A FIRST CENTURY OF NEW SPECIES OF v

STELIS OF ECUADOR

CONTENTS

New species and introduction to Stelis 1-6

New species of section Dialissa 7-8

Illustrations 27

New species of section Nexipous 8-9

Illustrations OR

New species of section Humboldtia 9-26

Illustrations 28-35

New species of section Stelis 36-64

Illustrations 65-78

New species of section Labiatae 79-83

Illustrations 84-85

Index to new species of Stelis 86

Addenda to the Lepanthes of Ecuador 87-93

Illustrations 94-96

Addenda to taxa previously treated 97-114

Illustrations 115-122

ACKNOWLEDGMENTS

Alexander C. Hirtz of Quito has been included as co-author of the majority of

species, because, without his ability to gather these collections, this work would

have been impossible. Recently, vast collections cultivated by José Portilla at

Ecuagenera, in Gualaceo, have also produced numerous new species.

Also, in addition to his collections, I am grateful to Stig Dalstrém for his untold

hours of meticulous inking of most of the illustrations, and for the generosity of the

Pleurothallid Alliance that made it possible.

As usual, I owe gratitude to Mrs. Amy McPherson for her hours of proof-

reading the manuscript.

Determinations of the status of about half of over 1,400 Ecuadorian collections

at MO and SEL, organized by Lorena Endara A. of Quito, have been made. These

determinations were made possible by examinations of type-collections graciously

permitted by the herbaria of AMES, BR, G, K, and W, for which I am immensely

grateful. With the remaining collections, a second century of new species of Stelis

of Ecuador is in preparation.

Stelis hirtzii Luer, province of Pichincha, Ecuador, altitude 1450 meters

above sea level, 1 April 1984, Photograph by Alexander C. Hirtz.

SO a ee A

A FIRST CENTURY OF NEW SPECIES OF STELIS SW. OF ECUADOR

Schlechter published in 1921 a list of 78 species of Stelis that had been reported

from Ecuador. About 43 of these names are presently accepted, the remainder

being misidentified, erroneously attributed to Ecuador, or represented by synony-

mous names. Twenty-two names have been added since that

One hundred new species chosen at random from a number approaching 200

found in Ecuador are described herein. Instead of presenting “ie me in alphabeti-

cal order, they are presented alphabetically in five artificial “* ”* groups of

more or less allied species to aid in orientation and iienatieation. "ieee the

habit is the most readily recognized key character of the genus, it is impractical if

used as the first couplet in a key. Species with similar flowers would be dispersed

among the three basic habits: caespitose, repent, and prolific. Although also artifi-

cial, key characters of connation and position of the sepals offer the best solution.

KEY TO THE SECTIONS OF STELIS

1 Sepals variably connate, forming a tube or globose sepaline cup............... Dialissa

1’ Sepals not connate into a tubular or globose flower. 2

2 Lateral sepals much more deeply connate to the dorsal sepal than to themselves....

Nexipous

2’ Lateral sepals not deeply connate to the dorsal sepal

3 Flower bilabiate, with the lateral sepals deeply connate into a —— ——

lip usuall

boldtia

3’ Flower not bilabiate with a synsepal

Stelis

4 Sepals widely expanded

4’ Lateral sepals directed forward, more or less simulating a concave synsepal; lip

usually type-B

abiatae

Section Dialissa

Fig. 1. Stelis hirtzii Luer, sp. nov.

Fig. 2. Stelis saccata Luer & Hirtz, sp. nov.

Section Nexi

Fig. 3. Stelis matula Luer & Hirtz, sp. nov.

Fig. 4. Stelis zelenkoi Luer & Hirtz, sp. nov.

Section Humboldtia

Fig. 5. Stelis acutilabia Luer & Hirtz, sp. nov.

Fig. 6. Stelis ann-jesupiae Luer, sp. n

Fig. 7. Stelis in a Luer & Hirtz, sp. nov.

Fig. 8. Stelis applanata Luer & Hirtz, sp. nov.

Fig. 9. Stelis bivalvis Luer & Hirtz, sp. nov.

Fig. 10. Stelis bolivarensis Luer & Hirtz, sp. nov.

Fig. 14. Stelis dromedarina Luer & Hirtz, ba nov.

Fig. 15. Stelis st ei Luer & Hirtz, sp. n

Fig. 16. Stelis fissurata Luer & Hirtz, sp. nov.

Fig. 17. Stelis flava Luer & Hirtz, sp. nov.

Fig. 18. Stelis flexilis | Luer & Hirtz, a nov.

Fi . Stel

Fig. 21. Stelis aie Luer & Hirtz, sp. nov.

ag 22,

. Stelis parvipetala Luer & Hirtz, sp. nov.

Fig. 44.

Stelis nycterina Luer & Hirtz, sp. nov.

. Stelis rosamariae Luer & Hirtz, sp. nov.

. Stelis sanchezii Luer & — sp. nov.

. Stelis santiagoi Luer & Hirtz, sp. nov.

Stelis si itulifera Luer & Sine Sp. nov.

acoénsis Luer & Hirtz, sp. nov.

5. Stelis velivolva Luer & Hirtz, sp. nov

. Stelis vollesii Luer, sp. nov.

on Stelis

. Stelis adrianae Luer, sp. nov.

Stelis congesta Luer & Hirtz, ms nov.

2 ICONES PLEUROTHALLIDINARUM

Fig. 45. Stelis cutucuénsis Luer & Hirtz, sp. nov. Fig. 74. Stelis pachyphyta Luer & Hirtz, sp. nov.

Fig. 46. Stelis dalessandroi Luer, sp. nov. Fig. 75. Stelis phil-jesupii Luer, sp. nov.

Fig. 47. Stelis dalstroemii Luer, sp. nov. Fig. 76. Stelis pilosa Luer & Hirtz, Sp. Nov.

Fig. 48. Stelis dirigens Luer & Hirtz, Sp. nov. Fig. 77. Stelis polycarpica Luer & Hirtz, Sp. nov.

Fig. 49. Stelis distans Luer & Hirtz, Sp. nov. Fig. 78. Stelis porphyrea Luer & Hirtz, sp. nov.

Fig. 50. Stelis i Luer, sp. nov.

Fig. 79. Stelis procera Luer, sp. nov.

g. 80. Stelis protracta Luer & Hirtz,

Fig. 52. Stelis exasperata - Fig. 81. Stelis & Hirtz, sp. nov.

Fig. 53. Stelis excavata Luer & Hirtz, SP. nov. Fig. 82. Stelis purpurella Luer & Hirtz, sp. nov

Fig. 54. Stelis exilis Luer & Hirtz, SP. nov. Fig. 83. Stelis sanctae-rosae Luer & Hirtz, Sp. NOV.

Fig. 55. Stelis falcifera Luer & Hirtz, Sp. NOV. Fig. 84. Stelis scalena Luer & Hirtz, sp. nov

Fig. 56. Stelis Luer & Hirtz, sp. nov. Fig. 85. Stelis serrulifera Luer & Hirtz, sp. nov

Fig. 57. sissima Luer & Hirtz, sp. nov Fig. 86. Stelis tridactyloides Luer & Hirtz, Sp. NOV.

Fig. 58. Stelis floresii Luer & Hirtz, Fig. 87. Stelis undulata & Hirtz, sp. nov

Fig. 59. St rmosa Luer & V. Fig. 88. Stelis uxoria Luer & Hirtz, Sp. nov

Fig. 60. Stelis gigantea Luer & Hirtz, sp. nov. Fig. 89. Stelis vesca Luer & no

Fig. 61. Stelis glaberrima Luer & Hirtz, Sp. nov. Fig. 90. Stelis villifera Luer, sp. nov

Fig. 62. Stelis globulifera Luer & Hirtz, sp. nov. Fig. 91. Stelis yanganensis Luer & Hirtz, Sp. Nov.

Fig. 63. Stelis hoeijeri Luer & . NOV. Hirtz, OV.

Fig. 64 is jimburae Luer & Hirtz, sp. nov.

Fig. 65. Stelis kuijtii Luer, Sp. nov, Section

Fig. 66. Stelis limonensis Hirtz, sp. nov. Fig. 93. Stelis acutula Luer & Hirtz, Sp. NOV.

Fig. 67. Stelis longihirta Luer & Hirtz, sp. nov. Fig. 94. Stelis coralloides Luer & Hirtz, sp. nov.

Fig. 68. Stelis longissima Luer & Hirtz, sp. nov. Fig. 95. Stelis exigua Luer & Hirtz, sp. nov

Fig. 69. Stelis macilenta Luer Hirtz, sp. nov. Fig. 96. is hispida & Dalstrém, sp. nov.

Fig. 70. Stelis micacea Luer & Hirtz, sp. nov. Fig. 97. Stelis juncea Luer & Hirtz, Sp. nov.

Fig. 71. ima Luer, sp. nov. Fig. 98. Stelis poculifera Luer & Hirtz, sp. nov.

Fig. 72. & Hirtz, sp. nov. Fig. 99. Stelis serpens Luer & Hirtz, sp. nov.

Stelis nepotula Luer z . NOV.

Fig. 73. Stelis ortegae Luer & Hirtz, sp. nov. Fig. 100. Stelis tempestuosa Luer & Hirtz, sp. nov.

ety COs

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pre sha attic Aeettl eaba

SUNS. Sere ype oem

STELIS OF ECUADOR a

For descriptions of flowers of Stelis, definitions of a few morphological features

are sir eniee Easeeag of these features of the petals and lip, that are not seen, or

e notc only seen, in other pleurothallid genera, simplifies descriptions and

fe litas wisi cheered Terms used in describing vegetative features ay

rhizomes, ramicauls, sheaths, and leaves) do not differ from other pleurothallids, th

genus of many species being indeterminate when sterile. Except for a rare fades

flowered species, the flowers are produced in a raceme.

Sect. Dialissa Sect. Nexipous

Sect. Stelis

Sect. Humboldtia Sect. Labiatae

Fig. la. SECTIONS OF STELIS

The sepals are variously connate, none being truly caudate. Degrees of conna-

tion are used to define five sections: Dialissa with the sepals connate into a tube or

a deep cup, Nexipous with nearly free lateral sepals deeply connate to the dorsal

sepal, Humboldtia with the lateral sepals deeply connate, Stelis with the sepals

expanded (encompassing a few subsections), and Labiatae with the lateral sepals

connivent and not deeply connate.

petal

= apical margin

transverse callus

Fig. 2a. CENTRAL APPARATUS AND PETAL

The central apparatus is the compact unit composed of the column surrounded

by the petals and lip. The usually transverse petals approach some of those of

Lepanthes, and the three-dimensional lip approaches that of Brachionidium. The

petals are usually concave with the apical or forward margin thickened and flattened

with various modifications. Somewhere across the middle of the petal, there is

often an additional transverse thickening, referred to as the transverse callus

4 ICONES PLEUROTHALLIDINARUM

‘ peo

top view (dorsum) front view

oblique view

Fig. 3a. THE LABELLUM

Although it cannot compare to the complexities of the lip of Lepanthes, the lip

of Stelis, a tiny, intricately sculpted, third dimensional organ, is also one of the most

inaccurately described floral parts of all the Pleurothallidinae. Old descriptions and

crude sketches of boiled flowers made with a hand lens, together with a lack of

common terminology, are incomprehensible and unreliable. Most lips are under two

millimeters in the greatest dimension. All floral parts illustrated herein were made

from flowers pickled while fresh, and drawn to scale while submersed in water.

Three dimensions are employed: length, width, and depth. Illustrations of the same

lip often appear differently depending upon the degree of rotation, or the angle from

which it is viewed.

The dorsum, the top of the lip appressed against the undersurface of the column,

is the length. It extends from the hinged base to the vertical front. The depth is the

distance from the dorsum to the tip of the apex. The width is obviously the distance

from side to side. The side view is more or less triangular, with the dorsum on top;

the front, the distance from the top to the tip; and the back, the hypotenuse. In some

elongated, flattened lips the length becomes the distance from the base to the tip.

The apical margin is commonly thickened, similar to the margins of the ,

The body of the lip is usually thick with a transverse callus across the center,

called here the bar. The margin of the bar usually stretches from the angle between

the junction of the dorsum with the front, to the opposite angle. The center of the

bar usually bears the upper reaches of the glenion, which is found on the front. The

glenion is the round or ovoid, often concave structure commonly seen on the lip in

several pleurothallid genera and infrageneric taxa (especially subsection Macrophyl-

lae-Fasciculatae). It is positioned beneath the rostellum, no doubt playing a part in

enticing a pollinator. In Stelis, it is sometimes absent, and sometimes so greatly

modified izable.

The front (the front of the bar) is usually concave to some degree below the

transverse margin of the bar between the apex and the base. The dorsum, behind

the margin of the bar and beneath the column, often bears one or more calli, which

are swellings along the veins of the lip. The glenion is sometimes seen as the

forward termination of the middle vein. The truncate base of the dorsum is hinged,

often snugly so, to the tip of the vestigial foot of the column.

STELIS OF ECUADOR 5

viscidium

anther cap pollinium

column clinandrium

Spee N rostellum

Pe ———_

petal —~

stigmatic lobe Bs

“———. stigmatic lobe

a column

Fig. 4a. CENTRAL APPARATUS AND COLUMN

The column is short and stout, commonly broader than long. The anther is

apical and deciduous. The pollinia, always two in number residing in the clinandri-

um, are shaped like tear-drops with their pointed tips either free or joined in a

detachable viscidium at the tip of the rostellum. The stigma is greatly variable,

from spherical to transversely oblong beneath the rostellum with the lobes united as

one, to dumbbell-shaped with the stigmatic lobes protruding to either side of the

anther. Sometimes the receptive surfaces are held out on elongated arms, with the

connection very slender, even sometimes lost, beneath the rostellum.

Lip, type-A

akdary "

ne Eee: -

Lip, type-B

Fig. Sa. TYPES OF LABELLUM

Two kinds of lip can be defined, but often they are not easily visualized. Type-A

is subquadrate or wider than long (excluding a sometimes long-acuminate tip of

the apex) with a distinct bar usually with a glenion, and about as deep as it is long.

The dorsum usually bears one, two, three or more calli, that are glabrous or pubes-

cent in varying degrees. Type-B is subquadrate or longer than wide, usually more

deeply concave anteriorly and not as deep as it is long. The dorsum is usually

pubescent, often densely so, or brush-like at the base, and without individual calli.

Type-B is usually seen in section Labiatae.

PACIFIC OCEAN

ICONES PLEUROTHALLIDINARUM

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STELIS OF ECUADOR e

1. Stelis hirtzii Luer, sp. nov.

Ety.: Named for Alexander C. Hirtz, noted ee of hears who collected this species.

P we patye breviter pn sae epalis obtucis papillosis

talis ot is ten sa ,et fsbetio orbiculato ‘ad basim

transverse calloso distinguitur.

Plant small, epiphytic, ascending-caespitose; roots slender. Ramicauls ascending, erect, slender,

1.5-2.5 cm long, proliferating, enclosed by 3-4 tubular, imbricating sheaths. Leaf erect, coriaceous,

narrowly elliptical-obovate, subacute to obtuse, long- -petiolate, 2-3 cm ey: paras a fia lcm

long, the blade 0.6-0.8 cm ps bearers narrowed below into the petiole. Inflor e an erect,

distantly 4- to 5-flowered raceme with 2-5 flowers open denitasconsy. rath cm na asiieg the

nga oa 3-5 cm e from “ annul below ae of the rnunicanl; floral bracts s oblique, acute,

minate, 3 mm long; pedicel mm long; o m long;

light green and pleiod 8 sates within posta ea ate. connate to the middle, reflexed above the

middle, the dorsal sepal ovate, obtuse, 3 mm long, 2.3 mm wide, 3-veined, connate to the lateral ~~

for 1.5 mm to form a —_ deep tube, the — — ovate, oblique, obtuse, 3 mm long, 2.6 m

wide, 3-veined; petals green, membranous, obovate, obtuse, shallowly concave, 1.3 mm long, 1.3 mm

wide; lip purple, biaegeecte elliptical, 1.3 mm lon son 1.6 mm wide, flat with the broadly rounded sides

slightly rincurved, the apex broadly rounded, the base with - saoagan: seine broadly =e A

lamella, he t p column terete, 1.3 mm long, 1.3

wide, th ae + i. + 4 «lh bilobed stigr

Pichincha: forest between San Miguel de los Bancos and ate alt. 1450 ~ 1 so 1984, C. Luer, S.

Dalstrém, T. Héijer & A. Hirtz 9872 (Holotype: MO); between Chiriboga an to Domingo, alt. 1650

m, 31 Mar. 1984, C. Luer, S. Dalstrém, T. Héijer, J. Kuijt & " Piet 9846 (0 same see alt. 1500 m,

28 May 1988, C. Luer & A. Hirtz 13715 (MO); farm road south of Tandapi 1600 m, 1 Mar. 1986,

Luer, C. Dodson, A. Hirtz & A. Embree 12063 (MO); near = eaten: os — m, 25 ae 2001, col-

sere = % Dalstrém & L. Jost, C. Luer 19529 (MO).

Los Cedros Reserve, alt. 1700 m, 21 Jan. 1993, S. Dalstrém, T. Héijer & H. Wanntorp 1708

Das Listens edros Reserve, alt. 1400-1800 m, 2 Mar. 1999, L. Endara 269 (QCA).

?Azuay: above Sayaus, alt. 3200 m, 14 Mar. 1958, DS. Correll E347 (AMES, TEX).

This small species is apparently endemic in northwestern Ecuador. It is distin-

guished by the ascending, proliferating habit and a distantly few-flowered raceme.

The obtuse sepals are connate below the middle into a broad, sepaline cup, and

reflexed above the middle. The inner surfaces are covered with minute papillae.

The petals are membranous, obovate and obtuse. The broadly elliptical lip is flat

with slightly incurved lateral margins. At the base a flat, transverse, broadly round-

ed callus stands erect above the hinge to the short foot of the column.

pa conte — Luer & Hirtz, sp. n

the Latin saccatus, ‘“‘saccate,”’ ett to the bag-shaped, sae tube.

ies haec S. lamellatae Lindl. similis, sed majore, petalis grandibus flabella

lari he differt.

t large, roadside terrestrial (epiphytic), ni and prolific; roots slender. Ramicauls erect,

stout, 4-11 cm long, enclosed by 2-3 loose, tubular sheaths. Leaf erect, coriaceous, elliptical, subacute

to ade + 14 cm long including a petiole less “ie 1 cm long, the blade 2-3 cm a cuneate below

nto the petiole. Inflorescence an erect, congested, secund, many-flowered with many flowers

ech ieee 25-38 cm long pene at the = uncle ca. 10 cm long, ~idented by a spathe 1.5-2

cm long, at an annulus below the apex of the ul; floral bracts oblique, acute, pedicels

mm long; ovary 3 mm long; sepals ¢ dull yellow to ow to yellow-green, — externally, ee

within obtuse, 10 mm lo: wide,

5- veined, connate to the lateral sepals for 2.5 mm to toon a emg whee tube, the lateral ied connate

into a concave broadly obovate, a a with the apices obtuse, 7.5 mm long, 8 mm ea 6-veined;

petals yellow or yellow-green, transversely cuneate- flabellate, Fae concave, 1.5 mm long, 2 mm

— the apex rounded, thickened; lip. yellow or yellow-green, triangular, 0.9 mm long, 1. f mm wide, |

m deep, the apex acute to subacute, concave cep the dorsum ih 3 rounded calli, the base

ance, hinged to the base of t e col umn; t, 1 mm long, 1.5 mm wide, the foot obsolescent,

8 ICONES PLEUROTHALLIDINARUM

re A L 7 1. | ee

Cc ial P Tulcdn and Maldonado, alt. 3000 m, 16

Mar. 1991, C. Luer, J. Luer, J. del Hierro, A. & X. Hirtz 15074 (Holotype: MO); same area, alt. 3400 m,

16 Mar. 1991, C. Luer, J. Luer, J. del Hierro, A. & X. Hirtz 15053 (MO).

More or less inflated below the middle, the sepaline tube of this species is simi-

lar to that of S$. lamellata. Vegetatively, it is characterized by stout, relatively short

ramicauls, often prolific, completely enclosed by two loose, tubular sheaths. A

secund raceme of numerous, simultaneous flowers far exceeds the elliptical leaves.

The petals and lip are out of sight deep within the recess of the flower. The petals

are flabellate, and the lip is triangular with the anterior surface concave.

3. Stelis matula Luer & Hirtz, sp. nov.

Ety.: From the Latin matula, “a chamber-pot,” for an obvious reason.

Inter speci Nexipoi, sepalis i a. on Po tel

. ser a . r ~ r rd b

inter se liberis, petal Se lahall 5 ile is

e LJ

o tr

Plan di epiphytic pit lender. Ramicauls erect, stout, 4-6.5 cm long,

enclosed by 3 loose, spotted, tubular sheaths. Leaf erect, coriaceous, linear-oblong, obtuse, petiolate, 5-

7.5 cm long including a petiole 0.7-1 cm long, the blade 0.7-1.1 cm wide, narrowly cuneate below into a

petiole 0.5-1 cm long. Inflorescence an erect, subcongested, secund, many-flowered raceme with most

flowers open simultaneously, 6-7 cm long including the peduncle ca. 1 cm long, subtended by a spathe 7

mm long, fro lus shortly below the apex of th icaul; floral t bular, acute, 3.5-4 mm

g:P 2. 8; ovary 2 mm long; sepals purple, glabrous, the dorsal sepal te to th

lateral sepals for 3.5 mm and adherent the rest of the way to the tips, forming an inflated, pyriform

from each other, forming a gaping entrance in the under side of the

panded, expanded the dorsal pal sut , 6mm long, 3-veined,

the expanded synsepal 7 mm wide, 9-veined, the apex of the lateral sepals oblique, Giese: petals

purple, transversely oblong, convex on both halves, rounded on the ends, 0.5 mm long, 1 mm wide; lip

purple, subdiscoid, 0.6 mm long, 0.6 mm wide, 0.3 mm

ae obtuse, shortly apiculate, the dorsum with a low suborbicular, minutely pubescent

ulus, base broadly “ate, hinged to the base of the column; column stout, 0.6 mm long, 1 mm

wide across expanded stigmatic lobes, the foot bsol , the anth 1 the bilobed sti ical

lone: ned ie 9 € 2. 1

ea = ae

Owered in cultivation at Ecuagenera, Gualaceo, 17 Apr.

lustr. 20043; above El Pangui, alt. 1900 m, May 1984, S.

Morona-Santiago: near Sinai north of Macas, fl

2001, A. Hirtz 7593 (Holotype: MO), C. Luer jl

Dalstrim & T. Héijer 872A (SEL),

: This species of section Nexipous is unique in the genus with an inflated, pyri-

—_ sepaline tube, with the lateral Tal sepals parted on the under surface of the flower.

€ saccate flower produced is reminiscent of a species of Dresslerella. The

several, purple, globular, nutant flowers are produced nearly simultaneously in a

eme

loose rac S and lip are very small i

sepeline cavity. Pp ery , Out of sight on the back wall of the

4. Stelis zelenkoi Luer & Hirtz, sp. nov.

Ety.: Named for Harry Zelenko, <0 fe

Pet qe dimidi tho

r MEIVOAO Qlitért,

Plant medium in size to large, epi i sa ORG

long, enclosed by a loose, tubular sheath fon baka nos Slender. Ramicauls erect, stout, 5-10 em

. elliptical-oblong, subacute obtus the base. Leaf

contracted below Arai ig es Petiolate, 7-9.5 cm long, the blade 1 5 cm wide

nto a peti 7-8 mm long. Inflorescence an erect. su se a :

flowered raceme with several flowers simultaneously, to 20 cm | beongested, distichous, many-

7? 4 le 5.6 om

oblique, acute, 3-3.5 mm lo se mm long, from an annulus below the apex of the ramicaul; floral bracts

within, glabrous externally, minutely pubescent withi RP pticdane purple externally, olive

the dorcal

STELIS OF ECUADOR 9

small glenion, th dt ie —— column stout, 0.8 mm

long, Imm wide, the foot obsolescent, the anther and the bilobed stigma apical

1400-1800 m, 18 Apr. 2001, A. Hirtz, X. Hirtz, H. Zelenko &

R. Zloch 7573 eo nh om c. pe 30044; above El Pangui, alt. 1400 m, May 1984, S. Dalstriém &

T. Héijer 871 (SEL).

0.6 mm long, 1 mn wide, 0.4 mm deep, th i lar, acute, concave, th gin of the bar with a

Known only from southeastern Ecuador, this species is closely allied to S.

aperta, but readily distinguished by the acute lip. Stelis zelenkoi is characterized by

stout ramicauls about as long as narrowly oblong leaves that are surpassed by the

multiflowered raceme. The shape of the connate sepals is similar to that of S.

aperta, but a little larger. The petals are transversely oblong and convex on each

half. The quadrate lip with an acute apex is more similar to the lip of S. nexipous.

5. Stelis acutilabia Luer & Hirtz, s

Ety.: From the Latin acutilabius, ‘ nani lipo” scinaiach to the ean of the labellum.

pecies haec S. Saag! oe we & = ) Willd. affinis ibus, inflores-

+ Aiffact

centia elongata multi y In latices) i ¢ lah iN p

Plant medium in size, — errs ag roots slender. Ramicauls wie slender, 6-10 cm long,

the middle “third enclosed by a tubul and another 1-2 sheaths at the base. Leaf erect, coria-

ceous, elliptical-ovate, subacute to a pelo 6-7 cm long including a petiole cm long, we _

1.5-2.2 cm wide, cuneate below into the petiole. Inflorescence an erect, congested, distichous, several-

flowered raceme in two ranks, with most flowers open simultaneously, 12 cm long including the aii

cle 2-3 cm long, ea a spathe 12-15 mm long, from an the apex of the ramicaul; floral

bracts eran acute, 4-5 mm long; pedicels 1.5 mm long; ovary | mm long; flowers purple, bilabiate;

sepals glabro ye + al sepal erect, broadly ovate, obtuse, 6 mm long, 4 mm wide, 5-veined, connate

basally to the eine for 2 mm, the lateral sepals connate into a seine ovoid, concave synsepal, 3

mm long, 4 mm wide unexpanded, 6 mm wide expanded, 6-veined; peta ee ee semilunate, 0.9

ng, 1 mm wide, 3-veined, with a broad, thick, rounded apex, concave basally below the transverse

callus; lip thickly triangular, 0.5 mm long, 0.8 mm wide, 0.75 mm deep, the cont cane acute,

deeply concave below the margin of the bar, the dorsum with a low, trilobed callous, the base truncate,

hinged to the base of the column; veers stout, 0.8 mm long, 0.8 mm wide, the foot obsolescent, the

anther and the bilobed stigma apic

Pichinch fi to Domingo, alt. 1900 m, 1 Mar. 1986, C. Luer,

G Dodson, A. Hirtz & A. Embree 12051 eed Aes

This species of section Humboldtia is characterized by a small habit with a

several-flowered raceme with the flowers arranged in two, opposite-facing rows.

The sepals are broad, the synsepal short, more than twice broader than long when

panded. The round apices of the petals are broad and flat. The apex of the lip is

concave, triangular, and acute.

6. Stelis ann-jesupiae Luer, sp. n

Ety.: Named for Ann Lauer Jesup of Bristol, CT, co-collector of this sp

Species citi 5. oe Luer affinis, sed p lo unifloro foliis perparvis longioribus, petalis

crassissimis e mi-orbiculari pl differt

Plant very small, epiphytic, caespitose; me Keine Ramicauls erect, slender, 4-6 mm long,

enclosed by 2-3 thin, tubular sheaths. Leaf ere riaceous, elliptical-obovate, subacute to obtuse, 10-

20% mm long including a petiole 3-6 mm long, on ‘ade 3-4 mm wide, cuneate below into the en

bor 1e by a slender pedunc e 20-30 mm lo ong, with a bract near the middle,

from an annulus below the apex of the ramicaul; floral aa infundibular, acute, 1.5 mm long; olin 1

mm long with a minute filament; ovary 1.5 mm long; flower bilabiate; sepals glabrous, the dorsal sepal

53mm — 6 mm wide unexpande: d, 8- vemned: petals purpl €, thick, oe aity ome rounded at the apex,

1 mm long, 1.3 mm wide, 3-veined, t papillose apex; lip

a suborbicular, 0.6 mm long, 1 mm wide, 0.75 mm deep, shallowly concave anteriorly with the

ee broadly rounded, the dorsum narrow, deflexed, with a rounded callus, hinged to the base of the

mn; column stout, 1 mm long and wide, the foot obsolescent, the anther and bilobed stigma apical.

10 ICONES PLEUROTHALLIDINARUM

7. Ohinchi. Daedill dal ©, a

east of Los Encuentros, alt. 1500 m, 21 Jan. 1989, C. Luer,

J. Luer, A. Jesup & P. Jesup 14027 (Holotype: MO).

This little species, apparently endemic in the Cordillera del Condor in southeast-

ern Ecuador, is one of only a few in the genus known to be naiigt ame The

flower is borne by a slender peduncle that is longer than the leaves. The pedicel

produces a “‘filament,” the aborted terminal bud of a pleurothallid Satis

Produced from near the end of the pedicel, the filament protrudes from the floral

bract.

A : £ a LI, L p pe Fey 4c 1 + = 431

y very small, elliptical

leaves longer than the ramicauls, and a ‘single flower with an erect dorsal sepal

peoei a concave synsepal that is inflated near the base. As with the others, the

| apparatus is on the posterior wall. The petals are thickly lunate. The much

ba lip is cuneate with the flat apex broadly rounded and the dorsum sloping

ard and downward.

7. Stelis anthracina Luer & Hirtz, s

Ety.: From the Latin anthracinus, “coal oa fering to the flower.

Planta mediocris pi prolificans, racemo lent

pal blongo longiore floribus fecundis,

0 dors q tis ot is. netalis rti dib latissime

semilunatis a tah uke lari} 1 4 i es i

pirmbrspoyiabr. iphytic, caespitos fleshy. Ramicauls erect, stout, 6-12 cm

long below to sed by a loose, tubular sheath and 1-2 shorter sheaths at the base

ee coriaceous, cllptica-ebiong. beie ee bee 5 cm long ts a petiole 1-1.2 cm long, the

1.5-2.2 cm wide, c1 ow into erect, subcongested, distichous,

many-Howered raceme, with ower open a tine, 9-11 cm long ‘ictadins tie peduncle 2-3 cm long,

<0 see Ves eae t pathe ca. Ic m lon, ng; floral bracts oblique, inflat-

black, glabrous externally,

pene : are with the he don dorsal sepa raat ovate, 4 mm me. 3 3 mm wide, 3-veined, connate to

ty Conduplicate, ov: se-

te se and — bifid at the apex, 3.3 mm long, 4.2 mm gel Rta 6-veined; Aine

ogg » transy : milunate, shallowly concave, the apex broadly rounded with a thickened

thickly subtr or vie faintly 3 veined, concave below a low, transverse callus; lip dark bywie,

thick a zai the es sie 1 , the epee 0.6 mm deep, shallowly concave anterior! orly with non-

glenion extending from the atime, ick, slightly p een eae tweldeveloped, round

to the base of the column; broadly

column hinged

bilobed stigma apical. ie Velececceac ee

Imbabura: sh 2062. Acosta, alt, 3300 m, 1 May,

spa: 2001, A. Hirtz & X. Hirtz 7741 (Holotype: MO), C

prolific habit, chleas leaves

; ell-deve' :

from a rounded callus on the loped glenion continuous

8. Stelis Luer & Hirtz, sp

SP. nov.

Ety.: From the Lati applanat + Pry ‘

cone insect. thao > Feferring to the expanded, nearly flat flower that is

ous sonnet . similis, sed habitu majore, inflc

bus ex: PEE site pepe ia congesta secunda flori-

Plant small sil aaa caespitose; slender. ae

by 2 tubular sheaths. Leaf capo slender, 1.5-2 cm long, enclosed

Coie fe ete ek cme Sem Sh see. ce

STELIS OF ECUADOR 11

secund, simultaneously several-flowered raceme, to 7 cm long including the peduncle 3 cm long, from an

annulus below the -_ of the ramicaul; floral bracts tubular, 2 mm long; pedicels 1 mm long; ovary 1

mm long; sepals rose with ie apices, glabrous, the dorsal s sepal erect, ovate, obtuse, 7m uns Bering, 4. e

mm wide, 3- ae connate , obtu

shallowly concave synsepal, Sn a 1 long, 5. 51 mm wide, G-star, petals purple, s — with sn

= broadly rounded, thicken m long, 1

wide, 3-veined; li ni thick, ovoid, 0. ut mm long, 0. 6 mm wide, 0. 5 mm dee ep, a anterior

ee shallowly concave, the apex y rounded g the bar with a shallow

glenion, the dorsum with a low, trilo b t te, hinged to the base of the

column; — stout, 0.5 mm se” o 75 mm wide, the foot obsolescent, the anther and the bilobed

Mates apic

Morona-Santiago: Cordillera del Cutuct, new road between Mendez and Morona, alt. 950 m, 17 Jan.

1989, C. Luer, J. Luer, A. Hirtz, S. Ortega, A. & P. Jesup 13987 (Holotype: MO).

This small species of section Humboldtia is apparently endemic in lowland,

eastern Ecuador. It is distinguished by the small habit with the flowers of the

raceme overlapping and facing one direction. The dorsal sepal is ovate and distinct-

ly longer than the shallowly concave synsepal that is held below in the same plane.

The petals are broadly semilunate. The lip is thickly ovate and shallowly concave.

The dorsum does not slope downward as in the closely allied S. memorialis.

9. Stelis bivalvis Luer, sp. no

Ety.: From the Latin bivalvis, “bivalved, ” referring to the ‘‘clam-like,”” bivalved flowers.

Inter species sect. Humboldtiae, habitu parvo, di ngesto folio ellipti ’

convexo lo simili sed concavo, petali is, et labello

brevi semilunato distinguitur.

Plant small, epiphytic ts slender. Ramicauls erect, stout, 3-4 cm long, enclosed by a

loose tubular sheath and another below. Leaf erect, coriaceous, elliptical, subacute to acute, . 7cm

long michaceny a apes 1 —. ree ong, the blade 1.2-1.5 cm wide, cuneate below into the petiole. In-

isti several-flowered raceme 5 cm long, ae the

peduncle ca. 2 cm long, . an a ont s below the apex of the ramicaul; floral bracts infundibular, in-

flated, obtuse, 4 mm long, 4 mm wide; pedicels Z mim kenge rad 1.5 mm long; sepals glabrous, the

dorsal sepal orange- “ie conve roadly rounded, 4.5 mm long, 6 mm wide, 7-

veined, connate basally to the syns bin for 1 mm, Beal eae “light yellow, connate into a trans-

versely ovate, broadly rounded synsepal, 3.5 mm long, 6.5 mm wide expanded, 10-veined, concave

centrally; petals purple on outer half, green on inner om thickly fleshy, transversely semilunate, with

the apex broadly rounded, cxiiakee, ~_—— above the base, 1 mm long, 1.8 mm wide, 3-veined; lip

brown, semilunate, broadly rounded at the apex, very actieally concave above the beasd maargan, « 5

mm long, 1.4 mm wide, 0.8 mm d Meh the bar oe a a“ — with a low,

dorsum, the base truncate, hinged to the base of the column, olumn stout, 0.5 mm long, 1.3 mm wide,

the foot obsolescent, the anther rand the ‘bilobed stigma pe

-Chinchipe: Cordillera del Condor, Paquisha, alt. ca. 1500 m, flowered in cultivation at Ecua-

genera, Gualaceo, 7 Mar. 2001, C. Luer 19797 (Holotype: MO)

This small species of section Humboldtia, apparently endemic in the Cordillera

del Condor, is distinguished by the small, caespitose habit with the inflorescence

shorter than the leaves. The flowers are borne in two ranks in a short, congeste

raceme, with all the flowers facing outward. The flowers are not widely spreading,

the convex, transversely kidney-shaped dorsal sepal held over the shallowly con-

cave synsepal of a similar shape. The petals are thickly reniform. The semilunate

lip is short, three times wider than long.

10. Stelis bolivarensis Luer & Hirtz, sp. n

Ety.: Named for the province of Bolivar in central — where the species was collected.

Spe c S. purpureae (Ruiz & Pav.) — affinis, sed habitu meer -_ ae ooipeee

abrupe oa ea sepalo dorsali 7-nervat

12 ICONES PLEUROTHALLIDINARUM

Plant large, epiphytic, caespitose; roots slender. Ramicauls Stout, erect, 11-13 cm long, mostly

concealed by a tubular sheath from below the middle third and 2-3 sheaths at the base. Leaf erect,

coriaceous, elliptical, obtuse, large, long-petiolate, the blade 11-13 cm long, 5-5.5 cm wide, abruptly

contracted below into a stout petiole 4-5 cm long. Inflorescence an erect, congested, distichous, many-

fl eme witt y fl imul usly, 12-23 cm long including the peduncle 5-6 cm

long, with a spathe 2 cm | g lus below the apex of th icaul; floral bracts infundibular,

oblique, 5-6 mm long; pedicels 3 mm long; ovary 3 mm long; flowers purple; sepals glabrous, the dorsal

erect, ovate, subacute, 11 mm long, 7 mm wide, 7-veined, connate basally 1.5 mm

the lateral epal i deeply ynsepal witl below the lip, 6 mm long, 9 mm

wide, 10-veined; petals transversely semilunate, 1.5 mm long, 2 mm wide, 3-veined, apex thick, broadly

rounded, shallowly concave basally below the transverse callus; lip thick, transversely subtriangular, 1

mm long, 1.4 mm wide, 1 mm deep, concave teriorly at he broadly ied margin, the margin of

the bar with 2 central swellings that flank a glenion, the dorsum mostly filled with a rounded callus, the

base truncate, hinged to the base of the col column stout, 1 mm long, 1.5 mm wide, the foot obso-

lescent, the anther and bilobed stigma apical.

var: forest west of Guaranda toward Balzapamba, alt. 2000 m, 26 Mar. 1984, C. Luer, S. Dalstrém,

T. Héijer, J. Kuijt & A. Hirtz 9733 (Holotype: MO).

is member of the S. purpurea alliance is distinguished by the large habit;

large, elliptical, abruptly long-petiolate leaves: and a congested raceme of relatively

large, purple flowers with conspicuous bracts produced in two ranks. The dorsal

Sepal is noticeably longer than the deeply concave synsepal, which in turn is deeply

concave with a mentum. The petals are protuberant with thickly fleshy apices. The

dorsum of the lip is filled with a single, large callus.

man

11. Stelis brachiata Luer, sp. nov,

Ety.: From the Latin brachiatus, “with arms

Species haec S. odobenellae Luer

&t stigmatis lobis brevioribus differt.

bya erent ripen psn : 2 at roots slender. Ramicauls erect, slender, 5-10 mm long, enclosed

acute, petiolate, 2-3 5 anita ese base. Leaf erect, coriaceous, elliptical-obovate, subacute to

seareeed beans into nee sconce g,> 1-5 em long, the blade 4-5 mm wide, gradually

: an erect, subdense, disti i

Several-flowered raceme, 3-5 cm long including the peduncle wins chan y uremia

fundibul , Obli , acute, 2 lone: + wal A

0.75 mm long: flowers le, bilabiate- = ti » «mm long; pedicels 1 mm long; ovary

tuse, 3 mm long, 3 oa na cea seb Sepal suberect, broadly ovate, ob-

connate into an ovoid, concay, synsepal, 2

»” referring to the elongated stigmatic lobes.

affinis, sed floribus multiminoribus, labello ad basim alticalloso,

sVMenueu at

: : : » O » 0.5 mm , the anteri hallow!

concave _ alow thickening centrally, the poleebiene the dorsum shallowly eee suc an

: . truncate, hinged to the base af ¢

stout, 0.75 mm long, 1.25 min wide across : Bed to ; column

stigma apical. the stigmatic lobes, the lobes elongated, antrorse, the anther

a 1

Zamora-Chinchipe: Cordillera del Condor

J. Luer, A. Jesup & P. Jesup 14959 (Holotype pda Snoventos, alt. 1800 m, 22 Jan. 1989, C. Lue,

blest — ee = ~ ig benella with tusk-like stigmatic lobes, is character-

sie .e ct ’ — habit with a simultaneously several-flowered raceme of

out dic lip is wie The sepals are broadly ovate, the petals thickly semilunate,

lobes are elongated perth. carved eatlts above the base. The stigmatic

odabentis P curved, an arms, but not nearly as long as

12. Stelis coeliaca Luer & Hirtz, sp, nov.

Ety.: From the Greek koiliakos, “cavity ferting to the

Petiolato Subsequandins eco it & Pa) Willd. affinis, sed habitu minore, racemo folio elliptico

differt. nervato, synsepalo profunde , et labello unicalloso

STELIS OF ECUADOR 13

+ 1

Ramicauls stout, erect, 6-7 cm long, mostly

concealed “ a plese att ye below the middle third and 2-3 sheaths at the base. Leaf erect,

coriaceous, elliptical-oblong, obtuse, petiolate, the blade 4-6 cm een 1.8-2 cm wide, contracted below

Infloresce!

into a petiole 2 cm long. rescence 1-2 erect, aet tichous, several-flowered racemes with

most flowers open simultaneously, 4-8.5 cm long includin pti ca. 1 cm long, with a spathe 6-7

mm long, from an annulus below the apex of the mene floral bracts infundibular, oblique, 3-4 mm

long; pedicels 0.75 mm long; ovary 2 mm long; flowers purple, non-resupinate Sepals mi mane 8 ae

cent within, the dorsal sepal erect, broadly ovate

~~ tuse to round at the apex, 8 mm long, 7 mm wide, 7 7-veined, connate basally 15 mm to the cca

y below the lip. Pp; 5mm rae

wide, 6?-veined; petals transversely semilunate, ii 5m mm long, 2 mm wide, 3-veined, shallowly

concave below the thick, broadly rounded aus concave basally below the transverse callus; lip thick,

transversely subtriangular, 1 mm long, 1.3 mm de, 0. 9 mm deep, very shallowly concave anteriorly

above the broadly rounded margin, the margin of tl ar protube rant with a glenion, the dorsum mostly

filled with a broad, rounded callus, the base t te, I i to the b f the column; column stout, |

mm long, 1.5 mm wide, the foot obsolescent, the anther and bilobed stigma ied

Azuay: east of Cuenca, forest above Cumbe, alt. 2800 m, 6 Mar. 2001, C. Luer, J. Luer, A. Hirtz, E

Sanchez & L. Suin 19761 (Holotype: MO).

This handsome member of the S. purpurea alliance from southern Ecuador is

distinguished by the medium-sized habit with one or two several-flowered racemes

more or less as long as elliptical-oblong, petiolate leaves. The sepals are purple,

gaping and minutely pubescent, with the margins of the dorsal sepal revolute. The

synsepal is shallowly concave with a deep excavation below a convexity beneath

the lip. The petals are protuberant with round, thickly fleshy apices. The dorsum of

the lip is filled with a single, large callus.

Stelis coeliaca is related to the much larger S. bolivarensis, but distinguished by

more or less flat flowers with revolute sides of the dorsal sepal, and a deeper exca-

vation of the synsepal.

13. aaa digitata Luer & Hirtz, sp. n

.: From the Latin digitatus, “with ings" referring to the stigmatic lobes.

denc

racemo Hexouso are coe elliptico

oideo supra

petilato bis vel ter longiore, ae lateralibus i in

L . | eee, 5 | - 1 sh hi

ta dictinonitur

medium in orm 2 erp, cae roots slender. Ramicauls crtct, ascending, —

sender, 412 em on ng below, 2-2.5 cm long above, with the r

yipaers with purple, erec “4 ain elliptical, — petiolate, 3-6 cm long va

tee jaw 5- m mm long, the blade 9-14 mm wide, cuneate below into the petiole. Inflorescence an

erect, sublax, distichous, flexible, flexuous, el red raceme, aie 3-6 flowers open simultane-

usly, 5-12 cm Nase including the peduncle 2-3 cm long, from an annulus below the apex of the rami-

oat floral bracts oblique, acute, infundibular, 3.5-4 mm m long; pedicels 1-2 mm long; ovary 1-2 mm

long; sepals red-purple, glabrous externally, cellular within, the dorsal sepal broadly ovate, obtuse,

broadly connate basally, 6 mm long, 5.5 mm wide, 5-veined, the lateral sepals connate into a concave

synsepal, 5 mm long, 7.5 mm wide expanded, ete the apex obtuse, split for 1-2 mm; petals red-

purple, transversely ovate-triangular, 1.25 mm long, 2.25 mm seo 3- veined, the apex broadly o a an

pas concave, with the margin slightly thickened; lip white, discoid, shallowly conca ong,

e, 2 Lanne deep, the apex rounded, minutely alegre dee bar prominently bilobed ae the

bases of the back f the avity, the dors siitdige a low, trilobed,

minutely en a callus, the the base broadly truncate, hinged . iis base of the saben column stout, 1.5

mm —_ 2.5 mm wide, with elongated stigmatic arms, the anther and the stigma apical, the foot obso-

lesc

El Oro: forest along the old road west cE Paccha, alt. 1850 m, 23 May 1988, C. Luer, A. Hirtz, W. Flores,

A. Andreetta & W. Teague 13616 (Holotype: MO); forest a bove Zaruma, alt. 1800 m, 23 Feb. 1986, C.

Luer, J. Luer, A. Hirtz, W. Flores & A. bien 12027 (MO).

This species, known only from southwestern Ecuador, is characterized by a

scandent habit with proliferating ramicauls. The long- -petiolate leaves are surpassed

by loose, flexuous racemes of dark purple, bilabiate flowers. The petals are triangu-

lar while the considerably larger lip is discoid and concave anteriorly with promi-

nent lobes at the base beneath elongated, protuberant, stigmatic arms.

14 ICONES PLEUROTHALLIDINARUM

14. Stelis dromedarina Luer & cag Sp. nov.

Ety.: From the Latin dromedarinus, “like a dromedary camel,” referring to the hump.

poo haec S. - purpureae (Ruiz & hi — tessa sed habitu floribusque minoribus, inflores-

sepalo dorsali et synsepalo latissimo differt

Plant medi epiphytic pit lender. Ramicauls erect, slender, 2-4 cm long,

th erect, coriaceous, siti , subacute, pe-

toe, 5: 5-8 cm mong oe a pel 1 dace mon the blade 1 .2-1.6 cm | wid into the

cm long

including the peduncle less than 1 ty sae annulus below the apex of the ramicaul; Pi bracts

oblique, acute, 3-4 light yellow-green, bila-

5? b fen,

a, T +

, broadly ovate, obtuse, 4 mm n long, 4 mm wide, shes

— anc to the synsepal for 1 mm, the lateral sepals connate into a transversel

oid, onca

ynsepal, 2.5 mm long, 6 mm wide expanded, oo petals thin, transversely oblong, shallowly

itn with the obtuse, 2 mm long, 3 mm wide, 3-veined: lip thickly su :

— a wide, 1 1.5 mm deep, concave within the rounded a apex, the bar with a high, ido eae

ike callus. t the +6

hump,” the base truncate, hinged to

the base of the column: column : stout, 1.5 mm long, 2 mm wide, “the foot obsolescent, the anther and the

bilobed stigma apical.

Carchi: between Tuledn — El Carmelo, alt. 3200 m, 6-7 A i

posaageayeag m, pr. 1985, C. Luer, J. Luer, A. Hirtz & W.

several-flowered raceme shorter than the leaf. The sepals are broad, the synsepal

thicke

Proportionately sted hump-shaped callus fills the center.

15. Stelis embreei Luer & Hirtz, Sp. ni

a se in honor of the late Al Embree, 2 an n old friend, co-collector of this species.

successi _ - eas affinis, sed habitu floribusque majoribus, inflorescentia laxe fractiflexa

to differt. : ig F quilongo, et sepalo dorsali 5- ad 7-nerva-

Plant small, epiphytic, cocspita a:

by a tubular rad and ano’ slender. Ramicauls erect, slender, 1-1.5 cm long, enclosed

long including apie 1-15 base. Leaf erect, coriaceous, broadly elli ptical, obtuse, 3-5 cm

exw, oe the blade 1.3-1.6 cm wide, cuneate below into the palinke In-

peduncle 25.3 cm long, from ; annulus below the ively ep ti ered raceme, =e cm long including the

Hgue,

the dorsal : sepal convex, us, cellular within,

connate basally to the = erect : beans oat, obtuse, 8 mm long, 7mm a wide, 5- to 7-veined,

<a. 6 ne 4.5 mm wide unexpanded, 9 mm w: de

, le, with the a x broadly rounded, cellul crea

center, concave above the eee | rou cellular, thicke

— thick, 75 mm long, 1.2 nisershplestern phiwheen sve ee eee ee ees

tral alone the Rese truncate. | » the dorsum narrow with a single, pubescent, cen-

wide, the foot Oot obsolescent, the anthes and ane ta oF the column; column stout, 0.5 mm long, 1.3 mm

Zamora-Chinchipe:-

Hirtz, W. Flores aA. Entree 1509 — a m, 20 Feb. 1986, C. Luer, J. Luer, A

re : ? (Holotype: MO); south of Zamora toward Zumba, alt. 1100 m, 3

. 1990, 5. Dalstrtn & 1. Ae fia north of Yan anzaiza, alt. ca. 1300 m, 6

Sf Ss mis taba

tiago: Rio Urpano

pais 13919 (MO) Candies ge Macas, alt. 1100 m, 15 Jan, 1989, C. Luer, J. Luer, A. Hirtz, A.

Luer, endez and

J. Luer, A. Hirtz, mA. & P.denwp 1397S eM and Morona, alt. 950 m, 17 Jan. 1989,

The sepals of this little s.

Humboldtia: a d tatty ay similar to many others in section

orsal sepal standin

8 erect ove

related to the larger S. tessmannii hin Peru, and the s oe ae hes a .

iid Ae Se

STELIS OF ECUADOR 15

nana of Ecuador. The raceme of S. embreei is lax and fractiflex, about as long as

the leaf; the dorsal sepal is incompletely 7-veined; the petals are thickened and

semilunate; and the lip is discoid, not convex, with the apex rounded, and with the

base truncate with a pubescent callus

16. Stelis fissurata Luer & Hirtz, sp. n

Ety.: Fro: meena spaptiginestg “fissured,” refering tthe ip

—_ 7 2: ape, Haase & Pav.) Pers. pl jue parvis, foliis

disticho breviorbus, sepals initits pubescentibus, p inibus i tis minut verruculosis,

teralihn lati 4 4: £, in Oe a 5 ea

superne pubescenti.

Plant small, epiph if Ramicauls erect, slender, 2-5 cm et sgn by

a tubular sheath from oe the middle and another sheath at the base. Leaf erect, coriaceous, narrowly

elliptical, acute, 2-7 cm long including an indistinct petiole ca. 1 cm long, the blade 46 mm oe ae

ally narrowed below into ihe petiole. bai flor’ escence an erect, subdense, distichous, several-flowered

raceme pen 6-9 cm 1 gth pedun uncle ca. 1

cm long, from an annulus below the apex of the ramicaul; floral bracts oblique, ye mm long; wipes 1

mm long; ovary 1 mm long; flowers bilabiate, perpendicular labrous

externally, shortly pubescent within, the dorsal sepal creat, OnIOne. obtuse, convex, 3. 6 mm tong 2mm

wide, 3-veined, connate less than 1 mm hs the synsepal d, deeply

concave synsepal, 2.5 mm long, 4 mm wide expanded, 2 mm wide acapeaied 8-veined; petals tr ans-

versely ovate-flabellate, concave, 0.8 mm a , 1.2 mm wide, the margins thick, minutely verrucose, the

base broadly truncate; lip thick, Pepa narrowly rounded at the apex, with the lateral margins

os angled, 0.9 mm long, 0.9 mm wide, 0.9 mm deep, the anterior surface deeply fissured, =

dor. m witha sang pubescent callus, the base truncate, hinged t to the base of the column; colum

stout, i" 8m

Carchi: above Maldonado toward Rio La Plata, alt. 2200 m, 11 Jan. 1993, A. Hirtz 5869 (Holotype:

MO), C. Luer illustr. 19477; plateau above Chical, alt. 1800 m, A. Hirtz 5857 (MO).

This species of section Humboldtia is apparently restricted to wet forests of

northwesternmost Ecuador. It is distinguished by the small, caespitose habit;

narrowly elliptical leaves; a several-flowered raceme of small flowers perpendicular

to the rachis in opposite ranks; sepals pubescent within; and a subquadrate lip with

obtuse, lateral margins, and a deep central fissure.

17. om — Luer & Hirtz, sp. n

Latin flavus, “yellow,” “te the color of the flowers.

is ter species sect Humboldtiae, planta gr

paca Longer sepalis flavis multinervis, — a —— subacuto, aa obtuso, petalis

rassissimis lunati

Plant large, epiphytic, a roots slender. Ramicauls erect, — athe cm _ with a loose,

tubular sheath from below the middle and another 2 sheaths at the base. Leaf erect, coriaceous, ellipti-

cal, subacute, long- _ ate, 10- - cm in including the petiole 2- 22 5 cm ie tie blade 2.7-3.3 cm

wide, cuneate below into the petiole. Inflorescence an erect, loose, distichous, many -flowered raceme

with many flowers open sinless —" 24 cm long including eh peduncle c a7 cm long, witha

sate 1.5-2 cm lon e apex of the ramicaul; floral bracts sete, oblique, 2-3

ong; pedicels I- ‘1. 5 mm long; ovary ine m long; sepals yellow, the dorsal sepal ne ovate,

eens 15 mm long, 7 mm wide, 5(7)-veined, state to the synsepal for 2 mm, the lateral sepals

connate ed a concave, broadly ovate synsepal, 11.5 mm long, 11 mm w wide, 10-veined; an yellow,

; 1 : :

c e

cakes ‘fleshy thickened, broadly rounded, concave basally below a sharp, broad, transverse callus; lip

agin rien boblong, the apex broadly rounded, subretuse, 1 mm long, 1.3 mm wide, 0.7 mm deep,

the an surface shallowly concave, the bar with a slender glenion, the dorsum concave with 3 calli,

se base t truncate, hinged to the base ua the column; column stout, | mm long, | mm wide, the foot

gma apical

Carchi: between Maldonado and Chical, alt. 1300 m, May 1992, A. Hirtz 5784 (Holotype: MO), C. Luer

illustr. 20063.

16 ICONES PLEUROTHALLIDINARUM

This large species, from a relatively low altitude in northwesternmost Ecuador is

similar to the vegetatively larger S. titanica from much higher altitudes. The flow-

ers are large and yellow with the elliptical dorsal sepal erect above the concave

synsepal. Most remarkable are the large, thickly fleshy petals that overlap above

the column and meet below under the labellum. The lip is transverse with a slender

glenion, and concave dorsally with three calli.

18. Stelis flexilis Luer & Hirtz, sp. n

Ety.: a Latin — Asian i wits to the rachis.

Inter sp ~ Hi di perp , racemo 0 laxo flexuoso successivo folio elliptico

tiolat giore, sy cu »fund »p is et labello ovoi-

deo apice rotundo distinguitur.

Plant very small, epiphytic, caespitose; roots slender. Ramicauls erect, slender, 4-6 mm long,

enclosed by two close, tubular sheaths. Leaf erect, coriaceous, narrowly elliptical, acute to subacute,

petiolate, 12-20 mm long including the petiole 3-5 mm long, the blade 3-4 mm wide, narrowly cuneate

into the petiole. I sohisipong sda many-flowered raceme with 2-3 flow-

ers open simultaneously, to 10 cm long including the le 3-4 cm oe from an annulus below the

a oe of the rami icanl, with an insignificant spathe floral ae gripe 1.5 mm

;o vary mm

sli eo concave, 3 J mm ty 3 mm wide, 3- — connate 1 mm asally to the synsepal, the lateral

deeply concave sande with an obtuse sear

xpended, 6-veined; relatively thin, transversely

x only ound rounded, 0 Gn mm long, 1 mm wide, 3- veined; ‘lip tick,

Si obscure, lngiendinal calli, with a a minute glenion o n the marg: n

bc fe ol a rounded callus, microscopic ally pubescent, = base rye hinged to the

a oe column stout, 0.6 mm long, 0.6 mm wide, the foo nther and the

Zamora- ae Cordillera del Condor, east of P.

uisha, alt. 12

Luer, A. Hirtz, W. Flores & A. Embree 11929 (Holotype: Mi M MO). ee een cee t

This tiny, caespitose species of section Hum

lera del Condor. The habit is similar to that

ly thin and shallowly concave. The

and rounded at the apex, with a rounded callus o

19, _Stelis lorianii Luer uer, Sp. nov.

Ety.: Named in honor of Flori re ren who collected this species.

Species haec

cum callo nie a Rchb.f. affinis sed petalorum cri x oe eek in doi ub

. shana small, epiphytic, caespitose: roots slender. Ramicaul

reat including a petiole 3-10 mm long, the blade $9 rao eorme, subacute, petiolate, 10-35

petiole. Inflorescence uneate i

blade 5-9 mm wide. narrow]

eek: 5 c low into the

slender peduncle 1-2 cm 2-3 f pevees Faceme, more or less erect, 3-5 cm long including the

floral bracts in: — ly, from below the ramicau

Ol ; 5 : ;

flowers translucent hite, bi 2 mm long, 2 mm wide; pedicels 1 mm long; ovary 1.5 mm long;

obtuse, connate 1.5 mm basally to the gibrous, the dorsal se adly ovate or orbicular.

connate into a deeply ne, oned cease sa long, 5-6 mm wide, 5-veined, the

Pears transversely ovate, the — iaeae long, 6-7 sa 6- eined; petals =

ticulate, 7 ann ay

vores call ! fringe, 2 mm wide, 3. veined, acd with concave below the trans

stent es ka

shears

STELIS OF ECUADOR 17

a indistinctly thickened, the bar _ an = bifid, conical callus, the corsum with a small, Lament

eoeiey arms, | mm long, 1 mm wide, i t, th I d the 2 d sti

Zamora-Chinchipe: San Francisco Biological Station between Loja and Zamora, alt. 2200 m, 27 Jan.

2000, F. Werner 218 (Holotype: MO), C. Luer illustr. 20051.

Apparently rare, this species is known only from the original collection in

southern Ecuador. It is closely allied to the relatively frequent S. dactyloptera, but

differs in having smaller, white flowers borne successively in a suberect raceme.

The crest of the petals is proportionately smaller than that of S. dactyloptera. In-

stead of orbicular, the lip is trilobed with the oblong, basal lobes overlying the basal

half of the petals. Centrally, an erect, grooved callus reaches behind the rostellum

positioned above.

20. ote — Luer & Hirtz, s

e Latin galeola, “a bucket ‘aon like a helmet,”’ referring to the concave synsepal.

see species sect. Humboldtiae, habitu parvo, foliis spathulatis minute verrucosis, racemo elongato

J r =

Plant small, epiphytic, caespitose; roots slender. Ramicauls erect, stout, 5-8 mm long, enclosed by

2 tubular sheaths. Leaf erect, coriaceous, smooth beneath with the dorsum minutely verrucose,

spathulate, ye sno a 20 mm — including a petiole 4-6 mm “at the blade 5-

8 cuneate below into the petiole. Inflorescence a lax, flexible, flexuous, successively flow-

ered raceme we § 12 cm long, satiate the Shiels a ie long, with 1 or 2 flowers open simultaneous-

ly, from an annulus below the apex "of the ramicaul; floral bracts siesta, obtuse, 1-1.5 mm long;

pedicels 0.75 mm long; ovary 1 mm long; flowers nig in pendent position; sepals glabrous,

the nr sepal purple, convex, ovate, obtuse, 6 mm long, 5.5 mm wide, 3-veined, connate basally to

the synsepal for 1 mm, the lateral sepals connate into an ovoid, dtm, deeply concave synsepal, dark

6 long, 6 i

transversel Ilular, thic

center, concave above the base, 0.8 mm long, 1.4 mm wide, 3-veined; lip thickly triangular, 0.5 mm

long, 0.9 mm wide, 0.6 mm deep, the bar taller than the sides, with a narrow glenion, the apical margin

thick and rounded, shallowly concave above, the dorsum w ith a low, rounded, eae callus at the

» the base tru truncate, hinged | to ari bases of the oman; coleman st tout, 0.5 mm long, 0.8 mm wide, the

Zamora-Chinchipe: Cordillera del Condor, forest east of re alt. 1200 m, 20 May 1989, C. Luer,

A. Hirtz, W. Flores, A. Andreetta & W. Teague 13496 (Holotype: MO).

This small species of section Humboldtia, apparently endemic in lowland, east-

ern Ecuador, is distinguished by the small, caespitose habit with spathulate leaves

that are minutely verrucose on the dorsum. The raceme is flexuous and flexible, the

flowers becoming pendent as the raceme lengthens. The synsepal is unusually

deeply concave. The petals completely enclose the column and lip. The lip is

characterized by a tall bar higher than the sides.

Fak Les luteria Luer & Hirtz, sp. n

From the Greek louterion, “‘a little Tatkocls % —a to the concave a

ureae (Ruiz & Pav. m brant affinis, sed ij ;: b

sepali dorsalis Sects revolutis, synsepalo tral petalis crassisimis et labelli

apice rotundo distinguitur.

Plant medium to large, epiphytic, — roots slender. Ramicauls erect, stout, 11-16 cm long,

enclosed by a tubular sheath on the middle third 2-3 sheaths at the base hes erect, coriaceous, ellipti-

cal, subacute, subpetiolate, 10-13.5 cm long a a Sapa gg petiole ca. | cm long, the blade 3-5

cm wide, cuneate below into the petiole. Inflorescence an erect, ack distichous, successively

-flowere

cle

8-10 cm long, with a spathe 1.5 cm long, from an annulus below the apex of the ramicaul; floral bracts

tubular, ae 4mm Jong below to rosie ye — above; pedicels 3 mm long; ovary 3 mm long; flowers

bilabiate g hortly ne aa within, the dorsal sepal erect, " oblong-

18 ICONES PLEUROTHALLIDINARUM

ovate, with the sides incurved, obtuse, 13 mm long, 5 mm wide, unexpanded, 7-veined, connate 2 mm to

the synsepal, the lateral sepals connate into an oblong, bifid synsepal, 9 mm long, 7 mm wide, unexpand-

ed, 8-veined, with a well-demarcated cavity in the center; petals purple, thickly reniform, cellular pubes-

cent, with apex broadly rounded with ill-defined margin, 1.3 mm long, 2.25 mm wide, 3-veined; lip

purple, subovoid, thick, 0.8 mm long, 1.25 mm wide, 1.1 mm deep, the bar shallowly teriorly

above the broadly rounded apex, with a superficial glenion, the dorsum with a low, broad callus, the base

truncate, hinged to the b f the col column stout, 1 mm long, 1.5 mm wide, the foot obsolescent,

iil

the anther and the bilobed stigma apical.

Morona-Santiago: without specific locality, cultivated at Ecuagenera, 7 Mar. 2001, C. Luer 19806

(Holotype: MO); new road between Mendez and Morona, alt. <1000 m, 19 Jan. 1989, A. Hirtz, A. An-

dreetia & S. Ortega 4131 (MO).

This distinctive species from southeastern Ecuador is characterized by a large

habit with large, elliptical, subpetiolate leaves that are far surpassed by a many-

flowered raceme. The purple, pubescent flowers are large and gaping with the

margins of the erect dorsal sepal recurved. The center of the oblong synsepal is

concave while the outer halves of both lateral sepals are convex; the petals are thick

and reniform without transverse Margins; and the lip is shallowly concave and

22. Stelis nycterina Luer & Hirtz, sp. nov.

Ety.: From the Greek nykteris, ‘‘a bat,” referring to the ear-like petals.

Species haec S. slobiflorae Rchb.f. affinis, sed petali jit i

distinguitur, tas

—J r

Plant small, epiphytic, caespitose; roots slende i

by two tubular sheaths Leaf erect, cori = Remi cree, slender, 5-10 mm long, enclosed

mm long including a petiole 5-10 mm long, the blade 6-8 mm

ole. Inflorescence a jax or subdense, several-flowered raceme mor

lar, > sepal erect, broadly ovate or orbic-

bes a : — a basally to the synsepal, 7 mm long, 5.5 mm wide, 3-veined, the lateral

: ply pei: ovoid synsepal, 5 mm long, 7 mm wide, 6-veined: petals purple,

cate with the incurved and irregular, 2.5 mm long, 1.5

mm wide at vei

: Y Concave at ‘ * ani

0.4 : : le, subdiscoid,

thictnneia Bt sheng Imm deep, shallowly concave anteriorly with the pittays sh th i

reflexed, truncate — oe em, the rescore with an erect, rounded, pubescent callus, the

mm long, 1 mm wide, the foot obsolescent. the a, er ee with short, stigmatic arms, |

ome apital,

Morona-Santiago: near Lim6n, cultivated at

, : : :

ar = ad pet lites and fringed in S. dactyloptera and S. florianii.

0 ers, the lip of S. nycterina is shallowly discoid with a tall,

™ MUIOUITL,

notched hb 3

Pecies haec S. purpureae (Ruiz & Pay :

dorsali plus minus Te pared. affinis, sed racemo folio acuto multilone;

do differt "reflect, petalis parvis partiale post labellum, et anon fae aioans ner

Plant medium in size to lar. Seay

8¢. epiphytic, densely itose-

slender, 10-17 cm long, enclosed ly caespitose; roots s}

Leaf erect, cori Z 4 close, tubular ; ;

ender. Ramicauls erect,

narrowly Mipticl, acute, petiolate, -11 em lone below at the base.

. and 2-3

including a petiole 2-2.5 cm

STELIS OF ECUADOR 19

peau sage blade 1.3-1. 8 cm wide, cuneate below into the petiole. Inflorescence an erect, subdense, dis-

many- -flowered rac raceme, 20-30 cm long including the peduncle 3-5 cm sig =

ote tia dibular, oblique, acute, 4 mm lon: g, 4 mm wide;

long; ovary 3 mm long; sipsen _ yellow suffused with sel to dark purple, glabrous, — rio

sepal more or less reflexed from the base, broadly ovate, obtuse, 9 mm lon, g, 6 mm wide, 5-(7-)veined,

connate oe to the synsepal I for 1 mn, the lateral sepals connate — a ans versely ovoid, concave

synsepal, 7 mm long, 9 mm wide, 5 mm deep, ca. 12-veined; petals white to purple, transversely ovate,

shallowly concave, with the apex broadly rounded, thickened, with an indisunet margin, 0.5 mm long,

1.25 mm wide, 3-veined; lip purple, protuberant, thickly subovoid, 1 mm long, 1.25 mm wide, 1 mm

deep, shallowly concave anteriorly with the rounded apex thickened, the wai ‘of the bar noe a am

glenion between a pair of swellings, the dorsum with a low, central, trilobed callus, the bas

hinged to the base of the column; column stout, 1.5 mm long, the anther and the bilobed sti pe paper

the foot obsolescent.

ee eas ae forest between G Limon, alt. 2150 m, 14 Jan. 1989, C. Luer, J. Luer, A.

P. &A. Jesup & S. Ortega 13893 se tes MO); same data, flowers of different color, C. Luer

et wen "13891 (MO).

This species of section Humboldtia is characterized by a slender, densely caespi-

tose habit with ramicauls longer than long-petiolate, acute, elliptical leaves. The

raceme far exceeds the leaf. The erect dorsal sepal sometimes reflexes above the

base. The synsepal is deeply concave with a chin protruding posteriorly. Within,

the floral parts are proportionately small, especially the petals that are partially

behind a twice larger, protruding lip with a bitumid bar and a central glenion.

24. Lo —— ie & Hirtz, sp. n

patens, “open, ee Puede to the flowers

ns er species sect. Humboldtiae eae parva, racemo laxe luo folio a petiolato plus

minisve duplolongiore, sepalo ‘0 do rsali | q , petalis semilunatis

# lahat e s: A ae ge

Plant small, on caespitose; roots slender. Ramicauls erect, slender, 1-2.5 cm a vee

by two loose, tubular sheaths. Leaf erect, coriaceous, narrowly elliptical, acute to subacute te, 2-

4cm long incl uding the ae 0.5- vie cm =. — — ae “a ae sine wide, narrowly cuneate =e into the

le. loose raceme with 3-4 flowers open simul-

petiole

taneously, 6-13 cm long including the nek oe cm - ong, on an annulus below the apex of the

micaul, with an insignificant spathe; floral bracts oblique, inflated, 2 mm long; pedicels 1 mm long;

ovary | mm long; flowers bilabiate; sepals pecten be rple, glabrous, the dorsal sepal erect, ovate,

btuse, 5.5 mm long, 4.5 mm wide, 5-veined, connate mmm basally | to the synsepal, the a ee

ed; petals

lunate, the rounded apical margin thick, well demarcated, shallowly concave basally below the eieitee

callus, 1 mm long, 1.5 mm wide, 3- veined; lip green, » thick, tri tiangular-sebqoa uadrate, 0.5 mm long, | mm

ide, 0.75 mm deep, th d, the bar very shallowly concave anteri-

— slightly higher than the sides, with a superficial glenion, the dorsum with a single, rounded callus,

cally pubescent, the base truncate, hinged to the base of the column; column stout, 1 mm

As mm ae, the foot obsolescent, the anther and the bilobed stigma apical.

ioe tg eg west of Macas, new road to Guamote, junction of Rio Colombo and Rfo Upano, alt.

1600 m, ar. 2001, C. Luer, J. Luer = A. Hirtz1 =o (Holotype: MO); north of Gualaquiza toward

San Juan = alt. 1250 m, 27 Apr. 1 985, C. Luer, J. Luer, A. Hirtz & W. Flores 10976 (MO); ae

lera del Cutucé, between Mendez and Morona. alt. 950 m, 17 Jan. 1989, C. Luer, J. Luer, A. Hirtz e

Napo: Rio Jatu tunyacu, above Shandia, alt. 650 m, 12 Aug. 1991, A. Hirtz & S. Ortega 5375 (Holotype:

MO), C. Luer illustr. 20059; west of near Tena, collected by C. Luer et al. 26 July 1975, cultivated at

SEL, Jan. 1976, C. Luer 371, 1043 (SEL); west of Tena, Rio Jatunyacu, alt. 600 m, 21 Feb. 1982, C.

Luer & A. Hirtz 6909 (SEL);

Tungurahua: Rio San F isco bet Bajios and Puyo, alt. 1700 m, 16 Feb. 1963, C.H. Dodson &

L.B. Thien 2264 (SEL).

Zamora-Chinchipe: Cordillera a Condor, east of Paquisha, alt. 1200 m, 23 Jan. 1992, C. Luer, J. Luer,

A. Hirtz, A. & P. Jesup 16122 (M

This small species of section Humboldtia occurs frequently on the eastern de-

clivity of the Andes of Ecuador at relatively low altitudes. The caespitose habit of

elliptical leaves borne by short ramicauls is similar to that of many other species of

20 ICONES PLEUROTHALLIDINARUM

the genus as well as many species of Pleurothallis. It is characterized by a loose,

several-flowered raceme of resupinate flowers about twice as long as the leaves.

The dorsal sepal is ovate, obtuse and five-veined. The lateral sepals are connate to

the apex into a concave synsepal. The petals are semilunate with a broad, thick

margin; the lip is subtriangular, shallowly concave and rounded at the apex, with a

relatively large, rounded callus on the dorsum.

eel spreygng Luer & rit sD. nov

“Bay: From the Latin patinarius, “ shallow pan,’ Serie tay at

ee haec S. purpureae (Ruiz ahd) Willd affinis, d fl 1

Ash re

tihy

is, et labello propor-

4 lH ey

Plant medium to large, epiphytic, caespitose; roots slender. Ramicauls en stout, 2-15 cm long,

enclosed by a loose, tubular sheath on the middle third and 2-3 sheaths at the base. Leaf erect, coria-

blade rie gine subacute to obtuse, petiolate, 4-13 em wie ees the its 1 is 5 cm long, the

many-flowered . . Inflorescence an erect, subcongested,

Gi ered raceme w loves ce

peduncle 3-5 cm long, neeped open simultaneously, 15-18 cm long including the

witha snes : 5-1.5 cm long, from an annulus below the a: of the ramicaul;

floral bracts oblique, acute, 6-9 mm lon below to 3 ‘ ae

long; flowers purple, non-resupinate. one Zig = long above; pedicels 2mm long; raves Bice

bs git long, 5-6 mm wide, S-veined (sometimes with accessory veins), connate 2 mm to the synsepal,

wide, 8-v eined: petals ce —_— ideuss shallowly concave synsepal, 4-6 mm long, 6-8 mm

Had 2-2.3 mm wide, 3- eined, cours the lip, 1.5 mm

: igs below the thickened, broadly rounded apex, and concave basally

ne dy me ee ays mgs mw

tru hinced ta the k Cat 1

wide, the foot obsol Sh Ati enone ye in; column st 0.75 mm long, 1.75 mm

apical.

Aawed W healer sl cabanas alt. 1850 m, 28 Mar. 1985, C. Luer, J. Luer,

Napo: El Chaco

sip Cu 50st 198.4, Hine & A Andreetia 1953 (MO), \Seagrd ese abne7.

Hirtz & W. Flores lee (MO): tare spool te south of Loja, alt. 2500 m, 21 Mar 5, C. Luer, J. Luer, A

Gualaceo, 7 Mar. 2001, C. Luer 1981 6 (MO). Loja and Zamora, flowered in salaries at = petal

Thi

‘3 s species i is distinguished from the other members of the section Humboldtia

middle-sized, flat, gaping flowers with a transverse, s

that gives the flower an appearance si

Species are variable with broad,

ramicauls bear a tubular sheath

hallowly concave synsepal

imilar to section Stelis. Vegetatively, the

petiolate leaves of various sizes and shapes. Th

a By: Pom he a ee Dalat Dalstrém, M, SP. Nov.

__ Inter species sect. Humboldtiae wide petals,” referring to the broad, flat petals.

giore, sepalo to: ‘

dorsali 7 ovato, synsepalo profunde conca — wake is nian multifloro folio Aomaayatorel lon

oe petali

cha dy one td ana Sau io eet, son, 2.5-4

below into the petiole. Inflone sn tin® #Petiole 1-1.5 cm long, the blade 18.2.3 cm pide onsen

long including the sieikeiene 3-5 cm long, sco sda congested, many-flowered raceme 12-20 cm

flowers open Open simultaneously, from an annulus below

STELIS OF ECUADOR 21

Tk Ce |

g; pedicels 2 mm long; ovary 2 mm lon

en glabrous, the dorsal sepal purple, sony case soba, 8 mm | long, 7.5 mm wide, 5- veined,

connate yellow 1,6m

long, 9. ; mm wide, 8- veined; petals purple, broadly ovate-suborbicular, with the surface flat, hie,

1.6 mm long, 2 mm wide, 3-veined, narrowly concave at the base; lip purple. thickly triangular, 1 mm

long, | mm wide, 1 mm deep, shallowly concave anteriorly with the apex acute with an acute apiculum,

a = of the bar mores witb the sides, with a minute, beater glenion, ad dorsum with 3 rounded

2g foot; column stout, | mm long, 1.2 mm wide,

. 4 +t ea San pe. ee |

EI Oro: north of Salvias, alt. 2000 m, 4 Feb. 1993, 5. Dalstrom & Tr. Hoijer = pei MO), C

ne illustr. 20061. road between Q m, 2 Feb. 1978,

C. Luer, J. Luer & A. Hirtz 2434 (SEL).

This species of section Humboldtia is apparently endemic in southwestern

Ecuador. It is characterized by the densely caespitose habit and a densely many-

flowered raceme surpassing an oblong leaf. The dorsal sepal is broadly ovate and

the synsepal is deeply concave. The petals are proportionately large, round and flat.

The apex of the lip is sharply acute.

ZT. Eien preclara Luer & — sp. n

rom the Latin preclarus, “very pretty,” atein to the —s —

ow see é purpureae ue & Pav ) Willd affi inis, sed folio grandi elliptico abrupte longip

lato, sepalo dors p d avo, petali i ilunatis et

labello antice concavo differt.

Plant large, epiphytic, eongae roots slender. Sg stout, erect, 8-15 cm long, with a close,

tubular sheath above the middle and 2-3 sheaths at the base. Leaf erect, coriaceous, a ciel. obtuse,

large, short- oe the blade 8- 10 cm long, 4-5 cm wide, pic contracted below into a stout petiole

ca. 1.5 cm long. Inflorescence an e erect, ‘conges oe distichous, many-flowered raceme with many flow-

ide open simultaneously, 15-18 ¢ the peduncle c ca. 10 om long, with a — 2 cm long,

of th oblique, 4 mm long; pedicels

3 mm long; ovary 2.5 m a flow ers ma poet — externally, minutely sie within,

the dorsal sepal erect, siially ovate, obtuse to rounded at th x, 9 mm long, 8 mm wide, 9-veined,

connate 2 mm basally to the synsepal, the lateral sepals mae into a =a ioe sc with a

mentum — the lip, 7 mm long, 9 mm wide, 10-veined; petals transversely semi Re ate, 1.2 mm long,

om e, 3-veined, apex thick, broadly rounded, shallowly seg basally below the transverse

wile nm thick, Same 1 mm m lon ng, 1.5 mm wid e, 1. 25 mm deep, concave ed above the

“ipo ; APEX, uigher margins 0 the lip, with a long, narrow glen-

ite call , the base truncate, hinged to the | base of the column,

1 £,

aii stout, 1 mm “ie 1 25 mm wide,

Tungurahua: slopes of Volcén Tungurahua, alt. 2400 m, Dec. 1983, A. Hirtz 1475 (Holotype: MO), C.

Luer illustr. 20064.

This member of the S. purpurea alliance is distinguished by the large habit,

large, elliptical, abruptly petiolate leaves; and a congested raceme of relatively

large, purple flowers produced in two ranks. The dorsal sepal is broadly elliptical to

suborbicular with nine veins; the synsepal is deeply concave with a mentum. The

petals are semilunate; the lip is triangular, subacute and concave with a narrow

glenion.

28. ae regina Luer & Hirtz, a. nov.

.: From the Latin regina, “the queen,” referring to the si er ae

pr haec S. purpureae (Ruiz & Pav. ) wale. affinis, sed floribus saslittenaieatiies, petalis grandis

Plant large, epiphytic, caespitose; roots slender. Ramicauls slender, erect, 13-20 cm long, enclosed

by a loose, tubular sheath near the middle and 2-3 other sheaths at the base. Leaf erect, coriace —

elliptical, cat with the apex acute to subacute, 10-14 cm long auletisy the petiole 2-2.5 cm long,

the blade 2.5-3.5 cm wide, cuneate below into the petiole. jorescence an erect, deactoms,' ae

several-flowered raceme with most flowers open simultaneously, 15-18 cm long including the peduncle

22 ICONES PLEUROTHALLIDINARUM

ca. 5 cm long, subtended by a slender spathe ca. 2 cm long, from an annulus below the apex of the

icaul; floral t inflated, infundibular. oblique, acute, 5-6 mm long; pedicels 3-4 mm long; ovary 4

mm long; flowers nutant, bilabiate; sepals white, suffused with pink, glabrous, the dorsal sepal erect,

ovate-triangular, subacute, 13 mm long, 10 mm wide, with 7 primary veins and ca. 4 accessory veins,

s or 3 mm, the lateral sepal into a broadly ovate, deeply concave synse-

pal, 11 mm long, 13 mm wide unexpanded, 10-(15-)veined; petals light yellow, transversely oblong, 2

mm long, 2.2 mm wide, concave at the base, the apex broadly rounded, subreniform, thickened; lip light

yellow, thick, su 1 mm long, 1.8 mm wide, 1.5 mm deep, the apex broadly rounded, with the

margins only slightly thickened, shallowly concave anteriorly with a small, narrow, glenion, the dorsum

with a low, rounded callus, the base truncate, hinged to the base of the column; column stout, 1 mm

y EMU, ‘ie 8 Jl Bile 3 - we.

long, 1.5 mm wide, the foot

orn plateau above Chical, alt. 1800 m, 11 Jan. 1993, A. Hirtz 5850 (Holotype: MO), C. Luer illustr.

This species was discovered in wet, relatively lowland of northwestern Ecuador.

It is similar to the widely distributed and frequent S. purpurea, but the ramicauls are

more slender, and the racemes of much larger flowers are shorter and more loosely

flowered. The sepals are white, suffused with pink, the dorsal being 13 millimeters

long, and the deeply concave synsepal being 11 millimeters long. The petals are

large and broad, and the lip is Proportionately small.

29. Stelis repanda Luer & Hirtz, Sp. nov.

as - "epandus, “repand,” referring to the uneven, wavy margin of the floral bracts.

Planta grandis, racemo folio elliptico multilongiore, bracteis floralit ] i b i

obtu

petalis eres Seria Ovato anguste ‘uso, sepalis lateralibus semiconnatis synsepalum formantis,

Plant medium t large, epiphyti iy: : d

enclosed by iii | rere: Caespitose; roots slender. Ramicauls slend

heath : ender er, erect, 5-12 cm long,

through the middle and 2-3 other sheaths at the base. Leaf erect, coria-

Oo obtuse

uccessi the petiole. Inflorescence an erect distichous, subdense,

S$ Vi %, * ’ ,

peduncle ed earn. - ie a with 3-4 flowers Open simultaneously, 15-20 cm long including the

annulus below th plein ce undulate bracts, subtended by a slender Spathe 1-1.2 cm long, from an

gressively sh t the ti Seay Rysads ny THOUS, Oblique, acute, ca. 8 mm long, pro-

dorsal se is hacnes ) Ong: Sepals dull purple, glabrous, the

margins cata: Prttena a the ly media Subacute, 4 mm ong, 3.5 mm ae: ue with the

connate into a broadly ovate, pt oe llores the lateral sepals oblique, ovate, obtuse, 3 mm long,

Breen, transversely semilunate, | mm ong. 2m ee Pat 3 mm wide, with the apices adjacent; petals

ith the in thi ; lip green, vl mM wide, concave at the base, the apex broadly rounded

apex broadly rounded, shallowly concave a, subquadrate, 0.8 mm long, 1.3 mm wide, 0.8 mm deep, the

thickened into low calli the base trunc; They With a minute glenion, the dorsum with the 3 nerves

| Wide, the foot obsolescent,

Zamora-Chinchipe: :

& W. Flores 10957 Giant a an boast of lee at. 2150 m, 26 Mar. 1985, C. Luer, J. Luer, A. Hirtz

A. Hirtz & W. Flores 10933 (MO). Withour nen? above Valladolid, 23 Mar. 1985, C. Luer, J. Luer,

25 May 1988 F

Uzhupud, 988, C. Luer 13660 (me ity, collected by Mario Portillo, cultivated at Hosteris

This Speci .

a many-flowered, Ecuador. It is characterized by

are held obliquely oe the rachis with undulating margins. The flowers

are Connivent into a shal] ma

a Ow, suborbi

the lip has a round apex and a : i The petals are semilunate, and

30. - : Named write Luce & Hinz, sp, nov.

eo Zloch, presently Of Quito Ecuador. ¢ "

cus, sepalo dorsal we (Ruiz & Pav.) Wittd. affinic ei fF hen Payee pis

ieraso-alloso differ, | "YM ¥tcpalo menwnan serie ae . et label-

Si i cmc all

STELIS OF ECUADOR 23

medium or large in size, epiphytic, caespitose; roots slender. Ramicauls erect, slender, 5-20 cm

— the middle third enclosed by a tubular sheath, and another 1-2 sheaths at the base. Leaf erect,

riaceous, elliptical, subacute to acute, petiolate, 6-13 cm long — a petiole 1.5-2 cm long, the

bade 1-2.5 cm wide, cuneate below ‘into the petiol €. Inflorese e lor2 an subcongested, dis-

"10-2

ee 1- os cm m long, with a spathe 6-10 mm long, from an annulus below the apex o of the tensioned, aes

Liat oblique, acute, widely 1 inflated, 5-7 mm long, 4-5 mm wide; pedicels 2 mm long; ovary 4-5 mm

llular-glandular within, the dorsal sepal ni ovate,

ea 10 mm 1 long, 8 mm n wide, incompletely 7- to 9-veined, connate basally to the synsepal for 2 ;

the lateral sepals connate except for the distal 2-3 mm into an ovoid, concave synsepal, 9 mm long, 10

mm wide yas enone 10-veined, with a broad mentum; petals light green, thick, transversely semi-

lunate, 1.5 mm long, 2 mm wide, 3- veined, the b broad, rounded apex with a low, conical swelling on the

— of the pies Ilus; lip thickly triangular, 1 mm long, 1.4

le, | mm deep, the anterior surface obtusey triangular, shallowly concave above a

pial thickening, = bar with | low glenion, the dorsum witha a

ged t the base oF the column; cohen is tout, 1.5 mm long,

ivan he f bsol t, th d

Tungurahua: between Lieto and Triunfo, alt. 2800-3000 m, 25 sian 2001, A. ane X. Hirtz, C. & P.

Dodson, H. Zelenko & R. Zloch 7766 (H oe a ee area, Leito, alt. 3000

m, 12 Oct. 1984, A. Hirtz 199] (MO); southeast of Patate, alt. 3070 m, sg tans e 1971, B B. MacBryde sy

(MO, SEL); Cordillera de los Llanganates, Leito, alt. 3100 m, 11 Apr. 1985, C. Luer, J. Luer, A. Hirtz

W. Flores 11148 (MO).

ce rona-Santiago: forest east of dng alt. 2750 m, 15 May 1988, C. Luer, A. ae et al. 13329 (MO);

n Gualaceo and Limé6n, alt. 2650 m, 16 Feb. 1986, C. Luer, J. Luer, A. Hirtz, W. Flores & A.

Enbre 11837 (MO); same area, a 2150 m, 14 Jan. 1989, C. Luer et al. 13889 (MO -

25 Km south of Cuenca, alt. ca. 3000 m, 12 Feb. 1978, C. Luer, J. Luer & M. Portilla 2650

(SEL .

Loja: road north of Loja, alt. 2900 m, 1 May 1973, L. Holm-Nielsen et al. 4648 (AAU, SEL); road to TV

tower east of Yangana, alt. 2650 m, 4 Mar. 1982, C. Luer et al. 7136 (SEL); above Yangana, alt. 2600 m,

27 Jan. 1986, S. Dalstrim ae . , 1108 (Mi

Zamora-Chinchipe: betw a and Zama, alt. 2800 m, 10 Mar. 1958, C.H. Dodson 215 (SEL);

paramo above Valladolid, ac wae. 20 Jan. 1982, D. D’Alessandro 128 (SEL); east of the pass east of

Loja, alt. 2600 m, 21 Jan. 1992, C. Luer, J. Luer, A. Hirtz, A. & P. Jesup 16066 (M

This species, widely distributed in southeastern Ecuador, has pinned been

included in the ubiquitous S. purpurea. Vegetatively, it is variable in size, but the

leaves are acute and petiolate. The racemes vary greatly in length, but the bracts are

more or less inflated. The flower is readily identified by the gibbous protrusion of

the synsepal, and the conical swelling on the inner margin of the petals. The latter

is sometimes nearly absent. The lip is also broad with a thick, obtuse apex. The bar

is bicallous with a rounded dorsal callus.

ai. bs sanchezii Luer & Hirtz, sp. n

Named for Dr. Eduardo Sanchez of Cuenca, weet aaa yas this species.

DI foali llintica multi

oe Ea

ai tic hi il + +.

longiore, pee latissime ovatis obtusis lateralibus ad

labello triangulari concavo distinguitur.

Plant small, epiphytic, caespitose; roots slender. Ramicauls erect, slender, less than | cm long,

Leaf ere

enclosed by 2-3 tubular sheaths. ct, coriaceous, narrowly elliptical, subacute, petiolate, 2.5-4

cm oe including a petiole 0.7-1 cm — ee blade 0.7-0.9 cm wide, narrowly cuneate below into the

. Inflorescence a flexible, flex ax, successively many-flow og raceme up to 18 cm long or

longer including th the A cnt ay cm aie eventually becoming pendent, with 1-3 flowers open simul-

eously, fro’ - floral ou — acute, 2.5 mm long;

meres 1 mm anions si a “ mm ene flowers suffused with purple; sepals incomp — pai

glabrous, the dorsal sepal broadly ovate, obtuse, 5 mm long, 4 mm wide, neanerdoirte connat ‘o the

lateral sepals, the lateral sepals fe ‘connate 1 mm into a shallowly concave, bifid s am 5 ; mm

long, each sepal 2.75 mm wide, obtuse; petals ewan obcuneate, the apex truncate with the margin

; pel

thickened, 1 mm long, 1.75 mm wide, 1-veined, sligh shallowly concave

below; lip thickly triangular, 1 narrow sic rounded ms the lott io 75 mm long, | mm n wide, - mm deep,

cal margin. it, the d. central

callus, the base cuneate, h ged t Pek mney abe stout. npeog se 1 mm wide,

J =

24 ICONES PLEUROTHALLIDINARUM

rn 4 Dien da bail (I imé6n), alt - 2200 m, 7 Jan. 1995, A. Hirtz,

Hirtz & E. Sanchez 6120 Holotype: MO), C. Lue illustr. 20049; vcultivated at

Ecuagenera, Gualaceo, 5 Mar. 2001, C. Luer 19723 (MO).

This species, known from southern Ecuador, is characterized by the small,

caespitose habit; elliptical leaves borne by short ramicauls; and an elon gating,

arching to pendent, flexuous, distantly and successively flowered raceme. The

sepals are broad and obtuse, the laterals semiconnate; the petals are truncate; and

the lip is thickly triangular, concave anteriorly below a prominent transverse bar

with a large, rounded callus on the dorsum

San see coating Lincs uer & Hirtz, sp. nov.

Ety.: N or Santiago Arévalo of Macas, Ecuador, who collected and cultivated this species.

enna ans 8. Ppurpureae (Ruiz & Pav. ) Willd. affinis, sed beartets floralibus site non-inflatis,

synsepalo mentoso intus brev init a labello brevi

bitumido antice subplano di

Plant large, epiphytic, caespitose; roots slender. Ramicauls Rpg pals stout, 12-15 cm long,

enclosed by a loose, tubular m below the middle, and another 2-3 t the base. Leaf

erect, coriaceous, elliptical, acute, 12-15 cm yoo 3-3.5 cm wide, cuneate ‘trier into a petiole less than 1

7 rescence an erect, ested, distichous, simultaneously many-flowered raceme, 20-30

cm long including the peduncle 10-12 cm long, with a slender spathe 2 cm long, from an annulus below

ramicaul; floral bracts , Short, 3 mm fee. 2 mm wide; siaieal 2 mm long; ovary

curved, 3 mm long lowers dark purple, sepals glabro externally, the d ellular-glandular

recurved sides, narrowly obtuse, 10 mm long, 5 mm wide, 7-veined, connate to the

ate into a age ovate, concave

synsepal, 8 mm long, 7 mm wide unexpanded, 6-veined, with, a shallo tum

ar,

the lat

w men ansversely

semilunate, 1.1 mm lon ig, 1.6 mm wide, 3-veined, the apex broadly rounded with tk margin rere

minutely pubescent, jaiicals concave basally; lip su’ €-subtriangular, 0.5 mm long, 1.1 m

wide, | mm deep, the apex ded, the bar bitumi til a small glenion, the dorsum descend-

ing with a low, more or less three-lobed callus,

column stout, 0.5 mm long, 1.2 mm wi the fi

stigma bilobed. . si

orona-Santiago: Cordillera del Cutucii, east of Macas, alt. 800 : as

Mac 20 Feb. 2001, C. Luer 19510 (Holotype: MO). ti m, collected by Santiago Arévalo in

ge The rounded Margins of

with the anterior surface plain, with only a b.

within the broadly rounded ya arely perceptible pair of low calli

33. Stelis situlifera Luer & Hirtz, sp. no

Ety.: From the Latin situlifer, “bearing a water Sete” referring to the a concave cece

- minasve ae haec S. purpureae (Ruiz & Pav.) Willd. affinis <4...

PT EIS gracili, bracteis floralibus acutis, sepalo dorsali obs ls mine 7

SIP, E oncavo, petalis crassis, et labello crasso calloso viff

begat esa mu ong. 7 mn wide, incompletely 7

deeply concave, ould epuiggan ote 5 mm long, ne 8mm wide werpeern coma no 8-veined: i eels

STELIS OF ECUADOR 25

rose, thick, transversely semilunate, ns - long, 2mm wile; ‘Sevcenett, the apex broadly rounded with the

margin alana concave aig’ ow a tran s; lip thickly quadrate- triangular, 0.8 mm

long, 1.5 mm wide, 0.8 m dl reat att obscure glenion higher than

the sides, the anterior surface voae if at all, poset peo sei with a broad, low calles, the base

truncate, hinged to the base of the column; column st ong, 1.5 mm wide, the foot obsolescent,

the anther and the bilobed stigma apical.

Loja: along the new road north of Loja, alt. 2600 m, 2 Feb. 1987, C. Luer, J. Luer & A. Hirtz 12579

(Holotype: MO).

This relative of the frequent S. purpurea is known from the province of Loja in

southern Ecuador. It is distinguished by a large, petiolate leaf as long as the rami-

caul. The several-flowered raceme is more or less as long as the leaf. In the only

collection known, the flowers appear to have been borne successively on a slender

peduncle and rachis. The dorsal sepal stands erect and the synsepal is short, trans-

verse and deep. The petals are not unusual. The lip is shallow without an obvious

glenion.

34. bag sumacoénsis Luer = — sp. n

ed for here the avi was ee

Species haec S. d l Rchb.f. affinis d

+ Letty Wy 1

Plant small, epiphytic ts slender. Ramicauls erect, slender, 6-10 mm long, enclosed

by two, tubular shea . Leaf erec , narrowly elliptical-obovate, ver petiolate

mm long — a petiole e 8- Ot mm mae. the blade 5-7 mm wide, narrowly cuneate below into the

petiole. Inflorescence a lax, successively oe raceme, more or les ess Sine. to 10 cm long

including the iaaier an ncle ca. 2. cm long, 2 imult from below the apex o

the ao floral bracts infundibular, a = -_ long; Paco 0.75 mm long; ovary 1.5 mm

lon al erect, broadly ovate, obtuse, connate

L; 31 mm basally to = synsepal, 5 vom lon ng, 4mm er nga the lateral sepals connate into a deeply

concave, ovoid synsepal, 4.5 mm long, 5 mm wide, 6-veined; petals purple, cellular, transversely ovate

milunate, c caer below the ro ae rounded apex, 1 mm long, 1.4 mm wide, 1-veined, concave

esalhs below the transverse callus; © purple, s subovord, 0.4 mm long, 0.8 mm wide, 0.6 mm —

shallow d, the margin indistinctly’ thickened, the bar w

pose slevited, channeled glenion, th the dorsum

hinged to the base of the column; column stout, 0.8 mm long, 0.8 mm wide, the foot obsolescent, the

anther and the bilobed stigma apical.

Napo: Sumaco Volcano, Rio Hollin, alt. 1200 m, 13 Aug. 1991, A. Hirtz & S. Ortega 5396 (Holotype:

MO), C. Luer illustr. 20066.

This little species of section Humboldtia is characterized by a caespitose clump

of narrowly obovate leaves that is practically indistinguishable from many others in

the genus, as well as from many species in other genera, such as Platystele or

Pleurothallis.

A loose, flexible, successively flowered raceme surpasses the leaves. With the

weight of the flowers the raceme bends, eventually becoming pendent with non-

resupinate flowers. The synsepal is deeply concave; the petals are thickly semi-

lunate, concave below the apical margin, and concave below the transverse callus.

he apex of the lip is round, and the glenion is elevated, narrowly oblong, and

minutely channel

35. Eick meee Luer & Hirtz, sp. n

e Latin velivola, ‘‘a sail boat,” sii to the vii like stitial with a sail.

Species fe oh pene (Ruiz & Pav.) Willd. affinis, sed h ynsepalo late profun-

toso differt.

i piphy der. Ramicauls erect, stout, 5-7 cm

ytic

ee enclosed rakes . a loose, pc sheath from the lower quarter, and another 1-2 sheaths at the

26 ICONES PLEUROTHALLIDINARUM

Leaf erect, coriaceous, ellipti i including a petiole 1-

. elliptical, subacute to acute, petiolate, 5-6 cm long inc be

rake long, the blade 0.9-1.2 zy wide, cuneate below into the petiole. Inflorescence an erect, dis

ly, 7-9 cm long includ-

many- Wii y flowers ; :

ing the peduncle 2-3 cm long, with a spathe 6-7 mm long, from an annulus below the apex of the pan

caul; ique, acute, inflated, 4 mm long; pedicels 2 mm long; ovary curved, 2.5 ong;

flowers

; glabrous externally, microscopically pubescent thin,

the dorsal sepal erect, ovate, obtuse, 8.5 mm long, 5.5 mm wide, incompletely 5- to 7-veined, connate

basally to the synsepal for I mm, the lateral sepals connate into a broadly ovoid, deeply concave ee

Pal, abruptly concave below the central apparatus into a broad mentum, 6.5 mm long, 7.5 —o

unexpanded, 10-veined; petals suborbicular, broadly semilunate, 1.25 mm long, 1.5 mm wide,

rounded callous, truncate, hin

mm wide, the foot obsolescent, the anther and the bilobed

Zamora-Chinchipe: east of the pass east of Loja, alt. 2750 m, 20 Feb. 1986, C. Luer, J. Luer, A. Hirtz,

W. Flores & A. Embree 11974 (Holotype: MO).

Related to S. purpurea and its many allies, this species of southern Ecuador is

distinguished from most of them by the small habit with a long, loose, tubular

sheath covering the ramicaul. The Taceme slightly exceeds the small, elliptical

oral te)

le with

t protrusion of the inner margin of the

in. The lip is ovoid and concave anteriorly above a thickened apex.

36. Stelis vollesii Luer & Dodson, sp. nov.

Ety.: Named for Hans Volles of Tdquerres, Colombia, Co-collector of this species.

Species haec §. globi; Rchb-f. ; sed es Itimi . . dib triangular-

is earn ee eos, con bf. —— floribus multiminoribus, petalis grandibus

vant Stall epiphytic, caespitose; roots slender Ramicauls slender 1-4 cm long, enclosed by

> . > €rect, cm ong,

a tubular sheath and another at the base. Leaf erect, coriaceous, elliptical, petiolate, acute to subacute,

15-5 cm long including the petiole ca, 1

aca a ‘rect to arching, di flexuous, successively many-flowered raceme

ee eumultaneously, 3-13 cm long including the peduncle 1-3 cm lon , subtended by a

og eis Seem a aU low the apc fra tel 3 on

a :

i : . igher than the sides and

the base of th , tae ae ith a , Callus, the base truncate, hinged to

the bilobed stigma apical. “mm long, 0.8 mm wide, the foot obsolescent, the anther and

rey Nae pobnsece oe Of Lita, alt. 1450 m, 13 Mar. 1991. CH Dodson, H. Volles

20068. * forested plateau above Chical, alt. 1800 m, 11 Jan. 1993, A. Hirtz 5853 (MO), C. Luer illustr.

fe ca — ogee dtia is 4pparently endemic in northwestern-

most or. Closely allied to 5 8lobiflora, it is distinguished from the latter by

comtively fioweteee vd ranBular petals. A 0ose, flexuous, suc-

me eventually far Surpasses :

broadly ovate while the shallow] the leaves, The sepal is

Proportionately large petals are trianoy

lip is ovoid with a broadly

the dorsum.

eh MI WSS

fan fo

Fig. 4. Stelis zelenkoi

tei ne ee eee ee pr MR

Fig. 11. Stelis brachiata Fig. 12. Stelis coeliaca

Fig. 19. Stelis florianii Fig. 20. Stelis galeola

Fig. 22. Stelis nycterina

Fig. 28. Stelis regina

Fig. 34. Stelis sumacoénsis

ears

Fig. 35. Stelis velivolva Fig. 36. Stelis vollesii

36 ICONES PLEUROTHALLIDINARUM

37. Stelis adrianae Luer, sp. nov.

Ety.: Named for Adriana, wife of A.P. Sijm, who collected and cultivates this species.

sf1 P b-parsieme, lei fl £32

i porpnyrea,

lis obtusis di longiput t petalis crassis. et labell

» P ’

Be. ef

lato distinguitur.

Plant medium in size, epiphytic, ascending-caespitose, rhizome thick; roots slender. Ramicauls

i stout, fascicled, 3-4 cm long, enclosed by 2 loose, tubular sheaths. Leaf purple, erect,

thickly coriaceous, elliptical, subacute to obtuse, 5-8.5 cm long including a petiole ca. 1 cm long, the

blade 1.8-2.7 cm wide te below i duplicate petiole within which the peduncl ts. In-

florescence to 15 cm long, the raceme 5 ng, congested ks, distichous, y

many-flowered, with the flowers touching, borne by a slender peduncle 9 cm long, from the base of the

leaf; floral bracts tubular, 2.5 mm long; pedicels 1 mm long; ovary 1.75 mm long; flowers dark purple;

sepals fleshy, glabrous externally, densely long-pubescent within, with margins more or less recurved,

the dorsal sepal ovate, obtuse, 5 mm long, 3.75 mm wide, 3-veined, connate less than 1 mm basally, the

lateral Is obl oblique. semiconnate. def] db h tt taini th tral ap aratus,

o .

subquadrate, 0.75 mm long, 0.75 mm wide, 0.6 mm deep, the anterior surface shallowly concave, the

roadly rounded, apiculate, transversely thickened across the middle with a longitudinal glenion,

the dorsum flat, densely short-put i the t the b , hing the base of the col-

umn; column stout, 0.75 mm long, 0.8 mm wide, the foot obsolescent, the anther and bilobed stigma

preg ee near _— sett 1650 m, 25 Feb. 2001, collected by S. Dalstrém & L. Jost, C. Luer

(Holotype: ; near Mindo, 1994, collected and culti i i

AP. Sijm 201231 (MO). c and cultivated in Venhuizen, the Netherlands, by

This handsome species is most notable for the deep purple pigmentation

thro ughout the plant. The habit is ascending with a fascicle of ramicauls trailing a

thick rhizome entwined in a mass of fine roots. The petioles of the ovate leaves are

38. Stelis amoena Luer & Hirtz, sp. nov

Ety.: From the Latin amoena, “pretty,” referring to the plant.

avec han’ — s oe affinis, sed ramicaulibus in vaginis laxis obtectis foliis ellipticis leviter

unum ao ee ssive multifloris, sepalis ovatis obtusis similibus grosse ciliatis, petalis

F tinguitur.

Plant medium in size to lar. i i i

8¢, epiphytic, densely caespitose: i :

slender, 3-11 cm long, encl by 2 loose, tubular hats nie saci sapien ee ae

ae ee a subacute, 6-11 cm long, 1-1.8 cm wide, narrowly cuneate below into the

delle Serre ‘ — an erect, subcongested, distichous, simultaneously many-flowered

pe ab nid reg inc uding the peduncle 6-10 cm long, from an annulus below the apex of the

: cts acute, oblique, 3.5 mm long; pedicels 2 mm long; ovary 1.5 mm long; sepals

iy enes ateces reer de middle, coarsely ciliate-denticulate, broad-

lateral sepals 3 mm long, 3 mm wid ly, the dorsal sepal 3.5 mm long, 3.5 mm wide, 3-veined, the

wide, 3-veined; petals transverse] ve :

concave belo’ broadly rounded : , ersely elliptical, 1.5 mm long, 2 mm wide,

mm wide, ecu deep va ike, apical margin; lip purple, subquadrate, 1 mm long, 2

a faint, superficial glenion, the dorsarn pre gin, shallowly teriorly with

: calenaie “ um pubescent, the base broad! i f th

— stout, 1 mm long, 2 mm wide, the foot obs sad ee Benged to Ge reser ca

Hirtz 12457 Postar — of Volcén Iliniza, alt. 2800 m, 23 Jan. 1987, C. Luer, J. Luer & A.

irtz 19871 (MO), Calacalf and Nanegalito, alt. 2000 m, 9 Mar. 2001, C. Luer & A.

STELIS OF ECUADOR 37

This pretty species, apparently endemic on the western slope of Pichincha, is

characterized by ramicauls concealed by loose sheaths, elliptical leaves, and a dis-

tichous raceme of purple flowers facing the same direction. The sepals are similar,

ovate and ciliate-denticulate. The petals are transverse and concave, and the lip is

rounded.

39. Stelis andreettae Luer & Hirtz, sp. no

Ety.: Named for Father Angel Andreetta of Paute Ecuador, co-collector of the species and well

known for h

fali kh ; ; }

wae 2 o

s haec S. lindenii Lindl _affinis

nervis, et labello leviter concavo differt.

Plant large, epiphytic, densely sige to hema repent, the rhizome ca. 7 mm thick; roots

slender. Ramicauls erect, relatively slender, 6-14 cm long, with a loose, tubular sheath above the middle

and another 2 ees at the base. Sxep ieee aa elliptical-ovate, subacute to obtuse, petiolate,

9-13 cm long ere the petiol 1.5-2 cm Suen the eens 3-4 cm wide, cune eate below into the petiole ole.

Inflorescence an t, distichous, : ae

taneously, 5-7 cm ape including the ee ca. “ cm cigars witha afore Ms cm long at an patent

below the apex of the ramicaul; floral bracts oblique, e, 3-4 mm long; pedicels 1-1.5 mm long; 0

1.5 mm long; sepals similar, Ae ee brandy. pee broadly obtuse, penn to — to a

middle, the erate sepal 3.2 mm wide, 3 veined, the lateral sepals 2.7 mm long, 3. 3 mmm

wide, 3-veined; sear E rown, abr Bante bl

ing, 1 mm long, 1. mm wide, 3- eee with the margin broadly rounded ‘and thickened; lip brown,

, 1mm

rary shale e, 0.75 mm long, 1.3 mm wi deep, the anterior surface shallowly concave

h the apex broadly rounded and thickened, the oer Pathe a small ph viens the dorsum with a low

bends callus, th column; column stout, 1 mm long, 1.2 mm

wide, the foot obsolescent, the anther and the bilobed nee apical.

Zamora-Chinchipe: new road east of th Loja and Zamora, alt. 2250 m, 22 May 1988, C.

Luer, J. Luer, A. Hirt et al. 13605 (Holotype: MO).

This large species is uncommon, known from only one collection in southeast-

ern Ecuador. Vegetatively, it is superficially similar to the frequent S. lindenii.

Both are vegetatively large with elliptical-ovate, petiolate leaves. The ramicaul of

S. andreettae is slender with a sheath above the middle, and the raceme is shorter

than the leaf. The sepals are expanded, membranous, broadly ovate, obtuse, and

three-veined, instead of fleshy, sensitive, and five-veined. The broadly obtuse

petals are not remarkably different from those of S. lindenii. The lip is much less

deeply concave.

40. bay aarp Luer & Hirtz, sp. n

the Latin apiculifer, Pagans, an sical,” cepeeade to the labellum.

ann haec S. gui is Rolf is, sed sepalis pory

fert.

Plant small, epiphytic, caespitose; roots slender. Ramicauls erect, slender, 4-10 mm long, enclosed

by 2-3 tubular sheaths. Leaf erect, coriaceous, narrowly obovate, subacute, 10-25 mm long, 4-6 mm

ide, gradually narrowed below into a petiole 3-5 mm lon an erect, distichous, subcon-

gested, aaa few-flowered raceme 3-5. a cm n long including the peduncle 2-2.5 cm long, from

lar, 1.5 mm long; pedicels 1 mm long; ovary

0.6 l bove the middl : 1 apares the middle, glabrous externally, cellular

papillose within, years ovate, omg: oe connate basally, the dorsal sepal 1.75 mm long, 2 mm

wide, the lateral sepals 1.25 mm long, 1.75 wide; petals purple, transv oe =e broadly round-

ed-truncate at the apex with a he ane ot concave, 0.5 m g, 0.8 mm wide, |-

veined; lip thick, Hs spemesip 0. 5 mm long, 0.6 mm wide, 0.3 mm deep, ue concave syirta

unded, nute, ~~ apiculum in the . center, the anterior surface co:

ed with a long, very narrow glenion, the dorsum roun unded, cellular-pubescent above the ‘ona res "i

ee nad a to an obsolescent column-foot; column stout, 0.6 mm long, 0.6 mm wide, with the

and the bilobed stigma apical.

38 ICONES PLEUROTHALLIDINARUM

Morona Morona, alt. 950 m, 17 Jan.

1989, C. Luer, J. J. Luer, A. Hirtz, S. aout A. & P. hak 13953 vented MO

This little species is closely related to S. guianansis, Seine only in a more

congested raceme with broadly ovate, cellular-papillose sepals. The petals are

single-veined. As in S. guianansis, the minute lip is thickly discoid with a tiny,

sharp apiculum in the center of the round apical margin.

41. Stelis barbellata Luer & Hirtz, sp. nov.

Ety.: From the Latin saceapoen ee bares” eaiiad to the tails of the <img

oes haec s. h y ij is prolif ti, sepalis

Plant large, epiphytic, i i nosey and prob; roots slender. Ramicauls — bee:

srg 10-22 cm a! below, 4-8 ig above, enclose a long, tubular sheath above the middle

heath at : ; elli iptic , Subacute to obtuse, peti va ‘Sl cm

a long including the est 0. = m m long, ine b blade 2 2 3. 5 cm wide, cuneate below into the petiole. In-

su many with several flowers open

simultaneously, 12-20 cm long ‘including the > peduncle 37 cm long, with a spathe 1-1.5 mm long, from

blique, acute, 3-4 mm lon ng; tigi

1.5 mm long; ovary i 5 mm fone: ers either es or varie sepals glabrous, ovate, obtus

vex, 5 mm long, 4 mm wide, 3-veined, aise mm

basally to the lateral sepals, the oails sepals 5 mm mm long, 4 mm wide, 3-veined, connate 2 mm; petals

ly obovate, 2 mm wide, 3 mm long, including the obtuse apex acuminate into a ig pro-

pin 1 mm long; lip thin, subquadrate, acute, 0.75 mm long, 1.5 mm wide, 2 mm deep, co above

third, the basal third occupied with a thick, densely pubescent callus, the base ae eee

to the base of the column; res BO 1.5 mm long, 2.75 mm wide, the foot obsolescent, the anther

apical, with the stigmatic lobes

oe — - x scons st on ut hemi alt. 3050 m, 21 Feb. 1986, C. Luer, J. Luer, A.

Although the petals and lip of this species suggest a relationship with the Boli-

vian S. antennata, the habit differs dramatically. This large species is characterized

by ramicauls both caespitose and prolific, and by elliptical, petiolate leaves sur-

ies by a subdense raceme of either yellow or purple flowers. From the center of

petals. The lip is oblong, concave, and acute. The basal third is filled with a trans-

verse, densely pubescent callus.

42. Stelis benzingii Luer & Hirtz, s

Ety.: Named for David ici Barn a of Botany,

Species haec S. argentatae Lindl.

aia ae similis, sed florib

Oberlin College, co-collector of this species.

us iescnig non-resupinatis, racemo laxifloro,

Plant medi

with a loose, hibeedar laze. city casts: —_ ree mapa erect, slender, 4-18 cm long,

ceous, ovat third -3 sheaths wer third. Leaf erect, coria-

2 13 Stash sae eas sti ‘igre long, 2-4 cm wide, eacdie below into an indistinct

most flowers < ulnar t, loose, distichous or secund, many-flowered raceme with

15 mm Mtoebcandenprenems : 15.2¢em| mms including the » peduncle ca. 5 cm long, with a

pedicels blique, a mm long;

cnk epee cnr cine 5mm long; flowers yellow, non-resupinate; sepals shor yes:

sepal 4 mm long, 4 mm wide ex se pau Sides recurved, co: ne basally | mm, the

veined; petals transverse sls oF > veined, the lateral cel sepals 3.5 mat sm long, 4

rounded, flat, shallowly pee : mn long, 1.5 mm wide, 3-veined, apex broadly pabtrancsto-

0.5 mm deep, shallow! W; lip thick, transversely su te, 0.3 mm long, 1 mm wide,

in the center, the Fie pone moe b t ¢ broadly inded margin, with 3 minute, sharp

inged to the base

of the

cme Tg a in ie ne

STELIS OF ECUADOR 39

Esmeraldas: wet forest west of Lita, alt. 750 m, 18 Jan. 1987, C. Luer; J. Luer, C. Dodson, A. Hirtz & D.

Benzing 12405 (Holotype: MO).

This species from lowland, coastal Ecuador is similar to the widely distributed

S. argentata and its numerous allies, but differing with larger, non-resupinate flow-

ers with elongated ovaries, and loosely arranged in a long raceme. The margin of

the lip is sharply tridentate, similar to that of S$. vulcanica Schltr., but the flowers of

the latter are smaller and glabrous.

43. Stelis bicarinata Luer & Hirtz, sp. n

Ety.: From the Latin bicarinatus, “‘bicarinate,” keine to the callus of the —

Pe ee Tei] ot & Paan Ts

Planta grandis, signin grail f

sepalis ovatis villosis, et labe

Pikes pes PASC densely caespitose; roots slender. Ramicauls erect, ord a te = long,

a short, and 2-3 others at the base. Leaf erec liptical,

pea re 10-13 cm long including the petiole 2-2.5 cm long, the blade rs 5. ven cm ee ict

below into the petiole. Inflorescence 1-3 erect, subdense, distichous, several-flowered racemes ee

m owers open simultaneously, 6-10 cm g the peduncle 2-3 cm — subtended b

slender — 10-13 mm een from an annulus below the apex of the ramicaul; ts in ee

lar, 2 mm long; pedicels 1.5 mm long; ovary 0.75 mm long; sepals expanded, light 3 elo ws Sols villous

within, glabrous externally, connate basally, ovate, rang the dorsal sepal 2.5 mm long, 2 mm wide, 3-

veined, the lateral sepals 2 mm — - mm ee 3- veined; petals light yellow, — ersely cuneate-

obovate, shallowly concave, 0.5 mm 1.1m , the apex broadly rounded wi

rs less irregular and thickened; lip igh yellow, thick, subquadrate, 0.9 mm long, 0.6 mm wide, 0.6 mm

cep, p g P ith short, acuminate eros, the bar cn

nts a pair of arinae, th a flat, ciliated he base broadly tru

cate, hinged to the faa of the ae column stout, 0.75 mm long, 0.6 mm ne the ome orn acer

anther and the bilobed stigma apical.

Zamora-Chinchipe: south of the pass south of Yangana, above Valladolid, alt. 1900 m, 23 Mar. 1985,

C. Luer, J. Luer, A. Hirtz & W. Flores 10839 (Holotype: MO).

This species, apparently endemic in southeastern Ecuador, is characterized by

long, slender ramicauls that bear much shorter petiolate leaves. One to three

slender, erect racemes of tiny flowers do not pass the leaf in length. The sepals are

ovate and long-villous within. The petals are transverse with a thickened margin.

The subquadrate lip is round at the apex with a small, incurved apiculum, and a

bicarinate, vertical callus occupies the center.

44. boing er Luer & Hirtz, s

sp. n

e Latin congestus, “aenee _ refering to the bor nein raceme.

sate mediocris cacspoam, | racemis | 1-3 d

lic la ‘i rs

2 = mS

a 1 ictineuitur

nt medium in size, epiphytic, densely caespitose; roots filiform. Ramicauls erect, slender, 5-9

cm a enclosed by 2-3 tubular sheaths. Leaf erect, coriaceous, narrowly elliptical, acute to subacute,

3- - cm long including a petiole 1-1.5 cm long, ton blade 1-15 cm wide, ee cuneate below into the

Inflo

mm long; pedicels 1 2% long; ovary 1 mm long; sepals expanded, purple, glabrous externally, cellular

anv ent within, broa y ova ate, — connate aie! the dorsal a 2.5 mm long, 2.5 mm w

wide, 3-veined; li le, thickly angular-subqu uadrate, 0.5 mm long, 1 mm wide, 0.4 mm iw Mat to to

barely convex anteriorly, bigs then margins thickened, the apex broadly rounded, the bar thick, with a

shallow glenion, the dorsum with a rounded, cellular pubescent callus, the base broadly truncate, hinged

to the base of the column; aheuihviee tout, 0.75 mm long, 1 mm ae vibe foot obsolescent, the anther an

the bilobed stigma apical.

40 ICONES PLEUROTHALLIDINARUM

M Santiago: Cordillera del Condor, east of Chuchumbletza, alt. 1750 m, 21 May 1988, C. Luer, A.

Hirtz, W. Flores, A. Andreetta & W. Teague 13547 (Holotype: MO); new road west of Macas toward

Guamote, alt. 1900 m, 6 Feb. 1987, C. Luer, J. Luer & A. Hirtz 12709 (MO).

This species, known from the Oriente of Ecuador, is distinguished by a caespi-

tose habit with narrowly elliptical leaves borne by slightly longer ramicauls, and

one to three inflorescences shorter than the leaves. The raceme is slowly succes-

sively flowered, becoming more congested above with the flowers overlapping.

The sepals are expanded and obtuse; the petals are transversely ovate; and the lip is

nearly flat or barely convex anteriorly with a small, superficial glenion.

45. Stelis cutucuénsis Luer & Hirtz, sp. nov.

fant?

te narve d ee : ee ea ~ aa *

Planta p racemis paucis p t to superanti,

a | ih . eral . arr a . : . . . .

se. u-

? ” ia ae vie!

natis trinervis, et labell j bp! i do disti

Plant small, epiphytic, densely caespitose; roots slender. Ramicauls erect, slender, 5-7 mm long,

enclosed by 2-3 tubular sheaths. Leaf erect, coriaceous, narrowly obovate, subacute, 15-20 mm long

includ ; :

petiole ca. 5 mm long, the blade 3-5 mm wide, gradually narrowed below into the petiole.

dorsal sepal 2 mm long, 2 mm wide, the lateral sepals oblique, 1.5 mm long, 2 mm wide; petals purple,

semilunate, broadly ded and thick i on th gi he apex, conc 0.75

wide, 3-veined; lip purple, subdiscoid, 0.25 mm long, 0.6 mm wide, 0.5 mm , nearly flat to shallow-

ly concave anteriorly with the apex rounded, the anterior surface featureless, the dorsum shallow, with a

rounded, central callus, the base tri hinged t fee

long, 0.6 oP tress hinged to a lesc foot; column stout, 0.5 mm

plekiyatonsa yet ng del Cutucti, between Mendez and Morona, alt. 950 m, 17 Jan. 1989, C.

- r, aa A. Hirtz, S. Ortega, A. & P. Jesup 13952 (Holotype: MO); along Rio Upano, north of

acas, alt. 1100 m, 15 Jan. 1989, C. Luer, J. Luer, A. Hirtz, S. Ortega, A. & P. Jesup 13920 (MO).

The lip is discoid with the anterior surface very shallowly concave and featureless,

while the narrow dorsum has a central, rounded callus.

46. Stelis dalessandroi Luer, Sp. nov.

Ety.: Named in honor of Dennis D’ Alessandro who collected

tali hors, Ss. breviracemae C.Schweinf. PRE geen ere on

md ) an 4 *« talbott kK Ag nm

this species.

fn}

Piant medium in size to | ‘ : :

arge, epiphytic, caespitose; :

stout, 10-11 cm long, with a close, te rr se the middle. seein ovarae. age SR

coriaceous, narrowly ellipti Cre s ;

eave uae ly ellipti = acute, 14-15 cm long including a petiole 1.5-2 cm long, the blade 2-2.5

Fieenna Seat now iaiio the: petiole. , an erect, congested, distichous, many-

long, from an annulus helow ae aneously, 8-9 cm long including the peduncle ca. 1 cm

meee eben 0 Imm long; ovary 1 mm long: is phate leaning yg aon an

jon ot hone spr 3-veined, the lateral sepals 2.25 mm long, 2 mm wide, 3-veined;

é fl ersely se . _ ng, 1.3 mm wide, 3-veined, concave below the broadly obtuse,

thickened, flattened, margin; lip thick, subquadrangular, 0.8 mm long, 0.8 mm wide, 0.5 mm deep,

STELIS OF ECUADOR 4]

shallowly CORCANG peatteaioaly with the apex i ~ bar with a glenion anteriorly, with 3 alow low, sie

angular

causaloeciumhieees foot ob d the bilobed stigma ap

da Hond th of Yangana, alt. ca. 1800 m, collected by D.

D’ Alessandro 71-0083, flowered in cultivation at SEL, July 1981, C. Luer 6449 (Holotype: SEL),

This species is similar to Peruvian S. breviracema with an acute, elliptical, pe-

tiolate leaf and a densely flowered raceme about half as long. The sepals are ob-

= the petals are transversely ovate, and the lip is thickly subquadrate. Vegeta-

vely, this species differs from S. breviracema with wider leaves. If it were not for

“t ditieeences 3 in the lip, they could represent the same species. The lip of S. da-

lessandroi is deeper from the base to the bar. This distance is illustrated as short in

S. breviracema. The width of lip of both species appears to be the same. The

length of the lip of S. dalessandroi is short, creating a flat, horizontal lip beneath the

column. The depth of the lip of S. breviracema is long, creating a flat, vertical lip.

47. Stelis dalstroemii at pe nov.

Ety.: Named for sine Dalstré f S ta, Florida, co-collector of this species.

As Ls lef a3 Ths cats: 1 Py dunialan

r a =

lM . “1 eo = ee ee 1

Plan

giore, “sn aes ovatis obtusis lateralibus minoribus pet

acuto distinguitu

Plant large, epiphytic, caespitose; roots numerous, slender. Ramicauls erect, ~~ 10-17 cm long,

enclosed by: a gon sheath on the middle third and another 1-2 sheaths at the base. Leaf erect, coria-

ceous, see iptic te, petiolate, 7-13 cm long including the petiole 1 ae m long, the

aie cuneate below into the petiole. Inflorescence a strict, erect, distichans, sere

sieabaneors sly many-flowered raceme,

Ain = long os nae 8 the stout peduncle 3-7 cm long, subtended by a spathe 10-12 mm long, fro fro

us below i of the 1 ramicaul; nit racts ts tubular, obtuse, = mm long; pedicels 1- “Ss mm

“i ovary I. ae ng panded cellular within, transversely

ovate, broadly hee, connate chon the pe? sepal 6 mm ong, 6.25 mm wide, 5-veined, the lateral

sepals oblique, 3.5 mm — 5 mm wide; petals thickly de ch Hag sea basally and below the

broadly obtuse apex, 1.5 mm long, 1.5 mm wide; lip thick, subtriangular, 1 mm long, 1.2 mm wide, 1

mm deep, concave naa a apex acute, the bar thick with a aa. the dorsum with a rounded

care the base broadly tr truncate, hing the b ase of the column column stout, | mm long, | mm

Carchi: forest between Tulcdn and Maldonado, alt. 2050 m, 2-4 Apr. 1984, C. Luer, 5. Dalstrim & T.

ois 9917 (Holotype: MO); same area, alt. 1950 m, 2-4 Apr. 1984, C. Luer, S. Dalstrim & T. Héijer

911 (MO); same area, alt. 1600 = C. Luer, J. Luer, J. del Hierro, A. & X. Hirtz 15090 (MO); east of

ata alt. 1700 m, 17 Mar. 1991, C. Luer, J. Luer, J. del Hierro, A. & X. Hirtz 15130 (MO).

Esmeraldas: wet forest west of er alt. 750 m, 18 Jan. 1987, C. Luer, J. Luer, A. Hirtz et al. 12404

(MO).

This species, apparently confined to northernmost Ecuador, is related to S.

purpurea. It is vegetatively similar; the flowers are purple- -black and arranged in

two rows, back to back, more or less overlapping in a congested raceme. The sepals

are very broad with the dorsal sepal distinctly larger than the oblique lateral sepals.

The petals are thickly semilunate, and the lip is triangular and acute.

48. hse chains jor’ Luer & Hirtz, sp. n

the Latin di oe “making sight” referring to the habit.

Planta mediocris ta, racemo elongato mein longipedicella-

to. sena - , ee + roy * Iahelli guiti

a a - z =

Plant medium to large, scandent, the rhizome erect, stout, 2-3 cm long between ramicauls, roots

slender at the base. Ramicauls erect, ascending, slender, 4-6 cm long, enclosed by 3-4 tubular sheaths.

rect, coriaceous, elliptical, subacute, ait 46 cm long including the atige - : cm long, the

ade 1-1.3 cm wide, cuneate below into the petiole. rescence an erect, sublax, distichous, slightly

pec many-flowered raceme, with 2-3 flowers open simultaneously, 18-30 cm si alain ie

42 ICONES PLEUROTHALLIDINARUM

peduncle 8-10 cm long, ry 1 PSL +, r f th i IE floral bracts oblique, acute, Z

mm long; pedicels 2-3.5 mm long; ovary 3 mm long; sepals purple, glabrous externally, pubescent

within, broadly ovate, obtuse, 5-veined, connate dorsal sepal ng, 4.5 mm wide, 5-

veined, the lateral sepals oblique, 3.5 mm long, 4 mm wide, 5-veined; petals green, transversely ovate,

0.9 mm long, 1.4 mm wide, 3-veined, the apex b lly obtuse with th gin thick and flat, shallowly

concave below; lip green, subquadrate, 0.5 mm long, 0.8 mm wide, 0.5 mm deep, shallowly concave

anteriorly with th ded, abruptly notched with a minute, sharp apiculum in the center of the

+ the kh. k ee “4 1 4 +h. |

Al. 4 1

ded with a low patch of short pubescence,

lender, 0.5 mm |

e 7 tants ee

thes be e h ged ays fo 1 nals ong, 0.8 mm wide, the foot

obsolescent, the anther apical.

Pichinch it Nanegali d Mindo, alt. 2000 m, 9 Sept. 2001, C. Luer & A. Hirtz 19896

(Holotype: MO).

This species stands tall with ramicauls ascending beside a stout, erect rhizome.

The raceme is elongate, subflexuous, loosely and successively flowering to far

exceed the leaf. The sepals are pubescent, five-veined and broadly ovate with

recurved margins. The round margin of the lip is abruptly notched with a central

apiculum as seen in the variation of S. argentata described as S. vulcanica.

49. Stelis distans Luer & Hirtz, Sp. nov.

Ety.: From the Latin distans, “distant,” referring to the remotely flowered raceme.

emon 1 A.

Planta parva caespi Tac li i I

: phonic i ee eae. : ee a ees : ral;

longiore, Sepalis latissime Ovatis trinervis lateralibus minoribus obtusis ad medium connatis, petalis

semilunati guitur,

Plant small, epiphytic, Caespitose; roots slender. Ramicauls erect, slender, 1-1.5 cm long, encl

by a tubular sheath and another | or 2 at the Leaf erect, cori

to acute, petiolate, 2-3 cm long including a petiole 1-1.5 cm lon

cuneate below into the petiole. Inflorescence an erect, lax, secund, several-flowered raceme 6-8 cm

long including the peduncle 3 ig, with most fl imul ly, from an annulus below

the apex of the ramicaul; floral bracts oblique, acute, 2 mm long; pedicels 1.25 mm long; ovary | mm

long; flowers red-purple; incompletely expanded,

the blade 0.4-0.5 cm wide, narrowly

ined,

1 mm into a shallowly concave, bifid s i

, ynsepal, 2.5 mm long, each sepal 2.25 mm wide, obtuse; petals

pra ovate, broadly rounded at the apex with the margin thickened, 0.75 mm long, 1.25 mm wide,

deep, shallowly cues ee rege mir iS gas sige

> in the rounded api argi i i

central callous, the base cuneate, hinged to the obsolecn nee eee the disc sb eel apc

mm wide, with the anth

ry sy Ameen Pada hee, oo alt. 1500 m, 20 Feb. 1986, C. Luer,

This species, known from southern Ecuador, is characterized by the small,

ae a narrow elliptical leaves somewhat longer than the ramicauls; and

seine oe " y eran owered, secund raceme. The dorsal sepal is broadly

siniited 2 sepals are smaller and semiconnate into a shallowly concave, bifid

“ne petals are transversely ovate, thickened on the rounded margin and

thickened i - :

cae aaa middle. The lip is ovoid, shallowly concave with a rounded

50. capes dodsonii Luer, Sp. nov.

— for Calaway Homer Dodson, co-collector of this Species.

TEND shes Soni bets rum

i habi ‘Species haec S. lindenii Linden persimilis, sed sepalis dense pubescentibus, petalo

Plant large, epiphyti : ‘ ° 3

25 cm long, with by ps Foes a nee Ramicauls erect, relatively slender, 14-

coriaceous, elliptical ; t the base. Leaf erect,

blade 3-4.5 cm wide nape tree petiolate, 9-15 cm long including the petiole 1.5 cm long, the

he into the le. Inflorescence 1-2 erect, strict, distichous, many-

STELIS OF ECUADOR 43

flowered racemes with several flowers It b loosely flowered below,

15-20 cm | g the es 4-5 cm long, with a na 1- 1.5 cm long, at an annulus below 7

apex of the ramicaul,; floral bracts oblique, acute, 2- 2. = mm long; pedicels 1 mm long; ovary 1.75 mi

fleshy, brown, with a dense, short, white pubescence within,

broadly oval te, erg tuse, none below the middle, the dorsal sepal 3 mm long, 2.5 mm wide, 3-

d, the lateral sepals 2 mm | i 25 mm jars ? veined; petals light | brown,

Scie obtuse, 0. 8 mm long, 1. 25 ta wide, 3-vein ith th d sub

concave bs _ — lip “ oo long aor ounsion, 1 -e long, 0.9 mm beat 0.5

ther side

J VY¥Yarw,

m dee} teriorly w

ofa pe aa Pi dorsum cee densely and minutely

hinged to the base of the column; column stout, 1 mm long, 1. 25 mm wide, the foot obsolescent, the

anther and the bilobed stigma apical.

sare west of Calacali and La Liberia, alt. 2100 m, 4 Apr. 1985, C. Luer, J. Luer, A. Hirtz, C.

& P. Dodson 11057 oo MO); same area, alt. 2200 m, 7 Apr. 1984, C. Luer, S. Dalstrém,

£ Hoijer & A. Hirtz 9996 (MO).

This large species is known from the western slopes of Pichincha near Quito.

Vegetatively, it is characterized by a broadly elliptical leaf, thrice shorter than the

ramicaul. Although the flowers are about the same size and shape as those of S.

lindenii, the sepals are densely short-pubescent within; the margins of the petals are

subverrucose; and the lip is oblong, longer than broad, concave above the middle

with the apex incurved and sharply apiculate.

iw ae. elongatissima Luer & Hirtz, sp. n

rom the Latin a, bind much h elongate, a seis to the —

a ta mediocris, racemo elong g giore is

ovatis obtusis, P : . . lahkell L ——

Pl i ts slender. Ramicauls — slender, 2-4 cm long,

mostly enclosed by a cballar Sa, and another sheath at the base. Leaf erect, coriaceous, narrowly

linear-elliptical, acute, obscurely ene 8-14 cm long including the bea 1- 2 cm long, the blade 0.5-

| cm wide, gradually narrowed below into the petiole. Inflorescence an elongated, flexi ible le, very distant-

ly many-flowered raceme = few eos open simultaneously, up to B cm long including the pedun-

cle 18- 20 cm a with a spath ee mm mites the sete of the ramicaul; floral bracts

long, 2 mm wide, 3-veined; petals purple, transversely oblong, broadly rounded at the apex, shallowly

concave below a slightly thickened margin, 0.75 mm long, 1 mm wide, 1-veined; lip purple, thickly

subquadrate, 0.75 mm long, 1 mm wide, 0.75 mm deep, shallowly concave anteriorly with the margin

wore rounded, the bar rounded with a shallow glenion, the dorsum densely and minutely pubescent

above the base, , the bane truncate, hinged to the base: of th Sones: column stout, 1 mm long, 1 mm

wide, th

orona-Santiago: Cordillera del Cutuct, along the ween Mendez and Morona, alt. 950 m,

18 Jan. 1989, C. Luer, J. Luer, A. Hirtz, S. Ortega, A. P Fa 14000 pe MO).

The inflorescence of this species found in the Cordillera del Cutucti is exceed-

ingly long with distantly spaced flowers. The leaves are narrowly linear, twice to

thrice longer than the ramicauls. The sepals are broadly ovate and similar, the

petals are single-veined, and the lip is semi-orbicular with a glanion, and pubescent

at the base.

52. asi a comepernts Luer, sp. no

From the Latin exasperatus, “atte shut, rigid points,” hasan to the petals.

: ellipticis longioribus, race-

indie haec S. hallii Lindl. affinis, sed a ese haan’ = iliatis pubes-

centibusque, talis i ¢ P ie differt

7 a

44 ICONES PLEUROTHALLIDINARUM

Pl ts slender. Ramicauls erect, slender, 3-11

cm aes enclosed by 2 ars eee and another 1-2 at the base. Leaf erect, coriaceous, narrowly

elliptical, acute, 4.5-6.5 cm long including a petiole 1- i cm long, the blade 0.8-1.5 cm wide, narrowly

cuneate below into the

1-3 erec' congested, distichous, ar uas es racemes,

uate nen he ie y, with a slender

the ca. 8 mm long, f th Saal Noval bli acute, 1.5-2

anests ovate, obtuse,

erect connate = iy ae sac the dorsal cata 2.25-3.6 mm tewntag 3.6 mm wide, 3-veined, the

lateral pe eh, 2 mm long, 2.3-3 mm wide, 3-veined ed; petals green of be transversely semi-

mm wide, broadly tt pex broadly rounded,

0 aa ag 073mm wide, 0.5 mm deep,

p y rounded g y a small glenion, the

dorsum with a central, round, pub Illus, the base b sg ce to the base of the

column; column stout, 0.6 mm long, 0.6 mm wide, the foot =e Sy the anther and the bilobed

apical.

SD e >

oh a Al. rT a Ee | ‘ h. 41

Carchi: forest between Tulcan and Maldonado, alt. 2050 m, 2-4 Apr. 1984, C. Luer, S. Dalstrém & T.

Héijer 99. MO) ta, C. Luer, S. Dalstriém & T. ee (MO); same

ace 1950 m, 2-4 Apr. 1984, C. Luer, S. Dalstrim &T. bier 9915 (MO).

Vegetatively, this species from northernmost Ecuador is distinguished by

slender ramicauls that are longer than the narrowly elliptical leaves. One to three

congested racemes with small, widely spread flowers basically similar to many

species surpass the leaves. The sepals are are broadly ovate and microscopically ciliate

and pubescent. The petals are broadly thickened and covered by minute, sharp

points along the rounded apex. In some specimens the points are reduced to cells.

ps lip is semi-orbicular with a small glenion, and a single, rounded callus on the

rsum.

53. Regge omaran ene & Hirtz, sp. n

avatus, “with a seaviy” i to the concave labellum.

Planta it

pie seta vata a lens pcp foliis ang is longiore, sepalis similibus

E gui

bb ne ne oo shortly ascending-repent, the rhizome stout, ca. 5 mm between ramicauls;

— acceiiat ge me. Ramicauls erect, slender, 6-11 cm long, enclosed by a tubular sheath

: 2 anal sg tubular sheaths below. Leaf erec coriaceous, narrowly elliptical, acute

Fit wate cm long, ne a nish 1- 15 cm long, the blade 1.8-2.4 cm wide, cuneate below

gested, subsecund, distichous, many-flowered raceme with

many fl taneous y

me ig ta i open si smal sly, 15- 1S- i701 = long including the peduncle ca. 1 cm long, with a spathe

ute

apo a a Prong above; pedicels 1 5-2 mm long; ovary 1 mm long; sepals yell iow, glabrous external-

Pads = cae — , oe y, the dorsal sepal 3 mm lon, ng, pee m wide, 3-veined,

ia ong, 2.8 mm wide, Seine petals purple, transversely ovate, with ie cae

rap Pyooage with a thick, flat

on mm

oan sey oe lip re i sabspheoi, 0. scan mm cn pl 0. 8 mm sigs £ 6mm deep, decoy

dorsum slightly convex with - bo , the

ace, truncate basally, hin to the h Poe . uous 1 the anterior

anther and the bilobed sti eeenties ppamminaly Aaceadierny the

Morona-Santiago: aroun.

Campamento G

Andreetta & L. tap = = alt. 1650 m, 24 Mar. 1991, C. Luer, J. Luer, A

Pei «deena oseren species is apparently restricted to southeastern

than the narrowly sm ipoeet yg a strict, crowded, many-flowered raceme longer

The lip is thick, subspherical, The sepals are equally ovate and pubescent within.

angled above, continuous aac anteriorly with protruding margins, obtusely

rounded apex, the bar deeply concave anteriorly.

STELIS OF ECUADOR 45

54. Stelis exilis Luer & Hirtz, sp. n

Ety.: From the Latin exilis, ee, shin,” = referring tc to the habit.

Di + di

multifloro foliis anguste ellip-

4 ba tr, lat, hinen et

ticis multilongiore, aioe latissime ovatis obtusis, petal

lahall hk A + bate “4 i As nee

= (—)

ot aan in size, epiphytic, —— caespitose; roots eee: ce Ne — Mae 1.5-2

sed by a tubular Leaf erect, arrowly

poet’ acute indistinctly petiolate, 3-5.5 cm long | including the petiole 1. 5-2 cm mong i blade 4-5

wide, narrowly cuneate below into the petiole. lax, distichous,

flexuous, ream} many-flowered raceme, 2-3 flow rs open simultaneous! , 15-30 em long includ-

ing the peduncle 10-11 cm long: floral bracts cighaten acute, 2 mm long; pedicels 1 mm long, enclosed

within the a ovary | als purple ] long-pubescent within,

transversely ovate, obtuse, th dors al sepal 1.8mm aaa 2 1 mm wide, 3-veined, connate basally to

— — the lateral sepals 1.5 mm long, 2 ma wide, 3- veined, petals purple, transversely ovate,

with the Os e, 0.8 mm lon. ong, 1 mm

wide, 3 ‘aah, shallowly concave b below the apex; lip purple subovoid, obtuse, 0. 6 mm long, 0.8 mm

ide, 0.4 mm deep, th concave with a small glenion, the dorsum with 3 low,

wae calli, the base truncate, hinged to the base of the column; column stout, 0.6 mm long, | mm

wide, the foot obsolescent, the anther and the bilobed stigma apical.

Zam sey gg Cordillera del Condor, east of Paquisha, alt. 1450 m, 19 Feb. 1986, C. Luer, J.

Luer, A. Hirtz, W. Flores & A. Embree 11937 (Holotype: MO).

This species is apparently endemic in southeasternmost Ecuador. It is character-

ized by a densely caespitose habit with narrowly elliptic, petiolate leaves that are far

surpassed by an elongating, flexuous, successively flowering raceme. The sepals

and petals of the tiny flowers are wider than long. The lip is shallowly concave

anteriorly with a small glenion, and with three small calli on the dorsum.

55. Stelis falcifera Luer & Hirtz, sp. n

Ety.: From the Latin pia ‘sickle- bering “ "refering to the —

Planta mediocris prolifican i : aie ee

spathis propartions aed acutis, ‘sepalis ellipticis obtusis otion oi liberis, petalis transversis apice

re Itilonoio

“Ss

Plan um in size, epiphytic, climbing; roots slender. Ramicauls erect, stout, prolific, 5-7

ct moar hens by a tubular sheath and another sheath at the base. Leaf erect, co cairo bl cipal

uneate

petiole ‘Inflorescence an erect, congested, distichous, many-flowered raceme, many canes i

simultaneously, 2 cm long — the peduncle 2-3 cm long, from an acute, slender spathe 1.5-2

cm long; floral bracts tubul mm long; pedicels 1 mm long, enclosed within the bract; ovary | mm

long; sepals eaeien ae aed ovate, obtuse, the dorsal sepal 3 mm long, 2.5 mm wide,

veined, shortly connate basally to the lateral sepals, the lateral sepals nearly free, 2.5 mm long, 2mm

wide, 3-veined; petals dark eae transversely elliptical, concave, broadly ob

thickened, 0.85 mm long, 1.1 mm wide, 3-veined, shallowly concave below the apex; lip dark purple

— €, obtuse, mm sei ‘0. 85 mm wide, 0.5 mm deep, the anterior surface shallo wly concave

with a genion, the dorsum with a low, —— oe; the base truncate, — to the base of the

t, 0.6 mm long, tt nt, the anther and the bilobed sti

Pchince west of Calacalf, alt. 2550 m, 4 Apr. 1985, C. Luer, J. Luer & A. Hirtz 1052 (Holotype:

This species is apparently endemic in west- central Ecuador. It is characterized

by a prolific habit that creates a series of ramicauls up to a meter long. From the

base of each elliptical, petiolate leaf the next ramicaul is produced, along with a

long, narrow, curved spathe, and the inflorescence. The cylindrical raceme, much

longer than the leaf, is tightly packed with numerous, small, overlapping flowers.

The sepals are white, the dorsal sepal being slightly larger than the lateral —

while the central figures are dark purple. The lip is subquadrate with a glenion an

a callus on the dorsum

46 ICONES PLEUROTHALLIDINARUM

56. inenoane es rbcneiediini Sp. nov.

the Latin ferax, “fruitful, th

ti . ne nent,

t fee

‘ . i abt . ngs | eee ‘ . ; i j

pr eee distich Itifl fol p longi petiolatis longiore, floribus

1 1. £, ech. 1 ape Bitine. a. rh =e. KI Ag

| r 7 —— + =. 7 Ties:

, o a A tos ie ne. ie

my i

Plant Nt dium in size, epiphytic, it lend . Ramicauls ane a 2-7

cm long ] bular sheath fr below th 2 sheaths at the base. Leaf

erect, thinly coriaceous, elliptical, acute, petiolate, 5-10 cm sent aaoke the petiole re 2 cm nih the

blade 1.5-2.5 cm wide, cuneate below into the slender petiole. Inflorescence an erect, congested, dis-

tichous, -flowered raceme 7-15 cm long, a, the peduncle 4-7 cm long, with few

fl imul mm long including the peduncle 5-9 cm

a thin spathe ca. 5 mm long, a at an annulus below the apex of the ramicaul; floral

tects srg acute, 2mm long; pedicels | 7 mm long; ovary 1 51 mm long; cite Opes ini ee

ong, 2.4

wide, 3-veined, | larger th lat Is, the lateral sepals obli sg | mm lon. 12 2 mm aie %: 3-

veined; petals yellow miia terns oblong, shallowly concave, 0.5 mm long, 1 mm wide, 3-

veined, d; lip ye ello w-white, thickly subquadrate, 0.

long, 0.6 mm wide, o 6mm deep, th the anterior surface convex P dly rounded, minutely

apiculate, the bar thic tudi i the di : pubes-

cent, the b d to the b f th nn; column stout, 0.6 mm long, 0. 6mm wae: the

amcor eee ee ae

: near the pass eet alt. 2400 m, 16 Mar. 1991, C. Luer, J. Luer, J. del

Hierro, A. & X. Hirtz 15084 (Holotype: M

This species is apparently a in northernmost Ecuador. It is peace

by an acute, elliptical, long-petiolate leaf about as long as the slender ramicaul.

slender raceme of numerous, tiny, successive flowers surpasses the leaf. The ei

ers are probably self-fertilized, because practically every flower develops a capsule.

The sepals are pubescent and broadly ovate, the laterals being smaller and oblique.

The petals are transversely oblong. The minute lip is similar to those species relat-

ed to S. argentata with a marginal apiculum.

57. Stelis flexuosissima Luer & Hirtz, Sp. nov.

Ety.: From the Latin flexuosissimus, “very flexuous,”’ eionring to the vigzag raceme.

Species haec S. concinnae Lindl. affinis, sed habitu mul j

cuosis differt, $ :

BP aiey medium i m size, epiphytic, caespitose; roots slender. Ramicauls erect, slender, 6-8 cm long,

tical, acute, 7-8.5 cm long including a petiol and 2-3 sheaths at the base. Leaf erect, coriaceous, ellip-

sie ie pedicle aan 8 nga petiole ca. 1 cm long, the blade 1.2-1.7 cm — cuneate below

flowe i, Pea ith flexible, flexuous, subcon nges

me with 1-4 flowers simultaneously, 12-23 em long including ts peduacie 6-7 cm long,

jue, acute, 5 mm lon cels 2-2.5

ane ee ents spa purple, al nthe margins, similar, ovate with ec te margins,

3-veined; petals — a n elliptic ena: basally, 3 mm long, 2 mm wide expanded,

wide, 3-veined, the apex tsely €: ipicaLoblong, thin, shallowly concave, 1 mm a ran 5 mm

1 mm long, 1 mm wide, 0s y rounded, narrowly thickened on the margin; lip purple, oblon

hegwheaiéanaes broadly rounded, thickened centrally, Siediowly ere

‘ow omar. minutely pubescent at the base, the base truncate,

hinged to the base

aavatice of cole: ¢aleme: st out, 1 mm long, 1.2 mm wide, the foot obsolescent, the

Zamora-Chinchipe: south

Hirtz, W. Flores & A. cave taeleqee jimbura, alt. 3050 m,21 Feb. 1986, C. Luer, J. Lue, A

This species is little more than a gigantic S$. concinna. Except for the larger size,

flexible, lex are similar with the sepals widely diverging with revolute sides. The

ey racemes are also similar, but the floral bracts at each angle are

and leaves are eee times larger. - Vegetatively, there is no similarity. The ramicauls

STELIS OF ECUADOR 47

58. Stelis floresii Luer & Hirtz, s

Ety.: Named for Washington Flores of Guin, Ecuador, co-collector of this ee

_ Species haec S. hallii L indl similis d P breviore, sepa-

Pe he «Ae

nt medium in size, — ee caespitose; ~ — Ramicauls erect, slender, st :

cm nie enclosed by 2 loose, tubular ths, and another at . Leaf erect, coriaceous, narro

tise acute ae saci petiolate, an cm ‘long oe = hei 1.5-2.5 cm long, the blade 1- 1 3

e below into the petiole. Inflorescence an erect, subcongested, distichous, simultaneous-

= me raceme, “<i 13 cm long including the sta 3-4 cm long, from an annulus below the

apex of the ramicaul; floral sikesiol acute, oblique, 3 mm long; pedicels i 5 mm long; ovary ie reek mm

diffusely inutely

ie ‘broadly ants obtuse subequal, connate basally, the decal sepal 4 mm long, 3.25 mm Stic. 3-

eined, the lat epals oblique, 3.5 mm long, 3 mm wide, 3-veined; petals yellow- green, semilunate,

0. 75 mm ng + ‘ mm wide, shallowly concave below th broadly

a — concave at the base; li ¢ purple, subquadrate, 0.75 mm long, | mm wide, 0.75

m deep, the apex — rounded, aura concave anteriorly with a faint, superficial glenion, the

oe with a low, rounded, pubescent callus, the base broadly truncate, hinged to the base of the

column; column ion. ‘t mm long, 1 mm wide, the foot obsolescent, the anther and the bilobed stigma

ical.

ua: Cordillera de los Llanganates, Leito, alt. 2870 m, 11 Apr. 1985, C. Luer, J. Luer, A. Hirtz

& W. Flores 11139 (Holotype: MO).

rona-Santiago: Coedilieee del Condor, east of pit alt. 1500 m, 18 Feb. 1986, C. Luer, J. Luer,

A. Hirtz et al. 11904, 11913 (MO); east of the pass between Gualaceo and Limén, alt. 2650-2750 m, 16

Feb. a ia Luer, J. Luer, A. Hirtz, W. Flores & A. pone 11829, 11838 (MO)

ve Yangana north of the pass, ais 2400 m, 23 Mar. 1985, C. Luer, J, Luer, A. Hirtz & W.

Flores 10871 (MO).

ra-Chinchipe: east of the pass east of Loja, alt. 2750 m, 21 Mar. 1985, C. Luer, J. Luer, A. Hirtz

& W. Flores 10721 (MO).

This species is very similar to S. hallii, and it possibly partly accounts for the

reportedly wide distribution of the latter. Vegetatively, S. floresii is distinguished

by wider leaves that are more distinctly petiolate. Both species produce a raceme of

flowers of similar qualities and quantity. The sepals of S. floresii are densely

pubescent, and the petals and lip are proportionately larger in comparison to the

sepals than those of S. hallii.

wt f£ the plant

59. Rca bpp Luer & ne 5 nov. ;

Latin formosus, “beauti Ter P &

anta randis, k Pyar | yd beet I Itil gio re, bracteis

orbs sconspiuis, =e diocrib palis ovatis obtusis breviter pubescenibs, petal telies

rse o b i

Plant ge epiphytic, caespitose; roots slender. Ramicauls erect, stout, 4-10 cm long, wi with a lo

tubular she nother 2 sheaths at the base. Leaf erect, coriaceous, elliptical,

ee ahaa: 5-8 cm long including the petiole 1-1.5 cm yon the blade 2-2.5 cm wide, ager

below into the e petiole. Inflorescence an erect, strict, c congested, distichous many-flowered raceme ae

pon pode rs open aeerrTETy 15-23 cm long including the “soaps 5-9 cm long, with a er

15cm sie at an annulus below the apex of the ramicaul; floral bracts c : nspicuou congas acute,

mm peli apg vary 2 mm long; sepals rosy tan, tan, tee iow, similar,

externally, — short-pubescent within, ovate, obtuse, connate basally into a short cup, the dorsal

sepal 4 mm long, 3.5 mm wide, 3-veined, the lateral sepals 4 mm ree 3mm _ 3-vel

yellow-green, transversely oblong, shallowly concave, 2 mm long, 2.25 mm wick,

apex broadly subtruncate and thickened; lip yellow-green, thickly subquadrate, 1.25 mm lon 2mm

wide, 1.5 mm , the anterior surface concave with the apex broadly sent the vende oe tae

large, oe concave ery the dorsum shortly pubescent, the base ee a

m long, 2 mm wide, the foot Pe the an es the bi

apical.

48 ICONES PLEUROTHALLIDINARUM

Loja: army road to TV tower east of Yangana, alt. 3100 m, 22 Mar. 1985, C. Luer, J. Luer, A. Hirtz &

W. Flores 10790 (Holotype: MO); llection data, C. Luer et al. 10785, 10788 (MO).

This species is apparently endemic in southeastern Ecuador. It is distinguished

by a stout ramicaul with a loose sheath near the middle, and a large, obtuse, ellip-

tical leaf with a distinct petiole. The floral bracts of the raceme are conspicuous,

creating a saw-like appearance. The sepals are densely pubescent, ovate and form a

basal cup. The lip is concave anteriorly and the surface of the bar has a deep, circu-

lar glenion, while the dorsum is flat and shortly pubescent.

60. Stelis gigantea Luer & Hirtz, sp. nov.

From the Latin giganteus, “gigantic,” referring to the size of the habit and flowers.

Planta inet reese On slecert £1 Con ne |

floribus grandissimis, sepali

cuneato concavo distinguitur.

Iti tate allineic. lel

menso Pp g ¥

eco s 8 ‘ ‘

t i: lab tal et labello

Plant very large, epiphytic to terrestrial, caespitose; roots slender. Ramicauls erect, stout, 35 cm or

more long, with a tubular sheath near the middle and another 2 sheaths at the base. Leaf erect, coria-

dly elliptical-ovate, obtuse, sessile, 25 cm long, 20 cm wide, the base rounded. Inflo nce

an erect, strict, congested, distichous, many-fl d itl fl pen simultaneously, 75

cm long including the peduncle ca. 15 cm long, with a spathe ca. 5 cm long, at the base of the leaf: floral

ts oblique, obtuse, 3 mm long; pedicels 1.5 mm long; ovary 3 mm long; flowers dark purple; sepals

fleshy, glabrous, ovate, obtuse, connate basally, the dorsal sepal of lower flowers 14 mm long, 7.5 mm

wide, of upper flowers 9 mm long, 6 mm wide, 7- to 9-veined, the lateral sepals of lower flowers 13 mm

concave, broadly rounded and thick at the apex, 1.75 mm long, 2.5 mm wide, 3-veined; lip subcuneate,

1 mm long, 1.5 mm wide, | mm deep, the anterior surface concave with the apex broadly rounded,

it pee transverse bar with a shallow glenion, the dorsum with a low, subtrilobed callus, the

base truncate, hinged t ft lu ut, 2 mm long, 2 mm wide, the foot obsoles-

cent, the anther and the bilobed stigma apical.

ee pc alt. ca. 1000 m, 11 Aug. 1991, A. Hirtz & S. Ortega 5351 (Holotype: MO), C. Luer il-

praees ; ; south of Baeza toward Tena, alt. 1500 m, 11 Aug. 1978, C. Luer, J. Luer, A. Hirtz & A.

‘ta 3242 (SEL); south of Baeza, alt. 1900 m, 20 Feb. 1982, C. Luer & A. Hirtz 6861 (SEL).

quarters of a meter long. The lowermost flowers are the largest, measuring 27

millimeters from the tip of the dorsal sepal to the tips of the lateral sepals. The

Sepals are ovate, glabrous and spreading; the petals are thickly fan-shaped; and the

lip is small, cuneate, and concave anteriorly.

61. Stelis glaberrima Luer & Hirtz, Sp. nov.

a ce - Latin glaberrimus, “very glabrous,” referring to the sepals.

anta parva. foliic ellini-3 . ave ed peti ‘ : é

duplolongiore, sepalis ovatis obtusis plaberrimis, er i pie ca folio plus minusve

Plant small, iphytic. itose: roots :

by a loose, ysl ie oe slender. Ramicauls erect, slender, 1-1.5 cm long, enclosed

cm long including a petiole ‘ — at the base. erect, coriaceous, elliptical, subacute, 2.5-4.5

lulliveicente 2 stating the blade 0.9-1.2 cm wide, cuneate below into the petiole.

. ae us, s¢veral-flowered race i fl imulta-

neously, 5-7 cm | ludin owers open sim!

a annals scing the peduncle ca. 2.5 cm long, subtended by a slender spathe ca. 5 mm

x oral bracts infundibular, oblique, acute, 3 mm

5 mm long, 4.5 mm ide i

lateral sepals obli wide, 5-veined, connate basally to the lateral sepals, the

1 mm long, 2 a mm wide, 4-veined; petals yellow, transversely obovate-oblong,

~ thickened and flat; lip yellow, thick,

ape, with « glenior nf 1-5 im Wide, 1 mm deep, shallowly

Pubescent, decurved along the shaft : he protruding bar, the base minutely

wid of inged toa the L oa —

1.5 mm wide, the bas, ob the the column hinged othe base of ; pis

STELIS OF ECUADOR 49

Napo: forest along the road from Baeza to Cosanga, alt. 1800 m, 14 Apr. 1985, C. Luer. J. Luer, A. Hirtz

& W. Flores 11273 (Holotype: MO).

This species, found at moderate altitudes on the eastern slope of Ecuador, is

characterized by a small habit; elliptical leaves considerably longer than the rami-

cauls; a several-flowered raceme exceeding the leaf; comparatively large flowers

with totally glabrous, broadly ovate, five-veined sepals; standard, transverse petals;

and a shallow, subquadrate lip.

62. ee globulifera Luer & Hirtz, sp. n

.: From the Latin globulifer, “‘bearing a little, aaa ball,” referring to the labellar callus.

ie anta parva oe nial pluri- successivifloro folio late era leviter excedenti, sepalis

obtusis, petalis cra

Plant small, oa caespitose; roots slender. Ramicauls erect, slender, 7-10 mm ae enclosed

by 2 tubular sheath . Leaf erect, thickly coriaceous, broadly elliptical, subacute to obtuse, 2.5-3 cm os

including a petiole % 10 mm long, the blade 1 cm wide, cuneate below into a petiole. I

erect to suberect, sehr “i es lanes several- flowered raceme, at cm long

including the peduncle 2-2.5 with 1 e leaf;

floral bracts oblique, mente 2. a8 mm mong ids 2 mm long; ovary 1.75 mm aes flowers expended,

thin, transversely ovate, connate basally, the dorsal sepal 3

mm long, 4mm w wide, 5- -veined, the: inten sepals oblique, 2.5 mm long, 3.5 mm wide, 4-veined; petals

thick, transversely oblong, 0.75 mm long, 1.25 mm wide, 3-veined, thickly rounded above the concave

base; lip thick, transversely oblong, 0.5 mm ong, | mm wi “ty 0.5 mm — ig anterior surface shallow-

ly concave, the apex thick, broadly rounded, the dorsum with a proporti y large, spheroid callus that

protrudes anteriorly instead of a glenion, the base cela hinged to ‘the pe of the column; column

stout, 0.75 mm long, 1 mm wide,

Es das: wet forest west of Lita, alt. 750 m, 18 Jan. 1987, C. Luer, J. Luer, C. Dodson, A. Hirtz & D.

Benzing 12413 (Holotype: MO).

This little species is best distinguished by a distinct, spherical callus on the

dorsum of the lip that protrudes anteriorly where a glenion would be expected to

occur. The callus is a modification of the midvein of the lip, which is frequently

seen as a callus, often between a smaller callus on either side, products of the lateral

veins. The plant is small and caespitose with broadly elliptical leaves surpassed

slightly by a successively several-flowered raceme. The flowers are “standard.”

The sepals are expanded and transversely obtuse. The petals are transverse and

thickly rounded above the base.

63. ane hoeijeri Luer & Dalstrém, sp. n

ty.: Named for Thomas es of Stockholm, Sweden co-collector of this —

Siegen prolificans, racemo disticho multifloro folio anguste ellipico malilongioe., eri

imbriacantibus, sepalis ovatis Dultees li i ilunatis et lat is

Plant large, epiphytic, proliferating; rooting at the base. Ramicauls stout, erect, to 25 ‘om besied

below, branching to slender above, 4-10 cm long, with a tubul ar sh

1-2 sheaths at atthe inn. Leaf erect, coriaceous, narrowly elliptical, acute, petio late, 7- 11 cm long includ-

ing the petiole 2 cm long, the blade 1-1.5 cm wile cuneate below into the petiole. Inflorescence an

ae — simultaneously many-flowered raceme, with the flowers overlapping, to 25 cm long

ding the peduncle 10 cm long, subtended by a spathe ca. 1 cm long, from an annulus below —

ofthe ramicaul; floral bracts thin, tubular, repand, = 6 mm long; pedicels 3-4 mm long; ovary om

long; sepals green, suffused with purple basally, expanded, glabrous, ervntt; csi connate em *

similar, 5 mm long, 4.25 mm wide, 5-veined; petals purple, thickly e bas : an

low the broadly obtuse apex, 0.6 mm long, 0.9 mm wide; lip thick, ime aon long, 0. ~

= 0.75 mm deep, concave anteriorly, the apex subacute, the bar thick with a glenion, the eye wi

rounded callous, the base bro. roadly truncate, a to the base of the column; column stout, 0.75 mm

rig 0.75 mm wide, the obsolescent.

anther

Carchi: fore st between a and Maldonado, alt. 1950 m, 2-4 Apr. 1984, C. Luer, S. Dalstrim & T.

Héijer 9912 (Holotype

50 ICONES PLEUROTHALLIDINARUM

This species, apparently endemic in northernmost Ecuador, is characterized by

the prolific, branching ramicauls with narrowly elliptical leaves. The flowers are

overlapping in a congested raceme that far surpasses the leaf. The sepals are ovate

and similar; the laterals reflex from their common union which is convex beneath

the central apparatus. The petals are thickly semilunate with a sharp transverse

callus. The lip is triangular with a glenion on the bar, and a callus on the dorsum.

64. gy —- Luer & Hirtz, sp. no

the remote community of prart near where the species was collected.

Diane As, ae Ninsi ee | 1 . e+ | 3 5 CR eanenL ee

- s * s a ~ m _s

. . edly Se .

Plant ~~ ee caespitose; roots slender. Ramicauls erect, , relatively stout, 9-20 cm long,

with a loose, t erect, coriaceous,

clipart acta: petiolate, 8-14 cm long ee the petiole 1.5-2 cm long, the blade 2-3 cm

eate below i into the petiole Inflorescen -4 erect, Strict, distichous, subcongested, many-

ith fl :

poner including the peduncle ca. 3 cm long, with a the 1 .2 cm long, at an annulus below the apex of

the the ramicaul; floral bracts aaanes dilated, acute, pee mm long; pedicels 2 mm ome ovary 2 mm long;

thickened

verse callus, the apex ly rounded, 1.3 mm long, 2 mm wide, $-veined: lip brown, obliquely a.

quadrate, 1 mm long, 1.8 mm wide, 0.8 mm deep, the anterior surface concave with the a apex broadly

h of Jimbura, alt aie m, 21 Feb. 1986, C. Luer, J. Luer,

pon Ww. Flores & A. Embree 11995 (Ho MO)

£7 Wier Jone Ono lotype: 2800 m, 26 Jan. 1986, S. foes

This large species is known only from southeasternmost Ecuador. A few

racemes of relatively large flowers are about as long as the elliptical, petiolate leaf.

The synsepals are deeply connate into a broad, non-concave, bifid synsepal. The

petals are broad and thick. The lip is concave above the rounded apex.

65. Stelis k Luer & Hirtz,

Ety.: Named for Job Kuijt of Victoria, British Columbia, co-collector of this species.

Species S. _atroviolaceae Rchb. f. affinis, sed sepalis plurinervis, petalis grandis columnam

Plant large. arge, epiphytic to terrestrial, dens

Y Caespitose; roots slender. Ramicauls erect, stout, 8-14

cm Jong, an a loose, tubular shea ‘ew below the middle, and 2-3 at the base. Leaf erect,

» Subacute to obtu otuse, petiolate, 9-13 cm long including a pomele 1.5-2 cm long, the

e. Infloresce

many-flowered neem, gig cm “ee including the peduncle 2-3 cm long, with a spathe

Ts mm

connate ah terals for 3 mm, the lateral sepals broadly ovate,

Proportionately inege, eat = for: 3 ae 3 mm long, 4. 5 mm aera , 4-veined; sb purple,

apical margin bro rounded, flatte ly semilunate, the

bi the eco lip ed and thickened, 1.5 mm long, 2 mm wide, Seine: concave

bisa rm amccory with the eed mekly subquadrate, 1.3 mm long, 1.3 mm wide, 0.6 mm

ow, min ely pubescent

1.5 mm long, the calli, the base truncate, , hinged to the base of the column; column stout,

foot obsolescent.

ancave

Bolivar: terrestrial on

‘hake cee te ead embankment ol road west of Guaranda toward Pueblo Viejo, alt. 3100

trim, T. Héijer, J. fos 8 A Atlee 9740 (Holotype: MO); old road

between Guaranda and and Caluma, alt. 3100 m, 10 Mar. 1982, C. Luer & § S. Dalstrim 7237 (SEL).

STELIS OF ECUADOR 51

This handsome species of high altitudes of south-central Ecuador is character-

ized by stout ramicauls clothed in a loose sheath and a large, elliptical, long-pe-

tiolate leaf. The raceme with large, purple bracts and flowers barely surpasses the

leaf. The lateral sepals are connate to near the middle into a broad, bifid synsepal

suggesting section Humboldtia. The proportionately large petals with a sharp,

transverse callus overlap above and below the column and lip. The lip is concave

above the round apex, with the dorsum tricallous.

66. Stelis limonensis Luer & Hirtz, sp. n

Ety.: Named for the town of Limén, above lich this species was found.

Species haec floribus S. atrae Lindl. similis, sed habitu dense caespitoso, foliis anguste ellipticis

nt medium in size, epiphytic, hinds caespitose; roots slender. Ramicauls erect, slender, 5-11

cm long, enclosed by a tubular sheath on the middle third and 2 others below. Leaf erect, coriac

narrowly ell oe acute, 6-7 cm long including a petiole 1.5 cm long, the blade 1-1.2 cm wide, narrow

ly cuneate below into the petiole. Inflorescence an erect, loosely oe raceme with several

e ped annulus below

the apex of the a floral bracts infundibular, oblique, acute Hie mm ne pedicels 2.5-3 mm

long; ovary 1.5 mm long; sepals expanded, densely short-pubescent within, gray-green, broadly ovate,

obtuse, connate <a than 1 mm below the pees! a eek, “i 35 mm long, 3. 5 mm wide, 3-veined,

the lateral ce 3 mm long, 3 mm wide, 3-vein

ansverse thickening, the apex broadly obtuse lym a ae margin, 1mm m long, 1.25 mm wide; lip

yellow, thick, mca ovoid, 0.5 mm long, | mm wide, 0.5 mm deep, the apex broadly rounded,

shallowly concave anteriorly with a small, indistinct glenion, the dorsum with a low, rounded callus

base broadly truncate, hinged to the base of the column; column stout, 0.75 mm long, | mm wide. the

oot obsolescent, the anther and the stigma apical, the stigma bilobed.

Morona-Santiago: Ave Maria Pass between Gualaceo and Limén, alt. 2900 m, 16 Feb. 1986, C. Luer, J.

Luer, A. Hirtz, W. Flores & A. Embree 11816 (Holotype: MO).

Except for the color and pubescence, the floral parts of this species are similar to

those of S. atra. Instead of deep blue-purple, the sepals are gray green. The flowers

are borne on a long, loose raceme, several times as long as the narrowly acute

leaves. The sepals are broadly ovate; the petals have a sharp transverse callus; and

the lip is ovoid, shallowly concave anteriorly with a rounded callus on the dorsum.

67. Stelis longihirta Luer & Hirtz, sp. n

Ety.: From the Latin cages en ee refering to the acent

Planta pary g is pl i pees longiore,

petalis navanh ae | - : ‘ “28 I ra : . . lakall -

Plant very small — very slender. Ramicauls erect, slender, 0.5-

I cm — — by a tubular sheath from below the middle and moat 1-2 other sheaths at the base.

riaceous, ree narrowly apie subacute, 2-3 cm long with an indistinct petiole,

0.5- Le cm m long, the blade 3-3.5 mm wide, gradually narrowed into the petiole. Inflorescence an erect,

lax, success al-flowered raceme to 5 cm ae including the _ 3-3.5 cm long, from an

annulus below a open of the ramicaul; floral bracts oblique, acute, 1.5 mm long; pedicels 1 mm long;

ovary | mm long; flowers red-purple; sepals long-villous, the dorsal sepal erect, 1.75 mm long, | 25 mm i:

wide, ——— 3-veined, convex, the lateral sepals ovate, obtuse, barely connate, 1.5mm —_ i.

mm wide, incompletely 3-veined, more or less held forward, opposite when expanded;

g-obov

stout, 0.75 mm long, 0.75 mm wide, the foot obsolescent, the anther and the bilobed stigma ap

Sealey ary ok cloud forest north of the pass north of — alt. 1700 m, 17 Feb. 1986, C-

J. Luer, A. Hirtz, W. Flores & A. Embree 11878 (Holotype: M

This little, caespitose species is characterized by a loose, few-flowered raceme

borne about twice as high as the narrow leaves. The purple sepals are narrowly

52 ICONES PLEUROTHALLIDINARUM

ovate with a long, villous pubescence along the margins. The lateral sepals spread

in opposite directions. The broad, transverse petals are thin and single-veined with

a anerrowly one er = Bp is oblong-triangular, shallowly subacute and

Ja?

Stelis longissima Luer & Hirtz, sp. n

me From the Latin longissimus, “the vie. desi of longus,”’ referring to the proportionately very

raceme.

Plent. oo L Fs] £12: = a

t -

nese netali . * . + lekall . en L J hr

f -

longiore, sepalis late ovatis

Atots: ne

Plant small, epiphytic, caespitose; roots filiform, Ramicauls erect, slender, 1-2.5 we pail enclosed

ode 2-3 tubular sheaths. Leaf erect, coriaceous, narrowly elliptical, acute, 3.5-5 cm long including an

tinct petiole ca. | cm long, the blade 0.5 cm wide, narrowly a below into the petiole. Inflor-

cane an erect, lax, » flexuous, successively many-flowered raceme, with a few flowers open

simultaneously, 20-45 cm pe including the peduncle 67 cm long, from an annulus below the apex of

ramicaul with a spathe oblique, acute, 2 mm were! pedicels

2 mm long; ovary 2 mm long; sepals expanded, dull yellow above the middle, dull purple-brown below

the middle, glabrous externally, minutely pubescent within, broadly ovate with edges ashes, obtuse,

connate basally, the dorsal sepal 2.6 mm lo ong, 3 mm wide, 3-veined, the lateral sepals oblique, 2 mm

long, 2.6 mm wide, 3-veined; petals purple-brown, thick, transversely triangular-ovate, concave at the

base, thickened across the middle, the st broadly obtuse with a thickened, subverrucose races a 75

mm long, | mm wide, 3-veined; lip purple-brown, thickly iliesilic-iibemeaanc 0.6 mm long, 1

wide, 0.6 mm deep, barely concave secon if at all, the apex triangular, obtuse, the bar thick, we a

mannons, shallow gl th indistinct, low callus, the base broadly truncate, hinged to

e Ih tout, 0.8 mm long, 0.8 mm wide, the foot obsolescent, the anther and

Morona-Santiago: Cordillera del Condor, east of Chuchumbletza, alt. 1750 m, 21 May 1988, C. Luer, A.

Hirtz, W. Flores, A. Andreetta & W. Teague 13554 (Holotype: MO); near Paute dam, alt. 1550 m, 25 Jan.

1000 m, 5 Feb. 1987, C. Luer, J. Luer & A Hirtz 12654 (MO): same are (

F a, alt. 950 m, 17 Jan. 1989, C.

Luer, J. Luer, A. — & P. Jesup 13960 (MO).

es, known from the Cordillera del Condor, is distinguished by a small,

thrice or more longer than the leaves, is loose, distichous, and flexuous. The se

are expanded and obtuse; the petals are thickly triangular with a transverse ic

and the lip is nearly flat or barely concave anteriorly with a narrow glenion.

69. Stelis a eactiente Luer & Hirtz, sp. no’

mas Latin Scag fase eel,” eens to agape of bees gates

mediocris dense P

loge ato is ob Hatis ti

apiculato cum callo rites €t margine bimammillata p-aese

Plant medium

dl mio nedium in size, epiphytic, densely caespitose; roots slender. _Ramicauls erect, very Lion =

cous, narrowly elliptical, acute, petiolate, +75 om long includin the petiole 1.5-2 cm long, the blade

05-06 ne cm wide, narrowed below i into the pet iole. . Inflorescence an not ce atic rosea gested,

et labello sgancncdiad

te he ede Sm ng, with v5 aia ca. 3mm long, at an annulus below the apex of the ramicaul;

sepals similar, glabrous, g change, 1mm ong, ovary 1 mm long; flowers light yellow;

sonererstiy a ond pee a7) on one, | ‘Ten wide, eet pty ea ar

subquadrate, 0. mam ong, 08 phan rag oon see 1-veined, with the apex thick externally; lip

flat, minutely pubescent, the Ota me ins il pf ow nde cli the dorsum

05mm wide thefet ont ea aia ee co t 0.5 mm long

: eta dvicdl.

STELIS OF ECUADOR 53

Pichincha: new road between Quito and Santo Domingo, alt. 2600 m, 31 Mar. 1984, C. Luer, S. Dal-

strom, T. Héijer, J. Kuijt & A. Hirtz 9809 (Holotype: MO).

Napo: hill south of Lumbaqui, alt. 850 m, 10 Feb. 1986, C. Luer, J. Luer, A. Hirtz, W. Flores & A.

Embree 11761, 11762 (MO).

This species occurs on both sides of the Andes of central Ecuador. Related to

the numerous variations of S. pusilla, it is distinguished by the much larger habit

with very slender ramicauls, and slightly longer, narrow leaves that are surpassed by

slender, secund, strict racemes of numerous, minute flowers. The sepals are ovate,

obtuse, and usually not widely expanded. The petals are cuneate, cucullate and

single-veined. The lip is concave anteriorly with a small, obtuse apiculum, and a

pair of prominent, rounded calli on the bar.

70. soo micacea Luer & Hirtz, sp. n

ty.: From the Latin mica, “‘a tiny bit,” fering to the very small habit.

Planta by Hf, +f] £85 i M duniol

sepalis piabeis late ovatis, 5 ech incrassatis semilunatis, labello crasso subovoideo superne cum callo

orbiculari distinguitur.

Plant very small, epiphytic, pees: Bar roots slender. Ramicauls — — 2-4 mm long,

enclosed by a tubular sheath and another at the base. Leaf erect, coriaceous, elliptical, subacute to

obtuse, 5-12 mm long including a oetiede i. 2 mm long, the blade 3-4.5 mm oid ead below into the

petiole. Inflorescence an erect, distichous, subflexuous, congested, several-flowered raceme with 2-3

flowers open parma sia 20-30 mm — includ me the peduncle ca. 1 cm long, from an annulus

below the apex of the aul; floral bracts proportionately large, oblique, acute, dilated, 1.5 mm long;

pedicels 0.5 mm ie paces ra 75 mm “sits ong; sepals aa or as glabrous, expanded, similar, broad-

ly ovate, obtuse, connate basally, the dorsal — 1.5-2 mm long, 1.75-2 mm wide, 3-veined, the lateral

sepals oblique, 1.25-2 mm long, 1.5-2 mm wide; petals sr poration semilunate, concave below

the thickened margin, concave at the base, . 4 mm long, 0.6 mm wide; lip purple, thick, subovoid,

0.4 mm long, 0.6 mm wide, 0.3 mm deep, concave anteriorly above the y rounded apex, the bar

with a denies continuous with a central, rounded callus on the dorsum, the base broadly truncate,

hinged to the base of the eee column stout, 0.6 mm long, 0.6 mm wide with the apical stigmatic

lobes, the foot obsolescen

Pastaza: north of Puyo, alt. 1000 m, 12 Apr. 1985, C. Luer, J. Luer, A. Hirtz & W. Flores 11209 (Holo-

type: = mae

N w road to Coca from Archidona, alt. 1200 m, 13 Apr. 1985, C. Luer, J. Luer, A. Hirtz

Flores i 226 mb

tiago: ceErNEN del vateegeryee sii ther

18 a 1989, C. Luer, J. Luer, riz, S. Ortega, A. & P. Jesup 13995 ros

south of Yangana, alt. ani m, il May 1981, C. Luer, J. Luer & A. Hirtz 6149 (SEL).

“This s tiny species occurs on the eastern declivities of the Andes of Ecuador. —

characterized by the minute, caespitose plant with elliptical leaves longer than °

Tamicauls, and a minute, closely subflexuous raceme about twice as long as

leaves. The sepals are broadly ovate and and the lip is concave and obtuse —

ly with a rounded callus on the dorsum continuous with the glenion as seen 1n

globulifera.

d Morona. alt. 950 m

71. Stelis minutissima Luer, sp. nov. cane

ty.: From the Latin cae minutissimus, one referring to the flow

Plant eeeit re iis ellipticis is longiore, sepalis ciliatis c ous obtusis,

petali ib Ss ene Ba kat $B Lene

= Ls

-5 mm long,

Plant very small, epiphytic, caespitose; roots slender. Ramicauls erect, slender, 2-5 nea &

enclosed by 2 tubular sheaths. Leaf erect, coriaceous, elliptical, Ce Oe tiolate,

long including a petiole 3-4 1 mm long, the blade 5-6 mm wide, cuneate below into t : petiole.

: capsul ary 1 mm

ramicaul; floral bracts tubular, 1.5 mm long; pedicels 0.5 mm long; ov : sega tat within the

dull white, suffused with rose within; sepals fleshy, glabrous externally, sparse!y

54 ICONES PLEUROTHALLIDINARUM

margins, ovate, subacute to obtuse, similar, 0. 8 mm sind = . mm wide, <a but with faint, incom-

suborbic hallowly concave, broadly — ‘ the

apex, 0 0s mm long, 0.5 mm wide; popaae 4 sabobovat, “05: mm long, 0. 3 mm w wide, 0. 3m

tel

p salt a es Scala te. hinged to th tcol foot: column stout, " 5 nly

saagre pein Ppassthaa ier eran -)

: scrub tation south of Cuenca ova Sang alt. 3000 m, 17 Mar. 1984, C. Luer, S. Dal-

strim, T. Héijer & J. Kuijt 9537 (Holotype: M

This species, found in southern Ratti is perhaps the smallest-flowered spe-

cies known, with sepals eight-tenths of a millimeter long and wide. The flowers are

borne slowly in a three- to four-flowered raceme. It is similar vegetatively to S.

exigua, but the latter is characterized by a longer, more congested raceme. The

sepals are thickly ciliate within the margins; the petals are thin and single-veined;

and the minute lip is concave anteriorly and callous on the dorsum.

72. Stelis se: tan uer & Hirtz, sp. n

Ety.: From the Latin nepotula, “a little Chere” "refering to to the “AR eeaedh with S. —

A ge haec S. morganii Dodson & Garay af!

Plant small, epiphytic, repent, the rhizome stout, branching, 5-15 mm long; roots slender. Rami-

cauls poate slender, 5-10 mm long, enclosed by a loose, tubular sheath and another at the

erect, y coriaceous, elliptical, acute, 15-25 mm long including a a 2-3 mm long,

the blade 5-6.5 mm wide, cuneate below into the petiole. Inflorescence an erect, congested, distichous,

several-flo raceme, with a few flowers orb apes simultaneously, 15-20 mm long including t dun-

cle 3-5 mm long, borne b the apex of the ramicaul; floral bracts

oblique, acute, 1.5 mm long; pedicels 0.75 mm long; ovary 0.75 mm cng: flowers yellow; sepals gla-

brous, expanded, y ovate, obtuse, the dorsal sepal 1.5 mm mm long, 1.75 mm wide, 3-

veined, the lateral sepals 1 mm long, 1.5 mm wide; petals transversely ovate, cellular shal-

lowly concave, 0.5 mm long, 0.6 mm wide, 1-veined, the apex broadly rounded, thickened at the tip and

aon g the > margin; lip sane psu thick, 0.3 mm long, 0.6 mm wide, 0.3 mm deep, the anterior

a small glenion Pease oadly rour ded, transversely thickened across the center, with

ithe cal , the base truncate, hinged to the base of

column; column stout, 0.5 mm long, 0.6 mm wide, the foot obsolescent, the anther and the bilobed

auntie apical.

Napo: forest along the new road to Coca, north co alt. 1200 m, 13 Apr. May 1985, C. Luer, J.

Luer, A. Hirtz & él ye 121 > Ciadslone Na

gmap ye: pb gon rant Rio Upano, ok of Macas, alt. 1100 m, C. Luer, J. Luer, A. & P. Jesup,

bee small species, known only from lowland, eastern Ecuador, is related to the

larger S. morganii, which is widely distributed on the western declivities.

vegetatively, S. nepotula is distinguished by a thick, creeping rhizome that gives

rise to short, ascending ramicauls with their small, thickly coriaceous leaves. A

Congested raceme of tiny flowers is borne behind the leaf. The sepals are glabrous;

the petals are minutely pubescent; and the li

callus on the dorsum. ip is more or less triangular without a

Stelis ortegae Luer & Hirtz, sp. no

= ned for Se Sentiap? Ontegs of Quin, Ccinice co-collector of this species.

SI uo ane is brevipetiolatis, sepalis late ovatis, petalis crassis lunatis et

Plant small,

amueeatat vc caespitose; steigecerasseras tiga uel se geben 0.5-1 cm long, enclosed

including a petiole coriaceous, elliptical, subacute to obtuse, 2.5-4 cm long

rnetigiimnieenae e053 emi nthe blade 08 13cm cm wide, narrowly cuneate below into the petiole.

EE ge ele a a flexuous, distichous, se several- wered raceme with 1-few

- = ig the F from an annulus below the

STELIS OF ECUADOR 55

apex of the ramicaul; floral bracts oblique, rei : ~ men long; eae 1.5 mm long; ovary 2 mm long;

sepals expanded, flat, ni glabrous ex ular within, broadly ovate, obtuse,

connate below the middle, the dorsal ot 4mm long, 4.5 mm wee . sepals oblique,

3 mm long, 4 mm wide, 5-ve ined; petals fleshy, red- ee transversely obovate, shallowly concave

below the broadly rounded, thick, ill-defined — mm long, 1.25 mm wide, 3-veined; lip red-pur-

ple, thick, transversely oblong-ovoid, 0.6 mm | ong, “* 9 mm wide, 0.8 mm deep, the apex thickened,

broadly rounded, shallowly concave anteriorly without a visible glenion, the dorsum with a rounded,

gy callus, the base broadly truncate, hinged to the base of the column; column stout, 0.75 mm

ong, | wide, the anther and the stigma apical, with the stigma bilobed, the foot

Morona-Santiago: ee bg Wad new road between Mendez and Morona, alt. 950 m, 17 Jan.

1989, C. Luer, J. Luer, A S. Ortega, A. & P. Jesup 13958 (Holotype: MO); east of Guismé, alt.

1650 m, 20 Jan. 1989, C. fit H Luer, A yt & P. Jesup 14018 (MO).

This species occurs in the lowlands of the Cordillera del Cutuct. It is distin-

guished by short ramicauls with proportionately large, broad, leaves and a few-

flowered raceme of proportionately large, flat flowers. The sepals are broadly

obtuse, the petals are thick, and the lip is obtuse with a callus on the dorsum.

74. baa t peachy playa Luer & Hirtz, sp. n

m the Greek gmk “a thick eanrel a * refering to to the habit.

ae a Mie xf *

—J

le }4 florib ] is imbricatis. cenalis obtusis a LI . int

z © Li r + =a r

Plant large, epiphytic, oe roots slender. Ramicauls erect, stout, 6 mm thick above, 3-9 cm

long, enclosed by a large, loose tubular sheath above the middle and 2-3 other sheaths at the base.

erect, thickly coriaceous, broadly. ovate, obtuse, 8-16 cm long, the blade 4-7 cm wide, conduplicate in

lower half, rounded at the sessile base. Inflorescence a thick, rigidly erect, very congested, many-flow-

ered raceme, secund with lowes overlapping, many flowers open simultaneously, 18-24 cm ‘dy in-

cluding the peduncle 10-15 cm long, from the base of the leaf, with a a spathe ca. lem long; floral bracts

broad, tubular, 3 mm long; pedicels 3 mm long; ovary 3 mm | fleshy

glabrous externally, mrs potas gona ae ovate. ate, obtuse, the dorsal ‘sepal 5 mm a 4.5

ined, connate 1

y, the | sepals 5 mm long, 3.5 mm wide, conn

petals Sater a Sas cam i® Pe mm ng 1.8 mm wide, 3-veined, sllowy concave

below the rounded, thickened margin; bes depen hye erga 08 mm long, 1.3 mm wide, sagt sie

hes eergh shallowly concave anteriorly, with a small glenion, the dorsum narrow t inged

et bilobed stigma apical.

Zamora-Chinchipe: south of Yangana above Valladolid, alt. 1900 m, 23 Mar. 1985, C. Luer, J. Luer &

A. Hirtz 10848 (Holotype: MO); same area, alt. 2000 m, 18 Mar. 1984, C. Luer, S. Dalstrém rom & T. —

9616 (MO); Quebrada Honda, alt. 1500 m, 17 Jan. 1982, D. D’Alessandro 114 (SEL); new road

Loja and Zamora, alt. 2250 m, C. Luer, A. Hirtz et al. 13608 (MO)

Napo: welaesees 1800 m, May 1982, A. Hirtz 307 (SEL).

s large species is apparently endemic in southernmost Ecuador. It is distin-

guished by short, thick ramicauls and large, thickly coriaceous leaves that are

conduplicate at the sessile base. From within the fold, the thick, stout, strict pedun-

cle rises to bear a very congested raceme of overlapping, dark purple flowers. The

flowers present in two rows, both facing the same direction. The floral parts are

obtuse and not remarkably distinctive.

75. Stelis phil-jesupii Luer, sp. n

Ety.: Named for H. Phillips Jesup of Bristol, CT, co-collector of this species.

Planta parva caespitosa, racemo laxo paucifloro, sepalo dorsali orbiculari, sepalis lateralibus brevi-

bus latioribus quam longis, petalis carnosis obtusis, et labello subquadrato apice p rotundo.

Plant small, epiphytic, caespitose; roots slender. Ramicauls erect, slender, 5-9 mm long, enclosed

bya heatt Leaf erect, coriaceous, elliptical, subacute to —

ssa 29-30 FRE Ee ap tiole 5-10 mm long, the blade 7-10 mm wide, cuneate below ir

the petiole. "i aden ay an erect, successive, distantly f fow-Shovenvet i raceme one"

the peduncle 20 mm long, fi 1

1a hilaclk

1-15 mm long; pedicels 1.5 mm long; ovary 1 mm long; fl flowers purp F

56 ICONES PLEUROTHALLIDINARUM

ly, d at th , 2.5 mm long, 2:5

mm wide, 5-veined, ¢ connate basally, he ly, the lateral pie ranvee ovate, oblique , obtus tuse, 1 mm mm long, 2

mm wide, 4-veined; petals thickly triangular, smooth, 0.75 mm long, 1.2 mm wide, thickened above the

k pex; lip thi mm long, 1 mm wide, 0.5 mm deep,

flat t anteriorly ith Il gleni ith th ded, the sida scsi a rounded, microscopically

the b hinged t bsol ot; column stout, 0.8 mm long,

Sion! ides iin siiewes (de Sietedatinese wicel

Zamora-Chinchipe: Cordillera del Condor, east of Los Encuentros, alt. 1500 m, 21 Jan. 1989, C. Luer,

J. Luer, A. Jesup & P. eee 14041 (Holotype: MO); Cordillera del Condor, alt. ca. 1500 m, cultivated at

Ecuagenera, Gualaceo, 7 Mar. 2001, C. Luer 19839 (MO).

This species is known from the Cordillera del Condor in southeastern Ecuador.

It is characterized by a small habit, with a loose, successively flowered raceme that

eventually surpasses the elliptical leaves. The sepals are convex with revolute

margin sepal is orbicular and more deeply connate to the half-as-large

laterals than they are to themselves. The petals are semilunate, and the lip shortly

oblong with a callus on the dorsum.

76. ~~ aLarsagy Luer & Hirtz, sp. n

m the see pilosus, “with der vaste ee to aed doped neon peice

Planta racemo = aes

Lom bk p

petalis tr is oblongis concavis, et labello subqua-

parapet seein cea:

(—

Plant small, epiphytic, a gp ts slender. Ramicauls erect, slender, 8-12 mm long, enclosed

by 2-3 tubular sheaths. Leaf erec | tae elliptical-obovate, subacute to obtuse, petiolate, 25-45

mm long oe a petiole 8-10 mm long, the blade 10-15 mm wide, cuneate below into the petiole.

Inflorescence a distichous, sublax, successively several-flowered raceme 5-7 cm long includ-

“ :

es he eneiriae sani eigeearie

with a long, narrow Ow glenion persist cc and densely zoho: the b te, hinged

obsole besce

slim apical apical = coke foo column stout, 1 mm long, 0. 3 mm wide, se the anther and the bilobed

Pastaza: forest north of Mera, alt. 1350

Caceres MO). m, 12 Apr. 1985, C. Luer, J. Luer, A. Hirtz & W. Flores 11202

> sepals are ovate, obtuse, more or less

nu slater & Hirtz, s ep; SOv-

oer polycarpicus, “fruiting many times,” referring to the prolific production of

iS Tracema =

1 . 15

sewn capsulas formantibus, sepalis g ok ants, Crpnic ez petiona is longiore, —

SOSAIES ULL

Plant medi large,

ee epic, caespitose; roots fine. Ramicauls erect, slender, aig cm long,

- Leaf erec

Petiolate, acute, the blade cm coriaceous,

1.5-2 cm long. blade 10-15 cm long, 2.5-3.5 cm wide, cuneate below into th oa

- a a Strict, ies son ted,

flowers bearing capsules simul ee secund, many-flowered raceme with most

9gnnniac Kal, pr simultaneously, 12 Zz 23 lon a

min long: ovary 25-3 mm long ipables toi iq ee eS oan bag: pedeel 15

eae ott JeMUW-E rose,

STELIS OF ECUADOR 57

glabrous, ovate, obtuse, the dorsal sepal 8 mm long, 7.5 mm wide, 5- to 7-veined, the lateral sepals

connate 4 mm, 7 mm long, 3.25 mm wide, 4- to 6-veined; petals yellow, transversely obovate, concave,

the apex thick and aed obtuse, 0.75 mm long, 1.5 mm wide, 3-veined; lip thick, ovoid, shallow-

. concave anteriorly with the apex i 0.5 mm long, 0.9 mm wide, 0.8 mm deep, the bar thick with

an indistinct glenion, ri dorsum with a low, rounded callus, truncate, —— to the base of the column;

column stout, 1 mm long, 1 mm wide, ri anther and the bilobed stigma apical

Carchi: forest east of Maldonado, alt. 1550 m, 17 Mar. 1991, C. Luer, J. Luer, J. del Hierro, A. & X.

Hirtz 15125 (Holotype: MO).

This species, known from northwesternmost Ecuador, is characterized by an

elliptical, acute, long-petiolate leaf far surpassed by a strict, secund, crowded

raceme that produces many capsules simultaneously. The sepals are ovate with the

laterals semiconnate. The central apparatus is proportionately small. The petals are

thick and concave. The lip is ovoid and concave anteriorly with a round apex.

78. a9 porphyrea Luer & Hirtz, s

From the Greek senilickis Raju % fering to the eee and flowers.

Plant: a

x rr Ls r r bail

Pt,

facut pannite fic ahimcie <tnctihh : h ran tali i tis, et labello

m r

hind Slag Anan ots

Plant small, epiphytic, caespitose; roots slender. Ramicauls erect, slender, 1-2.5 cm fe enclosed

by 2-3 tubular sheaths. Leaf purple a erect, a. beg cal, subacute to obtuse, 2.5-4 cm long, 0.7-

1.2 cm wide, eate below i 1 cm long. Inflorescence an erect, distichous, sublax,

sei a few- flowered raceme sto 6 cm long including the peduncle ca. 3 cm long,

belo apex of the ramicaul; , floral bracts acute, 1 5 mm long; | pedicels 1.25 mm long; ovary 1.5 mm

lon ng; fl 1 , expanded ly. sparsely sho! epeagar ieee =

the ‘sepals ovate, obtuse, 3: veined, ‘connate basally, th the dorsal sepal 2.5 mm long, 3 mm w

sepals 2.25 mm long, 2.5 mm besos petals transversely ovate, thick from above the base, teoadly round-

ed at the apex, pea , 0.9 mm long, 1.25 mm wide; lip thick, irene, 0.6 mm =~ 0.8 mm wide,

0.6 mm deep, shallowly concave pat with the apex rounded, with a sharp apiculum in the center,

the anterior surface of the bar — a long, narrow wen the d dorsum with ; postcnccen callus at the

base, the base truncate, hinged to m long, 0.6 mm wide,

Pichincha: forest between San Miguel de los Bancos and Mindo, alt. 2300 m, 1 Apr. 1984, C. Luer, S.

Dalstrém, T. Héije ijer & A. Hirtz 9879 (Holotype: MO).

This small species of northwestern Ecuador is closely related to the much larger

S. argentata. It is characterized by the caespitose habit with all parts of the plant

and inflorescence purple. The raceme is simultaneously several flowered, and

exceeds the leaf in length. The sepals are shortly pubescent, broadly ovate, obtuse,

and more or less equal; the petals are thick and convex; and the lip is thickly sub-

quadrate with a sharp apiculum in the center of the round apical margin.

79. Stelis procera ra sp. n

Ety.: From the Latin procerus, aan referring to the peduncle and raceme ieeiss

ase ean Esti racemo brevi laxo cum pera elongato folio eal elliptic

multilong: ideo subplano distinguitur.

Plant m epiphytic, lender. _ erect, slender, 3-4 os _

enclosed by a loose, ‘abulec - sheath, and 2 others below. Leaf erect, coriaceous, a iptic

subacute to obtuse, 6-7 cm long including a petiole 1-1. 5m long, ihe blade 1.2-1.5 cm

below into the petiole. Inflorescence an erec nsec, lax, several-flow

€rs open os , 5-12 cm long, total ‘a

Several, distant brac to m an ann eaten she tee aord of namic floral bracts ;

om long; pedicels 1-1.5 mm ag oka 2 mm long; sepals sae cream-co’

shortly ciliated margins, broadly ovate, obtuse, connate basally, 2. mm long, 2. oa

petals yellow with purple mete. fobesisisar oblong-suborbicular, vee pi“ 1 * veined,

concave below the broadly rounded, slightly thickened, apical margin; lip yellow with F P zi

1 flow.

58 ICONES PLEUROTHALLIDINARUM

discoid, shallowly subquadrate-oblong, 0.75 mm long, 1.4 mm wi ide, 1.4 mm deep, the apex broadly

bese oad

tI 5 mm long, 1 4 mm wide, the foot obsolescent, the anther and the stigma apical and

Loja: near Yangana, collected by D. D’ Alessandro 80-0107, flowered in cultivation at SEL, 1981, C.

Se SEL).

This es, apparently endemic in southern Ecuador, is allied to S. major, but

differs with narrower leaves; a proportionately ope Sif heen flowered

raceme borne by a thrice longer peduncle; and flowers half the size. The sepals are

ovate, obtuse, and minutely ciliate; the petals are ssoebicalar oe concave; and the

lip is discoid, or flat vertically, that is, much deeper than long.

80. Stelis protracta Luer & Hirtz, sp. no

Ety.: From the - abenese et a referring to the sshecen come rachis.

a pai mi rey

multilongiore floribus illic S tor, Peed 52) ee ee . es ee ee RS

| ae

Piant medi

1 meres:

, the rhizome stout, erect; roots fine, nu-

merous, Ramicauls erect, seek pope cm long, enclosed b by 2. 3 tubular sheaths. Leaf erect, coriaceous,

suonty elliptical, acute to subacute, 7-9 cm long including a pee ca. 2cm long, the blade 1-1.5 cm

wide, narrowly: cuneate below into the petiole. Inflorescence an erect, strict, distichous, successively

aay fl 1] dis tantly flowered below,

subcongested toward the tip, 15-22 cm long ‘including the peduncle 5-7 cm iong, wi a few bracts, from

125 1 ar, 1-1.5 mm mm long; pedicels 0.75 mm long;

sean purple, ciliate, shortly pubescent , connate basally, broadly

ovate, obtuse, the the dorsal non ‘1.75 mm long, 1.75 mm wide, 3-veined, the lateral sepals 1.4 mm long,

a brainer ors = 8 a : purple, transversely obovate, shallowly concave, 0.6 mm long, 0.8

by cellular , thic nsely covered

rl points, transversely : subquadrate, 0.4 mm long, 0.6 mm wide, 0. 5 mm deep, concave inside

below the middl » he margin thin, minutely apiculate, vex with a very small, round glenion

phi e este sa et roadly magen oo pubescent, anges to the base of the column;

column stout, 0.5 mm long, 1 mm wid Sea

stigma hate . and the stigma apical, the

Fl Orn ii eee ee

W. Flores 10684 (Holotype: MO). to Loja, alt. 830 m, 20 Mar. 1985, C. Luer, J. Luer, A. Hirtz &

; This tiny-flowered species is closely related to the common S. argentata. The

ri Slender raceme is strict and distantly flowered except toward the tip. Superfi-

eg y similar, the flowers are smaller than the smallest known of S. argentata. The

P is most distinctive, being covered by microscopic, spiculate prickles. The bar is

similar, but instead of a a narrow gle d

below the middle of the anterior glenion, the glenion is minute and roun

81. Stelis pubipetala Luer & Hirtz, s

Pp. nov.

area ae the Latin pubipetalus, “wi th pubescent petals,” ee

centibus, et Hace S. lanceae Lindl. similis, sed floribus \jori Soe, Mine

or o r

WOM MEICiL,

Pit i ten ge: nes en stout, 1-2.5 cm long between ramicauls;

the middle and 2-3 ieee the aes I bane 10-12 cm long, enclosed by a tubular sheath near

lon: coriaceous, elliptical, subacute, late, 10-12 cm

testes, sonnets cm long, the blade 2.5-3 em wide, c cca below in acpi. tater

simultaneously. § 8-13 cm long including the pe anks with many Huan open

oom hoot emda long, from an an annulus below the apex of

' iio

rial ener ene gtr cer

45, p

iene mi wie, Sic aoe nea the mide: petals purple, broadly

vey eee

STELIS OF ECUADOR 59

triangular, subacute, with a — flattened margin, densely short-pubescent, shallowly concave below

the middle, 1.5 mm long, 1.5 mm wide; lip purple, thick, triangular, obtuse, 0.8 mm long, 1.3 mm wide,

1.3 mm deep, shallowly concave mplowie 5 with a narrow glenion, the dorsum with 3 rounded calli, the

base broadly truncate, eat ed to the base of the column; column stout, 1 mm long, 1 mm wide, the

anther and the stigma a)

ee between Giidiaaies and San Juan Bosco, alt. 1250 m, 27 Mar. 1985, C. Luer, J.

riz & W. Flores 10980 (Holotype: MO).

With the large, repent habit, long ramicauls, and a petiolate leaf, this species is

inseparable vegetatively from S. lancea. However, the flowers are larger; the petals

are triangular with a thick, shortly pubescent margin; and the lip is subtriangular

and shallowly concave, with three distinct calli on the dorsum.

82. Stelis purpurella Luer & Hirtz, sp. no

Ety.: From the Latin purpurellus, “‘a little se one,”’ referring to the habit.

pecies haec S. —— Lindl. similis, sed bracteis floralibus minoribus et sepalis trinervis

pubescentibusque distinguitu:

Plant small, epiphytic, —— caespitose; roots slender. Ramicauls erect, slender, 1-2 cm long,

enclosed by a close, tubular sheath and 2-3 sheaths at the base. Leaf erect, coriaceous, narrowly ellipti-

cal, acute to subacute, 3-6 cm oe ne an indistinct i i -2 bas wh the vena 0.5-0.6 cm

wide, gradually narrowed below into the petiole. Infloresce ongested, distichous, many-

flowered raceme with se iain on open eerste 10-12 cr cm n long including the pedonct e 3-4

cm long, with a spathe 3 mm

ute, 2-3 mm long; ay 1 "S mm long; ovary 1 mm long; .; sepals expanded, purple, shortly pubes-

cent, transversely ovate, obtuse, deeply connate, the dorsal sepal 3 mm long, 4 mm wide, 3-veined, the

ong, 3.5 mm wi e purp

broadly rounded with a broad, flat margin, shallowly concave above the base, 0.75 mm long, 1.5 mm

wide; lip purple, thickly triangular, 0.75 mm lon ng, 1 mm wi ide, 0. 5 mm deep, concave anteriorly, =

the apex obtuse with a thick margin, the dorsur

waa broadly truncate, a to the base of the column; column stout, 1 mm ‘long, 1 mm wide, si

Napo: Guacamayo range between Baeza and Tena, alt. 1950 m, 13 Apr. 1985, C. Luer, J. Luer, A. Hirtz

& W. Flores 11253 (Holotype: MO).

This species is characterized by the small habit with short ramicauls bearing

considerably longer, very narrow leaves that are surpassed by a congested raceme of

small, purple flowers. The flowers, as well as the plant, are similar to those of S.

muscifera, but the floral bracts are distinctly smaller. The transversely ovate sepals

are deeply connate, minutely pubescent within, and three-vein

83. Stelis sanctae-rosae Luer & Hirtz, sp. n =

a for the community of Sancta Rosa in pera Ecuador, near where the speci

was c

giore, sepalis : 7. ed g is, et labello crasso

subquadrato tute

Plant small, epiphytic s slender. Ramicauls erect, slender, ty ines lon -

enclosed by 2 tubular ‘sheaths and another at the b a Leaf cnet, coriaceous, narrow —

subacute to acute, oe m long, the blade 0 cap cm w : :

preinndnl o the petiole. Inflorescence a congested, sec ont sola, an sg a - apr ie

low

cm long including the peduncle 3-4 cm long, with most flowers open simultaneously, fro

below the apex of the ramicaul; floral bracts oblique, acute, 1.5 mm long; ssnte .5 mm long, ovary

basally, glabrous externally, coarse

sepals expanded, con any a ee a as 1

mm long, 1.6 mm wide; petals purple, transversely oblong, subtruncate, concave, tg : sa s 07

thickened, 0.6 mm long, 1.1 mm wide, 3-veined; lip purple, mar, he arb _— papa Ley:

mm deep, shallow! concave ayn iperhemone acinar magn, | gl ’

base pubescent, truncate, h i th lnmn stout. 0.75 mm long, 1mm

60 ICONES PLEUROTHALLIDINARUM

oO Santa Rosa to Loja, alt. 850 m, 20 Mar. 1985, C. Luer,

J. Luer, A. Hirtz & W. Flores 10682 pa MO).

This species is known from relatively lowland in southwestern Ecuador. It is

characterized by the small, caespitose habit; elliptical leaves longer than the rami-

cauls; and a slender, congested, secund raceme of very small flowers. The sepals

are broadly ovate and coarsely pubescent; the petals are transverse, thin and con-

cave; and the lip is shallowly subquadrate

84. Stelis scalena Luer & Hirtz, sp. no

wt risciappaiiqaalees 07% a " refering tthe spl,

lat, one a

p i , Sepalis pap

Plant medium in size, epiphytic, caespit lender. Ramicauls erect, slender, 3-4.5 cm long,

1 Akh | ee | L s £ hel _7 ' 3.3) Ay eh. the base . Leaf erect, coria-

ceous, elliptical, sub te t te, 6-8 cm long i iol 1-1.5 cm long, the blade 0.8-1.4. cm

wide, cuneate below into the petiole. flan a et cng, c many-flow racem

with most flowers open simultaneously, 1 12-20 cm 1 4-5 cm _—_ from an

mm long; pedicels 2 mm long; ovary

1, 15m mm m long; sepals expanded, light yellow, patos extemal, minutely Papillose a fone the

-9 mm long, 1.8 mm

Be 3. veined, the lateral sepals shoal 2 mm eta z mm wide, 3-ve eined; petals thin, light

yellow, transversely cuneate, mm long, 0.75 mm wide, 1-veined; lip light yellow, —-

pegs mm long, 0.6 mm wide, 0. 8 mm om concave more the middle, Tounded at t the ¢ apex, then

wunrene: fel d to the k ly

ener ne oe gma pe

t, : mm long, 0.6 mm wide, the foot cbeclescent,

Pichincha: road to \ , betw Chirit d§$ Domi alt. 1300 m, Jan. 1989, A. Hirtz

3963 (Holoype: Mo).c. Luer illustr. 19493, a PE Ee

beneath. The petals are thin, truncate and one-veined. The lip is oblong and con-

cave above the middle with a large, pyramidal callus below the middle.

85. Stelis serrulifera & Hirtz, sp. nov.

Ety.: From the Latin poten bearng small small serrations,” Paige to eg — of the labellum.

Species haec S§. argentatae Lindl. similis. sed senal

ibus recurvis. labelli SE ae 2 oe acini

_ Plant aber in size, ceirinic, caespitose; Toots slender Ramicauls erect, slender, jin cm —

Clow and 2-3 others at the Leaf erect,

<= ofa ole. I including an indistinct petiole ca. 1 cm long, the blade 0.5 cm

petiol erebengihs ; erect, subdense, distichous, several-flowered

cena emt, sd Wn vaataat fasca long including the peduncle 3-5 cm long,

ae 1 sa mm long; ovary 2 mm long;

ovate, obtuse, minute pubescent-ciliate ate along revolute

8, +. ral

wide, the apex ly wanna ly concave be Ww thickening, 0.6 mm long, 1.2

subquadrate, 0.75 mm long, 1 te wide, cae any sebverracosc, ee eeeety

minutely, sharply, and irr » U.5 mm deep, concave inside the apical n ‘

hin. Sr Poe ae 2 ts a]

ae ween gs Stout, 1 mm long, | mm wide, the foot obso-

STELIS OF ECUADOR 61

Esmeraldas: wet forest west of im, a0 Foy m, 18 Jan. 1987, C. Luer, J. Luer, C. Dodson, A. Hirtz & D.

Benzing 12418 (H tion data, C. Luer et al. 12357, 12397 (MO).

This species of ‘oatlane northwestern Ecuador is related to the widely distribut-

ed S. argentata. Stelis serrulifera is characterized by a small habit with elliptical

leaves more or less as long as the ramicauls; several-flowered racemes with flowers

borne in two ranks that exceed the leaf. The sepals are more or less round with

revolute, pubescent margins that cannot be straightened. Toward the apices, the

hairs become long. The petals are semilunate with thick, flat margins. The lip is

small with a rounded apex with a thin edge that is irregularly serrulate with at least

three longer, pointed processes with several smaller processes between and to either

side. The margin is reminiscent of the triapiculate margin seen in S. vulacnica.

86. bogies a Luer & Hirtz, sp, —

similarity to Stelis tridactylon Lue:

sesh haec S. poner Luer similis, sed habitu minore, foliis petiolatis, floribus minoribus,

e, epiphytic der. Ramicauls erect, slender, 3-4 cm long,

with a loose, tubular haa from below the middle, and 2 other sheaths at the base. Leaf erect, coria-

ceous, eggs subacute to obtuse, 4-6 cm long noes a er less than 1 cm long, the blade 1-1.5

= into the base. Inflorescence an distichous, densely and nearly simultane-

ous: iy many-flowered raceme, 12-22 cm long including the nach 3-7 cm long, subtended by a spathe

5-10 mm long, with an annulus at the base of the leaf; floral bracts broad, short, 1.5 mm long; pedicels 1

mm long; ovary 2 mm long; sepals yellow-green, similar, coarsely minutely pubescent, connate basally,

llo

apex broadly rounded, ‘thickened along the margin, shallowly ec concave, 0.75 mm long, 1 mm wide, 3-

i lip yellow-green, hemispherical wi ith the © apex broadly rounded, 0.6 mm long, 0.8 mm

cep, lenion dorsally,

truncate,

mm wide, the foot obsolescent, the anther and

to sa base of the coleitine column stout, 0.75 mm long, 1

the bilobed stigma apical.

Morona-Santiago: Cordillera del Cutuct, 7 eh new road east of Mendez, alt. 700 . 5 re

C. Luer, J. Luer & A. Hirtz 12640 (Holotyp - same collection data, C. Luer, J. Luer

12639 Guo).

while the lowermost flowers of

timeter in length. The

coarsely short-pubescent.

ina densely flowered raceme, are all the same size,

S. tridactylon are much larger, the sepals reaching one cen!

Sepals are glabrous while those of S. tridactyloides are coarse

87. Stelis undulata Luer & Hirtz, s

Ety.: From the Greek undulatus, ‘“peidailabe: “a.”

Planr a:

as pee see UR oral bracts.

longiore bracteis floralibus magnis

illosis, labello crasso

pens iis undulatis, — ciliatis late ovatis, , petals semilunatis ; marginibus long-pap

eaecnitanca

der. a erect,

Plant medium in size, epiphytic, jamal caespitose; roots eget - t, coriaceous,

slender, Stem long, enclosed by a loose, tubular s: ne ada 1 p15-20m wide

elliptical, subacute to obtuse, 5-7 cm long including a petiole 1.5-2 cm pei ‘ab

cuneate below into the petiole. Inflorescence an erect, secu — pee ian: from an oe

below the apex of the ramicaul, with an inconspicuous spathe; floral

ed, undulate, 7-10 mm long; pedicels 1-1 5 mm long; ovary 2.5 mm gr ag ge

oor — within the — coir rt obtuse, connate arse mm long, thickened above

S; petals purple ersely oblong-semilunate, d long, 1.25 mm

the middle with the margin ei ana minutely densely papillose, 1 mm ‘ong,

62 ICONES PLEUROTHALLIDINARUM

wide, men ay i thick, subdiscoid, 0. 6mm long, | 1mm wide, 1mm deep, Gensely microscopi-

cally pu

a pair of mar-

» the dorsum ‘with a low, obscure, longi-

nal alte se oly ete i ged to the base of the t, 0.75 mm long,

Carchi: between Maldonado and Chical, alt. 1300 m, May 1992, A. Hirtz 5772 (Holotype: MO), C. Luer

illustr. 19474.

vio se apparently endemic in relatively lowland forest of northwestern-

s characterized by elliptical, petiolate leaves that are surpassed by a

pai a raceme of small, purple flowers subtended by large, dilated,

undulating floral bracts that protrude from one side of the raceme. The sepals are

obtuse and ciliate within the margins. The broadly rounded margins of the petals

are minutely and densely long-papillose. The rounded lip is densely short-pubes-

cent with longer hairs surrounding a central depression.

co r

nS 1 +1. 4.3

88. Stelis uxoria Luer & Hirtz, sp. no

Ety.: From the Latin aie “the wife’s,” referring to Jane Luer who collected the first eee

I ]

t Teil. ‘ dann £ 14} hii . : . . lit + res — sae

o 7 r r 7 -

lahell e 3 4 bd a a: 4

Plant large, epiphytic, repent, the rhizomes stout, 5-10 mm thick, 2-5 cm jon between ramicauls;

roots coarse. Rami stout, ii gies cm long, with a loose, tubular sheath near the middle, and 2

other sheaths at the base. Leaf erec oriaceous, elliptical, obtuse, 15-24 cm hs. including a petiole

ergo ge sep ay into mce an erect, congest-

distichous, progressively many-flowered raceme, with several flowers open paca 35-55

cm tong including the aero 5-20 cm long, wiht spathe 1-1.5 cm long, from an annulus below the

gpl ts oblique » acute, 10 mm — below to 5 mm long i pedicels 3-4

> Ovary 2.5 xternally, dull ish white or silv

pro hags pi es -glandular within, ae ate, obtuse, c te 2- : _ ye flat

flower, the dorsal sepal 5-10 ~ long, 5-5-9 5.5-9 mm wide, a dlacd the lateral sepals 5- “ap mm ia ~

ick nca

mm wide, 5-veined; petals ye

the thickened margin, 1 mm a LS am wide, ey to 5-veined; lip yellow, thick, sacs: ve

drate, with the apex broadly mm long, 1.5 mm wide, 1 mm deep, ly concave amerior-

ly witha small enon, the dorsum wih tow callus, flanked by 2 I

ly truncate, hinged

sient: 4 werclong. 1:5 men wide, ehacbenhaeeral

bilobed stigma apical, the foot obsolescent. _

Carchi: terrestrial on road-shoulder, between Tul an

i bas A Meh Pcs er Ic and El Carmelo, alt. 3200 m, 6 Apr. 1985, C. Luer,

15093 - area, alt. 1600 m, 16 Mar. 1991, C. Luer, A. Hirtz et al.

wen of El Play6n, alt. 31 3100 m, 20 Mar. 1991, C. Luer, J. Luer & A. Hirtz

15194, 15195 15200 (MO

This large Lage locally abundant at high altitudes in northern Ecuador, is

characterized by a repent rhizome with tall, stout ramicauls with large, ellipti-

cal leaves. Several . many flowers are produced simultaneously as anthesis as-

cends in a lengthening raceme. The flowers, variable in color, are large with

dimensions of some specimens double those of others. The sepals are broadly

Ovate, obtuse and five-, seven-, or rarely nine-veined. The tals are transverse with

a thick, rounded margin, and the lip is subquadrate with a broadly rounded apex,

and concave anteriorly.

STELIS OF ECUADOR 63

89. Stelis vesca Luer & Hirtz, sp. n

Ety.: From the Latin vescus, “little, weak.” "referring to the habit.

Species haec S. g isR , sed sepalis glabris

integro dorso calloso ) differt.

Plant very small, epiphytic, caespitose; roots filiform. Ramicauls erect, slender, 2-6 mm long,

enclosed by 2 tubular sheaths. Leaf erect, coriaceous, — subacute, 8-15 mm long including a

petiole 3-4 mm long, the blade 4-6 mm wide, cuneate below into the petiole. Inflorescence an eect lx

simultaneously several-flowered raceme, 4.5-6 cm long including the peduncle 2.5-3 cm long, from an

annulus below the -_ of the ramicaul, floral bracts tubular, ring! 1 mm long; pedicels 1- “1. 5 mm

long; ovary 0.75 m ng; fl glabro

broadly ovate, ie connate basally, 2 mm long, 4: 25 mm wide, pi ed, the lateral sepals oblique,

1.5 mm long, 1.8 mm wide, 3-veined; petals transversely oblong, shallowly concave below the broadly

nded apex with a thickened margin, 05 mm long, 0.7 mm wide, faintly 3-veined; lip thickly triangu-

ong, 0.6 mm wide, 0.6 mm dee shal-

low depression, the dorsum with a central, round, cellular callus, the base broadly truncate, hinged to the

base of the column; —— stout, 0.6 mm long, 0.6 mm wide, the foot obsolescent, the anther and the

bilobed stigma apic

Morona-Santiago: along Rio sr below Limé6n, 17 Feb. 1986, C. Luer, J. Luer, A. Hirtz, W.

Flores & A. Embree 11854 (Holotype: Mi

This little species of lowland, western Ecuador is superficially similar to S.

guianensis. It is distinguished by a small, caespitose habit with elliptical leaves

surpassed by a loose raceme of small, purple flowers. The sepals are glabrous and

suborbicular; the petals are three-veined; and the lip is shallowly concave anteriorly

with the margin entire.

90. Stelis villifera Luer, sp. n ee

Ety.: From the Latin villifer, heating villous hair,” referring to the pubescence of the sepals.

Planta mediocris caespitosa, racemo secundo multifloro folio elliptico superanti, sepalis. wea

obtusis villosis,

minute pubescenti distinguitur.

a | r

Plant medium in size, epiphytic, aE roots slender. Ramicauls oo°*e “ageing fae

enclosed by a tubular sheath on — ird and another 1-2 sheaths at the base le 2-2.5 cm

ceous, elliptical-ovate, subacute to obtuse, pean 6-8 cm long including a Liga aa snares pies

long, the blade 1-1.5 cm wide, ee below into the petiole. bag I avy cm long, subtend-

simultaneously many-flowered raceme, 7-13 cm long including ash

ed by a slender spathe ca. 8 mm long, foo an annulus below the apex ch page ight green,

infundibular, oblique, acute, 2 mm long; pedicels 1.5 mm anes fe sb ots. connate basally, the

0.75 mm deep, concave anterior

pubescent

wide; lip brian green, thick, ea. 1 mm long, | mm wide,

: F minutely

with a broad glenion, the aici x bro roadly sh as the bar ar thick with 0 broadly sper con aon a

mn stout.

ate, hinged

the anther and bilobed som apical, the foot obsolescent.

pe rehi: forest between Tulcén and Maldonado, alt. 2050 m, 2-4 Apr. |

Gijer 9922 (Holotype: MO).

Ecuador. It is ch haracterized

This species is apparently endemic in northernmost tly longer than the

by the caespitose habit with slender ramicauls that are sree about twice as

elliptical-ovate, petiolate leaves, which are s surpassed b flowers that are long-

Ong. The raceme is slender and congested with minute sce lip is oblong and

villous within. The petals are transversely oblong and thin. J

obtuse, longer than wide, with a pubescent callus at the base.

984, C. Luer, S. Dalstrim & T.

64 ICONES PLEUROTHALLIDINARUM

91. Stelis yanganensis Luer & Hirtz, sp. nov.

Ety.: Named for the —— of Yang: Ecuad I his species was found.

Species haec 5S. C. Schweinf. . similis, 4 lis latioribus, P lis ] g ; larit

. fd uff . Pe Ye Re oe | id rd TI, Ait - oy

Pla It d epiphytic, d amicauls erect, Fvirerk 1 7

4.5 cm long bular sheaths, and another at the base. a erect, coriac narrow

pon ce acute to subacute, sonra 2-4.5 cm long AP the te 0.5-1.5 cm nce on ‘blade 0. 4

0.8 cm the petio subcongested, subflexuous, succes-

sively many flowered sis 310 cm long including ¢ the peduncle 4 5-2.5 cm long, several flowers

the apex 0 the ramicaul Horal bracts ee - oblique 1-2

ar pubescent

mm long; pedicels 1.25 mm long; ovary 1 ]

within, expanded, broadly ovate, obtuse, su’ subequal, co nnate basally. Ges tect eg 25 vs tong.

wide, 3-veined, the lateral sepals oblique, 2 mm long, phages 3-veined; petals purple, triangular,

0.8 mm a 0. 8 mm wide, shallowly concave below the hee fl obtuse and thickened apical margin,

transverse angular, 0.5 mm long, 0. be mm wide,

0.66 mm deep, the apex obtuse, shallowly concave anteriorly i a shallow glenion, the dorsum with a

small, rounded aquired the base broadly truncate, hinged to the base of the column; ; column seit 0.75

mm long, 0.8 mm

Siig Adi of Yangana above Valladolid, alt. 2450 m, 23 Mar. 1985, C. Luer, J. Luer, A.

Hirtz & W. Flores 10932 co MO).

This little species is similar to many other small species, but it is distinguished

by the elliptical leaves about as long as the ramicauls. The many-flowered, crowd-

ed raceme exceeds the leaf with ramicaul about twice in length. The sepals are

broadly ovate and obtuse, about two millimeters long; the petals are triangular, as

long as wide, with the apex thick and narrowly obtuse; and the lip is subtriangular

with a shallow glenion and a small, round callus on the dorsum.

92. Stelis zarumae Lue: & Hirtz, sp. nov.

Ety.: Named for the prs of — near sheng the ee was desineccan

Species haec S. hallii | Lindl. similis f, “= * sepa-

lis intus i 7 4 * leak . kd i’

t TI. il re . oe

Plant small . ee in size, e c, densel 3

phen FE ca piphyti *» densely ‘caespitose; roots slender. Ramicauls erec

: ea nother 1-2 sheaths at the

Ptical, acute to subacute, petiolate, 5-7.5 cm long including the

petiole 1- 1.5 cm lon nthe blade Sem wide, Dp cahioehaabey assem rescence an erect,

subcongested, distichous. si me, 6-8

cm long including the peduncle

ca. 2 cm long, from an annulus below the apex x of th & gcranedy gaara

pedicels 1.5 mm long; ovary Imm mm long: 5 Sepals Acne with a dense, white Pubescence, broadly

Ovate to suborbicular, convex, obtuse, subequal, connate basally pel 5 casa & es

wide, 3-veined, the lateral ateral sepals 23 mm 2.3 M

1 mm long, 1.5 mm wide, shallo Ong, 2.5 mm wide, 3-veined; ed; petals purple, teeny gee

transversely, concave at the base; se purple, ‘subquadrate, 1 hi mm sei 1 mm wide, 0.3 mm | deep, the

tuse.

apex concave, eee

neni

of the ‘Shiai: cht ia

&*' with Pr Aveease

a distinct, cruci iform callus , the base truncate, hinged to the base

stout, 1 mm re 1 mm wide, the foot obsolescent, the anther and the bilobed

El Oro: forest above Zaruma, alt. 1800 m. ‘

Embree 12028 (Holotype: MO). m, 23 Feb. 1986, C. Luer, J. Luer, A. Hirtz, W. Flores & A.

Although not distinctive vegetatively with small-flowered racemes more or less

equal the elliptical leaves, this Species is characterized by suborbicular, convex, red-

purple sepals covered by a long, dense, white pubescence. The thick petals are

proportionately large. The glenion of the lip is prominent with elevated margins. A

t, cruciform callus is present on the dorsum of the lip

ICONES PLEUROTHALLIDINARUM 65

a ®

Fig. 42. Stelis benzingii

ICONES PLEUROTHALLIDINARUM 67

é =< ey =

\ y

] mm

Fig. 45. Stelis cutucuénsis

Fig. 47. Stelis dalstroemii

.RUM

ICONES

Fig. 53. Stelis excavata

Spe 0 |

eae, nae Siaiee

AE ie)

Fig. 58. Stelis floresii

Fig. 57. Stelis flexuosissima

ICONES PLEUROTHALLIDINARUM 71

ICONES PLEUROTHALLIDINARUM 73

ZOU

1mm

Fig. 70. Stelis micacea

Fig. 74. Stelis pachyphyta

ICONES PLEUROTHALLIDINARUM

Fig. 73. Stelis ortegae

ICONES PLEUROTHALLIDINARUM 75

& 5 cm

NG OW AOQO =

2 oa _mm_,

Fig. 77. Stelis polycarpica 7 Fig. 78. Stelis porphyrea

Fig. 80. Stelis protracta

ICONES PLEUROTHALLIDINARUM

6. Stelis tridactyloides

Fig. 8

Fig. 85. Stelis serrulifera

Fig. 87. Stelis undulata

78 ICONES PLEUROTHALLIDINARUM

Ren?

Seth heel

pe eee

Se : 4 oy aes

oe ee

Are

he = ¥%

rine, et ce

Dey An Tet

Wray ee AL rAd at

2mm

eee

STELIS OF ECUADOR 79

93. Stelis acutula Luer & Hirtz, sp. n

Ety.: From the Latin acutulus, “small and pete ” referring to the ee

Species haec S. spathulatae Poepp & Endl similis

Plant small, epiphytic, caespitose; roots slender. epee a erect, slender, 1.5-2.5 cm

~~ enclosed by 2-3 | Oose, tubular sheaths. Leaf erect, coriaceous, narrowly elliptical, subacute, 3-3.5

le ca 0.5 cm eo the blade 0.4-0. "08 cm wide, narrowly cuneate

en into the subpetiolate base. ‘Inflorescence an erect, congested, secund, many-flowered raceme,

many flowers open simultaneously, 5-5.5 cm long including the peduncle 2-2.5 cm long, from an

ts infundibular,

with m

annulus below the apex of the ramicaul; floral bracts » oblique, acute, 2 mm long, apie ae

0.75 mm long; w,

gla abrous, connate basally, ovate, subacute, the peat sepal 2: 5 mm long, 1 5 mm wide, 3-v atten pin

lateral sepals oblique, turned forward, 2.5 mm long, 1.25 mm wide; petals y ellow, cgi

shallowly concave, 0.75 mm ay: 0.6 mm 5 - veined, oak apex broadly re ickened; -

yellow, subquadrate, 0.6 mm long, 0.5 mm wide, 0.3 mm deep exc pipe 03 m m m lon jong, the

dorsum filled with a callus, bilobed anteriorly, shor tly P i p Howly concave

with the se deflexed, —— triangular, acute broadly t te, hinged to the base of th

mm long, + i ™ tig ip ]

Morona-Santi Gual d San Juan Bosco, alt, 2500 m, 28 Mar. 1985, C. Luer, J. Luer,

A. Hirtz & W. Flores 10999 (Holotype: MO).

Pichincha: west Men ptr _—, et Azul, alt. 2800 m, 8 Jan. 1987, C. Luer, J. Luer & A. Hirtz

12471 (MO). Loja: n of Loja, alt. 2600 m, 2 Feb. 1987, C. Luer, J. Luer & A. Hirtz 12575

(MO).

This little species is related to the common and variable S. pusilla. The habits

are similar, but it is distinguished from the latter by a slender, congested raceme of

superficially similar, small flowers. The sepals are ovate, shallowly connate, and

directed forward. The petals are thin, obovate, and single- -veined. The lip is sub-

quadrate, shallowly cleft anteriorly on the thick, bilobed bar, and with an acute, de-

flected apiculum.

94. —— —— Luer & Hirtz, sp. n

Ety.: F Latin coralloides, “like coral,” tie to the branching, crystal-like deposits in the

floral parts ae leaves.

1! = en D, & Endl affinis 4 1 .. i ee, |

os ee i+ ott +, She Asfteact y

Plant _— ~~ densely caespitose; roots a 5 sec pon i es 3.5 cm long,

enclosed by a close, tubular sheath and 1-2 sheaths at = . Leaf erect, con arrowly ellipti-

cal, acute, 3-5. ee cm was 35 mm wide, gradually narr a belo w into a vetiole LL. I: ae cm long. Inflor-

escence an erect, loose, successively several- uals raceme, with most flowers open simultaneously,

to 10 cm long including the peduncle 3-4 cm long, from an annulus below the apex of the ramicaul with

a small spathe 3 mm long; floral bracts infundibular, oblique, acute, 2 mm long; pedicels 1.5 mm long;

vary 1 mm long; flowers light green, sepals glabrous, ovate, obtuse, connate basally, the dorsal sepal 2

m lo m ique, i i

thickened margin, 0.75 mm long, 0.6 mm wide, 1-veined, but with a vestigial pair at the base; lip ovate,

narrowly obtuse, 1 mm long or deep, 0.6 mm wide, slightly convex, with a minutely pubescent callus

_— the base, the base truncate, hinged to the base of the column; column stout, 0.5 mm long, 0.5 mm

e, the foot obsolescent, the anther and the bilobed stigma apical.

Morona flowered in cultivation at Ecuagenera, Gualaceo, 17 Apr. 2001, A. Hirtz

7710 eae MO), C. Luer illustr. 20069.

This small species, found in enicouel Ecuador at moderate altitudes, is dis-

tinguished by a caespitose habit with narrowly elliptical leaves borne by slender,

slightly shorter ramicauls, and a delicate, loose, successively flowered raceme. The

tiny, light green flowers appear similar to those of the widely distributed S. spathu

80 ICONES PLEUROTHALLIDINARUM

lata with converging lateral sepals. The petals are similarly cucullate, single-veined

and concave, but the lip is entire, narrowly ovate, slightly concave, and featureless

save for a brush-like callus at the base. The anther cap is proportionately large,

nearly as large as one of the petals.

Within the substance of the floral parts and leaves are branching clumps of a

coral-like substance, similar to the scattered, crystalline deposits found in so many

species of pleurothallids.

95. Stelis exigua Luer & Hirtz, sp. nov.

Ety.: From the Latin exiguus, “small,” referring to the habit and the flowers.

eS ee . ak ee als oe eo RS ee NCE

TS ee “1 Lk a. | ae ¢t lahell

enhohiannon_cak yey . leet Pin ‘ m

giore q guitur.

Plant very small, epiphytic, caespitose; roots slender. Ramicauls erect, slender, 4-6 mm long,

enclosed by - tubular sheaths. Leaf iaceous, elliptical, sul to obtuse, petiolate, 10-18 mm

long including a petiole ca. 2 mm long, the blade 3-5 mm wide, cuneate below into the petiole. Inflores-

cence an flexuous, congested, distichous, successivel fl d bey oh eee Teka es teal

ing the peduncle ca. 1 cm long, with 2-3 fl r Se ed EE

of the ramicaul; floral bracts oblique, acute, 1.5 mm long; pedicels 0.5 mm long; ovary 0.5 mm long;

minute, purple fleshy, glabrous, ovate, subacute to obtuse, 1-veined, the dorsal sepal 1.3

mm long, 0.8 mm wide, connate basall to the | I Is, the lat 1 1 eH 1 W

facing inward, 1 mm long, 0.7 mm wide; thin, suborbicular, shall wly concave: broadly rounded

ya ag apex, 0.5 mm long, 0.5 mm wide; lip thickly subobovate, 0.7 mm long, 0.4 mm wide, 0.25 mm

thin the y obtuse, apical margin, the disc callous, minutely

hing ion at ey | + eer eee ae A en

long, 0.6 mm wide, with the anth pical, the sti vi fe : uta :

Loja: road to TV tower east of Yan alt. 2600 m. .

Ww Flores 10803 (Holotype: MO). gana, alt. m, 22 Mar. 1985, C. Luer, t 2 Luer, A. Hirtz &

ef o

This tiny species, apparently endemic in southern Ecuador, is characterized by

the caespitose habit: elliptical leaves somewhat longer than the ramicauls; and an

erect, congested, distichous raceme of minute, successive flowers. The sepals are

Ovate with a single vein. The dorsal sepal is larger than the laterals that are held

forward. The petals are suborbicular,thin, and single-veined. The lip is obovoid

and shallowly concave at the obtuse apex.

96. Stelis hispida Luer & Hirtz, Sp. nov,

From the Latin hispidus, “with short, rigid hairs,” referring to the sepals.

inept ediocris scande ns longirepens, racemo multifloro foliis ellipticis longiore, sepalis ovatis

t li at . +1 * takue tt a a SE

basim transverse calloso disti inguitur, = ees

Boy aa Shep ages the rhizome stout, erect, 2-5 cm long between ramicauls;

sheaths. Leaf erect, coriaceous, muse ni aig long, proliferating, enclosed by 2-3 tubular, imbricating

less than 1 cm long, the blade 1-1.3 cm wi ee — Petiolate, 3-5 cm long including a petiole

distichous, sublax, many-fl 5 » Cuneate relow into the petiole. Inflorescence an erect,

including the peduncle 5-8 cm long. from an «4.45 f .

£ ot,

ly, 12-14 cm long

1; floral bracts tubu-

lar, obtuse, 2-2.5 mm long: pedicels 1.5 smells,

ispidoes | “> Tam long; ovary 1 mm long; sepals brown, glabrous externally,

- on Outer halves, the dorsal sepal ovate, obtuse, 3,3 mm long, 2.3 mm pert gree ined, connate

obtuse,

1 _ forward; psd green, membranous, semilunate, broadly

: concave, 0.6 mm long, 1.5 mm wide, 3-veined; n,

See et 0.75 mim long, 0.9 mm wide, 0.9 mm ecibi taeoe ennai ean ome

apex, notched above a cleft glenion, the base with a ‘oun: callus, truncate, hinged to

. column-foot; terete, I mm long, 1 mm wide, with a short foot, the anther and

STELIS OF ECUADOR 81

Pichincha: f eet ie Ww ia: 3. e 1 Py ie a

A. Hirtz & X. Hirtz 220 (Holotype: SEL), C. Luer illustr. 20058. papas geciecceg mt.

This species is apparently rare and endemic in north-central Ecuador where it is

known from a single collection. It is distinguished by the scandent, ascending to

erect rhizome that bears ramicauls shorter that the intervals between them. The

elliptical leaves are surpassed by the sublax raceme of small flowers. The sepals

are ovate, hispid within, with the laterals free and held forward resembling a synse-

pal. The petals are membranous and transverse with a broadly rounded apex. The

lip is deeply concave anteriorly with the apex thin and rounded, and the bar is

deeply notched.

OF. on juncea Luer & Hirtz, sp. nov.

Latin junceus, “slender like bulrushes,”” Linseed to the oe — habit.

i ies haec S. pusillae H.B.K. affinis d habi j Itil gi

bus, et ie trinervi:

um in size, epiphytic nadie rani i ly caespitose; roots slender. Ramicauls ns

very slender, 9-16 cm long, with a tubular she.

Leaf erec riaceous, narrowly ca acute petiolate 5 5.5-8 cm long including the petiole 1 vty cm

le. Inflorescence an

long, the blade 0. 8-1 cm wide, narrowed into the petiok an erect, secund, congest-

ed, si simultaneous. i uSLY, i

ing the sehacle ei ca. re 5 cm long, with a sate ca. 8 mm lon ng, at an annulus below the apex of the

ramicaul; floral bracts tubular, acute, 2 mm long; pedicels 1 mm long; ovary 1 mm long; flowers light

yellow; glabrous, broadly ovate, broadly obtuse, connate basally into a short tube, dorsal sepal

2 mm long, 1.5 mm wide, 3-veined, the lateral sepals held forward, 2 mm long, 1.5 mm mi 3-veined;

petals — truncate, shallowly concave, 0.5 mm long, 0.75 mm wide, 3- veined, with the margin of

the apex narrowly thickened; lip subquadrate, 0.75 mm long, 0.6 mm wide, 0.3 mm deep, concave an-

teriorly see th the apex peoery niet and minutely apiculate, the bar: rage! into ioe caven spb the

base thickly pu

Bac” 1 mm wide, the sce por stigma oe with the stigmatic lobes pobicog

Tulcan ~ Maldonado, alt. 2400 m,

16 16 Mar. 1991, C. Luer, J. Luer, J. del Hiern, A A. &X. Hirtz 15086 cman MO).

This species has been found in northwesternmost Ecuador. It is characterized by

proportionately long, slender ramicauls, and narrow, elliptical leaves that are sur-

passed by slender, congested, secund racemes of numerous, minute flowers. The

sepals are ovate and obtuse with the laterals more or less pointing forward. The

petals are cuneate and truncate. The lip is concave anteriorly with a minute apicu-

lum, and with the bar cleft. The lobes of the stigma are elongate, protruding beyond

the margins of the lip to either side.

98. Stelis poculifera Luer & Hirtz, sp. n

Ety.: From the Latin Ai “cup-bearing,” refering to the yaar

Planta parva dense caespito g lliptico longiore,

sepalis ovatis glabris similibus | lateralibus non expansis, petali iformib i is, et labello

Plant small, epiphytic, ag caespitose; roots slender. a erect, slender, 2.5-5 cm long,

losed by 2-3 tubular sheaths. Leaf erect, coriaceous, narrowly elliptical, acute, petiolate, 2.5-5 cm

long including a petiole | a cm ca the blade 5 mm wide, attenuate below into the petiole.

rect, strict, very congested, simultaneously many-flowered raceme 8-10 cm long including th

ee 3-4 cm long, from an annulus below the apex of the ramicaul; floral bracts oblique, acute, 2.5

m long; pedicels 1.5 mm long; ovary | mm m long; sepals yellow, glabrous, ovate, subacute, 3-veined.

robe basally, the dorsal sepal 3 mm long, 2 mm w wide, the lateral sepals held forward, opposite when

expanded, oblique, 2.75 mm long, 1.75 mm wide; petals brown, broadly ¢ — ae 1- pei

shatiowly concave, 0.6 mm long, 0. iA mm a _ haves, blong-o¥0 oid, 0.8 mm long, 0.6 mm .

0.3 mm deep, de nterior surface « cxcarie tenes ae

shelf-like bar, the dorsum with a broad p

Po eel at the hase

inged

tout. 0.5 mm long prpanharleg SS a panel

stigma apical.

82 ICONES PLEUROTHALLIDINARUM

Pichincha: wet forest, west flank of ap Iliniza, Cerro Azul, alt. 2800 m, 23 Jan. 1987, C. Luer, J.

Luer & A. Hirtz 12453 (Holotype: Mi

This species is apparently es in east-central Ecuador. It is characterized by

the small, densely caespitose habit and a densely many-flowered raceme surpassing

the narrowly elliptical leaf. The sepals are ovate with the laterals held forward: the

petals are thin, cuneiform and single-veined; and the lip is longer than broad with

the anterior half deeply excavate.

99. Stelis serpens Luer & Hirtz, sp. n

slags From sa ccs Serpens, “crawling,” fering to the reer rhizome.

longiore, petalis trans-

versis incrassatis et labello trilobo distinguitur.

Plant medium, epiphytic, repent, 15 cm long - nla ee stout, 1.5-2 cm long between

+ roots slender. Ramicauls der, 3-5 cm long, enclosed by 2 tubular sheaths

and another at the base. Leaf erect, coriaceous, cllpical, iluaied to — Jagepad 6-7.5 cm long

including a petiole ca. 1.5 cm long, the blade jade 0.9-1.2 mm 0 the

le 1-1.5cm cm long, from an annulus below the apex of the ramica 1; floral bracts oblique, acute, 3

mm long; pedicels 1-1.5 mm long; ovary 1 mm long; flowers light asc nodding; sepals glabrous

externally, microscopically pubescent within, the dorsal — oblong, obtuse, convex, 3.25 mm long, 1.6

mm wide, 3-veined, connate bas st the | ls, the lateral ] te, obtus e, 2.8

sekeal babe coac 2 . lobed, 0.8 mm og 0.5 mm wide vig the lateral lobes, th

» concave, Seal ¢ lateral lobes below th

base of the midlobe, the base t “spinels ow the peddle, obtuse rse, overlapping the

ing. 5 we wide aia mn roa st stout, 0.5 m

a — one TV me ng of Yangana, alt. 2750 m, 22 Nov. 1985, C. Luer, J. Luer, A. Hirtz &

Chinchipe: Cajanuma Ran

psn sey pong —— prep gg south of Loja, alt. 2750 m, 21 Mar. 1985, C. Luer, J. Luer, A.

This species, apparently endemic in southeastern Ecuador, is characterized by

the repent habit; elliptical leaves about as long as the ramicauls; and a congested,

secund raceme slightly longer than the leaf. The flowers are very small with con-

rages lateral sepals; transverse sepals with the apex irregularly thickened external-

y; and a three-lobed lip. A pair of obtuse lateral lobes overhang the base of the

concave middle lobe. The lateral | appear to be a modification of a bar.

100. Stelis tempestuosa Luer & Hirtz,

rf aig the Latin tempestuosus, “stormy,” referring to the cold, wet, windy pass where the

reid vices mediocris, foliis is anguste oblongis ramicaulibus gracilibus ‘on minusve .

i senalic late ollinei~: : Se eal hk at

apice transverse incrassatis, labello cuneato anc Si i ieee Nae a

sender 2-7 em on medium in size, spiny. densely caespitose; roots slender. Ramicauls erect,

eae sities tubular sheath near the middle and 2-3 sheaths at the base. Leaf

> a to sul “ 4. i 1 ind: =

cm tale a the blade 1-1 se 4m ic —— be cm i including an indistinct oxen 1-2

con

prom : ce

uous, many-flowered raceme with several flowers open simultaneous! 312

ie tong including the rsa 1-2 cm long, subtended bya spp spathe 5 7 an long, from peaiitg

1.5-2 mm long; pedicels 0.75-1.5 mm long;

Ovary | mm long; sepals yellow. ellow, glabrous Fas

1.6-1.9 mm wide, 3-veined, connie eo. - obtuse, the dorsal sepal erect, 1.75-2 mm lon,

below the middle, 1.8 mm long, 1 1

a mm wide, ta ee gee yellow, thin, uadrate, shallowly

the subtruncate, transversely thickened 9 ig

ee margin, 0.6 mm long, 0.6 mm wide, 1-veined; lip

Saas ge Ome 0.6 mm wide, 0.2 mm thick, shallowly

the base with a flat,

STELIS OF ECUADOR 83

lal one ee Nn hi ge £ th column stout, 0.8 mm long 0.8 mm wide, the

1 the anther apical ionatel une eed obsolescent.

Bea Colechine: cloud forest east of the pass east of Loja, alt. 2750 m, 21 Mar. 1985, C. Lue

Luer, A. Hirtz & W. Flores 10711 (Holotype: MO); same area, alt. 2700 m, 21 Sept. 1980, C. ress

Luer & A. Hirtz 5522 (SEL); south of the pass south of Jimbura, alt. 3050 m, 21 Feb. 1986, C. Luer, J.

Luer, A. Hirtz, W. Flores & A. a 12014 ).

RiaaunKentia : east O een Gualaceo and Limé6n, alt. 2900 m, 16 Feb. 1986, C. Luer, J.

Luer, A. Hirtz, W. Flores & a Embree 11815 (MO); same area, alt. 3180 m, 16 Feb. 1986, C. Luer, J.

Luer, A. Hirtz, W. Flores & A. Embree 11806 (MO).

This species, — endemic in the mountains of southeastern Ecuador, is

characterized by slender ramicauls about as long as narrowly oblong leaves that are

more or — pease: in length by a congested raceme of very small, yellow seuss

The sepals are glabrous and elliptical with the laterals held forward. The pe

broadly ee The lip is cuneate, longer than wide, with the apex ae

rounded and the base with a broad, cellular-pubescent callus.

ICONES PLEUROTHALLIDINARUM

Fig. 97. Stelis juncea

Stelis acutilabia 9, 28

Stelis acutula 79, 84

Stelis adrianae 36, 65

Stelis amoena 36, 65

Stelis applanata 10, 28

Stelis barbellata 38, 66

Stelis bivalvis 11, 29

Stelis bolivarensis 11, 29

Stelis brachiata 12, 29

Stelis coeliaca 12, 29

Stelis congesta 39, 66

Stelis coralloides 79,84

Stelis cutucuénsis 40, 67

Stelis dalessandroi 40, 67

Stelis dalstroemii 41, 67

Stelis digitata 13,30

Stelis dirigens 41, 67

Stelis distans 42,68

Stelis dodsonii 42, 68

Stelis dromedarina 14, 30

Stelis elongatissima 43, 68

Stelis embreei 14, 30

Stelis exigua 80,84

Stelis exilis 45, 69

Stelis falcifera 45, 69

ICONES PLEUROTHALLIDINARUM

INDEX TO SCIENTIFIC NAMES

Stelis ferax 46, 69

Stelis fissurata 15,30

Stelis flava 15,31

Stelis flexilis 16, 31

Stelis flexuosissima 46, 70

Stelis floresii 47,70

Stelis florianii 16, 31

Stelis formosa 47,70

Stelis galeola 17, 31

Stelis gigantea 48,70

Stelis glaberrima 48, 71

Stelis globulifera 49, 71

Stelis hirtzii_ 7,27

Stelis hispida 80, 84

Stelis hoeijeri 49, 71

Stelis jimburae 50,71

Stelis juncea 81, 85

Stelis kuijtii 50,72

Stelis limonensis 51, 72

Stelis longihirta 51,72

Stelis longissima 52, 72

Stelis luteria 17,32

Stelis macilenta 52,73

Stelis matula 8, 27

Stelis micacea 53, 73

Stelis minutissima 53, 73

Stelis nepotula 54, 73

Stelis nycterina 18, 32

Stelis ortegae 54, 74

Stelis pachyphyta 55, 74

Stelis parvipetala 18, 32

Stelis phil-jesupii 55, 74

Stelis pilifera 56, 74

Stelis platypetala 20, 33

Stelis poculifera 81, 85

Stelis polycarpica 56, 75

Stelis porphyrea 57, 75

Stelis preclara 21, 33

Stelis procera 57,75

Stelis protracta 58,75

Stelis pubipetala 58, 76

Stelis purpurella 59, 76

Stelis regina 21,33

Stelis repanda 22, 34

Stelis rosamariae 22, 34

Stelis saccata 7,27

Stelis sanchezii 23, 34

Stelis sanctae-rosae 59,76

Stelis serrulifera 60,77

Stelis situlifera 24, 35

Stelis sumacoénsis 25, 35

Stelis tempestuosa 82, 85

Stelis tridactyloides 61,77

Stelis undulata 61, 77

Stelis uxoria 62,77

Stelis velivolva 25, 35

Stelis vesca 63,78

Stelis villifera 63, 78

Stelis vollesii 26, 35

Stelis yanganensis 64, 78

Stelis zarumae 64, 78

Stelis zelenkoi 8, 27

ADDENDA TO LEPANTHES OF ECUADOR

ABSTRACT

The following | 12 Sp i ise th ber of species of the genus Ley

Ecuador to 332. g bered i I Seer pI pry

New species:

uisepala Luer

Scouts! “et Luer & Portilla

es lynniana Luer

Lepanthes Luer

Lepanthes persimilis Luer & Sijm

Lepanthes tridactyla Luer

Lepanthes werneri Luer

cme bibarbullata Luer, sp. nov. Fig. 310.

From the Latin seer ox pct “with two stir a sepeassoat to the a of the els

snucve

hese rva q' : aera eS

aequilongo, sepals obtusis breviter acuminatis, F ] S

Plant small, epiphytic, <i roots slender. Ramicauls roa erect, gga mm ne.

enclosed by 3-4 microscopically scabrous, lepanthiform - broadly cuneate

ular, reticulated, rounded at he apex, 12 mm long, 11 mm wide, the we cial eg

rounded, contract ted into a ai 1 mm long. Inflor —". dps eae bove the leaf by a

successively several long; pedicels 1- -1.5 mm long;

slender peduncle 5 mm m lon ng: floral bracts 0.5 mm long; i

brvad obtuse, shortly acuminate, the

long; ll brous, ly denticulate, eo

g — ye ar po sparsely cage bl to the lateral sepals for 0.25 mm, the

ly oblong, obscurely bilobed, exter 0.3 mm long, 0.75 mm w Ee ce : vm

ean dulate, 1 mm long, ! mm wide, the abruptly

a emanatce of the column, column terete, 0.75

contracted into a slender claw, att

mm |

, the anther and stigma apical. aa

: . Novem ,

ECUADOR: Without locality, cultivated by J & L Orchids 799-593, Easton, CT, Nov

C. Luer 19440 (Hol : MO).

et terized by the small

site, It is charac

species is related to L. calodictyion. less spreading, non-crenate

Thi

habit with suborbicular, reticulated, more OF

leaves, and a congested raceme arching above the surface. a — a

sparsely denticulate, broad, obtuse and shortly acuminate. nab as tae

transversely oblong with a long, filamentous process a soe chanticolet,

of the lower lobe and passing behind the lip. The lip is

and attached to the column by a slender claw.

88 ICONES PLEUROTHALLIDINARUM

Lepanthes chilopsis Luer & Hirtz, sp. n Fig. 311.

Ety.: From ~ — cheilopsis, * = a ise se reference to the lace

me 4 nlite} conaln di te

acuto, petali im lobulati , labello gl b to-cordato ated lobis basalibus | columnam

amplectentubus sing

Plant ize for tt b pende: up to 30 1 e rhizomes

slender, 2.5-3 mm long between sh tet ms i nen long-ciliate, vA wai roots

slender, from nodes near the base. _Ramicauls 0. a long-ciliate, lepanthiform

sheath. Leaf pend y long-<iliate, elliptical, subacute, 9-10

mm lon ng. Infloresce

mm long, 3.5-5 mm wide, 2 mm thick, the bas :

iP nce a

single ve. borne by a peduncle ca. 1.5 1 floral bract long-

spiculate, 1-1.5 mm long; pedicel 1.5 mm — ovary ssp-sunig ce 0.5 mm m long; sepals mem-

the dorsal th purple, stripe, suborbicul ones concave, 5

translucent

mm long, 4 mm wide, 5-veined with faint accessories, “st connate to the lateral sepals, wee aes

S translucent yellow, faintly suffused with purple setae, connate 1.5 m

shortly bifid synsepal, 4.5 mm long, 3.8 mm wide, 4-veined; petals lide diverging, minutely =.

_ oblong, obtuse, 1.5 mm long, 0.25 mm wide, with the margins involute, the base with a lobule at

angles; lip yellow-orange, glabrous, ovate, obtuse, protuberant, with the base cordate, clasping the

column, 2.25 mm long, 1.75 mm wide expanded, the base connate to a minute Sibiu from the base

of the column; column 1.5 oe oe apical.

ECTIAnnNpD. A.

, at the junction between Rio Colim-

bo and Rio Upano, alt. ae 1 Mar. 2001, C. Luer, J. Luer & A, Hing 19588 (Holotype: MO).

This species is similar to L. pilosella and its relatives, but the leaves are thick

and elliptical, and covered by a fine, long pubescence. The sheaths are also finely

long-pubescent. The inflorescence is a single, erect flower produced successively.

The dorsal sepal is concave and, the synsepal is ovate and acute. The petals are

small, wrens tor and bilobed at the base. The lip is ovate and protrudes forward

beneath the col

Lepanthes co dila bia Luer , Sp. Nov. Fig. 312.

Ety.: From the Latin cordilabius, Pas rataped lip,” referring to the cordate labellum.

centibus. piles ses “i » Sed floribus majoribus, sepali o dorsali o ongipubes-

P » petalis breviter cileecastibus¢ et st glabro

cordato col umnam sauce differt.

Rib iei small, pioneer pendent, up to 10 cm m long or longer, the rhizome slender, 2-4 mm long

2 infundibular, lo: Remaiesuls

y coriaceous, glabrous,

0.5 mm long, peat wi I ciliate sheath. Leavy. pende

aes n distichous, thickl

broadly ovate, obtuse, the b' ade 3-5 mm long, 4.5 mm wide, the base abruptly

5 mm a successively 2- to 3 flowered renew up to 3 mm long, borne

g-ciliate, 1 mm long; pedicels 1.5 mm long; ovary long-ciliate-

ine

equal, triangular wit wih Ne fecromiice: Bo ote red, glabrous, cordate with the

sie ose ie column, 15mm long, 1 mm wide expanded the apex trosdly rounded the al

Webel west cere bsaurcacent J ; column terete, 1 mm

BCTIANRM?D. F.

Loj d Z It. 2200 m, 9

€ shortl t.

and cordate with the sides embracing the colusms oe ee Sete

———————— Eee ee

LEPANTHES OF ECUADOR 89

Lepanthes exigua Luer & Jost, sp. nov. Fig. 313.

ty.: From the Latin exiguus, “‘v —_ small,”’ in a to 9 —

_ Inter species subgen. B hycl g lis mi pice ciliati

Rast: tals 4 as ba vs 2 lahall — a

f Ls r

Plant very aie —— mami eS the thizomes slender, I 5-2 mm long between rami-

“sie enclosed by 2 long-ciliate, imbri

the sent Ramicauls 0.5 mm ep concealed by 1 long-ciliate, lepanthiform sheath. Leaf

a in the natural position, coriaceous, le lentiform, min utely short-ciliate, broadly lliptical, obtuse, 3-

; inal . Hae

ong. Inflor-

escence a single flower, borne by an ascending aa 1mm long from near the apex of ari ramicaul;

floral bract long- -iliate, 1 mm long; pedicel 1.2 mm long; = costate-dentate, 1 mm long; sepals

yellow with the veins purple, membranous, microscopically ci iate-pubescent, the season broadly

obovate, erect, obtuse, 3 mm long, 2 mm wide, 3-veined, | sepals, the lateral

sepals oblong, connate 1.5 mm into a bifid lamina, 3 mm long, 3 mm wide, 4-veined, the oblique,

ap ‘age petals yellow, suffused with purple, pubescent, obliquely bilobed, 0.8 mm long, |. 3

mm w e ey opposite, ays the upper lobe 1-veined; lip purple, minutely pubescent, nt, cordate-

iciabed, 2 mm long, 2.6 mm wide expanded, with the apical lobe triangular, acute, with the lateral or

basal lobes ie dhs ali rounded

the column; column 1.5 mm long, , the anther and the stigma apical.

ECUADOR: Morona-Santiag d between Mendez and Paute, alt. 1890, 14

May 2000, L. Jost 2042 rapa MO), C. Luer illustr. 19424,

This tiny species occurs in southeastern Ecuador at a medium altitude near 1900

meters above sea level. The minutely ciliate leaves are three millimeters or less

long, and the single-flowered peduncle is barely one millimeter long, i the

rhizome and peduncle with long-ciliate sheaths. The sepals are obtuse, :

apiculate; the petals are oblique and single-veined; and the lobes of the eine Pp

enclose the lip.

Lepanthes geminipetala Luer & Portilla, is, nov. Ai Fig. 314.

se From ge Latin Senteee with tw inned petals, in reference to the petals.

di foliis longipub tibus lo dorsali concavo, synsepalo sub-

petalis geminatis pubescentibus, labello glabro bilobo,

qu Mika apicibus obtusis breviter acuminatis,

lobis columnam amplectentibus distinguitur. to

th branches or

Plant average in size for the Opie epiphytic c, pendent, wi b 2 lon — epntirn

er, me thizomes slender, 5-7 mm long bet tween ramicauls, ¢ enclosed Y pe * concealed by a

shear

ac long-ciliate,

longi lepanthiform sheath. Leaf pendent in the eral pen. ic eons a petiole A 2mm

iptic se, 9-11 mm long, 4-5 mm wide, 2 mm thi tee elk from near the apex

ance’ a single flower, borne by an ascending 3 cae ovary minutely cellular-

ofa emitant: floral bract soe seine: 2-3 mm long; sLansaigne ~ , suborbicular, obtuse

glandular, 1 mm long; sepals light rosy brown, mem eined, barely connate to the lateral

oes = actetieete at the tip, concave, 4 mm long, 3.5 mm wide, aad 45 mm long, 3.75 mm

sepals, teral sepals ~~, 2mm into a subquadrate, left into two oblong, obtuse lobes, the

Outer lobes 2 mm long, 0.5 mm ere the inner lobes 1.5 mm qe subfalcate, clasping 4 poms

dark purple-brown, bilobed, “U-shaped,” wi b obtuse, ca. 1.5 mm

with an oblique, linear callus on the outer surface, the apex anther and on stigma

q th the

with

ed, the base connate to the base of the column; column 1.75 mm long,

apical

Gualaceo, 4

ECUADOR: R: Azuay: Soldado, Molleturo, southwest of Cuenca, cultivated at Ecuagenera,

Mar. 2001, C. Luer 19656 (Holotype: MO). ‘ d its relativ

Superficially, this species is similar to L. pilosella an

leaves are thick and elliptical, and covered by a fine, long erect flower produced

are also finely long-pubescent. The inflorescence - ees is S

successively. The dorsal sepal is concave and, the eee erect behind the lip and

widely bifid. Each petal, divided into two gmat the column.

column. The lip is broadly bilobed with the lobes

es, but the

The shi

90 ICONES PLEUROTHALLIDINARUM

Lepanthes inaequisepala Luer & Portilla, > nov. g. 315.

: From the Latin inaequisepalus, “‘with unequal sepals,”’ referring to the markedly tienen size

and shape of the dorsal sepal and synsepal.

Species haec L. Pe ase Schitr. affinis, sed Scipio oe age dorsali jae a

acinmi ami-

1 is 4: 4: 3° or

oe i a

lender. Ramicauls slender, le 5-8 cm long,

enclosed by 6-7 microscopically ci te, lepan orm sheaths. Leaf erect, coriaceous, elliptical, acute, 4-

cm — 1-1.2cm wide, , the base cuneate into a petiole ca. 5mm long. Inflorescence a, suberect,

y p to 10 cm long including - filiform peduncle

Pepeny 2; floral t Fe long; pedicels 4 05 mm long; y carinate, 3mm — sepals color?,

carinate glabrous the do ! casi anes as 8 mm long, the entire

length length 20 mm to a 8.5 mm wide, (puseualy connate ‘to the lateral al sepals for 3 mm, the lateral sepals

mm in : ng, 7 mm

base, the acute ips free 2 mm; petals pu pubescent, transversely sigmoid, 1 1 eset a emtese = lobes

obtusely , pubescent, triangular with

1.75 mm long, the an

the stigma apical.

ECUADOR: Morona-Santiago: Sinai near Macas, alt. 1800 m, flowered in cultivation at Ecuagenera,

Gualaceo, Oct. 2000, J. Portilla 2114 (MO), C. Luer illustr. 19445.

This species is related to L. elegantula. The relatively large flowers are borne in

a long, flexuous, distantly flowered raceme that far surpasses the acute leaf. The

sepal i is large and concave with a long-acuminate tip. The synsepal is nar-

Lepeetins laxa Lites & detcbonen. Sp. nov. Fig. 316.

Ety.: From the Lati: »” referring to the distantly flowered raceme.

___ Species ha haec L. done Set affinis, sed pedunculo fracti ynsepalo infra medium ovato

flexo

truncato, labelli laminis etiam ei differt. Ontiguis, petalorum lobis rig lobo superiore oblique

all to medium in ze, epiphytic, caespitose; roots sle 4-5

cm long, poor by 6-7 close, ‘ae lepanthif bes daa. pe emia clipica-obovae,

obtuse to rounded at the tom 2-2.5 cm a I- ie .2 cm wide. the hace c :

suberect,

uous, loose, successivel several-flowered raceme he cm

em og: floral | bracts 2 mm long; pedicels 3-6 mm long; ovary

long. Inflorescence an

— including the filiform p edn 7 cm

abo 25 mm long; sepals co!

: bove the mi — aera oe oly tra triangular apex, 13 mm long, 7.5 mm wide, concave basally,

rm a shallow, bowl-shaped, sepaline cup, the lateral sepals

ovate in lower half, narrowed above whan concave basally, 12-13 mm long, 7-8 mm wide together

expanded,

long, 4 mee contiguous, acu te; petals pubescent, transversely linear-oblong, 0.8-1 mm

ly obtuse; lip bilaminate, Pa with the upper lobe obliquely truncate, the lower lobe longer, narrow-

ciliate at the apex, 1.6-18 mm ra ao with rari ends, slightly convex, minutely pubescent,

» the connectives ae obli ue, cuneate , the body connate to the

at de the appendix narrow narrowly obl sts ™ rasa

hee eae tind oC

ECUADOR: Zamora-Chinchipe: Cordill i Gual

: era del C; Biren ec ee Ll ss

PERU: Hua — ontilla 2164 (Holotype: MO), C. Luer 9444 > :

re — d by P. i Se :

- =m by fae sa = ee cultivation at Hasselt, Belgium, 13 Aug.

Sa ete erm x talon are imi

sepsis somewhat similar to those of

L. nycteris and relatives. The lobes of the petals and lip are oblong.

LEPANTHES OF ECUADOR 9]

Lepanthes lynniana Luer, sp. n Fig. 317.

Named for Lynn O’ raped: shee se Howell, MI, who cultivates this species, and submitted it

for iAbatification.

Species haec ‘a ciliari Luer & Hirtz affinis, d sepalis reflexis duplomajoribus proporti g

tioribus 2. GSE SI Pa 2. : ee

Plant small to medium in size for the subgenus Brachycladiwm, epiphytic, jas the rhizome : 3

mm long between ramicauls, enclosed by 1 loose ig

Ramicauls 1 mm long, enclosed by | infundibular, long-ciliate sheath. Leaf pendent, distichous, thickly

coriaceous, glabrous, ovate, obtuse, 4-5 mm lon ng, 4-5. 5 mm wide, the base rounded, contracted into a

petiole ca. 0.25 m ——e istant, successively 2-flowered raceme, borne by

a slender, ascending peduncle 4-6 mm long; floral bracts 1 mm long; pedicels 2-2.5 mm long; ovary |

m long

vate

long, 3.2 mm wide, 3-veined, connate to the lateral or for 1 mm, the lateral sepals elliptical-ovate,

oblique, subacute, apiculate, 3.5 mm long, peor 1.5 mm, 3.5 mm wide together, 2-veined;

reniform, pubescent, 0.5 mm long, 1 mm wide, the pended obtuse; lip pubescent, long-ciliate, bi-

lobed-hippocrepiform, 1.5 mm long, 1.5 mm wide, "he apex broadly rounded, th the basal lobes obt

embracing the column, the body broad, connate terete, 1.5 mm long,

the anther and stigma apical.

ECUADOR: Without collection data, obtained from José Portilla, cultivated in Howell, MI, flowered

Jan. 2001 by Lynn O’ Shaughnessy 00689 (Holotype: MO), C. Luer illustr. 19462.

This species is distinguished by the small habit with ovate, glabrous leaves, and

short, ascending, successively two-flowered racemes. Except for being twice larg-

er, the flowers are very similar to those of L. ciliaris. The sepals of L. ciliaris are

broader, with the dorsal broader than long, and the lateral sepals are free and diverg-

ia... The senals of Le are proportionatel narrower and reflexed, and the

g- pals of L. lynniana are proporti sf mall, those of L. ciliaris

laterals are semiconnate. The petals of both species are s 7 :

being twice longer and lunate in shape. The lips of both species are “horse shoe-

shaped.”

Lepanthes seteccnees | Luer, sp. nov. Fig. 318.

petals and

Ety.: From the Latin octocornutus, “eight-homned,” referring to ss eS . phere ip.

Species haec L. Pls wi f. similis, sed petalis .

bh a ‘ , ntucladium, epiphytic, pendent, long-repent, up 10 St

e hee Pa Yee ent enclosed by 2 long-spiculate

long or longer, the rhizome 3-4 mm lon g between ramicauls, each segment : breviated, 0.5 mm

sheaths with dilated, long-ciliate ostia, roots —, along the the rhizome. ws og elliptical,

long, newt by a long- 1p aeaiie: sheath. mm long.

bac 7-11 mm long, 4-5 mm w de, 3- ba . the base cuneate in ae floral es long-

lon; ad

siege atapcss tore b eapaniny 9 seme with purple-brown

pubescent, | mm long; pedicel 1 mm hae pin 0.6 mm long, S€ Ae at the apex, 8 mm

stripes, glabrous, the dorsal sepal deeply ne suborbicular, Dupage ; rounded .

veined, the lateral or

the

3.5 mm wide, 4-veined,

long, 7 mm wide unexpanded, 7-

eo gen eae apices, 8.5 aed der, the middle lobe

3 mm into a narrowly ovate, bifid synsepal

i Is glabrous, narrowly trifurcate, 2. Sites g. 0.5 me peace eee oa

longest, with basal lobes ca. 0.5 mm long; slabrou : the is —— :

mm long, 1.5 mm wide expanded, 3 mm i a ee pant the base

column, the lobes expanded, slender, 1 mm long, pile ria cose

of the column; column terete, 2 mm long, the anther . alt. 2200 m, 9 June

pn ia and Zamora,

2000. FE sii 329A pene MO), C. C tser il. 19426.

ies, known only from this

L. aes because of the large, orbicular, ual cone 5 pei eat saan

The lateral sepals form a narrowly ovate, a a ak of the base. The lip is

a long, middle segment and a short segment at 0 saw. erect, basal IODES.

subquadrate with sides clasping the column, and wi

92 ICONES PLEUROTHALLIDINARUM

is Luer & Sijm, sp. nov. Fig. 319.

Ety.: From the Latin persimilis, “very similar,” referring to its similarity to L. pilosella.

Species haec L. pilosellae Rchb.f. similis, sed sepalo dorsali quinqui-nervoso, petalis magnitudine

Plant ge in si th genus, epiphytic, pendent-repent, with branches up to 10 cm long or

longer, the rhizomes stout, subflexuous, 5-7 mm long between ramicauls, enclosed by 2 long-spiculate,

imbricating, lepanthife heaths: roots slender, fr des along the rhi icauls 1 mm long,

concealed by | long-ciliate, lepanthiform sheath. Leaf pendent or prostrate, coriaceous, densely ciliate-

pubescent, elliptical-ovate, subacute to obtuse, 11-14 mm long, 8-9 mm wide, 5-veined, the base con-

tracted into a petiole less than 1 mm long. Inflorescence a successive, single flower, borne by an as-

cending peduncle 1.5-2.5 mm long, borne laterally fi icaul; floral b iliate, 1.5-2 mm long;

pedicel 3 mm long; ovary smooth, 1 long; sepals glabrous, the dorsal sepal translucent, prominently

striped in purple along the veins, orbicular, concave, 5 mm long, 5 mm wide, 5-veined, free from the

lateral the lateral sepals connate 2 mm into an ovate, bifid lamina, 5 mm long, 3.75 mm wide, 4-

veined, with the api petals glabrous, oblong-triangular, subacute, 0.8 mm long, 0.4 mm wide;

lip glabrous, ovate-cordate, 2.3 mm long, 2 mm wide expanded, with the apex broadly rounded, with the

basal angles obtuse, with th id i ling if 13 gt y 4 +} 1 , th te to th

Ls i} eh. el. 3 ot . 7

i 13

ECUADOR: without collection data, obtained from Johannes Pap, flowered in cultivation, 11 Aug. 2000,

YE LA 84

byA P. Sij 991112 (F O),C Luer illustr 193

cially, this species is similar to L. pilosella and its relatives with ciliate-

pubescent, elliptical-ovate leaves. The sheaths are long-spiculate. The flower is

similar with a large, rounded, concave, purple-striped, dorsal sepal, and a bifid

synsepal. The petals are minute, half the size of those of L. pilosella. The lip is

considerably larger and ovate with the sides incurved, surrounding the column, and

ie ee rounded apex protruding beyond the column. The lip is longer

Lepanthes tridactyla Luer, Sp. nov. Fig. 320.

Ety.: wines the Greek tridactylos, “tt -fingered,”’ referring to the shape of the petals.

5 Suerte tate L. platyse| . wu : : E sobs imilis, s lo angustiore, petalis trifurcatis, et

“al ium in size for the subgenus Brachycladium, epiphytic, pendent, long-repent, up to 10 cm

sheaths with dilated, lon agi a Poland eyo

the rhizome. Ramica’ viated, 1 mm

descendi i a) iptical,

» 8-10 mm long, 2-3 mm wide, 3-veined. the b = ooniacouds, tong-pubcscent, elliptic

rescence a single flower, borne by a

pubescent, 1 mm long: pedicel 0.5 mm I

stripes, glabrous, the dorsal

wide, 7-veined, connate to th

oo bifid synsepal with acute apices, 6 mm lon

long, 1.75 mm wide expanded, with the

ECTIANRND. 7.

2000, F: Werner 329B (Holotype: MO), C. Luer illustr. 19427,

Loja and Zamora, alt. 2200 m, 9 June

pratysepala because of the large, erect, obovate, obtuse, shallowly concave,

LEPANTHES OF ECUADOR 93

pnts werneri Luer, sp. no Fig. 321.

ed for oe Werner, Universi of Bayreuth, who collected this species at the Estacién

Bcd s on Franci

Species haec L. ciliari Luer & Hirtz affinis, sed florib ijoribus, sep list iter p

lateralibus acutis, p q gul | g

bus differt.

Plant s small to medium i in size for the — Brachycladium, epiphytic, — ~ to 20cm

long out, 3-4 mm long between ramicauls, enclosed by 2-3

iste chucallin. weeks sinha Ramicaul 0-1 mm long, enclosed by 1 | iat

dibular, ciliated sheath. Leaf pendent. distichous, hikly coriaceous, | commas

4.5-7 mm long and 3.5-7 mm wid 5 ss Aweae ang

mm long. Inflorescence an ty loose, successively - to Wi raceme,

by a slender, suberect peduncle 4-6 mm long; floral bracts 1-1.5 mm long; pedicels 1. egex mh

ovary 1-1.5 mm long, nies pe ~~ tomy: spread, the dorsal papers He fos 0.5 man

pubescent within, = - 5 5m g, 2.5-3 mm wide, 3-veined, connate . the ene sh ran

the lateral sepals ce goa ra ‘diverging, omg gular, oblique coal wiglk are Jandular, 0.6 mm

1.75-2 mm wide, Ebel connate 1 mm; petals rose, transversely yl li " citelar-giandala,

long, 1.3-2 mm wide, the lobes equal, triangular, acute, oblique, — id BY lobes triangular,

bilobed, 1.5-2 mm long, 1.5-3 sf connate to the

erect, oblique with the basal angles acute to obtuse, € mbracing the = for Sie the stigma

base of the column; column terete, 1.5 mm long, with a long rostellum, the an

apical

sisi ca. 03

ECU ' $on Francisco station between Loja and Zamors 0 TO pon

Werner pres ieee: MO), C. Luer illustr. 19429 ithout collection data, 'L00508 (MO), C. Luer

la, stage ag in Howell, MI, foward Sept. 2000 by Lynn O'Shaughnessy -

illustr. 19423. : bor-

Lepanthes werneri is distinguished by the small habit oo _ 2 widely

bicular, glabrous leaves; short, ascending, suecessively-flowered t, while the

spread, purple flowers. The dorsal sepal is broad, _— Jlular-glandular. The

lateral sepals are widely spread, acute, apiculate and only cellular x Ck ae

lobes of the petals are triangular and opposite. The lip is simply

tire without

lobes erect, triangular and either obtuse or acute, and with the apex en

an appen ndix.

94 ICONES PLEUROTHALLIDINARUM

96 ICONES PLEUROTHALLIDINARUM

haan

Fig. 319. Lepanthes persimilis

ADDENDA TO BARBOSELLA, DRACULA, DRESSLERELLA,

LEPANTHOPSIS, PLATYSTELE, PLEUROTHALLIS, RESTREP.

SCAPHOSEPALUM, TEAGUEIA AND TRICHOSALPINX

ABSTRACT

Ilan Srhle. Ty. 7 lla

The following Ra es of Bark Dracula Luer,

anthes Sw., Lepanthopsis Ames, Platystele Schitr., Pleurothallis R.Br., Restrepia ord

Soca dracon Pfitz., Teagueia Luer and Trichosalpinx Luer are added to the genera

New spec

Fig. EL: Markco portillae Luer

ig. 10 rella sijmi

Fig. 11. Lepanthopsis calva Dod ex Luer

Fig. 12. Lepanthopsis ex Luer

2 Dartilla

Fig. 16. Pleurothallis adrianae Luer & Sijm

7 Te 41a )it.

Fig. 17.

Fig. 18. Pleurothallis ihe aha Luer & Hirtz

Fig. 19. Pleurothallis rostratissima i Luer & Portilla

Fig. 20 enbeai sextonii Luer

. 21. Pleuro

Fig. 22. Restrepia portillae Luer

Fig. 23. Scaphosepalum bicristatum Luer

: ‘noi

Fig. 29. berate Luer

Fig. 30. Trichosalpinx werneri Luer

Barbosella portillae Luer, sp. nOV tei aa

Ety.: Named for José Portilla of Gialaceds in whose a this species

pedunculo foliis parvis multilon-

Pe ease

erect, 4-5 man long, comme

Plant small, epiphytic, caespitose; roots slender. eous, linear-obovate,

2 thin, loose, tubular sheaths. Leaf erect, thickly coriac age

long , gradually narrowed floral bract thin, tubular, 2 mm

flower, produced oy. peduncle slender, erect, Ladi ns 2.5 mm long; 0

long, enclosing the pedicel; pedicel s 1 mm long, sepal erect, wly ’

de er nearly | 180 narrowly tri-

f y linear, y

; olive

angular, acute, 15 mm long, 3.5 mm wide, 3-veined; | mm wide; lip olive se thick, fleshy,

obtuse, thickened above the middle, 6 mm long, 0.5- the base rounded, deeply concave, ee

a? 3: 5 mm long, 1.5 mm wide, = mapas disc with a central cavity, with 3 io paralie!

ated umn

98 ICONES PLEUROTHALLIDINARUM

calli above the middle; column greenish white, stout, longitudinally winged above the middle, 3.5

io the llate. the foot thick with tt f th , the apex free, bulbous

Mar, 2001, C. Luer 19796 (Holotype: MO).

This small, caespitose species, known only from southeastern Ecuador, is unique

in the genus. In only one other species, the lateral sepals part, but they remain

connate toward the base. A partial parting of the lateral sepals occasionally occurs

in other species, especially in old flowers. In B. portillae, the narrowly triangular

lateral sepals are totally free, and diverge nearly 180°. The flower is light brown

and held well above the thick, elliptical leaves. The petals are narrow and thick-

ened. The lip is basically similar to the other Andean species.

gui, alt. 1000 m, cultivated at Ecuagenera, Gualaceo, 7

antonii Luer, sp. nov.

Ety.: Named for A.P. Sijm, who culti hi ies at his Orchid kwekerij Venhuizen in

Holland.

s Species haec D. erythrochaete (Rehb.f.) Luer leviter similis, sed labello minore, epicheilio 45° de-

Plant medium in size, epiphytic, caespitose; roots coarse. Ramicauls erect, slender, 3 cm long,

enclosed by 2-3 loose, tubular sheaths. erect, thinly coriaceous, carinate, narrowly linear-obovate,

acute, 20-24 cm long, 1-2 cm wide, gradually 1 below to tt duplicate, subpetiolate base.

Infloresce essively few-fl , borne by a slender, horizontal to descendin pedun-

cle 9-11 cm long, from low on the ramicaul; floral bract tubular, 10-12 mm long; pedicel ca. 15 mm

long; ovary thick, 5 mm long, round in cross section; white, yellow-brown at the base, shortly

pubescent within, the dorsal sepal transversel vate, 9 mm long, 13 mm wide, connate to the lateral

sepals for 5 mm to form a broad, shallow, sepaline cup, the apex rounded, contracted into an erect,

slender, color tail nearly 5 cm long, the lateral sepal 8 i transversely oblong lamina, |

mm long, 22 mm wide, with a shallow mentum, the obtuse apices contracted into slender, descending

tails m marked wi x

roun

mm wide, the epichile deflexed 45°,

ae the central lamella taller than the margins, with an incomplete accessory lamella on either side,

Plus a few short, margin: the hypochile oblong, 3 mm long, 2.3 mm wide, with erect, obtuse

k he 5 L 1 Pape 1 1 1

ie ts,

marginal angles, deeply cleft centrally, Was aru:

low, stout, semiterete, 4 mm long, with a stout foot nearly as long.

-ULUIIIIE o dit

LOMBIA: Without collection data, purchased from Tsubota as “Eva-6,’’ Eva Gardens, Pereira,

Colombia, cultivated at Orchideeénkwerkerij Venhui ij

(Holotype: MO), C. Luer illustr. 20123, : i ie aaens tc nee

This species is known only from Colombia without collection data. Although it

was first thought to be another of the multitude of hybrids that have been encoun-

tered recently, it has distinctive fea

tea eee ii Luer, sp. nov.

this wie or Roy Barrow of Royden Orchids, Great Missenden, England, who cultivates

Inter species generis Draculae Lue

berecto brevi, flore Parvo caudibus

silhpsel ag

r, planta parva, foliis angustissimis, pedunculo crasso

See a CrasSsSissimis. senalic d, rs . : . pil

‘“ 5 o

UL 4: +. ge ow .

& b MUSUILUL,

De VN een Taae

‘

ee.

Zig

fe)

SYSTEMATICS OF PLEUROTHALLIDINAE 99

Plant small, epiphytic, caespitose; roots coarse. Ramicauls erect, stout, 2.5-3 cm long, en-

closed by 2-3 loose, tubular sheaths. Leaf erect, thin inly coriaceous, carinate, very narrowly obo-

vate, subacute, 12-15 cm long, 1.3-1.5 cm wide, gradually narrowed below to the conduplicate,

subpetiolate base. Inflorescence a successively few-flowered raceme, borne by as ar suberect

to horizontal peduncle 3.5 cm long, from low mm long;

pedicel 8 mm long; ovary thick, 6 mm long di y white, dif Spotted

with ru soi red-brown, with a short dense, papillary cess within, the dorsal sepal broadly ovate, 12

mm long, 12 mm wide, p d, shallow, sepaline cup, the

ce tg contracted in a 2.5 mm broad, erect, dark red-brown tail ca. 33 mm long, the lateral sepals

connie 10 mm into a ‘broad, bifid lamina, ca. 18 mm long, 20 mm wide, with a shallow mentum, | the

tice a

cream, marked with ‘brown, cartilaginous, oblong, 3 mm teeta 2mm wide, the apex | bivalvate, te, densely

white, suf-

— with rose, spathulate, 8 mm long, 4.75 mm wide, th ith 5 a

, 4.75 mm wide, shallowly concave with pao margins, the concavity with 3 semana

ae lamellae plus a few small, incomplete ts, the hypochile oblong, 3 mm long, 3.25 mm

wide, with erect, round, marginal angles, cleft comnelll fe concave base hinged to the column-foot;

column light yellow, stout, semiterete, 5 mm long, with a stout foot equally long

PERU: without pimenia data, obtained from Manuel Arias, of Lima Peru, cultivated at Royden

Orchids, Great Missenden, Buckinghamshire, England, Jan. 2002, by R. Barrow 0/0 (Holotype: MO), C.

Luer illustr, 20083.

This species is apparently endemic in Peru, but without known locality. With

the thick tails it is most similar to Ecuadorian D. navarroérum. Dracula barrowii is

distinguished by the small habit with narrow leaves and a short, stout, suberect

peduncle that bears a small flower. The sepals are densely papillose-pubescent

within, and diffusely spotted with red-brown. The tails are :

The petals are small and verrucose between a short inner valve and a roun oe

valve. The epichile of the lip is ovate and shallowly concave with incomplete

lamellae radiating from the margins to an incomplete central irregular!

disrupted lesser keel on either side.

Dracula christineana Luer, sp. n for identi-

Ety.: Named for Mrs. Christine Cieosink of Lexington, KY, who submitted this species

fication. CPt eo snoribus. sepalis late

— haec D. ——— Luer & Dalstré affi : 4

Aictinowitnr

et lahell; £'

roots slender. Ramicauls slender,

ee Tes eat ie sts Leaf eect, sang coriaceous, carinate,

erect, 3-4 cm long, enclos ed by 2- 3 loose, tu sinha 4 cm wide, gradually

Narrowly obovate, acute, 10 snd seach few- poses

cronies below into the condupicae base. In Le ilons.agroatre ramicaul; flora’

raceme, borne by a horizo ound ii en section; sepals eee

bract, tubular ts long; penee - mm long; aye ie Sea a idle, ent within, the

uffused and spotted ws between ne. 16mm oie connate to the lateral

dorsal ba transversely ovate, transversely 0 obtuse, = mm long, ccmtracted into a slender, erect, pUr-

widely g Ott 2 dae white basally, 13 mm long, 13 mm

le-bl ‘+ } long, the mentum, the transverse apices

wide, ack ae a a forma a synsepal tee tO en at the pex, cartila-

contracted into tails similar to that of papillose between the laminae, the inner

ginous, oblong, 3 mm long, I 75 mm wide, the apex 28 alvate, white, suffused with rose canon 8 mm

lamina rounded, denticulate, the outer ~ 4.5 mm long, St mm wide, set concave with the

long, 5.5 mm wide, the epichile suborbicular, ther side of the sata

terior margin incurved and inflated, with numerous, i, with st, incurv incurved, unc inate, marginal angles,

midrib, the hypochile oblong, 4 mm long, 3.25 5 wide, white, stout, semiterete, 3.5

cleft central base concave, hinged to

mm long, with a stout foot 2 mm long. be eee nae

ECUADOR: Without locality, f gate

Lexington, KY, flowered in cultivation, May 1996, C- ae (Holotype:

100 ICONES PLEUROTHALLIDINARUM

This species was purchased from Dr. Eduardo Sanchez of Cuenca, Ecuador.

With little doubt, it had been collected in southern Ecuador. The small flowers with

yellow sepals marked with brown below the middle are reminiscent of variations of

D. erythrochaete, but the small, deeply concave epichile with numerous radiating

veins is distinctive. Except for the smaller size, the lip is similar to that of D. la-

fleurii. The sepals of the aeiiee are much larger and acuminate into tails, while the

sepals of D. christineana e d abruptly contracted into tails.

Dracula | deniseana Luer, Sp. nov.

hids, G Mi den, England

ies haec eh — Luer . Andreetia affinis, sed | pedunculo pi plusmir ere viore,

i e infra medium ferrugineis su Pp caudibus divergentibus

oribus diciesaiien

Di. + At } 3 +t

piphytic Ramicauls wr stout, unt cm

po enclosed = 2-3 loose, faclar s sheaths. Leaf erect, thinly coriaceous, carinate, very narrowly

te, ee , 13- 18 cm long, 1.5-1. 8 cm wide, gradually songs below to ike ctindagiteaks,

on ew me by a stout, erect to suberect

peduncle mye cm fee from low on the ramicanl: floral bract tubular, mm long; pedicel stout, 12 mm

long; ovary thick, ua mm site ne re mm thick, round in cross section; sepals rusty red-brown below the

middle in confluent’ i pu

ateral sepals connate 14 mm into a transversel yo blong lami » 20 mm long, 27 mm wide, with a

tae mentum i i il to that of the dorsal sepal;

petals cream, marked with brown, cartilaginous, oblong, 3 mm op 2 mm wide, the apex ee

er lamina acute, minutely denticulate, the oute

ensign rounded, slightly revolute; lip white, pete with Eg _pempers 11 mm long, 5.5 mm w ak

d, 8 mm long, 5.5 m shallowly concave with

denticulate margins, the concavity with 3 tall, Gauneioas aterstevey panies plus a few shorter,

incomplete eee. the hypochile oblong, 3 mm long, 2.75 mm wide, with erect, obtuse marginal

angles, vcleft centra foot; column light yellow, stout, semiter-

ete, 4 mm long, ay a stout foot nearly as long.

PERU: Hudnuco: \ ith , 2 lity, al alt 1700 m, obtained from Manue’ 1 Arias of Lima, Peru, cultivated at

y Y t Mi i ire, England, Jan. 2002, by R. Barrow RO9 (Holotype:

MO), C. Luer illustr. 20084.

This species is apparently endemic in Peru, but without known locality. It is

most similar to D. hirsuta which is known from both Ecuador and Peru.

Dracula deniseana is distinguished by long, narrow leaves and a short, stout,

suberect peduncle that bears a medium-sized flower. The sepals are densely papil-

pubescent within, with coalescing rusty red-brown spots below the middle, and

whitish above. The petals are very small and minutely verrucose between the

valves. The epichile of the lip is oblong-ovate and shallowly concave with incom-

plete lamellae radiating from the margins to an incomplete central keel and an

arly disrupted lesser keel on either side.

nigritella Luer, sp. nov.

Ety.: — the Latin nigritellus, “a little black one,” referring to the color of the sepals.

Species nec D. an Luer & Escobar similis, sed mea minoribus, petalis propor-

small to medium in size,

Plant 2

epiphytic, d Is

iides: exctk $ ons ioe Piphytic, densely panes: a “ ae Ramicauls

narrowly

closed by 2-3 loose, tubular y coriaceous,

cae linear-elli liptical, acute, 14-16 cm long = an nds petri base,

r OE ied - In sted, ivel

w-flowered ae RS Retaiee onges successively

wre sn by a d le ca. 7 cm long, from low on

the ramicaul; floral bract, tubular, “anv

g; pedicel 10 mm lianas ovary 6 mm long, round in

cross

section; sepals deep purple-brown-black, shortly pubescent within, the dorsal sepal broadly

SYSTEMATICS OF PLEUROTHALLIDINAE 101

ovate, obtuse, 11 mm long, 11 mm wide, connate to the lateral sepals for 4 mm to form a widely

gaping flower, the apex obtuse, contracted into a slender, erect, purple-black tail 2 cm long, the

lateral sepals 11 mm long, connate 8 mm to form a synsepal 17 mm wide, with a shallow mentam,

the apices obtuse, contracted into slender tails 2 cm long, similar to that of the dorsal sep:

cream, black at the apex, cartilaginous, oblong, 3.25 mm long, 2 mm wide, the apex bivalvate,

densely papillose between the laminae, the inner lamina rounded, denticulate, the outer

rounded, recurved; lip pink, spathulate, 6 mm long, 4.5 mm wide, the epichile sorbic. 45 5

mm long, 4.5 mm wide, shallowly concave with the margins not incurved, with about 4 tortuous

lamellae on either side of the minutely verrucose midrib, slightly verrucose externally, the hy-

pochile oblong, 2.5 mm long, 2.75 mm wide, with erect, obtuse, marginal angles, cleft sapere

the base concave, hinged to the column-foot; column yellow-white, stout, semiterete, 3.5 m

long, with a stout foot 2 mm long.

ECUADOR: Pichincha: near Mindo, collected by M. and F. Navarro, flowered in cultivation in

Quito, 9 Mar. 2001, C. Luer 19866 (Holotype: MO).

This species, recently discovered on the western slopes of Pichincha in northern

Ecuador, is superficially similar to D. anthracina, which has also been found recent-

ly in the same area. The deep purple-brown, nearly black sepals are smaller than

those of D. anthracina, and with shorter tails. The petals with rounded, black,

prickly apices are larger than the column. The shallowly concave epichile of the lip

contains about four tortuous lamellae on either side. The lamellae are as tall as the

non-incurved edges of the epichile.

Dracula sijmii Luer, sp. nov.

Ety.: Named for A.P. Sijm of Venhuizen, Holland, who ae this eaprae

—e haec D. houtteanae (Rchb.f. ) Luer affinis, "

tis, sepal

pichilio leviter

ih : a:

Plant medium in size, ovine caespitose; roots slender. Ramicauls stout, erect, 2.5-3 cm long,

enclosed by 2-3 loose, tubular sheaths. Leaf erect, thinly coriaceous, carinate e, RaNTOW why. obovate, Lan

14-18 cm long, 1.8-2 cm cia gradually narro low into th te base

successive, distantly few-flowered raceme by an ascending to to horizon tal peduncle 10-15 cm sine

ist won the ramicaul; floral bract tubular, il —_ long; pedicel 11 mm long; ovary purple-black, 5

black, glat externally, — purple within,

to the lateral

spar peal, the dorsal sepal transversely ‘obovate, 14mm Sa 16 mm wide, connate

sepals fo!

gaping 9 nea apex transverse, memes contracted

i 3 cm long, the lateral se sepals broadly ov hss pepe mr

lamina, 17 mm long, 16 mm wide, with a broad, ioiee o tum, btuse, contracted

into slender jer ary to that of the dorsal sepal; petals white, heavily marked with purple-brown,

cartilaginous, oblong, 3.5 mm long, 2.5 mm wide, the apex bivalvate, papillose between the laminae, the

inner lamina Ae sequins tthe outer lamina obtuse, slightly recurved, denticulate, lip white,

spathulate, 7 mm long, 4 mm wide, the epichile subcircular, shallowly concave, 4 mm long, 4 mm wide

ary lamella flanked on either side a lesser lamella and

incomplete ther lamellae, the hypochile oblong, 3 mm long, 2.5 mm wide, with erect, rounded,

=, angles, cleft centrally, with the margins of the cleft papillose, the base concave, hinged to the

column-foot; column yellowish white, stout, semiterete. te, 4 mm long, with a stout foot nearly as long.

ECUADOR: without collection data, obtained from José Portilla, Feb. 1997, cultivated in Venhuizen,

Holland by A.P. Sijm 200811 (Holotype: MO), C. Luer illustr.

This species collected in Ecuador is closely related to the variable Dracula

houtteana of Colombia. In the key to the species (Icones Pleurothallidinarum- 10),

it keys out to D. houteana. With narrowly obovate leaves, the two species are indis-

tinguishable vegetatively. The racemes of D. sijmii are distantly flowered; the

sepals are deep purple-black externally, gray-purple and sparsely spiculate within,

and deeply connate into a conical, sepaline cup or tube. The round epichile of the

lip is proportionately slightly wider with incomplete lamellae flanking a central

lamella. Both margins of the cleft of the hypochile have a pair of papillae.

r7 mm t

102 ICONES PLEUROTHALLIDINARUM

Dracula tsubotae Luer, sp. nov.

Ety.: Named for Shigenobu Tsubota of Eva Orchids, Pereira, Colombia, who cultivates this species.

pee haec D. lafleurii Luer & Dalstrém affinis, sed floribus minoribus, sepalis late obtusis, et

lakall

medium in size to large, epiphytic, densely caespitose; roots slender. Ramicauls slender,

erect, ~ cm long, enclosed by 2- 3 loose, tubular she eat ths. L eafe erec ct, sage coriaceous, carinate,

narrowly obovate, acute, 1 12-17 petiolate base. cm W: ide, gradu: ally

narrowed below into the -conduplicate base. Inflorescence a ae sacsaatocy. few- flowered raceme,

bo l7c pha from low on the ramicaul; floral bracts,

oblique, 10 mm long; pedicel 15-171 mm long; ovary 5 mm | ong, round in wea section; sepals greenish

otted the middle, densely short-pubescent within,

ne broadly ovate, obtuse, 18 mm long, 3 20 mim W wide, c conna ~ > dai lateral sepals for 6 mm

form a widely spread flower, e-black tail 7 cm long, ie

lateral sepals broadly ovate, 18 mm long, connate 14 mm into a oe ica 30 mm vie: with a

shallow mentum, the apices obtuse, contracted into slender tails also 7 cm long; petals cream, dark

brown at athe aaa oblong, 3.5 mm long, 2 mm wide, the apex bivalvate, ae sees

between the laminae, the inner lamina rounded, denticulate, the outer lamina rounded; lip white,

ate, 9.5 mm ar, 5 mm long

arg

c base concave, hinged to the column-foot; column yellow-white, stout, semiterete, 4.5

mm long, with a stout foot 4 mm long.

COLOMBIA: v itk ion dat hased fi Tsubota of Orchids in 1997, ae ar si

Great Mi ngland, Feb. 2002, R. Barrow RO8 (Holotype: MO). Nari

Samaniego, collected in 1995 fa iL Aguirre, cultivated e saad a del Valle, Cali, Colombia, psa

ered Mar. 2002, J.C. Uribe s.n. (MO), C. Luer illustr. 201

This species is apparently endemic in southernmost Colombia where it was

discovered by José Luis Aguirre. Several clones have become known as “Dracula

tsubotae” in collections, apparently because of its having been obtained from

Shigenobu Tsubota’s Eva Orchids in Pereira, Colonbia.

Although vegetatively rather large, the widely spread flower of this species is

medium in size. The sepals are broadly ovate, obtuse, densely pubescent, and long-

tailed. With the basic whitish or greenish color densely spotted in dark purple-

brown below the middle, the flower i is reminiscent of D. vlad-tepes. The subspheri-

cal, deeply concave epichile of the lip with multiple, radiating veins is similar to

that of the Ecuadorian D. lafleurii, and also the presently described D. christineana.

portillae Luer & Hirtz, sp. nov.

~~ Named far I.

hi i E in Gual Ecuador.

h 3 i—

SE D. hirsutissi (C.Sct inf.) I ffinic cod x 1 lek : #1 Soo saas

sile synsepali » ae as 2 v

J et Iahell 4 ’ ci ies ee

t — P igi

Plant small epiphytic, caespitose; roots slender icaul

. s suberect, stout, 3-5.5 cm long, enclosed

= s spotted, meting — tubular, imbricating sheaths. Leaf green, suberect to horizontal, erect

Famucaul, thickly 1.

ne cm wide, tose broadly cuneate 1 genmeee i both surfaces, elliptical, obtuse obtuse, 3-4.5 th sont 5

tubular, long-pul et peduncle ca 1 mm long, with a basal sheath at the base of the leaf; floral bract

cence, the sepal purple spots, covered scahacmly. by a long ~ A pase

long, 3.5 mm wide, ovate, acute, acuminate, 11 mm long including the slender, a

essentially free from the lateral sepals, the late sepals connate hs the i sare a

” spcaeathade Pp 12 mm long, 6 mm wide une seamed. with the margins

f bel w the

€, oblique, 9 mm long, 2.8 mm wide, the pee aces send 2 pu ple s spots, obovate | ef 3mm

ate-pandurate or pyriform,

~pubescent, 5 pedicel p

long steaks anslucent dull yellow with dull

lip yellow.

Ne i 5 trilobed, obo

3mm wide, the 7 rounded, the margins long-fimbriate ean the middle, <n late -

SYSTEMATICS OF PLEUROTHALLIDINAE 103

lobes on the basal quarter, thin, ease uncinate, ca. 1 mm long, the disc with a low pair of converging

e claw w with an erect, obtuse callus, the base truncate with minute

eRe lobules, fi ly hinged to th foot; lu lender, semiterete oe the apex

ad Co

1 . thirl pedestal 28. ned cancels the

E a

eee Mores eee sen Chitin alt. 1400 m, cultivated at root Gualaceo, 5

This species, closely related to sympatric D. hirsutissima, has recently been

discovered in southeastern Ecuador. The leaves of both species are thickly fleshy

and covered by a long pubescence on both sides. The peduncle of D. portillae is

very short, causing the slightly smaller flower to rest in the sulcus at the base of the

leaf. The dorsal sepal and petals of D. portillae are similar to those of D. hirsutis-

sima, but they lack the clavate tips of the attenuated apices. The margins of the

boat-like synsepal meet a few millimeters below the apex to create an apical aper-

ture separate from the main opening. The lip is long-fimbriate above the middle,

but similar, uncinate, lateral lobes are present on the lower portion.

Dresslerella sijmiana Luer, sp. no

Ety.: Named for A.P. Sijm of Soe ‘Holland, who cultivated this species.

Species haec, D. hirsutissimae (C. Schweinf. ) Luer affinis, sed pee breviore, foli folio | late ellipti-

co ciliato,

Lal xf

Ramicauls erect, slender, 2-2.5 cm sophia enclosed

by 3-4 snes: ena pee tubular, mfr sheaths. Leaf green, Se ce be-

neath, erect, thickly fleshy, densely ciliate, also ciliate along the midvein on the igen ly ellipti-

broad!

cal to suborbicular, obtuse, 1-3¢ m long, 1.5 em wide, the base y cuneate, sessile. Inflorescence a

in a fas ach borne e by a suberect, pubescent Pesncl 6-7 mm long,

ith a basal sheath at the b t, 4-5 mm long; pedicel 7-

8 mm long; ovary densely pees 2 mm long; sepals dull yellow with irregular, = purple spots,

ovate, acuminate, 9mm — ——e

the slender apex, 3.5 mm wide, essentially free fi he lateral epals, th f

the apex into a deeply concave, obtuse, id s Ba tae aid cacapontet

year ap purple, obovate below the middle, » oblique, ‘As = long, 2.5 mm wide, the _ acute,

ith le, trilobed, obovate-pandu-

rate, 5 mm lo ial ng, 3mm 1 wide, the isthmus | mm wide, the ests lobe oblong, transversely ian

prominent veined, with the margins erose, the lateral lobes on the lower half, erec t, obtuse, with a

minute, process, the disc between the lateral lobes tricarinate, wi point at

inute marginal lobules, firmly hinged to the

dev

, uncinate,

the base of the middle carina, the base truncate with m

veloped into a

coterie TO0r; ro slender, ecmamerete, 2.5 mm long, the apex bidentate, the base

thick, p

ovary

ECUADOR: Without locality, obtained from José ot cultivated in 2000 in Hem, Venhuizen, Hol-

land, by A.P. Sijm 200424 (Holotype: MO); C. Luer, i 19402.

This species, closely related to sympatric D. hirsutissima, has recently been

discovered in Ecuador. It is characterized by suborbicular, densely ciliate leaves,

and a fascicle of short-pedunculate flowers. The dorsal sepal with a narrowly

acuminate apex is shorter than the synsepal. The petals are oblique with acuminate

tips. The lip is pandurate with a broadly expanded, erose, apical lobe, separated

from obtuse lateral lobes by a very narrow isthmus.

Lepanthopsis calva Dod ex Luer, sp. no

Ety.: From the Latin Calvus, “*bald,”’ referring to ms glabrous sepals.

Species haec L. —— Dod s similis, - ramicauliorum — crass

rotundis abrupte petio

e ciliatis, foliis ad basim

: roots slender. Ramicauls “or stout, 5-6 cm long,

Plant medium in size, epiphytic i

enclosed by 8 coarsely Jr ent, at, lepentiform s sheaths with anes, a te ostia. Leaf erect,

thickly coriaceous, conve the margins mt tely denticulate 1, obtuse oe a mm long, 11 mm

wide, the base t cun dintoa aa oY mm long. Inflorescence a sub-

104 ICONES PLEUROTHALLIDINARUM

dense, strict, distich ith ly several-flowered raceme, up to 35

mm long includin the capillary sr 17 mm long, from near the apex of the ramicaul; floral bracts

1.25 mm a ede 1.25 mm long; ovary costate, 0.6 mm long; color _ flowers not stated; sepals

‘ ovate, subacute, glabrous, the dorsal se sepal 2.5 mm long, 1.5 mm wide, incom-

pletely Salen fr the lat free to near - base, oe 2.5mm

long, 1. wide, i let ed; petals ovate, acute, slightly acuminate, 1.75 mm long, 0.75

mm wide, I-veined, glabrous; lina ovate-cordate, thick, 1 Len mm an fon ng, h 4 mm wide, the se rounded

yg , the disc with a ag

gl t the b Jumn 0.5 mm long, 0.5 mm wide, the anth sos I, the stigma bilobed, ap

HAITI: Massif de la Hotte, Les Cayes, Formond, alt. ges m, 1 Jan. 1983, flowered in cultivation 17

Oct. 1983, D.D. Dod s.n, (Holotype: MO), C. Luer illustr. 12294.

This species is similar to and sympatric with L. micheleae in one known area on

the Massif de la Hotte in southern Haiti. It differs from L. micheleae in stout,

coarsely pubescent-ciliate, lepanthiform sheaths; a thicker, convex leaf rounded at

the base and abruptly short-petiolate; and similar flowers but with glabrous sepals

and petals. The lips of both species are margined toward the apex, and with basal

lobes that encircle the column.

—— i Borperasn Dod ex Luer, sp. nov.

the Latin er. rooned purple,” referring to the flowers.

haec 1 th. fili i

vexis sepalis lat, th ee 4: tie Lee.

Cit,

sed floribus levit majoribus, sepalis con-

Plant small but average in size for tt lender. Ramicauls erect,

stout, 1.5-3 cm long, enclosed by 3-7 blackish “ates oo. sheaths on long-ciliate ostia. Leaf

erect, the margins smooth, elliptical, obtuse to rounded at the apex, 15-

20 mm long including a petiole 1 mm long, 8-9 mm wide, the base cuneate pare the peti Inflores-

cence a congested, distichous, simultaneous] ly several-flowered race:

de,

toe 3 oe Toeind lip colored as the sepals, circular-cordate, rounded at the apex, 1 mm

shall setae a lumn, the disc featureless except for a

sigma Gnen Sesion athe base; column stout, broad, 0.5 mm long and wide, the anther apical, the

HAITI: Massif de la H 22;

: eRe, = . Grog Formond, alt. 1100 m, Apr. 1992, D. Dod 2260 (Holotype:?;

This species i 1S known only from the Formond area of the Massif de la Hotte in

al leaves are smooth on the margins. A short peduncle

bears the tip of the crowded raceme of back-to-back flowers beyond the leaf. The

broadly Ovate sepals and suborbicular petals are deep purple and convex. The lip is

icular with short, basal angles embracing the column.

Lepanthopsis rinkei rinkei Luer, sp. no

Ety.: Named for Bryon Rinke of Wichita, KS, SS erepngg

Planta mediocris S Caespitosa inflorescenti

ticho parvifloro sepalis obtu

connatis leviter

. mo congestissimo dis-

— lateralibus elliptico ad basim

ibus, petalis Ovatis, labello : or suborbic

Plant medium to large for the

penis Stout, 15-6 cm cm long, enclosed by 5-6 lately uy epiphytic, caespitose; roots — a se

nauk 2-3 mm rt ar the apex, 20-35 mm long, 13-20 mm wide: the base Paes into a

57 4 congested, simultaneously several-flowered raceme in 2 opposite

ranks, bracts mes tong: pea Hacc Petutcle 20-23 mm on, from near the apex ofthe ramicaul;

g; Sepals translucent rose-purple,

SYSTEMATICS OF PLEUROTHALLIDINAE 105

white at the base, Pa pub i the apices, the dorsal | broadly le, obtuse, 2 m 2 mm

long, 1.75 mm wide, 3-ve he | 1 Is, the lateral M

ee ote oblique, obtuse, uae i 2.2 mm long, 1 mm wide, connate less than 0.5 mm, am, 1-

veined; petals dark purple, ape elliptical es obtuse, 0.6mm og, 0: 0.3 mm wide, faintly 1-

veined, if at all; lip dark purple, glabrous, broadly ovate, rounded at the apex, slightly concave, 1.2 mm

lon ng, 1. 5 mm wide, the base es bilobed, be asa lo bes Ss rounded, rir the column, the disc

fas

PANAMA?: without d fi dy's Orchids, ded ln om

Maduro in Panama, cultivated in Wichita, Kansas, as ranci Rinke,

fication, Sept. 2001, C. Luer 20048 (Holotype: Mi

Although obtained from Panama, I tae that this species is Andean in origin.

No other collections are known. All features of this species, except for the size, are

found in other species of the genus, but not in the present combination.

Vegetatively large for the genus, the leaf and ramicaul are similar to but larger

than those of L. floripecten. The leaf of L. rinkei is broader, being broader than the

leaf of any known species of the genus. Common to a few species (L. abbreviata,

L. atrosetifera, and L. ubangii), a congested, two-ranked raceme does not exceed

the leaf. The obtuse dorsal sepal; slightly diverging, oblong, lateral sepals; minute

petals; and broadly rounded, large lip are most similar to those of L. glandulifera

and L. melanantha. A proportionately large, rounded lip is seen in many species,

but the disc of the lip of L. rinkei is featureless.

Lepanthopsis woodsiana Dod ex Luer, sp. nov.

Ety.: Named for

eee ee ee Ca

Species haec L. stellaris Dod affinis, sed t Se ere ieiili

flexuoso, et labell borbiculato levi 8 r

differt.

. " ively fleshy. Ramicauls

Plant small but large for the genus, epiphytic F Pay Gt NS SEAR

rect, slender, 1. = cm long, enclosed by 3-7 sparsely long-pubes«

ciliate, dilated ostia. Leaf erect, coriaceous, lightly pusticulate, with

elliptical,

the base cuneate into the petiole. Infl

long, from near the

flowers open simultaneously, up to 40 mm_ long including the peduncle 10-20 abe g, fi

pex of the ramicaul; aby eat erie eee rrr , glabrous, sub-

yellowish, suffused with rosy brown, especially along the veins, sos ie uas nse 3.25 mm wide,

orl obtuse, “igh cl » ti “ 1: oneal oblique, subacute, 5.5 mm long,

: exe obli

3 mm wide, 2-veined an itor aN connate 1.5 mm; petals colored as the sepals, ov o _

acute, 3. He Ne ong, 1 ie +

lar-glandular, lr ovtorate frei 2 75 mm Tong, 2.25 mm wide, the base with lobes | that su sarod

column, “a disc featu

mm long, 1.5 sails nn aie ager pee

263 (Hol 2;

HAITI: Massif de la Hotte, Les Cayes, Formond, alt. 1100 m, Apr. 1992, D. Ded 2263 (Homtype-

Isotype: MO), C. Luer illustr. 16535. tilles.

Vegetatively and florally, this species is the largest of me a igd site of a

The flowers are nearly one centimeter across from tip o flowers that reaches

lateral sepal. The raceme is a congested array of oe slightly acuminate

Or surpasses the tip of the leaf. The sepals are ‘am arative large and also

with the tips of the laterals diverging. The Sec lalics that encircle the

ee acuminate. The lip is rounded with elongated

columnn

106 ICONES PLEUROTHALLIDINARUM

Platystele portillae Luer, sp. nov.

Ety.: Named for José Portilla of Gual , Ecuador, re seorssnn and cultivated this species.

par = be ao p E ] minusve duplo lon-

r . .

gioribus

h Hy Ly ter flavic lahell 7 ] + di

2 = a i

Plant small, epiphytic, sn roots slender, densely fasciculate. Ramicauls slender, erect, 4-6

0-3

mm long, with a close, thin, tubular sheath. Leaf erect, cori oriaceous, narrowly obovate, obtuse, 20-30 mm

long, 4.5-5.5 mm wide, gradually caachie below i 5 -10 mm | lon ong. Inflorescence

re , flexuou: ny successively few-fl d i

1 nuh hy ae ral bracts 1 mm | long; pedice

oe vided de 0.4 mm long; sepals long-ciliate, yellow, the dorsal sepal elliptical, obtuse to are at in

e, 1.5 mm long, 1 mm wide, L-vomed, the lateral sepals free, reflexed, elliptical-obovate,

cob 1S. Se mm long, 1 mm wide, 1-veined; petals long-c yew ce elliptical-subfalcate, the apex

acute, acuminate, incurved, 1.5 mm long, 0. 4 mm wide, 1-ve p purple, fleshy, ovate, ces

microscopically pees 1.5 mm long, 1 mm wide eae with a minute glenion, trunca'

oe attach base of the column; column clavellate, 0.4 mm long, the psoas ah bi lobed

stigma apical, the oe rudimentary.

ECUADOR: Zamora-Chinchipe: Loyola, east of Valladolid, alt. 1800 ode cultivated at Ecuagenera,

Gualaceo, 12 Apr. 2001, A. Hirtz 760] (Holotype: MO), C. Luer illustr. 2

This little species from southern Ecuador is similar in habit to many species of

the genus. A hairlike, flexuous raceme that bears a tiny flower successively, sur-

passes the little, narrowly obovate leaves. The sepals are obtuse and ciliate with the

laterals reflexed. The petals are narrower and incurved. The ovate lip, as large as

the sepals, protrudes.

Pleurothallis adrianae Luer & Sijm, sp. n

Ety.: Named for Adriana, wife of A.P. Sijm of Saale Holland, who collected this species.

Species haec P. obovatae (Lindl.) Lindl. . affinis, sed habitu multimajore, sepalis acutis, petalis

acuminatis.

VeIOW

_ Plant large, epiphytic, » caespitose; roots slender. Ramicauls erect, stout, 20-28 cm long, with a

lpia subacute, petiolate, 12-15 cm long, 3-4 cm wide, cuneate bel,

nce a fascicle of several » Secund, simultaneously aCe racemes, 2.5-3

cm cm long — the peduncle 0. 5 cm long, subtended by a spathe 3-4 mm long, from an annulus 2-3

mm long; pedicels 1.5 mm long; ovary 1 mm

long: sep sepals yellow-green, tee ob nt ate- -triangular, acute, the dorsal sepal 5 mm long, 1.75 mm wide,

low elow the column-foot, 4.5 mm

long, | 73 mm wide, 3-vein ed; petals yellow w-green, — ovate, oblong below the acuminate apical

third, acute, 3.75 mm long, 1 mm wide, with ically erose

le, oblon ye

ie seeps les, obtuse the ds shallowly slate th die tess sebescaneiel i u-

a hac above the middle,

long, the foot thick, 1 St mm “we the anther and the ventral. -yconeceitetnty cod es

ECUADOR: Without locality, collected ca. 1994, cultivated in Venh 5

tion 7 Aug. 2001, A.P. Sijm 210807 (Holotype: MO), C. Luer illustr, as ik ae tra aaea er gi

This species is the largest vegetatively in the subgenus Aciminatia, but closely

related to P. citrina, P. angustilabia and P. obovata. It is distinguished from them

by the much larger habit with a stout ramicaul over 20 centimeters tall and a large,

aeDaG — Several short, secund, few-flowered racemes are produced simulta-

neously. The sepals are yellow-green and acute, and the

tubing eae petals are acuminate. The

SYSTEMATICS OF PLEUROTHALLIDINAE 107

Pleurothallis cauda-hirundinis Luer & Portilla, sp. n

: Ety.: From the Latin cauda-hirundinis, “the tail of a sees in allusion to the lobes of the

lip. Hirundo, the genus of swallows

Species haec P. cauda- it Luer & Hirtz affinis, sed labelli lobis anguste triangularibus erectis et

callo minute toga

i fasciculate. Ramicauls slender,

erect, round in cross section, 28 ¢ cm eae! with close, capees sheaths above the base. Leaf erect, coria-

ceous, ovate, acute, I 5- 25 cm long, 0.8-1 cm wide, cuneate, sessile. Inflorescence a fascicle

of solit tary, the 3 mm long,

3-5 mm long; floral bract 3 mm long; pedicel 3-4 mm long; ovary 3mm long; sepals g labrous, pale

yellow with brown spots, the dorsal sepal ovate, acute, deeply concav: ein — lower half, 6 mm long, 3. :

ide unexpanded, 3-veined, th

concave srauuuk: 6 mm long, 5 mm = pave pc 4-veined; cae giabrows, age yellow wh

brown spots, obliquely sub-falcate, the acuminate, with a subacute angle lower m

gin, 5 mm long, 2.5 mm wide, eoat lip p we catiau bilobed, = lobes porsche erect, petal

around the column, 3 mm long, 1.2 mm wide, the apices acute, the body thick, | mm long, truncate cate, with

a minute, 2-pronged callus at the middle, the: base firml y ak above the base of the column; column

cylindrical, 1 mm long, th 8

ECUADOR: Carchi: Palén, west of Chical, alt. 1000 m, cultivated at Ecuagenera, Gualaceo, 5 Mar,

2001, C. Luer 19675 (Holotype: MO).

This species of northwesternmost Ecuador is related to P. cauda-phocae from

nearby. Although basically similar, P. cauda-hirundinis differs markedly in some

respects. It is characterized by ovate, acute leaves and slender ramicauls. The

solitary flowers are short-pedicellate, and borne in a fascicle at the base of the leaf.

The dorsal sepal and synsepal are concave and acuminate, the petals are large and

falcate with a lower marginal angle; the lip bears a minute, two- pronged callus,

while triangular lateral lobes curve upward to surround the cylindrical, essentially

footless column.

Pleurothallis cypelligera Luer & Hirtz, sp. n

.: From the Latin cypelliger, henies a beaker, or ae ’ referring to the lip.

li mhbhifarmibnuc

Inter apoers subgeneris Ancipitiae habitu mediocris, , sepalo dorsali : synsepalique cy

b Be eee

o —J

Plant medium in size, epiphytic, caespitose; roots slender. a — erect, compressed

ee m long, 2 mm d eep at at the leaf, with a tubular w the middle and 2-3

sheaths at the base. ie cm eo 1.5-1.8 cm tees

e base ie rounded to subcordate. Inflorescence a fascicle of pobril successive flowers

mm long, at the base of the leaf; peduncle slender, suberect 5-7 mm vie

ng; sepals alee white,

sie conaceous ding, ova

from a reclining spathe 3

floral bract thin, ae 4 mm long; pedicel 5-8 mm long; ovary 2-3 mm lo

the dorsal sepal ovate, narrowly obtuse, 7 mm long, pedigree

completely connate into a cymbiform — 7 mm long, 4

suffused with rose, oblong, acute, 7 mm long, mm wide, 1-veined; lip suffused with rose, subspher-

ical, obtuse, concave e, 3.5 mm long, 2.5 mm wide, the apex broadly rounded with the margin thickened,

with the —— margins minutely irregular with angles near the middle, the disc within with a shallow

depression base broad, subtruncate, hinged to the base of the column |

semiterete, 3. 5 mm long, the anther and stigma apical, with an obsolescent mane

orona-Santiago: w ff Macas, road toward Guamote, alt. 2300 m, C. Luer, J. Luer

& A. Hirtz 1 19625 bearer MO); . above Limén, alt. ca. 1500 m, cultivated at Ecuagenera, Gualaceo, 7

Mar, 2001, C. (MO).

This species, from southea

members of subgenus Ancipitia by concave,

suffused with rose; oblong petals; and a concave, su

irregular lateral margins.

stern Ecuador, is distinguished from the other

greenish white sepals and petals and lip

borbicular lip with minutely

108 ICONES PLEUROTHALLIDINARUM

rostratissima Luer & saci Sp. nov.

Ety.: From the Latin rostratissimus, “ th f th

Species haec P. trichostomae Luer Ss sed fois late ellipticis hues petites, ilbada

Plant small, epiphytic, densely caespitose; roots slender. Ramicauls stout, erect, 0.5-1.5 c

enclosed by eaf erect, coriaceous, broadly cpt

ci 25-3 bio long including a petiole 0 3 cm Pash 1. =o 1.8 cm — — contracted below

the petiole. I disticho! xuous, several-

» up g jing the peduncl "em long, borne below the ‘apex of the rami-

thin above the middle, glabro lly, the dorsal sepal oblong, obtuse, 7 mm lo ong, 2.25 mm wide,

3-veined, the lateral sepal the apiculate tips i into a sang concave, obovoid to pyriform,

obtuse synsepal, 7 mm long, 3 mm wide unexpanded, 3 mm deep, 6 veined; petals spa thulate, rounded at

the apex, unguiculate, 2 mm long, 1.6 mm wide, 3-veined, callous and minutel along the

midvein e xternall ys lip oblong, unguiculate, 2. 6 mm long, 1 mm wide, acute at the apex with a minutely

obdtusel » the disc with a pair of longitudinal carinae and a cen-

tral, al, shallowly ch channeled carina, itha 0.5 mm long, truncate,

te-fimbriate, 4 mm long including a long-aristate tip

. 5mm long, the foot 1.5 mm long with a mare patch of dense pubescence, the anther, rostellum and

stigma ventral

ECUADO) I] ta. cultivated il-

opts AOR wit ae at Ecuagenera, Gualaceo, Ecuador, Sept. 2000, J. Portil

This species is known from a single collection. Among the members of subge-

nus Effusia, this species is distinguished by the dense cluster of overlapping, broad-

ly elliptical, abruptly short-petiolate leaves; a loose, flexuous, successively flowered

raceme; sepals long-villous within; a deeply inflated, obovoid synsepal; spathulate

petals; and an unguiculate ep that is tricarinate and with a verrucose callus at both

the apex and above the claw

Sextonii Luer, sp. no

Ety.: Named for Dr. Laurence ry eat ef Batavia, IL, who cultivates this species.

Species haec P — Luer & Escobar similis, sed foliis ad basim cuneatis, floribus majoribus et

labello cordato integro differt

lon Plant large, phan — repent-caespitose; roots slender. Ramicauls stout, erect, 11-15 cm

8, with a short, close y fitting, tubular sheath below the middle, and another 1-2 sheaths at the base.

| coriaceous, clic ova, ingore: , acuminate, 9-12 cm long, 4-5 cm big the base cuneate,

: a loose, distichous a. 15 cm long includ-

31g ine Peduncle ca. 6 em long, subtended by arec lining spathe 1.5 cm long; floral bracts thin, tubular, 3

~> mm long; pedicels 3-4.5 mm long; sad 3-3.5 mm long; sepals translucent purple, glabrous, the

d ve, 12 mm long, 5 mm wide, 3-veined, the lateral

preety haga liptia-oblong, subacute synsepal, -eptonr te ong, 5 mm wide, 4-veined; petals

—

0mm!

ones acute, convex, entire, 2.5 mm long, 2.25 mm wide, the =

: : : column-foot; col terete, 2.5 mm long, the an

apical, the stigma Msc lary umn semitere g

COLOMBIA: without collection data, obtained from cultivated

L i Vv ‘olombi tivated by

This Species has affinities with ona members of Pleurothallis subsection

Pe Ee re, cordate lip. From P. dentipetala Rolfe ex Ames, it is distinguished by

ce larger Soothe ina oe longer than the leaf. From P. gomezii Luer &

erose-fimbriate hed by a raceme longer than the leaf; minutely but coarsely

os and 4 cordate lip. From P. rowleei Ames, it is distinguished

SYSTEMATICS OF PLEUROTHALLIDINAE 109

by a larger habit; erose-fimbriate petals; and a cordate lip. From P. longipedicellata

Ames & C.Schweinf., it is distinguished by twice larger flowers; short cellate

flowers in a raceme longer than the leaf; and erose-fimbriate petals.

Pleurothallis sijmii Luer, sp. no

Ety.: Named in honor of A.P. Sijm, eshte the ane en again ~ sepa

Species haec P yp hyll, H 1B K. affinis, 1 I i gi

} = 2 tr. Aitteart

4+ lalall ; =)

linearis,

Plant large, citi, qeuperk roots coarse. Ramicaals stout, erect, more than 15 cm long,

enclosed by a tubular Leaf erect, thickly

coriaceous, = lpia sal 15 cm 1 long, 4.5 cm wide, cuneate into the base. Enteric

suberect to horizontal, loose, secund, simultaneously several-flow raceme up to 20 cm long includ-

ing the esos ca. iO er cm aiiee. from a spathe 1.5 cm long, a at ie hese of the leaf; floral bracts thin,

pos 5 mm long; pedicels 7 ] with purpl

e the base, ene the dorsal sepal narrowly ovate, concave, acute, acuminate, 30 mm long, 8 mm

wide expanded, 3-veined f ynsepal g

panded, 6 | th | third, acute, 12-13

mm wide ex ned; petals fi mottled

ee 3-ve contracted in! Baa cisG ual 15 we ing, with small verrucae.

en 29 mm; lip purple, sig 14 or long, 4.5 mm wide, concave e below

les, elliptical above the middle, minutely spiculate-verrucose, with

ate, ag op apex obtuse, shortly acuminate, the disc smooth below the middle,

the base broadly truncate, attached to the aa gy aegis column terete, 9 mm long, the anther subapical

the rostellum and stigma ventral, the foot thick, 3 mm lon

PERU: Hudnuco: Nov. 1998, A.P. Sijm 200425 (Holotype: MO), C. Luer illustr. 19408.

This large, handsome species of subgenus Elongatia is apparently endemic in

northern Peru. It is characterized by a rigid, thickly coriaceous, elliptical, sessile

leaf that is surpassed by a loose raceme of large, showy flowers. The acuminate

sepals are mottled with purple. From an elliptical blade the apex of the petal is

narrowly terete and with small warts. The oblong lip is elliptical above the middle

with a minutely fringed margin, and below the middle the margins are smooth, erect

and broadly rounded.

— portillae Luer, sp. n

Named for José we of ae, Ecuador, in whose smear this species is cultivated.

Ss haec R. brachypodi Rchb f. ffini ynsef ; I fora hc (capa

= spe oc

r r

t medium in size, epiphytic, —. roots slender. Ramicauls erect, 4-6 cm long, —.

by 6 loose, iia imbricating sheaths, with only the lower rp minutely dotted. Leaf erec

suffused with purple be neath, coriaceous, elliptical, obtuse, 4-4.5 c

roadly cuneate into a twisted petiole ca. 2 mm long. ot

duced wkealeely) ina heodake behind the leaf; peduncle slender, 4-4.5 cm long; floral bract tubular, 5

mm long; pedicel stout, 2- 3mm — with a slender filament San mm long; ovary 4 mm long, sepals

ous, t, light ye nba ty in red, narrowly ovate in the lower

half, attenuated above with the nae See orm wide, 5-veined, the lateral sepals

connate to near the apex into a concave, nc shoaly » bifid synsepa. light yellow with red

spots, 19 mm Te bg mm wide, ee the api ces obtuse, free ie eg - epee

narrow! the base

third a Haro the apex vation 12 mm n long, 1.75 mm wide above the “pe lip yellow with purple

, pandurate, 9 mm long, the epichile cuneate-truncat i mm wide, 15 mm wide at

the isthmus, the hypochile oblong-subquadrate, 4 mm sige 4 mm wide, peti with minute, hair-like

lobes from the middle of the sides, the base subtrun e, connected to the column-foot bya short, thick

neck; column greenish white, slender, clavate, 5 mm ie. the base pedestal-like with a pair of small,

inconspicuous, pubescent calli.

110 ICONES PLEUROTHALLIDINARUM

ECUADOR: Morona Santiago: near Cuigiiinda, alt. ca. 2500 m, cultivated at Ecuagenera, Gualaceo, 5

Mar. 2001, C. Luer 19663 (Holotype: MO).

This species, known so far by only the original collection in southeastern

Ecuador, is characterized by the medium-sized habit with elliptical leaves that are

purple beneath. The flowers, about average in size for the genus, are borne near the

tips of the leaves. The red-spotted synsepal is broadly concave, more or less in-

curved near the middle with the broad apex shortly bifid. The petals are minutely

fringed toward the base. The panduriform lip is most distinctive with a cuneate,

spiculate epichile; a narrow isthmus; and a proportionately large, concave hypochile

with extremely slender, short lobules.

Scaphosepalum bicristatum Luer, sp. nov.

Ety. : From the Latin bicristatuc referri on thie wok p.

Planta parva, foliis late ellipticis longipetiolatis, elongato quam folio multilongiore,

pedunculo laevi, sepali mediani cauda incrassato, petalis oblongis apiculatis unguiculatis, et labello

iaioibar,

+ fal, 83.

Plant small, epiphytic, caespitose; roots slender. Ramicauls erect, proportionately stout, 8-10 mm

long, enclosed by a tubular sheath and anot! he base. erect, coriaceous, long-petiolate, 2.5-3

mm long including a petiole 9-12 mm long, the blade elliptical, obtuse, 10-12 mm wide, cuneate below

into the petiole. Inflorescence a loose, distichous, flexuous, flexible ively many-flowered

raceme up to 15 cm long or longer, including the slender, smooth peduncle, 12-13 cm long, from low on

the floral bracts 3 mm long: pedicels 3-4 mm long; ovary smooth, 3 mm long; sepals yellow

with darker veins, glabrous, carinate, the sepal ovate, concave, thickened at the narrowly rounded

apex, 6 mm long, 3 mm wide, barely connate to the lateral sepals, the lateral sepals minutely ciliate,

’ to an ovoid, scaphoid, carinate lamin ded, the apical

third of each lateral sepal occupied by a triangular cushion with indistinct margins, 2 mm long, 1 mm

wide, Spices contracted into acute tails 1 mm long; petals translucent, oblong, oblique with

the labellar margin broadly rounded, 3.5 mm long, 1.75 mm wide, the apex shortly acuminate, apiculate,

mar.

ly serrulate, the disc with a pair of

; Slender, semiterete, arcuate, winged above the middle with a crest at the

apex, 3 mm long, the foot thick, 2mm long.

Postings aes bee cultivated by Ecuagenera, Gualaceo, Apr. 2001, A. Hirtz 7629

This Species is characterized by the small habit with spathulate leaves, and a

ively lengthening, lax, flexuous, successively flowering raceme. The muc

longer peduncle is smooth, but the rachis is microscopically subverrucose. The

median sepal is ovate and concave with a thick apex. The synsepal is deeply

sora and boat-shaped with tails only one millimeter long. The lateral margins

Margin, the apex is apiculate, and the base is unguiculate. The lip is arcuate with

the Sides thin and dilated, the apex is round and minutely serrulate, and the disc has

a pair of tall, fringed lamellae.

Scaphosepalum merinoi Luer, sp. nov.

Ety.: Hanes for Gilberto Merino of Gualaceo, Ecuador, collector of this species.

Species haec S. sibberoso (Rchb.f.) Rolfe

labelli cristis majoribus differt.

non elongatis, et affinis, sed sepalo impari breviore, sepalorum callis

oa a nie ae Caespitose; roots slender. Ramicauls erect, slender, 1.5-2 cm long,

long including a Petiole 3-3 tres, ratenpirhatlon ype Leaf erect, coriaceous, long-petiolate, 8-10 cm

below into the slender petiol tech , the blade elliptical, acute, 1.2-1.7 cm wide, narrowly cuneate

Faceme borne by a sn Weenece a nce a loose to distichous, successively several-flowered

icaul, floral t 4mm | * m0fe or less horizontal peduncle, ca. 12 cm long, from low on the

6-7 mm long; ovary verrucose, 5 mm long: translucent

oF AS

SYSTEMATICS OF PLEUROTHALLIDINAE 111

| es ~r 1 mn. . be tT My bd

white purple strip spots, y : reagan the dorsal 1 ovate here concave,

acute, narrowed with t Z 1 i twi hick, terete tai th 18 mm,

5 mm wide, barely he lateral t the base, the lat it

CORERYE, carinate seas 10 mm jones § 8 mm wide, +h . t halt ry = = , } ao i J

= r iil =

, 5mm long, 3 mm w brupt! t d into slender, white

oo 22-25 n mm | 2; BP tal. } _ ovate, , oblique, poy are se eas 4.5 mm long, 2.5

b d h , arcuate pandurate, 35 5 } mm long, 1 5mm

wide, longitudinally des below the middle, ricted ab

pich ins, the disc with an erect pair of denticulate-

ith a mi icle i h hinged to

cies 1 lat “

the column-foot;column slend ncsiaa, Woniay wiped isle Wie akiae™s Sa face tae tare te 2

mm long.

ECUADOR: Zamora- 4 , near Paquisha, alt. ca. 1400 m, collected by G.

Merino, cultivated at Ecuagenera, Gualaceo, 17 Apr. 2001, A. Hirtz 7598 (Holotype: MO), C. Luer il-

lustr. 20038.

This species, presently known only from the Cordillera del Condor, is similar to

the Colombian S. gibberosum. It is characterized by narrow, petiolate leaves and a

coarsely verrucose, more or less horizontal peduncle. The tail of the dorsal sepal is

thick and elongate; the cushions of the synsepal are triangular with rounded bases

abruptly contracted into slender tails more than twice as long as the synsepal. The

petals are dilated on the labellar margin. The lip is arcuate and subpandurate with a

suborbicular, minutely erose apical lobe. Across the middle is a tall pair of fringed

lamellae.

4 apse lum portillae Luer, sp. nov.

: Named for José Portilla of Gualeceo, Ecuador, in whose establishment this — is cultivated.

P iocri » pedunculo

laevi suberecto, cic dorsali cauda incrassato, petalis pectin labello parvo epichilio wea

oblongo deflexo distinguitur

Plant medium in size, epiphytic, caespitose; roots slender. Ramicauls erect, slender, 1.5-2.5 cm

long, enclosed by 2 tubular sheaths. Leaf erect, qin — an tiolate, 8-9 cm long including a

petio ole 3-3.5 cm i. the blade elliptical, subacute, 1 cm wide, narrowly cuneate below into the slender

petiole. I a loose: ti me subdease, linichous, successively sngsd to many-flowered raceme

up to 10-11 cm a i cm aot from low on the

ramicaul; floral bracts 3 mm aor pe mn ing ovary smooth, 3m mm lon light green

with faint dots on the syns labo carinate, t jena. narrowed near the

9 mm long, 3 mm wide, aes to the lateral sepals

for 1 mm, the lateral sepals int ate, concave, ina, 8 mm long, 6 mm

wide, the apical portion ‘of each lateral a Sie by a triangular cushion, 3 mm long, 2 mm wide,

the rounded apices contracted into slender, greenish white tails 6 mm long; petals translucent, sao or

lar, 2 mm long, 1.5 mm wide, the apex obtuse, shortly acuminate, the labellar margin broadly r

the ee margin bilobulate; lip green, thin, the blade oblong, flat with the base pt ned with a a

thin , 2.5 mm long, | mm wide, the apical lobe oblong, deflexed, 1 mm long, 0.5 mm wide;column

alee semiterete, arcuate, winged above the middle, 3 mm long, the thick foot ile: ae 3

mm long.

ECUADOR: Pichincha: near Mindo, alt. ca. 1600 m, cultivated at Ecuagenera, Gualaceo, 5 Mar, 2001,

C. Luer 19729 (Holotype: MO).

This species, recently collected on a western slope of Pichincha, is distin-

guished by narrow, long-petiolate leaves, and an equally long, suberect, successive-

ly flowered raceme. The sepals and petals are not remarkably distinctive, but the lip

is proportionately small with an oblong blade and a narrow, deflexed, apical lobe,

similar to that of S. digitale.

112 - ICONES PLEUROTHALLIDINARUM

Teagueia portillae Luer, sp. nov.

jae

Ety.: Named for José Portilla of Gualeceo, Ecuador, who cultivates this species.

Inter species generis Teagueiae Luer, species haec minutis caespitosa, racemo laxe paucifloro foliis

petiolatis duplolongiore, sepalis longi-acuminatis, petalis filamentosis, labello ovoideo supra medium

fisso distinguitur.

Plant very small for the genus, epiphytic, caespitose; roots slender. Ramicauls slender, erect, 4-5

mm long, enclosed by 1-2 thin, tubular sheaths. erect, coriaceous, petiolate, 2.5-3 cm long includ-

ing a petiole 1-1.5 cm long, the blade elliptical, obtuse, 0.6 cm wide, the base acuminate into the peti

an erect, loosely few-flowered raceme with 2-3 flowers open simultaneously, up to 7 cm

. a: 1 ry 4 49 1 PERE ne * 339 £f sh . 7 tL 3°

= ine ir ie or

15 1 a a. a ee |

ig; p 5 long; ovary 0.5 long greenish white; sepals glabrous, the

dorsal sepal ovate, acute, 4.5 mm long, 0.6 mm wide, 1-veined, long-acuminate into a filiform tail, the

lateral sepals elliptical, oblique, acute, long-acuminate, 1-veined, 5 mm long, each 0.6 mm wide, connate

0.6 mm; petals narrowly linear, 1.8 mm long, 0.2 mm wide, reflexed; lip white, ovoid, 1.5 mm long, 0.6

mm wide, cellular-papular, the sides incurved above the middle to the narrowly obtuse apex, the disc

cleft above the middle, ciliate-pubescent, the base truncate, hinged to the base of the column; column 0.6

mm long and broad, footless, the pex d , the anth bapical, the sti ]

bit VOI dal,

BIRTIARnMOD. 7. Ohinnahi

4amora pe: Parqué Nacional Podocarpus, south of Nambija, alt. 2500 m, cultivat-

ed at Ecuagenera, Gualaceo, 4 Mar. 2001, C. Luer 19744 (Holotype: MO).

This tiny species, superficially resembling a Platystele, is the smallest known in

the genus Teagueia. It is characterized by an erect, hair-like rachis and peduncle

about twice longer than the petiolate leaves. The raceme bears a few, distant flow-

petals reflex. The proportionately large, ovoid lip is held perpendicularly below the

short, stout column. The lip is cellular-papillose with the cleft, common to all

species of the genus, ciliate and pubescent.

i inx acestrochila Luer, Sp. nov.

Ety.: From the Greek acestrochila, “with darning needle-like lip,”” referring to the labellum.

Species haec T. orbicularis (Lindl.) Luer affinis, sed sepalis lateralibus semiconnatis, petalis long-

acuminatis, et labello

Plant medium in size, epiphytic, caespitose; roots slender. Ramicauls erect, slender, 5-7 cm long,

enctosed by 7-8 coarsely ciliated, lepanthiform sheaths with dilated ostia, coriaceous, broad-

ly elliptical-ovate, obtuse to subacute at the apex, 2.5-3 cm long, 1.2-2 cm wide, round or cuneate below

I a cong simultaneously 2- to 3-flowered raceme, born by

a Pes 4-8 mm long, from near the apex of the ramicaul; floral bracts 1.5 mm long; pedicels 1-1.5

6c aoe — 0.75 mm long; sepals white, rose toward the base, glabrous, carinate, thickened toward

narrowly ovate- , 3.25 mm long, 0.7 mm wide, 1-veined, with the margins

lon a ef aye ellow-white at the tip, narrowly ovate, 3.25 mm long, 0.3 mm wide, the apex

the Sane akg ns TeVOlade, the disc With a mazrow, longitudinal Cocina in the comer tal at

peal shea base truncate with a lobule in each corner hinged to the column-foot: col ee

Pct i the middie, 1.75 mm long, denticulate at the apex, the anther and stigma ventral, the foot

Ci ‘ * ue

OLOMBIA: Antioquia: Rio Caldera, collected by E. Valencia, Feb. 1988, flowered in cultivation at

Colomborquideas 16 May 1993, C. Lue, 16908 (Holotype: MO).

; bite ctae is closely allied to T. orbicularis, the two species appearing similar

xcept for the petals and lip. The petals are narrowly ovate with the apex narrowly

acuminate. The lip is also narrowl i ; ? oo

apex revolute. y ovate with the margins of the long-acuminate

SYSTEMATICS OF PLEUROTHALLIDINAE 113

Trichosalpinx manningii Luer, sp. nov.

Ety.: Named in honor of Steve Manning of Tarporley, England, who collected this species.

Inter species Trichosalpinicis subgen. Tubellae, species haec mediocris prolificans, sepalis

Plant medi size, epiphyt ts slender pln eo

ramicaul at the apex, 2-4.5 cm ons. gree pepe 4-5 closely appressed, sheaths with dilated

ostia. Leaf erect, coriaceous, broadly elliptical, obtuse to rounded at the — 15-22 mm long, 10-16

wide, cuneate below into a pers petiole 2-3 mm long. oose, lightly flexuous,

flexible, simultaneously 5- to 7-flowered raceme, 5-6 cm long, including the tee 1-3 cm long, from

near the apex of the ramicaul; floral b ovary | mm long;

white, faintly suffused with rose, glabrous, the dere chs elliptical in the basal third, 15 mm _

including the acuminate, filiform apex, the blade ide, 3 eined, the lateral

veined; lip white faintly suffused with green, trilo bed, 3 mm long, 2 mm wide ex the apical lobe

ovate, subacute, half the length of the lip, with the margins thin, recurved, the basal lobes below the

middle, erect, uncinate, antrorse, the di sie with a narrow, longitudinal cavity in the center; the base

rounded, t e. slender, 2 mm long, bidentate the

iVVUl,

apical, the stigma ventral, the foot 0.5 mm sen

PERU: Without collection data, purchased and cultivated in Tarporley, England, Feb, 2000, 5S. Manning

980709 (Holotype: MO), C. Luer illustr. 19333.

This species is cultivated by Stephen Manning in the National Pleurothallid

Collection in Tarporley, UK, but without collection data. It is distinguished from

all other species of the genus by long-attenuate petals that are similar to the sepals.

Vegetatively, it is characterized by a prolific patie and broadly elliptical leaves,

that are surpassed by a loose, few-flowered raceme. The free d are

translucent white and long-acuminate. The lip is at ee with the lateral lobes

below the middle, uncinate and antrorse. In the center of the disc is a deep, longitu-

dinal, well-defined cavity.

barn inx sijmii Luer, sp. Nov.

Named in honor of A.P. Sijm, Venhuizen, the Netherlands, who collected this oe

tk fractiflexis, racemis paucis lax

cul hs pei ata a subgen. Ti cas sepal abi me om fo zs, sai

nauciflo

; ae | PY

Plant medium in size, epiphytic, caespitose; roots slender. Ramicauls ¢

ot enclosed by 5-6 lepanthiform rm sheaths with oblique, I — =, ron ng. Ifo : .

narrowly elliptical, acute, 5-6.5 cm long, | cm wide, cuneate below into — pe nflor

escence 34 loose, successively paaciay te — 1 heres on ni apd gee ee Cates

tie from aye the apex of the ramicaul; fl ci os ; aon

ovary 1 mm long; sepals orange, glabrous, i dorsal sepal ov moet

sitet 85mm ng, 5 mm wide, 3-v veined, the lateral paren See an Se olen

Ramicauls erect, stout, flexuous, 4-55

rast and thickened to toward

jalate from below the middle, den-

ticulate at the apex, x, 4mm long, the anther and st

ERU: Junin: Chanchamayo valley, alt. 1500 m, 3 Nov. 1995, cultivated in Tarporley, England, A.P.

Sijm 210115 (Holotype: MO), C. Luer illustr. 20031. tise

This species, without close relatives, is apparently pet

distinguished from all other members of the genus ia aaa cal leaf; and a

caul with long-ciliated ostia of the sheaths; a narrowly ellipt

114 ICONES PLEUROTHALLIDINARUM

fascicle of a few, successively few-flowered racemes. The colorful flowers

are relatively large with the lateral sepals forming a deep mentum with an elongated

column-foot. The petals are acute and ciliate. The lip is arcuate, reaching out of the

mentum with its obovate blade. It is ciliate below the middle and biauriculate at the

base

Trichosalpinx werneri Luer, sp. nov.

Ety.: Named for Florian Werner, University of Bayreuth, who collected this species at the Estacién

r Chinchipe.

Species haec T. hirtzii Luer affinis, sed racemo brevi _ fl gni , labelli lobo antico

anguste lineari ciliatis differt.

Plant small, epiphytic, pendent; roots slender at the base of the ramicaul. Ramicauls pendent,

slender, produci h ical from th m long, enclosed by 3 closely appressed, micro-

scopicall scabrous, lepanthiform sheaths. Leaf descending, thickly coriaceous, narrowly elliptical,

fusiform, acute, 28-30 mm long, 4 mm wide, 3 mm thick, cuneate below into a petiole 1-2 mm long.

Inflorescence a congested, simultaneous y 3- to 4-flowered raceme, 6-8 mm long, including the filiform

peduncle ca. 2 mm long, from near the apex of the ramicaul; floral bracts 1 mm long; pedicels 1 mm

long; ovary 0.8 mm long; sepals translucent dark purple, glabrous, ovate, with subacute, slightly thick-

ened apices, the dorsal i

mm, the lateral sepals 4 mm long, 1.2 mm wide, 1-veined, connate 1 mm; petals dark purple, ovate,

subacute, 2.75 mm long, 1.2 mm wide, 1-veined; lip dark purple, 3-lobed, 3 mm long, 2 mm wide

the apical lobe narrowly linear, ciliate, retuse at the tip, 1 mm long, the lateral lobes erect,

broadly rounded with an low, curved callus above the middl , the disc with a low, longitudinal, midline

callus, the base broadly truncate, hinged to the column-foot: column semiterete, 1.2 mm long, the anther

apical, the stigma ventral.

ECTIANND. > Ohinnk. a ss F ‘. a

2000, F. Werner 12017 (Holotype: MO), C. Luer illustr. 19378.

This species is closely related to T. hirtzii Luer. Both species are characterized

by long, flexible, pendent chains of prolific ramicauls with narrowly fusiform

leaves about as long as the ramicauls. The purple flower of T. werneri is produced

simultaneously in a short, congested raceme. The lateral sepals form a deep, round-

ed mentum with an elongated column-foot: the petals are large but simple; and the

lip 2d also deep ly trilobed. The anterior lobe is narrowly linear, ciliate, and retuse at

the tip. The lateral lobes are broadly rounded and erect.

Loja and Zamora, alt. 2200 m, 9 June