Lorence

Arachnothryx bourgaei (Standi.) Borhidi, Acta

Bot. Acad. Sci. Hung. 28; 69. 1982.

Rondeletia bourgaei Standi., N. Amer. FI. 32:

56. 1918. Type; MEXICO: Veracruz, region

de Orizaba, Monte San Cristobal, 8 Jul 1866,

E. Bourgeau 2618 (Holotype US 49964

(photos MEXU, PTBG); Isotypes P, 4 sheets

(photos MEXU, PTBG)).—Rondeletia

laniflora Benth.

Arachnothryx brenesii (Standi.) Borhidi, Acta

Bot. Hung. 33: 301. 1987. Rondeletia brenesii

Standi., Publ. Field Mus. Nat. Hist., Bot. Ser.

18: 1370. 1938. Type: COSTA RICA:

Camino de San Ramon a La Calera, 20 Mar

1929, A. M. Brenes 6757 (Holotype F 854972

(photo PTBG)).—Rondeletia brenesii Standi.

Arachnothryx brevicorolla (J. H. Kirkbr.)

Borhidi, Acta Bot. Acad. Sci. Hung. 28: 69.

1982 [as brevicollis]. Rondeletia salicifolia

Dwyer & Hayden subsp. brevicorolla J. H.

Kirkbr., Ann. Missouri Bot. Gard. 55: 383.

1963. Type: PANAMA: Bocas del Toro,

cloud forest above Quebrada Huron on Cerro

Bonyik, 166-^00 m, 13 Apr 1968, J. H.

Kirkbride, Jr. & J. A. Duke 597 (Holotype

MO 1946832 (photo PTBG); Isotypes NY,

n.v., US (photos MEXU, PTBG)).

—Rondeletia bertieroides Standi.

Arachnothryx buddleioides (Benth.) Planch., FI.

Serres 5: tab. 442. 1849. Rondeletia

buddleioides Benth., PI. Hartweg. 69. 1840.

Type: MEXICO: Oaxaca, in planitie Llano

Verde, T. K. Hartweg 502 (Holotype K (photo

PTBG); Isotypes NY, n.v. (fragment US

(photos MEXU, PTBG), P (photos MEXU,

PTBG)).—Rondeletia buddleioides Benth.

Arachnothryx buddleioides (Benth.) Planch, var.

aspera (J. H. Kirkbr.) Borhidi, Acta Bot.

Hung. 33: 301. 1987. Rondeletia buddleioides

Benth. var. aspera J. H. Kirkbr., Ann.

Missouri Bot. Gard. 55: 379. 1969. Type:

PANAMA: Bocas del Toro, between Criollo

(just above Buena Vista) and Quebrada

Higueron on Chiriqui Trail, 18 Apr 1968,

J. H. Kirkbride. Jr. & J. A. Duke 780

(Holotype MO 1942973 (photos MEXU,

PTBG)).—Rondeletia buddleioides Benth.

Arachnothryx calycosa (Bonn. Sm.) Borhidi,

Acta Bot. Acad. Sci. Hung. 28: 70. 1982.

Rondeletia calycosa Bonn. Sm., Bot. Gaz.

(Crawfordsville) 56: 59. 1913. Type: COSTA

RICA: Forets de la Palma, 1460 m, 25 Sep

1898, A. Tonduz 11633 (Holotype US 941406

(photos F, MEXU, PTBG)).—Rondeletia

calycosa Dorm. Sm.

Arachnothryx capitellata (Hemsl.) Borhidi, Acta

Bot. Acad. Sci. Hung. 28: 69. 1982.

Rondeletia capitellata Hemsl., Diag. PI. Mex.

28. 1879. Type: MEXICO: Without precise

locality, Jurgensen 355 (Lectotype K, here

designated (photo PTBG); Isolectotypes G-

DEL (n.v., fragment F), K (photo PTBG)).

Syntypes: MEXICO: Oaxaca, Joya, F.

Liebmann 202 (K (photo PTBG); without

precise locality, Jurgensen 376 (K (photo

PTBG)).—Rondeletia capitellata Hemsl.

Arachnothryx chinajensis (Standi. & Steyerm.)

Borhidi, Acta Bot. Hung. 33: 301. 1987.

Rondeletia chinajensis Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254.

1947. Type: GUATEMALA: Alta Verapaz,

Cerro Chinaja, between Finca Yalpemech and

Chinaja, above source of Rio San Diego,

150-700 m, 1-2 Apr 1942, J. A. Steyermark

45637 (Holotype F 1134210 (photos MEXU,

PTBG)).—Rondeletia chinajensis Standi. «fe

Steyerm.

Arachnothryx cooperi (Standi.) Borhidi, Acta

Bot. Acad. Sci. Hung. 28; 69. 1982.

Rondeletia cooperi Standi., Publ. Field Mus.

Nat. Hist., Bot. Ser. 4: 267. 1929. Type:

PANAMA: Bocas del Toro, Buena Vista

Camp, on Chiriqui Trail, 1250 ft., Jan-Mar

1928, G. P. Cooper 600 (yziXc 12233)

(Holotype F 579596 (photo PTBG); Isotype

NY, n.v.).—Rondeletia cooperi Standi.

Arachnothryx cordovana (Standi. 8c Steyerm.)

Borhidi, Acta Bot. Hung. 33: 301. 1987.

Rondeletia cordovana Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 26.

1943. Type: GUATEMALA: Zacapa, in

ravine bordering Quebrada Alejandria,

summit of Sierra de las Minas, vicinity of

Finca Alejandria, 2500 m, 13 Oct 1939, J. A.

Steyermark 29906 (Holotype F 1042952

. Nomenclator of Mexican and Central American Rubiaceae

(photos MEXU, PTBG); Isotype US (photos

MEXU, PTBG)).—Rondeletia

brandegeeana Lorence

Arachnothryx costaricensis (Standi.) Borhidi,

Acta Bot. Acad. Sci. Hung. 28: 70. 1982.

Rondeletia costaricensis Standi., N. Amer. FI.

32: 61. 1918. Type: COSTA RICA: Buena

Vista, San Carlos, 900 m, Apr 1903, H. Pittier

16696 (Holotype US 764141 (photos MEXU,

PTBG)).—Rondeletia costaricensis Standi.

Arachnothryx darcyi (Dwyer) Borhidi, Acta

Bot. Acad. Sci. Hung. 28: 70. 1982.

Rondeletia darcyi Dwyer, Ann. Missouri Bot.

Gard. 67: 467. 1980. Type: PANAMA: Colon,

road bank just W of Portobelo, 12 Apr 1970,

W. G. D’Arcy 4079 (Holotype MO 2238438

(photo PTBG); Isotype MO (photo PTBG)).

—Rondeletia secunda Standi.

Arachnothryx dariensis (Standi.) Borhidi, Acta

Bot. Acad. Sci. Hung. 28: 68. 1982.

Rondeletia dariensis Standi., N. Amer. FI. 32:

53. 1918. Type: PANAMA: Darien, Boca de

Paurando, on Sambu River, southern Darien,

20 m, Feb 1912, //. Pittier 5684 (Holotype US

867136 (photos MEXU, PTBG); Isotype F

(photo PTBG)).—Rondeletia dariensis

Standi.

Arachnothryx deamii (Donn. Sm.) Borhidi, Acta

Bot. Acad. Sci. Hung. 28: 68. 1982.

Bouvardia deamii Donn. Sm., Bot. Gaz.

(Crawfordsville) 49: 455. 1910. Type:

GUATEMALA: At base of Rillo de las

Nabajas, about Va mile front side of Fiscal,

3700 ft., 5 Jun 1909, C. C. Beam 6190

(Holotype US 579579 (photos MEXU,

PTBG); Isotypes A, n.v., F (photo PTBG),

GH, n.v., MO (photos MEXU, PTBG), US

(photos MEXU, PTBG)).—Rondeletia

deamii (Donn. Sm.) Standi.

Arachnothryx edwardsii (Standi.) Borhidi, Acta

Bot. Hung. 33: 301. 1987. Rondeletia

edwardsii Standi., Trop. Woods 37: 31. 1934.

Type: HONDURAS: Comayagua, Pito Solo,

Lake Yojoa, 2000 ft., 29 Aug 1932, J. B.

Edwards P476 (Holotype F 662645 (photos

MEXU, PTBG); Isotype A, n.v.).

—Rondeletia edwardsii Standi.

Arachnothryx elongata (Bartl. ex DC.) Planch.,

FI. Serres 5: tab. 442. 1849. Rondeletia

elongata Bartl. ex DC., Prodr. 4: 409. 1830.

Type: MEXICO: Guerrero, Haenke s.n.

(Holotype presumably PR, n.v.; Isotype W,

n.v. (photo MO)).—Rondeletia leucophylla

Kunth

Arachnothryx falciformis (Lundell) Borhidi,

Acta Bot. Hung. 35: 310. 1989. Rondeletia

falciformis Lundell, Wrightia 5: 324. 1976.

Type: MEXICO: Hidalgo, km 332 of Mexico

City-Laredo highway, above Chapulhuacan,

3800 ft., 13 Jul 1943, C. L Lundell 12234

(Holotype TEX-LL (photos MEXU, PTBG);

Isotype F (photo PTBG)).—Rondeletia

Arachnothryx galeottii (Standi.) Borhidi, Acta

Bot. Acad. Sci. Hung. 28: 70. 1982.

Rondeletia galeottii Standi., N. Amer. FI. 32:

59. 1918. Type: MEXICO: Without precise

locality, H. Galeotti s.n. (Holotype US

454830 (photos MEXU, PTBG)).

—Rondeletia galeottii Standi.

Arachnothryx gonzaleoides (Standi.) Borhidi,

Acta Bot. Hung. 33: 302. 1987 [as

gonzalaeoides]. Rondeletia gonzaleoides

Standi., Publ. Field Columbian Mus., Bot. Ser.

8: 160. 1930. Type: MEXICO: Chiapas,

Pueblo Nuevo, 2200 m, 1840, J. Linden 149

(Holotype LE, n.v. (fragment F, photos

MEXU, PTBG); probable Isotype P (photos

MEXU, PTBG)).—Rondeletia gonzaleoides

Standi.

Arachnothryx gracilis (Hemsl.) Borhidi, Acta

Bot. Acad. Sci. Hung. 28: 69. 1982.

Rondeletia gracilis Hemsl., Diag. PI. Nov.

Mexic. 53. 1880. Type: GUATEMALA: Alta

Verapaz, 3600 ft., H. von Tuerckheim 404

(Holotype K (photo PTBG)).—Rondeletia

gracilis Hemsl.

Arachnothryx gracilispica (Standi.) Borhidi,

Acta Bot. Hung. 33: 302. 1987. Rondeletia

gracilispica Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 8: 342. 1931. Type: MEXICO:

Oaxaca, Sierra San Pedro Nolasco, Talea, etc.,

1843-33, C. Jurgensen 935 (Holotype K

(photo PTBG; fragment F, photos MEXU,

PTBG)).—Rondeletia gracilispica Standi.

Escuintla, 14 Nov 1945, E. Matuda 6146

A Nomenclator of Mexican and Central American Rubiaceae

4: 31. 1940. Type: BRITISH HONDURAS

(BELIZE): El Cayo Distr., Vaca, 21 Apr

1938, P. H. Gentle 2504 (Holotype F 941520

(photos MEXU, PTBG); Isotypes A, n.v.,

TEX-LL ex MICH (photos MEXU, PTBG),

US (photos MEXU, PTBG)).—Rondeletia

stenosiphon Hemsl.

Arachnothryx macrocalyx (Standi. & Steyerm.)

Borhidi, Acta Bot. Hung. 33: 302. 1987.

Rondeletia macrocalyx Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254.

1947. Type: GUATEMALA: Izabal, Cerro

San Gil, 300-900 m, 25 Dec \9A\,J. A.

Steyermark 41864 (Holotype F 1134224

(photos MEXU, PTBG); Isotype F (photos

MEXU, PTBG)).—Rondeletia macrocalyx

Standi. & Steyerm.

Arachnothryx mexiae (Standi.) Borhidi, Acta

Bot. Hung. 33: 302. 1987. Rondeletia mexiae

Standi., Publ. Field Mus. Nat. Hist., Bot. Ser.

22: 55. 1940. Type: MEXICO: Oaxaca, Distr.

Choapan, Santa Maria, Montana Santa Maria,

1500 m, 1 Apr 1938, Y. Mexia 9261

(Holotype F 982856 (photos MEXU, PTBG);

Isotypes GH, n.v., MO (photos MEXU,

PTBG), US (photos MEXU, PTBG)).

—Rondeletia scabra Hemsl.

Arachnothryx minor (Lundell) Borhidi, Acta

Bot. Hung. 35: 310. 1989. Rondeletia minor

Lundell, Wrightia 5: 326. 1976. Type:

MEXICO: Veracruz, about 10 miles north of

Teziutlan, 6 Jan 1964, D. S. Correll & H. B.

Correll 28736 (Holotype TEX-LL (photos

MEXU, PTBG); Isotype MEXU).

—Rondeletia gracilispica Standi.

Arachnothryx myriantha (Standi. & Steyerm.)

Borhidi, Acta Bot. Hung. 33: 302. 1987.

Rondeletia myriantha Standi. & Styerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 288.

1940. Type: GUATEMALA: Suchitepequez,

Finca Moca, 3200 ft, 31 Oct 1934, A. F.

Skutch 1569 (Holotype F 813463 (photos

MEXU, PTBG); Isotypes A, n.v., US (photos

MEXU, PTBG)).—Rondeletia myriantha

Standi. & Steyerm.

Arachnothryx myriantha (Standi. & Steyerm.)

Borhidi var. armentalis (L.O. Williams)

Borhidi, Acta Bot Hung. 35: 310. 1989.

Rondeletia myriantha Standi. & Steyerm. var.

armentalis L. O. Williams, Phytologia 26:

127. 1973. Type: GUATEMALA: Quiche,

Nebaj, 5600 ft., 22 Nov 1934, A. F. Skutch

1776 (Holotype F 815811 (photos MEXU,

PTBG)).—Rondeletia myriantha Standi. &

Steyerm. var. armentalis L. O. Williams

Arachnothryx nebulosa (Standi.) Borhidi, Acta

Bot Hung. 33: 302. 1987. Rondeletia

nebulosa Standi., Trop. Woods 37: 32. 1934.

Type: HONDURAS: Tegucigalpa, La Aurora

Trail, Rosario, San Juancito, 5400 ft., 2 Apr

1932, J. B. Edwards PIO (Holotype F 662646

(photos MEXU, PTBG); Isotype A, n.v.).

—Rondeletia nebulosa Standi.

Arachnothryx nicaraguensis (Oerst.) Borhidi,

Acta Bot. Hung. 35: 310. 1989. Rondeletia

nicaraguensis Oerst., Vidensk. Meddel.

DanskNaturhist. Foren. Kjobenhavn 1852:

43. 1853. Type: NICARAGUA: Matagalpa,

Segovia, in monte Pantasmo, Jan 1848,

Oersted s.n. (Holotype C (photos F, MO, US),

fragment US).—Rondeletia nicaraguensis

Oerst.

Arachnothryx nitida (Hemsl.) Borhidi, Acta Bot.

Acad. Sci. Hung. 28: 69. 1982. Rondeletia

nitida Hemsl, Diag. PI. Nov. Mexic. 29.

1879. Type: MEXICO: Chiapas, FI. rosatres,

forets des pins, Chiapas, in fevr.. Linden 1660

(Lectotype K, here designated (photo PTBG);

Isolectotypes G-DEL, n.v., fragment F, P

(photos MEXU, PTBG)). Syntype: MEXICO:

Chiapas, Ghiesbreght 728 (K (photo PTBG),

GH, n.v.). Linden 1660 was designated as

more duplicates.—Rondeletia nitida Hemsl.

Arachnothryx oaxacana (Standi.) Borhidi, Acta

Bot. Hung. 33: 302. 1987. Rondeletia

oaxacana Standi, Field Mus. Nat. Hist., Bot.

Ser. 11: 255. 1936. Nomen superfl., based on

same type as Rondeletia mexicana (Turcz.)

Standi. Type: MEXICO: Oaxaca, without

precise locality, 3000 ft., H. Galeotti 2664

(Holotype P (photos MEXU, PTBG),

fragment F (photos MEXU, PTBG)).

—Rondeletia mexicana (Turcz.) Standi.

Arachnothryx ovandensis (Lundell) Borhidi,

Acta Bot. Hung. 35: 310. 1989. Rondeletia

A Nomenclator of Mexican and Central American Rubiaceae

GUATEMALA: Along road from Patln to

Escuintla, 1860, S. Hayes s.n. (Holotype GH,

n.v.).—Rondeletia secundiflora B. L. Rob.

Arachnothryx septicidalis (B. L. Rob.) Borhidi,

Acta Bot. Acad. Sci. Hung. 28: 69. 1982.

Rondeletia septicidalis B. L. Rob., Proc.

Amer. Acad. Arts 45: 403. 1910. Type:

MEXICO: Chiapas, Chicharras, 3000-6000

ft., 6 Feb 1896, E. W Nelson 3755 (Holotype

US 256722 (photos MEXU, PTBG); Isotype

GH, n.v.).—Rondeletia septicidalis B. L.

Rob.

Arachnothryx silvicola (L. O. Williams)

Borhidi, Acta Bot. Hung. 35: 311. 1989.

Rondeletia silvicola L. O. Williams,

Phytologia 26: 128. 1973. Type:

HONDURAS: Atlantida, Lancetilla Valley,

near Tela, 20-600 m, 6 Dec 1927-20 Mar

1928, P. C. Standley 56885 (Holotype F

581955 (photos MEXU, PTBG)).

—Rondeletia silvicola L. O. Williams

Arachnothryx skutchii (Standi. & Steyerm.)

Borhidi, Acta Bot. Hung. 33: 302. 1987.

Rondeletia skutchii Standi. & Steyerm., Publ.

Field Mus. Nat. Hist., Bot. Ser. 22: 289. 1940.

Type: GUATEMALA: Suchitepequez, Finca

Moca, 4200 ft., 10 Jan 1935, A. F. Skutch

2110 (Holotype F 934088 (photos MEXU,

PTBG); Isotypes A, n.v., US (photos MEXU,

PTBG)).—Rondeletia skutchii Standi. &

Steyerm.

Arachnothryx stachyoidea (Donn. Sm.) Borhidi,

Acta Bot. Acad. Sci. Hung. 28: 69. 1982.

Rondeletia stachyoidea Donn. Sm., Bot. Gaz.

(Crawfordsville) 42: 298. 1906. Type:

GUATEMALA: Alta Verapaz, Semococh,

600-900 m, 25 Apr 1905, R. S. Hay s.n.

(Holotype US 473726 (photos MEXU,

PTBG)).—Rondeletia stachyoidea Donn.

Sm.

Arachnothryx standleyana (A. R. Molina)

Borhidi, Acta Bot. Hung. 33: 302. 1987.

Rondeletia standleyana A. R. Molina, Ceiba

1: 262. 1951. Type: HONDURAS: Morazan,

in pine forest above Zambrano, 1200 m, 20

Jul 1949, L. O. Williams & A. Molina R.

7447 7 (Holotype US 2216018 ex EAP

(photos MEXU, PTBG); Isotype F (photos

MEXU, PTBG)).—Rondeletia standleyana

A. R. Molina

Arachnothryx stenostachya (Standi.) Borhidi,

Acta Bot. Hung. 33: 303. 1987. Rondeletia

stenostachya Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 18: 1372. 1938. Type: COSTA

RICA: La Hondura de San Jose, 1300 m, 15

Aug 1933, M. Valerio 769 (Holotype F

681816 (photos MEXU, PTBG)).

—Gonzalagunia brenesii Standi.

Arachnothryx tacanensis (Lundell) Borhidi,

Acta Bot. Hung. 33: 303. 1987. Rondeletia

tacanensis Lundell, Bull. Torrey Bot. Club 66:

603. 1939. Type: MEXICO: Chiapas, Volcan

Tacana West, 2800 m, 30 Mar 1939, E.

Matuda 2928 (Holotype TEX-LL (photos

MEXU, PTBG); Isotypes A, n.v., F (photos

MEXU, PTBG), GH, n.v., MEXU, US

(photos MEXU, PTBG)).—Rondeletia

tacanensis Lundell

Arachnothryx thiemei (Donn. Sm.) Borhidi,

Acta Bot. Acad. Sci. Hung. 28: 69. 1982.

Rondeletia thiemei Donn. Sm., Bot. Gaz.

(Crawfordsville) 42: 299. 1906. Type:

HONDURAS: Santa Barbara, Rio Chameleon,

400 m, 16 Dec 1888, C. Thieme 5276

(Holotype US 941403 (photos MEXU,

PTBG)).—Rondeletia thiemei Donn. Sm.

Arachnothryx torresii (Standi.) Borhidi, Acta

Bot. Hung. 33: 303. 1987. Rondeletia torresii

Standi., J. Wash. Acad. Sci. 18: 163. 1928.

Type: COSTA RICA: Alajuela, Viento

Fresco, 1600-1900 m, 13 Feb 1926, P. C.

Standley & R. Torres R. 47839 (Holotype US

1305295 (photos MEXU, PTBG)).

—Rondeletia torresii Standi.

Arachnothryx tuxtlensis (Lorence & Cast.-

Campos) Borhidi, Acta Bot. Hung. 35: 311.

1989. Rondeletia tuxtlensis Lorence & Cast.-

Campos, Biotica 13: l4S,fig. 7. 1988. Type:

MEXICO: Veracruz, Mun. Mecayapan,

Volcan San Martin Pajapan al sur del Ejido La

Valentina, 700 ra, 8 May 1985, J. I. Calzada

11183 (Holotype XAL; Isotypes F, MEXU,

MO (photo PTBG)).—Rondeletia tuxtlensis

Lorence & Cast.-Campos

Arachnothryx uxpanapensis (Lorence & Cast.-

Campos) Borhidi, Acta Bot. Hung. 35: 311.

Lorence

“Mexique,” photos MEXU, PTBG)).

—Randia monantha Benth.

Basanacantha pittieri Standi., Contr. U.S. Natl.

Herb. 18: 134. 1916. Type: COSTA RICA:

Gent Junction, pres Matina, Fev 1900, H.

Pittier 16036 (Holotype US 578473 (photos

K, MEXU, PTBG); Isotypes F, P (fragment F,

photos MEXU, PTBG)).—Randia pittieri

(Standi.) Standi.

Basanacantha pringlei S. Watson, Proc. Amer.

Acad. Arts 18: 98. 1883. Type: MEXICO:

Coahuila, Jimulco, mountains, 9 Apr 1885,

C. G. Pringle 142 (Holotype GH, n.v.;

Isotypes A, n.v., BM, F (photos MEXU,

PTBG), MEXU, P (photos MEXU, PTBG),

US (photos MEXU, PTBG)).—Randia

pringlei (S. Watson) A. Gray

Basanacantha reticulata S. Watson, Proc.

Amer. Acad. Arts 21: 379. 1886. Type:

MEXICO: Coahuila, S. Lorenzo de Laguna,

Feb-Oct 1880, E. Palmer 393 (Holotype US

47705 (photos MEXU, PTBG)).—Randia

pringlei (S. Watson) A. Gray

Basanacantha spinosa (Jacq.) K. Schum. var.

guatemalensis K. Schum. ex Loes., Verhandl.

Bot. Ver. Brandenburg 65: 110. 1923. Type:

GUATEMALA: “La Antigua, above

Alotengo,” E. Seler & C. Seler 2570

(Holotype B, presumably destroyed).

—Randia armata (Sw.) DC.

Basanacantha subcordata Standi, Contr. U.S.

Natl. Herb. 18: 133. 1916. Type: COSTA

RICA: Buissons et bois des chemins a Nicoya,

May 1900, A. Tonduz 13934 (Holotype US

861251 (photos MEXU, PTBG); Isotypes B,

destroyed (fragment F, photos MEXU,

PTBG), GH, n.v., LE, n.v. (fragment F, photo

PTBG), P (photos MEXU, PTBG; fragment F

photos MEXU, PTBG)). Randia.—Kzndiz

monantha Ben±.

Basanacantha tetracantha (Cav.) Hook, f in

tetracantha Cav., Icon. PI. 4: 20 pi 435

1799. Type: MEXICO: [Probably Guerrero]

cerca Acapulco, Nee s.n. (Holotype MA, n v

(photos F, MEXU, MO)).—Randia

tetracantha (Cav.) DC.

Bathysa C. Presl, Bot. Bemerk. 84. 1844. Type:

Bathysa stipulata C. Presl

Bathysa panamensis Dwyer, Ann. Missouri Bot.

Gard. 55: 34. 1968. Type: PANAMA: Darien,

Rio Ucurganti, 7 Jul 1967, N. Bristan 1187

(Holotype MO 1968219 (photo PTBG);

Isotype GH, n.v.).—Status uncertain.

Bathysa veraguensis Dwyer, Ann. Missouri

Bot. Gard. 67: 4Q,fig. 9. 1980. Type:

PANAMA: Veraguas, Cerro Tute, 1200 m 28

Sep 1972, E. A. Lao & A. Gentry 531

(Holotype MO 2164349 (photo PTBG)).

Belicea Lundell, Contr. Univ. Michigan Herb. 8:

86. 1942. Type: Belicea hoffmannioides

Lundell. [Herein treated as a synonym of

Morinda L.]

Belicea hoffmannioides Lundell, Contrib. Univ.

Michigan Herb. 8: 87. 1942. Type: BRITISH

HONDURAS (BELIZE): Toledo Distr.,

Swasey Branch, Monkey River, near falls, 7

Mar 1942, P. H. Gentle 3945 (Holotype TEX-

LL, n.v.; Isotypes A, n.v., MO (photo

PTBG)).—Morinda asperula Standi.

Berghesia C. G. D. Nees, Linnaea 20: 701.

1847. Type: Berghesia coccinea C. G. D.

Nees. [Genus of uncertain status. According

to Nees’s protologue description, Berghesia

belongs to “tribu Cinchoneae,” the capsule is

2-locular, and each locule is 1-seeded.]

Berghesia coccinea C. G. D. Nees, Linnaea 20:

702. 1847. Type: MEXICO, without precise

locality, De Berghes s.n. (Holotype possibly

LE, M, or GZU, n.v.).—Status uncertain.

Bertiera Aubl, Hist. PI. Guiane 1: 180. 1775, as

“Berthiera.” Type: Bertiera guianensis Aubl.

Bertiera aequaliramosa Steyerm., Mem. New

York Bot. Gard. 17: 319. 1967. Type:

PANAMA: Bocas del Toro, Region of

Almirante, Buena Vista Camp on Chiriqui

Trail at 1250 ft., Jan-Mar 1928, G. Proctor-

Cooper 586 (Holotype NY, n.v.; Isotypes F,

MO (photos MEXU, PTBG), US).—Bertiera

guianensis Aubl.

A Nomenclator of Mexican and Central American Rubiaceae

Bertiera angustifolia Benth., Bot. Voy. Sulphur

103. 1844. Type: COSTA RICA: Cocos

Island, Barclay s.n. (Holotype BM, n.v.).

Bertiera guianensis Aubl., Hist. PI. Guiane 1:

180,p/. 69. 1775. Type: FRENCH GUIANA:

“prope Aroura,” Aublet s.n. (Holotype

presumably BM, n.v.).

Bertiera guianensis var. leiophylla Steyerm.,

Mem. New York Bot. Gard. 17: 319. 1967.

Type: PANAMA: Chiriqui, vicinity of San

Felix, 0-120 m, 17 Dec 1911, H. Pittier 5178

(Holotype NY, n.v.; Isotype US (photos

MEXU, PTBG)).—Bertiera guianensis Aubl.

Bertiera oligosperma Wemham, J. Bot. 50: 162.

1912. Type: NICARAGUA: Seemann 129

(Holotype BM, n.v.).—Status uncertain.

Bertiera panamensis Standi., N. Amer. FI. 32:

180. 1934. Type: PANAMA: Darien, Cana

and vicinity, 2000-5600 ft., 17 Apr-8 Jun

1908, R. S. Williams 832 (Holotype US

678312 (photos MEXU, PTBG); Isotype NY,

n.v.).—Raritebe palicoureoides Wemham

subsp. dwyeranum J. H. Kirkbr.

Bertiera tenuis Lundell, Wrightia 4: 49. 1968.

Type: GUATEMALA: Alta Verapaz, Sebol,

in high forest about 800 m west of village,

bordering arroyo, 15 Apr 1964, E. Contreras

4318 (Holotype TEX-LL (photos MEXU,

PTBG); Isotype F).—Bertiera guianensis

Aubl.

Bertiera viburnoides (Standi.) J. H. Kirkbr.,

Brittonia 31:311. 1979. Raritebe viburnoides

Standi., Publ. Field Mus. Nat. Hist., Bot. Ser.

7: 56. 1930. Type: COLOMBIA: “Caucas, La

Gallera, Mica Valley,” Killip 7934 (Holotype

US, n.v., fragment F, n.v.; Isotypes G, n.v., Y,

n.v.).

Blepharidium Standi., J. Wash. Acad. Sci. 8: 59.

1918. Type: Blepharidium guatemalense

Standi.

Blepharidium guatemalense Standi., J. Wash.

Acad. Sci. 8: 59. 1918. Type: GUATEMALA:

Alta Verapaz, forest along Saclac River,

below Secanquim, 300 m, 7 May 1905, H.

Pittier 266 (Holotype US 472925 (photos

MEXU, PTBG); Isotype US (photos MEXU,

PTBG)).

Blepharidium mexicanum Standi., Publ. Field

Mus. Nat. Hist., Bot. Ser. 22: 108. 1940.

Type: MEXICO: Chiapas, Rvina [Ruina],

Palenque, 10-12 Jul 1939, E. Matuda 3670

(Holotype F 1011462 (photo PTBG); Isotypes

A, n.v., TEX-LL (photos MEXU, PTBG), US

(photos MEXU, PTBG)).—Blepharidium

guatemalense Standi.

Borojoa Cuatrecasas, Ser. Bot. Aplicada, Serv.

Devulg., Seer. Agric. Ganad. (Cali, Colombia)

2: 474. 1949. (Preprint of Rev. Acad.

Colombia Ci. Exact. 7: 474. 1950.) Type:

Borojoa patinoi Cuatrecasas.

Borojoa atlantica Dwyer, Ann. Missouri Bot.

Gard. 67: 46. 1980. Type: PANAMA: Colon,

2-3 miles up from the Rio Guanche, 10-20 m,

19 Jan 1973, H. Kennedy & R. Foster 2197

(Holotype MO 2163702 (photo PTBG);

Isotype MO (photo PTBG)).

Borojoa panamensis Dwyer, Phytologia 17:

446. 1968. Type: PANAMA: Code, Cerro

Pilon, El Valle, 4 Jan 1968, J. A. Duke & B. R.

Lallathin 15014 (Holotype MO 1968459

(photo PTBG); Isotype MO).

Borojoa patinoi Cuatrec., Ser. Bot. Aplicada,

Serv. Devulg., Seer. Agric. Ganad. (Cali,

Colombia) 2: 474. 1949. (Preprint of Rev.

Acad. Colombia Ci. Exact. 7: 474. 1950.)

Type: COLOMBIA: Valle, Rio Calima

(region del Choco); margen derecha, entre

Quebrada de la Brea y La Esperanza, 25 m, 19

May 1946, J. Cuatrecasas 21137 (Holotype F

1251558 (photo PTBG); Isotypes F (3 sheets),

US (photo PTBG)).

Borreria G. Mey., Prim. FI. Esseq. 79, t. 1. 1818.

Type: Borreria suaveolens G. Mey., excl. ref.

Browne Jam. 141, no. 4, nom. et typ. cons.

[Herein treated as a synonym of Spermacoce

L. In cases where generic transfers have not

yet been published, the correct genus is given

in boldface type.]

Borreria arenicola Brandegee, Univ. Calif

Publ. Bot. 10: 417. 1924. Type: MEXICO:

Chiapas, in sandy arroyo near Jalisco, Aug

1923, C. A. Purpus 8968 (Holotype UC

A Nomenclator of Mexican and Central American Rubiaceae

Borreria laevigata M. Martens & Galeotti, Bull.

Acad. Roy. Bruxelles 11: 130. 1844. Type:

MEXICO: Michoacan, Vallee de Morelia

(Valladolid de Michoacan), or Mexico, la

plaine de Jordana, pres Toluca, 8000 pieds, H.

Galeotti 2586 (Holotype BR, n.v.; Isotype P,

n.v. (photo MO)).—Spermacoce verticillata

Borreria laevis (Lam.) Griseb., Abh. Kon.

Gewiss. Gottingen 7: 231.1857. Spermacoce

laevis Lam., Tabl. Encycl. et Meth. Bot. 1:

273., n. 1435. 1791. Type: SANTO

DOMINGO: Without precise locality, Martin

s.n. (Holotype P-LA, n.v.).—Spermacoce

tenuior L. Note: Although Borreria laevis

sensu Griseb. equals Spermacoce assurgens

Ruiz & Pav., the type of the basionym,

Spermacoce laevis Lam., equals the type of

Spermacoce tenuior L.

Borreria latifolia (Aubl.) K. Schum. in Mart.,

FI. Bras. 6(6): 61., t. 80. 1888. Spermacoce

latifolia Aubl., Hist. PI. Guiana 55. 1775.

Type: FRENCH GUIANA: Cayenne, Aublet

s.n. (Holotype P, n.v.; Isotype MA, n.v.

Borreria longiseta M. Martens & Galeotti, Bull.

Acad. Roy. Bruxelles 11: 128. 1844. Type:

MEXICO: Veracruz, Zacuapan et Mirador

[label on isotype at P states “cordilleres pres

de Vera Cmz, 1000 pieds”], H. Galeotti 2611

(Holotype BR, n.v.; Isotypes K, n.v., P)).

—Status uncertain. According to C. D.

Adams (pers. comm. 1996) the description

does not agree with the type, which actually

represents a specimen of Spermacoce confusa

Rendle.

Borreria malacophylla Standi. & L. O.

Williams, Ceiba 1: 165. 1950, nom. illeg., not

Borreris malacophylla K. Schum., 1903.

Type: HONDURAS: El Paraiso, near Galera,

moist thicket, 900 m, Nov-Dec 1943, P. C.

Standley 15584 (Holotype US 2215979

(photos MEXU, NY, PTBG); Isotype F).

—Spermacoce assurgens Ruiz & Pav.

Borreria nesiotica B. L. Rob., Proc. Amer.

Acad. Arts 45: 409. 1910. Type: MEXICO:

Socorro Island (Revillagigedo Group), 1897,

A. W. Anthony s.n. (Holotype GH, n.v.;

Levin

Borreria oaxacana M. Martens & Galeotti, Bull.

Acad. Roy. Bmxelles 11: 130. 1844. Type:

MEXICO: Oaxaca, pres de la ville d’Oaxaca,

5000 pieds, H. Galeotti 2629 (Holotype BR,

Borreria ocymoides (Burm.f.) DC., Prodr. 4:

544. 1830. Type: Tropical Asia, collector

unknown, (Holotype presumably G, n.v.).

Spermacoce ocymoides Burm.f, FI. Ind. 34.

1768. This name has been erroneously applied

by many authors to tropical American taxa of

Spermacoce with small leaves and flowers,

including S. exilis, S. ovalifolia, and S.

Borreria ovalifolia M. Martens & Galeotti, Bull.

Acad. Roy. Bmxelles 11: 129. 1844. Type:

MEXICO: Veracruz, Mirador, 5000 pieds, H.

Galeotti 2606 (Holotype BR, n.v.).

—Spermacoce ovalifolia (M. Martens &

Galeotti) Hemsl.

Borreria patula M. Martens & Galeotti, Bull.

Acad. Roy. Bmxelles 11: 128. 1844. Type:

MEXICO: Veracruz, Mirador, 3000 pieds, H.

Galeotti 2608 (Holotype BR, n.v.; Isotype K,

n.v.).—Spermacoce prostrata Aubl.

Borreria podocephala DC., Prodr. 4: 542. 1840.

Type: MEXICO or CUBA: Without precise

locality or collector, based on living material

(IDC 800.719.II.1 (photo MO)).

—Spermacoce verticillata L.

Borreriaprocera Schltdl., Liimaea 9: 593. 1834.

Type: MEXICO: Veracruz, Hacienda de la

Laguna, Schiede & Deppe 250 (Holotype

HAL, n.v.).—Spermacoce suaveolens (G.

Mey.) Kuntze

Borreria pumilio Standi., Ann. Missouri Bot.

Gard. 27: 342. 1940. Type: PANAMA: Code,

Nata, ca. 50 m, 12 Sep 1938, P. H. Allen 822

(Holotype MO 1808165 (photos MEXU,

PTBG); Isotypes F (photo PTBG), GH, n.v.,

US (photo PTBG)).—Spermacoce

Borreria radicosa Brandegee, Univ. Calif Publ.

Bot. 10: 416. 1924. Type: MEXICO: Chiapas,

meadows near Jalisco, Aug 1923, C. A.

Purpus 9223 (Holotype UC 223392, n.v.;

Lorence

Isotypes GH, n.v., US (photos MEXU,

PTBG)).—Spermacoce densiflora (DC.)

Alain

Borreria rhadinophylla B. L. Rob., Proc. Amer.

Acad. Arts 45: 409. 1910. Type: BRITISH

HONDURAS (BELIZE): On dry sandy pine

ridges, 23 Oct 1905, M. E. Peck 180

(Holotype GH, n.v.).—Mitracarpus

rhadinophyllus (B. L. Rob.) L. O. Williams

Borreria scabiosoides Cham. & Schltdl.,

Linnaea 3: 318. 1828. Type: BRAZIL: Rio de

Janeiro, Bellows.n. (presumably B,

destroyed?).—Spermacoce scabiosoides

(Cham. & Schltdl.) Kuntze

Borreria setosa M. Martens & Galeotti, Bull.

Acad. Roy. Sci. Bmxelles 11: 131. 1844.

Type: MEXICO: Oaxaca, dans les forets de

palmiers et sur les dunes de la cote d’Oaxaca,

baignee par 1’Ocean Pacifique, pres du

Rancho du Rio Grande, Sep 1839, H. Galeotti

2627 (Holotype BR, n.v.; Isotype P, n.v.

(photo MO)).—Crusea setosa (M. Martens &

Galeotti) Standi. & Steyerm.

Borreria suaveolens G. Mey., Prim. FI. Esseq.

81.,/7/. y. 1818. Type: SURINAM: In arenosis

continentis circa Arouabischkreck, G. Meyer

s.n. (Holotype GOET, n.v.).—Spermacoce

suaveolens (G. Mey.) Kuntze

Borreria subulata DC. Prodr. 4: 543. 1830.

Type: MEXICO: Without precise locality, 3

Sep 1827, Berlandier 826 (Lectotype G-DEL

n.v. (photos MO, US), B (specimen annotated

by de Candolle but not cited in protologue);

Isolectotype BM, n.v.; designated by

Anderson, 1972). Crusea.—Crusen

diversifolia (Kunth) W. R. Anderson

Borreria tampicana DC., Prodr. 4: 544. 1830.

Type: MEXICO: Tamaulipas, Tampico, 1827,

Berlandier 37 bis (Holotype G-DC, n.v!

(photo MO); Isotype BM, n.v.).

—Spermacoce prostrata Aubl.

Borreria tenella (Kunth) Cham. & Schltdl.,

Linnaea 3: 317. 1828. Spermacoce tenella

Kunth in Humb., Bonpl. & Kunth, Nov. Gen.

& Sp. 3: 345. 1820. Type: Locality unknown

Humboldt s.n. (Holotype B-W, n.v. (photo

MO)).—Spermacoce suaveolens (G. Mey.)

Borreria tonalensis Brandegee, Univ. Calif.

Publ. Bot. 6: 191. 1915. Type: MEXICO:

Chiapas, Sierra de Tonala, C. A. Purpus 7369

(Holotype UC, n.v.; Isotype GH, n.v.).

—Spermacoce

Borreria vegeta Standi. & Steyerm., Publ. Field

Mus. Nat. Hist, Bot. Ser. 23: 21. 1943. Type:

GUATEMALA: Dept. Jutiapa, between

Jutiapa and La Burrera, NE of Jutiapa,

800-850 m, 1 Nov 1940, P. C. Standley

76007 (Holotype F 1091300 (photos MEXU,

PTBG)y—Spermacoce

Borreria verticillata (L.) G. Mey., Prim. FI.

Esseq. 83. 1818. Spermacoce verticillata L.,

Sp. PI. 102. 1753. Type: JAMAICA: Hort.

Cliff 33: (BM, n.v.).—Spermacoce

verticillata L.

Borreria verticillata (L.) G. Mey. var.

thymiformis B. L. Rob., Proc. Amer. Acad.

Arts 45: 410. 1910. Type: MEXICO: Oaxaca,

18 miles SW of city of Oaxaca, 7000-8000

ft., 10-20 Sep 1894, E. W. Nelson 1410

(Holotype GH, n.v.; Isotype US 574465

(photos MEXU, PTBG)).—Spermacoce

verticillata L.

Bouvardia Salisb., Parad. Lond.pl. 88. 1807.

Type: Bouvardia triphylla Salisb.

Bouvardia alexanderae A. Carter, Madrono 13:

\A2Jigs. 1, 2. 1955. Type: MEXICO: Baja

California: Distr. del Sur, Arroyo del Salto, E

of La Paz, 30 Mar 1949, A. Carter 2577

(Holotype UC 985926, n.v.; Isotypes F,

MEXU, US (photos MEXU, PTBG)).

Carterella .—Hedyotis alexanderae (A.

Carter) W. H. Lewis

Bouvardia angustifolia Kunth in Humb., Bonpl.

& Kunth, Nov. Gen. Sp. 3: 384. 1819. Type:

MEXICO: Moran (?), A. Bonpland s.n.

(Holotype P-Bonpland, n.v. (photo MO);

Isotype F (photo PTBG)).—Bouvardia

ternifolia (Cav.) Schltdl.

Bouvardia bicolor Kunze, Linnaea 20: 24. 1847.

Type: Based on plants grown from Mexican

seed (the description is taken as the type;

designated by Blackwell, 1968).—Bouvardia

multiflora (Cav.) Schult. & Schult. f

A Nomenclator of Mexican and Central American Rubiaceae

Bouvardia bouvardioides (Seem.) Standi., N.

Amer. FI. 32: 102. 1921. Hedyotis

bouvardioides Seem., Bot. Voy. Herald 296.

1856. Type: MEXICO: NW Mexico

[Durango], Sierra Madre, Seemann s.n.

(Holotype BM, n.v.: Isotypes C, n.v.,

fragment F (photo PTBG), GH, n.v.).

Bouvardia capitata Bullock, Hook. Ic. PI. t.

3296. 1935. Type: MEXICO: Mexico, Distr.

Temascaltepec, Palmar, barranca, 7 Jul 1934,

G. B. Hinton 6319 (Holotype K, n.v.; Isotypes

BM, n.v., F (photo PTBG)).

Bouvardia cataphyllaris Bullock, Hook. Ic. PI. t.

3297. 1935. Type: MEXICO: Mexico, Distr.

Temascaltepec, Temascaltepec, 1780 m, 28

Jul 1932, G. B. Hinton 1131 (Holotype K,

n.v.; Isotypes BM, n.v., F (photo PTBG), G,

n.v., GH, n.v., MICH, n.v., MEXU, MO

(photos MEXU, PTBG), US (photos MEXU,

PTBG)).—Bouvardia chrysantha Mart.

Bouvardia cavanillesii DC., Prodr. 4: 366. 1830.

Nom. superfl., based on Aeginetia multiflora

Cav.—Bouvardia multiflora (Cav.) Schult.

& Schult. f.

Bouvardia chrysantha Mart., Del. Hort.

Monac. Sem. 1848. 4. 1848. Type: MEXICO:

[Hidalgo], near Santiaguillo, Karwinsky s.n.

(Holotype presumably BR, n.v.).

Bouvardia conzattii Greenm., Proc. Amer.

Acad. Arts 39: 92. 1903. Type: MEXICO:

Oaxaca, San Felipe, 1750 m, Jul-Aug 1900,

C. Conzatti & V. Gonzales 1067 [published as

“7976"] (Holotype GH, n.v.).

Bouvardia cordifolia DC., Prodr. 4: 366. 1830.

Type: MEXICO: Without precise locality,

Sesse & Mogino s.n. (Holotype MA, n.v.; FI.

Mexic. Ic. G-DC, n.v. (photo MO)).

Bouvardia corymbosa Oerst., Vidensk. Meddel.

Naturhist. Foren Kjobenhavn 1852: 46. 1853.

Type: NICARAGUA: Without precise

locality. Oersted 11039 (Holotype C, n.v.).

—Bouvardia leiantha Benth.

Bouvardia deamii Donn. Sm., Bot. Gaz.

(Crawfordsville) 49: 445. 1910.

GUATEMALA: Fiscal, Rillo de las Nabajas,

3700 ft., 5 Jun 1909, C. C. Deam 6190

(Holotype US 579579 (photo PTBG);

Isotypes A, n.v., F (photo PTBG), GH, n.v..

MO (photo PTBG), US (photo PTBG)).

Arachnothryx .—Rondeletia deamii (Donn.

Sm.) Standi.

Bouvardia dictvoneura Standi., N. Amer. FI.

32: 109. 192f. Type: MEXICO: Chiapas,

Chicharras, 3000-6000 ft., 6 Feb 1896, E. W.

Nelson 3757 (Holotype US 256554 (photos

MEXU, PTBG); Isotype GH, n.v.).

Bouvardia discolor Hook. & Am., Bot. Beechey

Voy. 428. 1840. Type: MEXICO: Oaxaca,

inter Tehuantepec et Voca [Boca] del Monte

in dit. Oaxaca maxime australis. Sept., G.

Andrieux 334 (Holotype K, n.v.; Isotype P

(photos MEXU, PTBG)). Not Rondeletia

discolor Kunth in Humb., Bonpl. & Kunth,

1820.—Rondeletia leucophylla Kunth

Bouvardia dolichantha Loes., Verb. Bot. Ver.

Brandenburg 65: 106. 1923. Type:

GUATEMALA: Huehuetenango, Distr.

Nenton, near Uaxackanal and toward Quen

Santo, 1100 m, 9 Jul 1896, E. Seler & C. Seler

2883 (Holotype B, presumably destroyed

(photo MO); Lectotype US 1205579 (photos

MEXU, PTBG); Isolectotypes F (fragment),

GH, n.v.; designated by Blackwell, 1968).

—Bouvardia glabra Pol.

Bouvardia endlichii Loes., Repert. Sp. Nov. 18:

357. 1922. Type: MEXICO: Durango, Endlich

176a (Holotype B, presumably destroyed

(photos F, MO, US)).—Bouvardia ternifolia

(Cav.) Schltdl.

Bouvardia erecta (DC.) Standi., N. Amer. FI.

32: 110. 1921. Catesbaea erecta DC., Prodr.

4: 401. 1830. Type: MEXICO: Without

precise locality, Sesse & Mogino 560

(Holotype MA, n.v.; probable Isotype F

(photo PTBG); FI. Mexic. Ic., MA (photo

MO)).

Bouvardia flava Decne., FI. Serres 1: 215,p/.

38. 1845. Type: Based on a greenhouse-

grown plant (protologue illustration is the

type; designated by Blackwell, 1968).

—Bouvardia laevis M. Martens & Galeotti

Bouvardia flos-joannis K. Schum., Bull. Herb.

Boissier 3: 621. 1895. Type: MEXICO: Prope

Puebla, Jul, E. Seler & C. Seler 846 (Holotype

B, presumably destroyed; Isotype GH, n.v.).

—Bouvardia erecta (DC.) Standi.

Lorence

Bouvardia mollis Linden in Schltdl., Linnaea

26: 85. 1853. Type: MEXICO: Without

precise locality or collector, herb. Lipsky

(presumably LE, n.v.).—Bouvardia iaevis M.

Martens & Galeotti

Bouvardia multiflora (Cav.) Schult. & Schult.

f., Mant. Syst. Veg. 3: 118. 1827. Aeginetia

multiflora Cav., Anal. Ci. Nat. 3: 131, t. 28, f

2. 1801; Icon. 6: 52, t. 572, f 2. 1801. Type:

MEXICO: Queretaro, Nees.n. (Holotype MA,

n.v. (photo MO)).

Bouvardia myrtifolia Schltdl, Linnaea 26: 121

1853. Type: MEXICO: Hacienda de

Cocoyotle, Schiedes.n. (Holotype B,

presumably destroyed).—Bouvardia

Bouvardia nubigena Standi. & Steyerm., Publ.

Field Mus. Nat. Hist., Bot. Ser. 22: 382. 1940.

Type: GUATEMALA: Jutiapa, Volcan de

Suchitan, NW of Asuncion Mita, 2050 m, 18

Nov 1939, y. A. Steyermark 31901 (Holotype

F 1043221 (photos F, PTBG)).—Bouvardia

Iaevis M. Martens & Galeotti

Bouvardia oaxacana Standi, J. Wash. Acad.

Arts 13: 7. 1923. Type: MEXICO: Oaxaca,

Distr. Juquila, de Santa Cruz al Aguacate, 500

m, 24 Dec 1921, C. Conzatti 4513 (Holotype

US 1110842 (photos MEXU, PTBG); Isotype

GH, n.v.).

Bouvardia obovata Kunth in Humb., Bonpl. &

Kunth, Nov. Gen. Sp. PI 3: 385. 1820. Type:

MEXICO: Distr. Federal, entre Chapultepec et

Tezcoco, Jun 1803-04, A. Bonplands.n.

(Holotype presumably P-Bonpland, n.v.).

Bouvardia orizabensis Standi, Publ. Field Mus.

Nat. Hist., Bot. Ser. 8: 334. 1931. Type:

MEXICO: Veracmz, Orizaba, 1854, M.

Botteri 604 (Holotype P, n.v. (fragment F);

Isotype G, n.v.).—Bouvardia ternifolia

(Cav.) Schltdl.

Bouvardia ovata A. Gray, PI. Wright. 5: 67

1853. Type: MEXICO: Sonora, mountain

valley from San Pedro to Santa Cruz, Sep

1851, C. Wright 1117 (Holotype GH, n v ■

Isotypes MO (photos MEXU, PTBG), Us’

(photos MEXU, PTBG)).—Bouvardia

ternifolia (Cav.) Schltdl.

Bouvardia pachecoana Standi. & Steyerm.,

Publ Field Mus. Nat. Hist., Bot. Ser. 23: 22.

1943. Type: GUATEMALA: San Marcos,

Barranco Eminencia, road between San

Marcos and San Rafael Pie de la Cuesta,

2500-2700 m, 6 Feb 1941, P. C. Standley

86226 (Holotype F 1093896 (photo PTBG);

Isotypes A, n.v., US (photos MEXU, PTBG)).

—Bouvardia dictyoneura Standi.

Bouvardia pallida Standi, N. Amer. FI. 32: 102.

1921. Type: EL SALVADOR: Volcan de San

Salvador, 1600-1800 m, 7 Apr 1922, P. C.

Standley 22977 (Holotype US 1138686

(photos MEXU, PTBG); Isotype GH, n.v.).

—Bouvardia bouvardioides (Seem.) Standi.

Bouvardia purpusii Brandegee, Univ. Calif.

Publ Bot. 10: 415. 1924. Type: MEXICO:

Chiapas, Hacienda Monserrate, Sep 1923,

C. A. Purpus 9251 (Holotype UC 223423,

n.v.; Isotypes F, GH, n.v., MO (photos

MEXU, PTBG), US (photos MEXU, PTBG)).

—Bouvardia induta (B. L. Rob.) Standi.

Bouvardia quaternifolia DC., Prodr. 4: 365.

1830. Type: MEXICO: Circa urbem Mexici,

Alaman s.n. (Holotype G-DC, n.v.).

—Bouvardia ternifolia (Cav.) Schltdl.

Bouvardia quinqueflora Dehnh. in Schltdl,

Linnaea 26: 92. 1853. Type: MEXICO:

Without precise locality or collector (location

unknown).—Bouvardia chrysantha Mart.

Bouvardia quinquenervata Standi, N. Amer.

FI. 32: 108. 1921. Type: MEXICO: Chiapas,

San Bartolome, 22 Mar 1904, E. A. Goldman

769 (Holotype US 470574 (photos MEXU,

PTBG)).

Bouvardia rekoi Standi, N. Amer. FI. 32: 108.

1921. Type: MEXICO: Oaxaca, Cafetal

Montecristo (Cerro Espino), 1000 m, 7 Dec

1917, B. P. Reko 3650 (Holotype US 867149

(photos MEXU, PTBG)).

Bouvardia rosea Schltdl, Linnaea 26: 116.

1853. Type: MEXICO: Guanajuato, 22 miles

west of Xichu, road from Xichu to San Luis

de la Paz, 2300 m, 14 Jun 1957, R. McVaugh

14819 (Neotype MICH, n.v.; designated by

Blackwell, 1968).

Bouvardia rosei Standi, N. Amer. FI. 32: 109.

1921. Type: MEXICO: Durango, Sierra

Lorence

Isotype F (photo PTBG)).—Bouvardia

ternifolia (Cav.) Schltdl.

Bouvardia tolucana Hook. & Am., Bot.

Beechey Voy. 427. 1840. Type: MEXICO:

Mexico, Toluca, Andrieux 332 (Holotype K,

n.v.; Isotype W, n.v.).—Bouvardia ternifolia

(Cav.) Schltdl.

Bouvardia triflora Kunth in Humb., Bonpl. &

Kunth, Nov. Gen. Sp. 3: 386. 1820. Type:

MEXICO: In temperatis, Humboldt &

Bonplands.n. (Holotype P-Bonpland, n.v.

(fragment P, n.v.)). Bouvardia

multiflora (Cav.) Schult. & Schult. f.

Bouvardia triflora Kunth var. hirsuta M.

Martens & Galeotti, Bull. Acad. Bruxelles 1T

236. 1844. Type: MEXICO: Oaxaca,

cordillere orientale, dans les montagnes du

Rincon, 5000 pieds, H. Galeotti 2658 bis

(Holotype BR, n.v.).—Bouvardia multiflora

(Cav.) Schult. & Schult. f.

Bouvardia triphylla Salisb., Parad. Lond.,p/. 88.

1807. Nom. superfl., based on Ixora ternifolia

Cav.—Bouvardia ternifolia (Cav.) Schltdl.

Bouvardia venosissima Lundell, Bull. Torrey

Bot. Club 66: 602. 1939. Type: MEXICO:

Chiapas, Mt. Tacana, 1400 m, 17-23 Mar

1939, E. Matuda 2748 (Holotype MICH, n.v.;

Isotype MEXU).—Bouvardia dictvoneura

Standi.

Bouvardia versicolor Ker Gawl, Bot. Reg. 3: pi.

245. 1817. Type: Based on a greenhouse-

grown plant (the protologue illustration is

taken as the type; designated by Blackwell,

1968).—Bouvardia multiflora (Cav.) Schult.

& Schult. f.

Bouvardia versicolor Ker Gawl. var. graciliflora

A. Gray in S. Watson, Proc. Amer. Acad. Arts

22: 416. 1887. Type: MEXICO: Jalisco, Rio

Blanco, E. Palmer 154 (Syntype GH, n.v.);

Tequila, E. Palmer 369 (Syntype GH, n v )•

Chapala, E. Palmer 708 (Syntype GH, n.v.).

—Bouvardia multiflora (Cav.) Schult. &

Schult. f.

Bouvardia villosa Standi., N. Amer. FI. 32: 107.

1921. Type: MEXICO: Oaxaca, Alturas de

Matatlan, 1800 m, C. Conzatti I486

(Holotype US 764094 (photos MEXU

PTBG)); Isotypes G-DEL, n.v., fragment F

(photo PTBG), GH, n.v.).—Bouvardia

xylosteoides Hook. «fe Am.

Bouvardia viminalis Schltdl., Linnaea 26: 120.

1854. Type: MEXICO: Oaxaca, Monte Alban,

6000 ft., 4 Sep 1894, C. G. Pringle 4888

(Neotype MO 174074 (photos MEXU,

PTBG); Isoneotypes BM, n.v., BR, n.v., F,

GH, n.v., M, n.v., MEXU, 2 sheets, MSC,

n.v., P, n.v., UC, n.v., US (photos MEXU,

PTBG), W, n.v.; designated by Blackwell,

1968).

Bouvardia violacea Rzed., Ciencia (Mexico) 19:

82. 1959. Type: MEXICO: San Luis Potosi, J.

Rzedowski 7680 (Holotype MEXU; Isotype

ENCB).—Bouvardia rosea Schltdl.

Bouvardia xestosperma (B. L. Rob. &

Greenm.) Terrell & S. D. Koch, Novon 4:

181. 1994. Oldenlandia xestosperma B. L.

Rob. & Greenm., Proc. Amer. Acad. Arts 32:

41. 1896. Type: MEXICO: Oaxaca, open

glades. Sierra de San Felipe, 8000 ft., Jun

1894, C. G. Pringle 4692 (Lectotype GH,

n.v.; Isolectotypes BM, n.v., CM, n.v., F, K,

n.v., MEXU, 2 sheets, MICH, n.v., MO

(photos MEXU, PTBG), MSC, n.v., PH, n.v.,

US, 2 sheets, VT, n.v.; designated by Terrell,

1980).

Bouvardia xylosteoides Hook. & Am., Bot.

Beechey Voy. 428. 1840. Type: MEXICO:

Oaxaca, on slightly elevated mountains about

Mitlam [Mitla], Andrieux 335 (Holotype K,

n.v.; Isotype G-DEL, n.v., fragment F).

Buena Cav., Anal. Hist. Nat. (Madrid) 2: 278, t.

23. 1800. Type: Buena panamensis Cav. Not

Buena Pohl, 1826. [Herein treated as a

synonym of Gonzalagunia Ruiz & Pavon.]

Buena panamensis Cav., Anal. Hist. Nat.

(Madrid) 2: 279. 1800. Type: PANAMA:

Without precise locality. Nee s.n. (Holotype

MA, n.v.; Isotype F (photo PTBG)).

Gonzalea. —Gonzalagunia panamensis

(Cav.) K. Schum.

Buena skinneri Oerst., Vidensk. Meddel. Dansk

Naturhist. Foren. Kjobenhavn 1852: 48. 1853.

TYPE: NICARAGUA: Masaya, toppen af

Vulkanen Masaya, 2000-3000 ft.. Oersted

11134 (Syntypes C, n.v. (photos A, F), US);

Lorence

Type: GUATEMALA; Dept. San Marcos, Rio

Vega, near San Rafael and Guatemala-Mexico

boundary, Volcan de Tacana, 2500-3000 m,

20 Feb 1940, J. A. Steyermark 36237

(Holotype F 1043222 (photos F, PTBG);

Isotype F 1043229).

Chiococca vestita Lundell, Wrightia 5: 8. 1972.

Type: GUATEMALA: Dept. Peten, Poptun, 2

km east, in hammock in pineland, 13 Jul

1959, C. L. Lundell 16425 (Holotype TEX-

LL, n.v. (photo F)).—Chiococca semipilosa

Standi. & Steyerm.

Chiococca vestita Lundell var. glaberrima

Lundell, Wrightia 5: 8. 1972. Type:

GUATEMALA: Dept. Peten, Poptun, 2 km

east, in hammock in pineland, 13 Jul 1959,

C L Lundell 16425A (Holotype TEX-LL,

n.v. (photo F)).—Chiococca semipilosa

Standi. & Steyerm.

Chione DC., Prodr. 4: 461. 1830. Type; Chione

glabra DC.

Chione allenii L. O. Williams, Phytologia 25:

462. 1973. Type: COSTA RICA: Golfo Duke

area, vicinity of Esquinas Experiment Station,

sea level, 8 Jul 1949, P. H. Allen 5321

(Holotype F 1679433; Isotypes MO (photo

PTBG), TUL (photo PTBG)).—Chione

sylvicola (Standi.) W. C. Burger

Chione buxifolia Dwyer & Hayden, Phytologia

15: 55. 1967. Type: PANAMA: Panama,

Cerro Jefe, E. Tyson etal. 3291 (Holotype

MO 2210380 (photo PTBG); Isotypes F, NY,

Chione campanensis Dwyer, Ann. Missouri

Bot. Gard. 67: 92. 1980. Type: PANAMA:

Cerro Campana, trail to Campana Ridge, 2300

ft., 18 Sep \96%A. R. Wilbur. R. Foster.

H. Kennedy & R. Dressier 1020 (Holotype

MO 2224284 (photo PTBG)).

Chione chambersii Dwyer & Hayden, Ann.

Missouri Bot. Gard. 54: 138. 1967. Type:

PANAMA: Comarca de Bani, area near

Puerto Armuelles, between Canazo and

Cocos, 17 Jun 1957, W. /. Stern & K. L

Chambers 144 (Holotype MO 1739838;

Isotype GH (photo PTBG)).—Chomelia

microloba Donn. Sm.

Chione chiapasensis Standi., Publ. Field Mus.

Nat. Hist, Bot Ser. 22: 110. 1940. Type:

MEXICO: Chiapas, Palenque, Rvina [Ruina],

river side in virgin forest, Jul 1939, E. Matuda

3675 (Holotype F 1011456 (photo PTBG);

Isotypes A, n.v., US (photos MEXU, PTBG),

MEXU).

Chione costaricensis Standi., Publ. Field Mus.

Nat. Hist, Bot. Ser. 22: 111. 1940. Type:

COSTA RICA: Alajuela, Canton de San

Carlos, Villa Quesada, Caribbean rain forest,

825 m, 14 Mar 1939, Austin Smith FI 778

(Holotype F 980765 (photo PTBG); Isotype

US (photos MEXU, PTBG)).—Chione

sylvicola (Standi.) W. C. Burger

Chione dariensis Dwyer, Ann. Missouri Bot.

Gard. 67: 94. 1980. Type: PANAMA: Darien,

Rio Balsa, between Manene and Tusijuanda,

26 Jul 1967, /. Duke 13571 (Holotype MO

1968196 (photos MEXU, PTBG); Isotypes

MO, 2 sheets, US (photo PTBG)).

Chione guatemalensis Standi. & Steyerm.,

Publ. Field Mus. Nat Hist., Bot Ser. 22: 174.

1940. Type: GUATEMALA: Dept. Izabal,

Rio Duke, between Livingston and 6 miles up

river, on north side, 1-25 m, 14 Apr 1940,

J. A. Steyermark 39375 (Holotype F 1035017

(photo PTBG)).

Chione mexicana Standi., J. Wash. Acad. Sci.

17: 340. 1927. Type: MEXICO: Veracruz,

Zacuapan, open forests, Jun 1926, C. A.

Purpus 10757 (Holotype US 1266079 (photos

MEXU, PTBG); Isotypes A, n.v., F, 2 sheets,

MO, 2 sheets (photos MEXU, PTBG)).

Chionepanamensis Steyerm., Ceiba 3: 19.

1952. Type: PANAMA: Chiriqui, Boquete

region, Cerro Horqueta, 6500 ft., 1 Jul 1940,

C. von Hagen & W. von Hagen 2137

(Holotype F 1397595 (photo PTBG); Isotype

MO, n.v.).—Chione sylvicola (Standi.) W. C.

Burger

Chione sylvicola (Standi.) W. C. Burger,

Selbyana 12: 138. 1991. Chomelia 1 sylvicola

Standi., J. Wash. Acad. Sci. 18: 182. 1928.

Type: COSTA RICA: Heredia, Yerba Buena,

northeast of San Isidro, 2000 m, 22-28 Feb

1926, P. C. Standley & J. Valerio 49196

(Holotype US 1305902 (photo PTBG)).

A Nomenclator of Mexican and Central American Rubiaceae

Chomelia Jacq., Enum. PI. Carib. 12. 1760.

Type: Chomelia spinosa Jacq.

Chomelia atlantica Dwyer, Ann. Missouri Bot.

Card. 67: 96. 1980. Type: PANAMA: Colon,

swamp forest and bordering terra firma, ca.

0.5 km from the Rio Buenaventura, near

Portobelo, 30 Jan 1973, H. Kennedy & S. Gra

2242 (Holotype MO 2161971 (photo PTBG);

Isotype MO)).—Chomelia barbellata Standi.

Chomelia barbata Standi., J. Wash. Acad. Sci.

13: 7. 1923. Type: MEXICO: Oaxaca,

Juquila, vicinity of Chacahua, 5 m, 17 Dec

1921, C. Conzatti 4475 (Holotype US

1110841 (photo US)).

Chomelia barbellata Standi., Field Mus. Nat.

Hist., Bot. Ser. 13(7): 120. 1936. Type;

PERU: San Martin, Klug 3920 (Holotype F,

n.v.).

Chomelia brachyloba Standi., Contr. U.S. Natl.

Herb. 18: 136. 1916. Type: PANAMA:

Panama, Sabana de San Juan Corso, near

Chepo, 60-80 m, Oct 1911, //. Pittier 4673

(Holotype US 679833 (photo PTBG)).

—Chomelia spinosa Jacq.

Chomelia brachypoda Donn. Sm., Bot. Gaz.

(Crawfordsville) 47; 255. 1909. Type:

GUATEMALA: Alta Verapaz, Sasis, banks of

Rio Ogewaj, 900 m. May 1908, H. von

Tuerckheim II2253 (Holotype US 941389

(photos PTBG, US); Isotypes F (photos

MEXU, PTBG), US (photo PTBG), W).

Anisomeris.

Chomelia coclensis Dwyer, Ann. Missouri Bot.

Gard. 67: 97. 1980. Type: PANAMA: Code,

La Mesa above El Valle, 800 m, T B. Croat

25396 (Holotype MO 2210285; Isotype MO

2219529).-^Malanea colombiana Standi.

Chomelia englesingii Standi., Trop. Woods 16;

45. 1928. Type: NICARAGUA: Region of

Braggman’s Bluff, Kukalaya River bank west

of Camp 14, 2 Dec 1927, F. C. Englesing 49

(Yale 1233) (Holotype F 572606 (photo

PTBG)). Anisomeris. —Chomelia recordii

Standi.

Chomelia filipes Benth. in Oerst., Vidensk.

Meddel Dansk Naturhist. Foren. Kjobenhavn

1852: 41. 1853. Type: NICARAGUA:

Granada, Oersted s.n. (Holotype presumably

C, n.v. (photo US), fragment US (photo

PTBG)).—Chomelia spinosa Jacq.

Chomelia grandicarpa Dwyer, Ann. Missouri

Bot. Gard. 67: 97. 1980. Type: PANAMA:

Colon, 2-3 miles up Rio Guanche, lowland

rain forest, 10-20 m, 19 Jan 1973,

H. Kennedy & R. Foster 2174 (Holotype MO.

2465399 (photo PTBG); Isotype MO).

Chomelia leucophylla Dwyer, Ann. Missouri

Bot. Gard. 67: 98. 1980. Type: PANAMA:

Darien, slopes of Cerro Chucula, drainage of

Rio Pavarando, 11 Feb 1972, A. Gentry 4262

(Holotype MO; Isotype MO). Not Rondeletia

leucophylla Kunth, 1820.—Rondeletia

dwyeri Lorence

Chomelia microloba Donn. Sm., Bot. Gaz.

(Crawfordsville) 31: 114. 1901. Type:

COSTA RICA: Santo Domingo de Golfo

Dulce, Feb 1896, A. Tonduz 7048 (Herb. Mac.

Costa Rica 9874) (Holotype US 943460

(photo PTBG); Isotypes A, n.v., MO (photos

MEXU, PTBG), P (photos MEXU, PTBG),

US (3 sheets; photos PTBG)). Anisomeris.

Chomelia panamensis (Standi.) Dwyer, Ann.

Missouri Bot. Gard. 67: 100. \9%0. Antirhea

panamensis Standi., N. Amer. FI. 32; 264.

1934. Type; PANAMA: San Bias District,

21-28 Apr 1933, G. P. Cooper 238 (Holotype

F 686345 (photo PTBG); Isotype F 709955).

—Chomelia microloba Donn. Sm.

Chomelia peninsularis Dwyer, Ann. Missouri

Bot. Gard. 67: 101. 1980. Type: PANAMA:

Chiriqui, Burica Peninsula, along Quebrada

Punta de Piedra, 2 mi SE of Puerto Armuelles,

0-100 m, 1 Mar 1973, T. B. Croat 22440

(Holotype MO 2785727 (photo PTBG);

Isotype F).—Chomelia microloba Donn. Sm.

Chomelia pringlei S. Watson, Proc. Amer.

Acad. Arts 26: 137. 1891. Type: MEXICO:

San Luis Potosi, Tamasopo Canyon, 6 Aug

1890, C. G. Pringle 3209 (Holotype GH, n.v.;

Isotypes A, n.v., F (photos MEXU, PTBG),

MO (photos MEXU, PTBG), P (photos

MEXU, PTBG), US). Anisomeris.

Chomelia protracta (Bartl. ex DC.) Standi.,

Contr. U.S. Natl. Herb. 23: 1384. 1926.

Guettarda protracta Bartl. ex DC. Prodr. 4:

457. 1830. Type: MEXICO: Without precise

A Nomenclator of Mexican and Central American Rubiaceae

Condaminea petiolata Dwyer, Ann. Missouri

Bot. Card. 67: 117. 1980. Type: PANAMA:

Panama, Rio Maje, near new Bayano Lake,

<100 m, 5 May 1976, T. B. Croat 34706

(Holotype MO 2398904 (photo PTBG);

Isotype MO).—Rondeletia panamensis DC.

Cordiera A. Rich., Mem. Soc. Hist. Nat., Paris

5: 222, t. 20. 1830. (Herein considered as a

synonym of Alibertia A. Rich.)

Cordiera edulis (Rich). Kuntze, Rev. Gen. PI. 1:

279. 1891. Genipa edulis Rich., Act. Soc.

Hist. Nat. Paris 1: 107. 1792. Type: FRENCH

GUIANA: Guyane, without precise locality,

Richard s.n. (Holotype P, n.v.; Isotype F

971706 (photo PTBG)).—Alibertia edulis

(Rich.) A. Rich.

Cosmibuena Ruiz & Pav., FI. Peruv. 3: 2. 1802,

nom. cons. Type: Cosmibuena obtusifolia

Ruiz & Pav., nom. illeg., type. cons. [=

Cosmibuena grandiflora (Ruiz & Pav.)

Rusby]. Not Cosmibuena Ruiz & Pav., 1794,

nom. rej.

Cosmibuena grandiflora (Ruiz & Pav.) Rusby,

Bull. New York Bot. Gard. 4: 368. 1907.

Cinchona grandiflora Ruiz & Pav., FI. Peruv.

2: 54,;?/. 89. 1799. Type: PERU: Huanuco,

Andes de Peru, ad Pozuzo et San Antonii de

Playa grande, Ruiz Lopez & Pavon s.n.

(Holotype presumably MA, n.v.; Isotype B,

destroyed (photo F)).

Cosmibuena holdridgei Monach., Phytologia 3:

64. 1949. Type: GUATEMALA:

Huehuetenango, Barillas to Tokija, Jul 1948,

L. R. Holdridge s.n. {Monachino 519)

(Holotype NY 3187, n.v. (photo NY); Isotype

NY, n.v. (photo NY)).—Cosmibuena

matudae (Standi.) L. O. Williams

Cosmibuena macrocarpa (Benth.) Klotzsch ex

Walp., Repert. Bot. Syst. 6: 69. 1846. Buena

macrocarpa Benth., Bot. Voy. Sulphur 104, /.

38, f. 1-14. \%45.'blot Buena macrocarpa

Wedd. 1869. Type: COLOMBIA: Narino,

Island of Gorgona, Hinds 353 (Holotype K,

n.v. (photo NY)).

Cosmibuena matudae (Standi.) L. O. Williams,

Fieldiana, Bot. 31: 45. 1965. Hillia matudae

Standi., Publ. Field Mus. Nat. Hist., Bot. Ser.

22: 51. 1940. Type: MEXICO: Chiapas,

Volcan Tacana, 2000-4039 m, Aug 1938, E.

Matuda 2327 (Holotype F 980255 (photo F);

Isotypes A, n.v., NY, n.v., US (photos

MEXU, PTBG)).

Cosmibuena ovalis Standi., Contr. U.S. Natl.

Herb. 18: 137. 1916. Type: PANAMA: Code,

vicinity of Ola, 100-350 m, 7-9 Dec 1911, //.

Pittier 5074 (Holotype US 715202 (photos F,

MEXU, PTBG); Isotype NY, n.v.).

—Cosmibuena grandiflora (Ruiz & Pav.)

Rusby

Cosmibuena paludicola Standi., Contr. U.S.

Natl. Herb. 18: 137. 1916. Type: PANAMA:

Colon, vicinity of Viento Frio between Playa

Damas and Viento Frio, along beach and near

sea level, in mangrove formation, 7-8 Aug

1911,//. Pittier 4107 (Holotype US 679204

(photos F, MEXU, PTBG); Isotypes GH, n.v.,

NY, n.v.).—Cosmibuena macrocarpa

(Benth.) Klotzsch

Cosmibuena skinneri (Oerst.) Hemsl., Biol.

Centr.-Amer., Bot. 2: 12. 1881. Buena

skinneri Oerst., Vidensk. Meddel. Dansk

Naturhist. Foren Kjobenhavn 1852: 48. 1853.

Type: NICARAGUA: In monte ignivomo,

Masaya, Jan 1848, Oersted 11134 (Holotype

C, n.v. (photos A, F, MO); Isotype US (photos

MEXU, PTBG)).—Cosmibuena grandiflora

(Ruiz & Pav.) Rusby

Cosmibuena valerii (Standi.) C. M. Taylor,

Ann. Missouri Bot. Gard. 79: 897. 1992.

Hillia valerii Standi., J. Wash.‘Acad. Sci. 18:

164. 1928. Type: COSTA RICA: San Jose,

vicinity of La Palma, on the road to La

Hondura, 1500-1700 m, 17-18 Jul 1923,

W. R. Maxon & A. D. Harvey 8092 (Holotype

US 1181776 (photos F, MEXU, PTBG)).

Cosmocalyx Standi., Publ. Field Mus. Nat. Hist.,

Bot. Ser. 8: 56. 1930. Type: Cosmocalyx

spectabilis Standi.

Cosmocalyx spectabilis Standi., Publ. Field

Mus. Nat. Hist., Bot. Ser. 8: 56. 1930. Type:

MEXICO: Yucatan, without precise locality,

1917-1921, G. F. Gaumer 24270 (Holotype F

Lorence

552275 (photo PTBG), fragment US); Isotype

MO (photo PTBG)).

Coussarea Aubl, Hist. PI. Guiane 1: 98. 1775.

Type: Coussarea violacea Aubl.

Coussarea austin-smithii Standi., Publ. Field

Mus. Nat. Hist, Bot Ser. 18: 1286. 1938.

Type: COSTA RICA: Region of Zarcero,

4700 ft, 25 Aug 1937, A. Smith 4226

(Holotype F 905233 (photos MEXU, PTBG)).

Coussarea caroliana Standi., Publ. Field Mus.

Nat. Hist., Bot Ser. 22: 178. 1940. Type:

COSTA RICA: Alajuela, Canton San Carlos,

Suerre, 1000 m, 2 Mar 1939, A. Smith HI645

(Holotype F 981032 (photo PTBG); Isotype

US (photos MEXU, PTBG)).

Coussarea cerroazulensis Dwyer & Hayden,

Ann. Missouri Bot Gard. 53: 374. 1966.

Type: PANAMA: Panama, Cerro Azul, 600

m, 6 Aug 1961, J. D. Dwyer 1368 (Holotype

MO 1810161 (photos MEXU, PTBG)).

Coussarea chiriquiensis (Dwyer) C. M. Taylor

in W. C. Burger & C. M. Taylor, Fieldiana,

Bot., n.s. 33: 113. 1993. Rudgea chiriquiensis

Dwyer, Ann. Missouri Bot. Gard. 67: 476.

1980. Type: PANAMA: Chiriqui, Cerro

Colorado, along road above San Felix, 29 km

above bridge over Rio San Felix (7.9 km

above turnoff to Escopeta), 1500 m, 14 Jul

1976, T. B. Croat 37071 (Holotype MO

2798805 (photo PTBG); Isotype MO).

—Psychotria nebulosa (Dwyer) C. M. Taylor

Coussarea colonis Standi., Publ. Field Mus. Nat

Hist., Bot. Ser. 22: 179. 1940. Type:

PANAMA: Bocas del Toro, Buena Vista,

region of Almirante, Jan-Mar 1928, G. P.

Cooper 228 (Holotype F 579282 (photo

PTBG)). Nom. superfl. for Coussarea

euryphylla Standi., 1929.—Raritebe

palicoureoides Wemham subsp.

dwyerianum J. H. Kirkbr.

Coussarea curvigemma Dwyer, Phytologia 38:

215. 1978. Type: PANAMA: Barro Colorado

Island, Wheeler Trail 850, 1 Jun 1971, T. B.

Croat 14863 (Holotype MO 2050013 (photo

PTBG)).

Coussarea dariensis Steyerm., Ceiba 3: 20.

1952. Type: PANAMA: Darien, Rio Chico,

vicinity of Yaviza, 100 ft, 9-11 Jun 1947,

P. H. Allen 4576 (Holotype F 1397594 (photo

PTBG); Isotypes MO, NY, n.v.).—Coussarea

paniculata (Vahl) Standi.

Coussarea durifolia Dwyer, Ann. Missouri

Bot Gard. 67: 124. 1980. Type: PANAMA:

Panama, along El Llano-Carti Tupile road, 12

miles above Panamerican Highway, 200-500

m, 26-27 Mar 1973, R. Liesner 1180

(Holotype MO 2219526 (photo PTBG)).

Coussarea enneantha Standi., J. Wash. Acad.

Sci. 18: 282. 1928. Type: PANAMA: Darien,

Cana and vicinity, 2000-6500 ft., 17 Apr-8

Jun 1908, R. S. Williams 841 (Holotype US

678317 (photos MEXU, PTBG)).

Coussarea euryphylla Standi., Publ. Field Mus.

Nat. Hist., Bot. Ser. 4: 294. 1929. Type:

PANAMA: Bocas del Toro, Buena Vista

camp on Chiriqui trail, region of Almirante,

Jan-Mar 1928, G. P. Cooper 228 (Holotype F

579282). Dukea, Raritebe. —Raritebe

palicoureoides Wemham subsp.

dwyerianum J. H. Kirkbr.

Coussarea garciae Standi., Publ. Field Mus.

Nat Hist, Bot. Ser. 22: 180. 1940. Type:

COLOMBIA: Choco, Bahia Solano, along

Quebrada Jellita, 50-100 m, 22 Feb 1939,

E. P. Killip & H. Garcia 33524 (Holotype F

999258 (photo PTBG); Isotype US (photo

PTBG)).

Coussarea hondensis (Standi.) C. M. Taylor &

W. C. Burger, Selbyana 12: 138. 1991.

Psychotria hondensis Standi., J. Wash. Acad.

Sci. 18: 183. 1928. Type: COSTA RICA: Bois

de Rio Hondo, versant atlantique, 100 m, Aug

1901, //. Pittier 16161 (Holotype US 764151

(photos MEXU, PTBG)).

Coussarea imitans L. O. Williams, Phytologia

26: 488,yig. 1973. Type: GUATEMALA:

Izabal, along trail beginning from mile 33.23

between Dartmouth and Moralaes toward

Lake Izabal, Montana de Mico, 30-150 m, 7

Apr 1940, J. A. Steyermark 39013 (Holotype

F 1042963 (photos MEXU, PTBG)).

Coussarea impetiolaris Donn. Sm., Bot. Gaz.

(Crawfordsville) 37: 418. 1904. COSTA

RICA: La Colombiana, Llanuras de Santa

Clara, 200 m, Jun 1899, H. Pittier 7582

. Nomenclator of Mexican and Central American Rubiaceae

(Holotype US 943498 (photos MEXU,

PTBG); Isotype US 943497 (photos MEXU,

PTBG)).

Coussarea jefensis Dwyer, Ann. Missouri Bot.

Gard. 67: 128. 1980. Type: PANAMA:

Panama, Cerro Jefe, ca. 1000 m, 25 Aug

1975, 5. Mori & L. Joli 7921 (Holotype MO

2285634 (photo PTBG)).

Coussarea latifolia Standi., J. Wash. Acad. Sci.

18: 281. 1928. Type: COSTA RICA: Forets

de Tsaki, Talamanca, 200 m, Apr 1895, A.

Tonduz 9574 (Holotype US 938840 (photos

MEXU, PTBG)).

Coussarea loftonii (Dwyer & Hayden) Dwyer,

Ann. Missouri Bot. Gard. 67: 129. 1980.

Faramea loftonii Dwyer & Hayden,

Phytologia 15: 56. 1967. Type: PANAMA:

Panama, Cerro Jefe in Clusia forest at

2700-3000 ft, 27 Jan 1966, E. L. Tyson, J.

Dwyer & K. Blum 3323 (Holotype MO

1824505 (photo PTBG); Isotype MO (photo

PTBG)).

Coussarea mediocris Standi. & Steyerm., Publ.

Field Mus. Nat. Hist, Bot. Ser. 23: 248. 1947.

Type: GUATEMALA: Huehuetenango, Sierra

de los Cuchumatanes, vicinity of Maxbal,

about 17 miles north of Barillas, 1500 m,

15-26 Jul 1942, J. A. Steyermark 48732

(Holotype F 1134264 (photos MEXU,

PTBG); Isotype US (photos MEXU, PTBG)).

Coussarea mexicana Standi., Publ. Field Mus.

Nat Hist, Bot Ser. 17: 274. 1937. Type:

MEXICO: Oaxaca, Ubero, 30-60 m, Jun

1937, L. Williams 9442 (Holotype F 897721

(photos MEXU, PTBG); Isotypes P (photos

MEXU, PTBG), US (photos MEXU, PTBG)).

Coussarea morii Dwyer, Ann. Missouri Bot.

Gard. 67: 130. 1980. Type: PANAMA:

Darien, Rio Tuquesa, at middle Tuquesa

Mining Company Camp called Charco Peje,

tropical wet forest, 250 m, 8 Jul 1975, S. Mori

7058 (Holotype MO 2821790 (photo PTBG)).

Coussarea nebulosa Dwyer, Ann. Missouri Bot.

Gard. 67: 131. 1980. Type: PANAMA:

Chiriqui, uncut cloud forest at Monte Rey

near Boquete, 20 Jul 1971, T. 5. Croat 15868

(Holotype MO 2162995 (photo PTBG);

Isotype MO 4043108 (photo PTBG)).

—Psychotria nebulosa (Dwyer) C. M. Taylor

Coussarea neei Dwyer, Ann. Missouri Bot.

Gard. 67: 132. 1980. Type: PANAMA:

Veraguas, 6-7 km W of Santa Fe on new road

past agriculture school, 2900 ft., 17 Feb 1974,

M. Nee 9820 (Holotype MO 2241791 (photo

PTBG); Isotype MO 2241792 (photo PTBG)).

Coussarea nigrescens C. M. Taylor & Hammel,

Selbyana 12: \35,fig. /. 1991. Type: COSTA

RICA: Heredia, Finca La Selva, the OTS field

station near Puerto Viejo de Sarapiqui, near

the junction of the Rios Puerto Viejo and

Sarapiqui, 100 m, 5 Nov 1980, B. Hammel

10383 (Holotype DUKE, n.v.).

Coussarea paniculata Standi., J. Wash. Acad.

Sci. 18: 282. 1928, based on Froelichia

paniculata Vahl, Eclog. Amer. 1: 3. 1796. Not

Froelichia paniculata Moench., 1794. Type:

TRINIDAD: Ryan s.n. (C, n.v.).

Coussarea psychotrioides C. M. Taylor &

Hammel, Selbyana 12: U5,fig. 2. 1991.

Type: COSTA RICA: Heredia, Finca La

Selva, the OTS field station near Puerto Viejo

de Sarapiqui, near the junction of the Rios

Puerto Viejo and Sarapiqui, 100 m, 12 Jul

1979, J. H. Beach 1467 (Holotype DUKE,

n.v.; Isotypes CR, n.v., F, MO).

Coussarea roseocremea Dwyer, Ann. Missouri

Bot. Gard. 67: 133. 1980. Type: PANAMA:

Panama, El Llano-Carti Highway, 6-10 km

north of El Llano, 13 Apr 1973, R. L. Dressier

4334 (Holotype MO 2126258 (photo PTBG)).

—Psychotria roseocrema (Dwyer) C. M.

Taylor

Coussarea talamancana Standi., Publ. Field

Mus. Nat. Hist., Bot. Ser. 18: 1288. 1938.

Type: COSTA RICA: Talamanca Valley, Jun

1927, G. P. Cooper T4 (Holotype F 572698

(photo PTBG)).

Coussarea veraguensis Dwyer, Ann. Missouri

Bot. Gard. 67: 134. 1980. Type: PANAMA:

Veraguas, NW of Santa Fe, 1 km from

Escuela Agricola Alto de Piedra, on the slopes

of Cerro Tute, S. Mori & J. Kallunki 4800

(Holotype MO 2276605; Isotype MO

2276606).—Coussarea caroliana Standi.

Lorence

Juarez, Ruta 175 a 2.5 km al norte de Puerto

Eligio, 780 m, 4 Ago 1985, D. Lorence et al

4714 (Holotype MEXU; Isotypes ENCB,

MEXU, MO, UC, W, XAL).

Deppea miahuatlanica Lorence, Novon 7: 48,

fig. 2. 1997. Type: MEXICO: Oaxaca, Distr.’

de Miahuatlan, Mun. de San JCTonimo

Coatlan, 17.9 km al SW de San Jeronimo

Coatlan, carretera Miahuatlan-Piedra Larga,

1800 m, 16°12'N, 96°57W, 13 Dec 1987, R.

Torres C. A. Campos 10847 (Holotype

PTBG 15197; Isotype MEXU).

Deppea microphylla Greenm., Proc. Amer.

Acad. Arts 41: 249. 1905. Type: MEXICO:

Hidalgo, river bluffs below the Trinidad Iron

Works, 5700 ft., 26 Apr 1904, C. G. Pringle

8907 (Holotype GH (n.v.); Isotypes F (photos

MEXU, PTBG), MEXU, MO, P (photos

MEXU, PTBG), US, 2 sheets (photos MEXU

PTBG), W (photo PTBG)).

Deppea oaxacana Lorence, Allertonia 4:419

fig. 5. 1988. Type: MEXICO: Oaxaca, Distr.

de Mixe, a 9 km de la entrada a Villa Alta,

este entronque a 10 km al suroeste de

Totontepec, 2230 m, 13 Die 1985, R. Torres

C. & C. Martinez 7956 (Holotype MEXU-

Isotypes BM, ENCB, F, MEXU, MO, Uc’ W

XAL). ’ ’

Deppea obtusiflora (Benth.) Benth., PI

Hartweg. 349. 1857. Choristes obtusifiora

Benth., PI. Hartw. 63. 1840. Type: MEXICO:

Oaxaca, in planitie Llano Verde, T Hartweg

473 (Holotype K (photo PTBG); Isotypes F

(fragment from P), P (photos F, MEXU,

PTBG, US), W, 2 sheets (photos PTBG)).

Deppea panamensis Dwyer, Ann. Missouri Bot

Gard. 67: 145. 1980. Type: PANAMA:

Panama, 5-10 km northeast of Altos de

Pacora, ca. 750 m, 7 Mar 1975,5. Mori & J.

Kallunki 4965 (Holotype MO 2257147-

Isotypes MO, 2 sheets).—Elaeagia

myriantha (Standi.) C. M. Taylor & Hammel

Deppea pubescens Hemsl., Diagn. PI. Nov.

Mexic. 31. 1879; Biol. Centr.-Amer., Bot 2-

27. 1881. Type: MEXICO: Mexico, Santa

Gertrudis, F. Liebmann 65 (=Plantae

M^icanae Liebmann 11382) (Holotype K

(photo PTBG), fragment F); Isotypes GH,

n.v., MO, P (photos F, MEXU, PTBG, US),

US (photos MEXU, PTBG)).

Deppea purpurascens Lorence, Allertonia 4:

402, fig. 3. 1988. Type: MEXICO: Veracruz,

Mun. de Soteapan, Cumbres de Bastonal, ca.

10-11 km al sureste de Tebanca (lado sureste

de Lago de Catemaco) por camino, 1200 m,

26 Abr 1982, D. Lorence, R. Cedillo T & G.

Ibarra M. 4141 (Holotype MEXU; Isotypes

CHAPA, MEXU, MO).

Deppea purpusii Standi., N. Amer. FI. 32: 88.

1921. Type: MEXICO: Veracruz, Sierra

Madre between Misantla and Naolinco, Aug

1912, C. Purpus 6260 (Holotype US

464734 (photos MEXU, PTBG); Isotypes

BM, F (photo PTBG), GH, n.v., MO (photo

PTBG)).

Deppea schultzei Lorence, Allertonia 4: 422.

1988. Type: MEXICO: Guerrero,

Malinaltepec, Mar 1930, L. Schultze 363

(Holotype B, destroyed, fragment F (photos

MEXU, PTBG)). New name for Edithea

floribunda Standi., Trop. Woods 34: 1. 1933.

Not Deppea floribunda Hemsl., 1879.

Deppea scoti (J. H. Kirkbr.) Lorence, Allertonia

4: 398. 1988. Omiltemia scoti J. H. Kirkbr.,

Syst. Bot. 9: 411. 1984. Type: MEXICO:

Puebla, above Teotitlan del Camino on road to

Huautla de Jimenez, 2000-3250 m, 3 Aug

1961, C. £. Smith, Jr., F. A. Peterson & N.

Tejeda 4137 (Holotype US; Isotype GH

(photo PTBG)).

Deppea splendens Breedlove & Lorence,

Phytologia 63: 43,^g. 1. 1987. Type:

MEXICO: Chiapas, Mun. of Motozintla de

Mendoza, southwest (side) of Cerro Mozotal,

11 km northwest of junction of road to

Motozintla along road to El Porvenir and

Siltepec, 2100 m, 18 Sep 1976, D. E.

Breedlove 40258 (Holotype CAS; Isotypes

MEXU, MO (photo PTBG)).

Deppea stenosepala Standi., Publ. Field

Columbian Mus., Bot. Ser. 8:161. 1930.

Type: MEXICO: Oaxaca, Tlaxiaco a

Chicahuastla, 1844,//. Galeotti 2685

(Holotype LE, n.v., fragment F; Isotype P,

n.v., fragment F)).—Rondeletia jurgensenii

Hemsl.

A Nomenclator of Mexican and Central American Rubiaceae

Deppea tenuiflora Benth., PI. Hartw. 349.

1857. Type; MEXICO: Chiapas, San Bartolo,

Feb 1840, Linden 1644 (Holotype K; Isotype

LE, n.v., fragment F).

Deppea umbellata Hemsl., Diagn. PI. Nov.

Mexic. 31. 1879. Type: MEXICO: Veracruz,

region de Orizaba a Izhuatlancillo, 15 Mai

1866, E. Bourgeau 2428 (Lectotype P (photos

MEXU, PTBG); Isolectotypes F, K (photo

PTBG), P, 3 sheets (photos MEXU, PTBG),

US, 2 sheets; designated by Lorence &

Dwyer, 1988).

Didymaea Hook. f. in Benth. «& Hook, f, Gen.

PI. 2(1): 150. 1873. Type: Didymaea

mexicana Hook. f.

Didymaea alsinoides (Schltdl. & Cham.)

Standi., Publ. Field Mus. Nat. Hist., Bot. Ser.

18: 1291. 1938. Nertera alsinoides Schltdl. &

Cham., Linnaea 6: 413. 1831. Type:

MEXICO: Cuesta Grande de Jalacingo,

Schiede & Deppe 1271 (Holotype presumably

HAL, n.v.).

Didymaea alsinoides (Schltdl. & Cham.) Standi,

var. australis Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 18: 1292. 1938. Type: COSTA

RICA: Charpinteira [La Carpintiera], 1700 m,

Nov 1908, A. Brade & A. C. Brade 2143

(Holotype F 838717 (photo PTBG)).

—Didymaea alsinoides (Schltdl. & Cham.)

Standi.

Didymaea alsinoides (Schltdl. Cham.) Standi,

var. mollis Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 18: 1292. 1938. Type: COSTA

RICA: San Jose, Cerro de Las Vueltas,

2700-3000 m, 29 Dec 1925-1 Jan 1926, P. C

Standley & J. Valerio 43949 (Holotype US

1253422 (photos MEXU, PTBG); Isotype F

(photo PTBG)).—Didymaea alsinoides

(Schltdl. & Cham.) Standi.

Didymaea australis (Standi.) L. O. Williams,

Fieldiana: Bot. 24(11): 61. 1975. Didymaea

alsinoides (Schltdl. & Cham.) Standi, var.

australis Standi., Publ. Field Mus. Nat. Hist.,

Bot. Ser. 18; 1292. Type: COSTA RICA:

Charpinteira [La Carpintiera], 1700 m, Nov

1908, A. Brade & A. C. Brade 2143 (Holotype

F 838717 (photo PTBG)). 1938.—Didymaea

alsinoides (Schltdl. & Cham.) Standi.

Didymaea floribunda Rzed., Bol. Soc. Bot.

Mexico 44: 12, fig. 1. 1983. Type: MEXICO;

Distr. Federal, Del. Villa A. Obregon, Limbo,

2700 m, 4 Aug 1979, A. Ventura A. 3479

(Holotype ENCB).

Didymaea hispidula L. O. Williams, Fieldiana:

Bot. 24(11); 61. 1975. Type: GUATEMALA:

San Marcos, Sierra Madre Mountains about 6

km (airline) north of San Marcos, 2700 m, 13

Dec 1963, L O. Williams, A. Molina R. & T

Williams 25896 (Holotype F 1655451 (photo

PTBG))

Didymaea linearis Standi., Publ. Field

Columbian Mus., Bot. Ser. 4: 297. 1929.

Type: MEXICO: Jalisco, Real Alto, Pena

Gorda, Sierra Madre Occidental, 2500 m, 21

Feb 1927, Y. Mexia 1726 (Holotype F 579896

(photo PTBG); Isotypes A, n.v., MO (photos

MEXU, PTBG), US (photos MO, PTBG;

fragment F)).

Didvmaea mexicana Hook, f in Benth. &

Hook, f, Gen. PI. 2: 150. 1873. Syntypes:

MEXICO: Veracruz, Pico de Orizaba, 10000

ft.. Bates s.n. (K, n.v.); Orizaba, Linden 1403

(K, n.v.), Galeotti 4405 (K, n.v.), Bourgeau

3050 (K, n.v.), F. Mueller 1519 (K, n.v.), F.

Liebmann Rub. Mexic. 255 (K, n.v.).

Didymaea microflosculosa L. O. Williams,

Fieldiana: Bot. 24(11): 61. 1975. Type:

GUATEMALA: San Marcos, near Aldea

Fratemidad, between San Rafael Pie de la

Cuesta and Palo Gordo, west-facing slopes of

the Sierra Madre Mountains, 1800-2400 m,

10-18 Dec 1963, L. O. Williams, A. Molina R.

& T. Williams 25944 (Holotype F 1655450

(photo PTBG)).

Didymaea microphylla L. O. Williams,

Fieldiana: Bot. 24(11): 62. 1975. Type:

GUATEMALA: Quezaltenango, narrow ridge

of Volcan Santo Tomas, 2500-3700 m, 22 Jan

1940, J. A. Steyermark 34790 (Holotype F

1038453 (photo PTBG)).

Didymochlamys Hook, f., Icon. PI. pi. 1122.

1872. Type: Didymochlamys whitei Hook. f.

Lorence

(Holotype US 943468 (photos MEXU,

PTBG); Isotypes F, n.v., NY, n.v., US (photos

MEXU, PTBG)).—Gonzalagunia bracteosa

(Donn. Sm.) B. L. Rob.

Duggena hayesii (Wemham) Standi., Contr.

U.S. Natl. Herb. 18: 125. 1916, comb, inval.

Gonzalea hayesii Wemham, J. Bot. 51: 219.

1913. Type: PANAMA: Panama, in woods

near Panama, Hayes s.n. (Holotype BM, n.v.).

—Gonzalagunia panamensis (Cav.) K.

Schum.

Duggena nivea (Bartl. ex DC.) Standi., Contr.

U.S. Natl. Herb. 18: 125. 1916, comb, inval.

Gonzalea nivea Bartl. ex DC., Prodr. 4: 436.

1830. Type: MEXICO: Without precise

locality, Haenke s.n. (Holotype presumably

PR, n.v.). The specimen Lesta & Gomes 42

(W, n.v. (photo MO)) is not the type.

—Rondeletia buddleoides Benth.

Duggena ovatifolia (Donn. Sm.) Standi., Contr.

U.S. Natl. Herb. 18: 126. 1916, comb, inval.

Gonzalea ovatifolia Donn. Sm., Bot. Gaz.

(Crawfordsville) 27: 336. 1899. Type:

COSTA RICA: Cartago, Atirro, J. D. Smith

6599 (Lectotype US 943467 (photos MEXU,

PTBG); Isolectotypes F, n.v., GH, US).

—Gonzalagunia ovatifolia (Donn. Sm )

B. L. Rob.

Duggena panamensis (Cav.) Standi., Contr. U.S.

Natl. Herb. 18: 126. 1916, comb, inval. Buena

panamensis Cav., Anal. Hist. Nat. 2: 279.

1800. Type: PANAMA: Lancon, Nee s.n.

(Holotype MA, n.v.; Isotype F (photo

PTBG)).—Gonzalagunia panamensis (Cav.)

K. Schum.

Duggena rudis Standi., Contr. U.S. Natl. Herb.

18: 125. 1916. Type: PANAMA: Panama,

shady places along Rio Chagres at El Vigia,

12 Jan 1911, //. Pittier 2378 (Holotype US ’

676602 (photo PTBG)).—Gonzalagunia

rudis (Standi.) Standi.

Duggena thyrsoidea (Donn. Sm.) Standi., Contr.

U.S. Natl. Herb. 18: 126. 1916, comb, inval.

Gonzalea thyrsoidea Donn. Sm., Bot. Gaz.

(Crawfordsville) 13: 188. 1888. Type:

GUATEMALA: Alta Verapaz, Berge bei

Pansamala, 3800 ft.. May 1887, H. von

Tuerckheim 1249 (Holotype US 941409

(photos MEXU, PTBG)).—Gonzalagunia

thyrsoidea (Donn. Sm.) B. L. Rob.

Duhamelia Pers., Syn. 1: 203. 1805, nom.

superfl. Based on Hamelia patens Jacq. 1760.

[Herein treated as a synonym of Hamelia

Jacq.]

Dunhameliapatens (Jacq.) Pers., Syn. 1: 203.

1805. Type: SANTO DOMINGO: Jacguin

s.n. (location unknown).—Hamelia patens

Jacq.

Dukea Dwyer, Ann. Missouri Bot. Gard. 53:

360-367. 1966. Type: Dukea panamensis

Dwyer. [Herein treated as a synonym of

Raritebe Werhnam.]

Dukea blumii Dwyer, Ann. Missouri Bot. Gard.

53: 365. 1966. Type: PANAMA: Colon,

around Dos Bocas, Rio Fato Valley, H. Pittier

4223 (Holotype: MO 1227470 (photo PTBG);

Isotype US). Raritebe. —Raritebe

palicoureoides Wemham subsp.

dwyerianum J. H. Kirkbr.

Dukea darienensis Dwyer, Ann. Missouri Bot.

Gard. 53: 365. 1966. Type: PANAMA:

Darien, Chepigana District, Cana-Cuasi Trail

(Camp 2), M. E. Terry & R. A. Terry 1487

(Holotype MO 1196543 (photo PTBG);

Iso^e F)). Raritebe. —Raritebe

palicoureoides Wemham subsp.

dwyerianum J. H. Kirkbr.

Dukea euryphylla (Standi.) Dwyer, Ann.

Missouri Bot. Gard. 53: 366. 1966. Coussarea

euryphylla Standi., Publ. Field Columbian

Mus., Bot. Ser. 4: 294. 1929. Type:

PANAMA: Bocas del Toro, Almirante (or

Buena Vista Camp on Chiriqui Trail), G. P.

Cooper 228 (Holotype F, n.v.). Raritebe.

—Raritebe palicoureoides Wemham subsp.

dwyerianum J. H. Kirkbr.

Dukea panamensis Dwyer, Ann. Missouri Bot.

Gard. 53: 364. 1966. Type: PANAMA:

Darien, Cana and vicinity, 2000-6500 ft., 17

Apr-8 Jun 1908, E. L. Tyson, J. Dwyer & K.

Blum 3326 (Holotype MO 2084467 (photo

PTBG)). Raritebe. —Raritebe palicoureoides

Wemham subsp. dwyerianum J. H. Kirkbr.

A Nomenclator of Mexican and Central American Rubiaceae

ipecacuanha Brot., Trans. Linn. Soc. London

6: 137. 1802. Type: BRAZIL: Without precise

locality, Gomez s.n. (presumably LISU, n.v.).

Cephaelis .—Psychotria ipecacuanha (Brot.)

Stokes

Evea nana Standi., J. Wash. Acad. Sci. 15: 105.

1925. Cephaelis nana (Standi.) Standi. Type:

PANAMA: Canal Zone, hills N of Frijoles, 19

Dec 1923, P. C. Standley 27550 (Holotype US

1153871 (photos MEXU, PTBG); Isotype F).

Cephaelis .—Psychotria guapilensis (Standi.)

Hammel

Evea tomentosa (Aubl.) Standi., Contr. U.S.

Natl. Herb. 18: 123. 1916. Tapogomea

tomentosa Aubl., Hist. PI. Guiane 160. 1775.

Type: FRENCH GUIANA: Aublets.n.

(Holotype presumably P or BM, n.v.; Isotype

B, presumably destroyed (photo MO)).

Cephaelis .—Psychotria poeppigiana Muell.

Arg.

Exandra Standi., J. Wash. Acad. Sci. 13: 292.

1923. Type: Exandra rhodoclada Standi.

[Herein treated as a synonym of Simira Aubl.]

Exandra rhodoclada Standi., J. Wash. Acad.

Sci. 13: 292. 1923. Type: MEXICO: Oaxaca,

between La Venta and Niltepec, 200 ft., 14 Jul

1895, E. W. Nelson 2796 (Holotype US

229232 (photos MEXU, PTBG); Isotype F

(photos MEXU, PTBG)).—Simira

rhodoclada (Standi.) Steyerm.

Exostema (Pers.) Bonpl. in Humb. & Bonpl., PI.

Aequin. 1: 131. 1807 (“1808"). Lectotype:

Exostema caribaeum (Jacq.) Roem. 8c Schult.

(designated by Rogers, 1987: 165).

Exostema canescens Bartl. ex DC., Prodr. 4:

359. 1830. Type: MEXICO: Without precise

locality, Haenke s.n. (Holotype presumably

PR, n.v., fragment G-DC, n.v. (microfiche 4:

359.3)).—Status uncertain (“scarcely of this

genus” according to Standley, 1926).

Exostema caribaeum (Jacq.) Roem. & Schult.,

Syst. Veg. 5: 19. 1819. Cinchona caribaea

Jacq., Enum. PI. Carib. 16. 1760. Type:

DOMINICA: Without precise locality, De

Ponthieu s.n. (lectotype BM, n.v.; illustrated

in Lambert, Desc. Cinchona 24, t. 4. 1797;

designated by McDowell (1996b: 289)).

Howard (1989) stated that no type specimens

are known and a neotype needs to be

designated. Later, T. McDowell, in his review

of the genus, selected a lectotype from among

original material.

Exostema coulteri Hook. f. ex Hemsl., Diag. PI.

Nov. 32. 1879, [as Exostemma]. Type:

MEXICO: Hidalgo, Zimapan, Coulter 209

(Holotype K, n.v.; Isotype MO).

—Syringantha coulteri (Hook, f ex Hemsl.)

T. McDowell

Exostema indutum Standi., N. Amer. FI. 32:

126. 1921. Type: MEXICO: Oaxaca, Villa

Alta, 3000 ft., Aug, H. Galeotti 2664

(Holotype US 572997; Isotypes P, 2 sheets

(photos MEXU, PTBG)).—Exostema

Exostema longicuspe Oerst., Vidensk. Meddel.

Dansk Naturhist. Foren. Kjobenhavn 1852:

48. 1853. Type: COSTA RICA: In insula San

Lucas, Aug 1849, Oersted 11403 (Holotype

C, n.v. (photo US)).—Exostema caribaeum

(Jacq.) Roem. & Schult.

Exostema mexicanum A. Gray, Proc. Amer.

Acad. Arts 5: 180. 1861. Type: MEXICO:

[Presumably Veracruz] Prov. Huasteca,

Wartenberg, near Tantoyuca, 1858, L. C.

Ervendberg 125 (Holotype GH, n.v.; Isotype

P (photos MEXU, PTBG)).

Exostema occidentale Benth., Bot. Voy. Sulphur

104. 1844. Type: COLOMBIA: Narino, Isla

Gorgona, Barclay 354 or Hinds 354

(Holotype K, n.v.).—Rustia occidentalis

(Benth.) Hemsl.

Faramea Aubl., Hist. PI. Guian. 1: 192, t. 40.

1775. Type: Faramea corymbosa Aubl.

Faramea altipetens Dwyer, Ann. Missouri Bot.

Gard. 67: 162. 1980. Type: PANAMA:

Darien, Cerro Tacarcuna, south face of

westernmost peak, lower montane wet forest,

1500 m, 30 Jan 1975, A. Gentry & S. Mori

14013 (Holotype MO 2272402 (photo

PTBG)).

Faramea belizensis Standi., Carnegie Inst.

Wash. Publ. 461: 90. 1935. Type: BRITISH

HONDUDRAS (BELIZE): Camp 36, British

Lorence

Honduras-Guatemala survey, 2800 ft., 1 Jul

1934, fV. A. Schipp S-721 (Holotype F 733535

(photo PTBG); Isotypes A, n.v., F, G, n.v.,

MO (photos MEXU, PTBG)).—Faramea

occidentalis (L.) A. Rich.

Faramea bocataurensis Dwyer, Ann. Missouri

Bot. Gard. 67: 163. 1980. Type: PANAMA:

Bocas del Toro, eastern slope of La Zorra to

Divide, mossy cloud forest, 19 Apr 1968,

J. H. Kirkbride &J. A. Duke 806 (Holotype

MO 1968230 (photos MEXU, PTBG); Isotype

MO 2165813 (photos MEXU, PTBG)).

Faramea brachysiphon Standi., Publ. Field

Columbian Mus., Bot. Ser. 8: 62. 1930. Type:

BRITISH HONDURAS (BELIZE):

Middlesex, 200 ft., 17 Sep 1928, W. A. Schipp

345 (Holotype F 603047; Isotypes A, n.v.,

GH, n.v., MO (photos MEXU, PTBG), US

(photos MEXU, PTBG)).

Faramea bullata Standi., Publ. Field Columbian

Mus., Bot. Ser. 4: 294. 1929. Type:

PANAMA: Bocas del Toro, region of

Almirante, Kankintoe 10 miles above

Holstein, Jan-Mar 1928, G. P. Cooper 507

(Holotype F 579385 (photo PTBG); Isotype

NY, n.v.).—Faramea trinervia K. Schum.

Faramea capulifolia Dwyer in W. C. Burger &

C. M. Taylor, Fieldiana, Bot., n.s. 33: 132.

1993. Type: PANAMA: Darien, Cerro Pirre,

ridgetop at 1200 m, J. P. Folsom 4262

(Holotype MO 2620580). Note: the following

collections were also annotated as “type” by

J. D. Dwyer but are not types: COSTA RICA:

Puntarenas, Monteverde Cloud forest Reserve,

1500-1600 m, 8 Nov 1986, W. A. Haber ex E.

Bello C. 6235 (MO 36482259, photo PTBG).

PANAMA: Darien, Cerro Pirre, 10-20 Jul

1977, J. P Folsom 4435 (MO 2620712 (photo

PTBG)).

Faramea caputanguis Dwyer & Hayden, Ann.

Missouri Bot. Gard. 55: 29. 1968. Type:

PANAMA: Panama, Cerro Jefe, 2900 ft., 29

Jul 1967, J. D. Dwyer & G. Gauger 7375

(Holotype MO 1961869; Isotypes GH, n.v.,

K, n.v., MO (photos MEXU, PTBG), NY,

n.v., UC, n.v., US).—Faramea papirifolia

(Standi, ex Steyerm.) C. M. Taylor

Faramea cobana Donn. Sm., Bot. Gaz.

(Crawfordsville) 57: 422. 1914. Type:

GUATEMALA: Alta Verapaz, Coban, 1550

m, Jul 1912, //. von Tuerckheim 2474

(Holotype US 943628 (photos MEXU,

PTBG)).

Faramea correae C. M. Taylor, Novon 6: 289,

fig 1C. D. 1996. Type: PANAMA: Panama,

Parque Nacional Altos de Campana, sendero

de Interpretacion, 1 km al E del campamento

de los guardaparques de INRENARE, 8°40'N,

79°55'W, 800-900 m, 9 Sep 1993, M. D.

Correa A., E. Montenegro & E. Hidalgo 9983

(Holotype PMA 38229, n.v.; Isotypes MO

4641338, 4658116).

Faramea eurycarpa Donn. Sm., Bot. Gaz.

(Crawfordsville) 44: 113. 1907. Type:

COSTA RICA: Forets de Las Vueltas,

Tucurrique, Dec 1898,^. Tonduz 12880

(Holotype US 943627 (photos MEXU,

PTBG); Isotypes B, presumably destroyed

(fragment F), GH, 2 sheets (photos PTBG), P,

2 sheets (photos MEXU, PTBG)).

Faramea hondurae Standi., J. Wash. Acad. Sci.

18: 169. 1928. Type: COSTA RICA: San

Jose, La Hondura, 1400 m, 16 Mar 1924,

P. C Standley 37890 (Holotype US 1153105

(photos MEXU, PTBG)).

Faramea jefensis Dwyer & Hayden, Phytologia

15: 56. 1968. Type: PANAMA: Panama,

eastern slope of Cerro Jefe in Clusia forest,

2700 ft., 8 Feb 1966, E. L. Tyson et al. 3396

(Holotype MO 1936328 (photo PTBG);

Isotypes F, GH, n.v., NY, n.v., US (photo

PTBG)).

Faramea killipii Standi., Publ. Field Columbian

Mus., Bot. Ser. 7: 73. 1930. Type:

COLOMBIA: Norte de Santander, Killip &

Smith 20569 (Holotype US, n.v.).

Faramea liebmannii Standi., J. Wash. Acad.

Sci. 17: 343. 1927. Type: MEXICO: Oaxaca,

Tepitapa, F. Liebmann 11404 (Liebm.

Rubiaceae Mexic. 105) (Holotype presumably

C, n.v.; Isotype US).

Faramea liesneri Dwyer, Ann. Missouri Bot.

Gard. 67: 167. 1980. Type: PANAMA:

Panama, road to Carti (San Bias), 19 km north

of El Llano, ca. 500 m, 13 Mar 1973, P. Busey

A Nomenclator of Mexican and Central American Rubiaceae

913 (Holotype MO 2106781 (photos MEXU,

PTBG)).

Faramea loftonii Dwyer & Hayden, Phytologia

15: 56. 1967. Type: PANAMA: Panama,

Cerro Jefe, in Clusia forest, 2700-3000 ft., 27

Jan 1966, E. L. Tyson, J. D. Dwyer & K. Blum

3323 (Holotype MO 1824505 (photo PTBG);

Isotype MO (photo PTBG)).—Coussarea

loftonii (Dwyer & Hayden) Dwyer

Faramea luteovirens Standi., Contr. U.S. Natl.

Herb. 18: 138. 1916. Type: PANAMA: Colon,

Loma de la Gloria, near Fato (Nombre de

Dios), 10-104 m, Aug 1911, //. Pittier 4098

(Holotype US 679194 (photos MEXU,

PTBG)).

Faramea multiflora A. Rich., Mem. Soc. Hist.

Nat. Paris 5: 176. 1834. Type: FRENCH

GUIANA: Richard s.n. (Holotype presumably

P, n.v.).

Faramea myrticifolia Dwyer in W. C. Burger

& C. M. Taylor, Fieldiana: Bot., n.s. 33: 134.

1993. Type: COSTA RICA: Limon, Las

Brisas de Pacuarito, Siquirres, 300 m, 18 Apr

1985, L. D. Gomez, C. Herrera. E. Herrera &

W. Murillo 23401 (Holotype CR, n.v.; Isotype

MO 3648258 (photo PTBG)).

Faramea occidentalis (L.) A. Rich., Mem. Soc.

Hist. Nat. Paris 5: 176. 1834. Ixora

occidentalis L., Syst., ed. 10, 2: 893. 1759.

Type: JAMAICA: Based on P. Browne, Hist.

Jam. t. 6, f 2 (LINN, n.v.), and Plumier, Icon.

tab. 156, f 2 (ed. Burmannus). Steyermark

(1974: 947) suggested the type was from

Jamaica, thus selecting the Browne reference

over that of Plumier within the references

given by Linnaeus.

Faramea ovalis Standi., Contr. U.S. Natl. Herb.

18: 138. 1916. Type: PANAMA: Chiriqui,

forests along Rio Ladrillo and vicinity, above

El Boquete, 1200-1300 m, W. R. Maxon 5397

(Holotype US 675764 (photos MEXU,

PTBG)).

Faramea papirifolia (Standi, ex Steyerm.)

C. M. Taylor. Novon 6: 210. 1996. Cephaelis

papirifolia Standi, ex Steyerm., Acta Biol.

Venez. 4: 21. 1964. Type: COLOMBIA:

Valle, Rio Calima (region del Choco), margen

derecha, lomas frente a quebrada de la Brea,

30-50 m, 18, 22 May 1946, J. Cuatrecasas

21078 (Holotype US 2825318, n.v.; Isotypes

COL, n.v., VEN, n.v.).

Faramea papulata Dwyer & Hayden,

Phytologia 15: 57. 1967. Type: PANAMA:

Panama, Cerro Jefe, in Clusia forest,

2700-3000 ft, 27 Jan 1966, E. L Tyson. J.

Dwyer & K. Blum 3284 (Holotype MO

1831279 (photos MEXU, PTBG)).

Faramea parvibracteata Steyerm., Mem. New

York Bot. Gard. 17: 376. 1967. Type:

VENEZUELA: Terr. Fed. Amazonas, Cano

Tamatama, Rio Orinoco, 150 m, 12 Sep 1954,

5. Silverio Level 145 (Holotype NY, n.v.).

Faramea pauciflora Dwyer, Ann. Missouri

Bot. Gard. 67: 172. 1980. Type: PANAMA:

Veraguas, Valley of Rio Dos Bocas along

road between Escuela Agricola Alta Piedra

and Calovebora, 15.6 km northwest of Santa

Fe, 450-550 m, 31 Aug 1974, T B. Croat

27575 (Holotype MO 2201361 (photo

PTBG); Isotype MO (photo PTBG)).

Faramea permagnifolia Dwyer ex C. M.

Taylor, Novon 6: 19%, fig. IB. 1996. Type:

COSTA RICA: Puntarenas, Osa Peninsula, in

forest ca. 9 km W of Rincon on road to

Rancho Quemado, 8°39'N, 83°32'W, 200 m,

26 May 1986, B. Hammel, M. Grayum & G.

de Nevers 15212 (Holotype CR, n.v.; Isotypes

MO 4658114, MO 4658115).

Faramea quercetorum Standi., J. Wash. Acad.

Sci. 18: 168. 1928. Type: COSTA RICA: San

Jose, oak forest near Quebradillas, north of

Santa Maria de Dota, 1800 m, 24 Dec 1925,

P. C. Standley 42999 (Holotype US 1253066

(photos MEXU, PTBG)).—Faramea ovalis

Standi.

Faramea sanblasensis C. M. Taylor, Novon 6:

300,yzg. lA. 1996. Type: PANAMA:

Comarca de San Bias, Cerro Brewster,

headwaters of Rio Cangandi, 9° 1874,

79° 16'W, 630 m, 24 Apr 1985, G. de Nevers.

H. Herrera, B. Hammel & S. Chamley 5491

(Holotype PMA 38230, n.v.; Isotype MO

4658117).

Faramea scalaris Standi., Contr. U.S. Natl.

Herb. 13: 139. 1916. Type: PANAMA:

Chiriqui, humid forest between Altos de las

Lorence

Palmas and top of Cerro de la Horqueta,

2100-2268 m, 18 Mar 1911, //. Pittier 3265

(Holotype US 677656 (photos MEXU,

PTBG)).

Faramea schultesii Standi., Bot. Mus. Leafl.

Harvard Univ. 9: 194. 1941. Type: MEXICO;

Oaxaca, Distr. Choapam [Choapan], Cerro de

Lalana, near San Juan Lalana, in forest, long.

95°45', lat. 17°25', 1300 m, 11 May 1939,

R. E. Schultes & B. P. Reko 855 (Holotype F

1016864 (photo PTBG)).

Faramea sessifolia P. H. Allen, Rain Forests

Golfo Duke 409. 1956. Type: COSTA RICA:

Esquinas Forest, area between the Rio

Esquinas and Palmar, 200 ft., 22 May 1950,

P. H. Allen 5539 (Holotype US 2215891

(photos MEXU, PTBG); Isotypes F, 2 sheets).

Faramea standleyana L. O. Williams,

Phytologia 26: 490. 1973. Type:

GUATEMALA: Izabal, Cerro San Gil, rocky

slopes along Rio Frio, 75 m, 17 Dec 1941,

J. A. Steyermark 39962 (Holotype F 1134267

(photo PTBG)).—Faramea occidentalis (L )

A. Rich.

Faramea stenophylla Standi., J. Wash. Acad.

Sci. 18: 169. 1928. Type; HONDURAS:

Cuyamel, on hills, 29 Mar 1924, M. A.

Carleton 592 (Holotype US 1208399 (photos

MEXU, PTBG)).

Faramea stenura Standi., Publ. Field

Columbian Mus., Bot. Ser. 4: 331. 1929

Type: HONDURAS: Dept. Atlantida, in wet

forest near summit of mountains above

Lancetilla Valley, 600 m, 13 Mar 1928, P. C.

Standley 56743 (Holotype F 581582 (photo

PTBG); Isotype US 1409383 (photos MEXU

PTBG)).

Faramea suerrensis (Dorm. Sm.) Donn. Sm.,

Bot. Gaz. (Crawfordsville) 44: 112. 1907.

Faramea trinervia K. Schum. & Donn. Sm.

var. suerrensis Donn. Sm., Bot. Gaz.

(Crawfordsville) 31: 115. 1901. Type:

COSTA RICA: Suerre, Llanuras de Santa

Clara, 300 m, Feb 1896, J. D. Smith 6589

(Holotype US 943626 (photos MEXU,

PTBG)) 943624 (photos MEXU,

Faramea talamancarum Standi., Publ. Field

Columbian Mus., Bot. Ser. 4: 332. 1929.

Type: PANAMA: Talamanca Valley, 3 Feb

1921, M. A. Carleton 135 (Holotype F 578962

(photo PTBG); Isotype US (photos MEXU,

PTBG)).—Faramea multiflora A. Rich.

Faramea terryae Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 22; 189. 1940. Type:

PANAMA; Darien, Chepigana Distr.,

Cana-Cuasi Trail (Camp 2), 3000 ft., 10 Mar

1940, M. E. Terry & R. A. Terry 1479

(Holotype F 1034536 (photo PTBG); Isotype

MO (photo PTBG)).

Faramea trinervia K. Schum. & Donn. Sm.,

Bot. Gaz. (Crawfordsville) 31: 115. 1901.

Type: COSTA RICA: Talamanca, forets de la

Boca de Zhorquin, Mar 1894, A. Tonduz 8571

(Holotype US 943623 (photos MEXU,

PTBG)).

Faramea trinervia K. Schum. & Dorm. Sm. var.

suerrensis Donn. Sm., Bot. Gaz.

(Crawfordsville) 31; 115. 1901. Type:

COSTA RICA: Suerre, Llanuras de Santa

Clara, 300 m, Feb 1896, / D. Smith 6589

(Holotype US 943626 (photos MEXU,

PTBG); Isotype US 943624 (photos MEXU,

PTBG)).—Faramea suerrensis (Donn. Sm.)

Dorm. Sm.

Faramea woodsonii Standi., Aim. Missouri

Bot. Gard. 28: 470. 1921. Type: PANAMA:

Code, North rim of El Valle de Anton, near

Cerro Turega, 650-700 m, 30 Jun 1940, R. E.

Woodson, Jr. & R. W. Schery 164 (Holotype F

1085608 (photo PTBG); Isotypes F, GH, n.v.,

MO (photo PTBG)).

Faramea zetekii Standi., Contr. Arnold Arbor. 5:

147. 1933. Type: PANAMA: Canal Zone,

Barro Colorado Island, shore of cove W of

Fuertes House, 21 Feb 1932, R. H.

Woodworth &P. A. Vestal 641 (Holotype A,

n.v.; Isotypes F, MO (photo PTBG)).

—Faramea occidentalis (L.) A. Rich.

Ferdinandusa Pohl, PI. Bras. Icon. Descr. 2: 8.

1828-1829. (Substitute name for Ferdinandea

Pohl, 1827, not Ferdinanda Lagasca, 1816).

Type: Ferdinandusa speciosa Pohl

1 , Ceiba3:34. 1952. Type:

s del Toro, ^

Chiriqul Lagoon, Fish Creek, 15 Apr 1941, H.

von Wedel 2232 (Holotype F 1120423 (photo

Lorence

Acad. Arts 17: 203. 1882. Type: U.S.A.:

Arizona, Camp Crittenden, Rothrock 675

(Holotype GH, n.v.; Isotype NY, n.v.).

—Galium wrightii A. Gray

Garapatica H. Karst., Linnaea 30: 143. 1859.

Type: Garapatica edulis H. Karst.

Garapatica edulis (L. C. Rich.) H. Karst., FI.

Columb. 1: 57, t. 28. 1860. Genipa edulis

L. C. Rich., Actes Soc. Hist. Nat. Paris 1: 107.

1792. Type: FRENCH GUIANA: Guyana,

Richards.n. (Holotype P, n.v.; Isotype F

971706 (photo PTBG)).—Alibertia edulis (L.

C. Rich.) A. Rich.

Gardenia J. Ellis, Phil. Trans. 51: 935, t. 23.

1761 (nom. cons.). Type: Gardenia

jasminoides J. Ellis

Gardenia armata Sw., Prodr. FI. Veg. Ind. Occ.

51. 1788. Type: COLOMBIA: Cartagena,

Jacquin s.n. (Lectotype: Swartz’s plate;

designated by Taylor & Lorence, 1993^

—Randia armata (Sw.) DC.

Gardenia augusta (L.) Merr., Interpret.

Rumph. Herb. Amboin. 485. 1917. Varneria

augusta L., Amoen. Acad. 4: 136. 1759.

Type: Not cited; the description and

illustration may be taken as the type according

to Smith (1974). Frequently cultivated in

Mexico and Mesoamerica.

Gardenia edulis (Rich.) Poir. in Lam., Encycl.

Meth. Bot. Suppl. 2: 708. 1812. Genipa edulis

Rich., Act. Soc. Hist. Nat. Paris 1: 107.

17792. Type: FRENCH GUIANA: Guyane,

without precise locality, Richard s.n.

(Holotype P, n.v.; Isotype F 971706 (photo

PTBG)).—Alibertia edulis (Rich.) A. Rich.

Gardenia laccifera Sesse & Mo?., FI. Mexic

ed. 2, 54. 1894. Type: MEXICO: Habitat in

Ayecapixtla baud procul ab Eclesia. Floret

Aprili, Sesse & Moqiho s.n. (presumably MA,

n.v.).—Status uncertain, probably Randia

sp. (protologue states twigs have paired

spines).

Gardenia mexicana Sesse & Mo?., FI. Mexic.,

ed. 2, 54. 1894. Type: MEXICO: Habitat in

Ayacapixtla. Floret Aprili, Moqiiio

s.n. (presumably MA, n.v.).—Status

uncertain, probably Randia sp. (protologue

states twigs have 4 spines).

Gardenia schiedeana Schltdl., Linnaea 9: 601.

1834. Type: MEXICO: Veracruz, prope

Actopan, Schiede s.n. (presumably HAL,

n.v.).—Status uncertain, possibly

synonymous with Randia monantha Benth.

(1841); perhaps the oldest available epithet for

this species.

Genipa L., Syst. Nat., ed. 10, 2: 931. 1759.

Type: Genipa americana L.

Genipa americana L., Syst. Nat., ed. 10, 2:

931. 1759. Lectotype: BRAZIL: Janipaba Piso

& Marcgraf, Hist. Nat. Bras. 52, Jc. 1648.

(Lectotype designated by Howard, 1989.)

Genipa americana L. var. caruto (Kunth) K.

Schum. in Mart., FI. Bras. 6(6): 352. 1889.

Genipa caruto Kunth in Humb., Bonpl. &

Kunth, Nov. Gen. Sp. 3: 407. 1820. Type: In

riparia Orinoci et Fluminis Nigri, prope

Caracas [VENEZUELA] et Carthagena

[COLOMBIA], Humboldt & Bonplands.n.

(Holotype P-Bonpland, n.v.). For typification

see Steyermark, 1972: 349.—Genipa

americana L.

Genipa caruto Kunth in Humb., Bonpl. &

Kunth, Nov. Gen. Sp. 3: 407. 1820. Type: In

riparia Orinoci et Fluminis Nigri, prope

Caracas [VENEZUELA] et Carthagena

[COLOMBIA], Humboldt & Bonpland s.n.

(Holotype P-Bonpland, n.v.). For typification

see Steyermark, 1972: 349.—Genipa

americana L.

Genipa cinerea Femald, Proc. Amer. Acad. Arts

33: 93. 1897. Type: MEXICO: Guerrero,

Acapulco and vicinity, Oct 1894-Mar 1895,

E. Palmer 348 (Holotype GH, n.v.; Isotype

US (photos MEXU, PTBG)).—Randia

cinerea (Femald) Standi.

Genipa codonocalyx Standi., Contr. U.S. Natl.

Herb. 17: 446. 1914. Type: COSTA RICA:

Boca Matapalo, cordon littoral du Pacifique,

10 Apr 1898, H. Pittier 12085 (Holotype US

577536 (photos MEXU, PTBG); Isotype US).

—Genipa americana L.

Genipa curviflora Dwyer, Ann. Missouri Bot.

Gard. 64: 184. 1980. Type: PANAMA:

viTRAL American Rubiaceae

A Nomenclator of Mexican and Central American Rubiaceae

PTBG); Isotypes F, n.v., NY, n.v., US 943469

(photos MEXU, PTBG)). Duggena.

Gonzalagunia brenesii Standi., Publ. Field

Mus. Nat. Hist., Bot. Ser. 18: 1302. 1938.

Type: COSTA RICA: Near San Ramon,

Bosque a orillas del mar de Golfito de Osa, 12

Apr 1930, A. Brenes 12323 (Holotype F

856730 (photo PTBG); Isotypes GH, n.v., US

(photos MEXU, PTBG)).

Gonzalagunia chiapasensis (Standi.) Standi. &

Steyerm., Publ. Field Mus. Nat. Hist., Bot.

Ser. 22: 283. 1940. Rondeletia chiapasensis

Standi., Publ. Field Mus. Nat. Hist., Bot. Ser.

22: 54. 1940. Type: MEXICO: Chiapas,

Cascarada, 31 Dec 1936, E. Matuda 1064

(Holotype MICH, n.v.; Isotypes F (fragment),

TEX-LL (photo PTBG)). Rogiera.

Gonzalagunia exaltata Standi., Publ. Field Mus.

Nat. Hist., Bot. Ser. 22: 385. 1940. Type:

COSTA RICA: San Jose, basin of El General,

beside river, 675-900 m. Mar 1940, A. F.

Skutch 4747 (Holotype F 1044350 (photo

PTBG); Isotypes A, n.v., MO (photos MEXU,

PTBG), US (photos MEXU, PTBG)).

—Rondeletia brenesii Standi.

Gonzalagunia glabra S. Watson, Proc. Amer.

Acad. Arts 25: 152. 1890. Type: MEXICO:

Jalisco, in the mountains near Lake Chapala,

Dec 1889, C. G. Pringle 2442 (Holotype A,

n.v.; Isotype US (photo PTBG)).

—Citharexylum glabrum (S. Watson)

Greenm. (Verbenaceae)

Gonzalagunia kallunkiae Dwyer, Ann.

Missouri Bot. Gard. 67: 192. 1980 [as

kallunkii]. Type: PANAMA: Veraguas, NW

of Santa Fe, 11 km Escuela Agricola Alto de

Piedra, on road to Calovebora, near Rio

Coloveborita, 15 Nov 1975,5. Mori & J.

Kallunki 3141 (Holotype MO 2285895 (photo

PTBG)).

Gonzalagunia longithyrsa Fosberg, Sida 2: 387.

1966. Type: COSTA RICA: Cartago, Tapanti,

at edge of wet forest, 1200 m, 1 Nov 1964,

W. H. Hatheway & R. L Rodriguez 1241

(Holotype US 2562636 (photos MEXU,

PTBG)).—Gonzalagunia rosea Standi.

Gonzalagunia nivea (Bartl. ex DC.) Kuntze,

Rev. Gen. PI. 1: 284. 1891. Gonzalea nivea

Bartl. ex DC., Prodr. 4: 436. 1830. Type:

MEXICO: Without precise locality, Haenke

s.n. (Holotype presumably PR, n.v., fragment

G-DC, n.v. (microfiche 4: 436.3). Duggena.

—Rondeletia buddleioides Benth.

Gonzalagunia ovatifolia (Donn. Sm.) B. L.

Rob., Proc. Amer. Acad. Arts 45: 405. 1910.

Gonzalea ovatifolia Donn. Sm., Bot. Gaz.

(Crawfordsville) 27: 336. 1899. Type:

COSTA RICA: Cartago, Atirro, 600 m, J. D.

Smith 6599 (Lectotype US 943467, designated

here under main entry (photos MEXU,

PTBG); Isolectotype F). Syntypes: Costa

Rica, J. D. Smith 6600 (F, n.v., GH, US);

Costa Rica, Pittier & Tonduz 9171 (US, n.v.);

Panama, Hart 113 (US, n.v.). Duggena.

Gonzalagunia panamensis (Cav.) K. Schum. in

Mart., FI. Bras. 6(6): 292. 1889. Buena

panamensis Cav., An. Hist. Nat. 2: 279. 1800.

Type: PANAMA: Lancon, Nee s.n. (Holotype

presumably MA, n.v.; possible Isotype F

(photo PTBG)).

Gonzalagunia parviflora Cham. & Schltdl.,

Linnaea 6: 414. 1831. Type: MEXICO:

Veracruz, Cuesta Grande de Jalacingo,

Schiede & Deppe 1275 (Holotype presumably

HAL, n.v.; Isotype B, presumably destroyed

(photo MO)).—Rondeletia septicidalis B. L.

Rob., vel sp. aff

Gonzalagunia rojasii Standi., Publ. Field Mus.

Nat. Hist., Bot. Ser. 22: 114. 1940. Type:

GUATEMALA: Quezaltenango, El Muro,

below Santa Maria de Jesiis, ca. 1200 m, 1

Mar 1938, P. C. Standley 67145 (Holotype F

995085 (photo PTBG)).

Gonzalagunia rosea Standi., Ann. Missouri

Bot. Gard. 25: 836. 1938. Type: PANAMA:

Chiriqui, valley of the upper Rio Chiriqui

Viejo, 1300-1900 m, Jul-Aug 1937, G. IVhite

& P. White 7 (Holotype F 913064 (photo

PTBG); Isotypes A, n.v., MO (photo PTBG),

NY, n.v.).

Gonzalagunia rudis (Standi.) Standi., J. Wash.

Acad. Sci. 17: 170. 1927, comb, inval.

Duggena rudis Standi., Contr. U.S. Natl.

Herb. 18: 125. 1916. Type: PANAMA:

Panama, shady places along Chagras at El

A Nomenclator of Mexican and Central American Rubiaceae

(photos MEXU, PTBG); Isotype GH, n.v.).

Duggena. —Gonzalagunia thyrsoidea (Donn.

Sm.) B. L. Rob.

Guettarda L., Sp. PI. 2: 991. 1753. Type:

Guettarda speciosa L.

Guettarda armata Bartl. ex DC., Prodr. 4: 457.

1830. Type: MEXICO: Without precise

locality, Haenke s.n. (Holotype presumably

PR, n.v. (fragment G-DC, n.v. (microfiche 4:

457.19)).—Status uncertain, probably .

Chomelia spinosa Jacq.

Guettarda brenesii Standi., Publ. Field Mus.

Nat. Hist., Bot. Ser. 18: 1303. 1938. Type:

COSTA RICA: Golfo de Nicoya, Isla de

Caballo, Jun 1932, A. Brenes 15694

(Holotype F 857985 (photo PTBG)).

Guettarda chiriquiensis Standi., Ann. Missouri

Bot. Gard., 25: 838. 1938. Type: PANAMA:

Chiriqui, Valley of upper Rio Chiriqui Viejo,

1300-1900 m, 27 Jul 1937, P. White & G.

White 22 (Holotype MO 1177311 (photo

PTBG); Isotypes A, n.v., F).—Guettarda

crispiflora Vahl

Guettarda cobanensis Donn. Sm., Bot. Gaz.

(Crawfordsville) 47. 255. 1909. Type:

GUATEMALA: Alta Verapaz, near Coban,

1500 m, H. von Tuerckheim 11.2096 (Holotype

US 941415 (photo PTBG); Isotype US (photo

PTBG)).

Guettarda colubrinoides Standi., Publ. Field

Columbian Mus., Bot. Ser. 8: 358. 1931.

Type: MEXICO: Tabasco, Paso del Carrizal, 4

May 1890, J. N. Rovirosa 787 (Holotype K,

n.v., fragment F)).—Status uncertain,

probably Guettarda elliptica Sw.

Guettarda combsii Urb., Symb. Antill. 6: 48.

1909. Type: CUBA: Distr. Cienfuegos, Prov.

Santa Clara, prope Cieneguito, 2 Jul 1895, R.

Combs 270 (Syntype B, destroyed; Lectotype

F 358073 (photo PTBG), here designated;

Isolectotypes F, US). Syntype: Prov. Pinar del

Rio inter Candelaria et Artemisa, Sept.,

Wilson 1783 (B, destroyed).

Guettarda conferta Benth., Bot. Voy. Sulphur

106. 1845. Type: COSTA RICA: Cocos

Island, Barclay s.n. (Holotype BM, n.v.).

Guettarda crispiflora Vahl, Eclog. Amer. 1:

36, t. 6. 1796. Type: MONTSERRAT: Versus

surrunitates in insula Montserrat, Ryan s.n.

(Holotype presumably C, n.v.).

Guettarda dealbata M. Martens «fe Galeotti,

Bull. Acad. Roy. Soc. Bruxelles 11: 232.

1844. Type: MEXICO: Oaxaca, dans les

forets epaisses et humides de Llano Verde et

de Tanetze, dans le Rincon (cordillere

orientale d’Oaxaca), 4000-6000 ft., H.

Galeotti 7131 (Holotype presumably BR,

n.v.).—Status uncertain, “scarcely of this

genus” according to Standley (1934).

Guettarda deamii Standi., Contr. U.S. Natl.

Herb. 20: 209. 1919. Type: GUATEMALA:

Zacapa, Gualan, 620 ft., 15 Jun 1909, C. C.

Deam 6271 (Holotype US 796136 (photo

PTBG); Isotype GH, n.v.).

Guettarda dichotoma M. Martens & Galeotti,

Bull. Acad. Roy. Sci. Bruxelles 11: 233. 1844.

Type: MEXICO: Veracruz, savanes de

Consoquitla, pres la colonie de Zacuapan et

Mirador, 1500-2000 pieds, 1840,//. Galeotti

2601 (Holotype presumably BR, n.v.; Isotype

P (photos MEXU, PTBG)).—Guettarda

elliptica Sw.

Guettarda elliptica Sw., Prodr. Veg. Ind. Occ.

59. 1788. Type: JAMAICA: Campestris

siccis, Swartz s.n. (presumably S, n.v.).

Guettarda filipes Standi., Contr. U.S. Natl.

Herb. 20: 210. 1919. Type: MEXICO:

Durango, near Huasemote, 15 Aug 1897,/ N.

Rose 3498 (Holotype US 302475).

Guettarda foliacea Standi., Contr. U.S. Natl.

Herb. 18: 139. 1916. Type: PANAMA: Canal

Zone, along the Trinidad River, near sea level,

19-21 Jul 1911, //. Pittier 4031 (Holotype US

679116; Isotype F).

Guettarda galeotti Standi., Contr. U.S. Natl.

Herb. 23: 1384. 1926. Type: MEXICO:

Oaxaca, Prov. Pinotepa, H. Galeotti 2576

(Holotype US 572992, fragment F (photos

MEXU, PTBG)).—Chomelia protracta

(Bartl. ex DC.) Standi.

Guettarda gaumeri Standi., Publ. Field

Columbian Mus., Bot. Ser. 8: 58. 1930. Type:

MEXICO: Yucatan, without precise locality,

1917-1921, G. F. Gaumer 24239 (Holotype F

A Nomenclator of Mexican and Central American Rubiaceae

Guettarda tikalana Lundell, Wrightia 2: 63.

1960. Type: GUATEMALA: Peten, Tikal, on

Uaxactun trail, in center of escobal, 18 Jul

1959, C L. Lundell 16519 (Holotype TEX-LL

(photos MEXU, PTBG); Isotypes F, US).

Guettarda tournefortiopsis Standi., Publ. Field

Columbian Mus., Bot. Ser. 7: 293. 1931.

Based on Tournefortiopsis reticulata Rusby,

Bull. New York Bot. Gard. 4: 369. 1907. Not

Guettarda reticulata Griseb., 1863. Type:

BOLIVIA: Without precise locality. Bang

2230 (Holotype US, n.v.).

Guettarda turrialbana N. Zamora & Poveda,

Ann. Missouri Bot. Gard. 75: 1157, fig. 1.

1988. Type: COSTA RICA: Cartago,

Turrialba, CATIE, Bosque de Los Espabeles,

600 m, 26 Jun 1986, N. Zamora, M. M.

Chavarria & H. Gomez 1263 (Holotype CR,

n.v.; Isotypes F (photo PTBG), K, n.v., MO

(photo PTBG), NY, n.v.).

Habroneuron Standi., J. Wash. Acad. Sci. 17:

338. 1927. Type: Habroneuron mexicanum

Standi.

Habroneuron mexicanum Standi., J. Wash.

Acad. Sci. 17: 339. 1927. Type: MEXICO:

Oaxaca, Tintalcingo, F. Liebmann 11527

(Holotype C, n.v.; Isotype C, n.v., fragment

US (photos MEXU, PTBG)).—Habroneuron

radicans (Wemham) S. P. Darwin

Habroneuron radicans (Wemham) S. P.

Darwin, Brittonia 32: 343. 1980. Lindenia

radicans Wemham, J. Bot. 52: 226. 1914.

Type: MEXICO: Oaxaca, shaded places on

rocks at crossing of the river near

Tocotepeque, Jul 1839, T. Hartwegs.n.

(Holotype K, n.v. (photo MO)).

Hamelia Jacq., Enum. Syst. PI. 2: 16. 1760.

Type: Hamelia erecta Jacq. (Lectotype

designated by Britton & Millsp., Bahama FI.

441. 1920).

Hamelia axillaris Sw., Prodr. Veg. Ind. Occ.

46. 1788. Type: JAMAICA: Without precise

locality, Swartz s.n. (Lectotype S, n.v.;

designated by Elias, 1976).

Hamelia barbata Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 22: 115. 1940. Type:

GUATEMALA: Escuintla, Finca Monterrey,

S slope of Volcan de Fuego, 1140-1260 m, 5

Feb 1939, P. C. Standley 64569 (Holotype F

982608 (photo PTBG); Isotypes A, n.v., NY,

n.v., US (photos MEXU, PTBG)).

Hamelia calycosa Donn. Sm., Bot. Gaz.

(Crawfordsville) 12: 132. 1887. Type:

GUATEMALA: Alta Verapaz, Pansamala,

3600 ft., Jun 1885, H. von Tuerckheim 454

(Holotype US 943617 (photos MEXU,

PTBG); Isotypes GH, n.v., K, n.v., NY, n.v.,

US (photos MEXU, PTBG)).

Hamelia chiapensis Brandegee, Univ. Calif

Publ. Bot. 6: 71. 1914. Type: MEXICO:

Chiapas, Finca Mexiquito, Jul 1913, C. A.

Purpus 7030 (Holotype UC 173058, n.v.;

Isotypes F (photos MEXU, PTBG), GH, n.v.,

MO (photos MEXU, PTBG), NY (n.v.), US

(photos MEXU, PTBG)).—Hamelia calycosa

Donn. Sm.

Hamelia costaricensis Standi., Contr. U.S. Natl.

Herb. 20: 207. 1919. Type: COSTA RICA:

Sumlres [?] pres San Mateo, Aug 1890, P.

Biolley 2656 (Holotype US 764417 (photos

MEXU, PTBG)).—Hamelia xerocarpa

Kuntze var. costaricensis (Standi.) Elias

Hamelia corymbosa Sesse & M09., La

Naturaleza, ser. 2, 1: 37. 1888 (as

“Hamellia”). Type: MEXICO: Habitat in

montibus Haereditatis-Ahuesuyo, Sesse &

Mogiho s.n. (presumably MA, n.v.).

—Hamelia patens Jacq. var. patens

Hamelia declinata Sesse & M09., FI. Mexic.,

ed. 2, 60. 1894. Type: MEXICO: Sesse &

Mogiho s.n. (presumably MA, n.v.).

—Hamelia axillaris Sw.

Hamelia erecta Jacq., Enum. Syst. PI. Carib. 16.

1760. Type: COLOMBIA: Bolivar, vicinity of

Cartagena, Jacquin s.n. (location unknown).

—Hamelia patens Jacq. var. patens

Hamelia hypomalaca B. L. Rob., Proc. Amer.

Acad. Arts 45: 406. 1910. Type: MEXICO:

Durango, without precise locality, Aug 1897,

J. N. Rose 2304 (Holotype US 301215 (photos

MEXU, PTBG); Isotypes GH, n.v., MO

(photos MEXU, PTBG)).—Hamelia

xorullensis Kunth in Humb., Bonpl. & Kunth

Lorence

Hamelia lanuginosa M. Martens & Galeotti,

Bull. Acad. Roy. Sci. Bruxelles 11; 233. 1844.

Type: MEXICO: Veracruz, Mirador, Xalapa,

1841, //. Galeotti 2615 (Holotype BR, n.v.;

Isotypes MO (photos MEXU, PTBG), P

(photos MEXU, PTBG)).—Hamelia patens

Jacq. var. patens

Hamelia longipes Standi., Proc. Biol. Soc.

Wash. 37: 53. 1924. Type: GUATAMALA:

Izabal, vicinity of Escoba, on the bay opposite

Puerto Barrios, 150 m or less, 2 Jun 1922,

P. C. Standley 24829 (Holotype US 1150447

(photos MEXU, PTBG); Isotype GH, n.v.).

Hamelia lutea Rohr ex Rees, Cyclop. 17. 1811.

Type: Based on manuscript by von Rohr and

Jaquin’s (Collectanea, Ic. Rar. tab 335)

description and illustration of H. chrysantha

—Hamelia axillaris Sw.

Hamelia macrantha Little, Carib. Forester 9:

274. 1948. Type: ECUADOR: Chimborazo,

El Oro, Spruce 6193 (Lectotype K, n.v.

(photos F, MO); designated by Elias, 1976).

Hamelia magnifolia Wemham, J. Bot. 49: 210.

1911. Type: COSTA RICA: Puntarenas, bords

du Rio Corozal, pres Santo Domingo de Golfo

Duke, Mar 1896, H. Pittier 10091

(Lecototype BM, n.v.; Isolectotypes F,

fragment n.v., GH, n.v., P (photos MEXU,

PTBG), US (photos MEXU, PTBG);

designated by Elias, 1976).

Hamelia magniloba Wemham, J. Bot. 49: 209.

1911. Type: NICARAGUA: Chontales, Tate

200 (Holotype K, n.v.).—Hamelia xerocarpa

Kuntze var. xerocarpa

Hamelia nodosa M. Martens & Galeotti, Bull.

Acad. Roy. Sci. Bruxelles 11: 234. 1844.

Type: MEXICO: Veracmz, pres Mirador,

3000 pieds, H. Galeotti 2581 (Holotype

presumably BR, n.v.; Isotype P, n.v.).

—Hamelia patens Jacq. var. glabra Oerst.

Hamelia panamensis Standi., Contr. U.S Natl

Herb. 20: 208. 1919. Type: PANAMA: Above

Paca, 16 Apr 1908, R. S. Williams 744

(Holotype US 678258 (photos MEXU,

xerocarpa Kuntze

Elias

r. costaricensis (Standi.)

Hamelia patens Jacq., Enum. Syst. PI. Carib.

16. 1760. var. patens. Type: SANTO

DOMINGO (DOMINICAN REPUBLIC):

Jacquin s.n. (location unknown). Dunhamelia.

Hamelia patens Jacq. var. coronata Dorm. Sm.,

Bot. Gaz. (Crawfordsville) 40: 4. 1905. Type:

GUATEMALA: Alta Verapaz, Cubliquitz,

350 m, Aug 1903, H. von Tuerckheim 8532

(Holotype US 943525 (photos MEXU,

PTBG); Isotypes F, GH, n.v., NY, n.v., US

(photos MEXU, PTBG)).—Hamelia

rovirosae Wemh.

Hamelia patens Jacq. var. glabra Oerst.,

Vidensk. Meddel. Dansk Naturhist. Foren.

Kjobenhavn 1852: 42. 1853. Type:

NICARAGUA: San Juan, Oersted s.n.

(Holotype C, n.v.).

Hamelia pauciflora Standi., Contr. U.S. Natl.

Herb. 18: 139. 1916. Type: PANAMA:

Panama, forests on dry limestone around

Alhajuela, Chagres Valley, 30-100 m, 12-15

May 1911, //. Pittier 3469 (Holotype US

678513 (photos MEXU, PTBG)). Not H.

pauciflora 1825.—Hamelia

sanguinea Elias

Hamelia purpurascens S. F. Blake, Contr. Gray

Herb. II, 52: 105. 1917. Type: BRITISH

HONDURAS (BELIZE): On pine ridge near

Manatee Lagoon, 6 Aug 1904, M. E. Peck 104

(Holotype GH, n.v.).—Hamelia rovirosae

Wemham

Hamelia rostrata Bartl. ex DC., Prodr. 4: 442.

1830. Type: MEXICO: Guerrero, Acapulco,

Haenke s.n. (Holotype presumably PR (n.v.)).

According to Elias (1976: 136), “This may be

the oldest valid name for H. versicolor, but

the description is too brief for accurate

determination.”—Status uncertain, probably

Hamelia versicolor A. Gray

Hamelia rovirosae Wemham, J. Bot. 49: 211.

1911. Type: MEXICO: Tabasco, marshes on

banks of Rio Sebastian, 11 Jun 1889, J. N.

Rovirosa 499 (Holotype K, n.v.; Isotypes NY,

n.v., US (photos MEXU, PTBG)).

Hamelia rowleei Standi., J. Wash. Acad. Sci. 15:

7. 1925. Type: COSTA RICA: Livingston on

the Reventazon River, Jul-Aug 1920, W. W.

Rowlee & H. E. Stork 751 (Holotype US

A Nomenclator of Mexican and Central American Rubiaceae

Hedyotis sharpii (Terrell) G. L. Nesom, Syst.

Hot. 13: 434. 1988. Houstonia sharpii Terrell,

Brittonia 32: A90,fig. I. 1980. Type:

MEXICO: Hidalgo, camino entre Metepec y

Tenango de Doria, entre 16 y 31.5 km al NNE

de Tulancingo, 18 km NNE de Metepec, 29

Mar 1974, S. D. Koch 7423 (Holotype US

2754863 (photos MEXU, PTBG); Isotypes

CHAPA, ENCB, MEXU).

Hedyotis sinaloeae W. H. Lewis, Rhodora 63:

222. 1961, nom. nov. for Houstonia parvula

Brandegee, Zoe 5: 221. 1905. Not Hedyotis

parvula (A. Gray) Fosb., 1943. Type:

MEXICO: Sinaloa, vicinity of Culiacan, 12

Oct 1904, T. S. Brandegee s.n. (Holotype UC,

n.v.; Isotypes GH, n.v., US (photos MEXU,

PTBG)).

Hedyotis spellenbergii G. L. Nesom &

Vorobik, Syst. Bot. 13: 432. 1988. Type:

MEXICO: Chihuahua, Mun. Ocampo, area of

Cascada de Basaseachic at the confluence of

Rio Basaseachic and Rio Durazno, ca. 2 mi S

of village of Basaseachic, 2100-2200 m, 18

May 1985, y. Grimes, T. Ayers, M. Lavan &

R. Scott 2855 (Holotype TEX, n.v.; Isotypes

ARIZ, n.v., CAS, n.v., MEXU, n.v., NMC,

n.v., NY, n.v., US, n.v.).

Hedyotis spinescens Sesse & M09., FI. Mexic.

22. 1893. Type: MEXICO: Habitat in aridis

Tehuacani circuitibus. Floret Maio, Sesse &

Mogiho s.n. (presumably MA, n.v.).

—Bouvardia erecta (DC.) Standi.

Hedyotis teretifolia (Terrell) G. L. Nesom,

Syst. Bot. 13: 434. 1988. Houstonia teretifolia

Terrell, Brittonia 31: 166. 1979. Type:

MEXICO: Coahuila, 12 km NNE of Las

Margaritas on easternmost ridge of Sierra de

las Margaritas, 1300-1400 m, 24 Sep 1972,

M C. Johnston, F. Chiang & T. Wendt 9508E

(Holotype TEX, n.v.).

Hedyotis terrellii Lorence, Novon 7: 51, fig. 3.

1997. Type: MEXICO: Oaxaca, Distr. de

Mixe, Mun. de Totontepec, 3 km al SW de

Totontepec, carretera a Mitla, 1910 m, 6 Sep

1986, R. Torres C. & C. Martinez 9067

(Holotype PTBG 12314; Isotype MEXU).

Hedyotis umbratilis (B. L. Rob.) W. H. Lewis,

Rhodora 63: 222. 1961. Houstonia umbratilis

B. L. Rob., Proc. Amer. Acad. Arts 45: 401.

1910. Type: MEXICO: Nuevo Leon, shaded

cliffs and mountains, near Monterrey, 25 Apr

1906, C. G. Pringle 13877 (Holotype GH,

n.v.; Isotypes MO (photos MEXU, PTBG),

US (photos MEXU, PTBG)).

Hedyotis vergrandis W. H. Lewis, Rhodora 63:

222. 1961. Nom. nov. for Houstonia prostrata

Brandegee, Zoe 5: 105. 1901. Not Hedyotis

prostrata Blume, 1823, norKorth., 1851.

Type: MEXICO: Baja California, La Palma,

Cape Region, 25 sep 1899, T. S. Brandegee

s.n. (Holotype UC, n.v.; Isotypes GH, n.v.,

US, 2 sheets (photos MEXU, PTBG)).

Hedyotis watsonii W. H. Lewis, Rhodora 63:

222. 1961. Nom. nov. for Oldenlandia ovata

S. Watson, Proc. Amer. Acad. 18: 97. 1883.

Not Hedyotis ovata Thunb. ex Maxim., 1833,

nor Thwaites ex Hook, f, 1892. Type:

MEXICO: Nuevo Leon, Guajuco, Feb-Oct

1880, E. Palmer 399 (Lecototype GH, n.v.;

Isolectotype US 47919 (photos MEXU,

PTBG)).

Hedyotis wrightii (A. Gray) Fosberg, Lloydia

4: 290. 1941. Houstonia wrightii A. Gray,

Proc. Amer. Acad. Arts 17: 202. 1882. Type:

MEXICO: Nuevo Leon, shaded cliffs of

mountains, near Monterrey, 25 Apr 1906,

C. G. Pringle 13877 (Holotype GH, n.v.;

Isotypes MO (photos MEXU, PTBG), US

(photos MEXU, PTBG)).

Hedyotis xestosperma (B. L. Rob. & Greenman)

W. H. Lewis, Rhodora 63: 222. 1961.

Oldenlandia xestosperma B. L. Rob. &

Greenman, Proc. Amer. Acad. Arts 32: 41.

1896. Type: MEXICO: Oaxaca, open glades.

Sierra de San Felipe, 8000 ft., Jun 1894, C. G.

Pringle 4692 (Lectotype GH, n.v.;

Isolectotypes BM, n.v., F, K, n.v., MO

(photos MEXU, PTBG), MSC, n.v., PH, n.v.,

US, 2 sheets (photos MEXU, PTBG), VT

n.v.); designated by Terrell, 1980). Houstonia.

—Bouvardia xestosperma (B. L. Rob. &

Greenman) Terrell «fe S. D. Koch

Hemidiodia K. Schum. in Mart., FI. Bras. 6(6):

29. 1888. Type: Hemidioidia ocymifolia

. Nomenclator of Mexican and Central American Rubiaceae

Langlasse 234 (K, n.v.); Nayarit, Tepic,

Lumholtz s.n. (US); Puebla, Andrieux 337 (K,

n.v.), Pringle 8583 (K, n.v.), Seler 849 (US),

Rose & Hay 5932 (US), Rose. Painter & Rose

10123 (US); Oaxaca, Purpus 3279 (US), Reko

4253 (US), Pringle 7473 (US), Pringle &

Conzatti 27 (US). C. V. Morton annotated the

Nelson 6985 specimen at US as lectotype, but

this choice was not published.—Hintonia

latiflora (Sesse & Mo?, ex DC.) Bullock

Hippotis Ruiz & Pavon, FI. Peruv. Chil. 33

1794. Type: Hippotis trifolia Ruiz & Pavon

Hippotis albiflora H. Karst., FI. Columb. 1: 33.

1860. Type: COLOMBIA: Mountains E of

Bogota, Karsten s.n. (presumably W, n.v.)

Hoffmannia Sw., Prodr. 30. 1788. Type:

Hoffmannia pedunculata Sw.

Hoffmannia aeruginosa Standi., Publ. Field

Mus. Nat. Hist, Bot Ser. 18: 1313. 1938.

Type: COSTA RICA: Zarcero, 1500 m, 10

Aug 1937, Austin Smith A86 (Holotype F

941349 (photo PTBG)).

Hoffmannia affinis Hemsl., Diag. PI. Nov.

Mexic. 31. 1879. Type: COSTA RICA:

Without precise locality, Endres 150

(Holotype K, n.v.; Isotype W (photo PTBG)).

—Hoffmannia nicotianifolia (M. Martens &

Galeotti) L. O. Williams

Hoffmannia amplexifolia Standi., J. Wash.

Acad. Sci. 15: 8. 1925. Type: COSTA RICA:

Cartago, vicinity of Orosi, moist forest, 30

Mar 1924, P. C. Standley 39869 (Holotype

US 1153192 (photos MEXU, PTBG)).

Hoffmannia angustifolia Standi., Contr. U.S.

Natl. Herb. 20: 206. 1919. Type:

GUATEMALA: Zacatepequez, Acatepeque,

1290 m. Mar 1892, J. D. Smith 2747

(Holotype US 888492 (photos MEXU,

PTBG); Isotype GH, n.v.).—Hoffmannia

nicotianifolia (M. Martens & Galeotti) L. O.

Williams

Hoffmannia araneopedaria Dwyer, Ann.

Missouri Bot Gard. 67: 226. 1980. Type:

PANAMA; Darien, Cerro Mali on Colombian

border, S slope just below surmnit, 1400 m, 6

Feb 1975, A. Gentry & S. Mori 14166

(Holotype MO 2470655; Isotype MO

2470654).

Hoffmannia arborescens Donn. Sm., Bot Gaz.

(Crawfordsville) 37: 417. 1904. Type:

COSTA RICA: Forets de Santa Rosa del

Copey, 1800 m, Apr 1898, A. Tonduz 8121

[Herb. Inst. Fysico-geogr. nat. costaricensis

12230] (Holotype US 943523 (photos MEXU,

PTBG)).

Hoffmannia areolata Standi., Publ. Field Mus.

Nat Hist, Bot Ser. 22: 52. 1940. Type:

PANAMA; Chiriqui, Boquete District, near

Bajo Chorro, Jan 1938, M. E. Davidson 40

(Holotype F 915621 (photo PTBG); Isotypes

GH, n.v., MO).

Hoffmannia asclepiadea Standi., J. Wash.

Acad. Sci. 15: 7. 1925. Type: COSTA RICA:

San Jose, Las Nubes, 1500-1900 m, 20-25

Mar 1924, P. C. Standley 38349 (Holotype

US 1153122 (photos MEXU, PTBG)).

Hoffmannia bullata L. O. Williams, Fieldiana,

Bot 36: 52. 1973. Type: GUATEMALA:

Izabal, between Virginia and Lago Izabal,

Montana del Mico, 50-100 m, 4 Apr 1940,

J. A. Steyermark 38719 (Holotype F).

—Hoffmannia discolor (Lem.) Hemsl.

Hoffmannia calycosa Donn. Sm., Bot. Gaz.

(Crawfordsville) 40: 4. 1905. Type:

GUATEMALA: Alta Verapaz, Cubliquitz,

350 m.. May 1901 //. von Tuerckheim 7912

(Holotype US 576489 (photos MEXU,

PTBG); Isotypes F (photos MEXU, PTBG),

US, 2 sheets (photos MEXU, PTBG)).

Hoffmannia capillacea Dwyer, Ann. Missouri

Bot. Gard. 56: 274. 1969. Type: PANAMA:

Code, mountains N of El Valle, 2500-3000

ft., 28 May 1967, Lewis, MacBryde, Oliver &

Ridgway 1783 (Holotype MO; Isotypes MO

2214603, 2163471). Note: Between the time

Hoffmannia capillacea Dwyer was published

and the Rubiaceae treatment for the Flora of

Panama was published (Dwyer, 1980), the

wrong labels {T. Elias 1783) were

inadvertently mounted on the type collection.

Dwyer’s protologue lists the type as being

Lewis et al. 1783. The three collections in the

type folder at MO correspond with the

description of H. capillacea but bear Elias

LORENCE

/ 783 labels. A reexamination of the Lewis

and Elias field books by C. M. Taylor and me

in June 1996 revealed that Elias 1783 was

collected in San Bias, Panama, reputedly as a

species of Annonaceae: “headquarters of Rio

Mulatupo, tree 5 m, fruits green ripening

yellow.” The Lewis et al. 1783 collection was

said to be “Rubiaceae, shrub to 2.2 m, calyx

tube red, flowers white, fruit red.” Therefore,

the Lewis et al. 1783 label seems to be the

correct one.

Hojfmannia carlsoniae Standi. & L. O. Williams

ex A. R. Molina, Ceiba 14: 3. 1968. Type:

MEXICO: Chiapas, Los Lagos, 3 mi. NW of

Rancho San Jose, which is 34 mi. SE of

Comitan, 5,000 ft., 15-20 Apr 1949, M. C.

Carlson 1874 (Holotype presumably EAP,

n.v.; Isotype F 1412975).—Hoffmannia

huehueteca Standi. & Steyerm.

Hoffmannia carpinterae Standi., N. Amer. FI.

32: 199. 1934. Based on

macrophylla Standi. 1925, not H. macrophylla

Hemsl., 1881. Type: COSTA RICA: Cartago,

Cerro de la Carpintera, 1700 m, P. C. Standley

35636 (Holotype US 1152958 (photos

MEXU, PTBG); Isotype US 1152957 (photos

MEXU, PTBG)).—Hoffmannia leucocarpa

Standi.

Hoffmannia cauliflora Hemsl., Diag. PI.

Mexic. 30. 1879. Type: GUATEMALA:

Without precise locality, Skinner s.n.

(Holotype K, n.v.).

Hoffmannia cercidifolia Dwyer, Ann. Missouri

Bot. Gard. 56: 275. 1969. Type: PANAMA:

Darien, Ceao Campamento, S of Cerro Pirre,

20-22 Mar 1968,7. A. Duke 75597 (Holotype

MO 1956874 (photo PTBG)).

Hoffmannia chiapensis Standi., Contr. U.S.

Natl. Herb. 20: 206. 1919. Type: MEXICO:

Chiapas, Cerro del Boqueron, Jun 1914, C. A.

Purpus 7268, in part (Holotype US 567526

(photos MEXU, PTBG); Isotype F (photos

MEXU, PTBG)).—Hoffmannia

psychotriifolia (Benth in Oerst.) Griseb.

Hoffmannia confertiflora Standi., Contr. U.S.

Natl. Herb. 20: 206. 1919. Type:

GUATEMALA: Quiche, San Miguel

Uspantan, 2100 m, 1892, Heyde & Lux 3169

(Holotype US 888491 (photos MEXU,

PTBG); Isotype GH, n.v.).

Hoffmannia congesta (Oerst.) Dwyer in W. C.

Burger & C. M. Taylor, Fieldiana, Bot., n.s.

33: 171. 1993. Xerococcus congestus Oerst.,

Vidensk. Meddel. Dansk. Naturhist. Foren

Kjobenhavn 1852: 52. 1853. Type: COSTA

RICA: Turrialba, 3000 ft.. Oersted 11776

(Holotype C, n.v. (photos F, MO); Isotype F,

n.v.).

Hoffmannia conzattii B. L. Rob., Proc. Amer.

Acad. Arts 45: 406. 1910. Based on Coffea

obovata Schlecht. & Cham., Lirmaea 6: 412.

1831. Not Hoffmannia obovata (Ruiz &

Pavon) Standi., 1930 (based on Ohigginsia

obovata Ruiz & Pavon 1798). Type:

MEXICO: Veracruz, Canton de Cordoba,

Colonia Melchor Ocampo, 1200 m, 19 Jun

1896, C. Conzatti 168 (Holotype GH, n.v.).

Hoffmannia cryptoneura Standi., Publ. Field

Mus. Nat. Hist., Bot. Ser. 22: 53. 1940. Type:

MEXICO: Chiapas, Mt. Ovando, 4 Apr 1936,

E. Matuda 944 (Holotype F 898899 (photo

PTBG); Isotypes MICH, n.v., MO (photos

MEXU, PTBG), TEX-LL (photos MEXU,

PTBG)).

Hoffmannia culminicola Standi. & L. O.

Williams, Ceiba 1: 166. 1950. Type:

HONDURAS: Morazan, Cerro de Uyuca, ca.

1600 m, 2 Mar 1947, P. C. Standley 4797

(Holotype F 1256864 (photos MEXU,

PTBG); Isotype EAP, n.v.).

Hoffmannia cuneatissima B. L. Rob., Proc.

Amer. Acad. Arts 45: 407. 1910. Type:

MEXICO: Morelos, mountain canyon near

Cuernavaca, 6000 ft., 29 May 1898, C. G.

Pringle 7662 (Holotype GH, n.v.; Isotype US

334830 (photos MEXU, PTBG)).

Hoffmannia davidsoniae Standi., Publ. Field

Mus. Nat. Hist., Bot. Ser. 22: 53. 1940. Type:

PANAMA: Chiriqui, Boquete District, Bajo

Chorro, 6000 ft., 26 Jan 1938, M. E. Davidson

216 (Holotype F 915570 (photo PTBG);

Isotypes A, n.v., MO, US (photos MEXU,

PTBG)).

Hoffmannia decurrens Standi., Contr. U.S.

Natl. Herb. 20: 205. 1919. Type: COSTA

RICA: Forets de Santa Rosa del Copey,

A Nomenclator of Mexican and Central American Rubiaceae

1800-2000 m, Apr 1898, A. Tonduz 12230

(Holotype US 392077 (photos MEXU,

PTBG); Isotypes P, 2 sheets (photos MEXU,

PTBG)).

Hoffmannia discolor (Lem.) Hemsl., Biol.

Centr.-Amer., Bot. 2: 36. 1881.

Misc. 31. fig. 1847. Type: Not designated,

based on cultivated plants grown from seed

collected in Mexico by H. Galeotti. Note: A

specimen at P [“en herb. Houtte Juin 1857.

Hort. Parisei”] and a specimen at G [“Mexici,

Oaxaca, 3000', fl. albis, Galeotti 7183"] may

correspond with Galeotti’s seed collection and

represent authentic cultivated material.

Higginsia.

Hoffmannia dotae Standi., J. Wash. Acad. Sci.

18: 181. 1928. Type: COSTA RICA: San

Jose, vicinity of Santa Maria de Dota,

1500-1800 m, 26 Dec 1925-3 Jan 1926, P. C.

Standley & J. Valerio 43277 (Holotype US

1253172 (photos MEXU, PTBG)).

Hoffmannia eliasii Dwyer, Ann. Missouri Bot.

Gard. 56: 276. 1969. Type: PANAMA:

Darien, Rio Cuasi mainstream, 2.5 miles S of

Tres Bocas, 28 Apr 1968, J. H. Kirkbride, Jr.

& J. A. Duke 1129 (Holotype MO 1982892

(photos MEXU, PTBG)).—Hoffmannia

Hoffmannia excelsa (Kunth) K. Schum. in

Mart., Fl. Bras. 6(6): 327. 1889. Psychotria

excelsa Kunth in Humb., Bonpl. & Kunth,

Nov. Gen. Sp. 3: 355. 1819. Type: MEXICO:

Veracmz, Xalapa, A. Bonpland 1445

(Holotype presumably P-Bonpland, n.v.;

Isotypes B, n.v. (photo MO), P (photos

MEXU, PTBG)). Deppea.

Hoffmannia fimbrianthera Dwyer, Arm.

Missouri Bot. Gard. 56: 277. 1969. Type:

PANAMA: Rio Teribe, just below Puerto

Palenque, 350 m, 12 Apr 1968, J. H.

Kirkbride, Jr. & J. A. Duke 556 (Holotype

MO 1999724).—Hoffmannia inamoena

Standi.

Hoffmannia gentryi Dwyer, Ann. Missouri

Bot. Gard. 67: 230. 1980. Type: PANAMA:

Darien, Cerro Tacarcuna, summit camp, along

stream N of camp, 1550-1650 m, 1 Feb 1975,

A. Gentry & S. Mori 14061 (Holotype MO

2353753; Isotype MO 2423663).

Hoffmannia gesnerioides (Oerst.) Kuntze, Rev.

Gen. PI. 285. 1891. Ophryococcus

gesnerioides Oerst., Vidensk. Meddel. Dansk

Naturhist. Foren. Kjoebenhavn 1852: 53.

1853. Type: NICARAGUA: Segovia, Monte

Pantasmo, Jan 1848, Oersted 11823

(Holotype C, n.v. (photos F, MO); Isotype F).

Hoffmannia ghiesbreghtii (Lem.) Hemsl., Biol.

Centr.-Amer., Bot. 2: 36. 1881.

Campylobotrys ghiesbreghtii Lem., Illust.

Hort. 8: pi. 279. 1861. Type: Not designated,

based on cultivated material from seed

collected by Ghiesbreght, probably in

Chiapas, Mexico. Higginsia.

Hoffmannia hamelioides Standi., J. Wash.

Acad. Sci. 15: 8. 1925. Type: COSTA RICA:

San Jose, between Aserri and Tarbaca,

1600-1800 m, 12 Feb 1924, P. C. Standley

34149 (Holotype US 1152904 (photos

MEXU, PTBG)).

Hoffmannia hammelii C. M. Taylor in W. C.

Burger & C. M. Taylor, Fieldiana, Bot., n.s.

33: 174. 1993. Type: COSTA RICA:

Puntarenas, near Sirena, Corcovado National

Park, 0-200 m, 5 Jul 1977, R. L. Liesner 2907

(Holotype CR, n.v.; Isotype MO).

Hoffmannia haydenii Dwyer, Ann. Missouri

Bot. Gard. 56: 277. 1969. Type: PANAMA:

Chiriqui, Cerro Campana, Jul 1965, s. M. V.

Hayden 7/7 (Holotype MO 2030831). Note:

In the protologue the type locality is

incorrectly cited as “Finca Collins, ca. 5000

ft.” —Hoffmannia subauriculata Standi.

Hoffmannia hondurensis Standi., Publ. Field

Columbian Mus., Bot. Ser. 4: 329. 1929.

Type: HONDURAS: Atlantida, Lancetilla

Valley, 200-500 m, 6 Dec 1927-20 Mar

1928, P. C. Standley 52905 (Holotype F

582885 (photo PTBG); Isotype US (photos

MEXU, PTBG)).

Hoffmannia huehueteca Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 249.

1947. Type: GUATEMALA: Huehuetenango,

Sierra de los Cuchumatanes, Cerro Negro, 2

miles east of Las Palmas, 1600-2000 m, 31

Lorence

Aug 1942, y. A. Steyermark 51697 (Holotype

F 1132825 (photo PTBG)).

Hoffmannia inamoena Standi., J. Wash. Acad.

Sci. 18: 179. 1928. Type: COSTA RICA:

Guanacaste, Los Ayotes, near Tilaran, 600 m,

21 Jan 1926, P. C. Standley & J. Valerio

45421 (Holotype US 1254102 (photos

MEXU, PTBG)).

Hoffmannia ixtlanensis Lorence, Novon 4:

\2\Jig. 2a. b. 1994. Type: MEXICO:

Oaxaca, Distr. de Ixtlan, Sierra de Juarez; ruta

175 Tuxtepec a Oaxaca, ca. 5 km al N de

Vista Hermosa, 1260 m, 14 Apr 1982, D. H.

Lorence. R. Torres C. & R. Cedillo T. 4013

(Holotype MEXU; Isotypes MO, PTBG).

Hoffmannia josefina Standi., J. Wash. Acad. Sci.

15: 8. 1925. Type: COSTA RICA: San Jose,

between Aserri and Tarbaca, 1800 m, 12 Feb

1924, P. C. Standley 34143 (Holotype US

1152900 (photos MEXU, PTBG)).

—Hoffmannia arborescens Donn. Sm.

Hoffmannia kirkbridei Dwyer, Ann. Missouri

Bot. Gard. 56: 278. 1969. Type: PANAMA:

Bocas del Toro, Quebrada Huron, ca.

300-400 ft., 11 Apr 1968, J. H. Kirkbride, Jr.

& J. A. Duke 429 (Holotype MO 1968253).

—Hoffmannia vesiculifera Standi.

Hoffmannia iancistigma Dwyer, Ann. Missouri

Bot. Gard. 56: 279. 1969. Type: PANAMA:

Chiriqui, Boquete, Cerro Horqueta,

5000-6000 ft., 8 Aug 1967, J. D. Dwyer &

S. M. V. Hayden 7735 (Holotype MO

2065523).

Hoffmannia laxa Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 22: 116. 1940. Type: COSTA

RICA: Alajuela, Cantdn de San Carlos, Sucre,

1000 m, 30 Mar 1939, A. Smith F1851

(Holotype F 980999 (photo PTBG); Isotype

US (photos MEXU, PTBG)).

Hoffmannia lenticellata Hemsl., Diag. PI. Nov.

Mexic. 30. 1879. Type: MEXICO: Veracruz,

region d’Orizaba, Rio Blanco, 29 Aout 1866,

Bourgeau 2992 (Lectotype P, here designated

(photos MEXU, PTBG); Isolectotype P

(photos MEXU, PTBG)). Syntypes:

MEXICO: Veracruz, Orizaba, Botteri 974

(Syntype P, photos MEXU, PTBG); Jalapa,

Linden 1 (Syntype K, n.v.). The Bourgeau

2992 collection at P was chosen as lectotype,

because it is most representative, with flowers,

fmits, and full-sized leaves.—Hoffmannia

nicotianifolia (M. Martens & Galeotti) L. O.

Williams

Hoffmannia leucocarpa Standi., J. Wash.

Acad. Sci. 15: 9. 1925. Type: COSTA RICA:

San Jose, Las Nubes, 1300-1600 m. Mar

1924, P. C. Standley 38340 (Holotype US

1153121 (photos MEXU, PTBG)).

Hoffmannia lewisiana Dwyer, Ann. Missouri

Bot. Gard. 56: 279. 1969. Type: PANAMA:

Chiriqui, vicinity of Cerro Punta, 6800 ft., 31

May-1 Jun 1967, Ridgway & Solis 2413

(Holotype MO 1966156).

Hoffmannia liesneriana L. O. Williams,

Fieldiana, Bot. 36: 54. 1973. Type: COSTA

RICA: Limon, Guapiles, a 2 millas al norte de

Diamantes, 150 m, 9 Aug 1963, A. Jimenez

1054 M. (Holotype F 1690189 (photo

PTBG)).

Hoffmannia lineolata Donn. Sm., Bot. Gaz.

(Crawfordsville) 40: 5. 1905. Type:

GUATEMALA: Alta Verapaz, Cubliquitz,

350 m. May 1902, H. von Tuerckheim 8227

(Holotype US no 576514 (photos MEXU,

PTBG); Isotypes US, 2 sheets (photos MEXU,

PTBG)).—Hoffmannia regalis (Hook, f.)

Hemsl.

Hoffmannia longicalycina Dwyer, Ann.

Missouri Bot. Gard. 56: 280. 1969. Type:

PANAMA: Darien, Cuasi-Cana Trail between

Cerro Campamiento and La Escalera to

paramo, E of Tres Bocas, 30 Apr 1968, J. H.

Kirkbride. Jr. & J. A. Duke 1278 (Holotype

MO 2165098; Isotype MO 1686749).

—Status uncertain, probably Amphidaysa

sp.

Hoffmannia longipetiolata Pol., Linnaea 41:

567. 1877. Type: COSTA RICA: In sylvis

primaevis Carpinterae, 24 Jun 1875, H.

Polakowsky 134 (B, presumably destroyed

(photo F), lectotype fragment F, here

designated (photo PTBG)). Syntype: COSTA

RICA: In sylvis primaevis Carpinterae, 24 Jun

1875, H. Polakowksy 140 (B, presumably

destroyed). The fragment of Polakowsky 134

at F is here designated as lectotype.

. Nomenclator of Mexican and Central American Rubiaceae

Hoffmannia longipetiolata Pol. var. minor Pol.,

Linnaea 41: 568. 1877. Type: COSTA RICA:

In sylvis primaevis Carpinterae, Jun 1875, H.

Polakowsky 144 (Holotype B, presumably

destroyed).—Hoffmannia longipetiolata Pol.

Hoffmannia macrophylla (Planch.) Hemsl., Biol.

Centr.-Amer., Bot. 2: 37. 1881. Higginsia

macrophylla Planch., FI. Serres 5: 482d. 1849.

Type: Not designated, based on cultivated

material grown from seeds collected in

Guatemala.—Hoffmannia nicotianifolia (M.

Martens & Galeotti) L. O. Williams

Hoffmannia macrophylla Standi., J. Wash.

Acad. Sci. 15: 9. 1925. Not 7/. macrophylla

(Planch.) Hemsl., 1881. Type: COSTA RICA:

Cartago, Cerro de La Carpintera, 1500-1850

m, Feb 1924, P. C. Standley 35636 (Holotype

US 1152957 (photos MEXU, PTBG); Isotype

US 1152958 (photos MEXU, PTBG)).

—Hoffmannia leucocarpa Standi.

Hoffmannia macrosiphon. Standi., Field Mus.

Nat. Hist., Bot. ser. 17: 276. 1937. Type:

GUATEMALA: Quezaltenango, Volcan de

Zunil, 8000 ft., 4 Aug 1934, A. F. Skutch 937

(Holotype F 813468 (photo PTBG); Isotype

A, n.v.).

Hoffmannia manussatani Dwyer, Ann.

Missouri Bot. Gard. 67: 234. 1980. Type:

PANAMA: Bocas del Toro, Robalo Trail, N

slopes of Cerro Horqueta, 6000-7000 ft., 5-7

Aug 1947, Allen 4978 (Holotype MO

1636310).

Hoffmannia mexicana (Link, Klotzsch & Otto)

Hemsl., Biol. Centr.-Amer., Bot. 2: 37. 1881.

Higginsia mexicana Link, Klotzsch & Otto,

Ic. PI. Rar. Hort. Ber. 1: 57, t. 23. 1841. Type:

Not designated, based on material of Mexican

origin cultivated at the Berlin Botanical

Garden.—Hoffmannia excelsa (Kunth) K.

Schum.

Hoffmannia minuticarpa Dwyer & Lorence,

Novon 4: \22,fig. 2c, d. 1994. Type:

MEXICO: Veracruz, Mun. de Soteapan,

Cumbres de Bastonal, ca. 10-11 km al SE de

Tebanca, 1200 m, 26 Apr 1982, R. Cedilla T.,

D. Lorence & G. Ibarra M. 1280 (Holotype

MEXU; Isotype MO).

Hoffmannia montana L. O. Williams,

Fieldiana, Bot. 36: 55. 1973. Type:

GUATEMALA: Quezaltenango, Volcan

Zunil, 8000 ft., 4 Aug 1934, A. F. Skutch 936

(Holotype F 934132 (photos MEXU, PTBG);

Isotype A (photo PTBG)).

Gard. 67: 236. 198^^ T^q^e: PANAMA:

Chiriqui, N of San Felix at Chiriqui-Bocas del

Toro border, on Cerro Colorado copper mine

road along continental divide, 5000-5500 ft.,

5 May 1975, S. Mori & J. Kallunki 5915

(Holotype MO 2259023).—Hoffmannia

vesiculifera Standi.

Hoffmannia nebulosa Standi, in Yunck., Field

Mus. Nat. Hist., Bot. Ser. 17: 396. 1938.

Type: HONDURAS: Comayagua, above the

plains of Siguatepeque, near summit of range

above El Achote, 1850 m, 1 Aug 1936, T. G.

Yuncker, R. F Dawson & H R. Youse 6252

(Holotype F 857687 (photo PTBG)).

Hoffmannia nesiota Donn. Sm., Bot. Gaz.

(Crawfordsville) 61: 374. 1916. Type:

COSTA RICA: Cocos Island, Wafer Bay, 18

Jun 1898, H. Pittier 12387 (Holotype US

941381 (photos MEXU, PTBG)).

Hoffmannia nicotianifolia (M. Martens &

Galeotti) L. O. Williams, Fieldiana, Bot. 36:

56. 1973. Psychotria nicotianifolia M.

Martens & Galeotti, Bull. Acad. Roy. Sci.

Bmxelles 11: 229. 1844 [as nicotianaefolia].

Type: MEXICO: Veracruz, Cordillera pres

Xalapa, 3000 pieds, 1840, H. Galeotti 7066

(Holotype presumably BR, n.v.; Isotype P

(photos MEXU, PTBG)).

Hoffmannia nubium Standi. & L. O. Williams,

Ceiba 1: 167. 1950. Type: HONDURAS:

Morazan, drainage of the Rio Yeguare, at

about long. 87°W, lat. M^N, 6000 ft., 20 Jul

1948, S. F. Glassman 1996 (Holotype US

2215976 (photos MEXU, PTBG); Isotypes F,

GH (photo PTBG), MEXU, WIS).

—Hoffmannia cryptoneura Standi.

Hoffmannia oreogena L. O. Williams,

Fieldiana, Bot. 36: 57. 1973. Type:

HONDURAS: La Paz, Cordillera Guajiquiro,

Montana Verde, aldea Las Marias, 2100 m, 23

May 1964, A. Molina R. & A. R. Molina

Lorence

14052 (Holotype F 1616225 (photo PTBG);

Isotype EAP, n.v.).

Hoffmannia oreophila L. O. Williams,

Fieldiana, Hot. 36: 57. 1973. Type:

NICARAGUA: Matagalpa, Cordillera Central

de Nicaragua between Matagalpa and

Jinotega, Disparate de Potter, near Santa

Maria de Ostuma, 1500 m, 20-24 Feb 1963,

L O. Williams. A. Molina R. & T Williams

24709 (Holotype F 1606614 (photo PTBG);

Isotype EAP, n.v.).—Hoffmannia

nicotianifolia (M. Martens & Galeotti) L. O.

Williams

Hoffmannia orizabensis Standi., Contr. U.S.

Natl. Herb. 20: 205. 1919. Type: MEXICO:

Veracruz, region of Orizaba, 1855, Mueller

1359 (Holotype GH, n.v.).

Hoffmannia ostaurea Dwyer, Ann. Missouri

Bot. Gard. 56: 280. 1969. Type: PANAMA:

Bocas del Toro, Rio Terrible between

Quebrada Huron and Puerto Palenque,

120-140 m, 12 Apr 1968, J. H. Kirkbride. Jr.

& J. A. Duke 501 (Holotype MO 1982893).

Hoffmannia pallidijlora Standi., J. Wash. Acad.

Sci. 15:9. 1925. Type: COSTA RICA: San

Jose, La Hondura, 1300-1700 m, 16 Mar

1924, P. C. Standley 37877 (Holotype US

1153101 (photos MEXU, PTBG)).

—Hoffmannia viridis Rusby

Hoffmannia panamensis Standi., Contr. U.S.

Natl. Herb. 20: 204. 1919. Type: PANAMA:

Chiriqui, above El Boquete, forests along the

Rio Ladrillo and vicinity, 1200-1300 m, 6

Mar 1911, //. Pittier 3056 (Holotype US

677443 (photos MEXU, PTBG)).

—Hoffmannia psychotriifolia (Benth. in

Oerst.) Griseb.

Hoffmannia phoenicopoda K. Schum.,

Notizbl. Bot. Gart. Berlin 2: 276. 1899. Type:

Not designated, based on material cultivated

in the Berlin Botanical Garden of uncertain

origin, probably from Mexico or Central

America.

Hoffmannia piratarum Standi., J. Wash. Acad.

Sci. 18: 180. 1928. Type: COSTA RICA:

Cocos Island, Forets humides de la vallee de

Wafer, 10-200 m, Jan 1902, H Pittier 16259

(Holotype US 579835 (photos MEXU,

PTBG); Isotype GH (photo PTBG)).

Hoffmannia pittieri Standi., Contr. U.S. Natl.

Herb. 18: 140. 1916. Type: PANAMA:

Chiriqui, between Alto de las Palmas and top

of Cerro de la Horqueta, 2100-2268 m, 18

Mar 1911, H Pittier 3247 (Holotype US

677642 (photos MEXU, PTBG)).

Hoffmannia psychotriifolia (Benth. in Oerst.)

Griseb., FI. Brit. W. Ind. 321. \S6\. Higginsia

psychotriifolia Benth. in Oerst., Vidensk.

Meddel. Dansk Naturhist. Foren. Kjobenhavn

1852: 50. 1853 [aspsychotriaefolia]. Type:

COSTA RICA: In monte Barba (Volcan de

Barba), 6000 ft.. May 1847, Oersted 11507

(Holotype C, n.v. (photos F, MO); Isotype US

(photos MEXU, PTBG)).

Hoffmannia pusilla L. O. Williams, Fieldiana,

Bot. 34: 117. 1972. Type: GUATEMALA:

Alta Verapaz, between Sepacuite and

Secoyocte, 1100 m, 24 May 1905, H. Pittier

348 (Holotype US 1083004 (photos MEXU,

PTBG)).—Pinarophyllon bullatum Standi.

Hoffmannia pustulata Dwyer, Ann. Missouri

Bot. Gard. 67: 240. 1980. Type: PANAMA:

Colon, Rio Guanche, pressed from cultivation,

May-Jun 1972, R. L. Dressier s.n. (Holotype

MO 2178490 (photo PTBG)).

Hoffmannia quadrifolia Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 250.

1947. GUATEMALA: Zacapa, Sierra de las

Minas, around the summit of Monte Virgen,

ca. 2400 m, 12-13 Jan 1942, J. A. Steyermark

42600 (Holotype F 1133818 (photo PTBG)).

Hoffmannia racemifera Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 249.

1947. Type: GUATEMALA: Izabal, Cerro

San Gil, 300-900 m, 25 Dec 1941, J. /f.

Steyermark 41910 (Holotype F 1134162

(photos PTBG); Isotypes F 1134161 (photos

PTBG), US (photos MEXU, PTBG)).

Hoffmannia ramonensis Standi., J. Wash. Acad.

Sci. 18: 180. 1928. Type: COSTA RICA: Rio

Barranca a San Juan, pres de San Ramon,

1300-1400 m, 25 Apr 1913, A. Tonduz 17812

(Holotype US 861910 (photos MEXU,

PTBG)).—Hoffmannia dotae Standi.

A Nomenclator of Mexican and Central American Rubiaceae

Hoffmannia refulgens (Hook, f.) HemsL, Biol.

Centr.-Amer., Bot. 2: 37. ISEl. Higginsia

refulgens Hook, f., Bot. Mag. tab. 5346. 1862.

Type: Based on plants grown at Kew from

seed thought to be from South America, but

most probably from Central America or

Chiapas, Mexico (Holotype presumably K,

Hoffmannia regalis (Hook, f.) Hemsl., Biol.

Centr.-Amer., Bot. 2; 37. 1881. Higginsia

regalis Hook, f., Bot. Mag. tab. 5280. 1861.

Type: Based on plants grown at Kew

presumably from Mexican seed (Holotype

presumably K, n.v.). Campylobotrys.

Hoffmannia rexmontis Dywer, Ann. Missouri

Bot. Card. 67: 241. 1980. Type: PANAMA:

Chiriqui, NW side of Cerro Pando, 21 Jul

1971, T. Croat 15934 (Holotype MO

2692337; Isotypes MO 2692338, MO

2150879).—Hoffmannia pallidiflora Standi.

Hoffmannia rhizantha Standi., Carnegie Inst.

Wash. Publ. 461: 90. 1935. Type: BRITISH

HONDURAS (BELIZE): Esperanza Trail,

630 m, 26 Jun 1934, W. A. Schipp S-731

(Holotype F 729968 (photo PTBG)).

Hoffmannia riparia Standi., Field Mus. Nat.

Hist., Bot. Ser. 17: 214. 1937. Type:

GUATEMALA: Quezaltenango, Colomba,

720 m, 24 Sep 1934, A. F. Skutch 1291

(Holotype A, n.v. (photo and fragment F)).

—Hoffmannia regalis (Hook, f.) Hemsl.

Hoffmannia rosei B. L. Rob., Proc. Amer. Acad.

Arts 45: 407. 1910. Type: MEXICO: Nayarit,

Tepic, near Pedro Paulo, 3 Aug 1897, J. N.

Rose 1968 (Holotype US 300860 (photos

MEXU, PTBG); Isotype GH, n.v.).

—Hoffmannia cuneatissima B. L. Rob.

Hoffmannia rotata Donn. Sm., Bot. Gaz.

(Crawfordsville) 18: 204. 1893. Type:

GUATEMALA: Quiche, San Miguel

Uspantan, 7000 ft., Apr 1892, Heyde & Lux

3169B (Lectotype US 943496 (photos MEXU,

PTBG), here designated). Syntype:

GUATEMALA: Quiche, San Miguel

Uspantan, 7000 ft., Apr 1892, Heyde & Lux

3174 (P (photos MEXU, PTBG)). The US

sheet of Heyde & Lux 3169B was selected as

lectotype as it is most complete, with intact

leaves, flowers, and inflorescences.

Hoffmannia rotundata Standi., Contr. U.S. Natl.

Herb. 20: 204. 1919. Type: MEXICO:

Chiapas, Cerro del Boqueron, Jun 1914, C yf.

Purpus 7268 in part (Holotype MO 765057).

—Hoffmannia nicotianifolia (M. Martens &

Galeotti) L. O. Williams

Hoffmannia rubripigmenta Dywer, Ann.

Missouri Bot. Gard. 56: 283. 1969. Type:

PANAMA: Darien, between upper Rio

Membrillo and Camp 7 on construction road

to San Bias, 100-800 m, 26-28 Apr 1967,

J. A. Duke 10870 (Holotype MO 1981851).

Hoffmannia sessilifolia L. O. Williams,

Phytologia 25: 463. 1973. Type:

GUATEMALA: Baja Verapaz, Sierra de las

Minas, above Purulha, 1600 m, 2 Jan 1973,

L. O. Williams, A. Molina R. & T. P. Williams

41977 (Holotype F 1716220 (photo PTBG);

Isotype 1716221).

Hoffmannia steinvorthii Fosberg, Sida 2: 388.

1966. Type: COSTA RICA: Heredia, south

slope of Volcan Barba, 2000 m, 26 Jun 1965,

W. H. Hatheway & A. Jimenez 1461

(Holotype US 2562634 (photos MEXU,

PTBG); Isotypes F, GH, n.v.).—Hoffmannia

arborescens Dorm. Sm.

Hoffmannia steyermarkii Standi., Publ. Field

Mus. Nat. Hist., Bot. Ser. 22: 192. 1940.

Type: GUATEMALA: San Marcos, between

Canjula and La Union, near southeast portion

of Volcan de Tacana, 2000-3000 m, 22 Feb

1940, J. A. Steyermark 36426 (Holotype F

1035015 (photo PTBG); Isotype A, n.v.).

Hoffmannia strigillosa Hemsl., Diag. PL Nov.

31. 1879. Type: MEXICO: Without precise

locality, Salle s.n. (Holotype K, n.v.).

Hoffmannia subauriculata Standi., J. Wash.

Acad. Sci. 18: 179. 1928. Type: COSTA

RICA: Cartago, El Muneco, on the Rio

Navarro, 1400-1500 m, 6-7 Mar 1926, P. C.

Standley & R. Torres R. 50956 (Holotype US

1306554 (photos MEXU, PTBG)).

Hoffmannia teruae L. O. Williams. & A. R.

Molina, Fieldiana, Bot. 36: 59. 1973. Type:

GUATEMALA: San Marcos, near Aldea

Fratemidad, between San Rafael Pie de la

Lorence

Cuesta and Palo Gordo, west facing slopes of

Sierra Madre Mountains, 1800-2400 m,

10-18 Dec 1963, L O. Williams, A. Molina R.

& T. P. Williams 26028 (Holotype F 1651553

(photo PTBG); Isotype EAP, n.v.)-

Hoffmannia tetrastigma Donn. Sm., Bot. Gaz.

(Crawfordsville) 27: 336. 1899. Type:

GUATEMALA: Santa Rosa, Zamorora, 1800

m, Apr 1893, Heyde & Lux 4490 (Holotype

US 354989 (photos MEXU, PTBG); Isotypes

F, US, 2 sheets (photos MEXU, PTBG)).

Hoffmannia tonduzii Standi., Contr. U.S. Natl.

Herb. 20: 205. 1919. Type: COSTA RICA:

Forets de Las Vueltas, Tucurrique, 635-700

m. May 1899, A. Tonduz 133 73 (Holotype US

941386 (photos MEXU, PTBG)).

—Hoffmannia longipetiolata Pol.

Hoffmannia trichocalyx Standi., J. Wash.

Acad. Sci. 18: 181. 1928. Type: COSTA

RICA: Alajuela, vicinity of Fraijanes,

1500-1700 m, 12-13 Feb 1926, P. C.

Standley & R. Torres R. 47690 (Holotype US

1305242 (photos MEXU, PTBG); Isotype US

1305243 (photos MEXU, PTBG)).

Hoffmannia tuerckheimii Donn. Sm., Bot.

Gaz. (Crawfordsville) 47: 254. 1909. var.

tuerckheimii. Type: GUATEMALA: Alta

Verapaz, bei Coban, 1600 m. Mar 1908, H.

von Tuerckheim 11.2160 (Holotype US 941383

(photos MEXU, PTBG); Isotype US 941384

(photos MEXU, PTBG)).

Hoffmannia tuerckheimii Donn. Sm. var.

glabra Standi. & Steyerm., Publ. Field Mus.

Nat. Hist., Bot. Ser. 23: 250. 1947. Type:

GUATEMALA: El Progreso, Sierra de las

Minas, Montana Canahui, between Finca San

Miguel and summit of mountain, upper limits

of Finca Caieta, 1600-2300 m, 10 Feb 1942,

J. A. Steyermark 43807 {43806 in protologue]

(Holotype F 1133774 (photo PTBG)).

Hoffmannia uniHora Standi., Contr. U.S. Natl

Herb. 20: 204. 1919. Type: GUATEMALA:

Alta Verapaz, Coban, 1100 m, Feb 1908, H.

von Tuerckheim 11.2107 (Holotype US 941382

(photos MEXU, PTBG)).

Hoffmannia valerii Standi., J. Wash. Acad. Sci

18: 178. 1928. Type: COSTA RICA:

Guanacaste, El Arenal, 600 m, 20 Mar 1923,

J. Valerio 57 (Holotype US 1206194 (photos

MEXU, PTBG)).

Hoffmannia vesiculifera Standi., Publ. Field

Columbian Mus., Bot. Ser. 4: 285. 1929.

Type: PANAMA: Bocas del Toro, Buena

Vista Camp, Chiriqui Trail, 750 m, Jan-Mar

1928, G. P. Cooper 231 (Holotype F 579284

(photo PTBG)).

Hoffmannia viridis Rusby, Descr. N. Sp. S.

Amer. PI. 134. 1920. Type: COLOMBIA:

Santa Marta, H. H. Smith 2649 (Holotype NY,

n.v., photo and fragment F).

Hoffmannia vulcanicola Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 251.

1947. Type: GUATEMALA: Solola, southern

slopes of Volcan de Atitlan above Finca

Moca, 1000-1250 m, 20 Jun 1942, J. A.

Steyermark 47931 (Holotype F 1132754

(photo PTBG)).

Hoffmannia wilsonii Standi., Publ. Field Mus.

Nat. Hist., Bot. Ser. 22: 117. 1940. Type:

GUATEMALA: Alta Verapaz, Finca Los

Alpes, 1200 m, 21 Mar 1939, C L. Wilson

344 (Holotype F 968020 (photo PTBG)).

Hoffmannia witheringioides Dwyer, Ann.

Missouri Bot. Gard. 67: 244. 1980. Type:

PANAMA: Veraguas, Rio Segundo Braso, 8

km beyond Escuela Agricola Alto Piedra

beyond Santa Fe, 750 m, 24 Jul 1974, T. B.

Croat 25551 (Holotype MO 2214610; Isotype

MO 2207374).

Hoffmannia woodsonii Standi., Ann. Missouri

Bot. Gard. 28: 471. 1941. Type: PANAMA:

Chiriqui, vicinity of Callejon Seco, Volcan de

Chiriqui, 1700 m, 17 Jul 1940, R. E.

Woodson, Jr. & R. W. Schery 503 (Holotype

MO 1210619 (photo PTBG); Isotypes F

1085625 (photo PTBG), GH, n.v.).

—Hoffmannia longipetiolata Pol.

Holtonia Standi., Trop. Woods. 30: 37. 1932.

Type: Holtonia myriantha (Standi.) Standi.

Herein treated as a synonym of Elaeagia

Wedd.]

Holtonia myriantha (Standi.) Standi., Trop.

Woods 30: 37. 1932. Sickingia myriantha

Standi., Publ. Field Columbian Mus., Bot. Ser.

7: 27. 1930. Type: COLOMBIA: Magdalena:

A Nomenclator of Mexican and Central American Rubiaceae

Las Nubes road, region of Santa Marta, 1200

m, 3 Dec 1898, H. H. Smith 1810 (Holotype F

138660; Isotypes MO, NY, n.v., US, n.v., W,

n. v., Y, n.v.). Simira. —Elaeagia myriantha

(Standi.) C. M. Taylor & Hammel

Houstonia L., Sp. PI. 1: 101. 1753. Type:

Houstonia caerulea L. [Herein treated as a

synonym of Hedyotis L., although some

authors (e.g., Terrell, 1996) recognize

Houstonia as a genus.]

Houstonia acerosa (A. Gray) Benth. & Hook, f,

Gen. PI. 2: 60. 1873. Hedyotis acerosa A.

Gray, PI. Wright. 1: 81. 1852. Type: U.S.A.:

Texas, high prairies from Live Oak Creek to

Las Moras, C. Wright 237 (Holotype GH,

n.v.). Oldenlandia. —Hedyotis acerosa A.

Gray

Houstonia acerosa (A. Gray) Benth. & Hook. f.

subsp. polypremoides (A. Gray) Terrell,

Brittonia 31: 168. 1979. Houstonia

polypremoides A. Gray, Proc. Amer. Acad.

Arts 21: 379. 1886. Type: MEXICO:

Chihuahua, Santa Eulalia Mountains, 26 Sep

1885, C. G. Pringle 356 (Lectotype GH,

n.v.; Isolectotypes F, MO (photos MEXU,

PTBG), US (photos MEXU, PTBG);

designated by Lewis, 1968).—Hedyotis

acerosa A. Gray var. polypremoides (A.

Gray) W. H. Lewis

Houstonia angustifolia Michx., FI. Bor. Amer.

1: 85. 1803. Type: U.S.A.: Florida, in

submaritimus Floridae, collector not cited

(Holotype presumably P, n.v.).—Hedyotis

nigricans (Lam.) Fosberg

Houstonia angustifolia Michx. var. scabra S.

Watson, Proc. Amer. Acad. Arts 18: 97. 1883.

Type: MEXICO: Coahuila, Saltillo, Caracol

Mountains, 1-15 Apr 1880, E. Palmer 410

(Holotype GH, n.v.; Isotype US (photos

MEXU, PTBG)).—Hedyotis nigricans

(Lam.) Fosberg

Houstonia arenaria Rose, Contr. U.S. Natl.

Herb. 1: 70. 1890. Type: MEXICO: Baja

California, La Paz, 20 Jan-5 Feb 1890, E.

Palmer 28 (Holotype US 47260 (photos

MEXU, PTBG); Isotypes F, GH,

n.v.).—Hedyotis arenaria (Rose) W. H.

Lewis

Houstonia asperuloides (Benth.) A. Gray, Proc.

Amer. Acad. Arts 5: 158. 1860. Hedyotis

asperuloides Benth., Bot. Voy. Sulphur 19, p/.

13. 1844. Type: MEXICO: Baja California,

Cabo San Lucas, Barclay s.n. (Holotype BM,

n.v.).—Hedyotis asperuloides Benth.

Houstonia asperuloides (Benth.) A. Gray var.

brandegeana (Rose) Wiggins in Shreve 8c

Wiggins, Veg. FI. Sonoran Desert 2: 1399.

1964. Houstonia brandegeanna Rose, Contr.

U.S. Natl. Herb. 1: 83. 1890. Type: MEXICO:

Baja California, La Paz, 20 Jan-5 Feb 1890,

E. Palmer 31 (Holotype US 47259 (photos

MEXU, PTBG); Isotj^es F, GH, n.v.).

—Hedyotis asperuloides Benth. f.

brandegeana (Rose) W. H. Lewis

Houstonia bouvardioides (Seem.) Benth. &

Hook, f, Gen. PI. 2: 60. 1873. Hedyotis

bouvardioides Seem., Bot. Voy. Herald 2:

296. 1856. Type: MEXICO: Without precise

locality [Durango ?], Sierra Madre, Seemann

s.n. (Holotype BM, n.v.; Isotypes C, n.v.,

fragment F (photo PTBG), GH, n.v.).

—Bouvardia bouvardioides (Seem.) Standi.

Houstonia brandegeana Rose, Contr. U.S. Natl.

Herb. 1: 83. 1890. Type: MEXICO: Baja

California, La Paz, 20 Jan-5 Feb 1889, E.

Palmer 31 (Holotype US 47259 (photos

MEXU, PTBG); Isotypes F, GH, n.v.).

—Hedyotis asperuloides Benth. f

brandegeana W. H. Lewis

Houstonia brevipes Rose, Contr. U.S. Natl.

Herb. 1: 83. 1890. Type: MEXICO: Baja

California, Santa Rosalia, 24 Feb-3 Mar 1889,

E. Palmer 202 (Holotype US 47258 (photos

MEXU, PTBG); Isotypes F, GH, n.v., US

(photos MEXU, PTBG)).—Hedyotis

brevipes (Rose) W. H. Lewis

Houstonia coccinea Andr., Bot. Repos, p/. 106.

1800. Type: No herbarium specimens are

known to exist, only the drawing p/. 106.

—Bouvardia ternifolia (Cav.) Schltdl.

Houstonia drymarioides Standi., J. Wash. Acad.

Sci. 18: 162. 1928. Type: MEXICO:

Tamaulipas, on mountains south of Victoria,

1000 m, 9 Apr 1926, R. Runyon 870

A Nomenclator of Mexican and Central American Rubiaceae

(photos MEXU, PTBG)).—Hedyotis palmeri

(A. Gray) W. H. Lewis

Houstonia parvula Brandegee, Zoe 5: 221.

1905. Not Hedyotis parvula (A. Gray)

Fosberg, 1943. Type: MEXICO: Sinaloa,

vicinity of Culiacan, 12 Oct 1904, T. S.

Brandegee s.n. (Holotype UC, n.v.; Isotypes

GH, n.v., US (photos MEXU, PTBG)).

—Hedyotis sinaloae W. H. Lewis

Houstonia peninsularis Brandegee, Zoe 5: 160.

1903. Type: MEXICO: Baja California, Cape

Region, Sierra de la Trinidad, Nov 1902, T S.

Brandegee s.n. (Holotype UC, n.v.; Isotypes

GH, n.v., US (photos MEXU, PTBG)).

—Hedyotis peninsularis (Brandegee) W. H.

Lewis

Houstonia polypremoides A. Gray, Proc. Amer.

Acad. Arts 21: 379. 1886. Type: MEXICO:

Chihuahua, Santa Eulalia Mountains, 26 Sep

1885, C. G. Pringle 356 (Lectotype GH; n.v.;

Isolectotypes F, MO (photos MEXU, PTBG),

US (photos MEXU, PTBG); designated by

Lewis, 1968). Syntype: MEXICO: Chihuahua,

Santa Eulalia Mountains, 29 May 1885, C. G.

Pringle 16 (GH, n.v., US (photos MEXU,

PRBG)).—Hedyotis polypremoides (A.

Gray) Shinners

Houstonia polypremoides var. bigelovii

Greenman, Proc. Amer. Acad. Arts 32: 291.

1897. Type: U.S.A.: Florence Mountains, Jun

1852, Bigelow 437 (Holotype GH, n.v.).

—Hedyotis polypremoides (A. Gray) W. H.

Lewis

Houstonia prostrata Brandegee, Zoe 5: 105.

1901. Not Hedyotis prostrata Blume, 1823

norKorth., 1851. Type: MEXICO: Baja

California, La Palma, Cape Region, 25 Sep

1899, T. S. Brandegee s.n. (Holotype UC,

n.v.; Isotypes GH, n.v., US 382879 (photos

MEXU, PTBG), US 735791 (photos MEXU,

PTBG))—Hedyotis vergrandis W. H. Lewis

Houstonia prostrata Brandegee var. parvula

(Brandegee) Wiggins in Shreve & Wiggins,

Veg. FI. Sonoran Desert 2: 1399. 1964.

Houstonia parvula Brandegee, Zoe 5: 221.

1905. Not Hedyotis parvula (A. Gray)

Fosberg, 1943. Type: MEXICO: Sinaloa,

vicinity of Culiacan, 12 Oct 1904, T. S.

Brandegee s.n. (Holotype UC, n.v.; Isotypes

GH, n.v., US (photos MEXU,

PTBG)).—Hedyotis sinaloae W. H. Lewis

Houstonia rubra Cav., Icon. 5: 48, t. 474, fig. 1.

1799. Type: MEXICO: Hidalgo, Ixmiquilpan,

Nee s.n. (Holotype presumably MA, n.v.).

—Hedyotis rubra (Cav.) A. Gray

Houstonia serpyllacea (Schltdl.) C. L. Sm. ex

Greenman, Proc. Amer. Acad. Arts 32: 284.

1897. Hedyotis serpyllacea Schltdl., Linnaea

9: 599. 1834. Type: MEXICO: Mexico, inter

La Joya et San Salvador, Jun 1829, Schiede &

Deppe 265 (Holotype presumably HAL, n.v.;

Isotypes B (photo MO), F (photo PTBG),

MO).—Hedyotis serpyllacea Schltdl.

Houstonia sharpii Terrell, Brittonia 32: 49Q,fig.

1. 1980. Type: MEXICO: Hidalgo, camino

entre Metepec y Tenango de Doria, entre 16 y

31.5 km al NNE de Tulancingo, 18 km N.NE.

de Metepec, 29 Mar 1974, S. D. Koch 7423

(Holotype US 2754863 (photos MEXU,

PTBG); Isotypes CHAPA, ENCB, MEXU).

—Hedyotis sharpii (Terrell) G. L. Nesom

Houstonia teretifolia Terrell, Brittonia 31: 166.

1979. Type: MEXICO: Coahuila, 12 km NNE

of Las Margaritas on easternmost ridge of

Sierra de las Margaritas, 1300-1400 m, 24

Sep 1972, M. C. Johnston, F. Chiang & T.

Wendt 9508E (Holotype TEX, n.v.).

—Hedyotis teretifolia (Terrell) G. L. Nesom

Houstonia trifiora (Kunth) A. Gray, Proc. Amer.

Acad. Arts 4: 314. 1860. Bouvardia trifiora

Kunth in Humb., Bonpl. & Kunth, Nov. Gen.

Sp. 3: 386. 1820. Type: MEXICO: Without

precise locality, Humboldt & Bonplands.n.

(Holotype P-Bonpland, n.v. (photo MO)).

—Bouvardia multiflora (Cav.) Schult. &

Schult. f.

Houstonia umbratilis B. L. Rob., Proc. Amer.

Acad. Arts 45: 401. 1910. Type: MEXICO:

Nuevo Leon, shaded cliffs of mountains, near

Monterrey, 25 Apr 1906, C. G. Pringle 13877

(Holotype GH, n.v.; Isotypes MO (photos

MEXU, PTBG), US (photos MEXU, PTBG)).

—Hedvotis umbratilis (B. L. Rob.) W. H.

Lewis '

Houstonia wrightii A. Gray, Proc. Amer. Acad.

Arts 17: 202. 1882. Type: MEXICO: Nuevo

Lorence

Leon, shaded cliffs of mountains, near

Monterrey, 25 Apr 1906, C. G. Pringle 13877

(Holotype GH, n.v.; Isotypes MO (photos

MEXU, PTBG), US (photos MEXU, PTBG)).

—Hedyotis wrightii (A. Gray) Fosberg

Houstonia xestosperma (B. L. Rob. &

Greenman) Terrell, Brittonia 32: 492. 1980.

Oldenlandia xestosperma B. L. Rob. &

Greenman, Proc. Amer. Acad. Arts 32: 41.

1896. Type: MEXICO: Oaxaca, Sierra de San

Felipe, open glades, 8000 ft., Jun 1894, C. G.

Pringle 4692 (Lectotype GH, n.v.;

Isolectotypes BM, n.v., F, K, n.v., MO

(photos MEXU, PTBG), MSC, n.v., PH, n.v.,

UC, US, 2 sheets (photos MEXU, PTBG),

VT, n.v.; designated by Terrell, 1980).

Hedyotis .—Bouvardia xestosperma (B. L.

Rob. & Greenman) Terrell & S. D. Koch

Isertia Schreb., Gen. PI. 1: 234, n. 602. 1789.

Type: Isertia coccinea (Aubl.) Vahl

Isertia deamii Bartlett, Proc. Amer. Acad. Arts

43: 59. 1907. Type: GUATEMALA: Izabal,

Puerto Barrios, 24 Feb 1905, C. C. Deam 48

(Holotype GH, n.v.).—Isertia haenkeana

DC.

Isertia deamii Bartlett var. stenophylla Bonn.

Sm., Bot. Gaz. (Crawfordsville) 61: 374.

1916. Type: COSTA RICA: Boca Culebra,

plaines des Pacifique, 50 m, 20 Jan 1898, H.

Pittier 11989 (Holotype US 941413 (photo

PTBG); Isotypes F (photo PTBG), P, n.v.,

fragment F).—Isertia haenkeana DC.

Isertia haenkeana DC., Prodr. 4: 437. 1830.

Type: MEXICO: Without precise locality,

Haenkes.n. (Holotype presumably PR, n.v.).

Isertia laevis (Triana) B. M. Boom, Brittonia

36: 433. 1984. Cassupa laevis Triana, Ann.

Sci. Nat., Bot., Ser 4, 9: 44. 1858. Type:

COLOMBIA: Cundinamarca, Triana 1838

(Lectotype P; Isolectotypes BR, n.v., COL,

n.v., L, n.v., P, n.v., US, n.v.; designated by

Boom, 1984).

Isertiapanamensis (Standi.) Standi., Publ. Field

Columbian Mus., Bot. Ser. 8: 346. 1931.

Cassupa panamensis Standi., Contr. U.S.

Natl. Herb. 18: 135. 1916. Type: PANAMA:

Colon, along Rio Fato, 10-100 m, 8 Jul 1911,

H. Pittier 3889 (Holotype US 678963 (photo

PTBG); Isotype US (photo PTBG)).—Isertia

laevis (Triana) B. M. Boom

Isertia scorpioides B. M. Boom, Brittonia 36:

435. 1984. Type: PANAMA: Colon, Atlantic

side of Isthmus on road to Pina, 25 Aug 1967,

J. H. Kirkbride, Jr. & S. M. V. Hayden 316

(Holotype NY, n.v.; Isotypes MO (2 sheets;

photos PTBG), NY, n.v.).

Ixora L., Sp. PI. 1: 110. 1753. Type: Ixora

coccinea L.

Ixora alba Sesse & M09., La Naturaleza, ser. 2,

I. app. 16. 1887. Not Ixora alba L., 1753, nor

Roxb., 1814, nor Sus. ex Wight & Am., 1834.

Type: MEXICO: Habitat in India ad montibus

Mazatlan (presumably MA, n.v.).—Status

uncertain.

Ixora americana Jacq., Hort. Schonbr. 3: 4, pi.

257. 1798. Type: Cultivated at Schobmnn,

Vienna, said to have been brought from

Mexico.—Bouvardia ternifolia (Cav.)

Schltdl.

Ixora aphylla Sesse & M09., Pi. Nuev. Hispan.

16. 1887; La Naturaleza, ser. 2, 1. app. 16, ed.

2, 15. 1887. Type: MEXICO: Habitat prope

Xochistlan, Sesse & Mogino s.n. (presumably

MA, n.v.).—Status uncertain.

Ixora casei Hance in Walp., Ann. Bot. Syst. 2:

754. 1852. Type: CAROLINE ISLANDS:

Strong Island, Q. IV. Case s.n. (Isotype or

possible Holotype K, n.v.). For typification

see Fosberg & Sachet (1989a), who note that

var. casei is cultivated in Mesoamerica.

Ixora coccinea L., Sp. PI. 1: 110. 1753.

Lectotype: Rheede, Hortus malabaricus 2:17,

t. 13. 1679. Designated by Fosberg & Sachet

(1989b), who note that Ixora coccinea L. var.

coccinea and var. intermedia Fosberg &

Sachet are cultivated in Mexico and Central

America.

Ixora coccinea L. var. intermedia Fosberg &

Sachet, Baileya 23: 80. 1989. Type: VIRGIN

ISLANDS: St. Croix, Teague Bay, cultivated,

F. R.. Fosberg 55279 (Holotype US, n.v.)

—Ixora coccinea L.

Ixora finlaysoniana Wall, ex G. Don, Gen.

Hist. Dichlam. PI. 3: 572. 1834. Type: EAST

Lorence

C G. Pringle 6641 (Holotype GH, n.v.;

Isotypes A, n.v., MO, P (2 sheets; photos

MEXU, PTBG), US (photos MEXU, PTBG)).

—Machaonia acuminata Humb. & Bonpl.

Machaonia galeottiana Baill., Bull. Mens. Soc.

Linn. Paris 1: 204. 1879. Type: CUBA: Le

Havane, H. Galeotti s.n. (Holotype P (photo

MO)). On the holotype label the word

“Veracruz” is crossed out and “Le Havane”

written in. As noted by Standley (1934: 222)

the collection resembles no Mexican species.

—Machaonia havanensis (Jacq.) Alain

Machaonia hahniana Baill., Bull. Mens. Soc.

Linn. Paris 1: 204. 1879. Type: MEXICO:

Puebla, Acatlan, Juillet 1865, L Hahn s.n.

(Holotype P (photos MEXU, MO, PTBG)).

Machaonia havanensis (Jacq. ex J. F. Gmel.)

Alain, Phytologia 8: 370. 1962. Spermacoce

havanensis Jacq. in J. F. Gmel., Syst. Nat. 2:

234. 1791. Type: Based on Spermacoce

fruticosa Jacq., ined.? (type not located).

Machaonia lindeniana Baill., Bull. Mens. Soc.

Linn. Paris 1: 204. 1879. Type: MEXICO:

Campeche, Campeche, fl. en Juin, (Yucatan),

Linden s.n. (Holotype P (photos MEXU, MO,

PTBG)).

Machaonia martinezorum Lorence, Novon 4:

122 Jig. 3. 1994. Type: MEXICO: Chiapas,

Mun. de Ocosingo, a 50 km al S de Boca

Lacantum, 130 m, 24 Jun 1986, E. Martinez S.

19016 (Holotype PTBG 7356; Isotypes

MEXU, MO).

Machaonia martinicensis (DC.) Standi., Publ.

Field Mus. Nat. Hist., Bot. Ser. 22: 193. 1940.

Tertrea martinicensis DC., Prodr. 4: 481.

1830. Type: MARTINIQUE: Richard s.n.

(Holotype P, n.v.).

Machaonia martinicensis (DC.) Standi, var.

dodgei (Standi.) Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 22: 193. 1940. Machaonia

rotundata Griseb. var. dodgei Standi., Publ.

Field Mus. Nat. Hist., Bot. Ser. 18: 1326.

1938. Type: COSTA RICA: Beach between

Rio Sandio and Rio Tigre, 1 May 1930, C. W.

Dodge 7752 (Holotype F 890888 (photo

PTBG); Isotype MO (photo PTBG)).

—Machaonia martinicensis (DC.) Standi.

Machaonia pringlei A. Gray, Proc. Amer.

Acad. Arts 21: 380. 1886. Type: MEXICO:

Coahuila, limestone ledges, mountains,

Jimulco, 27 Apr 1885, C. G. Pringle 146

(Holotype GH, n.v.; Isotypes A, n.v., F

(photos MEXU, PTBG), P (photos MEXU,

PTBG), US (photos MEXU, PTBG)).

Machaonia rotundata Griseb. var. dodgei

Standi., Publ. Field Mus. Nat. Hist., Bot. Ser.

18: 1326. 1938. Type: COSTA RICA: Beach

between Rio Sandoval and River Tigre, 1 May

1930, C. W. Dodge 7752 (Holotype F 890888

(photo PTBG); Isotype MO (photo PTBG)).

—Machaonia martinicensis (DC.) Standi.

Machaonia tysonii Dwyer, Aim. Missouri Bot.

Gard. 53: 106. 1966. Type: PANAMA. Los

Santos, thicket near man-made dam, main

highway, Monagre Beach, J. D. Dwyer 5067

(MO, not located in 6/1996). Note: John

Dwyer (pers. comm. 24 March 1993) believes

citation of this collection was a typographical

error and the following collection actually

represents the holotype: PANAMA: Along

Quebrada Vital off Rio Chucanagre, km 16 de

Yavita, 4 Jul 1962, J. A. Duke 5067 (Holotype

MO 3725554 (photo PTBG)). In the Flora of

Panama Rubiaceae treatment, Dwyer (1980)

erroneously cited Dwyer 5061 as the holotype;

the latter specimen was not located at MO in

3/1993 or 6/1996.—Machanonia acuminata

Humb. & Bonpl.

Machaonia velutina M. Martens & Galeotti,

Bull. Acad. Roy. Sci. Bruxelles 11: 134. 1844.

Type: MEXICO: Veracruz, ravines de Puente

Nacional, pres Veracruz, H. Galeotti 7104

(Holotype BR, n.v.; Isotype P (photos MEXU,

MO, PTBG)).—Machaonia acuminata

Humb. & Bonpl.

Machaonia veracruzeana Baill., Bull. Mens.

Soc. Linn. Paris 1: 204. 1879. Norn, superfl.,

based on Machaonia velutina M. Martens &

Galeotti, 1844.—Machaonia acuminata

Humb. & Bonpl.

Macrocnemum P. Browne, Hist. Jam. 165.

1756. Type: Macrocnemum jamaicense L.

Macrocnemum candidissimum Vahl, Symb. 2:

38, p/. 30. 1791. Type: TRINIDAD: Vicinia

100

Lorence

Manettia Mutis ex L., Mant. PL: 553, 558. 1771,

nom. cons. Type: Manettia reclinata L.

Manettia arboricola Lorence, Novon 3: 59.

1993. Type: PANAMA: Comarca de San

Bias, El Llano-Carti Rd., km 16.7, 350 m, trail

west to waterfall 5 km from road, 16 Jun

1985, G. de Nevers & S. Charnley 5907

(Holotype MO 3623655 (photo PTBG)).

Manettia barbata Oerst., Vidensk. Meddel.

Dansk Naturhist. Foren. Kjobenhavn 1852:

47. 1853. Type: COSTA RICA: In Monte

Aguacate, 2000 ft., Feb 1846, Oersted 11521

(Holotype C, n.v. (photos MO, US); Isotypes

K, n.v., US (2 sheets; photos MEXU, PTBG)).

Manettia bocataurensis Dwyer, Ann. Missouri

Bot. Gard. 67: 278. 1980. Type: PANAMA:

Bocas del Toro, along RR (railroad) near

station at Milla 10, 27 Jul 1971, T. 5. Croat &

D. M. Porter 16234 (Holotype MO 2166138

(photo PTBG)).—Hedyotis lancifolia K.

Schum.

Manettia costaricensis Wemham, J. Bot. 41

(suppl.): 38. 1919. Syntypes: COSTA RICA:

Tucurrique, deboisements de Las Vueltas,

650-700 m, Dec 1898, A. Tonduz 12873 (B,

presumably destroyed (photos F, MO;

fragment F), BM, n.v., US 1390734 (photos

MEXU, PTBG), US 354369). COSTA RICA:

Without precise locality, 3500 ft., Endres 2

(W, n.v. (fragment F)).—Manettia reclinata

Manettia estrellae Standi., J. Wash. Acad. Sci.

15: 6. 1925. Type: COSTA RICA: Cartago,

La Estrella, 26-27 Mar 1924, P. C Standley

39224 (Holotype US 1153146 (photos

MEXU, PTBG)).—Manettia Oexilis

Brandegee

Manettia flexilis Brandegee, Univ. Calif. Publ.

Bot. 6: 196. 1915. Type: MEXICO: Chiapas,

Cerro del Boqueron, Jun 1914, C. A. Purpus

7118 (Holotype UC 174907, n.v.; Isotypes F,

GH, n.v., MO (photos MEXU, PTBG), US

(photos MEXU, PTBG)).

Manettia hirtella M. Martens & Galeotti, Bull.

Acad. Roy. Sci. Bruxelles 11: 238. 1844.

Type: MEXICO: Oaxaca, cordillere orientale,

au bords des rivieres, dans les forets de

Tonaguia, 4000-5000 pieds, Jun 1840, H.

Galeotti 2635 (Holotype BR, n.v.).—Status

uncertain.

Manettia hydrophila Dwyer, Ann. Missouri Bot.

Gard. 67: 280. 1980. Type: PANAMA:

Veraguas, mouth of Rio Concepcidn, in

swamp, 4 Dec 1967, W. H. Lewis, T. B. Croat

& J. L. Hawker 2822 (Holotype MO

2268375).—Sipanea biflora (L. f.) Cham. &

Schltdl.

Manettia liebmannii Standi., J. Wash. Acad. Sci.

17: 337. 1927. Type: MEXICO: Oaxaca,

Pelado [probably Cerro Pelon], Liebmann

11485 [=Liebmann Rubiaceae 147] (Holotype

C, n.v. (fragment US; photos MEXU,

PTBG)).—Hedyotis galeottii (M. Martens)

Terrell & Lorence

Manettia iongicalycina Dwyer & Lorence,

Novon 3: 61. 1993. Type: PANAMA: Code,

Alto Calvario, 800-900 m, 20 Apr 1977, J. P.

Folsom & Jaslon 2665 (Holotype MO

2608777 (photo PTBG)).

Manettia longipedicellata C. M. Taylor, Novon

5: 2^2, fig. I A. 1995. Type: COSTA RICA:

Limon, Canton de Matina, Cordillera

Talamanca, 200 m, aguas abajo de la

confluencia de la quebrada Canabral con el rio

Barbilla, 10°00T0"N, 83°25'30"W, 5 Nov

1988, G. Herrera 2290 (Holotype CR, n.v.;

Isotype MO 3649361).

Manettia microphylla Lorence & Dwyer,

Novon 3: 60. 1993. Type: PANAMA: Darien,

Pirre Massif, Alturas de Nique, above Cana

Mine, 850-1150 m, 2 Mar 1988, G.

McPherson 12189 (Holotype MO 3648261

(photo PTBG); Isotypes PTBG, SCZ).

Manettia orbifera Wemham, J. Bot. 57 (suppL):

41. 1919. Type: GUATEMALA: Retalhuleu,

Bernoulli & Cairo 1649 (Holotype K, n.v.).

—Manettia reclinata L.

Manettia panamensis Duchass. & Walp.,

Linnaea 23: 753. 1850. Type: PANAMA:

Duchassaing s.n. (presumably GOET, n.v.).

—Manettia reclinata L.

Manettia reclinata L., Mant. PI. 2: 558. 1771.

Type: Not designated. Note: this species is

actually based on Colombian material seen by

Mutis, not Mexican material. Apparently

Linnaeus believed that Santa Fe de Bogota

ANomenclatorc

AND Central American Rubi>^

A Nomenclator of Mexican and Central American Rubiaceae

Palicourea phanaerandra Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Dot. Ser. 23: 252.

1947. Type: GUATEMALA: Izabal, Cerro

San Gil, uppermost ridges and summit,

1200-1300 m, 26-27 Dec \9A\,J. A.

Steyermark 41952 (Holotype F 1132612

(photo PTBG); Isotype US).—Psychotria

phanaerandra (Standi. & Steyerm.) Lorence

Palicourea purpurea C. M. Taylor, Syst. Bot.

Monogr. 26: l\,fig. 33 c, d. 1989. Type:

PANAMA: Chiriqui, NW of Boquete along

road to Palo Alto Arriba, from 2 km until road

ends then another 0.5 km by trail, 1400-1600

m, 13 May 1984, C. M. Taylor 3201

(Holotype DUKE, n.v.; Isotypes F (photo

PTBG), MO (photo PTBG)).

Palicouraea purpusii Standi., Publ. Field

Columbian Mus., Bot. Ser. 8: 67. 1930. Type:

MEXICO: Veracruz, Zacuapam [Zacuapan],

shady forest. May 1928, C. A. Purpus WHO

(Holotype F 602907 (photos MEXU, PTBG);

Isotype MO (photos MEXU, PTBG)).

—Palicourea macrantha Loes.

Palicourea rigidifolia (Dwyer & Hayden)

Dwyer, Ann. Missouri Bot. Gard. 67: 313.

1980. Cephaelis rigidifolia Dwyer & Hayden,

Ann. Missouri Bot. Gard. 55: 37. 1968. Type:

PANAMA: Panama, Cerro Jefe, 2700 ft., 20

Aug 1967, J. D. Dwyer 8075A (Holotype MO

2165097 (photo PTBG)).—Status uncertain,

probably Psychotria subg. Heteropsychotria

Palicourea salicifolia Standi., J. Wash. Acad.

Sci. 18: 280. 1928. Type: COSTA RICA: San

Jose, Laguna de La Chonta, northeast of Santa

Maria de Dota, 2000-2100 m, 18 Dec 1925,

P. C. Standley 42174 (Holotype US 1252646

(photo PTBG)).

Palicourea seleri Loes., Verb. Bot. Vereins

Prov. Brandenburg 65: 114. 1923. Type:

GUATEMALA: Huehuetenango, zwischen

Trinidad und Rosario, 13 Aug 1896, E. Seler

& C. Seler 3049 (Syntype B, destroyed

(photos F, GH, MO, NY, US), fragment F,

GH, n.v.); Yalambahoch, E. Seler & C. Seler

2676 (Syntype B, destroyed).—Psychotria

galeottiana (M. Martens) C. M. Taylor &

Lorence

Palicourea skotakii C. M. Taylor, Syst. Bot.

Monogr. 26: 76, fig. 41. 1989. Type: COSTA

RICA: San Jose, along a road to the S from

the Panamerican Highway at km 67, about 1

km along the road, 2700-2800 m, 5 Mar

1985, C. M. Taylor & C. Skotak 4756

(Holotype DUKE, n.v.; Isotypes F (photo

PTBG), MO (photo PTBG)).

Palicourea spathacea C. M. Taylor, Syst. Bot.

9: 226. 1984. Type: COSTA RICA: Cartago,

along the trail about 6 km beyond the gate to

the meteorology station in Alto Patillos,

Tapanti Hydroelectric Project, 7 Apr 1976, J.

Utley & K. Utley 4538 (Holotype DUKE, n.v.;

Isotypes CAS, n.v., CR, n.v., F (photo PTBG),

US (photo PTBG)).

Palicourea standleyana C. M. Taylor, Syst.

Bot. Monogr. 26: SI, fig. 45 d-f 1989. Norn,

nov. for Psychotria brenesii Standi., Publ.

Field Mus. Nat. Hist., Bot. Ser. 18: 1347.

1938. Not Palicourea brenesii Standi., 1938.

Type: COSTA RICA: Alajuela, Los Angeles

de San Ramon (Cerros de La Balsa), Mar

1928, Brenes 6050 (Holotype F 854521

(photos MEXU, PTBG)).

Palicourea stevensonii Standi., Trop. Woods.

16: 42. 1928. Type: BRITISH HONDURAS

(BELIZE): Middlesex, Jul 1927, N. S.

Stevenson C(Yale 10683) (Holotype F

572673; Isotype NY, n.v.).—Psychotria

capitata Ruiz & Pav.

Palicourea subrubra Pol., Linnaea 41: 571.

1877. Type: COSTA RICA: San Jose, Cerro

de La Carpintera, 24 Jun 1875, Polakowsky

200A (Holotype B, presumably destroyed

(photo F)).—Palicourea padifolia (Willd. ex

Roem. & Schult.) C. M. Taylor & Lorence

Palicourea talamancana Standi. & L. O.

Williams, Ceiba 1: 250. 1951. Type: COSTA

RICA: Cartago, Cordillera de Talamanca, near

Vuelta Alta, Cerro de las Vueltas, 3200 m, 10

Apr 1949, L. O. Williams 16264 (Holotype

EAP, n.v.; Isotypes F (photo PTBG), GH,

n.v., US (photo PTBG)).—Palicourea

brenesii Standi.

Palicourea tilaranensis C. M. Taylor, Syst.

Bot. Monogr. 26: 84,^g. 3 a. b. 1989. Type:

COSTA RICA: Puntarenas-Alajuela border

112

Lorence

region, in and around Monteverde Nature

Reserve, mostly on Pacific watershed,

10° 18'N, 84°47’N, 1450-1650 m, 29 Jun

1977, R. O. Lawton 1189 (Holotype F

1885069 (photo PTBG)).

Palicourea triphylla DC., Prodr. 4: 526. 1830.

Type: FRENCH GUIANA: Guiana Gallica,

without precise locality, Patris s.n. (Holotype

G-DC, n.v. (photo MO)). Psychotria.

Palicourea tubiflora Dwyer, Ann. Missouri

Bot. Gard. 67: 316. 1980. Type: PANAMA:

Panama, Cerro Jefe, along trail on ridge

running NE from summit, ca. 1000 m, 11 May

1975,5. Mori & J. Kallunki 6082 (Holotype

MO 2271726 (photo PTBG)).

Palicourea tumidonodosa Dwyer, Ann.

Missouri Bot. Gard. 67: 317. 1980. Type:

PANAMA: Panama, Cerro Mali, base camp

near Colombian border, lower montane wet

forest, 1500 m, 22 Jan 1975, A. Gentry & S.

Mori 13787 (Holotype MO 2259310 (photo

PTBG)).

Palicourea veraguensis Dwyer, Ann. Missouri

Bot. Gard. 67: 318. 1980. Type: PANAMA:

Chiriqui, Cerro Colorado, 50 km N of San

Felix on the continental divide, 1200-1500 m,

17 Aug 1975, S. Mori & R. Dressier 7796

(Holotype MO 2777356 (photo PTBG)).

—Palicourea lasiorrhachis Oerst.

Palicourea vestita Standi., J. Wash. Acad. Sci.

18: 277. 1928. Type: COSTA RICA: San

Jose, near Quebradillas, about 7 km north of

Santa Maria de Dota, 1800 m, 24 Dec 1925,

P. C. Standley 42909 (Holotype US 1253015

(photos F, PTBG)).

Pavetta L., Sp. PI. 1: 110. 1753. Type: Pavetta

indica L.

Pavetta odoratissima Sesse & M09., FI. Mexic.

20. 1893; La Naturaleza, ser. 2, 2: app 20

1893. Type: MEXICO ?: Habitat in Praedio

Sacarino de la Langosta, Equitis Gonzalo de

Herrera, floret Agusto, Sesse & Mogino s.n.

(presumably MA, n.v.).—Status uncertain.

Pavetta racemosa Sesse & M09., FI. Mexic. 20.

1893; La Naturaleza, ser. 2, 2: app. 2, 18.

1893. Type: MEXICO: Without precise

locality, Sesse & Mogino 543 (presumably

MA, n.v.; probable Isotype F).—Rondeletia

villosa Hemsl.

Pentagonia Benth., Bot. Voy. Sulphur t. 39.

1804; 105. 1845 (nom. cons.). Type:

Pentagonia macrophylla Benth.

Pentagonia alba Dwyer, Ann. Missouri Bot.

Gard. 67: 321. 1980. Type: PANAMA:

Darien, Cerro Pirre, Bristan 1224 (Holotype

MO, not located 6/1996; Isotype NY, n.v.).

Pentagonia alfaroana (Standi.) Standi., J. Wash.

Acad. Sci. 17: 171. 1927. Watsonamra

alfaroana Standi., J. Wash. Acad. Sci. 15:

287. 1925. Type: COSTA RICA: Alajuela,

vicinity of Capulin, on the Rio Grande de

Tarcoles, 80 m, 2 Apr 1924, P. C. Standley

40194 (Holotype US 1152712 (photos

MEXU, PTBG); Isotype F).—Pentagonia

tinajita Seem.

Pentagonia brachyotis (Standi.) Standi., J.

Wash. Acad. Sci. 17: 171. 1927. Watsonamra

brachyotis Standi., Contr. U.S. Natl. Herb. 17:

443. 1914. Type: PANAMA: Darien,

Marraganti, 3 Apr 1908, R. S. Williams 999

(Holotype US 678351 (photos MEXU,

PTBG)).

Pentagonia costaricensis (Standi.) W. C.

Burger «fe C. M. Taylor, Fieldiana, Bot., n.s.

33: 213. 1993. Nothophlebia costaricensis

Standi., Contr. U.S. Natl. Herb. 17: 438. 1914.

T>pe: COSTA RICA: Collines de Moin,

littoral atlantique, Nov 1899, H. Pittier 16024

(Holotype US 578472 (photos MEXU,

PTBG)).

Pentagonia donnell-smithii (Standi.) Standi., J.

Wash. Acad. Sci. 17: 170. 1927. Watsonamra

donnell-smithii Standi., Contr. U.S. Natl.

Herb. 17: 442. 1914. Type: COSTA RICA: La

Emilia, Llanuras de Santa Clara, 250 m, Apr

1896, J. D. Smith 6590 (Holotype US 355176

(photos MEXU, PTBG); Isotype F).

Pentagonia gymnopoda (Standi.) Standi., J.

Wash. Acad. Sci. 17: 171. 1927. Watsonamra

gymnopoda Standi, Contr. U.S. Natl. Herb.

17: 444. 1914. Type: PANAMA: Colon,

Loma de la Gloria, near Fato (Nombre de

Dios), 10-104 m, 8 Jul 1911, //. Pittier 3858

(Holotype US 678935 (photos MEXU,

A Nomenclator of Mexican and Central American Rubiaceae

113

PTBG); Isotype US 678934 (photos MEXU,

PTBG)).

Pentagonia hirsuta Standi., J. Wash. Acad. Sci.

18; 165. 1928. Type: COSTA RICA:

Talamanca, forets au-dessus de Tsaki, 500 m.

Mar 1895, A. Tonduz 9415 (Holotype US

938648 (photos MEXU, PTBG); Isotype F).

Pentagonia lobata C. M. Taylor, Novon 5. 202,

fig. ID. 1995. Type; COSTA RICA:

Puntarenas. Canton de Golfito, Reserva

Natural de Vida Silvestre Golfito, final del

camino hacia las torres, 8°39'52"N,

83°10’00''W, 7 Dec 1992, B. Hammel 18619

(Holotype MO 4355639; Isotypes CR, n.v., F,

n.v., MO 4355638).

Pentagonia macrophylla Benth., Bot. Voy.

Sulphur 105, tab. 39. 1845. Type: PANAMA:

Panama, forests between Panama and Cruces,

Barclay s.n. (Holotype BM, n.v.).

Watsonamra.

Pentagonia nuciformis Dwyer, Ann. Missouri

Bot. Gard. 67: 326. 1980. Type: PANAMA:

Veraguas, 5 miles W of Santa Fe on the road

past Escuela Agricola Alto Piedra on Pacific

side of divide, 800-1200 m, 18 Mar 1973,

T. B. Croat 23093 (Holotype MO 2161493

(photo PTBG)).

Pentagonia pinnatifida Seem., J. Bot. Hooker

7: 566. 1848. Type: PANAMA: Darien, banks

of the river Cupica, Seemann 1019 (Holotype

BM, n.v. (photo MO)).

Pentagonia pittieri (Standi.) Standi., J. Wash.

Acad. Sci. 17: 170. 1927. Watsonamra pittieri

Standi., Contr. U.S. Natl. Herb. 17: 443. 1914.

Type: PANAMA: San Bias, forests around

Puerto Obaldia, San Bias coast, 0-50 m, Aug

1911,//. Pittier 4298 (Holotype US 679414

(photos MEXU, PTBG); Isotype US 679415

(photos MEXU, PTBG)).—Pentagonia

pinnatifida Seem.

Pentagonia pubescens (Standi.) Standi., J.

Wash. Acad. Sci. 17: 171. 1927. Watsonamra

pubescens Standi., Contr. U.S. Natl. Herb. 17:

441. 1914. Type: PANAMA: Canal Zone,

around Tabemilla, along the railroad, 20-25

m, 6 Jul 1911, //. Pittier 3822 (Holotype US

678894 (photo PTBG); Isotype NY, n.v.).

—Pentagonia macrophylla Benth.

Pentagonia tinajita Seem., Bot. Voy. Herald

134, / 28. 1854. Type; PANAMA: Veraguas,

western parts, Seemann 1595 (Holotype BM,

n.v. (photo MO)).

Pentagonia veraguensis Dwyer, Ann. Missouri

Bot. Gard. 67: 328. 1980. Type: PANAMA:

Veraguas, Valley of Rio Dos Bocas, 11 km

from Escuela Agricola Alto Piedra (above

Santa Fe) on road to Calovebora, 450 m, 30

Aug 1974, T. B. Croat 27504 (Holotype MO

2201425 (photo PTBG)).

Pentagonia wendlandii Hook, f, Bot. Mag. pi.

5230. 1861. Type: CENTRAL AMERICA

[probably Costa Rica or Panama]: Described

from cultivated material, type uncertain.

Pentas, Benth. in Hook., Bot. Mag. 4086. 1844.

Type: Pentas camea Benth.

Pentas lanceolata (Forssk.) Defiers, Voy.

Yemen 142. 1889. Ophiorhiza lanceolata

Forssk., F. Aegypt.-Arab. 42. 1775. Type:

AFRICA: Yemen, Hadie Mts., Forsskals.n.

(Holotype C, n.v.; Isotype BM, n.v.). Widely

cultivated in Mexico and Mesoamerica.

Pentodon Hochst., Flora 27: 552. 1844. Type:

Pentodon decumbens Hochst.

Pentodon pentandrus (Schumach. & Thonn.)

Vatke, Osterr. Bot. Zeitschr. 25: 231. 1875.

Hedyotis pentandra Schumach. & Thonn.,

Kongel. Danske Vidensk. Selsk.

Naturvidensk., Math. Afh. 3: 71. 1827. Type:

GUINEA: Aquapim, “S” [Haaslund Smith?]

(location unknown). Naturalized at least in

Mexico (Veracruz), Belize, Nicaragua, and

Costa Rica.

Petesia P. Browne, Civ. Nat. Hist. Jamaica 143,

t. 2, 3. 1756. Type not designated.

Petesia grandis Bartl. ex DC., Prodr. 4: 395.

1830. Type: MEXICO; Without precise

locality, Haenke s.n. (Holotype presumably

PR, n.v., fragment G-DC, n.v. (microfiche 4:

395.1)).—Sommera grandis (Bartl. ex DC.)

Standi.

Petesia minor Griseb., Bonplandia 6: 8. 1858.

Type: PANAMA: Without precise locality.

Lorence

Seemann s.n. (Holotype presumably BM,

n.v.).—Sommera donnell-smithii Standi.

Petesia repens Sesse & M09., FI. Mexic. 21.

1893. Type: MEXICO: Habitat in Ahualuci

pratis, florit Octobris, Sesse & Mogifio s.n.

(presumably MA, n.v.).—Status uncertain.

Petesia stipularis Sesse & M09., FI. Mexic. 21.

1893, La Naturaleza 2, 2: App. 2, 16. 1893.

Not Petesia stipularis L., 1759. Type (listed

twice): MEXICO: Habitat in Montibus

Mazatlanensibus, floret Maio, habitat in

Acahuitzotlae montibus, floret Julio, Sesse &

Mogino s.n. (presumably MA, n.v.).—Status

uncertain, possibly Chomelia or Sommera.

Protologue states leaves are prominently

lineolate beneath and inflorescence is once

dichotomous.

Pinarophyllon Brandegee, Univ. Calif. Publ.

Bot. 6: 71. 1914. Type: Pinarophyllon flavum

Brandegee

Pinarophyllon bullatum Standi., N. Amer. FI.

32: 87. 1921. Type: GUATEMALA: Alta

Verapaz, between Sepacuite and Secoyocte,

1100 m, 24 May 1905, H. Pittier 348

(Holotype US 473013 (photos MEXU, PTBG,

US)).

Pinarophyllon flavum Brandegee, Univ. Calif.

Publ. Bot. 6: 71. 1914. Type: MEXICO:

Chiapas, Finca Mexiquito, Jun 1913, C. A.

Purpus 6700 (Holotype UC 172447, n.v.;

Isotypes F (photo PTBG), GH, n.v., US

(photos MEXU, PTBG)).

Pittoniotis Griseb., Bonplandia 6: 8. 1858. Type:

Pittoniotis trichantha Griseb.

Pittoniotis protracta (Bartl. ex DC.) Griseb.,

Bonplandia 6: 8. 1858. Guettardaprotracta

Bartl. ex DC., Prodr. 4: 457. 1830. Type:

MEXICO: Without precise locality, Haenke

s.n. (Holotype presumably PR, n.v.).

Anisomeria.—ChomeMz protracta (Bartl ex

DC.) Standi.

Pittoniotis trichantha Griseb., Bonplandia 6: 8.

1858. Type: PANAMA: Without precise

locality, Duchassaing s.n. (possibly P, n.v.).

Antirhea.

Placocarpa Hook. f. in Benth. & Hook, f., Gen.

PI. 2: 107. 1873. Type: Placocarpa mexicana

Hook. f.

Placocarpa mexicana Hook. f. in Benth. &

Hook, f., Gen. PI. 2: 108. 1873. Type:

MEXICO: Veracruz, Orizaba, Apr 1855,

Botteri 913 (Lectotype P, (photo PTBG), here

designated; Isolectotypes F (fragment), K

(photo PTBG). Syntypes: MEXICO:

Veracruz, Orizaba, Botteri J/d (F, K (photo

PTBG)); Bourgeau 3022 (K (photo PTBG)).

The sheet of Botteri 913 at P, with duplicates

in two other herbaria and copious flowers and

fruits, represents the most appropriate choice

for lectotype.

Plocaniophyllon Brandegee, Univ. Calif. Publ.

Bot. 6: 69. 1914. Type: Plocaniophyllon

flavum Brandegee

Plocaniophyllon flavum Brandegee, Univ.

Calif Publ. Bot. 6: 69. 1914. Type: MEXICO:

Chiapas, Finca Mexiquito, Jun 1913, C.

Purpus 70/P (Holotype UC 173077, n.v.;

Isotypes F (photos MEXU, PTBG), US

(photos MEXU, PTBG)).

Pogonopus Klotzsch, Monatsber. Konigl.

Preuss. Acad. Wiss. Berlin 1853: 500. 1853.

Type: Pogonopus ottonis Klotzsch

Pogonopus exsertus (Oerst.) Oerst., Amer.

Centr. 17. t. 13. 1863. Macrocnemum

exsertum Oerst., Vidensk. Meddel. Naturhist.

Foren Kjobenhavn 1852: 45. 1853. Type:

COSTA RICA: Jaris, Oersted (Holotype C,

n.v.). Note: Pogonopus speciosus (Jacq.) K.

Schum., a species from northern South

America, was erroneously recorded from

Central America by Standley & Williams

(1975) and Dwyer (1980).

Portlandia P. Browne, Hist. Jamaica 164, /. 11.

1756. Type: Portlandia grandiflora L.

Portlandia ghiesbreghtiana Bail!., Adansonia

12: 300. 1879. Type: MEXICO: Oaxaca, pres

de Hacienda de Huijastla au bord des ravine,

1842-3, Ghiesbreght 27 (Holotype P (photos

MEXU, PTBG); Isotypes F, P (3 sheets;

ANOMENCLATOR OFl

I AND Central American Rubiaceae

117

Jose, La Hondura, 1300-1700 m, 16 Mar

1924, P. C Standley 37892 (Holotype US

1153107 (photos MEXU, PTBG)).

ANomenclator of]

I AND Central American Rubiaceae

A Nomenclator c

NTRAL American Rubiaceae

A Nomenclator of Mexican and Central American Rubiaceae

139

(Lectotype US 941393, here designated

(photos MEXU, PTBG); Isolectotypes F

(photos MEXU, PTBG), US 577282).

Syntype: COSTA RICA: Bords du Rio

Corozal, pres Santo Domingo de Golfo Duke,

Apr 1896, A. Tonduz 9982 (US, F (photos

MEXU, PTBG)). Tonduz 9878, with more

abundant flowers and two duplicates,

represents the better choice for lectotype.

Randia grayumii Dwyer & Lorence in W. C.

Burger & C. M. Taylor, Fieldiana, Bot., n.s.

33: 285. 1993. Type: COSTA RICA: Limon,

behind NAISA house, Gandoca, 9°36'N,

82°37'W, 3 m, 4 May 1985, M. H. Grayum &

G. Schatz 5287 (Holotype MO 3648263

(photo PTBG); Isotype ?CR, n.v.).

Randia guatemalensis Standi., Contr. U.S. Natl.

Herb. 20: 202. 1919. Type: GUATEMALA:

Alta Verapaz, Secanquim, 550 m, 6 May

1905, H. Pittier271 (Holotype US 472930

(photos MEXU, PTBG)).—Randia aculeata

L. var. aculeata

Randia guerrerensis Lorence & Rodr. Acosta,

Biotica 11: \95,fig. 1. 1986. Type: MEXICO:

Guerrero, Mun. de Coyuca de Benitez,

camino de El Zapote a Hacienda de Cabanas,

a 10 km al SO de la alberca El Dorado, cerca

de la Laguna de Mitla, 7 m, 21 Jul 1984, M.

Rodriguez Acosta sub Lorence 4584

(Holotype MEXU; Isotypes ENCB, MO

(photo PTBG)).

Randia habrophlebia Standi. & Steyerm., Publ.

Field Mus. Nat. Hist., Bot. Ser. 23: 25. 1943.

Type: GUATEMALA: Jutiapa, Valley of

Jutiapa, 850 m, 24 Oct-6 Nov 1940, P. C.

Standley 76321 (Holotype F 1091237 (photos

MEXU, PTBG)).—Randia cinerea (Femald)

Standi.

Randia hidalgensis Lorence, Bol. Soc. Bot.

Mexico 47: 3%, fig. la, b. 1987. Type:

MEXICO: Queretaro, Mun. de Arroyo Seco,

12 km NW de El Trapiche, camino a Rio

Verde, a 1 km SE de Puerto de Ayutla, 1200

m, 29 Abr 1982, P. Tenorio & R. Hernandez

M. 297 (Holotype MEXU; Isotype MO (photo

PTBG)).

Randia hondensis Karsten, Linnaea 30: 154.

1859. Type: COLOMBIA: Valley of Rio

Magdalena, 260 m, H. Karsten s.n. (Holotype

B, presumably destroyed).

Randia induta Standi., Contr. U.S. Natl. Herb.

23: 1377. 1926. Type: MEXICO: Guerrero,

Cayacol, 12 Apr 1903, E. W. Nelson 7018

(Holotype US 399362 (photos MEXU,

PTBG)).

Randia inermis Sesse & M09., FI. Mexic. 54.

1891. Not Randia inermis (Ducke) Ducke,

1932. Type: MEXICO: Habitat in Amatitani

clivo, ubi Testiculus Taurinus ob Fructus.

Figuram vulgo adpellatur. Florit Maio, Sesse

& Mogiiio s.n. (presumably MA, n.v.).

—Status uncertain.

Randia karstenii Pol, Linnaea 41: 568. 1877.

Syntypes: COSTA RICA: In sepibus vivis

prope San Jose, Jun & Sep 1875, H. Karsten

74 (B, presumably destroyed); Karsten 302

(B, presumably destroyed, BM, n.v.).

—Randia aculeata L. var. aculeata

Randia iaetevirens Standi, Contr. U.S. Natl.

Herb. 23: 1377. 1926. Type: MEXICO:

Nuevo Leon, Monterrey, Cerro la Silla, 20

Mar 1902, E. W. Nelson 6083 (Holotype US

347312 (photos MEXU, PTBG); Isotype F

(photos MEXU, PTBG)).

Randia laevigata Standi, Contr. U.S. Natl.

Herb. 20: 201. 1919. Type: MEXICO: Sonora,

Sierra de Alamos, 18 Mar 1910, J. N. Rose,

P. C. Standley & P. G. Russell 13051

(Holotype US 635870 (photos MEXU,

PTBG)).

Randia lasiantha (Standi.) Standi, Contr. U.S.

Natl. Herb. 20: 201. 1919. Basanacantha

lasiantha Standi, Contr. U.S. Natl. Herb. 18:

134. 1916. Type: PANAMA: Panama,

Alajuela, Chagres Valley, 30-100 m, H.

Pittier 3473 (Holotype US 678517 (photos

MEXU, PTBG)).

Randia latifolia Lam. var. micrantha Schltdl,

Linnaea 6: 723. 1831. Type: MEXICO:

Veracruz, prope Jalapam, Schiede & Deppe

276 (Holotype presumably HAL, n.v.).

—Randia aculeata L. var. aculeata

Randia letreroana Lundell, Wrightia 4: 126.

1969. Type: MEXICO: Chiapas, Letrero, near

Siltepec, 2000 m, 6 Jul 1941, E. Matuda 4358

(Holotype TEX-LL, n.v.; Isotypes F (photo

Lorence

PTBG), MEXU, MO (photo PTBG)).

—Randia thurberi S. Watson

Randia longiloba Hemsl., Biol. Centr.-Amer.,

Bot. 4: 101. 1887. Type: MEXICO: Yucatan,

Cozumel Island, Gaunter s.n. (Holotype

presumably K, n.v.).

Randia loniceroides Dwyer & Lorence, Bol.

Soc. Bot. Mexico 47: 40, fig. Ic, d. 1987 [as

lonicerioides]. Type: MEXICO: Veracruz,

Mun. de Minatitlan, Zona de Uxpanapa, 13.7

km E. de La Laguna sobre camino a

Uxpanapa, luego 8 km N sobre camino a

Belisario Dominguez, 17°21'N, 94°23'W, 130

m, 19 Apr 1981, T. Wendt & E. Lott 3207

(Holotype MEXU; Isotypes CHAPA, ENCB).

Randia lundelliana Standi, in Lundell, Contr.

Univ. Michigan Herb. 4: 30. 1940. Type:

BRITISH HONDURAS (BELIZE): El Cayo

Distrist, Vaca, 26 Apr 1938, P. H. Gentle

2524 (Holotype F 941519 (photo PTBG);

Isotype TEX-LL).—Randia monantha

Benth.

Randia malacocarpa Standi., Contr. U.S. Natl.

Herb. 20: 202. 1919. Type: MEXICO:

Nayarit, near Acaponeta, Tepic, 30 Jul 1897,

J. N. Rose 3298 (Holotype US 302274 (photos

MEXU, PTBG, US)).

Randia matudae Lorence & Dwyer, Bol. Soc.

Bot. Mexico 47: 42. 1987. Based on Casasia

chiapensis Miranda, Ceiba 4: \42, fig. p. 144.

1954. Not Randia chiapensis Standi., 1926.

Type: MEXICO: Chiapas, Rancho el Ocote,

30 km NW de Ocozocuautla, 650 m, 21 May

1950, F. Miranda 7734 (Lectotype MEXU;

designated by Lorence & Dwyer, 1987).

Syntype: MEXICO: Chiapas, same locality, F.

Miranda 6224 (F (photo PTBG)).

Randia mayana Lundell, Wrightia 5: 322.

1976. Type: GUATEMALA: Baja Verapaz,

Nino Perdido, in high forest on top of hill, E.

of km 150/151, 30 Aug 1975, C. L. Lundell &

E. Contreras 19758 (Holotype TEX-LL

(photos MEXU, PTBG); Isotypes F, MO

(photo PTBG), TEX-LL (photos MEXU,

PTBG)).

Randia megacarpa Brandegee, Zoe 5: 257.

1908. Type: MEXICO: Baja California,

Comondu, 2 Feb 1889, T. S. Brandegee s.n.

(Holotype UC, n.v.).—Randia capitata DC.

Randia megalocarpa Dwyer, Ann. Missouri

Bot. Gard. 67: 449. 1980. Type: PANAMA:

Panama, Cerro Penon, 3 km S of Alcade Diaz,

vicinity of television tower on Continental

Divide, 350-440 m, M. Nee 8865 (Holotype

MO 2208620 (photo PTBG); Isotype MO

2208621 (photo PTBG)).—Randia lasiantha

(Standi.) Standi.

Randia microcarpa Sesse & M09., PI. Nov.

Hispan. 33. 1888 (La Naturaleza ser. 2, 1:

app. 33. 1888) [as microcarpus]. Type:

MEXICO: Habitat in montibus Haereditatis-

Ahuesuyo, Sesse & Mogiho s.n. (presumably

Randia millspaughiana S. F. Blake, Proc. Biol.

Soc. Wash. 34: 45. 1921. Type: MEXICO:

Yucatan, Maxacanu, Mar 1916, G. F. Gaumer

et al. 23260 (Holotype F 446691 (photos

MEXU, PTBG); Isotypes GH, n.v., US

(photos MEXU, PTBG)).—Randia longiloba

Hemsl.

Randia mira Dwyer, Ann. Missouri Bot. Gard.

67: 450. 1980. Type: PANAMA: Panama,

5-10 km NE of Altos de Pacora at end of

road, 6 Mar 1975, S. Mori & J. Kallunki 4953

(Holotype MO 2247822 (photos MEXU,

PTBG); Isotype MO (photos MEXU, PTBG)).

Randia mitis L., Sp. PI. 2: 1192. 1753. Type:

Habitat in Jamaica. Lectotype: Lycium majus

americanum, jasmini fiore, foliis subrotundis

lucitus, Plukenet, Phytographic a, t. 97, fig. 6.

1691. (designated by Lorence in Jarvis et al.,

1993: 80).—Randia aculeata L. var.

aculeata

Randia mollifolia Standi., Field Mus. Nat.

Hist., Bot. Ser. 17: 219. 1937. Type:

MEXICO: Sonora, Saguaribo, Rio Mayo, 3

Nov 1935, H. S. Gentry 2139 (Holotype F

809526 (photos MEXU, PTBG); Isotypes GH,

n.v., MEXU, US (photos MEXU, PTBG),

WIS).

Randia monantha Benth., PI. Hartweg. 84.

1841. Type: GUATEMALA: Rio de los

Esclavos, T. Hartweg 582 (Holotype K

(photos F, MEXU); Isotypes B, presumably

. Nomenclator of Mexican and Central American Rubiaceae

141

destroyed (photo MO), BM, P [label states

“Mexique”] photos MEXU, PTBG)).

Randia montana L. O. Williams, Phytologia 24:

160. 1972. Type: HONDURAS, Intibuca,

Cordillera Opalaca, between Calaveras and El

Duraznillo, 1800 m, 12 Mar 1970, A. Molina

R. <& A. R. Molina 25550 (Holotype F

1695297 (photos MEXU, PTBG); Isotype

EAP, n.v.).—Randia aculeata L. var.

aculeata

Randia nelsonii Greemn., Proc. Amer. Acad.

Arts 34; 574. 1899. Type: MEXICO: Oaxaca,

Juchitan to Chivela, 150-900 ft., 1895, E. W.

Nelson 2635 (Holotype GH, n.v.; Isotype US

569319 (photos MEXU, PTBG)).—Randia

thurberi S. Watson

Randia nicaraguensis Lorence & Dwyer,

Novon 8: 241, fig. 1. 1998. Type:

NICARAGUA: Esteli, 4.9-7.6 km NE of

Hwy. 1 at Esteli along road to Yali, ca.

13°08-09', 86°19-20’W, 1100 m, 14 Nov

1979, W. D. Stevens & A. Grijalva 15553

(Holotype MO 2872306; Isotypes F, HNMN,

MEXU, MO).

Randia oaxacana Standi., Contr. U.S. Natl.

Herb. 32(3): 165. 1934. Type: MEXICO:

Oaxaca, Choapam [=Santiago Choapan],

2800-1500 ft., 28-29 Jul 1894, E. W. Nelson

842 (Holotype F 599712 (photos MEXU,

PTBG)).

Randia obcordata S. Watson, Proc. Amer.

Acad. Arts 24: 53. 1889. Type: MEXICO:

Sonora, Guaymas, 1887, E. Palmer 648

(Holotype US 47988 (photos MEXU, PTBG);

Isotype BM).

Randia obovata Kunth in Humb., Bonpl. &

Kunth, Nov. Gen. Sp. 3: 409. 1820. Not

Randia obovata Ruiz Pav., 1799. Type:

COLOMBIA; Ad ostia fluminis Sinu,

A. Bonplands.n. (Holotype P-Bonpland,

n.v.).—Randia aculeata L. var. dasyclada

Steyerm.

Randia panamensis Standi., Publ. Field

Columbian Mus., Bot. Ser. 4: 288. 1929.

Type: PANAMA: Prov. Bocas del Toro,

region of Almirante, Flat Rock, Jan-Mar

1928, a Proctor Cooper 213 (Holotype F

579675 (photos MEXU, PTBG)).—Randia

armata (Sw.) DC. subsp. panamensis

(Standi.) Lorence

Randia pepoformis Dwyer, Ann. Missouri Bot.

Gard. 67: 451. 1980. Type: PANAMA:

Darien, trail from Tacarcuna Village on Rio

Tacarcuna to Cerro Mali, 800-1300 m, 16 Jan

1975, A. H. Gentry & S. Mori 13614

(Holotype MO 2248019 (photos MEXU,

PTBG); Isotypes F (photos MEXU, PTBG),

MO (photos MEXU, PTBG)).—Randia

altiscandens (Ducke) C. M. Taylor

Randia petenensis Lundell, Wrightia 4: 127.

1969. Type: GUATEMALA: Peten, La

Cumbre, about 5 km from the village, on

Pusila River Road, 17 May 1967, E.

Contreras 6941 (Holotype TEX-LL (photos

MEXU, PTBG); Isotypes F, US (photos

MEXU, PTBG)).

Randia pittieri (Standi.) Standi., Contr. U.S.

Natl. Herb. 20: 201. 1919. Basanacantha

pittieri Standi., Contr. U.S. Natl. Herb. 18:

134. 1916. Type; COSTA RICA: Gent

Junction, pres Matina, Feb 1900, H. Pittier

16036 (Holotype US 578473 (photos K,

MEXU, PTBG); Isotype P (fragment F,

photos MEXU, PTBG)).

Randia pleiomeris Standi., Contr. U.S. Natl.

Herb. 20: 202. 1919. Type: GUATEMALA:

Santa Rosa, Santa Rosa, 3000 ft.. May 1892,

Heyde & Lux 3166 (Lectotype US 888490,

here designated [left- and right-hand

collections only; the center collection is R.

cookii]; Isolectotype GH, n.v.).—Randia

thurberi S. Watson

Randia pringlei (S. Watson) A. Gray, Proc.

Amer. Acad. Arts 21: 379. 1886.

Basanacantha pringlei S. Watson, Proc.

Amer. Acad. Arts 18: 98. 1883. Type:

MEXICO; Coahuila, Mountains, Jimulco, 9

Apr 1885, C. G. Pringle 142 (Holotype GH,

n.v.; Isotypes A, n.v., BM, F (photos MEXU,

PTBG), MEXU, P (photos MEXU, PTBG),

US (photos MEXU, PTBG)).

Randia pterocarpa Lorence & Dwyer, Bol.

Soc. Bot. Mexico 47: 43, fig. 2. 1987. Type:

MEXICO: Veracruz, Mun. San Andres

Tuxtla, Estacion de Biologia Tropical Los

Tuxtlas, jardin botanico, 95°04-09'W,

142

Lorence

18°34-36'N, 160 m, 6 Dec 1983, G. Ibarra

M. 1109 (Holotype MEXU; Isotypes BM, F,

MO, XAL).

Randia purpusii Greenm. & C. H. Thomps.,

Ann. Missouri Bot. Card. 1; 410. 1914. Type:

MEXICO: San Luis Potosi, Minas de San

Rafael, May 1911, C. Purpus 5208

(Holotype MO 4186182 (photos MEXU,

PTBG); Isotypes F, (photos MEXU, PTBG),

GH, n.v., MO (photos MEXU, PTBG), US

(photos MEXU, PTBG)).—Randia capitata

DC.

Randia retroflexa Lorence & M. Nee, Brittonia

39: 371, fig. 1. 1987. Type: MEXICO:

Veracruz, Mun. de Hidalgotitlan,

Campamento Hermanos Cedillo a 4 km de

Pancho Villa, 150 m, 25 Mar 1975, B. Ortiz &

J. Martiniano A. 56 (Holotype MEXU;

Isotypes BM, BR, ENCB, F (photo PTBG),

MO (photo PTBG), NY, UC, XAL).

Randia rhagocarpa Standi., Contr. U.S. Natl.

Herb. 23: 1376. 1926. Type: MEXICO:

Tamaulipas, vicinity of Victoria, 320 m, 1

Feb-9 Apr 1907, E. Palmer 38 (Holotype US

572266 (photos MEXU, PTBG); Isotypes F

(photos MEXU, PTBG), GH, n.v., MO

(photos MEXU, PTBG)).—Randia

obcordata S. Watson

Randia rosei Standi., Contr. U.S. Natl. Herb. 20-

203. 1919. Type: MEXICO: Sinaloa, Rosario,

7 Jul 1897, J. N. Rose 1551 (Holotype US

300395 (photos MEXU, PTBG); Isotype F

(photos MEXU, PTBG), GH, n.v.).—Randia

thurberi S. Watson

Randia sonorensis Wiggins, Contr. Dudley

Herb. 3: 15,pi. 19, figs. 4-6. 1940. Type:

MEXICO: Sonora, granitic hillside 12 miles E

of Ures on road to Babiacora, 2760 ft., 20 Sep

1934, 1. L. Wiggins 7360 (Holotype DS

263358, n.v.; Isotype US (photos MEXU,

PTBG)).

Randia standleyana L. O. Williams, Phytologia

24: 162. 1972. Type: GUATEMALA: Peten,

La Libertad, 29 May 1933, C. L Lundell 3474

(Holotype F 692287 (photos MEXU, PTBG);

Isotype MICH, n.v.).

Randia subcordata (Standi.) Standi., Contr. U.S.

Natl. Herb. 20: 201. 1919. Basanacantha

subcordata Standi., Contr. U.S. Natl. Herb.

18: 133. 1916. Type: COSTA RICA: Buissons

et bois des chemins a Nicoya, May 1900, A.

Tonduz 13934 (Holotype US 861251 (photos

MEXU, PTBG); Isotypes B, destroyed

(fragment F, photos MEXU, PTBG), GH, n.v.,

LE, n.v. (fragment F, photo PTBG), P (photos

MEXU, PTBG), fragment F (photos MEXU,

PTBG)).—Randia monantha Benth.

Randia tetracantha (Cav.) DC., Prodr. 4: 387.

1830. Mussaenda tetracantha Cav., Icon. PI.

4: 20, pi. 435. 1799. Type: MEXICO:

[Probably Guerrero], cerca Acapulco, Nee s.n.

(Holotype MA, n.v. (photos F, MEXU, MO)).

Basanacantha.

Randia tetracantha Sesse «fe M09., PI. Nov.

Hispan. 33. 1888 (La Naturaleza ser. 2, 1:

app. 33. 1888). 'Not Randia tetracantha (Cav.)

DC., 1830. Type: MEXICO: Habitat in

montibus Haereditatis-Ahuesuyo, fl. Dec,

Sesse & Mogiho s.n. (presumably MA,

n.v.).—Status uncertain.

Randia tetramera Loes., Verhandl. Bot. Ver.

Brandenburg 65: 109. 1923. Type: MEXICO:

Yucatan, Xkombec and Uxmal, E. Seler & C.

Seler 4035 (Syntype B, presumably destroyed,

GH, n.v.); E. Seler & C. Seler 5559a (Syntype

B, presumably destroyed).—Randia truncata

Greenm. & C. H. Thomps.

Randia texensis Lundell, Field & Lab. 13: 22.

1945. Type: U.S.A.: Texas, Cameron County,

about 5 miles north of Brownsville off Point

Isabel road, 21 Sep 1943, C. L. Lundell &

A. A. Lundell 12456 (Holotype SMU,

n.v.).—Randia obcordata S. Watson

Randia thurberi S. Watson, Proc. Amer. Acad.

Arts 24: 53. 1889. Type: MEXICO: Sonora,

Guayamas, 1887, ThurberinE. Palmer 229

(Holotype US 1393134 (photos MEXU,

PTBG); Isotypes BM, US).

Randia tomatillo Loes., Rep. Spec. Nov. 18:

360. 1922. Type: MEXICO: Veracruz, Puerto

de Alvarado, and Isla de Sacrificios, E. Seler

& C. Seler 4498 (Lectotype US 1205748, here

designated; Isolectotypes B, destroyed; GH,

n.v.). Same locality, E. Seler & C. Seler 5433

(Syntype B, destroyed). The US collection of

Seler & Seler 4498 is a good fruiting

Lorence

Loes., Verb. Bot. Ver. Brandenb. 65: 105.

1923. Type: GUATEMALA: Huehuetenango,

Nenton Distr., near Chacula, E. Seler & C.

Seler 3015 (Holotype B, presumably

destroyed (photos F, MO, US), fragment F).

—Rondeletia gratissima (Planch. & Linden)

Hemsl.

Rogiera stenosiphon (Hemsl.) Borhidi, Acta

Bot. Acad. Sci. Hung. 28: 67. 1982.

Rondeletia stenosiphon Hemsl., Diag. PI.

Nov. Mexic. 26. 1879. Type: MEXICO:

Yucatan and Tabasco, without precise

locality, E. P. Johnson s.n. (Holotype K

(photo PTBG)).—Rondeletia stenosiphon

Hemsl.

Rogiera strigosa (Benth.) Borhidi, Acta Bot.

Acad. Sci. Hung. 28: 67. 1982. Bouvardia

strigosa Benth., PI. Hartweg. 75. 1841. Type:

GUATEMALA: Quezaltenango, [Volcan]

Zunil, Nov., T. Hartweg 530 (Lectotype K

(photo PTBG), here designated; Isolectotypes

G-DEL, n.v. (fragment F), P (photos F,

MEXU, MO, PTBG), W, n.v. (photos F,

MO)). Syntype: GUATEMALA: Volcan de

Fuego, 6000-6400 ft., Salvin s.n. (K (photo

PTBG, fragment F)). The K specimen of

Hartweg 530 is the better choice for lectotype,

with duplicates in three other herbaria.

—Rondeletia strigosa (Benth.) Hemsl.

Rogiera subscandens (Lundell) Borhidi, Acta

Bot. Hung. 35: 311. 1989. Rondeletia

subscandens Lundell, Wrightia 5: 328. 1976.

Type: GUATEMALA: Peten, La Cumbre,

about 500 m west of km 142/143 of the

Cadenas Road, 10 Sep 1976, C. L. Lundell &

E. Contreras 19830 (Holotype TEX-LL

(photos MEXU, PTBG); Isotypes F (photo

PTGB), K (photo PTBG)).—Rondeletia

subscandens Lundell

Rogiera suffhitescens (Brandegee) Borhidi, Acta

Bot. Acad. Sci. Hung. 28: 67. 1982.

Rondeletia suffrutescens Brandegee, Univ.

Calif. Publ. Bot. 6: 70. 1914. Type: MEXICO:

Chiapas, Cerro del Boquerdn, Aug 1913, C. A.

Purpus 6704 (Holotype UC 172451, n.v.;

Isotypes F (photos MEXU, PTBG), MO, US

(photos MEXU, PTBG)).—Rondeletia

suffrutescens Brandegee

Rogiera versicolor (J. Sm.) Lindl. «fe Paxton,

Paxton FI. Gard. 2: 69. July 1851. Rondeletia

versicolor J. Sm., Bot. Mag. 7: tab. 4579.

1851. Type: Based on cultivated material

grown at Kew sent by Seemann from

Boquete, Veraguas, Panama (type not

designated).—Rondeletia amoena (Planch.)

Hemsl.

Ronabea Aubl., Hist. PI. Guiane 1: 154; 2

(Index): 26, /. 59. 1775. Type: Ronabea

latifolia Aubl. [Herein treated as a synonym of

Psychotria L.]

Ronabea erecta Aubl., Hist. PI. Guiane 1: 156.

1775. Type: FRENCH GUIANA: In sylvis

Oyao, Orapu, Sinemari, Aublet s.n. (Holotype

presumably BM, n.v.).—Psychotria erecta

(Aubl.) Standi. & Steyerm.

Rondeletia L., Sp. PI. 1: 172. 1753. Lectotype:

Rondeletia americana L. (Designated by

Howard, 1989.)

Rondeletia acuminata (Oerst. ex Standi.)

Lorence & Cast.-Campos, Biotica 13: 147.

1988. Sommera acuminata Oerst. ex Standi.,

J. Wash. Acad. Sci. 17: 340. 1927. Type:

MEXICO: Oaxaca, Amatlan, F. Liebmann

11712 (Holotype C, n.v. (photo MEXU);

Isotype US (photos MEXU, PTBG)).

Arachnothryx .—Rondeletia scabra Hemsl.

Rondeletia aetheocalymna Donn. Sm., Bot.

Gaz. (Crawfordsville) 42: 298. 1906. Type:

GUATEMALA: Alta Verapaz, trail between

Sepacuite and Secanquim, 550-1000 m, 14

Jan 1905, W. R. Maxon & R. Hay 3275

(Holotype US 473244 (photos MEXU,

PTBG); Isotype US (photos MEXU, PTBG)).

Arachnothryx.

Rondeletia affinis Hemsl., Diag. PI. Mex. 28.

1879. Type: MEXICO: Veracruz, Zongolica

[Sierra de Zongolica], Jun 1866, Sumichrast

[sub Botteri 1017] (Lectotype K, here

designated; Isolectotypes K (photo PTBG), P

(2 sheets, photos MEXU, MO, PTBG)).

Syntypes: MEXICO: Chiapas, Ghiesbreght

686 (K (photo PTBG), P (photos MEXU,

PTBG), MO). COSTA RICA: Endres 226

(K). PANAMA: Veraguas, Seemann 1594

A Nomenclator of Mexican and Central American Rubiaceae

(K). The K specimen of Sumichrast sub

Botteri Wilis the best choice for lectotype

and has duplicates in two other herbaria.

—Rondeletia buddleioides Benth.

Rondeletia albida Lundell, Wrightia 5: 323.

1976. Type: MEXICO: Chiapas, Carelas, near

Motozintla, 2176, Apr 1945, E. Matuda 5509

(Holotype TEX-LL (photos MEXU, PTBG);

Isotypes F, MEXU, TEX-LL).—Rondeletia

laniflora Benth.

Rondeletia amoena (Planch.) Hemsl., Diag. PI.

Mex. 26. 1879. Rogiera amoena Planch., FI.

Serr. Jard. 5: tab. 442, figs 1. 2. 1849. Type:

Based on plants cultivated in Belgium,

presumably from Guatemalan seed (type not

designated).

Rondeletia aprica Lundell, Contrib. Univ.

Michigan Herb. 7: 54. 1942. Type: MEXICO:

Chiapas, Saxchanal, Sierra Madre, in

advanced forest, 1-5 Jul 1941, £. Matuda

43/4 (Holotype TEX-LL (photos MEXU,

MO, PTBG); Isotypes A, n.v., F, MO, US

(photos MEXU, PTBG)). Rogiera.

—Rondeletia cordata Benth.

Rondeletia aspera Standi., N. Amer. FI. 32: 54.

1918. Type: COSTA RICA: San Jose, Pont du

Rodeo, Pacacol (Villa Colon), Jan 1891, //.

Pittier 3243 (Holotype US 941408 (photo

PTBG)). Arachnothryx.

Rondeletia atravesadensis Lorence, Novon 1:

137,y?g. lA, B. 1991. Type: MEXICO:

Oaxaca, Cerro Atravesada-Cerro Azul,

Arroyo Rana, 6 Dec 1956, T. MacDougall s.n.

(Holotype MEXU 253128 (photo PTBG)).

Rondeletia backhousii Hook.f, Bot. Mag. 103:

tab. 6290. 1877. Based on material of

unknown origin cultivated at Kew, obtained

from the Backhouse establishment in York

(type not designated). Rogiera .—Status

uncertain, resembles Rondeletia panamensis

DC. but is nearly glabrous.

Rondeletia belizensis Standi., Carnegie Inst.

Wash. Publ. 461: 91. 1935. Type: BRITISH

HONDURAS (BELIZE): Jacinto Hills, 300

ft., 28 Aug 1933, W. A. Schipp 1201

(Holotype F 683600 (photos MEXU, PTBG);

Isotypes GH, n.v., K (photo PTBG), MO).

Arachnothryx.

Rondeletia belizensis Standi, var. longiloba

Lundell, Wrightia 5: 323. 1976. Type:

GUATEMALA: Alta Verapaz, Rubelsanto,

2.3 km SEE [=ESE] of Balastrera, 20 Jul

1975, C. L. Lundell &E. Contreras 19545

(Holotype TEX-LL (photos MEXU, PTBG);

Isotypes F, MO (photo PTBG)).—Rondeletia

belizensis Standi.

Rondeletia bertieroides Standi., Publ. Field

Columbian Mus., Bot. Ser. 4: 267. 1929.

Type: PANAMA: Bocas del Toro, Buena

Vista Camp, Chiriqui Trail, 900 m, 1928,

G. P. Cooper 598 (Yale 12231) (Holotype F

579614 (photo PTBG); Isotype NY, n.v.).

Arachnothryx.

Rondeletia bourgaei Standi., N. Amer. FI. 32:

56. 1918. Type: MEXICO: Veracruz, Region

de Orizaba, Monte San Cristobal, 8 Jul 1866,

E. Bourgeau 2618 (Holotype US 49964

(photos MEXU, PTBG); Isotypes P (4 sheets,

photos MEXU, PTBG)). Arachnothryx.

—Rondeletia laniflora Benth.

Rondeletia brachistantha Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 25.

1943. Type: GUATEMALA: San Marcos, Rio

Vega, near San Rafael and Guatemala-Mexico

boundary, Volcan Tacana, 2500-3000 m, 20

Feb 1940, J. A. Steyermark 36286 (Holotype

F 1050045 (photos MEXU, PTBG); Isotype

US (photos MEXU, PTBG)). Rogiera.

—Rondeletia cordata Benth.

Rondeletia brandegeeana Lorence, Novon 1:

138. 1991. Based on Otocalyx chiapensis

Brandegee, Univ. Calif Publ. Bot. 6: 69.

1914. Not Rondeletia chiapasensis Standi.,

1940. Type: MEXICO: Chiapas, Cerro del

Boqueron, middle region, moist valley slopes,

Aug 1913, C. A. Purpus 7041 (Holotype UC

173047 (photo PTBG); Isotypes GH, US

(fragment)).

Rondeletia breediovei Lorence, Novon 4: 129,

fig. 6. 1994. Type: MEXICO: Chiapas, Mun.

of Tenejapa, paraje of Mahben Chauk, on

slopes along the Tana Te’ River near Shal

K’esh, 2900 ft., 27 Nov 1964, D. E. Breedlove

7682 (Holotype F 1617735 (photo PTBG);

Isotype CAS (photo PTBG)).

150

Lorence

Rondeletia brenesii Standi., Publ. Field Mus.

Nat. Hist., Bot. Ser. 18: 1370. 1938. Type:

COSTA RICA: Camino de San Ramon a La

Calera, 20 Mar 1929, A. M. Brenes 6757

(Holotype F 854972 (photo PTBG)).

Arachnothryx.

Rondeletia buddleioides Benth., PI. Hartweg.

69. 1840. Type: MEXICO: Oaxaca, inplanitie

Llano Verde, T. Hartweg 502 (Holotype K

(photo PTBG)); Isotypes B (destroyed, photo

MO, fragment G-DEL), K (photo PTBG),

NY, n.v. (fragment US, photos MEXU,

PTBG), P (photos MEXU, PTBG)).

Arachnothryx.

Rondeletia buddleioides Benth. var. aspera J. H.

Kirkbr., Ann. Missouri Bot. Gard. 55: 379.

1968. Type; PANAMA: Bocas del Toro,

between Criollo (just above Buena Vista) and

Quebrada Higueron on Chiriqui Trail, 18 Apr

1968, J. H. Kirkbride, Jr. & J. A. Duke 780

(Holotype MO 1942973 (photos MEXU,

PTBG); Isotype US). Arachnothryx.

—Rondeletia buddleioides Benth.

Rondeletia calycosa Donn. Sm., Bot. Gaz.

(Crawfordsville) 56: 59. 1913. Type: COSTA

RICA: San Jose, forets de La Palma, 1460 m,

25 Sep 1898, A. Tonduz 11633 (Holotype US

no 941406 (photos F, MEXU, PTBG)).

Arachnothryx.

Rondeletia capitellata Hemsl., Diag. PI. Mex.

28. 1879. Type: MEXICO: Without precise

locality, Jurgensen 355 (Lectotype K, here

designated (photo PTBG); Isolectotypes G-

DEL, n.v. (fragment F), K (photo PTBG).

Syntypes: MEXICO: Oaxaca, Joya, F.

Liebmann 202 (K (photo PTBG)). MEXICO:

Without precise locality, Jurgensen 376 (K

(photo PTBG)). Arachnothryx. The K sheet of

Jurgensen 355 is very floriferous and

represents the best choice for lectotype. Note:

Rondeletia capitellata and R. liebmannii

Standi, are synonymous. Material from

Veracruz and Hidalgo erroneously identified

as R. capitellata in many herbaria actually

represents a distinct species, R. pringlei

Lorence.

Rondeletia chaconii Lorence in W. C. Burger

8l C. M. Taylor, Fieldiana, Bot., n.s. 33: 296.

1993. Type: COSTA RICA: Heredia,

Magsasay, pasturas, bosque socolado, fmca de

Bernardo Gomez, 700 m, 1 Jan 1983, /. C.

Chacon 177 (Holotype CR 92850; Isotype

MO).

Rondeletia chiapasensis Standi., Publ. Field

Mus. Nat. Hist., Bot. Ser. 22: 54. 1940. Type:

MEXICO: Chiapas, Cascarada, 31 Dec 1936,

E. Matuda 1064 (Holotype MICH, n.v.

(fragment F); Isotype A, n.v.). Rogiera.

—Gonzalagunia chiapasensis (Standi.)

Standi. & Steyerm.

Rondeletia chinajensis Standi. «fe Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254.

1947. Type: GUATEMALA: Alta Verapaz,

Cerro Chinaja, between Finca Yalpemech and

Chinaja, above source of Rio San Diego,

150-700 m, 1-2 Apr 1942, J. A. Steyermark

45637 (Holotype F 1134210 (photos MEXU,

PTBG)). Arachnothryx.

Rondeletia chiriquiana Lorence, Novon 4:

\19,fig. 7. 1994. Type: PANAMA: Chiriqui,

Fortuna Dam area, along shores of Rio

Chiriqui where it enters reservoir, 1000 m,

8°45'N, 82°13'W, 2 Aug 1984, H. W.

Churchill 5924 (Holotype MO 3595277

(photo PTBG)).

Rondeletia cooperi Standi., Publ. Field

Columbian Mus., Bot. Ser. 4: 267. 1929.

Type; PANAMA: Bocas del Toro, Buena

Vista Camp, on Chiriqui Trail, 1250 ft.,

Jan-Mar 1928, G. P. Cooper 600 (Yale

12233) (Holotype F 579596 (photo PTBG);

Isotype NY, n.v.). Arachnothryx.

Rondeletia cordata Benth., PI. Hartweg. 85.

1841. Type: GUATEMALA: In praeruptis

prope Guatemala [Guatemala City], Jan 1841,

T. Hartweg 585 (Holotype K (photo PTBG);

Isotype G-DEL, n.v. (fragment F)). Rogiera.

Rondeletia cordovana Standi. & Steyerm., Publ.

Field Mus. Nat. Hist., Bot. Ser. 23: 26. 1943.

Type: GUATEMALA: Zacapa, in ravine

bordering Quebrada Alejandria, summit of

Sierra de las Minas, vicinity of Finca

Alejandra, 2500 m, 13 Oct 1939, J. A.

Steyermark 29906 (Holotype F 1042952

(photos MEXU, PTBG); Isotype US (photos

A Nomenclator of Mexican and Central American Rubiaceae

151

MEXU, PTBG)). Arachnothryx. —Rondeletia

brandegeeana Lorence

Rondeletia cornifolia Benth., PI. Hartweg. 39.

1840. Type: MEXICO: Michoacan, prope

Morelia, T. Hartweg 299 (Holotype K (photo

MEXU); Isotypes BM, G-DEL, n.v. (fragment

F), P (photos MEXU, MO, PTBG), US

(fragment, photos MEXU, PTBG), W (2

sheets, photos PTBG)). Choristes. —Deppea

cornifolia (Benth.) Benth.

Rondeletia costaricensis Standi., N. Amer. FI.

32: 61. 1918. Type: COSTA RICA: Buena

Vista, San Carlos, 900 m, Apr 1903, H. Pittier

16696 (Holotype US 764141 (photos MEXU,

PTBG); Isotypes B, presumably destroyed

(fragment F), US). Arachnothryx.

Rondeletia darcyi Dwyer, Ann. Missouri Bot.

Gard. 67: 467. 1980. Type: PANAMA: Colon,

road bank just W of Portobelo, 12 Apr 1970,

W. G. D ’Arcy 4079 (Holotype MO 2238438

(photo PTBG); Isotype MO (photo PTBG)).

Arachnothryx. —Rondeletia secunda Standi.

Rondeletia dariensis Standi., N. Amer. FI. 32:

53. 1918. Type: PANAMA: Darien, Boca de

Paurando, on Sambii River, southern Darien,

20 m, Feb 1912, //. Pittier 5684 (Holotype US

867136 (photos F, MEXU, PTBG, US);

Isotypes F (photo PTBG), US (2 sheets)).

Arachnothryx.

Rondeletia deamii (Dorm. Sm.) Standi., N.

Amer. FI. 32: 60. 1918. Bouvardia deamii

Donn. Sm., Bot. Gaz. (Crawfordsville) 49:

455. 1910. Type: GUATEMALA: Guatemala,

at base of Rillo de las Nabajas, about y4 mile

front side of Fiscal, 3700 ft., 5 Jun 1909, C. C.

Deam 6190 (Holotype US 579579 (photos

MEXU, PTBG); Isotypes A, n.v., F (photo

PTBG), GH, n.v., MO (photos MEXU,

PTBG), US (photos MEXU, PTBG)).

Arachnothryx.

Rondeletia dubia Hemsl., Diag. PI. Mex. 28.

1879. Based on Bouvardia discolor Hook. &

Am., Bot. Beechy Voy. 428. 1840. Not

Rondeletia discolor Kunth in Humb., Bonpl.

& Kunth, 1820. Type: MEXICO: Oaxaca,

Inter Tehuantepec et Voca [Boca] del Monte,

in dit. Oaxaca, maxime [?] australi. Sept., G.

Andrieux 334 (Lectotype K, here designated

(photo PTBG); Isolectotypes G-DEL, n.v.

(fragment F), P, (photos MEXU, PTBG), W,

n.v. (photos F, MO)). Syntype: MEXICO:

Sierra Madre, N. W. of Mexico, Seemann

2057 (K (photo PTBG)). The K sheet of

Andrieux 334 is richly flowering and has

duplicates in three other herbaria.

—Rondeletia leucophylla Kunth in Humb.,

Bonpl. & Kunth

Rondeletia dukei Dwyer & Hayden, Ann.

Missouri Bot. Gard. 54: 144. 1967. Type:

PANAMA: Darien, Rio Pirre, 24 May 1966,

J. A. Duke 8329 (Holotype MO, unnumbered

sheet (photo PTBG)).—Wittmackanthus

stanleyanus (R. H. Schomb.) Kuntze

Rondeletia dwyeri Lorence, Novon 1: 139.

1991. Based on Chomelia leucophylla Dwyer,

Ann. Missouri Bot. Gard. 67: 9%. jig. 21.

1980. Not Rondeletia leucophylla Kunth in

Humb., Bonpl. & Kunth, 1820. Type:

PANAMA: Darien, slopes of Cerro Chucula,

drainage of Rio Pavarando, 11 Feb 1972, A.

Gentry 4262 (Holotype MO 2097508; Isotype

MO 2161959).

Rondeletia edwardsii Standi., Trop. Woods 37:

31. 1934. Type: HONDURAS: Comayagua,

Pito Solo, Lake Yojoa, 2000 ft., 29 Aug 1932,

J. B. Edwards P476 (Holotype F 662645

(photos MEXU, PTBG); Isotypes A, n.v.,

US). Arachnothryx.

Rondeletia eherenbergii K. Schum. ex Standi.,

N. Amer. FI. 32: 50. 1918. Type:

GUATEMALA: Totonocapan, Ehrenberg

1033 (Holotype US 594426 (photos F,

MEXU, PTBG); Isotype B, presumably

destroyed (photos F, MO, US)). Rogiera.

—Rondeletia gratissima Planch. «& Linden

Rondeletia elongata Bartl. ex DC., Prodr. 4:

409. 1830. Type: MEXICO: Guerrero, ad

Acapulco, Haenke s.n. (Holotype presumably

PR, n.v.; Isotype W, n.v. (photos F, MO)).

Arachnothryx .—leucophylla

Kunth in Humb., Bonpl. & Kunth

Rondeletia evansii Lorence, Novon 7: 52, fig.

4. 1997. Type: HONDURAS: Dept. Lempira,

Parque Nacional de Celaque, sendero entre la

planta electrica y la primera casa abandonada

de Don Tomas, 14°36'N, 88°40'W, 1800 m.

presumably destroyed (photos F, MO, US), G-

PTBG),’ W, n.v. (photo F)). Arachnothryx.

Rondeletia latifolia Oerst., Vidensk Meddel.

5t. Foren. Kjobenhavn 1852:

43. 1853. Type; NICARAGUA: In monte

154

Lorence

Rondeletia liebmannii Standi, J. Wash. Acad.

Sci. 17: 336. 1927. Type: MEXICO: Oaxaca,

without precise locality, 1841-43, F.

Liebmann 11834 (Holotype US 1315231

(photos MEXU, PTBG); Isotypes C, n.v.

(photo F), P (photos MEXU, PTBG)).

Arachnothryx. —Rondeletia capitellata

Hemsl.

Rondeletia ligustroides Hemsl, Diag. PI. Nov.

Mexic. 26. 1879. Type: MEXICO: Veracruz,

Orizaba, Botteri 971 (Lectotype K, here

designated (photo PTBG); Isolectotypes P (2

sheets, photos MEXU, PTBG). Syntypes:

MEXICO: Veracruz, Orizaba, Bourgeau 2518

(K (photo PTBG), P (3 sheets; photos MEXU,

PTBG)). Region d’Orizaba, Bourgeau 2622

(K (photo PTBG), B (n.v.; fragment F), F, K

(photo PTBG), P (4 sheets; photos MEXU,

PTBG), US (2 sheets)). Orizaba, Botteri 474

(K (2 sheets [label on first states “coll. Salle"-,

second states “Sumichrast coll.”] photos

PTBG)), P (photos MEXU, PTBG). Orizaba,

Botteri 1951 (K, n.v., P (2 sheets; photos

MEXU, PTBG). Orizaba, Muller 969 (K, n.v.,

P (photos MEXU, PTBG). Orizaba, Muller

1308 (K, n.v.). Botteri 971 at K is selected as

lectotype because it has three leafy twigs with

an abundance of flowers at anthesis, and three

duplicates exist at P. Rogiera.

Rondeletia linguiformis Hemsl, Diag. PI. Nov.

Mexic. 29. 1879. Type: GUATEMALA:

Without precise locality, Skinner s.n. in 1857

(Holotype K (photo PTBG)). Arachnothryx.

Rondeletia longipetiolata Lundell, Wrightia 5:

325. 1976. Type: MEXICO: Chiapas, Mt.

Ovando, near Escuintla, 14 Nov 1945, E.

Matuda 6146 (Holotype TEX-LL; Isotypes

MEXU, TEX-LL). Arachnothryx.

—Rondeletia buddleioides Benth.

Rondeletia lundelliana Standi, Contr. Univ.

Michigan Herb. 4: 31. 1940. Type: BRITISH

HONDURAS (BELIZE): El Cayo District

Vaca, 21 Apr 1938, P. H. Gentle 2504

(Holotype F 941520 (photos MEXU, PTBG)-

Isotypes A, n.v., K (photo PTBG), TEX-LL ’

ex MICH (photos MEXU, PTBG), US (2

sheets; photos MEXU, PTBG)).

Arachnothryx.~KonAe\ei\2i stenosiphon

Hemsl.

Rondeletia macdougallii Lorence, Novon 1:

144, 3A, B. 1991. Type: MEXICO:

Oaxaca, Cerro Azul (top), N of Niltepec, 7000

ft., 7 Mar 1956, T. MacDougalls.n. (Holotype

MEXU 253131 (photo PTBG)).

Rondeletia macrocalyx Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254.

1947. Type: GUATEMALA: Izabal, Cerro

San Gil, 300-900 m, 25 Dec 1941, y. T.

Steyermark 41864 (Holotype F 1134224

(photos MEXU, PTBG); Isotype F (photos

MEXU, PTBG), US). Arachnothryx.

Rondeletia manantlanensis Lorence, Novon 1:

U5,fig. 4A. B. 1991. Type: MEXICO:

Jalisco, Mun. de Autlan, Canada del Laurelito,

Las Joyas, 1850 m, 7 Dec 1985, A. Vazquez

3750 (Holotype WIS (photo PTBG); Isotype

IBUG, n.v.).

Rondeletia megalantha Lorence, Novon 4:

\32,fig. 8. 1994. Type: HONDURAS:

Cuyamel, 23 Aug 1924, M. A. Carleton 653

(Holotype US 1208409; Isotype US

1208410).

Rondeletia mexiae Standi, Publ. Field Mus.

Nat. Hist., Bol Ser. 22: 55. 1940. Type:

MEXICO: Oaxaca, Distr. Choapan, Santa

Maria, Montana Santa Maria, 1500 m, 1 Apr

1938, Y. Mexia 9261 (Holotype F 982856

(photos MEXU, PTBG); Isotypes GH, n.v.,

MO (photos MEXU, PTBG), UC (photo

PTBG), US (photos MEXU, PTBG)).

Arachnothryx. —Rondeletia scabra Hemsl.

Rondeletia mexicana (Turcz.) Standi, Publ.

Field Mus. Nat. Hist., Bot. Ser. 22: 388. 1940.

Siphonandra mexicana Turcz., Bull. Soc. Imp.

Naturalistes Moscou 21: 581. 1848. Type:

MEXICO: Oaxaca, without precise locality,

3000 ft., Oct, H. Galeotti 2664bis (Holotype

G-DEL, n.v. (photos F, MO, US); Isotype P

(photos MEXU, PTBG; fragment F, photos

MEXU, PTBG)).

Rondeletia microdon DC., Prodr. 4: 408. 1830.

Type: CUBA: Havana, de la Ossa s.n.

(Holotype G-DC, n.v. (microfiche IDC

800.691 :III.7,8)).—Psychotria microdon

(DC.) Urb.

156

Lorence

Rondeletia pittieri K. Schum. & K. Krause ex K.

Krause, Bot. Jahrb. Syst. 40: 316. 1908. Type:

COSTA RICA: Bords du Rio Poros, 2000 m,

10 Jan 1890, H. Pittier & T. Durand 1729

(Holotype B, presumably destroyed (photos F,

MO); Isotype US (photos MEXU, PTBG)).

Rogiera .—Rondeletia amoena (Planch.)

Hemsl.

Rondeletia platysepala Standi., Ann. Missouri

Bot. Card. 27: 343. 1940. Type: PANAMA:

Code, El Valle, North Rim, 600-1000 m, 14

May 1939, P. H. Allen 1791 (Holotype F

989484 (photo PTBG); Isotypes F (photos

MEXU, PTBG), G, n.v., GH, n.v., MO

(photos MEXU, PTBG), NY, n.v.).

Arachnothryx .—Rondeletia bertieroides

Standi.

Rondeletia polycephala Standi, J. Wash. Acad.

Sci. 17: 337. 1927. Type: MEXICO: Oaxaca,

vk'ithout precise locality, F. Liebmann 11826

(Holotype C, n.v. (fragment US; photos

MEXU, PTBG)). Arachnothryx .—Rondeletia

mexicana (Turcz.) Standi.

Rondeletia povedae Lorence in W. C. Burger

& C. M. Taylor, Fieldiana Bot., n.s. 33: 299.

1993. Type: COSTA RICA: San Jose, Santa

Rosa de Puriscal, 17 Jan 1973, L. J. Poveda

441 (Holotype CR 51260; Isotype MO

2211249).

Rondeletia pringlei Lorence, Novon 7: 54,

5. 1997. Type: MEXICO: Veracruz, hills near

Orizaba, 5000 ft., 9 Feb 1895, C. G. Pringle

6135 (Holotype GH; Isotypes K, MEXU, MO,

WU).

Rondeletia purpurea Lorence, Novon 1: 148,

fig. 6A-F, 7A, B. 1991. Type: MEXICO:

Oaxaca, Distr. de Ixtlan, Sierra de Juarez,

Ruta 175 Tuxtepec a Oaxaca, a 5 km al N. de

Vista Hermosa, 1175 m, 14 Apr 1982, D. H.

Lorence cfe R. Torres C. 4009 (Holotype

MEXU (photo PTBG); Isotypes BM, F, MO,

PTBG).

Rondeletia pyramidalis Lundell, Wrightia 5:

327. 1976. Type: MEXICO: Chiapas, San

Luis, near Siltepec, 1300 m, 24 Jan 1945, E.

Matuda 5292 (Holotype TEX-LL (photos

MEXU, PTBG); Isotypes F, MEXU, TEX-LL,

n.v.). Arachnothryx.

Rondeletia rekoi Standi, Proc. Biol. Soc.

Wash. 8: 126. 1918. Type: MEXICO: Oaxaca,

Las Pilas (Cerro Espino), 400 m, 10 Oct 1917,

B. P. Reko 3490 (Holotype US 867145

(photos MEXU, PTBG)). Arachnothryx.

Rondeletia repens L., Syst., ed. 10, 2: 928.

1759. Type: JAMAICA: Shanes.n.

(Lectotype: Sloane herb, iv.l 11, n.v.;

designated by Howard, 1989).—Geophila

repens (L.) I. M. Johnst.

Rondeletia ricoi Lorence, Novon 1: 152, fig.

8E, F. 9A, B. 1991. Type: MEXICO: Oaxaca,

Distr. de Putla de Guerrero, a 10 km al NE de

Putla, 950 m, 9 Dec 1982, L Rico A., E. M.

Martinez & O. Tellez V. 566 (Holotype

MEXU (photo PTBG); Isotypes BM, F, MO,

PTBG).

Rondeletia roezlii (Planch.) Hemsl, Diag. Pi.

Nov. Mexic. 25. 1879. Rogiera roezlii

Planch., FI. Serr. Jard. 5: tab. 442. 1849.

Type: Based on cultivated material grown in

Belgium, probably of Guatemalan origin (type

not designated).—Rondeletia amoena

(Planch.) Hemsl.

Rondeletia rothschuhii Loes., Engl. Bot. Jahrb.

60: 369. 1926. Type: NICARAGUA:

Matagalpa, Canada Yucul, 1000 m, Rothschuh

592 (B, presumably destroyed (photos F, MO;

fragment F)).—Rondeletia buddleioides

Benth.

Rondeletia rubens L. O. Williams, Phytologia

26: 128. 1973. Type: GUATEMALA: Quiche,

Nebaj, 6000 ft, 19 Nov 1934, A. F. Skutch

1725 (Holotype F 815804 (photos MEXU,

PTBG)). Arachnothryx.

Rondeletia rufescens B. L. Rob., Proc. Amer.

Acad. Arts 45: 402. 1910. Type:

GUATEMALA: Alta Verapaz, Coban, 4800

ft. Mar 1881, //. von Tuerckheim 582

(Holotype GH, n.v.; Isotypes F, K (photo

PTBG), P (photos MEXU, PTBG), US (2

sheets; photos MEXU, PTBG)).

Arachnothryx .—Rondeletia gracilis Hemsl.

Rondeletia rufescens B. L. Rob. var. ovata B. L.

Rob., Proc. Amer. Acad. Arts 45: 403. 1910.

Type: GUATEMALA: Alta Verapaz, Tactic,

1800 m. Mar 1903, H. von Tuerckheim 8401

(Holotype GH, n.v.; Isotypes F (2 sheets), US

A Nomenclator of Mexican and Central American Rubiaceae

157

(photos MEXU, PTBG)).—Rondeletia

gracilis Hemsl.

Rondeletia rugosa Benth. in Oerst., Vidensk.

Meddel. Dansk Naturhist. Foren. Kjobenhavn

1852: 43. 1853. Type: COSTA RICA:

Cartago, 5600 ft., Oersted s.n. [^Plantae

centroamericanae Oersted 11828] (Holotype

C, n.v.; Isotype US (photo PTBG)).

—Rondeletia amoena (Planch.) Hemsl.

Rondeletia rzedowskii Lorence, Novon 1: 154,

fig. 10. 1991. Type: MEXICO: Guerrero,

Mun. de Atoyac, Puerto de la Piedra

Acanalada, 1930 m, 6 Jan 1984, Laboratorio

de Biogeografia 867 (Holotype MEXU

364217 (photo PTBG); Isotypes FCME,

MEXU (photo PTBG)).

Rondeletia salicifolia Dwyer & Hayden,

Phytologia 15: 58. 1967. Type: PANAMA:

Panama, Cerro Jefe in Clusia forest at

2700-3000 ft., 27 Jan 1966, E. L Tyson, J. D.

Dwyer & K. Blum 3319 (Holotype MO

1824503 (photo PTBG); Isotypes F, US

(photos MEXU, MO)). Arachnothryx.

—Rondeletia bertieroides Standi.

Rondeletia salicifolia Dwyer & Hayden subsp.

brevicorolla J. H. Kirkbr., Ann. Missouri Bot.

Gard. 55; 383. 1968. Type: PANAMA: Bocas

del Toro, cloud forest above Quebrada Huron

on Cerro Bonyik, 166-400 m, 13 Apr 1968,

J. H. Kirkbride, Jr. &J. A. Duke 597

(Holotype MO 1946832 (photo PTBG);

Isotypes NY, n.v., US (2 sheets; photos

MEXU, PTBG)). Arachnothryx.—

bertieroides Standi.

Rondeletia scabra Hemsl., Diag. PI. Nov.

Mexic. 29. 1879. Type: MEXICO: Oaxaca,

Sierra San Pedro Nolasco, Talea etc., C.

Jurgensen 926 (Holotype K (photo PTBG)).

Rondeletia schumanniana K. Krause, Bot.

Jahrb. 40: 315. 1908. Type: PANAMA:

Veraguas, Mar 1906,7. von Warszewicz 1168

(B, presumably destroyed (photo F; fragment

F))-—Rondeletia amoena (Planch.) Hemsl.

Rondeletia scoti Lorence, Novon 1: 155,//g.

8A. B. 1991. Type: MEXICO: Oaxaca, La

Glorieta-Rio Grande (N slope), 12 Mar 1953,

T. MacDougall s.n. (Holotype MEXU (photo

PTBG); Isotypes MO, PTBG).

Rondeletia secunda Standi., Contr. U.S. Natl.

Herb. 18: 141. 1916. Type: PANAMA: San

Bias, forests around Puerto Obaldia, San Bias

Coast, 0-50 m, Aug 1911, H. Pittier 4279

(Holotype US 679392 (photos F, MEXU,

PTBG); Isotype NY, n.v.). Arachnothryx.

Rondeletia secundiflora B. L. Rob., Proc.

Amer. Acad. Arts 45: 403. 1910. Type:

GUATEMALA: In woods along road from

Patin to Escuintla, 21 Jul 1860, S. Hayes s.n.

(Holotype GH, n.v.; Isotype K (photo

PTBG)). Arachnothryx.

Rondeletia seleriana Loes., Verb. Bot. Ver.

Brandenb. 65: 105. 1923. Type:

GUATEMALA: Huehuetenango, Nenton

Distr., near Chacula, E. Seler & C. Seler 3015

(Holotype B, presumably destroyed (photos F,

MO, US), fragment F). Rogiera. —Rondeletia

gratissima (Planch. & Lindl.) Hemsl.

Rondeletia septicidalis B. L. Rob., Proc. Amer.

Acad. Arts 45: 403. 1910. Type: MEXICO:

Chiapas, Chicharras, 3000-6000 ft., 6 Feb

1896, E. W. Nelson 3755 (Holotype US

256722 (photos MEXU, PTBG); Isotype GH,

n.v.). Arachnothryx.

Rondeletia silvicola L. O. Williams, Phytologia

26: 128. 1973. Type: HONDURAS: Atlantida,

Lancetilla Valley, near Tela, 20-600 m, 6 Dec

1927-20 Mar 1928, P. C Standley 56885

(Holotype F 581955 (photos MEXU, PTBG)).

Arachnothryx.

Rondeletia skutchii Standi. & Steyerm., Publ.

Field Mus. Nat. Hist., Bot. Ser. 22: 289. 1940.

Type: GUATEMALA: Suchitepequez, Finca

Moca, 4200 ft., 10 Jan 1935, A. F. Skutch

2110 (Holotype F 934088 (photos MEXU,

PTBG); Isotypes A, n.v., US (photos MEXU,

PTBG)). Arachnothryx.

Rondeletia spinosa K. Schum. ex Loes., Bull.

Herb. Boissier3: 620. 1895. Type: MEXICO:

Hidalgo, prope Huejutla, Apr, E. Seler & C.

Seler 668 (Holotype B, presumably

destroyed).—Status uncertain, possibly

Randia sp. The protologue diagnosis states

branches armed with paired spines and

flowers solitary or fascicled.

Rondeletia stachyoidea Donn. Sm., Bot. Gaz.

(Crawfordsville) 42: 298. 1906. Type:

158

LORENCE

GUATEMALA: Alta Verapaz, Semococh,

600-900 m, 25 Apr 1905, R. Hay s.n.

(Holotype US 473726 (photos MEXU,

PTBG)). Arachnothryx.

Rondeletia standleyana A. R. Molina, Ceiba 1:

262. 1951. Type: HONDURAS: Morazan, in

pine forest above Zambrano, 1200 m, 20 Jul

1949, L O. Williams & A. Molina R. 14417

(Holotype US 2216018 (photos MEXU,

PTBG); Isotypes F (photos MEXU, PTBG,

US), US). Arachnothryx.

Rondeletia stenosiphon Hemsl., Diag. PI. Nov.

Mexic. 26. 1879. Type: MEXICO: Without

precise locality, Yucatan & Tabasco [sic],

E. P. Johnson s.n. (Holotype K (photo

PTBG)). Rogiera.

Rondeletia stenostachya Standi., Publ. Field

Mus. Nat. Hist., Bot. Ser. 18: 1372. 1938.

Type: COSTA RICA: La Hondura de San

Jose, 1300 m, 15 Aug 1933, M. Valerio 769

(Holotype F 681816 (photos MEXU, PTBG)).

Arachnothryx. —Gonzalagunia stenostachya

(Standi.) W. C. Burger

Rondeletia strigosa (Benth.) Hemsl., Diag. PI.

Nov. Mexic. 27. 1879. Bouvardia strigosa

Benth., PI. Hartweg. 75. 1841. Type:

GUATEMALA: [Volcan] Zunil, Nov, T.

Hartweg 530 (Lectotype K, here designated

(photo PTBG); Isolectotypes G-DEL, n.v.

(fragment F), P (photos F, MEXU, MO,

PTBG), W, n.v. (photos F, MO)). Syntype:

GUATEMALA: Volcan de Fuego,

6000-6400 ft., Salvin s.n. (K (photo PTBG;

fragment F)). Rogiera. The K collection of

Hartweg 530 is a good, representative choice

for lectotype and in addition has duplicates in

three herbaria.

Rondeletia subscandens Lundell, Wrightia 5:

328. 1976. Type: GUATEMALA: Peten, La

Cumbre, about 500 m west of km 142/143 of

the Cadenas Road, 10 Sep 1976, C. L. Lundell

& E. Contreras 19830 (Holotype TEX-LL

(photos MEXU, PTBG); Isotypes F (2 sheets;

photos MO), K (photo PTBG)). Rogiera.

Rondeletia suffrutescens Brandegee, Univ.

Calif. Publ. Bot. 6: 70. 1914. Type: MEXICO:

Chiapas, Cerro del Boqueron, Aug 1913, C. A.

Purpus 6704 (Holotype UC 172451, n.v.;

Isotypes F (photos MEXU, PTBG), MO, US

(photos MEXU, PTBG)). Rogiera.

Rondeletia tacanensis Lundell, Bull. Torrey

Bot. Club 66: 603. 1939. Type: MEXICO:

Chiapas, Volcan Tacana West, 2800 m, 30

Mar 1939, E. Matuda 2928 (Holotype TEX-

LL (photos MEXU, PTBG); Isotypes A, n.v.,

F (photos MEXU, PTBG), GH, n.v., K (photo

PTBG), MEXU, US (2 sheets; photos MEXU,

PTBG)). Arachnothryx.

Rondeletia taylorae Lorence in W. C. Burger

& C. M. Taylor, Fieldiana, Bot., n.s. 33: 300.

1993. Type: COSTA RICA: Limon, Canon de

Limon, El Progreso. Fila Matama, Valle de la

Estrella, 9°47T8"N, 83°08'45"W, 1350 m, 19

Apr 1989, G. Herrera & A. Chacon 2656

(Holotype CR; Isotypes MO, PTBG).

Rondeletia tenorioi Lorence, Novon 1: 156,y2g.

11 A, fi. 1991. Type: MEXICO: Oaxaca, Distr.

de Juxtlahuaca, 19 km al SW de Juxtlahuaca y

a 5 km en la misma direccion de Sta. Ma.

Asuncion, 1900 m, 4 Apr 1982, P. Tenorio &

R. Torres C. 184 (Holotype MEXU (photo

PTBG); Isotypes BM, F, MO).

Rondeletia thiemei Donn. Sm., Bot. Gaz.

(Crawfordsville) 42: 299. 1906. Type:

HONDURAS: Santa Barbara, Rio

Chamelecon, 400 m, 16 Dec 1888, C. Thieme

5276 (Holotype US 941403 (photos MEXU,

PTBG)). Arachnothryx.

Rondeletia torresii Standi., J. Wash. Acad. Sci.

18: 163. 1928. Type: COSTA RICA: Alajuela,

Viento Fresco, 1600-1900 m, 13 Feb 1926,

P. C. Standley & R. Torres R. 47839

(Holotype US 1305295 (photos MEXU,

PTBG)). Arachnothryx.

Rondeletia tuxtlensis Lorence & Cast.-

Campos, Biotica 13: \4^,fig. 1. 1988. Type:

MEXICO: Veracruz, Mun. Mecayapan,

Volcan San Martin Pajapan al sur del Ejido La

Valentina, 700 m, 8 May 1985, J. /. Calzada

11183 (Holotype XAL; Isotypes F, MEXU,

MO (photo PTBG)). Arachnothryx.

Rondeletia urophylla Standi. & L. O.

Williams, Phytologia 26: 129. 1973. Type:

COSTA RICA: Puntarenas, Esquinas forest,

30 m, 3 Aug 1951, P. H. Allen 6291

A Nomenclator of Mexican and Central American Rubiaceae

159

(Holotype F 1562934 (photos MEXU,

PTBG); Isotype EAP, n.v.).

Rondeletia uxpanapensis Lorence & Cast.-

Campos, Biotica 13; 150, 2. 1988. Type:

MEXICO: Veracruz, Mun. Hidalgotitlan, zona

de Uxpanapa; Rio Solosuchil a 5-6 km ESE

de Hermanos Cedillo, 100-200 m, 24 Mar

1982, D. H. Lorence et al 3909 (Holotype

MEXU (photo PTBG); Isotypes BM, ENCB,

CHAPA, MO, NY, PTBG, UC, XAL).

Arachnothryx.

Rondeletia versicolor J. E. Sm., Bot. Mag. 7;

tab. 4579. 1851. Type; Based on cultivated

material grown at Kew, sent by Seemann from

Boquete, Veraguas, Panama (type not

designated). Rogiera .—Rondeletia amoena

(Planch.) Hemsl.

Rondeletia villosa Hemsl., Diag. PI. Nov.

Mexic. 27. 1879. Type: MEXICO: Oaxaca,

San Pedro Nolasco, etc., Jurgensen 806

(Lectotype K, here designated (photo PTBG);

Isolectotypes G-DEL, n.v. (fragment F), K

(photo PTBG), P (photos MEXU, PTBG)).

Syntypes: MEXICO; Oaxaca, Trapiche de

Concepcion, F. Liebmann 70 (K (photo

PTBG), G-DEL, n.v. (fragment F)). Tabasco,

arbuste a fleurs blanches en novembre et

Xbre., environs de Teapa (Tabasco), 1839,

Linden 145 (K (photo PTBG); probable

syntypes G-DEL, n.v. (fragment F), P (photos

MEXU, PTBG)). Arachnothryx. The K sheet

oi Jurgensen 806 is designated as lectotype,

because it is representative and has flowers

and fruits, with duplicates in three other

herbaria.

Rondeletia volubilis Sesse & Mo?., PI. Nov.

Hisp. 36. 1887. Type: MEXICO: Habit in

Acahuizotla, Sesse & Mogiho 1393 (Holotype

MA, n.v. (photo WU); Isotype F, fragment,

n.v.). Caique of FI. Mexic. Ic. 275 (G-DC,

n.v. (photo MO)). For typification, see Puff

(1991: 326).—Paederia ciiiata (Bartl. ex

DC.) Standi.

Rondeletia vulcanicola Standi. & Steyerm.,

Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 291.

1940. Type. GUATEMALA; Quezaltenango,

Finca Pireneos, lower south-facing slopes of

Volcan Santa Maria, between Santa Maria de

Jesus and Calahuache, 1300-1500 m, 31 Dec

1939, J. A. Steyermark 33220 (Holotype F

1041786 (photos MEXU, PTBG)).

Arachnothryx .—Rondeletia secundiflora

B. L. Rob.

Rondeletia wendtii Lorence & Cast.-Campos,

Biotica 13: \5A,fig. 3. 1988. Type: MEXICO:

Oaxaca, Distr. Juchitan, Mun. de Santa Maria

Chimalapa, Arroyo Chocolin, Poblacion de

Nicolas Bravo, cerca del Rancho de Agustin

Montero, 3^ km al sur de la Poblacion de

Rio Alegre (Veracruz), 150 m, 21 Oct 1983,

T. Wendt, A. Montero & 1. Almaraz 4244

(Holotype MEXU (photo PTBG); Isotypes

CHAPA, PTBG). Arachnothryx.

Rondeletia yucatanensis Lundell, Wrightia 5:

329. 1976. Type; MEXICO: Yucatan,

Chichen Itza, 28 Apr 1950, J. T. Baldwin. Jr.

14230 (Holotype TEX-LL). Arachnothryx.

Note: the type was likely collected from a

cultivated plant outside its normal range, as

this species is commonly cultivated in many

parts of Mexico for its nocmmally fragrant

flowers._Rondeletia leucophylla Kunth in

londeletia zolleriana Standi. & Steyerm Publ

Field Mus. Nat. Hist., Bot. Ser. 22: 292. 1940.

Type. GUATEMALA: San Marcos, between

Finca El Porvenir and Loma Corona, 9 mi.

NW of El Porvenir, southwest-facing slopes

of Volcan Tajumulco, 1300-2000 m, 14 Mar

1940, J. A. Steyermark 37738 (Holotype F

1039668 (photos MEXU, PTBG)).

.. - -I.__Rnnd(>letia brandeeeeana

Lorence

Rosenbergiodendron Fagerl., Svcnsk Bot.

Tidskrift. 42: 150. 1948. Type:

Rosenbergiodendron formosum (Jacq.)

^^rgiodendron

;erl., Svensk Bot. Tidsknft. 42; 152. 1948.

ssaenda formosa Jacq., Enum. PI- Canb.

1760 . Type: COLOMBIA; Bolivar,

L Nomenclator of Mexican and Central American Rubiaceae

161

Rudgea laevis C. M. Taylor, Novon 5: 101, fig.

IF, G. 1995. Type: COSTA RICA;

Puntarenas, Monteverde, Bajos de La Lindora,

Rio Guacimal, 10°18’N, 84°48'W, 19 Feb

1988, W. Haber &E. Bello 8271 (Holotype

CR, n.v.; Isotype MO no 3646480).

Rudgea magna Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 22: 56. 1940. Type;

PANAMA: San Bias Coast, hills of Sperdi,

near Puerto Obaldia, 20-200 m, Sep 1911, //.

Pittier 4350 (Holotype F 600258 (photo

PTBG)).

Rudgea mandevilliifolia Dwyer ex C. M.

Taylor, Novon 6: 304,fiig. 3C, D. 1996. Type:

PANAMA: Chiriqui, vicinity of Planes de

Homito beyond Gualaca, ca. 23 km E of Finca

Linares, 1400-1900 m, 28 Nov 1979, T. B.

Croat 48863 (Holotype MO 3615290).

Rudgea mcphersonii Dwyer ex C. M. Taylor,

Novon 6: 304,yig. 3E, F. 1996. Type:

PANAMA: Colon, Santa Rita ridge, SE of

Colon, 10-11 road-miles from trans-isthmian

highway, 9°25'N, 79°40'W, 450 m, 18 Sep

1987, G. McPherson 11771 (Holotype PMA,

n.v.; Isotype MO 3646497).

Rudgea monofructus Gomez-Laur. & Dwyer,

Novon 1: 50, fig. 1. 1991. Type: COSTA

RICA; Alajuela, Reserva Forestal de San

Ramon, Campamento Rio San Lorencito,

900-1100 m, 25 Sep 1989, J. Gomez-Laurito

& G. Vargas 11822 (Holotype UCR, n.v.;

Isotype MO).

Rudgea panamensis (Dwyer) C. M. Taylor,

Novon 6: 305. 1996. CephaHis panamensis

Dwyer, Ann. Missouri Bot. Gard. 67: 77.

1980. Type: PANAMA: Colon, Rio Guanche,

ca. 2.5 km upriver from bridge on road to

Portobelo, 10-100 m, 14 Dec 1974, S. Mori &

J. Kallunki 3714 (Holotype MO 2353037

(photos MEXU, PTBG); Isotypes MO

2353038 (photos MEXU, PTBG), PMA

22701, n.v.).

Rudgea pittieri Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 22: 211. 1940. Type:

PANAMA: Colon, Loma de la Gloria, near

Fato (Nombre de Dios), 10-100 m, Jul-Aug

1911,//. Pittier 3855 (Holotype F 636043

(photo PTBG); Isotypes F, US).

Rudgea raveniana W. C. Burger in W. C.

Burger & C. M. Taylor, Fieldiana, Bot., n.s.

33: 304. 1993. Type: COSTA RICA: Rincon

de Osa, W. Burger & R. Liesner 7251

(Holotype CR, n.v.; Isotype F, n.v.).

Rudgea reducticalyx Dwyer in W. C. Burger &

C. M. Taylor, Fieldiana, Bot., n.s. 33: 304.

1993. Type: COSTA RICA: Alajuela, along

road between Canas and Upala, 4 km NNE of

Bijagua on slopes leading into Rio Zapote,

400 m, 24 Jun 1976, T. B. Croat 36330

(Holotype CR, n.v.; Isotype MO 2481680

(photo PTBG)).

Rudgea sanblasensis C. M. Taylor, Novon 6:

220. 1996. Type: PANAMA: San Bias, Rio

Playon Chico, Dukandi, vecinidad de la Isla

Playon Chico, 09°15'N, 78°15'W, 30 m, 6

Sep 1993, H. Herrera et al. 1358 (Holotype

MO 4671394; Isotype PMA, n.v.).

Rudgea simiarum Standi. & Steyerm., Publ.

Field Mus. Nat. Hist., Bot. Ser. 22: 389. 1940.

Type: GUATEMALA: Izabal, between

Virginia and Lago de Izabal, Montana del

Mico, 50-500 m, 5 Apr 1940, J. A.

Steyermark 38839 (Holotype F 1042936

(photo PTBG)).—Psychotria cf eurycarpa

Standi.

Rudgea skutchii Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 18: 1374. 1938. Type: COSTA

RICA: San Jose, vicinity of El General, 915

m, Aug 1936, A. F. Skutch 2836 (Holotype

US 1642550 (photos MEXU, PTBG);

Isotypes GH, n.v., MO (photos MEXU,

PTBG)).

Rudgea thyrsiflora Donn. Sm., Bot. Gaz.

(Crawfordsville) 61: 375. 1916. Type:

COSTA RICA: Talamanca, Forets de Tsaki,

200 m, Apr 1895, A. Tonduz 9579 (Holotype

US 943477 (photos MEXU, PTBG); Isotype

BR, n.v.).—Psychotria stenostachya Standi.

Rudgea trifurcata Gomez-Laur., Brenesia 33:

139. 1990 (1991). Type: COSTA RICA:

Limon, Talamanca, 3 km al oeste de Volio,

9°37’40N, 82°53’50W, 100 m../. Gomez-

Laurito & G. Vargas 11957 (Holotype CR,

n. v.; Isotypes F, n.v., MO, US, n.v.))

162

LORENCE

Rustia Klotzsch in Hayne, Getr. Darstell. Gew.

14; sub tab. 14, 15. 1846. Lectotype: Rustia

formosa (Cham. & Schltdl.) Klotzsch

(=Exostema formosum Cham. & Schltdl).

Rustia costaricensis (Standi.) Lorence in W. C.

Burger & C. M. Taylor, Fieldiana, Bot., n.s.

33: 306. 1993. Stomandra costaricensis

Standi, Publ. Field Mus. Nat. Hist., Bot. Ser.

23; 247. 1947. Type; COSTA RICA; San

Jose, basin of El General, 675-900 m. Mar

1940, A. F. Skutch 4777 (Holotype F 1044344

(photo PTBG); Isotypes MO (photo PTBG),

US).

Rustia dressleri Delprete, Novon 5: 137. 1995.

Type: PANAMA: Panama, El Llano-Carti

highway, 8-10 km N of El Llano, 31 Aug

1974, R. L. Dressier 4703 (Holotype MO

2268783).

Rustia ferruginea Standi, Contr. U.S. Natl.

Herb. 18: 142. 1916. Type: PANAMA: Colon,

along Rio Fato, 10-100 m, Aug 1911,//.

Pittier 420J (Holotype US 679302 (photos

MEXU, PTBG)).

Rustia occidentalis (Benth.) Hemsl, Biol.

Centr.-Amer., Bot. 2; 14. 1881. Exostemma

occidentale Benth., Bot. Voy. Sulphur 104.

1844. Type: PANAMA: Without precise

locality, Barclay s.n. or Hinds s.n. (Holotype

K, n.v.).

Rustia panamensis Dwyer, Ann. Missouri Bot.

Gard. 67: 483. 1980. Type: PANAMA:

Veraguas, Valley of Rio Dos Bocas, 11 km

from Escuela Agricola Alto Piedra (above

Santa Fe), on road to Calovebora, 450 m, 30

Aug 1974, T. B. Croat 27508 {HoXotypcUO

2214602; Isotypes F (photo PTBG), GH,

n.v.).—Rustia costaricensis (Standi.)

Lorence

Sabicea Aubl, Hist. PI. Guiane 192. 1775. Type

Sabicea ambigua Standi, Publ. Field

Columbian Mus., Bot. Ser. 7; 49. 1930. Type:

COLOMBIA: Without precise locality, 1847,

Triana s.n. (Holotype K, n.v. (photo F)).

—Amphidasya ambigua (Standi.) Standi.

Sabicea costaricensis Wemham, Monogr.

Sabicea 31.1914. Syntypes: COSTA RICA:

Buenos Aires, buissons, 300 m, Jan 1893, H.

Pittier 6712 [on label “H. Pittier” is crossed

out and “Tonduz” is written in] (US 1393299

(photos MEXU, PTBG)). On the Savannah,

H. Pittier 4025 (BM, n.v.; Isosyntypes K, n.v.,

BR, n.v., G, n.v.).—Sabicea panamensis

Wemham

Sabicea flagenioides Wemham, Monogr.

Sabicea 57. 1914. Type; MEXICO; Yucatan,

Chichankanab, G. F. Gaumer 1432 (Holotype

F, n.v.; Isotype B, presumably destroyed).

—Status uncertain.

Sabicea hirsuta Kunth in Humb., Bonpl. &

Kunth var. adpressa Wemham, Monogr.

Sabicea 55. 1914. Syntypes: PANAMA:

Isthmus of Panama, Chagres, Jan-Mar 1850,

A. Fendler 180 (F, (photo PTBG), BM, n.v.,

W, n.v., fragment F); same locality, Hayes s.n.

(n.v.), same locality, Seemann 1073 (n.v.).

COSTA RICA: Limon, Kuntze 1995 (n.v.);

Punta Arena, Tonduz 6712 (n.v.), Tonduz

9955 (n.v.); six additional syntypes are cited

from St. Thomas, Trinidad, and Brazil.

Sabicea mexicana Wemham, Monogr. Sabicea

41. 1914. Syntypes; MEXICO; Oaxaca,

without precise locality, 3000 ft., H. Galeotti

2662 (K, n.v. (photo MO), B, n.v., BR, n.v.,

G, n.v., P, n.v., W, n.v.). Without precise

locality, F. Liebmann 43 (K, n.v., BR, n.v.).

Sabicea panamensis Wemham, Monogr.

Sabicea 30. 1914. Type: PANAMA: Chagres,

Isthmus of Panama, Mar 1850, A. Fendler 181

(Holotype K, n.v.; Isotypes US, 2 sheets

(photos MEXU, PTBG)).

Sabicea stellaris Dwyer, Ann. Missouri Bot.

Gard. 67: 489. 1980. Type: PANAMA:

Panama, Rio Pequeni, 10-15 minutes

upstream from hydrographic station by motor,

11 Dec 1974, R. L Dressier 4887 (Holotype

MO 2353169 (photo PTBG)).

Sabicea villosa Willd. ex Roem. & Schult,,

Syst. Veg. 5: 265. 1819. var. villosa. Type;

VENEZUELA: Orinoco River, A. Bonpland

s.n. (Holotype presumably B-WI, n.v.).

Sabicea villosa Willd. ex Roem. & Schult. var.

adpressa (Wemham) Standi, Publ. Field

Columbian Mus., Bot. Ser. 7: 52. 1930.

Syntypes; PANAMA: Isthmus of Panama,

A Nomenclator of Mexican and Central American Rubiaceae

163

Chagres, Jan-Mar 1850, A. Fendler 180 (F,

(photo PTBG), BM, n.v., W, n.v., fragment

F); same locality, Hayes s.n. (n.v.), same

locality, Seemann 1073 (n.v.). COSTA RICA:

Limon, Kuntze 1995 (n.v.); Punta Arena,

Tonduz 6712 (n.v.), Tonduz 9955 (n.v.); six

additional syntypes are cited from St. Thomas,

Trinidad, and Brazil.

Schradera Vahl, Eclog. Amer. 1: 35. 1796, nom.

cons. Type: Schradera capitata Vahl, nom.

illeg. i=Schradera involucrata (Sw.) K.

Schum.)

Schradera blumii Dwyer & Hayden,

Phytologia 15: 59. 1967. Type: PANAMA:

Panama, Cerro Jefe in Clusia forest at

2700-3000 ft., 27 Jan 1966, E. L Tyson, J.

Dwyer & K. Blum 3218 (Holotype MO

1831023 (photo PTBG)).

Schradera costaricensis Dwyer in W. C.

Burger & C. M. Taylor, Fieldiana, Bot., n.s.

33: 309. 1993. Type: COSTA RICA: Alajuela,

Bijagua, G. Herrera 2004 (Holotype CR, n.v.;

Isotypes F, n.v., MO).

Serissa Comm, ex Juss., Gen. 209. 1789. Type:

Serissa foetida Comm, ex Juss.

Serissa japonica (Thunb.) Thunb., Nov. Gen.

PI. 132, p/. 9. 1798. Type: JAPAN: Collector

unknown (presumably UPS, n.v.). Lycium

japonicum Thunb., Nov. Act. Roy. Soc.

Upsal. 3: 207. 1780. Cultivated at least in

Costa Rica and Panama.

Sherardia L., Sp. PI. 1: 102. 1753. Type:

Sherardia arvensis L.

Sherardia arvensis L., Sp. PI. 1: 102. 1753.

Type: Habitat in arvensis Scaniae, Germaniae,

Angliae, (Lectotype Sherard, Herb. Clifford

33: Sherardia 1 (BM, n.v.); designated by

Natali in Jarvis et al., 1993: 88). Naturalized

at least in Veracmz and Costa Rica.

Sickingia Willd., Ges. Naturf Freunde Berlin

Neue Schrift. 3: 445. 1801. Type not

designated. [Herein treated as a synonym of

Simira Axihl.]

Sickingia calderoniana Standi., Publ. Field Mus.

Nat. Hist., Bot. Ser. 22: 214. 1940. Type: EL

SALVADOR: Jayaque, finca del Dr. Zarate, 9

Jan 1939, M. L Van Severen 66 (Holotype F

1033327 (photo PTBG); Isotype F).—Simira

calderoniana (Standi.) Steyerm.

Sickingia klugei (Standi.) Standi., Trop. Woods

22: 10. 1930. Calderonia klugei Standi., J.

Wash. Acad. Sci. 15: 6. 1924. Type:

PANAMA: Panama, near Chepo, 1924, H. C.

Kluge 19 (Yale 7132) (Holotype US 1206424

(photos MEXU, PTBG); Isotype F (plus

another probable isotype at F, a fruiting

collection from type tree, Kluge s.n., also Yale

7132)).—Simira klugei (Standi.) Steyerm.

Sickingia lancifolia Lundell, Wrightia 4: 50.

1968. Type: GUATEMALA: Peten, Laguna

Petexbatun, in high forest bordering lake, on

Escarbadero, 2 Apr 1964, C L Lundell 18224

(Holotype TEX-LL, n.v.; Isotype US (photos

MEXU, PTBG)).—Simira salvadorensis

(Standi.) Steyerm.

Sickingia lancifolia Lundell var. puberula

Lundell, Wrightia 4: 50. 1968. Type:

GUATEMALA: Peten, Lacandon, in bajo,

about 1 km, 700 m south, 12 Mar 1962, E.

Contreras 3514 (Holotype TEX-LL, n.v.).

—Simira salavadorensis (Standi.) Steyerm.

Sickingia maxonii (Standi.) Standi., Trop.

Woods 14: 30. 1928. Genipa maxonii Standi.,

J. Wash. Acad. Sci. 8: 642. 1918. Type:

PANAMA: Panama, Canal Zone, forest along

the Rio Indio de Gahin, near sea level, 17 Feb

\9n, W.R. Maxon 4848 (Holotype US

675223 (photos MEXU, PTBG)).—Simira

maxonii (Standi.) Steyerm.

Sickingia mexicana Bullock in Hook. Icon., ser.

4, 4: tab. 3321. 1936. Type: MEXICO:

Mexico, Distr. Temascaltepec, Luvianos, 26

Sep 1933, G. B. Hinton 4818 (Holotype K,

n.v.; Isotype US (photos MEXU, PTBG)).

—Simira mexicana (Bullock) Steyerm.

Sickingia mollis Lundell, Contr. Univ. Michigan

Herb. 4: 32. 1940. Not Sickingia mollis

Standi., 1936. Type: MEXICO: Tabasco,

Reforma, on the Rio San Pedro de Martir,

above Balancan, 22-26 May 1939, E. Matuda

3161 (Holotype MICH, n.v.; Isotypes A, n.v..

166

Lorence

Sommera arborescens Schltdl., Linnaea 9: 602.

1835. Type; MEXICO: Veracruz, Hacienda de

la Laguna, Schiede 272 (Holotype presumably

HAL, n.v.; Isotypes F (photos MEXU,

PTBG), LE, n.v.(fragment F), MO (photos

MEXU, PTBG), US (photos MEXU, PTBG)).

Sommera chiapensis Brandegee, Univ. Calif.

Publ. Bot. 6: 196. 1915. Type: MEXICO:

Chiapas, Finca Irlanda, Jun 1914, C.

Purpus 7263 (Holotype UC 174914. n.v.;

Isotypes F (photos MEXU, PTBG), MO, 2

sheets (photos MEXU, PTBG), US (photos

MEXU, PTBG)).

Sommera donnell-smithii Standi., Contr. U.S.

Natl. Herb. 17: 436. 1914. Type: COSTA

RICA: San Jose, Alajuelita, 3000 ft.. Mar

1894, J. D. Smith 6592 (Holotype US 245836

(photos MEXU, PTBG)).

Sommera dunlapii L. O. Williams, Phytologia

26: 122. 1973. Type: PANAMA: Changuinola

Valley, 17 Jan 1924, V. C. Dunlap 351

(Holotype F 635718 (photo PTBG); Isotype

US (photos MEXU, PTBG)).—Sommera

donnell-smithii Standi.

Sommera fusca Oerst. ex Standi., J. Wash.

Acad. Sci. 17: 339. 1927. Type: MEXICO:

[Presumably Oaxaca] Jocoaltepec, Jun 1842,

F. Liebmann 11720 [=Liebmann Rubiaceae

36] (Holotype C, n.v.; Isotype US (photos

MEXU, PTBG)).

Sommera grandis (Bartl. ex DC.) Standi., N.

Amer. FI. 32: 145. 1921. Petesia grandis

Bartl. ex DC., Prodr. 4: 395. 1830. Type:

MEXICO: Without precise locality, Haenke

s.n. (Holotype presumably PR, n.v., fragment

G-DC, n.v. (microfiche 4; 395.1)).

Sommera guatemalensis Standi., Contr. U.S.

Natl. Herb. 17: 436. 1914. Type:

GUATEMALA: Alta Verapaz, Cubliquitz,

350 m. May 1902, H. von Tuerckheim 8225

(Holotype US 398487 (photos MEXU,

PTBG)).

Sommera mesochora Standi., Contr. U.S. Natl.

Herb. 17: 439. 1914. Type; PANAMA;

Chiriqui, vicinity of El Boquete, 1000-1300

m, 2-8 Mar 1911, IT. R. Maxon 4941

(Holotype US 675326 (photos MEXU,

PTBG)).—Sommera donnell-smithii Standi.

Sommera montana L. O. Williams, Phytologia

26: 124. 1973. Type: HONDURAS: Morazan,

margen de la Quebrada de La Paz, en medio

de la floresta de pino-liquidambar de la

Montana Zanquin, 1600 m, 14 May 1950, A.

Molina R. 2829 (Holotype F 1389080 (photo

PTBG); Isotypes EAP, n.v., US (photos

MEXU, PTBG)).

Sommera parva Lorence, Novon 4: 134, fig. 9.

1994. Type: MEXICO: Chiapas, Mun. de

Tila, Pie de Cerro K’abana, 900 m, 1 Jul

1982, A. Mendez Ton (A. Shilom Ton) 4367

(Holotype MEXU; Isotypes BM, F, MO

(photo PTBG), NY, PTBG, US, XAL).

Sommera rivularis L. O. Williams, Phytologia

26: 125. 1973. Type: COSTA RICA:

Puntarenas, Rio Esquinas, near kilometer 42,

30 m, 8 Dec 1949, P. H. Allen 5431 (Holotype

F 1679274 (photo PTBG); Isotypes EAP, n.v.,

F, US (photos MEXU, PTBG)).—Sommera

donnell-smithii Standi.

Sommera subcordata Standi., N. Amer. FI. 32:

144. 1921. Type; MEXICO: Sinaloa,

Colomas, 1897,7. N. Rose s.n. (Holotype US

866799 (photos MEXU, PTBG)).—Sommera

grandis (Bartl. ex DC.) Standi.

Sommera zygocalyx L. O. Williams, Phytologia

26: 126. 1973. Type: MEXICO; Oaxaca,

Distr. Choapam [=Choapan], Yaveo, Arroyo

de Culebras, 430 m, 21 Mar 1938, Y. Mexia

9191 (Holotype F 982858 (photos MEXU,

PTBG); Isotypes UC (photo PTBG), US

(photos MEXU, PTBG)).—Sommera

arborescens Schltdl.

Spermacoce L., Sp. PI. 1: 102. 1753. Type:

Spermacoce tenuior L.

Spermacoce ambigua Brandegee, Zoe 5: 180.

1904. Type: MEXICO: Veracruz, Cordoba,

Jan 1904, C. A. Purpus 4/7 (Holotype UC,

n.v.; Isotypes MO (photo PTBG), US (photos

MEXU, PTBG)).—Diodia brasiliensis

Spreng.

Spermacoce apiculata Willd. ex Roem. &

Schult., Syst. Veg. 3: 531. 1818. Type:

BRAZIL: Without precise locality,

Hoffmannseggs.n. (B-WI 2626,

n.v.).

r.3:lV25<5. 1798. Type: Cuba

from MEXICO], Jacquin s.n.

j type W,n.v., fragment F). For

n, see Anderson (1972: 110-111

.(Mill.)B. L. Rob.

. Gen. PI. 3: 123. 1

170

Lorence

Type: Habitat in plerisque Nov. Hisp.

temperatis agris. Floret Julio (presumably

MA, n.v.).—Status uncertain.

Spermacoce tetracocca M. Martens & Galeotti,

Bull. Acad. Roy. Soc. Bmxelles 112: 132.

1844. Type: MEXICO: Michoacan, Santa

Maria, near Morelia, H. Galeotti 2587

(Holotype presumably BR, n.v.; Isotype K,

n.v., P (photos MEXU, PTBG)).—Richardia

tricocca (Torr. & A. Gray) Standi, subsp.

tetracocca (M. Martens & Galeotti) W. H.

Lewis & R. L. Oliv.

Spermacoce tetraquetra A. Rich, in Sagra,

Hist. Fis. Cub. Bot. 11: 29. 1850. Type:

CUBA: Without precise locality, R. de la

Sagra s.n. (Holotype presumably P, n.v.;

probable Isotype F (photo PTBG)).

Spermacoce vegeta (Standi. & Steyerm.) C. D.

Adams in W. C. Burger 8c C. M. Taylor,

Fieldiana, Bot., n.s. 30: 320. 1993. Borreria

vegeta Standi. & Steyerm., Publ. Field Mus.

Nat. Hist., Bot. Ser. 23: 21. 1943. Type:

GUATEMALA: Dept. Jutiapa, between

Jutiapa and La Burrera, NE of Jutiapa,

800-850 m, 1 Nov 1940, P. C. Standley

76007 (Holotype F 1091300 (photos MEXU,

PTBG)).

Spermacoce verticillata L., Sp. PI. 1: 102.

1753. Type: JAMAICA: Hort. Cliff. 33 (BM,

n.v.). Borreria. For typification see Rendle

(1934: 331).

Spermacoce villosa Sw., Prodr. Veg. Ind. Occ.

29. 1788. Type: JAMAICA: Swartz s.n.

(Holotype presumably BM, n.v.).

—Mitracarpus hirtus (L.) DC.

Stachyarrhena Hook, f.. Hook. Icon. PI. 1068.

1870. Type: Stachyarrhena spicata Hook. f.

Stachyarrhena heterochroa Standi., Contr.

U.S. Natl. Herb. 18: 142. 1916. Type:

PANAMA: Colon, Loma de la Gloria, near

Fato (Nombre de Dios), 10-104 m, Jul-Aug

1911, H. Pittier 4078 (Holotype US 679174

(photos MEXU, PTBG); Isotype F).

Staelia Cham. & Schltdl., Linnaea 3: 364, pi. 3,

fig. 3. 1828. Type: Staelia thymoides Cham.

& Schltdl.

Staelia scabra (C. Presl) Standi., Publ. Field

Columbian Mus., Bot. Ser. 8: 385. 1931.

Diphragmus scaber C. Presl, Bot. Bermerk.

81. 1844. Type: MEXICO: Habitat in Mexico,

precise locality or collector not cited.

Stannia H. Karst., Auswahl Gewachse

Venezuela 27. 1848. Type: Stannia formosa

H. Karst.

Stannia panamensis Walp. & Duchass., Linnaea

23: 755. 1850. Type: PANAMA: Isthmo

Panamensis, Duchassaing ? (location

unknown).—Posoqueria panamensis (Walp.

& Duchass.) Walp.

Stenostomum C. F. Gaertn., Fruct. Sem. PI. 3:

69. 1806. Type: Stenostomum lucidum (Sw.)

C. F. Gaertn.

Stenostomum lucidum (Sw.) C. F. Gaertn.,

Fmct. Sem. PI. 3: 69. 1806. Laugeria lucida

Sw., Prodr. Veg. Ind. Occ. 48. 1788. Type:

JAMAICA: Without precise locality, Swartz

s.n. (presumably S, n.v.).

Steyermarkia Standi., Publ. Field Mus. Nat.

Hist., Bot. Ser. 22: 216. 1940. Type:

Steyermarkia guatemalensis Standi.

Steyermarkia guatemalensis Standi., Publ.

Field Mus. Nat. Hist., Bot. Ser. 22: 216. 1940.

Type: GUATEMALA: Izabal, Rio Duke, 2^

miles west of Livingston, on south side (left.-

hand side going up river), alt. 1-3 m, 16 Apr

1940, J. A. Steyermark 39520 (Holotype F

1035008 (photo PTBG); Isotypes F 1035009,

MO (photos MEXU, PTBG)).

Stomandra Standi., Publ. Field Mus. Nat. Hist.,

Bot. Ser. 23: 247. 1947. T3^e: Stomandra

costaricensis Standi. [Herein treated as a

synonym of Rustia Klotzsch.]

Stomandra costaricensis Standi., Publ. Field

Mus. Nat. Hist., Bot. Ser. 23: 247. 1947.

Type: COSTA RICA: San Jose, basin of El

General, 675-900 m, Mar 1940, A. P Skutch

4777 (Holotype F 1044344 (photo PTBG);

Isotypes MO (photo PTBG), US).—Rustia

costaricensis (Standi.) Lorence

176

Lorence

MASCARENE ISLANDS: Mauritius et

Bourbon, 1770-1780, P. Commerson s.n.

(Holotype P-JU, n.v.). Occasionally cultivated

in tropical America and sparingly naturalized

at least in Costa Rica. The name Vangueria

edulis Lam., a synonym, has sometimes been

applied to collections from the Neotropics.

Vameria L. Type: Vameria augusta L., Amoen.

Acad. 4: 136. 1759. [Herein treated as a

synonym of Gardenia L.]

Vameria augusta L., Amoen. Acad. 4: 136.

1759. Type: Not cited. According to Smith

(1974), the description and illustration may be

taken as the type.—Gardenia augusta (L.)

Merr.

Virecta L. f., Suppl. 17, 134. 1781. Type:

Virecta biflora L. f [Herein treated as a

synonym of Sipanea Aubl.]

Virecta biflora L. f, Suppl. 17, 134. 1781.

Type: SURINAM: Without precise locality,

Dalbergs.n. (presumably LINN, n.v.).

—Sipanea biflora (L. f) Cham. & Schltdl.

Warszewiczia Klotzsch, Flora 36: 716. 1853;

Monatsber. Kdnigl. Preuss. Akad. Wissensch.

Berlin 1853: 496. 1853. Type: Warszewiczia

coccinea (A. DC.) Klotzsch (Basionym:

Macrocnemum coccineum Vahl.)

Warszewiczia coccinea (Vahl) Klotzsch,

Monatsber. Konigl. Preuss. Akad. Wissensch.

Berlin 1853: 497. 1853. Macrocnemum

coccineum Vahl, Symb. Bot. 2: 38. 1791.

Type: TRINIDAD: Without precise locality,

von Rohr (presumably C, n.v.).

Warszewiczia pulcherrima Klotzsch, Monatsber.

Konigl. Preuss. Akad. Wissensch. Berlin

1853: 497. 1853. Type: PANAMA: Veraguas,

von Warszewicz (Holotype B, presumably

destroyed).—Warszewiczia coccinea (Vahl)

Klotzsch

Watsonamra Kuntze, Rev. Gen. PI. 1: 302.

1891. Type: Watsonamra macrophylla

(Benth.) Kuntze [Herein treated as a synonym

of Pentagonia Benth.]

Watsonamra alfaroana Standi., J. Wash. Acad.

Sci. 15: 287. 1925. Type: COSTA RICA:

Alajuela, vicinity of Capulin, on the Rio

Grande de Tarcoles, 80 m, 2 Apr 1924, P. C.

Standley 40194 (Holotype US 1152712

(photos MEXU, PTBG)). Pentagonia.

—Pentagonia tinajita Seem.

Watsonamra brachyotis Stand., Contr. U.S.

Natl. Herb. 17: 443. 1914. Type: PANAMA:

Darien, Marraganti, 3 Apr 1908, R. S.

Williams 999 (Holotype US 678351 (photos

MEXU, PTBG)).—Pentagonia brachyotis

(Standi.) Standi.

Watsonamra donnell-smithii Standi., Contr. U.S.

Natl. Herb. 17: 442. 1914. Type: COSTA

RICA: La Emilia, Llanuras de Santa Clara,

250 m, Apr 1896, J. D. Smith 6590 (Holotype

US 355178 (photos MEXU, PTBG)).

—Pentagonia donnell-smithii (Standi.)

Standi.

Watsonamra gymnopoda Standi., Contr. U.S.

Natl. Herb. 17: 444. 1914. Type: PANAMA:

Col6n, Loma de la Gloria, near Fato (Nombre

de Dios), 10-104 m, 8 Jul 1911, //. Pittier

3858 (Holotype US 678935 (photos MEXU,

PTBG); Isotype US 678934 (photos MEXU,

PTBG)).—Pentagonia gymnopoda (Standi.)

Standi.

Watsonamra macrophylla (Benth.) Kuntze, Rev.

Gen. PI. 1: 302. 1891. Pentagonia

macrophylla Benth., Bot. Voy. Sulphur 105,

tab. 39. 1845. Type: PANAMA: Panama,

forests between Panama and Cruces, Barclay

s.n. (Holotype BM, n.v.).—Pentagonia

macrophylla Benth.

Watsonamra pittieri Standi., Contr. U.S. Natl.

Herb. 17: 443. 1914. Type: PANAMA: San

Bias, forests around Puerto Obladia, San Bias

coast, 0-50 m, Aug 1911, //. Pittier 4298

(Holotype US 679414 (photos MEXU,

PTBG); Isotype US 679415 (photos MEXU,

PTBG)). Pentagonia. —Pentagonia

pinnatifida Seem.

Watsonamra pubescens Standi., Contr. U.S.

Natl. Herb. 17: 441. 1914. Type: PANAMA:

Canal Zone, around Tabemilla, along the

railroad, 20-25 m, 6 Jul 1911, //. Pittier 3822