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SOCIETE BELGE DE MALACOLOGIE

JAY & DRIVAS

Cerithiopsidae Reunion

The Cerithiopsidae (Gastropoda) of Reunion Island (Indian Ocean)

Maurice JAY

46 rue Eugène Dayot,

97434 Saint-Gilles-les-Bains, la Réunion

Jean DRIVAS

Tax Skriperon,

49083 Skripero, Corfu, Greece.

KEY WORDS. Cerithiopsidae, Reunion Island, Indian Ocean.

MOTS CLES. Cerithiopsidae, Réunion, Océan Indien.

ABSTRACT. The authors have collected 83 species of Cerithiopsidae on Reunion Island, Indian

Ocean. After a revew of literature, and after having examined material in the Museum collections in

Berlin, Cardiff, London, Manchester and Paris, 16 species were recognized as known species, 23

were left aside on account of unsufficient material.

- 44 species are described as new.

- 2 new genera are proposed: Belonimorphis (type species Belonimorphis belonimorphis); Koïilofera

(type species Koilofera koilofera).

- New synonymies: Cerithiopsis aeolomitres Melvill & Standen, 1896 — Cerithiopsis (Horologica)

balteata Watson, 1886.

- New name: Cerithiopsis (Mendax) melvilli nom. nov. = Cerithiopsis aurantiaca Melvill & Standen,

1896, preoccupied.

RESUME. Les auteurs ont pu rasssembler des spécimens de Cerithiopsidae de la Réunion,

appartenant à 83 espèces. Après une revue de la littérature et comparaison avec le matériel des

Museums de Berlin, Cardiff, Londres, Manchester et Paris, 16 espèces ont été répertoriées comme

déjà connues, et 23 ont été laissées en dehors de l’étude pour cause de matériel encore insuffisant.

- 44 espèces sont décrites comme nouvelles

- 2 nouveaux genres sont proposés: Belonimorphis (espèce type Belonimorphis belonimorphis):

NOVAPEX 3 (1): 1-45, 10 mars 2002

Koilofera (espèce type Koïilofera koilofera).

Synonymes nouveaux:

(Horologica) balteata Watson, 1886.

Cerithiopsis aeolomitres Melvill & Standen, 1896 — Cerithiopsis

- Nouvelle dénomination: Cerithiopsis (Mendax) melvilli nom. nov. — Cerithiopsis aurantiaca

Melvill & Standen, 1896, preoccupé.

INTRODUCTION

The family Cerithiopsidae A. & H. Adams, 1854 (type

genus Cerithiopsis Forbes & Hanley, 1850) was

established for turreted shells, small or very small,

with a conical elevated and narrow spire, or fusiform

or pyriform, the teleoconch whorls sculptured with

spiral cords and often with axial ribs, their

intersections nodulose: aperture oval with a short

anterior canal; protoconch variable, either short or

more often elevated and very fragile, either smooth or

finely sculptured.

Since the publication of The Cerithiopsidae of New

Zealand by B.A. Marshall (1978), a growing

importance has been given to the features of

protoconch and radula among the classification

characters. A. Nutzel (1992) proposed to give a family

rank to the Eumetulinae of preceeding authors,

considering thus two families among the

Cerithiopsoidea, namely the Eumetulidae and the

Cerithiopsidae. But none of the species collected in

Reunion, except one, was ever found alive: hence,

radulae were not studied, and we could not follow the

taxonomic concepts based on their characters. So,

following Marshall, we will consider a single family,

the Cerithiopsidae sensu lato.

The Cerithiopsidae on Reunion

The earliest catalogues of molluses in Mascarene

Islands (Bernardin de Saint-Pierre in 1773,

d’Argenville in 1780, Sganzini in 1843) do not include

any Cerithiopsidae.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Nor did Deshayes (1863) cite any in his “Catalogue

des Mollusques de la Réunion”.

Lamy (1909) described Cerithiopsis blandi Vignal,

after a single specimen from Madagascar, and several

specimens from Reunion, Saint-Pierre, all of them

syntypes. Protoconchs of the species remained

unknown. 2 boxes labelled ‘“syntypes” are registered

in the MNHN typotheque: all specimens except 2

match the figure of Lamy, and in spite of the lack of

protoconchs, can be certainly identified as Horologica

turrigera (Watson, 1886), of which Cerithiopsis

blandi is thus a junior synonym. Furthermore, the

boxes labelled C. blandi contain 2 other different

specimens: one of them, deprived of its protoconch,

matches the teleoconch of our Mendax penneyi, n.sp.,

but could nevertheless be Cerithiopsis hedista Melvill

& Standen, 1896: the other specimen, very worn and

without protoconch, cannot be identified with

certainty, but resembles our new species Joculator

keratochroma.

Descriptions of the marine fauna in the neighbouring

island Mauritius are more numerous: although neither

Lienard Elize (1877) nor von Martens (1880) quote

any Cerithiopsidae. Several samples from Mauritius

are present in the Melvill-Tomlin collection, namely

Cerithiopsis adelpha, C. aurantiaca, C. catenaria, C.

eutrapela, €. fosterae, C. mathildaeformis. Viader in

his Revised Catalogue of species from Mauritius

(1937), quoted 6 species of Cerithiopsidae, namely :

Cerithiopsis catenaria Melvill & Standen, 1896, C.

Jfosterae Melvill & Standen, 1896, C. mathildaeformis

Melvill, 1907 ( Metaxa, Triphoridae), C. pulvis

(Issel, 1869), C. subreticulata (Dunker, 1861) and

Seila alfredensis Bartsch, 1915.

During those last 40 years, we have collected in

Reunion specimens belonging to 83 species of

Cerithiopsidae. They consist mostly in specimens

found dead in hand-dredged sand from depths

accessible by scuba diving, that is from 10 to 70 m.

Many specimens lack their protoconch, in which case

only those with a particular coloration could be

identified. Among these 83 species, 16 could be

identified as known species; 44 are new species and

are described; for 23 species, material was not

sufficient to allow identification or an original

description. The latter will not be studied in this paper.

Most of species seem to be rare, with sometimes only

a few specimens collected. The commonest species

Horologica turrigera (Watson, 1886) was the only

species to be found alive. In spite of our research, we

have not found Cerithiopsidae specimens on or in

sponges.

Assigning our species to genera.

The earlier descriptions have most often been based on

specimens lacking a protoconch, and generic

characters relate only to the teleoconch. Thus

comparison and identification with earlier type

material, figures, and descriptions, are difficult.

However, in more recent studies, a growing

importance has been given to protoconch characters,

primarily its smooth (for genera Cerithiopsis,

Joculator, Horologica) or sculptured appearance

(genera Mendax, Dizoniopsis, Prolixodens).

Examination of the protoconchs at high magnification

and SEM, may show other characters, such as

granulose or strongly mamillated patterns, small and

low reliefs of various forms, or small and short axial

riblets. When more material is available, these

characters may be seen to be of generic significance,

but in the present paper, we will consider them as

minor characters.

Among the species with obviously beaded spiral cords

on the teleoconch, species with a smooth protoconch

are easily referred to genera.

Cerithiopsis Forbes & Hanley, 1850 (Type species:

Cerithiopsis tubercularis (Montagu, 1803) from

Europe), includes species with a high teleoconch, with

numerous whorls, not constricted at base, and with a

protoconch made of 3 to 6 smooth whorls that is

without cords or ribs, sometimes granulose, especially

on the first whorl. 15 of our species may be attributed

to this genus sensu lato, among which 2 are known

species, and 10 are new ones. The lack of radulae on

our specimens does not allow us to refer them to allied

genera based on radular characters.

Joculator Hedley, 1909 (type species: Cerithiopsis

ridicula Watson,1886, from Queensland) includes

small or very small species, bulbous or ovate in shape,

more or less constricted at base, bearing 3 beaded

spiral cords from the earliest whorls of the teleoconch,

with an elevated smooth or punctate protoconch, but

without any cords or ribs. 32 of our species are

assigned to this genus, among which 5 are known

species and 16 are new ones.

Horologica Laseron, 1956 (Type species Horologica

bicolor Laseron, 1956, from Queensland) was

established for ovate species constricted at base like

Joculator, With a smooth or punctate protoconch, but

with only 2 beaded spiral cords on teleoconch whorls.

Authors have considered this genus as poorly

separated from Joculator, some species with 2 beaded

spiral cords on earlier whorls having 3 of them on last

whorl. However following Marshall, we will consider

species showing this character as Horologica. 16 of

our species are assigned to this genus, among which 7

are known species, and 4 are new ones.

Our species with beaded spiral cords on teleoconch,

and with a ribbed protoconch, are not so easily

assigned to genera. These ribbed protoconchs may be

classified into 4 different types : Some of them show

axial ribs extending from suture to suture, with smooth

or punctate intervals (Type 1) ; other ones have the

same axial ribs and intervals, with a narrow row of

small axial riblets in suture (Type 2) ; other ones show

axial ribs extending from suture to suture, and finer

spiral cords in their intervals (Type 3) and lastly, other

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

ones bear axial ribs limited to the lowest two thirds of

whorls, the upper third being smooth or punctate and

appearing more or less concave (Type 4).

Dizoniopsis Sacco, 1895 (type species Cerithium

bilineatum Hôrnes, 1855, type locality the Tertiary

formations of Piemont) was created for fossil shells

with only 2 beaded spiral cords on teleoconch whorls,

but the original description did not mention

protoconch, and the figure does not show it. Some

authors have assigned to this genus shells with smooth

protoconchs (Gougerot & Le Renard). But the genus

was redefined by Nordsieck, (1968), the type of the

genus having, according to him, a protoconch with a

smooth first whorl, and following whorls with axial

riblets extending from suture to suture. This genus

was used by Glibert (1973) for shells with pupoid

shape, with 2 beaded cords on teleoconch whorls, and

protoconchs with axial ribs. We will follow these

authors, and consider as Dizoniopsis sensu stricto,

species defined as above. We provisionally assign to

the genus sensu lato, shells with 2 beaded cords on the

teleoconch, and a ribbed protoconch of type 3 (with

spiral cords in intervals) and type 4 (axial ribs limited

to the lower 2/3 of whoris). A total of 3 of our new

species are assigned to the genus Mendax, Finlay,

1927 (Type species Cerithiopsis trizonalis Odhner,

1924, from New Zealand). Marshall (1978) restricted

the genus to species with a short paucispiral

protoconch, weakly delimited from teleoconch, with a

smooth first whorl, following whorls with axial ribs

extending from suture to suture, and a more or less

high teleoconch, more or less constricted at the base,

with 3 beaded cords per whorl. We assign to the genus

4 of our new species, though their protoconchs are

rather high. We provisionally assign to the genus one

more species with a ribbed protoconch of type 3 (spiral

cords in intervals between ribs).

Prolixodens Marshall, 1978 (type species Cerithiopsis

infracolor Laseron,1951, type locality Long Reef,

NSW, Australia) was established for narrow and

slender shells with straight sides, a ribbed protoconch

of type 4 (axial ribs limited to the lower 2/3 of whorls),

and 3 beaded spiral cords on teleoconch whorls. 2 of

our species are assigned to this genus, as new species.

In due course, species with protoconch of type 3, or

species with protoconch of type 4, may be thought to

deserve new genera, based on protoconch characters

only, and including species with 2 or 3 beaded cords

on the teleoconch whorls.

Two of our species with beaded spiral cords show very

unusual characters.

One of them shows 2 beaded spiral cords per

teleoconch whorl, and a protoconch with a flat summit

and 3 whorls bearing a strong rounded spiral pad at

their lower part, the median and upper parts of whorls

being concave: this protoconch sculpture may recall

protoconchs illustrated by Marshall (1973) in the

genus Seila, With a quite different teleoconch. It is

nearer to the protoconch of /nella spina Marshall,

1983, which is sinistral. For this species, we propose a

new genus, Xoilofera (from Greek, meaning bearing a

concavity); type species: Koilofera koilofera, n.sp.

The second species is a very high and slender shell,

with teleoconch sides slightly convex and a reticulated

more or less strongly beaded sculpture, and with a

strongly elevated protoconch of 3.5 whorls bearing 2

saillant equal and well separated spiral keels, with

smooth intervalss We could not find such

protococonchs in former descriptions of

Cerithiopsidae. For this shell, we propose the new

genus Belonimorphis (from Greek, meaning needle-

shaped): type species: Belonimorphis belonimorphis

n.Sp.

Lastly, 4 of our species (2 of which as new species)

are assigned to the genus Sei/a A. Adams, 1861 (Type

species: 7riphoris dextroversus Adams & Reeve,

1860, from China Seas). This genus was established

for species with high conical or slightly convex

teleoconch, with smooth and unbeaded spiral cords,

very close-set fine axial threads in the intervals

between spiral cords and a variable protoconch.

Abbreviatons used

MM : Manchester Museum.

MNEN : Museum national d'Histoire naturelle, Paris.

MNK: Museum für Naturkunde, Berlin.

NHM: Natural History Museum, London.

NMVW: National Museum of Wales, Cardiff.

SEM: Scanning Electron Microscope.

SYSTEMATICS

Genus Cerithiopsis Forbes & Hanley, 1850

Type species: Cerithiopsis tubercularis (Montagu,

1803), Europe: elongate shell, not constricted at base,

with 3 beaded spiral cords per teleoconch whorl,

protoconch smooth or punctate, without any axial rib

or spiral cord, except sometimes for a spiral carina on

last half-whorl.

Cerithiopsis eutrapela Melvill & Standen, 1896.

Plate 1, A; colour plate I, Fig. 1)

Material examined. ! spmn MNHN; 30 spmns with

complete protoconch, coll. M.Jay; 11 spmns coll.

J.Drivas.

Description. Shell conical, elevated, and slightly

oval, somewhat wider than the related species; angle at

the summit of teleoconch 35°. Protoconch strong and

high of 3,5 convex smooth whorls, finely punctate

under SEM, with very fine close-set axial threads in

suture; its limit from teleoconch clear-cut and oblique,

its last 1/4 whorl with a median spiral carina.

Teleoconch of 8 or 9 convex whorls, bearing 3 beaded

spiral cords, the upper one slightly weaker. One weak

spiral thread between the cords, on the 2 last whorls.

JAY & DRIVAS

Axial ribs, weaker than cords, crossing them at right

angles, with one rounded bead at each intersection,

beads numbering 23-24 per whorl. A fourth weaker

and more finely beaded spiral cord emerging from

suture at base of last whorl. Base smooth. Aperture

circular. Colour creamy-white, the earlier whorls more

neatly white, protoconch brown.

Size: maximum total height 6.5 mm: width at base 2

mm; height of protoconch 0.42 mm; width of

protoconch at base 0.28 mm.

Locality. Found dead in hand-dredged sand at 10-20

m, off Saint-Gilles-les-Bains.

Remarks. Our specimens were compared and found

identical to the holotype of Melvill & Standen, in

MM: a single specimen without protoconch; and to

the syntypes in NMW (lot Nr Z.1955.158.02262) from

Lifu, Loyalty Isl. They resemble also another lot of

the Melvill-Tomlin collection, in NMW (lot Nr

Z.1955.158.02268) from Mauritius.

Cerithiopsis fosterae Melvill & Standen, 1896.

Plate 1, B; colour plate I, Fig. 2

Material examined. 1 spmn MNHN; 13 spmns with

complete protoconch coll. M. Jay; 40 spmns with

broken protoconch coll. M. Jay; 11 spmns coll. J.

Drivas.

Description. Shell elevated and conical, very elongate

and slender, summit angle 22°. Protoconch prolonging

the general shape of teleoconch, comprising 5 convex

smooth whorls, with under SEM some very fine axial

threads above suture on first whorl, and a weak spiral

cord in suture between the 4th and Sth whorls; limit

from teleoconch oblique and well marked, the last 1/4

whorl bearing a median spiral carina, more or less

visible. Teleoconch of 10 strongly convex whorls.

Suture deeply impressed. 3 beaded spiral cords per

whorl, the uppermost one somewhat weaker and

slightly constricted, but visible from first whorl. Axial

ribs weaker than cords, crossing them and joining the

beads, obliquely in the first interval, axially in second

one, and extending from upper to lower suture, but

discontinuous from one whorl to another. Axial ribs

and beads numbering 26 to 28 per whorl. A fourth

spiral cord, weaker and smooth, emerging from suture

at base of last whorl. One more weak and smooth

spiral cord on base. Aperture rounded. Colour white,

suture and upper beaded cord very pale brown in fresh

specimens, but this colour fades rather fast.

Protoconch white.

Size: maximum total height 6.1 mm; width at base 1.4

mm; height of protoconch 0.56 mm; maximum width

of protoconch 0.30 mm.

Locality. Found dead in hand-dredged sand at 30-55

m, off Saint-Gilles-les-Bains.

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Remarks. Our specimens appear identical to the

holotype of Cerithiopsis fosterae Melvill & Standen,

1896, type locality Loyalty Islands, in MM: a single

specimen that matches the figure of Melvill &

Standen, with a protoconch broken at summit and 3

whorls left. They are identical also to the syntypes of

Melvill & Standen in NMW (lot Nr Z.1955.158.00205 )

five specimens from Lifu: only one specimen has a

protoconch of 3 whorls, with broken summit.

Furthermore our specimens are identical to specimens

in two other lots of the Melvill-Tomlin collection in

NMW, from Mauritius, of which none has a complete

protoconch. The original description of Melvill &

Standen described a protoconch of 3 whorls instead of

5, without mentioning the broken summit, and

described 2 beaded spiral cords on teleoconch whorls;

the third and upper cord, weaker and constricted, not

mentioned in the original description, is nevertheless

visible on Melvill & Standen’s figure, and on the types

and syntypes.

Cerithiopsis boucheti n.sp.

Plate 1, C; colour plate I, Fig. 3

Material examined. 2 spmns MNHN; 37 spmns coll.

M. Jay, all with complete protoconch; 6 spmns coll.

J. Drivas.

Description. Shell conical, elevated, very slightly

oval. Protoconch nearly cylindrical, of 3 strongly

convex whorls, the earlier one rounded, the following

two equal, their diameter smaller than the first whorl

of teleoconch; whorls looking smooth, but finely

punctate under microscope; mamillated with close-set

hemispherical tubercles under SEM; last whorl well

separated from teleoconch by an oblique line with

change of colours, but the 3 spiral cords of teleoconch

have appeared and are weakly visible on the last 1/4

whorl of protoconch. Teleoconch of 7 slightly convex

whorls, with 3 beaded spiral cords per whorl, equal in

importance on last whorl, the upper cord a little

weaker and retracted on earlier whorls. Very fine axial

riblets crossing the cords at right angles, with a

rounded bead at each intersection, beads numbering

22-23 per whorl. Very close-set fine axial striae visible

in the intervals between cords, under strong

enlargement. A fourth finer and more weakly beaded

spiral cord emerging from suture at base of last whorl:

beads becoming weaker, and axial riblets more neatly

visible on last part of last whorl. One more weak

smooth cord on base. Aperture circular. Colour plain

glossy brown, protoconch white.

Size: holotype total height 4.5 mm, width at base 1.3

mm; height of protoconch 0.49 mm; width of

protoconch at base 0.36 mm.

Type locality. Saint-Gilles-les-Bains, in hand

dredged sand at 10-20 m, between Hermitage and

Saint-Paul Bay.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Type material. Holotype and paratype 1 in MNAN,

paratypes 2 to 15 coll. M. Jay: paratypes 16 to 18 coll.

J. Drivas.

Etymology. Dedicated to Philippe Bouchet, curator of

the Mollusca section in MNHN.

Remarks. Eight of our species look alike by their high

teleoconch, with 3 spiral cords, the upper one weaker

and retracted to penultimate whorl, and by their colour

more or less golden brown. They are undistinguishable

on teleoconch characters only, in spite of slight

differences in their width, and differ mostly by their

protoconch: 5 of them are Cerithiopsis, with smooth or

punctate protoconchs. C. hadfieldi n.sp. has a conical

protoconch of 5 whorls, and the widest teleoconch. C.

boucheti n.sp. has a more cylindrical protoconch of 3

whorls; C.pickeringae n.sp. has a conical protoconch

of 4 whorls with rounded apex. C. nutzeli n.sp. is a

shorter shell with a protoconch of only 2 whorls.

Cerithiopsis seddonae n.sp. differs from other ones by

its wider protoconch of 3.5 whorls, by its shorter

teleoconch, and by its upper cord more retracted.

Three other species may be confused with the five

preceeding ones but have protoconchs with axial ribs:

well visible for Mendax metivieri n.sp. and Mendax

ribesae n.sp.: for Mendax mascarenensis n.sp. axial

riblets on protoconch are low and have to be searched

under oblique light.

Moreover, these species resemble the species labelled

Cerithiopsis brunnea Thiele from SW Australia

(MNK lot Nr 67484), which has only the last smooth

whorl of its broken protoconch left, but the teleoconch

of which bears larger and less numerous beads (18 per

whorl instead of 22). C. exquisita Sowerby, 1897

(Natal, South Africa) differs from our species in that

its teleoconch whorls are more convex, and its upper

cord more retracted. The Joculator species with the

same colour are much smaller and constricted at base.

Cerithiopsis hadfieldi n.sp.

Plate 1, D: colour plate I, Fig. 4

Material examined. 2 spmns MNHN, 17 spmns coll.

M.Jay, (13 with complete protoconch).

Description. Shell conical, high, slightly fusiform,

summit angle of teleoconch 30°. Protoconch

prominent and obviously conical, of 5 smooth convex

whorls, appearing punctate under microscope: apex

rounded, suture wide bearing fine riblets, axial or

slightly oblique, stronger and more widely-spaced than

on other protoconchs of the same type and numbering

about 20 per whorl: suture slightly darker than

protoconch whorls: limit from teleoconch clear-cut

and oblique. Teleoconch of 8 convex whorls, bearing

3 beaded spiral cords per whorl, the uppermost one a

little weaker and constricted but visible from the first

whorl. Axial ribs, weaker than cords, crossing them at

right angles, with a bead at each intersection: beads

more or less strong and rounded numbering 23-24 per

whorl. Intervals between cords very finely axially

striated. A fourth smooth cord emerging from suture at

base of last whorl. Base excavated, smooth: last whorl

and aperture expanded on some specimens. Aperture

circular. Colour more or less dark brown, protoconch

white.

Size: maximum total height 5.6 mm; maximum width

1.4 mm: height of protoconch 0.82 mm: width of

protoconch at base 0.32 mm.

Type locality. Found dead in hand-dredged sand at 45

m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNEN,

paratypes 2 to 7 coll. M. Jay.

Etymology. Dedicated to the Rev. James Hadfield,

who collected the first specimens of that species in

Lifu.

Remarks. Our specimens were compared and found

similar to the syntypes of Cerithiopsis catenaria

Melvill & Standen, 1896, in NMW, (lot Nr

Z.1955.158.00193): 3 specimens from Lifu, one of

them with protoconch. They are identical also to the

specimens of another lot in the Melvill-Tomlin

collection in NMW (lot Z.1955.158.02263) from

Mauritius. However, the holotype of the species in

MM collection (Lot EE 3714) differs from the NMW

syntypes by its protoconch, only 1/4 of last whorl of

which remains, bearing 4 strong and rounded axial

ribs; and also in its teleoconch possessing elevated and

narrow spiral cords, darker than ground, and beads a

little more numerous: this specimen, using our criteria,

should be assigned to the genus Mendax. The

specimens labelled syntypes in NMW thus belong to

another and undescribed species, together with our

specimens from Reunion; for it, we propose the name

Cerithiopsis hadfieldi.

Cerithiopsis iochrous n.sp.

Plate 1, E; colour plate I, Fig. 5

Material examined. 2 spmns MNHN, 10 spmns

coll. M. Jay, (5 spmns with complete protoconch); 2

spmns coll. J. Drivas.

Description. Shell very high and slender with nearly

straight sides, summit angle 20°-22°. Protoconch

prolonging the general outline of teleoconch,

comprising 5 convex whorls, smooth under SEM,

without axial threads, but with a weak spiral cord in

suture between the 4th and Sth whorls: its limit from

teleoconch clear-cut and oblique, the last 1/4 whorl

bearing a spiral carina beginning the sculpture of

teleoconch. Teleoconch of 11 to 14 whorls, bearing 3

beaded spiral cords per whorl, high and narrow, the

uppermost one weaker and constricted. Axial ribs a

little weaker than the cords, numbering 21-22 on last

whorl; ribs slightly oblique in the first interval between

JAY & DRIVAS

cords, and axial in second one, and extending from

upper to lower sutures, but not from one whorl to

another. À small bead at each intersection. One fine

spiral thread between the cords on last whorl. A fourth

spiral cord emerging from suture at base of last whorl.

One additional weak smooth spiral cord on base.

Aperture cireular. Colour uniform pale violet, a little

darker on earlier whorls, protoconch white. This

colour fades more or less swiftly.

Size: holotype height 5.5 mm, width at base 1.2 mm;

height of protoconch 0.56 mm, maximum width of

protoconch 0.33 mm.

Type Locality. Saint-Gilles-les-Bains, found dead in

hand dredged sand at 10-20 m.

Type material. Holotype and paratype 1! in MNHN,

paratypes 2 to 6 coll. M. Jay, paratypes 7 and 8 coll.

J. Drivas.

Etymology. Named on account of its colour, from

Greek meaning violet.

Remarks. This species is very near to Cerithiopsis

Josterae Melvill & Standen, 1896, and differs from it

only in its colour and lack of fine axial threads on the

protoconch. It could perhaps be a colour variation of

this last species. One juvenile specimen among the

syntypes of C. fosterae in NMW has this colour and

belongs to this species.

Cerithiopsis jousseaumei n.sp.

Plate 1, F; colour plate I, Fig. 6

Material examined. 2 spmns MNHN, 86 spmns coll.

M.Jay, ( 21 with complete protoconch) 10 spmns

coll. J. Drivas.

Decription. Conical, high and slender shell, with

nearly straight sides, summit angle of teleoconch 20°.

Protoconch of 4.5 convex whorls, with a weak and

narrow spiral thread in suture, its last whorl with

progressive development of adult sculpture, but limit

from teleoconch marked by clear-cut oblique change

of colour. Teleoconch of 10 to 12 whorls bearing 3

subequal beaded spiral cords per whorl; axial ribs

weaker than cords, crossing them at right angles, with

a rounded bead at each intersection; beads appearing

close-set, and numbering 25 per whorl. The beads of

the uppermost spiral cord slightly stronger than others.

A fourth weaker and finely beaded cord emerging

from suture at base of last whorl. Another smooth very

weak cord on base. Aperture quadrangular with

rounded angles. Colour golden brown, glossy on fresh

specimens, becoming paler with age; protoconch paler,

base a little darker.

Size: holotype total height 6 mm, width at base 1.5

mm; height of protoconch 0.35 mm; width of

protoconch at base 0.25 mm.

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Type locality. Saint-Gilles-les-Bains, found dead in

hand-dredged sand at 10-30 m, off Boucan-Canot

beach.

Type material. Holotype and paratype 1 in MNHN,

paratypes 2 to 10 coll. M. Jay; paratypes 11 to 14

coll. J. Drivas.

Etymology. Dedicated to F. Jousseaume, who

described new species of Cerithiopsidae from Red sea

and Mascarenes Isl..

Remarks. This species differs from other Reunion

species in its 3 rows of subequal beads, its more

numerous and close-set beads, and its poorly marked

suture. Cerithiopsis orientalis Preston, 1905 (type

locality Ceylon) has a wider and deeper suture, and is

paler in colour. Cerithiopsis infracolor Laseron, 1951

(type locality Long Reef, NSW, Australia) has a

similar shape, size and colour, but is easily

distinguished from it by its costate protoconch.

Cerithiopsis lamyi n. sp.

Plate 2, A; colour plate I, Fig. 7

Material examined. 2 spmns MNHN, 6 spmns coll.

M. Jay (5 with complete protoconch).

Description. Shell conical, high, very narrow and

slender, base not constricted, summit with a wide

protoconch more shortly conical than the general

outline of the shell. Large conical protoconch of 5

smooth whorls, more convex at their lower part,

resembling a pagoda’s roof, with a wide suture; last

whorl equal in diameter to first whorl of teleoconch,

the limit between both marked by the sudden

appearance of the 3 adult cords and change of colour,

along a vertical line; earlier whorls tappering to

narrow rounded apex. Teleoconch of 8 whorls,

suture wide and deeply impressed; 3 spiral cords per

whorl, subequal, crossed at right angles by well

developed axial riblets, with one rounded bead at

each intersection; beads rather small, numbering 17

per whorl. A fourth spiral cord, weaker and finely

beaded, emerging from suture at base of last whorl. A

fifth smooth cord on base. Aperture circular. Colour

horn or dark cream, protoconch a little paler.

Size: holotype total height 1.5 mm, width at base 0.5

mm; height of protoconch 0.40 mm; width of

protoconch at base 0.32 mm.

Type Locality. Found dead in hand-dredged sand at

30m, off Souris-Chaude, Trois-Bassins.

Type material. Holotype and paratype 1 in MNHN;:

paratype 2 to 5 Coll. M. Jay.

Etymology. Dedicated to Edouard Lamy, French

conchologist.

JAY & DRIVAS Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

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PLATE 1. Fig. A. Cerithiopsis eutrapela Melvill & Standen, 1896; Off Saint Gilles les Bains,10-20 m:; height

6.5; MNAN. Fig. B. C. fosterae Melvill & Standen, 1896 ; Off Saint Gilles les Bains, 30-55m; height 6.1 mm:

MNAHN. Fig. C. C. boucheti n.sp. Off Saint Gilles les Bains, 10-20 m; holotype, height 4.5 mm; MNAHN. Fig .D.

C. hadfieldi n.sp. Off Saint Gilles les Bains, 45 m; holotype, height 5.6mm; MNHN. Fig. E. C. iochrous n.sp.

Off Saint Gilles les Bains, 10-20 m; holotype, height 5.5 mm; MNHN. Fig. F. C. jousseaumei n.sp. Off Saint

Gilles les Bains, 10-30 m; holotype, 2,6 mm; MNHN.

Scale bars: S (shells): 1 mm; P (protoconchs): 100 um.

JAY & DRIVAS

Remarks. The protoconch of this species resembles in

its size and its shape, /oculator granata Kay, 1979 and

Cerithiopsis vaurisi n.sp., but it does not bear the small

sutural axial riblets visible on those 2 species.

Furthermore, our new species differs from Joculator

granata by its more slender shape not constricted at

base and differs from Cerithiopsis vaurisi which has

the same shape, by its obviously paler colour.

Cerithiopsis nutzeli n.sp.

Plate 2, B: colour plate I, Fig. 8

Material examined. 2 spmns MNHN, 24 spmns coll.

M. Jay, all with complete protoconch.

Description. Shell high, fusiform, topped by the wide

and cylindrical protoconch. Protoconch of 2.5 convex

whorls, with a large rounded apex, appearing smooth

but finely punctate under microscope, and under SEM,

mamillated with small hemispherical close-set

tubercles; limit from teleoconch along an oblique line

marked by change of colour, but adult sculpture begins

1/4 whorl earlier by the progressive development of 2

beaded spiral carinas, continuing the 2 lower cords of

teleoconch. Teleoconch of 6 whorls, 3 beaded spiral

cords per whorl, subequal on last whorl, but upper

cord weaker and retracted on earlier whorls. Weaker

axial ribs crossing the cords at right angles, with a

rather small bead at each intersection; beads

numbering 24 on last whorl. Intervals occupied by

very fine axial striae visible only under microscope. A

fourth spiral cord emerging from suture at base of last

whorl, as strong as other ones, but unbeaded. Base

smooth. Aperture rounded. Colour plain orange-

brown, protoconch white.

Size : holotype total height 3.4 mm, maximum width 1

mm; height of protoconch 0.46 mm; width of

protoconch at base 0.29 mm.

Type locality. Saint-Gilles-les-Bains, between

Hermitage and Saint-Paul Bay, found dead in hand-

dredged sand at 15-30 m.

Type material. Holotype and paratype 1 in MNHN,

paratypes 2 to 12 coll. M. Jay.

Etymology. Dedicated to Alexander Nutzel (Berlin)

for his work on the family.

Remarks. This species differs from other allied

Cerithiopsis of Reunion, in its smaller size and its 2.5

whorled protoconch. It differs from Joculator species

of the same colour, in its larger size and unconstricted

base.

Cerithiopsis pickeringae n. sp.

Plate 2, C; colour plate I, Fig. 9

Material examined. 2 spmns MNHN,

coll. M. Jay, all with complete protoconch.

28 spmns

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Description. Shell conical, high, slightly fusiform,

protoconch extending the general outline of

teleoconch. Protoconch of 4 convex whorls, regularly

tapering towards the rounded apex, whorls smooth

with fine granulations appearing under microscope and

SEM, and very fine axial riblets in suture, hardly

visible under SEM: limit from teleoconch oblique and

clear cut, with change of colour and beginning of

teleoconch sculpture. Teleoconch of 8 whorls, 3

beaded spiral cords per whorl, equal on last whorl, but

the upper one weaker and retracted on earlier whorls.

Weaker axial ribs in their intervals, crossing them at

right angles, with a rounded bead at each intersection,

beads numbering 26-27 on last whorl. A fourth spiral

cord emerging from suture at base of last whorl,

weaker than other ones, retracted and unbeaded. Beads

becoming smaller at the end of last whorl. Base

excavated, smooth. Aperture rounded. Colour dark

golden brown, protoconch white.

Size: holotype total height 3.9 mm, maximum width

1.1 mm: height of protoconch 0.48 mm; width of

protoconch at base 0.27 mm.

Type locality. Saint-Gilles-les-Bains, found dead in

hand-dredged sand at 20-30 m.

Type material. Holotype and paratype 1 in MNHN,

paratypes 2 to 10 coll. M. Jay.

Etymology. Dedicated to Mrs Joan Pickering, curator

of Mollusca section in the BM.

Remarks. This species resembles Cerithiopsis

boucheti n.sp. but differs from it in its smaller size,

and its more conical protoconch of 4 whorls instead of

3. Other characters have been discussed under C.

boucheti.

Cerithiopsis seddonae n.sp.

Plate 2, F; colour I, Fig. 10

Material examined. 2 spmns MNHN, 20 spmns

coll. M. Jay, all with complete protoconch.

Description. Shell fusiform, rather elongate, base not

constricted. Protoconch prominent, rather wide, with

rounded apex, consisting in 3.5 convex whorls,

appearing smooth to the naked eye, finely granulose

under optical microscope, and mamillated with

rounded close-set tubercles under SEM; lower limit

oblique and clear-cut, marked by change of colour,

while on last whorl 2 cords progressively develop,

beginning the sculpture of teleoconch. Teleoconch of 5

slightly convex whorls, with 3 spiral cords per whorl,

crossed by finer axial ribs, with a rounded bead at each

intersection; beads numbering 20 to 22 on penultimate

whorl;: beads of the upper cord a little weaker than

others and constricted, including last whorl. Beads

always small, widely spaced, the reticulated sculpture

being more obvious than for other species. A fourth

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

spiral cord emerging from suture at base of last whorl,

smooth, and underlining angle with base. A further

smooth cord at mid-height on base. Aperture cireular.

Colour pale brown, protoconch paler or whitish.

Size: holotype total height 2.2 mm; width at base 0.8

mm; height of protoconch 0.60 mm; width of

protoconch at base 0.29 mm.

Type locality. Found dead in hand-dredged sand at

10-20 m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN;

paratypes 2 to 6 coll. M. Jay.

Etymology. Dedicated to Dr. Mary Seddon, curator of

Mollusca section in NMW.

Remarks. This species differs from Cerithiopsis

boucheti n.sp. in its smaller size (2.2 mm instead of 4.5

mm), and its wider protoconch (3.5 whorls instead of

3). It differs from Cerithiopsis pickeringae n. sp. in its

smaller size (2.2 mm instead of 3.9 mm) and wider

protoconch. From Cerifthiopis nutzeli n. sp. it differs in

its higher protoconch (3.5 whorls instead of 2.5); and

from Joculator species of the same colour, it differs in

not having a constricted base.

Cerithiopsis vaurisi n.sp.

Plate 2, E; colour plate 1, Fig. 11

Material examined. 2 spmns MNHN,

coll. M. Jay, 37 with complete protoconch.

38 spmns

Description. Shell conical, slender and high with

straight sides, protoconch more shortly conical than

general outline of teleoconch, giving its apex a

characteristic appearance. Protoconch strongly conical

of 5 whorls, last whorl as wide as first whorl of

teleoconch, their limit nearly axial marked by

development of adult sculpture; earlier whorls tapering

swiftly and regularly to the rounded and narrow apex;

whorls with flat sides, smooth, with a very narrow

spiral row of fine axial riblets in suture and

immediately under it, visible under optical microscope

and SEM; these riblets somewhat stronger than on

other species with the same type of protoconch, and

numbering about 24 on last whorl. Teleoconch up to 8

whorls with flat sides, wide and deeply impressed

suture; 3 spiral cords per whorl, crossed at right angles

by slightly weaker axial ribs, with a rounded bead at

each intersection, more or less large; beads more

widely spaced axially than spirally, numbering 25-26

on last whorl. A weaker unbeaded spiral cord

emerging from suture at base of last whorl. Base

excavated. Colour dark brown, protoconch paler at

apex but becoming darker on later whorls.

Size: holotype total height 3.6 mm; width at base 0.8

mm, height of protoconch 0.50 mm; width at the base

of protoconch 0.34 mm.

Type locality. Found dead in hand-dredged sand at

50-55m, Possession-Bay.

Type material. Holotype and paratype 1 in MNAN,

paratypes 2 to 8 coll. M. Jay.

Etymology. Dedicated to Daniel Vauris, skin diver

who collected sand for this study.

Remarks. This species has a protoconch of the same

type as Joculator granata Kay, 1979, but differs from

it in its base unconstricted, and in its more numerous

whorls. It is quite near Cerithiopsis lamyi n.sp. in

outline and shape of the protoconch, but differs from it

in the small axial riblets in the protoconch suture

(lacking in the C. /amyi), and in its darker colour. One

other Joculator species has a protoconch with ribbed

suture (Joculator myia n.sp.), but this is easily

separated by its constricted base, size and colour.

Cerithiopsis wayae n. sp.

Plate 2, D; colour plate I, Fig. 12

Material examined. 1! spmn MNHN, 2 spmns coll.

M. Jay, all with complete protoconch.

Description. Shell with a high spire, slightly fusiform,

base not constricted. Protoconch prolonging the

outline of teleoconch, made of 3 rather wide convex

whorls, smooth under the naked eye, but granulose

under optical enlargement, and mamillated by rounded

tubercles under SEM; apex widely rounded: lower

limit clear-cut oblique marked by change of colour,

but 2 spiral cords progressively develop on last 1/4

whorl, beginning adult sculpture. Teleoconch of 7

slightly convex whorls, suture weakly impressed, 3

spiral cords per whorl, rather strong, regularly swelling

into small and spirally elongate beads, numbering 26-

27 per whorl. Fine axial riblets joining the beads,

hardly visible even under microscope. The whole

surface covered with very fine lamellose axial threads.

A fourth unbeaded spiral cord, emerging from suture

at base of last whorl. Base excavated, a weak smooth

cord at mid-height. Colour blackish brown, protoconch

white.

Size: holotype height 4.1 mm; maximum width 1 mm;

height of protoconh 0.51 mm; width of protoconch at

base 0.38 mm.

Type locality. Off Souris-Chaude, Trois-Bassins,

found dead in hand-dredged sand at 30 m.

Type material. Holotype in MNHN, paratypes 1 and

2 coll. M. Jay.

Etymology.Dedicated to Ms Kathie Way, BM,

London.

Remarks. This species differs from other Cerithiopsis

species in its spirally elongate beads, and well

Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

PLATE 2. Fig. A. Cerithiopsis lamyi n.sp. Off Souris-Chaude, Trois-Bassins, 30 m; holotype, height 1.5 mm;

MNAN. Fig. B. C. nutzeli n.sp. Off Saint Gilles les Bains, 15-30 m; holotype, height 3.4 mm; coll. M.Jay.

Fig. C.C. pickeringae n.sp. Off Saint Gilles les Bains, 20-30 m; holotype, height 3.9 mm; MNHN.

Fig. D. C. wayae n.sp. Off Souris-Chaude, Trois-Bassins, 30 m; holotype, height 4.1 mm; MNHN.

Fig. E. C. vaurisi n.sp. Possession Bay, 55m; holotype, height 3.6 mm; MNAHN. Fig. F. Cerithopsis seddonae

n.sp. Off Saint Gilles les Bains, 10-20 m; holotype, height 2.2 mm; MNAN.

Scale bars: S (shells): 1 mm; P (protoconchs): 100 um.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

developed lamellose axial threads. The sculpture

resembles that of Belonimorphis belonimorphis n.sp.,

but the present species differs in its protoconch and its

earlier whorls.

Genus Joculator Hedley, 1909

Type species Cerithiopsis ridicula Watson.1886.

Queensland: Shell constricted at base; teleoconch

whorls with 3 beaded cords; protoconch smooth or

punctate, without ribs or cords.

Joculator albocinctum (Melvill & Standen, 1896)

Plate 4, A: colour plate I, Fig. 13

Material examined. 1 spmn MNHN: 30 spmns (13

with complete protoconch) coll. M. Jay; 11 spmns

coll. J. Drivas.

Description. Shell fusiform, constricted at base,

topped by protoconch. Angle at the summit of

teleoconch 40°. Protoconch nearly cylindrical, made of

4 convex whorls, smooth to the naked eye, under

SEM with fine granulations on first whorl and fine

axial riblets in suture, numbering about 20 on last

whorl; apex rounded, limit from teleoconch oblique,

marked by change of colour, adult sculpture

developing progressively on last whorl. Teleoconch

with 3 beaded spiral cords per whorl, and weaker axial

ribs, with a bead at each intersection, 19 to 20 beads

per whorl; beads of upper cord a little stronger and

more rounded than on lower cords. A fourth spiral

cord, undulose rather than beaded, emerging from

suture at base of last whorl. A fifth smooth cord on

base. Aperture oval. The upper beaded cord creamy-

white, the remainder of whorls pale brown, the 2

lowest cords appearing as brown lines, darker than the

beads. Protoconch white.

Size: maximum height 3 mm; maximum width 1 mm;

height of protoconch 0.57 mm; width of protoconch at

base 0.33 mm.

Locality. Found dead in hand-dredged sand at 10-20

m, off Saint-Gilles-les-Bains.

Remarks. The teleoconch of our specimens have the

same sculpture and same colour pattern as the

holotype of Bittium albocinctum Melvill & Standen,

1896, (type locality Loyalty Isl.) (MM lot EE 3702);

but this type has no trace of the protoconch left, and

is larger (4.3 mm without protoconch instead of 2.4

mm for the teleoconch of our larger specimen). On

account of the characteristic pattern of the

teleoconch, and in spite of the difference in sizes, we

propose this identification, specimens from Reunion

being considered as a dwarf variety.

Joculator granata Kay, 1979.

Plate 4, D: colour plate I, Fig. 14

Material examined. 1! spmn MNHN ; 15 spmns (9

with complete protoconch) coll. M. Jay:

Description. Shell fusiform, nearly cylindrical in the

middle, and slightly constricted at base. Protoconch

conical of 5 slightly convex whorls, smooth except for

a very narroW spiral row of very fine, short axial

riblets inside suture, projecting over the very

uppermost part of whorls, visible under optical

microscope and SEM, 22 riblets on last whorl; last

whorl as wide as the earlier whorl of teleoconch, the

limit between them obliquely marked by the

development of adult sculpture; earlier whorls tapering

swiftly to the narrow rounded apex. Teleoconch of 6

whorls, with 3 spiral cords per whorl, crossed at right

angles by weaker axial ribs, with a rounded bead at

each intersection: beads numbering 24 or 25 on last

whorl: the upper beaded cord a little weaker than the

other ones. A fourth beaded spiral cord emerging from

suture at base of last whorl. Aperture circular. Rare

specimens with one more whorl, wider and distorted,

but always constricted at base. Plain pale brown in

colour, protoconch paler.

Size: total height ranging from 2.3 mm to 2.9 mm;

maximum width from 0.7 to 0.9 mm; height of

protoconch 0.41 mm: width of protoconch at last

whorl 0.31 mm.

Locality. Found dead in hand-dredged sand at 10-20

m off Saint-Gilles-les-Bains.

Remarks. Our specimens match the description and

figure given by A. Kay, 1979 (Joculator granata, type

locality Hawaï) specifically the suture of the

protoconch, which A.Kay described as “erimped”, this

character given as distinctive from the related species.

3 species of Joculator among our material have such

protoconchs: Joculator myia n.sp. differs from J.

granata in its smaller size and in its 4.5 teleoconch

whorls instead of 6; Joculator skolix n.sp. differs from

it in its protoconch shape, size, and colour; Joculator

albocinctum (Melvill & Standen. 1896) differs in the

particular colour pattern of the teleoconch.

Cerithiopsis species with such protoconchs (

Cerithiopsis vaurisi n.sp., Cerithiopsis pickeringae

n.sp. Cerithiopsis hadfieldi n.sp.) differ from it in

having a higher teleoconch, not constricted at the base.

Joculator minima Laseron, 1955.

Plate 3, B; colour plate I, Fig. 15

Material examined. 1 spmn MNHN: 12 spmns (6

with complete protoconch) coll. M. Jay; 3 spmns

coll. J. Drivas.

JAY & DRIVAS

Description. Shell pupiform with constricted base and

summit topped by a high conical pointed protoconch.

Protoconch of 5.5 slightly convex smooth whorls,

under SEM with a very fine spiral thread just above

suture; last whorl rather large, equal to first whorl of

teleoconch. Teleoconch of 5 whorls, with 3 spiral

cords per whorl, crossed at right angles by axial ribs a

little weaker than the cords, with a rounded bead at

each intersection; beads numbering 19-20 on last

whorl. Beads of upper cord somewhat larger than

those of lower cords. A fourth beaded cord emerging

from suture at base of last whorl; base smooth and

concave, with 3 spiral threads on anterior canal.

Aperture circular. Colour pale brown to cream,

protoconch of the same colour. A few specimens with

one more distorted whorl, wider but constricted at

base.

Size: maximum height 2.3 mm, maximum width 1

mm; height of protoconch 0.45 mm; width of

protoconch at base 0.30 mm.

Locality. Found dead in hand dredged sand at 10-20m,

off Saint-Gilles-les-Bains.

Remarks. Our specimens match exactly the

description and figure given by Laseron (Joculator

minima Laseron, 1955, type locality Hope Isl .). This

species 1s distinguished by the width of the

protoconch, especially its last whorl, occasionally

wider on our specimens than in Laseron’s figure.

Joculator minutissima (Thiele, 1925).

Plate 3, C; colour plate I, Fig. 16

Material examined. 1 spmn MNHN; 26 spmns (4

with complete protoconch) coll. M. Jay; 3 spmns

coll. J. Drivas.

Description. Shell pupiform and very short, wide

compared to its height, its summit topped by a

prominent cylindrical but short protoconch.

Protoconch of 2.25 smooth whorls, the first one

rounded, the second one weakly convex. Teleoconch

of 3 whorls, with 3 subequal spiral cords, crossed at

right angles by weaker axial ribs, with a rounded bead

at each intersection: beads numbering 18 on

penultimate whorl. Colour plain brown, protoconch

paler.

Size: maximum height 1.5 mm; maximum width 0.6

mm; height of protoconch 0.25 mm; width of

protoconch at base 0.21 mm.

Locality. Found dead in hand-dredged sand at 10-54

m, off Saint-Gilles-les-Bains and Possession-Bay.

Remarks. Our specimens were compared with the

holotype of Cerithiopsis minutissima Thiele, 1925,

type locality Nias Isl. in MNK (Lot Nr 102724) and

found identical.

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Joculator pulvis ({ssel, 1869)

Plate 3, F; colour plate I, Fig. 17

Material examined. 1! spmn MNHN, 7 spmns (2 with

complete protoconch) coll. M. Jay. 7 spmns coll. J.

Drivas.

Description. Shell pupiform, tapering towards apex,

with base strongly constricted. Protoconch prominent,

almost cylindrical, made of 3.5 convex smooth whorls

with rounded apex, and under microscope a spiral row

of fine granules just above lower suture. Teleoconch of

6 whorls, with 3 subequal spiral cords, and distinct

axial ribs, equal to the cords, crossing them at right

angles, with a small rounded bead at each intersection;

beads widely spaced, numbering 16 or 17 on

penultimate whorl. A fourth spiral cord emerging from

suture, weaker and finely beaded, at base of last whorl.

Base with a fifth smooth cord. The upper cord and its

beads reddish brown, ground and the 2 other cords

very pale brown. Protoconch white.

Size: maximum height 2.9 mm; maximum width 1.2

mm; height of protoconch 0.46 mm; width of

protoconch at base 0.35 mm. Some variability in size

exists between specimens.

Locality. Found dead in hand-dredged sand at 10-20

m off Saint-Gilles-les-Bains.

Remarks. Our specimens were compared and found

identical to the specimen stored in the MNHN

typothèque, and labelled Cerithium (Cerithiopsis?)

pulvis, type locality Suez roadstead. Specimens match

Savigny’s figure (Bouchet & Danrigal, 1982). This

species resembles Joculator eudeli n.sp. which has the

same colour pattern, but differs in its greater width, its

base bearing one smooth spiral cord instead of 2, the

colour of the upper cord being more reddish and less

blackish, and the fact that this colour does not reach

the suture. The species may be compared with

Cerithiopsis insignis E.A.Smith, 1906, from Port

Shepstone, South Africa, but differs from it in having

6 teleoconch whorls instead of 8, its 3.5 whorled

protoconch instead of 4.5, and in its size (2.9 x 1.2

instead of 3.25 mm x Imm) (see discussion below

with Dizoniopsis herberti n.sp.). The other pupiform

species with a similar colour pattern have only 2 spiral

beaded cords, except on last whorl; Horologica

bicolor Laseron, 1956 and Horologica semipicta

Gould, 1861, are smaller; Dizoniopsis herberti n.sp.

with its 2 spiral cords per whorl, and a third cord on

the last whorl, is easily separated by its axially costate

protoconch.

Joculator christiaensi n.sp.

Plate 3, E; colour plate I, Fig. 18

Material examined. 1! spmn MNHN; 18 spmns (1

with complete protoconch) coll. M. Jay; 2 spmns

coll. J. Drivas.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Description. Shell pupiform, strongly swollen at

middle, strongly constricted at base, and topped by a

stick-shaped protoconch. Protoconch cylindrical and

narrow, Of 3 slightly convex smooth whorls, with a

rounded apex, limited from teleoconch by an oblique

line marking change of colour, and appearance of

adult sculpture. Teleoconch of 4 whorls, flat-sided,

with 3 spiral cords per whorl, and finer axial ribs

crossing them at right angles; one rounded bead at

each intersection, beads numbering 16 per whorl. On

the 2 middle whorls, beads of the upper row are

stronger. A fourth weaker cord, finely beaded,

emerging from suture at base of last whorl. On last

whorl, beads becoming smaller towards the outer lip of

aperture. À fifth smooth cord on base. Aperture small,

roundly quadrate. Colour dark brown, the upper cord

and its beads, and the whole first whorl and

protoconch whitish.

Size: holotype total height 1.7 mm; maximum width

0.6 mm height of protoconch 0.34 mm; width of

protoconch at base 0.24 mm. Paratype height 1.9 mm:

maximum width 1 mm.

Type locality. Found dead in hand-dredged sand at 55

m, Saint-Gilles-les-Bains, between harbour and

Boucan-Canot beach.

Type material. Holotype in MNHN, paratype 1 coll.

M. Jay.

Etymology. Dedicated to J. Christiaens (Belgium).

Remarks. This species is characterized by its swollen

shape, with its prominent narrow protoconch, which

resembles that of Joculator minutissima (Thiele, 1925)

from which it differs in its larger size, stronger beads,

and distinct colour pattern. The nearest species is

Joculator ovata Laseron, 1956, from NE Australia

(Hope Isl.), the illustrated type of which has a broken

protoconch with only 1.5 whorls left, but has a colour

pattern and general outline, including the last whorl of

protoconch, similar to the present species. However, /.

christiensi n.sp. differs from it in having a teleoconch

of 4 whorls on all specimens instead of 5, and 16

beads per whorl instead of 14. Joculator albordina

Laseron, 1956, (type locality Michaelmas Cay, NE

Autralia), which has a comparable colour pattern is

larger in size, With teleoconch of 6 whorls instead of 4

in mature specimens, and 20 beads per whorl instead

of 16.

Joculator eudeli n.sp.

Plate 3, D; colour plate I, Fig. 19

Material examined. 2 spmns MNHN: 41 spmns (3

with complete protoconch) coll. M. Jay.

Description. Shell very small, pupiform, with a

nearly cylindrical protoconch, base strongly

constricted. Protoconch of 3.5 whorls, slightly

convex and smooth, with a fine granulation under

optical microscope, distinct under SEM on the lower

half of first whorl; limit from teleoconch oblique,

marked by change of colour, adult sculpture arising

progressively one half whorl earlier. Teleoconch of 5

whorls, suture wide and deeply impressed. 3 spiral

cords per whorl, subequal, crossed at right angles by

equal axial ribs, with a strong rounded bead at each

intersection; beads close-set, numbering 20-21 per

whorl. A fourth beaded spiral cord, equal to the other

ones, emerging from suture at base of last whorl. 2

more spiral cords on base, weaker and smooth,

followed by 2 fine threads on anterior part. Aperture

quadrangular. Colour beige or fawn on fresh

specimens, suture and upper cord blackish-brown

with reddish gleam: fourth cord on last whorl and

base blackish-brown.

Size: holotype height 1.9 mm; maximum width 0.8

mm, height of protoconch 0.40 mm; widh of

protoconch at base 0.27 mm.

Type locality. Found dead in hand-dredged sand at

30 m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNEN,

paratypes 2 to 6 coll. M. Jay.

Etymology. Dedicated to Emile Eudel, merchant

ship captain in the last century, who was one of the

earlier collectors of Reunion fauna.

Remarks. This species resembles Joculator pulvis

(Issel, 1869), but differs in its smaller size, the 2

smooth spiral cords on base instead of 1, the blackish-

brown colour of the upper cord instead of reddish-

brown, and in its suture being tinted by this same

colour. It differs from Cerithiopsis (Joculator)

insignis E.A.Smith, 1906, as discussed above with

Joculator pulvis. Other pupiform species with similar

colour pattern (Horologica bicolor Laseron, 1956,

Horologica semipicta Gould, 1861, and Dizoniopsis

herberti n.sp.) have only 2 spiral beaded cords per

whorl, a third cord only on last whorl; Dizoniopsis

herberti n.sp. is easily distinguished by its axially

ribbed protoconch.

Joculator fischeri n.sp.

Plate 3, A; colour plate I, Fig. 20

Material examined 2 spmns MNHN: 8 spmns (4

with complete protoconch) coll. M. Jay.

Description. Shell fusiform, with a nearly cylindrical

protoconch, base constricted. Protoconch of 3.5

smooth convex whorls, with rounded apex, its limit

from teleoconch oblique marked by change of colour

and appearance of adult sculpture. Teleoconch of 7

whorls, 3 beaded spiral cords per whorl; axial ribs

distinct, equal to cords, crossing them at right angles,

with a rounded bead at each intersection; beads rather

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

small, appearing widely spaced, and numbering 19-

20 on last whorl; the middle cord reduced to a fine

thread on first whorl, obviously weaker than the other

ones on following whorls, and still a little weaker on

last whorl, the upper cord a little stronger than the

lower one. A fourth beaded spiral cord, weaker,

emerging from suture at base of last whorl. Base

excavated with a weak smooth cord at mid-height.

Aperture circular. Colour glossy brown, the upper

cord a little darker, protoconch paler.

Size: holotype total height 2.3 mm, maximum width

0.8 mm; height of protoconch 0.35 mm; width of

protoconch 0.23 mm.

Type locality. Found dead in hand-dredged sand at

30 m off Souris-Chaude, Trois-Bassins.

Type material. Holotype and paratype 1 in MNHN,

paratypes 2 to 5 coll. M. Jay.

Etymology. Dedicated to Pr.P.H. Fischer, late

professor in zoology at Saigon.

Remarks. This species is smaller than Joculator

keratochroma n.sp. and Joculator lozoueti n.sp. It

differs from other species of brown Joculator of the

same size, namely J myia n.sp., J. mygaki n.sp., J.

phtyr n.sp. J psyllos n.sp. and J. thielei n.sp. in its

weaker middle cord and stronger upper cord.

Joculator keratochroma n.sp.

Plate 3, G:; colour plate I, Fig. 21

Material examined. 2 spmns MNHN: 27 spmns (1

with complete protoconch) coll. M. Jay; 9 spmns (1

with complete protoconch) coll. J. Drivas.

Description. Shell pupiform with strongly constricted

base and prominent protoconch. Protoconch high,

slightly conical with wide and rounded apex, and 4

convex smooth whorls; the last whorl a little narrower

in diameter than the first whorl of teleoconch: its limit

from teleoconch oblique, marked by change of

sculpture. Teleoconch of 7 slightly convex whorls,

suture shallow, penultimate and last whorl constricted.

3 spiral cords per whorl, subequal, crossed at right

angles by rounded axial ribs, weaker than cords. A

rounded bead at each intersection, beads numbering 17

or 18 per whorl. A fourth beaded spiral cord emerging

from suture at base of last whorl. Last whorl strongly

constricted with 4 spiral cords, their beads becoming

much weaker towards aperture. 2 more cords, fine and

smooth, on base. Aperture circular. Colour plain

horny-brown, or dark cream, ïidentical on all

specimens.

Size: holotype total height 2.5 mm; maximum width 1

mm; height of protoconch 0.30 mm; maximum width

of protoconch 0.26 mm.

Type locality. Found dead in hand-dredged sand at

10-20 m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN;

paratype 2 and 3 coll. M. Jay; paratype 4 coll. J.

Drivas.

Etymology. Named after its colour, from Greek kera,

meaning horn.

Remarks. This species differs from Joculator

melanoraphis n.sp. which has a comparable shape, in

its plain coloration and its slightly higher protoconch

(4 whorls instead of 3.5); it differs from / ridicula

(Watson,1886) in its larger size (2.5 mm instead of 1.4

mm) and in having 7 teleoconch whorls instead of 3; it

differs from / tribulationis (Hedley, 1909) in its

slightly larger size (2.5 mm instead of 2.1 mm), its 4

protoconch whorls instead of 3 and in its 6 teleoconch

whorls instead of 5: Z minima Laseron, 1955,

minutissima, (Thiele, 1925), J. thielei n.sp. are much

smaller and have different protoconchs. Cerithiopsis

(Joculator) pupula Dunker (MNK lot Nr 6824) is

more ventricose.

Joculator laseroni n.sp.

Plate 5, A; colour plate I, Fig. 22

Material examined. 2 spmns MNHN;:

with complete protoconch) coll. M. Jay.

18 spmns (3

Description. Shell small, fusiform with constricted

base. Protoconch rather cylindrical with rounded apex,

comprising 3 convex smooth whorls, its limit from

teleoconch very oblique, marked by change of colour

and sculpture. Teleoconch of 5 whorls, suture poorly

visible, 3 spiral cords per whorl, crossed at right angles

by slightly weaker axial ribs, with a large rounded

bead at each intersection; spiral cords subequal, beads

numbering 16 per whorl, and appearing close-set. A

fourth beaded spiral cord emerging from suture at base

of last whorl. A fifth weak and unbeaded cord on base.

Aperture circular. Colour deep brown, the beads a

little paler than ground; protoconch white.

Size: holotype total height 2.3 mm, maximum width

0.9 mm; height of protoconch 0.26 mm; width of

protoconch at base 0.22 mm.

Type locality. Found dead in hand-dredged sand at 30

m, off Boucan-Canot beach, Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN;:

paratypes 2 to 4 coll. M. Jay.

Etymology. Dedicated to C.F. Laseron, on account of

its resemblance to Joculator subula described by him.

Remarks. This species matches the description of

Joculator subula Laseron 1956, (Bowen, Queensland)

except for its 3 protoconch whorls instead of 5.5

JAY & DRIVAS Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

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PLATE 3. Fig. A. Joculator fischeri n.sp. Off Souris-Chaude, Trois-Bassins, 30 m:; holotype, height 2.3 mm:

MNAN. Fig. B.J/. minima Laseron, 1955. Off Saint Gilles les Bains, 10-20 m; height 2.3 mm; MNAN.

Fig. C. J. minutissima (Thiele, 1925). Off Saint Gilles les Bains and Possession Bay 10-54 m:; height 1.5 mm:

MNAN. Fig. D. J. eudeli n.sp. Off Saint Gilles les Bains, 30 m:; holotype, height 1.9 mm; MNHN.

Fig. E. J. christiaensi n.sp. Off Saint Gilles les Bains, 55 m; holotype, height 1.7 mm; MNAHN. Fig. F. J. pulvis

(Issel, 1869). Off Saint Gilles les Bains, 10-20 m:; height 2.9 mm; MNHN. Fig. G. J. keratochroma n.sp. Off

Saint Gilles les Bains, 10-20 m; holotype, height 2.5 mm; MNHN.

Scale bars: S (shells): 1 mm; P (protoconchs): 100 um.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

whorls on Z subula. It differs from Joculator salvati

n.sp., which has the same shape and colour, in its

smaller size (2.3 mm against 3.6), in its protoconch (3

whorls against 2.25) and its plain colour.

Joculator lozoueti n.sp.

Plate 4, B; colour plate I, Fig. 23

Material examined. 2 spmns MNHN; 53 spmns (2

with complete protoconch) coll. M. Jay: 10 spmns

coll. J. Drivas.

Description. Shell pupiform with strongly constricted

base and prominent protoconch. Protoconch

cylindrical, made of 2.5 convex whorls, smooth and

translucent, with very rounded apex, its limit from

teleoconch very oblique, marked by change of colour

and sculpture. Teleoconch of 7 slightly convex whorls

with shallow suture. 3 spiral cords per whorl,

subequal, crossed at right angles by weak axial ribs. A

strong rounded bead at each intersection, beads very

close-set and numbering 17 or 18 per whorl. A fourth

spiral cord emerging from suture on base of last whorl.

On the 4 cords of last whorl, towards aperture, beads

becoming weaker and axially elongate, the lowest cord

unbeaded. A fifth fine smooth cord on base. Aperture

oval. Colour plain dark brown, protoconch white.

Size: holotype total height 3.2 mm; maximum width

1.3 mm; height of protoconch 0.42 mm; maximum

width of protoconch 0.34 mm.

Type locality. Found dead in hand-dredged sand at

10-20 m off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN:;

paratypes 2 and 3 coll. M. Jay; paratypes 4 and 5

coll. J. Drivas.

Etymology. Dedicated to Mr. Lozouet, MNHN Paris.

Remarks. This species differs from Joculator

keratochroma n.sp. in its slightly higher size, shorter

protoconch (2.5 whorls instad of 4), and darker

colour. It differs from J. melanoraphis n.sp. in its

shorter protoconch (2.5 whorls instead of 3.5), and

plain coloration; it differs from J. subula Laseron,

1856 in its larger size, less blackish colour, and

shorter protoconch (2.5 whorls against 5.5); it differs

from J continens Laseron, 1856 (Bowen,

Queensland) in the above characters and its slightly

shorter and wider protoconch; it differs from J

melania Laseron, 1856 (Bowen, Queensland) in its

shorter protoconch (2.5 whorls instead of 4.5).

Joculator megacephala n. sp.

Plate 5, B; colour plate I, Fig. 24

Material examined. 2 spmns MNHN; 20 spmns (13

with complete protoconch) coll. M. Jay.

Description. Shell very small and pupiform, strongly

constricted at base, with à prominent rather large

protoconch. Protoconch conical of 5.5 convex smooth

whorls, forming by itself about 1/3 of the total height

of the shell, with narrow and rounded apex, its last

whorl of large diameter compared with teleoconch, its

lower limit along an oblique line marked by change of

sculpture and colour; the specimen photographed

under SEM shows a worn surface. Teleoconch of 4

whorls with 3 spiral cords per whorl, crossed at right

angles by slightly weaker axial ribs, with a rounded

bead at each intersection; 20 beads per whorl. A fourth

weaker and more finely beaded cord emerging from

suture at base of last whorl. Aperture oval. Colour dark

cream, protoconch pale brown.

Size: holotype height 1.8 mm., maximum width 0.6

mm; height of protoconch 0.42 mm; width of

protoconch at base 0.35 mm.

Type locality. Found dead in hand-dredged sand at

10-20 m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN:;

paratypes 2 to 14 coll. M. Jay.

Etymology. Named after its shape, from Greek,

meaning “large head”.

Remarks. This species has the same shape as

Horologica macrocephala (Laseron, 1956), but differs

from it in the 3 spiral cords instead of 2 on the

teleoconch whorls, its weaker, more numerous and

more close-set beads, and its smaller size. It differs

from Joculator minima Laseron, 1956 in its smaller

size (1.8 mm instead of 2.3mm), its proportionally

larger protoconch (0.5 mm for the 2 species), its colour

(cream instead of brown), and its subequal beaded

cords, the beads of the upper cord being somewhat

larger on J. minima.

Joculator melanoraphis n.sp.

Plate 5, F; colour plate I, Fig. 25

Material examined. 2 spmns MNHN; 24 spmns (11

with complete protoconch) coll. M. Jay; 5 spmns (2

with complete protoconch) coll. J. Drivas.

Description. Shell pupiform, rather globose, with

strongly constricted base and high protoconch.

Protoconch of 3.5 smooth convex and translucent

whorls, rounded apex, well delimited from teleoconch

by an oblique line marking change of sculpture.

Teleoconch of 7 whorls with straight sides and shallow

suture. 3 spiral cords per whorl, the uppermost one a

little weaker on first whorl, but equal to the other ones

on following whorls. Rounded and widely-spaced

axial ribs, a little smaller than cords, crossing them at

right angles, with a rounded bead at each intersection,

beads close-set and numbering 16 or 17 per whorl. On

fresh specimens, very fine axial riblets visible under

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

microscope on whole surface including cords and ribs,

numbering 6 to 7 in each interval. A fourth spiral cord

emerging from suture at base of last whorl, with only

weak swellings rather than beads; a fifth smooth cord

on base of mature specimens. Aperture circular.

Colour plain beige to pale brown; suture, base and

protoconch darker brown. On fresh specimens, the

upper cord is a little darker than the other ones, but

remains clearly paler than suture.

Size: holotype total height 2.9 mm, maximum width 1

mm; height of protoconch 0.44 mm; width of

protoconch at base 0.30 mm.

Type locality. Found dead in hand-dredged sand at

10-30 m, off Saint-Gilles-les-Bains, between harbour

and Boucan-Canot beach.

Type material. Holotype and paratype 1 in MNHN:

paratypes 2 to 12 coll. M. Jay; paratypes 13 and 14

coll. J. Drivas.

Etymology. Named on account of its colour pattern

from Greek meaning brown suture.

Remarks. Dizoniopsis herosae n. sp. which has the

same colour and a brown suture, is easily separated

from this species by its 2 spiral cords instead of 3, and

its axially ribbed protoconch. The other species

discussed below lack the darker suture: Joculator

granata Kay, 1979, is higher and less constricted

basally, and its upper cord is weaker on all whorls; it

also has a crimped suture on the protoconch whorls.

Joculator keratochroma n. sp. is of the same shape,

but is smaller and has a plain coloration; Joculator

lozoueti n.sp. can be separated on account of its more

ventricose shape, its white protoconch and the plain,

darker colour of the teleoconch; Bittium (Joculator)

tenthrenois Melvill, 1896 from Bombay, differs in its

more ventricose shape and its plain brown colour:

Cerithiopsis (Joculator) ridicula Watson, 1866 from

Wednesday Isl., NE Australia, differs in having only 3

teleoconch whorls, and its upper cord is weaker on all

whorls, and its coloration plain; Joculator minima

Laseron, 1955, J/ niasensis (Thiele, 1925), and /

psyllos n.sp. are of similar shape, but are much

smaller.

Joculator mygaki n. sp.

Plate 5, E; colour plate I, Fig. 26

Material examined. 2 spmns MNHN; 15 spmns (10

with complete protoconch) coll. M. Jay; 5 spmns

coll. J. Drivas.

Description. Shell very small and pupiform, its

slender apex topped by protoconch, and constricted at

base. Protoconch of 5.5 convex smooth whorls, the last

one wide and ventricose, its limit from teleoconch

marked by beginning of sculpture along an oblique

line; the earlier whorls regularly tapering to the

rounded but very narrow apex. Teleoconch of 4.5

whorls, with 3 subequal spiral cords per whorl, crossed

at right angles by weak axial ribs, with a rounded bead

at each intersection; beads close-set, numbering 23-24

per whorl. A fourth beaded spiral cord emerging from

suture at base of last whorl, weaker than the other

ones. 2 weak and unbeaded spiral cords on base.

Aperture rounded. Colour plain orange-brown, vivid

when fresh, the 3 last whorls of protoconch brown, the

earlier 2 white.

Size: holotype total height 1.9 mm, maximum width

0.6 mm; height of protoconch 0.42 mm; width of

protoconch at base 0.30 mm.

Type locality. Found dead in hand-dredged sand at

30m, off Boucan-Canot beach, Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNAN;:

paratypes 2 to 10 coll. M. Jay.

Etymology. Named for its size, after Greek meaning

midge.

Remarks. This species resembles Joculator minima

Laseron in its shape, but differs from it in its smaller

size (1.9 mm instead of 2.3 mm) and its 3 rows of

equal beads. It differs from Bittium (Joculator)

uveanum Melvill & Standen, 1896, from Loyalty Isl.

in its smaller size (1.9 mm instead of 2.3 mm), its less

swollen shape, its finer and more numerous beads, and

its colour; it differs from Joculator myia n.sp. in its

protoconch (brown tinted and more acute) and in its

more orange colour.

Joculator myia n. sp.

Plate 4, C; colour plate I, Fig. 27

Material examined. 2 spmns MNHN: 10 spmns (7

with complete protoconch) coll. M. Jay.

Description. Very small pupiform shell, more pointed

and narrow towards apex than towards the constricted

base. Protoconch of 5 whorls, conical, the 2 last

whorls convex and ventricose, the 3 earlier tapering

regularly towards the rounded and rather wide apex:; a

narrow spiral row of fine and very short axial riblets in

suture, overlapping on the upper 1/8 of lower whorl:

the earlier whorls finely granulose under SEM. Limit

from teleoconch very oblique marked by change of

colour and appearance of adult sculpture. Teleoconch

of 4.5 whorls, with 3 spiral beaded cords per whorl,

crossed at right angles by fine axial ribs, with a

rounded bead at each intersection; the 3 beaded cords

subequal, beads close-set, and numbering 19-20 per

whorl. A fourth spiral cord, with smaller beads,

emerging from suture at base of last whorl. Aperture

circular. Colour pale golden brown, protoconch white.

Some specimens with one more wider whorl, but also

constricted at base.

Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

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PLATE 4. Fig. A. Joculator albocinctum Melvill & Standen, 1896. Off Saint Gilles les Bains, 10-20 m, height

3 mm; MNAN. Fig. B. J lozoueti n.sp. Off Saint Gilles les Bains, 10-20 m; holotype, height 3.2 mm; MNEN.

Fig. C. J. myia n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 30 m; holotype, height 2.2 mm; MNHN.

Fig. D. J granata Kay, 1979. Off Saint Gilles les Bains, 10-20 m; height 2.5 mm; MNEN. Fig. E. J. psyllos

n.sp. Off Saint Gilles les Bains, 10-20 m; holotype, height 2 mm; MNHN. Fig. F.J. salvati n.sp. Off Boucan-

Canot beach, Saint Gilles les Bains, 35 m; holotype, height 3.6 mm; MNAN. Fig. G. J. skolix n.sp. Off

Boucan-Canot beach, Saint Gilles les Bains, 35 m; holotype, height 3.2 mm; MNHN.

Scale bars: S (shells): 1 mm; P (protoconchs): 100 um.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Size: holotype total height 2.2 mm; maximum width

0.6 mm; height of protoconch 0.46 mm; width of

protoconch at base 0.34 mm.

Type locality. Found dead in hand-dredged sand at 30

m, off Boucan-Canot beach, Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN;:

paratypes 2 to 6 coll. M. Jay.

Etymology. Named for its size, after Greek meaning

fly.

Remarks. This species resembles Joculator mygaki

n.sp., but differs from it in its protoconch (wider at

apex and with a row of fine axial riblets in and under

suture), its slightly larger size (2.2 mm instead of 1.9

mm) and its darker colour. It differs from J/. granata

Kay, 1979, which has a protoconch with the same fine,

brephic riblets, in its slightly smaller size (2.2 mm

instead of 2.4 mm), its shorter and more swollen

shape, and its paler colour.

Joculator phtyr n.sp.

Plate 5, G; colour plate I, Fig. 28

Material examined. 2 spmns MNHN:; 48 spmns (5

with complete protoconch) coll. M. Jay.

Description. Shell small, pupiform, strongly

constricted at base, pointed at apex, with a high

conical protoconch; the greatest width at the lower

third of teleoconch. Protoconch conical of 5 whorls,

slightiy convex and smooth, their convexity more

marked abapically;: apex rounded, limit from

teleoconch marked by the development of the 3 cords

simultaneously in a vertical line, but without a clear-

cut boundary. Teleoconch of 4 flat whorls, 3 spiral

cords per whorl, bearing rounded beads, of equal

importance on the 3 cords, regularly diminishing in

size towards both extremities; beads numbering 19-20

per whorl. Axial ribs joining the beads poorly visible

in their intervals. A fourth beaded spiral cord,

obviously weaker, emerging from suture at base of last

whorl; beads of last whorl becoming weaker and

axially elongate towards outer lip of aperture, except

the beads of the upper cord which remain rounded.

One smooth cord on base, followed by 2 very fine

spiral threads on edge of anterior canal. Aperture

circular. Colour dark brown to blackish-brown, upper

spiral cord and its beads slightly darker. Protoconch of

the same colour.

Size: holotype total height 2 mm, maximum width 0.9

mm; height of protoconch 0.44 mm; width of

protoconch at base 0.26 mm.

Type locality. Found dead in hand dredged sand at 35

m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN;:

paratypes 2 to 6 coll. M. Jay.

Etymology. Named for its size, after Greek meaning

crablouse.

Remarks. Prolixodens sknips n.sp. has the same

shape, but is slightly smaller, with teleoconch whorls

more numerous, and axially ribbed protoconch.

Dizoniopsis gothica n. sp. bears only 2 beaded spiral

cords per whorl, instead of 3, and has a ribbed

protoconch. Joculator subula Laseron, 1956, lacks its

pointed apex. Joculator lozoueti n. sp. is more

regularly ovate and larger.

Joculator psyllos n.sp.

Plate 4, E; colour plate I, Fig. 29

Material examined. 2 spmns MNHN; 7 spmns (4

with complete protoconch) coll. M. Jay; 2 spmns

with complete protoconch coll. J. Drivas.

Description. Shell of very small size, pupiform with

strongly constricted base. Protoconch not prominent of

1.25 smooth whorls, appearing finely punctate under

microscope, apex rounded; limit from teleoconch clear

and oblique, marked by change of colour, but 2 very

fine beaded cords begin on last 1/4 whorl. Teleoconch

of 6 slightly convex whorls, suture moderately

impressed. Three spiral cords per whorl, subequal in

middle of shell, the upper cord slightly weaker on

earlier whorls. Axial ribs, weaker than cords, crossing

them at right angles, with a rounded bead at each

intersection, numbering 16 per whorl. A fourth weaker

spiral cord emerging from suture at base of last whorl.

On last whorl, strongly constricted at base, beads

become smaller towards aperture and obsolete on

lower cords. Several very fine spiral threads on base.

Colour pale brown, apex a little paler, protoconch

white.

Size: holotype total height 2.0 mm: maximum width

0.8 mm; height of protoconch 0.19 mm, maximum

width of protoconch 0.18 mm.

Type locality. Found dead in hand-dredged sand at

10-20 m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN:

paratypes 2 to 5 coll. M. Jay; paratypes 6 and 7 coll.

J. Drivas.

Etymology. Named on account of its small size, after

Greek meaning “flea”.

Remarks. This species differs from other Joculator

species in its very short protoconch. Besides, it differs

from Joculator thielei n.sp. in its more elongate

outline and slightly paler colour; it differs from

tribulationis (Hedley, 1909) in its less numerous

beads, smaller size, and paler colour; it differs from

Bittium (Joculator) uveanum Melvill & Standen, 1896

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

from Lovalty Isl. in its much shorter protoconch,

narrower shape and colour.

Joculator salvati n.sp.

Plate 4, F: colour plate I, Fig. 30

Material examined. ! spmn MNHN; 3 spmns (1

with complete protoconch) coll. M. Jay.

Description. Shell fusiform, slightly constricted at

base. Protoconch conical, extending the general

outline of teleoconch, made of 2.5 smooth whorls,

under SEM showing a worn surface; apex wide and

rounded, limit from teleoconch ill-defined by the

progressive development of adult sculpture.

Teleoconch of 7 whorls with flat sides, with 3 spiral

cords per whorl, crossed at right angles by axial ribs of

similar strength, with a rounded bead at each

intersection; beads rather widely-spaced and

numbering 18-19 per whorl. A fourth beaded cord

emerging from suture at base of last whorl. Base with

a weak smooth cord. Aperture roundly quadrate.

Colour dark brown, beads a little paler, mostly those of

the upper cord. First whorl and protoconch white.

Size: holotype total height 3.6 mm; maximum width

1.1 mm: height of protoconch 0.69 mm; width of

protoconch at base 0.61 mm.

Type locality. Found dead in hand-dredged sand at 35

m, off Boucan-Canot beach, Saint-Gilles-les-Bains.

Type material. Holotype in MNHN; paratype 1 coll.

M. Jay.

Etymology. Dedicated to B. Salvat, a French

malacologist.

Remarks. This species has the same shape and colour

as Joculator psyllos n. sp. but 1s twice as high, and its

protoconch has 2.25 whorls instead of 1.25. It differs

from J. lozoueti n.sp. in its slightly larger size, less

swollen shape, and wider, less elevated protoconch.

This species resembles J. tomacula negrita Laseron,

1955, from Michaelmas Cay, NE Australia, in its

shape, size and colour, but differs from it in its more

constricted aperture, and in the paler beads of the

upper cord. The holotypes of Z 1omacula and J.

tomacula negrita Laseron, 1955 have a broken

protoconch.

Joculator skolix n.sp.

Plate 4, G:; colour plate I, Fig. 31

Material examined. 2 spmns MNHN; 9 spmns (6

with complete protoconch) coll. M. Jay.

Description. Shell fusiform, narrow, nearly cylindrical

and constricted at base. Protoconch narrow and

slender, comprising 4 smooth slightly convex whorls,

the earlier two finely granulose under microscope and

SEM, the last one obviously narrower than the first

whorl of teleoconch; suture wide, with very short axial

riblets, limited to suture, and numbering 20-22 per

whorl: limit from teleoconch a distinct oblique line

marked by change of colour, but the 3 beaded spiral

cords of adult sculpture develop progressively on last

1/2 whorl. Teleoconch of 7 flat whorls, bearing 3

spiral cords per whorl, crossed at right angles by axial

ribs, with a rounded bead at each intersection; beads

numbering 20-21 per whorl; the 3 beaded cords

subequal in strength. A fourth spiral cord emerging

from suture at base of last constricted whorl, and

underlining angle of excavated base. Aperture broken

on the type, but circular on other specimens.

Columella smooth. Colour plain creamy-white.

Size: holotype total height 3.2 mm; maximum width 1

mm; height of protoconch 0.48 mm; width of

protoconch at base 0.29 mm.

Type locality. Found dead in hand-dredged sand at 35

m, off Boucan-Canot beach, Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN;

paratypes 2 to 7 coll. M. Jay.

Etymology. From Greek meaning maggot, to which

this species resembles by shape and colour.

Remarks. This species may recall Cerithiopsis

eutrapela Melvill & Standen, 1896, but differs from it

in its narrower Width, its more cylindrical shape

(constricted at base, instead of conical), and its 3 rows

of beads being equal.

Joculator thielei n.sp.

Plate 5, C; colour plate I, Fig. 32

Material examined. 2 spmns MNHN: 23 spmns (9

with complete protoconch) coll. M. Jay.

Description. Shell very small, pupiform, short, with

strongly constricted base. Protoconch prominent,

nearly cylindrical, slightly tapering to the wide,

rounded apex, and comprising 3.25 convex whorls,

looking smooth but worn on type under SEM; limit

from teleoconch obliquely marked by change of colour

and beginning of adult sculpture. Teleoconch of 5

whorls, with 3 spiral cords per whorl, crossed at right

angles by weaker axial ribs, with a rounded bead at

each intersection; beads numbering 16 on penultimate

whorl. The 3 cords subequal on the entire shell. A

fourth spiral cord, smooth and unbeaded, emerging

from suture at base of last whorl. On last whorl, the

beads of the lower cords become smaller and

disappear near aperture. 2 more fine smooth spiral

cords on base. Aperture cireular. Colour dark blackish-

brown, the upper cord a little darker, protoconch

whitish.

Size: holotype height 1.5 mm; maximum width 0.5

mm; height of protoconch 0.26 mm; width of

protoconch at base 0.22 mm.

JAY & DRIVAS Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

PLATE 5. Fig. A. Joculator laseroni n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 30 m:; holotype,

height 2.3 mm; MNAN. Fig. B. J/. megacephala n.sp. Off Saint Gilles les Bains, 10-20 m; holotype, height 1.8

mm; MNAN. Fig. C. J. thielei n.sp. Off Saint Gilles les Bains, 10-20 m; holotype, height 1.5 mm:

MNEN. Fig. D. J vignali n.sp. Off Souris-Chaude, Trois-Bassins, 30 m; holotype, height 1.4 mm; MNHN.

Fig. E. J. mygaki n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 30 m; holotype, height 1.9 mm:

MNAN. Fig. F. J. melanoraphis n.sp. Off Saint Gilles les Bains, 10-30 m; holotype, height 2.9 mm; MNHN.

Fig. G. J. phtyr n.sp. Off Saint Gilles les Bains, 35 m; holotype, height 2 mm; MNAN.

Scale bars: S (shells): 1 mm; P (protoconchs): 100 um.

JAY & DRIVAS

Type locality. Found dead in hand-dredged sand at

10-20 m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN;:

paratypes 2 10 10 coll. M. Jay.

Etymology. Dedicated to Pr. J. Thiele, who described

several species of Cerithiopsidae.

Remarks. This species was compared with the

holotype of Cerithiopsis niasensis, Thiele, 1925, from

Nias Isl. (MNK lot Nr 102725 ) which has same shape

and size, but has a broken protoconch with only a

piece of the last smooth whorl left, and has 2 spiral

cords per whorl, 3 cords on the last whorl only: thus,

following our criteria, this species should be attributed

to the genus Horologica. Our new species is less

ventricose than Joculator minutissima Laseron, 1955.

It differs from Joculator psyllos n. sp. in its more

swollen shape and its higher protoconch (3.25 whorls

instead of 1.5). It differs from Joculator phtyr n.sp. in

its more ovate shape and its protoconch more

cylindrical than conical of 3.5 whorls instead of 5.

Joculator vignali n.sp.

Plate 5, D; colour plate I, Fig. 33

Material examined. 1! spmn MNHN; 2 spmns (1

with complete protoconch) coll. M. Jay.

Description. Shell very small, fusiform, with

cylindrical protoconch, base constricted. Protoconch

of 3 moderately convex whorls, with a rounded apex,

smooth but granulose under microscope, well

delimited from teleoconch by an oblique line marked

by change of colour and beginning of adult

sculpture. Teleoconch of 4 whorls, with 3 subequal

beaded spiral cords per whorl, crossed at right angles

by axial ribs, a rounded bead at each intersection:

beads numbering 16 per whorl. A fourth weaker

beaded spiral cord emerging from suture at base of

last whorl. A fifth spiral cord still more weakly

beaded at mid-height of base. Aperture circular.

Colour golden brown, protoconch white.

Size: holotype total height 1.4 mm; maximum width

0.5 mm; height of protoconch 0.33 mm, width of

protoconch at base 0.22 mm.

Type locality. Found dead in hand-dredged sand at

30m off Souris-Chaude, Trois-Bassins.

Type material. Holotype in MNHN\: paratype 1

coll. M. Jay.

Etymology. Dedicated to Vignal, malacologist who

worked on material from Reunion.

Remarks. This species differs from Joculator

megacephala n.sp., which is smaller, in its very

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

different protoconch, cylindrical of 3 whorls instead of

conical of 5.5 whorls; It differs from JZ. mygaki n.sp., J.

myia n.sp., and J. phtyr n.sp. which are a little larger,

by its cylindrical protoconch instead of conical. II

differs from Z thielei n.sp. which is the most similar

species, in its smaller size, its more slender shape, and

its higher protoconch.

Genus Horologica Laseron, 1956

Type species Horologica bicolor Laseron, 1956,

Queensland: Shell fusiform with constricted base,

teleoconch with 2 beaded spiral cords, protoconch

smooth or punctate, without axial ribs or spiral cords.

Horologica balteata Watson, 1886.

Plate 7, A; colour plate IL, Fig. 34

Material examined. 1 spmn MNHN;: 3 spmns coll.

M. Jay; 3 spmns coll J. Drivas; none having a

complete protoconch.

Description. Shell pupiform, strongly swollen at

middle and constricted at base, apical angle 50°.

Protoconch known only by its 2 last smooth whorls,

extending the outline of teleoconch, separated from it

by an indistinct line marking the beginning of adult

sculpture. Teleoconch of 6 whorls, with 2 spiral cords

per whorl, crossed by finer axial ribs, with a large

rounded bead at each intersection; 17 or 18 beads per

whorl; beads of upper cord stronger, axially elongate

on penultimate whorl, then incised and divided by a

spiral furrow, forming thus a third cord on last whorl,

the lower cord always weaker. One more beaded cord,

much weaker than the other ones, emerging from

suture at base of last whorl. On last part of the very

constricted last whorl, the upper cord remains strong,

the lower 3 weaker, beads reduced to single swellings;

a fine smooth spiral thread on base. The 2 earlier

whorls plain white, protoconch brown; on the other

whorls of teleoconch, the upper beaded cord white, the

lower cord and base pale brown.

Size: maximum height with broken protoconch 3 mm:

width 1.1 mm.

Locality. Found dead in hand-dredged sand at 12 m,

off Saint-Gilles-les-Bains.

Remarks. This species, characterised by its shape and

colour pattern, matches Watson’s description and

figure in spite of a slight difference in size (2.21 mm

for Watson’s holotype instead of 2.30 to 3.4 for our

specimens). It also matches the description and figure

of Cerithiopsis aeolomitres Melvill & Standen, 1896,

which has the same size (2.26 mm for holotype);

Cerithiopsis aeolomitres Melvill & Standen, 1896, is

thus a junior synonym of Horologica balteata

(Watson, 1886). Our specimens differ from

Cerithiopsis perligera Thiele, 1925 (MNK lot Nr

67483 ) which has the same colour pattern, in is

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

obviously smaller size, and its 6 teleoconch whorls

instead of 9.

Horologica bicolor Laseron, 1956.

Plate 7, B; colour plate Il, Fig. 35

Material examined. 1! spmn MNHN, 7 spmns coll.

M. Jay; none having a complete protoconch.

Description. Shell small and pupiform, more slender

towards apex than towards the constricted base.

Protoconch of our specimens known only by its last

smooth whorl, separated from teleoconch by the

progressive growth of adult sculpture. Teleoconch of 6

whorls, with 2 spiral cords per whorl, crossed by axial

ribs of equal strength, with a more or less rounded

bead at each intersection; beads numbering 16-17 per

whorl; intervals between cords rather wide. On earlier

whorls, lower cord stronger, but the 2 cords equal in

the middle of the shell. On last whorl, beads of upper

cord incised then divided by a spiral furrow, the

median beaded cord so born remaining slightly weaker

than the other ones; a fourth spiral cord, with small

beads, emerging from suture at base of last whorl. On

last whorl near aperture, beads become weaker and

axially elongate. One more spiral cord, smooth, at

mid-height of base. Aperture quadrangular. Colour

cream, the upper cord and its beads chocolate brown.

Size: height without protoconch: 2.1 mm; width 0.9

mm.

Locality. Found dead in hand-dredged sand at 30m,

off Saint-Gilles-les-Bains.

Remarks. Our specimens match the description and

figure of Horologica bicolor Laseron 1956, type

locality Michaelmas Cay, NE Australie, but are

smaller (2,1 mm instead of 3.2 mm for Laseron); the

specimens from Réunion could be considered as a

dwarf variety of the species. The species may be

confused with several other bicolored species: it

differs from Joculator pulvis (Issel, 1869), Joculator

eudeli n.sp. and Cerithiopsis (Joculator) insignis

Smith in its 2 cords per whorl instead of 3 since

earlier whorls for the 3 last species: it differs from

Dizoniopsis herberti n.sp. which also has 2 cords, in

its smooth protoconch while D. herberti has a ribbed

protoconch, the ribs of which are well visible even on

a fragment. It differs from Horologica semipicta

(Gould, 1861) in the way the third cord is born, and

by its base.

Horologica macrocephala Laseron, 1956.

Plate 6, A; colour plate II, Fig. 36

Material examined. 1! spmn MNHN; 17 spmns, coll.

M. Jay; 5 spmns coll. J. Drivas; all with a complete

protoconch.

Description. Small shell, pupiform, constricted at

base, with prominent protoconch, its height about one

third of total height of shell. Protoconch of 5 convex

smooth whorls, the last one swollen, the earlier ones

tapering regularly to the fine and rounded apex; limit

from teleoconch oblique, marked by change of colour

and appearance of adult sculpture. Teleoconch of 4

whorls, penultimate one widest. Two beaded spiral

cords per whorl, their interval wider than the interval

at suture; axial ribs crossing the cords at right angles,

with a rounded bead at each intersection; beads

numbering 16 per whorl. A third spiral cord, unbeaded

or very weakly beaded, emerging from suture at base

of last whorl. Aperture quadrangular. Colour golden

brown.

Size: height of shell 1.7 mm, maximum width 0.5 mm;

height of protoconch 0.45 mm; width of protoconch at

base 0.28 mm.

Locality. Found dead in hand-dredged sand at 20-

30m, off Saint-Gilles-les-Bains.

Remarks. This species matches on every point the

figure and description of Horologica macrocephala

Laseron, 1956, type locality Darwin, but also found in

Indian Ocean (Christmas Isl.). The difference of size

(1.7 mm for our specimens and 1.9 mm for Laseron’s

decription) seems unimportant.

Horologica minareta Laseron, 1955.

Plate 6, B; colour plate II, Fig. 37

Material examined. ! spmn MNHN; 51 spmns (23

with complete protoconch) coll. M. Jay: 9 spmns

coll. Drivas.

Description. Shell fusiform, slender, slightly

constricted at base, apical angle 30°. Protoconch

conical, prominent, comprising 5 convex, smooth

whorls, and forming about 1/5 of total height of shell:

its penultimate whorl swollen, earlier whorls tapering

to the pointed apex; limit from teleoconch marked by

an ill defined oblique line. Teleoconch of 7 whorls,

with 2 subequal beaded spiral cords, crossed by

slightly weaker axial ribs, with a slightly axially

elongate bead at each intersection; beads numbering

22-23 on last whorl, close-set, except on base. Suture

weakly impressed. Last whorl and base constricted, the

beads of the 2 cords more axially elongate, the axial

ribs becoming more conspicuous; a third weakly

beaded spiral cord emerging from suture at base of last

whorl; one more spiral thread, smooth, at mid-height

of base. Aperture quadrangular. Colour plain dark

brown including protoconch. Some specimens ill

formed or twisted, with supplementary whorls

widening after a normally constricted whorl.

Size: maximum height 2.6 mm, maximum width 1.1

mm; height of protoconch 0.42 mm; width of

protoconch at base 0.30 mm.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Locality. Found dead in hand-dredged sand at 10-20

m, off Saint-Gilles-les-Bains.

Remarks. This species is characterized by its shape,

and by the shape of its protoconch. Our specimens are

slightiy smaller than the holotype of Horologica

minareta Laseron, 1955, (type locality Michaelmas

Cay, NE Australia), (2.6 mm instead of 2.8).

Horologica purpurea Laseron, 1955.

Plate 7, D; colour plate II, Fig. 38

Material examined. ! spmn MNHN:; 50 spmns coll.

M. Jay: 15 spmns coll. J. Drivas; 2 with a complete

protoconch.

Description. Shell fusiform, swollen, slender towards

apex, slightly constricted at base, angle at apex of

teleoconch 30°. Protoconch of 2.5 convex smooth

whorls, cylindrical in shape, with rounded apex; SEM

reveals very fine axial striae on lower part of whorls;

limit from teleoconch clear-cut and oblique, marked

by change of colour and appearance of adult sculpture.

Teleoconch of 7 or 8 whorls, bearing 2 spiral cords per

whorl, crossed by finer axial ribs, with at each

intersection, a slightly axially elongate bead; 18 or 19

beads per whorl. The 2 rows of beads subequal on

earlier whorls, but on penultimate whorl, the beads of

the upper cord are more axially elongate, becoming

incised and then divided on last whorl by a spiral

furrow. À fourth spiral beaded cord, a little weaker

than the other ones, emerging from suture at base of

last whorl: one more spiral cord, weak and smooth, at

mid-height of base. Aperture roundly quadrate. Colour

of teleoconch plain violet, paler near apex, protoconch

white.

Size: maximum height 3.1 mm; maximum width 1.2

mm; height of protoconch 0.25 mm; width of

protoconch at base 0.24 mm.

Locality. Found dead in hand-dredged sand at 10-20

m off Cap La-Houssaye, Saint-Gilles-les-Bains.

Remarks. Our specimens match the decription and

figure of Horologica purpurea Laseron, 1955, type

locality Heron Isl., but Laseron did not describe the

protoconch of the species, remarkable by its size and

its colour. All our specimens are smaller than

Laseron’s holotype (3.1 mm instead of 3.9 mm), and

are less cylindrical; but Laseron had noted size and

shape variations, with one specimen from Hope lIsl.

smaller than ours. Our specimens likewise show large

variations, with adult teleoconch height ranging from

3.1 mm and 6 whorls, to 1.4 mm and 4 whorls, with all

intermediates, each of them with a similar protoconch.

Horologica cf semipicta (Gould, 1861)

Plate 7, C: colour plate II, Fig. 39

Material examined. ! spmn MNHN; 23 spmns coll.

M. Jay; none of them having a complete protoconch.

Description. Shell small and pupiform, more slender

towards apex than towards the constricted base.

Protoconch of our specimens broken and reduced to

the last smooth whorl, limited from teleoconch by a

clear-cut oblique line. Teleoconch of 6 whorls, with 2

beaded spiral cords per whorl, widely spaced, distinct

axial ribs crossing cords at right angles, with one bead

at each intersection, beads of upper cord always a little

weaker and transversally elongate, beads of the lower

cord a little stronger and rounded. On penultimate

whorl, a spiral thread develops in interval between the

2 cords, growing into a third beaded cord on last

whorl. A fourth spiral cord emerging from suture at

base of last whorl, only weakly swollen at intersection

with axial ribs. Base smooth. Aperture roundly

quadrate. Colour cream, the upper cord and its beads

reddish-brown.

Size: height of teleoconch 2.1 mm; width 0.8 mm.

Locality. Found dead and worn in hand-dredged sand

at 30 m, off Saint-Gilles-les-Bains.

Remarks. This species resembles Aorologica

bicolor Laseron, 1956, and in the absence of

protoconchs, differs from it only in its base being

smooth instead of bearing one spiral cord, and in the

way the third spiral cord is born on last whorl. Our

specimens match the figure of Cerithiopsis semipicta

Gould, 1861 provided from Hawaii by A. Kay,

showing a third spiral beaded cord on last whorl,

without visible incision of the upper row; but

according to Kay, the species is known from Fiji as

Cerithiopsis balteata Watson, 1886 from Levuka,

Fiji, which we believe is a distinct species. The

holotype of Cerithiopsis semipicta Gould, 1861,

(type locality “China seas”), 1s, following the original

description, a little higher than our specimens (2.4

mm instead of 2.2 mm), with 7 whorls instead of 6

for our specimens; but its protoconch is not

described, nor the way the third cord is born. The

holotype of Gould figured by Johnson looks very

worn, more elongate, and does not show an incision

of the upper row of beads. The identification of our

species with Cerithiopsis (Horologica) semipicta

Gould, 1861 remains doubtful. However, this species

could be confused with other pupiform bicolor

species: Joculator pulvis (Issel, 1869), Joculator

eudeli n.sp. and Cerithiopsis (Joculator) insignis

Smith, 1906, are easily separated, having 3 beaded

cords from the earlier whorls; Dizoniopsis herberti

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

n.sp.. with its 2 beaded cords, is easily distinguished

by its axially ribbed protoconch, the ribs being

visible even on a small piece of the protoconch.

Horologica turrigera (Watson, 1886)

Plate 7, E; colour plate II, Fig. 40

Material examined. ! spmn MNHN; more than 200

spmns coll. M. Jay; more than 50 spmns coll. J.

Drivas; all with complete protoconch. This is the

only species sometimes found alive.

Description. Shell fusiform, or pupiform elongate,

base constricted. Protoconch prominent of 5.5 whorls,

slightly convex and smooth, the 2 last of equal width,

the earlier ones tapering to the rounded and narrow

apex; its limit from teleoconch axially marked by the

growth of adult sculpture. Teleoconch of 8 whorls,

with 2 beaded spiral cords per whorl, separated by an

interval wider than sutural area, and crossed by axial

ribs, a little weaker than the cords but distinct. A

strong rounded bead at each intersection, beads

numbering 18-19 on penultimate whorl. On the 3 last

whorls, beads of upper cord stronger and axially

elongate, beads of the lower cord remaining rounded.

On penultimate whorl, the upper beads incised by a

spiral furrow, then divided on last whorl, thus forming

a third cord. A fourth cord emerging from suture at

base of last whorl, weaker with slight swellings rather

than beads. One more spiral cord, smooth, at mid-

height of base. Some specimens twisted, with one

more whorl, widening after the normal constricted

whorl. Colour very pale brown, the apex of the shell

and the upper beaded cord yellowish white,

protoconch white or weakly tinted.

Size: maximum height 3.7 mm; maximum width 1.2

mm; height of protoconch 0.45 mm; width of

protoconch at base 0.30 mm.

Locality. Commonly found dead in hand-dredged

sand at 10-20 m, off Saint-Gilles-les-Bains.

Remarks. Our specimens have been found identical to

the type of Cerithiopsis turrigera Watson,1886, type

locality Honolulu, in NHM (Lot Nr 1887.2.9.1643.4).

Horologica anisocorda n. sp.

Plate 6, C; colour plate II, Fig. 41

Material examined. 1 spmn MNHN; 15 spmns (2

with complete protocench) coll. M. Jay.

Description. Shell fusiform with strongly constricted

base. Protoconch continuing the general outline of

teleoconch, conical, comprising 4 smooth whorls

tapering to rounded apex; a small spiral thread above

lower suture; limit from teleoconch oblique marked by

change of colour and beginning of adult sculpture.

Teleoconch of 6 whorls, 2 spiral cords per whorl,

crossed at right angles by subequal axial ribs, with a

rounded bead at each intersection; beads numbering 18

per whorl. From the second whorl, beads of upper cord

clearly stronger, and becoming progressively more

axially elongate towards anterior end; on last whorl,

beads of upper cord incised, then divided in 2 parts by

a spiral furrow, beads of upper cord remaining larger.

A fourth spiral cord, weakly beaded, emerging from

suture at base of last whorl. Two more spiral unbeaded

cords on base. Aperture rounded. Colour blackish-

brown, protoconch white.

Size: holotype total height 2.2 mm, maximum width

0.7 mm; height of protoconch 0.35 mm, width of

protoconch at base 0.27 mm.

Type locality. Saint-Gilles-les-Bains, in hand-

dredged sand at 30 m.

Type material. Holotype in MNHN, paratypes 1 and

2 coll. M. Jay.

Etymology. Named on account of its 2 strongly

unequal beaded cords.

Remarks. This species resembles ÆHorologica

telegraphica Hedley, 1909, (type locality Hope Is.) in

size, shape, and its 2 unequal beaded cords, but it

differs from it in its 6 whorls teleoconch instead of 5,

in its 2 beaded cords being more widely spaced, its

more numerous beads per whorl, and in the beads of

the upper row being divided on last whorl. The

protoconch of FH. telegraphica Was not described by

Hedley nor by Laseron who figured the species. Our

new species differs from Horologica glaubrechti n. sp.

which is of the same size and colour, in its 2 rows of

beads being strongly unequal on the whole height, and

in its protoconch of 4 whorls instead of 5. It differs

from Horologica minareta Laseron, 1956 in its 2

unequal rows of beads, its slightly smaller size, its

darker coloration and its 4 whorled protoconch instead

of 5.

Horologica glaubrechti n.sp.

Plate 6, D; colour plate II, Fig. 42

Material examined. 2 spmns MNHN; 9 spmns (7

with complete protoconch) coll. M. Jay; 8 spmns

coll. J. Drivas.

Description. Shell fusiform, surmonted by a wide and

conical protoconch. Protoconch of 5 whorls, convex

and looking smooth, but with very fine granulations

under SEM; last whorl as wide as the first whorl of

teleoconch, separated from it by an oblique line

marking the appearance of adult sculpture; earlier

whorls regularly tapering towards the rounded apex.

Teleoconch of 5 whorls, with 2 spiral cords per whorl,

crossed at right angles by axial ribs equal to the cords,

with a rounded bead at each intersection; beads

numbering 19-20 per whorl, of equal strength on the 2

cords, becoming axially elongate on last whorl; beads

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Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

contiguous on cords, but separated axially from the

other row, leaving axial ribs clearly visible. On last

whorl beads of upper cord become slightly stronger,

then incised on last half-whorl, then divided near

aperture by a spiral furrow, while all beads become

progressively smaller; one more spiral cord, weaker

and finely beaded, emerging from suture at base of last

whorl; a fourth spiral cord on base, unbeaded, and

followed by 4 very fine spiral threads. Aperture

circular. Colour plain dark brown, with the upper cord

and its beads a little darker, the earlier whorls paler:

protoconch white.

Size: holotype total height 2.3 mm; maximum width of

shell 1 mm; height of protoconch 0.45 mm; width of

protoconch at base 0.30 mm.

Type locality. Off Saint-Gilles-les-Bains, between the

harbour and Boucan-Canot beach, in hand-dredged

sand at 30-50 m.

Type material. Holotype and paratype 1 in MNHN,

paratypes 2 to 7 coll. M. Jay: paratype 8 coll. J.

Drivas.

Etymology. Dedicated to Dr. Glaubrecht, MNK,

Berlin.

Remarks. This species differs from Horologica

minareta Laseron, 1956 in its more swollen shape, its

larger beads, the sculpture of base, its darker colour,

and its white instead of brown protoconch: it differs

from Horologica macrocephala Laseron, 1956 in the

smaller size of its protoconch, its larger size (2.3 mm

instead of 1.7 mm), and the sculpture of base. It differs

from Horologica anisocorda n.sp. which is quite

similar, in its more swollen and shorter shape, in the

beads of the 2 cords being equal on earlier whorls

instead of strongly unequal. It differs from Dizoniopsis

gothica n.sp. in its smooth instead of ribbed

protoconch.

Horologica konops n. sp.

Plate 6, E; colour plate II, Fig. 43

Material examined. 2 spmns MNHN; 9 spmns (6

with complete protoconch) coll. M. Jay; 5 spmns

coll. J. Drivas.

Description. Very small shell, elongate, nearly

cylindrical, slightly constricted at base. Protoconch

conical, wide and high, of 5.5 whorls, convex and

smooth, its 2 last whorls swollen; limit from

teleoconch along an oblique line marking the change

of colour and start of adult sculpture; earlier whorls

tapering towards rounded apex. Teleoconch of 5

whorls, 2 beaded spiral cords per whorl, interval

between them wider than the interval at suture; weaker

axial ribs crossing them at right angles, with a rounded

bead at each intersection; beads numbering 17-18 per

whorl, subequal on the 2 cords on early whorls, beads

of upper cord larger and axially elongate on

subsequent whorls. A third beaded cord emerging

from suture at base of last whorl. A fine smooth cord

on base. Aperture rounded, broken on type. Colour

plain cream, protoconch and base a little darker on

fresh specimens.

Size: holotype total height 1.7 mm; maximum width

0.5 mm; height of protoconch 0.43 mm; width of

protoconch at base 0.31 mm.

Type locality. Found dead in hand-dredged sand at 30

m, off Boucan-Canot beach, Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNAN,

paratypes 2 to 7 coll. M. Jay; paratype 8 coll. J.

Drivas.

Etymology. Named from Greek, meaning mosquito,

after its size and slenderness.

Remarks. This species resembles Horologica

macrocephala Laseron, 1956 in its size and the shape

of the protoconch, but differs from it in its more

cylindrical shape, less swollen at the middle and less

constricted at base, its somewhat smaller protoconch

(1/4 of total height instead of 1/3), and its more

numerous beads (18-19 instead of 16). This new

species may be compared to Horologica martini

n.sp. which has a similar protoconch and colour, but

differs from it in being less swollen and of smaller

size (1.7 mm instead of 2.2 mm).

Horologica martini n. sp.

Plate 6, F; colour plate II, Fig. 44

Material examined. 2 spmns MNHN; 58 spmns coll.

M. Jay: 23 spmns coll. J. Drivas: all with complete

protoconch.

Description. Shell fusiform with constricted base,

apex of teleoconch topped by the prominent

protoconch. Protoconch conical of 5 convex smooth

whorls, the last 2 whorls of equal size, their limit from

teleoconch unprecise, marked by the progressive

development of adult sculpture; the earlier whorls

regularly tapering to the rounded apex. Teleoconch of

5-6 whorls, with 2 spiral cords, crossed by slightly

finer axial ribs, with a rounded bead at each

intersection; beads close-set, subequal on earlier

whorls, and numbering 20-21 per whorl. On the 2 last

whorls, beads of upper cord stronger and axially

elongate, then incised by a spiral furrow, then divided

into two beaded cords on last whorl. A fourth beaded

cord emerging from suture at base of last whorl. A

smooth finer spiral cord at mid-height of base,

followed by 4 very fine spiral threads. Towards

aperture, the beads progressively reduced to weak

axial swellings. Aperture rounded. Colour white, base

very pale brown, the lower cord tinted with pale brown

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NOVAPEX 3 (1): 1-45, 10 mars 2002

on the 2 last whorls, the upper cord remaining white;

this colour pattern disappearing on worn specimens.

Size: holotype total height 2.2 mm; maximum width

0.8 mm; protoconch height 0.44 mm; width of

protoconch at base 0.32 mm. Height of specimens

ranging from 2.0 to 2.5 mm.

Type locality. Found dead in hand-dredged sand at

30m off Boucan-Canot beach, Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN,

paratypes 2 to 15 coll. M. Jay; paratypes 16 to 20

coll. J. Drivas.

Etymology. Dedicated to J.C. Martin, skin diver and

collector, who collected the sand in which the first

specimen was found.

Remarks. This species has the same outline as

Horologica turrigera Watson, 1886, of which it

could be considered as a dwarf form (2.2 mm instead

of 3.7mm), but no intermediate exists between the 2

forms, even though many specimens of the 2 species

could be found. This species differs from

Horologica balteata Watson, 1886, which has a

similar colour pattern, in its smaller size (2.2 mm

instead of 3 mm), less rounded shape, and its less

neatly marked colour pattern. It differs from

Horologica macrocephala Laseron, 1956, in its

larger size (2.2 mm instead of 1.7 mm), in the

proportionally smaller size of its protoconch, and its

colour pattern. Horologica konops n.sp. which has a

comparable protoconch, is less swollen and smaller

(1.7 mm instead of 2.2 mm).

Genus Dizoniopsis Sacco, 1895

Type species: Cerithium bilineatum Hôrnes, 1855,

type locality Tertiary banks of Piémont: teleoconch

with 2 beaded spiral cords per whorl, protoconch

bearing axial ribs, extending from suture to suture,

with smooth intervals (Dizoniopsis ss). We

provisonally consider as Dizoniopsis (sl) the species

with 2 spiral beaded cords per teleoconch whorl, with

protoconchs bearing spiral cords in intervals betwen

the ribs (type 3) or with axial ribs that do not reach the

upper suture (type 4).

Dizoniopsis gothica n.sp.

Plate 9, C; colour plate II, Fig. 45

Material examined. 2 spmns (1 with complete

protoconch) MNHN: 7 spmns (1 with complete

protoconch) coll. M. Jay; 2 spmns (1 with complete

potoconch) coll. J. Drivas.

Description. Teleoconch pupiform, with pointed apex,

topped by the prominent protoconch, general outline

recalling that of a gothic tower; apical angle of

teleoconch 55°; maximum width at the penultimate

whorl; base slightly constricted. Protoconch high,

nearly cylindrical, consisting in 4.5 convex whorls; its

earliest 1.5 whorls smooth, the subsequent whorls with

their upper 1/3 concave, smooth and finely frosted,

their lower 2/3 convex and bearing slightly prosocline

axial ribs (ribbed protoconch of type 4). Teleoconch of

6 whorls, rather low, suture moderately impressed; 2

spiral cords per whorl, crossed at right angles by axial

ribs weaker than cords, reaching upper and lower

suture, but discontinuous from one whorl to another; a

rounded bead at each intersection, beads rather small

and widely-spaced, numbering 20-21 per whorl. On

last whorls, beads of upper cord progressively axially

elongate, incised on last whorl and divided near

aperture by a spiral furrow. A fourth spiral cord

emerging from suture at base of constricted last whorl,

finer and bearing finer beads. Beads of the 4 cords

becoming smaller towards outer edge of aperture, axial

ribs becoming predominant. A fifth fine, smooth,

unbeaded cord on base. Aperture roundly quadrate.

Colour blackish-brown, protoconch and the first 2

whorls of teleoconch clearly paler or creamy-white.

Size: holotype total height 2.7 mm; maximum width

1.2 mm; height of protoconch 0.42 mm; maximum

diameter of protoconch 0.33 mm.

Type Locality. Found dead in hand-dredged sand at

10-20 m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 (without

protoconch) in MNHN; paratype 2 coll. M. Jay:

paratype 3 coll. J. Drivas.

Etymology. Named after the general shape and the

prominent protoconch, recalling gothic churches.

Remarks. This species differs from Dizoniopsis

herosae n.sp. in its more swollen shape, and its

protoconch, the protoconch of D. herosae n.sp. being

shorter with axial ribs extending from suture to suture.

D. herberti n.sp. which has à similar protoconch of

type 4, is easily separated by its colour pattern.

Dizoniopsis herberti n.sp.

Plate 9, B; colour plate II, Fig. 46

Material examined. 2 spmns MNHN; 10 spmns (3

with complete protoconch) coll. M. Jay.

Description. Shell pupiform with constricted base,

and pointed apex surmonted by the prominent

protoconch. Protoconch of 4.5 convex whorls, the first

1.5 smooth, the following ones with axial riblets,

numbering 18-19 per whorl, slightly prosocline,

present only on the lower 3/4 of whorls, leaving

smooth and concave the upper 1/4 (ribbed protoconch

of type 4). Teleoconch of 6 whorls, with 2 beaded

spiral cords per whorl, crossed at right angles by

distinct flattened axial ribs, with a rounded bead at

each intersection; beads numbering 19 or 20 per

whorl; beads of lower cord slightly stronger and

Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

PLATE 6. Fig. A. Horologica macrocephala Laseron, 1956. Off Saint Gilles les Bains, 20-30 m; height 1.7

mm; MNAN. Fig. B. H. minareta Laseron, 1956. Off Saint Gilles les Bains, 10-20 m; height 2.6 mm; MNHN.

Fig. C. Æ. anisocorda n.sp. Off Saint Gilles les Bains, 30m; holotype, height 2.2 mm; MNEHEN.

Fig. D. Æ. glaubrechti n.sp. Off Saint Gilles les Bains, 30-50 m; holotype, height 2.3 mm; MNEN.

Fig. E. Æ. konops n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 30 m; holotype, height 3ämm; MNAN.

Fig. F. A. martinin.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 30 m; holotype, height 2.7 mm;

MNAN.

Scale bars: S (shells): 1 mm; P (protoconchs): 100 um.

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axially elongate. On the Sth whorl, a third finely

beaded spiral cord develops from a fine thread

between the 2 earlier cords, and becomes comparable

to them but slightly weaker on last whorl; a fourth

weaker beaded spiral cord emerging from suture at

base of last whorl, the beads of all cords becoming

smaller towards outer edge of aperture. A fifth

flattened smooth cord on base. Aperture circular.

Colour cream to pale orange-beige, the upper cord and

its beads reddish-brown, the median cord on last whorl

between these two colours. Protoconch brown.

Size: total height 2.4 mm; maximum width 0.9 mm;

height of protoconch 0.4 mm; width at base of

protoconch 0.3 mm.

Type locality. Found dead in hand-dredged sand at

10-20 m off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN;

paratypes 2 to 4 coll. M. Jay.

Etymology. Dedicated to Dai Herbert, Natal Museum.

Remarks. The teleoconch of our species resembles a

specimen labelled Cerithiopsis insignis E.A. Smith,

1906, from Southern Africa, in NHM (lot Nr.

1906.6.23.10, type material, syntype?); this specimen

has no protoconch, 3 beaded spiral cords only on last

whorl, and 2 cords only on earlier whorls. But Smith

described the species (type locality Port Shepstone,

South Africa) as bearing 3 beaded spiral cords, the

median one weaker, and his figure shows 3 subequal

spiral cords from the first whorl of teleoconch.

Topotypic specimens from Port Shepstone, in Natal

Museum, have no protoconch and 3 beaded spiral

cords on all whorls of teleoconch, equal, or the median

cord slightly weaker on a few specimens; other

specimens from Durban area, similar to the specimens

from Port Shepston, have à smooth 4,5 whorled

protoconch (personal communication from D. Herbert,

Natal Museum): these specimens match the

description and figure of Smith, but differ from the

type material in NHM in the number of spiral cords on

the teleoconch. We think that these specimens are the

real Cerithiopsis insignis, and that the syntype in

NHM is another species. Our specimens with the

costate protoconch are also another species, for which

we propose the name Dizoniopsis herberti.

Furthermore, Dizoniopsis herberti n.sp. differs from

other bicolored species, namely Horologica bicolor

(Laseron, 1955), Horologica semipicta (Gould, 1861),

Joculator pulvis Issel, 1869, Joculator eudeli n.sp., in

its axially ribbed protoconch (instead of smooth). This

species is referred to the genus Dizoniopsis on account

of its ribbed protoconch and its 2 spiral beaded cords

on the earlier whorls of the teleoconch.

Dizoniopsis herosae n.sp.

Plate 9, A; colour plate II, Fig. 47

Material examined. 2 spmns MNHN; 32 spmns (12

with complete protoconch) coll. M. Jay; 11 spmns

coll. J. Drivas.

Description. Shell small and fusiform, strongly

constricted at base, apical angle 25°, the greatest width

at the lower third of total height. Protoconch

elongating the general outline of teleoconch,

consisting in 2.5 convex whorls, well delimited from

teleoconch by the clear-cut change of sculpture; apex

rounded and smooth, subsequent whorls with strong

axial ribs, extending from suture to suture, numbering

15-16 on last whorl, their intervals smooth.

Teleoconch of 8 whorls, suture shallow. 2 beaded

spiral cords per whorl, situated near upper and lower

sutures, With a wide interval; 18 to 20 axial ribs per

whorl, wide and low, crossing the cords at right

angles, with a rounded bead at each intersecton. A

third beaded spiral cord on last whorl, developed from

a fine thread between the 2 main cords, and remaining

weaker than them. A fourth weak, unbeaded spiral

cord emerging from suture at base of last whorl,

followed immediately by a fifth spiral cord equal to it.

Beads becoming smaller towards aperture. Aperture

circular. Colour plain pale brown, suture a little darker,

protoconch and the 3 earlier whorls of teleoconch

creamy-white.

Size: holotype total height 3.3 mm; maximum width

1.3 mm; height of protoconch 0.49 mm; width of

protoconch at base 0.36 mm.

Type locality. Found dead in hand-dredged sand at 30

m, Souris-Chaude, Trois-Bassins.

Type material. Holotype and paratype 1 in MNEN,

paratypes 2 to 12 coll. M. Jay; paratypes 13 to 15

coll. J. Drivas.

Etymology. Dedicated to Mrs Virginie Heros,

MNHN.

Remarks. This species could be confused, in its shape

and colour, with Joculator keratochroma n. sp., but

differs from that in its larger size, its 2 cords instead of

3, and its ribbed protoconch instead of smooth. It

differs from Horologica turrigera (Watson, 1886) and

Horologica martini n.sp. in its ribbed protoconch. It

differs from Dizoniopsis gothica n.sp. in its colour

(pale brown instead of black), and its ribbed

protoconch of 2.5 whorls instead of 4.5. It differs

from other species of Mendax and Prolixodens in its 2

beaded cords on teleoconch whorls instead of3.

Genus Mendax Finlay, 1927

Type species Cerithiopsis trizonalis Odhner, 1924,

North Island, New Zealand: spire high, teleoconch

with 3 beaded spiral cords per whorl. Protoconch with

axial ribs extending from suture to suture. 2 species

with spiral cords in the intervals between protoconch

ribs are provisionally attributed to the genus.

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Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Mendax mascarenensis n.sp.

Plate 8, C; colour plate II, Fig. 48

Material examined. 2 spmns MNHN:; 4 spmns (3

with complete protoconch) coll. M. Jay; 3 spmns

coll. J. Drivas.

Description. Shell with high spire, slightly fusiform,

base not constricted. Protoconch rather cylindrical of 4

convex whorls, the earlier 1.5 appearing smooth by the

naked eve, but bearing fine close-set granules under

SEM, the subsequent whorls with axial riblets,

extending from suture to suture, weak but clearly

visible under oblique light; their intervals smooth.

Limit from teleoconch oblique marked by change of

colour and start of adult sculpture. Teleoconch of 9

whorls, with 3 beaded cords per whorl, equal on last

whorl, but upper cord smaller and contracted on all

preceeding whorls. Intervals between cords rather

wide, with distinct axial ribs joining the beads at right

angles. A fourth weaker cord emerging from suture at

base of last whorl. Aperture rather quadrangular.

Colour plain bright orange-brown, protoconch white.

Size: total height 4.2 mm; maximum width 1.1 mm;

height of protoconch 0.52 mm; width at its base 0.33

mm.

Type locality. Found dead in hand-dredged sand at 30

m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN;:

paratypes 2 to 4 coll. M.Jay: paratype 5 coll. J.

Drivas.

Etymology. Named after Mascarene Isl. since Melvill

knew this species from Mauritius.

Remarks. Our specimens were compared with the

specimens of a lot labelled Cerithiopsis aurantiaca

Melvill & Standen, 1896, in NMW ( lot N°

Z.1955.158.02268) from Mauritius: this lot includes 4

specimens, only one of them with its axially ribbed

protoconch, and quite identical to ours; 2 other

specimens With identical teleoconch but without

protoconch: the fourth specimen, with only the last

whorl of a smooth protoconch, and the upper spiral

cord of teleoconch weaker and darker, is a quite

different species. Another lot in NMW from Lifu (lot

N° Z.1955.158.00189), labelled syntypes of

Cerithiopsis aurantiaca Melvill & Standen, 1896,

includes 2 specimens without protoconch, which look

identical to ours after their teleoconchs, but could be

attributed to any of our 9 species discussed above with

Cerithiopsis boucheti n.sp. But the holotype of

Cerithiopsis aurantiaca Melvill & Standen, 1896, type

locality Loyalty Isl. in MM has no protoconch left,

and a teleoconch obviously wider than our specimens

and specimens in NMW. The specimens labelled

syntypes in NMW are then another species, difficult to

identify without protoconch. Our specimens, identical

to the specimens from Mauritius in NMW, belong

certainely to another species, which seems

undescribed, and for which we propose the name of

Mendax mascarenensis.

Besides, the name Cerithiopsis aurantiaca attributed

by Melvill & Standen to the holotype in MM, is

preoccupied by Cerithiopsis aurantiaca Gould, 1861,

which is a quite different species. For the species of

Melvill and Standen, we propose here a new name,

Cerithiopsis melvilli nom. nov. in honour of J.C.

Melvill; but without a protoconch, assigning it to a

genus is quite impossible and it will provisionnally be

left in the genus Cerithiopsis.

Mendax metivieri n. sp.

Plate 8, B; colour plate II, Fig. 49

Material examined. 2 spmns MNHN; 105 spmns

Coll. M. Jay; 20 spmns coll. J. Drivas; all with

complete protoconch.

Description. Shell fusiform, protoconch extending

general outline of teleoconch, last whorl not

constricted; all whorls strongly convex giving the shell

a characteristic profile. Protoconch of 2.5 whorls, with

axial ribs extending from suture to suture, numbering

18-19 per whorl, their intervals with much finer, close-

set spiral cords (ribbed protoconch of type 3); limit

from teleoconch progressive along 1/4 whorl.

Teleoconch of 9 convex whorls, suture deeply

impressed, 3 spiral cords per whorl, crossed at right

angles by axial ribs, with a big rounded bead at each

intersection; beads numbering 19 or 20 on penultimate

whorl. The upper spiral cord hardly visible on first

whorl, very weak on the 2 following whorls, but equal

to the other ones on last whorls. Last whorl with a

fourth spiral cord emerging from suture, weaker than

the other ones, and underlining angle with the

excavated base; a fifth spiral cord, weak and

unbeaded, at the upper part of base. On last whorls,

suture marked by a fine spiral thread. Aperture

trapezoidal with its upper and outer angle acute.

Colour plain golden brown, with a reddish gleam on

fresh specimens; protoconch brown or white.

Size: maximum height of shell 4.3 mm; width at base

1 mm; height of protoconch 0.40 mm; width at its base

0.35 mm.

Type locality. Found dead in hand-dredged sand at

30-50 m, off Saint-Gilles-les-Bains, between harbour

and Boucan-Canot beach.

Type Material. Holotype and paratype 1 in MNEN,

paratypes 2 to 20 coll. M. Jay; paratype 21 to 25

coll. J. Drivas.

Etymology. Dedicated to Mr. Bernard Metivier,

MNEHN.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Remarks. This species recalls Mendax ribesae n. sp.

in its shape and colour, but differs in that its

protoconch bears spiral cords.

Mendax penneyi n.sp.

Plate 8, A; colour plate II, Fig. 50

Material examined. 1! spmn MNHN; 4 spmns (1

with complete protoconch) coll. M. Jay; 2 spmns

coll. J. Drivas.

Description. Shell fusiform, with moderately

constricted base, apex surmonted by the prominent

cylindrical protoconch. Protoconch of 4.5 whorls, the

earlier 1.5 smooth with rounded apex, the following

ones with axial ribs extending from suture to suture,

their intervals smooth (ribbed protoconch of type 1);

limit from teleoconch oblique, marked by change of

colour, but adult sculpture develops progressively on

last half-whorl. Teleoconch of 7 whorls, 3 spiral cords

per whorl, crossed at right angles by a little weaker

axial ribs, with a rounded bead at each intersection;

beads numbering 18 or 19 per whorl, the 3 spiral cords

subequal. À fourth unbeaded spiral cord emerging

from suture at base of last whorl. A strong smooth

cord at mid-height of base. Colour white, the upper

beaded cord very pale brown on fresh specimens.

Size: total height 4.1 mm; maximum width 1.4 mm;

height of protoconch 0.58 mm; width of protoconch at

base 0.32 mm.

Type locality. Found dead in hand-dredged sand at

10-20 m, off Saint-Gilles-les-Bains.

Type material. Holotype in MNHN, paratype 1 coll.

M. Jay.

Etymology. Dedicated to Dr. David Penney, assistant

keeper of Zoology, Manchester Museum.

Remarks. The teleoconch of this species is similar to

that of Cerithiopsis hedista Melvill & Standen, 1896

(type locality Lovyalty Is.) in its size, sculpture and

colour pattern; but the holotype of Cerithiopsis hedista

in MM (lot EE 3743) has 4 whorls of protoconch left,

with a broken summit, these 4 whorls slightly convex

and smooth; this character is quite distinctive from our

species. The colour pattern of Mendax penneyi n.sp.

recalls that of Cerithiopsis eutrapela Melvill &

Standen, 1896, from which it is easily separated by its

smaller size, its 3 equal cords, and ribbed protoconch.

Mendax ribesae n.sp.

Plate 8, D; colour plate II, Fig. 51

Material examined. 2 spmns MNHN:; 1! spmn coll.

M. Jay; 1 spmn coll. J. Drivas; all with complete

protoconch.

Description. Shell small and slender, fusiform,

slightly constricted at base. Protoconch conical of 4.5

whorls, slightly convex, apex rounded, the first whorl

smooth, the following 3 with fine axial ribs, extending

from suture to suture, numbering 20 per whorl, their

intervals smooth; suture with very fine close-set axial

threads (ribbed protoconch of type 2); limit from

teleoconch oblique, marked by change of sculpture

and colour. Teleoconch of 5 slightly convex whorls

and impressed suture. 3 beaded spiral cords per whorl,

the uppermost one weaker and recessed except on last

whorl where the 3 cords are subequal; finer axial ribs,

crossing them at right angles, with one bead at each

intersection, beads numbering 20-21 per whorl. A

fourth spiral cord weaker, and bearing finer beads

emerging from suture at base of last whorl. One more

fine smooth cord at mid-height of the excavated base.

Aperture rounded. Colour pale orange-brown, base

slightly darker, protoconch white.

Size: holotype total height 2.6 mm; maximum width

0.8 mm; height of protoconch 0.5 mm; width of

protoconch at base 0.4 mm.

Type locality. Found dead in hand-dredged sand at 55

m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNAN,

paratype 2 coll. M. Jay; paratype 3 coll. J. Drivas.

Etymology. Dedicated to Mrs Sonia Ribes, curator of

the Natural history Museum of Saint Denis, La

Réunion.

Remarks. This species has the same size as Mendax

theodosiae n.sp. but differs from it in its narrower

width, weak and recessed upper cord, and colour. The

other species of Mendax collected on Reunion are

much larger. This species is easily separated from

Joculator species of the same size, by its ribbed

protoconch.

Mendax theodosiae n.sp.

Plate 8, E; colour plate II, Fig. 52

Material examined. 2 spmns MNHN; 9 spmns coll.

M. Jay; 3 spmns coll. J. Drivas.

Description. Shell pupiform with strongly constricted

base, more slender towards apex, topped by the

prominent and pointed protoconch, its maximum

diameter at the lower 1/3 of teleoconch. Protoconch of

4.5 convex whorls, the earlier 1.5 whorl with rounded

apex, looking smooth but showing under SEM, a

narrow spiral row of close-set axial riblets, numbering

about 50 per whorl, situated in suture and extending

shortly on the upper part of lower whorl: following

whorls bearing strong axial ribs numbering 18-20 per

whorl, with the same very narrow row of axial riblets

in and under suture (ribbed protoconch of type 2);

limit from teleoconch oblique marked by change of

colour, but adult sculpture develops progressively on

the last half-whorl. Teleoconch of 6 slightly convex

whorls, suture shallow. 3 spiral cords per whorl,

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

except on the 2 earlier whorls where there are only 2

cords, the other one originating on the second whorl,

between the 2 earlier cords, becoming equal to the

other ones on the third whorl. Axial ribs weaker than

cords, crossing them at right angles, and numbering 18

on penultimate whorl, with a rounded bead at each

intersection. Last whorl strongly constricted at base, a

fourth weakly beaded spiral cord emerging from

suture at its base. On the last half-whorl, beads become

weaker and more axially elongate towards the outer lip

of aperture. A fifth spiral cord, smooth, on base.

Colour dark brown to golden brown, the earlier whorls

of teleoconch paler, protoconch white.

Size: holotype total height 3.3 mm; maximum width

1.2 mm; height of protoconch 0.73 mm; width of

protoconch at base 0.42 mm.

Type locality. Found dead in hand-dredged sand from

10-20 m., off Saint-Gilles-les-Bains, between harbour

and Boucan-Canot beach.

Type material. Holotype and paratype 1 MNHN,

paratypes 2 to 7 coll. M. Jay; paratype 8 coll. J.

Drivas.

Etymology. Dedicated to Mrs Theodosia Drivas.

Remarks. This species differs from other Mendax

species in its characteristic protoconch, bearing under

suture the narrow row of fine axial riblets; besides, it

differs from Mendax ribesae n.sp. which has a similar

colour, in its shorter and more compact shape, and its

size (2.7 mm instead of 4.1mm); it differs from

Mendax metivieri n.sp. which has the same colour, in

its shorter and more compact shape and its size (2.7

mm instead of 4.5 mm). It is easily separated from

Prolixodens sknibs n.sp. in its higher and differently

sculptured protoconch (protoconch of type 2 instead of

type 4), and its colour less blackish. /oculator lozoueti

n.sp., which has a similar shape, is somewhat larger

and has a smooth protoconch.

Genus Prolixodens Marshall, 1978

Type species: Cerithiopsis infracolor Laseron, 1951,

Long Reef, NSW, Australia: teleoconch with 3

beaded spiral cords per whorl, protoconch with axial

ribs on the lower 2/3 of whorls, the upper 1/3 smooth

or punctate (ribbed protoconch of type 4).

Prolixodens nicolayae n. sp.

Plate 8, F; colour plate II, Fig. 53

Material examined. 2 spmns MNHN; ! spmn coll

M. Jay: 1! spmn coll. J. Drivas; all with complete

protoconch.

Description. Shell very small, fusiform, with

constricted base, and relatively high protoconch.

Protoconch conical with pointed apex, comprising 4

strongly convex whorls, the first one slightly granular,

the following ones with axial ribs on the lower 2/3 of

whorls, the upper 1/3 slightly concave and granulous;

no spiral sculpture on protoconch; limit from

teleoconch clear-cut and oblique, marked by change of

colour and sculpture. Teleoconch of 4 whorls, bearing

3 beaded spiral cords per whorl, the upper one weaker

and recessed, becoming nearly equal to the other ones

on last whorl; weaker axial ribs crossing them at right

angles, with a rounded bead at each intersection; beads

numbering 16 per whorl. A fourth spiral cord, finely

beaded, emerging from suture at base of last whorl; a

fifth smooth spiral cord on base. Colour creamy-white,

protoconch more milky white.

Size: holotype total height 1.4 mm; maximum width

0.6 mm; height of protoconch 0.40 mm; width of

protoconch at base 0.29 mm.

Type locality. Found dead in hand-dredged sand at

30m, off Souris-Chaude, Trois-Bassins.

Type material. Holotype and paratype 1 in MNHN,

paratype 2 coll. M. Jay; paratype 3 coll. J. Drivas.

Etymology. Dedicated to Mrs K. Nicolay, foundress

of “La Conchiglia”.

Remarks. This species differs from Prolixodens

sknips n.sp. the protoconch of which is of similar type,

in its less swollen shape, smaller size (1.4 mm instead

of 1.8 mm) and colour (cream instead of blackish-

brown); it differs from Dizoniopsis gothica n.sp. and

from Dizoniopsis herberti n.sp. the protoconch of

which is also of similar type, in its teleoconch with 3

beaded spiral cords instead of 2, its shape and colour.

Prolixodens sknips n.sp.

Plate 8, G; colour plate II, Fig. 54

Material examined. 2 spmns MNHN; 7 spmns (1

with complete protoconch) coll. M. Jay; 2 spmns

coll. J. Drivas.

Description. Shell pupiform, strongly swollen, the

maximum diameter at the middle of teleoconch, base

strongly constricted, and apex topped by the prominent

protoconch. Protoconch of 3.5 whorls, the first one

rounded and finely granular, the following ones with

axial ribs numbering 20-21 per whorl, appearing only

on the lower 2/3 of whorls, the upper 1/3 being

concave and finely granular; limit from teleoconch

axial, marked by change of colour and development of

spiral sculpture. Teleoconch of 6 slightly convex

whorls, suture shallow. 3 spiral cords per whorl,

crossed at right angles by weaker axial ribs, numbering

16 on penultimate whorl. A strong rounded bead at

each inersection, beads more close-set axially than

spirally; the 3 cords subequal, in spite of the narrowing

of both ends of shell. A fourth spiral cord emerging

from suture at base of last whorl, with only weak

swellings; a fifth unbeaded smooth cord on base, and

JAY & DRIVAS Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

PLATE 7. Fig. A. Horologica balteata (Watson, 1886). Off Saint Gilles les Bains, 12m; height 3 mm; MNAN.

Fig. B. Æ. bicolor Laseron,1956. Off Saint Gilles les Bains, 30 m; height 2.1 mm; MNAN. Fig. C. Æ. cf

semipicta (Gould, 1861). Off Saint Gilles les Bains, 30 m; height 2.1 mm; MNAN. Fig. D. F. purpurea

Laseron, 1955. Off Cape La Houssaye, Saint Gilles les Bains, 10-20 m; height 3.1 mm; MNHN.

Fig. E. Æ. turrigera (Watson, 1886). Off Saint Gilles les Bains, 10-20 m; height 3.7 mm; MNAN.

Fig. F. Belonimorphis belonimorphis n.sp. Off Saint Gilles les Bains, 10-30 m; holotype, height 6 mm:

MNAN. Fig. G. Koïlofera koilofera n.sp. Off Saint Gilles les Bains, 10-30 m; holotype, height 2.7 mm:

MNEHN.

Scale bars: S (shells): 1 mm; P (protoconchs): 100 um.

JAY & DRIVAS

below it, several fine spiral threads visible under

microscope. Aperture circular. Colour plain dark

brown to blackish-brown, with reddish gleam on fresh

specimens, protoconch paler brown with white apex.

Size: holotype total height 1.8 mm; maximum width

0.9 mm; height of protoconch 0.25 mm; width of

protoconch at base 0.25 mm.

Type locality. Found dead in hand-dredged sand at

10-30 m, off Saint-Gilles-les-Bains, between harbour

and Boucan-Canot beach.

Type material. Holotype and paratype 1 in MNHN,

paratypes 2 coll. M. Jay: paratype 3 coll. J. Drivas.

Etymology. Named for its size after Greek, meaning

small fly.

Remarks. This species differs from Prolixodens

nicolavae n. sp. in its size (1.8 mm instead of 1.4 mm),

shape, equal spiral cords instead of unequal, and

shorter protoconch (3.5 whorls instead of 4.5). It

resembles Dizoniopsis gothica n. sp. in the sculpture

of the protoconch, but differs from that species in its 3

beaded cords on teleoconch whorls instead of 2: it is

easily separated from Koïlofera koilofera n.sp. and

from small species of Joculator in its ribbed

protoconch. The Prolixodens species of Marshall

(1978) are much larger.

Genus Belonimorphis n. gen.

Teleoconch conical elevated and narrow with 3 spiral

cords. Protoconch high, cylindrical, each whorl convex

with 2 spiral keels, and without axial ribs. Type

species: Belonimorphis belonimorphis n.sp.

Belonimorphis belonimorphis n.sp.

Plate 7, F; colour plate II, Fig. 55

Material examined. 2 spmns MNHN; 200 spmns

(100 with complete protoconch) coll. M. Jay; 50

spmns coll. J. Drivas.

Description. Shell fusiform, high and slender, tapering

regularly to the apex of teleoconch, protoconch more

elevated and pointed. Protoconch of 3.5 whorls, nearly

equal in diameter, and thus cylindrical; each whorl

strongly convex bearing 2 prominent and well

separated spiral Kkeels, without any other sculpture;

limit from teleoconch ill-defined, marked by the

development of axial ribs. Teleoconch of 11 or 12

whorls with fine, prominent, widely spaced spiral

cords, numbering 2 on earlier whorls, 3 on the

following ones, the new cord originating immediately

under the upper suture; this upper cord remaining

weaker except on last whorl. Rounded axial ribs, a

little weaker than the cords, crossing them at right

angles, and numbering 18-19 per whorl; each

intersection with a small bead, more or less rounded,

sometimes reduced to a faint swelling. Numerous

close-set axial riblets well visible under microscope,

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

on the whole surface including spiral cords, axial ribs

and their intervals, numbering 18 or 19 on each axial

rib and the near interval, totalling 400 per whorl. A

fourth spiral cord emerging from suture at base of last

whorl, as strong as the other cords, but bearing only

faint swellings rather than beads. A fifth spiral cord,

smooth and unbeaded, at mid-height of base. Aperture

circular, anterior canal markedly oblique. Colour dark

blackish brown to golden brown, the 3 earlier whorls

of teleoconch and protoconch white.

Size: holotype total height 6 mm; width at base 1.5

mm; height of protoconch 0.74 mm; maximum width

of protoconch 0.40 mm.

Type locality. Found in hand-dredged sand at 10-20 m

off Saint-Gilles-les-Bains, between Hermitage and St

Paul Bay.

Type material. Holotype and paratype 1 in MNEN,

paratypes 2 to 20 coll. M. Jay; paratypes 21 to 30

coll. J. Drivas.

Etymology. Named after its shape, from Greek

meaning needle-shaped.

Remarks. This species is easily dinstinguished from

other species of Cerithiopsidae by its slender shape,

and by its protoconch bearing 2 spiral keels. Species of

the genus Cerithiopsis with a slender shape have a

smooth protoconch. Species of the genus Mendax have

an axially ribbed protoconch. Some species with a

similar shape and similar teleoconch sculpture have

been described by Laseron (1956) in the genus

Cerithiopsis, and by Marshall (1978) in the genus

Laskeya, but their protoconchs are very different. A

protoconch with a similar sculpture has been

illustrated by Marshall (1978) in the family

Triphoridae (/nella gigas) but with a sinistral coiling.

This species seems to have some variable characters:

its colour ranging from blackish brown to golden

brown; the size of its teleoconch beads, (either well

rounded on some specimens or obsolete on other ones,

the sculpture being thus reduced to a cancellate

pattern); and its size, with specimens smaller and

narrower though in their adult state; however all these

specimens have identical protoconchs, and many

intermediates exist between these variations.

Genus Koilofera n. gen.

Teleoconch pupiform with 2 beaded spiral cords per

whorl as Horologica, but protoconch with a strong

spiral swelling just above suture, and a strong

concavity above this, without any other sculpture.

Named from Greek meaning “bearing a concavity”.

Type species: Koilofera koilofera n.sp.

Koilofera koilofera n.sp.

Plate 7, G; colour plate II, Fig. 56

Material examined. 2 spmns MNHN; 2 spmns coll.

M. Jay; 1 spmn coll. J. Drivas; all with complete

protoconch.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Description. Shell pupiform with strongly constricted

base, the maximum width just under mid-height.

Protoconch of 2.5 whorls, with a flat apex, delimited

by a well marked angle; whorls of protoconch with a

very strong rounded spiral swelling, situated just

above lower suture, its upper part appearing very

slightly beaded under oblique light on fresh

specimens; whorl strongly concave above this

swelling, without any other sculpture either axial or

spiral; limit from teleoconch clear-cut and oblique,

marked by change of colour and development of adult

sculpture. Teleoconch of 6 slightly convex whorls,

shallow suture. 2 spiral cords per whorl, crossed at

right angles by fine axial ribs, with at each intersection

a slightly axially elongate bead; beads numbering 17-

18 per whorl. On the fourth and fifth whorls, beads

become larger on the upper cord than on the lower

one; on last whorl, these upper beads are more axially

elongate, and partially incised in their middle by a

spiral furrow, but are never quite divided on our

specimens. Last whorl strongly constricted at base,

with a third spiral cord, unbeaded, emerging from

suture; a fourth unbeaded spiral cord on base. Aperture

circular. Colour very dark brown to blackish-brown,

protoconch distinctly paler.

Size: holotype total height 1.7 mm; maximum width

0.6 mm, height of protoconch 0.29 mm; width of

protoconch at base 0.36 mm.

Type locality. Found dead in hand-dredged sand at

10-30 m, off Saint-Gilles-les-Bains, between harbour

and Boucan-Canot beach.

Type material. Holotype and paratype 1 in MNHN;:

paratype 2 and 3 coll. M. Jay; paratype 4 coll. J.

Drivas.

Etymology. Named for the shape of protoconch

whorls, from Greek meaning “bearing a concavity”.

Remarks. In its shape and colour, this species

resembles Dizoniopsis gothica n.sp. but is easily

separated from it by its protoconch. Some species with

a more or less similar protoconch have been described

by Marshall (1978) in the genus Sei/a, but these have a

quite different adult sculpture.

Genus Seila A. Adams, 1861

Type species Triphoris dextroversus Adams &

Reeve, 1860, China seas. Shell high and slender,

teleoconch with 3 or more smooth spiral cords per

whorl and fine close-set axial lamellae in their

intervals. Protoconch smooth or axially ribbed.

Seila bandorensis (Melvill, 1893).

Plate 9, D; colour plate II, Fig. 57

Material examined. ! spmn MNHN; 19 spmns coll.

M. Jay; 5 spmns coll. J. Drivas.

Descripton. Shell elongate pyriform; base strongly

constricted. Rounded protoconch of 1.5 smooth

convex translucent whorls; limit from teleoconch ill-

defined on 1/4 whorl, marked by development of the 3

spiral cords. Teleoconch whorls with 3 smooth spiral

cords, equal in strength on last whorl, the median cord

weaker on earlier whorls. Very fine lamellose axial

threads in their intervals, well distinct under

microscope. À fourth smooth, weaker spiral cord

emerging from suture at base of last whorl, followed

immediately below it by a fifth cord, and a sixth one at

mid-height of base. Colour plain orange-brown, the 3

earlier whorls paler and whitish, protoconch white.

Size: maximum total height 3.8 mm; maximum width

1.2 mm; height of protoconch 0.30 mm; width of

protoconch at base 0.33 mm.

Locality. Found dead in hand-dredged sand at 10-20

m, off Saint-Gilles-les-Bains.

Remarks. Our specimens were compared and found

identical to the holotype of Cerithiopsis (Seila)

bandorensis Melvill, 1893, type locality Bombay

(NHM lot Nr. 1893.2.16.7).

Seila hinduorum (Melvill, 1898)

Plate 9, E; colour plate II Fig. 58

Material examined. 1 spmn MNHN; 3 spmns coll.

M. Jay; 2 spmns coll. J. Drivas.

Description. Shell conical, slightly fusiform, elongate,

base not constricted. Protoconch of 3 smooth convex

whorls; limit from teleoconch ill-defined on 1/4 whorl,

marked by development of the 3 adult cords.

Teleoconch up to 14 whorls, suture weakly impressed,

whorls with 3 smooth spiral cords, subequal on the

whole height of the shell. Fine very close-set lamellose

axial threads in their intervals. A fourth much weaker

spiral cord emerging from suture at base of last whorl.

Base flat and smooth. Colour plain orange brown to

golden brown; summit paler to white, protoconch

white.

Size: total height 5.5 mm; maximum width 1.2 mm:

height of protoconch 0.58 mm; width of protoconch at

base 0.38 mm.

Locality. Found dead in hand-dredged sand at 40-50

m,; off Saint-Gilles-les-Bains.

Remarks. Our specimens were compared and found

identical to the specimens labelled syntypes of

Cerithiopsis (Seila) hinduorum Melvill, 1898, type

locality Karachi, in NMW (lot Nr Z.1955.158.00206),

18 specimens.

LU)

JAY & DRIVAS Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

PLATE 8. Fig. A. Mendax penneyi n.sp. Off Saint Gilles les Bains, 10-20 m; holotype, height 4.1 mm; MNHN.

Fig. B. M. metivieri n.sp. Off Saint Gilles les Bains, 30-50 m; holotype, height 4.3 mm; MNEN.

Fig. C. M. mascarenensis n.sp. Off Saint Gilles les Bains, 30 m; holotype, height 4.2 m;, MNHN.

Fig. D. M. ribesae n.sp. Off Saint Gilles les Bains, 55 m; holotype, height 2.6 m, MNAHN. Fig. E. M. theodosiae

n.sp. Off Saint Gilles les Bains, 10-20 m; holotype, height 3.3mm; MNHN. Fig. F. Prolixodens nicolayae n.sp.

Off Souris-Chaude, Trois-Bassins, 30 m; holotype, height 1.4 mm; MNAN. Fig. G. P. sknips n.sp. Off Saint

Gilles les Bains, 10-30 m; holotype, height 1.8 mm; MNHN.

Scale bars: S (shells): 1 mm; P (protoconchs): 100 um.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

Seila chenui n.sp.

Plate 9, F: colour plate II, Fig. 59

Material examined. 2 spmns MNHN; 16 spmns (10

with complete protoconch) coll. M. Jay; 5 spmns

coll. J. Drivas.

Description. Shell elongate conical slightly fusiform,

base not constricted, protoconch a little more

prominent than the general outline of teleoconch.

Protoconch conical of 6 whorls, all of them since the

first one bearing axial ribs, numbering 16 or 17 per

whorl, strictly axial on earlier whorls, then more and

more prosocline on following whorls; the 3 last

whorls of protoconch bearing 3 very weak spiral

cords in lower half of whorls, in the intervals

between ribs; limit from teleoconch clear-cut, marked

by change of colour and start of 2 adult cords.

Teleoconch of 13 or 14 whorls with straight sides:

the 5 earlier whorls with 2 spiral cords per whorl; a

third spiral cord beginning at the sixth whorl,

originating from a fine spiral thread in the interval

between the 2 cords; a fourth cord beginning at the

twelfth whorl, originating between the second and

the third cord, but remaining obviously weaker up to

the last whorl; 4 unequal cords on last whorls. AI the

cords prominent and bearing regularly spaced

undulations, sometimes with a slight swelling. Suture

marked by a fine spiral thread. Intervals between

cords bearing very numerous close-set lamellose

axial riblets, and a very fine spiral thread on last

whorls. Last whorl with a fifth weaker cord,

emerging from suture, followed immediately by a

sixth cord; the remaining part of base smooth.

Aperture quadrangular; anterior canal short and very

oblique. Colour plain orange, protoconch a little

more brownish.

Size: holotype total height 5.4 mm: width at base 2

mm. Height of protoconch 0.66 mm; maximum

diameter of protoconch 0.40 mm.

Type locality. Found dead in hand-dredged sand at

10-20 m off Saint-Gilles-les-Bains, between

Hermitage and Boucan-Canot beach.

Type material. Holotype and paratype 1 in MNHN,

paratypes 2 to 11 coll. M. Jay: paratypes 12 to 16

coll. J. Drivas.

Etymology. Dedicated to Dr. J.C. Chenu, French

conchologist in last century.

Remarks. This species resembles the following one

(Seila reunionensis n.sp.) from which ït is

undistinguishable on teleoconch characters alone, but

is easily separated from it by its protoconch of 6

whorls instead of 2.5. Other remarks will be

discussed with the following species.

Seila reunionensis n.sp.

Plate 9, G; colour plate II, Fig. 60

Material examined. 2 spmns MNHN: 19 spmns (10

with complete protoconch) coll. M. Jay: 5 spmns

coll. J. Drivas.

Description. Shell elongate conical, slightly fusiform,

not constricted at base. Protoconch of 2.5 whorls,

regularly tapering to apex; first whorl rather wide, the

earlier 0.25 whorl smooth, the following ones with

axial ribs extending from suture to suture, and 4 finer

spiral cords in the lower half of their intervals; limit

from teleoconch ill-defined on 1/4 whorl, marked by

development of 2 spiral cords deforming axial ribs.

Teleoconch of 11 whorls with straight sides, suture

weakly impressed marked from earlier whorls onwards

by a fine undulose spiral thread: 2 spiral cords per

whorl on the 3 earlier whorls: then a third spiral cord

arising at the end of fourth whorl, developing from a

fine thread situated between the 2 cords, this third cord

remaining weaker than the earlier two except on last

whorl. At the beginning of the tenth whorl, a fine

spiral thread emerges between the second and third

cords, developing into a fourth cord, distinct on last

whorl but remaining weaker than the three other ones.

A fifth spiral cord, weaker than the fourth one,

emerging from suture at base of last whorl,

immediately followed by a sixth cord still a little

weaker than the fifth one. AI these spiral cords with

wide undulations, sometimes with slight swelling. The

intervals between cords with numerous close-set very

fine lamellose axial riblets, and a fine spiral thread on

last whorls. Base smooth. Aperture roundly quadrate,

anterior canal wide and strongly oblique. Colour plain

orange brown.

Size: maximum height 7.6 mm; width at base 1.9 mm:

height of protoconch 0.56 mm: width of protoconch

0.43 mm.

Type locality. Found dead in hand-dredged sand at

10-20 m, off Saint-Gilles-les-Bains.

Type material. Holotype and paratype 1 in MNHN:

paratypes 2 to 8 coll. M. Jay: paratypes 9 and 10 coll.

J. Drivas.

Etymology. Named after Reunion island.

Remarks. This species resembles the preceeding one

(Seila chenui n.sp.) in that its teleoconch bears spiral

cords with distinct undulations, and differs from it

only in its 2,5 protoconch whorls instead of 6. Both

species are provisionally attributed to the genus

Seila, though their spiral cords are undulose and

occasionally slightly swollen. Some specimens

resembling ours have sometimes been identified as

Seila laqueata (Gould, 1861) (type locality China

Seas) but Gould did not describe the protoconch of

his species, and the figure given by Johnson shows a

Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

PLATE 9. Fig. A. Dizoniopsis herosae n.sp. Off Souris-Chaude, Trois-Bassins, 30 m; holotype, height 4.8 mm:

MNAN. Fig. B. D. herberti n.sp. Off Saint Gilles les Bains, 10-20 m; holotype, height 2.4 mm; MNEN.

Fig C. D. gofhica n.sp. Off Saint Gilles les Bains, 10-20 m; holotype, height 2.7 mm; MNEAN. Fig. D. Sei/a

bandorensis (Melvill, 1892). Off Saint Gilles les Bains, 10-20 m; height 3.8 mm; MNAHN. Fig. E. S. hinduorum

(Melvill, 1898). Off Saint Gilles les Bains, 40-50 m; height 5,5 mm; MNAN. Fig. F.S. chenui n.sp. Off Saint

Gilles les Bains, 10-20 m; holotype, height 5.4 mm; MNAHN. Fig G.S. reunionensis n.sp. Off Saint Gilles les

Bains, 30-50 m; holotype, height 7.6 mm; MNAHN.

Scale bars: S (shells): 1 mm; P (protoconchs): 100 pm.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

shell broken at both ends, bearing 3 spiral cords per

whorl, unequal, smooth, neither undulose nor

swollen. A lot labelled Sei/a laqueata (Gould, 1961)

in NMW (lot N° Z.1955.158.02266 , 2 specimens

from Hong-Kong) shows only 3 smooth spiral cords.

Our new species differs from Seila crocea Angas,

1878 (type locality NSW, Australia), in its fourth

spiral cord and the undulations on the cords; Angas

did not describe the protoconch of his species. One

specimen labelled Seila crocea in NMW (lot

Z.1955.158.02267 ) shows four spiral cords on last

whorl, but all of them are smooth without any

undulation, and have a colour pattern with scattered

white blotches, very different of our species. Our

specimens were compared with Seila cinctum Dunker

(MNK lot Nr 3237 ), Seila capitata Thiele, 1925

(type locality Agulhas bank) (MNK lot Nr 102723);

Seila alfredensis Bartsch, 1915 (type locality Port

Alfred, South Africa) (NMW, not material type); and

Seila dissimilis Sutter, 1908, type locality New

Zealand (NMW, not material type) but all these

species differ from ours by their smooth spiral cords

instead of regularly undulose or slightly swollen.

ACKNOWLEDGEMENTS

We are particularly grateful to Dr. Philippe Bouchet,

MNHN, Paris, for his aid and remarks, and for

allowing us to examine material in MNHN; without

him, this study would not have been performed; and

to Dr. Dai Herbert, Natal Museum for his aid, and the

re-reading of the manuscript.

We thank similarly Mrs J. Pickering and Mrs K.

Way, NHM, London; Dr M. Glaubrecht, MNK,

Berlin, Mr. D. Penney, MM, Manchester; Dr. M.

Seddon and Mrs H. Wood, NMW, Cardiff, for their

aid and for having given us the opportunity to

examine collections. Mr Lozouet and Mr. Metivier,

MNEN, for their aid in bibliography; Mr. B.

Fontaine and A. Abdou, MNEHN, for their aid in

photos SEM; Mr. Claude Michel, late curator in the

Museum of Port Louis, Mauritius; Mr. Pierre Viader,

Cardiff for their advice.

REFERENCES

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Leiostraca and Cerithiopsis from Japan. Ann. and

Mag. Nat.Hist., 1861: 9-16.

Angas F. 1877. Description of two genera and twenty

species of Marine Shells from NSW. Proc. Zool.

Soc. London 1877: 35-40.

Barnard K.H. 1963. Contributions to the knowledge

of South African Marine Mollusces, Part 3

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199.

Bartsch P. 1915. Report on the Turton collection of

South African marine mollusces. United States

Nat.Mus. Bull. 81:1-305.

Bouchet P. et Danrigal F. 1982. Napoleon’s egyptian

campaign (1798-1801) andthe Savigny

collection of shells. The Nautilus, 96 (1): 9-19.

Cotton B. 1951. Australian recent and tertiary

Mollusca, family Cerithiopsidae. Rec. S. Aust.

Mus. 9(4): 383-395

Dunker G. 1861. Mollusca japonica. Stuttgartiae

Types et sumtibus E. Schweizerbat

Dushane H & Draper B. 1975. The genus Seila in the

Eastern Pacific. Veliger 17 (4): 335-345

Finlay H.J. 1927: A further commentary on New

Zealand Mollusca Systematics. Transactions of the

N.Z.Institute 57: 320-485

Finlay 1928 The recent Mollusca of the Chatham

islands. 7rans. N. Zeal. Inst. 59:232-286

Fischer P. 1887. Manuel de conchyliologie. Paris,

Savy éd.

Forbes E. & Hanley $S. 1850. A history of British

Mollusca and their shells. Vol 3 London, Van

Voorst ed.

Glibert 1973. Révision des Gastropodes du Darien et

du Moustien de la Belgique. Inst.Royal Sciences

Nat. Belg. Mémoire N° 173

Gougerot & Le Renard 1980. Clé des genres fossiles

du bassin de Paris. Cahier des Naturalistes 36: 17-

Gould A.A. 1861. Descriptions of new shells,

collected by the United States North Pacific

Expedition. Proc. Boston Soc. Nat. Hist. 7: 387-

388

Grundel J. 1989. Bemerkungen zur überfamilie

Cerithiopsacea H. & A. Adams, 1854

(Gastropoda). So wie zur Fassung etniges iher

Gattungen. Zool. agischen Anseige. 204 (3-4):

209-264

Habe T. 1970. Two new Cerithiopsid species from

Japan Venus 29(2):55-57

Hedley C. 1899. The Mollusca of Funafuti Part 5

Gasteropoda. Mem. Aust. Mus. part 7 (3): 337

Hedley C. 1909. The Mollusca from the Hope

Island. Proc. Linn. Soc. NSW. 34: 441

Hedley C. 1911. Report on the Mollusca obtained by

the FIS Endeavour Part 1 Zoological Results of the

fishing experiments carried out by the FIS

Endeavour, 1909- 10, Part 2:90-114

Issel A. 1869. Malacologie del Mar Rosso, Pisa.

Johnson R. 1964. The Recent Molluses of A.A.

Gould; illustrations of the types described by

Gould. United States Nat. Mus. Bull. 239.

Kay A. 1979. Hawaïian Marine Shells. Bishop

Museum Press, Honolulu, pp 125-128

Kilburn R.N. Taxonomic notes on South African

marine Mollusca (5). Ann. Nat.Mus. 22(2):577-

622

Kuroda T. Habe T & Oyama K. 1971. The sea shells

of Sagami Bay. Tokyo, Marusen ed.

Lamy 1909. Les coquilles marines recueillies par M.

Geay à Madagascar. Mem. Soc. Zool. France

XXII (3-4). Bulletin du MNHN 1909 6: 368

Laseron C.F. 1951. Revision of the New South Wales

Cerithiopsidae. Austr. zool. 11(4):351-367

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NOVAPEX 3 (1): 1-45, 10 mars 2002

Laseron C.F. 1956. The family Cerithiopsidae

(Mollusca) from the Solanderian and Dampierian

zoogeographical provinces. Autr. J. mar.

freshwater research 7(1):151-182

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malacologique de l'ile Maurice et ses

dépendances. Paris, Tremblay ed.

Marshall B.A. 1978. Cerithiopsidae (Mollusca:

Gastropoda) of New Zealand. New Zeal. J. of

Zoology 5: 47-120

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Gulf. Proc. Mal. soc. Lon. 1896: 10-11

Melvill J.C. 1896. Descriptions of new species of

minute marine shells from Bombay. Proc. Malac.

Soc. Lon. 2(3):108-116

Melvill J.C. 1898. Further investigations into the

Molluscan Fauna of the Arabian sea, Persian

Gulf and Gulf of Oman, with description of 40

species. Mem. Proc. Manchester Liter. Philos.

Soc 42(4): 1-185

Melvill J.C. 1907. Description of 31 gastropoda and

one scaphopod from the Persian Gulf and Gulf of

Oman, dredged by M.F.W. Townsend. Proc.

Malac. Soc. Lon. 7:69-80

Melvill J.C. & Standen R. 1896. Notes on a

collection of shells from Lifu and Uvea, Loyalty

Islands formed by the Rev. James and Mrs

Hadfield with list of species. Journal of

Conchology 8(2): 273-303

Monterosato 1874. Recherches conchyliologiques

effectuées au cap Santa Vita de Sicile par le

Marquis de Monterosato. Traduit de l'Italien par H.

Crosse. Journal de Conch, 1874 : 243-282

Nordsieck F. 1968. Die europaïschen Meres

gehauseschecken von Eismer bis Capverden und

Mittelmeer. Tafel XI Stuttgart, Fischer ed.

Nutzel A. 1997. Über die Stammes geschichte der

Ptenoglossa (Gastropoda). Berliner Growissens

schaftliche Abbarlungen 220 p.

Odhner N. 1924. New Zealand Mollusca. Papers

from Dr. Th. Mortensen’s Pacific Expedition

1914-1916. Videnskabelige Meddelelser fra

Dansk Naturhistorisk Rorening i Kjobenhavn

77:1-90

Oliver B.1915. The Mollusca of the Kermadec

Islands. 7rans. N.Z. Institute 47: 509- 568

Powell A.W.B. 1940. The marine Mollusca of the

Ampourian province. 7rans. Proc. of Rose Soc.

70 (2):209-248

Preston H.P. 1905. Description of new species of

Marine Shells from Ceylon. Journal Malacology

XIL:1-10

Sacco 1895. I Molluschi dei terreni terziarii del

Piemonti 1 della Liguria. Torino, Carlo Clausen

ed. Parte XVII.

Smith E.A. 1906. South African Marine Mollusca.

Ann. Nat. Mus. 1:19-71

Smith E.A. 1910. On South African Marine

Molluscs with description of new species. Ann.

Nat. Mus. 11(2):175-219

Sowerby G.B. 1897. Marine shells of South Africa et

Appendix. London, Sowerby ed.

Suter H. 1918. Manual of the New Zealand Mollusca.

Wellington, Mackay ed. pp: 244-254

Thiele J. 1925. Gastropoda der deutschen Tiefsee

Expedition. Fisher; Iena

Viader R. 1937. Revised catalogue of the testaceous

Molluses of Mauritius and its dependancies.

Mauritius Inst. Bull. 21(2): 1-111

Watson R.B. 1885: on the Cerithiopsidae from east

side North Atlantic, with 3 new species from

Madeira. Linnean Soc. Journal. Zoology Vol

XIX: 89-95

Watson R.B. 1886. Report on the scientific results of

the voyage of HMS Challenger. Zoology

XV:525-530

JAY & DRIVAS Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

COLOUR PLATES

JAY & DRIVAS Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

COLOUR PLATE 1

erithiopsis eutrapela Melvill & Standen, 1896. Off Saint Gilles les Bains, 10-20 m; height 6.5; coll. M. Jay.

= foster…ae Melvill & Standen, 1896. Off Saint Gilles les Bains, 30-S5m; height 6.1 mm; coll. M. Jay.

=. boucheti n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 4.5 mm; coll. M. Jay.

| hadfieldi n.sp. Off Saint Gilles les Bains, 45 m; paratype 2, height 5.6mm; coll. M. Jay.

= iochrous n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 5.5 mm; coll. M. Jay.

=. jousseaumei n.sp. Off Saint Gilles les Bains, 10-30 m; paratype 2, 6 mm, coll. M. Jay.

. lamyi n.sp. Off Souris-Chaude, Trois-Bassins, 30 m; paratype 2, height 1.5 mm; coll. M. Jay.

. nutzeli n.sp. Off Saint Gilles les Bains, 15-30 m; paratype 2, height 3.4 mm; coll. M.Jay.

. pickeringae n.sp. Off Saint Gilles les Bains, 20-30 m:; paratype 2, height 3.9 mm; coll. M. Jay.

10. Cerithopsis seddonae n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 2.2 mm; coll. M. Jay.

11. C. vaurisi n.sp. Possession Bay, 55m; paratype 2, height 3.6 mm; coll. M. Jay.

12. C. wayae n.sp. Off Souris-Chaude, Trois-Bassins, 30 m; paratype 2, height 4.1 mm; coll. M. Jay.

13. Joculator albocinctum Melvill & Standen, 1896. Off Saint Gilles les Bains, 10-20 m, height 3 mm; coll. M.

Jay.

14.J. granata Kay, 1979. Off Saint Gilles les Bains, 10-20 m; height 2.5 mm; coll. M. Jay.

15.J. minima Laseron, 1955. Off Saint Gilles les Bains, 10-20 m; height 2.3 mm; coll. M. Jay.

16..J. minutissima (Thiele, 1925). Off Saint Gilles les Bains and Possession Bay 10-54 m; height 1.5 mm; coll.

M. Jay.

17.J. pulvis (Issel, 1869). Off Saint Gilles les Bains, 10-20 m; height 2.9 mm; coll. M. Jay.

18./. christiaensi n.sp. Off Saint Gilles les Bains, 55 m; paratype 1, height 1.9 mm; coll. M. Jay.

19. J. eudeli n.sp. Off Saint Gilles les Bains, 30 m; paratype 2, height 1.9 mm; coll. M. Jay.

20.J. fischeri n.sp. Off Souris-Chaude, Trois-Bassins, 30 m; paratype 2, height 2.3 mm; coll. M. Jay.

21.J/. keratochroma n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 2.5 mm; coll. M. Jay.

22.J. laseroni n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 30 m; paratype 2, height 2.3 mm; coll. M.

Jay.

23.J. lozoueti n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 3.2 mm; coll. M. Jay.

24.J. megacephala n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 1.8 mm; coll. M. Jay.

25.J. melanoraphis n.sp. Off Saint Gilles les Bains, 10-30 m; paratype 2, height 2.9 mm; coll. M. Jay.

26.J. mygaki n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 30 m; paratype 2, height 1.9 mm; coll. M.

Jay.

27.J. myia n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 30 m; paratype 2, height 2.2 mm; coll. M.

Jay.

28.J. phtyr n.sp. Off Saint Gilles les Bains, 35 m; paratype 2, height 2 mm; coll. M. Jay.

29.J. psyllos n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 2 mm; coll. M. Jay.

30.J. salvati n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 35 m; paratype 1, height 3.6 mm; coll. M.

LOIR AAA Are 'œ

Jay.

31.J. skolix n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 35 m; paratype 2, height 3.2 mm; coll. M.

Jay.

32. J. thielei n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 1.5 mm;

coll. M. Jay.

33..J. vignali n.sp. Off Souris-Chaude, Trois-Bassins, 30 m; paratype 1, height 1.4 mm; coll. M. Jay.

Le)

‘a

—

JAY & DRIVAS

JAY & DRIVAS Cerithiopsidae Reunion NOVAPEX 3 (1): 1-45, 10 mars 2002

COLOUR PLATE 2

34. Horologica balteata (Watson, 1886). Off Saint Gilles les Bains, 12m; height 3 mm; coll. M. Jay.

35. H. bicolor Laseron,1956. Off Saint Gilles les Bains, 30 m:; height 2.1 mm; coll. M. Jay.

36. H. macrocephala Laseron, 1956. Off Saint Gilles les Bains, 20-30 m; height 1.7 mm; coll. M. Jay.

37. H. minareta Laseron, 1956. Off Saint Gilles les Bains, 10-20 m; height 2.6 mm; coll. M. Jay.

38. H. purpurea Laseron, 1955. Off Cape La Houssaye, Saint Gilles les Bains, 10-20 m; height 3.1 mm; coll.

M. Jay.

39. H. cf semipicta (Gould, 1861). Off Saint Gilles les Bains, 30 m; height 2.1 mm, coll. M. Jay.

40. FH. turrigera (Watson, 1886). Off Saint Gilles les Bains, 10-20 m; height 3.7 mm; coll. M. Jay.

41. H. anisocorda n.sp. Off Saint Gilles les Bains, 30m; paratype 2, height 2.2 mm; coll. M. Jay.

42. H. glaubrechti n.sp. Off Saint Gilles les Bains, 30-50 m; paratype 2, height 2.3 mm; coll. M. Jay.

43. H. konops n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 30 m; paratype 2, height 3mm; coll. M.

Jay.

44. H. martini n.sp. Off Boucan-Canot beach, Saint Gilles les Bains, 30 m; paratype 2, height 2.7 mm; coll.

M. Jay.

45. Dizoniopsis gothica n.sp. Off Saint Gilles les Bains, 10-20 mi; paratype 2, height 2.7 mm; coll. M. Jay.

46. D. herberti n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 2.4 mm; coll. M. Jay.

47. D. herosae n.sp. Off Souris-Chaude, Trois-Bassins, 30 m; paratype 2, height 4.8 mm; coll. M. Jay.

48. Mendax mascarenensis n.sp. Off Saint Gilles les Bains, 30 m; paratype 2, height 4.2 m; coll. M. Jay.

49. M. metivieri n.sp. Off Saint Gilles les Bains, 30-50 m; paratype 2, height 4.3 mm; coll. M. Jay.

50. M. penneyi n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 1, height 4.1 mm; coll. M. Jay.

51. M. ribesae n.sp. Off Saint Gilles les Bains, 55 m; paratype 2, height 2.6 m; coll. M. Jay.

52. M. theodosiae n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 3.3mm,; coll. M. Jay.

53. Prolixodens nicolayae n.sp. Off Souris-Chaude, Trois-Bassins, 30 m; paratype 2, height 1.4 mm; coll. M.

Jay.

54. P. sknips n.sp. Off Saint Gilles les Bains, 10-30 m; paratype 2, height 1.8 mm; coll. M. Jay,

55. Belonimorphis belonimorphis n.p. Off Saint Gilles les Bains, 10-30 m; paratype 2, height 6 mm; coll. M.

Jay.

56. Koilofera koilofera n.sp. Off Saint Gilles les Bains, 10-30 m; paratype 2, height 2.7 mm; coll. M. Jay.

57. Seila bandorensis (Melvill, 1892). Off Saint Gilles les Bains, 10-20 m; height 3.8 mm; coll. M. Jay,

58. S. hinduorum (Melvill, 1898). Off Saint Gilles les Bains, 40-50 m; height 5,5 mm; coll. M. Jay.

59.S. chenui n.sp. Off Saint Gilles les Bains, 10-20 m; paratype 2, height 5.4 mm; coll. M. Jay.

60. S. reunionensis n.sp. Off Saint Gilles les Bains, 30-50 m; paratype 2, height 7.6 mm; coll. M. Jay.

JAY & DRIVAS

Cerithiopsidae Reunion

NOVAPEX 3 (1): 1-45, 10 mars 2002

GUILLOT DE SUDUIRAUT

Scabricola lavoisieri n.sp.

NOVAPEX 3 (1): 47-49, 10 mars 2002

Description d’une nouvelle espèce de mitre des Philippines

(Sous-famille des Imbricariinae)

(Gastropoda:Prosobranchia:Mitridae)

Emmanuel GUILLOT de SUDUIRAUT

P.O. Box 104

Lapu Lapu City, Cebu, Philippines

conchyliologie(@eurasiashells.net

MOTS CLES. Gastropoda, Mitridae, Imbricariinae, Scabricola (Scabricola), nouvelle espèce,

Philippines.

KEY WORDS. Gastropoda, Mitridae, Imbricariinae, Scabricola (Scabricola), new species,

Philippine Islands.

RESUME. Scabricola (Scabricola) lavoisieri n. sp. est décrite des Philippines et comparée avec

S.(S.) potensis (Montrouzier, 1858) de Nouvelle-Calédonie.

ABSTRACT. Scabricola (Scabricola) lavoisieri n. sp. is described from the Philippine Islands. It

is compared with S. (S.) potensis (Montrouzier, 1858) from New Caledonia.

INTRODUCTION.

Dans la sous-famille des Imbricariinae, le genre

Scabricola (Scabricola) était représenté par 10

espèces (Cernohorsky 1991) : S. variegata (Gmelin,

1791), S. desetangsii (Kiener, 1838), S. potensis

(Montrouzier, 1858), S. caerula (Reeve, 1844), S.

vicdani Cernohorski, 1981, S. eximia (A. Adams,

1853), S. coriacea (Reeve, 1845), S. padangensis

(Thiele, 1925), S. alabaster (Sowerby, 1900). Une

espèce a été récemment décrite: S. condei Guillot

de Suduiraut, 2001. Avec la description de S. {S.)

lavoisieri n. sp. le nombre d’ espèces appartenant à

Scabricola (Scabricola) connues à ce jour est de 11.

ETUDE SYSTEMATIQUE

Famille MITRIDAE Swainson., 1831

Sous-famile IMBRICARIINAE Troschel, 1867

Espèce type : Voluta variegata Gmelin, 1791

Scabricola (Scabricola) lavoisieri n. sp.

Figs 1-5

Materiel type. Holotype Muséum national

d'Histoire naturelle, Paris, France; paratype 1 : coll.

de l’auteur; paratype 2 : coll. R. Salisbury; paratype

3: coll. A. Deynzer.

Localité type. Philippines, au sud de l'Ile de

Balicasag, Panglao-Bohol, au filet, 140 — 180 m,

sur fond de sable.

Description. Coquille de petite taille pour le genre.

Protoconque inconnue. Tours de spire fusiformes,

légèrement convexes; dernier tour cylindrique.

Sutures peu profondes. Epaulement crénelé. Les

deux avant-derniers tours portant cinq cordons

spiraux arrondis, interspaces striés axialement.

Dernier tour portant 17 ou 18 cordons spiraux.

Base de l’ouverture plus ovoïde, sa hauteur

représente 47% de la hauteur de la téléoconque.

Lèvre exterieure ondulée. Fasciole siphonale

droite,distincte, orné de quatre plis columellaires.

Couleur blanche, maculée irrégulièrement d’une

bande jaune-orange au-dessus des sutures. Dernier

tour portant une large bande de même couleur au-

dessus de l’angle parietal, avec des taches blanches

sur le cordons spiraux. Ouverture blanc-crème,

periostracum transparent.

Discussion. Scabricola (Scabricola) lavoisieri n.

sp. diffère de S. potensis (Montrouzier, 1858) par sa

spire plus fusiforme, son avant-dernier tour portant

5 cordons spiraux, le dernier tour 17 ou 18 au lieu

de 6 ou 7 cordons sur l’avant dernier tour et 30 à

40 sur le dernier tour chez S. (S.) potensis. S.

lavoisieri diffère également par la hauteur de son

ouverture (47 % de la hauteur de sa téléoconque au

lieu de 67 % chez S. potensis), par sa couleur

blanche, par le nombre de ses plis columellaires (4

au lieu de 6 ou 7 chezS. potensis) .

Dimensions

Holotype : 16, 7 x 6, 5 mm ; hauteur de l’ouverture:

8,4 mm.

Paratype 1: 13,2 x 5, 3 mm; hauteur de l’ouverture:

72 Sn:

Paratype 2: 12, 7 x 5, 3 mm; hauteur de l’ouverture:

6,2 mm.

Paratype 3: 15,2 x 5, 6 mm; hauteur de l’ouverture:

7,9 mm.

GUILLOT DE SUDUIRAUT

Scabricola lavoisieri n.sp.

NOVAPEX 3 (1): 47-49, 10 mars 2002

Etymologie. Cette nouvelle espèce est nommée en

mémoire de Antoine Laurent de Lavoisier (1743 —

1794), père de la chimie moderne et aïeul de

l’auteur.

REMERCIEMENTS.

Je suis particulièrement reconnaissant à Virginie

Héros, à Pierre Lozouet Muséum national

d'Histoire naturelle, Paris pour le cliché de l

holotype de Mitra potensis, à R. Houart (Landen,

Belgium) pour ses commentaires et corrections du

manuscrit et à G. Poppe (Bruxelles, Belgium) pour

la mise en page de la planche.

REFERENCES.

Cernohorsky, W. O. 1991. The Mitridae of the

World. Part 2 The subfamily Mitrinae

concluded and subfamilies Imbricariinae and

Cylindromitrinae. Monographs of Marine

Mollusca. Trophon Corporation, Silver Spring,

U.S.A. 164 pp

Guillot de Suduiraut, E. 2001. Description de

Scabricola (Scabricola) condei n. sp.

(Gastropoda: Prosobranchia : Mitridae) des

Philippines. Novapex 2 (1) : 21-23.

1-5. Scabricola (Scabricola) lavoisieri n. sp. Philippines, Ile de Balicasag, Panglao-Bohol, 140-180 m.

1-2 Holotype MNHN: 16,7 mm x 6,5 mm; hauteur de l’ouverture: 8,4 mm.

3. Paratype collection de l’auteur: 13,2 mm x 5,3 mm; hauteur de l’ouverture: 7,3 mm.

4. Paratype collection R. Salisbury: 12,7 mm x 5,3 mm; hauteur de l’ouverture: 6,2 mm.

5. Paratype collection A. Deynzer: 15,2 mm x 5,6 mm; hauteur de l’ouverture: 7,9 mm.

6-7. Scabricola (Scabricola) potensis (Montrouzier, 1858): 24,5 mm x 11,4 mm; hauteur de l’ouverture: 16,5

mm. Île de Pot, Nouvelle-Calédonie. Holotype MNHN.

GUILLOT DE SUDUIRAUT Scabricola lavoisieri n.sp. NOVAPEX 3 (1): 47-49, 10 mars 2002

à ‘

284

POPPE & TERRYN

Lyria pauljohnsoni sp. nov.

NOVAPEX 3 (1): 51-53, 10 mars 2002

Description of a new volute (Gastropoda: Volutidae)

from southern Madagascar

Guido T. POPPE

Stanislas Leclefstraat, 8, 2600 Berchem, Belgium

Yves TERRYN

Acaciastraat, 44, 9000 Gent, Belgium

KEY WORDS. Gastropoda, Volutidae, Lyria (Indolyria) pauljohnsoni sp. nov., Madagascar.

ABSTRACT. ZLyria (Indolyria) pauljohnsoni sp. nov., from Madagascar, is hereby described and compared

with its closest relatives Lyria (Indolyria) brianoi Poppe, 1999 and ZLyria (Indolyria) patbaili Bouchet, 1999.

RESUME. Zyria (Indolyria) pauljohnsoni sp. nov. de Madagascar, est décrite et comparée avec les espèces

proches, Lyria (Indolyria) brianoi Poppe, 1999 et Lyria (Indolyria) patbaili Bouchet, 1999.

INTRODUCTION

Two years after the description of Lyria (Indolyria)

brianoi and L. (1) patbaili, fishermen from Fort

Dauphin, Madagascar, discovered a third species of

Indolyria living in the waters of southern

Madagascar. The new species definitely belongs to

the subgenus /ndolyria and is very closely related to

L. (1) brianoi.

SYSTEMATICS

Class GASTROPODA

Family VOLUTIDAE Rafinesque, 1815

Subfamily VOLUTINAE Rafinesque, 1815

Tribe LYRIINI Pilsbry & Olsson, 1954

Genus Lyria Gray, 1847

Type species: Lyria nucleus (Lamarck, 1811)

Subgenus /ndolyria Bail & Poppe, 2001

Type species: Lyria (1) lyraeformis (Swainson,

1821)

Lyria (Indolyria) pauljohnsoni sp. nov.

Figs 1-6

Type Material. Holotype MNHN, Paris. Length:

37.3 mm. Maximum width: 17.1 mm. ex coll. P.

Johnson, Great Brittain.

Paratype 1 Coll. G. T. Poppe. Length: 35.5 mm.

Maximum width: 17.5 mm. Animal and operculum

dried inside.

Paratype 2 Coll. J. Conde. Length: 32.5 mm.

Maximum width: 16.1 mm. Animal and operculum

dried inside.

Type Locality. Madagascar, Fort Dauphin area.

Range. Only known from the type locality.

Habitat. Unknown.

Description. The shell has an ovate, elongate shape

and is thick and solid. The spire is very small and

covers less than a third of the shell length. The

protoconch is broad and has about 2 whorls and

presents in fresh specimens a silky gloss. The

transition into the teleoconch is gradual and only

visible due to the occurrence of faint axial plicae.

The holotype has a protoconch of 3.2 mm at its base.

This is very large compared to the total shell width.

The teleoconch consists of 4.5 flattened whorls. On

the first teleoconch whorl, just behind the

protoconch, there appear a few faint axial ribs which

rapidly become very weak and irregular on the next

whorl. The overall appearance of the body whorl is

smooth. On the back of the last whorl, situated close

to the siphonal canal, all three type specimens show

7-9 hardly discernible spiral grooves. The outer lip is

thickened all along the peristome and flares at its

lower half. The parietal path bears several plicae of

which 3, situated near the siphonal canal are very

strong. On the posterior end of the aperture, there 1s a

channel, which is about 1.2 mm in width and 5 mm

in length. The suture is deep, almost channelled. The

outer surface has a silky gloss and the interior of the

aperture is very glossy. The parietal callus is thick for

the genus and covers the fasciole. The colour pattern

is complex: the protoconch whorls are light brown

with a tinge of purple, the teleoconch whorls have a

purple-brown base colour with 3 spiral bands of

interrupted cream and black blotches. Below these

spiral bands there is another, parallel, spiral band, in

which the black blotches are mirrored by cream

blotches. This is especially visible in the holotype.

Both the paratypes are not so dark coloured: the

shells are cream-chocolate and the spiral bands are

POPPE & TERRYN

Lyria pauljohnsoni sp. nov.

NOVAPEX 3 (1): 51-53, 10 mars 2002

less obvious. On the thickening along the peristome

appear 7-9, very dark coloured spiral lines. They

become very prominent and continue on the

thickened outer lip and form brown spots on the

inside of the lip. The outside of the siphonal canal is

darker coloured than the rest of the shell.

Animal and radula. Unknown. Animal dried inside

in both paratypes. Operculum present, about 10 mm

in length.

Differential diagnosis. L. (1) pauljohnsoni differs

from L. (1.) brianoiï at first glance by the presence of

an obvious much larger channel on the posterior side

of the aperture. L. (/.) brianoï has distinct axial ribs

while the shell is smooth in Z. (Z.) pauljohnsoni. The

number of plicae on the parietal shield is much larger

and more obvious in L. (I ) brianoi. While L. (1)

brianoi most often has an orange-brown colour, Z.

(1) pauljohnsoni is much more chocolate with a

pinkish shine. The spire in the latter species is much

smaller compared to the total shell length.

L. (1) patbaili is, in general, twice as large as an

average L. (1) pauljohnsoni, it has only a faint

channel on the posterior end of the aperture, is

axially ribbed, has a different colour pattern and a

much longer spire.

Remarks. This new species, at present known from

the type material and one very beach-worn shell,

belongs to the subgenus 7/ndolyria, Which is

characterised by thick shells with a very large

protoconch. This subgenus is limited to the Indian

Ocean.

Derivatio nominis. The shell is named in honour of

Paul Johnson, Great Britain, active volute-collector

who donated the holotype to the MNHN, Paris.

ACKNOWLEDGEMENTS

We thank Michel Charles for providing us the

paratypes, Javier Conde for critically rereading and

commenting the manuscript.

SELECTED REFERENCES

Bail, P. 1993. Le genre Lyria Gray, 1847. Part I.

Xenophora 61 : 6-23.

Bail, P. 1993. Le genre Lyria Gray, 1847. Part II

Xenophora 64 : 4-19.

Bail, P. & Poppe, G.T. 2001. À Conchological

Iconography : A Taxonomic Introduction to the

Recent Volutidae, Conchbooks, Hackenheim,

Germany: 50 pp.

Poppe, G.T. 1999. New Volutidae of the Indian

Ocean and Australia. Malacologia, 29 : 3-7.

Poppe, G.T. & Goto, Y. 1992. Volutes, Ancona,

Italy: 348 pp.

Weaver, C.S. & duPont, J.E. 1970. The Living

Volutes, a Monograph of the Recent Volutidae of

the World., Delaware Museum of

Natural History, Greenville, USA: 375 pp.

1-6. Lyria (Indolyria) pauljohnsoni Poppe & Terryn, sp. nov.

1-3. Holotype MNHN, Madagascar, Fort Dauphin area. 37.3 mm.

4-5. Paratype 1, coll. G.T. Poppe. 35.5 mm.

6. Paratype 2, coll. J. Conde: 32.5 mm.

7-8: Lyria (Indolyria) brianoi Poppe, 1999. Madagascar, Fort Dauphin area, 80-150 m.

7. 31.2 mm. 8. 28.1 mm.

9. Lyria (Indolyria) patbaili Bouchet, 1999. Madagascar, Fort Dauphin area, 80-150 m, 65.1 mm.

. 10 mars 200

-53

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NOTE AUX AUTEURS

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La liste des références, en ordre alphabétique, respectera la forme suivante (les titres des publications ne devraient pas être abrégés):

Keen, A.M. & Campbell, G.B. 1964. Ten new species of Typhinae (Gastropoda : Muricidae). TheVeliger 7(1): 46-57.

Powell, A.W.B. 1979. New Zealand Mollusca. Marine, land and freshwater shells. William Collins Publishers Ltd: xiv + 500 pp.

Mayr, E. 1989. Attaching names to objects. In: What the philosophy of biology is : essays for David Hull (M. Ruse, ed.),

Klumer Academic, Dordrecht: 235-243.

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Keen, A. M. & Campbell, G.B. 1964. Ten new species of Typhinae (Gastropoda : Muricidae). TheVeliger 7(l): 46-57.

Powell, A.W.B. 1979. New Zealand Mollusca. Marine, land and freshwater shells. William Collins Publishers Ltd: xiv + 500 pp.

Mayr, E. 1989. Attaching names to objects. In: What the philosophy of biology is : essays for David Hull (M. Ruse, ed),

Klumer Academic, Dordrecht: 235-243.

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E. Meuleman

R. Houart et

C. Vilvens

R. Houart

C. Vilvens

Vie de la Société — Life of the Society

(suite)

Une récolte éclair en Espagne

Quoi de neuf ?

La Fête de l'Eau

Quelques nouvelles publications

Nous avons reçu

Prochaines activités

Novapex / Société 3(1), 10 mars 2002 1

VIE DE LA SOCIETE LIFE Of Te SOCIETT

11 L'exposition 2002 de la SBM

Promenade au pays des malacologues de la SBM

Claude VILVENS et

Christiane DELONGUEVILLE, Ralph DUCHAMPS, Koen FRAUSSEN, Roland HOUART,

Annie LANGLEIT, Jeannine et René MASSON, Etienne MEULEMAN, Roland SCAILLET, Rita

SENDERS et Edgar WAIENGNIER

Cette tradition bien ancrée dans l'esprit des membres de la SBM a une fois de plus été respectée : la

première réunion de l'année 2002 a été consacrée à l'Exposition (avec un 'E' majuscule). Pour rappel, l'idée est

d'inviter les membres à présenter un thème malacologique, qu'il fasse partie de leurs préoccupations habituelles

ou, au contraire, qu'il appartienne à un domaine dans lequel on n'est pas habitué à les voir évoluer. Le public

visé ? Tout qui est intéressé par les coquilles, bien sûr !

La cuvée 2002 pourrait être qualifiée de variée et diversifiée. Si certains ont présenté un sujet somme

toute assez habituel pour eux (Roland, Claude, Etienne, Annie, Koen, … }, d'autres se sont aventurés dans des

terrains où l'on ne les attendait pas (Ralph, Jeanine et René, Fernand et Rika, ..). Cela donnait une formidable

promenade au pays des Mollusques dont les pages suivantes vont tenter de vous rendre compte. Suivez les

guides !

Précisons encore que les auteurs cités ci-dessus ont rédigé personnellement, avec

brio, l'article présentant leur thème d'exposition. Doublement merci à eux © !

2 NoOVAPEX / Société 3(1), 10 mars 2002

1. Etienne Meuleman : « Freak » spécimens chez Strombidae

Il arrive de trouver des coquilles qui diffèrent légèrement de la forme normale

de l'espèce à laquelle ils appartiennent. En effet, une cicatrice peut apparaître suite à une

1 blessure du manteau. Une maladie ou une carence alimentaire peut également engendrer

une déformation de la coquille qui lui donne parfois une forme bizarroïde. De telles

déformations se présentent régulièrement chez les Zambis. Certains Strombus et Tibia

présentent parfois des formes spéciales.

Sur la table d'exposition, on pouvait découvrir, quelques spécimens tels Lambis lambis (Linné, 1758),

Strombus urceus urceus ustulatus Schumacher, 1817 ou 7ïbia fusus (Linné, 1758). Ces coquilles aux formes

étranges ont parfois été nommées sous un autre nom en pensant qu’il s'agissait d’une nouvelle espèce.

E À

Freak: spécimens

chez Strombidac

NOVAPEX / Société 3(1), 10 mars 2002

mm rt, en

Dans cette grande famille des Buccinidae, le genre Beringius Dall. 1877 est un

{;. des plus célèbres. Tout comme chez Ancistrolepis Dall, 1895. Parancistrolepis Azuma.

+ O4 1965. Clinopegma Grant & Gale. 1931 ct surtout Neptunea Bolten in Rôding. 1798. les

coquilles de Beringius sont grandes et belles. garnies d'une sculpture souvent très

prononcée. La variabilité est étonnante et c'est un plaisir de collectionner les diverses

formes des espèces communes. C'est aussi une plaisir bien sûr d'essayer d'obtenir les

espèces très rares.

Les descriptions présentées par Habe & lio. 1965 des genres Neoberingius

(espèce type: Beringius frielei Dall. 1895) et Beringion (espèce type: Beringius marshalli Da. 1919) ne sont pas

très claires. Pour écarter ce problème J'ai choisi d'utiliser ici le nom génerique Beringius sensu stricto.

Golikov & Starobogatov (1975) ont décrit 1a famille des Beringiidae et l'ordre des Beringioidea (pour contenir les

Beringiidae et les Anachidae). Dix espèces actuelles sont connues, avec bien sûr un nombre de formes et de

synonymes. Plus de 15 espèces fossiles (Cenozoïque Supérieur, Eocène, Miocène. Pliocène) sont déja décrites

dans le genre Beringius, et 2 fossiles du Miocène dans le genre Tyrannoberingius (espèce type. attention. ne

tombez pas à la renverse: Tvrannoberingius rex Marincovich. 1981).

ee 2. Koen Fraussen : Le genre Beringius Dall, 1877 (Buccinid:

ill

IR ARLES LES

Beringius crebicostatus

Mr 2157

rdsmi

4 Novapex / Société 3(1), 10 mars 2002

Neuf espèces, 1 sous-espèce et plusieurs formes ont été exposées, toutes actuelles, listées ici par ordre de rareté :

Beringius turtoni (Bean, 1834) (Ocean Atlantique boréal-arctique) avec 4 formes: f. ossiania Friele, 1879; f

attenuata Simpson in Marshall, 1902: f minor Harmer, 1914 et la forme grande et large d'Islande.

Beringius beringii (Middendorff, 1848) (Océan Pacifique Arctique et la Mer de Bering) avec plusieurs formes:

grandes et petites, lisses ou fortement spiralées, fragiles ou très lourdes, et ce qu'on appelle généralement la forme

indentatus Dall, 1919 et kobelti Dall, 1902.

Beringius polynematicus Pilsbry, 1907 (Japon et Kamtchatka), avec periostracum jaune ou brun foncé. Cette

espèce est souvent présentée dans la litterature et dans les listes d'échange sous le nom de B. marshalli Dall, 1919.

Beringius frielei Dall, 1894 (Mer de Bering occidentale). Un exemplaire d'eau profonde avec une coquille âbimée

par l'acidité de l'eau de mer dans ces grandes profondeurs était également exposé.

Beringius frielei miyauchii Habe & Ito, 1972 (Mer de Okhotsk). Représenté à l'exposition par des exemplaires

fragiles ou robustes.

Beringius stimpsoni (Gould, 1860) (Alaska).

Beringius kennicottü (Dall, 1907) (Alaska). Représenté ici par la forme incisus Dall, 1907 et par une forme

presque lisse.

Beringius eyerdami AÀ.G.Smith, 1959 (Washington). Une forme sans cordons spiraux était exposée.

Beringius crebricostatus (Dall, 1877) (Iles Aleoutiennes). Extrêmement rare.

Beringius aleuticus Dall, 1894 (Iles Aleoutiennes bien sûr). Extrêmement rare. La coquille et surtout l'opercule

font penser au genre Japelion Dall, 1916.

Beringius undatus (Dall, 1919) n'était malheureusement pas exposé, c'est la seule espèce que je recherche encore.

Elle n'est pas très rare, mais apparemment j'ai eu plus de chances pour me procurer les autres espèces.

Beringius marshalli Dall, 1919 (Alaska) n'était pas exposée non plus (il s'agit probablement d'une forme de B.

polynematicus) ni Strombella malleata (Dall, 1884) (qui ressemble à une forme de B. beringii, mais on m'a fait

remarquer que l'holotype ressemble également à un freak de Buccinum glaciale).

NovarEx / Société 3(1), 10 mars 2002

L 3. Roland Houart : Muricidae : Les genres Murex s.s. et Haustellum

Se « J'avais choisi pour cette exposition, pour ne pas sortir de ma famille préférée, de

vous présenter les genres Murex sensu stricto et Haustellum. L'un et l'autre ont fait

récemment l'objet d'une révision par Ponder et Vokes (1988). Mes conclusions et mon

classement ne sont pas toujours identiques aux leurs mais cela n'enlève rien à la beauté

quelque peu irréelle! de ces coquillages. Le genre Murex est représenté par 31 espèces ou

sous-espèces possédant chacune une coquille très épineuse pouvant, comme chez M.

troscheli, parfois atteindre près de 20 cm. Le canal siphonal est très long et fin, épineux

également. Le dernier tour de spire est orné de 3 varices et le bord externe de l'ouverture

est pourvu d'une dent labrale. Sa distribution est largement Indo-Pacifique. Une espèce, Murex forskoehlii

Rôding, 1798 a pénétré en Méditerranée via le canal de Suez. Le genre Haustellum contient 11 espèces et sous-

espèces et à la différence du genre Murex s.s. il ne présente pas d'épines. Le canal siphonal est également très

long et fin. Le dernier tour de spire, large et globuleux, possède également 3 varices axiales. L'ouverture est

arrondie et large et ne présente pas de dent labrale. La distribution géographique est uniquement Indo-Pacifique.

Ci-dessous je vous propose de découvrir la liste de toutes les espèces actuelles. En gras les spécimens

exposés ce 12 janvier. F

be” 7

Wurex (Murex) scolopax Dillwyn. 1847

INDO-W PAC (D Southern part of the Red Sea. 1h

Gulf of Aden and the Persian Gulf

se | ; +979

Murex (Murex) poppei Mouart,

ed LV PAC LE, À small area DOtween thaland.

Samira nl Bornoo

! NDLR : notre ami se laisserait-il entraîner à des déclarations peu objectives :-) :-) ?

NOVAPEX / Société 3(1), 10 mars 2002

| MUREX 5.5.

| acanthostephes Watson, 1883

| aduncospinosus Sowerby, 1841

| africanus Ponder & Vokes, 1988

altispira Ponder & Vokes, 1988

brevispina brevispina Lamarck, 1822

brevispina macgillivrayi Dohrn, 1862

brevispina ornamentalis Ponder & Vokes, 1988

brevispina senilis Jousseaume, 1874

carbonnieri (Jousseaume, 1881)

concinnus Reeve, 1845

coppingeri E. A. Smith, 1884

falsitribulus Ponder & Vokes, 1988

Jorskoehlii Roding, 1798

hystricosus Houart & Dharma, 2001

kerslakae Ponder & Vokes, 1988

megapex Neubert, 1998

occa Sowerby, 1834

pecten pecten Lightfoot, 1786

pecten soelae Ponder & Vokes, 1988

Philippinensis Parth, 1994

poppei Houart, 1979

queenslandicus Ponder & Vokes, 1988

salomonensis P

Référence

scolopax Dillwyn, 1817

somalicus Parth, 1990

spectabilis Ponder & Vokes, 1988

spicatus Ponder & Vokes, 1988

surinamensis Okutani, 1982

tenuirostrum Lamarck, 1822

ternispina Lamarck, 1822

trapa Roding, 1798

tribulus Linnaeus, 1758

troscheli Lischke, 1868

HAUSTELLUM

barbieri Houart, 1993

bondarevi Houart, 1999

fallax (E. A. Smith, 1901)

franchii Bozzetti, 1993

haustellum (Linnaeus, 1758)

kurodai kurodai (Shikama, 1964)

kurodai langleitae Houart, 1993

kurodai vicdani Kosuge, 1980

longicaudum F. C. Baker, 1891

tweedianum (Macpherson, 1962)

wilsoni D'Attilio & OI

PONDER, WE. & VOKES, EH. 1988. Revision of the Indo-West Pacific fossil and Recent species of Murex 5.5.

and Faustellum (Mollusca: Gastropoda: Muricidae). Records of the Australian Museum, suppl. 8: 1-160.

NovarEx / Société 3(1), 10 mars 2002 7

4. Edgar Waiengnier : Les Facsimiliidae

De Quelle chance extraordinaire que d’assister à une « première » !

A À l’occasion de l’exposition annuelle de la S.B.M. chacun à pu découvrir avec

intérêt une nouvelle famille qui n’a pas fini de faire parler d’elle.

Cette famille, exclusivement limitée aux copies plus ou moins artistiques,

d’escargots nous a été dévoilée pour la première fois par LPG. (Evidemment !)

Le placement systématique de niveau supérieur à la famille est encore à l’étude. Nous

avons donc découvert la famille et tous les niveaux taxonomiques inférieurs. À savoir

Famille :

Facsimiliidae fam.nov. (copie)

S-famille :

Ludoninae s-f nov. (Jouets)

Xyloninae s-f. nov. (à base de bois)

Vitreaninae s-f. nov (à base de verre)

Terraninae s-f nov. (d’origine terrienne Ls.)

Metaluminae s-f. nov. (à base de métal Ls.)

Mixtusinae s-f. nov. (toutes les matières qui n’ont pas trouvé places plus haut)

Comme on peut le voir, la systématique de cette famille (nous dirons le fil conducteur) est basée sur la

composition de ia reproduction (fac-simili). Ceci a obligé l’auteur à plusieurs remaniements, surtout au niveau

des genres. Pas moins de 33 actuellement, avec plus de 200 espèces décrites. Contrairement aux idées reçues, la

composition n’est pas toujours évidente. Une révision est déjà envisagée au niveau des Ludoninae, trop

d’incohérence vis-à-vis des autres sous familles. II n’est pas exclu qu’elle disparaisse purement et simplement.

Chacun a pu également observer une nomenclature très recherchée. (pour les puristes 1). Ce fut

l’occasion pour l’auteur de faire preuve d’imagination dans les descriptions des nouvelles espèces. Excellent

exercice de relaxation qu’il recommande à tous.

NovaPEx / Société 3(1), 10 mars 2002

5. Annie Langleit : Les Psammobiidae (Superfamille: Tellinoidea -

Ordre: Veneroïda)

Bivalves très proches des Tellinidae, ils s’en distinguent généralement par une

charnière à nymphes épaisses, souvent proéminentes postérieurement aux sommets des

valves, sous le ligament externe, et aussi par une coquille non arquée à l’arrière. Ces

coquilles sont généralement inéquivalves et légèrement baiïllantes. Elles présentent 1 à

3 dents cardinales et des dents latérales faibles ou absentes. Le sinus palléal est assez

grand. À l’origine, ces bivalves ont été nommés comme Tellina, Solen ou même Venus.

On distingue 3 sous-familles : Psammobiinae, Sanguinolariinae et Novaculininae.

NovarEx / Société 3(1), 10 mars 2002 9

Les Psammobiinae sont ovales à trapézoïdales et comptent 5 genres récents : les Gari sont majoritaires

et divisées en plusieurs sous-genres, puis les Asaphis, les Ascitellina, les Heterodonax et les Orbicularia.

Les Sanguinolariinae sont ovales, souvent plus ou moins rostrées et comptent les genres Sanguinolaria

avec plusieurs sous-genres, puis les Nuffallia et les Soletellina ou Hiatula. Ce dernier genre pose des problèmes

de classement que chaque auteur essaie de résoudre à sa façon, ce qui ne simplifie pas nécessairement les choses.

Les Novaculininae comportent les genres Novaculina et Sinonovacula.

Le classement généralement suivi est celui de Myra Keen dans Moore repris et modifié, entre autres.

par Vaught en 1989 et par Millard en 1996. Vaught inclut les Solecutinae dans la famille des Psammobiidae,

Millard en refait une famille séparée.

10 Novarex / Société 3(1), 10 mars 2002

6. Ralph Duchamps : Les Heteropoda

ù Les /leteropoda étaient dans le passé nommés Aflantacea. Si nous voulons

ps = trouver leur place dans la systématique, nous dirons qu’ils appartiennent au Phylum des

CPR Mollusques, à la Classe des Gastéropodes, à la Sous-Classe des Prosobranches et à

4 | l'Ordre des Mesogastropodes. Mais cette superfamille à beaucoup voyagé dans la

SV. classification, à partir du moment où l’on a voulu construire l’arbre généalogique de ces

animaux. La sacro-sainte phylogénie des différents cladisticiens a transporté nos

4 mesogastropodes chez les Cypraeacea, les Naticacea, sans compter le groupe des

Echinospira. Pour corser le tout, les Æeteropoda ont été confondus avec les Pteropoda

(ou fusionnés), alors que ces derniers appartiennent à l’ordre des Thécosomes (Saccoglosses). Les Hétéropodes

vivent du niveau zéro à -1300m. Cette superfamille, se subdivise en 3 familles et 8 genres.

Ces gastéropodes sont bien adaptés à la nage et on en trouve dans toutes les mers du globe, toutefois

plusieurs genres ne vivent que dans les mers tropicales. Environ 60% des espèces connues possèdent une

coquille, mais 25% n’ont qu’une coquille de dimension réduite, et 15% n’ont pas de coquille du tout.

Dans la famille des Aflantidæ, toutes les espèces possèdent en principe une coquille dans laquelle

l’animal peut entièrement se rétracter et refermer l’ouverture grâce à un opercule. Les sexes sont séparés, le

développement des larves véligères varie selon les genres. Ceux-ci sont au nombre de trois.

Atlanta: 16 espèces, 3 sous-espèces. La coquille est dextre, pas entièrement planorboïde et mesure de

1,7 à 1 Imm, sa carène est cartilagineuse et se prolonge jusqu’à l’ouverture. L’opercule est ovale, et son nucleus

est situé sur la ligne médiane.

Protatlanta: 1 espèce qui mesure de 1 à 1,5mm. Les caractéristiques générales sont semblables au genre

précédent, toutefois la carène totalement transparente se continue jusqu’à l’ouverture et l’opercule est de forme

circulaire. La radula est légèrement différente de celle d’ Atlanta.

Oxygyrus: 1 espèce qui atteint 10mm. La coquille est planorboïde, cartilagineuse et transparente à l’état

adulte. Il en va de même pour la carène. L'ouverture est triangulaire, le sommet dirigé vers l’extérieur, tandis

que la base est concave et située vers le côté interne. L’opercule est de forme trapézoïdale, le nucleus ne se

trouve pas sur la ligne médiane.

La seconde famille appelée Carinariidæ comprend trois parties, la tête ou proboscis, le corps et la

masse viscérale. Cette dernière supporte à sa pointe extrême une coquille fragile qui ne peut contenir toutes les

parties molles. La famille se divise en trois genres.

Carinaria: 4 espèces, 6 formes qui possèdent les plus grandes coquilles de la superfamille. La plus

grande espèce a un corps qui peut atteindre 500mm pour une coquille de 85mm tandis que la plus petite a un

corps de 50mm.

Cardiapoda: 2 espèces dont la coquille est plus petite que chez Carinaria et insère une plus petite part

de la masse viscérale. La partie molle est prolongée par une sorte de queue courte se terminant en forme de

flèche, ou longue et filiforme, selon l’espèce. La longueur totale de l’animal est respctivement de 50 et 20mm.

Pterosoma: 1 espèce dont le corps est de forme ramassée, déprimée dorso-ventralement. Cet ensemble

disquoïde est terminé par une courte queue. La coquille est petite, enroulée et plate en forme de clou, dont la

carène relie le bord de l’ouverture à la coquille embryonnaire. L'animal mesure 80mm.

La troisième et dernière famille dénommée Prerotracheideæ est très comparable en forme et en

dimensions aux Carinariidæ (40 à 260mm). Toutefois elle s’en distingue par l’absence de coquille et un nucleus

visceral plus petit. Elle se subdivise en deux genres.

Pterotrachea: 4 espèces qui ne possèdent pas de tentacules antérieurs aux yeux et dont les femelles ne

sont pas munies de nageoire avec ventouse. Présence d’une queue bilobée postérieure au nucleus. Selon les

espèces la longueur varie de 50 à 260mm.

Firoloida: 1 espèce très commune de 40mm au corps long, cylindrique et transparent comportant un

proboscis frontal court et un nucleus viscéral globuleux à l’autre extrémité. L’espèce présente un dimorphisme

sexuel. Le mâle possède une nageoire avec ventouse, des tentacules implantés avant les yeux, un volumineux

pénis, une queue rudimentaire se terminant par un long filament. Le tout se trouvant sous le nucleus viscéral.

Chez la femelle, la queue est absente, seuls deux lobules obtus en forme de crochet sont présents et servent à

enfiler les œufs lors de la ponte.

Novapex / Société 3(1), 10 mars 2002 11

Inutile de dire que les coquilles des espèces de cette superfamille sont toutes fragiles, et souvent petites.

Les parties molles doivent être conservées en alcool, et tout examen microscopique nécessite fréquemment la

coloration préalable des organismes, ce qui facilite l'observation. En ce qui concerne la détermination des

espèces, les taxa principaux concernent la formule de la spire, l'examen de la radula, la grandeur et la forme de

l’opercule ainsi que son attache musculaire, l’examen des yeux et du nucleus viscéral, sans compter la

morphométrie.

Les Hétéropodes ne sont pas souvent présents dans les collections, et pourtant ils ne sont pas rares. Cela

provient du fait qu’ils n’ont pas un caractère esthétique et que l’observateur ne les imagine pas en tant que

mollusque.

Heteropoda

(Atlantacea)

Atlantidae Pterotracheidae

Carinaria

Protatlanta

Pterotrachea

Cardiapoda id

Le Museum d'Histoire naturelle de Paris possédait en 1819 un exemplaire de cette coquille, qui lui avait été

donnée par M. HUON et provenaïit de l’expédition d'ENTRECASTEAUX.

(Cfr. Lamarck : « Anim. s/vert. VII p.673 » 1822)

Le second exemplaire connu à Paris provenait de la vente de M. PIERRE LYONET qui fût acquis par CH. H.

HwAss

(Cfr. Cat. Raison. du célèbre cabinet de coquilles de Pierre Lyonet, publié à La Haye en1796)

NOVAPEX / Société 3(1). 10 mars 2002

lectiun #2

arinariid

NovaPEX / Société 3(1), 10 mars 2002 13

7. Claude Vilvens : Grands Trochidae et cladistique

Pour être honnête, je dois bien dire que je ne savais pas trop quoi exposer cette

fois : il me semblait avoir déjà bien exploité les familles qui m'intéressent, c'est-à-dire

les Turbinidae, Littorinidae et Cerithiidae (les Strombidae m'étant à présent interdits en

public ;-) — pas vrai, Etienne ?). Et tout naturellement, je suis donc revenu vers ma

famille de prédilection qui fait hurler de jalousie les Muricidae eux-mêmes : j'ai nommé

les Trochidae.

Première réflexion à leur sujet : un Trochidae, ce n'est pas grand. De fait, les

grands Trochidae sont assez peu nombreux. Tout le monde connaît bien sûr 7rochus

maculatus Linné, 1758 ou 7rochus niloticus Linné, 1767. Mais il en est d'autres assez grands, comme certains

Maurea ou certains Bathybembix. Evidemment, la notion de grande taille est relative : on peut être considéré

comme grand dans une sous-famille alors que l'on serait catalogué petit dans une autre. Etaient donc exposées les

espèces suivantes :

Tegula regina (Stearns, 1892) [Californie] Diloma tigrina (Anton, 1839) [Afrique du Sud]

Tegula (Chlorostoma) atra (Lesson, 1830) [Chili] Cittarium pica (Linné, 1758) [Guadeloupe]

Norrisia norrisi (Sowerby, 1838) [Californie] Cantharidus opalus (Martyn, 1784) [Nouvelle Zélande]

Turcica coreensis (Pease, 1860) [Japon] Tosatrochus attenuatus (Jonas, 1845) [Nouvelle Calédonie]

Ginebis argenteonitens (Lischke, 1872) [Japon] Calliostoma scotti Kiburn, 1973 [Afrique du Sud]

Lischkeia alwinae (Lischke, 1871) [Taïwan] Calliostoma formosense E.A. Smith, 1907 [Taïwan]

Bathybembix macdonaldi (Dall, 1890) [Chili] Maurea tigris (Gmelin, 1791) [Nouvelle Zélande]

Margarites ochotensis Philippi, 1846 [Russie] Pseudostomatella papyracea (Gmelin, 1791) [Philippines]

Gaza superba (Dall, 1881) [Cuba] Stomatia phymotis Helbling, 1779 [Nouvelle Calédonie]

Gaza olivacea Quinn, 1991 [Surinam] Solariella zaccaloides Schepman, 1908 [Philippines]

Trochus niloticus Linné, 1767 [Philippines] Ethalia guamensis (Quoy & Gaimard, 1834) [Philippines]

Tectus dentatus (Forskäl, 1775) [Mer Rouge] Umbonium giganteum (Lesson, 1831) [Japon]

Tectus pyramis (Born, 1778) [Japon] Bankivia fasciata (Menke, 1830) [Australie Sud]

Clanculus undatus (Lamarck, 1816) [Australie Ouest] | Monilea lentiginosa À. Adams, 1851 [Queensland]

La notion de sous-famille m'amène à la deuxième réflexion. Car, puisque nous parlons de sous-familles,

sans doute n'est-il pas inutile de repréciser celles qui constituent la famille des Trochidae. Il est surprenant, en

effet, de constater que nombre de revues malacologiques "grand public" semblent ignorer qu'une révision

systématique au niveau supra-générique a été réalisée il y a plus de10 ans par Hickman & Mc Lean. Cette

révision est basée sur l'ensemble des caractères taxonomiques, donc bien sûr ceux concernant la coquille, mais

aussi et surtout ceux relevant de l'anatomie interne et externe (branchies, epipodium, radula, tentacules, etc). Les

Turbinidae y sont à présent considérés comme plus primitifs que les Trochidae (on considérait plutôt l'inverse

jusqu'alors), les Skeneidae constituant la troisième et dernière famille la plus évoluée au sein des Trochoidea. Un

autre résultat systématique, exposé dans le tableau ci-dessous (avec quelques ajouts postérieurs à cette révision),

est la disparition de certaines sous-familles (comme les Monodontinae) et l'apparition de nouvelles (comme les

Lirulariinae), ou encore le passage à l'état de tribu (comme les Gibbulinae qui deviennent des Gibbulini, tribu de

la sous-famille des Trochinae) et la migration de certaines vers une autre famille (comme les Angarinae qui sont

classés dans les Turbinidae malgré leur opercule corné). Les mêmes auteurs proposent en fin d'ouvrage plusieurs

cladogrammes, dont l'un reflétant l'évolution des Trochidae les plus primitifs (les Tegulinae) vers les plus

évolués (les Umboniinae). J'ai reconstruit, pour l'occasion de l'exposition, le schéma de ce cladogramme avec les

coquilles citées plus haut comme exemples. On trouvera aussi dans les deux pages suivantes une liste des genres

et sous-genres Récents qui me sont connus chez les Trochidae, taxons classés par sous-familles et tribus.

! Hickman, C.S. & Mc Lean, JH. 1990. Systematic revision and suprageneric classification of trochacean

gasteropods. Natural History Museum of Los Angeles County Science Series. VI+169 pp.

NovapeEx / Société 3(1), 10 mars 2002

1. Tegulinae Kuroda, Habe and Oyama, 1971

Tegula Lesson, 1835

(Agathistoma) Olsson & Harbison, 1953

(Chlorostoma) Swainson, 1840

(Omphalius) Philippi, 1847

(Promartynia) Dall, 1909

(Stearnsium) Berry, 1958

Norrisia Bayle, 1880

2. Eucyclinae Koken, 1897

Agathodonta Cossman, 1918

Danilia Brusina, 1865

Dentistyla Dall, 1889

Euchelus Philippi, 1847

(Vaceuchelus) Iredale, 1929

Granata Cotton, 1957

Herpetopoma Pilsbry, 1889

Hybochelus Pilsbry, 1889

Mirachelus Woodring, 1928

Synaptocochlea Pilsbry, 1890

Tallorbis G.&H_.Nevill, 1869

Turcica A.Adams, 1854

Calliotropis Seguenza, 1903

(Solaricida) Dall, 1919

Bathybembix Crosse, 1893

Bathymargarites Waren & Bouchet, 1989

Cidarina Dall, 1909

?Echinogurges Quinn, 1979

Convexia Nodia, 1975

Ginebis Taki and Otuka, 1942

Lischkeia Fisher in Kiener, 1879

(Turcicula) Dall, 1881

Putzeysia Sulliotti, 1889

Vetulonia Waren & Bouchet, 1993

3. Margaritinae Stoliczka, 1868

(Cantharidoscops) Galkin, 1955

(Valvatella) Gray, 1857 = (Pupillaria)

Dall, 1909

Antimargarita Powell, 1951

Omphalomargarites Habe & Ito, 1965

Tibatrochus Nomura, 1940

Tropidomarga Powell, 1951

Gaza Watson, 1879

(Callogaza) Dall, 1881

Kaiparathina Laws, 1941

4. Trochinae Rafinesque, 1815

Trochus Linné, 1758

(Belangeria) Fischer, 1880

(Camelotrochus) Marshall, 1998

(Coelotrochus) Fischer, 1879

({nfundibulops) Piülsbry, 1889

(Infundibulum) Montfort, 1810

(Kanekotrochus) Habe, 1958

(Praecia) Gray, 1857

(Thorista) Iredale, 1915

(Thoristella) Yredale, 1915

Clanculus Montfort, 1810

(Camitia) Gray, 1847

(Clanculopsis) Monterosato, 1879

(Euclanculus) Cotton & Godfrey, 1934

(Euriclanculus) Cotton & Godfrey, 1934

({soclanculus) Cotton & Godfrey, 1934

(Macroclanculus) Cotton & Godfrey,

1934

(Mesoclanculus) redale, 1924

(Microclanculus Cotton & Godfrey, 1934

(Paraclanculus) Finlay, 1927)

Oligomeria Galkin & Golikov, 1985

Pseudotalopia Habe, 1961

Rubritrochus Beck, 1995

Tectus Montfort, 1810

(Cardinalia) Gray, 1847

(Rochia) Gray, 1857

Gibbula Risso, 1826

(Colliculus) Monterosato, 1888

(Forskalena) Kredale, 1818

(Phorcus) Risso, 1826

(Pseudodiloma) Cossman, 1888

(Steromphala) Gray, 1847

(Tumulus) Monterosato, 1888

Agagus Jousseaume, 1894

Austrocochlea Fischer, 1885

Cantharidella Pilsbry, 1889

Chrysostoma Swainson, 1840

Cittarium Philippi, 1847

Diloma Philippi, 1845

(Cavodiloma) Finlay, 1927

(Fractarmilla) Finlay, 1927

(Oxystele) Philippi, 1847

Enida À. Adams, 1860

Eurytrochus Fischer, 1880

Fossarina Adams & Anpgas, 1864

(Minopa) Iredale, 1924

(Clydonochilus) Fischer, 1890 =? Pagodatrochus

Margarella Thiele, 1893

Melagraphia Gray, 1847

Monodonta Lamarck, 1799

Nanula Thiele, 1921

Notogibbula Xredale, 1924

Osilinus Philipp, 1845

Pagodatrochus Herbert, 1989

Priotrochus Fischer, 1879

Cantharidus Montfort, 1810

({wakawatrochus,) Kuroda & Habe, 1954

(Mawhero) Marshall, 1998

(Plumbelenchus) Finlay, 1927

Alcyna À. Adams, 1860

Calthalotia Xredale, 1929

Calliotrochus Fischer, 1879

NovapreEx / Société 3(1), 10 mars 2002 15

Clelandella Winckworth, 1932 Bathymophila Dall, 1881

Jujubinus Monterosato, 1884 Ethaliopsis Schepman, 1909

(Manotrochus) Fischer, 1885 Ilanga Herbert, 1987

(Pictijubinus) Nordsieck, 1973 Lamellitrochus Quinn, 1991

(Mirulinus) Monterosato, 1917 Microgaza Dall, 1881

(Gravijubinus) Nordsieck, 1973 Minolia A. Adams, 1860

(Scrobiculinus) Monterosato, 1889 Minolops lredale, 1929

Kanekotrochus Habe, 1958 Spectamen lredale, 1924 (-Zeminolia Finlay, 1927)

Komaitrochus Kuroda & Taki, 1958 Suavotrochus Dall, 1924

Micrelenchus Finlay, 1927 Zetela Finlay, 1927

Phasianotrochus Fischer, 1885

Prothalotia Thiele, 1930 8. Halistylinae Keen, 1958

Thalotia Gray, 1847 Halistylus Dall, 1890

(Odontotrochus) Fischer, 1880 Botelloides Strand, 1928

Tosatrochus Mac Neil, 1960 Charisma Hedley, 1915

(Cavostella) Laseron, 1954

5. Stomatellinae Gray, 1840 (Cavotera) Laseron, 1954

Gena Gray, 1850 Fucaria Waren & Bouchet, 1993

Microtis A Adams, 1850

Pseudostomatella Thiele, 1924 9. Lirulariinae Hickman and Mc Lean, 1990

(Stomatolina) Iredale, 1937 Lirularia Dall, 1909

Stomatella Lamarck, 1816

Stomatia Helbling, 1779 10. Umboniinae Adams and Adams, 1854

Monileini Hickman and Mc Lean, 1990

6. Calliostomatinae Thiele, 1924 Monilea Swainson, 1840

Calliostoma Swainson, 1840 Antisolarium Finlay, 1827

(Alertalex) Dell, 1956 Camitia Adams & Adams, 1854

(Ampullotrochus) Monterosato, 1890 Conotalopia Iredale, 1929

(Benthastelena) Iredale, 1936 Ethalia Adams & Adams, 1854

(Elmerlinia) Clench & Turner, 1960 Ethaliella Püsbry, 1905

(Eucasta) Dall, 1889 Ethminolia Xredale, 1924

(Fautor) Iredale, 1924 Isanda Adams & Adams, 1854

(Kombologion) Clench & Turner, 1960 Parminolia \redale, 1929

(Laetifautor) Yredale, 1929 Rossiteria Brazier, 1895

(Otukaia) Ikebe, 1942 Talopena Xredale, 1918

(Salsipotens) Xredale, 1924 Vanitrochus Yredale, 1929

(Sinutor) Cotton & Godfrey, 1935 Zethalia Finlay, 1927

(Spikator) Cotton & Godfrey, 1935 Bankiviini Hickman and Mc Lean, 1990

(Tristichotrochus) Ykebe, 1942 Bankivia Krauss, 1848

Astele Swainson, 1855 Leiopyrga Adams & Adams, 1863

(Astelena) lredale, 1924 Umboniini Adams and Adams, 1854

(Coralastele) (redale, 1930) Umbonium Link, 1807

(Omphalotukaia) Yoshida, 1948 (Suchium) Makiyama, 1924

Bathyfautor Marshall, 1995 Inkaba Herbert, 1992

Callumbonella Thiele, 1924 Pseudominolia Herbert, 1992

Dactylastele Marshall, 1995

Fautrix Marshall, 1995 11. Cataeginae Mc Lean and Quinn, 1987

Falsimargarita Powell, 1951 Cataegis Mc Lean & Quinn, 1987

Maurea Oliver, 1926

Photinula H.Adams & A.Adams, 1854 12. Trochaclidinae Thiele, 1928

Photinastoma Powell, 1951 Trochaclis Thiele, 1912

Selastele Marshall, 1995 Acremodonta Marshall, 1983

Venustatrochus Powell, 1951

13. Thysanodontinae Marshall, 1988

7. Solariellinae Powell, 1951 Thysanodonta Marshall, 1988

Solariella Wood, 1842 (= Machaeroplax Carinastele Marshall, 1988

Friele, 1877) Herbertina Marshall, 1988

(Micropiliscus Dall, 1927)

Archiminolia \redale, 1929

16 NovaprEx / Société 3(1), 10 mars 2002

NovarEx / Société 3(1), 10 mars 2002

8. Rika et Fernand De Donder : Les ouvrages

malacolosiques concernant le Japon et les Philippines

Etaient ici exposés un nombre impressionnant d'ouvrages

traitant des mollusques de la zone géographique allant du Japon aux

Philippines. Les deux vedettes sont les deux récents ouvrages traitant

des coquilles du Japon :

Okutani, T. 2000. Marine Mollusks in Japan, Tokai University Press.

Tokyo, Japan, 1173 pp.

Higo, S., Callomon. P. & Goto, Y. 1999 + part Type figures 2001. Catalogue and bibliography of the Marine

Shell-Bearing Mollusca of Japan. Gastropoda. Bivalvia. Polyplacophora. Scaphopoda.

Il s'agit certes de références incontournables dans leur genre, mais Ô combien onéreuses (pour ne pas dire

monstrueusement coûteuses) et donc d'un accès difficile au malacologue prolétaire …

1. Morisla norisi / y A 314 (Calitornis). 2,3. Gars supebaz #9 75229 2 (Surinam). 4. Calhostoma monile= 1 # LEA (Aria Ses)

S. Lischhoia roginamaris 7 7 + © ZE A 214 {Holotype Philippines). 6. Akoys shinsyakaz 7 ZE 2 74 (Japan). 7. Ethakopsis kate D 7

29 #7 (Holotype. Japan). 8. Salsipotens nobilen UTC AN Tnatichotrochus scotti7 > 7 2 A 714 © {Sue Alrical

10. Maures solecta7 1) LEA M1 (New Zealand) 11. M. tignis + 5727742ZÉ2 71 (Neu Zealand}

FE NOVAPEX / Société 3(1), 10 mars 2002

9. Christiane Delongueville et Roland Scaillet :

Crassadoma pusio (Linnaeus, 1758

Crassadoma pusio (Linnaeus. 1758), plus connu sous le nom

de Chlamys (Hinnites) distorta (Da Costa, 1778), est un représentant

européen de la famille des Pectinidac. A l’état juvénile ce bivalve est

parfaitement symétrique. En grandissant, 1l se fixe par sa valve

inférieure (la droite) sur des substrats solides et prend dès lors les

formes les plus bizarres et les plus tourmentées. Des spécimens de

Crassadoma pusio incrustés sur différents substrats font l’objet de

cette exposition. [Substrats minéraux : granite, silex, basalte - substrats animaux : Bryozoaires comme Porella

compressa (J. Sowerby, 1805) et Pentapora fascialis (Pallas, 1766); Scléractiniaires comme Dendrophyllia

cornigera (Lamarck, 1816) ou encore sur d’autres mollusques comme par exemple des valves de Pecten

maximus (Linnaeus, 1758) - substrats végétaux : crampons de laminaires.]

Ulh

Distribution : de l'Islande et du Nord de la Norvège, en passant par les îles Britanniques, les côtes de Bretagne,

la péninsule ibérique, jusqu’au sud dans le golfe de Guinée, avec une incursion sur les côtes de la Méditerranée

voisines du détroit de Gibraltar.

PR TU

NovarEx / Société 3(1), 10 mars 2002 19

10. Rita Senders : "COQUILLES … ART"

Les artistes sont, sans cesse, à la recherche de nouveaux objets susceptibles

d'éveiller leur inspiration. Les coquillages ne sont-ils pas un matériel de choix tant par

leurs formes élégantes et variées que par l'évocation immédiate d'instant heureux, de

vacances insouciantes ?

C'est pourquoi, on découvre dans toutes les contrées touristiques des créations,

plus ou moins réussies, d'assemblages de ces petits hôtes marins. Je me suis contentée

de réunir et de présenter quelques parures et bijoux ramenés de nos séjours au bord de

mers proches ou lointaines. II s'agit le plus souvent de créations purement artisanales

mais où pointent parfois un véritable sens esthétique. A chacun le plaisir de jeter un œil simplement curieux ou,

pourquoi pas, un regard admiratif à ces "coquilles. art".

Bracelets :

- Fragments d'Haliofis sur argent. YUCATAN (Mexique).

Broches :

- Fragment d'Haliotis poli sur monture argent. NOUVELLE ZELANDE.

- Centre de Nautilus pompilius et feuillage argent. MIKONOS (Grèce).

Ceinture :

- Ceinture de nassaridae et sommets de cônes, sur bande de fibres végétales.

SUMBAVW A (Indonésie)

Colliers :

- Cypraea obvelata, Cypraea mauritiana et deux caputserpentis.

TAHITI.

- Naturelles

- Peints TAHITI.

- Centre et deux fragments de Nautilus pompilius cousus sur lien de fibres.

BALI.

- Trivias simplement enfilées. BALI.

- Umbonium vestiarum. VIETNAM.

- Trois fois trois Callista species sur gros lien de fibres. THAILANDE.

Couronnes :

- Coquillages terrestres. TAHITI.

- Trois rangs de Fragum fragum (Cardiidae). "Hereheretue« en polynésien.

TUAMOTU. Polynésie Française.

Ces couronnes posées sur les chapeaux d'hommes (elles remplacent les rubans des chapeaux de feutre de chez

nous) servent à les maintenir en place par leur poids. Les femmes mettent quelquefois une couronne légère faite

de mollusques terrestres directement sur leurs cheveux.

Pendentifs :

- Double face: deux patelles polies. INDONESIE.

- Valve d'Amusium cerclée d'or. JORDANIE.

- Ostreidae sur argent. PHILIPPINES.

- Double face: deux chitons polis et cerclés d'or. REPUBLIQUE DOMINICAINE.

- Coupes transversales de Strombe sur argent. PHILIPPINES.

NOVAPEX / Société 3(1), 10 mars 2002

npilius

- Centre de Nautilus

argent.

MIKONOS (Grèce).

Pompilius et fe

f :

NovapEx / Société 3(1), 10 mars 2002 21

11. Simone Maenhaut : La rétrospective

Notre artiste malacologue ne nous a pas, cette fois, proposé une œuvre

nouvelle mais, au contraire, nous a invité à nous souvenir de ses réalisations

successives créées à l'occasion des dernières expositions.

F' ie, vw}

nn. |

L CYSTISCIDAE Stimpson.,,, 1865

SOUS-FAMILLES :

Cystiscinae Stimpson.,,, 1865

Genres

Granulininae G.A. et HK. Coovert,., 1995

Genres

Persiculinae

Genres

GA. et B.K. Coovert,,, 1995

Plesiocystiscinae G.A. et H.K. Coovert,, 1995

Genre

IL MARGINELLIDAE J. Fleming,,, 1828

SOUS-FAMILLES :

Marginellinae J. Fleming,,, 1828

TRIBU : Marginellini J. Fleming,,, 1828

Genres

TRIBU : Austroginellini G.A. et B.K. Coovert,,, 1995

Genres

TRIBU : Prunini C.A. et H.K. Coovert,, 1995

Genres

Marginelloninae Coan,, 1965

Genres

Ces Ccystiscidae étaient représentés par :

NovarEx / Société 3(1), 10 mars 2002

12. Jeannine et René Masson : Les Marginelles

Jeannine et René Masson ont présenté les Marginelles en

“= deux familles : CYSTISCIDAE et MARGINELLIDAE, selon la

scission opérée par G.A et H.K. COOVERT. Les deux familles ainsi

créées comportent plusieurs sous-familles, tribus et genres.

Cystiscus Simpson, 1865

Crithe Gould, 1860

Extra Jousseaume, 1894

Granulina Jousseaume, 1888

Pugnus Hedley, 1896

Persicula Schumacher, 1817

Canalispira Jousseaume, 1875

Gibberula Swainson, 1840

Plesiocystiscus G.A. et HK. Coovert, 1995

Marginella Lamarck, 1799

Dentimargo Cossmann, 1899

Eratoidea Weïnkauff, 1879

Clabella Swainson, 1840

Austroginella Laseron, 1957

Alaginella Laseron, 1957

Balanetta Jousseaume, 1875

Closia JE. Gray, 1857

Hydroginella Laseron, 1957

Mesoginella Laseron, 1957

Ovaginella Laseron, 1957

Protoginella Laseron, 1957

Serrata Jousseaume, 1875

Serrataginella G.A. et H.K. Coovert, 1995

Prunum Herrmannsen, 1852

Bullata Jousseaume, 1875

Cryptospira Hinds, 1844

Hyalina Schumacher, 1817

Rivomarginella Brandt, 1968 (eau douce)

Volvarina Hinds, 1844

Marginnellona von Martens, 1904

Afrivoluta Tomlin, 1947

NovapEx / Société 3(1), 10 mars 2002 23

Persicula blanda (Afrique de l'Ouest)

Persicula accola (Panama)

Persicula cingulata (Sénégal)

Persicula imbricata (Mexique)

Persicula persicula (Sénégal)

Persicula avellana (Afrique de l'Ouest)

Persicula pulchella (W. Australia)

Persicula interruptolineata (Vénézuéla)

Gibberula chudeaui (Sénégal)

Granulina occulta (Afrique de l'Ouest)

Les coquilles étaient disposées de manière telle que l'on pouvait repérer les 10 Cystiscidae (marqués

d'un point rouge) parmi une centaine de la grande ex-famille. Dans l'ensemble Marginellidae et Cystiscidae

confondus, nous avons pu observer des coquilles de tailles diverses allant de 1 mm à 50 mm. La plus grande

espèce mesure 150 mm et provient de la mer de Chine (Marginellona gigas (von Martens,, 1903)). Le deuxième

plus grand spécimen connu de 120 mm se rencontre sur la côte de l'Afrique du Sud (4frivoluta pringlei (Tomlin,

1947)). Bullata bullata (Born, 1778) du Brésil (100 mm) est le troisième plus grand.

Une photocopie couleur reproduisait les 10 nouvelles espèces attribuées à 5 genres différents du niveau

bathyal de Nouvelle Calédonie, décrites récemment par Franck Boyer.

Nous savons que la plupart de ces animaux se nourrissent surtout d'autres mollusques. Quelques espèces

peuvent percer la coquille d'autres bêtes puis aspirer à l'aide de leur trompe le tissu de leur victime par l'orifice

qu'elles ont créé.

été (1), 10 mars 2002

NOVAPEX / S

haine — EXPO

année proc

Rendez-vous

NovAPEx / Société 3(1), 10 mars 2002 23

Une récolte éclair en Espagne

Etienne MEULEMAN

La plage de Vilanova ila geltrü, à une quarantaine de kilomètres de Barcelone, ne s’attendait certainement pas à

voir débarquer, un 19 décembre en fin d’après-midi, un collectionneur de coquillages. En effet, les coquilles

rejetées sur la plage n’ont pas eu le temps de se cacher.

Quel bonheur, après une journée de dur labeur de se retrouver sur la plage au coucher du soleil ! Le bonheur est

d’autant plus grand lorsque celle-ci est remplie de coquilles (photo 1). Sur une heure de temps quelques 35

espèces différentes récoltées. Il suffisait de se baisser pour remplir son sac de trésors fraîchement rejetés.

La récolte a commencé lorsque j’ai buté contre une natice, Naficarius cruentatus (Martyn, 1784). Vu le temps

dont je disposais, j’aurais déjà été heureux de ne trouver que cette coquille. Mais plus je me rapprochais de la

mer, plus les coquilles étaient nombreuses, un Murex à gauche, une Telline à droite, une Natice, deux Natices,

trois Natices, dix Natices, je n’en croyais pas mes yeux.

Même les Bernard-l’ermite étonnés ont accepté de poser pour la photo (photo 2).

La récolte s’est terminée lorsque le soleil s’est caché et que la lumière ne me permettait plus de distinguer les

coquilles sur la plage. Bref, c’est heureux et les poches pleines de coquilles que je suis rentré à l’hôtel.

Le soir en m’endormant, je me suis dit que la nature pouvait nous réserver encore bien des surprises, et qu'il

était encore possible de trouver de beaux spécimens pas trop loin de chez soi.

Ci-dessous vous trouverez la liste des espèces récoltées sur la plage :

Gastéropodes

Famille: APORRHAIDAE Famille: NASSARIIDAE

Aporrhais pespelecani (Linné, 1758) Nassarius mutabilis (Linné, 1758)

Nassarius reticulatus (Linné, 1758)

Famille: BUCCINIDAE

Buccinulum corneum (Linné, 1758) Famille: NATICIDAE

Naticarius cruentatus (Martyn, 1784)

Famille: CANCELLARIIDAE Naticarius punctatus (Chemnitz in Karsten, 1798)

Cancellaria cancellata (Linné, 1767) Neverita josephina Risso, 1826

Famille: CASSIDAE Famille: PATELLIDAE

Phalium undulatum (Gmelin, 1791) Patella caerulea Linné, 1758

(coquille abîimée)

Famille: THAIDIDAE

Famille: CERITHIDAE Thais haemastoma (Linné, 1758)

Cerithium vulgatum (Bruguière, 1792) (coquille abîmée)

Famille: CREPIDULIDAE Famille: TROCHIDAE

Crepidula unguiformis Lamarck, 1822 Calliostoma zizyphinum (Linné, 1758)

Gibbula magus (Linné, 1767)

Famille: MURICIDAE

Bolinus brandaris (Linné, 1758)

Hexaplex trunculus (Linné, 1758)

Bivalves

Famille: ANOMTIIDAE

Inomia ephippium Linné, 1758

Famille: ARCIDAE

Arca noae Linné, 1758

Famille: CARDIIDAE

Acanthocardia tuberculata (Linné, 17538)

Famille: CARDITIDAE

Cardites antiquata (Linné, 1758)

Famille: GLYCYMERIDIDAE

Glycymeris insurbica (Brocchi, 1814)

Famille: MACTRIDAE

Mactra stultorum (Linné , 1758)

Famille: MYTILIDAE

Mytilus edulis Linné, 1758

Coquilles terrestres

(aux abords de l’hôtel)

Famille: SUBULINIDAE

Rumina decollata (Linné, 1758)

NovapEx / Société 3(1), 10 mars 2002

Famille: OSTREIDAE

Ostrea edulis Linné, 1758

Famille PECTINIDAE

Pecten jacobeus (Linné, 1758)

(des valves isolées)

Famille SPONDYLIDAE

Spondylus gaederopus Linné, 1758

(des valves isolées)

Famille TELLINIDAE

Tellina planata Linné, 1758

Tellina nitida Poli, 1795

Famille: VENERIDAE

Callista chione (Linné, 1758)

Dosinia lupinus (Linné, 1758)

Famille: HELICIDAE

Theba pisana (Müller, 1774)

Cernuella virgata (da Costa, 1778)

Eobania vermiculata (Müller, 1774)

Photo n° 1 : La plage de Vilanova ila Geltrü au coucher du soleil.

NovapEx / Société 3(1), 10 mars 2002 27

Photo N° 2 : Un Bernard-l’ermite dans Bolinus brandaris (Linné, 1758) qui a accepté de poser pour la photo.

Quelques espèces récoltées sur la plage:

En haut à gauche: Bolinus brandaris (Linné, 1758)

En haut au centre : Cancellaria cancellaia (Linné, 1767)

En haut à droite : Hexaplex trunculus (Linné, 1758)

En bas à gauche: Callista chione (Linné, 1758)

En bas à droite: Naticarius punctatus (Chemnitz in Karsten, 1798) et Neverita josephina Risso, 1826

28 NOVAPEX / Société 3(1), 10 mars 2002

Quoi de neuf ?

Roland HOUART et Claude VILVENS

EXPOSITION DE LA SOCIÉTÉ BELGE DE MALACOLOGIE A

GENVAL

En octobre 2001 nous avons reçu une demande du responsable du Musée de l'Eau et

de la Fontaine à Genval en vue d'organiser une exposition de coquillages au sein de

leur week-end "fête de l'eau" les 29, 30, 31 mars et ler avril 2002.

Leur exposition/animation rassemblera, dans le cadre et le prolongement de la

Journée Mondiale de l'Eau (le WE des 23 et 24 mars) des associations ayant des

projets liés à l'eau, des spécialistes du domaine de l'eau, des animations sur le thème de

l'eau, des expositions et des stands sur les activités culturelles ou ludiques liées à l'eau

(plongée, maquettisme, malacologie, etc.)

Nous serons présents les deux week-end, soit:

- Journée mondiale de l'Eau: WE des 23 et 24 mars de 10 à 18 heures

- Fête de l'Eau: (WE de Pâques), les 30, 31 mars et le 1° avril, également de 10 à

18 heures

Nous y exposerons diverses jolies pièces et des vitrines contenant les mollusques d'eau douce de Belgique, les

mollusques d'eau douce du lac Moero, des coquillages exotiques, etc. D'autres réalisations tels des panneaux

didactiques y seront aussi présentées

M. et Mme Senders y exposeront également des photographies sous-marines.

La Société Belge de Malacologie aura son stand (pas très loin de l'entrée) avec une permanence assurée au moins

durant le week-end de Pâques.

Soyez nombreux à venir nous rendre une petite visite!

Situé dans un environnement pittoresque, près du lac de Genval, le Musée de l'Eau et de la Fontaine vaut

vraiment le détour!

ENTRÉE: adultes: 3.00 €; enfants de 6 à 12 ans: 1,50 €; moins de 6 ans: gratuit.

Voies d'accès: VOIE FERREE

Ligne Ottignies-Louvain-La-Neuve-Bruxelles

Ligne Bruxelles-Ottignies-Louvain-La-Neuve

Descendre à la gare de Genval

RESEAU ROUTIER: notamment E411

De Bruxelles

sortie n° 3 Genval-Terhulpen-Overijse (ou)

sortie n° 4 Rosières

De Namur

sortie n° 4 Rosières (ou)

sortie n° 5 rixensart-Bierges

Suivre les indications "Lac de Genval" ou "Musée de l'Eau et de la Fontaine"

Musée de l'Eau et de la Fontaine

l Avenue Hoover, 63 — B-1332 Genval — Belgique

www.pixelsbw.com/musee-eau-fontaine

(Ouvert les week-ends et jours fériés de 10h à 18h

ou sur rendez-vous en semaine (02/654.19.23) )

NoOVAPEX / Société 3(1), 10 mars 2002 29

#Æ Vis RE À Quelques nouvelles publications

S 4 Roland HOUART

1. Quelques livres

——

THE MOLLUSCS OF THE SOUTHERN

GULEF OF THAILAND

par G. Swennen, R.G. Moolenbeek, N. Ruttanadakul,

H. Hobbelinck, H. Dekker & S. Hajisamae

pp. 1-ix, 1-210, 530 figures (photographies en couleur) Publié et distribué par "the Biodiversity Research and

Prix: 25 USD (couverture souple); Training Program (BRT)", Bangkok, Thailande.

40 USD (couverture cartonnée).

Encore un livre en plus me direz-vous, oui mais... un conseil,

découvrez-le vite, il en vaut la peine.

L'introduction comprend plusieurs chapitres très intéressants

telle l'exploration de la région étudiée, une vue d'ensemble des

mollusques locaux et de leurs habitats, les espèces introduites

récemment, la systématique (taxonomie et nomenclature,

classification, etc.), écologie et historique (locomotion, nourriture,

croissance, couleur...), les mollusques et les humains. Les premières

figures du livre nous montrent une carte de la région explorée, des

pontes et diverses photos tirées de la vie locale en rapport direct avec

an les mollusques et les coquillages.

The Molluses of the Les auteurs nous emmènent ensuite vers les divers sites, 1ls

Southerà Gülf of Thailand nous décrivent aussi les différentes façons de nettoyer, d'étiqueter, de

1 récolter et d'identifier des mollusques et d'entreposer une collection.

qi La partie systématique débute par une liste des espèces, chacune d'elle

est ensuite décrite: genre, espèce, auteur(s), date de description, courte

diagnose et distribution dans la région explorée. Quarante-quatre

|| planches couleurs de très bonnes qualité illustrant polyplacophores,

| FE RobaIrR. 1. Der an Same bivalves, dentales, céphalopodes, gastéropodes précèdent un glossaire

et l'index final.

Sans hésitation, c'est un livre que je conseille à tous nos

membres ! Les descriptions sont explicites, les photos sont très bonnes et... le prix du livre très démocratique.

À acheter immédiatement pour inclure dans toute bonne bibliothèque.

A CONCHOLOGICAL ICONOGRAPHY

1. A Taxonomic Introduction to the Recent VOLUTIDAE

par Patrice Bail et Guido T. Poppe

2. Genus Amoria

par Patrice Bail et Allan Limpus

Publié et distribué par ConchBooks

Mainzer Str. 25, D-55546 Hackenheim, Allemagne.

conchbooks(@conchbooks.de

Tout le monde connaît à présent cette série fabuleuse qui petit à petit s'agrandit pour, on l'espère, former

une série de monographies absolument indispensables à toute bibliothèque.

30 NOVAPEX / Société 3(1), 10 mars 2002

Les derniers nés sont écris par des professionnels en la matière. Texte, dessins et photographies (couleur

evidemment) sont à la hauteur de ce que l'on attend de ces spécialistes: du beau, du bon et de l'indispensable.

"A taxonomic introduction of Volutidae" nous conduit vers une vue d'ensemble de la famille et des

différents genres. Chaque (sous)genre est briévement décrit et est accompagné de la liste des espèces. Les

espèces types sont indiquées et sont illustrées à l'aide d'excellentes photographies couleurs sur 5 planches.

Dans "The genus Amoria" par P. Bail et A. Limpus, en guise d'introduction, les auteurs nous brossent

d'abord quelques traits importants sur la position systématique, les sous-familles proches, les sous-genres, les

fossiles, la distribution et l'habitat, la morphologie de la coquille, et d'autres points.

Chaque espèce est ensuite décrite avec localisation du matériel type, la distribution géographique,

l'habitat et quelques remarques. Chaque description est accompagnée d'une photographie noir et blanc de

l'espèce et de la localisation de l'espèce sur une carte géographique. Le volume se termine par une courte

bibliographie, un index et 93 splendides planches couleurs où chaque espèces est illustrées à l'aide de nombreux

spécimens, montrant ainsi la variabilité (infinie) des coquilles. Les planches 85 à 93 sont consarées aux

photographies de quelques espèces in situ.

Les fêtes de fin d'année sont déjà (ou encore) bien lointaines mais il n'est pas toujours nécessaire

d'attendre un événement heureux pour s'offrir (ou se faire offrir) LE CADEAU (the must...).

Bonne lecture!

4 ! i de

Genÿs Amoria

Edited by ConchBooks Edited by ConchBooks

2. Quelques publications

Pour rappel, il s'agit ici de publications ne se trouvant à la bibliothèque de la SBM, mais

qu'il est possible de consulter à l'IRSNB et le plus souvent à l'ULB. On peut consulter

Roland Houart à ce sujet.

Moluscos del Plioceno Superior Marino de Isla Guafo, Sur De Chile, Parte Il. Gastropoda, par D. Frassinetti.

Boletin del Museo Nacional de Historia Natural, Chile, 49: 131-161 (2000).

L'auteur décrit les gastéropodes marins fossiles de l'Ile Guafo au Sud du Chili. Dix-huit espèces sont recensées

dont 8 nouvelles: Trophon covacevichi, T. huilliche, T. parcus, T. vetulus, Phos chilensis, Penion crassus,

Penion diversum et Adelomelon reconditus.

Catalog of Recent type specimens in the division of invertebrate zoology, American Museum of Natural History.

V. Mollusca, Part 2 (class Gastropoda [exclusive of Opisthobranchia and pulmonata], with supplements to

Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History, n° 262,

2001.

NOVAPEX / Société 3(1), 10 mars 2002 31

Je pense utile de reproduire ici le résumé:

À complete, annotated listing of all primary and secondary type specimens of gastropod mollusks (exclusive of

opisthobranchs and pulmonates) in the collection of the Division of Invertebrate Zoology at the American

Museum of Natural History (AMNH) (as of December 31, 1999) is presented. Supplementary listings for

bivalves and opisthobranch gastropods (previously covered in Part 1) are also given. This catalog includes 1309

type lots, 18 lectotype designations, and illustrations for 12 previously inadequately illustrated type specimens

and five species never before illustrated. New synonymy information is given for nine species, one species is

shown to be a valid name, one new replacement name is introduced, and one neotype is suppressed with

rediscovery of type material. An appendix of AMNH specimens previously incorrectly cited as types is also

given. An index of genera and species is included for each class and subclass.

Contribution to the marine molluscan fauna of Kerguelen Islands, South Indian Ocean, par N. Troncoso, J.L.

Van Goethem and J.S. Troncoso. /berus 19 (1): 83-114 (2001).

Résumé original:

The present work contributes to the knowledge of the mollusc fauna of the Kerguelen Is., on the basis of a

collection of the Institut Royal des Sciences naturelles de Belgique. This collection include 32 species of

gastropods and 12 of bivalves collected in shallow waters of the Morbihan Bay, among the gastropods

Margarites cf. porcellana and Perissodonta mirabilis are the most abundant, whereas among the bivalves the

commonest species are Gaimardia trapesina and Laternula elliptica. Most of the species in this collection have

a wide distribution, although some species are endemics of Kerguelen Is. or of the Kerguelen-Heard platform

and another species circumantartic.

LCL

Proceedings of the 10" Congress & Workshop Tropical Marine Molluse Programme (TMMP). Ministry of

Fisheries, Vietnam. Hanoi & Halong Bay 20-30 October 1999. Phuket Marine Biological Center Special

Publication n° 21: i-xû, 1-316 (part 1), 317-537 (part 2), 539-644 (part 3).

NovAPrEXx / Société 3(1), 10 mars 2002

TABLE OF CONTENTS

Part 1

Jorgen Hylleberg: Joyce Dangebun’Inmemoriam.. "M rise rer nes de

Co:operating'institutionséss fs fier ee RS EN ER RER

Table: of contents’: emtiomee ARR ee 2 ASE RER UT El

Jorgen Hylleberg: General conclusions regarding the 10th TMMP Congress & Workshop

ADDRESSES

Ho Van Hoanh Chairman of the Organising Committee: Welcome address .…........................

Jorgen Hylleberg Programme Director of the Tropical Marine Mollusc Programme:

Outlook on the occasion of the 10th TMMP Congress & Workshop ..….................................

Nguyen Thi Hong MinhVice Minister for Fisheries, Socialist Republic of Vietnam:

Opening addré Rev nee ER RS De

ECOLOGY and ENVIRONMENT

Ib Svane & Stephen J. Hall: Stable isotope analysis to determine trophic relationships

in the Spencer GUulEe SOUTIAUS TANIA ARR

Pitiwong Tantichodok: Detritus as a food source in marine ecosystems: An overview of

reSéarch in the TOBDS SRE ARE Re" à LE RE

Ing Try: Mollusc Species in the diets of dab (Limanda limanda), flounder (Platichthys

flesus) and plaice (Pleuronectes platessa) in Aarhus Bay, Denmark

Nur Taufiq SPJ & Retno Hartati: Relationships beween organic matter in sediment and

abundance, condition index, and growth of cockle (Anadara granosa L.). in three

IndoneS An eSATA à nee à à ee nn nn eee eee

Jintana Nugranad, Supot Chantrapornsilp & Thani Varapibal: Feeding and spaw-

ning behaviour of the trumpet triton, Charonia tritonis (L., 1758) in captivity..............….

Le Duc Minh: Reproductive biology of Haliotis varia Linné, 1758 in Cam Ranh Bay,

South Central Wietname "222.

Sharifuddin Bin Andy Omar & Fredinan Yulianda: Distribution of Anadara granosa

(L.) atRawameneng Beach, Subang, West Java, Indonesia .…...............................

Surapong Banchongmanee & Tipamat Upanoï: Giant clams in the Andaman Sea,

Thailand. Part 1: Distribution and abundance at Rawi, Butang, and Kata Islands...

Tipamat Upanoi & Surapong Banchongmanee: Giant Clams in the Andaman Sea,

Thaïland. Part 2: Distribution and abundance at Adang, Bitsi, Hing-Ngam, and Ta

Lang Islands tnt tie he RSR A A ER AE

Mickmin Charuchinda & Panya Asawangkune: Nursing of giant clam Tridacna

SGUAMOSCANCABÉS". NTM AE ser innenee es en AIR 2e ec a D SLR Eee eee

G. J. Fontje Kaligis: Distribution in relation to environment of the marine snail

Littorina littorea (Gastropoda:Prosobranchia) atHelgoland (North Sea)...

Jaruwat Nabhitabhata & Pitiporn Nilaphat: Behaviour of juvenile cephalopods:

preference for texture and brightness of substrata.....…............................. PR

Zulfigar Yasin & Aileen Tan Shau-Hwaïi: Quantitative and qualitative effects of light

on the distribution of giant clams at the Johore Islands in South China Sea...

Aïleen Tan Shau-Hwai & Zulfigar Yasin: Photoacclimation responses by zooxanthallae

ofgiantclam Fridaenarsquamosates 2 Le Jui RES ERRE PR ETRNT ISERE EUR

Ake Granmo: Effects of organotin on marine bivalves

Michael Bech: Acute toxicity of tributyltin to the larvae of Thais bitubercularis

Sunan Yuaycharoen, Wanloap Kumsupa & Sunit Pattapong: Heavy metal residues

in razor clams, sea water, and sediment in coastal areas of Samut Songkhram

Province, Thailand

Novarex / Société 3(1), 10 mars 2002

Retno Hartati & Jusup Suprijanto: Quantitative and qualitative studies on bio-

accumulation of lead (Pb) in cockles (Anadara inflata Reeve)

Isdradjad Setyobudiandi: Abundance and biomass of gastropods and bivalves in a warm

APE MPOIULedarenorCIesOn, WESt JAVA. se eee Locmtes ceutanerenesee ocre ipe ice re iorritenc

Yusli Wardiatno, Majariana Krisanti & Poppi L. Wahjuhardini: Occurrence of

bacteria in cockles, Anadara granosa Linné in Jakarta Bay, Indonesia ...........................

À. Murugan & J. K. Patterson Edward: Factors threatening biodiversity of marine

mollraanluancanaCuton Enr MR ne

Ngo Anh Tuan: Note on distribution and exploitation of the babylon snaïl Babylonia

ARCOLALAO NP BTE LIU ANS VITE ET AT 22.222212 === asus eee eds e ep eee se see ec tscecsenn=s ces een e sen ens

AQUACULTURE

Vo Si Tuan: Status and solutions for farming and management of the clam Meretrix

MratatGolConsaDone Ment(Ciane Pro MINCE, VIe ORAN

Jocelyn A. Madrones-Ladja & Wenresti Gallardo: Restocking of window-pane shell,

Placuna placenta in a depleted bed off Tigbauan, Iloilo, The Philippines

Nguyen Huu Phung: Distribution and yield of commercial gastropods and bivalves

(Mollusca) in coastal waters of Vietnam ..…................................... PRES DR RER nl PACE

Fredinan Yulianda & Agus Soleh Atmadipura: Growth of oyster Crassostrea sp.

setrlecionishellandstoneltcollectorsie 2e ne enr A ee Ne nt

Retno Hartati & Chrisna A. Suryono: Oyster spatfall at Mlonggo waters, Jepara,

OO en Mi ee fit (4

Delianis Pringgenies, M. Murdjani, S. Soelaksono & Noorsalam R. Nganro: A first

attempt at culturing the squidLoligo duvaucelii Orbigny, 1835 in Indonesia...

Jocelyn A. Madrones-Ladja & M. R. de la PeOa: Hatchery management for the window-

paneshelPlocura placenta linnaeuss 1758422. Mousse. Demi paie. 5" Tru.

Armando C. Fermin, Rolando S.J. Gapasin & Myrna B. Teruel: Spontaneous

spawning, fecundity and spawning periodicity in the donkey's ear abalone Haliotis

CARO Iron cocon poor oo ont tel AE sen ER RSR

Jintana Nugranad, Siriya Noodang, Wichulada Ratanachurdchai, Kanchanee

Promchinda, Sompit Punna, & Supatcha Chantara: Breeding of the oriental hard

Cam ererneomrene tr ET ES)

Nguyen Van Chung: Note on culture of abalone Haliotis ovina (Gmelin) in Khanh Hoa

PEOVIMCeN IE ERA RE AR M en Men ot mb den ete Len e le RU

Nguyen Thi Bich Ngoc: Culture and maintenance of microalgae for mollusc larviculture.…

Nur Taufiq SPJ & Retno Hartati & Endang S. Susilo: Ecology, morphology and the use

CUpanen MAUSCLLISLA SDL IMAONESTAE eee erreroereeenr ce eue eoeeenussc ans aroemasaonnru sep anenaumegntecerve sance

Gunarto Latama: Growth and survival of juvenile giant clam Tridacna squamosa as a

OUT CHID AA O ETS LE ee re nee uen ee Dee ou es ne do dt de. meiout AU. Lie

Nguyen Chinh & Nguyen Thi Xuan Thu: Status of biodiversity and aquaculture of

marine molluses in Vietnam - the present scenario and future prospects

Armando C. Fermin & Rolando S.J. Gapasin: Postlarvae density and photoperiod

effects on the settlement and metamorphosis of the donkey's ear Abalone, Haliotis

CE De GE RE Re cannot ou avan dau duree

Hua Thai Tuyen & Vo Si Tuan: Growth of the silver-lip pearl oyster restored in the

waters around Cu Lao Cau Island, Binh Thuan Province, Vietnam

Nguyen Trong Nho & Ngo Anh Tuan: Note on biological characteristics and resources of

the scallop Chlamys nobilis in the coastal areas of Binh Thuan Province...

195

203

NoOVAPEX / Société 3(1), 10 mars 2002

Fredinan Yulianda & Edward Danakusumah: Growth and gonad development of

babylon snail Babyolnia spirata (L.) in culture...

Tanate Poomtong, Jintana Nugranad & Wichulada Ratanachurdchai: Mass

Transportation of Live Marine Molluses: Case stories on success and failure.

POSTHARVESTING

J. K. Patterson Edward & A. Murugan: Screening of cephalopods for bio-activity.…......…

Jamila Patterson: Utilization of gastropod meat for the preparation of flakes..…............

ABSTRACTS

Azanza, Rhodora V., Maria Patricia V. Azanza & Arlynn I. Gedaria: Ultraviolet-

assisted PSP toxin and microbial depuration of Perna viridis mu

Bautista-Teruel, Myrna, Oseni M. Millamena & Armando Fermin: Broodstock nutri-

tion of abalone, Haliotis asinina, Linne: Effects of diet on reproductive performance...

Benzie, John A. H.: Genetic tools for assessing diversity: Patterns of genetic variation

A DATINE ED AUISENS en Ne

Chau Thanh, Nguyen Chinh, & Tran Mai Kim Hoa: Reproductive biology of green

musselPerné, vins (Lines 158) a ere Ce le RES

Ha Le Thi Loc: Reproductive biology of the black lip pearl oyster Pinctada margaritifera

(Linne, 1758) in Nha Trang - Khanh Ho Vietnam St PEU A NEA AT SPA

Kastoro, Woro W. & Mudjiono: Bivalvia collected from the islands of Sunda Straits.

Kilburn, R. N.: Biogeography, Biomes and the Mollusca of South East AfipAr. same F2

Kittiwattanawong, Kongkiat: Correlations of multilocus heterozygosity to growth rate,

oxygen consumption, and morphological characteristics in Cerastoderma edule

Kohn, Alan: Molluscan Castle-Building: How molluses make their shells.….......

Mamangkey, Gustaf: Shipworms (Bivalvia: Teredinidae) in the Indo-Pacific region:

Places where thèv'are well dispersed ONE Re

Mantiri, Desy M. H. & Farnis B. Boneka: Pigments of a polymorphic species, Littoraria

pallescens (Prosobranchia-Bittorimdae) INSEE NOR

261

201

Novaprex / Société 3(1), 10 mars 2002

Nguyen Chinh: General features of the research work and handling of molluscan

resources in Vietnam and the orientation of development ..…...........................................

Nguyen Chinh: Morphological Characteristics and geographical distribution of the

commercial bivalve species Tegillarca granosa and T. nodifera in Vietnam

Nguyen Chinh, Nguyen Van Hung & Phung Bay: Results of vellow-lipped pearl oyster

Pinctada maxima artificial seed breeding experiments in Van Ninh Sea waters, Khanh

Éoa Érovince and'inViune Ro Bay, PDU Yen PTOVINCE nr rienemnnssessennnie nn ose

Nguyen Huy Dien: Distribution of some economic bivalve species in intertidal areas in

Nene nm Ie, Le. PU, NOR EI A RNCS CRU bee Er Te enter ttles se

Nguyen Thi Xuan Thu, Hua Ngoc Phuc, Phan Dang Hung, Mai Duy Minh, & Kieu

Tien Yen: Biological spawning characteristics of babylon snail, Babylonia areolata..….……

Nguyen Thi Xuan Thu, Hua Ngoc Phuc, Hoang Thi Bich Dao & Thai Ngoc Tran:

Effect of salinity on hatching, larval growth and survival of babylon snail, Babylonia

CRAOUCIC nebasooosco oups benne de dé ASIE LE LE ES er

Nguyen Van Hao, Nguyen Dinh Hung, Pham Cong Thanh, Tran Quang Minh &

Nguyen Thanh Tung: Environmental parameters, biology and stocks of Meretrix lyrata

In REMeRGDe TAN Ron Ame AM CAR RM ent ee dance apennas sente sen ne

Nguyen Xuan Duc: Species composition and distribution of cephalopods in Vietnam

Pripanapong, Suparp: Cement pole culture of oyster (Crassostrea belcheri) using

VaTousiSIZeS 0H hAtCheERV=-PrOdUCE NS DA AE PRE IE TR Re.

Singhagraiwan, Tanin, Vicharn Ingsrisawang, Saowanee Singhagraiwan &

Somnuk Kabinrum: Biodiversity of molluses on artificial reefs in Phe Bay, eastern

Coastotne CulTo EN aAIANA 2.82... DOMAINES EPMIONS CAR RER ECTANTEREE Le APE

Soekendarsi, Eddy, M.Iqbal Djawad & Yulianus Pangaongan: Growth rate of Trochus

HIIOUICUS TE Nath eMmphasis On MacrO-alpal fO0d ITEMS"... soe dede creer seen

Strack, Hermann Leberecht: Malacological results of the Rumphius biohistorical

ÉXPECIONLOrAMOONMOLOOO) ER RL nn de ann sn ea ane ea:

Sunoto, Hadi, Ninis Trysjani, & Rahmadi Prasetyo: The commercial aspects of harve-

sting Solen grandis in the coastal waters of East Surabaya, Indonesia .…............................

Tan, K.S. & Y.Y. Loo: Molluscan species identification using artificial neural networks...

Tandavanitj, Sanchaï: Feasibility study on culture of cockle Anadara troscheli Dunker…….

Thorarinsdottir, Gudrun G: Size and age at sexual maturity and annual gametogenic

cycle in ocean quahog, Arctica islandica (Linnaeus, 1767), off North-West Iceland

THE TENTH CONGRESS & WORKSHOP CONCLUDED

Jorgen Hylleberg: 107" Congress & Workshop Concluded

SL OR ACTA NT RE AT CR Mr RM een de PAM MENU. REA UE hs

Jorgen Hylleberg: The Tropical Marine Mollusc Programme (TMMP) activities during

DESERT 0 RE RER MER ET IR RE OR ER RAGE TS ES

Jorsenilvilehers Status Neport en sn UML LR UE Rens.

ER LE TR ne PRE RE SRE MP En ER RE ER ASE PE

269

270

271

272

273

274

275

276

PAU

NOvAPEx / Société 3(1), 10 mars 2002

En E _ a

Part 2

Jorsen Hyheberg Joyce Dengebun. In mena

Co-operating RESORT

Table of contents «insu. t-ne. deu iles CA MEN Re

BIODIVERSITY

Vibeke Simonsen & Kongkiat Kittiwattanawong: Marine molluscs: Application of

molecular markers for estimation of effective population size and gene flow...

David G. Reid: The use of the radula in the taxonomy and phylogeny of gastropods:

Cautionary cases of convergence, intraspecific variation and DIREHICIEYS CE RETR RES

Konstantin A. Lutaenko: Russian contribution to the studies of Vietnamese bivalve

molluscs. Part 1. Historical and bibliographical review with emphasis on faunal

SEUdeSE SA A OR NN ARENNE Pau ee Se

Konstantin A. Lutaenko: Russian Contribution to the studies of Vietnamese bivalve

molluses. Part 2. List of species recorded by Russian authors or stored in museums.

Bui Quang Nghi: Mollusca in sea grass of Van Phong Bay and Cam Ranh Bay, Khanh

H09 PrORGÉ D

Fred E. Wells & Clayton W. Bryce: Molluscan surveys of offshore coral reefs in

Northwestern Australia and adjacent biogeographical areas

Eddy Soekendarsi & Yulianus Pangaongan: The algal communities around Samalona

Island, Ujungpandane, South Sulawess "PP PNR

Sazlina Md. Salleh, Zulfigar Yasin & Aileen Tan Shau-Hwaï: The distribution and

diversity of chromodorid nudibranchs at Sipadan [Stand Bornéo) M UN ON

Bernard Tursch & Dietmar Greifeneder- Colour patterns in the genus Oliva: An

attempt to simple analysis and'interpretation.….. UNE

Jacques Vidal: Genus Vepricardium Iredale, 1929 (Bivalvia, Cardiidae), with description

of a new species from Thailand, Vepricardium albohamatum Hylleberg & Vidal...

Peter Middelfart: Taxonomic study of micro-molluscs: À case study using the Condylo-

carqidas. D

Wantana Yoosukh & Charuay Sukhsangchan: Electrophoretic studies on tuberculated

oysters.from-Ranong. Province Mhaïlandises. one CRU ITS

319

329

347

361

391

395

405

409

413

427

431

447

465

477

483

Novarex / Société 3(1), 10 mars 2002 37

—

Part 3

Jorgen Hylleberg: Joyce Dangebun. In memoriam........................................... res ii

Go Opera ENS AEUTIONSÉE ARR EE LR Tir Ter ee rennes \

RADIETOMEONTE DES EN RS PR Sn duree descendre core DS vil

LECTURE NOTES

Richard Kilburn: Basic rules of scientific nomenclature................................................s…… 539

Richard Kilburn: Principles of nomenclature: Worksheet (test your knowledge)... 547

Jorgen Hylleberg: Illustrated dictionary. Technical terms and common words used in

TÉÉRATIMOS DIVAIVE SR RE eee scies c oué 551

David G. Reid: Preservation and curation of marine molluscan specimens.….…........…........... 583

Alan J. Kohn: Molluscs: À microcosm of invertebrate zoology. An introduction to the great

payium/Molluseas se". 2... + hs As cet ne M Rene SAN 2 DIT 591

Richard Kilburn: Shallow-water "Archaeogastropods” of South-East Asia:

ATNTIGEO AUCH ONE MR RE NE ER ON PR RS RM rennes enseeccteseene cesse 595

David G. Reid: Introduction to Mesogastropods’…..........… RER Ps PAL: 2 PL 603

Alan J. Kohn: Introduction to Neogastropoda. RS 0 Ur SE EE 611

Kathe R. Jensen: Classification of the superorder Heterobranchia: Overview and

LACS Re ee RD Sn EDR r tent ett Dec node ec op Fe ir. 613

Richard Kilburn: Family Veneridae in South-East Asia. ....................... D CS AA ATRE 629

Jacques Val Class Meation ot Car ae re ener esp 639

www.sbm.be tt - Redirect by ulimit com - Netscape

SRE RS = œ

ESS SERRE Fe

Le site Internet de la SBM :

http://www.sbm.be.tf/

38 NoOVAPEX / Société 3(1), 10 mars 2002

Nous avons reçu

Claude VILVENS .

AMERICAN CONCHOLOGIST

(U.S.A. Sud-Est)

Vol. 29, N° 3, septembre 2001

M.W. JOHNSON : Hammock hopping and the reward of the day

J. COLTRO : New Caledonia : A French Paradise

E.K. SHEA : The Ommastrephid Proboscis

M. ERLENDUR & D. STRATMANN : Buccinum superangulare : freak or fact ?

J. MILLER : The search fo Melongena corona "sprucecreekensis"

L. BRUNNER & L. SCHROEDER: Coutdown at the Cape : Convention 2001

N.E. FABY : The ancient Maya and their modern land snails

K. & L. SUNDERLAND : Liguus fasciatus of South Florida — Part one

R.L. ALEXANDER : From the art of the scientific paper to paper arts by a scientist

P. MONFILS : The old shell game / Tusk shells : an unusual and unique class of mollusks

Diverses notes, revues et annonces …

++ + + + + + + + + +

KEPPEL BAY TIDINGS

(Australie — Queensland)

Vol. 40, N° 3, septembre-novembre 2001

J. OFFORD : Our trip to Clairview

JF. SINGLETON : Conus neptunus.

J. OFFORD : Oyster farming

P. BAIL & A. LIMPUS : A new species of Amoria

E. COUCOM : Surprise from the deep.

J. OFFORD : Cockling in England

Quelques notes, revues et annonces …

++ + + + + +

LES NATURALISTES BELGES

(Belgique)

Vol. 82, hors-série 2001

Comme cela arrive souvent : numéro exclusivement consacré aux Orchidées

GLORIA MARIS

(Belgique néerlandophone)

Vol. 40, N°2-3, novembre 2001

+ Ch.KRIJNEN, A.DELSAERDT, N. SEVERIINS, M. VERHAEGHE : Genus Werita (part 4) É

+ W. SEGERS & J.J. VAN AARTSEN : Aclis verduini Van Aartsen, Menkhorst & Gittenberger, 1984 : a

junior synonym of Aclis trilineata Watson, 1897.

ACTA CONCHYLIORUM

(Allemagne)

N°5

Beiträge zur Kenntnis der Ovulidae (Mollusca: Cypraeoidea)

VIII. Einleitung zur Familie sowie Katalog, Taxonomie und

Bibliographie und Bemerkungen zu verwandten Gruppen.

FEHSE, D.

NOvAPEXx / Société 3(1), 10 mars 2002

BASTERIA 7 }

(Pays Bas) ES =

Vol. 65, N° 4-6, décembre 2001

DAAN, R,, G.C.A. DUINEVELD, MSS. LAVALEYE & M. MULDER: Four marine

molluse species new 10 the Dutch recent fauna

GITTENBERGER, L.: Basteria accused, a reply

BRUGGEN, AC. VAN: Studies on the Streptaxidae (Mollusca, Gastropoda,

Pulmonata) of Mala@i 7. Gulella hildae Spec. nov., a rarity from the Mt. Mulanje

complex

PERRONE, AS: À new species of Nudibranchia of the genus Aldisa Bon

(Gastropoda, Opisthobranchia) from Thailand

AARTSEN, JJ. VAN, & J. GOUD: European marine Mollusca: notes on less well-

known species. XIL. Parvicardium carrozzni spec. nov., with notes on P minimum

(Philippi, 1836) and Cardium perrieri Dautzenberg & Fischer 1897 (Bivalvia,

Heterodonta, Cardiidae)

GITTENBERGER, E. & JJ. VERMEULEN: Oospira (O) pyhnosoma spec. nov.

(Gastropoda. Pulmonata. Clausiliidae), an impressive clausilüd species from

Vietnam

WESSELINGH. F: Book review

HORI, S., & R. KÜRODA: Spermatophores in {olava scitula Fe Ha 1860)

(Gastropoda, Heterobranchia, Pyramidellidae)

WINKLER PRINS, C.F: Book review

SMRIGLIO. C., & P MARIOTTINI: Pre Don SpEC. nov. esse

Prosobranchia, Fissurellidae}, a new bathyal species from the Mediterranean Sea 139

BRUGGEN, A.C. VAN: Boekbespreking NNITAA

AARTSEN, JJ.VAN: On the enigmatic Djeddilia Paie jénsserte, 1894

(Gastropoda, Caenogastropoda, ?Vermetidae) . 145

JANSSEN, A.W., & L ZORN: Notes on the systematics, or pre Descats

raphy of fossil holoplanktonic Mollusca. 11. Limacina Bosc, 1817: precedence over

Spiratella Blainville, 1817 (Mollusca, Gastropoda, Euthecosomata) 7

DIJRKSTRA, HH: Notes on taxonomy and nomenclature of Pectinoidea (Bivalvia,

Propeamussiidae & Pectinidae) 3 Nomina nova (rectifications) 100

MAASSEN, WLM: Nachträge zur Helicidenfauna der Insel Rhodos (Ctropode

Pulmünata, Hehcidae

NROMCEMOICONTRIBETORS. TL

Inhoud van Vol. 65

BULLETIN OF THE INSTITUTE OF MALACOLOGY TOKYO

Vol. 3, N° 8, septembre 2001

GOODWIN, Daniel R. A new species of Fusinus from the north-western Hawartan Islands

(Éasiopodasrastialanidhe) (Plate AIM RENCUEMENTMLT CRUE: * * à DMIUTrS

GOODWIN, Daniel R. The genus Ceflana H. Adams. 1869 from the Island of Kaho'olawe.

HasamMéastropadas Patelhdae)(Plare42,) 4,25 «50,52, + + + + pe MES

DA MOTTA. Mauricy Alves, Elaine Barros CARVALHO and

Ana Mara Mendonca DE ALBUQUERQUE MELO

DLcof *Co Gamma rays applied on Biomphalaria glabrata (Say,

Transfer of the holocvpes :

40 NOVAPEXx / Société 3(1), 10 mars 2002

LA CONCHIGLIA

(Italie)

Vol. XXXIII, N° 300, juillet-septembre 2001 The: Shell: |

À significant range extension for Epitonium auriculatum..to continental South America

BRUCE NEVILLE & SYBiL BURGER 11

The Genus Planktomya in the East Atlantic

JOHN JACOB VAN AARTSEN & WINFRIED ENGL 14

Errato: À New Species of Marginella from Northern Transkei.

MIKE HART

Further Notes on Conus vicweei Old, 1973 (Range extension and Color Variation)

MIKE HART

Shells from Bunaken (Celebes) d

FRANCESCO FONTEMAGGI 27

À New Species from Capraia Is. (Tuscan Archipelago): Alvania elisae sp. nov.

ALESSANDRO MARGELLI 43

4ïh International Workshop of Malacology: "Systematics, Phylogeny and Biology of Polyplacophora”

ANDREA BIDDITTU

Presence of Nodlitiorina punctata (Gmelin, 1791) Along the Coast of Lower Salerno Province

GiANNI D'ANNA ne Fr 53

New Data About Ischnochiton (Stenosemus) dolii Van Belle & Dell’ Angelo, 1998 n

BruNO DEu’ANGE10, G. Di PACO, R. ROCCHINI 55

Report of a Specimen of Buccinulum corneum Linné, 1758 With Three Eye Tentacles

PAOLO GiuLID ALBANO & FLAVIO FAVERO 57

SPIRULA

(Pays-Bas)

N° 322, 2001

Inhoud

Van de redactie

Agenda .

Rectificatie

J.G.I. Kuiper

À. Meckel

{Slurkie, cartoon]

F. Zandvoort Inventarisatie-excursie in het dal van de Ruiten-Aa (Groningen)

[ANM mededeling: 16]

H.K. Mienis Mollusken op waterlelies, 1. Slakken op Gele Plompen in

de Molslaan, Delft (met enkele notities betreffende Ferrissia)

[ANM mededeling: 17}

RH. de Bruyne &

S. van Leeuwen

C.M. Neckheirm, RH. de Landslakkeninventarisatie in de kuststreek van Zeeuws Vlaanderen

Bruvne. K. Oosterkerk & (Zeeland). [ANM mededeling: 19]

AN. van der Biil

R.H. de Bruyne, H. Wall- De Rijn-Glasslak Vitrinobrachium breve (Férrussac, 1821) bi]

brink & C.M. Neckheim Nieuwegein [ANM mededeling: 20]

H.J. Kwant. H. Wallbrink De Grootmondpluimdrager Valvara macrostoma (Férussac, 1821)

R.H. de Bruyne nog steeds in Nederland [ANM mededeling: 21]

H.K. Mienis Een tweede verkenning van de malacofauna van het natuurreser-

vaat ‘Het Heitje van Katham' [ANM mededeling: 22]............ 96

Oproepen/Mededelingen

R.A. Bank

R_A. Bank Artikelen in tiydschriften

R.A. Bank International Code of Zoological Nomenclature

R_A. Bank Nieuwe boeken

[Red.] Schelpenbeurzen, exposities en bijeenkomsten

{Larry Cook, column] Heavy taalstrijd

NovaPEXx / Société 3(1), 10 mars 2002 41

SPIRULA

(Pays-Bas)

N° 323, 2001

Voorplaat

Penmngmeester

Agenda

Excursiecommissie

Wolf, P. de

Mienis, HK.

Müenis, H.K.

Rijken, A.C.

Veen, B.v.d. & P.v.d.Kooi]

Cadée, G.C.

Wolf, P. de

Moolenbeek, R.G.

Meeuse, A.D.J.

Cadée, G-C.

Mienis, HK.

Hopman, HI.

Moolenbeek, RG.

Meeuse. A.D.J.

GCK

Meeuse, A.D.J.

TU Delft

Faber, W.

Dekker, H. & Faber, W.

Faber, W. & Walker, T.M.

Diverse bronnen

[Bestuursmededelingen]

Excursies 2002

Fossiele schelpen van het Texelse strand…….…........…....…........

De gezusters M.J. en A.J. de Graag:

een schelpenverzamelaarster en een kunstenares

Wratparels in Peronaea planata

met een notitie betreffende het gebruik van Peronaea

Galeodea bicatenata van de Kaloot

Verslag van een excursie naar Schotland

Scholekster eet Japanse oester (Crassostrea gigas)

Gruis van Brora (Sutherland), Schotland

Strombus gigas bewerkt tot steekwapen

Uit de literatuur

Juveniele Crassostrea gigas op adulte exemplaren, Waddenzee....116-117

Mariene mollusken uit het oostelijk deel van de Middellandse Zee

5. De eerste vondsten van Canarium mutabilis

6. De eerste vondsten van Cerithium egenum Gould, 1849

Zandsuppleties op Texel en de gevolgen voor de malacoloog

Alaba punctostriata Gould, 1861 nieuw voor de Turkse kust

Nieuws uit de diepzee

What is in the name”? (Column)

Vindplaatsen en verspreiding

Als uw been in schrven ligt, pakken we straks gewoon de lijmpot.

Nieuwe weekdiersoorten (schelpen)

Tijdschriftartikelen

Nieuwe boeken

Weekdieren op postzegels

Schelpenbeurzen en bijeenkomsten

Mutaties

.106-107

107-108

L'ESCARGOT OBSERVATEUR

(Belgique)

N°13

Le mot du Président — Vie associative

Séjour aux Guézoux

{ l’utile à l’agréable , diversification, exploitation, tarif des produits et

présentation de quelques produits )

Les conseils du chef - Timbale d'escargots aux fanes de radis

Chaussons aux petits escargots

Les propos de l’oncle Jules - Amendement des parcs

Les étiquettes de certaines conserveries française

TVA: achats à l'étranger

Cette année voici quelques belles réalisations de parcs chez nos membres (4)

Visité pour vous d'un parc d'engraissement au Pays des Collines (Belgique ;

Calendrier hélicicole et petite annonce (gratuite pour les membres)

SPIXIANA

(Allemagne)

Vol. 24, N°3, novembre 2001

Kreipl. K. & À. Alf:

Page 01

Pages 02 à 05

Page 06

Pages 07 et 08

Page 09

Page 10

Page 11

A new Oocorys from Western Australia (Mollusca. Gaästropoda.

Cassidae)

42 NOVAPEX / Société 3(1), 10 mars 2002

DOCUMENTS MALACOLOGIQUES

(France)

Vol. 1, juillet 2000

| H.GIRARDI et M. WIENIN : Les Hydrobiidae (Mollusca : Gastropoda : Prosobranchia) du massif

karstique de Saint-Julien-des -Rosiers (Gard)

S. CLANZIG et A. BERTRAND : 7rochoidea penchinati en France

A. BERTRAND : Atlas préliminaire des mollusques d’ Andorre

À. BERTRAND : Argna ferrari dans l’Hérault et |’ Aveyron

A. BERTRAND et C. MAUGE : Vertigo substriata en Ariège

H. GIRARDI : Inventaire des mollusques potamologiques, benthiques et hypogés de l’ensemble des

Gardons et de ses affluents

DOCUMENTS MALACOLOGIQUES

(France)

Hors série N°1, octobre 2000

Révision du Genre

Bythinella (MOQUIN-TANDON, 1855)

(Gastropoda Prosobranchia Hydrobiidae :

Amnicolinae Bythinellini)

de la France du Centre-Ouest,

du Midi et des Pyrénées

R. BERNASCONI

DOCUMENTS MALACOLOGIQUES

(France)

Vol. 2, novembre 2001

R. Bernasconi : Nouvelles données sur Valvata bourguignati Letourneux, 1869. (Prosobranchia :

Hydrobiidae : Horatiinae)

R. Bernasconi : Compléments à la connaissance du genre Bythinella (Prosobranchia : Hydrobiidae :

Amnicolinae) en France (Corrèze, Charente-Maritime)

H. Girardi et M. Rosello : Notes sur Bythiospeum kemmi (BOETERS 1969)

A. Bertrand : Quelques récoltes de mollusques continentaux de la faune de France

P. Geniez, À. Bertrand : Abida escudei sp. nov. du Sud de la France

H. Girardi : Moitessieria wienini sp. nov. Des eaux de l’aquifère de la montagne de la Selette (France,

Hérault) ( Mollusca : Gastropoda : Moitessieriidae)

A. Bertrand : Moitessieria fontsaintei sp. nov.

A. Bertrand : « Helicella » subcantabrica FAGOT 1882 en France

S. Clanzig et À. Bertrand : Mercuria emiliana (PALADILHE 1869) en France

S. Clanzig et À. Bertrand : Orala punctata (O.F. MULLER 1774) en France

MOLLUSCAN RESEARCH

(Australie)

Vol. 21, septembre 2001

B. Morton & E.M. Harper

Cementation in Cleidothaerus albidus (Lamarck, 1819)

(Bivalvia: Anomalodesmata: Pandoroidea)

D. S. Brown

Freshwater snails of the genus Gyraulus

(Planorbidae) in Australia: taxa of the mainland

NoOvaPEx / Société 3(1), 10 mars 2002 43

NOTICIARIO DE LA SOCIEDAD ESPANOLA DE MALACOLOGIA Te AS

(Espagne) DRE CC CPR GR 5: a s

N°36, décembre 2001 Nous 4

Editorial

Secretaria

Correspondencia de la SEM... PE DL En ER

Tesoreria

Noticias Malacolôgicas

Recensiones Malacolôgicas

Protecciôn de Moluscos y Espacios Naturales

Colaboraciones

Preguntas a... José Templado

Indices de revistas

Pasatiempos

Malacohumor

RESENAS MALACOLOGICAS

(Espagne)

Vol. XI

PROTECCIÔN DE MOLUSCOS EN EL

CATÂLOGO NACIONL 2° ESPECIES SMENSZRPAS

THE FESTIVUS

(U.S.A. — Californie)

Vol. XXXIII, N° 10, octobre 2001

Ciub news

Notes on the validity of Stramonita delessertiana (d'Orbigny. 1841) and Cancellaria (Massyla) Cumingiana

Peut de la Saussaye. 1844

Valentin Mogollén Avila

New Panamic Province distribution records for two species of Parviurbo Mollieca: Skeneidae)

Carol Skoglund

Photographer's note

David K. Mulliner

Book news: Monograph of the Living Zoila by F. Lorenz, reviewed

Terry S. Arnold & Hugh Bradner, reviewers ...................

44 NOVAPEX / Société 3(1), 10 mars 2002

THE FESTIVUS

(U .S.A. — Californie)

Vol. XXXIIL, N° 11, novembre 2001

Club news

É d8e (Nicema) subrostrrata found Withe egg cases

Jules Hertz .

New Panamic Province code for de deep-water buccinid Neptunea amianta (Dall. 1890)

(Gastropoda: Buccinidae)

Kent D. Trego

2002 winter and spring shell shows and other events (compiled by Donald Dan) .

À selected index to Volume XXXIII (2001)

THE CONCHOLOGISTS' NEWSLETTER

(Grande-Bretagne)

Vol. 10, N°159, décembre2001

Du Feu, Chris How to record slug abundance?

Lindlev, David Field meeting to the Yorkshire Dales

Alexander, Keith Wanted - images of British land and

freshwater mollusces

Verdcourt, Collectors in East Africa No. Hi: Robert

{Robin] Kemp (1871-72) (Part 1}

Gillard, Colin Marine field trip to Kilve, Somerset and

September

some North Devon sites, 27

1 October 200

Carr, Ron Fhe e-mail file

Sumner, Adrian l.. Responses to: ‘Future developments for

Hartield, David, the Conchological Society"

Longrigg, Sarah

VENUS

(Japon)

Vol. 60, N° 3, septemebre 2001

Takashi Okutani: Six new bathval and Shelf trochoïdean species in Japan

Yasunori Kano. Takenori Sasaki and Hiroshi Ishikawa: Neriütilia ntimotoi, a new nenulid

species trom an anchialine lake and estuaries in southwestern lADAN RE ES STE

els rs enr pe eriesie na saisie « FRET

PP DEC OC IOOICICIOICICI ICI CEE

gew venus from Kuroshima Island. Okinawa. Japan ent

Den uessrsmnmsaensesnslas ess rinnisie einiviele se 2ieeiv pie)» the ete er

OREILLES

Hiroyuki Ozawa: Reproductive cycle and spawmne of Modiolus philippinarim (Bivalvia:

Mytilidac) in à seagrass bed in Kin Bay, Okinawa Island. southern Japan "7"

Rvuji Onoyama, Yoshiro Noda. Hitomi Takada and Hiroshi leyama: Gonad structures in two

à D Hre Te £ « RO Re a DU On LU A AE COCO tac on g3

species of Pisidium (Bivalvia! Sphacridue) 18:

Kazutaka Amano. Toshikazu Hamuro. Masui Hamuro and Shoji Fujit The oldest vesicomyid

bivalves from the Japan Sea borderland

Harumi Ohtaki. Eiko Maki and Kivonori Tomivarna: Seasonal changes in the distribution and

: SN AIT . < Son fe aAtamididaer scosscsceumss OC

matine behavior of Cerithideu rhizophorarum {Gastropoda: Potamididae) 199

oo im a

NovaPEx / Société 3(1), 10 mars 2002

TRITON

(Israël)

N°4, septembre 2001

MARINE MOLLUSCS

Buzzurro G.

&Russo P. FUSINUS DALPIAZI (COEN, 1918), A CONTROVERSIAL SPECIES......... 1

Hein ET ABNORMALITIES IN COWRIES 2. AN UNIQUE ROSTRATED SHELL OF

: CEPRAP ADI RIS ES ere rananin na an ons ee se ours en an le Le dede en du Te 4

Mienis, HK. ON THE IDENTITY

L OF GALEOMMA RUEPPELLI F.HAAS, 1935. esse SEA 6

Ra CYPRAEA PANTHERINA RASNASRANIE NSIS NEW SSP.

& Mienis H K FROM THE EAST COAST OF SINAI (GASTROPODA:

PROSOBRANCHIA:CYPRAEIDAE:CYPRAEAINAE)............... {)

5 FRS VARIABILITY OF COWRY POPULATIONS

IG ABOL'RNHE TERRES COMPLEXE SR Resa one eva nessunscnrasc esse 12

Heiman E. L. VARIABILITY OF COWRY POPULATIONS 17. LEPORICYPRAEA

MAPPA: TEETH COLOR AS A SHELL CHARACTERISTIC................... 17

Dekker Henk ADDITIONS TO THE KNOWLEDGE OF LACRA (BIVALVIA; EMELIDAE)

WITH COMMENTS ON THE TAXONOMYOF THE SPECIES................ 19

Singer B.S. SCAPHOPODA OF EAST SINAÏ 1. AN INTRODUCTION ss. 24

LAND-SNAILS AND FRESH-WATER MOLLUSCS

THE MOLLUSC FAUNA

QuiR © OF THE NA'AMAN CATCHMENT AREA, ISRAEL

| 1. A REVIEW OF THE RECORDS OF THE INLAND MOLLUSCS......... 27

LISE A PRELIMINARY SURVEY OF ALBINARIA POPULATIONS

AROUND KUSADASI BAY, TURKEY

Dern nsnnmn en nnannsn amas ssnense

XENOPHORA

(France)

N°96, octobre-décembre 2001

3- Editorial par P.Bail

4- Droit de réponse de E. Guillot de Suduiraut

5- Le coin du Débutant par G. Jaux

8- Sur les traces de E. & H. Vokes par J. Morin

12- Crépidules en sursis, article lu par M. Gueguen

13- Quelques fleurs de la Caraïbe de N. Lauranceau

14- Remarks on Glycymerididae from French Guyana

and Antilles par G. Paulmier

27- Retour sur un « Identifiez- moi » de J. Marronnier

28 & 2- Journée de grande marée à Port-Gentil

par J. Gourayeb

28- Petites annonces

29- La plongée en apnée par JL. Delemarre

33 & 43- Tombola de l'AFC 2001/2002 par T. Dhainault

34- À la recherche des porcelaines seychelloises

par D. Touitou

37- Couleurs des Antilles (séries 2 & 3),

photos de G. Boltz

39- Vie des sections

40- Echo. quillages

42- Ile Maurice-Pollution {suite}, lu par T. Dandrimont

44- Ramassage enfantin (série 2), photo de M. Hary

46 NOVAPEXx / Société 3(1), 10 mars 2002

Prochaines activités de la SBM

Claude VILVENS

Lieu de réunion : Médiathèque de l'Institut St Joseph - Rue Félix Hap 14 - 1040 Bruxelles

à partir de 14h. Sonnez et l'on vous ouvrira !

SAMEDI 23 MARS 2002

Georges VAUQUELIN et Isabelle VAN LIEFDE : L'Australie Occidentale et le groupe Zoiïla

(Cypraeidae)

Notre cycle de conférence nous emmène décidément un peu partout de par le monde (par la pensée seulement,

d'accord, mais tout de même © !). Cette fois, ce presque-continent qu'est l'Australie nous est évoquée par notre

“docteur es venin de Cônes" — si ce n'est qu'il s'agira de Porcelaines …

XX X

SAMEDI 20 AVRIL 2002

Jacques et Rita SENDERS : Madagascar

Existe-t-il un coin du monde où nos infatigables globe-trotters n'ont pas posé leurs pieds ? De toute manière, les

voici dans cette grande île de l'Océan Indien où tant de richesses malacologiques attendent d'être découvertes.

* XX

SAMEDI 12 MAI 2002

Claude VILVENS : Les Cantharidini : des Troques mal connus.

"L'homme qui aimait les Trochidae" (je vous laisse deviner ce qu'aime le Président ;-) ...) envisage donc de nous

parler de Trochidae que les collectionneurs n'affectionnent que modérément à cause de leur (relative) petite

taille : les Cantharidus, Jujubinus, Phasianotrochus et autres Thalotia.

XX *

SAMEDI 8 JUIN 2002

L'excursion de la SBM.

Où allons-nous aller cette fois ? Aucune idée pour l'instant ! Nos équipes de reconnaissance (—Claude et Etienne)

vont le déterminer au retour du printemps. Comme d'habitude, le plus simple est de contacter quelques jours

auparavant soit Claude (cvilvens@prov-liege.be ou 04/248.32.25), soit Roland (Roland.Houart@skynet.be ou

016/78.86.16). Quoi qu'il en soit, emportez votre bonne humeur … et prévoyez aussi bottes et vêtements de pluie

(au cas, totalement improbable, où il pleuvrait légèrement ;-) ...).

kX%X

SAMEDI 22 JUIN 2002

Etienne MEULEMAN : Hommes et coquillages

Nous savons que ce thème passionne Etienne (et il n'y a pas que lui !). Il nous propose ici une fabuleuse histoire

à travers l'espace et le temps. Voici la 1ère partie : coquillages et monnaies.

NoOVAPEX / Société 3(1), 10 mars 2002 47

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NOVAPEX

Trimestriel de la société Belge de Malacologie

association sans but lucratif

Quarterly of the Belgian Malacological Society

DOS 12) 10 JUILLET

SOMMAIRE CAEN

JUL 2 9 2002

—— © — HARVARD

Articles originaux — Original articles UNIVERSITY

L. R. L. Simone Three new deepwater species of Eulimidae (Caenogastropoda) 55

from Brazil

C. Vilvens Description of Zetela alphonsi n.sp. (Gastropoda: Trochidae:

Solariellinae) from Chile

A. Wakefield Review of the genus Pachybathron Gaskoin, 1853

F. Boyer (Gastropoda : Cystiscidae)

T. McCleery

K. Kreipl A new species of Galeodea Link, 1807 (Mollusca:

A. AIf Gastropoda: Cassidae) from the Philippine Islands

F. Boyer Description of five new marginellids from bathyal levels

of southern New Caledonia

Vie de la Société — Life of the Society

C. Delongueville et Bivalves et Crustacés - illustrations d’une association

R. Scaillet

R. et J. Senders Madagascar II

(suite du sommaire en dernière page de couverture)

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SOCIETE BELGE DE MALACOLOGIE

L.R.L. SIMONE

Three new eulimids

NOVAPEX 3 (2-3): 55-60, 10 juillet 2002

Three new deepwater species of Eulimidae

(Caenogastropoda) from Brazil

Luiz Ricardo L. Simone

Museu de Zoologia da Universidade de Säo Paulo

Cx. Postal 42594

04299-970 Säo Paulo, SP, Brazil

Irsimone@usp.br

KEY WORDS. Eulimidae, Annulobalcis procera n.sp., Batheulima lutescens n. sp., Batheulima

epixantha n.sp., Brazil, deepwater.

ABSTRACT. Three new deepwater species are described, belonging to the family Eulimidae,

based on shells collected in the Marion-Dufresne expedition and Revizee in southeast coast of

Brazil. Annulobalcis procera sp. n. has the shell longer and slenderer than the remainder congeners

and 1s the second described species of the genus for the South Atlantic. The other 2 species belong

to genus Batheulima, both have characteristic yellow band in each whorl and are the first record of

the genus to the south Atlantic. B. lutescens sp. n. is larger, and has a longer aperture and thinner

inner lip than B. epixantha sp. n.

INTRODUCTION

The MD-55 expedition performed by the R. V.

Marion-Dufresne in 1987, dredged in the southeast

coast of Brazil, and collected several deep-water

molluses, most of them still undescribed. In the

present paper, 2 new species of Eulimidae

(Caenogastropoda) are described based on shells

collected in depths of more than 600 m. The third

species was collected by two Brazilian projects,

which also have collected several undescribed

species in the southeastern coast, as following. The

Revizee project, ‘“Programa de Avaliaçäo do

Potencial Sustentavel de Recursos Vivos na Zona

Econômica Exclusiva”; and the PADCT project,

"Importância e Caracterizaçäo da Quebra de

Plataforma Continental para Recursos Vivos e Näo

Vivos".

The Eulimidae are parasitic on echinoderms. Warén

(1983) reviewed its genera, and this systematic

arrangement is adopted herein.

The genus Annulobalcis Habe, 1965, encompasses 5

species: À. shimazui Habe, 1965 (type species from

Japan); À. yamomotoi Habe, 1974 (also from Japan):

A. marshalli Warén, 1981 (from New Zealand); À.

ptilocrinicola (Bartsch, 1907) (from British

Columbia): and À. aurisflamma Simone & Martins,

1995 (from Brazil). The latter is the only species

described for the Atlantic Ocean, and of which the

anatomy is known. The members of the genus

Annulobalcis have been found to be ectoparasites of

crinoids.

The genus Batheulima Nordsieck, 1968, has been

described for B. fuscoapicata (Jeffreys, 1884)

(Bouchet & Warén, 1986), from Portugal, and was

reported for deep waters. The host of Batheulima

species 1s still unknown.

The Marion-Dufresne material is deposited at the

mollusk collection of the Musée National d'Histoire

Naturelle of Paris, France (MNHN), while Revizee

and PADCT material is deposited the Museu de

Zoologia da Universidade de Säo Paulo, Säo Paulo,

Brazil (MZSP). Except for some paratypes, that were

distributed among these. Comparison with shell

photo of Dall’s type specimen was possible under

courtesy of the National Museu of Natural History,

Smithsonian Institution (USNM).

SYSTEMATICS

Annulobalcis procera, Sp. nov.

Figs. 1-4

Type material. Holotype. MNHN, shell.

Paratypes. MZSP 34512 + 34513, 2 shells from type

locality. BRAZIL; Säo Paulo: off Säo Sebastiäo

Island, 23°47°S 42°10°W, 610 m depth, 3 shells

(MDS55, sta. CB105, Bouchet, Leal & Métivier col.

02/vi/1987).

Type locality. BRAZIL: Espirito Santo; off

Conceiçäo da Barra, 19°36°S 38°53°W, 640 m depth

(MDS55, sta. CB93, Bouchet, Leal & Métivier col.

v/1987).

Diagnosis. SE Brazilian deepwater species. Shell

long, slender, translucent, white. Aperture long,

almost half of spire length.

Description. Shell up to 15 mm, turriform, slender,

narroW, pointed (apical angle about 27°), up to 12

weakly convex whorls. Color translucent white,

semi-transparent. Protoconch with 2.5 whorls; first

whorl flat, remainder whorls narrow, almost of same

size. Transition between protoconch and teleoconch

L.R.L. SIMONE

Three new eulimids

NOVAPEX 3 (2-3): 55-60, 10 juillet 2002

marked by distinct sigmoid, shallow, axial furrow.

Spire very long, slender. Suture shallow, but distinct.

Shell surface almost smooth, glossy; marked by

several extremely thin, somewhat uniform spiral

striae; subsutural striae slightly deeper. Axial,

shallow furrows sparsely present, apparently

randomly distributed. Aperture long, narrow; length

almost half of spire length. Inner lip weakly concave:

callus somewhat broad, more conspicuous anteriorly,

preceding inferior area. Outer lip with sharp edge,

middle region slightly projected, and convex. Inferior

region somewhat wide, with lip deflected, projected

forwards. Columella simple. Umbilicus absent.

Measurements (in mm). Holotype: 14.6 by 4.6;

Paratypes (from type locality): MZSP 34512) 13.4 by

4.4; MZSP 34513) 5.8 by 2.4; (from sta. CB105): 7.7

by 2.7.

Distribution. S. E. Brazilian coast, from Espirito

Santo to Säo Paulo.

Habitat. The depth ranges from 610 to 640 m. No

information about the host is available.

Etymology. The specific name refers to the

elongated aspect of the shell, from Latin procera,

meaning slender, tall.

Batheulima lutescens, sp. nov.

Figs. 5-8

Type material. Holotype. MNHN, shell.

Paratypes. BRAZIL, (MDSS, Bouchet, Leal &

Métivier col.). Espirito Santo; off Säo Mateus,

18°59°S 37°50°W, 637 m depth, 1 shell (sta. CB76).

Säo Paulo: off Säo Sebastiäo Island, 23°47°S

42°10°W, 610 m depth, MZSP 34514 + 34515, 2

shells, MNHN, 2 shells (sta. CB105, 02/vi/1987).

Type Jlocality. BRAZIL; Espirito Santo; off

Conceiçäo da Barra, 19°36°S 38°53°W, 640 m depth,

(MDS55 sta. CB93, Bouchet, Leal & Métivier col., v/

1987).

Diagnosis. SE Brazilian deepwater species. Shell

suture shallow. Color translucent white, with yellow

spiral band in middle region of each whorl. Aperture

almost half of spire length.

Description. Shell up to 10 mm, turriform, slender,

narrow, pointed (apical angle about 22°), up to 14

Figures 1-8

weakly convex whorls. Color translucent white, with

a broad spiral band, yellow, running along middle

region of each whorl. Protoconch with 2 whorls, first

whorl mamillated, pale brown, remainder whorls

narrowW, increasing weakly. Transition between

protoconch and teleoconch marked by distinct

sigmoid, shallow, axial furrow. Spire very long,

slender. Suture very shallow, distinctively to weakly

marked. Surface almost smooth, glossy; delicately

marked by growth lines. Last whorl weakly to

distinctively angulated. Aperture long, narrow; length

almost half of spire length. Inner lip slightly concave:;

callus very narrow. Outer lip with sharp edge, middle

region slightly projected and convex. Inferior region

somewhat wide, with lip deflected, projected

anteriorly. Columella simple. Umbilicus absent.

Measurements (in mm). Holotype: 10.0 by 3.2;

MNEN (#1) (from type locality): 8.6 by 2.7; MZSP

34514: 10.7 by 3.2; MZSP 34515: 6.8 by 1.9; MNHN

(from sta. CB5S): 7.8 by 2.5.

Distribution. S. E. Brazilian coast, from Espirito

Santo to Säo Paulo.

Habitat. The depth ranges from 610 to 640 m. No

information about the host is available.

Etymology. The specific name refers to the

yellowish color of the shell band, from the Latin

lutescens, meaning yellowish.

Batheulima epixantha, sp. nov.

Figs. 9-12

Type material. Holotype. MZSP 34518, shell.

Paratypes. MZSP 34519 + 34520, 2 shells, MNHN, 1

shell (#3), from type locality. BRAZIL: Säo Paulo,

off Ilha de Santo Amaro, 24°17.939°S 44°35.983°W,

133 m depth, MZSP 34516, 1 shell (Sta. 6673,

11/1/1998); off Santos, 24°40.747°S 44°50.822°W,

137 m depth, MZSP 34517, 1 shell (sta. 6677,

12/1/1998).

Type locality. BRAZIL; Santa Catarina; off Itajai,

27°10.380°S 47°27.540°W, 129 m depth (sta. 6635,

09/x11/1997).

Diagnosis. S. E. Brazilian deepwater species with

shell suture shallow. Color semi-transparent, whitish,

with a narrow yellow spiral band in supra-sutural

region of each whorl. Aperture almost 1/3 of spire

length.

1-4. Annulobalcis procera. 1. Holotype, SEM, scale = 0.5 mm 2. Same, optical photo, scale = 1 mm 3. Paratype

1 (MZSP 34512), scale= 1 mm 4. Detail of holotype protoconch, SEM, scale= 50 um. Figs. 5-8. Batheulima

lutescens 5. Paratype 1, SEM (MNHN, Sta. 105), scale= 0.5 mm 6. Paratype 2, SEM (same data), scale—0.5 mm

7. Holotype (MNHN, Sta. 93), scale= 1 mm 8. Protoconch of paratype 1, SEM, scale = 0.1 mm.

L.R.L. SIMONE Three new eulimids NOVAPEX 3 (2-3): 55-60, 10 juillet 2002

L.R.L. SIMONI

lhree new eulimids

NOVAPEX 3 (2-3): 55-60, 10 juillet 2002

mm, turriform, slender,

narrow, pointed (apical angle about 26°), up to I1

Description. Shell up to 7

weakly convex whorls. Color translucent white, with

narrow yellow spiral band, running along supra-

sutural region of each whorl. Protoconch with 2

whorls, first whorl mamillated, pale brown,

remainder whorls narrow, increasing weakly.

lransition between protoconch and teleoconch

marked by distinct sigmoid, shallow, axial furrow.

Spire very long, slender. Suture very shallow,

distinctively to weakly marked. Surface almost

smooth, glossy; marked only by delicate growth

lines. Last whorl weakly to distinctively angulated.

Aperture moderately long, length almost 1/3 of spire

length. Inner lip slightly concave: callus somewhat

broad. Outer lip with sharp edge, middle region

slightly projected and convex. Inferior region

Figures 9-13

rounded, with lip not deflected. Columella simple.

Umbilicus absent.

Measurements (in mm). Holotype: 5.7 by 1.7;

Paratypes (from type locality): MZSP 34519: 6.6 by

1.8; MZSP 34520: 3.5 by 1.2; MNHN (from type

locality): 3.6 by 1.2.

Distribution. S. E. Brazilian coast, from Säo Paulo

to Santa Catarina.

Habitat. The depth ranges from 129 to 137 m. No

information about the host is available.

Etymology. The specific name refers to the

yellowish color of the shell band, from the Greek

epixantha, meaning part yellow.

9-12. Batheulima epixantha. 9-11. Holotype 9. Frontal view 10. Detail of apex 11. Detail of the aperture

12. Paratype (MZSP 34519). Scales = 1 mm 13. Srrombiformis elata Dall, 1927, type USNM 108381, courtesy

USNM, length 8 mm.

L.R.L. SIMONE

Three new eulimids

NOVAPEX 3 (2-3): 55-60, 10 juillet 2002

DISCUSSION

The three species described here differ from any

Brazilian species of eulimid (Rios, 1994), except

Annulobalcis aurisflamma, in having well marked

suture. Most eulimids from this region have the

suture in a straight profile.

Annulobalcis procera shell differs from remainder

congeners in being longer and narrower.

Additionally, it differs from À. aurisflamma in being

less transparent (maybe an artifact in being dead

collected), in having a shorter aperture (almost half

of spire length, while À. aurisflamma has the aperture

length only slightly shorter than the spire), and more

spiral striae. The depth where they occur are also

very different, as À. procera was collected at a depth

of more than 600 m, while À. aurisflamma occurs

from the intertidal zone to 8 m depth.

Batheulima lutescens and B. epixantha Were assigned

to this genus due to their similarity with the type

species, the Northern Atlantic B. fuscoapicata, in

characters such as the shell and aperture shape,

impressed suture, and the darker protoconch. They

differ from the type species in having a longer

aperture (from almost half to 1/3 of spire length,

while B. fuscoapicata has aperture length with about

1/4 of the spire), and by the color. B. epixantha

differs from B. lutescens by being smaller (around 6

mm, while B. lutescens reaches more than 10 mm),

the color pattern (the yellow spiral band is narrower

and situated in the supra-sutural region of each

whorl, while B. lutescens has the band broader,

situated in the middle region of each whorl). Also,

the aperture of B. epixantha is shorter (about 1/3 of

spire length, while B. lutescens the aperture 1s about

half of spire length), has thicker lip, the inner lip has

a broader callus, and the anterior (siphonal) region 1s

simpler and blunter (while this region of B. lutescens

is clearly projected forwards). The outline looks

different, B. epixantha is broader, with spire angle of

about 26°, while B. lutescens is sharper pointed, with

spiral angle of about 22°. The depth of occurrence

appears to be also different, as B. epixantha occurs at

about 130 m and B. lutescens about 600 m deep.

B. lutescens resembles Strombiformis patula (Dall &

Simpson, 1901), differ in having less projected outer

lip, shorter aperture and pointed apex<. B. lutescens

also resembles Srrombiformis fusus (Dall, 1889),

differs in having broader aperture, wider spire and

broader protoconch. The three species described here

differ from those eulimids described by Dall (1927)

and Watson (1886), mostly by well marked suture.

The single which can be confused is Srombiformis

elata Dall (Fig. 13), from which the species here

described differ in having the aperture proportionally

longer, broader outline, apex more pointed, and in

being larger.

ACKNOWLEDGMENTS

The material collected by the Marion-Dufresne

expedition was sent for this study as courtesy of Dr.

Philippe Bouchet and Philippe Maestrati, at the

MNEHN. The Revizee material was sent for study by

Dr. Cintia Miyaji, IOUSP (Instituto Oceanogräfico

da Universidade de Säo Paulo) and Dr. Antonia

Cecilia Amaral, UNICAMP (Universidade de

Campinas), who send the material of the south

regional sub-committee. The photo of the

Strombiformis elata type was courtesy of Tyjuana

Nickens and Dr. M.G. Harasewych, from USNM.

The Revizee project is supported by “Ministério do

Meio Ambiente, dos Recursos Hidricos e da

Amazônia Legal” (MMA), “Instituto Brasileiro do

Meio Ambiente e dos Recursos Naturais

Renoväveis” (IBAMA), “Comissäo Interministerial

para os Recursos do Mar” (CIRM) and “Programa de

Apoio ao Desenvolvimento Cientifico e

Tecnolégico” (CNPq). The PADCT material was

responsibility of Dr. Cintia Miyaji and Dr. Airton

Tararam, IOUSP. For Paulino José S. Souza Jr. for

reviewing the manuscript. For Lara M. Guimaräes for

helping the SEM studies. This project is carried on

by the CNPg, and is performed by the IDUSP. This

study 1s part of a project developed under a support

of Fapesp, Fundaçäo de Amparo a Pesquisa do

Estado de Säo Paulo (procs. # 00/11074-5 and

#00/11357-7).

REFERENCES

Bartsch, P. 1907. A new parasitic mollusk of the

genus Eulima. Proceedings of the United States

National Museum 33(1548): 555-556.

Bouchet, P. & Warén, A. 1986. Revision o the

northeast Atlantic bathyal and abyssal Aclididae,

Eulimidae and Epitonnidae. Bollettino

Malacologico suppl.2: 299-576.

Dall, W.H. 1889. A preliminary catalogue of the

shell-bearing marine mollusks and brachiopods of

the SE coast of the U.S. with illustration of many

of the species. Bulletin of the United States

National Museum 37: 1-232 + 95 pls.

Dall, W.H. 1927. Small shell from dredging off the

SE coast of U.S. by the “Albatross” in 1885/6.

Proceedings of the United States National

Museum 70(18): 1-134.

Dall, W.H. & Simpson, C.T. 1901. The Mollusca of

Puerto Rico. United States Fish Commission

Bulletin 20: 351-524 + pls. 53-58.

Jeffreys, J.G. 1884. On the Mollusca procured during

the Lightning and Porcupine expeditions, 1868-

1870. 8. Proceedings of the Zoological Society of

London 24: 341-372.

L.R.L. SIMONE Three new eulimids NOVAPEX 3 (2-3): 55-60, 10 juillet 2002

Habe, T. 1965. Description of Annulobalcis

schimazui n. gen. Et sp. (Eulimidae). Venus 24:

106-107.

Habe, T. 1974. Five new gastropodous species

parasitic in the Japanese Echinoderms. Venus 32:

117-123.

Nordsieck, F. 1968. Die Europäischen

Meeresgehäuseschnecken. G. Fischer. Stuttgart,

273 pp.

Rios, E.C. 1994. Seashells of Brazil, second edition.

Fundaçäo Universidade do Rio Grande. Rio

Grande, 368 pp. + 113 pls.

Simone, L.R.L. & Martins, C.M. 1995. Annulobalcis

aurisflamma, a new species of Eulimidae

(Gastropoda, Prosobranchia) parasitic on a

crinoid from Brazil. Journal of Conchology 35:

223-235.

Warén, A. 1981. Eulimid gastropods parasitic on

echinoderms in the New Zealand region. New

Zealand Journal of Zoology 8(3): 313-324.

Warén, A. 1983. A generic revision of the family

Eulimidae (Gastropoda, Prosobranchia). Journal

of Molluscan Studies suppl. 13: 1-96.

Watson, R.B. 1886. Scaphopoda and Gastropoda.

Report on the Scientific Results of the Voyage of

the “Challenger” During the Years 1873-1876

15(42): 1-756 + 53 pls.

VILVENS

Zetela alphonsi n.sp.

NOVAPEX 3 (2-3): 61-64, 10 juillet 2002

Description of Zerela alphonsi n.sp.

(Gastropoda: Trochidae: Solariellinae)

from Chile

Claude VILVENS

Rue de Hermalle, 113 - B-4680 Oupeye, Belgium

cvilvens(@prov-liege.be

KEY WORDS. Gastropoda, Trochidae, Chile, Zetela alphonsi n. sp.

ABSTRACT. Zetela alphonsi n. sp. is described and compared with similar Solariella-like species

from deep waters around the world.

RESUME. Zerela alphonsi n. sp. est décrite et comparée avec des espèces analogues à l'aspect de

type Solariella et provenant d'eaux profondes du monde entier.

INTRODUCTION

A few month ago, once again, Guido T. Poppe, well

known shell collector from Belgium, entrusted me

with shells from deep water off Chile. This reminded

me that Trochidae from this area are relatively poorly

known and seldom well illustrated. Especially, only a

few authors seem to have studied the trochids from

off Chile and, in this group, it seems a bit easier to

find numerous data and illustrations about shells

from the magellanic province (Dell, 1971, 1990;

Forcelli, 2000) as to get some information about

species from central (Rehder, 1971; Mc Lean &

Andrade, 1982) and north Chile (Santa Maria, 1982).

In this later case, it is often necessary to look for

papers talking about neighbouring areas, like Peru or

Galapagos Islands (Dall, 1919; Keen, 1971; Finet,

1995)

Nevertheless, the peculiar shape of the shells that

initiated this paper, obviously Solariella-like, 1s very

striking and reminds only weakly any known species

from this area and belonging to this group of

trochids. After further studies, it appears these shells

belong to a species different from all described

species.

Abbreviations

Repository

IRSNB : Institut royal des Sciences naturelles de

Belgique, Bruxelles.

MNEN : Muséum national d'Histoire naturelle, Paris.

Other abbreviations (Text Fig.)

D : diameter

H : height

HA : height of aperture

BIMP2A12% : primary cords (PI is the most

adapical)

SI S2,159

adapical)

lv : live-taken specimens present in sample

: secondary cords (SI is the most

NORMES - |

T :

Tec. je

CLR CR e

ss sr LEP P4 FR

SOS Î REC à

h | CEST ET US —

ARTS d

RaReT

SYSTEMATICS

Family: TROCHIDAE Rafinesque, 1815

Subfamily : SOLARIELLINAE Powell, 1951

Genus: Zetela Finlay, 1927

Type species: Minolia textilis Murdoch & Suter,

1906 (by original designation) - Recent, New

Zealand.

Zetela alphonsi n.sp.

Figs 2-6

Type material. Chile, off Chiloé, trawled in

800 m on muddy bottoms, holotype MNEN,

15.5 x 11.9 mm, (Iv);. paratype, .IRSNB,

11.5x9.8 mm (lv); paratype, 10.0 x 8.8 mm

(1v), author's collection; paratype, 8.3 x 7.7 mm

(1v), collection G.T. Poppe .

” Stanislas Leclefstraat, 8, 2600, Berchem, Belgium.

VILVENS

Zetela alphonsi n.sp.

NOVAPEX 3 (2-3): 61-64, 10 juillet 2002

Other material. Chile, off Taltal, Antofagasta,

trawled in 900-1000 m on muddy bottoms, 2 1v, coll.

G.T. Poppe.

Diagnosis. A Solariella-like species, green

irridescent, with a wide umbilicus, conspicuous

tubular and shouldered whorls bearing three

spiral cords and strong axial ribs.

Description. She/! rather large for the genus (height

up to 15.5 mm, width up to 11.9 mm), higher than

wide, thin, scalariform; spire high, 2x to 3.8x higher

than aperture, widely umbilicate.

Protoconch of about 1.5 whorl (first whorl damaged

in all specimens examined), sculptured by granules

and a thick abapical spiral cord; apical fold rounded.

Teleoconch up to 4.5 convex tubular whorls, bearing

spiral cords and prosocline threads, with one

subsutural shoulder. Suture visible, slightly

canaliculated.

First teleoconch whorl convex, sculptured by 2

granular primary cords and strong prosocline ribs,

producing axially elongated nodules at intersections:

PI and P2 similar in size and shape, evenly

distributed on whorl; number of axial ribs of about

25. Subsequent whorls with sutural ramp with

rounded rim, gently sloping on second whorl,

becoming almost horizontal on last whorl and

producing obvious shoulder at first quarter. On

second whorl, P3 appearing near suture, weaker than

P1 and P2. On third whorl, P3 staying weaker than

others; axial ribs becoming slightly finer and more

numerous by intercalation, producing therefore

smaller granules. On last whorl, axial ribs

evanescent; very weak secondary spiral cords S1 and

S2 appearing between P2 and P3.

Aperture subcircular, lip thin at rim; peristome

complete in fully adults specimens: outer lip with

weak angulations corresponding to external spiral

cords; inner lip meeting outer lip at a strong basal

angulation.

Base convex, sculptured as last whorl, with 3 or 4

spiral cords, cord around umbilical area strongest:

crowded axial riblets, weakly lamellose, producing

weak nodules at intersections with spiral cords.

Umbilicus very wide, funnel-shaped, with angulate

rim bordered with most internal spiral cord of base:

wall convex; sculptured within by fine axial threads,

occasionally (paratype 2) by 1 granular spiral cord

near rim.

Colour of protoconch and first whorl of teleoconch

dark green, subsequent whorls light green

irridescentaperture nacreous within.

Operculum horny, multispiral, with short growing

edge and about 12 volutions.

Discussion. The new species belongs undoubtely to

the subfamily Solariellinae. The genus

Lamellitrochus Quinn, 1991 could be a right choice

for it, but I follow Marshall (1999) who state, based

on conchological and anatomical arguments, that this

genus is a junior synonym of Zetela Finlay, 1927.

The peculiar shape of Zetela alphonsi n.sp. implies

that it can hardly be confused with another known

species from Chile.

The description of the new species remember

nevertheless Calliotropis illota (Watson, 1886) (Fig

1) from South Chile and Patagonia, but this species

has a spire less elevated without distinct shoulder,

bears 4 spiral cords instead of 3 with nodules

horizontally, not axially, elongated (ide original

description).

Zetela alphonsi n.sp. may be compared to

Calliotropis nyssona (Dall, 1919), but this species

from Japan and China is smaller and bears about 8

spiral cords; moreover, the granules of the subsutural

cord are pointed.

The new species is also superficially similar to Zetela

annectens Marshall, 1999 from New Zealand and

Lamellitrochus pourtalesi (Clench & Agayo, 1939)

from West Atlantic, but these species are smaller and

show a very different shape with 3 keels.

Etymology. The new species is named after our

friend Alphonse Thielemans, Belgium, who was a

faithful member of the Belgian Malacological

Society and who unfortunately passed away in 2000.

ACKNOWLEDGEMENTS

I would like to thank G. T. Poppe (Berchem,

Belgium) who draw my attention to the specimens

upon which the present work is built. I am also very

grateful to P. Bouchet (Muséum national d'Histoire

naturelle, Paris) for access to the malacological

ressources of the MNHN, to V. Heros (MNHN) for

the kind attention she gave to all my enquiries for

searching various scientific papers, and to R. Houart

for his judicious advices.

Fig. 1 Calliotropis illota (Watson, 1886) (from

Watson)

VILVENS

Zetela alphonsi n.sp.

NOVAPEX 3 (2-3): 61-64, 10 juillet 2002

Table 1. - Zetela alphonsi : Shells measurements in mm — sample of 6 specimens.

REFERENCES

Dall, W.H. 1909. Report on a collection of shells

from Peru, with a summary of the the littoral

marine Mollusca of the Peruvian zoological

province. Proceedings of the U.S. National

Museum 37 (1074):147-294.

Dall, W.H. 1919. Descriptions of new species of

Mollusca from the north Pacific Ocean in the

collection of the United States National Museum.

Proceedings of the United States National

Museum 56 (2295): 293-371.

Dell, R.K. 1971. The marine mollusca of the Royal

Society Expedition to Southern Chile, 1958-1959.

Records of the Dominion Museum 7 (17):155-

233;

Dell, R.K. 1990. Antarctic mollusca, with special

reference to the fauna of the Ross Sea. Royal

Society of New Zealand Bulletin 27:1-311.

Forcelli, D.O. 2000. Molluscos magallanicos.

Vazquez Mazzini Ed. Buenos Aires. 200 pp.

Finet, Y. 1995. Marine molluscs of the Galapagos

(Monographs on Galapagos Mollusca N°2).

L'Informatore Piceno. Ancona. 139 pp.

Hickman, C.S. & Mc Lean, J.H. 1990. Systematic

revision and suprageneric classification of

trochacean gasteropods. Natural History Museum

of Los Angeles County Science Series VI+169 pp.

Keen, A.M. 1971. Sea shells of tropical West

America (2"* ed.). Stanford University Press.

Stanford. x + 1064 pp.

Marshall, B.A. 1999. A revision of the recent

Solariellinae of the New Zealand region. The

Nautilus 113 (2):4-42.

Mc Lean, J.H. & Andrade V. 1982. Large

archibenthal gastropods of central Chile. Natural

History Museum of Los Angeles County,

Contributions in Science, 342:1-20.

Quinn, JF. Jr. 1979. Biological results of the

University of Miami deep-sea expeditions. The

systematics and zoogeography of the gasteropod

family Trochidae collected in the Straits of

Florida. Malacologia 19 (1):1-62.

Quinn, JF. Jr. 1991. Lamellitrochus, a new genus of

Solariellinae, with description s of six new

species from the western Atlantic Ocean. The

Nautilus 105 (3):81-91.

Rehder, H.A. 1971. À molluscan faunule from 200 m

off Valparaiso, Chile, with descriptions of four

new species. Proceedings of the Biological

Society of Washington 83 (51):585-596.

Santa Maria, J.B. 1982. Moluscos marinos del norte

de Chile. Catalogo ilustrativo. Vina del Mar.

49 pp.

Watson, R.B. 1886. Report on the Scaphopoda and

Gasteropoda collected by HMS Challenger during

the years 1873-1876. Report on the scientific

results of the voyage of HMS Challenger, 1873-

1876. Zoology 15:1-680.

VILVENS Zetela alphonsi n.sp. NOVAPEX 3 (2-3): 00-00, 10 juillet 2002

Figures 2-6. Zetela alphonsi n.sp.

2-4. Holotype MNHN, Chile, off Chiloé, 15.5 x 11.9 mm.

5-6. Paratype IRSNB, Chile, off Chiloé, 11.5 x 9.8

WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

Review of the genus Pachybathron Gaskoin, 1853

(Gastropoda : Cystiscidae)

Andrew WAKEFIELD

14 Forest Side, Buckhurst Hill, Essex, 1G9 5SL, UK.

Franck BOYER

110, Chemin du Marais du Souci, 93270 Sevran, France.

Tony McCLEERY

The Moat House, Fort Road, St. Peter Port, Guernsey, GY1 1 ZU.

KEY WORDS. Cystiscidae, Pachybathron, external anatomy, radulae, Caribbean Sea, Panamic

Province, new species, sibling species, biogeography.

ABSTRACT. The species of the genus Pachybathron Gaskoïin, 1853 are studied. Four species are

revised, all recorded from the southern Caribbean sea, and a new species, P. olssoni sp. n., is

described from Panama. The external anatomy and the radula of the majority of the animals are

presented. Information is given on the habitat and distribution of each species. The

biogeographical aspects are discussed.

RESUME. Les espèces du genre Pachybathron Gaskoiïin 1853 sont étudiées. Quatre espèces sont

révisées, toutes rapportées du sud de la mer Caraïbe, et une nouvelle espèce, P. olssoni sp. n., est

décrite du Panama occidental. L’anatomie externe et la radula des animaux sont présentées. Des

informations sont données sur l’habitat et la distribution des espèces. Les aspects

biogéographiques sont discutés.

INTRODUCTION

The genus Pachybathron was described by Gaskoin

(1853, p.356-358), and was based upon two species :

P. cassidiforme Gaskoin, 1853 and P. margi-

nelloideum Gaskoin, 1853. P. cassidiforme was

described first, and must therefore be considered as

the type species. This species remained very elusive

for more than a century, with even the holotype being

apparently lost. It was neither figured nor recorded in

collections after its description until a photograph of

a shell from the Coen Collection (HUJ) was

presented by Coomans (1973, p. 12). In the same

paper, Coomans demonstrated that the type locality

recorded by Gaskoin (‘Island of St. Vincent’) should

be interpreted as a Caribbean locality as opposed to a

West African one.

Since its description by Gaskoin, the familial

placement of the genus Pachybathron has been much

discussed. Gaskoin himself (1853, p.356) declined to

associate the genus with the Marginellidae, Cassidae

or Cypraeidae. Coomans (1972) presented, in

chronological order, the different stages of this

discussion by previous authors but failed to add the

important comments of Sowerby (1852, re-ed ) to

this debate. Sowerby stressed the resemblance of the

species he illustrated (P. marginelloides) to those of

the genus Marginella (sensu lato), and he

characterised the original features of the genus

Pachybathron thus : ‘...the columellar lip spread

over the body whorl and the teeth continued across it

in folds.” Sowerby also noted that Marginella

kieneriana Petit, 1838 was close to P. marginelloides

and commented that this could associate the genus

Pachybathron with the ‘Marginellae”.

Six taxa, all described from Caribbean localities,

present a shell morphology which allow their

placement into the genus Pachybathron :

Marginella kieneriana Petit, 1838

Erato cypraeoides C.B. Adams, 1845

Pachybathron cassidiforme Gaskoin, 1853

Pachybathron marginelloideum Gaskoin, 1853

Pachybathron tayrona Diaz and Velasquez, 1987

Microcassis colettae Paulmier, 1997

They all share the shell features of a very long and

narrow aperture, a moderate to heavily callused

columella, callused parietal surface traversed by

continuations of columellar plaits, a strongly

denticulated inner lip, marked transverse lirae

regularly spaced along the internal apertural wall,

and a tendency to form a pattern of chevrons of

various configurations on the body whorl.

During the last 30 years, several partial revisions of

the genus have been published. Coomans (1972 and

1973) recognised two species ; P. cassidiforme

Gaskoin and P. cypraeoides C.B. Adams (Synonym :

P. marginelloideum Gaskoin) assigning them to ‘the

WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

Marginellid genus Pachybathron, related to

Persicula’ (1973, p. 12).

Diaz and Velasquez (1987) attributed the same two

species 10 the ‘Marginellid genus Pachybathron" and

described as new a third species, P. tayrona, from

northern Colombia.

More recently, however, Coovert and Coovert (1995)

declined to assignate generic status to Pachybathron,

for the moment preferring instead to place it within

the genus Persicula as part of the family Cystiscidae

Stimpson, 1865.

The obtaining and observation of living specimens

corresponding to the six taxa quoted above, together

with the re-evaluation of the available type material,

has enabled the current authors to propose a general

revision of the genus Pachybathron.

During the preparation of this work, several new

Pachybathron populations have been discovered off

the coasts of Panama. One is possibly a western

population of P. fayrona, Whereas the other has

sufficient original features to allow its description as

a new species.

MATERIAL AND METHODS

Living specimens of four species of Pachybathron

were collected by several means:

- P. kieneriana Was collected by small boat dredging

in Central Venezuela by F. Boyer and later by F.

Hennequin. T. McCleery also dredged this species in

Central Venezuela, and in addition scuba dived for it

(hand sieve technique) in Los Testigos Islands.

Jamaica

CARTBBEAN SEA

Bocas del

Toro

Archupelago

PSE San Elas

Fig. 1.

Netherlands Antilles 1,1.

- P. cypraeoides Was collected by diving and

snorkelling in the Netherlands and Venezuelan

Antilles by T. McCleery (a suction technique being

used to extract sand and sediment, which was then

hand sieved).

- P. cassidiforme Was collected by diving in

Martinique by P. Clovel, and in St. Vincent,

Grenadines and Grenada by G. Mackintosh.

- P. aff. tayrona was collected by snorkelling (using

suction apparatus and hand sieve technique) in the

San Blas Islands, Eastern Panama by T. McCleery.

Numerous shells of a new species were obtained

from Marcos Alvarez (Panama City). These shells

were presented as several lots, variously labelled as

coming from either the Caribbean region of Bocas

del Toro, the Veraguas region with coasts both on

the Caribbean and Pacific sides of Panama, or three

localities from West Panama namely the Azuero

Peninsula, and the Montuosas and Ladrones Islands.

Taking natural variability into account, these shells

were found to be conchologically conspecific. Due

to the referred localities and to the depths and dates

of the samples taken, most of these data are

considered to be credible, and they are used as such

for the determination of a specific distribution on

both sides of the isthmus of Panama.

Field sketches and photography of the living animals

were performed in most cases, and several specimens

of each species were preserved in 70% isopropyl

alcohol for later radular studies.

The radulae were extracted by E. Rolän and scanning

electron microscope images produced (C.A.C.T.E.,

University of Vigo, Spain.)

ATLANTIC OCEAN

Puerto +

Rico

St. Vincent » bEztado:

Aruba Curacao # The Grenadines

à Roques I] Elanquila @ Grenada

Dr rw > I: Los D

Cabo #æ# Testigos RFES

EBonarre ”

Trinidad

SOUTH AMERICA

Map showing the area of distribution of the genus Pachybathron

WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

ABBREVIATIONS AND TERMINOLOGY

The following abbreviations are used in this paper:

NHM: Natural History Museum, London, England.

HUJ: Hebrew University of Jerusalem, Israel.

IRSNB: Institut royal des Sciences naturelles de

Belgique, Bruxelles.

MCZ: Museum of Comparative Zoology, Harvard,

Massachusetts, U.S.A.

MHNLR: Muséum d'Histoire Naturelle, La Rochelle,

France.

MNHN: Muséum national d'Histoire naturelle, Paris,

France.

MOL: Instituto de Investigaciones Marinas de Punta

de Betin, ‘INVEMAR, Santa Marta, Colombia.

SMF: Seckenberg Museum, Frankfurt, Germany.

ZMA: Zoologische Museum, Amsterdam, The

Netherlands.

AWC: Andrew Wakefield Collection.

FBC: Franck Boyer Collection.

TMC: Tony McCleery Collecton.

JCC: Jacques Colomb Collection, Marseilles, France

LBC: Luigi Bozzetti Collection, Milan, Italy.

ABC: The Islands of Aruba, Bonaire and Curacao

which together comprise the Netherlands Antilles.

For the purposes of this paper, ‘specimen’ refers to a

live taken specimen, whereas ‘shell’ refers to a dead

collected shell.

SYSTEMATICS

Family CYSTISCIDAE Stimpson, 1865

Genus Pachybathron Gaskoin, 1853.

Type species : Pachybathron cassidiforme Gaskoin,

1853 (subsequent designation by Adams and Adams,

1858).

Pachybathron kieneriana (Petit, 1838)

Figs. 2-3, 18, 22-27, 50-51.

Marginella kieneriana Petit, 1838, p.20.

Type material. One syntype, considered to be the

holotype, in MNHN, 11.5 x 7.6mm (Figs. 2-3).

Other material examined. - Puerto Frances, Cabo

Codera, Central Venezuela, dredged and dived

September 1999 on coarse muddy sand at 20 m. 8

adult specimens (8.3 x 5.2mm to 12.0 x 8.7mm),

AWC.

- Puerto Frances, Cabo Codera, Central Venezuela,

dredged February 1997 in sticky black mud at 25 m,

1 adult specimen (9.65 x 5.9 mm), AWC.

- Puerto Frances, Cabo Codera, Central Venezuela,

dredged and dived September 1999 on coarse muddy

sand at 20 m, 16 adult specimens (9.3 x 5.9 mm to

13.3 x 9.5 mm), TMC, (Figs. 22-24).

- Puerto Frances, Cabo Codera, Central Venezuela,

dredged February 1997 from coarse coral sand at 15

m to sticky mud at 25 m, 12 adult specimens (9.1 x

5.7mm to 13.4 x 9.1mm), 3 juvenile specimens (6.0 x

4.0mm to 12.5 x 8.5 mm) and 1 adult shell (12.0 x

7.85mm), FBC.

- Isla Grande, Is. Los Testigos, Venezuela, dived

June 1999 on coarse clean sand at 20 — 21m, 4 adult

specimens (8.0 x 4.5mm to 8.9 x 5.3mm), AWC, and

3 adult specimens (7.5 x 4.4 mm to 8.6 x 5.1 mm),

TMC, (Figs.25-27).

- Tobago, 2 large shells, Dautzenberg Collection

(IRSNB).

Original description. * Marginella kieneriana, Petit.

Long:12 mill. Larg:9 mill. — Testa parva piriforme,

fulva, maculis albis transverses per quarto series

dispositis ornate : spira brevissima, exsertiuscula,

labro crasso, vix intus crenulato plicis columella

octonis.”

Complementary Notes. Shell glossy, rarely

completely smooth, usually with heavy growth lines.

Medium to large sized for the genus (length 8mm to

12.5mm, possibly larger) comprising 2.5 whorls

(excluding protoconch of 1.5 whorls) and presenting

variety of outlines from globular, ovoid or pyriform,

to sub-cylindric (Los Testigos). Spire slightly

elevated and smooth with glazed suture. Aperture

curved, narrow and parallel sided with very little

anterior flare. Inner aspect of lip with 22-24

denticles. Lip thickened internally with no external

callus formation. 8 columellar plaits. Weak parietal

callus traversed by 8 plaits. Strong parietal callus

ridge at entrance to aperture. AIl of parietal callus

blends into glaze of body whorl with no posterior

groove to separate the two.

Colour variable. Base colour tan. Olive green bands

encirele body whorl either concentrated as one main

central band, or as three or four narrower ones. These

bands form the background to the spiral chevron

pattern. Spiral chevron pattern of 5-7 evenly spaced

rows from sub-sutural level to anterior extremity.

Pattern comprises white blotches with rounded to

pointed black chevrons pointing towards the growing

edge of the shell. Nearer to the lip, extra white marks

and associated chevrons are formed between the

regular rows, disrupting the regular pattern. Spire

colour reddish tan, becoming paler towards

protoconch, which is translucent at the nucleus (Figs.

22-27). Specimens which retain the basic pattern but

are weakly coloured, appearing yellowish to pinkish

tan overall, are often found amongst the typical

darker shells.

The following description of the animal is based on

the study of several specimens from Puerto Frances

(Fig. 18): Typical Cystiscid Type 4 animal (after

Coovert & Coovert 1995). Foot reddish brown with

opaque white blotches, some fusing together to form

larger irregular edged patches. Bilobed head opaque

white, each half having a streak of reddish brown

running from slightly behind the small black eye,

WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

medially and anteriorly to meet the other at its tip.

Eyes situated at the base of the two short reddish

brown tentacles. Siphon short, opaque white, and

fringed at its tip with reddish brown.

The radula was extracted from a specimen from

Puerto Frances (Figs. 50-51) : Type 3 Cystiscid

radula (after Coovert & Coovert 1995). Very thin (10

um), cord like, uniserial radula with approximately

400 strongly arched and overlapping rachidian plates.

Each plate has 9 sharply pointed cusps, the strongest

being the single central cusp. The lateral cusps are

shghtly smaller, with the most lateral tooth being

diminutive.

Type locality. Les Plages de la Guayra, Venezuela.

Distribution. The species has been found from La

Guayra (central mainland coast of Venezuela) to Los

Testigos Islands and Tobago. Due to its various

habitats, it could well have a wider distribution in

Venezuelan mainland coastal locations as well as in

offshore island groups. Unconfirmed reports that this

species has been found in Brazil require verification.

Habitat. Available records indicate an important

varlability of habitats, from sticky black mud to

coarse clean white coral sand. The bathymetric range

of distribution of the species is currently known to be

from 15 to 25 m.

Remarks. This species has the most distinctive shell

of all the group, and has until now been considered to

be a member of the genus Persicula by most authors.

It does possess, however, all of the defining

characteristics of the genus Pachybathron (ovoid to

pyriform shell shape, slightly elevated spire, parietal

callus with horizontal plaits, labial denticles,

chevronned pattern, body whorl often with numerous

growth lines, and similarities in the chromatism of

the soft parts of the animal), and therefore the authors

consider that its correct taxonomic assignation should

be within this genus.

The central Venezuelan coastal populations exhibit

considerable variability of size, morphology and

colour pattern. Specimens from Los Testigos Islands,

however, all appear to have more constant characters

(size, very regular rows of chevrons, and a very dark

brown parietal callus and anterior notch). This may

be a phenotypic expression of the smaller genetic

pool found within relatively smaller isolated colonies

in the offshore island localities. The minor

differences in shell shape and colour pattern between

Figures 2-10.

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

both populations are not considered of sufficient

magnitude to permit separation into different species,

unless further analysis of the animals suggests this is

indicated.

Pachybathron cypraeoides (C.B. Adams, 1845)

Figs. 4-10, 19, 28-37, 52-53, 56-58.

Erato cypraeoides C.B. Adams, 1845, p.1-2.

Pachybathron marginelloideum Gaskoin, 1853, pp.

357 — 358, figs. 4-6.

Type material. Two syntypes are deposited in MCZ

and were examined by the authors. Though badly

worn and faded, one syntype is clearly recognizable

as being representative of the species (Figs. 5-7).

This specimen was figured as a black and white

photograph by Clench and Turner (1950, p.271, fig.

11) in their revision of the species described by

Adams. This selection by Clench and Turner must be

considered as the designation of a lectotype. The

paralectotype is in fact the shell of another species,

Persicula chrysomelina (Redfield, 1848). The

lectotype, ref. No. 186065, measures 7.9 x 5.2mm.

The holotype of P. marginelloideum is recorded as

being deposited in Cabinet Gaskoin in the NHM but

no trace of it can be found there and so it is presumed

lost. It measured 7.1 x 4.1mm.

Other material examined. - Aruba harbour,

Netherlands Antilles. Suction, followed by hand

sieving, September 1999 from grass and rubble at

0.5-Im, numerous adult specimens (6.5 x 4.5mm to

8.5 x 5.5mm.), TMC, (Figs. 28-30).

- Aruba harbour. Dived and hand sieved September

1999 from sand and coral rubble at 0.5 — 1m, two

adult specimens (measuring 8.4 x 5.4mm and 7.4 x

4.8mm), and one juvenile specimen, used for radular

examination (5.2 x 3.2mm), AWC.

- East of Aruba harbour. Snorkelled and hand sieved

September 1999 from sand and coral rubble at 2-

2.5m, one adult specimen, TMC.

- Malmok bay, Aruba. Dived and hand sieved

September 1999 from sand and coral rubble at 7- 8m,

seven adult specimens (6.0 x 4.0mm to 7.4 x 4.6mm),

TMC, (Figs 31-33).

- Malmok, Aruba. On clean sand at 7m. One

specimen (6.8 x 4.Imm) and one shell (8.0 x

5.15mm), FBC.

- Arasji, Aruba. Two adult shells and two juvenile

shells, FBC.

2-3. P. kieneriana Petit, 1838, syntype (MNHN), 11.5x7.6mm; 4. P. cypraeoides C.B. Adams, 1845, La

Blanquilla, Venezuelan Antilles, 4.35x2.8mm; 5-7. P. cypraeoides C.B. Adams, 1845, lectotype (MCZ),

7.9x5.2mm; 8-10. P. cypraeoides C.B. Adams, 1845, original figures.

WAKEFIELD, BOYER, MCCLEERY Genus Pachybathron NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

- Lac Bay, Bonaire, Netherlands Antilles. Snorkelled

and hand sieved 1998 from grass and rubble at 2m,

one adult specimen (7.0 x 4.1mm), TMC.

- Curacao, Netherlands Antilles. Eight slender shells

(5.3 x 3.2mm to 6.2 x 3.6mm), LBC, (Figs. 34-35).

- Los Roques, Venezuean Antilles. Snorkelled and

hand sieved July 1999 from white sand at Im, one

adult specimen measuring (6.5 x 4.1mm), TMC,

(figs. 36-37).

- La Blanquilla, Venezuelan Antilles. Dived at 12m

on sand, one subadult specimen (4.35 x 2.8mm), JCC

(Fig. 4).

- Many specimens and shells in ZMA, all listed in

Coomans (1972, p.91).

Original description. ‘Erato (?) cypraeoides M. t.

solida, alba; anf. ifra suturam fusco-canaliculatis:

spira plana, parva; apertura lineari, pro (2) funde

emarginata; labro extra crasso, intus exile crenulato:

labio per totum transversum exile plicato. Long. .325

poll.; lat. .2 poll. Jamaica.”

Original figures. See Figs. 8-10.

Complementary notes. Shell glossy, medium to

heavily callused and ribbed with growth lines.

Medium sized (length from 5.2 mm to 8.5 mm) for

the genus. Comprising 2 whorls (excluding

protoconch), and presenting an ob-ovate outline.

Spire slightly depressed with a low lens-shaped,

translucent white protoconch, comprising 1.5 whorls.

Suture irregular, glazed over.

Aperture narrow, slightly curved and parallel sided

except for a slight flare anteriorly. Inner aspect of lip

has 15-18 denticles, obliterated and uncountable in

some specimens due to very heavy labial callus,

which extends posteriorly slightly beyond spire

height. 5-9 columellar plaits. Strong parietal callus

with 12-14 plaits lying horizontally across it,

commencing from the parietal callus ridge. Edge of

parietal callus demarcated in its posterior half by a

deep groove. Anterior half gradually merges into

glaze of body whorl.

Base colour of shell pure white. General appearance

is yellowish off-white, due to presence of fine axial

pattern which extends to cover the whole of the body

whorl except the internal and external labial callus,

and the parietal callus and columella which are all

white. Microscopically the fine axial yellow brown

lines are drawn sharply towards the lip forming 4-5

rows of sharp arrowheads. Macroscopically these are

visible as spiral bands which appear slightly darker

than the area in between. In many specimens the

Figures 11-17.

pattern can be extremely faint, and is then reduced to

4 -5 fine double rows of tiny brown spots

representing the bases of the arrowheads. Occasional

shells are almost pure white but there is rarely

absolutely no trace of a pattern. Spire colour varies

according to locality ; specimens from the

Aruba,Malmok Bay population all have dark brown

spires (excluding the protoconch which is always

translucent white), whereas other populations from

Aruba have colour absent from the spire (Figs. 28-

33). The specimen from Klein Curacao (Coomans

1972, p. 92) also has a dark spire. This feature 1s

therefore in need of further study.

The following description of the animal is based

upon the study of several specimens from Aruba

(Fig. 19) : Typical Cystiscid Type 4 Animal (after

Coovert & Coovert, 1995). Bilobed head, and foot

peachy — orange, finely fringed with bright orange.

Midline and posterior part of foot also marked with

bright orange. Tentacles short, completely orange.

Eyes tiny, black, situated medially at base of

tentacles.

The radula was extracted from a sub-adult specimen

from Aruba (Figs. 52-53) : Type 3 Cystiscid radula

(after Coovert & Coovert 1995): Very thin (9 um),

cord like, uniserial radula with approximately 400

strongly arched and overlapping rachidian plates.

Each plate has 7 - 9 sharply pointed cusps along the

cutting edge, the strongest being the single central

cusp. The lateral cusps can be of unequal size and are

slightly smaller, the most lateral tooth being

diminutive.

Type locality. Jamaica.

Distribution. The occurrence of P. cypraeoides in

the quoted type locality of Jamaica has never been

verified, and in fact remains distinctly unlikely. This

can be concluded for the following reasons. Firstly,

all the records from the date of the original

description to the present day are from the ABC

islands of the Netherlands Antilles, except two new

recent records (Figs. 4, 36-37) from the neighbouring

Venezuelan Antilles (Los Roques and La Blanquilla).

Secondly, the paralectotype of P. cypraeoides 1s in

fact a shell of Persicula chrysomelina, well known

as a Netherlands Antillean endemic species. Finally,

in the same paper C.B. Adams described another

species (Volvarina rubella C.B. Adams, 1845) citing

the same type locality as P. cypraeoides. V. rubella is

well known from the Southern Caribbean but like P.

cypraeoides has never been recorded from Jamaica

since its original description.

11-13. P. cassidiforme Gaskoin, 1853, original figures; 14-15. Microcassis colettae Paulmier, 1997, original

figures; 16-17. P. tayrona Diaz & Velasquez, 1987, paratype MOLS95, 8.7x5.0mm.

WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

It can therefore be assumed that C.B. Adams

described P. cypraeoides from a lot of shells

collected in the ABC islands. At this period in

history, there was an active shipping trade between

Jamaica and the ABC Island group, and Adams could

have procured this particular lot of shells from a

merchant vessel or similar source. The lectotype has

a distinctly dark spire, and it is possible that it could

have originated from the dark spired Malmok

population (Aruba).

Based upon current collecting records, and the above

albeit circumstantial evidence, we propose to

provisionally consider that the distribution of P.

cypraeoides 1s restricted to the Southern Antilles

(ABC Islands to La Blanquilla). The apparent

scarcity of the species in the Venezuelan Antilles

(although this area has been reasonably well sampled

during the last decade) is probably an indication that

this archipelago could represent the extreme eastern

edge of the geographical range of the species. The

practically unexplored Paraguana Peninsula of

mainland Venezuela is in close proximity to the

Netherlands Antilles, and the presence of supposed

endemic species from the ABC Islands must be

investigated there. For example, two specimens of

Volvarina vokesi de Jong & Coomans, 1988 were

collected in the Park of Moroccoy, State of Falcon,

100km south of Bonaire Island (AWC).

In the collection of one of the authors (FBC) is a

shell resembling P. cypraeoides which belonged to a

lot of dead collected material from an old collection,

with the label ‘Callao, Peru’. This shell is identical to

the material examined from the ABC islands, and

was mixed with shells of several undetermined

species of Volvarina, apparently not from the ABC

islands. Future sampling of the molluscan fauna

along the western coasts of Colombia, Ecuador and

Peru should reveal if a sibling species of P.

cypraeoides really does occur in the Panamic

province.

We propose the Islands of the Netherlands Antilles

(Aruba, Bonaire, Curacao) as the revised type

locality of P. cypraeoides.

Habitat. In coral rubble and Thalassia beds.

Compact communities were observed in Aruba at

0.5-3m and again at 7-8m. Coomans (1972, p. 92)

quotes a fresh specimen taken at 60m near Klein

Curacao (deposited in NNM).

Figures 18-33.

Remarks. The lot of small slender shells from

Curacao (Figs. 34-35) could simply represent a local

form of P. cypraeoides. The obtaining of live

specimens of this population will be necessary to

determine if this is the case or if they are in fact

morphological intergrades between the eastern P.

cassidiforme and the western P. tayrona. In addition,

the two specimens from Los Roques and La

Blanquilla (Figs. 4, 36-37), though assigned here to

the taxa P. cypraeoides, are certainly not typical of

that species. They have some morphological affinity

with P. cassidiforme, have a translucent white shell

with seven narrow opaque spiral bands and only a

faint suggestion of the usual zigzag pattern. A picture

appears to be emerging, therefore, of a cline of

Pachybathron populations distributed right across the

Southern Caribbean region, comprising several

‘ sibling species distributed at geographic intervals.

The original description and the type figures of P.

marginelloideum (Figs. 8-10) are a perfect match for

the species P. cypraeoides, thereby making P.

marginelloideum its junior synonym. This synonymy

has been previously stated by Coomans (1972).

Pachybathron cassidiforme Gaskoin, 1853

Figs. 11-13, 20, 38-39, 54.

Pachybathron cassidiforme Gaskoin, 1853, pp. 356-

357, figs 1-3.

Microcassis colettae, Paulmier 1997, pp. 733-748.

Figs. 1-4, 9.

Type material. P. cassidiforme is recorded as being

deposited in Cabinet Gaskoin, British Museum, but

there is no trace of the type specimen (nor any other

specimen of this species) in the collections at the

NHM. The holotype (which measured 6.3 x 3.8mm)

is therefore presumed lost. The specimen from HUJ

(ref. No. 31.587/1 ex. Coll. Coen 2008), figured in

Coomans (1973, p. 12) is here designated as neotype.

Length 6.8mm. Locality: Grenadines, Lesser

Antilles.

The type material of M. colettae is as follows:

- Holotype.(Ref. No. MG1): Insular shelf, Martinique

14 36° 06 N-60 46° 44 W dredged in white sand at

60m. Adult shell (7.55 x 4.35mm), MHNLR.

- Paratypes (Ref. No. MG2): Same locality as

holotype. Six adult shells (from 5.75 x 3.5mm to 7.4

x 4.2mm), MHNER.

18. P. kieneriana Petit, 1838, live animal; 19. P. cypraeoides C.B. Adams, 1845, live animal. 20. P. cassidiforme

Gaskoin, 1853, live animal; 21. P. aff. tayrona Diaz & Velasquez, 1987, live animal; 22-24. P. kieneriana Petit,

1838, Puerto Frances, Cabo Codera, Venezuela, 10.5x6.8mm. 25-27. P. kieneriana Petit, 1838, Isla Grande, Los

Testigos, Venezuela, 8.7x5.1mm; 28-30. P. cypraeoides C.B. Adams, 1845, Aruba Harbour, Netherlands

Antilles, 8.5x5.5mm; 31-33. P. cypraeoides C.B. Adams, 1845, Malmok, Aruba, Netherlands Antilles,

7.4x4.6mm.

WAKEFIELD, BOYER, MCCLEERY Genus Pachybathron NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

Other material examined. - Isle de Ronde, Grenada

(north end of bay), dived 1998 in coral rubble at 7m,

one adult shell (5.8 x 3.6mm), TMC, (Figs. 38-39).

- Anse d’Arlet, Martinique. Scuba and hand sieved

January 1999 under flat coral rock in sand at 22-30m,

one adult specimen (5.2 x 3.6mm), TMC ; two adult

specimens (6.0 x 3.8mm and 5.65 x 3.5mm), AWC ;

one specimen (3.3mm long) and four shells (from 5.1

to 5.35mm in length), FBC.

- Baliceaux Island, St. Vincent. Dived and sieved

May 1997 from sand at 13m, one specimen

(measuring 6.1 x 3.9mm), AWC.

- Frigate Is, Grenada. Dived at night in sand at 8m.

One specimen (5.3mm long), FBC.

Original description. ‘Shell subcylindrico-ovate,

opake-white colour, three continuous bands a few

shades darker than the shell traverse the dorsum ;

dorsum coarsely striated longitudinally ; spire rather

depressed, subacuminated, volutions four, irregularly

crenulated ; the posterior edge of the last whorl forms

a coronated ridge at the base, of which a deep

depression surrounds the shell terminating at the

outer part of the aperture : base rather round, broad

and very thick, abrupt at its outer border, and extends

over the anterior third of the side of the shell,

terminating on the columellar extremity ; aperture

rather narrow, slightly curved ; outer lip thick and

finely denticulated along the inner edge ; columellar

side, about twelve or fourteen distant linear teeth

transvers the entire base and terminate on the inner

margin of the columellar groove ; columellar groove

shallow ; extremities flat (perpendicularly), rather

prominent and keeled, posterior end of aperture not

rostrated ; channel deep and rather short.”

Original figures. See Figs. 11-13 for P. cassidiforme

and Figs. 14, 15 for M. colettae.

Complementary notes. Shell small for the genus

and has the original feature of an unglazed body

whorl, even in fresh specimens. The regular growth

lines are the most numerous of the genus, and result

in a distinctly axially costate body whorl.

The following description of the animal is based

upon a study of a single specimen from Martinique

(Fig. 20) : Typical Cystiscid Type 4 animal (after

Coovert & Coovert 1995). The foot, which does not

extend very far out of the shell, is translucent grey

with yellowish patches of various sizes, and on the

specimen examined, two large yellow patches

Figures 34-49.

situated either side of the midline on the posterior

part of the foot. Tentacles translucent grey except the

middle third which is bright orange. Lateral surfaces

of each head lobe also bright orange, this extending

to surround the tiny black eyes which are situated on

the medial aspect of the base of the tentacles.

The radula was extracted from a specimen from

Martinique (Figs. 54) : Type 3 Cystiscid radula (after

Coovert & Coovert 1995): Very thin (7 um), cord

like, uniserial radula with 425 strongly arched and

overlapping rachidian plates. Each plate has 9

sharply pointed cusps, the strongest being the single

central cusp. The lateral cusps are slightly smaller,

the most lateral tooth being diminutive.

Type locality. Island of St. Vincent.

Distribution. Known only from the southeastern

Antilles, from Martinique to Grenada.

Habitat. Found by divers in coral rubble and sand

under coral slabs at 7-30m. The species seems to be a

sand dweller, moving at night (G.Mackintosh, pers.

comm.). Its bathymetric range of distribution must be

considered as deeper than this as M. colettae was

found as deep as 123m (Paulmier, 1997).

Remarks. Living specimens of this species were

only discovered in the Grenadines as recently as

1997 by G. Mackintosh, and a short time later by P.

Clovel in Martinique. The axial costae, unglazed

body whorl and dark blotch at the base of the

columella enable this species to be easily identified,

and there does not appear to be any distinguishing

feature by which specimens from Martinique can be

separated from Grenadian examples.

Paulmier (1997) makes no mention of the possibility

of the animal being a Marginelliform gastropod, but

instead considers it to be associated with the

Cassidae. The study of dead shells only may be

partly responsible for this. Paulmier therefore not

only assigns an invalid species name, but also

erroneously creates a new Genus, Microcassis.

Nevertheless, Paulmiers work is useful in that it

presents new information on the bathymetric range of

the species, which can be considerably extended to

123 metres. It has been confirmed herein that the

species is in fact a Cystiscid, and is the same as that

described in 1853 by Gaskoin as Pachybathron

cassidiforme. Microcassis colettae Paulmier, 1997 is

therefore a junior synonym of P. cassidiforme,

Gaskoin, 1853.

34-35. P. cypraeoides C.B. Adams, 1845, Curacao, Netherlands Antilles, 6.2x3.6mm; 36-37. P. cypraeoides

C.B. Adams, 1845, Los Roques, Venezuelan Antilles, 6.5x4.1mm; 38-39. P. cassidiforme Gaskoin, 1853, Isle de

Ronde, Grenada. 5.8x3.6mm; 40-42. P. aff. tayrona Diaz & Velasquez, 1987, Porvenir, San Blas Archipelago,

East Panama, 6.5x3.9mm; 43-45. P. olssoni sp. n., Escudo de Veraguas Is, Bocas del Toro, East Panama.

Holotype (NHM) 5.0x3.4mm; 46-47. P. olssoni sp. n., Escudo de Veraguas Is, Bocas del Toro, East Panama.

Paratype 6, 5.4x3.5mm,; 48-49. P. olssoni sp. n., Peninsula de Azuero, West Panama, Paratype 7, 5.9x4.0mm.

WAKEFIELD, BOYER, MCCLEERY Genus Pachybathron NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

Pachybathron tayrona Diaz & Velasquez, 1987

Figs. 16-17, 21, 40-42, 55.

Pachybathron tayrona Diaz & Velasquez, 1987, pp.

217-221.

Type material. - Holotype (Ref. no. 305966): Bahia

de Chengue, Colombia, in Thalassia meadow at

0.5m. Adult specimen (10.2 x 5.8mm), SMF.

- Paratypes (Ref. nos. 305967 / 305968): Bahia de

Chengue, Colombia, in Thalassia meadow at 0.5m.

One adult shell (8.2 x 4.7mm), and one juvenile shell

(no dimensions given), SMF.

- Paratypes (Ref. nos. 895 / 896): Bahia de Chengue,

Colombia, in Thalassia meadow at 0.5m. One adult

specimen (9.4 x 5.2mm), and one adult shell (11.5 x

6.3mm), MOL.

Other material examined - Porvenir, San Blas

Archipelago, East Panama. In erosions in Thalassia

beds in 1.5-2m. 36 adult specimens (from 5.6 x

3.4mm to 7.3 x 4.3mm), TMC, (Figs. 40-42) ; 11

specimens (from 6.4 x 3.6mm to 7.0 x 4.1mm),

AWC; two adult and one juvenile shells, FBC.

Original Description. ‘Shell medium sized (up to

11.5mm total length), obovated, thick shelled, glossy,

with a rather blunt, low but unconcealed spire.

Nucleus rather blunt, tan, apparently with two

whorls, and covered by an irregular line. Surface of

the spire may present some enamellous bulges along

the suture. Body whorl smooth and polished; some

fresh specimens (over 5.0mm long) bear four to six

fine longitudinal scissulations on the body whorl,

which can be better recognised on the dorsal side and

near the outer lip.

Aperture rather narrow, extending about 6/7 of total

length. It becomes anteriorly somewhat wider, twists

lightly to the left and forms a distinct anterior notch,

visible from the dorsal view. A callous formation

along the parietal wall is poorly developed but

visible: it extends above the end of the aperture

towards the upper suture of the penultimate whorl

and is minutely pitted.

There are nine to thirteen linear folds on the pillar

whorl, the upper ones becoming nearly or completely

obsolete. These pillar folds continue undiminished

into the interior of the columella. Outer lip thickened,

minutely pitted and bearing 18 — 20 denticulations.

Shell tan to dirty white, highly polished, spire

occasionally with light brown to chestnut streaks or

mottlings at the suture. Body whorl shows numerous

microscopical wavy or interrupted longitudinal

Figures 50-55.

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

brown lines, which become widely spaced towards

the outer lip. Some side by side running and

uninterrupted lines form simultaneously a single

arrow — pointed mark at regular intervals. Some other

lines are weaker coloured and interrupted at the same

intervals where the darker ones form the arrow — like

flexures, leaving thus whitened spaces between two

arrow series. Macroscopically this colour pattern is

expressed as eight to nine spiral rows of whitened

mottlings alternating with brown stains on a tan

background. Some rows are usually more

conspicuously coloured than others. In young and

worn specimens, the colour pattern fades away,

leaving a light tan or pure white shell.”

Original figures. The holotype (SMF 305966) and

two paratypes (SMF 305967 and SMF 305968) are

figured in the original paper.

Another paratype (MOL 895), 8.7x5.0mm, is figured

here (Figs. 16-17).

Complementary notes. Northern Colombian shells

are large for the genus (up to 11.5mm long) whereas

San Blas specimens (Figs. 40-42) are much smaller

in size (up to 7.3mm). Both of these populations have

a L/W ratio of 55-57% which is slender for the genus

and comparable only to the two specimens of P. cf.

cypraeoides from Paloe Lechi, Bonaire (Coomans,

1972 ) at 56.5% and the eight slender shells of P. cf.

cypraeoides from Curacao in LBC (Figs. 34-35).

These may be found to form a link in a cline of

morphologies from P. cypraeoides to P. tayrona.

It is well known that shells from continental

mainland coastal areas can be larger than those of

populations from offshore islands where food supply

is less abundant.

P. tayrona from Northern Colombia is typically light

tan to cream across the whole of the body whorl

(Diaz & Velasquez 1987). This is the impression

when viewed from a distance, but on closer

inspection it becomes apparent that it is the pattern of

axial wavy lines which gives this effect, the base

colour being either white or off white. In the vast

majority of the San Blas specimens this pattern 1s

reduced to a central band occupying 70% of the body

whorl. The pattern is completely absent from the

posterior 20% of the shell and also from the anterior

10%. The axial wavy lines in the central section lie

parallel to each other and at 6 or 7 places along their

length are drawn forwards to form an extra large

arrowhead shape. This has the effect of creating 6 or

7 spiral rows of arrowheads encircling the central

50-51. P. kieneriana Petit, 1838, Radula, scale bar 5 um; 52-53. P. cypraeoides C.B. Adams, 1845, Radula, scale

bar 5 um; 54. P. cassidiforme Gaskoin, 1853, Radula, scale bar 5 um; 55. P. aff. tayrona Diaz & Velasquez,

1987, Radula, scale bar 10 um.

PETRE SN PATES

WAKEFIELD, BOYER, MCCLEERY Genus Pachybathron NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

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WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

band. There are generally 2 or 3 irregular blotches of

the same colour on the arrowhead row closest to the

posterior end, and the same on the penultimate row at

the anterior end. When viewed from a distance, these

blotches appear to merge somewhat, giving the

impression of two rows of brown markings. On

occasional specimens, very faint traces of axial

pattern can be detected in the light zones, but never

on the spire itself, yet even then the overall effect of

a darker central band remains. The intensity of the

pattern varies slightly with the growth lines which

form slightly raised ribs. There are only two

specimens from San Blas which do not follow the

above rule and exhibit a full length light brown axial

pattern with 8 or 9 rows of chevrons (as in

Colombian examples).

There are 16 or so countable labial denticles, fading

out anteriorly into 6-8 internal lirae, as the lip begins

to flare.

The following description of the animal 1s based on a

study of several animals from San Blas (Fig. 21) :

Typical Cystiscid Type 4 animal (after Coovert &

Coovert 1995). AI soft parts are medium grey-brown

with small paler blotches. There are two larger pale

patches situated either side of the midline on the

posterior part of the foot. The short tentacles are a

darker brown colour. The overall appearance 1s of

rather a drab looking animal compared to the bright

orange markings of some other species in the genus.

Radulae were extracted from two specimens from

San Blas (Fig. 55) : Type 3 Cystiscid radula (after

Coovert & Coovert 1995): Very thin (12 um) cord

like uniserial radula with, in the two specimens

examined, 298 and 310 strongly arched and

overlapping rachidian plates. Each plate has 9

sharply pointed cusps, with the central cusp being the

strongest.

Type locality. Bahia de Chengue, Parque Nacional

Natural Tayrona, Caribbean coast of Colombia.

Distribution. Initially thought to be restricted to the

Northern Caribbean coast of Colombia, between the

Bays of Santa Marta and Nenguange, but recently a

population has been found in the San Blas Islands

(East Panama). Once the population from San Blas

was discovered at Porvenir, a search for other

populations occupying similar habitat was instigated

throughout the rest of the northwestern part of the

archipelago, with no success. This population

therefore seems quite isolated. However, it remains

possible that other populations occur between the San

Blas Archipelago and Santa Marta, as the intervening

coastline has never been sampled for micromollusces.

Habitat. In Northern Colombia, living specimens are

found only in turtle grass beds (Thalassia

testudinum) and under corals in shallow water to 3

metres. The specimens from the San Blas

Archipelago were found in the banks of pit-like

erosions In 7halassia beds, amongst its roots in 1.5-2

metres.

Remarks. The differences between the populations

of San Blas and Santa Marta are mainly based on

features not considered to be of major taxonomic

importance (minor colour variation and size

differences). The authors therefore feel that further

exploration of the Colombian coastline West of Santa

Marta is required before the specific status of both

populations can be clarified. It is therefore proposed,

for the time being, to refer to the San Blas population

as P. aff. tayrona.

Pachybathron olssoni sp. n.

Figs. 43-49.

Type material. -Holotype. Escudo de Veraguas

Island, Bocas del Toro Province, East Panama

(Caribbean). Dredged 40-490 ft. in sand and red mud

between 1976 and 1982. Adult shell (5.0 x 3.4 mm),

NHM ref. no. 20001283, (Figs. 43-45).

-Paratype 1. Same locality as holotype. Adult shell

(5.3 x 3.45 mm), NHM ref. no. 20001284.

-Paratypes 2 - 5. Ladrones Island, Gulf of Chiriqui,

West Panama (Pacific). Dredged 200-300 ft. 1988.

Four adult shells (5.6 x 3.5 mm, 5.4 x 3.4 mm, 5.5 x

3.5 mm, 5.0 x 3.2 mm), TMC.

-Paratype 6. Same locality as holotype. Adult shell

(5.4 x 3.5 mm), TMC, (Figs. 46-47).

-Paratype 7. Peninsula de Azuero, West Panama.

Dredged 200-400 ft. on sand and mud. 1986. Adult

shell (5.9 x 4.0 mm), TMC, (Figs. 48-49).

-Paratype 8. Ladrones Island, Gulf of Chiriqui, West

Panama. One adult shell (5.6 x 3.7 mm), AWC.

-Paratype 9. Ladrones Island, Gulf of Chiriqui, West

Panama. One adult shell (5.25 x 3.1mm), FBC.

Other material examined. - Veraguas Province,

without further precision. Also Peninsula de Azuero

and Montuosas Islands, West Panama (see section on

Materials and Methods). Dredged in 30-120m.

Eighty nine shells (from 5.0 x 3.2 mmto 7.3 x 4.5

mm), IMC.

Description. Shell glossy and small for the genus

(length from 5 to 7.3 mm, averaging 5.4mm) with an

average L/W ratio of 65%, comprising two whorls

(excluding protoconch), presenting an outline

varying from globular to oval-pyriform. Spire

slightly elevated with a distinct, slightly irregular

suture. Smooth, translucent, slightly raised

protoconch of 1.5 whorls.

Aperture narrow and curved, approximately 9/10 of

the total length of the shell, parallel sided for the

posterior half, then flaring a little anteriorly. Inner

aspect of lip has approximately 22 fine denticles

which vary in size. Anteriorly the denticles are fine

and small, gradually becoming larger and coarser

centrally, then are fine and irregularly spaced

WAKEFIELD, BOYER, MCCLEERY Genus Pachybathron NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

56 57

10 um

Figures 56-58

56. Variation in a series of 3 rachidian plates (non-repeating pattern) of P. cypraeoides C.B. Adams, 1845.57.

End view of single rachidian plate of P. cypraeoides C.B. Adams, 1845; 58. three dimensional representation of

a single rachidian plate of P. cypraeoides C.B. Adams, 1845.

WAKEFFIELD, BOYER, MCCLEERY

Genus Pachybathron

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

towards the posterior end of the lip. Internal lirae

visible through the thin shell of the body whorl on

dorsal aspect.

Outer aspect of lip weakly callused, with a shallow

external groove running its full length, and

continuing around the siphonal notch. Siphonal notch

‘V’ shaped and deep, curving towards central axis of

shell when viewed from anterior end.

Anterior half of columella internally bearing 5 or 6

distinct plaits, which fade out in the posterior half.

Plaits emerge more clearly on apertural parietal

callus ridge and extend across parietal callus almost

horizontally as 10-12 evenly spaced plaits.

Edge of parietal callus demarcated along its posterior

third by a deep groove. Anterior two thirds merges

smoothly into body whorl as a clear glaze.

Colour pattern consists of light yellow-brown zig-

zag pattern on a pale background, extending the full

length of the shell giving an overall impression of a

golden brown shell (in the majority of unfaded

shells). The pattern varies in detail but generally

comprises 6 or 7 (occasionally 8) evenly spaced rows

of arrowhead marks which point towards the outer

lip. Lip, columella and columellar half of parietal

callus opaque white. Base of columella with a

distinct brown blotch.

Animal and radula not seen.

Type locality. Bocas del Toro Province, East

Panama (Caribbean).

Distribution. À number of identical shells selected

from several tens of examples were labelled as

coming from Ladrones Islands and Peninsula de

Azuero on the Pacific side of Panama, more or less

opposite Bocas del Toro Province. These are almost

identical to the Caribbean shells, falling within their

size range and exhibiting comparable shell features.

In the present state of our documentation, it seems

that the new species is distributed along both

Caribbean and Pacific shores of Panama. Many

species (or sibling species) of marine mollusces are

known to range on both sides of Panama, so the case

is quite likely. One point remains to be clarified, that

is to know whether the Caribbean and the Panamic

populations of P. olssoni sp. n. are truly conspecific,

or 1f they belong to two sibling species. Further

investigations of the soft parts of the animals

(chromatism, variability of proteins, etc.) would

provide the answer to this question.

Habitat. Unknown.

Remarks. P. olssoni sp. n. differs from other

Pachybathron species in several respects. The body

whorl is usually devoid of strong growth lines and is

smooth and glossy. Due to the lack of growth marks

the colour pattern remains very clear. Morphometric

analysis reveals a L/W ratio of about 65%, which is

very high for the genus and represents a very

globular profile. The brown blotch at the base of the

columella is a feature shared by only one other

species in the genus, P. cassidiforme, from which it

is clearly separable on other grounds. The

combination of the locality data, a distinct shell

profile, the columellar blotch, and the intricate axial

pattern on a smooth surface are sufficient characters

to enable the species to be described as new. P.

olssoni Sp.n. is named in honour of Axel A. Olsson, a

pioneer in the study of the marine molluscan faunas

of both the Caribbean and Pacific sides of Panama.

He began his fieldwork in Panama as early as 1917

and published the majority of his work during the

1950’s and 60°s.

BIOGEOGRAPHY

During the Pliocene period, the Panamanian area

constituted the southern part of the Limonian sub-

province of the Gatunian. At this period in geological

history, Panama and Northern Colombia formed a set

of archipelagos of large and small islands separated

by wide seaways. These allowed free exchanges of

marine faunas between Pacific and Caribbean zones.

At the end of the Zanclian and beginning of the

Piacenzian stages of the Pliocene (Petuch, 1988),

approximately 3 million years ago, the isthmus of

Panama was formed, initially at the level of the

Balboa Seaway, situated at the present level of Costa

Rica and West Panama (close to the type locality of

P. olssoni sp. n.). By the end of the Piacenzian era at

the end of the Pliocene, only the Atrato Seaway

remained open, situated at the level of the Darien

Peninsula (close to the area of distribution of P.

tayrona). The Isthmus of Panama was finally

completed at the Pliocene-Pleistocene boundary,

around 2 million years ago.

Due to the relatively recent (in geological terms)

completion of the land bridge, it is hardly surprising

that within many molluscan groups, similar phenae

occur on both the Caribbean and Pacific sides. Some

of these separated populations have been subject to

gradual genetic drift (or even sudden mutation)

which has led to reproductive incompatability and

specific separation. These newly evolved species,

though genotypically different from each other may

have retained similar phenotypic characteristics and

therefore appear almost identical to each other.

Alternatively there may have been little or no genetic

change and the separated populations could still have

retained their mutual reproductive potential, despite

being physically isolated from each other. Both of

these scenarios could apply to populations of sibling

P. olssoni and P. cypraeoides (if the both are

confirmed as currently living on the Pacific side).

WAKEFIELD, BOYER, MCCLEERY

Genus Pachybathron

NOVAPEX 3 (2-3): 65-81, 10 juillet 2002

TAXONOMIC DISCUSSION

As expected, Pachybathron shares the same features

of its radular apparatus with Persicula, namely

strongly arched, overlapping, multicuspid rachidian

plates, arranged as a long, thin cord (Figs.50-58). A

significant difference however seems to be the

relative lengths of the radula. According to

observations under light microscopy undertaken by

Coovert & Coovert (1995) the genera Persicula,

Gibberula and Canalispira have rachidian plate

counts of 80 to 209. The current study on the genus

Pachybathron, utilising SEM observation of all but

one of the currently known species, produced plate

counts from 293-425. This dramatic difference could

prove to be a reliable diagnostic feature at the generic

level.

It has been observed that the various species of

Pachybathron can be found in a wide variety of

environments and habitats. P. kieneriana seems to be

able to tolerate substrates of both sticky mud and

coarse coralline sand. This could be responsible for

its wide area of distribution along the mainland coast

of Venezuela, as well as some offshore islands (Los

Testigos and Tobago). Other species, eg. P.

cassidiforme, are restricted to offshore islands and

found in places where fluvial sediments are absent.

At least two species (P. cypraeoides and P. tayrona)

appear to be restricted to environments comprising of

coral sand and seagrass. The habitat of P. olssoni sp.

n. is presently unknown (all the available material

being dredged dead). As P. olssoni sp. n. 1s currently

understood to be found on Pacific as well as the

Caribbean sides of Panama, it demonstrates its ability

to survive in two somewhat different environmental

conditions, since the Pacific coast of Tropical West

America nearly always has a Pacific swell which

makes for turbid shallow water with a notable

absence of Thalassia beds. The absence of P. olssoni

sp. n. from Caribbean Southeast Panama (San Blas

Archipelago) seems to be well established by recent

enquiries. However, the general distribution of the

species remains to be clarified, both along the

Caribbean side (towards the coasts of Costa Rica and

Nicaragua) and along the Panamic side (towards

Costa Rican protected bays and the Gulf of Panama),

as all these areas have not been checked for

micromolluscs.

ACKNOWLEDGEMENTS

Thanks are due to Regis Delannoye (Martinique,

France) who provided illustrations of the live animal

of P. cassidiforme, and to Rodolfo and Julia Santiago

(Caracas, Venezuela) who provided logistic

assistance for research into live P. kieneriana.

We are also indebted to Adam Baldinger (Museum of

Comparative Zoology, Harvard), Kathie Way and

Joan Pickering (Natural History Museum, London),

and Virginie Heros (Museum national d'Histoire

naturelle, Paris) for access to type material, and to

Juan M. Diaz (Instituto de Investigaciones Marinas y

Costeras, Santa Marta, Colombia) for information on

the Colombian population of P. tayrona.

Special thanks to Emilio Rolän (Vigo, Spain) who

extracted all the radulae and provided SEM

photographs.

REFERENCES

Adams C.B. 1845. Specierum Novarum

Conchyliorum, in Jamaica Repertorum, Synopsis.

Proceedings of the Boston Society of Natural

History, 2 : 1-2.

Adams H. and Adams A. 1858.. The genera of

recent Mollusca 1(1-40) : 1-484. London.

Boyer F., 1998. Dragages à Puerto Francés.

Xenophora, 83:10-14.

Clench W.J. and Turner R.D. 1950. The Western

Atlantic Marine Molluses Described By C.B.

Adams. Occasional Papers On Molluses, 1(15) :

233-404. Cambridge, Massachussets.

Coomans H.E. 1972. The genus Pachybathron

(Gastropoda). Basteria, 36 (2-5) : 89-96.

Coomans H.E. 1973. Additional notes on the genus

Pachybathron (Gastropoda, Marginellidae).

Argamon, 4(1) : 11-14.

Coovert G.A. and Coovert H.K. 1995. Revision of

the Supraspecific Classification of

arginelliform Gastropods. The Nautilus, 109 (23),

43-110.

De Jong K.M. and Coomans H.E. 1988. Marine

Gastropods of Curacao, Aruba and Bonaire. Ed.

J. Brill, Leiden, 1-261.

Diaz J.M. and Velasquez L.E. 1987. À new species

of Pachybathron from the Caribbean coast of

Colombia. Archiv für Molluskenkunde 117 (4-6) :

217-221.

Gaskoin J.S. 1853. On the genus Pachybathron and

some New Species of Marginella. The Annals and

Magazine of Natural History, (2) 11 : 356-360.

Olsson A.A. and McGinty T.L. 1958. Recent marine

molluscs from the Caribbean coast of Panama

with the description of some new genera and

species. Bulletins of American Paleontology, 39

(177) : 1-58.

Paulmier G. 1997. Trois mollusques nouveaux du

plateau insulaire Martiniquais. Annales de la

Société des Sciences Naturelles de la Charente

Maritime, 8 (6) : 733-748.

Petit de la Saussaye, S. 1838. Description de trios

espèces nouvelles des genres Carocolle,

Pleurotome et Marginelle. Revue Zoologique,

1220:

Petuch E.J. 1958. Neogene History of Tropical

American Molluses. The Coastal Educational and

Research Foundation, Charlottesville, Virginia, 1-

DATE

Sowerby G.B. 1852, re-ed. À Conchological

Manual. 4". Edition. PI. XXVIII, No. 600.

Talavera F.G. and Princz D. 1984. Marginella

lasallei spec. nov. y revision de la familia

Marginellidae en el mar Venezuelano. Bolletino

Malacologico, 20 (9-12) : 273-282.

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KREIPL & ALF

Galeodea beui sp. nov.

NOVAPEX 3 (2-3): 83-85, 10 juillet 2002

A new species of Galeodea Link, 1807 (Mollusca: Gastropoda: Cassidae)

from the Philippine Islands

: Kurt Kreipl

Meeresmuseum Ohringen, Hühenweg 6, 74613 Ohringen, Germany

e-mail : meeresmuseum(@t-online.de

Axel AIf

University for Applied Sciences Weihenstephan, 91746 Triesdorf, Germany

e-mail : axel.alf@fh-weihenstephan.de

KEY WORDS. Cassidae, Galeodea, Philippine Islands, new species

ABSTRACT. A new species of Galeodea Link, 1807 is described from the Philippine Islands and

compared with the three other species of Galeodea occuring in the same area :

Galeodea

leucodoma Dall, 1907, G. echinophorella Habe, 1961 and G. nipponica Sakurai & Habe, 1961.

INTRODUCTION

In September 2001 the senior author got a Galeodea

from a British shell dealer who had obtained this

shell from the Philippine shell dealer Quirino Hora.

At first glance it looked like an aberrant specimen of

Galeodea nipponica, but it is clearly distinguishable

from this species.

After having discussed this species with Dr Alan

Beu, Lower Hutt, New Zealand we have decided to

describe it as a new species.

List of Recent Galeodea Link, 1807 and their

geographical distribution :

Galeodea echinophora (Linné, 1758); Mediterranean.

Galeodea echinophorella Habe, 1961; southern Japan to

Western Australia.

Galeodea ferrarioi (Bozzetti, 1989); Somalia.

Galeodea hoaraui Drivas & Jay, 1989; Reunion Island.

Galeodea keyteri (Kilburn, 1975); southern Mozambique

to Natal, South Africa, and Madagascar.

Galeodea leucodoma Dall, 1907; Japan to Western

Australia, and southern Madagascar.

Galeodea maccamleyi Ponder, 1983; Central Queensland,

Australia.

Galeodea nipponica Sakurai & Habe, 1961; Japan to

Western Australia.

Galeodea rugosa (Linné, 1771); British Isles to West

Africa and western part of Mediterranean.

Galeodea triganceae (Dell, 1953); eastern coast of New

Zealand.

SYSTEMATICS

Family CASSIDAE Swainson, 1832

Genus Galeodea Link, 1807

Type species: Buccinum echinophora Linnaeus,

1758, Mediterranean.

Galeodea beui, spec. nov.

Figs 1, 2

Type material. Holotype MNHN, Paris

81,1 mm; greatest width : 51,8 mm.

height :

Type locality. Off north-western Panglao, Philippine

Islands; by tangle nets in 450-550 m, live-taken.

Description. Shell medium-sized for the genus,

elongate-ovate, thin but solid; taller than wide (h/w =

155);

Protoconch consisting of 2 > whorls. Teleoconch of

five whorls sculptured by regularly spaced spiral

cords (about 55 on body whorl), bearing fine nodules

on the early whorls. Shoulder of body whorl slightly

angulate with 28 not very prominent but distinct

nodules; another three nodule-bearing cords at

midbody of body whorl. These cords always consist

of three crowded spiral striae. Suture not distinctly

incised; one stronger spiral cord just below the

suture.

Parietal shield smooth, very thin. Aperture wide,

semilunate; outer lip reflected backward, almost

smooth. Anterior canal short, rather straight. No

posterior nodule on outer lip present.

Basic colour yellowish to orangish white with very

thin dirty greenish-brown translucent periostracum.

Protoconch whitish; inside of outer lip and interior of

aperture porcelain white: parietal callus transparent.

Operculum typical for Galeodea: corneous, ovate

with a marginal nucleus; reddish-brown.

Discussion. Three other species of the genus

Galeodea Link, 1807 are known in the Philippines:

leucodoma (Figs 3, 4), echinophorella (Figs 5, 6) and

nipponica (Figs 7, 8). These species differ from

Galeodea beui, spec. nov. in the following features :

(see Table 1)

KREIPL & ALF Galeodea beui sp. nov. NOVAPEX 3 (2-3): 83-85, 10 juillet 2002

LE sehnophorele — echinophorella

up to 65 mm

chalky white with a | chalky white chalky white

distinct orange patch

Anterior canal straight curved backward curved backward rather straight

on the back

Posterior nodule on | not present present not present not present

the outer li

Inside of outer lip smooth bearing obscure smooth bearing distinct

denticles denticles

thin thick

G. beui, spec. nov.

81,1 mm

yellowish to

orangish-white

Adult size

Colour

Spiral rows of

nodules on the body

whorl

Number of nodules

on the shoulder

spiral cord

Table 1. Comparison of Galeodea species.

Etymology. This new species is named after Dr Alan Kreipl, K. 1997. Recent Cassidae. Verlag Christa

Beu, Institute of Geological and Nuclear Sciences, Hemmen, Wiesbaden, Germany: 1-151.

Lower Hutt, New Zealand. Springsteen, F. J. & F.M. Leobrera 1986. Shells of

the Philippines. Carfel Seashell Museum, Manila,

ACKNOWLEDGEMENTS Philippines: 1-377.

Wilson, B. 1993. Australian Marine Shells vol. 1.

We want to thank Dr Alan Beu, Lower Hutt, New Odyssey Publishing, Kallaroo, Western

Zealand for his constant help and advice, as well as Australia: 1-408.

Simon Aiken, Rossett, GB and Quirino Hora, Bohol,

Philippines for useful information. Remarks

SELECTED REFERENCES AIT photographs by Uschi Damaschke, Môckmühl,

Germany.

Habe, T. 1964. Shells of the Western Pacific in Color

vol. 2. Hoïkusha Publishing, Japan: 1-233.

Figures 1-8

1-2. Galeodea beui spec. nov., 81, 1 mm x 51,8 mm, Panglao, Bohol, Philippines. Dorsal and ventral view.

Holotype MNHN. 3-4. Galeodea leucodoma Dall, 1907, 61,8 mm x 40, 9 mm, Honshu, Japan. Dorsal and

ventral view, coll. K.Kreipl. 5-6. Galeodea echinophorella Habe, 1961, 36, 9 mm x 25, 5 mm, Panglao, Bohol,

Philippines. Dorsal and ventral view, coll. K.Kreipl. 7-8. Galeodea nipponica Sakurai & Habe, 1961,

102, 7 mm x 69, 3 mm, Rowley Shoals, NW-Australia. Dorsal and ventral view, coll. K.Kreipl.

KREIPL & ALF Galeodea beui sp. nov. NOVAPEX 3 (2-3): 83-85, 10 juillet 2002

F. BOYER

Five new marginellids

NOVAPEX 3 (2-3): 87-96, 10 juillet 2002

Description of five new marginellids from bathyal levels

of southern New Caledonia

Franck BOYER

110, chemin du Marais du Souci

F-93270 Sevran, France.

franck.boyer6(@wanadoo.fr

KEY WORDS : Cystiscidae, Marginellidae, Gibberula, Dentimargo, Protoginella, bathyal levels,

New Caledonia.

ABSTRACT : One species of Gibberula, three species of Dentimargo, and one species of

Protoginella are described as new from bathyal levels south from New Caledonia. Dentimargo

caledonicus (Cossignani, 2001) is redescribed and a new type locality is proposed. Some elements

are given about the apparent distribution of the six species.

RESUME : Une espèce de Gibberula, trois espèces de Dentimargo et une espèce de Protoginella

sont décrites comme nouvelles du bathyal au sud de la Nouvelle-Calédonie. Dentimargo

caledonicus (Cossignani, 2001) est redécrit, et une nouvelle localité type est proposée. Quelques

éléments relatifs à l'apparente distribution des six espèces sont présentés.

INTRODUCTION

In a precedent article (Boyer, 2001), the author

presented a first set of ten bathyal marginellids

collected south from New Caledonia and conserved

in the Malacology department of the Muséum

national d'Histoire naturelle. In view of this first

article, few stations, as parts of four benthic

campaigns, were checked.

A further work on the general marginellids

collections from New Caledonia in MNHN led to

have a better knowledge of this fauna, especially

about some well-marked trends, like a very high

diversity at the specific level (around 100 species of

marginellids are represented from bathyal levels and

apparently as many in infralittoral), numerous

complexes of sibling species, and pronounced

situations of endemism, even at the bathyal levels.

The diversity of marginellids from New Caledonia is

also important at the generic level, the genus Cysticus

(Cysticidae) being dominant in shallow water and the

genus Dentimargo (Marginellidae) being dominant at

the bathyal levels. This pattern is original compared

to that one represented in the surrounding southwest

pacific archipelagos (Vanuatu, Fiji and Tonga

collections in MNHN) where the bathyal

marginellids fauna is characterized by a wide

diversity into the group Serrata. This contrasted

situation will require to be interpreted on the ground

of further inquiries.

This article is conceived as a complement to our

previous paper. It devotes to the description of five

new species which are especially well represented at

the bathyal levels south from New Caledonia, or

which present close affinities with such ones.

Cossignani (2001) described as new six species of

marginellids said to come from Northern New

Caledonia. One of these species, named as Prunum

caledonicum, belongs in fact to an area ranging south

from New Caledonia, where it is rather common in

upper bathyal. The original description of this species

being somewhat summary, a redescription 1s given

hereunder together with full collecting references

from MNHN material, and a new type locality is

proposed.

The holotypes of these six new species were initially

deposited in a private museum from Cupra Maritima,

Italy. They were removed from this place by Dr

Cossignani in December 2001 and deposited in

MNAN.

Abbreviations

L = length; W = width; Iv = live collected specimen:

dd — dead collected specimen; ad — adult; juv =

juvenile; fr = fragment; stn = station; N.C. = New

Caledonia.

AMS = Australian Museum, Sydney; MMM =

Mostra Mondiale Malacologia, Cupra Maritima:

MNHN = Muséum national d' Histoire naturelle,

Paris; NMNZ = National Museum of New Zealand:

NSMT = National Science Museum, Tokyo.

SYSTEMATICS

Genus Gibberula Swainson, 1840

Type species : Gibberula zonata Swainson, 1840 (—

Volvaria orvza Lamarck, 1822), by monotypy.

F. BOYER

Five new marginellids

NOVAPEX 3 (2-3): 87-96, 10 juillet 2002

The genus Gibberula is not represented by many

species in the bathyal levels of N.C. The species

described as new hereunder is the largest sized

Gibberula found off Southern N.C.

Gibberula nebulosa Sp. nov.

Figs 1-3

Type material. Holotype (6.05 x 4.15 mm), MNHN

(Fig. 1) : SMIB 8, stn DW 197-199.

Paratypes : 11 ad (L = 5.95 to 7 mm) and 2 juv,

MNAHN (Figs 2-3), same stn.

Material examined. ” Vauban" 1978-1979 : stn 15,

22°49'S, 167°12'E, 390-395 m,1 Iv. — stn 16, 22°46'S,

167°12'E, 390-400 m, 3 dd. -— stn 37, 22°32'S,

166°26'E, 175-250 m, 1 dd. — stn 40, 22°30'S,

166°24'E, 250-350 m, 2 dd, 2 juv dd.

BIOCAL : stn DW 37, 23°00'S, 167°16'E, 350 m,

Idd. — stn 44, 22°47'S, 167°14'E, 440-450 m, 1 dd.

MUSORSTOM 4 : stn DW 210, 22°44'S, 167°09'E,

340-345 m, 1 dd. — stn DW 226, 22°47'S, 167°22'E,

390 m, 1 dd.

SMIB 8 stns DW 197-199, 22°52'S-22°53'S,

168°12'E —-168°13'E, 408-436 m, 1lv (holotype Fig.

1), 2 juv lv (paratypes), 11 dd (paratypes Figs 2-3).

BATHUS 2 : stn DW 729, 22°52'S, 167°12'E, 400 m,

1 1v, 1 juv Iv, 5 dd, 1 fr — stn DW 739, 22935'S,

166°27'E, 465-525 m, 4 dd.

Type locality. SMIB &, stns DW 197-199, 22°52'S-

22°53'S, 168°12'E-168°13'E, 408-436 m, off Ile des

Pins, southeast of Grand Récif Sud, N.C., bathyal.

Description. Shell ovate subcylindrical, moderately

slender; protoconch small, paucispiral, hyalinous

grey, very faintly produced and partially covered by

callous enamel of the last whorl of the spire, spire

mostly sunken; aperture as long as the entire shell,

narrowed, slightly widened towards the base, labrum

faintly sinuous, labial denticles faint or lacking,

faintly developed lirations within the inner lip, thin

but distinct labial margin, making a strong oblique

varix at the base of the dorsum; parietal border

faintly convex, 3 visible columellar plaits, the two

first ones being the strongest, one to several lirations

on the parietal wall, well-incised siphonal notch;

ground colour whitish, spire whorls brownish, body

whorl covered by a lattice pattern of crossing

diagonal flammules, making a nebulous appearance.

Distribution. Southern N.C. (22°30'S to 23°00'S;

166°24'E to 167°22'E). Bathyal : Iv in 395-436 m, dd

in 250-465 m.

Remarks. The morphology and the decoration of the

shells are very constant. Their size range is L = 5.4 to

7.1 mm, W =3.4 to 4.1 mm.

G. nebulosa sp. nov. must be considered as a sibling

species of the northern G. quemeneri (Cossignani,

2001), which shows stronger lirations but a fainter

labial margin and a whitish concave spire, whereas

G. nebulosa has a brownish faintly convex one. The

decoration of G. quemeneri presents two extreme

phases : a basic pattern of zic-zac axial yellowish

lines, which may coexist with several blade-like

spiral bands of the same colour. The axial lines may

be lacking, a pattern of spiral bands and flammules

may ornate the entire shell. However, this last pattern

does not show any trend of crossing or merging and

appears as structurally different from that one of G.

nebulosa.

G. nebulosa is not represented in the MNHN

collections from Northern N.C., and the two sibling

species appear as being non-sympatric at both ends

of N.C.

The scarce samplings made off Western N.C. do not

allow to verify if both species are succeeding,

overlapping or lacking in this area.

Etymology. Referring to the nebulous aspect of the

shell decoration.

Genus Dentimargo Cossmann, 1899

Type species : Marginella dentifera Lamarck, 1803,

by original designation.

Dentimargo caledonicus (Cossignani, 2001)

Figs 3-6

Prunum caledonicum - COSSIGNANI, 2001, p. 15.

Holotype 14.33 x 7.00 mm, MNHN. Type locality :

Grand Passage, Northern N.C.

Materiel examined. SUD POINTE GRAND RÉCIF,

1976 : 200 m, 1 dd.

"Vauban" 1978-79 : stn 15, 22°49'S, 167°12'E, 390-

395 m, 2 dd. — stn 16, 22°46'S, 167°12°E, 390-400 m,

1 dd.

BIOCAL : stn DW 38, 23°00'S, 167°15'E, 360 m, 1

dd. stn DW 44, 22°47'S, 167°14'E, 440-450 m, 1 Iv,

1 juv Iv, 9 dd, 2 juv dd.

MUSORSTOM 14 : stn DW 212, 22°47'S, 167°10'E,

375-380 m, 7 dd, 1 juv dd. — stn DW 222, 22°58'S,

167°33'E, 410-440 m, 1 Iv, 4 juv Iv, 1 dd. stn DW

229, 22951'S, 167°13'E, 445-460 m, 1 dd.

SMIB 1 : stn DW 2, 22°52'S, 167°13'E, 415 m, 2 dd.

SMIB 2 :stn DW 1, 22°53'S, 167°13'E, 438-444 m, 2

dd. — stn DW 3, 22°56'S, 167°15'E, 412-428 m, 2 juv

Iv, 2 dd. -stn DW 6, 22°56'S, 167°16'E, 442-460 m,

1 dd.— stn DW 14, 22°53'S, 167°13'E, 405-444 m, 1

Iv.

SMIB 3 :

dd.

SMIB 8, stns DW 197-199, 22°52'S-22°53'S,

168°12'E-168°13'E, 408-436 m, 3 lv (Figs 4-6), 1 juv

lv, 22 dd, 1 juv dd.- stn DW 200, 23°00'S, 168°2l'E,

514-525 m,1 dd.

stn DW 29, 22°47'S, 167°12'E, 405 m, 2

F. BOYER

Five new marginellids

NOVAPEX 3 (2-3): 87-96, 10 juillet 2002

BATHUS 2 : stn DW 719, 22°48'S, 167°16'E, 444-

445 m, 8 dd, 1 juv dd. — stn DW 729, 22°52'S,

167°12'E, 400 m, 5 dd. — stn DW 730, 23°03'S,

166°58°E, 397-400 m, 1 dd.

Description. Shell subcylindrical, slender;

protoconch medium sized, low, moderately wide,

paucispiral; spire moderate, regularly conical, non-

stepped, made of 3 whorls, representing around 30 %

of the total length; aperture widely opened towards

the base, labrum straight and vertical, the small upper

labial denticle is principally made by the depression

of the anal canal, inner labrum smooth or very faintly

denticulate, external margin moderately thickened,

bordered by a deep groove on the backside, shoulder

of the labrum stepped; parietal border oblique and

straight, the columellar zone being faintly concave

and bearing 4 columellar plaits, the first two ones

being oblique and subequal, the third one almost

horizontal, and the fourth nearly perpendicular to the

parietal border; a pronounced wide varix is generally

apparent towards the ventral base of the body whorl,

at the level of the columellar plaits; ground colour

horny to amber, with one orange spiral band at the

top of the body whorl, one orange spiral line a short

distance below, another spiral line behind the basal

callosity and a lower spiral line are present on the

spire whorls, all these bands and lines are

discontinuous, only the lines are extending on the

labial margin.

Distribution. Southern N.C. (22°46'S to 23°00'S,

167°10'E to 167°33'E). Bathyal : 1v in 436-440 m, dd

in 200-514 m.

Remarks. The morphology and the decoration of the

shells are very constant. Their size range is : L = 12.2

to 16.8 mm, W = 6.2 to 7.6 mm. The species is not

represented in the MNHN collections from Northern

N.C. and seems to be restricted to the Southern N.C.

The replacement of the species within the genus

Dentimargo is based on the presence of a sharp

denticle at the upper part of the inner labrum and of

small denticles below.

Dr Cossignani founded this species on two specimens

reported from a northern station. This record seems

to be erroneous, and the type locality is changed here

as "south of N.C., bathyal".

Dentimargo tropicensis sp. nov.

Figs 7-9

Type material

Holotype (13 x 5.7 mm), MNHN (Figs 7-8) : SMIB

8, stn DW 182-184.

Paratypes : 3 ad (L = 11 to 11.6 mm) and 10 juv,

MNAN (Fig. 9): same stn.

Material examined. SMIB 8 : stns DW 182-184,

23°18'S, 168°05'E, 305-367 m, 3 lv ( holotype Figs

7-8, 2 paratypes Fig. 9), 10 juv Iv (paratypes), 1 dd

(paratype). — stn DW 189, 23°18'S ,168°06'E, 400-

402 m, 2 1v, 2 juv Iv. — stn DW 190, 23°18'S,

168°05'E, 305-310 m, 5 juv lv, 1 juv dd.

BATHUS 3 : stn DW 830, 23°20'S, 168°0l'E, 361-

365 m, 1 dd.

Type locality. SMIB 8, stn DW 182-184, 23°18'S,

168°05'E, 305-367 m, Banc Aztèque, Northern

Norfolk ridge.

Description. Shell slender biconical, narrow;

protoconch domed, medium sized, paucispiral; spire

high and produced, made of 3 *% whorls faintly

convex and representing more than 35 % of the total

length; aperture very narrowed, labrum faintly

arched, the small upper labial denticle fastens an

enclosed anal canal, the produced inner labrum

enveloping the aperture, bearing 15 smaller denticles,

the lower one making like a button-like varix inside

the aperture, beside the siphonal canal ; external

margin moderately thickened, bordered by a deep

groove on the backside, shoulder of the labrum

stepped; parietal border oblique and straight, the

columellar zone being faintly concave and bearing 4

columellar plaits, the first two ones being oblique and

subequal, the third one almost horizontal, and the

fourth nearly perpendicular to the parietal border;

ground colour flesh beige, with one orange spiral

band at the top of the body whorl, one orange spiral

line a short distance below, another spiral line and a

basal band at the level of the columellar plaits, the

upper spiral band and the lower spiral line being also

present on the spire whorls, all these bands and lines

are generally discontinuous, the extensions of the

lines on the labial margin are present even when the

lines are lacking.

Distribution. Northern Norfolk ridge (23°03'S to

23°20'S, 166°58'E to 168°06'E). Bathyal : 1v in 310-

400 m, dd in 310-397 m.

Remarks. The morphology of the shell is somewhat

variable in its restricted range of distribution. The

specimens collected on Banc Aztèque (SMIB 8, stns

182-184 and stn 189) are more variable, as well for

their outline (from very narrow and slender fusiform

to squat biconical) than for their decoration (from

uniformly deep greenish yellow or golden amber to

milky white with dark orange marks) and other

features (as the relative thickness and denticulation of

the inner labrum). The size range of the shells is : L =

11 to 13 mm, W =5 to 5.7 mm.

D. tropicensis Sp. nov. shows close similarities with

D. caledonicus.

It differs from this last species principally on the

ground of its much-narrowed aperture, attenuated

base, narrow outline and length/width ratio.

F. BOYER

Five new marginellids

It must be underlined that both species are not

overlapping, but on the contrary their geographic

ranges are following according to a very steep

manner : between the specimen of D. caledonicus

from SMIB 8, stn DW 200 (23°00'S, 168°2l'E) and

the specimens of D. tropicensis from SMIB 8, stn

DW 189 (23°18'S, 168°06'E), which are in the

closest vicinity, we do not verify any tendency to

intergrading. So, it seems that we are in the case of

an "insular isolation", D. tropicensis having speciated

on the submarine relieves ("guyots") situated at the

north of the Norfolk ridge, from an ancestral

population belonging to the D. caledonicus lineage.

A very small shell of D. caledonicus, collected in

BATHUS 2, stn DW 730, is somewhat comparable to

D. tropicensis, but not with a so slender outline and a

so attenuated base. Its ground colour is horny and the

inner lip 1s smooth below the upper denticle.

D. tropicensis is also very close to D. alisae Boyer,

2001 and to D. virginiae Boyer 2001, these three

species living sympatrically in the stn 190 of SMIB

8. D. tropicensis differs from D. alisae principally by

its larger shell and protoconch, by its less produced

spire and by its discontinuous pattern of decoration

which better remembers that one of D. caledonicus.

These four species show like representants of a

descendant lineage.

Etymology. Referring to the range of distribution of

the species, approximatively situated under the tropic

of Capricorn.

Dentimargo cingulatus sp.nov.

Figs 10-12

Type material. Holotype (9.4 x 5 mm), MNHN

(Figs 10-11) : SMIB 8.

Paratypes : 12 ad (L = 9 to 12.2 mm) + 3 juv, MNHN

(Fig. 12) : same stn.

Material examined. BIOCAL : stn DW 44, 22°4T'S,

167°14'E, 440-450 m, 4 juv Iv, 8 dd, 4 juv dd.

MUSORSTOM 14 : stn DW 222, 22°58'S, 167°33'E,

410-440 m, 4 dd. — stn DW 229, 22°51'S, 167°13'E,

445-460 m, 1 1v, 2 juv lv. — stn DW 230, 22°52'S,

167°12'E, 390-420 m, 2 I1v, 3 juv lv.

SMIB 2 : stn DW 3, 22°56'S, 167°15'E, 412-428 m, 2

dd, 1 juv dd. — stn DW 6, 22°56'S, 167°16'E, 442-460

m, 1 dd. — stn DW 9, 22°54'S, 167°15'E, 475-500 m,

Figures 1-9

NOVAPEX 3 (2-3): 87-96, 10 juillet 2002

1 1v, 2 dd. — stn DW 17, 22°55'S, 167°15'E, 428-448

m, | 1v.

SMIB 3 : stn CP 4, 24°54'S, 168°22'E, 530 m, 2 lv.

SMIB 8 : stns DW 197-199, 22°52'S-22°53'S,

168°12'E-168°13'E, 408-436 m, 2 Iv (holotype Figs

10-11, 1 paratype Fig 12), 2 juv lv (paratypes), 11 dd

(paratypes), 1 juv dd (paratype). — stn DW 201,

22°59'S, 168°21'E, 500-504 m, 1 dd.

BATHUS 2 : stn DW 719, 22°48'S, 167°16'E, 444-

445 m, 4 dd.

Type locality. SMIB 8, stn DW 197-199, 22°52'S-

22°53'S, 168°12'E-168°13'E, 408-436 m, off Ile des

Pins, southeast of Grand Récif Sud, N.C., bathyal.

Description. Shell biconical somewhat oval,

moderately slender; protoconch small, narrow and

tall, paucispiral; spire regularly conical, non-stepped,

representing around 30% of the total length; aperture

moderately opened, labrum not arched, faintly

oblique, one small denticle at the upper part of inner

labrum, lower denticles very faint, external margin

thick, shoulder of the labrum faintly stepped; parietal

border moderately concave, 4 strong columellar

plaits, the first two ones being oblique and subequal,

the third almost horizontal, and the fourth nearly

perpendicular to the parietal border; ground colour

white, 2 well defined spiral orange lines on the body

whorl recovering the labial margin, a third orange

line above the lower suture, a fourth one suggested

above the next suture.

Dry animals are orange coloured.

Distribution. South from N.C. to Northern Norfolk

ridge (22°47'S to 24°54'S; 167°12'E to 168°22'E).

Bathyal : Iv in 420-530 m, dd in 428-500 m.

Remarks. The morphology of the shell is very

constant all along its wide range of distribution. The

size range is : L = 9 to 12.2 mm, W = 4.5 to 6 mm.

The spiral decoration may be very erased or lacking.

A group of similar forms, possibly polyspecific, is

represented in Northern N.C. It can be distinguished

from D. cingulatus sp. nov. by the presence of much

wider protoconchs and stronger stepped spires.

Dentimargo guionneti Cossignani, 2001 belongs to

this northern group.

Etymology. Referring to the spiral decoration of the

shell.

1. Gibberula nebulosa, holotype, 6.8 x 4.15 mm. 2. Gibberula nebulosa, paratype, 6.15 x 3.65 mm.

3. G. nebulosa, paratype, 7.1 x 4 mm. 4-5. Dentimargo caledonicus, 15 x 7 mm, SMIB 8, stn DW 197-199.

6. D. caledonicus 16 x 7.25 mm, SMIB 8, stn 197-199. 7-8. D. tropicensis, holotype, 13 x 5.7 mm.

9. D. tropicensis, paratype, 11.45 x 5.6 mm.

F. BOYER Five new marginellids NOVAPEX 3 (2-3): 87-96, 10 juillet 2002

F. BOYER

Five new marginellids

Dentimargo biocal sp. nov.

Figs 13-15

Type material. Holotype (6.4 x 3.8 mm), MNHN

(Figs 13-14) : BIOCAL, stn DW 44.

Paratypes : 72 ad (L = 6.2 to 8.8 mm) + 20 juv, 14 ad

+ 8 juv (alc), MNHN; 2 ad, AMS; 2 ad, NMNZ,; 1 ad,

NSMT; 1 ad, MMM. Same stn.

Material examined. "Vauban" 1978-79 : stn 2,

22°17'S, 167°14'E, 425-430 m, 1 Iv, 1 juv lv, 14 dd, 2

juv dd.- stn 3, 22°17'S, 167°12'E, 390 m, 4 dd. — stn

4, 22°17'S, 167°13'E, 400 m, 1 dd.

BIOCAL : stn DW 44, 22°47'S, 167°14'E, 440-450

m, 17 lv (holotype Figs 13-14, 16 paratypes), 13 juv

IV (paratypes), 79 dd (paratypes), 12 juv dd

(paratypes). — stn DW 77, 22°15'S, 167°15'E, 440 m,

L Iv, 1 juv Iv, 14 dd, 1 juv dd.

MUSORSTOM 4 : stn DW 212, 22°47'S, 167°10'E,

375-380 m, 1 dd. — stn DW 222, 22°58'S, 167°33'E,

410-440 m, 2 dd, 1! juv dd. — stn DW 230, 22° 52'S,

167°12'E, 390-420 m, 1 dd.

SMIB 8 stns DW 197-199, 22°52'S-22°53'S,

168°12'E-168°13'E, 408-436 m, 1 1v, 6 dd.

BATHUS 2 : stn DW 719, 22°48'S, 167°16'E, 444-

445 m, 19 dd, 1 juv dd. — stn DW 723, 22°50'S,

167°27'E, 430-433 m, 2 dd. — stn DW 724, 22°48'S,

167°26'E, 344-358 m, 1 dd. — stn DW 729 22°52',

167°12'E, 400 m, 1 dd.

As D. cf. biocal :

BATHUS 3 : stn DW 818, 23°44'S, 168°16'E, 394-

401 m, 1 juv Iv, 4 dd (Fig. 15).

Type locality. BIOCAL, stn DW 44, 22°47'S,

167°14'E, 440-450 m, Southern N.C., bathyal.

Description. Shell somewhat biconical-oval, squat;

protoconch wide and low, paucispiral, spire short,

rather obtuse and blunted, representing 22% of the

total length, 2 thin spire whorls, the upper one being

very short and somewhat indistinct; body whorl

rounded and inflated; aperture moderately opened,

labrum arched, faintly oblique; one very small

denticle at the upper part of inner labrum, lower

denticles faintly

distinct; external margin thick, shoulder of the

labrum non-stepped; columellar border somewhat

concave, bearing 4 plaits, the first two ones oblique

and subequal, the third one almost horizontal and the

fourth one nearly perpendicular to the parietal border:

ground colour light greenish-yellow.

Figures 10-18

NOVAPEX 3 (2-3): 87-96, 10 juillet 2002

Wet animals are creamy beige (BIOCAL, stn DW

44), dry animals are orange (SMIB 8, DW 197-199).

Distribution. South from N.C. to Northern Norfolk

ridge (22°15'S to 22°58'S, 167°10'E to 167°33'E).

Bathyal : Iv in 430-440 m, dd in 358- 444 m.

Remarks. Despite its variation in relative length of

spire , the shell morphology of D. biocal is very

constant along its range of distribution. The size

range 1s : L = 6.2 to 8.8 mm, W = 3.6 to 5 mm.

Depending on localities, the shells may show 2

orange marks on the backside of the margin, on a

creamy-white ground.

The single lot collected on the northern slopes of the

Norfolk ridge (BATHUS 3, stn DW 818) displays

shells with a similar morphology, but with a smaller

protoconch, a more produced spire, a shorter

aperture, a more receding labial shoulder and a

pronounced orange decoration (Fig. 15). The entire

shell is light orange, with 2 thin orange bands on the

body whorl, another one behind the lower suture, and

a dark orange zone covering all the base of the shell.

This lot, labelled as D. cf. biocal, may represent a

sibling species restricted to a part of Northern

Norfolk ridge.

D. biocal is not represented in MNHN collections

from Northern N.C.

Etymology. Referring to the campaign BIOCAL

which yielded the most important lot of specimens.

Genus Protoginella Laseron, 1957

Type species : Marginella lavigata Brazier, 1877, by

original designation.

A phena labelled as P. cf. caledonica in Boyer (2001)

can be confirmed as constituting a new species

widely distributed in Southern N.C. and Northern

Norfolk ridge. This species is described hereunder.

Protoginella maestratii sp. nov

Figs 16-20

Type material. Holotype (8.3 x 4.7 mm), MNHN

(Fig. 17) : MUSORSTOM 4, stn DW 212.

Paratypes : 6 ad (L = 7.4 to 8.3 mm), 7 ad (alc),

MNEN,; 2 ad, AMS; 2 ad, NMNZ; 1 ad, NSMT; 1

ad, MMM. Same stn.

10-11. Dentimargo cingulatus, holotype, 9.4 x 5 mm. 12. D. cingulatus, paratype, 9.35 x 5.05 mm. 13-14. D.

biocal, holotype, 6.4 x 3.8 mm. 15. D. cf biocal, 7.3 x 4.05 mm. 16. Protoginella cf. maestratii, 7.25 x 4.5 mm,

MUSORSTOM 4, stn DW 224. 17. P. maestratii, holotype, 8.3 x 4.7 mm. 18. P. maestratii, 7.6 x 4.9 mm,

BATHUS 2, stn DW 729.

F. BOYER Five new marginellids NOVAPEX 3 (2-3): 87-96, 10 juillet 2002

F. BOYER

Material examined. "Vauban" 1978-79 : stn 15,

22°49'S, 167°12'E, 390-395 m, 2 dd. — stn 16,

22°46'S, 167°12'E, 390-400 m, 1 Iv, 5 dd, 1 fr.

BIOCAL : stn DW 37, 23°00'S, 167°16'E, 350 m, 3

dd. — sin DW 38, 23°00'S, 167°15'E, 360 m, ljuv Iv,

1 dd. — stn DW 44, 22°47'S, 167°14'E, 440-450 m, 36

lv, 3 juv lv, 82 dd, 8 juv dd. — stn DW 46, 22°53'S,

167°17'E, 570-610 m, 2 Iv, 1 juv Iv, 142 dd, 46 juv

dd. — stn DW 49, 23°03'S, 167°32'E, 825-830 m, 1

dd. — stn DW 53, 23°09'S, 167°43'E, 975-1005 m, 2

dd. — stn DW 77, 22°15'S, 167°15'E, 440 m, 5 Iv, 2

dd.

MUSORSTOM 4 : stn DW 212, 22°47'S, 167°10'E,

375-380 m, 7 Iv (paratypes), 13 dd (holotype Fig. 17,

12 paratypes). — stn DW 219, 23°02'S, 167°33'E, 750

m, 1 dd. — stn DW 220, 22°58'S, 167°38'E, 505-

550m, 1 dd. — stn DW 221, 22°59'S, 167°37'E, 535-

560 m, 1 dd. — stn DW 222, 22°58'S, 167°33'E, 410-

440 m, 7 Iv, 4 dd. — stn DW 223, 22°57'S, 167°30'E,

545-560 m, 1 dd. — stn DW 224, 22°55'S, 167°27'E,

575-595 m, 3 dd (Fig. 16). — stn DW 230, 22°52'S,

167°12'E, 390-420 m, 1 dd.

SMIB 1: stn DW 2, 22°52'S, 167°13'E, 415 m, 2 Iv (1

used for radula extraction), 1 dd.

SMIB 2 : stn DW 3, 22°56'S, 167°15'E, 412-428 m, 1

dd.— stn DW 5, 22°56'S, 167°14'E, 398-410 m, 2 dd.

— stn DW 6, 22°56'S, 167°16'E, 442-460 m, 11v. — stn

DW 9, 22°54'S, 167°15'E, 475-500 m, 1 dd. — stn DW

16, 22°5SL'S;167°12'E, 390 m, 2dd-.

SMIB 3 : stn CP 4, 24°54'S, 168°22'E, 530 m, 1 Iv, 2

dd.

BERYX 11: stn DW 27, 23°37'S, 167°4]'E, 460-470

m, 2 Iv.

SMIB 8 : stn DW 166, 23°38'S, 167°43'E, 433-450

m, 6 dd. — stn DW 167, 23°38'S, 167°43'E, 430-452

m, 3 dd. — stn DW 169, 23°37'S, 167°42'E, 447-450

m, 3 Iv, 1 dd. — stns DW 182-184, 23°18'S-23°19'S,

168°05'E, 305-367 m, 1 juv lv, 5 dd. — stn DW 189,

23°18'S, 168°06'E, 400-402 m, 11 dd. — stn DW 190,

23°18'S, 168°05'E, 305-310 m, 1 dd. — stns DW 197-

199, 22°52'S-22°53'S, 168°12'E-168°13'E, 408-436

m, 38 dd.

BATHUS 2 : stn DW 719, 22°48'S, 167°16'E, 444-

445 m, 9 dd. — stn DW 720, 22°52'S, 167°16'E, 530-

541 m, 1 Iv, 36 dd, 1 juv dd. — stn DW 721, 22°54'S,

167°17'E, 525-547 m, 28 dd, 6 juv dd. — stn DW 723,

22°50'S, 167°27'E, 430-433 m, 3 Iv, 6 dd. — stn DW

7129,22°52/S,167°12'E, 400/m.11IV Fis: 18) 2 uv

lv, Idd.

BATHUS 3, stn DW 830, 23°20'S, 168°0l'E, 361-

365 m, 2dd.

Type locality. MUSORSTOM 4, stn DW 212,

22°4TS, 167°10'E, 375-380 m, west-southwest off

Ile des Pins, east of Grand Récif Sud, N.C., bathyal.

Description. Shell white, subpellucid, triangular,

eratoiform, squat, very widening at the level of the

shoulder and very attenuated towards the base; spire

short, narrow and pointed, non turriculated;

Five new marginellids

NOVAPEX 3 (2-3): 87-96, 10 juillet 2002

protoconch narrow and produced, paucispiral;

aperture, moderately widened, faintly but regularly

curved, inner labrum thickened, somewhat

enveloping, bearing around 25 very faint denticles,

upper labrum sticking out backwards and producing

above the anal canal, outer margin very thickened;

parietal callus making an axial crest continuous and

pronounced; 5 columellar plaits making faint

crenelations in the parietal callus, the first plait is

long and very oblique, the second and the third ones

are thick and faintly biplicate and incised at the level

of the parietal callus, without visible external varix,

the fourth and fifth are smaller and simple, but well

distinct.

Animal : based on 3 well-conserved dry specimens

(BATHUS 2, stn DW 723), foot yellowish with

violet marks, mantle light violet with small violet

stains on the border, siphon light violet with a darker

tip.

Radula : examined and pictured from one specimen

(SMIB 1, stn DW2, L = 6.8 mm). Type 5 modified

(Figs 19-20), plates bearing 20 to 23 cusps, large and

small alterning, with a large central cusp very

produced, frontline of the cusps almost rectilinear.

Distribution. South from N.C. to Northern Norfolk

ridge (22°46'S to 24°54'S, 167°12'E to 168°22'E).

Bathyal : Iv in 360-570 m, dd in 310-975 m.

Remarks. P. maestratii Sp. nov. presents a very

variable shell morphology, depending on the

populations considered : the spire may be very

narrow and produced in some populations, or squat

and stepped in others; the labrum may be arched,

highly produced and very thickened, without visible

denticles, or more straight, angular and widening at

the level of the shoulder and distinctly denticulated,

or low streamlined, deeply enveloping the aperture in

its medium part, with strong denticulations; the

columellar plaits may be very faintly produced or

sharply standing out, the fourth plait being always

limited to a simple liration, whereas the fifth one 1s

often strongly biplicated and sharply stepped. Very

callous shells are frequent (Fig. 18).

Even if rarely occurring within such or such

population, some sympatric specimens intergrading

these forms are however observed.

The size range of the shell is : L = 5.05 to 10 mm, W

= 3.3 to 6.05 mm. The size of the shells tends to be

larger near from N.C. mainland and to become

smaller southward (the 2 very large shells from the

deep station DW 53-BIOCAL constituting an

exception), reaching its minimal on Banc Stylaster,

situated west from the Norfolk ridge ( SMIB 8, stns

166, 167 & 169). On the eastern side of the Norfolk

ridge (Banc Aztèque : SMIB 8, stns DW 182-184,

189, 190), P. maestratii presents larger specimens.

P. maestratii differs from P. laseroni Boyer, 2001,

principally by its more produced and sharper spire,

its more elevated labial shoulder, its smaller and

F. BOYER

more numerous labial denticles, and the presence of 5

columellar plaits instead of 4.

P. maestratii differs from P. caledonica Boyer, 2001,

principally by its more heart-shaped outline and its

smaller and sharper protoconch, its less slender base,

its enveloping and more elevated labrum, and its

slightly more widened aperture. P. maestratii is

generally squatter and more callous.

Although P. maestratii and P. caledonica must be

considered as sibling species and despite the

important material at hand, very few shells can be

considered as intergrading. These few shells come

from BIOCAL, stn DW 49 and MUSORSTOM 4,

stns 220, 223 & 224 (Fig. 16).

The smallest specimens of P. maestratii have the

same size than the specimens of P. laseroni, and the

largest ones have the same size than many P.

caledonica, both being sympatric with P. maestratii

in very few stations :

P. maestratii was found with P. laseroni in SMIB 8,

stns DW 182-184 & 190;

P. maestratii was found with P. caledonica in

MUSORSTOM 4, stns DW 219, 220 & 221, each

time as single shells (the one from stn DW 220 is

considered as intergrading).

The scarce sympatry of P. maestratii With P. laseroni

is due to the fact that P. /aseroni is apparently

endemic from one seamount from the Norfolk ridge,

where P. maestratii might be at the upper and/or

southern limit of its range of distribution.

The scarce sympatry of P. maestratii with P.

caledonica is more surprising, as both species present

more or less the same ranges of geographic and

bathymetric distribution from Southern N.C. to the

northern slopes of the Norfolk ridge. This kind of

pattern is generally representative of a polymorphic

species ranging along a bathymetric and/or a

geographic cline. However, a better look on the

compared bathymetry of both phenae shows that 70%

of the stations where P. maestratii was sampled are

situated in 350-500 m, whereas 70% of the stations

where P. caledonica was sampled are situated in 500-

700 m. Despite these different bathymetric

preferences, the scarcity of sympatric occurrences

must have a complementary explanation. The

compared geographic distribution of both phenae

does not reveal any coherent pattern. The dominant

light and slender form of P. caledonica, With narrow

aperture and large bulbous protoconch, is found all

along the bathyal plains situated south from N.C., as

is found the dominant callous and heart-shaped form

of P. maestratii With more widened aperture and

small protoconch.

As the cases of close similarities between both

phenae are very limited and as the differences

between both phenae are not based on a bathymetric

or a geographic ground, the possibility of a cline of

forms linking two phenae belonging to a single

species cannot be accepted. The simplest explanation

is that P. maestratii and P. caledonica are sibling

Five new marginellids

NOVAPEX 3 (2-3): 87-96, 10 juillet 2002

species which have any kind of different requirement

which leads them to occupy different places.

Even if reaching also the north of the Norfolk ridge,

P. caledonica does not settle the upper levels of the

banks, as P. maestratii does on Banc Aztèque (310-

400 m) and on Banc Stylaster (450-460 m).

Like P. laseroni and P. caledonica, P. maestratii is

not represented in the MNHN collections from

Northern N.C., where the diversity of Protoginella

species seems to be less important.

The chromatism of the animal of P. maestratii seems

to be very close to that one of P. /aseroni.

The radula of P. maestratii (Figs 19-20) differs

strongly from that one of P. laseroni (BOYER 2001 :

168) as the later presents the classic corner-patterned

plate with reduced number of cusps found in the

group Mesoginella (sensu COOVERT & COOVERT,

1995), whereas the former shows a pattern of plates

much closer to that one of some species belonging to

the complex Volvarina-Serrata (comblike pattern).

So, the study of the radulae of all the species of

Protoginella might be of interest for the

understanding of the possible relationship between

the group Mesoginella and the complex Volvarina-

Serrata.

Etymology. The species is dedicated to Philippe

Maestratii (MNHN), who valuably contributed to the

building up of the New Caledonian collections of

molluses, and who sorted out most of the shells with

such a patience and such a discriminating eye.

19-20 : radula from Protoginella maestratii (x 835),

SMIB 1, stn DW 2.

F. BOYER

Five new marginellids

DISCUSSION

On the ground of the observations reported in the

present article and of the data from the previous one

(BOYER, 2001), we can distinguish the outlines of an

original pattern of distribution in the bathyal

marginellid gastropods from N.C.

Each of the 16 species presented by us from Southern

N.C. appears as lacking in Northern N.C., and it

seems that none of the phenae examined from the

northern stations is also found in the southern area,

except the large Volvarina armonica Cossignani,

1997 which is apparently distributed all around the

New Caledonia mainland at circalittoral and upper

bathyal levels. So, a steep separation seems to occur

at the specific level between the northern and the

southern bathyal fauna from N.C., at least for small

neogastropods having a direct (non-planktotrophic)

development such as marginellids. The choppy

relieves laying off the west and east coasts of N.C. at

bathyal levels are apparently the place for special

microenvironments favourable to the development of

local endemisms and working like barriers

preventing the diffusion of northern and southern

populations. This situation remains to be fully

documented and interpreted.

Superposed to this general pattern, a normal tendency

to endemism seems to occur on the seamounts

ranging upon the Northern Norfolk ridge. Depending

on the species, the distribution of the seamounts

marginellids seems however to be irregular and may

be the result of very complex influences. The

evolution history of this kind of endemism may be

the matter of another process than the one working

about the specific separation between northern and

southern new-caledonian faunas.

NOVAPEX 3 (2-3): 87-96, 10 juillet 2002

The development of the study of MNHN

marginellids collections may allow to interpret also

the respective influences in the formation of the

biodiversity observed at bathyal levels between New

Caledonia and surrounding archipelagos.

ACKNOWLEDGEMENTS

[ have to thank collectively the Malacology

department team of the Muséum national d'Histoire

naturelle (Paris) for its permanent help and support.

Pr Philippe Bouchet encouraged the production of

this article, and Philippe Maestratii performed the

iconography. The radula was extracted and pictured

by Dr Anders Waren from Rijksmuseum

(Stockholm).

Thanks are also due to Robert & Nicole Hasselot

(Jouques) who nicely typed the texts and to Roland

Houart (Landen) for his kindness as editor.

REFERENCES

Boyer, F. 2001. Espèces nouvelles de Marginellidae

du niveau bathyal de la Nouvelle-Calédonie.

Novapex 2 (4) : 157-169.

Coovert, G. A. & Coovert, H. K. 1995. Revision of

the Supraspecific Classification of Marginelliform

Gastropods. The Nautilus 2-3 : 43-110.

Cossignani, T. 1997. Descrizione di tre nuove

marginelle (Gastropoda : Prosabranchia,

Marginellidae e Cysticidae). Malacologia 24 : 16-17.

Cossignani, T. 2001. Descrizione di sei nuove

marginelle (Gastropoda : Prosabranchia,

Marginellidae e Cysticidae) della Nuova Caledonia.

Malacologia 35 : 12-17.

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NOTE AUX AUTEURS

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ou une Institution scientifique publique.

Les auteurs devront suivre strictement les règles du Code de Nomenclature Zoologique (quatrième édition).

Manuscrits. Les manuscrits seront rédigés en français ou en anglais. Ils doivent être dactylographiés, justifiés à gauche, avec double

interligne, sur une seule face de papier A4 et sur une colonne. Les marges doivent être de 25 mm minimum. La séquence des sections

respectera l'ordre suivant : titre, nom de(s) auteur(s), adresse(s) de(s) auteur(s), mots-clés et résumé en anglais (et éventuellement en

français). Les noms de genre et des (sous) espèces seront en caractères jfaliques. Les références dans le texte auront la forme: Keen &

Campbell (1964) ou (Keen & Campbell, 1964). Consultez un numéro récent de Novapex pour l'organisation du texte.

La liste des références, en ordre alphabétique, respectera la forme suivante (les titres des publications ne devraient pas être abrégés):

Keen, A.M. & Campbell, G.B. 1964. Ten new species of Typhinae (Gastropoda : Muricidae). TheVeliger 7(l): 46-57.

Powell, A. W.B. 1979. New Zealand Mollusca. Marine, land and freshwater shells. William Collins Publishers Ltd: xiv + 500 pp.

Mayr, E. 1989. Attaching names to objects. In: What the philosophy of biology is : essays for David Hull (M. Ruse, ed.),

Klumer Academic, Dordrecht: 235-243.

Illustrations. Les photographies doivent être de bonne qualité (couleur ou noir/blanc), imprimées sur papier brillant et montées sur un

support adéquat dans le format final souhaité (max. 16 X 21 cm). Des photographies en couleur peuvent être soumises pour une

reproduction en noir et blanc. Les illustrations peuvent également être fournies sur un support informatique (CD-ROM, ZIP) en format

BMP, JPG ou TIFF avec mention du programme utilisé. Elle doivent être montées et ne peuvent contenir aucun texte, sauf la