FACULTAD DE CIENCIAS NATURALES

Y OCEANOGRAFICAS

UNIVERSIDAD DE CONCEPCION

CHILE

DIRECTOR DE LA REVISTA

Andrés O. Angulo

REEMPLAZANTE DEL DIRECTOR

Oscar Matthei J

REPRESENTANTE LEGAL

Sergio Lavanchy Merino

PROPIETARIO

Universidad de Concepción

DOMICILIO LEGAL

Víctor Lamas 1290, Concepción, Chile

EDITOR EJECUTIVO GAYANA BOTANICA

Carlos M. Baeza

COMITE EDITORIAL

MIREN ALB

anesidal. ea de Chile

KRISLER ALVEAL

Universidad de Concepción, Chile

GREGORY ANDE

University of ias

SerGIO A

Universidad A Valparaíso

CarLos Bicu

Instituto de Botánica. Sao Paulo

NGEL L. CAB

da de san Darwinion

Luis CORCUERA

Tinivercidar 4

Concepción

DanteL CRAWFEC

The Ohio State PA

MARIELA GONZALEZ

Universidad de Concepción

JURKE GRAU

Ludwig-Maximilians Universitát Miinchen

Mary T. KaLin ARROYO

O ivemidad de Chile

GINE

Real Jardín] Botánico de Madrid

DOMIRO MARTICORENA

Universidad de Concepción

GLORIA MONTENEGRO

Universidad Católica

University of Reading

AN C. ORT

Univ ersidad A Concepción, Chile

RLOS RAMI

ridad pa de Chile

Parricio Riv

Universidad de Concepción

CLAUDE SASTRE

Museum National d'Histoire Naturelle. París

Top F. STUESSY :

Institut fúir Botanik der Universitát, Wien

CHARLOTTE Tay

Missouri Botanical Garden

GUILLERMO TEL

Universidad de Buenos Aires

CAROLINA VILLAGRA

Universidad de Chile

Para las instrucciones a los autores, ver reglamento en Gayana Botánica 54(1), 1997.

Accredited with the International Association for Plant Taxonom

vascular plants (excluding fossils), fungi and fossil fungi

Indexada en: Kew Records (Kew Garden, L

ondon); Bulletin of the Torrey Botanical Club cam Biological Abs-

tracts (BIOSIS): Ulrik's International Periodicals Directory; Botanico-Periodicum-Huntian

Diseño y diagramación

Débora Cartes $.

y for the purpose of registration of new names of

ISSN 0016-5301

CAVANA ISOVAINICA

VOLUMEN 55 NUMERO 2 1998

CONTENTS

PEÑñanLILLO, P. New combinations in genus Nassella E. Desv. Emend. pa

worth (1990) (Poaceae, Stipeae)

EccL1, E. « B.E. LEUENBERGER. Eulychnia castanea Phil. (Cactaceae): pco

graphical distribution and variation

ARROYO, M.T.K., C. CasTOR, C. MARTICORENA, M. Muñoz, L. CAVIERES, O.

Ma .«A. SQUEO, M. GROSJEAN Á R. RODRIGUEZ. The flora of Llullailla-

co National Park located in the transitional winter-summer rainfall area of the

northern Chilean Andes 93

GALLOWAY, D.J. S£ W. QuiLuor. Checklist of chilean Lichen-Forming and Li-

chenicolous Fungi

CORENA, C., T.F. Sruessy £C.M. Barza. Catalogue of the vascular Pe

ra of the Robinson Crusoe or Juan Fernandez Islands, Chile

MISSOURI BOTANICAL

JAN O 4 ÑEBa

GARDEN LIBRARY

UNIVERSIDAD DE CONCEPCION-CHILE

ISSN 0016-5301

CAVANA BOTANICA

VOLUMEN 55 NUMERO 2 1998

CONTENIDO

PeÑamLLO, P. Nuevas combinaciones en el género Nassella E. Desv. Emend.

Barkworth (1990) (Poaceae, Stipeae) 85

EccLt, E. € B.E. LeuENBERGER. Eulychnia castanea Phil. (Cactaceae): Distri-

bución geográfica y variación 89

ARROYO, M.T.K., C. Castor, C. MARTICORENa, M. Muñoz, L. CAvIERESs, O.

Marrhel F.A. Squeo, M. GROSJEAN $ R. RobriGuEz. La flora del parque na-

cional Llullaillaco ubicado en la zona de transición de las lluvias de invierno-

verano en los Andes del norte de Chile 93

GALLOWwaY, D.J. £ W. QuiLnor. Lista patrón de los líquenes y hongos ..

nícolas de Chile

MARTICORENA, C., T.F. Sruessy £C.M. BAEZA. Catálogo de la flora ea

de las islas Róbinson Crusoe o Juan Fernández, Chile

UNIVERSIDAD DE CONCEPCION-CHILE

“Los infinitos seres no podrán perfectamente conocerse sino luego que

los sabios del país hagan un especial estudio de ellos”.

CLAUDIO Gay, Hist. de Chile (1847)

PORTADA: Lactoris fernandeziana Phi., único representante de la familia Lactori-

daceae, endémica del archipielago de Juan Fernández, Chile (W. Hems-

ley, Report on the botany of Juan Fernández, the southeastern

Moluccas, and the Admiralty Islands. Rep. Sci.Results Voyage H.M.S.

Challenger. Botany 1(3): 1-275, lám. 59.

ESTA REVISTA SE TERMINO DE IMPRIMIR

] DE 1999

EN LOS TALLERES DE

IMPRESOS ANDALIEN,

ROZAS 1591, CONCEPCION, CHILE.

EL QUE SOLO ACTUA COMO IMPRESORA

PARA

Gayana Bot. 55(2): 85-88, 1998.

ISSN 0016-5301

NUEVAS COMBINACIONES EN EL GENERO NASSELLA E. DESV.

EMEND. BARKWORTH (1990) (POACEAE, STIPEAE)

NEW COMBINATIONS IN GENUS NASSELLA E. DESV.

EMEND. BARKWORTH (1990) (POACEAE, STIPEAE)

Patricio Peñailillo'-? to!

RESUMEN

Después de revisar el 90% de las especies sudamerica-

nas de la tribu Stipeae (Poaceae), se ha llegado a la

conclusión que un concepto amplio del género Nasse-

lla, tal como aquel propuesto por Barkworth en 1990,

es más natural que el propuesto por Parodi en 1947.

En el presente trabajo se transfieren 14 especies trata-

das como Stipa L. s.!. al género Nassella E. Desv.

emend. Barkworth, realizándose las nuevas combina-

ciones.

PALABRAS CLAVES: Poaceae, sao Nassella, com-

binaciones nuevas, Sudaméric

INTRODUCCION

Los recientes estudios en la tribu Stipeae han

llevado a subdividir el género Stipa L. s.l. en va-

rios géneros monofiléticos (Barkworth y Everett,

1988; Barkworth, 1990; Barkworth, 1993; Rojas,

1993; Jacobs et al., 1995; Peñailillo, 1996; Torres,

1997; Jacobs y Everett, 1997; Rojas, 1998).

La interpretación de Parodi (1947) del género

Nassella s.str. ha predominado en la literatura

agrostológica (Hitchcock, 1927; Burkart, 1969;

Roig, 1978; Nicora y Rúgolo, 1987; Muñoz

Schick, 1990; Tovar, 1993; Torres, 1993). Parodi

(l.c.) incluye en este taxón a aquellas gramíneas es-

tipoídeas de antecios pequeños, lateralmente com-

primidos, lema de márgenes sobrepuestos, pálea

rudimentaria, arista fácilmente caediza, cañas pluri-

'Instituto de Biología Vegetal y Biotecnología, Uni

versidad de Talca, Casilla 747, Talca, Chile; ai

ppenaili Cpehuenche.utalca.cl

"Trabajo realizado en el Departamento de enga

como alumno becado de la Escuela de Graduados

la Universidad de Concepción, PI N” 9232.06-6.

ABSTRACT

pei reviewing about 90% of South American species

the tribe Stipeae (Poaceae), it has been concluded

e Barkworth's concept of Nassella s.l. in 1990 is

more consistent that proposed by Parodi in 1947. In

this paper, 14 species of Stipa s.l. are transferred to ge-

nus Nassella E.Desv. emend. Barkworth, and the new

combinations are proposed.

KEYWORDS: Poaceae, A Nassella, new combi-

nations, South Americ

nodes generalmente ramificadas, y vainas común-

mente menores que los entrenudos. En 1990, Bark-

worth publica una revisión del género Nassella, en

donde se proporciona una detallada historia taxo-

nómica y se realiza una enmienda de la diagnosis.

El género enmendado por Barkworth incluye 79

especies del continente americano, pero principal-

mente de América del Sur, de las cuales 65 corres-

ponden a nuevas combinaciones.

El presente trabajo tiene por objeto dar a co-

nocer 14 nuevas combinaciones en el género

Nassella para América del Sur.

MATERIALES Y METODOS

Se estudiaron 2.000 especímenes que inclu-

yen 225 especies y 9 géneros (Aciachne, Ach-

natherum, Lorenzochloa, Nassella, Ortachne,

Oryzopsis, Piptatherum, Piptochaetium y Stipa).

os especímenes estudiados provienen de los si-

guientes herbarios, BAA, CONC, LPB, MVFA,

MO, SGO, SI, US y W.

Para los estudios morfológicos, las espigui-

Gayana Bot. 55(2), 1998,

llas se disectaron y montaron en líquido de Holler.

Los embriones maduros se prepararon siguiendo la

técnica propuesta por Barkworth (1982) y los pre-

parados epidérmicos de la lema según el método

de Thomasson (1978a). Las observaciones ma-

croscópicas se realizaron en una lupa estereoscópi-

ca Zeiss y las microscópicas en un microscopio

fotónico Zeiss y en un microscopio electrónico de

barri tec modelo Autoscan de la Universidad

de Concepción. El secado a punto crítico de los an-

tecios se realizó en un secador Balzers Unions, los

cuales previo a su observación, fueron sometidos a

un sombreado con oro.

RESULTADOS Y DISCUSION

Al revisar una gran cantidad de material de

herbario de las especies de la tribu Stipeae cita-

das para América del Sur sobre la base de carac-

teres de la morfología vegetativa y reproductiva,

como asimismo del embrión maduro y de la epi-

dermis de la lema, se ha llegado a la conclusión

que el género Nassella E. Desv. emend. Bark-

worth (1990) es un grupo monofilético. En Suda-

mérica, este concepto es compartido por Torres

(1997) en su revisión de la tribu Stipeae del no-

roeste de Argentina y por Rojas (1998) en su tra-

tamiento de las especies de la tribu en Bolivia.

Nassella s.!. incluiría a todas las especies de

la tribu Stipeae con las siguientes características:

Lema coriácea, de márgenes convolutados a la

madurez, ocultando la pálea y ápice fusionado for-

mando una corona notoria o inconspicua. Pálea

hialina, glabra, emarginada, generalmente enervia,

de 1/3 de la longitud de la lema o menor. Embrión

de epiblasto largo. Patrón epidérmico de la lema

tipo “Nassella pungens” sensu Thomasson (1976,

1978b), es decir, formado por células largas acor-

tadas en longitud cuyas paredes son gruesas y sili-

ficadas; ganchos presentes o ausentes.

Las nuevas combinaciones propuestas son las si-

guientes:

Nassella argentinensis (Speg.) Peñail.. comb. nov.

BASIONIMO: Stipa argentinensis Speg., Revista

Argent. Bot. 1: 45. 1925. Tipo: “Hab. en las pra-

deras cerca de la Estación Gálvez, Prov. Sta. Fe,

Ener. 1906”.

Nassella ACA e Zanin et Longhi-Wag-

ner) Peñail., comb. n

BASIONIMO: Stipa ori A. Zanin et Longhi-

Wagner, Bradea 5 (33): 342, fig. 1. 1990. Tipo:

“BRASIL -Río Grande Do Sul: Cambará Do Sul,

Itaimbezinho, 18 dez. 1987, A. Zanin 17”. (Holo-

tipo: CEN, Isotipo: MVFA).

Nassella coquimbensis (Matthei) Peñail., comb. nov.

BASIONIMO: Stipa coquimbensis Matthei, Gayana,

Bot. 13: 35, f. 5A-G. 1965. Tipo: “CHILE. Prov. Co-

quimbo. Carretera dre 8 Km al Norte de

la Quebrada del T e. Leg. Marticorena-Matthei

163. 13-X-1963”. pad CONC 29361!).

Nassella entrerriensis (Burkart) Peñail., comb. nov.

BASIONIMO: Stipa entrerriensis Burkart, Bol.

Soc. Argent. Bot. 12: 285, f. 1. 1968. Tipo: AR-

GENTINA, prov. Entre Ríos, dep. La Paz, Bov-

ril, J. M. Jozamí 8”. (Holotipo: SI!).

Nassella ibarrensis (Kunth) Peñail., comb. nov.

BASIONIMO: Stipa ibarrensis Kunth, in Humb.

Bonpl. et Kunth, Nov. Gen. Sp. 1: 125. 1816. Tipo:

“Crescit ad muros urbis Quitensis, Villa Ibarra, in

subfrigidis, alt. 1184 hexap, perennis, Floret Janua-

rio, Humboldt et Bonpland”. (Holotipo: P. Clasto-

tipo: US 866103).

Nassella karstenii (Hitchc.) Peñail., comb. nov.

BASIONIMO: Stipa karstenii Hitchc., Contr. U.S.

Natl. Herb. 24(7): 274. 1925. Tipo: “Collected at

Bogotá, Colombia, by H. Karsten”. (Holotipo: US

1126745)

Nassella pittieri (Hitchc.) Peñail., comb. nov.

BASIONIMO: Stipa pittieri Hitchc., Contr. U.S. Natl.

Herb. 24(7): 289. 1925. Tipo: “collected below Pi-

taió, Río Palo Basin, Tierra Adentro, State of Cauca,

Colombia, altitude 2,400 meters, February, 1906, by

H.Pittier (no. 1438)”. (Holotipo: US 531634).

Nassella psittacorum (Speg.) Peñail.. comb. nov.

BASIONIMO: Stipa psittacorum Speg., Anales

Mus. Nac. Montevideo 4(19): 165. 1901. Tipo:

“Hab. Abunde in praeruptis collinis Arroyos de

Nuevas combinaciones en Nassella: PEÑAILILLO, P.

los papagallos prope Mendoza, Febr. 1901 (C.S-

pegazzini)”. (Holotipo: LP ex LPS 2445. Isotipo:

US 141663, 1721313).

Nassella planaltina (A. Zanin et Longhi-Wag-

ner) Peñail., comb. nov.

BASIONIMO: Stipa planaltina A. Zanin et Long-

hi-Wagner, Bradea 5(33): 344, fig. 2. 1990. Tipo:

“BRASIL-Río Grande do Sul: Cambará do Sul,

Itaimbezinho, 18 nov. 1988, A. Zanin 4 F.A. Sil-

va Filho 151a”. (Holotipo: ICN, Isotipo: CEN).

Nassella rhizomata (A. Zanin et Longhi-Wag-

ner) Peñail., comb. nov.

BASIONIMO: Stipa rhizomata A.Zanin et Longhi-

Wagner, Bradea 5(33): 345, fig. 3. 1990. Tipo:

“BRASIL-Santa Catarina: Irani, 28 nov. 1986,

VALES et al. 10561”. (Holotipo: CEN, Isotipo:

MVFA)

Nassella sellowiana (Nees ex Trin. er Rupr.) Pe-

ñail., comb. nov.

BASIONIMO: Stipa sellowiana Nees ex Trin. et

Rupr., Sp. Gram. Stipac. 38. 1842. Tipo: “Brasi-

lia merid. (SELLOW!)”. (Holotipo: LE, TRIN.

Clastotipo: US 157500).

Nassella tenuiculmis (Hackel) Peñail.., comb. nov.

BASIONIMO: Stipa tenuiculmis Hackel, Verh.

Zool.-Bot. Ges. Wien 65 (1-2): 75. 1915. Tipo:

“BRAZIL, Río Grande do Sul: Mun. Soledade:

in campis prope Lagao, Dec 1910, C.Juergens”.

(Holotipo: W, Hack. Herb.; Clastotipo: US

3168612).

Nassella vallsii (A.Zanin et Longhi-Wagner) Pe-

ñail. comb. nov

BASIONIMO: Stipa vallsii A.Zanin et Longhi-Wag-

ner, Bradea 3(33): 346, fig. 4. 1990. Tipo: “BRA-

SIL-RIO GRANDE DO SUL: S,.o Francisco de

Paula, 18 nov. 1988, A. Zanin £ F.A.Silva Filho

148”. (Holotipo: CEN, Isotipo: MVFA).

Nassella vargasii (Tovar) Peñail., comb. nov.

BASIONIMO: Stipa vargasii Tovar, Phytologia

47(6): 445. 1981. Tipo: “Collected at laderas de

Tarpata, prov. Urubamba, depto. Cuzco, Perú, al-

titude 2860-3000, March 8, 1963, by César Var-

gas (n. 14127)”. (Holotipo US 2474222!).

MATERIAL ADICIONAL ESTUDIADO

Nassella argentinensis (Speg.) Peñail. ARGEN-

TINA: Prov. Chaco. Distrito General Donovan,

La Escondida, 3-111-1958, Costany 5, (SD.

Nassella coquimbensis (Matthei) Peñail. CHI-

LE: IV Región. Prov. Choapa. Illapel, Huente-

lauquén, Estancia La Cebada, 60 m, 22-X-1971,

Jiles 5754, (CONC)

Nassella entrerriensis (Burkart) Peñail. ARGEN-

TINA. Prov. Entre Ríos. Dep. Concordia, Arroyo

Robledo a Redomom, 8-XI-1965, Burkart 25898,

(SD.

Nassella ibarrensis (Kunth) Peñail. ECUADOR:

Andes, SPRUCE 5819 (US s.n. Ex Herb Boissier).

Prov. Cotoxapí. Ad basin Monte Cotoxapí, Sodiro

s.n., 1890 (US). Prov. Pichincha. Pomarqui, dry

slope, 2500 m, 6-111-1939, Asplund 6688, (US).

Nassella karstenii (Hitchc.) Peñail. COLOMBIA:

Int. Putumayo-Corregimiento El Encano, Laguna

de la Cocha, páramo El Tabón, 3300 m, 10/11-

VII1-1939, García Barriga 7771, (US).

Nassella pittieri (Hitchc.) Peñail. COLOMBIA:

Edo. Cauca. Las Escaleretas, Moras Valley, río

Paez basin, tierra adentro, 2500-3000 m, II-1906,

Pittier 1334, (US). Edo. Cundinamarca. Zipaquirá,

2700 m, 5-1-1851, Triana 3345, (US). Int. Putu-

mayo-Corregimiento El Encano, Laguna de la Co-

cha, páramo “El Tábano”, 3300 m, 10/11-VIH-

1939, García Barriga 7762, (US).

Nassella planaltina (A. Zanin er Longhi-Wag-

ner) Peñail. BRASIL: Edo. Paraná. General Car-

neiro, 2-XII-1987, Valls et al. 11474, (MVFA).

Nassella sellowiana (Nees ex Trin. ef Rupr.) Peñail.

BRASIL: Edo. Paraná. Faxinal dos Souza (Mun.

Gal. Carneiro), 7-XIL-1971, Hatschabach, Smith,

Klein 28351, (MVFA). Ponte Grossa, Desvio Ribas

in Campo, 29-X1-1910, Dusen 12102A, (SD. Cam-

de Palma (Mun. Agua Doce), 30 km SE of Ho-

rizonte (26"45"S, 51%24"W), 1000-1200 m,

Gayana Bot. 55(2), 1998.

Klein 28351, (MVFA). Ponte Grossa, Desvio Ribas

in Campo, 29-XI-1910, Dusen 12102A, (SD. Cam-

un. Agua Doce), 30 km SE of Ho-

rizonte (26%45”S, 51%24*W), 1000-1200 m,

3-XII-1964, Smith et Klein, ARGENTINA: Prov.

Misiones, Apóstoles a San Javier, 11-1922, Molfino

s.n., (SD.

Nassella tenuiculmis (Hackel) Peñail. URU-

GUAY": Dep. Treinta y Tres. Quebrada de Las Cue-

vas, 8-X-1966, Rosengurtt et al. 10288, (MVFA).

ARGENTINA: Prov. Buenos Aires. Pdo. Balcarce,

Sierra la Brava, ladera SW a cumbre, 11-X1-1962,

Boelcke, Matthei y Correa 9385, (BAA, CONC).

Prov. La Pampa. La Pampa, Parque Luro, 29-XI/2-

XIP-1959, Rosengurtt B-7801, (CONC, MVFA). La

Pampa, Utracan, entre General Acha y Los Caran-

chos, 16-XII- 1952, Ragonese y Piccinini 8860,

(BAB). Dep. Lihuel- Calel, 29-1/2-11-1959, Rosen-

gurtt B-7856, (MVFA). Dep. Santa Rosa, 30-x-

1972, Kaprovickas et al. 22299, (CONC). Entre

Anguil y Santa Rosa, 22-X1-1955, Ragonese-Picci-

nini 9231, (BAB). Prov. Río Negro. Pichi Mahuida,

puente sobre el río Colorado, 7-X1-1965, Correa y

Nicora 3157, (BAB, CONC)

AGRADECIMIENTOS

El autor agradece a los directores y curado-

res de los herbarios consultados por las facilida-

des otorgadas, como asimismo a los profesores

Clodomiro Marticorena, Marcelo Baeza y Rober-

to Rodríguez por sus sugerencias. También ex-

preso mi reconocimiento por su asistencia técnica

al personal del Laboratorio de Microscopía Elec-

trónica de la Universidad de Concepción.

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07 de mayo de 1999

Gayana Bot. 55(2): 89-92, 1998.

ISSN 0016-5301

EULYCHNIA CASTANEA PHIL. (CACTACEAE): GEOGRAPHICAL

DISTRIBUTION AND VARIATION

EULYCHNIA CASTANEA PHIL. (CACTACEAE): DISTRIBUCION GEO-

GRAFICA Y VARIACION

Urs Eggli' 8 Beat Ernst Leuenberger”

ABSTRACT

The geographical range of tiene castanea (Cacta-

ceae) has been Pr in th and was found

be a rather restricted area do a small fraction of

the central Chilean coast. Populations om scattered

and sometimes reduced in number of individuals. The

conservation status is interpreted as being e of dan-

ger, since a y and viable population occurs within

Fray Jorge na Park. A case of UTA intro-

rhnin nrirdr +

also reporte

KEYWORDS: Eulychnia, conservation, Cactaceae.

INTRODUCTION

The genus Eulychnia is a small genus of sh-

rubby to arborescent columnar cacti, and 4 of the

presumably $5 species are restricted to Chile (the

exception being E. ritteri from southern Peru).

The two species Eulychnia acida and E. iqui-

quensis form a conspicuous element in the vege-

tation of the semi-arid and arid Norte Chico. The

history and typification of Eulychnia taxa, as

well as some preliminary notes on possible sy-

peca have been the subject of a recent con-

tion (Leuenberger 4 Eggli, 1998).

Entrinda castanea was described by R. A.

Philippi in 1864, based on a collection by Land-

beck from the coast near Los Molles (then in

Aconcagua Province, today in Petorca Province).

It was named for the diagnostic pungent thin spi-

'Stádtische Sukkulenten-Sammlung Ziirich, Mythen-

quai 88, CH-8002 Zúrich, Switzerland.

“Botanischer Garten und Botanisches Museum B

Dahlem, Kónigin-Luise-Strasse 6-8, D-14191 Berlin,

Germany.

RESUMEN

Se estudia la distribució a a less

pe direc lead resultados dema que. su distniba

de Chile. Las poblaciones s son más o menos aisladas y

frecuentemen: úmero deci de plantas.

El estado de niga se i “fuera de

peligro” a base de una población siendida y vado den-

tro del Parque Nacional Fray Jorge. Se menciona un ca-

A DA sim da Prairie 5d,

PALABRAS CLAVES: Eulychnia, conservación, Cactaceae.

nes produced from the areoles of the pericarpel,

and the spiny fruits were compared with the

fruits of the chestnut by Philippi (Lat. castaneus

= chestnut). Although Philippi did not publish an

illustration of E. castanea, and despite some pro-

blems associated with the type material, the iden-

tity of the taxon does not present any problems,

and its typification was recently resolved (Leuen-

berger « Eggli, 1998).

GEOGRAPHICAL DISTRIBUTION: The type locality

of Eulychnia castanea is at Los Molles (32*14"S,

Región de Valparaiso, Prov. Petorca), where the

taxon can still be observed today on the coastal

rocks below the houses. The northern limit of the

range of this strictly coastal taxon is given as “Ta-

linay” in the literature (Ritter, 1980; Hoffmann,

1989), and this is taken to represent the region of

the Quebrada Talinay and Cerro Talinay (c.

305 1'S, Región de Coquimbo, Prov. Limarí) (the

“Altos de Talinay” further N cannot qualify since

E. castanea has a strictly coastal distribution).

Our collecting activities in connection with

a study of the Cactaceae for the “Proyecto Flora

de Chile” have now considerably extended the

Gayana Bot. 55(2), 1998.

known geographical range for E. castanea to-

wards the North, just into Prov. Elqui. Together

with several additional collections, we have esta-

blished a more or less continuous distribution of

E. castanea along the coast between Los Molles

in the South and near Puerto Aldea on the Len-

gua de Vaca peninsula W of the Bahia Tongoy

(3018.51”S) in the North (Map 1).

Whether this represents the true geographi-

cal range of the taxon remains open to debate,

and subsequent searches may well reveal additio-

nal localities. Our own observations indicate that

E. castanea does neither occur to the north of the

known range at Punta Teatinos N of La Serena,

nor at Quintero to the south of the known range.

All our observations as well as remarks on

the labels of the few available herbarium speci-

mens indicate that E. castanea is a strictly coastal

taxon, occupying but a narrow fringe along the

coast. Its ecological amplitude appears to be consi-

derable, and apparently well-established plants can

be observed on coastal rocks, among coastal peb-

bles (e.g. near Pichidangui) or in coastal semi-sta-

bilized dune sands (e.g. just N of Puente El

Teniente). In only one place (near Huentelauquen)

could we observe E. castanea planted as compo-

nent of a “living fence” primarly made up by one

of the numerous forms Echinopsis chiloensis.

CONSERVATION: Since the coast strip ape

Api Papudo and Zapallar is of easy access from

, the area is currently heavily ios

ps erckend and summer tourism. Considerable

areas between the Panamerican Highway and the

coast are already fenced off and divided into lots

for sale and house construction. The remaining area

suffers from grazing impact and there are also scat-

tered plantations of Eucalyptus and/or Pinus. As a

consequence, the already narrow distribution of

Eulychnia castanea appears to be under severe

pression from these developments. Its conservation

status needs to be assessed, although Benoit (1989)

classified it as out of danger without giving further

details. Hoffmann (1989) does not comment on the

conservation status of this species.

While the number of populations of E. cas-

tanea remains unknown, the distribution map im-

plies a more or less continuous occurrence along

the coast. Despite the threats posed by the increa-

sing urbanization of the area and increasing gra-

zing pressure, the species may have a good chance

to survive, as we have observed the plant growing

in moderately disturbed areas, e.g. at Los Molles,

Pichidangui, in the Quebrada del Teniente, and al-

so growing in hedges at Huentelauquen. Moreo-

ver, a very vigorous population consisting of a

large number of indidivuals is located within Fray

Jorge National Park, and E. castanea therefore en-

joys a fair amount of protection. lts conservation

status as “out of danger” seems to be a sound inter-

pretation, and its survival does not seem to be im-

mediately threatened as a whole. Populations in

the southern part of the natural range are, however,

under some pressure as outlined above, and it

would be a good idea to suggest to landowners and

communities to protect the natural populations as a

local biodiversity treasure, and even to propagate

the plant at appropriate sites, e.g. in hedgerows. A

study of the biology of this species by a local bio-

logist would seem very promising for a more de-

tailed assessment of its conservation status.

HYBRIDIZATION INVOLVING EULYCHNIA CASTANEA:!

At the N-most limit, R. Kraus (pers. comm.) obser-

ved typical Eulychnia castanea as well as plants

which appear intermediate between E. castanea and

one of the larger growing species of the genus, jud-

ging from the woolly-bristly indumentum of the pe-

ricarpel and the larger stature of the plants (R.

Kraus photos nos. 1988 and 1989, ZSS). Similar

plants were also collected back in 1947 by B. Spa-

rre (Sparre 1974). While E. castanea cannot be dis-

tinguished vegetatively from either E. acida or E.

breviflora (the only other species of the genus likely

to be present in the area), the pericarpel spination is

denser and more bristly instead of pungent, as

would be typical for E. castanea (Figs. 1, 2)

According to our observations, the range of

E. breviflora does not extend significantly to the

south of Coquimbo, while E. acida was found

with a few individuals growing in the immediate

vicinity of E. castanea on the coastal terrace in

Fray Jorge National Park (near 30? 37.94” S / 71"

42.45” W). In the absence of better evidence, we

interpret the occurrence of the densely bristly flo-

wers in E. castanea as introgression from E. aci-

da, whose flowers are characterised by a short

softly woolly tomentum (Figs. 1, 2).

MATERIAL EXAMINED: CHILE: IV Región: Prov.

Elqui: 2 km E of Puerto Aldea, 20 - 50 m, 31-X-

1997, EGGLI £ LEUENBERGER 3083 (B,

CONC, SGO, ZSS); Prov. Limarí: Parque Nacional

Fray Jorge, coastal plain, 20-100 m, 1-X1-1997,

Distribución de Eulychnia castanea: EGGLI, U. € B.E. LEUENBERGER

EGGLI € LEUENBERGER 3086d (B, ZSS: pho-

tos only); Fray Jorge, boca del Río Limarí, 11-X-

1947, SPARRE 2974 (SGO); first hill N of the

mouth of the Quebrada San Pedro, 80 - 140 m, 18-

X-1997, EGGLI £ LEUENBERGER 2945 (B,

CONC, SGO, ZSS); Prov. Choapa: Caleta Oscuro

3 km W of Puerto Oscuro (Panamericana), 20 - 100

m, 2-XI-1997, EGGLI £ LEUENBERGER 3088

(B, CONC, SGO, ZSS); Fundo Corral de Julio, km

315 de la Carretera, 1 -2 km desde el mar, 6-XI-

1976, Muñoz 921 (SGO); 7.3 km on Panamericana

N of turnoff from Panamericana to Los Vilos, on

top of coastal rocks, 40 m, 10-X1-1991, EGGLI «

LEUENBERGER 1669 (B, SGO, ZSS); Los Vilos,

s.a., Ritter 241 loc. 1 (SGO, ZSS); Pichidangui, ex-

treme tip of peninsula, coastal rocks, 10 - 50 m, 17-

X-1997, EGGLI £ LEUENBERGER 2933

(CONC, B, SGO, ZSS); Pichidangui, s.a., Ritter

241 loc. (SGO); V Región: Prov. Petorca: c. 1 - 1.5

km S of Los Molles, N side of Puente El Chivato,

20 m, 10-XI-1991, EGGLI £ LEUENBERGER

16561 (B, SGO, ZSS); Los Molles, coastal rocks

below houses, 10 - 80 m, 17-X-1997, EGGLI «

LEUENBERGER 2932a (B, ZSS: photos only).

ACKNOWLEDGMENTS

We are grateful to the authorities at the Botany

Institute of the Universidad de Concepcion for provi-

ding infrastructure and a vehicle for our studies. The

Swiss National Science Foundation (project no. 31-

39176.93 “Study of Cactaceae for the Flora of Chile

project”) as well as the Swiss Academy of Natural

Sciences are gratefully acknowledged for financial

support to the first author for field work in Chile Li-

kewise, the second author is grateful to the Research

Committee of the Cactus and Succulent Society of

America, Inc., for a grant towards travel costs in con-

nection with the above project. B and ZSS have

kindly provided support and infrastructure, and we

are further grateful to the curators at SGO and

CONC to allow us free access to their collections.

REFERENCES

gos IL L. (ED.) 1989. Red Book of Chilean terres-

ial flora (Part One). Santiago. (vi), 151 pp., 22

P< incl. en el texto, 1 mapa.

HOFEMANN, A A. E. 1989, Cactáceas en la flora silvestre

LEUENBERGER, B. E. «

Eulychnia (Cactaceae) in Chile: Notes on the ta-

nomy, types, and other old specimens. Hasel-

nia 5 (submitted).

ies R.A. 1864. Plantarum novarum chilensium.

Centuriae inclusis quibusdam Mendocinis et Pa-

taonicis. Tinnaea 33: 1-308.

RITTER, F. 1980. Kakteen in Sudamerika. Band 3. Chi-

le. Spangenberg. Pp. 857-1238. (Cactaceae).

MAP 1. Known geographical range of Enlychnia castanea. Base map drawn with Geodat V2.0 (Jacquez, 1995),

using a Lambert Azimuthal Projection. L = La Sere

na, S = Santiago, V = Valparaiso

Gayana Bot. 55(2), 1998.

FIG. 1. Advanced buds and flowers of typical Eulychnia castanea (Eggli £« Leuenberger 2945, N of

mouth of Quebrada San Pedro)

FIG. 2. Recently spent flowers and young fruits of Eulychnia castanea showing (arrow) introgression

from E. acida (EGGLI £ LEUENBERGER 3086d [photos only], coastal terrace of Fray Jorge National

Park).

Fecha de publicación: 07 de mayo de 1999

Gayana Bot. 55(2): 93-110, 1998.

ISSN 0016-5301

THE FLORA OF LLULLAILLACO NATIONAL PARK LOCATED IN THE

TRANSITIONAL WINTER-SUMMER RAINFALL AREA OF THE

NORTHERN CHILEAN ANDES

LA FLORA DEL PARQUE NACIONAL LLULLAILLACO UBICADO EN

LA ZONA DE TRANSICION DE LAS LLUVIAS DE INVIERNO-VERANO

EN LOS ANDES DEL NORTE DE CHILE

Mary T. Kalin Arroyo', Carmen Castor', Clodomiro Marticorena?, Melica Muñoz”, Lohen-

grin Cavieres”, Oscar Matthei”, Francisco A. Squeo*, Martin Grosjean' y Roberto Rodríguez?

ABSTRACT

The vascular plant flora of Llullaillaco National Park, si-

tuated mostly above 3500 m elevation in the high Andes

of the II Region of Chile (24930"S - 25%10”S; 68%30"W -

69*15”W) in the transitional zone between the summer

and winter rainfall regimes, is given, along with analysis

of endemism, presence of distributional eros and spe-

cies-accumulation tendencies. The known flora stands at

91 species and subspecies (90 species), in 58 genera and

26 families, with an additional 3 species likely to occur

in the park. Thirteen (14%) of taxa are endemic to the

Chilean , with two entirely restricted to the park.

Outstanding eb of taxa (35 in total: 38%) reach

their northern, southern or north-south distributional li-

mits Sci or very close to the park boundary. Know-

ledge of the park's flora has accumulated steadily since

R. A. Philippi's landmark expedition to the Atacama de-

sert in 1860. By 1925 over half of the known taxa had

been collected, and by 1975 over 2/3 of the species were

red. Llullaillaco National Park plays an important role in

pro: marginal populations of many high Andean

species of subtropical and

the park constitutes a critical area for understanding cli-

matic evolution in the northern Chilean Andes.

o de Biología, Facultad de Ciencias, Uni-

versidad de Chile, Casilla 653, Santiago, Chile. E-mail:

southernO abello.dic.uchile.cl.

“Departamento de Botánica, Facultad de Ciencias Natu-

rales y Oceanográficas, ac de Concepción,

Casilla 160-C, C

“Sección Botánica, Museo cial de Historia Natural,

Casilla 787, Santiago, Chile.

de Biología, Facultad de Ciencias, Uni-

versidad de La condo Casilla 599, La Serena, Chile.

de sao of Physical Geography, University of Ber-

ne, Switz

le, northern Chilean rainfall, winter rainfall,

Andean flora, endemics, north and south distributional li-

mits lysi catalog of flora.

cds

KEYWORDS: Llullaillaco National Park, HI Region of Chi-

flora, summer

RESUMEN

Se presenta la flora vascular del Parque Nacional Llu-

llaillaco, situado principalmente por sobre los 3500 m

en los Andes de la II Región de Chile (24*30'S-25*

10”S; 6830"W - 6915” w), en la zona transicional de

cular consta de 91 especies y subespecies (90 especies)

distribuidas en 58 géneros y 26 familias, con 3 especies

adicionales que posiblemente se encuentran en el par-

que. Trece (14%) taxas son endémicos a los Andes chi-

lenos, con dos restringidos en su distribución al

mismo. Un número destacable de taxa (35 en total:

38%) alcanzan sus límites de distribución norte, sur o

sur y norte dentro o muy cerca de los límites del parque.

El conocimiento de la flora del parque se ha acumulado

gradualmente a partir de la histórica expedición de R.

A. Philippi al desierto de Atacama en 1860. En 1925,

más de la mitad de la flora había sido vigoroso y en

1975 se conocía más de 2/3 de la flora. La curva de acu-

mulación de especies del parque predice que es proba-

ble el descubrimiento de algunas especies adicionales.

El Parque Nacional Llullaillaco juega un papel impor-

tante en la protecci las poblaciones marginales de

un número apreciable de especies andinas de afinidad

subtropical La composición y caracte-

y mediterránea.

na de la fora del parque ria e. oran did un

cormmmrención de

e a dd a CUR

VES: Parque Nacional Llullaillaco, IMM Re-

andina,

distribución norte y sur, análisis de acumulación de es-

pecies, catálogo florístico.

Gayana Bot. 55(2), 1998,

INTRODUCTION

Continental Chile possesses 30 National

Parks, 47 National Reserves and 13 Natural Mo-

numents. These components of the National Pro-

tected Area System of Chile (SNASPE) cover an

area Of 14.4 million hectares (Muñoz et al. 1996),

the equivalent of 19% of total land area (Arroyo $

Cavieres, 1997). Over 99% of the area under pro-

tection pertains to national parks and reserves.

Despite the high percentage of land area that is un-

der protection in Chile in comparison with a world

average of 6.3% (Heywood, 1995), published flo-

ristic lists and complete floras are available for re-

latively few protected areas. This situation is

particularly evident for the northern Chilean An-

des where there is no published floristic list for

any protected area, other than the partial list provi-

ded by Arroyo et al. (1982) for Lauca National

Park. The lack of floristic knowledge constitutes a

serious handicap for evaluating the conservation

value efficiency of the SNASPE as a whole, and

for developing management plans for individual

protected areas

In this paper we provide a working checklist

of the vascular plant flora of Llullaillaco National

Park, II Región Chile, information on endemism

and geographical ranges, and a quantitative analy-

sis of the level of floristic knowledge of the park.

The present publication was stimulated by the CO-

NICYT-Chile Sectorial Program: “Biomas y Cli-

mas Terrestres y Marinos en el Norte de Chile”

and a “Cátedra Presidencial en Ciencias” -1997

held by the first author. The “Biomas” program tar-

geted at the regional level, enabled ample floristic

exploration of poorly known areas of the II Región

of Chile over the period 1996-97, development of a

comprehensive data base of all vascular plant co-

llections available for the Region, and establish-

ment of a regional floristic list of close to 1000

taxa (Marticorena et al. 1998). The second initiati-

ve supports studies aimed at determining the

conservation efficiency of protected areas in conti-

nental Chile situated within the area of mediterra-

nean-type climate influence.

LLULLAILLLACO NATIONAL E Llullaillaco

NP, comprising 262,000 as incorpora-

ted into the National Protected Mia System of

Chile in 1995 (Muñoz et al.. 1996). The area com-

prises one of the “Sitios Prioritarios la Con-

servación de la Diversidad Biológica de Chile”

(Muñoz et al., 1996). Located latitudinally bet-

ween 24*30"S and 25*10”S, and longitudinally bet-

ween 6830” W and 69%15”W, with elevations

mostly over 3500 m (Figure 1), this remote loca-

tion is situated in the hyperarid transitional zone

between the summer and winter rainfall regimes

(Arroyo et al., 1998, Messerli et al., 1993; Fuen-

zalida 8 Rutllant, 1986, Table 1). The extremely

dry climate characterizing the park, where annual

precipitation is estimated to lie between 20-50 mm

(Messerli et al., 1993), results from the synergistic

interaction between subsiding anticyclonic air

masses of the southeast Pacific High Pressure

Belt, the drying effect of the Humboldt current,

and the rainshadow effect of the high Andes

(Arroyo et al., 1988; Messerli et al., 1993). Winter

cyclonic precipitation associated with the typical

mediterranean climate in central Chile is normally

blocked around 30%S, and the tropical convective

summer precipitation of the “invierno boliviano”

is restricted mainly to the eastern slopes of the An-

des. Based on Fuenzalida $ Rutllant (1986) and

Aravena et al. (1989), both summer and winter

rainfall in the transitional zone is of Amazonian

origin. Winter precipitation results from the colli-

sion of wet and warm tropical and cold extra-tro-

pical air masses. Summer precipitation is linked to

an anticyclonic flow pattern in the upper troposhe-

re over the eastern Altiplano. Vuille (1991) sug-

gests that the amount of precipitation received

during the winter in the transition zone is highly

underestimated because of its solid form. The cri-

tical transition zone falls some where around

24'00'S -24%30"S, corresponding to the northern

border of Llullaillaco NP. According to Vuille

(1991), salt lakes north of this limit show increa-

sed water volume during the summer months; tho-

se south of 24%S show water increase as a response

to winter precipitation. Based on Messerli et al.

(1993), probably over 50% of annual precipitation

received in Llullaillaco NP is of winter origin.

METHODS

The floristic information provided here is a

product of botanical exploration by: a) early wor-

kers such as R.A. Philippi, C. Reiche, E. Werde-

prior to the present silo: Cc) the present resear-

Flora of Llullaillaco National Park: ARROYO, M.T.K., ET AL.

chers during the 1997 summer season. All resul-

tant collections were incorporated into the compre-

hensive data base of the Flora of the II Region of

Chile where they are indexed according to precise

latitude and longitude, and elevation where possi-

ble. Specimens are deposited at CONC and SGO.

Presence of a species in the park was deter-

mined by defining the rectangular area: latitudes

25"30"-25*15”S and longitudes: 68%15'- 69%15"W

and eliminating all specimens from the rectangle

not collected within the exact boundary of the

park as per in Figure 1. Some collections made

very early on in the botanical history of the area

are not as precise with regard to location as would

be desired. The important Philippi expedition to

the Atacama desert (Philippi, 1860) which resul-

ted in many new described species for Chile, is a

typical case. Philippi (1860) explored localities on

the western side of the park, a number of which

are problematical. For example, the elevation re-

ported for the locality Río Frío, the latter which is

clearly entirely within the limits of the park,

would appear to be out by some 500 m according

to present cartographic values. Collections from

this locality have been accepted as pertaining to

the park boundaries. It would also appear that the

quoted elevations of around 3000 m for many co-

llections from Valle or Río Zorras are underesti-

mated. Judging by the fact that most of the plants

collected at this last locality pertain to Andean

bogs developed at much higher elevations, it is

probable that the correct elevation is around 3500-

4000 m. With this adjustment, many of the plants

collected at the locality Zorras by Philippi (1860)

would fall within the park boundaries.

In several instances Philippi (1860) (see al-

so Muñoz 1960) reports visual records of species

at park localities. In other cases more localities

than collection numbers are cited, making it im-

possible to determine the precise locations of the

collections. Species falling into these last two ca-

tegories, that could potentially exist in the park,

have been included in an Addendum to the cata-

log. The presence of northern and southern distri-

butional limits within the park boundaries or

very close to the latter was determined through

consultation of the Chilean flora data base and li-

terature records. The category “very close” co-

rresponds to distributional limits found within 5”

of the park boundary.

To assess the present state of floristic know-

ledge of the park, a species accumulation curve

was constructed using accumulation intervals of

25 records, beginning with R. A. Philippi's collec-

tions made in 1854. The clench model (Soberón +

Llorente, 1993) was used to mathematically fit the

accumulation curve and to determine the effects of

further collecting efforts on the park's flora.

Short descriptions and exsiccatae are given

for each species.

RESULTS

HISTORICAL DEVELOPMENT OF FLORISTIC KNOW-

LEDGE: Figure 3 shows that species registered for

the park have accumulated gradually over time.

The earliest botanical collections date back to the

last century when the German naturalist R. A.

Philippi, then Director of the Museo Nacional de

Historia Natural was directed by the government

of Chile to undertake a study of the Atacama De-

sert (Philippi, 1860). First records of species for

the park represented with a fair amount of cer-

tainty in R. A. Philippi's collections are:

- Adesmia erinacea Phil.

- A. frigida Phil.

- Carex maritima Gunnerus

- Chenopodium frigidum Phil.

- Deyeuxia robusta Phil.”

- Menonvillea frigida (Phil.) Rollins

- Montiopsis glomerata (Phil.) D.I. Ford

- M. modesta (Phil.) D.I.Ford

- Nastanthus caespitosus (Phil.) Reiche

- Oxychloe andina Phil.

- Phacelia cumingii (Benth.) A.Gray

- Potamogeton strictus Phil.

- Puccinellia frigida (Phil.) LM.Johnst.

- Ranunculus cymbalaria Pursh f. exilis (Phil.)

Lourteig

- Senecio chrysolepis Phil.

- Viola frigida Phil.

Over the period 1901-1903 the German bota-

nist Karl Reiche, also associated with the Museo

Nacional de Historia Natural, effected botanical

expeditions in the north of the country, including

the vicinity of Volcan Llullaillaco (Reiche, 1907),

leading to the following new taxa reported for the

first time within the area of park

Gayana Bot. 55(2). 1998.

-23*00'

q Erio

- 4822

OCEANO

(6 DA co

7 6739

;

PORTEDELO

UNLLAILLACO SUR

DE ARGENTIMA

REPUBLICA

FIGURE 1. Details of Llullaillaco National Park and its location in the II Region of Chile

(

inset). Source CONAF - January, 1994,

Flora of Llullaillaco National Park: ARROYO, M.T.K..,, ET AL.

- Artemisia copa Phil.

- Atriplex imbricata (Moq.) D.Dietr.

- Descurainea stricta (Phil.) Prantl ex Reiche

var. minutiflora (Phil.) O.E. Schulze

- Fabiana bryoides Phil.

- Fabiana denudata Miers

- Junellia digitata (Phil.) Mold.

- Moschopsis monocephala (Phil.) Reiche

- Phacelia pinnatifida Griseb ex Wedd.

- Senecio eriophyton J.Remy

- Senecio sundtii Phil.

- Senecio xerophilus Phil. var. xerophilus

- Triglochin concinnum Burtt Davy

- Urbania pappigera Phil.

Between 1923 and 1927 Erich Werdermann

(Werdermann 1927), later Director of Berlin Botani-

cal Garden, undertook an expedition in the high An-

des of Bolivia and northern Chile which resulted in

41 collections made in the present area of Llullailla-

co NP (Looser, 1960). Species registered for the first

time in the park by Werdermann include:

- Acantholippia deserticola (Phil.) Mold.

- Adesmia caespitosa Phil.

- Arenaria serpens Kunth

- Astragalus cryptobotrys 1.M.Johnst.

- Calandrinia compacta Barnéoud

- Catabrosa werdermannii (Pilger) Nicora et Rú-

golo

- Cistanthe minuscula (Añon) Peralta

- Cistanthe picta (Gillies ex Arn.) Carolin ex

Hershk.

- Colobanthus quitensis (Kunth) Bartling

Chaetanthera revoluta (Phil.) Cabrera

- Cristaria andicola Gay

- Gilia crassifolia Benth.

- E eremophila (Phil.) Burkart ex

- ad gossypifera (Gillies ex Benth.) V.

Grant

- Mancoa hispida Wedd.

- Montiopsis copiapina (Phil.) D.LFord

- Mulinum crassifolium Phil.

- Nicotiana ro (Griseb.) Millan

- Oxalis hypsophila

- Parastrephia RA (Meyen) Cabrera

- Phacelia setigera Phil. var. EG

- Pycnophyllum bryoides (Phil.) R

- Senecio hirtus

- Senecio AS Meyen et Walp.

- Sisymbrium philippianum 1.M.Johnst.

- Stipa chrysophylla E.Desyv.

- Viola llullaillacoensis W.Becker

In 1944, Carlos Muñoz, then Director of the

Museo Nacional de Historia Natural, explored se-

veral areas of the Atacama desert with the aim of

visiting sites explored previously by R. A. Philippi

(Muñoz « Pisano, 1950). Species collected within

the park boundary for the first time from a total of

32 collections made, include:

- Chaetanthera sphaeroidalis (Reiche) Hicken

- Deyeuxia eminens J.Presl

- Festuca deserticola Phil.

- Gentiana sedifolia Kunth

- Perezia atac is (Phil.) Reiche

ta (Graham) Hook.

- Nototriche auricoma (Phil.) A.W.Hill

- Nototriche clandestina (Phil.) A.W.Hill

- Senecio puchii Phil.

- Nicotiana

In 1992 G. Arancio, Universidad de La Sere-

na, made 23 collections within the park boundary,

discovering for the first time:

- Adesmia melanthes Phil.

- Adesmia occulta (R.E.Fries) Burkart

- Baccharis tola Phil. subsp. altiplanicola F.H.

Hellwig

- Deyeuxia crispa Rugolo et Villav.

- Festuca chrysophylla Phil.

- Stipa frigida Phil.

- Werneria pinnatifida J.Remy

In 1994 M. T. Kalin Arroyo and collabora-

tors, Universidad de Chile, made 82 collections

within the park boundary over a period of 3 days,

discovering the following species for the first time:

- Adesmia spinosissima Meyen

- Astragalus bustillosii Clos

- Chaetanthera minuta (Phil.) Cabrera

- Deyeuxia deserticola Phil.

- Eleocharis albibracteata Nees et Meyen ex Kunth

- Festuca werdermannii St.-Y ves

- Hypsela reniformis (Kunth) K.Presl

- Lycium humile Phil.

- Nicotiana longibracteata Phil.

- Opuntia atacamensis Phil.

- Ruppia filifolia (Phil.) Skottsb.

- Scirpus atacamensis (Phil.) Boeckeler

- Sisymbrium lanatum (Walp.) O.E.Schulze

Gayana Bot. 55(2), 1998.

Finally, in 1997, G. Arancio and F. Squeo,

Universidad de La Serena made 29 collections

within the park boundary, revealing the following

previously undiscovered species:

- Anatherostipa venusta (Phil.) Peñail.

- Baccharis tola Phil. subsp. tola

- Ephedra breana Phil.

- Lenzia chamaepitys Phil.

- Opuntia camachoi Espinosa

An undated specimen of Haplopappus rigi-

dus Phil. was collected at Río Frío by an unknown

collector. This specimen possibly was collected by

R. A. Philippi who states that it was seen only se-

veral occasions in the Atacama (Muñoz, 1960).

SPECIES ACCUMULATION ANALYSIS: A good fit

of the species accumulation data was obtained

with the Clench model (Figure 3; number of taxa

—S(1 MRARQT 4 PEO MALOIOACTOMUNYT

TANYA ARODO Y ARA

may be seen that the species accumulation curve

for Llullaillaco NP does not yet reach a clear

asymptote, as would be expected if the flora were

exhaustively known. Mathematical extrapolation

of the curve in Figure 3 gives an expected increase

in the total number of species to 110 with a dou-

bling of the collection effort.

ENDEMISM: Endemic taxa, according to three ca-

tegories of endemism are shown in Table IL The

park possesses 2 local endemics: Menonvillea fri-

gida (Phil.) Rollins and Deyeuxia robusta Phil.

Menonvilla frigida is a very distinctive species of

genus. It was found by us growing abundantly

at around 4700 m on the slopes of Volcan Llullai-

llaco, but would also appear to grow at lower ele-

vations (see description). Deyeuxia robusta is a

poorly known species, which perhaps, upon furt-

her study, will not prove different from one of the

other species of Deyeuxia in the park. A total of

13 taxa (14.3%) of the flora is endemic to Chile.

DISTRIBUTIONAL LIMITS: Twelve species reach

their northern limits in the park (Table HD. Seven-

teen species reach their southern limits in the park.

An additional 8 species reach their southern limits

immediately outside the park boundary. In all 35

taxa (38.5% of the flora) are represented by their

in the park. Relatively more species found in the

zonal vegetation reach their southern limits in the

park than for azonal bog habitats (Figure 4).

ADDENDUM

The following species referred to in Philippi

(1860) may prove to exist in the park.

Triglochin palustris L. “Lo vi en Aguas Profetas,

después en Zorras y en otras localidades semejan-

tes” Two specimens cited (SGO 37012; 45409)

(Muñoz, 1960) but it is unclear as to which locali-

ties they pertain (Muñoz, 1960).

Triglochin striatum R et P. “Lo vi más arriba de Zo-

rras” (Muñoz, 1960). Other localities are also cited

in relation to the single specimen (SGO 45410) ci-

ted.

Juncus articus var. andicola - was observed by

R.A. Philippi at 2900 m in Zorras, which would

place on the border of the park on the basis of ele-

vation. A single specimen (SGO 37866) is cited in

Muñoz (1960) as representative of the latter and

other localities.

DISCUSSION

Although botanists have visited the area of

Llullaillaco National Park since 1860, prior to this

study no comprehensive account of the flora of

Llullaillaco NP has been available (Muñoz et al.,

1996). Richter (1995) citing 19 species observed

in a vegetation transect on Volcán Llullaillaco, is

the only published reference to the botany of the

park. Interestingly, however, as early as 1925,

over half of the species known to occur in the park

had been collected, whereas as early as 1950, over

2/3 of the flora was known.

The 90 species (91 taxa) constitute a mixture

of zonal and Andean bog elements. Zonal ele-

ments include high Andean elements (e.g. Perezia

atacamensis, Chaetanthera sphaeroidalis, Stipa

igida, Fabiana denudata. Baccharis tola, Noto-

triche spp., Senecio sundtii, S. eriophyton, S.

puchii, S. xerophilus, Viola llullaillacoensis, Me-

nonvillea frigida, Mulinum crassifolium, Pya-

nophyllum bryoides, Parastrephia quadrangularis,

Lenzia chamaepitys) and species representative of

Flora of Llullaillaco National Park: ARROYO, M.T.K., ET AL.

the desert margin (Acantholippia deserticola, Atri-

plex imbricata, Chenopodium frigidum, Cistanthe

minuscula, Ephedra breana, Opuntia atacamensis,

O. camochoi, Haplopappus rigidus, Gilia crassifo-

lia, Hoffmannseggia eremophila, Ipomopsis gossy-

pifera). Most of the very depauperate high Andean

bog flora known for this latitude is found in the

park (e.g. Calandrinia compacta, Colobanthus

quitensis, Deyeuxia spp., Hypsela reniformis, Scir-

pus atacamensis, S. deserticola, Oxychloe andina,

Potamogeton strictus, Ruppia filifolia, Gentiana

sedifolia). Regionally, nevertheless, the park lacks

a reliable record for some of the more typical plant

species associated with salt lakes (e.g. Triglochin

striatum). Such species can be found in the Salar

de Pajonales, immediately to the south of the park

boundary. Inclusion of this salar in the park clearly

would increase its conservation coverage.

Known species richness for the park clearly

indicates an impoverished flora for a land area of

hectares in relation to other areas in the

Chilean Andes. For comparison, the high Andean

flora of the much smaller area of Laguna Grande

and Laguna Chica in the HI Región of Chile con-

tains 281 species (Arroyo et al., 1984). Monumen-

to Nacional El Morado in the mediterranean-type

climate Andes of central Chile supports 280 spe-

cies in area of 3009 hectares (Tellier et al.. 1994).

The alpine flora of Parque Nacional Torres del

Paine (242,242 ha; Muñoz et al., 1996) stands at

179 species in 94 genera (Arroyo et al., 1992),

with only a fraction of the park area supporting al-

pine vegetation. Nevertheless, the 90 species (91

taxa) recorded in Llullaillaco NP are within the

range expected for the hyper-arid Andes. An in-

tensive study of an altitudinal gradient slightly to

the north at latitude 24*S where precipitation is

slightly higher, revealed 77 species in 55 genera

and 30 families over a similar elevational range

(Arroyo et al., 1988).

While Llullaillaco NP is not overly rich in

species, it clearly plays a very important role in

protecting the limiting distributions of a very large

number of high Andean species. Anywhere in the

Andes it is very unusual to find such a high pro-

portion of species represented by their northern or

southern ANTE limits. This situation undoub-

tedly reflects the transitional climatic setting of

Liuilaillaco pa between the summer and winter

rainfall regimes. Here, effectively, many northern

po ments adapted to a summer ae

regime, and southern elements of mediterrane

influence adapted to a winter rainfall regime reach

ir southern and northern limits, respectively. In-

terestingly, proportionally more species reach their

southern distributional limits in the park than vice-

versa. The significance of this trend can only be

speculated upon at this time. Around twenty seven

species in the park are considered to be northern

Andean elements, while 23 are considered to be of

mediterranean affinity (Arroyo et al., unpublished

data), with the remaining species distributed in

both the mediterranean and subtropical sectors of

the Andes. With a predominance of northern sub-

tropical elements in the flora, relatively more sout-

hern limits could be expected. Alternatively, the

trend may simply reflect the fact that Llullaillaco

NP is located slightly south of the presumed 50-

50% summer-winter rainfall zone. That there are

relatively more northern summer rainfall than

southern mediterranean elements in the flora, sug-

gests the hypothesis of relictual distributions of

summer rainfall species. This hypothesis finds

strong support in the fact that summer rainfall ele-

ments tend to be more locally distributed in the

park and found in specialized habitats, such as

north-facing warm rocky slopes. This last pattern

is consistent with the recent findings of Grosjean

et al. (1995) who concluded, on the basis of lake

level studies, that precipitation rates between 22*-

24”S in the northern Andes of Chile were in-

tensified during the late Glacial and Holocene.

According this study, annual precipitation would

have been 400-500 mm higher than at present.

Species accumulation analysis indicated that

some new species records can indeed be expected

for the park over the 91 given here. Indeed, inclu-

ding the species given in the Addendum would

increase the number of taxa to 94. That few addi-

tional species have been revealed in several well

collected areas immediately outside the boundary

of the park (e.g. Cerro Guanaqueros to the north,

Quebrada de Zorras, region of Salar de Punta Ne-

gra, Sierra de Varas to the east, Salar de Pajonales,

south face of Cerro del León, and Laguna de Azu-

frera to the south-east, and Valle Sandon to the

south-west) indicates that new records are likely to

be relatively few. Future botanical exploration

should concentrate on the still poorly explored

areas of Cerro Aguas Calientes, the south-eastern

sector of Sierra de Varas, the dry pampas in the

north-west sector of the park, and the northern side

of Cerro del León, any small patches of high An-

dean bog in the north-eastern sector of the park,

Gayana Bot. 55(2), 1998.

and any small patches of salar vegetation that wed Presidential Science Appointment (Cátedra

might exist within the park boundaries. Presidencial de Ciencias -1997) (MTKA). Explo-

ration by the first author in 1994 benefited from

partial field support from the project “Climate

ACKNOWLEDGMENTS Change in the Arid Andes” (Bruno Messerli-

Martin Grosjean) financed by the Swiss National

Research supported by CONICYT Grant Science Foundation.

No. Biomas-S960016 and carried out on a Endo-

TABLE L. Precipitation data for the closest climatic stations to Llullaillaco National Park after Hajek £ di Castri

(1975). CIREN y Dirección General de Aguas (DGA).

Annual Summer

Station Lat. Long. Elevat. Yrs. Source Precipitation Precipitation (mm)

(m) mm

Peine 23418 6804 "W 2480 17 CIREN 2) 49.3

Socaire 2393 $ 67"52"W 3251 18 CIREN 45.6 54.5

Monturaqui 24%02'S 68"26"W 3450 7 A 58.5 90.0

Socompa 24%6"S 68”18'S 3915 8 CIREN 20.7 78.7

Catalina 25"14"S 6944 "W 2180 8 CIREN 0 0

Refresco 25"19'S 6952 W 1850 19 H£D 9:2 E,

TABLE HI. Endemic vascular plants in Llullaillaco National Park. A: entirely restricted in distribution to within the

boundary of the park; B: endemic to the Andes of the II Región; C: endemic to the Andes of Chile in a broader sense.

Menonvillea frigida (Phil.) Rollins

Deyeuxia robusta Phil.

Descurainea stricta (Phil.) Prantl ex Reich var. minutiflora (Phil.) O.E.Schulz

ia ata is Phil.

mint

Adesmia frigida Phil.

Baccharis tola Phil. subsp. altiplanicola F.H. Hellwig

Festuca werdermannii St.-Y ves

Nototriche auricoma (Phil.) A-W. Hill

Nototriche clandestina (Phil.) A.W. Hill

Opuntia camachoi Espinosa

Oxalis hypsophila Phil.

Senecio hirtus Cabrera

Viola llullaill is W. Becker

00 oOo0O!a»y-> >

Flora of Llullaillaco National Park: ARROYO, M.T.K., ET AL.

TABLE III. Species with northern or southern distributional limits in, or immediately outside the border of Llullai-

llaco National Park. N: northern limit in park; S: southern limit in park: N and S: northern and southern limits in

park; S-close: southern limit close to park bound:

Adesmia frigida Phil. N

haetanthera minuta (Phil.) Cabrera N

Festuca werdermannii St.-Y ve N

Montiopsis copiapina (Phil.) D.L Ford N

Montiopsis glomerata (Phil.) D.L For N

a rompe dy ) Reiche N

Senecio Ji JR N

Senecio pel Phil. N

Viola fri N

Viola Undalacocnt W.Becker N

Deyeuxia robusta N and S

Menonvillea frigida ¿Phi ) Rollins N and S

Adesmia caes, S

Adesmia melanthes Phil. S

Anatherostipa venusta (Phil.) Peñail. S

Astragalus cryptobotrys 1.M.Johnst S

Baccharis tola Phil. subsp. altiplanic ola F.H.Hellwig S

Chaetanthera revoluta (Phil.) Cabrera S

Descurainea stricta (Phil.) Prantl ex Reich var. minutiflora (Phil.) O.E.Schulz S

Fabiana denudata Miers S

Junellia pi (Phil.) Mold. S

Mancoa hisp S

Mischa! scale (Phil.) Reiche S

Nototriche auricoma (Phil.) A.W.Hill S

Senecio puchii Phil. S

Sisymbrium lanatum (Walp.) O.E.Schulz S

Urbania pappigera Phil. Ss

Adesmia occulta (R.E.Fries) Burkart S-close

Artemisia copa Phil. S-close

Deyeuxia crispa Rúgolo et Villav. S-close

Opuntia atacamensis Phil. S-close

Parastrephia que ciloril (Meyen) Cabrera S-close

Phacelia setigera Phil. var. Setigera S-close

Pycnophyllum bryoides (Phil.) Rohrb. S-close

Senecio xerophilus Phil. var. Xerophilus S-close

S 60

Percent of flora

annuals perennials subshrubs shrubs

Life forms

FIGURE 2. Life forms in the flora of Llullaillaco National Park. Perennials = perennial herbs. Facultatively annual

species are included in the annual category.

101

Gayana Bot. 55(2), 1998.

1998(n:39/)

100 F ha.

E 1975(n=66) x

80 Fr 1995(n=86)

o 60+

x 1875(n=16) 1950 (n=66)

le] AS

Lead 40 1925 (n=549)

20t =€ 1900 (n=16)

o A 1 L L L L L

0) 50 100 150 200 250 300

Collections

RE 3. Species accumulation curve for Llullaillaco National Park, based on fitting of the Clench Model to co-

io data. Data points are at 25 collection intervals. The number of taxa (species and 1 subspecies) accumulated

E N=70

ZE ES North

30F N=9]|] EJ] South

Percent of flora

total zonal azonal

FIGURE 4. Taxa reaching their northern or southern distributional limits in the Chilean Andes within the boundaries

of, or immediately outside (see methods) Llullaillaco National Park.

Flora of Llullaillaco National Park: ARROYO, M.T.K., ET AL.

CATALOG OF VASCULAR PLANTS

GYMNOSPERMAE

EPHEDRACEAE

1. Ephedra breana Phil.

Xerophytic shrub to ca. 1.5 m with stiff, yellow-green

stems; fruits fleshy, red. Exc.: Antofagasta: Llanos Ce-

rro de la Pena, 3900 m, Arancio «£ Squeo 10494 (ULS).

ANGIOSPERMAE: DICOTYLEDONEAE

CACTACEAE

2. Opun

duos ormúlll species, composed of bright green

ovoid or conical segments with reddish spines; flowers

yellow to brownish-yellow. Endemic to the II Región of

vas where it is found only at mid to high elevations in

the southern portion of the Región. Exs.:Antofagasta:

Volcán Llullaillaco, 4100 m, Arroyo et al. 94025

(CONC)

3. O. camachoi Espinosa

Cushion forming species, closely related to O. ataca-

mensis, but differing in its smaller size and grey-brown

spines. Endemic to the high Andes of Chile (24*5-

27%S). Exs.: Antofagasta: Pampa Las Carretas km

28.092, 3950 m, Arancio « Squeo 10436 (ULS)

CAESALPINIACEAE

4. Hoffmamseggia eremophila (Phil.) Burkart ex Ulib.

pe ea herb to 5 cm high. Flowers orange-red-

h. On sandy soils. Exs: Antofagasta: Cordillera Vol-

cán ed 4000 m, Werdermann 1014 (CONC):

Antofagasta: Volcán Llullaillaco, 4000 m, Arroyo et al.

94035 (CONC); pa Quebrada de la Pena,

3800 m, Arancio € Squeo 10467 (ULS); Antofagasta:

Llanos Cerro de la Pena, 3900 m, Arancio £ Squeo

10499 (ULS); Antofagasta: Base Cerro de la Pena, 3680

m, Arancio £ Squeo 10405 (ULS)

CALYCERACEAE

5. Moschopsis monocephala (Phil.) Reiche

Succulent, glaucous, rosette-forming perennial herb.

Flowers whitish. Exs.: Río Frío, Reiche w/o no. (SGO

057228); Antofagasta: Vega Zorra, 3150 m, Arancio

92338 (ULS); Antofagasta: Llullaillaco, camino a Mina

Esperanto, 3740 m, Muñoz C. 3806 (SGO); Folomcard

ta: Cordillera Volcán Llullaillaco, 4400 m

1016 (CONC); Antofagasta: Volcán Llullaillaco, 4500

m, Arroyo et al. 94009 (CONC).

6. Nastanthus caespitosus (Phil.) Reiche

Succulent, rosette-forming perennial herb. Flowers whi-

te. On edge of cushion bogs. Exs.: Río Frío, 3292 m,

Philippi w/o no. (SGO 43609, 57221); Antofagasta: Va-

lle de Río Frío, 3200 m, Muñoz C. 3891 (SGO); Anto-

fagasta: Río Fo, 3500 m, Werdermann 1028 (CONC).

CAMPANULACEAE

7. Hypsela dridiae (Kunth) K.Pres

Interweaving perennial herb. Flowers e tinged blue-

violet. In cus alta Dore Exs.: eii de Las

Zorritas, 4150 m, Arroyo et al. 94068 (CO

CARYOPHYLLACEAE

8. Arenaria serpens Kunth

Mat forming to interweaving perennial herb. Flowers

white. Common in cushion bogs. Exs.: Antofagasta: Ve-

ga Las Zorritas, 4200 m, Arroyo et al. 94054 (CONC);

Antofagasta: Cordillera Volcán Llullaillaco, 3800 m,

Werdermann 1018 (CONC); Antofagasta: Volcán Llu-

llaillaco, 3950 m, Arroyo et al. 94042 (CONC)

9. Colobanthus ee Bartling

Dwarf perennial herb. Flowers green. In cushion bogs.

Exs.: Antofagasta: Volcán Llullaillaco, 3800 m, Johns-

ton 6214 (S); Antofagasta: Volcán Llullaillaco, 3400

m, Werdermann 1452 (S).

10. Pycnophy cn. bryoides (Phil.) Rohrb.

ommon herb forming loose cushions. Flowers

inconspicuous, q, Bas: Apr Lla-

nos en camino Quebrada de Las Zorrritas, 4150 m

cio 248 (CONC); Antofagasta: Llullaillaco, camino a

Mina Esperanto, 3740 m, Muñoz C. 3804 (SGO); Anto-

fagasta: Cordillera Volcán Llullaillaco, 4300 m, Werder-

mann 1026 (CONC); Antofagasta: Volcán Llullaillaco,

4400 m, Arroyo et al. 94004 (CONC); Antofagasta: Ce-

rro del León, 4150 m, Arancio £ Squeo 10412 (ULS).

CHENOPODIACEAE

11. Atriplex imbricata (Moq.) D.Di

White-stemmed shrub up to 7Scm High: leaves with

glaucous borders. At lower elevations on desert border.

ExS.: Región del Llullaillaco, Reiche w/o no. (SGO

048299). (SGO 048301): Ari Cordillera Vol-

cán Llullaillaco, 3800 m, rmann 1003 (CONC):

Antofagasta: Faldeos S Ba ñ Quebrada Llullaillaco,

00 m, Arroyo et al. 94081 (CONC).

12. Chenopodium frigidum Phil.

Spreading annual herb with glaucous foliage. On sandy

sites. Rare. Exs.: Antofagasta: Volcán Llullaillaco, 4000

m, Arroyo et al. 94030 (CONC); Río Frío, Philippi w/o

no. (SGO 48176); Antofagasta: Valle de Río Frío, 3200

m, Muñoz C. 3859 (SGO)

Gayana Bot. 55(2), 1998.

COMPOSITAE

13. Artemisia copa Phil.

Aromatic shrub up to 60 cm high with whitish foliage.

Locally abundant on rocky north-facing slopes. Exs: Re-

gión del Llullaillaco, Reiche w/o no. (SGO 066433); An-

tofagasta: Vega Zorra, O m, Arancio 92340 (ULS);

Antofagasta: Volcán Llullaillaco, 4100 m, Arro

m, Werdermann 1007 (CONC); Antofagasta: Cerro del

León, 4240 m, Arancio € Squeo 10423 (ULS).

14. Baccharis tola Phil. subsp. altiplanicola F.H.Hellwig

Yellow-stemmed, dioecious, resinous shrub usuall y

>40 cm high. Leaves strongly dentate with 2 or more

the altiplano of northern Chile. Exs.: Antofagasta:

Quebrada de Las Zorritas, 3150 m, Arancio 243

(CONC); Antofagasta: Volcán Llullaillaco, 4100 m,

Arroyo et al. 94024 (CONC); Antofagasta: Faldeos S

de la Quebrada Llullaillaco, 3850 m, Arroyo et al.

94078 (CONC); Antofagasta: Llanos Cerro de la Pena,

900 m, Arancio $: Squeo 10493 (ULS).

15. B. tola Phil. subsp. tola

Differs from subsp. altiplanicola principally in its sma-

ller stature (< 40 cm) and leaves that are entire or with

1, or exceptionally 2 teeth on each side of the lamina.

Exs: Antofagasta: Base Cerro de la Pena, 3680 m,

Arancio $: Squeo 10402 (ULS).

16. Chaetanthera minuta (Phil.) Cabrera

Decumbent annual herb, covered with a white tomen-

tum. Heads radiate, white. On sandy soils. Exs.: Anto-

fagasta: Volcán Llullaillaco, 4000 m., Arroyo et al.

94033 (CONC).

17. Ch. revoluta (Phil.) Cabrera

Inconspicuous perennial herb growing at base of

rocks. Heads radiate, white. Common. Exs: Antofagas-

ta: Vega Zorra, 3150 m, Arancio 92337 (ULS); Anto-

fagasta: Llullaillaco, camino a Mina anto, 3740

m, Muñoz C. 3809 (SGO); Antofagasta: Cordillera

Volcán Llullaillaco, 4200 m, Werdermann 1020

(CONC); Antofagasta: Volcán Llullaillaco, 4500 m,

Arroyo et al. 94013 (CONC); Antofagasta: Faldeos S

de la Quebrada Llullaillaco, 3850 m, Arroyo et al.

94079 (CONC); Antofagasta: Base Cerro de la Pena,

3680 m, Arancio 4 Squeo 10407 (ULS).

camino a Mina Esperanto, 3740 m, Muñoz C. 3808

SGO): Antofagasta: Volcán

yo et al. 94017 (CONC); Antofagasta:

co, 4350 m, Arroyo et al. 94003 (CONC); Antofagasta:

S de Llullaillaco, 4800

104

19. Haplopappus rigidus Phil.

Resinous, yellow-stemmed shrub. Heads radiate, ye-

llow. The collection data upon which inclusion of this

species in the park is based are very limited. Its pre-

sence needs further verification. Exs.: Antofagasta:

Río Frío, w/o collector and no. (BM).

20. Parastrephia quadrangularis (Meyen) Cabrera

Small shrub to 50 cm with tightly imbricate greenish-

yellow leaves; heads discoid, yellow. Mostly restricted

to warmer rocky slopes. Exs: Antofagasta: Valle de

Zorritas, 3720 m, Muñoz C. 3830 (SGO); Antofagasta:

Cordillera Volcán Llullaillaco, 4200 m, Werdermann

1012 (CONC); Antofagasta: Volcán Elullaillaco, 4150

m, Arroyo et al. 94026 (CONC).

21. Perezia atacamensis (Phil.) Reiche

Perennial herb to ca. 30 cm. Heads large, radiate, var-

ying in color from pinkish to bluish. On rocky sites.

Exs.: Antofagasta: Llullaillaco, camino Mina Esperanto,

40 m, Muñoz C. 3805 (SGO); Antofagasta: Volcán

Llullaillaco, 4500 m, Arroyo et al. 94014 (CONC).

22. Senecio chrysolepis Phil.

Erect, lightly tomentose subshrub with oblong leaves, to

20 cm. Heads discoid, yellow. Exs.: Antofagasta: Vol-

cán Llullaillaco, 4500 m, Arroyo et al. 94007 (CONC);

Antofagasta: Valle del Río Frío, 3200 m, Muñoz C.

3893 (SGO): Antofagasta: Río Frío, Philippi w/o no.

(SGO 060675); Antofagasta: Sierra de Varas, Punta del

Viento, 4200 m, Werdermann 1040 (CONC).

23. $. eriophyton J.Remy

Densely branched, lanose aromatic roundish dwarf sh-

rub to 20 cm with conspicuously dentate-curled leaves.

Heads radiate, yellow. Exs.: Antofagasta: Región del

Llullaillaco, 4200 m, Reiche w/o no. (SD; Antofagasta:

Región del Llullaillaco, 4700 m, Reiche w/o no. (SI).

24. S. hirtus Cabrera

Small densely branched shrub with sessile, clasping

glandular-pubescent leaves, to 30 cm igh. Heads ra-

diate, yellow. Endemic to the high Andes of Chile

(25-288). Exs.: Antofagasta: Sierra de Varas, Punta

del Viento, 4000 m, Werdermann 1036 (CONC).

25. S. puchii Phil.

Tightly branched low subshrub to 10 cm with succulent

glabrous dentate leaves. Heads discoid, yellow. Exs.:

Antofagasta: Faldeos Volcán Llullaillaco, Zorritas, 3800

m, Muñoz C. 3799 (SGO); Antofagasta: Volcán Llullai-

llaco, 4600 m, Arroyo et al. 94001 (CONC).

26. $. scorzoneraefolius Meyen et Walp.

Lax, rhizomatous perennial with conspicuously long-

linear leaves. Heads discoid, dirty white. Exs.: Antofa-

gasta: Cordillera Volcán Llullaillaco, 4200 m,

Werdermann 1029 (CONC).

27..S. sundtii Phil.

Low, tomentose subshrub. Head discoid, yellow. Exs.:

Región del Llullaillaco, Reiche w/o no. (SGO 062330);

Flora of Llullaillaco National Park: ARROYO, M.T.K.., ET AL.

Región del Llullaillaco, 4700 m, Reiche w/o no. (SD).

28. $. xerophilus Phil. var. xerophilus

Erect tomentose subdichotomously branched subshrub

to 25 cm. Heads discoid, yellow. Exs.: Región del Llu-

laillaco, Reiche w/o no. (SGO 062354); Antofagasta:

Vega Zorra, 3150 m, Arancio 92334 (ULS); Antofagas-

: Faldeos Volcán Llullaillaco, Zorritas, 3800 m, Mu-

ñoz C. 3801 (SGO); Antofagasta: Cordillera del Volcán

Llullaillaco, 4200 m, Werdermann 1037 (CONC); An-

tofagasta: Región del Llullaillaco, 4200 m, Reiche w/o

no. (SI); Antofagasta: Volcán Llullaillaco, 4100 m,

Arroyo et al. 94020 (CONC); Antofagasta: Volcán Llu-

llaillaco, 4700 m, Zoellner (CONC 49070)

29. Werneria pinnatifida J Remy

Rhizomatose perennial herb with fleshy pinnatisect roset-

te forming leaves. Heads discoid, white. Exs.: Antofagas-

ta: Quebrada de las Zorritas, 4150 m, Arancio 335

(CONC)

CRUCIFERAE

30. Descurainia stricta (Phil.) Prantl ex Reiche var.

minutiflora (Phil.) O.E. Schulz Annual to biannual,

kr Entro leaves. Flowersminute, yellow

o Chile. Exs.: Región de Llullaillaco, Reiche w/o

no. Per sGO 071497).

31. Mancoa hispida Wedd

Small, hispid perennial herb. Exs.: Antofagasta: Vol-

cán Llullaillaco, 3800 m, Werdermann 1008 (CONC).

32. Menonvillea frigida (Phil.) Rollins

Perennial herb with aero brown-reddish

stems and leaves and showy white flowers. Grows in

rock crevices. Locally common. A rare Seta endemic,

thus far only known from high elevations in Parque

Nacional Llullaillaco, and thus endemic to the II Re-

gión of Chile. Exs.: Antofagasta: Volcán Llullaillaco,

4700 m, Arroyo et al. 94006 (CONC); Antofagasta:

Faldeos Volcán Llullaillaco, camino Mina Iris, 3920

m, Muñoz C. 3798 (SGO); poa Río Frío, Phi-

lippi w/o no (SGO 063964

33. Sisymbrium lanatum (Walp.) O.E.Schulz

Perennial herb, woody at the base, to ca. 30 cm, with

glaucous tomentose to glabrous leaves. Flowers ye-

llow. Exs.: Antofagastá: Vega Las Zorritas, 4150 m,

Arroyo et al. 94076 (CONC); Antofagasta: Volcán

Llullaillaco, 4000 m, Arroyo et al. 94031 (CONC).

34. $. philippianum 1.M.Johnst.

Subs cm branched at the base with tomento-

se stems and tomentose to glabrous leaves. Flowers

yellow to purplish. Exs.: Antofagasta: ro del

Volcán Llullaillaco, 3500 m, Werdermann 1457 (Pho-

to: BAA ex B).

GENTIANACEAE

35. Gentiana sedifolia Kunth

Small, glabrous annual to probably biannual herb. Flo-

wers blue, with yellow centers. Grows in cushion

bogs. Exs.: Antofagasta: Valle de Río Frío, 3200 m,

Muñoz C. 3862 (SGO).

HYDROPHYLLACEAE

36. Phacelia cumingii (Benth.) A.Gra

Small erect o prostrate glandular-viscid annual herb

with reddish stems, o leaves and cymose inflo-

rescences. Flowers white. Common on sandy re

Exs.: Antofagasta: Volcán Llullaillaco, 4000 m, Arro

et al. 94029 (CONC); Antofagasta: Valle de Río Frio,

3200 m, Muñoz C. 3860 (SGO); Río Frío, Philippi w/o

no. (SGO 42186, 42188, 54299) Antofagasta: Base Ce-

rro de la Pena, 3680 m, Arancio £ Squeo 10403

(ULS).

37. Ph. pinnatifida Griseb. ex Wedd.

Erect to decumbent robust perennial herb with strongly

pinnatifid leaves. Flowers blue, in dense scorpioid cy-

mes, with exerted styles and stamens. Exs.: Cajón de

Llullaillaco, Reiche w/o no. (SGO 054313).

38. Ph. setigera Phil. var. setigera

Robust, annual to perennial glandular-viscid herb with

lobulate leaves and bluish-purple flowers with included

styles and stamens. Exs.: Antofagasta: Cordillera Vol-

cán Llullaillaco, 4000 m, Werdermann 1006 (CONC).

MALVACEAE

39. Cristaria andicola Gay

Perennial herb with yellowish stems and bright green

pubescent leaves. Flowers pink. Common, in sandy

soils. Exs.: Antofagasta: Quebrada de Las Zorritas,

50 m, Arancio 241 (CONC); Antofagasta: Cordillera

de Llullaillaco, 4700 m, Werdermann w/o no. (CONC

23195); Antofagasta: Volcán Llullaillaco, 4000 m,

Arroyo et al. 94028 (CONC); Antofagasta: Cerro del

León, 4240 m, Arancio £ Squeo 10418 (ULS); Antofa-

gasta: Pampa Las Carretas, 3950 m, Arancio «£ Squeo

10440 (ULS): Po Pampa Las Carretas km

28,092, 3950m, Arancio £ Squeo 10435 (ULS); Anto-

fagasta: Cuesta Casa de Lata, 3700 m, Arroyo et al.

94100 (CONC): Antofagasta: Llanos Cerro de la Pena,

3900 m, Arancio £ Squeo 10498 (ULS); Antofagasta:

Base Cerro de la Pena, 3680 m, Arancio «£ Squeo

10404 (ULS)

40. Nototriche auricoma (Phil. a A-W.Hill

tome al herb to ca. 3 cm dia-

Antofagasta: Región de Llullaillaco, Muñoz C. 3796

(SGO):; Volcán Llullaillaco, 4500 m, Arroyo et al.

105

Gayana Bot. 55(2), 1998.

41. N. clandestina (Phil.) A.W.Hill

Compact, cushion forming perennial herb, densely co-

vered yellowish-white tomentum. Flowers white. Grows

close to the upper vegetation limit. Endemic to the high

Andes of Chile (24*5-26%S). Exs.: Región de Llullailla-

co, Muñoz C. 3797 (SGO); Antofagasta: Volcán Llu-

llaillaco, 4500 m, Arroyo et al. 94018 (CONC).

OXALIDACEAE

42. Oxalis hypsophila Phil.

Caespitose perennial herb. Flowers yellow. Endemic to

the high Andes of Chile (23-31%S) Exs.: Antofagasta:

Cordillera Volcán Llullaillaco, 4000 m, Werdermann

1010 (CONC)

PAPILIONACEAE

43. Adesmia passe Phil.

Woody, flat, ushion-forming species, with short thick

par ueda spines; cushions pere immersed in sand;

flowers yellow-orange, rt shoots. Exs.: An-

tofagasta: Faldeos Volcán pus Zorritas, 3800 m,

Muñoz C. 3803 (SGO); Antofagasta: Cordillera Volcán

Llullaillaco, 4000 m, Werdermann 1025 (SI, S)

44. A. erinacea Phil.

Small glandular shrub forming open cushions with gree-

nish-white, densely interwoven 3-6 bifurcate Spines.

Flowers yellow, generally produced on spines. Exs.:

Antofagasta: Zorras, Desierto de Atacama, 3500 m, Phi-

(CONC); Antofagasta: Valle de Zorritas, 3720 m, Mu-

ñoz C. 3828 (SGO); Antofagasta: Volcán Llullaillaco,

rroyo et al. 94015 (CONC); Antofagasta:

Volcán. Llullaillaco, 4150 m, Arroyo et al. 94027

(CONC); Antofagasta: Volcán Llullaillaco, 4200 m,

Baumann 286 (CONC).

45. A. frigida Phil.

Small caespitose glutinose-villose subshrub with 1-2

branched spines. Flowers yellow. E to the high

Andes of Chile (25"-27%S). Exs.: pasmado Taltal,

Punta del Viento, 4200 m, Werdermann 1023 (CONC):

Río Frío, Philippi w/o no. (SGO 40179, 50241).

46. A. melanthes Phil.

Erect shrub to Im with thick reddish-brown branches

and spines. Leaves very small, borne on distant short-

shoots. Flowers yellow. Exs.: Antofagasta: Quebrada

de las Zorritas, 4150 m, Arancio 251 (CONC).

47. A. occulta (R.E.Fries) Burkart

Flat, hard, woody, glandular-pilose cushion forming

Ape, partially immersed in substrate, with browni

tofagasta:

92242 (ULS):; Antofagasta: Volcán 'Lltullsllaco, 4400

m, Arroyo et al. 94005 (CONC): Antofa ta: Volcán

Llullaillaco, 4300 m, Arroyo et al. 94023 (CONC).

48. A. spinosissima Meyen

Erect shrub to 1 m, with reddish brown stems and thick,

yellowish brown to white spines. Leaves borne on short-

shoots. noe abundant on sunny north-facing slopes and

in dry water courses. Flowers yellow. Exs.: Antofagas-

ta: Volcán: Llullaillaco, 4000 m, Arroyo et al. 94022

(CONC); Antofagasta: Subida de Río Frío a Aguas Ca-

lientes, 3500 m, Arroyo et al. 94083 (CONC).

49. Astragalus bustillosii Clos

Slender, glabrous perennial herb with pale blue flo-

wers. Á cushion bog species. Exs.: Antofagasta: Vega

Las Zorritas, 4150 m, Arroyo et al. 94067 (CONC).

. A. cryptobotrys 1.M.Johns

Perennial herb forming pra knit cushions partially

immersed in substrate, with brownish-grey pubescent

foliage and pinnate leaves. Flowers bluish-white. Exs.:

Antofagasta: Llanos en camino a Quebrada de las Zo-

rritas, 4450 m, Arancio 635 (CONC); Antofagasta:

Sierra de Varas, Punta del Viento, 4000 m, Werder-

mann 1034 (CONC).

POLEMONIACEAE

51. Gilia crassifolia Benth.

Robust viscid annual herb to ca. 20 cm. Flowers blue

to pan phd: white. Mostly on desert border of pa

Exs.: Antofagasta: Taltal, Punta del Viento, 3900 m

Werdermann 1030 (CONC).

52. Ipomopsis gossypifera (Gillies ex Benth.) V.Grant

Robust puberulent-lanose annual herb with somewhat

succulent entire to pinnatisect leaves. Flowers in heads

separated from the leaves; corolla rose-violet. Exs.:

Antofagasta: Cordillera Volcán Llullaillaco, 4000 m,

Werdermann 1041 (CONC); Antofagasta: Volcán Llu-

laillaco, 4000 m, Arroyo et al. 94039 (CONC).

PORTULACACEAE

53. Calandrinia compacta Barnéoud

Rhizomatose perennial herb forming extensive mats,

with succulent leaves q. white flowers on reddish pe-

tioles hidden among leaves. Common on edge of cus-

hion bogs. Philippi (1860) described Calandrini

occulta, which he considered to be similar to C. com-

pacta and C. caespitosa, based on cd collected

from the sa area of Río

her part of quebrada Zorras, finding only 1 species of

Calandrinia in cushion bog. Philippi's C. occulta is

ery ;

Werdermann 1035 (CONC).

o Cistanthe minuscula os Peralta

annual f a few b long at-

Flora of Llullaillaco National Park: ARROYO, M.T.K., ET AL.

tenuate, spathulate leaves. Flowers in simple or double

spikes: petals white to lilac-purple. Exs.: o

Cordillera Volcán Llullaillaco, 3800 m, Werdermann

1015 (CONC); Antofagasta: Río Frío, sd m, Werder-

mann 1033 (CONC).

55. C. picta (Gillies ex Ar. ) Carolin ex Hershk.

, SO!

times tinged red; sepals wide, with conspicuous

red-black markings; petals white. Exs.: Di sta:

Cordillera Volcán Llullaillaco, 3800 m, Werdermann

1038 (CONC); Antofagasta: Volcán oe 4000

m, Arroyo et al. 94038 (CONC).

56. Lenzia chamaepitys Phil.

Diminutive perennial herb to 4 cm with closely ad-

pressed, oda pots and slender roots. Flo-

wers yellow ge, e, borne at tips of stems. A

rare socia tn in park, alo found on gravelly subs-

trates. Exs.: Antofagasta: Base Cerro de la Pena, 3680

m, Arancio £ Squeo 10406 (ULS).

57. Montiopsis copiapina (Phil.) D.I.Ford

Spreading, densely sericeo-pubescent perennial herb

with thick roots. Flow

mose racemes. Ens: Antofagasta

Quebrada de las Zorritas, 4150 m, Arancio 246

pri sb Antofagasta: Vega Zorra, 3150 m, Arancio

36 (ULS); Antofagasta: Cordillera Volcán Llullailla-

CO, pote Werdermann 1021 (CONC).

58. M. glomerata da ) D.L Ford

Small, prostrate annual herb with pubescent oblanceola-

white-pubescent calyx, petals small magenta to white.

Exs.: Río Frío, 3353 m, Philippi w/o no. (SGO 48522).

se 2d NA (Phil. ) D.LFord

with pubescent, linear leaves. Calyx

ma le “glandular nn flowers inconspi-

orescences. Exs.: Antofa-

ñoz C. 3861 (SGO); Antofagasta: Río Frío, Philippi w/o

no. (SGO 048661).

RANUNCULACEAE

60. Ranunculus cymbalaria Pursh f. exilis (Phil.) Lourteig

Semi-aquatic, glabrous rhizomatous perennial herb

y 1 leaves. Flowers yellow with shiny,

waxy petals. Grows only in cushion bogs, preferen-

tially at the edge of water courses or standing pools.

Exs.: Valle Zorras, 3351 m, Philippi w/o no. (SGO

.4200 m, Arroyo er al. 94053 (CONC).

SOLANACEAE

61. Fabiana bryoides Phil.

Stout shrub to ca. 50 cm, with twisted, dirty green, coral-

like branches covered in scale-like leaves. Flowers white

to lilac, very ommon i arm sunny

slopes. Exs.: Región del Llullaillaco, Reiche w/o no.

Zorra, pa m, Aran-

cio 92250 (ULS); Antofagasta: Quebrada de las Zorritas,

50 m, Arancio 639 (CONC); Anota: Valle de

Zorritas, 3720 m, Muñoz C. 3829 (SGO): Antofagasta:

Volcán Llullaillaco, 4150 m Paca et al. 94021

aaa Antofagasta: Cordillera Llullaillaco, 4000 m,

rmann w/o no. (CONC ció Antofagasta: Cor-

pios de Llullaillaco, 4000 m, Werdermann w/o no.

(CONC 22149); Antofagasta: Cerro del León, 4150 m,

Arancio «£ Squeo 10410 (ULS); Antofagasta: Llanos Ce-

rro de la Pena, 3900 m, Arancio « Squeo 10496 (ULS).

62. Fabiana denudata Miers

trongly resinous shrub to ca. 1 m, with somewhat si-

nuous cti o eo and small, distant lea-

amis] ite. Locally distributed. Exs.:

Región del Lisllilaco: Reiche w/o no. (SGO 055649);

Antofagasta: Cerro Zorritas, 4450 m, Arancio 252

(CONC); Antofagasta: Cordillera Volcán Llullaillaco,

4000 m, Werdermann 1005 (CONC); Antofagasta: Fal-

deos S de la Quebrada Llullaillaco, 3850 m, Arroyo et

al. 94077 (CONC); Antofagasta: Llullaillaco, 4000 m,

w/o collector (SGO 55640).

63. ea humile Phil.

abrous subshrub with greyish branches, for-

2. a dense cushion. Flowers white. Restricted to the

drier edges of cushion saline soils. Exs.: Anto-

fagasta: Volcán Llullaillaco, 3950 m, Arroyo et al.

94040 (CONC).

64. Nicotiana acuminata (Graham) Hook.

i ing annual herb with viscid stems, and

distant, ovate, acuminate leaves. Flower large (5-9 cm

long) greenish with purplish lines on corolla; white within.

Exs.: Antofagasta: Valle de Río Frío, 3200 m. Muñoz C.

3895 (SGO).

65. Nicotiana longibracteata Phil.

Stout, several-stemmed and densely leaved, pubescent to

viscid annual herb with long-acuminate leaves. Flowers

e. (ca. less than 2 2 - Cm long) : pame Poli cru)

crikhed A

nara

e: 85 HP A, 1ne.

O X

rs 3 BLIQASAS

rate form of N. longibracteata, but that further study is

required to ascertain this possibility. This last possibility

po mora recognizing the name N. frigida, which was des-

cribed before N. arsapristans Further work on these

species necessary. Exs.: : Volcán Llullailla-

co, 4000 m, Arroyo et al. unan a

b.) Millán

with conspicuous abs narrow

wers greenish-white. Common in sandy soils. Exs.:

107

Gayana Bot. 55(2), 1998.

Antofagasta: Cordillera Volcán Llullaillaco, 3800 m,

Werdermann 1039 (CONC); Antofagasta: Pampa Las

Carretas km 28,090, 4030 m, Arancio £ Squeo 10433

(ULS); Antofagasta: Cuesta Casa de Lata , 3700 m,

Arroyo et al. 94084 (CONC); Antofagasta: Querida

de la Pena, 3800 m, Arancio 8 Squeo 10466 (ULS):;

Antofagasta: Llanos Cerro la Pena, 3900 m, Arancio 4

Squeo 10500 (ULS)

UMBELLIFERAE

67. Mulinum crassifolium Phil.

Softly spinose, semi-hemispheric subshrub with bright

yellow-green, somewhat succulent leaves and whitish

older stems. Flowers inconspicuous, greenish-yellow.

Exs.: Antofagasta: Quebrada de las Zorritas, 4450 m,

Arancio 640 (CONC); Antofagasta: Vega Zorra, 3150

m, Arancio 92249 (ULS); Antofagasta: Faldeos Vol-

cán Llullaillaco, Zorritas, 3800 m, Muñoz C. 3800

( ); Antofagasta: Llullaillaco, camino a Mina Es-

peranto, 3740 m, Muñoz C. 3807 (SGO); cp

Cordillera Volcán Llullaillaco, 4300 m, Werdermann

1009 (CONC); Antofagasta: Volcán Llullaillaco, 4500

m, Arroyo et al. 94010 (CONC)

VERBENACEAE

68. Acantholippia deserticola (Phil.) Mold.

branc ngly aromatic, spiny, white-

hispid shrub to ca. 1 m with small adpressed scale-like

leaves. Flowers subsessile, lilac. Mostly at lower ele-

vations in park on desert margin. Exs.: o

Cordillera Volcán Llullaillaco, 3500 m,

1024 (CONC); Antofagasta: Faldeos S ce la Quebrada

Llullaillaco, 3500 m, Arroyo et al. 94082 (CONC).

69, Junellia En (Phil.) Mold.

Prostrate hrub forming isolated patches to

ca. 3-5 cm, with cree cecilia .. rust-coloured, lo-

se bark. Flowers abundant, sweetly fragrant, dark pink.

Exs.: Región del Llullaillaco, pa Reiche w/o

no. (SGO 054781); Antofagasta: Faldeos Volcán Llu-

llaillaco, Zorritas, 3800 m, Muñoz C. 3802 (SGO).

70. Urbania pappigera Phil.

eri shortly branched subshrub forming flat, ¡ in-

ONspicuous, pink. A

common species at mid-high elevations in park. Exs.:

Antofagasta: Llullaillaco, camino a Mina Esperanto,

3740 m, Muñoz C. 381 di

ra Volcán Llullaillaco, 4500 m, Werdermann 1019

(CONC); Antofagasta: Volcán Llullaillaco, 4600 m,

Arroyo et al. 94012 (CONC); Antof:

; Antofagasta: Pampa Las Carretas

28,092. 3950 m, Aracio de Squeo 10437 (ULS).

108

VIOLACEAE

71. Viola frigida Phil.

Rosette-forming annual herb. Flowers bluish-white.

Exs.: Río Frío, 3231 m, Philippi w/o col. (SGO 052337).

72. Viola llullaillacoensis W.Becker

Inconspicuous, rosette-forming annual herb, leaves

brownish, elliptic with ciliate margens. Flowers light

with darker lines. Typically occurrs in sandy soils.

A rare species, endemic to the high Andes A Chile (24*-

26"S). Exs.: Antofagasta: Vega Las Zorritas, 4150 m,

Arroyo et al. 94075 (CONC); poes Cordillera

Volcán Llullaillaco, 4000 m, Werdermann 1017

(CONC); Antofagasta: Volcán Llullaillaco, 4000 m,

Arroyo et al. 94034 (CONC); pr a eee de

las Zorritas, 4200 m, Baumann 289 (CONC

ANGIOSPERMAE: MONOCOTYLEDONEAE

CYPERACEAE

73. Carex maritima Gunnerus

Rhizomatose perennial herb. Restricted to high eleva-

tion cushion bogs. Exs.: Valle Zorras, Philippi w/o no.

PsP 046175); Antofagasta: Vega Las Zorritas, 4200

m, Arroyo et al. 94052 (CONC); Antofagasta: Vega

Las Zorritas, 4150 m, Arroyo et al. 94073 (CONC).

74. Eleocharis albibracteata Nees et Meyen ex Kunth

Caespitose, a perennial herb. Restricted to

cushion bogs. : Antofagasta: Vega Las Zorritas,

4150 m, Arroyo et 5 94069 (CONC); Antofagasta: Ve-

ga Las Zorritas, 4150 m, Arroyo et al. 94072 (CONC).

75. Scirpus atacamensis (Phil.) Boeckeler

Perennial herb forming large, tight cushions. Typically

found at the drier ados of cushion bogs, where common.

Exs.: Antofagasta: Vega Las Zorritas, 4200 m, Arroyo

et al. 94065 (CONC); Antofagasta: Vega Las Zorritas,

4150 m, Arroyo et al. 94071 (CONC); Antofagasta: Ve-

ga Las Zorritas, 4200 m, Arroyo et al. 94064 (CONC);

Antofagasta: Volcán Llullaillaco, 3950 m, Arroyo et al.

94041 (CONC), 94043 (CONC); 94045 (CONC).

GRAMINEAE

76. deba venusta (Phil.) Peñail.

Perennial grass to 30 cm with rigid convolute glabrous

yellow cai pa open; aristas long, reddish and

curved. Exs.: Antofagasta: Llanos Cerro de la Pena,

3900 m, Arancio de Squeo 10495 (ULS).

77. Catabrosa werdermannii (Pilger) Nicora et Rúgolo

Slender, inconspicuous perennial species forming lose

mats. Restricted to alpine bogs. Exs.: Antofagasta: Valle

de Río Frío, 3200 m, Muñoz C. 3868 (SGO); Antofagas-

ta: Taltal, Río Frío, 3500 m, Werdermann 1027 (CONC).

78. Deyeuxia crispa Rúgolo et Villav.

Flora of Llullaillaco National Park: ARROYO, M.T.K., ET AL.

nnial grass forming isolated clumps to 15 cm, with

pia twisted yellow leaves. Uncommon. Typically

asta: Cerr

Zorritas, 4450 m, Arancio 244 (CONC); Antofagasta:

Volcán Llullaillaco, 4400 m, Arroyo et al. 94002

79. Deyeuxia deserticola Phil.

Perennial grass forming isolated tufts to 30 cm gro-

wing on large cushions, with yellowish-grey, rigid lea-

ves. Restricted to high alpine bogs. Exs.: Antofagasta:

Vega Las Zorritas, 4200 m, Arroyo et al. 94056

(CONC), 94057 (CONC), 94058 (CONC).

80. Deyeuxia eminens J.Presl

Stout perennial species with junciforme leaves for-

bogs. Exs.: Antofagasta: Valle de Zorras,

3500 m, Muñoz C. 3819 (SGO); Antofagasta: Quebra-

da de Zorritas, 4150 m, Baumann 299 (CONC); Anto-

fagasta: Vega Las Zorritas, 4200 m, Arroyo et al.

94066 (CONC); Antofagasta: Volcán Llullaillaco,

3930 m, Arroyo et al. 94049 (CONC), 94050 (CONC).

81. Deyeuxia robusta Phil.

A poorly known species. Further study might show

that this species is the same as the former. Perennial

herb. Endemic to Chile.Exs.: Valle de las Zorras

2926-3048 m, Philippi w/o no. (SGO 068418).

82. Festuca chrysophylla Phil.

Glabrous perennial tussock species with rigid leaves

and glabrous sheaths. Apparently not common in park.

Exs.: Antofagasta: Llanos en Quebrada de las Zorritas,

4450 m, Arancio 636 (CONC)

83. Festuca deserticola Phil.

Caespitose perennial species to ca. 35 cm. Occasional

toward the desert margin. Exs.: Antofagasta: Valle de

Río Frío, 3200 m, Muñoz C. 3865 (SGO).

84. Festuca werdermannii St.-Y ves

Perennial species with greyish foliage forming small tus-

socks. At edge of cushion bogs. Endemic to the high Andes

of Chile (24”-29*S) Exs.: Antofagasta: Vega Las Zorritas,

4200 m, Arroyo et al. 94061 (CONC), 94062 (CONC).

85. Puccinellia frigida (Phil.) LM.Johnst.

= O species growing at drier edge of

cushion bogs. Exs.: Valle Río Frío 3292 m, Philippi w/o

no. (SGO 037544, 63510,62606): Antofagasta: Vega

Las Zorritas, 4200 m, Arroyo et al. 94060 (CONC),

94063 (CONC); Antofagasta: Taltal, Cordillera Volcán

Llullaillaco, 3500 m, We E

tofagasta: Volcán Llullaillaco,

(SGO); Antofagasta: Volcán Llullaillaco, 3950 m, Arro-

yo et al. 94046 (CONC).

86. Stipa chrysophylla E.Desyv.

Caespitose perennial species with erect yellow rigid

leaves to ca. 45 cm. Panicle contracted, usually exser-

ted above leaves. Exs.: Antofagasta: Cordillera Volcán

Llullaillaco, 4500 m, Werdermann 1013 (CONC).

e Dear frigida Phil.

nnial species with narrow rigid yellow leaves to ca.

300 cm. Panicle narrow, violet, not strongly exserted

el Las Zin itas,

4450 m, Arancio 637 (CONC), 638 (CONC); Antofa-

gasta: Volcán Llullaillaco, 4500 m, Arroyo et al. 94011

(CONC); Antofagasta: Cerro del León, 4150 m, Arancio

Arancio £ Squeo 10439 (ULS);

Antofagasta: Cuesta Casa de Lata, 3700 m, Arroyo et al.

94099 (CONC); Antofagasta: Quebrada de la Pena, 3800

m, Arancio « Squeo 10464 (ULS); Antofagasta: Base Ce-

rro de la Pena, 3680 m, Arancio £ Squeo 10399 (ULS).

JUNCACEAE

88. Oxychloe mt es

rming hard. el ted hi with

rigid, leaves. Fruits pan at maturity. Dominant cushion

species in alpine bogs. Exs.: Antofagasta: Valle de Zo-

rras, Philippi w/o no. (SGO 063055); Valle de Zorritas,

3500 m, Muñoz C. 3821 (SGO); Antofagasta: Vega Las

Zorritas, 4200 m, Arroyo et al. 94055 (CONC); Antofa-

gasta: Valle de Río Frío, 3200 m, Muñoz C. 3867 (SGO).

JUNCAGINACEAE

89. Triglochin concinnum Burtt Davy

Slender perennial herb often forming extended mats,

with short, carnose leaves, brownish stems, and erect,

contracted inflorescences partially exserted above the

leaves. Typically found in alpine cushion bogs. Exs.:

Prov. de Antofagasta, Llullaillaco, Reiche w/o no. (SGO

045420); Antofagasta: Cordillera Volcán Llullaillaco,

3500 m, Werdermann 1011 (CONC); Antofagasta: Vol-

cán Llullaillaco, 3950 m, Arroyo et al. 94044 (CONC).

POTAMOGETONACEAE

90. Potamogeton strictus Phil.

Aquatic perennial herb with submerged elongate cilin-

drical stems and filiform, membranous leaves; flowers

disposed in small spikes. In pools in high alpine bogs.

Exs.: Valle de Zorras, 3261 m, Philippi w/o no. (SGO

45443); Antofagasta: Valle de Río Frío, 3200 m, Mu-

ñoz C. 3863 (SGO).

RUPPIACEAE

91. Ruppia pene (Phil.) Skottsb.

Submerged tie peros nnial hai. o in slow-

flowing qe : Antofa-

gasta: Vega Zorritas, m, al. 94051

(CONC); presea Dése Llullaillaco, 3950 m,

Arroyo et al. 94047 (CO!

Gayana Bot. 55(2), 1998.

REFERENCES

ARAVENA, R., H. PEÑA, A GRILLI, O SUSUKI £ M.

MORDECKAI. 1989. Evolución isotópica de las llu-

vias y origen de las masas de aire en el Altiplano

chileno. L[AEA-TECDOC-502. Isotope Hydrology

Investigations in Latin America, pp. 129-142.

ARROYO, M. T. K. £ L. CAVIERES. 1997. The medite-

rranean-type climate flora of central Chile - What

, C. MARTICORENA éz C.VILLAGRAN.

1984. La flora de la Cordillera de los Andes en el

área de Laguna Grande y Laguna Chica, III Re-

gión, Chile. Gayana Bot. 41(1-2): 3-46.

ARROYO, M. T. K., C. VILLAGRAN, C. MARTICORENA

£ J. J. ARMESTO. 1982. Flora y relaciones bio-

geográficas en los Andes del norte de Chile (18-

1995). In: Alberto Veloso £ Eduardo Bustos

(eds.): El Ambiente Natural y las Poblaciones

Humanas de los Andes del Norte Grande de Chi-

le (Arica, Lat. 18? 28” S), Volume 1. pages 7192.

Rostlac, Montevideo.

ARROYO, M. T. K., F. A. SQUEO, J..J. ARMESTO é

C.VILLAGRAN. 1988. Effects of aridity on plant di-

versity in the northern Chile Andes: results of a na-

CAVIERES 6 C. MARTICORENA. 1992. Surv ey of

the alpine flora of Torres del Pg National

Park, Chile. Gayana Bot. 49(1-4):47

FUENZALIDA, H. 4 J. RUTLLANT. 1986. sc sobre

el origen del vapor de agua que precipita en el in-

poes altiplánico. Unpublished Informe Final.

ersidad de Chile. 51 pages

Goose, he S. 1954. The genus Nisaticn: Cronica

16:1-536.

nda “M. , M. GEYBUS, B. MESSERLI $ U-

HOTTERER. 1995. Late-glacial and early Holo-

cene lake sediments, ground water formation and

climate in the Atacama Altiplano 22-248. J. of

Paleolim.14:241-252.

a E. £ F. DICASTRI. 1975. Bioclimatografía de Chi-

le. Dirección de Investigación, Vicerrectoría Acadé-

mica, Universidad Católica de Chile, Santiago.

HEYWOOD, V.H. (Ed.). 1995. Global Biodiversity As-

sessment. Camb. U. Press, Cambridge

LoOOSER, G. 1960. El botánico Dr. Erich eS

Rev. Univ.(Stgo.) 44-45: 231-234, 1 retr.

MARTICORENA, C. UN

Catálogo de la flora vascular de la II Región de

Chile. Gayana Bot. 55:23-93.

MESSERLI, B., M. GROSJEAN, G. BONANI. A. so.

Rom

1993. Climate change and natural resource dyna-

mics of the Atacama Altiplano during the last

18,000 years: a preliminary synthesis. Mount.

Res. Devel. 13:117-127.

MUÑoz, C. á E. PISANO. 1950. Botánica. En Memoria

de Siete años de Investigación Agrícola, Impren-

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MUuÑoz, M., H. NUÑEz k J. YAÑEZ. (Eds.) 1996. Li-

bro Rojo de los Sitios Prioritarios para la Conser-

vación de la Diversidad Biológica en Chile.

Ministerio de Agricultura. Corporación Nacional

Forestal, Santiago.

Muñoz, C. 1960. Las especies de plantas descritas po

R. A. Philippi en el siglo XIX. Ediciones de la

Universidad de Chile, Santiago.

PHILIPPI, R.A. 1860. Viage al desierto de Atacama he-

cho de orden del gobierno de Chile en e verano

1853-54. Halle viii, 236 pp., 1 mapa, 27

REICHE, C. 1907. Grundziige der Pawzcwvetriuns

in Chile. (Traducido por G. Looser en 1934 co

Geografía Botánica de Chile, Tomo L, Imp. Univ.

424 pp.)

RICHTER, M. 1995. Klimaokologie Atacama. Absch-

lubbericht DFG-Projekt.

SOBERON, J. $ J. LLORENTE. 1993. The use of species

É Parque Nacional El

Morado oa Metropolitana, Chile). Gayana,

Bot. 51:

VUILLE, M. ca pa Seen und Salare im nordchile-

nischen Altiplano. Eine hydrologische Untersu-

e mit LANDSAT/TM- und LANDSAT/

MSS-Daten. Geographisches Institut, Universitat

Bern. 99 pages.

WERDERMANN, E. 1927. Beitráge zur Kenntnis der

Flora von Chile. Notizbl. Bot.Gart. Berlin-Dah-

lem 10(92):135-155.

Fecha de publicación: 07 de mayo de 1999

110

Gayana Bot. 55(2): 111-185, 1998.

ISSN 0016-5301

CHECKLIST OF CHILEAN LICHEN-FORMING

AND LICHENICOLOUS FUNGI

LISTA PATRON DE LOS LIQUENES Y HONGOS

LIQUENICOLAS DE CHILE

David J. Galloway' and Wanda Quilhot?

ABSTRACT

This Checklist of the lichens and lichenicolous fungi of

pags fungi (in 23 genera). Full bibliographic citations

are given for both accepted taxa and for synonyms and

references to relevant htesature are included for most ge-

The following ne 1: Ca-

pa deci (Zahlbr.) D.J. Galloway É

Quilhot, Dendriscocaulon calithamnion (Taylor) D.J.

Galloway £ Quilhot, Neuropogon durietzii (Motyka)

Neuropogon su-

Quilhot,

bantarcticus (F.J. Walker) D.J. Galloway £ Quilhot

Ss: Checklist, classification, lichens, licheno-

logy, eii fungi, Chilean lichens, plant taxo-

nomy.

INTRODUCTION

Chile, situated on the Pacific coast of South

America, is a long, narrow country extending 4350

km southwards from latitude 17?30"S to Cape

Horn at 56%S. It is bounded by Peru and Bolivia to

the north, by Argentina for much of its long eas-

tern border, and to the west by the Pacific Ocean.

Chile has sovereignty over Easter Island and the

Juan Fernández Archipelago and also has Antarc-

tic Territory between 53” and 90W. The country

is dominated by the Andean Cordillera which

'Department of Botany, The Natural History Museum,

Cromwell Road, London SW7 5BD, United Kingdom.

Present address: Landcare Research, Private Bag 1930,

Dunedin, New Zealand

“Escuela de Química y Farmacia, Facultad de Medicina,

Universidad de Valparaíso, Casilla 5001, Valparaíso,

Chile.

RESUMEN

La lista patrón de los líquenes y hongos liquenícolas de

Chile, incluídos el Territorio Antártico y las islas de

Juan Fernández y Pascua, contiene 1415 taxa en 304 gé-

neros, de los cuales 1383 son líquenes (en 281 éneros)

y 32 son hongos liquenícolas (en 23 géneros). Incluye

los taxa aceptados, los sinónimos y las referencias bi-

bliográficas. Se p nen las siguientes combinaciones:

Caloplaca austroshetlandica (Zahlbr.) D.J. Galloway $

Quilhot, Dendriscocaulon calithamnion (Taylor) D.J.

Galloway £ Quilhot, Neuropogon durietzii (Motyka)

D.J. Galloway £ Quilhot, Neuropogon patagonicus

(F.J. Walker) D.J. Galloway $: Quilhot, y Neuropogon

subantarcticus (F.J. Walker) D.J. Galloway $ Quilhot.

PALABRAS CLAVES: Lista patrón, clasificación, líque-

nes, liquenología, hongos liquenícolas, líquenes chile-

nos, taxonomía.

forms its entire eastern margin, the highest point

reaching to 6880 m (Ojos del Salado). Subject to

the influence of winds, storms and ocean currents,

as well as to violent tectonic activity related to the

presence of active plate margins along the floor of

the Pacific Ocean to the west, and to volcanic

eruptions and earthquakes, Chile is a land of clima-

tic and geographic extremes. Thus hot deserts in

the north and cold deserts in Antarctica, coastal

rocks and cliffs, alpine barrens, lava fields, grass-

land, scrub, temperate rainforest, Magellanic tun-

dra, urban and agricultural landscapes, parks,

gardens and plantations all provide a wealth of mi-

croclimates and microhabitats for lichens, making

Chile in many ways, a lichenologist's paradise.

Not surprisingly, Chile has one of the most

diverse and in places in terms of biomass, one of

the best developed lichen mycobiotas of any re-

gion in the world, but one which is alas, still in-

111

Gayana Bot. 55(2), 1998.

completely known in many parts. The history of li-

chenological exploration and discovery in Chile is

a long and varied one spanning over 200 years

(Galloway 8: Marticorena, 1991; Quilhot, 1995). It

is now over 30 years since Gerhard Follmann

compiled his catalogues of the lichens of Chile

(Follmann, 1962, 1963, 1965, 1967) from what

was then known either in the earlier literature or

from his own extensive field investigations. Since

that time, lichen taxonomy has undergone great

changes and many modern taxonomic revisions ci-

te Chilean material, with a number of recent mo-

nographs dealing specifically with Chilean lichens

(see Checklist Bibliography). The present Chec-

Klist includes 1416 taxa in 305 genera (comprising

1386 lichen taxa in 282 genera, and 30 taxa of li-

chenicolous fungi in 23 genera). This number

compares with 1415 taxa in 308 genera known

from New Zealand (Galloway, 1985; Malcolm é

Galloway, 1997); 2602 taxa in 393 genera known

from Sweden and Norway (Santesson, 1993):

1721 taxa in 267 genera from the United Kingdom

and Ireland (Purvis ef al., 1993); 3799 taxa in 477

genera known from the continental United States

and Canada (Esslinger $: Egan, 1995); 2145 taxa

in 305 genera from Italy (Nimis, 1993); 2657 taxa

in 393 genera from Australia (Filson, 1996); and

745 taxa in 157 genera from the Venezuelan An-

des (Marcano et al., 1996). Since much of Chile is

still “terra incognita” in lichenological terms, the

number of genera and of infrageneric taxa will un-

doubtedly increase over the next few years as more

studies are undertaken. In an excellent and refres-

hing discussion on lichen checklists and their va-

lue Nimis (1996: 15) states “...checklists can have

different nature, scope and contents, and they

should always be judged while considering the si-

tuation of floristic and taxonomic research that

they reflect...They offer an indispensable basis for

specimen revision, for the critical re-appraisal of

poorly known taxa, and for the further exploration

of under-investigated areas. In this sense, chec-

klists may and should be catalysts for new, more

intensive investigations. The best criterion for a

checklist to have accomplished its task as a facility

to the scientific community is the speed of its be-

coming outdated...”. It is hoped that the publica-

tion of this Checklist will further stimulate a range

of studies on Chile's lichens and lichenicolous

fungi, all of which deserve to be far better known

than they are at present.

In the Checklist taxa are arranged alphabeti-

112

cally by genus and species. Lichenicolous taxa are

indicated by an asterisk (*). Accepted genera are

printed in bold type followed by a bracketed syste-

matic arrangement of order and family designa-

tion. Citation of genera and dates of publication

follow the 8th edition of Anisworth 4 Bisby”s

Dictionary of the Fungi (Hawksworth et al., 1995).

Order and family affiliations of genera follow

Eriksson £ Hawksworth (1993) and Hawksworth

et al. (1995), and author abbreviations accord with

Kirk £ Ansell (1992). Names based on material

collected from Chile are indicated with a capital T

(in bold) after the citation, indicating a Chilean ty-

pe. Literature surveyed for this Checklist was that

current until 31 January 1998.

Any errors and omissions are the authors” so-

le responsibility. The authors realise that this is in

still a preliminary Checklist, but feel that in the ab-

sence of a modern Lichen Mycobiota of Chile, it is

important to have available a list of modern lichen

names for Chile to encourage lichenological inves-

tigations in this country. We would be grateful if

errors and omissions could be pointed out to us for

incorporation into a future, updated version. We

agree with Nimis (1996), that future biodiversity

inventories, of which this checklist is an example,

be made available on-line via the World-Wide-

Web, to be continuously updated as both explora-

tion of unknown or poorly-known areas, and local

and regional taxonomic studies are undertaken. It

is hoped to follow this checklist shortly with a key

to Chilean lichen genera, the bibliography (Gallo-

way $ Marticorena, 1991), checklist and keys

bridging the gap until a modern Lichen Mycobiota

of Chile is produced in the not too distant future.

ACKNOWLEDGMENTS

We are grateful to the following colleagues

and friends who assisted us in locating literature,

and in many other ways over the years: Prof. Teu-

vo Ahti (Helsinki); Prof. Braulio Araya (Viña del

Mar); Prof. Mary T.K. Arroyo (Santiago); Prof.

Dr. Gerhard Follmann (Kúln); Sr. Gerardo Guz-

mán (Valparaíso); Prof. Dr. Hannes Hertel (Miin-

chen); Prof. Henry Imshaug (East Lansing); Prof.

Per Magnus Jórgensen (Bergen); Izabella Kruger

(Landcare Research, Lincoln); Prof. Clodomiro

Marticorena (Concepción); Prof. Roland Moberg

(Uppsala); Dra. Melica Muñoz (Santiago); the late

Prof. Ed Pisano (Punta Arenas); Prof. Jorge

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

Redon (Viña del Mar); Dr. Soili Stenroos (Helsin-

ki); Ngaire van der Velde (Landcare Research,

Lincoln); and Dr. Mats Wedin (London). This

work was begun in 1988 when the senior author

was on the staff of the Natural History Museum in

London, and the Keeper of Botany, Prof. Steve

Blackmore, is thanked for his enthusiastic support

of Chile-based research in the Department of Bo-

tany.This work was also supported in part by

FONDECYT Proyecto N* 1960353, and we wish

to thank FONDECYT for their help in publishing

the Checklist.

REFERENCES

ERIKSSON, O.E. ££ HAWKSWORTH, D.L. 1993, Outline

of the Ascomycetes-1993. Systema Ascomyce-

1-217.

tum 12:

ESSLINGER, T.L. $ EGAN, R.S. 1995. A sixth checklist

of the lichen-forming, lichenicolous and allied

fungi of the continental United States and Cana-

da. Bryologist 98: 467-549,

FILSON, R.B. 1996. Checklist of Australian lichens and

allied fungi. Flora of Australia Supplementary

Series No.7. Canberra: Australian Biological Re-

sources Study. 204 pp.

FOLLMANN, G. 1962. Catálogo de los líquenes de Chi-

le. Parte I. Pyrenocarpae. Revista Univ. (Santia-

go) 46: 173-203.

FOLLMANN, G. 1963. Catálogo de los líquenes de Chi-

le. Parte IL Coniocarpiidae y Graphidiidae. Re-

vista Univ. (Santiago) 47: 63-97.

LLMANN, G. 1965. Catálogo de los líquenes de Chi-

le. Parte HI. Thelotrematales y Cyanophilales.

Revista Univ. (Santiago) 49: 17-65.

FOLLMANN, G. 1967. Catálogo de los líquenes de Chi-

le. Parte IV. Stictaceae y Peltigeraceae. Revista

Univ. (Santiago) 50-51: 33-75.

GALLOWAY, D.J. 1985. Flora of New Zealand Li-

; ORENA, C. 1991. A biblio-

graphy of Chilean lichenology. Gayana Bot. 48:

7-66.

HAWKSWORTH, D.L., KIRK, P.M., SUTTON, B.C. £

PEGLER, D.N. 1995. Ainsworth €: Bisby's Dic-

tionary of the Fungi (Sth edition). Wallingford:

CAB eme XII + 616 pp.

KIRK, P.M. $2 ANSELL, A.E. 1992. Authors of fungal na-

mes. A list ms autor of scientific names of fungi,

with recommended standard forms of their names,

including aa: Index of Fungi Supple-

ment. Wallingford: CAB nus 95 pp.

MALCOLM, W.M. £ GALLOWAY, D.J. 1997. New Zea-

land lichens. Checklist, pia and glossary. We-

llington: Museum of New Zealand, Te Papa

Tongarewa. VI + 192

MARCANO, V., MORALES MENDEZ, A., SIPMAN, H. 4

CALDERON, L. 1996. A first checklist of the li-

chen-forming fungi of the Venezuelan Andes.

Tropical Bryol. 12: 193-235.

NiMIS, P.L. 1993. The lichens of Italy, an annotated

catalogue. Monografia XIL Torino: Museo Re-

, P.W. 1993.

Checklis of lichens of Great Pr and Ireland.

Lichen Soc. Bull. 72 (Suppl.): 1-75.

action W. 1995. Liquenes. /n: J.A. Simonetti,

M.T.K. Arroyo, A.E. Spotorno € E. Lozada (ed)

Diversidad biologica de Chile. Santiago: CO-

37:

- pp. 26-

SANTESSON, R. 1993. The lichens and lichenicolous

fungi of Sweden and Norway. Lund: SBT-fórla-

get. 240

113

Gayana Bot. 55(2), 1998.

THE CHECKLIST

*Abrothallus De Not. (1849)

Enron incert, sed.]

ae Wedin, Lichenologist 26: 304 (1994).

poh acid (Sommerf.) Arnold, Flora 57: 102 (1874).

Wedin K R. Sant., Lichenologist 26: 301 (1994).

Lit.: Zahlbruckner (1925); Hertel (1971); Wedin (1994).

ra A. Massal. (1852)

e cinisc Acarosporaceae]

ndina H. Magn., K. Vet. O. Vitterh. Samh. Handl. f. 6. ser. B. 6 (17): 26 (1956).

Jnibiafimol (Nyl.) Th. Fr., Nova Acta Regiae Soc. Sci. Upsal., ser. 3,3: 190

(1861).

Acarospora deceptionis C.W. Dodge, Publ. Inst. Antart. Chile 6: 6 (1965) T.

?catamarcae H. Magn., Acta Horti Gothob.17: 64 (1947

convoluta Darb., Wiss. Ergebn. Schwed. esop Pe 1903, 4 (11): 16 (1912).

macrocyclos Vain., Résult. Voy. Belgica, Lich.: 34 (190

plumbeocaesia Zahlbr., Acta Horti ne 1 hos T.

?punae 1.M. Lamb, Lilloa 14: 231 (194

sanguinascens Zahlbr., Acta Horti dr 2 Ras Y

schleicheri (Ach.) A. Massal., Ric. Auton. Lic. Crost.: 27 (1852).

Acarospora bella (Nyl.) Jatta, Malpighia 20: 10 (1906

Lecanora bella Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 156 (1855) T [non. Ach.]

Acarospora chilensis H. Magn., Kongl. Svenska Vetenskapsakad. Handl. II, 7 (4).: 367 (1929) T.

br. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 4 (13): 453 (1926) T.

skotts.

smaragdula (Wahlenb.) A. Massal., Ric. Auton. Lich. Crost:.: 29 (1

socialis H. Magn., acera 21 65): 252 (1929).

sparsiuscula H. Magn., Acta Horti Gothob. 17: 65 (1947).

strigata (Nyl.) Jatta, Pas 20: 10 (1906

852

Lecanora strigata Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 155 (1855) T.

odium rm (Nyl.) Mill. Arg., Flora 72: 510 (1889).

Plac

subcastanea (Nyl.) Hue, Nouv. Arch. Mus. Hist. Nat., sér. 7 1: 162 (1909).

canora subcatanea Nyl., Lich. Nov. Zel.: 145 (1888

nora cervina sensu Nyl., Ann. Sci. Nat. Bot. sér.4, a 155 (1855) [non Ach.]

5, 1: 162 (1909

?subglebosa (Miill. Arg.) Hue, Nouv. Arch. Mus. Hist. Nat., sér.

Placodium subglebosum Miill. Arg., Flora 72: 510 (1889).

?theleomma 1. M. Lamb, Lilloa 14: 233 (1948).

pen (Mill. Arg.) Hue, Nouv. Arch. Mus. Hist. Nat., sér.5,1: 163 (1909).

um trachyti Mill. Arg., Hedwigia 31: 279 (1892).

Lit.: Zahlbruckner (1924); Magnusson (1929, 1947, 1956

Jacobsen £ Kappen (1988); Hafellner (1993); Castello £ Nimis (1994

2. Agyrium Fr. (1822)

premeras Agyriaceae]

ens Miill. Arg., Nuovo Giorn. Bot. Ital. 21: 50 (1889).

Lt: porra Argoviensis (1889).

3. Alectoria Ach. Asa

[Lecanorales: Alectoriac

nigricans (Ach.) NyL. Ele Scand.: 71 (1861).

Alectoria nigricans var. implexiformis Rásánen,

aaa (Hoffm.) A.Massal., Sched. Crit. Lich. Exs. Ital.: 47 (1855).

(Rásánen) D. Hawksw., tg 72: 249 (1969).

var. citrina

Alectoria ochroleuca f. citrina Rásiinen Ann

4. Amandinea M. enc ex Scheid. £¿ H. Mayrhofer (1993)

[Lecanorales: Physci

petermannii

114

);, Weber (1962, 1968); Follmann (1965c); Redon (1985);

nn. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2(1): 12(1932) T.

- “Vanamo” 2 (1): 121 (1932) T.

30)

(Hue) Ma Msn, H. Mayrhofer é: Scheid., Lichenologist 26: 39 (1994).

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

unctata (Hoffm.) Coppins « Scheid., Lichenologist 25: 343 (1993).

Lit.: Lamb (1968); Redon (1985); Jacobsen $ Kappen (1988); Aptroot £ van der Knapp (1993); Scheidegger

(1993); Matzer et al. (1994).

ecium Hampe ex A. Massal. (1860)

¡yremes:3 Pyrenulaceae ae]

pyrenuloides (Mont.) Mill. Arg., Linnaea 63: 44 (1880).

Lit.: Follmann (1961).

6. Anzia Stizenb. (1861) nom. cons.

[Lecanorales: Parmeliaceae]

montana R. Sant., Thunbergia 2: 1 (1986).

Lit.: Moberg (1986); oshirenra (1995); Calvelo (1996).

7. Arthonia Ach. (1806) nom. cons.

Dl ari Arthoniace ae]

is Follmann, Willdenowia > 365 (1968) T.

e (Zahlbr.) C.W. Dodge, (196

Arthonia subdiffusa var. australis od Acta Horti Gothob. 2: 2 (1925) T.

austrolitoralis Follmann, Willdenowia 4: 368 (19

*badia Wedin 8 Hafellner, Lichenologist 30: 65 (1998) T.

berberina Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11): 325 (1924) T.

catillaria Vain., étud. Lich. Brésil 2: 165 (1890).

chilensis Follmann, Willdenowia 4: 370 (1968) T.

998) T

cytisi A. Massal. v ar. meridionalis Zahlbr. In C. polar Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3

(11): 324 (1924) E

dispersa (Schrad.) Nyl., Lich. Scand: 261 (1861).

*epiphyscia Ny]l., Flora 58: 361 (1875

*flavicantis Wedin $: Hafellner, Lichenologist 30: 72 (1998) T.

follmannii C.W. Dodge, Nova Hedwigia 12: 310 (1967) [*1966”] T.

fuscescens Fée, Essai Crypt. Ecorc.: 56 (1824).

Ps (Tul.) R. Sant., Svensk Bot. ignora 54 (4): 501 (1960).

yl., Ann. Sci. Nat. Bot. sér. 4,3: 172 (1855) T

ilicinoides Stein., Sitzungsber. Kaiserl. Akad. Wiss. Math. -Naturwiss. CL, Abt. 1, 106 (1): 231 (1897).

*minuta Wedin £ Hafellner, lacas 30: 75 (1998).

miserula Nyl., Acta Soc. Sci. Fenn. 7: 484 (1863

palmicola Ach., Syn. Meth. Lich.: 5 (1814).

arpoides (S. Kondr. $: D.J. Galloway) Wedin « S. Kondr., Lichenologist 29: 97 (1997).

Dactylospora plectocarpoides S. Kondr. £ D.J. Galloway, Acta Bot. Fenn. 150: 95 (1994) T.

*pseudocyphellariae Wedin, Lichenologist 25: 301 (1993).

Pe Miill. Arg., Nuovo Giorn. Bot. Ital. 21: 50 (1889).

radiata (Pers.) Ach., Kongl. Vetensk. Acad. Nya Handl.: 131 (1808).

ramulosa Nyl., Mém. Soc. Sci. Nat. Cherbourg 2: 335 (1854),

rufella Nyl., Amn. Sci. Nat. Bot. sér. 4, 3: 170 (1855) T.

>*santessoniana Wedin £ Hafellner, Lichenologist 30: 81 (1998).

*subaggregata Wedin 8: Hafellner, Lichenologist 30: 84 (1998) T.

subantarctica Vvstedal, Norsk Polarinst. Skr. 185: 36 (1986).

subexcedens Nyl., Flora 62: 221 (1879).

subdiffusa Willey var. australis Zahlbr., pcia Horti Gothob. 2: 2 (1925) T.

s Nyl., Lich. Nov. Zel.: 148 (188

subnebulosa Zahlbr. In C. Skottsberg, Nat. AS Juan Fernández Easter Isl. 2 (Bot.), 3 (11): 324 (1924) T.

trilocularis Miill. Arg., Flora 64: 233 (1881

atte Lich. Fueg. Patag.: 18 quo

Lit: eE (1855); Múller e ti (1889); Zahlbruckner cie 1925, 1926); Santesson (1952); Follmann

(1962, 1968a, 1968b); Dodge (1966); Wvstedal (1986); Aptroot « van der Knapp (1993); Egea £ Torrente (1995);

Kondratyuk £: Galloway (1994); a 82 Kondratyuk (1997); Wedin « Hafellner (1998).

315

Gayana Bot. 55(2), 1998.

8. Arthoniactis (Vain.) Clem. (1909) = Lecanactis

[Arthoniales: Roccellaceae]

chilena C.W. Dodge, Nova Hedwigia 12: 326 (1967) [“*1966”] T.

Lit.: Dodge (1967).

9. Arthopyrenia A. Massal. (1852)

[Dothideales: Arthopyreniaceae]

adnexa var. leptosperma Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter sl. 2 (Bot.), 3 (11): 320

(1924) T

australis Múll. Arg., Miss. Sci. Cap Horn, Lich.: 172 (1888) T.

brachyspora Miill. Arg., Nuovo Giorn. Bot. Ital. 21: 52 (1889).

cinchonae (Ach.) Mill. Arg., Flora 66: 289 (1883).

itima Dvstedal, Norsk Polarinst. Skr. 185: 37 (1986).

planorbis (Ach.) Mill. Arg., Mém. Soc. Phys. Genéve 30: 27 (1888).

pa e. ) A. Massal., Ric. Auton. Lich. Crost.: 168 a

subfallax ) Mill. Arg., Nuovo Giorn. Bot. Ital. 21: 51 (18

Lit.: Miller Pe de (1888, 1889); Zahlbruckner (1924, 1925); pea (1986).

10. Arthothelium A. Massal. (1852)

[Arthoniales: Arthoniaceae]

chilense C .W. Dodge, Nova Hedwigia 12: 311 (1967) [*1966”] T.

cingulatum R. Sant., Symb. Bot. Upsal. 12: 93 (1952) T

follmannii C.W. Dodge, Nova Hedwigia 12: cds (1967) [“1966”] T.

968) T.

spilomatoides (Nyl. ) Zahlbr., Cat. Lich. Univ. 2: 135 (19 22).

Arthonia spilomatoides Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 169 (1855) T.

igrense C.W. Dodge, Nova Hedwigia 12: pre [“1966”] T.

taediosum (Nyl.) Mill. Arg., Flora 63: 287 (1880).

Arthonia taediosa Nyl., Ann. Sci. Nat. Bot. sér.4, 3: 171 (1855) T

Lit.: Zahlbruckner (1925); Santesson (1952); Dodge (1967); Follmann (19680); Makhija € Patwardhan (1995);

Grube « Giralt (1996).

11, Arthrorhaphis Th. Fr. (1860) nom. cons. prop.

[Patellariales: Arthrorhaphidaceae]

pep (Schaer.) R. Sant. In D.L. Hawksworth et al., Lichenologist 12: 106 (1980).

lla (Ach.) Poelt, Bestimmungs. Eur. Flecht.: 126 (1969).

Eat: 2 (1971); Galloway £ Bartlett (1986); Jóorgensen 8: Santesson (1993); Obermayer (1996); Ihlen (1998).

12. Aspicilia A. Massal. (1852) nom. cons.

[Lecanorales: Hymeneliaceae]

calcarea (L.) Mudd, Man. Br. Lich.: 161 (1861).

Note: It is likely that the taxon Lecanora masafuerensis Zahlbr. [In C. Skottsberg, Nat. Hist Juan Fernández Easter

Isl. 2 (Bot.), 3 (11): 378 (1924) T.] is also referable to Aspicilia, but as we have not seen material of it we hesitate

to name it in Aspicilia

Lit.: Jatta (1906); Zahlbruckner (1924).

13. Asterothyrium Miill. de em. Kalb € Vezda (1890)

[Ostropales: Thelotrematac:

rotuliforme (MúllL. poe ) ¡Séros. In Sérusiaux £ de Sloover, Veróff. Geobot. Inst. Ziirich 91: 268 (1986).

Gyalectidium rotuliforme Miill. Arg., Rev. Mycol. 38: 65 (1888).

Lit.: Santesson (1952); Farkas $: Sipman (1993).

14. Austrolecia Hertel (1984)

[Lecanorales: Catillariaceae]

antarctica Hertel, Beih. Nova Hedwigia 79: 453 (1984) T.

Lit.: Hertel (1984).

15. Bacidia De Not. (1846)

[Lecanorales: oi

(1924) < Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3 (11): 366

116

Checklist of Chilean lichen: GALLOWAY, D.J. € W. QUILHOT

delapsans Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter 15]. 2 (Bot.), 3 (11): 366 (1924) T.

dodgei Follmann, Nova Hedwigia 14: 248 (1968) [“1967” E dd da

filiformis Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 57 (1932) T.

laurocerasi (Delise) Vain., Acta Soc. Fauna. Fl. Fenn. 53 (017150922

pallida (Mill. Arg.) Darb., Wiss. Ergebn. Schwed. ie Mcg 1901-1903, 4 (11): 49 (1912).

Patellaria pallida, Miss. Sci. Cap Horn, Lich.: 167 (188

lidocarnea (Mill. Arg.) Zahlbr., Cat. Lich. Univ. 4: 231 , ee

*plumbina (Anzi) R. Sant., Svensk Bot. Tidskr. 54 (4): 513 (1960

rhodochroa (Hue) Darb., Br. Antarct. (“Terra Nova”) Exp. 1910, Nat. Hist. Rept. Bot. 3: 47 (1923).

stipata 1.M. Lamb, Rhodora 56: 126 (1954

subluteola (Nyl. a Sitzungsber. Kaisedl Akad. Wiss. Math.-Naturwiss. pee e 1, 111 (1): 395 (1902).

tuberc, Darb., s. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4 (11): 5 (19

Lit: Zahlbrackner (1924) dansaion (1959. Follmann (1968a); Aptroot $ van der ias Ekman (1996).

16. Bacidina Vezda (1991) nom. cons. prop.

[Lecanorales: Lecanoraceae

apiahica (Miill. Arg.) Vezda, Folia Geobot. Phytotax. 25: 432 (1991) [**1990”].

Lit.: Vezda (1990); Ekman (1996).

17. Bactrospora A. Massal. (1852)

[Arthoniales: Roccellaceae

1 C.W. Dodge) Egea « Torrente, Lichenologist 25: 219 (1993).

smc acicularis C.W. Dodge, Nova Hedwigia e (1969) [**1968”] T.

ea £ Torrente, Lichenologist 25: 235 (199

Lit: ques e eel (1980); Egea £ Torrente (1993, Aa

18. Baeomyces Pers. (1794)

[Leotiales: Baeomycetaceae]

chilensis (Nyl.) Cromb., Bot. J. Linn. Soc. 16: 223 (1876).

Baeomyces rufus var. chilensis Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 177 (1855).

Biatora byssoides var. chilensis Mont. In C. Gay, Hist. Chile, Bot. 8: 173 (1854) T.

ramalinellus Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 146 (1855) T

Lit.: Jahns (1971); Imshaug (1972); Gierl € Kalb (1993).

19. Bellemerea A «£ Cl. Roux (1984)

[Lecanoraceae: Porpidiaceae]

alpina (Sommerf.) Chande $ Cl. Roux, Bull. Soc. Bot. Centre-Ouest, sér. 15: 129 (1984).

subsorediza (Lynge) R. Sant. In R. Moberg, Thunbergia 5: 2 (1987).

Lit.: Hertel (1989); nó et al.(1992).

20. Biatora Fr. (1817)

[Lecanorales: Lecanoraceae]

albopraetextata (C. Knight) Hellb., Bih. Kongl. Svenska Vetenskapsakad. Handl. 21 (3) 13: 105 (1896).

Lecidea albopraetexta C. Knight, Trans. N.Z. Inst. 12: 376 (1880).

glaucopa (Hook. f. £ Taylor) Rásinen, pu Soc. Bot. Zool.-Bot. Fenn. “Vanamo” 2 (1): 58 (1932).

Lecidea glaucopa Hook. f. ££ Taylor, Lond. J. Bot. 3: 637 (1844) T.

Lecidea conflectens Nyl., Lich. Fueg. Patag.: 15 (1888).

SE epichlorotica Miill. Arg., Miss. Sci. Cap Horn, Lich.: 164 (188

var. oculans (Nyl.) Rásinen, Ann. Bot. Soc. Zool.-Bot. Fenn. cti 2 (1): 58 (1932).

pco oculans Ny]l., Lich. Fueg. Patag.: 13 (1888) T.

Lecidea epichlorotica var. expallens Miill. Arg., Miss. Sci. Cap Horn, Lich.: 165 (1889) T.

Lecidea conflectens var. expallens (Mill. Arg.) Mill. Arg., Nuovo Giorn. Bot. Ital. 21: 46 (18

Note: It is possible sorda the taxon Lecidea lividula Miill. Arg. [Miss. Sci. Cap Horn, Lich.: 164 pa in should al-

so be referred to Bia

Lit.: Printzen (1995 sm.

1. Biatorella De Not. (1846)

app Biatorellaceae]

ica C.W. Dodge, Trans. Amer. Microscóp. Soc. 84: 512 (1965) T.

Ln: ur e 1965b); Hafellner € Casares-Porcel (1992); Aptroot $ van der Knapp (1993); Hafellner

(1994, 1995),

117

Gayana Bot. 55(2), 1998.

22. Biatorina A. Massal. (1852) nom. rej. prop. = Catinaria

[Lecanorales: Bacidiaceae]

chilena C.W. Dodge, Nova Hedwigia 16: 490 (1969) T

coccinea C.W. Dodge, Nova Hediwgia 12: 333 (1967) [* 1966”] T.

Lit.: Jorgensen $ Santesson (1993).

*Biatoropsis Risánen (1934)

[Platygloeales: incert. sed.]

usnearum Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 5 (9): 8 (1934).

Lit.: Diederich £ Christiansen (1994); Diederich (1996).

Blastenia A. Massal. (1852) = Caloplaca

23. Brigantiaea Trevis (1853)

[Lecanorales: Brigantiaceae]

chrysosticta (Hook. f. £ ro Hafellner $ Bellem., Nova rra si e (1981).

Biatora berteroanum Mont., Ann. Sci. Nat. Bot. sér. 3, 18: 310 (185

Heterothecium nal (Mont.) Mont. In C. Gay, Hist. Chile, a qe 176 (1852).

Brigantiaea berteroana (Mont.) Trevis., Spighe e Paglie: 9 (1853).

ium leucoxantha var. albidius Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.)

3 (11): 368 (1924) T.

fuscolutea (Dicks.) R. Sant. In J. Poelt £ A. Vezda, Bestimmungsschl. Eur. Flecht. Erg. II: 116 (1981).

phaeomma (Nyl.) Hafellner, Symb. Bot. Upsal. 32 (1): 61 (1997).

Lecidea phaeomma Nyl., Lich. Nov. Zel.: 90 (1888).

rete res var. austroamericanum Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 20

7 (1944).

Rrionantinen

á Hafellner, Lichenologist 15: 265 (1983).

Lit.: Hafellner « Bellemere (1981); Hafellner (1983, 1997).

[Lecanorales: Parmeliacea

chalybeiformis (L.) Brodo £ D. Hawksw., Opera Bot. 42: 81 (1977

Lit.: Hawksworth (1972); Brodo £ Hawksworth (1977); Redon (1985); Jacobsen $ Kappen (1988).

25. Buellia De Not. (1846)

[Lecanorales: Physciaceae]

agelaeoides Miill. Arg., Hedwigia 31: 283 (1892) T.

albula abeto ) Múill. Arg., Bull. Herb. Boissier 2, App. 1: 71 (1894).

Mill. Arg., Hedwigia 31: 282 (1892).

purecogén Mill. Arg. In Zahlbr., Acta Horti. Gothob. 2: 25 (1925).

anisomera Vain., Résult. Voy. Belgica, Lich.: 26 (190 se

augustata Vain., Résult. Voy. Belgica, Lich.: 26 (190

Buellia brabantica Vain., Résult. Voy. Belgica, ae 26 (1903).

pos oa (Hook. f. £ Taylor) IM. Lamb ex Dodge, Rep. B.A.N.Z.A.R.E., ser. B, 7: 248 (1948).

anora babingtoni [sic.] Hook. f. 8: Taylor. In Hook. f., Fl. Ant. 1 (ID): 535 (1847).

huron Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter 1sl. 2 (Bot.), 4 (13): 459 (1926) T

art iento var. oceanica Zahlbr. In C. Skottsberg, Nat. Hist Juan Fernández Easter Isl. 2 (Bot.), 3(11):

(

24. Bryoria Brodo € D. Hawksw. (1977)

coniops (Wahlenb. ex < Ach. ) Th.Fr., Nova Acta Regiae Soc. Sci. ei ser. 3, 3: 331 (1861).

cog nsis C.W. Dodge, Nova oa 12: 349 (1967) [*1966”] T

darbishirei 1.M. Lamb, Br. Antarct. Surv. Sci. Rep. 61: 23 (1968).

Rinodina crassa Darb., Wiss. Erecbn Schwed. Siidpolarexp. 1901-1903, 4 (11): 13 (1912).

eVanescens Darb., Br. Antarct. (“Terra Nova” ) Exp. 1910, Nat. Hist. Rep. Bot. 1 (3): 36 (1923).

Zahlbr., K. Sv. Vetensk.-Akad. Handl. 57(6): 53 (1917) T.

pom vareolata (Nyl.) Mill. Arg.. propos 283 (1892).

acotada se Arg., Flora 71: 542 (188:

Dodge, Nova Hedwigia 07 350 (1967) [*1966”] T.

ga Nat. Antarct. Exp. 1901-

(Kremp.) Múill. Arg., Flora 63: 19 (1880).

EW.

-A.N.Z.A.R.E. ser. B, 7: 244 (1948).

Múll ei e Herb. Boissier 1(2): 52 (1893).

Skottsberg, Nat. H: Hist. Juan Fernández Easter ls]. 2 (Bot.), 4 (13): 459 (1926) T.

118

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

halophiloides var. pruinosula Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 4 (13):

455 (1926) T.

illaetabilis 1M. Lamb, Br. Antarct. Surv. Sci. Rep. 61: 29 (1968).

inordinata (Hue) Darb., Br. Antarct. (“Terra Nova”) Exp. 1910, Nat. Hist. Rep. Bot. 1 (3): 63 (1923).

isabellina (Hue) Darb., Br. Antarct. (“Terra Nova”) Exp. 1910, Nat. Hist. Rep. Bot. 1 (3): 63 (1923).

jattana Mill. Arg., Nuov. Giorn. Bot. Ital. 21: 48 (1889) T.

jorgensis Zahlbr., Acta Horti Gothob. 2: 23 dad

kitensis (Stirt.) Zahlbr., Cat. Lich. Univ. 7: 374 (193

Lecidea kitensis Stirt., Proc. Phil. Soc. Glasgow e 111 (1879) T.

latemarginata Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4 (11): 15 (1912).

melanostola (Hue) Darb., Br. Antarct. (“Terra Nova”) Exp. 1910, Nat. Hist. pi Bot. 1 (3): 63 (1923).

nelsonii Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, sde 15 (1912).

olivacea Mill. Arg., Verhandl. Zool.-Bot. Ges. Wien 43: 298 (18

papillata (Sommerf.) Tuck., Lichens of California, Oregon 4 us Mts: 26 (1866).

paschalis Zahlbr. In C. Skottsberg., Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 4 (13): 456 ( a

perlata (Hue) Darb., Br. Antarct. (“Terra Nova”) Exp. 1910, Nat. Hist. Rep. Bot. 1 (3): 64 (1923

pirionii de Lesd., Bull. Soc. Bot. France 81: 767 (1934) T.

protothallina (Kremp.) Vain. var. pygmaea Risánen, Ann. Soc. Bot. Zool.-Bot. Fenn. “Vanamo” 2 (1): 31

(1932) T.

Era Darb., Br. Antarct. (“Terra Nova”) Exp. 1910, Nat. Hist. Rep. Bot. 1 e 41 (1923).

arb., Br. Antarct. (“Terra Nova”) Exp. 1910, e Hist. Rep. Bot. 1 (3): 64 (1923),

cal (Schaer.) Kórb., Syst. Lich. Germ.: 228 (185

siphoniatula Zahlbr. In C. Skottsberg, le Hist. Juan lada Easter Isl. 2 (Bot.), 3(11): 400 (1924) T.

sordidula Jatta, Malpighia 20: 12 (1906

spuria (Schaer.) Anzi, Cat. Lich. Sondr.: a ).

stellulata (Taylor) Mudd, Man. Brit. Lich.: 216 (1861).

subdisciformis (Leight.) Vain. var. americana Jatta, Malpighia 20: 12 (1906) T.

PR (Hue) Darb., Br. Antarct. da Le Exp. 1910, Nat. Hist. Rep. Bot. 1 (3): 64 (1923).

subsquamescens Jatta, Malpighia 20: 11 (1906

taltalensis C.W. Dodge, Nova Hedwigia 12: 351 Gon 1966”] T.

tristicolor Zahlbr., Acta Horti Gothob. 2: 21 (192

Lit.: Jatta (1906); Zahlbruckner (1917, 1924, 1925, dodo: Magnusson (1955); Dodge (1967); Follmann (1965c);

Lamb (1968); Redon (1985); Jacobsen £ Kappen (1988); Scheidegger (1993).

26. Bunodophoron A. Massal. (1861)

[Lecanorales: Sphaerophoraceae

agnetae Wedin, Mycotaxon 53: 33 (1985).

australe (Laurer) A. Massal., Mem. Imp. Reale 1st. po Sci. 10: 76 (1861).

dodgei (Ohlsson) Wedin, Pl. Syst. Evol. 187: 232 (199

Sphaerophorus dodgei Ohlsson. In Wedin, pora 24: 121 (1992) T.

imshaugii (Ohlsson) Wedin, Pl. Syst. Evol. 187: 233 (1993).

haerophorus imshaugii Ohlsson. In D.J. Galloway, N.Z. J. Bot. 21: 197 (1983).

insigne (Laurer) Wedin, Pl. Syst. Evol. 187: 233 (1993).

Sphaerophorus insignis Laurer, Linnaea 2: 45 (1827).

macrocarpum (Ohlsson) Wedin, Pl. Syst. Evol. 187: 233 (1993).

Sphaerophorus macrocarpus Ohlsson. In D.J. Galloway, N.Z. J. Bot. 21: 197 (1983).

patagonicum (C.W. Dodge) Wedin, Pl. Syst. Evol. 187: 234 (1993).

Pleurocybe patagonica C.W. Dodge, Nova Hedwigia 16: 484 (196

Sphaerophorus que ao (C.W. Dodge) Ohlsson. In D.J. mc N.Z. J. Bot. 21: 197 (1983).

ramuliferum (LM. Lamb) Wedin, Pl. Syst. Evol. 187: 234 (1993).

ph ropholia o LM. Lamb, Farlowia 4: 426 (1955).

serobiculatum (C.Bab.) Wedin, Pl. Syst. Evol. 187: 234 (1993).

Sphaerophorus australis var. scrobiculatus C. Bab. In Hook. f., Fl. Nov. Zel. 2: 305 (1855).

LO scrobiculatus (C. Bab.) M. Sató, Misc. Bryol. Lichenol. (Nichinan) 4: 151 (1968).

tibelli (Wedin) Wedin, Pl. Syst. Evol. 187: 234 (1993).

> Mete tibellii Wedin, Lichenologist 24: 129 (1992).

e (Wedin) Wedin, Pl. Syst. Evol. 187: 234 (1993).

phacio plo whakapapensis Wedin. In Wedin 4: Tibell, N.Z. J. Bot. 29: 287 (1991).

Lit.: Tibell (1987, 1994); Wedin (1993, 1994, 1995b).

27. Bulbothrix Hale ld

[Lecanorales: Parmeliaceae]

119

Gayana Bot. 55(2), 1998.

goebelii (Zenker)Hale, Smithsonian Contr. Bot. 32: 14 (1976).

imshaugii (Hale) Hale, Phytologia 28: 480 ( 1974).

isidiza (Nyl.) Hale, Phytologia 28: 480 (1974).

tabacina (Mont. £ Bosch) Hale, Phytologia 28: 481 (1974).

Lit.: Hale (1976); Elix (1994a).

Byssocaulon Mont. (1835)

[Arthoniales: Chrysotrichaceae]

andidum Miill. Arg., Nuov. Giorn. Bot. Ital. 21: 49 (1889). = Sagenidium

niveum Mont., Amn. Sci. Nat. Bot. sér. 2, 3: 355 (1835) = Sagenidium.

Lit.: Montagne (1835); Follmann (1968a).

28. Byssoloma Trevis. (1853).

[Lecanorales: Pilocarpaceae]

leucoblep (Nyl.) Vain. em. R. Sant., Symb. Bot. Upsal. 12: 483 (1952).

Lit.: Santesson (1952); Vezda (1987); Kalb $: Vezda ( 1990); Malcolm £ Vezda (1995); Elix er al. (1995).

29. Calicium Pers. (1794)

[Caliciales: Caliciaceae]

abietinum Pers., Tent. Disp. Meth. Fung. (Suppl.): 59 (1797).

- SSp. Tibell, Publ. Herb. Univ. Uppsal. 12: 1 (1984).

glaucellum Ach., Methodus: 97 (1803).

hyperelloides Nyl., Syn. Meth. Lich. 1: 153 (1860).

salacinum Pers., Usteri Ann. Bot. 7: 20 (1794).

trabinellum (Ach.) Ach., Methodus Suppl.: 14 (1803).

tricolor F.Wilson, Vict. Nat. 6: 64 (1889).

viride Pers., Usteri Ann. Bot. 7: 20 (1794).

Lit.: Follmann (1962); Tibell (1987).

30. Calopadia Vezda (1986)

[Lecanorales: Ectolechiaceae

fusca (Mill. Arg.) Vezda, Folia Geobot. Phytotax. 21: 215 (1986).

Lit.: Vezda (1986).

31. Caloplaca Th.Fr. ( 1860) nom. cons.

[Teloschistales: Teloschistaceae]

ammiospila (Wahlenb. in Ach.) H. Olivier, Mém. Soc. Nat. Sci. Nat. Cherbourg 37: 136 (1909).

athallina Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 4(11): 9 (1912).

austroshetlandica (Zahlbr.) D.J. Galloway £ Quilhot, comb.nov.

Basionym: Blastenia austroshetlandica Zahlbr., K. Sv. Vetensk.-Akad. Handl. 57 (6): 47 (1917) T.

buelliae Olech 82 Sochting, Lichenologist 25: 261 (1993) T.

carnella (Nyl.) Jatta, Bull. Bot. Soc. Ital.: 255 (1911).

cerina (Ehrh.) Th.Fr., Lich. Arctoi: 1 18 (1860).

cinericola (Hue) Darb., Br. Antarct. (Terra Nova”) Exp. 1910, Nat. Hist. Rep. Bot. 3: 54 (1923).

cirrochrooides (Vain. ) Zahlbr., Cat. Lich. Univ. 7: 225 (1931)

citrina (Hoffm.) Th.Fr., Nov. Acta Regiae Soc. Sci. Upsal. ser. 3, 3: 218 (1861).

clandestina - In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3(11): 394 (1924) T.

coralligera (Hue) pc Cat. Lich. Univ. 7: 274 (1931).

ma coralligera (Hue) Gyeln., Acta F. Fl Univ., ser. HL, L, No 5-6: :

cribrosa (Hue) Zahlbr.. Cat. Lich. Univ. 7: 274 ( 1931). A

issima (Nyl.) Zahlbr., Cat. Lich. Univ. 7: 238 (1931).

Follmannia rufa CM. Dodge, Nova Hedwigia 12: 335 ( 1967) [“1966”] T.

as 7 pls a ££ Redon, Willdenowia 6: 448 (1972).

enta fernandeziana br., Kongl. Svenska Y apsak : 4,

Blastenia Sernandeziana f. validior ps a corneal $ rro

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

peter Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is. 2 (Bot.), 3 (11): 396 (1924) T.

jorgensis C.W. Dodge, Nova Hedwigia 12: 347 (1967) [“1966”] T.

lucens Ny. ) Zahlbr., Deutsch. Siidpolexp. 1901-1903, 7: 29 (1906).

Lecanora elegans f. lucens Cromb., Bot. J. Linn. Soc. 15: 184 (187

microphylla (Hue) Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández msi Isl. 2 (Bot.), 3 (11): 397 (1924).

Lecanora microphylla Hue, Ann. Mycol. 13: 77 (1915) T.

millegrana (Miill. Arg.) Zahlbr., Cat. Lich. Univ. 7: 247 (1931).

Amphiloma millegranum Mill. Arg., Flora 69: 12 (1886).

orthoclada Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter 1sl. 2 (Bot.), 3 (1 1): 398 (1924) T.

pe: Zahlbr., Acta Horti Gothob. 2: 20 (1925) T

pergracilis f. compactior Zahlbr., Acta Horti Gothob. 2: 21 (1925) T.

psoromatis Olech $: Sóchting, Lichenologist 25: 265 (1993) T

pulverulenta (Mill. Arg.) Zahlbr., Cat. Lich. Univ. 7: 168 (1931).

Callopisma pulverulentum Mill. Arg., Flora 72: E (1889).

regalis (Vain.) Zahlbr., Cat. Lich. Univ. 7: 274 (193

Placodium regale Vain., Résult. Voy. Belgica, pe 23 (1903).

rubina Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11): 396 (1924) T

rubina var evolutior Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter 1sl. 2 (Bot.), 3 (11): 396 (1924) T.

rugulosa (Nyl.) Zahlbr., Cat. Lich. Univ. 7: 263 (1931).

Placodium rugulosum Nyl., Amn. Sci. Nat. Bot. pre 4. 35153 (18559) T.

sanguinea (Miill.Arg.) Zahlbr., Cat. Lich. Univ. 7: (19 31).

saxicola (Hoffm.) Nordin, Caloplaca, sect. Gasparrinia 1 Nordeuropa: 87 (1972).

selkirkii Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández East Is]. 2 (Bot.), 3 (11): 395 (1924) E

siphonospora Olech $: Sóchting, Lichenologist 25: 267 (1993) T

pr (NyL.) Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3 (11): 397 (1924).

subcerina var. aurantiaca (Mill. Arg.) Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fe rnández Easter Isl. 2 (Bot.),

311) 397 (1924).

Premi cos Kongl. Svenska Vetenskapsakad. Handl. 57 (6): 47 (1917).

subdimorpha var. leprascens Zahlbr., Kongl. Svenska Vetenskapsakad. Handl. 57 (6): 48 (1917).

subherbidella paritecats C.W. Dodge. In G. Follmann, Nova Hedwigia 14: 264 (1968) [” 1967”].

sublobulata (Ny1.) Zahlbr., Cat. Lich. Univ. 7: 267 (1931).

Caloplaca portó var. striolata Zahlbr., Kongl. Svenska Vetenskapsakad. Handl. 57 (6): 49 (1917).

Callopisma lucens var. leprascens (Zahlbr.) Malme, Ark. Bot. 230A (9): 42 (1926).

tetraspora (Nyl.) H. Ollivier, Mém. Soc. cra es Nat. Cherbourg 37: 140 (1909).

pos . Mycol. 1: 360 (190:

Lit.: Jatta (1906); Zahlbruckner (1917, 1924, cp Santesson (1944); Follmann (1965c, 1968); Poelt $ ap

(1984) Redon yodo Jacobsen $: Kappen (1988); Kirnefelt (1989, 1998); Sgchting $ Vvstedahl (1992); Ott

Sancho (1993); Aptroot $: van der Knapp ( 1993); Poelt € Hinteregger (1993); Olech £ Sechting (1993); >

$ Olech (1995).

alycidium Stirt. (1877)

ctas Calycidiaceae]

cuneatum Stirt., . Phil. Soc. Glasgow 10: 292 (1877).

Lit.: Galloway (1985); Tibell (1987, 1994); Wedin (1993).

33. Candelaria A. Massal. (1852).

[Lecanorales: Candelariaceae]

concolor (Dickson) B. Stein. In F.J. Cohn, Kryptog. -Fl. Schlesien 2(2): 84 (1879).

crawfordii (Mill. Arg.) P. .ó Jorg. é: D.J. Galloway, Lichenologist 24: 407 (1992).

murrayi (C.W.Dodge) Poe

Lit.: Jacobsen £ Kappen Al Joórgensen £ Galloway (1992b); Filson (1992).

4. Candelariella Miill. nn (1894)

eo Candelariaceae]

andicola (Zahl EZ Cat. Lich. Univ. 5: 791 (1928).

antennaria Rásinen, Anales Soc. Ci. Argent. 128: 137 (1939).

aurella (Hoffm.) Zahlbr.. Cat. Lich. Univ. 5: 790 (1928).

coralliza (Ny1.) H. Magn., Svensk Bot. Tidskr. 29: 122 (1939).

121

Gayana Bot. 55(2), 1998.

fuegiensis C.W. Dodge, Nova Hedwigia 12: 344 (1967) [*1966”] T.

lambii Hakul., Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 27: 36 (1954).

ossicola C.W. Dodge, Nova Hedwigia 12: 345 (1967) [“1966”] T.

Poe (NyL ) Lettau, Hedwigia 52: 196 (1912).

isánen, Anales Soc. Ci. Argent. 128: 137 (1939).

vainioana Hakul., Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 27: 100 (1954).

cta (Hoffm.) Miúill. Arg., Bull. Herb. Boissier 2, e 1: 47 (1894).

nthostigma (Pers.) Lettau, Hedwigia 52: 196 (1912

Lit: - Paloalidl (1954), Thomson (1973); Aptroot £ van cd Knapp (1993).

35. Candelariopsis sem pa (1959) comb. inval.

[Teloschistales: Teloschistac

chilena C.W. Dodge, o Hedwigia 12: 348 (1967) [*“1966”] T.

Lit.: Dodge (1967).

36. Canomaculina Elix se Hale (1987)

eii ago Parmeliacea:

ria (Zahlbr.) Elix, Mycotaxon 63: 477 (1997).

Lit.: Elx £ Hale (1987); Elix (1993; 1997).

7. Canoparmelia Elix e Hale (1987)

Um penis ae]

aus dd Ai Mycotaxon 28: 251 a

texana (Tuck ) Elix £ Hale, Mycotaxon 27: 279 (1986).

Lit.: Hale (1976); Elix £ Hale (1986); Elix et al. (1986), Adler (1987); Elix (1993, 1994b).

*Capronia Saccardo (1883

[Chaetothyriales: do eran,

normandina D. Hawksw., Notes RBG Edinburgh 46 (3): 395 (1990).

Lit.: Hawksworth ae

38. Carbonea or Hertel (1983)

[Lecanorales: Lecan e]

assentiens (Ny]l. )Hentel Lecideac. Exs. No. 103 (1984).

eyanosarca (Zahlbr.) Knoph € Leuckert, Nova Hedwigia 59: 000 (1994

Lecidea cyanosarca Lali In C. Skottsberg, Nat. Hist. Juan Fernández Easter IsI. 2 (Bot.), 3 (11): 359 (1924) T.

Lecidella cyan hlbr.) Hertel, Herzogia 2: 254 (1971).

inactiva (Zahlb ) Hertel, Beih. Nova Hedwigia 79: 443 (1984).

Lecidea inactiva Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3 (11): 358 (1924) T.

latypizoides (Ny1.) Knoph £ Rambold,

Lecidea montevidensis Mill. Arg., Rev. Mycol. 37: 4 (1888).

osa (Flórke) Hertel, Mitt. Bot. Miinchen 19: 442 2 (1983).

Lit: my (1984, 1989); Rambold (1989); Knoph £ Leuckert (1994).

39. Catapyrenium Flotow (1850)

[Verrucariales: Verrucariaceae

cinereum ras ap Syst. Lich. Germ.: 325 (1855).

Kremp.) B. Stein. In F.J. Cohn, Krypt.-Fl. aa 2(2): 312 (1879).

Lit.: Breuss EA 1995, 1996b); Breuss € McCune (1994); Pérez (1997).

40. Catillaria A.Massal. (1852).

[Lecanorales: E

ens (Mill. Arg.) Zahlbr., Cat. Lich. Univ. 4: 27 (1926).

Patellara aeruginascens Mill. Arg., Nuovo Giorn. Bot. Ital. 21: 47 (1889).

tatorina aeruginascens (Miill. Arg.) Darb., Wiss. Ergebn. Schwed. qe .

coccinea (C.W. Dodge) Follmann 4 Redon, e 437 (197 ple

Biatorina coccinea C.W. Dodge, Nova pda q 331967 E 19667] T.

coquimbensis Ibr., Acta Horti. Gothob. 2: 10 (19

corymbosa (Hue) IM. Lamb, Rhodora, 56: bi

endochroma (Fée) Zahlbr. In Engler-Prantl, Natiirl. Pflanzenfam. l( de 134 (1905).

(NyL.) Arn., Verhandl. Zool.-Bot. Ges. Wien 20: 455 (1870).

Lecidea intermixta Nyl., Amn. Sci. Nat. Bot. sér. 4.3: 161 (1855).

122

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

ras (Mont.) Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3 (11): 363 (1924).

rmelia varia Fr. var. leucochlora Mont., Amn. Sci. Nat. Bot. sér. 2 , 4: 91 (1835) T

Purmelia leucochlora (Mont.) Mont., Ann. Sci. Nat. Bot. sér. 3.18: 310 (1852).

Berengeria leucochlora (Mont.) Trevis., Spighe e Paglie: 5 (1835).

Lecidea leucochlora (Mont.) Nyl., Amn. Sci. Nat. Bot. sér. 4, 3: 184 (1855).

Bayrhofferia leucochlora (Mont.) Trevis., Riv. Period. Lavori Accad. Padova 5: 69 (1857).

leucochlora ¡gli odos In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is. 2 (Bot.), 3 (11): 364 (1924) T.

melastegia (Nyl.) Zahlbr. f. coto ce Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter 1sl. 2

(Bot.), 3 (11): 362 (1924)

obludens (Nyl.) Zahlbr., Cat. bl Univ. 4: 21 (1926).

Lecidea obludens Nyl., Lich. Fueg. Patag.: 12 (1888) T.

praepallida (Miill. Arg.) Zahlbr., Cat. Lich. Univ. 4: 63 (1926).

Patellaria praepallida Miill. Arg., Miss. Sci. Cap Horn, Lich.: 166 (1888) T.

Biatorina praepallida (Mill. Arg.) Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4 (11): 48 (1912).

theobromina Zahlbr. In C. Skottsberg, Nat. Hist. Juan Pa Easter Isl, 2 (Bot.), 3 (11): 364 (1924) T.

tremellula (Mill. Arg.) Zahlbr., Cat. Lich. Univ. 4: 80 (192

Patellaria tremellula Mill. Arg., Nuovo Giorn. Bot. a 21: 47 (1889).

violascens (Múill. Arg.) Zahlbr., Cat. Lich. Univ. 4: 84 (19

Patellaria violascens Miill. Arg., Nuovo Giorn. Bot. + 21: 46 (1889).

tatorina violascens (Múill. Arg.) Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4 (11): 48 (1912).

Lit.: Zahlbruckner (1925); Follmann (1968a); Follmann $: Redon (1972c); Jacobsen €: Kappen (1988); Quilhot et

al. (1989).

41. Cenozosia A. Massal. (1854)

[Lecanorales: Ramalinaceae]

inanis (Mont.) A. Massal., Neagen. Lich.: 5 (1854).

Ramalina inanis Mont., Ann. Sci. Nat. Bot. sér. 2, 18: 266 (1842).

Dufourea inanis (Mont.) Nyl., Mém. Soc. Imp. Sci. Nat t. Cherbourg 3: 171 (1855).

Desmaziera inanis (Mont.) Follmann £ Huneck, Willdenowia 5: 208 (196 9).

Lit.: Follmann £ Huneck (1969); Bowler (1981); Spjut (1996)

42. Cetraria Ach. (1803) nom. cons.

[Lecanorales: Parmeliaceae]

aculeata (Schreb.) Fr., Nov. Sched. Crit.: 26 (1826)

ericetorum Opiz ssp. patagonica Kirnefelt, Opera Bot. 46: 82 (1979).

islandica (L.) Ach. ssp. antarctica Kárnefelt, Opera Bot. 46: 90 (1979) T.

Cetraria tenuifolia var. pseudoislandica Rásánen, Ann. Soc. Bot. Zool.-Bot. Fenn. “Vanamo” 2 (1): 15 (1932)T.

Cetraria tenuifolia var. spinosa Rásiánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (D): 15 (1932) T.

uric h.) Eckf., Bull. Torrey Bot. Club 22: 240 (1895).

Lit.: Rásánen (1932); Kiirnefelt (1979, 1986); Kiárnefelt et al. (1993).

43. Cetrariella ni áz Thell (1993)

A Parmeliaceae]

lisei (Bory ex Aer Kárnefelt £ Thell, Bryologist 96: 403 (1993).

es Kárnefet et al. (1993b).

44. Cetrelia W.L.Culb. £ C.F.Culb. (1968)

[Lecanorales: Parmeliaceae]

olivetorum (Nyl.) W.L. Culb, £ C.F. Culb., Contr. U. S. Natl. Herb. 34: 515 (1968).

Lit.: Culberson $: Culberson (1968); Elix (1993).

45. Chaenotheca (Th.Fr.) Th.Fr. (1860)

proa e:

la (Ach.) Múill. Arg., Mém. Soc. Phys. Genéve e 360 (1862).

pirenad (Harm.) Lettau, Festschr. Preuss. Bot. Ver.: 27 (1912).

chrysocephala (Turner ex Ach.) Th.Fr., Nova Acta Regiae da Sci. Upsal. 3 (3): 250 (1860).

deludens Tibell, Symb. Bot. Upsal. 27: 95 (1987).

ferruginea (Turner ex Sm.) Migula, Krypt.-Fl. Deutschl., Flecht. 2: 479 (1931).

hispidula eq Cat. Lich. Univ. 1: 567 (1922).

Ach.) Th.Fr., Nova Acta Regiae Soc. Sci. Upsal. 3 (3): 351 (1860).

Lit.: Follmanmn (1962); Tibell (1987, 1994).

Gayana Bot. 55(2), 1998.

46. Chaenothecopsis Vain. (1927)

[Caliciales: Mycocaliciaceae]

haematopus Tibell, Symb. Bot. Upsal. 27: 126 (1987).

sanguinea Tibell, Symb. Bot. Upsal. 27: 151 (1987).

Lit.: Tibell (1987).

47. Chiodecton Ach. (1814)

[Arthoniales: Roccellaceae]

chilense Zablbr., Acta Horti Gothob. 2: 8 (1925) T.

Sollmannii Riedl, Sydowia 17: 87 (1964) T.

hypoleucum Nyl., Amn. Sci. Nat. Bot. sér. 4, 3: 172 (1

48. Chrysothrix Mont. (1852) nom. cons.

[Lecanorales: Chrysotrichaceae]

candelaris (L.) JR. Laundon, Lichenologist 13: 110 (1981).

granulosa G. Thor, Bryologist 91: 362 (1988).

pavonii (Fr.) JR. Laundon, Lichenologist 13: 117 (1981).

Peribotryon pavonii Fr., Syst. Mycol. 3: 288 (1832).

Cilicia noli-tangere Mont., Ann. Sci. Nat. Bot. sér. 2, 2: 375 (1834) T. nom illeg.

Chrysothrix noli-tangere (Mont.) Mont., Ann. Sci. Nat. Bot. sér. 3, 18: 313 (1852) nom illeg.

Arthonia noli-tangere (Mont.) Nyl., Mém. Soc. Imp. Sci. Nat. Cherbourg 3: 189 (1855) nom. illeg.

Crocynia noli-tangere (Mont.) Kremp., Ges. Litt. Lich. 2: 247 (1869) nom. illeg.

Lit.: Laundon (1981); Thor (1988).

49. Cladia Ny]. (1870)

[Lecanorales: Cladiaceae]

aggregata (Sw.) Nyl., C.R. Hebd. Séances Acad. Sci. 83: 88 (1876).

schizopora (Nyl.) Nyl., Rev. Bot. Bull. Mens. 6: 161 (1888).

Lit.: Filson (1992); Stenroos (1995).

50. Cladonia P. Browne (1756) nom. cons.

[Lecanorales: Cladoniaceae]

anserina Ahti, Ann. Bot. Fenn. 29: 68 (1992).

arbuscula (Wallr.) Flot. In Wendt, Thermen Warmbrumn: 94 (1839).

Cladina arbuscula (Wallr.) Hale £ W.L. Culb., Bryologist 73: 510 (1970).

asahinae JW. mson, J. Jap. Bot. 51: 364 (1977) [“1966”].

bacilliformis (Ny.) Gliick, Verhand. Naturhist.-Med. Vereins Heidelberg, N. F. 6: 97 (1899).

bellidiflora (Ach.) Schaer.. Lich. Helv. Spicil.: 21 (1823).

borealis S. Stenroos, Ann. Bot. Fenn. 26: 160 (1989)

cariosa (Ach.) Spreng., Syst. Veg. 4: 272 (1827).

carneola (Fr.) Fr., Lich. Eur. Reform.: 233 (1831).

cenotea (Ach.) Schaer., Lich. Helv. Spicil.: 35 (1823).

cervicornis (Ach.) Flot. ssp. mawsonii (C.W. Dodge) S. Stenroos éz Ahti, Ann. Bot. Fenn. 27: 320 ( 1990).

chlorophaea (Flórke ex Sommerf.) Spreng., Syst. Veg. 4: 273 (1827)

confragosa S.Stenroos, Mycotaxon 59: 270 ( 1996) T.

corni, Ahti 8: Kashiw. In H. Inoue, Stud. Cryptog. S. Chile: 136 ( 1984).

cornuta (L.) Hoffm., Descr. PL. Crypt. 1: tab. 25 (1791).

didyma (Fée) Vain., Acta Soc. Fauna Fl. Femn. 4: 137 (1887).

a. E e A. Evans, Rhodora 52: 95 (1950).

nia furcata var. farinacea Vain. In P. Hariot, J. Bot. (Morot) 1: 2 DT

fimbriata (L.) Fr.. Lich. Eur. Reform.: 222 (1831). : riguida

Surcata (Huds.) Schrad., Spic. Fl. Germ.: 107 (1794).

galindezii Dvstedal, Cyrptog. Bryol. Lichénol. 9 (2): 137 (1988).

gracilis (L.) Willd., FL Berol.: 363 787).

gracilis ssp. (Wulfen) Vain., Acta Soc. Fauna Fl. Fenn. 53 ( 1): 92 (1922).

124

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

Lichen elongatus Wulfen. In Jacquin, Misc. Austriaca 2: 368 (1781) T.

gracilis ssp. vadiviensis Ahti, Ann. Bot. Fenn. 27: 322 (1990

granulosa (Vain.) Ahti, Ann. Bot. Fenn. 23: 205 (1986).

humilis (With.) JR. Laundon, Lichenologist 16: 220 (1984).

laevigata (Vain.) Gyeln., Lichenotheca Parva. No. 148 (1937).

Cladonia sylvatica var. laevigata Vain. In P. Hariot, J. Bot. (Morot) 1: 284 (1887) T.

Cladina laevigata (Vain.) C.W. Dodge, Lich. Fl. Antarct.: 97 (1974) [*1973”].

Cladonia rangiferina var. patagonica Kremp., Verhandl. Zool.-Bot. Ges. Wien 26: 437 (1871).

lepidophora Ahti 8% Kashiwad. In H. Inoue, Stud. Cryptog. S. Chile: 140 (1984) T

luteoalba Wheldon $ A. Wilson, Fl. W. Lancashire: 450 (1907).

macilenta Hoffm., Deutsch. Fl. 2: 126 (1796).

macrophyllodes Nyl., Flora 58: 447 (1875).

merochlorophaea var. no care Sipman, Acta Bot. Neerl. 22: 496 (1973).

mitis Sandst., Clad. Exs. No. 55 (19

Cladina mitis (Sandst.) Hustich, pe Geogr. (Helsinki) 12 (1): 27 (1951).

Cladonia arbuscula ssp. mitis (Sandst.) Ruoss, Bot. Helv. 97: 260 (1987).

Cladina arbuscula ssp. mitis (Sandst.) Burgaz, Nova Hedwigia 59: 400 (1994).

norvegica Tonsberg £ Holien, Nord. J. Bot. 4: 79 (1984).

ochrochlora Flórke, De. Cladon.: 75 (1828).

phyllophora Hoffm., Deutsch. Fl. 2: 126 (1796).

pleurota (Flórke) Schaer., Enum. Crit. Lich. Eur.: 186 pues

pocillum (Ach.) Grognot, Pl. Crypt. Sáone-et-Loire: 82 (1863

pyenoclada (Pers.) Nyl., Bot. J. Linn. Soc. 9: 244 (1967) pi

Cno pyenociada Pers. In Gaudich., Voy. Uranie Bot.: 212 (1828) [“1826”].

) Leight., Ann a Nat. Hist. Ser. 3, 19: 122 (1867).

Po (L.) Hoffm., Deutsch. Fl. pi 121 (179

ramulosa (With.) J.R. Laundon, ld 16: 225 (1984).

rangiferina (L.) F.H. Wigg., Prim. Fl. Holsat.: 90 (1780).

Cladina rangiferina (L.) Nyl., Not. Sállk. Fauna Fl. Fenn Fórh. 5: 110 (1866).

rigida (Hook. f. 8: Taylor) Hampe, Linnaea 28: 216 (1856

robbinsii A. Evans, Trans. Connecticut Acad. Arts 35: 611 (1944).

sarmentosa (Hook. f. £: Taylor) C.W. Dodge, Rep. B.A.N.Z.A.R.E. n.s., 7: 129 (1948).

Cladonia gracilis f. propagulifera Vain., Résult. Voy. Belgica, Lich.: 30 (1903) T.

Cladonia aspera Ahti £ Kashiw. In H. Inoue, Stud. Cryptog. S. Chile: 132 (1984) T.

scabriuscula (Delise) Nyl., Compt. Rend. Hebd. Séances Acad. Sci. 83: 88 (1876).

squamosa Hoffm., Deutsch. F1.2: 125 (1796).

subchordalis A. Evans, Rec. Bryol. Lichénol. 24: 133 (1955).

subdelicatula Asah., J. Jap. Bot. 38: 1 (1963).

subradiata (Vain.) Sandst., Abh. Naturwiss. Vereine Bremen 25: 230 (1922).

subsquamosa Kremp., Vidensk. Meddel. Dansk. Naturhist. Foren, ses 5: 366 (1874).

subsubulata Nyl., Compt. Rend. Hebd. Séances Acad. Sci.83: 88 (187

Cladonia auerii Rásinen, Ann. Bot. Soc. Zool.-Bot. Fenn. da 53 (1952) T:

Cladonia auerii f. crassa Rásinen, Ann. Bot. Soc. Zool-Bot Fenn. “Vanamo” 2 (1): 53 (1932) T.

sulphurina (Michx) Fr., Lich. Eur. Rele 237 (1831).

symphycarpa (Flórke) Fr., Nov. Sched. Crit. Lich. 8-9: 20 (1826).

tessellata Ahti £ Kashiw. In H. Inoue, Stud. Cryptog. S. Chile: 145 (1984) T.

umbricola Tónsberg 8 Ahti, Norw. J. Bot. 27: 307 (1980

ustulata (Hook. £.£ Taylor) Leight., Ann. Mag. Nat. Hist. Ser. 3, 19: 109 (1867).

weymouthii F. Wilson ex A.W. Archer, Muelleria 6: 94 (1985).

Cladonia pleurota var. digitiiformis Rásinen, Anmn. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 51 (1932).

Lit.: Ahti (1961, 1993); Ahti € Kashiwadani (1984); Ahti et al (1990): Stenroos £: Ahti (1990, 1992); Stenroos

(1993, 1995, 1996); Stenroos et al. (1992, 1997); Hyvónen et al. (1995).

51. Coccocarpia Pers (1827)

[Lecanorales: Coccocarpiaceae]

domingensis Vain., An. Acad. Sci. Fenn. ser. 2, 6: 104 (1814).

erythroxyli (Spreng.) Swinscow €: Krog, Norw. J. Bot. 23: 256 (1976).

palmicola (Spreng.) Arv. £ D.J. Galloway, Bot. Notiser 132: 242 (1979).

Lit.: Arvidsson (1983).

52. Coccotrema Miill. Arg. (1889)

[Pertusariales: Coccotremataceae]

Gayana Bot. 55(2), 1998.

cucurbitula (Mont.) Miill. Arg., Nuovo Giorn. Bot. Ital. 21: 51 (1889).

Pertusaria cucurbitula Mont. In C. Gay, Hist. Chile, Bot. 8: 200 (1852) T.

33. Coelopogon Brusse $: Kiárnefelt (1991)

[Lecanorales: Parmeliaceae]

epiphorellus (Nyl.) Brusse € Kárnefelt. Mycotaxon 42: 39 (1991).

etraria epiphorella Nyl. In J. Crombie, Bot. J. Linn. Soc. 15: 227 (1876) T.

Cornicularia epiphorella (Nyl.) DR., Ark. Bot. 20A (11): 34 (1926).

Coelocaulon epiphorellum (Nyl.) Kárnefelt, Opera Bot. 86: 64 (1986).

Cetraria aculeata var. gracilenta Krempelh., Verhandl Zool.-Bot. Ges. Wien 18: 315 (1868) T.

Cetraria gracilenta (Krempelh.) Vain., Résult. Voy. Belgica, Lich.: 13 (1903).

Lit.: Du Rietz (1926); Lamb (1964); Kárnefelt (1986); Brusse £ Kiáirnefelt (1991).

54. gonium Ehrenb. ex Nees (1820)

[Gyalectales: Gyalectaceae]

implexum Ny]l., Ann. Sci. Nat. Bot. sér. 4, 16: 92 (1862).

interplexum Nyl., Ann. Sci. Nat. Bot. sér. 4, 16: 92 (1862).

linkii Ehrenb, Horae Phys. Berol.: 120 (1820).

velutinum Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is, 2 (Bot.), 3 (11): 332 (1924) T.

Lit.: Zahlbruckner (1924); Hosseus (1939).

55. Collema Wiggers (1780) nom. cons.

[Lecanorales: Collemataceae]

Pycnocarpoides (Jatta) Zahlbr., Cat. Lich. Univ. 3: 45 (1924).

Synechoblstus pychnocarpoides Jatta, Malpighia 20: 13 (1906) T.

santessonii Degel., Symb. Bot. Upsal. 20: 84 (1974) T

tenax var. substellatum (H. Magn.) Degel., Symb. Bot. Upsal. 20: 47 (1974).

Lit.: Degelius (1954, 1974, 1986).

56. Cresponea Egea £ Torrente (1993)

[Arthoniales: Roccellaceae]

follmannii (C.W. Dodge) Egea de Torrente, Mycotaxon 48: 317 (1993).

Lecanactis follmannii C.W. Dodge, Nova Hedwigia 12: 327 (1967) [“1966”] T.

Lit.: Egea $: Torrente (1993b, 1994).

57. Cyphelium Ach. (1815)

[Caliciales: Caliciaceae]

inquinans (Sm.) Trevis., Flora 45: 3 ( 1862).

Lit.: Tibell (1987).

58. Cystocoleus Thwaites (1849)

[Mitosporic fungi]

beneus (Dillw. waites,

Lit.: Follmann £ Redon (1972a); Jacobsen 4 Kappen (1988).

*Dacampia A. Massal. ( 1853)

[Dothideales: Dacampiaceae

rufescentis (Vouaux) D. Hawksw., Notes Roy. Bot. Gard. Edinburgh 43 (3): 497 (1986).

Lit.: Hawksworth ( 1986); Alstrup $: Hawksworth (1990); Wedin (1994).

*Dactylospora Kórb. (1 855)

[Ascomycota - incert. sed. Dactylosporaceae]

126

Checklist of Chilean lichen: GALLOWAY, D.J. € W. QUILHOT

australis Triebel £ Hertel, Biblioth. Lichenol. 35: 205 (1989).

frigida Hafellner, Herzogia 7: 116 (1985).

lobariella (Nyl.) Hafellner, Beih. Nova Hedwigia 62: 118 (1979).

rygmaea (Nyl.) S. Kondratyuk £ D.J. Galloway, Biblioth. Lichenol. 57: 336 (1995).

Lit.: Hafellner (1979, 1985); Triebel (1989); Kondratyuk £ Galloway (1995).

59. Degelia Arv. £ D.J. ceo (1981)

[Lecanorales: Pannariac

duplomarginata (P. pes $: Henssen) Arv. € D.J. Galloway, o 13: 39 (1981).

gayana (Mont.) Arv. £ D.J. Galloway, Lichenologist 13: 44 (1981).

Parmelia gayana Mont., Ann. Sci. Nat. Bot. 3: 58 (1849) T.

Pannaria gayana (Mont.) A. Massal., Geneac. Lich.: 19 (1854).

Coccocarpia gayana (Mont.) Nyl., Syn. Meth. Lich. 2: 41 (1869).

Trachyderma gayanum (Mont.) Trevis., Lich. Veneta: 98 (1869).

Parmeliella gayana (Mont.) Miill. Arg., Flora 69: 286 (1886).

Pannaria chilensis Fée. In A. Massalongo, Ric. Auton. Lich. Crost.: 111 (1852) T.

annaria pycnophora var continentalis Rásiinen, Ann. Bot. Soc. Z agil Fenn. “Vanamo” 2 (1): 46 (1832) T.

uviatilis P.M. Jórg. £ P. James, Biblioth. Lichenol. 38: 272 (1990

Lit.: Arvidsson € Galloway (1981); nn £ James (1990); dis K£ Galloway (1992a).

60. Dendriscocaulon Nyl. (1888)

[Peltigerales: Lobariaceae]

callithamnion (Taylor) D.J. Galloway £ Quilhot, comb. n

Basionym: Sticta calithamnia Taylor, Lond. J. Bot. 6: 183 (1847) T

pei calithamnium (Taylor) Nyl., Syn. Meth. Lich. 1 ea 126 (1860).

na callithamnia (Taylor) Miill. Arg., Flora 71: 164 (188

E rentar 1847) Zahlbruckner (1924); James € Henssen (1976).

61. Dibaeis Clem. (1909)

[Lecanorales: Icmadophilaceae]

soluta (Tuck.) Kalb £ Gierl, Herzogia 9: 613 (1993).

Lit.: Gierl 8: Kalb (1993); Rambold et al. (1993).

62. Dictyonema C.Ag. ex Kunth (1822)

[Aphyllophorales: ae Cora

glabratum (Spreng.) D. Hawksw., Lichenologist 20: 101 (1988)

poes prriasd Spreng., Kongl. Vetensk. Acad. Handl. 1820: 51 (1820).

Thelephora pavonia Sw., Fl. Ind. Occid. 3: 1930 (1806).

am sena iia (Sw. ) Parmasto. In Follmann £ Redon, Wilkdepowria 6: 456 (1972) nom. inval.

rei (Nyl.) Henssen, Symb. Bot. Upsal. 18 (1): 109 (1963).

Lite teta (1978); Hawksworth (1988); Brodo (1995); PioVano et al. (1995).

Norman (1853)

[becanorales: vee

oreina (Ach.) Norman, Nyt. Mag. Naturvidensk. 7: 231 (1852).

Lit.: Mayrhofer et al. (1996).

64. Dimerella Trevis. pe

[Gyalectales: Gyalectaceae]

lutea (Di ido) Trevi. Ric. Ist. Lomb. Sci. Lett. 13: 65 (1880).

Lit.: Vezda £ Farkas (1988).

65. Diploschistes Norman (1853)

[Pertusariales: Thelotremataceae]

anactinus (Nyl.) Zahlbr., Hedwigia 31: 14 (1892).

actinostomus Pers. ex Ach. Zahlbr., Hedwigia 31: 34 (1892).

conceptionis Vain., Hedwigia 38: 189 (1899) T

diacapsis (Ach.) Lumbsch, Lichenologist 20: 20 (1988).

euganeus (A. Massal.) aa biqpuan Zool. Bot. Ges. Wien 69: 96 (1919).

muscorum ssp. bartlettii L h, Herzogia 7: 602 (1987).

127

Gayana Bot. 55(2), 1998.

scruposus (Schreb.) Norman, Nyt. Mag. Naturvidensk. 7: 232 (1853).

Lit.: Zahlbruckner (1926); Follmann (1965b); Lumbsch (1987, 1989, 1993); Guderley « Lumbsch (1996).

66. Diplotomma Flot. (1849)

pepsi Physciaceae]

mae Follmann, Bol. Univ. Chile 7 (6): 60 (1965) T.

LE: peta (1965a, 1968a).

67. Dirinaria (Tuck.) Clem. (1909)

[Lecanorales: Physciaceae]

picta (Sw.) Clem. á Shear, Gen. Fung.: 323 (1931).

Lit.: Zahlbrukner (1926); Awasthi (1975); Aptroot (1987); Swinscow 4 Krog (1988).

68. Dolichocarpus R. Sant. (1949)

[Arthoniales: Roccellaceae]

sis R. Sant., Svensk Bot. Tidskrift 43: 552 (1949) T.

Lit.: Santesson (1949); Follmann (1968a).

69. Echinoplaca Fée (1824)

[Ostropales: Gomphillaceae]

heterella (Stirt.) R. Sant., Symb. Bot. Upsal. 12: 372 (1952).

Lit.: Santesson (1952); Follmann (1965b).

70. Endocarpon Hedwig (1789)

feo smc: Verrucariaceae]

um Zahlbr., Cat. Lich. Univ. 8: 74 (1931).

Endocarpon Malme, Ark. Bot. 22A (1): 10 (1928) non Vain.

pallidulum Ny]. In Hue, Nouv. Arch. Mus. sp e sér. 3, 4:106 (1892).

pallidum Ach., Lichenogr. Universalis: 301 (18

pusillum Hedw., Descr. Micr. Anal. Musc. a 2: 56 (1789).

peon? (Jatta) Zahlbr., Cat. Lich. Univ. 1: 249 (1921).

ocarpon scottianum Jatta, Malpighia 20: 13 (1906) T

Lit.: Jasa (1906): McCarthy (1991).

71. Endocena Cromb. (1876)

poor; Acarosporaceae]

Cromb., Bot. J. Linn. Soc. 15: 226 (1876).

Lit.: - Coabie (1876),

*Endococcus Nyl. (1855)

[Dothideales: fam. incert. sed.

propinquus (Kórb.) D. Hawksw., Bot. Not. 132: 287 (1979).

Lit.: Hawksworth (1979); Triebel (1989).

72. Enterographa Fée (1825)

[Arthoniales: Roccellaceae]

albinea (Nyl.) C.W. Dodge. In G. Follmann, Nova Hedwigia 14: 230 (1968).

Stigmatidium albineum Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 169 (1855) T.

Chiodecton albineum (Nyl.) Zahlbr., Cat. Lich. Univ. 2: 473 (1924).

atacamensi: . Dodge, Nova Hedwigia 12: 323 (1967) [“1966”] T

chilensis (Zahlbr. ) C.W. Dodge. In G. Follmann, Nova Hedwigia 14: 230 ( 1968).

ropa chilense Zahlbr., Acta Horti. Gothob. 2: 8 (1925) T

MW. ge, Nova Hedwigia 12: 324 (1967) [1966 7.

Lit.: e ( o Follmann (19684); Follmann $: Redon (1972c).

Mill. Arg. = Thelotrema (see Salisbury 1972).

cerei eS Do In Riedl £ Schiman-Czeika, Sydowia 17: 89 (1964) T.

Melam, poe cerei (Schiman-Czeika) Redon 4 Follmann, Philippia 1: 186 (1972).

Lit.: Salisbury (1972

73. Ephebe Fr. (1825)

[Lichinales: Lichinaceae]

128

Checklist of Chilean lichen: GALLOWAY, D.J. $ W. QUILHOT

ocellata Henssen, Symb. Bot. Upsal. 18(1): 51 (1963).

LjE: dera (19630).

74. Erioderma Fée 0

[Lecanorales: Pannariaceae]

leylandi (Taylor) Mal. Arg., Flora 71: 24 (1888).

Sticta leylandi Taylor, Lond. J. Bot. 6: 179 ón

tedio chilense Mont., Amn. Sci. Nat. Bot. sér. 3, 18: 309 (1852) T.

m D.J. Galloway « P.M. Jorg., ileso? 139 (1975).

E: A et al. (1983); Jorgensen $: Galloway (1992a).

75. Everniastrum Hale ex Sipman (1986)

[Lecanorales: Parmeliaceae]

americanum (Meyen K Flot.) Hale ex Sipman, Mycotaxon 26: 239 (1986).

cirrhatum (Fr.) Hale ex Sipman, Mycotaxon 26: 239 (1986)

sorocheilum (Vain.) Hale ex Sipman, Mycotaxon 26: 242 (1986).

Lit.: Culberson € Culberson (1981); Sipman (1980, 1986); Elix et al. (1994c).

76. Everniopsis Nyl. (1860)

[Lecanorales: Parmeliaceae]

trulla (Ach.) Nyl., Syn. Meth. Lich. 1 (2): 375 (1860).

Parmelia trulla Ach., Meth. Lich.: 256 (1803).

ticta rugulosa Taylor, Lond. J. Bot. 7: 177 (1847).

Lit.: Nylander (1860).

77. Farnoldia Hertel (1983)

[Lecanorales: Porpidiaceae]

dissipabilis (Nyl.) Hertel, Mitt. Bot. Miinchen ce 443 (1983).

Lecidea pei Nyl., Flora 57: 314 (1874).

Lecidea nelsonii C.W. Dodge, Publ. Inst. Antart. Chileno 6: 5 (1966).

cf. micropsis (A. Massal.) Hertel., Mitt. Bot. Miinchen sá 443 (1983).

similigena (Nyl.) Hertel, Mitt. Bot. Miinchen 19: o 89).

Lit.: pra (1981, 1984, 1989); Pietschmann (1990

78. Fellhanera Vezda Ada

[Lecanorales: Pilocarpace

uteillei (Desm.) Vezda, Folia Geobot. Phytotax. 21: 214 (1986).

Lit.: Vezda (1986); Farkas $ Sipman (1993); Liicking ef al. (1994); Ferraro € Vezda (1995); Sérusiaux (1996).

79. Flavoparmelia Hale (1986)

[Lecanorales: Parmeliaceae]

caperata (L.) Hale, Mycotaxon 25: 603 (1986).

ferax (Miill. Arg.) Hale, Mycotaxon 25: 604 (1986).

gerlachei (Zahlbr.) Hale, Mycotaxon 25: 604 (1986).

tidota (Hook. f. 8: Taylor) Hale, Mycotaxon 25: 605 (1986).

Lit.: Hale (1976, 1986); Elix (1993, 1994d); Elix £ Johnston (1988b).

80. Flavopunctelia Pai Hale (1984)

[Lecanorales: Parmeliac

eel a ) Hale, Mycotaxon 20: 681 (1984)

rmelia caperata f. crispa Meyen á Flot., Nov. Actorum Acad. Caes. Leop.- Carol. Nat. Cur. 19: 229 (1843) T.

Pismelí alutacea Zahlbr., Ann. Mycol. 29: 85 (1931) T.

Parmelia soredica var. purata Zahlbr., Amn. Mycol. 29: 85 (1931) T.

ci looseri Rásánen, Revista Univ. (Santiago) 21: 141 (1936) T

dica (Nyl.) Hale. Mycotaxon 20: 682 (1984).

Lit.: “Hale (1980, 1984).

81. Follmanniella Peine £ B. Werner (1995)

[Arthoniales: Roccellaceae

llata Peine £ B. Werner, Flechten Follmann: 289 (1995) T.

Lit.: Peine 8: Werner (1995).

Gayana Bot. 55(2), 1998.

82. Fuscidea V. Wirth € Vezda (1972)

[Teloschistales: Fuscideaceae]

asbolodes (Nyl.) Hertel £z V. Wirth, Beih. Nova Hedwigia 79: 454 (1984).

Lecidea asbolodes Nyl. In J. Crombie, J. Bot. 14: 21 (1876).

pegar Ago Arg.) Hertel, Beih. Nova Hedwigia 79: 454 (1984

impolita Mill. Arg., Miss. Sci. Cap Horn, Lich.: 165 ( 1 888) T.

M.: e pepe 1987b, 1989).

83. Graphina Miill. Arg. (1880)

[Ostropales: Graphidaceae]

Jfissurinoidea (Vain.) Zahlbr., Cat. Lich. Univ. 2: 406 (1923).

?gracillima (Kremp.) Mill. Arg., Rev. Mycol. 10: 120 (1888).

Graphis gracillima Kremp., Flora 61: 520 (1878).

hololeuca (Mont.) Miill. Arg., Flora 65: 386 (1882).

insignis (Vain.) Zahlbr., Cat. Lich. rl 2: 410 (1923).

?lorentzii Múll. Arg., Flora 72: 67 (188

platycarpa (Eschw.) Zahlbr., Sitzun o Kaiserl. Akad. Wiss. Math.-Naturwiss. CL, Abt. 1, 111 (1): 385 (1902).

da Zahlbr., Acta Horti Gothob. 2: 4 (1925) T.

Lit.: Follmann (1962).

84. Graphis Adans. (1763)

[Ostropales: Graphidaceae]

brevissima Fée, Bull. Soc. Bot. France 21: 27 (1874).

chilensis Zahlbr., Cat. Lich. Univ. 2: 297 (1923).

Graphis petraea Nyl. Ann. Sci. Nat. Bot. sér. 4, 3: 186 (1855) T. non Wallr.

dumastii (Fée) Spreng., Syst. Veg. 4 (1): 254 (1827).

duplicata Ach., Syn. Meth. Lich.: 81 (1814).

duplicata Ach. var. peruviana (Fée) Zahlbr., $ Lich. Univ. 2: 303 (1923).

elegans (Sm.) Ach., Syn. Meth. Lich.: 81 (18

Sollmannii C.W. Dodge, Nova Hedwigia 12: o ts [“1966”] T.

intricata Fée, Essai Crypt. Ecorc.: en (1834).

lineola Ach., Lich. Univ.: 264 (181

seripta (L.) Ach., Kongl. Vetensk. pea ss Handl.: 145 (1809).

tenella Ach., Syn. Meth. Lich.: 81 (181

zapallarensis e, Nova Aia 12: 321 (1967) [“1966”] T.

Lit.: Zahlbruckner ( 1926, 1933) Dodge (1967); Follmann (1962, 1968a).

85. Gyalecta Ach. (1808)

[Gyalectales: Gyalectaceae ]

skotts. ezda, Annot. Zool. Bot. 13: 3 (1965) T.

Lit.: Vezda (1969).

86. Gyalectidium Miill. Arg. (1881)

a Gomphillaceae]

filicinum Miill. Arg., Flora 64: 101 (1881).

Lit: Mica (1952): Follmann (1965b); Farkas é£ Sipman (1993).

87. Gyalectina Vezda (1969)

[Gyalectales: Gyalectaceae

saxatilis Vezda, Folia Geobot., Phytotax. 4: 445 (1969).

Lit.: Vezda (1969).

88. Gymnographopsis C.W. Dodge (1967)

[Ostropales: ?Graphidaceae]

cerei Follmann, Nova Hedwigia 14: 225 (1967) T.

chilena C.W. Dodge, Nova Hedwigia 12: 320 (1967) [**1966”] T.

Sollmannii C.W. Dodge, N ova Hedwigia 12: 320 (1967) [1966] T.

Lit.: Dodge (1967); Follmann (1968a); Follmann €: Redon (1972c); Egea « Torrente (1996).

89. Haematomma A. Massal. (1852)

[Lecanorales: Haemato:

chilenum C.W. Dodge, Nova Hedwigia 12: 343 (1967) [*1966”] T.

130

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

erythromma (Nyl.) Zahlbr., Kongl. Svenska Vetenskapsakad. Handl. 57 (6): 37 (1917).

fenzlianum A. Massal., Mem. I. R. Ist. Veneto 10: 58 (1861).

matomma campanaense Redon £ Walk., Anales Mus. Hist. Nat. Valparaíso 11: 30 (1978) T.

hilare Zahlbr., Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 104: 350 (1941).

nothofagi Kalb K Staiger, Biblioth. Lichenol. 59: 139 (1995).

Haematomma puniceum ssp. magellanicum Asah., J. Jap. Bot. 39: 213 (1964).

Lit.: Jacobsen £ Kappen (1988); Staiger £ Kalb (1995); Vobis et al. (1995).

90. Heppia Nágeli ex A. Massal. (1854)

[Lecanorales: Heppiaceae

chilensis Jatta, Malpighia 20: 8 (1906) T.

lutosa (Ach.) Nyl., Syn. Meth. Lich. 2: 45 (1869).

phylliscum (Nyl.) Hue, Mém. Soc. Nac. Sci. Nat. Cherbourg 36: 19 (1907).

Lecanora phylliscum Ny]l., Ann. Sci. he Bot. sér. 4, 3: 154 (1855) T

subglebosa (Mill. Arg.) LM. Lamb, Anales Parques Nac. 7: 48 (1959) [* 1958”.

Lit.: Lamb (1959); Follmann (1965b); Wetmore (1970); Henssen (1994, 1995).

91. Hertella Henssen (1985)

[Peltigerales: Placynthiaceae]

chilensis Henssen, Mycotaxon 22: 393 (1985) T.

Lit.: Henssen (1984).

92. Heterodermia Trevis. (1868)

[Lecanorales: Physciaceae]

chilensis (Kurok.) Swinscow $ Krog, Lichenologist 8: 115 (1976).

dactyliza (Nyl.) Swinscow £ Krog, Lichenologist 8: 117 (1976).

Physcia speciosa Fr. var. dactyliza Nyl., Sym. Meth. Lich. 1(2): 417 (1860).

Anaptychia dactyliza (Nyl.) Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11):

3 (1924)

follmannii Sipman, Flechten Follmann: 333 (1995) T.

galactophylla (Tuck.) W.L. Culb., Bryologist 69: 482 (1967).

lamelligera (Taylor) Follmann 4 Redon, Willdenowia 6: 446 (1972).

Parmelia lamelligera Taylor, Lond. J. Bot. 6: 169 (1847).

leucomelos (L. ) Poelt ssp. boryi (Fée) Swinscow 8 Krog, Lichenologist 8: 124 (1976).

) Follmann € Redon, Willdenowia 6: 446 (1972).

Heterodermia neoleucomelaena f. cireinalis (Zahlbr.) Follmano $: Redon, Willdenowia 6: 446 (1972).

magellanica (Zahlbr.) Swinscow £ Krog, Lichenologist 8: 130 (1976).

Anaptychia magellanica Zahlbr., Kongl. Svenska Vetenskapsakad. Handl. 57 (6): 54 (1917) E

Anaptychia dactyliza f. pectinata Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is. 2 (Bot.), 3

(11): 403 (1924) T.

Anaptychia pectinata (Zahlbr.) R. Sant., Ark. Bot. 31A (7): 14 (1944).

Anaptychia magellanica var. pectinata (Zahlbr.) Kurok., Beih. Nova Hedwigia 6: 67 (1962).

multiciliata (Kurok.) Follmann € Redon, Willdenowia 6: 446 (1972).

Anaptychia multiciliata Kurok., Beih. nd Hedwigia 6: 72 (1962) T.

pinnata Sipman, Flechten Follmann: 330 (199

vulgaris (Vain.) Follmann £ Redon, ad ES 6: 447 (1972).

Lit.: Zahlbruckner (1924); Kurokawa (1962, 1973, 1998); Follmann £ Redon (1972c); Swinscow £ Krog (1976,

1988); Aptroot (1987); Sipman (1995).

93. Heteroplacidium Breuss (1996)

[Verrucariales: Verrucariaceae]

acarosporoides (Zahlbr.) Breuss, Ann. Naturhist. Mus. Wien 98 B Suppl.: 40 (1996).

podolepis (Breuss) Breuss, Ann. Naturhist. Mus. Wien 98 B Suppl.: 40 (1996).

Lit.: Breuss (1996b).

94. Himantormia 1.M. Lamb (1964)

[Lecanorales: Parmeliaceae]

lu (Hue) LM. Lamb, Br. Antarct. Surv. Sci. Rep. 38: 18 (1964).

Lit.: Lamb (1964); Redon (1985); Jacobsen £ Kappen (1988).

131

Gayana Bot. 55(2), 1998.

95. Homothecium A. Massal. (1855)

[Lecanorales: Collemataceae]

chilense (Rásánen) Henssen, Lichenologist 3: 36 (1965).

Leciophysma chilense Rásánen, Revista Univ. (Santiago) 22: 206 (1937) T.

? intermedium Henssen, Bot. Notiser 132: 272 (1979).

opulentum (Mont.) A. Massal., Alc. Gen. Lich.: 8 (1853).

Collema opulentum Mont., Ann. Sci. Nat. Bot. sér. 3, 18: 313 (1852) T.

opulentum var. redonii Henssen, Bot. Notiser 132: 274 (1979) T

patagonicum (Rásánen) Henssen, Lichenologist 3: 36 (1965).

Leciophysma patagonicum Rásiánen, Anales Soc. Ci. Argent. 128: 145 (1939).

orediosum Henssen, Bot. Notiser 132: 271 (1979) T. :

Lit.: Henssen (1965, 1979).

96. Hubbsia W.A.Weber (1965)

[Arthoniales: Roccellaceae]

langei Follmamn, Biblioth. Lichenol. 67: 17 (1997).

Lit.: Follmann (1997).

97. Hyperphyscia Miill. Arg. (1894)

[Lecanorales: Physciaceae]

adglutinata (Flórke) H. Mayrhofer $ Poelt, Herzogia 5: 62 (1979).

Lit.: Scutari (1991, 1992).

98. Hypogymmnia (Nyl.) Nyl. (1896)

[Lecanorales: Parmeliaceae]

lugubris (Pers.) Krog, Norsk Polarinst. Skr. 144: 99 (1968).

mundata (Nyl.) Rassad., Bot. Mater. Gerb. Bot. Inst. Kamarova Akad. Nauk SSSR 11: 11 (1956).

subphysodes (Kremp.) Filson, Vict. Nat. 87: 325 (1970).

turgidula (Bitter) Elix, Brunonia 2: 238 (1980).

Lit.: Elix (1980, 1992).

99. Hypotrachyna (Vain.) Hale (1974)

[Lecanorales: Parmeliaceae]

bogotensis (Vain.) Hale, Smithsonian Contr. Bot. 25: 23 (1975).

brevirhiza (Kurok.) Hale, Smithsonian Contr. Bot. 25: 26 (1975).

dactylifera (Vain.) Hale, Smithsonian Contr. Bot. 25: 30 (1975).

densirhizinata (Kurok.) Hale, Smithsonian Contr. Bot. 25: 31 (1975).

Jlavovirens (Kurok.) Hale, Smithsonian Contr. Bot. 25: 37 (1975).

Parmelia flavovirens Kurok. In Hale de Kurok, Contr. U.S. Natl. Herb. 36:176 (1964) T.

Sormosana (Zahlbr.) Hale, Smithsonian Contr. Bot. 25: 38 (1975).

imbricatula (Zahlbr.) Hale, Smithsonian Contr. Bot. 25: 41 (1975).

laevigata (Sm.) Hale, Smithsonian Contr. Bot. 25: 44 (1975).

Parmelia boliviana var. cephalota Zahlbr.. Acta Horti Gothob. 2: 16 (1925) T.

oostingii (Dey) Hale, Smithsonian Contr. Bot. 25: 51 (1975).

psuedosinuosa (Asah.) Hale, Smithsonian Contr. Bot. 25: 58 (1975).

(197

4 66

Parmelia chilena Nyl., Flora, Jena 68: 612 (1885) T.

Parmelia ec. cens Kurok. In M. Hale £ S.Kurokawa, Contr. U.S. Natl. Herb. 36: 167 (1964) T.

subphysodalica (Hale) Hale, Smithsonian Contr. Bot. 25: 66 (1975).

Parmelia subphysodalica Hale, Phytologia 28: 268 (1974) T.

Lit.: Hale (1975); Elix (1993, 1994e).

100. Icmadophila Trevis. (1852)

[Leotiales: Icmadophi aceae]

ericetorum (L.) Zahlbr., Wiss. Mitt. Bosnien $: Herzegovina 3: 605 (1895).

Lit.: Rambold er al. (1993).

132

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

101. Ingaderia Darb. (1897)

gracillima (Kremp.) Feige £ Lumbsch, Mycotaxon 48: 386 (1993).

Roccella gracillima Kremp., Verhandl. Zool.-Bot. Ges. Wien 26: 442 (187

Dictyographa gracillima (Kremp.) Darb., Ber. Deutsch. Bot. Ges. 15: irciia nom. illeg.

Darbishirella gracillima (Kremp.) Zahlbr. ex Darb., Bibl. Bot. 9 (45): 13 (1898

Lit.: Huneck Follmann (1967b); Follmann (1968a); Follmann € Redon (19720): Tehler (1990); Feige «

Lumbsch (1993).

102. Immersaria Rambold «€ Pietschmann (1989)

[Lecanorales: Porpidiaceae]

ovocarpa (Ach.) Rambold £ Pietschmann. In G. Rambold, Biblioth. Lichenol. 34: 239 (1989).

idea athroocarpa Ach., Meth. Lich.: 41 (1803).

Lit.: Hertel (1989); Rambold (1989).

103. Ingvariella Guderley £ Lumbsch (1997)

[Ostropales: Thelotremataceae]

bispora (Bagl.) Guderley £ Lumbsch, Nova Hedwigia 64: 152 (1997).

Urceolaria bispora Bagl., Nuovo Giorn. Bot. Ital. 3: 246 (1871).

oschistes bisporus (Bagl.) Steiner, Sitzungsber. Kaiserl. Akad. Wiss. Math.-Naturwiss. Cl. 102: 155 (1893).

Dipl

Lit.: Follmann (1965b); Lumbsch (1989); Guderley et al. (1997).

104. Ionaspis Th.Fr. (1871)

Sis Hymeneliaceae]

fuegensis P.M. Jórg. 8 R. Sant., Nord. J. Bot. 9: 431 (1989).

Ets pon $ Santesson (1989).

105. Josefpoeltia S. Kondr. £« Kárnefelt (1997)

Pra Teloschistaceae

op S. Kondratyuk £ Kárnefelt, Biblioth. Lichenol. 68: 23 (1997).

SO. redliasa. SK dratyuk £ Kárnefelt, Biblioth. Lichenol. 68: 24 (1997) T.

Lit.: Kondratyuk E Kamefel (1997).

*Lauderlindsaya J.C. David € D. Hawksw. (1989)

[Verrucariales: Verrucariaceae

borreri (Tul.) J.C. David € D. Hawksw., Sydowia 41: 116 (1989).

Lit.: David € Hawksworth (1989); Diederich € Sérusiaux (1993).

06. Lecanactis Kórb. (1855) nom. cons.

dera Roccellaceae

?abietina vane ) Kórb., Syst. Lich. Germ.: 275 (1855).

pse ylacea Redon $: Follmann, Philippia 1: 187 (1972) T

Lit.: Redon £ rima (1972b); Tehler (1992); Egea £ Torrente ( 1994).

107. Lecania A. Massal. (1853)

[Lecanorales: Bacidiaceae]

andina C.W. Dodge, Nova Hedwigia 12: 341 (1967) [“1966”] T.

brialmonti (Vain.) Zahlbr. In Engler-Prantl, Natiirl. Pflanzenfamil. (1): 205 (1907).

Lecanora brialmontii Vain., Résult pr Belgica, Lich.: 17 (1 :

cariosa (Hue) Darb., Br. Antarct. (“Terra Nova”) Exp. 1910, Nat. Hist. Rep., Bot. 3: 57 (1923).

chilena C.W. Dodge, Nova Hedwigia 12: 338 (1967) [“1966”] T

gerlachei (Vain.) Darb., Br. Antarct. (“Terra Nova”) Exped. 1910, Nat. Hist. Rep., Bot. 3: 57 (1923).

hookeri C.W. Dodge, Nova Hedwigia 12: 341 (1967) [“1966”].

Baer (Mont.) Zahlbr., Cat. Lich. Na 5: 7353 (1928).

¡a varia var. leucochlora Mont., Ann. Sci. Nat. Bot sér. 2, 4: 91 (1835

Lit.: da (1967); Redon (1985); Vvstedal (1986) Jacobsen Kappen (1988): poca et al. (1989).

108. Lecanographa Egea £: Torrente (1994)

[Arthoniales: Roccellaceae]

aff. arinosa (Hepp) Egea 82 Torrente, Biblioth. Lichenol. 54: 130 (1994).

grapha farinosa Hepp. In Stizenb., Nov. Actorum Acad. Caes. Leop.-Carol. German. Nat. Cur. 32 (4):

133

Gayana Bot. 55(2), 1998.

subhydrophila (Follmann $ Vezda) Egea £ Torrente, Biblioth. Lichenol. 54: 157 (1994).

Lecanactis subhydrophila Follmamn £z Vezda, Philippia 3: 271 (1977) T.

Lit.: Follman 8: Vezda (1977); Egea £ Torrente (1994); Tehler $ Egea (1997).

109. Lecanora Ach. (1810)

[Lecanorales: Lecnoraceae]

addubitata Kremp., Flora, Jena 56: 473 (1873) T.

Lecanora herteri Zahlbr., Revista Sudamer. Bot. 1: 56 (1934).

Lecanora homalea H. Magn., Acata Horti Gothob. 18: 217 (1950).

Lecanora subatra Rásánen, Revista Sudamer. Bot. 5: 66 (1938).

albellina Mill. Arg. var. validior Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter 1sl. 2 (Bot.), 3

(11): 380 (1924) T.

argentata (Ach.) Degel., Ark. Bot. 24A :78 (1931).

atrynea (Ach) Róhl., Dtsch. Fl. 3 (2): 82 (1813).

icanum (Rásiánen) Zahlbr., Cat. Lich. Univ. 10: 487 (1940).

Squamaria austroamericana Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 25 (1932) T.

austrolitoralis Zahlbr., Acta Horti Gothob. 2: 14 (1905T.

avium (Zahlbr.) Hertel, Beih. Nova Hedwigia 79: 447 (1984).

Lecidea avium Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández East Is]. 2 (Bot.), 3(11): 357 (1924) T.

Lecidea aeruginosa Nyl., Amn. Sci. Nat. Bot. sér. 4,3: 164 (1955) T.

Lecidea areuginosa Borrer.

Lecidea chilena Zahlbr., Cat. Lich. Univ. 3: 533 (1925).

Lecidea chilensis G. Follmann, Nova Hedwigia 14: 250 (1967) nom. nov. illegit.

caesiorubella Ach., Lich. Univ.: 366 (1810) E

Lecanora espinosae Zahlbr., Ann. Mycol. 29: 83 (193 DT.

carpinea (L.) Vain., Medd. Soc. Fauna Fl. Fenn. 14: 23 (1888).

cenisea Ach., Lichenogr. Universalis: 361 (1810).

chlarona var. fuegiensis Rásánen, Amn. Soc. Bot. Zool.-Bot. Femn. “Vanamo” 2 (1): 27 (1932) T.

coquimbensis Zahlbr., Acta Horti Gothob. 2: 13 (1925) T.

dancoénsis Vain., Résult. Voy. Belgica, Lich.: 20 (1903).

dispersa (Pers.) Sommerf., Suppl. Fl. Lappon.: 96 (1826).

epibryon ssp. broccha (Nyl.) Lumbsch. In Lumbsch et al., Pl. Syst. Evol. 191: 229 (1994).

Lecanora broccha NyL. In J. Cromb., J. Bot. 14: 21 (1876).

epibryon ssp. xanthophora Lumbsch. In Lumbsch et al., Pl. Syst. Evol. 191: 231 (1994).

Lecanora subfusca var. pelidnocarpa Mont. In C. Gay, Hist. Chile, Bot. 8: 150 (1852) T.

Lecanora aeruginosa Nyl., Amn. Sci. Nat. Bot. sér. 4,3: 157 (1855) T.

follmannii Klem.. Rev. Bryol. Lichénol. 31: 256 (1963) T.

Srazieri C.W. Dodge, Nova Hedwigia 12: 339 (1967) [“1966”] T.

garovaglii (Kórb.) Zahlbr., Ann. Naturhist. Hofmus. Wien 15: 208 (1900).

hagenii (Ach.) Ach., Lichenogr. Universalis: 367 ( 1810).

hiulca (Nyl.) Hue, Nouv. Arch. Mus. Hist. Nat. Paris, sér. 3, 3: 59 (1891).

Squamaria hiulca Nyl.. Ann. Sci. Nat. Bot. sér. 4, 3: 153 (1855) T.

Placopsis hiulca (Nyl.) NyL., Bot. J. Linn. Soc. 9: 25 (1865). Non est Placopsis (Lamb 1948).

Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3 (11): 379 ( 1984) T.

leprosa Fée, Essai Crypt. Ecorc.: 118 (1825).

masafuerensis Zahlb. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is. 2 (Bot.), 3 (11): 378 (1924) T.

Jatta, Malpighia 20: 10 (1906) T.

microphthalma Hook. f. 8: Taylor, Lond. J. Bot. 3: 642 (1844) T.

Lecanora dentilabra Tuck., Proc. Amer. Acad. Arts. Sci. 12: 173 (1877) T.

Lecanora praedolosa Nyl., Lich. Fueg. Patag.: 9 (1888) T.

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

Lecanora albellina Múll. Arg., Miss. Sci. Cap Horn, Lich.: 162 (1889) T.

muralis (Schreb.) Rabenh., Deutschl. Krypt.-Fl. 2: 42 (1845).

poe Vain. var. persorediosa Rásinen, Ann. Bot. pon Zool.-Bot. Fenn. “Vanamo” 2 (1): 27 (1932) T.

C.W. Dodge, Nova Hedwigia 12: 340 (1967) [*1966”] T

di Lumbsch. In Lumbsch et al., Bot. Acta 107: 34 (1993).

peculiaris Zahlbr., Acta Horti Gothob. 2: 15 (1925) T

polytropa (Hoffm.) Rabenh., Dtsch. Kryptfl. 2: 37 (184 5).

skottsbergii Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4 (11): 9 (1912).

9).

xantholeuca (Mill. Arg.) Hertel, Beih. Nova Hedwigia 79: 448 (1984).

Lecidea xantholeuca Mill. Arg., Nuovo Giorn. Bot. Ital. 21: 45 (1889).

Lecidea interrupta Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4 (11): 3 (1912).

Lit.: Hooker £ Taylor (1844); Nylander (1855); Zahlbruckner (1933); Klement (1963); Dodge (1967); Hertel

(1971, 1984); Brodo (1984); Jacobsen € Kappen (1988); Aptroot « Van der Knapp (1993); Lumbsch (1994); Ryan

£ Nash (1997); Lumbsch et al. (1997).

110. Lecidea Ach. (1803)

[Lecanorales: Lecideaceae]

atrobunnea (Ramond ex Lam. £ DC.) que Lich. Helv. Spicil. 3: 134 (1828).

auriculata Th.Fr., Lich. Arct.: 213 (1860

"urella Miill.Arg., Miss. Sci. Cap Horn, de 165 (1888) T.

ueclinanda Nyl., Lich. Nov. Zel.: 146 (1888).

Lecanora declinis Nyl., Lich. Fueg. Patag.: 12 (1888) non Tuck.

1865).

laboriosa Mill. Arg. Flora, Jena 57: 187 (1874).

Inpec pe Methodus: 37 (180 3).

eci, Miill. Arg. In Zahlbr., Kongl. Svenska Vetenskapsakad. Handl. 57 (2): 21 (1917).

icida var. pantherina Ach., Kongl. Vetensk. Acad. Nya Handl.: 232 (1808).

lygomma Nyl. In J.Cromb., J. hue? 13: 334 (1875).

lygomma var. crassilabra (Múll. Arg.) Hertel « Rambold, Symb. Bot. Upsal 32 (1): 104 (1997).

Lecidea crassilabra Mill. Arg., Hedwigia 32: 127 (1893).

mannii Tuck., Syn. N. Amer. Lich. 2: 75 (18

Lecidea kalbii Hertel, Beih. Nova Hedwigia 79: 418 (1984

paschalis Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández o Is1. 2 (Bot.), 4 (13): 452 (1926) T.

philippii Zahlbr., Ann. Mycol. 29: 82 (1931) T.

promiscens Nyl., Flora 55: 358 (1872).

santessonii Hertel, Symb. Bot. Upsal. a 107 (1997) T.

noides Kórb., Syst. Lich. Germ.: (1855).

sciatropha Hue, Deux. Expédit. Antarct. cn Lich.: 110 (1915).

tessellata Flórke, Deutsche Lich. No. 64 (1819).

Lecidea andina Rásinen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 20: 25 (1944) nom. inval.

, 2 1

umbonata (Hepp) Mudd, Man. Br. Lich:: 204 (1861) nomen sed non planta.

variegatula Nyl., Flora, Jena 48: 6 (1865).

verruca Poelt, Mitt. Bot. Miinchen 4: 187 (196

Lit.: Cengia Sambo (1926, 1930); a Hertel (1971, 1984, 1987a, 1987b, 1989, 1995, 1997); Ja-

cobsen € Kappen (1988); Rambold (1989); Aptroot $: Van der Knapp (1993).

111. Lecidella Kórb. (1855)

[Lecanorales: Lecan oraceae

135

Gayana Bot. 55(2), 1998.

ridans (Flot.) Kórb., Syst. Lich. Germ.: 242 (1855).

wulfenii (Hepp) Kórb., Parerg. Lich.: 216 (1861).

Lit.: Hertel (1984); Rambold (1989); Knoph (1990); Leuckert £ Knoph (1993, 1997); Knoph £ Leuckert (1994,

1997); Knoph £ Mies (1995); Knoph et al. (1995).

112. Lecidoma G. Schneider € Hertel (1981)

[Lecanorales: ?Pso:

demissum (Rutstróm) Gott. Schneid. £ Hertel, Herzogia 5: 460 (1981).

Lit.: Hertel (1981); Pietschmann (1990).

113. Leifidium Wedin (1993)

[Lecanorales: Sphaerophoraceae]

tenerum (Lauurer) Wedin, Pl. Syst. Evol. 187: 235 (1993).

Sphaerophorus tener Laurer, Linnaea 2: 45 (1827).

Sphaerophorus tener f. compactus Cromb., Bot. J. Linn. Soc. 15: 223 (1876) T.

Sphaerophoron globiferum var. polycladum Miill. Arg., Bot. Jahrb. Syst. 4: 53 (1883) T.

Sphaerophoron polycladum (Miúill. Arg.) Mill. Arg., Flora 71: 17 (1888).

pon ri polycladum var. depauperatum Mill. Arg., Miss. Sci. Cap Horn, Lich.: 145 (1888).

erophorus tener f. depauperatum (Múill. Arg.) Follmann, Revista Univ. (Santiago) 47: 71 (1962).

Eit: Wedin (1993, 1995b).

114. Leioderma Ny]l. ai:

[Lecanorales: Pannariac

pyenophorum Nyl., Lich. Nov. Zel.: 47 (1888).

Coccocarpia gayana var. suvdivisa Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl, 2 (Bot.),

3 (11): 343 (1924) T.

Parmeliella pycnophora var. subdivisa (Zahlbr.) R. Sant., Ark. Bot. 31A (7): 15 (1944).

Lit.: Galloway é: Jórgensen (1987); Jóorgensen $ Galloway (1992a).

115. Lempholemma Kórb. (1855)

[Lichinales: Lichinaceae]

polyanthes (Bernh.) Malme, Lich. Suec. e. No. 883 (1924).

Lit.: Follmann (1965b); Schiman-Czeika (1988).

116. Lepolichen Trevis. Loco

[Pertusariales: Coccotremataceae]

coccophorus (Mont.) Trevis. Spighe e Paglie: 5 (1853).

Parmelia coccophora Mont., Ann. Sci. Nat. Bot. sér. 3, 18: 309 (1852) T.

Lecanora coccophora (Mont.) Nyl., Flora 38: 674 (1855).

Thelocarpon coccophoron (Mont.) Nyl. Mém. Soc. Imp. Sci. Nat. Cherbourg 5: 135 (1858) [“1857”].

Pertusaria coccophorum (Mont.) Nyl. In a Bot. J. Linn. Soc. 15: 233 (1876).

ertusaria thamnoplaca Tuck., Proc. Amer. Acad. Arts Sci. 12: 175 (1877) T.

Lit.: Montagne (1852); David € Hawksworth (1991): Galloway £ Watson-Gandy (1992); Castillo et al. (1995).

117. Lepraria Ach. (1803) nom. cons.

[Ascomycota: inc.

incana (L.) Ach., Meth. Lich.: 4 (1803).

bificans Nyl., Flora, Jena 56: 196 (1873).

Lit.: Laundon (1989, 1992).

118. L perdido (1878)

[Mitosporic fungi

ans (Th! LM ex Hue, Lichenes Exotici a Professore W. Nylander descripti vel recogniti. Nouv. Arch.

Mus. His. Nat. Paris, sér. 3, 2: 248 (1890).

arbuscula (Nyl.) Nyl., Lich. Ins Guineensium: 8 (1889 yA

congestum tdci Lamb é: Ward, J. Hattori Bot. Lab. 38: cio

Stereocaulon subalbicans LM. Lamb ex a Bryologist 60: 220 (1957) T.

uck.) Nyl., Lich. Japoniae: 19 (1890).

Lit.: Lamb € Ward (1974).

136

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

119. Leproloma o ex Cromb. (1894)

[Lecanorales: ? Pann

angardianum (Dvstedal J.R. Laundon, Lichenologist 21: 19 (1989).

polo pdas (Dicks.) Vain., Természetr. Fiiz. 22: 293 (1899).

uauxii (Hue) J.R. Laundon, Lichenologist 21: 13 (1989).

E iaa (1989); Leuckert £« Kiimmerling (1991).

120. Leptogium (Ach.) S.F. Gray (1821)

[Lecanorales: Collemataceae

australe (Hook.f. £ Taylor) Miill. Arg., Flora 70: 268 (1887).

a saturninum var. australe (Hook. f. 8 Taylor) Hook. f., FL. Antarct. 2: 541 (1847).

azureum (Sw.) Mont. In Webb á Berth., Hist. Nat. lles Canar. 3 (2): 129 (1840).

reia Mont. In Webb á Berth., Hist. Nat. Iles Canar. 3 (2): 130 (1840).

britannicum P.M. Jórg. 8: P. James, Lichenologist 15: 110 (1983).

cochleatum (Dicks.) P.M. Jórg. £ P. James, Lichenologist 15: 113 (1983).

ideum (Meyen á Flot.) Vain., Ann. Acad. Sci. Fenn., Ser.A 6 (7): 110 (1915).

restó? (Rabenh.) Kórb., Syst. Lich. Germ.: 420 (185 5).

ns P.M. Jorg., Flechten Follmann: 237 (1995) T

rus ata M.E. Mitch. £ P.M. Jorg., Herzogia 2: 462 (1973).

juressianum Tav., Portugalia Acta Biol. Ser. B, Sist. 3: 68 (1950).

laceroides de Lesd., Ann. Crypt. Exot. is Las es

malmei P.M. Jórg., Herzogia 3: 442 (19

menziesii (Ach.) Mont., Ann. Sci. Nat. cid SEL. 3: o 313 (1852).

Parmelia menziesii Ach., Meth. Lich.: 221 (1803).

Lichen menziesii Sm. In Ach., Meth. Lich.: 212 pe nom. inval.

Collema menziesii (Ach.) Ach., Lich. Univ.: 645 (181

Leptogium granuligerum Hue, Lichens Deux. Exp. Pt Fr.: 16 (1915)

Leptogium menziesii var. subcaesia Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 47 (1932) T.

phyllocarpum (Pers.) Mont., Ann. Sci. Nat. Bot. sér. 3, 10: 134 (1848).

valdivianum M. Lindstróm, Lichenologist 28: 91 (1996) T.

Lit.: ponen 8: Kappen (1988); Galloway « Jórgensen (1995); Lindstróm (1996); Jórgensen (1997).

*Lichenoconium Petr. 8 Syd. (1927)

[Mitosporic fungi]

ecanorae (Jaap) D. Hawksw., Bull. Br. Mus. (Nat. Hist.) Bot. 6: 183 (1979).

Lit.: Hawksworth (1981); Aptroot £ Van der Knapp (1

121. Lichenothelia D. Hawksw. (1981).

[Dothideales: Lichenotheliaceae]

minor Henssen, Biblioth. Lichenol. 25: 261 (1987).

patagonica Henssen, Biblioth. Lichenol. 25: 261 ( 1987).

Lit.: Henssen (1987).

2. Lichina Agardh (1817) nom. cons.

(Lichinale Lichinaceae]

co! mdd (O.F. Mill.) Agardh, Spec. Alg. 1: 105 (1821).

sulans Henssen, Lichenologist 4: 95 (1969).

Lito H aos (1969).

123. Lithothelium Miill. Arg. (1885)

dps Pyrenulac psa

um Mill. Arg., Bot. Jahrb. Syst. 6: 386 (1885).

Lit.: abc (1917); Aptroot (1991a).

124. Lopadium Kórb. dns nom. cons.

[Lecanorales: Ectolechiacea

diffluens (Nyl.) Darb., Wiss. Ergebn. Schwed. Súdpolarexp. 1901-1903, 4 (11): 49 (1912).

Lecidea diffluens Nyl., Lich. Fueg. Patag.: 17 (1888

Heterothecium diffluens (Nyl.) Múll. Arg., Nuovo Giom. Bot. Ital. 21: 48 (1889).

137

Gayana Bot. 55(2), 1998,

inutum C.W. Dodge, Nova Hedwigia 12: 336 (1967) [“1966”] T.

Lit: pcs (1952).

125. Loxospora A. Massal. (1852)

pocotas rre

ES Ach.) A. Massal., Ric. Auton. Lich. Crost.: 138 (1852).

Lit.: Rogers £: Hafellner (1988); Tónsberg (1992); Brodo £ Henssen (1995); Ekman (1996).

126. Maronea A. Massal. (1856)

[Lecanorales: Aeon

W. Dodge, Nova Hedwigia 16: 491 (1969) T.

W. Dodge Nova Hedwigia 12: 336 (1967) [“1966”] T.

EJE: Pal (1934).

127. Massalongia Kórb. (1855)

bici incert. sed.

sa (Dicks.) Kórb., Syst. Lich. Germ.: 109 (1855 5).

Lit: BSAS o (1924); Henssen (1963a); Redon (1985); Jacobsen Kappen (1988).

128. Mazosia A. Massal. cb

poes e Phragmopelthecaceae]

ma (Nyl. Abed Cat. Lich. Univ. 2: 503 (1923).

Lit.: Eee Kalb 8 Vezda (1988); Liicking £ Matzer (1996).

129. Megalaria Hafellner (1984)

[Lecanorales: Megalariaceae]

pe (Pers. ex. Nyl.) Hafellner, Beih. Nova Hedwigia 79: 302 (1984).

putverea (Borrer) Hafellner £ Schreiner, Biblioth. Lichenol. 45: 146 (1992).

Ext: eee (1984).

130. Megaloblastenia a (1983)

[Lecanorales: Megalospo ]

marginiflexa (Hook. E. E Taylor) Sipman, Biblioth. Lichenol. 18: 87 (1983).

Lit: Sipman (1983).

131. Megaspora (Clauzade > Cl. Roux) Hafellner $ V. Wirth (1987)

[Pertusariales: Megasporace

verrucosa (Ach.) Hafellner « V. eps o Baden-Wiirttemberegs 1: (1987).

Lit.: Wirth (1987); Jacobsen 4 Kappen (1

132. Melanaria san 1936)

[Pertusariales: Pe: ae]

atacamae CW. o Mona Hedwigia 12: 337 (1967) [“1966”] T.

Lit.: Dodge (1967); Follmaoa CO hi

133. Melanelia Esslinger (1978)

pro p pa aceae]

br.) Essl., Mycotaxon 7: (1978).

pe D.J. Galloway £ P.M. Jo pre... dpi Pa Bot. 28: 10 (1990).

78).

Parmelia subaurifera y var. sabolals a. Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 19 (1932).

ushuaiensis (Zahlbr.) Essl., Pe dor 7: 49 (1978).

Parmelia ushuaiensis Zahlbr., Kon ngl. Svenska Vetenskapsakad. Handl. 57 (6): 42 (1917).

Parmelia roivainenii Rásánen, Ann. Bot. Soc. Zool. -Bot. Fenn. “Vanamo” 2 (1): 19 (1932) T.

pro (Ess1.) Essl., Mycolaxon 7: 49 ad

zopheroa J. Hattori Bot. Lab. 42: 96 ( 1977) T.

pens A (1917); Sentesaon (1944); Esslinger (1977, 1978); Galloway $ Jórgensen (1990); Elix (1993,

134. Melanothecopsis C.W. Dodge (1967)

[Pyrenulales: incert. sed.]

coactella (Stirt.) C.W. Dodge, Nova Hedwigia 12: 310 (1967) [*1966”].

138

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

follmannii C.W. Dodge, Nova Hedwigia 12: 309 (1967) [*1966”] T.

Lit.: Dodge (1967).

135. Melaspilea Nyl. (1858)

pgppEtias. incert. sed.: Melaspilaceae

atacamensis C.W. Dodge, Nova Hedwigia 12: 318 pl [“1966”] T.

lied C.W. Dodge, Nova Hedwigia 16: 485 (196

coquimbensis C. W. Dodge, Nova Hedwigia Zo e sent 1966”] T.

(1969) T

ocarpa Mill. Arg., Miss. Sci. Cap Horn, Lich.: 168 (1889) T.

¡Es Deben 1969).

136. Menegazzia A. Massal. (1854)

[Lecanorales: niego

albida (Zahlbr.) R. Sant., Ark. Bot. 30A (11): 12 (1942).

Parmelia cincinnata var. albida Zahlbr., Kongl. Svenska Vetenskapsakad. Handl. 57 (6): 38 (1917) T.

Parmelia wilsonii Vain., In V. Rásánen, Amn. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 17 (1932) T.

cincinnata (Ach.) Bitter, Hedwigia 40: 172 (1901).

A cincinnata Ach., Meth. Lich.: 252 (1803)

e lo Ach., Meth. Lich.: 252 (1803) nom. inval.

Lichen bullatus ed In Ach., Lich. Univ.: 495 (1810) nom. inval.

dispora (Nyl.) R. Sant., Ark. Bot. 30A (11): 13 (1942).

Parmelia dispora Nyl., In J. Crombie, Bot. J. Linn. Soc. 15: 228 (1876) T.

fumarpr trarica Calvelo £ Adler, Mycotaxon 59: 369 (1966).

globulifera R. Sant., Ark. Bot. 30A (11): 30 (1942) T

hollermayeri (Rásánen) R. Sant., Ark. Bot. 30A (11): 22 (194

Parmelia hollermayeri Rásánen, Revista pat ot ss 196 (1937) T.

magellanica R. Sant., Ark. Bot. 30A (11) 23 (194

Genaro (Rásánen) R. Sant., Ark. Bot. 304 da 21 (1942).

rmelia megalospora Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2(1): 18 (1932) T.

ur ctaboin | Zahlbr.) P. James, Fl. Australia 54: 313 (1992).

Parmelia nigrescens Stirt., Scott. Naturalist ng 4: 253 a non. Ach.

disc neozelandica Zahlbr., Cat A Univ. 6: 53 (19

negazzia circumsorediata R. San ., Ark. Bot. 304 ( 1 On

Por in Adler 4 Calvelo, es ie 59: 368 (1996).

opuntivides (Miill.Arg.) R. Sant., Ark. Asa 30A (11): 25 (194

Parmelia opuntioides Miill. Arg., Miss. Sci. Cap Horn, pr 158 (1888) T.

sanguinascens (Rásiánen) R. Sant., Ark. Ba 30A (11): 28 (1942).

Parmelia sanguinascens Rásinen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2(1): 18 (1932) T

Parmelia opuntioides var. vulgaris Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2(1): 18 (1932) T.

Parmelia opuntioides var. violascens Rásiinen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2(1): 18 (1932) T.

Parmelia dispora var. alboffi Zahlbr., Kongl. "Svenska Vetenskapsakad. Handl. 57 (6): 39 (1917) T.

Parmelia alboffi (Zahlbr.) DR. In R. Sant., Ark. Bot. 30A (11): 28 (1942).

tenuis R. Sant., Ark. Bot. 30A (11): 27 (1942) T

Pese (Rásánen) R. Sant., Ark. Bot. 304 ( 11): 26 (1942).

rmelia valdiviensis Rásánen, Revista Univ. (Santiago) 22: 197 (1937) sE

Ext: inician (19420); Quilhot et al. (1975); James £: Galloway (1992); Calvelo £ Adler (1994); Adler £: Calve-

lo (1996)

137. Metus D.J. Galloway £: P. James (1987)

[Lecanorales: Cladoniaceae

efflorescens D.J. Galloway $: P. James, Notes RBG Edinburgh 44 (3): 569 (1987) T.

pileatum (Mont.) D.J. Galloway $ P. James, Notes Roy. Bot. Gard. Edinburgh 44 (3): 571 (1987).

Cladonia pileata Mont., Ann. Sci. Nat. Bot. sér. 3, 18: 3210 (1852) T

Pilophoron pileatum (Mont.) Zahlbr., Cat. Lich. Univ. 10: 381 (1939).

Lit.: Galloway $: James (1987); Stenroos (1996).

138. Micarea Fr. (1825)

[Lecanorales: Micareaceae]

Coppins é Kantvilas, Lichenologist 22: 277 (1

990).

erratica (Kórb.) Hertel, Rambold $: Pietschm.. In G. Rambold, Biblioth. Lichenol. 34: 227 (1989).

139

Gayana Bot. 55(2), 1998.

isabellina Coppins $ Kantvilas, Lichenologist 22: 284 (1990).

prasina Fr., Syst. Orb. e es (1825).

turfosa (A. Massal.) Du Rietz., Svensk Bot. Tidskr. 17: 94 (1923 ).

Lit.: Coppins (1983); Coppins z ¡la (1990); Aptroot $: Van der Knapp (1993).

ras (1927)

[Caliciales: Microcaliciaceae]

onversum Tibell, Bot Not. 131: 237 (1978).

Lit.: Tibell (1987, 1994).

Microglaena Kórb. (1855) = Thelenella Nyl.

140. Miltidea Stirt. (1877)

[Lecanoraceae: Miltideaceae]

ceroplasta (C. Bab.) D.J. Galloway 8: Hafellner, Beih. Nova Hedwigia 79: 308 (1984).

Biatora ceroplasta C. Bab. In J.D. Hook., Fl. Nov. Zel. 2: 300 (1855).

Lit.: Hafellner (1984); Galloway (1985).

141. Minksia Miill. eric Poem

[Arthoniales: Roccellace:

chilena (C.W. Dodge) Redon 4 Follmann, Philippia 1: 133 (1972

Medusulina chilena Dodge, Nova Hedwigia 12: 322 e [“1966”] E

saxicola J. H ancock Pacific Exped. 3 (9): 184

Lit.: Miller Argoviensis (1882); Follmann (1968a); Redon es dido (1972a); Mies « Printzen (1997).

142. Multiclavula R.H. Petersen (1967)

geo: Clavariaceae

mucida -H. Petersen, Amer. Midl. Naturalist 77: 212 (1967).

Lit.: Petersen e Kantvilas (1986): Poelt £« Obermayer (1990).

143. Mycoblastus Norman (1853)

[Lecanorales: Mycoblastaceae]

dissimulans (Nyl.) Zahlbr., Cat. Lich. Univ. 4: 3 (1926).

Lecidea dissimulans Nyl., Amn. Sci. Nat. Bot. sér. 4, 3: 167 (1855) T.

Lit.: Zahlbruckner (1926).

144. Mycob bia Rehm (1890)

palco puta

Note: It is possible that the taxon Lecidea dolichospora Ny]., Pra Sci. Nat. Bot. sér. 4, 3: 162 (1855) T] should be

transferred to Mycobilimbia. lts realted synonyms are listed below

Bacidia dolichospora (NyL.) Trevis., Linnaea 28: 293 (1856).

O dolichospora (Nyl.) Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.),

(11): 394 (1924).

Lit.: Nylander (1855); Zahlbruckner (1924); Sipman (1983); Hafellner (1984); Timdal (1991).

145. Mycomicrothelia Keiss]. (1936)

[Dothideales: Arthopyreniaceae]

para (Ach.) D. Hawksw., Bull. Br. Mus. Nat. Hist. (Bot.) a Pa A

rrucaria cinerella NyL., Amn. Sci. Nat. Bot. sér. 4,3: 174 (1855) T

Lit.: dt 1985).

146. Myxodictyon A. na (1860) = Brigantiaea

diem Brigantiaceae]

chil + Nova Hedwigia 16: 492 (1969) [*1968”] T.

Ext: Dodge ( (1969): Hafeliner A: Bellemére (1981).

147. Neofuscelia Essl. (1978)

[Lecanorales: Parmeliaceae]

delisei (Duby) Essl. , Mycotaxon 7: 50 (1978).

Parmelia elivocea var. delisei Duby, a Gall. 2: 602 (1830).

glabrans (Nyl.) Essl.. Mycotaxon 7:50 (197.

Parmelia glabrans Nyl.. Flora 58: st

140

Checklist of Chilean lichen: GALLOWAY, D.J. € W. QUILHOT

imitatrix (Taylor) Essl., Mycotaxon 7: 50 (1978).

Parmelia imitatrix Taylor, Lond. J. Bot. 6: 161 (1847).

squamans (Stizenb.) Essl., Mycotaxon 7: 52 (1978).

Parmelia squamans Stizenb., Ber. Thátigk. St. Gallischen ¿raro Ges. 1887-88: 164 (1889).

Parmelia degasperii (Sambo) C.W. Dodge, Trans. Amer. Microscop. Soc. 84: 507 (1965).

Parmelia acetabulum var. degasperii Sambo, Boll. Soc. Bot. la. 1926: 90 (1926).

stygiodes (Nyl. ex Cromb.) Essl., Mycotaxon 7: 52 (1978).

Parmelia stygiodes Nyl. ex Cromb., J. Bot. 13: 333 (1875).

ea (Essl.) Essl., Mycotaxon 7: 52: (1978).

rmelia subhosseana Essl., J. Hattori Bot. Lab. 42: 148 (1977).

Mica ie Es ) Essi. , Mycotaxon 7: 53 (197

a a tatimirix Essl., J. Hattori Bot. Lab. 42: 151 (1977) T.

Lit.: Esslinger (1977, 1978); Elix (1993,1994g).

148. Nephroma Ach. (1810)

io cirr A OA ae]

m Ny]l., In J. Crombie, Bot. J. Linn. Soc. 15: 231 (1876) T.

Opisteria crol (Nyl.) Vain., Ark. Bot. 8 (4): 93 (1909

Nephroma patagonicum Krempelh., Verhandl. Zool.-Bot. Ges. Wien 26: 439 (1876) T.

Opisteria patagonica var. macrospora Rásinen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 33

1932) T.

Nephroma analogicum var. macrospora (Rásiinen) Follmann, Revista Univ. (Santiago) 50-51: 62 (1966).

Opisteria patagonica var. squamosa Rásánen, Amn. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 34 (1932)

cc patagonicum var. squamosum pr rgirió paid Cat. Lich. Univ. 10: 284 (1939).

ephroma sublaevigatum f. ryptog. Exot. 4: 136 (1931) T.

ido (Jacq.) Nyl., Syn. Meth. Lich. 10): 317 ¡cora

Lichen antarcticus Jacq., Miscell. Austriac. 2: 370 (1781) T.

antarcticum var. lobuligerum Mill. Arg., Miss. Sci. Cap Horn, Lich.: 154 (1888) T.

24 (1

Nephroma tema (Miill. Arg.) Gyeln., Magy Bot. Lap. 29: 930).

igor d odio var. lobuligera (Mill. Arg.) Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2

att 3 (1932).

Nephroma homanii Gyeln., Ann. Cryptog. Exot. 4: 129 (1931) T

Opisteria homanil (Gyeln.) Rásánen, Revista Univ. (Santiago) 2 (1): 202 (1937).

cellulosum (Ach.) Ach., Lich. Univ.: 52 23 (1810

Peltidea cellulosa Ach., Meth. Lich.: 289 (1803

Opisteria cellulosa (Ach.) Vain., Ark. Bot. 8 (43): 93 (1909).

cellulosum var. isidioferum J. S. Murray, Trans. R. Soc. N.Z. 88: 385 (1960).

Nephroma is lepidonicilad Rásiinen ex Gyeln., Ann. Cryptog. Exot. 4: 147 (1931) T.

Nephromium lepidophyllum (Rásánen ex Gyeln.) Rásiinen, Ann. Soc. Bot. Zool.-Bot. Fenn. “Vanamo” 2

1): 34

cin li f. hypomelaena Rásinen ex L. M. Lamb, Farlowia 4: 439 (1955).

chubutense 1.M. Lamb, Farlowia 4: 438 (1955).

kuehnemannii 1.M. Lamb, Farlowia 4: 437 y

microphyllum Henssen, Mycotaxon 7: 358 (1978).

papillosum F.J. White 8 P. James, Lichenologist 20: 146 (1988) dE.

parile (Ach.) Ach., Lich. Univ.: 522 (1810).

Lichen parilis Ach., Lich. Suec. Prodr.: 799).

plumbeum (Mont.) Mont, in Gay, Hist. Chile, Bot. 8: 100 (1852).

Peltigera plumbea Mont., Ann. Sci. Nat. Bot. sér. 2, 4: 87 (1835) T.

Nephromium plumbeum (Mont.) Nyl., Mém. Soc. Sci. Nat. Cherbourg 5: 101 (1857).

Nephroma lya -D.

Nephromium byallii (C. Bab.) Nyl., Syn. Meth. Lich. 1 bl 322 (1860).

Opisteria lyallii (C. Bab.) Vain., Ark. Bot. 8 (4): 93 (1909

Nephroma chilensis Gyeln., Ann. Cryptog. Exot. 4: 132 jet E

?Nephroma pubescens var. minutiphyllinum Gyien., Ann. Cryptog. Exot. 4: 136 (1931).

ar. isidiatum (3.S. Murray) FJ. White éz P. James, Lichenologist 20: 154 (1988).

141

Gayana Bot. 55(2), 1998.

Nephroma lyallii f. isidiatum J.S. Murray, Trans. Roy. Soc. New Zealand 88: 387 (1960).

pseudoparile (Rásánen) Zahlbr., Cat. Lich. Univ. 10: 285 (1939).

Nephromium pseudoparile Rásánen, Revista Univ.(Santiago) 21: 143 (1936) T.

Opisteria homanii var. gyelnikii Rásánen, Revista Univ. (Santiago) 22: 202 (1937) T.

Nephroma gyelnikii (Rásánen) IM. Lamb, Anales Parques Nac. 7: 81 (1959) [*1958”].

skottsbergii F.J ite 8 P. James, Lichenologist 20: 162 (1988) T.

Lit.: White £ James (1988).

149. Nimisia Kirnefelt $ Thell (1993)

[Lecanorales: Parmeliaceae]

egiae Kárnefelt £ Thell, Lichenologist 25: 370 (1993).

Lit.: Kárnefelt € Thell (1993a).

50. Neuropogon Nees £ Flotow (1835)

[Lecanorales: Parmeliaceae

acromelanus (Stirt.) LM. Lamb, Bot. J. Linn. Soc. 52: 218 (1939).

Usnea fasciata Torrey, Am. J. Sci. 6: 106 (1823) T.

Neuropogon antennarius Nees $ Flot., Linnaea 9: 497 (1835) T.

Usnea taylori var. kranckii Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 10 (1932) T.

Usnea trachycarpa var. eciliata Risánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 10 (1932) T.

durietzii (Mot.) D.J. Galloway £ Quilhot, comb. nov.

Basionym: Usnea durietzii Mot., Lich. Gen. Usnea Stud. Monogr. 2: 503 (1937) T.

patagonicus (F.J. Walker) D.J. Galloway $: Quilhot, comb. nov.

Basionym: Usnea patagonica F.J. Walker, Bull. Brit. Mus. (Nat. Hist.) Bot. 13: 82 (1985).

perpusillis LM. Lamb, Bot. J. Linn. Soc. 52: 234 (1939) T.

icus (F.J. Walker) D.J. Galloway $: Quilhot, comb. nov.

Basionym: Usnea subantarctica EJ. Walker, Bull. Brit. Mus. (Nat. Hist.) Bot. 13: 99 (1985).

us Stirt., Scott. Nat. (Perth) 6: 105 (1881).

Usnea taylori var. subciliata Rásinen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 11 (1932) T.

Lit.: Lamb (1964); Walker ( 1985); Jacobsen £ Kappen ( 1988); Quilhot et al. (1991a, 1996)

151. Normandina Nyl. (1855)

[Ascomycota: incert. sed.]

pulchella (Borrer) Nyl., Ann. Sci. Nat. Bot. sér. 4, 15: 382 (1861).

Lit.: Aptroot (1991b).

[Ostropales: Thelotrem

Note: No Chilean material seen was referable to this genus, but the taxon Leptotrema schizoloma Mill. Arg.[ Nuo-

vo Giorn. Bot. Ital. 21: 49 ( 1889)] may be referable to Ocellularia.

Lit.: Miiller Argoviensis (1889); Follmann (1964); Hale (1981).

153. Ochrolechia A. Massal. (1852)

[Pertusariales: Pertusariaceae]

antarctica (Múll. Arg.) Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4, 11: 50 (1912).

ertrusaria antarctica Mill. Arg., Flora 69: 125 (1886).

Lecanora antarctica (Miill. Arg.) Miill. Arg., Rey. Mycol. 9: 135 (1887).

Lecanora hypotartarea Ny]l., Lich. Fueg. Patagon.: 9 (1888).

ue hypertartarea (Nyl.) Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4 (11): 50

152. Ocellularia G.Meyer (1825) nom. cons.

ataceae]

, Ann. Soc. Bot. Zool.-Bot. Fenn. “Vanamo” 2 (1): 28 (1932) T.

iformis (Vain.) Verseghy, Beih. Nova H igia 1: 54 (1962).

Ochrolechia pallescens var. occelliformis Vain., Résult. Voy. Belgica, Lich.: 21 (1903) T.

¡(L.) A. Massal., Nuovi Amn. Sci. Nat. Bologna 7: 212 (1853).

Ochrolechia blandior (NyL) Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4, 11: 50 (1912).

142

Checklist of Chilean lichen: GALLOWAY, D.J. $ VW. QUILHOT

Lecanora parella * blandior Nyl., Lich. Fueg. Patagon.: 8 (1888).

parella (L.) A. Massal., Ric. Auton. Lich. Crost.: 30 (1852).

diremos (Vain.) Verseghy, Beih. Nova Hedwigia 1: 77 (196

Ochrolechia pallescens var. pseudotartarea Vain., Résult. Voy. EN Lich.: 21 (1903) T.

Lit.: Verseghy (1962); Huneck £ Follmann (1971); Brodo (1991).

154. Omphalodiella Henssen (1991)

Epscaas Parmeliaceae]

patagonica Henssen, Lichenologist 23: 335 (1991).

Lit: Sapa (1991).

155. Omphalodium Meyen « Flot. (1843)

[Lecanorales: Parmeliaceae]

pa mn... Rásinen, Anales Soc. Ci. hn 131: 98 (1941).

nse Meyen á Flot., Nov. Act Acad. Caes. Leop.-Carol. Nat. Curr. Suppl. 19: 223 (1843).

Lit.: A 1973); Ho (1992a); Nash et al. (1990); Elix (1993).

6. Opegrapha Ach. (1810) nom. cons.

[Arthoniales: Roccellaceae]

atra Pers., Usteri's N. Ann. Bot. 1: 30 (1794).

atacamensis C.W. Dodge, Nova o 12: 313 (1967) [“1966”] T.

atratula. Miill. Arg., Flora 63: 41 (1880

betulina Sm. In Sm. £ Sowerby, Bot. ¡ea 0 32: tab. 2281 (1811).

bonplandii Fée, Essai Crypt. Ecorc. 1: pe 25).

cactacearum Riedl, Sydowia 17: 85 (1964

decussata C.W. Dodge, Nova Hedwigia 12: en (1967) [“1966”] T.

dirinoides Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 168 (1855) T.

eulychniae C.W. Dodge, Nova Hedwigia 12: 314 (1967) [* ta

euphorbiae C.W. Dodge, Nova Hedwigia 12: 315 (1967) [“1966” ]

*leuckertii S. Kondratyuk $ D.J. Galloway, e qlo + 332 (1995) T.

medusuliza Nyl., Lich. Fueg. Patag.: 17 (188

oxalidis C.W. Dodge, Nova Hedwigia 12: sos ae [“1966”] T.

paschalis Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter 1sl. 2 (Bot.), 4 (13): 450 (1926) T.

robustula Nyl., Lich. Nov. Zel.: 147 (1888) T

tigrensis C.W. Dodge, Nova Hedwigia 12: 317 (1967) [* 1966”] T.

larensis C. W. Dodge, Nova Hedwigia 12: 317 (1967) [“1966”] T.

Lit.: Zahlbruckner (1926); Follmann (1962); Dodge (1967); Riedl € Schiman-Czeika (1964); Tehler (1983); Kon-

dratyuk £ Galloway (1995).

E o

[Lecanorales: Ophioparm

pales (Follmann) Kalb á rada Biblioth. Lichenol. 58: pq

Haematomma araucariae Follmann, Bol. Univ. Chile 7: 44 (1965) T

Lit.: Follmamn (1965d); Rogers 4 Hafelloer (1988) Kalb et al. (1995).

158. Oropogon Th.Fr. (1861)

[Lecanorales: Alectoriaceae]

rolobic Essl., Syst. Bot. Monogr. 28: 81 (1989).

Lit.: Esslinger (1989).

159, Lónnr. (1858)

(Graletles: Gyalectaceae

? carneola (Ach.) Am., Flora 54: 50 (1871).

Ls Nimis (1993).

60. Pannaria Delise ao

ccanonls Emugi=ss

hilaris Zahlbr. In C. Skottsberg, Nat. Hi

hookeri (Borrer ex Sm.) Nyl., Mém. Soc. Nat. Sci. Nat. e ER 109 (18

143

Gayana Bot. 55(2), 1998.

immixta Nyl., Bot. J. Linn. Soc. 9: 249 (1866).

pac rda 0 Ny]l., Ann. Sci. Nat. Bot. sér 4, 3: 150 (1855) T.

rubiginosa var. vulcanica Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3(11): 338

os 4) T

mienta Ny]1. In J. Crombie, Bot. J. Linn. Soc. 15: 231 (1876) T

Lit: Nylander (1855); Zahlbruckner (1917, 1924); Jóorgensen (1994); Jorgensen £ Galloway (1992a).

Pannoparmelia (Mill. Arg.) Darb. (1912)

Peret Parmeliaceae]

angustata (Pers.) Zahlbr. In H. Handel-Mazzetti, Symb. Sin. 3: 195 (1930).

Parmelia angustata Pers. In C. Gaudichaud-Beaupré, Voy. Uranie Bot.: 195 (1827).

as angustata (Pers.) Miill. Arg., Flora 72: 507 (1889).

noparmelia anzioides Darb., Wiss. Ergebn. Schwed. Siidpolarexp 1901-1903, 4 (11): 11 (1912).

Anzia anzioides (Darb.) IM. Lamb, Anales Parques. Nac. a 141 (1959) [*1958”].

zia darbishireana Zahlbr., Cat. Lich. Univ. 6: 276 (192

Lit.: Calvelo £ Adler (1992); Elix (1993); Galloway € Sammy ed:

162. Parmelia Ach. (1803) nom. cons.

[Lecanorales: Parmeliaceae

cunninghamii Cromb., Bot. J. Linn. Soc. 15: 228 (1876) T.

protosulcata Hale, Mycotaxon 16: 162 (1982).

saxatilis (L.) Ach., Meth. Lich. 204 (1803

sulcata Taylor In J.T. Mackay, Fl. Hibern. 2: 145 (1836).

on cruenta Darb., Wiss. Ergebn. Schwed, Siidpolarexp. 1901-1903, 4 (11): 13 (1912) T.

rmelia sulcata f. aberrans Zahlbr., Ann. Mycol. 29: 84 (1931) T

Lit: a (1987); Stenroos (1991); Elix (1993, 19941).

163. Parmeliella Múll. Arg. (1862)

[Lecanorales: Pannariaceae]

concinna 1M. Lamb, Farlowia 4: 431 (1955).

granulata 1.M. Lamb, Farlowia 4: 429 (1955).

nigrocincta (Mont.) Múll. Arg., Flora 44: 86 (1881).

Parmelia nigrocincta Mont., Ann. Sci. Nat. Bot. sér. 2, 4: 91 (1835) T.

HE nes (Stirt.) Zahlbr., ro a Univ. 3: 225 (1925).

rmeliella major Darb., s. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4 (11): 17 (1912).

Est: bl 1955, 1959); => % Galloweyí 1992a); Jórgensen (1998).

164. Parmelina Hale (1974)

[Lecanorales: Parmeliaceae

Zahlbr.) Elix £ J. Johnst., Brunonia 9: 160 (1987) [“1986”].

Lit: Elix « Johnston (1987b); Elix £ Hale (1987); Elix (1993, 1994;).

165. Parmeliopsis Nyl. (1866)

[Lecanorales: Parmeliaceae]

h pta (Ach.) Vain., Meddeland. Soc. Fauna Fl. Fenn. 6: 127 (1881).

Lit.: Elix (1993, 1994k).

66. Parmentaria Fée (1824)

[Pyrenulales: aia

ai Crypt. Ecorc.: 70 (1825).

Lit.: Follmann (1961) Galloway (1985).

167. Parmotrema A. Massal. (1860)

[Lecanorales: Parmeliacea

arnoldii (Du Rietz) Hale, Phytologia 28: 335 (1974).

chinense (Osbeck) Hale £ Ahti, Taxon 35: 133 (1986).

crinitum (Ach.) M. Choisy, Bull. Mens. Soc. Linn. Soc. Bot. Lyon 21: 175 (1952).

cristiferum (Taylor) Hale, Phytologia 28: 335 (1974).

(Vain.) Hale, Phytologia 28: 335 (1974).

praesorediosum (Nyl.) Hale. Phytologia 28: 338 (1974).

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

rampoddense (Nyl.) Hale, Phytologia 28: 338 (1974).

sancti-angelii (Lynge) Hale, Phytologia 28: 339 (1974).

subarnoldii (Abbayes) Hale, Phytologia 28: 339 (1974).

sulphuratum (Nees á Flot.) Hale, Phytologia 28: 339 (1974).

tinctorum (Despr. ex Nyl.) Hale, Phytologia 28: 339 (1974).

zollingeri (Hepp) Hale, Phytologia 339 (1974).

Lit.: Hale (1965); Krog € Swinscow (1981); Swinscow $: Krog (1988); Elix (1993, 1994m).

168. Peltigera Willd. (1787) nom. cons.

[Peltigerales: Peltigeraceae]

andensis Vitik., Flechten Follmamn: 372 (1995).

austroamericana Zahlbr., Cat. Lich. Univ. 3: 456 (1925).

canina (L.) Willd., Fl. Berol. Prodr.: 347 (1787)

collina (Ach.) Schrad., J. Bot 1801: 78 (180 a...

With.) J.R. Laundon, Lichenologist 16: 217 (1984).

dolichorhiza Nyl., Lich. Nov. Zel.: 43 (1888).

frigida R. Sant., Ark. Bot. 31A (7): 11 (1944) T.

laciniata (G. Merr. ex Riddle) Gyeln., Nyt. Mag. Naturvidensk. 68: 270 (1930).

lepidophora (Vain.) Bitter, Ber. Deutsch. Bot. Ges. 22: 251 (1

neckeri Hepp ex Miill. Arg., Mém. Soc. Phys. Hist. Nat. Genéve 16: 370 (1862).

? patagonica Rásinen, Anales Soc. Ci. Argent. 128: 142 (1939).

polydactylon (Neck.) Hoffm., Descr. Adumbr. Lich. 1: 19 (1790).

pulverulenta Taylor, Lond. J. Bot. 6: 184 (1847

rufescens (Weiss) Humb., Fl. Friberg: 2 (1793).

scabrosa Th. Fr., Lich. Arct.: 45 (1860).

?spuriella Vain., étud. Lich. Brésil 1: 180 (1890).

?truculenta De Not., Mem. R. Accad. Soc. Torino, ser. 2, 12: 134 (1851).

Lit.: Santesson (1944); Thomson (1955); Lindahl (1962), Holtan-Hartwig (1993); Goffinet £ Hastings (1994); Vi-

tikainen (1992, 1994, 1995)

169. Peltula Ny]. (1853)

[Lecanorales: Lecanoraceae]

euploca (Ach.) Poelt ex Ozenda $: Clauzade, Les Lichens: 324 (1970).

Note: It is possible that the taxon Heppia chilensis Jatta renga 20: 8 (1906) T] is referable to Peltula, but not

having seen Jatta?s material we hesitate to name it in Pel

Lit.: Biidel (1987); Filson (1988).

170. Peltularia R.Sant. (1944)

[Lecanorales: Coccocarpiaceae]

A (Rásinen) R. Sant., Ark. Bot. 31A (7): 4 (1944).

cocarpia gyrophoroides Rásánen, Anales Soc. Ci. Arg. E. IIL T. 78: 143 (1939).

Coc

Lit.: Santesson (1944); Jóorgensen « Galloway (1984).

171. Pentagenella Darb. (1897)

[Arthoniales: Roccellaceae]

ima Darb., Ber. Deutsch. Bot. Ges. 15: 5 (1897).

Lit.: Darbishire (1897, 1898a, 1898b); Follmann « Schultz (1993); Follmann (1997).

172. Pertusaria DC. (1805) nom. cons.

[Pertusariales: Pertusariaceae]

arthoniaria Ny]l., Lich. Fueg. Patag.: 10 (1888).

chilena Zahlbr., Ann. pida 29: 83 (1931) T

ova Hedwigia 16: 491 (1969) [*1968”] T.

¡ Nat. Cherbourg 5: 117 (1858).

Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11): 376 (1924) T.

jamesii Kantvilas, Lichenologist 22: 296 (1990).

Nyl. le DEI _ Bot. sér. 4, 3: 160 (1855) T.

leioplaca var. turgida Miill. Arg. (1884).

mammata Nyl., Lich. Fueg. Patag.: 10 (1888).

145

Gayana Bot. 55(2), 1998.

melanospora Ny]l., Ann. Sci. Nat. Bot. sér 4, 3: 159 (1855) T.

papillulata Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 159 ( e E

E Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 160 (1855) T

carpa Krempelh. var. monospora Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3

an Eb AS

rugifera Miill. Arg., Miss. Sci. Cap Horn, ce 163 (1888) T.

velata re Nyl., Lich. Scand.: 179 (186

Pertusaria skottsbergii Zahlbr. In C. ¡ente Nat. Hist. Juan Fernández Easter Is. 2 (Bot.), 3 (11): 377

24) T.

Lit.: Nylander (1855); Zahlbruckner (1924, 1933); Kantvilas (1990b); Archer (1993, 1997); Archer 4 Elix (1993);

Archer £ Messuti (1997).

*Phacopsis Tul. (1852)

eras Lecanoraceae]

ra (Tul.) Triebel ££ Rambold, Nova Hedwigia 47: 300 (1988).

Elt: *Hertel (1971); Triebel et al. (1995).

173. Phaeographina Miill. Arg. (1882)

ao Graphidaceae]

ahlbr., pea Mycol. 29: 75 (1931).

ride (Ach.) Miill. Arg., Flora 65: 399 (1882).

Et: oia (1950

. Phaeographis Miill. Arg. (1882)

Ostopals o ae]

a Zahlbr., e Mycol. 5: 129 (1908).

patagonica var. holoplaca Zahlbr., Revista Chilena Hist. Nat. 37: 166 (1933) nom. nud.

Lit.: Zahlbruckner (1933); Follmann (1962).

175. Phaeophyscia e (1977)

[Lecanorales: Physcia ae]

hirsuta (Mereschk.) Essl., Mycotaxon is 302 2d 978).

rn (Ach.) Essl., Mycotaxon 7: 305 (1978).

sctastra (Ach.) Moberg, Symb. Bot. Upsal. 22: 47 (1977).

Et: Pr ore 1993, 1994, 1995); Scutari (1992).

*Phaeosporobolus D. Hawksw. $: Hafellner (1986)

[Mitosporic fungi]

alpin - Sant., Alstrup 8 D. Hawksw., Medd. om Grónl. ., Biosience 31: 15 (1990).

Lit.: Alstrup € Hawksworth (1990); Wedin (1994).

176. Phlyctis Wallr. (1850) nom. cons.

[Ascomycota, incert.sed.: Phlyctidaceae]

chilensis D.J. Galloway $: Guzmán, Lichenologist 20 (4): 393 (1988) T.

Lit.: Galloway £ Guzmán Grimaldi (1988).

Phyllopsora Miill. ho (1894)

erro Bacidiaceae]

bibula (Taylor) a á Krog, Lichenologist 13: 239 o 1).

Lecanora bibula Taylor, Lond. J. Bot. 6: 160 (184

onfusa Swinscow $: Krog, Lichenologist 13: 229 ( ete

Lit: « Krog (1981, 1988); Brako (1989).

178. Phyllisciella Henssen (1984)

[Lichinales: Lichinaceae]

morpha Henssen, Beih. Nova Hedwigia 79: 387 (1984).

Lit.: Henssen 8 Biidel (1984).

179. Physcia papada Michaux (1803)

[Lecanorales: Ph:

adscendens (Fr.) H. Olivier, Flore Lich. L'Orme 1: 79 (1882).

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

alba (Fée) Miill. Arg., Rev. Mycol. 9: 136 (1887).

albata (F. Wils.) Hale, Bryologist 66 : 72 (1963).

biziana (A. Massal.) Zahlbr., Ósterr. Bot. Zeitschr. 51: 348 8 (1901).

caesia (Hoffm.) Fiirnr., Naturhist. e Regensburg 2: 250 (1839).

callosa Ny]., Flora, Jena 52: 119 (1869

convexa Miill. Arg., Rev. Mycol 10: 57 ( 1888).

dimidiata (Arn.) Nyl., var. ornata (Nádv.) Moberg, Symb. Bot. Upsal. 22: 74 (1977).

dubia (Hoffm.) Lettau, Hedwigia 52: 254 (1912 12).

lopezii Moberg, Nord. J. Bot. 10: 333 (1990).

poncinsii Hue, Bull. Soc. Bot. Fr. 63, Mém. 28: 10 (1917).

stellaris (L.) Nyl., Acta Soc. Linn. pes 21: 307 (1856).

tribacia (Ach.) Nyl., Flora 57: 307 (187

undulata Moberg, Nord. J. Bot. 6: 861 e E

aff. vainioi Rásáinen, Medded. Soc. Fauna Fl. Fenn. 47: 166 (1921)

Lit.: Moberg (1977, 1986, 1990, 1994, 1996); Swinscow k: Krog ( 1988); Scutari (1992).

180. Physconia Poelt (1965)

[Lecanorales: Physciaceae]

muscigena (Ach.) Poelt, Nova Hedwigia 9: 30 (1965).

Lit.: Moberg (1977); Swinscow é Krog (1988): Jacobsen £ Kappen (1988).

181. Physma A. Massal. (1854)

[Lecanorales: Collemataceae]

chilense Hue, Bull. Soc. Linn. Normandie sér. 9, 5: 126 (1906) T.

Physma tricolor Zahlbr., Ann. Mycol. 6: 132 (1908) T.

Lit.: Hue (1906); Dughi (1946); Verdon (1992); Verdon « Elix (1994).

182. Placidiopsis Beltr. (1858)

ie soe Verrucariaceae

eusiana Gyeln., Ann. Mus. Nat. Hungar. 32, Pars Bot.: 62 (1939).

EXE: o (19964).

183. Placidium A. Massal. (1855)

[Verrucariales: Verrucariaceae]

analogicum (Breuss) Breuss, Ann. Naturhist. Mus. Wien 98B ( pun 38 (1996).

Catapyrenium analogicum, Breuss, Pl. Syst. Evol. 185: 20 (1993).

andicola (Breuss) Breuss, Ann. Naturhist. Mus. Wien 98B (Suppl.): 38 (1996).

atapyrenium andicolum Breuss, Pl. Syst. Evol. 185: 20 (1993

chilense (Rásánen) Breuss, Ann. Naturhist. Mus. Wien 98B (Suppl.): 38 (1996).

Dermatocarpon chilense Rásinen, Revista Sudamer. Bot. 5: 71 (1938) T.

Catapyrenium chilense (Rásánen) Breuss, Pl. Syst. Evol. 198: 22 (1993).

lachneoides (Breuss) Breuss, Ann. Naturhist. Mus. Wien 98B (Suppl.): 39 (1996).

Catapyrenium lachneoides Breuss, Pl. Syst. Evol. 185: 26 (1993).

ruiz-leali¡ (Rásánen) Breuss, Ann. Naturhist. Mus. Wien 98B (Suppl.): 39 (1996).

Dermatocarpon ruiz-lealii Rásánen, Anales Soc. Ci. Argent. 128: 147 (1939).

squamulosum (Ach.) Breuss, Ann. Naturhist. Mus. Wien 98B (Suppl.): 39 (1996).

Catapyrenium squamulosum (Ach.) Breuss, Ber. Deutsch. Bot. Ges. 98: 389 (1985).

Lit.: Breuss (1993, 1995, 1996b).

ce Placomaronea Risinen (1944)

PR Candelariaceae]

candelarioides Ráisinen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 20 (3): 29 (1944).

candelarivides var. lacinulata R. Sant., Ark. Bot. 31A (7): 9 (1944).

Lit.: Rásánen (1944); Santesson (1944); Hakulinen (1954); Osorio (1974).

185. Placoparmelia od 1992)

[Lecanorales: Parmeliac

Patagonica Henssen, iricio dia 24: 134 (1992).

Lit: Henssen (1992 992b).

147

Gayana Bot. 55(2), 1998.

186. Placopsis (Nyl.) Linds. (1866)

[Lecanorales: Trapeliaceae

baculigera 1.M. Lamb, Lilloa 13: 220 (1947) T.

bicolor (Tuck.) de Lesd., Ann. Cryptog. Exot. 4: 100 (1931).

brevilobata (Zahlbr.) IM. Lamb, Lilloa A 276 (1947).

chilena 1M. Lamb, Lilloa 13: 213 (1947

contortuplicata 1.M. Lamb, Lilloa 13: std (194

contortuplicata f. fuegiensis 1M. Lamb, Lilloa >: a (1947) T.

cribellans (Nyl.) Rásánen, J. Jap. Bot. 16: 1d 940).

dusenii 1[M. Lamb, Lilloa 13: 215 (1947) T

effusa 1.M. Lamb, Lilloa 13: 216 (1947) T.

fuscidula 1.M. Lamb ex Rásánen, Anales Soc. Ci. Argent. 128: 138 (1939).

Lecanora gelida f. leprosula Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11):

382 (1924) T.

gelida (L.) Linds., Trans. Linn. Soc. 25: 536 (1866).

gelida var. subreagens 1.M. Lamb, Lilloa 13: 206 (1947) T

Lecanora patagonica f. sorediosula Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.),

3 (11): 382 (1924) T

Lecanora sorediosula (Zahlbr. ) Gyeln., Acta Fauna Fl. Univ. 2 (1): 10 (1933).

parellina LM. Lamb, Res. Norweg. Sci. Exp. Tristan da Cunha 58 38, 1 (3): 3 (1940) T.

ina var. carnea (Rásánen) I.M. Lamb, Lilloa 13: 253 (194

ca gelida var. carnea Rásánen, Ann. Bot. Soc. Zool. pue Fenn. “Vanamo” 2(1): 25 (1932) T.

carnea f. subcribellans 1.M. Lamb, ree 13: 254 (1947) T.

pad (Zahlbr.) LM. Lamb, Lilloa 13: 261 (194

Lecanora patagonica Zhalbr., Kongl. Svenska IA Handl. 57 (6): 35 (1917) T.

perrugosa (Nyl.) Nyl., Lich. Nov. Zel.: 57 (1888).

perrugosa f. activa 1.M. Lamb, Lilloa 13: 272 (1947) T.

pyenotheca 1.M. Lamb ex Rásinen, Anales Soc. Ci. Argent. 128: 139 (1939) T.

rhodophthalma (Mill. Arg.) Rásánen, Ann po Bot. Zool.-Bot. Fenn. “Vanamo” 2 (1): 25 (1932).

stenophylla (Hue) LM. Lamb, Lilloa 13: 257 (1947).

Lecanora stenophylla Hue, Ann. Mycol. 12: E (1914) T.

terricola (Cromb.) LM. Lamb, Lilloa 13: 263 (1947).

Lecanora gelida f. terricola Cromb., Bot. J. Linn. Soc. 15: 232 (1876) T.

tuberculifera (IM. Lamb) Follmann. In Follman et al. , Crypt. Bot. 2 as 303 (1991).

Placopsis cribellans f. tuberculifera IM. Lamb, Lilloa 13: 228 (1947) T.

Lit.: Lamb (1947); Redon (1985); Jacobsen é: Kappen (1988); Follmamn et al. (1991); Ott et al. (1997).

187. Placynthiella Gyelnik (1939)

[Lecanorales: Trapeliaceae]

uliginosa (Schrad.) Coppins $: P. James, Lichenologist 16: 244 (1984).

Lit.: Coppins £ James (1984); Nimis (1997).

8. Placynthium (Ach.) S.F. Gray (1821)

cera Placynthiaceae]

arachnoideum Henssen, Lichenologist 16: 265 (1984).

rm (Ach.) Trevis., Lich. Veneta, No. 98 (1869).

ni, Huds.) S.F. Gray, Nat. Arr. Brit. PL: 395 (1821).

Lit.: Henssen (1963b, 1984).

189. Placynthiopsis Zahlbr. (1932)

[Peltigerales: Placynthiaceae]

ollmannii C.W. Dodge, Nova Hedwigia 15: 306 (1968) T.

Lit.: Dodge (1968).

W.L. Culb. £ C.F. Culb. (1968)

fhecisanles: Parmeliaceae

(L.) W.L. Culb. $ C.F. Culb., Contr. U. S. Natl. Herb. 34: 530 (1968).

Lit.: Culberson € Culberson (1968); Elix (1993 y

“Plectocarpon Fée (1825)

[Arthoniales:

lichenum (Scontrto) D. Hawksw.. Lichenologist 16: 86 (1984).

148

Checklist of Chilean lichen: GALLOWAY, D.J. £ VW. QUILHOT

pseudosticta (Fée) Fée, Essai Crypt. Ecorc., Suppl.: 147 (1837).

Lit.: Hawksworth $ Galloway (1984); Diederich $ Etayo (1994); Kondratyuk $ Galloway (1995).

191. Pleopsidium Kórb. (1855)

[Lecanorales: Lecanoraceae]

Ed rea (Wahlenb.) Zopf., Ann. Chem. 284: 117 (1895).

rella antarctica J.S. Murray, Trans. Roy. Soc. New Zealand Bot. 2: 60 (1963).

TAL: Redon proa Hafellner (1993a, 1995); Castello $ Nimis (1994).

192. Pleurotrema Miill. Arg. (1885)

[Pyrenulales: Pleureotremataceae

leptosporum Mill. Arg., Miss. Sci. Cap Horn, Lich.: 170 (1888) T.

Lit.: Miller Argoviensis (1888).

193. Poeltiaria Hertel (1984)

[Lecanorales: Porpidiaceae]

corralensis (Rásánen) Hertel, Beih. Nova Hedwigia 79: 431 (1984

Lecidea corralensis Rásinen, Revista Univ. pp 22: 00 (1937) eE

Lecidea patagonica 1.M. Lamb, Farlowia 4: 440 (1955).

Lit.: Hertel (1984); Rambold (1989).

194. Poeltidia Hertel $ o (1984)

[Lecanorales: Porpidiace

perusta (Nyl.) el 2 Hafellner, Beih. Nova Hedwigia 79: 463 (1984).

Lecidea perusta Nyl. In J. Crombie, J. Bot. 13: 334 (1874).

Lit.: Hertel (1984, 1987b, 1989); Rambold (1989).

*Polycoccum Sauter ex me (1865)

[Dothideales: Dacampiac

pulvinatum (Eitner) R pd Lich. Sweden, Norway: 175 (1993).

gulosarium (Linds.) D. Hawksw. In Pegler et al., Kew Bull. 35: 513 (1980)

LA: egos £ Diederich (1988); Triebel (1989); Aptroot € Van der Knapp (1993); Santesson (1993); Wedin

(199

195. Porina Miill.Arg. (1883) nom. cons.

[Trichotheliales: Trichotheliaceae]

depressula Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11): 322 (1924) T.

epiphylla (Fée) Fée, Essai Crypt. Ecorc.: 76 (1825).

Sernandeziana Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11): 320 (1924) T.

fulvella Mill. Arg., Flora, Jena 66: 335 (1883).

leptalea (Durieu ££ Mont. ) A.L. Sm., Monogr. Brit. Lich. 2: 333 (1911).

Porina rufocarpella Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3 (11): 321

(1924) T

limbulata (Kremp. ) Vain., Ann. Acad. Sci. Fenn. Ser. A, 15: 363 (1921).

micromma (Mont.) A. Massal., Framment. Lich.: 26 (1855).

nitidula Miill. Arg., Flora 66: 336 (1883).

rubrosphaera R. Sant., Symb. Bot. Upsal. 12: 261 (1952) T.

macroverrucosa P.M. McCarthy, Biblioth. Lichenol. 52: 67 (1993).

Porina depressula Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11): 322

24) T - nom. illeg. non P. depressula Múll. Arg 0 Jahrb. 6: 399 (1885)].

tetracerae, (Ach.) Miill. Arg., Bot. Jahrb. Syst. 6: 401 (1885

. McCarthy, Biblioth. Lichenol. 52: 116 (1993) T

Lit.: Santesson ( 1952); Follmann (1961); McCarthy (1993).

196. Porocyphus Kórb. (1855)

[Lichinales: Lichinaceae]

ispersus Á. E.Dahl, Meddel. Gronland, Biosci. 150 (2): 51 (1950).

Lit: Henssen (1963c, 1977).

197. Porpidia Kórb. (1855)

[Lecanorales: Porpidiaceae]

149

Gayana Bot. 55(2), 1998.

albocaerulescens (Wulfen) Hertel £« Knoph, Beih. Nova Hedwigia 79: 433 (1984).

austroshetlandica Hertel. Beih. Nova Hedwigia 79: 434 (1894) T.

crustulata (Ach.) Hertel £ Knoph, Beih. Nova ms 79: 435 (1984).

Lecidea oblonga Riedl, Sydowia 20: 343 (196

rita follmannii Riedl, Sydowia 20: 345 e E.

ea magellanica Riedl, Sydowia 20: 346 (1968) T.

Lit.: Hertel 1971. 1984, 1987a).

198. Protoparmelia M. Choisy (1929)

[Lecanorales: Lecanoraceae

badia (Hoffm.) Hafellner, Beih. Nova Hedwigia 79: 292 (1984).

Lecanora badia var. fuscobrunnea Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 156 (1855) T.

Lit.: Hafellner (1984); Nimis (1993).

199. Protoroccella Porres

[Arthoniales: Roccellaceae]

nnii San SP á: Schulz, Flechten Follmann: 319 (1995) T.

minima (R. Sant.) Follmann, Crypt. Bot. 5: 230 (1995).

Roccella minima R. Sant., Ark. Bot. 31A (7): 6 (1944) T.

Lit.: Santesson (1944); Follmann (1995); Sánchez-Pinto £ Schulz (1995).

200. Protousnea (Mot.) Krog (1976)

[Lecanorales: Parmeliaceae]

alectoroides (Mot.) Krog, Norw. J. Bot. 23: 99 (1976).

Usnea alectoroides Mot., Usnea 1: 13 ion de

dusenii (DR.) Krog, Norw. J. Bot. 23: 99 (1976).

Usnea dusenii DR., Svensk Bot. Tidskr. 20: 93 (1926).

Usnea barbata var. oxygona Miill. Arg., Hedwigia 34: 139 ais E

Usnea oxygona (Múll. Arg.) Zahlbr., Cat. Lich. Univ. 6: 588 (1930).

Usnea santessonii Rásánen, Arch. Soc. Zool.-Bot. Ho “Vanamo” 2: 45 (1949).

magellanica (Mont.) Krog, Norw. J. Bot. 23: 100 (1976).

Evernia magellanica Mont., Ann. Sci. Nat. Bot. sér. 2, 20: 356 (1843) T.

000 desea vpo ) Mot., Usnea 1: 9 (1936

eth ar. tonellii Sambo, Contr. Sci. Miss Sales. Ven. Don Bosco: 40 (1926) T.

Pes (Stirt.) Krog, a J. Bot. 23: 101 (1976).

Chlorea malacea Stirt., Scott. Naturalist (Perth) 7: 74 (1883).

Usnea malacea (Stirt.) Zahlbr., Cat. Lich. Univ. 6: 600 (1930).

Usnea campestris R. Sant., Ark. Bot. 30A (6): 9 (1942).

poeppigii (Nees £ Flot.) Krog., Norweg. J. Bot. 23: 102 inca

Neuropogon poepiggii Nees € Flot., Linnaea 9: 497 (18

Chlorea poeppigii (Nees € Flot.) Ny]l., Syn. Meth. Lich. pres 275 (1860).

Usnea poeppigii (Nees £ Flot.) Vain., Résult. Voy. Belgica, Lich.: 12 (1903).

Letharia poeppigii (Nees $: Flot.) Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, 3 (11): 33

(1912).

pet poeppigii (Nees £ Flot) Elenkin, Izv. Imp. Bot. Sada Petra Velikago 16: 266 (1916) nom.

nval.

scrobicula Sambo) pp Norweg. J. Bot. 23: 102 (1976).

ar. scrobiculata Sambo, Contr. Sci. Miss. Sales. Ven. Don Bosco: 40 (1926) T.

Ps Krog, ER J. Bot. e 103 (1976) T

sp. in Krog, Norw. J. Bot. 23: 105 (197

Lit.: Santesson (1942a); Krog (1976); 00d et al. (1985); Garbarino et al. (1987); Quilhot et al. (1987).

201. Pseudephebe M. pines (1930)

[Lecanorales: Parmeliacea:

minuscula (Nyl. ex A Brodo £ D. Hawksw., Opera Bot. 42: 140 (1977).

pubescens (L.) M. Choisy, Icon. Lich. Univ. ser. 2: sine pag. (1930).

Lit.: Brodo £« Hawksworth (1977); Redon (1985); Garbarino et al. ( 1993).

pea er reiciós (1890) nom. cons.

os jariaceae]

argyracea ra Vain.. Hedwigia 37: 35 (1898).

: > Srcta agyracea Dels, Mém. Soc. Linn. Calvados 2: 91 (1825).

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

aurata (Ach.) Vain., Acta Soc. Fauna Fl. Fenn. 7: 183 (1890).

bartlettii D.J. Galloway, Lichenologist 17: 303 (1985).

Stictina mougeotiana f. isidiosa Múill. Arg., Bull. Herb. Boissier 4: 89 (1896).

berberina (G. Forster) D.J. Galloway 8 P. James, Lichenologist 9: 95 (1977).

Lichen berberinus G. Forster, Comment. Soc. Sci. Gotting. 9: 44 (1787).

Sticta orygmaea Ach., Methodus: 278 (1803).

Lobaria orygmaea (Ach.) Hellbom, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 21 (3/13): 36 (1896).

Pseudocyphellaria orygmaea (Ach.) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25 (3/6): 28

(1899)

Sticta durvillei Delise, Mém. Soc. Linn. Calvados 2: 599 (1825).

Lobaria urvillei (Delise) Trevis, Lich. Veneta Exs. 75 (1869).

Sticta endochrysa var. urvillei (Delise) Miill. Arg., Miss. Sci. Cap Horn, Lich.: 157 (1889).

Pseudocyphellaria durvillei (Delise) Vain., Hedwigia 38: 187 (1899).

Pseudocyphellaria orymaea var. urvillei (Delise) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25

(3/6): 28 (1899).

Sticta endochrysa sense Hook. f., Fl. Antarct. 2: 525 (1847) non Delise

Sticta endochrysa var. angustiloba Mont. In C. Gay, Hist. Chile, Bot. 8: 105 (1854) T.

Sticta urvillei var. orygmaeoides Nyl., Syn. Meth. Lich. 1 (2): 360 (1860) T

Sticta endochrysa var. orygmaeoides (Nyl.) Múill. Arg., Miss. Sci. Cap Horn, Lich.: 157 (1889).

Pseudocyphellaria endochrysa var. orygmaeoides (Nyl.) Follmann, Revista Univ. (Santiago) 51: 42 (1966)

nom. inval.

Sticta endochrysa var. compacta Miill. Arg., Miss. Sci. Cap Horn, Lich.: 157 (1889) T.

Sticta urvillei var. orygmaeoides f. compacta (Mill. Arg.) Stizenb., Flora 81: 117 (1895).

Pseudocyphellaria durvillei var. laciniatula Rásiinen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 36

(1932) T

Pseudocyphellaria durvillei var. normalis Rásiinen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 36

(1932) T.

Pseudocyphellaria durvillei var. canescens Rásiinen, Revista Univ. (Santiago) 22: 204 (1937) T.

bereteroana (Mont.) Redon, Anales Mus. Hist. Nat. Valparaíso 10: 21 (1977).

Sticta coerulescens Mont., Amn. Sci. Nat. Bot. sér. 3, 18: 306 (1852) T. :

Pseudocyphellaria coerulescens (Mont.) D.J. Galloway $: P. James, Lichenologist 12: 294 (1980) nom. su-

rfl

pertt.

Pseudocyphellaria nitida var. curta Rásánen In Gyeln., Lich. Parva No. 5 (1937).

compar (Ny1.) H. Magn., Acta Horti Gothob. 14: 8 (1940).

Stictina compar Ny]., Flora 49: 135 (1866) E.

Sticta compar (Nyl.) Zahlbr., Cat. Lih. Univ. 3: 375 (1925

Pseudocyphellaria orwayensis (Jatta) H. Magn., Acta Horti Gothob. 14: 7 (1940).

Cyanisticta otwayensis (Jatta) C.W. Dodge, Trans. Amer. Microscop. Soc. 84: 506 (1965).

Sticti 7 ill. Are.. Flora 74: 111 (1891) T.

cda mp S h. Kongl. Svenska Vetensk.-Akad. Handl. 25

Sticta endochrysoides (Miill. Arg.) Hue, Nouv. ergo 0000 degrco po 3: 87 (1901).

Pp ¡cti ¿des (Múll. Arg.) Clem., Gen. Fung.:

memos od 7 dos ánen, Revista. Univ. (Santiago) 22: 204 ( 1937).

ifera Hi ¡ sr. 4, 3:86 (1901)

Sticta granulifera Hue, Nouv. Arch. Mus. Hist. Nat. sér. 4, 3:86 ( )

Pseudocyphellaria granulifera (Hue) H. Magn., Acta Hortt Gothob. 14: 8 (1940).

Pseudocyphellaria compar var. granulifera (Hue) R. Sant. In S. Huneck et al., J. Hattori Bot. Lab. 37: 545

(1973)

coppinsii D.J. Galloway, Biblioth. Lichenol. 46: 85 (1992) T. :

coriifolia (Miill. Arg.) old Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25 (3/6) erecta

Sritina coriifolia MúN. Arg.. Bot. Jahrb. Syst.4: 55 (1889 E

Sticta coriifolia (Mill. Arg.) Zahlbr., Cat. Lich. Univ. - : “ ”2 (1): |

Cyanisticta coriifolia (Múll. Arg.) Rásánen, Ann. Bot. Soc. Zool tot e. bucal a (1): 40 (1932).

Stictina coriifolia var. hypomelaena Miill. Ag. Miss. Sci. Cap Eno Ku Varese Acad: Hand. 25

Pseudocyphellaria coriifolia var. hypomelaena (Múll. Arg) Malme, Bib. £- Sy. YeErse O

(3/6): 25 (1899). E a

Sticta coriifolia var. hypomelaena (Mill. Arg.) Zahlbr., Cat. Lich. Univ. 3: 376 (1925).

151

Gayana Bot. 55(2), 1998.

po coriifolia var. hypomelaena (Múll. Arg.) Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo”

2 (1): 40 (1

crocata (L.) Vain., Hedwigia 37: 343 (1898).

dissimilis (Nyl. )D .J. Galloway £ P. James, Lichenologist 12: 297 (1980).

divulsa (Taylor) Imshaug, Adapt. Antarct. pp pra 964 (1977).

Sticta divulsa Taylor, Lond. J. Bot. 6: 182 (184

Sticta billardierei var. divulsa CoN) Hook. E és _Antarct. 2: 527 (1847).

Sticta fossulata f. divulsa (Taylor) Stizenb., Flora 81: 114 (1895).

Sticta richardi f. divulsa (Taylor) Hue, Nouv. Arch. Mus. Hist. Nat. sér. 4, 3: 56 (1901).

Pseudocyphellaria richardi var. divulsa nda Follmann, Revista Univ. (Santiago) 51: 53 (1966).

Rietz, Ark. Bot. 20 B (1): 2 (1926

Cyanisticta dubia (Du Rietz) Rásiánen, Amn. Bot. Soc. Zool.-Bot. os “Vanamo” 2 (1): 42 (1932).

Sticta (Stictina) dubia (Du Rietz) Zahlbr., Cat Lich. Univ. 8: 309 (193

Cyanisticta dubia var. arrhiza Rásinen, Ann. Bot. Soc. Zool.-Bot Fon apo 2 (1): 42 (1932) T.

Sticta dubia var. arrhiza (Rásánen) Zahlbr., Cat. Lich. Univ. 10: 274 (19

Pseudocyphellaria dubia var. arrhiza (Rásánen) F Revista aria O 51: 42 (1966).

encoensis R. Sant. In D.J. Galloway, Lichenologist 21: 88 (1989) T

endochrysa (Delise) Vain., Résult. Voy. Belgica, Lich.: 28 (1903).

Sticta endochrysa Delise, Mém. Soc. Linn. Calvados 2: 43 (1825).

Ricasolia endochrysa (Delise) Trevis., Le Veneta Exs. 75 A

Lobaria endochrysa (Delise) Kuntze, Rev. Gen. Pl. 2: 876 (1891

Sticta orygmaea var. endochrysa (Delise) Zahlbr., Cat. Lich. Univ. 8: 312 (1932).

Sticta feei Delise, Mém. Soc. Linn. Calvados 2: 44 (1825).

Sticta endochrysa var. feei (Delise) Zahlbr., Cat. Lich. Univ. 3: 343 (1925).

Parmelia pubescens Pers. In M. Pra pe Voy. Uranie Bot. 1: 199 (1827).

Sticta endochrysa var. pul Nyl., Syn. Meth. Lich. 1 (2): 359 (1860).

Sticta lechleri Flot. In W. Lechler, ano Amer. Austral.: 49 (1857) nom. nu

Pseudocyphellaria leptospora Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” o» (1):37 (1932) T.

Pseudocyphellaria latiloba C.W. Dodge, Trans. Amer. Microscop. Soc. 84: 503 (1965).

exanthematica 1.M. Lamb, Farlowia 4: 435 (1955).

Saveolata (Delise) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25 (3/6): 23 (1899).

Sticta faveolata Delise, Mém. Soc. Linn. Calvados 2: 102 (1825) T

Stictina faveolata (Delise) Nyl., Syn. Meth. Lich. 1 (2): 337 (1860).

ota os (Delise) Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 42 (1932).

(J.D. Hook. £: Taylor) Vain., Philipp. J. Sci. 8: 115 (1913).

Sticta endochrysa sensu Hook. f., Fl. Antarct. 1: 525 (1847) non Delise.

Sticta urvillei var. flavicans (Hook. F. 8: Taylor) Nyl. Syn. pac Lich. 1 (2): 360 (1860).

Sticta endochrysea [sic] var. flavicans (Hook. F. £ Taylor) M 11. Arg., Flora 71: 136 pri

Pseudocyphellaria durvillei var. flavicans (Hook. F. €: Taylor) Vain., Hedwigia 38: 187 (18

Pseudocyphellaria orygmaea var. flavicans (Hook. F. €: Taylor) Malme, Bih. Kongl. a Vetensk.-

Akad. Handl. 25 (3/6): 28 (1899).

Sticta desfontainii .

Stictina mallota (Tuck.) Stizenb, Flora, Jena 81: 131 (1895).

Stictbd stauromatica Flot. In W. Lechler, Pl. Chilensem 562a (?1854

A E

freycinetii (Delise) Malme, Bih. Kongl. Svenska Vetensk.-Akad. + 25 (3/6): 34 (1899).

152

Sticta freycinetii Delise, Mém. Soc. Linn. Calvados 2: 124 (18

Lobaria freycinetii (Delise) Trevis., Lich: Veneta Exs. 75 ds

Phanosticta freycinetii (Delise) Clem., Gen. Fung.: 175 (1909).

Parmelia lactucaefolia Pers. In M. Gaudichaud-Beaupré, Voy. Uranie Bot.: 199 (1827).

Sticta lactucaefolia (Pes.) Nyl., Lich. Nov. Zel.: 39 (1888).

Sticta freycinetii var. lactucaefolia (Pers.) Miill. Arg., Miss. Sci. Cap Horn, Lich.: 157 (1888).

pra de freycinetii var. lactucaefolia (Pers.) Mame, Bih. Kongl. Svenska Vetensk.-Akad. Handl.

Sticta fulvocinerea Menk:. Voy. Pole Sud Bot.1: 184 (1845) T.

Sticta freycinetii var. fulvocinerea (Mont.) Nyl., Mém. Soc. Sci. Nat. Cherbourg 5: 103 (1859).

fulvocinerea (Mont.) Trevis., Lich. Veneta Exs. 75 (1869).

Sticta freycinetii var. fimbriata Mont. In C. Gay, Hist. Chile, Bot. 8: 121 (1854) T.

Ps Uria freyinetii var. fimbriata (Mont.) Follmann, Revista Univ. (Santiago) 51: 44 (1966).

Sticta patagonica Miill. Arg., Hedwigia 34: 140 (1895).

Pseudocyphellaria patagonica (Mill. Arg.) LM. Lamb, Anales Parques Nac. 7: 66 (1959) [“1958”].

Cyanisticta antarctica Rásánen, Ann. Bot. Soc - Zool.-Bot. Fenn. “Vanamo” 2 (1): 39 (1932) T.

Checklist of Chilean lichen: GALLOWAY, D.J. $: VW. QUILHOT

Sticta (Stictina) antarctica (Rásánen) Zahlbr., Cat. Lich. Univ. 10: 272 (1939).

gilva (Ach.) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25 (3/6): 32 (1899).

Sticta erythroscypha Taylor, Lond. J. Bot. 6: 181 (1847) T

Stictina subfaveolata Nyl. In E. Stizenberger, Flora 81: 129 (1895) T.

glabra (Hook. f. £« Taylor) C.W. Dodge, B.A.N.Z.A.R.E. Rep. B, 7: 79 (1948).

Sticta glabra Hook. f. £ Taylor, Lond. J. Bot. 3: 647 (1844).

Sticta freycinetii var. glabra (Hook. f. £ Taylor) Zahlbr. Cat. Lich. Univ. 3: 347 (1925).

Pseudocyphellaria nio var. glabra (Hook. f. £ Taylor) Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn.

“Vanamo” 2 (1): 3 a

Cetraria (sect. Platy. e la Zahlbr., Kongl. Svenska Vetenskapsakad. Handl. 57 (6): 43 (1917).

Pseudocoptellria burkholderi C.W. Dodge, Trans. Amer. Micros cop. Soc. 84: 502 (1965).

granulata (C. Bab.) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25 (3/6): 21 (1899).

guillemini (Mont. ) D.J. Galloway, Lichenologist 18: 127 (1986

Sticta guillemini Mont. Ann. Sci. Nat. Bot. sér. 2, 4: 87 ( 1835) T ;

Sticta hirsuta var. guillemini (Mont.) Mont. In C. Gay, Hist. Chile, Bot. 8: 107 (1854).

Sticta obvoluta var. guillemini (Mont.) Nyl., Mém. Soc. Sci. Nat. Cherbourg 5: 103 (1857).

Crocodia guillemini (Mont.) Nyl. In Hue, Nouv. Arch. Mus. Hist. Nat. sér. 3, 2: 297 (1890).

Cyanisticta guillemini (Mont.) Rásinen, Revista Univ. (Santiago) 22: 203 (1937

Sticta guillemini var. stictica Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11):

248 (1924) T.

guzmanii D.J. Galloway, Biblioth. Lichenol. 46: 145 (1992) T.

hillii (C.W. Dodge) D.J. Galloway, Biblioth. Lichenol. 46: 149 (1992).

Cyanisticta hillii C.W. Dodge, Nova Hedwigia 19: 492 (1971) T

hirsuta (Mont.) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25 (3/6): 18 (1899).

Sticta hirsuta Mont., Ann. Sci. Nat. Bot. sér. 2, 4: 88 (1835) T.

Stictina hirsuta (Mont.) Nyl., Ann. Sci. Nat. Bot. sér. 4, 15: 41 (1861).

Cyanisticta hirsuta (Mont.) Rásánen, Revista Univ. (Santiago) 21: 144 (1936).

Pseudocyphellaria hirsuta f. leucosticta 1.M. Lamb, Farlowia 4: 436 AP

Stictina scholanderi C.W. Dodge, Nova Hedwigia 19: 493 (1971) T

imbricatula (Taylor) D.J. Galloway, Lichenologist 17: 305 (1985).

Sticta imbricatula Taylor, Lond. J. Bot. 6: 180 (1847) T.

Sticta endochrysa f. imbricatula (Taylor) Nyl., Syn. Meth. Lich. 1 (2): 359 (1860).

imshaugii D.J. Galloway $ Redon. In D.J. Galloway, Biblioth. Lichenol. 46: 158 (1992) T.

intricata (Delise) Vain., Hedwigia 37: 35 (1898).

lechleri (Mill. Arg.) Du Rietz. In H. Magnusson, Acta Horti Gothob. 14: 27 (1940).

Stictina lechleri Miill. Arg., Flora 66: 354 (1883) T.

Cyanisticta lechleri (Miill. Arg.) C.W. Dodge, Trans. Amer. Microscop. Soc. 84: 505 (1

Sticta carpoloma f. latifolia Kremp., Verhandl. Zool.-Bot. Ges. Wien 18: 316 (1868) nom. nud.

Stictina latifolia Stizenb., Flora 81: 130 (1895) T

Pseudocyphellaria carpoloma var. latifolia (Stizenb. ) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl.

25 (3/6): 33 (1899).

Sticta latifolia (Stizenb.) Zahlbr., Cat. Lich. Univ. 3: 39 (1925).

Pseudocyphellaria latifolia (Stizenb.) Follmann, Revista Univ. (Santiago) 51: 48 (1966).

Sticta (Stictina) carpoloma var. benovei Sambo, Lich. Terr. D. Fuoco: 31 (1926)

Cyanisticta wilkseii C.W. Dodge, Nova Hedwigia 19: 472 (1971).

mallota (Tuck.) H. Magn., Acta Horti Gothob. 14: 7 (1940).

Sticta crocata var. mallota Tuck., Syn. N. Amer. e 1: 101 (1882) T.

Stictina mallota (Tuck.) Stizenb, Flora 81: 131 (189

Sticta mallota (Tuck.) Zahlbr., Cat. Lich. Univ. 3: ras

Cyanisticta mallota (Tuck.) C.W. Dodge, Trans. Amer. Microsc cop. Soc. 84: 506 (1965).

Cyanisticta hirsuta var. sorediifera Rásinen, Revista Univ. (Santiago) 22: 203 (1937) T.

Pseudocyphellaria hirsuta var. sorediifera (Rássánen) Follmann, Revista. Univ. (Santiago) 51: 46 (1966).

eana D.J. Galloway, Biblioth. Lichenol. 46: 172 (1992) T.

meyenii (Trevis.) D.J. Galloway, Lichenologist 21: 89 (1989).

Stictina meyenii Trevis, Lich. Veneta Exs. 75 Aj ada

Stictina faveolata var. cervicornis Nyl., Syn. ic

po can faveolata var. cervicornis (Nyl.) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl.

25 (3/6): 23 (1899).

Sticta faveolata var. cervicornis (Nyl.) Zahlbr., Cat. Lich. Univ. 3: 380 (1925).

mooreana (Zahlbr.) Imshaug. Adapt. Antarct. a cdo ett

lecta (Mill. Arg.) H. Magn., Acta Horti Go!

ade hn var. isidiifera Rásiinen, Revista Univ. (Santiago) 22: 204 (1937) T.

153

Gayana Bot. 55(2), 1998.

Pseudocyphellaria gilva var. isidiifera (Rásánen) Follmann, Revista Univ. (Santiago) 51: 45 (1966).

nitida (Taylor) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25 (3/6): 26 (1899).

Sticta nitida Taylor, Lond. J. Bot. 6: 178 (1847) T.

Lobaria nitida (Taylor) Trevis, Lich. Veneta Exs. 75 (1869).

Sticta flabellata Mont., Ann. Sci. Nat. Bot. sér. 3, 18: 306 (1852) T.

Sticta horrida Hue, Nouv. Arch. Mus. Hist. Nat. sér. 4, 3: 50 (1901) T.

Pseudocyphellaria horrida (Hue) 1.M. Lamb, Anales Parques Nac. 7: 62 (1959) [“1958”].

Sticta carpoloma f. ornata Zahlbr. In C.Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11):

349 (1924) T.

Pseudocyphellaria carpoloma f. ornata (Zahlbr.) Follmann, Revista Univ. (Santiago) 51: 36 (1966).

Pseudocyphellaria nitida var. subglauca Rásiinen, Revista Univ. (Santiago) 22: 205 (1937) T.

Pseudocyphellaria nitida var. substraminea Rásá Revista Univ. (Santiago) 22: 205 (1937) T.

norvegica (Gyeln.) P. James. In B.J. Coppins « P.W. James, Lichenologist 11: 172 (1979).

nudata (Zahlbr.) D.J. Galloway, Lichenologist 21: 89 (1989).

Sticta coriifolia var. nudata Zahlbr., Ann. Mycol. 29: 81 (1931).

Pseudocyphellaria coriifolia var. nudata (Zahlbr.) Follmann, Revista Univ. (Santiago) 51: 38 (1966).

obvoluta (Ach.) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25 (3/6): 17 (1899).

Parmelia obvoluta Ach., Methodus: 218 (1803).

Cyanisticta piloselloides Rásinen, Amn. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 39 (1932) T.

Sticta piloselloides (Ráisánen) Zahlbr., Cat. Lich. Univ. 10: 278 (1939).

pluvialis R. Sant. In D.J. Galloway, Lichenologist 18: 150 (1986) T.

Pseudocyphellaria nitida var. mollis Rásinen, Revista Univ. (Santiago) 22: 205 (1937) T.

Sticta faveolata var. richardi (Mont.) Linds., Trans. Roy. Soc. Edinburgh 22: 197 ( 1859).

1 J (1860).

Lobaria fossulata var. richardi (Mont.) Hellbom, Bih. Kongl. rencia Vetensk.-Akad. Handl. 21 (3113): 38

1896

scabrosa R. Sant. In D.J. Galloway, Lichenologist 18: 155 (1986) T.

skottsbergii D.J. Galloway, Biblioth. Lichenol. 46: 224 (1992) T

subrubella Rásánen, Anales Soc. Ci. Argent. 128: 142 (1939).

vaccina (Mont.) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25 (3/6): 27 (1899).

Sticta vaccina Mont., Ann. Sci. Nat. Bot. Sér. 3, 18: 307 (1852) T.

Sticta endochrysea [sic.] var. vaccina (Mont.) Cromb., Bot. J. Linn. Soc. 15: 230 (1876).

Pseudocyphellaria albidopallens Vain.. Résult. Voy. Belgica, Lich.: 28 (1903) T.

Sticta albidopallens (Vain.) Zahlbr., Cat. Lich. Univ. 3: 330 (1925).

Pseudocyphellaria vaccina var. albosticta Rásánen, Anales Soc. Ci. Argent. 128: 142 (1939).

ges cias albidopallens f. albosticta (Rásánen) LM. Lamb, Anales Parques Nac. 7: 64 (1959)

vil ( Nyl.) Follmann, Revista Univ. (Santiago) 51: 54 (1966).

Sticta valdiviana Nyl. In E. Stizenberger, Flora 81: 115 (1895) T.

verrucosa (Mont.) D.J. Galloway, Lichenologist 21: 89 ( 1989).

Sticta argyracea var. verrucosa Mont. In C. Gay, Hist. Chile, Bot. 8: 116 (1854) T.

Stictina argyracea var. verrucosa (Mont.) Zahlbr. In C. Skottsber Nat. Hist. Juan Fernández Easter 1s]. 2

(Bot.), 3 (11): 346 (1924). ., sudan

Pse yphellaria argyracea var. verrucosa (Mont.) Follmann, Revista Univ. (Santia 0) 51: 35 (1966

wandae D.J. Galloway, Biblioth. Lichenol. 46: 238 (1992) T. a nó

Lit.: Galloway (1986, 1988, 1989, 1992).

203. Pseudoparmelia Lynge (1914)

[Lecanorales: Parmeliaceae]

154

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

subambigua Hale, Smithsonian Contr. Bot. 31: 50 (1976) T.

Lit.: Hale (1976, 1986); Elix (1993); Elix £ Nash (1998).

204. Pseudopyrenula Miill. Arg. (1883)

[Pyrenulales: ?Trypetheliaceae]

diluta (Fée) Mill. Arg., Flora 66: 249 (1883).

Lit.: Follmann (1961).

205. Psilolechia A. Massal. (1860)

[Lecanorales: Pee

lucida (Ach.) M. Choisy, Bull. Mens. Soc. Linn. Lyon 18: 142 (1949).

Lit.: Coppins 4 Purvis (1987).

206. Psiloparmelia Hale (1989)

[Lecanorales: Parmeliaceae]

distincta (Ny!l.) Hale, Mycotaxon 35: 42 (1989).

flavobrunnea (Miill. Arg.) Elix £ Nash, Bryologist 95: 386 (1992).

salazinica Elix £ Nash, Bryologist 95: 388 (1992).

Lit.: Hale (1989); Elix € Nash (1992).

207. Psora Hoffm. (1796) nom. cons

[Lecanorales: Psoraceae]

icterica (Mont.) Mill. Arg., Flora 71: 45 (1888

Biatora icterica Mont., Ann. Sci. Nat. Bot. 2: 373 (1834) T.

Lecidea icterica (Mont.) Taylor, Lond. J. Bot. 6: 150 (1847).

Schaereria argentina Gyebe, Repert. Spec. Nov. Regni Veg. 33: 304 (1934).

Lecidea (Gyeln.) Zahlbr., Cat. Lich. Univ. 10: 346 (1939).

diiformis (Hue) Darb., Br. Antarct. (“Terra Nova”) Exp. 1910, Nat. Hist. Rep. Bot. 3: 48 (1923).

NE: remtrbióa Timdal (1984, 1987).

208. Psoroma Ach. ex Michaux (1803)

[Lecanorales: Pannariaceae

angustisectum Zahlbr. In C. prod mo cs Juan Fernández Easter Is. 2 (Bot.), 3 (11): 342 (1924) T.

aphthosum Vain., Hedwigia 38: 188 (18

caliginosum Stirt., Proc. Phil. Soc. o po 295 (1877).

calophyllum Miill. Arg., Hedwigia 31: 278 (1 1892) T.

cephalodinum Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández

cinnamomeum Malme, Ark. Bot. 20A (3): 11 (1925) T.

contortum Mill. Arg., Miss. Sci. Cap Horn, Lich.: 160 (1889) T.

cristatulum Mill. Arg., Hedwigia 31: 278 (1892)

dasycladum Zahlbr. In C. Skottsberg, Nat. Hist. lem Fernández Easte

dimorphum Malme, Ark. Bot. 20A (3): 11 (1925).

durietzii P. James 8: Henssen, Lichenologist 7: 143 (1975).

fruticulosum P. James €: Henssen, Mycotaxon 18: 31 (1983).

hirsutulum Nyl. ex Cromb., J. Bot. Lond. 13: 333 (1875).

hispidulum Nyl., Flora, Jena 38: 674 (1855) T.

Pannaria hispidula (Nyl.) Hue, Bull. Soc. Bot. France 48: LVH (1902).

hypnorum (Vahl) S.F. Gray, Nat. Arr. Br. Pl. 1: 445 (1821).

Seibo follmannii C.W. Dodge, Trans. Amer. Microscop. Soc. 84: 510 (1965) T.

pnorum var. paleaceum (Fr.) Rostr., Bot. Tidsskr. 4: 96 (1871

implezumo Str Trans. 8: Proc. New Zealand Inst. 6: 236 (1874).

incisum Vain., Hedwigia 38: 188 (1899) T.

Easter Isl. 2 (Bot.), 3 (11): 339 (1924) T.

r Isl. 2 (Bot.), 3 (11): 341 (1924) T.

leprolomum (Nyl. ) Rásiinen, Ann. Bot. Soc. Zool.-Bot. F

microlepideum Malme, Ark.Bot. 2304 (3): 17 (1925) T. ;

microlepideum var. pallida Rásinen, Ann. Soc. Bot. Zool.-Bot. Fenn. “Vanamo” 2 (1): 45 (1932) T.

microphyllizans (Ny1.) D.J. Pia pe es J. Bot. 21: 196 (1983).

m Ny]l., Syn. Meth. Lich. 2: 25 (186

Pannaria pallida (Nyl.) Hue, Bull. Bot. qe France 48: LVI (1902).

7 enssen, Mycotaxon 18: 98 (1983).

155

Gayana Bot. 55(2), 1998.

patagonicum Malme, Ark. Bot. 20A (3): 13 (1925) T.

Pannaria sphictrina var. dilatata Hue, Nouv. Arch. Mus. Hist. Nat. sér. 4, 8: 267 (1906) T.

Psoroma sphictrinum var. dilatatum (Hue) Zahlbr., Cat. Lich. Univ. 8: 303 (1932).

lidotoides (Nyl.) Trevis. Lich. Veneta: 98 (1869).

Pannaria reticulata Hue, Nouv. Arch. Mus. Hist. Nat. sér. 4, 8: 261 (1906) T.

Psoroma reticulatum (Hue) Zahlbr., Cat. pepris Univ. 3: 274 (1925).

pholidotum (Mont.) Miúill. Arg., Flora 71: 45 (188

Parmelia pholidota Mont., Amn. Sci. Nat. ey sér. 2, 4: 91 (1835) T.

Pannaria pholidota (Mont.) Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 182 (1855).

pulchrum Malme, Ark. Bot. 20A (3): 12 (1925).

rubro. m P. James £ J.S. Murray, Mycotaxon 18: 40 (1983).

sphinctrinum (Mont.) Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 181 (1855).

Parmelia sphinctrina Mont., Amn. Sci. Nat. Bot. sér. 2, 4: 90 (1835) T.

Parmelia rubiginosa var. sphinctrina (Mont.) Hook. f. £ Taylor, Fl. Ant. 2: 533 (1847).

Lecanora sphinctrina (Mont.) Nyl., Bull. Soc. Linn. Normandie, sér. 2, 2: 62 (1868).

Pannaria sphinctrina (Mont.) Hue, Bull. Soc. Bot. France 48: LVI (1902).

vulcanicum Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3 (11): 339 (1924) T.

Lit.: Zahlbruckner (1924); Malme (1925); Lamb (1959); Follmann $ Huneck (1976); Henssen £ Renner (1981);

Henssen et al. (1983); Henssen (1983); Galloway (1985); Quilhot et al. (1989); Calvelo (1992).

209. Psoromidium Stirt. (1877)

[Lecanorales: Pannariaceae

versicolor (Hook. f. £ Taylor) D.J. Galloway, New Zealand J. Bot. 21: 196 (1983).

Lecanora versicolor Hook. f. £ Taylor, London J. Bot. 3: 642 (1844).

Psoroma versicolor (Hook. f. € Taylor) Miill. Arg., Flora 71: 534 (1888).

Psoroma subdescendens Nyl., Lich. Fueg. Patag.: 7 (1888).

Pc subdescendens les ) Nyl. In Hue, Nouv, Arch. Mus. Hist. Nat. Paris sér. 3, 3: 45 (1891).

a subdescendens (Nyl.) Hue, Mém. Soc. Nat. Hist. Nat. Math. Cherbourg 38: 49 (1911).

Lit: Cllóma l K James (1985).

210. A. Masal. (1855)

[Lichinales: Lichinaceae ]

argentinica Miill. Arg., Flora 72: 142 (1889).

Lit.: Lamb (1959); Follmann (1965b).

211. Psorula Gotth. Schneid. (1979)

[Lecanorales: Psoraceae]

rufonigra (Tuck.) Gotth. Schneid., Biblioth. Lichenol. 13: 136 (1979).

Lit.: Huneck $: Follmann (1976); Schneider (1979).

212. Pterygiopsis Vain. (1890)

per Lichinaceae

a Henssen, Symb. Bot. Upsal. 18: 76 (1963) T.

Lit.: e (1963c); Jorgensen (1990).

213. Punctelia Krog (1982)

[Lecanorales: Parmeliaceae]

borreri (Sm.) Krog, Nordic J. Bot. 2: 291 (1982).

reddenda (Stirt.) Krog, Nordic J. Bot. 2: 291 (1982

peras (Delise in Duby) Krog, Nordic J. Bot. 2: 291 (1982).

(Nyl.) Krog, Nordic J. Bot. 2: 29 1 (1982).

E -Krog (1982) a dl 00d, Stenroos (1991); Elix (1993, 1994); Adler (1996).

*Pyrenidium Ny]l. (1865)

[Dothideales: Dacampiaceae

actinellum Nyl.. an dd 210 (186:

Lit: Alerces (109: coo Wedin (1994); Ihlen (1998).

214. Pyrenocollema Reinke (1895)

[Dothideales: Arthopyreniaceae]

156

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

halodytes (Nyl.) R.C. Harris. In R. Egan, Bryologist 90: 164 (1987).

Lit.: Santesson (1939).

215. Pyrenula Ach. (1810) nom. cons.

[Pyrenulales: Pyrenulaceae

apistea (Afzel.) Ach., Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck.

Gesammten Naturk.. 6: 17 (1814).

cereina Eschw., Syst. Lich.: 25 (1824).

coryli (Nyl.) A. Mans. Ric. Auton. Lich. pe e (1852).

kunthii Fée, po Essai Crypt. Ecorc.: 80 (18.

mamillana (Ach.) Trevis., Conspect. Verruc.: La lol

nitida (Weigel) Ach., Syn. Meth. Lich.: 125 (1814).

nitidella (Schaer.) Múill. Arg., Bot. Jahrb. Syst. 6: 414 (1885).

occulta var. leucomma (Nyl.) Zahlbr., Cat. Lich. Univ. 1: 448 (1922

Verrucaria micromma var. leucomma Nyl., Amn. Sci. Nat. Bot. sér. 4, 3: 176 (1855) T.

tdi micromma var. leucomma (Ny]l. Ai Conspect. Verruc.: 13 (1860).

vernicosa (Kremp.) Miill. Arg., Flora 72: 68 (1889).

Ext: dra (1924); Follmann (1961); Follmann € Huneck (1976)..

216. Pyrrhospora Kórb. (1855)

los: Lecanoraceae]

russula (Ach.) Hafellner. In Kalb £ Hafellner, Herzogia 9: 86 (1992).

Lit: desrernat (1993b).

217. Pyxine Fr. (1825)

pc Physciaceae]

br. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11): 402 (1924) T.

Ext: pio (1987).

218. Raciborskiella Hóhn. (1909)

clar incert. sed.: Strigulaceae]

ina (Múll.Arg.) R. e. Symb. Bot. Upsal. 12: 197 (1952).

Lit: a da

19. Racodium Fr. (1829) nom. cons.

[Mitosporic fungi]

pestre Pers, Tent. Disp. Meth. Fung.: 76 (1797).

Lit.: Hawksworth (1970); Hawksworth £ Riedl (1977).

220. Ramalina Ach. (1810) nom. cons.

[Lecanorales: Ramalinaceae

cactacearum Follmann, Nova Hedwigia 14: 256 (1968) [”1967"] T. = .

Desmazieria cactacearum (Follmann) Follmann, Philippia 3: 86 (197

Ramalina follmannii C.W. Dodge, Nova Hedwigia 16: 493 (1969) [' dl sd

canariensis Steiner Ost. Bot. Z. 9: 8 (1904

celastri (Spreng.) Krog £ Swinscow, Norwe eg. J. Bot. 23: 159 (1976).

amalina striatula Nees £ Flot., Linnaea 9: 501 (1835) T

Ramalina ecklonii var. ambigua Mont. In C. Gay, Hist. Chile, Bot. 8: 79 (1852)

ceruchoides H. Magn. In Bendz, Santesson £ Wachtmeister, Acta pampa Scand. e 1185 (1965) T.

chilena (Nyl.) Kashiw., Bull. Natn. Sci. Mus. Tokyo ser. B, 16 (1): 2 (1990).

Ramalina pollinaria f. chilena Nyl., Bull. Soc. Linn . Normandie Pe 150 (1870) T.

chilensis Bertero ex Nyl., Bull. Soc. Linn. Normandie sér. 2, 4: 124 (1870) T.

Desmazieria chilensis (Bertero) Follmann 4 Huneck, Willdenowia 5: 208 (1969).

dissecta Kashiw., Bull. Natl. Sci. Mus. N. S. 14: 125 (1988).

pilulifera Ta lor, London J. Bot. 6: 190 (1847) T.

pea scopulorum var. linearis (Sw.) Mont., Ann. Sci. Nat. Bot. sér. 2, 4: 86 (1835).

Ramalina subulata Mont., Ann. Sci. Nat. Bot. sér. 3, 18: id

terebrata Hook. f. 8: Taylor, London J. Bot. 3: 654 (1844

unilateralis Vils., Vict. Nat. 6: 69 (1889).

usnea (L.) Howe, Bryologist 17: 81 (1914).

157

Gayana Bot. 55(2), 1998.

Lit.: Huneck £ Follmann (1967a); Follmann £ Huneck (1969); Follmann (1976); Stevens (1987); Kashiwadani

(1989); Kashiwadani £ Kalb (1993); Blanchon et al. (1996); Quilhot et al. (1996).

221. Redonia C.W. Dodge (1973)

[Lecanoraceae: Physciaceae]

hile C.w. Dodge, Lich. Fl. Antarct. Continent $ Adj. Ids: 353 (1973) T.

cladocarpiza (1.M. Lamb) C.W. Dodge, Lich. Fl. Antarct Continent: 354 (1973).

uellia cladocarpiza 1.M. Lamb, Br. Antarct. Surv. Sci. i. Rep. 61: 24 (1968).

Lit: Dodge (1973), Redon (1982).

222. Reinkella Darb. (1897)

[Arthoniales: Roccellaceae]

irellina Darb., Bull. Herb. Boissier 5: 764 (1897).

Lit.: Darbishire (1897, 1898a); Tehler et al. (1997).

223. Rhizocarpon Ram. ex DC. (1805)

[Lecanorales: Rhizocarpaceae]

disporum (Hepp) Mill. Arg., Rev. Mycol. 1: 170 (1879).

distinctum Th. Fr., Lich. Scand.1: 625 (1874).

geographicum (L.) DC. In Lam. £ DC., Fl. Fr. 3 ed., 2: 365 (1805).

lindsayanum Rásinen, Revista Sudamer. Bot. 7: 87 (1942

obscuratum (Ach.) A. Massal., Ric. Auton. Lich. Crost.: 103 (1852).

pepe microspermum Zahkbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3 (11):

368 (1924) T.

superficiale (Schaer.) Malme, Sv. Bot. Tidskr. 8, 3: 282 (1914).

De ce barilochense Rásánen, Arch. Soc. Zool.-Bot. Fenn. “Vanamo” 5 (1): 85 (1952).

iridi Wulfen) Kórb., Syst. Lich. Germ.: 262 ( 1855).

Lit: Rima 1956a, 1956b); Hertel (1971); Henssen (1977); Timdal « Holtan-Hartwig (1988); Feuerer (1991).

4. Rhizoplaca Zopf (1905).

pto Parmeliaceae]

ra (Vain.) Redon, Líquenes Antárticos: 81 (1985

Lecanora aspidophora Vain., Résult. Voy. Belgica, Lich.: 19 (1903).

Omphalodina aspidophora (Vain.) Follmamn $ Redon, Willdenowia 6: 422 (1972).

chrysoleuca (Sm.) Zopf, Ann. Chemie 340: 291 (1905).

johowii (Follmann €: Redon) Follmann, Philippia 4 (1): 35 (1979).

phalodina johowii Follmann €: Redon, Willdenowia 6: 419 (1972) T.

(Ramond) Leuckert $: Poelt, Nova Hedwigia 28: 72 (1977).

odina melanophthalma var. obscura (J. Steiner) Follmann £ Redon, Willdenowia 6: 422 (1972).

Lit.: Follmann £ Redon (1972b); Leuckert et al. (1977); Follmann (1979); Redon (1985); Jacobsen é: Kappen

(1988).

*Rhymbocarpus Zo cala

[Ostropales: Odontotrematac

histophorus (NyL) TriebeL Biblioth. Lichenol. 35: 145 (1989).

Lit.: Triebel (1989).

225. Rimelia Hale € Fletcher (1990)

[Lecanorales: Parmeliaceae

cetrata (Ach.) Hale £ Fletcher, Bryologist 93: 26 (1990).

reticulata (Taylor) Hale € Fletcher. A o 93: 28 (1990).

Lit.: Hale dz Fletcher (1990); Elix (1993, 1

226. rom ens

s: Rimulariac

presentaría Hertel 2 id Biblioth. Lichenol. 38: 176 (1990).

insularis (Nyl.) Rambold £ Hertel. In H. Hertel, Lecideac. Exs. Fasc. 8, No. 159 (1985).

psephota (Tuck.) Hertel £ Rambold. In H. Hertel, Mitt. Bot. Miinchen 23: 334 (1987).

Lecidea psephota Tuck, Proc. Amer. Acad. ¿ey Sci. 12: 181 (1877) T.

ella psephota (Tuck.) Hertel, Beih. No va Hedwigia 79: 460 (1984).

Lit.: (Hertel 1985, 1987b); Rambold (1989); Hertel 8 Rambold (1990).

158

Checklist of Chilean lichen: GALLOWAY, D.J. $ W. QUILHOT

227. Rinodina (Ach.) : F. Gray (1821)

[Lecanorales: Physciaceae]

conradii Kórb., Aral Lich. Germ.: 123 (1855)

endochrysodes (Nyl.) Miill. Arg., Nuovo e Bot. Ital. 21: 43 (1889).

Lecanora endochrysodes Nyl. In J. Cromb., Bot. J. Linn. Soc. 15: 232 (1876).

deceptionis 1M. Lamb., Br. Antarct. Surv. Sci. Rep. 61: 65 (1968) T

endophragmia 1.M. Lamb, Br. Antarct. Surv. Sci. Rep. 61: 71 (1968).

gennarii Bagl., Comment. Soc. Crittogam. Ital. 1: A 1).

infuscata (Nyl.) Zahlbr., Cat. Lich. Univ. 7: 522 (1931).

Lecanora infuscata Nyl., Amn. Sci. Nat. Bot. a 4, 3: 156 (1855) T.

nimbosa (Fr.) Th.Fr., Lichenogr. Scand.1: 193 (1871).

occulta (Kórb.) Sheard, Lichenologist 3: 349 (1967).

olivaceobrunnea C.W. Dodge £ G.E. Baker, Ann. Missouri Bot. Gard. 25: 659 (1938).

peloleuca (Nyl.) Múill. Arg., Nuovo Giorn. Bot. Bi 23: 125 (1891).

Lecanora peloleuca Ny]l., Flora 48: 338 (18

Lecanora endochrysodes Ny]l. In J. es ¿a J. Linn. Soc. 15: 232 (1876) T.

Rinodina endochrysodes (Nyl.) Múll. Arg., Nuovo Giorn. Bot. Ital. 21: 43 (1889).

Rinodina antarctica Mill. Arg., Miss. Sci. Cap Horn, Lich.: 163 (1888) T.

Rinodina deceptionis 1.M. Lamb, Br. Antarct. Surv. Sci. Rep. 61: 65 (1968) T.

aff. subcrustacea (Mill. Arg.) Zahlbr., Cat. Lich. Univ. 7: 567 (1931).

thiomela (Nyl.) Mill. Arg., Flora 64: 515 (1881).

turfacea (Wahlenb.) Kórb., Syst. Lich. Germ.: 123 (185

Lit.: Lamb (1968); Mayrhofer (1983, 1985); Redon (1985); e et al. (1998); Jacobsen £ Kappen (1988); Ap-

troot £ Van der Knapp (1993).

228. Roccella DC. (1805)

[Arthoniales: Roccellaceae]

arboricola Follman, Nova Hedwigia 15: 334 (1968) T.

atacamensis Follmann, Herzogia 9: 658 (1993) T.

babingtonii Mont., Ann. Sci. Nat. Bot. sér. 3, 18: 305 (1852).

Roccella montagnei var. peruensis Kremp., Verhandl. Zool.-Bot. Ges. Wien 26: 442 (1876).

Roccella peruensis (Kremp.) Darb., Biblioth. Bot. 45: 47 (1898)

gayana Mont. In C. Gay, Hist. Chile, Bot. 8: 88 (1852) T

portentosa (Mont.) Darb., Biblioth. Bot. 45: 29 (1898).

occella tinctoria var. portentosa Mont. In C. Gay, Hist. Chile, Bot. 8: 84 (1852) T.

Lit.: Darbishire (1897, 1898b); Huneck 4: Follmann (1967b); Follmann (1962, 1968a); Follmann er al. (1993).

229. Roccellaria Darb. (1897)

[Arthoniales: Roccellaceae]

mollis (Hampe) Zahlbr., Cat. Lich. Univ. 2: 511 (1923).

Roccella mollis Hampe, Linnaea 17: 122 (1843) T.

Roccella intricata Mont., Ann. Sci. Nat. Bot. sér. 3, 11: 57 (1849) T.

Roccellaria intricata (Mont.) Darb., Ber. Deutsch. Bot. Ges. 15: 6 (1897).

Lit: Follmann (1962) Follmanmn et al. (1993).

230. Roccellina Darb. em. Tehler (1983)

ps proa

m. (1967)

accedens (NyL ) Tehler, Opera Bot 70: 76 (1983).

Lecidea accedens Nyl., Amn. Sci. Nat. Bot. sér 4, do a a e

east cedens (Nyl.) Zahlbr., Beih. Bot. Cen 1

Schism ' mma pu Bro (Nyl.) Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3

a 327 (1924)

Roccellina olivacea Follmann, Philippia 4 (2) o A 1979) T.

cerebriformis (Mont.) Tehler. Opera Bot. 70: 51

ca cerebriforme Mont., Ann. Sci. Pe Bot. sér. 3, 18: 311 (1852) T.

Lobodirina cerebriformis (Mont.) Follmann, Nova Hedwigia 14: 234 (1968) [*1967”].

Roccellina condensata Darb., Ber. Deutsch. Bot. Ges. 16: 11 (1898) T

Dirina condensata (Darb.) Zahlbr., Acta Horti Gothob. 2: 6 (1925). nomen sed non planta.

eos ont.) Tehler f. o Tehler, Opera Bot. 70: 53 (1983) T.

Tehler, ia Bot. 70: 57 (1983).

prota (C.W. Dodge) Tehler, Opera Bot 70: 59 (1983).

159

Gayana Bot. 55(2), 1998.

Dirinastrum chilenum C.W. Dodge, Nova Hedwigia 12: 324 (1967) [*1966”] T.

Dirinastrum chilense (C.W. Dodge) Follmann, Nova Hedwigia 14: 233 (1968) [“1967”].

Jlavida Tehler, Opera Bot. 70: 54 (1983) T

inaequabilis Tehler, Opera Bot. 70: 63 (1983) ¿A

pcs (Ny].) Tehler, Opera Bot. 70: 59 (1983).

Dirina limitata Nyl., Ann. Sci. Nat. Bot. sér 4, 3: 158 A E;

luteola Follmann, Nova Hedwigia 12: 239 (1968) [**1967”] T

lutosa (Zahlbr.) Tehler, Opera Bot. 70: 65 (1983).

Dirina lutosa Zahlbr., Acta Hoti Gothob. 2: 5 (1925) T.

mahuiana (Follmann) Tehler, Opera Bot. 70: 55 (1983),

Lobodirina mahuiana Follmann, Nova Hedwigia 14: 234 (1968) ESIOT IT.

Dirina condensata sensu Zahlb., Acta Horti Gothob. 2: 6 (1925)

nigricans Tehler, Opera Bot. 70: 72 (1983) T

nigrocincta Tehler, Opera Bot. 70: 61 ( 1983).

obscura Tehler, Opera Bot. 70: 69 (1983) T.

suffruticosa Tehler, Opera Bot. 70: 75 (1983) T.

terrestris Te ra Bot. 70: 71 (1983) T

Lit.: Tehler (1983); Quilhor et al. (1983); Follmann et al. (1993).

231. Roccellinastrum dore (1967)

[Lecanorales: ?Micareace

candidum (Mill. pri MO Nordic J. Bot. 2: 592 (1982).

Byssocaulon candidum Miill.-Arg., Nuovo Giorn. Bot. Ital. 21: 49 (1889).

Sagenidium candidum (Mill. Arg.) Follmamn, eos 2: 284 (1975)

ht Henssen £ Vobis. Nordic J. Bot. 2: 592 (19

¡oideum Follmann, Nova Hedwigia 14: a or

Lit: a (1968a); Follmann €: Crespo (1975); Henssen et al. (1982); Kantvilas (1990a); Follmann é: Schulz

(1993

* Roselliniella Vain. (1921).

[Sordariales: ai sed.]

coccocarpiae (Pat.) Matzer £ R.Sant. In M. Matzer $2 J. Hafellner, Biblioth. Lichenol. 37: 66 (1990).

LE: Mat Hafellner (1990).

32. Sagenidium Stirt. (1877)

pi Roccellaceae

niveum (Mont.) Follmann, Bot. Jahrb. Syst. 96: 49 (1975).

reas niveum Mont., Ann. Sci. Nat. Bot. 2,3: 355 (1835) T.

patagonicum Henssen, Lichenologist 11: 264 (1979

E: Polla (1975); Follmann $: Huneck (1976); Penis et al. (1979).

233. Santessonia Hale ba Vobis (1978)

[Lecanorales: Physciaceae]

cervicornis (Follimann) Follmann, Crypt. Bot. 5: (1995).

Roccella cervicornis Follmann, Ber. Deutsch. Bot. Ges. 79: 453 (1967) T.

Lie: es (1967, 1

234. Santessoniella Henssen (1997)

[Lecanorales: Pannariaceae int.

chilensis (Hue) Henssen, nal Bot. Upsal. 32 (1): 83 (1997).

Placynthium chilensis Hue, Bull. Soc. Linn. Normandie sér. 5, 9: 158 (1906) T.

elongata Henssen, Symb. Bot. Upsal. 32 (1): 83 (1997).

polychidioides (Zahlbr.) Henssen, Symb. Bot. Upsal. 32 (1): 77 (1997).

perl ea Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11):

Polychidium polychidioides (Zahlbr.) Henssen, Symb. Bot. Upsal. 18 (1): 105 (1963).

Lit.: Henssen (1997); Jórgensen (1998).

235. Schaereria Th.Fr. (1860)

[Ascomycota incert sed.: Sc

haereriaceae]

fuscocinerea (Nyl.) Clauzade € Cl. Roux, Bull. Soc. Bot. Centre Ouest, sér., N 7: 829 (1985

LL: Rambold (1989): Lunke er al. (1996). pa : "

160

Checklist of Chilean lichen: GALLOWAY, D.J. $ W. QUILHOT

236. Schismatomma Flot. € Kórb. ex A. Massal. (1852)

[Arthoniales: Roccellaceae]

accedens (Nyl.) Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3 (11): 327 (1924).

z ,T.

atacamensis C.W. Dodge, Nova Hedwigia 12: 327 (1967) [“1966”] T.

chilense C.W. Dodge, Nova Hedwigia 16: 489 sá ea JE.

epileucum (Nyl.) Zahlbr., Cat. Lich. Univ. 2: 556 (19

Jfuegiensis C.W. Dodge, Nova Hedwigia 12: 328 pul [“1966” PE

graphidioides (Leight.) Zahlbr., Cat. Lich. Univ. 2: 557 (1923).

gymnophorum (Nyl.) Zahlbr., Cat. Lich. Univ. 2: 558 (1923).

Lit.: Zahlbruckner (1924); Follmann (1965); Dodge (1967, 1969).

*Scutula Tul. (1852) nom. cons. prop.

[Lecanorales : era aa

allr.) Trevis., Spighe e Paglie: 10 (1853).

Peziza cnc Wallr., Fl. Crypt. Germ. 2: 499 (1833).

Lit.: Rambold « Triebel (1992); Triebel et al. (1997).

237. Siphula Fr. (1831) nom. cons.

[Lecanorales: incert. sed.

comata (Nyl. ex Cromb.) R. Sant. € Kantvilas, Herzogia 12: 14 (1996).

Siphula ramalinoides var. comata Nyl. In J. Crombie., Bot. J. Linn. Soc. 15: 226 (1876) T.

complanata (Hook. f. £ Taylor) R. Sant. In D.J. Galloway, New Zealand J. Bot. 21: 197 (1983).

Sphaerophoron complantum Hook. f. £ Taylor, London J. Bot. 3: 654 (1844) T

Siphula subcoriacea Miill. Arg., Miss. Sci. Cap Horn, Lich.: 151 (1888) T.

Siphula patagonica Vain., Résult. El is Lich.: 39 (1903) T.

pei Nyl., Lich. Nov. Zel.: 14 (188

noides Nyl. In J. Crombie, Bot. J. e n. Soc. 15: 226 (1876) T.

EJE: pd (1937); Galloway (1985): oobiinó (1987, 1994, 1996).

238. Siphulastrum coi e (1889)

pomos Pannariac

m (Hook. £ 4 Taylor) D.J. Galloway, New Zealand J. Bot. 21: 197 (1983).

¡E mamillata Hook. f. € Taylor, London J. Bot. 3: 637 (1

Parmeliella mamillata (Hook. f. £ Taylor) 2 Cat. Lich. Univ: 35212: 019235%

Lecidea adumbrans Ny]l., Lich. Fueg. Patag.:

Parmeliella adumbrans (Nyl.) Múll. Arg., AN HE 539 (1888).

Parmeliella adumbrans f. nigrata Miill. Arg., Nuovo Giorn. Bot. Ital. 21: 40 (1889).

PR P.M. Jórg., Lichenologist 00: 000 (1998) T

úill. Arg., Flora 72: 143 (1889).

Lit.: gesta (1985); Jóorgensen « Galloway (1992a); Jórgensen (1998).

*Skyttea Sherwood, D.Hawksw. € Coppins (1981)

[Ostropales: Odontotremataceae]

nitschkei (Kórb.) Sherwood, D. Hawksw, € Coppins, Trans. Brit. Mycol. Soc. 75: 488 (1981).

Lit.: Sherwood et al. (1981); Triebel (1989); Kondratyuk 4 Galloway (1995).

* Skyttella D. Hawksw. 8 R. Sant. (1988)

[Leotiales: incert sed.]

mulleri (Willey) D. Hawksw. £ R. Sant., Graphis Scripta 2 (2): 34 (1988).

Lit.: Hawksworth £ Santesson (1988).

9. Solorina Ach. (1808)

[Peltgeals. Peltigeraceae]

Spo. sa (Sm.) Anzi, Comment. Soc. Crittogam. Ital. 1: 136 (1862).

Lit.: Galloway (1998).

*Sphaerellothecium Zopf (1897)

[Dothideales: Mycosphaerellaceae]

161

Gayana Bot. 55(2), 1998.

minutum Hafellner, Herzogia 9: 760 (1993).

Lit.: Hafellner (1993c); Wedin (1994).

240. Sphaerophorus Pers (1794) nom. cons.

[Lecanorales: Sphaerophoraceae

globosus (Huds.) Vain., Résult. Voy. Belgica, Lich.: 35 (1903).

Sphaerophorus globiferus var. lacunosus Tuck., U.S. Expl. Exped. 1838-1842, 17. Bot. 1: 116 (1874)

[“1862”] T.

Sphaerophorus globosus var. lacunosus (Tuck.) Vain., Résult. Voy. Belgica, Lich.: 35 (1903).

Sphaerophoron globiferum var. versicolor Miill. Arg., Bot. Jahrb. Syst. 4: 145 (1888) T.

phaerophorus sae var. versicolor (Miill. Arg.) Zahlbr., Cat. Lich. Univ. 1: 693 (1922).

Lit.: Wedin (1993, 1995

241. Spilonema Bornet (1856)

eproniora Po quere ]

m Bornet, Mém. Soc. Imp. Sci. Nat. Cherbourg 4: 226 (1856).

NA Flora 48: 601 (1865).

Et: oa (0 1

*Spirographa Zahlbr. (1903)

[Ostropales: Odontotremataceae]

fusisporella (Nyl. ) Zahlbr. Cat. Lich. Univ. 2: 285 (1923).

Lit.: Santesson (1993).

242. Squamarina Poelt (1958)

[Lecanorales: Bacidiaceae

squamulosa (Nyl.) Follmamn, Philippia 2: 74 (1974).

Squamaria squamulosa Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 152 (1855) T.

Lit.: Follmann (1974); Timdal (1983): Feige et al. (1997).

243. Staurothele Norman (1853) nom. cons.

pipi Verrucariace: ae]

244. Stephanocyclos Hertel (1983)

¡sides Lecideaceae]

nssentanus Hertel., Lecid. Exs. Fasc. 5, No. 96 (1983).

Lit.: Hertel (1984 1987b).

245. Stereocaulon Hoffm. (1796) nom. cons.

[Lecanorales: Stereocaulaceae]

antarcticum Vain., Résult. Voy. Belgica, Lich.: 16 (1903) T.

corticatulum Nyl., Flora, Jena 41: 117 (1858).

corticatulum Nyl. var. humile (Mill. Arg.) LM. Lamb ex Frey, Bot. Jahrb. Syst. 86: 244 (1967).

corticatulum Nyl. var. subcorticatu; m (Rásánen) LM. Lamb, Canad. J. Bot. 29: 581 (1951).

Stereocaulon subcorticatum Rásinen, Ann. Bot. Soc. Zool. -Bot. Fenn. “Vanamo” 2 (1): 24 (1932) T.

LM. Lamb., J. Hattori Bot. Lab. 43: 290 (1977).

curtum (Rásánen) LM. Lamb, Can. J. Bot. 29 : 565 (1951).

Stereocaulon pi ito var. curta Rásánen, Revista Univ. (Santiago) 22: 200 (1937) T.

ltd

eocaulon ara var. recen Rásinen, pee Bol Soc. pul. -Bot. Fenn. “Vanamo” 2(1): 24 (1932).

Pilophoron colensoi var. reagens (Rásánen) Zahlbr., Cat. Lich. Univ. 10: 381 (1939).

Stereocaulon patagonicum f. reagens (Rásinen) LM M. Lamb, Anales Parques Nac. 7: 99 (1959) [**1958”].

162

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

Stereocaulon patagonicum f. subirregulare 1.M. Lamb. Farlowia 4: 456 (1955

glabrum (Mill. Arg.) Vain. f. flabellans (1.M. Lamb) LM. Lamb, J. Hattori Bot. Lab. ed 218 (1977).

Stereocaulon paschale var. alpinum f. flabellans, LM. Lamb, Farlowia 4: 457 (1955).

implexum Th.Fr., Stereoc. Piloph. Comm.: 23 (1857) T

Stereocaulon Peine var. implexum (Th.Fr. ) Nyl., Syn. Meth. Lich. 1 (2): 236 (1860).

Stereocaulon proximum var. compactius Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2

(Bot.) 3 (11): 373 (1924) 7

melanopotamicum 1.M. Lamb., J. Hattori Bot. Lab. 43: 296 (1977).

Stereocaulon corticatulum var. procerum 1.M. Lamb, Farlowia 4 : 462 (1955).

meyeri B. Stein, Jahrsb. Schles. Ges. Vaterl. Cult. 66: 134 (1888).

meyeri B. Stein var. farinosum (Th.Fr.) LM Porra Canad. J. Bot. 29: 582 (1951).

?microcarpum, Mill. Arg., Flora 62: 162 (1879

?obesum Th.Fr., Stereoc. Piloph. Comm.: 28 ed

ramulosum (Sw.) Ráuschel., Nomenclat. Bot. Ed. ds 328 (1797).

ramulosum (Sw.) Ráuschel, f. elegans Th.Fr., Stereoc. Piloph. Comm.: 11 (1857).

ramulosum f. subcompressum 1.M. Lamb, J. Hattori Bot. Lab. 43: 282 (1977).

ramulosum (Sw.) Ráuschel var. macrocarpum (A. Rich.) C. Bab. In Hook. f., Fl. Nov. Zel. 2: 294 (1 855).

ramulosum (Sw.) Ráuschel var. pulvinare (C.W. Dodge) LM. Lamb, J. Hattori Bot. Lab. 43: 283 (1977).

Stereocaulon pulvinare C.W. Dodge, B.A.N.Z.A.R.E. Rep. 7: 139 (1948).

ramulosum (Sw.) Ráuschel var. pulvinare (C.W. Dodge) LM. Lamb f. crebratum LM. Lamb, J. Hattori Bot.

Lab. 43: 283 (1977).

stenospermum (1.M. Lamb) LM. Lamb., J. Hattori Bot. Lab. 43: 304 (1977).

Stereocaulon argus var. stenospermum 1.M. Lamb, Farlowia 4: 460 (1955).

? strictum Th.Fr., Stereoc. Piloph. Comm.: 24 (1857).

tomentosum Fr., Sched. Crit. Lich. Exs. HI: 20 (1825).

Stereocaulon tomentosum ssp. magellanicum Th.Fr., Stereoc. Piloph. Comm.: 31 (1857).

Stereocaulon paschale var. magellanicum (Th.Fr.) Nyl., Mém. Soc. Sci. Nat. Cherbourg 5: 96 (1857).

Stereocaulon magellanicum (Th.Fr.) Zahlbr., Cat. Lich. Univ. 4: 651 (1927).

verruciferum Nyl., Sym. Meth. Lich. 1 (2): 248 (1860).

Stereocaulon speciosum 1.M. Lamb, Farlowia 4: 458 (1955).

verruciferum Ny]l. var. surreptans (IM. Lamb) 1.M. Lamb, J. Hattori Bot. Lab. 43: 256 (1977).

Stereocaulon speciosum Ny]. var. surreptans 1.M. Lamb, Farlowia 4: 459 (1955).

vesuvianum oi var. kilimandscharoense B. Stein., Jahrber. Schles. Ges. Vaterl. Cult. 66: 134 (1888).

vesuvianum var. efflorescens (Rásánen) LM. Lamb, Anales Parques Nac. 7: 100 (1959) [“1958”].

iia obesum var. efflorescens Rásinen, Anales. Soc. Ci. Argent. 128: 136 (1939).

Stereocaulon violascens Mill. Arg., Rev. Mycol. 1: 164 (1879

Stereocaulon vesuvianum var. violascens (Mill. Arg.) LM. Lamb, Anales Parques Nac. 7: 101 (1959)

58”

[

Lit.: Lamb (1977, 1978).

246. Sticta (Shreb.) Ach. (1803)

¿pora Stictaceae

D.J. Galloway 4 J. Pickering, Biblioth. Lichenol. 38: 92 (1990) T

caulescens De Not., Osserv. Sticta: 12 (1851)

Stictina caulescens (De Not.) Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 43 (1932).

Sticta biatora De Not., Osserv. Sticta: 13 (1851) T

Stictina filicina f. latissima Nyl. In J. Cromb., Bot. J. Linn. Soc. 15: 229 (1876) T.

Stictina caulescens f. pallidoglauca Rásánen, Revista Univ. (Santiago) 22: 202 (1937) T.

Stictina caulescens f. brunnea Rásánen, Revista Univ. (Santiago) 22: 303 (1937) T.

fuliginosa paren, Methodus: 280 (1803

gaudichaldia Delise, Mém. Soc. Linn. Normandie 2: 80, Tab. VIL fig. 23 (1825).

Sticta gaudichaldiana [sic.] Pers. In C. Gaudichaud-Beaupré, Voy. Uranie Bot.: 200 ( 1827).

Sticta filicina var. gaudichaudii (Delise) Mont., Ann. Sci. Nat. Bot. sér. 3, 18: ne a

Sticta freycinetii var gaudichaudii (Delise) Linds., Trans. Roy. Soc. Edinburgh 22 22: 197 (1859).

Stictina gaudichaudii (Delise) Nyl., Syn. Meth. Lich. 1 (2): 345 (1860).

Stictina quercizans var. gaudichaudii (Delise) ce Flora 48: 297 (1865).

Sticta malovina Fr., Syst. Orb. os E ss

Stic Vain., Hedwigia 38: 1

pon at Vain pinitos Wiss. Ergebn. poe Siidpolarexp. 1901-1903. 4 (11): 55 (1912).

Vain., Résult. Voy. Belgica, Lich.: 29 (190:

Ear fuliginosa f. pens a Bot. J. Pad o. 15: 229 (1876) T.

163

Gayana Bot. 55(2), 1998.

Stictina quercizans var. glaucovirens Jatta, Malpighia 20: 6 (1906) T.

Stictina filicinella var. ochraceofusca Rásánen, Anales Soc. Ci. Argent. 128: 143 (1939).

hypopsila (Mont.) Cromb., Bot. J. Linn. Soc. 16: 223 (1877).

Sticta filicina var. lineariloba f. hypopsila Mont., Ann. Sci. Nat. Bot. sér.3, 18: 308 (1852) T.

Sticta var. hypopsila (Mont.) Nyl., Amn. Sci. Nat. Bot. sér. 4, 11: 213 (1859).

best lineariloba var. hypopsila (Mont.) Nyl., Syn. Meth. Lich. 1 (2): 355 (1860).

icta laciniata var. denudata sensu Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.),

3 (11): 353 (1924) non. E e Meth. Lich. 1 (2): 354 (1860)].

limbata (Sm.) Ach., Methodus

lineariloba (Mont.) Nyl., Syn. Meth. y 1 (2): 355 (1860).

Sticta filicina var lineariloba Mont., Ann. Sci. Nat. Bot. sér. 3, 18: 308 (1852) T.

Lobaria lineariloba (Mont.) Trevis., Lich. Veneta Exs. No. 75 (1869).

longipes (Miill. Arg.) Malme, Bih. Kongl. Svenska Vetensk.-Akad. Handl. 25 (3/6): 11 (1899).

Stictina longipes Miúill. Arg., Flora, Jena 65: 303 (1882) T

Stictina filicinella var. pallescens Rásánen, Revista Univ. (Santiago) 22303 (1937WE.

Stictina filicinella var. roseola Rásánen, Revista Univ. (Santiago) 22: 303 (1937) T.

santessonii D.J. Galloway, Lichenologist 26: 268 (1994) T

sublimbata (Steiner) Swinscow $ Krog. In D.J. Galloway, New Zealand. J. Bot. 21: 198 (1983).

weigelii (Ach.) Vain., Acta Soc. Fauna Fl. Femn. 7: 189 (1890).

Lit.: Galloway (1994, 1995, 1997).

*Stigmidium Trevis (1860)

[Dothideales: Mycosphaerellaceae]

peltideae (Vain. ) R. Sant., Svensk Bot. Tidskr. 54 (4): 510 (1960).

pumilum (Lettau) Matzer 8: Hafellner, Biblioth. Lichenol. 37: 115 (1990).

Lit.: Santesson (1960, 1993); Hawksworth (1975); Matzer € Hafellner (1990).

247. Strigula Fr. (1823)

[Ascomycota incert. sed: Strigulaceae]

e Fr., Linnaea $5: 550 (1830).

rigula ee (Fée) Miill. Arg., Flora 63: 41 (1880).

1: bi (1952)

248. Tapellaria Miill. deL (1890)

[Lecanorales: Ectolechiaceae]

epiphylla (Mill. Arg. A R. Sant. In C.A. Thorold, J. Ecol.: 129 (1952).

Lit: serena (1952).

249. Teloschistes Norman (1853)

adigrer Teloschistaceae]

rysophthalmus (L.) Th.Fr., Gen. Heterolich. Eur.: 51 (1861).

Physcia chrysophthalma f. flavoalbida Kremp., Verhandl. Zool.-Bot. Ges. Wien 18: 322 (1868) T.

Teloschistes chrysophthalmus f. flavoalbida (Kremp.) Malme, Ark. Bot. 20A (3): 46 (1926).

vicans (Sw.) Norman, Nyt ue Naturvid. 7: 16 (1852).

deceo flavicans var. crocea f. maxima Meyen $: Flotow, Nov. Actorum Acad. Caes. Leop.-Carol.

t. Cur. 19 Suppl.: 210 (1843) T nom nud.].

ia flavicans var. maximus (Meyen $ Flotow) Zahlbr., Cat. Lich. Univ. 7: 323 (1931).

hosseusianis Gyeln., Borbásia Nova 9: 1 (194 mes

nodulifer (Nyl.) Hillm., Hedwigia 69: 326 (1930

Physcia nodulifera Nyl., Syn. Meth. Lich. 1 sa 407 (1860).

Borrera porn ont AS Flora 44: 52 (1861).

peruensis (Ach.) JW. Thomson. In Thomso: n « Itis, Bryologist 71: 32 (1968).

Borrera pus var. al Ach., Syn. Meth. Lich.: 224 (1814)

Physcia peruensis (Ach.) NyL., Syn., Meth. Lich. 1 (2): 407 rte

stellatus (Meyen $: Flotow) Miill. Arg., Jahrb. Kónigl. Bot. Gart. Berlin 2: 309 (1883).

Dr >. Meyen é Flotow, Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19 Suppl.: 210

Teloschisies llena Risioca. Revista Univ. (Santiago) 22: 201 (1937) T.

villosus (Ach.) Norman, Nyt Mag. Naturvidensk. 7: 17 (1 (1852).

Lit.: Hillmann (1930); Zahlbruckner (1931); Follmann (1968a); Thomson éz lltis (1968); Almborn (1989, 1992).

164

Checklist of Chilean lichen: GALLOWAY, D.J. € W. QUILHOT

250. Tephromela M. Choisy (1929)

[Lecanorales: Bacidiaceae

ds.) Hafellner. In K. Kalb, Lich. Neotrop. Exs. Fasc. 7: 15 No. 297 (1983).

Lit.: Rambold (1989); Poelt (1993); Poelt £ Grube (1993); Ekman (1996).

Thamnolecania (Vain.) Gyeln. (1933)

[Lecanorales: Bacidiaceae

fuegensis C.W. Dodge, Nova Hedwigia 12: 342 (1967) [**1966”] T.

Lit.: Dodge (1967); Ekman (1996).

252. Thamnolia Ach. ex Schaer. (1850) nom. cons.

[Lecanorales: incert sed.]

vermicularis (Sw.) Ach ex Schaer., Enum. Crit. Lich. Europ.: 243 (1850).

Thamnolia subuliformis (Ehrh.) W.L. Culb., Brittonia 15: 144 (1963).

nolia subvermicularis Asah., J. Jap. Bot. 13: 317 (1937).

Lit.: Kárnefelt € Thell (1995).

253. Thelenella Nyl. (1855)

[Ascomycota i incert. sed: Thelenellaceae]

-M. Lamb) O.E. Erikss., Opera Bot. 60: 96 (1981).

Microglaena antarctica 1.M. Lamb, Disc. Rep. 25: 24 (1948) T.

fernandeziana (Zahlbr.) H. Mayrhofer, Biblioth. Lichenol. 26: 33 (1987).

Microglaena fernandeziana Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3 (11):

318 (1924) T.

kerguelena (Nyl.) H. Mayrhofer, Biblioth. Lichenol. 26: 43 (1987).

Lit.: Lamb (1948); Mayrhofer (1987); Aptroot 8 van der Knapp (1993).

254, Thelidium A. Massal. (1855)

[Verrucariales: Verrucariaceae]

inaequale C.W. Dodge £: G.E. Baker, Ann. Missouri Bot. Gard. 25: 524 (1938).

Note: It is possible that the taxa; Porina (Segestria) chilena C.W. Dodge [Nova Hedwigia 12: 307 (1967) [“1966”]

T]; and Porina (Segestria) mahuana C.W. Dodge [Nova Hedwigia 16: 483 (1969) [“1968”] T]; are both referable

to en but not having seen the material we hesitate to namew them in Thelidium.

t.: Dodge 8: Baker (1938); Dodge (1967, 1969).

255. Thelocarpon Ny]l. (1853).

pipes Acarosporaceae]

reri (Flotow) Nyl., Mém. Soc. Nat. Sci. Cherbourg 3: 191 (1855).

Lit.: iron

256. Thelopsis Nyl. (1855) nom. cons.

ciao ?Stictidaceae]

: a Stizenb., Ber. St. Gallischen Naturw. Ges.: 262 (1895).

Ext: Egca 8 £ Torrente (1996).

257. Thelotrema Ach. (1803)

[Ostropales: Thelotremataceae]

lepadinum (Ach.) Ach., Meth. Lich.: 132 (1803).

osornense C.W. Dodge, Nova Hedwigia 12: 331 (1967) [”1966"] T.

subdenticulatum (Zahlbr.) G. Salisb., Lichenologist 5: 267 (1972).

laria subdenticulata Zahlbr. In C. Skottsberg, Nat. Hist. Juan Fernández Easter Is. 2 (11): 329 (1924) T.

Ocellu

Lit.: Zahlbruckner (1924); Salisbury (1972a, 1975, 1978), Galloway (1985).

258. Thyrea A. Massal. (1856)

[Lichinales: Lichinaceae]

pu Schaer.) A. Massal., Flora 39: 211 (1856

Lit.: Lamb (1959); Follmann (1965b); Henssen (1986), Moreno $: Egea (1991, 1992).

259. Toninia A. Massal. (1852) nom. cons. prop.

[Lecanorales: Lecanoraceae

Lobiona Kilias £ Gotth. Schneid., Lichenologist 10: 27 (1978).

auriculata Timdal, Opera Bot. 110: 44 (1991).

165

Gayana Bot. 55(2), 1998.

Lobiona albomarginata Kilias £« Gott. Schneid., Lichenologist 10: 27 (1978), non Toninia albomarginata

e Lesd.

australis Timdal, Opera Bot. 110: 45 (1991

bullata (Meyen £ Flot) Zahlbr., Beih. Bot. Teil, 19, 2:76 (1905).

Lecidea bullata Meyen £ Flot., Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 17, Suppl. 2: 227

1843).

inia badia Rásinen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 20, 3: 21 (1944) T.

?cinereovirens (Schaer.) A. Massal., Ric. Auton. Lich. Crost.: 107 (1852).

?hosseusiana Gyeln., Ann. Hist.-Nat. Mus. Natl. Pa 35 Bot.: 98 (1942).

sedifolia (Scop.) Timdal, Opera Bot. 110 (93 (1991

iospora (Nyl.) H. Olivier, Bull. Géogr. Bot. 21: 197 (1911).

Lit.: Timdal (1991); Jóorgensen (1996).

260. Tornabea Vsth. ai

[Lecanorales: Physci

scutellifera (With) ) UE R. Laundon, Lichenologist 16: 226 (1984).

Lichen scutelliferus With., A Botanical Arrangement of all the Vegetables naturally growing in Great Bri-

tain: 728 (1776).

Tornabea ephebea (Ach.) Kurok., J. Jap. Bot. 37: 293 (1962).

: Santesson (1942d); Kurokawa (1962); Huneck $ Follmann (1966); ODsthagen € Sunding (1980); Nimis $

Trad (1997).

261. Trapelia M. Choisy (1929)

dijorames ento

m.) M. Choisy. In R.G. Werner, Bull. Soc. Sci. Nat. Maroc 12: 160( 1932).

Eit.: Hola! np Aptroot $ van der Knapp (1993).

remella Pers (1794) nom. cons.

Pd Tremellaceae]

eliarum Diederich, Biblioth. Lichenol. 61: 125 (1996).

psoromicola Diederich, Biblioth. Lichenol. 61: 150 (1996).

Lit.: Diederich (1996).

262. Tremolecia M. Choisy (1953)

ape denon ron

atrata (Ach.) Hertel, Ergebn. Forsch. Unternehmens Nepal Himal. 6 2 351 (1977).

Lit.: erat Hertel (1984, 1987b, 1989); Jacobsen € Kappen (1988).

263. Tricharia Fée (1825)

[Ostropales: Gomphillaceae]

vulgaris (Mill. Arg.) R. Sant., Symb. Bot. Upsal. 5 oc

Lopadium ere Mill. Arg., Flora 64: 109 (18

Lit.: Santesson (1952).

264. Trichothelium Miill. Arg. (1885)

[Pyrenulaceae: Trichotheliaceae

epip: Mill. Arg., Bot. Jahrb. Syst. 6: 418 (1885).

Lit.: Santesson (1852).

*Trichothyrina (Petr.) Petr. (1950) = Lichenopeltella Hóhn. (1919)

265. Trimmatothelopsis era (1934)

[Verrucariales: ? Verrucariaceae]

antarctica C.W oder] Pp Hedwigia 15: Se ( iS [19681 T.

Lit.: Dodge (1969); Aptroot $ rece

266. Trypethelium Spreng. (1804) nom. cons.

deca Trypetheliaceae]

um (Eschw.) Zahlbr. In Engler-Prantl, Natiirl. Pflanzenfamil. 1 (1 70 (1

Lit.: patio Se

166

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

267. Tuckermannopsis Gyeln. (1933)

[Lecanorales: Parmeliaceae]

chlorophylla (Willd.) Hale. Bryologist 90: 164 (1987).

Ce traria eto (Willd.) Vain., Meddel. Soc. Fauna Fl. Fenn. 6: 121 (1881).

etr scutata var. pallida Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 16 (1932) T.

xt: Kairnefelt h al. (1992, 1993).

268. Turgidosculum Kohlm. 4 E. Kohlm. (1972)

[Ascomycyota, incert. sed.: Mastodiaceae]

complicatulum (Nyl.) Kohlm. $ E. Kohlm., Marine Mycology. The higher fungi: 361 (1979).

astodia tessellata auct.

Lit.: Galloway (1985); Aptroot €: van der Knapp (1993).

269. Umbilicaria Hoffm. (1798)

[Lecanorales: Umbilicariaceae]

africana (Jatta) Krog £ Swinscow, pesas: J. Bot. 6: 79 (1986).

aprina Nyl., Syn. Meth. Lich. 2: 12 (1863).

cristata C.W. Dodge £ Baker, Ann. Missouri. Bot. Gard. 25: 565 (1938).

decussata (Vill.) Zahlbr., Cat. Lich. Univ. 8: 490 (1942).

Umbilicaria eximia Hue, Deux. Exp. Antarct. Fr. 1908-1910, Sci. Nat. Doc. Sci.: 55 (1915) T.

Umbilicaria parvula Hue, Deux. Exp. Antarct. Fr. 1908-1910, Sci. Nat. Doc. Sci.: 56 (1915) T.

Umbilicaria leiocarpa var. nana Vain., Résult. Voy. Belgica, Lich.: 9 (1903) T

Agyrophora leiocarpa var. nana (Vain.) Llano, Monogr. Umbil.: 56 (1950).

diffissa (Nyl.) Llano, Monogr. Umbil.: 196 (1950).

Umbilicaria miihlenbergii (Ach.) var. diffissa Nyl., Ann. Sci. Nat. Bot. sér. 4,3: 149 (1855) T.

Umbilicaria dillenii Tuck. var. diffissa (Nyl.) Nyl., Syn. Meth. Lich. 2: 10 (1869).

Gyrophora diffissa (Nyl.) Nyl., Lich. Fueg. Patag.: 23 (1888).

haplocarpa Ny]l., Ann. Sci. Nat. Bot. sér. 4, 9: 217 (1859).

hyperborea (Ach.) Hoffm., Dtsch. Fl. 2: 110 (1796).

krascheninnikovii (Savicz) Zahlbr., Cat. eee ud 10: 405 (1939).

krempelhuberi Mill. Arg., Flora 72: 64 (188

nylanderiana (Zahlbr.) H. Magn., Lich. Sel. cl Exs. No. 252 (1937).

polyphylla (L.) Baumg., Fl. Lips.: 571 ( 1790

polyrrhiza (L.) Fr., Sched. Crit., Fasc. 5-6: 3 (1825).

propagulifera (Vain.) Llano, Monogr. Umbil.: 162 (1950).

Gyrophora cylindrica f. propagulifera Vain., Résult. Voy. Belgica, Lich.: 10 (1903) T.

Gyrophora propagulifera (Vain.) a Revist. Univ. (Santiago) 23: 195 (1937).

rufidula (Hue) Filson, Muelleria 6: 343 (19

Charcotia rufidula Hue, Bull. Soc. Bot. 24 sér. 4, 15: 17 (1915).

Umbilicaria antarctica Frey €£ LM. Lamb, Trans. Br. Mycol. Soc. 22: 270 (1939).

Umbilicaria dillenii var. solida Frey, Ber. Schweiz. Bot. Ges. 45: 217 (1936).

umbilicarioides (B.Stein) Krog £ Swinscow, Nordic J. Bot. 6: 83 (1986).

zahlbruckneri Frey, Ber. Schweiz. Bot. Ges. 45: 221 (1936)

Lit.: Llano (1950); Krog £ Swinscow (1986); Filson (1975, 1987); Swinscow £ Krog (1988); Quilhot et al.

(1991 a); Sancho et al. (1992).

270. Usnea Dill. ex Adans. (1763)

[Lecanorales: Usneaceae]

andina Motyka, Usnea 2 (1): 531 (1937).

aureola Motyka, Usnea 2 (1): 542 (1937) T.

chilensis Motyka, Usnea 2 (2): 578 (1938) T

eladoblephara (Miill. Arg.) Motyka, Usnea 2 (2): (1938

Usnea dasypogoides var. cladoblephara Miúill. Arg., E 69: 254 (1886) T

Usnea dasypoga var cladoblephara (Mill. Arg.) Zahlbr., Cat. Lich. Univ. 6: 556 (1930).

cladocarpa Fée, Essai Crypt. Ecorc.: XCVII (1824).

coccinea Dix, Bryologist 56: 277 (1953

columbiana Mont. In Rásánen, Revista Univ. (Santiago) 21: 138 (1936

comosa (Ach.) Róhl. var. chilensis Rásánen, Revista Univ. (Santiago) 21: 138 (1936) T.

comosa f. fuegica Rásánen, Ann. Bot .Soc. Zool.- Bot. Fenn. “Vanamo” 2 (1): 9 (1932) T.

comosa t. isidiella R Amn. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 9 (1932) T.

'omosella Ráisánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 9 (1932) T.

167

Gayana Bot. 55(2), 1998.

concreta Mont., Ann. Sci. de Bot. sér.3, 18: 302 (1852) T

condensata Mont. In Rásinen, Revista Univ. (Santiago) 21: 138 (1936) T.

costata Motyka, Usnea 2 Mm: -402 (1937) T

crassula Motyka, Usnea 2(2): 590 (1 938).

dasycera (Nyl.) Motyka, Usnea 2(1) 552 (1937).

Usnea florida var. dascyera Nyl. In Cromb., Bot. J. Linn. Soc. 15: 226 (1876) T.

denudata (Vain.) Rásiánen f. sorediella Rásiinen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 9 (1932) T.

durietzii Motyka, Usnea 2 (1): 503 ( 1937) T.

elongata Motyka, Usnea 2 (1): 411 (1937) T.

eulychniae Follmann, Nova Hedwigia 14: 261 Joe [”1967"] T.

eyerdami Herre, Rev. uba oa 29: 274 (196

eyerdami var. nigro-rubra e, Rev. Bryol. ao 29: 274 (1961) T.

Jastigiata Motyka, Usnea ao 580 (1938) T

Jlavocardia Risinen, Revista Univ. (Santiago) 21: 139 (1936) T.

fuegiana Motyka, Usnea 2 (1): 453 (1937) T

Jlexuosa Taylor, London J. Bot. 6: 192 (1847).

gaudichaudii Motyka, Usnea 2 (2): 600 (1938) T.

gigantea Vain. In Rásánen, Ann. Bot. Soc. Zool.-Bot. Fenn. “Vanamo” 2 (1): 7 (1932) T.

igniaria Motyka, Usnea 1: 108 (1936).

Jamaicensis Ach., Lich. Univ.: 619 (1810).

Jelskii Motyka, Usnea 2 (2): 595 (1938) T.

? kuehnemannii Motyka. In Lamb, dr 4: 467 (1955).

lacerata Motyka, Usnea 1: 76 (1936

maculosa Stirt, Scott. Nat. 7: 75 tra ¡E

meridionalis Zahlbr. , Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. KI. 83: 183 (1909).

miliaria Taylor, London J. Bot. 6: 192 (1847) T

ka, Usnea 2 (1): 512 (1937).

Ppusilla (Rásinen) Rásinen, Revista Univ. (Santiago) 22: 199 (1937

Usnea florida var. pusilla Rás e Revista Univ. (Santiago) 21: 139 (1936) T.

radiata Stirt., Scott. Nat. 6: 103 (188

rubicunda Stirt., Scott. Nat. 6: eo

scaberrima (Mont.) Motyka, Usnea 2 (2): 607 (1938).

Usnea ceratina var. scaberrima Mont. In C. Gay, Hist. Chile, Bot. 8: 63 (1852) T.

simplex Motyka, Usnea 2 (2): 582 (1938) T.

pee (Vain.) Motyka in de Lesd., Ann. pu Exot. 6: 121 (1933).

eri Zahlbr. var. tincta Zahlbr. , Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. KI. 83: 183 (1909).

reo (Vain.) Vain., Kongel. Danske pes Selsk. Naturvidensk. Math. Afh. 8(6): 392 (1924).

subtorulosa (Zahlbr.) Motyka, Usnea 2 (2): 603 (1938).

A sil subtorulosa Zahlbr. In E Skottsberg, Nat. Hist. Juan Fernández Easter Is]. 2 (Bot.), 3

a 92 (1924) T

sulcata Motyka, Usnea 2 ( 1): 478 (1937).

wirthii P. Clerc, Saussurea 15: 984).

Lit.: Zahlbruckner (1924, 1926); Motyka (1936-1938); Herre (1960); Clerc (1997).

271. Vainionora Kalb (1991)

[Lecanorales: Lecanoraceae

flavovirens (Fée) Kalb, Lich. N Neotrop. Fasc. 12: 3 (1991).

e. pierna Fée, Essai Crypt. Ecorc.: 115 (1825

nora aeruginosa Ny]l., Ann. Sci. Nat. Bot. sér. 4,3: 157 (1855) T.

Lit.: ro plo. te et al. (1996).

272. Vermilacinia Spjut (1995)

[Lecanorales: inaceae]

cephalota (Tuck.) Spjut 8: Hale, Flechten Follmann: 347 (1995).

168

Checklist of Chilean lichen: GALLOWAY, D.J. £ W. QUILHOT

Ramalina ceruchis f. cephalota Tuck., Syn. N. Am. Lich.: 21 (1882).

Desmazieria cephalota (Tuck.) Follmann £ Huneck, Willdenowia 5: 208 (1969).

Niebla cephalota (Tuck.) Rundel ££ Bowler, Mycotaxon 6: 498 (1978)

ceruchis (Ach.) Spjut € Hale, Flechten Follmann : 347 (1995

Parmelia ceruchis Ach, Meth. Lich.: 260 (1803).

1810).

Usnea ceruchis (Ach.) Mont., Ann. Sci. Nat. Bot. A > 368 (1834).

Desmazieria ceruchis (Ach.) Trevis., Flora 4: 52 (18

Niebla ceruchis (Ach.) Rundel $ Bowler, ebria 4 498 (1978).

Epa (Nyl.) Spjut € Hale, Flechten Follmann: 348 (1995).

na flaccescens Nyl., Bull. Soc. Linn. Normandie, sér. 2, 4: 109 (1870).

e. flaccescens (Nyl.) Follmann £ Huneck, Willdenowia 5: 208 (1969).

Niebla flaccescens (Nyl.) Rundel £ Bowler, Mycotaxon 6: 499 (1978)

tigrina (Follmann) Spjut € Hale, Flechten Follmann: 348 (199

Ramalina tigrina Follmann, Willdenowia 4: 227 (1966) T.

Desmazieria tigrina (Follmann) Follmann £ Huneck, Willdenowia 5: 208 (1969).

iebla tigrina (Follmann) Rundel £ Bowler, Mycotaxon 6: 498 (1978).

Ek: los (1966b); Follmann £ Huneck (1969); Spjut (1995, 1996).

273. Verrucaria Schrad. (1795) nom. cons.

[Verrucariales: Verrucariaceae

aethiobola Wahlenb. In Ach., Methodus Suppl.: 17 (1803).

brasiliensis Múll. Arg., Flora 67: 467 (1884).

ceuthocarpa Wahlenb. In Ach., Methodus: 22 (180

ceuthocarpa var. deformata Vain., Ark. Bot. 8 (4): a 909).

chiloénsis Zahlbr., Kongl. Svenska. e Handl. 57 (6): 4 (1917) T.

confinis Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 173 (1855) T

cylindrophora Vain., Résult. Voy. Belgica Lich.: 38 (1903).

dispartita Vain., Résult. Voy. Belgica, Lich.: 38 (1903).

durietzii 11M. Lamb, Lilloa 14: 205 (1948).

durietzii LM. Lamb f. rhabdota 1.M. Lamb, Lilloa 14: 206 (1948).

elaeoplaca Vain., Résult. Voy. Belgica, Lich.: 37 (1903).

famelica Darb., Wiss. Ergebn. Schwed. Siidpolarexp. 1901-1903, 4 (11): 18 (1912) T.

glaucoplaca Vain., Résult. Voy. Belgica, Lich.: 37 (1903).

? glaucoplacoides Darb., Wiss. Ergebn. Schweds, Siidpolarexp. 1901-1903, 4 (11): 18 (1912).

lacustris 1.M. Lamb, Farlowia 4: 424 (1955).

macrostoma Dufour Ex DC. In Lam. 4 DC., Fl. Fr. ed.3: 319 (1805).

pacas (Wahlenb.) Wahlenb, Fl. Lappon.: 465 (1812).

ahlenb. In Ach., Methodus: 19 (1803).

psico var. ecrustacea Nyl., Ann. Sci. Nat. Bot. sér. 4,3: 175 (1855) T.

pod halophila Nyl., Ann. Sci. Nat. Bot. sér. 4, 3: 175 (1855) T.

mucosa Wahlenb. In Ach., Methodus: 23 (1803).

muralis Ach., Methodus: 115 (1803

nigrescens Pers., Usteri Ann. Bot. 14: 36 (1795).

psychrophila 1M. Lamb, Discovery Rep. 18 (1948) T.

inctrina Ach., Syn. Meth. Apo 91 (1814

striatula ssp. australis R. Sant., Ark. Bot. 29 (10): 29 (1939).

tessellatula Ny!. In J. Cromb., 3 Bot. 13: 335 (18

tristis (A. Massal.) Kremp., Flora 40: 376 (185

umbrin . monospora Nyl., Ann. Sci. Nat Bot. sér. 4, 3: 175 (1855) T.

Lit.: Nylander (1855); Lamb (1948, 1959 1973); Follmann (1961); Redon (1985); Jacobsen $: Kappen (1988); Ap-

troot £ van der Knapp (1993).

*Vouauxiomyces Dyko $: D. Hawksw. (1979)

[Anamorphic Dothideales]

granulatae Wedin, Lichenologist 26: 306 (1994).

Lit.: Wedin (1994).

274. Wawea Henssen £ Kantvilas (1985)

[Lecanorales: Arctomiaceae]

Gayana Bot. 55(2), 1998.

? fruticulosa Henssen £: Kantvilas, Lichenologist 17: 86 (1985).

Lit.: Henssen € Kantvilas (1985).

275. Xanthomendoza S. Kondr. € Kiirnefelt (1997)

[Teloschistales: dias

mendoza ánen) S. Kondr. £ Kárnefelt, Biblioth. Lichenol. 68: 26 (1997)

Xanthoria an eiéd Rásánen, Anales Soc. Ci. Argent. 128: 140 (1939).

Lit.: Kondratyuk £ Kárnefelt (1997).

276. Xanthoparmelia (Vain.) Hale (1974)

[Lecanorales: Parmeliaceae]

australasica D.J. Galloway, N.Z. J. Bot. 18: e (1981).

conspersa (Ach.) Hale, Phytologia 28: 485 (1974).

cordillerana (Gyeln.) Hale, Phytologia 28: 487 (1974).

cotopaxiensis Nash, Elix £ J. Johnst., Mycotaxon 28: 288 (1987).

Jarinosa (Vain.) Nash, Elix 8 J. Johnst. ro a 286 (1987).

hypopsila (Mill. Arg.) Hale, Phytologia 28: 488 (197.

mexicana (Gyeln.) Hale, Phytologia 28: 488 (1974).

microspora (Miill. Arg.) Hale, Phytologia 28: 488 (1974).

mougeotii (Schaer.) Hale, Phytologia 28: 488 o :

scabroas (Taylor) Hale, Phytologia 28: 488 (1974).

skottsbergiana Nash £ Elix, Biblioth. Lichenol. 56: 98 (1995).

standaertii (Gyeln.) Hale, Phytologia 28: 489 (1974).

subramigera (Gyeln.) Hale, Phytologia 28: 489 (1974).

subsorediata Hale, Mycotaxon 30: 332 (1987).

taractica (Kremp.) Hale, Phytologia 28: 489 (1974).

tasmanica (Hook. f. £ Taylor) Hale, Phytologia 28: 489 (1974).

ulcerosa (Zahlbr.) Hale, Phytologia 28: 490 (1974).

Parmelia ulcerosa Zahlbr., Ann. Mycol. 29: 84 pit y

Parmelia tegulicola Zablbr., Ann. Mycol. 29: 85 (193

Parmelia conspersa var. obtecta Zahlbr. In C. Ace Nat. Hist. Juan Fernández Easter Isl. 2 (Bot.), 3

(11): 386 (1924) T

vagans (Nyl.) Hale, Phytologia 28: 490 (1974).

wildeae (C.W. Dodge) Hale, Phytologia 28: 490 (1974

xanthomeleana (Miill. Arg.) Hale, Mycotaxon 30: 334 ( 1987).

Lit.: Hale (1990); Elix (1993, 1994p); Nash et al. (1995).

277. Xanthopeltis R. Sant. (1949)

[Teloschistales: ego

ico! ., Sy. Bot. Tidsk. 43: 560 (1949) T

Lt: Santesson: 19 Huneck £ Follmann (1971); Fúllmana (1973).

278. Xanthoria (Fr.) Th.Fr. (1860) nom. cons.

[Teloschistales: Teloschistaceae]

ascendens S. Kondratyuk, Rms 29: 431 (1997).

candelaria (L.) Th.Er. Gen. Heterol. Eur. e 61 (1861).

elegans (Link) Th.Fr., Lich. pei 69 (18

?gallowayii S. Kondr. £ Kárnefelt, a Lichenol. 68: 28 (1997).

parietina (L.) Th.Fr. Lich. Arct.: 69 (18 60).

polycarpa (Hoffm.) Th.Fr. ex Rieber, Jahresh. Vereins Vaterl. Naturk. Wiirttemberg 42: 252 (1891).

Xanthoria polycarpa f. santiagoénsis Rásiinen, Revista Univ. ( Santiago) 21: 143 (1936) T.

Xanthoria santiagoénsis (Rásánen) C.W. Dodge. In Follmann, Nova Hedwigia 14: 267 (1969) [1968”].

bp 2. o gee poo ( 97; Redon (1985): Jacobsen é Kappen (1988); Kárnefelt (1989); Poelt

g ptroot £ Van der Kn. 1993); Kondr.

Lino (O ira Ec Sis app ( ); atyuk (1997); Kondratyuk é: Poelt (1997);

279. Xenolecia Hertel éter

[Lecanorales: Porpidiaceae

PEPUIRS (NyL) a Beih. Nova Hedwigia 79: 440 (1984).

Lecidea spadicomma Nyl. In Cromb.. Bot. J. Linn. Su do 233 (1876) T.

P PROS ero (Nyl.) Darb., Wiss. Ergebn. Sc - Súdpolarexped. 1901-1903, 411): 49 (1912).

170

Checklist of Chilean lichen: GALLOWAY, D.J. € W. QUILHOT

280. Xylographa (Fr.) po (1836)

Laponia Agyriace

usta (Stirt.) Mall Arg., Bull. Herb. Boissier 2, App. 1: 76 (1894).

perangus

Lit.: Follmann (1962); Galloway (1985).

281. puro Herre (1912)

[Lichinales: Lichinace

penis Poio Symb. Bot. Upsal 18: 80 (1963) T.

patagonica Henssen, Lichenologist 9: 41 (1977).

Lats Henssen (19630, 1977).

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Fecha de publicación: 07 de mayo de 1999

185

Gayana Bot. 55(2): 187-211, 1998.

ISSN 0016-5301

CATALOGUE OF THE VASCULAR FLORA OF THE

ROBINSON CRUSOE OR JUAN FERNANDEZ ISLANDS, CHILE

CATALOGO DE LA FLORA VASCULAR DE LAS ISLAS

ROBINSON CRUSOE O JUAN FERNANDEZ, CHILE

Clodomiro Marticorena', Tod F. Stuessy?, and Carlos M. Baeza'

ABSTRACT

The Róbinson Crusoe (=Juan Fernández) Islands are a

Chilean National Park and a World Biosphere Reserve

ated 667 kms W of continental Chile at 339S latitude,

the archipelago consist of three principal islands: Mas

atierra Glsla Róbinson Crusoe; 48 km?), MA

Ace and have extended to the present time. Numerous

of the archipelago. Interest in conservation of these en-

demic taxa is also increasing. It is timely, therefore, for

the production of a new flora of vascular plants of the

islands, and this catalogue is a necessary step towards

that goal. Herein are documented 423 species of vascu-

lar plants, including 55 Pteridophyta, 289 Dicotyledon-

eae, and 79 Monocotyledoneae. Of these, 31.2% are

endemic, 18.7% native, and 50.1% introduced. Of ad-

ventive taxa, the highest level if found on Santa Clara

(71.4%), followed by Masatierra (55.7%), and lastly by

Masafuera (50%)

KEYWORDS: Juan Fernández/Róbinson Crusoe Islands,

flora, endemism, conservation.

INTRODUCCION

The Róbinson Crusoe (=Juan Fernández) 1s-

lands are located in the Pacific Ocean at 33? S la-

titude 667 kms W of continental Chile. There are

two principal islands, Isla Róbinson Crusoe

(=Masatierra) and 181 kms further west Isla Ale-

'Departamento de Botánica, Universidad de Concep-

ción, Casilla 160-C, Concepción,

“Department of Higher Plant Systematics and Evolu-

tion, Institute of Botany, University of Vienna, Vienna,

Austria.

RESUMEN

E Islas repni Pe (=Juan Fernández), Chile,

onstituye: arque nacional y una reser-

va mu do de la healer SAO a 667 km al oeste de

Chile continental, el archipiélago consta de tres islas:

Masatierra (=Róbinson Crusoe), de 48 km”, Masafuera

(Alejandro Selkirk), de 50 km”, y Santa Clara, de 2

km. Los inventarios botánicos del archipiélago empe-

zaron en 1823 y han continuado hasta el presente. Du-

rante las recientes décadas se han hecho numerosas

y patro:

divergencia evolutiva entre los taxa endémicos papas plan-

tas vasculares del archipiélago. También se ha acrecen-

tado el interés en la conservación de estos taxa

endémicos. Es tiempo, entonces, de producir una nueva

flora de las plantas vasculares de las islas, y este catálo-

go es un paso necesario hacia esa meta. En él están do-

cumentadas 423 especies de plantas vasculares,

incluyendo 55 Pteridophyta, 289 Dicotyledoneae y 79

Monocotyledoneae. Del total de especies, 31,2% son

endémicas, 18,7% nativas y 50,1% adventicias. De es-

tos últimos taxa el mayor porcentaje se encuentra en

Santa Clara (71,4%), seguido por Masatierra (55,7%), y

finalmente por Masafuera (

PALABRAS CLAVES:Juan OS Crusoe,

flora, endemismo, conservac

jandro Selkirk (=Masafuera). The former names

have been officially adopted by the Chilean

government, but the latter have been used histori-

cally and still continue to be used in much scien-

tific literature (e.g., Stuessy and Ono, 1998).

There is also a third smaller island, Isla Santa

Clara, that lies 1.8 km SW off the western end of

Masatierra. These three islands together measure

only 102 km', with Masafuera being the largest at

50 km?, Masatierra nearly the same size at 48 km",

and Santa Clara with 2 km? (Stuessy, 1995).

In this small Pacific archipelago is contained

one of the world's most interesting endemic vas-

cular floras. The distinctiveness of the vascular

187

Gayana Bot. 55(2), 1998.

plants is the main reason for designation of the

three islands as a Chilean National Park in 1935

and more recently as a World Biosphere Reserve.

There are many endemic species, several endemic

genera, and an endemic family, Lactoridaceae, the

only case known of a entire family of vascular

plants being restricted to an oceanic island (Stuessy

et al., 1998a, 1998d).

During the past 20 years, studies on the evo-

lution of the vascular flora of the Róbinson Crusoe

Islands have been carried out by collaborators at

the University of Concepcion, The Ohio State

University, and now at the University of Vienna.

These investigations have resulted in more than 50

publications on different aspects of the endemic

taxa, including cytology, phylogeny, modes of

speciation, phytochemistry, biogeography, mole-

cular divergence, and genetic variation within and

between populations (for recent summaries see

Stuessy et al. 1998b, 1998c; Crawford et al.,

1998). Studies have also focused on the introdu-

ced flora and impacts on conservation of the nati-

ve taxa (Matthei et al., 1993; Stuessy et al., 1997,

1998d; Swenson et al., 1997).

These previous collaborative investigations

on the vascular taxa have led to the initiation of a

synthesis of all available information to be pub-

lished as a book entitled “Flora of the Róbinson

Crusoe Islands”. Previous descriptive inventories

include those of Hemsley (1884), Johow (1896),

Christensen and Skottsberg (1920), and Skottsberg

(1921, 1951). Additional information has been ad-

ded by others, such as Ricci (1996), Macaya and

Faúndez (1997) and Barrera (1997). The only

comprehensive flora is that of Johow (1896), now

more than a century old, and lacking keys to taxa.

For improved perspectives on conservation of the

native and endemic species, plus management of

the introduced taxa, a new comprehensive flora is

urgently needed.

The present catalogue of the vascular flora is

the first step toward publication of the new flora

of the archipelago. Contents of the catalogue

derive from previous literature reports, our own

collections made during eight expeditions to the

islands during 1980-1997, and additional collec-

tions in the herbarium of the University of Con-

cepcion (CONC). We harbor no illusion that this

list is complete nor fully accurate in all respects. It

188

is a stage in preparation of the flora that should

stimulate preparation of individual generic treat-

ments and serve as an impetus for completion of

the entire volume. In fact, at this time manuscripts

for the ferns (R. Rodríguez), grasses (M. Baeza),

and miscellaneous other genera have already been

prepared.

We list all known species of vascular plants

that have at one time or another been reported to

occur in at least one of the three islands. We

include also distribution on the three principal is-

lands (T=Masatierra; F=Masafuera; S=Santa Cla-

ra), and biogeographic status (E=endemic;

N=native; A= adventive). This list builds, ob-

viously, on the catalogue of vascular plants for the

entire country (Marticorena and Quezada, 1985),

which includes the island floras of Róbinson Cru-

soe, Islas Desventuradas, and Isla de Pascua.

The catalogue does not include cultivated

plants. Swenson et al. (1997) listed species that

eventually may escape from cultivation and have

the potential to become aggressive weeds, but we

have not included these here. Previous information

regarding cultivated plants in the archipelago also

can be found in Johow (1893).

The catalogue also provides the opportunity

to summarize new statistics on the vascular flora

of the Róbinson Crusoe Islands. In our own studies

we have found it difficult to maintain agreement on

numbers of taxa of different biogeographic cate-

gories, due largely to numerous publications using

different sets of data. For the first time, this new

catalogue sets baseline data for any additional

evolutionary or biogeographic investigations.

NEW STATISTICS OF THE VASCULAR FLORA

All manner of statistics on the composition of

the vascular flora have been published previously in

Johow (1896), Skottsberg (1956), Marticorena

(1991), Stuessy et al. (1992), etc. This paper is not

the place to compare all different previous estimates,

but rather the intent is to provide a new baseline of

the total size and general categorization of the vas-

cular flora. Statistics for the entire archipelago and

for each of the three islands separately are given in

the following tables.

Vascular flora of the Róbinson Crusoe Islands: MARTICORENA, C. ET AL.

MASATIERRA

PTERI | DICOT | MONOC | Total %

E 21 64 10 95 1275

N 21 16 21 58 1163.

A 0 155 7 19 | 557

Total| 42 68 345 |

% 122 19 |

SJ |

345 taxa, of which 177 (51.3%) are restricted to the island.

MASAFUERA

PTERI | DICOT | MONOC | Total %

E 19 37 8 64 1271]

N 27 15 12 22.9

A 0 95 23 118 150.0

Total | 46 147 - A e E

% 195 A

236 taxa, of which 78 (33%) are restricted to the island.

SANTA CLARA

PTERI | DICOT | MONOC | Total %

E 0 4 (3 AOS E 11.4 |

N 2 A A io JO!

0 22 A A!

Total 3 Ed: OR Y AS os Ly

% 86 80 114

35 taxa, of which 1 (2.9%) is restricted to the island.

FLORA OF THE ARCHIPELAGO

' PTER | DICOT | MONOC | Total | %

26 LE A

N MIA LO

A 01 M4 (22 ML

o 5 1 9 | WI 14M -

% 1d SS 18,7 |

ACKNOWLEDGMENTS

Partially supported by Oesterreichische Na-

tionalbank Jubiláumsfondsprojekt Nr. 6880. Spe-

cial thanks go to Gerhard Jakubowsky, Vienna,

for locating and copying needed literature and

1 . W give thanks to the CONAF

guides of the Robinson Crusoe National Park,

who have always accompanied and helped us in

the field, and who have alerted us to interesting

variations in the native and introduced flora.

LITERATURE CITED

BARRERA, E. 1997. Helechos de Juan Fernández. Publ.

as. Mus. Nac. Hist. Nat. Chile 51: 1-104

CHRISTENSEN, C. AND C. SKOTTSBERG. 1920. The Pte-

ridophyta of the Juan Fernández Islands, pp. 1-46.

ln €: Skottsberg (ed.), The Natural History of

Juan Fernández and Easter Islands, vol. 2, Botany.

Almgyvist € Wiksells: Uppsala.

Eesti D.J., T. SANG, T.F. STUESSY, S.-C. KIM

D M. SILVA. 1998. Deiiraseris Úlilbnibcos:

pera and Robinsonia (Asteraceae: Senecio-

neae) on the Juan Fernández Islands: similarities

and differences in biology and phylogeny, pp. 97-

119. In T. F. Stuessy and M. Ono (eds.), Evolution

and Speciation of Island Plants. Cambridge Univ.

Press: Cambridge.

HemsLEY, W.B. 1884. Report on the botany of Juan

Fernández, the south-eastern Moluccas and the

Admiralty Islands, pp. 1-96. In C.W. Thomson

and J. Murray (eds.), Report on the Scientific Re-

sults of the Voyage of HMS Challenger During

the Years 1873-76, vol. 1, part 3, Botany. HMSO:

JOHOW, F. 1893. Las plantas de cultivo en Juan Fernán-

dez. Anales Univ. Chile 84: 939-970.

JoHow, F. 1896. Estudios sobre la Flora de las Islas de

Poaceae para el archipiélago de Juan Fernández y

nuevas localidades para cuatro especies adventi-

cias. Notic. Mens. Mus. Nac. Hist. Nat. 327: 13-15.

MARTICORENA, C. 1991. Contribución a la estadística

de la flora vascular de Chile. Gayana, Bot. 47: 85-

13.

MARTICOREN D M. QUEZADA. 1985. Catálogo de

la flora apt Gayana, Bot. 42: 1-157.

MATTHEL O., C. MARTICORENA AND T.F. STUESSY.

1993. La flora adventicia del archipiélago de Juan

rmández. Gayana Bot. 50: 69-102

Ricci, M. 1996. Variation in distribution and abun-

dance of the endemic flora of Juan Fernández Is-

lands, Chile. Pteridophyta. Biodivers. Conserv.

5 1521-1532.

SKOTTSBERG, C. 1921. The phanerogams of the Juan

Fernández Islands, pp. 95-240. In C. Skottsberg

(ed.), The Natural History of Juan Fernández and

Easter Island, vol. 2, Botany. Almqyvist € Wik-

sells:

SKOTTSBERG, C. 1951. A supplement to the pterido-

phytes and phanerogams of the Juan Fe rmández an

Easter Islands, pp. 763-792. In C. Skottsberg (ed.),

The Natural History of Juan ape and Easter

Island, vol.2., Botany. Almqyist €: Wiksells: Uppsala.

SKOTTSBERG, C. 1956. Derivation of the flora and fauna

of Juan Fernández and Easter Island, pp. 193-439.

189

Gayana Bot. 55(2), 1998.

In C. Skottsberg (ed.), The Natural History of Juan

Fernández and Easter Island, vol. 1. Almgyvist $

Wiksells: Uppsala.

STUESSY, T. F. 1995. Juan Fernández Islands-Chile, pp.

565-568. In S.D. Davis, V.H. Heywood and A.C.

Hamilton (eds.), Centres of Plant Diversity: A

Guide and Strategy for Their Conservation. WWF,

IUCN. Cam pepa

STUESSY, T.F., D.J. CRA D, G. ANDERSON AND R.

JENSEN. 1998a. Systematic, ada and

conservation of Lactori erspectives in Pl.

Ecol. Evol. Syst. 1(2): 267. 290.

STUESSY, T.F., D.J. CRAWFORD, C. MARTICORENA AND

R. RODRIGUEZ 1998b. Island biogeography of an-

lution and Speciation of Island Plants. Cambridge

Univ. Press, Cambridge

STUESSY, T.E...D. 3 CRAWFORD, C. MARTICORENA

AND M. SILVA 1998c. Isolating mechanisms and

modes of speciation in endemic angiosperms of

the Juan dora Islands, pp. 79-96. In T. F.

Stuessy and M. Ono (eds.), Evolution and Specia

tion of Island Plants. Cambridge Univ. e

Cambridge.

STUESSY, T.F., C. MARTICORENA, R. RODRIGUEZ, DI

CRAWFORD, AND M. SILVA. 1992. Endemism in

the vascular flora of the Juan Fernández Islands.

Aliso 13: 297-307

STUESSY, T.F. AND M. ONO (eds.). 1998. Evolution and

Speciation of Island Plants. Cambridge Univ.

Press: Cambridge.

STUESSY, T. F., U. SWENSON, D.J. CRAWFORD, G. AN-

DERSON AND M. SILVA O. 1997. Plant conserva-

tion in the Juan Fernández archipelago, Chile.

Aliso 16: 89-102

STUESSY, T. F., U. SWENSON, C. MARTICORENA, O.

MATTHEI, AND D. J. CRAWFORD. 1998d. Loss of

plant diversity and extinction on Robinson Crusoe

Islands, Chile, pp. 243-257. In C.-I. Peng and P. P.

Lowry II (eds.), Rare, Threatened, and Endangered

Floras of Asia and the Pacific Rim. Institute of

Botany, Academia Sinica Monograph Series No.

A L

SWENSON, U., T.F. STUESSY, M. BAEZA AND D.J.

me ORD. 1997. New and historical plant intro-

ductions, and potential pests in the Juan Fernández

Islands, Chile. Pacific Sci. 51: 233-253.

Vascular flora of the Róbinson Crusoe Islands: MARTICORENA, C. ET AL.

FLORA OF JUAN FERNANDEZ ISLANDS

Col. 1, Status: E = endemic, e = endemic to JF and continental Chile, N = native, A = adventive/

natural ized.

Col. 2, Life form: A =:annual, B = biennial, H = perennial herb, S = undershrub, F = shrub, T = tree,

R = rosette shrub/tree

Col. 3, Distribution: T = Masatierra, S = Santa Clara, F = Masafuera

PTERIDOPHYTA

ADIANTACEAE

ADIANTUM

NH TSF chilense Kaulf., Enum. Filic. 207. 1824.

NOTHOLAENA

EH TF chilensis (Fée ex J.Remy) Sturm, Abh. Naturhist. Ges. Núrnberg 1(2): 166. 1858.

("Nothochlaena")

ASPLENIACEAE

NH TF dareoides Desv., Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk.

SU SE. TB EL.

EH TF macrosorum Bertero ex Colla, Herb. Pedem. 6: 205. 1836.

NH TSF obtusatum G.Forst. var. sphenoides (Kunze) C.Chr. ex Skottsb., Kongl. Svenska

Vetenskapsakad. Handl. 56(5): 167. 1916.

EH TF stellatum Colla, Herb. Pedem. 6: 206. 1836.

BLECHNACEAE

BLECHNUM

NH TF cordatum (Desv.) Hieron., Hedwigia 47: 239. 1908.

EH TF cycadifolium (Colla) Sturm, Abh. Naturhist. Ges. Núrnberg 1(2): 173. 1858.

NH TSF hastatum Kaulf., Enum. Filic. 161. 1824.

EH F longicauda C.Chr., Ark. Bot. 10(2): 10, t. 1. 1910.

EH TF mochaenum G.Kunkel var. fernandezianum (Looser) De la Sota, Bol. Soc. Argent. Bot.

14(3): 196.. 1972.

schottii (Colla) C:Chr:,' Ark. Bot. 10(2): 7. 1910.

—

DENNSTAEDTIACEAE

HISTIOPTERIS

NU. TF. incisa (hunb.) 1%. Hist. FIL 299. 1875.

HYPOLEPIS E

NH TF poeppigii (Kunze) R.A.Rodr., Gayana, Bot. 46(3-4): 202. 1990. ("1989")

DICKSONIACEAE

DICKSONIA

1 : bb.

EF 7 perrecoma Ccotla) Hook, Sp. FM. 87, E. 2 os

ETE extema Skottsb., Mar. Mist. Juan Fernandez 2: 763. ;

ese in Skottsb., Nat. Hist. Juan Fernandez 2: 16. 1920.

N HF TF quadripinnata (J.F.Gmel.) C.Chr.,

191

Gayana Bot. 55(2), 1998.

THYRSOPTERIS (E)

elegans Kunze, Linnaea 9: 507. 1835.

—

DRYOPTERIDACEAE

MEGALASTRUM

END 04 imaequalifolium (Colla) A.R.Sm. et R.C.Moran var. data (C.Chr. et Skottsb.)

R.A.Rodr., Gayana, Bot. 46(3-4): 206. 1990. (19891)

EH T - inaequalifolium (Colla) A.R.Sm. et R.C.Moran var. imaequalifolium, Amer. Fern y.

77(4): 128. 1987.

POLYSTICHUM

tetragonum Fee, Mém. Foug. 8: 99. 1857.

pan]

RUMOHRA

EH TF berteroana (Colla) R.A.Rodr., Bol. Soc. Biol. Concepción 45: 150. 1972

GLEICHENIACEAE

GLEICHENIA

EH F. lepidota R.A.Rodr., Gayana, Bot. 47(1-2): 40, f. 1-5. 1990.

NH F quadripartita (Poir.) T.Moo oore, Index Fil. 382. 1862.

NH T squamulosa (Desv.) T.Moore var. squamulosa, Index Fil. 383. 1862.

GRAMMITIDACEAE

GRAMMITIS

.magellanica Desv., Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten

Naturk. 5(2): 313... 1811:

—

HYMENOPHYLLACEAE

HYMENOGLOSSUM (e)

eH TF cruentum (Cav.) K.Presl, Hymenophyllaceae 35. 1843.

HYMENOPHY LLUM

Nh TF caudiculatum Mart. var. productum (K.Presl) C. Chr., Index Filic. 623. 1906.

NH TF. cuneatum Kunze var. cuneatum, Analecta Pteridogr. 50. 1

E H T— cuneatum Kunze var. rariforme C.Chr. et Skottsb

Fernandez 2: 8, f. 3 e-g. 1920.

F —— falklandicum Baker var. falklandicum, in Hook. et Baker, Syn. Fil

TF. ferrugineum Colla var. ferrugineum, Herb. Pedem. 6: 190.

TF fuciforme Sw., Sym. Fil: E 1806.

pectinatum Cav., Descr. Pl. 275. 1801.

TF plicatum Kaulf., Enum. Ped 268. 1824.

TF rugosum C.Chr. et Skottsb., in Skottsb

192

LO Skotrab. : Nat. Hist. Juan

. ed. 2, 68. 1874.

MEZZZZ

TITTT

—

—m

-, Nat. Hist. Juan Fernandez 2: A EA

NH F secunmdum Hook. et Grev., Icom. Filic. 2: t. 133. 1829.

NH F tortuosum Hook. et Grev. var. tortuosum, Icom. Filic. 2: e. 129. 1829.

SERPYLLOPSIS

caespitosa (Gaudich.) C.Chr. var. fernmandeziana C.Chr. et Skottsb., in Skottsb

Hist. Juan Fernandez 2: 5. 1920.

-. Nat.

192

Vascular flora of the Róbinson Crusoe Islands: MARTICORENA, C. ET AL.

—_

—

—Ú

“nm

+ —

" .

TRICHOMANES

exsectum Kunze, Analecta Pteridogr. 47, t. 29, f. 2. 1837.

imgae C.Chr. et Skottsb., in Skottsb., Nat. Hist. Juan Fernandez 2: E

philippianum Sturm, Abh. Naturhist. Ges. Núrnberg 1(2): 188. 1858.

LOMARIOPSIDACEAE

ELAPHOGLOSSUM

squamatum (SwW.) T.Moore, Index Fil. 15, 367. 1857.

LYCOPOD IACEAE

LYCOPOD IUM

gayanum J.Remy, in Gay, Hist. Chile, Bot. 6: 545. 1854.

magellanicum (P.Beauv.) Sw. var. magellanicum, Syn. Fil. 180. 1806.

OLEANDRACEAE

ARTHROPTERIS

altescandens (Colla) J.Sm., Hist. Fil. 225. 1875.

OPHIOGLOSSACEAE

OPHIOGLOSSUM

fernandezianum C.Chr., in Skottsb., Nat. Hist. Juan Fernandez 2: 44, f. 7. 1920.

POLYPOD IACEAE

PLEOPELTIS

macrocarpa (Bory ex Willd.) Kaulf.,

POLYPOD IUM

intermedium Colla subsp. intermedium, Herb. Pedem. 6: 229. 1836.

intermedium Colla subsp. masafueranum-C.Chr. et Skottsb., in Skottsb., Nat. Hist.

Juan Fernandez 2: 38, f. 3

masafuerae Phil., Linnaea 29: 107. 1858.

Berlin. Jahrb. Pharm. Verbundenen Wiss. 21: 41.

PTERIDACEAE

PTERIS

berteroana J.Agardh, Recens. Spec. Pter. 66. 1839.

chilensis Desv., Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk.

semiadnata Phil., Linnaea 29: 106. 1858.

WOODS IACEAE

CYSTOPTERIS

fragilis (L.) Bernh. var. apiiformis (Gand.) C.Chr., Index Filic. Suppl. Prelim. 11.

193

Gayana Bot. 55(2), 1998.

DICOTYLEDONEAE

AIZOACEAE

TETRAGONIA

AA TSF tetragonoides (Pall.) Kuntze, Revis. Gen. Pl. 1: 264. 1891.

AMARANTHACEAE

AMARANTHUS

deflexus L., Mant. Pl. 295. 1771.

AGA: :T... hybridus;L...,. Sp. PL. 990... ¿1733

APOCYNACEAE

VINCA

AH. 7 omajeris, Sb-+PL. 20%: 1/03

BERBERIDACEAE

BERBERIS

E FT T corymbosa Hook. et Arn., Bot. Misc. 3: 135. 1832.

E FT F masafuerana Skottsb., Nat. Hist. Juan Fernandez 2: 125, f. 9 a-b. 1921.

BORAGINACEAE

CYNOGLOSSUM

AB T creticumMill., Gard. Dict., ed. 8, Cynoglossum 3. 1768.

MYOSOTIS

A AH T laxa Lehm., Pl. Asperif. Nucif. 83. 1818.

AH T sylvatica Hoffm., Deutschl. Fl. 61. 1791.

SELKIRKIA (E)

E3F. T berteroi (Colla) Hemsl., Rep. Challenger, Bot. 1(3): 48, t. 57. 1884.

CALLITRICHACEAE

CALLITRICHE

AA T lechleri (Hegelm.) Fassett, Rhodora 53(632): 191, t. 1174, f. 18 a-m. 1951.

CAMPANULACEAE

LOBEL IA

NA TF alata Labill. var. alata, Nov. Holl. Pl. 1: 51, t. 72. 1805.

SO tipa La, "Sp PE ed 2 ITA

WAHLENBERGIA

ES TS berteroi Hook. et Arn., J. Bot. (Hooker) 1: 2er, -€.. 157.. 1834.

ESSF: T fernandeziana A.DC., Monogr. Campan. 160. 1830.

ES T' grahamiae Hemsl., Rep. Challenger, Bot. 1(3): 66, t..36, Y. 3-10. 1 e

ES F masafuerae (Phil.) Skottsb., Kongl. Svenska Vetenskapsakad. Handl. 51(9): 6. 19164.

ES F tuberosa Hook.f., Bot. Mag. 101: 6155. 1875.

Vascular flora of the Róbinson Crusoe Islands: MARTICORENA, C. ET AL.

CARYOPHYLLACEAE

CERAST IUM

AH F fontanum Baumg. subsp. vulgare (Hartm.) Greuter et Burdet, Willdenowia 12(1): 37

1982

—=

glomeratum Thuill., Fl. Env. Paris, ed. 2, 2: 226. 1799.

PARONYCHIA

AH T franciscana Eastw., Bull. Torrey Bot. Club 28: 288. 1901.

POLYCARPON

tetraphyllum (L.) L., Syst. Nat., ed. 10, 881. 1759.

SAGINA

AA F chilensis Naudin, in Gay, Hist. Chile, Bot. 1: 282. 1846,

SILENE

AA TSF gallica L., Sp. Pl. 417. 1753.

SPERGULARIA

EH TF confertiflora Steud. var. confertiflora, Flora 39: 425. 1856.

EH TF confertiflora Steud. var. polyphylla (Phil.) Skottsb., Nat. Hist. Juan Fernandez 2:

123. 1921

EH F —— masafuerana Skottsb., Nat. Hist. Juan Fernandez 2: (Ego e alas Tio al a

A AH T rubra J.Presl et K.Presl, Fl. Cech. 94. 1819.

STELLARIA

AH T chilensis Pedersen, Bonplandia (Corrientes) 5(22): 204, f. 1 A-E. 1983.

AA TF media (L.) Cirillo, Essent. Pl. Char. Comment. 36, t. 2, f. 9. 1784.

CHENOPODIACEAE

BETA

AH T— vulgaris L. subsp. maritima (L.) Arcang., Comp. Fl. Ital. 593. 1882.

CHENOPOD IUM

TF ambrostioides L., Sp. Pl. 219. 1753.

T— crusoeanum Skottsb., Nat. Hist. Juan Fernandez 2: 119, f.6b, 7 f-k; t. 11. 1921.

multredum to. Se PE. 220: 55:

TSF murate £..,, Sp. PL. 219. 1753:

F nesodendron Skottsb., Nat. Hist. Juan Fernandez 2: 119, f. 6 c, 7 l-o. 1921.

TS sanctae-clarae Johow, Estud. Fl. Juan Fernandez 119. 1896. (Introduced in

Masatierra).

mma>oss>m iy»

"M1. WYPS<SN

pa]

bed]

SARCOCORNIA

NH TSF fruticosa (L.) A.J.Scott, Bot. J. Linn. Soc. 75(4): 367. 1978.

COMPOSI TAE

ABROTANELLA

linearifolia A.Gray, Proc. Amer. Acad. Arts 5: 137. 1861. ("1862")

AMBLYOPAPPUS

AA TS pusillus Hook. et Arn., J. Bot. (Hooker) 3: 321. 1841.

195

Gayana Bot. 55(2), 1998.

ANTHEMIS

cotuta to, Sp. PL. 994. 1/53.

ARTEMISIA

AN TY absinthiumL., Sp. Pl. 848. 1753.

BAHIA

AF F ambrosioides Lag., Gen. Sp. Pl. 30. 1816.

BIDENS

AA TF: pitoes L:, Sp. Pl. 332. 153.

CALENDULA

KA... T. offñematis Ll... Sp Pl 920. 1703.

CARTHAMUS

AGA. FF. tanatus:L:. Sp. PL 890. 1733.

CENTAUREA

AA. .TSF metitensis L., Sp. PL. 917. 1/53.

CENTAURODENDRON (E)

E TR T dracaenoides Johow, Estud. Fl. Juan Fernandez 63. 1896.

E TR T palmiforme Skottsb., Bull. Jard. Bot. Belg. 27(4): 587, f. 59-61. 1957.

CHRYSANTHEMUM

KA E coronaria E... 34. PL.0890:) 1/53.

CI CHOR IUM

A AB TF "ntybus-L., sp PL: 813. 1753.

CIRSIUM

AA F vulgare (Savi) Ten., Fl. Nap. 5: 209. 1835-38.

CONY ZA

AA TSF bonariensis (L.) Cronquist, Bull. Torrey Bot. Club 70: 632. 1943.

COTULA

australis (Sieber ex Spreng.) Hook.f., Fl. Nov.-Zel. 1: 128. 1852.

AH OT corenepitolia L., Sp. PL. -892.. 1753;

CREPIS

AA T capillaris (L.) Wallr., Erst. Beitr. Fl. Hercyn. 287. 1840.

CYNARA

A >. TF: carduncutus L. Sp. PL, 827.017593.

DENDROSERIS (E)

E FTR T— berteroana (Decne.) Hook. et Arn., Companion Bot. Mag. 1: 32. 1835.

gigantea Johow, Estud. Fl. Juan Pernia 69. 1896.

E FTR TS litoralis Skottsb., Nat. Hist. Juan Fernandez 2: 204, f. 37 k-l, p; t. 19, t. 20, f

—m

—

- TEE

macrantha (Bertero ex Decne.) Skottsb., Nat. Hist. Juan Fernandez 2: 202, f. 37 b-d,

n. 1921

E FTR FE macrophylla D.Don, Philos. Mag. Ann. Chem. 11: 388. 1832.

Vascular flora of the Róbinson Crusoe Islands: MARTICORENA, C. ET AL.

mo "momomomom

mo mimim

mom mom mim

ABH

—

—m

—

—m

—

—m

—

pa]

—

marginata (Bertero ex Decne.) Hook. et Arn., Companion Bot. Mag. 1: 32. 1835.

micrantha (Bertero ex Decne.) Hook. et Arn., Companion Bot. Mag. 1: 32. 1835,

neriifolia (Decne.) Hook. et Arn., Companion Bot. Mag. 1: 32. 1835.

pinnata (Bertero ex Decne.) Hook. et Arn., Companion Bot. Mag. 1: 32.

pruinata (Johow) Skottsb., Nat. Hist. Juan Fernandez 2: 207, f. 39 d- E ae

regia Skottsb., Nat. Hist. Juan Fernandez 2: 209, 5.385 tc 17. 2. 10d

ERIGERON

fernandezianus (Colla) Solbrig, Contr. Gray Herb. 191: 30. E. 2h 1962.

ingae Skottsb., Nat. pe Juan Fernandez 2: 184, f. 30 a-d; t. 16, 1. 1921.

luteoviridis Skottsb., Nat. Hist. Juan Fernandez 2: 183, f. 29 f-i. 1921.

rupicola Phil., Ass Univ. Chile 13: 165.

GAL INSOGA

paruitlora:Cav.. “Icon. 45. 61:00 281-1295.

GAMOCHAETA

coarctata (Willd.) Kerguélen, Lejeunia, n.s. 120: 104. 1987.

fernandeziana (Phil.) Anderb., Opera Bot. 104: 157. 1991.

stachydifolia (Lam.) Cabrera, Bol. Soc. Argent. Bot. 9: 382. 1961.

GNAPHAL IUM

aldunateoides J.Remy, in Gay, Hist. clas Bot. 4: 232

cheiranthifolium Lam., Encycl. 2: 752. 1788. ("cheranthifol jun")

HYPOCHAERIS

glabra L., Sp. PL..841. 1/53.

radiata... Sp. PE 801. 105,

LAGENOPHORA

ariotij Franchet, Miss. Sci. Cap Horn 5, Bot.: 344. 1889.

LAPSANA

conmmnis:L., Sp BC. 31. 1753.

MADIA

sativa Molina, Sag. Stor. Nat. Chili 136, 354. 1782.

MATRICARIA

tecutita Loy Sp. PL. M2 20 NA

MICROPSIS

nana DC., Prodr. 5: 460. 1836.

ROBINSONIA (E)

berteroi (DC.) Sanders, Stuessy et Martic., Gayana, Bot. 47(3-4): 79. 1990.

evenia Phil., Anales Univ. Chile 13: 166. 1856.

gayana Decne., Ann. Sci. Nat., Bot., A dr EA A

gracilis Decne., Ann. Sci. Nat., Bot., ser. 2, 1529. 19346

macrocephala Decne., Ann. Sci. Nat., Bot., sér. 2 Pos EU a

1834.

Ea Y

masafuerae Skottsb., Nat. Hist. Juan Fernandez 2: 1951.39. ps, 36. 8, 3).€,

thurifera Decne., Ánn. Sci. Nat., Bot., Sér. £,.1: 28. 1834.

SENECIO :

mikanioides Otto ex Walp., Allg. Gartenzeitung 13: 42. 1845.

197

Gayana Bot. 55(2), 1998.

AR TF. Wigeris L., Sp. PC. 007. 1133.

SILYBUM

AA TSF marianum (L.) Gaertn., Fruct. Sem. Pl. 2: 378, t. 162, f. (2). 1791.

SOL IDAGO

AH T chilensis Meyen, Reise 1: 311. 1834.

SONCHUS

AA TSF oleraceus L., Sp. Pl. 796. 1753.

A AB TSF tenerrimus L., Sp. Pl. 794. 1753.

TARAXACUM

NH TF fermandezianum Dahlst., in Skottsb., Hist. Juan Fernandez 2: 226. 1922.

NH T subspathulatum A.J.Richards, Rhodora lo: 692. 1976.

XANTHIUM

RA TF 'spinosun L., Sp. Pl. 987. 1733,

YUNQUEA (E)

E TR T tenzii Skottsb., Acta Horti Gothob. 4: 163, f. 14-15. 1929.

CONVOLVULACEAE

CALYSTEGIA

tuguriorum (G.Forst.) R.Br. ex Hook.f., Fl. Nov.-Zel. 1: 183, t. 47. 1853.

CONVOLVULUS

AH TF arvensis L., Sp. Pl. 153. 1753.

DICHONDRA

NH TF sericea Suw., Prodr. 54. 1788.

IPOMOEA

indica (Burm.) Merr., Interpr. Herb. Amboin. 445. 1917.

—

CRUCIFERAE

BRASSICA

ACAB TSF. napus:L., Sp. Pt. 666

nigra (L.) W.D.J.Koch, in Róhling, Deutschl. Fl., ed. 3, 4: 713. 1833.

RAT rapa Eo, Sp. Pt. 666.. 1733.

—

CARDAMINE

NA T chenmopodiifolia Pers., Sym. Pl. 2: 195. sa issue dis

flaccida Cham. et Schltdl., Linnaea 1: 21. 1826.

kruesselii Johow ex Reiche, Anales Univ. Chile 9077126. 1895.

m =

HIRSCHFELDIA

AAB T incana (L.) Lagr.-Foss., Fl. Tarn Garenne 19. 1847.

LEPIDIUM

AA T bonmariense L., Sp. Pl. 645. 1753.

MATTHIOLA

AH TSF incana (L.) R.Br., in W.T. Aiton, Hortus Kew. 4: 119. 1812.

198

Vascular flora of the Róbinson Crusoe Islands: MARTICORENA, C. ET AL.

RAPHANUS

A.AB- TF. sativus"L:,:+Sp. PL. 669. 1733.

RORIPPA

nasturtium-aquaticum (L.) Hayek, Sched. Fl. Stiriac. 3-4: 22. 1905.

-—)

—m

SISYMBR IUM

AAB T officinale (L.) Scop., Fl. Carniol., ed. 2, 2: 26. 1772.

DIPSACACEAE

DIPSACUS

AB T sativus (L.) Honck., Verz. Gew. Teutschl. 374. 1782.

SCABIOSA

AH T atropurpurea L., Sp. Pl. 100. 1753.

ELAEOCARPACEAE

ARISTOTELIA

AT TF chilensis (Molina) Stuntz, U.S.D.A. Bur. Pl. Imdustr. Invent. Seeds 31: 85. 1914.

EMPETRACEAE

EMPETRUM

NS F rubrum Vahl ex Willd., Sp. Pl. 4: 713. 1806.

ERICACEAE

PERNETTYA

EE 4 rigida (Bertero ex Colla) DC., Prodr. 7: 587. 1839.

EUPHORBIACEAE

DYSOPSIS

E O hirsuta (Múll.Arg.) Skottsb., Nat. Hist. Juan Fernandez 2: 781. 1951.

EUPHORBIA

AH lathyris L., Sp. Pl. 457. 1753. ("lathyrus")

AA TF peplus L., Sp. Pl. 456. 1753.

RICINUS

AF TF communis L., Sp. Pl. 1007. 1753.

FLACOURTIACEAE

AZARA

ER serrata Ruiz et Pavón var. fernandeziana (Gay) Reiche, Anales Univ. Chile 90: 884.

1895.

FUMAR IACEAE

FUMAR IA

RA T capreolata l., Sp. Pl. 701. 1753.

Gayana Bot. 55(2), 1998.

mmmm

TATI

—

—W

NAAA

—

-n

3 mn

—

—m

GENTIANACEAE

CENTAUR IUM

cachanlahuen (Molina) B.L.Rob., Proc. Amer. Acad. Arts 45(17): 396. 1910.

- GERANTACEAE

EROD I1UM

cicutarium (L.) L'Hér. ex Aiton, Hort. Kew. 2: 414. 1789.

GERAN IUM

core-core Steud., Flora 39: 438. 1856.

dissectum L., Cent. Pl. 1, 21. 1755.

robertianum L., Sp. Pl. 681. 1753.

GUNNERACEAE

GUNNERA

bracteata Steud. ex Bennett, Pl. Jav. Rar. 75. 1838.

bracteata x peltata, Nat. Hist. Juan Fernandez 2: 148, f. 1606-55 t, 3, 2. 1922.

glabra Phil., Anales Univ. Chile 13: 164. 1856. (= bracteata)

masafuerae Skottsb., Kongl. Svenska Vetenskapsakad. Handl. 51(9): 14, f. 3; t. 3 et

. 1914

peltata Phil., Anales Univ. Chile 13: 165. 1856.

GUTTIFERAE

HYPER I CUM

perforatum L., Sp. Pl. 785. 1753.

HALORAGACEAE

HALORAGIS

masafuerana Skottsb. var. asperrima (Skottsb.) Orchard, Bull. Auckland Inst. Mus. 10:

81. 1975.

masafuerana Skottsb. var. masafuerana, Nat. Hist. Juan Fernandez 2: 156, f..20. 1921.

masatierrana Skottsb., Nat. Hist. Juan Fernandez 153. 10 192%

LABIATAE

CUMINIA (E)

eriantha (Benth.) Benth. var. eriantha, in DC., Prodr. 12: 258. 1848.

eriantha (Benth.) Benth. var. fernandezia (Colla) Harley, Kew Mag. 3(4): 155, f. B.

1986.

MARRUB IUM

vulgare L., Sp. Pl. 583. 1753.

MELISSA

officinalis L., Sp. Pl. 592. 1753.

MENTHA

aquatica L., Sp. Pl. 576. 1753.

pulegium L.. Sp. Pt. 577. 1753.

Vascular flora of the Róbinson Crusoe Islands: MARTICORENA, C. ET AL.

—

ORI GANUM

majorana L., Sp. Pl. 590. 1753.

PRUNELLA

vulgaris -L. , Sp. PL. 600. 1753.

STACHYS

arvensis (12) 'L.,' Sp. Pb., 6d. 2, 814. 1/83

LACTORIDACEAE (E)

LACTORIS (E)

fernandeziana Phil., Anales Univ. Chile 26: 642. 1865.

LARDIZABALACEAE

LARDIZABALA

biternata Ruiz et Pavón, Syst. Veg. Fl. Peruv. Chil. 286.

LINACEAE

LINUM

usitatissimum L., Sp- PL. 27. 193.

LORANTHACEAE

NOTANTHERA

1798.

heterophylla (Ruiz et Pavón) G.Don, Gen. Hist. 3: 429. 1834.

TRISTERIX

corymbosus (L.) Kuijt, Syst. Bot. Monogr. 19: 20, f. 6. 1988.

LYTHRACEAE

LYTHRUM

hyssopifolia L., Sp. Pl. 447. 1753.

MALVACEAE

ANODA

cristata (L.) Schltdl., Linmaea 11: 210. 1837.

MALVA

nicaeensis All., Fl. Pedem. 2: 40. 1785.

parviflora L., Demonstr. Pl. 18. 3

MOD IOLA

caroliniana (L.) G.Don, Gen. Hist. 1: 466. 1831.

MIMOSACEAE

ACACIA

dealbata Link, Enum. Pl. Hort. Berol. Alt. 2: 465: 1822.

melanoxylon R.Br., in W.T. Aiton, Hortus Kew. 5: 462. 1813.

201

202

—

bas |

A —-

—

ás]

—

+ —

ll y

ALBIZIA

lophantha (Willd.) Benth., London J. Bot. 3: 86. 1844.

MORACEAE

FICUS

carice Lo, Sp. PE. 1059 1753:

MYRTACEAE

MYRCEUGENIA

ernandeziana (Hook. et Arn.) Johow, Estud. Fl. Juan Fernandez ye, €. 2. 1996.

schulzei Johow, Estud. Fl. Juan Fernandez 96. 1896. ("schulzii")

MYRTEOLA

nummularia (Poir.) O.Berg, Linnaea 27: 396. 1856.

UGNI

molinae Turcz., Bull. Soc. Imp. Naturalistes Moscou 21(1): 579. 1848.

selkirkii (Hook. et Arn.) O.Berg, Linnaea 27: 392. 1856.

ONAGRACEAE

FUCHSIA

magellanica Lam., Encycl. 2: 565. 1788.

OENOTHERA

affinis Cambess., in A.St.-Hil., Fl. Bras. Merid. 2: 269. 1830.

picensis Phil., dj items PEC. 22. 189).

rosea L'Hér. ex Aiton, Hort. Kew. 2: 3. 1789.

OXAL IDACEAE

OXALIS

corniculata:L., Sp. PL.:435.. 1753.

micrantha rimel ex Colla, Nuovo Giorn. Lett., Sci. 24: 145. 1832.

PAPAVERACEAE

PAPAVER

somiferum L., Sp. Pl. 508. 1753.

PAPILIONACEAE

MEDICAGO

arabica (L.) Huds., Fl. Angl. 288. 1762.

tupulina L., Sp. Pl. 779. 1753.

polymorpha L., Sp. Pl. 779. 1753.

sativa L., Sp. Pl. 778. 1753.

MELILOTUS

indicus (L.) All., Fl. Pedem. 1: 308. 1785.

SOPHORA

fernandeziana (Phil.) Skottsb. var. fernandeziana, Nat. Hist. Juan Fernandez 2: 140,

> >>>

m mmm

E E E E

YITYTA

—

—W

—_

—

—_

-—

f. 128. 11.

Fernandez 2: 140, f. 11 c, 12 CAN - SE ar Ia OE

fernandeziana (Phil.) Skottsb. var. reedeana (Phil. : Skottsb

1921.

masafuerana (Phil.) Skottsb., Nat. Hist. Juan Fernandez 2 Me fe

2. 1921

TELINE

monspessulana (L.) K.Koch, Dendrologie 1: 30. 1869.

TRIFOLIUM

campestre Schreber, in Sturm, Deutschl. Fl., Abt. 1(4) Heft 16, t. 253. 1804.

dubium Sibth., Fl. Oxon. 231. 1

pratense L.., Sp. PL. 768. 1753.

repens L.., Sp. Pt. 76f. 1753.

PIPERACEAE

PEPEROMIA

berteroana Miq. subsp. berteroana, Syst. Piperac. 114. 1843.

fernandeziana Miq. f. fernandeziana, Syst. Piperac. 139. 1843.

fernandeziana Mig. f. oblongifolia Skottsb., Acta Horti Gothob.

margaritifera Bertero ex Hook. var. margaritifera, Icon. Pl.

ar. umbraticola Skottsb. ex Yunck.,

margaritifera Bertero ex Hook. v

Lilloa 27: 282, .t. 163. 1955. -(u1953u)

skottsbergii C.DC., Kongl. Svenska Vetenskapsakad. Handl . 51(9): 20.

PLANTAGINACEAE

PLANTAGO

australis Lam., Tabl. Encycl. 1: 339. 1792.

fernandezia Bertero ex Barnéoud, Monogr. Plantag. 47.

firma Kunze ex Walp., Nov. Actorum Acad. Caes. Leop.-Carol.

402. 1843.

tanceclata 12, Sp. Pl... 113:051753.

major "L.... Sp. PLLFTFIZ, 1753

POLEMONIACEAE

GILIA

valdiviensis Griseb., Syst. Bemerk. 43. 1854.

MICROSTERIS

gracilis (Douglas ex Hook.) Greene, Pittonia 3: 300.

POL YGONACEAE

POL YGONUM

avicutare ote, Sp. PL. 362: TH

hydropiperoides Michx., Fl. Bor.-Amer. 1: 239. 1803.

lapathifolium L., Sp. Pl. 360. 1753

RUMEX

acetosetla £.. Sp. PL. 338. 1/53.

conglomeratus Murray, Prodr. Stirp. Gott. 52. 1770.

crispus Lo. (Sp pt: 0335. 1793.

fotiosus Rech.f., Ark. Bot. 264(3): 42, t. 6. 1933.

1845.

1898.

A A AA

, 4. 49. 1947.

sine desc. lat.,

Nat. Cur. 19, Suppi. Ts

203

204

TSF pulcher L., Sp. Pl. 336. 1753.

ml 9

PORTULACACEAE

CALANDRINIA

monandra (Ruiz et Pavón) DC., Prodr. 3: 359. 1828.

PORTULACA

oleracea L., Sp. Pl. 445. 1753.

PRIMULACEAE

ANAGALLIS

arvensis L., Sp. Pl. 148. 1753.

minima (L.) E.H.L.Krause, in Sturm, Deutschl. Fl., ed. 2, 9: 251. 1901.

RANUNCULACEAE

ANEMONE

decapetala Ard., Animadv. Bot. Spec. Alt. 27, t. 12. 1764.

RANUNCULUS

caprarum Skottsb., Nat. Hist. Juan Fernandez 2: 123, f. Ba-€e.: 1921:

suricatus L., Sp. Pt. 555. 1153.

RHAMNACEAE

COLLETIA

spartioides Bertero ex Colla, Nuovo Giorn. Lett., Sci. 24: 146. 1832.

ROSACEAE

ACAENA

argentea Ruiz et Pavón, Fl. Peruv. Chil. 1: 67, t. 103, f. b. 1798.

masafuerana Bitter, Biblioth. Bot. 17(74): 45, t. 2. 1911.

ovalifotia Ruiz et Pavón, Fl. Peruw. Chil. 1: 67, t. 103, f. €. 1798.

FRAGARIA

chiloensis (L.) Duchesne, Hist. Nat. Frais. 165. 1766.

MARGYRACAENA

skottsbergii Bitter, Repert. Spec. Nov. Regni Veg. 17: 239. 1921.

MARGYRICARPUS

digynus (Bitter) Skottsb., Nat. Hist. Juan Fernandez 2: 779. 1951.

RUBUS

gegides Sm., Pt. Icom. Ined. E: t. 19. 17/89.

ulmifolius Schott, Isis (Oken) 2(5): 821. 1818.

RUBIACEAE

COPROSMA

oliveri Fosberg, Brittonia 20(4): 292. 1968.

pyrifolia (Hook. et Arn.) Skottsb., Nat. Hist. Juan Fernandez 2: 173, f. 24 d.

1921.

E TF

—m

—

—.

+= -

GAL 1 UM

aparine L., Sp. Pl. 108. 1

masafueranum Skottsb., Nat. H

HEDYOTIS

133

ist. Juan Fernandez 2: 174, f. 25. 1921.

salzmannii (DC.) Steud., Nomencl. Bot., ed. 2, 1: 728. 1840.

NERTERA

granadensis (Mutis ex L.f.) Druce, Bot. Soc. Exch. Club Brit. Isles 4: 637. 1917.

FAGARA

RUTACEAE

externa Skottsb., Nat. Hist. Juan Fernandez 2: 143, f. 14 e. 1921.

mayu (Bertero ex Colla) Engler, Nat. Pflanzenfam. 3(4): 118. 1896.

RUTA

chalepensis L., Mant. Pl. 69. 1767.

SANTALUM

fernandezianum F.Phil., Anales Mus. Nac. Chile, Secc. 2, Bot. 5, t

(Extinct)

ESCALLONIA

SANTALACEAE

SAX IFRAGACEAE

callcottiae Hook. et Arn., Bot. Misc. 3: 342. 1833.

CYMBALARIA

muralis P.Gaertn.,

DIGITALIS

SCROPHULARIACEAE

purpurea L., Sp. Pl. bat. 31795.

EUPHRASIA

formosissima Skottsb., Nat.

6-7. 1921.

KICKXIA

elatine (L.) Dumort., Fl.

MIMULUS

glabratus Kunth,

VERBASCUM

Beto. 35. 1827.

B.Mey. et Scherb., Oekon. Fl. Wetterau 2: 39%. 1800.

1892.

Hist. Juan Fernandez 2: 169, f. 23; t. PEFTO Eu, ES

in Humb., Bonpl. et Kunth, Nov. Gen. Sp. (ed. qu.) 2: 370. 1818.

thapsus L., Sp. Pl. a Y pb

virgatum Stokes, in With., Bot. Arr. Brit. Pl., ed. 2, 1: 227. 178.

VERONICA

anagallis-aquatica L., Sp- PL. 12, 13%

arvensis L., Sp. Pl

33.

1753.

205

P>PE>GoSmo>

TIXC_I>OCT

206

T persica Poir., in Lam., Encycl. 8: 562.. 1808.

SOLANACEAE

CESTRUM

1F-.parqui L'Heér., SETED: Now. (3,.t. 36..1/88.

DATURA

L-” setramobiun E... Sp. PE 17%. 1793.

NICOTIANA

F cordifolia Phil., Anales Univ. Chile 13: 167. 1856.

Y -Cabacun Li. Sp. PU. 180. 17035

PHYSALIS

TF peruviana L., Sp. Pl-, ed. 2, 1670. 1163.

SOLANUM

TF argenteum Dunal ex Poir., in Lam., Encycl., Suppl. 3: 755. 1814.

T —— fernandezianum Phil., Linmaea 29: 23. 1858.

TSF furcatum Dunal ex Poir. var. furcatum, in Lam., Encycl., Suppl. 3: 750. 1814.

T . margihatun L.f., Suppt. PU. 147. 1782.

F —— pentlandii Dunal subsp. interandinum (Bitter) Edmonds, Kew Bull. 27(1): 110. 1972.

PT... «tuberosum Es, Sp. Pb. 185 "1753.

TROPAEOLACEAE

TROPAEOLUM

TT. Majes Eo Sp PE 345. 1753.

UMBELLIFERAE

AMMI

TT... visnaga (L-) Eam., El. Franc. 3: 462. 1779.

ANETHUM

F . graveolens Ex, Sp. PE. -263..1753.

AP 1UM

T— australe Thouars, Esquisse Fl. Tristan d'Acugna 43. 1808.

T fernandezianum Johow, Estud. Fl. Juan Fernandez 101. 1896.

T graveolens L., Sp. Pl. 264. 1753.

CENTELLA

T— asiatica (L.) Urb., in Mart., Fl. Bras. 11(1): 287, t. 78, f. 1. 1879.

CON 1UM

TF maculatum L., Sp. Pl. 243. 1753.

COR! ANDRUM

FT” sativue to. Sp. PL: 256. 1753.

CY CLOSPERMUM

T leptophyllum (Pers.) Sprague var. leptophyllum, J. Bot. 61: 131. 1923.

—n

177

—

DAUCUS

montanus Humb. et Bonpl. ex Spreng., in Schult., Syst. Veg. 6: 482. 1820.

ERYNGIUM

bupleuroides Hook. et Arn., Bot.

sarcophyllum Hook. et Arn., Bot.

Xx fernandezianum Skottsb., Kongl.

914.

FOEN I CULUM

vulgare Mill.

PETROSEL I NUM

crispum (Mill.) A.W.Hill, Hand-l

SANICULA

crassicaulis Poepp. ex DC., Prod

TORILIS

nodosa (L.) Gaertn., Fruct. Sem.

Misco 3302. 1033-

Juan Fernandez 2: 158, 21 h-p, t. 12, 2. 1921.

Misco” 3: 3592. 1835.

Svenska Vetenskapsakad. Handl. 51(9): 17, f. 5.

, Gard. Dict., ed. 8, Foeniculum 1. 1768.

ist Herb. Pl. Kew, ed. 3, 122. 1925.

r..6< 84. 1830.

PA 4. 20 de 0 EOA:

TICACEAE

BOEHMERIA

excelsa (Bertero ex Steud.) Wedd.,

" PARIETARIA

debilis G.Forst. var. debilis, F

URTICA

berteroana Phil., Linnaea 33: 235

glomeruliflora Steud., Flora 33:

masafuerae Phil., Anales Univ. €

urensk. ¿Sp Pl. 984 113%

Ann. Set. Nat. Bot, Ser. 6, 1: 202. 1956,

l. Ins. Austr. 73. 1786.

1864.

257. 1850. ("glomerulaeflora")

hile 13: 167. 1856

LERIANACEAE

CENTRANTHUS

ruber (L.) DC., in DC. et Lanm.,

FL. Franc, ed... 3, 4: 23%. 1805.

RBENACEAE

RHAPHI THAMNUS

venustus (Phil.) B.L.Rob., Proc.

VERBENA

ltitoralis Kunth, in Humb., Bonpl.

1818

DRIMYS

confertifolia Phil., Anales Univ.

Amer, Acad. Arts 51(10): 531. 1916.

NTERACEAE

Chite 13: 163.. 1856.

et Kunth, Nov. Gen. Sp. (ed. qu.) 2: 276, t. 137.

207

e mmaz

TIN ZA

—

n —

pes

—

MONOCOTYLEDONEAE

ARACEAE

ZANTEDESCHIA

aethiopica (L.) Spreng., Syst. Veg. 3: 765. 1826.

BROMELIACEAE

GREIGIA

berteroi Skottsb., Nat. Hist. Juan Fernandez 2: 109. 1921.

OCHAGAVIA (e)

elegans Phil., Anales Univ. Chile 13: 168. 1856.

CYPERACEAE

CAREX

banksii Boott, Trans. Linn. Soc. London 20: 119. 1846.

berteroniana Steud., Flora 25: 604. 1842.

CYPERUS

eragrostis Lam., Tabl. Encycl. 1: 146. 1791.

reflexus Vahl, Enum. Pl. 2: 299. 1805.

ELEOCHARIS

fuscopurpurea (Steud.) H.Pfeiff., Repert. Spec. Nov. Regni Veg. 28: 19. 1930.

MACHAER INA

scirpoidea (Steud.) Koyama ex M.T.Strong, Novon 7(3): 318. 1997.

OREOBOLUS

obtusangulus Gaudich. subsp. obtusangulus, Ann. Sci. Nat. (Paris) 5: CO ar

1825

SCIRPUS

cernuus Vaht,. Enum. Pt. 2: 245. 1805.

nodosus Rottb., Descr. Icon. Rar. Pl. 52. 1773.

UNCINIA

brevicaulis Thouars, Esquisse Fl. Tristan d'Acugna 35, t. 6. 1808.

costata Kuk., Repert. Spec. Nov. Regni Veg. 16: 433. 1920.

douglasii Boott, in Hook.f., Fl. Antarct. 369. 1

tenuis Poepp. ex Kunth, Enum. Pl. 2: 525. 1837.

GRAMINEAE

AGROSTIS

masafuerana Pilger, Repert. Spec. Nov. Regni Veg. 16: 388. 1920.

stolonifera L., Sp. Pl. 62. 1753

AIRA

caryophyllea L., Sp. Pl. 66. 1753.

pEaecox' E... Sp. PL..65. 1753.

Do>p>p>Z

—

—.

—_

—=

—

pu]

—

soil

ANTHOXANT HUM

odoratum L., Sp. Pl. 28. 1753.

AVENA

barbata Pott ex Link, J. Bot. (Schrader) 1799(2): 315. 1800.

BRACHYPOD IUM

distachyon (L.) P.Beauv., Essai Agrostogr. 101, 156. 1812.

BRIZA

maxima L., Sp. Pl. 70. 1753,

minor L., Sp. PL. 70. 1753.

BROMUS

berteroanus Colla, Herb. Pedem. 6: 68. 1836. ("'berterianus")

catharticus Vahl, Symb. Bot. 2: 22. 1791.

diandrus Roth, Bot. Abh. Beobacht. 44. 1787.

hordeaceus L., Sp. Pl. 77.

lithobius Trin., Linnaea 10: 303. 1836.

stamineus E.Desv., in Gay, Hist. Chile, Bot. 6: 440. 1854.

CHAETOTROPIS

chilensis Kunth, Révis. Gramin. 271, t. 47. 1830.

imberbis (Phil.) Bjorkman, Symb. Bot. Upsal. 17(1): 14. 1960.

CHASCOLYTRUM

subaristatum (Lam.) Desv., Nouv. Bull. Sci. Soc. Philom. Paris 2: 190. 1810.

CHUSQUEA

fernandeziana Phil., Anales Univ. Chile 43: 577. 1873.

CYNOSURUS

echinatus L..., Sp. Bl. 72. 17353.

DACTYLIS

glomerata L.,. Sp. Plis 71. 1753

DANTHONIA

chilensis E.Desv. var. chilensis, in Gay, Hist. Chile, Bot. 6: 360, t. 80, f. 3,

1854.

malacantha (Steud.) Pilger, Notizbl. Bot. Gart. Berlin-Dahlem 10(97): 759. 1929.

DIGITARIA

sanguinalis (L.)» Scop., Fl. Carniol.;, ed. 2, Ti: 52. 172.

ELEUSINE

tristachya (Lam.) Lam., Tabl. Encycl. 1: 203. 1792.

FESTUCA

arundinacea Schreb., Spic. Fl. Lips. 57. 1771.

GASTRI1DI1UM A

ventricosum (Gouan) Schinz et Thell., Vierteljahrsschr. Naturf. Ges. Zúrich 58: 39.

1916.

210

—

—Á4

—Á

€

—

A Ms

HORDEUM

chilense Roem. et Schult., Syst. Veg. 2: 796. 1817.

murinum L. subsp. murinum, Sp. Pl. 85. 1753.

secalinum Schreber, Spic. Fl. Lips. 148. 1771.

LEPTOPHYLLOCHLOA

micrathera (E.Desv.) C.E.Calderón ex Nicora, in M.N.Correa, Fl.

1978.

LOL IUM

multiflorum Lam., Fl. Franc. 3: 621. 1779.

perenne L., Sp. Pl. 83. 1753.

MEGALACHNE (E)

berteroniana Steud., Syn. Pl. Glumac. 1: 237. 1854.

Patag. 3: 70, f. 32.

masafuerana (Skottsb. et Pilger ex Pilger) Matthei, Bol. Soc. Biol. Concepción 48:

Mi KT EE DA

NASSELLA

laevissima (Phil.) Barkworth, Taxon 39(4): 610. 1990.

neesiana (Trin. et Rupr.) Barkworth, Taxon 39(4): 611. 1990.

PASPALUM

dasypleurum Kunze ex E.Desv., in Gay, Hist. Chile, Bot. 6: 242.

distichum L., Syst. Nat., ed. 10, 855. 1759.

PHALARIS

amethystina Trin., Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér.

Seconde Pt. Sci. Nat. 5(2): 56. 1839.

angusta Nees ex Trin., Sp. Gram. 1: t. 78. 1827.

PIPTOCHAET IUM

bicolor (Vahl) E.Desv., in Gay, Hist. Chile, Bot. 6: 273. 1854.

POA

annua L., Sp. Pl. 68. 1753.

pratensis L.,.Sp. Pl. 07. 1/35.

PODOPHORUS (E)

bromoides Phil., Anales Univ. Chile 13: 169. 1856. (Extinct?)

POLYPOGON

australis Brongn., in Duperrey, Voy. Monde Phan. 21. 1829.

SETARIA

parviflora (Poir.) Kerguélen, Lejeunia, n.s. 120: 161. 1987.

viridis (L.) P.Beauv., Essai Agrostogr. 51, 171, 178, t. 13, f.

TRISE TUM

caudulatum Trin., Linmaea 10: 300. 1836.

VULPIA

bromoides (L.) Gray, Nat. Arr. Brit. Pl. 2: 124. 1821.

1854.

6, Sci. Math.,

S. JB.

myuros (L.) C.C.Gmel. var. hirsuta Hackel, Cat. Rais. Gramin. Portugal 24. 1880.

myuros (L.) C.C.Gmel. var. myuros, Fl. Bad. 1: 8. 1805. ('"myurus

Vascular flora of the Róbinson Crusoe Islands: MARTICORENA, C. ET AL.

IRIDACEAE

CROCOSMIA

AH Tx ocrocosmiiflora (Lemoine ex Burb. et Dean) N.E.Br., Trans. Roy. Soc. South Africa

20: 2664. 1932

LIBERTIA

NH TF chilensis (Molina) Gunckel, Revista Chilena Hist. Nat. 31: 87. 1928. (19271)

JUNCACEAE

JUNCUS

AR =P BUON L., Sp. PL. 328. 1753.

N H TF capillaceus Lam., Encycl. 3: 266. 1789.

NH T ¡imbricatus Laharpe, Essai Monogr. Jonc. 61. 1823

NH T pallescens Lam., Encycl. 3: 268. 1789

NH T planifolius R.Br., Prodr. Fl. Nov. Holl. 259. 1810.

NH T procerus E.Mey., Linnaea 3: 367. 1828.

LUZULA

EH. F masafuerana Skottsb., Nat. Hist. Juan Fernandez 2: 775. 1951.

ORCHIDACEAE

GAVILEA

EN F insularis M.N.Correa, Revista Mus. La Plata, Secc. Bot. 11: 75, f. A-E. 1968.

PALMAE

JUANIA (E)

ET T— australis (Mart.) Drude ex Hook.f., Rep. Prog. Condition Roy. Bot. Gard. Kew 1882:

57. 1884

Fecha de publicación: 07 de mayo de 1999

211

NUEVOS TAXA Y COMBINACIONES PUBLICADOS EN ESTE VOLUMEN

LICHENES

Caloplaca austroshetlandica (Zahlbr.) D.J.Galloway £ Quilhot, com. nov. 120

Dendriscocaulon calithamnion (Taylor) D.J.Galloway £ Quilhot, com. nov. 127

Neuropogon durietzii (Motyka) D.J.Galloway $ Quilhot, com. nov. 142

Neuropogon patagonicus (F.J. Walker) D.J.Galloway £ Quilhot, com. nov. 142

Neuropogon subantarcticus (F.J. Walker) D.J.Galloway $ Quilhot, com. nov. 142

ANGIOSPERMAE

Poaceae

Nassella argentinensis (Speg.) Peñail., com. nov. 86

Nassella brasiliensis (A. Zanin et Longhi-Wagner) Peñail., com. nov. 86

Nassella coquimbensis (Matthei) Peñail., com. nov.

Nassella entrerriensis (Burkart) Peñail., com. nov. 86

Nassella ibarrensis (Kunth) Peñail., com. nov. 86

Nassella karstenii (Hitchc.) Peñail., com. nov. 86

Nassella pittieri (Hitchc.) Peñail., com. nov. 86

Nassella psittacorum (Speg.) Peñail., com. nov. 86

Nassella planaltina (A. Zanin et Longhi-Wagner) Peñail., com. nov. 87

Nassella rhizomata (A. Zanin et Longhi-Wagner) Peñail., com. nov. 87

Nassella sellowiana (Nees ex Trin. et Rupr.) Peñail., com. nov. 8

Nassella tenuiculmis (Hackel) Peñail., com. nov. 87

Nassella vallsii (A. Zanin et Longhi-Wagner) Peñail., com. nov. 87

Nassella vargasii (Tobar) Peñail., com. nov.

213