodon ; Crotapho-
peltis; Himantodes; Rhinobothryum; Dipsas.
Attenuate. Cladophis; Oxybelis; Dryophis (has a diverticulum simulating a bifur-
cation of the hemipenis) ; Tragops ; Passerita.
The CHRYSOPELEIN® includes only the genus Chrysopelea, so far as
I have examined.
Genera of Colubride in which the calyces are not papillose, are
Phyllorhynchus, Hypsiglena, Dianodon, Proterodon, Coronella, Sym-
phimus, Dendrophis, Crotaphopeltis and Dipsas.
XENODONTID£.
The arrangement of the genera of this family is difficult, and what is
presented here is only tentative. It seems probable that some genera
with a grooved posterior tooth are more nearly allied to others with a
smooth tooth than to each other; but on the other hand the external
form of the animal is a poor guide, as all forms pass into each other. So
also the reticulate or plicate character of the integument of the hemi-
penis. Many of the genera may be distinguished by details of structure.
As before, I designate the general forms as fusiform, colubriform,
attenuate, ‘and dipsadiform ; and for the present I adopt two sub-
families. a
Aglyphodont; ; Xenodontine.
Glyphodont ; i Scytaline.
XENODONTIN &.
Fusiform. Catostoma; Carphophiops; Farancia* ¢; Pseudoeryx; Ninia.
Colubriform. Homalosomat; Grayia; Theleus g. n.; Diadophis; Rhadinæa*
Pliocercus ; Srema Liophis*; Dromicus; Alsophis; Lianthera g. n. ;
Hypsirhynchus*; Amastridium ; a Xenodon* ; Acantebpiadat s
Lystrophis* ; Heterodon.
482 The American Naturalist.
Attenuate. Uromacer.
Dipsadiform. Leptognathust, Mesopeltist.
The genera of this subfamily in which the surface is exclusively
gitudinally plicate are marked with a star; on the other genera
more or less reticulate. The genera in which the extremity is spir
are marked with a f. In Theleus the hemipenis is simple, and
apex is covered with short separate papille, below which it is coar
spinous; type Boodon virgatum Hallow. from W. Africa. Li
is established on Herpetodryas bernieri of Madagascar, and rel
species, on account of the absence of calyculi, and weak develo
of spines of the hemipenis, and the isodont dentition. The
Acanthophallus is designed to include the species formerly refe
to Xenodon in which the hemipenis is spinous to the extremity.
pe is X. colubrinus Gthr; the anal plate is entire. In Xen
the extremity is smooth and plicate, and the anal plate is divided.
SCYTALIN®.
Fusiform. Wydrocalamus.t
Colubriform. Pseudophis; Erythrolamprus*; Tachymenis+; Coniophanes;
phis; Rhinostoma; Scytale; Oxyrrhopus; Tomodont; Thamnodynastes
cludes Tachymenis hypoconia M.), Tropidodryas; Philodryas; Jaltris.*
Altenuate. Langaha.
As in the Xenodontine, genera with the apex plicate only at
marked with a star, and those with a spinous apex are marked with
dagger. The only species of Tomodon I have examined is the T.
latus, which is not the type. The position of Langaha I am not
fied about, as it may be an attenuate form of the Natricide.
NATRICID#.
This family, which I first defined from vertebral characters, is g
ally easily distinguished from the others by the characters of its
1893.) Zoology. 483
Of the HoMALoprsin& only the genera Homalopsis, Herpeton, and
Cerberus have come under my observation ; but I suppose that Hypsir-
hina and other allied Asiatic genera will be found to present the same
characters. In both genera the sulcus is bifurcate, and the hemipenis
divided.
I have been able to examine but one genus of the PsEUDASPIDIN»,
viz., Pseudaspis Cope (type Coronella cana L.), but I suspect that
Ablabes (A. rufulus) which is African, belongs here. In Pseudaspis
the sulcus is bifurcate, and the hemipenis is divided almost to the base,
quite as in the most specialized Solenogly pha.
NATRICINE.
This natural group includes fusiform and colubriform genera, and
presents great variations in the form of the hemipenis. The Asiatic
species have that organ Maa itt while in the European and American
water snakes it is simple.
Fusiform. (Hemipenis simple). Haldea; ia ss (the apex with a pair
of robust age as in Oligodon;) Virginia
Colubriform . Sulcus and hemipenis undivided. Sto ; Eutenia; Natrix;
Clonephies sie II. Sulcus and api divided. ’ Dipl ophallus, g. n.
(type. Zropidonatus piscator Schneid. ; syncranterian dentition, and no
apical papilla); Amphiesma; camel “(established on Tropidonotus vittatus
on account of the presence of a rigid papilla on the apex of each branch of the
hemipenis.)
SA Cs PRE I EEDE R A
Similar gradations in the characters of the hemipenis are to be seen
in the types of venomous snakes. Thus in the Proteroglypha this organ
is spinous to the tip, on a calyculate basis, in Hydrophis, Elaps, (surina-
mensis); Dendraspis. It is reticulate at the extremities and spinous
below, in Callophis (bivirgatus) ; ; Naja; Acanthophis; Bungarus and
Sepedon ; the apex smooth in the two genera last named. In Elaps
nigrocinctus the organ is usually smooth, with a few spines at the apex.
In Solenoglypha the genus Atractaspis is spinous to the apex, appar-
ently on alongitudinally laminate basis. In the Viperide and Crota-
lidæ the spines are on a flounced basis. The apices are calyculate in
Bitis, Clotho, and Vipera, and spinous in Cerastes. They are calycu-
late in Crotalide in Bothrops, Ancistrodon, Crotalophorus, Crotalus
and Uropsophus. In Crotalus (durissus of the Neotropical fauna),
the median spines are replaced by papillæ, in all the other genera they
are spinous.—E. D. Corr.
484 The American Naturalist.
Zoological News. Prorozoa.—Prof. August Gruber, ina
on nuclear multiplication and swarm formation in the Fresh
Rhizopoda * describes an Arcella in which milotic cell division oce
He also figures Arcella with 19 and 32 nuclei, and Leeythium hyo
with eightnuclei. These facts are adduced in evidence that amor
fresh-water Rhizopods reproduction by spore formation coexists
with simple fission. In other cases Gruber found small bodies in A
tests which at first sight might be taken for swarm spores but
are none other than parasitic Amoebæ,
Worms.—Nachtrieb and Barrows are studying the leeches
Minnesota. In a preliminary note’ Mr. Barrow states that they
found 11 species of Gnathobdellidx, 10 of Rhynchobdellide and 1 of
Branchiobdellide. It is noted that two species of Aulostoma can
distinguished by the fact that in the one the right sperm duet
under the nerve cord; in the other the left. A final report
promised. : id
Henry B. Ward reports’ the host of Nectonema agile’ Dr.M
rich found a worm, like Nectonema except that it lacked the
bands of sete, characteristic of the adult, in the thoracic cav
Palæmonetes. This when studied by Dr. Ward, proved to be af
Nectonema. |
Mottusca.—H. Suter enumerates 142 species of land moll
fluviatile, and 18 brackish water species, in New Zealand.
According to ©. Hedley, Parmacochlea fischerii collected by
“Challenger ” expedition is most nearly allied to Helicarion.
5 Ber. Naturf, Gesellsch. Freiburg, VI, 114, 1892.
ê Quarterly Bulletin, Univ. Minnesota I, p. 87, 1893.
T Proc, Amer. Acad. Arts and Sci. 1893, p. 260.
*See Amer. NAT. XXVI, 1037, 1892.
1893.] Embryology. 485
EMBRYOLOGY.’
Germ-layers of Vertebrates.’—Basilius Lwoff has extended
his work upon the germ-layers of Amphioxus’ to other vertebrates and
put forth a preliminary paper that arouses in us a lively interest in the
final detailed account that is to contain both his own discoveries and a
discussion of the literature of the subject.
In the present paper he makes statements directly contrary to some
Opinions that up to the present time we have supposed to be well-
founded in fact.
Besides Amphioxus, he has studied the formation of the germ-
layers in Petromyzon, the Axolotl, Pristiurus, Torpedo, Labrax, Julis,
Gobius, and Lacerta. He regards the Chordata as derived from a gas-
trula-like ancestor. Gastrulation is defined as the process by which
the gut is formed, and the entoblastic cells are those which form the
ut, whatever else they may produce. In Amphioxus and Verte-
brates with holoblastic eggs, cleavage results in a blastula, one-half of
which is composed of micromeres, the other of macromeres. Owing to
the continued more rapid multiplication of the micromeres, they cover
and grow around the macromeres. When the blastula is single
layered, as in Amphioxus, this results in the invagination of the macro-
meres, where it is several Jayered, as in Petromyzon and the Amphibia,
they are simply surrounded by the micromeres.
The macromeres form the gut and are therefore entoblast, while the
micromeres form the outer covering and are therefore ectoblast. But
there can be no distinction between entoblast and ectoblast until the
micromeres have surrounded the macromeres. Besides this process of
gastrulation there is a dorsal invagination that has nothing to do with
the formation of the gut, but forms the whole ectoblastic rudiment of
the chorda and the mesoblast.
The blastula of Petromyzon contains an extensive cleavage cavity,
the roof of which is composed of micromeres and the floor of macro-
meres. The former multiplying more rapidly, grow around the latter.
At the same time a new cavity is sunk into one side of the embryo,
the roof being formed of invaginated ectoblast, while the floor is com-
1This department is edited by E. A. Andrews, Johns Hopkins University.
* Biol. Centblt. 13, 1893, 40-50, 76-81.
3 See AMERICAN NATURALIST, March, 1893, p. 228.
a
486 . The American Naturalist. Diy
posed of entoblast that has not been invaginated. As in Amphioxus,
this invaginated ectoblast forms the chorda and part of the mesoblast, —
while the entoblast forms the rest of the mesoblast as well as the gut.
Nearly the same process, according to Lwoff, takes place in the |
Axolotls. But here the dorsal invagination begins, while the micro-
meres are growing over the macromeres, and the ectoblastic rudiments H
of the mesoblast never form part of the roof of the enteric cavity. n
the frog, however, Morgan‘ and Robinson and Assheton? seem to have —
found good evidence that the micromeres do not grow around the
macromeres, but that the superficial ones of the latter are themselves
gradually split up into micromeres, so if we are to accept Lwoff’s inter-
pretation of the dorsal invagination, the gastrulation in this form may —
better be described as being by delamination than by an epibolic ee
cess. Moreover, according to the apparantly accurate work of R
inson and Assheton, there is no invagination of the ectoblast in the ‘
frog. |
In Selachians and Teleosts, Lwoff believes the entire entoblast arises
from the yolk-elements, that is, the periblast. This is surprising in
view of the many statements to the contrary, especially H. V. Wilson's
work on the sea-bass. Lwoff finds the chorda and part of the meso
blast to be formed from the same elements as the nervous system. The —
entoblast also takes part in the formation of the mesoblast. 7
In Lacerta, cleavage results in a two layered germinal dise. The —
inner layer becomes the entoblast. An invagination from the outer —
_ layer forms the chorda and part of the axial mesoblast. The rest of | a
the axial and the peripheral mesoblast is of entoblastic origin. a
The process in the formation of the germ layers throughout the —
Vertebrates is practically the same as in Am phioxus, except that there i a
is no invagination to form the entoblast, and there are, therefore, 20
gastrula lips. There is but one point in the embryo that is homo-
logous in all Vertebrates, and that is the point where the ectoblastie —
invagination begins. The so-called neurenteric canal does not lie —
between the neural canal and the archenteron, but between the former i
and the chorda, and shows their intimate connection—R. P. B. y
ny ATA E ae eae Oe: a ESAE SEA eT > IS A e EN ew A ETE E oe ee eee a a E A
The Mantle of Ascidians,—In studying live specimens of
larval Phallusia, A. Kowaleysky’ at first thought with Hertwig, Sem-
* AMER. NAT., Aug., 1891, p- 753.
“Quart. Jour. Mis. Sci., 32, 1891, p. 451.
‘Bul. U. S. Fish. Com., 9, 1891, p. 209.
7 Mem. l'Acad. Imp. Sci. St. Petersburg, 7, 37, No. 10, 1892.
1893.] Embryology. 487
per, Maurice and others, that the mantle of this ascidian should be
regarded as an epidermis-like structure, a thickening of the ectoderm
with large amounts of intercellular material between the ectodermal
cells.
On studying sections of this larva, however, he obtained evidence
that mesodermal cells migrate out through the epidermis of the larva
into the thick hyalin cuticle or secreted mantle matrix, and thus supply
mesodermal cells as the fundamental cells of the mantle.
These migrating cells the author would regard as a sort of phagocytes
and imagine to have a primary function in destroying the injurious
parasites, bacteria, etc., that would easily lodge in the secreted mantle
matrix.
In the same paper there is an interesting account of the degenerative
changes that the tail of the larva undergoes when free life is given up.
After peculiar histological transformations in the notachord and mus-
cles of the tail, these cells and the epidermal cells pass into the body.
The last of the tail is drawn in by an actual hollow invagination that
forms a closed vesicle in the body. The ultimate changes in the mus-
cle cells are accompanied by the activity of clusters of phagocyte-like
mesoderm cells.
488 The American Naturalist.
PROCEEDINGS OF SCIENTIFIC SOCIETIES,
Boston Society of Natural History.—April 5.—The
lowing paper was read by Professor George Lincoln Goodale:
some aspects of Australian vegetation.” It was illustrated
stereopticon views.
SAMUEL HENSHAW, Secreto
New York Academy of Sciences. Biological Sectic
March 13, 1893.—Professor T. D. Quackenbos, in a paper on the
ling of Lake Sunapee, distinguishes, in this, a fourth variety of
England Charr, demonstrating that the present abundance of
Salvelinus is accounted for not from its introduction and 1
increase, but from destruction of inimical forms within recent
which has given a greater available food supply. Professor
Huntington, on “Anomalies of Pectoralis major and minor,” ref
to the value of these as often presenting reversions. He emph
the evolutionary tendency in man to proximalization of the poir
attachment of the shoulder muscle group, referred to cleavage
tions in anterior portion of brachiocephalic sheet, and compared |
with ontogenetic characters in Anthropoids. Human anomalies
Romeyn B. Hough, for the State Forestry exhibit at the
Exposition, as mentioned at the meeting of October 15, 1892.
Pany’s water, from a sample taken from a hydrant at the c
New York Avenue and Cliff Street, and found 185 individual b
per cubic centimetre, The temperature of the water was 34° Far,
Previously stated, the New York City Croton water contains
12,000 per cubic centimetere, :
1893.] Proceedings of Scientifie Societies. 489
Mr. Arthur Hollick presented a piece of ferruginous sandstone, con-
taining impressions of dicotyledonous leaves. The specimen resembles
those found in the Drift, at Tottenville and Princes Bay, which are
known to be derived from the Cretaceous clays. The specimen in
question, however, was found as a Drift rock on the Serpentine hills,
to the north of the Cretaceous area. It is possible that it may have
been brought to the place where it was found by human agency, but
if not, its occurrence there is more or less of a problem which future
discoveries may solve. It is desired, at present, merely to place the
fact upon record.
The Biological Society of Washington.—March 11.—The
following communications were made: Dr. Frank Baker, “ Recent
Discoveries in the Nervous System ;” Mr. Vernon Bailey, “ The Bur-
row of the Five-toed Kangaroo-Rat;” Mr. E. M. Hasbrouck, “The
Breeding of the Bald Eagle near Mount Vernon ”—with exhibition of
Eggs
March 25.—The following communications were made: Mr. L. M.
McCormick, “A Hybrid between Pyranga rubra and Pyranga erythro-
melas;” Prof. E. W. Doran, “ Development of the Intestine of Tad-
poles ;” Dr. Theobald Smith, “ The Bacteriology of Potomac Water
and its Bearing on Sanitary Problems ;” Mr. B. T. Galloway, “ Experi-
ments in Preventing Rusts Affecting Cereals.”
Freperic A. Lucas, Secretary.
Anthropological Society of Washington.—Feb. 21.—The
following papers were read: “ The Foundation of the Zuni Cult,” Mrs.
Matilda Coxe Stevenson ; “ Dual Civic Functions: a study in the evo-
lution of institutions,” Miss Katie Foote; “ Early Man in the Missis-
sippi Valley,” Mr. Thomas Wilson.
Wesron Fut, Sec’y Board of Managers.
SCIENTIFIC NEWS.
At the meeting of the Geographical Club in the Hall of the Acad-
emy of Natural Sciences of Philadelphia, it was announced that the
result of the balloting among the members of the club upon the ques-
tion of active co-operation in the proposed new expedition to the Arctic
regions under the lead of Lieut. R. E. Peary, stood in favor of the
project.
490 The American Naturalist.
The proposed expedition will involve an expenditure of from
$20,000 to $25,000. The expenditure covers two northern vo
The first to land Peary and his party, and the second to bring i
upon the completion of its work. An auxiliary summer expe di
to be organized to take charge of the ship on her second voya
movements of the vessel to be under the control of the Geog
Club
The design of the expedition is the complete determination a
delineation of the detached land masses lying to the north of main
Greenland. The second object will be the determination of th
unknown portion of the east coast of Greenland, extending from n le
pendence Bay to Cape Bismarck, the most northern point sigt by
the German expedition in 1870. The third object will be the
mination of the transverse profile of the Greenland inland ice ca
Cape Bismarck to Inglefield Gulf.
Other aims of the expedition will be the complete detailed st
Whale Sound and Inglefield Gulf, with studies and measurem
the glaciers in that region, and their parent ice cap ; a continua
the study of the little tribe of Arctic Highlanders. As comp
lections as may be practical will be made of the natural objects
region, and more or less extended meteorological and tidal ok
tions carried on.
The amount of funds to be contributed by the Geographical C
will be from $8,000 to 10,000.
Botanists will hear with regret of the death at Brighton, Jan. I
Dr. Benjamin Carrington, the highest authority on British E
The State of Iowa has assumed the expense of the publication sie
papers and proceedings of its Academy of Sciences, the vt
having made an appropriation for that purpose at its last mee
Professor A. M. Miller has been appointed to take charge
department of Geology which has just been established in the
College of Kentucky, at Lexington.
Mr. John Eyerman has presented to the Princeton Museum
_ easts of Rhytina gigas Linn, after the originals in the British M
‘The number of casts is 20, that of the cranium, measuring 4
_ -M, Johnston-Lavis has been appointed Professor of Valea
the University of Naples. A similar chair existed at Cat
was abolished at the death of its oceupant, M, Silvestri.
1893.] Scientific News. 491
Mr. G. W. Lichtenthaler, one of America’s most eminent con-
chologists, died in San Francisco. Feb. 20, 1893. For 20 years he had
been a traveler and collector in almost every sea and country. During
this time he brought together a large collection of specimens, which he
bequeathed to the Illinois Wesleyan University, in his home city,
Bloomington, Ill.
The collection embraces mainly shells, sea algae and ferns. His shells
have been estimated conservatively at from six to ten thousand species,
including thousands of duplicates. They embrace the Hemphill col-
lection which he purchased, a large collection from the Hawaiian
Islands, which he visited often; many from the Micronesia, Australia,
Japan, the coasts of Europe, and America, and from every part of the
United States.
The sea algae contain about a thousand species, all mounted in
books and identified, and a large number of duplicates. They were
collected from both coasts of America, from north to south, a large
number from England, and many from other countries.
There are about 500 species of ferns, including the many duplicates.
They represent a complete collection of the ferns of the Hawaiian
Islands, and almost a complete collection of those of the United
States, besides many from Europe, Asia, Australia, South America,
te.
He has also brought together many mosses and lichens, the number
of which cannot be given with precison, but there are several hundred
species.
The Marine Biological Laboratory of the Johns Hopkins University
will be located this Summer at Port Henderson, Kingston Harbor, in
the island of Jamaica.
Dr. R. P. Bigelow, present holder of the Bruce Fellowship with
Messers Lamb, Siegirfuss and Lefevre left Baltimore, April 20, by a
fruit steamer for Port Antonio, whence they will go to Kingston to
open the Laboratory. Others with the Director, Professor W. K.
Brooks expect to go down there later.
CAMBRIDGE, Mass., April 15, 1893.
Tue Faculty of the Museum or Comparative Zoooey will receive
applications from candidates desiring to oecupy the table at the
NAPLES ZOOLOGICAL Station, which has been placed at its disposal
“from October 1, 1893.
The applicant must be (or have been recently) a student or instructor
492 The American Naturalist. [May
at some American University, preferably a person who has taken the
degree of Ph.D. or S.D.; he must have published some creditable
original investigation, and should be recommended as an able investi- —
gator by the professor under whom he has studied. :
Applicants will please forward to the undersigned, before May 10 —
their recommendations and a statement of their qualifications and of —
the subject to which they hope to devote themselves.
In order that the Faculty may make the most satisfactory disposi- |
tion of the table during the whole year, the applicants are requested —
` to state the length of time they desire to remain at Naples, and also —
the earliest and latest dates within which they can avail themselves of —
the appointment. :
The Faculty will, at its meeting in May, nominate to the Corporation
of Harvard College for approval the incumbent or incumbents for the
year 1893-1894. |
ALEXANDER AGASSIZ,
Director, —
The North American Review. Table of Contents for a
April, 1893.—Charges at the World’s Fair—by Director-General —
Davis. Brain Surgery—Dr. William A. Hammond. Shipbuilding —
Here and Abroad—Naval Constructor Hichborn, U.S.N. Good and —
Bad Mothers—Mrs. Amelia E. Barr. How Shall the Pension List be —
Revised ?—by Representative Wilson, of Missouri, Chairman of the —
Committee on Pensions ; Gen. S. S. Burdett, Past Commander-in-Chief —
of the Grand Army of the Republic, and Col W. C. Church, editor of
the “Army and Navy Journal.” Art of Mystery in Fiction—George —
Manville Fenn. The Interior of the Earth—George F. Becker, of the
—1. The Currency and the Democratic Party—by the Hon. R. P.
Bland, Chairman of the Committee on Coinage, etc. ; 2. The Brussels
Conference Reviewed—by the Hon. Charles Foster, Ex-Secretary of
the Treasury. Notes and Comments.—*“ By Women for Women ”—
Lillian A. Mercur. Do the Fittest Survive?—W. A. Croffut. The
Abuse of Secrecy—Clara Dixon Davidson. English Poor Law Reform
—Edward Porritt.
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THE
AMERICAN NATURALIST
Vor. XXVI. June, 1893. 318
RECENT RESEARCHES UPON THE SUCCESSION OF
THE TEETH IN MAMMALS.
By Henry FAIRFIELD OSBORN.
While American paleontologists have been making rapid
advances in the phylogenetic history of mammalian teeth,
the English have made notable additions to our comparative
anatomical knowledge, and the Germans to the embryogene-
sis. The recent studies of Kükenthal, Röse and Taeker in
the discovery of the complete double or milk dentition in the
Marsupials, and in the discussion of its relation to that of the
reptiles, also in the ontogenesis of the crowns of the teeth
among the Cetaceans, Edentates, Primates and Ungulates are -
of the greatest interest and importance. They involve a
complete revolution in our ideas as to the interpretation of the
dentition in the three orders first mentioned above.
The latter authors are perhaps inclined to lay too great
stress upon the testimony of ontogenesis in the order of
appearance and consequent homologies of the mammalian
cusps, and Rése’s work is to a certain extent rendered less
useful by the fact that he has substituted the conclusions of
ontogeny for those based upon the more certain foundations of `
Phylogeny. It isa subject of congratulation, however, that
ese authors, who constitute the rising school in Germany,
Tecognize the value of the paleontological work done in this
33
494 The American Naturalist. [June,
country, and are ready to join hands in pushing forward these
investigations upon a joint basis.
KUKENTHAL’S RESEARCHES.
Kikenthal’s studies upon toothed whales were partly
directed upon the theory of Weber, Julin and others that
these animals were formerly heterodont. In the embryo of
the Porpoise (Phocena communis,) he found traces of a
heterodont condition quite sharply marked. There were
twenty-five teeth in each half of the jaw, and the posterior
seven were found to have two or three cusps. Having thus
supported the opinion that the homodont toothed whales were
primitively heterodont, Kiikenthal has also proved that they
are diphyodont, and that the dentition of the toothed whales
is a true milk dentition, while the second or permanent denti-
tion is represented by rudiments which display a distinct
crown of enamel and even the enamel pulp, yet does not
reach the surface. i
In the whalebone whales in which germs of teeth had been
found in the first third of foetal life, Kükenthal does not con-
firm the opinion of Weber that the dentition is heterodont.
He denies that the posterior teeth are more complex than the
nine anterior teeth, and holds that throughout they are simply
conical, excepting in cases where two external teeth are fused
together. This fusion of teeth does not follow any definite
rule; in some cases it occurs in the anterior nine teeth. The
course of embryonic development shows that these fused teeth
represent an original condition, and in the opinion of the
author, are to be regarded as molars. This conclusion was
reached by the comparison of younger and older embryos, the
number of fused teeth constantly diminishing in the latter.
Kiikenthal advances the hypothesis that this was the method
by which numerous homodont teeth arose from a small num-
ber of heterodont, namely, by the splitting apart of cusps.
This hypothesis he promises to support by paleontological a A
evidence. He also shows that these embryonic teeth in the
whalebone whales also represent the first or milk dentition;
and that the rudiments of a second series of teeth develop — 4
1893. Succession of the Teeth in Mammals. 495
beneath them. Kükenthal refers these transformations in the
dentition to natural selection, terminating with diminished
calcification connected with the advantage of diminished spe-
cific gravity.
His general conclusions are, that all the earliest mammals
were diphyodont. This is based upon his discovery of suc-
cessional teeth in Marsupials, Edentates, Odontocetes and
Mystacocetes. The monophyodont and homodont condition
of many mammals, such as the toothed whales, he believes
has been secondarily acquired. Within the higher members
of the mammalian class the second dentition is developed
progressively, both as regards form and funetion; while in
` the lower divisions, the first or milk dentition is predominant.
“Tn the rudimentary stage both dentitions are of equal value.
Embryology gives us no support for the often expressed asser-
tion, that one of the two dental rudiments has arisen in
dependence upon the other. They are both sisters, whose
mother is the simple invagination in the jaw, which we term
the dental fold, (Zahnleiste.)” He continues, that there are no
absolute differences between mammalian and reptilian teeth ;
that not one of the characters of the mammalian teeth is
perfectly constant, and that the derivation of the dentition of
mammals from that of the reptiles does not appear to be too
hazardous. Of the several series of teeth which are found in
reptiles, only two persist in the mammals.
From this I would dissent in part. The three differences
between the mammalian and the reptilian teeth are shown in
the capacity for the multiplication of cusps upon the crown,
in the division of the fang whenever the crown becomes mul-
ticuspid, in the acquisition of the cingulum. The frequent
Succession of teeth in the reptiles, may be the cause of the
non-progression of reptilian as compared with mammalian
teeth. In the reptilia among the Theromorpha, we find true
triconodont crowns, as for example in Galesaurus, and a
heterodont dentition which closely imitates that of the
Mammals; but the class differences appear in the fact
that in the mammalia a development of lateral cusps upon
the protocone, and the stages from the protodont toward the
496 The American Naturalist. [June,
triconodont type, are marked step by step by the division of
the fang. This law was advanced hypothetically by Cope and
Wortman, and I regard it as absolutely proven by the evi-
dence I have adduced in the study of Dromotherium, Microcon-
odon and Amphilestes.
The hypothesis of Kiikenthal and Rose that the numerous
single pointed or homodont teeth of the whales, have arisen
by the splitting up of the three cusps of a triconodont crown
is an ingenious one, upon which paleontology at present
throws no favorable light. Amphilestes of the middle Triassic
with its seven triconodont molars, might by such a splitting
process, furnish twenty-one homodont teeth; nevertheless,
this seems to me highly improbable; while the converse
hypothesis suggested by Kiikenthal and developed by Rose,
that multicuspid crowns have originated by the fusion of
single cusps, is capable of direct disproof by paleontological
evidence.
The multiplication of teeth accompanying the elongation of
the jaw in Cetacea, can be much more simply explained by
the supposition that the dental fold was carried backward,
and gave rise to new teeth caps at definite intervals. I may
add that the rapid reduction of the molars in the Mesozoic
period from behind forward, which reduced their number in
the Triconodontide from seven to four, between the middle
and upper Jurassic periods, is against the supposition that
the Amphilestes molar, for example, furnished the material for
the multiple Cetacean series.
Kikenthal’s researches upon the dentition of the opossum,
published in 1891, mark another great step in advance in
our knowledge of the dentition of the mammals. We may
refer to the later researches of Röse for details, and simply
quote one passage from Kiikenthal:
“The permanent dentition of the Marsupials, belongs te the
first series or milk dentition. Rudiments of the second den-
tition are actually present in an embryonic condition, but
with the exception of the third premolar, they do not cut
the gum.” . Again, the two first so-called true molars, of the
*Anatomischer Anzeiger, Nos. 23, 24.
1893.] Succession of the Teeth in Mammals. 497
upper jaw, and the first three similar teeth of the lower jaw,
also belong to the first dentition and have rudimentary
successors. Therefore, excepting the last upper and lower
molars which appear at a late stage of development, neither
dentition of Didelphys belongs to the so-called permanent
series. This opinion has been somewhat emended by Röse.
Rosek’s RESEARCHES.
MARSUPIALS. Dr. Rose gives the following summary of his
principal results. “ When we combine the results of the fore-
going researches, we find that the development of the teeth in
the Marsupials follows exactly the same principle as that of
man and the other mammalia. The first matrix is indicated
by the dental fold, i.e. a part of the epithelium of the jaw
which grows into the mesoderm aided by an extensive fold.
Upon this dental fold are formed the dental caps which belong
to the first series; in the case of Didelphys, the incisors, the
canine, two premolars and the first molar. These dental caps
are then immediately constricted off from the dental fold, and
this ridge grows both inwards below the dental caps, and
backwards behind the molar above mentioned. The posterior
molars arise in exactly the same manner as I have described ;
inthe case of man, through lateral extension of the dental
e.
“ While, however, in the case of man, the permanent ridge
extends beneath the ten anterior teeth constituting the milk
series and gives origin to ten permanent teeth, which cause
their predecessors to be absorbed, in the Marsupials only the last
premolar of the adult arises from the dental ridge of the per-
manent teeth. It is nevertheless, more than probable that the
. Outer (5th) incisors of Perameles, as well as of Macropus and
Phalangista, are built up from the permanent ridge, i.e. they
belong to the second dental series. The last upper premolar
belonging to the second series in some cases simply pushes its
. Way into a gap in the first series without causing the usual
absorption of the first tooth in this row. This type we find to
be shown in Didelphys, in Perameles, in Phalangista cookii, as
498 The American Naturalist. [June,
well as in Macropus. In other cases the last premolar of the
first series is absorbed, and in its place enters the premolar of
the second series; this more primitive type is found in an
undetermined species of the genus Phalangista ; also in Macro-
pus lugens and M. gigantius; also, according to the figures of
Oldfield Thomas, in Phascogale, and in the fossil Triacantho-
don serrula.
“ Further researches are necessary to determine which type is
the most common among the Marsupials. It also remains to
ascertain through sections and models, whether the last incisor
(fifth or lateral, i°) of the upper jaw really arises from the sec-
ond series, and in what species this occurs. Possibly in some
Marsupials other teeth also rise from the second series.
Whether this, however, is the case or not, the principal theory
of Marsupial dentition is not thereby affected. It is certainly
well established by my researches, as well as by those of
Kikenthal, that the teeth of the Marsupials, with the excep-
tion of the last premolar and probably of the last superior
incisor of some species, belong to the first series, and are
analogous with the milk teeth of man and other mammalia.
“ This shows that in the entire vertebrate series the principle
holds good, that by the better development of the single tooth,
the frequent succession of teeth of the Selachian type, is
gradually limited. The Marsupials, however, in the reduction
of the multiple succession of the reptilian-like ancestors of
existing mammals, have together gone past the limit, and
have wandered into a cul de sac of evolution out of which there
was no return. Upon this rests the remarkable constancy
of this order, from the Mesozoic period to the present time.
The reduction of the former multiple succession of teeth of the
vetrebates to a single series of teeth, appears therefore, only to
be of service when the teeth at the same time attain a perma- i
nent growth. This stage was only attained among the
Marsupials by Phascolomys.”
In the same paper the author restates his theory of the
origin of premolars and molars by the fusion of several single -
teeth, and is led far astray from the actual conditions which we
observe in such development. Originally, he says, the pre-
1893.] Succession of the Teeth in Mammals. 499
molars, like the molars, embraced three single cusps, which were
arranged in the triconodont type; in Triacanthodon the pre-
molars are altogether formed like the molars. (This observa-
tionis an error. The only Mesozoic mammal in which the
premolars are even approximately similar to the molars, is
Phascolotherium.) After the original three cusps of the pre-
molars had been arranged in the triconodont order, he supposes,
arose the degeneration of the forward cusp, leaving a tooth type
which’ we observe especially among the Carnivora. (The
cusps left here are simply the protocone and talon.)
In an earlier paper upon human dentition, the author is
also led astray by ontogeny to false conclusions as to phylo-
geny, and at the conclusion of his extremely interesting paper
upon the embryogenesis of the human dentition, he says:
“The typical form of upper molars in man is unquestionably
the four cusped type; while the typical form in the lower
molars is the five cusped type.”
EDENTATES. Dr. Röse has also contributed interesting obser-
vations upon the rudimentary development of teeth among
the Edentata. He quotes Max Weber to the effect, that the
reduction of the teeth in Manis is so absolute, that not a single
rudiment remains, and that there is, so to speak, no place left
for the layer either of dentine or enamel. He then goes on to
say: “ After, in my earlier studies, I recognized the morpho-
logical importance of the dental fold, and finding it ended as
a last rudiment of an earlier dentition both in the birds and
turtles, I did not doubt that also in Manis, at least the first
stages of a dental ridge must be present in the early embryonic
development.” With material received from Professor Max
Weber himself, Dr. Röse demonstrates the beginnings of the
dental fold both in the upper and lower jaws of Manis ; and
_ in the lower jaw on both sides he finds even an unmistakable
rudiment of the tooth layer, in the form of a swollen portion
of the common dental fold, i. e. an abortive tooth cap. Such
Tesults were obtained in Manis javanica.
The forms investigated were the Nine-banded Armadillo
(Dasypus novemcinctus), Dasypus hybridus, the Pangolin (Manis
javanica), and the {Anteater (Myrmecophaga.) In the Nine-
500 The American Naturalist. eer
banded Armadillo most careful researches were made upon
the dental ridge in a series of 420 sections, leading to a dis-
covery which was confirmed in the other species, that the
dentition of the Edentata has arisen by degeneration from the
dentition of a more highly organized mammalian type. This
result is directly contradictory to the hypothesis of Oldfield
Thomas, that the Edentata were sharply marked off as Para-
theria by the exceptionally simple features of their dentition
among other characters.
Dr. Röse continues, that the enamel of the teeth has so far
retrogressed, that only an upper capping of the enamel layer
is formed, which lies directly upon the dentine. “ In all cases
in which we have instituted exact microscopical researches, in
three species of Dasypus, and in Orycteropus, it is proved that
invariably in the Edentata the typical two dentitions of the
mammalia are exhibited in the embryonic stages. In every
case as the tooth is constricted off, the dental ridge grows
further backwards, as the basis of the successional tooth, as it
has been observed in the development of the teeth in man and
in the Opossum ; and as Kükenthal observes, from a morpho-
logical standpoint it is of no moment whether this replacing
ridge gives rise to teeth or not. ”
Dr. Röse anticipates that the Bradypodide will also prove
to be diphyodont.
This author further shows that there are proofs of heterodonty
which weigh against the opinions of Parker and Thomas, that
the Edentata stand entirely aside from the other mammalia.
In the early stages of development, a continuous dental ridge
is found in the whole jaw, and in the anterior part of the jaw
two rudimentary teeth are observed which must be regardet
as incisors. According to Rheinhardt, the number of rudi-
mentary incisors in the Nine-banded Armadillo is still greater
(four). The lateral tooth in Bradypus may be either an
incisor or canine. Dr. Röse agrees with Tomes and Osborn
that the canine should be considered as the most anterior pre
molar. Another question arises as to whether the posterior
teeth of the Edentata are to be regarded as molars or 9%
divided into molars and premolars. As Kiikenthal be
1893.] Succession of the Teeth in Mammals. 501
observed in the whalebone whale, we also find in young
embryos of the Edentatesa double pointed or biconodont tooth,
which later divides secondarily into its single pointed compo-
nents. The author considers it doubtful whether this bicono-
dont type is primitive, or as seems to him more probable, is
derived by degeneration from a triconodont type !
As regards the bearing of paleontology and comparative
anatomy upon these facts, we find rudimentary incisors in the
living Dasypus setosus and in the fossil Chlamydotherium.
The latter shows the fusion of two conical points. The author
refers also to Glyptodon, which has prismatic back teeth with
a very evident triconodont type. He concludes by saying
that these circumstances afford a very marked support for the
opinion, that the present biconodont teeth of these Edentata,
at least the posterior members of the series, have arisen by
reduction of a typical triconodont type. If this is confirmed
by further observation, it will be a fact of the very greatest
importance in the phylogeny of the mammalia. For the tri-
conodont type has been shown to prevail in the upper Triassic
and lower Jurassic periods. There was unquestionably a vast
period in the evolution of mammalia in which a triconodont
molar was the only type.
SAUROPSIDA. In the conclusion of his work upon the
development of the teeth in the crocodile, he reaches the fol-
lowing result: “The cause of the existence of the thecodont
(fangless) teeth is to be found in the continuous growth of the
epithelial sheath of Hertwig. . . . -+ . . The fant-
tional tooth of the crocodile is altogether analogous with a
rooted mammalian tooth, the growth of which is not yet com-
plete. The first embryonic tooth series of the crocodile, on,
the other hand, exhibits a development which we hitherto
have only observed in the Selachia and Urodela through the
labors of Hertwig. The theory of Hertwig as to the genesis
of the mouth skeleton out of the cement sockets of tooth
structure, has received an unexpected and weighty support by
our researches, ”
The types examined by the author, included Sterna Wil-
sonii, the Ostrich, Struthio camelus and the turtle, Chelone
502 The American Naturalist. [June,
midas. He anticipated from his studies upon the tooth
development of the reptiles, that only traces of the dental fold
would be found, and if any really rudimentary dentine teeth
were found, they could only represent those primitive
Selachian-like teeth which constitute the first series of the
crocodile. In the crocodile as above noted, a first dental
layer exhibits itself altogether similar to
the placoid scales and the first teeth of the
Selachians in the form of free papille upon the surface
epithelium of the jaw.
TAEKER’s RESEARCHES.
Dr. Taeker of the Veterinary Institute of Dorpat, has com-
pleted a most interesting series of studies upon the embryonic
form of the teeth in the Ungulates.
His technical methods were an improvement on those
introduced by Klever, and his material included embryos of
the horse as a type of the most complex form of perrissodactyl,
of the pig as a modern bunodont, and of Selenodonts he
selected the embryos of the Elk, Deer, Ox and Sheep, and of
greater rarity, an embryo of one of the group of Tragulids.
His conclusions are summed up as follows: :
1. As a result of my investigations I find that both the Sui-
dæ with rounded or bunodont cusps, and the Ruminants with
their crescentic or selenodont cusps, arise from a similar initial
bunodont stage; that is,-all the highly complex forms of
modern cusps spring from the simple ancestral hillock in
the embryonic stages.
« 2. The order of differentiation soon follows, in which the
Separate cones and conids (cusps of the lower jaw) are trans-
formed into pyramids in the case of the pig, and into cres-
cents in the case of the ruminants.
3. The transformation of the cones is not effected simultan-
eously, but successively. In the upper teeth it does not begin
with the ‘protocone (antero-internal cusp which is first
developed in the paleontological history), but with the
external cusps, the paracone and metacone.
1893.] Succession of the Teeth in Mammals. 503
Order of Appearance of the Cusps. The author observes
that while in the lower jaw the order and appearance of the
cusps is the same in the embryo as in the paleontological his-
tory, in the upper jaw this ontogenesis is no longer parallel
with phylogenesis, (as regards the upper teeth, this exception
is confirmed by Röse). In fact the external cusps not only
appear before the internal cusp, which paleontology shows
to be more primitive but they assume the crescentic form
earlier. In other words, their development is accelerated.
The development of the premolars is also traced and
it is interesting to recall the fact that in the paleontological
history, the order of evolution of the cusps of the premolars
is not the same as that of the molars. Taeker’s results there-
fore show a parallelism between Ontogenesis and Phylogene-
sis, in that he proves that the embryonic cusp order in the
‘premolars is different from that in the molars, and is approxi-
mately similar to that in the ancestral history as recently
worked out by Scott.
Upon the whole this parallelism between embryogenesis and
palingenesis is most striking, and I think we can explain the
exceptions which Taeker and Röse have shown to occur in the
upper teeth both in the Primates and Ungulates, in the follow-
ing manner: In the most primitive types of trituberculates,
the protocone was the most prominent cusp in both jaws.
This is seen in all the known Triassic and Jurassic trituber-
culates without exception. During the Cretaceous period a
change took place, in which the upper molars rapidly diverged
in pattern from the lower molars. The lower were more con-
servative, retaining the trigonid or triangle in its primitive
proportions. The upper molars, on the other hand, must have
undergone a marked change, consisting mainly in the depres-
sion of the protocone below or to the level of the paracone and
metacone, as seen in the primitive Carnivores, Creodonts,
and Insectivores of the Puerco period. Thus we find both in
the Ungulates and sub-Ungulates or early Primates, that in
the upper molars the outer two cusps are slightly more ele-
vated and decidedly more progressive in the acquisition of
new forms than the older internal protocone. For example
504 The American Naturalist. [June,
the outer cusps often became crescentic, while the inner cusp
remained rounded. And it was therefore during all the lower
Eocene, that the external cusps surpassed the internal cusps in
progressive development.
The bearing of these facts is this: in Embryogenesis, we
are dealing with repetition of ancestral history; we should
not expect, however, that this repetition would invariably
extend back of the form characteristic of the Eocene period.
As a matter of fact, the upper molars repeat their Eocene but
not their Mésozoic form. The lower molars also repeat the
Eocene form, and this, as explained above, owing to the con-
servative proportions of the cusps, is also the Mesozoic form, the
protocone still being the most prominent cusp in the crown.
While, perhaps, not thoroughly satisfactory, there is a great
deal of probability that the discrepancy between the embryonic
and phyletic order in the upper molars, is due to these differ-
ences in their phyletic history.
In the accompanying table I havesummed up the phylo-
genetic order observed by Cope and myself, and the ontogene-
tic order observed by Röse and Tacker.
THE Fusion THEORY or Cusp ORIGIN.
As we have seen, the fusion theory was first proposed by
Kükenthal; it was afterward independently advanced by
Rose. We find in addition to the grounds given above, one
very strong argument against this theory derived from
Paleontology, is in the law of molar evolution, namely, that
the cusps appear at or near the apex of the crown, and
development takes place from above downward. Thus, so far
as we can judge from Dromotherium and Microconodon, the
lateral cusps first appear on the sides of the protocone, and
much later, the fang subdivides; the lateral cusps are at the
outset very much smaller than the medium cusp, and it is
only after a long course of evolution that they attain the same
size. i
Now, if the fusion theory were correct, and the triconodont
crown, for example, were constituted by the fusion of three —
Succession of the Teeth in Mammals. 505
1893.]
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506 The American Naturalist. [June,
distinct cones, these cones should certainly appear sub-equal
from the first, and they should present a division extending to
the base and indicated in the fang. From the fact that
Kükenthal does not mention the above genera, we infer that
his attention has not been called to them. In one of his
latest addresses he speaks as if the multituberculates were the
only Triassic mammals. We know that the trituberculates
were as old or older than the multituberculates, and reasoning
by analogy, I have attempted to show elsewhere that the
multituberculates will be found to have a trituberculate
origin.
As I have lately criticised this theory in some detail in the
“Anatomischer Anzeiger,” I here refer to the well-known
evidence which renders the Kiikenthal-Rése hypothesis both
superfluous and untenable; when we find in the series such
as is represented by the genera Dromotherium, Microconodon,
Spalacotherium, Amphit , Miacis, exhibiting in the lower
jaw all the stages between the sectorial and bunodont crown,
and when in the early Ungulates we are able to trace one by
one the successive additions of cusps to the bunodont molar,
often in the center of the crown, it certainly is not necessary to
attempt to establish any new hypothesis as to cusp origin.
SUMMARY OF RESULTS.
The general bearing of these researches upon the stem forms
of the pro-mammalia is extremely important. To summarize,
it is rendered probable that the earliest forms of mammals
were homodont and had two or more series of successional
teeth. Then within the mammalian stem the teeth were
differentiated, and there arose a great heterodont group with
teeth at least of three kinds—incisors, premolars and molars,
all successional. From the most anterior premolar arose the
canine. Then came the division between the Marsupials and
the Placentals, the former tending to suppress the develop
ment of the second series of teeth, the latter retaining the
second series as far back as the first molar. 3
We can see an obvious advantage in the line of succession
P:
1893.] Succession of the Teeth in Mammals. 507
being drawn at the first molar, for upon the molars rested the
necessity of complex development, and such development was
best effected in permanent crowns.
1. All the so-called “milk molars” plus the so-called “true
molars” constitute the first series. Beneath one or more of the
“true molars” in lower mammals are rudiments of a second
series. The second series consists therefore of these sub-molar
rudiments plus the successional or permanent premolars,
incisors and canines.
2. In the stem Marsupials the entire first series persisted
and became mainly permanent (non-deciduous); the second
series became rudimentary and non-successional with the
exception of the fourth upper and lower premolars and
possibly one or two other teeth which either replaced or were
intercalated between members of the first
series. One or more premolars were suppressed and one
more molar retained than typical in the Placentals. Thus is
explained the apparently atypical dental formula of Marsupials.
3. In the stem heterodont Placentals (excepting the Cetacea
and Edentata) the entire first series persisted and all the
incisors, canines and premolars remained deciduous. The
successional second series persisted as far back as the first
molar. r
4. In the stem Cetacea the entire first series persisted and
the second series became rudimentary and non-successional.
The tooth form changed from a heterodont to a homodont
type.
5. In the stem Edentates, which also transformed from the
heterodont to the homodont type, the first series became rudi-
mentary and the second series persisted in the succession even
behind the region of the first molar.
Finally, there is evidence that a primitive succes-
Sion inthé region of the molarteeth, lost
1The law of molar evolution is that complication is most rapid in teeth which are
longest in use. Thus the first molar is the most progressive tooth of the true molar
series and the last premolar is the most progressive of the premolar series. The
apparent exception that the third milk premolar is always an advance type of the third
permanent premolar is = by the fact that the milk premolars are formed to
assume the molar functio
508
The American Naturalist. [June,
in the Marsupials andin the Placentals, was
more or less fully retained in the Cetacea and Edentates.
Biological Department, Columbia College, April 10, 1893.
Kükenthal.
Röse.
Taeker.
BIBLIOGRAPHY.
“Das Gebiss von Didelphys, ” Anatomischer
Anzeiger, 1891, Nr. 23, 24.
“ Einige Bemerkungen über die Säugethierbe-
zahnung ” Anat. Anzeiger, 1891, Nr. 13.
“Uber den Ursprung und die Entwickelung
der Säugetierzähne, ” Jena, 1892.
“ Ueber die Enstehung und Entwickelung des
Säugetierstammes, ” July 15, 1892, Biolo-
gisches Centralblatt. '
“ Uber die Entstehung und Formabänderungen
der Menschlichen Molaren.” Anat. Anz.,
1892, Nr. 13, 14.
“Ueber die Zahnentwickelung der Krokodile”
Verh. d. Anat. Gesellschaft, June, 1892.
“ Beiträge zur Zahnentwickelung der Edenta-
ten” Anat. Anz., 1892, Nr. 16, 17.
“Ueber die Zahnentwickelung der Beutel
tiere” Anat. Anz., 1892, Nr. 19, 20.
“ Ueber rudimentire Zahnanlagen der Gattung
Manis.” Anat. Anz., Nr. 19, 20.
“Ueber die Zahnleiste und die Eischwiele der
Sauropsiden.” Anat. Anz., 1892, Nr. 23, 24
“Zur Phylogenie des Siiugetiergebisses. ” Biol.
Centralb. Nov., 1892. :
“ Zur Kenntniss der Odontogenese bei Ungula-
ten.” Inaug. dissert, Dorpat, 1892.
1893.] Symbiosis and Mutualism. 509
SYMBIOSIS AND MUTUALISM.:!
By Roscor Pounp.
Symbiosis and mutualism, in the vegetable kingdom at least,
are phenomena accompanying parasitism. Parasites have
various effects upon their hosts, according to the nature of the
parasite, its mode of life and method of attack. In some cases
the host is quickly killed and the parasite becomes a sort of
saprophyte upon the remains. In others the host lives longer
or is only partially affected. In still others the host lives on
side by side with the parasite indefinitely. A further develop-
ment is attained in cases where the parasite and host not only
live together, but are mutually beneficial, and, perhaps, even,
in extreme cases, inter-dependent. To the first phenomenon
—namely, the living together of parasite and host—DeBary,
in 1869, in a work entitled Die Erscheinung der Symbiose, gave
the name of Symbiosis. The latter phenomenon—i. e., mutual
assistance or inter-dependence of parasite and host—was named
mutualism in 1873 by Van Beneden in his “Animal Parasites
and Messmates.” Symbiosis in the strict sense and mutualism
are often confounded, that is, the term symbiosis is often used
to mean mutualism as such; but, in strictness, while mutual-
ism, in the case of plants, can only exist with symbiosis, in the
larger proportion of cases of symbiosis there is no mutualism. ©
At the outset it should be noted that the mutualism of which
we are here speaking is mutualism of parasite and host—not
mutualism of independent organisms. Of the latter, we have
many examples in the animal kingdom, and, indeed, the
human race furnishes examples of it. There is a sort of
mutualism between man and wheat, for example. Wheat is
cultivated by man and enabled to grow in quantities, and in
localities which, under ordinary conditions, would be impos-
sible. It gains this partial exemption from the struggle for
existence only at the expense of an immense number of indi-
1Read before the Botanical Seminar of the University of Nebraska, December 17,
1892. :
510 The American Naturalist. [June,
viduals sacrificed, but it is, nevertheless, a great advantage
which it gains. This may be called mutualism. But there is
a case of mutualism of plant and animal much more closely
resembling the mutualism of parasite and host in the vege-
table world. The mutual inter-dependence of Yucca and a
moth of the genus Pronuba, is probably the most unique and
interesting case of mutualism to be found anywhere. This is
well described by Mr. Webber in the AMERICAN NATURALIST
for September, 1892. In this case the plant and the moth, if
not strictly sustaining the relation of parasite and host, live
together for a long period, and it approaches much closer to
mutualism as found between vegetable organisms than phe-
nomena like entomophily where animals and plants are
mutually beneficial, without any approach to symbiosis. In
the vegetable kingdom, mutualism is a relation of mutual
benefit between organisms living together as parasite and host.’
The most conspicuous and earliest observed instance of
mutualism in the vegetable kingdom is the relation of the
Lichen fungi to their gonidia or algal hosts. The relation of
the lichen thallus to its contained gonidia was, at one time,
the subject of no little ridicule, not only because its discovery
overturned many established ideas, but because it really did
seem at variance with common sense. A parasite of far larger
size than its host, controlling the growth of its host—not grow-
ing within or upon the host, and following its growth at a dis-
tance, but growing outside of the host, spreading in all direc-
tions of its own motion, and being followed by the slower
growth of the host—such a parasite was indeed a novel phe-
nomenon. We cannot blame the lichenologists of the old
school for their facetious remarks about the horse parasitic
upon the bot and the symbiotic relations of Jonah and the
whale.
If all lichens were the large, robust parasites that the com-
moner lichens are, we should have reason to hesitate long
before accepting so remarkable a phenomenon as established.
` ?The case of the bacteria in the « pitchers” of Nepenthes and other carnivorous
plants seems, according to the investigations of Tischutkin, to been exception. See
AMERICAN NATURALIST, May 1893.
1893.] Symbiosis and Mutualism. 511
Fortunately, the lichens exhibit several intermediate forms,
and enable us to see the relation between the phenomenon
found in the commoner lichens and ordinary parasitism.
Lichenologists have, for a long time, distinguished, under
one name or another, two classes of lichens. In the one group
the thallus is entirely or substantially homogeneous—there is
no differentiation into rind, medulla, etc. In the other there
there is a well-defined rind, and gonidial and other zones are
differentiated. The former have been called homcomerous
lichens, the latter heteromerous lichens. In the first group
the alga is the principal part of the lichen. The hyphe grow
within the mass of algal cells and follow them in their growth.
To this class belong Collema and like genera, which are fungi
parasitic upon Nostoc, Scytonema, etc., and growing within the
gelatinous membranes and sheaths enveloping those alge.
Here there is symbiosis—a living together of parasite and host
—but no one will contend that there is mutualism.
In the second group the fungus is the principal part of the
lichen. It contains in its thallus a zone of alge, but they fol-
low the growth of the thallus, and their bulk is a small pro-
portion of the whole lichen. In these lichens the alge are
Protococcoidex or Palmellacex, ete., and to the different mode of
growth of these alge the difference is largely to be attributed.
Between these groups there are a number of forms, usually
classed as heteromerous lichens, which, nevertheless, show no
differentiation of medulla and rind, and in which the thallus
consists of a weft of slender hyphe growing around filaments
of Chroolepus and like forms. Still another fact is important
in this connection. Some of the genera of this intermediate
group have species which contain no gonidia and are sapro-
phytes upon bark, and indeed the parasitic species are often
saprophytes during a part of their existence. Many genera of
fungi exhibit the same phenomenon.
It is seen, then, that mutualism does not exist in all lichens,
and that the steps from an ordinary case of parasitism, such as
that exhibited by the homceomerous lichens, which consist of
a mass of algal cells permeated by the hyphe of a fungus and
often distorted by it, to the peculiar case of the heteromerous
512 The American Naturalist. [June,
lichens, where the fungus forms an extensive thallus in a
zone of which are contained the alge upon which it subsists,
may be traced in existing species. Not only this, but there are
genera, as has been said, in which there are species that do not
attack alge, but live independently as saprophytes, and the
point to be noticed here is that these genera belong to the
intermediate group of what I may call, pseudo-heteromerous
lichens. a cept pi
These considerations, of course, do not prove the existence of
mutualism in lichens, but they deprive it of much of its seem-
ing unreasonableness. Other facts, now well established,
make it certain that this relation really does exist in the
heteromerous lichens.
Arthonia is one of the pseudo-heteromerous lichens. More-
over, it is one of those genera in which certain species, during
their entire existence, live independently as saprophytes. Of
its development, De Bary says: “ . . . . the hyphe of
the thallus make their way into the outer layers of the peri-
derm in the smooth stems of oaks and ashes and there grow as
saprophytes independently, that is, without alge, into a thallus
formed of an abundance of slender hyphe which spread
through the cells of the periderm. Then its proper alga,
Chroolepus umbrinum, finds its way from without through the
cell walls of the peridermis into the previously formed hyphal
thallus and is seized by it. The cells of the Chroolepus are in
rows forming filaments with apical growth, and it is by means
of this growth that they penetrate into the thallus in the same
way as mycelial hyphe pierce through membranes. Thealga
is a frequent inhabitant of the bark of trees, and makes its way
into the periderm for its own purposes. Its penetration into
the thallus of the fungus can scarcely be supposed to be caused
by the fungus, but is merely an adaptation which favors the
formation of a lichen.” This is plainly an ordinary case 0
parasitism’ on the part of the lichen, but it not only throws
light on the origin of the true heteromerous lichens, but it
shows in what manner the fungus may be of benefit to the
alge. In the heteromerous lichen the thallus takes the place
of the bark of the tree in these pseudo-heteromerous lichens
1893.] Symbiosis and Mutualism. 513
The gonidia of the heteromerous lichens are usually Palmel-
laceze, which, from their different structure and mode of growth,
have not the power of getting beneath the bark as does Chroo-
lepus. The thallus of the lichen serves the same purpose
with them—protecting the colony of alge and absorbing
and retaining unusual quantities of moisture, and enabling
them to live and multiply in places where, under ordinary
conditions, life would be impossible.
That the thallus does do this is shown by the fact that
lichens grow in places where alge could not maintain them-
selves unaided, and by the fact that the gonidia multiply with
great rapidity in the thallus, often more so we are told than
without, and the individual cells attain a larger growth within
the thallus than without, as has been shown by taking algae
from the thallus and cultivating them independently. That
the fungus does not do all this for nothing, the numerous
exhausted cells to be found in the gonidial zone of any ordin-
ary lichen abundantly testify.
There is another curious phenomenon exhibited in some
lichens. In these species the alge are not confined to the
gonidial zone, but grow up into the tissues of the sporocarps
between the paraphyses and among the asci, so that when the
ascospores are ejected, cells of the alge are ejected with them
and are promptly seized upon by the germinating spores. This
can hardly be accidental, and it should be observed that it is
the alga which is the moving party, not the fungus. Surely
some benefit must result to the alga or it would not be done.
It is possible, also, that there are other adaptations resulting
from the symbiosis of fungus and alga in the lichen. Frank
claims to have discovered several, one of which deserves men-
tion. It is well known that alge can be separated from the
lichen, and that they will then vegetate in the ordinary way
independently. Frank asserts that certain species of alge
ave become so adapted to life in the lichen and so accustomed
to it, that they have partially or wholly lost the power of inde-
pendent growth. No examples of this, however, are certainly
known.
514 The American Naturalist. [Tune,
Frank also claims that the fungus exhausts the protoplasm
of algal cells without entirely destroying them. If by this
is meant that it does not always entirely destroy the cells it
attacks, it is probably so, but if anything more is meant, it
seems, like some other theories of Frank, which I shall have
occasion to mention presently, if I may say so, decidedly
“fishy.” Such a thing is not necessary to mutualism. The
alga can purchase the protection of the thallus only by the
sacrifice of a large number of individual cells. If it gets quid
pro quo, why should it not prefer to sacrifice them to the fungus
in return for the shelter of the thallus rather than to leave
them victims to natural conditions without compensating
advantage. To put the matter in another way, if the energy
spent by the alga in producing cells to be destroyed by the
fungus were put to making a shelter of its own, could it effect
as much as it does by taking advantage of the thallus?
Two other cases in the vegetable kingdom where mutualism
is thought to exist remain to be examined. These are the
cases of “ Pilzsymbiosis” or “ Wurzelsymbiosis” of the roots of
anthophytes and certain fungi. The first noticed was what is
termed “ Mycorhiza,’ and of this first.
T. Hartig, in 1840, and others since, had noticed mycelia,
apparently parasitic on the roots of trees. In 1885, Frank
published the results of investigations of mycelia growing upon
the roots of various Cupulifere in which he claimed that the
sustenance of these trees depends upon fungi symbiotic with
their roots. The title of his paper indicates his claim: “ Ueber
die auf Wurzelsymbiose beruhende Ernaehrung gewisser Baewme
durch unterirdische Pilze.” To begin with, Frank found that
certain Cupulifere have almost the whole of their root system
covered with mycelium associated symbiotically with the root
and he claimed that these fungi took the place of root hairs,
and were the only means of absorbing water, etc., possessed by |
the roots, though, of course, like the gonidia of lichens, the
roots could be grown independently in water cultures for years.
The mycelia, of the existence of which there is no doubt,
are probably connected with some of the Gasteromycetes OF — A
Tuberacez. But Frank observes that the presence of a my eb oy
1893.] Symbiosis and Mutualism. ; 515
ium does not necessarily imply the presence of the perfect
fungus fructification, as mycelia may, and often do, go on
growing in a sterile condition for years.
Frank did not stop here. He found symbiotic fungi on the
roots of many other trees, and others after him found mycelia
on the roots of various plants to which he attributes the same
relations of mutualism. His final statement is that this phe-
nomenon belongs “ to all trees under certain conditions ; ” that
“the Mycorhiza is formed only in a soil which contains
humous constituents or undecomposed vegetable remains; ”
that “the development of Mycorhiza increases or diminishes
with the poverty or richness in these constituents ; ” and that
“the fungus of the Mycorhiza conveys to the tree, not only the
necessary water and the mineral nutritive substances of the
soil, but also organic matters taken direct from the humous
and decomposing vegetable remains.” Finally, he claims that
only through the fungus can the tree employ such organic
matter directly.
If the fungus develops only in soil containing undecomposed
vegetable remains, we might ask why it takes the trouble to
attach itself symbiotically to the root and give the tree the
benefit of its saprophytism; especially, as Frank says that
the protoplasm of the cells and the fungus live together “ with-
out the former being parasitically affected or its vital phe-
nomena disturbed.” This reminds one of the exhausted gon-
idial cells which are still uninjured, and is not the only one
of Frank’s statements calculated to try our patience. and cred-
ulity.
In 1886, Warlich (Botanische Zeitung, 1886, p. 481, et seq.)
investigated certain fungi on the roots of orchids. He
examined several hundred species, all of which he found
affected on both aerial and subterranean roots with the
mycelia of what he showed to be a species of Nectria. The
-hyphe of this fungus affect spots here and there, forming
knots or coils in certain cells and causing them to enlarge,
but, as a rule, only partly filling the cell and not destroying
the protoplasm. Frank, of course, took this up, and he claims
that the protoplasm of the cell is not affected or disturbed by
516 The American Naturalist. [June,
the fungus; that the fungus is strictly connected with that
part of the plant which absorbs the food materials; and that
those orchids which are chlorophyll-less, and therefore depend
on the humus of the soil for carbonaceous matter, always
exhibit this fungus highly developed. Accordingly, he includes
this too in Mycorhiza, calling it “ endotropic Mycorhiza” (i. e.
the hyphe live in the cells) as opposed to “ ectotropic Myco-
rhiza ” in which the fungus is entirely outside of the cells.
As to Frank’s statement that the’ protoplasm of the cell is
not affected by endotropic Mycorhiza, Marshall Ward, in the
Annals of Botany for February, 1888, says: “ This can only be
an assumption, and the impression I gather from the study of
what is known of this orchid fungus is in favor of the view
that the fungus does disturb or ‘ parasitically affect’ the proto-
plasm of the cell, and that an outward and visible sign of some
such action exists in the hypertrophy of the cells affected and
in the turning yellow of the chlorophyll-grains.”
R. Hartig, a more sober and trustworthy writer than Frank,
said the last word so far on Mycorhiza in 1891. He admits
that the mycelia of some of the Tuberacex or Gasteromycetes are
found symbiotic with the roots of certain trees. But his con-
clusion is that they are of no use to the tree, and are probably
injurious by taking nourishment properly belonging to the
tree. It would seem that they must do this, even were there
mutualism between them and the roots—else why are they
there? Organisms are not given to gratuitously assisting one
another. © Mycorhiza undoubtedly exists—i. e., mycelial stages
of many fungi of different groups are parasitic upon and in
the roots of anthophytes. But that there is, in any of these
cases, more than the ordinary symbiosis of parasite and host,
has not been shown, and is improbable. That every tree has — a
its root system covered with mycelia, proves nothing. Every
tree has its bark covered with lichens, its twigs with black 4
fungi, and its leaves with parasitic fungi of every description. =
The second case of “ Wurzelsymbiosis” is the root tubercles
of the Leguminose. These tubercles have long been known
upon clover, and of late years—since 1885, in fact—have been —
found upon nearly all of the Leguminosæ. Naegeli found &
1893,] Symbiosis and Mutualism. 517
Chytridium-like parasite in the cells of Jris which has never
been seen since, and named it Schinzia. So when, in 1879,
Frank first worked upon clover tubercle, he considered it sim-
ilar to Naegeli’s Schinzia, and named it Schinzia leguminosarum.
Subsequently, a tubercle was found on the roots of Alnus by
Woronin, called by him Schinzia alni. Tubercles have been
found in this country on the roots of Ceanothus, and are known
on a few other plants besides the Leguminose.
There has been considerable uncertainty as to the cause of
clover tubercle and the nature of the parasite to which it is
due. Schroeter took the parasite for a Myxomycete similar to
Plasmodiophora and named it Phytomyza. Marshall Ward, in
the article cited, compares it to the yeast fungi. De Bary, in
1884, dismissed the matter with a sneer. Frank now puts the
parasite among the Schizomycetes, and, indeed, the best view
seems to be that the parasites are bacteria pure and simple.
There are, in some tubercles, hyphe, or something very like
hyphe, which Frank now calls “ Infektionsfaden.” Marshall
Ward considered these the hyphe of which what some call the
“ baktroiden ”—i. e., the bacteria—were spores. Schroeter saw
in them a plasmodium. Frank, always unique and startling,
has finally (1891) decided that the “ Infektionsfaden” have
nothing to do with the fungus, but are products of the host for
the purpose of self infection! These hyphe are usually filled
with the “ baktroiden,” and Thaxter’s recent discovery of Myzxo-
bacteria may throw some light upon their true nature. In an
article in the Torrey Bulletin for July, 1892, Mr. Schneider
concludes that these tubes have nothing to do with the bac-
teria, or Rhizobia, as Frank now calls them, and considers them -
hyphal fungi related to the parasite of Alnus tubercle. As
these tubes often contain the bacteria, this seems improbable.
-From all that I have read and seen, I am satisfied that the
parasites are bacteria, and I see no reason for separating them
from the rest of the Schizomycetes as Schneider does. I even
doubt the necessity of creating a separate genus for them, as
Frank did in 1890, under the name of “ Rhizobium” (Pilzsym=
biose der Leguminosen).
518 The American Naturalist. [June,
_ These tubercles are fine examples of symbiosis, and it has
recently appeared probable that they exhibit mutualism of an
unexpected kind, analogous to that claimed by Frank for his
Mycorhiza. I can only go into this briefly. It is known that
the plant cannot directly assimilate free nitrogen. Yet, as
Marshall Ward puts it, “ For a long time it has been generally
known that the Leguminosex, especially, have what we may
term a special aptitude for seizing large quantities of nitrogen-
ous substances from the soil, and this problem has become a
classical puzzle in vegetable physiology.” In 1886, Hellriegel
and Wilfarth published some investigations of this matter.
Subsequent experiments founded on theirs have been very
numerous, and are yet in progress, but their researches remain
our principal authority on the subject. Without detailing
them, I may say that these researches seem to demonstrate
that this power of taking up large quantities of nitrogen
depends entirely upon the presence or absence of the tubercles
—that without them it does not exist, and that it exists in
greater or less degree according to their abundance. Conced-
ing this, two theories are. possible as to the cause.
In 1888, Marshall Ward appeared to think that the parasite
stimulated the cells to extraordinary metabolic activity, and
that was probably all it did. This view has had no followers
so far as I can find.
The other possible theory is that the parasite does this work
and the host takes advantage of it. Frank, as might be
expected, takes this view. The most recent observations seem
to have settled pretty thoroughly that the tubercles do assist
the plant in some way in assimilating free nitrogen, and that
here is a case of mutualism analogous to that of the lichens.
The bacteria (as I assume that they are) are parasites. They
are there for their own purposes, and are incidentally benefi-
cial to the plant. The plant, it is generally admitted, can exist
and thrive without them. In some cases it appears, and the
analogy of the lichens makes this probable, that the bacteria
“are purely parasitic, and that there is symbiosis without
mutualism. But, in most cases of the Leguminose, it seems to
be shown that the plants affected do better than those unaf
1893.] Symbiosis and Mutualism. 519
fected. Much research is needed in this matter. The manner
in which the parasite acts and the host takes advantage of its
work are not known with any certainty.
To these probabilities, Frank adds certain characteristic
improbabilities. One has already been spoken of, namely,
that the plant develops tubes or hyphee for the purpose of self-
infection which it sends through its tissues. This is somewhat
like the algæ in some lichens which grow up among the asci
in the sporocarp and are ejected with the spores. Only the
latter is an established fact, the former a feat of the imagina-
tion. Another of his ideas, pronounced a “settled fact” by
Schneider in the article cited, is that “at the close of vegeta-
tion and on other special occasions, the plant reabsorbs the
protoplasm of the fungi.” After all the trouble of self-infec-
tion to which the host has been, this seems rather like killing
the hen that laid the golden egg. There is no sufficient evi-
dence to establish so remarkable a phenomenon. Finally,
Frank thinks that the roots of the Legwminose possess the
power of attracting Rhizobia, due, as he considers, to some
secretion. This is too much for his followers, and I think all
will agree that it is the last straw of an unsupportable load
with which he has already burdened our credulity. The
exuberance of Frank’s enthusiasm, however, should not blind
us to the fact that some relation of mutualism between the
Leguminose and the tubercle parasites probably—almost cer-
tainly—does exist.
It is not necessary, as Frank seems to think, in order to
establish mutualism to show that the organisms do no injury
to each other. Mutualism of the kind we meet with in the
vegetable kingdom involves sacrifices on the part of the host.
The parasite is not there gratuitously. It is there to steal from
its host the living it is hereditarily and constitutionally indis-
posed to make for itself. If the host gains any advantage from
the relation, it can only do so by sacrificing—by giving the para-
site the benefit of its labor that it may subsist. If the plant or
the plant colony benefits as a whole, it purchases the benefit by
the sacrifice of certain parts or individuals. Mr. Webber, ina
note on the Yucca moth in the American Natura ist for Sep-
520 The American Naturalist. [June,
tember, 1892, makes a significant remark to the same effect:
“The larva of Pronuba uses up only from 10 to 12 seeds, s0
that even in those capsules where the most abundant larvæ
develop, hundreds of good seeds are nevertheless developed.
The few seeds destroyed may well be sacrificed to insure the
pollination and development of the others.”
Ethieally, there is nothing in the phenomena of symbiosis
to justify the sentimentalism they have excited in certain
writers. Practically, in some instances, symbiosis seems to
result in mutual advantage. Inall cases it results advantage-
ously to one of the parties, and we can never be sure that the
other would not have been nearly as well off, if left to itself.
1893.] Evolution in the Genus Megascops. 521
EVOLUTION AND DICHROMATISM IN THE GENUS
MEGASCOPS.
By E. M. HASBROUCK.
INTRODUCTION.
The subject of Dichromatism has ever been a stumbling-
block to scientific research, and from the time of its earliest
admittance into zoological discussion, has been without
apparent cause and without explanation. Few, if any, theo-
ries have been published as to what it really is, or as to the
possible influence governing it, while any theories that have
been advanced stand without proof of their correctness; a
new theory, therefore, cannot be considered as conflicting with
any settled doctrine upon the subject. When we find a peculi-
arity common to all forms of animal life, it becomes apparent
that if thoroughly understood in one form, a flood of light
will be thrown upon the same perplexing problem affecting
the rest of the animal kingdom.
In attempting to explain the causes and effects of dichrom-
atism in the Screech Owl, no stone has been left unturned to
prove the correctness of the views herein set forth. The writer
has labored lovingly and patiently, following a course of rea-
soning which he believes to be correct, and in submitting his
work and its results to fellow-workers would ask for it their
kindly consideration. The collection of the necessary data
has been an extremely difficult task, and more information is
still desirable from certain localities, although sufficient has
been received to make possible the construction of a compara-
tively accurate map of the color distribution. It is a lament-
able fact that scientific observers of bird life are exceedingly
scarce in the southern states and in the lower part of the Mis-
sissippi Valley. The data obtained from these regions,
although to a certain extent more scanty than could be
desired, has been used. to the best possible advantage. In
obtaining information, copies of the following circular letter
522 The American Naturalist. [June,
were sent out by the National Museum, and the matter
received in reply was turned over to me for study and investi-
gation, together with the privilege of preparing the results for
publication.
UNITED STATES NATIONAL MUSEUM
DER DIRECTION OF
THE SMITHSONIAN INSTITUTION.
Dear Sir:
The National Museum is engaged in a systematic investigation of the color-
hases of the Screech Owl ( Megascops asio), with the view of determining the relative
abundance of the red and gray plumages in different parts of the country, the plu-
mage of the young, produced by parents of known character as to plumage, and the
other related questions.
rder that the investigation may be as thorough as possible, assistance in
the matter is requested; and if you can give information on the subject, your kind
co-operation will be much appreciated.
Should you favor us with any information on the subject, please arrange the
data in the following order:
ame your locality, including County and State.
(2) State about how many specimens have come under your notice, and how
many of them were in the gray plumage.
(3) If you have obtained or observed a pair of old birds with their young, state
the character of plumage of the former (whether both red, both gray, or one red an
one gray—and if the latter, the sex represented by each phase), and also the plumage
of the young.
The above are the principal points upon which information is desired, but any
additional memoranda will be very acceptable.
ours truly,
ROBERT RIDGWAY,
Curator, Department of Birds.
In conclusion I wish to express my gratitude to Professor
G. Brown Goode, Assist. Secretary of the Smithsonian, and
Mr. Robert Ridgway, Curator of Birds, who rendered every
assistance and courtesy in their power; while to all who
responded to the request for information, without which the
task would have been impossible, is due the sincere thans
of The Author
Part I.
RELATIONSHIP OF DICHROMATISM TO EVOLUTION,
So universally adopted has the theory of evolution become, —
that it scarcely seems necessary to refer to it as forming thé
Ste ee eee ee a
ep ee ae all) Rete
1893.] Evolution in the Genus Megascops. 523
foundation of the theory here advanced, regarding the dichro-
matic phases of the common Screech Owl (Megascops asio).
“In the Systema Nature (Vol. I, 12 Ed., 1766, p. 132), the
Red Owl is first described by Linnzus under the name of
Striz asio. Twenty-two years later (1788) Gmelin described
(Systema Nature, Vol. I, 13 Ed., p. 289) the Mottled Owl as
Strix nevia. In 1812, in the fifth volume of his American
Ornithology, Alexander Wilson re-describes the two under the
same names, also as distinct species, and not until 1828 does it
appear to have been publicly hinted that the two were really
identical, when Prince C. L. Bonaparte united them, he con-
sidering the red bird as the young, and the gray the old.
Audubon, in 1832, sustains this view; one of the red birds he
figures as the young, being one he reared from a fledgling.
Nuttall, a few years later, supports the same view. In 1837,
Dr. S. Cabot, Jr. (Journ. Bost. Soc. Nat. Hist., Vol. II, p. 126),
while considering the two birds identical in species, reverses
the order, making the red plumage the old and the gray the
young, and in confirmation of his views exhibited as seeming
conclusive evidence, an old red bird which he shot while in
the act of feeding some gray young, which he also exhibited.
In July of the same year, Dr. Ezra Michener, in a paper (Phila.
Acad. Nat. Sci., Vol. 7, p. 53) entitled, A few Facts, in Relation
to the Identity of the Red and Mottled Oul, states that he had
seen young Screech Owls, accompanied by their parents, after
leaving the nest, of both red and gray colors, the parents
being always of thesame color as the young. “The conclu-
sion is, therefore,” he says, “ evident, either that the color of
both old and young is variable or uncertain, or that they are
specifically distinct.” The latter opinion he adopts, ignoring
the then sole known case of different colors in the young and
parent exhibited by Dr. Cabot, very positively concluding
that there are two species, and that Wilson was right.
Dr. P. R. Hoy, in his Notes on the Birds of Wisconsin (Proc.
Phil. Acad. Nat. Sci., 1853, Vol. 6), gives them as two distinct
species, while Cassin, in his various papers on the owls,
adopts the conclusion of Bonaparte; considering them as one
species, and the gray the adult.” *
Am. Nat., II, 1866, 327-328.
524 The American Naturalist. [June,
Again, Dr. W. Wood (Am. Nat., Vol. II, 1868, p. 371), men-
tions two cases :—the one, a Mottled Owl, taken from the nest
with one young, neither of which had a red mark on them;
the other, a Red Owl, taken with four young, all red. His
deductions are, that there are two adults, one red without a
gray feather, the other gray without the slightest trace of red;
also, that there are young of each before they are able to fly,
one gray and white without a red feather, the other with a
reddish tinge to all the feathers. These facts he is unable to
reconcile, unless, as he says, “ It is admitted that the color of
the plumage is either ‘ variable or uncertain, or else there are
two distinct species as described by Wilson.”
The whole is admirably summarized and described in
Baird, Brewer and Ridgway (Vol. III, 1874, p. 51), in the fol-
lowing words, “ That these two very different plumages are
entirely independent of age, sex or season, and that they are
purely individual there can be no doubt, since in one nest
there may often be found both red and gray young ones, while
their parents may be either both red or both gray, the male
red and the female gray, or vice versa. Occasionally specimens
are exactly intermediate between these two plumages, it being
difficult to decide which predominates.”
This difference in plumage has been termed Dichromatism,
and while a hundred and twenty-five years have succeeded in
establishing the fact that the red and gray screech owls are
one and the same species, no satisfactory theory known to
the writer has been advanced as to what dichromatism is due
in this case or as to the possible causes governing it.
Naturally enough, this is far from satisfactory, while the opin-
ions of various authors just quoted, and which comprise the
most of what has been written upon the subject, tend but to
raise innumerable queries for which there appear to be no sat-
isfactory answers. Why, for instance, is the gray form doml-
nant in one part of the country and the red in another? Why
are both forms equally common in a third? Why is the red
phase peculiar to the eastern members of the genus, while 1n
the western forms it is unknown? Above all, why, at the
northern boundary of asio proper, is the red form entirely
PLATE XIII.
Fe i
Liobunum ventricosum hyemale Weed. Immature.
1893.] Evolution in the Genus Megascops. 525
absent, while at the southern limit it is greatly in the major-
ity? These and many other questions of equal importance
present themselves, and but confuse and leave the reader
more than ever in the dark.
While collecting screech owls in the District of Columbia
during the winter of 1890-91, these questions arose very forci-
bly, and upon investigating, I found that while the matter
was apparently settled to the satisfaction of ornithologists, as
regards the identity of the two forms, nothing further
appeared to be known aboutit. Further investigation showed
that dichromatism is principally confined to the typical form
of Megascops asio, appearing but slightly in the Florida form
(Megascops a. floridanus), and barely reaching the Texan sub-
species, mccallii. The remaining members of the group, four
in number, remaining true to their normal color.
Three phases of plumage were clearly defined—the red, the
gray, and the intermediate (the last less frequently seen than
the others). Three distinct phases, represented indiscrimi-
nately by individuals of one-species, regardless of age, sex or
season! Such an unusual circumstance that the possibility of
some great and important change gradually taking place at
once suggested itself, which has been proved to be in reality
the case.
An attempt will be made to show, first, that while the red,
the gray and the intermediate phases are at present but indi-
vidual variations of the same species—the gray was the ances-
tral stock ; second, that from the gray bird has evolved the
red, which at some future time will be a recognized sub-spe-
cies with a range peculiar to itself, and thus dichromatism 1s
one step in the evolution of the Screech Owl, while the vari-
ous phases exhibited are the transitorial stages of develop-
ment of one species from another; third, that this condition of
affairs is influenced by four powerful factors (two of which
; sub-speci ibed by Brewster (Auk, Vol. VIII, 1891, 139-
1) wel make piorar para aipha are al yet admitted to the check
EAT a lennie exkibits a dict tic phase, but it is a decided dark brown, and
“Rot a reddish phase, and is not included in the present treatise. '
35
526 The American Naturalist. . [June,
temperature and humidity, are dominant powers in geograph-
ic distribution), the most potent of which is temperature;
fourth, that the predominating distribution of the respective
colors is largely confined to the faunal divisions of the East-
ern United States, and as such is approaching the sub-specific
differentiation of the two phases; lastly, that the Darwinian
theory of ‘Reproduction with variation and the survival of
the fittest,’ is well exemplified in our common little Megascops
0.
The Screech Owl has been made the subject of investigation
simply because it offers one of the best known examples of
dichromatism : if explained and disposed of in one case, much
light will be thrown upon the same problem relating to many
forms of animal life.
To begin with, a short description of the young, together
with the manner of attaining the adult plumage of both
phases, may be of service to many not thoroughly acquainted
with the species. When first hatched they are covered with.
pure white down, which in a week or ten days begins to show
faint traces of transverse dusky bars.. At the age of four
weeks they are fully barred with gray and white alternately,
which markings they retain until the appearance of the first
full plumage. As a usual thing, those destined to be red
assume a rufous cast in the down shortly before the appear-
ance of the feathers, but this is not always the case, as
instances are known where the first intimation of red was the
appearance of a rufous feather pushing its way through the
gray or mottled down. Such instances, however, are rare, and
but few are known to the writer. At the period when feathers
first appear, the future color of the young bird is settled
beyond a doubt; both the gray and the red birds rapidly
attain their mantle, and the colors once assumed are never
changed. Much confusion has arisen from the fact that some
observers apparently make no distinction between the bird in
the gray or mottled down and the gray or mottled plumage
and upon the appearance of the red feathers through the gray
down, the assertion has been made that the gray feathers
changed to red.. The gray birds, on the other hand, attain
1893.] Evolution in the Genus Megascops. 527
their plumage at precisely the same time and in the same
manner as the reds, only that their feathers are gray instead
of rufous. If the above simple facts are borne in mind, no
trouble need be experienced in undertanding the appearance
of the two plumages.
The first step in the work was to ascertain the relative pro-
portion of the red and gray phase in every section of the area
covered by the asio group of Megascops affected by dichroma-
tism. For this purpose nearly one thousand copies of the cir-
cular letter referred to in the introduction, were sent to the
ornithologists distributed over this territory. Over two hun-
dred replies were received, a few of which failed to be suffici-
ently explicit to make their contents available, but enough
were obtained to compile in a reasonably accurate manner the
map showing the distribution of the two color phases (map 2).
On the northern border of the range of Megascops asio
where gray is the only form known (see map 2), corre-
spondents from southern Ontario state that at occasional rare
intervals a red specimen has been taken in the spring; all these
birds were taken close to Lakes Erie and Ontario—while at
Toronto, one pair composed of a gray and red have been
known to breed, which necessitates considering the red phase .
in Ontario. MclInraith states that the species is migratory
there, to a certain extent, and it is probable that some of the
red birds may be chance visitors that have come north with
others. With this exception the gray belt is unbroken from
Picton, Nova Scotia, through New Brunswick; northern
Maine, New Hampshire, Vermont and New York; the south-
ern part of Quebec and Ontario; northern Michigan and Wis-
consin; a strip through central Minnesota, thence south-west-
ward across, Dakota and most of Nebraska, and ending
in a long point near Leavenworth, Kansas. On the south
this gray belt is somewhat broken as shown on map 2; in the
‘uture more complete data may show it as unbroken here from
east to west as it is on the north. Beginning at a point near
St. Augustine, in Florida, it encompasses nearly the whole of
that state and much of the range of the Florida form, Megascops
a. floridanus; runs northward and westward to Eufaula in
528 The American Naturalist. [June,
Alabama, and terminates near Mobile (this terminal point is
arbitrary); while in Texas it is resumed in the southwestern
portion of the range of mecallii?
Next to this belt comes that wherein both phases occur, but
where the gray predominates. It begins near Belfast, Maine,
follows the exclusive gray belt with irregular outline to its ter-
mination in northern Kansas, and continues southward ina
broad belt into Louisiana, where it is a question whether or
not it becomes broken at New Orleans. Immediately north
of New Orleans it begins again at Mandeville, La. (based on
specimens in Am. Museum), and continues eastward to the
coast in the neighborhood of Savannah. In its northern half
it sends a long arm southward, through eastern Massachusetts,
into central Rhode Island and Connecticut, again in Western
Pennsylvania, it reaches in a narrow strip far down the Alle-
ghanies into Virginia and West Virginia, terminating near
latitude 37°, covers practically the whole of Ohio, with a good
share of Indiana, and continues thence to Leavenworth, Kan-
sas, the delta of the Mississippi, and to the Atlantic.
Within this area lies the region in which both forms occur,
but in which red largely predominates (which in turn includes
the exclusive red areas), and which occupies nearly one-half
the area inhabited by asio proper. It of course conforms to
the outline of the last described belt, and extends along the
entire Atlantic coast from New England to Savannah.
Lastly are the two areas where red is the exclusive form.
The one of considerable size and importance based upon sub-
stantial evidence, lies wholly within the Mississippi Valley:
the other, a small strip, extends from a little northwest of
Oakdale, North Carolina, to Variety Mills, in Virginia, and
including Wytheville. At present the grays are apparently
unknown from these areas. It has been impossible to examine
material from either, and to those who may have the oppor
tunity, I would point out the desirability of collecting all the
specimens possible.
* The intermediate region is mapped upon very insufficient data, it being next to
impossible to learn anything of this territory, but the best has been done that was pos-
sible under the circumstances,
1893.] Evolution in the Genus Megascops. 529
The following table shows the data arranged by states, upon
which the map (No. 2) of color distribution is based. The
original letters had to be consulted in many cases as to the
proper placing of the lines marking the several divisions; the
table itself shows but two things: the areas where reds or
grays are the exclusive form known, and where each predomi-
nates in the mixture of the two; the finer distinctions it is of
course impossible to show in tabular form.
It will be remembered from quotations at the beginning,
that the young produced by parents of different colors were
said to be either all red, all gray, or both; that those produced
by parents both of which were red were the same; this much
is true, but not a single record can be found of the offsprings of a
pair of gray birds showing the slightest trace of red. That the
grays invariably breed true even in a region where red. is the
predominating color, and where the individuals in question may
themselves be the offspring of red parents, is of itself a strong point
in support of the theory that the gray was the ancestral color.
Further, taking into consideration that there are certain areas
where a red bird is unknown, it is evident that the grays do
certainly produce young the color of their parents; and that
where we take the red birds over a very large area and find
them continually producing gray birds together with red, and
compare them with the gray birds which never produce a red ; it
would appear that the gray bird was the original stock, and that
the red was an offshoot—a branch, so to speak, which, owing to
certain climatic conditions, or certain elements in the environ-
ment, gave it the supremacy in the struggle for existence, and
that the producing of gray birds by red parents is an evident
tendency to revert to ancestral characters. Nor is that old
maxim in natural history, natura non facit saltum to be ignored ;
that natural selection played and is playing an important
part in the matter is evident, else how came that area in
which the grays are entirely absent? That the red birds
were a ‘sport,’ or a freak, and suddenly acquired their charac-
ters, is not to be supposed for a moment; but on the theory of
‘reproduction with variation’ and ‘natural selection,’ I hold
that some slight deviation from the parent stock did at one
530
The American Naturalist.
[June,
TABLE SHOWING DATA BY STATES, UPON WHICH MAP OF COLOR
DISTRIBUTION IS BASED.
STATE. Rep Gray, SOURCE oF INFORMATION.
Alabama. Eufaula. E. L. Bro
Arkansas. Cerro Gordo. Chas. A. ike
Clinton, Mrs. Lillie ct (Oologist, VIII, 1891,
195).
Connecticut. Portland. H. Sage
East Hartford.| Willard E. Treat.
: Wilmington. Walter D. Bush
Florida. alatka Jas. Sands.
Thonotosassa.| W. H. Steacy.
Citronelle Wn. H. Jeremiah
Georgia. Faceville, W.B. MeDaid.
Illinois. Mt, Carmel. R. Ridgway (Nat. Hist, Surv. Il., I,
1889, 417.)
Odin. C. B. Vandycook
Jacksonville, Chas. W. Tindall
arsaw. has. K. Worthen
Rockford. J. E. Dickinson
Glen Ellyn. . T. Gault,
Indiana, Camden Trans. Ind. Hort. Soc., 1890, 52.
Terre Haute, Trans. Ind. Hort. Soc., 1890, 52.
Dunreith. Pleas
ond. Mrs. Lillie E. Pleas
Indianapolis. Fletcher M. N
Iowa. La Porte City. o. D. Peck
Sioux City. Dr. Guy C. Rich
Grinnell. Lynds Jones
Kansas, tine, G. E. Stilwell
Manhattan, ntz.
Kentucky Ellis, Dr. Louis Watson
Morganfield. ~ Lemen.
Garrard Co. G. V. Young.
Versailles. L. O. Pindar.
Bell. Carrington C. Bacon.
Louisiana, New Orleans, ustave Kohn.
Mandeville. Collection Aus. Mus. Nat. Hist.
Maine. Bangor. Merrill.
Maryland & Dist. of g
Columbia. Washington AE a
ashington. m. Palm
Washington. C W. DARIE
Washington. J. D. Figgins.
Washi ; E. M. Hasbrouck
Sandy Spring James P. Stabler.
; Laurel. Mars
Massachusetts. North Stoughton C. A. Rum
Boston. Wm. oa E. W. Ricker.
Holyoke, W. F. La
: aunton, A.C. Pe
Michigan. Grand Rapids. art E. White.
ù Morris Gibbs.
East Saginaw. Lepp.
1893.] Evolution in the Genus Megascops. 531
STATE, Rep. Gray. SOURCE oF INFORMATION.
Minnesota. Rochester, W.D.H her
Lake City. E. A. Wis
Mississippi. Waverly. EN: Cound
Missouri. Fayette. J. W. Kilpatrick
Butler. Harvey Clark.
Independence.| Charles W. Tindall.
Nebraska. Ong. oel Nelson.
London. Geo. oe
New Hampshire. Milford. J. P. Mel
Webster. Charles F. Gecthes (F. & S., VILI;
113).
New Brunswick. New Castle. Phillip Cox.
New Jersey. Morristown. Specimens in Am. Mus. Nat. Hist.
New York. Albany . Arnold.
Oneida Co. B. W. Arnold
Lowville. C. Fred.
Lockport J. L. Davison.
Owego. . Alden ing.
Binghampton.| Willard N. Clute.
Port Byron. . M. Hasbrouck.
North Carolina, e. Robt. J. Thompson.
Weaverville. J. S. Cairns
i H. H. & CS. B
Nova Scotia. Pictou (?)- Prof. A.M. Sortanee Prof.McCulloch
Ohio. Ney. Jno. O. Snyder.
Hamilton. Harbron & Dr. F. W. Langdon.
Canton R. H. Bulley.
peice be John. W. (Jour. Cin. Soc. Nat.
er Hist., VIII, 1885, 52).
Lebanon. Raymond W. Smith
Salineville. m. A. e
Doni Plover Mills. | Robert Elliott.
Lansdowne. C. J. Young.
arker. John Ewart.
Danville. Dr. G. A. McCallum.
Hami T. Mclluraith.
Bradford. A. C. Sloam
Listowell. Wm. D. Kells
Annan. . C. Sloam.
Ardtrea. John Blair.
Toronto. James R. Thurston.
—— pss ys Del. Val. Orn
i ‘: Chas. A. roc. -
Pennsylvania Philadelphia. Club, I, ’90-’91, p. 12).
Ss August
a T Arthur B. Williams, Jr.
Rhode Island Providence. seu E. Doe.
_A
Pawtucket. EAT HA.
ith i ur. ayne.
South Carolina. Mount Pleasant. Walter Hoxie.
J. B. Cathey.
Tennessee, Duck River. B. F. Schulte.
Teraz Giddings. J. O. Singley.
‘ San Antonio. H. P. Attwater.
Gainesville. G. H. Ragsdale.
532 The American Naturalist. (Tune,
STATE, Rep. Gray. | Source oF INFORMATION.
|
Texas. Tyler. W.-L. McDaniel.
i is Co. Chas. D. Oldright.
Vermont, East Berkshire G. B. Hopkins.
Strafford. Chas. P. Collins.
Lunnenburg. W. E. Balch.
Castleton. A. O. Johnso
St. Johnsbury.| Franklin Fairbanks.
Virginia, Variety Mills. H. M. Micklem
ytheville. John B. Barrett, Jr
Eagle Rock. J. T: Paxton.
New Market. Geo. M. Neese.
West Virginia. Charlestown. B. W. Mitchell.
French Creek.| Earle A. Brooks.
White Sulphur
Springs. Thaddeus Surber.
Wisconsin. Madison. Chas. F. Carr.
Madison. : herari e Vol. I, 188.
Racine. Dr. F: Hoy.
Jefferson. Ludwig tao
Pewaukee. B. F. Goss
time occur, which in some way better fitted the individual for
the struggle for existance. Through countless generations this
has been perpetuated by the inter-breeding of those possessing
- it, in consequence, the grays, as has been seen (see map), have
entirely disappeared in at least one portion of the country, and
ve become extremely rare in others. Many difficulties exist,
both in showing the original condition, and in explaining the
present state of affairs. In early scientific work, as is well known,
no attention whatever was paid to matters of this nature, con-
sequently it is impossible to ascertain the proportion of
and gray birds at a period say four hundred years ago—still,
I hope to reason by analogy. As for the possible causes influ-
encing the change, they will be found fully treated in their
` proper place; at present the theory is either to be proved or
disproved.
Assuming that the gray was the ancestral stock, and that
the producing of gray birds by red parents is a tendency to
revert to ancestral characters, an aneigons case will be found
in Darwin’s own experiments.
As is well known, many of these were performed with pig-
eons, which proved to be the best subjects, and the following
is quoted from the “Origin of Species: ”
*Origin of Species, pp. 17-19.
1893.] Evolution in the Genus Megascops. ` 533
“Great as are the differences between the breeds of the pig-
eon, I am fully convinced that the common opinion of natu-
ralists is correct, namely, that all are descended from the rock
pigeon, Columba livia * * * * The rock pigeon
is of a slaty blue with white loins, * * The tail has a
terminal dark bar, with the outer feathers externally edged at
the base with white. The wings have two black bars. * *
Now in every one of the domestic breeds, taking thoroughly
well bred birds, all the above marks, even to the white edging
of the outer tail feathers, sometimes occur perfectly developed.
Moreover, when birds belonging to two or more distinct breeds
are crossed, none of which are blue or have any of the speci-
fied marks, the mongrel offspring are very apt suddenly to
acquire these characters.” (If this be true of distinct species,
how much more isit true of different color phases of the same
species). “To give one instance out of several I have
observed: I crossed some white fantails, which breed very true,
with some black barbs—and it so happens that blue varieties
‘of barbs are so rare that I never heard of an instance in Eng-
land, and the mongrels were black brown and mottled. I also
crossed a barb with a spot, which is a white bird with red tail
and red spot on the forehead, and which notoriously breeds
very true; the mongrels were dusky and mottled. I then
crossed one of the mongrel barb-fantails with a mongrel barb-
spot, and they produced a bird of as beautiful a blue color,
with the white loins, double black wing bar, and barred and
-white edged tail feathers, as any wild rock pigeon! We can
understand these facts, on the well known principle of rever-
sion to ancestral characters, if all the domestic breeds are
descended from the rock pigeon.”
Now if this reversion to ancestral characters occurs in breeds
so far removed from the parent stock and from each other—
how much more is it reasonable to suppose that a color phase
of the screech owl, when found continually producing gray
birds, while gray birds produce grays alone—is but a repeti-
tion only between birds more closely related, of the same per-
formance?
: (To be continued.)
. 534 The American Naturalist. [June,
THE CINNAMON HARVEST-SPIDER AND ITS VARIA-
By CLARENCE M. WEED.
In his monograph of the Phalangiide, published’ in 1868,
Dr. H. ©. Wood described a well-marked harvest-spider as
Phalangium ventricosum. The description was drawn up from
a single female taken near Philadelphia, and specimens of a
male Phalangium collected in West Virginia supposed to
belong to the same species. Besides these Dr. Wood had a
‘number of female harvest-spiders from Nebraska “which pre-
sent apparently the same specific characters as the former,
except that the legs are a little shorter. Suites of specimens
from the two localities would however probably show them to
be distinct.”
In the same paper, Dr. Wood described as Phalangium
formosum another well-marked form found in spring.
Aside from a mention in 1869 by Dr. Packard in his Guide
to the Study of Insects (p. 657), and a bibliographical reference
in 1885 by Professor Underwood,’ these species appear not to
have been noticed in our literature for nearly twenty years.
Since then, however, I have referred to them in a number of
papers.
My first mention was published in 1887* when I provision-
ally referred P. formosum to the genus Liobunum, and con-
jectured, without having seen specimens, that P. venéricosum
also belonged to that genus. Two years later I published’
extended descriptions of P. formosum referring it to Liobunum
“with considerable hesitancy, as it does not strictly belong
-there on account of the projecting inner angle of the palpal
patella. ”
1Comm. Essex Institute, Vol. VI.
‘Bull. Ill. St. Lab, Nat. Hist., III, 91.
1893.] Harvest-Spider and its Variations. 535
These forms were next discussed in my paper on “The
Harvest-spiders of North America,” in which P. ventricosum is
definitely placed in Liobunum, and P. formosum together with
a southern form not before described is referred to the new
genus Forbesium. The new form is named F. hyemale. “The
former (F. formosum) is a distinctly northern species, ranging
from New York to Colorado, while the latter (F, hyemale) is
evidently its southern representative. ”
Finally in the American Naturatist for September, 1892,
I announced that by keeping specimens of F. formosum taken
in early spring in vivaria, I had determined that it was the
immature form of L. ventricosum; and about the same time I
published® extended descriptions of adults of both sexes of L.
ventricosum.
For the purposes of the present study I have had a consider-
erable number of adult specimens from nine widely separated
states, viz.; Maine, New Hampshire, New York, Ohio, Michi-
igan, Illinois, Nebraska, North Carolina and Mississippi.
Besides the adults I have had an immense number of the
immature form (formosum) and a few of the form at present
called F. hyemale.
The first glance at the adult specimens shows that there is a
great variation in the size of the body and length of legs in
different localities. The Mississippi forms are twice as large
as those from New Hampshire; while those from the states
between present intermediate sizes. This is shown by the
measurements in millimeters in the following table; numbers
1 to 6 are males, and 7 to 12 females.
To show graphically the gradual lengthening of the legs of
_ this harvest-spider as it goes southward I have reduced the
length of the second pair, as given in this table, to the
straight lines reproduced on the opposite page. The Miss-
issippi specimens were taken at the Agricultural College in the
central part of thestate. I regret that I have not more speci-
mens from the region between Ohio and Mississippi to show
the transition more completely.
*Am. Nat., Oct., 1890.
*Trans, Am. Ent., Soc., XIX, 188.
The American Naturalist. [June,
536
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ÁyesoT jo
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‘1UASOIAJUIA WNUNGOLT JO SUOTILUBA—'T Iq",
1893.]
Harvest-Spider and its Variations.
Males.
New Hampshire.
Michigan.
Ohio.
North Carolina,
Mississippi.
or
~q
Females.
Maine.
Michigan.
New York.
Ohio.
Mississippi.
538 The American Naturalist. (June,
Like other harvest-spiders this species also varies consider-
ably in a given locality. It is so rare that it is difficult to get
long series from one place so that I cannot tabulate at present
these variations as fully as has been done for Phalangium cin- .
ereum and Liobunum vittatum dorsatum. To indicate the vari-
ation that sometimes occurs in a single state I may record that
of two specimens from Dover, New Hampshire, measured
since the above tables were prepared, the body of the male
was 8 mm. long, and the second legs 63 mm., while the body
of the female was 10 mm. long, and the second legs 50 mm.
The second legs of this male specimen were 15.5 mm, longer
than the one from Hanover recorded in the table. It should
be stated however that Dover is distinctly within the region
of the Alleghanian fauna, and Hanover is on the border
between the Alleghanian and Canadian faunas.
A study of all the specimens of my Forbesium hyemale now
accessible leads to the conclusion that this is the immature
stage of the southern form of L. ventricosum, just as Wood’s P.
formosum is the immature stage of the northern form of this
species. The dissection of specimens shows that they are not
FIGURE 1.—Liobunum ventricosum. Immature: a, body; 4, eye eminence, side
view: c, same, front view; æd, palpus; e, palpal claws; all magnified.
sexually mature, and the dates of capture indicate that they
disappear late in spring or early in summer at the time the
TAMERICAN NATURALIST, Vol, XXVI, pp. 34-36.
1893.] Harvest-Spider and its Variations. 539
adult L. ventricosum appear. This is precisely the condition in
the northern states. An idea of the similarity of the two
immature forms in structural details may be obtained by com-
paring Fig. 1 with Plate XIV, Fig. 2.
The southern form of L. ventricosum should evidently be
considered a geographical race sufficiently distinct for sub-
specific name. Inasmuch as hyemale has already been applied
to the stage immediately preceding the adult it may well be
used for this race, and the form be known as Liobunum ventri-
cosum hyemale. Just where the line dividing the two forms
should be drawn is difficult to say without more material, but
it probably occurs near the latitude of southern Ohio.
LAws oF VARIATION IN HARVKST-SPIDERS.
Comparatively little definite investigation of the geographi-
cal variation of North American invertebrates has yet been
undertaken. In the vertebrates—especially birds and mam-
mals—much attention to the subject has been given by Baird,
Allen and other well-known zoologists, so that a number of
general laws have been formulated To determine to what
extent these laws hold true for the invertebrates, as repre-
sented by the Phalangiide, I have ventured to formulate some
of the results obtained in my studies of this group—results
which in part have already appeared in the Narura.ist and
in part are yet unpublished. At'the present stage of investi-
gation such formule cannot be considered as final by any
means, but if they serve no other purpose they will be useful
in determining the direction of future work.
(1.) In mature individuals of the same species and sex from
a given locality there may be found a decided variation in
size of body and length of legs. In a series of fifty or a hun-
dred such individuals taken at random the body of the
largest specimen will usually be from one-fifth to one-fourth
longer than that of the smallest; and the legs of the longest-
legged individual are likely to be from one-fifth to one-third
longer than those of the shortest-legged specimen.
See the AMERICAN NATURALIST, Vol. XXVI, pp. 87-89.
540 The American Naturalist. [June, :
(2.) As a rule, not without exceptions, the legs vary together
in a given direction ; that is, if in a certain individual the
first pair are longer than the first pair of another individual,
the other legs of the former are likely to be longer than the
corresponding legs of the latter. The hind legs are the most
irregular in their variation.
(3.) There is a gradual increase in size of body and length
of legs in the individuals of a species from the north, south-
ward at least as far as latitude 37° in the Mississippi Valley
near which region the sub-family Phalangiine appears. to
attain its maximum development.
(4.) The proportionate increase in the length of the legs to
the southward appears to be greater than that of the body.
These last two propositions are true of all the harvest-
spiders having an extended northern and southern distribution
studied with reference to their variations, viz; Liobunum
vittatum, L. ventricosum, L. politum and L. longipes. Probably
the most important factor in determining this increase in size
is to be found in the climatic conditions which permit a
longer period of growth and feeding at the south than in the
north. The arachnids undergo no definite series of molts,
simply casting their skins as increase in size requires. Such
climatic conditions, combined presumably with a more
abundant food supply, may tend, apparently, to increase the
size of these harvest-spiders by at least three methods:
(a.) By the direct effect of a long warm period of growth
with abundant food upon the individual before attaining its
full size.
(b.) By the effect of a long period of development of the eggs
in the ovaries of the female.
(c.) By the action of the eliminating principle in natural selec-
tion in fostering those individuals which attain the maximum
size compatible with their environment.
This increase in size of individual may possibly be of bene
fit to the species (1) by enabling it to prey upon larger insects,
thus increasing the source and variety of its food-supply ; (2)
by enabling it better to elude predaceous enemies, or (3) to
catch fleeing prey.
PLATE XIV.
BiG. t,
Fic. 2.
Liobunum ventricosum (Wood.)
1893.] Harvest-Spider and its Variations. 541
(4.) In the plains region of the Northwest (Dakota, Nebraska)
the legs are considerably shorter than in similar latitudes in
New England.
For example, the second legs of a male Liobunum longipes
from Brookings, South Dakota, measure 62 mm., while those
of a Hanover, New Hampshire specimen measure 74 mm.
The case of the Striped Harvest-spider (L. vittatum dorsatum)
is still more striking; the second legs of a New Hampshire
specimen measure 71 mm., while the normal length in South
Dakota seems to be only 35 mm.
In addition to ‘the above the following laws of vertebrate
variation appear to hold good in the Phalangiide.
(5.) “ The maximum physical development of the individual
is attained where the conditions of environment are most
favorable to the life of the species.
(6.) “The most typical or most generalized representatives
of a group are found near its center of distribution, outlying
forms being generally more or less aberrant or specialized. ”
Explanation of Plates.
Plate XIII. Fig. 1—Liobunum ventricosum (Wood), male.
Natural size.
“ 2—Parts of same. Magnified.
“ 2a—Body.
“ 2b.—Eye eminence. Side view.
“ 2¢¿—Eye eminence. Front view.
“ 29d.—Palpus. Side view.
“ 2e—Claw of palpus. Side view.
“ 9f--Maxillary lobe of second pair of legs.
Plate XIV. Fig. 1—L. ventricosum hyemale. Immature.
- Natural size.
“« 2—Parts of same. Magnified.
“ 2a—Body.
“ 2b—Eye eminence. Side view.
“ 9¢—Eye eminence. Front view.
“ 9d—Palpus. Side view.
“ 2e—Claw of palpus. Side view.
542 The American Naturalist. [June,
EDITORIALS.
EDITORS, E. D. COPE AND J. 8. KINGSLEY.
—We have received from the Secretary of the American Philo-
sophical Society, a programme of the exercises on the occasion of the
celebration of the 150th anniversary of the foundation of the Society.
The celebration commences on Monday, May 22d, at 8 P. m., and con-
tinues until Friday the 26th inclusive, and consists of sessions com-
mencing at 11 o’clock a.m., excepting on Monday, when the session
opens at 8 o’clock, P. M. A
The programme is the work of a committee, and was not submitted
to the Society until its terms could not be altered without discourtesy
to the persons who had been invited to participate in it. This much is
due to the Society, since it is, under such circumstances, not responsi-
ble for the committee's work. As to the committee, its work is disap-
pointing, for since there is one member of it who is presumably com-
petent for the work it undertook, he should have been able to so influ-
ence the other members as to have produced a widely different result.
The adoption of such a pro me is to misrepresent the position
which Philadelphia holds in the wide field of labor covered by the
Society, since it does not include any paper or address by any one of
its citizens presenting his own original work in art, science or philosophy.
On the contrary, the best that can be said of the programme is that its
subject-matter, so far as contributed by Americans, consists of those
generalities to which popular assemblies are wont to be treated, and,
perhaps, entertained, if not satisfied. For a Society whose fundamen-
tal object is the “ increase of knowledge,” to occupy its time in plati-
tudinous disquisitions of this kind, displays a misconception of its own
position and, a lowering of the standard of achievement which it is of at
the societies in this city called upon to bear.
When an active Society, such as the one in question, celebrates 40
anniversary of significance, it usually presents to the world some evi
dence of its activity by securing for the occasion the services of such
of its members as are competent so to do. Such occasions are usu Ri
selected for the announcement of the results of their work before a
wider audience which the occasion is expected to attract. A memoris
+
volume of permanent value is the usual result.
1893.] Editorials. 543
As an illustration, in 1880, the Boston Society of Natural History
celebrated the fiftieth anniversary of its foundation. A memorial
volume was issued, which includes the papers read on that occasion,
and it is a monumental work. The same course should have been pur-
sued by the American Philosophical Society. There is no deficiency
of workers or of work. The members of this Society produce original
work in Prehistoric Archeology, Philology and Ancient and Modern
History ; in Chemistry, Geology Biology; in Political Science, and
in pure Philosophy. We emphasize the words original work. That
none of these men and none ‘of this work appear in the programme
will be cause of astonishment both in Philadelphia and out of it.
544 The American Naturalist. [June,
RECENT BOOKS AND PAMPHLETS.
ALLEN, J. A.—The Geographical Distribution of North American Animals, Bull.
Am. Mus. Nat. Hist., Vol. IV, No. 1, Art. XIV, 1892. From the author
Bascock, E. J.—Clays of Economic Value in North Dakota. laine by H. L
Helgesen, State Commissioner of Agriculture and Labor, Bismark, 1892. From the
author.
Bulletin No. 19, Iowa Agricultural Experiment Station, 1892.
Rae me 87, North Carolina Agri. Exp. Station, Nov., 18
, E. W.—On the Structure of the American Pteraspidian, rege
(Claypole) eit remarks on the Family. Extr. Quart. Journ. Geol. . Vol.
ee 1892. From the author
AFERT, E. W.—On the Hinataa z Minerals of High 2 Gravity. Extr.
a Foha Acad. of Sci., Vol. II, 1893. From the a
Duces, Dr. A.—Un Nuevo Ixodideo. Eiti. La Naturaleza, pe ‘ead: S. 2a. From
the author.
DUMELE, E. T.—Notes on the Geology of the Rio Grande. Bull. Geol. Soc. Am.,
Vol. III, EAE í
Do.; Re n the Brown Coal and Lignite of Texas. Character, Formation,
Occurrence en Fuel Uses, 1892, From the Texas Geological Survey.
EA .—Revision of the Species of Coryphodon. Extr. Bull. Am. Mus. Nat.
Hist., Vol. IV, No. 1, Art. XII. From the author
EVERMANN, B. W.—Description of a New Sucker, Pantosteus jordanit, from the
Upper Missouri Basin. Extr. Bull. U. S. Fish Com., 1892. Date of pub., Jan. 27,
1893. From the author.
FARQUHAR, H.—Competition and oo in Nature. Extr. Proceeds. A. A
A. S., Vol. XLI, 1892. From the a
FRAZER, P.—Biographical Sketch P ioni Sterry Hunt. Extr. Am. Geol., Jan.,
1893. From the author. f
GALLOWAY, B. T.—Report on the Experiments made in 1891 in the Treatment ©
Plant Diseases. Bull. No. 3. Div. Veg. Path. U. S. Dept. Agri. From th he Sain
sonian Institution,
HATCH gi .—The Ceratops Beds of Converse County, Wyoming. Extr. Am.
Journ. Sci., Vol. XLV, 1893. From the author i
HENDERSON, C. H.—The First Cretaceous Fold of the Alps between the Linth
and the Sihl. D Dissertation Presented to the Philosophical Faculty of the
University of Zürich, for the Degree of Doctor of Philosophy, 1892.
author.
Herrick, C. L.—Additional Notes on the Teleost Brain. Separat-Abdruck Anat
Anzeiger, 1892, No. 13 and 14.
——Notes upon the Histology of the Central Nervous System of Vertebrates. es
arat-Abdruck aus der Festschrift zam siebenzigsten Geburtstage Rudolf Leuckart —
From the author a
Hicks, L. eee elements of Land Sculpture. Extr. Bull. Geol. Soc.
Vol. IV, 1893. From the Society.
From the
1893.] Recent Books and Pamphlets. 545
KINGSLEY, J. S.—The Pike Steed of Limulus. Extr. Journ. Morph. Vol. VII,
1892.—From the author.
UNTZ, G. F.—Precious Stones. Extr. from meas on Mining Industries in the
United States at the Eleventh Census, 1890. New American Meteorites.
——Mineralogical Notes on Brookite, Osha satni and Ruby. Extrs. Am.
Journ. Science, Vol. XLIII, 1892.——B an Garnets. Extr. Trans. Am. Inst.
Mining EIE. as From isi aut ae
Kuntz, G. F. unD E. WEINSCHENK. Maronad. Separat-Abdruck aus
Tochermak’s Mineraog u. soba Mittheilungen, herausgegeb. von F. Becke,
XII, Band 3, Heft Wei rig TSIEN armington, Washington Go., Kansas Aerolite,
——On two Meteoric ro Extrs. Amer. Journ. Sci., Vol. XLIII, 1
Lewis, T. H.—De Soto’s soa in the Chicksaw Country in 1540-41. Extr.
National Mag., Nov., ar The “ Old Fort” of Saline County, Missouri. Extr.
a ERPE iey 1892. From the autho
B. S.—Report on the Shippen and ‘Wetherill Tract, Schuylkill Township,
aaah Co., Pa. Phila., 1893. From the author.
Morris, R. T.—Is Svobuive trying to do away xen the Clitoris? Extr. Am,
Journ. i Vol. XXVI, No. 6, 1892. From the
Ossory, H. F. AND J. L. ianiai N.—Characters of Pian (Marsh), a New
eara from the Lower Miocene. Extr. Bull. Am. Mus. Nat. Hist., Vol. IV,
No. 1, Art. XVIII, 1892.—From the authors
. PouLTON, E. B.—Further Experiments tre the Colors of certain Lepidoptera.
Trans. Ent. Soc. London, 1892, Part IV. From the author.
Proceedings of the American Association for re 2 Kikii of Science, 1892.
— The Thickness of the Devonian and Silurian Rocks of Central New York.
Extr. Bull. Geol. Soc. Am., Vol. IV, 1893. From the Society. .
Putnam, F. W.—Report of the Peabody Museum of American Archaeology and,
Ethnology, 1892.
Ruoaps, S. N.—Hybridism Exemplified in the Genus Colaptes. Extr. Science,
Dec., 1892. i
——Notes on Certain Washington and British Columbia Birds. Extr. The Auk,
Vol. X, Sep From the author.
, W. B.—The Evolution of the Premolar Teeth in the Mammals. Extr.
Hasek Phila. Acad. 1892. From the author.
SCUDDER, S. H.—Some Insects of Special Interest from Florissant Colorado, and
other points, in the Tertiaries of Colorado and Utah. Bull. 93 U. S. Geol. Survey,
From the Smithsonian Institution.
SCHEN Common Errors in the Physical Training, orate and Dress
CK, J.
of Girls. Extr. Trans. p nye Med. Soc., 1892. From the au
» H. Reptile from Welte Vreden, pet ae africanus
preda Extr. Quart. ee Geol. Soc., Nov., 1892. From the author.
SHUFELDT, R. W.—Comparative Notes on the Swifts and Pilot Binds. Extr.
Ibis, Jan., 1893.
—Notes on the American Bittern. Extr. The Auk, Jan., 1893. From the
546 The American Naturalist. [June,
SIEBENROCK, F.—Ueber Wirbelassimilation bei den Saurien. Ann. des K. K.
Naturhistorischen Hofmusems; Separat abdruck aus Band VII, Heft 4, 1892. From
SNYDER, M. B.—Report of Special Correspondence See at the Feb. Meeting
of the Pr Club of Do 1893. From the aut
TOWNSEND, C. H. T.—Catalogue of the described South Keii Species of
Calyptrate PET Extr. Ann N ew York Acad. Sciences, Vol. VII, 1892.
—A Blood-su zy Gnat of the Family fc Extr. Psyche, Vol. VI,
1892.——A Cone al Summary of the Known Larval Food- Habits of the Acalyptrate -
Muscidae; a Deptutnary Grouping of the Described Species of Sapromyza of North
America, with one new species; Notes on North American Tachinidae with descrip-
tions of New Genera and Species; = a on New Mexico Insects. Extrs.
Canadian Entomologist, 1892. From the a
Watcort, C. D.—Notes on the Cam a one of Pennsylvania and Maryland,
from the Susquehanna to the Potomac. Extr. Am. Journ. Science, Vol. XLIV, 1892.
——Preliminary Notes on the Discovery of a ane. Fauna in Silurian (Ordovi-
cian) Strata. Extr. Bull. Geol. Soc. Am., Vol. III, 1892. From the Society,
Warp, L. F F.—-Address before the Section of Economic Science and Statistics, A.
A. A. S., 1892. From the author.
ILCOX, J.—On the Evolution of the Earth and the Heavenly Bodies. Read before
the Delaware Inst. Sci., Dec., 1892. From the author
WINCHELL, H. V.—The Mesabi Iron Range. Extr. Twentieth Annual Rep. Minn,
Surv.
1893.] Recent Literature. 547
RECENT LITERATURE.
Evolution of the Colors of North American Land
Birds.'—This octavo of 361 pages, published by the California
Academy of Sciences, the author Mr. Charles A. Keeler, modestly
calls a study. The views set forth are more or less provisional and
tentative, and are intended to direct research into this new field of
ornithological inquiry. The subject-matter of the essay is preceded by
a discussion of the various views held as to the laws conditioning
evolution, such as inheritance of acquired characters ; the nature of
species ; natural and sexual selection, and isolation, as factors in the
evolution of species. In concluding this rather lengthy introduction,
the author remarks “ that life has evolved in accordance with tolerably
definite and unvarying laws, and that the element of chance, if any
such there be, is a very limited one.”
In Part II the colors of North American birds, are treated of under
the following heads: Modes of plumage changes; General principles
of color in Birds ; The proportion and distribution of the colors in the
North American genera; The pattern of markings; Variation of color
with sex, age and season; The direct influences of the environment ;
Geographical distribution as a factor in the evolution of colors; Orders,
families and genera of North American Birds, considered from the
stand-point of their evolution.
The theory of bird colors which Mr. Keeler seeks to establish is as
follows: Pigment is a chemical composition thrown off from the sys-
tem of the bird, probably as a product of waste, and lodged in the
integument. The chemical substance thus generated varies in different
groups of birds, but is probably generally constant in the species of
one genus, or frequently in an entire family. A certain genus would
thus be capable of generating only a given number of fundamental
colors, but natural and sexual selection by combining and rearranging
this limited assortment can produce a variety of effects. :
Associated with this theory is the Law of Assortment of Pigments ;
that is, that the primitive color is a composite which, when more or less
completely resolved into its component elements gives the specialized
tints of the species or genus. If the system of the bird from which
1Evolution of the Colors af North American Land Birds. Occasional Papers of the
California Academy of Sciences III. By Charles A. Keeler. San Francisco, Jan-
uary, 1893.
548 The American Naturalist. [June,
some of the present highly colored genera were evolved produced
normally two pigments, unless some distributing force were brought to
bear upon them, they would naturally be combined at first. It is only
after long selection that the component colors become apparent. The
examples cited are, as might be expected, among the most highly
specialized genera. The woodpeckers are colored black, white and
scarlet. The combination would produce brown, which is still the
body color of some species. _In the genus Tyrannus the original color
was probably olive green, the special colors are black and yellow:
In the blue jays that are blue in the adult together with black and white, |
the mixture of these colors gives the grays of the young, and of the less
specialized forms, as Perisoreus and Picicorvus. Besides black and
white, most North American genera appear to contain but two fanda-
mental colors, but in the tropics three frequently occur in a single
genus.
In discussing protective coloring, repetitive marks, and recognition
marks, the most recent literature upon this subject is cited and illus-
trated with numerous facts of the author’s own observation.
Having considered the factors that influence the evolution of colors
of North American birds, Mr. Keeler gives a brief account of the
families and genera with an application of the principles discussed.
Mr. Keeler’s review of the factors of organic evolution is comprehen-
sive and intelligent. He appreciates the fundamental difference
between the origin of characters and the selection of characters, though
occasionally he slips into the customary confusion on this point by
ascribing the origin of some color marks to natural selection. T
reviewer takes this opportunity to refer to one misunderstanding into
which hein common with many others has fallen, in his reference to
effort as prior to use in developing structure. He says (p. 75) “All
that can be said to the above [the theory of effort] is that`it may be
true but that it has not yet been demonstrated. -There is indeed a b
difference between the assumption that use can modify a part which a
already exists, and the assumption that desire or effort can originate
something which does not exist. Moreover, evem if effort be a vañd
factor in creation, it cannot it seems to me have the general applica- a
tion ascribed to it by Professor Cope. For example, it could appa
ently have no influence on the origination of new colors. Does —
bird desire to be protectively colored? If so it must decide what cok
ors would be most in harmony with its surroundings, and then make -
an effort of will to have these colors developed ; all of which is on the
face of it inconceivable. Or by what imaginable sort of effort cm”
ee eee it Pe ee
pon
A z
RS ee Se ee ae
1893.] Recent Literature. 549
feathers be originated? Effort then, if it can be shown to have any
creative power, must be relegated to a very special field, and cannot
be considered as the sole or even principal originator of the fittest.”
The above paragraph indicates a radical misconception of the
proposition.
Effort is simply the conscious preliminary to motion, and motion is
the fundamental efficient cause of the leading modifications of
structure. No knowledge or intention as to the result or effect (final
cause) of the motion is to be supposed. It is motion which changes
the environment of an animal, and which is thus at the bottom of
whatever results from that change, let the immediate efficient agency
be physical, chemical or mechanical. The psychic cause of this effort
isa sensation. Of course in purely reflex acts, effort (which is assumed .
to be conscious by its definition), is wanting, but the hypothesis
sustained in the work quoted by Mr. Keeler, (The Origin of the
Fittest), is that reflex acts have had their origin in conscious acts,
and are the result of automatization, which is the ordinary process of
education. Reflex motions then have had their origin in effort as
well as the so-called voluntary acts, but at an earlier period. For
this reason they have relatively little to do with the molar move-
ments of animals at the present time, and therefore little to do with
the present origin of specific characters. It is present effort which
precedes most of the motions of animals, and which thus has everything
to do with the environment, to which evolutionists of all shades appeal
as an efficient cause. es
The most important contribution towards the discovery of the origin
of colors in birds by Mr. Keeler, is his demonstration of the law of the
Assortment of Pigments. His classification of our birds in accordance
with their color relations, is a valuable preliminary to further research.
‘When we reach the final stage of the subject, the origin of the tints
themselves, no definite progress is made in the book before us. As one
of the most difficult problems in organic molecular physics, it requires a
very special mode of experimental treatment, and one which Mr.
Keeler has not attempted. The origin of color patterns is less difficult
of approach, and some progress has been made in this direction, but
the subject is yet in a very primitive stage. i
The illustrations of the book are numerous and often in colors, and
they add greatly toits value. The work is gotten up ın excellent
style, and is a credit to all concerned in its publication.
550 The American Naturalist. [June,
Wright’s Man and the Glacial Period.'—This book is the best
synopsis of present knowledge of the glacial epochs and its relations to
human history which has yet appeared. The compass of the work
necessarily does not permit as great detail as would he appropriate to a
technical monograph, but it is admirably adapted for the purpose for
which it is designed, i. e., that of giving the greatest amount of informa-
tion in a readable form in the smallest space. The treatment of dis-
puted topics is generally judicial, and the author has brought to bear
on the subject a great wealth of facts not only from all published
sources, but also from his own original research in North America
and Europe. A question of much general interest is that of the
age of the great ice period. He brings together evidences from
various observers to show that its close cannot have been more than
15,000 years ago, and that its duration may have been twice as long.
The basis of this estimate isthe rate of cutting of various post-glacial
gorges, of which well-known examples are that of the Niagara River,
and that of the Mississippi below the falls of St. Anthony. This
shortened time is in remarkable contrast to the estimate made by the
geologists who first attacked the problem.
The portion of the book relating to the antiquity of man is the
smaller half, but the conclusive evidences of man’s existence during the
glacial epoch are necessarily local. Such evidence as this is handled
judiciously, and all objections are duly considered. Professor Wright
is of the opinion that some of the finds which indicate the existence of
man during the glacial epoch are trustworthy evidence to that effect.
He cites especially as American localities, Trenton, New Jersey,
(Abbott) ; Newcomerstown and Madisonville, Ohio; Little Falls, Min-
nesota, Miss Babbitt; Nampa, Idaho, and Calaveras Co., California.
These finds will be mentioned again below. He considers the supposed
finds of human implements in beds of Neocene age as not established.
Dr. Wright’s book has been made the object of a vigorous attack by
the geologists of the U. S. Geological Survey in a way which shows an
animus on their part not strictly scientific. President Chamberlin 1n
the Chicago Dial, charged the author with improperly alleging on the
title page that he was an assistant on the U. S. Geological Survey. To
this Dr. Wright replied that he was so employed at the time the book
assistant. One of the other criticisms was regrettably free from the
amenities which should characterize scientific discussion, while others —
*Man and the Glacial Period by G. Frederick Wright, D.D. LL.D. International i
Scientific Series No. LXIX. New York, D. Appleton & Co., 1892.
was written and demonstrated satisfactorily his right to use the title ae
Se ae Bog ay eS oy RI 2 rth OS eR
1893.] Recent Literature. 551
exaggerated unimportant details, and ignored the general value and
utility of the book. To all this Dr. Wright has replied temperately and
convincingly.
The ethnologists of the Bureau at Washington have made destruc-
tive criticisms of the evidence for glacial man contained in the book.
Probably the most expert makers in the world of human imple-
ments of the stone age are Messrs Holmes and Maguire of Washing-
ton. They show convincingly that it is easier to make neolithic or
pecked and polished stone implements, than to make fine chipped
flints of paleolithic type. Hence they conclude that either the order
of age should be reversed, or that paleoliths and neoliths are of con-
Fic. 1;
Fig. 1. Argillite implement found by Dr. C. C. Abbott, March, 1879 at K. K.
Rowan’s farm, Trenton, N. J., in gravel 16 feet from surface. From G. F. Wright’s
Man and the Glacial Period.
temporary age, and that the absence of neolithic implements from
some deposits is simply due to accident or to the soft material of which
such implements were made. This conclusion, if correct, revolutionizes
prehistoric archeology. These gentlemen think that it should be
552 The American Naturalist. [June,
revolutionized, and that paleolithic man in both Europe and North
America is a myth. The great collections of paleoliths of the
turtle-back and Chelléen types they look upon as cores and rejects of
pieces from which better implements have been made and taken away.
This view leads them to look with suspicion on the alleged discoveries
of glacial man, and Mr. Holmes has accordingly written articles dis-
crediting the finds described in Dr. Wright’s book.
It may be remarked apropos of the observations of Messrs
Holmes and Maguire, that though it may be true that pecked and
ground implements are more easily made than well chipped flints
their actual relations in time can only be ascertained by stratigraphic
and paleontologic research. A flint broken once or twice so as to pro-
duce an edge is more easily made than a neolith, and gives a great
deal better edge, so that such implements may very probably have
antedated the latter, while the finer ones are well-known to have been
neolithic, and have been made up to the present day. The question
is however, not which implement ought to have come first, but which
actually did come first.
As regards the finds in Europe, those of the caves are the result of
so much careful investigation, and are characterized by such satisfac-
tory stratigraphic conditions, that they cannot be impeached by obser-
vations made in this country. The paleoliths and human bones
have been conclusively shown to belong to the age of the glacial
fauna. In North America the paleontologic evidence is not 50
good, but such as there is, indicates strongly that the earliest known
American was not more modern than the paleolithic European.
Those who saw the Calaveras skull when first found, allege that it was
more or less covered with the adherent cement so characteristic of the
gold bearing gravel of California. The age of this gravel is not
exactly determinable, since data respecting the finding of fossils in it
are not generally reliable. But that it is of approximately glacial age no
one doubts. Mr. Holmes believes that the implements of the Abbott and
Babbitt finds occur only in the talus, and are not from the undisturbed
glacial gravels (American Geologist) ; but so far as regards part
Babbitt, and all of the Abbott finds, other observers hold a different
opinion. In the Journal of Geology he shows that the evidence for the — 4
stratigraphic position of the finds at Madisonville and Newcomerstown,
Ohio, is defective. It may be added here that the Nampa image, .
whatever may be its real stratigraphic origin, displays in its form 9
artistic skill on the part of its maker, not to be looked for in primitive ©
man ; nevertheless it is time that the name of the person who ates™
of the
Werte Nia on sa eee ee
Pr pees Tea gs aoe ea
1893.] Recent Literature. 553
FIG. 2.
Fig. 2. Smaller figure, paleolith from Newcom wn, Ohio; larger figure do
from Amiens, France; both one-half size. From G. F. etn s Man and the Glacial
Period.
that he intentionally deceived Professor Wright in this matter, be pro-
duced ; and the authority for the statement that such an assertion was
made, should make himself known. The image cannot well be the
work of any existing Indian tribe, as has been asserted. In any case it
seems that the evidence for Plistocene man in America, must be
further investigated with careful methods, and under more favorable
circumstances than are furnished by most of the so-called glacial
gravels,
We give figures of two characteristic types of paleoliths; one of
argillite found by Abbott at Trenton, N. J.; and one flint from
Newcomerstown, Ohio. E. D. COPE.
554 The American Naturalist. [June,
Some Recent Books on Bacteriology.—One of the most
recent works dealing with the bacteria is Dr. Sternberg’s' Manual of
Bacteriology. He is well known as an investigator in this field, and
is moreover one of the pioneers in the United States. The work isa
most comprehensive one, the best published in the English language.
While perhaps not so full on pathogenic organisms as Baumgarten’s
Lehrbuch der Pathologischen Mycologie, yet it is sufficiently full and
complete to make it one of the best books of its kind. It moreover
contains accounts of the most recent discoveries in regard to pathogenic
organisms. :
One hundred and one pages are devoted to history, classification,
morphology and general bacteriological technology. The author
adopts the classification of Baumgarten (1890) with slight modifica-
tions. He divides the species into (*) relatively monomorphous and
(?) pleomorphous. Under the first micrococci, bacilli, and spirilla are
grouped ; under the second spirulina of Hueppe, leptotrichee (Zopf)
and cladotrichee. Hauser’s Proteus also belongs here. Part second
is devoted to general biological characters; including an account of
the action of antiseptics and germicides. Dr. Sternberg’s work on
antiseptics and germicides is so well known that it will not be necessary
to refer to this part of the work at length. Three hundred pages are
devoted to pathogenic bacteria, 158 species are carefully described, and
many are illustrated. In some cases the colors of the growth when
grown in different nutrient media are given. Botanists will be chiefly
interested in the account given of the saprophytic bacteria. Chapters
are devoted to the bacteria of the air and water, bacteria found in the
stomach and intestinal canal, articles of food and soil, in which the
usual methods of culture are given. Three hundred and thirty one
species are described. This feature of the book makes it especially
commendable. Those who have had occasion to use the classification
of De Toni in Saccardo’s Sylloge Fungorum or that of Crookshank in
his Manual of Bacteriology have found the descriptions of the species
far from satisfactory. The descriptions given by Sternberg are ample
for the determination of the species. ;
The biological characters are very full. An excellent bacterio-
logical diagnosis is a great aid in the determination of the species.
few errors have made their way into this part of the book. On page
758, Bacillus citreus-cadaveris is said to be motile, but turning to pag?
7A Manual of Bacteriology. Illustrated by heliotype and chromolithographi¢
plates and two hundred and sixty-eight engravings, 886 pp. New York, Wm.
and Co., 1892.
ieee
oe
i :
ae
A
ae
pss
ea
ate
X
1893.] Recent Literature. 555
633 where the species are described it is said to be non-motile. The
same for Bacillus fulvus, which p. 758, is said to be motile, but on p. 629
is described as non-motile. Notwithstanding that a few of these defects
. occur, this part will certainly be appreciated by working bacteriolo-
gists. In this key the following characters are used. (1) Morphologic;
Micrococci, Bacilli, Spirilla, Leptotricheze and Cladotrichee. Forma-
tion of spores, independent motion. (2) Physiological characters ;
relation to oxygen, xrobic, strictly anwrobic, facultative anzrobic,
growth in gelatin, liquefy or do not, no growth in gelatin, growth on
potato, in milk, coagulate milk, do not coagulate milk, color of growth,
chromogenes, not chromogenic, pathogenesis.
The bibliography has been gotten together with great care, and is
very full, containing the titles of 2,582 papers grouped under historical
classification, staining methods, culture media, sterilization of culture
media, etc. A very large number of papers are cited on the physiolog-
ical properties, such as lactic acid fermentation, viscous fermentation,
putrefaction, ete. The literature on pathogenic microorganisms is,
especially full. Bacteriologists should feel grateful to Dr. Sternberg
for this work, and also to the publishers for the excellent manner in
which their part also has been performed.
Wm. Wood & Co. have during the past four years issued two other
text books on bacteriology; the English translation of Salomonsen’s
Bacteriological Technology, by Professor Trelease is well known and
needs no introduction. A second admirable work, antedating Stern-
berg’s Manual, is Frinkel’s Grundriss der Bakterienkunde translated
by J. H. Linsley.? It has been translated into six different languages,
and those who have been fortunate enough to use the German edition
know its admirable qualities. The translation before us is an admir-
able one. The style is clear and there is no difficulty in understanding
the author. The work contains a great deal on the biology of many
species that hitherto has not been found its way into English works.
It is to be regretted however that references to literature are not given,
but these may readily be obtained from Sternberg’s Manual. No
figures are given in the book, but the accounts are clear. Future
editions should have figures.
For a limited amount of work both of the books noticed above are
too large. One of the best of the small books with which the writer 1s
familiar is Abbott’s? Principles of Bacteriology.
*Text-book of Bacteriology, third edition, translated and edited by J. H. Linsley,
Pp- 376. New York, Wm. Wood & Co., 1891. :
The Principles of Bacteriology. A manual for students and physicians. Lea
_ Brothers & Co., Philadelphia, 1892, pp. 263
556 The American Naturalist. [June,
This book is well suited for class room work. Its technique is
stated in clear and concise language, accompanied with numerous
illustrations. ‘The writer having used this for a large class is prepared
to say that it is an excellent work of its kind. .
Another hand-book intended for a larger circle of readers is Wood-
head’s' small volume on Bacteria and their Products. It treats the
subject in a somewhat different manner than the others. It gives con-
siderable attention to historical matters, and treats quite fully the
different systems of classification used by Ehrenberg, Cohn, Van
Tiegham, Zopf, De Bary, Hueppe and Fliigge. The chapter on fer-
mentations contains a great deal on chemistry, but it is far from being
a popular exposition of the question. The author has of course largely
drawn from Hansen, Pasteur, and Schiitzenberger. The author uses
the word parasite and saprophyte in a somewhat peculiar way. He
speaks of the bacteria of the mouth.as being parasites, contradicting &
previous statement made with reference to parasitic bacteria. The
work has been edited rather carelessly, the word “ parasiticism ” for
parasitism occurs in several places as well as the word “ saprophyti-
cism. ”—L. H. PAMMEL.
Report on the Fish and Fisheries of the United States
for 1888.°—This volume, an octavo of 902 pages, contains in addition
to the Reports of the Commissioner, Hon. Marshall McDonald and his
two assistants, 11 important papers bearing upon the Fish Industry-
The results of the sea-coast inquiries conducted during the year 1888
are embodied in the reports of Mr. J. W. Collins and Lieutenant
Commander S. L. Tanner. The work plished at the laboratory of
Woods Holl, Massachusetts is reported on by Mr. J. A. Ryder. Notes
on Entozoa of Marine Fishes, with descriptions of new species and the -
anatomy of Thysanocephalum crispum Linton are given by Mr. E
n.
Number 8 in the series of Appendices is a review of the Fresh-water
Sunfishes of North America by Mr. C. H. Bollman. Itis the beg
ning of the systematic investigation of interior waters by volan oe 2
naturalists under the direction of Dr. D. S. Jordan. The Apoda
Fishes inhabiting the waters of America and Europe are re iewed by
*Bacteria and their Products. The contemporary Science series, edited by ee :
lock Ellis, pp. 459, with 20 photo-micographs. London, Walter Scott, 24 Want
Lane, Charles Scribner Sons, New York, 1892.
SU. S. Commission of Fish and Fisheries. Part XVI. Report of the commission
for 1888. Washington, 1892. n
1893.] Recent Literature. 557
Dr. Jordan and Mr. B. M. Davies. The work undertaken by Prof.
W.O. Atwater ten years ago, viz.,a comprehensive series of experi-
ments upon the chemical composition and nutritive values of the
American food-fishes, has been completed, and his report upon the sub-
ject is published as Appendix 10 to this volume. The last paper is J.
W. Collins’ Report upon the participation of the U. S. Fish Commission
in the Centennial Exposition held at Cincinnati, Ohio, in 1888.
The numerous illustrations and maps give additional interest to this
comprehensive report.
37
ee i A
eae
fee
558 The American Naturalist. [June,
General Notes.
aa a:
pales a te eee Se a a a y N
GEOLOGY AND PALEONTOLOGY.
Oneonta and Chemung Formations in Eastern Central
New York.—The Oneonta formation comprises a thick mass of red
shales and red and gray sandstones, similar in character to the rocks
of the Catskill Mountains. They overlie the Chemung formation in
southern New York and northern Pennsylvania. The recent investi-
gations of Mr. N. H. Darton confirm Professor Hall’s supposition that
the Oneonta beds represent the eastern extension of the Portage forma-
tion. The former is characterized by a large quantity of red shale
which occurs as streaks in the lower beds, a thick mass a little higher
in the series, and constitutes elongated lenses in the gray flags and red
sandstones of the upper member of the formation. Toward the western
termination of the Oneonta formation the red material rapidly disap-
pears and its place is taken by gray shales and thin bedded sandstone,
There can be no doubt of the continuity of sedimentation throughout.
Toward the east the formation increases in thickness until it comprises
the lower thousand feet of beds in the Catskill Mountains. The Che-
mung fossiliferous shales which overlie the Oneonta formation south of
Franklin grade upward through a series of flags into hard, coarse,
cross-bedded gray sandstones with intercalated red shale layers.
Toward the east the fossiliferous shales merge into flags, and then "n
hard, coarse sandstone with flaggy layers along the eastern front of the
Catskill Mountains.
Since the rocks of the Catskill Mountains comprise the Chemung
and Portage horizons, Mr. Darton proposes to discard the usé of
Catskill to designate a formation, and to use the term Catskill group s
include the Chemung and Portage formations. (Am. Journ. Science,
March, 1893.)
ee Pa) TENE SE el OAN
Tertiary Insects from Colorado and Utah.—Bulletin No. 93
of the Geological Survey comprises descriptions of eight spe o i
Oli insects from Florissant and other points in Colorado mé
Utah, by Samuel H. Scudder. sa
Of this group of interesting fossils six are referred to new ge" a
In a short introduction the author gives the special claim which each —
1893.] Geology and Paleontology. 559
has for consideration. The Trichocemsis represents a type hitherto
considered exclusively gerontogeic; the Stenogomphus is the first
Gomphine fossil found in this country ; the Cicada for its great size
and for being the first member of its family known from American
rocks; the Hymenoptera is curiously related to Oriental forms; the
Diptera are interesting departures from the modern types to which
they are most nearly allied ; the two Coleoptera present some curious
features ; the butterfly is of exceptional interest as belonging to a
waning type which probably flourished remarkably in Oligocene times,
if the published figures are to be regarded as having any weight at all.
A Supposed New Order of Gigantic Fossils from
Nebraska.—In “ University Studies,” published by the University of
Nebraska, 1893, Mr. E. H. Barbour describes and figures a number of
gigantic mineral bodies of such anomalous form and structure, that the
author offers as a merely provisional classification until their place can
be more definitely determined.
These fossils are found in the Miocene beds which follow the divide
between the White and Niobrara rivers in Sioux County, Nebraska.
They are very abundant and increase in size toward the southern lmit
of the beds. In appearance they resemble colossal corkscrews varying
from two to nine feet in length. A transverse piece corresponding
to the handle of the corkscrew, for which the term rhizome is adopted
by the author, is often three feet in diameter. The fossil corkscrew
is invariably vertical, while the so-called rhizome as invariably curves
rapidly upward and extends outward an indefinite distance. Some of
the screws coil about an axis, others are unsupported. The author is
positive as to their organic origin, and is inclined to believe them to be
_ sponges. Fiye microscopic slides show certain smooth spindle-shaped
rods suggestive of sponge spicules, but a sixth section reveals unmis-
takable plant cells. A third evidence of their organic nature is the
characteristic intricate network of minute silicious tubes in the stems.
To add to the difficulty of the problem, a well-preserved skeleton of a
rodent was found in the great stem of one specimen. This rodent is
the size of a “jack rabbit,” with proportionately large incisors, sagittal
and occipital crests high and sharp, and a mole-like shoulder girdle.
In order to preserve the appropriate name—* Devil’s Corkscrew —
bestowed by the ranchmen, Mr. Barbour calls these strange fossil forms
Daimonelix.
The most probable explanation of
are the casts of the burrows of some large roden
. tion will then be the entrance; the enlargemen
these objects seems to be that they
t. The horizontal por-
ts the position of the
560 The American Naturalist. [June,
nests, and the spiral vertical portions shafts for safety or escape of the
occupants, or for the admission of air; the spiral being necessary for
the convenient ascent of the animal. It is well known that species of
Thomomys make spiral burrows; and remains of this genus are not
rare in the Plistocene beds of Kansas.—C.
Mammalia from the ‘“‘ Pits of Gargas.’’—In the grotto of
Gargas, not far from Montréjean, a number of pits have been found
about 100 metres from the entrance. These pits are 20 metres in
depth and have mouths so narrow that it is with difficulty that a man
ean force himself through the openings. M. Regnault has recently
explored these pits and found them rich in fossil remains of bears
( Ursus speleus, the small variety), wolves (Canis lupus), and hyaenas
(Hyaena crocuta). M. Gaudry notes the unusual circumstance of the
fine state of preservation of the fossils, particularly of a hyaena and a
wolf, of which almost complete skeletons were obtained. These finds
have been made the subject of a joint paper by MM. Gaudry and Boule,
in which they discuss the affinities of these cave animals, and give
tabular statements of the genealogy of the bears and of the hyaenas.
The latter differs from the one published by Schlosser in 1890.
A series of fine plates illustrates the paper. (Materiaux Pour.
P Histoire des Temps Quaternaires. Quatrième Fascicle, Paris, 1892.)
Mr. G. F. Matthew reports a new genus, Protolemus, of Trilobites
from the St. John group of the lower Cambrian beds. The new genus
is represented by two species, P. elegans and P. parodozoides. (Bull.
New Brunswick Nat. Hist. Soc., 1892.) ee
The Cleveland shale of Ohio has yielded a new Coccostean, which is
described by Professor Claypole under the specific name Co
cuyahogae. This species comes from a higher horizon than either C.
hercynius or C. occidentalis, and is remarkable for its large size. (Am.
Geol., March, 1893).
Paleozóic.—Mr., H. E. Sauvage has been studying the fish fauna
of the Permian of France. He finds that of 24 species, 14 are, for the
present, peculiar to France. This fauna is, undoubtedly, that of the
Lower Permian, and is characterized by a predominance of species
belonging to the genera Amblypterus. (Revue Scientifique, Avril,
1893.)
Mesozoic.—According to Chapman, the Phosphatic Chalk of
Taplow, England, has yielded 5 species and varieties of Ostracoda, p
previously known, and 98 species and varieties of Foraminifera, .
which the following are new to science: Nubecularia jonesiana, 40°”
1893. | Geology and Paleontology. 561
laria decurrens, Textularia serrata, Bulimina trigona, and Bolivina
strigillata. (Quart. Journ. Geol. Soc., Nov., 1892).
Cenozoic.—The Pliocene fauna of Rousillon, which now numbers
over 30 species of vertebrata, has recently had a second Proboscidian,
Mastodon borsonii, added to the list of mammalia. The specimen, found
near Villeneuve-de-Raho, consists of the series of upper molars, a tusk,
and a part of the right occipital region. This discovery is important,
as it gives a wide geographical range to M. borsonti, which has not
before been known from any pliocene bed of southern France. (Revue
Scientifique, Mars, 1893.)
Dr. Noetling, in a report on Jade in Upper Burmah, says that jade
is found in association with and enclosed in an eruptive rock closely
resembling serpentine, and that this serpentine pierces strata of perhaps
lower, but more probably Upper Miocene date. The jade found in
Burmah belongs to a group of eruptive rocks of late Tertiary age.
(Nat. Sci., April, 1893.)
The prevalence of lake basins in glaciated countries is accounted for
by Mr. J. C. Hawkshaw by the following conditions:
Earth movements in limited areas tend to form basins; these move-
ments are gradual, and, under ordinary circumstances, the basins are
obliterated by water-borne detritus, growth of vegetation and erosion.
In glaciated regions the basins are not only protected by the ice from
such destructive action, but they are still further deepened by its
grinding action. (Nature, April 13, 1893.)
M. M. Boule has described and figured the Hyaena brevirostris found
by M. Aymard in the Pliocene deposits of Sainzelles near Puy (Haute
Loire). Although this fossil has been referred to by Pomel, Gervais,
Gaudry and Weithofer, no detailed description of it accompani
figures has ever before been given. (Ann. Sci. Nat. Zool. t. XV, 1893.)
According to Mr. R. T. Hill, the Hematite and Martite iron ores of
Mexico occur in the Cretaceous and Tertiary rocks, and are undoubtedly
of later age than the rocks in which they occur. This is an unusual
geologic age for ore. (Am. Journ. Sci., Feb., 1893.) :
Mr. Warren gives, as a result of a comparison between pleistocene
and present ice-sheets, an opinion that the Ice Age was a continuous
and geologically brief period. (Bull. Geol. Soc. Am., Vol. IV j 1893.)
Mr. E. T. Dumble reports the occurrence of Grahamite in two
localities in Texas. The first specimen came from the Fayette beds in
Webb County, the second from the lowest strata of the Eocene exposed
on the Rio Grande River in Fayette County. (Trans. Am. Inst, Mine
Engineers, 1892.)
562 The American Naturalist. [June,
MINERALOGY AND PETROGRAPHY:.
The Petrography of the Abukuma Plateau, Japan.—The
northern half of the Japanese Archean area, the Abukuma Plateau, is
thought by Koto? to consist of a series of Laurentian granites and
“pressure gneisses, cut by younger granites and other eruptives, and
overlying these a series of schists, divided by the author into lower and
upper Huronian. The Laurentian granitic are an older amphibole-
biotite variety and a younger, intruding biotite granite. The former
contains, in addition to the usual granite components, microcline,
and a bluish-green, weakly pleochroic hornblende, that very frequently
plays the role of an ophitic groundmass for the other constituents.
This granite passes by dynamo-metamorphism into foliated phases, in
which the various minerals have been compressed, and the quartz, in
addition, granulated. The Huronian (?) beds are principally schists
and gneisses, that differ from the Laurentian gneisses in having the
plane parallel structure, i. e., they are composed of bands of different
composition. The most important schists of the lower division are:
gneissic mica schists, containing andalusite and sillimanite, two-mica
schists, one of whose constituents is margarite, garnet-biotite schist and
hornblende schist. A peculiar member of the series is a titanite
amphibole schist, consisting of bands whose structure is granular. Its
black bands are made up of green-hornblende, plagioclase and a little
biotite, and its white ones of sphene and granular sahlite in a ground-
mass of altered feldspar. The upper Huronian seriesembraces foliated
amphibolites, mica-schists, and green schists that may be tufas. The
distinction between the lower and upper members of the group seems
to be based mainly upon petrographical characteristics. Among the
rocks cutting these various schists may be mentioned an amphibole-
picrite, pegmatites, and several varieties of diorite-porphyrite.
The Leucite-Tephrite of Hussak, from New J ersey.—The
eleolite syenite eruption of Beemerville, N. J., was accompanied by
basic extrusions now represented by the smaller dykes associated with
the large eleolite-syenite dyke in this region. One of the most interest-
ing of the basic dykes is the one at Hamburg, Sussex Co. It is from
15 to 20 feet wide and consists of a dark, tough, biotite rock, holding
‘Edited by Dr. W. S. Bayley, Colby University, Waterville, Maine.
? Jour. Coll. Science, Imperial University, Japan, v, 3., p. 197.
1893.] Mineralogy and Petrography. 563
spheroidal inclusions, that have been taken for Hussak* to be leucites.
Kemp‘ has recently examined this rock very carefully, and now
describes it as composed of biotite and pyroxene imbedded in an iso-
tropic groundmass that is chiefly analeite. The biotite is dark brown
and the pyroxene of a faint yellow color, with an extinction of 33°.
The spheroidal inclusions are analcites, about whose ruins are often
grouped grains of biotite and crystals of sphene. An analysis of one
of the spheroids, after deducting 3.886 % of Ca Co,, gave:
HO SiO, 1,0, Fe0, Ca0 KO Nao
631 5244 2644 43 194 854 890
As to the origin of the analcite the author is not certain. It may
have been derived either from leucite, in which case the rock would be
a leucite-tephrite, as considered by Hussak, or it may be alteration
product of nepheline.
A Sodalite-Syenite from Montana.—In the mountains form-
ing the northern portion of Montana, Lindgren® and Melville have dis-
covered post-cretaceous quartz-porphyrites, lamprophyres, augite-
trachytes, analcite basalts, and a peculiar sodalite-syenite, somewhat
resembling certain rocks described by Chrustschoff from Russia. The
Montana syenite is from Square Butte, situated thirty miles southeast
of Fort Benton. It is a light gray eruptive, associated with sheets of
theralite and analcite basalt. Macroscopically it consists of lath-
shaped feldspars, prisms of hornblende and pale brown grains of soda-
lite. In addition, analcite and plagioclase are discoverable under the
microscope. Many of the feldspar crystals are corroded in an extraor-
dinary manner and the cavities thus formed in them are filled with
-analcite which is believed to be an alteration product of albite. The
hornblende is very dark brown, almost opaque, with a strong pleo-
chroism, an extinction of 13° and a density of 3.437. Its analysis
indicates its identity with the variety barkevikite : °
H,O SiO, Al,0,(TiO,) Fe,0, FeO NiO MnO CaO MgO Na,O K,O
24 38.41 17.65 3.75 21.75 tr 19 10.52 2.54 2.95 1.95
The sodalite is quite fresh. It forms irregular grains that are bounded
by faces when in contact with analcite. It was evidently
formed after the feldspar but before the analcite. The composition of
3 AMERICAN NATURALIST., March, 1893, p- 274.
t Amer. Journ. Sci., Apr., 1893, p. 298.
5 Amer. Journ. Sci., June, 1893, p- 286.
6 AMERICAN NATURALIST, June, 1890, p. 576.
564 The American Naturalist. [June,
the rock as calculated from its analysis is: 23% hornblende, 50% ortho- r
clase, 16% albite, 8% sodalite, and 3% analcite.
The Anorthosites of Canada.—The Canadian geologists have
long considered the Laurentian of northern North America as consist-
ing of an upper and a lower division, of which the latter rests uncon-
formably upon the former. This upper division is made up largely of
basic schists to which the name Novian was given by Hunt. Adams
has examined all of the important occurrences of the supposed schists,
and has discovered that in all cases they show an irruptive contact
with the surrounding gneisses, which they evidently cut. They are
thus unquestionably post-Laurentian, and, from their relations to the
overlying rocks, they are thought to be pre-Cambrian. The dark
rocks are anorthosites—aggregates of plagioclase, with a little pyroxene,
olivine and some accessories—which are in places schistose, and in
other places are connected genetically with gabbros. The schistosity
of the rock is accompanied by the possession of cataclastic structure,
regarded by the author as due not to dynamic processes, but to the
movement of the magma just before final consolidation. The plagio-
clase of the rock which is by far its most prominent component, 18 &
labradorite so filled with tiny inclusions of microlites, thought by the
author to be ilmenite tables, that fragments of the mineral are dar
and often show the play of colors so beautifully seen in the labradorite
of Labrador. The pyroxenes are a weakly pleochroic green augite,
and a strongly pleochroic hypersthene. Hornblende, biotite, quartz,
garnet and zircon are also present in small quantities in all specimens
of the anorthosite. In the Saguenay river occurrence, olivine E
enclosed in the plagioclase, and between it and the latter mineral is &
reaction rim, composed of an inner zone of hypersthene, and an outer
one of actinolite, including many small, green spinels. All the occur-
rences of the rock in Canada are briefly described, and with them arè
compared similar occurrences found elsewhere.
The Melibocus “ Massiv ” and its Dyke Rocks.—The peak
of Melibocus’ in the Odenwald consists mainly of a medium- ag
white granite to the West, and a complex of schists and gneisses t0 p A
East. The granitic constituents, orthoclase and quartz, are Usua =
aggregates of small grains variously orientated, and the biotite ge a
evidence of having been subjected to pressure. Near the contact WE?
* Neues Jahrb. f. Min., etc., B. B. viii, p. 419.
3C. Chelius: Notizbl. d. Ver. f. Erdk. z. Darmstadt, 1892, iv, F. 13 H., p.t
1893.] ; Mineralogy and Petrography. 565
the surrounding rocks the granite becomes gneissic, and everywhere it
is cut by dykes of aplites, porphyries and lamprophyres. Where the
aptites penetrate the gneisses they possess the usual characteristics of
these rocks, but where they pass from the schists into the granite they
become porphyritic, showing a fine grained groundmass of quartz,
orthoclase and mica and numerous phenocrysts of the same minerals
and garnet. Like the granite the aplite components exhibit evidences
of the effect of pressure. The large crystals are granulated and the
rock’s structure is more or less schistose. For this aplitic rock with
porphyritic crystals the author, Chelius, used the name Alsbachite.
An analysis of an alsbachite from the northwest side of the mountain
= gave:
SiO, AlO, Fe,0, FeO MnO CaO MgO K,O NaO H,O
7413 12.61 287 86 16. 1.60, 28, 213 .456. 6
The dioritic aplites, malchite, luciite and orbite are also represented
among these dyke rocks—the malchite being the panidiomorphic
diorite aplite, the luciite the hypidiomorphic granular forms, and the
orbite the corresponding porphyritic phases. One of the luciites is
described as made up almost exclusively of plagioclase and horn-
blende. Among the lamprophyric dykes, mention is made of a
gabbrophyre, or odinite, which differs from the gabbro-aplite, beer-
bachite, in consisting of phenocrysts of plagioclase and colorless augite
in a matrix of plagioclase laths and hornblende needles, while the
aplite is a panidiomorphic aggregate of diallage and feldspar, with the
addition, sometimes, of hornblende crystals that enclose the other con-
stituents. The descriptions of all these rare rocks are very brief.
of 185 hand specimens of stock and dyke granites and pegmatites, cuts
ting the archean and paleozoic beds of Argentina, and of younger
granites cutting these older ones,
the origin of the micropegmatitic
exhaustive, it contains no points of spe
of the paper is a list of the specimens exa
° Neues Jahrb. J. Min., etc. B. B. viii, p. 275.
mined, with their localities,
566 The American Naturalist. [June,
and appended to it are twelve plates containing seventy-two micro-pho-
tographic reproductions of their sections.
New Minerals.— Geikielite.°—This mineral was found as pebbles
in the gem washings near Rakwana, Ceylon. It is essentially a mag-
nesium titanate, MgTiO,, corresponding to the calcium compound,
perofskite. The mineral is bluish-black and opaque, with a brilliant
lustre, and possessing two cleavages at right angles to each other. Its
density is 3.98 and hardness 6.5. In thin section it is translucent
with a purplish-red tint, and in converged light it shows a uniaxial
re.
Baddeleyite,” also occurring as pebbles in the above-mentioned
locality, is a black substance with a density of 6.02 and a hardness of
6.5, thus strongly resembling columbite. Under the microscope small
fragments are seen to be dichroic in greenish-yellow and brown tints,
and to possess a biaxial symmetry. The crystallization is thought to
be monoclinic, though only a few plans could be detected on the speci-
men. In chemical composition the substance is zirconia ZrO,.
Folgerite, blueite and whartonite are all nickel-iron-sulphides from
the Sudbury nickel mines at Algoma, Ontario. Emmens” describes
the first named as a massive, bronze-yellow substance, with a grayish-
black streak, a density of 4.73 and hardness 3.5. Its composition (Fe
= 33.70; Ni = 35.20; S = 31.10) corresponds to Ni Fe $,
The blueite is also massive. Its color’is olive-gray or bronze ; its
streak black, density 4.2 and hardness 3-3.5. Its analysis yielded Fe
= 41.01; Ni = 3.70; S = 55.29, corresponding to pyrite with a
thirteenth of the Fe replaced by Ni. Unlike pyrite, however, it dis-
solves easily in nitric acid, without the precipitation of sulphur.
Whartonite differs from blueite in containing more Ni. Its composi-
tion is Fe = 41.44; Ni — 6.27; S= 52.29, corresponding to (Fe Ni)
8, in which Fe: Ni=7:1. Its hardness is 4, density 3.73, and color
and streak like those of blueite.
Hauchecornite is another nickel mineral. It is described by Scheibe”
from the Friederich mine in the Hamm mining district, Germany. It
is found in bronze-yellow tetragonal crystals, with a hardness of 5, and
4 density of 6.4. It is thought to have the composition corresponding
to the formula Ni (Bi. Sb. S), though analysis yields discordant results.
Cuprocassiterite was described by Ulke” from the Etta mine, South
1 Fletcher: Nature, Oct. 27, 1892, p. 620.
11 Jour. Amer. Chem. Soc., Vol. xiv, No. 7.
12 Jahrb. A. preuss. geol. Landeranst, 1891, p. 91.
13 Proc. Amer. Inst. Min. Engineers, Feb. 1892.
1893.] Mineralogy and Petrography. 567
Dakota, but the author’s data were so scanty that Headdon™ has
thought it advisable to add a small additional contribution -to the
literature of the mineral. This last-named writer obtained a small
quantity of what he supposed to be Ulke’s new mineral from both the
Etta and Peerless mines, and found upon examination that in the
interior of a small mass from the Peerless mine is a nucleus of stannite
containing a little cadmium. Intergrown with this and also forming
an envelope around it is a green clayey substance, which, upon its
exterior, passes into a yellow earth. The green substance has a density
of 3.312-3.374. Its analysis shows it to be a mixture of about 7 SnO,,
6 CuO, 2 FeO and 11 H,O. The author regards it is an alteration
product of stannite, but not as a well-defined mineral species.
New Edition of Rosenbusch’s Volume on Minerals.—The
new edition of Professor Rosenbusch’s Microscopic Physiography of
the Rock-forming Minerals is an enlargement rather than a revision of
the second edition. There is no material difference in the arrangement
of the matter in the two editions, but there have been large additions
made in the later volume in the shape of descriptions of new petro-
graphical apparatus and methods, and in the number of minerals
treated. The plates illustrating the text have been decreased by one.
The remainder are much better executed than was the case in the
earlier volume.
Mineral Syntheses.—Michel” has obtained melanite garnets and
sphene crystals by cooling slowly a mixture of 10 parts titanic iron, 10
parts calcium sulphide, 8 parts silica and 2 parts carbon, that had
been heated to 1200° for five hours.
Crystallized leucite, potassium eryolite and potassium nepheline results
from the fusion of silica or of fluosilicate of potassium and alumina
with an excess of fluoride of potassium. Prolonged heating produces
leucite, and potassium cryolite. Less prolonged treatment yields
a potassium nepheline, which crystallizes in negative orthorhombic
prisms.”,
Instruments.—For measuring the curves of isotherms on mineral
plates Jannetaz® has constructed a new ellipsometer, which it is
l4 : Wee Š 105. ;
15 ae Sea Oe eee Physiographie der petrographisch wichtigen
Mineralien, Stuttgart, 1892, pp. 712, Fig. 239, ete.
18 Bull. Soc. Franc. d. Minn., Vol. xv, p. 237-
568 | The American Naturalist. [sede #
believed will enable its user to measure accurately the axes of the
isothermal ellipses, and to determine rapidly in each case whether
apparently circular isotherms are in reality circles or slightly eccentric
ellipses.
A new machine for cutting and grinding thin sections of rocks and
minerals, with stored electricity as the motive power, is described by
G. H. Williams”
Rock Separations.—Thallium-silver-nitrate [Tl Ag (NO,) .],
according to Retgers,” is an excellent medium for the separation of
mineral grains of great density. The double salt fuses at 75°, and in
the fused condition is clear and mobile. In this condition its specific —
gravity is 5, and this may easily be lowered by the addition of water.
Its manipulation is simple. A small beaker containing the solidfsalt
is placed in a water bath and heated. Upon its liquefaction the pow-
der to be separated is added and the mixture is allowed to stand for a
short time. As soon asa layer of clear liquid forms between the pre-
cipitated and the floating grains the beaker is plunged into cold water.
The salt thus consolidates rapidly. The beaker is now broken and the
heavy grains are collected by scraping and washing.
A new method of separating the constituents of rock powders, whose
densities are above 2.60, has been devised by Dafert and Derby.” The
principle involved is the suspension of small particles in gentle cur-
rents of water. The apparatus necessary for the operation is fully
described by the authors. Separation is not complete between pow-
ders of nearly the same density, but there is a strong concentration of
the heavier and the lighter ingredients in the two resulting portions of
the separated material.
19 Amer. Jour. Sci., Feb. 1893, p. 102.
Neues Jahrb. f. Min., etc., 1898, Vol. I, p. 90.
1 Proc. Roch. Acad. Sci., Vol. II, p. 122.
=
ERR] Botany. 569
BOTANY.
Cæoma nitens.—The development of Cwoma nitens has recently
been studied by Mr. H. M. Richards, and the results published in an
interesting paper in the Proceedings of the American Academy .of
rts and Sciences, The particular question investigated was whether
or not the spermogonia are developed within the cavities of the
epidermal cells of the host. By means of carefully made sections, Mr.
Richards demonstrates that they arise as masses of hyphæ which push
up between the epidermal cells, and that later the walls of some of these
cells become absorbed. The spermogonia are therefore at first inter-
cellular, but by the absorption of the walls they become intracellular.
CHARLES E. BESSEY,
Our Naiads.—Thomas Morong’s monograph “The Naiadacex of
North America ” has been brought out in the Memoirs of the Torrey
Botanical Club. It contains descriptions and plates of 54 species
distributed as follows: Triglochin, 3 species; Scheuchzeria, 1; Lilea,
1; Potamogeton, 37 ; Ruppia, 2; Zannichellia, 1; Naias, 4; Zostera,
3; Phyllospadix, 2. Among these we find one new species Potamoge-
ton faxonit from Lake Champlain, and several new varieties of pre-
viously described species. Quite a number of changes have been made
in the nomenclature of the species. vs
Thus comparing Dr. Morong’s list with that in the 6th edition of
Gray’s “ Manual,” we note the following changes: Potamogeton penn-
sylvanicus Cham. and Sch. becomes P. nuttallii Cham. and Sch., since
the latter was described on an earlier page of Linnea than the former
(II. 1827) ; P. hybridus Mx. (1803) being preoccupied (by Thuillier
in 1790), gives way to P. diversifolius Raf. (1808) ; P. rufescens Schrad.
(1815) is antedated by P. alpinus Balbis (1804) ; P. fluitans Roth.
(1788), gives way to P. lonchites Tuck. (1848), inasmuch as it is
highly improbable that the European and American species are
identical ; P. zizii Mert. and Koch. of the “ Manual” appears to have
included two species which are now to be known as P. thulaformis
(Robbins) Morong, (P. gramineus, var. (?) spathulaformis Robbins, P.
spatheformis Tuck., and P. varians Morong), and P. angustifolius
Berch. and Presl. (P. lucens, var. minor Nolte); the var. lanceolatus
Robbins (1867) of P. perfoliatus L. being preoccupied by Blytt (1861)
- Must give way to var. richardsonii Ar. Bennett (1889) ; P. pauciflorus
Pursh. (1814) must be replaced by P. foliosus Raf. (1808) ; P. mueron-
570 The American Naturalist. l [June,
atus Schrader, being uncertain, the species is to bear the name of P,
major (Fries) Morong; P. tuckermani Robbins (1856) gives way to
the earlier P. confervoides Reichb.; P. marinus L. turns out not to be =
that species, and must take the name P. filiformis Pers. (1805). ;
CHARLES E. BESSEY. ;
Pe ots) EE A eE
Hough’s American Woods.—The third part of R. B. Hough's
« American Woods” has recently been distributed. The twenty-five
species in this part are Magnolia glauca, Ilex opaca, Acer rubrum, A.
negundo, Prunus pennsylvanica, P. avium, Pyrus communis, Oratægus
punctata, Amelanchier canadensis, Liquidamber, styraciflua, Diospyros
virginiana, Fraxinus sambucifolia, Morus rubra, Hicoria suleata (Carya
sulcata), H, glabra (Carya porcina), Quercus bicolor, Q. prinus,
muhlenbergii, Q. coccinea, Betula populifolia, Salix amygdaloides, Pop-
ulus tremuloides, P. dilatata, Chamæcyparis thyoides, Pinus mitis.
Each species is represented by three sections of the wood, transverse,
radial and tangential, each 44 by 2inches. A good descriptive text
accompanies the set of specimens. The parts are sold by the author at
Lowville, N. Y., for the low price of five dollars each.
: CHARLES E. BESSEY.
Allen’s Characee of America.—Five years ago Dr. T.F.
Allen of New York City brought out Part I of a promising work on
the Characez of America, consisting of an introductory chapter on the
structure, followed by the keys to the species of all our genera. e
has now brought out the first fascicle of Part II in which he begins the
work of carefully describing and illustrating every species. The illus-
` trations are ample, there being no less than fourteen plates for the eight
species of Nitella included. The descriptions are full and apparently
well drawn up, measurements being fully given. The following are
the species described : oe
N. opaca Ag.—New England and Canada to California and Mexico.
N. obtusa Allen.—A new species from Lake Tamiscouata, Canada.
N. montana Allen.—A new species from Montana.
N. blankinshipii Allen—A new species from Missouri.
N. missouriensis Allen.—A new species from Missouri.
N. flexilis Ag.—Across the continent.
N. subglomerata A. Br.—N. Y. and N. J. to Oregon, Texas and :
Missouri and the var. brachyteles A. Br. of this species occurs ae
Mexico Alabama.
ni glomerulifera A. Br—Mass. to N. J., Ohio and Louisiana.
Every botanist will hope for the early a ance of the succeeds —
fascicles. A : T ARLES E. BESEY.
1893.] Zoology. 571
ZOOLOGY.
A Medusa from Lake Tanganyika.—In the “Annals of
Natural History,” for the present month, will be found an account of
a very interesting zoological novelty. Mr. R. T. Giinther describes
and figures a remarkable new form of Medusa, or jelly-fish, that occurs
in Lake Tanganyika. Until recent years, when the little Limnoco-
dium was found living in the Victoria lily-tank of the Botanic Gar-
dens, Regents Park, it was believed that the Medusæ were nearly
exclusively oceanic. It is now shown that the freshwater lake Tanga-
nyika is the home of a peculiar member of this group. The existence
of such an organism in Tanganyika was asserted some years ago by the
German naturalist, Dr. Boehm, and Professor v. Martens, of Berlin,
even went so far as to name it Tanganjice, although he had never seen
a specimen. Mr. Giinther now supplies us with a full description of
this singular Hydrozoon, which he refers to a new genus, Limnocnida,
adopting the suggestion of v. Martens as to its specific name. Lim-
nocnida tanganjice is, as might have been anticipated, perfectly differ-
ent from all the members of the group hitherto known, and probably
represents a distinct family, but its exact position cannot be settled
positively until the mode of its development has been ascertained.
(Nature, April 13, 1893.)
The Air-Bladder and Weberian Ossicles in the Siluroid
Fishes.—A study of the physiology of the Weberian ossicles and of
the air-bladder in general has been made by Professors T. W. Bridge
and A. C. Haddon, for the purpose of discovering the physiological
relation of the Weberian mechanism to one of the several functions
that have been ascribed to the auditory organ or to the air-bladder.
With this object in view they discuss (I) how far the function of the
Weberian mechanism is conditioned by the anatomical structure of
_ the air-bladder and auditory organs as well as by the character of the
mechanism itself; (II) to which of the known functions of the air-
bladder and auditory organ the Weberian ossicles are to be regarded.
as accessory structures; and (IIT) the utility of the mechanism to the
fish possessing it.
The alee find (I) that from the anatomical structure of the parts
the Weberian apparatus is better adapted to register the more forcible
distentions or contractions of the anterior chamber of the air-bladder
So pee
572 The American Naturalist. [June,
ee
rather than the slight or rapidly recurring vibrations of its lateral
walls. (II) The ossicles under consideration are accessory to the
hydrostatic function of the air-bladder. (IIIT) The Weberian mechan-
ism is of great functional importance to the fish possessing it, since it
confers on them an exceptional capacity for freedom of locomotion in
a vertical direction. The possession of this mechanism permits all
movements to be made with the maximum economy of muscular effort
and tissue metabolism.
In regard to the evolution of the Weberian mechanism the authors
reach the following tentative conclusions: i
1. The special feature of a fresh-water habitat that has conditioned
the development of the Weberian mechanism in the Ostariophysex is
the occurrence of seasonal or periodic quantitative variations in the
food supply, variations to which the Ostariophysex, from their herbiv-
orous or omnivorous habits are specially liable. ;
2. In view of such unfavorable nutritive conditions, the special
advantage which is conferred upon the Ostariophysex by the posses-
sion of the Weberian mechanism is a capacity for executing locomotor
movements in any plane, with an almost irreducible minimum 0
muscular effort and tissue metabolism.
3. If a variable and inconstant food supply is to be regarded as one
of the inevitable conditions of a fresh-water existence, and necessitates
strict economy in the expenditure of muscular energy, any mechanism
which secures this result must be of unquestionable importance to the
species, and hence it may be that the Ostariophysee owe their domi-
nant position among fresh-water fishes to the possession of the Weber-
ian mechanism.
4, The evolution of the Weberian mechanism has not only condi-
tioned the predominancy of the Ostariophyses, but, indirectly, has
favored the existence in fresh water of a large number of purely car
nivorous fishes, which depend on the former for their food, and there-
fore may also be regarded as one of the primary causes of the anomal-
ous abundance and diversity of fresh-water piscine life, as compared
with the remarkable poverty of all other groups of fresh-water organ-
isms. (Proceeds. Roy. Soc. Vol. LII, 1892).
Age Modifications of the Mucous Lining of the Stomach
of Ruminants.—In a study of the Comparative Anatomy of the
Stomachs of Ruminants, Mr. J. A. Cordier has discovered the follow-
ing interesting facts. es
The interior of the stomach of a young adult is covered with papille
closely packed, which are larger in the region which Wilkins calls the
SEN Zoology. 573
“col” of the paunch, than on the rest of the surface. In the stomach
of a very old animal a different condition exists. The large papillze
of the “col” are few and far between—two or three times as far apart
as in the younger animal, much smaller, twisted once on the base, evi-
dently becoming atrophied. Between these papille can be seen traces
of many others which have disappeared, leaving their bases only as
vestiges, and these are entirely covered by the ordinary epithelium of
the stomach.
That these papills do not renew themselves is shown by histological
sections of the papille. In a bison, thirty years of age, the papille
were almost entirely wanting. An antelope and a ram showed the
process of degeneration very clearly.
The second age modification is the appearance of a black coloration
more or less intense over the entire surface of the stomach. Observa-
tions so far, however, show that this modification takes place in domes-
tic animals only. [Bull. Soc. Zool., T. XVII, 1892.]
. Zoological News.—According to C. H. Eigenmann, the develop-
ment of the Point Loma blind fish (Typhlogobius californiensis Stein-
dachner), is a striking example of the degeneration of the eyes. The
embryo, before it is hatched, has eyes developed as well as the embryo
of any other fish. When the individuals have reached the length of
an inch they can still see a short distance, but it is evident that the eye
has stopped growing long before this age is reached. In the adult con-
dition the eye has become degenerated and covered with a thick skin,
and the fish is totally blind. (Proc. U. S. Natl. Mus., 1892.)
In his Comparative Notes on Swifts and Humming-Birds, Dr.
Shufeldt submits 61‘important structural differences existing between
the Cypseli and the Trochili, These differences, in the author's
opinion, establishes the fact that these two groups, morphologically
speaking, are not related. (The Ibis, Jan., 1893).
A new rat, Perognathus merriamii, is described by J. A. Allen, from
southeastern Texas. It is allied to P. flavus, but differs from it in col-
oration, in the general form of the skull, and in the relative size and
proportions of special parts of the skull. The species is based on 17
specimens from Brownsville, Texas. (Bull. Am. Mus. Nat. Hist., 1892.)
Mr. Amos Butler reports that the “ Least Shrew,” Blarina parva
(Say), the smallest mammal in the United States, is rather common in
the Whitewater Valley in Indiana. (Proc. Indiana Acad. Science,
1891, . : 63, 1893.) , ,
574 The American Naturalist. [June,
Mr. Witmer Stone notes the occurrence of the genus Neotoma in
Pennsylvania. The specimens, which were secured near the top of
South Mountain-in Cumberland Co., evidently belong to a new species,
and are described by Mr. Stone under the name Neotoma pennsyl-
vanica. This species is distinguished from N. floridana by its larger
size, its densely hairy and distinctly bicolored tail, and by certain well-
marked cranial characters. (Proceeds. Phila. Acad., 1893.)
ENTOMOLOGY.
North American Cosmetidæ—A recent study of a considerable
collection of Cosmetidæ from the Southern States sl that three well-
marked species occur in our fauna. This family belongs to the sub-
order Mecostethi of Simon or Laniatores of Thorell, of the order
Opileonea. Say described oné species, Wood another and Sorensen
the third. The three species are closely related and all belong to the
genus Cynorta. They may be separated by the following key:
Posterior pair of abdominal tubercles very prominent ; four or five
ai ‘
Posterior pair of abdominal tubercles little larger than anterior pair.
Dorsum with a distinct yellow Y connected posteriorly with a
transverse yellow line. C. albolineata. i
Dorsum without or with very little yellow marking. C. sayt.
orta ornata is abundant in Florida and probably occurs in the
South Atlantic States ; C. albolineata is found in Louisiana and Miss-
issippi; and C. sayiin Texas. An illustrated descriptive synopsis of
these species is now in the hands of the American Entomological
Society for publication in the Transactions CLARENCE M. WEED.
An American species of Sabacon.—In 1879, the French
Arachnologist, M. Eugene Simon established? the genus Sabacon for &
very peculiar species (S. paradoxus) of the family Ischryopsalide—
one of the smaller families of the Opileonea. The specimen descr!
was immature and the lateral pores were not distinct. No other
species of the genus appear to have since been described.
‘The genus Sabacon is especially characterized by the peculiar form
ofthe palpi, the joints of which are large and swollen, and the short
‘Edited by Prof. C. M. Weed, New Hampshire College, Hanover, N. H.
Arachnides de France, VII, 266.
1893.] Entomology. 575
tarsus is not provided with a claw, but is capable of being turned back
against a depression in the tibia. The mandibles are shorter than the
body.
During the autumn of 1892, I found under a piece of driftwood
along a small creek, a fully developed male belonging to this genus.
The species is evidently rare for I have never been able to find another
although I have searched persistently.
Sabacon spinosus, n. sp.
Male—Body 3 mm. long, 2 mm. wide ; palpi, 8mm. long. Legs:
first, 11.5 mm. ; second, 19 mm. ; third, 12 mm.; fourth, 16.5 mm.—
Body testaceous with dusky markings; the markings on dorsum
arranged transversely and following segmentation. Palpi and legs
light testaceous with almost continuous dusky blotches. Ocular
tubercle black, very near front margin of cephalothorax, much wider
than long, low, with a deep longitudinal sinus but no spines on carine.
A small round, not very distinct pore on each cephalo-lateral angle of
the dorsum, not isolated by distinct oblique sinuses. On the dorsum
of the cephalothorax and the ocular tubercle are many short, acute,
black spines arranged more or less irregularly ; back of ocular tubercle
on cephalothorax are two transverse rows of similar spines, and on
Fig. 1—Sabacon spinosus. Male: A, palpus; B, eye eminence; C, distal tarsi
of first leg. All magnified.
576 The American Naturalist. [June,
abdomen are many more similar rows. The ventrum of abdomen is
also provided with such rows, and the ventrum of the cephalothorax
including coxæ is covered with these spines. Palpi large and longer
than the body ; thickly covered with long black spines ; femur enlarging
a little from base to apex ; patella thick, subcylindrical, with a conical
tubercle on ventral surface near distal end; tibia slightly petiolated,
curved and attenuated distally, hollowed out on under side to receive
tarsus; tarsus petiolated, swollen, a little more than half as long as
tibia, rounded at end and having no claw; capable of being turned
_back upon the tibia like a thumb. Mandibles short, much shorter
than body ; first joint having a large, truncate, wart-like tubercle on
dorsal surface near distal end; top of tubercle and dorsal surface of
distal portion of the joint thickly furnished with stiff spinose hairs;
second joint short and thick, provided dorsally with similar hairs;
claws curved, unequal. Legs rather slender, with rows of spinose
hairs on proximal joints, Shaft of genital organ long, flattened;
toward tip enlarging into a spoon-shaped portion, from which there
projects forward a long slender piece gradually coming to very acute
point.
Described from one specimen taken at Hanover, New Hampshire.
CLARENCE M. WEED.
The Puparium of Jurinia.—In a paper recently sent to the
American NATURALIST, I described the puparium of Blepharipeza.
The present paper describes the puparium of Jurinia, which genus,
while it belongs to the same group as Blepharipeza (Hystriciine),
shows considerable difference in the puparium. The description is
drawn from a puparium of Jurinia algens Wd., from which issued a 2
specimen of the fly, bred by Professor C. P. Gillette from Hadena
lignicolor, in Colorado.
Puparium of Jurinia algens Wd.—Length, 12 mm.; greatest width
(8th segment), 54 mm. Color reddish brown, capital tubercles and
anal stigmata blackish. Puparium consisting of 12 segments, including
capital and anal plates, more or less cylindrical, bulging a little pos-
teriorly, the anterior end being less in diameter than the posterior en@,
while the eighth segment is the widest portion. The rugose belts
described in Blepharipeza are absent, the whole surface being more OF —
less fluted, the flutings showing most plainly on the three anterior 568-
ments next the capital plate, becoming less distinct in the middle oF —
giving way to an almost smooth surface, and reappearing in irregular e
flutings, or minute furrows and ridges, on the last three segments and a
1893.] Entomology. 577
anal plate. The larval mouth parts are represented on a portion of
the capital plate forming part of the single anterior flap which is pres-
ent, by two small erect tubercles projecting straight out from the sur-
face of the integument, situated at edge of capital plate, and on
opposing sides. The remaining portion of the capital plate, which is
absent, the other flap being detached and missing, doubtless bears a
third similar erect tubercle, the three being so arranged that they
represent the corners of a nearly equilateral triangle. These tuber-
cles are erect, about as high as their basal diameter, ending in a blunt
but laterally compressed apex, the apical diameter one way equalling
the basal while the other way it is much less than the basal, the apical
surface with about 4 faint transverse notches leaving 5 faint transverse
ridges. The longitudinal axis of the compressed apex of each tubercle
is at an angle of about 45 degrees with the margin of the capital plate.
The surface of the capital plate is more or less irregularly transversely,
not circularly, fluted. Anal stigmata situated in center of anal plate,
consisting of two erect raised organs, not as far apart as the diameter of
either, each consisting of 3 ridge-like sections separated by deep
notches, the flap-like ridges of each stigma quite closely approximated
at their inner ends and widely divergent outwardly, the superior and
inferior ones diverging at nearly a right angle. Each of these keel-
like ridges bears a longitudinal median suture or fissure its whole
length. These stigmatic organs, unlike those of Blepharipeza and
many other tachinids, project straight out from the surface of the
integument, being about as high as their diameter. A little distance
ventrally (? dorsally) of the pair of anal stigmata, being situated on
anterior border of 10th segment, is a small slit-like opening in the
integument resembling a spiracle, its longer diameter being longitudi-
nal tothe puparium. The vent-like anal tubercle described in Blepha-
ripeza is not present. : :
Supplementary note to description of puparium of Blepharipeza.—
In the description above referred to, of the puparium of Blepharipeza
adusta Lw., I mentioned only one tubercle on the capital plate. This
was all that was present on the nearly detached anterior flap of the
puparium, the correspondihg flap on the opposite side being A
It should have been mentioned that the absent portion of the capital
plate doubtless bore two more tubercles similar to the one described,
situated near the edges of the plate as was also the latter, the three
being arranged in a triangle, but not so far apart asin Jurinia, being
of larger size.—c. H. TYLER TOWNSEND.
578 The American Naturalist. [June,
Notes.—In Bulletin No. 19 of the Iowa Experiment Station, Prof.
Herbert Osborn reports further experiments in destroying leaf-hoppers ;
Mr. H. A. Gossard discusses the Clover-seed Caterpillar ( Grapholitha
interstinctana) ; and Mr. F. A.Sirrine treats of the Potato-stalk-weevil
(Trichobaris trinotata). The latter species has been found breeding in
“ground cherries” (Physalis).
Prof. S. W. Williston contributes to the third number of the Kansas
University Quarterly the third part of his Diptera Brasiliana, and an
illustrated paper on the Apioceride and their allies.
Mr. F. M. Webster publishes’ extended accounts of the insects
affecting the blackberry and raspberry, and the underground insect
destroyers of wheat. The former article enumerates 88 species.
Mr. M. V. Slingerland’ calls attention to the fact that the black
peach aphis (Aphis persice-niger) is being introduced into New York
through peach trees purchased in Delaware. The author describes the
indications of the presence of the pest; its past history ; its classifica-
tion, appearance, and life history and remedial measures.
Professor J. B. Smith presents’ an interesting illustrated discussion
of the grasshoppers, locusts and crickets affecting cranberries. He
shows that contrary to the usual belief it is the katydids and not the
common grasshoppers that attack this fruit.
A catalogue of the South American species of Calyptrate Muscide
by Prof. C. H. Tyler Townsend has lately appeared in the Annals of
the New York Academy of Sciences. (Vol. VII, Dec., 1892).
5Ohio Agr. Expt. Station, Bulletins 45 and 46.
‘Cornell Univers. Agr. Exp. Station, Bull. 49, p. 325.
5New Jersey Agr. Exp. Station, Bull. 98.
1893.] Archeoloyg and Ethnology. 579
ARCHEOLOGY AND ETHNOLOGY.
The International Congress of Americanists.—(Continued
from page 305)—Firra CONGRESS AT COPENHAGEN, 1883.—The fol-
papers were read and addresses made :
Observations upon some of the skulls and human bones of the Min-
asgeraes in Brazil, by M. Lutken; Note on the Quaternary Fossil
Animals, Fauna of the Plateaux of the Andes; An Examination of
the Discovery of the American Continent by Christopher Columbus,
and the assistance given to him by Martin Alonzo Pinzon, by M. Her-
rera; Aboriginal American Literature, by Dr. Brinton; The Vineland
Excursions of the Ancient Scandinavians, by M. Lofer ; The Pre-
Columbian Relations of the Gaels with Mexico, by M. Beauvois;
Old Scandinavian Ruins in the District of Julianehaab, South Green-
land, by M. K. Steenstrup, with two plates; Three Ancient Charts of
the North, by Baron Nordenskiöld, explained by M. Christian Bahnson ;
M. Lucien Adam, upon the theory announced by M. Hale, relative to
the Origin of America; M. Bamps, On the Traditions Relative to the
White Man and the Sign of the Cross in America, prior to the Dis-
covery; and a Criticism upon the Theory of Abbe Schmidt, which had
been announced in the former Congress ; The Lost History of America,
by Mr. Steven Blackett; To what Point did the Ancient Scandinav-
ians Penetrate near the North Pole in their Expeditions over the
Glacial Sea? by M. Brynjulfson ; The Voyages of the Brothers Zeni in
the North, by M. J. Steenstrup ; Nautical Remarks about the Zeni
Voyages, by M. Irminger ; Cartography of the American Continent,
; The Voyages of the Danes in Greenland, by
Polynesian Antiquities, a Link Between
sia and America, by Mr. Francis A. Allen ;
Fabrication, by Anatole Bamps; On the
Paleolithic Implements of the Valley of the Delaware near Trenton, by
Dr. C.C. Abbott ; Ancient Ornaments of Pottery, by E. Barber ; Memoir
on the Shell Mounds of the Chesapeake Bay and Potomac and Wyco-
mico Rivers, by Elmer T. Reynolds; Prehistoric Archeology, by
Baron Joseph de Baye; The Art of Ornamentation Among the Ameri-
can People, by M. Stolpe; The Dialect of the Eskimo Language, with
a synoptical table of words arranged after the system of the Greenland ©
homas Wilson, of the U. S. National Museum.
Madrid, by M. de la Rada;
the Ancient Civilizations of A
American Pottery in Pate and
1This department is edited by Dr. T
is ER
tae
580 The American Naturalist. (June,
|
:
Dictionary by M. Rink.. The author divides the Eskimos into six
groups: the Greenlander, the Labradorian, the Eskimos of Makenzie,
the Eskimos of the West, and the Aleutes—Wherein does the Eskimo ?
Language Differ Grammatically from the other Languages of North
America, by M. Lucien Adam; M. Leon de Rosny had made a Vol-
ume on the Deciphering of the Maya Inscriptions, the which M. Rada
Translated and Presented to the Congress with an Analysis and
_ Criticism; The Language Timucua, by M. Vinson; M. Vahl pre- i
sented an ethnographic chart of North America for the Danish Mis- |
sionary Society, which he explained; The Coloring Matter Employed he
by the American Indian, by M. Vera; The Variations in the Physical a
Geography of the American Continent, from the discovery to the
present—M. Vera; and the Formation of the Words of the Maya
Language, by the Count de Charencey.
The Fourth and Sixth Sessions were held respectively at Madrid,
and Turin. I have no report of these Congresses.
The Seventh Session was held at Berlin from the 2d to the 5th of
October, 1888. The papers read and addresses delivered were substan-
tially as follows: A 3
The Basques, Britons and Normans on the Coast of North America me
during the early part of the XVI Century, by M. Gaffarel, 9 pages; a
Publication of the Writings and Documents Relative to Christopher a
Columbus and his Time on the Occasion of the Celebration of the
Fourth Centenary of the Discovery of America, by M. Cora; An His-
torical Essay on the Primitive Legislation of the Spanish State of
America, by M. Fabie; On the Nahuatl Version of Sahagun’s Historia
de la Nueva Espana, by Dr. Brinton, with its discussion, 6 pages; On
Certain Archzologic Objects of Mexico and South America, by M.
Heger, 5 pages; The Stone Colliers of Porto Rico, by Jimenez de la
Espada; Antiquities of Vera Cruz; Archzologic Results of the Later
Voyages in Mexico, by M. Seler; An Ancient Mexican Mosaic, by M.
Andree; Notes on the Origin, Working Hypothesis and Primary
Researches of the Hemenway Southwestern Archzlogical Exposition,
_ by Mr. F. H. Cushing ; The Antiquities of Nicaragua, by M. Bovallius; —
The Ceramic Antiquities of the Isle of Marajo, and also on Nephrite and
Jadeite, by M. Netto; Place of Origin of Nephrite and Jadeite, by M. ‘
Virchow; The Aztecs and their Probable Relation to the Pueblo :
Indians of New Mexico, by Mr. S. B. Evans; The Employ of Cocoa —
in the Northern Part of South America; American Craniology, by —
Virchow; An Anatomical Characteristic of the Hyoid Bone of Pre-
Columbian Pueblo Indians of Arizona, by Drs. Wortmann and Ten
1893.] Archeology and Ethnology. 581
Kate; The Chronology of Dilluvial Man in North America, Emile
Schmidt; Indication of the Vestiges of the Pre-Columbian Population
of Nicaragua, M. Desiré Pector; Human Sacrifice in America During
Pre-Columbian Times, by M. Grossi; Cremation in America Before
and After Christopher Columbus, by M. Grossi; Anthropology of the
People d’Anahuac in the Times of Cortez, by M. Hartmann; Was
America Peopled from Polynesia? by Horatio Hale; Study of the
Mam Language, by the Comte de Charency ; Vocabulary of the Lang-
uage Timucua, by M. Raoule de la Grasserie; The Linguistic Family
of Pano, by the same; The Historic Archives of the Hemenway South-
western Archeological Expedition, by M. Bandelier; On the Sam-
baquis of Brazil, by M. H. Muller; Ancient Map of America,
by Gaffarel ; Three Linguistic Families in the Amazon and Orinoco
Rivers, by M. Adam; Bibliography of Recent Linguistic Investiga-
tions in South America, by the same; Maya Handwriting by M.
-Férstemann ; A Chronologic Classification of the Architectural Monu-
ments of Ancient Peru, by M. Borsari ; Contribution to Americanism
of Cauca (United States of Columbia) by M. Douay ; The Language
of the People of the Center of South America by M. Von Den Steinen ;
Peruvian Figures in Silver, by M. Luders.
Language vs. Anatomy in Determining Human Races.—
“Anthropologie,” replying to criticisms of Dr. Sergi’s work, thus states
the position of French Anthropologists on this controverted subject :
The Anthropologists of France are unanimous that some or a few
(anatomic) characters are not sufficient to determine a type of race ;
but there should be an investigation of all or as many as possible, and
Anthropology does not interfere with Ethnography. They are occu-
pied with different things. Anthropology does not say that physical
characters are superior or inferior to linguistic characters ; it says the
two sciences are of a different order and for a different purpose. The
first relates to the physical element constituting peoples; the second to
the classification of these peoples. Language grows, loses, borrows,
changes, transforms, and all this independent of the Anthropological
characters, such as beliefs, customs, industries. Physical characters are
hereditary and inherent in the blood, while linguistie characters are
not. A red Indian, born among strangers and without the society of
his parents or race, will speak, not his own language, but the language
of those who rear him, and, nevertheless, retain all the physical char-
acters of his race. Different and opposing races may speak the same
language, and per contra, the same race may speak different languages.
582 The American Naturalist. [June,
Much of the contest which has developed in the United States as to
the relative value of language arises from the confusion of the terms
Anthropology, Ethnology, Ethnography. The French writer makes a
distinction easily understood and maintained, and quite harmonious
with the “ Nomenclature of Anthropology ” as presented by Dr. Brin-
ton and commented on by Major Powell before the Anthropological
Society of Washington (Amer. Anthrop., July, 1892, Vol. V, No. 3, p.
265, et seq).
Language may be of great value, and should never be neglected in
determining living or historic races. In determining prehistoric races,
it cannot be studied because in most cases it is unknown, and so we are
driven to consider the physical characters. Because language assists
in determining historic races, it does not follow that in the prehistoric
races no other means can be used.
The true rule seems to require the employment of all possible means,
and even then the decisién may not be either harmonious or correct.
The Nephrite of New Zealand.—MM. Duparc and Morazee
have published in the Archives of Natural and Physical Science,
Geneva, a paper on Nephrite of New Zealand. Speaking of the num-
ber of hatchets and other objects from the Swiss Lakes in the museums
which have passed as Nephrite, they express doubt and counsel circum-
spection. They do right and are to be approved. But when they say
that they have found but one piece which from its appearance could be
identified as nephrite, their proposition is doubtful. There are many
polished specimens of the hard, greenish stone in the Swiss Museums,
which stand tests of hardness and density required for Nephrite, and
which have been called so. If not Nephrite, they are still some varie-
ties of Jade. One or more of their components may be soda or alumi-
num instead of lime or magnesia ; and they may be Jadeite, fibrolite,.
saussaurite or another variety of Jade.
While many of these polished objects can be determined with rea-
sonable certainty on inspection and from appearance, yet there are
many which possibly cannot be. That is to say, they correspond in
appearance, and they stand the tests of hardness and density, and may:
be either the one or the other, depending upon their chemical combi-
nation, and this similarity may be so great that it is doubtful if, e
disputed cases, anyone can determine without analysis or microscopie
examination of thin sections. If MM. Dupare and Morazec examined —
the doubtful or contested specimens of the Swiss Museums in this w8Y>
their determination would be accepted, but if from mere externat-
oe
1893.] | Microscopy. 583
appearance, the subject is still open. Their conclusions may be quite
correct, but examination from external appearance does not prove it.
One great trouble arising in the discussion of this subject is that the
destruction of the object consequent upon analysis and thin section
prevents a satisfactory and competent determination.
MICROSCOPY.
A Method for Injecting the Blood-Vessels in Birds.—
I presume all who give a laboratory course in Comparative Anatomy
have, in common with myself, experienced difficulty in injecting the
arterial and venous systems of birds. The usual directions are to
inject the arterial system through the pectoral artery, and the venous
system either through the pectoral vein or through the coecygeo-mesen-
teric vein. The blood of birds coagulates very rapidly, and even when
the procedure is carried out as expeditiously as possible, failure fre-
quently occurs from the inability of the operator to remove the greater
part of the blood from the animal, or from vexatious delays in tying
a canula in the divided vessel. For some time I have been using the
following method, and have not failed in securing the most satisfactory
results. It possesses the additional advantage of being quickly per-
formed.
The bird, a pigeon for example, is placed under a bell-jar and chloro-
formed in the usual way. While the bird is getting under the influence
of the anesthetic, the operator should see that there is, within conven-
ient reach, a scalpel, a pair of stout scissors with both points rounded
off, two canulae and the necessary ligatures for tying them in place.
As soon as the pigeon is thoroughly under the anesthetic, the bell-
jar is removed, a towel is wrapped about the head and a few drops of
chloroform poured over it; the feathers are rapidly plucked from the
breast and abdomen to a point just anterior to the cloaca; the skin is
divided the entire distance over the keel, and the pectoral muscles dis-
sected off from their attachment to the keel and body of the sternum
for a distance of half an inch on either side of the keel.
The muscles attached to the posterior margin of the sternum are
next divided elose to the sternum for the distance of half an inch on
either side of the median line; now raise the sternum carefully and
divide the attachment of the heart. Beginning at the outer angle of
the divided abdominal muscles, the body of the sternum is cut through
ìThis department is edited by C. O. Whitman, University of Chicago.
584 The American Naturalist. [June,
with the stout, blunt-pointed scissors, in a direction forward and
inward, until a point is reached directly over the origin of the vascular
trunks; repeat on the opposite side. Now grasping the keel firmly
with the left hand, make traction directly upward, at the same time
steadying the bird with the right hand; by this procedure the sternum
is broken equally across, and the heart exposed without causing any
damage to it or to the vascular trunks. Next, tear the pericardium |
open as quickly as possible, seize the heart, still strongly beating,
between the thumb and index finger of the left hand, invert the bird
over a sink and cut off with the blunt-pointed scissors the apex of the
heart. In this way the blood-vessels are thoroughly emptied and the
bird killed without any unnecessary suffering.
Although the description of the procedure may make it seem long,
I have repeatedly done it in less than three minutes. i
canula is now inserted through the left ventricle into the aorta,
and a ligature placed around it close to the point where the aorta
leaves the heart. In passing the ligature around the canula, care
should be exercised not to injure the vena cava or the right auricle. A
second canula is passed through the right ventricle into the right
auricle and secured by tying a stout ligature about the whole heart.
Ihave found that the best injecting mass is Gage’s modification of
Pausch’s, colored with vermilion or ultramarine blue ; this does not
pass through the capillaries, sets quickly, leaving the vessels well-dis-
tended with a firm mass.
I employ a constant pressure apparatus in injecting, using a pressure
of 100 mm. for the artery, but only 60 mm. for the vein; more than
this will usually cause a rupture.
W. S. MILLER.
Univ. of Wisconsin.
1893.] Proceedings of Scientific Societies. 585
PROCEEDINGS OF SCIENTIFIC SOCIETIES,
National Academy of Sciences.—The following papers were
read at the meeting commencing April 18, 1893: On the Systematic
Relations of the Ophidia, E. D. Cope; Biographical Memoir of Gen-
eral Montgomery C. Meigs, H. L. Abbott; On the Nature of Certain
Solutions, and on a New Means of Investigating Them, M. C. Lea;
The Relations of Allied Branches of Biological Research to the Study
of the Development of the Individual, and the Evolution of Groups, A.
Hyatt; The Endosiphonoidea (Endoceras, etc.), considered as a New
Order of Cephalopods, A. Hyatt; A New Type of Fossil Cephalopods,
A. Hyatt; Results of Recent Researches upon Fossil Cephalopods of the
Carboniferous, A. Hyatt; Biographical Memoir of Julius Erasmus
Hilgard, E. W. Hilgard; Monograph of the Bombycine Moths of
America, North of Mexico: Part I.—Notodontide, A. S. Packard;
Intermediary Orbits, G. W. Hill; The Relations Between the Statis-
tics of Immigration and the Census Returns of the Foreign-born Pop-
ulation of the United States, Richmond Mayo-Smith ; Statistical Data
for the Study of the Assimilation of Races and Nationalities of the
United States, Richmond Mayo-Smith ; Telegraphic Gravity Deter-
minations, T. C. Mendenhall ; Comparison of Latitude Determinations
at Waikiki, T. C. Mendenhall; A One-volt Standard Cell; H. S. Car-
hart (introduced by T. C. Mendenhall); Fundamental Standards of
Length and Mass, T. C. Mendenhall; Peptonization in Gastric Diges-
tion, R. H. Chittenden; Helen Kellar, Alexander Graham Bell; On
a Potentiality of Internal Work in the Wind, S. P. Langley; On a
Bolograph of the Infra-red Solar Spectrum, S. P. Langley .
No election of members took place. The following were elected
foreign correspondents: Tissandier, ere Paris; Rammelsberg,
chemist, Berlin ; Ludwig, physiologist, Leipzig.
Boston Society of Natural sie G aie follow-
in rs were read: Mr. J. B. Woodworth : Traces of a Fauna in
the Cambridge Slates; Mr. Charles P. Bowditch: Ruins of Central
America. The annual meeting was held May 3d. The following
. paper was read: Dr. R.T. Jackson—Notes on the Development of
Palms. SAMUEL HENSHAW, Secretary.
586 The American Naturalist. (June,
New York Academy of Sciences.—April 10, 1893.—Biologi-
cal Section—H. F. Osborn, on “ The Evolution of Teeth in Mammalia
in its Bearing Upon the Problem of Phylogeny,” reviewed the recent
researches and theories of Kükenthal, Röse and Taeker upon the forma-
tion and succession of the dental series in mammalia, and pointed out
that, especially in marsupials, cetaceans and edentates (with other
placentalia) the existence of two series of teeth was now abundantly
proven, as well as the fact that Homodynamous forms were derived from
early Heterodont. He then showed that recent discoveries demonstra-
ted that in marsupials, teeth of the second series might be interposed in
the first series—to explain the typical dentition of such forms as Didel-
phys. This transposition enables a comparison of dentition of mar-
supials with that of Jurassic mammalia (=i, $, c, t, p, 4, m, $). It
was further noted that the triconodont type (as Amphilestes) was
probably the hypothetical point of divergence of placental mammalia.
As to the form of crowns, the theory (Kükenthal-Röse) that complex
mammalian types were made by concrescence of simple reptilian
cusps was upon the evidence of the Jurassic mammalia shown unten-
able, as well as the converse theory that cetaceans have derived their
homodynamous form by the splitting of the cusps of triconodont.
Bashford Dean, in “ Contributions to the Anatomy of Diniththys,”
correlated the parts of this Devon-Lower Carboniferous Arthrodiran to
those of Coccosteus. Notes were made upon the (1) disposition and
character of the lateral line organs, (2) pineal foramen, (3) nasal cap-
sules, (4) dentary plates (homologies), (5) ginglymoid articulation of
lateral shoulder plates, (6) character of shagreen, (7) probable disposi-
tion of paired and unpaired fins.
N. L. Britton presented a “Note on the Genus Lechea.” This
genus of Cistinee is entirely American, and from the investigations of
Mr. Wm. H. Leggett and Dr. Britton appears to consist of about four-
teen species.
Natural Science Association of Staten Island.—March 18. | o
—Mr. Wm. T. Davis exhibited specimens of the Leopard Moth and
read the following communication : ,
On the 23d of June, 1888, I found on the sidewalk, under & partly
decayed white maple on Fort Hill, a large white and black moth that
was rendered helpless by having one of its fore wings broken in two
_ It was not until last year, when Col. Nicholas Pike’s article and —
accompanying figures on the ravages of the Leopard Moth—Zeuzer@
esculi (Linn.)—in Brooklyn, appeared in “ Insect Life,” that the moth
1893.] Proceedings of Scientific Societies. 587
found by me was proved to be the same species from Staten Island.
Col. Pike found in 1889 that it had bored nearly all of the trees,
mostly maples, from Carlton Avenue to the entrance of the park, and
that in 1891 it had extended its ravages throughout the city. He also
mentioned it as having appeared in Astoria, New Rochelle, Jamaica,
New Lots and Flatbush.
In August, 1887, Mr. J. B. Engelman took three specimens of the
Leopard Moth in Newark, N. J., and in 1888 many more were found.
Previous to this the very rare occurrence of the moths in this country
had been credited to the importation of wood containing the larve.
In 1889, Mr. Beutenmuller mentions it as having been found not
uncommonly in Central Park. From these facts it will be seen that
this destructive insect from the Old World has become quite generally
distributed in this vicinity.
Mr. Davis also read the following note :
With Messrs Kerr and Leng, on the 26th of last February, I observed
about twenty-five Snow Buntings—Plectrophenax nivalis (Linn.)—in a
field on Todt Hill, near the highest point of the island. They were
feeding in the few places where there was no snow, and flew occasion-
ally in a remarkably compact flock, from one to another of these
isolated spots. This bird has not been reported on the island for a
number of years, and its present occurrence is undoubtedly due to the
severity of the winter.
Mr. J. H. Bowles, in “Science,” for January 13th, 1893, comments
upon the abundance of the Pine Grosbeak—Pinicola enucleator (Linn.)
—and the scarcity of the Snow Bunting about Ponkapoag, in eastern
Massachusetts, and gives as a cause the unusually cold weather. The
Grosbeaks and the Buntings have each ranged further south than for
several years past. Though the weather has been severe, robins have
been seen sparingly on the island during every month of the winter.
Mr. Arthur Hollick remarked that the last time he remembered see-
ing the Snow Buntings in abundance in this vicinity was during the
winter of 1872-73, when they appeared in great flocks, accompanied
by the Horned Lark—Otocoris alpestris (Linn.). That — ay
were particularly numerous on Fort Hill, New oe many
days in December and January, and large num killed by
gunners. .
Mr. Walter C. Kerr exhibited, under the ‘microscope, sity fon
Quercus nigra L., Q. ilicifolia Wang., and Q. brittonii Davis, an
the following paper in connection with them :
588 The American Naturalist. [June,
In describing the new hybrid, Quercus brittonii, in our Proceedings
for September 10th, 1892, Mr. Davis refers to the pubescence of the
under surface of its leaves being intermediate between that of Q. ilici- _
folia-and Q. nigra. This observation was made from casual inspection,
without reference to the exact nature of the pubescence. Under the
microscope it will be seen, as in specimens submitted, that the Q. ilici-
folia has so dense a pubescence that the epidermis. is completely
hidden, while in Q. nigra the tufts are quite separated, even isolated.
In Q. brittoni, the hybrid between these species, the intermediate char-
acter of the pubescence is most striking, the under surface of the leaves
being starred at regular intervals by the tufts, which are usually suffi-
ciently close to allow their spreading hairs to touch one another.
These tufts seem to be composed of six to twelve hairs spreading from
a common base and occupying a space about .15 to .2 mm. in diameter.
An average specimen contains about sixteen tufts per square mm.,
while in any other, representing the strongest pubescence which the
hybrid seems to attain, about twice this number were present and
somewhat smaller in size, indicating considerable variation in these
appendages.
The wide difference in pubescence of ilicifolia and nigra, however, is
such that a variation of even one hundred per cent. between different
specimens of Q. britonii is not sufficient to materially affect the dis-
tincti of this characteristic. -
Mr. Arthur Hollick called attention to the fact that an earthquake
shock had been experienced on the island shortly after midnight on
March 8th, which had been sufficiently severe to awaken many people,
especially those who resided on the hills in New Brighton, but that it
did not seem to have been felt generally throughout the rest of the
island.
Mr. Jos. G. Thompson stated than he found, on South Beach, a dead
specimen of the Tom-Cod (Gadus tomcodus), about eight inches long,
which had made a meal of eight mud-killies ( Umbri limi). Those that
were in the foremost part of the stomach were quite perfect, while the
others that were further down had begun to be dissolved by the digest-
ive fluids. t.
_ April 8—Mr. Arthur Hollick exhibited specimens of Indian rub- :
bing stones and read the following paper concerning them :. ee
Since our last meeting, while on a tramp across the island from Please
ant Plains to Rossville, I was struck by the fact that the shell heaps,
which have been noticed in that region at different times, form pracy
tically a continuous chain or trail from shore to shore. They follow
1893.] Proceedings of Scientific Societies. 589
the course of Sandy Brook up to Woodrow road, and from thence may
be found in nearly every field or piece of cleared ground until we reach
the village of Rossville. Indian implements of various kinds, with
fragments of pottery have been picked up at many points and are
likely to be met with in any part of the region. The distance of some
of these accumulations from the shore, their elevation above the water,
and the fact that in many places they are spread thinly and evenly on
the surface has led me to think that the more distant ones from the
salt water may have been carried there in recent times by the farmers
for purposes of fertilization, as is frequently done. Whether this be
so or not, implements may be found wherever the shells occur, prov-
ing their origin indisputably.
Amongst a number of relics found at the time mentioned was an
unusually fine specimen of a rubbing or polishing stone—an imple-
ment of which few have turned up on Staten Island, and in regard to
which I believe, the attention of the Association has never been called.
Amongst the hundreds of implements which our members have col-
lected during the past ten or twelve years from all parts of the island,
I find but seven which can be classed in this category—all, with the
exception of this one, from Tottenville.
Mr. William T. Davis presented nine plants, new or rare to the flora
of the Island, with the following memoranda :
Rubus odoratus L. Side of Todt Hill road near the highest point.
Spreading from introduced plants. Aethusa cynapium L. Abundant
in a field corner Crescent and Jersey streets, New Brighton. Pre-
viously reported only from Clove Lake swamp. Oxycoccus macrocarpus
Pers. Near Sprague avenue, Tottenville, and accompanied by the
“Cotton grass” (Eriophorum) as in the swamp near Richmond village.
Vaccinium pennsylvanicum Lam. Watchogue and Arlington. Stachys
palustris L. var. cordata. Abundant in field near Eltingville station.
Pinus mitis Michx. Abundant at Linoleumville. Habenaria bleph-
ariglottis Hook. Sparingly at Watchogue along Merrill's road. Calo-
pogon pulchellus R. Br. Merrill’s road near trap ridge. Collected by
C. Leng. Smilacina stellata Desf. Borders of salt meadow at
‘Great Kills.
Mr. Walter C. Kerr exhibited a large fragment from the broken
trunk of a horse chestnut tree, showing profuse budding, and read the
followi :
dees ena sur i:no the adventitious budding of a horse chest-
nut, Æsculus hippocastanum L., on Tompkinsville Hill may be worthy
5 eastern brow of the hill there stands,
of passing notice. On the bleak
39
590 The American Naturalist. [June,
among the few scattered trees remaining, a horse chestnut about 18 ae
inches in diameter, branching about three feet from the ground into
two trunks, each about ten inches in diameter. ‘The trunk leaning to
the eastward is thriving and about the normal height for its diameter.
The one leading to the westward has broken off, with an irregular
splintery fracture, about five feet from the ground, perbaps on account
of its lesser resistance to our easterly gales, and the stump has rotted-
badly. The bark, however, retains its vigor, and from the cambium
layer, where exposed along the irregular edges of the ruptured section,
adventitious buds have sprouted profusely. They are also found
where the bark has split, and in the crotch where the tree has forked,
where the bark of the two trunks unite. Some of the thickest colonies.
of buds were at the apex of the splintered stump, and I therefore sawed
off about fifteen inches with its buds and three young shoots which have
seemed fortunate enough to grow to the length of some eight inches.
It will be noted that the bark is in a good state of preservation, the
inner layers alive, while the wood is not only dead, but far gone in
decay. The thickest cluster of buds is 8 inches long by 1} inches
wide, and within this space I have counted 200, about 30 of which are
alive, the others being mostly the remains of a previous crop. There
is no evidence of accidental destruction of the buds, though a few small
shoots may have been cut off. Cows would scarcely browse on them,
‘and one cluster occupied an inaccessible position in the fork.
One is not surprised at the dense growth of shoots which rise from
adventitious buds on a decapitated willow or the spraying branches of
elm which are of similar origin, but I have never before noticed so pro-
fuse a crop of buds whose mission seems to be entirely futile. They
seem to represent an especially vigorous effort of the broken organism
to survive, and under the circumstances this effort might be very per-
-sistent because of the opportunity afforded the unfortunate trunk,
deprived of means of assimilation, to draw on the sister trunk, fully
developed, for the requisite nourishment. It therefore might seem to
have more opportunity to thus maintain its life than had it been @
single trunk snapped off and dependent only on the residual nourish-
ment within its roots. This supposition is, in some degree, supported
by the fact that this particular tree stands on a hill some 300 feet high,
rooted in the thin, dry soil covering a barren serpentine ledge, and
hence would scarcely be expected to show a vigor which compares
favorably with the sprouting of the adventitious shoots of the brookside
willow. The specimen from the stump shows only three small surviv-
ing shoots, and may it not be fairly surmised that if the budding effort
1893.] Proceedings of Scientifie Societies. 591
of the cambium layer of this old stump could have been concentrated
into a few buds and their resultant shoots, the struggle for existence
would have been more successful. It would seem, even in the absence
of positive evidence, as though the prolificness with which the buds
were formed seriously retarded the survival of any, and thus the abil-
ity of the cambium layer of this stump to restore the interrupted
growth was handicapped by the opportunity afforded for abnormal
budding effort through the supply of nourishment obtainable from the
uninjured half.
Mr. George Dupuy exhibited a series of about 400 drawings repre-
senting Staten Island pond life—diatoms, desmids, algae and infusoria
—all drawn to scale on cards, handsomely colored and arranged.
The Biological Society of Washington.—April 8.—The fol-
lowing communications were made: Professor J. W. Chickering—The
Botanical Landscape; Frederick V. Coville—Characteristics and
Adaptations of a Desert Flora; Dr. C. W. Stiles—Notes on Parasites ;
The Cause of “ Measly Duck,” with|microscopic demonstration ; Dr.
R. R. Gurley—Natural Selection as Exemplified by the Cackling of
Hens.
April 22.—The following communications were made: Mr. O. F.
Cook—Notes on the Natural History of Liberia; Dr. J. N. Rose—
Two New Trees of Economic Importance from Mexico; Dr. V. A.
Moore—Observations on the Distribution and Specific Characters of
the Streptococci Group of Bacteria; Dr. Erwin F.Smith—Peach Yel-
lows and Plant Nutrition. FREDERICK V. COVILLE, Secretary.
Geological Society of Washington.—A Geological Society has
recently been organized in Washington, D. C., for the presentation and
discussion of topics of interest to geologists. The constitution and
standing rules were subscribed to by 109 founders at the first public
meeting, March 8th, 1893. Its members are of two classes, active and
corresponding. The annual dues of the first are $2, and of the second,
$1. Meetings will be held on the second and generally also on the
fourth Wednesday of each month from October to May, inclusive. —
The journals and bulletins of the various societies appear to furnish
sufficient opportunity for the publication of papers read before the
Society, so that for the present the Society will not undertake to pub-
lish the papers presented. It will probably issue one bulletin each
year containing the address of the retiring President, and such other
matter as the Council directs. J. S. DILLER, Secretary.
592 The American Naturalist. [June,
SCIENTIFIC NEWS.
Correction.—Dr. D. S. "Jordan informs me that the name albeolus
is not available for a new species of Notropis since it has been used in
that connection. The species called Notropis albeolus in the AMERI-
can NATURALIST, January, 1893, p. 152, may stand as Notropis jor-
danii. C. H. & R. S5. EIGENMANN.
Alphonse Louis Pierre Pyramus DeCandolle, born in Paris,
October 27, 1806, died in Geneva, April 4, 1893. The son of an emi-
nent botanist (Augustin Pyramus DeCandolle, 1778-1841), with whom
he labored many years, and himself the father of one who ranks high
among living botanists (Casimir Pyramus DeCandolle, 1836 ), the
subject of this sketch lived all his days a botanist. His first paper,
Monographie des Campanulées, appeared in 1830, and from that date
until within a short time, his brain and pen have been busy. In 1839
he began his contributions to the “ Prodromus ” which his father had
inaugurated fifteen years earlier, continuing until the final abandon-
ment of the work in 1873. A few years later (1878) he and his
son Casimir brought out the first volume of Monographiae Phanero-
gamarum, which was intended as in part, a continuation and revision
of the Prodromus. Of this great work, seven stately volumes have
appeared.
Aside from this systematic and descriptive work, DeCandolle will be
remembered for his treatise on geographical botany, Geographie
Botanique Raisonnée (2 vols., 1855), and his well known Lois de la
Nomenclature Botanique (1867), of which an English translation
appeared the year following. The latter, in these days of discussion,
is not likely to be forgotten or overlooked. La Phytographie, which
appeared from his pen in 1880, is not as well known as its merits
deserve. The much more popular Origin du Plants Cultivés (1885),
of which an English translation appeared in the International Scien-
tific Series is one of the most widely read of botanical works.
Lectures at the Paris Museum.—The following course of lec-
tures will be given at the Museum of Natural History during 1893:
April 25, Preliminary Lesson, M. Milne-Edwards; April 27,
Anthropology, M. Hamy; April 29, Ethnography, M. Verneau ;
May 2, Mammalia, M. Oustalet ; May 4, Birds, M. Oustalet ; May 6,
1893.] Scientific News. 593
Reptiles and Fishes, M. Vaillant; May 9, Molluscs, M. Perrier; May
13, Worms and Zoophytes, M. Bernard; May 16, Insects and Crus-
taceans, M. Ch. Brongniart; May 18, Comparative Anatomy, M. Pou-
chet; May 20, Botany (Phanerogams), M. E. Bureau; May 23, Bot-
any (Wood, Cryptogams), M. Van Tieghem ; May 25, Living Plants,
M. Cornu; May 27, Paleontology, M. Albert Gaudry; May 30, Geol-
ogy, M. Stanislas Meunier ; June 1, Meteorology, M. Daniel Berthelot ;
June 3, Mineralogy, M. Lacroix; June 6, Hygiene for Travelers, M.
Gréhaut. (Revue Scientifique, April, 1893.)
The Nottingham Meeting of the British Association for the Advance-
ment of Science will be held under the presidency of Professor Burdon-
Sanderson, the well-known physiologist. The presidents of the
Natural History Sections are to be: Geology, J. J. H. Teall; Biology
the Rev. Canon Tristram; Geography, Mr. Henry Seebohm; Anthro-
pology, Dr. Robert Monro.
Dr. Dall has done a good work in ascertaining the dates of the late
Timothy A. Conrad’s books upon the Tertiary fossils of the United
States. The quarrel which was their cause or their effect has fortun-
ately passed into ancient history, and these unfortunate volumes show
the ground of its permanent settlement.
The cousins Sarasin, so well known for their expedition to Ceylon
to study the development of the Cacilians, are about to start for a
several years’ stay in the Celebes.
The Rev. H. N. Hutchinson, 30 Vincent Square, Westminster, Lon-
don, S. W., England, has, for sale, plaster casts of a model of the horned
saurian Agathaumas, 12 inches long, for ten shillings.
Professor Karl August Lossen, Chief Geologist of the Prussian
Survey and Professor of Geology in the University of Berlin, died
February 24, 1893. Most of his geological work was done in the
Hartz Mountains. He was born January 5, 1841.
Professor Karl Prantl, of the University of Breslau, died February
24, 1898, in his forty-fourth year. He is best known for his text-book
translated by Professor Vines, and for his share in the “ Pflanzenfamil-
ien” of which he and Professor Engler of Berlin were joint editors.
This work is, however, so near completion that it will probably be but
slightly delayed. He also edited the Moss magazine, “ Hedwigia.”
Edward Parfitt, a student of the Geology of Devonshire, England,
died June 15, 1893, in his 73d year.
594 The American Naturalist. [June,
The Marine Biological Laboratory at Woods Hall, Mass,
INCORPORATED IN 1888. SrxTH Season, 1893.
OFFICERS OF INSTRUCTION.
C. O. Whitman, Director, Head Professor of Zoology the University
of Chicago ; Editor of the Journal of Morphology.
ZOOLOGY.
A. Investigation.
Howard Ayers, Director of the Allis Lake Laboratory, Milwaukee ;
J. Playfair McMurrich, Professor of Biology, University of Cincinnati ;
E. G. Conklin, Professor of Biology, Ohio Wesleyan University ; E
R. Lillie, Fellow in Zoology, Chicago University.
B. Instruction.
H. C. Bumpus, Professor of Comparative Anatomy, Brown Univer-
sity; W. M. Rankin, Instructor in Zoology, Princeton College; Pierre
A. Fish, Instructor in Physiology and Anatomy, Cornell University ;
A. D. Mead, Fellow in Zoology, University of Chicago.
BOTANY.
W. A. Setchell, Instructor in Botany, Yale University; W. J. V.
Osterhout, Brown University.
PHYSIOLOGY.
Jacques Loeb, Assistant Professor of Physiology, University of
Chicago.
Ryoiche Takano, Artist; F. W. Walmsley, Collector; G. M. Gray,
Laboratory Assistant.
In addition to the regular courses of instruction in Zoology, Botany,
Embryology and Microscopical Technique, consisting of lectures and
laboratory work under the constant supervision of the instructors,
there will be a number of lectures on special subjects by members of the
staff. A course of lectures in Embryology will be given by Professor
Whitman; on the Morphology of Vertebrate Head, by Dr. Ayers, and
two or more courses in Invertebrate Zoology, by Drs. Bumpus, Mc-
-Murrich, Rankin and Morgan.
There will also be ten or more evening lectures on biological subjects
of general interest. Among those who may contribute these lectures
may be ee in addition to the instructors above-named, the fol-
a ieee
Tee ae I? | GR RT Sa a eae Ra
1893.] Scientific News. 595
Drs. E. A. Andrews, Johns Hopkins University; Howard Ayers, of
the Allis Lake Laboratory; Professors W. G. Farlow, Harvard Uni-
versity ; William Libbey, Jr., Princeton College; J. M. Macfarlane,
University of Pennsylvania; C. S. Minot, Harvard Medical School ;
E. S. Morse, Salem ; H. F. Osborn, Columbia College ; John A. Ryder,
University of Pennsylvania; W. T. Sedgwick, Mass. Institute of Tech-
nology; E. B. Wilson, Columbia College.
The Laboratory is located on the coast at Woods Holl, Massachu-
setts, near the Laboratories of the United States Fish Commission.
The building consists of two stories, and has 33 private laboratories for
investigators and 5 general laboratories—two for beginners in investi-
gation in Zoology, one for teachers and students receiving instruction
in Zoology, one for Botany, and one for Physiology. The Laboratory
has aquaria supplied with running sea water, boats, a steam launch,
collecting apparatus and dredges; it is also supplied with reagents,
glassware and a limited number of microtomes and microscopes. No
aleohol can be supplied beyond what is required for work in the labora-
tory.
> the munificence of friends, the library will be provided not only
with the ordinary text-books and works of reference, but also with the
more important journals of zoology and botany, some of them in com-
plete series.
THE LABORATORIES FOR INVESTIGATORS
will be open from June 1 to August 30. They will be equipped with
aquaria, glassware, reagents, etc., but microscopes will not be provided.
In this department there are 33 private laboratories for the exclusive
use of investigators. ;
Those who are prepared to begin original work under the guidance
of instructors, will occupy tables in the general Laboratories for Inves-
tigators, paying for the privilege a fee of fifty dollars. The number of
such tables is limited to 20. ; >
An Elementary Course in vertebrate embryology will be introduced
this season, designed to meet the needs of those who have completed
the general courses in the Students’ Laboratory. The study will be
confined mainly to the fish egg, as the best type for elucidating verte-
brate development. Each member of the class will be supplied with
material, and be expected to work out each step in the development
from the moment of fecundation. The aim will be not only to master
the details of development, but also to acquire a thorough knowledge
of the methods of work. Methods of preparing surface views, imbed-
596 The American Naturalist. [June,
ding in paraffin and celloidin, various methods of staining and mount-
ing, drawing, reconstructing, modeling, etc. The course will thus
combine just what is needed as a preparation for investigation.
This course will open Wednesday, July 5, and continue six weeks,
and it will be conducted by Mr. Lillie and Professor Whitman. The
fee for this course will be fifty dollars, and the class be limited to ten.
Applicants should state what they have done in preparation for
such a course, and whether they can bring a complete outfit, viz.: a
compound microscope, a dissecting microscope (the Paul Mayer pat-
tern made by Zeiss is the best), camera-lucida, microtome, etc. In
case these instruments are furnished by the Laboratory, an additional
fee of ten dollars will be charged therefor. No application for less
than the whole course will be granted.
THE ZOOLOGICAL LABORATORY FOR TEACHERS AND STUDENTS
will be opened on Wednesday July 5, for regular courses of six weeks
in Zoology and Microscopical Technique. The number admitted to
this department will be limited to fifty, and preference will be given to
teachers and others already qualified. By permission of the Director
and by the payment of additional fees, students may begin their indi-
vidual work as early as June 15, but the regular instruction will not
begin before July 5.
` Though more advanced students who may wish to limit their work
to special groups will have an opportunity to do so, the regular course
in Zoology, in charge of Professor Bumpus, will embrace a study of
the more typical marine forms and elementary methods of Microscopi- `
eal Technique. The Laboratory work, outlined below, will be accom-
panied by lectures.
July 5-8, Study of the Lobster (general anatomy, methods of inject-
ing, preparation of histological material) ; July 10-15, Coelenterates
(Campanularia, Tubularia, Metridium, Mnemeopsis); July 17-22, -
Vermes (Nereis, Balanoglossus and Phascolosoma, Polyzoa, Bdelloura) ;
July 24-29, Echinoderms (Asterias, Arbacia, Echinarachinus, Thyone);
- Molluscs (Venus, Sycotypus, Loligo); July 31-Aug. 5, Crustaceans
(Branchipus, Pandarus, Lepas, Idotea, Talorchestia, Cancer, Limulus) ;
August 7-15, Vertebrates (Amphioxus, Raja, Teleost).
The tuition fee is thirty-five dollars, payable in advance. Appli-
cants should state whether they can supply themselves with simple and
compound microscopes, or whether they wish to hire. Microscope
slides, dissecting and drawing instruments, bottles, and other supplies,
-to be finally taken away, are on sale at the Laboratory. Further
1893,] Scientific News. 597
information in regard to this department may be had by addressing
Professor Hermon.C. Bumpus, Woods Holl, Mass., to whom applica-
tions for admission should also be made.
THE BOTANICAL LABORATORY FOR TEACHERS AND STUDENTS
will be opened on Wednesday, July 5. The laboratory work in Botany
will be restricted to the study of the structures and development of
types of the various orders of the Cryptogamous Plants. Especial
attention will be given to the study of the various species of Marine
Algz which occur so abundantly in the waters about Woods Holl,
and students desiring to give their entire attention to these plants will be
encouraged to do so, The Fungi and Higher Cryptogams will receive
less attention than the Algae, and will be studied in fewer types. Lec-
tures will accompany the laboratory work. The course may be out-
lined somewhat as follows :
First week, Cyanophycee—Lyngbya, Calothrix, Rivularia, Stigo-
nema, Tolypothrix, Anabena. Second week, Chlorophycee—Spiro-
gyra, Ulva, Enteromorpha, Chetomorpha, Bryopsis, Vaucheria,
(Edogonium ; Phaeophyceae, Ectocarpus, Mesogloia, Leathesia, Lami-
naria, Fucus, Sargassum. Third week, Rhodophycee—Batrachosper-
mum, Nemalion, Callithamnion, Chondriopsis, Rhabdonia. Fourth
week, Phycomycetes — Mucor, Sporodinia, Peronospora, Cystopus,
Achlya; Uredinei, Æcidium, Uredo, Puccinia, Uromyces. Fifth
week, Basidiomycetes—Agaricus, Lycoperdon; Ascomycetes—Micro-
sphæra, Sordaria, Peziza, Physcia. Sixth week, Muscinee—Riccia,
-= Madotheca, Marchantia, Mnium, Tetraphis, Hypnum; Filicineæ—
Dicksonia, Adiantum, Equisetum, Lycopodium, Marsilia, Selaginella.
The tuition for students in the regular course of laboratory work
and lectures is thirty-five dollars, payable in advance; for students
engaged in investigation the tuition is fifty dollars.
Students are expected to supply their own instruments, or to pay an
extra fee for those borrowed from the Laboratory. Applications
should be addressed to William A. Setchell, 2 Hillhouse Ave., New
Haven, Conn.
The Physiological Laboratory will be open from June 1 to Septem-
investigators.
bine ai two persons may be obtained near the Lab-
oratory, at prices varying from $2.00 to $4.00 a week, and board from
from $4.50 to $6.00. By special t, board will be supplied
to members at The Homestead at $5.00 a week.
8 The Ameritun Naturalist. (June,
z A DEPARTMENT OF LABORATORY SUPPLY
has been established in order to facilitate the work of teachers and
others at a distance, who desire to obtain materials for study or for
Ra instruction. Certain sponges, hydroids, starfishes, sea-urchins,
ne worms, crustaceans, molluscs, and vertebrates are generally —
peut in stock, though larger orders should be filled some time before
the material is needed. Circulars giving information, prices, etc., may
be obtained by addressing the Collector, Mr. F. W. Walmsley, Woods
Holl, Mass. ;
ADVERTISEMENTS.
A Tonie
For Brain-Workers, the Weak and
bilitated.
Horsford’s Acid Phosphate
is without exception the Best
Remedy for relieving Mental
and Nervous Exhaustion; and
where the system has become
debilitated by disease, it acts as
a general tonic and vitalizer,
affording sustenance to both
brain and body.
Dr. J. C. Wilson, Philadelphia,
Pa., says: “Ihave used it as a gen-
eral tonic, and in particular in the
-debility and dyspepsia of overworked |
men, with satisfactory results.”
Descriptive pamphlet free.
Rumford Chemical Works, Providence, R. I.
yore of Substitutes and Imitations.
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ll. Animal Effigies and Emblematic Mounds.
IH. Native Moths and Symbols—Unfinished.
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