JUL 2 4 1984

FIELDIANA

Botany

Published by Field Museum of Natural History

Volume 36, No. 1 August 18, 1972

New Species From South America, I

DONALD R. SIMPSON

ASSISTANT CURATOR, PERUVIAN BOTANY

FIELD MUSEUM OF NATURAL HISTORY

Over the past few years several valuable collections from the

forested regions of Peru have been sent to Field Museum in exchange

for identification. One of these, sent by Dr. Theodore Dudley of the

U. S. National Arboretum Herbarium, contained the new species of

Demosthenesia described below.

A most valuable set of collections of tree species from a dendrol-

ogy project conducted by the Peruvian Forest and Wildlife Service

has been sent to this museum for identification. Dr. Louis 0. Wil-

liams has already described several new taxa from this material,

publishing them in previous issues of this journal. In continuing the

study of this material begun by Dr. Williams, I have encountered

several novelties, one of which is the following new species of Ery-

throxylon from the Department of Tumbes.

The other two novelties are based on collections from a Museum-

sponsored project on which it has been my privilege to work since

joining the staff of Field Museum. This project has been supported

by a continuing grant from the National Science Foundation. Most

of the collections for this project have been made by Mr. Jos£

Schunke Vigo whom I have memorialized in the specific epithet of the

following new species of Tabebuia. Mr. Schunke is a resident of

Tingo Maria, Peru, a son of the natural history collector, Mr. Jos6

Schunke Lopez (given as Jos6 M. Schunke on many herbarium

labels), and grandson of the German botanist and collector, Karl

(Carlos) Schunke.

BIGNONIACEAE

Tabebuia schunkevigoi Simpson sp. nov.

Arbor 30 m. alta; diametro trunci 27 polices. Ramuli aliquantum complanati

longitudinaliter praesertim in internodiis extremis; epiderme ferruginea ad fuli-

Library of Congress Catalog Card Number: 72-

Publication 1152 1

FIELDIANA: BOTANY, VOLUME 36

FlG. 1. Tabebuia schunkemgoi Simpson. A, branchlet with leaves, X %',

B, branchlet with inflorescence, X H; C, flower opened lengthwise, X 1^;

D, anther, X 5; E, bilobed style tip, X 5; F, fruit, X \i; G, seeds, X y±. Drawn

by Marion Pahl.

ginea. Folia opposita, decidua, palmatim composita; foliolis supra olivaceis,

subtus palide subviridibus et flavinerviis, in statu sicco nitide fumosis supra sed

obscure fumosis subtus, ovatis vel ovato-ellipticis, integris, abrupte longe acumi-

natis, ad basim late obtusis vel truncatis et in foliolis lateral! bus parum obliquis;

laminis 65-140 mm. longis et 35-60 mm. latis; petiolulis 10-40 mm. longis, supra

profunde canaliculatis, laevibus, glabris sed minute furfuraceis in sulco et ad

extremis, articulatis ad petiolum et basim folioli, in statu sicco nigris; petiolis

plerumque 7-15 cm. longis, striatis longitudinaliter, canaliculatis in facie adaxiali,

SIMPSON: NEW SOUTH AMERICAN SPECIES, I 3

glabris vel leviter furfuraceis, in statu sicco micantibus nigris; pubibus furfuraceis

pilis dendriticis concistans. Inflorescentia apicalis, thyrsum cum rache et ramis

condensatis, dense tomentosis; pubibus pilis minutis et fusco-flavis. Calyx atrovio-

laceus, 8-11 mm. longus, campanulatus, dense breviter tomentosus, subanthesi ir-

regulariter circumscissilis. Corolla lilacina, plerumque 60-65 mm. longa et 2 cm.

lata ad faucem; lobisS, breviter villosis in am bis paginis, duobus supernis ca. 15 mm.

longis et 12-15 mm. latis. truncatis vel parum emarginatis, tribus infernis ca.

20 mm. longis et 10-15 mm. latis; tubis ca. 4-4.5 cm. longis, extus breviter vil-

losis, intus praeter pilosis in latis uno glabris; basibus tuborum constrictis, 3 mm.

latis et 8-10 mm. longis, et intus et extus glabris, in statu sicco nigris. Stamina

fertilia 4, inclusa, didynama; filamentis longioribus ca. 15 mm. longis; filamentis

brevioribus 10-12 mm. longis; antheris 4-4.5 mm. latis; sacculis antherarum un-

usquisque 2-2.3 mm. longis, vertice connatis, oppositis et parum sursum curvatis

extremetes versus. Stylus 22-28 mm. longus, constrictus ad basim, tune pro parte

ca. 2-2.5 mm. supra basim tumidus, illinc apicem versus angustatus; stigmate

bilobato, lobis ca. 1.5 mm. longis et 0.8 mm. latis. Ovarium ca. 4 mm. longum,

diametro 1.8 mm., cylindricum, obtusum ad apicem, parum contractum ad basim.

Discus cupulatis ca. 1.2 mm. altus, ad basim ovarii arete adpressus atque includens.

Fructus capsulam bicellularem, atropurpureus, in statu sicco nigricans, ca. 22 mm.

latus et usque ad 55 cm. longus, linearis, ad ambo extremo acutus, teres vel septo

contrarium modice complanatus. Semina numerosa, complanata, ca. 12 mm. lata

et 5-6 cm. longa; alis membranaceis.

Tree 30 m. tall; trunk with a d.b.h. of 27 inches (ca. 68.5 cm.). Branchlets

somewhat flattened lengthwise in the ultimate internodes; epidermis light brown

to sooty brown. Leaves opposite, deciduous, palmately compound; leaflets olive-

green above, very light green and with yellowish nerves beneath, in a dried state

lustrous grey-brown above, dull grey-brown beneath, ovate or ovate-elliptic, en-

tire, abruptly long acuminate, broadly obtuse or truncate at the base, in lateral

leaflets slightly oblique at the base; blades 65-140 mm. long and 35-60 mm. wide;

petiolules 10-40 mm. long, deeply grooved above, smooth, glabrous except min-

utely scurfy in the groove and at the ends, black in dried condition, articulate with

the leaflet base and with the petiole tip; petioles mostly 7-15 cm. long, glossy black

in a dried state, striate lengthwise, shallowly grooved on the abaxial surface, gla-

brous or slightly scurfy; scurfy pubescence of tiny dendritic hairs. Inflorescence

apical, a thyrse with rachis and branches condensed, tomentose; pubescence of tiny

brownish yellow hairs. Calyx dark violet, 8-11 mm. long, campanulate, densely

short tomentose, irregularly circumscissile at anthesis. Corolla lilac, mostly 60-

65 mm. long and ca. 2 cm. wide at the throat; lobes 5, short villous on both sur-

faces, two upper lobes ca. 15 mm. long and 12-15 mm. wide, truncate or slightly

emarginate, three lower lobes ca. 20 mm. long and 10-15 mm. wide; tube ca. 4-4.5

cm. long, short villous outside, glabrous within except pilose along one side; the

tube base constricted, 3 mm. wide and 8-10 mm. long, glabrous within and with-

out, black in a dried state. Fertile stamens 4, included, didynamous; longer fila-

ments ca. 15 mm. long; shorter filaments 10-12 mm. long; anthers 4-4.5 mm.

across; anther sacs each 2-2.3 mm. long, apically united, opposite and slightly

upturned at the tips. Style 22-28 mm. long, constricted at base, inflated for about

2-2.5 mm. above the base, then tapering upward ; stigma bilobed, lobes ca. 1.5 mm.

long and 0.8 mm. wide. Ovary ca. 4 mm. long and 1.8 mm. in diameter, cylindri-

cal, obtuse at apex, slightly constricted at base. Disc cupulate, ca. 1.2 mm. high,

enclosing and closely appressed to base of the ovary. Fruit a 2-celled capsule, dark

4 FIELDIANA: BOTANY, VOLUME 36

purple, in dried state blackish, ca. 22 mm. wide and up to 55 cm. long, linear,

acute at both ends, terete or moderately flattened contrary to septum. Seeds

numerous, flattened, ca. 12 mm. wide and 5-6 cm. long; wings membranous.

PERU: Dept. Huanuco: Prov. Pachitea; Dist. Honoria; in "high

forest" (i.e., mature forest) and "low forest" (i.e., successional forest)

across the Pachitea River from Tournavista, alt. ca. 300 m., Jose

Schunke V. 2596 (holotype F; isotypes US, NY, G, COL, USM, and

others). Tahuari de flor morada.

The genus Tabebuia is accepted here as defined by Sprague and

Sandwith (Kew Bull. 1932:18) to include the genus Couralia Splitg.

This species is probably most closely related to C. toxophora (Mart.)

Benth. & Hook., or at least to the herbarium materials to which that

name is usually attached. The nomenclatural validity of the name

C. toxophora (Mart.) Benth. & Hook, and its application could both

benefit from a careful study — a matter which I hope to pursue in a

future publication.

ERICACEAE

Demosthenesia dudleyi Simpson sp. nov.

Frutex scandens, ligneus; ramuli juvenes internodiis ultimis complanati longi-

tudinaliter, tomentosi implexis albis pilis, nodis inflatis; vetior ramulus glabre-

scens, epiderme in reticulum elongatum longitudinaliter findens, exposetens lae-

vem nitentem corticem quae in veteribus ramis aliquantum asperescens lichenibus

et hepaticis tectis. Folia subsessilia; petiolis usque ad 2 mm., plerumque minori-

bus; laminis late ovatis, subcoriaceis, supra pilis glandulosis dispersis inter sparsum

tomentum, subtus fere glabris praeter pilos disperses glandulosos, basi truncata ad

leviter cordata. apice late acuto usque obtuso vel rotundato, nervis 9-11 palmatim

divergentibus e petiolo, reti venularum secondario supra non manifesto, subtus

inconspicuo, marginibus integris sparse ciliatis. Gemma axillaris usque ad 2.5

mm. longa, inclusa a perulis duobus linearilanceolatis carinatis et longiacuminatis.

I nfl orescent! a axillaris, racemosa; rachidi usque ad 2 cm. longa, tomentosa; omni

pedicello subtento a bracteam et ferenti duas bracteolam ad locum circa tertia

parte longitudionis supra basem; bracteis circa 4 mm. longis, elliptico-oblongis, ad

basim truncatis, ad apicem rotundatis ad obtusis, multinervatis e basibus, integris

et ciliatis; bracteolis ovato-lanceolatis, 4-5 mm. longis, ad basim truncatis, longi-

acuminatis, ciliatis, distichis et suboppositis; pedicellis usque ad 18 mm. longis,

sparse tomentosis strictis attenuatis pilis, his in summis tertia parte pedicelli et in

partibus inferne calycum interspersis pilis clavatis et glanduliferis. Calyx usque

ad 1 cm. longus, gamosepalus2-3mm. ebasi; Iobis6-8mm. longis, oblongis, acumi-

natis, glabris vel subglabris, ciliatis. Corolla "carnosa et ceracea intense rubra"

(fide collectoris), 17-22 mm. longa, extus tomentosa, intus glabra, tubulosa, leviter

inflata ad basim; lobis valvatis, 4-6 mm. longis, linearibus, acutis. Stamina 10, ca.

16 mm. longa; filamentis ca. 1 mm. longis, complanatis, marginibus conjunctis ad

FIG. 2. Demosthenesia dudleyi Simpson. A, branch bearing leaves and flowers,

X V->- B flower bud, X 1H; C, pedicel and flower with corolla removed, >

D, corolla tube opened lengthwise showing stamens, X 3'/*; E, filament with

attached anther sacs, adaxial surfaces, X 10. Drawn by Marion Pahl.

6 FIELDIANA: BOTANY, VOLUME 36

kisim: conectivo antherae et parte summa filamentorum vittam constantem ex

pilis serioeis ferenti extus; sacculis antherarum 2.5-3 mm. longis, papillatis; tubulis

antherarum ca. 12 mm. longis laevibus glabris, ad apicem dehiscentibus cum fis-

suris 2.5 3 mm. prolongatis. Stylus indivisus, antheras non excedens. Fructus

mihi ignotus.

A woody vine; young branchlets flattened lengthwise in the ultimate inter-

nodes, tomentose with rather matted, white hairs, nodes inflated; older branch-

lets becoming glabrous, the epidermis splitting longitudinally into an elongated

network, exposing a smooth, shiny bark that on older branches becomes somewhat

roughened and covered with lichens and leafy liverworts. Leaves subsessile, the

petiole not exceeding 2 mm., usually less; blade broadly ovate, truncate to slightly

cordate at the base, broadly acute to obtuse or rounded at the apex, subcoriaceous,

glandular hairs scattered amid a sparse tomentum above, nearly glabrous below

except for the scattered glandular hairs, nerves 9-11, palmately divergent from

the petiole, secondary network not visible above, faint beneath, margins entire,

sparsely ciliate. Bud in the leaf axil enclosed in 2 linear-lanceolate, carinate

and long-acuminate bud scales, to about 2.5 mm. long. Inflorescence axillary,

racemose, the rachis to about 2 cm. long, tomentose, each pedicel subtended by

a bract and bearing 2 bracteoles at a point about one-third of length from pedi-

icel base; bracts elliptic-oblong, about 4 mm. long, truncate based, rounded to ob-

tuse at apex, several-nerved from the base, margin entire and ciliate; bracteoles

ovate-lanceolate, 4-5 mm. long, truncate at the base, long-acuminate, ciliate, dis-

tichous and subopposite; pedicels to about 18 mm. long, sparsely tomentose with

straight attenuate hairs, these interspersed in the upper one-third of the pedicel

nnd the lower part of the calyx with clavate, glandular hairs. Calyx to 1 cm. long,

gam >sepalous for 2-3 mm., the lobes 6-8 mm. long, oblong, acuminate, g'abrous

or subglabrous, ciliate margined. Corolla "fleshy and waxy deep red" (fide col-

lector), 17-22 mm. long, tomentose outside, glabrous within, tubular, slightly

inflated at base; lobes valvate, 4-6 mm. long, linear, acute. Stamens 10, ca. 16

mm. long; filaments ca. 1 mm. long, flattened, their edges joined at the base, tip

of filament and connective bearing a band of sericeous hairs externally; anther

sacs 2.5-3 mm. long, papillate roughened; anther tubules about 12 mm. long,

smooth, glabrous, opening apically by elongated clefts 2.5-3 mm. long. Style

undivided, not exceeding the anthers. Fruit unknown.

PERU: Dept. Cuzco: Prov. La Convencion; fairly common vine,

in enclaves of elfin forest scattered amid the wet grasslands on the

summit ridges at Camp 7, at ca. 3,400 m. alt., ca. 28 km. walking

distance northeast from Hacienda Luisiana and the Apurimac River,

T. R. Dudley 11199 (holotype at F, isotypes at NA and USM).

This species is readily distinguished by the leaves which are very

broad, often slightly cordate, and have a rounded or very broadly

acute apex. Mature leaves are mostly over 2.5 cm. wide and usually

have a length-width ratio of from 1:1 to 1:1.2. The corolla is pube-

scent and in this respect somewhat resembles D. speclabilis (Rusby)

A. C. Smith, but the flower is considerably smaller than in that spe-

cies. It is perhaps most closely related to the group composed of

SIMPSON: NEW SOUTH AMERICAN SPECIES, I 7

D. graebneriana (Hoer.) A. C. Smith, D. mandoni (Britton) A. C.

Smith, D. microphylla (Hoer.) A. C. Smith, and D. fabulosa (Sleu-

mer) A. C. Smith, although the assumed affinity with the lastspecies

is based only on comparison with the description since I have seen no

specimens of it.

ERYTHROXYLACEAE

Erythroxylon pacificum Simpson sp. nov.

Arbor ca. 20 m. alta; cortice ramulorum palide griseo. Folia subcoriacea;

laminis ovatis vel orbicularibus, glabris, supra nitidis praesertim ad maturitatem,

subtus hebetatis, in specimine florifero 3-4 cm. longis et 2.0-2.4 cm. latis, in spe-

cimine fructifero 3-6 cm. longis et 2.1-4.2 cm. latis; petiolis 3-6 mm. longis, supra

canaliculatis, glabris; stipulis 1.8-2.5 mm. longis, non striatis. Pedicelli floriferi

2.5-4.5 mm. longi, valde longistrorsum procrati; pedicelli fructiferi 5-8 mm. longi.

Flores (4) 6-10 in fasciculis axillaribus; lobis calycis 0.8-1.0 mm. longis, late tri-

angularibus; petalis 2.5-3.2 mm. longis, 1.6 mm. latis, oblongis vel leviter obovatis,

late acutis; tubo staminum lobos calycis aequanti vel leviter excedenti; filamenti-

bus in parte discreta ca. 3.5 mm. longis; antheris 0.8 mm. longis, apiculatis, api-

culo usque ad 0.3 mm. longo; ovario tubum staminum excedenti per tamquam 0.8

mm.; stylis per totam longitudinem discretis, ca. 1.2-1.5 mm. longis; stigmatibus

capitatis. Fructus 1.3-1.6 cm. longus, asymmetrice oblongus, uniseminalis. Se-

men endospermia abundantes continens.

Tree, ca. 20 m. tall, d.b.h. 14.7 inches. Branchlets with light grey bark;

vascular trace of stipule on either side apparent as a low ridge extending from

base of stipule to next lower node. Leaves subcoriaceous; blade 3-4 cm. long by

2.0-2.4 cm. wide in flowering specimen, 3-6 cm. long by 2.1-4.2 cm. wide in fruit-

ing material, shiny above especially at maturity, dull beneath, ovate to orbicular,

glabrous; petiole 3-6 mm. long, channelled above, glabrous; stipules 1.8-2.5 mm.

long, nonstriated. Flowering pedicles 2.5-4.5 mm. long, in fruit 5-8 mm. long,

noticeably longitudinally ridged. Flowers borne in axillary clusters of (4) 6-10,

each subtended by a bract; calyx lobes 0.8-1.0 mm. long, broadly triangular; petals

2.5-3.2 mm. long by 1.6 mm. wide, oblong or slightly obovate-oblong, broadly

acute; staminal tube equaling or slightly exceeding the calyx lobes; free filaments

about 3.5 mm. long; anthers 0.8 mm. long, apiculate, apiculus up to 0.3 mm. long;

ovary exceeding the staminal tube by about 0.8 mm.; styles separate throughout,

about 1.2-1.5 mm. long; stigma capitate; fruit 1.3-1.6 cm. long, asymmetrically

oblong, 1-seeded. Seed with abundant endosperm.

PERU: Dept. Tumbes: Prov. Zarumilla; Forest Service Dendrol-

ogy Project Tree no. T-39: flowering collection, Canales 17 (holotype

F, isotypes LIM, MOL, USM, US, NY, P, K) fruiting collection

Canales 6 (paratype F, isoparatypes LIM, MOL, USM, US, NY, P,

K) wood samples at LIMw, MADw, USw, BWCw, CLPw, CTFw,

PRFw, FPAw, CHITw, MEXFw, MERw, TOFOw, RPPRw.

FIG. 3. Erythroxylon pacificum Simpson. A, branch bearing fruiting clusters;

B, flowering cluster; C, fruit; D, flower; E, leaf base, petiole, and stipule. Scale in

A and B = 3 cm., C and D = 1 cm. Drawn by Richard W. Roesener.

SIMPSON: NEW SOUTH AMERICAN SPECIES, I 9

According to Schulz's treatment of Erythroxylaceae in the "Pflan-

zenreich," this species would be assigned to Erythroxylon, section

Archerythroxylon. The section is characterized by stipules nonstri-

ate, flowers perfect, and styles free. There seem to be about seven

species of section Archerythroxylon that occur in the coastal regions

of northern Peru and Ecuador. The following artificial key may

serve to separate those seven species.

1. Leaves obovate, apex rounded to emarginate.

2. Leaves markedly bilineate beneath, the lines paralleling the midrib.

E. carthagenense.

2. Leaves not bilineate or only weakly so.

3. Filament tube distinctly shorter than the calyx.

4. Stipules half the petiole length E. havanense.

4. Stipules as long as petiole or longer E. cumanense.

3. Filament tube equalling or exceeding the calyx.

5. Leaves coriaceous, glaucous beneath, borne on stubby spur

shoots; lateral branches rigid and at right angles to main

branch E. glaucum.

5. Leaves membranous to subcoriaceous, light colored but not glau-

cous beneath; lateral branches erect-spreading.. . .E. hondense.

1. Leaves not obovate, usually elliptic or oblong, apex acute or acuminate.

6. Filament tube distinctly shorter than calyx.

7. Leaf apex obtusely acuminate E. ruizii.

7. Leaf apex rounded to broadly acute E. havanense.

6. Filament tube equalling or exceeding the calyx E. pacificum.

RUBIACEAE

Pentagonia rubriflora Simpson sp. nov.

Arbor 14-15 m. alta. Ramuli, petioli, stipulae et inflorescentiae dense lanatae,

pilis palido-ferrugineis ad rubris. Folia 52-75 cm. longa; laminis 48-65 cm. longis

et 19-22 cm. latis, subtus velutinis, supra sparse tomentosis, elliptico-oblanceo-

latis, ad apicem acutis, ad basim rotundatis ad obtusis, leviter decurrenti in 5-7

cm. longo petiolo; stipulis binatis, gemmam includentibus, marginibus imbricatis,

dextrorsum hemitorsivis, caducis, 4.5-5.5 cm. longis, anguste triangularibus. In-

florescentia cymam ebracteatam, axillaris usque ad 2.5 cm. longae (flores exclusi).

Calyx ca. 22 ( — 30) mm. longus, bifidus (raro trifidus) per % longitudinis, extus

lanatus, intus glaber, viridoluteus, in statu sicco fuliginosus. Hypanthium dense

lanatum, in statu sicco argenteum cinnamomeo suffusum. Corolla usque ad 35

( — -45) mm. longa, rubra, extus glabra vel sparse pubescens sed versus apicem

loborum sublanata, intus praeter inter basibus filamentorum dense pilosis glabra;

lobis ca. 6-10 mm. longis et 4-5 mm. latis. Stamina 5; filamentis 11-14 mm.

longis, glabris; antheris ca. 3 mm. longis, caudatis, affixis ad basim. Discus cupu-

latus, ca. 2.5 mm. altus. Stylus ca. 18 mm. longus, bifidus; lobis 2.5 mm. longis.

Fructus mihi ignotus.

FIG. 4. Pentagonia rubriflora Simpson. A, leaf, bud, and flower cluster;

B, flower; C, flower with corolla tube opened lengthwise. Scale in A = 5 cm.,

in B and C = 3 cm. Drawn by Richard W. Roesener.

10

SIMPSON: NEW SOUTH AMERICAN SPECIES, I 11

Tree 14-15 m. tall. Branchlets, petioles, stipules, and inflorescences densely

woolly with a light rusty to rufous pubescence. Branchlets strongly 4-ribbed in

at least the ultimate 2 in tern odes; pubescence becoming dark and sloughing off

older parts of twigs. Leaves opposite, 52-75 cm. long; blades 48-65 cm. long by

19-22 cm. broad, velvety tomentose beneath, velvety sparse tomentose above,

olive green above, lighter beneath, elliptic-oblanceolate, apex acute, base rounded

to obtuse and slightly decurrent onto the 5-7 cm. long petiole, some blades nearly

pandurate in the lower half; stipules paired and enclosing the bud, margins over-

lapping, dextrally half-spiraled, light straw- or ochre-colored, caducous, 4.5-5.5

cm. long, narrowly long-triangular, when fallen, exposing a sheathing, 3-5 mm.

long, ciliate collar that encircles the stem at the nodes. Inflorescence an axillary,

ebracteate cyme, to 2.5 cm. long (excluding flowers). Calyx to 22 (rarely 30) mm.

long, bifid or rarely trifid for two-thirds its length, woolly pubescent outside, gla-

brous within, greenish yellow drying to a sooty grey-brown. Hypanthium densely

woolly, a cinnamon tinged silver color in the dried material. Corolla to 35-45 mm.

long, red, glabrous or very sparsely pubescent outside except becoming almost

woolly toward the tips of the lobes, glabrous inside except densely pilose between

the filament bases; lobes about 6-10 mm. long by about 4-5 mm. wide. Stamens

five, filaments glabrous, 11 to 14 mm. long; anthers basally attached, caudate,

about 3 mm. long. Disc cupular, about 2.5 mm. high. Style ca. 18 mm. iong,

bifid; lobes ca. 2.5 mm. long. Stigma lining the inside of the stylar lobes. Fruit

unknown.

PERU: Dept. San Martin: Prov. Mariscal Caceres; Dist. Cam-

panilla; Rio Sion, southwest of the village of Sion, in high forest, alt.

ca. 450 m., Oct. 3, 1969, Jose Schunke V. 3472 (holotype F, isotypes

LIM, US, COL, NY, G, USM, MO, K, P, MG, INPA).

This species is closely related to P. spathicalyx Schum. and P.

velutina Standl. The following key may serve to separate the three

species.

1. Leaf blades glabrous or subglabrous on both surfaces; bracts or bracteoles

present, red colored; calyx glabrous to sparsely pubescent; corolla white to

yellowish white, occasionally with pinkish lobes; widespread in Amazonian

Colombia, Peru, and Brazil P. spathicalyx.

1. Leaves densely pubescent at least beneath; bracts absent or inflorescence

densely congested; calyx various; corolla various.

2. Leaf blades densely pubescent beneath, glabrous above, 40-45 cm. long by

20-25 cm. wide; calyx sparsely puberulous; corolla unknown; known only

from type locality at Puerto Arturo, near Yurimaguas, Dept. Loreto, Peru.

P. velutina.

2. Leaf blades densely pubescent beneath, puberulous above, 48-65 cm. long

by 19-22 cm. wide; inflorescence ebracteate; calyx pilose, greenish yellow;

corolla red; from near Si6n in the middle valley of the Huallaga River below

Tocache. . P. rubriflora.

UNIVERSITY OF ILLINOIS-URBANA