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C. VILVENS

INTRODUCTION +

NOVAPEX I18CHS 11): 1-67, 10 mars 2017

New species and new records of Chilodontidae (Gastropoda:

Vetigastropoda: Seguenzioidea) from the Pacific Ocean

Claude VILVENS

Rue de Hermalle, 113 - B-4680 Oupeye, Belgium

Scientific Collaborator, Muséum national d'Histoire naturelle, Paris.

vilvens.claude @skynet.be

KEY WORDS. Gastropoda, Chilodontidae, Indo-Pacific, Vaceuchelus, Herpetopoma, Euchelus,

Ascetostoma, Clypeostoma, Pholidotrope, Danilia, Perrinia, new records, new species.

ABSTRACT. New records of Chilodontidae species described from various Pacific localities are

listed, extending their distribution.

15 new species are described from New Caledonia, Fiji, French Polynesia, Solomon Islands and

Taiwan, and compared with similar species: Vaceuchelus cavernoïides n. sp. V. phaios n. sp., V.

rapaensis n. Sp., Herpetopoma pantantoi n. sp., H. vitilevuense n. Sp., H. hivaoaense n. sp.,

Euchelus polysarkon n. sp., Ascetostoma pteroton n. sp., Clypeostoma chranos n. sp., C. adelon n.

Sp., Pholidotrope asteroeides n. Sp., P. choiseulensis n. Sp., Danilia stroggylon n. sp, Perrinia

cantharidoides n. sp. and P. guadalcanalensis n. sp.

Two new synonymies are established: Vaceuchelus saguili Poppe, Tagaro & Dekker, 2006 from

the Philippines is synonymized with V. favosus (Melvill & Standen, 1896), and V. vangoethemi

Poppe, Tagaro & Dekker, 2006 from the Philippines is synonymized with V. clathratus (A.Adams,

1853)

RESUME. De nouveaux relevés de Chilodontidae décrits de diverses régions de l'Océan Pacifique

sont listés, étendant leur aire de distribution.

15 nouvelles espèces sont décrites de Nouvelle-Calédonie, des Iles Fidji, de Polynésie française,

des Iles Salomon et de Taïwan, et sont comparées avec des espèces similaires: Vaceuchelus

cavernoides n. Sp., V. phaios n. Sp., V. rapaensis n. Sp., Herpetopoma pantantoi n. sp., H.

vitilevuense n. Sp., H. hivaoaense n. Sp., Euchelus polysarkon n. sp., Ascetostoma pteroton n. sp.

Clypeostoma chranos n. sp., C. adelon n. sp., Pholidotrope asteroeides n. Sp., P. choiseulensis n.

sp., Danilia stroggylon n. Sp., Perrinia cantharidoides n. sp. et P. guadalcanalensis n. sp.

Deux nouvelles synonymies sont établies : Vaceuchelus saguili Poppe, Tagaro & Dekker, 2006

des Philippines est placé en synonymie de V. favosus (Melvill & Standen, 1896), et V.

vangoethemi Poppe, Tagaro & Dekker, 2006 des Philippines est placé en synonymie de V.

clathratus (A. Adams, 1853).

Hickman & McLean (1990) brought a first major

Chilodontidae species are probably one the less

known Seguenzioidea and Trochoidea species. Apart

from some remarkable taxa such as some Euchelus or

Herpetopoma species, the others are often confused or

simply ignored. Fortunately, since the 2000s, some

authors undertook the clarification of the knowledge

of some genera (Poppe et al., 2006: Herbert, 2012 &

2015), even also with comparison to fossil taxa from

Jurassic and Cretaceous (Bandel, 2010).

Besides this fact, even the taxonomic position of this

group of taxa has changed many times during these 30

late years. Without any aim of exhaustiveness, one can

note the following steps:

+ Vaught (1989) still considered that genera as

Euchelus, Herpetopoma and Calliotropis as well

belonged to the Margaritinae subfamily among

Trochidae Rafinesque, 1815 species within the

Trochacea Rafinesque, 1815 superfamily:

change while ranking these taxa into the Eucyclinae

Koken, 1897 family, grouping Euchelus,

Herpetopoma and related in a tribe Chilodontini

Wenz, 1939, while ranking Calliotropis and related

into a new tribe Calliotropini:

+ Bouchet et al. (2005) produced a second

disruption while moving the Chilodontini and

Calliotropini taxa into the subfamilies Chilodontinae

Wenz, 1938 and Calliotropinae Hickman & McLean,

1990, forming with Cataeginae McLean & Quinn,

1987 the family Chilodontidae Wenz, 1938 attached to

the Seguenzioidea Verrill, 1884 superfamily:

+ Finally, various accurate studies (Kano, 2008:

Kano et al., 2009) leaded to identify 6 phylogenetic

groups in Seguenzioidea among those were

Seguenzlidae, Chilodontidae, Calliotropidae and

Cataegidae, giving to this three latter groups a full

family status: this latter contribution is taken in

account on WoRMS (World Register of Marine

Species).

C. VILVENS Chilodontidae from the Pacific Ocean

The present work describes several new species goal to study the biodiversity of invertebrates,

oine ‘ 1d: Fr ari : È : : : 7 - of x

belonging to the Chilodontidae from various Indo especially mollusces, in a wide range of habitats of the

Pacific areas and confirms and/or extends the

distribution area from known chilodontid species. This

iS also another attempt to enlighten the discriminant

Indo-Pacific (e.g. Crosnier et al., 1997; Bouchet et al.,

2008). These campaigns have covered among others

features to use in the study of these species. New Caledonia, Loyalty Islands, Fiji, French

Polynesia, Tonga Islands, Vanuatu, Solomon Islands,

MENÇGHALANEMENRONS Indonesia, Philippines and Taiwan. The following

table lists the expeditions that have collected the

The material studied in this paper was collected by udied samples:

French IRD-MNHN expeditions started up with the

; North Pactfic

1 AIN

À à 6) Fe 4 , RATS

PE PE CO CE PAP"

CR ME TA M 4 £, ja

TX 2 NA

À VE | \

CRÉENT 2e

Tara) uen Re

Figure 1. Map of approximate locations of MNHN expeditions referenced in this paper :

: TAIWAN 2000, TAIWAN 2001, TAIWAN 2002;

: KARUBAR;

: SALOMON 1, SALOMON 2, SALOMONBOA 3;

: MUSORSTOM 8, SANTO 2006, BOA 0, BOA |;

: MUSORSTOM 10, BORDAU 1, SUVA 2, SUVA 4;

: MUSORSTOM 9, MARQUISES, POLYNESIE FRANCAISE;

: BORDAU 2;

CORAIL 2:

: PALEO-SURPRISE, EBISCO, BATHUS 3;

: CHALCAL 2, SMIB 3, SMIB 8, BERYX 11, BATHUS 3, NORFOLK 1, NORFOLK 2;

*% : VAUBAN 1978-1979, LAGON, BIOCAL, MUSORSTOM 4, BIOGEOCAL, Campagnes d'essais 1987,

BATHUS 1, BATHUS 2, MONTROUZIER, BATHUS 4;

#%: AURORA, MUSORSTOM 3, PANGLAO 2004, PANGLAO 2005;

% : MUSORSTOM 6, VOLSMAR, LIFOU 2000;

+ : BENTHAUS, RAPA 2002;

x : MUSORSTOM 7.

HS+0Umo©on+*es+

C. VILVENS NOVAPEX 18CHS 11): 1-67, 10 mars 2017

5-0/2007

3/1993

Southern New Caledonia

Norfolk Ridge and Loyalty Ridge

Southern New Caledonia and Loyalty Islands

Loyalty Basin

Vanuatu

BOA !

5/1987

Norfolk Ridge

9-10/1999

TAIWAN 2002 North-eastern and southern Taiwan 5-9/2002

VAUBAN 1978-1979 New Caledonia 1978-1979

VOLSMAR Matthew and Hunter 5-6/1989

Table 1. List of the Indo-Pacific MNHN expeditions mentioned.

C. VILVENS

Regarding the distribution of the new species and the As for

Chilodontidae from the Pacific Ocean

the description methodology, the main

extension of the distribution of known species, the conchological features used are (Figs 2 & 3):

range 1s taken from the internal intervals of the two

extremes values.

+ general shape of the shell: spire depressed or depressed -

conical, cyrtoconoidal (=convex cone-shaped), coeloconoiïdal

(=concave cone-shaped);

+ size and shape of the protoconch:;

+ shape of the whorls: convex, concave, straight - with or without

shoulder or keel;

+ spiral cords of the whorls: ontogeny, number, beaded or

smooth, distance between cords, especially features of the

subsutural cord;

+ axial sculpture of the whorls;

+ shape of the aperture, denticles in the outer lip;

+ shape of the base and spiral cords: number, beaded or smooth,

distance between cords;

+ features of the umbilicus: open or covered with a callus, relative

size, presence/lack and shape of a parietal inducture;

+ columella: excavated or not, with or without columellar tooth or

teeth;

+ colour: background colour, pattern with possible flames, spots

Or marks.

colummella

adapical

columellar

tooth

abapical

columellar

tooth

apertural

denticles

Figure 2. Useful features of chilodontid shells

(1) (shell: Clypeostoma adelon n. sp., Loyalty

Islands, LIFOU 2000, stn 1461, 100-120 m,

5.8 x 3.8 mm).

apertural

denticle

columellar

tooth

parietal

inducture

umbilicus

Figure 3. Useful features of chilodontid shells (2): H: height; W: width; HA: height of the aperture (shell:

Euchelus dampierensis Jansen, 1994, Western Australia, Broome, intertidal area, 10.8 x 8.3 mm, coll C.Vilvens).

C. VILVENS

Abbreviations

Repositories

MNHN: Muséum national d'Histoire naturelle, Paris,

France;

NHMUK: Natural History Museum of the United

Kingdom, London, England;

USNM: National Museum of Natural

Smithsonian Institution, Washington, USA.

History,

Other abbreviations

H: height;

W: width;

HA: height of the aperture;

TW: number of teleoconch whorls;

PI, P2, P3, ..: primary cords (PI is the most adapical);

AE E. S3, secondary cords (SI is the most

adapical);

T1, T2, T3, ...: tertiary cords (numbered following

appearance order);

stn: station;

lv: live-taken specimens present in sample:

dd: no live-taken specimens present in sample;

sub: subadult specimen;

juv: juvenile specimen:;

est: estimated;

coll: collection;

CV: Claude Vilvens collection.

SYSTEMATICS

Superfamily SEGUENZIOIDEA Verrill, 1884

Family CHILODONTIDAE WENZ, 1938

NOVAPEX I8CHS 11): 1-67, 10 mars 2017

Remarks. Herbert (2012) has given a very accurate

characterization of the Chilodontidae from

conchological and anatomical points of view, with in

addition a useful key to genera of Chilodontidae in the

south-western Indian Ocean.

Moreover, the study of the radula has finally led him

to move the genus 7ibatrochus from Chilodontidae 10

the Calliotropidae (Herbert, 2015). Earlier, Kano et al.

(2009) have already established that Turcica H. & A.

Adams, 1854, belongs to the Calliotropidae. So these

two genera are excluded from the present study

focusing only on Chilodontidae.

Mainly, the shells of the family can be characterized

+ a protoconch with a diameter usually between

200-300 pm, exsert and rather globose to, on the

contrary, sunken and almost at the same level as the

first teleoconch whorl:

+ a rather elevated teleoconch (H/W>=1), without or

with a rather narrow umbilicus;

+ a first whorl with axial threads: spiral cords

appearing on second or third whorls; last whorls with

+ a more or less marked reticulate (sometimes even

foveolate) pattern produced by crossing of axial and

spiral sculpture:

+ a columella without or with one or two teeth, very

weak to very strong, the abapical one rounded, pointed

or squared;

+ possible denticles in the outer lip of the aperture.

The table 2 on the next page lists the taxa (known

species and new species [in bold type]) examined in

this study:

C. VILVENS Chilodontidae from the Pacific Ocean

Vaceuchelus Iredale, 1929

Vaceuchelus scrobiculatus (Souverbie in Souverbie & Montrouzier, 1866) ss

Vaceuchelus foveolatus (A.Adams, 1853) .sssassennrrneneteresmmeneren metiers ere t etat terrier 8

Vaceuchelus favosus Melvill & Standen, 1896)... esseenssrmemeenererresssasonsnennnossmensernerems ressens éaa sang ete 10

Vaceuchelus auricatris Huang & Fu, 20052... évasion pentes 11

Vaceuchelus Cayernoidés n, SD. uoonnrnnenatnreennnresmcsertetn ee TE NE RER 1]

Vaceuchelus DAS DRSD..... 2 ssae sé snnenneotiense tant tt dame rare tre te RE ENEES 14

VaceuGhelns TADAERSIS De SD remember EEE 16

Herpetopoma Pilsbry, 1889

Herpetopomasemmatum (God Mes) nn eeereneraau their re DT 19

Herpetopomainsmetum(Goud SO Re rer eee EE 20

Herpetopoma*entolun (Mel TOR ee ane annnrasnersarrree ee CO Re 22

Herpelopomapantantol. D: Sp: rrimencemennremmruub torrent 22

Herpetopomavernracat(GonidMB6l) sise nseesmremmanntatrntme eo ie RE 24

HerpétonomatdiwimaePoppe Tasaro @ Dokker, 2006)... rer 26

HerpetopomanoicnidmNiulvens, 2012 nn er ea tr 26

Herpelopomapuilepuense he SD: RE ut CE RO 28

HerpetonomanaokoaePoppe, Taparo @ DekKer 2006... re 30

HÉRDOODOMUHIVAOAENRS CIS DR RP ARE a reset us rer Rte nn ST ne D De lee TRE PE 30

HerpétopomacCorrUe dires MOOD M daurercesstescectersie eee ce een ne er ee 0 TE CO OO 31

Harnetopoma rubrume CASAdaMS SSP MR orienter ie red 32

Euchelus Philippi, 1847

ProhRelus DONSUTRONNNE SDE 2 srecnesmmonesonereacasiioerseernte dau tr einer A UT 34

Puchels ares (@imeltes 70) SR Re MO cc rusanete dre cd nen este se sepeser cree ee AR A 2 35

Ascetostoma Herbert, 2012

ASCaloomanneens(SCheEpDmaAn A OS nn sers dec e tee -crepheres 35

ASCCLOSIONTE DICTOLON SDS nee sn nee Ne eu nn Te ee nl Rene Passe dcr nnsees Lo sre ne 38

Hybochelus Pilsbry, 1890

Hbochelascancelani RSS MR ne etre den eee 39

Clypeostoma Herbert, 2012

Chhpeosionanortont MOMIE Re rene mn re encr tipo nie reel 42

CIPEOSOMA CHTAROS SD ane nn ne Re ee SR nee unes anses ti RE Se nn 42

ChpeostomaecancelammmaBenepnan MO ER een e etes et ie 43

Clpeostomacedleae (Reopperhsao CDEkKer 200 ec Liévin en di

Clpéostomaade lon ÉQReUn eee en e e D e e cernes eine es 46

Pholidotrope Herbert, 2012

BRONTOOPEUSIOT CIO ES D nr ta I nd cette tenter 48

Pholidotrope:ChROISCHERSIS ES DE Re rss ste er 49

Danilia Brusina, 1865

Danilia ansulosa VINS Hé OS 200 ed RER cernes renier ete nnereniaes S1

Danilia discordata Vie alé 200 su

D'anilid euchelnonms (Nour SO ue 52

Dana SO EVION 1: SD PR ee ee Un Ne eee A M duerv errant end attcRne 52

Chilodonta Etallon, 1862

Chilodonia suduiraut BODpE MAS AO DER ne 56

Perrinia H. & A.Adams, 1854

Perrine ansulifera (A ADS MS) a ren ec nee 60

Péerrmaschsa (Gould, D Re ee ete ea ce 61

Perrine Canthar lo le SD annee sert 61

Perrinia guadalcanalenSiS Sp Re RE ecran ira ne ed 62

Perrinia SquañocarmaliS ea

Table 2. List of the Chilodontidae species studied in this paper.

C. VILVENS NOVAPEX 18(HS 11): 1-67, 10 mars 2017

Genus Vaceuchelus lredale, 1929, labral denticles inside the outer lip of the aperture,

Type species: Euchelus angulatus Pease, 1867 [= others lack these denticles. Especially, Herbert (2012)

Monodonta foveolata À. Adams, 1853] (by monotypy) pointed out as the main difference between

— Recent, Pacific Ocean. Herpetopoma and Vaceuchelus genera the presence

(Herpetopoma) or the lack (Vaceuchelus) of a peg-like

Remarks. The main features for Vaceuchelus are a tooth coupled with a U-shaped notch on the basal lip

Small size (H < 10 mm), a whitish colour and a rough, near the columella.

cancellate or foveolate sculpture. Some species have

Vaceuchelus

shell size number which Si | denti- | num- open colour frontal

shape (mm) of cords present ? | clesin | berof | umbi- view

on penul- aper- cords licus ?

timate ture ? a

whorl

re yellowish white

white to

scrobiculatus

rounded | 6.5x5.5

conical

roundly

conical abapical

to SL. cord

Cyrto- covered by

conoiïidal EE

favosus cyrto-

conoidal

auricatris Cyrto-

conoidal

foveolatus

yellowish white

Re:

nacreous white

to bronze

white with

reddish brown

cavernosus* | roundly

conical

cavernoides sl.

n. SP. cyrto-

conoidal

sl.

cyrto-

conoidal

rapaensis cyrto- FX £ ss. Y (partly | yellowish

n. SP. conoidal covered nacreous white

nacreous brown

to pinkish white

phaios n. sp.

clathratus * conical 9,3x7.8

Table 3. Comparison of conchological features of some central Indo-Pacific Vaceuchelus species (sl. = slightly.

poss. = possible, * = for comparison only, not studied in this paper).

Vaceuchelus scrobiculatus 165°17'E, 45-56 m., 2 dd. — Stn 1272, 20°41'S,

(Souverbie in Souverbie & Montrouzier, 1866) 164°15'E, 82-120 m, 1 dd. — Stn 1273, 20°5056,

Figs 4A-J, Table 3 165°23'E, 20 m, 1 dd, 1 dd juv. — Stn 1315, 20°41'S

164°15'"E, 66-87 m, 4 Iv, 3 sub dd. — Stn 1323,

Trochus scrobiculatus Souverbie in Souverbie & 20°4/1'S, 164°15'E, 82-120 m, 6 dd. — Stn 1331,

Montrouzier, 1866: 140, pl. 6, fig. 9. Type locality: 20°50'S, 165°20'E, 10 m, 1 dd. — Stn 1350, 22°18'S,

Art Island, New Caledonia. 166°20'E, 3-6 m, 2 dd, 2 dd juv. - BATHUS |: stn

Vaceuchelus scrobiculatus — Herbert, 1996: 424-426, DW692, 20°35'S, 164°S9J'E, 140-150 m, 2 dd. — Stn

fies. 32-33. DW1233, 22°24'S, 166°4$8'E, 45-50 m. 40 1v. -Menou

Vaceuchelus scrobiculatus — Poppe, Tagaro & coll. Baie St Vincent, ilot Tenia, 42 m, 1 dd sub, 1 dd

Dekker, 2006: 47. juv.

Loyalty Islands. LIFOU 2000: stn 1445, 20°51'S,

Material examined. New Caledonia. 167°10'E, 10-12 m. 1 dd.

MONTROUZIER: stn 1261, 20°46'-20°47TS, 165°15"-

C. VILVENS

Chilodontidae from the Pacific Ocean

Vanuatu. MUSORSTOM 8: stn DW1094, 15°08'S,

167°12'E, 312-314 m, 1 dd. —- SANTO 2006: stn

DB25, 15 538$, 107 1LE 10m dd, "SD SS0

15°25'S, 167°1d'E, 6-43 m, 2 dd, 1 dd juv. — Stn

DB6S, 15275, 1671CE. 2m 2 (id =#SinDB60;

15°24$, 167°13'E, 38 m, l dd, 1 dd juv. — Sin DB75,

523$ 1070 E 20m

Fi. MUSORSTOML. 10:- sin. CP1366, “182135

178°33'E, 149-168 m, 1 dd juv. -SUVA 4: stn DWO8,

18922S;17S°02/E; 28-30hm, L'Ad

Philippines. PANGLAO 2004: stn B12, 09°36'N,

123°43'E, 24-27 m, 2 dd. — Stn B24, 09°29'N,

125S0E; 38m L'dd, 2 ddsub/ 1Fdd iv — Sim 525,

OOPA2IN, 123 S1E; 21m, dd — Sin 149% "9287IN;

123°45'E, 90 m, 1 dd, 1 dd juv. - AURORA 2007: stn

DW2750, 15°53'N, 121°54'E, 538 m, 2 dd.

Distribution. New Caledonia, 6-140 m, 1v at 50-66 m:;

Loyalty Islands, 10-12 m (dd); Vanuatu, 10-312 m

(dd); F1, 30-149 m (dd); Philippines, 21-538 m (dd).

Remarks. The main characteristics of this species are:

+ height up to 6.5 mm, width up to 5.5 mm;

+ rather elevated spire;

+ teleoconch rounded conical, of about 4 convex

whorls with 3 granular spiral cords on penultimate

whorl; rounded periphery; more or less orthocline fine

threads on first whorl, thickening on next whorls, very

thick on last whorls; P3 at first whorl, P2 at second

whorl!, PI at half third whorl; P2 and P3 the strongest;

on last whorl, P4 peripheral, distance between P2 and

P3 the largest, at least 2x width of cords;

+ sculpture initially cancellate, becoming strongly

foveolate with growth;

+ aperture subcircular to weakly elliptical, with

denticles in outer lip;

+ base convex, with 3 thick spiral cords; distance

between cords about 1.5x width of cords;

+ no umbilicus;

+ yellowish white.

Vaceuchelus foveolatus (A.Adams, 1853)

Figs 4K-S$S, Table 3

Monodonta foveolata A.Adams, 1853: 176. Type

locality: Lord Hood's Island (nowadays Marutea

Atoll), Tuamotu group, French Polynesia, 15-18 m.

Euchelus angulatus Pease, 1867: 283, p1.23a, fig. 27.

Syn.

Euchelus (Herpetopoma) foveolatus - Marshall, 1979:

524-525, figs. 2A-E.

Euchelus foveolatus — Kaicher, 1990: card # 5790.

Euchelus angulatus — Herbert, 1996: 426, figs. 74-75.

Syn.

Vaceuchelus foveolatus — Poppe, Tagaro & Dekker,

2006: 48, p1.16, fig. 2.

Vaceuchelus foveolatus — Vilvens, 2012: 8, figs 24-25.

Vaceuchelus foveolatus — Herbert, 2012: 447, 451.

Material examined. Philippines. PANGLAO 2004:

stn B41, 09931'N, 123°41'E, 17-19 mm, Ldd, 1dd juv. —

PANGLAO 2005: stn DW2374, 0O8°44'N, 123°14'E,

105-109 m, 1 dd.

Solomon Islands. SALOMON I: sin DWI1742,

11°29'S, 159°57'E, 366-421 m, 1 dd. - SALOMON 2:

stn DW2234, 6°51'S, 156°24'E, 192-277 m, 1 dd.

French Polynesia, Australes Archipelago.

BENTHAUS: sin DW2001, 22°27%S/151720W, 200;

2S0malhdd

Distribution. Solomon Islands, 277-366 m (dd);

French Polynesia, Australes Archipelago, 200-250 m

(dd); Philippines (using also data of Poppe et al.

2006), 4-105 m (dd).

Remarks. The main characteristics of this species are:

+ height up to 6 mm, width up to 6 mm;

+ moderately elevated spire;

+ teleoconch roundly conical to weakly

cyrtoconoidal, of about 4 convex whorls with 4 strong,

similar in size, moderately thick, granular spiral cords

on last whorl; P3 appearing near end of first whorl,

quickly strong on second whorl and making keel; P2

appearing on second whorl, quickly almost as strong

as P3; PI resolving and P4 emerging from suture on

third whorl; last whorl very convex, the three abapical

cords making keel, distance between them 2x width of

cords;

+ prosocline, rather thick axial threads: sculpture

cancellate, more or less foveolate on last whorls:;

+ aperture subcircular, with denticles all around the

outer lip;

+ base convex with 3 spiral cords, the innermost a

bit weaker and bordering the umbilicus; distance

between cords 1x to 1.5x width of cords;

+ narrow umbilicus, deep, possibly partly covered

by a callus on mature shells;

+ white to yellowish white.

Figure 4 (scale bar: 1 mm).

A-J. Vaceuchelus scrobiculatus (Souverbie in Souverbie & Montrouzier, 1866). A-C. New Caledonia.

MONTROUZIER, stn 1272, 10 m, 5.5 x 4.2 mm. D-F. Philippines, PANGLAO 2004, stn B24, 38 m, 5.6 x 5.3

mm. G-J. New Caledonia, BATHUS 1, stn DW1233, 45-50 m. G-H. 3.9 x 3.3 mm. I-J. 4.9 x 4.4 mm.

K-S. Vaceuchelus foveolatus (A.Adams, 1853). K-M. French Polynesia, Australes Archipelago, BENTHAUS.

stn DW2001, 200-250 m, 4.2 x 4.3 mm. N-P. Solomon Islands, SALOMON 2, stn DW2234, 192-277 m. 2.9 x

3.0 mm. Q-$. Philippines, PANGLAO 2005, stn DW2374, 105-109 m, 7.0 x 3.5 mm.

C. VILVENS

Chilodontidae from the Pacific Ocean

Marshall (1979) gave an accurate description of the

conchological main features. We follow here Herbert

(1996, 2012) who considers that Vaceuchelus

angulatus (Pease, 1867) 1s a likely synonym of V.

foveolatus.

Vaceuchelus favosus (Melvill & Standen, 1896)

Figs 5A-J, Table 3

Euchelus favosus Melvill & Standen, 1896: 311,

pl.XI, fig. 74. Type locality: Lifou, Loyalty Islands.

Vaceuchelus favosus — Herbert, 2012: 447, figs. 70E-

Vaceuchelus saguili Poppe, Tagaro & Dekker, 2006:

50-51, pl.18, fig.3. Syn. n.

Material examined. New Caledonia.

MONTROUZIER "sin" 1250, "20747S IGSAE, 3-

On dd SinUi2SSs 20848) CS208IE Lim, dd:

SU AS POS NICE 2526:m 2 dd Sin

1259, 20°45S, LESOIAE, 15-35 m, 11 dd, 1 ddjuv. —

Stn 1268, 20°45'S, 165°08'E, 9-1 1m, 2 dd. — Stn 1269,

2UBSS MC ONE MS 20m. 25 dd Sin 1270,

20°45'S, 165°17'E, 10-35 m, 3 dd, 4 dd juv. — Stn

1271, 20°45'S, 165°08'E, 5-25 m, 10 dd. — Stn 1272,

20°50$, 165°20'E, 10 m, 16 dd, 1 dd juv. — Stn 1273,

20850S, 165°23)E, 20m. 201 dd Sim 1291/2022,

164°07'E, intertidal, 1 dd. — Stn 1306, 20°39%,

NORME ES in, AN Id Sn 1816 "20040!S

GAME 2 un Tdid = Sn1822/208 IIS MCE,

53-71 m, 2 dd sub. — Stn 1323, 20°41'S, 164°15'E, 82-

120 m, 1 dd. — Stn 1331, 20°40'S, 164°11'E, 55-57 m,

SO AINIIAUT. StnnsS 10 22220S; AICO26E

intertidal, 1 dd, 2 dd sub, 2 dd juv. — Stn1351,

22220/S, 166°26'E, intertidal, 1'dd sub. — Sin 1352;

2222 160 16!E, 27-35 mn, 12 dd, 2 dd sub, 91vjurv.

= Stn 1354, 22°22S, 166°16!E, 27-37 m, 5 dd. — Sin

1284, 20°34'S, 164°11'E, intertidal, 5 1v, 1 dd juv. —

NORFOLK 1: stn DW1727, 23°17'S, 168°14'E, 190-

212 m, 3 dd.

Loyalty Islands. LIFOU 2000: stn 1410, 20°57S,

lOTMOSIE im, 2 dd = Sms, 205S5S)S AG/MOSIE,

3-10. 2 dd. Sin JA, 20P47S; 167 081E; 1-S'm, 1

HS 22 20 PS ICONE, in, dd dd

juv. — Stn 1425, 20°47'S, 167°07'E, 4-5 m, 3 dd, 1 dd

juv. — Stn 1429, 20°48'S, 167°07E, 8-18 m, 5 dd. —

Stn 1430, 20°48'S, 167°07E, 20-25 m, 1 dd, 1 dd ju.

= Sin 1492, 20PS%S C0 E 12-2m; 52 dd Sin

1434, 20°53'S, 167°08'E, 5-20 m, 15 dd. — Stn 1435,

DUPSSIS, ICONE S#20m.-22 dd "Sin 1420,

DUPSCS. ICT OTE, 1020 m0, 2 dd Soul

20°46'S, 167°02'E, 20 m, 2 dd, 1 dd juv. — Stn 1442,

20°46'S, 167°02'E, 47 m, 2 dd. — Sin 1444, 20855S,

167°05'E, 9-20 m. 1 dd. — Stn 1448, 20°46$S,

167°02'E, 20 m, 7 dd, 3 dd sub. — Stn 1449, 20°46%,

16 02/E im. l'add Sin 1450; 2016) SMIO7AUZIES

Dyélu S dd, = Sin AS I 20 RS MCAOMENIIOE1

m, 2 dd, 1 dd juv. — Stn 1453, 20°55'S, 167°02'E, 21-

30 m, 2 dd, 2 dd sub. — Stn 1454, 20°57S, 167°02'E,

15-18 m, 9 dd. — Stn 1457, 20°47'S, 167°03'E, 5-10 m,

9 dd.

Fiji MUSORSTOM 10: stn DW1381, 187185,

177°54'E, 275-430 m, 10 dd, 9 dd juv. - BORDAU I:

stn DW1451, 16°45'S, 180°00'E, 400-460 m, 4 dd. —

Stn DW1469, 19°40'S, 178°10'W, 314-377 m, 2 dd. —

Stn DW1494, 18°55'S, 178°29'W, 240-319 m, 1 dd. —

SUVA 2: stn BS18, 18°11'S, 178°28'E, 83 m, 1 dd.

Solomon Islands. SALOMON 1: stn DW1767,

8°19'S, 160°40'E, 98-200 m, 9 dd, 9 dd sub.

Philippines. PANGLAO 2004: stn B42, O9°37N,

123°46'E, 30-33 m, 1 Iv.

Vanuatu. SANTO 2006: DB25, 15°38'S, 167°1l'E,

lOm l'add:

Wallis Island. MUSORSTOM 7:

19 219/S 176 TV 550 Grid

Marquesas Archipelago. MUSORSTOM 9: stn

DR1247, 10°34'S, 138°42'W, 1150-1250 m, 1 dd. —

Stn DR1298, 8°49'S, 140°17'W, 305 m, 2 dd.

stn DW 601,

Distribution. New Caledonia, 0-190 m, lv at intertidal

level; Loyalty Islands, 4-47 m (dd); Fiji, 83-400 m

(dd); Solomon Islands, 98-200 m (dd); Philippines,

30-33 m (lv); Vanuatu, 10 m (dd); Wallis Island, 350

m (dd), Marquesas Archipelago, 305-1150 mm (dd).

Remarks. The main characteristics of this species are:

+ height up to 3 mm, width up to 2 mm;

+ protoconch sunken, lower than the first whorl;

+ teleoconch of about 4 convex whorls,

cyrtoconoidal in shape, with 3 granular spiral cords on

penultimate whorl; moderately elevated spire; rounded

periphery; strong axial folds on first whorl, distance

between folds 2x width of them; P3 at first whorl, P2

at second whorl, PI at half third whorl; P2 and P3 the

strongest; distance between cords 2x width of cords;

P4 peripheral; the three abapical cords producing keel

on last whorl; beads of cords possibly rather sharp;

axial threads thickening to become very thick on last

whorls;

+ sculpture initially cancellate, becoming strongly

foveolate on last whorls;

+ base convex, with 3 thick spiral cords, the two

outermost strong and thick, the innermost one very

weaker, bordering umbilicus; distance between cords

about 1.5x to 2x width of cords;

+ aperture subcircular, without denticles inside the

outer lip;

+ rather wide, deep, funnel shaped umbilicus:

+ white.

Herbert (2012) pointed out that this species from the

Loyalty Islands is poorly known. He designated a

lectotype in the Manchester Museum type material.

Because of the ignorance of this species, there has

been probably many confusions with V. foveolatus

(A.Adams, 1853), that is much larger regarding the

same number of whorls, has apertural denticles and an

umbilicus partly covered by callus on adult shells.

C. VILVENS

V. saguili Poppe, Tagaro & Dekker, 2006 described

from the Philippines based on a single dead specimen,

is only compared in its original description to another

new Philippine species (V. abdii, Poppe, Tagaro &

Dekker, 2006), ignoring V. favosus. Clearly the

description and the figure match very well with the

lectotype figured by Herbert (2012) and it can be

concluded that V. saguili is a synonym of V. favosus.

Vaceuchelus auricatris Huang & Fu, 2015

Figs 6GH-I, Table 3

Vaceuchelus auricatris Huang & Fu, 2015: 66, pl. 1,

figs 1-3. Type locality: South China Sea, Taiwan

Strait, 220-350 m.

Material examined. Solomon Islands.

SALOMONBOA 3: stn DW2775, 9°23S, 160°57'E,

282-427 m, 1 dd.

Distribution. South China Sea, Taiwan Strait, 220-

350 m (original description); Solomon Islands, 282-

427 m (dd).

Remarks. The main characteristics of this species are:

+ height up to 8.4 mm, width up to 9 mm;

+ moderately elevated spire;

+ teleoconch cyrtoconoidal, of about 4 convex

whorls with 3 granular spiral cords on penultimate

whorl; rounded periphery;

+ 3 spiral cords PI, P2 and P3, sometimes with

already additional SI, on the penultimate whorl; S2

possible; P4 visible on the last whorl; axial threads

between cords, making a cancellate sculpture:

+ aperture subcircular to weakly elliptical, with

about 14 couple of denticles on outer lip:

+ base convex, with 4 thick spiral cords: distance

between cords about 1.5x width of cords:

+ rather wide umbilicus, partly covered by a small

parietal callus:

+ teleoconch first whorls nacreous

whorls bronze on Taiwanese samples.

Ivory, last

This single specimen from Solomon Islands,

coupled with a single record in the Philippines

(G.Poppe, personal communication) extends

greatly the distribution of this recently

described species.

Vaceuchelus cavernoiïdes n. sp.

Figs SK-S, Tables 3,4

Type material. Holotype (4.1 x 4.8 mm) MNHN (IM-

2000-32778). Paratypes: 4 MNHN (IM-2000-32779,

MNHN IM-2000-32780 as listed below).

Type locality. New Caledonia, Grand Récif de

Koumac, MONTROUZIER, stn 1318, 20°41',

164°15'E, 20-30 m.

NOVAPEX I8CHS 11): 1-67, 10 mars 2017

Material examined. New Caledonia. BATHUS 2:

stn DW717, 22°44'S. 167°17'E, 350-393 m, 1 dd sub,

| dd juv. - MONTROUZIER: stn 1260, 20°445,

165°14'E, 49-59 m, 30 dd. — Stn 1269, 20°355,

165°08'E, 15-20 m, dd, 1 dd juv. — Sin 1270,

20°45'S, 165°16.5'E, 10-35 m, 4 dd. — Sin 1271,

20°45'S, 165°08.0'E, 5-25 m, 20 dd juv. — Stn 1272,

20°50'S, 165°20'E, 10 m, 1 dd. — Sin 1273, 20°50,

165°23'E, 20m, 3 dd juv. Stn 1292, 20°225,

164°07'E, 0-1 m, 1 dd sub, 3 dd juv. — Sin 1310,

20°40'S, 164°15'E, 15m, 3 dd juv. — Stn 1312,

20°40'S, 164°15'E, 26-40 m, 1! dd, 6 dd juv. — Sin

1318, 20°4l'S, 164°15'E, 20-30 m, 4 Iv, 10 1v sub, 10

lv juv (holotype and paratypes MNHN I! and 2 IM-

2000-32779). — Stn 1319, 20°45S, 164°16E,

intertidal, 3 1v, 1 Iv juv (paratypes MNHN 3 and 4 IM-

2000-32780). — Stn 1339, 22°22'S, 166°15'E, 20 m, 1

lv. — Stn 1340, 22°21'S, 166°14'E, 30 m, 1 dd. — Sin

1352, 22°22'S, 166°16'E, 27-35, 1 dd. — Stn 1354,

22°22'S, 166°16'E, 27-37 m, 1dd sub, 2 dd juv. — Stn

1356, 22°20'S, 166°15'E, 20-23 m, 1 dd juv. — Stn

1358, 22°19'S, 166°28'E, 14 m, 1 dd. — Stn 1373,

22°20'S, 166° L3'E, 9-10 m, 1 dd.

Loyalty Islands. LIFOU 2000: stn 1419, 5 m,

20°56'S, 167°05'E, 5 m, 1 dd sub. — Stn 1434, 20°53'S,

167°08'E, 5-20 m, 1 dd. — Stn 1435, 20°555,

167°00'E, 5-30 m, 1 dd. — Stn 1454, 20°575,

167°02'E, 15-18 m, 1 1v sub. — Stn 1455, 20°57S,

167°03'E, 15-20 m, 1 Iv sub.

Vanuatu. SANTO 2006: stn DS04, 15°31',

167°14'E, 25 m, 1 dd, 1 dd juv. — Stn DBO8, 15°345,

167°13'E, 2 m, 2 dd, 1 dd juv. — Stn ZBO9, 15°41'S,

167°05'E, 5-7 m, 6 Iv, 1 dd juv. — Stn NB12, 15°33',

167°10'E, 20 m, 1 dd. — Stn LD20, 15°42'S, 167°15'E,

2-5 m, 2 dd sub. — Stn ZS22, 15°33'S, 167°10'E, 27-29

m, 3 dd. — Stn NB43, 15°36'S, 167°16'E, 6-30 m, 1 dd,

2 dd juv. — Stn DB46, 15°29'S, 167°15'E, 2-3 m, 1 dd.

— Stn AT52, 15°32'S, 167°13'E, 52-62 m, 1 Iv. — Stn

DB65, 15°26'S, 167°13'E, 13 m., 1 Iv. — Stn DB69,

15°24'S, 167°13'E, 38 m, 2 dd.

Solomon Islands. SALOMON l!:

8°20'S, 160°41'E, 295-381 m, 1 dd.

Philippines. PANGLAO 2004: stn B7, 09°36'N,

123°52'E, 4-30 m, 2 dd, 1 dd juv.

sin DW1776.

Distribution. New Caledonia, 1-350 m., Iv at 1-20 m:

Loyalty Islands, 5-15 m, 1v at 15-18 m; Vanuatu, 2-52

m, lv at 7-52 m; Solomon Islands, 295-381 m (dd):

Philippines, 4-30 m (dd).

Diagnosis. A small, wider than high, whitish

Vaceuchelus Species With a rather depressed,

cyrtoconoidal spire, a weakly subangular periphery, 4

strong, granular spiral cords on the penultimate whorl

and additional thinner cords on last whorl, the three

abapical cords making keel with strong axial ribs

between them giving a foveolate pattern, a moderately

convex base with usually 3 strong spiral cords and an

inner cord sinking into the narrow umbilicus.

C. VILVENS

Chilodontidae from the Pacific Ocean

Description. Shell of rather small size for the genus

(height up to 5.6 mm, width up to 6.3 mm), wider than

high, rather depressed, rather thick, slightly

cyrtoconoidal; height 0.8x to 0.9x width, height 1.8x

to 2.2x aperture height; narrowly umbilicate.

Protoconch about 100 um, of 1 whorl, at the same

level as first teleoconch whorl, with a thin terminal

varix.

Teleoconch up to 4.7 convex whorls, bearing 4 thick,

similar in size, granular, spiral cords on the

penultimate whorl, with additional intercalated thinner

cords on last whorl; strong axial prosocline ribs

between cords; periphery weakly subangular. Suture

visible, not canaliculated.

First whorl convex, sculptured by about 20 prosocline,

smooth, rather thick threads, interspace similar in size

to threads; P3 appearing at mid whorl, thickening

quickly. On second whorl, P3 strong, making a

median keel; P2 appearing at begin of whorl; P4

emerging from suture at mid whorl, suprasutural; axial

threads a little stronger, distance between them greater

than width of threads. P1 appearing at begin of the

third whorl; all the 4 cords similar in size at end of

whorl; axial threads becoming strong ribs, distance

between about 2x width of cords; SI appearing at end

of the whorl or at begin of fourth whorl, quickly as

thick as the other cords; interspace between ribs

strongly excavated, producing a foveolate pattern on

the whole whorl. S2 and S3 appearing at end of last

whorl, remaining thinner than the other cords. On last

whorl, P2, P3 and P4 making moderate keel.

Columella curved, slightly oblique.

Aperture oval, slightly elongated and slightly oblique;

outer lip without denticles inside, crenulated by inner

folds corresponding to external spiral cords; inner lip

flaring in its basal part.

Base moderately convex, with 3 (exceptionally 4 on

very large specimen - see paratype 1) granular,

prominent spiral cords similar to the Pi of the last

whorl and an additional inner spiral cord nearly

smooth, sinking into the umbilicus.

Umbilicus rather narrow (about 1/10 base width),

funnel shaped.

Colour of teleoconch nacreous white; primary spiral

cords with one per two to one per four brown beads;

protoconch whitish.

Operculum corneous, multispiral with central nucleus,

light brown, somewhat translucent.

paratype 3

paratype 4

1.82

Table 4. Vaceuchelus cavernoides n. sp.: Shells measurements in mm for types.

Discussion. The new species is rather close to

Vaceuchelus cavernosus (Sowerby II, 1905) from

Ceylon (nowadays Sri Lanka), of which two different

NHMUK syntypes are figured by Kaicher (1990: card

# 5781) and by Herbert (2012: figs. 70A-B). The main

differences of this species with V. cavernoides are a

greater size (height up to 7.3 mm), a higher spire with

a greater H/W ratio (about 1), primary cords not so

prominent on the last whorls and no visible Si.

There is also a huge gap between the large area where

the samples of the new species were collected and the

type locality of V. cavernosus (Sri Lanka). One can

wonder that no book about shells from India, Sri

Lanka (and also Vietnam, Thailand, Indonesian,

Australia and even Philippines) mention any

occurrence of this definitively poorly documented

species.

Figure 5 (scale bars: 1 mm).

A-J. Vaceuchelus favosus (Melvill & Standen, 1896). A-C. New Caledonia, Touho, MONTROUZIER., stn

1270, 10-35 m, 2.2 x 2.1 mm. D-F. New Caledonia, Koumac, MONTROUZIER, stn 1316, 12 m, 3.2 x 2.9 mm.

G-H. Wallis Is, MUSORSTOM 7, stn DW 601, 350 m, 3.6 x 3.1 mm. I-J. Fiji, MUSORSTOM 10, stn

DW1381, 275-430 m, 2.3 x 2.2 mm, dd.

K-S. Vaceuchelus cavernoides n. sp. K-M. New Caledonia, MONTROUZIER, stn 1318, 20-30 m. N-P.

Holotype MNHN (1M-2000-32778), 4.1 x 4.8 mm. Q-S. Paratype MNEN 1 (I1M-2000-32779), 5.6 x 6.3 mm. S-

U. LAGON, stn 1340, 30 m, 5.2 x 5.4 mm.

C. VILVENS

V. cavernoides n. Sp. resembles V. foveolatus

(A.Adams, 1853) from French Polynesia, Solomon

Islands and Philippines (figs 4K-S), of which a

NHMUK syntype 1s figured by Kaicher (1990: card #

5790), but this similar in size species has no Si, has a

more elevated spire with a greater H/W ratio (at least

about 1, often greater than 1), much more foveolate

sculpture on the whorls and a tiny, funnel shaped

umbilicus partially covered by a callus.

The new species may be compared to V. auricatris

Huang & Fu, 2015 from Taiwan Strait (figs 6H-T), but

this slightly greater species has a more elevated spire,

a spiral cord SI appearing only on last whorl, a

rounded, not elongated, aperture and a wider

umbilicus.

The new species may also be compared to V. ampullus

(Tate, 1893) from eastern and southern Australia, but

this much greater species (height up about 11 mm) has

much thicker, weaker granular spiral cords and

stronger axial ribs, and a narrower or closed

umbilicus.

Etymology. Shaped (Ancient Greek: -@ônç, suffix) —

with reference to shape of the shell, that is rather

similar to the one of Vaceuchelus cavernosus

(Sowerby IL, 1905).

Vaceuchelus phaios n. sp.

Figs 6A-G, Tables 3,5

Type material. Holotype (4.9 x 4.9 mm) MNAN (IM-

2000-32781). Paratypes: 5 MNHN (1M-2000-32782,

MNHN-IM-2000-32783 as listed below), 1 paratype

CV:

Type locality. New Caledonia, Grand Récif de

Koumac, MONTROUZIER, stn 1318, 20°41'S,

164°15'E, 20-30 m.

Material examined. New Caledonia.

MONTROUZIER: stn 1237, 20°47S, 165°14'E, 0-

1 m, 1 dd, 1 dd juv. — Stn 1251, 20°46'S, 165°13-14'E,

Figure 6 (scale bars: 1 mm).

Chilodontidae from the Pacific Ocean

6-15 m, 3 dd. — Stn 1271, 20°45'S, 165°08'E, 5-25 m,

11 lv. — Stn 1278, 20°348, 164°16E, 0-2m, 2 lv

(holotype et paratype MNHN 1 IM-2000-32782). —

Stn 1297, 20°35'S, 164°16'E, 3-7 m, 19 dd (with

paratypes MNHN 2-5 IM-2000-32783 and paratype

CV). — Stn 1299, 20°34'S, 164°13'E, 12-14 m, 1 dd, 1

dd juv. — Stn 1313, 20°39'S, 164°16'E, 33-38 m, 1 dd.

— Stn 1314, 20°40'S, 164°15'E, 30-63 m, 1 dd sub, 3

dd juv.

Loyalty Islands. LIFOU 2000: stn 1423, 20°545,

167°07'E, 12 m, 1 dd. — Stn 1451, 20°47'S, 167°07E,

10-21 m, 2 dd, 1 dd juv.

Distribution. New Caledonia, 1-33 m, 1v at 2-5 m;

Loyalty Islands, 12 m (dd).

Diagnosis. À small, as high as wide or shightly higher

than wide, reddish Vaceuchelus species with a

moderately elevated, weakly cyrtoconoidal spire, a

rounded periphery, 5 rather thin, granular spiral cords

on penultimate whorl with close axial ribs between

them, a moderately convex base with 3 (sometimes 4)

strong spiral cords and a rather wide umbilicus.

Description. Shell of rather small size for the genus

(height up to 5.4 mm, width up to 5.2 mm), as high as

wide or slightly higher than wide, moderately

elevated, moderately thick, weakly cyrtoconoïdal;

height 1.0x to 1.2x width, height 1.9x to 2.3x aperture

height; umbilicate.

Protoconch about 150 um, of 1 whorl, under the level

of first teleoconch whorl, with a very weak terminal

varix.

Teleoconch up to 4.1 convex whorls, with 3 rather

thin, similar in size, granular, spiral cords on the

penultimate whorl, with an additional, similar in size

peripheral cord and a slightly thinner cord intercalated

between the two most adapical cords on last whorl;

axial prosocline ribs between cords; periphery

rounded. Suture visible, not canaliculated.

A-G. Vaceuchelus phaios n. sp., New Caledonia, Koumac, MONTROUZIER, stn 1278, 2 m. A-C. Holotype

MNAN (IM-2000-32781), 4.7 x 4.8 mm. D-G. Paratype MNHN 1 (IM-2000-32782), 4.9 x 5.2 mm.

H-I. Vaceuchelus auricatris Huang & Fu, 2015, Solomon Is., SALOMONBOA 3, stn DW2775, 282-427 m, 6.0

x 6.3 mm.

J-L. Vaceuchelus vangoethemi Poppe, Tagaro & Dekker, 2006, paratype MNHN 5305, Philippines Mactan Is.,

Punta Engano, 5.8 x 4.6 mm.

M-R. Vaceuchelus rapaensis n. sp., French Polynesia, Australes Archipelago, RAPA 2002, stn 38, 2 m. M-O.

Holotype MNHN (I1M-2000-32784), 3.9 x 3.0 mm. P-R. Paratype MNHN (IM-2000-32785), 3.2 x 2.6 mm.

C. VILVENS

First whorl convex, sculptured by about 25 prosocline,

smooth, rather thin threads, interspace similar in size

to threads; P3 and P2 appearing near mid whorl, P4

close to end of whorl, suprasutural, partly covered by

next whorl; all cords thickening quickly. On second

whorl, PI appearing at begin of whorl; axial threads a

stronger, distance between them from 1.5x to 2x width

of threads. On third whorl, PI, P2 ad P3 similar in

size, moderately thick, P4 thinner; SI appearing

between mid and end of whorl; axial threads

becoming strong ribs; interspace between ribs

somewhat excavated, producing a foveolate pattern on

the whole whorl. S2 then S3 appearing on fourth

whorl, remaining thinner than the other cords. On last

whorl, Pi and SI making weak keel; TI appearing

between SI and P2; additional Ti possibly appearing

at end of whorl.

Chilodontidae from the Pacific Ocean

Columella only weakly curved, slightly oblique, with

a small parietal inducture.

Aperture oval, axially oriented; outer lip thickened

internally and weakly flaring, with about 15 elongated

denticles inside; inner lip a bit flaring in its basal part,

with a strong denticle at meeting point with columella.

Base moderately convex, with 3 or 4 (large

specimens) prominent spiral cords similar; innermost

cord nearly smooth, thick; other cords granular; thick

axial ribs between cords, distance between about 1.5x

width of ribs.

Umbilicus deep, moderately wide (about 1/8 base

width).

Colour of teleoconch nacreous brown to pinkish

white, with first whorl darker and possibly spiral cords

on the last whorls lighter; protoconch whitish.

Operculum corneous, multispiral with central nucleus,

light brown translucent.

paratype 2

paratype 3

Table 5. Vaceuchelus phaios n. Sp.: Shells measurements in mm for types.

Discussion. The new species is slightly resembles

Vaceuchelus cavernoides n. sp. from western Indo-

Pacific (figs 4K-S) but this shightly greater species 1s

more depressed, has much thicker, much prominent

spiral cords, a transversally elongated aperture without

denticles inside and an inner basal spiral cord sinking

into the umbilicus.

Etymology. Brown (Ancient Greek: @a1oc, &, ov) -

with reference to the background colour of the shell.

Vaceuchelus rapaensis n. sp.

Figs 6M-R, Tables 3,6

Type material. Holotype (3.9 x 3.0 mm) MNEN (IM-

2000-32784). Paratypes: 5 MNHN (1M-2000-32785),

1 paratype CV.

Type locality. French Polynesia, Australes

Archipelago, Rapa Island, north of Anatakuri Bay,

RAPA 2002, stn 38, 27°37'S, 144°18'W, 2 m.

Material examined. French Polynesia, Australes

Archipelago, Rapa Island. RAPA 2002: stn 10,

27°34'S, 144°23'W, 16-18 m, 2 dd, 1 dd sub, 3 dd juv.

= Sin 11,27 37SM4P T8 We 2m, dd Suns,

27°36'S, 144°19'W, 2 m, 1 dd juv. — Stn 19, 27°38%,

144°19'W, 3 m, 1 dd, 1 dd juv. — Stn 20, 27°35'S,

14423 W, Sun, 1LIN, 2 dd juv. = Sin 210 27875,

144°21'W, 5 m, 6 dd, 2 dd sub, 3 dd juv. — Stn 21,

27°34'S, 144°21'W, 5 m, 1 dd sub. — Stn 25, 27°38%S,

144°19'W, 3 m, 1 dd. — Stn 27, 27°39'S, 144°19'W, 6

m, 3 dd. — Stn 27, 27°39'S, 144°19'W, 6 m, 2 dd sub.

— Stn 29, 27°34'S, 144°21'W, 2-4 m, 1 dd. — Stn 32,

2135S, 14423 WW, 15-20 m1" SMS.

27°37.4'S, 144°18.4'W, 2 m, 15 Iv (with holotype and

paratypes), 3 Iv juv. — Stn 44, 27°36'S, 144°18'W, 30

m, 3 dd juv. — Stn 51, 27°36'S, 144°21'W, 1-1.5 m, 1

dd sub. — Stn 53, 27°37'S, 144°20'W, 1-5 m, 1 dd, 1

dd juv. — Stn 60, 27°37'S, 144°19'W, 1-1.5 m, 2 dd

juv. — Stn 61, 27°37S, 144°19'W, 10-15 m, 2 dd, 2 dd

juv. — Stn 67, 27°35'S, 144°22'W, 3-4 m, 4 dd sub. —

Stn 68, 27°35'S, 144°21'W, 1-4 m, 1 dd sub. — Stn 69,

27°38'S, 144°19'W, 3-4 m, 2 dd juv. — Stn 70,

27°37TS, 144°20'W, 15-20 m, 6 dd, 3 dd sub, 1 dd juv.

— Stn 81, 27°40'S, 144°19'W, intertidal zone, 1 dd.

Distribution. French Polynesia, Australes

Archipelago, Rapa Island, 1-30 m, 1v at 2-5 m.

Diagnosis. À small whitish Vaceuchelus species with

a moderately elevated, cyrtoconoidal spire, a

subangular periphery, 3 strong, granular spiral cords

on penultimate whorl, making keel with strong axial

C. VILVENS NOVAPEX ISCHS 11): 1-67, 10 mars 2017

ribs between them giving a foveolate pattern, a weakly

convex base with 3 strong spiral cords and à narrow,

sometimes closed, umbilicus .

Description. Shell of rather small size for the genus

(height up to 3.9 mm, width up to 3.0 mm), slightly

higher than wide, rather thick, cyrtoconoidal; spire

moderately elevated, height 1.0x to 1.3x width, 1.9x to

3.2x aperture height; narrowly

anomphalous.

Protoconch about 120-150 um, of 1 whorl, at the

same level as first teleoconch whorl, with a thin

terminal varix.

Teleoconch up to 4 moderately convex whorls, with 3

thick, similar in size, granular, spiral cords and an

additional thinner sutural on last whorl; strong axial

prosocline ribs between cords; subangular periphery.

Suture visible, not canaliculated.

First whorl convex, sculptured by about 40 prosocline,

smooth, thin threads; interspace between smaller than

width of threads; P3 appearing at mid whorl,

thickening quickly. On second whorl, P3 strong,

making a median Keel: P4 emerging from suture at

begin of whorl, P2 appearing at mid whorl; axial

threads of adapical part a bit stronger, distance

between similar in size to threads; threads of abapical

part between Pi much stronger, less numerous than

adapical threads. On third whorl, P3 moving

abapically, P2 more or less median; axial threads

between suture and P2 still rather thin, strongly

umbilicate or

prosocline; Pi and threads between them very strong,

making a foveolate pattern; threads becoming ribs,

only weakly prosocline and connecting beads of

cords; distance between cords smaller to cords

thickness. On last whorl, PI possibly appearing,

subsutural, rather weak; P2, P3 and P4 similar in

strength, each making keel and giving a tricarinated

shape to the whorl; distance between the three

abapical PT about 2x width of cords, distance between

abapical ribs about 2x width of ribs; SI possibly

appearing at end of whorl.

Columella more or less straight, vertical.

Aperture almost circular to weakly elliptical, without

denticles; outer lip thickened internally and weakly

flaring, largely crenulated by inner folds

corresponding to external spiral cords; inner lip flaring

in its basal part.

Base weakly convex, with 3 spiral cords similar to the

main Pi of the last whorl, the innermost one possibly

partly or even fully hidden by columellar callus:

distance between cords about 1.5w width of cords:

strong axial ribs between them, producing the same

foveolate pattern as on the last whorl.

Umbilicus narrow, usually partly covered by a narrow

callus, possibly fully closed on mature shells.

Colour of teleoconch yellowish nacreous

protoconch white.

Operculum corneous, multispiral with central nucleus,

light brown.

white:

Table 6. Vaceuchelus rapaensis n. sp.: Shells measurements in mm for types.

Discussion. The new species 1s rather close to

Vaceuchelus favosus (Melvill & Standen, 1896)

from Coral sea (figs 5A-J), but this similar in size

species has weaker and more numerous abapical

axial threads, lacks the thin, adapical, strongly

prosocline threads different from the abapical ribs

and has an open wide umbilicus.

V. rapaensis n. sp. remind a little of V. foveolatus

(A.Adams, 1853) from French Polynesia (figs 4K-

S) but this larger species for a similar number of

whorls is more depressed, has a PI appearing much

earlier, all the axial ribs similar on last whorls and

denticles in the aperture.

V. rapaensis n. Sp. may also be compared to V.

clathratus (A.Adams, 1853) from the Philippines,

but this much larger species for a similar number of

whorls is more elevated and has a very different P2,

thinner and more numerous abapical axial ribs and

a closed umbilicus.

Remarks. Apart a relative smaller size, there are no

differences between V. clathratus (A.Adams.

1853), of which two different NHMUK syntypes

are figured by Kaicher (1990: card # 5789) and by

Herbert (2012: figs. 70:C-D), and V. vangoethemi

Poppe, Tagaro & Dekker, 2006 from the

Philippines (figs 6J-L). This is probably the reason

of the unexpected lack of V. clathratus noted by

Poppe et al. (2006), although they noted that

Kosuge & Chino (1998) mentioned it. This lead to

C. VILVENS

Chilodontidae from the Pacific Ocean

conclude that V. vangoethemi has to be considered

a synonym of V. clathratus.

Etymology. Named after the type locality.

Genus Herpetopoma Pilsbry, 1890.

Type species: Euchelus scabriusculus A.Adams &

Angas in Angas, 1867 (by original designation) -

Recent, Australia.

Remarks. The main features of the Herpetopoma

species are ridge-like denticles inside the outer lip,

a basal columellar tooth and another tooth on the

basal lip near the columella, the two teeth making

an U-shaped notch (Herbert, 2012). Considering the

type species, Herbert add to this features an open

umbilicus, characterizing Herpetopoma sensu

stricto. 1 use here an extending characterization for

a Herpetopoma sensu lato in which Herbert locate

for instance H. naokoae Poppe, Tagaro & Dekker,

2006.

Herpetopoma | shell size number | Pi appearing which Si, base num- | periphe | open frontal

shape (mm) | ofcords | simulta- Ti present ? | shape ber TY umbili view

neously ? shape -Cus ?

P1 a bit later SAS 2S SE

+ Sd; T1,

T2 + poss

conical 8x7

to sl.

CyrtoCco-

gemmatum

Convex

noidal Ti

instrictum conical 12x10 | 4-5 W. roun-

COnvex ded to

sub-

angular

xeniolum conical 4.5x 5 M S2,S4 mod. 6-8 roun-

35 convex ded

pantantoi conical sub-

n. SP. angular

verruca convex roun-

cyrtoco- ded

noidal

ludiviniae si. convex

cyrtoco-

noidal

poichilum conical convex 5 sub-

angular | Y nar-

vitilevuense cyrtoco Y except P4 convex |4 roun- N

n. SP. noidal ded

naokoae SE mod. S roun-

Cyrtoco- convex ded

noidal |

|Hivaoaense conical 4x3 SR N: P3 + P2 + SIES2) W. 3 sub-

ñ: Sp} PI + P4 convex angular

corrugatum conical 3.8x Y

to sl. 3.1

Cyrtoco-

noidal

rubrum cyrtoco- | 8.5x8 Y

noidal

5 M NS2: mod. 3 sub-

1 convex angular

8 P1 later SI SPMSE): COnvex 4-5 roun-

+ S4; T1, ded

Table 7. Comparison of conchological features of small central Indo-Pacific Herpetopoma s. L species (sl. =

slightly, mod. = moderately, poss. = possible, w = weakly).

C. VILVENS

Herpetopoma gemmatum (Gould, 1845)

Figs 7A-K, Table 7

Trochus gemmatus Gould, 1845: 27, pl.15, fig.10.

Type locality: Sandwich Islands (nowadays Hawaïian

Islands).

Monodonta fischeri Montrouzier in Souverbie &

Montrouzier, 1866: 142, pl. fig. 7. Syn.

Herpetopoma fischeri (Montrouzier in Souverbie &

Montrouzier, 1866) — Kaicher, 1990: card #5704. Syn.

Euchelus gemmatus (Gould, 1845) — Jansen, 1994: 63,

pl.1, figs. 5-6.

Herpetopoma fischeri (Montrouzier in Souverbie &

Montrouzier, 1866) — Herbert, 1996: 411-412, figs. 1-

Herpetopoma gemmata (Gould, 1845) — Vilvens &

Héros, 2003: 62, fig.6.

Euchelus gemmatus — Kay, 1979: 51, figs 13, 14B.

Herpetopoma fischeri (Montrouzier in Souverbie &

Montrouzier, 1866) — Vilvens & Héros, 2003: 62,

fig7.

Herpetopoma gemmatum — Severns, 2011: 44, pl. 6,

fig 5.

Herpetopoma gemmatum — Herbert, 2012: 433, 439.

Material examined. New Caledonia. CHALCAL:

stn DW60, 21°49'S 159°28'E, 45 m, 1 dd. - CORAIL

2: stn DW84, 19°12'S 158°57'E, 16-26 m, 1 dd sub. —

BATHUS 1: stn DW692, 20°35'S, 164°59'E, 140-150

m, 2 dd. — Stn DW1233, 22°24'S, 166°48'E, 45-50 m,

1 dd sub. — MONTROUZIER: stn 1237, 20°47'S,

165°14'E, 0-1 m, 2 dd juv. — Stn 1242, 20°46.2S,

165°14.5'E, 0-2 m, 1 dd sub. — Stn 1238, 20°47S,

165°13'E, intertidal zone, 2 1v. — Stn 1254, 20°465,

165°15'E, 7 m, 4 dd sub, 3 dd juv. — Stn 1255,

20°43'S, 165°08'E, 11 m, 20 dd sub, 10 dd juv. — Stn

1259, 20°45'S, 165°14'E, 15-35 m, 9 dd sub, 9 dd juv.

— Stn 1259, 20°45'S, 165°14'E, 15-35 m, 4 dd juv. —

Stn 1266, 20°40'S, 165°15'E, 10-15 m, 1 dd. — Stn

1268, 20°45'S, 165°08'E, 9-11 m, 1 dd sub. — Stn

1268, 20°45'S, 165°08'E, 9-11 m, 15 dd sub, 1 dd juv.

— Stn 1269, 20°35'S, 165°08'E, 15-20 m, 20 dd. — Stn

1270, 20°45'S, 165°17'E, 10-35 m, 4 dd, 29 dd sub. —

Stn 1271, 20°45'S, 165°08'E, 5-25 m, 19 dd. — Stn

1272, 20°50'S, 165°20'E, 10 m, 10 dd sub, 20 dd juv.

— Stn 1273, 20°50'S 165°23'E, 20 m, 1 dd, 9 dd sub. —

Stn 1284, 20°34'S, 164°1 l'E, intertidal zone, 5 Iv sub,

25 Iv juv. — Stn 1289, 20°29'S, 164°10'E, intertidal

zone, 3 dd, ! dd juv. — Stn 1291, 20°22'S, 164°07'E,

intertidal zone, 1 dd sub. — Stn 1298, 2-4 m, 20°35'S,

164°17'E, 2 dd sub. — Stn 1316, 20°40'S, IGYIT'E,

12 m., dd, 5 dd sub, 15 dd juv. — Stn 1318, 20°41'S,

164°15'E, 20-30 m, 1! dd, 4 dd sub, 3 dd juv. — Stn

1319. 20°45'S, 164°15.5'E, intertidal zone, 10 dd, 1 dd

juv. — Stn 1331, 55-57 m, 20°40'S, 164°1 l'E, 1 dd sub.

_ Stn 1346, 22°22S, 166°16'E, 5-6 m., 1 dd, 1! dd sub.

_ Stn 1350, 22°18'S. 166°20'E, 3-6 m, 1 dd, 3 dd sub,

1 dd juv. — Stn 1352, 22°22'S, 166°16E, 27-35, 1 dd,

3 dd sub, 10 dd juv. — Stn 1354, 22°22S, 166°16E,

27-37 m. 4 dd sub, 8 dd juv. — Stn 1357, 22°30'5,

NOVAPEX IS8CHS 11): 1-67, 10 mars 2017

166°26'E, 25-35 m, 1 dd. — Stn 1362, 22°215,

166°25'E, 6-8 m, 2 dd sub. — Stn 1371, 22°20$,

166°18'E, 12-16 m, 1 dd, 5 dd sub, 4 dd juv. — Stn

1373, 22°20'S, 166°13'E, 9-10 m, 1 dd. - PALEO-

SURPRISE: stn DW1381, 18°29'S, 163°04'E, 32-36

m, 1 dd. —- NORFOLK 1: sin DW1727, 23°175,

168°14'E, 190-212 m, 1 dd juv.

Loyalty Islands. LIFOU 2000: stn 1413, 20°55'S,

167°05'E, 3-10 m, 2 dd. — Stn 1420, 20°4855,

167°09'E, 4-5 m, 3 dd juv. — Sin 1421, 20°525,

167°09'E, 4 m, 1 dd. — Sin 1422, 20°47'S, 167°07E,

4 m, 2 dd. — Stn 1423, 20°54'S, 167°07'E, 12 m, 2 dd,

3 dd juv. — Stn 1424, 20°55'S, 167°03'E, 4 m, 1 dd. -

Stn 1429, 20°485'S, 167°07'E, 8-18 m,3 dd, 1 dd sub.

— Sin 1432, 20°54'S, 167°03'E, 12-32 m, 3 dd. - Stn

1434, 20°53'S, 167°08'E, 5-20 m, 2 dd, 1 dd juv. — Sin

1435, 20°55'S, 167°00U'E, 5-30 m, 3 dd, 6 dd juv. — Sin

1436, 20°56'S, 167°04'E, 10-20 m, 2 dd. — Sin 1444,

20°55'S, 167°05'E, 9-20 m, 1 dd. — Stn 1449, 20°465,

167°02'E, 17m, 1 dd juv. — Stn 1450, 20°465,

167°02'E, 27-31 m, 2 Iv juv. — Stn 1451, 20°475,

167°07'E,10-21 m, 2 dd, 1 dd juv. — Stn 1453,

20°55'S, 167°02'E, 21-30 m, 1 dd. — Stn 1453,

20°55'S, 167°02'E, 21-30 m, 2 dd juv. — Stn 1454,

20°57'S, 167°02'E, 15-18 m, 1 dd juv. — Stn 1456,

20°49'S, 167°10'E, 25-30 m, 1 dd juv. — Stn 1457,

20°47'S, 167°03'E, 5-10 m, 3 dd, 1 dd juv. — Sin 1461,

20°54'S, 167°02'E, 100-120 m, 1 dd. — Sin DW1650,

20°54'S, 167°02'E, 120-250 m, 1 dd.

Fiji. SUVA 4: DWO9, 18°21'S, 178°06'E, 37-41 m, 2

dd. — Stn DWI10, 18°2l'S, 178°08'E, 39-43 m, 1 dd

sub. — Sin DW12, 18°21'S, 178°10'E, 39 m, 4 dd sub.

— Stn DW16, 18°26'S, 178°07'E, 32-36 m, 2 dd. — Stn

DW22, 18°22S, 177°59'E, 32-36 m, 3 dd. — Stn

DW25, 18°27%S, 178°0l'E, 48-51 m, 2 dd. — Stn

DW74, 17°49'S, 177°12'E, 38 m, 1 dd.

Wallis and Futuna Islands. MUSORSTOM 7: sin

DW494, 14°19'S 178°03'W, 100-110 m, 1! dd. — Stn

DW601, 13°19'S, 176°17'W, 350 m, 1 dd sub, 1 dd

juv.

French Polynesia, Australes Archipelago.

BENTHAUS: DW1961, 23°21'S, 149°34'W, 470 800

m, 2 dd sub.

Vanuatu. SANTO 2006: stn DB6S, 15°265,

167°13'E, 13 m, 1 dd.

Solomon Islands. SALOMON ]I: sin DW1742,

11°29'S, 159°57'E, 366-421 m, 1 dd juv. — Stn

DW1762, 8°40'S, 160°04'E, 396-411 m, 1 dd juv. —

Stn CP1764, 8°37'S, 160°07'E, 1327-1598 m., 1 dd. -

SALOMON 2: stn DW2198, 7°43'S, 158°30' E, 273-

300 m, 1 dd.

Indonesia, Tanimbar Islands. KARUBAR: stn

DW49, 08°00'S, 132°59'E, 206-210 m.1 dd.

Philippines. PANGLAO 2004: stn B10, 09°37N.

123°46'E, 3-14 m, 1 dd, 1 dd juv. — Stn S13, 09°37'N,

123°52'E, 8-15 m, 2 dd. — Stn B38. 09°3]'N.

123°4l'E, 17-18 m, 2 dd. —- PANGLAO 2005: stn

DW2340, 09°29'N, 123°44'E, 271-318 m, 1 dd juv. —

Stn DW2401, 09°3l'N, 123°40'E, 397-410 m. 2 dd.

C. VILVENS

Distribution. New Caledonia, 1-190 m, 1v at 0-1 m:;

Loyalty Islands, 4-120 (dd); Fiji, 36-48 m (dd); Wallis

and Futuna Islands, 110-350 m (dd); French

Polynesia, Australes Archipelago, 800 m (dd);

Vanuatu, 13 m (dd); Solomon Islands, 300-1327 m

(dd); Indonesia, Tanimbar Islands, 206-210 m (dd):

Philippines, 14-397 m (dd).

Remarks. The main characteristics of this species are:

+ height up to 8 mm, width up to 7 mm;

+ conical, with a rather elevated spire;

+ teleoconch of about 5 convex whorls with at least

8 granular, rather thin, beaded spiral cords on

penultimate whorl, up to 10-12 on last whorl; beads of

cords rounded and small;

+ first whorl with prosocline, thick axial threads; P2

and P3 after mid first whorl, PI at begin of 2" whorl:

S1 and S2 at 3 whorl; S3, T1 and T2 at 4° whorl, T1

between PI and SI and T2 between S1 and P2; P4 and

S4 visible on last whorl; additional Ti cords possibly

appearing;

+ columella with one strong basal columellar tooth

with a second weaker upper tooth; an other tooth on

basal lip, a deep U-shaped notch between the two

teeth; a basal glossy expansion bordering the aperture

bearing short ridges.

+ denticles inside the outer lip of the aperture;

+ 4, sometimes 5, spiral cords on the base;

+ moderately wide umbilicus, with perspective to

apex, partly covered by a callus:

+ whitish with reddish spots.

I follow here Jansen (1994) and Herbert (1996) who

consider that Herpetopoma fischeri (Montrouzier in

Souverbie & Montrouzier, 1866) is a likely synonym

of H. gemmatum. 1 can't agree with Poppe et al. (2006)

who, ignoring apparently Æ. gemmatum, consider H.

Jischeri as a synonym of H. exasperatum (A. Adams,

1853) from the Philippines (maybe by comparison

with juvenile or subadult samples of A. fischeri = H.

gemmatum). For a similar size, H. exasperatum :s

indeed clearly much more depressed than A.

gemmatum, With P2 and P3 producing a double Kkeel,

giving a cylindrical shape to the abapical part of the

body whorl, instead of the convex shape of }H.

gemmatum Whorls; moreover, H. gemmatum has

Chilodontidae from the Pacific Ocean

similar in size spiral cords on the body whorl, instead

of spiral cords of alternating size.

Herpetopoma instrictum (Gould, 1849)

Figs 7L-T, Table 7

Trochus (Monodonta) instrictus in 1846-50: 107

(1852: 190, pl. 13, fig. 225a-c). Type locacility:

Pacific Islands.

Monodonta alveolata A.Adams, 1853: 176. Syn.

Monodonta bourcierei Crosse, 1863: 178, pl. 4, fig. 6.

Syn.

Euchelus instrictus — Kaicher, 1990: card # 5705.

Euchelus alveolatus — Kaicher, 1990: card # 5706.

Euchelus bourcieri — Kaicher, 1990: card # 5707. Syn.

(misspelling for bourcierei).

Herpetopoma instricta — Wilson, 1993: 68.

Herpetopoma instricta — Vilvens & Héros, 2003: 61,

fig. S.

Herpetopoma instrictum — Poppe, Tagaro & Dekker,

2006: 37, pl. 10, fig. 4.

Herpetopoma instrictum — Herbert, 2012: 429-433,

fig. 23:A-H.

Material examined. New Caledonia.

MONTROUZIER: sin” 1245, 207455, 165416 E.

intertidal, 24 dd, 2 dd sub, 7 dd juv. — Stn 1254,

20°46'$S, 165°15E, 7m, 1 'dd. = Sn 1270/2045,

165°15'E, 10-35 m, 1 dd, 1 dd sub, 2 dd juv. — Stn

1271, 20°45'S, 165°08'E, 5-25 m, 1 dd. — Sin 1272,

20°50'S 165°20'E, 10m, 1 dd sub. — Stn 1279,

20°35'S, 164°16'"E, intertidal zone, 1 dd, 2 dd juv. —

Stn 1284, 20°34'S, 164°11'E, intertidal zone, 3 dd. —

Sin 1297, 205$, 16416 6-7 4édd sub, Sin

1316, 20°40$S, LEE 12m, Itddsub:

Loyalty Islands. BIODGEOCAL: stn DW307, 20°35'S,

166°55'E, 470-480 m, 5 dd. —- BATHUS 4: stn

DW929;, 187525; 163 23E/ 6025160 dde

LIFOU:2000::stn 1455, 20555; LGTPOOUE, 5-30 1

dd juv. — Stn 1438, 20°48'S, 167°09'E, 16 m, 1 dd. —

Stn 1436, 20°56'S, 167°04'E, 10-20 m, 1 dd juv. — Stn

1441, 20°46'S, 167°02'E, 20 m, 1 dd juv.

Distribution. New Caledonia, 1-12 m (dd); Loyalty

Islands, 16-502 m.

Figure 7 (scale bar: 5 mm).

A-K. Herpetopoma gemmatum (Gould, 1845). A-C. New Caledonia, Koumac, MONTROUZIER, stn 1319, 7.9

x 6.9 mm. D-I. Loyalty Islands, Lifou, LIFOU 2000, stn 1429, 8-18 m. D-F. 7.3 x 6.2 mm. G-I. Juvenil, 5.0 x

5.4 mm. J-K. Broken specimen showing internal structure, New Caledonia, Touho, MONTROUZIER. stn1 266.

015 ni 66 77x5.6 mm.

L-T. Herpetopoma instrictum (Gould, 1849), New Caledonia. L-N. Koumac, MONTROUZIER, stn 1284,

intertidal, 8.9 x 7.5 mm. O-Q. Touho, MONTROUZIER, stn 1254, 7 m, 9.2 x 7.7 mm. R-T. Subadult. Touho.

MONTROUZIER, stn 1272, 10 m, 6.7 x 6.6 mm

C. VILVENS

Chilodontidae from the Pacific Ocean

Remarks. The main features of this species are:

+ height up to 12 mm, width up to 10 mm;

+ spire rather elevated, conical; sculpture foveolate;

+ protoconch of about 180-200 um;

+ teleoconch of up to 6.5 almost flat to weakly

convex whorls; 4 to 5 granular spiral cords on the

penultimate whorl, 6 on the last whorl; first whorl

with rather strong prosocline threads, distance

between threads similar in size to threads; P2 and P3

appearing at nud first whorl, PI at end of first whorl,

SI at half of third whorl, T1 at half of fourth whorl,

between PI and S1; on last whorl, P4 peripheral, cords

similar in size except P3 and P2 slightly stronger;

distance between cords 1x to 1.5x size of cords;

+ columella with a low bulge in the mid region and a

small, peg-like, basal tooth;

+ aperture D-shaped, with a dozen ridge-like

denticles; margin of outer lip somewhat flaring; U-

shaped notch between the basal tooth and first denticle

of basal part of outer lip;

+ base weakly convex to almost flat, with 5 to 6

strong, granular spiral cords, the innermost forming

umbilical margin; distance between cords similar to

size of cords;

+ large umbilicus, funnel shaped with convex wall:

+ light brown with reddish brown markings.

All the New Caledonian samples are dead taken; many

of them lack the labral denticles in the aperture except

one or two weak ones near the columella.

Herpetopoma xeniolum (Melvill, 1918)

Figs 8A-T, Table 7

Euchelus xeniolum Melvill, 1918: 154, pl. 5, fig. 27.

Type locality: Chabar, Gulf of Oman [= Chah Bahar,

Iran|,

Turcica (Perrinia) waiwailevensis Ladd, 1982: 23, pl.

24, figs 10-13. Syn.

Herpetopoma eboreum Vilvens & Héros, 2003: 61-64,

figs. 1-4. Syn.

Herpetopoma eboreum — Poppe, Tagaro & Dekker,

2006: 36, pl. 10, fig.2.

Herpetopoma xeniolum — Herbert, 2012: 444-446, fig.

36, 37:A-H, 38:A-C. )

Material examined. New Caledonia. LAGON: stn

830, 20°49'S, 165°19'E, 105-110 m, 50 dd. -BATHUS

1: stn DW692, 20°35'S, 164°59'E, 140-150 m, 2 dd.

Es MUSORSTOM 10: *CP1576, 18195178 09E,

497-504 m, 1 dd.

Tonga Islands. BORDAU 2: stn DW1567, 20°025,

175°19'W, 351-356 m, 2 dd.

Vanuatu. SANTO 2006: stn DS99, 15°33%,

lOTTE 100-105" Lrdd Sin DSID03 15% 111$.

167°16'E, 70-80 m, 1 dd.

Indonesia. KARUBAR: stn DW29, 05°365,

132°56'E, 181-184 m, 5 dd. — Stn DW49, 08°005,

132°59'E, 210-206 m, 1 1v.

Solomon Is. SALOMON 1: stn DW1767, 8°195,

160°40'E, 98-200 m, 3 dd. — Stn DWI1850, 10°28'5,

161°59'E, 139-261 m, 1 dd.

Philippines. AURORA 2007: stn DW2738, 16"04N,

121°56'E, 111-113 m, 2 dd. — Sin DW2757, 15°34%,

121°50'E, 169-176 m, 1 dd, 3 dd sub.

Distribution. New Caledonia, 110-140 m (dd); Fiji,

497-504 m (dd); Tonga Islands, 351-356 m (dd);

Vanuatu, 80-100 m (dd); Indonesia, Kai Islands, 184-

210 m (dd); Solomon Islands, 139-200 m (dd);

Philippines, 113-169 m (dd).

Remarks. The main characteristics of this species are:

+ height up to 4.5 mm, width up to 3.5 mm;

+ celevated, spire rather thick, roundly conical;

sculpture cancellate;

+ protoconch of about 150 um;

+ teleoconch of 4.5 to 5 convex whorls with 5

granular spiral cords on penultimate whorl: rounded

periphery; on 1° whorl, only thin prosocline threads,

distance between threads about 1.5x larger than

threads; at begin of 2"! whorl, 5 spiral cords (4 Pi and

S2) resolving almost simultaneously (one can detect

on some specimens P1 dividing into 2 cords but this is

visible on other samples); S4 peripheral on last whorl,

thinner than the other cords; distance between cords

smaller than cords ;

+ columella with one basal, small but sharp tooth:

+ moderately convex base with 6 to 8 rather thick,

nearly smooth to weakly subgranular, spiral cords;

interspace between cords as broad as cords:

+ strong labral folds all around the outer lip of the

aperture;

+ no umbilicus;

+ light beige to ivory, without maculation.

Herpetopoma pantantoi n. sp.

Figs 8J-O, Tables 7, 8

Type material. Holotype (4.1 x 4.8 mm) MNHN (IM-

2000-32786). Paratypes: 4 MNHN (IM-2000-32787),

1 paratype CV.

Type locality. Southern New Caledonia, Norfolk

Ridge, NORFOLK 1, stn DWI1684, 24°55%,

168°22'E, 508-541 m.

Material examined. New Caledonia. BIOCAL: stn

KGU6,20735S, 166°53:E; 735 1m, 2'dd. = SDS:

20°34'S, 166°54'E, 435 m, 3 dd. — Stn DW83,

20°35'S, 166°54'E, 460 m, 1 dd. - CHALCAL 2: stn

DW72, 2455 S 168228; 527m, l'dd. -SMIB 5°°m

DW17, 23 41S 167 S9E: 022% mn Loue

VOLSMAR: stn DW38, 22°22'S, 168°44'E, 380-420

m, 1 dd. - BERYX 11: stn DW10, 24°53'S, 168°21'E.

565-600 m, 6 dd. — Stn DW35, 23°33'S, 167°16'E.

550-570 m, 12 dd. — Stn CH41, 23°39'S, 168°00'E.

230-360 m, 7 dd. — SMIB 8: stn DW146-147, 24°55'S.

168°22'E, 508-532 m, 4 dd. — Stn DW153, 24°54'S.

C. VILVENS

NOVAPEX I18CHS 11): 1-67, 10 mars 2017

À À mine

168°22'E, 547-560 m, 1 dd. — Sin DW166, 23°385S,

I67°43E, 433-450 m, 4 dd. - Sin DW193-19%6,

23°59'S, 168°21'E-168°23'E, 491-558 m, 11 dd, 1 dd

sub, [ dd juv. — Sin DW201, 23°59'S, 168°2l'E, 500-

504 m, 1! dd. - BATHUS 1: stn CP654, 21°175,

165°5TE, 237-298 m, 1 dd, 2 dd sub, 5 dd juv. -

DW838, 23°0l'S, 166°56E, 400-402, 1 dd. -

NORFOLK 1: stn DW1666, 23°42'S, 167°44'E, 469-

860 m, 5 dd. — Sin DW1684, 24°55'S, 168°22'E, 508-

541 m, 7 Iv (with holotype and paratypes). — Stn

DW1692, 24°56'S, 168°21'E, 507-967 m, 1 dd. — Sin

DW1732, 23°20'S, 168°16'E, 347-1063 m, 8 dd, 3 dd

sub, 3 dd juv. — Sin DW1733, 23°56'S, 167°15'E, 427-

433 m, ! dd. -NORFOLK 2: stn DW2036, 23°38'5,

167°39'E, 571-610 m, 14 dd, 4 dd sub. — Stn DW2057,

24°40'S, 168°39'E, 555-565 m, 1 lv. —- EBISCO: stn

DW2603, 19°36'S, 158°43'E, 568-570 m, 12 dd.

Loyalty Islands. BIDGEOCAL: stn DW291, 20°34'S,

166°54E, 510-520 m, 1 dd, 1 dd sub, 1 dd juv. — Sin

DW307, 20°35'S, 166°55'E, 470-480 m, 2 dd sub. —

Stn DW253, 21°32'S, 166°29'E, 310-315 m, 2 dd sub.

Fiji MUSORSTOM 10: stn DW1334, 16°515,

178°14'E, 251-257 m, 1 dd juv

Solomon Islands. SALOMONBOA 3:

10°27'S, 162°02'E, 240-572 m, 15 dd.

stn CP2823,

Distribution. New Caledonia, 238-737 m, 1v at 541!-

555 m; Loyalty Islands, 315-510 m (dd); Fiji,

251-257 m (dd); Solomon Islands, 240-572 m (dd).

Diagnosis. A small light brownish Herpetopoma

species with an elevated, conical spire and a

subangular periphery, 4 granular spiral cords on the

whorls, a strong columellar basal tooth, strong fold-

shape denticles inside the aperture and an almost flat

base with 5 spiral cords and without umbilicus.

Description. Shell of rather small size for the genus

(height up to 5.6 mm, width up to 6.3 mm), higher

than wide with a subangular periphery, rather thick,

conical; spire elevated, height 1.2x to 1.3x width, 2.3x

to 4.3x aperture height; anomphalous.

paratype 3

paratype 4

paratype 5 CV

Protoconch about 150-200 um, of 1 whorl, with a thin

terminal varix.

Teleoconch up to 5.6 weakly convex to almost straight

whorls, bearing 4 spiral granular cords similar in size,

with an additional smaller peripheral cord on last

whorl; nodules from cords at intersections with strong

axial prosocline ribs. Suture hardly visible, weakly

canaliculated.

First whorl convex, sculptured by about 20 prosocline

smooth, rather thin ribs; interspace between similar in

size to ribs. On second whorl, primary cords PI, P2,

P3 and P4 appearing after mid whorl, granular, similar

in size; distance between cords similar in size 10

cords; axial ribs stronger, connecting beads of cords.

On third whorl, cords and ribs much stronger,

producing a reticulate pattern; distance between cords

smaller to cords thickness.

On next whorls, cords and ribs very strong, producing

an almost foveolate pattern; sulcus at sutural level,

made by subsutural cord and suprasutural cord of

preceding whorl: beads of cords slightiy axially

elongated.

On last whorl, S4 emerging from suture, thinner than

the other cords; P4 peripheral, slightly stronger then

other Pi; additional cords possibly appearing, e.g. T1

under PI and next T2 ender P2 (holotype) or T2 and

T3 respectively under P2 and P3 (paratype 1).

Columella straight, almost vertical, with a strong,

basal tooth; tooth 45° toward bottom oriented.

Aperture almost circular; outer lip thickened internally

and weakly flaring, especially near the columella; 9 or

10 inner fold-shaped labral denticles, innermost

separated from basal columella tooth by a U-shaped

notch.

Base almost flat, with 5 subgranular to nearly smooth.

similar in size spiral cords: distance between cords

slightly greater than cords: thin axial threads between

Spiral cords.

No umbilicus.

Colour of teleoconch pinkish white to light brown :

protoconch pinkish white.

Operculum corneous, multispiral, brown.

Table 8. Herpetopoma pantantoi n. Sp.: Shells measurements in mm for types.

Discussion. The new species is rather close to

Herpetopoma xeniolum (Melvill, 1918) from Indo-

Pacific (figs 8A-I), but this species has more convex

whorls, 5 (not 4) spiral cords on penultimate whorl, Pi

appearing at the latest at begin of 2" whorl (not after

the mud whorl), a smaller basal columellar tooth, a

C. VILVENS

Chilodontidae from the Pacific Ocean

more rounded periphery and a more convex base with

more numerous spiral cords (6 to 8 instead of at most

5).

H. pantantoi n. sp. may also be compared to H.

naokoae Poppe, Tagaro & Dekker, 2006 from the

Philippines (figs 10G-L), but this species has more

convex whorls, a smaller protoconch, P2, P3 and P4

appearing at begin of 2" whorl while PI appears later

on 3 whorl, granular, a rounded periphery and a more

convex base with the distance between cords smaller

then cords.

The new species reminds also H. norfolkensis Jansen,

1994 from Norfolk Island (off western Australia), but

this similar in size species is more or less as high as

wide (H/W ratio from 0.95 to 1.05), has a

cyrtoconoidal shape, much more convex last whorls, a

different ontogeny of the cords (P3 and P4 first, next

PI and finally P2), a more rounded periphery and only

three spiral cords on the base

Etymology. Cords together (Ancient Greek: together

= noavta and cord = avtoû, o1 - contracted) - with

reference to the spiral cords appearing simultaneously

on the second whorl.

Herpetopoma verruca (Gould, 1861)

Figs 9SA-U, Table 7

Diloma verruca Gould, 1861: 18. Type locality: Coral

Sea, China.

Herpetopoma verruca — Herbert, 2012: 439, fig. 69:

E-F.

Material examined. Indonesia, Kai Islands.

KARUBAR:" Sin "DWIS, "212-221 "1m, "OSIS,

132°41'E, L dd. — Sin DW18, 05°18'S, 133°01L'E, 205-

212 m, 5 dd. — stn DW44, 07°52'S, 132°48'E, 291-295

m, 3 dd. — Stn DW49, O8°00'S, 132°59'E, 206-210 m,

Ædd. = Sin DWS0, OTSOSMIS3 02; 184186" 1

dd.

Solomon Islands. SALOMON 1: stn DW1762,

8°40'S, 160°04'E, 396-411 m, 17 dd, 5dd juv. —

SALOMON 2: stn DW2191, 6°51'S, 156°24'E, 182-

277 m, 20 dd. — Sin DW2234 6°51$S, 156°24E, 182-

Figure 8 (scale bar: 1 mm).

277 m. 18 dd. — Sin DW2235, 6°51/5, 156°23E, 162-

196 m, | dd.

Vanuatu. SANTO 2006: DR89, 15°34'S, 167°17E,

751; l'dd = Sn DS99, 157338 /167°17ÆE/100-108

m, 2 dd.

New Caledonia. LAGON: stn 830, 20°49%5,

165°19'E, 105-110 m, 35 dd. - MONTROUZIER: stn

1321, 20°457'S, 164°15'E, 90-115m, 1 dd. — Stn

1323, 20°41'S, 164°15'E, 82-120 m, 3 dd. - BATHUS

1: stn DW692, 20°35'S, 164°59'E, 140-150 m, 2 dd.

Loyalty Islands. MUSORSTOM 6: sin DW406,

D0P4ANS 167 O0TE 573 01 l'ad,

Fiji. BORDAU 1: stn DW1465, 18°09'S, 178°39'W,

290-300 m, 1 dd. — Stn DW1469, 19°.40'S, 178°10'W,

314-377 m, 4 dd.

South-western Pacific, Wallis Island.

MUSORSTOM 7: sin'DW6O01, 137208 TIGE,

350 m, 1 dd. — Stn DW610, 13°21'S, 176°09'W, 286

m, 1 dd, 1 dd sub.

Tonga. BORDAU 72: stn

175°00'W, 300-302 m, 2 dd.

DW1523,. 217185,

Distribution. Indonesia, Kai Islands, 186-291m (dd);

Solomon Islands, 196-396 m (dd); Vanuatu, 75-100 m

(dd); New Caledonia, 110-140 m (dd); Loyalty

Islands, 373 m (dd), Fiji, 300-314 m (dd); Wallis

Island, 286-350 m (dd); Tonga, 300-302 m (dd).

Remarks. The main characteristics of this species are:

+ height up to 3.2 mm, width up to 2.7 mm:

+ moderately elevated spire; slightly cyrtoconoïidal;

sculpture cancellate;

+ teleoconch of about 4.5 convex whorls with 3

granular spiral cords on penultimate whorl; rounded

periphery; P2 and P3 appearing first, P1 a bit later; on

last whorl, P2 and P3 widely spaced, S1 and S2

appearing and S4 peripheral,;

+ aperture With denticles all around outer lip;

+ columella with one small basal tooth with U-

notch below;

+ convex base with 4 granular spiral cords; distance

between cords 1x to 1.5x the size of cords;

+ no umbilicus:

+ whitish.

A-I. Herpetopoma xeniolum (Melvill, 1918). A-C. New Caledonia, Poindimié area, LAGON, stn 830, 105-110

m, 4.2 x 2.8 mm. D-F. Vanuatu, SANTO 2006, stn DS99, 100-105 m, 3.5 x 2.4 mm. G-I. Tonga Islands,

BORDAU 2, stn DW1567, 351-356 m, 4.7 x 3.4 mm.

J-O. Herpetopoma pantantoi n. Sp., southern New Caledonia, Norfolk Ridge. J-L. Holotype MNHN (IM-2000-

32786), NORFOLK 1, stn DW1684, 508-541 m, 5.2 x 4.2 mm. M-O. SMIB 8, stn DW201, 500-504 m, 5.5 x 4.4

mm.

P-T. Herpetopoma poichilum Vilvens, 2012, BENTHAUS, stn DW1876, 150-160 m. P-R. 5.4 x 3.6 mm. S-T.

4.6 x 3.8 mm.

C. VILVENS

Chilodontidae from the Pacific Ocean

Herbert (2012) pointed out that this species, described

from "Coral seas, China”, is poorly Known and that the

two widely spaced main spiral cords are distinctive.

Using the little available information about this

species, it seems difficult to distinct the numerous

samples from the lectotype of the USNM: at most can

be said that the spiral cords of the whorls seem thinner

for some samples than those of the Coral sea type.

Considering this impossibility to differentiate the

specimens only on a conchological base,

Herpetopoma verruca seems to be the oldest available

name to gather all these samples from Solomon

Islands to New Caledonia and even to Tonga Islands.

In this last area, the only two sample collected are a

little stockier: strangely, the columellar tooth is only

fully visible from a three-quarters view.

Herpetopoma ludiviniae

(Poppe, Tagaro & Dekker, 2006)

Figs 9V-X, Table 7

Vaceuchelus ludiviniae Poppe, Tagaro & Dekker,

2006: 48-49, pl. 17, fig. 1-3. Type locality: Punta

Engaño, Mactan Island, Philippines.

Material examined. Philippines. PANGLAO 2004:

stn P1, 9°36'N, 123°45'E, 90-200 m, 1 dd, 6 dd sub, 2

dd juv. — Stn L50, 9°37'N, 123°46'E, 120 m, 2 dd. —

AURORA 2007: stn DW2741, 16°03'N, 121°54'E,

194-203 m, 2 dd. — Stn DW2759, 15°55'N, 121°55'E,

122-139 m, 13 dd, 1 dd juv.

Distribution. Philippines (Mactan, Balicasag and

Bohol Islands), 56-250 m (dd - using also data of

Poppe et al., 2006).

Remarks. The main characteristics of this species are:

+ height up to 4.4 mm, width up to 3.4 mm;

+ slightly cyrtoconoidal; rather elevated

sculpture cancellate;

+ teleoconch of about 4 convex whorls with 3

granular spiral cords on penultimate whorl; rounded

periphery; on 1* whorl, only thin prosocline threads:

on 2% whorl, P1, P2 and P3 appearing at begin of

whorl, granular, with round beads distinctly separate;

distance between cords about two times size of cords;

P4 peripheral on last whorl;

+ aperture With labral fold-shaped denticles that are

the ends of in-running spiral ridges;

spire;

Figure 9 (scale bar: 1 mm).

+ columella with one short basal tooth;

+ convex base with 4 granular spiral cords; distance

between cords about 1.5 times size of cords;

+ no umbilicus;

+ cream white.

This species was described by Poppe et al. as a

Vaceuchelus species, but the typical U-notch under the

columellar tooth shows that Herpetopoma is a better

genus for it. This species seems very close to

Herpetopoma verruca (Gould, 1861). Comparison of

the types of H. verruca and H. ludiviniae should

establish if these two close species are the same one or

not. An accurate comparison With Æ. elevata Jansen,

1994 from Queensland should be also needed.

Herpetopoma poichilum Vilvens, 2012

Figs 8P-T, Table 7

Herpetopoma poichilum Vilvens, 2012: 6,8, figs GI-

G2, 16-19. Type locality: Society Islands, Tahiti,

455-650 m.

Material examined. French Polynesia, Australes

Archipelago. BENTHAUS: stn DW1868, 28°59'%,

140°14'W, 173-250 m, 1 dd. — Stn DW1876, 28°59'S,

140°15'W, 150-160 m, 2 dd. — Stn DW1884, 27°54S,

143°33'W, 570-620 m, 1 dd. — Stn DW1998, 22°25SS,

JS1-22%, 250-02m.41rdad:

Distribution. French Polynesia, Société Islands,

460-700 m, Iv at 455-650 m (fide original

description); Australes Archipelago, 160-570 m (dd).

Remarks. The main characteristics of this species are:

+ height up to 4.8 mm, width up to 4.3 mm;

+ elevated, spire rather high, conical;

+ _protoconch of about 200 um;

+ teleoconch up to 5.1 convex whorls with 4

granular spiral cords on penultimate whorl; subangular

periphery; on first whorl, thin prosocline threads,

distance between threads interspace between similar in

size to threads; at end of whorl, 4 spiral cords Pi (i=1

to 4) appearing almost simultaneously, P1 smaller; on

last whorl, all cords similar in size, S4 peripheral,

slightly smaller than other cords; distance between

cords smaller than cords;

A-U. Herpetopoma verruca (Gould, 1861). A-C. Indonesia, Tanimbar Islands, KARUBAR, stn DW49, 210-206

m, 4.1 x 3.1 mm. D-F. Solomon Islands, SALOMON 2, stn DW2234, 182-277 m, 3.2 x 2.4 mm. G-I. Wallis

Island, MUSORSTOM 7, stn DW601, 350 m, 3.0 x 2.8 mm. J-L. New Caledonia, Koumac, MONTROUZIER.

stn 1323, 82-120 m, 3.0 x 2.2. M-O. New Caledonia, LAGON, stn 830, 105-110 m, 3.0 x 2.3 mm. P-R. Fiji,

BORDAU 1, stn DW1469, 314- 377 m, 2.4 x 2.2 mm. S-U. Tonga Islands, BORDAU 2, stn DW1523, 300-302

ie 222002 SUN

V-X. Herpetopoma ludiviniae (Poppe, Tagaro & Dekker, 2006), PANGLAO 2004, stn P1, 90-200 m, 2.8 x 2.2

mm.

C. VILVENS

Chilodontidae from the Pacific Ocean

+ strong labral folds all around the outer lip of the

aperture;

+ columella with one small basal tooth;

+ base convex with 5 subgranular to granular,

similar in size Spiral cords; distance between cords

smaller than cords;

+ closed or very narrow umbilicus (reduced to a

small chinK);

+ white to yellowish white.

Herpetopoma vitilevuense n. sp.

Figs I0A-F, Tables 7, 9

Type material. Holotype (2.7 x 1.9 mm) MNHN (IM-

2000-32788). Paratypes: 6 MNHAN (IM-2000-32789),

1 paratype CV.

Type locality. Fiji, southeast of Viti Levu,

MUSORSTOM 10, stn CP1376, 18°19'S, 178°09'E,

497-504 m.

Material examined. Fiji: MUSORSTOM 10:

SCP1566, 1812S, 1782335, 149-168 m, dd. >

Stn CP1376, 18°19/S, 178°09E, 497-504 m, S0“Iv

(with holotype and paratypes). — Stn DW1383,

18°18.4S, 178°02.6E, 230-251 m, 60 dd. — Stn

DW1390, 18°18.6'S, 178°05.1'E, 234-361 m, 16 1v. —

Stn DW1384, 18°19'S, 178°06'E, 260-305 m, 5 dd. —

SUVA 2: sin BS18, 18°11'S, 178°28'E, 83 m, 1 dd. —

BORDAU 1: stn DW1469, 19°40'S, 178°10'W, 314-

Sn dd

Solomon Islands. SALOMON 1: stn DW1820,

9525, 160°51E, 256-329 m, 1 dd. — Sin DW1825,

9°50'S, 160°58'E, 340-391 m, 1 dd.

Distribution. Fiji, 83-497 m, lv at 361-497 m;

Solomon Islands, 329-340 m (dd).

Diagnosis. À small yellowish white Herpetopoma

species with a moderately elevated, cyrtoconoidal

spire and a rounded periphery, 3 granular spiral cords

on the penultimate whorls with separate beads, a week

Keel made by the third abapical cord, a small, obtuse

columellar basal tooth, strong fold-shape denticles

inside the aperture and a convex base with 4 spiral

cords and without umbilicus.

Description. Shell of small size for the genus (height

up to 3.1 mm, width up to 2.4 mm), higher than wide,

rather thick, slightly cyrtoconoidal, with a rounded

periphery; spire moderately elevated, height 1.1x to

1.6x width, 2.3x to 3.4x aperture height; anomphalous.

Protoconch about 200-250 um, of 1 whorl, with a thin

terminal varix.

Teleoconch up to 4.2 weakly convex whorls, bearing 3

spiral granular cords similar in size, with an additional

smaller peripheral cord on last whorl; beads from

cords at intersections with strong axial prosocline ribs,

rather strong, clearly separate. Suture visible, not

canaliculated.

First whorl convex, sculptured by about 25 prosocline

smooth, rather thin threads; interspace between

threads about 1.5x size of threads. On second whorl,

primary cords P1, P2 and P3 appearing at begin of

whorl, granular, similar in size; distance between

cords similar to size of cords; axial threads connecting

beads of cords. On third whorl, cords and threads

much stronger (threads becoming ribs), producing a

reticulate pattern; distance between cords about 1x to

1.5x size of cords; no Si. On last whorl, beads of cords

and ribs strong; distance between cords about 1.5x to

2x size of cords; P3 peripheral, making a week keel;

P4 emerging from suture, thinner than the other cords.

Columella straight, vertical, with a small, blunt basal

tooth.

Aperture almost circular; outer lip thickened internally

and weakly flaring, especially at columella side; 9 to

12 inner long fold-shaped labral denticles that are not

the ends of in-running spiral ridges; innermost

denticle separated from basal columella tooth by a U-

shaped notch.

Base convex, with 4 subgranular, similar in size spiral

cords; distance between cords 1.5x to 2x size of cords.

No umbilicus .

Colour of teleoconch and protoconch white to

yellowish white.

Operculum corneous, multispiral with central nucleus,

light brown.

Figure 10 (scale bar: 1 mm).

A-F. Herpetopoma vitilevuense n. sp., Fiji, southeast of Viti Levu, MUSORSTOM 10, stn CP1376, 497 504 m.

A-C. Holotype MNHN (1M-2000-32788), 3.1 x2.1 mm. D-F. Paratype MNEN (IM-2000-32789), 2.7 x 1.9 mm.

G-L. Herpetopoma naokoae Poppe, Tagaro & Dekker, 2006, Philippines. PANGLAO 2004. G-I. Stn T13. 90-

100 m, 3.4 x 2.5 mm. J-L. Stn T31, 100-140 m, 3.3 x 2.3 mm.

M-R. Herpetopoma hivaoaense n. sp., Marquesas Island, MUSORSTOM 9, stn 1204, 60-62 m. M-O. Holotype

MNAN (1M-2000-32790), 4.0 x 2.9 mm. P-R. Paratype MNHN (1M-2000-32791), 3.6 x 3.0 mm.

S-X. Herpetopoma corrugatum (Pease, 1861), French Polynesia, Marquesas Islands, Ua Huka, MARQUISES.

S-U. Stn 12, intertidal zone, 4.2 x 3.4 mm. V-X. Stn 35, 25 m, 3.1 x 2.9 mm.

C. VILVENS Chilodontidae from the Pacific Ocean

TW H W HA H/W H/HA

holotype 0.9 1.42 3.00

paratype | 0.8 142 | 3.38

paratype 2 (ol 63) 2.82

paratype 3 0.9 1.29 3.00

Jparatype 4 |" 4 | [| 116) 270

26 1.8 0.8 1.44 3.25

Table 9. Herpetopoma vitilevuense n. sp.: Shells measurements in mm for types.

Discussion. The new species is rather close to

Herpetopoma ludiviniae (Poppe, Tagaro & Dekker,

2006) from the Philippines (figs 9V-X) but this

species has much thinner spiral cords with smaller and

more closely packed beads, lacks the weak keel made

by P3 and labral fold-shaped denticles that are the

ends of in-running spiral ridges.

H. vitilevuense n. Sp. may also be compared to H.

verruca (Gould, 1861) from north-western Info-

Pacific (figs 9A-U) but this slightly smaller species

has very prominent spiral cords with more closely

packed beads, a spiral cord P1 appearing later than P2

and P3, and spiral cords S1 and S2 present.

Etymology. Named after the type locality.

Herpetopoma naokoae

Poppe, Tagaro & Dekker, 2006

Figs 10G-L, Table 7

Herpetopoma naokoae Poppe, Tagaro & Dekker,

2006: 37-38, pl.10, fig.1,3. Type locality: Mactan

Island, Philippines.

Herpetopoma naokoae — Herbert, 2012: 443-444, fig.

35:A-EF, 36.

Material examined. Philippines. PANGLAO 2004:

stul2, DOPS2/N 1252481520, LLdd, dd juy. —

SIMS COPAIN 25 S 0 000 dd = Sin

T31, 09°33'N, 123°42'E, 100-140 m, 3 dd. — Stn L44,

9O°31'N, 123°41'E, 85-100 m, 5 dd. — Stn L69-73,

0931'N, 123°41'E, 90-98 m, 14 dd. —- AURORA

2007: stn DW2750, 15°53'N, 121°54'E, 538 m, 6 dd. —

Guphil, 150m, 1 dd (MNAN IM-2000-5298).

Solomon Islands. SALOMON ll: stn DW1762,

8°40'S, 160°04'E, 396-411 m, 4 dd, 1 dd.

Wallis Island. MUSORSTOM 7:

13°21'S, 176°08'W, 415-420 m, 19 dd.

stn DW604,

Distribution. Philippines (Mactan, Balicasag and

Bohol Islands), 100-538 m (dd); Solomon Islands,

396-411 m (dd); Wallis Island, 415-420 m (dd).

Remarks. The main characteristics of this species are:

+ height up to 3.5 mm, width up to 3 mm:

+ rather elevated spire, slightly cyrtoconoidal;

sculpture cancellate;

+ protoconch of about 100-150 um;

+ teleoconch of about 4.5 convex whorls with 4

granular spiral cords and subsutural horizontal ramp;

rounded periphery; on first whorl, only thin prosocline

threads, distance between threads at least 2x larger

than threads; on second whorl, P2, P3 and P4

appearing at begin of whorl, granular, P4 the

strongest; on third whorl, PI resolving slowly; S4

peripheral on last whorl;

+ columella with one basal, sharp, small tooth;

+ moderately convex base with 5 rather thick,

granular, spiral cords; distance between cords smaller

then cords;

+ labral folds all around the outer lip of the

aperture;

+ no umbilicus;

+ yellowish white.

Herpetopoma hivaoaense n. sp.

Figs 10M-R, Tables 7,10

Type material. Holotype (4.0 x 2.9 mm) MNHN (IM-

2000-32790). Paratypes: 5 MNHN (IM-2000-32791),

1 paratype CV.

Type locality. French Polynesia, Marquesas Island,

Hiva Oa Island, MUSORSTOM 9, stn 1204, 9°53'S,

139°03'W, 60-62 m.

Material examined. French Polynesia, Marquesas

Island. MUSORSTOM 9: stn 1204, 9°535,

139°03"W, 60-62 m, 50 1v (with holotype and

paratypes).

Distribution. Only known from the type locality.

Diagnosis. À small light brownish. Herpetopoma

species with an elevated, conical spire and a

subangular periphery, 3 granular spiral cords on the

penultimate whorl with separate beads and a stronger

C. VILVENS

NOVAPEX 18CHS 11): 1-67, 10 mars 2017

ZA rent

abapical cord with sharp beads, 6 or 7 cords on the last

whorl, a small, obtuse columellar basal tooth, strong

fold-shape denticles inside the aperture and a weakly

convex base with 3 strong, prominent spiral cords and

a narrow umbilicus.

Description. Shell of small size for the genus (height

up to 4.0 mm, width up to 3.0 mm), higher than wide,

rather thick, more or less conical, with a subangular

periphery; spire elevated, height 1.2x to 1.5x width,

2.7xX 10 3.6X aperture height; narrow and deep

umbilicus.

Protoconch about 100-150 um, of 1 whorl, flat and

somewhat sunken, with a terminal lip hard to

distinguish.

Teleoconch up to 4.0 straight to very weakly convex

whorls, with 3 granular spiral cords, the abapical one

the strongest and 6 granular spiral cords of different

size on last whorl; beads from cords at intersections

with strong axial prosocline ribs, rather strong,

somewhat pointed. Suture hardly visible, not

canaliculated.

First whorl convex, sculptured by 20-25 prosocline

smooth, poorly marked, rather thin threads; interspace

between threads similar to size of threads; P3

appearing after mid whorl, granular, quickly rather

strong; P2 appearing near the end of whorl, weaker

than P3. On second whorl, PI appearing at first half of

whorl, very weak and weakly granular; beads of P3

slightly pointed; distance between cords similar to size

of cords; axial threads connecting beads of cords. On

third whorl, cords and threads much stronger, threads

becoming ribs, producing a subreticulate pattern;

beads clearly separate, beads of P3 sharp; distance

between cords similar in size to size of cords. On

fourth whorl, Pi more or less similar in size, with P3

slightiy stronger; SI and S2 appearing after half of

whorl. On last whorl, P4 emerging from suture,

slightly stronger than SI and S2: S3 possibly

appearing at extreme end of whorl.

Columella straight, vertical, with a small, blunt basal

tooth.

Aperture roundly elliptical, very slightly transverse;

outer lip thickened internally and flaring; about 9 long

fold-shaped labral denticles as the ends of in-running

spiral ridges; two basal denticles separated from basal

columella tooth by a U-shaped notch.

Base weakly convex, with 3 strongly granular,

elevated, similar in size spiral cords: axial threads

between cords, connecting beads; distance between

cords similar in size to cords.

Deep and rather narrow umbilicus, funnel shaped,

with angular rim crenulated by axial threads.

Colour of teleoconch and protoconch light brown or

beige.

Operculum corneous, multispiral with central nucleus,

light brown, deeply retracted into the aperture.

Table 10. Herpetopoma hivaoaense n. sp.: Shells measurements in mm for types.

Discussion. The new species is rather close to

Herpetopoma poichilum Vilvens, 2012 from the

Society Islands (French Polynesia) (figs 8P-T), but

this slightly greater species has a different ontogeny of

cords with all the Pi appearing almost simultaneously,

lacks the SI, S2 and S3 cords, has 5 spiral cords on

the base and a closed (or reduced to a small chink)

umbilicus.

H. hivaoaense n. sp. may also be compared to H.

pantantoi n. sp., from New Caledonia (figs 8J-0), but

this slightly greater, strictly conical in shape species

has again a different ontogeny of cords with all the Pi

appearing almost simultaneously, has 5 spiral cords on

the base and no umbilicus.

Etymology. Named after the type locality.

Herpetopoma corrugatum (Pease, 1861)

Figs 10S-X, Table 7

Euchelus corrugatus Pease, 1861: 435. Type locality:

Sandwich Island (nowadays Hawaïi).

Euchelus fimbriatus — Kay, 1965: figs 12-13.

Euchelus corrugatus — Kay, 1979: 49, fig 14-A.

Herpetopoma corrugatum — Severns, 2011: 44, pl. 6,

fig 3.

Herpetopoma corrugatum — Vilvens, 2012: 5-6, figs

270%,

Herpetopoma corrugatum — Herbert, 2012: 439, figs

69A, B.

Material examined. French Polynesia, Marquesas

Islands. MUSORSTOM 9: stn DW1203, 9°535,

C. VILVENS

139°02'W, 60-61 m, 1 Iv. —- MARQUISES: stn 12,

8505, 139°33/W, intertidal zone. Sr dd "Sri,

8°56'S, 140°06'W, 15-30 m, 1 dd. — Stn 23, 8°56'S,

13993 l'W, intertidal zone, 1 dd. — Stn 24bis, 8°54'S,

139°37'W, 20-34 m, 1 dd. — Stn 30, 8°56'S, 139°32'W,

20-30 m, 2 dd. — Stn 35, 8°56'S, 139°31'W, 25 m, 11

dd. — Stn 36, 8°56'S, 139°32'W, intertidal zone, 1 dd.

— POLYNESIE FRANCAISE: stn 02, 8°56'S,

140°06'W, 10-20 m, 4 dd, 5 dd juv. — Stn 03, 8°56'S,

140°07'W, 10-20 m, 3 dd. — Stn 20, 9°21'S, 140°06'W,

10-15 m, ldd.

Distribution. Hawaïi, Marshall Islands, subtidal (Kay,

1979; Severn, 2011); French Polynesia, Marquesas

Islands, 1-60m, 1v at 60-61 m.

Remarks. The main characteristics of this species are:

+ height about 3.8 mm, width about 3.7 mm

(lectotype);

+ moderately elevated spire, conical to slightly

cyrtoconoidal; sculpture cancellate;

+ protoconch smooth;

+ teleoconch of about 4 slightly convex whorls with

5 granular spiral cords on penultimate whorl;

subangular periphery; P1, P2 and P3 clearly visible at

2" whorl, P3 the strongest; axial threads between

cords, connecting rounded beads of cords; distance

between cords similar to cords; on 3" whorl, SI

resolving with S2 possibly later; P4 peripheral on last

whorl, possible S3, thinner;

+ columella with a weak basal tooth;

+ moderately convex base with 3 rather thick,

granular, spiral cords; distance between cords larger

than cords; axial threads between cords;

+ labral folds all around the outer lip of the

aperture;

+ narrow deep umbilicus;

+ whitish with dark spots.

Differences with Æ. fimbriatum (Pease, 1861) (type

locality: Sandwich Island, nowadays Hawaï) seem

not obvious. Kay (1965) mentions the lack of

umbilicus and, apparently ("four cords on last whorl"

2) the presence of the-simgle Si. -Severns (2011)

Chilodontidae from the Pacific Ocean

figures the NHMUK holotype, unfortunately without a

basal view; only SI is visible as secondary cord.

Herbert (2012) also figures the holotype and mentions

a more angular periphery and three spiral cords on the

penultimate whorl (instead of how many for /.

corrugatum ?). Moreover, Kay (1979) points out that

there are no recent records of this species in Hawaïi

area. Further studies are clearly needed to establish the

specific features of this poorly known species. AIT the

samples from French Polynesia here examined have

an umbilicus and are considered as HA. corrugatum,

although some of them have an earlier SI and a more

elevated spire (figs 10V-X).

Herpetopoma rubrum (A.Adams, 1853)

Figs 11A-F, Table 7

Monodonta rubra A. Adams, 1853: 176. Type

locality: not specified.

Euchelus hachijoensis Pilsbry, 1904: 406, pl. 27, figs.

39-40. Syn.

Euchelus ruber brunneus Pilsbry, 1901: 53. Syn.

Herpetopoma rubra - Wilson, 1993: 68.

Euchelus rubrus — Jansen, 1996: 11, fig. 27.

Euchelus rubra — Kosuge & Chino, 1998: 77, pl.25,

jipenle

Euchelus rubra — Higo, Callomon & Goto, 1999: 52

(giving China for type locality).

Euchelus rubra — Sasaki, 2000: 57, pl. 28, fig. 15.

Herpetopoma rubrum — Poppe, Tagaro & Dekker,

200639 pl Mies"125;

Material examined. Philippines. PANGLAO 2004:

stn L51-60 (collected in 10/2003), 9°38'N, 123°48'E,

62 m, 1 dd, 1 dd juv. — Stn L69-73 (collected in

10/2003), 9°31'N, 123°41'E, 90-98 m, 1 dd, 1 dd juv.

— Stn B15,.957N 125753E,5 5m, L'dd' Ste,

J°30N; 123%5S5E, 60 1m, l'dd'juv. - Sin LAG OS IIN,

123°41'E, 90-110 m, 1 dd.

Distribution. Philippines, 5-92 m; Australia (from

Western Australia to Queensland), subtidal; Japan,

Korea, China and Indonesia, subtidal.

Figure 11 (scale bar: 5 mm).

A-F. Herpetopoma rubrum (A.Adams, 1853). A-C. Philippines, PANGLAO 2004, stn L46 , 90-110 m, 8.2 x

8.0 mm. D-F. Western Australia, Broome, intertidal zone, 8.2 x 8.0 mm, coll. C.Vilvens.

G-I. Euchelus polysarkon n. sp. Taiwan, off Tashi, TAIWAN 2001, stn CP109, 24°48'S, 122°84'E, 246-256 m.

holotype MNEN (1M-2000-32792), 7.2 x 6.5 mm.

J-L. Euchelus asper (Gmelin, 1791), Singapore, Changi Point, intertidal zone, 11.4 x 11.2 mm. coll. C.Vilvens.

C. VILVENS

Chilodontidae from the Pacific Ocean

Remarks. The main characteristics of this species are:

+ height up to 8.5 mm, width up to 8 mm;

+ cyrtoconoidal in shape, rather elevated spire;

+ teleoconch of about 4.5 convex whorls with 8

granular, beaded spiral cords on the penultimate

whorl, up to 10 on the last whorl; beads of cords

rounded on first whorls, axially elongated on the last

whorls; first whorl with numerous prosocline, thin

axial threads; P2 and P3 after mid first whorl, PI at

begin of 2" whorl; SI at end of 2" whorl; S2 then S3

at begin of 3" whorl, P2 and P3 temporarily stronger;

T2 between S1 and P2, next T1 between P1 and Sl;

P4 and S4 visible on last whorl; distance between

cords similar in size to the width of cords;

+ aperture subcircular, with about a dozen ridge-

like denticles inside;

+ columella with one strong, prominent basal

columellar tooth and, on the Philippine samples only,

a second weaker upper tooth; deep U-shaped notch

between the basal tooth and the first labral denticle.

+ base convex, with 4, sometimes 5, strong granular

spiral cords; distance between cords similar in size to

the width of cords,;

+ rather wide umbilicus, with perspective to apex

and possibly (on the Philippine samples only) an

inductural callus only bordering the umbilical area;

+ light to dark red.

This species should be highly variable regarding the

columellar teeth system. Neither the second adapical

columellar tooth nor the partial covering of the

umbilicus by the inductural callus are indeed

mentioned in the original description. Australian

authors (e.g. Wilson, 1993; Jansen, 1996) and

Japanese authors (e.g. Kosuge & Chino, 1998; Sasaki,

2000) always figure more stocky samples with a

single columellar basal tooth. But nonetheless, the

cords ontogeny of the Pi-Si-Ti cords are the same on

the few Australian and Philippine samples I have

examined.

Comparison of the Philippine A. rubrum samples with

H. gemmatum (Gould, 1845) specimens is also clearly

needed: the general shape and the colour are very

different (whitish with reddish spots) but the cords

ontogeny is only slightly different, with S1 occurring

much later (and with S2) on H. gemmatum samples.

Genus Euchelus Phihppi, 1847.

Type species: Trochus quadricarinatus [Chem.]

Holten, 1802 [=7rochus asper

Gmelin, 1791], by subsequent designation

(Herrmannsen, 1847) — Recent, Indo-Pacific.

Remarks. The main features for Euchelus are a rather

big size (H up to 23 mm), numerous finely beaded

spiral cords, a poorly cancellate and never foveolate

sculpture. The main feature is a single weak columella

tooth, lacking the peg-like tooth coupled with an U-

shaped notch on the basal lip, distinctive for

Herpetopoma.

Euchelus polysarkon n. sp.

Figs 11G-I

Type material. Holotype (7.2 x 6.5 mm) MNHN (IM-

2000-32792).

Type locality. Taiwan, off Tashi, TAIWAN 2001, stn

CP109, 24°48'S, 122°?84'E, 246-256 m.

Material examined. Taiwan. TAIWAN 2001: stn

CP109, 24°48'S, 122°84'E, 246-256 m, 1 dd

(holotype).

Distribution. Only known from the type locality.

Diagnosis. À small whitish Euchelus species with a

moderately elevated, roundly conical spire and a

rounded periphery, 4 granular spiral cords on the

penultimate whorl with separate beads and, 6 cords on

the last whorl, a straight vertical columella without

tooth, a half-moon shaped aperture and a weakly

convex base with 6 spiral cords and without

umbilicus.

Description. Shell of small size for the genus (7.2

mm, 6.5 mm), higher than wide, roundly conical, with

a rounded periphery; spire moderately elevated, height

1.1x width, 2.2x aperture height; anomphalous.

Protoconch about 200 um, of 1 whorl, flattened,

without visible terminal lip.

Teleoconch of 4.8 convex whorls, with 4 granular,

similar in size spiral cords on penultimate whorl and 6

cords on last whorl; distance between cords greater

than cords;: beads from cords rounded, isolated, at

intersections With strong axial prosocline ribs. Suture

visible, not canaliculated.

First whorl convex, sculptured by about 20 smooth,

rather large, poorly marked, prosocline threads;

interspace between threads similar to size of threads.

On second whorl, P2, P3 and P4 appearing, P1 half a

whorl later, all with rounded, isolated, granular beads:

PI weaker than the other cords; distance between

cords slightly greater than size of cords; axial threads

connecting beads of cords. On third whorl, cords and

threads much stronger, threads becoming ribs; SI

resolving neat end of whorl. On fourth whorl, PI as

strong as the other Pi; beads of cords slightly axially

elongated. On last whorl, S4 emerging from suture,

weaker than all the other cords.

Columella straight, vertical, with two weak bulges.

Aperture half-moon shaped: outer lip rather thick,

without but with short notches corresponding to the

external spiral cords.

Base moderately convex, with 6 granular, similar in

size Spiral cords; axial threads between cords,

connecting beads; distance between cords similar in

size to cords.

No umbilicus.

Colour of teleoconch and protoconch nacreous white.

except the reddish brown first teleoconch whorl.

C. VILVENS

NOVAPEX ISCHS 11): 1-67, 10 mars 2017

Discussion. Among the known Chinese

Chilodontidae (Dong, 2002; Qi. 2004) or even

Southeast Asia (e.g. Dharma, 1988; Swennen et al.

2001; Thach, 2007), no Euchelus or Herpetopoma

species seem close to the new species, except

thesimilar in shape Euchelus scaber (Linnaeus, 1758)

that is much bigger with alternating strong and weaker

spiral cords and has an umbilicus.

On the same way, the new species can be compared to

E. asper (Gmelin, 1791) (syn. Æ quadricarinatus

(Holten, 1802)) from western Indo-Pacific (figs 11J-

L) that is much bigger with much stronger, more

widely spaced spiral cords and has also an umbilicus.

Etymology. Stout, plump (Greek: noAVONPYOË, Ov)

— With reference to the very large and very convex last

whorl of the shell.

Euchelus atratus (Gmelin, 1791)

Figs 12A-0

Turbo atratus Gmelin, 1791: 3601, No 53. Type

locality: Nicobar Islands.

Trochus canaliculatus Lamarck, 1818. Syn.

Euchelus atratus — Dharma, 1988: 32, pl1.2, fig.18.

Euchelus atratus — Kaicher, 1990: card # 5062.

Euchelus atratus — Jansen, 1996: 11, fig.26.

Herpetopoma atrata — Wilson, 1993: 68, pl.10, fig.3;

Euchelus rubra — Higo, Callomon & Goto, 1999: 52.

Herpetopoma atratum — Poppe, Tagaro & Dekker,

2006: 34, p1.9, fig.2,4.

Euchelus atratus — Herbert, 2012: 426-427.

Material examined. Philippines. PANGLAO 2004:

stn M3, 09°33'N, 123°45'E, 0-2.5 m, 1 Iv. — Stn M7,

0936'N, 123°45'E, 0-3 m, 1 Iv sub. — Stn B18,

09°39'N, 123°50'E, 3-5 m, 5 1v, 12 1v sub, 15 dd juv. —

Stn S22, 09°29'N, 123°56'E, 15-20 m, 4 dd juv. — Stn

B31, 09°39'N, 123°50'E, 1-2 m, 1 dd sub, 15 1v juv. —

Stn S38, 09°38'N, 123SlE, 3-4 m, 3 dd. -

PANGLAO 2005: stn CP2380, 8°4l'N, 123°18"E,

163-271 m, 1 dd, 1 dd sub.

Vanuatu. SANTO 2006: stn LMI15, 15°375,

167°11l'E, intertidal zone, 3 Iv sub, 1 Iv sub. — Stn

VM52, 15°26'S, 167°13'E, intertidal zone, 6 1v, 2 Iv

sub, 1 Iv juv. — Stn VMS3, 15°3/'S, 167°12'E,

intertidal, 9 1v, 2 Iv sub.

Indonesia, Moluccas Islands. KARUBAR: Ambon,

stn Seith, intertidal zone, 1 dd.

Distribution. Philippines, 1-163 m (using also data of

Poppe et al., 2006); Vanuatu, Iv in intertidal area:

Australia (from Western Australia to Queensland),

intertidal and shallow subtidal; Japan, Korea, China

and Indonesia.

Remarks. The main features of this species are:

+ upto 23 x 21 mm

+ spire rather elevated,

cyrtoconoidal;

conical to slightly

+ teleoconch up to 5.5 moderately to very Convex

teleoconch whorls; rounded periphery; P3 and P2 at

half of first whorl, PI a bit later; S2 and S3 at the

beginning of second whorl, SI from half to end of

second whorl; Ti at fourth whorl between Pi and Sr:

on last whorl, P4 peripheral; cords similar in size:

beads of cords high, axially elongated from third

whorl; distance between cords smaller than cords;

+ columella with a weak basal tooth; denticles all

along the outer lip;

+ base convex with 4-5 spiral cords, similar to

teleoconch whorls;

+ rather large umbilicus, funnel shaped,

perspective to apex and weak axial lines;

+ reddish brown.

with

On the large specimens (H>20), SI appears with Pi

and is stronger than P1, while T1 appears very early.

Poppe et al. (2006) consider that the large specimens

from north-western Australia belong to another

species: Herpetopoma sulciferum (A.Adams, 1853).

The WoRMS's position is apparently only based on

these authors. But surprisingly, no Australian marine

malacofauna book that I know mentions this species.

Moreover, the Australian Faunal Directory (Beechey

& Ponder, 2014) considers it under the global name

Euchelus atratus (Gmelin, 1791). A further study 1s

clearly needed and so, waiting for such a survey, I

adopt here a "conservative" position, keeping E.

atratus as the only used name around the Indo-Pacific

area.

Genus: Ascetostoma Herbert, 2012

Type species: Euchelus providentiae Melvill, 1909

(by original designation) — Recent, Seychelles Is.

Remarks. The basic features for Ascetostoma are a

moderate to rather big size (H up to 16 mm). a reddish

colour, rounded whorls with numerous, finely beaded

spiral cords. But the main features of Ascetostoma are

above all (Herbert, 2012):

+ a columella with a strong, squarish, basal tooth and

a smaller, more rounded, median tooth, with

pronounced U-shaped notch between basal tooth and

first outer lip denticle:

+ a rather large umbilicus deep down a cavity.

apically closed by a callus, with a margin bearing

several small denticles and especially one larger in

parietal region:

+ a parietal glossy, translucent callus deposit

bordering the aperture (such a callus is much more

extensive in Clypeostoma) and bearing short oblique

ridges.

Ascetostoma ringens (Schepman, 1908)

Figs 13A-O

Euchelus ringens Schepman, 1908: 71-72, pl. VI, fig.

6. Type locality: Sulu Archipelago, Philippines, 350

C. VILVENS

Chilodontidae from the Pacific Ocean

Herpetopoma ringens — Poppe, Tagaro & Dekker,

2006: 38-39, pL.8, figs.1-2.

Ascetostoma ringens — Herbert, 2012: figs. 68:A-B.

Material examined. New Caledonia. LAGON: stn

DW267, 2222$, JGOMSE MON MAI St

DW320, 22°32$, 166°54E, 70 m, 2 dd: — Sin

DW602, 22°165S, 167°03'E, 43-48 m, 1 dd. — Stn

DW677, 21°37$S, 166°22'E;-32 mm, -L'ddsub. — Sin

DW830, 20°49S, 165°19'E, 105-110 m, 10 dd, 15 dd

juv. — Stn DW1139, 19°24S, 163°47'E, 39 m, 1 dd

sub. —- MUSORSTOM 4, stn DWI162, 18°355,

163°10'E, 525 m, 2 dd. —- BATHUS 1: stn DW692,

20835S, ACPSOE A0 TS 0 un Sn

MONTROUZIER: stn 1260, 20°44'S, 165°14'E, 49-

59 m,1 dd. — Stn 1269, 15—201m, 20°35'S, 165°08'E,

49-59 m, 1 dd. — Stn 1315, 20°41'S, 164°15'E,

66 Sad Sin 1828 20P4NS IGAUISIE,

82-120m 3 dd, 2 ddnsub. 2,dd'qjuv. Sin 1331;

20°40'S, 164°11'E, 55-57 m, 1 dd sub, 4 dd juv. —

BATEAUS 4: stn DW882, 22°02'S, 165°56'E, 250-350

ul dd NORFOER IE Sn D IMPT 220;

IGN EE 190212 1m Jtdid sub. = /EBISCO:. sin

DW2555, 21°04'S, 158°35'E, 500-614 m, 1 dd. — Baie

St Vincent, ilot Tenia, 42 m, coll Menou, 1 dd sub.

Loyalty Islands. MUSORSTOM 6: stn DW442,

20SÆS IGPITE, 200m, dd, 1ddsub. = MIEOU

2000: stn 1461, 20°54'S, 167°02'E, 100-120 m, 1 dd. —

Stn DW1648, 20°54'S 167°03'E, 150-200 m, 2 dd.

Ein MUSORSTIONPID Sin CPI563, 1812:

178°33'E, 144-150 m, 1 dd sub. —- SUVA 4: stn

DWO8, 18°22'S, 178°02'E, 28-30 m, 5 dd.

French Polynesia, Australes Archipelago. RAPA

2002: stn 5, 27°06'S, 144°19'W, 8 m, 1 dd, 1 dd sub. —

Stn 8, 27°37'S, 144°18'W, 52-57 m, 1 dd sub. — Stn 9,

27°37'S, 144°22'W, 3-24 m, 1 dd, 1 dd juv. — Stn 10,

27°35'S, 144°23'W, 16-18 m, 1 dd. — Stn 14, 27°36$,

144°14'W, 2 m, 2 Iv. — Stn 34, 27°39'S, 144°19'W, 2-8

m, 2 dd. — Stn 42, 27°37'S, 144°18'W, 2 m, 1 dd. — Stn

44, 27°36'S, 144°18'W, 30 m, 2 dd sub, 10 dd juv. —

Stn 46, 27°37'S, 144°20'W, 10-42 m, 1 dd. — Stn 47,

27°3T'S, 144°19'W, 33 m, 1 dd, 4 dd juv. — Stn 61,

27°37T'S, 144°19'W, 10-15 m, 1 dd, 10 dd juv. — Stn

NO DS TS IP ON IS 20 nn lu dd

BENTHAUS: stn DW1927, 24°39'S, 146°02'W, 95-

105 m, 1 dd sub. — Stn DW1939, 23°50'S, 147°42'W,

100 m, 2 dd sub. — Stn DW1968, 23°23'S, 150°44'W,

100-120 m, 2 dd, 2 dd juv. — Stn DW1985, 23°265%,

150°44'W, 100-107 m, 1 dd, 1 dd sub. — Stn DW1997-

2001 22 26/27S, 15020 200-1000 did Sn

DW2013, 22°39'S, 151°50'W, 80-93 m, 1 dd sub. —

Stn DW2018, 22°37'S, 151°49'W, 770-771 m, | dd.

Vanuatu. SANTO 2006: stn DS99, 157335,

167°17'E, 100-105 m, 1 dd, 1 dd sub. — Stn DS103,

15°31'S, 167°16'E, 70-80 m, 1 dd.

Solomon Islands. SALOMON 1: stn DWI1770,

8°20'S, 160°39'E, 453-542 m, 1 dd sub. — Stn

DW1823, 9°50'S, 160°53E, 82-83: m, 1 dd. —

SALOMON 2: stn DW2198, 7°43'S, 158°30'E, 273-

300 m, 1 dd. — Stn DW2237, 6°53'S, 156°22'E, 400 m,

4 dd, 9 dd sub, 2 dd juv.

Indonesia. KARUBAR: stn

I38801E, 8200 dd,

Philippines. PANGLAO 2004: stn B7, 09°36'N,

123°52'E, 4-30 m, 8 dd, 10 dd sub, 10 dd juv. — Stn

B40, 09935'N, 123°50'E, 22 m, 1 dd, 2 dd sub, 6 dd

juv. — Stn LA49 , 9937/N, 12345E,/ 90m, 1rdd,1rdd

juv. — Stn T4, 09933'N, 123°49'E, 82 m, 1 dd. — Stn

T38, 09°32'N, 123°42'E, 80-140 m, 2 dd, 4 dd juv.

DW22,4:1059225;

Distribution. New Caledonia, 32-525 m (dd); Loyalty

Islands, 120-200 m (dd); Fiji, 30-144 m (dd); French

Polynesia, Australes Archipelago, 2 -770 m, IV

at 2 m; Vanuatu, 80-100 m (dd); Solomon Islands, 83-

453 m (dd); Indonesia, 82 m (dd); Philippines, 22-90

m (dd).

Remarks. The main characteristics of this species are:

+ height up to 8.5 mm, width up to 7.5 mm;

+ rather elevated spire, cyrtoconoidal in shape:

+ _protoconch of 1 whorl, about 200 um;

+ teleoconch of up to 5 very convex whorls with up

to 5 or 6 granular, beaded spiral cords on penultimate

whorl and about 10 cords on last whorl; rounded

periphery; first whorl with rather strong axial threads,

P2 and P3 resolving at end of whorl, PI a bit later; S1

and S2 at beginning of third whorl, S3 at end of third

whorl or at begin of fourth whorl; P2 and P3 stronger

than other cords; on fourth whorl, T1, T2 and possibly

T3 appearing respectively between PI and S1, SI and

P2, P2 and S2; P4 peripheral on last whorl, cords more

or less similar in size; distance between cords similar

in size to the width of cords; possibly a very thin S4

under P4;

+ on last whorls, sulcus around the suture, made by

subsutural cord and suprasutural cord of preceding

whorl;

+ aperture more or less circular, with 12-14 ridge-

like denticles inside;

Figure 12 (scale bars: 5 mm).

A-O. Euchelus atratus (Gmelin, 1791). A-C. Vanuatu, SANTO 2006, stn VM53, intertidal zone. 14.3 x 13.2

mm. D-F. Indonesia, KARUBAR: stn Seith, intertidal zone, 9.9 x 9.8 mm, coll. C.Vilvens. G-I. Philippines,

PANGLAO 2004, stn S38, 3-4 m, 12.2 x 11.5 mm. J-L. Malaysia, intertidal zone, 18.1 x 15.8, coll. C.Vilvens.

M-O. Western Australia, Port Hedland, 5 m 22.4 x 21.3 mm, coll. C.Vilvens.

C. VILVENS

Chilodontidae from the Pacific Ocean

+ columella with a well developed, squarish, basal

tooth and a smaller, more rounded, upper one; U-

shaped notch between basal tooth and first outer lip

denticle.

+ base convex, with 4 strong granular spiral cords;

distance between cords from 1x to 1.5x the width of

cords; possibly a thin additional cord between the two

outermost cords; parietal region with spreading,

glossy, translucent callus deposit, bearing short

oblique ridges;

+ umbilicus closed by a callus, bordered by a strong

parietal tooth projecting over umbilical area and

additional smaller denticles;

+ brown or orange to brownish white, possibly with

darker brown spots.

Herbert (2012) considers that Ascetostoma ringens

could be conspecific with A. providentiae (Melvill,

1909) from western Indian Ocean, the only obvious

differences being that the African species is slightly

more elevated and have a wider umbilicus.

On the same way, the samples coming from Australes

Archipelago (Rapa Island) show minor differences

with the specimens from the other areas: the whorls

are less convex, the parietal tooth on the callus 1s

much weaker and thin additional spiral cords are

present between the main cords of the base. But this 1s

a bit lightweight to consider that these Polynesian

species justify a new (sub)species.

Ascetostoma pteroton n. Sp.

Figs 13P-U, Table 11

Type material. Holotype (16.5 x 15.4 mm) MNHN

(IM-2000-32793). Paratypes: 2 MNHN (IM-2000-

32794), paratype CV.

Type locality. Australes Archipelago, Rapa Island,

Mei Point, RAPA 2002, stn 31, 27°38'S, 144°18'"W, 6

Material examined. French Polynesia, Australes

Archipelago. RAPA 2002: stn 5, 27°06'S, 144°19'W,

Son dd And Sub, 2 "ddr. Siné 27070)

144°18'W, 52-57 m, 3 dd, 6 dd sub. — Stn 10, 27°35S,

144°23'W, 16-18 m, 5 1v sub, 3 dd juv. — Stn 11,

27°37TS, 144°18'W, 2 m, 1 dd. — Stn 16, 27°36S,

144°18'W, 5 m, 1 dd, 1 dd juv. — Stn 17, 27°35S,

144°23%W, 9. mm, L dd juv. — Sin 19, 27°38S,

MON ENS nn du "Sin 20 2755,

14423, 5 mi. L''dd_juv. Sin 22 2734$,

144°22'W, 18-22 m, 1 dd juv. — Sin 28, 27°385,

IAA 50h, ds = Sin 29/2752,

144°21'W, 2-4 m, 3 dd ju. — Sin 30, 27°38$,

144°18'W, 16-20 m, 1 dd juv. — Sin 31, 27°38$,

144°18'W, 6 m, 4 dd (with holotype and paratypes) . —

Stn 32, 27°35'S, 144°23'W, 15-20 m, 1 dd, 2 dd juv. —

Stn 33, 27°35'S, 144°19'W, 30 m, 1 dd juv. — Sin 34,

27°35'S, 144°19'W, 2-8 m, 1 dd sub. — Sin 34,

27°39'S, 144°19'W, 2-8 m, 2 dd sub, 6 dd juv. — Stn

43,27°37'S, 144°18'W, 45 m, 5 dd sub, 5 dd juv. — Stn

44, 27°36'S, 144°18'W, 30 m, 2 dd, 15 dd sub, 10 dd

juv. — Stn 48, 27°34'S, 144°22'W, 36 m, 9 dd juv. —

Stn 56, 27°37'S, 144°18'W, 25-30 m, 1 dd juv. — Stn

62, 27°37'S, 144°21'W, 20 m, 1 dd, 4 dd juv. — Stn 70,

27°37'S, 144°20'W, 15-20 m, 1 dd. — Stn 93, 27°355,

144°21'W, intertidal area, 1 dd juv. - BENTHAUS:

stn DWI1885, 27°52'S, 143°33'W, 700-800 m, 2 dd, I

dd sub.

Distribution. French Polynesia, Australes

Archipelago, 1-700 m, 1v at 16-18 m.

Diagnosis. À big pinkish with dark spots Ascetostoma

species with an elevated, rounded conical spire and a

rounded periphery, up to 18 thin granular spiral cords

on the whorls, a transversally elongated aperture, a

strong columellar basal tooth, strong fold-shape

denticles inside the aperture, a convex base with about

15 thin spiral cords, an umbilicus deep down within a

cavity in a glossy callus deposit.

Description. Shell of great size for the genus (height

up to 16.5 mm, width up to 15.4 mm), higher than

wide, rather thick, roundly conical; spire elevated,

height 1.0x to 1.1x width, 1.7x to 1.9x aperture height:

anomphalous.

Protoconch about 200 um, of 1 whorl, with a thin

terminal varix.

Teleoconch up to 6.6 weakly convex whorls, bearing

about 8 spiral granular cords on penultimate whorl,

about 18 thin cords of various size on the last whorl:

distance between cords slightly smaller than width of

cords; rounded periphery. Suture visible,

canaliculated.

First whorl convex, sculptured by about 20 strong,

smooth, prosocline threads; interspace between similar

in size to ribs; primary cords P2 and P3 appearing at

end of whorl. PI appearing at begin of second whorl,

thinner than the two other cords; all cords granular:;

distance between cords about 1x to 1.5x width of

cords; axial ribs stronger, connecting beads of cords.

On third whorl, all cords similar in size; beads of

cords rounded. SI, S2 and S3 appearing at begin of

fourth whorl; SI and SI quickly similar in size to Pi,

S3 thinner; beads becoming a bit pointed; sulcus

appearing at sutural level, made by subsutural cord

and suprasutural cord of preceding whorl. On fifth

whorl, 6 Ti appearing between the Pi and Si according

to a random order, each Ti between two succeeding

existing cords. On last whorl, additional cords

appearing between all the existing cords, giving a total

amount of spiral cords of about 18.

Aperture almost circular, obliquely elongated; outer

lip thickened internally and weakly flaring, with about

30 elongated, ridge-shaped denticles; possibly

additional smaller ridges between the main ridges on

the upper part of lip.

Columella more or less straight, slightly oblique, with

a basal tooth, rather strong on fully mature shells

C. VILVENS

NOVAPEX I8CHS 11): 1-67, 10 mars 2017

(more than 5 whorls); innermost denticle separated

from basal columella tooth by a U-shaped notch.

Base convex, with about 15 granular spiral cords,

similar to cords of the body whorl; on fully mature

shells, parietal region with spreading, glossy,

translucent callus deposit, bearing strong ridges.

paratype CV

Table 11. Ascetostoma pteroton n. sp

Discussion. The specific features of Ascetostoma

pteroton n. sp. make it difficult to confuse it with

other chilodontid species. At most could one compare

the new species to the well-known A. ringens

(Schepman, 1908) from western Indo-Pacific (figs

13A-O), but this smaller species has a very different

shape, stronger and less numerous spiral cords on the

whorls, a stronger columellar tooth and stronger, less

numerous inner apertural ridges.

Subadult samples (5 whorls or less) look rather

different of the fully mature shells: the umbilicus is

still open, there is no callus extending on the parietal

wall and the columellar tooth, when present, is

reduced. Their shapes evoke some Euchelus species,

but the cords ontogeny is strictly the one of the bigger

shells.

Etymology. Winged (Ancient Greek: ntEpatob, n,

OV) — with reference to the elongated, finely striated

aperture of the shell.

Genus Hybochelus Pilsbry, 1890

Type species: Stomatella cancellata Krauss, 1848 (by

original designation) - Recent, Indo-Pacific.

Remarks. The basic features for Hybochelus are a

moderate to rather big size (H up to 16 mm, W upto

20 mm), a thin shell, a depressed spire, an expanded

last adult whorl (but less than in Granata) and an open

umbilicus bordered by a strong spiral cord.

Hybochelus cancellatus (Krauss, 1848)

Figs 1I4A-L

Stomatella cancellata Krauss, 1848: 93, pl. 5, fig. 26.

Trochus (Euchelus) fossulatulus Souverbie in

Souverbie & Montrouzier, 1875: 39, pl.4, fig.5. Syn.

(Herbert, 1996).

Hybochelus fossulatulus — Poppe, Tagaro & Dekker,

2006: 40, pl.12, figs.7.

Hybochelus cancellatus — Wilson, 1993: 69, pl.10,

fig. 1;

Umbilicus deep down a cavity, apically closed by a

callus, bordered by a strong, flat parietal tooth and

additional smaller denticles.

Colour of teleoconch nacreous white with brownish

orange dashes on the spiral cords; protoconch pinkish

white.

.: Shells measurements in mm for types.

Material examined. New Caledonia.

MONTROUZIER: stn 1303, 20°38'S, 164°16'E, 0-8

m, 1 Iv, 1 dd juv. — Stn 1270, 20°45'S, 165°17'E, 10-

35 m, 1 dd. — Stn 1308, 24°40'S, 164°15'E, 15-20 m, 1

dd.

Loyalty Islands, Lifou. LIFOU 2000: stn 1421,

20°52'S, 167°09'E, 4 m, 1 dd. — Stn 1422, 20°475,

167°07'E, 4 m, 1 dd, 1 dd juv. — Stn 1423, 20°545S,

167°07'E, 12 m, 1 dd. — Stn 1424, 20°55'S, 167°03'E,

4 m, 1 dd. — Stn 1430, 20°48'S, 167°07'E, 20-25 m, 1

dd. — Stn 1432, 20°54'S, 167°03'E, 12-32 m, 3 Iv, 2 dd

sub, 1 dd juv. — Stn 1434, 20°53'S, 167°08'E, 5-20 m,

1 dd juv. — Stn 1448, 20°46'S, 167°02'E, 20 m, 1 dd

juv. — Stn 1449, 20°46'S, 167°02'E, 17 m, 1 dd, sub 1

dd juv.

Distribution. New Caledonia, 8-15 m (dd); Loyalty

Islands, Lifou, 4-20 m (dd), 1v at 12-32 m.

Remarks. The main characteristics of this species are:

+ height up to 16 mm, width up to 20 mm;

+ first whorl with axial threads: P3 almost

immediately, P2 almost simultaneously, PI later:

+ SI at or after beginning of second whorl, as

strong and other cords at end of whorl: axial threads as

thick as cords, giving a regular reticulate pattern:

+ at third whorl, P4 merging from suture: one

additional thinner Ti cord appearing between each

existing spiral cord, also between suture and PI and

under P4 on the last whorl, giving an amount of 11

spiral cords on the last whorl:

+ base with 5 main spiral cords, the most internal

one bordering the umbilicus: additional thin spiral

cords possibly between the most external main cords:

space between internal cords deeply excavated:

+ large, deep, funnel shaped umbilicus,

perspective to apex;

+ arcuate columella without tooth; no denticles in

the aperture.

with

I follow here Herbert (1996) who considers that

Hybochelus fossulatulus (Souverbie in Souverbie &

Montrouzier, 1875) 1s a synonym of Hybochelus

C. VILVENS Chilodontidae from the Pacific Ocean

cancellatus (Krauss, 1848). It can be only noted that, Remarks. The main features for Clypeostoma are à

on the New Caledonian species, S1 appears apparently slightly cyrtoconoidal to almost conical rather

earlier than on Australian A. cancellatus samples 1 elevated spire with flat to weakly convex, a reticulated

have examined. sculpture, a half-moon shaped aperture with an outer

lip flaring beyond an internal thickening and above all

Genus Clypeostoma Herbert, 2012 a) two prominent, rounded columellar teeth;

Type species: Turcica salpinx Barnard, 1964 (by b) a large inductural callus shield covering the

original designation) — Recent, South-eastern Africa. umbilical and parietal areas.

fron-

tal

view

colour

number

of cords

on base

which

Si/Ti

present

spiral lirae

in aperture

)

proto-

conch

exserted

number

Spiral

cords

penul-

timate

Clypeostoma

whorl (S)

nortoni s. Si, S4 We 5 E brownish

Cyrtoconoi- (denticles) white to

dal brownish

grey

brownish

white

(denticles)

chranos Cyrtoconoi-

n. SP. dal

cancella- conical, iVOrY /

tum spire mod. reddish

elevevated brown

flames

brown or

cream

with

brown

flames

yellowish

white to

light

brown

(denticles)

W 5 E

(denticles)

Table 12. Comparison of conchological features of small central Indo-Pacific Clypeostoma. species (sl. =

slightly, mod. = moderately).

cecileae conical

conical,

spire

elevated

adelon

Hi. Sp.

Figure 13 (scale bars: 5 mm).

A-O. Ascetostoma ringens (Schepman, 1908). A-F. New Caledonia. A-C. LAGON, stn 602, 43-48 m, 7.7 x 6.2

mm. D-F. MONTROUZIER, stn 1315, 66-87 m, 7.9 x 5.7 mm. G-L. French Polynesia, Australes Archipelago,

RAPA 2002. G-L stn 47, 33 m, 7.5 x 6.5 mm. J-L. stn 14, 2 m, 7.2 x 6.3 mm. M-O. Philippines, PANGLAO

2004, stn S8, 28-32 m 8.9 x 7.2 mm.

P-U. Ascetostoma pteroton n. Sp., French Polynesia, Australes Archipelago, Rapa Island, RAPA 2002, stn 31.6

m. P-R. Holotype MNHN (1M-2000-32793), 16.1 x 15.6 mm. S-U. Paratype MNHN (IM-2000-32794), 13.8 x

13.3 mm.

C. VILVENS NOVAPEX ISCHS 1!

C. VILVENS

Chilodontidae from the Pacific Ocean

Clypeostoma nortoni (McLean, 1984)

Figs 1SA-U, Table 12

Agathodonta nortoni McLean, 1984: 122, figs 1-3.

Type locality: off Baltazar Is., Philippines.

Agathodonta nortoni — Vilvens, 2001: figs 11-13;

Agathodonta nortoni — Vilvens & Héros, 2003: figs

10215:

Agathodonta nortoni — Poppe, Tagaro & Dekker,

2006: 30, pl. 7, figs 3, 4;

Clypeostoma nortoni — Herbert, 2012: 409, figs

8,12H-L.

Material examined. Philippines. PANGLAO 2004:

stn P1, 9°36'N, 123°45'E, 90-200 m, 25 1v, 4 lv sub. —

SULP3, OSTN, 123°410E,100m; 301, d'Ivsub. —Stn

MIO GIN IP SE 08m dd "Sim Er,

SN 2522408 20m id = PANGLAO 2005!

SHMCP2S81P OS OINMIPSSIE 255268 ni LI. = Sin

CR2552 0 SOIN PS AGE 6064118 1m, IN. "Sin

CPISAO OSOIN APS ARE 201-818 4m, 21. "Sin

CPISAS OP AN IPS AOIE 278 -S66um, 2 IN = Sin

CRISE OSON PSE SSCS96 um, 21 — Sin

GR2S97% OP OIN PS PMDIENAY70-SCO mm, 8 1. Sin

CP2S95 PS 6N, IPS 4PE, 382-45%4%m, 15 "1v. Sin

DW2401, 09°31'N, 123°40'E, 397-410 m, 2 dd. — Stn

CP2406, 9°AI'N, 123°47'E, 334-387 m, 8 Iv. —

AURORA 2007: stn DW2716, 14°30'N, 121°4]'E,

335-356 m, 1 dd, 1 dd sub, 3 dd juv. — Stn DW2670,

14°52'N, 121°49'E, 180-187 m, 20 dd, 10 dd sub, 15

dd juv.

Solomon Islands. SALOMON 1: stn DW1768,

8°21'S, 160°41'E, 194-286 m, 13 dd. - SALOMON 2:

SDW PIONEER 28IS ISO TE S00m, dd. -—

SALOMONBOA 3: stn DW2811, 09°42'S, 161°30'E,

228-238 m, 1 dd. -SALOMONBOA 3: stn DW2829,

10°46'S, 162°20'E, 220-292 m, 1 dd.

Taiwan. TAIWAN 2000: stn DW36, 21°55%,

l2086E, 305/ar 2 dd

Indonesia. Kai Islands. KARUBAR: stn DWI15, 212-

DDR OSMIS M2 NE rod Stn CIC OSAINS,

132°50E, 315-349 m, 1 1v. — Sin DW44, 07°52%S,

132 2481E 291-2951 dd:

Distribution. Philippines, 3-470 m, 1v at 100-470 m:;

Solomon Islands, 238-300 m (dd); Taiwan, 305 m

(dd); Indonesia, Kai Islands, 221-315 m (dd), lv at

315-349 m.

Remarks. The main characteristics of this species are:

+ height up to about 12.5 mm, width up to about 9.5

mm; up to 5.5 whorls;

+ first whorl with thin axial threads; on second

whorl P2, P3 and P4 almost simultaneously, PI later;

S1 at near end of third whorl, quickly as strong and

other cords at end of whorl; axial threads connecting

beads of cords, thinner than cords, giving a regular

reticulate pattern; on fourth whorl, beads pointed,

vertically elongated; S4 partially emerging from

suture; on last whorl, S4 fully visible, weaker than the

other cords; distance between cords from 1.5x to 2x

the size of cords;

+ two strong, rounded columellar teeth; columella

weakly excavated with 2 (possibly 3) denticles, the

adapical one thicker;

+ aperture with about 10 elongated denticles on the

outer lip and a basal tooth producing a notch with the

abapical columellar tooth:;

+ base with 5 strong, granular main spiral cords;

distance between cords similar to the size of cords;

+ callus shield covering the umbilical and parietal

areas, with weak superficial ridging on the parietal

zone;

+ brownish white to brownish grey.

In the examined material, one can notice that:

a) some Philippine samples have unexpected reduced

columellar teeth, despite their great size (H = 12.5

mm) and 5.5 whorls indicating maturity of the

specimens;

b) Taiwanese samples are more conical with almost

flat whorls and have additional tertiary cords, with

always at least T1 between P2 and P3;

c) on some Indonesian samples, S1 appears later on 4"

whorl;

d) one sample from Solomon Islands has small

granules on its callus shield.

Clypeostoma chranos n. sp.

Figs 15V-X, Table 12

Type material. Holotype (8.0 x 5.9mm) MNAN (IM-

2000-32795).

Type locality. Southeastern coast of Taiwan,

TAIWAN 2002, stn DW149, 22°19'N, 121°29'E, 258

Material examined. Taiwan. TAIWAN 2002: stn

DW149, 22°19'N, 121°29'E, 258 m, 1 dd (holotype).

Distribution. Only known from the type locality.

Diagnosis. À medium size chilodontid species with a

rather elevated, cyrtoconoidal shape and a subangular

periphery, with 4 granular cords on the penultimate

whorl, a subelliptic, slightly transversally elongated

aperture, a convex base with 3 spiral cords and a

granular inductural callus covering completely the

umbilicus.

Description. Shell of medium size for the genus

(height 8.0 mm, width 5.9 mm), higher than wide,

rather thick, cyrtoconoidal; spire elevated, height 1.4x

width, 2.8x aperture height; anomphalous.

Protoconch almost totally missing on the single

available specimen.

Teleoconch up to 5.1 convex whorls, bearing 4 strong

spiral granular cords; distance between cords similar

C. VILVENS

in size to cords; subangular periphery. Suture visible,

impressed.

First whorl convex. sculptured by about 20 rather

Strong, slightly prosocline threads:

interspace between them similar in size to threads: P1

and P3 appearing almost immediately, P2 and P4 a bit

later, all granular; beads of cords produced by

intersection between cords and threads. On second

whorl, axial threads stronger, interspace between them

about 2x width of threads; P3 and P4 the strongest, P1

the weakest; beads of cords no more close but

separated. On third whorl, all cords much stronger;

distance between cords ay most similar in size to

cords; beads of cords becoming pointed. On fourth

whorl, S4 partially emerging from suture; beads of

cords axially elongated: axial threads becoming ribs,

weaker than cords, connecting no more beads of

cords. On last whorl, S4 fully visible, slightly weaker

than the other cords.

Aperture subelliptical, axially elongated; outer thick,

with 8 strong, horizontally elongated denticles

corresponding to the external spiral cords of last whorl

and base; inner lip with 3 smaller basal denticles.

Columella straight, slightly opisthocline, weakly

excavated, without two teeth, the squared abapical one

much stronger;

Base weakly convex to almost flat, with 3 thick

granular spiral cords, similar to the cords of the last

whorl; very thin, close axial threads between cords.

Umbilicus completely filled by columellar callus:

umbilical and parietal areas covered by a large,

granular inductural callus shield, some (but not all)

granules corresponding to the beads of the underlying

basal spiral cords.

Colour of teleoconch brownish white to ochre; base

whitish.

Discussion. The new species is rather close to

Clypeostoma nortoni (McLean, 1984) from central

Indo-Pacific (figs 15A-U), but this similar in size

species has a more elongated shape, a SI spiral cord,

thinner spiral cords with greater interval between them

and usually 5 (rarely 4) spiral cords on the base.

C. chranos n. sp. is also rather similar to C.

reticulatum Herbert, 2012 from Mozambique, but this

proportionally greater size species has a more

elongated shape, more numerous and thinner spiral

cords on the whorls, similar columellar teeth and 4-6

spiral cords on the base.

Etymology. Helmet (Ancient Greek: kpavoc), used in

apposition — with reference to the shape of the shell,

reminding an Oriental helmet.

Clypeostoma cancellatum (Schepman, 1908)

Figs 16A-X, Table 12

Calliostoma cancellatum Schepman, 1908: 69-70, pl.

VI, fig. 5. Type locality: Samau strait, Timor,

Indonesia, 390 m.

NOVAPEX I18CHS 11): 1-67, 10 mars 2017

Perrinia cancellata - Poppe, Tagaro & Dekker, 2006:

40-41, pl.11, figs.4-5, pl.12, fig. 1,3.

Material examined. Philippines. PANGLAO 2004:

stn P1, 9°36'N, 123°45'E, 90-200 m, 1 dd juv. — Sin

P3, 993/l'N, 123°4/'E, 100 m, 2 dd. — Stn T1, 09°32'N,

123°47'E, 83-102 m, 20 dd, 5 dd juv. — Stn T4,

0933'N, 123°49'E, 82 m, 7 dd, 2 dd sub, 10 dd juv. —

Sin T36, 09°29'N, 123°52'E, 95-128 m, 4 dd sub, 6 dd

juv. — Sin T38, 09°32'N, 123°42'E, 80-140 m, 2 dd.

Taiwan. TAIWAN 2001: stn CP109, 24°48'N,

122°04'E, 246-256 m, 1 dd. - TAIWAN 2002: stn

DW117, 24°59'N, 122°03'E, 126-153 m, 1 dd. -Stin

DW149, 22°19'N, 121°29'E, 258 m, 1 dd.

Solomon Islands. SALOMON 1: stn CP1758, 8°49'S,

159°52'E, 180-187 m, 1 dd. — Stn CP1760, 8°475,

160°01'E, 172-179 m, 1 dd. — Sin DW1762, 8°40'5,

160°04'E, 396-411 m, 1 dd, 2 dd juv. — Stn CP1848,

10°27'S, 161°58'E, 159-169 m, 2 dd. - SALOMON 2:

stn CP2169, 9°0['S, 159°06'E, 100-200 m, 1 dd. — Sin

DW2254, 8°28S, 157°03'E, 150 m, 1 dd. -

SALOMONBOA 3: stn CP2804, 09°15'S, 161°21'E,

150-175 m, 1 dd.

New Caledonia. VAUBAN 1978-1979: stn 10,

22°17'S 167°05'E, 80 m, 1 dd. - LAGON: stn 830,

20°49'S, 165°19'E, 105-110 m, 8 dd. -MUSORSTOM

4: stn DW203, 22°36'S, 167°05'E, 105-110 m, 1 dd

sub. — CHALCAL 2: stn DW71, 24°42'S 168°IU'E,

230 m, 1 dd. —- BATHUS 1: stn CP688, 20°335,

165°00'E, 270-282 m, 10 dd, 1 dd sub. — Stn CP692,

20°35'S, 164°59'E, 140-150 m, 10 dd, 2 dd, 1 dd sub.

— BATHUS 2: stn DW724, 22°48'S, 167°26E, 344-

358 m, 5 dd, 4 dd sub, 5 dd juv. — Stn DW732,

22°50'S, 166°25'E, 236-264 m, 1 dd. —- BATHUS

2/MUSORSTOM 8 (mélange accidentel), 4 dd. —

MONTROUZIER: stn 1323, 20°4l'S, 164°15'E,

82-120 m, 19 1v. - BATHUS 3: stn CP827, 23°225,

168°0l'E, 381-469 m, 1 dd. -BATHUS 4: stn DW882,

22°02S, 165°56'E, 250-350 m, 1 dd. - NORFOLK 2:

stn DW2040, 23°41'S, 168°0l'E, 285 m, 1 dd. -

EBISCO: stn DW2549, 21°07S, 158°38'E, 330-331

m, 1 dd. — Stn DW2622, 20°04'S, 160°21'E, 291-323

m, | dd.

Loyalty Islands. LIFOU 2000: stn 1461, 20°548,

167°02'E, 100-120 m, 1 dd sub.

Fiji. MUSORSTOM 10: stn CP1366, 18°125,

178°33'E, 149-168 m, 1 dd sub 2 dd juv. - BORDAU

1: stn DW1435, 17°11'S. 178°45'W, 170-183 m., 1 dd.

— Stn DW1440, 17°11'S, 178°43'W, 190-308 m, 1 Iv.

Wallis and Futuna. MUSORSTOM 7: stn DWS512,

14°13'S 178°10'W, 210-245 m, 1 dd sub. — Stn

DW604, 13°21'S 176°08'W, 415-420 m. 1 dd sub, 1

dd juv.

Tonga. BORDAU 2: stn

174°57'W, 200-487 m, 1 dd.

DWI601, 20°50'5,

Distribution. Philippines, 82-152 m (dd - using also

Poppe et al., 2006 data); New Caledonia, 80-381 m

(dd), 1v at 82-120 m; Loyalty Islands, 100-120 m (dd):

C. VILVENS

Fiji, 168-190 m, Iv at 190-308 m; Wallis and Futuna,

245-415 m (dd); Tonga, 200-487 m (dd).

Remarks. The main features of this species are:

+ height up to 11 mm, width up to 8 mm;

+ spire rather elevated, conical;

+ teleoconch of up to 6.5 almost flat to weakly

convex Wwhorls; 5 granular spiral cords on the

penultimate whorl, 6 on the last whorl; first whorl

with thin prosocline threads, distance between threads

much greater threads; the four Pi at second whorl,

usually SI at third whorl; on last whorl, P4 the

strongest, peripheral, PI slightly weaker with strong

beads, other cords similar in size except S4 much

weaker; distance between cords similar to cords;

rather thick, possibly a bit lamellose, prosocline ribs

between cords;

+ columella with one strong, peg-like, basal tooth

and a weaker tooth above it;

+ aperture elliptical, with a small translucent

inductural callus shield; outer lip rather thick, with

about 15 inner folds; possibly one denticle at the base

of inner lip making a U-notch under the lowest

columellar tooth;

+ base rather convex, with 6 granular spiral cords;

distance between cords similar to size of cords; axial

thin threads between cords:

+ no umbilicus;

+ 1ivory with reddish brown flames and markings.

AI the studied samples match the Schepman's original

description very well, including the parietal callus and

the lamellose axial threads on the last whorls. The

only difference is that a second adapical columellar

tooth is not mentioned.

It seems indeed that there is à rather high variability

for this species. First, regarding the columella, the

original description and also apparently figures of

Poppe et al. (2006) mention only one pointed tooth,

for a type being 7 x 5 mm. But some medium and

bigger samples show an adapical swelling that can

give a second tooth. The small size of the type could

maybe explain the lack of mention of this second

tooth. à

Next, while the Philippine samples have always a SI

spiral cord, all the samples from Taiwan to Solomon

Islands and Tonga lack this S1. Some samples from

New Caledonia have this SI or not, sometimes on the

same station. Moreover, some specimens have an

accretion of axial sculpture elements along the

Chilodontidae from the Pacific Ocean

apertural border, reminding a very low varix weakly

similar to the one of Danilia species.

Finally, some variability is observed regarding the

spiral cords on the whorls, which can be thin to a

clearly thicker, with axial threads between them

lamellose or not.

Also to be noticed is the presence of the inductural

callus shield, which doesn't match with the Perrinia

characterization just like the secondary columellar

apical tooth. Herbert (2012) gives an exsert

protoconch as a distinctive feature of his new genus

Clypeosotoma but also noticed that Perrinia konos

(Barnard, 1964) has an exsert and rather globose

protoconch. Considering the columellar U-notch and

the parietal callus, is seems likely to put "Perrinia”

cancellata into the Clypeostoma genus, although the

protoconch of the studied samples is sunken, not

exsert.

AI the records here reported extend the distribution of

this species, splitting it apparently into two separated

areas.

Clypeostoma cecileae

(Poppe, Tagaro & Dekker, 2006)

Figs 17A-C, Table 12

Perrinia cecileae Poppe, Tagaro & Dekker, 2006: 42,

pl.13, figs.i-3. Type locality: Mactan Island,

Philippines.

Material examined. PANGLAO 2004: stn L51-60,

9938'N, 123°48'E, 62 m., 3 dd. - AURORA 2007: stn

CP27581 15 SSN,121FS0E4SI178 1m; d'dd.

Distribution. Philippines, 81-100 m (dd - using also

Poppe et al., 2006 data);

Remarks. The main characteristics of this species are:

+ height up to about 6.6 mm, width up to about 4.4

mm, about 6 whorls;

+ protoconch exsert, globose;

+ first whorl with thin axial threads; on second

whorl, P3, P4 almost simultaneously, Pland P2 later:

no Si; axial threads between cords, connecting beads

of cords; on next whorls, spiral cords stronger, P4

gradually the strongest with pointed beads on last

whorls, making stellate shape from a basal view:

distance between cords at least similar to the size of

cords;

Figure 14 (scale bars: 5 mm).

A-L. Hybocheius cancellatus (Krauss, 1848), New Caledonia. A-F. Loyalty Islands, Lifou. A-C. LIFOU 2000.

stn 1432, 12-32 m, 10.8 x 12.4 mm. D-F. LIFOU 2000, stn 1421, 4 m, 9.2 x 12.4 mm. G-I. Touho,

MONTROUZIER, stn 1270, 10-35 m, 7.2 x 8.6 mm. J-L. Western Australia, Broome, 6 m, 15.5 x 18.8 mm.

coll. C.Vilvens.

C. VILVENS

+ aperture flaring, with 8 elongated denticles on the

outer lip and a basal tooth producing a notch with the

abapical columellar tooth;

+ two strong columellar teeth, the adapical one

rounded, the abapical one squared; columella weakly

excavated with a few different in size denticles:

translucent callus shield covering the umbilical area

and partially the parietal area, with weak superficial

ridging on the parietal zone

+ base with 4 granular spiral cords; distance between

cords similar to the size of cords:

+ no umbilicus;

+ brown or cream with brown flames.

This species was originally described as belonging to

the genus Perrinia. But it is clear that some features of

this species, especially the two strong columellar teeth

and the inductural callus shield, are discordant with

the Perrinia characterization (see below) and match

on the contrary the main Clypeostoma features.

Clypeostoma adelon n. sp.

Figs 17D-L, Tables 12,13

Type material. Holotype (5.2 x 3.7 mm) MNAN (IM-

2000-327906). Paratypes: 4 MNHN (IM-2000-32797),

1 paratype CV.

Type locality. New Caledonia, Koumac,

MONTROUZIER, stn 1323, 20°41'S, 164°15'E, 82-

120 m.

Material examined. New Caledonia. BATHUS 2:

stn DW739, 22°35'S, 166°27'E, 465-525 m, 1 dd. —

StnnDW717, 22%44$, 1674178, 350-393, 1 dd —

MONTROUZIER: stn 1331, 20°40'S, 164°1 l'E, 55-57

m, 2 dd juv. — Stn 1315, 20°41'S, 164°15.E, 66-87 m,

1 dd. — Stn 1323, 20°41'S, 164°15'E, 82-120 m, 14 Iv

(with holotype and paratypes). — Stn 1321, 20°45',

164°15'E, 90-115 m, 1 dd, 1 dd juv.

Loyalty Islands. LIFOU 2000: stn 1461, 20°545,

167°02'E, 100-120 m, 3 dd sub. — Stn 1462, 20°47'S,

167°03'E, 70-120 m, 1 dd.

Bi SUV BST ÉMIS AT 28 'E 65 ul

dd. - BORDAU 1: stn DW1440, 17°11'S, 178°43"W,

190-308 m, 1 dd.

Figure 15 (scale bar: 5 mm).

Chilodontidae from the Pacific Ocean

Vanuatu. MUSORSTOM 8: sin DW1070, 15°375,

167°16'E, 184-190 m, 2 dd. —- BOA O0: stn CP2326,

15°4]'S, 167°03'E, 260-313m, 1 dd. - SANTO 2006:

stn EP28, 15°38'S, 167°05'E, 90-110 m, 1 dd. — Stn

ATS8S. 15°32!S, 16715, 87-150 d'dd

Solomon Islands. SALOMON 2: stn DW2169,

9C01'S, 159°06'E, 100-200 m, 2 dd. — Stn DW2234,

6°51'S, 156°24'E, 182-277 m, 3 dd, 8 dd'sub. =

SALOMONBOA 3: stn CP2840, 10°25'S, 161°22E,

121-180 m, 1 dd sub.

Distribution. New Caledonia, 57-465 m, 1v at 82-120

m; Loyalty Islands, 100-120 m (dd); Fiji, 83-190 m

(dd); Vanuatu, 110-260 (dd); Solomon Islands, 180-

182 m (dd).

Diagnosis. A rather small chilodontid species with an

elevated, conical shape and an angular periphery, with

an exsert dome-shaped protoconch, 5 granular cords

on the penultimate whorl, a subrounded aperture with

two teeth, a weakly convex base with 5 spiral cords

and an inductural callus covering completely the

umbilicus.

Description. Shell of small for the genus (height 5.2

mm, width 3.7 mm), higher than wide, rather thick,

conical; spire elevated, height 1.4x to 1.6x width, 3.2x

to 3.7 aperture height; anomphalous.

Protoconch about 200 um, of 1 whorl, translucent,

exsert dome-shaped, with a straight, thin terminal

Varix.

Teleoconch up to 5.9 almost flat whorls with 5

granular spiral cords on last whorls; distance between

cords similar in size to cords; angular periphery.

Suture canaliculated.

First whorl convex, sculptured by about 15 rather

strong prosocline threads;

interspace between them 2x to 3x size of threads; P3

and P4 appearing near end of whorl, granular. On

second whorl, axial threads of same size and same

interspace, connecting beads of cords. On third whorl,

PI and P2 resolving, thin, granular; beads of all cords

separated. On fourth whorl, P1, P2 and P3 similar in

size, P4 stronger; beads of P1 slightly vertically

elongated, beads of P4 stronger, horizontally

elongated; axial threads stronger, becoming thick ribs;

A-U. Clypeostoma nortoni (McLean, 1984). A-I. Philippines. A-F. Balicasag Is., PANGLAO 2004, stn P3, 100

m. A-C. 11.0 x 8.3 mm. D-F. Juvenil, 8.0 x 6.7 mm. G-I. Bohol Is., PANGLAO 2005, stn CP2340, 291-318 m,

12.5 x 10.1 mm (Marbol 35489). J-L. Taiwan, Bashi channel, TAÏWAN 2000, stn DW36, 305 m, 9.8 x 6.7 mm.

M-O. Indonesia, Kai Is., KARUBAR, stn CP16, 315-349 m, 9.5 x 7.6 mm. P-U. Solomon Is., SALOMON 2. stn

DW2191, 300 m. P-R. 8.9 x 6.7 mm. S-U. 6.9 x 6.1 mm.

V-X. Clypeostoma chranos n. sp., southeastern coast of Taiwan, holotype MNHN (IM-2000-32795), TAIWAN

2002, stn DW149, 258 m, 8.0 x 5.9mm.

C. VILVENS

Chilodontidae from the Pacific Ocean

S 1 appearing near half whorl. On fifth whorl, all cords

similar in size, except P4 stronger,; distance between

cords similar in size to cords. On last whorl, P4 the

strongest, making keel, their strong pointed beads

giving a stellate shape from a basal view.

Aperture subrounded; outer lip flaring beyond an

internal thickening, with 9 horizontally elongated

denticles corresponding to the external spiral cords of

last whorl and thinner similar denticles between them;

inner lip With 1 to 3 rounded denticles.

Columella straight, opisthocline, weakly excavated,

without two teeth, the adapical one rounded obtuse

and the abapical stronger and squared.

holotype

paratype |

Base moderately convex flat, with 5 granular spiral

cords, the most internal one partly covered by an

expansion of the inner lip; distance between cords 1%

to 2x size of cords; strong axial threads between cords.

Umbilicus completely filled by columellar callus;

umbilical area and internal parietal area covered by an

inductural callus shield, parietal part translucent.

Colour of teleoconch yellowish white to light brown;

protoconch off white.

Operculum corneous, multispiral with central nucleus,

light brown, translucent.

paratype 2

paratype 3 À

paratype 4

paratype CV

Table 13. Clypeostoma adelon n. Sp

Discussion. The new species is superficially close to

Clypeostoma cancellatum (Schepman, 1908) from

central Indo-Pacific (figs 16A-X), but this greater

species has a smaller H/W ratio (about 1.2-1.3) and a

sunken protoconch.

C. adelon n. Sp. is also similar to C. cecileae (Poppe,

Tagaro & Dekker, 2006) from the Philippines (figs

17A-C), but this slightly greater species lacks a SI

spiral cord, has spiral cords with less numerous beads

and has only 4 spiral cords on the base.

Etymology. Invisible (Ancient Greek: aônAoc, ov) —

with reference to the possible confusion of the new

species with two others, making it difficult to detect.

Genus Pholidotrope Herbert, 2012

Type species: Pholidotrope gloriosa Herbert, 2012

(by original designation) — North-western

Madagascar, west of Banc of Leven, 35-150 m.

Remarks. The main features for Pholidotrope

(Herbert, 2012) are a conical shape, a very small size

(H < 5 mm), a cancellate sculpture, a columella with a

single basal tooth and a U-shaped notch under it, a

parietal glossy inductural callus shield and especially

a strong, subterminal labral varix behind the outer lip.

Pholidotrope asteroeides n. sp.

Figs 17M-S, Table 14

Type material. Holotype (3.8 x 2.7 mm) MNAN (IM-

2000-32804). Paratype: 1 MNEN (1M-2000-32805).

4.9 >

4.8 3.4

53 4.6 1.4

4.5 1.4

.: Shells measurements in mm for types.

Type locality. New Caledonia, Norfolk Ridge,

NORFOLK 1, stn DW1675, 24°45'S, 168°09'E, 231-

25 3.

Material examined. New Caledonia. SMIB 8: stn

DW159, 24°46'S, 168°08' E, 241-245 m, 1 Iv

(paratype). —- NORFOLK 1: stn DW1675, 24°45S,

168°09'E, 231-233 m, 1 Iv (holotype).

Distribution. Southern New Caledonia, alive in 233-

241 m.

Diagnosis. À very small, much higher than wide,

chilodontid species with a conical shape, an angulated

periphery, 3 granular spiral cords, the abapical cord

much stronger with strong pointed beads and making

keel, an additional thinner cord on last whorl, a weak

columellar tooth, an almost flat base with 3 spiral

cords and no umbilicus.

Description. Shell of moderate size for the genus

(height up to 3.9 mm, width up to 2.7 mm), higher

than wide, rather thick, conical; height 1.4x to 1.5x

width, height 2.8x to 2.9x aperture height;

anomphalous.

Protoconch unknown (broken on the two available

samples).

Teleoconch up to 5.7 weaklÿy convex to nearly flat

whorls, bearing 3 spiral cords on the penultimate

whorl, the abapical cord much stronger than the other

cords, making keel, stellate with thick pointed beads:

strong axial prosocline ribs between cords; periphery

angulated. Suture visible, not canaliculated.

C. VILVENS

NOVAPEX I18(HS 11): 1-67, 10 mars 2017

First whorl convex, sculptured by prosocline, smooth,

moderately thick threads, interspace from about 2x the

width of threads; P2 and P3 appearing at end of whorl

or at begin of next whorl, weakly granular. On second

whorl, P2 and P3 quickly stronger, with rounded

beads made by intersection of cords and threads. On

third whorl, PI appearing, thinner than the other

cords; beads of P2 and P3 stronger, those of P3 a little

pointed; distance between cords similar to width of

cords. On fourth whorl, P3 much stronger than the

other cords, with sharply pointed beads and making

keel; PI and P2 similar in size; axial threads

thickening, making ribs; distance between ribs about

2x width of ribs. On last whorl, P4 visible, much

thinner than the other cords; beads of P3 very thick

and strongly pointed; periphery carinate, stellate in

shape because the peaks of P3.

holotype

Columella almost straight, more or less vertical, with a

small, weakly prominent basal tooth.

Aperture subelliptical, without angle; interior

nacreous, exterior with a thickened, rather low varix;

outer lip slightly thickened inside, with 10 to 12 ridge-

like denticles, that nearest columella making a U-

shaped notch between it and columella tooth; inner lip

somewhat flaring in its lower part.

Base almost flat, with 3 rather strong granular spiral

cords, more or less similar in size 10 P4; distance

between cords 1x to 1.5x width of cords; rather strong

axial ribs between cords, distance between them

similar to width of ribs; upper parietal part covered by

a thin inductural callus.

No umbilicus.

Colour of teleoconch brown with some

(especially those of P3) white; protoconch white.

beads

Table 14. Pholidotrope asteroeides n. sp.: Shells measurements in mm for types.

Discussion. Pholidotrope asteroeides n. sp. is rather

close to P. gloriosa Herbert, 2012 from Glorieuses

Islands (Madagascar area), but this similar in size

species has a smaller H/W ratio (1.18 - compare with

1.4 to 1.5 for the new species), three spiral cords of

the whorls more or less similar in strength with a

distance between cords greater (not similar) than the

cords, a much wider inductural callus and a shightly

stronger, more pointed columellar tooth.

The new species reminds a little Perrinia morrisoni

(Ladd, 1966) from Bikini Atoll, but this similar in size

species is much more depressed, has 4 spiral cords on

the penultimate whorls and lacks the external apertural

varix.

The new species weakly resembles Perrinia stellata

(A.Adams, 1864) from western Indian Ocean and Red

Sea (figs 20V-X), but this much greater species

(height up to 15 mm) has 4 spiral cords on the

penultimate whorls with a stronger PI with thicker

beads and a peripheral P4 with much blunter beads,

and stronger spiral cords on the base: it also lacks the

external varix on the border of the aperture.

Etymology. Star shaped (Ancient Greek: aotTEpoElon,

eç adjective) — with reference to the stellate shape of

the periphery of the shell.

Pholidotrope choiseulensis n. sp.

Figs 17T-W

Type material. Holotype (2.7x1.8 mm) MNHN (IM-

2000-32806).

Type locality. Solomon Islands SALOMON 2, sin

DW2234, 6°51'S, 156°24'E, 192-277 m, | dd.

Material examined. Solomon Islands. SALOMON

2: stn DW2234, 6°51'S, 156°24'E, 192-277 m, 1 dd

(holotype).

Diagnosis. A very small, much higher than wide,

chilodontid species with a conical shape, a more or

less rounded periphery, 3 granular spiral cords with

rather thick bead on the penultimate whorl, the

abapical cord slightly stronger with weakly pointed

beads, a weak columellar tooth, an almost flat base

with 3 spiral cords and no umbilicus.

Description. Shell of rather small size for the genus

(height 2.7 mm, width 1.8 mm), higher than wide,

rather thick, conical; height 1.5x width, height 3.4x

aperture height; anomphalous.

Protoconch bulbous, prominent, about

with a thin, weakly convex terminal varix.

Teleoconch of 5 weakly convex to almost flat whorls,

with 3 more or less similar in size spiral cords on the

penultimate whorl, the abapical cord a little stronger

on last whorl: rounded periphery. Suture visible, not

canaliculated.

First whorl convex, sculptured by rather indistinct

smooth prosocline threads; interspace similar in size

to width of threads. On second whorl, threads

thickening and more marked: P2 and P3 appearing at

mid whorl. On third whorl, PI appearing, thinner than

the other cords; all cords with rounded beads made by

intersection of cords and threads; P3 slightly stronger

than the other cords. On fourth whorl, all cords more

or less similar in size; interspace between cords

smaller than cords; beads of cords rather strong,

isolated from each other. On last whorl, P3 slightly

stronger than the other cords, with slightly pointed

1.25 whorl.

C. VILVENS

Chilodontidae from the Pacific Ocean

beads; P4 visible, thinner than the other cords; wide

axial ribs still present, connecting beads of the cords.

Columella almost straight, sightly opisthocline, with a

small, weakly prominent basal tooth.

Aperture subelliptical, without angle; interior

nacreous, exterior With a moderately thickened varix

rather far from the aperture rim; outer lip slightly

thickened inside, with 10 ridge-like denticles.

Base almost flat, with 3 rather strong granular spiral

cords, slightly thinner than P4; distance between cords

similar in size to width of cords; rather strong axial

ribs between cords, distance between them similar to

width of ribs; upper parietal part covered by a thin,

poorly visible inductural callus.

No umbilicus.

Colour of teleoconch cream with brownish flames on

the whorls; protoconch white.

Operculum corneous, multispiral with central nucleus

and short growing edge, light brown, somewhat

translucent.

Discussion. Pholidotrope choiseulensis n. sp. is rather

close to P. gloriosa Herbert, 2012 from Glorieuses

Islands (Madagascar area), but this similar in size

species has a smaller H/W ratio (1.18 against 1.5 for

the new species), much thinner spiral cords on the

whorls with a much greater distance between them

and much smaller beads, a much wider inductural

callus and a stronger, more prominent columellar

tooth.

The new species reminds only weakly P. asteroeides

n. sp. from New Caledonia (figs 17M-S), this slightly

greater species having a very different shape with

much more pointed beads on P3 and an angulated

periphery.

Etymology. Named after Choiseul Island (Solomon

Islands - native name Lauru), the type locality area.

Genus Danilia Brusina, 1865

Type species: Monodonta limbata Philipp, 1844 (by

monotypy) [= Monodonta tinei Calcara, 1839] (by

monotypy) — Recent, Mediterranean Sea.

Remarks. The main features for Danilia are a

turbinate shall with a cyrtoconoidal to almost conical

elevated spire, more or less convex whorls, a flat to

even sunken protoconch, a reticulated sculpture, a

half-moon shaped aperture, two columellar teeth with

the abapical one usually stronger and an exterior

subterminal varix along the outer lip of the aperture.

Danilia shell size number | which spiral number | columella | colour

shape of S1/T1 lirae in |ofcords | -

cords present | aperture | on base | columellar

on } teeth

penul-

timate

whorl

angulosa sl. deeply yellowish or

Cyrtoco- excavated | ochre beige

noidal - ab.

strong

squared,

ad. weak

discordata conical “e deeply brown +

excavated | possibly

- ab. brown

strong maculations

squared,

ad. very

weak

eucheliformis | cyrtoco- | 9.6x7.6 S1, 52 deeply off white +

noidal + S4 excavated | greyish and

- ab. brownish

strong marks

rounded,

ad. weak

stroggylon sl. TS 50 6 S1, S2 D 5-6 weakly light-brown

n. SP. Cyrtoco- + S4 (strong) excavated | + axial

noidal - ab. reddish

strong, ad. | brown

weak flames

frontal

view

Table 15. Comparison of conchological features of small central Indo-Pacific Danilia species (sl. = slightly.

ab. = abapically, ad. = adapically).

C. VILVENS

Danilia angulosa Vilvens & Héros, 2005

Figs 18A-H, Table 15

Danilia angulosa Vilvens & Héros, 2005: 54-55, figs.

1-7. Type locality: New Caledonia, 220 m.

Danilia angulosa — Poppe, Tagaro & Dekker, 2006:

L FA

Danilia angulosa — Herbert, 2012: 414-415.

Material examined. New Caledonia. EBISCO: sin

DW2547, 21°06'S, 158°36'E, 356-438 m, | dd.

Vanuatu. SANTO 2006: stn DS99, 15°335,

167°17"E, 100-105 m, 1 dd, 1 dd sub.

Solomon Islands. SALOMON 2: stn DW2173,

9°06S, 159°02'E, 201-231 m, 1 dd. -

SALOMONBOA 3: stn DW2790, 08°19'S, 160°37'E,

314-586 m, 1 dd juv. — Stn CP2804, 09°155,

161°2l'E, 150-175 m, 1! dd.

French Polynesia, Australes Archipelago.

BENTHAUS: stn DW1957, 23°19'S, 149°29'W, 558-

1000 m, ! dd. — Sin DW1894, 27°40'S, 144°22'W, 100

m, | dd.

Distribution. New Caledonia, 150-356 m, Iv at (dd -

using data from the original description); Loyalty

Islands, 120-150 m, 1v at 70-120 m; Vanuatu, 469-472

m (dd); Fiji, 105-469 m (dd); Solomon Islands, 175-

314 m (dd); French Polynesia, Australes Archipelago,

100-558 m (dd); Tonga, 555-581 m (dd).

Remarks. The main characteristics of this species are:

+ height up to about 8.0 mm, width up to about 7.5

mm; about 5 whorls:

+ protoconch sunken:

+ on first whorl, prosocline threads; PI and P3

almost immediately, P3 stronger, P2 à bit later: on

second or third whorl, P3 producing a shoulder;

subsutural ramp oblique and slightly convex; S1, S2

and S3 appearing later; on fourth whorl, additional

tertiary cords T2 and T3 appearing respectively

between P2 and S2, and between S2 and P3; P4

becoming visible on fifth whorl:

+ on last whorl, P3 peripheral, making keel: S4

visible; possibly a tertiary cord between S3 and P4 or

between P4 and S4; beads of P3 especially pointed:

+ aperture subcircular, outer lip with 20 to 25

plications with an exterior subterminal varix along it;

+ columella straight, opisthocline, deeply

excavated, with two teeth, the adapical one much

weaker, rounded (possibly reduced to a swelling), the

abapical one squared:

+ base moderatly convex with 7 to 8 granular spiral

cords;

+ anomphalous; whitish, translucent callus shield

covering the umbilical area and partially the parietal

area;

+ yellowish or ochre beige, with brown dots on

some spiral cords or brown flammules on whole

whorls.

NOVAPEX I8CHS 11): 1-67, 10 mars 2017

The additional samples from Solomon Islands and

Australes Archipelago extend the distribution area to a

larger area of the the Central Pacific. But this species

seems rather rare and found only at rather great

depths.

Danilia discordata Vilvens & Héros, 2005

Figs 181-T, Table 15

Danilia discordata Vilvens & Héros, 2005: 60, figs.

25-27. Type locality: Vanuatu, 375-397 m.

Danilia discordata — Poppe, Tagaro & Dekker, 2006:

32.

Danilia discordata — Herbert, 2012: 419.

Material examined. Vanuatu. BOA !, sin DW2459,

16°11'S, 167°20'E, 336-353 m, 1 dd. — Stn CP2465,

16°43'S, 1 dd.167°59'E, 770-799 m.

Solomon Islands. SALOMON I: stn CPI831,

10°12'S, 161°19'E, 135-325 m, 1 dd. — Stn DW1856,

9°46'S, 160°52'E, 254-281 m, 1 dd. - SALOMON 2:

stn DW2183, 8°17'S, 160°O0'E, 489-491 m, 2 dd juv.

— Stn CP2200, 7°44'S, 158°30'E, 325-331 m, 1 dd. -

Stn DW2204, 7°14'S, 158°29'E, 286-423 m, 3 dd. -

Stn DW2229, 6°36'S, 156°20'E, 315-418 m, 1 dd. —-

Stn CP2248, 7°42'S, 156°25'E, 650-673 m, 1 dd. —

SALOMONBOA 3: stn DW2811, 09°47'S, 161°30'E,

228-238 m, 2 dd. — Stn CP2853, 09°47'S, 160°54'E,

264-285 m, 1 dd. — Stn DW2859, 09°19'S, 160°19'E,

320-330 m, 1 dd.

Philippines. MUSORTOM 3: stn CP105, 13°52'N,

120°30'E, 398-417 m, 1 dd. —- PANGLAO 2005: stn

DW2362, 08°57'N, 123°33'E, 679-740 m, 1 dd. — Stn

DW2364, 0O9°0]'N, 123°26'E, 427 m, 1 dd.

New Caledonia. LAGON: stn DW475,

163°11'E, 415-460 m, 1 dd.

Loyalty Islands. MUSORSTOM 6: stn DW398,

20°47'S, 167°06'E, 320-370 m, 1 dd.

18°36'S,

Distribution. Vanuatu, 353-770 m (dd - using data

from the original description); Solomon Islands, 238-

650 m (dd); Philippines, 417-679 m (dd): New

Caledonia, 415-460 m (dd); Loyalty Islands, 320-370

m (dd).

Remarks. The main characteristics of this species are:

+ height up to 12.8 mm, width up to 9.5 mm:

+ about 5 to 5.5 convex whorls;

+ _ protoconch sunken into first teleoconch whorl:

+ conical in shape:

+ on first whorl, prosocline threads: interspace

between ribs 2 times larger than width of ribs; on

second whorl, P2, P3 and P4 almost simultaneously,

similar in size; PI a bit later, weaker: SI and S2 from

end of second whorl to end of third whorl, S3 absent:

all cords granular with small beads, except P4 with

scaly, sharp pointed beads; on third whorl, beads of P3

and P4 sharp; on fourth whorl, Ti appearing, usually

T1 appearing between PI and SI and T2 between S1

and P2; P3 and P4 the strongest; beads of P3 and P4

C. VILVENS

Chilodontidae from the Pacific Ocean

very sharp, horizontally oriented; beads of PI pointed,

adapically oriented; beads of other cords rounded,

reduced, sometimes even flattened; axial threads still

present, visible but rather thin; S4 emerging from

suture; on last whorl, P3 and P4 peripheral, making

weak keels; possibly additional tertiary cords;

+ aperture rounded to elliptical, flaring; outer lip

with rather strong varix; inside flaring, lirate within

with about 20 rather weak plications, the 3 ridges near

inner lip stronger;

+ columella straight, weakly opisthocline, deeply

excavated, with two teeth, the adapical one much

weaker and rounded, the abapical one stronger and

squared;

+ base moderately convex, sculptured with 6 to 8

granular spiral cords, similar in size; interspace

between cords less than or equal to cords, filled with

broad axial threads.

+ anomphalous: whitish, translucent callus shield

covering the umbilical area and partially the parietal

area;

+ brown, sometimes with large brown strip-like

maculations.

At the time of the original description, this species

was only known by the holotype (with protoconch

missing). The additional samples from Solomon

Islands, a neighbouring area of the original description

zone (Vanuatu), are useful to confirm or clarify the

features of this species (including the protoconch), and

also to highlight an intraspecific variability of the

shell. These new records extend the distribution area

to a larger area of the Central Pacific, from Philippines

to New Caledonia. But this species seems rather rare

and only found at great depths.

Danilia eucheliformis (Nomura & Hatai, 1940)

Figs 19A-D, Table 15

Monodonta eucheliformis Nomura & Hatai, 1940: 61,

pl IL figs la-b. Type locality: Japan (north-eastern

Honshu).

Danilia eucheliformis — Vilvens & Héros, 2005: 56,

figs. 15-18.

Material examined. Taiwan. TAIWAN 2001: stn

CPYA DAS IS PISE 2200 Ir dd PAIN AN

2002: stn DW149, 22°19'N, 121°29'E, 258 m, 3 dd.

Indonesia, Kai Islands. KARUBAR: sin DWI8,

05°18:S,13320LE, 205-212, l'dd)

Distribution. Japan, 50-200 m (fide Habe, 1961:

Kuroda, Habe & Oyama, 1971); Taiwan, 220-420 m

(dd); Indonesia, Kai Islands, 205 212 m (dd).

Remarks. The main characteristics of this species are:

+ height up to 9.6 mm, width up to 7.6 mm; up 5.5

whorls;

+ _ protoconch sunken into first teleoconch whorl;

+ cyrtoconoidal in shape, with convex whorls;

rounded periphery;

+ on first whorl, prosocline threads, distance

between at least 2x wider than threads; on second

whorl, P4, P3, P2 and P1 successively; S1 and S2 at

third whorl, quickly similar in size to Pi; S3 absent; S4

visible on last whorl; all cords granular with small,

largely separated, pointed beads; distance between

cords about 1.5x size of cords; axial threads still

visible, connecting beads of cords, producing a

reticulate pattern;

+ aperture rounded to ovate; outer lip with an

external rather strong external varix and about 20-25

rather weak plications inside;

+ columella straight, vertical, deeply excavated,

with two teeth, an abapical rounded, stronger one and

an adapical weaker one;

+ base moderately convex, sculptured with 5-6

granular spiral cords, similar in size; interspace

between cords Similar in size to cords, with thin axial

threads;

+ anomphalous; whitish, translucent callus shield

covering the umbilical area and partially the parietal

area;

+ off white with greyish and brownish marks.

Danilia stroggylon n. sp.

Figs 19E-L, Tables 15,16

Type material. Holotype (8.0 x 5.9mm) MNHN (IM-

2000-32798). Paratypes: 2 MNHN (IM-2000-32799).

Type locality. Southeastern coast of Taiwan,

TAIWAN 2002, stn DW149, 22°19'N, 121°29'E, 258

Figure 16 (scale bar: 5 mm).

A-X. Clypeostoma cancellatum (Schepman, 1908). A-I. Samples with S1 spiral cord. A-F. Philippines. A-C.

Pamilacan Is., PANGLAO 2004, stn T36, 95-128 m, 7.4 x 6.2 mm. D-F. Aliguay Is., 100-150 m, 9.6 x 6.9, coll.

C.Vilvens. G-L. Eastern New Caledonia, BATHUS 1, stn CP688, 270-282 m, 6.6 x 5.2 mm. J-X. Samples

without SI spiral cord. J-O.Solomon Islands. J-L.S ALOMON 1, stn CP1758, 180-187 m, 8.4 x 6.7 mm. M-O.

SALOMON 2, stn DW2254, 150 m, 9.4 x 7.6 mm. P-R. New Caledonia, BATHUS 2, stn DW732, 236-264 m.

7.6 x 5.5 mm. S-U. Taiwan, TAIWAN 2002, stn DW117, 126-153 m, 6.9 x 5.6 mm. V-X. Tonga Islands,

BORDAU 2, stn DWI1601, 200-487 m, 4.3 x 3.1 mm.

C. VILVENS

Chilodontidae from the Pacific Ocean

Material examined. Taiwan. TAIWAN 2002: stn

DW149, 22°19'N, 121°29'E, 258 m, 1 dd (holotype

and paratypes).

Indonesia, Kai Isl. KARUBAR, stn DWIS8, OS°I18SN,

IS 0E 205-2120 Liv:

Distribution. Taiwan, 28 m (dead); Indonesia, Kai Is.,

205-212 m (living).

Diagnosis. À small size chilodontid Danilia species

with a rather elevated, weakly cyrtoconoidal shape,

almost flat whorls and a rounded periphery, 6

granular, similar in size cords on the penultimate

whorl, a subrounded aperture with a strong external

varix on the border, a convex base with 5 spiral cords

and a granular inductural callus shield covering

completely the umbilicus.

Description. Shell of small size for the genus (height

up to 7.7 mm, width 5.5 mm), higher than wide, rather

thick, weakly cyrtoconoidal to almost conical; spire

elevated, height 1.3x to 1.4x width, 3.0x to 3.2x

aperture height; anomphalous.

Protoconch of about 1 whorl, diameter about 200 um,

rather flattened, sunken into the first teleoconch whorl,

without terminal varix.

Teleoconch to 5.2 weakly convex to almost flat

whorls, with 6 rather strong spiral granular cords on

penultimate whorl; distance between cords similar in

size to cords; rounded periphery. Suture visible,

canaliculated.

First whorl convex, sculptured by about 20 rather

strong, slightly prosocline threads:

interspace between them 2x size of threads; P4

resolving before end of whorl, P3 and P2 a bit later,

all granular. On second whorl, PI appearing near

holotype

paratype |

suture; P4 the strongest, PI the weakest; beads of all

cords produced by intersection between cords and

threads, separated; beads of cords quickly pointed

after mid of whorl, except those of PI. On third whorl,

all cords much stronger, P4 the strongest with very

pointed beads; axial threads becoming strong ribs;

distance between cords about 1.5x size of cords; S1

and S2 appearing near end of whorl; S3 absent. On

fourth whorl, pointed beads of cords becoming blunt,

beads of P1 slightly vertically elongated; axial ribs

still present and strong, making with cords a reticulate

pattern. On last whorl, all the 6 cords similar in size,

distance between them similar to the size of cords; S4

fully visible, emerging from suture, weaker than the

other cords.

Aperture subrounded, flaring from internal thickening;

strong external varix making the border of the outer

lip (paratypes) or separated from this border by a

thickening made by accumulation of growth lines;

outer lip with 10 rather strong plications on the

border; inner lip with a stronger basal denticle,

producing a notch with the columella.

Columella straight, slightly opisthocline, weakly

excavated, with two teeth, the abapical one stronger,

pointed and possibly squared, the adapical one much

weaker, rounded, possibly only inflated.

Base moderately convex, with 5-6 thick granular

spiral cords, weaker than the cords of the last whorl,

distance between cords 1x to 1.5x size of the cords;

axial threads between cords.

Umbilicus completely filled by columellar callus;

umbilical and parietal areas covered by an inductural

callus shield (only visible on holotype).

Colour of teleoconch light-brown, with rather wide

axial reddish brown flames; protoconch and base

whitish.

paratype 2

Table 16. Danilia stroggylon n. sp.: Shells measurements in mm for types.

Figure 17 (scale bars: 1 mm)

A-C. Clypeostoma cecileae (Poppe, Tagaro & Dekker, 2006), Philippines, Mactan Is., paratype MNEN (Moll

5299), 50-100 m, 6.4 x 4.2 mm.

D-L. Clypeostoma adelon n. sp. D-F. Holotype MNHN (1M-2000-32796), New Caledonia, Koumac,

MONTROUZIER, stn 1323, 82-120 m, 5.6 x 3.8 mm. G-I. Loyalty Islands, Lifou, LIFOU 2000, stn 1462, 70-

120 m, 5.3 x 3.1 mm. J-L. Fiji, Viti Levu, SUVA 2, stn BS18, 83 m, 4.7 x 2.9 mm.

M-S. Pholidotrope asteroeides n. sp., New Caledonia. M-P. Holotype MNHN (IM-2000-32804), Norfolk Ridge,

NORFOLK 1, stn DW1675, 231-233 m, 3.8 x 2.7 mm. Q-$. Paratype MNHN (1M-2000-32805), SMIB 8. stn

DW159, 241-245 m, 3.9 x 2.6 mm.

T-W. Pholidotrope choiseulensis n. sp., holotype MNHN (1M-2000-32806), Solomon Islands, SALOMON 2.

stn DW2234, 192-277 m, 2.7 x 1.8 mm.

C. VILVENS

Chilodontidae from the Pacific Ocean

Discussion. Although the ontogeny of the spiral cords

of the new species sounds very similar to the one of D.

eucheliformis (Nomura & Hatai, 1940) from Western

Pacific (figs 19A-D), the new species 1s different by

its size (smaller for a similar number of whorls), the

global shape of the shell (much more conical), the

shape of the whorls (much less convex) and the shape

of the spiral cords (similar in size with interspace

similar in size to cords).

D. stroggylon n. Sp. may also be compared to D.

galeata Vilvens & Héros, 2005 from Indonesia, but

this proportionally greater size species has a more

elevated spire, more numerous spiral cords on the last

whorls, a spiral cord P4 making keel giving to the

shell a subangular periphery and a more reduced,

obtuse abapical columellar tooth.

One can wonder about the huge gap between the two

distribution areas (Taiwan and Indonesia), but it's a

fact that the Indonesian sample match perfectly the

description of the Taiwanese new species.

Etymology. Stocky (Ancient Greek: 6tpoyyvAog, n,

Ov) — with reference to the stocky shape of the shell,

producing a feeling of solidity.

Genus Chilodonta Etallon, 1859

Type species: Chilodonta clathrata Etallon, 1862 (by

subsequent designation Bayan, 1874, Bulletin de la

Société Géologique de France, ser. 3, 2: 335) —

Jurassic (Oxfordian), France.

Remarks. The main features for Chilodonta are

(Cossmann, 1918) a solid shell with a rather elevated

spire, a reticulated sculpture all over the whorls and

especially a complex apertural teeth system: one

strong, prominent columellar tooth, a parietal tooth, an

strong upper palatal tooth opposite to the columellar

tooth and one or two weaker basal palatal teeth.

Chilodonta suduirauti Poppe, Tagaro & Dekker, 2006

Figs 19M-X

Chilodonta suduirauti Poppe, Tagaro & Dekker, 2006:

30-31, pl.7, figs.1-2.

Figure 18 (scale bars: 5 mm).

Material examined. Philippines. PANGLAO 2005:

stn DW2364, O9C01'N, 123°26'E, 427 m, 3 dd, 4 dd

sub, 2 dd juv. — Stn CP2384, 08°46'N, 123°16'E, 624-

647 m, 1 dd. — Stn DW2401, 0993 l'N, 123°40'E, 397-

410 m, 1 dd.

Solomon Islands. SALOMON 1: stn DWI1825,

9°50'S, 160°58'E, 340-391 m, 1 dd. - SALOMON 2:

stn DW2183, 8°17'S, 160°00'E, 489-491 m, 1 dd. —

Stn CP2199, 7°43'S, 158°30'E, 296-304 m, 1 dd. —

SALOMONBOA 3: stn CP2786, 08°31'S, 160°4l'E,

320-520 m, 1 dd.

New Caledonia. EBISCO: stn DW2555, 21°045,

158°35'E, 500-614 m, 1 dd.

Indonesia, Kai Islands. KARUBAR: stn DWI13,

05°26'S, 132938'E, 417-425 m, 1 dd, 1 dd sub. — Stn

DW28, 05°31'S, 132°54'E, 448-467 m, 1 dd 1 dd sub.

Distribution. Philippines, 410-624 m (dd - including

data from Poppe et al., 2006); Solomon Islands, 304-

489 m (dd); New Caledonia, 500-614 m (dd);

Indonesia, Kai Islands, 425-448 m (dd).

Remarks. The main features of this species are:

+ height up to 8.4 mm, width up to 7.0 mm;

+ spire rather elevated, weakly cyrtoconoidal;

+ teleoconch of up to 6 weakly convex whorls;

+ first whorl with thin prosocline threads: second

whorl with Pi (i=124), P4 sunken under the suture;

S1 and S2 appearing quickly, similar in size to Pi, all

granular with pointed, separated beads, connected by

axial threads, giving a reticulated pattern; P4 emerging

from suture on fourth whorl; possibly Ti between Pi

and Si near end of last whorl;

+ one very strong columellar tooth projecting out of

columella border;

+ aperture elliptical, with a translucent inductural

callus shield on the parietal wall; outer lip with a

strong palatal tooth and 4-5 denticles; basal lip with a

strong tooth with an U-shaped notch between it and

the columellar tooth:;

+ base almost flat, with 5 to 7 granular spiral cords;

distance between cords similar to size of cords; axial

thin threads between cords;

+ no umbilicus;

+ white to light brownish white.

A-H. Danilia angulosa Vilvens & Héros, 2005. A-D. New Caledonia, Bellona, EBISCO, stn DW2547, 21°065,

7.1 x 6.1 mm. E-H. Solomon Islands, SALOMON 2, stn DW2173, 201-231 m, 8.0 x 3.1 mm.

I-T. Danilia discordata Vilvens & Héros, 2005. I-L. Solomon Islands, SALOMON 2, stn DW2229, 315-418 m,

12.8 x 9.6 mm. M-P. SALOMONBOA 3, stn DW2811, 228-238 m, 12.1 x 9.2 mm. Q-T. Vanuatu, BOA 1, stn

DW2459, 336-353 m, 11.4 x 8.4 mm.

C. VILVENS

Chilodontidae from the Pacific Ocean

The records here reported extend substantially the

distribution of this species, especially considering the

single record in north-western New Caledonia

(Bellona Reef). It can be now considered that

Chilodonta suduirauti 1s a Western Indo-Pacific

widespread species. No variability per area is

observed: it can only be noticed that some Solomon

samples are slightly less elevated and light brownish

white instead of white coloured, while the Indonesian

samples have slightly more convex whorls and are

brownish coloured.

Genus Perrinia H. & A.Adams, 1854

Types species: Monodonta angulifera A.Adams, 1854

in 1853 (by subsequent designation Pilsbry, 1889) —

Recent, Philippines.

Remarks. The main features for Perrinia are a rather

small to medium size (H < 20 mm), an elevated spire

with slightly convex to almost flat whorls, an axial

sculpture on the first whorls with addition of axial

cords on the next whorls producing a cancellate to

almost foveolate pattern, a single basal columellar

tooth without U-shaped notch under it and the absence

of any inductural callus shield (Herbert, 2012).

Perrinia shell size number axial colour frontal

shape (mm) |ofspiral | threads vieW

cords

on penul-

timate

angulifera conical, 16 5 d'A SE greyish

spire xs il white to

elevated pale buff

elisa conical, 20 3 M off white to

spire x14.5 pale beige

elevated 2

cantharidoides | conical, : 5 NA SI + with with

n. SP. spire very x5.6 S4 possible

elevated bownish

flames

guadalcana- coeloconoiïi- | 7x5.2 |5 de S1 + off white

lensis n. Sp. dal on S4 with

adapical possible

part and bownish

conical on flames

abapical

part

squamocari- conical, 9.2 3 (4) nf - nacreous

nata spire mod. x6.5 white with

elevated pinkish

flames

Table 17. Comparison of conchological features of some central Indo-Pacific Perrinia species

(mod. = moderately).

Figure 19 (scale bars: 5 mm).

A-D. Danilia eucheliformis (Nomura & Hatai, 1940), Taiwan, off Tashi, TAIWAN 2001, stn CP74, 24°51'S

121°59'E, 220 m, 7.6 x 7.6 mm.

E-L. Danilia stroggylon n. sp., Taiwan, TAIWAN 2002, stn DW149, 258 m. E-H. Holotype MNHN (IM-2000-

32798), 7.1 x 5.4 mm. I-L. Paratype MNHN (I1M-2000-32799), 7.7 x 5.5 mm.

M-X. Chilodonta suduirauti Poppe, Tagaro & Dekker, 2006. M-R. Solomon Islands. M-O0. SALOMONBOA 3.

stn CP2786, 320 520 m, 8.6 x 7.0 mm. P-R. SALOMON 1, stn DW1825, 9°50'S, 160°58'E, 340-391 m. 9.2 x

6.6 mm. S-U. New Caledonia, Western Bellona, EBISCO), stn DW2555, 500-614 m, 6.1 x 5.3 mm. V-X.

Indonesia, Kai Islands, KARUBAR, stn DW28, 448-467 m, 7.3 x 6.4 mm.

C. VILVENS

Chilodontidae from the Pacific Ocean

Perrinia angulifera (A.Adams, 1853)

Figs 20A-I, Table 17

Monodonta angulifera A.Adams, 1853: 176. Type

locality: Puerto Galero, Mindoro Is., Philippines, 11

Tectaria montrouzieri P.Fischer, 1878: 212-213. Syn.

Cantharidus pliciferus Schepman 1908: 43, pl. 3, fig.

3. SyYn.

Perrinia plicifera — Poppe, Tagaro & Dekker, 2006:

45, pl. 15, fig. 7. Syn.

Perrinia angulifera — Herbert, 2012: 465-470, figs 52-

S5r

Material examined. New Caledonia. LAGON: stn

WP 22 00020 nt dd Sin DWE,

22 NS MGM ES dd SnD\N PS 222210S

IOOPTE DE mn IN Sin DWS2 0240S

l'OMS nn eddsub = Sim iD\WS6 22410IS

COS TS MDN PIE 22205

ICOSLEND2 im, 2,4 = Sin” DWSO22%)S;

IOGPSIE 35 un INdd "Sin DW8S, 22232;

COS 220 NI Sin DV O2 22002715

LOGSTE, 24 m0 1 dd. =" Sin DWO4, 22230;

CCS AE En, AN = Sin IDDN 104 22%6!$S;:

COCO 2 on IR Sin DMC 22253:

IOOZÆTE 45 on dd =" Sin DW125, 22%30$;

IG AUDE INT Sd Sin DIS, 22252S;

166°48'E, 31-33 m, 2 dd. — Sin DW154, 22°335,

NOGAOIE, “29 mm, 1" dd.….—-Sin-DW201,-22°00$,

CSS nn Id Sin DW233, 22255,

IGOAGE 50m did Sin DW264 "224190$,

COPIE MOMENT ANG Sin DN266, 22222);

ICE I in IN dd Sin DW269 22218,

ICONE 20 un ANG = Sin, DW2 T0 275$;

IGG2RE 22 un dd Sin DW27S, 22444;

166°18'E, 19 m, 1 dd. - Stn DW285, 22°24%,

166°26'E, 19 m, 1 dd. — Stn DW313, 22°40$S,

166/50E, 3000 “1 dd.."Stn-DW348, 224%,

COS SES nn NO Sn D WST2022252%;

IOHSOUE 65 ox AN dd” Sin. DWYOS 211,

IG DE Sn dd Sin DWTSS 2102

IOSSSE 63 ni dd Sir DNre921E05'S;

IGSAE 29, ml dd Sin DWS63 20730,

I65S°07E, 28 m, l'dd'sub. Sin DWO012, 20575,

IOMIE IS on dd Sin DWAIDID 20500;

IGSSHE IG on Aid = SnADMIOIr "200$;

IG SE 21 IN dd = SmADMIU27 20085;

I6SIE 29 on dd Sn DM I0S AL MOSES

IOSSAIE,. 25 nn dd = Sin D MIOS MOSS.

164°00'E, 25 m, 1 dd. — Sin DW1094, 19°54S,

IGSPNE 260, INA Sn DNA IRS,

IGSAAIE, 53 mn, Ad CORALIE 2""stn DW21

20°36'S, 161°02'E, 86 m, 1 dd. — Campagnes d'essais

1987: stn DE29, 22°35'S, 166°41'E, 29-33 m, 1 dd. —

Stn DES58, 22°1l'S, 166°11'E, 17-18 m, 2 dd. — Stn

DE6S, 22210S, 166222 ét Add = BATAUSAE

stn DW1233, 22°24'S, 166°48'E, 45-50 m, 40 Iv. —

MONTROUZIER: stn 1260, 20°44'S, 165°14'E, 49-

59 m, 4 dd, 1 dd juv. — Stn 1261, 20°46-4TS, 165°15-

L6'E, 45-56 m, 6 1v, 1 dd juv. — Stn 1277, 20°345,

164°16'E, 0-2 m, 1 dd. — Stn 1299, 20°34'S, 164°13'E,

12-14m, 1 dd juv. — Stn 1304, 20°395,

164°13'E,12-15m, 1 dd — Sin 1505, 207368,

IGAALE, 12-15, 2 dd ju. — Sin 1307,-203%8;

164°10'E, 12m, 6 dd juv. — Stn 1308, 24°405,

164°15'E, 15-20 m, 1 dd. — Stn 1309, 20°415,

164°13'E, 18m, 1 dd juv. — Stn 1312, 20°405,

164°15'E, 26-40 m, 2 dd sub. — Stn 1314, 20°405,

164°15'E, 30-63 m, 2 dd, 2 dd sub, L dd yuv. = Sin

1315, 20°41'S, 164°15'E, 66-87 m, 2 dd. — Stn 1321,

20°45'S, 164°15'E, 90-115 m, 1 dd juv. — Stn 1372,

22°20'S, 166°13'E, 50 m, 1 dd juv. — Stn 1492,

22°20'S, 166°26'E, 20 m, 1 dd. - NORFOLK I: stn

DW1673, 23°30/S .ACSOUNE, 0278 1 MINUTES

EBISCO: stn DW2608, 19°33'S, 158°40'E, 393-396

m, | dd juv.

Loyalty Islands. LIFOU 2000: stn1648, 20°54S

167°03'E, 150-200 m, 3 Iv juv.

En SUVA 2:%stin BSI8, 18" 105 1628 FE 65105

dd. — Sin CP65. 17P48S, L77AS/0TE, 57, Sad

SUVA 4: stn DW26, 18°24'S, 178°04'E, 42-43 m, 4

dd, —Stn DW44,17°52$, 197419, 88000 dd, 244

sub. — Stn DW62, 17°48'S, 177°13/04'E, 32 m, 9 dd.

Wallis Island. MUSORSTOM 7: stn DW2, 13°22'S,

176°11'W, 52-55 m, 1 dd

Vanuatu. MUSORSTOM 8: stn CP1131-1132,

15°38'S, 167°03-04'E, 140-182 m, 1 dd. — Stn deep

water "mixed", 1 dd.

Indonesia, Kai Islands. KARUBAR: syn DWOI,

05°46'S, 132°10'E, 156-305 m, 1 dd sub. — Stn DWI18,

OSATSS 1SSO0IE 205212 md;

Philippines. PANGLAO 2004: stn L69-73, 09°31"N,

123°41'E, 90-98 m, 9 dd juv.

Distribution. New Caledonia, 2-393 m, lv at 45-50 m:;

Loyalty Islands, 1v at 150-200 m; Fiji, 32-83 m (dd);

Wallis Island, 52-55 m (dd); Vanuatu, 140 182 m

(dd); Indonesia, 13-205 (dd - using also data of

Schepman, 1908); Philippines, 90-98 m (dd - using

also data of Poppe et al., 2006 /P. plicifera]).

Remarks. The main characteristics of this species are:

+ height up to about 16.0 mm, width up to about

IS tn:

+ protoconch nearly flat, at the same level as the

first teleoconch whorl;

+ teleoconch up to 7.5 almost straight whorls;

+ first whorl with axial threads only; PI, P2, P3

appearing at end of whorl, granular; P4 covered by

suture; SI appearing about half a whorl later, quickly

as strong as other cords; TI at third whorl, between P1

and Sl;

+ penultimate whorl with 5 spiral cords with axial

threads between them; PI with sharp pointed,

triangular, adapically oriented beads, possibly a bit

scaly;, P3 as strong as Pl, with slightly axially

elongated beads; occasionally, T2 may appear

between PI and T1;

C. VILVENS

+ _P4 peripheral on last whorl, thin, at the beginning

of the base;

* angular periphery without keel:

* moderately convex base with 5-6 (sometimes up

to 8) spiral cords, axial threads between them:

+ one moderately prominent columellar tooth at

second abapical third;

+ outer lip thin, thicker internally, with up to 16

Spiral lirae running inside, abapical ones possibly

terminating near columella and producing small

denticles below columella tooth:;

+ no umbilicus;

+ greyish white to pale buff.

Perrinia elisa (Gould, 1849)

Figs 20J-L, Table 17

Trochus elisus Gould, 1849:

Pickering, Singapore, dredged.

Calliostoma (Perrinia) Elisa [sie] — Schepman, 1908:

68.

Perrinia elisa — Poppe, Tagaro & Dekker, 2006: 43,

pl.15, fig. 4,6.

92. Type locality:

Material examined. Philippines. MUSORSTOM 3:

stn DR140, 11°43'N, 122°34'E, 93-99 m, 12 dd sub. —

PANGLAO 2004: stn T26, 09°43'N, 123°49'E, 123-

135 m, 1 dd. —- AURORA 2007: stn DW2758,

1S"55N, 121508, 151-173 m, 1 dd.

Distribution. Philippines, 50-200 m (using also data

of Poppe et al., 2006); Indonesia, 32-90 m (using data

of Schepman, 1908); Vietnam (fide Poppe et al.

2006).

Remarks. The main characteristics of this species are:

+ height up to about 20.0 mm, width up to about

14.5 mm:

+ protoconch nearly flat, at the same level as the

first teleoconch whorl;

+ teleoconch up to 7.5 almost straight whorls;

+ first whorl with poorly distinct, low axial ribs, PI,

P2, P3 appearing at end of whorl, granular; P4

covered by suture; on next whorls, P3 the strongest

with horizontally elongated beads, PI almost as string

as P3 with axially elongated beads, P2 the weakest:;

axial ribs connecting beads of cords, more or less

orthocline on first whorl, clearly prosocline on the last

whorls; suture strongly channelled: P4 peripheral on

last whorl, thin, at the beginning of the base: P3

making keel:

+ angular periphery without keel:

+ moderately convex base with 5-6 spiral cords,

distance between smaller then cords; rather thick,

close axial threads between them:

+ one median, poorly prominent columellar tooth

with a second abapical tooth below:

+ outer lip thin, thicker internally, with up to 11

spiral lirae running inside, never reaching aperture

TIM;

NOVAPEX I8CHS 11): 1-67, 10 mars 2017

+ no umbilicus (covered by columellar callus);

+ off white to pale beige.

Perrinia cantharidoides n. sp.

Figs 20M-U, Tables 17, 18

Type material. Holotype (8.4 x 5.4 mm) MNHN (IM-

20000-32800). Paratypes: 3 MNHN (1M-2000-32801).

Type locality. New Caledonia, Chenal de Koumac,

MONTROUZIER, stn 1315, 20°4l'S, 164°15'E, 66-87

Material examined. New Caledonia. VAUBAN

1978-1979: Sud NC, stn DWI10, 22°17'S 167°05'E, 80

m, ! dd. - MUSORSTOM 4: sin DW231, 22°345,

167°10'E, 75 m, 2 dd, 2 dd sub. —- CORAIL 2: stn

DW26, 20°22S, 161°05E, 62 m, 1 dd. -

MONTROUZIER: stn 1315, 20°41'S, 164°15'E, 66-87

m, 7 dd (with holotype and paratypes).

Vanuatu. BOA !: stn CP2476, 16°42'S, 167°53'E, 91-

103 m, 3 dd. —- SANTO 2006: stn AT74, 15°375,

167°10'E, 50-64 m, 2 dd.

Distribution. New Caledonia, 62-80 m (dd): Vanuatu,

64-91 m (dd).

Diagnosis. À medium size Perrinia species, much

higher than wide, with a conical shape, an angulated

periphery, 5 granular spiral cords on penultimate

whorl and an additional thinner cord on last whorl, the

abapical cord much stronger, making keel, two

columellar teeth, the abapical one much stronger, an

almost flat base with 4 rather thin spiral cords and no

umbilicus.

Description. Shell of moderate size for the genus

(height up to 8.7 mm, width up to 5.6 mm), much

higher than wide, rather thick, conical; height 1.6x to

1.7x width, height 3.5x to 3.9x aperture height:

anomphalous.

Protoconch about 150 um, of 1.25 whorl, prominent

above the first teleoconch whorl, without clearly

defined terminal varix.

Teleoconch up to 7.9 convex whorls, bearing 5 spiral

cords on the penultimate whorl, the abapical cord

much stronger than the other cords, making keel, with

thick pointed beads: strong axial prosocline ribs

between cords: periphery angulated. Suture visible,

not canaliculated.

First whorl convex, sculptured by about 20 weakly

prosocline, smooth, thin threads, interspace from 1.5x

to 2x the width of threads. On second whorl, P3 and

P4 rather weak, with pointed beads formed at

intersection of cords and threads. On third whorl, PI

and P2 very weak, P3 and P4 thickening quickly,

similar in size, making two similar keels. On fourth

whorl, P4 slightly stronger than P3, both much

stronger than PI and P2 and still making keel: S]

appearing between end of 3% and middle of 4* whorl,

C. VILVENS

similar in size to PI and P2; axial threads thickening,

making ribs; distance between ribs at least 2x width of

ribs. On fifth whorl, PI thickening, stronger than SI

and P2; beads of P4 transforming into blunt prickles;

interspace between P3, P4 and axial ribs excavated.

On last whorls, beads of PI axially pointed; on last

whorl, S4 visible, weaker than all other cords;

periphery carinate, stellate by the prickles of P4.

Columella straight, vertical, with a rather strong to

very strong rounded basal tooth and an inner part

axially excavated.

Aperture subcircular, without angle; outer lip slightly

thickened inside, without denticles but with up to 8

Chilodontidae from the Pacific Ocean

inner folds corresponding to external spiral cords;

possibly one denticle at the base of inner lip making a

U-notch under the columellar tooth.

Base almost flat, with 4 rather thin, subgranular to

granular spiral cords, similar in size to S4; distance

between cords about 1.5x width of cords, distance

between S4 and outermost cord about 3x width of

cords; rather close, thin axial ribs between cords.

No umbilicus.

Colour of teleoconch off with possible brownish

flames on the whorls; spiral cords of the base with

regular brownish spots; protoconch white.

holotype

paratype |

paratype 2

paratype 3

Table 18. Perrinia cantharidoiïdes n. sp.:

Discussion. The especially high H/W ratio combined

with the columellar teeth and spiral sculpture of the

whorls make it difficult to confuse the new species

with the other Indo-Pacific Perrinia species, except

two species.

First, the new species is rather close to Perrinia

stellata (A.Adams, 1864) from western Indian Ocean

(figs 20V-X), but this slightly greater species (height

up to 15 mm) has a lesser H/W ratio, a much stronger

peripheral Kkeel, lacks SI, has a much weaker

columellar tooth and stronger spiral cords on the base.

Secondly, P. cantharidoides n. sp. resembles

Clypeostoma cancellatum (Schepman, 1908) from the

Philippines (figs 16A-X), but this slightly greater

species has especially a much more depressed shape

and 6 spiral cords on the base.

Etymology. Shaped (Ancient Greek: -mônç suffix) —

with reference to shape of the shell, that is rather

Figure 20 (scale bars: 5 mm).

Shells measurements in mm for types.

similar to one of Cantharidus (Trochidae:

Cantharidinae) species.

Perrinia guadalcanalensis n. sp.

Figs 21A-G, Tables 17, 19

Type material. Holotype (6.8 x 5.2 mm) MNHN (IM-

2000-32802). Paratypes: 2 MNHN (IM-2000-32803).

Type locality. Solomon Islands, off Guadalcanal

Island SALOMON 1, stn DW1744, 10°32'S,

159°39'E, 49-55 m.

Material examined. Solomon Islands. SALOMON

l: sin DW1744, 10°32'S, 159°39'E, 49-55 m, 3 dd

(holotype and 2 paratypes).

Distribution. Only known from the type locality.

A-I. Perrinia angulifera (A.Adams, 1853), New Caledonia. A-C. BATHUS 1, stn DW1233, 45-50 m, 11.5 x

8.6 mm. D-F. LAGON, stn DW271, 22 m, 16.2 x 11.5 mm. G-I. Juvenile, Koumac, MONTROUZIER, stn 1309,

18 m, 3.2 x 2.4 mm.

J-L. Perrinia elisa (Gould, 1849), Vietnam, 40 m, 19.5 x 15.0 mm, coll. C.Vilvens.

M-U. Perrinia cantharidoides n. sp. M-R. New Caledonia, Koumac, MONTROUZIER, stn 1315, 66-87 m. M-

O. Holotype MNHN (IM-2000-32800), 8.4 x 5.4 mm. P-R. Paratype MNHN (1M-2000-32801), 8.7 x 5.6 mm.

S-U. Coral Sea, Lansdowne-Fairway Banks, CORAIL 2, stn DW26, 62 m, 9.7 x 5.7 m.

V-X. Perrinia stellata (A.Adams, 1864), Red Sea, Dahab, Sinaï, 8.3 x 6.4 mm. coll. C.Vilvens.

C. VILVENS

Chilodontidae from the Pacific Ocean

Diagnosis. À rather small size, higher than wide,

Perrinia species with a more or less conical shape, an

angulated periphery, 5 granular spiral cords on

penultimate whorl and an additional thinner cord on

last whorl, the abapical cord much stronger and

making keel, a strong basal columellar tooth, an

almost flat base with 6-7 rather thin spiral cords and

no umbilicus.

Description. Shell of moderate size for the genus

(height up to 7 mm, width up to 5.2 mm), higher than

wide, rather thick, coeloconoiïidal on the adapical part

and more or less conical on the abapical part, height

1.3x to 1.4x width, height 3.8x to 3.9x aperture height;

anomphalous.

Protoconch missing on all available samples.

Teleoconch up to 7 almost flat whorls, with 5 spiral

cords on the penultimate whorl, the abapical cord

much stronger than the other cords, making keel, with

moderately thick pointed beads; strong slightly

prosocline ribs between cords; periphery angulated.

Suture visible, channelled, not canaliculated.

First whorl convex, with some weakly prosocline,

smooth, low, wide threads. Second whorl less convex,

with 4 Pi, with beads formed at intersection of cords

and threads; P3 and P4 stronger. Third whorl almost

holotype

)

6.8

paratype 2

flat, with S1 appearing; P3 and P4 thickening quickly:

distance between cords similar to size of cords. On

fourth whorl, P4 stronger than the other cords, making

a weak keel. On fifth and sixth whorl, PI thickening,

stronger than S1, P2 and P3, weaker than P4; beads of

PI axially pointed, beads of P4 horizontally pointed;

distance between P3 and P4 greater than cords,

interspace between P3, P4 and axial ribs excavated.

On last whorl, S4 visible, weaker than all other cords;

periphery carinate, moderately stellate by the pointed

beads of P4.

Columella straight, slightly prosocline,

moderately strong rounded basal tooth.

Aperture subcircular, without angle; outer lip slightly

thickened inside, with up to 9 inner folds (not clearly

visible on holotype).

Base almost flat, with 6 to 7 thin, subgranular to

nearly smooth spiral cords, much closer in the inner

part; distance between cords from 1.5x to 2x width of

cords, close, thin axial ribs between cords.

No umbilicus.

Colour of teleoconch off white with possible brownish

flames on the whorls; spiral cords of the base with

regular brownish spots.

with a

Table 19. Perrinia guadalcanalensis n. sp.: Shells measurements in mm for types.

Discussion. The new species is only close to P.

cantharidoides n. sp. from New Caledonia (figs 20M-

Ü), but this slightly greater species has a much higher

H/W ratio, P3 making two keels with P4 from the

third to the fifth whorl, two columellar teeth and only

4 wider and subgranular (not almost smooth) spiral

cords on the base.

Etymology. From Guadalcanal Island (Solomon

Islands), off which the samples were collected.

Perrinia squamocarinata (Schepman, 1908)

Figs 21H-P, Table 17

Calliostoma (Perrinia) squamocarinatum Schepman,

1908: 68-69, pl. VL fig. 3 [squamicarinatum]. Type

locality: Madura Bay, East Java, Indonesia, 69-91 m.

Perrinia squamicarinata [sic] — Poppe, Tagaro &

Dekker, 2006: 45-46, , figs 38-39, pL.IS, fig.S5.

Perrinia squamocarinata — Herbert, 2012: 474, figs.

71E-F.

Philippines. AURORA 2007: stn CP2653, 16°06'N,

121°59'E, 83 m, 1 dd. — Stn CP2654 “LG 04N,

121°57E, 98-107 m, 2 dd. — Stn CP2666, 15°58'N,

121°45'E, 199-307 m, 3 dd, 1 dd sub. — Stn DW2670,

14°52'N, 121°49'E, 180-187 m, 1 dd. — Stn CP2738,

I6°0O4'N, 121°56'E, 111-113 m, 9 dd. — Stn CP2739,

16°05'N, 121°58'E, 96 m, 21 dd, 5 dd sub, 4 dd juv. —

Figure 21 (scale bar: 5 mm).

A-G. Perrinia guadalcanalensis n. Sp., Solomon Islands, off Guadalcanal Island SALOMON 1, stn DW1744,

49-55 m. A-C. Holotype (1M-2000-32802), 6.8 x 5.2 mm. D-F. Paratype 1 (1M-2000-32803), 6.6 x 5.2 mm. G.

Paratype 2 (IM-2000-32803), 7.0 x 5.0 mm, view of the apertural inner folds.

H-P. Perrinia squamocarinata (Schepman, 1908), Philippines. H-M. AURORA 2007, stn CP2654, 16°04'N.

121°57'E, 98-107 m. H-J. 5.5 x4.4 mm. K-M. 7.4 x 5.5 mm. N-P. AURORA 2007, stn DW2670, 14°52'N,

121°49'E, 180-187 m, 9.2 x 5.9 mm.

C. VILVENS

Chilodontidae from the Pacific Ocean

Stn CP2747, 15°56'N, 121°42'E, 120-124 m, 5 dd. —

Sin CP2758, IS°S5SIN 121FS0 ES ITS mn 210

PANGLAO 2004: stn T4, 09°33'N, 123°49'E, 82 m, 3

dd, 2dd juv. — Stn T6, 09°35'N, 123°S l'E, 34-82 m, 4

dd, 3 dd sub. — Stn T9, 09°34'N, 123°50'E, 97-120 m,

15 dd, 15 dd juv. — Stn T14, 9°42'N, 123°49'E, 101-

110 m, 4 dd, 2 dd juv. — Stn T18, 09°42'N, 123°5S0'E,

80-100 m, 1 Iv. — Stn 127, 09°33/N; 123°5 VE; 106-

137 m, 1 dd, 1 dd sub, 2 dd juv. — Stn T44, 09°34'N,

123°49'E, 83-86 m, 3 dd. —- PANGLAO 2005: stn

DW2370, 08°34'N, 123°09'E, 92-96 m, 1 dd.

Solomon Islands. SALOMON 2: stn DW2294,

STIS IS TS 0E T0S-12$1m dd

Distribution. Philippines, 50-199 m, Iv at 80-100 m;

Indonesia, East Java, 90 m (using data of Schepman,

1908); Solomon Islands, 105-128 m (dd).

Remarks. The main features of this species are:

+ height usually about 9.2 mm, up to 6.5 mm, width

usually about 4.2 mm, up to 5.2 mm;

+ spire elevated, conical;

+ protoconch glossy, very weakly convex (almost

flat);

+ teleoconch of up to 6.5 almost flat to weakly

convex whorls; 3 granular spiral cords fully visible

and a fourth usually partially covered by suture on the

penultimate whorl; 4 cords on the last whorl;

+ first whorl with prosocline threads, distance

between threads about 2x width of threads; P2 and P3

at end of whorl, very thin; PI at second whorl; on

third whorl, threads thickening into ribs; spiral cords

stronger, similar in size, beads at intersection of cords

and ribs; on next whorls, P3 the strongest with pointed

beads, P2 the weakest; P4 partially emerging from

suture, visible on last whorl, peripheral, weaker than

P2; on last whorl, pointed beads of P3 sharp and scaly;

+ columella with a small basal tooth;

+ aperture subelliptical; outer lip thickened inside,

with about 8 inner folds;

+ base moderately convex, with 5 granular spiral

cords; distance between cords greater than size of

cords; axial thin threads between cords;

+ narrow umbilicus restricted to a slit or even closed

by columellar callus; :

+ nacreous white with pinkish flames on the whorls

and brownish spots on the cords.

The single record here reported from Solomon Islands,

with a single dead adult sample, extends the

distribution of this species to the Solomon Islands.

Regarding the Philippine samples, the only different

feature is the number of spiral cords on the base (4

instead of 5).

ACKNOWLEDGEMENTS

I would like to thank P. Bouchet (MNHN) for reading

the manuscript, constructive advice and access to the

malacological resources of the MNHN, and V. Héros

(MNHN) for checking carefully the manuscript, for

her help in finding various scientific papers and for

accurate management of registration numbers and

locality data. We also thank P. Maestrati (MNHN) for

his help to locate and sort the numerous samples

studied in this paper.

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Mayr, E. 1989. Attaching names to objects. In: What the philosophy of biology is: essays for David Hull (M. Ruse, ed.), Klumer Academic, Dordrecht: 235-243.

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