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NOVITATES ZOOLOGIOAE.
Vol. XIII., lyOG.
NOVITATES ZOOLOGICAE
H Jouvnal of Zooloi3\!
IN CONNECTION WITH THE TRING MUSEUM.
EDITKD BY
The Hon. WALTER UOTHSCHILD, Ph.D.,
1)k. ERNST IIARTRRT, .\nd Dr. K. JORDAN.
Vol. XTIT., 190G.
(WITH TKX PLATES.)
Issued at the Zoolo(ik'al Museum, Tring.
rniNTED BY HAZEI.L, WATSON & VINKY, Ld., LONDON AND AYLESBURY.
190G.
CONTENTS OF VOLUME XIII. (lOOO).
GENERAL SUBJECTS.
(See notes on nomenclature of species, geographical and other varieties, etc., pp. 42.3—4.3:2.)
AVES.
P.\GES
1. On the Birds of the Ishvnd of Trinidad. 0. E. Hellmayr .... 1—60
2. On the Birds of the Island of Babber. Ernst Hartert .... 288—302
3. On the Birds of Luang. Ernst Hartert 302—304
4. Critical Notes on the types of little-known species of Neotiopical Birds.
C. E. Hellmayr 305—352
5. Notes on a second collection of Birds from the District of Pari, Brazil.
C. E. Hellmayr 353—385
6. Miscellanea Ornithologica. Part III. Ernst Hartert .... 386 — 405
7. Additional Notes on Birds from N.W. Australia. Ernst Hartert.
(Plate I.) 754—755
8. Notes on Birds from the Philippine Islands. Part I. Ernst Hartert.
(Plate II.) 755—758
9. Erklarung. Anton Reiciienow .......... 762
llEPTILIA.
1. A new species of Giant Tortoi.se. Walter Rothschild .... 753—754
COLEOPTERA.
1. Some n^w Anthribid.ae from the collection of H. E. Andrewcs. Karl
Jordan 408-409
J. Two new Xenocerxs in the collection of R. von Bennigsen. Karl .Jordan 410
r
( vi )
LEPIDOrTKHA.
PAGES
1. New DrepanuVid'te, Tkyrid'ulae, Um/iliilue, and GeometriihiP, from liritisli
New Guinen. William Warren Gl 161
2. Two new A i/nrhikUe. Kaul Jordan. IGl
3. On a new paiasitie Tiiieid Moth from Quoensluid. Walter Kotiischild 102 — 100
4. New Sphinyidae. Walter Rothschild and Karl Jordan . . . 178 — 185
5. Two new Satwnidae. Walter Rothschild ...... 1S9 — 190
G. New Noctuidcie from British New Guinea. G. T. Bethune-Baker . 191 — 287
7. Some Sphingidae in the British Museum. Walter Rothschild and Karl
Jordan 400 -407
8. A Revision of the American Papilios. Walter RoTBSCiuLn ami Karl
Jordan. (Plates IV.— IX.) 411-752
9. Notes to Plates III. and X 759-761
SIPHONAPTEKA.
1. Notes on the Siphonnpte'i-a from the Argentine described by tlie late Prof<s.s(ir
Dr. Weycnbcrgh. Karl Jordan and N. Charles Rothschild . 170 — 177
2. Notes on Bat Fleas. N, 0. Rothschild 186-188
LIST OE PLATES IN VOLUME XIII.
Plate I. Aiitstralian Birds. By J. G. Keulemans.
II. Philipiiiiie BiiJs. By J. G. Keulemans.
III. Lepidoptera from New Guinea.
IV. ^
V.
VI.
VII.
VIII.
IX.
American Papilios.
X. Lepidoptera from New Guinea, Surinam, and Africa.
Vii
ERRATA.
p. 8, line 3 from above, read : vol. 38 instead of vol. 30.
p. 'J: the heading of species 14 ought to be; Cijaiicri)cs ci/niicn ryniica.
p. 33, species 99 : the authority of Gluucis Uirsula should be : (Gm.).
p. 58, species fi, read : dirdinnlh instead of Orrdueltx.
p. 3'20, line 17 from below, read: adult ^ (J instead of adult $ ?.
p. 351, line 33 from above, read : p. 3J2 instead of 'iii.
p. 351, last Une, read: p. 332 instead of 322.
p. 369, line 14 from below, read: "the main difference" instead of "the only difference.'
p. 387, line 16, 18, 22 from above, read : yeiiissieimis instead of yumseei/isis.
tut
NOVITATES ZOOLOGICAE.
Vol. XIII. FEBRUARY, 1900. No. 1.
ON THE BIRDS OF THE ISLAND OF TRINIDAD.
By 0. E. HELLMAYR.
ri"^HE iire.sL'iit jiaper is primarily liased ou the extensive collectiou.s made l)j'
-L Mr. Andre or his collectors in different parts of the island. Besides these
series, which amount to upwards of 1500 skins, the Tring Mnseum received a
number of birds collected bv Dr. Percv Rendall in the districts of Savannah
Grande and Tacarigua.
The greater part of Mr. Andre's collections was brought together on the
C'aparo and ( 'aroni Rivers, in the province Chaguauas ; bnt a good many specimens
have also been sent from Chaguararaas, on the north-western peninsula, and i'rom
Valencia, ])rovince of Arima. A few skins were obtained at Mount Aripo, at an
elevation (if from 150U to 2■^W feet. Among these are several species not obtained
elsewliere : such as, Titnagra cijcitiocephala suhrinerea ScL, Colihri (lelphinrie (Less.)
and Steatnn/is caripensis Humb. Thus, it would appear that the avifanna of the
mountainous district is to a certain extent different from that of the plains.
The best account of the birds of Trinidad has been published by Mr. Frank
Chapman in the Bulletin of the American Museum, vol. vi. (1894) pp. 1 — SO. This
elaborate paper contains a review of the existing literature, and the faunal position
of the island is so fully discussed that I need not go into the question at great
length. Mr. Chapman also called attention to the fact that most of the so-called
" Trinidad " skins are collected on the Venezuelan mainland. Many of these
skins in continental museums are labelled " Trinidad," " Orinoco delta," or even
" (luiaua," but their exact localities are as yet unknown. I suspect, however, that
they mostly come from that part of Venezuela south of the Orinoco delta adjoining
British Guiana, inhabited by the Warraw Indians. It appears that the Indian
linnters visit the plains as well as more elevated districts (perhaps the Serra de
Imataea), as typical (iniaiia forms, sui^h as Xi-nopipu atronite/is, and at the same
time characteristic high bind species, for instance ('nt/iarus tnelpomene /jirc//ul/i Heeh.,
occur in these trade collections. The most remarkable type, however, is JL/lon</mjj/M
macroceica Gonld. This has been attributed to Brazil by Salviu and Hartert,
but the make of the skins leaves not the slightest doubt as to their origin. The
many specimens examined are all of the nsnal '' Orinoco" m;ike, as is also the type
of C/tloropkanes purpurascens Scl. & Salv.
As well known, Trinidad belongs faunistically to Venezuela, and has no
relations whatever to the West Indies. Only one species, a swift, Chactura
cinereiiientiix laicrem-ci Kidgw., is peculiar to Grenada, Tobago and Trinidad, being-
represented in Guiana by the nearly allied C. c. yuianeims Hart. All other species
are of undoubted South American origin.
^ The greater part of the resident land birds appears to have limnd their way to
1
(-^ )
Trinidad throngh the Paria peuiiisulu. Quite si nnniliei- of forms characteristic of
the north coast of Venezuela extend tlieir range into the island, and several others
are strictly confined to Trinidad and tlie neighbouring district of Vcneznola, viz.
the state of Cumanu (and sometimes the northera portion of the Orinoco delta).
Sucii s])ecics are tlie following : Tnrdns phaeopijtjui pliaeopi/yokles, Uampliocelus
jacapa magnirostris, ['hocnicothmiipis rubra, P/iact/ioniis gmj gui/, Agijrtriu
ckionopectus cliionopectiis, etc.
Other siiecies show decided Guiana affinities, and are not known to occur
in the coast region of Venezuela. Hore belong Molothni.s iitroniti'ns, Sclateria
naevia, Pliaethoruis Ionguemareu$, Ceryle superciliosa miperciliosa, Trogon violaceus
riolaceus, llampliastos citellinus, Amazona amazonica, Coluniha rtifina rufna, Rallus
longirostris longirostris, etc. Thns the avifauna of Trinidad is composed of two
elements, Lnt the true Venezuelan forms are much in preponderance.
Besides, there are a number of species and subspecies peculiar to tiie island of
Trinidad. Mr. (ihapman {l.c. p. 7) enumerates twelve species, four of which were
since shown to occur also on the mainland (Phelps, Ati/t 1897, p. 302). The
careful comparison of onr Trinidad series with the material from Cnmana* and
the Orinoco region f in the Tring Museum brought to light several other forms,
which, though nearly allied to their mainland representatives, have sufficiently
marked characters as to warrant subspecific separation. The following is the list
of these forms with their mainland representatives :
Of these ten peculiar forms three : Dgsitkanuus affiitis undrci, Calospizu
mexieana vieilloti, and Celens elegans Icotaudi, are of undoubted Guianan origin ;
Pipile pipile finds its nearest ally in a bird inhabiting the environs of Paril,
while five others are of decided Venezuelan affinities, lliplranuam t. rujimentum
has close allies in the West Indies and in the (iuiauas, being therefore of no
importance for the present consideration.
it now remains to say a few words about the relations between Trinidad and
Tobago. The only recent account about the birds of Tobago is Comte de Dalmas'
excellent paper in the Mcmoires dc la Socicfc Zoologique de France, xiii. (19U0)
• This collection, nnmbering abont 1200 specimens, wa-s made by a Mr. Caracciolo, one of Mr. Andr6"s
collectors.
t This comprises the extensive collections made by Mr. Cherrie on the Orinoco, that of Messrs.
Klages and Andr6 on the t'aura River, and a sm:tll collection from tJuanoco in tlic Orinoco ilelta, received
from Mr. Andr6.
( 3 )
pp. l:S2-44, wlierein S5 species are recorded for the island. Mr. Andre sent
a collection of 4U0 specimens from Tobago to the Tring Museum containing
all the species mentioned by Dalmas, with the exception of two or three. Ten
subspecies are peculiar to the island of Tobago ; only four of these have repre-
sentatives on Trinidad, but of the others nearly allied races are found on the
Venezuelan mainland. Three (or perhaps fonr) species are common to Trinidad
and Tobago, while allied races take their place on the Paria coast.
The following lists give a better idea of these sjjecies and their mainland
representatives :
Peculiar to Tobago.
Represented on Trinidad by
Represented on the mainland by
Tiirthis .raiflhoxcefu^ Jard.
.1/imMs fi'ilvHA /ohagenais Dalra.
Tfof/lodi/tea 7nu>!mliiA tohagenals
Lawr
li llhipliihiiy flui'ipea hiatihiris Scl." .
7\inugi'a cpUciij)Un herlepsflt/ Dalm.
Chirtj.r/jihia ptireohi tilhiiiticaDnlm.
Si/)ialhixis terreslris ierrexlris Jard.
SitUyittnttis griaeiis fir'isfiAtK Javd.
Form/civot'a hiternteilia tohiuiPii^'is
Dalm
Sauceriittea fri/thrntiot't.^ vclhi
Bouc.
[Turdas leiicojix subsp. (Brit.
Guiana).]
Mimus gilviis melajiopleriix'La'WT.
T. m. rhn-im Berl. & Hart, j T. m. chiriis Berl. & Hart.
H. jUwipes ucuik-anda Lawr.
T, episcopus sclaliri Berl.
.S, li'nrslri.i rmri Chapm.
S. fr}/thrntiof(}ft eri/tliri'ttotoa
(Less.).
T. episcojyus sclateri Berl. (?).
Chh'oxiphia lyai'Pola jtarfuht
(Linn.) in Brit. Guiana, ete.
S. tm-enti-is HfrkitipeciKu (Jhapni,
tS". fjrlseu.'i phdp^i Chapm.
F. tiiUrmedia intermedia Cab.
S, evi/thruwiloii feliciue (Less.).
Besides the ten forms jieculiar to Tobago, there are six species which are
known to occur on the mainland, but have not been fonnd on Trinidad. These are
the following species :
Sj/orojiliihi aiitericariii (Gra.) (= Vmcatii auct.) Found also in ]>rit. (Juiana,
Cayenne, Surinam ; but not in Venezuela.
Eiietlieia hicolor omissii (Jard.). Fonnd also in the state of Cumana, etc.
DysiOiamniis meidalis meiitfdis (Temm.). Found alsn in the state of
Cnmansl, etc.
Cumpyloijterus ensipcnnis (8ws.). Known also from (Aimami.
Melaiterpeii ferricolor Berl. Known also from (Jumana, Orinoco, etc.
Ortalis riificatif/a (Jard.). Also fonnd on the Orinoco, but not yet recorded
from Cumana.
The occurrence on ToViago of these l)irds is difficult to explain. Even if we
admit that the two Finches and the Ortalis might have been introduced, any such
* llylophilus insularis Solater, P. Z. S. 18C1. p. 128 (Tobago).—//, pallidi/rons Dalmas, Af<hn. .S'w.
Zoiil. France ,xiii. VMt). p. Ilj5 (Tobago). I compared Sclator's type in the British Museum, and fmnul it
to be an immature bird of the large island race of aciitirmtdtf, named II. paUidi/rimn by Dalmas.
( 4 )
slippositimi would lie qtiito impossible with regaiil to the Humming-ljird and the
Woodpeokcr.
The fiist chapter of my pa])er eontains the acconut of the species collected bj-
Messrs. Andre, Perc_v Hendall and Chapman, the second a list of the sj)ecie8 found
on the islands between Trinidad jirojier and the Paria coast, and the third an
enumeration of those the oeenrrence of which is doubtful or has been wrongly given.
I have not included a number of Water-birds once or twice recorded for Trinidad,
as they arc carefully enumerated in C'hajuuau's paper referred to above.
I.-LIST OF THE BIRPS ACTUALLY FOUND ON TRINIDAD.
1. Turdus phaeopygus phaeopygoides Seeb.
[Turdua pharojii/f/us Cabanis ; in Schomburgk, Tfcwf Brit. Guuimi, iii. (1S4S) p. GGl! (Brit. Guiana).]
T. pl,aeoj>;/f/oule« Seebohm, Cat. Birds Brit. Mim. v. (1881) p. 404 (Tobago).
T. phoeopiijiia Leotaud, Ois. Trinidad, p. 197.
2 (J ad., 1 S jr., 1 ? jr., Valencia, March : 1 c? ad., Cajmro, April ; 1 t? ad.,
Aripo, 20IJU ft., May.
This series agrees with two topotypical Tobago-skins, in iiaviii'^- the back
idive-brown with a slight greenish tinge, which is never to be seen in tyjiieal
'/'. jt/KWOji'/t/i/.i from British Guiana, etc. The specimen from Aripo, although
nither browner than the others, is still decidedly more olive, less nal'ons brown
on the crown and nape than Gniana examples. One c? ad. from Cnmana and
one ? from Gnanoco in the Orinoco delta belong likewise to 7'. p. phacopi/goiilc.'i,
while the specimens collected by Mr. S. M. Klagos on the ('aura River represent
typical phaeopyyus. Notwithstanding some individual variation, tlie distinctness
of T. jj. pkaeojjygoides is fully confirmed by the present series. Its distribution
is as follows : Tobago : Mariah and Castare, May (Andre coll. in Mus. Tring) ;
Tiiniflad {tide supra); N.f'J. Venezuelu : Santa Ana Valley, near Cumanii,
March (Caracciolo coll. in Mus. Tring) ; Gnanoco, in the Orinoco delta, February
(Andr(5 coll. in Mus. Tring).
The adult specimens from the various localities have the ft)llowing measure-
ments : —
? (probably cJ), Mariali, Tobago . Wing,
¥ Castare, Tobago . . . „
J ( 'ajiaro, Trinidad . . . „
S N'alencia, „ . . . „
o ,, ,> . . . ,,
? (iuanuco, Orinoco-delta . . „
cJ Sta Ana, Cnmand . . . „
According to Ltotaud {Lc.) this species is
in .hum. J. Oriiitli. 19U2, pji. (i:5-5, CD.
2. Turdus fumigatus Licht.
Tardus fninKjulKS Lichtenstcin. I'ei:. Duld. (IHi:!) p. 38 (Brasilien).
T. casius (nee Bonaparte !) LiJotaud, Ois. Trinidad, p. 204.
8 adults ((? and ?) and 2 juv. from Caparo, March and April; 1 c? ad.
Valencia, March.
These specimens are above much lighter, not so dark rn-ily brown as
( ^ )
others I'rum I'aiVi (coll. hftwre), iiml on tlic Inwcr parts also cousidcnilily jalcr,
more ocbveous-brown, less rusty. A specimen from Duaca (near Tocuyo iu
N.W. Veneznela) is similar. Two examples from the Orinoco (Maipnres,
Munduapo) are everywhere darker, but not so intensely coloured as those from
Brazil. Cf. my remarks in Jonrii. f. Ornith. 1902, pp. 65-(i.
3. Turdus gymnophthalmus Cab.
Tiiirlns ijt/iiiiii'/JdIialiiiiis Cabauis : in Schuniburgk, R'ji«e lir'it. Guuiim, iii. (1848) p. fiS.i (ex
La Guaira, Caracas, and Cayenne).
T. iwdii/iiiis Lrataud, Ois. Triiiidml, p. 201.
•,' <? ad,, 3 ? ?, from Caparo, March, Aiiril ; 1 cJ from 8eelet, April; 2 ? ?
from C'aroni, May.
With a series of 'S'^ adult birds from Ciimana, Surinam, Britisii (Juiana,
Trinidad, Tobago, and tlie ( )rinoco region, I can see no difterence whatever Ijetween
the skins from the various localities. All, whether in fresh plumage or not, have
the base of the bill dark horn-colour or jilumbeous, the apical portion yellow
or greenish yellow. The naked space behind the eye is bright orange-yellow in
freshly killed specimens, but is also very well jaonouuced in old skins.
The nearest ally of 7'. (ji/mmphthalmus is T. maculirostrin Berl. it Tacz.,
of West Ecuador, as I have pointed out in Journ. f. Ornith. lOii'J, p. 53, where
also a short review of the allied forms is given.
1 have since had an opportunity of inspecting the series of " T. maculirostrix"
of the " Monograph of the Turdidae " in the British Museum.
As suggested in my former article, it is a mixtiim comiJOsitKin of three
different species. The following is the identification of the specimens listed by
Mr. U. B. Sharpe {I.e. i. p. 240) s.n. " T. niMulirostri.'s" :—
(a) Turrl>(s maculirostris Berl. & Tacz.
8pec. b, c. Balzar (lllingworth) ."i
„ ,/, r/. Pallatanga (Fraser). . - All these places are in West Ecmdor.
„ /. Santa Rita (Villagomez) ..'
The above specimens have the base of the bill dusky and the apical portion
yellow ; behind the eye there is a small naked spot.
(J)) T/irc/ifs iqiiohlU.-s debilis Hellm.
Spec. a. Rio Napo (Jameson) .\
,, (I, e. Sarayacu (Buckley) .j- East Ecuador.
„ h. Zamora (Fraser) .1
„ /■ — 0. from various localities in Peru.
These nine si)ecimcns, as well as those collected by Messrs. (Joodfellow &
Hamilton at Archidona and on the Rio Napo, East Ecuador (Mus. Tring) have
the bill entirely black and no trace of the naked spot behind the eye. Most of
the above e.xamples have a pure white patch on the fore-neck below the strijied
throat, but sometimes this character is obsolete. T. i debilis can, however, always
easily be distinguished from the typical form by its much shorter and weaker bill,
and the pure white throat with the dusky stripes much more distinct.
I have e.xamined three Bogota skins which, undoubtedly, belong to T. i. debilis.
They probably came from the eastern slopes of the Andes, while typical T. ////inbU's
seems to be confined to the mountains north of Bogota, and evidently m-curs only
on hi"h elevations.
{'■) Tu/ffus fimaurochalixu^ Cab.
Spec, ji, ij. Mapiri and Buganti (Buckle\}.
No. >i is yonng. Imt specimen // a fnlly adult bird. It Las a yellow bill,
and is nothing else than a very worn example of T. amawoclialinus. Count
Berlepsch has a large series of beantiliil skins from various places in Eastern
Bolivia whicli are in no way different from the Brazilian ones.
4. Troglodytes musculus clarus Berl. ik. Hart.
Triiriltifli/lcs iiiuxniliis r!nrn)< Berlepsch & II:utcit, Xur. Zo-//. ix. (1902) p. 8 (typ. ex Barlica Grove,
Brit. Ouiana).
T. nifuliis (nee Cabaiiis !) Chapman, BM. Amer. Mux, vi. (1894) p. 2,S (Trinidad).
1 S ad. from Caparo, April, agreeing with the pale form whicii inhabits
British Guiana, Cayenne, Surinam. Cumami, and the Orinoco region.
T. m. tobn(/ciii<i» Lawr.* from the island of Tobago, however, is quite distinct,
having the lower surface pure white, only the sides and under tail-coverts being
washed with isabelline-rufous. The differences have been well pointed out by
Comte de Dalmas.f It has also much longer wings and a rather longer tail.
In T. m. clurus the whole lower surface is pale isabelliue-rufous, only the
fore part of the throat and the middle of the abdomen inclining to creamy-whitish.
5. Thryothorus rutilus Vieill.
Thiiiolliunis nililiis Vieillot, Xoiu: Ui.i. xxxiv. (181'.i) p. :ib (" AmiTique septentrionale " — errore !
we substitute Trinidad as the typical habitat).
Sixteen specimens of both sexes from Chaguaramas (.January ), Caparo (April),
Lavantille (March), Valencia (March), Pointe Gourde (January), and Seelet (April).
They agree with a large series from Tobago, Cumana, and Ejido (near Jlerida),
in having the whole nndersurfaee strongly tinged with rufous except the middle
line of breast and belly, which is white. In some specimens, however, this white
8trij)e is almost wanting, and they come nearer to T. i-Ktilus hijijenjthras from
Costa Ilica and Panama, but the latter form is still recognisable by the decidedly
lighter, more orange-rufous tint of the lower parts.
7". riitihi.-i fii/pospoilins Salv. k Godm., from Bogota, has .only the chest
bright rufous, the rest of the lower jiarts being dirty gre\ ish with a Ijrownish
wash on tlie flanks.
C. Compsothlypis pitiayuini ( Vieill.).
Sijhiii intiiniiiiiii Vieillot, Nuno. Dirt. xi. (1817) p. 276 (Paraguay — ex A/.ara, No. 109).
Comiisiilhli/jiix jiilitiijiniii Chapman, Bull. Amu: Mas. vi. (1894) p. 24 (Trinidad aud Monos Ibluud).
i S 6 from Chagnaramas, January ; h <S <? I'rom Pointe Gourde, January ; aud
1 (? ad. from Caparo, Ai)ril.
These specimens are generally richer coloured than typical Paraguay skins.
~. Dendroica aestiva (Gm.).
iloidciUa 'ifMii-a Gmulin, SiiM. Xal. 1. ii. (1788) p. 996 (Cayenne and Canada : the latter locality
accepted as the typical habitat).
I c? ad. and 1 ? from Laventille, 1',). iii., i). xi. ; 1 ? (Jhagnaramas, ]. i.
* Tn'ijhilifUg iohatjfiigis Lawrence, Auk v. (ISss) p. iiij (Tobago),
t Mem. Soe. Zool. France, xiii. (1900) p. 133.
( ' )
^. Seiurus noveboracensis (Gin.).
iloluciUa iiovchovacemis Gmelin, Syst. Nut. i. ii. (1788) p. [158 (Louisiana).
1 c?, Caparo, 19. iv. ; 1 c?, Ohaguavamas, 6. \. ; I i , Seelet, 14. iv.
!». Geothlypis aequinoctialis (Gm.).
MoktciUa aeiiiihitntiulis Gmelin, Sjxt. Nul. i, ii. (1788) p. 97'2 (ex Dauljcntou, /'/. enl. 1)85, fig. 1 —
Cayenne).
Althoiii;h Taylor atul Chapman met with this species, and Leotand says it
is common, Mr. Audre did not send it in his numerons collections.
111. Setophaga ruticilla (Linn.)
MiilwiUu. Riilirilhi Linnaeus, f<ijsf. Nat. x. (17.58) p. 186 (Virginia: e.'c Catesljy).
2 c?cJ, Pointe Gourde, 13, l.si. ; 1 c? ad.,',' ¥ ?, Laventilie, 0 xi., :,'m xii. ;
one pair, (.'haguaramas, January.
Winter visitor.
11. Basileuterus auricapillus olivascens riiapni.
[Setaphaijii iiiniiapiUu Swainson, Aiiim. Mciiaij. (1838) p. 2'X3 ("Mexico & Brazil."— We accept
Brazil as typical locality.*).]
Basileuterus vertnirorns nlivasmis Chapman, Auk x. (1803) p. 343 (Trinidad).
3 c?c? and 1 ?, Caparo, March and April; 1 adult (not sexed), Lavcntille,
March.
This subspecies dififers from typical B. ai(ricapUli(s (Sw.) ex Rio, S. Paulo
and Paraguay by its slightly more greyish, less greenish back, and in having
the outer webs of the tail-feathers and quills olive-grey instead of olive-green.
Specimens from Cumana (a large series in the Tring Museum) are practically
identical with those from the island. .
12. Coereba luteola (Cab.)
Ccrthiola lulcolit Cabanis, J/«s. IleiiKun, I. (1850) p. Ulj ("Puerto Cabello?")
Sixteen specimens of both sexes (adult and young) from t'aparo, iii. iv. ;
] c? ad. Valencia, iii. ; one pair from Seelet, iv. They agree perfectly with a
large series from Cumana, S. Esteban and the Orinoco River. One S ad. from
Englishman Bay, Tobags, is a shade blacker on the back, but otherwise not
different. All have the whole upperside dull blackisli, deeper on the head
(only the immature birds with a slight olivaceous tinge) and a large white
speculum at the base of the primaries.
C. (juianensis Cab., ijuite erroneously synonymised with ('. chloropi/ifi, by
Mr. Sclater, differs from ( '. luteola by its much paler, smoky grey upper surface,
and by the want of the white speculum, which is but barely indicated under
the primary coverts by some traces of whitisli ; the rump-band is as brighl
yellow as in C. luteola.
C. cMofopiiya Cab., which ranges from Rio to Para, is again much i)aler,
light olivaceous grey above, the crown decidedly duller and less intense than in
C. guianensis, and the rump-band much paler, dull olive-yellowish (instead ot
bright chrome-yellow); the wing-speculum is, likewise, nearly wanting. In fact,
the three forms are very distinct one from another.
* Swaiiison's description evidently refers to the Brazilian :>pecies, for he sa,\^> "above, oUve-greeu."
( « )
lo. Cyauerpes caerulea trinitatis (Bii.)
[Certhia aienihn Linnaeus, %</. Sm. Id. i. (17.')8) p. 11« (ex Edwards— Surinam).]
Coereha Irimhills Bonaparte, Cnm/.t. lb ml. Ar. Sc. vol. 3'J. (1854) p. 258 (" e.\ Insula Saucla-
Trinit;ui.'' — Mus. Verreau.'i)
ArhelwhiiM aieniha Chapman, Bull. Amer. Mux. vi. (18114) p. 25 (Trinidad).
8 t? ad., C <S jr. or ? ? fioiii Vulcncia, March. Besides, we buvc two
adnlt males, collected by Dr. Percy Reiuliill in the Savannah Grande district, in
Febrnary INOT.
The series dillers Irom ail the ulher subspecies in liavnig a much loiij;er,
and at the base considerably broader bill. In the males the anterior part of
the crown, too, is decidedly darker blue, there beini^ no trace of the distinct
liglit blue patch to be seen in the other forms. The females from Trinidad have
the upper parts of a darker, duller green, and the uchreous colour of the throat
distinctly darker.
Birds of the so-called " Orinoco "-make agree jierfectly with those from
Trinidad, and raise once more the question whether some of these skins may not
really come from some part of that island. This sii[iposilion is strengthened by the
fact that specimens collected by Mr. Andre at Guauoco in the Orinoco-delta do
not belong to the hmg-billed island race, but agree in every way with typical
C. merulca from Surinam and Cayenne.
The Trinidad form is apparently entitled to the name C. trinituti.'i of
Bonaparte, which is entirely omitted from the Cat. of Binis, vol. .\i. Ferhajis
A. longiio/s/iis ("ab.* is an earlier name for the same form, but then the locality
must be wrong, since specimens from S. Esteban and Cumana both represent
the short-billed typical <'. caernlca.
With a series of 8U specimens before me, I can easily recognise the following
subspecies south of the isthmus of Panama :
{a) C)/anerpes caerulea caerulea (Linn.)
Typical locality : Siirin/i))/ (ex Edwards).
6 ad. Bill slender at the base, 18— 2(1^ mm. long, .\nterior part of crown
slightly tinged with clear azure bluish.
I cannot find any constant difference between tyjncal Surinam skins and
others from Puerto tJabello upon which the name Arhclorliiiia brerirostris (.'ab. has
been bestowed. The latter have not shorter bills, as claimed by Professor Caliauis :
on the contrary, one of my specimens from Puerto C'abello has the bill slightly
longer than those from Cayenne and Surinam. A series from Cumana also agrees
with those from the two latter localities.
The following measurements of some adult males out of my series may be
useful to students of this dilHcnlt group:
•Z from (Juanoeo, Orinoco-dell a: wing .3(j —57 ; tail 28 —20; bill 10 — 2U mm.
„ 29 —31: „ 10 -211 „
„ 20 -27; „ 10 -105 „
• Mus. llcinean. i. (ISTiU) y. '.IC— Caracas.
( 9)
As will be seen from (1k' above, tlie speciineiis tVimi Parii, on the average,
have slightly shorter bills, but some of them are not ditfereiit in this respect from
Cayenne skins.
Distribution : Surinam, (Jayenne, Brit. Guiaiia, westwards through the Orinoco-
delta along the north coast of Venezula as far as Puerto Tubello, southwards
extending to Para on the mouth of the Amazons.
(6) Ci/aHcr/M'g caerulea microrhijncka (Berl.)
Coereba cui-nilen mierorhyncha Berlepscli, Jmnn. J', (hii/th. 1.SS4, \k '^^~
(Bucaramanga).
d ad. This form differs from the tyjiical one by having the anterior ])arf
of the crown as well as the cheeks of a much clearer and paler azure blue ; the
rest of the pileum is also brighter blue. Bill slender and short, sometimes even
shorter than in ' '. c. crtrruliM.
20 Bogota skins . . Wing .56— 58 ; tail 28 —31 ; bill 18— lOAmm.
1 from Jlerida, W. Venezuela „ 57 ; „ 28i ; ,, 10 ,,
2 „ Beni, E. Bolivia . „ 58i : „ 3U ; „ 17i
Distribution : C!olonibia (Bucaramanga, Bogota coll.) ; Merida in X.W.
Venezuela; thence through the Upper Amazonian region of East Ecuador and
E. Peru to N.E. Bolivia.
(c) Cyanerpes caerulea cherriei Berl. & Hart.
JS'or. Zool. ix. 09112) p. 10 (Munduapo, Upper Orinoco).
i ad. exactly like that of No. 2, but consideriilily smaller, with the bill si ill
shorter and more slender.
Two males (types) measure : Wing 52— 5o ; tail 2(J — 2(U ; bill 10 — 17 mm.
Distribution, known only from the type locality : Munduapo, Upper Orinoco.
{il) CijaiK'rpes cacndi:u triiiifotin (Bonap.)
5 ad. Differs from No. 1 in its ranch longer and stronger bill, and in having
no trace of the light azure blue tinge on the front of the head, which is dark
bine like the occiput.
Fifteen S ad. measure : wing, 58-50 ; tail about 3o ; bill, 2;H— 25 mm.
Distribution : A])pareutly confined to Trhi'ukul. Also found in the so-called
"Orinoco-delta" collections, but their exact locality is, of course, not known.
Mr. Andre, however, collected in the Orinoco delta another form (cfr. sub a).
14. Cyanerpes cyanea (Linn.)
Certliia cyaiicu Liuuaous, Si/st. Nat. xii. i. (17G6) p. 188 (e.x Edwards, Urisson, etc. — Vfv Hx
Surinam (ex Edwards, whom Linnaeus quotes first) as the typical locality).
6 c? ad., 1 (? jr. and 1 ? from Valencia, iii. ; 1 (^ ad. Seelet, iv. ; 7 r( ad.
audi ?,raiiaro, iii. and iv. ; 1 ?,Aripo, l8t)ii feet elevation, v.: Id ad. from
Savannah Grande, collected by Dr. Percy Kendall.
In the length of the bill this series agrees best with specimens from Cayenne,
and Guanoco in the Orinoco-ilelta. Two c? c? from Toliago are exactly alike.
The birds from the interior of British (luiana apjicar to be referable to the same
form, although some specimens have very short bills. A series from Cumana
and San Esteban, as well as a specimen from Ejido, however, certainly represent
a different form, having the bill very much longer. In the so-called " Orinoca-
delta" collections, both forms occur, hence it seems that these skins come from
different places.
The long-billed form from N. Venezuela is evidently entitled to the name
( 1" )
r. c. erimia (I'ali.), described from Puerto Cabello. IMr. Oberholser* adopted tor
it the name C c hrcripes (Cab.), based on a sjieciiueu from the same jilace.
However, I am not qaite sure whether the locality is reliable, and as the description
of Arbelorkina eximia snits much better the long-billed race, I ])refer the latter
designation.
It is very interesting to note that the specimens from C'umana belong to
the long-billed form, while those from the Orinoco-<U'ltii, Trinidad and Tobago,
agree with the typical bird from < 'avenue. Bogota skins belong also to the
trne C. cyanea (Linn.),
The following measurements may heli> to distinguish the two races :
(a) Cijuncvpes ri/aiica cjarira (Linn.).
3 (J ad. Cayenne (Gherrie coll.) . "Wing, 01— 04; tail, 33— 37; bill, i:,ls,lNmni.
6 „ Mines district, Britisli
Guiana . . . ,, 63 — (io^; ,, 36 — 37; ,, lo — 17J „
2 ,, (Juanoco, Orinoco-delta ., 6oi, 67 ; ,, 36,38; „ l^A, 1'-' „
15 „ Trinidad . . . „ 64— 6S ; „ 354—40;,, 17^—10 „
2 „ Tobago . . . . ,,67,68; „ 38,40; „ 19 „
5 „ "Orinoco-delta"' skins . „ 63A — 65; „ 37 — 39; „ 161 — 18 „
a „ Bogota coll. . . . „ 64—66; „ 36-41; „ 161-18 „
1 „ Bahia . . . . „ 65; „ 39; „ 171 „
(J)) Ci/ancrjii'x cijaKfn cjchuiu (Cab.).
Arbrlorliiiia c.riiiiiii Cabanis, Mii». Ueiiicon. i. (IsoU) p. '.i6 (" I'orto Cabello").
4 i ad. y. Esteban, near Puerto
Hah. : North coast of Venezuela from Cumaua to Puerto Cabello, thence to
Ejido, near Me'rida.
lo. Chlorophanes spiza spiza (Linn.).
ilvlocHla spi-M Liunaeus, Si/nt. Sat. x. (17.')8) p. 188 (ex Edwards— Surinam) [exol. var /i].
4 c? ad., 1 ? from Caparo, iv. : 8 c^ ad. and 2 ? ? from Valencia, iii.
They belong to the tyi)ical form, agreeing with a large series from Cumauii,
S. Esteban, Gnanoco in the Orinoco-delta, British Guiana, Canra Kiver and
Cayenne. <\ -s. ijiiatfmalciisis !Scl. from Central America is larger, with a longer
bill, and has the j)bimagc of a lighter, more greenish tinge.
16. Dacnis cayana cayana (L.j.
Mniocilln cayniw Liuuacus, Sijsl. .\,il. xii. 1. (ITiKi; p. ri3G (Cayenue.— ex Bris.son f).
2 S S from Laventille, iv., v. ; 1 J from Seelet, \v.; I S from Chaguaramas,
i. ; 1 J jnv. Valencia, iii. ; 1 S ad. and 2 ? ? from Caparo, iv.
The adult c? c? agree in colour and dimensions with others from Cayenne,
British Guiana, Para, t!umana and the Orinoco region.
• Aiili xvi. (1899) p. 3.3.
t " Elotototl " of Hernandez, also cited by Linnaevis :is a .-.vnonym o£ his MolacilU eai/aiia, does not
belong to this specicy.
( 11 )
IT. Dacuis bicolor (Vieill.)-
Sylvia hh-i>hii- Vieillot, 0/». Amir, xcjil. ii. (1807) p. 32. tab. 90 bis. [•' tn>s rarement sous la zone
bore'ale, et plus communemciit eiitre les tropiques."— We substitute Onjenne as the typical
locality].
DacnU phnnhen- auct.*
3 6 ad., Seelet, Ai)ril, :uk1 1 S :ul. frum the Caroni Hwamp, I\Iarcli.
These skins agree with one S from L'umaiul, except in being rather more brightly
coloured above and in having the throat slightly mixed with greyish (instead of
pure buft'). Specimens from Cayenne are partly like those from Trinidad, and partly
like the one from Cumaua, tlius proving that the said differences are of no
significance.
is. Cyclarhis flavipectus flavipectus Scl.
diclorhis jlariperiiis Sclater, /'. Z. S. 1H.')8. p. 44H (•' Trinidad. Veoezuel^i. aud Colombia lltt.").
df. UhMalis AUen, Bull. Amn: ilus. ii. (1880) p. 131 (Trinidad).
C.f. runiicns 0. Bangs, Pi;u\ Biul. Soc. )l'»»/i. xii. (1898) p. 192 (Santa Marta).
A large series from C'ai)aro, March and April ; Laveutille, December and
March ; Chagnaramas, January ; Puinte Ciourde, January ; Valencia, March :
Seelet, April. These skins are practically identical with an extensive series from
the mainland of Venezuela (12 Cumauii, in Orinoco, 4 from N.W. Venezuela:
Bucarito, San Esteban and S. Carlos). ^Vitll two dozen Bogota skins before me,
I can no longer maintain C. /. ca/ific^/s as distinct. Many of the Colombian
specimens have the yellow colour on the underparts unite as pale as a number from
Trinidad, though some are certainly richer coloured and of a more golden yellow
beneath, as claimed by Mr. Bangs. All of the many specimens examined have the
lilumbeous sjiot at the base of the lower mandible very strongly pronounced.
l'». Vireo chivi (Vieill.) subsp. ?
Si/li-iri rhiri Vieillot, Xuut: Dirt. xi. (1817) p. 174 (ex Azara.— Paraguay I.
One <J and 3 ¥ ? from Caparo, March and April ; 1 ? Valencia, March. These
four skins, as well as a good series from Tobago, differ from a large number of
true l'. c/iici t in their larger size, especially longer wings and considerably longer,
heavier bill. The ujiper mandible is always blackish, while in ty[iical T. ckiei it is
much jialer, more brownish horn-colour. Specimens trom (hiinaiul (a large series in
Trlug) approach the Trinidad form in size, but the colour of the bill is exactly like
that of frne e/u'ri. Very likely the birds inhabiting Trinidad and Tobago constitute
a different race — which, however, cannot be called V. c. agiUf, as has been done by
Chiipman and other American writers. Lanim affilis Licht. is based on Bahia
skins which agree in every way with topotypical examjiles from Paraguay. If
separable, the northern form must have a new name.
• As alreatlj pointed out by Berlepscb (Jbix, 1881, p. 212), Sijlcia jjliimbrii I.alb. cannot apply to tl,c
Daciu-s jilnmbea auct., since in this species the upper parts are certainly not "deep lead-colour, nearly
blacli," nor are the lower parts " pale ash-colour."
t Fifty specimens from I'araguay, Rio, Baliia, Ceara, Orinoco, Caura, Soutli Fern. British Guiana and
Western Ecuador.
( 12 )
-". Pachysylvia aiirantiifrons saturata ikhu. uov.*
Ilylojihiltis i/milarif (nee Sclater 1) Li'otaud, Oi.s. Trinidad, p. 186.
//. amimtiifrons (nee Lawrence !) Chapman, Bull. Amei: Muh. vi. (1H'J4;' p. 21 (Trinidad).
Piii-hy^ijhiu aiimutiifi'iii.i hyjioj-aiithu (nee Pelzeln !) Ridgway, BinU X. and M. America, iii. (11104)
p. 215 (Venezuela, Trinidad).
Eight £?(? and ? ? from Oaparo, April arid May. Tlicy agree with a series of
14 sjieciiuens from the vicinity (if Ciimana, the dift'crences mentioned by Jlr. Kidgway
being ajuiarentiy not constant.
Professor Allen, as well as Count Berlepsch and Mr. Hartert, have conclusively
shown that P. auranfiifrons and P. wutkuuihi are spei-ifieally distinct. 1 have
now before me a series of both from near Cumana.
P. (iiircHtnfroHs is at once distinguishable by Laving the foreliead and lores
light yellow, the ear-coverts and sides of the neck pale rufous isabelline, and the
lower surface clear yellow. In P. f. acaticauda, on the other hand, forehead, lores,
sides of the Iicad and lower j)arts are uniformly liufly brownish. (Jenerally, there is
no trace of yellow on tlie under surface, but some specimens from tiie Orinoco
(Maipures, etc.) have the flanks slightly washed with yellowish, thereby apjiroaching
P. f.Jlaripoi (Lafr.) from Bogot.i.
As pointed out by Mr. Kidgway, the yellow-bellied form, inliabiting N.E.
Venezuela and. Trinidad, differs from tyjiical /'. luifdiiHifiom aKiantii/rons (Lawr.)
in having the cliest distinctly washed with ochraceous or butfy. It is, however,
ipiite distinct from /'. li>/poxantha (Pek.), with which it has been identified by
the same author. The typical specimens in the Vienna Museum are readily
distinguishable liy the following characters. Head above and back are dull sepia
brown, only the rump and upjier tail-coverts being pale green ; the sides of the head
and neck dark brownish (not rufous isabelline) ; no trace of yellow on forehead
or lores ; whole lower surface uniform pale yellow without any ochraceous wash on
the chest, the latter being greyish yellow, sliglitly flammulated with brighter yellow.
In the coloration of the upper parts, and in lacking the yellowish tinge on the
ibrehead and lores, the types of P. hijpoxantha. agree exactly with those of
P. fuscicapillu (Sd. & hialv.) (ex Eastern Ecuador), but the latter differ in having
the lower parts (except the dirty whitish throat) much darker, deep greenish yellow.
I am, however, not certain wjiether this difference will hold good when a larger
series of both races is compared. For the present, the following species and sub-
species ought to be recognised :
1. P. aura lit a fro itx anntHtiifroiis (Lawr.). Panama to Santa Marta.
2. P. ainaiitiifronK saturata Hellm. Nortliern coast of Venezuela, from
San Esteban to Cumana and Trinidad. (The locality " (iuiana" is erroneous, the
so-called Guiana skins being of the well-known "Orinoco" make.)
o. P. hi/jioxaiitlia h'ipoxantka (Pelz.). North Brazil : Kio Icanna and
1!. A'anjie, tributaries of the njijier Rio Negro.
4. P. h;! poxaiithu J'ii»i:irapilla (Scl. it Salv.). East Ecuador : Sarayac;u.
2 1 . Tachycineta t albiventer (Bodd.).
Hiriimlo nlljirenler Boddaort, Tuhl. PI. eiil. (178a) ]). 31' [luiscd on Daubenton. /'/. rid.lah.bi6.
fig. 2. — Cat/i'iiw],
One ? ad. from Seelet, April. •• Iris brnwn." Identical witli examjiles from
British (iuiana and Cayenne, only the bill being rather larger.
* TjiK ; .Muj. Tring, No. il7*S, Caracciolo coll. '■ 5 " a4., San Antonio, Cuniami, .Marcli Isth, 18it8.
t There is not the slightest ground for separating this species gcnerically as IridljinnHe.
( i:^ )
2'2. Progne chalybea chalybea (Gm.)
Jliruiiilri I'htilyhea Gmelin, Sysl. Nttt. 1. ii. (178S) p. 102i! [ha,sed on Brisson & Daubenton, PI. enl.
545. fig. 2. — Cayenne].
One ? ad. from Seelet, Ajiril. " Iris brown.'' Not different in any way from
Brit. Guiana and Pari'i examples. An adnlt male has been collected by Mr. Andre,
near Castare, Tobago, May lu, 19U3. It also agrees with mainland specimens.
23. Stelgidopteryx ruficollis aequalis Bangs.
IHiriiiidn rtificollis Vieillot, Nam: Did. xiv. (1817) p. 523 (ex Azara No. 30(j. — Paraguay)].
Stelqiddpteryx rufirnllis ueqiiaVm Ontram Bangs, Pioc. Ni')o Engl. Zaol. CI. ii. (1901) p. 58 [type
from Santa Marta].
1 rj ad. Laventille, May ; 1 ? Chaguaramas, Jannary : 1 <? ( 'aroni, April ;
and 1 S from Cangrejal, May.
These specimens have the rump ashy whitish, in marked contrast to the dark
brown back, agreeing in that respect with a good series from Cnmana, Garthagena,
British Guiana, Merida, and the Orinoco and ('aura Rivers. Tin's form, which has
been well described by l\Ir. Bangs, differs from S. r. aropyf/ialiii (Lawr.) in its
decidedly paler rufous throat anil paler brownish grey chest and sides. S. r. nijicollis
(Vieill.) is easily recognisable from both by having the whole upper surface
UMJfornily dark brown without any trace of the ashy whitish rnnip band.
I have but a few words about the geograiihical distribution to add to Mr. Bangs'
excellent account.
1. .S'. n/ficoliis 1-nfcolh's (Vieill.).
Distribntion.— Paraguay ; Brazil : Uio, S. Paulo, Minas Geraes, Bahia,
Mattogrosso, Gnyaba, Gaicara (Natterer coll.), west to the eastern
slopes of the Andes in Peru and Ecuador ; and east to Para on the
month of the Amazons.
Obs. I have before me 2 adult SS from Nauta (N.E. Pern) and 2 cJc?
and 1 ? from Archidona, East Ecuador, which agree perfectly
with specimens from Eastern Brazil, and show no approach to the
pale-rumped form found on the western side of the Ecuadorian Andes.
Three examples from Parii are also qnite identical.
2. S. ruficollis uropjujiuUs (Ijawr.).
Distribution. — Gosta Bica ; Ghiriqui, Panama ; and soutli wards thrnugh
Western Golombia (Remedios, etc.) to AVestern Ecuador.
Obs. A good series from West Ecuador and 1 i jr. from Remedios,
Antio(|uia, agree with others from Gosta Rica, etc., and avi' mui'lt
darker everywhere than .S'. /■. iicipud'n.
3. S. nificollis ueqnalis 0. Bangs.
Distribution. — North coast of (lolombia (Garthagona, Santa JIarta) :
Bogota coll. ; Venezuela : Jlerida, Cumand, Orinnco and ( 'aura Rivers ;
British Guiana (Roniima); Trinidad.
24. Euphonia trinitatis Strickl.
Eiijilmiki Iriiiilati.-! Strickland, Omlrih. Ornith. \Ki\. p. 72 ["Trinidad, Cuminu, Venezuela, St.
Thomas." — We accept Trinidad as typical locality].
1 S ad., ( 'hagnaramas, .January ; 1 ? , Pointe Gourde, January. They ayree in
every resjiect with a series from Gumana and the Orinoco River.
( 14 )
'i->. Euphonia violacea lichtensteinii*(C'ali.)
IFiiiKjillii tv'.i/mr/' Linnaeus, Si/xi. Xal. x. (1768) p. IS2 ('in calidis regionil)Us").]
Phniiaxi-ti lii-hleimtcinii Cabanis, Joiiin. Oniilh. 18G0. p. 331 [Cayenne].
One hniidrfd siipcimons of liotli sexes from fiiparo, Mareli : 1 r? ad., Poiiite
Gourde, January : 'i S ad., Valencia, March ; and one pair from Chagnaramas,
■Tannary. This series agrees perfectly with mainland specimens.
The name E. v. riolacra (Linn.) i.s to be retained for the Brazilian form, since
Gabanis, when first distingnishiug the two races, separated the northern snbspecics.
In the tenth edition, I^innaens gave no locality, and Cabanis was, thus, justified in
restricting the original name. In the Cat. Birds, xi., the names are jnst reversed.
~0. Calospiza desmaresti (Gray).*
Cdlliilf Desmiiirxt; (}. R. Gray, Gnt. BiriU ii. (June 1844), genus Calliste, Xo. (! [based on
"Tanagra gyrola" Swainson, XAiiil Ilhist. n. ser. tab. 28.— No locality. We substitute
Tiinklad. ]
Twenty-one adults and young from Cajiaro, March and April ; '-' S ad. from
< 'haguararaas, January. They agree with a large series from the vicinity of < 'umand,
and Estanijues near Mcrida, Venezuela.
2T. Calospiza mexicana vieilloti (Scl.)
[Tifutu/rd nie.c/rinni Linnaeus, Sij^t. Xul. .\ii. i. (ITtjii) j). ^Jlfi [ex Brisson ; Cayana. — (excl. .Syn.
Hernandez — Mexico).]
CalHsIt fielllnti Sclater, P. Z. H. 185f>. p. =>:,', [Trinid.ad].
Fifty specimens of both sexes i Caparo, March and k\m\ : Laventilli>, March :
Valencia, March.
All have the belly bright yellow, dift'eriug thereby very markedly from the pale-
bellied ('. m. media, Berl. & Hart.f which inhabits the Orinoco region. A series
from Guanoco, Orinoco-delta, belongs also to the latter race, but one specimen
approaches C. m. eieilloti in the colour of the under parts.
No Calospiza of this section has as yet been found on the north coast of
Venezuela, and ('. m. eieilloti appears to be confine<l to the island of Trinidad.
:i8. Calospiza guttata (Cab.)
Cullhpiza yulliita Cabanis, Man. Ilein. i. (1850) p. 20 ["Roraima, Guiana.'']
One ¥ ad. from t'hagnanas, July oth, agreeing with a large series from
('nmana.
■J'.). Tanagra episcopus sclateri Hcvl.
[TaiKii/ra episciijnix Linnaeus, Sijsl. Nat. xii. i, (ITOl'i) p. 310 : ex " LKvesque." — Rrisson.]
Tmnifim sclateri Beilepscli, Ibix, 1880, p. 11'2 (" Orinoco District oder Trinidad ").
A series of 'i\ specimens from Caparo, March and April : Valencia, Jlarcli ;
Chagnaramas, January; Fointe Gourde, January ; and Seelet, April.
ISjjecimens from (Juanoco (Orinoco-delta) and Cindad Bolivar on the lower
Orinoco agree in every way with those from Trinidad, while the birds i'rom Cumand
* Altliougli Swainson's description and synonymy e\idently refer to C. gijrola (l.imi.), tlie plate
seems without doubt to be intended fur tlie above si>ccics, since there is no trace of tlie gt-ilden yellow
huraoml \K\Xrh to be seen I So we may retain tin- iianie i\ tlcitimin:i1i for it.
t yov. Zmil. ix. 1902, p. la.
( 15 )
and from the npiier Orinoco (Altagrania, Maipures, etc.) form the transition between
sclateri and canu. Some skins from Cnmanu, however, are scarcely different from
the Trinidad series.
7". e. berlfpsrlii Dalmas,* from Tobago, agrees with T. e. cana in the bright
bbie colonr of the shoulder ])atcli, but differs in the more bluish, less greenish,
median and greater upper wiug-coverts, and iu having a decided violet or bluisli
wash on the under parts.
30. Tanagra palmarum melanoptera Scl.
[Taiiaf/m iialmariiiH Wied, Heise Brusil. ii. (1821) p. 76 (Canavieras, Bahia).]
Tanagra melaiinptera Sclater (ex Hartlaub MS.), P. Z. S. 1856. p. 235 (East Peru, etc.).
A large series from Caparo, March and April ; 1 S ad. from Ijaventille,
December; 2 i S , Chagnararaas, Januar}- ; 2 ??, Poiute Gourde, January;
3 ? , Valencia, March \ \ S , Seelct, April.
None of these specimens have any trace of olive edgings to the wing- and fiiil-
feathers, thus agreeing with topotypical melanoptera from East Peru. The latter,
however, show a decided violet suffusion ou the back and breast, almost or altogether
wanting in the Trinidad birds.
31. Tanagra cyanocephala subcinerea Scl.
\_Aglaia ciitinocephda Ijafresuaye & D'Orbigny, Siju. Ai\ i. in Maq. Znol. 18.37. cl. ii, p. 32 (Yiingas,
Bolivia).]
Tanagra subcinerea Sclater, P. Z. S. 1861. p. 129 (Venezuela).
One S ad. from Aripo, 2000 ft.. May 13, 1903. It agrees with a good series
from Cumana except in having the under parts a shade darker grey.
T. c. subcinerea is a strongly marked subspecies and easily recognisable by
its pale dingy grey under-surface, passing into dull whitish grey on the middle
of the abdomen. Throat and foreneck are slightly speckled with paler greyish.
This form is strictly confined to Trinidad and the mountains near Cumana, N.E.
Venezuela.
T. c. auricrissa Scl. has the under parts much darker : schistaceons with a
faint bluish tinge ; throat and foreneck are uniform dark schistaceons, crissnm and
thighs much brighter yellow. It occurs in Colombia (Antioquia and Bogota-coll.)
and in the Andes of Venezuela near Merida.
T. c. cijanocepliala (Lafr. & D'Orb.) differs from the foregoing form in its
paler grey under-surface (but not nearly as pale as in T. c. suda/ierea), much
lighter, lemon-yellow crissnm and thighs, as well as by its much more gieenish
back. It is found in Northern Bolivia, Pern, and West Ecuador.
32. Ramphocelus jacapa magnirostris Lafr.
[TatMgr/i jacapa Linnaeus, .S'l/s/ Nut. xii, i. 1766) p. 313 (Surinam— ex Edwards ; et Cayenne— ex
Brisson).]
liampliiicchis magnirostris Lafresnaye, Rev. Zool. 1853. p. 243 (Trinidad).
RanqihiiceJus alroserireus capifalis Allen, Bull. Aiucr. Mas. iv. (1892) p. 51 (El Pilar, near Cumana),
More than sixty specimens, adults and young, from Caparo, Valencia,
Chagnaramas, Seelet, and Laveutille. Examples from Cumana and from Gnanoco
in the Orinoco-delta are practically identical with those from Trinidad ; hence
B. a. ciijjitdlis becomes a synonym of nun/nirostris.
§
* Mini. Sue. Xoiil. Fruiicf xiii, (1900), p. l.ii;.
( I'i )
1 can recoguise the following races of the It. jacapa gronji :
(a) R.jacapajacapa (Linn.)
Distribution : ('iii/i'inic, Sini/iam, Bn'f. Guiarta : Lower Amazonki : Para (Steere
III)!, ill -Mils. Tring), JIanaos (Natterer); Vpjicr Amazonia : "^waia, Pebas, Ljuitos
ill N.E. Pern; (Jnayaliamba, North Peru (Baron coll.); F.aisteni Kcaador : Najio
(Gloodfellow coll. — Mus. Tring); Vcni'zaela : on the middle and ujtper Orinoco and
its tributary, the Canra River.
{U) L'. Jacajj'i )t/(iffmrosfri,s Lafr.
Distribution : Tiinidod ; and N.E. Venezuela: Cnmana (( 'aracciolo coll.),
and Gnanoco in the Orinoco-delta (Andre coll. — Mus. Tring).
This form differs from the typical one by its larger size (wing 80 — 85, instead
of 74 — 8U mm.) and larger, stronger bill.
(<■) li. jacajKi renezueleii.-ti.s Lafr.
Tliinijiliocehi.i: renezuelensi.'i Lafresiiaye, Ucc. Zool. lS.""v5. ]). ^43 (Venezuela).
Distribution : North coast of Venezuela, near San Estebau.
It differs from the two foregoing races in having the wliole ventral surface
nearly uniform dark crimson ; only a small patch on the lower abdomen being
blackish. In R.j.jucnpu and R. j. mayiiirostris only the throat and the chest are
bright crimson, while the breast is decidedly duller and the blackish jiatcli in the
middle of the belly more extended. The back is also more washed witli crimson
in /?./". cenezxelertsis than in its allies.
I have a good series of this furni from S. Estebaii, near Puerto Cabello
(A. Mocquerys coll.), which answers well the original desc^riptimi.
(r/) li. jacn/i'i nnicolor Scl.
RiimpliocelKs unicolor Sclater, P. Z. S. 1850. p. 128 (Bogota).
Distribution: Colomhiu : Bogota aoW. ; Mcridu, Venezuela.
Perhaps barely separable from R. j. cenezneleHsis. Seven e.xamjiles before
me differ only by the dorsal surface being almost uniform crimson, while in the
jirecedent subspecies the back is always rather duller than the pileuni.
((■) It. jacajKi coii/iecteii-i Bcrl. it Stolzm.
Rliiimiilioceliiii jacapn connectens Berlepsch it Stolzmami, I'.Z.H. Ln'.m;. p. ;')44
(( 'eiitral Pern).
Distrilintioii.— rVv(//Y</ and Soitth Peru, extending eastwards through the
inferior of Brazil (Mattogrosso, Goiaz) to Baliia and southwards to the Rio I'uraini,
wliich forms the nurthern frontier of the state S. Paulo.
This form ditfcrs from y?./'. />/«//;« and R. j. magrdro.'ftri.^ in having only the
throat and chest crimson, the breast and abdomen being dull blackish, with but
a faint crimson hue. The whole back, too, is uiiil'orm velvety blackisli.
Jl. atro.v'riri'iis (Lafr. it D'Orli.) from Eastern Bolivia is still ]iurer black
above and beneath, and the female is so very dilferent from that of the subsijccies
of R. jacapa that it may be regarded as sjiecilically distinct.
33. Piranga rubra rubra (Linn.).
Frin(jiUu nihrii Linnaeus, Si/xt. Xiil. ed. x. (176S) p. 181 (ex Catesby. Xul. HIkI. Cnnijinii i. p. 50
tab. 66. — CaroliiM, A'irginia).
Taiiagra aentiva Gmelin, Sygt. Naf. 1. ii. (1788) p. 88'.i (based on the samp).
Fjirantjd uestica Sclater, Cm. 13. lirit. .\liis. xi. (1880), p. 182.
t)ue ¥ ad. from Laveiitille, taken November 6, 11(03.
( 1- )
34. Phoenicothraiipis rubra (Vicill.).
Tiirhiii,h,n,m nihrr Vieillot, Nonr. Dirt, xxxii. (181'.i) p. 3r,0 (" ile de la Triniti' ").
:i c? !i(l. irom Caparo, iv. ; 2 <S ail. from Laventille, xii. ami iii. ; 1 S aJ.
and one liinl in female dress (marked " tj jr"), Chagiiaramas, i. ; 1 J ad., 1 J juv.,
1 ? ad. from Valencia, iii. ; 1 tS and 1 ? ad. from Aripo, 20n0 ft., v.
The males agree in every respect with three others from the neighbonrhood of
(Uiraana. The range of this species is restricted to the extreme north-east of
Yene/.nela, north of the Orinoco delta and the inland ol Trinidad. As far as I
I; now it has not }'et been recorded from Caracas.
35. Tachyphonus luctuosus Lafr. A: D'Orb.
Tiiclii//>!ioiiii.i JiirlKiisiix Lafresnaye & D'Orbigny, Sijn. Av. i. in Mn;/. ZnnJ. 18.17. cl. ii. p. 29
(Guarayos, Bolivia).
r. alliixpcciilarh Le'otaud, Ols. Triukhd 186G, p. .30(1 (Trinidad).
T. nlricajiillim Lawrence, Prnr. Anal. Philad. 1868. p. ?)6() (Trinidad) [= J jnv.] und,"- :
Lanio (.') lawrmni Sclater, Ihls 18H5. p. 27-', tab. C. fig. 2.
CMnrnspiiir/iis Untaudi Chapman, Aid- x. (1893) p. :i43 (Trinidad) [= ?] ; idL-m, Bull, Amer. Mint.
vi. (1894) p. 31 (Trinidad).
Two (J ad. from Valencia, March, agreeing with Bolivian skins, but rather more
gloss}' on the lower suri'ace. The female of this species lias been described as
CIdorospingus IcoUuuli. Chapman's careful description leaves no doubt, and
the differences from Cklorospiiigiis chryHogaster Tacz., pointed out b}* him, are quite
obvious when an actual comparison of both species is made.
Although I have not seen the type of Lanio lawrencci I do not hesitate to
pronounce it to be nothing else Imt a young male of the same species, changing
from the juvenile to the adult plumage I The blackish head and the intermixed
lilack feathers on the interscapnlium clearly indicate that the specimen wonld have
become black in more advanced plumage. A skin from Bogota agrees perfectly
with the description and figure in the Ibia. It is very unfortunate that the three
stages of plumage of this bird not only should have been described as three distinct
species, but also placed in three different genera !
3(i. Tachyphonus rufus (Bodd.).
Tii/mi/fii rtifii Bndd.aert, Tiihl. PI. rn>. (1783) p. 44 ( ? . — Cayenne : ex Buffon).
TtirliifphoiiHn mrial' tiriirt (Sparrm.) auct.
Many specimens from ('aparo, iv., v. ; 1 J ad. from Pointe Gourde, i. ; one pair
from Valencia, iii. ; 1 d ad. from Seelet, iv. ; one pair from Chaguararaas, i. ; and
4 <S6 from LaventiUe, xii, iii.
These liirds, as well as a series from Tobago, do not differ from others from
Cumana, Cayenne, and the Orinoco delta.
3T. Saltator striatipictiis Lafr.
Sallalor str'mtipirliis Lafresn.aye, Rrr. Zwil. 1847. p. 7.3 (Caly, W. Colombia).
<S'. albiioll'is auct. nee Vieillot ! *
One S ad. from ( 'aparo, March ; one from Chaguaramas, January ; and another
from Pointe (iourde, January.
They agree well witli .some siiecimens from near Cali, W. < 'olomliia (Hiia]] coll.).
* Cf. my paper on litllc-kuown types o£ neotropical birds wliidi will be published shortly.
2
( 18 )
The <liisky streaks on the lower parts are jjerliaps ratlier liroader, but this dues not
seem to he a constant character, for in a series from ("uraaiia botli broadly and
n;irrii\vly striped examples an' to be lound.
3S. Saltator olivasceus < 'nb.
S. ullrasceim Cabanis: in Scliomburgk, ii-'iae Biit. Gulmm iii. (1848) p. (570 (Brit. Guiana).
Seven adults of both sexes from ("aparo, April ; 1 c? ad. Valencia, March ; and
one i)air from Seelet, April.
The series agree jierfectly with others from Siirinrim and Cumana.
A young bird, collected near Caparo, April 12, is above olive-green (instead
of dark ashy grey), the snpraloral and superciliary stripes are pale sulphur-yellow,
the blackish malar stripe barely indicated, and the lnwcr jiarts stronsriy tinged
with yellowish.
39. Volatinia jacarina splendens (Vieill.).
[Tiimii/ni Jnrtiriim Linnaeus, Si//:!. Niil. xii. i. (1700) p. 314 (ex Marcgrave.— East Brazil).]
FringiUa xphndens Vieillot, Num. Diet. xii. (1817) p. 173 (Cayenne), i
0 S ad., S i imm. and 4 ? ? from Caparo, April.
The males have the axillaries and under wing-coverts entirely bhuk, and
there is but a small white spot on the shoulilers. Tobago specimens are identical.
4ii. Sporophila grisea (Gm.).
Liirla f/rhni fimelin, S;/.^l. Xul. 1. ii. (1788) p. 8.^7 (based on Daubenton, PI. enl. 393. fig. 1.
"Virginie" — errore ! AVe substitute Cnyenur as typical locality).
•i iS, ~ ? ? from Caparo, March ; 1 c? Chaguaramas, January : 1 cJ, Pointe
Gourde, January; 2 S S, Seelet, April ; and 2 63, Valencia, March.
The series is not very uniform, some specimens having a large white patch
on the sides of the neck, entirely wanting in others.
41. Sporophila gutturalis gutturalis (Licht.).
FriiHiiUri f/iiHiirnlia Lichtenstein, I>)-;. Didil. 18l'.3. p. i>l> (S. Paulo).
(Jne J ad., Caparo, Ajiril ; .and another from Aripo, Isno ft., May. Not different
from typical Brazilian specimens.
42. Sporophila miuuta luinuta (Linn.).
Lnrlii miiiiihi Tiinnarus. .S;/s/. Xnl. x. (17.')8) p. Hi'i (Siirinani).
2 6 ;i(l. and 2 V ? from Caparo, JIarcli, April ami ;\riiy ; 1 S from Seelet,
A]iril.
The (^(Jagri'c with speeinieiis IVom ( 'nmana and I'ritisli (Jniana. Two SS
from Surinam are rather darker below, but an example from Antionuia, Cdloinbia, is
rjuite as dark.
43. Euetheia fiiliginosa (W'ied).
FnmiiUiifiiliiimnna Wied, Ihllr. Xatunj. BiaxU, ?,. i. (1830) p. 028 [Camaniu in lialiia].
I'kiiiiiiiiirafiimnm Lawrence, Ann. Li/c. Ken- V.irl: x. (1874) p. 3110 [Trinidad].
4 cJ(? and 2 ? ? from Caparo, Ajjril ; and 1 ? from Valencia, March.
These specimens (topcjtypes of P. /iii/io.m) are practically identical with two
( ly )
Bahia skins (E. fuliyinosn). A series from (Jniuaua is also not ilifferent. This
species is not included in Chapman's paper on the birds of Trinidad, thongli the
type oi P. f/imvsn has lieeu collceted on that island.
44. Oryzobonis augolensis (Linn.).
Loxia migiilensU Linnaeus, Si/nt. Xnl. xii. i. (ITOfi) p. .'30:1 (ex Edwards ; " Aiignl.i " — prrnve !
We substitute Eaut Brazil as typical locality).
L. Iiirridu Scopoli, Ami. Nat. Wxt, i. (1769) p. 140.
2 <i<S and 3 ? ? from ( "ai)aro, March and April; S S <S from Scelet, April ;
and 1 S each from Laventille, December, and Chagnaramas, Jannary. This series
agrees e.xactly with specimens from Surinam, British Guiana, and Cnmami. I have
3") adnlt males from different localities before me, and none of them shows any
trace of a chestnut cross-band on the throat. Therefore it seems very probable
that 0. speciihiris Finsch,* if not au accidental variety, may yet turn out to lie a
different race.
45. Oryzoborus crassirostris (Gm.).
Loxia crussirnslris Gmelin, Si/xt. A'tit. I. ii. (I7SM) p. Hi'd [ex Latham : PTab. ign.— We substitute
Cai/fime.']
Two (?c? ad. from Seelet, March.
They agree with specimens from the Orinoco (Qniribana de C'aicara) and Upper
Rio Negro (Lamalonga and Marabitanas) in having rather large and strong bills.
The birds found in British (iniana have mostly smaller bills, but this is not quite
constant.
40. Ostinops decumanus (Pall.)
XaulliurmiK ihcumunns Pallas, Spicil. fasc. vi. (1769) p. 1 (Surinam).
Oatinops deatmanus insuhii-is Dalmas, Mini. Snc. Zool. France xiii. (1900) p. I.'i7 (Tobago).
Two c? and S ¥ ? from Caparo, April, and Seelet, April.
Count Dalmas separated the Tobago birds as a subspecies on account of their
smaller size and paler castaneous rump, but there must be some mistake here.
On comparing 4 cJcJ and 3 ? ? from Tobago with a very large series from
different localities on the mainland, I cannot find the slightest difference, either in
size or in colour. The females are always considerably smaller than the males,
the crest is much shorter, and the rump usually paler castaneous. These arc just tlie
characters by which 0. d . hi.'^iildiiti is said to differ from the typical form, and I
strongly suspect that ( 'ount Dalmas was misled by wrongly sexed specimens.
Measurements :
• /'. /.. .i. 1870, p. 5S3.
{ 20 )
47. Cacicus cela (Jjiun.).
PuniA Cela Linnaeus. Si/sl. Xnl. .\. (IT58) p. 191 ("in Indiis" — errore ! We substitnte SiiriiMin).
Tfiiui/im albimxlri.'! Linnaeus, .1A««. Ad. Friil. ii. Prodi: (ITG4) p. 31 (" America").
Oiioht.t pemh-us Linnaeus, S'//»7. .Y"^ xii. 1 (ITiii'i) p. Kil ('■ in .\ineiiea mcriiliona'i ").
Citaxit'tis pet'AicKn auct.
Five adnlts I'rom ('a])aro, Ainil and Jlay ; imf iiair t'runi Seclot, Apiil. Tlit\v
agree perfectly with spcrimons from Surinam ami ( 'iimana.
N.B. — Tlio above is tlie proper name for tlie species commonly called C. pcrsiciis,
as has been brony:lit to my attention by Count Berlepsi-h. Linne's diagnosis of
Parus cela reads as follows : " P. niger, rostro albo, macula alarnm basiqne caiidae
flavis, Mus. Arl. Fi: 2. p. . . . Habitat in Indiis." Turning to the " Museum Ad.
Frid.", we find but one species, T<i/iai/iri nlhiroxtris, to which this description can
a]iply. The diagnosis given ibr that bird is the same, word for word, but a more
detailed account is added that renders the ideutilication absolutely certain : (!f.
" corpu.s sturno fere mains, nigrum, basi pennarum in tergo albo."
48. Xanthornus xanthornus xanthornus (lira.).
Orinhiii raiilhorniia Gmelin, Sijst. A'nt. 1. i. (17>I8) p. 3;il [(excl. Edwards and habitat : Jamaica) ex
Bris.son & Buffon, '■ Mexico " — errore ! We fix OniPiine ex Buffon as the typical locality.]
A good series from Chaguaramas, January ; Seelet, April ; C'ai>aro, March and
April ; Pointe Gourde, January; Laventille, March.
These examples have generally longer and stronger bills than others from
( 'umana and the Orinoco valley, and the wings average 2 to 3 ram. longer. One
specimen from Margarita Island is somewhat intermediate. X. x. curaqoensis
(Ridgw.) has the bill still longer and more slender.
Immature birds are much less brightly coloured tlininghonf, more greenish
yellow aljove and less orange below.
4'.t. Molothrus atronitens < 'al
Muhithms iitriimti II'! Cali.inis : in Schomburgk, Heism Brit. O'liiaiia iii. (184H) p. G8'2 [coast of
British Guiana].
^f. minimiiK Dalmas, Mcni. Hoc. Zvul. Fniiirr xiii. (litUO) p. 1.S8 (Tobago).
3 cJcJ from ("ai)aro. April ; 1 c? ad., C'aroni K., April; A 66 and 3 ¥ 9 from
Seelet, April.
Tiiese sjjecimens agree in coloration and size with a series from Tobago
{M. >ni)iimn.s Dalm.) and with the type of J/, iit/onitens Cab. in the Berlin Museum.
I suspect that Ualmas, when describing his M. mini/mis, compared it with
M. renezuelethvs Stone, which he might have mistaken for M. atronitens.
M. renezuelethv.'i is indeed mucli larger than the latter species, as will be seeu from
the following measurements.
M. atronitens Cab.
(J ad., British Guiana (Schomburgk coll.)
Type of the species. Mus. Berlin . Win^
8 sexed 6 6 from Trinidad ......
9 „ „ „ Tobago {M. minimus Dalm.)
1 .. c^ ad. from Para (Xatterer coll.) . ,,
1 ,, „ from K. Cauanie, Hio Branco
(Natterer coll.) „ lUU ; „ 77
( 21 )
M. cenezuelemis Stone.*
4 sexed c?(?, Puerto Cabellii, Venezuela . . Wing 114— llf,; tail 86— 0(1 mm.
1 ,, i ad. Lagunillas . . . . ,,114 ; „ 89 „
6 „ c? c? from Merida „ 111)— 118; „ 84—01 „
4 ,, S ad. from Qniriljauu de Caicara,
Orinoco ,, \\--i — 110 ; „ 88— ',_)4 „
50. Agelaius icterocephalus (Linn.)
Oriijluf irlirorrjihalH.'i Liimauus, Si/sl. Xat. xii. 1 (17GG) p. 163 [•■ Cayaiia," ux lirissou A
Edwards].
A good series of ailnlts and yonng birds from Seelet, April, and ('anini, I\[aveli
and April, not different from (Jayeune specimens.
51. Leistes militaris (Linn.)
Emheriza lu/likiris Linnaeus, S;/»l. yul. x. (1758) p. 178 (ex "America" — "Asia," orr. ! — We
suljstitute Surhiam).
Oriolus (juiaiifitxh-\- Taiiiiijra militaris Linnaeus, .S'</.s7. Nut. ed. xii. i. (lyCiCi) pp. 162, M16.
Leistes f/iiitiiieiisis Sclater, Oil. B. Bril. Miis. x\. p. 348.
2 SS and 2 ? ¥ from (!aroni, iv. ; and 1 cf from El Socorro, iv. These
specimens agree perfectly with others from Surinam, British Gniana, and the
Orinoco Uiver.
52. Quiscalus lugubris Sw.
Qiiiscalns hii/iiliris Swaiusou, Aiiim. in Mvmuj. (18.S8) p. 299 ("Brazil," err. ! — We substitute Bi-it.
Guiana).
One c? ad. from (.'aroni, 25 iv. 1003, agreeing with specimens from t'umana, and
the Orinoco region. One adult from Lagunillas near Merida, W. Venezuela, is
apparently also not different.
53. Cassidix oryzivora (Gm.)
Orhilus arji-Jeorus Gmelin, Sust. Xat. 1. i. (1788) p. 386 (Cayenne.— ox Jjatbain I
1 S ad. from Seelet, iv.
54. Arundinicola leucocephala (Linn.).
Pipra kncori-jihala Liuuaeus, .l/«s. .Id. Fiid. ii, Prodr. (1764) p. 33 {Sariiiam : cfr. Si/xi. Xat. xii 1.
p. 340).
One <S from Caroni, iv., agreeing in colour and size with a series from Brit. Guiana
and the Orinoco Uiver.
55. Fluvicola pica (Bodd.)
.Miiniiaiiupira Boddaert. Tal,i. PI. ml. (1783) p. 42 (ex Daubeiiton. tab. 675 f. i.— Cayeime).
6 c?c? from Seelet, iv., 1 S ad. (sexed ? 1) and 1 S jr. from Garoni Swamp, ii.
Tliey are in no way dift'erent from a very large series from the Orinoco, Brit-
Guiana, ('uman;i, Maracailid and I'ogota coll.
* Auk 1S',I1.-|,. 3j7 (Venezuela— Goeiing- e,.)I.)
( 22 )
oO. Platyrhynchus mystaceus insularis Allen.
Plai>jrh<j,irhii> i,is„l,n-i.'< Allen. BnII. Amrr. M,,^. ii. (iss'.i) p. 143 [Tobago].
1 c? from Laveiitillc, j\[arili, aud 1 ? from Caiiaro, April.
They agree with three adults from Tobago (toj)otypical F. i/tsular/s) in every
respect. Specimens from Cumaiia and Tocuyo in Nflrthern Veueznela belong also
til tliis form, whieh dillers from tlie typical race of Paraguay and S.E. Brazil in its
dull greenish (instead of warm oil-brown) upjier ]iarts, less brownish crown and
much paler nnder-snrface, the chest being but slightly washed with brownish. Tlie
birds from the upper Orinoco (Caieara) and the Horainia Jits., Brit. Guiana, are
somewhat intermediate between the tyjiieal and the northern race, Ijut nearer the
former ; yet they differ in having the back slightly paler brown and the under jiarls
not (juite so dark. Very likely they rei)resent another distinguishable form, but 1
would like to see more specimens before describing it. The difference in size upon
which Jlr. i\llen laid much stress is not constant.
ST. Orchilus* ecaudatiis (Lair. A M'Orb.).
Tnfliriislniin rciiudnlii in Lafresnaye & D'Orbigny, S////. Ai\ i. in Mwj. Zi«iL IS.TT. d. ii. p. 47
(Yuracares, E. Bolivia. — type in Mus. Paris examined).
One pair from Savannah Grande, February 7, 1897, collected by l>r. I'erey
Kendall (Nos. 35, 36)— " Iris dark brown (?), yellowish brown (S), feet steel-
colour, bill greyish horn-colonr, tipped with dirty white."
The specimens from Trinidad are in no way different from others, collected
by Natterer on the Rio Madeira. The species is for the first time recorded from
Trinidad. The most northerly locality hitherto known was San Esteban, in
N.AV. Venezuela, where Prof. Goeriug collected a specimen. (Jherrie sent an
e-xamjile from Muaduapo, ]J. Orinoco, and Klages another from Snajiure on the
('aura U., to the Tring Museum.
08. Mionectes oleagineus pallidiventris nov. subsp.
[Jlii.fiycajin oleai/hira Lichten.st(:in, ]'ii::. Diilil. (18:i3) \>. .')."i (Baliia).]
A good series from Caparo (April) and Valencia (March).
These specimens, as well as a good many from Cnmani'i, and mw. male from
Castare, Tobago, differ from 3i) skins from more southern localities in the decidedly
])aler orange colonr of the belly, much less greyish green suffusion of the throat,
and duller olive green ujiper i)arts. The differences are so obvious when a series is
compared that 1 propose to sejiarate the northern race as
Mionectes oleagineus pallidiventris Ilellm.
Type in Mus. Tring : i ad. IliKcnii o'i San Aiitnnio, Ciiinniiii, March 14, lMt8.
Caracciolo coll.. No. 850.
M. 0. ax.si'milif Scl. and ^f. 0. parcHg Bani;s, from ( 'entral America, are at once
known by their much paler and duller colonr of the abdomen, considerably darker
greyish green throat, and darker green upper parts.
* Pn-hxiifrhruA Ohovliolser [Pror. L\ S. Xa/, .}fi/s. xxv, li)02. p. I>1 (Type : Todirosh'inn trftudatiint
I.afr. & IVOib.)]. This f,'enus is not, .separable from Orchilus, sinre 0. alUvrntrh lierl. & Stolzn). connects
O. aiiririiluri.i witli I lie two sliort-taileil species, 0, ccaudatux and 0. atricajnlliia.
( 23 )
50. Ornithion pusillum (<'al>. & Ileiue).
Mijinimthpmilla Cabanis & Heine, J/«». Ilrui. ii. (ISoO), p. 5S (Cartagena, N. Colombia).
3 ? ? , 1 ? from Caparo, iv. ; 1 ? , Seelet, iv.
They agree with a ver)' large series from Cnmana, tlu; Urinuco region,
Snrinam, and Bogota coll. Costa Rica birds differ somewhat by their slightly
greener back and generally brigliter yellow nnder-parts, and may stand as a
subspecies, U. p. suhjlamnn Cherrie.
<iO. Elainea gaimardii CD'Orb.).
iliisckaiiara GiilmunU't D'Orbigny, l'".'/",7'' (P"W- between 1838 and 1817), Ok. p Mf> [Yuraeares,
North-east Bolivia].
Fonrteeu specimens of both sexes from Ciij)ar(>, March and Ajiril.
They are mostly paler yellow liclow thaQ a large series from the Orinoco and
Canra R., Guiana, (Jiimana, and Bogota coll. ; but some are hardly different in
that respect.
01. Elainea pagana pagana (Lcht.).
Miisinipii j«i;/iimi Lichteustein, Vn-:. Diihl. (1823), p. hi [Bahia].
Three females from .Seelet, Ajiril, and three adnlts of both sexes from C'aparo,
Aiiril. They have usually more white in the crest than typical Brazilian liird-;,
aiH'eeing in that respect with others from Venezuela, Tobago, Guiana, and Bogot,
collections, though this character is not iiuite constant.
a
02. Legattis albicollis (Vieill.).
Tyraimii.H alhlrollls Vieillot, Xmii: Did. xxxv. (1810), p. 80 [ex Azara— Pavagua)].
One female from Caparo, April, and a male from Seelet, April, agreeing with
a series from British Guiana, Snrinam, and Bahia. Typical Paraguay skins were not
available for comparison.
03. Myiozetetes sulphureus (S])ix).
Miisn.aiM siiljihi'irn Spix, Ar. Bias. ii. (1825), p. 1(5, tab. xx. ["in Brasilia"].
BIr. Andre did not send this species, and its occurrence in Triuidail rest.s on (he
single specimen collected by Mr. Chapman at La Brea.
04. Rhjaichocyclus flaviventris flaviventris CWied).
Musriprl,, fririrndns Wied, Bcilr. Xuhny. Brasd. .3 ii. (1831), p. '.)2'.l [Mucuri and AIcnba.;a :
Espiritu Santo, S.E. Brazil].
!d i S and ? ? iVom Cupam, Aiiril, ] 3 fnuu Valencia,. ;\ia.rcli, and 1 ? from
Seelet, April. They are not different from Bahia skins, and one adult from Tocuyo,
Venezuela, is also identical. N(ine of the many specimens before me apiiroaches
Ix'.f. hnrbac Ilcllm. in the colour of the under parts.
05. Rhynchocyclus sulphurescens sulphurescens (Spix).
riatyrhnnrhuH mlphiiresrciiH Spix, Ai\ Bras. ii. (182.5). p. 10, tab. xii. fig. 1 (" S ") 1^'" de Janeiro,
Piauhy, etc. — We accept liin as typical locality].
8tJ(?and ?? from Caparo, March and April. Besides, the Tring Jlnscnm
has one " ? " collected at Tacarigua in February by Dr. Percy liendall.
( -'^ )
Tlie series agrees in every iletiiil with sjieciiuciis i'rom Ciiinaiui mul S. Paiilo in
S. Brazil, the pileiim beinfr olive u;reen like tlie Ijack, and but slif;lit]y mixed with
grey. A nunilier of sl< ins from X.E. Peru (Xeberos and ( 'hamicuros, E. Barth>tt
t'oll.) have the pilenm grey with very little greenish admixture, and the nnder-
snrface distinctly pale yellow. They ought to be called A', s. u/asimilix I'elz.,
although the types of the latter form have a duller, more greyish greeu back.
06. Pitangus sulphuratus trinitatis nov. subs]).
[Lniiiiis sulphumtii.^ Linnaeus, .%s7. A'//, xii. i. (ITliil), p. lo7 (ux Brisson : '' Cai/miia ''].
Pilaiigus nulpliuiiitics (nee Linno) Chapman, litill. Antri: Mux. X. Y. vi. (1894), p. 40 (TriuiJail).
8 c?(? and ?? from Caparo, Marcli and April, 1 S from ( 'haguaramas,
January, and 1 ? from Seelet, Ai>ril.
The birds from Trinidad are exactly intermediate lietween typical /'. .s. sul-
jj/tuiafi'S and P. s. rufpcnnis (Lafr.). The differences of tiie three suljsjiecios may
be expressed as follow.s : —
((/) 1'. .s. m/i'/i'iraOfx (Linn.)
Uj)per wing- and tail-coverts with scarcely any trace of rnf(jus margins ; (piills
narrowly edged with rufous; outer tail-feathers with the extreme base of the inner
weiis only rufous.
Ilab. Surinam, Cayenne, Brit. (Juiana, and Amazouia, from I'ara to N.E. Peru
aud East Ecuador.
(b) P. s. trinitatis Hellm.
Upper wing- and tail-coverts distinctly margined with rufous ; quills more
broadly edged with rufous, these edges being separated from the rufous colour of
the inner webs by a distinct dusky stripe along the shaft; tail-feathers with broad
rufous margins occupying about half the lireadth of the inner web.
Type in Mus. Tring : " ? " ad., Caparo, Trinidad, April Id, E. Andre coll.
Ilf/i. Trinidad.
(c) P. s. rufpennis (Lafr.).
SftitnijiIuKjiis riiji2>eniih Lafresnaye, Rcr. Zunl. 1851, p. 471 (Gankas).
Upper wing- aud tail-coverts almost entirely rufous, only a narrow central
streak loeing dusky ; both webs of the quills (except the tips of the primaries and
a broad shaft-stripe on the apical half of the secondaries) and inner web of theonter
tail-feathers entirely rufous.
Ildlj. Venezuela: from C'umana to S. Esteban, south to the Orinoco \alley
and its tributaries; Colombia: Cartagena, Santa Marta, Cauca valley, aud Bogota
collections.
The above differences are ipiite constant in a series of .i4 adults.
•57. Myiodynastes audax (Gm.).
MHtt:icapaiiuda.rGmfs\\n,Sij^l. Xnl. 1. ii. (1788), p. 034 [ex Uaubenton, PI. en!. 453. fig. 2 : Cayennel -
1 ?, Valencia. Jlanh ; 1 ¥, Puinte (tourde, January; and 1^ adults of botli
sexes from Caparo, March. They agree witli a large series from Tobago, ( 'umana,
Merida, and British Guiana. It is certainly wrong to treat .1/. solitarius (Vieill.)
( 25 )
as a siilispecies of .^f. fu/f/n.r, siiifu tliey iicc.iii- t,u;;etla'r at vaiimis Incalitics. In the
Tring Museum there are both sjiecies from tlie ]{io HujiurLimi, British Guiana, and
Natterer collected both at Barra do Rio Negro, N. Brazil. M. solitariitH has never
any trace of rufous on the tail ; the black stripes ou the under-surface are also more
extended and broader, especially on the throat, which in M. aiuhix is almost devoid
of dusky markings. The back, too, is blacker, and the rufous edges of the upper
tail-coverts much darker.
68. Megarhynchus pitangua (Linn.).
Liuiina p'llnnijiin Linnaeus, StjM. Nat. xii. i. (1766) p. 13ii (ex Brisson : " Brosil ").
Eleven adults and one young bird from ( 'a])aro, March and April ; 1 c? ad.
Laventille, December; 1 (?, Pointe (iourde, January; 'Z $ i , Chaguaramas
January. They are not different in any way from topotypical Brazilian skins.
6tt. Myiobius fasciatus (P. L. S. Mull.) .
Muxi-irrijin/ufC/atn P. L. 8. JliiUer, ,Y(//«)■^//^7. Suppl. (177(5) p. 172 ["Ca/r«/)( "— e.x Daubentuu, PI.
enl. hii, fig. 3).]
Jfn.tcioajxi iiarria Boddaert, Tab/. PI. ml. ( 1 7x3) p. 34 [based on Daubentoii].
Mlf'n}hiuf< uafcf'fs auct.
1 c? ad. from C'aparo, April, and I ? from Seelet, April, agreeing with
examples from (Himana, Cayenne, and Brazil. In the male the crest is lemon
yellow, in the female ferrnginous.
'". Empidonax lawrencei Allen.
Orhlhnrrii tl,n-im,lris Lawrence, Ami. .%'<■/'■ Ynrk Ar. Sri. iv. (1887) p. (57 (loc. ign.) uiide ;
Empidniiiix htirrfwri Allen, Bull. Aiiui: .!/«.<. .V. 1'. ii. (18811) p. 150: Chapman, Bull. Aiiiry. .Mkx.
vi. (1894) p. 4-2 (Trinidad).
E. olim Sclater, Cal. Birih Biil. .Miis. xiv. (1888) p. 224.
Seven specimens of both sexes from Caparo, April. They agree in every
respect with a series from Camana and the Caura River, Venezuela.
E. luwrenrei is the bird commonly called E. olivri or /:,'. plleata (cfr. Sor. Zool.
ix. 101(2, p. 174) ; but it is almost certain that the " Gobe-raonche olive, de Cayenne "
of Daubenton, upon which both names have lieen based, does not refer to the species
in (piestion, which is not known to occur in Cayenne. It seems, therefore, best to
retain Mr. Allen's name.
Measurements : \Ving 01— Uts ; tail .J.J— 05 mm.
Tl. Blacicus brachytarsus (Sd.) (subsp. ?j
Eiii/iirlniiii.i- bnirlii/tarsiin Selater, Jhix, I85',l. p. 441 [Cordova, South Mexico].
A series of •,'0 specimens {S d and ? ?) from Caparo, April; Pointe (iourde,
January, and Valencia, IMarch ; agreeing with others from Cumanu and Mount
Bucarito, Venezuela.
Two skins from Vera Paz, Guatemala, hnve decidedly darker under ]iarfs. but
are not otherwise different. Perhai)s the South American birds could be sejiarated
subspecitically. EMjiii/ona.r ai)r/i//>t.t Tacz. seems to be the same as ('. >■//■<'/'.•<
(Liim.), so they would re((nire a new name. A larger series, however, should lie
examined.
( 2(1 )
T'.'. Myiarchus tubeixulifer ( t-atr. ^, J)'l)rli.).
Tyraiiiiiin litherculifer Lafresnaye & D'Orbigny, %/i. Ae. i. in Mag. Znnl. 1837. cl. ii. p. 43
[Guarayos, East Bolivia].
1 6 and 2 ? ? from Caparo, April : 1 J from Aripo, 2niiii ft., 5Iay.
These examples agree e.xaetly with a large series trom Cnmaiid, Brazil, British
Guiana and Bogota coll. In Hie C'tl. Birds Brit. Mus. most of the specimens of
this species have been jilaced under M. /liffrice/is, while two others arc registered
as a different species : .1/. Irii-olor !' As a matter of fact, however, the former are
absolntel)- identical with those from Baliia and Uio Tocantins. The distribution of
the two species or subspecies is as follows :
(a) Mi/iarchiM tuberculifer (Lafr. & D'Orb.)
Brazil: ]{io, Bahia, Mattogrosso, Para, I'.orba ; /■.'".v/ Bnlirin : (Jnarayos ;
Ujjjier Amozonia : Iquitos, N.E. Peru ; Sarayaen, East Ecuador ; Coloinhia : Bogota
coll.; Venezuela: Merida ; S. Estelian ; Cumana : Orinoco valley (Quiribana de
Caicara, Nericagna) ; Trinidad: Brit, (iniaim : Koraima, etc.
Obs. Having not seen specimens from Santa Marta and Panama, I cannot say
whether they belong to this or to the next form.
(4) Myiarchus nigriccps Scl.
Western Ecuador : Pallatanga, Esmeraldas, Chimbo, Bngnac, Nicbli, etc. ; and
yprfherH Peru: Tambilln, Callucate, Cnlervo, Ilnaiubo, Tamiapampa,
C'ajabamba, Malea, etc.
'■'•>. Myiarchus tyrannulus tyrannulus (P. L. S. Milll ).
.Uusr.iriipa li/nni,iii/i(g P. L. S. Miillor, Xalin:'<i/iil. Siqipl. p. 1«0 (1771',.— Cayenne— c.k U'Aubcnton,
/'/. (•»/. 57I,fig. 1).
Mij'ianhis cfi/llii-na'rciis Sclater & Salvin, P. Z. .S. 18l'iS. pp. 6-28, 631 (CarAoas).
1 tj ad. from ( 'liaguaramas, January : 1 i, Laventille, December ; and 3 ? ?
from Seelet, April.
These specimens, as well as some others from Tobago, are practically identical
with a series from ('nmana and the Orinoco Biver. They evidently represent
typical M. ti/rannulu.^.
.\r. t. ohcri Lawr., from the Lesser Antilles, differs in its larger and longer bill,
decidedly darker njiper parts, and in having the whole inner web of the outer tail-
feathers rnfons. .1/. t. brccijjcnnis Hart., from Curacao, etc., on the other hand, is
much smaller, especially witli a considerably shorter and weaker bill.
T4. Tyrannus melaucholicus satrapa (Cab. ^V, Heine).
[ TyramiuH mrl„i,-h;lkiis Vieilint, .Y""c. Did. .x.x.w. (1819) p. 84 (ex A/.ani : Paraguay).]
Laphjrtex Hnlrnjiii Cabanis & Heine, .!/»«. Ilrinmn. ii. (1850) p. 77 (" Guiana and Caracas ").
Many s[.ecimens from Caparo, Aiiril, agreeing in size and colour with a series
from Tobago, Panama, Costa Bica and British Guiana. The nortliern form differs
from typical T. m. melancliolims only by its somewhat larger size. On the other
hand, T. m. couchi Baird is a strongly marked race, easily known from those of
South America by its largrr size, paler, more whitish throat, much less greenish
suffusion on the foreneck and ninch paler yellow belly.
( 27 )
T.s. Tyrannus rostratus Scl.
Tymn/iKK mxlmlns Sclatcr, Ibis, 18(i4. p. 87 [Port of Spain, Triuidad (Taylor) and Guiaua.]
1 S acl. from Seelet, Aiiril ; ami auuther 6 i'rom Lavuiitillc, Ueceiiiljer.
They are absolutely identical with a large series from the Antilles (tjanta Lucia,
Dominica, etc.). Although originally described from Trinidad, tiiis bird is certainly
only a winter visitor on the island.
TC). Muscivora tyrannus (Linn.).
Miisacojia li/raiuim Liuuauus, Si/d. ym. .\ii. 1. (17Glij p. 325 [tx Brissou : '• Canada" (orrore !) and
Cayenne].
One adidt i'rom Caroni, A[iril.
77. Pipra erythrocephala (Linn.).
Parus enjthnKfplmhis Linnaeus, Si/>l. yal. x. (17i8) p. I'.H (ux Edwards.- Surinam).
Pipra anrnvnpiUa Lcht.
A good series from ('aparo, March and .Vi)ril ; Valencia, March ; Pointe Gourde
and t'haguaramas, January.
Like others from Cumana, and (inanoco in the Orinoco delta, the S S have the
cap brighter, more orange than tliose from Cayenne and Surinam. Specimens from
the Orinoco and t'aura are, however, so variously intermediate tliat I do not venture
to separate them.
78. Chiromachaeris manacus manacus (Linn.).
Pipia iiiaittiriif: Linuacus, %.s(. \at. .xii. 1. p. 240 (ITGU.— e.x Kdwards ; "supposed to be Sumntin,"
and ex Brisson : part).
A very large series of both se.\es from (_'ai)aro, March and April ; one pair
from Laventille, December; 2 (i S from Valencia, March; and :2 JcJ, 1 ? from
C'haguaramas, January.
The males agree witli several specimens from Surinam in having only the sides
of tlie belly and the under tail-coverts grey. 2^6 from Archidona (East Ecuador),
one from the (Jaura River, and several from tlie Hio Negro (Natterer), belong also
to this form.
C. III. jjxrm (Bangs) from the Lower Amazons, is easily recognisable by its
white under tail-coverts and narrow black area on the back.
711. Tityra cayana (L.).
Lnn/Hs mi/iiniis Linnaeus, Si/sL Xnt. xii. 1. (17G6) p. 137 (ex Brisson.— Caycnni).
-! c?c? and 2 ? ? from Caiiaro, iii., iv.; and 1 6 ad. from Valencia, iii.
Apparently not ditt'erent from (Cayenne and Cumana specimens, Imt tiie males
are a shade more cinereous, both beneath and above.
•sti. Pachyrhamphus niger cinereiventris Sd.
[Piiiiiijvijiirhiis H/'^r,- Spix, /I '■. />'/</^. ii. (IS-J.'i) p. 3:;. tal>. 4.'i. tig. I (no locality given.— We subslilute
Fuiitrbua^ Amazon.i, as the typical locality).]
Paclit/rluiiiqihiit fiiirnirriiliis Sclater, Oil. Aiiar. IJhds, p. -Hi (1862.— Santa Marta).
Parlu/rhamphus nii/cr (nee Spix !) Chapman, /.''///. .!«»,■, .)/»«. vi. (1894) p. 47 (Trinidad).
2 (?c? from ('aparo, Ajirij, niid 1 c^ ad. from I'ointe (Gourde, January.
These specimens agree with a scries from Cnmnna, Tobago, the Orinoco valley,
and Bogota-collections. I compared also the type of /'. ciiieificen/ria from Santa Marta
(28 )
ill till' I'litisli Museum, ami liniiul il tn bolnug tn the sauu' Inrm. The luidei' surface
is (lark cinerous, more or less freckled with dull blackish, especially on the throat
and I'oreiieck. The true P. n. niijer from N.E. Pern and East Ecuador differs at a
glance by having the lower parts uniform deep black, the rumji and npper tail-
coverts black like the back (never grey as in ci iiereiventris), and by its larger bill.
81. Chasmorhyuchus variegatus (Gm.).
Ampdis mritgiila Gmelin, Syxt. Xal. 1. ii. (1788) p. 841 [ex Brissoii, etc. — " Brasilia "—erroic 1
We substitute Caijcmir.']
1 S ad., 1 S jr., and 1 ? from Aripo, L'tMil) ft., May ; 1 S ad. Caparo, Jlay ;
] i ad., 2 SS }i: and a ? ? from Valencia, March.
This series agrees well with several specimens from the Itoraima Sluuutains,
British Guiana. The species has not yet been recorded from the " Cote de I'aria,"
but Gocriiig collected it at S. Esteban, near Puerto f'abello.
s:,'. Sclerurus albigularis Scl. & Salv.*
Schrurus alhiijiilarls Sclater & Salviu, P. Z. S. 1868. p. 630 [Caracas.]
1 ? ad. from Valencia, March 25 ; and 1 c? from ('aparo, Ajiril 1~.
Identical with specimens from Tobago and Cnmaua.
S. caniyuluris Ridgw. has i|nite erroneously been united with the above species
by Dr. Sclater. It is a very much darker bird throughout. The ui)per parts are
dark rufous brown (instead of pale olive brown), the foreneck deep chestnut rufous
(instead of pale ferruginous), breast and abdomen dark rufous brown mixed with
blackish, and the whole throat is uniform dark ashy grey (chin and ujipcr throat
not whitish, as in S. idbiijiilarix).
83. Synallaxis cinnamomea (Gm.).
Certliia cbinamnmeu Gmelin, Si/.sV. X'lt. 1. i. (178S) p. 480 [ex Latham: loc. ign. — We accept
Cayemie as the typical locality].
2 SS from Caroni Swamp, March : '^ S S ad., 1 ? ad. and '~ juv. from Seelct,
April ; agreeing with a good series from Surinam, Uritish Guiana mid the ()rim)co
Hiver.
84. Synallaxis albescens albigularis Scl.
[Hnwillajia albescens Temminck, /'/. n,l. lirr. 88 (Sept. 1823) tab. 227. fig. -1 (S. Brazil i].
,S albignlaris Sclater, /'. Z. S, 1808. p. 63 [Rio Napo, East Ecuador].
.S. ncr'ipilalU Madara-sz, Ann. Mux. Xat. Hunij. i. (1903) p. 463 [Merida, Venezuela].
Three S S from Caparo, Blareh and Ajtril ; and one young bird from Laventille,
Blarch.
The adults agree with a large series irom the Orinoco, Merida, Bogota, and
East Ecuador. Through Sclater's " lumping " of S. albescens albescens and S. a
albiiiiilnris in the Cut. JJ. .\v., Dr. von JIadaiasz was misled to create a new name
for the northern form. The latter may be distinguished from the Brazilian bird by
the much broader, greyish brown frontal band, paler, more greyish brown (instead
of reddish brown) tail, and, as a rule, lighter rufons, less castaneoiis colour of the
crown and shoulder patch.
♦ Tlie ii;imc i> commuiily atuibiited to SSw:iiiifOn (Bhils Jliazil, vlc, iAh. 7sj, but that platf is not
to be found in any u£ the copies 1 cuusulted.
( --i^ )
■s."). Synallaxis terrestris carri (hapm.
Si/, Ill/Ill. lis rurri r!li;i|iman, Hull. Ami-r. Mim. vii. (1895) p. I!'i3 [Caparo : Ti-iniciailJ.
One S iiil. IVom Cajiaro, coUeftcil April 4, VMYZ.
This is a vny distinct I'orra, at once known from .V. t. fi'nvsfi-i.i .Tard. In- the
following characters : tlie n])iier [larts are much darker, mnmmy brown (instead
of i)ale olivaceous brown), the wings darker chestnut, and tlie breast and abdomen
also much more intensely colourgd, scarcely lighter than the back. The whitish
spots on the throat are greatly restricted, the blackish ground-colour becoming
much more apparent. Only the anterior portion of the chest shows a few narrow
fulvous shaft-lines, while in -S'. t. terre.stria from Tobago the whole breast is
covered with distinct pale stripes, which are slightly margined witli dusky
laterally.
8. t. strir/tijii'i'tus ( 'hapm. from Cumana is another close ally, but differs in
the decide<lly rufous sides of the neck and superciliaries, and in having tiie whole
lireast and abdomen marked with broad fulvous shaft-stripes.
■SO. Xenops rutilus rutilus Licht.
Xeiiups riil'ilua Lichtenstein, Vei-~.. Diibl. 18J3. p. 17 [Baliia].
One (? ad. and one specimen not sexed, from Laventille, December and April.
They agree with examples from Bahia, S. Paulo ami Cnmana.
87. Dendrocincla meruloides (Lafr.).
Deiidrocnps meviilniih-s Lafresnaye, Rev. Zmil. 1851. p. 4lj7 [" Cote ferme " coll. Beauperthuys. — This
means the north coast of Venezuela near Ciiiiuind '].
Dfiidriiciiirla iiii'mhiidea riphanln Oberholser, Proc. Acail. Phihitl. 11IIJ4. p. 4GU (Tobago).
Thirteen specimens of both sexes from Caparo, April ; ] S ad. from Pointe
Gourde, January.
The series is fairly uniform, and agrees perfectly with a good many skins from
Gumanaand Puerto Oabello. Two specimens from Tobago are also in no way ditferent.
It is certainly wrong to treat D. ineruloidfs as a subspecies of r>. inenda (Lciit.).
The latter is readily distinguished from all South American species of Dendrocincla
by its deep chestnut upjier wing- and lower tail-coverts and bright orange axillaries.
D. meruloidex, on the other hand, agrees in both respects with J ). jihaeochroa {v{\\\ch.
occurs together with ]>. mcritla at different localities, e.g. on the Orinoco), but
differs in its more ciunamnmeous-brown upjier parts, more rusty under-snrface, and
in the throat being uniform with the breast (instead of buffv).
88. Dendrornis susurrans susurrans (.lard.).
DciHlroi'nlapies s«s//mi«s Jardine, Ann. .Vu(/. iVii/. /fiat. xix. (1847) p. 81 [Tobago].
Nasica albhqiiaiiui Lafi'osnayo, liir. Ziinl. 1852. p. 4(55 [loc. ign.].
Deiiiliiiriiis mnxdhi-iimx Dalmas, M^iii. Soi: Zoul. France xiii. (190II) p. 140 [TriniJad].
Fourteen specimens (adult and young) from (Japaro, April ; 1 S ad.
Ohaguaramas, January ; 1 c? ad., Pointe Gourde, January ; 1 ¥ jr., Seelet, April ;
4 (?(?, Laventille, April and May ; 1 S, Valencia, March.
* I compared two of the typical specimens, cullectcd by M. l!eaupeitlniy.s, in the I'.iris Museum.
They are both labelled •' Ciimaiiu," ami as lieaupeitluiys never collected anywhere else, tills locality must
be regarded as the typical one.
( o" )
On corapariiijj this scries with fifteen specimens IVdiii Tobago I tail to see .any
ilifierence, and cannot, therefore, admit the validity oi 1>. co/isol/rim/s Ualni.
The form fonnd on the opposite mainland, near (Inmana, Matnrin, and in the
Orinoco delta ((tuanoco), however, is ajiparently dillerent and worthy of recognition.
Ir ditVers in its more fnlvons nnder-jiarts, tliis lieing especially nutiecahle on tlie
throat, which is distinctly bntV (instead of wiiitish), and more rnfesient back. This
subspecies has to stand as A susurransjardinei Dalm.
At my reipiest, Mr. Oberholser examined t\\o types of yasicK ulbisquama
in the Lafresnaye collection, and found them to be identical with tlie white-throated
Tobago form. Jlost probably, Lafresnaye's types came from Trinidail.
80. Picolaptes alboliueatus (Lafr.).
Vewlri'Culaj.tm (illuJhiriilim Lafresnaye, lifi: Znnl. ItSiCi. p. 208 [" Colonihie on Mexique.''— We
accept liiiijiild !Lt tlie typical loc.ility].
2 S (S and 1 ? from Caparo, April : exactly agreeing with three adults from
Cumauii.
Bogota skins and several examples from West Colombia differ slightly in
having the back rather more rnfescent and the under-jiarts darker brown, with the
whitish longitudinal streaks somewhat broader. Two s]iecimens from Valencia
appear to be identical with those from Bogota.
A much larger series is necessary to decide whether there are any geographical
races of this .sjiecies or not.
00. Thamnophilus major semifasciatus (Cab.).
[Thamnophihis major Viuillot, .V<///i'. Did. iii. (181IJ) p. .'513 (ex Azara : Paraguay).]
Diallactes neiiii/iisiititiis Cabanis, J on in. f. Oniith. I87'2. p. 234 ['' Para, Guiana and Venezuela "].
Thamniiplidiis iilbhris.<ux Ridgway, Prnr, C.S. .!/«<. xiv. (1891) p. 481 [Trinidad].
6 S ad. and o ? ? from Cajiaro, April.
In the markings of the tail they agree with a series from Venezuela (Cnmana
and E. Orinoco), liritish Guiana, Cayenne, and Para. Typical 7'. m. imijor \ie\\\.
of Paraguay, Mattogrosso and Bahia has much more white in the tail, broader white
edges to the primaries and, as a rale, a smaller bill.
01. Thamnophilus doliatus (Linn.).
Liiiiiiis ihiVHitun Linnaeus, Mim. Ail. Fmli . ii. I'lodr. p. 12 (17t'i4) [Inc. ij^n.— We substitute Surinam.
as the typical locrdity].
Thamnoj)liilus (loliiilu* fr'iterrulun IJerlepsch Ji Hartert, Xov. yinnl. ix. (10o2) p. 70 [Orinoco River].
25 (J ad. and jnv., IT ? ? from Caparo, April and May; and Seelet, April ;
1 6 ad., 1 ?, Pointe Gourde, January ; 1 J ad. from Laventille, December.
With a series of seventy adult males liefore me, I can no longer distinguish the
form named T. d. J'niterculus by Berlejjsch and Hartert. In some cases, the
specimens from Cayenne, Surinam and Trinidad have rather broader bars on
the lower surface tiian those from Venezuela, but so many of them are ipiite
indistinguishable from the latter that 1 cannot make any separation. Birds from
Tobago agree better with the narniwly-barrcd form of the mainland.
(1^1 )
02. ThamnophiUis canadensis canadensis (Linn.).
Lniiiiis miKirli'iiiiK Linnaeus, Si/mL XuI. xii. 1. p. I'M. desrr. f [l")')!'. : ex Rrisson, '^ Cumnhi" :
errore ! — Wesulistitute Ciii/i ii/ir as typical locality].
T/iamuophi/us tn,i',l,ilis Ridgw.ay, I'm: I'.S. .Uiis. xiv. (IW91) i>. 4«I [Triniilad].
T. firi-huliis auct.
9. 3 6 and 1 ? from Soclet, April ; •") S ad., 1 S jr. and 1 ? from Pointe
Gourde, January ; 1 c? ad. from Chagnararaas, .January ; and 1 J jr. from
Laventille, Manb.
Besides these specimens, I have examined a series of 40 adults from ('aycnne,
British Guiana, Surinam, (!nmana, the Orinoco and the Rio Branco (Natterer coll.),
and I fail to see any constant differences between them. Dark and pale-backed
specimens are to be found in Trinidad as well as on the mainland, and some of the
(typical) Cayenne skins are even darker on the upperside than those from Trinidad.
I can, therefore, no longer maintain T. trinitatis as different.
93. Dysithammis affinis andrei n. suhsp.
Di/sithamniis meiiliilis spoiliniiohis (nee D. xpiuliniintns Salv. & GoJni. I) flliapman. Bull. Amer. Mus.
vi. (1894) p. an [Trinidad].
4 c? ad. and 3 ? ? from Claparo, April.
(?. Differs from />. n. afflids oi Mattogrosso only in having the under tail-
coverts white.
?. Quite different from that of /A a. tijfniis and D. a. sjiodioiiotn.'i by having the
back olive-grey (not cinnamomeous or warm olive-brown) and the under-surface
much paler without any cinnamomeous tinge on sides or foreneck, only the flanks
and crissnm being pale olive yellowish.
Type in Mus. Tring : ? ad., Cajiaro, Trinidad, April 12, 19(12. E. Andre coll.
It is strange that the cj c? of the Trinidad form are much more like D. a. ajjuiis
of Mattogrosso than B. a. spodionotiis from the Roraima Mountains. Comparing
three males of each, I can hardly perceive any differences between them, but the
females are strikingly different. The distinguishing characters of the three
subspecies may be summarised as follow :
(rt) D. a/fniis afii/ii.i Pelz.— Distribution : MidtiKj rosso, ('. Brazil : Villa Jlarin
(Natterer) ; Chaj)ada (Smith).
5 ad. Ujiper parts puro schistacenus gri'y, pileum rather darker, rump slightly
washed with pale greenish. Throat and middle of the breast and ab<loni('n white,
sides and foreneck washed with pale grey, flanks and under tail-coverts distinctly
tinged with pale brownish or greenish. Wing, 61 ; tail, 39 mm.
? ad. Ijack warm olive-brown : upjtcr wing-coverts and outer weli of the
remiges light rufeseent brown. Throat and middle of the lielly pure white, chest
and sides pale cinnamomeous brown. Wing, 01 ; tail, 39 mm.
{b) D. affinis andrei Hellm. — Distribution : Trinidad.
(J ad. Exactly like the S of the i>. a. uj/iiti.i, but the under tail-coverts [)nre
white, only the flanks being faintly shaded with greenish. Wing, (jl — 03; tail,
40—43 mm.
? ad. Back olive-grey, passing into pale greenish on the rump. Upper wing-
coverts and outer webs of the remiges pale olive brownish. Throat and middle of
( :i2 )
tlie belly imre wliite, fcircneck soavce]_v shaded with biifty ; tiauk.s and niulev tail-
coverts olive-yellow with a lirownish hue. Wing, On — G:J ; tail, 39 — 41 ram.
(<:•) I). (i//i/'/s s/iiii/io/'oh/s i>a\v. ^ (iodm. nisfi-ibutimi : Il(>r<i.iniii Jit., British
(iiiiana.
<i ad. Ditt'ors from a and li by the darker, more slaty j:;rey nppor parts without
any trace of greenish on the nimi). The grey colour below is al.so rather darker and
more extended, and the flanks are always washed with brownish. Wing, 05 — 08 ;
tail, 40 mm.
? ail. Exactly like that of D. u. ii/fi/i/,i, perha]is a shade paliT, more rnfcsceut
olive-browu on the lower parts. Tail distinctly longer. Wing, 0"2 — 04 ; tail,
44 — 47 mm.
These three snlispecies form a natnral grou]i, and are quite different from
/). i/ii'iitalis me/iti'I'x (Teniui.j and /). m. olicaci'iis (Tsch.) which have the belly
in both sexes sulphur-yellow without any white, ('fr. Journ. f. Ornith. 19u5,
p. 14 li".
D. u. (uidrci is evidently confined to the island of Trinidad, since two specimens
from Tobago (? ?) and several adults from ('umana Itelong to the yellow-bellied
/'. III. Illflltulis.
'.»4. Myrmotherula axillaris (Vieill.).
Mijintdlhera a.ciUaiis Vieillot, Xotir. Did. xii. p. 113 (1817) [ex ■' La Guayane," sc. Cayenne].
;i (S ad. from ( 'aparo, April, agreeing in every resjiei't with typical ('ayenne
specimens.
'•i-">. Ramphocaenus melanurus trinitatis Less.
[liu mphocinms meUinuncs Tieillot, Noiir. Dirt. xxix. (181'.)), p. 0 (" Bri'sil "— coll. Delalande :
sc. Hio de Janeiro) ].
Ramphnraemts triuitatis Lesson, Tin: Znol. 1839. p. 42 (" in insula Irinihilis ").
S and ? from Caparo, April and March 1902.
The specimens agree with a good series from (Jnmaua and the Orinoco and ( 'aura
Rivers in having the under-parts strongly washed with ochraceous buff (Hidgway,
NomcDcl. v. fig. 10). The clieeks, ear-coverts and sides of the neck are uniform
deep ochraceous (liidgway v. fig. 7), and the back dull smoky brown in decided
contrast to the dull rufous brown pileum. Two specimens from Cumamlhave only a
slight ochreous buffy wash on the innermost sides of the belly, thereliy appro:iching
Jt. m. ((Ihiceidii.s iScl. from I'ritish (Juiana and Surinam ; but the sides nf the head
:ire deep ochraceous, as in It. in. trinitatis.
11. lit. )iicl(ini<rus of East Brazil (S. Paulo to Para) resembles the brightly
coloured specimens of R. m. trinitatis on the under parts ; but the ear-coverts are
much paler buff, and the back Is warm rufescent brown, scarcely different from the
colour of the crown.
It is extremely doubtful wlu^ther Lesson's name can be accepted for the
northern form of R. niflaniaus. Although Trinidad is given as the typical locality,
the description does not fit very well, and the terms " corpore infra niveo, lateribns
griseis," apply much better to A', m. uthircntris. Perhaps Lesson described a
specimen of the latter race with a wrong locality, lietore giving a new name,
however, the type of trinitatis should lie examined.
( 33 )
90. Myrmeciza longipes lougipes (Sw.).
Dri/iim/iliihi hiii(/i/)cs Swainson, Ztmhxj. Jniirii. ii. (July I^i'25). p. 152 [" somo part of Brazil (I was
told from Rio ile Janeiro) " — errore ! We subititute Triniihnl as ilie typical locality].
Myriiieciza siniiiiniiiii Berlepseb, IhU 1HK8. p. liiO [Puerto Cabello and Trinid.ad].
Myriitei'ha loiif//pc>i uVi/i-ndrix Cbapnian, .I"/' x. (18*'.'i), p. 1^4.3, and Itiill. Amcr. J\[ii>i. vi. (1804),
p. 51 [Trinidad].
2 c?c? and 1 ? , Chagnaraiuas, Jannarv ; one pair from Laventille, April ami
Deccmlier ; one c^ ad., (laparo, April.
The series is aliscilntidy identical witli another from Cnmani'iand Puerto C'aliello,
Venezuela. I fully aijree with Mr. Ohapman that the term longipes Sw. should
be ae.cepted for the bird named ^f. tiwaimoiii by P>erlo[tseh, but I cannot follow him
in separating the Triuidad form nnder a new name. One S ad. from Pauama (coll.
Harris) differs from the latter in having the sides of the chest ashy grey (instead of
l)alc fulvous brown like the flanks), as poflited out by Chapman. In Swaiuson's
descrijjfion, however, no mention is made of this colour ; hence his name becomes
strictly applicable to the Triuidad race. Moreover, ^[. siminsoiii Pierlepsch being
based upon specimens fnun Puerto Cabello !ind Trinidad, antedates in any case
(Jhai)man's later designation, while the birds from ('artagena and Panama, if really
se])arable, must have a new name.
In the Orinoco valley and in British Guiana a very distinct subspecies takes its
|ilac'e. This has been named ^f. I. (/riscijtcctKS Berl. & Hart., and is easily known
by a broad grey band across the chest just below the black throat.
'.IT. Sclateria* iiaevia (Gm.).
Siltd Mipriii Gmelin, .S.(/s(. Nal. i. 1 (1788) p. 4t'J [ex Edw.ards : Surinam],
4 t?c?, 1 ?, from ('aparo, March and April ; one ?, (!haguaramas, January.
Identical with a series from Cayenne.
'••8. Formicarius hoifmanni saturatus Uidgw.
[Mi/niiiiniif: tli'll'miiinii Cab.anis, Jcnni. Uniilh. ISGl. p. '^f> (C'osita Rica).]
Fiiniiii'iiriui futliiniliii Ridgway, Fm : U. S. Mas. xvi. (I8'.)4), p. ('i77 [type ex Trinidad].
2 S d from ('ajiaro, April; I S from Vali'iiciii, March; and 1 '? iVom Aripo,
2000 ft., May.
The Triuidad specimens agree perfectly widl with others from Curaana, Tocuyo,
V'^enezuela, and from the Cauca valley in W. Colombia. F. h. saturatus is easily
recognisable from /'. //. ciixxulis ( 'ab. (ex Cayenne and Brit. Guiana) by its
paler, more olive-brown colour above, smaller (u- (jlisolefe white loral siuit, darker
grey under jiarts, and paler lower tail-coverts.
0!'. Glaucis hirsuta (Linn.).
Trochihts h'muttis Gmelin Sifsi. Nul. i. 1 (1788), p. 4'.i0 ['' Brasilia " — ex Marcgrave].
Two pairs from Caparo, April, and three si)ecimens from Laveutille, November.
They agree with a scries from Puerld Cabello, the Caura River, and Tobago.
Topotypical East Brazilian skins are not available for comparison. Accordiui,' to
Phelps {Ault 1897, p. 30G) the species occurs also in the state of ('umana,.
* Ulim Httt'rociiemitt (preoccupied).
( 34 )
loo. Phoethornis longuemareus (Less.).
Trnchihis Jmiguemareus Lesson, Ilisl. Xat. Tinrh;!.. p. If), pi. 2 (1832— Cayenne).
Three c?(? fi'oin Cai)aro, Ajjril. Tlic Ti-injr Miisoum lias also two skins (r? ?)
collected by Dr. Percy Kendall at ('ajiaro.
Tliev differ iVom a series of ( 'ayenue skins (coll. Cliorrie) liy liaving the
hiackish subterniinal bar on the tail-feathers less distinct and the njijier lail-eoverts
not so broadly margined with rufous. The blackish ^nlar spot, too, seems to be
larger. Tlie.se slight differences, however, are not qnite constant.
P. lonqueimiri'us is hitherto only known from Cayi'nne, British (Jniaiia
(Schombnrgk), and Trinidad.
101. Phoethornis guy guy (Less.).
Trneliihix ijiiji Lesson, Hist. Xal. Tmckilid. p. ll'.l. tab. 44 [18.32: no locality gircn. Wc substitute
Ti-iii/ihtfl].
3 c?(? from Caparo, March and April ; 1 J ad. from Aripo, 2000 ft., May.
Specimens from ( 'iuiian;i are practically identical. All have the base of the
tail and the njiper tail-coverts bronze green, withont any bluish gloss. This form
is confined to Trinidad and N.E. Venezuela (state of Cnmaua).
102. Florisuga mellivora (Tjinn.).
Trtn-hiliiti ittfUirnri/^i Linnaeus, Si/\t. Xuf. x. p. 121 (17.'i8 — ex Edwards: " in Indiis " : sc. Snr/iKtni
ex Edwards).
J? ad. frmn ( 'ap;iro, April : 1 d fmm Valencia, March: and 1 cJ imm. from
Aripo, KiiiO ft.. May.
Though occurring in Tobago, the species has not yet been found on the opposite
Venezuelan coast.
I'l;'. Agyrtria chiouopectus chionopectus (Gould).
T/iiitiiitalidH rliiniiiijirrtus fiould, Monaf/r. Ti-iii/iil. V. (l.SiJ'.l), tab. 2'.i:i [Ti'inidad].
A large series from (laparo, March and Ajiril, and Laveutille, March. It occurs
alsd on the opposite Venezuelan coast iu the state of Cumana, where it has been
collected by Mr. Phelps (Aid 1807. p. 300).
In British (Tiiiana and on the ('aura liiver it is represented by a smaller Ibini
with a less coppery tail, ,1. c. /r/ii/c/i/i (Bone.).
Iii4. Saucerottea erythronotos erythronotos (Less.).
Oriiiionyn eri/lhrniiolos Le<son, Jlixl. Xiil. Oi.i. .1/"h<7««, pp. .\x.\ii., 180. tab. Ixi. [1829: "Bresil" —
crrore ! We substitute TniihluJ].
Twelve specimens of both se.xes from (!aparo, April ; 1 S from Ohaguaramas,
January ; 1 i from Laventille, November ; and 1 i from Arijio, 2000 ft., May.
In most specimens the under tail-coverts are dark coppery with ciunamou
edges, but in one S from C'aparo they are dark violet edged with whitish, and in
another from Laventille ])ale greyish with whitish margins.
The Tobago form, S. <'. wells/ Bone, is rather larger, with the tail luori' deeply
i'orked, and with the back and crown darker, purer green. On the Venezuelan
mainland it is represented by another close ally, S. e.fi'liciuc. ((Jf. the excellent
remarks liy Count Dalmas in Miin. Soc. Zool. France xiii. 1900. pp. 142-3.)
( 35 )
10."). Chlorestes coeruleus (ViCill.).
Trorhilit^ nu'riilfiis Vieillot, A^utir. lUrl. vii. (1S17), p. .^*U [Cayenne]
30 d <i and 3 ?? from ( ';i,paro, Miircli and April, and 1 cJ ad. from
(jbaguaramas, .Tannary.
The males ai^n'i' perfectly with a jjood series from Cayenne, British Guiana,
and the Orinoco region. The species has not yet been taken in the ( 'nmana region.
106. Chlorostilbou caribaeiis (Lawr.).
Chlunisl'illiiii ciiiihiniis Lawrence, Ann. N. Y. Li/c. x. (1871 ), Ji. lo [Cura^'ao].
2 c?c? from Laventille, May. We have also 2 cJc?, collected in March 1S07 at
Tacarigua by Dr. Percy Kendall.
These four specimens agree with several ollicrs from Curacao in having tlie
throat and breast pure glittering green. The birds from Curaanu liave been
separated by Simon & Dalraas as ('. c. Irxaoni, but their chief cluiracter, the bluish
hue on the throat, occurs sometimes also in specimens from ('iiracao, and I doubt
the possibility to distinguish them.
It IT. Colibri delphinae (Less.).
()riiismii<i th'liiliiniic Le^scm, Rev. Znnl. IS.TO. p. 44 [loc. ign. — We substitute Trtiiidinl].
Eight iulidts from Aripo, IC.Dil (o 2(Hlil ft., ftfay, fully agreeing with a good
series from ( 'ninana and Columbia ( l'>ogot;i coll.).
108. Anthracothorax gramineus (Gm.).
Trnchihut i/ra III /liens Gmelin, Si/sl. Niil. i. 1 (1788), p. 488 [based on the " Haussecol vert " of Buffon.
No locality given ; we substitute Ciiiji iinf'\.
Ln-iiijiin'iih iji'diiiincus auct.
1 i ad., Seelet, April S ; 1 (J) ad., Caroni Swamp, February In ; 1 ?, (Jaroni,
March 22, l'.i(i2.
The Trinidad birds agree perfectly with a series from Cayenne and Surinam.
I'lo. Anthracothorax nigricollis fVieill.).
Tnichilm niiirlciiUin Vieillot, Nmir. Did. vii. (1817), p. 34'.l (" Biv.sil ").
Lniiipoi'Hin iiiijrivuJIh auct.
1 (? ad., Valencia, March; 2 ??, Laventille, November 8, March 11);
1 cJ juv., Seelet, April ; 1 ?, Ari|io, IsOO ft., May; 26 adults and young from
Caparo, April. We have also 2 cJcf, 1 ?,froin ('ai>aro valley, and 2 i 3 from
Tacarigua, February, collected by Dr. Percy Hendall.
The series agrees e.xactly with other examples from Cumana and S. Brazil.
The female oi A. niyricolUs dilfers from that of the preceding species by having
a broad velvety black stripe along the middle line of the underside from the chin to
the anal region. In .1. (ivaminCKs there is only a dull blackish stripe to be seen .m
the middle of the throat and foreneck.
( 30 )
llii. Chrysolampis mosquitus (Linn.).
TnichihiK mii.'niiiiliix Linnaeus, Si/al. X,il x. p 120 [1758: ex " Indiia " — urrore ! Wo .substitute
Siiriiiaiii as typical locality].
A larjje series of lioth sexes from Ciiparo, April ; Lavi'ntilie, Deei'iulier ; T'ninte
Gonrde, Janiian- ; antl .Seolet, April.
111. Polytmus thaumantias chrysobronchos (Shaw).
[Triirhilun IhiiiiuKiiitiiis Linnaeus, Si/i.1. A'<il. xii. 1 (ITliii), p. I'.Mt ^e.\ Urissrm : " Bru^il ").]
Triichiliis rliri/xcibi-inirhos Shaw, Gen. Znnl. 8. 1 (1812), p. 287 [ba.sed on Audebert .t Vieillot, Oh.
Doles i. tab. 41 — "Guiana," sc. Cayenne].
1 (c?) ad. and 1 ( ? ) from Caroni Swamp, February. Thoy agree with speeiiuens
from British Gniana and ( 'ayenue.
This species has not. yet been recorded from the Paria peniiisnla.
11-. Anthoscaenus lougirostris longirostris (Andeb. & Vieill.).
TrncUhm longirostris Audebert & Vieillot, 0«. /)„;■. i. (1802), p. 128. tab. 59 [" Indes occidentales."
AVe substitute Tritiitlii<i\
1 6 ad. from < 'aparo, Ajiril 13, 1902, agreeing with a specimen I'rom (Uimana.
113. Lophornis ornatus (Rodd.).
Trnchilus ornatus Bodd.aert, Tnlil. PL cut. p. .'j'.l [178:) : ex Daubenton, /'/. mi. 040. fig. IS, Cayenne].
A fine series of eleven si)ecimcus from Laventille, October and November. The
S ad. are not different from other.s collected by Mr. Andre on the ( 'anra Uiver in
Veneznela. Typical Cayenne skins are not available for comparison.
114. Chaetxira zouaris albicincta (Cab.).
{IlirmuJo zimaris Shaw, Cimelia Phys. (I7'JG) p. 100, tab. 55. (No locality).]
Hemiprortie iiltiicim-ta CabanU, Joiirii.f. Ornith. l.siVi. p. 1G5 (part : Guiana).
Acanthylis coUuris L^otaud, Ois. Trinlihiil, p. 83 (Trinidad).
Two specimens, luarlced S and J jnv., collected by Mr. Carr at Cliagnanas,
September 24, 1804.
Botli are young and have the white neck-band still incomplete, bnt agree with
specimens from British Guiana in dimensions. The wings measure 177, 187, the
tail 01, ()'.) mm. The length of the wing in skins from Guiana (Annai & Horairaa)
varies from 170 to 18G, that of the tail from (iU to 05 mm. According to Leotand,
the sj)ecios does not bre( d on the island, but Carr's specimens seem to indicate that
such may be the ease. Anyhow, further observations are required to establish its
claims as a breeding liird.
117p. Chaetura sp. (near t'. iicUnjint (L.)).
Mr. E. Andre sent a female, collected March 20, lOii;!, near Valencia, of a
species allied to ('. jn'layica, but evidently distinct. 1 refrain, however, lor the
])resont from naming it, since it may be a migrant from some more ncn-thern locality,
in which case it would be difiicult for future writers to identify the species.
I 37 )
It agrees with ('. pelfiyica and C. fan.ri in general form and size, Imt ilill'ei's
very conspicnously in having no trace of the whitish throat-patch, and thi' pih'nni
and l);u'k of a ninch darker, metallic lilack colonr.
11 II. Chaetura cinereiventris lawrencei Ridgw.
\Chaelura chiereivcnl ris Scliter, Cat. Aiiier. Birds (1802) p. 2.S3 (ex Wiud & Burniuisler. -naliiii).]
Clmeltmi lawreiirri Eidgway, /'nw. C/..S. Nat. .\lif<. xvi. (1894) p. ^'^ (Grenada, W. I.).
Aciiiithijlh ii.r.i)urfi (nee Vioillot !) Leotaud, Ois. Tr'mhhul p. 84.
1 9 ad., collected liy Mr. Carr, 'Z'l. iii. LSili, without exact locality.
Tliis specimen as well as a ¥ I'roiu Toiiago are practically idenlical wilh a
(J from Grenada. All have the upper tail-eoverts Idackish with a slight steel-green
gloss, only a very narrow edge along the inner web being pale greyish ; tlic lower
surface is pure schistaceous, in strong contrast to the steel-black under tail-coverts.
C. c. quianeitsis Hart, of which form there are three examples from British
Guiana in the British Museum, differs in having the upper tail-coverts or at least
tiieir inner webs light schistaceous grey like the rump.
According to Leotaud, the species is sedentary on the island.
117. Chaetura brachyura (Jard.).
.iruiitlii/lix tiriii-Iiijiira Jardine, Aim. .!/.(</. .V.i/. Ilisl. xviii. (18iG) p. 120 (Tobago).
Araiil/ii/lis jiiiliiiiinis (nee Temminck !) Leotaud, Oh. Tyinidml, p. 8G.
Chiirtitm p'llium auct. (nee Temminck !).
1 i ad. from Laveutille, 111. v. 1003.— E. Andre coll.
It agrees perfectly with a typical specimen from Tobago. A series ot twenty
examj)les from Cayenne and Surinam is also absolutely identical.
This is the bird commonly called C. poUura, but Temminck's name certainly
does not refer to the species in question. Ci/pselua poliouii.is Temminck * is
exclusively based on D'Aubenton's plate 720, fig. 2. Both the figure and the
description of the " Hirondelle brune acutipenne de la Louisiane" suit much better
the common North American Chaetura pelagica (Linn.); and, moreover, in the
letterpress Moutbeillard expressly states : " L'individu dont il est ici question a 6te
cnvoye de la Loiiisiani' par M. Lelieau." Tiience there is not the slightest donbt
that ('j/psdi(s poliourias must be regarded as a pure synonym of C. pclfujicn, which
is the only species found in Louisiana. Tlie next available name for the South
American s])ecies is A. hradnjHra.
118. Panyptila cayennensis (Gm.).
Hindnl') cni/cnnfiisis Gmc\m, Sy.it. Mtit. 1. ii. (1788) p. 11124 [based on l)3,uhcntoii, PL nil. 1'2'>
fig. 'J.—Oii/cniic].
Although Ghapman found the species common at La Brea, Mr. Andre's
collectors never met with it.
119. Nyctibius griseus griseus (Gm.).
Ciipriinulgus ijrisctis Gmelin, Syst. .\al. 1. ii. (IT^H) p. 102'.l [ex BufTon. — '' Cayenna ").
Nijctibius jamaiccnsis griseus Hartert, Tierreich, Lief. 1. (18'J7) p. Ui.
1 S from Chaguanas, May 17, 1894, collected by Mr. t'arr, received from
Mr. E. Andre ; and 1 ? from Savannah Grande, February 15, 18U7, collected by
Mr. Percy Rendall.
* I'nM. Viiili. ri. col. p. 57.
( 38 )
(J : AVing, 2o3 ; tail, 178 ; ? : wiiiir, 240 ; luil, ICT mm.
One adult from Buliia measures: wing, 20(J ; tail, ISO. One from ('ayenue :
wing, 265 ; tail, 180 ; and another from British Guiana: wing, 2o9 ; tail, 185 mm.
Tlins the Trinidad l)irds belong to the smaller .southern form. Several skins
from Jamaica are niueli larger : wing, 2!I0 — lilO ; tail, 210 — 225 mm., and
aj)iiarentlv sejiaralile as a subs|)ecies. Its proper name is S. griscus jamaiccnsis
(Gm.)
It is strange that all authors nsed to call the species -A', jamaiccnxis, liut as
C. griseus stands first in (inielin's work, it must be adojited as the specific name.
120. Lurocalis semitorquatus semitorquatus (<im.).
Ci'iirimulgus semitorqiiatiis GiiKYm, Sijil. Xc.t. I. ii. (178H) p. 1031 [based ou Daubciiton, /V. (•«/.
734.— "Caycniia"].
1 (J and 2 ¥ ? from Cluiguanas, July, May and Se|iteniber, collected by
Mr. (."arr, received from l\Ir. E. Andre.
These specimens agree in coloration with a Cayenne skin, but have rather
shorter tails — 77 to 84 instead of 92 mm. There is, however, no difference iu the
length of the wing.
Skins from S.E. Brazil are much larger, and I'orui an easily recognisable race,
which has to be called L. xi'mi/orij'/afu-s iiattereri (Temm.).
121. Nyctidromus albicoUis albicoUis ((Jni).
Cnprimulgwi albicoUis Gmeliii, Sijsl. Nat. 1. ii. (1788) p. 103U (ox Latham.— C'fl,'/fH«<').
3 (?<? and 1 ? from Laventille, November and April ; 1 ? from ( 'haguanas,
January 20, 1897, collected by Mr. ("arr. Agreeing in colour and size with others
from Cayenne.
122. Steatornis caripensis Humb.
Steatontis mrijiriinis Humboldt, Hull. ."<,„■. riiilnm. Pans (:'.) xvii. (1810) p. -I'X, [Caripi-, Vi-iiczuela"].
3 ii, 2 ? ? and 1 witliont indication of the sex from Aripo, 2400 ft.,
collected May 13—15, 1903. " Iris black."
They agree with some specimens I'mm Venezuela ('iMiH'<[nerys coll.).
123. Chlorouerpes rubiginosus rubiginosus (Sw.).
Piciis ruhii/iiinsii.s SwainsoD, Zmil. llluslr. i. (1S2II-1) I'l. 14 ["Spanish Maiu." — Wc substitute
Cumana a.s typical locality],
12 (JJand ??,(;aiiaro, Aju-il ; 1 S, Valencia, JIarcli ; 1 S, Seelet, March;
and S ? , Chaguunas, June.
Some of them are strikingly small, but others ipiite as large as birds from
Cnmanii. In the Andes of Merida and near Bogota a ranch larger form occurs,
which, liiiwever, is hardly different in enloratiun finm tvpiciil iuhi(]i)io.%iiii.
i ad., Los I'almales, ('nni,Tn:'i . Wing, lOs ; tail, tis ; bill, 22 mm.
4 c? ad., Cajiaro, Trinidad . . „ 99_l0(; ; „ (JO— 57 ; „ 21— 22i „
6 ad., Valencia, „ . „ 108 ; „ (i8 ; „ 22
5 ad., Seelet, „ . „ 104 ; „ 04 ; „ 21
Unfortunately, 1 have no s])ecimeiis from Tobago to com])are; from the remarks
of Mr. Hargitt* and ( 'onnt Dalmas f it would seem tiiat they are diiferent from
those of Trinidad.
• Cat. liirdt Bril. Mu»., xviii. p. 88. f Mem. ,Soi\ Ziutl. l-'rniiir, xiii. (I'JOO) p. HI.
( 39 )
124. Veniliornis kirkii kirkii (Malli.j.
r. {Oihn-;jwns) l.nlii Malherbc, lire. Zn„l . 184.',. p. 400 (Tobago).
1 i, ( 'liaj^nanas, May; 1 S ad., Poiiite Gonrdc, Jaimarv ; 1 6 a.il., .Vri|ii>,
21.11111 Ct., May; 1 S jnv., and 1 ■»' ail., Caparo, April; ami I ¥ ad., liaveiitiilr,
Noveiiibcr.
Tills series a,y;roes, in ucduratiou and si/.e, witli a fair luimber of skins IVoni
Tolia,i,'ci, while 3 ? ? from the Verieziielau coast (Caripe and 8au Esteban) are so
mnch smaller that I do not iiesilate to separate tlieiu as
Veuiliornis kirkii coutinentalis nov. subs[i.
Type in Mus. Trinir, ? ad., Caripe near Ciiinana, Venezuela, Jannary Is',t4. A.
Mocipierys coll.. No. 243 : wing, 83 ; tail GO ; bill, ISJ mm.
Auotiier ? from the same locality (Mus. Tring) : wing, 82 ; tail, o(j ;
bill, 19 mm.
1 ?, IS. Estelian, near Puerto ('abello (coll. Goeriug. — Mus. Brit.) : wing, 81 ;
tail, M ; bill, 18 mm.
Measurements of T. kirkii kirkii (Malb.).
~ SS ad. from Tobago . . Wing, 89—00 ; tail, .38—03 ; bill, 22 — 23J, mm.
3?? ad. „ ,, . . „ 88-80; „ 50—61; „ 2U— 23 „
ASS-m\. „ Trinidad. . „ 91—93; „ 00-03; „ 23—24 „
2 ? ? ad. „ „ . . „ 01 ; „ 00-61 ; „ 20,1—22 „
125. Celeus elegaus leotaudi nom. nov.
One ? ad., Valencia, March; one ? ad., (iaparo, May ; ? juv. from Chaguanas,
.July. Besides these, we have another ? ad., collected Febrnary 11, 1807, at
Savannah Grande, by Dr. Percy Rendall.
The Trinidad form has erroneonsly been called C. elegants by all iirevious
authors. Fictts cliyan.^ Mfdl., as well as /'. J'n.'iroftilcm Bodd. and V. ciniiamomcus
Gm., are based on Daubenton's PL nil. 524, " Pic janue tachete de Ca>/eime.''
Four adult specimens (3 (?c?, 1 ? ), collected in Cayenne by Mr. G. K. Cherrie,
are very diflTereiit from our Trinidad series, being much larger and darker thronghoiit,
with the back dislinctly spotted. In fact, they agree very closely with the characters
of ('. rcicbenbacJd (as given in the Cat. B.) to which Mr. Hargitt actually refened
several examples from Cayenne. It is the more surprising to iind the name clegans
used as the specific term of another s])ecies which does not at all occur in Cayenne ! !
It seems also that two distinct forms have been united hitherto under the name
C. reichrnhnchi, but to which of them the latter is strictly ajiidicable, cannot be
decided with certainty from Mallierbe's description. The distinguishing cliaracters
of the three races are as follows :
(a) Celi'us cleyans clcgaiix (P. L. S. Milll.).
Picus elegam P. L. S. Midler, yatiir.v/sf. Siipi>l. (ITTO) p. 02 [based on
Danbenton, f'l. cnl. 524 — Cayenne].
I'icmfusco /ulcus Boddaert, Tatd. I 'I. ml. (17s3; p. 30 [liased on the same].
J'icus ciiiiiamomeuii Gmelin, Sgst. Sat. 1. i. (1788) p. 428 [based on the same].
Top of the head and crest ochraceons-bulf (Hidgw. V. 10); liack and upper
wing-coverts deep chestnut brown (rather darker than Kidgw., IV, (ig. 0), willi
( 40 )
very distinct pale yeliowisli spots ; iiiiderparts deep chestnut brown, still darker
than the back. Win;; Hil — 17(1; l.iil UKi— 117, iiill, L".i— ;il mm.
Hab. Cai/etiiie, A]ipron'a,i;uc (('lierrii' roll.).
(Ji) Celeiix rlfqans n-irhciihnrhi (l\l!illi.).
CelenpicHS rekhenbnrlii Mulin'rlM', MoiiKiir. I'iri,!. ii. (lSii'2) ]i. -'><, tab. ."iii,
figs. 4, .5 [•' le Bri^sil, hi Coloiubic."'- The hitler hicalilv is dmibtless erroneous].
Top of tlie iiead and crest cinnanion ; back and iipi)er wing-coverts deep
cinniinion-rufons witliont any pale markings, only the greater series of the latter
sometimes with obsolete jiaie cinnamon cross-lines; nnilerparts chest nnt-rnfons.
Wing lOU— l(i«, tail, 102— ll.">, bill -JS— :iO mm.
llab. Surinam, Paramaribo (Chiinkoo coll.— Mns. Tring) ; Hritixk Guiana,
]{. Carimaug, ( 'amacusa, (^inonja, etc. (II. Whitely coll.); S.i:. Venezuela,
Guaiioco in the (_lrinoco dilla (.\iidre coll.— Mns. Tring).
Specimens from Guanoco and Surinam are exactly like those from British
Guiana, except the head and crest being slightly paler, more ochraceons-cinuamon.
Whether this form is really entitled to the name C. irirjicnbaclii, can be ascertained
only by an examination of MaUierbe's types. I susjiect that the birds obtained by
Natterer at I'.arra do liio Negro and on the Kio Branco will prove to belong to the
present subspecies, but I have not yet had an ojiportunity to compare them.
((■■) Cclrii.i dcgans h'otauili nom. nov.
Celcus elcgans (nee Mirller) llargill, Cat. Birds Brit. Mns. xviii. (Is'.ni) \>. 426.
Top of the head and crest pale cinnamon ; back an<l upper wing-coverts bright
cinnamon-rufous, the former as well as the smaller wing-coverts with indistinct
pale yellowish spots; underjiarts bright cinnamon-rufous. Wing 139-142, tail
Sf)— 08, bill 20—28 mm.
Type of subsi)ecies: " ?" ad., ^■ah■nc•ia, Trinidad, March 20, l'.MJ3. E. Andre
coll. — l\[us. Tring.
I lab. Triiiidnd.
This form is evidently confined to the island of Trinidad. The specimens
collected by Goeriug at Guarauuo in the jjrovince of N. Andalucia, Venezuela, and
those obtained by Mr. Andre's collectors in the Orinoco delta, belong to the large
race found in British Guiana and Surinam.
I take great pleasure in naming this bird after Dr. Leotaud in recognition of
his vahialile contributions towards our knowledge of the avifauna of Trinidad.
12(1. Ceophloeus lineatus (Linn.).
PicuH Umiiliis Linnaeus, .%.s7. Xnl. .\ii. 1. p. 174 (17(10.— ex Brissoii : Cayiina).
One cJ and 2 ? ? from Seelct, Ajn-il l'."i:i. In bad ]dumage, with the
abdomen much stained. Tliey are apparently nut dillereut from Cayenne si)ecimens.
127. Momotus bahamensis bahamensis (Sw.).
Pr'viiiilrx litilinniniiis Swainsoii, Aniin. ni Miiiii'j. (ISliS) p. '.V.Vl [•■ Bahama IslaiiiLs "— urrore !].
Moinolus smiimoni Sclater, Cat. Amer. Birds (1802) p. L'(il (num. emend, on grouiids of puri.siu].
3 ? ? from Cai)aro, A]iril ; 1 (?, (Uniguaramas, danuary : t? ? , Fointe (ionrde,
January ; and 3 ? ?, Cliaguanas, May and June.
This series agrees in every way with 14 specimens from Tobago. All have
the whole under surface from the chin to the under tail-coverts deep ciunamon-
rnfous.
( 41 )
M.h /wA'<OT('//.s/.s is eviilently coiitiiiid In the islands of Tnnidad and Tdbago.
Ill till' V'l'iieznelan coast region (Cinnana ami Puerto ('abello), ,1/. Ii. ri'iiezuelne
Sliarjie * takes its i)lacc. It is distingnislied by its muck 2>iiler luider-iiarts, only
liic breast and abdonuMi lieiiig light cinnamon-rutbiis, while tlic tliroat and loreneck
are ociireons washed with greenish. There are five specimens of this form in the
Tring Musenm, collected by A. Mocijuerys at El Gnacharo (near Ciimanaj, S. Oarlos
and San Esteban (near Puerto Cabello).
}f. /i. si//iiy/frsceng ^d. of northern llolombia (Santa Marta and Cartagena) is
another very chjse ally, but differs from M. h. r,'ne.:uclae by its generally more
greenish uii<K>r-p;irts and the colour of thr nuchal cincture as pointed out by
Mr. Sluu-pe.
l~!s. Ceryle americana americaua (Gra.).
Ah-ritn tunrnntmiGmeWn^Syst. Xnl. 1. i. (I7W«) p Ifil [Ijasod on Daubeiiton, P/. c/(/. 591 . fig. l.'-'.—
1 ?, Points Gourde, January; 1 (?, ( 'haguaramas, January; 2 cJcJ, Seelet,
April ; and 1 tj, Caroni Swam[i, March.
Identical with specimens from Tobago and t'ayenne.
N.B. Alcedo hrasiliensis Gm., (pioted in the Cat. Birds as a synonym of
C. a)Hcricana,ca,\motm any way be referred to tlie present species. This will be
easily conceived by a glance at the description.
129. Ceryle superciliosa superciliosa (Linn.).
Akah siipcri-iUnsii Lianaeus, Si/st. Nat. xii. 1. (170(j) p. 17'J [ex Edwards and Hrissoii.— " ill
America." — We substitute Surinam as typical locality].
<J ? from Seelet, April ; c? ? , Caroni Swamp, March ; and 1 ¥ , Chaguaramas,
January. Absolutely identical with speiwiuens from British Guiaiui and Surinam.
The species does not occur in Tobago nor on the " Cote de Paria."
130. Trogou curucui curucui Linn.
Trnijun riiru-ni Liiiuacus Sijsl. Nat. .\ii. 1. (ITOii) p. 1G7 [ox Brissou : ex Marcgrave. — " Urasilia"
(e.\cl. Syn. Hern. — Mexico)].
Trujnn CiiUiirU auct.
One S and two ? ? from Caparo, April. They agree with a scries from
Tobago, Cumana, and Puerto (!abello. Typical Brazilian skins are not available
for comjiarison.
The above is the proper name of T. collaris auct., as has been pointed out
long ago by Cabanis and Heine. f By all recent writers, however, the term
7'. cjiriicui has been ignored, apparently owing to the erroneous description of
Linne, who calls the nnder-jiarls " fulvus," although they are exactly described as
red both by Brisson and Marcgrave, upon whose accounts the name has been based.
Brissou's description is so clear iind full as to leave no doubt of his liird being the
same as T. collaris auct.
In Western Ecuador an easily recognisable race, T. cnrucid cirfiinall.'i Cab. &.
Heine, takes the jilace of the typical form (cfr. Salvadori & Festa, Boll. }[//■•<. Tor/no,
XV. No. 36s (190U) p. 17).
* Momntiix irnc:iielae Sliarpc, Cat. liirds xvii. (1S'.I2) p. 321 in text (S. Esteban ami I'licilo t.'abuUo,
Venezuela).
f Museum Ih'ui. iv. (lsij;j) p. 177.
( -12 )
13). Trogon viridis Liim.
TrorjiMi ihiilis Linnaeus, Si/st, Xul. xii. 1. (17156) p. 167 (ex Brisson : " C'li/miin")
Tidijmi ci/niiiiriis Finscb, P. Z. S. 1870. p. fi.'i',' (Cayenne 1).
6 (J (J iiiiil 2 ? ?, Ciii>aro, April ; 1 ?, Valoiiciii, Miuvli ; cj ? , Cliagiiaiuis, May.
Not diflereiit from a scries from Vcuenuela and iSuriuam.
1;!,'. Trogon violaceus violaceus (im.
Trillion rinhiciHs (imulin, !<ijsl. Xnl. 1. i. (1788) p. 4(l4 (ex Knelieuler, Xnr. Ail. Pclnip. ii. p. 436.
No. 7, tab. 16, fig. 8. — As terra typica accepted Siiriimin ; cfr. Xot\ Znul. ix. 1902. p. 106).
Triifjon mcridinmtlis Swaiiison, Aiihn. in Mcmrif. (183H) p. 332 ('' Uahanias." — Type in Mus. Cautabr.
examined by me).
3 c?t? from Cnjiaro, March and Ajiril, perfectly agreeing witli specimens
from Britisli Guiana. Wc received also 1 ?, cnllocted l)y Dr. Percy Reiidali, at
Savannah Grande, Trini(hid, Felirnary 14, Is'.tT.
This sjiecies has not yet been recorded from I he north coa.st of Vene/.nela, and
I greatly donbt, therefore, whether the siieciniens from llemedios, W. Colomliia,
referred to 7'. riolarf's by Grant,* are correctly determined.
In U]iper Amazonia the typical form is represented by 7'. dola-ceus ramoiiinims
Dev. it Des Mors.
133. Galbiila ruficauda ^'n^.
Giilbnhi nifii-iiudii Cuvicr, IHijiir Anim. i. (1817) p. 420 (ex Lcvaillaut, O/.s. Piiniilis. ii. tab. L. —
" La Giiyane," sc. Cayeiuie).
3 cT(?, 4 ??, from Caparo, April and Slay; 1 ?, Valencia, March; J?,
Seelet, March ; c? ? , Laventille, Marcli and April ; 1 cJ, Chagnaramas, Jannary : and
1 ?, Clnignanas, May. We have also a series of 14 specimens from Tobago whicli
are jierfectly similar. As a rule, the birds from tlie islands are rather darker on
the abdomen Iban those fonnd on the nortli coast of Veneznela (Cnmana, Valencia,
Puerto C'abello) and in the Orinoco valley ; but some from Puerto Gabello are not
different, one being even more intensely coloured than the darkest from Trinidad.
Hence I prel'er, for the j)resent at least, to include all under the name riiticnuilii.
The above observations are based iijion a splendid series (d' (14 adults in the Tring
Museum.
134. Ramphastos vitellinus I^cht.
Riunphiistfi.-i rilrllhiux Licbtcnstein, I'm-.. /'///./. (IS23) p. 7 ((':iyaria).
1 ? I'rom Valencia, March; 1 (^, Cliugnanas, I\Iarfh ; -^ S S from Cajiaro.
April. Besides these s|iecimciis we have I rf IVom Savannah (iraiule, February,
collected by Dr. Percy Kendall.
They agree in size and culour wilh a g 1 series from Surinam, (Juanoeo
(Orinoco delta), and Bi'ilish (iuiaua. Tliis sjiecies does not oeeiir on the " Gote de
Paria," nor on the island of Tobago.
135. Crotophaga major Gm.
Ci-iiln/iliiiijii niiij.ir Cmelin, S,/.^l, Xnl. I. i. ( I7S,S) p. .'■.r,.', (ox I!i-is.soii and liulTon.— Cayenne).
1 cf friini Lavenlilli', March; ajid another <? I'rom Caroui Swamp, February.
• C'ul. UmU xvii. (Is:i2) p. 46S,
( 43 )
I'M. Crotophaga ani Limi-
Crotopluuja aui Linnaeus, Syd. Nal. x. (175S) p. 105 (ex Brown & Sloanc, Jamaica ; and Marcgrave,
Eastern Brazil).
3 ? ? from t'aparo, April ; 1 ? , Seek't, March ; and J ? , Cliagtiaramas,
June.
137. Tapera naevia (Ijiiu].).
Cucidm imei'lus Linnaeus, %s(. N.U. xii. ! (17i;C,) p. 170 (ex Briason.-Cayenue).
8 c?(??? from Caparo, April aud May; 1 immature bird from liaventillo,
November ; 1 i from (_"ha,^iiauas, May ; and 1 pnllus from Valencia, JIarch.
138. Piaya rutila (111.).
Cticiilus mlilux Illiger, AhhiiM. Akad. Berlin, 1812. p. -224 (ex Graeliu, C. mj/iwis var. (i (and y
errore !; : (ex Brissou, Cayenne).
riiiija miittffa auct.
1 S from Caparo, April 14, l!i02, aj;reeiiig with a series from Surinam, but
rather darker rufous on throat and foreneck.
130. Piaya cayana iusulana n. subsp.
[Cuculus caymms Linnaeus, Syit. N,U. xii. 1 (17(;i;) [i. 170 (ex Brissuu— Cayeimf)!
Si.x adults of both se.xes from Caparo, April ; ■^ d d iVoiii Chajruaramas,
January; 1 J, Chaguanas, Blay ; S 'i from Valencia, March. In addition to
these we have some specimens from Savannah Grande, February, collected by
Dr. Percy Kendall.
Similar to F. cayaiia guiuiieiisis (Cal). & Heine), but ditfers by its much liirhter
and brighter cinnamon-rufous (instead of dark rufous chestnut) upper parts, and in
having the outer web of the second and third rectri.x entirely rufous (except a
distinct blackish subapical band). The lower jiarts are just of the same pale ashy
grey as in J', c. ijiiiaiiciisis, the under tail-coverts Ijeing only a shade darker than
the iibdomen.
Type in Tring Museum : " c? " ad. (Uiaguaramas, Trinidad, Jaimary 11, 1003 :
E. Andre coll. Wing, 13.j ; tail, '^37 ; bill, '^8! mm.
This interesting race is evidently contined to the island of Trinidad. A large
series from northern Venezuela (Oumana and Uuaca) is easily distinguished by the
still paler cinnamon-rufous upper parts and by having the outermost tail-feather
on both webs rufous (except tlie blackish subterminal band) ; the rufous colour on
the two next rectrices is also more or less extended over the inner web, which is
never the case in the Trinidad form.
With about one hundred adults from suutli of the isthmus of Panama before lue,
I am able to distinguish the following races occurring north of the Amazon :
{fi) Fiatja cai/ana cai/ana (Linn.).
Cuculus cai/anus Linnaeus, tSijat. Sat. xii. 1 (17(50) p. 170 (ex Brisson. —
Cayenne).
Hab. : < '(ii/niiir * and f.o/rrr An/moiiia : vicinity of Parat (Sleere, Pobert coll.).
Under-surfnce of the tail-feathers dull blackish, without any rntbus tinge. Upper
* Tluee specimens. f Four specimens.
( 44 )
jiiirts (lull (■iiiiiaiiioii-riifuiis, liivii.st Mid alidoiJU'ii very ]>iile whitish givy, iiudi'r
tiiil-coverl.s [liilo ;ishy i,'i'fyish.
(6) I'/iii/a cinjniia ijukiiininix (( 'iib. k lloiiie).
J'i/rrliocoiTi)xf/iiian('niiisV;\.\yM\\» k, lli'iiii', Ma.s. U,iii. iv. (18(32) p. So [(luiaiiii].
Hiili. Biitixh (Juidiin : CJiionju,* Aiiiiai * ; Suriiiam : Paramaribo,! Kwata t ;
Diinnro: Altagracia, f (.'aicara, t t'iinhul P.nlivar^; ('aura li.: Snapnre, + La
I'ricion. %
1,\Vv P. I'. r,ii/,iHii, with tlio tail-feat hiM-.s iiiuK'riu'adi iiiiil'ornily blackish, Imt.
the breast ami abiloiDeii are distinct ly darker ashy grey and tlio under tail-coverts
more sooty greyish. ll|ipcr parts distinctly darker, rulniis chestnut.
(r) I'iai/a atijnnn cohnnbidwi (Cal).).
r;/rrliococ<:)/x coliniibi<(ini!i C&h-AWK, Jdiini. f. (hiiil/i. 18(1:.'. p. 170 (Cartagena,
and Aragna, Venezuela).
Hab. Sorth Vcnczndn : from CiuiiaMaH to Duaca * ; and yortli Colomhl'i. :
Cartageua.*
IJeadily known from (n) and {h) by its luuch lighter, bright ciiinaiuon-rufous
upper parts, and by having the three outer tail-feathers (except the blackisii
subterruinal l)ar) mostly rufous underneath. Under parts just as in P. c. giiianensis.
One adult from l)uaea is exactly similar to those from Cnmana, the crissum
being jialo grey, not Idack as in P. c. nitjiicriam, which occurs i'arther west, near
Merida.
{(I) Piai/a cai/niia innuht na Ilellm.
Hab. : Triniilail (for characters see above).
{e) Piaya cayana niyricrixm Scl.
I'iaya p/i/r/cr/xsa Sclatcr, P. X. .S'. 18C(I, p. 28.") (Pabahoyo, W. Ecuador).
Easily known from all the foregoing races by having the crissum blackish, in
marked contrast to the ashy grey abdomen.
llab. : Culomhia : Bogota coll.H ; Venezuela: Merida t; yiV-Mrtf/or on both sides
of the Andes •* ; JSoith Peru: Guayabamba, f Loreto,* Ucayali.*
140. Amazona amazonica (Linu).
Psillni:i(s nma^uniiii.t Liiiiiavus, %v(. Xut, xii. 1 {\1(,[\) p. 117 (Suriiiam-cfr. .Vor. Znnl. ix. (1002)
p. 110).
One pair from Seelet, April, and one S from Chaguanas, December.
They .igree with a large series from Hurinam and (iiianoco (Orinoco delta).
The species docs not occur on the Paria jieninsnla, and reached Trinidad evidently
from British Guiana through the Orinoco delta.
MI. Piouus menstruus (Mnn).
Pnitlitcua iiinisfi-iiiis Liiiiiauis, Nys^ S'nl, xii. 1 (170tj) p. 14« (ex lirisson— Cayenne).
Sir. Andre did not send ns this species, but Chapman says it was "common in
pairs or small flocks of four to seven birds."
* One siieeimen. % Three specimens. || Nine specimens.
f Two specimens. § Four specimens, ^ Six specimens.
** Sixteen from W. Ecuador, four from E. Ecuiidor,
( 45 )
142. Urochroma batavica (Bodd.).
r.silliini Biilnrwa Boddacrt, Tnhl. IV. nil. (\7H?,) p. 49 [based ou Daubonton, /'/. rii!. Till, fig. 1.
'* Batavia " — errore ! We substitute Surinam as typical locality].
Urochroma chi<jiilttta auct.
2 cJcJ, 4 ? ? from C'angregal, May ; aud 2 ? ? from Cliagnanas, September.
This series agrees perfectly' with two specimens collected in the stat(? of
Ciimand In- A. Mocqnerys. Several skins from the interior of Britisli Gniana differ
slightly in having the feathers of the nape more distinctly margined with lilael;ish,
but this is not liUely to be a constant character.
143. Pulsatrix perspicillata (Lath.).
Strir perspic.ilUUn Latham, I ml. Oni. i. (17'.H)) p. L'l (Cajana).
i ad., ? ad. and S juv. from Lavcntille, October and .Inly.
The adnlts are mnch paler- bnif (not ochraceous) on the belly — than a series
from Snrinara, British Gniana and North Ecuador. A larger series from Trinidad is
rc<iuired to confirm the constancy of this difference.
144. Ciccaba virg'ata (Cass.).
Syniiiim firf/ati(iri Cassin, Prnc. Arwl. I'liihn/. iv. (IS.'")0) p. 1-24 ("South America").
1 c? fere ad. from Cajiaro, May; 1 d juv., and one young bird, not sexed, from
Cliagnanas, October and .June.
The first specimen, being nearly adult, agrees best with another from Guatemala,
while an adult male from Curaana is mm-h more suffused with ochraceous below.
14.5. Glaucidium brasiliauum phalaenoides (Hand.).
[Slrij:biiiiiilkimi Gmelin, .S'//.s7. .V<i/. 1. i. (17ss) p. I'S'.I (ex Brisson ; ex Marcgrave. EuNterii Brazil —
cfr. Berlepsch, Bull. Bril. Orn. CI. xii. lilill. p. 8 11).]
Sirh phihininiilrx Daudin, Trtiitf iTih-u. ii. (ISOO) p. iOO (Trinidail).
4 Si, dajiaro, March ami A]iril ; (1 SS, 2 ??, Laventille, December and
March; 2 cJc?, Valencia, March ; 1 cJ, I'ointe Gourde, .January; 1 c?, (Jhagnara-
nias, .lannary ; 1 ?, (Jangregal, May ; J ?, Chagnanas, May and .lime.
In the series both the rufous and the brown ])hases are represented. They
agree with sjiecimens from British Guiana in size and coloration. AVhile the fo.\-
red birds from Cumana are exactly like those from Trinidad, the brown phase is
rather more greyish on the upjier surface; but some specimens are hardly distin-
guishable, and I believe that the birds fnnn Venezuela and British Guiana cannot
be separated from G. h. plialaenoides, a view already expressed by l!erle|)sch and
Hartert (New. Zool. ix. l'.H)2. p. 110).
14(1. Pisorhina choliba crucigera (Spix).
\Slrix chnliha Vieillot, Nmw. Dirt. vii. (1817) p. ii'.t (ex Azara No. 48.— Par.aguay).]
Stri.r cruriijera Spix, Ar. Unix. i. (1824) p. 22, tab. ix. (" juxta flumeii Amazonum ').
3 SS, 2 ? ?, 2 young birds fnitn Laventille, Ai)ril, May and December;
1 S from Cajiaro, April ; and 1 6, Chagnanas, May.
The specimens agree with a good series IVdui Merida and the (Iriniico vaUey
(Ciudad Bolivar, Quiribana de ( 'aicai'a ami I'eiico), wliieii I inid comiiared with the
( 46 )
t_V[M? (if .S'//v> rr>iei</rr(i Spix. Like P. c. dcciisstitd (Ijclit.), this i'orui has no fox-red
jilia.se, lint (lid'ers hy its slight!}- snialler size, piiKr njiper jiarls in all plumages,
mid less Imfiy mixture on the lower surface.
147. Buteo albicaudatus albicaudatus (Viciii.).
Iltiltn tilhitiiatliiliia Vieillot. Xoiir. iini. iv. (IKlii) p. 477 (ex Azara- Paraguay).
One S from ('hajfiianas, August ; and one jiair from the outskirts of Port of
Spain, Slareii.
None of the three siie('iraena is (piite adnlt, thougii in the two males the
wlioje hreast and abdomen are luaiuly white, only the throat being blackish. In all
tiiree examples there is a large rnfous jiatch on the shoulders.
So far as I am aware, this is thf^ first reeord of the sjiecies for Trinidad.
Leotand clearly describes Buteo ubbieriatux Cab. s. nom. B. zoiiocfrciis, but Mr.
Andre has not sent ns the latter species.
1-ls. Urubitinga albicoUis (Lath.).
Falro albicoUis Latham, //«/. ()r>i. i. (17'J0) p. .'10 (■' Cayaua").
A single ? adnlt from (Miaguauas, Ajjril, exactly agreeing with other specimens
from ('nujana and N.E. Pern.
14'.i. Elanoides furcatus (Linn.).
Fiihofiirciiliix Linnaeus, Si/xl. A'lit. xii. 1 (IViifi) p. l'J9 (ex Catesby — Carolina).
One adult, not sexed, from T'hagnauas, July.
15(1. Harpagus bidentatus (Lath.).
Fdlcn hideiilaliishathim, Tnd. Orii. i. (17yO) p. :!8 (" C.ayana").
Two immature birds, one marked cJ, the other not sexed, from Chagnanas,
April an<l May.
ini. Ictinia plumbea ((Jm.).
Fiilmplumhius Gmelin, Si/sl, A^at. 1. i. (1788) p. 283 (ex Latli.am— Cayenne).
One d and one adult not sexed, from Chaguanas, April ;iuil May ; and one
c? from Seelet, Ajirii.
152. Falco peregrinus anatum P>p.
l^Falcn peregriinis Tunstall, Oni. lirilatm. (1771) p. I. (ex Brit. Zool.).]
Ftilin (iiiuliim Bonaparte, Omiji. Lixl Binh Eiir. and jV. Amei: p. 4 (18118. — ex Auilubou, pi. llj.
" Northern parts," sc. of North America).
0[\r Y ad. from Srelet, April 5, 1905.
l.');i. Columba speciosa (im.
Cnhimha xperiiim Gmelin, S;/st. A'al. I. ii. (1788) p. 78.3 (ex Bu Hon— Cayenne).
One i ad. from Aripo, 21)00 ft , May, aiul a female, collected by Mr. Carr near
Ohagnaiias, November 15, 1894.
Idcniicid with examjiles from I'ritish (iniana and ( 'ayenne. The s|)('ci('s has
as yet not been found on tlii' ojjposite Venezuelan coast, and might be looked upon
as an inlrmli'r from Guiana.
( 47 )
1.".4. Columba rufiua rufiua Toraiu. & Knip.
Columha rufim, Temmiiic.k & Kiiip, /V""' '• f-i'"- ^ec. p. 59. tab. L'4 [1808-11: "La Guiane
f ran^aise "].
One J und two ? ? from Reolct,, A]ii-il. Tlio former has the nape pure
mctallie purple, instead of bronze -jjreen witli golden reflections, Imt tliere is
sometimes a sliglit pnrple adrai.xtnre to lie seen in sjieciraens from Rritisli (Jniiuiu.
In northern birds the tail is always nniformly pale brownish .C'rey, wliile those i'rom
Sonth 15razil have the basal portion blackish, in strons' contrast to the bidwiiish
grey apical band. The sonthern form ajipears to be entitled to the name ('. riijbia
si/lrestris Vieill.,* but I have not yet been able to compare typical Paraguay
skins.
C. r. nifiiiu is not found on the north coast of Venezuela, but it occurs also on
the island oif Tobago, wlience Mr. Andre sent us a fair series.
-ir);"). Cohxmbigalliua t rufipennis (Bp.).
Taljianilia nifiitciiiiis Bonaparte, 0'ii>ip. Ar. ii. (1S54) p. 70 (Carthagena).
Seven c?cJ ? ? from Caparo, April, and a single ? from Seelet, March.
They agree with a large series from (Uiniana, Tobago, and Bogota. From
Surinam tlie Tring Museum has lately received C. tnlpacoti (Temm. & Kuip).
In the Cat. Birds xxi. p. 4s7, the specific name nifpennis is wrongly attributed
to G. H. Gray, who never described the species. The first description was ]iublished
by Bonaj)arte ; consequently, spec, v", w" of the Brit. Museum cannot be the types !
156. Claravis pretiosa (Ferrari-Perez).
Perislern irretimn Ferrari-Perez, Proc. U.S. Mns. ix. 18SC (1887) p. 175 (ex Jalapa, Mexico).
One S ad. each from ( 'hagnaramas, January, and Laventille, A])ril. They
a^ree . perfectly with a large series from ( 'nraana and Brazil. I am unable to
separate the South American form from typical ( '. pretiosa e.x Mexico.
157. Leptotila verreauxi Bp.
Lejilnpiila verreauxi Bonaparte, Cmisp. Av. ii. (18£14) p. 73 (" Nova Granata ").
LqiUilila himlaris Richmond, Proe. U.S. .V«-'. xviii. 18'.l5 (18'JG) p. G59 (Margarita lit).
Two ? ? from Laventille, March, and one ? each from Pointe Gourde, January,
and Seelet, April.
They agree in colour and measurements with a series from Tobago, t'umana,
Margarita Island (L. insuluris) and the Orinoco valley. The differences between
( 'olombian and Venezuelan specimens, alluded to by Berlepsch & Hartert (A'ov.
Zool. ix. 1902. p. 118) do not appear to be constant; yet a larger series from
Bon-ota may jirove them to be separable. Anyhow, the birds from Trinidad, Tobago
and Venezuela are absolntely identical with the one from Margarita Island.
• Coltimha sijlirnlris Viuillot, A'l.HC. Diet. xxvi. (1818) p. 30(1 (ex Azara — I'aiagu.ay).
I t'l'Iiniihitjiiirttiu Boie, I.-rh ISLMl. p. !)77 (Species iinica : C. jjitssf-rinu Linn.).
(48)
l-">8. Leptotila rufaxilla (Hicli. >t Bern.).
Cohimhn nifttoriVa Richard & Bernard, Ail. Soc. Uist. Nat. Paris i. (1792) p. 118 (Cayenne).
c? ? ?, Caparo, April ; c? ? , Cliagiianas, May ; ?, Valencia, Marcli.
Most of tlu'so specimeii.s are rather darlcer, more rufuns on foreneck and breast
than others from the mainland of Veneznela, lint the dill'irence is nut ijiiite
constant.
liiO. Geotrygou sp.
Pet-htrra frnmid (nee Tscliudi I) .Tardiiie, Ann. Miiij. Nal. Hid. xx. (1847) p. 374 (Tobago).
P. liiiearia (nee Prevost <t Knip !) Lrotand. 0/.v. Trinnhitl, p. .'i7.*i (Trinidad).
There can be scarcely any donlit that a species of (Jcofri/i/o/i inhabits the
monntainnns jiarts of Trinidad, althonn'ii it has not been collected there of late years.
Kirk sent it from Tobago, bnt, nnlbrtunately, his specimens seem to have been
lust — at least they are not in the Britisli nor in the Tring Jlnseum. As the nearest
allies occur in the Andes of Merida (G. liiiciiri.s i:('/ic:/tr/e/i.vs Salvad.) and
Colombia {G. I. linearis), the birds inhabiting Trinidad and Tobago form most likelv
a dillerent sjiecies or subspecies.
160. Pipile pipile (Jacqn.).
Criir pipilr Jacquin, Bri/li: GexiJi. Viigel (1784) p. 20. tali. 11. (1784: ''Am Orenokofluase bei
Kumana ") unde :
Pipile Jacqiiini Reicbenbach, Oiliimli. p. 154.
Penelope cuinanrnxix (uec Jacquin !) Li'otaiul, Oix, Triniihiil (ISl)!)) p. .'i8.^ (Trinidad).
r, pile pipilr Cliapman, Bull. Amn: Miis. vi. (1804) p. 74 (Trinidad); Ilvllmayr, Hull. Bril. Orn. CI.
xiv. No. 10-, (March 1904) p. f>;t (Trinidad).
One ?, killed at Caparo, Trinidad, Ajiril I'.i, 1'.)U2. "Iris brown, liill blade,
feet red." Wing, 345 ; tail, 30(J ; bill, 331 mm.
This bird is, no doubt, of the same species as those fignred by Jaciinin and
described by Leotaud and Chapman. As I have shown in a note, jinblished in the
Bull. Brit. CI. .\iv. p. 59, it is quite distinct from /'. CKm(nu'iini.'<, with which it has
generally been confounded. In fact, it is more nearly related to P. cujiilii (VAz.),
bnt there are some differences whicli serve to distingnish it at a glance. The
feathers of the pilenm are mainly blackish brown with only narrow while edges
laterally, which in P. ciijiili Ava much broailer; the middle upper wing-coverts are
])nre white except the extreme base and a large blackish brown tiji, while in
/'. r//y///!i/ they are blackish brown willi a short white stripe on each margin ; the
white on the greater series, too, is much more extended in the Trinidad bird,
occupying nearly the whole outer web except the dark-colonred tiji and reaching
almost to the shaft. In P. ci/Jiihi, however, there is only a narrow white margin
along the second third of the outer web. In the jiurplisli gloss of the jilnmage
both species agree, as they do also in the amonnt of the naked skin on the sides of
the head and on the throat. In both there are but very faint indications of white
edgings on some of the feathers on the foreneck. I may add that the bare skin of
the throat in the Trinidad specimen looks uniform dark grey, whih' in /', nijnhi
the lower jiart is decidedly reildish tlesh-coloiired. A better series of both species
is rctpiircd to confirm the constancy of this character or otherwise.
i'.7W«//fly« (Spix) is ijnite distinct from P. jii/iilc and /'. nijuhi, having the
chin and anteriiir ]iart of the throat as well as the sides of the head densely i-overed
with ileep black feathers. Moreover, the forehead is broadly black, and the feathers
of the |iili'nm have only hair-like blackish shaft-lines.
( 49 )
The four known species oi Pi/Jile may be distingnislieJ liy the following key.
A. Sides of the head (except a narrow rim round the eye) as well as chin and
upper part of the throat thickly covered with deep black feathers. Fore-
head and superciliaries deep black. Pileum and crest with narrow,
liair-like blackish shaft-lines. Plumage dark brown, glossed with {mrjile.
Foreneck and breast strongly edged with white. . . . P.jacutiii[/a (Spix).
B. Lores, superciliary — and eye-region, cheeks and the whole throat naked,
only a broad malar stripe being feathered. Forehead never black.
a. Plumage glossed with green. Pileum and crest nniform huffy white,
or with hair-like blackish shaft-lines ... P. cumanensis (Jacqu.).
h. Plumage glossed with purple. Pileum and crest with broad blackish
brown stripes.
i'. White margins of the crest-feathers broader. Median series of
ujjper wing-coverts purplish brown, with a white margin on
each side ; greater series with a narrow white margin along the
second third of the outer web 1'. cujiihi (Pelz.).
b-. Crest-feathers blackish brown with only very narrow white edges.
Median series of upper wiug-coverts pure white, except the
extreme base and a distinct blackish brown tip ; greater series
on the outer webs white to the shaft excej)t the tip.
P. pipile (Jaciiu.).
101. Crypturus soui (Ilerm.) (subsj). ?)
Tinamiis sunt Ilcrmauu, Tab. Aff. Aniiii. 1783. pp. 11)4, 255 (ex Daubeuton, I'l. cid. 8"J'.l. — Oii/iiiiii).
Cnjpiuran pileidus auct.
One ? ad. and d juv. from Ca])aro, April ; two ? ¥ ad. and one S juv. from
Chaguanas, -June and January.
Tlie adult birds agree best with others from the Orinoco delta (Guauoco), wiiile
those from Cumana and British Guiana, as a rule, are rather lighter and brighter
beneath. A large series from different parts of its range is reqirircd for defining
the various geographical races into which C. soui is doubtless separable.
162. Eudocimus ruber (Linn.).
Taittalus riihcr hinnncus, Syst. Nat. xn. 1 (17(JG) p. 241 (syn. part.) ["in America": ex Bri^son
Cayecne) and Catesby (Bahamas). — We accept Cayenne as typical locality].
A single ? adult in red plumage, from Garoni Swamp, October.
1G3. Ajaja ajaja (Linn.).
riataka AJaJa Linnaeus, Syit. Nul. .xii. 1 (1706) p. 231 [ex Brisson (Cayenne), etc.].
A single young ? from the month of the Oaroni River, July.;
164. Herodias egretta (Gm.).
Airka Eijirtia Gnielin, Si/kI. Nat. 1. ii. (1788) p. 02i) [" iu insula S. Dominici, insulis Falkland
et America australi ad Louisianam." — We take Cayeinif (e.\ Buffou, Umelin's first and
principal source) as the typical locality].
Three S ad. from Caroni liiver, March and August. Identical with skins frem
Surinam.
4
( 50)
IC)."). Florida caeriilea (Linn.V
^ )■(/(■« '(/en; /<a Linnaeus, Sijsl. XhI. x. 1 (17o8) p. 143 ["in America scptontriouali." — We accept
as the typical habitat Carulina ex Catesby],
One ? ad. in the shity bliiisli, and c? ? in the white plnmage, all from
('aroui Swamp, Blarch. We have also a good series of both phases from Tobago.
106. Hydranassa tricolor rufimentum n. snbsp.
Agrees with 77. tricolor tricolor (P. L. S. Miill.) in size, but differs at a glance
by its much darker, blackish instead of slate-grey colour of the upper parts and
neck, and by having the chin like the whole middle line of the throat chestnut,
while the chin is always white in 77. t. tricolor and II. t. rujicollis. The chestnut
colour on the throat is also decidedly darker than in the two last-named subspecies.
Type in Tring Museum : " c? " i)erad. Caroni Swamp, Trinidad, March 22,
1UU2, E. Andre coll.— Wing. 214 ; tail, 73 ; bill, 85 mm.
Unfortunately Mr. Andre sent only one male of this interesting new form.
It differs, however, so strikingly from ten examples from Cayenne and Surinam
(true 77. t. tricolor), and from a good many specimens of 77. t. rujicollis that 1 feel
quite coutident as to its distinctness, and have no hesitation in describing it as
a different race. In both of its allies, the chin and more or less also the anterior
portion of the throat are white. The upper ])arts and the foreneck are always
bluish or slaty grey, while they are deep blackish in the Trinidad form,
and the elongated feathers on tlie rump are decidedly darker, more umber-brown,
in the latter.
We have thus to distiuguisli the following subsjjecies :
(a) lli/dranassa tricolor tricolor (P. L. S. Miill.).
Ardea tricolor P. L. S. Miiller, yatarsi/st. Sup pi. p. Ill (1 776 : ex Daubcnton,
PL eiil. 350.— Cayenne).
Arrlea leucogaster Boddaert, Tabl. PI. cnl. [i. 21 (1783 : based on the same).
H;ib. ('(njeniic : Roche Marie, etc. (Clierrie coll.) ; Surinam : near Paramaribo
(Pcnard coll, — Mus. Tring); J'.Zs. Bnizil: Cajutuba, near Para (Natterer coll.).
2 (? ad., Surinam. . Wing, 2i(t ; tail, 65 ; bill, 82, 85 mm.
8 from Cayenne . . „ 204 — 218; ,, OS — 78; „ 78 — 80 „
(Ij) Ilijdranassa tricolor rujimcnlum llellm.
Hab. Trinidad.
(c) 77. tricolor ruficollis (Gosse).
Ec/rctta rujicollis Gosse, JJirds Jamaica, j). 338, tab. 03 (1847 — Jamaica).
Hab. ^V'armer portions of Eastern JS'ortk America ; south tiirough Ccidrul
America to Panama and JV'. TP. Ecuador ; and the West Indies south U) Aruba, off
the north coast of Venezuela.
2 ad. (c? ? ) from Georgia .
I c? ad., Jamaica
1 ? ad., l.sje of Pine, Cuba
cj ad., Mexico . . . .
2 ((? ? ), Andros IsL, ]5ahamas .
1 S jr., Panama (Harris coll.) .
1 ad., Vaqueria, N.W. Ecuador.
1 ? ad., Aruba (Hartert coll.) .
^&j
( 51 )
107. Nyctanassa violacea violacea (Linu.).
Anlea riuliircu Linnaeus, Si/tit. Nat, x. p. 143 [1758 : '' in Aiuorica septcutrionali." — Carolina (ex
Catesby) as typical locality accepted].
Three c? ? ? and one yoniig bird from Caroui Swaiii|), March, and Poiute
Gourde, January. We have also a fair series from Tobago.
168. Agamia agami (Gm.).
Ardea Aijaini Gmelin, Sjsf. Nut. 1. ii. (1788) p. IJ2'J [ex Biiffonand Daubenton. — Cayenne].
Two cJ ad. from Caroni kSwamp, August and September. Tiiey are rather
larger than others from Guiana and the Gaura lliver, Venezuela, but not otherwise
diiferent.
16'J. Nycticorox nycticorax naevius (Bodd.).
[Ardea Ni/dicora.c Linnaeus, Si/st. Nid. x. p. 142 (1758 : " in Europa au.strali ").]
Ardca naevia Boddaert, Tabl. Pi. cnl. p. 50 (1783 : ex Daubenton, PI. enl. 939. — Caijr.iiue).
One ? ad. and one young bird from Caroni Swamp, March.
170. Cancroma cochlearia Linn.
Cancmimi C'ic/ilmria Liun;iciis, Sysl. Xnl. xii. 1. p. 233 (17liG : ex Brisson. — Cayana).
2 t? ad., 1 d juv. and 1 ? juv. from Caroui Swamp, March. Tliey are
ajiparently not ditl'erent from Cayenne skins.
171. Butorides virescens (Linn.) an snbsp. ?
Ardea virescens Linnaeus, Si/st. Nat. x. (1758) p. 141 [ex Sloane (Jamaica) and Catesby (Carolina)].
Three immature birds from Caroni, March, and Chaguaramas, January.
As far as I know, this sjiecies has not been taken before on tlie island of
Trinidad ; but since Mr. Andre sent only young birds, I cannot determine to which
of the numerons subspecies they should be referred. I may remark, however, that
the Tring Museum received from Tobago a good series of what appears to be true
B. virescens,
172. Ardetta erythromelas (Vieill.).
Ardea enjthromchiH (err. typ.) Vieillot, Nniw. Diet. xiv. (1817) p. 422 (ex Azara No. 3(i0. —
Paraguay).
1 ? ad., c? juv., and a young bird not sexed, all from Caroni Swamp,
March and April. The adult specimen agrees well with others from S. Paulo and
Surinam.
173. Botaiirus pinnatus (Wagl.).
Ardea piniiuliis Wagler, his 1829. p. 602 (" in Brasiliae in'ovincia Bahia ").
A single adult c? from Caroui Swamp, killed March 14, 1002. " Iris golden
yellow." It agrees with another adult S from Aunai, British Guiana.
174. Plotus anhinga Linn.
.V.((. xii. 1. p. 218 (17*
A single 6 from Caroni Swamp, February.
Plotus anhinga Linnaeus, Sijsl. Nai. xii. 1. p. 218 (1700 : ex Marcgrave, Brazil ; and Brisscc
Cayenne).
(52 )
175. Podilymbus podiceps (Liun.)-
Cobjmhus Podii-rps Linnaeus, Syst. Nat. x. p. 136 (1758 : ex Catesby. — Carolina).
A single ? from Caroni Swamp, February.
170. Rallus longirostris longirostris (BoiW.).
Rallus loiigirodrh Boddaert, Tabl. I'l. cnl. p. 52 (1783 : ex Daubenton, PI. eiil. 840— Cayenne).
6 c?t? ? ?, from Caroni and Caroni Swamp, March and April.
Unfortunately there are no Cayenne skins available for comparison.
Several specimens from Western Ecuador, collected in February and September,
differ at a glance by their much paler, pnre olive-grey upper parts, without any
trace of the brownisli tint to be seen in the Trinidad series. They agree well with
Taczanowski's description of R. ci/percti* which therefore has to stand as a distinct
subspecies. I am not so confident as to the distinctness of R. I. crassirostriti Lawr.
of Eastern Brazil, for one specimen from S. Paulo does not show the characters
assigned to this form by Mr. Sliarpe, but a good series may prove it to be another
recognisable race. The distribution of the three forms, then, would be as follows :
1. R. longirostris longirostris (Bodd.).
Cayenne, British Guiana, Trinidad.
2. R. lonijirostris crassiro.stris Lawr.
Eastern Brazil : Bahia, Pernambuco, S. Paulo : Iguape (spec, in Mus. Tring).
3. R. longirostris ajpereti Tacz.
West Ecuador : Vacqueria (Mus. Tring) ; N.AV. Peru : Tumbez (Stolzmanu).
177. Limnopardalus maculatus (Bodd.).
Rallus maculatus Boddaert, Tail. PI. ait. p. 48 (1783 : ex Daubenton, PI. oil. 775 — Cayenne).
1 c? from Caroni, April, and another c? from Caroni Swamp, March.
Identical with specimens from Surinam and British Guiana.
178. Aramides axillaris Lawr.
Aramkles nxilhiria Lawrence, Proc. Pintail. ^Irail. 1803. p. 107 (Baranciuillu, North Colombia).
1 (?, ~ ? ?, from Caroni, May ; and 1 ? juv. from Chaguaramas, January.
The adults agree with another old male from Merida, Venezuela, in every
respect; but I have not been able to compare tojiotypical skins from Northern
Colombia.
The sexes of this si)ecies do not differ in size or in colour.
179. Aramides cajanea (P. L. S. Miill.).
Fulka cajaiiea P. L. S. MuUer, Niitur.ii/sl. Sujijd. (177C) p. 110 (ex Daubenton, PI. int. 352 —
Cayenne).
1 c? ad. from Caroni Swamj), March. In addition to it, the Tring Museum
possesses an adult ? from Savannah Grande, collected in February by Ur. Percy
llendall.
These specimens agree very well with others from Surinam and British Guiana.
* P. Z. S. 1877, p, 717 (Tumbez, N.W. I'uru).
( ^^ )
180. Gallinula galeata galeata (Lcht.).
Crex gtileala Lichtenstein, Verz. Duhl. p. 80 (1823— San Paulo).
One S ad. from Caroni, April ; and another S from C'aroni Swamp, March.
They agree iu colour and diiiiension.s with several examples from South Brazil.
G. (J. fjiirmnni Allen, of Lake Titicaca, Fern, is easily known by its much larger
size, stouter and larger bill, and p-eDerally blacker plumage.
LSI. Porphyriola martinica (Linn.).
Fulica iiKirliinra Linnaeus, Syni. Nat. xii. 1. (170C), p. 259 ("in Martinicae inundatis").
1 (J ad. and 1 ? juv. from Caroni, March and April.
182. Parra jacana jacana (Linu.).
Parra Jacana Linnaeus, Syst. Nat, xii. 1. (1766) p. 259 [(excl. quot. Hernandez) ex Marcgrave and
Edwards. — Surinam : ex Edwards].
One c? and two young birds from Caroni Swamp, April, and a single ? ad.
from Seelet, April, agreeing best with specimens from Valencia, Venezuela.
Typical skins from Surinam are perhaps rather lighter on the back, but are
otherwise not different. P. j. melunopi/f/ia Scl. and P. j. variabilis Linn, are
undoubtedly but geographical representatives of P.j. jacana, and thus more pro-
perly designated by trinomials. Strange to say, one of them (variabilis) has been
made the type of a separate genus, Asarcia Sharpe ! ! -which, we trust, will only be
recognised by those affected with furor ffeneriacs.
183. Arenaria interpres (Linn.).
Tihir/a Tiili'i-jires Linnaeus, Si/U. Nal. x. p. 148(1758: "in Europa et America septentrionali." —
We accept Sweden as typical locality : ex Fauna suec. 154).
1 ? from Seelet, April.
184. Ochthodromus wilsonia rufinucha (Ridgw.).
[Charadriun viUonia Ord : in Wilson, Amcr. Oni. ix. (1814), p. 77. tab. 73, fig. 5 ("Cape Island
New Jersey").]
Acffialilh Wils'iuius var. riifinurha Ridgway, -twice. Nat. viii. (1874), p. 109 (.Jamaica).
4 <S ad. and 2 $ ? from Seelet, April.
All the males have a broad black jugular band, slightly mixed with pale
rufous, and a black patch on the crown just behind the white forehead. The sides
of the head are bright rusty. In the females the blackish crown patch is wanting,
and the jugular cross-band is bright rusty like the cheeks and ear-coverts.
Tills bird is by no means only a winter visitor to the tropics, as Mr. Sharpe seems
to think, for Mr. Hartert found it breeding in numbers on the islands Arnba and
Bonaire, Dutch West Indies.
18.5. Aegialitis semipalmatus (Bp.).
Cliaradrius snnijHiImalus Bonaparte, " Obs. Wilson, 1825, No. 219."*
3 (JcJ, 1 ?, from Seelet, April.
' I have not been able to verity' the .ibove quotation.
( '4 )
ISO. Symphemia semipalmata ((Jm.).
Scoloixix semipulmata Gmelin, Sysl. Nat. 1. ii. p. GjO (1788—" in Noveboraco ').
Two ? ? from Seelet, April.
1»T. Totanus flavipes (Um.).
Scolopax flavipes Gmelin, ^ijft. Nut. 1. ii. p. G59 (1788 ; "in Noveboraco").
1 cJ and 5 ? ? from C'aroni Swamp, March.
188. Totanus macularius (Linn.).
Ti'inya maniJaria Linnaeus, Syst. Nat. xii. 1. p. '240 (17G6 : ex Edwards — Europa (errore !), " et
America septentrionali," ac. Pennsylvania (fide Edwards).
A small series nf both sexes from Pointe Gourde, January; Seelet, April;
Caroui, March ; and tUiagnaramas, January.
189. Barti'amia longicauda (Bechst.).
Triiign loiKjimuda Bechstein, Lcithams Uhcrs. ViJijel iv. (1811), p. 453 (Nordamcrika).'
A single c? from Seelet, April.
190. Ereunetes pusillus pusillus (Linn.).
Tringa jmsilla Linnaeus, Si/st. Nat. xii. 1. p. 2b2 (1760 : ex Brisson — Domingo).
3 <?(? and 1 ? from Seelet, April ; and 2 ? ? from Caroui Swamp, March.
191. Ereunetes pusillus occidentalis Lawr.
Ereunetes occiihiitalh Lawrence, Proc. Acad. I'liilad. 1804. p. Iu7(" Pacific Coast, Oregon, and
California ").
Seven specimens (cJ cJ and ? ?) from Seelet, April.
192. Tringa minutilla Vieill.
Ti-inga minntilhi Yieillot, Nain: Dirt. xx.\iv. (181!)) p. 400 [" a Halifax, ct dans la Nonvelle-
Eco.sse '').
Two ? ? from Seelet, A])ril.
193. Querquedula discors (Linn.).
Anas dixi'ors Linnaeus, Sy.i/. A'lit. xii. 1. p. '205 [1700: "in America septentrionali": ex Brown
(J.imaica) and Catesby (Carolina).]
1 (? and 3 ? ? fr(im ('aroni Swamp, Octolier.
194. Nomonyx dominicus (Linn.).
Anas dominica Linnaeus, Syst. Nat. xii. 1. p. 201 [1700 : " in America meridional " : ox Brisson,
Dominica, and Hernandez, Mexico],
One cJ ad. and one young bird from C'aroni Swaraj), March.
The old bird agrees exactly with another from Peruambuco.
( 55 )
195. Sterna fluviatilis Naura. (an subsp. ?).
Slfrmifliiviiililis Naumann, Isin 1819. p. 1848 (no locality).
A single ? from Seelet, March,
The form inhabiting the tropical portions ofSonth America is perhajis separable
as a subspecies. (Cfr. Hartert, Ibis 1893. p. 309.)
lOCi. Phaetusa magnirostris (Lciit.).
Steriiii miii/iiirot/ris Lichtenstein, Verz. Dubl. p. 81 (1823 — Brazil).
Three immature ? ? from Port of Spain, .June, agreeing with mainland
specimens in corresponding plumage.
197. Ryiichops nigra cinerascens Spix (?)
[_Ri/nrlinj)s nigra lAnntiens, SysL Naf. x. p. 1.^8 (1758: "in America." — We accept Carolina (ex
Catesby) as typical habitat).]
R. cinerascens Spix, Av. Bras. ii. p. 80. tab. cii. (1825 : " in locia ripariis Hum. Ama-imnm ").
R. melanuru auct.
One nearly adult S and two young birds marked c? and ? from Port of Spain,
Juno.
The specimen in most advanced plumage agrees with the adult male from
Caicara, Orinoco, spoken of by Berlepsch and Hartert,* in having pale wliitish ashy
under wing-coverts and broad white tips to the secondaries. An adult bird from
Manaos, Amazons, which doubtless represents typical i?. cinerascens, has much
darker, smoky brown under wing-coverts, and very narrow whitish apical
margins on the secondaries. One S from Cumanji is quite similar, and therefore
I doubt the possibility of separating the Venezuelan form as a dilferent subspecies.
198. Larus atricilla Linn.
Lnriis atricilla Linnaeus, Sijst. Nat. x. p. 1.3G (1758 : " in America," sc. Carolina — ex Catesby).
One pair of adults and one immature c?, all obtained at Port of Spain
harbour, June 1903. Mr. Andrd sent also a good series from Tobago.
II.^LIST OF THE SPECIES OCCURRING ON THE ISLANDS OF MONOS
AND CHACACHACARE, BUT NOT FOUND ON TRINIDAD.
Unfortunately, we know as yet very little about the ornis of the small islands
lying between Trinidad and the Paria peninsula. It is not improbable that some of the
sj)ecies treated of in the next chapter might yet be discovered on one of these islets.
So far as I am aware, the only naturalists who ever visited them were Chapman and
Taylor. The former collected several birds on Monos Island, the latter spent a few
days on Chacachacarc. Both collectors obtained a number of species which have
never been taken on Trinidad, but are common residents on the opposite coast of
Venezuela.
* Aia: Ximl. ix. (1302) p. \?.2.
(50 )
1. Spinus cucullatixs (Swains.).
Chapman observed two specimens on Monos Island. Tlio Tring Musenm
received a large series from the state of Cumana, Venezuela. There is no record of
its occurrence in Trinidad. The " Trinidad " specimens in the r.ritish Museum are of
the well-kuuwu " Orinoco "-make, and came eertainlv not from the island.
-'. Phaeomyias incomta (('ah. & Heine).
Myiopalix nemifiixca Chapman, Jliil/. Aiiier. Mns, vi. (1804) p. .'58 (Mouos Island).
P/ii/llomi/ids semi/ti6ca Taylor, Jbis 18G4. p. 86 (Triuidad).
Chapman found this species only on Monos Island. Its occurrence on Trinidad
proper requires confirmation.
About the nomenclature of the species cfr. Tor. Zool. ix. (1902) p. 41.
3. Sublegatus fasciatus glaber Scl. & Salv.
Siihlegiittis ylaber Chapman, Bull. Amn: Mas. vi. (1894) p. .j'j.
Found by Chapman on Monos Island.
4. Empidochanes fascatus cabanisi (Ldotaud).
Emphloclumes cabanisi Chapman, Bull. Amer. Mas. vi. (18ll4) p. 41 (Monos Island).
Chapman found it on Monos Island not uncommon. Leotaud had only a single
specimen before him, which perhaps was also obtained on one of the outlying-
islands.
Chapman {Bull. Amer. Mus. vi. p. 42) separated the Tobago form as E. c.
canescens [= zireoninus Ridgw. 1886 ! ex Tobago], but the differences pointed out
by him do not exist. "We have both from the Orinoco (Altagracia) and from Tobago
brown-backed specimens with pale yellow uuderparts as well as grey-backed ones
with a nearly white belly. The same observation has been made by Conut Dalmas
{Mem. Soc. Zool. France xiii. lUUO, p. 139 : E. arenaceus).
5. Formicivora intermedia intermedia Cab.
Taylor, Ih,^, 18(U. p. 85 (Chacachacare).
H. Taylor found it " abundant in the low, dry bush of the small island
Chacachacare," though he did not tiud it elsewhere.
We have a large series from Cuman^. On Tobago it is represented by the
much larger and darker /''. intermedia tobagensis Dalmas {Mem. Soc. Zool. France
xiii. 19U0, p. 141).
III.— HYPOTHETICAL LIST.
In the following pages I give a critical list of those species which have
erroneously been recorded for Trinidad, or the occurrence of which has yet to be
proved. Some of them observed by Leotaud may be occasional stragglers from the
mainland, others may yet be discovered in the mountains of the northern part of
the island.
( ^-' )
1. Turdus flavipes venezuelensis (Sharpe).
Merida venezuelensis Sharpe, Moiinqr. Tunl. ii. (lUOO) p. 83 [Venezuela].
J/, mehiimpleum, idem, I.e. p. 87 ["Trinidad "— errore !]
Turdus flavipes (nee Vieillot !) Taylor, Ibis 18G4. p. 80 ; Leotaud, Ois. TrinuUi!, p. lOO.
L^otand states that the species does not breed on the island and occurs only
between the months of Augnst and April, It is strange that neither Chapman nor
Andre's collectors ever met with it.
I have shown* that specimens similar to that described by .Sharpe as
M. melanopleiva may be found in Venezuela as well as in South Brazil, the
distinguishing characters of the supposed species being only those of the very adult
male. Having examined the type in the British Museum, the last doubts as to its
identity with vcncx-iu'li'iis/s are now removed, for the type turns out to he a skin of
the well-known " Orinoco "-make, and certainly never came from Trinidad !
2. Turdus xanthoscelus Jard.
Timhis xanthoscelus Jardine, Ann. Mag. Nat. Hist. xx. (1847) p. 329 [Tobago].
According to Leotaud, this species is an occasional visitor to Trinidad, but
neither Ohaiuuau nor Audrey collected it on the island. This tlirush is strictly
confined to Tobago. The specimens from the Roraima Mts., Britisli Guiana,
referred by Sharpe to 7'. xanthoscelus, differ from the typical Tobago scries by their
much smaller size (wing 110— 112, instead of 117 ; tail 83—88, instead of 98 mm.),
weaker and shorter bill and much more glossy plumage. They are perhaps not
separable irom T. lencojis Tacz., but a better series should be examined.
3. Euphonia nigricoUis (Vieill.).
Although Leotaud says it is pretty common, it has not been met with by any
of the more recent travellers who visited the island. It is also included in Taylor's
list with the remark : "much less abundant than on the main."
4. Piranga testacea faceta Bangs.
Piranf/afii(!et(( Bangs, rmr. Biol. Sor. Was!,, xii, (1898) p. Ul [Santa Marta, Colombia].
Pyranija hepatim (nee Swainson !) Lrotand, Ois. Trinidad, p. 291 [Trinidad].
I'iranya liaemalea (nee Salvin & Godman !) Chapman, Bull. Amey. Mas. vi. (1894) p. 30 [Trinidad] ;
Allen, Ball. Amer. Mas. xiii. (1900) p. 168 [Santa Marta],
I have not seen as yet any authentic Trinidad specimen, but Ldotand describes
a bird which seems to belong to the above species, of which there is a good series
from Cumana in the Tring Museum. It is easily known from P. t. testacea by its
much lighter, orange-vermilion underparts, and is in no way to be confounded with
P. t. liaemalea of the Roraima Mountains, British Guiana, as has been done by
Allen and Chapman. The latter ditiers very little from typical testacea, while
P. t. faceta is much paler and brighter everywhere than its two allies.
Ldotand adds that this species is only a visitor to the island.
• Juurii.J'. Oriiith. 1902, p. 07.
( 5S )
5. Sporophila bouvronides (Loss.).
PyrrhiUi hniin-nnides Lesson, Triuli- il'Orn., 1S31. p. 4.50 [no locality given].
Spermojihila Icxxotii Finsch, P. Z. .S. 1870. p. 582 [" Trinidad "— errore !].
Sj>ermoj>liila trhiilulis Sharpe, Cut. Ilh-ih lirit. yrus. xii. {1888) p. 132 [" Trinidad '—errore !].
S. bottvronoides L^otaud, Ois. Trinkkul, p. 318 (Trinidad).
Noitbcr C'liaiiman nor Andre pollorted this species, and the type of S. trinitatis
Sharpe did also not come from Trinidad, but is of the usual " Orinoco "-make.
The only anthority for its occnrrence on the island i.s Leotand, who says tliat it is
fonnd in company with S. jninuta, but less common.
I cannot see any reason for not accei)ting Lesson's term hoiaronuks. The
description answers well to the species with uniform black pilenm (cfr. " tete et cou
noir "), no mention being made of a white crown stripe. Even if Lesson's name
should be rejected, then Fiusch's designation lessoni would have precedence over
trinitatis Sharpe.
S. bourronides is certainly only a straggler from the mainland, but an adult
male, collected by Leotand, on Trinidad, is preserved in the Paris Museum.
As I will show in another paper, S. ocellatn and S. amazonica arc identical
with S. bouvronides, which, however, apjiears to be specifically distinct from
S. lincola, both species being found together at various localities. For want of
material, these birds are very unsatisfactorily treated of in the Catalogue of Birds.
'i. Carduelis phoeniceus Bonap.
Erroneously recorded from Trinidad in the Cat. Birds, xii. p. 166. The Tring
Museum received a large series of both sexes from the vicinity of Cumaiui.
The species is not known to occur elsewhere, unless C. grajiudensis Lafr. of New
Granada be the same.
7. Mionectes olivaceus Lawr.
Elnnia strkilicfiUis Wotaud, Ois. TiiiiidiiJ, p. 238 (Trinidad).
Mionectes olivaceus Chapman, Bull. Amer. Miis. vi. (1894) p. 38.
The claims of this species to be included in the avifauna of Trinidad rest
upon the single specimen recorded by Leotaud. It has been examined and
identified by Chapman. Li the Tring Museum there is a good scries of M. o/iraeciis
from Cumaui'i.
8. Pachyrhamphus albogriseus Scl.
Taylor, Ibis 18i;4. p. 87 (" Trinidad ").
There can be no doubt that the bird obtained by Taylor has been wrongly
identified as F. albogriseus. This species occurs only in the Andes of C'olombia
and Ecuador, and its occurrence on Trinidad is simply impossible. 1 suppose the
specimen really belonged to P. ru/us (Bodd.) (= cincreus auct.), of which there is
an adult male from Cumana in the Tring Jluseum. In the Catalogue of Birds
two 6 ad. of this species are recorded from Trinidad, but they are most probably
so-called "Orinoco" skins, which, in many collections, are labelled "Trinidad."
Neither C!hapmau nor Ai.dn' met with any l'ar/,i/r/aimp//u.f of this grou|i, hence
the occurrence of B. ru/us remains to Ijc confirmed.
( ;J9 )
0. Dendrocolaptes altirostris Leotaud.
0/s. Tnnhhnl, p. IOC.
The description of tbis species, as given hy Leotand, agrees well with Dendrorms
susiz/'raiii susiirraiis (Jard.), so far as c'oloration is concerned, but the bill is said
to be " blanc de corne." I do not know what to make of this bird ; I can hardly
believe, however, that there is a species of Dendrocolaptes on Trinidad, not found
again since Ltotand's time.
As the type is now in the collectiim of the Boston Society of Natnral History,
the matter could easily be settled by our American fellow-workers.
10. Dendroplex pious (Gm.).
Although Leotaud says it is common, it has not been obtained by any of the
recent collectors.
n. Cercomacra maculicaudis (i^cl.) [= nigricans Scl.].
There cannot be the slightest doubt that this species has been wrongly
assigned to Trinidad. It occurs only on the western slopes of the Andes in Ecuador
and C'olombia, extending northwards to Panama.
1:^. Campylopterus ensipennis (Swains.).
In the Catalogue of Birds xvi. p. 290, and again in the Tierreich, Trochilid.
p. 31, this species is recorded from Trinidad, evidently on the strength of some
specimens in the British Museum labelled " Trinidad." These are skins of the
so-called " Orinoco "-make, and the occurrence of C. ensipennis on the island is not
yet ]iroved, although it is common both on the Paria coast and on Tobago.
13. Agyrtria albiventris (Less.).
The two examples in the British Museum from " Trinidad " are also so-called
"Orinoco" skins (cfr. Salvin, Cat. B. xvi. p. 186: Agyrtria riridissima). As yet
there is no proof that the species ever occurs in Trinidad, nor is it found on the
Paria coast.
14. Chrysuronia oenone oenone (Less.).
This species has without doubt been wrongly indicated as occurring in Trinidad,
although Lesson described it from that island. The examples in the British
Bluseum labelled " Trinidad " are again so-called " Orinoco " skins.
15. Thalurania refulgens Gould.
In nearly all works on Humming-birds we find Trinidad as the only locality
mentioned for this species, but as neither Leotaud nor any of the more recent
collectors ever met with it there, we may safely exclude it from the ornis of the
island. The Triug Museum received a very large number of skins from the state of
Cumand, N.E. Venezuela, and this is the only exact place known hitherto.
10. Chaetocercus jourdani (Bourc).
Calnlhornr enicuniR (neo Vieillot !j Luotaud, Ois. Trinidad, p. 143 (Trinidad).
Leotaud's bird belonged without doubt to the above species, as may easily be
seen from the description of the tail : " rocttrices noires ; dn roux snr la barbe
( «' )
interne de la base an milieu . . . ; la pins ezterieure est conrte et finit en
pointe."
This can never refer to CalUphlox ametlii/stina with which Chapman and Salvin
identified Leotand's description.
Besides Leotand's, we have no other record of this species occnrring on
Trinidad. The only specimens with exact localities I have seen are those in Tring
(1 c?, 1 ?), which were collected near CumanA b_v one of Mr. Andre's men.
IT. Chaetura spinicauda (Temm.).
Chapman, BiiV. Amrr. Mus. vi. p. rjS (Trinidad).
I have not seen any specimens from Trinidad, and I jj^reatly donLt its occnrrence
there. It is not obvions from Ciiapraan's account whetlier lie got an example of
the species, and its claims to be included in the ftiuna of Trinidad cannot be
considered as established.
IS. Cypseloides rutilus (Vieill.).
Neither Chapman nor Andre's collectors ever met with this remarkable bird,
bnt according to the former ornithologist, there is one specimen in Leotand's
collection. The French naturalist, however, leaves it an ojten question whether it
breeds on the island or not, and after all it may be only a straggler there.
19. Chordeiles acutipennis acutipennis (Bodd.).
Althongh Leotand (p. 7(1) says that this species is the most common of all
Goatsuckers, it has not been observed either by Chapman or Andre. From his
observations it is evidently only a winter visitor.
20. Podager nacunda (Vieill.).
According to Leotaud (]). 79) it is a common winter visitor between the months
of July and October.
21. Chrysoptilus punctigula (Bodd.).
Erroneously recorded from Trinidad in the Cat. Birds xviii. p. \\C^. The
species occurs only in Cayenne and Surinam. On the Orinoco River and in
Veneznela, near Valencia, a nearly allied form, C. p. puKCtipcctus Cab. & Heine,
takes its place.
22. Coccyzus pumilus Strickl.
This species, although descrilied as coming from " Trinidad," is now ascertained
to be only an inhabitant of the upper Orinoco valley and the eastern slopes of the
Andes in Colombia. I have seen a large number of Bogota skins. Tiic original
locality was doubtless erroneous.
23. Urochroma hueti (Temm.).
According to Leotaud (p. 332) this species, " est de passage ici, et encore
ne vient-elle nons visiter qn'5,de long intervalles." I have never seen a specimen
from Trinidad.
( «1 )
NEW BEEPANULIDAE, THYBIDIDAE, TJRANIIDAE, AND
GE03IETIIIDAE, FROM ERITISH NEW GUINEA.
BY W. WARREN, M.A., F.E.S.
ri'^HE insects were collected by A. S. Meek at the Aiigabiuiga River, an udlnent
-L of the St. Joseph River, British New Gninea, at an elevation of OUOLI ft.
and upwards, from November 19U4 to February 1905. It is a marvellous
collection, the number of new species being beyond all expectation.
Family DREPANULIDAE.
1. Callidrepana discipunctata spec. uov.
Forewing : rather deep yellow ; from two-thirds of inner margin a band of
three lines runs straight and oblique to below vein 8 ; the central line broadly
brown, the inner and outer slender, the pale interval before the outer slightly
lustrous ; below 8 the inner and central lines are retracted to costa and the outer
bent outwards ; dark spots on the veins represent the submarginal line ; ape.\ and
fringe above it brownish fulvous ; cell-mark formed of two brown-edged aunuli.
Ilindicing : with the band central, but not reaching above vein 4 ; costal area
paler yellow.
Underside fulvous yellow, paler towards inner margins ; both wings with a
ronud brown cell-spot, and traces of a straight brown outer line, running from
apex in forewing, and from three-fourths of costa in hindwing, hardly reaching
below middle.
Thorax, abdomen, and legs like wings ; upper half of face black, lower half
yellow.
Expanse of wings : 50 mm.
1 ?.
2. Callidrepana superba si)ec. nov.
Forewimf : jjale nnspeckled ochreous; from three-fifths of inner margin an
oblique olive-brown band runs straight to below vein 7 ; this band consists of a dark
olive fine line on the inner edge, separated by a pale interval from a broader and paler
olive line, which is edged externally by a somewhat diffuse line of silvery scales ;
before vein 7 the inner dark line is widely retracted to costa and becomes obsolete ;
the outer line develops into a wide shallow red-brown costal blotch with jialer
centre, the lustrous line forming a large bright spot below it; submarginal line
represented by a series of olive-brown wedge-shaped marks on the veins, the
whole marginal .area, especially towards apex, slightly deeper tinged ; apex and
fringe dark olive-brown, (lie fringe becoming jialer towards anal angle ; cell-sjiut
double, formed of a bent line of silvery scales on the upper arm of discocellular,
and a spot at the lower end.
Ilindwintj : with the olive band central, not reaching above vein G; the
submarginal spots more distinct, the marginal area beyond them distinctly olive-
brown.
( ea )
Underside with the band nmber-brown, very thick on hindwiug, not reaching
inner margin in either wing, filling up the apex of forewing, the inner edge
forming a pale-centred loop on costa followed by a round i)ale sjwt ; suhmnrginal
line vi.silde only in forewing and there not below vein 2 ; a, brown marginal
line on forewing to vein 2, broader on hiudwing ; fringe brown on forewin;:, jiale
on hindwing ; costa of forewing towards base yellowish.
Head, thorax, abdomen, and legs ochreons ; upper half of ftice brown ;
forelegs tinged with l)rown.
Expanse of wings : 00 mm.
1 ?.
Hyalospectra gen. nov.
ForewitH/: costa curved ; ajjcx dejiressed, bluntly rounded ; hindmargin bluntly
elbowed in middle, concave above, oblique below.
Uindw'nuj : bluntly angled at middle of hindmargin.
Antennae pectinated in ?, with stiff parallel pectinations to two-thirds, ai)ical
third simple ; palpi quite short ; tongue and frenulum present ; hiudtibiae with
two spurs only.
Neuration : forewing, cell more than half as long as wing ; upper and lower
third of discocellnlar vertical, middle oblique ; first median uervule at one-half,
second at four-fifths, both curved downwards at origin ; lower radial from the
lower angle of discocellnlar; 6 stalked with 7, 8, 9, from the depressed end of
.subcostal; lO and 11 close together from cell, 10 anastomosing wath 8, 0:
hindwing, 7 from long before end of cell, shortly anastomosing with 8.
^Vings sparsely scaled, with large hyaline discal patches.
Type : Ilijalospectra grisca spec. nov.
3. Hyalospectra grisea spec. nov.
Forewing ; hyaline ; the basal area, the costal area above subcostal vein, and
the hindmarginal border clothed with sparse grey scales ; at about one-third from
base a dark paler-edged lunulate-dcutatc line, toothed outwards on the veins ;
in the grey marginal area a lunulatc-dentate dark line, running out from costa
at three-fourths and below middle forming the limit of the hyaline space ; this
space is roughly four-lobed, comprising the outer half of cell and the inner half
of all the intervals between veins 2 and 8 ; a round hyaline marginal spot between
3 and 4, and the marginal area below apex semibyaline ; a dark grey marginal
line, spotted on the veins ; fringe pale grey.
Ilinchciny: grey, with the base, an angled band beyond middle, and the
marginal spot between 3 and 4 hyaline.
Underside hyaline, the marginal area grey, the l)asal only showing through.
Head, thorax, and abdomen dark grey, ftice and antennae jialer.
Exi)ause of wings : 30 mm.
1 ?.
4. Oreta patiens spec. nov.
i. Forewinij: dark fawn-grey, with numerous transverse darker striae and
si)eckles ; costa with two blackish blotches, one nearly at middle, the other at
two-thirds ; from the first a diffuse dark central shade arises, which is bent at
cud of cell ; from the second a fine outer line arises, angled at vein C then
( fi3 )
oblique inwards (in the type S this line is very indefinite) ; discal mark a line
of faint bUiisli shining scales ; along vein 5 a pale chestnut-brown streak extends
to an oblique subiuarginul whitish mark ; a dark blotch at anal angle ; this blotch,
as well as the apical area, is stndded with flaky white scales ; fringe rufous
brown, tipped in places with shining white scales.
Hindtving : redder, with a broad darker central band, and with dark s])ccks
between the veins ; fringe deep red-brown.
Underside deep dull reddish, coarsely black-spotted between the veins ; the
black outer line on forewing and central on hindwiug well marked, as well as
the white submarginal streak.
Head, thorax, and abdomen dark red-brown ; underneath and legs reddish.
¥. Much larger and paler, with scarcely any red tinge ; the outer line
on forewing black and well defined, produced as a central line across hindwing ;
underside pale whitish brown, with coarse black speckling and black thick
oblique line.
Expanse of wings : c? 30 mm. ; ¥ 40 mm.
1 c?,l S.
Family 77/ ) 7? IDT DAE.
5. Addaea maculata spec. nov.
Forewniq : yellowish straw-colour, with some faint pale brownish striae between
the veins ; costal edge brown dotted with pale ; base of cell brown ; a broad brown
band from just bej'ond middle of costa to before middle of inner margin, followed
below middle of wing by a grey-brown blotch reaching anal angle ; traversed
at middle by a narrow darker band, the upper part of which starts faintly from
costa just beyond middle shade ; a series of brown triangular spots along margin ;
fringe straw-colour.
Hindwing : with central shade thick, unaccompanied by a blotch ; in marginal
area one brown and three paler lines visilde.
Underside yellower, with the bands and markings pule red-brown and very
regular.
Head, shoulders, and base of patagia deep brown ; thorax and abdomen
straw-colour ; fifth dorsal segment deep brown, sixth brownish.
Expanse of wings : 26 mm.
1 ?.
0. Banisia quinquelineata sj)ec. nov.
Forctciiig : jiale uchreous, rather glossy; traversed by very fine grey wavy
lines or striatious, and by five brown lines or thin bands, oblique outwards fnim
costa, angled on subcostal vein, then oblique inwards and waved; down the pale
intervals between them a darker central line forming small black patches between
the veins ; a series of large marginal brown spots ; fringe brown.
Hindwing : with only the two inner lines clear ; fringe brown.
Underside with the straw ground-colour paler, and the brown lines and black
speckling much brighter.
Head and forelegs dark brown ; vertex, thorax, and al)domen straw-colour.
Expanse of wings : 34 mm.
3 33.
( M )
7. Brixia quartaria sj)ec. nov.
Forewing : oclircous, nuiformly snffnsed, except along costa, with pale brown :
crossed b)' four or five darker bands more or less interlacing laterally, two
antemcdian and two postmedian being plainer, and brown-edged on costa ; tlie
paler intervals with faint traversing lines ; fringe pale.
Himhciiii/ : with the costal area above median vein like forewing ; below with
three broad deep brown bands, basal, median, and marginal, the intervals whitish
with a i)ale line in middle ; fringe ochrcons, darker at base, especially towards
anal angle.
Underside of forewing snfFnscdly pale brown, sluiwing no markings ; of
liindwing with the pale and dark markings amplified and very distinct.
Head, thorax, and abdomen ochrcons, sometimes tinged with jiale brown ;
dorsum marked with brown ; face, palpi, j)ectus, legs, and underside of abdomen
deep brown.
E.xpanse of wings : S 22 mm. ; ? L'6 mm.
2 cJcJ.l ?.
8. Letcheiia satelles spec. nov. and ab. coeca nov.
Foretcinq : fawu-colonr in c?, with obscnre dark transverse striae and
reticulations ; dull brick-red in ? , with the striae developed into dull lines and
bands ; costal edge with pale linear intervals ; between veins 2 and 3 at middle
of wing a large hj'aline patch, generally elongate and oblong, sometimes shortened
and nearly round, in one case divided into two by a vertical middle line; with
two small spots one above the other in the interval beneath, sometimes confluent ;
fringe, dark brownish or reddish grey. In 1 y , ab. coeca, the hyaline spots are
entirely absent.
IJindwi/u/ : like forewing, bnt without hyaline spots ; fringe with apical half
white ; inner margin and fringe pinkish ochreous.
Underside of forewing with a diffnse brownish postmedian cloud and sub-
marginal spot ; hindwing pink ; inner margin of both wings pale.
Thorax and abdomen concolorous with wings; head generally darker ; forelegs
dark brown, with jiale joints ; hindleg with tuft of hair internally fuscous.
Expanse of wings : 4U mm.
4 (?c?,3 ? ?.
Microbelia gen. nov.
Forewing : elongate ; costa straight, convex only before apex, which is slightly
Iirominent ; hindraargin somewhat bulged at middle, very obliqne below.
liindwing : narrow, oblong ; the apex rounded to a slight elbow at 7 ; hind-
margin bulged at middle and inbent on each side ; inner margin very short.
Paljii obliiiuely porrect upwards; second segment thick, rough-haired, terminal
short; antennae bimeliatc, thick ; legs long ; neuration siinjile.
ItitlVrs from Baninia in having the termiiuil segment of jtalpi siiort.
Underside of forewing without metallic or hairlikc scales.
Type : Microbelia curvinota spec. nov.
To this group belong intimalis Moore, cotnpunctalis Warr., and ulterior ^\'ar^.
( 65 )
9. Microbelia curvinota sjiec. nov.
Foir/rl/ii/ : ophreons, witli five snlir|uadrate lirowii costal jiatolios, tlie commence-
ment of fasciae which in and lielow tlie cell hoconic black-bmwn ; costal edije dotted
light and dark ; the pale intervals lietween the hands each traversed by a brown line,
that beyond the second band palest and most prominent ; the third and fourth bauds
unite to form a Y-sha])ed mark ; the fifth becomes black and is bent round at right
angles to margin at vein 4 : veins towards margin fnlvons, the intervals with
vertical brown-black striae ; fringe ochreous, chequered with black beyond veins
below middle, and at vein 7.
Hindwing : with basal third pale straw-colour or whitish, with a brown-black
band near base and limited by another running from above anal angle to niidwing,
followed by a dark fulvous mark at anal angle ; rest of wing paler, as in forewing.
Underside with all the tints, both pale and dark, much brighter and clearer.
Head, thorax, and abdomen, fulvous ochreous ; metathorax and anal half of
dorsum black-brown ; palpi and legs ochreous.
? with the markings fulvous brown instead of black, except the ))pnt mark to
middle of hindmargin ; the jiale costal s[iaces dotted black and white ; hindwing
with base more largely pale, the basal band being slender; head, thorax, and
abdomen rich fulvous.
Expanse of wings : c? 2il mm. ; ? 20 mm.
1 (?, 1 ?.
li'. Pharambara basalis sjiec. nov.
Forewing : whitish ochreous with pale red-brown reticulations ; the markings
dark chestnut ; basal two-fifths deep chestnut edged by a vertical line ; an
interrupted postmedian fascia formed by a dependent bar from costa and a bifurcate
mark from before anal angle ; hindmargin narrowly chestnut-brown, with obliipie
Y-shaped streaks running into it ; costal area brown, with pale rounded intervals
dotted with dark on costal edge ; all the dark markings more or less connected
laterally along subcostal vein and inner margin ; in the paler spaces the veins
are jiale reddish ; fringe pale ochreons with brown mottling.
Himhrimj : with three brown bands near base running into inner margin, the
third broad and double ; the hindmargin red-brown, with two mcn-e, partially double,
streaks running into it, with single lines between.
Underside the same, with the dark brown markings all clearer, and with some
bluish silvery scales along subcostal area.
Head and thorax grey-brown ; abdomen brown, the anal tuft of cf ochreous ;
face deep brown ; i)alpi and forelegs deep brown with the joints pale; antennae
annnlated lirown and ochreous.
Expanse of wings : S 18 mm.; ? 22 mm.
1 ^, 1 ?.
11. Siculodes fumiceps spec. nov.
Foreiviny : white, traversed by numerous contorted bronzy grey lines and
reticulations, which cut up the white ground into rounded siwts or blotches of irregular
size and shape : costal edge minutely dotted with dark ; subcostal vein with white
spots alternating with bronzy-grey intervals ; some short oblique lines along hind-
margin running out grev into the white fringe.
5
( <i6 )
Ilinclwing : similar ; many of the spots appearing as if embossed, as a few of
those in tlie forewiug also do.
Underside of forewiiig similar; of liindwint; wliite with tlie markings showing
tlmingh; costa df forewing nnilormly dotted tjjaclc and white, with a yellowish streak
below it.
Head ami jirothorax smoky grey ; metathorax and abdomen wliite.
Expanse of wings : 2ii mm.
1 S.
12. Siculodes nigrithorax spec. nov.
Forewing : glossy white, with very indistinct grey-brown reticnlations, excejit
towards hindmargiu, where they become dark brown and complicated ; a fine black-
brown marginal line swollen into spots at the veins, which project into the white
fringe ; a large brown spot at base of discoeellular, and another below it on snb-
median vein, forked obliquely towards inner margin ; costal streak dark-brown ;
costal edge dotted with orange ; subcostal vein with white spots.
Ilimhciiiq : with a hyaline central space ; the rest of the wing with bosses of
white scales arranged in rows between the veins; a black spot on inner margin
at middle, and some dark markings towards hindmargin; marginal line as in
forewing.
Underside white, glossy, the markings as above ; costa of forewing gilded
yellow, speckled with black, and with a deep yellow line running into apex.
Head and prothorax black-brown ; metathorax, apical half of patagia, and
abdomen white; dorsal segments, except basal, banded with brown; abdomen
beneath, pectus, and legs silvery white; foretarsi black, with white joints.
Expanse of wings : 2(3 mm.
2 ??.
13. Siculodes subauratalis spec nov.
Foreicinq : snow-white ; the costa finely dotted with dark ; the base with thick
purplish si)Ots and striae between veins ; the marginal area with some longer and
more angular marks as well ; the space from apex to end of cell and the middle of
inner margin remaining imre white ; large dark marginal spots at the vein-ends below
middle ; fringe white.
Ilimlwinq : with thick irregular markings except along costal area; marginal
spots at the vein-ends throughout.
Underside like upper, btit most of the markings of hindwing and many on
forewing pale instead of dark; costa of forewing broadly gilded yellow; costa of
both wings black-dotted.
Thorax and abdomen white : dorsum with black spots ; the head and jjrothoras
are discoloured, but appear to have been grey; foretarsi black, with joints white.
Expanse of wings : 20 mm.
1 6.
Family URANIIDAE.
Subfamily MICRONIINAE.
14. Cyphura pardata spec. nov.
Vorewiiiq : cream-white ; the costal area above subcostal vein crossed by stout
black striae, those in the apical portion more variable in shape and size, sometimes
forniii)'^' a iVw long streaks, soniclinies more numi-rons anil short, reaching as far as
( 67 )
vein 4 ; three olive-fulvous, brown-edged, fnnuel-shapod bands cross the wing from
lOsta ; tlie inner edge of the first from base of costa to before middle of inner
margin, the outer from one-sixth of costa to tliree-fiftlis of inner margin ; inner edge
of tlie second from one-fonrth of costa to three-fonrtiis of inner margin, the outer from
middle of costa to four-fifths of inner margin ; the inner edge of the third from
three-fifths of costa to anal angle, the outer edge not defined by a line but
only marked by a darker shade, which joins the inner edge at anal angle ; a broad
velvety black sulnnarginal line nut reaching costa and ending in a point before anal
angle ; fringe with basal half black, outer half whitish.
Ilindwing : white, with a bright fulvous suffusion in the outer half of wing from
vein 4 to anal angle, containing five black submarginal spots, a hirge one on each
side of vein 3, a smaller one on each side of the submedian fold and a smaller one
still below vein I ; a broad abbreviated submarginal black stripe from vein 5 to 2, and
a shorter fine one close before margin from 6 to 4 ; fringe white from apex to
the end of tooth, thence to au.al angle fulvous, tipped with grey and with a velvety
black basal line interrupted at the veins : fringe of inner margin fulvous in outer half,
white in basal half of wing.
Underside of forewing white with two broad blackish oblique streaks, cor-
responding to the two outer dark lines of upperside, the inner not reaching below
middle; some dark brown sjiccks and striae on costa before apex, and a grey linear
shade before hiudmargin from apex to middle ; fringe grey-brown, the basal half
darker; hindwiug with the fnlvous area more restricted; the submarginal black
stripe from 5 to 2 broader and continued to anal angle, including the three smaller
spots.
Palpi white, with second segment black above ; face white with a black spot
above; vertex, antennae, and collar white; shoulders and patagia pale fulvous;
thorax and abdomen white with a broad black dorsal stripe : abdomen below and legs
white ; forelegs with a black line in front.
Expanse of wings : 56 — 65 mm.
0 c?c?,2 ? ?.
The largest examjjle has an additional short brown middle line on hindn-ing
from vein 4 bent round to inner margin above anal angle.
15. Stesichora pura spec. nov.
Fore- and hindwintia, above and below, and all the parts of the liody pure white,
except the upper half nf face, the tips of the palpi, and the front of the forelegs,
which are blackish.
Expanse of wings : 4n mm.
1 S.
10. Stesichora strigifera spec. nov. and ab. crassistriga nov.
Foreioiiiy : white, covered with irregular pale brown strigae ; those on costa
and subcostal vein fine and black ; marginal line pale brown ; fringe white with
a faint brownish tinge ; besides the striae there are sometimes traceable three
more or less continuous pale grey-brown linos across wing, antemedian, median,
and postmedian ; of these the median is generally the most prominent, and in
certain cases developed into a broad conspicuous band, ab. crassistrii/a.
IJiiuhving: with the strigae confined to the marginal area; the nxdiiin anil
( (i8 )
liostnieiliiiii liaiuls fjcnerally well miirked, the latter sometiiucs angiilated below
niiddlo : inargiiial line fine, hlaclc-l)ro\vn, interrupted at the slight tooth, which
(X)iitains an oval black spot ; frinj;i' wliite, tinircd with brnwii. In tlie !ilierrafiiin
tlic fringes are darker in both wings.
Underside nniforiuly white.
Head, thorax, abdomen, and legs :ill wliite; face and palpi black, whitish
below ; forelegs dark grey in I'ront.
E.xpanse of wings : c?, 4i< miii. ; ?, .")'-i niiu.
4 t? J, 2 ? ?, one of these being the aberration cnissistriyu.
SuBiWMiLY EPirLEMlNAE.
IT. Dirades corrasa sjiec. nov.
Fmrtring : pale grey, freckled with darker; a dark rell-spot followed by n
larger dark spot above vein 7, edged with ferrnginous externally ; a row of snb-
niarginal black Inuules between the veins from apex to vein 2 ; a dark marginal
line ; fringe grey with a pale basal line.
Ilimhininj : dark brown above vein 4 ; white and withont markings, exeejit on
vein 1, below it; costa with a silvery white blotch from liase to apex, where it
is narrowed off to a point ; across the middle a donble dark brown line with
yellowish scales between ; along hindmargin from vein 7 to 4 a leaden, blue band
edged inwardly by black white-edged Innnles ; fringe brown to vein 4, wliite
below it.
Thorax and abdomen pale grey ; vertex white; lace and paljii black.
Expanse of wings : 22 mm.
1 S.
Hindwing with a tooth at vein 7, and the inner margin below vein 3 lobed ; an
elongate hyaline fovea near base of wing beneath the median vein.
18. Dirades niveicosta spec. nov.
Foreiring : cinereons grey, striated with darker ; the lines fnscons, interrnpted ;
first cnrved, at one-third; second at two-thirds, ontcnrved below middle, where
it is thickened ; a cloudy dark cell-spot ; an apical short streak of three black
lininles; costal edge distinctly speckled black and white ; fringe grey.
Iliiulwiiig : similar ; but the costa with a bro:ul silvery white streak from
close to base to outer line, where it is bluntly rounded ; the inner edge of this
white streak is jierfectly straight, and contains three black spots, one at base, a
small one at middle, and a third at the end, these last forming parts of the
inner and outer lines ; a row of leaden-blue dark-edged lunules along margin from
vein 7 to 2.
Underside of forewing grey, of hindwing whitish ; costa of hindwing with two
tnfts of dark scales, a large one at base and smaller one at apex.
Thorax and abdomen grey like wings ; vertex white ; face, pal])i, and forelegs
black.
Expanse of wings : 22 mm.
2 ? ?.
Near JK (itiibaaiti Warr. iVom Fergusson Island, but the white streak of
hindwing is not jagged below.
10. Diradopsis alberta spec. nov.
FoirwiiKj : pale grej', cloiuleJ and speckled with dark grey ; costa miuutely
dotted l)lack and white ; lines dark with paler edge ; first ontcurved from one-third
of costa to median vein, second from three-fifths of costa to vein 4, both interrnpted
below middle, bnt really joining the edges of a dark brown blotch on inner margin
beyond middle ; this blotch is narrower and rounded at summit in the S, broader
and flattened in the ? ; a row of black snbmarginal spots from apex to anal
angle ; a dark grey triangular cloud on costa beyond outer line, and a slighter one
at anal angle : fringe brown.
lliiulwimj : browner; outer line in i nearly straight from middle of costa to
vein 4, where it is bluntly angled and becomes obsolete, in the ? curved above
middle, straight below, with a lilnnt tooth on vein 4, dark l)rown edged with pale ;
in the ? lielow 4 the marginal space is whitish grey nearest the line, the rest dark
grey with snbmarginal dark spots between veins ; fringe brown with pale tips ; in
the S the submedian interval is whitish, without scaling ; in the ¥ whitish speckled
with black; in both sexes the veins of hindwing are sometimes pale.
Underside dull brownish with Idack s]>eckling, whiter towards base of liindwing.
Thorax and abdomen grey; vertex pale grey ; face, palpi, and forelegs black.
Expanse of wings : c?, 22 mm. ; ? , 23 mm.
4 cJd',4 ??.
"-'II. Epiplema canibrunnea spec. nov.
Forewiuij : liver-brown, dusted at base, along costa, and before hindmargin with
bluish grey scales ; costal edge ochreous with slight brown dots ; the lines finely
jiale ; first from one-fourth of costa, obli(]uely curved outwards to median, then
(ibli(pie and straight inwards ; second line straight from just beyond middle of costa
to l)eyond middle of inner margin ; the interval lietween the lines deeper brown,
forming a fascia, tlie veins across it paler ; a pale dark-edged kuuilate line from
near apex, curved to hindmargin at submedian fold, enclosing a darker brown
marginal area widest beyond cell, edged inwardly with bluish grey; a fine bluish
grey marginal line ; fringe broadly brown at base, [laler and mottled towards tips.
HIndtriiuj : with the two lines curved and parallel ; the lunulate submarginal
line entire from ai)ex to anal angle, enclosing a dull leaden-blue band; fringe as in
forewing.
Underside of forewing purjilish grey, with darker speckling ; of hindwing white
with purplish speckles along hindmargin ; fringes grey chequered with purplish.
Vertex white ; shoulders and jmtagia pale bluish grey, powdered with dark
scales; abdomen brown with jiale rings; anal tuft ochreous; face, pulpi, and
forelegs brown-black ; antennae ochreous.
Expanse of wings : 17 mm.
1 S.
Hindmargin of forewing entire, with sligiit indentation beyond cell ; of hindwing
with slight blunt teeth at 4 and T ; costa of hindwing fringed with hair ; antennae
with clavate teeth.
21. Epiplema casbiata spec. nov.
Forewing : ochreous or whitish grey, densely covered with fine short striae ;
cell-spot blackish, in a cloud of dark striae; the lines accompanied liy shades of
dark striae ; inner line marked by a dark sjiot above and below median vein ; outer
( 70 )
line liy small brown liiiiatc marks between veius at. two-thirds, parallel to hiiitl-
margin ; a black spot before apex above vein 7 ; fringe concolorons.
Ilnidiriiui : cell-sjiot white, followed by a cloud of striae which extends
towards anal angle ; onter Hue faintly expressed, bluntly angled on vein 4 ; an
oval leadeu-blne, somewhat embossed spot, edged with yellowish, on liiMdmargin
below lower tooth.
Underside white, sliglitly discoloured iu forewing, with nuiucrons blackish
striae between the veins; a black spot before lower tooth.
Thorax and abdomen like wings ; vertex whitish ; face and palpi black.
Expanse of wings : 38 mm.
2 ? ?.
Forewing with hindmargin simple ; hindwing slightly toothed at 7, fi, and 4,
the last finer and longer.
22. Epiplema cinereella spec. nov.
Fori'iriiiq : ash-grey with a slaty tinge, and covered with indistinct dark striae ;
costa dotted with dark grey; lines darker, but indistinct, and interrniited ; first
at one-third, bent in middle ; second from two-thirds of costa to three-fourths of
inner margin, inbeut below middle; both phiinest at their extremities; a blackish
marginal blotch from apex to vein 'i, inwardly edged by a black line, bluntly
angled basewards on vein u ; fringe grey.
Hiiidwiiiij : with equally indistinct curved lines ; a dark marginal lunate-edged
shade between teeth.
Underside uniform dull dark grey.
Thorax and abdomen grey ; vertex white ; shonlders whitish grey ; face, jialpi,
and forelegs blackish.
Expanse of wings : ](j mm.
1 ?.
Forewing indented slightly beyond cell ; hindwing with small teeth at 4 and 7 ;
cell of forewing very short; vein .5 from lielow ujjper angle of cell.
23. Epiplema configurata spec. nov.
ForewiiKj : pinkish ochreous, thickly dusted with grey, at liase and along
costa ; the central fascia and anal region below middle and the hindmargin suffused
with black ; lines velvety black ; first at one-third, acutely angled outwards on
median vein, then oblique inwards, broadly edged internally with ferrnginons, the
costal portion above cell almost obsolete ; outer line from three-fifths of costa to
four-fifths of inner margin, forming a small projection outwards on vein 7 and
a prominent blunt one on vein 4, then incurved, and vertical from vein 3, edged
externally finely with ferruginous ; a line of lilack dots from apex, limiting the
black hindmargin, toothed inwards on vein .j ; fringe black-brown.
Ihndwiiig : black ; the lines edged with ferruginous conversely as in forewing ;
the inner at one-third angled outwards on both folds, the outer on vein 4, and
marked by white dots at veins ; the submedian interval diffnsely white with black
speckling, the anal region below vein 3 whitish ; the Innules along margin oliscurely
edged with ferruginous; fringe black.
Underside of forewing smoky fuscous with black striae ; costa whitish with
black dots; inner margin whitish; hindwing dull white, grey-tinged towards costa
and aj)ex ; the fringe brown.
( 71 )
Face, jiali>i (wliich are lung and porrcct), anil forelegs black ; vertex snow-
white ; shoulders and patagia grey-brown ; abdomen whitish, the basal half of
dorsum black, the anal grey ; abdomen beneath and legs white.
Exjiause of wings : 27 mm.
1 ?.
Hindmargin of forewing simple; of hindwing with two lougish teeth at veins 4
and 7, and a shorter one at end of 0.
24. Epiplema dealbata spec. nov.
Forcwimj : glossy stone-grey, iinely dusted with dark scales ; costa marked
with grey; lines starting from costa at two-fifths and three-fifths; first very
obscnre, passing over a linear chestnut-brown discal spot ; second outcurved,
reddish grey with a paler limiting line, incurved at 4, joining the edges of a
large flattened pale-centred chestnut-brown blotch at middle of inner margin ;
a chestnut-brown narrow shade before margin from ajiex to vein 3, broken into
two lunules above vein 6 ; a submarginal grey cloud running into anal angle ;
fringe grey.
IHndiriiKj : inner line straight, chestnut-brown, its middle formed by the
discocellular mark ending in a round blotch between the origin of veins 3 and 4 ;
outer line brown, with paler edge, waved and bent on vein 4, ending in a square
brown blotch between 3 and 4 ; a dark, pale-edged lunular line before teeth ;
Bubmedian interval cream-white without markings, which reappear on vein 1.
Underside white, slightly discoloured in forewing, with a few dark specklings
towards hindmargin.
Vertex, thorax, and abdomen like wings ; face and palpi brown-black.
Expanse of wings : 27 mm.
1 S.
Forewing simjile ; hindwing toothed at 4 and 7.
25. Epiplema despecta spec. nov.
Forcwimj : pale grey, thickly covered with grey-brown striae ; lines dark
brown and fairly distinct; first well curved at one-third; second from just beyond
middle of costa to two-thirds of inner margin, projecting outwards on the two
folds, angularly beyond cell and bluntly on the submedian ; a browner grey
marginal shade from apex to vein 3, prominently projecting inwards between
4 and 6, where it is edged by two black lunules, and above vein 0 by another
black Innule nearer margin ; fringe grey.
IlinihciiKj : a brown line near base; a faint ochreons line along median vein
joined by a faint brown mark on discocellular ; outer line sinuous, brown edged
with i)aler, bluntly angled or rounded on vein 4 ; a leaden-grey shade along margin
from upper to lieluw lower tooth, with a sinuous darker inner edge cut by a whitisli
dash on vein 4, with a whitish edge between 3 and 4.
Underside of forewing dull brownish grey with darker freckling and a dark
shade at base; of hindwing much paler, becoming greyer and freckled only towards
hindmargin.
Vertex, antennae, thorax, and abdomen grey ; face, palpi, and forelegs black.
Expanse of wings : 25 mm.
1 c?.
Hindmargin of forewing entire ; of hindwing toothed at 4 and 7, with a blunt
tooth at G ; submediau fold scaled like the rest of wing.
( T2 )
','(). Epiplema detecta spec. nov.
Foreicinq : as in the last siiecies, pale grey with sliyhtly darker striae; no
distinct lines ; in their places slight diti'iise olive sliades ; a darker marginal space,
not so prominently projecting beyond cell, edged by a single obliiiuely curved black
line, with two dark Inniiles above, one on each side of vein T ; the pale grey fringe
with darker specks at base beyond veins.
WnuUcimi : with a donble brown spot on mtidian vein near base, a distinct
thick brown streak on discoeellular, and no ochreons line along median vein ; outer
line tine, acutely angled outwards on vein 4 ; the submedian interval whitish,
without markings ; the dark marginal sjjace from upper to lower tooth narrower
and with straight inner edge ; three small brown-edged lunules below lower tooth,
with a fine white dash on vein -t.
Underside as in desjjecta, as well as the parts of the body.
Expanse of wings : 23 mm.
1 ¥.
Hindmargin of hindwiug between teeth straight, with no tooth at vein 6.
■-'7. Epiplema eupeplodes spec. nov.
Forewlnij : ]iale ]iearl-grey towards base and ahmg costa, speckled with brown
and dark grey, with a difl'nse brown cloud on hindmargin below middle ; the veins
throughout marked paler; costa finely brown ; the lines red-brown, darker on casta;
first at one-third, oblique outwards, sharply angled on subcostal, then sinuous
inwards, marked by brown spots on tlie veins and fold ; outer line from before
two-thirds, obliijue outwards, and bent on the subcostal, then waved to inner margin
at two-thirds, jiarallel to inner line; a l)rown lunate-edged streak along excision,
with two black spots above it ; fringe brown, exce])t at apex; cell-spnt formed of
two black semi-confluent spots.
Hiixhcing : with costal margin, the cell, and central space to inner margin
whitish; the veins yellowy three short black curved lines with yellow between
them on inner margin near base ; an outer pale brown-bordered line acutely angular
above vein 4 ; two black spots on discoeellular ; marginal area lilac-grey, with
white black-edged submarginal spots between 4 and G, and a dove-coloured blotch
below 4 containing a black dot.
Underside of forewing pale brown, coarsely black-speckled, jiale grey towards
apex and along inner margin ; cell-spot black ; hindwing white, with black costal
sj)ots, a black discal mark, and scatlered black speckles ; a brown-black marginal
border, broad at costa, narrowing downwards to vein 2, containing white marginal
jiatches.
Vertex, thorax, and abdomen pale jiearl-grey, the dorsum with slight black
and yellow markings; face, palpi, and forelegs deep brown; antennae and lilkt
yellowish.
Expanse of wings : 20 mm.
2 cJcJ.
Hindmargin of forewing excised between 7 and 4, straight and oblicpie below,
the apex depressed ; of hindwiug shortly toothed throughout, with a long slender
tooth at vein 4 and slightly excised above it; anteunae with distinct clavate teeth.
( 73 )
28. Epiplema falcigera spec. uov.
ForawiiKj : lilac-grey, speckled with black, most thickly along inner margin;
first line indicated by a brown oblii[ue streak from costa at one-third, second by a
brown cloud at two-thirds ; patches of black scales between veins before hiud-
margiii below middle, and a larger patch on inner margin at two-thirds ; a black
discal spot ; fringe ferrnginons.
llui(hriii<i : with the costal half sutfused with wood-brown, the whole irregu-
larly spotted and striated with dark ; a black spot before margin below lower
tooth ; fringe brown.
Underside of forewing dull smoky brown with black strigae, the costal area
pale grey ; hindwing i>ale yellowish grey, coarsely black-speckled, with a black
discal dot and slight submarginal shade.
Face, Jialpi, and forelegs lilack ; vertex and shoulders jiale pearl-grey : thora.x
brownish grey ; abdomen paler.
E.xpanse of wings : 35 mm.
1 ?.
Forewing with falcate apex, and a deep excision between it and vein 4, straight
and obliijue below ; hindwing crenulate, with longer teeth at veins 4 and 7 ;
antennae with short clavate teeth.
29. Epiplema flexifascia spec. nov.
FcircwiiKj : jiale grey, with dark grey suffusion and strigae, sometimes with
a reddish tint ; lines thick, blackish : first at one-fifth, bent in cell, then oblique,
the uj)per arm almost obsolete ; second from quite two-thirds of costa, oblique
inwards to median, then curved outwards to three-fourths of inner margin : the
area from base to this line darker grey, especially in the ?, and with dark striae
along inner margin ; outer line curved from apex to submediau fold, enclosing a
semi-oval marginal space of dark grey ; tlie area between secDiid and tliiril lines
l)ale, with slight striations and a central dark cloud lielow middle running into anal
angle ; fringe dark grey, pale above anal angle.
Uuidwiny : dark brown ; a deeper brown, pale-edged outer line, bluntly rounded
in middle, not angled, towards hindmargin ; basal line abbreviated, confined to
costal half; a dark brown thick line along median vein and vein 4, interruj)ted
by an oblique white mark, where the dark discocellular line joins it ; a dark brown
lunular line before margin from upper to below lower tooth, on each side of which
it is swollen, i)receded by a white spot below vein 6 ; fringe brown, with a paler
base ; space along the fold iVom base pale in both sexes.
Underside yellowish ochreous in forewing and costal hall' of hindwing, with
dark brown strigae, lines, and suffusion ; llie veins also dark ; inner-marginal half
of hindwing whiter.
Head, thorax, and abdomen pale or dark grey ; face, palpi, and forelegs black ;
vertex white ; collar black ; antennae ferruginous, with thick, rather long, clavate
teeth in the cf , simply lamellate in ? ; anal tuft in S pale ochreous.
Expanse of wings : cj 22 mm. ; ¥ 24 mm.
2 (?(?,2 ? ¥.
Hindmargin of forewing entire, but I'aiutly indented beyond cell ; of hindwing
with two small teeth at veins 4 and 7, concave between, with a blunt tooth at
vein 6.
The ? , as a rule, is darker than the $.
( 74)
3n. Epiplema innocens spec. nov.
Forpwing : whitish, densely striated with jiale lilac-groy ; the lines a little
darker, but obscurely marked aud interrupted ; first at two-filths, curved ; second
from three-fifths of costa to three-fourths of inner margin, slightly outciirved above;
a slightly darker marginal sjjace from apex to vein 3, edged inwardly liy a dark
waved line, prominent on vein 'i : fringe pale grey, beyond some faint whitish dots
at the ends of veins.
lUndwing : first line curved close to base ; second forming u long blunt
projecting beak on vein 4 ; some dark grey marginal lunulcs from upper to lower
tooth.
Underside lilac-whitish ; forewing with a faint grey tinge.
Thorax and abdomen grey like wings ; vertex and shoulders whitish ; face
dark brown above, whitish at base.
Exi)anse of wings : 17 mm.
1 S.
Forewing with hindmargin entire ; hindwing incurved between 4 and 7, with
minute blunt teeth at their ends,
31. Epiplema planimargo spec. nov.
ForririiH] : dull pur]>lish grey, this tint being formed by darker grey striae on
a paler ground : lines fine and distinct, olive-brown ; first curved, from i[uite one-
third of costa to fully middle of inner margin : second from two-thirds of costa to
three-fourths of inner margin, forming a beak outwards on vein 4, then incurved
and parallel to inner line ; fringe dark grey, the margin before it also ditfnsely
dark.
JlhuhciM/ : with tiie outer line only distinct, a little curved.
Underside uniform jiale lilac-grey, slightly darker in forewing.
Head, thorax, and abdomen concolorous with wings ; vertex whitish ; face and
palpi dark brown.
Expanse of wings : ^1 mm.
1 ?.
Margins of both wings simjde ; hindwing with a scarcely jierceiitible bend at
veins 4 and 7.
32. Epiplema taminata spec. nov.
Forciciii;/ : dull white, with a lilnish tint ; tiie lower outer half of wing suffused
with pale brownisji ; c(jsta finely dotted black and white ; the whole wing with a
few dark striae ; lines purjilish black, interrupted ; first at about one-third, angled
bluntly on median vein ; second from a dark costal mark at two-thirds, bent in
below vein 4 and intcrrnpted throughout ; the central space below vein (i filled
in with brown striae ; a sulimarginal line of black spots from apex to vein 2, the
margin beyond it purplish; fringe black-cluMiuered at base, pale towards tips, with
a pale basal line.
Iluiilwiny : white, brown-sj)eckled ; the two lines curved, blackish, interrupted,
the outer angled on vein 4 ; two distinct black sjjots at the ends of discocellular ;
a brown black-edged marginal shade, cut by a pale line before lower tooth.
Underside of forewing washed with pale brown ; the submarginal line marked ;
the fringe chequered black and jtale ; hindwing white, with a few dark freckles ;
two black discal dots and black luuules before the excision between the teeth.
( 75 )
Vertex, (horax, and abdomen white; lace, jialjii, and forelegs dark brown ;
legs white, tarsi and spnrs brown-mottled.
Expanse of wings : 1.") mm.
1 S.
Hiudmargin of forewing simply indented beyond ecll ; of hindwing with short
teeth at 4 and 7 ; antennae with clavate teeth.
:5;3. Epiplema vialactea spec. uov.
Forewiiiij : deep liver-oolonr, crossed by two deeper brown shades, one at
one-third, the other from three-fourths of costa to two-thirds of inner margin ;
costa and all the veins dotted with white ; beyond the outer shade between
veins 4 and 6 a large white crescent-shaped blotch of coalescent white spots,
these spots lying between, as well as on, the veins; the spots above it on
vein 7 and in the interval below also being larger but not confluent ; a row
of white submarginal dots between the veins, those above vein 4 large and
conspicuous ; a marginal row of white spots between veins, above vein 4 running
out into the fringe, which is liver-brown.
Hindwing: similar; the spots beyond cell and those in the whole lower
outer half of wing larger and more numerous, but not forming a crescent,
and coalescing along submedian fold ; marginal white spots between veins
throughout.
Underside of forewing pale liver-colour freckled with white, of hindwing
white freckled with brown; the fringes chequered white and liver-colour.
Head, thorax, and abdomen dark liver-colour; dorsum with white points;
vertex and shaft of antennae snow-while ; abdomen beneath and legs whitish ;
pectus and forelegs liver-colour.
Expanse of wings : 26 mm.
1 ?.
Hiudmargin of forewing simple ; of hindwing prominent, but hardly toothed,
at veins 4 and 7.
Family GEOMETRIDAE.
Subfamily OENUCHROMINAE.
Dicyclodes gen. nov.
Forewini/ : costa straight, with a faint curve at ba>e and belbre apex ;
apex minutely produced: hiudmargin strongly gibbous above, oblii|ue below, in
the ? faintly indented below ajiex.
Uindtviny : with apical angle and hiudmargin rounded ; anal angle rectangular.
Thorax woolly ; antennae of S uniseriate pectinate for two-thirds ; of ?
filiform ; forehead protuberant ; paljii stout, upturned, short, terminal segment
indistinct; frenulum and tongue present; hindtibiae not thickened, with
four spurs.
Neuration : forewing, cell longer than half wing ; discocellular obliijue
inwards to near median vein, then sharjily angled at the lower fork and oblique
outwards ; first median nervule at four-fifths, second close before end ; radials
normal ; 7, 8 stalked from end of cell, 'J, IK stalked a little before them, 9
( 7fi )
iuiastomusiiig siil)sei[ucntly with 8, 11 t'loiu cell; liiiidwiiij; with discocellnlar
augled outwards Id middle, the radial tioiii the aiigidatioii ; H, T and 3, 4 from
angles of cell, 7, 8 apiu'oximating near basi-.
Tvjic Dici/clodcx liicroqbjphicu spec nov.
Allied to Sarcinodes Guen.
34. Dicyclodes hieroglyphica spec. miv.
Forririnri : dark pearly grey, alonu; tlie costa beymid middle deep garnet-red,
shading lielow into deej) olive; costawith two snow-white streaks ; one antemediau
cnrviug in towards eell-spot, then apparently bent inwards and marked by black
dashes nn the median and snbmedian veins; a broad olive snbbasal band; cell-
mark garnet-red ; a very fine dark onter line, Innulate-dentate, the teeth pointing
inwards, parallel to hindmargin, at two-thirds, starting belnw the ]iostmedian white
costal streak ; a silvery white submarginal line, edged inwardly with black, from
costa just before apex, bent inwards and sinuous below vein 4, to inner margin
dose to outer line ; marginal area a mixture of shining grey and rosy, becoming
olive along the margin itself ; the veins whitish and bearing arrow-headed points
before the margin ; fringe dark olive.
nindtriiiy : with a broad central olive band, outwardly edged by a darker
sinuous line, containing a sinuate hyaline cell-mark ; a wavy grey submarginal line,
jircceded and followed by olive clouds ; fringe dark olive.
Underside of forewing silvery whitish, with transverse olive-brown striae ;
suffused with olive towards anal angle ; a broad straight central olive shade and an
(iblic|ne onter line of spots on veins starting from a costal streak at four-fifths ;
liindwing jicarly white only along abdominal margin, the rest olive-brown finsbed
with red, paler before the dark antemedian line ; an indistinct vyaved line beyond
cell-spot ; submarginal line lunulate-dentate, edged with grey at costa, forming white
teeth on veins 2, o, and 4, followed by two crimson blotches.
Head, thorax, and abdomen pearl-grey tinged with pale olive; alidomcn
beneath broadly blotched with crimson ; legs brown with pale joints ; pectus and
femora woolly, pale grey and pink.
Expanse of wings : i 4.S mm. ; ? 48 — 52 mm.
.5 cJcJ,3 ??.
8;"). Noreia pulverosa sjiec. nov.
Forewing: ]iale njouse-grcy, dustc(l with fuscous; costal edge brown; lines
dark brown ; first nearly straight, at one-third, hardly reaching costa ; second from
jnst before apex, and ajiparently retracted to costa, to two-l birds of inner margin ;
cell-sj)ot dark ; submarginal line indicated by dark sjiots lietwecn veins.
[linflicini/ : with onter line only.
Underside pale grey in forewing, whilish grey in liindwing ; the outer lines
dark ; the cell-spots marked ; costa of forewing yellowish brown ; hindwiug
8j)eckled with grey.
Thorax and abdomen dark grey; lillet fulvous; face dark brown; palpi
and legs grey marked with reddisli.
Expanse of wings : 38 mm.
1 S.
( 77 )
Subfamily PSEUDOTERPNINAE.
3(1. Hypochroma purpurissa spei\ nov.
Foreici iiq : pale bluish white, shailfd witli purplish slate-colour aud with
greenisli slaty striations ; costa with purple striae aud tour ])laclv l)lotchcs at
eveu distances apart; lines black; first ontcurved above aud below median to
quite one-third of inner margin, preceded there by a purplish shade ; outer luuulate-
dentate, from the third costal sjiot, ontcurved above aud incurved from 4 to middle
of inner margin, where it approaches inner line ; a large purplish black ear-
shaped cell-spot beneath the second costal spot; submarginal' shade cloudy
imrplish, darker and ontcurved beyond cell and between 1 and 3, the marginal area
at apex and between 3 and 4 pale bluish white ; black marginal luunles ; fringe
pale blue with black cheiiueriug beyond veins, its base beyond the Innules white ;
close to base of wing are three purplish sjiots, indicating a basal line.
IliiHliciiiy : more thickly striated ; no basal line, but a short dentate line on
discocellular; the rest as in forewiug ; inner margin and fringe golden yellow.
Underside deep dark purple, the apex of forewiug and a broad band just beyond
middle of hindwing pale bluish wliite, the baud showing slightly on inner margin
of forewiug ; lines and cell-s])ots indistinctly deeper purple ; inner margin of
hindwing more broadly yellow.
Pali)i externally and lower half of face purplish black ; upper lialf of face and
vertex slaty white; shoulders aud base of patagia the same; tips of shoulders and
a broad band reflexed to sides of patagia velvety black ; thorax and dorsum pale
slaty blue, the segments with pairs of lateral black spots ; anal segments and
sides and nnderneath of abdomen golden yellow ; tarsi black with yellow spots.
Expanse of wings : 52 mm.
1 (?.
In Xoc. Zool. x. p. 350 mention was made of 3 ? ? from the Upper Aroa
River (which were there referred to satarutaria Wlk.), in which the usual green
upperside of that species had acquired the slaty coloration of the underside ; the
last mentioned of the three certainly, and the other two probably, should be referred
to the present species. I am also inclined to keep H. caesia, from Fergusson and Ron
Islands, distinct from the New Guinea species.
RtiBFAMiLY GEOMETRINAE.
37. Anisog^amia albifusa spec. nov.
Foi'etcing : hyaline green ; the basal third, the discocellular and space beyond,
and the three onter lines of distinct conjoined Innules powdery white ; across the
green central area the veins are strongly dotted green and white ; costa fuscous,
irregularly mottled with white ; marginal white dots at the ends of the veins ;
fringe green flecked with whitish.
Iliiidwiiiii : mottled all over with white scales from base, without any central
green space ; all the veins throughout uniformly dotted green and white.
Underside whitish green, iridescent ; ajiical half of costa of forewings and
marginal dots of hindmargin below apex bronzy fuscoiw.
Head, thorax, aud abdomen green si>eekled with white ; tillet narrowly white ;
( 78 )
face green above, the lower half white ; abdomen beneath, pectus, and legs white,
foretarsi brownish.
Expanse of wings : Sn nun.
2 JcJ.
3>^. Anisogamia albiseriata .spee. nov.
Fori'wi III/ : diirk green ; costal edge white, with dark brown flecks, thickeneil
towards apex ; lines represented by series of white spots ; first from one-fourth
of costa to one-third of inner margin, consisting of small sjwts, on the folds as
well as on the veins ; middle line of larger spots, two somewhat diffnse on each
side of vein 7, a partially donble ipiadrate spot beyond cell, a large one on inner
margin at middle, with sometimes a smaller one above it on the fold ; exterior
series placed on the veins, those on veins 4 and T) coalescing into a blotch and
displaced basewards ; snbmarginal series more elongate and diffuse, situated
between the veins, those on the folds larger : fringe pale green with distinct
white mottling beyond veins.
IliiKhving : without basal series of spots ; the middle series often enlarged into
a broad white band, containing two dark green spots on discocellnlnr : fringe
of inner margin white.
Underside whitish green, the white spots showing through ; costa of Ibrewing
yellowish.
Head, thorax, and abdomen deep green ; dorsum with conspicuous white s[)ots:
fillet white : antennae greenish white; abdomen beneath, jiei'lus, and legs white;
forelegs greenish.
Expanse of wings : 10 to 24 mm.
5 c?(?.
Hindmargins witlmut crenulations ; hindwing slightly elbowed at veins 4 and fi.
3'.i. Anisogamia batis spec. nov.
Forewikg : dull grass-green ; costa broadly brown, speckled with ochreons ;
marginal line thick, jmrplish brown, finely edged externally with flesh-coloured
ochreons, which below apex is swollen into spots between the veins ; fringe pale
grey with a purplish grey basal line ; hindmargin marked with two flesh-coloured
ochreons blotches, edged and centred witli purjilish brown ; one rounded between
veins '.J and 5, the other flattened and elongate at anal angle below vein 2.
lliiidwiny : with the middle marginal blotch ranch snniUer ; cell-spot white.
L'nderside pale blue-green, the marginal blotches showing white ; costa of
forewing i'uscons sjieckled with pale.
Head, thorax, and abdomen green ; palpi externally, vertex, fillet, and
antennae reddish varied with paler ; dorsum with more or less coalescent patches
of flesh-colonred ochreons and grey scales ; abdomen beneath, pectus, and legs
whitish ; forelegs pinkish Ijrown, with pale joints.
Expanse of wings : 2>i mm.
:i ? ?.
411. Anisogamia commaculata spec. nov.
Fofi'iriiiy : semi-transparent, deeji green, with snow-white blotched lines ; a
white blotch at base of costa ; an oblique one ou inner margin near base and
a (juadiate white blotch on costa at one-fifth ; a s]iot before middle of inner margin
( 79 )
and an irregular wliite liloteh in end of ci'll, eonneeted with a white lilotch on eosta
before it, represent a middle line ; three series of broad white Innnlar blotches before
hindmargin, not reaching eosta, and all iiiteri'npted between veins 2 and 3 ;
veins bevond eell dotted alternately green and white ; costal edge fnscons,
irregularl}' interrnpted by white marks ; fringe grey cut by white flecks from the
white marginal dots at the vein-ends.
IliiidiciiKi : with a large white blotch at base of cell, one on discocellnlar, and
three below eell ; followed by the three series of lunnles as in forewing ; fringe
paler ; veins near base deep green.
Underside pale green, with the wliite markings showing through ; eosta of
forewing broadly fuscous marked with white, towards apex projecting toothlike
into the green area ; marginal dots between veins large, fuscous ; a fuscous snbapical
spot on hindwing.
Head, thorax, and abdomen green ; abdomen with silvery white dorsal and
subdorsal blotches; fillet and lower half of face white; abdomen below, pectus, and
legs white; foretarsi mottled fuscous and white.
Expanse of wings : 34 mm.
1 6.
41. Anisogamia decorata spec. nov.
J. Forrwiiifi : semi-transparent, grass-green ; costal edge dark brown freckled
with creara-colonr, extending across liase nearly to inner margin ; first line starting
from a cream-coloured costal spot at one-fourth to two-fifths of inner margin,
lunnlate outwards, dentate inwards, pale above median vein, i'uscons below ; outer
line lunulate-dentate, from nearly three-fourths of eosta to two-thirds of inner
margin, obscurely marked across wing, but from eosta to vein 6 edged with
white, and followed by a dull whitish blotch containing pinkish scales and
outwardly shaded with black scales ; submarginal line marked on the outside edge
of this blotch by two white luuules and below by spots of pinky browu between
the veins, ending in a flattened blackish blotch at anal angle, containing some
reddish scales beyond outer line ; cell-spot a white crescent ; white marginal spots
at the vein-ends ; fringe rufous grey.
Hindwing : with only the two outer lines ; a large blotch of smoky black
and pink scales at apex, the submarginal line marked across it by three black
pale-centred wedge-shaped marks.
Underside pale iridescent green ; eosta of forewing yellowish, sjieckled with
dark brown ; two brown costal blotches before aj)ex, with a yellowish space
between them; a blackish blotch at annl angle, and a large oval black blotch at
apex of hindwing ; fringes green tipped with brown; brown marginal s])(its b(dow
ai)ex of forewing.
Head, thorax, and abdomen dark green ; thorax and metathorax marked
with dark and light grey scales, with a green space in the centre ; second and
third dorsal segments with small white spots, fifth and sixth with large grey
blotches ; anal tuft pale oehreous ; face with two white spots below ; fillet
whitish ; palpi externally dark brown ; abdomen beneath, pectus, and legs whitish ;
foretarsi blackish, with white joints.
? . Forewing : with costal edge more broadly but paler brown, thickly
speckled with white ; first line fine, simply waved ; outer line finely white,
forming two large lunnles to vein 4, along which it runs out to close to hind-
( 80 )
margin, skirting this to vein 2, then cnrving inwards to inner margin before
anal angle, where it is followed liy dark grey scaling; the sqnare apical l)lot(^li
is filled ni> witli rnfons grey externally, siqianited by three green j)iile-edged
InnidfS from the pale edging nt' the onter line : niuvginul line dark, swoJIen into
spots between veins.
Ili)i<hciiiq : like fnrewing, the ii|iii'!il lilntih consisting of dark and light grey
scaling.
Undersidr wirli flie hhuk lj!cit<-lu's larger, tiie liircwing with a single pear-
shaped blotch before apex.
Dorsum wholly reddish grey, except the basal segment, wliicli alone is green.
Exi)anse of wings : cJ, 4n mm. ; ? , 36 mm.
3 <^cJ,3 ? ?.
4:.'. Auisoganiia flavilinea spec. nov. and ab. albinata nov.
Foreivinq : semihyaline dee]> green ; costal edge narrowly fnscons, uniformly
dotted with pale; the lines pale yellowish ; first from one-thinl of costa to two-fifths
of inner margin irregnlarly zigzag ; outer somewhat irregularly deutate-lunnlate,
from three-fonrths of costa to two-thirds of inner margin, bent inwards below vein 3,
and above it followed by a blotch of pale scales ; a submarginal line of yellowish
spots between veins, that between 3 and 4 biaugnlate, almost obsolete below :
a row of jiraemarginal wedge-shaped yellowish sj>ots between veins, that between
3 and 4 large, obsolete below ; yellowish white marginal spots at the vein-ends ;
fringe grey-green, finely chequered with yellowish ; in the basal and median
areas are traces of yellowish cross-lines marked by vein-dots only ; across the
discoeellular and reaching vein 2 is a diffuse grey cloud.
Tlindwing : similar, without basal line or central cloud, but with a small
grey patch at apex.
Underside pale iridescent green, the lines showing through ; costa of forewing
yellow, with fine brownish speckling, underlined before apex by three deep green
blotches ; a brown spot at apex. ^
Head, thorax, and abdomen deep green ; dorsum with white s])ots, a patch
of brown scales on the penultimate segment, and the anal tuft whitish ; face with
two white dots, one on each side below; antennae red-brown with liasal segment
white; palpi externally pale red-brown; forelegs red-brown with white joints;
abdomen beneath, pectus, and legs white.
Expanse of wings : 4it mm.
.-. Jc?,4 ??.
The aberration albiiiatit has all the lines white, generally much less marked
than in the ty})e form, except between veins 3 and 4, where the expansion of
the three lines forms a conspicuous whitish patch. It is also, on the average, a
little smaller than the type form.
6 cJ<?,l ?.
In the single ? the white markings are greatly reduced, while the brown
apical blotch of hindwing is amplified.
43. Anisogamia griseonotata s])ec. nov.
Foirivhig : dull grass-green, semi-truns|)arent ; costal edge fuscous, with fine
white striations ; the lines very fine, whitish ; first from one-fifth of costa to
one-third of inner margin, Innulnte-deutate, the teeth pointing inwards, followed
( 81 )
by a (lull t'nscou.s shade ; outer line from two-Uiircls of costa to two-thirds of
inner margin, lunulate-dentate, tlie teeth pointing outwards and marked b)- white
dashes on veins, preceded b}' a fuscous shade : a submarginal row of white spots
between the veins ; a row of large white marginal dots at the voin-cnds ; fringe
green ; cell-mark a tine white crescent, outwardly edged with fuscous.
Ilinduim/ : with some white spots on veins near base; the rest as in forewing,
but the outer line ends in a large white spot on inner margin.
Underside pale iridescent green ; costal edge of forewing yellow with fine
black specks ; a dark speck at apex.
Head, thorax, and abdomen deep green; centre of thorax and metathoracic
tuft fuscous ; abdomen with white dorsal spots ; lower half of face white ; terminal
segment of palpi brown with pale tip ; antennae annulated brown and wiiite.
Expanse of wings : 30 mm.
8 ??.
44. Anisog'amia iridescens spec. nov.
Forewhu/ : hyaline green, the wing membrane being covered with very fine
hairlike scales ; costal edge fuscous, uniformly cut with white d<its ; all the veins
deeper green alternated with pale points, representing the dark and light lines
crossing the wing, whicli, however, are not visible except on the veins ; the only
visible lines .ire an inner interrupted line from one-fonrth of costa to near middle
of inner margin; an outer line of ])ale lunules from four-fifths of costa to two-
thirds of inner margin, followed by a similar submarginal line and a series of
pale marginal spots, the spots of these three lines all lying in the intervals;
a deep green marginal line containing white ]ioints at the vein-ends ; fringe
mottled green and white.
Hind icing : the same, but without any basal line.
Underside pale iridescent green, the markings merely showing through ;
extreme costal edge towards apex brown.
Head, thorax, and abdomen green s]irinkled with white scales ; face with
white cheeks ; abdomen with whitish dorsal siiots ; antennae fnscons : pectus and
legs whitish green; forelegs fnscons with white joints.
Expanse of wings : :^it mm.
1 cJ.
4;"). Anisogamia sciutillaus spec. nov.
Farnring : deep gri'cn, sprinkled with ilaky white scales, which in some
lights are scintillating ; costa narrowly white ; its inner edge with some brown
striae, which towards apex reach the costal edge; first line marked by white
dots on folds as well as veins ; a white dot on discocellular and another on
vein T) beyond cell ; an exterior and submarginal row of white sjjots between
the veins, forming each a white lunulc on submedian interval and a white sjnit
on inner margin ; the spots beneath vein .5 large and displaced basewards ; tlie
apical spot of the submarginal row large ; on each side of the last series a
series of white dots on the veins ; fringe green with conspicnous white cherpiering.
Ilindiring : without basal line.
Underside jinle green ; the forewing darker except on hindniargin ; costa of
forewing yellow.
6
(. «2 )
Head, tlumix, and abilomt'ii deep green; (illcl lnoadly siiuw-white ; dorsum
with snow-wliite spots; underside of ahdonien, pectus, and lej;s wliite ; I'oretarsi
fiiscons witli pale joints ; antonnal shaft white, the pectinations green.
Expanse of wings : '2^ mm.
1 3.
4(). Anisogamia semiuivea spec. nov.
Forein'iii/ : deep green, irvegnhirly i'rctkled with white ; costal edge white,
witli slight obli(|ne fuscous striae ; basal and subcostal areas powdered with
bluish white ; an ill-defined oblique line near base, marked by white spots on
veins and folds upon a deep green space ; median area occupied by diffuse white
blotches below costa, and above inner margin containing deep green granular
speckling ; outer Hue acutely luunlate-dentate, the lunnles on the folds only
showing white ; marginal area filled with white and green speckling ; a dark
green margintil line; fringe pale green chei[uered with white.
IIiii(hriii(j : white sjieckled with green, except on costal area; a distinct white
Innnlate-dentate outer line, preceded by a deep green shade ; cell-spot deep green.
Underside whitish green, deeper green towards costa of forewing, which is
white.
ITc;id, thorax, and abdomen deep green ; palpi beneath, fillet, antennal shaft,
and dorsal sjiots white ; antennal ]icctinations and forelegs greenish ; abdomen
beneath, pectus, and legs white.
Expanse of wings : 28 mm.
1 cJ, 1 ?.
47. Auisogamia triseriata spec. nov.
Forciving : grass-green; the costal edge finely but conspicuously white;
inner line close to base, very indistinct, ontcurved above and lielow median
vein, and ending in a white spot on inner margin ; discal spot dark, preceded
by a few white scales ; outer line deeply Innulate-dentate, the white lutmles
appearing as an interior line and the white teeth as an exterior line ; large
marginal white spots at the vein-ends ; fringe concolorons.
UiiidiriiKj : similar, but without basal line.
Underside pale bluish green, the fringe showing rather darker.
Head, thorax, and abdomen grass-green; fillet white; dorsum with small
wliito spots; aiiteMiiae with shaft white, and ])e(-tinations greenish ochreous ;
abdomen l)ene!ith, pectus, and legs white; forelegs greenish.
Expanse of wings : ;!4 mm.
2 a.
The jialpi, especially the terminal segment, nuuh sluirtor than usual.
48. Anisogamia viridissima sjjoc. nov.
Foi-eir'uKj : very deep green, the costal edge whitish at base, tlien reddish
In-own : lines bluish white, very slender ; first close to base, dentate inwards
on veins, Innulate outwards ; outer line at four-fifths, dentate-lunulate, the teeth
marked on veins, ontcurved from costa to snbmedian fold, then vertical ; in the
middle of wing, from subcostal vein to snbmedian fold, a broad greenish black
blotch : fringe dee]! green ; cell-spot black.
Iliiiiliriiiii : without inner line.
Underside glossy bluish green ; cost.a of forewiug yellow.
Head, thorax, and alidoraen dark green ; fillet and dorsal spots white ;
alidomen beneath, iiectns, and legs white ; foretarsi green : jialpi green externally.
Expanse of wings : 28 mm.
1 ?.
40. Chlorochroma discata spec. uov.
Foreichnj : deep apple-green, frosted with jialer; eostal edge snow-white,
nnderlined by a fawn-colonred streak ; the two lines huinlate-ilentate, slightly
[laler, edged conversely with deejjer green : first from one-fifth of costa to l.ieyond
one-third of inner margin, obscurely toothed inwards on the veins ; the outer
from three-fourths of costa to two-thirds of inner margin, outcnrved above ;
cell-spot a large round brown disc with deep smoky black centre ; deep red
marginal spot;* at ends of veins ; fringe yellow.
lliiKhviiK] ; the same, the cell-spot somewhat larger.
Underside shining whitish green, deeper green below costa of forewing, which
is snow-white underlined with fawn-colour: fringes whitish ; marginal dots showing
only below apex of forewing.
Head, thorax, and abdomen green ; vertex white, edged behind with ]iink ;
abdomen with yellow dorsal line ; antennae whitish green ; palpi above and foreiegs^
in front tinged with fawn-colour ; abdomen beneath, pectns, and legs wliit('.
Expanse of wings : 36 mm.
Hindmargin of forewing straight and vertical, anal angle square-cut.
50. Chlorochroma flavilimes sjiec nov.
Fm'ewiiig : grass-green ; costal edge white, underlined with yellow, reddish
at base ; lines fine, yellowish white, plainer towards inner margin ; first from
one-si.xtli of costa to one-third of inner margin, bluntly angled on median vein ;
outer parallel to hindmargin, from three-fourths of costa to two-thirds of inner
margin ; fringe vellow, bevond minute red dots at the ends of the veins : rell-spnf
black.
J/i/u//ri/itf : the same, but the outer line slightly curved.
Underside pale green, deeper towards costa of forewing, which is yellow.
Head, thorax, and abdomen dark green ; fillet broadly silvery white ; basal
segment and shaft of antennae white, the pectinations greenish : a yellowish white
dorsal line down thorax and abdomen ; anal segment white : abdomen beneath
and legs whitish green, forelegs green-tinged.
Expanse of wings : 40 mm.
1 cJ, 3 ? ?.
ol. Chlorochroma geminipuacta spec.
nov.
Fori'/r/m/ : ileep sea-green, semi-transparent; costal edge finely dull red;
lines very faint, marked by tlie difference of tint : first oblique from one-sixth
of costa to before middle of inner margin ; second Innulate-dentate, from two-thirds
of costa to three-fifths of inner margin, preceded, as the inner line is followed,
by a deeper green tint; cell-spot black, small, preceded by a reddish streak on
ujiper arm of discocelliilar ; fringe yellowish.
(84)
Hindwhuj : similar.
Underside paler green.
Tliorax and abdonion jircon ; face, palpi, and collar red ; vertex snow-white ;
iintcnnae reddish; paljii and face helow jialer : lorelegs iu front reddish.
Expanse of wintrs : :!ri mm.
1 ?.
.")2. Chlorochroma laticostata spec. nov.
Forewuifi : dull grey-green, snl)tran.^]jareut ; costa broadly white, before apex
underlined with red ; lines dentate-lnnnlate, very faint, the teeth minntely whitish
on the veins ; first from one-sixth of costa to one-third of inner margin, dentate
inwards ; second from three-fonrths of costa to two-thirds of inner margin ; cell-spot
small, brown : marginal line vinons red, interrnjited at the vein-ends : fringe
yellow.
Ilinthcing : like forewing.
L'nderside whitish green : costa of both wings yellow.
Head, thorax, and abdomen green ; tillet and antennae white, apical third red ;
abdomen beneath, pectus, and legs white.
Expanse of wings : liO mm.
ij,r>??.
;")3. Chlorochroma latistriga spec. nov.
Foretriiig : blue-green, frosted with paler; costal edge white, nnderlined
thronghont with fawn-colonr ; a narrow pale inner line from one-fifth of costa
oliliijue to one-third of inner margin ; an outer nearly straight broad j)ale yellowish
line from three-fonrths of costa to two-thirds of inner margin, widening downwards;
cell-spot black, large ; marginal spots black, triangular ; fringe pale yellow.
lUndirhui : similar ; the cell-spot .short and linear ; outer line broad, slightly
curved.
Underside pale green, the broad outer line sliowing through ; costa of forewing
yellow, dnsted with dark ; marginal black spots.
Head, thorax, and abdomen green ; vertex white ; antennae greenish white ;
abdomen beneath, pectus, and legs white; forelegs green in front; palpi external ly red.
Expanse of wings : -i^) mm.
1 (J,2 ? ¥.
spec.
•">4. Chlorochroma obsoleta
Foreiriiiii : darker greyer green than pimrtillificni ; the two Innnlate-dentatc
lines almost equally indistinct; costal edge more broadly and distinctly yellow;
cell-spot green, not black ; marginal dots smaller, darker ; fringe yellowish.
Hi lid wing : like forewing.
Underside whitish green ; costa of forewing yellow.
Head, thorax, and abdomen deep green; fillet white; antennae greenish;
anal tufts and sides of jirae-anal segments while; abdomen beneath, jiectus, and
legs whitish ; forelegs greenish in front.
Expanse of wings : 44 mm.
1 <?, I ?.
The apex of forewing is minntely produced.
( ss )
o.). Chlorochroma punctilligera spec. nov.
ForciriiKi : dnll pale green, the same tint as in C. minor ^Val r. ; (lie costiil
edge white at base, pale fawn-colour beyond ; lines dentate-liinulatc, bnt scarce]}-
perceptible : cell-spot a red-brown point on a grey bunt line ; marginal [loints
minute, reddish ; fringe pale yellow.
lUmhrimi : similar : the lines rather more plain.
Underside uniform pale green.
Thorax and abdomeu green like wings, with a fine yellowish dorsal line ; face
and pali)i deep green ; vertex and antennae white ; abdomeu beneath, pectus, and
legs whitisii ; forelegs greenish.
E.\j)anse of wings : 40 mm.
3 ? ?. One of these is smaller, measuring only ;3o mm.
."it>. Chlorochroma ruficosta spec. nov.
ForewiiKj . grass-green, the marginal half paler; costa rufous witii tine dark
speckling; an obscure pale line from costa near base to nearly one-third (if inner
margin, the green beyond it deeper; a straight ])ale line, parallel to hindmargin,
from two-tliirds of costa to three-fifths of inner margin, where it is pbiiiier,
inwardly edged with darker green; marginal line dull dark pink, still darker at
the vein-ends ; fringe paler pink.
Uiiidwimi : similar.
Underside jiale green ; costa of lioth wings pale, yellowish ; fringe yellowish.
Face, thorax, and dorsum green ; thorax and dorsum with a jiinkish yellow
central line; vertex snow-white; collar crimson; abdomen laterally and beneath,
also the whole anal segment, pectus, and legs whitish green ; forelegs tinged
with pink.
Expanse of wings : 40 mm.
1 S.
Hindwing with hindmargin bluntly elbowed at middle.
57. Chlorochroma rufistriga spec. nov.
Vnn'irimf : [lale green, frosted with whitish-green scales ; costal edge white,
finely underlined towards base with fawn-colonr : inner line from costa close to
base to one-fourth of inner margin, straight and oblique, whitish outwardly edged
with fawn-colour ; outer line parallel to hindmargin from three-fourths of costa
to two-thirds of inner margin, white internally edged with fawn-colour ; w. fawn-
coloured linear mark on discocellular. bent below njiddle and marked with a daiker
sjiot at the bend; fringe fawn-colour with slightly paler base.
Uiiidwimj : like forewing ; the cell-spot green.
Underside whitish green; furewing towards costa speckled, and at base snlinsed,
with fawn-colour.
Head, thorax, and abdomen green, face deep green ; fillet and antennae wliite,
the former edged behind' with crimson ; shoulders at sides pink ; a1)diiinen witii
broad yellowish dorsal line becoming white towards end, where the anal segments
are themselves wliite; jialpi above and forelegs in front tinged with fawn-colour.
Expanse of wings : 44 mm.
2 ? ?.
Hindwing with hindmargin rounded.
( «r, )
;iS. Chlorochroma vestigiata spec. nov.
Voiiiriiui : dark grass-green, the same tint as in ''. mfdiotiiiclo Warr., Imt
nut ([uite so tlarlv ; eosta yellowish white ; lines marked by whitish vein-dots, on
a taint, scarcely jierceptible line ; first oblique, from one-sixth ol' costa to one-third
of inner margin ; second line at three-fii'tlis, sliglitly angled on vein 'i, and bent
on vein 4 ; fringe yellowisli ; cell-spot green,
llimhrinq : like forewing, with distinct dark marginal dots; the outer line
angled on vein 4.
Underside whitish green ; marginal dots distinct on both wings.
Head, thora.x. and abdomen dark green ; dorsum with small while dots; liliel
and antennae white: abdomen beneath, i)ectus, and legs whitisli.
K.\])anse of wings : 3o mm.
2 ? ?.
l)iflcrs from (.'. mcdiotim-tu Warr. in the lines, the fringe, and the dorsal spots.
o'.i. Comostola fiavifimbria spec. nov.
Vorcivuiq : ])ale apple-green ; costal edge finely yellow, with a few brown
speckles towards apex ; a very obscure curved line near base, and a dentate-lumdate
inwardly darker-edged line beyond midille: a triangular red-brown cell-s]iot with
some lustrous scales on it ; a thick red-brown marginal line, interrupted by yellow
dots at the vein-ends, overlaid with shining scales ; fringe yellow.
Ifiixhrinij : with the cell-spot larger, iiyriform.
b'nderside jiale green, with the fringes and costa of forewing yellowish ; a
red-brown sjiot at apex of forewing.
Face red above, whitisli below; fillet and antennae white ; vertex yellowish;
thorax and abdomen green ; abdomen beneath pale green ; legs whitish ; forelegs
tinged with reddish.
Exjianse of wings : 22 mm.
1 ?.
t'loselv allied to C. nUilHinlo-hi Warr.. from the Khasias.
'ill. Comostola rufimargo spec. nov.
Forcicinii : ]iale a]ijilc-grcen : costa deep pink, freckled with shining ilark
scales; hindraargin and fringe pink; thick red marginal lunnles between veins,
overlaid with lustrous scales ; a jiink yellow-edged sjiot on inner margin before
one-third indicates the inner line; a similar, but larger, spot at two-thirds ends
the outer line, which is marked across wing by small white vein-dots, starting
from a dark costal spot ; discal spot like those on inner margin.
UiiKhriMj : the same, but without the sj)ot indicating inner line
Underside pale shining green, the fringes pinkish tinged.
Face, ])alpi, vertex, and basal half of dorsum deep pink; fillet while; antennae
reddish ; thorax, patagia, anal half of abdomen, and its base laterally green :
underside and legs pale greenish white.
Exjianse of wings ; 1<5— IS mm,
4 (?cJ, 4 ? ?.
( 87 )
liJ. Comostolodes castaneata sj>l'c. nov.
J'arr/riiK/ : apple-greini ; costal area paler, the costal edge pale t'awii-ioliMir ;
lines bright chestnut-brown, double, starting from subcostal vein ; first vertical at
one-fourth, the two arms divergent aliove middle ; median beyond middle, waved,
bent inwards along vein 2, then vertical to middle of inner margin ; space between
the arms in lioth lines green ; a brown marginal shade, obliquely swollen at apex,
ibrniing an inward triangle on vein 4, and swollen into a large blotch before
anal angle, which touches middle line as a ronndish blackish blotch, externally
edged by a white spot, the red-brown above it nearly reaching vein '?> ; a row of
deejier brown dashes between veins along margin ; fringe brown, faintly che(piered
with greenish ; cell-spot black.
lUiulirimj : with only the brown border, which is much swollen at apex and
anal angle, containing a i)ale pink blotch above vein 4, between which and vein 'Z
the green ground-colour rnns out nearly to hindmargin ; inner margin narrowly
brown nearly to base.
Underside dull green, with all the brown markings showing through.
Head and paljii mixed, red-brown and whitish; sides of metathorax, tips
of jiatagia, and abdomen red-brown ; thorax otherwise green ; abdomen beneath,
pectus, and legs pale green ; forelegs red-brown mottled with i)ale.
Exi)anse of wings : 22 mm.
2 Si.
62. Comostolodes viridifimbria spec, no v.
Forcicinr/ : grass-green, rather thinly scaled; costal edge jiale brown; lines
denoted by white dots on veins, the lowest in each series, on vein 1, being larger
and ringed with bi-own ; first line marked otherwise only by a dot on median vein
at the origin of vein 2 ; the onter line at four-fifths, parallel to hindmargin, angled
on vein (1 and starting from a white spot at three-fourths of costa, with the spot
on vein 4 like that on vein 1 ; white marginal dots at the ends of the veins ;
fringe green ; cell-spot brown.
Hindiciitg : without lines ; cell-spot brown ; marginal spots white, large, that
at anal angle ringed with brown.
Underside paler green : the spots of onter line of forewing showing throngh.
Head, thorax, and abdomen green ; fillet and front of shoulders whitish.
Expanse of wings : 2ii mm.
2 SS.
i;:!. Gelasma imitans sjiec. nov.
Forcicinf) : dull grey-green; costal edge finely speckled, fuscous and white:
lines white, Innulate-dentate ; hrst from one-fifth of costa to two-tifths of inner
margin, dentate inwards on veins ; outer line froin five-sixths i)f costa to two-thirds
of inner margin, rnnning parallel to hindmargin to vein 2, then rnnning in
basewards ; the hmules on each side of vein 4 broadly white and disi)la(ed
basewards, followed by a similar double white Innule touching hindmargin :
marginal spots large and white ; fringe ]iale green ; cell-s])ot dark green.
HhidwiMj : like forewing, but with no basal line.
Undersiilc jiale green, the white lunules showing throngh ; costa of forewing
yellow.
( 88)
Head, iLidrax, and abdoiiii'ii <;ii.x'ii ; autfuiiae wIuIl' ; tillet sliglitly jialer green.
Expanse of wings : 40 lum.
1 ?.
In tlie ,;;riuitL'i' dcvcluimient of llir white markings at middle i>f liiudtnargia
of botb wings this speeies mimics several sj)ecies (if A/t/.i()i/u////fi.
04. Gelasma invidens spec. nov.
FoiririiKi : ilull greyish green ; cosia fnscons sjieekleil wilh wliite; lines while,
distinct, dentate-liinulate ; first from one-tifth of costa to two-lifths of inner margin,
dentate inwards on veins ; outer line from tbree-foiuths of costa to two-tliirds of
inner margin, the limnles on the folds stronger; cell-s])0t dark green; fringe
pale green, ]ialer at tips.
IlinihciiKj : without inner line.
Underside paler green.
Head, thorax, and abdomen green ; fillet and aiitennal shaft white, the
pectinations greenish ; dorsum with white dots ; forelegs greenish.
Expanse of winns : 38 mm.
I S.
(')."). Gelasma spumata sjiee. nov.
Forewiiii/ : semi-transparent, jiale dull green frosted with white scales ; a
slightly curved inner and an obliijue broader postmediau band of uuspeckled green ;
a tine waved submarginal line ; cell-spot dark green ; fringe jiale green.
Iliiulidni/ : similar, but the outer band narrow, and acutely angled on vein 4 ;
cell-spot large, black.
Underside whitish green.
Face and jialpi above olive-green ; thorax and abdomen pale green speckled
with whitish ; vertex whitish, with a few green scales ; aVidouien beneath, pectus,
and legs whitish ; forelegs olive-green in front.
Expanse of wings : 30 mm.
1 ?.
00. lodis bicolor spec. nov.
I'oreuiiig : semi-transparent hoary green ; the costal edge narrowly fawn-
colour, green at base ; lines luuulate-dentate, indicated only by the difference of
colour ; first vertical, from one-fourth of costa to one-third of inner margin, dentate
inwards on veins, followed by a deep green shade ; second from two-thirds of costa
to two-thirds of inner margin, the teeth on veins :> and 4 projecting, preceded by
a deep green shade ; fringe green ; cell-spot black.
IJiiiihriii)/ : without basal line.
Underside jiale green, with the cell-spots and outer band showing llirough.
Head, thorax, and abdomen green; antennae greenish wliite; abdomen beneath,
pectus, and legs whitish ; forelegs greenish.
Expanse of wings : 22 mm.
'MS.
Very much like I. JrayiUa Warr., but rather smaller; the cell-spots black,
not green.
( 89 )
117. lodis conimixta j^pte. iwv.
Forewing : deep moss-green, varied along middle of central fascia and along
liindmargin with whitish striae ; costal edge finely fawn-colour ; two broad dittuse
dark shades, the first cnrved at cue-third, the onter at two-thirds ; on the outer
side of this last can be seen a Innulate-dentate line, with the teeth marked white on
tlie veins ; a fine dark marginal line ; IVinge paler green ; cell-mark greeu.
IliMliriiKj : like forewing.
Underside whitish green, deeper beneath the dark bands.
Head, thorax, and abdomen green ; fillet and antennae snow-white ; abdomen
beneath, pectus, and legs white ; forelegs greenish iu front.
Exjiause of wings : -!'.) mm.
2 ? ?.
Larger than l./raijiU.^ Warr., wilh more confused markings.
nov.
08. Pyrrhoi'achis ruficeps spec.
Foieicing : briglit ajiple-green ; costa, liindmargin, and fringe deep pink : the
costal streak freckled with fuscous ; the liindmargin with some grey scales and
inwardly edged with yellowish.
lliiiihciiii/ : with the hindmargin and fringe jiiiik from l)efore apex to above
anal angle.
Underside pale greeu ; fringes and costa of forewing pale pink.
Upper half of face, vertex, and dorsal streak deep pink ; antennae pink and
white : lower half of face and fillet white; patagia and sides of abdomen green ;
abdomen beneath, pectus, and legs whitish : forelegs pink.
Expanse of wings : c^, 17 mm. ; ¥, ".'2 mm.
3 S6, 1 ?.
*i'J. Rhomborista exililinea spec. nov.
Fori'iciufi : deep emerald greeu ; costal edge snow-white : lines very fine, jjali'
green ; first from near base of costa obliipiely cnrved to one-third of inner margin :
onter line from two-thirds of costa t<i two-thirds of inner margin, irregularly
crenulate, the teeth on veins 3 and 4 jirojeetiug outwards, running in strongly
along vein 2, an<l again obliiiuely outwards, jiarallel to inner line; cell-spot deep
green ; marginal spots white and hii'ge, between the veins ; fringe purplish grey,
with a whitish middle line,
llimhriiuj : like forewing.
Underside almost white; costa of forewing yellowish; fringe white wilh the
outer half purplish grey and with purplish grey spots beyond the vein-ends.
Head, thorax, and abdomen deeji green; dorsum with silvery white spots:
sides of the iirae-anal segments and anal segment itself white : fillet and antenna!
shaft white, the pectinations greenish ; abdomen beneath, pectus, and legs shining
white : forelegs tinged with greeu in front.
Expanse of wings : c?, 4n — 44 niui. : ?,44mm.
0 t? c?, 2 ? ? .
This species stands i|uite alone, and will probably reipiire a new genus for
its reception.
r 90 )
Til. Thalassodes zebrata spec. nov.
ForcuiKg : green, crossed olili'|uely by lliive pairs nf broad gretniisli wliile
bands ; two near base, two in middle, bro;idev and coalesccnt Ijelow median vein,
separating again towards inner margin, containing above a dark green crescentic
cell-spot; two towards hindmargin, ending above anal angle : fringe green ; costal
edge brownish oolireons : the inner edge of the outside band is obscurely lunulate,
outlined by a dentate line which is only clear below middle, running in basewards
above vein 2, and again angulated outwards on vein 1.
Jlindirinij : almost wholly greenish white ; a curved dark green line IVom near
base of costa to one-third of inner margin, which is dark green throughout; a
dark green streak from beyond middle of costa, angled on vein 4, becoming dentate-
Innuhite and narrow to two-thirds of inner margin, bounded outwardly by a
greenish-white line; marginal area frosted with greenish white; a dark green
discal line.
Underside mealy whitish green : costa of lorewing yellowish.
Head, thorax, and abdomen green; fillet and hiwer jiart of face while;
abdiimen beneath, pectus, and legs white ; forelegs greenish.
Expanse of wings : 44 mm.
Srm--A5aLY STERRHINAE.
Tl. Perixera (?) bisecta spec. nov.
J'lin/n'iii/ : greyish stone-colour, dusted with fine dark atnms and tingeil
with pale fawn; first line curved, close to base, and marked by duts on veins;
second at five-sixths, similarly marked : marginal dots minnte ; cell-sjuit hardly
marked; median shade re])resented by a brown-red straight line a little beyond
middle ; fringe j)aler.
Hiitdicinij : similar ; cell-spot whitish ; outer line jilainer.
Underside whitish, slightly rosy-tinged, especially in the forewing ; luedian
band and outer line marked; dark marginal triangles.
Head, thorax, and abdomen like wings ; face fuscous above, whitish below ;
l)alpi reddish above; abdomen laterally with some red marks.
Expanse of wings : 3.") mm.
:-' V ?.
''-. Perixera C') festiva spec. miv.
I'on'iriiKj : ]i;i,le pearl-grey, darker at base, wilh imnierdiis purpli>h sjieckles ;
first line obscure, marked by dark points on veins and folds, and when visible
ontcurved above and below median vein, olive ; median shade olive, thick, incurved
below median ; irregularly lunulate-dentate ; cell-spot blackish, small ; outer line
marked by jMirjile spots on veins ; praesubterminal shade marked by three olive
clouds, at costa, beyond cell, and above inner margin ; marginal sj)ots purj)le ;
fringe concolorous.
llimhviiHj : like forewing, but the cell-s]iot is feri'Uginous, with some pale
scales in centre, the whole within a dark purplish ring.
Underside whitish : forewing sntVnsed and speckled witli dull reddish ; outer
line and cell-s]iot dull led ; marginal line purplish; liindwing with costa and cell-
spot red ; outei' and marginal lines purplish.
( '-n )
Head, tljciiax, and aliilniueii like wiii^s. tlic abdoiucu tiiiucd willi viudiis :
face and palpi deep red-lmiwii ; abdomen beneatb wliitisb ; le^'s in front reddish.
Exjianse of wings : 34 mm.
4 ¥ ?. ^
7;'>. Perixera (h iudigens spee. nov.
Foiciri/n/ : greyish stone-colour, dnsted with grey ; basal line very obscure,
marked by dark dots on veins; cell-spot a small dark jwint; median shade from
two-thirds of costa, curved to submedian fold, then vertical ; outer line at five-
sixths, marked by dark dots on veins ; marginal spots black ; fringe concolorous
or slightly paler.
Illndirimi : the same ; in one example the cell-spots eilged with black scales.
Underside i)ale stone-colour, with slight rosy suffusion in forewiiig ; outer line
marked in both wings; black marginal triangles in both.
Head, thorax, and abdomen like wings ; face above dark fuscous, jale below;
palpi reddish fuscons above, pale below ; abdomen at sides with rosy stains.
Expanse of wings : 3-5 mm.
2 ? ?.
T4. Perixera 0) stabilata spec. nov.
ForririiKi : stone-colour, tinged with i)aie fawn and finely dusted with ibirk
atoms ; basal line marked only by minute dots on veins; outer Hue from three-
fourths of costa to two-thirds of inner margin Innulate-dentatc, but only the dark
teeth on the veins distinct, preceded by a faint darker median shade parallel to it,
and followed at the same distance by the shade preceding submarginal jiale line,
marked by slight dark dots between the veins, more conspicuous on each side
of vein 5 and forming on the subniedian fold a black V-shaped mark ; the shade
following the snbmarginal line being also marked with dark scales above inner
margin ; cell-spot small, pale, with a few dark scales round it ; marginal dots
black ; fringe concolorous.
HimlwiDq : similar ; hindmargin with tine dark dots at the ends of veins
as well as between.
Underside jialer, with fine dark striations ; cell-sjwt of forewing dark ; only
the outer line distinct, marked with black ])oints on veins.
Head, thorax, and aljdomen concolonius with wings : face and pulfii dark
fuscous above.
Expanse of wings : 4>> mm.
IV. •
T.i. Perixera subrosea spec. nuv.
Foi-inriiKj : dull brick-red, thickly dusted with olive fuscous; basal line
generally obscure, marked by darker points on veins and slightly curved outwards
above and below median ; cell-spot whitish, surrounded with a few dark scales;
median shade at three-fifths, iui'urved below middle, obscurely lunulate-dentate ;
outer line at five-sixths, marked by distinct black vein-dots ; black marginal dots
between veins ; fringe concolorous.
Uimla-imi : similar: the cell-dot snmetimes ringed with black scales from
subcostal vein.
( n2 )
Umlersidc iU'f|i ilull rusy, sniiicwlint piilcr, luorc ycllo\vif;li, in Ijindwiiic,
especially towards abilniuinal luargiii ; tli(^ liiiidiUiirLiiri with dull red triangles
between veins ; the onter line alone marked.
Head, thorax, alidoiuen, and antennae concoloruns ; ]iai|ii litdow, extreme base
of face, abdomen beneath, and legs ]iale yellowish ; Ibrelegs red ; liindtibiae with
a thick fringe of red hairs, the femora with yellower hairs.
Expanse of wings: ;i.j mm.
4 66,4 ??.
76. Problepsis magna s|i('c. nov.
Fnrr/r/m/ : white: costa grey ; markings as in /'/7)/>/ry«/(;f/c.'( i:()i'J'///rtivaWa,rr.
from India; the ocellns large, nearly round, the outer orbit olive ochreons ;
its interior above vein 4 white, with tine blatk scales ; the lower part velvety-
black, crossed by the ochreons veins :> and 4 ; a clondy grey median shade,
visible at middle of costa, rnns vertically from the ocellus to three-tifths of inner
margin; the base of inner margin largely spangled with silvery scales; outer line
ochrcous-grey, followed by a macnlar grey band ; a thin grey shade before the
very fine black marginal line ; fringe silvery-white.
Iliiidirimi : with outer half of ocellus only, the orbit being sim])ly exterior ;
the inner edge straight and formed of silvery scales; tiie outer area also silvery,
but mixed internally with a few brown scales ; the orbit continued to two-thirds
of inner margin, where there are some silvery scales.
Underside white.
Palpi and face white below, black above; vertex black; thorax white;
abdomen grey above with brown semi-lustrous dorsal spots, white below ; antennae
ferruginous.
Expanse of wings : 46 mm.
77. Ptychopoda bipartita spec. nov.
Fore/ri/ii/ : glossy whitish ochreons overlaid with pale grey; the outer line
and two sulmiarginal shades sinuous and distinct; the basal curved and obscure;
a red-brown straight line at middle touching the black cell-spot ; fringe pale, with
dark dots at base beyond veins.
Iliiiilirimj : similar ; the brown line in front of the cell-spot.
Undersidi' )iale oclireous, with the lines broadly greyer; median shade olive-
brown, hardly touching cell-spot in forewing; in hindwing angled at oell-spot.
Thorax an<l abdomen like wings, but darker grey ; face and pal[ii black.
Ex()anse of wings: 17 mm.
1 cJ,-' ? ?.
78. Ptychopoda exempta spec. nov.
Forewing : shining greyish ochreons ; basal line curved, at one-fourth ; median
oblique from two-thirds of costa to before middle of inner margin, preceded liy a
black cell-spot : onter line finely Innnlate-dentate ; praesubmarginal shade swollen
and Innate below middle : fringe paler, with dark dots beyond veins at the hase.
lUiulwiKij : without basal line; cell-spot hirge, black; outer line strongly
insinuate on the folds.
( !'3 )
Underside yellow ochreous, with the lines and shades grey ; cell-siiot» black,
distinct.
Vertex, thorax, and abdomen lilce wings ; face and imlpi black.
Expanse of wings : Ifl mm.
3 ? ? .
Subfamily HYDT?I0MENINAE.
TO. Anticlea semiflava spec. nov.
Forewing : purple, the lines green ; edge of basal patch formed by a sligiitly
curved green line with a fine purple centre ; inner edge of central ftiscia sharply
angled inwards on median vein, and as sliarj)ly outwards on the two folds, green
like the basal line, with a purple line near its enter edge ; outer edge of fascia
sharply indented on vein 7, insinuate beyond cell, with a blunt double projection on
vein 3, vertically waved from vein 2, the green band following with its inner edge
white ; a vertical whitish band with a waved jmrple line on its outer edge from
middle of costa to median vein, enclosing the linear black cell-spot, and joining
three greenish streaks between submedian fold and vein 4 ; snbmarginal line
waved, greenish, very faint, preceded and followed in U])per half of wing bv
a large greenish patch with a bluisli-white centre; pairs of black s])uts at the
end of veins ; fringe purple mottled with green.
lliiidiviiKj : deep yellow, with base greenish and fringe dark grey.
Underside of forewing mouse-colour, dusted with pale along costa, the inner
margin whitisli ; cell-spot a blackish curved line ; hindwing mousc-colonr tinged
with reddish and thickly dnsted with yellowish ; a dark curved snbmarginal line ;
cell-spot dark.
Head and thorax green ; abdomen purple sprinkled with green ; vertex, collar,
shoulders, and patagia green with spots of purple ; abdomen beneath and legs
purplish sprinkled with green scales.
Expanse of wings : 30 mm.
2 S3. ~
^'1. Chaetolopha ('0 antennata sjiec. nov.
Voreiriiiij : cream-colour, covered with pale green scales ; the lines violet ;
basal line at one-fourth, angled outwards ou median vein, and incurved below ;
edges of central fascia comjiosod of narrow violet bands; the inner curved, at two-
fifths, projecting slightly basewards at median and submedian veins; the outer
i'rom three-fourths of eosta to two-thirds of inner margin, toothed outwards on
veins 3 and 4, then incurved, the fascia below middle wholly violet : some violet
scales along the conrse of submarginal line.
Hiridtrinfi : paler, with a single violet line across middle.
Underside like upper, but less distinct.
Head, thorax, and abdomen pale varied with green: dorsum with indications
of violet bands.
Expanse of wings: 15 mm.
, 1 S.
The single example is much worn : I have placed it temporarily in Chactolojj/ia
Inmi its superficial likeness to ('. sj//f/ii/e/is, but it will probably require a separate
( !»4 )-
^eiius ; till' liindiuargiii of torewing is siunatp ami cueniiliito, the auteiiiiae long,
with angled segments, and long cilia. The anal segments of abdomen are swollen,
as often in ('liiirtnlniili<t, lint the neuration is that of F.iicijmntogi-.
>>!. Chaetolopha flexilinea spec. nov.
Vorcivimj : brownish fawn-e.olonr, the cell and si)ace beyond to hindmargin
jialer ; a slight brown line at one-fifth, vertical, inbent on costal and snbmedian
veins ; inner and onter lines jiarallel thronghont to each other, from costa at
two-fifths and two-thirds, oblicjne inwards above subcostal and below median veins,
bent ontwards between, dark brown, edged conversely with white, the space
between them darker brown: cell-spot dark brown, vertical; siibmarginal line
forming dark bi'own lunules, interrupted beyond cell and below costa; a dark
marginal line interrnpted by the veins ; fringe concolorons, the tips white-spotted.
liuidirhn] : wliitish ochreons, browner along hindmargin, with a faint outer
line; fringe dark grey.
Underside duller, tlie hindwing speckled ; cell-spots and outer lines jilain in
both wings.
Head and thorax like forewings ; abdomen paler, more sjieckled.
Expanse of wings : •i\ mm.
2 ??.
82. Chaetolopha fulgurata spec. nov.
Voreinmj : bright fulvous, suffused with darker : the cell and space beyond
to subniarginal line j)alcr ; a slight spot at base and a narrow waved vertical line
near base brown with a few white scales ; inner line at one-third, snow-white,
outwardly edged with black-brown, minntely dentate outwards on the veins ; outer
line at two-thirds, parallel to hindmargin, but with a strong shoulder on vein 4,
snow-white, inwardly edged with black-brown ; submarginal line brown-black,
marked by an oldiqne snow-white dash above vein (!, and below vein 4 acutely
dentate, followed by some white brown-speckled scaling, the veins across it brown ;
above vein 4 the line itself is interrupted, but followed by horizontal brown streaks
between the veins ; fringe whitish, with brown chequering beyond veins ; cell-spot
dark brown, vertical.
Ilii<(hrhui : fulvous, much paler towards base, with a darker angled middle
line and traces of a submarginal line, especially at apex, which is speckled with
grey.
rnderside dull fulvons, the lines showing brown, and only the snbapical dash
white ; hindmargin grey-brown, with ]iale dusting ; hindwing with large brown
cell-Kpof and brown angled jiostmedian line ; the marginal area brown with grey
dusting, and a dark snbmarginal line: the brown-grey scales form a shade also
along snbmedian \\\]i\.
Head, thorax, and abdomen deej' fulvous; metafliorax with a snow-white
vertical line; first segment of dorsum brownish.
In the i the basal two-thirds of hiiidwiiig beneath is cldtlied with fine
furry hairs.
Expanse of wings : ".'(i mm.
•■) 66, 1 ?.
Superficially greatly resembling CIhh'IoIojjIhi huwojjliruymu Meyr. from Australia,
but in that species the snbapical white mark is horizontal.
( 95 )
^o. Chaetolopha pictipennis spec. imv.
Forewiiii/ : purple plnm-colonr ; the markings deep lirowii, edged with orange
n,iid yeUow lines ; basal patch witli the edge vertical ; inner edge of central fasria
nearly ])arallel to it ; outer edge sinnous, from three-fourths of costa to three-fourths
of inuer margin, bent in below vein 0 ; submarginal line waved, orange, from costa
just beyond outer edge of ftiscia-, outcnrved and reaching inner margin before anal
angle, preceded by a plum-coloured band, the inner edge of which is convex
basewards and runs to apex, crossing and interrupting submarginal line, with a
yellow spot at apex and another where tlie lines separate on vein (i ; fringe
]dum-colour.
Hiiuhriiuj : coppery red, dusted with blackish fuscous ; a straight dark
jiostmedian line, edged by an nnspeckled space of red before the dark hindmargin ;
fringe dark.
Underside dull coppery red, the markings brown-black edgeil with brighter red.
Head, thorax, and abdomen purple.
Expanse of wings : ".'U' mm.
84. Chaetolopha rectilineata.
Pnh.,„„<( icri;i;,„'al,i Warr., Xar. Z.,..l. v. p. 2411 ^ (1898),
This s]iecies must be transferred to Chaetolopha.
85, Chaetolopha ruptistriga spec. nov.
ForeiciiKj : dull chestnut-brown ; a large wedge-shaped area of dull purplish
grey running through wing from base of cell to hindmargin, interrupting the lines ;
a snow-white inwardly oblique bar at base, edged with black-brown, from costal
to median vein ; inner and onter lines at two-fifths and two-thirds, snow-white,
edged conversely with black-brown, oblique from costa to subcostal vein,
reajjpearing below median vein, as similar, but more oblique, streaks to inner margin ;
submarginal line denoted only by a more obscure white streak from inuer margin ;
veins close to hindmargin chestnut, interrupting the dark marginal line ; fringe
dark grey, the onter half whitish.
Ilindwiny : pale ochreous, with a curved grey central line, becoming jiale
fulvous before the darker hindmargin.
Underside dull grey-brown, greyer along hindmargins ; costa of forewing
fulvous ; hindwing with two bent brown lines, the space between them fulvous.
Head, thorax, and abdomen dark lirown ; a snow-white, jiartially interrupted,
line from tips of pali)i along face, thorax, and dorsum.
Expanse of wing.s : 24 mm.
2 $S,\ ?.
In the ? the brown is ])aler and brighter.
86. Chaetolopha splendens spec. nov.
Forewiny : glossy pale straw-yellow ; lines and veins bright brown, also the
costa at base ; basal line vertical at one-fourth ; inner edge of central fascia at
two-fifths, outcurved on submedian fold; outer edge at two-thirds, projecting
bluntly at vein 4, and insinuate below, the two edges closely approximated on the
fold ; space between, like the basal patch, with brown suft'usion and lines : space
beyond basal patch traversed by a brown line, a thick subaiai'ginal line, forking at
( 96 )
vein li to ajiox, followed liy an obscurely edged line of lirijiht liiiiales ; fringe
glossy yellow with brown lines beyond veins.
IHiwUrinq : pale yellow, lines of uudersiJe sliowing through; fringe ]>alo
yellow.
Underside of foiewing Hushed with pale tawny : costa black-spoekled : the
lines fnlvons ; hindwiug with tliree ('(piidistant fulvous lines, the outermost forked
to apex.
Head, thorax, and abdomen pale yellow flushed here and there with brown ;
tarsi dee]) brown with pale joints.
Expanse of wings : S IS mm. ; ? 22 mm.
1 cJ,2 ? ?.
87. Chaetolopha tristriata spec. nov.
Voi-i'u-imi : pale olive-grey, dusted with darker ; a dark olivo-green shade at
one-third, inw.ardly darker an<l edged with pale, outwardly diifuse ; a similar shade
at two-thirds, slightly nearer first on inner margin than on costa; a submarginal
irregularly waved dark line, inwardly pale-edged; veins towards hindmargin dull
])ink : fringe pinkish ochreons, with darker chequering beyond veins ; costal edge
and snb('ostal vein dull pink ; cell-spot olive.
Illntlwinq : dnll greyish pink, more olive towards base, crossed about niidille
by a thick dark paler-edged shade ; fringe pink.
Underside of forewiug ))inkish brown, the veins pinkish brown, those below
middle reaching margin as wedge-shaped marks ; hindmargin between the veins
olive-green speckled with whitish scales, at apex more whitish ; middle line
showing on costa as a yellowish streak, outer line as a row of wiiite spots ; fringe
brown-pink with dark checpiering : hindwiug white, in basal area densely sprinkled
with olive scales, the white showing clear only before the dark brown cross-line,
which is followed by a brownish ochreons and then an olive shade ; marginal area
olive, with a dark waved submarginal line preceded by a whiter spac'e ; inner
margin with its fringe brown.
Head and thorax grey ; dorsum dull red, with the margins ol' segments aN<l
anal segment greyish white ; legs red-brown, with pale joints.
Expanse of wings : c? 2(i mm. ; ? 30 mm.
1 c?,3 ? ?.
The ? is a little paler than the c?. The species is luucli like I'lOjiitliiw
allci iKilii W'arr. from l!ou Islaiiil, liut liirger.
8><. Coenocalpe augustipeunis siicc. uov,
l\trv)ni((j : greyish white, the costal half dift'usely fawn-brown ; the subcostal
vi-in with splashes of white to middle; crossed by obscure oblique dark lines,
lunulate outwards and dentate inwards on the veins, and only distinct towards
inner margin; the veins faintly dotted dark and light: the three ]irinciiial lines
double ; a paler costal space in centre of fascia continuing the small lilaek cell-sjiot ;
in the enter line a black dash on snbmedian fold: nuirginal festoon dark; fringe
chequered dark and light grey.
Ilimlwimj : wliiter, with no fawn-coloured suffusion ; the lines fairly distinct ;
fringe whitish.
Underside fawn-grey, shining ; costa whitish ; lines dark only on costa of
forewiug ; fringes white with. dark mottling.
( 97 )
Head, thorax, ami abdomen like wings ; the head and thorax somewhat
darker ; abdomen beneath, jjectns, and legs whitish ; Ibretarsi blackish, with the
joints pale ; palpi externally dark.
Expanse of wings : 30 mm.
4 66.
The wings are peculiarly narrow ibr the genus.
89. Coenocalpe hirtivena spec. nov.
Forewiiuj : pale fawn-grey, dnsted with dark atoms ; crossed by a succession
of brownish grey lines all hinulatc outwards and dentate inwards on the veins ; the
inner, outer, and jjraesul (marginal double and accompanied each by a brownish
grey shade ; the luuules along the costal area all clear and well-detined ; all the
lines forming blackish dashes across the veins, alternating with the paler inter-
spaces ; median vein with linear patches of raised black scales to the end of cell ;
cell-spot black, oblic^ne ; snbmarginal line wavy, whitish, sometimes with a darker
shade on both sides ; marginal festoon black, joined between the veins by black
dashes with the teeth of snbmarginal line ; fringe concolorous.
In the ¥ the shade beyond outer line forms a dark blotch beyond cell.
llindidmj : similar ; the hindmargin strongly crennlate.
Underside much paler, especially towards base, with two postmedian and a
submargiual dark curved shade ; cell-spots black ; costa of forewing cream-colour.
Head, thorax, and abdomen like wings.
Expanse of wings : 3s mm.
1 c?,3 ¥ ?.
The costal shoulder of forewing is roughly fringed with hairs.
Ot). Coenocalpe semirufata spec. nov.
ForciciiKj : pale vinous ; the lower half of central fascia fuscous, also the
marginal area between veins 4 and 5 ; marginal area between veins 2 and 4 white ;
inner edge of central fascia from one-third of costa, outcurved above median and
slightly again below, then oblique inwards to one-third of inner margin ; outer edge
from three-fourths of costa, lunulate-dentate, the teeth pointing basewards, to three-
fourths of inner margin, roundly biiobed between 2 and 4 ; closely preceded and
followed by the darker line of the pale fasciae ; snbmarginal line marked by white
dots between the veins ; marginal line crenulate, dark red, interrupted by pale spots
at the veins and preceded there by patches of pale scales ; fringe vinous.
Ilinthcing : with the central fascia much broader, only its edges fuscous.
Underside grey, glossy ; cell-spots and lines darker ; fringe vinous.
Head, thorax, and first two segments of abdomen pale vinous ; rest of ulidomen
fuscous and vinous mixed, the third segment sharply blackish fuscous ; face and
palpi deej) vinous ; legs externally fuscous, the tarsi black, with pale joints.
Exjianse of wings : 33 mm.
3 66.
'.)I. Coenocalpe ustimacula spec. nov.
Forewing : dark ashy grey, the costal half blackish grey, with the pale intervals
forming white costal patches ; cell-spot black, lying in the middle pale sj)ace ; lines
all Innulate outwards and dentate inwards, the outer and praesnbmargiual blackest,
( 98 )
forming black jiuiiits lowarcls inner margin ; siibmargiiial jiaif, iireci'ilcd and lollowid
by black sagittate markings ; fringe black bevDud veins, jialer between.
Uimhi-incj : with costal half pale grey, inner-margiual half blackish.
Underside shining, iron-grej' ; costa of forewing pale ; cell-spot and the ends
of lines of costa black ; fringe black and white.
Head, thorax, and abdomen blackish ; basal segment of abdomen and meta-
thorax pale grey, the metathorax with a thick black angular mark ; shoulders and
patagia with a pair of minute white dots; basal half of abdomen beneath, pectus,
and legs white ; foretarsi black, with white joints.
Expanse of wings : 32 mm.
2 c?c?.
In the tyi)e specimen the costa of forewing and centre of hiudwing are tinged
with reddish fawn-colour.
92. Coenocalpe xylinata spec. nov.
Forewing : wood-brown, the markings slightly darker, but very indistinct ; the
lines forming dark and pale dashes on the veins ; a paler brownish space at apex ;
submarginal line marked by jiale dots between veins, followed by black dashes to
the margin ; fringe brown.
llinduiiig : paler, with the lines more plain ; two before the dark cell-spot
and two beyond, closer together, followed by a blackish band projecting outwards in
middle ; three outer lines also angled and marked black and light on veins ; fringe
brown.
Underside greyish ochreous ; the lines all darker and the cell-spots black.
Head, thorax, and abdomen pale and dark brown ; palpi and face below
blackish ; anal segment of abdomen paler.
Expanse of wings : 35 mm.
1 ?.
93. Collix examplata spec. nov.
Forcicing : fuscous, with darker fuscous lines and shades ; all the markings
confused ; the darker lines forming blackish dashes on veins alternating with pale
ochreous ones ; cell-spot large and blai:k ; the base of veins 3 and 4 shows as a
pale ochreous space ; the submarginal line is preceded by blackish patches at costa,
beyond cell, and towards inner margin ; a paler space from outer line runs between
veins 0 and 7 towards apex ; black curved marginal lines between veins, with a
pale dot at the veins ; fringe dark fuscous.
Ilimhclng : with less dark suH'nsion, and the markings therefore plainer ; a
curved dark antemedian line followed by a sinuate black cell-spot ; three waved
central lines, followed by a pale band with dark centre ; marginal area dark fuscous
with obscure pale submarginal line ; the disc reddish tinged.
Underside rufous ochreous ; cell-spots black ; two outer series of dull blackish
semi-coniiuent spots on veins ; the outer one interrnpted between 3 and 4 ; fringe
blackish.
I'alpi reddish grey, terminal segment and the base of second segment fuscous
head, thorax, and aliduinen like wings.
Expanse of wings : 4U mm.
1 ?.
( 99 )
94. Crasilogia dispar ab. simplex uov.
Aloiii; wi(.li a fair number of t^vpical ('. dispar Warr. of both sexes 1 find 2 cj cC
of remarkably aberrant structnre ; of tbese one, of normal size, has the forevving
exactly like that of the S type, the other, much smaller, has the forewing almost
exactly like that of the ? ; in both cases, however, the hindwings are shaped and
coloured as in the ? , without a sign of any of the secondary sexual characters that
apjicar in the normal S, and their neuration is that of the ?. In both cases the
retinaculum of the forewings and the fine frenulum of the hindwings is distinct ;
besides which the condition of the two insects precludes any suspicion of mending.
95. Crasilogia? fumipennis spec. nov.
Foreiomg : ochrcous, suffused throughout with brownish grey ; the basal area
and central fascia brownish fuscous, the lines of the latter forming three dark bands
with fainter intervals ; the edges crenulate, the outer deeply insinuate beyond cell
and projecting on vein 3, finely edged with whitish ; three dark waved Hues alter-
nating with paler ones before the submarginal, which is yellowish and lunulate-
dentate, both the luunles and teeth projecting outwards ; a black blotch before it
between veins 6 and 7, passing into apex, and the lines before it blackened and
confluent between 3 and 4; the spaces between 5 and 6, and 2 and 3 filled with
black and white scales ; pairs of square black spots at the ends of the veins, which
are yellowish ; fringe chequered pale aud dark brown ; cell-sj)ot black ; the dark
inner edge of central fascia extended to basal patch above submediau.
Ilimlwing : smoky fuscous, with the postmedian area fulvous ; traces of two
or three curved shades from inner margin ; costal area whitish.
Underside ochreous, suffused in forewing with grey, with all the lines dark
grey, becoming black on the costa, which is yellowish ; hindwing clear ochreous,
with three thick grey lunulate shades sejiarated by finer lunulate lines.
Head, thorax, and abdomen brownish fuscous intermixed with jjaler ; abdomen
beneath and legs ochreous ; fore and middle tibiae and tarsi black, with yellow joints.
Expanse of wings : 39 mm.
1 ¥.
Though very different in a])pearance above from Crasiloijia disjxir, I refer it to
the genus from the great resemblance in the marking of the underside.
96. Diactinia intromissa spec. nov.
Forewing : with the lines snow-white, the basal patch and central fascia
blackish fuscous ; the former edged by a fine white line, and in one specimen
crossed by a whitish line ; the edges of central fascia liroadly white, meeting on
the submediau fold, then separating again and forking symmetrically to inner
margin, as a double curve denticulate inwards on vein 1 ; the inner edge is also
dentate into the fascia along the cell-fold ; the interval between basal ])ateh and
fascia, aud the lower imrtion of the fascia itself, are olive-grey speckled with black ;
below costa beyond the fascia the commencement of a line is shown by two white
angulated marks filled in with blackish, continued below as an olive ochreons
somewhat lunulate shade edged by a black clond ; between veins 7 and 4 on the
margin is a bilobed blackish blotch edged by a white line, which runs out into
the fringe between 3 and 4, and appears again as a narrower line curving to anal
( 100)
anj;le : marjjiiial line ilurlc, iiitcrrnptcHl a,t thu veius : Cringe olive ocbrcou.s, with
dark middle line and dark mottling bcvond veins.
Ilindicing : wLitisb, tinged witL olive ochreous, and striated with grey; traces
of dark lines oq iuuer margin and a thick streak before margin from aual angle to
vein 4.
Underside yellowish speckled with Maekish ; I'urewing with inner margin
whitish.
Head, thora.x, and abdomen wliitish ochreou.s spoeklud with olive ; underside
of abdomen like that of wings ; legs black and yellow.
E.Kpanse of wings : 34 mm.
1 c?, 1 ?.
The ? is somewhat paler than the S. In the i'orewing tlie hiudmargin is
slightly indented from vein 7 to 4.
'JT. Ochyria dilataria spec. nov.
Foiricing : basal patch and central fascia dark brownish fu.scous, their edges
deeper ; the intervals slaty grey ; the central fascia broader than in O.fulcistriga,
its edges more waved, and margined by first a lustrous grey acd then a ferruginous
orange line ; from this line above vein 0 a narrow ferruginous streak runs oblii^nely
to apex ; centre of fascia dull violet-grey, containing a rather large blackish cell-spot
surrounded with dull rust-coloured scales ; submarginal line waved, dull bluish,
preceded by a darker clond at costa : fringe (worn) slaty grey.
llinthcuKj : pale grey, darker towards hindmargin.
Underside jiale grey, darker in outer half of wing, without distinct markings.
Head, thorax, and abdomen dark fuscous ; segmental rings of abdomen deeji
ferruginous.
Expanse of wings : 17 mm.
1 <^.
The neuratiou is abnormal ; vein 11 appears to form the coulinuation of 10,
and is joined to 12 by a bar ; vein 'J is coincident with lO, and subsequently
anastomoses with 7, 8.
^"i. Ochyria fiilvistriga spec. nov.
ForewiiKj : basal patch, central fascia, and praesubmarginal band ilark fuscous ;
the intervals dull purplish grey; all the dark markings edged by broad dull
ferruginous violet lines ; centre of central fascia traversed by a broad band of
purplish grey, containing the black ccll-sjwt ; a ferruginous violet obliijue streak
above vein 0 from outer to submarginal line, which is waved and dull lilue ;
marginal line black ; fringe fnseous, with the basal half darker.
Ilindwimj : purjilish grey.
Underside dull shity cinereous; hindwing slighlly spriuldod willi bluish scales,
with a black cell-spot, and traces of lines.
Head, thorax, and abdomen dark fuscous, like wings.
Expanse of wings ; 6 10 mm. ; ? IS mm.
3 cJ d, 5 ? ? .
!)'••. Ochyria pulchella spec. nov.
Fori'icing : pale olive-green, crossed by broad white slightly lustrous lines ;
two, indistinct, near base; two on each side of the middle of wing, bent outwards
above, then vertical; and one .submarginal, interrupted by (he veius; the pale lines
( 101 )
are preceded irregularly by dull iintroiis purplish scales, wijicli beyond cell form
a blotch before the submarginal ; fringe pale green ; cell-sjiot linear, oblique,
purplifli.
UimhciiKj : whitish grey, darker and reddish tinged towards hindmargin.
Underside dnll rosy, speckled with grey ; forewing with v/hite snbmarginal
line; hindwing more speckled, with dark cell-spot and three dark white-edged lines.
Head, thorax, and abdomen pale olive-green.
Expanse of wings : l(i nun.
1 ¥.
100. Ochyi'ia rubecula spec. nov.
Forewing: pale brick-red; the markings very dark green, miuntely edged
with white, the basal patch small ; the central fascia broad, swollen below median,
with two outward-jjrojecting teeth between 2 and 4, and much narrowed on inner
margin ; the centre of the baud preceding it is slightly marked with green on
costa and inner margin, and a large costal blotch with a smaller one below it
on vein 0 precedes the submarginal line, which is finely white, the luiiules followed
also by some dark green scales above inner margin ; a very fine dark marginal
line inwardly white-edged ; fringe brick-colour.
llimliciii;/ : pale grey ; the fringe jjinkish.
Underside of forewing dull pinky grey, with the markings showing throngh ;
hindwing with faint grey antemedian, postmedian, and submarginal bands
alternating with paler spaces each intersected by a fine line.
Head and abdomen pale brick-red ; palpi, thorax and forelegs blackish green.
Expanse of wings : 17 mm.
1 c?.
101. Ochyria sixbcaesia spec. nov.
Forewing : slaty grey ; the basal patch and the central fascia dark fuscous ;
the paler bauds sprinkled with greenish yellow scales; the edges of the dark
markings dull blnish; central fascia crossed also by two dull bluish lines; snb-
marginal line waved, dull blue; fringe like the intervals greenish grey ; sometimes
a dark shade precedes the submarginal line.
JHndicirig : dull slaty grey ; fringe as in forewing.
Underside of forewing dull blurred cinereous ; costal and hindmargin with
some bluish scales ; lines visible along costa only ; hindwing sprinkled with dull
blue scales, forming a double outer and waved snbmarginal line ; the dark lines
clear across wing ; cell-spot blackish.
Head, thorax, and abdomen like wings, all sprinkled with blnish scales.
Expanse of wings : ,J IV, mm. : ? IS mm. One small 6 expands only 13 mm.
5 c?c?,3 ?¥.
102. Ochyria unitaeniata spec. nov.
Forewing : purplish fnscons, crossed by some fine wavy darker lines ; at
one-fourth from base is a slightly paler band, bent on subcostal, separating basal
patch from central fascia, which contains a large cloudy dark cell-spot, and is
edged at four-lifths by a dark line followed by a white line and a pale green
band ; marginal area beyond dark jiurplish fuscous ; a pale green patch at anal
angle, separated from the green band by a narrow dark space edged by the whiti.sh
( 10-' )
waved snbmarginal line, wliioh is not visilile aliovc uiiddlo of winf^ ; some dark
margiual liuiules before the gre}' fringe.
Hind winy : greenish white, without markings except ou inner margin lielow
the median and vein 3, the band above anal angle being prominently dark.
Underside of forewing dnll grey ; the onter line pale ; the marginal area
darker ; liindwing whitish green, crossed by six waved grey lines, of which the
onterraost is dark before anal angle ; cell-spot black.
Head, thorax, and abdomen fnscons.
Expanse of wings : 18 mm.
1 <3.
Spectrobasis gen. nov.
ForeiciMi : costa straight, but with a fiiint shoulder at base fringed with hair,
and curving before apex, which is blunt ; hindmargin simply curved.
Iliiidwinji : with hindmargin well rounded ; both angles rounded.
Antennae of S bipectinate to near apex ; of ? simple, tlie segments angulate ;
palpi rostriform, porrect, rough-haired, the terminal segment smooth, decumbent ;
tongue slight ; frenulum present ; hindtibiae with four spurs.
yeuratioii : forewing, cell about half of wing; discocellular vertical ; all three
median nervnles rising close together; radials normal ; 7, 8, 9 stalked; In, 11
separate, 10 anastomosing with 11 and again with 8, 0, forming a double areole :
biiidwing, discocellular biangulate, the radial from tbc lower angulation ; 7, 8
anastomosing for three-fourths of cell ; (!, 7 stalked ; vein ~ from nearer base
than in forewing.
In the (? the space between base of 7 and 8 contains an elongate hyaline fovea.
Type : Spectrobasis nifa spec. nov.
An endemic genus without any apparent close affinities.
Iu3. Spectrobasis rufa spec. nov.
Forewing: dark olive-grey ; with obscure darker cross-lines, visible chiefly on
costa, which at base is dull brick-red to median vein ; basal patch, central fascia,
and centre of band between them rather darker; the pale bands edging them pinkish
white on costa, with a darker central line ; the outer edge of central fascia from
just beyond middle of costa curved outwards below middle to two-thirds of inner
margin ; submarginal line wavy at costa only ; a dark cell-spot ; fringe dnll pink,
with dark olive-grey mottling.
lUmJwintj : dark olive-grey, without markings ; fringe as in forewing.
Underside of forewing blurred purjilish grey ; the costa and hindmargin
red-brown, the submarginal line showing white teeth below costa ; costa from base
more ocbreous in c? : hindwiug red-brown speckled with pale, with darker ante-
median and median reddish bands ; a waved submarginal line with red-brown
blotch above anal angle, and showing white spots between veins; in the S
the basal half is flesh-coloured ochreons speckled with brown ; a flesh-coloured
blotch on inner margin beyond middle band, and often a second towards costa ;
fringe reddish.
Head parts reddish mixed with pale ; thorax and abdomen dark olive, the
dorsal segments often broadly pale.
Expanse of wings : 20 mm.
A long series,
( 103 )
104. Spectrobasis viridis spec. nov.
Foreicing : reddish and olive-gre}% the tints mneh mixed, the liaes and bands
being darker and the intervals ou costa whitish ; costa at base dnil moss-green,
beyond dotted, dark green and pale ; a darlc cell-spot ; outer edge of central fascia
from two-thirds of costa, ontcnrved, to two-thirds of inner margin ; fringe whitish
with dark green cheqnering.
Bindwing : dnll greenish gre}-, with a reddisli tinge, paler along costal area,
the bands of underside showing throngli ; fringe as in forewing.
Underside of forewing blurred ; the costal and hindmargins green, with
paler markings ; hindwing white sprinkled with green, forming three green bands
separated b}- whitish bands, tlie green bands often varied with dark purplish scales.
In the larger ? the costal and hindmargin of forewing and most of the scaling
of hindwings is reddish instead of green.
Head and abdomen pinkish grey, with Vlarlc speckling ; thorax mainly dark
green, varied with paler.
Expanse of wings : 20 mm. ; the redder ? 28 mm.
2 c?c?,2 ??.
Evidently a very variable species.
lii.j. Xanthorhoe albiapicata spec. nov.
Fon'w/'nr/ : blackish fuscous from base to outer edge of central fascia, the
band beyond basal area always greyer, sometimes witli pale dots along its edges ;
the baud beyond central fascia brightly white to below vein 4, where it joins a
white lunule in the submarginal line, sometimes running pale to hindmargin and
out into tlie fringe; submarginal line rarely distinct; the lunule above vein 6
always pale, bluish grey ; marginal line black ; fringe blackish, mottled with
jialer ; the marginal area round apex, beyond cell, and broadly below vein 3
dark fuscous, sometimes blackish. In the c? the white of the pale band is more
extensive than in the ¥ .
Uiixlwiiig : dull blackish fuscous, with the lines obscure, but paler on inner
margin.
Underside glossy cinereous, sprinkled with whitish in hindwing ; the lines all
dark but the submarginal, wliich is represented by white spots, that below 4 large.
Head, thorax, and abdomen blackish fuscous.
Expanse of wings : 20 mm.
A long series of both sexes.
106. Xanthorhoe bifulvata spec. nov.
Forewing : with basal patch, central fascia, and shade preceding submarginal
line brownish fuscous ; basal patcli edged and crossed by a darker shade ; central
fascia edged with darker fuscous, with a deeper band on its inner edge and two
wavy dark lines before the outer, with paler grey lines between ; basal patch and
central fascia edged by silvery grey bands with a dark centre ; space between the
inner two filled up with fulvous grey ; band before snbmarginal line fulvous
tinged in its inner half, paler between veins 3 and 4; snlimarginal line wavy,
pale grey, broadly silvery grey just below middle ; marginal area dark and light
grey ; a row of dark marginal lunules between veins ; fringe mottled dark and
light grey.
( 104 )
Tliiiihrinq : flark fuscous, with traces of curved wavy lines, whicli arc clear
only oil inner margin, where the paler intervals are sometimes silvery grey ; fringe
of inner margin whitish.
Underside of forewing dnll olive fnscnns, tinged with bine-grey ; the lines
shdwing darker; hiiidwing with all the intervals dull lilnish white, the shades
olive fuscons.
Head, thornx, ami alidonien fuscons, varied with liliiisli grey scales ; nietathorax
bine-grey.
Expanse of wings : 28 mm.
5 c?(?, 3 ??.
107. Xanthorhoe cerasina spec. nov.
Foreicing : with basal patch and central fascia velvety blackish fuscons, the
interval between them and the marginal area bright cerise ; the blackish areas
are finely edged with white ; in the antemedian band are three fuscons grey linos,
and the fascia is followed by three ; the extreme hindraargiii beyond the waved
snbraargiual line is fuscons grey, and all the lines and shades of the marginal area
are fuscons at costa ; marginal spots black, followed by pink spots running out into
the fuscous fringe ; cell-spot deep black.
Hindicing : fuscous, with the lines of central fascia darker, as far as outer
line, then dull cerise ; the apical half smoky fuscous, and all the lines waved.
Underside dark fuscous, with all the lines indicated ; the outer lines marked
pale across wings, and white on costa of forewing.
Head, thorax, and abdomen all dark fuscons ; anal tuft of <S black.
Expanse of wings : 44 mm.
A long series of both sexes.
Both surfaces are glossy.
I<i8. Xanthorhoe coenileata spec. nov.
Forewiiiq : dark fuscons, crossed by three bright bluish white double lines :
one close to base, the second at one-third, both incurved in middle of wing ; the
third at two-thirds, almost interrnjited between 2 and 4 by the projection of the
central fascia ; submarginal line single, whiter between 3 and 4 and preceded there
by a fulvous patch ; dark marginal lunnles, followed by white sjiots at the base
of the fuscous fringe; the spot between 3 and 4 and that below apex large and
white, 1 nulling throngli the fringe, and each preceded by a submarginal whitish
spot ; a faint jiale line at middle of central fascia.
liniilirintj : greyish fuscous, with traces of pale and dark lines on inner margin
only ; fringe dark, with the two white spots at middle and apex.
Underside of forewing blurred greyish fuscous ; fringe with two white sjiots;
hindwirig with faint bluish white markings ; two diffuse lines at base, a double
waved postmedian, and a single submarginal line.
Head, thorax, and abdomen brownish fuscous, the tips of shniildcrs and jiatagia
and the segmental rings of abdomen with bluish white scales.
Expanse of wings : 30 mm.
1 (?.
K'9. Xanthorhoe fulvinotata spec. nov.
Forewing: white; the markings grey and black, the basal patch and central
fascia darkest ; basal patch edged and crossed by black lines ; pale band following
( 10.'5 )
with iiion; than its iiuifv half dark grey, the ontor half with a grey thread ; central
fascia narrow at inner margin, both edges irregularly Innnlate-deiitate, its centre
often blue-grey, containing a black cell-spot and two dark lines and a pale one
before outer edge ; pale band following very distinct, interrupted between 3 and 4
by a fulvous patch ; submarginal line white, preceded by blackish blotches on
costa, beyond cell, and above inner margin ; followed by a dark blotch at apex,
and by grey ones beyond cell and above inner margin ; marginal space between
3 and 4 generally paler; black marginal crescents between veins ; fringe whitish,
mottled with dark grey.
In the ? , as usual, the central fascia is broader, and the fulvous patcli beyond
outer line more extensive.
Ilindwing : dark grey, with the lines darker, the intervals on inner margin
white.
Underside pale, speckled with grey in hind wing, suffused with dark grey in
lower half of forewing ; all the lines indicated.
Head, thorax, and abdomen dark and light grey, the thorax mixed with
blackish; palpi externally fuscous; tarsi blackish, with the joints ochreous ;
segmental rings of abdomen white.
Expanse of wings : c?, 2G mm. ; ? , 28 mm.
5 66,4 ¥¥.
110. Xanthorhoe interrufata spec. nov.
Closely akin to A', ceras/iui, but the dark areas paler fuscous ; the central
fascia showing the two dark lines and the edges and cell-spot plainer ; the cerise
tints are confined to the pale bands edging the fascia, and even these are duller ;
the band before submarginal line is fuscous as well as the margin beyond.
In the hindwing the whole surface is fuscous, the cerise tints of the outer |]ale
band being restricted to the inner margin.
Underside fuscous, with all the pale cross-lines plainer.
Head, thorax, and abdomen fuscous; anal tuft white.
Expanse of wings : 44 mm.
A long series.
The wings are all glossy, as in crras/iia.
111. Xanthorhoe monastica spec. nov.
Forewiny : blackish fuscous, with all the lines darker; the central fascia with
three dark lines and edged with whitish, which at costa is white and conspicuous ;
fringe concolorous.
Ilindwing : wholly dark fuscous, paler beyond outer edge of fascia; jiale dots
along margin at the ends of the veins.
Underside dark glossy fuscous, the outer line marked by whitish dots on veins,
and the submarginal at costa only.
Head, thorax, and abdomen all blackish fuscous.
Expanse of wings : 40 mm.
A long series, not varying inter xc, except a pair apparently dwarfed, the 6 o(
which expands 3U mm. only and the ? 28 mm,
( lOG )
II"-. Xanthorhoe plumbiliuea spec. nov.
Foren-iiig : basal jiatcli and central fascia dark fiilvons fnscons, edged by broad
dull lenden-grc}- lines; the fascia narrow, with a leaden-grey line down its centre,
generally broken np into spots ; the broad bands on each side of fascia greyish
olive ; snbmarginal line fine, waved, leaden-blne, with the Innules filled up with
dark fnscons, sometimes only those beyond cell and on the snbmcdian fold ; a dark
obliqne streak from apex, the apex above it generally paler; marginal area and
fringe dark fnscons.
Uimluing : brownish fnscons, with a few indications of lines on inner mnr^in.
Underside greyish fnscons, paler beyond the onter edge of fascia.
Head, thorax, and abdomen dark fnscons.
Expanse of wings : 27 mm.
3 c?c?, 3 ? ?.
113. Xanthorhoe urbana Meyr.
(J. Forewing : grey, with a slight mixtnro of brownish, and speckled with
dark grey ; a small dark basal patch with ronnded edge ; the grey band beyond
it traversed by three darker lines ; central fascia occnpying middle third of wing,
the inner edge curved, the outer bluntly projecting in middle of wing; two lines
beyond inner edge and two before outer form bands filled np with darker grey ;
in these bands and in the basal patch some brown scales occur below median aiid
vein 4; below middle the bands imito and form annuli ; a paler grey band follows
with a grey line down it; snbmarginal lino obscure, preceded and followed by
darker tints, especially at apex ; a black marginal festoon interrupted at veins ;
fringe grey varied with paler.
Ilindwing : with base dark grey, followed by three blackish waved lines, plain
on inner margin, not reaching costa ; snbmarginal line and shades more distinct.
Underside pale grey, darker towards margin, with all the Hues blackish.
Head, thorax, and abdomen dark grey, like basal patch.
Expanse of wings : 24 mm.
5 6S.
Very near to A', subidaria Gnen. from Australia, for which I at first mistook
it. Two examples exactly like the present were received jireviously from tiie Aroa
River and referred to siibidana ; but with these 5 cJ <? there have now come fi ? ?
not distinguishable from the dark grey ? ? which Meyriek referred to subidaria as
a probable variety under the name of urbana ; in both sexes the wings arc greyer
and darker than in subidaria proper.
Subfamily ASTHENINAE.
114. Acolutha canicosta spec. nov.
Forncing : whitisii yellow ; the costal area above median vein sprinkled with
wiiite scales, the lines across it dark lirown; all the markings interrupted below
middle, reappearing much paler on inner margin ; these marks arc two brown
bands near base ; a central fascia jnst licyond middle, angled outwards on vein 0, and
a darker brown snbmarginal line angled on vein 7 towards apex, and reappearing
before anal angle as an upright brown bar ; fringe pale yellow.
Umdwi-ng : with four obscure curved brownish grey lines ; the antemedian
( 107 )
and median narrow, containing a lilaclc coll-spot lictweon thnni ; tlie postmcdian
and sulimarginal broader and distinctly double.
Underside pale yellow ; only the costal half of forewing dnll brown.
Head, thorax, and abdomen pale yellow ; face with a broad brown bar in middle
and a narrow one at top: vertex brown with a pale line in middle; shoulders brown.
E.xpanse of wings : 26 mm.
1 ?.
Near pictnria Moore.
115. Asthena argentipuncta spec. nov.
Forewinq : pale yellow, tinged with deeper yellow along costa and hindmargin ;
crossed by bands of round dull silvery spots ringed with brown-red ; in the ? these
spots are small and separate except along subcostal vein ; in the S they are large,
with the brown rings thicker, and therefore more or less confluent, especially in basal
half; there are four antemedian series, including one at base itself ; one postmediau
and one submarginal ; in the postmediau the two spots on vein 3 and 4 are displaced
outwards ; a larger discal spot, with costal spot above it ; a marginal line of deep
brown dashes between veins ; fringe yellow.
Hinchoing : with the two outer series only ; the cell-spot and two joined spots
below it.
Underside of forewing irregularly brown along costa ; only the outer and
submarginal lines represented as brown spots, not reaching inner margin ; of
hindwing with the outer lines marked only on costa; both wings with fine
marginal line ; in the ? the lines are not shown, and the brown along costa
is much reduced.
Face deep yellow ; vertex white ; thorax and alidomen yellow, the dorsal
segments deep yellow marked with silvery spots ; anal segment and underneatii
with legs pale yellow.
Expanse of wings ; S 24 mm., ? 26 mm.
3 c?c?, 4 ??.
The double arcole is very narrow. Antennae of c? with fine long fascicles
of cilia.
116. Asthena subditaria sjiec. nov.
This is almost a reproduction of the preceding species, but smaller in size and
paler in colour ; the spots, in proiiorti(ui, larger and more coalescent in both sexes ;
the outer line more sinnous ; the marginal line of dashes absent.
Expanse of wings : 22 — 24 mm.
1 c?,2 ??.
117. Hastina viridata spec nov.
Forewing : greenish white, the lines dark green, all wavy, and in the main
parallel to hindmargin ; two near base, two in middle, and two towards hindmargin
form darker bands, separated by pale sjjaces, each traversed by paler green lines ;
all alike are darker on the veins, followed by paler dashes ; the median band
generally shows two deeper areas, one beyond cell, the other above inner margin ;
a dark green marginal line preceded by pale spaces between the veins ; fringe
whitish, chequered with green at the vein-ends,
( 108 )
Ilimhving : without tlic basul lines.
Uiulersicle jiale greenish, witli tlie darker tints of npperside sliowing throni^h.
Head, thorax, and abdomen green, mottled with whitish ; fiice with lower half
whitish, npper half olive-brown ; tips of paljii brown.
E.xpanse of wings : 22 mm.
0 <?c?, 2 ? ?.
118. Hydrelia papuensis spec. nov. and ab. dorsinotata nov.
Forewing : pale greyish ochreons, tinged with darker, and crossed by a
succession of slightl}- darker luiuilate-dentate lines, the teeth all pointing inwards
and forming slight brown points on veins, tlie liiniiles of the outermost series also
marked at their aiiices with a brown jmiat between the veins; a row of large brown
marginal spots ; the line marking the outer edge of central fascia is more strongly
exjiressed and followed closely by two grey lines, forming together a narrow band ;
between veins 3 and 4 the outer of ithese three lines and the two praesnbmarginal
lines are marked, sometimes strongly, with Ijrown-black sealing ; cell-spot small,
black ; fringe pale oclireons.
Hindwing: similar, without the basal lines, and with no dark scaling between
3 and 4.
Underside paler, tlie lines much fewer, but darker, and only the outer ones
distinct ; cell-spots black, also the marginal spots, those of forewing being much
swollen.
Head, thorax, and abdomen dusky grey ; dorsal segments with dark middle
spots, second segment with a pair, one on each side.
Expanse of wings : 26 mm.
4(?c?.
The species is nearest to // atrostrignta Warr. from Qneensland.
It shonld be noted that though 1 have jilaced this sjiecies in ]li/<hrU(t, as
having apparently a single areole, the areole is actually double, vein id running
into 11 close to its origin, so forming one very small triangular inner and a large
outer areole. I am inclined to think that this small inner areole, situated as it is
far bascwards, is often overlooked.
In the aberration a large lilack-brown sinuous lilotch rises from middle nf inner
margin of forewing, where it is liroad, and is bent (lutw^ards at lower end of cell
between veins 2 and 4.
no. Poecilasthena paucilinea spec. nov.
Dill'ers from /'. thalassias Meyr. in two respects ; tlie hindwings ot //i<i/(issias
arc at most bluntly elbowed at vein 4 ; in paucilinea the hindmargin is distinctly
angled, vein 4 forming a sliort tooth ; secondlv, in thahiKsius the lines forming the
central fascia are four in number ; in the present species they are but three ; the
pale band on each side of the central fascia is traversed along its centre by a green
waved line in thalassias ; in paucilinea the bands themselves are broader, the
traversing line keeps close to the edge farthest from the fascia, which is therefore
limited by a broader space of pale ground-colour. In all otlicr jioints the species
appear to agree.
2 cJcJ, sent along with a series of the true thalassias Meyr.
( 109 )
Subfamily TRICHOPTERYGINAE.
120. Anthierax aroensis.
Auisociditiauriiensis'Wsi.VT., Nov. Zuol. x. p. 382 $ (I'JUS).
S. Forewinij : wbitisb, with grey suffusion and fine Ijliickisli lines, the lines
jilacecl as in the ¥ , but the whole wing much paler ; tlio band before snljmarginai
line as far as vein 4, and the edges of central fascia towards costa alone showing
fuscous with a slight greenisli tinge ; the cleft in hiudmargin between veins 2 and 3
rnns in for one-third of wing, the part below it being produced into an acutely
jiointed lobe, reaching well beyond the upper portion of hiudmargin ; the distortion
below at middle of inner margin at the extremity of vein 1 is strongly marked, and
the lobe clothed with white liairs.
llindwiny : white, with a large basal lobe, a fulvous grey twisted middle lobe,
and a dark grey anal projection.
Hindlegs shortened and swollen, with tufts of ochreons hair ; the abdomen
also roughened with hair below. Antennae fulvous, darker at base ; paljji pale grey
speckled with darker, second segment rough-haired, third spatnlate.
Exjjanse of wings : 35 mm.
1 S.
121. Episteira delicata spec, no v.
Forewinij : wliite ; Itasal area witli six irregularly waved thick greyish olive
lines, mainly vertical, the costal vein with three fine black dashes ; some black scales
along inner margin at base of all the lines ; cell-spot linear, black, in a narrow
pure white central space ; four postmedian olive-grey dentate lines, of which the
first two are marked with black and the third is thick and ditfase ; a partially double
blackish dentate submarginal line ; a marginal row of black swollen spots between
the veins, and a row of large black spots Ijeyond them in the white fringe.
liindwinij : very pale grey, slightly darker before hindmargin.
Underside olive-grey, paler in hindwing and along inner margin of forewing ;
fringe white.
Palpi and antennae blackish ; face dark brown ; vertex olive-grey ; fillet, collar,
and tijis of shoulders whitish grey ; thorax grey, with a black lateral line, thickened
behind ; abdomen olive-ochreous.
Expanse of wings : 20 mm.
1 S.
Meg'aloba gen. nov.
ForiH-iidi : large ; costa straight for four-fifths, then strongly convex ; apex
minutely j)roduced ; hindmargin gibbous or elbowed in middle ; anal angle distinct.
Hind winy : narrow, elongate ; the apex truncate; hindmargin crenulate; in the
$ with a large ear-shaped lobe reaching half-way down inner margin, the lobe itself
edged with a scale-bearing flap, which at the base forms a small additional lobe.
Abdomen in cJ long and slender, with a stout keel beneath at base, in ¥ stout ;
antennae simple, lamellate ; jialpi porrect, long, second segment long-haired, third
short and smooth ; tongne and frenulum present; legs long and slen<ler ; hindtibiae
in ¥ with terminal sjjurs ; in i thick, the femnr, tibia, and tarsus of equal length,
without spurs.
Neuralion : forewing, cell ipiite half as long as wing ; discocellular obliipie ;
first median nervnlejust bejond middle, second at six-sevenths; lower radial from
( lio )
above aiiddle of discouollultir, npper from npper end of cell in ? , stalked with 7, S, 'J
iu cJ ; lU, 11 coiucideut, 10 anastomosing with 8, U ; liiiidwiug, costal and sulicostal
anastomosing to near end of cell, C, 7 long-stalked iu c?, shurt-stalked in ? ;
discocellnlar biangnlate, the radial from tlie lower outward angulation, vein '^ absent
iu c?, 3 and 4 short; veins 2, 3, 4 at equal distances iu ?.
Tyjie Megaloba rubripicta sjjec. nov.
The large subercct lobe iu the S liindwiug is something like that iu Ti/injiniio(a
Warr., but in that genus the costal and subcostal are sei)arate but uuitod by a bar.
122. Megaloba leucocyma spec nov.
Forewing : sage-green, with darker and paler waved bands, all edged more
or less distinctly with bluish-white lines, which on the dark green costa form
snow-white spots ; the bands on each side of central fascia paler than the rest,
having an interrupted white Hue aloug their centre ; submarginal line forming
white crescents ; a line of large black crescents aloug margin at the ends of the
veius, edged with white scales ; fringe pale green.
Hindwing : pale brownish grey, paler towards base, with dark linear cell-mark,
curved postmedian Hue with paler edge and dark spots in fringe beyond the veins.
Underside of forewiug greyish olive, with the lines faint; the subnuirginal
shown by a curved row of white spots : hindwing without the olive tinge ; cell-spot
and postmedian line dark.
Palpi pale olive, the tips of each segment whitish ; face green, with the
lower part white ; a snow-white dot in front at the base of each antenna, and
one in the middle of the crown ; one behind each eye and one at the base of each
of the patagia ; a white curved line on thorax and metathorax, and a white mark
on basal segment of abdomen ; abdomen fawn-grey, the basal and anal segments
olive-green ; antennae greenish above, rufous below; pectus and femora olive-green;
tibiae and tarsi blackish, with suow-white joints, the hindtibiae grey.
Exjianse of wiugs : 44 mm.
1 ?.
123. Megaloba? rhododactyla spec. nov.
Forewing : olive-green, crossed by waved and crenulate lines of shell-pink ;
along the costa the edges of the green bauds become blackish green ; basal patch
olive, with three pinkish-white lines, with some black scaling above and below
the submedian vein ; central fascia with four pinkish-white lines, the dark
cell-spot lying in the first ; the inner band with a dark blotch ou the submedian
fold, the outer edge irregularly crenulate and marked by black spots on the veins ;
both pale bands shell-piuk, the outer more vividly, with an olive-grey central Hue ;
submarginal line lustrous white, rosy tinged, j)receded by a black line swollen
into teeth between the veius and externally tipped with black ; a margituil row of
black truncated crescents at end of veius alternating with spaces of shell-pink ;
fringe pale olive, with white spots beyond the veius.
IJiudtcimj : pearly cream-colour towards base, grailually becoming jiale green
towards hiudmargin ; fringe rufous beyond a dark marginal line.
Underside of both wiugs olive, the markings showing through.
Palpi olive-green, with the tips of each segmeut white ; face green, with the
sides below white ; base of antennae, a spot before each, ami a sjjot before each
eye white ; collar, thorax, and abdomen olive mi.xed with shell-piuk and while ;
( ill )
antennae deep red beuoaHi and at sides, aliovc unnulated with red-brown and
white ; pectus white ; femora and underside of abdomen olive ; forctarsi l)hi,ck-, with
white joints ; foretibiae bright red.
Expanse of wings : 30 mm.
1 ?.
The position of this very beantifnl insect is doubtful ; it agrees with Meqaloha
in ueuratiou, and in the slightly produced, pointed, not rounded, apex of forewing.
1~M. Megaloba rubripicta spec, no v.
c?. Forewing : dark green, with blackish green lines and shades ; a dark shade
close to base, followed by a paler grey-green baud ; then a broad outwardly obliipie
band, its edges dark, outcurved externally above submedian vein ; next a thin
band of pale bright green, mixed with bluish-white scales ; central fascia with
inner edge dark, above submedian blackish and projecting towards inner band so
as nearly to touch ; outer edge irregularly dentate, angled below vein 4, then
concave inwards, preceded by two dark green lines, the inner edge followed by one,
all interspersed with bluish-white scales ; cell-spot dark green ; the foscia is
followed by a narrow band of bluish-green scales, straight from costa to below
veiu 4, then incurved and lunnlate ; submarginal line pale bluish green, edged
outwardly with blackish and preceded by a dark green shade, which forms a
blackish blotch beyond cell ; small blackish angled spots at ends of veins ;
fringe green.
IliinhLiiiij : basal half and lobe whitish, scakdess ; outer half reddish fuscous,
with the veins and fringe bright red ; flap of the lobe with olive-green scales.
Underside of forewing bronzy olive-green, with the veins and hindraargin
rust-red ; costa from near base broadly bright ferruginous ; fringe pale yellow-green ;
hindwiug like upperside, but paler ; the veins and fringe red.
Head and thorax green like forewing ; abdomen yellow-green ; terminal
segment of palpi brownish with the tip white ; legs pale olive-green or ochreous ;
foretarsi black-brown with the joints ochreous ; antennae anuulated with green.
? much gayer; the dark edgings of the inner band and central fascia of
forewing, and the band preceding submarginal line broadly deep dark green ; the
intervals bright jjale green with ochreous edging ; the submarginal line white ;
cell-spot large, crescentic ; fringe red ; hindwing bright red, more ochreous towards
base, with two postmediau waved grey lines and the margin dark.
Underside bright ferruginous, mixed in forewing with greenish, the marginal
area green with the fringe ferruginous ; forewing with six dark shades, hindwiug
with two postmedian lines and marginal border.
Head, pal])i, shoulders, and patagia olive-greeu ; metathorax and base of
abdomen red-brown ; tips of patagia red ; metathoracic tuft bright i)ale green ;
antennae anuulated with red ; antennae beneath and legs bright ferruginous.
Expanse of wings : $ 35 mm. j ¥ 40 mm.
The difference between the sexes is remarkable.
1 (J,2 ¥ ?.
125. Remodes parviplaga spec. nov.
Forewing : greenish white, with a succession of waved and dentate greeu lines
crossing from base to margin, exactly as in li. lobata Warr. from Padaug, anil
R. pallulipluga Warr. from Java ; the only dark markings are the spots on margin
( 112 )
at the ehdx of the veins, and two or three dashes above the snbincdian vuiii in
central fttscia, as in lobata ; the pale band beyond tlic fascia ends in a bright
whitish fleck on margin above the end of the subraedian, smaller and whiter tlian
tlie blotch in pallidiplaqa.
Ilimlicuig: whitish grey, witli the outer lunrgin greyer; the terminal lobe
and the ni)i)er folded edge of the middle lobe still darker grey ; the lobe at base
ijnite small ; this agrees with palli(Upla<ia, but not with lobata.
Underside olive-green, becoming reddish grey towards hindmargin ; the IViiigc
of the two outer lobes of hindwing blackish.
Abdomen with lateral spreading tuft of hairs ou second segment and ui}curved
tuft from sides of penultimate segment; hindlegs twisted, with dull ochreous tuft
of hairs on femora; antennae olive-green externally, blackish internally.
Expanse of wings : 35 mm.
1 c?.
I have seen a pair before from the Ujipcr Aroa River, which 1 at the lime
referred to lobata.
126. Sauris atrilineata spec. nov.
Fon'wing : olive-green, crossed by darker augled waved lines, all thickly
marked with black except in the interspace between veins 3 and 4 and the costal
area above vein G, where the lines are finer and greenish black ; the line close
to base and the central threads of the two ])a]e bauds are also dark green; all
the lines are angled outwards in cell and beyond, incurved below middle, and again
inclined obliquely outwards towards inner margin ; beyond the green basal area are
two pairs of black lines ; tl)e bands of central fascia consist of three and four lines
each, and there are two dark lines before the snbmargiiial ; marginal spots at ends
of veins square and black, followed by square white spots in the Islackish fringe.
Hindiciny : grey, paler at base ; the small lobe with a black dash on its
upper edge near base.
Underside dark smoky-grey with the markings slmwing through ; cell-spot
and marginal sjiot of forewiug blackish.
Head, thorax, and abdomen olive-green, the last with black dor&al marks;
patagia black ; tibiae and tarsi black with pale joints ; hindlegs paler.
Expanse of wings : 2<J mm.
1 c?.
Neuration of hindwing as in (jriseolauta.
1~T. Sauris griseolaiita spec. nov.
Forcwiiiy : pale green; the transverse lines and shades dark olive-green, in
parts olive-brown, edged and alternating with pearl-grey lines, which are paler
in (? than in 9 ; a dark green i>atch at base, followed by a pale band ; a broad
dark green band of three ditfnse lines, the intervals grey, followed by the usual
pale band with darker centre before the central fascia, its enter edge pearl-grey ;
central fascia formed of two bands of three lines each, dark grceu and brown,
alternating with ])earl-grey, followed by tlie usual ])ale band ; a double dark green
band before the pearl-grey submargiual line, which is followed by a single dark
green shade ; marginal spots at end of veins dark green, siibi|uadrate, alternating
with jiearl-grey spots which project into the pale green fringe.
lliiKliriiKj : jialc grey, whiter towards base.
( 113 )
Underside greenish cinereous.
Head and thorax green ; palpi paler green with whitish tips ; abdomen
greenish ochreons ; antennae oclircous, more rufous beneath ; a white spot behind
each e)'e.
Exjjanse of wings: c?, 30 mm.; ?, 34 mm. One dwarf paler c? measures
only 26 mm.
3 c?c?, 1 ?.
Th ? is always darker than the S-
6. Hindwing : cell barely one-fourth of wing, broad; discocellnlar angled;
costal and snlicostal ajipniximating well beyond cell ; 6, 7 stalked, 7 anastomosing
and becoming coincident with 8; 5 and 4 close together from end of cell; 1,
2, 3 absent.
128. Steirophora violacea spec. nov.
Forewing : pale olive-green, crossed by darker green waved and angulate lines,
all more or less marked with blackish, except the dark green thread of the two pale
bands ; the dis])Osition of the lines is almost identical with that in <S. auratisquama
Warr. from Java; but the gilded yellow scales of that insect are less conspicuous ;
instead the present species is characterised by a violet shade, which suffnses the
submedian interspace and the outer pale band, and sometimes encroaches on the
outer half of central fascia, but not above vein 6.
lliiulwiug : grey with a distinct violet tinge, the outer half darker with a paler
curved postmedian band ; a small grey cell-spot.
Underside cinereous olive, with the markings showing through.
Head and thorax green ; patagia grey with black scales ; abdomen greenish
ochreons, sometimes with darker patches along d(irsum ; metathorax and two
basal segments of abdomen with a broken black lateral line ; palpi greeu dusted
with darker.
Expanse of wings : S 30 — 35 mm. ; ? 35 mm.
5 SS, 1 ?.
S. punctatissima Warr. from S. Celebes is also like this species, but has
narrower, more pointed forewings.
Subfamily TEPHROCLYSTIINAE.
129. Adeta confusa spec. uov.
ForctciiKj : dark olive-fuscous ; the lines pale, greenish, very faint ; basal patch
liuite small, the usual space beyond it broad, of the same dark grey tint, edged by a
pale line, which forms a strong angle above median and a fainter one below it,
followed by the broad blackish inner baud of central fascia; the white centre of the
fascia makes the thiee preceding areas appear to form one large basal patch ; outer
edge of central fascia at two-thirds, crenulate, oblicjue outwards to below 4, then
inwards, preceded by three dark lines, forming its outer band, and followed by a pale
greenish band with darker centre ; submarginal line pale greenish grey, preceded and
follcjwed l)y a dark band ; the outer one with a pale spot between 3 and 4 ; a dark
marginal line ; fringe mottled dark and light.
HindiciiKj : similar, but the paler green intervals broader.
Underside dark greenish cinereous, paler in hindwing, the dark shades showing
through.
8
( ni )
llciul, tliorax, iiiiil iilKlomcn dark and jiule grceii ; vortrx [laler.
ExiJaiksc of wings : 2"J mm.
1 ?.
Distinguished liy the dark ground-iulour and wiiite central streali.
130. Auiserpetes fasciata spec. nov.
Foreiciiu/ : dull green ; the central fascia tlnll violet ; the whole wing with
minute black dusliug; first line curved, black, at one-fonrth; outer line at two-
thirds, creniilate, outcnrved above, folhnved by a white line ; submargiual line faint,
waved, preceded by a violet-grey band, which is darker at costa, beyond cell,
and above inner margin ; marginal line black; fringe greenish grey.
linulwing : the same ; marginal area with a large scjuare white siwt between
3 and 4, which is merely a pale patch in forewing.
Underside pale greenish cinereous with dark bauds.
Head, thorax, and abdomen green and violet.
Expanse of wings : 20 mm.
1 J.
131. Ardonis deutifera spec. nov.
Forewing : bright pale green, crossed by wavy green lines ; basal patch marked
liy a black costal spot, with a tuft of raised black scales on it, and a dark point
on snbmedian vein ; inner edge of central fascia marked by a black tooth-shaped
blotch, its ajiex bearing a large tuft of raised black scales above the black cell-sjiot ;
outer edge formed by a small triangular black blotch, the lines edging the fascia
marked across wing by slight black dots on veins ; submargiual line white, wavy, the
lunules slightly filled with dark scales ; fringe mottled pale and dark green, beyond
slight dark marginal spots.
Hindwing : green, paler towards costa, with the lines alternately green and
whitish green, the edges of the central fascia shown by black dots ; a double black
spot at anal angle, and the marginal lunules black; costa of forewing of c? shining
white with an oval patch of black scales beyond middle.
Underside green, with the black markings distinct and neat; S with inner
margin of forewing glossy white, with au oval patch of rongh black scales.
Head, thorax, and abdomen green, the last with a blackish band on second
segment ; shoulders black-spotted laterally.
Expanse of wings : 23 mm.
1 cJ,2 ? ?.
132. Chloroclystis cuneilinea spec. nov.
Forewing: greenish wliitc, tlie intervals all reddish on costa; crossed by a
succession of pale grey-green lines ; the bands limiting the central fascia rather
paler green ; central fascia slightly greyer green, the lines towards its outer edge
reddish ; the outer edge itself marked by black wedge-shai)ed spots on veins ; inner
edge also with black points on median and submedian ; submargiual line pale green,
waved, preceded by a grey-green baud of two lines, tinged in j)arts with red ; the
margin grcj'-grecn ; marginal line black, interrupted by the veins ; fringe glossy
jiale green.
Hindwing : paler ; the fascia without reddish scaling ; the spots and all the
lines very delicately marked.
( 115 )
Underside of forewing greenish grey, except tlic inner margin, wliiuli, like the
liiiulwing, is jiaJe green, with thick outer and black marginal line.
Head, (horax, and abdomen jialc green ; abdomen with a [link spot on sides of
third segment ; forelegs dark green with pale joints.
Expanse of wings : 10 mm.
1 (?, I ?.
133. Chloroclystis semiscripta spec, no v.
Forctriiii/ : whitish green; the markings grey ; band edging basal patch at one-
fourth, curved; inner edge of central fascia at two-fifths jjarallel to basal line,
marked with some brown below middle : both these reach the inner margin ; outer
edge at two-thirds, dark at costa, then forming pairs of grey dots on veins to
4 ; praesubmarginal band dark grey to 4 ; submarginal line of the i)ale ground
colour, followed by grey diamonds on veins ; fringe grey ; all the pale intervals
crossed by pale grey lines dotted on veins ; below vein 4 the outer markings are
pale and blurred.
Hindtcinf/ : whitish green, with a faint grey cloud at anal angle.
Underside whitish green, with the markings only showing through.
Head, thorax, and abdomen whitish green ; dorsum brownish-tinged at base, and
with a black dot at side of second segment; forelegs blackish, with pale joints.
Expanse of wings : 23 mm.
1 ¥.
134. Eucymatoge albicristata spec. nov.
S. Fomwimj : puriilish brown, the two pale bands and centre of fascia whitish
at costa ; basal patch with a dark vertical edge ; inner edge of central fiiscia
crenulate and nearly vertical, outer edge crenulate, inangled on vein 7, insinuate
beyond cell, projecting on vein 4, and followed by a white spot on veins ; the
outer baud of fascia dark throughout, the inner band dark at costa only, both
containing three creuulated lines; cell-spot linear, black; marginal area purple-
brown ; all the lines marked on veins by dark and light dashes ; marginal line
black ; fringe reddish, mottled with brown ; the whole wing below median darker
than above.
Hindidng: grey, darker along hindmargin, with traces of grey postmediau
and submarginal lines ; pale dots at the vein-ends ; fringe dark grey.
The ? is slightly paler, and has the middle of central fascia above median
vein white.
Underside dark grey, jialer towards base, with black cell-spots and dark
outer line,
Head, thorax, and abdomen like wings, purplish brown with paler speckling;
metathoracic tuft wliite.
Eximnse of wings : i 26 mm. ; ¥ 3U mm.
1 <?, 1 ¥.
135. Eucymatog'e albimedia spec. nov. and ab. brunneotincta nov.
Forewing : chalk-white, the basal and marginal areas dark pnr])le ; basal patch
edged by a thick velvety black shade, outcurved from costa to submedian fold,
vertical on inner margin ; inner band of central fascia curved parallel to edge of
basal jiatch, starting from a pnridish triangle on costa and ending in a smaller
one on inner margin, always interrupted between veins 1 and 2, the costal i>art
( 116 )
joiiicil horizuiitnlly abovo median vein witli the elongated pnrplisb discal spot ;
outer band uiiiiiterniptcd, of uniform width, deep jnirplc-black, finely edj;ed
externally by a luuulate whitish-yellowish Hue ; the white band beyond basal
patch has two grey middle lines, only visible above median vein, where they are
bent, and is interrupted between snbmedian fold and vein by a brown clond, which
in a manner unites the basal ])atch to the lower jiart of central fascia; the white
area of the fascia is traversed by three lines, marked mainly by dark spots on veins,
the outermost immediately preceding the outer band ; the usual pale band beyond
fascia is filled up with purplish above middle and with brown below; snbmarginal
line indicated only in lower half of wing, followed by a pale grey patch at apex,
between veins 3 and 4, and below vein 2 ; marginal white dots at end of veins ;
fringe purple-black, paler beyond the three pale spaces, with two dark lines and
dark spots beyond veins.
Ilindwing : purplish black, paler in basal half; small white dots at end of
veins, and sometimes traces of a postmedi:in line.
Underside dark purplish, dusted with bluish white ; inner margin of both
wings whitish ; cell-spots dark : basal area paler ; outer area with the veins marked
dark and light ; fringe as above.
Face, palpi, and inside edge of collar, shoidders, and patagia snow-white ;
base of each segment of palpi purple ; thorax, shoulders, patagia, and antennae
purple ; abdomen purplish mixed with whitish ; dorsal baud and crests purple ;
legs purple with the joints white.
Expanse of wings : 3U — 32 mm.
5 S6.
In the aberration hmnncothicta only the interrupted band before fascia is
white ; the white of the fascia itself, except at the costal edge, being suffused
with brown, and the outer baud of the fascia is narrowed.
1 cJ.
13C. Eucymatog'e brunneata spec, no v.
Foreiriiuj : sufTused with pale brownish flesh-colour; the markings brownish
fuscous and black ; basal patch darkest, the edge curved outwards to submedian
fold, along which it runs shortly inwards, then vertical to inner margin ; inner
edge of central fascia 2)firallel to it, angled outwards in cell before cell-spot and
inwards on submedian vein ; outer edge acutely inaugled on vein 7, forming a
double lobe outwards on vein 4, then incurved, wavy, dentate ; the fascia crossed
by five minutely crenulate blackish lines and containing a vertical black linear
cell-spot ; the pale band beyond narrow, flesh-coloured, with white teeth inwardly
on veins, and wholly while at costa ; marginal area beyond it filled up witli dark
fuscous above vein 4, and brownish flesh-colour, with the lines dark, beneath ; paler
altogether between veins 3 and 4 ; the submargiual line pale and waved ; a dark
marginal line interrupted at the veins ; fringe brownish with dark chequering.
llindicimj : grey, witli wavy outer and snbmarginal lines, the marginal border
darker ; fringe grey, spotted with dark beyond veins.
Underside dark grey, with the cell-spots black and the outer lines indicated.
Vertex, paljji, thorax, and abdomen blackish fuscous ; face, fillet, and collar
whitish.
Expanse of wings : 26 mm.
1 i.
( iiv )
137. Eucymatoge constellata spec. nov.
ForeiriiHf : dark brown, dusted with pale ochreoiis ; lines shown by series
of round pale ochreous spots ; three larger costal spots at one-fifth, two-fifths,
and two-thirds, from which the three series start ; a black cell-spot in an ochreous
patch ; a submaruiual line of spots, starting from three-fourths of costa and bent
on vein 0, followed on costa by an intense black shade, which reappears above
anal angle ; miante orange dots at the vein-euds ; fringe brown, dark-chequered
beyond veins, with the tips pale.
Uimiivlmj : dull creiimy-white, with the dark postmediau and submarginal
lines of underside showing through ; fringe with a rufous tinge, chequered with
dark brown.
Underside of forewing paler, blurred ; the spots yellowish ochreous ; hindwing
yellowish ochreous with brownish speckling ; a black cell-spot ; a brown cloudy
postmediau shade starting from a dark brown costal blotch, and a macular
submarginal line.
Head and thorax brown, with pairs of yellow spots ; abdomen ochreous with
brown dorsal marks ; anal segment above and praeanal segment below black-browu ;
legs ochreous, with black tarsi.
E.xpanse of wings : 18 mm.
A long series of c?cJ, without a single ? .
138. Eucymatoge contaminata spec. nov.
Foreicinq : dirty white; the markings in the main the same as in E.rujilunata,
described below, but all tlie cross-lines, which are pale grey in that species, are
here pale rufous, the whole wing having a tinge of rufous over it ; the edge of
the basal area is evenly curved, not projecting in the middle, and the tooth of enter
edge of central fascia between veins 3 and 4 is much sharper and longer ; instead
of the chestnut streak of rufihimifa this species has a black or blackish brown mark,
followed by a diffuse dark cloud ; cell-spot generally broken up into two dots.
Ilindwiny : dirty white, with an ochreous tinge.
Underside dirty ochreous grey, without any distinct markings.
Head, thorax, and abdomen as in rnflunata, but the white scales all discoloured.
Expanse of wings : 27 mm.
4 S$.
The difference in the undersides will serve to distinguish the two species.
139. Eucymatoge discinota spec. nov.
Forewing : deep velvety brown ; the veins towards margin reddish, running
out into the reddish fringe ; some bluish-white scales at base, along subcostal vein,
and inner margin ; a white spot at base of wing ; lines marked by snow-white
spots ; five spots in a curve close to base, on the folds as well as on the veins ;
three larger spots in an oblique line to middle of inner margin, lying between
the veins, the middle one largest ; discal spot large and bent, finely edged with
pale, with a round white spot on costa above it ; a costal spot with one or two
smaller ones below it, and a large double spot below median followed by a dot
on inner margin represent outer line ; a row of submarginal and marginal white
spots between the veins ; three costal spots before apex ; a pale blurred wedge-
shaped space reaches from base through cell to hiudmargin.
( 118 )
Ilinthritifi : dull whitisli, with jjroy postmodian linn ninl martjiiiiil linrcler, the
space between paler.
Underside of forewing dnll brown, speckled with whitish ; the two large costal
spots and the corresponding pair above inner margin white ; also the scries before
marginal line: hindwiiig thickly speckled with brick-red; postmodian line and
veins red ; marginal border dark brown.
Head, thorax, and abdomen deep brown with white dusting; tln.irax with four
large white spots.
Expanse of wings : S 21 mm.; ¥ 24 mm.
4 c?(J,4 ??.
140. Eucymatoge disrupta si)ec. uov.
(?. Foraciiu/ : greenish white; the markings lilack, shaded with vi(det and
very dark green ; basal patch of the gronud-colonr, edged by a black shade whicli
is angled in middle of wing, interrnpted below, and ending in a broad blotch on
inner margin ; inr.er edge of central fascia interrnpted, parallel to edge of basal
patch, marked by blotches on costa and inner margin and spots on veins between ;
outer edge at three-fourths, prominently angled at veins 6 and 4, interrupted
between 2 and 4 and marked on 3 by a black spot ; cell-spot black, large and
vertical ; two lines of black vein-spots across fascia ; traces of three dark lines
along inner pale band at costa; some faint clouds along costa, before the cell-spot,
and in the interrupted part of onter band; submarginal line preceded by a violet
and olive band, edged with blackish, especially on veins, and interrnpted between
veins 3 and 4 and partially above 6 ; some violet clonds along margin above vein 4
and below vein 3; pairs of black dots at the vein-ends, followed by olive cheriuering
in the pale fringe.
Iliiulicing : pale grey, darker towards hindmargin, with an angled postmedian
and curved submarginal line marked darker on veins ; fringe whitish, mottled
with dark.
Underside dark grey, with cell-spots dark and fringes white with dark mottling.
Head, thorax, and abdomen greenish white ; centre of shonlders, patagia, and
thorax blackish green ; intermediate segments of dorsum dark green with whitish-
green sagittate marks, the basal and anal segments wholly pale green ; segments
of palpi with their bases green.
? whitish, without the green tint ; all the dark mnrkings and shadings more
developed and intense, (he pale bands before and Ijcyond the central fascia filled
in with dark scales.
Expanse of wings : c? 24—20 mm. ; ? 3ii mm.
3 (Jc?,l ?.
141. Eucymatoge distorta spec. nov.
Foretving : dnll brown, finely pale-sj>eckled ; three large white costal sjiots
at one-fifth, two-fifths, and two-thirds, three corresponding small white spots on
inner m.argin ; a white dot at base of wing ; a white spot near base of cell ; a large
white spot in submedian interspace below base of vein 1, and a donblc spot beyond
on each side of vein 1 ; cell-spot linear, dark ; a blurred pale brown wedge-shaped
mark from base of cell to hindmargin ; snbmarginal line showing as white dots
only above and below this space; a very fine dark marginal line; fringe rufons,
with white tijjs, and chei^uered with brown beyond veins,
( 119 )
lliiidiriiiii : white, with l)roatl bmwnisli maivinal border ; friiiije paler l)rown. ■
Underside of ibrcwiiig bhirred, much paler ; all the white spots shown :
hindwing redder brown, thieklj- speckled with pale; the marginal border darker;
the central space generally paler ; a dark cell-spot ; the veins often pale.
Head, thorax, and abdomen brown witli pale dusting; dorsum marked with
white spots.
Ex[>anse of wings : 24 mm.
3 $$.
Closely allied to, but ipute distinct from, E. discinota. The hiudmargin of
forewing is clearly indented just below vein 4.
142. Eucymatoge griseata spec. nov.
Foir/riiiff : ashy grey ; the lines darker grey ; basal patch dark grey, crossed
by two darker lines and edged by another, which is only slightly curved; inner
edge of central fascia curved parallel to it, not waved or projecting into the fascia
as in most of the allied species, blackish throughout; outer edge at two-thirds,
forming two sharp teeth between veins 2 and 4, and blackish beyond cell ;
snbmargiaal line pale grey, preceded by a darker grey band ; dark grey marginal
spots at ends of veins ; fringe grey ; all the pale spaces traversed by waved darker
lines ; cell-spot blackish.
Ilimlwimj : greyish white, grey along hindmargin, with traces of two waved
lines and two more before them on inner margin only.
Underside dull grey, the cell-spots black, the outer line indistinct.
Head, thorax, and abdomen pale grey, thickly speckled with dark grey ; palpi
and face blackish, the extreme tip of palpi whitish.
Expanse of wings : 30 mm.
1 V.
143. Eucymatoge licheuosa spec. nov.
Forewing : pale lichen-green, the markings moss-green, mixed in places with
violet ; basal area pale green, crossed by two waved dark lines, marked with
violet at costa and edged by a violet band, which projects shortly below subcostal
and submedian veins and is interrupted above the submediau ; inner band of
central fascia, outcnrved in middle, darker on costa and inner margin, where it is
marked with violet and black scales, interrupted below middle, followed by three
moss-green lines, the outermost of which is sharjjly dentate outwards l)eyond
the linear black cell-spot ; outer band of three lines, partially filled in with
uioss-green, the outermost marked with blackish and strongly wavy dentate, wilh
a prominent sharp tooth between veins 3 and 4 ; the pale band before fascia with
a double line of moss-green scales, that beyond it with a single line, its outer
half beyond cell occupied by a broad red-brown patch ; submarginal line pale
green, preceded by a deep violet and green band, which is strongest at costa
and beyond cell, and on the veins is marked by a strong pale green dash ; veins
towards margin violet ; pairs of dark spots at ends of veins ; fringe greenish
white, mottled with olive-brown beyond veins ; all the lines form blackish green
spots on the costa.
Ilindtrliu/: whitish in basal half, with a dark grey curved border, in which
the veins are marked with alternate dark and pale dashes, indicating three lines,
( 120 )
preceded by an angiilatcd jiostraedian line ; fringe whitisli, with dark dashes
beyond veins.
Underside dark grey, with the outer angulated lines and eell-siiots clear, as in
E. riifiliinutu.
Head, thorax, and basal segment of abdomen pale green like wings, sprinkled
■with moss-green and violet scales, especially the thorax ; second segment of
palpi externally, and base of terminal segment dark green ; abdomen a luixtnre
of pale green and violet-fnscons scales, the second segment brownish, the dorsal
crests large, olive-green ; fore and middle tibiae and tarsi black, with the joints
white ; pectus white.
Expanse of wings : 26 mm.
1 3.
144. Eucymatoge nigribasis spec. nov. and ab. mediotaeniata nov.
Foreiving : greenish grey ; basal patch lilack-green, finely white-edged ;
three black linear costal marks ; the first indicating the origin of the dark
centre of the jmlo band that precedes the central fascia ; the second the central
fascia itself, of which only faint traces are visible ; the third is contioned across
wing as a dark shade filling up the Innules of the waved white submarginal line ;
fringe grey with pale marks.
Hindwing : paler grey, darkening towards hindmargin, with a dark postmedian
line.
Underside of forewing dull greenish grey, with the markings darker and pale-
edged, visible only towards costa : hindwing paler at base, more speckled ; a dark
line beyond middle with paler baud beyond it.
Head, thorax, and basal segment of abdomen black-green ; rest of abdomen
like wings ; pectus and forelegs blackish.
Expanse of wings : 17 mm.
7 <?t?, 1 ?.
In the aberration the central fascia is completely lilack- green from costa to
inner margin.
8 <?<?.
There appear to be no intermediates.
145. Eucymatoge placens spec. nnv.
Forewing : whitish green, overlaid with grey ; the lines black ; edge oi basal
]iatcii black, close to base ; band beyond it broad, especially towards costa, its
centre filled up with dark grey; inner edge of central fascia at one-third, angled
strongly in cell and less so on submedian fold ; outer edge at three-fifths, indented
below subcostal, projecting on veins 4 and 3 and lunulate-dentate inwards ; both
edges black, and starting from black costal spots; cell-spot black, small ; two
grey lines traverse the fascia ; outwardly it is edged with a pale green baud
with darker centre, which is followed by a jiinkish lilac band, not always clear ;
submarginal line pale green, waved, filled up with blackish grey scales ; the
marginal area dark green mixed with pinkish ; both it and the praesubmarginal
shade, as well as the line itself, are interrnpted between 3 and 4 by a white
blotch ; marginal festoon black ; fringe mottled dark and light grey.
Hindwing : whitish grey, with traces of lines, and a paler submarginal band.
( 1-^1 )
Uiiilcrsido of forewing glossy cinereous, ol' hiiidwiiig wliitisli ; tlii' linos all
shown, lint mnch more clearly in hindwing.
Head, thorax, and abdomen grey mixed with green and i)iiik scales.
Expanse of wings : 22 — 24 mm.
2 ? ?.
146. Eucymatoge rufilimata spec. nov.
Forewing : white, dnsted with grey ; the Hues grey, darker on the veins ;
basal patch dark grey or brown-grey, edged by a brown-black band with a slight
projection ontwards above median, and crossed by two dark lines ; bands of central
fascia tinged with grey, darker towards costa, where the inner band forms a
triangular blotch ; the inner band curved parallel to edge of basal patch, the
outer insinuate beyond cell and forming a beak projecting between 3 and 4,
interrupted by the pale ground-colour above 0 ; the pale bands on each side and
the fascia itself crossed by waved grey lines ; the outer half of outer pale band
beyond cell marked by a chestnut-brown curved streak ; submarginal line whitish,
waved, preceded by a dark grey band above middle, paler below, its Innules
followed by blackish streaks to margin above middle, and grey ones below ; the
veins swollen into brownish spots between them ; pairs of black marginal spots
at the vein-ends ; fringe white, grey-mottled ; cell-spot blackish, linear.
Hindwing : whitish, with angled grey postmedian and curved submarginal
line, the margin itself grey ; marginal spots dark grey ; fringe white.
Underside of forewing dark grey, of hindwing whitish ; cell-spots, outer
lines, and marginal clouds distinct.
Head and jjalpi white, the base of each segment externally black ; thorax
and abdomen white speckled with blackish ; the liasal segments of dorsum dark
grey.
Underside of abdomen dark grey like the legs ; fore- and middle tibiae and
tarsi black, with white joints ; pectus white.
Expanse of wings : 27 mm.
4 cJc?,l ?.
147. Eucymatoge subriibescens spec. nov.
Forewing : dull green ; basal patch and central fascia dark fuscous, with fine
black waved lines along their edges followed by an equally fine white one ; edge
of basal patch slightly curved ; inner edge of central fascia at two-fifths, outer
edge projecting outwards at vein 4 ; the outer portion of fascia green from vein 5
to inner margin, the fuscous-scaled portion narrowing to margin ; a black cell-sjiot
and two dark wavy lines across it ; green bands preceding and following fascia
with their centres grey ; submarginal line indistinct, wavy, whitish, preceded by
a reddish grey band, with grey beyond it ; marginal festoon black ; fringe reddish
grey.
Hind wing : rufous grey, without markings, except a dark cell-spot.
Underside of both wings rufous, with a greenish tinge towards base ; outer
line grey, angled in middle on forewing, curved and wavy on hindwing.
Head, thorax, and abdomen pale green, speckled with dark ; second dorsal
segment with a black metallic ring ; dorsal tufts of radiating black hairs.
Expanse of wings : 22 mm.
2 ¥?.
( 122 )
14S. Gymnoscelis crassifemur sjn-c. nov.
Forcu-iiuj : smooth pale green, with pale violet shading ; the lines black ;
inner line at nearly one-third, angled ontwards on each fold, to inner margin near
base; followed by a broad curved violet b:ind; outer edge at two-thirds, angled
on 6 and 4 and insinuate between, then inwardly oblique to tliree-fourths of inner
margin, preceded by some violet shading, the interval with one or two violet lines ;
snbmarginal line green, dentate, the teeth filled up with dark ; the marginal area
violet ; a fine dark marginal line ; fringe violet,
Iliiidwini/ : with the base violet; the rest as in forewing ; veins in l)oth
wings dotted with black scales.
Underside pale green, overlaid with violet-grey in forewing, except along
inner margin, witli two dark curved outer lines and cell-spot, all much i)lainer in
hindwing.
Head, thorax, and abdomen green, more or less obscnrcd by violet-grey
scales. Underside of abdomen, and the legs shining greenish white ; fore-femora
heavily clothed with rnst-colonred furry scales.
Expanse of wings : 17 mm.
1 c?.
Micromia gen. nov.
Forewing: costa with a prominent slioidder at one-fonrth from base, thence
incurved to nearly three-fonrths ; hindmargin curved ; inner margin convex.
Ilimlirinq : with well-rounded hindmargin.
Abdomen with short dorsal tufts; luetathoraeic tuft erect, bifid; forehead
with a bifid tuft ; ]ialpi porrect, first and second segments hairy beneath, second
twice as long as first, third half as long as second, slender, pointed, from the toji
of second ; antennae simple, in the c? scarcely pubescent ; tongue and frenulum
present ; hindtibiae with four spurs.
Neuration as iu Tcphroclystia and Cosmorlioc—ihc areole simple ; in the
hindwing 3, 4, and 6, 7 from the cell, not stalked.
Typo : Micromia fulviptinctn spec. nov.
14',i. Micromia fulvipuncta spec. nov.
Forcwiiuj : deep olive-green ; the basal i)atch, the bands forming the edges of
central fascia, and the praesnbmarginal band still darker green, especially at costa,
where they widen out ; costa along the paler intervals whitish ; inner band of
central fascia swollen on inner margin as well as on costa ; the dark green markings
finely edged with whitish green ; the jiracsubmarginal band with a round fulvous
spot between veins 4 and 0, and two smaller sjiots on veins 1 and 2 ; sulimarginal
line indistinct, showing some paler green lunules ; marginal lino fine, black-green,
with blackisli dashes between veins to the submarginal lunules, those l)eyond cell
strongly marked ; fringe green, in the outer half mottled dark and light.
Hindtcimj : grey, darkening to margin; black lunules between veins at
margin ; fringe grey.
Underside shining grey, with traces of lines ; costa of forewing pale fulvous,
darker at the lines ; hindwing with five dark curved crosslines.
Head and thorax green ; metathorax paler ; abdomen dull fulvous with dark
dorsal markings ; underside of abdomen pale; tarsi dark, with pale joints.
Expanse of wings : <? 24 mm. ; ? 20 mm.
5(J(J,1 ?.
( 123 )
Polysphalia gon. nov.
c7. Foreidng : costa rough-haired and funning a i)rae-apical lobe ; apex ronrKicd ;
hindmargin obliquely curved.
llindirinij : narrow ; hindmargin indented beyond cell.
Antennae lamellate ; palpi porrect, decumbent ; hindtibia with four spurs.
Neuration of Chlorocb/stis, but distorted ; the cell-fold raised above and
furrowed below towards hindmargin, the lower radial being obscured; veins 1'2
with 11 and 10, running into the lobe, 9 and 8 into the costa at its end, and 7 into
apex ; the median vein is thickened and contorted at liase, the wing memlirane
beneath it partially hyaline ; the fold in cell bears a ridge of black erect scales.
Neuration of hiudwing, and of both wings in ? , normal.
Type : Fohjsphalia cristigcra spec. nov.
loO. Polysphalia cristigera spec. nov.
Fcirriciiuj : pale greyish green, with numerous waved greyish lines across it;
four in basal area, the fourth ])eing the inner edge of central fascia, angled on
median and marked by blackish dots on veins ; outer line at two-thirds, excurved
aliove, marked by blackish teeth inwardly on veins, preceded by two grey lines ;
snbnmvginal line pale, waved, the teeth filled up with dark grey, blackish below
costa and beyond cell ; margin between 3 and 4 slightly paler ; marginal lunules
black ; fringe chequered green and greenish white. In the c? the cell-fold bears
a ridge of black scales iu cell, and the furrow beyond is marked with rough
reddish scales.
llindniiii/ : with a dark bar at base ; the rest as in forewing ; the outer line with
its black vein-dashes distinct and followed by a pale green band with dark centre.
Underside grej'-green in forewing, whitish green in hindwing, with the
lines dark.
Head, thorax, and abdomen green, speckled with dark ; the face and vertex
clearer.
Expanse of wings : cJ 15 mm. ; ? 2ti mm.
1 c?, 1?.
The ? is more thickly speckled with grey.
Prosthetopteryx gen. nov.
Forewing: with neuration of Tcphroelg.^tin, i\w arcole large and single; the
structnre of wing normal.
Hindwing: in ? normal also ; in c? subject to numerous variations of structure.
In the typical section, including raexiata and rfltundata, a deep cleft runs up
towards base below vein 3, forming a large triangular anal lobe, like an additional
wing ; this lobe is densely fringed, and below covered with woolly hair ; median
vein at base with a pencil of hairs; the wing above vein 3 rounded. In another
section, comprising catilinea and barbata, a second cleft, less deep, occurs below
vein G, so that the wing is divided into three parts, all more or less contorted and
hairy, the wing in one case, j)arvijien?iatii, being quite dwarfed; or again, as in
liridisecta, the lower cleft becomes a short rounded excision and the anal lobe is
quadrate and bifid. Again, in a third sectiun, latistriga, the cleft is short and
below vein 2, and the upperside of wing is hairy.
In the typical section vein 8 anastomoses with subcostal to near end of cell ;
( 124 J
veins 6, 7 are stalked ; the discocellular has a long ohlicjue lower arm, and the
radial (vein 5) is stalked with 4 from the lower end of ceil, vein 3 rising mnch
nearer base, veins 1 and 2 being left in the anal lobe ; the neuratioQ is the same,
as far as can be seen, in the second section ; but in the third vein 3 rises closer to
the stalk of 4, 3, and vein 2 some distance before, vein 1 alone passing into the
anal lobe. In all alike the cell is broad at extremity and veins 0, 7 cnrved
downwards and divergent. In the hairior forms, such as harhnta, the abdomen
beneath is tufted with hairs.
Type : Prosthetopteri/x caesiata spec. nov.
In Tri ptcridia Warr., which also has the hiudwing divided into three lobes,
the forewing too has a deep cleft in hindmargin.
151. Prosthetopteryx barbata spec. nov.
Forcirimj : dull green ; the markings dark green : viz. the basal patch with
its edge obli(|ue outwards ; the central fascia broad, oblique outwards to middle,
then inwards, its inner edge approaching basal patch on inner margin; the pale
green space between them with dark green centre, broad at costa and narrowing to
a ])oint at inner margin, angled outwards in cell like the fiiscia itself ; beyond the
pale green band edging the fascia externally is a dark fascia with a rufous tinge
before the indistinct pale submarginal line ; fringe probably green.
Ilindwing : deeply three-Iobed ; all the lobes dingy grey and fringed with
long curled hairs ; the middle lobe pointed and darker grey, with a curled wisp of
ochreons grey hairs on its inuer edge.
Underside of forewing pale greenish, with the dark markings showing through :
hindwing with the middle and inner lobe thickly covered as well as fringed with
long fine woolly hair ; a wisp of black hairs at the end of the short npper cleft.
Head, thorax, and abdomen greenish.
Expanse of wings : 28 mm.
1 c?.
The apex of forewing is acute.
152. Prosthetopteryx caesiata spec. nov.
Foretving : deep brown-black, with a slight reddish tinge ; the lines blackish,
indistinct, except in certain lights ; a curved line near base ; another, double, at
about one-third ; an outer line, also double, and dentate outwards ; a dentate
submarginal line ; fringe dark brown.
Ilindwing : bluish slate-colour, the fringe included ; the inner lobe with its
fringe shining white.
Underside of forewing dull brown overlaid with dull bluish slate; the lines
darker in costal half; fringe brown with paler mottlings ; hiudwing slaty blue,
with dark cloudy cell-spot and cnrved outer line ; the anal lobe white as above ;
median vein with a pencil of purple hairs.
Head, tliorax, and abdomen like wings ; anal tuft blackish ; underside of
abdomen, legs, and pectus whitish ; dorsal tufts erect, long, bluish black.
Expanse of wings : 22 mm.
2 SS.
The hindmargin of hindwiugs is curved from costa to end of vein 5, where it is
bluntly angled, and again incurved to base of anal lobe ; in one example the angle
is much more prominent than in the other.
I
I
( 125 )
153. Prosthetopteryx cavilinea spec. nov.
<S. Forcwing : pale green; the markiugs reddish l)rowu ; the lines Ijlackish ;
first Hue near base ; inner edge of central fascia oblique outwards and forming a
small projection into the fascia above median vein, then oblique inwards ; outer
edge obli([ue outwards to below vein 4, then oblique inwards, followed by a pale
green baud with darker line in it, the fascia filled up with dark, and between 3 and 4
overlaid by a red-brown shade which extends to submargiual line, obscuring the
markings ; a dark patch before subniarginal line bayond cell ; marginal area green ;
some dark marginal spots before the green fringe. The inner margin is very strongly
lobed beyond middle.
IlimhciiKj : with the anal lobe grey, darker towards margin, where it is fringed
witli very long dark grey hairs ; the other portion of wing with a slight cleft
below vein 0, the inner part twisted over and fringed with long grey erect spatulate
hairs.
Underside of i'orewing with the dark markings showing through ou a dull green
ground, the inner margin pale : hiudwing blackish in outer lobe, the anal lobe
whitish, clothed with very long yellowish-white hairs, and dark only in the fringe
at end.
Head, thorax, and abdomen greenish; the anal segment dark grey.
¥ . Forewing : paler green ; the fascia at costal end darker ; the whole of the
markings cut off below vein 4, leaving only their edging lines rufous ; the basal
patch blackish and edged by a straight line.
llindwiiig : pale green, with grey marginal border.
Underside of forewing with all the lines diffusely dark grey and wavy on a pale
green ground, not reaching below middle, except the submarginal lines : hiudwing
with the base grey, then with two simple and two double curved and wavy
grey lines.
Head, thorax, and abdomen pale green ; the head and thorax much mixed with
fuscous scaling ; the abdomen with dark dorsal spots.
Expanse of wings : 25 mm.
1 t?,l ¥.
Neither example is in good condition, and the description is necessarily
deficient.
154. Prosthetopteryx latistriga spec. uov.
tJ. Forewing : greenish brown, towards hindmargin more greenish grey ; basal
patch dark, edged by a black line ; inner edge of central f;iscia bluntly angled
outwards in cell, then vertical, blackish ; the centre of the band between it and
basal patch grey-brown, edged on each side by a pale greenish line; outer edge of
fascia oblique and nearly straight outwards to below vein 4, then obli(juo inwards
to before anal angle ; tlie fascia is dark brownish fuscous, crossed by one or two
dark lines and with a black cell-spot ; baud beyond fascia uniformly broad, pale
green to vein 5, then dark green, almost hidden below by a blackish patch, which
includes the lower outer half of central fascia and the iiraesnbmarginal shade; this
last is edged by the pale waved submarginal line, and is interrupted along with the
line by an elongate flesh-coloured oblique streak running from vein G to apex ;
marginal area and fringe greenish grey.
In the ¥ the green spaces are all more prominent, and the broad green outer
band complete ; a marginal line of black dashes.
( 126 )
liindwing : jiale ulive-gre}' ia costal half, dark purplish or bluish grey in the
inner half, iuchuling tjie anal lobe, which has the fringe greenish grey ; the ilark
part of the apical portion covered witli woolly hair. In the ? the wing is jiale grey,
with three darker grey central lines and broad dark grey margin ; an interrupted
dark marginal line ; fringe mottled dark and light grey.
Underside dull greenish grey, rather shining ; the markings obscure in the S,
paler and regular in tlie ? .
Head, thorax, and abdomen dnll greenish; tlie jiatagia blackish; eruire of
basal segments of abdomen and the i)racanal segments blackish ; anal lul'L of i
white.
Expanse of wings : 2S mm.
1 J,3 ? ?.
loo. Prosthetopteryx parvipennata spec. nov.
S. Vorcwiiig : pale green; markings dark fuscous; basal iiaf<h narrow, the
edge oblique ; edges of central fascia fuscous, the inner slightly bent outwards on
median, the outer angled below 4, showing a deep indentation on vein 7; some
indistinct lines across it, esjiecially a dark one near inner edge ; on each side of fascia
a pale green band with darker thread ; praesubmarginal shade brownish, darker
below costa; submarginal line very obscure; black marginal triangles; fringe
clieqnered dark and pale greenish grey. In the ? the wing is altogether greener,
the bands being inconspicuous, and grey-green edged with blackish.
Hinduiiig : in <? minute, whitish ochreous with a greenish tinge, fringed with
white ; iu S normal, greenish grey, with traces of lines on inner margin only, the
outer line clear.
Underside of forewing dnll greenish, with all the lines grey ; of liindwing
iu ¥ whitish, with five curved grey lines, of which the postmedian is distinct, and
a marginal border.
Head and thorax pale greenish, the patagia with black scaling ; abdomen jiinky
ochreous, more fulvous on dorsum.
Expanse of wings : S 17 mm. ; ? 21 mm.
1 cf, 3 ??.
In hindwing of J the anal lobe is quite small and rcuuided, the cleft oidy
reaching halfway ; the cleft below vein (1 is slight, the lobes inconspicuous.
156. Prosthetoptei'yx rotiuidata spec. uov.
cJ. ForcwiiKj : dark brown, with the intervals j)aler, more reddish, than in
caesiata, and towards the costa greenish ; the band beyond basal patch and a
semi-oval patch on hindmargin above middle, containing a black blotch beyond
cell, dull pinkish ; the intervals on costa iu middle indistinctly paler, with a dark
line in each.
llinihniKj : greyer, not so bluish as in raesiata, the extreme hindmargin and
fringe whitish ; hindmargin rounded, not angled, at vein .').
Underside much as iu caesiata.
?. Forewing: with the dark spaces much clearer, and the green and reddish
intervals distinct.
llindiciiig : pale grey, darker aloug the hindmargin, which is limited by a
curved submarginal line.
( 1-^7 )
Underside pale gre)-, without any blue or slaty tinge, with all the lines fairly
distinct, dark grey.
Head, thorax, and abdomen paler in ? than S, the aual half of abdomen
entirely red-brown.
E.xiianse of wings : c? ~2 mm. ; ? 20 mm.
1 <S,2 ? ?.
loT. Prosthetopteryx viridisecta si)ce. nov.
S. FoiriviiKj : dnll purple-brown, with darker markings, and mi.\ed in places
with pale green ; a dark curved line near base, followed by a paler brown space
before the curved and waved inner edge of central fascia, which is succeeded by
a broad pale green and white band from middle of costa to three-fourths of inner
margin, which leaves only the costal half of the outer edge of central fascia to
appear beyond it ; an indistinct dark brown praesubmarginal shade, paler in the
middle ; submarginal Hue indistinct, the margin beyond it pale greenish brown ;
fringe brown and greenish, mottled.
llimhciiig : brownish grey ; the cleft before anal lobe not running up to base,
but only halfway and wide ; the anal lobe not triangular and large, but small and
bifid at end, each tail upturned and rough-fringed ; a slight cleft below vein (3, the
middle lobe so formed contorted and hairy at extremity-.
Underside dark grey, hindwing more brownish ; forewing with traces of lines.
Head and shoulders pale brownish ; thorax dark brown ; abdomen redder
brown ; the anal tuft whitish ochreous.
In the ? the paler intervals of forewing are tinged with green ; the green
band is more clearly divided into an inner green and outer pale half, and the outer
edge of central fascia is shown by a fine pale green line ; the submarginal line is
paler, waved and greenish, and the marginal space greener, exce)it the dark apex,
which is edged by an oblique line to central fascia.
lUmlwing: of normal structure, rather narrow, with hindmargin indented
beyond cell ; pale grey, darkening to hindmargin, with faint traces of lines ; fringe
pale.
Underside dark grey, with a paler middle band ; hindwing with two curved
I)aler bauds separated liy a dark curved lino and with a faint line at middle of each.
Head, palpi, and shoulders greenish ; thorax greenish, with the patagia dark ;
abdomen with basal half and anus dark, the praeanal segments greenish.
Expanse of wings : 26 mm.
1 c?, 1 ?.
Ptychotheca gen. nov.
A development of i'ldorodi/stis.
The forewing of the c? has the membrane from base of cell nearly to hind-
margin folded longitudinally so as to form a double pocket : on the u])perside this
pocket is open towards costa, on the underside towards inner margin ; the veins
beyond cell are partially distorted ; the fold is much more prominent beneath,
appearing as a double pleat with a furrow in the middle.
Type Ptychotheca pallidicircns Warr., described from the ? only as a
Chlorocli/stis, ]^oc. Zool. x. p. 378 (100;5).
Of this species 0 ? ¥ and 1 S have been received.
( 1^8 )
Pycnoloma gen. nov.
Forewhitj : costii strongly slionldered near base and curved tlirougliout ; aj)ex
jiroruinent, but blunt ; liindiuargiii obli(|ne, not curved.
Uiiulwing: narrow; apex bluntly rectangniar ; biudmargin vertical to vein 4,
then oblique to anal angle.
Forehead veith a cone of scales : jialpi porreet, rongh-scalcd, terminiil joint
dej)resscd ; tongue and frenulum ])resent ; antennae broken.
Costa of hiadwing convex and thickened, with a bed of mealy scales on its
upjiersidc near base.
Neuratiou as iu Chloroclystis ; the cell of forcwing short.
Type Fycnoloma rufibasalis spec. nov.
158. Pycnoloma rufibasalis spec. nov.
Forewing : grey-green ; basal patch quite small, its edge marked by black dots
on veins ; inner edge of central fascia dark grey, oblique outwards from one-fourth
of costa, angled in cell, then oblique, marked by black dots on veins; outer line
from three-fifths of costa to middle of inner margin, angled on vein 4, and marked
by vein-dots ; fascia followed by a rather wide pale green band with grey centre ;
snbmarginal line waved, preceded by a grey-green baud, the lunules at costa, bcyund
cell, and above inner margin slightly blackish ; a blackish marginal line interru2)ted
at the veins ; fringe green.
Ilindwing : whitish, tinged with green, with traces of lines.
Underside of forewing greenish .grey, of hindwing jJale green, with the whole
basal area dull pink.
Head, thorax, and abdomen pale green.
Expanse of wings : 15 mm.
1 c?.
159. Syncosmia colorata spec. nov.
Foreimig : pale green ; the markings brownish, edged or varied with fuscous ;
a spot at liase of costa ; a brown basal band slightly curved ; inner edge of central
fascia broad at costa and double, only the outer dark line reaching inner margin ;
outer edge marked only by a black line, often obscured ; bands on each side green
with a faint line down tiiem, slightly jialer than fascia itself: praesubmarginal
band brown, iu costal half double and edged by waved black lines : snbmarginal
line acutely dentate, whitish green ; marginal line fine, blackish; fringe pale, broadly
chequered with brown.
lliii<hciiig : of S dirty ochreous, without markings ; of ? ochreous grey with
darker marginal border, and traces of curved lines.
Underside greenish grey, with the markings of forewing showing through.
Palpi externally ferruginous, above with long green and metallic black hairs ;
vertex, collar, and shoulders pale green ; thorax and abdomen greenish ochreous
brown; metathorax edged with metallic black scales ; dorsal segments ringed with
black ; dorsal tufts of radiating metallic black hairs.
Expanse of wings : S 21 mm., ¥ 23 mm.
2 Jc?,0 ??.
( 129 )
I'll). Tephroclystia albibisecta spec. nov.
Forcwmg : deep brown-lilack, with darker but very indistinct cross-markings ;
a cnrved line near base, and a second, .geminate, at abont one-third ; from jnst
beyond middle of costa to beyond middle of inner margin a broad straight white
line, dark-bordered on each side, followed Iiy a thicker dark line ; outer line double,
forming black angular markings outwardly, interrupteil, like the lines before it,
between veins 2 and 4 ; snbmarginal line represented by a row of small white spots,
often obscured, except those on each side of vein C, wliich are larger ; fringe con-
colorons ; the intervals between the lines are tinged in certain lights with reddish,
especially along costa.
Jlituhcitig : jieariy-wliite, becoming grey towards liindmargiu ; marginal line
black, interrupted by pale spots at the vein-ends : fringe pale grey.
Underside dull brownish grey, jialer towards inner margin, the pale cross-line
and the snbmarginal sjKits showing through ; hiadwiiig as above.
Head, thora.N, and abdomen blackish ; the two segments beyond middle dull
reddish. In the ? these segments are pale ochreous and swollen.
E.xpanse of wings : 20 mm.
2 cJc?,2 ??.
In one ? the white bar is bent inwards at costa and swollen on inner margin.
161. Tephroclystia coerulescens spec. nov.
Forewiiiit : deeji velvety brown, towards costa paler and brighter brown ; basal
patch edged by a shining white band, interrupted below median, and yellowish at
inner margin ; central iascia broad, crossed by paler and darker shades, which at
costa are marked with jiale shining scales ; the subcostal vein marked at intervals
by bluish silvery scales ; central fascia edged outwardly by an obscure paler band,
marked witii cream-colour on costa and beyond cell, and by a double blue and
whitish line on inner margin ; submarginal line formed of bluish spots or Innules ;
intermixed lilack marginnl dashes, ])ree,eiled in jilaces by jjale ochreous scales;
an oblique ochreous ai)ical streak ; fringe deej) brown, its ajiical half with small
whitish spots, the spots at ape.\-, between veins .0 and (1, and 3 and 4 large, square,
and running to base of fringe.
Hindwiny: brownish fuscous ; fringe brown, with jiale spots at base.
Underside of forewing blurred grey-brown, the costal and hindmargins bright
brown with bluish spots; fringe as above: hindwing brown, crossed by curved
double bands of spots of blue scales.
Head, thorax, and abdomen dark brown dotted and spangled with Idue scales;
palpi paler brown; abdomen beneath and leg.s dark brown sprinkled with blue; the
joints of the legs pale ochreous.
Expanse of wings : S 2U mm., ? 24 mm.
9 c?c?,l ?.
102. Tephroclystia curvimacula spec. nov.
Fon'wimi: olive-green; the edges of the usual markings blackish, augulated
and interrupted ; costa dotted with black at the commencement of all the lines ;
basal jiatch crossed and edged by a thick black line; inner edge of central fascia
angled in cell, and thickened below median ; a similarly angled thin line through the
baud before it; onter edge forming a large (juadrate "black blotch fi'om vein 7 to 5,
( 130 )
with a curve from its inner edge to vein 4, a black spot on 3, and blotch from 2 to 1 ;
snbiuar<rinal line acutely dentate, the teetli filled np with dark green preceded by a
fulvous shade which rnns obscurely to apex ; a fnlvous streak above median along
cell, and one along inner margin ; marginal festoon blackish ; fringe green ; a small
black celI-si)ot; fringe of inner margin green dusted with black scales.
lliixltcinii : pale grey, darker towards hindmargin.
Undersiile of forewing cinereous, pinkisii ahmg costa, with the lines obscure;
hindwing juiler towards base, with an angled median and waved submargiual
dark line.
Head and thorax green spotted with liiaek; abdomen flesh-colour.
Expanse of wings : ~!G mm.
3 ??
There is a remarkable superficial resemblance between this species and
Micromia fulcipuncta Warr.
I(i3. Tephroclystia decens spec. nov.
Forewiitii : jiale green, with the markings liglit purplish grey ; these are the
baud limiting the basal jiatch, the central fiiscia, and a costal blotcli before apex ;
outer edge of basal patch from one-fourth of costa to one-third of inner margin,
angled on subcostal vein ; inner edge of central fascia from one-third of costa to
two-fifths of inner margin, bent on subcostal, and preceded by a fine dark line in the
green band ; outer edge of fascia at nearly two-thirds, outcurved beyond cell; the
fascia contains a dark cell-spot and is traversed by two or three wavetl dark lines ;
the green band beyond has a dark line dnwn tlie middle ; submargiual line very
obscure, jireceded by a very faintly darker band from the grey costal spot; a slight
dark marginal line; fringe greenish rufous.
HindwiiKj : pale greenish grey, darker tnwards hindmargin ; fringe paler.
Underside of both wings dull pale green, with the markings faintly showing ;
fringe paler.
Head, thorax, and abdomen pale green ; thorax and abdomen slightly varied
with purplish grey ; forelegs purplish witli the joints greenish.
Expanse of wings : 21 mm.
1 ?.
104. Tephroclystia flavicorpus spec nov.
Foreu-ing : deej) velvety brown, slightly paler at base and extreme apex:
markings pale buff or cream-coldur, edged witli silvery white; interval between
basal iiatch and central fascia represented by two triangular marks at one-third,
one on costa reaching the median, the other from inner margin reaching snbmedian
fold ; outer edge of fascia indented in cell and bulged at middle, followed by a
silvery white line edged above and below the bulge with buft'; a silvery whitisli
submargiual line of points ; two yellowish oblique streaks from outer line above
vein 0, one to ajjcx, the other to costa before apex ; a buif marginal spot between
veins 3 and 4 ; marginal Innules dark brown followed by white spots at the base
of fringe, which is brown, its outer half dotted with white.
Hindinng : dark grey.
Underside of forewing blurred brown ; subcostal streak orange ; apex tinged
with fulvous, with some white and bluish spots : hindwing brown sprinkled with
blue scales forming four double carved Hues.
I
( 131 )
Head, thorax, and alulomon pale oclirpons, tinged with fulvous ; dorsum
fnlvoiis, tbe seiimeiitiil rings broadly wliite ; abdomen below and legs dark brown
speckled with blue.
Expanse of wings : c? 18 ram. ; 9 20 mm.
4c?c?, 2 ¥ ?.
165. Tephroclystia oruatipennis spec. nov.
ForewiiK) : dark l)rown, witli an admixture of fulvous towards hiudmargin ; the
interspaces cream-colour or white ; costa strigulated with white ; basal patch crossed
by a pale line generally resolved into points ; central fascia containing a white bar
on discocellnlar and two lines of whitish jjoints, preceded and followed by a band
of cream-colour ; the inner band broad, with pale ochraceous centre and brown on
costa and inner margin, sometimes interrupted below median vein by the junction
of basal patch and central fascia ; outer band narrower, tinged with ochraceous on
its outer edge, almost interrupted Vietween ?> and 4 by tiie projecting tooth of central
fascia; snlimai'ginal line waved, white, outbont between veins 6 and 2; a fulvous
apical blotcli ; a slight marginal line of dark lunules ; fringe brown with broad
flecks of wliite between veins.
HiiulwiiK/ : dull whitish, tinged with pale grey and ochreous ; markings of
underside showing through.
Underside of forewing with basal two-thirds blurred ; an orange subcostal
streak: hindwing white, thickly brown-speckled, and with five darker bands, often
more or less obscure, three central and marked cliiefly towards inner margin, a
submarginal before the wliite line, and a marginal.
Head, thorax, and abdomen dark brown, speckled with white ; segmental rings
white ; abdomen in ¥ fulvous, sometimes the head and pal])i also.
Expanse of wings : d 22 mm. ; ? 23 mm.
5 cJc?,3 ¥ ¥.
160. Tephroclystia stabilis spec. nov.
ForewiiK/ : grey-green ; the bands edging liasal [latcli and central fascia double,
blackish, dentate inwards on veins and marked throughout by raised scales; the
green intervals with a dark waved central line ; praesubmargiual shade also double,
blackish, but without raised scales; submarginal line waved, whitish, followed by a
dark cloud at ajiex ; marginal lunules black ; fringe mottled pale and greyish green.
Hi ml wing : pale grey with a slight reddish tinge, and traces of dark lines;
lunules along margin black and prominent.
Underside greenish grey, with all the lines in both wings dark grey, waved,
and distinct ; liiudwiiig redilish-tinged.
Head, thorax, and abdomen olive-green ; metathoracic and dorsal tufts large,
of erect hairs.
Expanse of wings : 25 mm.
1 ?.
107. Tripteridia albimixta spec. nov.
Forewing : dark olive-brown, crossed by two shining white lines ; the first at
one-third, the second before two-thirds ; both starting narrow at costa and widening
below it ; the first angled outward in cell, reaching inner margin as a dark shade
edged by two fine lines ; outer line indented below middle and rniiiiing straight
( 132 )
to iuner margin pear inner line; joined to inner line by a whilish smudge below
median vein, which is continued obliqncly across onter lino to ajiex : tlie basal
patch, therefore, is entire; the central fascia is broadly interrnjited below middle;
tiie dark marginal area ibrraing a triangular costal blotch before apex (through
which the beginning of the wavy white submarginal line is visible) and a square
blotch before anal angle, the margin between being paler brown ; fringe cher|nered
dark and light.
Ilimlwinij: yellowish white, j.aler in costal half; the middle lobe with black
scales down its centre.
Underside yellowish ; the dark markings diffusely dull olive ; hindwing
without markings.
Head, (liorax, and abdomen olive-brown, tiie last at sides and bcneatli mixed
with jialer.
¥ wilh the pale markings amplified ; two double pale curved lines near base; a
pair of curved ])ale lines in central fascia from costa ; space between veins 'Z and 4
jiale bud; blurring all markings alike ; outer line distinctly double, white with a
thin brown centre, reaj)iiearing on inner margin, like the two paler lines in the
fascia ; sul)marginal line visible throughout, but not waved, tiie veins across the
praesubmarginal space jiale, continued obliquely as a small blurred space to margin
below apex ; marginal lunules black, distinct, fringe brown, chequered with eream-
colour ; hindwing whitish yellow, with a grey marginal border.
Underside straw-colour, with the dark and light cross-lines showing, but
interruptedly.
Abdomen pale ochreons, the dorsum yellower.
Expanse of wings : ITmni.
2 J<^, 1 ¥.
Xeuoclystia gen. nov.
Agreeing in neuration with Tcijhrochjstiu, 10 and 11 being stalked, and lo
anastomosing with S, !), forming a long simple areole; but the antennae of the i
are bipectinated, the pectinations well separated anil stiff, ciliated laterally and
apically ; in the ? with angulated segments; abdomen of c? laterally tufted towards
anus, the anal segment cut square and ending in a dorsal projection, which is
especially conspicuous in the ?.
Type : Xeiioclyiitia (lelect((ns sjiec. nov.
108. Xenoclystia delectans spec. nov.
Forewiiif/: brightly coloured, brownish grey on a whitish ground, crossed by
numerous minutely waved dark and light lines, two of which form in the middle
of the central fascia a darker space containing the obliijue black cell-mark, and
widening basewards below the middle; lines edging the fascia and basal patch
bright green and white ; submarginal line Innulate, brightly white, jireceded by
a dark band of three lines, the innermost much darker; some green scales before
margin; fringe pale grey, with a dark dividing line and triangularly chequered
beyond veins.
JlimlwuKj: whitish, with grey marginal border; crossed by faint wavy lines
and a jiale band before the border.
Underside of forewing blurred for two-thirds from base: hindwing whitish,
thickly jiepjiered with brcnVn. with three dark wavy lines and paler interspaces.
( 13"^ )
Head, thorax, aud abdomen brown marked witli wliitc and green ; metafchoracic
tnft green ; segmental rings of aljdomen white.
Expanse of wings : 6 ~2 mm. ; ? 20 mm.
6 c?cJ,5 ? ?.
160. Xenoclystia delicata spec. nov.
Forcwiiifi : jiale grey, tinged with vinnus, es[)ecialiy along cosia; the markings
red-brown, edged with a deeper line; basal jiatch oblii|neIy edged ; centra! fascia
broad above, narrow on inner margin, the inner edge indented in cell, tiie onter
exciirved at middie ; botli, as well as basal patch, edged by a pale grey band with
dark central line; s]iace between the two inner pale bands grey; cell-spot linear,
black in a central pale sjiace; marginal area dark grey, with only the teetli of
snbmarginal line siiown, wliite in lower half of wing, blackish above; fringe
reddish, with black chequering beyond veins and black lines at the vein-ends.
Iliudwing : (piite {)ale grey, with a faint reddish tinge ; a dark cell-spot and
curved jwstmedian and snbmarginal line ; fringe reddish beyond pairs of dark
marginal points.
Underside of forewing dnll red, witli the lines greyish on costa and a dark
blotch before hindmargin beyond cell : iliudwing whitish, red-speckled, witli black
cell-sjiot and onter row of black spots.
Face and palpi above white ; palpi externally dark fnscous ; vertex reddish
with a black speck at middle ; thorax grey ; abdomen reddisli with white segmental
rings and jiairs of l)lack dorsal spots.
Exj)anse of wings : 17 mm.
2<?c?.
One of these is much darker and less red than the other.
171). Xenoclystia humerata spec. nov.
Forewing: almost exactly tlie same as in A'. /<«//y^//(cte, described below; the
metallic lines are not so strongly marked, and the white spot in submargiuul line
is less conspicuous.
Uimhoing : dark grey, witli no traces of lines, and the fringe grey, not rufous.
Underside of forewing darker and of hiiidwing paler tlian in miijiimcta.
Head, thorax, and abdomen all dark.
Expanse of wings : 20 mm.
As far as markings go, this species miglit well be treated as a form of
X unipuncta, but the hindwing of the cj is strongly shouldered at base and the
costal and subcostal veins are curved upwards and bent parallel to it ; while in
tinipuneta the costal edge is jierfectly straight.
171. Xenoclystia unipuncta spec. nov.
Fninriiiij : vinous grey ; the cross-markings vinous brown ; the edge ol basal
jiatch, at one-fourth, the bands edging central fascia, :it abont one-third and two-
tiiirds, and to a less extent the praesubmarginal band, are dark brown, all in the main
parallel to hindmargin, but the outer edge of the central fascia projecting below
middle ; all the dark markings are edged with a broad line of dull metallic scales ;
the liasal ])atch atid central fasciaare traversed at middle liy a band of jialer ground-
colour, in the former case with a dark middle line ; the lower part of the middle
( 134 )
of leiitral fascia is clothed with metallic scales ; snbinargiiial line formed of the
same scales, and a few before the black marf:;iiial line ; frinj^c dark brown with
paler tij)s ; the submargiiial line shows a small white spot between veins 3 and 4.
In one ? the middle of central fascia is flesh-colour, showing a faint dark cell-spot.
Ilindiviiig : grey, darkening towards outer margin, with two obscure dark
lines before margin ; marginal line black ; fringe bright rufous.
Underside of forewing dark blurred grey, reddish towards costa and hind-
margin ; the lines slightly visible, esjjecially on costa: hiudwiug greyish ochreons
tinged with red, with two dark grey lines ; fringe rufous.
Head, thorax, and abdomen dark brown.
Expanse of wings : 2U mm.
6 <?c?, 4 ??
Subfamily DEILINIINAE.
172. Borbacha carneata spec. nov.
Foreiviny : deep flesh-colour ; the costal edge snow-white ; all the markings,
which are much as in B. pare/aria Gnen., greenish-grey finely edged with purj)le ;
the veins in marginal area thickly greenish-grey, excejit vein 5 ; in the broad
outer fascia its outer edge is preceded by a series of ])urple lunules, and between
veins 3 and 4 the submargiual line contains a ])urple blotch ; a row of purple
marginal triangles ; cell-sj)ot ])urple ; fringe flesh-colour, grey beyond veins ; two
purple dots near base, one on each side of the median vein.
llimlwiiHi : with a grey liar with luinilate edges near base; marginal area
between veins 2 and 4 greenish-grey ; the puri)le lunules less conspicuons.
Underside pale yellowish, with all the markings dull purple.
Vertex, upper half of face, and ])alpi above dull brown-red ; shoulders and
base of patagia greenish-grey ; rest of patagia flesh-colour, marked with i)nriilish ;
thorax and abdomen greenish-grey, with flesh-coloured dorsal spots and edges of
segments ; the metathorax wholly flesh-coloured ; fillet and shaft of antennae
snow-white.
Expanse of wings : 48 mm.
1 ?.
173. Syntaracta albibasis spec. nov. and ali. pUimbea nov. and atriplena nov.
?. Foiririiig : yellow, suftused with orange-red and thickly sjieckled with
brown-grey; the costal area and the lines brown-grey; the lines and shadings
almost precisely as in & clathrata, described below, the yellow ground-colour
showing only, as the ochreous in that species, as a line before first line and after
the outer and submargiual.
HindicitH/ : similar.
Underside paler, more ochreous, with all the niarkiiigs dull greyish orange.
Head and ])alpi yellow and orange; shoulders and base of jiatagia like costal
streak ; thorax orange and yellow ; abdomen red-brown with the white bar at
base, as in clathrafa, but no pale dorsal marks.
Expanse of wings : 40 mm.
2 ??.
This si)ecie8 may be distinguished from the corres])onding type form of
clathrata ? by the orange-red tinge, as well as by the absence of the iKile dorsal
spots of abdomen ; no <? of this form was received.
( 135 )
Tlie aberration plumbea is analogous to the aberration fasciata of clathrata,
the inner and outer lines being swollen into dull leaden-grey bands only about
half as wide as those in fasciata, the outer band edged inwardly with blackish,
a streak of the same grey filling up the interval between veins 3 and 4 to the
margin, and another streak, bnt much narrower, running along vein 0 ; the space
between this band and submarginal line being filled in with deep red-brown ;
the costa is of the same tinge as the bands.
Head, thorax, and abdomen as in the type form.
3 ? ?.
While the type form and its aberration plnnihca are represented only by ? ? ,
the aberration atriplena is a c? only.
Forcwing : yellow, dotted witji orange, and with the veins orange ; costal
area brownish grey with dark speckling ; the lines swollen into bands, as in the
ab. jilnmbca, but the outer one twice as broad, and both suffused with black, which
to a great extent obliterates the leaden-grey tints ; the bands as well as their
extensions to the hiudmargin are more conspicuous owing to the paler ground-
colour.
Uindwiiuj : the same.
Underside pale ochreons, witli the black liaiids both shown.
Abdomen orange-red with black bars at the hinder edge of each segment.
Expanse of wings : 3s mm.
1 (?.
1 have no doubt that the ground coloration of this c? is the normal ground-
colour of the cj in all its forms ; and it is possible that the aberration afri/j/cna
may be the corresponding c? form to the ? aberration pliunbea ; though in that
case the difference in the coloration of the sexes is much greater than in the
corresponding aberration of clathrata and its type form.
The antennae of the S of both alhihasis and clathrata differ from true Si/iit.aracta
in being serrate, with short I'ascicles of cilia, these being much shorter in albibasis
than in clathrata,.
174. Syntaracta clathrata spec. nov. and ab. fasciata nov.
(S. Forcwing : jiale ochreo\is, speckled with rusty-edged brown granules ; the
veins and lines brown ; costal area above subcostal vein dull grey-brown, the costal
edge dotted with ochreons and brown ; first line thick, curved, at one-third, luiinlate-
dentate, toothed inwardly on the veins and outwardly between them, jirecedcd
by a i)ale ochreons space ; outer line at two-thirds, parallel to hiudmargin, also
double and lunulate-dentate, followed by a line of unspeckled ochreons ; submarginal
line single, indented on submedian fold, followed by a [lale space ; all three lines
start from the subcostal vein ; veins 3, 4, and I) thickened to hiudmargin ; a dark
fine marginal line ; cell-si)ot black ; fringe ochreous, chequered with brown beyond
veins.
Jlii/diciiig : without basal line.
Underside (piite pale ochreous, with all the markings dull grey.
? suffused with grey-brown, the lines and veins grey-brown; only the (hrec
pale edges of the lines remaining clear ochreous, the rusty speckling being much
thicker ; costa ochreous grey, dusted with black.
Face, vertex, and paljii yellow and fulvous ; shoulders concolorous with costa ;
patagia fulvous ; dorsum marked with brown saddle-shaped marks with pale
( 13C )
centres : basal segment with a snow-white bar ; anal segment and abdomen
beneath anil legs jiiile oi'hreons, the legs dnsted with grey.
Expanse of wings : S 42 — 44 mm. ; ? 44 — 48 mm.
4 (?c?,4 ? ?.
In the aberration fasciata the inner line is expanded into a broad red-hrowu
or grey-brown band, and the whole s])ace between onter and snbmargiiial line
tilled nj) with red-brown, forming a still broader band, extending to liindniargin
below apex and beyond cell, and on inner margin uniting with the inner baud ;
these bands show paler on the underside.
3 (?(?, 5 ? ?.
The coloration of'tiie sexes differs as in the type-form.
175. Syntaracta fiilvata spec. nov. and ab. tephrospila nov.
<?. ForcwiiKj : dull deep yellow, covered witli more or less confluent dull
fulvous granular dots, and towards hindmargin speckled with blackish ; costal
urea dull fulvous brown, dusted witii darker ; lines formed of hoary-grey scales ;
first from one-fourth of costa to one-third of inner margin ; second from three-
fourths of costa to two-thirds of inner margin ; both dentate-lnnulate, the teeth
finely blackish, pointing inwards in the first, outwards in the second ; a similar,
but less distinctly outlined snbmarginal line ; space between veins 3 and 4, and a
narrow streak along vein 0, darker ; small dark marginal sjxits at the ends of the
veins ; fringe dull yellow ; cell-sjiot black.
Himlwiitg : similar ; fringe of inner margin and the margin itself narrowly
whitish. In the ¥ the grey scales of the lines are much less distinct, and on the
other ha!id the black dusting is much stronger, and the black teeth of all the lines
and the marginal spots larger and conspicuous.
Underside dnll ochreous, with the lines and speckling grey.
Head, thorax, and abdomen dark and light fulvous ; shoulders and base of
])atagia grey-black ; metathorax and abdomen with pairs of grey or black spots,
which become confluent towards anus, with oblinue divergent markings grey or dark
fulvous ; abdomen laterally and beneath, pectus, and legs i)ale ochreous.
Expanse of wings : c? 38 mm. ; ? 4U mm.
2 c?c?,2 ??.
The ? to which I give the name trj>/o-o.yj//a ab. has on each wing between
vein 4 and the inner margin a large grey blotch with cloudy blackish edges, across
which the black teeth of the second line are visible ; the blotch is etpially marked
on the underside. On the dorsum the spots on second segment are swollen into
a grey and black belt.
1 ?.
ITii. Syntaracta nigrellata sjicc. nov. and ab. scixtigera nov. and
ab. semifascia nov.
Foicwing : yellowish straw-colour, covered with dull greyish fulvous spots;
costa brownish fulvous, the edge dotted pale and dark ; lines and markings exactly
the same as in S. fulmta ; but, owing to the paler ground-colour, the lines are more
distinct, and in all cases the black spots at the teeth and the marginal spots are
large and black.
( 13T )
Hindwing : like iorewing.
Underside jialn sf raw-colour, witl) the lines distinct, dark grey.
Head, thorax, and abdomen pale and dark yellowish ochreous ; shoulders
brownish fulvous with a dark line; base of patagia black; dorsum with pairs of
dark brown or fulvous spots.
Expanse of wings : c?40mm.; ? 4i)— 46 mm.
3 (?(?, 4 ? ?. One of these males is a dwarf, 3n mm., witliout black dots
or spots.
In the aberration scutigera the second segment of dorsum bears a black
cruciform blotch and the anal segment a black sjiot, the others being marked
merely with fulvous. In one example the submarginal line of both wings bears
a large black blotch beyond cell, which is Iblhiwed by another beyond the line, and
in the forewing there are two smaller blotches below costa.
3c?^.
In the aberration scmifnscia the second segment ot dorsum is marked as in
ab. scutigera, and the other segments also bear smaller black marks of a similar
form; the forewing between outer and submarginal lines is tilled in with a black
blotch reaching from vein 4 to inner margin ; in this example, as in the dwarf
specimen of the type-form, there are no black marginal spots or scales denoting
the teeth of the lines.
1 (?.
Subfamily PLUTODINAE.
177. Plutodes connexa spec. nov.
c?. Forewing: pale yellow and red-brown; the costal area above subcostal
vein yellow, with two rounded blotches depending from it and reaching median
vein, one at one-fourth, small, the other at <M)e-half, larger ; extreme liindmargin
and fringe yellow, more widely at apex and anal angle ; on the inner margin
beyond middle another roundeil yeUow blotch reacliing vein '2 ; the edge of the
brown area is ferruginous throughout ; the large brown blotch at liindmargin is
edged with a line of dull lustrous scales, and is traversed throughout by a dark
paler-edged waved submarginal shade, extending as a small jirojection to inner
margin before anal angle ; the fringe is tinged with brown between veins 3 and 4,
and slightly also below apex.
Hindiviiig : with the Ijasal area red-brown, connected along vein 1 by a narrow
streak with the marginal blotch, which is irregularly crescentic, with a yellow
indentation at anal angle ; inner margin and fringe whitish.
In the ? the dark areas are more extensive, and grey-brown in tint rather
than red-brown.
Underside paler yellow, witli only the marginal blotches marked, and these
reduced in size, the other dark areas only showing througli.
Face, thorax, and dorsum concolorous with the dark areas of the wings ;
shoulders, vertex, antennae, paljii beneath, pectus, legs, and underside of abdomen
yellow ; palpi above ferruginous-tinged.
Expanse of wings : J, 34 mm. ; ? , 36 mm.
1 c?, 1 ?.
In the irregularity of outline of the marginal blotch tiiis species resembles
P. nilgirica Ilmpsn.
( 138 )
Subfamily BRAC'CINAE.
1T8. Arycanda commixta f^iioc. nnv.
Forewing : dark pnriilish slate-colour; tlu' lines jiurjilisli lilack ; the paler
spaces thickly striiited with black; a diffuse dark ]iatcb at base; a thick double
antemcdiiui band, the anus separating towards costa ; central fascia formed of a
])air of doid)le bands unitinjf below middle, scjiarating towards costa, the outer anus
huuilate-dentate ; a double submarginal band, and a row of marginal liinuley ;
along the course of the two folds and between veins 3, 4 and (>, 7 in the outer
arm of the submarginal band are patches of dull olive-fulvous scales ; fringe
Iiur]ilish.
Ilimhc'nnj : without the basal lines.
Underside dull jjurplish slate-colour, with dull black cell-sjints.
Head and thorax like wings ; three basal segments and the anal segment of
dorsum black, tlie rest bright yellow ; abdomen beneath, jiectus, and legs slaty
fuscous. In the ¥ the anal segment of dorsum remains yellow.
E.xpanse of wings : c?, .j^ mm. ; ?, Ot) mm.
2 (?c?,3 ? ?.
179. Arycanda fulviradiata sjiec. nov.
ForewiiKj : dark jiurplish blue, covered with long slender black striations ;
two black spots on costa at one-fiftli and one-fourth indicate a curved inner line ;
a spot before middle and tv/o beyond indicate the beginning of the curved inner
and double liuiuhite-dentate outer band of central fascia; an indistinctly double
submarginal line, and a tliick marginal line of conlluent lunules ; from tiie base
of cell a bright fulvous streak runs along the ibid as far as submarginal line,
tilling up the lunules on e;icli side of vein -J ; cell-sjiot bright fulvous ; along the
submedian Ibid a broad dull I'ulvous-olive shade runs, and a jiatcli of the same
colour lies on submarginal line between veins o and 4; fringe purple, with pale
base and broad dark middle line.
Uindwing : with a straight pale band close to b;ise, broadly edged on each
side with black, iiuri)lish at the e.xtremities and fulvous between ; the other dark
bands as in forewing, without fulvous streaks, but the cell-spot and a large spot
between 3 and 4 before hiadmargiu are bright orange ; basal line of fringe orange.
Underside dee]i ])urple slate-colour, with the cell-spots black.
Head, thorax, and abdomen dark pnriilish i'uscons ; dorsum marked with
deeper black spots in middle, containing jiale scales ; underside and legs slaty
fuscous.
Expanse of wings : 58 mm.
1 S.
18U. Bursada radicata al). flavifrons nov. and ab. carens nov.
In a lenglhy series of li. raiUnUn Warr., in which the size and shape of the
three yellow blotches of the forewing vary considerably, two extreme forms single
themselves out.
In the one the yellow markings are broader than in the average typical form,
and (he face as well becomes yellow ; in the other the opposite tendency is seen,
and one or other, sometimes all three, of the yellow bloti'hes becomes nearly or quite
obsolete. For these two forms I 2)roiiose the names ab. Jlaci/rons and ab. carens
respectively.
( 139 )
181. Bursada velata spec. uov.
Foreicing : velvety lirown-black, with two yellow spots ; cue round and small,
near base of cell, generally clear yellow, the other forming- an elongated blotch
beyond cell from subcostal vein to vein 2, overclouded with brown scales ; fringe
concolorons, with the extreme tips pale.
lllndwimi : bright yellow, with thick black margins, that along costa and
round apex twice as broad as the rest.
Underside of forewing with both spots clear yellow ; hiiidwing as above, but
the inner margin above anal angle witii a small quadrate projection towards apex.
Head, thorax, abdomen, and legs all black.
Expanse of wings : 3U — 32 mm.
3 ? ?.
182. Craspedosis costimaciila spec. nov.
ForctriiK/ : Ijrown-bhu'k, with traces of obscure fulvous lines across wing,
somewhat more plainly marked on costa; one from one-fonrth of ciista to one-tliird
of inner margin, one at two-thirds, and two beyond it; cell-spot white, linear;
jnst bej'ond it- a broad wiiite blotch from costa to vein 3, its outer edge curved,
edged on both sides by a thick black line, the outer one strongly incurved to
submedian fold; obscure fulvous marginal spots between veins, tliat between
3 and 4 white, preceded by fulvous marks and another wiiite one ; fringe
concolorons.
Hindwiiig : deeper black, with traces id' fulvous lines on inner margin only;
marginal yellowish dots bi4ween veins : fringe black.
Underside slaty Idack, witliout lines ; tlic white blotch of forewing yellowish
on costa ; the marginal spot between veins 3 and 4 m;uked.
Head, thorax, abdomen, and legs all Mack.
Expanse of wings : 3.5 mm.
1 ?.
Subfamily ASCOTINAE.
183. Alcis conciuna siiec. nov.
Foreirhuj : white, thickly black-sjieekled ; lines black, thick; first from
one-fonrth of costa to one-third of inner margin, liluntlv bent in cell, with some
black scaling before it ; outer line from two-thirds of costa to three-fifths of
inner margin, bent on vein (J, and insinuate on submedian fold, thickened
at veins ; a black median line, obsolete below middle, bent ont round the black
cell-spot ; subniarginal line preceded by blackish shading that fills up the
lunules from costa to vein G, and vein 3 to inner margin, followed by a double
black blotch between 4 and 6 and a small one above vein 2 ; large black marginal
luanles ; fringe black and white, wholly black towards apex.
Iluulwiiuj : with the black sjjeckling much finer; a fine ill-marked black
onter line and lilack cell-spot : a black marginal festoon ; fringe white, thinly
chequered with dark.
Underside the same.
Head and thorax white ; abdomen with the white almost hidden by bro.ad
black dorsal belts, the basal segments only spotted ; anal tuft white; legs black
mottled with white.
( 1^" )
Expanse of wiugs: 24 mm.
1 (?.
The forewiiig luis iiu uuiisually largu lijaliiie I'ovt'a.
1S4. Alcis irrufata .s|iec. nov.
I'omciiK/ : whitish, irroratuil with olive scales; basal area greyer ; first line
okscme, at nearly one-t.hinl, angleil in cell, then oblii|ne, ])re('eded by an olive-
grey shade ; outer line black, sinnous, from two-thirds of costa to three-fifths of
inner margin, jirojecting at vein 5 and insinuate below it, and there blackest,
followed by a broad olive brownish shade : median line brown, ditfuse and
iuterrujited, ajii)roaching onter line on inner margin ; submarginal line whitish,
waved, ])receded and followed by brown shades, the interior marked with black
scales below costa, beyond cell, and in lower third ; the dark edgings of the Innules
connected by dark scales with the black marginal spots ; fringe pale ; no distinct
cell-spots.
lliudiruKi : without first line; a dark cell-spot immediately following ante-
median line ; the rest as in forewing.
Underside dingy whitish ; the nnukiugs and siieckling of the nppcrside
showing throngh ; a grey snbmarginal shade bi'tbre apex of forewing.
Head, thorax, and abdomen like wiugs; tips of shoulders brown ; face above
olive, below whitish; palpi fuscous; forelegs black, with white joints.
Expanse of wings : 44 mm.
1 ?.
Distinguished from Clmijada .■ii'm/i/ixcxfc, which it greatly resembles by the
difference iu the discal spots of hindvving.
Atmoceras gen. nov.
ForeiciiKj : costa slightly sinnons ; apex blniit ; hiiidniargin curved below.
Ilindwiitff : narrow; apex bluntly rectangular; hindmargiu slightly indented
lieyond cell, obliquely rounded below, rnnning into iiuier margin without forming
a distinct anal angle.
I'allii npcnrved, rough-haired, terminal segment oliscure ; antennae of S w'ith
pedicellate fascicles of cilia, at right angles with the shaft ; each fascicle double,
forked at apex, symmetrically ciliated laterally and at apex, the ciliations interlacing
and forming a film; tongue and iVcnulnm present; hindtibiae of S much swollen,
with a pencil of fluffy hairs.
Iu the forewiugs 10 and 11 are coincident.
Type : Atmoceras ptunoxn sj)ec. nov.
Allied to Paralcits.
1<S.">. Atmoceras plumosa spec. nov.
Fomriiui : fawn-colour, irrorated with olive fuscous, when fresh with a
slight pinkish tinge; lines olive fuscous; first from before one-third of costa
to one-third of inner margin, ontcnrved above and below median, toothed inwards
on the veins, preceded by an exactly similar line ; median line lnnulate-dentat«,
the teeth pointing basewards, outcurved above round the deep green cell-spot to
middle of inner margin ; outer line at two-thirds, angled on vein 0 and marked by
(lark spots on veins, rnnning parallel to median line, followed by a greenish shade
( 141 )
which is distinct and separ.ate only at costa and on inner margin ; snhmarginal line
furniing sharp whitish tcefii, tilled iiji inwardly with dark greon, except between
3 and 4, and followed by dark streaks to the dark marginal spots ; fringe fawn-
colonr, mottled with green.
Ilindicinq: dnll fawn, grey-speckled, showing a cell-spot, a mi-dian line, an
outer, and a snbmarginal.
Underside with the dark markings indistinctly showing through.
Head, thorax, and abdomen fawn-colour ; vertex and part.s of thorax mottled
with green ; ]ialpi externally dark; abdomen and face fawn-cnlnnr.
Expanse of wings : i 35 mm. ; ? 3;) — 4(J mm.
C (?cJ, 2 ? ?.
186. Chogada compectiuata s])ec. nov.
Foreimnq : white, with slight grey speckling, chiefly along costa ; lines black,
fine; first from one-fourth of costa to one-fifth of inner margin, rounded in cell,
slightly indented on snbmediau fold, preceded by a macular grey shade ; outer
line at two-thirds, Innnlate-dentate, nniformly distinct, followed by a shade formed
of grey Innulcs, fmm which it is separated by a white line ; median line fine,
strongly projecting outwards in cell, and less strongly on submedian fold, blacker
on median and submedian veins ; cell-s])ot formed of raised white scales within the
projection ; praesubmarginal shade grey, lunnlar, interrupted between veins 3, 4, and
6, 7, the lunules blackened on each side of vein 5 ; the shade following snbmarginal
line much interrupted, black from 0 to below 5, and farming a (piadrate deep black
spot on margin between G and 7, running out into the white fringe ; marginal
spots black.
Ilimhi'iiiy : like forowing, but without inner line ; a straight line from the
white cell-spot to the submedian fold ; no marginal black blotch.
Underside white, greyer towards apex of fore wing, where the outer lines and
snbapical blotch are marked ; costa of forewing striated with black.
Head, thorax, and abdomen greyish white ; face with a black bar across
middle; vertex sometimes whitish; fringe on basal segment of abdomen white;
dorsal segments with pairs of black spots ; the segments following hasal flecked
with red.
Expanse of wings : 3U — 34 mm.
5 cJcJ.S ? ?.
The ? ? are slightly greyer than the cJ 6. The pectinations of the antennae
of the i are interlaced.
187. Chogada semidiscata spec. nov.
Forewing : white ; the costa shortly and neatly marked with brown striae ; a
brown-black sjjot on costa close to base ; the four lines starting also from brown-
black costal spots at one-fourth, two-fifths, two-thirds, and five-sixths respectively;
first line black, slender, outcurved above median to one-fifth of inner margin,
preceded by a slight rufous cloud ; outer line black, sinuous, ending at middle of
inner margin, projecting on vein 5, insinuate below it at the origin of veins 3
and 4, and followed by a rufous shade, which is deep brown between 3 and 4; median
line rufous, excurved, and projecting on each side of vein -J, below median running
close to outer line; submarginal line waved, white, preceded and followed by shades
of brown striae, mixed with black scales below costa, beyond cell, and below vein 3,
( 142 ;
and between '.i ami 4 swollen info a white lilotch ; marginal spots distinct, black ;
fringe jiale brown, clioqnered with wiiitish ; no distinct cell-spot.
Ilimlu'ing : with a brown-edged crescentic cell-mark ; a black antemedian and
post median line ; the rest as in forewing.
Underside dnll wliitisl], tlie markings showing throngli and discolouring tlie
forewing.
Palpi dark brown, the third joint oclireons ; face ochreons, brown above ;
vertex, thorax, and abdomen white speckled with brown ; tips of shoniders
brown; dorsal segments with jiairs of brown dots.
Exjianse of wings : 4ii mm.
1 S.
ISS. Myrioblephara adumbrata sjiec. nov.
Fonn-hiij : i}ale grey, dusted with fawn-colonr; basal third jialcr, but the
costa grey-brown ; inner line brown, with a slight angle outwards on submedian
fold, and inwards on submedian vein; outer line brown, at two-thirds, faintly
Innnlate-dentate, incurved across submedian interval; followed by a fainter, but
more strongly dentate line, lieyond which the space between 3 and 4 is tilled with
deep black and dark grey scales; from before middle nf costa to before niiddle of
inner margin a black-brown thick median line runs, followed by a broad brown
shade, incurved across submedian interval, where it all but touches the angle of
inner line; space between 3 and 4 at base filled with whitish scales; snbmarginal
line waved, pale, the lunnles filled in with lirown ; marginal area darker fawn-colonr,
blolehed with brown beyond cell and towards anal angle ; fringe pale fawn, beyond
some slight lilackish marginal spots.
HhuhriiKj : the same, without inner line, or black blotch beyond outer line ;
base whitish.
Underside ])aler, with all the lines grey-brown.
Face and palpi dark brown ; vertex and thorax dark grey ; abdomen jialer, like
liindwing, with a double brown band at middle; legs pale fawn; foretarsi black,
with i)ale joints.
Ex|iause of wings : 32 mm.
1 ?.
1^''. Myrioblephara complicata spec. nov.
Fornvinij : dark olive, with the veins fulvous ; the lines formed of contiguous
while irregular blotches; basal j)atch olive, the base of inner margin ])ale, and
traces of a pale line running round the foveal jmstnle ; edged from one-third of
costa by three white blotches between veins, that on inner margin nmniiig in
basewards ; central sjiace olive-green, with a small round white spot marking
discocellnlar, edged from three-fifths of costa by a curved white blotch to vein 4,
and a spot below beyond the rise of 3 and 4 ; a broad white band from apex
bii'urcating below vein (i and reaching inner margin as two series of irregular
lunnles, sei)arated by a widening dark olive S2)ace ; vein 4 with a white blotch to
margin ; the ends of veins 1, 2, 3 broadly rusty fidvous between the black marginal
lunules ; fringe dark grey, with white chetjuering between the veins, the apex
broadly white ; the costa is dark olive flecked with jiule, all the white markings
commencing from the subcostal vein.
lliii(/irii<y : yellowish bchreous, with a greenish tinge, with a cell-sjiot and
( 143 )
three waved lines bejond middle, jjrcrnisli ,£:i'ey ; fringe yellow oclireous, chequered
with olive beyond veins 2 and 3.
Underside yellowish oclireous, in tlie forewiiig with tlie inarl<iiigs of njjjier-
side showing throngli ; in hindwiiig with cell-sjiof, outer line, and two interrnpted
snbniarginal lines, and a lot of striae at apex dull olive-green.
Head and thorax mixed, darli olive-green and oehreous ; abdomen oi'lireons
with brownisli dorsal belts; legs niollli'd dnrk olive and oehreous.
Expanse of wings : 35 mm.
1 <?.
10(>. Myrioblephara fulvivena spee. nov. and ab. defulvata nov.
Fomriiig: white, speekled with dark brown atoms; all tlie veins thickly
fulvons ; a macular basal band; a double macular band at one-fourth, of which
the inner arm is browner and the outer marked by black spots ; a double outer
band at two-thirds, of which the inner arm is black-spotted on veins and the outer
brownish; median line fine, obliijiie outwards from a dark costal spot above the
blackish cell-spot, then reimrved, joining inner Iiand in a large blotch on inner
margin; submarginal line white preceded by blackish blotches at costa, beyond
cell, and on inner margin; marginal spots Mack; fringe with basal half i'ulvous,
outer half whitish, with Idack motllings lieyond veins.
Ilindwiii;/ : white, with grey curved antemodiau, postmedian, and subniargiiml
lines, ending in thick oblique black streaks on inner margin ; the postmedian dotted
on veins and followed by a grey shade; the submarginal with the lunules filled up
with dark grey ; marginal area and fringe fulvous ; cell-spot grey.
Underside fulvous white, paler in hindwing, whrt'e the markings are prominent;
costa of forewing fulvous with black speckling.
Head, thorax, and abdomen fulvons, the thorax with olive and pale scales
intermixed ; tarsi fuscous with the joints jiale.
In the aberration <l('fi(lciita the veins are without any fulvous tint and the
whole wing much less gay ; the fringes are also dark and light grey; the underside
grey, without any fulvous tint.
Expanse of wings : 30 mm.; the aberration only 2S mm.
2 cJJ, 2 ? ?, of the type form ; 2 ? ? only of the aberration.
I'.il. Myrioblephara iuquinata spec. nov.
Forewiny : dull white, speckled with dark grey ; base of wing and costa at
base blackish-grey ; a black dash on median vein at base ; inner line outeurved
from costa at one-third, closely followed hy a somewhat waved median line ;
outer line marked by black wedge-shaped marks on veins pointing basewards,
angled at vein 6 and below 3 ; the space between inner and outer lines blacker,
especially along costa ; the inner line is preceded and the outer followed by a
thick fulvous brown shade ; submarginal line wavy, white, between blackish shades;
the marginal area pale at anal angle and again between 3 and 4 ; marginal spots
black ; fringe mottled black and pale grey.
Ilimkmny : with extreme base black, a double antemedian line and black
cell-spot; the rest as in forewing ; both wings, except along marginal area, have a
slightly rufous tint.
Underside dull whitish, grey in the forewing; the Hues rather darker grey;
marginal border grey.
( 1-44 ;
Ili'iiil and thorax blackish ; colhiv wliitish ; base aiul anal hall" of abdomen
blackish, third and fourth segments whiter.
Expanse of wings : 3U mm.
1 ¥.
192. Myrioblepliara lucidata s|icc. nov.
Fommx/ : white, with slight (dive and fiiscons si)ec,kling ; the shading olive,
but more linear and condensed, the linos blackish, less waved, than in the
allied species ; basal line almost touching base; inner line nearly straight, vertical,
jirecedcd by a narrow olive band ; median shade olive, angled beyond the black
cell-spot, to two-fifths of inner margin, followed by a second shade; outer
line at two-thirds, slightly bent ont beyond cell and inbcnt across snbmedian
interval, accompanied by a well-dc(ined olive band; snbmarginal line broadly
wliitc, waved, ])receded and followed by distinct olive shading except between 3
and 4, where the shading is grey, and the line itself flattened inwards; marginal
lunnles black; fringe with the basal half brownish grey, the apical half whitish.
IliiuhciiH) : white, withont speckling ; a black antemedian and postmedian
line less sinuous than in other species, each followed by a pale olive-brown shade ;
snbmarginal line white, waved, between two olive-brown shades.
Underside fawn-grey, darker in forewing, with all markings clear, but the
lines not waved nor sinut)us.
Head and ]ialpi dark olive; thorax wliitish dusted with olive; abdomen
bioken off.
Expanse of wings : 31 mm.
] ?.
I'.t3. Myrioblepliara olivacea spec nov. and ub. albiviridis nov.
Foreu'iiiy : white speckled with dark atoms; the shadings olive; the lines
black; costa very finely dotted black and white ; lines and shades almost precisely
as in M.fidrirdid, but the shading olive instead of fulvons ; the whole wing paler ;
fringe white spotted with grey ; cell-mark black, but small.
llimltciiiy : with more than the costal half whitish ; the lines as m fiihieciia,
but less distinct.
Underside dirty whitish with a faint fawn-coloured fringe, especially in fore-
wing ; markings all grey, but finer, less numerous than in fuh'i re iHi.
Head, thorax, and alidomen whitish-grey speckled with olive.
Expanse of wings : S 30 mm. ; ? 32 — 35 mm.
1 (?,2 ? ?.
This apjiears to be really a distinct species, thougli in markings so much like
fulvivena.
The aberration (ilbiriridis has the gnmnil-colour greenish white, with the
olive shading developed, forming a broad shade occupying the second fourth of
inner margin and nearly reaching costa ; the underside greenish white.
I'.i-t. Myrioblephara submarginata sjiec. nov.
Forcwiny : whitish, thickly dusted with grey; all the lines and shades dark
grey ; basal lini; close to base, with a spot at base ; inner line at one-fourth, curved
to snbmedian fold, then iidient, preceded by a similar shade ; outer line at
two-thinls, lunnlate-deiilate, uniformly curved, and insinuate on submedian fold,
( 145 )
more strong!}' marked on veius, followed liy a thick dark grey shade ; median
shade well marked, outcurved round the black cell-spot, a little before middle;
snbmarginal line pale, wavy, preceded and followed by dark grey shading, which
]iales off at anal angle ; the marginal area pale grey between 3 and 4, preceded
by a darker blotch ; marginal sjiots !)lack ; fringe white, with dark grey chequering.
llimliciinj : with antemedian line thick, straight ; the rest as in forewing.
Underside wliilisli grey, the base greyer; lines and cell-spots black and
distinct ; both wings with smoky-black marginal band.
Head, thorax, and abdomen like wings.
Expanse of wings : \l'J mm.
1 c?, 1 ?.
Paralcidia geu. uov.
Forcwim/ : elongate; costa shouldered at base, slightly curved before apex,
which is rectangular ; hindmargin vertical above middle, obliijue below.
Ilindwinq : with upper half of hindmargin vertical, slightly indented beyond
cell, lower half oblique; costa straight.
Antennae lamellate, flattened, withont pubescence ; forehead protuberant ;
palpi obliquely porrect upwards, witli a}iprcssed scales, second segment long,
terminal depressed ; tongue and frenulum present ; abdomen of J elongate, with
large anal tufts.
Neuration : forewing, cell longer than half of wing ; discocellular vertical ;
(irst median uervule at five-eighths, second at seven-eighths ; radials normal ;
subcostal vein deflexed at extremity; 7,8, 9 stalked; li), 11 separate; hindwing,
costal and subcostal shortly approximated near base ; 7 and 3 from before angles
of cell.
Scaling smooth and tine.
Type : I'aralcidia crrabunda spec. nov.
I'Jo. Paralcidia errabimda spec. nov.
Forcwuuj : piukish ochreous, but the basal four-fifths of wing is suffused with
dark olive-fuscous, as far as the Innulate-dentate submarginal line, which rises at
three-fourths of the costa and runs obli(jnely outwards to vein 6, cutting off a
small olive costal blotch beyond ; the pale ochreous ground-colour of the marginal
fifth is striated with olive, and shows the veins across it pink ; along vein 4 this
pale ground projects inwards to cell, where it is joined by an oblique pale streak
from before middle of costa, which is incurved from vein 5 aud runs vertically
to the submedian fold ; the cell-spot ochreous ; along inner margin are some fine
ochreous striae, and au ochreous spot at one-fourth suggests the termination of
an inner line; costa with fine ochreous points; the lunules of the submarginal
line contain externally some olive-green spots, from which streaks of green scales
run to the dark olive marginal line ; fringe pinkish ochreous, mottled with olive
beyond veins.
liindwiny : pinkish ochreous, with the dark markings of underside showing
through ; fringe ochreous.
Underside of forewing pinkish ochreous, with the dark olive area showiug
through ; costa striated with olive : hindwing tinged and striated with olive as far
as a thick deep olive crenulated outer line ; ccll-sjiot dark olive ; abdominal margin
below fold ochreous.
10
( 140 )
Head, shoulders, patagia, and a broad lah'ral streak <iii abdumeu dark olive ;
tborax and abdomea above and below, ami tips of palpi pinkish ochreous ; legs
ochrcoiis dajijded with <;reeu.
E.N]iaiise of wings : 27 mm.
1 cJ,4 ? ?.
196. Paralcidia marginata spec. uov.
Forewing : olive-green as far as snbmarginal line, which is mnch nearer the
margin than in errahtimla ; the pale jjinkish ochreous area therefore narrower; the
line runs obliquely ont from three-fourths of costa and is blnntly bent at vein 6,
slightly indented on both folds ; the marginal area sprinkled with green scales,
the veins pink ; at anal angle the ochreous tint runs in along iuuer margin to
near base; fringe pinkish ochreous, with slight green chequering ; discal spot dark,
obscure.
Jlitidicing : pinkish ochreous, appearing grey to near margin from tlie dark
area beneath.
Underside of both wings dull olive, blurred in forewing, deeper in liiiulwing ;
cell-spots dark ; marginal area and fringe pinkish ochreous with slight green
speckling.
Head, shoulders, patagia, abdomen beneath, the segmental divisions, and top of
anal segment dark olive; thorax and abdomen ochreous.
Expanse of wings : c?, 27 mm. ; ? , 32 mm.
3 c?cJ, 1 ?.
197. Paralcidia rufivenata spec. nov.
Forewing: green thickly speckled with pink and grey scales; the veins deep
pink : costa dotted, pink and green ; lines white, edged with deep green ; first
from beyond one-third of costa, oblique outward and curved round at end to median
above the rise of vein 2, continued between median and snbmedian as a vertical
white lunule much nearer base, and below submedian by a white oblique streak
Dearer still ; followed by deep green shades between the veins ; in the mottled
basal area are a dark green sjiot in cell and another oblicjuely below it near base;
outer line from two-thirds, obliquely curved outwards to 0, insinuate beyond cell
and in submedian interval, outcurved between, below submedian sliarpiy oblique
basewards, edged internally by dark green Innnles ; in the marginal area are three
green blotches ; one on costa, broad, reaching to vein 6 ; one at anal angle reaching
vein 2, narrow; the third towards margin between them, separated by hoary grey
scaling ; dark marginal lunules between veins ; fringe pink, chequered with green.
llindwing : whitish ochreous, becoming grey towards hindmargin ; the veins
at extremity and the fringe pink ; a grey cell-spot and waved postmedian line.
Underside of forewing blurred cinereous-green to outer line, which is dark ;
costa pink striated with green ; the veins pink ; marginal area paler than basal :
hindwing sjieckled with green up to the tiiiek dark green outer line, which is
lunulale-dentate; cell-sjiot dark; abdominal margin paler, more broadly in c?.
Face and jialpi deep pink ; palpi externally dark green ; vertex, shoulders, and
patagia deep green ; thorax and abdomen pinkish grey ; abdomen below and legs
pink ; terminal curl of hindtibia shining olive ; antennae piuk.
Expanse of wings : 30 mm.
2 JJ,2 ?¥.
( 14?)
Tlie (? of tlie .sjiecies, wliile in geueral structure it agrees with tyi)ical Paial-
cidia, differs iu three particulars : the costa of hiudwing is convex and roughly
haired; the hindtibiae ave much swollen towards extremity and fringed with hair,
and at the end bear a curled tuft of shining hairs ; and thirdly the antepenultimate
segment of the abdomen beneath bears an abrupt jirojeetion, and is clothed with
tufts of rough hair.
198. Paralcis coerulescens spec. nov.
Forcwinq : mixed fulvous brown and purplish grey, irrorsited with bluish
grey scales ; the veins fulvous, the median broadly so ; lines velvety black ; first
at one-fourth, angled outwards in cell and on submedian fold, and swollen on
costa, median vein, and inner margin ; outer line crcuulate, outcurved, at two-
thirds, thickened on veins ; median line similar, nearer outer than inner line ;
cell-spot ronnd and black, with a linear blue-grey centre ; the outer line is edged
with bluish grey scales, and followed from costa to middle by a black shade ;
submarginal line blue-grey, preceded by black blotches between the veins ;
interrupted between 6 and 7 and 3 and 4 ; marginal black spots between veins ;
fringe purplish grey.
Ilitidwing : dark slaty grey ; the cell-spot, postmedian, and submarginal line
showing darker ; veins towards margin fulvous ; fringe slaty gray with a fulvous
sheen.
Underside dark slaty cinereous, the fringe and cell-spots deeper ; ajjex of
forcwiug pale.
Head and thorax purplish ; abdomen shining dark cinereous.
Expanse of wings : 38 mm.
109. Paralcis curvilinea spec. nov.
ForctciiHj : dark reddish brown, overlaid with a purplish shade in the central
area ; first line at one-fifth, black, angled outwards in cell, then oblique inwards,
with a paler inner edge ; outer line from two-thirds of costa to four-fifths of
inner margin, outcurved from below costa to submedian fold, where it is indented,
black with a pale outer edge; a black median line, waved, and projecting just
below median vein, followed by the black cell-spot ; submarginal line very obscure,
grey with a black edging, preceded by a black blotch on inner margin beyond
outer line ; the apex bluish grey, with some black scaling along hindmargiu
beneath it ; marginal spots black ; fringe brown.
lUndwing : dirty whitish ochreous, speckled with grey, with a dark cell-siwt,
and a strongly curved grey postmedian line ; fringe pale.
Underside of both wings like upi)ersidc of hindwing, the forewing blurred.
Head olive-ochreous ; base of shoulders, patagia, and a lateral line on thorax
purplish black ; tips of shoulders, thorax, metathorax, and basal segments of
abdomen pale ochreous ; abdomen greyish ochreous with dull brown-grey dorsal-
blotches ; tarsi mottled brown and ochreous.
Expanse of wings : 34 mm.
2 ??.
At the base of forewing, in the usual place of the fovea, there is visible a
small pustule.
( 148 )
2UU. Paralcis deformis spec. nov.
Forcwiiiij : pale wood-brown, smeared with greyish fuscons as far as outer
line; first line at oue-fburtb, black and interrupted, bent in cell; outer line from
twii-tbirds of costa to tbree-fonrtlis of inner margin, bhxclcisb, outenrvcd from 0 to
submedian fold ; cell-spot large and black ; traces of a median shade, plainest on
costa and above inner margin ; snbmargiual line pale, waved ; the lunnles filled in
with blackish ; marginal area tinged with deeper brown ; some black marginal
dots ; fringe brown.
llii)ihciii</ : ]iale brown, with grey speckling; a black cell-spot and grey
postmediau line.
Underside darker ; both wings dark grey to outer line ; cell-spots black.
Head and thorax brown varied with black ; abdomen like hiudvviiigs.
Expanse of wings : 30 mm.
1 6.
The single example is considerably worn.
2i)l. Paralcis discata spec, nov., and ab. alterata nov., and ab. albiclausa nov.
Forcwiiig : dull olive-lirown, generally densely covered with bluish grey and
olive scaling, so that the dark ground-colour is lost ; a large oval Idackish
cell-spot ; a dark blotch at extreme base ; lines darker, but obscure ; first at
one-fourth, bent in cell, then vertical ; outer from three-fifths of costa to two-
thirds of inner margin, obli(pie outwards to vein 5, then inwards to submedian
fold, then again outwards ; submarginal line obscure, parallel to outer line, preceded
by darker scaling in the lunules and with the whole marginal area beyond dark
brown, except between 3 and 4, where the bine-grey mottling rnns through to
margin ; marginal lunules black, edged outwardly with {)alc ; fringe brownish
grey ; median shade always obscure.
Ilindwing : dull greyish white, speckled with olive-grey ; a blackish cell-spot,
a sinuous dark outer line, and traces on inner margin only of antemedian and
submarginal lines ; fringe concolorous, beyond a dark marginal festoon.
Underside like npperside of hindwing ; cell-sjiots black ; the lines indicated
on forewing ; costa of forewing distinctly striated with dark.
Head and thorax like forewing, abdomen like hindwing.
Expanse of wings : 30 — 3.j mm.
A long series, all ? ? .
In the aberration alterata the broad central and narrow marginal areas are of
the brown ground-colour, unspeckled by white ; but the intervals preceding and
Ibllowing the fascia become altogether bluish grey or whitish, and the submarginal
line shows more broadly white ; in these cases the dark central area is broader
than in typical dittcata, the brown suffusion not only including the space between
inner and outer lines, but also extending to the dark shades which precede the
inner and follow the outer lines, and which in the typical form are more or less
obsolete ; median shade distinct and black.
Expanse of wings : 28 — 36 mm.
4 ? ¥.
The aberration (dlndausa is the extreme form of ab. a/tt'nita, in whicli the
brown central fascia is bordered on each side by broad white bands.
2 ??.
( 140 )
2112. Paralcis Mvisecta spec. nov.
cJ. Foreicing : olive-grej' sprinkled with blackish aud reddish scales in basal
two-thirds, beyond dark olive-fuscons ; the lines black ; first from one-fifth of
costa to one-fourth of inner margin, obliqne outwards above and angled botli above
and below median, preceded by a distinct wliitish line following patches of dark
olive between the veins ; traces of inner dark blotches ; outer line at two-thirds,
distinctly lunulate-dentate above middle, angled bluntly on vein 4 and thence
simply waved, followed by a whitish line ; cell-mark vertically oblong, large and
l)lack, with a pale linear centre ; above it a medinn line, thick aud black, rises,
running outwards obli(£uely to 6, then luuulate-deutate, parallel to outer line to
vein 3, thence incnrved to middle of inner margin, the space between it and outer
line above middle whitish ; a broad curved cream-coloured band from apex to
inner margin before anal angle, meeting an arm from three-fourths of costa, both
more or less dusted with rufous-olive scales ; submarginal line fine, white, sinuous,
from costa before apex, approaching hindmargin above vein 4, then toothed inwards
on veins and broadly lunate between them ; median vein fulvous throughout,
this tint broadened along vein 4 nearly across the interval between it and 'i ; the
other veins fulvous in places, ending as dull fulvous lunules on hindmargin
alternating with the black marginal lunules ; fringe yellowish ochreous, Ijroadly
mottled with olive-fuscous beyond veins, and with a dark dividing line.
In the ? the discal spot is much larger, velvety-black, scarcely showing the
pale centre, aud followed by a quadrate white blotch ; the curved white band from
ajiex is absent, being rejjresented by an obscure band of olive ; the apical space
between submarginal line aud the oblique outer edge of the band is white, and the
pale fulvous marginal area runs in as a rounded pale-scaled tooth between veins 3
and 4 ; fringe darker.
Ilindwing : cinereous, darker along hindmargin ; with a thick dark postmedian
line, edged with white on inner margin, traces of a snljmarginal line, and a dark
cell-spot; fringe dark cinereous beyond black marginal lunnles.
Underside dark slaty cinereous with an olive tinge ; apex of forewing aud
inner margin of hindwiug paler ; in the ? blackish, with large diffuse black
cell-spots followed by white patches, that in forewing large ; apex of forewing
white.
Head and thorax dark olive and fuscous ; abdomen cinereous ; jialpi externally,
pectus, and forelegs black.
Expanse of wings : (?, 40 mm. ; ? , 44 mm.
5 (?(?, 6 ? ?.
203. Paralcis junctilinea spec. nov. and al). punctata nov.
Foi-ciciiig : fawn-drab, slightly dusted with darker ; lines fuscous, fine ; first at
one-fourth, nearly straight ; outer from three-fourths of costa to middle of inner
margin, parallel to hindmargin, but slightly bent in on submedian fold ; cell-spot
blackish with grey centre, followed by a brown median line, approximated to outer
Hue on inner margin ; a blackish submarginal line from costa to vein 5, forming
slight black lunules between veins, edged with white from G to 4, the uppermost
largest, aud followed by a slight brown cloud to apex ; fringe concolorous ; a row
of black dots between veins just before margin ; a brown horizontal streak nuuiing
from cell to submarginal line above vein 4,
( 150)
Hindtrhifi : with the two brown lines wider apart at eosta, the outer more
sinnons, approximating on inner margin ; cell-spot small, black.
Underside whitj'-brown with the lines faiut ; ecll-s])iits black ; tlirco or four
black spots before apex.
Head, thorax, and abdomen concolnrons ; face darker ; anal segment paler.
In the aberration punctata the gronnd-colonr is darker ; the lines are repre-
sented b}" black spots on veins, the median shade Ijeing a ditl'iise cloud ; the
submarginal line is more distinct across wing, with the white spots larger, coalescing
above ; in the 9 develoj)ing into a whitish apical blotcli ; the cell-spots are larger
in the ?.
There is no trace of the dark horizontal line of tlie type-form.
Expanse of wings : S, 30 ram. ; ? , 34 mm.
2 (?c? of the type-form ; 1 (?, 1 ? of the aberration.
204. Paralcis latimedia spec. nov.
Foretcing : sepia-brown with dark striae ; costa with black striations ; a
blackish blotch at base ; first line thick, velvety-black, projecting outwards above
and below median, from before one-fonrth of costa to one-fonrth of inner margin ;
the area beyond it sntfnsed with brownish fnscons deepening to black at the outer
edge, which runs from two-thirds of costa to four-fifths of inner margin, irregularly
waved and forming a bitid projection on veins 3 and 4, edged first by a pale and
then a dark brown line ; in the outer third of fascia a dark lino runs parallel to
outer edge beyond a blackish vertical linear cell-sjiot ; submarginal line obscured
by cloudy brown marginal shading, whieh is interrupted between 3 and 4 ; a fine
marginal festoon, connecting the black spots ; fringe brown.
IIi)i(lwi))g : dirty whitish, dnsted with dull grey; a dark grey waved post-
median line from two-thirds of costa to inner margin above anal angle, with traces
of two other grey lines from the angle ; a dark Innulate marginal line ; fringe pale,
with a reddish tinge.
Underside of both wings like uj)perside of hindwing ; basal half of forewing,
except along inner margin, dull cinereous.
Head, thorax, and abdomen brownish grey ; paljii and slmuidcrs blaik ; jiatagia
and dorsal blotches of abdomen brown-black ; fiice brown.
Expanse of wings : 44 mm.
2 ? V.
205. Paralcis pallidimargo spec. nov.
Fornripfi : pinkish ochreous flushed with pale brownish in the S, jialer
ochreous flushed with olive-grey in the ? ; a sliglit dark brown patch at base ;
a donble black basal line, the interval filled in with lirown, irregularly bracket-
sliaped ; outer lino black, lunulatc-dentatc, at two-thirds, indented below costa,
running outwards to vein 5, forming a blunt projection between 5 and 4, then
inwards, followed by a thick dark brown shade ; median line from two-fifths of
costa, sinuate outwards to cell, tiien incurved, and again below middle running
outwards ; space between median and outer lines partially filled with brownish
black and with a patch of hoary grey scales on discocellular ; submarginal line
waved, preceded on costa by a double brown Innulate blotch, by two smaller brown
Innules beyond cell, and by a jiniminent black-brown sjmt on inner margin ;
marginal area black-brown from 4 to near apex, below 4 brown in the c?> olive
( IM )
or olive-ochrcous in the ? ; marginal sjiots large, black ; fringe dark brown with
ochreous flecks in the S ; ochreons with brown flecks in ? .
Ilindwing : ochreous 'at base, becoming grey-speckled and darker to the dark
grey snbmarginal line, containing a grey cell-spot and fine waved postmedian line ;
marginal area ochreons with slight speckling, and some dark marginal spots ; fringe
ochreous.
Underside of forewiug dull dark brown to turee-fonrths, the marginal area
ochreons, and some patches of ochreous along costa ; hiudwing as above, with a
dark cell-spot.
Abdomen ochreous speckled with brown ; face and palpi black-brown ; vei-tex
and thorax brown mixed with ochreous ; antennae ochreous spotted with brown.
Expanse of wiags : S 35 mm. ; ? 35 — 40 mm.
1 c?, 3 ??.
The two larger ? ? are much paler in tint throughout than the other ? and
the type S, but the markings are identical.
206. Paralcis ruptilinea spec. nov.
Forewincj : dark brown tinged with purplish grey; costa spotted with
yellowish ; lines blackish brown ; basal from one-fourth of costa to before one-
third of inner margin strongly outcurved above median ; outer line at three-fifths,
angled outwards on vein 4, concave above, marked chiefly by black vein-spots ;
a black cell-spot, over which a cloudy median line runs, approaching outer line
on inner margin ; snbmarginal line sinuous, indistinct, but marked by the deeper
tinge beyond it ; an oblique angular line from apex to snbmarginal line at vein (i,
edged outwardly with deep black ; the included costal triangle sometimes white,
sometimes like ground-colour.
Iliiuhmig : slaty bluish, ferruginous at anal angle, above which there are
traces of lines.
Underside dull slate-colonr, the costa of forewing yellow-spotted ; cell-spots
black ; apical triangle concolorous with upperside.
Head and thorax like forewing ; abdomen greyer ; antennae whitish.
Expanse of wings : 35—40 mm.
2 ? ?.
Most probably with other variations.
207. Paralcis venusta spec. nov.
S. Forewinr/ : pearl-grey, bluish grey at base and along costa, cream-colour
before snbmarginal line ; costa thickly striated with blade ; first line at one-fourth,
bracket-shaped, vertical between the folds and retracted at each end, with a grey
shade before it tinged with rosy brown below median ; outer line from two-thirds
of costa, sinuous, being incurved below middle and again outcurved before inner
margin, black, sharply dentate on veins, followed by a rosy brown shade ; median
line also black and dentate, more outcurved from costa, closely approximated
to outer line, which it touches forming a black blotch, on submediau fold ; a
large brown difl'use patch on discocellular lies within its upper curve; snbmarginal
line waved, interrupted ; from costa to vein 7 edged inwardly with black, and
distinct, passing through the grey costal shading, more or loss obliterated below
by the cream-coloured area, which between 3 and 4 j)rojects to hindmargin, inter-
( 152 )
rnjiting the brownish grey margin.al area; marginal spots largo niul black, joinod
by black streaks between 7 and 4 to the Innnles of submarginal line, and to each
other by a fine marginal line ; fringe mottled dark and light grey.
Ilincln-ing : blnish grey, with a dark ba.<al mark, an indistinct grey antemcdiau
line and small dark ocellns on discocellnlar ; the rest as m forewing. but the jialo
snbmarginal space not so wide and greyer, not interriiptiug the marginal area.
Underside dark slaty fnscons with a greenish tinge ; lines hardly visible ;
fringes and apex of forewing pale grey.
Falpi black; head, thorax, and abdomen dark grey ; basal segment of abdomen
black, second whitish, third with a pair of black spots ; alxlomen beneath and legs
slaty fnscons ; foretarsi blackish, with the joints pale.
?. Snlfascd throughont with jmrplish grey and brown, and thickly black-
speckled, especially along the veins, so that the whole wing surface assumes a
mottled appearance, the underside becoming black.
Expanse of wings : 4U mm.
1 cJ, 2 ? ?.
Subfamily SELIDOSEMINAE.
208. Tolmera albibasalis AVarr.
When this species was described, S'oe. Zool. x. p. 404 (1903), only the S
was known. Now along with 7 cJ(? a single ? has been sent; iu this sex of
this species the antennae are bipectinate, the pectinations thicker and shorter than
iu the (?.
Both wings are jjaler brown, and the outer line is not so near the hindmargiu
as in the c?.
209. Tolmera marcescens spec. nov.
S. Foreicimj: white, washed with pale brown and speckled with black ; the
veins pale ; lines white ; first at one-fifth, acutely angled below subcostal vein, and
oblique inwards, edged outwardly (and iu S inwardly also) with dark ; outer liue
from three-fifths of costa, below which it is incurved slightly, parallel to inner line
as far as submedian fold, then outcurved to two-thirds of inner margin, inwardly
edged with dark : the space between the lines with a dark central shade ; sub-
marginal line waved, the Innnles filled up with dark ; a dark oblique blotch beyond
it to ai)ex from vein 6, the apex itself jialer; dark marginal Innnles; fringe pale
chequered with dark brown.
In the only ? seen, and of which the ground-colour is paler, being more mixed
with white, the second line starts from nearer middle, forms two concavities inwards,
and two outward teeth on the median and submedian veins.
Iliiuhcing : pale ochreous, slightly speckled ; cell-spot and curved outer line
grey ; marginal spots dark.
Underside of forewing pale ochreous, yellower towards costa, which is spotted
with brown ; outer and submarginal lines brown towards costa ; marginal Innnles
and chequering of fringe black-brown: hindwing yellowish, paler towards abdominal
margin, speckled with brown, and with brown outer line and cell-spot.
Palpi ochreous, brown externally ; face pale ochreous with two brown spots ;
vertex, shoulders, and ])atagia jjale brown, black-speckled; abdomen pale ochreous
without any speckling ; legs pale brown, black-mottled ; ? with vertex white,
( 153 )
Expanse of wings : S 35 mm. ; ? 38 mm.
4 d'c?, 1 ?. ^
The ? differs from the ? of 7\ albihusalis, the type species, in having the
antennae not bipeetinate, bnt simi)Ie, ail the segments sharply angulate, witli a
bristle from each angulation.
210. Trochistis albivertex .spec. nov.
Forewiny : fawn-colour speckled with black; costa striated with black; a
blacki.sh dot at base of submedian vein ; three brownish outwardly oblique lines
from costa at one-fifth, two-fifths, and two-thirds to one-fifth, one-half, and four-fifths
of inner margin respectively ; the inner line waved, incomplete ; the second
followed by a small black cell-spot ; tlie third a little waved below middle, followed
by a pair of lilack spots, one on each side of vein 3 ; a cloud of black striae at apex,
in which the commencement of a submarginal line is visible ; marginal dots black ;
fringe paler.
lUivhviMj : with dark central line ; the outer lino not visible, though the two
black spots are present, but duller, beyond it.
Underside whity-brown, black-speckled ; marginal dots black.
Head, thorax, and abdomen like wings ; palpi with terminal segment black,
its tip white ; vertex snow-white ; antennae annulated black and brown ; basal
segment white.
Expanse of wings : 37 mm.
1 ?. ""
211. Trochistis asinina spec. nov.
Forewitig : dull fuscous, s^jcckled with dark ; the lines darker ; the costa with
a few very fine white markings, distinct only in the S ; three outwardly inclined
dark lines at one-fifth, two-fifths, and two-tliirds, and a broader but more diti'nse
submarginal line ; in the cJ these are partially accompanied by white scaling, and
the submarginal line is followed by a patch of white scales from costa to vein 4 ;
the marginal dots white ; in the ? all these are absent, the marginal dots being
minute; cell-spot black.
Iliiidwing : without inner line; cell-spot white; in the S the jiostmedian is
accompanied by bright white scales as well as the submarginal and marginal lines ;
fringe with pale tips.
Underside of forewiug dull olive-fuscous dusted with pale, the costa and
hindmargin darker ; cell-spot and outer line marked : hindwing whitish dnsted
with dark, the outer line and hindmargin only fuscous ; all the tints darker
in ¥ than in cJ.
Head, thorax, and abdomen like wings ; tips of palpi and anal tuft of c? pale
ochreous ; antennae black, the shaft annulated ; pectinations of S plumose.
Expanse of wings : 35 mm.
1 c?, 5 ?¥.
212. Trochistis rufoliva spec nov.
Forcwinij : deep red-brown, with an oval snow-white cell-spot; costa in S
blotched with olive-green, in ¥ with paler green, the green sj)aees with brown
striations ; three ouLvvurdly inclined thick red-brown lines at one-fifth, two-fifths,
( 154 )
and two-thirds, most distinct in the V ; from the groon intervals lines and bands of
pale green scales run across the wing between and edging the lines, plain only in
the ?,the middle space being broad and pale throughout; a submarginal row of
green or gre3--greeu spots on veins, and a marginal series of smaller green spots
round the black spots, these last showing also in the S ; fringe red-brown flecked
with greenish white.
Hindwing : with the two outer series only jilain, the greenish white speckling
being more scattered ; cell-spot small, white.
Underside whitish ochreous, greyer in forewing, grey-speckled; a grey
snbmarginal shade, slight in forewing ; white spot of forewing showing through ;
the coloration deeper in the ? .
Head, thorax, and abdomen red-brown ; palpi tipped with pale ; abdomen
dusted with greenish white ; anal tuft of S ochreous, red beneath ; antennae black
and pale ; pectinations in S short and stifi'.
Expanse of wings : 37 mm.
4 cJc?, 4 ? ?.
213. Trochistis vulpina spec. nov.
Forewing: rufous brown, peppered with blue-grey scales in the J, and with
black spots and striae in the ? ; cosfa black-brown ; lines dark brown, all obliqne
outwards, from costa at one-fifth, two-fifths, and two-thirds to one-fourth, one-half,
and three-fourths of inner margin respectively, the outer line twice bent; in the ?
all the lines become black at inner margin ; submarginal line in ? preceded and
fcilldwed by a black shade from costa to vein 4, and with one on inner margin before
it ; in c? not marked ; marginal spots blue-grey in S, black in ? ; fringe con-
colorous ; cell-spot minute, black.
Hindwing : similar ; the markings much clearer in ¥ ; cell-spot black in a
grey ring.
Underside whitish with a pink tinge and grey-speckled ; apex of forewing
rufous ; fringe rufous ; marginal and cell-dots black ; ? with the markings stronger,
the fringe deep red.
Head and palpi deep brown, tips of palpi ochreous ; thorax and abdomen like
wings ; abdomen in S sj)eckled with blue-grey, in ? with black; anal segment
in 6 with a white black-edged dorsal blotch and the tufts ochreous, the ? with a
black blotch and one at base ; abdomen beneath, pectus, and legs pinkish white ;
antennae black, the shaft in cJ annulated with white; the pectination full.
Expanse of wings : 35 mm.
1 c?, 1 ?.
Subfamily SEMIOTHISINAE.
214. Gonodela papuensis spec. nov. and ab. densinotata, ochrimixta,
albisparsa, and caesiata nov.
S- I'Ofi'iriiig : fawn-colour, dusted with grey; costa finely dotted dark and
yellowish ; lines brown, somewhat indistinct : first at one-fifth, right-angled on
subcostal vein ; second at one-half, angled on vein 0, then vertical, parallel to first ;
outer at three-fourths, also angled on vein (5, then incurved to two-thirds of inner
margin, thick and dark brown, followed by a i)ale and then a dark line, before the
dark brown marginal area, the u])per edge of which runs obliquely from angle of
( 155 )
outer lino to below apex ; a clomly waved darker .submargiual shade ; marf;iiial
black spots between veins; fringe dark brown ; cell-spot small, black.
Illiu/winq: without basal line; antemedian incurved in cell before the cell-
si)ot; outer double line straight, at thrce-fiftlis ; the marginal brown area paler.
? without the dark brown marginal area ; often with a double black blotch
before outer line from 5 to 3, with a single black blotch beyond it between 3 and 4.
Underside of forewing of S brown along costa and in marginal area; basal liaU'
ochreous spotted with brown ; middle and outer lines Ijrown ; some dashes of bluisii
white before apex and bluish-white sjiots at base of fringe : hiudwing bluish white
speckled with brown, with the lines brown and fulvous, the veins fulvous ; in
the ? the bluish-white tint is much more developed in both wing.s ; cell-spots
black.
Head, thorax, and abdomen fawn-colour ; face and palpi mottled with Itrown ;
antennae ciliated.
Expanse of wings : 30 mm.
7 c?c?, 4 ? ?.
The aberration (lensitiotata occurs in the ? only ; all three lines are distinct,
thick and brown-black or black ; the outer is followed by dark blotches between
veins, interrupted between 4 and 0 in the forewing, less marked but not interrupted
in hindwing ; the ground-colour is pale lilac-grey, or whitish, or pinky ochreous,
with more evident speckling; underside more variegated, the lines more prominent.
3 ??.
In ab. ochri mixta the ground-colour in both sexes is dirty whitish speckled with
darker ; the S S, as in the type form, with darker marginal areas.
3 SS, 3 ? ?.
In ab. albisparsa , confined to the ? ? only, the ground-colour is white, more or
less densely powdered with grey ; the marginal area darker grey with a whitisii
blotch above vein 0 ; the lines dark from costa, and often marked by dark dashes on
veins ; the black blotches of outer line nearly always present, and in some instances
largely developed and laterally confluent; the underside much whiter.
8 ? ?.
Lastly, the ab. caesi/ifa, which is restricted to the c?t?, has the whole upperside
slaty purplish ; the lines dark, tinged with ferruginous : the underside with fewer
whitish scales.
3 c?c?.
21.5. lulocera albinigra .spec, no v.
Forciciiiij : uniform dark purplish slate-colour; costa with fine pale dots;
lines blackish ; first at one-fifth, slightly bent at median vein ; second before
one-half, angled below subcostal, and touching the black cell-spot ; outer line at
two-thirds, bluntly bent at vein 5, preceded by yellowish scales, thickened between
3 and 4 and followed by a dark cloud ; a white subapical spot above vein 0 ;
fringe concolorous, with white spots along base beyond the black marginal spots.
Himlwimj : with antemedian and outer lines close together and i)arallel, the
outer edged internally with yellow scales ; cell-spot black ; submargiual line pale,
very indistinct.
Underside with the lines and bands dark, the intervals paler, with a few bluish-
white specks, all much more prominent and numerous in the hindwing ; subapical
spot of forewing and basal spots of fringe white and largely developed.
( 156 )
Head, thorax, and abdomen concoloroiis.
Expanse of wings : 20 mm.
1 S.
21C. Petrodava gibbosa ali. subradiata nov.
This form is distinguished at once by the underside of the wings, which have
both folds marked from base with snow-white streaks, the cell-fold in the hindwing
being white from base to margin, and the subniedian being often acconijiiuiifd by
another on each side ; in some cases tlie intervals between veins towards liiiidiuargin
are also streaked with white and black scales.
3 Jc?, 2 ¥?.
Sl'bfamily ENNOMINAE.
217. Azelinopsis amaura spec. nov.
Forewiiig : pearl-grey, largely snft'nsed with olive and darker grey; basal patch
qnitc small, olive ; band following it broad on costa, narrow on inner margin, dull
pearl-grey with a darker centre ; inner edge of central fascia from one-fourth of
costa to one-fourth of inner margin, bent in cell ; a similar median line ; outer edge
of fascia from two-thirds of costa to three-fourths of inner margin, minutely dentate,
edged with pale ; all three lines olive, starting from triangular costal blotches ;
costa whitish between the lines, dark-speckled ; space between median and outer
lines pearl -grey ; marginal third olive-brown, with a waved dark p.ale-edged
snbmarginal lino and a yellowish blotch on each side of it beyond cell ; fringe
olive-brown.
Hindwing : pale grey, speckled with darker ; marginal area olive-grey ; a
curved median and Innulate-dentate grey snbmarginal line, both starting from
dark brown blotches on inner margin ; cell-spot grey.
Underside dark olive-brown, with blackish speckling, the forewing greyer and
blurred ; all the lines dark brown.
Head, thorax, and abdomen olive-brownish.
Expanse of wings : 3n mm.
1 ?.
Differs from typical Azelinopsis in not having hyaline cell-spots ; the excision
in hindwing oblique, and reaching only from vein 1 to 2. In forewing, veins 7, 8, 9
are stalked, but rise near end of cell ; and veins 10, 1 1 arc coincident, not separate.
Epitherapis gen. nov.
Foi-ewinij : narrow ; costa faintly arched and slightly inbent beyond middle ;
apex blunt ; hindmargin oblique outwards with faint creuulation to vein <'<, thence
curved inwards ; anal angle obtuse.
llindwinfi : ample, broader than forewing, with rounded crenulate hindmargin.
Abdomen (c?) long, slender; anal tuft bifid; antennae jilumose, the extreme
apex simple ; palpi short and broad, the terminal segment minute ; tongue and
frenulum present; hindlegs broken off; forewing with fovea at base.
Neuration : forewing, cell half as long as wing ; discocellnlar vertical, curved ;
first median nervule at fmr-fifths, second close before third ; radials normal ; 7, 8, '.»
stalked ; 10, 11 stalked: hindwing, costal and subcostal closely approximated for
half of cell ; veins 7 and 3 from before angle.
Tyj)e : Epitherajns stramincata spec. nov.
( 157 )
218. Epitherapis stramineata spec, no v.
ForeiciiKj : straw-colour, washed here and there with jiale oehreou.s ; costal
edge pale, with oblique dark streaks and grey-speckled towards base; first line at
one-fourth, angled on median, marked by black spots on veins ; outer line from
just before apex, oblique to vein 4, then still more oblique to inner margin before
middle, black, with deeper black points on veins ; followed closely by a dark smoky
grey shade from vein 7 to 1 ; snbmarginal line shown by whitish wedge-shaped
marks between veins, those above middle followed by black scaling; a grey-brown
triangle on hiiuhuargin from apex to vein 0, the costal space before apex whitish ;
cell-spot brown ; a slight brownish median shade, and a browu llusli before
snbmarginal line.
Ilindiciiiij : with single obscure antemediau line ; the browu cell-spot followed
by a difl'use triple grey postmedian shade, bent outwards on submedian fold ;
snbmarginal line preceded by a row of brown patches between veins ; the margin
grey-brown ; fringe greyish ochreous ; inner margin and fringe grey-brown.
Underside paler, with all the brown markings distinct.
Head, thorax, and abdomen straw-colour Hushed with pale brown ; palpi
externally and forelegs blackish.
Expanse of wings : 48 mm.
1 c?.
219. Garaeus papuensis spec. nov.
Foreiviiig : rich chestnut-brown ; costa white at base, and with white patches
beyond all the lines ; the brown area with transverse blackish striae ; lines broad,
dark grey, varied with pale grey and whitish scales : first from one-fourth of costa
to one-third of inner margin, with dark spots on veins; outer line from three-fourths
of costa, oblique outwards and angled on vein G, then oblique inwards, widening
downwards, to two-thirds of inner margin, lunulate-dentate externally, the teeth
marked with white-pointed black spots on veins, followed on costa by a snow-white
streak ; the costa towards apex blackish, traversed by a snbmarginal white line, finely
waved, to a yellow spot above vein 6, then diffusely blackish and ending in a zigzag
whitish grey streak at anal angle ; a deeper brown median shade, angled at the
black cell-spot, then obliijue; fringe bright chestnut, with a yellow spot below apex.
Hiniliciixj : greyer, mixed with blue-grey scales, and whitish along inner
margin ; a broad red-bruwu median shade, and a wide brown band above anal
angle, traversed by a bluish white lunulate-dentate snbmarginal line.
Underside dark brown thickly sprinkled with bluish scales ; basal patch of
forewing, and median shades of both wings brown ; outer lines blue.
Head, thorax, and abdomen cliestnut-brown ; vertex and shaft of antennae
ochreous.
Expanse of wings : 30 mm.
1 c?.
220. Heterodisca castanea spec. nov.
Foretciiuj : bright chestunt-brown, with thick darker brown transverse striae;
but the iJaler ground-colour is only visible in the lower half of central area, all
the rest being clouded with dcej) brown ; lines marked by black white-tijipcd dots
on veins ; first nearly straight from one-fifth of costa to one-third of inner margin ;
outer straight and oblique from five-sixths of costa to two-thirds of inner margin ; a
( 158 )
(luik bliitcli oil (liscocellular ; costa strignlated with chirk brown and Ncllow; fringe
browu, pale sfjottt'd in ajiical half.
Uindiciitg : fuscous tinged with reddish, with a slightly curved line just beyond
middle, marked by black dots on veins ; fringe red-brown.
Underside of forewing like iqiperside of hindwing, with dim dark striae e.xcejit
on inner-marginal half: hindwing redder; costal area and cell with black and
grey striae; inner margin paler, with dark striae; a curved postmedian dark- lino
and black cell-spot ; veins and parts of hindmargin red.
Head and thorax woolly, deep chestnut ; abdomen duller red with a grey tint.
Expanse of wings : 36 mm.
1 ?.
Lobophysa gen. nov.
A development of (loiwphaga, characterised by tiie invscnco of a lobe at base
of inner margin of hindwing in the S , reaching about one-third, and preceded at
base by a double hyaline elongate space on each side of vein 1, which is swollen.
Type : Lobophijsa vagilinea spec. nov.
GoHOphiiga straminca Warr., Nov. Zool. x. }>. 408, must be removed to this
geuns.
~21. Lobophysa vagilinea spec, nov., ai.d ab. obsoleta nov., perstrigata nov.,
and colorata nov.
Forewing : pale fawn-colour (when faded, grejish white), sparsely black-
speckled ; lines blackish, sharply defined ; first bracket-shaped, at one-fonrth,
vertical from subcostal to snbmedian vein, both extremities retracted; outer line
from three-fourths of costa oblique and slightly curved to vein 7, thence oblique
inwards with a slight curve to before middle of inner margin, preceded below
middle by the median line, of which the upper portion is obsolete ; the angle
of outer line at 7 touches the apex of a brown-grey triangular blotch on hind-
margin extending from apex to vein 4 ; submarginal lino hardly marked except
by a black blotch at anal angle ; a row of dark dots just before margin ; fringe
concolorous ; cell-spot annular, small.
Hindwing : with a dark mark at base, a difl'uso dark lino of scales from
cell to end of lobe on inner margin; a black median line, generally curved
inwardly round the annular cell-spot, joined on inner margin by iiu uutcurved
postmedian line, which is sometimes completely lunulate-dentate, at others marked
only by the dark teeth on the veins : submarginal line from apex to anal angle,
angled between 4 and G and outwardly paler margined, with black spots on the
veins.
Underside paler, with the lines, cell-spots, and speckling blacker ; submarginal
lines wavy ; apex of forewing whitish ; a short brown line from origin of vein 2
to inner margin, and a curved brown line from two-thirds of costa to three-fourths
of inner margin.
Head, thorax, and abdomen concolorous ; palpi externally black ; face
browner; shoulders brown; metathorax with a pair of black spots; some black
spots on middle dorsal segments.
Expanse of wings : 40 mm.
In the aberration obsoleta the Hues are more or less obscured ; the basal
line being represented by dark dots; a brown blotch edged by two black lines
( 150 )
from inner margin to snljmedian foUl represents tlic ends of the median and
onter lines ; and in one instance tlic submarginal line is marked liy a large black-
brown blotch at anal angle and another beyond cell on the edge of the brown
triangle; the hindwing and underside remain as in the type.
The aberration jjersfiiifafa is much more abnormal : the ground-colour is pale
pearl-grey ; all lines and markings are nearly absent; a thick donble brown-black
streak, well-defined basewards but diffuse outwards, runs from jnst below apex
of forewing to middle of inner margin, and is produced across hindwing as a
broad band occupying the second fourth of the wing, with the black cell-spot on
its outer edge ; underside the same ; the dorsum with a broad brown band across
third and fourth segments.
2 c? c? of the type form ; 5 t? (? of ab. ohsoleta ; and 1 c? of jjerstrigata.
The two brown lines of the underside of forewing are visible above only in
worn specimens.
In the third aberration, colorafa, of which a single c? only has been sent,
the colour, instead of being uniform as in the others, is varied ; the ground-
colour is pale grey striated with olive ; the basal area and most of the marginal
area beyond outer line are dull green ; the blotcli on inner margin at the end of the
fine outer line is jiale brown, while the cell-spot is rufous and the hiudmarginal
blotches are rnfous brown ; ia the hindwing only the base and the space between
onter and submarginal line are pale, all the rest being tinged with rufous and
olive, the antemediau line and the inner edge of submarginal being ferruginous.
Underside pale pearly grey with all the lines shown distinctly.
1 c?.
Moneta gen. nov.
Forcwiiii) : costa curved at base and shortly before apex, which is bluntly
produced ; Iiindmargin obli(|ue outwards to vein 4 with two deep excisions, and
blunt teeth at apex and veins 6 and 4 ; below 4 obliijne inwards and crenulate.
Hindwing : with well-rounded crenulate hindmargin.
Antennae simple in both sexes ; palpi stout, obliquely porrect upwards, third
segment short ; forehead slightly prominent ; tongue and frenulum present, slight ;
thorax and jiectus hairy ; hindtibiae swollen, with four sjjurs ; abdomen of S
slender, with slight lateral tufts.
Neuration : forewing, cell half as long as wing ; discocellular oblique ; first
median nervule from one-half, second shortly before end ; subcostal vein strongly
beut down at extremity to meet the discocellular; vein 5 from the top of discocellular,
vein fi long-stalked with 7, 8, 9 from the bend, lU, 11 coincident, lU subsequently
anastonjosing shortly with 8, 9, these last two veins separating quite close to
costa : hindwing with costal and subcostal shortly approximated at base ; 3 and 7
before ends nf cell.
Type : Moneta plenicolor sjiec. nov.
-".'-. Moneta plenicolor spec. nov.
Foretciiitj : deeji red-brown, the ground-colour, which only shows through here
and there, being bright red-brown ; costa deep orange, spotted with brown ; first
line obliiinc, from one-sixth of costa to one-third of inner margin, interrupted,
running from subc<istal vein to median, and from snbmedian fold to inner margin,
white, with orange edging ; outer line obliij^ne and straight, from costa close before
( 160)
iipex to three-fourths of ioner maru:in, also interrn])teil, from vein 0 iicavly to 4,
aud between 3 and 2, where visibh', luniilatc-dontatc ; bntli lines slightly toothed
on veins ; an obscure dark cell-s])ot ; fringe brown, ti])i)cd with yellow.
Hindwinq : vinous red, deeper in outer half and black-speckled ; cell-spot
dark ; traces of an obscure Ininilatc-dontate postmedian line; fringe dark, with pale
yellow tips.
Underside paler ; forewing blurred brown, with fuscous speckling ; cell-spot
and sjwts on veins along outer line blackish ; marginal area dull yellowish in
iiround-colour ; ai)ex grey : hindwing deep brown in marginal area and along
inner margin; inner area sprinkled with lilac-grey aud coarsely black-speckled ; a
large black cell-spot ; spots of outer line black.
Head and thorax like forewiug ; abdomen like liindwing ; shoulders aud p;il|ii
tipped with orange ; tarsi rich brown, mottled with orange.
Expanse of wings : 48 mm.
7<?c?,l ?.
223. Nadagara cuneigera spec. nov.
Forcwinq : fawn-grey, in the S sometimes with a reddish tinge, covered with
blackish speckles ; the costal edge slightly darker, aud the subcostal vein pale
whitish ; lines dark brown edged with whitish, the inner inwardly, the outer
outwardly; both start from subcostal vein ; first at about one-third, oblique outwards
to two-fifths of iuuer margin ; outer from three-fourths of costa to fourth-fifths of
inner margin, its white edging diverted along subcostal to a])ex ; a tine subniarginal
white line forming wedge-shaped marks between veins, between 3 and 4 filled in
with dark scales ; fringe fawn-colour ; a black cell-spot.
Bindiring : without basal line ; the outer curved.
Underside whitish grey, speckled with darker ; the lines and cell-spots dark.
Head, thorax, and abdomen concolorous ; face, shoulders, and dorsum darker
I'rey in ?, ferruginous in S ; patagia paler.
Exiianse of wings : 34 mm.
4 (?(?,4 ? ?.
224. Prionia pulchricolor s]iec. nov.
Foreiriiiq : in 3 pearl-grey, in the ? darker, with a reddish brown suffnsion,
and in both sprinkled with black ; basal area darker grey, followed at one-third by
a deep "reen band, which above median is narrow and funnel-shaped, and below it
swells out into an oblong sometimes rounded blotch ; this lower part in some cases
becomes quite jjale green, or yellowish grey ; the band is edged with pale and
sometimes ])receded on costa by a dark spot; cell-spot blackish green ; snbmarginal
line dark "rcy or dark olive, lunulate between the veins, connected with ajiex by
an obliijue streak ; from three-fourths of costa a dark green streak curves outwards
to vein 0, where it is sharply angled, and runs obli(iuely to two-thirds of inner
margin ; tliis outer line is rarely ]ilain ; fringe deep olive, with pale flecks beyond
the veius.
Iliiidwimi: bright rosy in costal half, grey-green belnw median and along
margin beyond the waved snbmarginal line; from end of cell to inner margin a
dark green band, crossed by the ])alc veins.
Underside of forewiug blurred reddish fulvous along inner margin, ociireons
with dark brown striae along costa ; marginal border and outer line filled with
( 161 )
hoary grey, witli a yellow patch between them ; in tlie ? the ochrcons is suffused
with tawny brown : hindwing with base ochreons and outer half tawny brown
with dark striations ; the margin as in forewing ; ? wholly suffused with tawny ;
fringe in both wings dark brown with pale tips.
Face blackish green ; head and thorax dark grey, the vertex paler ; abdomen
pinkish grey, darker towards base ; in ? all the parts are darker, more suffused
with reddish grey ; palpi, pectus, and legs bright fulvous.
Expanse of wings : c? 28 mm. ; ? 32 — 35 mm.
5 c?c?, 4 ? ¥.
TWO NEW AGABISTIDAE.
By KARL JORDAN.
1. Aegocera leighi spec. nov.
<?. Side of palpus and the forecoxa chocolate ; head grey, a dot between antennae
brown, centre of frons and the crest of hairs at eye chocolate ; thora.x chocolate
.above, densely irrorated with grey and pale yellow scales ; abdomen orange above,
with central row of black spots, grey washed with chocolate on underside; tibiae
orange on upperside, fore- and midtibiae with two black dots each ; tarsi black, tips
of segments white ; bases of abdominal side-tufts chocolate beneath.
Wings, upperside. Forewing chocolate, densely irrorated with huffish grey
scales ; an irregular streak on median vein somewhat as in rectiUrica Boisd. (1S36),
irregularly sinuose, creamy white, not ijuite reaching a short costal creamy white
band which extends from lower angle of cell obliipiely to costa ; submarginal
area with an indistinct flexuose creamy line ; a row of minute admarginal chocolate
dots. Hindwing orange ; a small dot on cross-veins and a row of submarginal
spots black, the middle spots of the row minute.
Underside orange, apices of both wings somewhat washed with chocolate ;
forewing with a black dot in cell near apex and another on cross-veins ; hindwing
with a black spot on cross-veins and three or four submarginal spots, which are much
smaller than above, situated from anal angle forwards.
R- of forewing from apex of areole, as iu ^1. rectilinea ; areole longer than in
that species.
Length of forewing : 15 mm.
Ilab. Durban, Natal, October 1904 (G. F. Leigh) ; three specimens.
2. Argyrolepidia pamphila ombiranus.
?. y^'mgs, upperside ; blue tcaling reduced to a few dots on forewing and a
minute spot in centre of hindwing. Forewing white, cell-spot minute ; white
discal band very much broader than in p. pamphila, widening behind, stopping short
at M", measuring 3 mm. in width at M^
Underside: blue scaling also reduced, but not so much as above; central sjJot
of hindwing isolated.
Ilab. Ubi, Moluccas (J. WaterstradtJ ; one ?.
11
( 162)
ON A NEW PARASITIC TINEID MOTH EROM QUEENS-
LAND, DISCOVERED BY P. F. DODD.
By the HON. WALTEIt ROTHSCHILD, Pii.D.
QIINCE Westwood* pnblisLcJ au account of a Chiuese moth of which the laivu
K--^ is parasitic oa Fahjoridae, several lustauces of such parasitism have been
recorded from ( 'entral America, New Mexico, India, and Jajian. So far no snch
jiarasitic larvae have been ibnnd in Al'rica, but it is highly iirobable that they occur
all through the tropical and subtropical countries. Some years ago Mr. P. F. Dodd,
wlio has added so much to our knowledge of the early stages of Qneenslandian
Le]]ido]itera, discovered also a jiarasitc of this kind in tjueeusland. He has sent
us an interesting account of this discovery, which we a])])end here in full. Tlie
insect is closely allied to that figured by Westwood, but ({uite distinct. Dyarf
places these moths among the Tiueids, and we perfectly agree with what he says.
Epipyrops doddi spec. nov.
(??. Legs and shaft of antenna luteous. Wings olive above, the scales of
the forcwing partly silver grey, especially on disc ; no distinct markings ; underside
drab-ljrown.
Ncuratiou : nine veins from cell of forewing, 4* of Westwood's figure being
absent and i^ very weak near cell ; cell of hindwing mnch narrower than in
Westwood's figure, without inner spur {x of Westwood's figure), only five veins
from cell, b- of Westwood's figure being absent, upper cross-vein straight, obliijue.
Length of forewing : 4 to 0 mm.
The mandible of the larva is bifid, as in Westwood's figure ; but the second
lobe is more hook-shaped than in that figure, being w-idest in centre. The abdominal
legs have a complete ring of hooks, the anterior hooks of each ring being much
smaller than the posterior ones ; the anal leg has a posteriorly ojien half-ring of
large hooks, instead of a complete ring.
Mr. Dodd found this insect on several species of Falyoridac, since identified by
Mr. AV. L. Distant as Dictijophora prac.f errata Dist., Olonia s])ec., and a species of
the subfamily Flatinae.
Mr. Dodd's observations on the parasite arc as follows : —
"Upon June 15th, 1903, I received my copy of the EutomologUt for May, and
was very jdeased to read Mr. Kirkaldy's instructive 'Current Notes' therein. Those
in connection with parasite moths were of absorbing interest to me, for 1 have
been observing a carnivorous caterpillar here, breeding out the moth for some
time, and hoped I had a great surprise in store for entomologists. I have had
my account of its history, so far as 1 knew it in June, almost ready for some weeks,
but had delayed its completion and desjiatch until I could obtain the necessary
larvae upon pupae of the Ftdyoridae, so as to give fuller particulars concerning
the development of the white coating, for I had not been satisfied with my
observations upon those on the winged insects, owing to the great difficulty in
obtaining even a temporary view of a larva before its larger growth. Whilst it
is small it is not easily examined, as it is i)artly hidden, and the hosts object to
» Traim. Eiit. Soc. Land., p. 519. t. 7 (187G>. f Proo. EiU. Sw. Washiwjt. v. p. 13 (1902).
( 163 )
close senitiii)', becomiug restless and ofteu wiudiuy iiji with an aggravating- hop
when least desired. However, I have since obtained the renuisite specimens, and
have ascertained mnch that I wished to know.
" My larvae are certainly parasitic. I have kept them with tlieir hosts for
several weeks, and, though 1 have had many, I have never detected the slightest
movement in any of them. A web is spun upon their hosts to cling to, and there
they remain motionless once they commence to develop the white covering. The
very young larvae are usually very close up to the thorax, and may sometimes
lie across the al)domeu ; but, as they become larger, and require more space,
change their position and lie parallel to it, the head being directed towards the
end. The body presses closely down upon the host, and the head is drawn in
very much, generally resting upon the edge of a segment. As a larva grows, the
wings of the homopteron are forced outwards and upwards a little. The abdomens
of many of the parasitised ' hoppers ' are usually much siirunken or depressed where
the larvae adhere. The snouted Fulgorid produces fine large caterpillars, and it
will be noticed that it possesses very little indeed of the waxy substance. The
pupae, as well as the imagos, of the black and the green species carry the
caterpillars, yet the former has little or no ' wax ' at the pupal stage ; therefore
it cannot for a moment be conceived that there is anything else than the juices
of the Fulgoriilac for the caterjiillars to live upon.
" The moths, like various carnivorous species with which I am acquainted, soon
become greasy.
" I may mention that 1 had not the faintest idea that the species of moths
referred to by Mr. Kirkalily were known to exist, nor had I ever seen or heiird
of the Japanese publication, the Insect World.
" Some four years ago, during a visit to Mr. H. Tryon, our State Entomologist,
in Brisbane, he drew my attention to a small white cocoon, which he had found
attached to a twig in his garden, and which he aptly compared to a rosette.
The moth tlierefrom also was shown to me. 1 had not seen the species before,
nor, I believe, had Mr. Tryon. Not long afterwards I came to Towusville, some
800 miles north of Brisbane, and, during my entomological wanderings here,
have found similar cocoons at intervals, and obtained moths therefrom. The
cocoous usually were upon narrow blades of grass, but I did not search for the
larvae, considering any attempt to find them as utterly hoi)eless. However, during
April this year, I found several of the cocoous upon grass, under some shrubs,
and having during the previous year (1902) discovered that a caterpillar, which
constructs a somewhat similar cocoon, and passes the earlier part of its existence
upon certain homopterous insects (adhering to them and much resembling small
bloated ticks, and undoubtedly imbibing their juices), I at once commenced a
careful search amongst the several species of Homojitera in the vicinity. Ere
long I noticed a small whitish Fnlgorid, with wings somewhat apart, and looking
decidedly uncomfortable ; upon boxing this I found, to my extreme gratification,
that there was attached to it a stout, oval, and almost snow-white lepidopterous
larva, which I felt confident was the insect I was seeking for. Soon after several
more of these 'frog-hoppers,' each with a larva attached, were taken, tlio larva
being under the wings and a little upon the side of the abdomen. The caterpillar
upon the first hopper crawled off' that night, and next morning proceeded to
construct its charming little cocoon. The moth emerged early in the afternoon
thirteen days later.
( 164)
" Many hours were spent during the ensuing tew weeks iu searching for these
strange calerpiihvrs, wliicli were also t'onnJ uiioii tliree other species of Fuli/orifhe ;
some more cocoons, too, were discovered, but throe-fourths of tliese, and of otiiers
collected later, produced small jumping ichneumons, six, eight, or eveu twelve
emerging from a cocoon, according to its size. Presumabl}' the larvae are stung
after leaving their hosts— in all probabilit_v whilst sjiinning, for none of those that I
have taken from time to time proved to be victimised ; maybe tiiey are safe under
the wings of the hoppers, though, as they ajiproach maturity, they cause sjome
displacement of the wings, and though easily seeu may present no weak point
to be attacked.
"The larvae are not rare, but there are only special localities where I ol)tain
them ; small shrubs where ants abound are j)roduclive, the hoppers often being
numerous in such places, even where the green Oecopliylla ants are. The auts do
not fraternise with these Ftdgoridae, like they do with so many species of Jassidue,
Fgi/Uidae, etc. ; but they do not molest them, and pass by in scores without seeming
to notice their presence. I have seen the green ants traversing branches where
there were no less than three species oi Fahjoridae dotted along them.
" I have never found two larvae of the same size upon a hoi)per, but not
infrequently have noticed two of difl'ereut sizes, the second, in every case, being very
much smaller than the other, and upon the ojijiosite side. With but few excei)tious,
all the larvae I have seen were upon winged insects, the exceptions, as alreiuly
stated, being ujmu pupae of the green and the black sjiecies only. I have kept
these for many days, supplying succulent shoots regularly to the hosts, until the
larvae changed to white, and finally departed to spin. UiKin another page I give
particulars of several larvae which I obtained some time ago, two of them when
they were exceedingly small, and which I was enabled to carefully observe for
many days ; the hosts are still alive, one with a second caterjiillar upon it, but,
as the larger portion of my material was despatched five weeks ago, I cannot
delay my notes any longer, or would give fuller particulars concerning these.
{}\ote. — I kept my notes back for seven weeks, by which time two of the hosts
had died, the third dying several days after despatch of notes.)
" A full-grown caterpillar is in reality a very small creature, with a thick,
eveu coating of a waxy-white and light-as-down substance covering the dorsal
surface, the sides, and extending well underneath ; it is so thick that the cater-
pillar ajipears to be double its real size. This coating parts slightly at the segments
when the insect is in motion or curled up, — wheu parted from tiie host, if touched or
alarmed, it curls up almost into a ball, and can easily roll from whatever it may be
upon, and generally dangles by a thread, and thus suspended bears a strong resem-
blance to a round hanging egg-bag of a spider. The head and adjoining segment
are (Mily slightly covered with this white matter, liut are drawn closely to the
thickly clothed segments, leaving the face alone visible ; wheu the insect is extended
the head can be seen.
" I have collected larvK of various sizes, some being exceedingly small,
evidently not long from the egg, and, owing to their colour, scarcely discernible, the
lens revealing their presence. Very young specimens arc much the colour of orange
pulp. In about ten days this gradually alters to didl purplish red, after which
it daily becomes paler until almost colourless; then the skin slowly assumes a
whitish hue, as if caused liy a thin deposit, and becomes somewhat distended,
and the head is raised a little ; it is now perceived that a moult is about to take
( 105 )
place. As the skin whitens, and nbont the second, or eaidy on the third day, it bursts
underneath, but is not thrown off for some time, the period depending greatly
upon the host, which, if lively and inclined to hop, or open and close its wings
freely, may loosen it and cause it to iall after the third or fourth day, but it,
may remain attached to the larva's back for a week. As the old skin bursts, and is
forced outwards and upwards, only a portion of the new one can at first be seen,
but that which is exposed is perfectly naked, and the colour is temporarily darker;
the whole new sMu is at this time hare, for I have lifted the old one to make
sure, and, as the old one comes away at the sides, in the course of a few hours
the gradual whitening process is repeated, this time, however, developing into
the unique and dense coating. It is impossible to ascertain how long this is in
forming, once it becomes quite white, but it evidently takes several days, for by
then the larva appears to have doubled its size. During this development the
old skin is slowly pushed and folded up along the sides, bnt it retains its shape
above, and comes away when the new covering admits of its being dispensed with.
This extraordinary growth is doubtless intended for various protective purposes,
several of which I may suggest, as follows : as the naked larva, secure under
the ample and close-fittiug wings of the host, increases in growth, it would
probably open the protecting wings a little, and thus be rendered liable to the
attentions of ichneumons, so though the waxy growth opens the wings, in some
cases considerably, before they are actually apart, the coating has become suffici-
ently dense to eft'ectually shield the larva from those insects ; being a soft little
thing, it may require some protection from the wing pressure, which the covering
aftbrds ; further, it keeps the larva dry in all weathers, assists to whiten the
cocoon and render it waterproof, and gives it, when parted from the host, the
appearance when moving of a mealy bag, or when still of a waxy scale. Upon
removing and examining some of this waxy coating it is found to compare with
that borne by various species of Ftilyoridae, and with which several of them
cover their ova deposits; if touched with a pointed instrument it comes away in
little tufts ; these viewed tlirough a lens prove to be comjiosed of numbers of
delicately thin, short, and glistening white hairs, which, rubbed between tlie
fingers, leave the merest trace of white — are almost rubbed out of existence.
" A blade of grass, a thin twig, or a leaf, may be selected to spin the cocoon
upon ; the larva could drop there by a thread ; however, the overloaded host is
often in the grass, — perhaps tumbles there in essaying a leap. So the larva spins
upon whatever it may drop or crawl on to. It is very sluggish, and not meant
for much wandering, a distance of several inches being the usual extent of its
journeys ; it often remains for hours in the one spot. Many times I have dis-
turbed a larva when in the act of spinning, causing it to cease instantly, work
in some cases not being resumed for a few (up to fifteen) hours ; only lately a
large larva which, through being shaken, ceased spinning late one afternoon, when
the sides and ends only of the cocoon were finished, did uot resume operations
until 11 a.m. next day ; these long stoppages have invariably included all the
hours of night.
" The cocoon is a pretty and neat little object, which requires the powerful
glasses, and the pen, of the advanced entomologist to view and describe it and
its construction in a satisfactory manner ; however, I shall attempt the task. It
may pass equally well for a waxy scale, a mealy bug, or a little flower ; j)erhaps
it is best compared to a flower, for the ornamentation certainly suggests one,
( 1C6 )
with two petals exjiaiidpil and several others partly o])ene(l ; these (litTer in number,
the totiil varying t'roni four to six. After ]irepariny the fonndation of the cocoon
the caterpillar quickly builds up the sides ; after these one end, or the first " petal "
is spun, then, turning about, it spins another, thus forming the other end.
Commencing abont half way down one of these, a portion of the top of the cocoon
is added, the web being continued on and outwards to form the third "petal."
Finally, there are generally five of these, the centre two being as if abont to
open. This ornamental work ends in the centre, the last two jietals, which are
somewhat curled, being drawn tugether and fastened. The spinning of the " flower,"
after completion of the foundation, takes some ninety or a hundred minutes, rather
longer by the larger larvae, but there are several stopjiages, for the insect in
its various turnings pays much attention to the bottom and ends of the structure.
80 far this web is composed of row after row of loops, or rings ; each row runs
very evenly, and is spun under the preceding one, and joined to it all along about or
a little below the centre, so that a row scarcely jirojects luilf its height beyond the
one spun before. When a row of loops is finished the next is commenced where
that ended, — the work going on alternately from side to side. The web is formed at
a rapid rate, 95 to 115 loops being spnn per minute by small larvae, a lesser
number liy larger. When sjiinning, the actions of the larvae are so peculiar that
they cannot fail to attract attention, for the head, in moving up and down, is
tliriiwn much farther back, and thrust much more underneath, than apparently
necessary ; but it is soon ascertained tliat these extended movements are made
with a s])ecial object, and that is to give a partial coating of the waxy substance
to the web, for it will be seen that the head is supjjlied with short hairs, and as
it is thrown Ijack they touch the wax — already becoming slightly loosened, and
in wooll3'-looking tnfts, owing to the insect's various twistings and turnings, —
and little masses are caught upon them, which gradually transfer the substance
to the loops Ijy being purposely and regularly brought into contact with them as
the head moves up and down. A little of the stuff, here and there, also becomes
dislodged, and adheres to the fabric wherever it tonches. By the time the " flower "
is corai)leted the waxy matter has disappeared from the first thickly clothed
segment, and often quite from the next, as well as small portions from the sides.
The whole of the balance is then worked into the threads of the cocoon proper
nnderneath ; not a particle seems to be left upon the larva when the spinning is
completed. When the "flower" is finished the larva can be seen but indistinctly,
therefore its actions cannot be followed, but the restricted space would scarcely
admit of the head work referred to, so the waxy matter must be rublied or pressed
between the threads as the cocoon progresses.
" Hliould a cocoon be knocked sharply, or torn asunder, little clouds of white
jiowder fly oft', or a cocoon will leave patches of white upon anything with which it
comes in contact. The ichneumons all issue from the one hole, and the earlier
ones to emerge are finely powdered.
"It will be noted that the cocoon has a wide "frog-mouthed" slit, and
that the pupa shell protrudes therefrom. The mouth is forced open by the
pupa, but closes down njion the end segments, holding them securely enough to
enable the moth to burst the shell and escape. Several other species, including
the one referred to in these pages, which I have bred, but which, however,
are ant associates, form these " frog-mouthed " cocoons, and the moths emerge in
a similar manner.
( 167 )
" The moths emerged from thirteen to eighteen daj's from the time they
commenced to spin, appearing abont midda}'. They expand from 5 mm. in c? c? np
to 14 mm. in the hirger ? ? . Those from tlie white Fnlgorid are smaller than
exam])les from the others.
" It is most difficnlt to canse the larvae, or the little white waxy Falqoridao, to
adhere to eard, but by rubbing the thin deposit from the latter they can be fastened
down on their backs, with wings spread to show the attached larvae. The larvae with
their hosts will not sink in formalin solution ; owing to the lightness of the " wax,"
and to its being imjiervions to moisture, they float like corks, so can only be thrust
under and held there : a piece of wadding pushed down upon them in the tubes
keeps them immersed."
" The caterpillars do not appear to cause the death of their hosts (or, if so,
not for some weeks), which, when free from them, become as lively and healthy-
looking as ever ; then does not the presence of the second caterpillars on some
specimens help to bear this out ? Many times, when a matured larva has departed
to spin, I have found a much smaller one, of whose presence I had not been aware ;
but there is another parasite, an external ichneumon, which exists upon the lilaek
Fulgorid and several larger species (there are usually three to five, each eni'losed
in a thin dark shell), and when they break through their shells and drop off the
host dies. I have actually had one of these parasites upon a Fulgorid carrying
a fair-sized moth larva, but mislaid it or inadvertently sent it away. {Note. — These
ichneumons remained in pupae about four months; one is now sent with other
specimens — July 1905.)
" I have yet to learn where the eggs* of the moths are deposited. If adjacent
to spots where hoppers may then be, the newly hatched larvae would frequently
have to wander for some distance to find them, for they would scarcely remain
stationary for days ; besides, the larvae are too soft and sluggish to be fitted for
much wandering, so I believe that it will eventually be ascertained that the eggs arc
deposited, singly only, upon the hoppers ; however, the snouted and black species
average many more larvae in proiiortion to their numbers than the white, and abont
as many as the green, moreover they are not such stationary insects as the last
two. The white species is extremely common, the green and the black mach less
so, and the snouted one is comparatively quite rare. The last I meet with njion
two kinds of small trees only, seldom more than three or four upon a tree, and
usually far apart, yet it is the most productive species for caterpillars. Again,
there is a fifth Fulgorid, which is common, and often in company with the white and
green insects, yet I have never found a larva upon this one ; so, if the young things
have to search for their friends, it seems strange that this species shoulil escape,
lor it is rather a stationary one. Again, out of six snouted insects on one tree,
four were taken possession of by fine large caterpillars ; and another tree, whereon
were green hoppers, also furnished a number of large sj)ecimens, all of these two
lots being very even in size. If the eggs are deposited on leaves (ir twigs, of
course by various moths and at different periods, why is it that several larvae
of different sizes are not often met with on the same host? or how can the tiny
larvae distinguish between the suitable and unsuitable specimens of species of
Fidgovidac'i one never carrying larvae, and, of the four favoured species, only
* Ihe egg is about twice as long as wide, ovate, being narrowest at the micropyle end, wiilest at
the opposite pnlc ; apart from the circular groove at tlie micropyle pole, the egg is smooth but not
polished.
( 168 )
tbe pnpae of two also being burdened with them. Pupae oi' tlie white insect are
evidently too small and rounded to be patronised, and though the pupae of the
snouted insect are rather too rare to admit of a decided ojnnion being formed,
they wonld not, 1 think, make suitable hosts. I liave examined likely foliage and
twigs, and also many of the hoppers, without finding eggs; those sent were from
my boxes. In support of my belief I may add that the caterpillars of the other
parasitical lepidopterou, referred to several times, are found promiscuously on larvae,
pupae, and images, all being likely to carry several of dilferont sizes, some of the
imagos having as many us six or even eight atlixed to them ; but the moths lay
their eggs on the trees where the hosts abound, and the little caterpillars are
active, and often travel considerable distances to find their friends. These cater-
pillars have two stages, the first being passed on Homoptera in the open air, the
second being passed in ants' nests underground, where they live, still by suction,
upon tlie ant larvae.
" I now give particulars of three Fulgoridae, with attached larvae, which I
kept in glass jars for several weeks for the jiurpose of close investigation. Though
I often changed the jars, to ensure pure air for hosts and parasites, and snpjilied
succulent shoots regularly for the former, the long imjjrisoument evidently atfected
the caterpillars, and doubtless somewhat shortened the lives of the hosts. It will
be observed that the caterpillars abandoned the hoppers within a week of their
final moult, bnt many quite white examples, which were taken in the bush, have
not left to spin until ten or even twelve days after their capture.
"No. 1. PUPA OF GREEN FULGORID.*
1903.
July 12. Taken with large white caterpillar.
,, 15. ('ateri)illar left and spun cocoon.
Aug. 8. Host died.
" No. 2. ALSO PUPA OF GREEN FULGORID.*
July 8. Taken with reddish larva. Perhaps age about 12 to 14 days.
:; :?;)-'■
„ 12. V^ery much paler.
„ 13. Skin whitish and distended.
„ 14. ,, „ „ very much distended.
„ „ 3 j).m. Skin split underneath, partly exposing naked new one.
„ 15. 9 a.m. Skin opened a little more, naked parts whitening.
„ 1(5. „ Skin further opened and rising. Exposed parts of new one well
whitened.
„ 17. ,, Skin adhering only at terminal segments. New one; very white.
„ 19. Larva dropj)ed by a thread and spun cocoon.
Aug. K). Host still alive.
„ 12. „ sickly.] Pa]ier was desj)atched on Itith instant, these particnlai.^
J) 13. ,, died. J being sent on afterwards.
* Belonging to the subtamily Flatinae,
( lf59 )
"No. 3. SNOUTED Oil CLEAR-WINGED FULGORID IMAGO.*
July 8. Taken with pale reddish larva near thorax. Estimated age 13 or 14 days.
Lying lengthways.
Paler. Curled against thorax.
Larva again lengthways, nearer extremity of host's alidomen.
9 a.m. Head raised ; moulting.
Skin whitening and distended. A small object, which I could scarcely
discern for several days, now proves to be a caterpillar.
Head lowered, skin burst ; a little of new one showing.
Old skin rising. Exposed parts of new one whitening. 2 p.m., skin oil".
Very white.
Abandoned host and commenced to sjiin.
Small larva noticed on 14th instant now darkening; it cast a skin five
days ago, which lies under the wings and is not obtainable.
Preparing to moult.
Skin cast.
Moulting again.
Skin cracked underneath and raised from head. Host died, and witli
larva preserved in formalin.
" The pupae possessed a good deal of the waxy matter during and after the
presence of the caterpillars, and one twice put forth two white wavy and delicate
tails, which in time fell away and adhered in fragments to the leaves in the jars.
" I omitted to mention that I had never seeu frasx of the caterpillars, but in all
probability the discharges are liquid, and would at once roll off the waxy bodies of
the hosts. The other parasitical caterpillar, to which I have referred more than
once, emits only li(juid matter." — Frederick P. Dodd.
* ZHctyophora praeferrata Dist .
Mr. Dodd is to be heartily congratulated on these very interesting observations.
( 170 )
NOTES ON THE SIPIIONAPTEBA FROM THE ARGENTINE
DESCRIBED BY THE LATE PROFESSOR DR. WEYENBERGH.
By K. JOEDAN, Ph.D., and the HON. N. CHARLES ROTHSCHILD, M.A.,
F.L.S., F.E.S.
TTTEYENBERGH iu 1881 * described seven species of fleas. Thongt ever}-
VV name is ennmerateil in the Zooloi/ical Record (of the Zoological Society
of London), they have all been overlooked by subseqnent authors. The descriptions
in qnestion do not give those characters which are essential for the recognition
of species, the absence of all figures being, moreover, a great disadvantage in this
publication. We have, therefore, abstained in former papers from trying to identify
any of Weyenbergh's species with those we have from South America.
Professor Dr. Doeriiig, of the University of Cordoba, Argentina, has most
kindly sent us a set of cotypes of all the species described by Weyenbergh, and
we take this opportunity of reiterating our thanks to him. The specimens were
glued on cardboard, as was formerly the custom. In order to examine them
carefully we had them mounted in balsam. Some of Weyeubergh's names, as
we expected, refer to species described later by other authors, while some others
apply to species already described before Weyenbergh, one other again designating
a species which has not been rediscovered.
In the present pajier we discus-s in detail all the following species described
by Weyenbergh as new in the paper mentioned and in a i)revious one:t
Pulex yrossitentris, Pulcx nasuae,
Ceratophijllus i-ufulus, „ obscurus,
„ isidori, „ concoloHs,
Pulex {IIectopsijlla?)testudo, „ caricohi.
Family SARCOPSYLUDAE.
The family comprises three genera : Dertmtophilus Guer. (= Sarcopsylla
Westw.) ; Ecltidnophriga Olliff (= Aiyopsylld Enderl. = Xe.stopfi>jU(( Caker), and
Ilectopsijlln Frauenf. (= JUii/ncIiopsi/llux Haller).
Karsten (1804)emjiloyed the name Rhi/nclioprion Oken instead of Dennntophiltts
(= Sarcopsijlla) for the dhigoe, and Baker followed suit iu 1904 and 1005, calling
the family Jibjnchopnonidae. We remark (1) that Phynchoprion Oken is
])rcoccnpied by Rb/nclioprion Herm. |), and (2) that Rhi/iicho/iricw Oken was
not a term proposed for the Cliigoe, but for mites— the Chigoe, which Oken knew
only from figures and descriptions, being provisionally put into the same genus,
not as Rhynchoprion penetrans, but as Pulex penetrans. Wliatevcr rules of
nomenclature one adheres to, Rliijnclmprion cannot possibly be retained for Linn6's
Pulex penetrans. Baker calls Rhynchoprion " the older and only correct name " ;
but did he compare Oken, Natury. iii. p. 402 (1815) ?
• Periodwp Zoolojicit iii. pp. iifil-"" (1S81),
t Vol. Ao. Nat. Cicnc. Argent, iii. p. 188 (1879).
j 1804. Ucmoirc apteroloi/iqw. The name is proposed tor ccrtnin raitcs.
( in )
Onr SdirojJSijllidae comprise Baker's THii/nchoprionidac and Ilectopsyllidae,
Baker's family distinctions being erroneous, as we have pointed out in a paper
on the Sarcopsijllidae now in press. Wcyenbergh described one species of this
family as Pulex {Ilectops>jlla /) testudo. The species is the same as Fraueufeld'a
psittaci.
1. Hectopsylla psittaci.
IlectopsyJJa psiitaci Frauenfeld, SHz.-Ber. K. Ah. Wiss., Math. Natiivw. Clause xl. p. 462 (1860).
Pulex {[levtnpsijVa ?) testudo Weyenbergli, Feridd. Zool. iii. p. 2G7 (1881).
RhynehopsyUa pulex, Taschenberg, Die Flohe p. 56. t. 1. fig. G. 6a. 7 (1880) (partim ; II. psittaci =
It. pulex, mistake).
The species is easily recognised by the non-angnlate head, the comparatively
short maxillae, which are somewhat curved forward, and by the legs. The fifth seg-
ment of the fore- and midtarsi bears on each side seven or eight spines, the number
being on the hindtarsns sometimes reduced to six. The two specimens of testudo
which we received differ from other specimens in the first hindtarsal segment
bearing on the hinder side a pair of bristles beyond the middle, these bristles
being, however, absent from one hindtarsus in one of the two specimens. As the
individuals which Frauenfeld described as psHtacA have no hindtarsi preserved, it
is not possible to say if typical psittaci possess those bristles or not. There are
perhaps two geographical races, one inhabiting Chili and Northern Argentina
{psittaci = testudo) and the other found in Brazil. However, the material we
have is not sufficient to decide the question.
The insect described by Haller as Rhijnchopsyllits ptclex, though belonging to
the genns Ilcctopsylla, is very different from psittaci.
IF. psittaci has been found on American birds {e.g., Psittacus ; Sfrix perlata).
In the aviary of the Zoological Gardens in London the species has been found on
Shama and Dhyal birds, the insect doubtless having been introduced with some
American bird.
Family PULICWAE.
Genus MALACOPSYLLA.
M(iUi<;,j,siilhi Weycnbergh, ririuilim Zonl. iii. p. 271 (1881) (type : grossiventris).
Miijiipyslh, Baker, .Jnurn. N. York Eiit. Sue. vi. p. 53 (1895) (type : f/rossivciUHs + ar/eiioris).
The genus was proposed by Weyenbergli for Pulex grossirentris Weyenbergh.
This species has not been recognised witli certainty by any of the authors who
have written on the genns MalacopSjiUa. We have a male &nA tfio females ixora
Weyenbergh's collection, and j ossess also types or cotypes of all the other species
hitherto described, and are therefore able to revise the genus. We propose giving
a short review of what has been written about this genus after Weyenbergh's
publications. The first author who had an opportunity of examining specimens of
Malaco},.v/ll<i was Baker (1808). Not being aware that there was a valid generic
name fur these peculiar Siphonaptera, Baker created the generic title Mcgapstjlla
for the same, basing his description on what he called Pulex grossiccnti-is Weyenb.
This grossiventris of Baker, however, consisted of two species, one with a thoracic
comb and one without such a comb. Baker erroneously believed this difference to
be sexual, his female having a comb and his male being without it. We pointed
( 172 )
ont this mistake ia 10O4, describing a comb-bearing species as Malacop.v/l/a
agenoris iind a combless species as anilrodi, adding that these species were
presnmablN' tlie same as Bakei''s. We had some donbts about androcli being
distinct from Weyenbergli's qromtcntrh ; bnt as we conid not be certain from
tlie description whetlier onr insect agreed with grossirentris or not, we thought
it wiser to consider it new rather than to describe it as grosxivciitris, and thns
perhaps render the synonymy of the latter more involved. On now comjiariug
Weyenbergh's specimens with ours, we find that there is no difference whatever
between grossivcntris and aiidrocU.
The year before we {)ublished the descriptions of M. agenoris and androcli
{= yrossiventriis) another MalacopsijUa had been described by Wahlgren as
Megapyslla incrmia. ^\'e have a ])air of this inermis, cotypes, received from
the Stockholm Musenm. Though the author described the head of inermis as
not being armed with a frontal tubercle, we find that the tubercle is present.
We fail to detect any differences between inermis and grossirentris (= androcli).
In a recent paper on American SipLonaptera, Baker enters into the question
of the specific distinctness of agenoris, grossirentris, inermis, and androcli. He
agrees with us that the comb-bearing agenoris stands quite apart, but we do not
understand his statement that what he originally called the male of gro.ssircniris
"turns out to be agenoris"; for in 1898, and again in 1904, it was said to be
the female which had a prothoracic comb. We gather from the remark " a projier
male was found for \\^it female previously called gros.'iiventris'" that the material
referred to as grossivcntris by Baker in 1808 and 1904 consisted of both sexes
of two species. The non-combed species Baker accepts in 1905 to be the true
grossivcntris. The assumption appears to be correct according to Weyenbergh's
specimens before us. He is further right in considering inermis Wahlgren to
be identical with grossirentris. Bnt Baker is in error when treating androcli as
distinct from grossirentris. His opinion is based on two points : the rostrum of
androcli is said by us to reach almost to the apex of the forecoxa, which is not
the case in gro.ssivcntris. Now, we were misled to make that statement by a
mounted /ewff/c in which the labial palpi reacli, in consequence of pressure, beyond
the apical third of the forecoxa. The rostrum of androcli is not longer than that
ot g?-ossiventris. The second distinguishing character on which Baker relies refers
to the genitalia. The finger of grossirentris is said by Baker to be obliquely
truncate, while that of androcli, according to our fignre, " evenly narrows to a
rounded tip," Baker adding that the figure given by AVahlgren exactly " represents
this condition of grossirentris.'" Our figure and
that of Wahlgren are indeed very different, but
the finger in tlie specimens of inermis, androcli,
and grossivcntris is nevertheless the same. We
find our fignre quite correct, except that the finger
is a trifle too long. Wahlgren and Baker did not
see the jjroper outline of the finger. What they
considered to be the ventral outline of the finger
does not belong to the finger. We give here
a diagram taken from a eotyj)e of inermis. Apart from some detail not shown
in our previous figure, it will be observed on comparison that there is no
difference.
In June 19uu a fifth specific name for a MalucojjS>/lla was published by
( ns )
Enderlein for a species supposed to be diiFerent from ffrossifentris (and anJrocU).
The figures accompanyiug the description of tliis M. (ohjpeutii are very good,
although there are some errors in the details — as, for instance, the wrong jrosition
of the dorsal bristles of the hindtibia in the c?, the cylindrical shape of the second
and third foretarsal segments, the absence in the J of a line of separation between
the eighth abdominal tergite and sternite, the number of bristles in the i)ronotum,
etc. The finger is drawn essentially as in our figure (of 1904), but the ventral line
is a little too straight, and hence the ventral distal angle rather too pronounced.
This tolypeutis is again the same as grosskentris. By the description of tolypeutis
our attention has been drawn to an inaccuracy in our figure of the fifth tarsal
segment of grossirentris (= androcli). We described the segment as bearing on
the dilated apical portion a bristle, a spine, and a subajiical bristle ; instead of this
spine there is in the figure a slender bristle. We regret not having noticed the
mistake when correcting the plate.
The result of our comparison of types and cotypes of all the Malacopxi/lla
described is that so far only two species are known to exist : one with a lu-onotal
comb — agcnoris, and one without — grossiventn's ; the synonymy of grotisucntris
being as follows : —
2. Malacopsylla grossiventris.
Pukx grossireiili-h Weyenbergh, Bui. Ac. Nal. Cienc. Avr/ent. iii. p. 188 (1870) ; TascUuub., Die
FlOhe p. 101 (1880).
Mulacopni/lla groxsivenliis Weyenbergh, Periud. Zool. iii. p. 271 (1881) ; Rotbsch., Nov. ZooL xi.
p. go;! (1904) ; Baker, Prw. U. S. Nat. .Uus. xxix. p. 120 (1905).
Sarcoj>s>/lla grossiveiitris, Baker, Can. Eid. xxvii. p. 21 (1895) (partim).
MegnpsijUa fjrosahentns, id., Jount. N. York Eiit. Sac. vi. p. 53 (1898) (partim) ; id., Proc. U.S.
Nat. Mils, xxvii. p. 370 (1904) (partim).
.Mcrjapsijlla iiiermis Wahlgren, Arkiv Zool. i. p. 194. t. 9. fig. 11-15 (1903).
Mulacojisylla androcli Rothschild, Nov. Zool. xi, p. Ii04. no. 2. t. vii. fig. 10, t. viii. fig. II. 12. 14. 15
(1904) ; Baker, Proc. U.S. Nal. JIu.s. xxix. p. 125 (1905).
Malacopsylla tolypeutis ^niex\e,in, Zool. Am. p. I.'i9. fig. 1-6 (1905).
The s]K'cies has been found in Argentina and Brazil on Dasgpus sexcinctas,
cotiurus, and on Canis griseus.
Genus RHOPALOPSYLLUS.
Iihopalop.-<i/Ui,:i Baker, Proc. U.S. Nat. Mus. .xxix. p. 128 (1905) (type : lulzi).
The genus is characterised by Baker as follows : —
" Head without ctenidia, broadly rounded above and in front. Labial palpus
four-jointed. Prouotum without ctenidial spines. Legs stout and thick-set ;
Lindco.xa on innerside without a comb of minute teeth. Female with one aute-
pygidial bristle on either side."
This definition covers a great many Old and New World Pulicidae which are
by no means nearly related to one another. We think a classification of such a
difficult group of insects as the Pulicids should be based on the examination of the
greater jiroportiou of the species so far known, not on a portion of the American
Siphouaptera only, as is practically the case in Dr. Baker's pa])er. However,
no harm is done by the creation of a number of generic terms as long as
tiicy are accomjianied by some kind of definition and the indication ■ of the typo
( 174 )
si)ecies. At onr present state of knowledge of tlie existing forms of Sii)hona]itora
any detailed classification of the fleas mnst be prcmatnre. Wc know only such a
small {lercentage of the species actually in existence that to try to divide the Order
into a series of families and subfamilies must sooner or later prove itself to be but
an abortive attempt.
Dr. Baker, however, in differentiating his JUiOjjalopsyllas from Euderlein's
Parapsi/llus, does so only by the number of the segments of the labial palpus. Wo
are, in fact, presented with a puzzle, botli nomenclatorially and morphologically.
Rhopalopsi/llm is said to be characterised by a "four-jointed" labial palpus, and
Parripxi/H/is by a five-jointed one. Now, the type of lUiajjalopsi/Uatt, namely lutzi)
is described by l?aker as having ap])areutly a six-jointed labial palpus ! Therefore
the position is this : (1) The American Pulex are certainly generically different from
in-itans : (2) The character by which Rhopalojisi/llns is said to be distinguished
from the nomenclatorially older genus ['arapsi/llus does not exist in the species
specified as ty])e.
The confusion has further been intensified by treating, as Baker tloes, all
American Pulex without genal and thoracical combs as having a " fonr-joiuted "
labial palpns, regardless of the descriptions. Now, of the nine species enumerated
liy Raker under Kliopalop»i/Ui(s only one single one has a four-jointed labial palpus.
In our descriptions of coajti, australis, etc., it has been expressly stated that the
labial palpus consists of five segments. We did not mention the rostrum of
cleophofitis in our description. We now add that the labial palpus of this species
has six segments, the uuderlip itself being very short. In one of the specimens the
fourth segment is further divided on the hinder side only, indicating tliat there
may exist specimens of cleophontis with seven segments in the labial palpus.
As we do not know Baker's lutxi, we are not in a position to give a rectified
diagnosis of Baker's Rhopalopsijllus, We also abstain from proposing a new
generic term for any of the eight American Pulex we have before us. The erection of
new genera in this order of insects should be limited as far as possible ; at any rate,
the characterisation of new genera should not be attempted without comparison of
the various allied species from difterent faunistic regions. In this regard we are
in perfect agreement with what Dr. Baker says on p. 123 of his recent paper.*
Weyenbergh described one species of the group of Pulicidae under discussion.
AVe have twofcmales of this Pulex cacicoln, which, on examination, prove to belong to
the species which wc have described in 1904 as Pulex concitus from [.wo females
obtained at Sucre, in Bolivia, the synonymy being as follows :
3. Pulex cavicola.
Pulex cuvicola Weycnborgli, Period. Zmil. iii. p. 274 (1881).
Pulej; condtus Kothschild, Nov. Zool. xi. p. 615. n. 10. t. 10. fig. 38. 40 (1904).
Weyenbergh's specimens were obtained off Cadia leucopijya Bl., while ours
were found on Ilerodon boliviensis. We take the ojjportuuity of supplementing and
correcting the descriptions. The rostrum reaches well beyond tlic trochanter of the
foreleg, the labial palpus consisting of five segmeuts. The third segment of
the maxillary pal [Mis is about one-third the length of the fourth, or a little over
one-third, the fourth being a little longer than the second. The frons of the head
bears a groove near the oral angle, appearing as an incrassation of the skeleton in
• I'm; U.a. Nat. A/us. xxix. p. VS.i (1906).
( 175 )
a side-view (on slide). In this groove, which is present in all the species we
have from America, there is an acute, somewhat heart-shaped, tubercle, which lies,
in most species, usually concealed in the groove, but often projects a little when the
specimen is comjiressed. There is one (not two) long bristle beneath the eye near
the ventral edge of the head.
The fifth segment of the foretarsns is oblong, being hardly half as long again
as it is broad. The basal jjrojectiou of the claw of this segment is very largo, being
at least half tlie length of the claw. There are ten bristles on the outer surface of
the hindtiliia.
We do not know the male.
Pulex lutzi Baker, Proc. U.S. Nat. Mus. xxvii. pp. 378 and 380 (19U4), from
iSao Panlo, off Grisoii. vittata, apjiears to be a closely allied species. The second
anil third abdominal tergites, however, are described as bearing no minute apical
teeth, and the anterior row of bristles on the abdominal tergites 3 to 7 is described
as being represented by only one or two bristles on each side.
Genus CTENOCEPHALUS.
Clciiucrpluihis Kolenati, Fauiut Alirat. p. CG (1859).
This genus comprises only the dog and cat fleas. After having pointed out the
differences between these two insects [see Ent. Rec. xiii. p. 126 (IDOl) ; Xov. Zool.
xi. p. 192 (190.5)] we are rather surjirised to see in Baker's recent paper a note to
the effect that he is convinced that canis and fclis are the same insect. As the
differences in the head of these two fleas are so very conspicuous that they can be
distinguished at a glance from each other, we can explain Baker's contrary opinion
only by assuming that he has seen but one of the two fleas — probably C.feli.i.
Weyenbergh had specimens of both canis and felis. We have received two
specimens labelled as canis. They belong to the short-headed species — i.e. they are
true canis. We have not seen specimens named felis by him in Periudico Zool. iii.
p. 276. No less thaafour of his new species, however, belong io felis, according
to the specimens received ; the synonymy being as follows :
4. Ctenocephalus felis.
Piilcx fclis Bouch^, Nova Arid Ac. Leap. Carol. Gcs. iVal. xvii. 1. p. b05 (183.5).
I'ulcj: ijuniccpa Weyenbergh, Bui. Ac. Nac. Sc. Ai-f/eiit. iii. p. 202. (187'J) (nom. nud.).
Ceratopsijlliis riifulus id., Pcr'Mko Zool. iii. p. 265 (1881) (on C'enius rufus\ " P. parviccps this
species ").
Pulex iiasiKW id., I.e. p. 272 (1881) (on Nama sorkdis).
Pulex ohscurus id., l.c., p. 273 (1881) (ou Canis asarac and ijnuilix).
Pulex concoloris id., t.c., p. 274 (1881) (on Fclis cnnrolor).
As the insect varies a good deal in size and depth of colour, Weyenbergh was
led astray by such individual distinctions. Weyenbergh laid also too much stress
ou the general outline of the body, describing tlie fleas much in the same way as
beetles, for instance, are described. The descriptions accompanying the above-
quoted names contain many inaccuracies, though Weyenbergh apimrently examined
the specimens microscopically. Wliat he calls the comb of the metanotum is of
course that of the pronotum.
^\'e have received ii female of rnfnlus, two females of nasuae, a pair of obsciirus,
( 176)
and II pair t<i' concoloris. All these specimens have the long jioiuteil head oi C. frlis.
'Yhvfcmnh; oi' obsciiruis shows an interesting anomaly, tlie sjiinu at the apex of the
genal process being double on one side of the head.
Family (ERA TOPSYLLICLAE.
CcriiiiipstjUiduc Baker, Pror. U.S. Xal. .!/««. xxix. p. 124 (iyo5).
The bat fleas are so diftereat in organisation from the otiier Siphonajjtera, and
infei- se so much alike, that we agree with Dr. Baker in treating them as a distinct
family. There is only one genns of bat fleas recognised at present. The first
valid name for this genus is Ischnojjsi/llus Westwood. Hitherto Curtis's name
Ccratopsi/llus — altered into Ceratopsi/lla by most authors (philologically perhaps
a correction, but nomenclatorially a misspelling) — has been employed instead,
Kolenati being usually cited as author of Ceratopsi/llus. As nobody appears to
have looked up the literatnre on this point, accepting without i)rotest the erroneous
nomenclature for the bat fleas, we quote here what Curtis wrote about Ceratopsi/llus
and Westwood about hchnoj/syllus in 1833, 1838, and 184U. Criticising Curtis's genus
Ceratoplii/llus in Ent. Mag. i. j). 359 (1833), Westwood states that Cfratfipln/lliis
can scarcely be considered to be well founded, the distinctions based on the antennae
uot holding good. And he proceeds to say, on page 3G2 : " The species, however,
figured by Mr. Curtis, C. elongatu.i, as well as C. tespertilionis, and probably
C. bifascintus and Ptth'x muscull Dng., together with a Chinese species, which has
been kindly presented to me by the Rev. Leonard Jenys, exhibit a general form so
difiereut to that of the other fleas, that I cannot help thinking them, on that account,
entitled to form a distinct group, for which (as the name Ceminphi/llus must
likewise be rejected in consequence of having been previously employed in botany)
the generic name Lichnopsijlbis may not be deemed inajiplicable ; the characters of
which 1 propose to detail in a memoir, upon which I am at present occupied, upon
Bat Parasites." This promised memoir was never completed, only a monograph of
the genus ^'ijcteribia appearing in 1835 {Trans. Zool. Soc. LoiuL). But we find a
more precise statement of what Ischnopsijllm was meant to stand for in Westwood's
Introd. Classif. Ins. ii. p. 124 (1840). There we read :
" Obs. — Ceratopsyllus Curt. (Ceratophyllus Curt., B.E.) consists of species in
the type of which the antennae are inserted on each side of the head, concealed in a
cavity behind the eye when at rest, and as long as the head, four-jointed"; but other
si)ecies introduced into the genus are described by Curtis as diflering entirely in
this respect, hence the character derived from the antennae appears to me to be
merely specific. Some of the species, however (P. respertilionis and elongalus)
being of a much more slender general form, I have proposed for them the generic
name of Ischiiopsi/llus {Ent. Mag. No. 4)."
Meanwhile (1838) Curtis had altered the original spelling of the name
Ceratophyllus into Ceratopsyllus ; this explains the appearance of " Ceratopsyllus
Curt." in the above (jnotation from Westwood's book. In volume xv. of the
British Entomology Curtis says,* under Errata and Addenda : t
"Folio 417 for Ceratophyllus read Ceratopsyllus. This name, wjiidi was
compounded to express the peculiar structure of the horned fleas, was misprinted
* Jiritisk Entomology xv. Indijx p. 2 (1838).
t In a letter dated Sept. 2Gtb, lUUl, Mr. CJ. A. Verrall drew my alteiitiou lo Ibis note.— N. C. li.
( 177 )
when tkt; gt'ims was established in this work, and it was not corrected, as the
author intended, in the Guide."
From these quotations it will be clear (1) that Ceratopsijllus was merely a
well-meant correction of the term Ceratoplnjllus; (2) that Ccratopsi/llus, thervi'oret
is an absolute s}'nonym of CeratophijUiis, the type being stated by Curtis to be
C. hinmdiitis ; * and (3) that the correct name for the bat fleas is Westwood's terra
Iscknopsijllus.
Dr. Baker gives GeratopsijUtis priority over [schnopsyllus, dating the former
erroneously 1832. He quotes in Proc. U.S. iVat. Mks. xxvii. p. 432 (l'JU4) as
follows : —
1832. Cei'utopsi/llus Curtis, Britisk Eiitomolof/i/ x.
1833. Ceratopsyllus AVestwood {Ischiopsi/llus), Eiit. Mo. Mag.., I. p. 359.
Both quotations are wrong. There is no reference to a bat flea in British
Entomolwjij x., which appeared in 1833, not 1832. Westwood in the Entomological
Magazine (not Ent. Mo. Mag., the first volume of which was pul>lished in 1804-5)
does not mention the term Ceratopsgllus, which, as said above, dates only from
1838.
Weyenbergh described one species of Iscknopsglliis :
5. Ischnopsyllus isidori.
Ceralophyllus isiduri Weyenbergh, Period. Zool. iii. p. 271 (1881).
The species is closely allied to I. irolff'solud Rothsch., Nov. Zool. x. p. 321. n. 5.
t. '..I. fig. 13, t. lU. fig. 14. 15. 16 (10U4). The frous of the head is rather longer
and move pointed ; the posterior row of bristles of the metanotum of the c? is
normal, being similar in the sexes ; the metanotum and first abdominal tergite are
devoid of the two short spines standing in woljfsohni on each side at the apex of
these segments, etc.
Found on Vespertilio isidori Gerv.
* Haker, Proc. U.S. yat. Mit.'<. xxix. p. 129 (11)U5), erroneously makes gaUitiar the type of
C't'rtittiji/iijllu.'i.
12
( 178)
NEW SPniNGIDAE.
By the HON. WALTER ROTHSCHILD, Ph.D., and KARL JORDAN, Pu.D.
1. Protoparce fosteri spec. uov.
(J. Close to P. lichenea Bnrm. (1856). No sliarply defined line on mesothoracic
tegiila. Abdomen beneath pnre white, mesial dots black ; side-patches of tergites
white, large. Pulvillns of claw-segments present, bnt small.
Wings, iippt'rsidc. Forewing narrower than in both lichenea and Jlore^tan ;
the interspaces between the black lines purer white ; hairy patch at base restricted,
more extended white; black longitudinal discal streaks 11^ — M^ rather longer than in
lichenea ; first and second discal line heavy, merged together as in lichenen, inter-
space between second and third line white from costal to inner margin, the tiiird
line crenate, well marked ; interspace between third and fonrth lines buffisii white,
especially the posterior portion of this interspace much purer huffish white than
in the allied species ; fifth line widened at costal margin to a large triangular
patch ; interspace between this patch and the oblique apical line huffish white.
Hindwing more elongate than in lichenea ; white from base to first band,
except a large patch between cell and SM-, this patch brown-black, contiunons with
the first band ; interspaces between the three browu-black bauds white posteriorly,
sharply defined, the white colour gradually shaded over with brown anteriorly, but
the interspace remaining distinct to costal margin.
Underside. Forewing deeper black-brown than in lichenea ; cell more
washed with grey ; two rather distinct brown-black lines on disc, the interspace
between them and a band at the distal side of the second line greyish white,
distinct; submarginal area also more extended grey than m Jloredan vmCl lichenea.
Hindwing : greyish-white ; a heavy black line touching cell, curved basad on
abdominal fold, more proximal behind than the corresponding line of the allied
species ; just outside this line a second one, dentate, rather thin, joining the first
line at M' ; a third line farther distad, thin, partly obsolescent between the veins,
very strongly dentate ; marginal baud well defined, about 5 mm. broad in middle,
j)aler brown-black than first line ; interspace between the marginal band and the
third line about as wide as the baud.
Tenth tergite broader vertically than in the iillied species. Harpe in shape
similar to that of lichenea and Jiorestan, rather smaller, emargiuate ventrally,
its edge slightly irregular, non-dentate. Tooth of penis-sheath longer than in the
allied sjjccies.
Length of forewing : 04 mm.
Ilah. >Sapucay, Paraguay, January S2, I'JUu (^V. Foster).
One <f .
2. Eui'yglottis albostigmata basalis sulispcc. nov.
Eiu-i/gloitis albostigmala Rothschild & Joid., Xui: Zuul. ix. Supijl. p. I'S (IWKt) (partim ;
S.E. Peru).
? . We received this form when our revision of the Sphingidae was in press.
On comparing again typical ? ? of albostigmata we now find that the Peru
(ll'j)
individuals dill'ur iii thu hindwiDj? being all white from the liase to the first discal
line, the costal area excepted.
flab.- S. Domingo, Carabaya, S.E. rem, OUOD I't., March I'JOl (G. R.Oekenden).
Two ? ? .
3. Protambulyx xanthus spec. nov.
r?. Ujipi-rside : forewing as in P. eurijcles Herr.-Sch. (1854), but posterior
subbasal patch narrower, more oblique. Hiudwing : very pale chrome, jialer
than in euri/df» and ciiryalus; lines as in curi/alus R. & J. (1903), but the first
liue broader, standing closer to cell, and the marginal band narrower.
Undeiside deej) maize-yellow ; discal lines faint, except the first which
is strongly marked on both wings ; marginal band of forewing as in euri/cles,
extending to hinder angle as in that species, being much wider between R' and R-
than in eunjalas, and tapering to a point, ending at tip of SC\
Eighth sternite with short obtuse mesial lobe. Patch of friction-scales on
clasper obliquely transverse, widest dorsally, similar to that oi curi/cles but longer.
Length of forewing : 6U mm.
Hab. Tnis, Costa Rica.
One (?.
Perhaps a northern form of curijclcs. Easily distinguished from euri/clcs by
the third line of the hindwing above being curved as in e'ln/aln.s, suJpkurea, and
astij(joniu (see Sovi. Zool. ix. Sujipl. p. 176 ii'.), differing from the latter insects
especially in the marginal band of the forewing being shaped as in cunjcUs.
4. Compsogene panopus celebensis subspec. nov.
S ¥ . Forewing, above : interspace between subbasal band and patch of cell-
bars narrower at costal margin than in Malayan and Indian specimens, the proximal
bar of that patch more straight and heavier ; brown marginal border symmetrical,
evenly tajiering at both ends, the black line bordering it non-dentate, the olive
proximal border of this liue much wider and much more evenly rouuded ; black
discal band more oblique, almost parallel to margin, crossing subcostals distally of
base of SC°.
Underside : the proximal suljajiical costal spot larger, the line extending from
this spot to inner angle less distinct or vestigial ; marginal band almost evenly
rounded, being much less strongly narrowed from R- forward and backward.
Hal). Tondauo and Sawangan, North Celebes.
Two pairs.
6. Polyptyclius anochus spec. nov.
c?. Upperside ot palpus, head and thorax grey, of abdomen bnffish grey;
underside pale vinaceous cinnamon ; tibiae grey on upperside, the grey streak of
midtibia sharply defined, interrupted before middle. Legs, jjalpus, and antenna
similar to those of F. andosa Walk. (1856) iu structure.
Wings, upperside. Forewing: apex strongly produced, distal margin deej)ly
concave below apex, convex behind middle, inner angle projecting backwards ;
pinkish grey, markings clayish vinaceous cinnamon ; a broad antcmedian band
consisting of several lines with the interspaces filled in, narrower behind, about as
broad as the grey median interspace ; this intersxiace denticulate, being bordered by
( 180)
,a lliiii brownish crenate lino wliicb cnrves costad ; the whole area between this
line ami distal margin clayisli vinaceous cinnamon, bearinj); a row of ill-defined and
rather indistinct grey patches which are situated jiroximally of a second discal
crenate line ; this line extremely faint, being accentuated by vein-dots ; one small
subbasal dot ; stigma a small ring with grey centre. Hindwing pale salmon-
buff, greyish at anal angle, indistinctly marked with brown ; anal angle more
produced than in P. andosa.
Underside clayish salmon-buff ; two brown lines on disc "of each wing, jjavallel,
denticnlate, curving costad.
Neuration : SC- and R' of hindwing on a very short stalk ; D'- almost four
times as long as D^, very oblique, sliglitly angiilate ; D'' a little shorter than D^ ;
lower angle of cell acnte.
(Jenitalia : tenth tergite very different from that of the other sjiecies of
Pohjptychm in being completely divided into two very slender, pointed processes,
whicli stand widely separate; tenth sternite broken, apparently broad, rounded-
truncate. Clasper with elongate-ovate friction-patch dorsally ; no friction-scales on
inner side of eighth tergite ; clasper strongly narrowed apicad, ending in an acute
point which is bent downwards ; harpe with two processes ; one ventral, reversed,
beak-shaped, being carved upwards and pointed ; the other much longer, irregularly
triangular, obliipiely longitudinal, tapering, somewhat irregular, curved inward. No
distinct armature on penis-sheath.
Length of forewing : 28 mm.
Uab. Sierra Leone.
One S.
Similar in colour to P. andosa, compar, and consimilis (see 2^oc. Zool. ix. Snppl.
p. 250), but widely different in the genitalia. Apex of forewing much more
jiroduced.
Libyoclanis gen. nov.
?. Palpus much slenderer than in (.'lanis and Pseudoclanis* segments 1 and 2
nearly equal in length, the joint between them not open. Antenna cylindrical.
Tibiae spinose at apex ; si)nrs much shorter than in allied genera mentioned ; two
jiairs to hindtibia, the short spur a little shorter than tlie tibia is broad, half the
length of the long one ; cell of hindwing trnncate, both the upper and lower angles
being about 90' ; SC- and R' on a short stalk.
Antevaginal ridge asymmetrical, produced into a process on each side, the
processes being unequal in size.
Type : L. bainbridgei gen. nov.
Here belongs also Clanis bicolor, from Sierra Leone,t which we left
provisionally in Clanis when revising the Sphingidax.
fl. Libyoclanis bainbridgei spec nov.
?. Body tawny olive above, wood-brown beneath, sides of abdomen somewhat
creamy, with an ill-defiued white patch at base, upperside of tibiae and tarsi
muminy-brown, stiff hairs at tiji of abdomen yellow.
• See lievision of Sjihingidac, in .Vof. Zvol. is. Kiippl. p. 220 (1903).
t L.C. p. 219.
( 181 )
Wings, upperside, Fore wing produced at apex into a prominent lobe, which is
about 5 mm. long, measured from tip of Sf'° ; greenish clay-colonr (more green when
alive ?) ; two faint straight oblique parallel lines, greyish, the second a little beyond
base of M'^ the first about (i mm. distant from it, these lines being the proximal
borders of indistinct clay bands, a pale band across cross-veins, from costal to inner
margin, widest between R' and W, including a small transparent stigma bordered
with brown ; an oblique brown line from apex to R', ending here iu an indistinct
brown patch ; a small brown subapical patch at l)end of costal margin preceded
by a cloud of glossy grey scales ; a small rosy red basal spot at inner margin. ■
Hindwing rosy red, costal margin creamy white, distal margin tawny olive, this
marginal band gradually widening behind and becoming paler, extending along
abdominal margin to base, the hairs at base being creamy white.
Underside greenish buff, paler than upperside, creamy towards base.
Forewing rosy red centrally from base beyond apex of cell ; a brown line from
apex obliquely beyond R-. Hindwing with large rosy red patch along abdominal
margin ; two greenish yellow lines across disc, at R' the distances between these
lines, cell and distal margin, about equal ; R^ from centre of cell.
Vaginal ridge raised into a triangular tooth on each side, the right tooth being
longer and broader than the left one.
Length of forewing : 67 mm.
Ilah. Sierra Leone (Major Bainbridge).
One ?.
We name this fine species in honour of the collector, from whom we have
received several interesting captures.
7. Pholus vitis fuscatus subspec. nov.
Phnlns vilh vith, Rothschild & Jord., Noe. Zunl ix. p. 494. n. 418. a. (1903) (partim ; St. Vincent ;
Grenad.a).
Having received a series of fresh specimens, we now propose a separate name
for the form from the Lesser Antilles.
?. Wings, upperside : pale markings of forewing more or less washed with
brown, therefore less contrasting with the greenish olive-brown markings than in
nearly all Continental specimens ; subbasal band not distinctly edged with white ;
brown line situated within oblique grey discal band less distinct ; marginal band
broadm-, not edged with white.
HkI). Santa Lucia (S. Branch), ti/pe ; Grenada ; St. Vincent.
A series of ? ? .
8. Deilephila placida salomonis subspec. nov.
DeiUphila placida placida Rothschild & Jord., Noi\ Zool. ix. Suppl. p. 512. n.430. a. (190.3) (partim;
Guadalcanar and Florida).
c?. Blore uniformly olive, the markings less prominent ; subbasal patch of
forewing, upperside, hardly deeper olive than the ground-colour ; median band less
oblique than in pi. placida ; its proximal edge less curved.
Hab. Solomon Islands; N. Georgia, tijpe ; Choiseul; Guadalcanar; Florida,
4 <Jcf, collected by A. S. Meek,
( 182)
9. Temnora angulosa spec. nov.
?. Body rnssct-brown above, tawny-olivo beneath. Wings, iijipersidc :
forewing rnsset-brown, tliis area sharply limited on disc, indented on the veins,
dilated before and behind li', reaching posteriorly to inner angle and costally a
little beyond apex of cell ; disc ontside the rnsset-brown area fawn-colour, apex
washed with mars-brown ; a thin brown line ontside the basi-discal area, nearly
following the curve of the outer edge of this area, which it joins behind M';
distal margin biconcave, strongly aiignlate at K-, the angle somewhat nmnded.
liindwing mummy-brown, entire.
Underside tawny olive, washed with mumm3'-brown. Forewing mummy-
brown in centre from base to disc ; a faint line on disc ; an equally faint line
obliquely from apex, bordering a faint marginal band which is slightly deeper
brown than the distal portion of the disc. Hindwing with a fiiiut denticulate
line on disc, curved like distal margin, but standing farther away from it in front
than behind.
Length of forewing : 32 mm.
Ilnh. Luluabnrg, Kassai, Congo Free State, September 1902 (Landbeck).
One S.
10. Xylophanes fosteri spec. nov.
S ? . Head and thorax olive above, with a pinkish grey side-stripe extending
from tip of palpus to apical edge of mesothoracical tegula, edged with white
behind, a brownish grey mesial streak on thorax only ; a central line on tegula
tawny ; abdomen wood-brown above, greyish at base, clayish towards apex ; this
dorsal area bordered by an olivaceous line which somewhat widens basally ; three
indistinct dorsal lines ; underside creamy bufif in middle, buff shaded with tawny
at sides. Outer spur of midtibia half the length of the inner.
Wings, uppcrside. Forewing wood-brown shaded with sepia-colour ; apex
acute, distal margin entire, concave below apex ; seven lines between cell and distal
margin, first and second from inner margin to R^, heav}', more or less merged
together, the second being continued costad, but the upper portion very thin, third
line thin, fourth accentuated by minute vein-dots, the line distinct from R' to apex
of wing, fifth also distinct from R^ to apex, indistinct from R' backwards, the npper
]iortions of these two lines more or less contiguous, sixth feeble, seventh vestigial,
lines 4 to 7 converging to tip of wing, lines 2 and 3 not reaching costal
edge, disapjiearing in a pale Wdod-brown costal subapical space; a small black
stigma ; ontside apex of cell a brown cloud which is continued behind R- to
lines 4 and 5; a small blackish costal dot distally of subcostal fork, this dot
being the costal portion of the first line. Hindwing blackish brown ; a dirty
pinkish band on disc from anal angle to costal edge, which it does not (juite reach,
palest behind, being more or less washed with brown in front.
Underside pale ochraceous, with a faint tint of pink, irrorated witji wood-brown.
Forewing brown centrally from base beyond apex of cell ; a row of conspicuous
black-brown vein-dots parallel to margin, curving costad in front, between this row
and the cell a more or less vestigial line ; distal marginal area pale wood-brown,
this border angulate at R^, reaching here the line of dots. Hindwing: a wood-
brown line at lower angle of cell, jiarallel to distal margin, curving costad in front,
the lino broad Imt rather faint ; distally of this line and parallel with it a vestige of
( 183)
a second line and then a row of small bnt distinct dots ; distal marjjinal l)and
wood-brown, tapering behind, brown behind M-, the veins being more or less of the
ground-colonr.
Tenth tergite sinuate at apex, steruite long, with rounded apex. Harpe short,
slender, tai)ering, with the apex curved upwards; a few very minute teeth at tip.
Penis-sheath with an oblique row of teeth, the central portion of the row situated
on a slightl)^ elevated ridge, the portions at the right situated at the edge of an
obtuse short process. Friction-scales large.
Length of forewing : cj, 30 mm. ; ?, 33 mm.
Hab. Sapucay, Paraguay, February and October 1903 (W. Foster).
One pair.
Near A', turbala Edw. (1887) and robiimni Grote (1865).
11. Xylophanes dolius spec. nov.
S. Body as in fostcri, pale buff beneath. Outer spur of midtibia less than half
the length of inner.
Wings resembling in shape those of A', hi/drata R. & J. (1903). Upperside
of forewing wood-brown, washed with olive, lines in the same position .as in
A', hjdrata, fourth thin, deeper brown in upper half than the others, fifth vestigial
between R- and M-, ending anteriorly in a brown spot situated at apex ; costal
marginal area paler !it apex than rest of wing ; a rather large black stigma.
Hindwing : black-brown, with a narrow buff band from an.al angle to costal margin,
which it does not quite reach, being shaded with brown anteriorlj'.
Underside pale buff, somewhat pinkish on disc, irrorated with brown.
Forewing black-brown centrally from base beyond apex of cell ; a rather heavy line
just distally of lower angle of cell, somewhat incurved behind M', thin in front,
ending in a costal dot ; parallel with this line a row of dots, the costal dot being
situated halfway between first line and apex ; a marginal spot below tip of wing,
produced discad into an oblique line ; a heavy marginal double spot M' — SM- ; all
these lines and spots deep brown. Hindwing : a carved line and a row of dots
on disc ; distal margin brown.
Clasper with about seven large friction- scales. Harpe very slender, curved
upwards at apex, tapering, denticulate distally. Tenth tergite and sternite long, the
latter with almost parallel sides and rounded apex.
Length of forewing : 31 mm.
Hob. Zamora, Ecuador (0. T. Baron).
One i , somewhat faded.
12. Xylophanes cosmius spec. nov.
$. Body as in A. amadis Stoll (1782) ; basal lateral patch of abdomen less
distinct, quite gradually fading away ; abdominal tergites irrorated with brown
scales, without line or dots. External spur of midtibia one-third shorter than
inner.
Wings, vpperside. Forewing : apex produced, distal margin obtusely
denticulate, deeply concave below apex, strongly convex in middle ; wood-brown,
washed with green, costal marginal area, a triangular space at outside of discal
line from U- to apex, and inner margin more distinctly green than rest of wing ;
three greenish brown lines in basal half, curved, not reaching inner margin, outer
two close together ; a black stigma, outside which there is a greenish black patch,
( 184)
transverse, rounded distall)-, straiglit ]iro.\imiilly ; ii very conspiciioiis line fmin
tip of wing to middle of inner margin, continued along the latter to base, apically
more curved than in amadix, crossing W at one-third the distance from cell to
outer margin, greenish black, edged with grey proximally ; traces of two lines
proximally of it, these additional lines accentuated at costal edge by a spot each ;
a submargiual row of black vein-dols ; fringe black at veins. Hindwing black,
a greenish buff band from anal angle to costal margin, which it does not reach,
slightly incised at the veins ; distal margin narrowly green ; fringe sj>otted black
at posterior veins.
Underside ochraceons salmon-buff, irrorated with brown ; markings as in
amadis, distal marginal band rather wider ; lilack fringe-spots conspicuous on
both wings.
Tenth tergite feebly spatulate, truncate ; steniite triangular, with the apex
rounded. Seven large friction-scales on clasper. Harpe slender, tapering, curved
upwards at apex faintly denticulate at tip. Penis-sheath with a broad streak
of numerous minute teeth on left side, the convex area on which this streak is
situated rounded at apex and at right side produced into an obtuse process, which
is as broad as long, being covered all over with teeth and being directed proximad.
Length of forewing : 4U mm.
Ilrib, La Union, llio Hnacamayo, Carabayn, S.E. Pern, ^Oiio ft., November
1904, wet season (G. R. Ockenden).
One <J.
13. Xylophanes chiron lucianus snbspec. nov.
Xylnphanes chirnn chiron, Rothschild & Jord., Nov. Zool. ix. Suppl. p. G99. n. Go2. b. (1903) (partiiu ;
Sta. Lucia).
Having now seen several specimens from Santa Lucia, we find that this island
is inhabited by a special subspecies, which, though similar to the one found in
Jamaica, differs in several points.
?. Upperside of body and forewing green, as in ordinary Continental individuals
of A', chiron chiron ; a brown or black mesial patch anteriorly on mesonotum.
External spur of midtibia a little shorter than internal.
Wings, njjperside. -Forewing broader than in the Jamaica and Continental
forms; a green vestigial line from apex, proximally of this line a row of transverse
black-brown dots on the veins from SC'' to M-, followed at inner margin by a
triangular black-brown patch as in the other forms, these vein-dots partly connected
with one another by vestiges of a brown line ; no trace of a clayish patch either
at costal margin or on disc. Hindwing with three spots on disc, a fourth,
anterior spot being faintly indicated ; the anal and subanal spots as in the
other forms.
Underside similar to that of the Continental subspecies. One heavy line on
disc of forewing, the proximal line of the other two subspecies not being marked;
brown proximal edge of marginal band heavy, the costal intersjiace between this
band and the discal line wider than in tlie other forms. Hindwing : a brown line
on disc accentuated by vein-dots, a vestige of a second line close to cell ; marginal
band as broad as in the Continental subspecies.
Hub. Santa Lucia (S. Branch).
Several ? ¥ .
( 185 )
14. Xylophanes chiron cubanus subspec. nov.
Xylophanes chimn nechtm, Rothschild & Jord., !.<■. p. G'J8. n. Cul.a. (1903) (partim ; Cuba).
Fresh material from Cuba shows that there fire several jioiuts of difference
between Cuban and Continental specimens.
c??. Smaller than the (,'ontinental form; antenna pinkish; nnderside of
l)ody more rnfous.
AV'iugs, ujjperside. Forewing proportionately shorter, the costal and discal
clayish patches much smaller. Hindwing : discal spots smaller, huffish.
Underside: on the whole more extended rufous ; the costal discal luteons
patches of the forewing more restricted, the latter patch replaced by a dot.
Hub. Holiiuin, Cuba (Tollin ; Parish).
A small series.
15. Xylophanes pyrrhus spec. nov.
(??. Closely resembling A', thijeha L. (ITHs). Much larger. Forewing,
ahove : the pale band bearing the double line wider than in thi/clia, the line
bordering this band distally gradually widening from tip of wing to R^ ; distal
margin less rounded than in thi/elia.
Underside. Forewing : with three lines across disc, the first heavy, curved
in front, reaching costal margin abont 4 mm. beyond subcostal fork, the second
and third thin, close together but separate, curving costad in front, more proximal
than in thyelia, the third being at iSC° abont 3 mm. distant from oblique apical
line ; posterior portion of third line slightly dentate, tlie luteons streak standing
at its distal side more irregular than in tliyeVm and thinner. Hindwing :
proximally of the double line a vestige of another line.
Length of forewing : 33 to 37 mm.
Hah. 8. Domingo, Carabaya, S.E. Peru (G. U. Ockenden), a long series ;
Merida, Venezuela (Briceno), one S.
Wo have a series of A. thyelia from various places, the range extending from
S.E. Peru to Colombia, and eastwards to the Guianas and Trinidad.
( 186 )
NOTES ON BAT FLEAS.
By the Hon. N. C:. ROTHSCHILD, M.A., F.L.S.
I.
IN 1850 Kolenati described and figured an eight-combed flea which he had off
various species of bats. The figure represents a c? bearing a luaue of long
hairs on the thorax. This species, named by Kolenati octactenus, is commonly
found on the Contiueut and in England on the pipistrelle {Vcspcrugo i>ipistrellus),
whicli is one of the hosts mentioned by Kolenati.
AVhen Wagner, in L'^OS, published the result of his examination of the
Siphouaptera contained in the Imperial Museum at St. Petersburg, where some
of Kdlenati's sjiecimens are preserved, he made a mistake in respect to octactenus
Kolen. Wagner found in the collection a ? of an eight-combed bat flea from
Kolenati's collection. This ? he considered to be a typical specimen, which it
may be. Now, Wagner had in his own collection two closely allied species of
bat fleas — namely, a pair of one species and a single c? of the other. The pair
Wagner identified with the above-mentioned ? from Kolenati's collection, while
he described the single cj as a new species under the name of C. jubata. In this
he was wrong. His jubata is the maued species figured in 1S.56 by Kolenati as
octactenus.
Following Wagner, we have in some {)revious papers recorded the true
octactenus (with the maned S) as jubata, and the other species as octactenus, both
occurring in England. This second species being witliont a name, wo now propose
to call it
Ischnopsyllus simplex,
the synonymy of the two insectfi being as follows : .
1. Ischnopsyllus octactenus.
Ceratopsyllus oclacleniis Kolenati, Parnx. Chirojit. p. 31. t. .3. fig. 31 (1857) ; Rothsch., Ntir. Znul. v.
p. 543. n. G. t. 14. figs. 7. !> (1898).
Cemtopsijlla jubata Wagner, IIoi: Sue. Ent. Ross. xxxi. p. 584. n. 4 (1898).
2. Ischnopsyllus simplex spec. nov.
CeralopnijUa ndactcna Wagner («o« Kolenati), I.e. p. 580. n. 1 (1898).
The differences between the c? c? of this species and the preceding have been
ably described and figured by Wagner. The ? ? of the two species are best
recognised by the difference in the length of the apical bristles of the metathoracical
epimernm. Tliis plate bears in /. octactenus Kol. one long and two short bristles,
while there are in /. simplex one long and one short bristle.
The I'uleu- eespeitiUonis of Duges (1832) is an eight-combed bat flea whicli
may be identical with one of the later-described species. But Ungcs's description
is quite insufficient for recognition.
( 187 )
II.
In 10(i3 wc (lescrilied two North American bat fleas as Cerntopst/lla h)m'(//v's
from Ontario, and ('rratoj/si/Ua paljiosKS from British Columbia. Dr. Carl Baker,
in his recent jiaper,* has added a third, named Ceratops'/llus croshyi, from Missouri.
This insect is shortly described in a " key " to the American bat fleas. As this
"key " is misleading, we hope Dr. Baker will not take it amiss that we make the
following comments. The insect is characterised by Dr. Baker as follows : —
" Ctenidia on segments I — VII of abdomen ; metatarsal article I with spines
7 — 8j and this segment as long as segments II, III and IV togctlier ; mesonotum
nearly twice the length of metanotum on the dorsal line."
Now, taking the characters in the same order as in this short diagnosis of
crosbiji, we find that in the description of insignis the abdomen is stated to bear
seven combs of modified bristles. The mesonotum of insignis we figured as being
about twice the length of the metanotum. The tarsi were not described, as they
did not present any striking differences from the other American species. On
re-examination we find that the hindtarsus of insignis agrees with Baker's
description of the hindtarsus of croshiji. Therefore the diagnosis of croshyi as it
stands in Baker's paper offers nothing which does not also apply to insignis. Baker
himself differentiates insignis from croshyi by erroneously attributing only fonr
abdominal combs to insignis instead of seven, as stated in our description. It
is qnite possible, however, that croshyi is not identical with insignis.
III. — A New Bat Flea from Italy.
Ischnopsyllus gestroi spec. nov.
This species is allied to /. martialis Rothsch.,t but differs in many essential
points.
Thorax. — There are fewer bristles on the meso- and metanotum than in
I. martialis, there being no bristles immediately above the pleural plates. The
metanotum and the first and second abdominal tergites bear one short apical
spine on each side. The metathoracical epimernm has five or six bristles
(2 or 3, 2, 1).
Abdomen. — On the abdominal tergites 2 to 6 there is one bristle below the
stigma and another above it, and two more bristles on each side on the back some
distance from the one standing above the stigma. In front of the dorsal bristles
there are one or two small hairs on each side. The seventh tergite bears one
or two more bristles on the back than the preceding segments do. There is
one long apical bristle on each side of this tergite, accompanied in the ? by two
small hairs, there being in the S .a row of short apical hairs in between the
two long ajiieal bristles.
Legs. — The mid- and hindcoxae bear posteriorly at the ape.x one long and
one very short bristle. The mid- and hindfemora have two ventral and two lateral
bristles at the apex on the outer side, there being, besides, on the hindfomur
five short bristles situated in the basal fourth. The midtibia has on the outer
side one lateral row of six bristles, and on the inner side only one or two bristles.
The hindtil)ia bears a lateral row of eight or nine bristles on the outer side and
* rnic. U. S. A'at. .Viis, xxix. p. i:i7 (1005).
f A'm: Zunl. x. p. 322. n. fi. t. 10. fig. 17-20 (190:!).
( 188 )
a row of six or seven on tlie inner. Tliere arc only one or two sniiill luiirs
at the ventral edge of the mid- and hindtibiae, apart from the apical bristles.
The bristles of the tarsi are mnch less numerous than in /. martialis. The iirst
midtarsal segment bears four or five short bristles on one edge and five or six
at the other, while there is one single row of six hairs on the ventral surface,
and a row of three or four on the dorsal side. The first hindtarsal segment has
also only one row of hairs each on the dorsal and ventral surfaces.
Modified segments. — 6. Tbe eighth tergite is sinuate at the apex, the edge
below tlie small sinus being somewhat undulate (Fig. 1, viii. t.) There are about
nine bristles between the stigma and the apical edge. The eighth stcrnite
(Fig. 2, viii. st) is truncate, bearing a row of four bristles on each side veutrally at the
Fig. 1.
Fig. 2.
ajjex. The clasper (Fig. 1, CI) is oblong, as in 7. martialis, being widest at the
apex ; its upper angle is strongly rounded, while the lower angle is somewhat
produced, bearing a pair of long bristles. The finger (Fig. 1, F) is about two-
and-a-half times as long as it is broad, being of nearly even width throughout.
It bears a number of bristles at and near the apex, which is rounded. The
exact lengtli and number of these bristles we cannot clearly mahe out from the
single mounted specimen we have.
?. The seventh stcrnite is distinctly but slightly emarginate veatrally at
the apex. The eighth tergite resembles that of I. martialis. Tliere are ten stout
bristles on the outer side and no short hairs. On the inner side the row of
five or six short spine-like bristles found in mai-tialis is represented in the new
species by one or two bristles only.
Dr. R. Gestro, director of the Mnseo f!ivico at Genoa, sent us two pairs of
this flea taken ofi" Kyctino7nus ccstonii at Genoa. Dr. Gestro has kindly given
us one pair.
I
( 1«9 )
TWO NEW SATURNIIDAE.
By THE HON. WALTER ROTHSCHILD, Pii.D.
1. Dysdaemonia fosteri spec. nov.
? . Body cream-colour ; antenna, head, palpns and legs chestnut.
Wings, upperskle, cream-colour, olivaceous green in distal area, basal area
feebly washed with green. Forewing : lines and bands green ; a straight line
from costal to inner margin at base of M-, obtusely angulate at SM- ; a shadowy
band outside this line ; two transparent spots at cross-veins, the upper one small,
both with green proximal border; a band from costal to inner margin, touching
second transparent spot, slightly curving proximad between R' and SM-, sharply
defined proximally, gradually shading off distally ; a line between this band and
distal margin as in horeas, but a little more curved, brown behind ; hinder angle
of wing more rounded than in boreas. Hindwing : transparent spot encircled
with green ; a minute green dot at upper angle of cell ; two lines on disc as in
boreas, both obtusely angulate, but the angle of the first line more obtuse than
the angle of the second; first line green, broad, gradually shading off distally,
widened to a band behind, second line green iu front, brown behind ; outside the
second line a brown band irrorated with grey from tail backwards, as in boreas.
Underside cream-colour, washed with green ; two lines on disc of each wing,
both faint.
Hab. Sapucay, Paraguay, January 7, 1905 (W. Foster) ; one ? .
2. Bunaea macrothyris spec. nov.
?. Body hair-brown, thorax and abdomen densely irrorated or washed with
ochraceous, anterior part of mesonotum, collar, forecoxa and head almost uniformly
brown, collar edged with ochraceous ; antenna russet.
Wings, iipperside, hair-brown, densely irrorated with ochraceous. Forewing :
basal area bordered by a blackish hair-brown line angulate-sinuate upon M and
again at SM- ; along this line a broad white band gradually shading off distally,
the white scaling extending at costal margin to submargiual line ; a large trans-
parent spot, rounded distally, straight proximall.v, lO mm. long, thinly edged
with black, this ring followed by a broad red ring which is widened to a half-
cresceut proximally, bordered externally by a pinkish white line which is regularly
elliptical iu shape, being broader proximally than distally ; distally of this spot
a broad blackish brown line from costal to inner margin, very distinct within the
white costal area, becoming faint farther back, evenly curving costad in front,
slightly angulate between M- and SM- ; a second blackish brown line parallel
to distal margiu from costal edge to II-, being hardly vestigial I'arther baok^
8 mm. from distal margin at 11'. Hindwing : base washed with ochraceous ;
an autcmediau line white, widening behind ; a white line parallel to distal margin,
1 1 mm. distant from margin, edged with blackish brown on distal side ; area
between the two white lines without ochraceous irroration ; distal marginal area
( 190 ;
densely irrorated with ochraceons ; eye-sj)ot a little loujjer tliau broad, 13 luiu.
long, consisting of a minute snbtransjiarent bar, a black central sfiot of G mm.
diameter, a broad orange-red ring, edged ontside with a pinkish white line.
Underside : both wings densely irrorated with white from base to submargiual
lino, the veins being more or less ochraceons ; distal marginal area blackish browni
irrorated with ochraceons. Forewing : jiosterior area hair-brown from base to
discal line ; eye-spot as above, the black line bordering the transparent spot
heavier ; blackish brown discal line tajiering behind, distinct, except from M-
backwards. more jiroximal between li' and M' than above, not standing distally
of eye-spot, bnt being interrupted by it : area between discal and submargiual
lines slightly washed with maroon between SC and M'. Hindwing : a broad black
line from costal to abdominal margin, crossing eye-spot, couvex ; eye-spot much
smaller than above, consisting of a black central i)atch bearing a semi-transjiarent
bar, a red ring edged with pinkish white, the ring much narrower than above
and of the same deep red colour as in forewing ; wool at abdominal margin
yellowish ; interspace between discal and stibmarginal lines whitish at abdominal
margin.
Length of forewing : tS2 mm.
Ilab.: CJiissamba, Bilic, Angola, November 10, I'Ji.)4 (Ur. W. 1. Ansorgc);
one ?.
28 JUL 1906
NOVITATES ^OOLOGICAE
Vol. XIII. JULY, 1906. No. 2.
NEW NOCTUIDAE PROM ERITISH NEW GUINEA.
By G. T. BETIIUNE-BAKER, F.L.S., F.Z.S.
IN continuation of my tli'scriptions of the new species of Mr. Pratt's collections
from the Owen Stanley IJange, and of a portion of Mr. Meek's collections,
I am now able to add those of the Noctuidue, omitting the last sectiofi, viz. the
Deltoidinae, which I propose to deal with later on. The number of new species
sent us by Mr. Pratt is beyond our most sanguine expectations. Mr. Meek's
collections have alreaily received their well-earned praise. Each liave in their
own lines of work done magniticeutly, and both these well-tried collectors are to
be highly congratulated on the resnlts of their expeditions. I have again to thank
Sir George Hampson for constant help. My measurements are taken by doubling
the distance from the centre of the thorax to the tip of the primary.
I mnst make a correction in iny previous paper. I was informed at the last
moment, after the proofs had been all corrected, that there was no description for
Fig. 42, PI. VI., vol. xi., and in the absence of all papers, being away from home,
I wrote the description of No. 108, p. 429 (Colli/sa ekeiltei) ; the insect had,
however, been described on p. 4u3 as Anthela ekciliei, and this description and
generic name is correct; the descrijition on p. 429 sinks to the earlier one, and the
legend on Plate VI. for Fig. 42 should be Anthela ehcikei*
NOf'TUlPAE.
1. Amphipyra bicolora spec. nov.
(J. Head and collar very dark ]inrj)lish brown, collar tipped broadly with
paler brown ; thorax and abdomen brown, legs very hairy, ])ale brown with dark
tarsi jialely ringed. Primary with two or three dark basal dots followed by a short
curved costal line, a spot at the end of the cell laterally edged with darker brown ;
three strongly serrate dark postmcdial lines close together, followed by a sub-
terminal interrupted irregular line; termen darkly and finely dotted. Fringes
brownish grey. Secondary with the basal two-thirds yellow : terminal third dark
lirown, slightly iridescent at certain angles. Undersurfaco : primary uniform brown
with a dark cell-spot ; secondary as above.
? . Like the male, but larger.
Exjianse : i 44—46 mm. ; ¥ 48 — .^)il mm.
The type is in my collection. The sjiocies flies from January to March.
Huh. Ekeikei and Aroa lliver.
2. Protagrotis rufalis spec. nov.
i . Palpi pale reddish, head darker, antennae pale buff-colour, collar reddish buff,
thorax reddish brown with bright red patagiae darkly edged, abdomen pale yellowish
* The naniu on the lal)L'l of tlie spccinu-n fiiiuitnl was CnUiiAa /■krilici. Tlie name t'heih'i recurring so
frequently in Mr. Baker's article. I did not suspect thai Ctilhatf rhrikei was the same as Anthfla ehnhci. —
K.J.
13
( 192 )
brown, forelegs red with darker tibiae palely rinajed, mid and hind pair ])ale bnft'-coloiir.
Primary brightish red up to the postmedial line, dark reddish grey beyond ; ante-
medial line double, the enter one the darker, botli interrupted at the subcostal vein,
beyond whicli is a small jiale red round spot darkly but finely eneiroled ; outside tliis
a band of dark shading curved from the costa to the lower margin of the cell,
whence to the inner margin it is obliquely straight ; beyond this the reniform
stigma shows slightly grey in a paler red area, which is sharply margined by a
double postraedial irregular subdcntatc line, the rest of the wing being reddish-
brownish grey, iu whicli is the siibterminal obscure recurved row of dark dots,
which have an obscure trace of a reddish external edging; termen iiah-, fringes
dark. Secondaries jiale yellowisli grey wifh the terminal area pinkish. Underside
of both wings quite jiale, with tiie posi marginal area darker.
Expanse : 40 — 41 mm.
Ildh. Ekeikei, where the species flies from January to Ajiril.
Type in my collection.
3. Protagrotis novaguinensis spec. nov.
c?. Head greyish brown, face pale ochreous grey, palpi dark brown, the scales
at the tip of the second segment ])alely edged, third segment i)aler with a dark
patch below ; collar brownish grey, thorax darker ; patagiae brownish grey with a
dark central stripe from the shoulder, below whicli is a broad pale stripe ; abdomen
pale greyish, the last segment but one dark brown, the neighbouring segments
being also suffused with that colour. Primary greyish brown, darker along the
costa to the lower margin of the cell, an obscure pale stripe extending from near
the apex, where it rises in a small pale apical patch, to the inner marginal area,
which is also slightly paler than other parts of the wing; a snbbasal jiatch of
raised i)ale scales from the costa to the inner margin of the coll, antemedial line
broken and obscure, postmedial line irregular, strongly incurved towards the end
of the cell, beyond which is a curved row of dark spots ; termen darkly dotted ;
orbicular stigma obscure ; reniform evanescent (in pale specimens, especially
females, both these stigmata are moderately distinct); cilia pale with a darker
central line. Secondary uniform dark brownish grey, somewhat lustrous witii
paler fringes.
Undersurface : Primary dark grey with a broadish reddish costa and a very
broad whitish inner margin. Secondary whitish, with costa slightly irrorated with
reddisii ; apex and termen blackish brown, broad at the apex, but rapidly tapering
to vein 1.
?. Like the male, bnt the primary ochreous grey above ; in all other respects
just like the male above and below.
Expanse : c? 44 — 4(1 mm. ; ? 40 — 44 mm.
JIab. Ekeikei, January to April.
Tyjie in my collection.
This species differs somewhat inter se ; the males are sometimes paler, and the
females slightly reddish. 1 have one specimen of the latter of a decided reddish
ochre colour.
Epa gen. nov.
cJ. Head roughly haired ; eyes smooth, lashed ; antennae finely, very shortly
ciliate ; palpi npturned, end segment minute, deflexed, not reaching the vertex of
( 193 )
the head, first ami second seyiui'iits clothed witli hug hair, with mctatliovMcic
crest ; abdomen crested ; legs with the tarsi of each spined for all their len^jth :
niidtibiae with one pair of spnrs, hindtihiae with two pairs of spnrs ; femora and
tiliiae of all legs densel}- haired.
Nenration : Priraar\' with veins 3, 4, and Ti from close to the lower angle of the
ceil, vein C> from below the upper angle, 8 and '.) stalked from a verj- short stalk
close to tlie areole. Secondary, 5 from near the lower angle, G and 7 stalked from
well beyond the cell.
Type : Epa priitii ]'>-T?.
4. Epa pratti s|.ec. imv.
c?. Frons greenish yellow, capnt greenish grey, collar and thorax greyish
green, jiatagiae edged and bisected with black, abdomen cream-colonred. Legs
and femora yellowish grey, tibiae blacdvish grey, tarsi chestunt-red ; the femora and
tibiae arc clothed with longish greenish-grey hairs. Primary with a basal and
snbterniinal snil'nsion of yellowish green, the mediiiii area pale dirty gre)' np to the
u|ii)er margin of the cell ; costa and snbapical area black ; termen pale grey, basal
line ronghly U-shaped, from the costa to the lower margin of the cell ; beyond the
greenish sntlnsion and extending into it a broad black jiatch, which is invaded by
the grey median area ; beyond this a tine donble slightly iiLterrnj)ted black median
line, followed by a similar postmedial one, edged externally with white ; this line
is very irregnlar and strongly projected ontwards above vein 5 ; a qnadrangnlar
deep black patch at the end of the cell, a snbterrainal pale line in the greenish area,
four white jioints on the costa in front of the apex. Secondary whitish grey, termen
broadly dark grey. Underside of both wings creamy greyish with terminal area
dark grej'. In the primary the terminal area is invaded by a pale line, and in the
secondary there is a dark postmedial line.
Expanse : S 30 mm.
lUih. Dinawa, September.
Tyjie in my collection.
Yula gen. nov.
Head rough, vertex with short projecting tnft of liairs, antennal sockets fringed
with longish hairs, antennae shortly and finely ciliated ; ])alpi nptnrned, almost
erect, first and second segments thickly scaled with hairs, end segment short,
slightly deflexed ; eyes smooth- lashed, proboscis fnlly developed ; legs with mid-
til)iae with one pair, hindtibiae with two pairs of spnrs, all the tarsi strongly spined
tlu; whole length, fore- and niidtibiae with a tnft of hairs below the femoral joint ;
thorax with a central prothoracic tnft, a pair of tufts and a central tnft nn the
metathorax ; abdomen with the proximal segments tufted.
Tyjie : Yuln noracgvinme B-B.
•1. Yula novaeguineae spec. nov.
cJ. Frons grizzled brown, vertex creamy white, jialpi dark reil-brown with end
of second segment ringed with whitish, end of third segment tipped with whitish;
collar metallic brassy bronze, j)rothoracic and metathoracic tufts bronzy chestnut,
patagiae shining creamy, slightly bronzy, edged with dark brown with brassy
scales intermixed ; abdomen dark brown, with the proximal segments laterally
creamy whitish ; anal segments browni.sh cream-colour ; legs dark brown ringed
with whitish. Primary with a general metallic bronzy lustre all over, varying from
( IW )
silvpry and pale greenish into dark steel)- lirown. Basal area pale greenisli with
blackish interrupted fine basal line followed by a large blackish costal patch into
the cell, below which, extending as far as the cell, is a shining silvery oblong patch
intersected by the fine medial dark line ; jiostiuedial area dark bronzy brown, with
a small round pale greenish sj)0t at the end of the cell followed by a largish silvery
somewhat irregular sqnarish patch ; the i)0stmedial excnrved crennlate line defines
this area, and is followed by a narrowisli paler line, apical patch dark bronzy brown,
tornal area, and extending n]i to vein S dark bronzy brown ; terminal area jiale
greenish cream-colour scjiarating the dark patches just mentioned ; termeu very finely
black intersected at tlie veins. Secondary brownish, becoming cream-coloured at
the base.
?. Like the male in the primary, but witli the dark parts blacker; in the
secondary the wings are blackish instead of brownish.
Expanse: c?, 31— 33 mm. ; ?,3.'5-38mm.
Ilab. The type is in my collection from Ekeikei. I have specimens also
from the Kebea Range, Diuawa, and other localities, where Mr. Pratt collected.
The species is, however, rare apparently, only a few having been taken in each
jilace. It flies in March and April and again in duly and August ; but there is no
difference in colour in the two broods.
C. Perigea dinawa spec. nov.
S- Head, palpi, and thorax reddish brown, abdomen paler ; legs reddish
brown with darker tarsi palely ringed. Primary reddish brown, with a subbasal
whitish costal point followed by a second below it, beyond which are two more
whitish points below each other; an indistinct pale waved anfemedial line, beyond
which in the cell tlie orbicular stigma is palely but indefinitely visible, reniform
distinct pale, with whitish dots : jiostmedial line pale, strongly curved outwards,
obscurely darkly edged laterally ; snbterrainal line distinct, composed of a series
of indefinite whitish spots almost confluent, the central ones edged darkly
internally ; termeu darkly seallojied between the veins, where it is intersected
by whitish points ; costa with two whitish dots, one nearly over each stigma,
beyond which are three finer jmiiits nearer the apex. Secondary pale uniform
reddish brown.
Exj)anse : 37—39 mm.
Ildb. Dinawa, September; Kebea Range and Aroa River, March and April.
Type in my collection.
T. Perigea aroana spec. nov.
(J. Head and thorax mottled greyish, abdomen greyish. Primary jiale grey with
dark grey irrorations all over, a small white costal basal spot, an antemedian white
waved line ; reniform whitish, ]ireccded by some black nuirkings ; a postmcdial
whitish band interrupted l)y the veins; a fine sulitcrniinal dentate irregnlar line,
edged internally in fresh specimens by blackish short dashes. Secondary white
with broadish dark-grey termen.
Expanse : 30 mm.
Ilab. Aroa River, March and April.
Type iu my collection ; other sj)ecimeus in the Tring Museum.
This S])ecies is somewhat near 1'. confundens Wlk.
( 195 )
8. Euplexia viridacea spec nov.
(?. Heiul ochreoiis, collar greenish, thorax and abdomen reddish ochreous,
jiatagiae greenish, legs brownish oohreons with tiiu tibiae darkly spotted, tarsi
darkish, palely ringed. Primary pale bronzy green, with a dark snbbasal patch
edged with white, an antemedial white line edged Hnely with black ; medial area
dark browiush, with an irregnlar wliitc; M-sliaped mark below vein ID, with
a greenish spot between the two outer lines of the letter ; below vein 2 a white
inverted V-shajied mark ; this dark area is edged with a fine cremdate doable line
of black and white ; terraen darkly spotted, preceding which (spots) is a series
of spearliead marks finely ontlined with white ; along the costa are three dark
dashes with fine white centres, two being before the end of the cell and one
beyond the cell. Secondary with tlie basal three-fifths greyish with a dark
hinule at the end of the cell, the terminal two-fifths pale bronzy-green. Under-
snrface : Primary ochreons grey tiiiged with green, with a dark spot in the cell
and a dark postmedial line ; secondary ochreous with a dark spot at the end of
the cell and a dark postmedial line. In fresh specimens the dark patches of the
primary are covered with a greenish Instre.
Ex]ianse : 33 — 30 mm.
Hub. Dinawa, Ekeikei and the Kebea Range.
Type in my collection.
9. Euplexia smaragdina spec. nov.
S. Frons ochreous slightly tinged with green, palpi ochreous, head pale green,
collar and thorax bright green, patagia witli two grey stripes, abdomen ochreous.
Primary pale emerald green, with a short costal basal blackish diagonal dash with
a fine white internal edging; below this a second similar dash invading the cell,
a blackish snbbasal patch interrupted by the veins from the costa to well below
the cell edged externally with a y -shaped white line, a curved line of green below
it, followed by two blackish dashes, each edged internally finely with white; latter
half of cell covered with a blackish irregular subijuadrate patch, edged externally
with white, in the centre of this patch a fine diagonal ^-shaped mark ; reniform
stigma beyond the cell marked out with white, beyond which is a wedge-shaped
dark brown jiatch ascending to the costa, where its broad end is, a dark sjiot on the
costa in front of the a]iex, confluent with this ; below, nearer the base, between
veins 1 and 2, a black dash edged externally with white, a terminal series of black
spear-head marks increasing towards the tornus.
Secondary golden yellow without marks. Undersurface of both wings ochreous
yellow.
Expanse : 35 ram.
Hab. Kebea Range, March and April.
Type in my collection.
111. Euplexia dinawa spec. nov.
Head and frons ochreous, collar dark olive-greenish, thorax dark brown with
dark olive-green patagiae and tufts, abdomen greyish with orange dorsal tnfts, legs
darkish grey with reddish tarsi. Primary brown, with a small dark lirown basal
patch bisected by a pale <-shaped mark, a dark brown spot below the cell, outer
half of cell occupied by a broad dark brown stripe in which is placed a broad
r i9(i )
U-shaped mark ; at tin- fiid of llu' diirk striiH' llio |';iler iiostrneilial line extends
riy;bt across tlie winj; ; Vieluw tlie green mark are two small dark spots; terminal
area pale lironzy green edged internally with a broad tapering patch of dark
velvety brown, tinged with bronzy green ; termcn dark brown, inner margin dark
brown tinged with green with a green spot a third from the base. Secondary
brownish grey, paler at the base. Fringes of both wings green. Undersnrface :
Primary brownish grey, witli a snbterminal row of small jiale <l(its ; secondary,
basal half pale grey with a dark spot in the cell and a broad dark very serrated
postmedial line, outer half i^lnsely irrorated with dark grey.
Exjianse : 34 mm.
Ilah. Dinawa, September; Avo'a, August; Aroa IJiver, .lanuary ; Kelna
Range, March and April.
Type in my collection ; other sjiecimeus in the Tring Museum.
A rare species, the series sent home being very short.
1 1 . Em'ois dinawa .sjiec. uov.
(?. Head and thorax pale ochreotis brown, collar witli a dark line across tlie
middle, abdomen reddish ochreons, leg.s chestnut-colour with dark tarsi palely
ringed. I'rimary j)ale ochreons brown, with a fine basal line in the cell and on
the costa, a double medial line from the inner margin into the cell, the inner
line being obscure, strongly serrate, forming a double v; be3-ond this is a dark
patch edged partially by a dark line, reniform stigma pale with a white spot on
its exterior, and a dark v-shaped ]iatch on its interior ; above this the costa is
darkly spotted, postmedial line only visible i'roni the inner margin to vein :i;
from the lower jiart of the reniform a double dark dash extends to the termen.
Secondary pale greyish with a darker terminal area.
Expanse : 42 mm.
llnh. Dinawa, August.
Type in my collection.
12. Ancaroides kebea spec. uov.
cj. Head, paljii and thorax dark rich red-brown, palpi pale chestnut interuiilly,
thorax and patagiae mottled with still darker velvety brown and interspersed with
pale spatulate scales ; on the prothorax is a central tnft of raised dark hairs and
scales; abdomen chrome yellow with the terminal segments dark brown and a
series of dark dorsal tufts on the ycdlow segments, legs dark brown encircled with
j)ale chestnut-colour. I'rimary dark reddish brown with a large darker brown
patch below the cell and another occupying the tornal and terminal areas up to
vein 5 ; the basal and antemedial lines represented by irregular scries of dark dots,
the orbicular and reniform stigmata distinct, finely encircled with pale yellow,
])Ostmcdial line distinct, double, acutely angled externally above vein .i,froni whence
the double line is filled in with pale yellow and margins in an arc the dark tornal
area ; from the reniform stigma a fine pale chestnut line runs above vein o to
the termeu, a subterminal curved line extends from vein 5 to the toruus, below
the apex is a patch void of the large rough scales that cover most of the wing
(this ]]atch is in some specimens pale rusty chestnut-colonr), the costa is darkly
dotted where all the lines commence. Secondaries pale chrome yellow with very
broad brown ai)ieal and terminal areas tapering rapidly towards the tornns ; a dark
sex-patch nearly fills tlie s]iaee between veins 2 and :i, invading well into the yellow
( 1!'7 )
area. Underside : Primary brown, inner margin broadly pale yellow, cell closed
by a dark spot beyond which are somewhat obscure dark postmedial and sub-
ferminal lines. Secondary as above, but paler, with a dark spot closing the cell,
and the sex-patch pale yellow invading into the dark terminal area.
Exp.'inse : ;")(! ram.
Hab. Kebea Range, Jnly; Ekeikei, January and February; Anja Hiver, March.
Type in my collection.
i;^. Ancaroides griseola spec. nov.
(J. The [iriniaries (litter from A. kvhi:,i. in that liolh stigmata and the area
below them are filled in with jiale grey, there is no angled postmedial line, there
are no dark patches on the wings, whilst the subterminal line is fairly distinct
from near the apex to the tornus, and is strongly serrate ; in the secondaries the
dark border extends somewhat up the inner margin.
E.xpanse : 56 mm.
Hah. The type is in my collection, and was taken at Ekeikei in March
or April.
This may possibly be a variety of the j)revions species, A. hehca, but the
large grey area and the difference in the subterminal lines give it quite a distinct
appearance, which coupled with the fact that I have several from this one neigh-
bourhood makes me think it advisable to name it.
14. Dipterygia kebeae spec. nov.
S. Head ochreous grey ; collar dark grey, palely irrorated ; thorax jiale greyish,
with the jirothoracic tuft jiale ochreous brown ; patagiae blackish ; abdomen dark
grey. Primary blackish, with the costa broadly dark grey, increasing slightly near
the end of the cell, with' three pale dashes beyond the cell ; inner margin greyish,
broadly greyish for the posterior third and at the tornus, where it is invaded by
the black area; from near the base is a broad palish dash finely margined with
black. Secondary white, with the apex and termeu broadly blackish brown^
decreasing rapidly at the tornus.
? . Like the male, but not quite so dark.
Expanse : S and ? , 44 mm.
Hab. The species occurs at Mount Kebea and Ekeikei in March and April :
a long series was taken by Mr. Pratt ; other specimens are in the Tring Museum
from the Aroa River, taken at the same time.
The type is in my collection.
15. Dipterygia babooni spec. nov.
S. Head and thorax ochreous grey ; collar intersected with two dark lines
irrorated with white-ti]iped scales ; abdomen brownish grey. Primary ochreous grey,
with a trace of a subbasal strongly serrated fine line, the postmedial line having
an outward curve at the end of the cell, and then receding in a waved nearly
perpendicular line to the inner margin; reniform stigma pale whitish grey, below
which from the costa is a dark obliijue stripe having a deep curve iu the post-
medial area, but ascending to the middle of the termen ; a white dash below the
costa in the apical third ; from near the base of the inner margin is a broadish
pale dash darkly margined. Secondary white, broadly margined witli hrowiiish,
tapering towards the toruus.
( 198)
?. like the male, bnt with a subterruiiial very stroingly serrated line edged
outwardly with whitish.
Expause : S and ? , 44 mm.
Ilab. The type from Babooni is iu my collection ; other .specimens in tlie Tring
Museum. The sjiecies occnrs in September, and 1 have a good series from all the
localities tliat Mr. I'ratt collected iu. I believe this insect is a variety of the
previous one, and that it will prove to be the antuiun brood of J>. Iieheae ; bnt it is
80 distinct in general coloration that it is obviously advisable to name it.
Hi. Acronycta crenulata spec. nov.
S. Head and thorax irrorated pale and dark grey, abdomen greyish. Primary
pale grey, with a dark, double, obliipie, obscure iu parts, antemedial line, and a
blackish finely crenulate postmedial line, edged internally with wliite, (lutwardly
curved above vein 3, below which it recedes in a deep curve to the inner margin,
lieyoud this is an indistinct stripe of pale shading ; orbicular a whitish distinct
round spot, reniform rather obscure, between these two a dark costal patch.
Secondary pale grey, darker at the termeu.
E.xpanse : 42 mm.
Hab. The type is in my collection from the Kebea Range, where it was taken
in July.
17. Acronycta ekeikei spec. nov.
S . Head and thorax whitish grey, finely irrorated with pale green ; abdomen
grey, rapidly deepening into reddish brown ; legs pinkish grey, with blackish tarsi.
Primary pale whitish grey, more or less finely irrorated all over with pale green ;
a small dark costal patch near the base followed by a dark dot, a dark costal jiatch
above the end of the cell extending into a green spot closing the cell, two small
costal spots, followed by a longer dash on the costa in front of the apex, a trace of
an antemedial irregular whitish line, and a faint trace of two pale postmedial lines,
the outer one dotted white ; a broad blackish dash below vein 2, distinctly black
along the vein edged with white above, which (white) curves down nearly into the
tornus, intersecting the black dash which extends to the termen ; the postmedial
area beyond the cell-spot is darker along the costa to the apex and down to the
tornus than the rest of the wing; termen finely black. Secondary reddish brown,
whitish and somewhat diaphanous towards the base. Undersurface : Primary
lustrous reddish brown, white below the cell and vein 2, with a trace of a dark
postmedial line. Secondary whitish with costa and apex broadly reddish brown,
and a distinct red-brown irregular postmedial stripe from the costa to the inner
margin.
Expanse : 38 mm.
Ilab. The type from Ekeikei is in my collection. The species flies in March
and April.
18. Acronycta dinawa spec. nov.
S ■ Head and thorax pale olive green, abdomen pale brownish tinged with red,
anal extremity greenish, legs dark red-brown, jialcly spotted. Primary pale olive
green, costa spotted with black, the spot above the end of the cell large and
extending down to vein .^i ; a short black basal dash along the lower margin of
the cell and a black dash along vein 3 to near the termeu ; a trace of an antemedial
( 199 )
line, and a more definite trace of a curved black dotted postmedial lioe; terminal
area mottled, fringe crenulate. Secondary reddish brown, }ialer towards the base.
Uudersnrface : Primary red-brown, whitish below the cell and vein 2. Secondary
cream-coloured, with the costa and upjier part of termen and npox broadly red-brown,
a dark sjiot at the end of the cell, and a very irregular j)ostmedial line to vein 1 h.
? . Like the male, but with the pattern and especially the black dashes aii<l
spots decidedly more definite and rather larger. Secondary uniform reddish brown.
Expanse : c? 37, ? 4'i — 43 mm.
Hah. The type is in my collection from Diuawa ; other specimens from the
Kebea Range. The insect flies in May and June.
10. Ilattia aroa spec, no v.
$. Head, thora.x and abdomen dull brown. Primary dull brown, with a trace
of a basal and antemedial dark line rising in a white costal dot ; a white costal spot
over the white reniform, the latter being pupilled with an ochreous sjiot darkly
encircled ; postmedial line very tine, finely crenulate, a subterminal irregular line
of dark shading from a white apical spot, three white costal points in front of the
ape.v, termen with fine black points. Secondary uniform greyish brown, slightly
paler at the base.
Expanse : 32 mm.
Hnh. The type is in my collection from the Aroa River ; other specimens in
the Tring Museum from the same locality. The species will come next /. renalis
Moore.
20. Berresa rufa spec. nov.
S. Head and thorax pale reddish brown, abdomen greyish. Primary pale
reddish brown, with an' indistinct curved dark antemedial line across the shining
sex-mark, an obscnre dark median band meeting the dark palely edged postmedial
serrate irregular line about vein 2, where the latter is decidedly incurved ; a dark
irregularly waved subterminal line ; termen with tine creamy points. Secondary
pale brownish grey.
Expanse : 29 mm.
Hah. The type is in my collection from the Aroa River, where it occurred in
March.
21. Berresa meeki spec. nov.
5. Head and thorax ])ale piidiisli grey, abdomen grey. Primary uniform
j>urplish dull brown, with obscure excurved antemedial and less curved median dark
lines ; postmedial line white, sharply crenulate, a trace of a pale subterminal line ;
costa dotted with white. Secondary sooty grey, very dark at the termen.
Expanse : 23 mm.
Ilab. The type from the Aroa River is in my collection, other specimens in the
Tring Museum from the same locality being taken in March.
22. Thyria aroa ^pec. nov.
6. Head and thorax sejiia brown strongly and finely irrorated with pale greyish
brown ; abdomen yellow with dark dorsal tufts. Primary sepia brown with pale
basal stripe and a patch of pale greenish scales ; antemedial line fine, pale, evenly
excnrved in the fold with a pale costal fine dash to the base of the orbicular, which
( 200 )
is dark and palely encircled ; reniliinu olisciiic wilh ;i {nitcli of |iak' lironzy greenish
rongli scales above and liclow it ; jiDstmcdiu! liin' [lale wuvod and scalloped, a rich
dark broad siibtermiiial hand with three whitish cxtcinal spots by the apex ; teriuen
darkly dotted. Secondary yellow wilh very broad brownish terminal area.
Expanse : 3S mm.
Ilah. Aroa Kiver (March).
Type in Tring Museum.
23. Dissolophus ochraceus spec. uov.
S ■ Head and thorax ochraceous, abdnmen oclircoiis grey. Primary [lale oehreons
grey, with a trace of a dark basal line, an irregular waved autemedial line rising in
a small dark costal patch ; a postmedial pale line gently waved with dark edging
followed by an irregular line of dark shading ; a trace of a snbterminal serrate line ;
termen darkly dotted. Secondary pale greyish brown.
?. Like the male, but the secondary is dark brown witii ochreous fringes.
Expanse : 25 mm.
Ilah. The type is in my collection from the Aroa River, where the species
occurs in February and March. Other specimens are in the Tring Museum from
the same locality. The species will come next 1>. nluensis Btl.,. but the shajie and
position of the antemedial and postmedial lines differ.
24. Elusa dinawa spec. nov.
(J. Head reddish pink ; thorax red with jiinkish chestnut collar ; thorax
ochreous. Primary red, with two whitish yellow spots at the end of the cell, the
lower of which is oval and larger than the uj)per one ; the basal, antemedial,
medial, and postmedial dark grey lines are somewhat irregular ;ind confluent, the
postmedial line being the broadest and suti'ased, and the wing is spotted between
each of the lines with pale yellowish spots, one at the apex being more prominent
than the others ; the termen is dark red, almost crimson red. Secondary pinkish
brown, quite pale towards the base.
Expanse : 30 — 31 mm.
Ilab. The type is in my collection from Dinawa, where the insect flics in
September.
25. Elusa iucertans spec. nov.
?. Head and collar jiinkish chestnut; thorax reddish; abdomen brownisii
grey Primary red, with antemedial, medial, and jiostmedial lines of dark grey.
Secondary pinkish brown.
Expanse : 30 mm.
Hab. The type is in my collection from Dinawa, where the insect flies in
Angust ; other specimens in the Tring Museum from the Aroa River.
26. Elusa pratti sjiec. nov.
S . Head and thorax reddish brown ; abdomen pale brown. Primary brownish
with a very slight reddish tinge ; antemedial and postmedial lines dark grey and
irregular, medial line broader, somewhat indefinite ; a twin white spot at the end
of the cell ; a snbterminal row of dark dots. Secondary uniform jiale brownish.
?. Exactly like the male, except that it is decidedly darker, and has no tinge
of rufous colour.
( 201 )
Ex|iai]se : c? atid ? , 32 — 33 mm.
Jhih. Tlie tyjie is in my collcctinii from Diiiawa ; other sjieoimciis from
Ekeikei. The species flies in September ami in April.
27. Caradrina subpartita spec. nov.
S. Head, tliorax and abdomen cinnamon-brown, finely irrorated witli grey.
Primary cinnamon-brown with fine dark lines ; basal line only discernible on the
costa; antemedial line witli two sharp serrations [irojected inwardly; medial lirn;
broad, indefinite, obli(ine from the costa to the end of cell, then vertical ; postmcdial
line crennlate, followed by a band paler than the gronnd-colour ; snbterminal line
irregnlar ; termen finely dark, interrnpted at tlie veins. Secondary ])aIo whitish
grey, subhyaline darker at the apex.
Jjxpanse : 30 mm.
IJah. The typo is in my collection I'rom the Keliea Range, where it flies in
March and April ; other specimens in the Tring Musenm from the Area River.
This species is near C. paiiita AV^alker ; but the lines differ, especially the
median line in its strong angle.
2s. Caradrina unipunctata spec. nov.
S . Head and thorax dull brown, abdomen slightly darker, legs pale greyish
with dark tarsi palely ringed. Primary narrow, dull brown, very slightly lustrous ;
antemedial line fine, black, irregular and obscure, medial line dark curved outwardly,
postmcdial line very slightly waved then straight to the inner margin, pale
distinct, darkly and obscurely edged on each side, snbterminal line dark waved
palely edged externally, a small dark sjwt above vein 5 between the postmcdial
and subterminal lines, termen very finely darkly dotted. Secondary snbhyaliue
grey.
Expanse : 33 mm.
Ilah. The type from Ekeikei is in my collection.
29. Caradrina funerea spec. nov.
S . Head and thorax dull brown with a tinge of red, abdomen j)aler. Primary
rather narrow, dull slightly reddish brown ; a fine dark antemedial line, a band
of dark shading at the end of the cell from the costa to the inner margin, a small
l)ale ochreous dot at the end of the cell with three whitish points below it jilaced
as it were at the three jioints of the triangle, a very tine dark crennlate post-
medial lino followed by a line of paler shading, a subterminal row of fine irregnlar
dark points, termen very finely jiale. Secondary subhyaline pale grey, except at
the apex and the termen, where it is darker.
Expanse : 30 mm.
llab. The type is in my collection from the Kebea Range. The species flies
in May and June.
3ii. Caradrina insignifica spec. nov.
S. Head and thorax greyish brown, abdomen paler. Primary greyish brown,
with a dark fine irregnlar antemedial line, a dark medial line of shading produced
forward by the lower margin of the cell, a postmcdial pale line sliglitly curved
outwards, an irregular somewhat serrate subterminal line of dark shading edged
( 202 )
by a paler external line, terminal area ]ialer. Secondary snbhyaline, whitish grey,
darker at ajiex and teriueii. On the niulersnrf;ue of the secondaries, the costa
is ochreous grey finely darkly irrorated ; there is a distinct trace of a iiostmediul
line.
Exjianse : 30 mm.
//«/;. The type is in my collection from the Kebea Ran^e, where the species
flies ill May and Jnne ; other specimens are in the Tring Museum from the Aroa
River.
31. Caradrina gemini sjicc. nov.
S ■ Head, thorax, and abdomen pale grey, pal]ii pale grey with a dark brown
spot on the first and lower half of second segment. Primary pale grev, very
finely but sparingly irrorated with dark grey, giving the wing a rongh appearance ;
three dark points on the costa, two near the base, one nearer the apex, all the lines
obsolete, a dark brown twin spot in the postmedial area, divided by vein 4 ; termen
with a distinct dark point at the end of each vein. Secondary nniform pale grey.
Exjianse : 33 mm.
J/ah. The type is in my collection from Ekeikei, where the species flies in
March and April.
32. Caradrina crenulata spec. nov.
(?. Head and thorax ochreous grey, abdomen pale grey, palpi grey with a dark
spot on the first and basal three-fifths of second segment. Primary pale ochreous
grey, with a fine dark antemedial line with three lunulas between the costa and the
lower margin of the cell, a medial grey band of shading receding basewards above
the lower end of the cell with two whitish dots at the base of veins 3 and 4, and
a pale small spot above them, the postmedial line fine, sharply crennlate, the
crennlations ending in a blackish point, terminal area greyish nearly up to the
postmedial line, a dark point between veins 3 and 4, two blackish spots divided
by vein 5, above which are traces of three black spots to the costa; termen slightly
crennlate, very finely dotted with blackish between the veins. Secondary, sub-
hyaline brownish grey, slightly darker at the apex and termen, veins showing dark.
Uudersurface, both wings ranch paler than above, with the postmedial line
marked out by a series of fine distinct blackish dashes right through both wings.
Expanse : 37 — 38 mm.
Ilriij. The type is in my collection from the Kebea Range, where the species
flies in July.
33. Caradrina pratti spec. nov.
(?. Head and thorax greyish brown, abdomen pale grey. Primary greyisli
brown, antemedial dark line waved, jireceded by a broadish band of jiuler groiuid-
colour, medial band of dark shading indefinite and angled by the cell, at the upper
angle of which are two i)ale dots, the lower one being shifted inwards, postmedial
dark line curved, an indefinite waved subterminal lino. Secondary brownish grey,
with a large tuft of dark brown longish hairs on the underside of tlie inner margin ;
the thorax also has a similar heavy lateral tuft below.
Expanse : 31 — 33 mm.
J/aO. The type from Dinawa is in my collection, and was taken in August. I
have a specimen taken in Sej)tember of the same year which is much paler in
colour and has a distinct dark sjiot near the cud of the cell.
( 203 )
ACONTIINAE.
34. Dipthera kebeae ^pei'. nov.
S. Frons pinkish, slightly tiugecl with green, with three horizontal dark
spots ; palpi reddish brown, palely ringed at each segment, end of third segment
pale. Head and thorax pale yellowish green, patagiae darkly tipjied and darkly
spotted behind the collar, tuft darkly tipjiod ; abdomen pale greyish brown with
slight dorsal tnfts darkly tijiped, anal tnf't jjale ; legs dark reddish brown palely
spotted at all the joints, tarsi ])alely ringed. Primaries yellowish green with a
short very dark irregnlar dash from tlic base along the lower margin of the cell,
three dark small costal spots near the base and funr nearer the apex ; between the
two series of spots is a large dark mark extending across the end of the cell,
in front of the apex a short diagonal dark line on the costa ; between veins 4 and (5
are two whitish spots ontwardly edged with dark brown followed by a similar pair
nearer the termen ; between the lower margin of the cell and the turnns is a dark
brown irregnlar mark liisectcd and edged ontwardly with whitish, witli a whitish line
between it and the basal dash ; inner margin darkly dotted. Fringes intersected
greenish and dark brown. Secondaries creamy white, more or less snft'used with
pale pinkish brown for the terminal half.
Undersurlace, both wings creamy white. Primaries snlFnsed with pinkish
brown along the costa and in the medial area, whitish along the inner margin ;
secondaries snfFnsed with pinkish brown broadly along the costa and in the apical
area, with a dark costal patch towards the base and a cnrved dark interrnpted
postmedial band.
Expanse : 39 — 40 mm.
Ifnb. The type is in my collection from the Kcbea IJange ; tlie insect flies
in March and April.
Diptheroides gen. nov.
Palpi nptnrned, first and second segments thickly clothed with longish hair, end
segment moderately long, slightly deflexed ; antennae very minntely ciliated, collar
erect, large ; thorax with large central metathoracic tnft ; abdomen with small
tnfts on proximal segments. Nenration : Primary with vein Obehuv the njiper angle
of cell, 7 anastomosing with lO to i'orm the arcole, which is most minnte, 8 given
oif from 9 near the apex, 9 given olT fmm 10 well beyond the areole. Secondary
normal.
Ty|ie : JUpthcroidcs henricki B-B.
35. Diptheroides kenricki spec. nov.
c?. Head pale greenish bine, jialpi cream-coloured, dark above, collar dark
grey at base with ochreons centre and green tip ; thorax blnish green, with a black
stripe below the collar on the patagia, which arc also tijjpod with black ; meta-
thoracic tuft with black stripe at base and almost bluish tip ; abdomen creamy
grey, with green dorsal spot next to thorax lollowed by two black spots and another
black spot on anal segment. Primary blnish green, with a black costal jiatch
at base, followed by a white dash, another larger black costal patch and another
short white dash ; subbasal blacdc line irregnlar and interrnpted, the median black
line rising in a large black costal patch forms a black spot at the end of the cell and
recedes beneath it, whence it is irrcgiihir and has a second Mack interrupted line
( 204 )
rather nearer the base below the cell ; the postraodiiil liiif fine, rising in a small
black cdstal spot, is irregular anil follows ronglily a similar course to tlie tni'(li;in
line ; the subtern)inal line, edged externally with white and rising in a black
triangular costal patch in whicli are two white points, is irregular with an indefinite
black dash to the terinen above vein •> and on vein 2. Between the costa and the
cell tlie gnmnd-colour is whitish ; there is a white spot on the interior of the
black spot at the end of the cell. Tcrnien black to vein 7. Fringes green,
intersected with black. .Secondary darkish grey, broadly yellowish cream-colour
along the inner margin and at the base.
?. Like the male, but the colour is a delicate lavender blue shaded slightlv
with greenish in the median and terminal areas, with all the black lines sliar[icT
in outline.
Expanse : cf 47 mm. : ? 40 — .M) mm.
//(id. The type is in my collection tVom Ihiunva, and was talcen in August ;
another specimen from Kkeikei was cajitured in .binuary or Felniiary : other
specimens are in the Tring Museum from the Aroa Uiver.
30. Diptlieroides serrata spec. nov.
c7. Frons ochreous, caput lavender-colour, collar ochreous at base, then broadly
ilark grey edge.l with greenish grey, with the extreme tip lavender-grey ; thorax
lavender-grey, jiatagiae tipped with ochreous grey, tuft tijiped in the centre with
reddish o<-hreous ; abdomen creamy ochreous, with a recldish proximal small dorsal
tuft followed by a darker one, central segments bandcil with black. Primary didl
grevish lavender-colour, with a basal costal black jiatch enclosing a small lavcndci'
sjiot ; subbasal line irregular, rising in a large black costal dash; aiitemedial,
medial, ]Histmedial and suhterminal Hues line, very frcipiently and strongly serrated,
the antemedial line being interrupted in the cell, the snbterminal rising in a large
black costal patch and being followed by a row of spear-point marks, the one
at the tornus being heavy, teimen finely black. In the cell is a small round
wliitish spot, cell closed by a thick black line which is edged internally by a double
whitish spot. The large black patch in which the snbterminal line rises e.xtends
obliquely to the termen, thus enclosing an oval ajiical lavender-grey ])atch. Fringe
lavender-grey intersected with black. Secondary dark grey, with base and inner
margin ]iale ochreous.
Expanse : 47 — 48 mm.
/f/i/j. The type is in my collection from Dinawa, where tlie insect (lies in
August ; other specimens are in the Tring Bluseum from the Aroa River.
Lasiosceles gen. nov.
Paljii upturned, second segment rea<'hing nearly to the vertex thickly scalecl, third
segnu'iit long and smooth; antennae minutely <;iliate ; jiroboscis fully develojied ;
thorax tufted, both jiro- and metathorax ; abdomen with long proximal tuft ; legs
spined, femora thickly clothed with long hair, tiliia.e clothed with long hair, with a
lai'ge expanding tuft on the tibial joints of both the mid- and hindlegs, whose tarsi
are also finely spined. Neuration : Primary, vein \i from the lower angle, 4 and •)
from (dose to the lower angle, 0 from immediately below the njiper angle, 7 from
off 8 at the end of the areole which anastomoses with 9, which is given oil l(i to
form the areole. Secondary, veins 3 and 4 from the lower angle, Ti from near the
( ^l>5 )
middle of tlie discocellnlars, 6 and 7 from the u|i|i('r angle. AViurjs: rosfa nearly
straight, depressed at the ajjex ; apex acute, teniieii very sliglitly exeavateil to
vein 5, prodiieed outwards at 4 to form a slight angle from whence it recedes ni]iiilly
to the tornus ; wing broad, secondaries comparatively small, terraen evenly rounded
to vein 2, where there is a slight angle. Tcrmen of both wings slightly crennlate.
Type : Lasiosceles pt-atti B-B.
:37. Lasiosceles pratti spec. nov.
6. Head and thorax red-brown lined witji white; the collar has two lines
right across, and is tipped with white ; the jiatagiae are edged with white, and have a
central white line; abdomen orange-yellow, proximal tuft dark red-brown. Primary
reddish browvi, with the space below the coll ])ale ochreons greyish ; all the veins
are white, very ])rorainently so beyond tlie cell, the three main veins being pale
ochreons greyish uj) to that point : the basal C(diular and costal area is tesselated
with white lines; below the cell tlu're is a short deep red dash edged with white,
followed by a curved white streak edged with deej) reddish ; the renifomi is edged
with white, its internal edge being a broudish sfraiglit white dash; above it is a
short costal streak ; jiostmedial line curved broadly white, snliterminal line straight
oblique, broadly white from the apex well beyond the tornus ; termen finely wiiite,
between this and the obli(jue stripe there is a dentate white line from vein 5 to the
tornus. Secondary jiale orange-brown, the base almost yellowish orange.
¥. Like the male, bnt the secondaries are darker.
Expanse : cj 44, ? 4(5 mm.
Hub. The type is in my collection. The species occurs at Ekeikei an<l tin-
Kebea Range in March and April, and in duly.
Reticulana gen. nov.
Palpi with second joint heavily scaled, long, reaching above the vertex, third
joint long, smooth; antennae minutely and finely ciliate in both sexes; all the
legs thickly scaled to the end of the tibiae ; head with short projecting tufts from
the base of the antennae ; collar with a high erect tuft ; abdomen tufted on the
proximal segment. Neuration : Primary with vein 2 from the basal side of the centre
of cell, 3 and 4 below the angle, o from the angle, 0 from below the upper angle,
7 from 8 at the end of the areole ; 8 and 9 stalked, given oif from 10 by a bar to
form the areole; 11 from basal side of the centre of cell. Secondary with 3 and 4
from the angle, 5 from just below the centre, C and 7 from the angle. Costa
straight, slightly dei)ressed at apex ; termen receding very gradually to the tornus.
Type : Rdicidana, costiliitca B.-B.
38. Reticulana costilinea spec. nov.
c??. Head blackish brown, with white tufts from the antennae; thorax
blackish brown, collar and patagiae edged broadly with whitish grey; mctathoracic
tuft small, edged with grey; abdomen dark greyisii, with jiroximal tuft on dorsum.
Primary dark grey, costa blackish with a series of short oblique whitish daslies,
a small snbbasal blackish spot ringed with whitish grey, a blackish very irregular
median oblique area edged and invaded by whitish grey lines, a posterior series of
blackish spots interrupted by the vein-space of vein 2, edged with whitish grey,
a snbterminal smaller series, and a complete terminal series of similar spots :
( 206 )
termen whitish crennlate. Secondary dark greyish lirown, with a short darkish
terminal dash above the tornns and a short white dash from the termen between
veins 1« and 2.
Expanse : 44 mm.
Ilah. Ekeikei, March; Babonni and Dinawa, September; Aroa River. Type
in my collection ; other specimens in the Tring Museum.
:^0. Perciana bistrigata sjiec. nov.
S. Head and thorax pale pinkish brown, patagiae dark purplish brown,
abdomen greyish with dark pnrplish dorsal tufts. Primary pale pinkish brown,
with antemedial and postmedial creamy lines, the former curved and very o])lique,
the latter erect, enclosing an area of dark purjilish brown below the cell ; the
postmedial is excurved and very tine above the lower angle of the cell, a dark
band of shading in the postmedial area. Secondary uniform brownish.
Expanse : 3U mm.
Hab. The type is in my collection from Dinawa, and was captured in September.
41 L Perciana meeki si)ec. nov.
i. Head and thorax dark ijnr]ilish lirown, abdomen greyish. Primary pnrplish
brown, with a nearly erect antemedial line and an incurved medial creamy line,
enclosing a dark area below the cell between them ; a trace of a postmedial creamy
line in the costal area, with a broad stripe of very dark jmrplish beyond it ; termen
irrorated with pale lilac. Secondary darkish brown.
Exj)anse : 38 mm.
Hub. The type is in my collection from Owgarra.
Vittappressa gen. nov.
Pali>i upturned, thickly scaled ; first segment with longish liairs, second
segment somewhat thickened at the end, third segment small, reaching beyond
the vertex of the head ; antennae of c? finely ciliated ; thorax with a short
metalhoracic tuft; abdomen tufted on second segment.
Nenration : with vein 0 from below the angle of the cell, 7 given off from 8,
which anastamoses with 9, which is given off from 10 to form the areole, beyond
which 1(1 is ajipressed to 9 to form a spurious areole beyond, whilst 11 is appressed
on to tlie areole j)roper and forms a spurious areole above the cell, lying touching
vein lo to the end of the false areole nearer the apex. Secondary with 3 and 4 on
a very short stalk, .') from just below the centre of the discocellnlara, G and 7 from
the angle of the cell.
Type: \'il/'/ppr/'.i.m riijij/liu/a. R-l?.
This genus is allied to J'rrc/<i»a AVlk.
41. Vittappressa rufiplaga spec. nov.
S. Frons pale grey ; vertex reddish, thorax pale grey with small tnft
reddish ; abdomen jiale grey with red tnft, and a reddish dorsal patch on
segments 5 to 7. Primary pale whitish grey, with two dark jwints on the costa
before the centre and beyond it, three fine costal dashes in front of the apex; a small
reddish dash closes the cell ; beyond the dark postmedial line from vein (i the
outer part of tlie wing is reddish brown, with a grey patch at the tornns and in
( 207 )
the middle of the termeu ; snbterminal pale line creniilate, somewhat indistinct.
iSecoudar}' pale whitish grey, with a red-brown irregular patch on the inner margin
extending ohliquely upwards with an ochreons short stripe near its inner margin ; a
trace of a white medial crenulate line, with slight darker shading beyond it, margin
dotted with red. Termen and fringes of both wings crenulate.
9 . Similar to the male in all i)articulars.
E.xpanse : <? 32, ? 34 mm.
Hab. The type is in my collection from Dinawa, where the insect flies in
September.
42. Borbotana kebeae spec. nov.
c?. Head, antennae and thorax reddish nmber-brown, abdomen pale ochreons
grey, dorsal tufts tipped with reddish brown. Primaries dark somewhat lustrous
brown, with a broadish creamy white basal stripe, with a prominent projection on
the outer margin below the cell and a very slight one above the cell, a postmedial
oblique rather irregular narrow whitish stripe very finely intersected in the centre ;
bej'ond this the ground-colour is jialer with a very irregular outer edge, bej'ond
which it is quite dark to the termen, which is spotted with blackish between the
veins ; between the white postmedial stripe and the apex are three fine white points
on the costa followed by a short white very fine W-shaped mark, an obscure dark
dot at the end of the cell. Secondaries whitish with brownish costa and termen,
which latter tapers rapidly to the tornns. Undersurface : Primaries somewhat
lustrous brown with a whitish dash at the upper angle of the cell but beyond it ;
secondaries whitish, densely irrorated with reddish brown along the costa and
the termen t'aj)ering rapidly to the tornns ; in front of the apex there is a distinct
dentate short line on the costa with a trace of a continuation towards the inner
margin.
? . Just like the male, only more mottled in the brown colour of the
primaries.
Expanse : c? 31 mm., ? 33 mm.
Hab. The type is in my collection from the Kebea Range ; it flics in March
and April ; other specimens from Ekeikei.
This is no doubt a local race of B. nicifascia Walker, but all the New
Guinea specimens are uniform in their narrow, almost even, postmedial line.
43. Borbotana ekeikei spec. nov.
? . Head, thorax, antennae and palpi reddish brown, abdomen pale brownish
grey. Primary very dark brown for the basal half, with a very broad whitish
antemedial band slightly curved on its inner margin, but with two projected blnnt
dentate marks on its outer margin, the one at the lower margin of the cell being
much the larger ; the dark area is n:argined by a fine white line, with a sharp
toothlikc projection at the end of the cell inclosing a small whitish spot ; posterior
half of the wing very pale pinkish brown, darker towards the termen, with an
oblique short reddish brown dash from the apex. Secondary pale reddish brown,
becoming whitish towards the base. Undersurface : Primary uniform lustrous
pinkish brown ; secondary as above, but with a postmedial dark line. .
Expanse : 28 mm.
Ilab. The type is in my collection from Ekeikei. This insect flies in March
and April,
14
( 208 )
44. Borbotana dinawa spec. nov.
(?. Head, thorax, antennae and palpi blackish brown, abdomen pale lirowu
with dark dorsal tufts. Primary very dark blackish brown, with a paler dash below
the costa at the end of the cell, and the terminal area paler, termeu narrowly very
dark ; from the base of the costa a white-toothed very oblique dash extends to
the lower margin of the cell about halfway along, and is slightly toothed at the
costa, on the outside more strongly before its centre and yet more strongly at
its lower extremity where it is broadest ; at the ujiper end of the cell is another
dash almost hook-shaped at its inner end ; secondary whitish, dark towards the
termen.
Expanse : 27—28 mm.
Hab. The type is in my collci'tion from Dinawa, and the species tlies in
September.
45. Tarache kebeae spec. nov.
cj. Head white, collar white liuely divided in the centre with black and with
a broad central black tip, thorax white, patagiae with a black lateral dash, abdomen
grey, almost black l)elow with a white abdominal patch ; legs blackish, a white
spot at the femoral and tibial joints. Primary white with a short basal black
dash in the cell, a subbasal black point on the costa followed by a black wedge-
shaped spot beyond which is a large irregular wedge-shaped black patch, with
a small s(inarish spot beyond it, followed by a black point, an apical black patch,
touching a black spot on the termen, a black ])atch at the tornns, and a black
wedge-shaped patch nearer the base with a small black spot between the two.
Secondary blackish grey, whitish on the inner margin and at the Itase, fringe
whitish except in the apical area, where it is blackish. Undersurface, both wings
sooty black. Primary with a white costal spot before the apex, and on the termen
below the apex and above the tornus ; secondary, innermarginal and basal areas
whitish.
Expanse : 32 mm.
Ilab. The type is in my collection from the Kebea Range, where the insect
flies in March and April.
4(i. Bagada dinawa spec. nov.
S. Head and thm-ax pale ochreous grey darkly irrorated, abdomen lirownish.
Primary pale ochreous grey, irrorated in parts with darker pinkish grey ; the
autemedial, medial and postmedial lines are present in the costal area, but are not
continued below the cell ; a large dark pinkish grey patch occupies the end of the
cell and beyond the cell, in which the reuiform stigma shows as an orange spot,
and the orbicular stigma precedes it and is i)alely encircled, a darkish dash from
the apex to the dark patch. A subterminal serrated grey line is obscurely
discernible at the end of the patch previously mentioned, three dark dots above
the tornus. Secondary pinkish grey, with a darker termen and a pale costa.
Undersurface : Primary pinkish, with a dark subcostal stripe and a darkly
spotted postmedial line, terminal area grey. Secondary greyish, darkly irrorated
in parts with an irregular dark postmedial line.
Expanse : 28 mm.
Ilab. The type is in my collection from Dinawa, where the insect flies in
August. The species will come next to B. spicea Guen,
( 209 )
47. Bagada fuscostrigata spec. nnv.
(?. Head and pal{)i reddish lirown, collar grey, patagiae reddish, abdomen
brownish grey. Primary reddish with the inner margin slate grey, an obli(jue
dark grey stripe from just below the apex to the middle of the inner margin,
outside which the ground colour is greyish, the termen between veins 3 and 6
being paler ; the red area has a mottled appearance, the orbicular and reniform
stigmata being pale with a paler spot below the latter, whilst above them on the
costa is an indefinite patch of grey, and a pale spot at the apex. Secondary uniform
dark grey.
Expanse : 42 mm.
Hah. The t3-pe is in my collection from Ekeikei, where the insect flies in March
and April. The species will come next to B. lignigera Walker.
48. Bagada incertissima spec. uov.
$. Frons pale ochreous mixed with pink, head reddish mixed with pale
greenish scales, collar reddish with two grey lines across, thorax reddish ochreous
with greenish patagiae, abdomen pinkish brown with an ochreous extremity.
Primary chestnut brown with a pale snbbasal patch below the cell, below which
is a dark dash followed by a pale dash on the inner margin ; near the centre of the
costa are three short dark broad lines to the cell ; posterior half of wing darkly
blotched, with the orbicular and reniform stigmata more or less distinct, with a
creamy white narrow quadrate spot emitted just beyond the latter ; posterior line
dark and dentate, followed by a broader pinkish line ; tornns to middle of inner
margin and termen green ; termen above the centre paler, tinged with green ; on
the costa above the whitish quadrate mark is a pale patch. Secondary uniform
brown with a slight lustre.
Undersurface : Primary brown, costa paler, inner margin quite pale ; a
trace of a postmedial dark line, followed by a paler one ; a distinct creamy
subterminal dentate line. Secondary pale, with an interrnpted dark postmedial
line, beyond which the wing is rather darker, with a pale termen darkly spotted ;
a dark spot at the end of the cell.
? . Like the male, but darker, with the exception that in the primaries the
pale ochreous parts in the male are green in the female, and the green parts of
the male are ochreous in the female.
Expanse : S and ? 38—40 mm.
Hah. The type is in my collection from Ekeikei. Its general time of flight is
in March and April ; but I have specimens taken in January and also in Jnly.
Other specimens are in the Tring Museum from the Aroa River.
This species is most variable, scarcely two specimens being alike. I have
specimens quite dark all over, with dark green patches as described and a large pale
apical patch. Again, others are strongly suffused with green, and have an almost
black apical patch. I have a male which is pale ochreous with no green at all, the
only constant mark apjiarently being the small whitish ijuadrate spot just beyond
the reniform stigma ; this appears in all the specimens that we have received.
49. Bagada nigrostrigata spec. nov.
$. Head and thorax ochreous, patagiae tinged with pink, collar broadly
edged with dark brown, abdomen ochreous grey. Primary pale ochreous, tinged
( 210 )
with jiinkish in jiarts ; basal lino very obscure, almost obsolescent, except across
cell ; a double antemedial interrajited line, curved and waved, the outer line darker
than the inner ; reniforra stigma jmle greenish, above it a dark costal jiatch with
a blackish s])ot in front of the upper part of the reniform ; jiostmedial line iuter-
rn]ited, serrate ; termen slightly fuscous to this line ; snbterminal line distinct,
waved and irregular with a pale exterior edging ; a blackish dash between
veins 3 and 4 from the reniforra stigma to the termen ; fringes darkly dotted at
their extremities. Secondary pale brownish.
Expanse : 38 mm.
Ilab. The type is in my collection from Dinawa.
5ii. Xanthoptera dinawa spec. nov.
(?. Head and thorax ochreous grey, abdomen darker grey. Priniary ochreons
grey, finely irrorated with blackish scales; basal, antemedial and medial lines suflFused,
interrupted and angled about the cell ; postmedial suffused for the costal ]iart and
angled on vein 7, from which it recedes to the inner margin in a double line
and is crenulate ; a dark grey subapical patch ; a dark grey snbterminal spot below
vein 3 ; termen darkly dotted. Secondary pink, with a dark spot closing the cell ;
a dark postmedial line ; termen broadly dark grey ; fringes to both wings ochreous.
?. Similar to the male, but paler.
Expanse : S 29—32 mm.; ? 36—37 mm.
Hab. The type is in my collection from Dinawa, where the species fly in
August and September.
.51. Cosmia inconspicua spec. nov.
Head and prothorax greyish white tinged with pink, patagiae ochreous grey,
abdomen pale brownish ochreons, legs grey, tarsi ringed with white. Primary
ochreous pale grey, sparingly irrorated with dark grey, with a dark medial and
postmedial line projected outwards in the cellular area, the interspace being darkly
filled in ; an obscure grey serrated snbterminal line ; the terminal area brownish
ochreons, with a broad dash of the same colour between veins 5 and 7 into the
postmedial line ; termen finely darkly dotted ; fringes tipped with grey ; an obscure
grey spot on the costa before the apex. Secondary pale subhyaliuc greyish,
ochreons near the termen.
Expanse : 32 mm.
Ilab. The type is in my collection from Ekeikei, where the insect flies m
March and April.
52. Acontia aroa sj)ec. nov.
i. Head and thorax very pale canary-yellow, the latter with a red fawn-rolour
central stripe ; abdomen yellowish grey. Primary pale canary-yellow, with a
reddish fawn-colour central stripe expanding gradually from the base into the apex,
and nearly into the tornus at the termen ; a slight shading of the same colour along
the inner margin. Secondary yellowish cream-colour.
Expanse : 46 mm.
Ilab. The type is in my collection from the Aroa Kiver. It will come nest to
A. emboloscia, but it entirely lacks all transverse lines.
( 211 )
53. Zalissa ekeikei spec. nov.
(J. Head pinkish buff, cullar purplish red-brown with a central triauguhxr
jiatch of pale cinnamon-brown, thorax pale cinnamon-brown with pale whitish
grey patagiae mottled with chocolate-brown, abdomen yellowish. Primary warm
ciunamon-brown with a fine pale costa, inner margin broadly purplish brown,
irrorated with whitish, with a pnre white internal line extending round the tornus
in a small angle, termea broadly purplish, confluent with the inner margin, the
terminal area being pale, irrorated with whitish (but to a less extent than the inner
margin), and margined internally by a curved dark purplish line, which is again
edged internally by a broader and darker stripe of the same colour, having an
almost straight internal border ; termeu finely dark. Secondary yellow with a
dark brown termen very broad at the apex, tapering narrowly to the toruus,
where are two short waved pale lines. Fringes white for the apical half, dark
below vein 3.
Expanse : 44 mm.
Ilab. Ekeikei, where the species flies in March and April ; we have it also
from other localities. ,
The type is in my collection.
54. Zalissa kebeensis spec. nov.
S . Differs from B. ekeikei in that the primary is pink above the cell, gradually
toning down into a pale yellowish olive ; the borders are similar to the previous
species, but darker. The secondary is yellow with the termen broadly brown from
vein (5, increasing in width slightly at the tornus. The underside is uniformly pale
yellow in both wings with only a narrow darkish termen, but having none of the
dark pattern of ekeikei.
Expanse : 45 mm.
JIab. Kebea Range, Aroa River, February and March.
Type in my collection ; other specimens in the Triug Museum.
55. Rivula aroa spec. nov.
<?. Head and collar whitish, thorax tinged with grey, abdomen ochreous grey.
Primary ochreons grey, with antemedial and postmedial pale lines excurved at the
cell, reniform dark spotted with black, subterminal line dotted, terminal area
darkish, termen preceded by a fine whitish line, Costa finely dark, with three
whitish points before the apex. Secondary greyish, darker near the termen.
Expanse : 20 mm.
Hab. The type is in the Tring Museum from the Aroa River, where it was
captured in March.
The species comes near li. ommatopis Hpsn.
56. Rivula meeki spec. nov.
(?. Head and collar sepia brown, thorax lavender grey, abdomen grey. Primary,
base very restricted greyish, a large median sepia-brown patch with a lavender
centre ; this patch does not extend over the cell ; a small sepia-brown costal patch
with lavender-grey edging precedes the large patch ; postmedial and subterminal
whitish irregular linos enclosing a lavender-grev band ; subapical area with dark
dashes ; termen with dark dashes edged internally with white. Secondary greyish.
(212 )
Expanse : 20 mm.
Ilah. The type is in my collection from the Aroa River ; other specimens are
in the Tring Mnsenm from the same locality in March.
57. Tathodelta aroensis spec. uov.
$. Head, thorax and abdomen pale lilac brown. Primary pale lilac brown,
with olive-brown bands and lines ; an irregular autemedian and median Hue, the
latter broadly edged externally with a band of slightly paler olive brown. Position
of reniform marked by two small blackish spots ; a darkish irregnlar snbterrainal
Daud of shading ; termen broadly olive brown. Secondary pale greyish.
Expanse : 26 mm.
Hab. Aroa River, March.
Type in the Tring Museum.
Eublemmoides gen. nov.
Palpi upturned, thickly clothed with long hair ; third segment minute, reaching
above vertex of head ; antennae minutely ciliate. Legs short, stout, with tufts of
long hair on tibiae, and to a less extent on tarsi of the hind pair ; both mid- and
hiudtibiae with two pair of spurs. Wings : costa of primary depressed before apex,
which is strongly excised below, making it almost falcate, produced into a point
between veins 3 and 4, and again excised slightly to the tornns. Secondary
angular, with costa nearly straight, termen rapidly receding to tornus. Neuration :
Primary, veins 6 and 7 from the angle ; 8, 9 and 10 stalked from in front of the
angle. Secondary with cell very short, not more than a (juarter of the wing; veins
3 and 4 on a short stalk, 5 from below the centre, 0 and 7 from the augle.
Type : Eublemmoides dinawa B-B.
58. Eublemmoides dinawa spec. nov.
S. Head and collar dark red-brown ; thorax and alidomeu whitish grey.
Primary ochreons grey for the basal half of the wing margined by the medial line,
which is darkly dotted, obliquely produced externally to beyond the cell, whence it
recedes in an angle to the inner margin ; a trace of a snbbasal line below the costa ;
outer half of wing darker grey, with a brown fine postmedial line followed by a row
of whitish i>oints, and a band of yellowish brown shading. Secondary whitish grey,
slightly darker towards the termen, with termen rather broadly yellowish brown.
Both wings are finely but sparsely irrorated with dark scales.
Expanse : 24 mm.
Hab. The type is in my collection from Dinawa, where it was taken in the
mouth of August.
This genus appears to be most nearly allied to Hampsou's Section 111. of
Eublemma Hb.
50. Magulaba nigra si)ec. nov.
<J. Head, thorax, and both wings dark slaty grey, with an interruiitod dark
subbasal line ; a blackish jiostmedial line excurvcd beyond the cell, internipU'd by
the paler veins ; reniform darkly outlined, a subterminid irregular line of darkish
sliadiug ; termen with fine dark points. Secondary witli a dark jwstujedial line,
( ai3 )
with a paler external edging, beyond wliicli is a short similar line in the tornal area
only.
Expanse : 'M mm.
Ilab. Aroa River, March.
Type in ray collection ; other specimens are iu the Tring Museum.
60. Zagira bicolora spec. nov.
c?. Head and collar dark grey, thorax creamy white, abdomen grey. Primary
with a broad creamy white costal stripe ; the rest of the wing dark grey, in which
is a postmedial waved whitish line, edged internally by a dark line ; a trace of a dark
snbterminal line ; both tliese lines are carried through the secondaries. Secondary
with very restricted whitish base, the rest of the wing being dark grey.
Expanse : 21 mm.
Hab. The type is in my collection from the Aroa River, where it occurred in
March ; other specimens are in the Tring Museum from the same localit)'. Two of
the Tring specimens differ somewhat from my type ; in one the costal stripe is
snow-white, and in the other almost rusty white.
*>1. Zagira pratti spec. nov.
c?. Differs from bicolora in that the postmedial line is broadly white, with a
fine dark central line in it ; the subterminal line is pale, projected forwards on
vein 4, and slightly curved inwardly above and below it ; the shape of the secondaries
is much more angular.
Expanse : 21 mm.
Hab. The type is in my collection from Dinawa, where it occurred in August.
G2. Corgatha bipunctata spec. nov.
¥ . Head dark grey with a few pink scales intermixed, thorax jjinkish
ochreous pale brown, abdomen pinkish ochreous pale brown with the anal half
blackish on the dorsum. Primary pinkish ochreons pale brown, with the costa
blackish for the basal half and two white costal spots near the centre ; three black
points from the first white costal spot marking the course of the absent antemedial
line ; a trace of the medial line from the second white spot ; a trace of the
postmedial line in a'series of black points with white externals, the two points near
the inner margin being larger and suffused ; three black terminal dots below the
apex with a blackish grey internal suffusion ; below the apical dots are four black
terminal points on the veins. Secondary similar in colonr to the primary, with au
antemedial dotted line and a jwstmedial dotted line, as also a dotted termen as iu
the primary ; a strong blackish snflfusion in the tornal area, developing mainly
in two serrated short innermarginal lines.
Expanse : 29 mm.
Hab. The type is in my collection from Dinawa, where it was taken in
September.
63. Corgatha pulchra spec. nov.
cj. Head and thorax lavender grey, abdomen brownish. Primary with base
pale lavender grey, with a double dark antemedial line filled in with lavender grey;
the outer line is waved below the cell ; median area bright orange-red, edged
externally by au irregular double postmedial line, the inner line of which is blackish
(214)
aud tbe outer serrate and dark ; this is followed by an irregular indefinite area of
bright orange-red ; subtermiual line irregular, with a large apical lavender-grey
patch edged darkly internally ; below this the tornus is duller reddish. Secondary
orange-brown with double medial and iiostuiodial lines, which are irregularly
waved ; termen crennlate with fine dark points.
Expanse : 22 mm.
Ilab. Aroa River, March.
Type in my collection. Other specimens in Triug Museum.
The Tring Museum has one specimen which is quite dull purplish in tone of
colour.
04. Corgatha albolineata spec. nov.
?. Head, thorax, and abdomen chocolate-brown, the former irrorated with
grey ; both wings chocolate-brown. Primary, base with a short white dash ;
antemedial dark line with pale grey indefinite margins ; postmedial pale line waved,
irregular ; an oblique white line from the inner margin near base to upper margin of
cell and continued along vein 7 ; veins 8 and 9 outlined with whitish grey, a white
spot in the cell crossing the white line, a white curved line from the costa enclosing
a mottled apical patch, at the top and bottom of which is a white spot ; from the
lower edge of the white curved line the subterminal pale serrate line descends to
near the tornus ; termen dotted with black, costa dotted with white, cilia tesselated
with cream below the apex. Secondary with restricted white base ; a black
dot in the cell ; medial and subterminal pale lines waved and irregular ; termen
with a row of fine black points edged internally with fine white points just in front
of it ; inner margin creamy white.
Expanse : 20 mm.
Hab. Aroa River, Januar}'.
Type in the Tring Museum.
This species will come next to C. pltujiostola Hpsn.
0.5. Corgatha mediopallens spec. nov.
c?. Head, thorax, and abdomen pale pinkish grey with a central ])rothoracic spot
of rusty reddish ; abdomen rusty reddish on tlie proximal segment. Both wings
pale slaty-grey. Primary mottled with rusty reddish, edged by a dark line ;
reniform dark with a dark angled dash to the costa ; postmedial line indi.stinct,
incurved on the fold ; a rusty-reddish patch at this jioint and a large rustv-reddish
patch iu front of the apex, with an irregular white subapical curved line ; termen
darkly dotted. Secondary with an obscure trace of a medial and jrastmedial line ;
termen with fine dark points.
Expanse : 24 mm.
Hab. Aroa River, March.
Type in the Triug Museum.
0(1. Corgatha nigropunctata spec. nov.
cj. Head and thorax pale umber-brown. Hoth wings ])ale und)er-brown ; a
trace of an antemedial, medial, and jiostmedial line, the two latter excurved over
the cell ; one or two blackish basal points ; a black spot in the cell ; subterminal
indistinct line, rising from a white waved short costal stri[)e ; teruien with line
(215)
black jmints. Secondary with the Hues as in primary, carried tlirongh, and the
termen with fine black points.
Expanse : 24 mm.
Hab. Area River, February.
Type in the Tring Mnseiim.
67. Oruza dinawa spec. nov.
S- Head, thorax, abdomen, and both wings pale reddish brown. Primary
with a broad inwardly obliqne dark purplish brown stripe, beyond which is the
irregular fine snbterminal darkish line ; termen angled between veins 3 and 4.
Secondary like the primary, but th(; bniwu broad line is slightly outwardly
oblique.
Expanse : 31 mm.
Hab. Dinawa, August.
Type in my collection.
68. Oruza aroa spec. nov.
i. Head, .thorax, and abdomen dull reddish brown; collar very pale, pinkish
ochreous grey. Both wings dull reddish brown, with an obliipie dark strijie carried
through both wings. Primary with a trace of a waved irregular fine dark line,
which is scarcely perceptible in the secondary.
Ex])anse : 26 mm.
Hub. Aroa River, March.
Type in my collection ; also specimens iu the Tring Museum.
69. Oruza variegata spec. nov.
c?. Head, thorax, and abdomen ochreous, densely covered with purjjlish pink.
Both wings yellow ochre, densely irrorated all over with purplish red, except in
the primary ; a subtriangular costal patch over the cell and a large apical patch ;
an antemedial fine line, a medial oblique broad darker stripe, a postmedial fine
serrate line. Secondary with basal ochreous, a broad medial darker stripe followed
by an inuermarginal ochreous patch, a reddish irregular postmedial line edged
externally with ochreous ; an ochreous tornal area.
Expanse : 24 mm.
Hab. Dinawa, August ; Aroa River, January.
Type iu my collection ; also specimens iu the Tring Museum.
TO. Oruza unipunctata spec. nov.
cJ. Head, thorax, abdomen, and both wiugs pale pinkish. Primary with
slightly waved antemedial and postmedial oclireous lines; reniform rusty reddish
brown. Secondary with a waved postmedial ochreous line ; a dark spot in the cell.
Expanse : 22 mm.
Hab. Aroa River, February.
Type iu the Tring Museum.
Tl. Phanaspa ochracea spec. uov.
?. Head, thorax, and abdomen ochreous grey. Both wings ochraceous.
Piimary witli a fine dark antemedial line, a median dark obscure cloudy band,
a tine crenulate dark postmedial line ; subtermiual line pale irregular dentate.
(216 )
indefinitel.v edged laterally uu each side with dark grey, with a small ajjical dark
patch ; termeu with black dot. Secondary with greyish base, a dark dot iu the
cell, a crenulatc waved dark postmedial line ; termen very broadly darkish grey,
with a pale creniilate line in it ; termeu with black dots.
Expanse : ',W mm.
J/ab. Dinawa, September ; Aroa l?iver, Febrnary.
Type in my collection ; also specimens in the Tring Mnseum.
72. Phanaspa rubra spec. uov.
cJ. Head, thorax, and abdomen pinkish ochreous. Primary pinkish ochreons,
with a trace of a basal dark line ; an irregular fine dark antemedial and medial
line, an obscure postmedial line terminating iu a blackish spot above the tornns, an
obscure subterminal line ; termen with fine black points ; a black dot in the cell.
Secondary pinkish, with obscure medial, postmedial, and subterminal lines ; termen
crenulate, with fine black points.
Expanse : 22 mm.
Had. Aroa River, February.
Type in the Tring Museum.
This species will come next to P. usttila Hpsn.
73. Phanaspa punctata spec. nov.
c?. Head, thorax, and abdomen greyish. Both wings i)ale ochreous greyish,
with the lines marked out by blackish dots ; two submediau blackish dots iu the
cell, two blackish dots at the end of the cell ; a curved line of postmedial dots
incurved on the fold ; an interrupted row of subterminal dots ; termen dotted.
Secondary with median, postmedian, and terminal row of dots.
Expanse : 31 mm.
Hab. Aroa River, January.
Type iu the Tring Museum.
74. Phanaspa dinawa spec. nov.
(J. Head, thorax, abdomen, and both wings Very pale cinnamon-brown.
Primary with a trace of a jiale subbasal line ; postmedial line obscure, pale, crenulate,
excurved at the cell ; between it and the very irregular subterminal line is another
indefinite i)ale serrate line ; subterminal line darkly edged, and with a dark patch
at the inner margin ; one or two dark jioiuts in the cell; termen with blackish dots.
Secondary with two dark points across the cell ; a pale medial dentate line, followed
by an indefinite similar line ; postmedial pale line crenulate ; termen with dark
points ; on the inner margin above the tornus are three dark chocolate sjiots.
Expanse : 28 mm.
IJab. Dinawa, August ; Aroa River, January.
The type is in my collection ; other specimens in the Tring Museum.
7.5. Pseudacidalia unilineata s]iec. nov.
S. Thorax, abdomen, and Ijoth wings pale dull cinnamon-brown. Primary
with a tine outwardly obliipie antemedial dark obliiine line, a broad inwardly
oblique dark postmedial line jialely edged externally ; termeu with fine black
points ; cell with a dark point. Secondary exactly like the jirimary, but witliout
( 217 -.)
the antemedial Hue and with a small oeliraceous patch from the ohlicine line
by the cell.
Expanse : 25 ram.
Hab. Aroa River, January.
Type in the Tring Museum.
The species will come next P. albicosfa Moore.
76. Microeschus parva spec. nov.
(?. Both wings creamy whitish. Primary with traces of antemedial and
postmedial lines, the latter being fairly distinct ; reniform darkly edged internally
with a dark costal dash over it ; termen dark. Secondary with traces of antemedial
and postmedial lines ; termen dark.
Expanse : 12 mm.
Hab. Aroa River, January.
Type in the Tring Museum.
PALINDIINAE.
77. Dinumma hades spec. nov.
S ■ Head, jialpi, and thorax dark brown ; abdomen greyish, with dark brown
tufts on the dorsum. Primary dark brown, with a slightly paler antemedial,
strongly dentate line having a fine dark linear outer edging ; postmedial line
excurved slightly crenulate ; a trace of a hue pale grey subterminal line; a fine
terminal series of blackish short Inuular linos in the vein interspaces. Secoudary
uniform greyish brown, with a pale bmad band across the medial part of the cell
which is interrupted by the veins.
Expanse : 31 mm.
Hah. The type is in my collection from Dinawa, where the species flies from
May to July ; other specimens in the Tring Museum.
78. Dinumma mediobrunnea sp. nov.
?. Head and thorax chocolate-brown finely irrorated with greyish, abdomen
greyish with choci date-brown dorsal tufts. Primary with basal area pale greyish,
a very small purjilish brown basal costal patch, antemedial line waved irregularly,
postmedial line pale tortuous, the area between these lines uniformly dark purplish
brown ; following the jiostmedial line is a band of pale reddish chestnut, beyond
which the area is paler lilac brown with a dash of reddish above the two deep
black spots below the apex and above the tornns near the termen ; a pale spot
edges the former spot below ; subterminal line pale and crenulate; in front of the
termen is a row of silvery white spots, termen finely pale. Secondary greyish
brown, slightly paler basewards.
Expanse : 32 mm.
Hab. The type is in my collection from Ekeikei, where the insect flies in July.
70. Callyna biplagiata spec. nov.
c?. Head and collar deep chocolate-brown, the latter finely tipped with lilac-
brown, thorax lilac-brown, abdomen brown. Primary pale lilac-brown with an
irregular subbasal costal patch, extending just below the cell, and a large costal
( -^18 )
patch from the end of the cell nearly to the apex of deep chocolate-brown ; the
latter has an evenly carved interior margin, and both are palely edged ; the ante-
medial line forms the onter edge of the dark subbasal j)atch, and is projected forwards
to the inner margin; tlie medial line consists of au indefinite band, somewhat cnrved,
of darker brown ; the ])OStmedial line consists of the margin to the large costal
jiatch nearly to its middle, when it descends perpendicularly to the inner margin ;
three dark spots edged externally with whitish are below the outer edge of the
large patch ; the orbicnlar and reniform stigmata are edged with whitish ; there is
a small white ajiical patch ; fringes white i)y this patch, brown below it.
Secondary uniform dark brown, slightly paler from the end of the cell up to
the costa.
Expanse : 43 mm.
Hah. The type is in my collection from Ekeikei, where the species flies in
January and February.
80. Callyna fuscantaria spec. nov.
(J. Head and thorax reddish brown sparingly irrorated with pale grey, abdomen
greyish brown, legs brownish palely ringed. Primary reddish brown, with the
antemedial line waved and outwardly oblique ; the postmedial line is excurved and
crenulate ; both of these lines rise in a pale costal dot ; between the two, crossing
the end of the cell, is an angled band of dark brown somewhat indefinite as to its
margins ; a large dark reddish brown patch exte ds from the apex along the termeu
almost to the tornus ; immediately in front of the apex is a trace of a round s|)ot
obscurely outlineil with ])ale grey. Secondary uniform dark brown, slightly
lustrous.
Expanse : 30 — 38 mm.
Hub. The type is in my collection from Ekeikei. The species flies in March
and April.
81. Westermannia longiplaga spec. nov.
c?. Head pale grey with a pair of dark grey patches, collar j)ale lavender-grey,
darker at the tip, thorax dark smoky grey with whitish grey patagiae and meta-
thoracic tuft, abdomen grey. Primary very pale lavender-grey with a trace of a
medial and postmedial lines which are slightly oblique ; the whole of the cell ami
expanding somewhat to the termen is dark greyish bronzy brown, with an interior
blackish grey stripe to well beyond the cell, whence it ascends obliquely to the
apex ; below the angle of this stripe is a whitish dotted line to tlie inner margin,
with which a snbterminal row of similar spots is roughly parallel. Secondary
whitish grey, darker at the termen.
Expanse : 50 mm.
ILib. The type is in my collection from Dinawa, where the species flies
in August.
The species is nearest ir. argentea Hamps.
82. Homodes punctistriga spec. nov.
S . Head, thorax, and abdomen dull lirick-red. Primary dull brick-red, with
antemedial, medial, postmedial, and snbterminal spotted lines of silvery lead colour ;
between the two latter are two lines of darker red ; all the lines are somewhat
( 219 )
waved ; termen finely black with precediug fine black dashes. Secondary exactly
like the primary without the postmedial line.
Expanse : 30 mm.
Hah. The type is in my collection from Ekeikei, where the insect flies in
March and April.
SARROTHRIPINAE.
S3. Earias uniplaga spec. nov.
(J. Head, thorax, and primaries j)alest sulphur yellow, abdomen whitish.
Secondary white, somewhat hyaline. Primary with the costa at the base deep
reddish brown ; an orange spot at the end of the cell, somewiiat of the shape of a
figure 8, margined with deep rusty red.
? . Like the male, but without the spot at the end of the cell.
Ex]ianse : S and ? 32 mm.
Hub. Dinawa, the Kebea Range, July and August; Aroa River, February
and March.
Tyjie in my collection, and in the Tring Mnsenm are other specimens.
84. Earias rufopunctata sjiec. nov.
c? and ? . Similar to E. loioplaga, but somewhat paler and with no dark costa
at the base of the primary, whilst instead of the peculiar spot at the end of the
cell the primary has a very pale round slightly rusty spot.
Expanse : 30 mm.
Hab. Dinawa and Aroa River, July.
Type in my collection, and other specimens in the Tring Museum.
85. Earias novoguineana spec. nov.
(J and ? . Head, thorax and primary chrome yellow ; abdomen whitish tinged
with sulphur. Secondary whitish tinged with sulphur-colour. Primary with a
dark rust)'-red round spot at the end of the cell, preceded by two obliquely placed
dots, a trace of an obtusely angled posterior line having a dark point below the
costa, another at the apex of the angle, and two more near the inner margin.
Expanse : S and ?, 38 mm.
Hab. Dinawa and Kebea Range, July and August.
Type in my collection.
86. Hylophila rubromarginata sjiec. nov.
?. Head and collar slightly tinged with green; thorax greenish white,
abdomen yellowish white. Primary white, with the least greenish tinge at the
base and in the postmedial area, with oblique yellowish antemedial and postmedial
lines, the latter being very oblique and straight ; this last point is constant in all
our specimens (a good series). In H. orientalis, its nearest ally, this line is curved.
Costa and inner margin narrowly pink ; termen deep pink and somewhat wider.
Secondary pure white.
Expanse : 30 mm.
Hab. The type is in my collection from the Kebea Range, where it occurred
in March and April. Other specimens are in the Triug Museum from the Aroa
River,
( 220 )
87. Gadirtha costipallens spec. nov.
? . Head ochreous grey, collar greeuish ; thorax pale grev witL greenish
patagiae ; abdomen pale ochreous gre3^ Primary grey, with a slight suffnsion of
greenish ; a blackish snbbasal ])atch, within wliich is a small ochreous short dash ;
apical two-thirds of costa broadly whitish grey, median dark line fine, very obscure,
with a dark dot in the cell ; reniform pale brown, darkly pnpilled and finely
encircled ; jiostmedial line linear black, waved, produced rapidly outwards to vein 6,
receding rapidly to inner margin from vein 2, beyond which the ground-colour
is pale reddish, a dark dash below the ai)ex, a row of snbterminal obscure short
white dashes. Secondary dark grey with paler inner margin.
Expanse : 42 mm.
Uah. The type is in my collection from Ekeikei, where the species occurs
in January and February.
88. Gadirtha inconspicua spec. nov.
? . Collar pale chocolate-brown finely irrorated with dark grey ; thorax and
abdomen pale grey irrorated with dark grey. Primary p;ile olive-grey for the basal
half, with the costa pale chocolate-brown, terminal half pale chocolate-brownish,
the inner margin being strongly oblique, a trace of a snbbasal line, snbterminal line
dark, very irregular, angled, rising in a whitish grey costal patch ; apex with an
oval whitish-grey patch from which descends the serrate snbterminal interruiited
line. Secondary dark grey, pale in the basal area.
Expanse : 40 mm.
llab. The type is in my collection from the Kebea Range, where the insect
flies in May and June.
89. Ochthophora turneri spec. nov.
?. Collar grey, very broadly banded for its upjier edge with dark purplish
grey ; patagia ochreous grey with the metathoracic tnft darker. Primary lilac grey
with the base ochreous grey, palest on the costa ; subViasal area dark grey, margined
by the irregular serrated fine autemedial line ; a small raised dull chestnut spot
in the cell followed by a larger similar one at the end of the cell ; postmedial line
broadly serrate, very irregular, receding inwards and encircling irregularly the
larger raised spot, below which it recedes to the inner margin; a snbterminal row
of fine dark spear-shaped marks palely infilled, interrupted in the nerve spaces,
followed by a row of fine dark dashes. Termen darkly dotted. Secondary pale
grey, paler towards the base and inner margin.
Expanse : 40 mm.
Ilab. The tyi)e is in my collection from Bubooiii, where the species flies in
September.
This species is near 0. sericina Turner, but differs in the pale base and dark
subbasal area, and the snbterminal line is quite diti'erent.
90. Labanda viridescens spec. nov.
?. Head brown, collar dark brown with frontal of green scales ; thorax greyish
brown ; abdomen ochreous. Primary greyish brown, with a suifusion of green
scales all over the wing with the exception of the median area, where it is scanty ;
this suffusion is most prominent in the basal and postmedial areas, also over the
( 221 )
cell; the subbasal liuo is wliitish in the costal area and greenish below ; the dark
postmedial line is palely edged, or greenish in parts, verj- irregnlar and strongly
serrate ; in the postmedial area there is a dark band of the ground-colour and
a smallish dark apical patch. Secondary orange-yellow, with termen finely dark
and a])ex somewhat more broadly dark.
E.xpanse : 35 mm.
Hub. The type is in my collection from Mount Kebea, where it was taken
in the month of Jul}'.
91. Labanda crenulata spec. nov.
? . Head and thorax grey, darkly irrorated ; abdomen darker grey. Primary
grey, very finely irrorated with darker grey ; basal line dark, irregnlar, with a
pale grey exterior edging ; medial dark line oblique dentate, with a pale grey
interior edging ; posterior line excurved, crennlate, with a fine exterior paler
edging ; snbterminal line pale, interrupted, above the tornus a blackish patch
between the two latter lines, above which is another smaller blackish ])atch over
vein ti ; in front of the ajiex a pale grey patch on the costa, with a dark small
patch on its inner edge ; termen blackish. Secondary uniform dark grey.
Expanse : 30 mm.
Hab. The type from Ekeikei is in my collection. The species flies in March
and April. In the Tring Museum are other specimens from the Aroa River.
This insect is near L. griseonigra, but the lines all differ, especially the basal
and medial ones.
93. Alibama kebea spec nov.
c?. Head and thorax russet brown, collar banded with dark brown, abdomen
greyish brown. Primary rnsset brown, with darker brown lines and markings ;
basal line short and irregnlar, antemedial line bluntly dentate, medial line almost
obsolete ; postmedial line strongly excurved and serrate for its costal hall", receding
in a deep arch from vein 5 ; a series of blackish dashes in the terminal area edged
by short pale cream-coloured lines ; between the postmedial and antemedial lines
is a large very dark indefinite patch, inner margin broadly dark ; the position
of the reniform stigma is marked by an obscure cream-coloured exterior edging.
Secondary uniform dark brown.
Expanse : 31 mm.
Hab. The type is in my collection, being taken in the Kebea Range in March.
93. Barasa griseola spec. nov.
S. Head grey, collar dark grey broadly tipped with white, thorax pale grey,
darker on the metathorax, abdomen pale-greyish. Primary whitish grey for the
basal two-fifths, brownish grey for the terminal three-fifths ; antemedial black line
fine and interrujitod, broadly serrated ; jjostmedial line waved and crenulate ; a
trace of a strongly angled indefinite snbterminal line ; a large pale brownish patch
immediately precedes the crenulate line. Secondary subhyaline whitish, greyish
at the termen, more broadly so at the apex.
? . Like the male, but paler.
Expanse : 34 mm.
( 222 j
Hub. The \.\\)c is iu m}- collectiou from Ekeikei, where the sjiecies flies in
March and April. I have other specimens from higlier up in the Kebea Range
captured in May nud June.
'■'4. Barasa pura sjiec. no v.
? . Head white, finely irrorated with jiale ochreons brown, thorax and collar
white with fine ochreons scaling, abdomen whitish gre}'. Primar}* white with very
pale fine ochreons scaling in the median and postmedian areas ; costal half of
base pale chestnnt-brown edged by an oblicjne black line ; medial black line
irregnlar and interrupted in the cell, postmcdial black line j)rojected outwards
very obliquely from the costa to vein 5, whence it recedes irregularly to the
inner margin ; an indistinct snbterminal curved line, two black dots on the costa
before the apex, an apical dark rufous brown patcii exfemling to the postmedial
line, a palish rufous brown spot at the tornus ; the subterniinal area is slightly
greyish. Fringes white, darkly intersected. Secondary subhyaline white, terraen
dark brown, broad at apex, tapering rapidly to vein \b.
Expanse : 32 — 34 mm.
Ilab. The type is in my collection from Dinawa, where the species flies in
September. Other specimens I have are from Ekeikei and other localities, taken
in January and February.
95. Barasa nigrescens spec nov.
?. Head and thorax dark mottled grey, abdomen paler grey. Primary dark
grey, thickly irrorated with black ; basal area dark, margined externally with an
irregular black line ; medial line serrated, somewhat obscure, jiostmedial line
projected forwards from the costa to vein 7, from whence it descends slightly
receding to the inner margin as a dotted line ; beyond this is the snbterminal
line, broad from the costa to the angle of*vein 7, along which it extends considerably
and descends in a waved line to the tornns, indefinitely margined with palish grey;
termen dark, a short blackish dash at the end of the cell. Secondary subhyaline
white, termen dark grey, broad at the apex, rapidly tapering to vein \b.
Exjianse : 34 mm.
Hab. The type is in my collection from Dinawa, where the insect flies in May
and June.
9(5. Blenina owgarra sjiec. nov.
?. Head and thorax whitish grey, collar irrorated with cinnamon-colour,
abdomen cinnamon-grey, both wings almost pure white. Primary with base dark
grey ; median and postmedian lines irregular, nearly jiarallel, and enclosing a dark
grey area ; snbterminal line very irregular, preceded by a darkish grey shading ;
termen darkly dotted with a rusty red strijjc in front of it ; a rnsty red broad
indefinite stripe from the end of the cell to the termen. Secondary pure white
with a dark grey termen and broad dark apex.
Expanse : 38 mm.
Hab. The type is in my collection from Owgarra.
97. Blenina nigrans spec. nov.
cJ. Head pale grey mixed with green, collar and thorax l)lackish, abdomen
greyish brown with dark dorsal tufts. Primary lilackish grey; apical area, in
an oblique line from vein 3 across the end of the cell, jiale greyish ; basal black
( 223 )
line interrupted ; medial donble line interrn]ited, traceable below the cell ; post-
medial line more distinct, edged for the costal half broadly with white ; subterminal
line distinct, irregular dentate, behind which the costa is pale, followed below by
a broad short band of dark greenish ; a suffnsion of very dark green at the base.
Fringes green with white extremities, intersected with black. Secondary dark
brownish grey, with dark fringes except between veins 2 and 4, where they are
whitish.
Expanse : 32 — 34 mm.
Hah. The type is in my collection from Dinawa, where it was taken in
September. Other specimens are in the Tring Mnsenm from the Aroa Kiver.
_ 98. Blenina nigrans varians ab. nov.
$. Differs from nigrans in that the base of the primary is bright grass-green ;
the black basal line has a broad white external edging to the lower margin of the
cell ; the median area is rnsty brown with a broad edging of darker green, beyond
which is a broad band of whitish mixed with rusty grey, followed by a considerable
suffusion of green in front of the subterminal line ; the thorax and collar are pale
grey largely intermixed with green.
Hab. The type from Dinawa is in my collection. We have a fair series of
both forms.
99. Blenina viridata spec. nov.
c?. Head and thorax green ; abdomen dark yellowish. Primary green, paler
and mixed with white for the basal half, darker and mixed with blackish for the
outer half ; basal line interrupted, medial line very irregular ; postmedial line
irregular and interrupted, followed by broad obscure dashes of blackish between
the veins ; subterminal line irregular and dentate, obscurely palely edged ; area
beyond very dark ; near the middle of the inner margin is a whitish patch.
Secondary dark brownish grey, yellowish at the inner margin and near the base.
? . Similar to the male, but with au admixture of grey in the medial and
postmedial areas.
Expanse : S 32 mm., ? 34 mm.
IJah. The type is in my collection. The species was captured at Ekeikei
(15l>0 ft.) flying in January and February, and from high (0000 ft.) in the Kebea
Range in March and April. Other specimens are in the Tring Blusenm from the
Aroa River.
100. Blenina babooni spec. nov.
¥. Head, thorax, and abdomen dark grey. Primary dark slaty grey, with a
subbasal costal pale grey patch ; basal line indistinct ; medial, postmedial, and
subterminal lines edged with dark ochreous, each being somewhat interrupted
and irregular ; between the two latter is a curved darkish line; the ochreous edging
is prominent in the subterminal area. Secondary dark brownish grey.
Expanse : 28 mm.
IJab. The type is in my collection from Babooni, where the species flies in
September.
lol. Blenina smaragdina spec. nov.
(J. Head, collar, and thorax jiale green liarred with black; centre of thorax
whitish; abdomen dark grey. Primary pale green with a basal black dash; a
Bubbasal black costal spot, followed by a white spot ; medial line black, interrupted,
15
( 224 )
edged internall\- with white, aud with a broad white dash below the cell, in a line
with which, on the external side of the medial line, is a broad black dash, edged
above with white to tlie postmedial line, which is very irregular and edged
externally finely with white, a black mark at the end of the cell continent with
the postmedial line ; this mark is also edged with white internally ; a broad
subterminal white band to the tornns, where there is a black spot edged with green
externally : an apical and subapical black patch and a small black spot on the
termen aliont vein 3 ; fringes pale greenish, intersected with white and with black.
Secondary darkish grey, paler near the base.
Expanse : 40 mm.
Ilab. The type is in my collection from Babooni, where the species flies in
September.
102. Hypothripa distincta spec. nov.
?. Head white with tips of scales piuk ; thorax whitish with an admixture of
pink, barred with black ; abdomen grey. Primary white, witii a suftusion of pale
brownish pink in the antemedial and postmedial areas and a suffusion of dull lilac
in the lower part of the terminal area ; two blackish spots at the base and a
blackish dash near the base on the inner margin : a broad oblique blackish invaded
band across the median area ; a blackish patch, followed by a small black spot on
the costa beyond its centre ; a black waved subterminal line, a dark suffused patch
at the apex and just below it, a white costal spot immediately in front of it.
Secondary whitish, darker grey at the apex and termen.
Expanse : 22 mm.
Bab. The type is in my collection from Dinawa, where the insect flies in
September.
1U3. Cletthara multilinea spec. nov.
?. Head and thorax whitish, abdomen grey. Primary whitish grey, with a
subbasal black irregular and serrate line ; medial black line very strongly dentate,
connected by two oblique dashes with the very irregular postmedial line ; sub-
terminal black line strongly waved ; termen black with spearhead marks ; fringe
white, intersected with grey. Secondary whitish grey, pale at the base.
Expanse : 24 mm.
Hab. Area River, March.
Type in the Tring Museum.
Iit4. Sarrothripa aroa spec. nov.
(?. Head, thorax, and abdomen blackish, the thorax barred with white.
Primary white with black lines ; base darkish with a basal black line ; antemedial
line oblique, serrate, almost touching the double medial line at the inner margin ;
medial lines rise in a black costal patch ; postmedial line double dentate, rising
in a black costal patch and ending in a dark cloud ; subterminal line very
irregular, strongly serrate and dentate in part; beyond this is a grey line of
shading nearer the termen ; termen with a fine black line. Secondary greyish,
darker near the termen.
Expanse : 22 mm.
Hab. Area River, February.
Type in my collection. Other specimens are in the Tring Museum.
This species will come near S. mesoplaya Hpsn,
(225 )
105. Sarrothripa brunneicosta spec. nov.
S- Head and thorax greyish, abdomen darker. Primary darkish grey with a
broad white serrate costal line ; a fine double dentate median line, a double
irregular dentate postmedial line — both these two double lines filled in with whitish ;
a serrate snbterminal line rising in a whitish apical patch ; from this patch an
oblique pale area crosses the wing ; reniform darkly ontliued ; costa broadly
brownish grey with a dark grey spot touching the reniform. Secondary grey, pale
at the base.
Expanse : 24 mm.
Hab. Aroa Biver, .January.
Type in the Tring Museum.
This species will come next S. grisea Hpsn., but the medial and postmedial
lines differ.
ion. Sarrothripa arcuosa spec. nov.
? . Head, thorax and abdomen grey, collar brownish grey. Primary dark
grey, with a considerable suffusion of dark olive-green scales ; near the base is a
fine arc of whitish, beyond which the ground-colour is dark olive-green ; two dark
dots in the cell, in a patch of pale greyish ; postmedial line dark greenish, irregular,
and produced outward beyond the cell, edged below with whitish ; following the
postmedial line is a row of dark dashes on the veins, subapical area whitish grey ;
snbterminal line somewhat indistinct in the tornal area. Secondary dark grey.
Expanse : 20 mm.
Hah. The type is in my collection from Dinawa, where the species occurs in
August.
107. Sarrothripa kebea.
J. Head whitish ; collar brownish at base, black beyond tipped with white ;
thorax whitish, largely jiatched with blackish ; abdomen dove-grey. Primary
blackish, with an antemedial and medial oblique bar of whitish invaded and serrated
by the black ground ; extreme base whitish ; two oblique costal dashes of white at
the apex, the hindmost of which touches nearly at right angles a white dash beyond
the cell ; snbterminal line obscure ; tornal area up to vein 5 pale grey. Secondary
white for basal half, becoming dark grey beyond.
Expanse : 30 — 32 mm.
Hrib. The type is in my collection from the Kebea Range, where the species
occurs in March and April. I have specimens from other localities taken in August.
108. Sarrothripa avola spec. nov.
c?. Head and thorax wliite, slightly tinged with green and finely barred with
black ; abdomen pale grey. Primary greenish white, with about eight very fine
oblique irregular dentate or serrate black lines ; the innermarginal and terminal
areas are darker greyish ; the termen is spotted alternately with black and white.
Secondary white for the basal half, becoming dark grey beyond.
Expanse : 29 mm.
Ilab. The type is in my collection from Avola, where the species occurs in
August.
( 226 )
109. Dendrothripa venalis spec nov.
(S. Head and collar white mixed with ochreous, tlie latter with a darkish
patch on each side ; thorax and abdomen sjrey. Primary dark grey of a speckled
appearance ; hasal line very obscure ; medial line somewhat waved, but angular ;
postmedial line irregular, projected outwards into an acute angle about vein 3 ;
innermarginal area between these two lines filled with whitish into the angle just
mentioned ; postmedial line somewhat palely edged, followed by a band similarly
angled of very dark brownish grey ; subterminal line irregular, edged externally
with whitish. Secondary greyish.
Expanse : 22 mm.
flab. The type is in my collection from Dinawa, where the insect occurs in
September.
110. Exyra dinawa spec. nov.
(?. Head ochreous intermixed with dark brown scales, collar ochreous with
a dark brown band ; thorax ochreous intermixed with dark brown scales ; abdomen
ochreous grey with a fine dorsal ridge of dark lirown. Primary pale greyish
ochreous, with interrupted lines of dark brown scales : medial line waved, pale,
edged with crimson brown on its lower external side; a pale ochreous brown patch
on the coBta over the cell edged with whitish, the outer edging being part of the
postmedial line, which is interrupted and has on its inner side a pale dark brown
curved mark ; a pale whitish costal patch adjoining the postmedial line, beyond
which the whole wing is very dark, with the subterminal line irregular, very dark
edged palely as to its upper portion. Secondary pale greyish brown, darker towards
the terminal area.
Expanse : 22 mm.
Bab. The type is in my collection from Dinawa, where it was taken in August.
111. Ingura pratti spec. nov.
(?. Head, frons purplish, vertex pale brown ; collar pale brown broadly tipped
with lilac ; thorax brownish lilac ; abdomen lilac-brown. Primary jiale lilac up to
the median line and np to the postmedial line along the costa ; postmedial area
brownish ; terminal area pale greyish for the apical half, pale brownish for the
tornal half ; basal and median lines very fine, almost obscured ; postmedial line
double, dark brown, projected outwards rapidly from the costa to below vein 7,
receding along vein 5 and descending irregularly to the miildle of the inner margin,
where there is a dark grey patch on the outside of the line ; a faint trace of a
subterminal pale line. Secondary darkish grey, slightly paler near the termen.
? . Like the male, but greyer.
Expanse : 6 33—3(5, ? 35—36 mm.
Hab. The type is in my collection. The species occurs in the Kebea Range
and Babooui in March and April, and again in August and September.
The lilac colour ai)pears to fade raiiidly, for we have some specimens which are
very much paler and greyer than the types : the species will come next cyanescens
Turner.
11 J. Ingura costistrigata spec. nov.
S. Head and collar jjale chestnut-brown, the latter barred with slate-grey;
thorax darker chestnut-brown, sparingly mixed with slate-grey ; abdomen dark
slate-grey. Primary pale chestnut-brown, with the base, and the costa broadly
( 227 )
purplish shite-coloiir ; medial line dark brown, fine; postmedial line projected
outwards into an acute angle about veins 0 and 7, receding rapidly in a dentate line
to the middle of the inner margin, with a small grey patch on the margin on the
outside of the line ; apex with a whitish lilac patch ; snbterminal line dotted.
Secondary greyish brown, witli a ])aler, somewhat pinkish, termen, with a row of
dark points.
? . Exactly like the male.
Expanse : c? 34, ? 36—37 mm.
Hnb. The type is in-my collection fiom Diuawa, where the species occurred iu
August; we have sjiecimens from other localities in January and February.
113. Calling'ura kebeae spec. nov.
(?. Head lilac-grey, crested ; collar lilac-grey, narrowly barred with purplish
brown and broadly tipped with jiale ochreous brown ; thora.x purplish slate-colour ;
patagiae with a triangular shoulder-patch of ochreous cream-colour ; abdomen pale
silvery grey, pale straw-colour beneath. Primary dark olive-green, with a broad
horizontal ochreous cream-colonred stripe, edged above with greenish, and for its
outer half edged above also with a cnrved pink strijie ; a third from the base, on
the inner margin, rises a broadish strongly curved ochreous cream-coloured stripe,
meeting and coalescing with the horizontal one : both these stripes are finely
margined internally with black ; termen broadly pinkish lilac with a fine double
darker line. Secondary pinkish brown, pale at the base.
? . Like the male, but paler, and greeu more ochreous.
Expanse : S and ? , 34 — ^36 mm.
Hab. The type is in my collection. The species occurs higli (0000 ft.) in the
Kebea Range in March and April, other specimens from Dinawa being taken in
August. In the Tring Museum are others from the Aroa Kiver, taken in February
and March.
114. Targalla catephioides ekeikei snbspec. nov.
Differs from Hampson's species in its melanism ; the base up to the median
line is very dark blackish brown, and the rest of the wing is dark brown ; the
reniform stigma has a greenish hue in fresh specimens, degenerating into ochreous
brown. The secondary is blackish grey, with a small white basal innermarginal
patch.
In quite fresh specimens there is a dark bronzy-green scaling over the
thorax and abdominal tufts, and to a less extent over the primary, but this
evidently soon wears off. This melanic form is quite constant in all the collections
received, and has evidently developed into a local race.
Hab. All the localities in New Guinea.
Type is in my collection, and in the Tring Museum are other specimens.
115. Targalla pratti spec. nov.
S. Head whitish grey; thorax grey, barred with dark grey; patagiae with a
white line across, followed by a broader brown band ; thorax grey, darker on the
proximal segments. Primary pale grey ; base very dark grey, margined externally
by the antemedial black line ; medial line represented by an obscure line of darker
shading produced outwards at the end of the cell and then receding somewhat ;
postmedial line double, the inner one fine black, beyond which is a small triangular
( 228 )
dark patch on the costa, which is the origin of an interrni.ted snbtermiiial dark
line edged externally with whitish ; termen cronulate, finely blackish. Secondary
whitish grey with dark veins, termeu broadly darkisli grey.
Expanse : c? 30, ? 37 mm.
Hub. The type is in my collection from Ekeikci.
110. Eutelia hampsoni spec, no v.
? . Head greenish grey and thorax greenish grey, with patagiae rusty-red ;
abdomen dark brownisli. Primary of a general tone of greenish grey or brown ;
base dark, followed by a pale band, succeeded by a broad dark band ; median area
pale dirty olive-green, with two fine darker lines, that near the base reaching only
to vein 1 ; this pale area is margined externally by a broadish indefinite line of
very dark greenish brown ; postmedial line irregular, edged externally with white,
produced into a sharp acute angle on vein 3; a trace of a subterminal Hue
interrupted, white in the costal area ; apex up to postmedial line whitish i,'rccn.
Secondary dark grey, with a trace of a medial line.
Expanse : 22 mm.
Hab. The type is in my collection from Ekeikei, where the insect flies in
January and February.
117. Eutelia mediofusca spec nov.
S. Head greyish pinkish ochreons, collar jiinkish ochreous, thorax pinkish
ochreons with pinkish patagiae, abdomen dark grey. Primary pale ochreons o-rev
upper part of base narrowly dark ; median area dark brownish, receding slightly
below the greenisli-reniform stigma, edged laterally with greenish brown ; a white
point in the cell ; postmedial area whitish, intersected by the oblique dark line
edged with white from the costa to the termen on vein 2 ; the lower half of this
line is edged with pale ochreous grey, a dark triangular spot is above it on the
costa ; termen broadly greyish, crennlate, with a linear blackish edging. Secondary
greyish, paler towards the base.
Expanse : 25—28 mm.
Hab. The type is in my collection from Dinawa, where the species occurs in
September.
lis. Eutelia olivaceiplaga spec. nov.
S. Palpi rusty-red for the basal third, lilac-grey for the apical two-thirds;
head lilac-grey tipped slightly with rufous ; thorax lilac-grey with a rnfous central
stripe ; abdomen rnfous lirowc. Primary pale jiinkish lil;ic, with a short basal
white stripe ; antemedial line at an opjiosite angle rnfous brown, edged internally
with white ; median line broad, rnfous, angled outwards at the cell : postmedial
line rnfous, edged externally with ochreous; orbicular stigma olivaceous, jinjiillcd
with ochreous; reniforra olivaceons, outlined finely with ochreons; a whitish round
patch on vein 2 outside the jiostmedial lino, and a triangular olive-brown costal jiatch
edged with white, below which are two obli(iuely placed similar coloured dashes
reaching nearly to the termen above vein 3, above which the apical area is [lale
lilac-grey.. Secondary pinkish grey, jiale greyish at the inner margin and base.
Expanse : 28 mm.
Hab. The type is in my colh'ction from Diaawa, when' the sjiecies occurs in
September.
( 229 )
119. Eutelia dinawa spec. nov.
c?. Head pale cbestnut-brown, thorax chocolate-brown, abdomen paler than
thorax. Primary purplish bruwu, paler at base with an olivaceous costal mark;
antemedial line olivaceous, broad, irregular, and serrate ; medial line represented
by a broad band of darker shading ; postniodial and snbtermiual lines olivaceous,
serrate, the former almost crenulate in the radial area, the crennlations being
marked by white jioints ; both these lines rise in a triangular costal patch, which
is edged with ochreous, margined on the inner edging with white ; a white point in
the cell followed by the olivaceous reniform stigma. Secondary brown with pinkish
fringes ; the fringes of both wings are crenulate.
? . Like the male, but paler and greyer.
Expanse : <? 31, ? 28 mm.
Hub. The type is in my collection. This species was taken at Ekeikei in
January and February, and Dinawa in September. In the Tring Museum are other
specimens from tlie Aroa River.
The species is near E. ajiij'ascia Hamjis., but it lacks the ochreous spot beyond
the reniform, and the lines differ in direction.
120. Eutelia kebeae spec. nov.
cJ. Head crimson-brown ; thorax and abdomen dark jjurplish brown, the latter
reddish beyond the middle. Primary dark purplish brown, with traces of basal,
antemedial, and medial lines on the costa, the latter two of which develop below
the cell into broad irregular dentate very dark brown lines with pale interiors ;
adjoining the medial line is an oblique band of pale pinkish ochreous extending
into the olive-green reniform stigma, edged internally with white ; postmedial line
double, very dark, with ochreous interior, angled outwards about vein 7, followed by a
fine dark line roughly parallel with it ; subterminal line obscure, rising in a dark
subapical costal triangular patch darkly dotted below ; apex and the whole of the
tornal and radial areas lilac grey ; a white dot in the cell in front of the reniform ;
termen with white points at the end of the veins. Secondary dark greyish brown,
pale towards the base ; termen with white points at the ends of the veins.
? . Like the male.
Exi)ause : c? 3U, ¥ 29 mm.
Hah. The type is in my collection. This species occurs in the Kebea Range
and Ekeikei in March and April.
121. Eutelia albicomma spec. nov.
(J. Head ochreous tipped with dark crimson, thorax and abdomen dark
crimsonish brown. Primary sepia brown, with pale basal and antemedial lines,
very irregular, subdentate and somewhat obscure ; a broad pale pinkish medial
band, followed by a fine double dark postmedial line angled outwards beyond the
cell, followed by a fine whitish interrupted line in which is a large white comma at
the tornus ; a trace of an anteterminal line close to the previous one, both of
which arise in a dark triangular white-edged costal patch ; terminal area dark ;
termen with white points alternating with dark lunular marks ; a white point in
the cell in front of the pale greenish reniform stigma, which is edged internally
with white. Secondary brownish grey, pale' at the base ; a trace of a pale waved
( 230 )
line at the end of the cell ; termeii with wiiite jioints ; fringes somewhat creuulate.
Expanse : 30 mm.
Hub. The type is in my collection from Eljeikei, where the species occurs in
January and February.
122. Eutelia purpureonigra spec. nov.
?. Head, thorax, and al«louieu rich iiui]ile brown intermingled with crimson
scales. Primary rich darlc crimson brown for the basal half, with slightly paler
purplish base ; subbasal and medial lines dark, the latter waved, postmedial line
double, somewhat waved, and produced outwards beyond the cell, followed by a
fine crenulate line ; snbteriuinal line arising in a dark costal triangular patch,
serrate, somewhat obscured near the tornus ; reniform stigma somewhat ochreous.
The terminal half of the wing is paler and purplish ; teruien with white points
at the ends of the veins. Secondary tiark greyisli brown with crinisonish fringes.
Expanse : 34 mm.
llab. The type is in my collection from Ekeikei, where the species occurs in
January and February.
123. Eutelia viridata s])ec. nov.
?. Head aud collar greenish, thorax and abdomen jiale ochreous brown.
Primary pale yellowish green, with a small greyish patch margined by the basal
and by the double antemedial line ; medial line marked by an indefinite line of
grey, produced outwards at the cell ; jjostmedial line of the same shape, blactkish
near the inner margin ; apex pale bluish white with a dark dash in it, a small
pale brownish patch between veins 5 and 6 ; the whole of the medial area is very
pale whitish ochreous brown, margined laterally by the antemedial and postmedial
lines ; reniform stigma greenish ; termen finely dark with white jioiuts at the
veins. Secondary darkish grey, pale at the base, with an obscure pale medial line.
Expanse : 26 mm.
Ilab. The type is in my collection from Dinawa, where the species occurs
in August.
I am doubtful if this species is a Eutelia ; the end segments of the jialpi are
as long as the second ; the thorax and abdomen are quite slight in build ; the
wings, i.e. the primaries, are narrow, expanding considerably towards the termen.
As 1 have only one sex, aud that the female, I refrain from creating a new genus
for it at present.
124. Eutelia deletoides spec. nov.
?. Head grey, collar grey mixed with ochreous, thorax grey, abdomen grey
with black dorsal marks. Primary pale grey, with darker obscure basal line;
median line from a dark cell-spot to the inner margin, a small dark dot precedes
this spot ; jiostmedial line broad, somewhat indefinite, slightly brown, especially
at the inner margin ; subtermiual line broad, dentate and waved, edged externally
with whitish ; termen finely dark ; fringes whitish, intercepted with blackish.
Secondary dark grey, paler at the base. Fringes white intercepted with black.
Expanse : 24 — 29 mm.
llab. The type is in my collection from Ekeikei, where the insect occurs in
March and April ; other specimens in the Tring Museum, from the Aroa River,
February.
(231 )
125. Eutelia ochreiplaga spec, no v.
d. Head, thorax, auil abdomeu brownish grey, with short ochreous dashes.
Primary brownish grey, witli an ochreous line along the fold ; base with an
ochreous spot, an irregular dark subbasal stripe ; median line dark, produced
outward at the cell, tlien obliquely basewards ; postmedial line excnrved beyond
the cell, with ochreous patches on each side of it. Secondary brownish grey,
paler towards the base.
Expanse : 27 mm.
Uab. Aroa River, February and April.
Type in Tring Museum.
This species comes near E. ocularis Btl.
12(']. Penicillaria meeki spec. nov.
S. Head, thorax, abdomen, and primary dark purplish grey, the latter with
an oblique straight line darkly centred ; an indistinct angled (near the costa)
postmedial line ; a short fine j)ale curved dash in front of the apex and at the
tornus. tSecondary pure white, with a very broad purplish termen and a dark
spot in the cell.
Expanse : 24 mm.
Ilab. Aroa River, April.
Type in Tring Museum.
127. Penicillaria dinawa spec. nov.
c?. Head, collar, thorax, and abdomen pale uniform lilac-brown. Primary
pale lilac-brown, with a double dark oblique median line. Reniform stigma dark ;
postmeiliai line dark, angled outwards on vein 7, thence descending obliquely into
the tornus ; apex with a short curved white stripe across it ; tornus with an obscure
whitish erect line from vein 1 to vein .3. Secondary white, with a dark spot in the
cell ; termen broadly pinkish grey.
Expanse : 26 mm.
Ilab. The type is in my collection, from Dinawa, where the species flies iu
September.
Tibiocillaria gen. nov.
Differs from Fenicillaria or Kutelia in that the second segment of the palpus
is fringed with a long brush of hair ; end segment short and thickly clothed
with hair ; collar developed into a pyramidal prominent tuft, standing high above
the thorax and extending over it ; tiiorax with a metatlioracic tuft ; abdomen
with a tuft on the proximal segment and a long tubular tuft on the anal
segments. The fore- and midtibiae have each a very long and heavy tuft of hairs—-
a tuft on the end of the hindtibia is small ; mid- and hiudtarsi tufted. Wings :
primary, a])ical part of termen nearly erect to vein 3, where it is angled and
somewhat excavated below. Secondary, vein 2 produced into a slight tooth,
excavated slightly below.
Type : TihiociUaria pratti B.-B.
128. Tibiocillaria pratti spec. nov.
c?. Head purplish, tufted with chestnut l)rown ; tufted collar and thorax deep
purplish plum-colour, deep reddish brown or iiurjilish brown ; abdomen dark grey;
legs deep purjilish, with the long tibial tufts jiale chestnut ; puri)lish below at the
( 232 )
base of the underside ; tarsi ringed witli wliite. Primary deej) purjilish or ])nr]ilisb
grey ; onter half of oosta and cell, also the ajicx and teriuen to vein (i, finely
dusted with ochreons; lower part of base dark rusty-red ; medial and postmedial
lines, each double, finely whitish lilac, strongly angled externally about the upper
margin of the cell, the latter slightly angled in the opposite direction on vein 2 ;
a dark velvety spot at the ajiex ; a curved dark broad dash above vein 3, with an
inner deep rusty-red dash, both interrupted by the jiostmedial line ; an erect curved
whitish obscure line from the toruus to vein 2. Secondary white with a dark
cell-spot ; termen broadly deep purplish, becoming grey at the apex.
?. Exactly like the male, but the tufts are smaller.
Expanse : 3 and ? , 35 — 36 mm.
Hab. The type is in my collection, from the Kebea Range, taken in July ;
other specimens from Dinawa, August. 1 have specimens also from the former
locality that were taken in March and April.
120. Tibiocillaria aureiplaga spec. uov.
(?. Head, collar, and thorax purple, abdomen ochreons, legs and tufts purplish,
tarsi ringed with white. Primary purple, with a trace of a basal line ; medial
and postmedial lines obscure for their costal portion ; a bright golden yellow patch
in the middle of the inner margin and in front of the apex ; beyond the former
the ground colour golden yellow finely irrorated with superimposed purple scales ;
a curved white line in front of the apex on the costa, and also one rather obscured
rising from the tornus to vein 2. Secondary white with a dark cell-spot ; termen
broadly purjile with a white line and a yellowish patch at the tornus.
? . Like the male.
Expanse : cj 2o, ¥ 27 mm.
Hab. The type is in my collection, from Ekeikei, where the species occurs
from January to April.
130. Tibiocillaria dinawaensis sj)ec. uov.
c?. Head reddish chestnut, thorax reddish orange, abdomen pale brownish.
Primary pale reddish with an oblique straight dark median line confluent with a
dark cell-spot ; jiostmedian line fine, dark, produced outwards beyond the cell ;
termen purplish ; a yellow patch in the middle of the inner margin, a yellow
curved line before the apex, a white line rising out of the tornus to vein 2.
Secondary white, uo spot in the cell ; termen broadly purplish, with a white sj)ot
in the tornus below an orange one.
Expanse : 20 nun.
Hah. The type is in ray collection from Dinawa, where tlie species occurs in
September. Other specimens are in the Triug Museum from the Aroa Itiver,
March.
1;'>1. Maceda rufescens spec nov.
?. Head, thorax, and abdoiuen uniform fuscous. Primary purplish grey, with
base, costal part of apex, and tornal area orange red ; in the latter is a dark dash
at the tornus ; a basal dark line ; antemedial line bordering the red area angled
inwards and broadly dark,; postmedial line serrate, burderiiig inwardly both the
other orange-red areas, between which is a patch of dark purplisli grey expanding
( 233 )
to the termen ; a diivk point in tlie ct^ll. Secondary davlc brownish grey, paler
in the cell, which has a dark spot ; fringes wliite from vein 5 to vein 2.
Expanse : 38 mm.
Hab. The type from the Kebea Range is in my collection. The species flies
in March and April ; and in the Triug Mnsenm from the Aroa ]?iver are other
specimens.
132. Risoba delicata spec. nov.
c?. Head very pale greenish ; collar white, with intermixed green and grey
scales ; thorax very delicate and pale green, barred with white ; abdomen pale
grey. Primary white, snifnsed with very delicate pale green ; snbbasal area
suffnsed with pale green ; basal line very obscnre ; a short fine black basal costal
dash ; reniform greenish ; postraedial line very finely dark and creunlate, beyond
which is a short serrated black line from below the costa, and a short broad
erect blackish dash from before the tornns ; apex darker green, with an internal
black dash ; a row of fine black scallops in front of the termen ; fringes grey
intersected with white. Secondary white with a dark cell-spot ; termen broadly
dark, tapering somewhat to the tornns.
Expanse : 39 — 40 mm.
Ilab. The type is in my collection from Dinawa, where it was taken in
September.
133. Risoba viridata spec. nov.
c?. Head bright green; collar pale brown, with a fine black basal stripe
surmounted with a broad bright green band interrnpted in the middle ; thorax
green, barred finely with black, with a prothoracic brown patch ; abdomen pale
brownish, greenish laterally ; dorsum green in front of the anal segments.
Primary, base bright green, with a black basal serrated line, followed by a
broad whitish area, beyond which the wing is mainly grey, with the reniform
green ; a broad green dash above vein 1 and beyond the reniform, from whence
lip to the apex the ground is suffused with green; postmedial line finely dark,
crennlate ; a more or less obscure green subterminal stripe ; a row of short dark
dashes in front of the termen. Secondary creamy white, with a dark cell-spot
and a broad dark termen.
Expanse : 40 — 42 mm.
Hab. The type is in my collection from Dinawa, where the species occurs in
May and June.
134. Risoba kebea S2)ec. nov.
c?. Head greenish brown ; collar greenisli brown, with greenish base ; thorax
brown, with greenish patagiae ; abdomen jiale brownish, darker laterally. Primary
olive-brown, with traces of greenish suffusion in parts ; base whitish, with a short
black dash at hase of cell, below which is a pale brownish patch ; reniform greenish,
edged with white, below which is a white stripe darkly edged externally ; beyond
the reniform is a pale greenish ochreous siripe, with a fine internal white line
almost confluent with the dark apical jiatcli, which is edged below with white;
a subterminal row of dark dashes edged internally with white. Secondary creamy
white, with a dark cell-spot and a very broad dark termen.
?. Similar to the male, but much paler and without the white stripe below
the reniform.
( 2:^-t )
Expanse : c? 42, ? 44 mm.
Huh. The type is in my collection from tlic Kebea Rang'e, where it was taken
in July ; other specimens from Dinawa, August, and in the Tring Mnseum from
the Aroa River, January to April.
135. Risoba avola spec. nov.
cJ. Heed and collar very pale lavender-brown, the latter edged with rusty-
red; thorax greyish ; patagiae edged with wlute ; abdomen greyish, with a reddish
tuft. Primary pale brown, snffnsed with greenish— below the cell this suffusion
is almost absent ; base narrowly and obliquely whitish ; a trace of a dark serrated
medial line, followed by a second not extending below the cell ; beyond tlie cell
(not extending above it) is a broadish white stripe, edged by a fine dark line,
beyond which the radial area is slightly rufous ; a dark apical patch, edged
externally by an irregular whitish line and below by a curved greenish line ;
the usual row of dark subterminal dashes, edgeil internally witli white. Secondary
subhyaline white, with a dark cell-spot and a slight dark short broadisli termen.
Expanse : 34 — 36 mm.
Hab. The type is in my collection from Avola, where it occurs in August.
Other specimens are in the Tring Mnseum from the Aroa River, February.
136. Eisoba olivens spec. nov.
c?. Head and collar lilac-grey ; thorax dark grey, with pale jjatagiae ; abdomen
dark lilac-grey. Primary uniform olive-green, except at the base, which is white
with an olive-green innermarginal ])atch, and beyond the cell, where it is grey up
to the dark green ai)ical area ; reniform finely encircled and pnpilled with dark
brown ; apical area edged internally with a dark curved broadish stripe, below
which is a short fine serrated dark line ; below the apical patch the ground colour
is dashed with white ; the usual subterminal row of dark dashes ; the postmedial
line is finely serrated and strongly curved inwards below the cell. Secondary
creamy whitish, with a dark cell-spot and a broad dark terraen.
Expanse : 36 mm.
Hah. The type is in my collection from Dinawa, where the species occurs in
September ; and in the Tring Museum from the Aroa River, March, are other
specimens.
137. Risoba grisea spec. nov.
¥. Head and thorax grey, i>atagiae whitish, abdomen greyish. Primary with
base narrowly and obli(iuely white; the rest of the wing grey, darker near the
base ; a dark dot in the cell ; reniform finely and darkly encircled and pui)illed ;
postmedial line waved oblique, slightly serrated, on each side of which the area is
somewhat whitish ; apex brown, edged internally with white, which edging has a
dark internal streak below it ; two dark dashes from the aj)ex ; subterminal area
pale brownisli ; the usual subterminal row of dark dashes. Secondary subhyaline
creamy-white with no cell-spot, the broad dark termen evenly cnrvcd internally.
Expanse : 33 mm.
Ilah. The type is in my collection from Ekeikei, where it occurs in March and
A]iril ; other s])eciniens are in (he Tring Museum from the Aroa River, .January
to March.
( 236 )
138. Risoba pratti spec. nov.
c?. Head brownish, collar j)ale brownish grey with a rufous base surmounted
by a greenish band, thorax greyish brown, abdomen pale Ijrown-grey laterally.
Primary greyish brown, with an oliliqne cliestnut-red base ; a broad creamy white
subcostal stripe ; costa greenish ; renit'orm obscure greenish with an oblicjue white
stripe below it, edged externally by the postmedial fine oblii|ue dark line, palely
edged externally below, with its costal half edged broadly with greenish ; apical
dark area small with a whitish suffusion below it ; the usual subterminal row of
dark dashes very distinctly edged internally with white. Secondary creamy whitish
with a dark cell-spot and very broad dark termen.
Expanse : 41 mm.
Hab. The type is in my collection from Dinawa, where it occnrs in August
and September ; other specimens are in the Tring Museum from the Aroa River,
March.
13'.>. Stictoptera ekeikei spec. nov.
Head and thorax very dark brown, abdomen blackisli brown. Primary rufous
brown, with a dark costa ; medial, postmedial, and subterminal lines very fine and
obscure, finely crenulate ; the terminal half of the wing has the appearance of
having a series of these finely crenulate lines over it ; the costal area is dark.
Secondary opalescent hyaline, with termen broadly blackish.
Expanse : 34 mm.
Hab. The type is in my collection from Ekeikei, where it occurs in March
and April.
140. Stictoptera rufa spec. nov.
c?. Head and thorax dark reddish brown, abdomen sooty brown. Primary
pale rnfous, with antemedial, meilial and postmeilial darker oblique lines, the latter
slightly waved ; a trace of a subterminal line of darker shading. Secondary dark
uniform sooty grey.
Expanse : 32 mm.
Hab. Aroa River, February.
Type in the Tring Museum.
141. Stictoptera meeki spec. nov.
(?. Head and thorax pale rufous, collar deep crimson, abdomen sooty grey.
Primary with base orange, very bright at its margin with the oblique antemedial
line ; rest of wing reddish, inclined to orange in the postmedial area ; medial line
waved ; postmedial line waved inwardly dentate ojiposite the reniform, which is
somewhat obscure ; subterminal line dentate, irregular. Secondary sooty black.
Expanse : 38 mm.
Hab. Aroa River, February.
Type in the Tring Museum.
142. Sadarsa purpurascens spec. nov.
(?. Head and collar deep purplish black, nietathorax purplish grey, abdomen
purplish black. Primary with the basal two-fifths of costa broadly deep jinrplish
black, paler beneath ; remainder of costa broadly pale fawn-brown tinged with
purplish ; innermarginal outer three-quarters purj)lish ; a medial line e.xcurved on
( 236 )
the cell ; a postmcdial waved line ; a trace of a subtermiiial and teimiual dotted
Hue ; termen finely blackish. Secondary dark greyish brown, paler in the
cellular area.
Expanse : 30 mm.
Hah. Aroa River, JIarch.
Type in the Tring Mnsenm.
Subfamily GONOPTERINAE.
143. Cosmophila kebeensis sjiec. nov.
(J. Head and thora.x pinkish red, abdomen grey. Primary grey suffused more
or less with red, especially on the outer two-thirds ; medial line fine dark grey from
the upper margin of the cell, slightly waved below the cell ; postmedial line almost
straight from the darkish reniform ; a fine waved line from the costa to vein 4,
where it is angled sharply inwards, ending abrujitly on vein 3 ; beyond this is a
slight grey scaling. Secondary uniform pale brown.
Expanse : 37 mm.
Hub. The type is in my collection from the Kebea Range, where it was taken
in July ; other specimens are in the Tring Museum from the Aroa River, March.
This species is near C. vitiemis Btl.; but, irrespective of the difference in colour,
the shape and direction of the lines differs also.
144. Cosmophila aroa spec nov.
(J. Head and thorax cinnamon-brown, abdomen brown with dark grey anal
tufts. Primary cinnamon-brown, sparingly irrorated with dark brown ; the dark
medial line oblique, slightly curved up to the lower margin of the cell, above which
it recedes and is V-shaped, the point of the V being basewards ; postmedial line
broadish oblique, slightly curved, being angled slightly outwards about veins 8 and 9;
a slight snbterminal shading from before the apex to veins 3 or 4 ; the orbicular
is a small white spot ; the reniform is represented by a darkish dash with a black
external dot. Secondary uniform jiinkish brown.
?. Like the male, but paler, the primary being pale ochreous brown.
Exj)anse : 3 40 mm. ; ? 43 mm.
Hub. The type is in my collection from tlie Kebea Range ; other sjiecimens
from the Aroa River, taken in March or Ajiril from both j'laces.
145. Cosmophila angulata spec. nov.
cj. Head, thorax and abdomen greyish ociireous. Primary greyish ochreous
with yellowish brown lines ; medial line angulated sharply from the costa to tlie
lower margin of the cell, whence it descends in a slight curve to the inner margin ;
postmedial line obtusely angled at vein il, whence it descends obliquely inwards to
near the lower angle of the cell, below where it takes a sliglit curve to the inner
margin ; beyond this line the ground colour is paler ; a subtermiual stripe of darker
shading roughly parallel with the termen, which is yellowish brown. The wing is
slightly irrorated with brownish. Secondary jiiukish brown, paler towards the base.
¥ . Like the male, but paler.
Expanse: J 44mm.; ¥ 41 mm.
Hab. The type is in my collection from the Kebea Range, where it was taken
iu= July.
( 237 )
14(5. Cosmophila subpurpurea spec. nov.
(?. Head and collar ochreons pink, the latter with fine brown edging palely
tipped ; thorax reddish brown ; abdomen gre3-ish brown. Primary junkish brown,
with pale sepia brown shading bordering internally all the lines ; medial line
fractnred at the lower margin of the cell, below where it takes an irregnlar curve,
but is shifted inwards above the cell-margin, and ascends obliquely to the costa;
postmedian line oblique, nearly straight, e.xcept at the costa and inner margin, where
there is the least bend ; from the costa to vein 3 is a curved line, slightly angled
from the costa at vein 9; subterminal line with an ontward bend between veins
3 and 5, from whence it recedes gently to each margin ; a white point in the cell.
Secondary pinkish, becoming cream-coloured on the inner margin and towards the
base. Underside pinkish purple, exce])t at the inner margins, whicli are cream-
coloured.
Expanse : 46 mm.
Flab. The tyjie is in my collection from Dinawa, where it was taken in
September. I have specimens from other localities taken in July and also in March ;
and in the Triug Museum are some from the Aroa River, February.
147. Cosmophila ekeikei spec. nov.
cJ. Head and thorax orange-red ; abdomen pinkish brown. Primary orange-
red ; antemedial line grey, with a slight outward curve, beyond which the ground
colour is less bright; medial line waved, descending from the prominent dark
reniform to the inner margin and receding slightly basewards ; an irregnlar fine
postmedial line from the costa to vein 2 or 3 ; in the type it terminates on vein 3 ;
a subterminal indefinite line of grey shading, orbicular, a white dot darkly encircled.
Secondary uniform brownish pink.
Expanse : 53 mm.
Hub. The type is in my collection from Ekeikei, where the species occurs in
January and February. I have specimens from other localities taken in March and
April high in the Kebea Range (6000 ft), also from lower places in July.
The reniform occasionally is almost white instead of dark grey.
148. Cosmophila mafalui spec. nov.
i. Head rufous ochreons ; thorax warm reddish brown ; abdomen pinkish
brown with pale laterals. Primary with the basal half of the wing purple-brown,
roughly margined by the medial line, beyond wliich the colour is paler cinnamon-
brown, rather darker in the terminal area ; au obscure grey basal line with an
outward curve ; medial line from the upper margin of the cell very irregular and
finely serrated ; postmedial line from the costa to vein 3, very irregular ; a sub-
terminal oblique indefinite stripe of darker shading ; a white point in the cell ;
reniform obscurely ontlined, followed by a blackish point beyond its upper
extremity, and by two blackish points beyond its lower extremity. Secondary
uniform brownish red.
? . Like the male, but with but little purple suffusion.
Expanse : 00 mm.
llab. The type is in my collection from Mafalu, where it was taken in August.
1 have specimens from other localities taken in March and April.
( 238 )
Sinariola srf^n. nov.
Palpus, second segment well scaled, long, reaching above vertex ; third joint
very long, nearly bare, clnb-shaped ; antennae finely shortly ciliatc ; legs, middle
with one, hind with two pairs of spnrs. Neuration : Primary with veins 3, 4 and 5
from close to the lower angle, G from the n]iper angle, 7, 8, 0 and 10 stalked,
11 long from below the middle of the cell. Secondary, veins 3 and 4 from the
angle, 5 directly above the angle, 6 and 7 from the npper angle, 8 bent down to
touch the cell before its middle,
Costa of primaries dejiressed about the middle and again at tlie apex, below
which it is excavated, after which the termeu is strongly i)roduced forward, receding
gently from vein 4 to the toruns.
Type : Sinariola owgarra B-B.
140. Sinariola owgarra.
?. Head and thorax very dark nmlier-brown, abdomen dark brownish grey.
Primary very dark uniform umber-brown, with a trace of a dark waved medial line
and of a pale postmedial waved line incurved at vein 3. Secondary pale greyish,
becoming dark towards the toruns and less dark at the apex ; tornus and terminal
area irrorated with lavender-grey.
Expanse : 6U mm.
Hah. Owgarra, British New Guinea.
Type in my collection.
150. Sinariola rufa spec. nov.
S. Head and thorax jiale rufous, abdomen greyish, antennae with white scales
at their base. Primary dull uniform rufous, with an e^cnrved waved antemedial
line ; postmedial line obliipie with a short angle directly below the costa ; this
is followed by a similar fine almost parallel dotted line ; terminal area darker ;
reniform palely outlined. Secondary pale reddish greyish towards the base.
Ex]ianse : 50 mm.
llah. Owgarra.
Type in my collection.
151. Pseudogonitis pratti spec. nov.
?. Head, thorax and abdomen dull chocolate-brown. Primary dnll chocolate-
brown, with a trace of a whitish basal line ; a whitish ])i)int at the orbicular, and
two such jioints at the reniform ; postmedial line whitish, strongly serrate; a trace
of a dark snbterniinal line with three bhu'k points on veins 4, 5 and (! ; ajiex whitish
grey. Secondary dark grey, slightly paler towards the base.
Expanse : 42 mm.
Hub. The type is in my collection from the Kebea Range, where it was taken
in .July.
Paragonitis gen. nov.
Differs from Psntdogonitis in the jiiilpi being j)orrcct, the third segment being
very long and exjiauded at the tip, whilst veins 3, 4 and 5 of the secondary are
from the lower angle of the cell.
Type : Parayonitis strigocrenulata B-B.
( 239 )
152. Paragonitis strigocrenulata spec. nov.
(?. Head and collar bright chestnut-red, thorax reddish brown, abdomen
brownish grey. Primary uniform reddish brown, with the autemedial dark line
fractured at the lower margin of the cell, the longer and lower part being almost
in the median area ; postmedial line fine dark crenulate, edged externally with
ochreons grey, followed immediately by a straight oblique dark line, edged externally
with pale ochreons grey; an ochreons point at the orbicular; reniforra ochreons
grey, with two black internal dots and the ochreons grey colour suffused outwards
between veins 3 and 6 to the oblique line ; a slight grey costal scaling in front of
apex, and a trace of a subterminal irregular stripe of darker sliading just beyond
the ol)li(pie line. Secondary reddish brown, becoming cream-coloured baseward.s.
Expanse : 45 mm.
Ilab. The type is in my collection from the Kebea Eange, where it was taken
in July.
Gonopteronia gen. nov.
Palpi porrect, thickly scaled, second segment reaching above vertex, end
segment minute, proboscis well developed ; legs, tibiae clothed thickly with long
hair, midtibiae witli one pair of spines, hindtibiae with two j)air. Wings : Primary,
underside of cell clothed with long hair ; costa excavated at the base, then produced
forward into a shoulder, after which a slight excavation occurs again, with another
shoulder with a slight depression to apex, which is produced acutely ; termen crenu-
late, excised immediately below apex, produced to a point about vein 4, and excised
below slightly to toruus. Secondary long, with costa straight, receding rapidly from
the apex to the tornus, before which it is slightly excised. Neuration : Primary with
veins 7, 8 and 9 stalked, 9 given off from 10 to form the areole. Secondary with 5
from near lower angle, 3 and 4 and 6 and 7 from their respective angles. Genital
organs prominently exposed.
Type : Gonopteronia albopunctata B-B.
153. Gonopteronia albopunctata spec. nov.
(?. Head and collar che.st nut-red, the latter tipped with ochreons; thorax
rather darker red ; patagiae slightly tipped with ochreons ; abdomen pinkish brown.
Primary pale reddish brown, slightly ochreons at base, and with an ochreons patch
below the angle of vein 2 ; medial line dark, fine, outwardly oblique, irregular, and
strongly dentate ; postmedial line from near the lower angle of the cell to the inner
margin waved, bordering externally the ochreons jiatch ; a fine dark line from the
second costal shoulder to vein 3 dentate, preceded by some red patches between
veins 6 and 3, followed by a trace of a dark subterminal shading ; termen slightly
dashed (horizontally) with reddish ; the orbicular is a prominent small white spot ;
the reniform red, with a blackish dot beyond its upper margin and two such dots
beyond its lower margin. Secondary uniform brownish pink.
Expanse : .57 mm.
Hah. The type is in my collection from Ekeikoi, where it was taken in May.
We have specimens from other localities in July and also in March and April, and
in the Triug Museum are otliers from the Arua River.
154. Gonopteronia fulminans spec. nov.
(?. Head and collar crimson-brown; thorax deep crimson-brown; abdomen
slate-brown, pinkish brown laterally ; third segment of palpi moderately long,
16
( 240 )
Primarv dark red-browu, slightly crimson at the base, with a trace of a dark basal
line of shading from in front of the first costal shonlder, which (shoulder) is very
prominent ; medial line from this shoulder outwardly obli(ine, with a short
semicircular curve in the middle ; postmedial line inwardly oblique from the
reniform, slightly waved, bordering a small orange-red patch below vein 2 ; beyond
this is a very irregular jagged fine dark lino from the costa to vein 2 ; an obscure
subterminal band of dark shading, a white point in the cell. Secondary dark
brownish grey.
? . Like the male.
Expanse : (? and ? 60 — 02 mm.
Bab. The type is in my collection from the Kebea Range, where it was taken in
March and April. In the Tring Museum are other specimens from the Aroa Kiver.
155. Capotena albicosta spec. nov.
c?. Head and thorax dark lilac-grey ; abdomen dark grey, with anal segment
pinkish white. Primary pinkish lilac-grey, with the costa narrowly but prominently
white ; two oblique pale parallel lines from one-third and two-thirds of the costa, the
inner line straight, the outer one curved just above the tornus to the inner margin ;
a subajiical curved series of four black dots, followed below by two more such dots.
Secondary dark grey, paler below the cell.
? . Like the male.
Expanse : S and ? , 46 mm,
Bab. The type is in my collection from Ekeikei, where it was taken in March ;
other specimens from the Kebea Range, July.
This species is a close ally to C. elaina Swinhoe, which we have from the
same localities ; but the dull lilac-colour and the marked whitish costa will easily
difterentiate them.
156. Capotena subflava spec nov.
$ . Head and collar dark grey-brown, eyelashes rufous brown ; prothorax dark
grey-brown, with a short pale yellow central line ; patagia pale grey, with a dark
grey patch on its inner fore edge ; metathorax rufous brown ; abdomen velvety
black, with fine grey segmental divisions ; anal segment white ; thorax and abdomen
entirely whitish grey below. Primary pale lavender-grey, with the lower part of
the base suffused with greenish brown, followed by four oblique straight bands
of greenish olive-brown, the fourth band before the apex being short ; all these
bands are indefinite, the brown being suffused over the ground colour, but the first
oblique band is definite on its outer edge, and the third and fourth on their inner
edges ; the whole wing is finely irrorated, and is also finely streaked with short
oiive-browu dashes ; fringes whitish for the inner half, tipped with olive-brown.
Secondary blackish, pale grey and subhyaline below the cell.
Underside : Primary black, with the costa, apical area and termen bright
orange-yellow. Secondary blackish, subhyaline between veins 1 and 'I ; a hyaline
spot beyond the cell ; costa and apex bright orange-yellow.
? . Like the male.
Expanse : $ and ? , 55 — 66 mm.
Hub. The type is in my collection from the Kebea Range, where it was taken
in March and April ; other specimens from Pinawa, August,
(241 )
157. Capotena albotessellata spec, no v.
c?. Head dark reddish brown ; oollur orunge-brown ; thorax red-brown, with
metatliorax edged fine]}' with white; alidomen grey. Primary red-browu, extreme
base white ; a wliite streak along the fold to the tornns, where there is a white
patch ; two white outwardly oblique nearly parallel lines, the inner one straight
from the costa near the base, the outer one from a third from the apex, angled at
vein 4, where it curves back into the white fold streak ; termen finely white ; a
dark dot at the reniform. Fringe white at apex, brown below. Secondary dark
grey, pale between veins la and 2.
? . Like the male, but darker.
Expanse : S o2, ? 4.5 mm.
I/ab. The type is in my collection from Babooni, where it was taken in
September ; I have other specimens from Dinawa, where the species occurred
from May to July.
Sphingiforma gen. nov.
Differs from Capotena in the third segment of the palpi being long.
Neuration : primary, veins 6 and 7 from the areole ; a bar from 7 to 10 to form
the areole ; 8, 9 and 10 stalked from beyond the areole. Secondary with two
internal veins; cell very long, deeply excavated, especially in the male ; veins 3, 4
and 5 in the male not stalked as usual, but given off together from the common
origin, as in anastomosis ; in tlie female they are stalked as usual ; 6 and 7 joined
by a short bar ; wing excavated in male beyond vein 3 to near the toruus.
Type : Sphingiforma pratti B.-B.
158. Sphingiforma pratti spec. nov.
S- Head, thorax, and abdomen dark brownish grey, the latter with anal
segment pale lavender-grey ; the genital organs have long lateral whitish
retractible tufts. Primary lilac-grey, dull and dark for the basal half, paler
for the outer half ; the whole of the wing is finely irrorated with blackish ; a
trace of a dark medial line; a dark point in the cell; beyond this a dark, nearly
quadrangular costal spot ; a subterminal row of dark points. Secondary dark grey.
? . Similar in colour to the male, but the primary has an irregular oblique dark
basal line ; a broad oblique dark median band, angled outwards about vein \a, and
reaching the termen above the tornns — this stripe is very broad and dark in the
radial area ; a broad obli(iue baud from the costal spot bent round about vein 4 to
meet the median band; a short waved dark oblique stripe from in front of the apex.
Expanse : S 58, ? 52 mm.
Hah. The type is in my collection from Dinawa, where it was taken in
September; other specimens from Ekeikei in January.
Parelydna gen. nov.
Differs from Pseudeli/rliia in the third segment of the palpi being moderately
long, in the abdomen of the male being short ; the tibiae have no tufts of long hair ;
the midtibiae have one jiair and the hindtibiae two pairs of spurs ; veins 2 and 5
of the secondary from close to the lower angle ; veins 3 and 4 stalked. The primary
is broad, expanding considerably near the termen. The secondary is ample, has the
apex sharply angled, is excised sliglitly below it, and has a waved termen.
Type : Parelt/dna mirabilis B,-B,
( 242 )
150. Parelydna mirabilis spec. nov.
rj. Head and tliora.x greenish yellow ; aliduinen smalt blue, witli white anal
segment. Primary a peculiar yellowish green, with a small oval pateh of raised
scales in the cell ; a broad white jwstmedinl stripe edged with a small blue jiatch
at the toruus, above which the stripe is finely edged with green, from whence to
the termen the colour is Indian red ; fringes whitish, blue at the tornus. Secondary
uniform smalt blue. Underside, both wings blue ; secondary with a white ape.x.
?. Exactly like the male.
Exjjanse : c? 32, ? 38 mm.
I/a/j. The type is in my collection from Ekeikei, where it occurred from January
to April. The species occurred at Mafalu (over 6000 ft.) in August, and is a
common species, as we have a long series from the various localities. Other
specimens in the Triug Museum from the Aroa River.
The species is a close ally of Pseudebjdna cijanea Snell., but is adiflPerent colour,
and the termen is quite diiferent. P. cyanea should be referred to this genns, as it
is not cogeneric with Pxeaclebidiui rufojiara.
160. Carea parallelaria spec. nov.
cJ. Head and thorax crimson-red, abdomen brownish. Primary crimson-red,
slightly lustrous, with darker red oblique median and postmedian parallel lines ;
the latter from the inuer margin to a dull redder cloud right across the wing.
Secondary pale reddish, with greyish base and inner margin.
? . Much greyer and paler than the male, with the medial line curved
obliquely and not parallel with the postmedial line, and with two dark jioints
in the cell, of which there is but the least trace in the S.
Expanse : S 37, ? 38 mm.
llah. The type is in my collection from tlie Kebea I'ange ; it was taken in
March and April. Other specimens from Ekeikei were captured in the same
months.
101. Carea unipunctata spec. nov.
$. Head and thorax reddish chestnut; abdomen yellowish, witli reddish anal
segment. Primary reddish chestnut, with the dark medial and jiostmedial lines
oblique and somewhat waved, with a dark spot at the end of the cell ; subterminal
area paler ; termen dark purplisli, apex acute, slightly falcate. iSi'condary orange-
yellow, paler on the inner margin.
¥ . Just like the male, but darker.
Expanse : i and ?, 40 mm.
Ilab. The type is in my collection from tlic Kebea Kauge, wliere the sjiccies
occurs in March and April, and in July.
162. Carea flava spec. nov.
?. Head and thorax pale bnff-colour, abdomen pinkish. Primary, basal half
of wing pinkish ; terminating obliquely in this area is a short oblique blackish
costal dash near the base ; a broad greenish dash below the cell and two dark dots
in the cell ; between this pink area and the postmedial line is a triangular patch of
yellow, the costa being the base of the triangle and the apex in front of the tornus ;
iu this beyond the cell is a short double greenish black dash with an iudetinita
( 243 )
greenish slifwliiig above and a little in advance of it ; postmedial line donble, with
a white centre, nearly erect, but slightly hollowed, and extending basewards along
the costa for a sliort distance ; from this line nearly to the termen the ground colour
is pale lilac-pink ; termen yellowish. Secondary uniformly yellow, slightly paler
at the base ; fringes reddish.
E.xpanse : 48 mm.
Ilab. The type is in my collection from the Kebea Range, where it was taken
in Jaly.
This species is near C. rliodophila ; but I believe that it is not a true Carea,
but should be jjlaced in the genns Careades, a new genus, described later on in this
paper.
163. Carea pratti spec. nov.
S. Head and collar velvety dark greenish brown, thorax and abdomen pale
ochreous. Primary pale oehreous with a broad dark oblique straight greenish
brown band from before the middle of the inner margin into the apex, beyond
which the ground is pinker ; the inner costal angle of the oblique band is filled
in with pale greenish brown ; termen greenish brown, tapering finely to vein 2.
Secondary pale yellowish orange, deeper towards the termen.
? . Exactly like the male in all particulars.
Expanse : 46 mm.
Hah. The type is in my collection from the Kebea Range, where the species
occurs in July.
Careades gen. nov.
Differs from Carea in that both the mid- and hindlegs are densely clothed with
long hair, the mid pair being clothed down to the end of the first segment of the
tarsus, the hind jiair down to nearly the end of the tarsus ; in the female the hair
is but little longer than usual. The costa (c?) of the primary is strongly waved ;
the apex is shortly . falcate ; termen excavated below the apex, then produced
outwards in a bold Curve. Secondary strongly excavated in front of the fold, which
(fold) is jn'oduced outwards into a slight lobe ; below, lying in the fold, is a long tuft
of silky hair.
Neuration with veins 6 and 7 from the areole, 8 and 9 forked, the latter given
ofi'from lU to form the areole. Secondary with veins 3 and 4 strongly curved from
the angle, lying touching each other for a short distance. Both cells very short.
Type : Careades sanguinea B-B.
164. Careades sanguinea spec. nov.
<?. Head and thorax deep crimson, abdomen dark grey. Primary dull magenta
brownish pink, the area between the antemedial and the obli(iue lines being quite
pale ; antemedial line fine, irregular, finely toothed inwardly on the upper margin of
the cell, with an ontward curve below this; an oblique broad dark stripe, margined
outwardly with blackish, beginning before the middle of the costa and ending
almost in the tornus ; a trace of an erect line of dark dentate shading from in
front of the a])ex into tiie tornus ; terminal area greenish brown : a fine black point
in the cell. Secondary basal half dark greyish brown, outer half red ; beneath in
the iunermarginal fold is a long tuft of dark red silky hairs.
? . Like the male in general colour and pattern, except that the oblicpie line
( 244 )
is reversed, being sliglitly oliliqne in the reverse direction, whilst the secondaries
are uniformly brown.
Expanse : c? 4S mm., ? 43 ram.
Hab. The type is in my collection from Ekeikei, where the species flies in
March and April.
105. Careades ekeikei spec. nov.
3 . Head, collar and i)rothoras dark bronzy greenish grey, rest of thorax pale
ashy grey ; abdomen ashy grey, darker for the anal portion, with a buff anal tnft.
Primary, a pocnliar shade of pale olive grey, with a fine lunulate antemedial line,
and a very broad dark bronzy greenish obliqne line, edged outwardly by a double
line, the first blackish, the second very pale ashy grey ; beyond this the ground
colour is rather darker and greener, with a trace of an erect line of dark shading
in front of the lermen, which is tinged with greenish grey ; the oblique line is
shorter and less oblique than in the previous species. Secondarj' uniform very dark
grey ; beneath, lying in the fold, is a long tuft of long jiinkish hairs.
?. Like the male, except that the oblique line is nearly erect, and in front of
it is a large area of dark bronzy greenish colour.
Expanse : c? and ? , 44 mm.
Hab. The type is in my collection from Ekeikei, where the species flies in
March and April.
100. Careades falcataria spec nov.
c?. Head and thorax deep crimson, abdomen dark grey. Primary very dark
blackish crimson ; the antemedial line double, distinct on the costa, obscure below ;
the double oblique line, with a paler centre, is almost angled near the costa, below
which it is slightly waved into the tovnus ; beyond this the ground colour is
decidedly paler, except in the costal angle made by the oblique line ; the apex
is strongly hooked, and the termeu much produced near veins 3 and 4. Secondary
palish bright orange-red ; beneath, lying in the fold, a long tuft of carmine hairs.
Underside rufous in both the wings ; in the secondaries there is a long tuft of
red silky hair.
Expanse : 48 mm.
Hab. The type is in my collection from Ekeikei, where the insect occurs in
January and February.
107. Careades fulva spec. nov.
c?. Head and tliordx reddish fawn-colour, the latter with a pale central stripe ;
abdomen dark greyisli. Primary reddisli fawii-colnur with two dark dots in the
cell ; no antemedial line ; the postmedial lino dark, palely edged, externally angled
on vein 0 to the costa ; beyond this line there is a broadish stripe of silvery lilac
shading. Secondary dull pinkish red with the fold dark greyish, in which beneath
lies a long tuft of ochreous hairs.
?. Similar in colour to the male, but pinker, with a fine slightly darker
antemedial line ; the posterior double line is evenly curved to below vein 2, where
it is angled inwards, and proceeds somewhat obliquely to the inner margin ; the
lilac shading is whiter and is freckled rather than simple shading. The secondaries
are paler than in the male.
( 245)
Expanse : (? 50 mm. ; ? 45 ram.
Bab. The type is in my collection from the Kebea Range, where the insect
occurs in Marcli and April, and also in -January and February ; I have other
specimens I'rom Ekeikei captured in the same month.
168. Careades obliqua spec. nov.
i. Head and thorax pale reddisli, abdomen dark grey. Primary pale yellowish
red, with the antemedial line irregular and curved outwardly ; the medial line
oblique from the costa before the middle straight into the tornus, beyond which the
ground colour is paler ; a snbterminal erect interrupted line of dark grey shading
from the costa into the tornus ; a dark point in the cell. Secondary orange
reddish with the marginal fold dark grey, in which beneath lies a long tuft of bright
red hairs.
? . Like the male but darker, with the second line, and the postmedial one not
oblique but gently waved.
Expanse : c? 41 mm., ? 38 mm.
Hub. The type is in my collection from the Kebea Range, where the species
flies from March to June.
Kif). Careades subrubra spec. nov.
6. Head and thorax dark red, abdomen lilac-grey. Primary lilac-grey, with
the base dark to beyond the antemedial line, which is slightly angled at the
lower margin of the cell; postmedial line oblique, beat at the upper margin of
the cell ; two dark points in the cell ; au irregular band of pale snbterminal
shading ; termen dark bronzy brown. Secondary dull brick-red, with a dark grey
fold in which beneath lies a long tuft of deep crimson hairs.
? . Similar to the male but redder, and both the lines are double, the post-
medial one being less oblique with its inner costal angles filled in with orange
reddish.
Expanse : c? and ? , 40 mm.
Hab. The type is in my collection, from Mount Kebea, where it was taken in
July ; I have other specimens from Dinawa captured in September.
170. Careades rubricosa spec. nov.
¥. Head and collar deep crimson, abdomen dark grey. Primary dark crimson,
with the antemedial line slightly curved and waved ; the postmedial line double,
slightly receding as in the previous species, but also somewhat hollowed internally ;
this line has a broad internal patch of darker dull red, the rest of the ground
colour being slightly shining ; termen dark red. Secondary uniform dark grey
with red fringes.
Expanse : 42 — 44 mm.
Hab. The type is in my collection from Ekeikei, where the species occurs in
March and April.
Paracarea gen. miv.
Differs from Careades B-B. in that the third segment of the palpi is very
long, nearly us long as the first and second together, and that it gradually
thickens to the tip, forming a slight club. The neuration of the primary is
similar to Carmdc.^, but in the secondary veins :5 and 4 are stalked.
Type : Paracarea rttbiyinea B-B.
( 246 )
171. Paracarea rubiginea spec. uov.
?. Head and thorax bright red, abdomen reddish ochreons. Primary uniform
bright red, with a taint obliijne antoinedial line ; postmedial line faint, curved and
sliglitly waved ; a trace of a subtenuinal greyish shading ; two dark dots iu the
cell. Secondary reddish orange, with the base creamy ochreous.
Expanse : 40 mm.
Hab. The type is in my collection from Dinawa, where the species occurs in
Angnst. I have specimens also from the Kebea Range, in July.
Subfamily QUADRIFINAE.
17-'. Catocala dinawa sjjec. nov.
? . Head, thorax and abdomen grey. Primary grey, finely irrorated witli blackish ;
antemedial line obscured below the cell ; postmedial line strongly angled beyond
the cell, very deeply serrate, obscure below vein 5 ; a whitish irroration is on the
costa near both these lines. Secondary yellow, with a curved broad blackish
termen from the costa to the tornus, tapering rapidly below vein 4 ; a small white
terminal patch below the apex.
Expanse : 07 mm.
Hab. The type is in my cullection from Dinawa, where it was taken iu
August.
Erygansa gen. nov.
S. Palpi, second segment reaching vertex, heavily scaled ; third segment long,
smooth ; antennae of (J with very fine short bristles ; thorax and prothorax with a
large spreading tuft, metathorax smoothly scaled ; abdomen with slight tufts on
proximal segments, anal tuft moderately large ; legs with all the tibiae heavily
clothed with hair ; mid- and hindtibia spined ; hindtarsi fringed with long hair to
the end segment. Neuration : Primary, veins 3, 4 and .5 from close to the angle.
Secondary, veins 3 and 4 stall<ed, 5 from the angle ; ternicn of both wings
crenulate.
Type : Enjgansa kebea B-B.
173. Erygansa kebea spec. uov.
3- Head and thorax brown, mottled with dark brown ; al)d<)mon dark grey.
Primary umber-brown, with the veins, except veins 1 and 12, broadly lined with
dark grey ; a short double basal pale line on the costa ; antemedial line double
and dark to abont vein 1 ; medial dark line double, interrnpted below the cell;
postmedial line dentate, produced outwards beyond the cell, beyond which are two
faint dentate lines ; a subterminal row of dark spots palely edged on the exterior ;
termen finely pale ; all the lines are somewhat difficult to trace. Secondary dark
grey, with the basal half jialer; fringes unevenly dark and pale alternately.
Expanse : 48 mm.
Hab. The type is in my collccti(jn, from the Kebea Range, where the species
occurs in March and April.
174. Erygia avola spec. nov.
?. Head and thorax brown, slightly rufous, barred with black; abdomen
brown, with tufts slightly reddisii. Primary brown, tinged with rufous ; base
dark margined by a double dark line ; medial line faint, slightly angled at each
(247 )
margin of the cell, lieyond which the area is ])ale ochveons brown, extending deeply
outwards along vein 3 up to 4, whence it recedes gently to the costa ; this area is
margined indefinitely with black, very broadly so below vein 3 ; subterminal line
black, irregular, somewhat spotted, angled outwards about vein 4, from whence it
takes a rough curve to the a])ex and the tornus ; a subterminal row of dark spots
palely edged externally. Secondary brownish, ochrcous along the costa, with a dark
line crossing the cell, followed externally by two similar lines, the outermost one
palely edged for the tornal half; termen darkly spotted ; fringes strongly crenulate.
Underside : both wings somewhat ochreous, the secondary strongly irrorated with
four dark lines across the wing, the third serrate, the fourth very broad.
Exjianse : 53 mm.
Ilab. The type is in my collection, from AvoJa, where it was taken in August.
175. Erygia kebeensis spec. nov.
? . Differs from E. acola m that it has no pale patch ; the postniedial line
is double, and only exists below the cell, beyond which is a patch of a greenish
olive colour, and above which on the costa is a large dark brown j)atch ; subterminal
line irregular, blackish, taking an iuward curve from tlie apex to vein 5, from
where it proceeds in a dentate line close to the termen into the tornns ; the
reniform is large and ochreous. On the underside the secondary has but two
transverse waved lines, which are more definite than in the previous species.
Expanse : 58 mm.
Hah. The type is in my collection from the Kebea Range, where it was taken
in Julj'.
170. Sypna angulilinea spec. nov.
6- Head and thorax dark brown, collar with a pale creamy basal line, patagiae
palely edged. Primary dark iimber-browu, witli a i)ale basal line to vein 1 ; medial
pale line, deejily angled outwards below the cell ; postmedial line confluent with
the medial at its angle, thus enclosing two large roughly triangular patches of the
dark ground, whose bases are respectively the costa and the inner margin ; almost
adjoining this line is a broadish pale brown line projected outwards along vein 4
to 5, whence it rises erectly to the costa ; from this line to the termen the ground-
colour is paler. Secondary greyish brown, with two dark transverse bands ; a pale
terminal spot below the apex and several smaller ones near the tornus.
Expanse : 00 mm.
IJab. The type is in my collection from Dinawa, where the species flies in
August. Other sjjecimens are in the Tring Museum from the Aroa River.
177. Sypna bella spec. nov.
Head and thorax warm chocolate-brown, the latter barred slightly with greyish,
having two large lateral tufts and a small central metathoracic one ; abdomen
brownish, with grey segmental divisions. Primary warm rufous brown, with a
basal grey curved line ; a double antcmedial bluish-grey broadly scalloped line with
a somewhat paler infilling, an irregular bluish-grey postmedial line followed by one
or two obscured lines, a large patch of olive-greenish occupying all the rest of the
wing except above vein 6, wluch is rufous brown ; a subterminal more or less
obscure serrate line ; termen with internervular bluish dashes. Secondary duller
brown, darker beyond the postmedial serrate indistinct line, with a darker tornal
( 248)
cloml. Apex witU three creamy white sjiots through the fringe, below which are
internervular pale l)luisli dashes.
Expanse : 7U mm.
Type in my collection from Mafiiln. August.
The greenish patch appears to he somewhat evanescent.
178. Ercheia certa spec. uov.
?. Head and thorax sepia brown, with a paler central stripe through each ;
abdomen dark grej-, with sepia brown dorsal tnft. Primary sepia brown, with the
main veins dark grey, thus the basal area is mostly dark grey ; an obscure trace
of a postmedial dotted line, and traces of a double serrate 1 line beyond it : sub-
terminal line pale evanescent in the apical paler half, but visible below vein 5 ;
a darkish apical costal patch; a ]iale triangular small patch about vein 4, showing
plainly in the dark terminal area ; inner margin rather jialer than the ground
colour. Secondary dark grey, with a good-sized white central spot ; fringes dark,
white between veins 1 and 2 and between veins u and 0.
Underside : Primary dark, with a broad, postmedial j)ale band ; termen pale
chestnut brown. Secondary, basal half pale ochreous with a dark cell-sjjot and a
dark waved median band ; outer half dark grey.
Expanse : 56 mm.
The type is in my collection from ])inawa, where it occurred from May
to July.
17'J. Ercheia ekeikei spec. nov.
S. Head and thorax dark slate-grey, abdomen grey. Primary pinkish ochreous
above vein 1 ; outer half of wing somewhat smoky, below vein 1 dark lilac-grey ;
a basal line bordering a slightly darker area ; a strongly excnrved dark postmedial
line to vein 4, outside which is a darker smoky triangular costal patch ; reuiform
large, whitish grey ; subterminal line ])ale to vein 5, darker below ; a small dark
spot at the apex, and a dark iudeliuite curved band from half along vein 1 up to
vein 5 on the termen, in which below the origin of vein 2 is a pale creamy yellow
dash ; the iunermarginal grey area is edged above by a whitish line. Secondary
dark grey, with three i)ale sjjots.
Expanse : 54 mm.
Hab. The type is in my collection from Ekeikei, where the species flies in
January and February. I have specimens also from the Kebea Range taken in
July.
The species is somewhat near K umirosa Btl., but can be readily recognised
therefrom.
180. Ercheia styx si)ec. nov.
c?. Head brown, thorax dark brown with a paler central stripe, abdomen
dark grey. Primary purplish brown, with a broad central stripe along the cell
and nearly into the termen of a pale manve-grey, all the lines except the postmedial
being more or less interrui)te(l by it; antemedial blackish line serrated on the costa,
medial line obsolescent, jiostmedial line strongly jiroduced outwardly beyond the
cell, below vein 5 broad and dark ; subterminal line pale, similar in shape, receding
to the inner margin, where it is almost white ; below the cell is a paler dash finely
margined with dark brown ; beyond this is a short pale creamy dash, from which
( 249 )
a dark stripe extends to the termen ; termen with a pale spot about vein 4.
Secondary dark grey, with a small central creamy spot. Fringes dark, pale between
veins 1 and 2 and between 5 and 6.
Expanse : 46 mm.
Hab. The type is in my collection from Ekeikei, where the species flies in
March and April.
This species will come next to E. churon Btl.
ISl. Ercheia kebea spec. nov.
Head dark brown ; collar and thorax pale chestnut-brown, darkly irrorated, a
dark central prothoracic spot; abdomen dark grey, with a dark chestnut tuft on
the proximal segments. Primary brown ; basal area dark purplish to the lower
margin of the cell, below wliich it is jjale ochreous at the base, shading into dark
irrorated chestnut-red brown, the whole edged by the wavy dark antemedial line,
at the end of which the inner margin is produced into a lobe deeply excavated
on the exterior; beyond this line the area is paler, with a slight lavender-grey
suffusion, with a dark spot below the angle of vein 2 ; postmedial line faint,
produced well forward to vein 4, where it recedes to the inner margin ; beyond
this line the ground colour is darker, sepia brown, very dark on the costa up to
the apex ; two creamy points at the end of the reniform; termen finely dotted
with blackish. Secondary dark grey, with an interrupted j^ale medial baud from
the cell to the inner margin, beyond which the colour is very dark ; fringes blackish,
creamy from vein 1 to 2 and from 5 to 0.
Expanse : 58 mm.
Hab. The type is in my collection from Mount Kebea, where the insect occurs
in March and April. We have specimens from other lower localities in January
and February, and from the high localities in May and June. In the Tring
Museum from the Aroa River are others.
1S2. Hypocala kebeae spec nov.
? . Head and thorax dull rnfous brown ; abdomen alternately banded black
and yellow. Primary uniform dull rufous brown, with a darker shading beyond the
cell ; sparingly irrorated with black points edged internally with grey ; a small
bluish grey spot at the tornus with a black pupil, above which is a whitish jjoint ;
termen palely dotted. Secondary black, with a double yellow spot at the tornus ;
a yellow streak in the cell expanding into a spot beyond it ; a yellow stripe on the
fold, separated from the yellow cell by the black ground colour.
Underside black, reduced, especially on the secondary, where it is but a small
spot in the cell and a bar at- the tornus.
Expanse : 49 mm.
Hab. The type is in my collection from the Kebea Range, where the species
flies in March and April.
183. Hypocala pratti spec. nov.
$. Head and central stripe of thorax pale cinnamon-brown, thorax purplish
brown, abdomen yellow with narrow dark grey dorsum. Primary reddish fown-
brown, redder below the cell ; a short double basal line, an oblique antemedial
( 250 )
(lonlile line strong!}' cxcnrved in tlie fold, postmcdial line double, stronglj- excui'ved
beyond the cell and sliglitly crennlatc ; both stigmata darkly tilled in and encircled ;
snbterminal line whitish snbcrenulate, a])ical half of terineu whitish grey. Secondary
pale yellow with very broad black termen tapering down at the tornns, a white
terminal patch below vein 7.
Expanse : 40 mm.
Ilalj. The type is in my collection from Dinawa, where the species occurred in
May and Jnne ; it is also in the Tring Mnseum from the Aroa River in March.
Pseudozalissa gen. nov.
Palj)i n{)turued, end segment erect, reaching above the vertex, third segment
scaled. Nenration : Primary, veins 3 and 4 near the angle, 5 from below the
middle of the discocelliilars, 6 from directly below the npper angle, 7 from just
above the angle, bent down at a third and sliglitly depressed to the termen ; 8
and 9 on a short stalk given off from 10 to form the areole. Secondary, with
3 and 4 from the angle, 5 from below the middle of the discocellulars, C and 7
from the upper angle.
Type : I'semlozulisHa bella B-B.
184. Pseudozalissa bella spec. nov.
(J ? . Head pinkish cinnamon, collar dark red fawn-colonr, thorax jiale
cinnamon-brown with palest greenish-white shining patagiae ; abdomen yellow
with banded dark and pale terminal segments. Primary pale lilac grey with
waved dark antemedial line, outside which is a dark spot ; postmedial line
irregular, arising in a small snow-white triangular costal patch ; inside this line is
a patch of rich dark purplish red colour extending down towards the inner margin,
au apical patch of very pale greenish grey, snbterminal line serrate, white ; termen
below vein fi dark purplish, between veins 4 and 5 a purplish spear-shaped dash.
Secondary pale yellow with a very broad dark termen tapering down to the anal
angle, a subapical white terminal narrow patch.
Expanse : 50 mm.
Hab. The species occurred at Dinawa and Ekeikei in March and April, and
again in August.
The type is in my collection ; it is also in the Tring Museum, from the Aroa
River.
184. Nyctipao dentifascia obscura subspec. nov.
Similar to N. dentifascia Wlk., but entirely witliout any white stripe or white
spots on either wing.
Hab. Ekeikei (1500 ft.), May.
AVe have the ordinary form of N. detitifascia Wlk. from the mountains, hut
all the specimens from the low distri(tts are of this dark form.
l8.">. Polydesma pratti spec. nov.
?. Frons and pal|)i reddish brown; collar brown tipped with |)ale green;
thorax ])ale green ; patagiae with a brown stripe across near the tips ; abdomen
pale greyish brown ; legs dark brown ringed with cream, a pale green patch on the
fore- and midtibiae. Primary pale jiea-greeu, with two brown basal points ; an
( 251 )
irregular tapering subbasal brown patch, widest on tbe costa, edged with white
externallj-, followed by an irregular white antemedial line ; a triangular brown patch
above the end of the cell, edged with wliite internally, from which is an irregular
white line to the inner margin, which is edged internally by a brown line, witii brown
spots near the margin and short brown dashes by the cell ; a postmedial irregular
white line rising from a dark costal dot; a large brown patch at the apex, and
a smaller one at the tornns ; termen finely dark. Fringes pale greenish, intersected
with pale brown, slightly crenulate. Secondary brown, paler near the base, with an
interrupted pale terminal line.
Undersurface dark brown ; primary with a pale costal patch in front of the
apex and a postmedial darker line ; secondary, a dark spot in the cell and a medial
and postmedial dark line.
Expanse : 46 mm.
I [ah. The tyjie is in my collection from the Kebea Range ; the species flies in
March and April.
18(i. Polydesma brunneistriga spec. nov.
Head, tliorax, and abdomen greyish. Both wings whitish grey. Primary with
two small rusty-browu basal spots, an interrupted double antemedial rusty-brown
line ; a rusty-brown broad dash into the reniform ; an irregular costal short streak
before the extremely broad and irregular postmedial area of rusty brown ; termen
with fine dark points. Secondary with no marks except the very broad postmedial
area as in the primary.
Expanse : 55 mm.
Hah. Aroa River, February and March.
Tyi)e in my collection, and other specimens in the Triug Museum.
188. Panilla aroa spec. nov.
S- Head, thorax, and abdomen ochreous grey. Primary ochreous grey, with
an antemedial and medial line enclosing a slightly darker area with centre
paler ; costa darkly clouded ; postmedial pale line irregular, edged externally with
a dark costal j)atch, below which is a second similar patch ; subterminal pale line
serrate , an anteterminal row of dark points. Secondary with the basal two-thirds
ochreous grey ; terminal third pinkish, with a broad dark medial stripe with two
blackish dashes across the cell, followed by a pale greyish area ; postmedial line
very obscure ; an anteterminal row of black points.
?. Like the male, but olivaceous grey with the markings generally somewhat
obscured.
Expanse : c? 24 mm., S 28 mm.
Hab. Aroa River, February and March.
Type in the Tring Musenm.
189. Catephia rufostrigata spec. nov.
(J. Head and thorax dark reddish brown, irrorated with chocolate brown ;
patagiae with a blackish line edged with rusty-brown ; metathoracic tuft greyish ;
abdomen dark brownish with reddish dorsal tufts. Primary dark reddish brown,
with a broad rusty stripe from the base through the coll to the termen, slightly
( ^'^2 )
angled at the loner eud of the cell ; this stripe interrupts all the markings ; basal
line slightly rust}' and obscure ; a trace of an antemedial and postmedial stripe, bnt
it is obscure : subtermiiial rnsty stripe dentate ; below the cell tliere is a velvety
blackish shading, and the wing is more or less mottled with similar coloured spots
or j)atches. Secondary blackish, with a white centra! patch to beyond the ceU.
Expanse : 52 mm.
Hah. The type is in my collection from Ekeikei, where it was taken in January
and February.
190. Catephia alboplagiata spec. nov.
S. Head and thorax dark mottled brown, the latter with a large white tnft on
the metathorax ; abdomen dark brownish grey with a white dorsal ridge on the
proximal segments. Primary dark brown, somewhat mottled with blackish ;
antemedial line double, dentate, with its centre paler ; medial line obscure, post-
medial line double, dentate with a ])aler brown centre ; subterminal line obscure ;
a large creamy white oval spot below veins 10 and 11, below it on the inner
margin a greyish white smaller spot. Secondary blackish, with a central white
patch to beyond the cell.
Exjianse : 53 mm.
Hub. The type is in my collection from Ekeikei, where the insect flies in March
and Ajiril.
191. Catephia mediogriseata spec. nov.
? . Heiid and thorax mottled dark brown ; abdomen blackish grey. Primary
base dark brown spotted with velvety blackish, the spots finely edged with reddish ;
the whole of the median area greyish white, grizzled with dentate lines, with a black
dot in the cell followed by the black reniform ; beyond this area the wing is paler
reddish brown, with the dentate postmedial line on its inner edge obscured ;
the subterminal line irregular and paler; termen jmlely dotted. Secondary blackish
with a white central jiatch to beyond the cell.
Expanse : 4G mm.
Hab. The type is in my collection from Ekeikei, where the insect flies in
January and February.
192. Catephia dinawa spec. nov.
<?. Head and thorax olive-brown, sparingly irrorated with whitish; metathoracic
trift slightly chestnnt-colour ; abdomen pale grey with reddisli brown tufts
Primary, base dark greyish ; rest of wing slate-grey ; botli areas somewhat
shaded in parts with olivaceous. Antemedial dark line bordering the dark basal
area very irregular, projected outwards below the cell, then receding rapidly ;
postmedial line black, strongly dentate in parts; subterminal line angled below the
apex and becoming obsolescent ; reniform darkly outlined ; termen finely dark and
darkly dotted Secondary with basal half white; terminal half blackish grey.
Expanse : 34 mm.
Ilnh. The type is in my collection from Dinawa, where it was taken in August;
we have specimens also from other localities.
The species is near C. perdricipi'nnix Mooi-e, but diti'ers in the shape and
position of both the antemedial and postmedial lines.
( 253 )
193. Catephia pulchristrigata s]>ec. nov.
<?. Head and thorax purplish brown, abdomen purplish grey. Primary dark
velvety purjilisli brown, witli a short basal line into the cell ; autemedial line pale
ochreous brown, with a dark centre ; sharjily waved and oblique from the costa
into the cell, where it is very acutely angled, the angle being formed by a short
inward curve, after which it curves outwardly to the inner margin ; beyond this is
a broad median band of cream-colour, shading into crimson at the two margins and
edged externally by a white evenly curved line ; jjostmedial line broad, dark, finely
edged with ochreous brown, oblique to vein 6, where it is angled suddenly and
acutely inwards ; at vein 5 it assumes an oblique direction to about vein 2, where it
is acutely angled by a sharp inward curve to the inner margin; a steely-blue ocellus
with an olive iris and encircled finely with blackish occupies this curve up to the
tornus ; a large subapical costal jiatch is edged finely with ochreous brown, followed
by three smaller similar oval spots below it ; terminal area broadly pale purplish ;
termen finely ochreous, preceded by a fine double dark waved line. Secondary
blackish grey, with a broad median oblique band of bluish white into the tornus ;
fringes dark, with a white patch below the a])ex.
?. Like the male, but paler; all the lines whitish and very distinct, the pale
median band represented only by a white line ; the internal middle angle of the
postmedial line has two small ocelli somewhat below it, obliquely placed.
Expanse : c? 80, ? 87 mm.
Hab. The type is in my collection from the Kebea Range, where the species
flies in March and April ; other examples from Dinawa in the same months, and in
August from various localities ; and in the Tring Museum from the Aroa River are
other specimens.
194. Catephia splendens spec. nov.
<J. Similar in its main pattern to the previous species, but the area between
the white curved line and the postmedial, instead of being filled in with deep
velvety purplish brown, is slightly cream-coloured at first, shading rapidly into
a beautiful pale olive ; near the white line in this area is a dark purplish C-mark ;
the purplish pale terminal area is broader than in the previous species.
?. Precisely the same differences between the sexes obtain as in 0. pulchri-
strigata B-B., but the area outside the white curved line up to the postmedial line
is whitish bine with the same dark C-mark as in the male.
Expanse : t? 81, ? 83 mm.
llab. The type is in my collcotioa from Babooni, where the species occurs in
July to September, and again in March and April; other examples from Ekeikei,
taken in the same months. It is a rare species.
195. Catephia hampsonia spec. nov.
6. The general pattern of the j)rimary is as in 0. pulchristrigata B-B., except
that the broad creamy median band is reduced to a cream-coloured curved line,
whilst the acute angle of the autemedial line is extended outwards into this line ;
in the secondary the bluish-white band is altered to a large bluish-white patch.
The sexes are both exactly the same in every particular, and the insect is decidedly
smaller than either of its allies.
( 254 )
Expanse : <? and ?, 74 mm.
Hub. Tlio type is in my collcctioQ i'roni the Kebea Range, where the insect
oeenrrod iu Jnly ; other specimens from Babooni taken same mi>nth ; also from
other high hicalities in July anil August.
These three beautiful species are closely allied, but 1 have no doubt they are
distinct ; we have both the sexes represented in each case, and in a long series of
pulch-istrigata there are no intermediate forms.
190. Aedia melas spec, no v.
c?. Head, thorax and abdomen darkly mottled brown, thorax and abdomen
with chestnnt-colonred tufts. Primary dark reddish brown ; antemedial line waved,
beyond which the medial area is very dark ; postmedial line almost crennlate ;
oblif^ne from the costa to about vein 5, where it is sharply angled, and from wliere
it descends to the inner margin, receding basewards very slightly ; beyond this line
the costal area is pale reddish brown ; apex darkish brown ; subtermiual line pale
red-brown, white below vein 6 and interrupted; a whitish spot just above the
tornus ; reniform greyish, from whence there is a jiale greyish band to the termen.
Secondary black, with a white central patch to beyond the cell ; fringes black,
white at the tornus and below the apex.
Expanse : 37 mm.
Hah. The type is in my collection from Ekcikei, where the species flies in
March and April ; other examples in the Tring Museum from the Aroa River.
197. Aedia costiplaga spec. nov.
?. Head and thorax grizzly brown; abdomen brown. Primary lilac-brown;
base finely irrorated with pale grey scales slightly raised ; two subtriangnlar
nmber-brown patches on the costa at the middle and before the apex ; at the
base of the costa and midway between it and the central patch is a dark point ;
below the former on vein 2 is a similar coloured small spot. Secondary uniform
darkish brown.
Expanse : 36 mm.
Hab. The type is iu my collection from Ekeikei, where the species flies in
Jaunary and February.
198. Cyclodes pulchra spec. nov.
c^. Frons whitish ochreous ; vertex of head ochreous brown ; collar i>ale brown
tijiped with wliitisli : thorax brownish, barred with lilac-grey ; metathorax and tijis
of jiatagia yellowish brown, barred with whitish ; abdomen whitish grey with fine
dark segmental divisions, with a triangular reddish brown dorsal patch. Primary
greyish brown, with bronzy green irrorations over the basal half ; the outer half
is paler and strongly irrorated with lilac-grey ; a bronzy green ocellus below the
costa encircled successively with fine lines of bluish, of black and of greenish white,
edged externally by the antemedial j)ale line, which is angled and double below
the ocellus : a small patch of bluish scales is also below the ocellus ; medial line
darkish, slightly obscure ; postmedial line strongly waved and irregular to vein 3 ;
edged indefinitely internally by whitish ; a broad blackish strijie angled in the
middle from the end of the costal vein to vein 3 on the termeu ; a dark subterminal
( 255 )
line, a whitish patch above the tormis. Secondary with base pale lavender-grey,
followed by a broad dark transverse band with goldeu-bi'own liairs ; a broad pale
lavender-grey band, in wliich is a cnrved dark line, fuilovved by a broad blackish
area nearly up to the ternien, with a shortish whitish-grey tornal stripe ; termeu
lavender-grey, in which is a fine black subterminal line ; fringes golden brown.
Underside with the ])ectns orange. Secondary strongly irrorated with lavender-
grey, with a postmedial dark sinuate line ; lower margin and end of cell sharply
outlined witli black.
?. Like the male but darker, and all the lines more defined, especially the
subterminal, which iu that sex is very evanescent.
Expanse : c? 84, ? 97 mm.
Hah. The type is in my collection from Mafalu, where the insect occurs in
August.
Paragarista gen. nov.
Palpi with second segment reaching the vertex scaled, end segment long,
smoothly scaled ; antennae simple ; legs with tiliiae spined. Neuratiou : Primary
with vein 4 from the angle, 3 and 5 close to angle, 6 from the upper angle; 7, 8
and 9 stalked, the latter given ofl:' from 10 to form the areole. Secondary with
veins 3 and 4 from the lower angle, 5 from close to the lower angle, 6 and 7 from
the upper angle.
Type : Paragarista alboiitriata B-B.
199. Paragarista albostriata spec. nov.
c?. Head and thorax blackisli, collar with greyish irmrations and broadly edged
with deep yellowish ; thorax with centre entirely deep yellow and proximal segment
of abdomen ileep yellow, rest of abdomen black with white anal tuft. Primary
black with short subbasal bluish-white line ending in a white spot in the fold,
medial curved line pale bluisli, ending in a small whitish spot and edging
internally, a brown pear-shaped mark which adjoins a deep black ovate [)atch
edged finely with jiale bluish, a large white patch on the inner margin confluent
with a pure white postmedial stripe expanding broadly into the tornus, having a
black patch on the inner margin, an irregular bluish white line from the apex into
the white stripe at vein 2 ; secondary wholly black.
Expanse : 58 mm.
Hab. Aroa River ; March.
Type in my collection ; other specimens in the Tring Museum.
~'ii). Ophiusa kebea spec. nov.
cj. Head and thorax cinnamon-brown ; abdomen pale ochreous brown.
Primary pale cinnamon-brown; antemedial line fine, oblique; postmedial line
curved outwards to about vein 4, then receding rapidly basewards almost to meet
the antemedial line ; subterminal line pale lilac, with a dark edging on each side,
slightly oblique, nearly straight ; termen with a fine .scalloped line close to it ;
terminal area slightly lilaceous ; the whole wing is sparingly and finely irrorated
with black ; the area beyond the postmedial line is darker ; reniform distinct, darkly
outlined. Secondary pale ochreous grey, with a broad dark postmedial area tapering
towards the tornus, but not extending up to the termen.
17
( 256 )
Expanse : 64 mm.
Hub. The type is in my collection from the Kebea Range, where it was taken
in July ; otlier examples in the Tring Mnscum from the Aroa River in Fobrnary.
This species is somewhat near 0. certior Walker.
201. OpMusa lilacea spec. nov.
(?. Head and thorax pale cinnamon greyish brown ; abdomen greyish brown.
Primary with the area np to the medial line suffused with pale lilac, without
any lines ; medial line dark, crossing the end of the cell, slightly obliipie, (juite
straight, followed by a pale line which is edged by a dark line, slightly inter-
ru2jted beyond the cell ; this is edged by a fine lilac line np to vein .5, where it is
produced forward and is angled nearer the costa, becoming finely brown ; the wing
is cinnamon-brown beyond this line, but shades very gradually into pale lilac-grey
at the termen ; three white points on the costa before the apex ; a dark spot between
veins 6 and 7 below the third white point. Secondary pale greyish to tlie end of
the cell, from whence it is dark to the apex, there being a jjale short tornal stripe ;
termen whitish up to vein 6.
Expanse : 66 mm.
JIab. The type from the Kebea Range is in my collection, where the insect
occurs in July.
202. Ophiitsa subumbra spec. nov.
2 . Head and collar cinnamon-brown, finely and sparingly irrorated with black ;
thorax pale brownish grey, finely irrorated with black ; abdomen brown. Primary,
base j)inkish brown, becoming greenish brown in the median area and quite dark
at the postmedial line. A trace of a fine short basal line, and also of an indefinite
medial line ; postmedial line slightly oblique, double, the inner one being dark
reddish and the outer one dark grey ; the line is curved slightly from vein 7 to the
costa ; a trace of a pinkish serrate double-angled subterminal line ; area beyond
the postmedial line darkish grey ; termen deep nmber-brown, except at the apex.
Secondary uniform dark brown. Underside, both wings dark umber-brown, without
any lines or marks.
Expanse : 66 mm.
The type is in my collection from the Kebea Range, where the species occurs
in July.
203. Ophiusa ekeikei spec. nov.
c?. Head and thorax pale grej-ish brown, abdomen pale brownish grey. Primary
with base np to medial line ochreons brown, irrorated with pale grey ; a short basal
line from the costa into the cell ; medial line outwardly oblique, waved, double, with
its centre ])ale ; beyond this line the wing is pali' lilac-grey, shading rapidly into
brownish ; postmedial line consisting of three or four golden-brown serrated indefi-
nite lines, edged externally by a fine blackish line ; the whole is very broad and
darkish, and is strongly excurved from about vein 7 to vein 2 ; a trace of a dentate
pale subterminal line ; termen lilac-grey ; the costa is broadly irrorated with
ochreous grey to well beyond the postmedial line. Secondary blackish, paler at the
base, a broad oblique medial stripe ; termen with a white patch above the apex, and
( 257 )
below it, and also at the tornns ; the latter, with the medial strijie, is slightly bluish
in parts.
Underside : Primary with a broad whitish, slightly interrupted, oblique
band; secondary strongly irrorated with grey, and with a blackish patch at the
tornns.
Expanse : 65 mm.
Hub. The type is in my collection from Ekeikei, where the species flies in
March and April.
This .species is probably a local race ot O. melicerte Drury, but the colour differs
considerably, whilst the composite postmedial baud is also diverse.
204. Ophiusa serratilinea spec. nov.
(?. Head and tiiorax reddish brown, with a plum-colonred bloom over all ;
abdomen slate-grey. Primary pale reddish brown, with a jinrplish suflfusion, more
or less over all the wing, but specially over the basal half; a short waved pale
chestnut stripe with a dark centre ; antemedial line irregular, broad, waved and
indented, nearly erect ; postmedial lino serrate, angled about vein 9, followed by
a similar fine dark reddish line, with a pale external edging with whitish points at
each crennlation on the veins ; apical patch as in 0. iUibata Fab. ; termen broadly
bright lavender, barely reaching the toruus. Secondary dark blackish grey, with a
steely blue reflection ; a pale medial stripe angled at vein 2 ; termen bright lavender,
not extending quite to the apex.
? . Like the male, but altogether darker.
Expanse : S 75, ? 80 mm.
Ilab. The type is in my collection from Ekeikei, where the species occurs from
January to April. We have specimens from all the localities, but it is by no means
common.
205. Ophiusa dentilinea spec. nov.
? . Head and thorax dark purplish brown, finely irrorated with greyish ;
abdomen greyish brown. Primary dark reddish brown ; basal line short, whitish
from the costa ; antemedial and medial lines whitish, waved, parallel, enclosing
a white dot in the cell ; postmedial line whitish, dentate, strongly produced
outwards on vein 7, receding rapidly below, and produced forwards again to a less
extent to vein 2, below which it again recedes to the costa ; area between medial
and postmedial lines very dark, area beyond the latter paler and purplish brown ;
a dark oblique dash from the apex ; a dark spot on veins 3 and 4, below one another,
and a dark spot near the tornus on vein 1 ; termen finely edged by a dark scalloped
line. Secondary uniform dark brown.
Expanse : 52 mm.
Ilab. The type is in my collection from the Keliea Range, where the species
occurs in March and April.
20G. Ophiusa acutissima spec. nov.
? . Head, thorax and abdomen greyish brown. Primary with base dark umber-
brown, edged by a whitish strongly waved line ; median area, basal part pale pinkish
grey, gradually shading into darkish olive-brown, and edged externally ; the dark
( 258 )
postmedial line is edged by a whitish line ; this line is very acutely angled along
vein fi, where it is liighly produced outwards, receding in a short curve, and being
jiroduced forward to a less extent between veins 4 and 5, below where it recedes to
vein 2, and is again angled forwards ou vein 1 ; area from this line greyish brown,
with a creamy whitish patch on the costa above the acute angle; subterminal line
interrujited, obscure, with a small dark jiatch at the apex and a smaller one below
it; reniform dark grey. .Secondary dark greyish brown, with a broad iudetinite
medial line and a dark straiglit oblique postmedial line from the tornus to about
vein 6, the ground-colour being very dark on the inside of this line and pale greyish
on the outside, being edged with whitish near the tornus.
Expanse : 40 mm.
Hah. The type is in my collection from Ekeikei, where the insect flies in
January and February.
2117. Ophiusa subacuta spec. nov.
S. Head and thorax olive-brown, abdomen with a greyish tinge. Primary
with base olive-brown, edged by a double line, the first being dark, the second jiale
pinkish ; a short basal line on the costa ; median area jiinkish, edged internally by
the antemedial line, but extending over the medial dark curved line, beyond where
the area gradually shades into olive brown, becoming very dark on the postmedial
line, this line being produced strongly forward into a very acute angle about vein 6,
where it recedes in an irregular curve, and is obtnsely angled forwards about
vein 2 ; area from this line pale pinkish brown, a dark dash from the apex into the
acute angle ; termen edged by a fine scalloped dark line. Secondary uniform dark
brown, with a short tornal whitisii dentate dash in a darker subterminal area.
Ex])anse : 52 mm.
Jhtb. The type is in my collection from the Kebea Range, where the species
occurs in March and April.
208. Ophiusa insig'nifica sjiec. nov.
i. Head and thorax dull olive-bruwn, abdomen greyish brown. Primary
purplish brown, darker from the medial to the postmedial line, reddish beyond
the latter ; antemedial line waved, slightly oblique, followed by a white dot in the
cell ; medial line dentate near the costa, waved below the cell ; postmedial line
dark, sharply defined, edged externally with white, angled obtusely outwards about
vein 6 ; apex dark, with a dark dash to the angle ; termen pale lilac, edged with a
fine dark purjdish scalloi)cd line. Secondary uniform dark brown.
Underside with a crenulate postmedial line in the primary. Secondary with
three lines across the wing, the postmedial being strongly crenulate.
Expanse : 44 mm.
Jl((b. Tiie type is in my collection from the Kebea Range, where the species
occurs in March and A])ril ; other examples are in the Tring Museum from the
Aroa lliver.
209. Ophiusa vulgaris siiec. nov.
?. Head and thorax pale brown, abdomen greyish brown. Primary pale
lilac-brown up to the end of the cell, becoming bronzy-brown and very dark at the
postmedial line, beyond where the wing is wiiitish lilac-grey, except at the apex,
( L^59 )
which is lustrous ciiestiint-brown ; iiutemciliul ihirk line straight, slightly ()l)li(iiie ;
postmedial line whitish, curved near the costa, then straight to the inner margin.
Secondary dark greyish witli a medial whitish line ; a whitish small patch on the
termen near the toruns ; fringes pnrc white below the apex.
Expanse : 50 mm.
Hub. The tyi)e is my collection from Ekeikei, where the species occurs in
January and February.
210. Ophiusa medioalbata spec. nov.
?. Head ami tliorax bright orange, abdomen dark brown. Primary dark
brown with two small basal whitish spots ; a broad white median band and a
white subreniform small costal patch midway between the white band and the
aj)ex ; fringes white. Secondary uniform blackish brown, with white fringes.
Expanse : 48 mm.
Ilab. The type is in my collection from Dinawa, where the insect occurs in
May and Juno.
This sjiecies will come between 0. froniiiuis Drury and latizonu Btl.
211. Ophiusa sublutea spec. nov.
?. Head and thorax pale orange-brown, abdomen orange-yellow with a large
triangular black dorsal patch in front of the anus. Primary deep reddish with an
indefinite black basal line not lower than the cell, a grey dot in the cell ; reniform
grey for its lower half; an interrnpted deeply serrated pale subterminal line, ending
in a suboval pale grey tornal ])atch ; a black spot in the deep serration below the
apex, and a smaller one in the angle below vein 4. Secondary uniform chrome-
yellow, with two black confluent spots on veins 5 and G.
Expanse : 60 mm.
Ilah. The type is in my collection from Ekeikei, where the species occurs in
March and April ; other examples are in the Tring Blusenm from the Aroa River.
1 have a male from Mafalu (GOOO ft.), but the primaries are creamy colour ; it has,
however, every appearance of being worn, and I liave therefore described the fresh
female. The species will come next to O. Jigcrimiuans.
212. Ophiusa kenricki spec. nov.
cJ. Head and thorax bright chrome-yellow, the latter with a broad central
stripe of purplish red ; abdomen yellow. Primary chrome-yellow, with the
orbicular a small grey spot finely encircled with red ; reniform grey, finely
encircled with red ; a reddish purplish patch on the costa by the ape.x ; a much
larger similar coloured patch at the tovnus, in which is an oval greyish sharply
defined patch occnjiying the tornus itself ; the pur|)lish reddish colour extends
along the inner margin, tapering rapidly basewards to a fifth from the base ;
terminal area between the two patches very broadly i)ale reddish ; termen spotted
with grey ; a few grey spots in front of the ai)cx rcjiresenting the subterminal line.
Secondary pure spotless chrome-yellow.
Expanse : 78 mm.
Ilab. The type is in my collection from Mafalu, where it occurred in August ;
it is a very beautiful insect, and a rare one.
( 260 )
213. OpMusa pallida subplaga subspec. uov.
c?. Similar to 0. pallida Walker, but in the secondary there is no broad black
band, its place being partially occupied by a largish black oval patch in the
postmedial area between veins 3 and G.
Hab. Mafaln, 6000 ft., August.
Type in my collection.
214. Lagoptera pratti spec. nov.
(J. Head and thorax greenish lilack, with mctathoracic tuft tipped with red;
abdomen chrome-yellow, with three aual segments blackish. Primary deep black,
with a small basal whitish spot; a creamy ovate spot in the cell, followed by a
larger snbovate spot at the lower angle of the cell, above which is a similar coloured
dash. Secondary with the basal two-thirds chrome-yellow, terminal third black.
Exjiause : 98 mm.
Hab. Mafaln, 6000 ft., August.
Type in my collection.
Pseudophyx gen. nov.
Palpus roughly scaled, second joint reaching vertex of head ; third joint
smoothly sealed, of moderate length : antennae long, very finely and minutely
ciliate ; proboscis fairly developed ; legs hairy, hindlegs with retractile tufts on
the femora and tibiae ; thorax smoothly haired, abdomen without dorsal tufts.
Neuration : Primary with vein 2 from about the middle of the cell, 4 from tiie
angle, 3 from well before, S from directly above the angle, 6 from below the
angle, 7, 8, 9 and 10 stalked ; areole minute, vein 7 from its extremity, 8, 9 and 10
lieing on a long stalk. 11 lying close above 10, touching it for some distance.
Secondary with two internal veins, 4 from the angle, 5 directly above the angle,
G from the upper angle, 7 from above the angle, 8 bent down near the base to
touch 7, then rising imujediately in a bold curve. Primary of male with a large
cell on its nndersurface between veins la and 2, filled with long downy soft hairs ;
a fine pencil of coarser hairs between veins 2 and 3. Secondary of male with
a large tuft of long hairs from the underside of the coSta, and on the npperside
a large horny glabrous patch occujiying the basal and median areas to below
vein 2. Primary with costa straight, then slightly depressed, rising rapidly to the
apex ; termen slightly produced to vein G, then receding greatly to the toruus
at vein la, which is considerably shortened. Secondary with costa strongly curved
to vein 7, where an obtuse angle is formed ; termen gently curved from that point
to the tornus.
Type : Pseiidopln/j- jirutti B-B.
This genus is probably near Ophyx Guenee.
21."). Pseudophyx pratti spec. nov.
i. Head iind thorax darkisli reddish, colliir deep crimson-brown, abdomen
greyish. Primary pale rufous ochreous, somewhat muttled; three fugitive darkish
subbasal spots ; orbicular small, finely encircled ; a dark irregular jmstmedial jiatch,
edged internally by a curved dark line, which crosses the obscure reuiforra ;
snbterminal line (edging externally the aforementioned patch) crenulate, deeply
excised below vein 3, adjoining which is a very dark broad stripe confluent at
( 261 )
the torims with the equally dark outer lialt' of the inner margin ; termeu broadly
dark, with a pale spot below vein 7 well inwards and a pale streak between veins 4
and 2 edging the excised part of the previous band ; apex slightl}' paler. Secondary
with a glabrous patch occupying the basal and medial area ; termen and abdominal
fold mottled dark and paler rufous.
? . With head reddish, thorax and dorsum of iirominal segments pale orange-
reddish, abdomen dark grey. Primary with base and medial area orange-fawn-
colour ; postmedial area dark reddish, edged by a fine irregular line of white
scales ; terminal area broadly reddish orange, mottled with brown. Secondary,
orauge-fawn ; medial and postmedial area reddish, edged by an irregular line of
whitish scales ; terminal area very broadly reddish orange mottled with brown.
Expanse : c? 62 ; ? 64 mm.
Hab. Mafalu, Babooni, Ekeikei, and Aroa River, March, August, and September.
Type in my collection, and other examples in the Tring Museum.
Thoracolophotos gen. nov.
Palpi upturned, second segment reaching above vertex, thickly and closely
scaled, end joint moderately long, scaled ; antennae of male long, fasciculate ; thorax
tufted from the prothorax, spreading widely out over the metathorax ; legs thickly
scaled ; midtibia with a tuft at the femoral joint. Neuration : Primary, vein 2
from near middle of cell, 3 and 5 from close to the angle, 4 from the angle, 6 from
the upper angle, 7 from 8 at the end of the areole, which latter is given off from 10
to form the areole, 9 absent ; discocellulars very deeply recessed from the upper
angle ; cell covered with a tuft of hairs. Secondary, cell very short, not more
than a third ; vein 2 from near end of cell, 3 and 4 from angle, 5 from just above
the angle, 6 and 7 from upper angle ; costa straight to near apex, suddenly
depressed to apex, which is acute ; termen straight, oblique. Secondary fairly
ample ; termen evenly rounded.
Type : Thoracolophotos ekeikei B-B.
216. Thoracolophotos ekeikei spec. nov.
cJ. Head and thorax lavender-grey, witli a few pale ochreous scales ; thorax
with a large crest from the prothorax sjireading widely backwards; abdomen greyish.
Primary lavender grey, sparingly irrorated with ochreous, more abundantly on the
fold and inner margin ; a trace of a dark basal line ; two autemedial fine dark lines
angled in the cell ; three more or less obscure similar medial lines ; an obscure
postmedial one ; subterminal line dentate, rather palely edged externally, a small
whitish patch below vein 2 at aliout midway ; a large subtriangular dark purplish
brown cloud from midway along the costa to vein 3, and up to the costa about
a quarter from the apex. Secondary greyish, darker towards the termen.
Expanse : 51 mm.
Jlab. The type is in my collection, from Ekeikei, where the species occurs from
January to April.
217. Isoura pratti spec. nov.
£?. Head and collar dark umber-brown, the latter finely tipjied with whitish ;
thorax pale brownish with a slight lilac reflection ; abdomen yellowish, darker
towards the anal segments. Primary uniform j)ale brownish, with a sliglit lilac
reflection ; antemedial line waved, oblique, obscure ; postmedial lino nearly straight
( 262 )
to vein 9, where it curves slightly to the costa ; a hlaek spot is in this line on
vein 1 ; termen with black points ; reniform jnst visible as a slightly darker line ;
the wing is very sparingly irrorated with fine dark points. Secondary yellowish
with termen broadly dark grey, tapering rapidly to the toriins below vein 2, fringe
yellow.
Expanse : 52 mm.
IfaL The type is in my collection from Ekeikei, where the species occurred in
lilareh and April.
218. Hypaetra pratti spec. nov.
cj. Head and thorax deep crimson-brown, abdomen dark greyish brown, with
deep crimson tuft on the proximal segments. Primary, base and postmedial area
imrplish brown with lilac irrorations ; a small costal dark red subbasal s]iot with
a smaller one below it ; a large triangular dee]) velvety red-brown patch, occupying
the lower basal half of the wing, edged with white externally ; above its apex a
similar coloured costal spot almost confluent with it ; a broad oblique medial band
olive-green for its lower half, becoming pinkish towards the costa : in this band
is a waved crimson line, a large inverted triangular deep velvety red postmedial
patch with its base on vein G ; above its inner angle is a costal spot, and from
its outer angle rises a short line to the costa ; area from vein 6 to the costa
crimson : this is all margined by the broad pale lilac-grey postmedial area ; termen
broadly olive-brown. Secondary uniform darkish brown.
Expanse : 54 mm.
Hak The ty]ie is in my collection, from the Kebea Range, where the sjiecies
occurs in March, April, Mny and June. We have it also from other localities.
210. Hypaetra novoguineana spec. nov.
cJ. Head and thorax pale reddish brown, collar with pale lilac irrorations,
abdomen brownish. Primary reddish purjilish brown, at the base, between the
spots on the costa and in the postmedial area, especially on veins in the latter
part; a subbasal and a jiostmedial large deep velvety red-brown patch mostly
edged with creamy whitish ; above these patches to tlie costa the colour is
crimsonish brown, lietween them it is pale cinnamon red ; the postmedial waved
grey line edges externally the latter spot ; snbterminal line grey with two deep
angles, the lower of which has its apex about vein 5, from where it takes a curve
to the tornus ; termen with fine black points. Secondary uniformly brown, with
a trace of a ])ale postmedial line, apex and tornus, with fringes palely sjwtted.
? . Just like the male iu all particulars.
Expause : (? 46 mm. ; ? 50 mm.
I/ab. The type is in my collection from Ekeikei, where the insect occurs from
January to April, ^\'e have it from other localities, and other examides are iu the
Tring Museum from the Aroa Itiver. This sjiecies is one of the commonest in
British New Guinea.
221). Hypaetra subpunctata spec. nov.
cJ. Head and thorax greyisli brown, iibddnieii brownish. Primary brownish
grey with a tinge of lilac iu it ; base irrorated with ]iiMkish lilac, with a small dark
costal sjiot ; a deep red-brown broad subbasal strijie expanding rapidly below the
cell, and paler iu the cell ; the external edge of this stripe is straight, nearly erect
( 203 )
and palely edged ; medial greyish line obsoui-e, very irregular, immediately followed
by the fine dark interrupted highly irregular dentate postmedial line, which is
strongly produced forwards about vein S and rises in a small costal triangular
dark patch ; beyond this the ground-colour is finely irrorated witii whitish grey ;
snliterminal line sharply serrate ; termen with a fine scalloped dark line. Secondary
uniformly brown.
Undersurface : Both wings nnifcn'mly brown, the secondary with four dark dots
beyond the cell, a small one on the costa, and another between veins 7 and 8,
between 5 and 0, and below vein lb.
? . Like the male, but darker, and frequently has no dark dots on the under-
surface of the secondary.
Expanse : <? 48 ? 52 mm.
Nab. The tyj)e is in my collection from Ekeikei, where the insect occurs from
January to April ; other specimens in the Tring IMuseum from the Area Kiver.
Lasiopoderes gen. nov.
Palpus with second segment upturned, heavily haired, reaching above the
vertex ; third segment moderately long, naked ; antennae minutely bipectiuate ;
legs heavily haired, mid- and hindtibiae with large heavy tufts. Neuration :
Primary with vein 3 from well before the angle, 4 from the angle, 5 from just above,
6 from a little below the upper angle, 7, 8, 9 and li.» stalked. Secondary with two
internal veins, 2 from the middle of the cell, -i from three-quarters, 4 and 5 from the
lower angle, 6 absent. Wings triangular, expanding widely at the termen. Termen
produced outwards at vein 7, then receding gently to tornus ; inner margin hollowed
below the cell. Secondary small, with a tuft of long hairs from the costa ; the
greater part of the underside occupied by a patch of thick androconia.
Type Lasiopodf/rs pratti B-B.
221. Lasiopoderes pratti spec. nov.
cJ. Head greyish, tliorax dark brown ; abdomen greyish brown. Primary
greyish brown, with the area below the cell ; the median and postmedian areas dark
brown, deeply hollowed twice at the outer margin, and with a paler costal patch, with
dark antemedial and medial lines. Secondary uniform pale brownish.
Expanse : .'55 mm.
Hab. Babooni, September; Aroa Kiver, March.
Type in my collection, and other examples in the Tring Mnsenm.
Hirsutipes gen. nov.
Palpi with second segment heavily fringed with hair reaching above the vertex,
third segment long, smoothly scaled ; antennae of male with very minute and fine
hairs, so fine as to aj)pear simple except under a strong lens ; legs, forefemora and
tibiae heavily haired ; mid- and hindfemora and tibia of male densely clothed with
very long hairs ; midtarsi clothed with similar hairs, spined to the end of the tarsi.
Neuration : Primary, vein 4 from the lower angle, ft from just above, 0 from the
upper angle, 7 from just beyond the angle, 8, 9 and 10 stalked, the latter from
about midway between the angle and the apex, 8 to below apex, 9 into apex,
10 bent down and approximating to 8 just beyond 7 ; no areole. Secondary, 4 and 5
from lower angle, (1 and 7 from nppcr angle.
Type : llirmtijirs trifusciata Swinh.
( 264 )
222. Hirsutipes trifasciata Swiuli.
This species was described by Swiulioe as a Ihjpaelra, but it does not really
belong to that genus, and I can find no genus similar to it.
llab. New Guinea, Kebea Range, and Aroa Kiver, March and April ; specimens
in my collection and in the Tring Museum.
223. Baniana costiplaga spec. nov.
(?. Head and collar dark red-brown, finely freckled ; thorax and abdomen pale
chocolate-brown. Primary pale lilac-brown, with two dark chocolate-brown
triangular patches on the costa, from which rise the antemedial and postmedial
lines ; the former is nearly perpendicular, but slightly irregular, the latter is
angled sharply outwards from the dark patch, receding basewards from the angle ;
the area between these lines has a slight reddish tinge ; termen darkly spotted ;
orbicular represented by a dark point in the cell, and the reniform by two dark dots.
Secondary uniform cinnamon-brown.
Expanse : 32 mm.
Hab. The type is in my collection from Dinawa, where it was taken in
September.
224. Acantholipes bilineata spec. nov.
cj. Head, thorax, and abdomen pale ochreous grey. Primary pale brownish
ochreous ; antemedial line indistinct, angled inwards in the cell, with dark points
at each angle and on the costa ; postmedial line more distinct, darkly pointed,
projected strongly forward up to vein 6, receding rapidly below vein 4 ; a j)ale
irregular subterminal line of shading, from which np to the termen the ground
colour is rather darker ; termen darkly dotted ; reniform large and dark. Secondary
whitish ochreous, with a dark spot in the cell ; termen darkly dotted.
Underside : both wings pale ochreous grey. Primary with a large dark oval
ring at the end of the cell ; secondary with a dark cell-spot and a dark waved
postmedial line.
Expanse : 26 mm.
Ilab. The type is in my collection from Ekeikei, where the species occurs in
March and April.
225. Acantholipes ochreovenata spec. nov.
? . Head and collar pale chestnut-red, thorax pale ochreous grey with a dark
central line Primary pale ochreous grey; antemedial line whitish, angled outwards
in the cell, edged externally by a broadish stripe of sepia-brown interrupted at the
veins ; postmedial line whitish, angled outwards along vein 7, receding slightly
to 3, then erect to the inner margin ; a largish costal sepia-brown patch edges
internally this line and also fills the interior of the angle, but is interrupted at the
veins ; a subterminal dentate line of pale shading ; in front of the apex the costa
is finely dark, with three pale intersecting dots ; termen rather darkly shaded, and
darkly dotted ; all the veins stand out pale ochreous, interrupting all the marks
except the whitish lines. Secondary uniform pale brownish.
Expanse : 25 mm.
Ilah. The type is in my collection from Dinawa, where it was taken in
September.
( 265 )
226. Acantholipes dinawa spec. nov.
c?. Head, thorax, and abdotneu cream-colonr ; both wings creamy white, with
a broad dark brown obIi(pie stripe from the apex of the primary carried through
both wings, but beginning in the secondary at vein 7. Primary with a double
ochreous external edging to the brown stripe ; a subterminal line of dark shading ;
termen with fine dark points ; cell watli two terminal black points below each
other. Secondary with lines exactly as in the primary.
Expanse : 30 mm.
Hah. Dinawa, September ; Aroa River, March.
Type in my collection, and other specimens in the Tring Museum.
227. Acantholipes aroa spec. nov.
(?. Head, tliorax, and abdomen and both wings pale whitish yellow. Primary
with a trace of an angled medial line and an obscure darkish subterminal line
inclined inwards at vein 3 or 4 ; two dark dots at the end of the cell below each
other. Secondary paler and clearer than the jjrimary.
Expanse : 25 mm.
Hab. Aroa River, January and March.
Type in my collection ; other examples in the Tring Museum.
228. Acantholipes hampsoni spec. nov.
?. Head, tliorax, and abdomen pale greyish. Primary ]iale ochreons grey,
with basal and antemedial lines obscure ; medial and postmedial line oblique
dark, the ground between them being darkly filled in ; beyond the latter is a
broad pale grey stripe, edged obliquely externally by three lines — the first darkish
grey, the second pale ochreous, the third finely dark brown ; beyond this the
ground colour is darkish, edged by a tine double line of pale and darker shading;
termen darkly dotted; frmges silvery grey, with a darker central line. Secondary
paler than the primary, with all the primary markings carried more or less
distinctly through.
Expanse : 29 mm.
Hah. The type is in my collection from Dinawa, where it was taken in
September.
229. Acantholipes purpurascens s])ec. nov.
(?. Head and thorax purplish reddish brown, abdomen greyish. Primary
purplish chocolate-brown, with the antemedial and postmedial lines pale, interrupted
and obscure ; a broadish median band of paler ground-colour ; subterminal line
whitish, waved, bordering the dark ground colour, from whence the terminal area
is lilac-grey ; termen darkly dotted. Secondary piniiish brown, paler towards the
base.
Expanse : 24 mm.
Hab. The type is in my collection from Dinawa, where it was taken in
September.
230. Acantholipes arcana spec. nov.
(?. Head, thorax, abdomen, and both wings pale rufous, more or less finely
irrorated with darkish grey. Primary with a basal white point, three snbbasal
white points in a greyish line, a medial slightly angled grey line, a postmedial
( 266 )
row of white dots in .1 grey line, ii ]iali' rnl'ous dciitiitc snliteriuiMal line, ti>rnicii
with black dots. Secondary with a medial grey line, postiuedial grey line with
a row of white dots, a terniinal row of Mack dots, inside which is a scalloiied
grey line.
Expanse : 38 mm.
Ilab. Aroa llivcr, Fcl)rnary.
Type in my collection. There is a specimen in tlie Tring Mnseum wliicli is
very dull in colour, with but little of the red tone in it.
231. Acantholipes alboserrata spec. nov.
c?. Head, thorax, and both wings greyish brown, with a basal white point;
an antemedial dark line, edged internally with white on the fohl ; medial dark
line irregular ; reniform with two black points ; a broad white serrate irregular
postmedial line, a subterminal i)ale dentate line; termen with black jioints.
Secondary with a medial grey line, a serrate white postmedial line; termen with
black points.
Expanse : 34 mm.
Hab. Aroa River, March.
Type in the Tring Mnseum.
232. Acantholipes nigriplaga spec. nov.
tJ. Head and tliorax reddish oclireuus, abdomen blackish ; both wings reddish
ochreons. Primary with blackish antemedial spots on the costa and inner margin,
connected together by a tine line; a reddish irregular medial line; two black
dots in the reniform ; three large blackish postmedial spots, connected by a fine
erenulate dark line ; a jiale reddish subterminal line, with small blackish spots
on the external edge ; termen with black dots. Secondary with a grey medial
indefinite line, a postmedial erenulate line strongly marked with blackish, a pale
.snbterminal line; termen with lihuk dots.
Exjianse : 37 mm.
Hab. Aroa River.
Type in my collection.
233. Fodina kebea sj)ec. nov.
9. Head and thorax deep velvety reddish brown, patiigia and tufts tip))ed
with a double line of crimson and white, abdomen reddish brown. Primary with
the basal four-fifths deep velvety reddish brown, edged above externally with
])ale cream-colour; the costa is broadly pinkish up to the cell, which colour
makes an irregular deep excavation into the dark area at and beyond the end
of the cell ; terminal area iiinkish lilac. Secondary uniform orange vermilion.
Expanse : 46 mm.
Hab. The type is in my collection from the Kebea Range, where it was taken
in Jnly.
234. Ischyja kebeae spec. nov.
?. Head and thorax greyish brown, abdomen darker. I'rinuiry dnil umber-
brown, darkish up to the oblii|ne straight medial line ; a trace of an antemedial
line on the costa; an obliipie chestnut-red patch from the apex expanding inwards,
and descending in a dentate fascia to vein 3, edged outwardly with blackish.
( 267 )
Secondan', base pale brownish, termea and inner margin very broadly blackish, a
large central jiatch of white.
Expanse : 104 mm.
Hab. The type is in my collection from the Kebea Hange, where the species
occnrs in July.
This species will come next to /. cbttxa Swinh.
~35. Platyja arg'snteopunctata spec. nov.
S. Head and thorax dark purplish brown, finely but very sparingly irrorated
with grey. Primary very dark purplish brown, almost blackish ; a double silvery
grey dot on the costa near the base, followed by another farther out in the cell,
beyond and above which are three similar dots on the costa below each other in
a cnrve, two dots at the end of the cell and one small one in the middle, below
which on vein 1 is another ; a curved postmedial line of small silvery grey spots
terminating in a larger dash on the inner margin ; a fine dentate subterminal
silvery grey line. Secondary uniform dark brown.
Expanse : 58 mm.
Hab. The type is in my collection from Dinawa, where the insect occurs in August.
236. Platyja porphyrodes spec. nov.
S. Head and thorax dark chocolate-browu, with a few fine whitish scales;
abdomen dark greyish brown ; legs densely haired, dark purplish brown with a
few whitish scales ; tarsi encircled with cream-colour. Primar}- dark greyish
brown, witli a slight purplish lustre ; antemedial line short, irregular, brownish
ochreous, represented below the cell merely by a single small spot on vein \b ;
medial line represented by two ochreous brown dots on veins \b and 2 ; a small
ochreous brown spot above the cell near the end ; a fine postmedial crenulate
ochreous brown line, terminating in a black-edged ochreous brown blotcli between
veins 2 and 4; fringes ochreous yellow. The wing is sparingly and finely irrorated
with minute whitish scales. Secondary dark brown with a slight purple lustre
confined below and in the cell for tlie basal three-ijuarters ; a fine crenulate and
dotted ochreous brown postmedial line from the inner margin to vein 0. Fringes
pale ochreous yellow.
Exjmuse : 69 mm.
Hab. The type is in my collection from the Kebea Range, where the species
flies in March and April.
237. Platyja pratti spec. nov.
S . Similar to P. porplii/rodcx B-B., liut in the primary there are three ante-
medial ochreous brown spots, a medial waved scalloped line from the angle of
vein 2 to the inner margin, with an ochr(!ous brown spot at each scallop, the spot
above the cell as in jjorphjrodes; the jwstmedial line is broader, and edged with
black internally ; the termen is finely dotted with ochreous brown, and the fringes
are black. In the secondary the whole wing is sufi'used with puriilisii ; the post-
medial line is broader, edged internally with black ; and the fringes are black.
? . Like the male.
Expanse: 68 mm.
Hab. The type is in my collection from Ekeikei, where the s]iecies flies from
January to April. I have other exami}les from the Kebeii Kange.
It is a close ally to /', jiorj/lii/rodc.-<, but is without doubt a distinct s[iecies.
r 268 )
Plasmaticus <^en. nov.
Palpi with second segment thickly scaled reaching to the vertex ; end segment
short, shortly scaled ; antennae simple ; legs roughly scaled, spiued. Wings :
costa curved for the basal fifth, straight for tliree-fifths, highly arched at a fifth
from apex ; apex acute ; termen excavated below apex to vein 5, which is prolonged,
and below which the termen is angled, receding gently obliquely to the toruns.
Secondary with termen straiglit from vein 7 to vein 2, where it is angled in an
upward curve to the tornus at \b. Neuration : Primary with veins 3, 4 and 5
from close to the lower angle ; vein 5 ascending to well above the middle of the
termen. Secondary with veins 2, 3, 4 and 5 from the lower angle.
Type : Plasmaticus angulata B-B.
I have ventured to diagnose this new genus from the female moth as far as
possible, there being no doubt that though somewhat near Platijja it is quite
distinct from it.
238. Plasmaticxis angulata spec. nov.
? . Head and thorax chocolate-browu with scattered white scales ; abdomen
dark grey. Primary chocolate-brown, sparingly irrorated with fine white scales ;
a basal costal patch of white scales, another patch at a third, and another patch
darkly centred at three-fifths ; a white spot in the cell ; a trace of an antemedial
line ; postmedial line waved below vein 4, dark, a broadish paler area beyond it ; a
subterminal row of white dots ; three small greenish grey patches of scales by the
apex, placed in a triangle ; a larger patch between veins 3 and 5, and another
below 2. Secondary brownish grey ; termen with lavender scales below vein 3,
edged internally broadly with reddish ; a white dash in the angle between veins lb
and 2.
Underside lavender-grey, with a postmedial and subterminal ochreous grey
spotted line in both wings, the postmedial in the secondary being bisected by a
dark dividing line.
Expanse : 58 mm.
Hah. The type is in my collection from Dinawa, where the insect flies in
August.
239. Episparis angulatilinea spec. nov.
S and ? . Head, thorax, and abdomen jiale cinnamon-brown ; both wings pale
cinnamon-brown. Primary with a slightly darker small basal costal patch ;
submedian line twice highly angulated above the lower margin of the cell, then
strongly oblique basewards ; a black point in the cell, two snow-white small spots
at the end of the cell, the lower one the larger of the two ; an irregular indefinite
angled dark line ; postmedial line whitish, rising by a white costal dash angled at
vein 9 ; below where it is irregularly curved outwards to vein 2, where is a slight
internal angle ; beyond this line a slightly curved band of whitisii, the area outside
which is pale below vein 6, and has a darkish crcnulate subterminal line in it.
Secondary with a dark dash closing the cell, an angled crenulated dark medial
line followed by the darkish postmedial line, with a white centre, which is angled
between veins 3 and 4, subterminal area greyish in the excised area.
Expanse : c? and ? , 58 mm.
( 269 )
Hab. The type is in my collection from Dinawa, where the insect occurs in
the month of May. We have specimens from other localities also; and in the
Tring Mnseum, from the Aroa River, are other examples.
Parepisparis gen. nov.
Palpi small, scaled; third segment minute; antennae of male loipectinate ;
legs with forefemora expanded at base and hollowed ; a tuft of longish hairs at
the tibial joint. Neuration : Primary with vein 3 from before the angle, 4 from
the angle, 5 from the centre of the discocellulars, 6 from the upper angle, 7 from
the end of the areole, 8 and 9 stalked, given off from 10 to form the areole, 10
anastomosing with 11, forming a spurious areole, 11 anastomosing with 12 shortly.
Secondary with vein 3 from before the angle, 4 from the angle, 5 from the centre
of the discocellulars, 6 from the angle, 7 behind the angle, 8 touching 7 to half
the cell. Primary, costa nearly straight ; termen excavated below apex, angled
slightly at veins 6 and 5, receding rapidly to toruus. >Secondary with termen
crennlate, strongly so at veins 4 and 6.
Tj'pe : Parepisparis crenulata B-B.
240. Parepisparis crenulata spec. nov.
(?. Head and thorax greyish lilac, abdomen lilac-grey; both wiugs greyish
lilac, with very dark olive markings. Primary with two siibbasal spots, one near
the costa and the other on vein 1 ; two short outwardly oblique costal dashes,
one near the middle and one be3'ond ; an inwardly oblique fine line from the apex
slightly curved below the cell to the inner margin, with spots on each side of
it in the fold; from the tornus upwards two jJair of spots — viz., one pair between
veins 1 and 2 and the other on each side of vein 3 — and a pair of small spots
well beyond the cell near the termen ; a small dark spot at the end of the cell.
Secondary with a submedial indefinite waved line, a medial almost crennlate line,
three small spots above the tornus ; most of the wing is finely irrorated with grey.
Expanse : 45 mm.
Hab. The type is in my collection from the Kebea Eange, where it was taken
in July.
241. Eugorna alboarcuata spec. nov.
(?. Head slate-brown, thorax reddish brown, abdomen sooty grey. Primary
reddish brown, with a creamy white costa extending into a broad white arc across
the postmedial area into the tornus, margined externally by a chocolate-brown
curved line, vrith a second obscure line beyond; apex and terminal area dark greyish
brown, paler and tinged with lilac above the tornus. Secondary black, with a
white tornal patch ; on the underside the white arc is yellow.
Expanse : 65 mm.
Hab. Aroa Kiver, March.
Type in the Tring Museum.
242. Ophideres dinawa s|)ec. nov.
(?. Palpi and antennae bright carmine-red, head and thorax dark crimson
brownish, abdomen brownish. Primary darkish lilac-pink nj) to an oblique line
from the apex to a third from the base, beyond which the colour is pale greenish
golden brown, except for a triangular red patch below vein 2 in the angle made
( 270 )
by the obliiine line; in fVniit (if the apex there is also a triangular costal pale
red jiateh; a trace of an aiiteuiedial reddish line : renifurni deep crimson ; fringes
golden greenish brown. Secondary dark brownish grey.
?. Like the male, but darker, whilst the golden brown colour is less golden
and jialer.
E.xpanse : c? T4, ? 60 mm.
Hal). The type is in my collection from Dinawa, where it occurs in August.
243. Plusia kebea sjhc. nov.
S. Head and thorax beaver-grey, abdomen j)ale greyish. Primary beaver-grey,
with two subtriaugular dark bronzy lines ; the larger one, enclosing the larger
triangle of the ground colour, is from the costa, and meets the base of the
innermarginal one on vein 2 ; the terminal area is very strongly suftused with
dark bronzy colour nearly up to these lines. Secondary dark grey, becoming
creamy at the base.
Expanse : 33 mm.
Jlab. The type is in my collection from the Kebea Range, where the insect
flies in JIarch and April ; other examples are in the Tring Museum from the
Aroa Kiver.
244. Plusia dinawa sjiec. nov.
5 . Head and thorax lilac-grey, abdomen grey. Primary, basal area pale lilac-
grey margined ; a slightly oblique antemedial line from the inner margin extending
up into a silvery c^ mark, behind which is a silver spot, and margined costally by
a very obliipie line extending into the npj)er stroke of the silvery c^ mark ; post-
medial line waved, angled inwards on the fold ; the median area between these
lines is dark and suffused with rich bronzy brown ; a large terminal suffusion of
the same colour from the apex to vein o, terminating along vein 2 ; tornal area
j)ale lilac-grey ; the subterminal line is distinct near the toruus, but very obscure
in the dark area. Secondary greyish brown.
Expanse : 35 mm.
Ilah. The tyjie is in my collection from Dinawa, where the insect occurs in
August.
245. Plusia ekeikei spec. nov.
(?. Head and thorax jiale golden brown ; jiatagia pinkish brown, with white
scales ; abdomen pale ochreous brown. Primary, basal area pale pinkish bnflf,
mottled with golden brownish ; the area from the tcrmen to the cell below the
angle of vein 2 brilliant pale golden, divided by the postmedial and subterminal
lines, which are deep golden reddish, with a dasli along vein 5 connecting the
two ; reniform dark golden brown. Secondary ochreous at the base, broadly dark
grey beyond the cell.
Exjjanse .• 38 mm.
Hub. The type is iu my collection from Ekeikei, where the insect occurs in
July.
•,'l(i. Plusia babooni siiec. nov.
6 . Head and thorax darkish grey, abdomen somewhat jialer. Primary dark
bronzy grey, with obli(jue subbasal stripe angled near the costa; jiostmedial stripe
ol]lii|ue, slightly waved, shortly angled above the inner margin ; subterminal area
(271 )
with a very large ami irregular cloudy patch of dark bronzy brown iu a ])aler
area ; a small silvery spot near the angle of vein 2. Secondary dark brownish
grey, with the basal half somewhat paler.
Expanse : 34 mm.
Ilah. The type is in my collection from Babooni, where it was taken in
September.
247. Polychrisia kebeana spec. nov.
c?. Head pale grey, thorax and collar dark brownish grey, thoracic tuft pale
chestiuit-brown ; abdomen dark brownish grey, ventral tufts reddish browa.
Primary with basal area pale grey uj) to the whitish subbasal line which terminates
on the lower margin of the cell ; from this line up to the angled and irregular post-
medial line the colour is dark brassy brown, the brassy lustre showing more or less
according to the angle of light ; a posterior costal patch in this area of pale grey;
in the angle of vein 2 a golden (J-shapcd mark with its internal part also goldenish,
followed by a golden spot ; tornus with a brassy brown patch ; rest of wing pale
grey, with a short oblique irregular brassy brown streak from the apex; termen
finely whitish with a narrow dark bronzy brown internal edging. Secondaries
brownish grey.
Expanse : 48 mm.
Ilah. The type is in my collection from the Kebea Range (6000 ft.), where the
insect occurs iu March and April.
248. Deva multicolora spec. nov.
i. Head chestnut-brown, thorax chestnnt-brown tinged with purplish ;
abdomen greyish brown. Primary golden brown, restricted base lilac, followed by
a snbtriangular costal patch of golden brown and by an irregular broadish band
of metallic silvery brassy colour; median line obli(pie ; renifiirni distinct, slightly
metallic golden brown, the ground colour being tinged witli purplish ; postmedial
line highly angled just below the costa and very oblique, touching which are
several silvery brassy patches, a large one up to the tornus, above which is a small
one, followed above by another slightly larger ; an angled, curved and irregular
short line from near the apex to about vein 3, edged externally with golden.
Secondary brownish, pale near the base.
Expanse : 39 mm.
Hub. The type is in my collection from the Kebea Range, where the insect
occurs in July. This species will come next to 1). caclonoUi Koll., which I have
from the same district.
249. Marcipa tripuncta spec. nov.
$ . Head, thorax, and abdomen grey. Primary mauve grey, with three silver
spots at the base ; a trace of an antemedial line ; rcniform jialely outlined ; post-
medial line from near apex very oblicjue ; beyond this the area is paler, and has a fine
line near the postmedial slightly waved to about vein 4, where it is highly angled
and recedes to the inner margin in a deep curve. Secondary greyish brown.
Expanse : 39 mm.
Hub. The type is in the Triiig Jlusi^uni iVdUi the Aroa liiver, where it was
taken in March ; there is also a siu^cimin in the British Museum from Milne Bay.
18
( 272)
Subfamily FOCILLINAE.
250. Mecodina novoguineana spec. nov.
S. Head and thorax dark brownish gre)', tlie latter finely irrorated with brown ;
alidonipn rather ])alcr. Primarj- brnwnisli, finely irrorated with whitish grey;
a small dark brown eostal basal iiatch ; a jiale inwardly dentate anteniedial line
edged externally with a broad brown band ; a very broad irregular clouded brown
band at the end of the median area, edged externally by a pale serrate line ; a white
s])ot beyond this line in the tVild ; a snbterininal cloudy dark inwardly serrate line
edged internally with yellowish ; ternien (inely brown ; fringes brown intersected
with yellowish. Secondary brownish, with a deep velvety black ottellus (above the
tornus) having a small white pnpil, and being encircled externally with yelluwisli,
the encircling being doubled internally.
Exjjanse : 46 mm.
I/d/i. The type is in my colleetiun from the Aroa Klver; other sjiecimens are
in the Tring Museum.
2.51. Seneratia albopunctata spec. nov.
c?. Head and collar fuscous, tiiorax jiur|ilish brown ; abdomen greyish brown.
Both wings chocolate-brown. Primary with indistinct basal and snbbasal curved
waved lines ; a medial broad obliipie dark band diffused on its outer edge ; a sinuate
crennlate postmedial line ; a siunate subterminal row of white dots ; termen with
a very fine crennlate line. Secondary with all the markings of the primary carried
through except the basal and snbbasal lines.
? . Like the male, but with the pattern less defined.
Expanse : cJ 40 mm., ? 48 mm.
l/d//. The type is in the Tring Museum from the Aroa River, where it was taken
in February.
252. Zethes lilacea spec. nov.
?. Head, collar, and thorax pale fawn-brown, head and collar the palest ;
abdomen fawn-brown. Both wings lilac-brown up to the oblique postmedial
line, beyond which the colour is pale pinkish lilac, being slightly darker near the
termen ; nearly touching the jiostmedial line is a series of dark dots with a
jiale grey spot on the costa in the primary only ; reniform darkish. Secondary
like the primary.
Expanse : 30 mm.
I/td. The type is in my collection from the Kebea Range, where it was taken
in .Inly.
253. Zethes fuscomarg'iuata spec nov.
?. Ileud, thorax, and abdomen cinnamon-brown. Primary pale cinnamon-
brown uj) to the postnieilial line, and narrower along the outer ])art of the costa to
the apex, beyond this line dark sepia-brown ; a trace of a fine basal line ;
autemedial line fine, waved, and curved ; reniform pale, with a fine oblique dash
from it to the costa; postmedial line oblique, highly angulated just below the costa ;
a trace of a pale submarginal irregular lino, with a dilFused small pale patch between
veins 4 and 5. Secondary cinnamon-brown, finely darkly irrorated to the end of
tlie cell ; postmedial line fine, just beyond which is an irregidar deep sepia-brown
( 273 )
band from the tornns to vein (1, edged with pale fawn-eolonr, and with the veins
intersecting it with tlie same colour ; above this, below the costa, are two white
spots ; apical area darkly diffused.
Expanse : 51 mm.
Tliib. The type is in my collection from Dinawa, where the species occurs in
May and June.
The species will come near Z. shit-ida Guen.
254. Zethes aroa spec. nov.
Head and thorax pale straw-colour, collar yellowish ; abdomen pale straw-
colour. Both wings pale straw-colour. Primary with antemedial, medial, and
jwstmedial lines fine waved, somewhat curved, and the postmedian angled below the
costa; posterior part of wing darkly irrorated, more densely at the termen ; a pale
patch at. the apex, preceded by a dark angled dash. Secondary with a diffused
median band and a curved irregular iiostmediul line ; from the former the whole
of the wing is more or less densely and darkly irrorated, v«ry much so in the
terminal area.
Expanse : 51 mm.
Ihd). The type is in my collection from the Aroa Iliver ; other examples are
in the Tring Museum from the same locality.
255. Zethes griseistriga spec. nov.
¥. Head, thorax, abdomen and both wings dull sooty grey. Primary with
dark snbbasal line angled outwards; a medial line of dark shading angled outwards
below the costa ; postmedial dark line very oblique, angled shortly outwards just
below the costa. (Secondary with obliipie waved medial and straightish oblique
postmedial lines.
Expanse : 42 mm.
Ilab. The tyjje is in my collection from Dinawa.
256. Zethes meeki spec. nov.
S. Head and thorax lavender-grey, collar tipped with dark brown, abdomen
pale reddish grey. Both wings pale greyish. Primary with two snbbasal waved
lines filled in with pale olive-brown ; a medial irregular and a postmedial angled
line, the interspace darkly suffused with brownisli, and having a costal jiatch of
reddish brown ; an apical brownish snfTnsion, an irregular serrated submarginal
whitish grey line, followed by a subterminal row of white dots. Secondary with
the median suffusion edged by the postmedial obli(jne line ; terminal area with
markings as in the primary ; a dark spot at the tornns, and two dark points in
the cell.
?. Like the male, but without the brownish suffusions in both wings.
Expanse : 38 mm.
Ildlj. The type is in the Tring Museum from the Aroa Uiver, where it was
captured in April.
Tills species will come next to Z. jjhimipes Hpsn.
257. Zethes megaspila Warr.
?. Head, thorax, abdomen and both wings pale bluish grey ; costa with three
subdued small brownish patches, the third exteniling into a large brown patch
below it ; a slightly darker apical suffusion on the termen ; a trace of a basal dark
( ii74 )
spotted lino, .1 pale waved postmedial line, an iire.ixnlar serrated snbmarginal dotted
line ; tcrnien with black jioints. Secocdai'}- with a black point in the cell ; the
postmedial and other onter lines as in the jirimary ; a dark spot at the tornns.
Expanse : 42 mm.
II(i6. The type from British Now Gninea (Lo Ilnntc) is in the Tring Mnsonm.
Mr. Meek sent it from the Aroa Rivor, where it was taken in March.
258. Zethes nigra spec. nov.
?. Head and thorax blackish, collar and patagia greyish ; both wings blackish
grey. Primary with costa whitish grey to l)eyond the cell; basal and subbasal
lines interrnjited, waved ; medial line very obscured ; jiostmedial line serrate,
produced forwards beyond the cell ; snbterminal line dark, serrated above the
tornns; a white spot in the cell, followed by the creamy reniform ; apex whitish.
Secondary with the basal area darkly mottled; the jiostmedial line strongly crenn-
late, with dark spearhead crenulations ; snbterminal line very prominent and dark,
preceded by a row of pale reddish brown spots touching the line.
Expanse : 34 mm.
JIab. The type is in the Tring Musenm from the Aroa River, where it was
taken in JIarch.
259. Zethes tessellata spec. nov.
5. Head, thorax and .abdomeu pale ochreons grey, thorax marked with dark
brown ; botli wings pale ochreons grey with dark brown lines. Primary with a
short basal dash confluent into the basal line ; antemedial line curved ; two angloil
medial lines enclosing a slightly darker aroa; reniform darkly outlined and divided ;
postmedial line angled below vein.'); above the angle is a greyish patch up to
the costa, adjoining the irregular snbterminal line ; terminal aroa from this line
yellowish ; tormen darkly lunnlod. iSocondary with antemedial and very obscure
medial linos curved slightly inwards; postmedial line straight; subterminal line
very obscure ; termen darkly lunulod.
Expanse : 39 mm.
I/iiL The type is in the Tring Musenm from the Aroa River, whore it was
taken in March.
This species will come next to Z. n/f/riliiwa Wlk.
200. Zethes ochreistriga spec. nov.
6. Head and collar pinkish ochreous, thorax crimson ochreons, abdomen
crimson grey, laterally ochreons. Primary reddish ochreous, with antemedial Hue
pale ochreons edged with pink ; medial line irregular dark reddish, a small ochreous
sjiot in the coll ; rcnifonii ochreous with two black dots ; postmedial lino cronulato,
roughly angled outwards ojiposite tiie reniform ; subterminal line spotted dark groy,
with internal ochreous edgings to the spots ; angled at vein 5 ; termen with fine
dark points. Secondary crimson irrorated with grey, with ochreous base and a
yellow medial stripe from vein G to the tornns ; postmedial line obscure, with
strong grey irrorations in jiarts beyond it ; termen very finely black and crenulate.
?. Much paler than the male, with the lines finer and greyer.
Expanse : 34 mm.
JIab. The tyjic is in the Tring Museum from the Aroa River, whore it was
captured ill March.
f ii75 )
201. Zethes apicebnxnnea spei-. nov.
d . Head and collar jiale rusty lnowii irnnated witli grey, thorax and abdomen
pale jirey ; both wings j)ale grey. Primary with eosta sprinkled with jialo rnsty
brown ; a trace of an obscure medial and postmedial line ; a large apical sub-
triangular rusty-brown patch occupying nearly half of the costa, with a spot on the
termen at its extremity ; orbicular just discernible by its faint encircling ; both wings
are sparingly irrorated with darker scales.
Expanse : 4'2 mm.
Ilab. Ekeikei and Area River, IMarch and April.
T^'pe in my collection, and other specimens in the Tring Museum.
262. Zethes multilinea spec. nov.
c?. Head, thorax and abdomen sepia brown, thorax and abdomen with a dark
slaty grey central stripe; both wings dark sepia-brown, with all tiie veins palely
outlined; termen pale slaty grey, with a tine dark crenulate line ; fringes dark
sepia-brown.
E.xpanse : 42 mm.
Hab. The type is in my collection from Dinawa, where it was taken in August.
263. Zethes (Cultripalpa) ekeikei s])ec. nov.
?. Head and thorax reddish chocolate-brown, abdomen chocolate-brown; both
wings reddish chocolate-brown. Primary with a narrow creamy white costa; a
broken slate-grey snbbasal line ; a small slate-grey cell-dot ; a postmedial crenulate
slate-grey line projected outwards about veins 3 and 4, inside which projection is a
slate-grey roundish patch ; a waved snbmarginal row of slate-grey dots ; termen
with fine points. Secondary greyish in costal area : a trace of a medial crenulate
line ; three submarginal grey dots near the tornns ; termen finely dotted ; fringes
to both wings whitish.
Expanse : 42 mm.
Ilai. The type from Ekeikei is in my collection; the species occnrred in January
and February.
264. Zethes (Cultripalpa) rufalia spec. nov.
?. Head and abdomen reddish brown, thorax purplish brown; both wings
reddish brown. Primary with base pale buff; subbasal line dark, edged by a
broad median band of purplish brown; reniform palely outlined; an indistinct
jwstmedial line, a more distinct and spotted submarginal line ; termen with
fine black points ; a costal snbapical cliocolate triangular patch jialely edged.
.Secondary paler and redder, with the postmedial line dotted with white ; two
blackish dots above the tornus ; median and terminal area finely irrorated with
dark grey.
Expanse : 37 mm.
Ilab. The type is in the Tring Museum from the Aroa River, where the species
occurs in April.
265. Zethes (Cultripalpa) albisigillata Warren.*
?. Head reddish, thorax and abdomen pale greyish buff, both wings pale
greyish ochreous. Primary with a reddish suffusion beyond tlie median area ;
base with two dark dots ; a broad blackish median band with a silver dot in the
• A'iw. Zool. X. p. 123 (190:i).
( 27(i )
cell and u waveil silvt'i' liiif cld.siiig tlie i-i'll ; a oistal suliapical blackish triaiij^nlar
jiatcli, 11 !julj;uar<;iiial incj^ularly waved dotted liue : termeii with fine bhick points.
tSecoiuhiry with a trace ol" a jiale spotted medial line and a fine crennlate suhmarjjiiial
line; termen with fine black points ; the wing is in parts tiuelj' irroratcl with grey.
Expanse : 44 mm.
llah. The tyjie is in tiie Tring ^luseiim iVom British New Gninoa (Le Hnnte)
Mr. Meek sent the species from the Aroa Uiver, where it was taken in February.
200. Zethes (Cultripalpa) albopunctata spec. nov.
6. Head, thorax, and aiidomen reddish brown. I'rimary brownish, densely
irrorated with dark grey, coslu sjKitted with snow-white, a trace of a subliasal
dark line; postmedial line dark, crennlate, with white points at the tips of the
crenidations ; an indistinct paler snbterminal line, termen with black jmiiits.
Secondary slightly reddish brown, finely and darkly irrorated; subbasal and post-
medial lines as in the primary, with an indefinite dark medial line of shading;
terminal area as in the primary.
Exjiause : 4t) mm.
Hah. The type is in the Tring Miisenm from the Aroa River, where it was
taken in Februar}'.
207. Zethes (Cultripalpa) bella spec. nov.
?. Head and paljii dark grey, thorax pinkish grey, abdomen pink; both
wings pale pinkish with very pale lavender-grey lines. Primary with the costa
broadly very pale lavender-grey, edged internally near the base with white ; a
trace of the medial and postmedial lines on the inner margin ; above the latter
a large dark grey blotch extending in two lities at the toj) and bottom through
the fringes, and preceded on its inner edge by an interrupted short curved white
line margined internally with a jiale brown Innule. Secondary with a very restricted
white base, a dotted submedial line, the medial and submedial and terminal lines
pale lavender-grey, the medial with a series of central white dots in it, the terminal
line consisting of a spotted row.
Expanse: 34 mm.
llah. The type is in my collection from the Aroa River, where the Rpecies
occurs in March ; other sjjecimens are in the Tring Museum from the same locality.
20s. Zethes (Cultripalpa) rubraria spec. nov.
?. Head, thorax, abdomen, and both wings reddish ochreoas, costa with six
white points, snbmedial line red-spotted ; a trace of a similar medial line ;
postmedial line sinuate, almost crennlate, palely pointed at the crenulations ; an
irregular waved pale snbterminal line, beyond this the termen is darker and with
black points. Secondary with the markings of the primary carried through.
E.\panse : 30 mm.
Hab. The type is in the Tring Museum from the Aroa River, where the insect
occurs in April.
269. Pangrapta pratti spec. nov.
S. Head and collar reddish brown, thorax paler reddish brown, abdomen
dark grey. Primary with the costal half reddish brown, lower half dark sepia-
brown in the median area ; l)ase paler, edged by a pale irregular subbasal line ;
a medial dark line below the cell ; postmedial line white, strongly produced
( 277 )
outwards from vein G to 3, where it is deeply excavated to the inner margin
with a white Bufrusion iii the excavation ; an oblique crenulate paler subtcriuinal
line ; termen with a dark suffusion abont veins 3 to 5. Secondary with the white
postmedial line followed by the crenulate subterminal line, the median dark
diffusion more or less restricted.
Expanse : 36 mm.
Ilab. The tyi)e is in my collection from the Kebea Range, where it was taken
in March and Ajiril. We have examples from several localities, and others are
in Tring Museum irom the Aroa lliver.
27U. Pangrapta aroa sjiec. nov.
t?. Head and collar dark sepia-brown, thorax pale iiinkish brown, abdomen
darkish grey. Primary with the basal half pale pinkish grey, edged by a broad
oblique dark sepia-brown fascia produced outwards at vein 6, with a costal pale buff
triangular jiatcb above ; the rest of the wing bluish grey with a trace of a dotted
subterminal line ; a subbasal irregular fine pale lino, succeeded by a dark dot in
the cell. Secondary with a dark median band, a dark cell-spot encircled with
whitish ; a white postmedial line edged externally by a diffused dark sepia-brown
stripe with white serrations above the tornal area ; terminal area bluish grey, with
a dark suffusion across it to veins 3 and 4. Margin of both wings crenulate and
finely dark.
Expanse : 38 mm.
Hah. The type is in my collection from the Aroa Iviver, where it was taken in
March ; other specimens in the Tring JIuseum from the same locality.
-71. Disticta hampsoni spec. nov.
(?. Head, thorax, abdomen, and both wings cream-colour. Primary with a trace
of a greyish medial oblique line, two black points at the end of the cell; jiostmedial
line oblique from the apex, greyish, edged by an external oehreous stripe with a
blackish crenulate outer margin; a trace of a subterminal line roughly jtarallel
with the termen, termen with fine black points. Secondary with a black point
in the cell, postmedial lines as in the primary; terminal area broadly pinkish
with a greyish stripe, and fine black jioints to the termen.
? . Like the male, but darker, with all the lines accentuated.
Expanse : S and ? , 37 mm.
Hah. The type from Ekeikei is in my collection. We have examples also
from other localities taken in March and April, and there are several specimens
in the Tring Museum from the Aroa River.
272. Disticta tornopunctata spec. nov.
c? ? . Differs from D. Iirunp.toiu in that there are two blackish spots in each
wing above and a little inwards from the tornus.
Hah. The type is in my collection from the Kebea Range ; and we have the
insect from other localities, including the Aroa River.
I am doubtful whether this may not be a variety of hampsoni B-B., but it
obtains in the same localities as that insect and at the same time ; we have a good
series of the one, but only a small number of the other; there are, however, no
intermediate forms, and as wc have the spotted species in both sexes it is probable
that it is distinct.
( 278 )
2T3. Disticta kebea spec nov.
cJ. lleail, thorax, and abdomen pale pinkish cinnamon-brown ; snbbasal line
obscure, pale ; a dark point at the base, another in the cell, and a tliird at the lower
margin of the reniform ; an olilii[ne line from the apex basewards, the lower part
of whieh is edged externally witli white, a pale costal snbapical jiatch ; a trace
of a pale subterminal Hue with three dark dots from veins 2 to 4 ; termen with
fine black points ; all the lines and dots carried through the secondary except the
snbbasal one.
Expanse : 4U mm.
Ilab. The type is in my cnllection from the Kebea Range, where it was taken
in Jnly.
274. Axiocteta rufa spec. nov.
3. Head, thorax, and primary pale reddish lirown, abdomen and secondary
creamy white. Primary with double autemedial greyish indefinite lines somewhat
waved ; medial and jiostmedial lines obscure ; snbterminal line double waved.
Secondary with pink fringes and with an antetermiiial row of dark jioints.
?. Like the male, bnt lines more distinct, witii a white sjiot on the fold below
the end of the cell.
Expanse : S 34 mm., ? 33 mm.
Ilab. Owgarra and Kebea Range, July.
Type in my collection.
This species will come next to .1. cenophx Turner.
2~b. Axiocteta turneri spec. nov.
(?. Head and thora.x warm brick-red, abdomen creamy with reddish dorsum.
Primary warm brick-reddish, with the postmedian area greyish ; two hyaline spots
in the cell, the outer one the larger ; a trace of a snbbasal line ; medial line grey,
obliqne, obscure : iiostmcdial line frec^ueutly waved for the costal two-thirds ; sub-
terminal Hue obscure ; termen white in the upper and lower excision. Secondary
creamy white, pink at the toruns and termen.
Expanse : 28 mm.
Ilab. Ekeikei, Arua River, Marcli and April.
Type in my collection, and other specimens in the Triug Museum.
276. Axiocteta babooni spec. nov.
(?. Head and thorax dull reddish brown; abdomen creamy with reddisli
dorsum. Prinuiry dull reddish brown, with antemedial and postmedial grey lines
obscure; a creamy white spot in the fold below the end of the cell; reniform
obscurely grey with a similar coloured indefinite dash to the costa ; a subterminal
row of dark jioints. Secondary creamy white with pinkish termen.
Expanse : 3IJ mm.
Ilab. Babooni, September ; Aroa Kiver, March.
Type in my collection, other specimens in the Tring Museum.
277. Axiocteta obliqua sjiec nov.
S. Head and thorax jiale pinkish grey, abdomen greyish white. Primary
pinkish grey, with antemedial grey Hue somewhat curved; jwstmedial line straigiit,
oblique ; a creamy white sjwt at the end of the cell ; a reddish line of postmedial
(279 )
shading ; a grey dotted snbterminal line. Secondary creamy white, pinkish at the
tornus, termen with a row of fine dark points in front.
?. Like the male, but paler, and with a pale reddish spot at the end of the
cell.
Expanse : cJ 33 mm., ? 30 mm.
Ilab. Dinawa, September ; Aroa River, February to April.
Type in my collection, and other examples in the Tring Museum.
278. Axiocteta concolora spec. nov.
cJ. Palpi reddish brown, head and thorax jiiiikish ochreous, abdomen creamy
white. Primary pale ochreous cinnamon-brown, with antemedial, medial, postmedial,
and subterminal wavy grey, more or less obscure lines ; reniform obscurely grey.
Secondary, creamy white for the basal half, ]iinkish for the terminal half.
Expanse : 33 mm.
Hah. Kebea Range and Aroa River, March and April.
T3'pe in my collection, and other specimens in the Tring Museum.
279. Axiocteta flava spec. nov.
tj. Palpi darkish grey, head ochreous, tiiorax ochreous grey, abdomen creamy
whitish. Primary ochreous grey, with cell yellowish, in which the orbicular and
reniform are grey ; a trace of a grey medial line below the cell; an oblique grey
postmedial line; a trace of a serrated yellowish subterminal line; area below
vein 3 yellowish ; a subterminal series of fine dark points ; termeu finely dark.
Secondary creamy whitish, termen pinkish, especially just below the apex.
Expanse : 36 mm.
Hab. Aroa River, March.
Type in my collection ; one specimen only obtained.
280. Egnasia delicata spec. nov.
c?, Head, thorax, abdomen, and both wings pale lavender-grey. Primary
with a trace of a basal costal sjiot, a subbasal short dash from the costa, and a
broadish dark oblique dash from the inner margin ; two short oblique costal
dashes, viz. one at the middle and one beyond ; costa in front of apex dotted with
white ; a faint trace of a pale postmedial line ; all the dashes are of chestnut
brown ; a hyaline spot in the cell. Secondary with a large postmedial purple
blotch, extending into tlie tornus; termen strongly crennlate, pale chestnut with
white centre.
?. Similar to the male, but the markings on the primary much more prominent,
and with a large blotch from the postmedial line to the termen.
Exj)anse : S 3'.t mm., ? 4-1 mm.
Ilab. The type is in my collection from Ekeikei, where the species flies from
January to March ; we have it from other localities also, and examples are in
the Tring Museum from the Aroa River.
281. Egnasia chloroplaga spec. nov.
?. Head, thorax, and abdomen pale dove-grey, collar tipped with dark sei)ia-
brown. Both wings jiale grey tinged with lilac Primary with subbasal line
waved, somewhat obliquely curved ; medial line double waved and irregular ; cell
filled with pale green between these two lines with a black point ; a postmedial
( 280 )
broad diflFiised l)an<l with a short dark serrated line (i|i|iosite the cell, the costal ]>art
of the hand hciii'j pale greenish ; subniarginal line tine, irregular, anglcil at vein 4,
followed by a second line of dark dots, the interspace being faintly greenish, with
a small darkish subapical terminal patch. Secondary with a trace of the medial
and jiostmedial lines ; submarginal lines as in primary ; a greenish tornal patch
edged externally by a short dark brown line.
Expanse : 45 mm.
[lab. The ty[ie is in my collection from Ekeikci, where it was taken in March
and April.
282. Egnasia dinawa spec. nov.
S. Head and thorax grizzled jiale yeUowisli grey. I'oth wings grizzled
grejish brown. Triniary, with basal and snlib:isiil creamy irregular lines enclosing
an olive-brown area ; a diffused median band of daik shading preceded by a blackish
dot in the cell and followed by the reniform ]ialely outlined; post medial creamy line
deeply angled forwards about vein 0 ; a trace of a submarginal serrated fine
pale line; termon with dark points, and clouded with brownish below the aj)ex.
Secondary with the postmedial creamy line straight and oblique ; terminal area
as in the primary.
Expanse : 47 mm.
Hal. The type is in my collection from Dinawa, where it was taken in
September.
Crenularia gen, nov.
Differs from Aroatia in that vein 7 is free from the end of the areole, 8 and 0
stalked and given ofi' from 10 to form the areole, II appressed closely on to 10 at
a quarter of the areole, so as to look like anastomosis, becoming free beyond the
areole. Costa of primary depressed in the centre, falcate at the apex ; termcn
excised slightly below the apex, slightly hollowed to vein 3, below where it is
excised and crenulate. Secondary produced to a fine point at vein 7, and to a longer
point at vein 3, above and below which it is strongly crenulate. Antennae of male
bipectinate, ? very finely ciliate. Palpi porrect scaled, third segment shortish.
Type : Crenularia concolor B-B.
~>i3. Crenularia concolor spec. nov.
S. Head and collar whitish bufl', thorax, abdomen, and both wings uniform
pinkish grey. Primary with an angled irregular tine dark snbbasal line; orl>icular
grey, finely encircled by a dark line ; reniform dark grey with a small pale centre ;
postmedial line fine, irregular ; a short pale grey fascia before the apex. Secondary
with the lines carried through as in the primary.
¥ . Dark slate-grey, paler in the postmedial area of the primary, with the lines
and markings as in the male.
Expanse : S 32 mm., ? 38 mm.
JJab. The type from Ekeikei is in my collection, where the species flies in
March and Ajiril.
Aroana gen. nov.
Palpi sealed, upturned, second segment reaching above the vertex, third
moderately long; ]iroboseis, moderately developed; antennae very finely and
minutely ciliated in 6. Legs : one pair of spurs on the midtibia, two pairs on the
( -'81 )
hind. Nenration : Primary, vein 3 from just below tlie lower an.ijle, -I and .5 from
tlie angle, (5 from below the ujiper angle, 7, 8 and 9 stalked, given off from 10 to
form a most minute areole, 1 1 from well below the middle of the cell and extending
to a quarter from the ajiex. Costa straight ; ajiex excised ; termen jiroduced
strongly out at veins 5 to 3, excised below 3, and crenulate nearly throughout.
Secondary termen produced at vein 7, and strongly produced at 4 and 3, excised
and crenulate below.
Type : Aroana olicacea B-B.
284. Aroana olivacea :<pec. nov.
S and ?. Talpi red, head purplish black; thorax blackish olive-green;
alidonicu dark greyish with very dark olive dorsum. Both wings very dark blackish
grey with lavender lines. Primary, witii tine basal and subbasal internally crenulate
lines ; a broad dark olive median band angled just below the costa ; reniform
blackish, finely outlined with lavender ; postmedial line serrate, ])roduced outwards
about veins 4 and 5, and strongly serrate, receding below ; median and jiostmedial
areas olivaceous ; subterminal line very dark olive-brown, with an onter margin
spotted with lavender ; a brownish costal subapical patch. Fringes olivaceous.
Secondary with restricted base greyish ; antemedial line lavender, edged by a broad
dark olive band ; median area olive-green with a dark spot in the cell, edged by
a double line of dark brown and lavender; postmedian area olive-green ; subterminal
area dark with a broad band of fine lavender irroratious.
Expanse : 31 mm.
Ilab. The type is in my collection from the Aroa River, where it was taken in
April ; other examples are also in the Tring Museum from the same locality, but
taken in March.
285. Aroana rubra s])ec. nov.
cJ. Head, thorax, and abdomen brownish pink. Both wings jjinkish with olive
and golden-brown lines and marks. Primary with a short fine twice-angled basal
line ; an antemedial fine line shortly angled just below the costa, then nearly
straight but slightly outwardly oblique ; a dark broad median band edged by a
filling in of darker ground colour; reniform irregular, sublunular, very dark, palely
encircled, from whence a dark band descends to the inner margin ; an interrnj)ted
dentate postmedial line ; a golden-brown diffused subapical jiatch from the costa to
veins 4 and 5 ; termen finely dark, subcrenulatc. Secondary with a broad oblicjue
median band ; a dark sublunular spot in the cell ; a fine irregular postmedial line,
and a fine dark termen.
? . Like the male in all particulars.
Expanse : S 28 mm., ? 30 mm.
Ilab. The type from the Aroa River is in my collection ; I have specimens also
from Ekeikei, and other examples are in the Tring Museum from the former
locality. The species flies in March and April.
Focillodes gen. nov.
Palpus upturned scaled, second segment reaching vertex, third segment
short, antennae of S bipectinate to end, of ? minutely ciliate. Legs hairy ; foreleg
with tuft on n])perside of femora, and a small pencil of hairs on underside of tibiae ;
midtibia with one pair of si)urs, hiudtibia with two pairs of spurs. Neuration :
( 282 )
Primary with vein 2 from jnst beyontl middle of cell, 3 from just below the angle,
4 from tlie mifjle, 5 from jnst uhove the angle, 0 from the ui)iier angle, 7 atiasto-
mosing with lu to form the areole, 7, 8 and 9 stalked, 11 depressed on to lu, which
tenches 7 for its greater part. Secondary with veins H and 4 from the angle, 5 from
well above tjie angle, 6 and 7 from the npjier angle ; costa of primary straight,
slightly depressed over the cell ; apex depressed, almost acuminate : termoii
slightly angled about vein 5, receding from thence ra})idly to tlie toruus. Secondary
with costa straight ; termen fairly rounded ; wing broad.
Type Focillodcti (linaiva B-B.
280. Focillodes dinawa spec. nov.
t?. Head, thorax and abdomen dark i)arplish sooty brown. Both wings dark
slaty brown ; primary with a broad pale costal dash at and beyond the centre, below
and beyond which is a reddish area ; lower ])art of termen pinkish brown ; a dark
waved basal line, a dark waved angled medial line ; postmedial line obliipie, sub-
terminal line waved, orbicnlar ochreous. Secondary with a reddish median band
edged externally by a waved line, beyond which the area is wholly dark slaty grey.
Underside with both wings having the postmedial and terminal area dark grey,
edged internally by a pale line.
?. Wholly pale grey, with lines and costal dash as in the male ; subterminal
line spotted.
Expanse : c? 34, ? 40 mm.
Ilab. Dinawa, August and September ; Aroa River.
Type in my collection, and other specimens in Tring Mnsenm.
287. Focillodes brunnea spec. nov.
cT. Head, thorax, abdomen and both wings entirely dnll brown. Primary with
antemedial dark line waved, medial line angled at the cell ; jjostmedial line less
angled than the median, edged palely externally, a dark obscure subapical costal
cloudy patch ; snbterminal line dotted with white. Secondary with a dark medial
line, jiostmedial line as in the primary ; a dark dot above the tornus.
¥. Paler than the male, with rusty brown lines as in the male; in the
secondary there is a dark dotted subterminal line, which is also present in some
specimens of the male. Undersurface with no dark marks or area.
Expanse : c? 34, ? 4i.) mm.
Ilah. Ekeikei, Mount Kebea, and the Aroa River, March.
Tyjie in my collection, and other specimens in the Tring Museum.
288. Focillodes griseata s])ec. nov.
6. Head, thorax, abdomen and both wings entirely greyish. Primary with
waved and slightly angled obscure lines ; jiostmedial line edged laterally and
indelinitely with blackish, shortly angled at the costa; a black dash i'rom the angle
to the apex; a white dotted subterminal line; a costal pale obscure small patch
outside the angle of the medial and jiostmedial line. Secondary with median area
wholly blackish, edged externally by a blackish line with a pale edging ; an obscure
dotted subterminal line.
?. Like the male. Underside with a small ajiical blackisli patch.
Expanse : c? 36, ? 40 mm.
(283 )
Ilab. Dinawa, August; Kobea Range, May and June; and the Aroa River,
March.
Type in my collection, and other examples in the Triiig Museum.
2S9. Focillodes fulva spec. uov.
S • Head, thorax, abdomen and both wings uniform fulvous brown. Primary
with basal and medial lines waved and slightly angled ; postmedial line edged
indeiinitely with darker fulvous, oblique, straight, broadly angled below the costa ;
a fine dotted subterminal lino, a white costal apical dash ; apex acuminate, strongly
angled at vein 5, Secondary with a dark median fine line ; a dark broadly waved
postmedial line, edged externally with white and internally broadly darker fulvous ;
postmedian area jialer fulvous, with a trace of a fine pointed snbterminal line ;
termen angled at vein 4. Underside : Primary with apex having a whitish patch.
?. Wholly greyish, with lines as in tlie male. In the primary the inters])acc
of the postmedial and strongly dotted subterminal is fulvous to vein 2. Secondary
with the postmedial area whitish, with a black spotted subterminal line; termen
fulvous.
Expanse: S 42, ? 41 mm.
Ilab. Kebea Range, July ; Dinawa, August.
Type in my collection.
290. Focillodes medionigra spec. nov.
(?. Head, thorax, abdomen and both wings pale fulv<ius brown. Primary with
an obscure waved basal and medial fine line, the latter excurvcd across the cell ;
medial line slightly oliliijue, shortly curved just below the costa ; a black indefinite
band (if sluiding edges this line, being very broad internally on tlio inner margin and
extending in a broadish dash into tlic apex ; a pale dotted subterminal line. Secondary
with the whole of the median area blackish ; deep black at the oblique postmedial
line ; a pale spotted subterminal line ; termen dark fulvous. Underside of primary
with a dark apical patch.
Expanse : 4ii mm.
Ilab. Kebea Range and the Aroa River, March.
Type in my collection, and other specimens in the Tring Musenm.
Contortivena gen. nov.
Differs from Focillodes in that tlic arcole terminates at the end of the cell ;
veins 7, 8, 9 and 10 are stalked on a long stalk, 10 being given off nearer the apex
than 7 ; veins 4 and .5 are given off from tlie angle ; 2, 3, 4, 5 are waved and
contorted down towards the tornus. Tlie costa is waved and depressed at apex ;
tlie primary is more even in width and longer. The secomlary has tlie costa
produced into a lobe at the base, and is depressed in the middle.
Type : Contorticcna umbrosa B-B.
291. Contortivena umbrosa spec. nov.
3. Head, thorax, abdomen and both wings greyish brown. Primary with
basal line scalloped; medial line waved; jiostmedial blackish, angle at vein 7
obli(jnc and straight from tliat point; orbicular and reniform ochreons, darkly
encircled ; subterminal area [lale ochreons brown, reddish on its inner edge ;
subterminal row of dots. Secondary witii a medial dark line; postmedial line
( -^84 )
oblique, blackish, with pale external edging ; snliterminal dotted brown, ronghlj-
jiarallel with the jirevious line. On the undersurface the stigmata of the primary
are blaekish brown, palely encircled.
?. Both wings quite pale grey ; lines iukI pattern as in the male.
E.xpanse : 42 mm.
Hub. Rabooni, .Sejitember,
Type in my eollcctiou.
202. Capnodes costiplaga spec. nov.
(?. Head and thorax jmvplish, collar with a reddish tinge, abdomen greyish.
Both wings pnrplisli with pale olive-grey marks. Primary with three large white
spots on the costa, the third just beyond the end of the cell; three small obscure
snbbasal dots ; reniibrm marked by a double s[iot ; an irregular dentate waved
dotted i)ostmedial line ; an interrnpted subterminal line of larger spots, with a small
jiatch in the tornns and at the apex ; termen with two rows of ])oints. Secondary
with the jiattcrn exactly n'peated as in tlie primary, except as to the white
costal spots.
Expanse : 40 mm.
Ilab. Kebea liange, March and April.
Type in my collection.
203. Capnodes minuta spec. nov.
c?. Head white, thorax and abdomen pale reddish. Both wings pale reddish.
Primary with two large white costal spots ; a trace of a dark antemedial line at
the outer edge of the tirst spot ; a postmedial tine dark line, edged externally with
white points ; a small dark spot in the cell. )Secondary with a dark point in tiie
cell, the wing very finely irrorated in parts with grey.
Expanse : 18 mm.
JIub. Ekeikei, March ; Aroa River, January.
Type in my collection, and other specimens in the Trlng Museum.
294. Ceromacra babooni spec. nov.
S. Head, thorax, and abdomen greyish brown. Primary pale greyish brown,
somewhat paler on the terminal half, with a broadish indefinite darker brown basal
band ; snbbasal line narrower and irregular; median band very broad, followed by
a narrow irregnlar line ; jiostmedial band very broad indeed, and darker, beyond
which is an outwardly curved dark line which rises and terminates in the broad
I)ostmedial. Secondary greyish brown, pale greyish on the costal and abdominal
margins. The secondaries are much distorted below the cell, indicating the presence
of a sexual patch ; veins 1 and 2 are also distorted, bnt I can only trace a moderately
strong pencil of hairs lying above vein 1, and certainly insufficient to account
for the contortion of either the wings or the veins.
Expanse : .50 mm.
Ilab. The tyi)e from Babooni is in my collection, and was taken in Sei>tember
10(j:j.
20."i. Ceromacra ekeikei spec. nov.
?. Head and thorax yellowish brown, jiatagiae and collar irrorated with grey,
abdomen greyish brown. Primary ochreons brown with a broad indetiiiite greyish
baud beyond the cell ; terminal area paler ; antemedial line dark and irregnlar,
( 285 )
be3'ond which is a dark dot on the njiper margin of the cell ; a white point
terminates both the upper and lower angle of the cell, beyond which is a recurved
broadish darker indefinite baud ; the fine postmedial serrated recurved line lies in
the iiale greyish band, beyond which is a broadish area of darker ground colonr^
the outer margin of which is very irregular ; termen fine, very ^lale grey. Secondary
greyish brown with a fine very pale grey termen.
Expanse : 49 — .50 mm.
Hah. The tyi)e is in my collection, and was taken in January or February at
Ekeikei.
20(1. Ceromacra diuawa spec. nov.
? . Head pale brown, thorax brown ; patagiae and collar densely but loosely
scaled, the scales being paler tipped at their extremities, giving the thorax au
irrorated ai)pearance ; abdomen greyish brown. Primary brown, darker for the
basal half and at the termen, leaving the postmedial area broadly jiale ; lines
purplish brown ; the antemedial, rising on the costa in front of the centre, recedes
rapidly to the lower margin of the cell, whence it is angled outwards on to the
inner margin ; medial line broader, indefinite, irregular, beyond which the reniform
stigma is prominent ; postmedial lino strongly serrate (almost crennlate), out-
wardly produced from the costa, then receding gradually to the middle of the
inner margin, beyond which (postmedial line) is a trace of another similar one ;
subterminal line very irregular, forming the inferior edge of the darker terminal
area. Secondary uniform greyish brown.
Exjjanse : 49 mm.
Ilab. The type is in my collection from Dinawa, where the species occurs in
May and .June.
29T. Ceromacra purpurea spec. nov.
i . Head and thorax brown with a jmrple gloss, thorax with a central com-
pressed metathoracic tuft of strongly metallic purplish scales ; abdomen brown,
slightly puri)lish ; anal tuft tipped with white. Primary purplish brown, with
a j)ale yellowish brown antemedial line ; postmedial line pale, finely but darkly
edged internally, beginning on the costa a third from the apex, strongly oblique
outwardly to below vein 0, where the dark edging is very strongly serrated twice ;
the lower serration receding to below the cell, thence to the inner margin ; beyond
the middle the double line is resumed in an outwardly oblique direction ; a trace
of a subochreous subterminal line. Secondary blackish brown, slightly paler
towards the costa.
Expanse : 4i) — 41 mm.
Ilab. The type is in my collection from Ekeikei, where the insect occurs in
March and April.
2f»8. Ceromacra aroa spec. nov.
cJ. Head and thorax olive-brown, abdomen greyish brown. Primary olivaceous
brown, with basal, snbbasal and three somewhat indefinite medial lines, an oblique
broad jwstmedial dark band of shading, a subterminal dotted line, an obscure
terminal band of golden brown. Secondary uniform dark brownish grey.
Expanse : 82 mm.
Hub. Aroa River, February.
Type in my collection, ami in the Tring Museum are other examples.
( 286 )
299. Dierna lilacea spec. nov.
c?. Head and thorax very pale ciunamon-brown ; both wings pale cinnamon-
brown ; primaries with a dotible fine oblique dark postmedial line, the narrow
interspace filled in with lilac-grey and with a broad indefinite edging of lilac-grey ;
a dark point in the cell, followed at the end of the cell by a white point. Secondary
with the medial donble lino as that in the primary, and with a similar lilac-grey
edging.
Expanse : 46 mm.
Ilab. Ekeikei, January ; Diuawa, September.
Type in my collection.
Tornosinus gon. nov.
Palpus with second segment long, reaching above the vertex, scaled, fringed
above and below with hair; third segment minute, almost globular, fringed with a
long tnft of hairs above ; antennae simple ; legs with fore pair having femora and
tibiae heavily fringed with hair, mid and hind pair slightly fringed. Neuration :
Primary with vein 2 from middle of the cell, 3 from below the angle, 4 from the
angle, o from just above the angle, (i from below the upper angle, 7 from the
angle, 8, 9 and 10 stalked, 11 from near base of cell ; cell very short. Secondary
with two internal veins ; cell short ; 2 from near base of cell, 3 and 4 from lower
angle, .5 from directly above it, 6 and 7 from upper angle, 8 curved strongly upwards
towards costa and running parallel therewith. Primary with tornns deeply excavated
from vein 3 to middle of inner margin ; a heavy tuft of long hair on the underside
of inner margin. Secondary moderately ample; terraen slightly flattened ; tornns
shortened, with margin flattened from vein 3.
Type : Tornosinus riiger B-B.
3ii0. Tornosinus niger spec. nov.
S . Tiie whole insect is sooty black. Primary inclined to greyish beyond the
jiostmedial area, a short subterniiual whitish dotted line from in front of the apex,
a round white spot at the end of the cell ; termeu darkly dotted ; fringes white at
the excavated part of the tornns ; tornus deeply excavated from vein 3.
Expanse : 34 mm.
Hub. Aroa River, March.
Type in Tring Museum.
Rectangulipalpus gen. nov.
Palpus with second segment porrect, long, reaching above the vertex, heavily
scaled, fringed with long hair; third segment long, at right angles to second,
heavily fringed with lung liair. Antennae very fine, minutely ciliate ; head tufted ;
legs with forefemora and tibiae scaled, a rose-shaped tuft of long spatnlate scales
from the base of the secondary below. Neuration : Primary with vein 4 from the
angle, 3 and Ti from close to the angle, 0 from tiie upper angle, 7, 8, 9 and lU
stalked. Secondary with two internal veins, 3, 4 and S from close to the lower
angle, G and 7 from the uj)per angle. Costa depressed at base, then evenly curved,
with a short tuft of hair at the top of the depression on its underside ; wing very
briiad, termen evenly rounded from a subacute apex. Secondary less ample, with
terraen somewhat flattened ; a tnft of hair from the upper side of the costa.
Type : Rectangidipalpus mecid B-B.
(287 )
3nl. Rectangulipalpus meeki spec. nov.
cj. Thorax, abdomcu, and priiuary niiifonu dull brovvii, the latter with a small
pale ochreous cream}' spot at the end of the cell, and a trace of a waved subterminal
line. Secondary blackish browu, oraiigc-browu iti the mediau area aud iu a broad
baud to the tornus.
Expanse : 32 mm.
llab. Area River, February.
Type ill the Tring Bluseum.
liotli these last two genera are like iioue other with which 1 aiu tic'iiuiiiited,
bill I think their best position will be in I'rtmt uf (he Dclfoidimie, with which they
will forui a ci.uiiieeting' link.
Pseudodeltoida geu. nov.
Paljii with second segment long, well above vertex, thickly scaled so as to be
»l)atulate above ; third segment shortish, smoothly aud shortly scaled ; auteuuae
finely and minutely ciliate. Neuration : Primary with vein 4 from the augle,
3 and 5 from close to the angle, 6 from the angle, 7 given off from 8 before the
end of the areole, 8 and 9 stalked near apex, given off from 10 to form the areole,
lU with a short bar from II before the eud of the areole, which is small. .Secondary
with 3 and 4 from the angle, .3 from the middle of the discocellulars, U aud 7 from
the angle.
Type : I'seudodcUoida aroa B-B.
302. Pseudodeltoida aroa spec. nov.
S. Head and collar greyish; thorax and abdomen pale brownish, the latter
blackish on the dorsum for the anal half. Primary pale brownish with a small
dark subbasal costal spot; the outer part of the median aud the postmedian areas
dark brown ; cell and fold greyish, invading somewhat the dark area ; a dark brown
small subapical costal triangular indefinite patch ; a waved dark subterminal line
somewhat obscure ; costa with three ochreous jioints before the ajiex ; termen with
black points. Secondary brownish grey, darker beyond the cell.
Expanse : 3U mm.
Ilab. Aroa River, Februarj-.
Type in the Tring Museum.
le
( 288 )
ON THE BIRDS OF THE ISLAND OF BABBER.
By Dr. EKNST HAKTERT.
HAVING returned to the East; ouco more, our valued corrcsjioudenl BIr. Heiuricli
.Kiihu has agaiu commenced to collect birds on the less known islands
of the great Eastern Archipelago.
In Sor. Zoo/. l'JU4. jiji. 174— ::i21 I gave a list (if the birds collected by
ilr. Kidiu on the Honth-West Islands, and, as I said on ]i. 176, collections were
not made on the more eastern i.slauds, Sermatta, Luang and Babber. Mr. Kiihu
sent a parly of Malayan collectors to ISabber, who made a collection of 7t) sjiecies.
Unfortunately the collection does not seem to be a comideto one, as the work
on Babber had suddenly to be terminated, on account of the hostile behaviour
of some of the natives.
A few Babber specimens have long ago reached the Dresden Museum, and were
duly recorded and jiartly described as new by Ur. A. B. Meyer. Also Mr. Hchiidler
sent a collection of 31 species (about half of those sent by Mr. Kiihn) from
Babber to the Leyden Museum, and the latter has been recorded by Dr. Otto Finsch
in his valuable article "Systemat. Ueliers. der VOgel der Siidwcst-Iuseln " in Ao/ca-
Leijden Museum, xxii. pp. 225— 3U0 (1901).
The ornis of Babber differs considerably from that of the more western islands,
being in many instances the same as, or closely allied to, that of the Tenimber or
Timorlant group.
1. Megapodius dupeiTeyii Less. & Garn.
Cf. Not-. Zu,A. 10O4. p. 170.
3 S ad., Tejia, 27. viii., 5, 0. ix., I'JOo (Nos. 6788, 6849, 6886).
2. Turnix maculosus (Temm.).
Cf. Noe. Z.,ol. 11104. p. 17(i.
1 ? ad., Tepa, 2U. x. I'.jn.j (No. 7005).
3. Ptilinopus cincta ottonis Hart.
Ptilinopus cincta otloiits Hartcrt, Nuc. Zuul. VM\, pp. ITS, 179 (DammtT ami Babber. Terra typica :
Dammer).
A fine series was obtained near Tejia, on Babber Island, in August and
September IVii.io (Nos. UGU5, (3Gi)U, (1(122, 6024, (1628, 0040, OO'JO, 0700, 6703, 0755,
6760, OSIO). "Iris burnt sienna (brick-red in most specimens). Feet dark red
(chocolate-brown, purplish red, dark purple). Bill : tip yellowish, base dark
(olivaceous).
4. Ptiliuopus xanthogaster xanthogaster Wagl.
Cf. y<n: Zml. I'J04. p. 17'J. no. 8.
Eleven examples killed at Tepa in August and September (Nos. GOoO, 0015,
(J019, 0020, (j030, 0631, 0053, (1737, 0753, OtSOl, OSOIa).
It is interesting to find that P. :r. xanthogaster, the same as found on Banda,
Key, Dammer, Taam, Teour, Manggoer, Koer and the Timorlaut (Tenimber) group,
occurs on Babber, and not L', x. roscipileum {^ov. Zool. 1004. p. 170^, which inhabits
Roma, Moa, Kisser, Letti and Wetter.
( 289 )
o. Ptilinopus wallacei Gray.
Ptiliiinpus wallacei Gray, P. Z. S. 1858. pp. 185, 195. pi. 136 (Aru Lsland.s).
We have no sjiecimeu from Aru, but a large series from the Key and other
islands. Kiihu's men obtained fifteen siiecimens in Angnst, ISejitember, October
(Nos. 0603, 6G54, 0602, 0734, OToS, 0781, 0830, 0843, 0887, 0'JU4, O'JU.j, 6907,6942,
0963, 0998).
0. Carpophaga rosacea (Temm.).
Cf. Nuv.Zuol. i;i04. p. 1811.
Common near Tepa in Angnst (Ncs. 6604, 0636, 6657, 6658, 6692, 6752).
7. Carpophaga coiiciuua conciiina \Vall.
Cf. Xov.Ziiol. 1904. p. 181.
Very numerous near Tepa in Angn.st and September (Nos. 0008, 0018, 0(520,
0047, 6698, 6699, 0741, 0751, 6812, 0838, 0847, 6848, 6852, 6944).
8. Turtur tigrina (Temm. & Kuip).
Cf. Nnc. Znol. 19(14. p. 18i
This ubiquitous Turtledove was common on Babber Island (Nos. 0744, 0702,
6817, 0823, 0915, 0918, 0921, 0923, 0927, 0928, 0929, 0935).
9. Geopelia maugeus (Temm. & Knip).
Cf. Nov. Zuul. 1904. p. 182.
Seven specimens were sent (Nos. 0004, 0037, 6675, 6894, 6948, 6952, 6955).
111. Chalcophaps chrysochloi-a (Wagl.).
Cf. N,n\ Zuul. 1904. p. 183.
A series of specimens from Babber are somewhat intermediate between
C. c. chnjsochlofu and C c. timorcii>>is, but few have the hiudncck so continuously
grey as in our typical iimore/isis. The bill, however, is rather lougish, which is
more or less characteristic for tlmoreitsis, and the extent of white on the shoulders
varies somewhat. Mr. Kiihn sent fifteen specimens, of which, however, very few
are adult (Nos. 0017, 0035, 0049, 6659, 6681, 0740, 0794, 6834, 6837, 0839, 6860,
6855, 0861, 6U14, 0914 a).
11. Sterna bergii pelecanoides King.
Of. Nov. Zoul. 1904. p. 185.
One skin, August 1905 (No. 6078).
I have not heretofore tried to distinguish between the various races of
Sterna hergii, as our material was not sufficient for a comjilete study of them ;
but Mr. Rothschild, inspired by Mr. Bangs' article in Bull. Mas. Compar. Zool.
Harvard Coll. xxxvi. 1901. \). 257, suggested to me that there were probably as
many races of Sterna hergii as of other widespread birds, and I find this really to
be the case, although these races are closely allied and not recognisable by a
confirmed lumper, who has not trained his eye for careful comparison. Probably
the name for the Sterna hergii form from all these islands is to be S. b. pelecanoides
King {Surv. Int. Austr. ii. p. 422, 1820).
( -MO )
\~. Cliai'adrius squatarola (L.)-
Cf. -V.--. z„ui. i'jo4. p. iKi;.
? ud. Tqiii, I -J. viii. IDUo (No. 0607).
i-'i. Ochthodromus geofFroyi (Wagl.).
Of. A'of. Zuol. 1004. p. 186.
Three females, shot in September (Nos. G9S0, d'Jb'J, GD'.Jl).
14. Numenius phaeopus variegatus (Seup.).
Cf. .V"C. ZjuI. VM-i. p. 18G.
Si.x sijecimens (Nos. 0'.)['2, (j'.tl'.i, (i',i-^(;, GOT."), G9«;3, OU^:i .\).
lo. Limosa limosa melanuroides GtiiiKl.
IScolojiaj; Liiiwsa Linnaeus, %»(. Xnt. ed. -\. p. 147 (1758— ''Habitat iu Eiiropa." Tui-ra typica ;
Sweden, e.\ Fauna Suecica 144), Linnaeus' only quotation.]
Liintisu riiehiniiruides Gould, P.Z.S. 184IJ. p. 84 (Port E.ssington, Australia).
Tlie eastern Black-tailed Godwits are constantly ver)' much smaller (wing one
to two inches shorter, bill shorter and not so thick at base) than the European
I'orm, so that they must be separated subspecitically. Mr. Kiihu sent two luoultini;'
specimens, a 6 and a ?, shot on September Vii\i and ^Uth near Tejui (Nos. 01)21 1,
0'JS:i).
10. Totanus hypoleucos (L.).
Triiiijii. HupiiUiinis Liunauus, Sijxt. Nat. ud. X. p. 14U (1768— " Habitat iu Eui-opa." Tuna typica :
Sweden ; from first quotation).
<Jf. Xuc. Ziiul. 1004. p. 187.
Three sjiecimeus, August and September (Nos. 605>'.>, 0703, G'JTU).
IT. Glottis nebularius (Gunu.).
Sculii^iuj: ncbfilarliis Guuucrus, Lctitt, Lapi). IJv-^kf. 17li7. p. '261 (Lapplaiid).
Six specimens, August and September (Nos. GTU'J, GTOo, GSlo, &M2, 0'JU3,
OIHIO).
18. Heteractitis brevipes (Vieill.).
Cf. .V."-. Z„„l. I'.iOl. p. 187.
) 6, Tepa, I. i.\. I'.Mi.j (No. 06^!:.'),
I'.i. Terekia cinerea (Giild.).
L'f. .V.,1-. Zwd. 1U04. p. 187.
1 d, Tepa, 1:.'. ix. l',)n.5 (No. O'Jl Uj.
2<i. Limonites ruficollis (Pall.).
Cf. .Xni: Zuol. I'JUl. p. 187.
Si.\ specimens, August and Sejitember (Nos. 070.5, O'.IUS, GU13, O'JlT, O'.l.jG,
O'.I'JT).
'~;i. Heteropygia acuminatus (Horsf.).
Cf. Nor. Z'jol. 1904. p. 187.
1 ?, Tepa, 12. i.\. I'Jn.j (No. 0010).
( 291 )
~'2. Tringa crassirostris Temm. & Schl.
Of. iVor. Z.ml. 1004. p. 187.
1 c?, Tepn, 12. viii. 10115. "Iris dark brown; feet dark slate; bill black."
(No. 60(18.)
23. Stiltia isabella (Vieill.).
Cf. Nor. Z.,,,1. 100.5. p. 202.
Wc did not reooivo sjiecimens from the other Sonth-Wost Islands. Six
specimens were sent from Tcpa, on Babber (Nos. 6038, 6C4S, 0832, 08'.)."), 0047,
6958).
24. Lobivanellus miles (Bodd.).
Cf. Nor. Zool. 1905. p. 200.
This species too was not received from the other South-West Islands. One
female was shot at Tepa ou September 5th (No. 0851).
25. Esacus magnirostris (Vieill.).
Cf. Nor.ZanJ. 1004. p. 187.
1 c? iuL, Tepa, 10. viii. 1005 (No. 6668).
20. Ibis molucca Cnv.
Cf. Nor. Zoo}. 1005. p. 20.3.
1 $ jnv., 1 ? ad., Tepa, Anp:iist and September (Nos. 6677, 6923). No ibis
was obtained bv Mr. Kiilm on the other Sontli-West Islands.
27. Platalea regia Gnnld.
(M. Nor. Zool. 1004. p. 188.
1 (?, Tepa, 11. ix. 1005 (No. 6909).
28. Demiegretta sacra (Gm.).
Cf. Nor. Zool. 1004. p. 188.
Two grey examples and one white one, Anfjiist 10ii5 (Xos. 0759, 0701, Os74).
29. Notophoyx uovaehollandiae (Lath.).
Cf. .V'"-. Zoul. 1004, p. 188.
1 ? ad., Tepa, 24. viii. 1905 (No. 6742).
30. Nycticorax caledonica (Gm.).
Cf. Nor. Zool. 1004. p. 188.
2 c?c?, 1 ?, September 1905 (Nos. 0821, 6829, 6833).
31. Plotus novaehollaudiae Gould.
riolus mirae Jiolhuuloir Could, P. Z. N. 1847. p. .34 (S. An.slrali:i).
(??, 17. viii., 1. ix. 1905 (Nos. 0001, 0818).
( 292 )
32. Pelecanus conspicillatus Temm.
Pelecamm eonsplrilhiliis Temminck, PI. C"l. v. T;if. JTr, (Ifii'l : Au<tralia).
1 ad., Tejia, August lUO:) (No. dsO.")). We received no pelican from the other
South-West Islands.
33. Haliastur Indus intermedius (!urn.
Cf. /V<.)'. Znol. 1904. p. 180.
3 ad., 2 jnv., Tepa, Aui;iist and September (Xos. C.Ciio, (IGiiO, CGIM, OOSO, C0S4).
Tlie ailuil birds are typical intermedhis, having blaok slial'ts to the white featiiers of
tlie underside.
34. Astur albiventris polionotus (Salvad.).
Urnspiziax polionntiis Salvadori, Agyiuntc Oni. Papiinsia i. p. 19 (IH.S'.I : Timorlaut).
1 c? ad., 1 ? ad., 1 ? jnv., Teiia, August and September (Nos. 0041, 0899,
0957). " Iris, S ad. orange, ? ad. brown ; feet sulphureous, bill black."
The adult female agrees perfect!}- with our specimens from Tenimber, Banda,
and Dammer, while the male differs from our single male (from Banda) in having
only the jugular region vinous, the lower breast and alxlomen as well as the thighs
white, with barely a tinge of vinous and an indication of bars on the sides of the
abdomen. Whether this is an individual character, or peculiar to all Bal)bor
oxamjdes, cannot be guessed without more material. Possibly both nlbiri'iitris
from the Key Islands and Manggoer and polioiiotim may eventually be looke<l upon
as subs]iecies of A. torqtiatiis, but it reijnires more time than I can at present ad'ord
to study the relations of these forms at full length.
35. Tinnunculus moluccensis occidentalis Mey. & Wigl.
Cf. Nop. Znol. 1904. p. 190.
The Kestrel of the South-'Westeni Islands is 7'. m. ocridrntalis, and the
specimens, four in all, sent from Babber agree with those from Kisser, Letti, and
Moa, showing no approach to the darker T. m. moluccensis (Nos. 0032, 0819, 0827,
0911).
30. Tinnunculus cenchi'oides (Vig. k Horsf.).
Faho cem-hrohUx Vigors & IIorsBeld, 7V<(»s. Linn. Sue. Lond. xv. p. 181? (1821; ; Australia).
We did not receive this species from the other South- West I.slands. Kiiim's
bnnters obtained 1 c?, 2 ? ? at Tepa, 24, 29. viii., and 1. ix. 1905 (Nos. 6735,
0790, 08211). " Iris brown ; feet snI|ihureous ; bill slate with black tip."
37. Palco peregrinus ernesti Sharps.
FaUo ernesli Sharpe, Ihin 1894. p. (>ij (Borneo).
1 ? ad., Babber, 25. ix. 1005. " Iris brown ; feet chromcons ; bill slate-colour."
(No. 0953.)
Cf. Nor. Zool. 1904. p. 100, where I recorded a young female from Wetter as
F. jiereijiimis melanogenys'. I am now convinced that this is rather F. /^. c/v/^'.s//
than /'. p. melanogenys.
The Falcon from Babber is a very interesting specimen. It agrees with (he
specimens of F. ernesti from Borneo, N.W. Luzon, New Guinea and Mallikolh;
( 293 )
in having groy (not, ochracoons) under tail-coverts, with wide slaty-blackish l)ars,
very grey (not ochraoeons) flanks, with very wide bars, though the breast is not
quite as grey as in the males of eniesfi. The adult female in the British Museum
and the one iu Triog (from New Guinea) are still darker grey on the flanks and
more cinnanion-rufons on the ehost, than our Babber specimen, bnt in all essential
characters they agree.
38. Ninox boobook cinnamomina subspec. nov.
Formis Xinn.r hoobnnl; honhunk et Xim.r hoobn,,/: occUnln appellatis similia sed multo minor et
colore laete cinnamomino an rufo-cinnamomeo, neo grisescente, nee brunneo, nee rufo.
The four specimens from Babber, obtained at Tepa in August and September,
are almost alike, and their colour above is bright cinnamon, lighter on the neck,
darker and more brownish on the crown. The scapulars and greater upper wing-
coverts are rufous-cinnamon, with white or whitish bars and round spots. The tail
is rnfoas-ciunamon, with more or less ill-defined, sometimes, however, rather distinct
brown bars to the inner webs, which are generally much paler. The anricular
region is covered with a large, very conspicuous slaty-brown patch. The under-
surface varies from light reddish cinnamon with darker longitudinal patches and
whitish spots on the abdomen to whitish with rufous-cinnamon longitudinal
patches. Wing 209 — 213, tail about 120 — 130 mm. "Iris chromeous or whitish
yellow ; feet whitish ; bill dull black or slate with yellow or whitish tip." Type of
^Ino.c boohook ciitnamomina : S ad., Tepa, 22. viii. 1005 (No. 0000).
It is particularly interesting to find on Babber an apparently very constant
distinct form of the booboo/i-gmw]) of owls, as the birds from Moa, Letti and Roma
are much nearer to N. boobook boohook and X. boobook occllata. C!f. Nor. Zool.
1904. p. 191, and 1905. p. 210. It must be said that Australian specimens vary
enormously in coloration and size. Some small specimens from Derby and other
places in N.W. Australia are hardly larger than my cinnamomina, and some are
quite pale cinnamon, but not bright rnfous-cinnamon. I have, however, not seen
any scries of specimens from one place that are alike inter se (we received five
examples from Babber), that are so bright rufous-cinnamon, or so small. It is,
therefore, not only advisable but necessary to name the Babber form. We received
five skins, f.mr marked as males, one doubtful (Nos. 0006, 0007, UUll, 0808, 6892).
39. Eos reticulatus (S. Midi.)
PsillKi'iis rrliridatun S. Miiller, Yrrh. Lriinl- cii Vulhcnhumlc, pp. 107, 108 (Moluccas, no e.xact
locality).
Not known from the more westerly islands of the group, but already obtained
on Babber by Schildler (cf. Finsch, Notes Lei/den. Museum xxii. p. 285), and on
Damnier by Heinrich Kiihn. On Babber it is evidently quite common, as Kiihu's
collectors sent thirteen specimens obtained near Te]ia in August and .September
(Nos. 0612, 0C44, GG46, 0650, 6052, 6656, 0065, 6743, 6748, 6807, 6809, 6810, 6872).
" Iris burnt sienna (lirick-red) ; bill red ; feet blackish."
4ii. Trichoglossus euteles ( Temni.).
Cf. Nnr. Zaul. 1904. p. I'.i:).
Mr. Kiibn sent four specimens from Tepa (Nos. 6615, 6622, 6625, 6789),
Schiidler sent to Ticyden from Babber,
( 294 )
41. Tanygnathus megalorhynchos subaffinis i>d.
r„n,/g,„illu,s siibaffinh Sclatcr, I'ro,: Z.,,,1. So.: /..„„/..», IsS:;. pp. .".l, r,3, 194, 200 (Timorlaut) ;
Finsch, Notee Leijden Mm. xxii. p. 200 (Babbei).
Mr. Kiihn sent seven specimens shot in August and September at Tepa.
These birds appear to be all adnlt males in perfect iilnmage, with the exception of
one (No. 684S) which is an immature male. This last one has the lesser npper
wing-coverts dark green, not blue, and the outer edges to the median and greater
wing-coverts are narrow and dull lemon-3'ellow. All the others have the lesser
wing-coverts, bend of wing and tips of scapulars and median and greater wing-
coverts bright blue, the borders to the wing-coverts broad and of a very bright
yellow. They surpass thus in brightness of the wing onr series of T. m. suhaflims
from Tenimber (Timorlaut) ; but as only a few of the latter are adult males in fresh
jilnmage, and one closely approaches our Babber specimens, I believe that the
differences are due to the different season and age of the birds, Imt not peculiar fo
the localities. " Iris chromeous ; bill scarlet ; feet slaty l)lack." (Nos. OOol, (iTTn,
6806, 6840, 6846, 6945, 6946.)
The home of T. m. sulxifUnis is otherwise Timorlaut (Tenimber). It does not
occur on Moa, Wetter, Letti, or Kisser.
42. Alcyone azurea yamdenae Rothsch.
Cf. A''"!'. Zor,J. 1904. p. 19G.
"What I said {I.e.) about two .specimens from Roma applies exactly to two
examples from Tepa, Babber Island. They may be said to agree with the type of
A. II. ynimlenac, except that their wings are slightly longer, measuring 73 and
7.'>-5 mm., while the wing of the type of A. a. yawdnmr measures about 7(l-.5 mm.
It is very probable that the type of ijiinuhniiif is an exceptionally small specimen of
the same subspecies, but more material from Tcninilicr is necessary fo dec^ide this
question finally.
43. Halcyon australasia dammeriana ?
Cf. iV"i-. Z„o}. 1904. pp. lOli, 197.
The specimens from Baljber are rather puzzling. They are as dark cinnamon
below and on the crown and liindneck as //. australasia (Inmmcriaiui. From the
twelve specimens of //. a. (hnnmeriana before me they differ as follows : the
crown, instead of having a fairly large, rarely small, bluish green patch, has
generally a rather smaller jiatcli, and four examples have no patch whiifever,
but have the crown uniform dark cinnamon ; flic wing is generally som(^\vhat
shorter, varying from 75-5 to 79r), instead of 78 to 84 mm. in //. a. ilnmmeriami.
The question therefore is: are our series sufficiently large to prove an average
of size or more or less largely patched crown in one of these forms or not ? I do
not wish, at present, to decide, but probably the Babber form deserves a name.
Babber lies between the Letti-Moa group and Tenimber, and we slionld therefore
expect either the Letti-Moa form (//. a. interposita) or the Tenimber form
(//. a. mi/ior). But witli neither of these do our Babber birds agree ; they are
darker underneath (and jirobably smaller) than //. it. hi/rrpo.s/fn, and they are
ninch darker on the throat and in the middle of the abdomen and somewhat larger
ihan //. a. minor. That the Dammer form shnnld inhabit Babber as well is very
strange— but nothing is impossilile.
( 295)
Jlr. Kiiliu's collectors sent, twelve si)eciiiieiis killed near Tejia in Angnst and
.Sei)tember, of wliicli six are marked as males, six as females — whether correctly
or not I cannot say. " Iris dark brown ; bill black and white ; feet blackish."
(Nos. (J(;i2, 6010, (ji;iG!i, GOTU, 0696, 6880, 0881, 0882, 6894, 6040, 0004, 0071'.)
44. Halcyon sancta Vig. & Horsf.
Cf. iV-m. Z'liil. 1004. p. 19G.
1 ?,!>]«, 20. ix. lOOo (No. 0087).
4;"). Halcyon cliloris chloris rBodd.).
Cf. Nnv. Zunh 1904. pp. 197, I'.iS, 19<.i.
Thirteen specimens from Tepa (Nos. 6024, C033, 6034, 6683, 6684, 6603, 6700,
6776, 6786, 6798, 6890, 6898, 6959).
40. Misocalius palliolatus (Lath.).
Cf. Nnr. Znnl. UM14. p. 201.
1 ?, 0. ix. 1005 (No. 0801).
47. Cuculus variegatus Vieill.
Cf. N,ir. Zonl. 1005. p. 217.
1 ?, Tepa, 4. ix. 1905 (No. 6845).
48. Chrysococcyx spec, an snbsp. nov. ?
1 cJ ad., Tepa, Babber Island, 15. ix. 1905. " Iris bnrnt sienna ; bill and feet
black." (No. 6939.)
i ad. Upper surface dark green with metallic gloss, on the crown a bine
tinge. Some of the median upper wing-coverts have white terminal edges and
spots, so that an irregular white patch or ring is formed, showing an ajiproach
to the Key-Islands form, C. crmsii-oxtris, which has a large white patch on the
wing-coverts. Tail without any rufous colour ; only the outermost pair with white
crossbars, the next with only a ronnd white spot at the tip of the inner web,
central pair withont white. Undersnrface white with somewhat narrow greenish
bronze crossbars, which are few and not very consi)icuons on the throat, the
feathers there having only one bar instead of two.
Dr. Finsch describes a similar bird from ITalmahera, whence Dr. Vdrdcrnian
described C. nieuwcnhiiisi. Unless the latter is a somewhat j-onnger bird it is not
the same as Finsch's ! It is quite possible, or rather evident, that a sjiecial form
inhabits the Moluccas, but whether that extends from Halmahera to Aniboina or
not is doubtful, and whether the Babber bird is the same or not can only be jiroved
by more material.
40. Scythrops novaehoUandiae Lath.
Cf. .V,,,-. Zool lOOo. p. 219.
A young bird (No. 6709) just from the nest, unable to fly, shot at Tepa 20. viii.
10(15, proving that this species is not an accidental visitor, but is hatched on Babber
Island. These yonng Scj/tlirojts arc peculiar and pretty things. Their head and
neck is rusly buff, the feathers of the ui)perside with ashy-grey l)ases ; ear-coverts
( 290 )
light grey ; feathers ol' back and upper wing-coverts slaty gre\- with large buffisli
rnst-eolonred tips. Rectrices as in the adnlt birds, as far as visible ; breast and
abdomen white with butf)- yellow tinge and narrow dark grey crossbars which
are very wide apart. " Iris dark brown with a whitish ring ; feet slate ; bill dull
brownish."
50. Eurostopus argus Hart.
Cf. .Y'.i'. Z;nl. 1904. 11. 202.
Seven specimens (Nos. fiS56, 0859, 6802, 0803, 6870, 0S70, GSTT).
51. Caprimulgus macrourus Ilorsf.
Gqirimnhjiix macrourus Horsfield, Trniix. Linn. S'ic. .\iii. p. 14'2 (1821 — Java).
? immat., Tepa, 28. viii. 1005 (No. 0791).
This bird certainly agrees better with C 7»acroun/s than with C. maeronrus
cp/eii'iisis, which was received from Wetter (Nof. Zool. 10ri4. p. 2ii2). I come,
however, to the conclusion that C. macrourus, manilensis and celebensis arc all
three only subspecies, and should therefore be called Capr. macrourus macrourus,
C. m. mariileHsis and C. m. cclebi'iisis, while on ('eylon we have C. m. atripcrinis,
farther nortli C m. amhiguus, C. m. alhonotatus, and C. m. nipalcnsis.
52. Pitta vigors! (ionld.
VWhi r/fim-si Gould, B. Aiislralio, iv. pi. 2. (part II.) (1.S41 — Believed to come somewhere from
Australia ! Errore. As the terra typica we may accept Banda, the first known exact
locality).
1 ?, Tepa, Babber, 21. viii. 1005 (No. 0005).
53. Hirundo javauica Sparrm.
Cf. m^r. Zoo!. 1004. p. 20.'?.
1 cJ, 3 ? ?, Tepa, 24, 20. viii. 1005 (Nos. 0738, 0700, 0708, 0773).
54. Petrochelidou nigricans (Vieill.).
Cf. A'"!-. Zool. V.m. p. 204.
2 cJcJ, Tepa, 27. viii. 1005 (Nos. 0778, 0770).
55. Muscicapula hyperythra audacis subsp. nov.
rT. Similar to ^1/". Inipcrythra lii/pcri/thra in general appearance, but larger,
especially the bill much longer, the metatarsus longer, wing longer. The inner
edges to the qnills are lighter, more whitish, the under wing-coverts have more
buflfy-white colour. Wing 03-7— 65-4, tail about 42 — 45, cnlmen 14 — 14-5,
metatarsus about 20 mm. (Type <? ad. No. 0804, Tepa, G. ix. 1905). " Iris
brown, blackish, or wliitish yellow." This last statement, however, is probably
erroneous. " Feet pale plumbeous, dull black, blackish ; bill black."
5 c?(?, Tepa, Babber, August, Seiitcmber 10ii5 (Nos. 0780, 0850, 0854, 0804,
G98.5).
1 " ?," Tei)a, 0. ix. 1905 (No. OSGO). Dilfers from those of M. h. In/pcri/thra
in having the back and scapulars distinctly greyish, not olive like the head, aud
the breast buft'y rufous, the middle of the abdomen and the under tail-coverfs
( 207 )
wLife, with onl;- a faint ting-e of bnfF ; lint iioKsibl3- this supposed ? may be a
young S , in which case the valne of these ditferences is of course doubtful.
Note. — Dr. Fiiisch {Notes Lpi/rlen Mhs. xxii. pp. 201, 202) places my Iiummcriu
henrici (of. Nov. Zool. 1900. p. 14) from Dammer in the genus I'occilodn/as.
I cannot agree with this decision. I readily believe that it is desirable to diminish
the genera of Miiscicapidae very considerably ; but if this is done, and IJammeria
should be suppressed, it would far better be united with Muscicapula than with
Poecilodnjas, a genus which inhabits Australia and the Papuan Islands, but nut
the Sonth-West Islands. Moreover, the structure is different, the tail oi Poecilodiyas
being softer and shorter, the bill weaker. The sexes of Pammcrin differ considerably,
as iu Muscicaimld.
56. Gerygone dorsalis fulvescens A. B. M.
Gerygnne fidvcaceuH A. B. Meyer, hix 1884. pp. 2, 27 (Babber).
The Gerijqonc from Babber has been separated by Meyer, I. c. Undoubtedly
all these forms — i.e. kiihniivom Dammer (which is cjnite different from both dorsalis
and fulvescens), fukesccns from Babber, sequens from Roma, and others — are
representative forms, and Dr. Finsch's surmise, from the comparisons of spirit-
specimens, that " Iriihni " as well as fideescei/s are found on Baliber, is erroneous.
Dr. Finsch's /dssereitsis (cf. Wotes Leijden ^^lls. xx. p. 133, Nov. Zool. 1904. p. 2(J.i)
is merely a synonym of fulvescens — at least I cannot separate our specimens from
Kisser, Moa, Letti and Babber. Mr. Kuhn sent six specimens from Babber.
3 S ad., 1 c?, 1 ? aet. med., 1 ? juv., Tepa, September 190.J (Nos. 0s41, 0844,
6853,0875,6909,6976). "Iris burnt-sienna brown; bill black in adults, brown
in young ; feet slate-black."
57. Ehipidura fiiscorufa Scl.
Rhipkhtra fuscoriifa Sclater, Pi-oc. Zonl. Sue. Lnmhm, 1883. p. 197 (Tenimber).
10 c?? ad., Tepa, August and September 19()5 (Nos. 0088, 0701, 0820, 0873,
0883, 6937, 6955, 6902, 6968, 6971). " Iris blackish ; bill black ; feet durk
brown or blackish."
Dr. Finsch has already made known the lact, that the Babber form is the same
as the one from Tenimber (Timorlant).
58. Rhipidura elegantula reichenowi Finsch.
Hhipidura Reich etimri Finsch, Notes Lei/Ji n Museum sxii. p. 257 (Babber).
(Cf. Nm\ Zool. 1',I04. p. 20l"i, sub nomine Rh. elcgaiilnhi.)
1 c? ad., Babber, 28. ix. 1905 (No. 6981).
Dr. Finsch separated li/i. reicheyioici from eleeiantuki on account of its rusty-
cinnamon, instead of white, forehead. I ventured {I. c.) to unite reichenowi with
elegantula, because among our twenty-nine examples from Moa, Letti, Roma and
Dammer I fonnd two sjiecimens which agreed with Dr. Finsch's tyjie of veichcnowi.
Perhaps my conclusion has been too hasty, because the specimen received from
Kuhn has also a cinnamon forehead. If all specimens from Babber have a
rusty-cinnamon forehead, the Babber form would bo a subspecies, iu spite of the
occurrence of single indiviiluals on other islands which are similar in colour.
Only more skins from Babber can decide whether reichenowi is really separable
from elegantula or not. I have termed this form 1th. elegantula reichenoivi, but
this nomenclature may not be final : thej-e is little doubt that other forms are
( 298 )
also subsjiecies of this same species, Imt 1 cannot work this (inestion out iinally
at this moment, and it serves m_v pnrposc at present if 1 show in my nomenclature
that " lih. rriclii'itoiri " is a subspecies of ctrgantidu, if separable at all.
59. Pratincola caprata caprata (L.)-
Cf. A'"!'. Znol. 19U4. p. 209.
1 c?, 2 ? ?, Tepa, Angnst and September 1905 (Nos. OSoo, r,951, Oi)<):5).
On. Lalage timoriensis (S. Mull.).
Cf. X„r. Zoo/. 1904. p. -208.
1 c?, 1 ?, Tepa, Angnst and September 1905 (Nos. 077;"), 0938). Tlie males
vary xvifli regard to presence or absence of a white superciliary line. In most cases
this is indicated, bnt sometimes it is well developed, sometimes absent.
01. Graucalus melanops (Lath.).
Cf. Nor. Zor.l. 1904. p. 208.
Twelve specimens from Tepa, Angnst and Seijtember 190.") (Nos. 00"23, G71ti,
G797, 0811, 0813, 6824, 68.31, 68!):, 0869, 6944, 6955, 6960).
62. Heteranax mundus (Scl.).
irmmrclm nmmhis Sclater, Prnr. Zonl. Soc. Lnndim 188.^. p. 54. pi. xii. fig. 2 (Tinrorliiut or
Tenimber).
Heffrnim.r mfiii(hi>! Hartert, iVoi'. Zihil. 1900. p. 14 (Dammor) ; Finsch, .V'/frs Let/ilni .l/»s. xxii.
p. 2i'i7 (B.ibbcr, Dammer).
4 c? ad., 2 t? .jnv., Te]ia, Angnst and Se]iteniber lOii.'i (Nos. rj(i64, 0674,
0803,0930, 6!)33, 0041).
Dr. Finsch (l.r.) declares that this interesting binl is not; a Flycatcher, bnt
belongs to the Laniiehie or Prior>opi(hic of Sliar[ie. This conception is quite
erroneous. Dr. Finsch .apparently based it only on a superficial comparison of the
strange bill ; but had he t.iken into consideration the plumage, style of coloration,
strong rictal bristles, etc., combined, he would agree that Ifetemmtx is a Flycatcher,
and that it is best placed in the neighbourhood of Monarrha.
03. Geocichla peronii audacis Hart.
Cf. Nur. Znol. 1004. p. 2llS.
Nine adult birds from Tej.a (Nos. 0010, 0782, 07S3, 0787, 0787a, 0.<«!3, 0900,
09.")4, O'.iOl).
04. Cisticola exilis (Vig. \: TTorsf ).
Cf. Nm: Z,»,\. 1904. p. 210.
2 $S, 1 ?, Tejia, Angnst and September 1905 (Nos. 0610, 0835, 6932).
05. Orthnocichla subulata advena subsp. nov.
Rubspcciei 0. mihulnla xuhujala dictae .similis, sed m.njnr, onloro supra rufcRcentiore, rostrn
nigricantiore baud difficile distinguenda.
Differs from < >. mdmlaUt subulata from Timor by its smaller size, more rnfes-
cent colour of the ujiporside, and darker, blackish, not light brown, ujiper bill.
Wing, cJ 58, ? 57; tail, cj 27-5, ? 20; culraen, 10; metatarsus, 21-0 ram,
Type No. 0858, Tepa, 0. i.x. 1005, S ad.
( 299 )
Mr. Kilhn seut only a jjair, t? ? , Tepa, 6, 7. ix. lOOo. "Iris bluckish : fuet
colourless ; bill black, pale below." (Nos. C858, 0871.)
The occurrence of an Urt/iHocickla so far eastwards is of extreme interest, and
we may imagine from this fact, tluit all the islands between Babber and Timor have
forms of Oi-thuociclda as well, either our adceiia, or undescribed ones. Probably
new forms are still to be discovered on Roma and Dammer.
The wings of onr 0. s. ■■ntbalata measure in the S uo — 56'5, ? .52 — 53-3 mm.
60. Pachycephala melanura tepa subsp. nov.
This most interesting new form of l'<MliijccpImla is aiijiarently nearest to
and stands between those from Dammer and Wetter. These forms may be
distinguished as follows :
Pachycephala melanura dammeriana Hart.
Dammer. — Edges to remiges grey with an olive-green tinge. Tail black with
greenish grey-brown tip and greenish outer margins towards the base ; bill slender
and strongly hooked.
Pachi/cephala melanura tepa subsp. nov.
Babbcr. — Edges to remiges greenish. Tail olive-green, with a blackish shaft-
line and black anteajucal area of variable extent, and an olive-green tip ; bill stouter,
and, as a rule, not so strongly hooked.
Pachi/ccphala melanura arthuri subsp. nov.
Wetter. — Light grey margins to the distal halves of primaries, broad greenish
ones to the secondaries, entirely green tail, bill elongated. Differs from P. m.
calliope by having a longer bill, cnlmen 21'7— 233 mm. (20 — 21 in calliope), and in
being more yellowish above, esi)ecially on the nape. Named after Arthur Goodson
of the Triug Museum.
Pachi/cephala melanura calliope Bp.
Timor. — Like P. m. arthuri, but the bill a little shorter, the u^jperside less
yellowish.
The female of P. m. tepa difl'ers also from those of its allies. The top of the
head, sides of head, ear-coverts and nape are greyish brown, contrasting with and
sharply separated from the olive-brown liack ; the rump and upper tail-coverts are
greenish ; tail greenish with blackish shafts ; throat white, separated from the
breast by a lavender-buflf chest-band ; abdomen saffron-yellow, sides with a brownish
wash. S : " Iris dark brown ; bill black ; feet dark plumbeous grey." ? : " Iris
dark brown ; bill Ijlack ; feet plumbeous or slate-grey."
Type of P. in. tepa : No. 0044a, c? Tepa, Babber Island, 15. ix. 19U6. — Type
of 7^ m. arthuri : No. 541)8, S, Wetter, 14. ix. 1002.
Wing of cJ P. m. tepa 91 — 94 mm. (12 specimens).
Wing of c? P. m. dammeriana, 90 — 94 mm. (5 specimens).
AVing of c? P. m. calliope 87 — 89 mm. (4 specimens).
Wing of c? P. m. melanura about 79 — 84 mm. (5 specimens).
Wing of i P. VI. arthuri about 88 — 93'5 mm. (5 specimens).
BIr. Kiihn sent twenty specimens, shot in August and (September 19U5 (Nos.
6621, 6642, 6644a, 6645, 0651, 6660, 6672, 6676, 6702, 6733, 6739, 6757, 6774,
0804, 6808, 0820, 6830, 0880, 0890, 0897).
( 300 )
UT. Pachycephala arctitorquis kebirensis Mey.
CtXoi:Z;ul. I'Ml. p. 2V2.
At last wo have received a series from Bablier, i.e. topotypical kchimixig.
They bear out what I wrote two years ago {I. c). The females have the crown
brownish or rnfons grey. Though occasionally examples of 1'. a. arctitonjuis
a])]iroach them, as a rnle the two forms are easily separable in the female sex.
Jlr. Klilin sent from ISabber :
4 ? ad., 1 S juv., o ? ¥, Te]ia, August and September TJiKJ (Nos. 6(i:.'0, 0(597,
6707, U771, 0777, 091^4, 0930, 0943).
68. Zosterops bassetti Sharpe.
Cf. Noc.Zuol. i;i04. p. 214.
(As 1 have said before, I cannot decide, without a complete study of the genus
ZoMerops, how many forms can bo named trinomial ly, and tliercl'ore employ
l)iuomials for them at present. As long as we distinguish minutely, our labour
will not be lost.)
The specimens from Babber and Luang a2)pear to be ijuite similar to each
other, and I cannot distinguish them from e.xamples from Dammer and Roma
(cf. Soc. Zool. 19U4. p. 214). The bills of the Babber and Luang speeiraens are
as large as in those from Roma and Dammer — some, if .anything, oven larger.
Z. letticims has a much smaller bill and is more white underneath, and Z. citrinella
of Timor is still smaller, with a much smaller bill. Mr. Kiihn sent twelve
Z. bassetti from Babber (Nos. 0085, 0080, 0740, 0747, 6792, 0842, 0S07, 0901, 090o,
0907, 0979, 0990).
69. Dicaeum mackloti salvadorii A. B. Bleyer.
Dicaeum salvadorii k.'B.M.eyev, Ahh. his. 1S84. pp. 7. ^8 (Babber); of. Fiusch, Xutcx Lrijihni
Museum xxii. p. 274.
In Nov. Zool. 1904. p. 214 1 enumerated specimens from Roma and Jloa as
Dicaeum inackloti snbsp. nov. ?, saying that they were somewhat apparently
intermediate between D. m. mackloti and I), m. salvadorii, of which I had only a
single male in jioor condition from Dammer, and none from the " terra tyjiica "
(Babber), and that the black band surrounding the rod throat was " nearly always
wider in the Roma birds." Now we have received topotypical specimens from
Babber, and I find that I cannot separate the specimens from Moa from those of
Bablier, while those from Roma have the red of the throat darker (as dark as in the
"tyj)ical" mackloti from Timor), and surrounded by a wide black liand, in adnlt
birds ; while in U. m. salvadorii this baud is absent or indicated, but never wide
and well developed.
I distingaish, therefore, the following forms :
Dii-aciim mackloti mackloti Mlill.
[Smaller, c^ ad., wing about o4'o to oO'o, abdomen white with a creamy tinge,
dark red throat-patch surrounded by a broad black line : Timor and .Savu.
Dicaeum mackloti romac subsp. nov.
Larger, c? iwl., wing about 58 — 59-5, abdomen yellowisii crcam-colonr, dark
rod throat-jiatcli surrounded by a broad black line : Roma. Type No. 5399, Roma,
10. viii. 1902, H. Kiihn coll.
( 301 )
Dicaeum mackloti salcadorii A. B. Me}'.
About as large as l>. m. romac, wing S ad. about 57 — 59, abdomeii yellowish
cream-colonr, the somewliat lighter, more fiery red throat-patch not surrounded by
a wide black line, but only by an indicated or narrow band, il' present at all. LJabber
and Moa Islands.
The one male we have from Dammer seems to belong to salvadorii. Perhaps
it does not occur there regularly. The specimen is not in very good condition.
Mr. Kiihu sent from Babbcr h S S and 7 ¥ ? (Nos. 6610, 0618, 6021, 6087,
6691, 0729, 0828, 686o, 0884, 6931, 6973, 0999). " c? ? Iris black, feet black,
bill black."
7i). Myzomela boiei aunabellae Scl.
\_Mijzomela hoki S. Mullur, \'i rli. LumU tn \'ulhiiliiiuli. p. 172 {lH'i'J-ii, Banda).]
Myzomela auiiabdlae Sclatcr, P.Z. S. 1883. p. 50 (Lulur, Tuiiiiubor).
M. hoki ^OiVv' differs from .1/. b. annnhellai: -a^ follows : It is larger, and the black
area limiting the red throat is much wider. Otherwise they agree in all essential
characters.
Mr. Kiihn sent a fine series from Babber (Nos. 0613, 6014, 6017, 60.55, 6003,
6667, 6670, 6671, 6082, 6728, 6731, 6732, 0730, 6749, 0750, 0977, 6980, 6990).
71. Stigmatops squamata Salvad. ('I'subsp.).
Cf. A'oi-. Z,jul. 1904. p. L'I5.
The wings of the Babber examples measure from 61 to 71 mm. Probably the
smaller specimens are females. About the somewhat doubtful -S'. s. salcadorii
and litibirensis, a name given to the Babber birds (A. B. Meyer, Zeitschr. Ges. Orii.
1884. p. 218), see A'of. Zool. 1904. p. 215, and Fiusch, AVtw Leijden Mtis. xxii.
p. 270. BIr. Kulm sent nine skins from Babber (Nos. 6027, 0785, 6799, 0874, 0902,
0974, 0988, 6995, 7U04).
72. Erythrura tricolor (Vieill.).
Cf. Nov. Zoul. 1904. p. 217.
c? ? ad. and juv., Tepa, August and September (Nos. 6013, 0714, 6730, 0793,
6798, 6825, 6966, 0994).
None of these specimens have any blue on the hindneck.
73. Munia quinticolor (Vieill.).
Luxla quinlicuhi- Vieillot, (Jls. Clwnt. pi. 5-1 (1805— Finsch Noles Leydcii Jliis. x.\ii. p. 277 (one
specimen from Baljber).
2 3 ad. (one worn and both monltiug), 1 c? med., 2 juv., Tepa, August and
September 1905 (Nos. 0772, 6878, 6888, 0949, 6950). The young bird seems to be
undescribed; it is in colour exactly like that of .17. /^a^^/r/a, being above cinnamon,
below yellowish buff. The wings are longer than in young .17. jxtllida. The adult
female dilfers only slightly from the adnlt male.
74. Munia punctulata nisoria (Temm.).
Cf. Noi: Zunl. 1904. p. 218.
Tepa, Babber, August, September and October 19U5, young birds in moult
(Nos. 07.54, 0879, 6992, 7000, 7001, 7002).
( 302 )
Tri. Artamus leucorhynchus (L.).
Cf. Xnf.Zw.l. 1001. p. lilil.
Tcjia, BiibbLT, Ally-list lUU.") (Nos. 0025, 0020, 0740, 0704, 0784, 0SU2, 002(5,
G978).
70. Corvus latirostris ? subsp.
Corcus lalirtistris A. B. Meyer, Ztitschr. Gci. Oni- i. p. 1911 (1884 : Timoi-hiut).
Cf. Nuo. Zuol. I'JOl. pp. 173, 174.
Tim sjiecimcns from Bablier agree in all ossoiitial characters with Corciis
latlroiilris from Timorhiut (Teiiimber), but they are somewhat laru'er. They
measure : c? ad. wing SoU, S ail. wing 342, ¥ ail. wiug 318, ? ail. wing 310 mm.,
while specimens from Timorlaut measure S ad. wing 328, 6 ad. wing 317, ¥ ad.
wiug 322, ¥ jun. wing about 3U0 mm.
Possibly the Timorlaut specimens are not all correctly sexed. If we accei)t
that the alleged males are all adnlt males, the Babber form would be a larger,
and should be separated subspecifically, but I should like to see a better series
before doing this.
Mr. Kiihu's four e.xample.s from Tepa bear the Nos. 6611, 6023, 0027, 0004.
The iris is described as white, bill aud feet black. In Xoc. Zool. 19U1. jip. 173,
174, I have discussed this form, which has ignite erroneously been united with
C. macrorltijnchas.
ON THE BIRDS OE LUANG.
By Dii. ERNST IIAETERT.
WE had hoped that Mr. Kiihu's collectors, after their work on Baiiber, would
visit the large island of Hermatta, but unfortunately they chose the little
islet of Luaug, near Sermatta. They collected there 41 S2)ecies of birds, mostly of
little interest to iis. Undoubtedly Sermatta would contain many more species.
1. Syiiokm raalteni (S. Miill.) {Xoc. Zool 1904. p. 177). S 0. xi. 1905
(No. 7031).
2. PtilinojMS cinr.ta lettiensis Sch. {Xov. Zool. 1904. p. 179) (Nos. 7063, 7U00,
7072, 7076, 7093, 7096, 7097). It is interesting to find that P. c. lettieitsis and not
ottotiis is found on Luang.
3. Pttlinopus xanthogastcr xantkogaster Wagl. (cf. antca) (Nos. 7013, 7u26,
7038, 7051).
4. Carpopkaya rosacea (Temm.) (cf. antea) (Nos. 7032, 7092).
6. Geopelia maugeus (Temm. & Knip) (cf. antea) (Nos. 7028, 7030).
6. Geloc/telidonlnilotica (Gm.) (Nov. Zool. 1904. p. 199) (No. 7021). This bird,
marked S, is very young. It appears to be the European small-billed form,
and not macrotarsa. Adult birds should, however, be compared, to be certain
about this.
7. Sterna bcrgii pelecanoides King, (cf antea) (Nos. 7022, 7091).
( 303 )
8. Morimlla interpres (L.) {Nov. Zool. 1904. p. 186) (Nos. T045, 7040, 7047,
7049, 7052, 7101, 7103, 7104, 7105).
9. Charadrius dominicus fuhus (Gm.) {Noi\ ZooL 1904. p. 18(5) (No. 7045a).
10. Ochthodromus geoffroyi (Wagl.) (cf. antea) (Nos. 7080, 7088).
11. Numenius phaeopus variegatus (Scop.) (cf. antea) (Nos. 7027, 7068).
12. Numenius ctjanopus Vieill. (cf. Noc. Zool. 1905. p. 201) (Nos. 7048,7079).
13. Limosa limosa melanuroides Gould (cf. antea) (Nos. 7082, 7087, 7089).
14. Totanus glareola (Gm.) (cf. Nov. Zool. IU04. p. 187) (No. 7094).
15. Totanus hypoleucos (L.) (cf. antea) (Nos. 7006, 7007, 7011, 7025).
10. Limonites ruficolUs (Pall.) (cf antea) (No. 7090).
17. Glottis nebularius (Gunn.) (cf. antea) (Nos. 7083, 7084).
18. Heteropygia acuminutus CHorsf.) (cf. ««tof) (Nos. 7012, 7014, 7015, 7016,
7017, 7023).
19. Stiltia isahella (Vieill.) (cf. antea) (Nos. 7018, 7034. 7050, 7102).
20. Haematopus longirostris Vieill. (cf. Nov. Zool. 1905. p. 200), 1 " ? " ad.,
1 "c?"jiiv. (Nos. 7106, 7107). The bills of these examples are very long, that
of the adult bird (? ¥ ) measnriug 90 mm. from the end of the frontal feathering
to the tip.
21. Esacus ■magnirostris (Vieill.) (cf. antea) (No. 7009).
22. Demiegretta sacra (Gm.) (cf antea). Two white and three grey examples
(Nos. 7020, 7043, 7044, 7064, 7071).
23. Notophoyx novaehollandiae (Lath.) (cf antea) (Nos. 7010, 7032).
24. Garzetta nigripes (Temm.). Ardea nigripes Temminck, Man. d'Orn. 2ud
ed. iv. p. 376 (1840 : " I'Archipel des Indes ").
1 ? (?) Luang, 4. xi. 1905 (No. 7019).
25. Nycticorax caledonica (Gm.) (cf. antea) (Nos. 7024, 7029, 7069).
26. Fregata aqnilus (L.) — Pelecanus Aqiiilus Linnaeus, Sy.Ht. Nat. ed. x. p. 133
(1758— terra typica : Ascension Island). ? ad., 12. xi. 1905 (No. 7085). It is
interesting to find the large species and nut the small one (ariel) on these islands.
27. Phalacrocorax melanoleucos (Vieill.) {Soc. Zool. 1904. p. 189) (Nos. 7035,
7036).
28. Spizaetus limnaetus Jloris Hart. {Nov. Zool. 1904. p. 189). 1 c^ juv.
12. xi. 1905 (No. 7086).
29. Trichoglossus euteles (Temm.) (cf. antea) (No. 7095).
30. Eun/stomus orientalis austrulis Hw. (Nov. Zool. 19(J4. p. 195) (Nos. 7008,
7073).
31. Misocalius pulliolatus {La.i\i.) (cf. antea). ¥ 9. xi. 1905 (No. 7058).
20
( 304 )
32. Cuculus saturatus Blyth. (cf. Xoi: ZooL VM)\. p. 20l). 1 c? juv. 13. xi.
1905, seems to beloug to tliis furm (No. 7098).
33. Lalage timoriensis (S. Miill.) (cf. antea). 2 cJ <? 9. xi. 1905 (Nos. 7056, 7057).
34. Gniucalus melanopg (Lath.) (cf. antea). Five specimeus (Nos. 7037, 7039,
7040, 7041, 7042).
35. Graucalus hypoleucus Gonkl (cf Finscb, Xotes Leyden Museum xxii.
p. 250). <? ? Luang, 11. xi. 1905 (Nos. 7077, 7078)
36. Locustella ochotensis (Midd.).
Sylvia {Loaislelki) orholeiisis MiiidenJorff, Slbii: lieise II. 2, p. 185, Taf. xvi. 7—8 (185.3 : Mouth
of the River Ouda into tlic Sea of Ochotsk).
1 (J jnu. Luaiig Island, 14. xi. 1905 (No. 7098). '-Iris blacki.sh ; feet light
brown ; bill dull bhiclc, pale beli)W."
37. Grallina picata (Lath,).
Gracula pkata Latham, Ind. Oni. Stijjjil. p. 29 (1801 : Avistralia).
1 ? ad., Lnang, 10. xi. lOO."). " Iris black ; feet black ; bill yellowish white."
(No. 70n7.)
This Australian species is new to tlie Sonth-West Islands. Mr. Kuhn formerly
obtained a single specimen on tlie island of Koer, in tlie South-East Islands.
Probably it straggles occasionally northwards, being as a rule resident in Australia.
38. Zosterops bassetti Sharpe.
Cf. antea.
2 6S, November lOiio (Nos. 7053, 7075).
39. Stigmatops squamata Salvad.
Cf. antea.
c??, Luang, 12, 15. xi. 1905 (Nos. 7081, 71i)0). Wings : cj 72, ? 63 mm.
40. Taeniopygia castauotis iusularis Wail.
Cf. Nov. Zo,d. 11104. p. 218.
Ten specimens obtained on Luang (Nos. 7046, 7054, 7055, 7059, 7060, 7061,
7062, 7005, 7070, 7070a).
41. Oriolus viridis Latli.
Gracula ririclis Latham, Ind. Urn. ShjijiI. p. xxviii (1801 : Australia).
1 (? jnv. Luang Island, 11. xi. 1905 (No. 7074). " Iris scarlet ; feet brownish
red ; bill blackish."
The occurrence of tliis species is probably an irregular one on Luang.
( 305 )
CRITICAL NOTES ON THE TYPES OF LITTLE-KNOWN
SPECIES OF NEOTROPICAL BIRDS.
By C!. E. HELLMAYR.
PART I.
"VTTHEN I began, six }'ears ago, to study the neotropical aviiauna and to
' V collect material for my proposed work on tlie Birds of Brazil, it soon
became clear to me that there was a great uncertainty in the nomenclatnre of
many species, and that a thorough revision of those types about which any doubts
existed would be very useful ; and notwithstanding its many apparent difficulties,
I decided to undertake this. Thus, during the last four years, I have carefully
-examined a great number of types in various museums of Europe, and I intend
to publish the results of these stndies in a series of papers of which this f(.irms
the first instalment.
Remarks on some donbtful or little-known species have already been published
by me in various periodicals, also a ])aper dealing with certain genera oi Fritigillidae
in the Verhamllungen der zoolog.-hotan. Gesellscha/t , of Vienna, 19(J4. pp. olG-S".
A more extensive memoir, in which many types of Tschudi, Heinhardt, Cabanis
and Pelzeln are discussed, appeared in the Journal fiiv Ornitlwloyie, I'JOo. pp. 1-33,
nnder the joint authorship of Count Berlepsch and myself An account of the birds
described by Spix in his great work, Avium Brasiliae Species Novae, has lately
been issued ; and critical notes on the types of the Tracheophonac in the Paris
Museum, by Meuegaux and myself, have been published in some French
periodicals.*
The main object of the present paper is to clear up a number of the many
doubtful and little-known species preserved in the British Museum, but several
types belonging to the Museum d'Histoire Naturelle of Paris are also discussed.
A special paper will be devoted to many additional types of the French naturalists,
chiefly those of Lafresnaye and D'Orbiguy.
My conclusions are, wherever possible, based on a large amount of material,
for the use of which I am much indebted to several kind friends, particularly
Count Berlepsch, Dr. H. 0. Forbes, Dr. E. Hartert, Dr. L. von Lorenz-Liburnau, of
Vienna, and the Hon. L. W. Rothschild, to all of whom my sincere thanks are
due. Neither must I forget to mention the late Professor Oustalet, and Dr. A.
Meneganx, who not only gave me free access to the treasures of the Paris Museum,
but also facilitated my work in every possible way. To Mr. Rothschild I am
specially obliged for kindly offering to publish these notes in the Novitates
Zoologicae.
1. Cyanocorax inexpectatus Elliot = C. caeruleus (Vieill).
Pica caenilm Vieillot, A'omc. Did. .vxvi. (1818) p. 126 (ex Azara : Paraguay).
Cyanocorax Ileclelii Pelzeln, Sitz.-Ber. Akad. Wien xx. (185(i) p. 103 [Rio Boraxudo, near
Paranagua, Parana],
Cyanocorax inexpectatus Elliot, Ibis, 1878. p. 55 [''south of Sao Paulo "].
1. Mus. Paris, av. juv. " Capt. de St. Paul. M. A. de Saint Hilaire. C. inex-
pectatus Elliot. Type de I'esp^ce." Wing 197 ; tail 172 mm.
* Bull. Mus. Paris, 190,5. pp. 372-81; Mem. Soo. delist, nat. Autuii, xix, (I'Mii) pp. 43-12G; Hull.
Sao. Philom. Taris, 1906. pp. 21-58.
(,306 )
2. Mus. Vindob. ? ad., Rio Boraxudo, Dear Paraiiagm'i, Parana, December
1820. Natterer coll. Wiug 190 ; tail 170 mm.
3. Mus. Viudob. c? ad., Rio Boraxndo, December 1820. Natterer coll. Wing
195 ; tail 174 mm.
4. Mus. Viudob. ? , Rio Boraxudo, December 1820. Natterer coll. Wing 183 ;
tail IGO mm.
(Nos. 2 to 4 types of C. heckelii Pelz.)
5. Mus. Tring, No. 697, ? ad., Roca Nova, Serra do Mar, Parana, October 27,
19111. A. Robert coll. Wing 192 ; tail 166 mm.
6. Mils. Tring, No. 718, ? ad., Roia Nova, Serra do Mar, Parana, November 4,
191)1. A. Robert coll. Wing 193; tail 165 mm.
Besides the above specimens I have examined : 2 c? J, 1 ? , Ignape, S. Panlo ;
one adult, y])auema, 8. Panlo; 2 c?c?, 2 ??, Scaramuza ; 3 ??, Ytarare,
S. Paulo; 1 (?, Jaguaraiba, all collected by Natterer, except those from Ignajie,
which were obtained by Mr. Krone, and the one from Ypanema. I compared also
a series from Taqnara and Arroio Grande, Rio Grande do Snl, and two adults from
Blumeuan, Sauta Catharina (Mus. H. v. Berlepsch).
The examination of this material proves, beyond doubt, that the shade of the
blue colour, on which the "species" C. heckelii aud C. inexpectatiis have been based,
is of no geographical significance, since we find, at the same localities, verditer-blue
{inexpectatiis} aud purjale blue {hechelii') specimens along with typical caeruleus.
This is especially illustrated by the series from Rio Grande do Snl • in Count
Berlepsch's collection. Three skins from Iguapd, and two others from Blumenau,
agree perfectly with the types of C heckelii, and I at first thought the latter
might be a difterent form confined to the coast region. Two ? ad. from the Serra
da Mar (almost topotypes of C. heckelii), however, do not belong to tliis purple-blue
form. One (No. 5) is, like the type of C. iiie.cpectatus, verditer-blue, the other
(No. 6) is typical caeruleus ! One adult from Ypanema, on the other hand, is
exactly intermediate in colour between the i)urplc-bluo {heckelii) and the caeruleus
phase, aud a female from Scaramuza has the lower [larts almost as purple as the
types of C heckelii 1 Moreover, the latter " form " has been lately recorded from
Paraguay ,-f the tj^pical locality of C. caeruleus I There is no constant diti'erence
in the colour of the throat or in the shape of the frontal feathers ; nor are the
dimensions diflerent. The types of C. heckelii being in very worn ])lumage, the
throat appears dull brownish, black, without any gloss. I may also mention that
the typical specimen of C. inexpectatus is an immature bird, with the under jiarts
mainly brownish grey, but some verditer-bhiish feathers are just coming out.
I append the measurements of Natterer's series and some other specimens, to
show that there is no difierence in size.
C. heckelii-^ha,se (purple-bine).
1 ? ad., Tguap^. Wing 180 ; tail 152 mm.
2 cJ ad. (sex ?), Iguape. Wing 184, 191 ; tail 150, 162 mm.
Intermediate between heckelii and caeruleus.
Ad., Ypanema. Wing 192 ; tail 175 mm.
? ad., Scaramuza, August 2, 1820. Natterer coll. Wing 192; tail 180 mm.
' Cf. also Berlepsch, Zfitschr. gea. Ornitk. ii. p. 127.
f Oberholser, Proc. V. S. Nat. Mus. xxv. p. 140.
( 307 )
C. c«r?<fe««-phase. Natterer coll.'
(?, Scaramnza. Wing 205 ; tail 190 mm.
5, Scaramuza. Wing 203 ; tail 192 mm.
?, Scaramnza. Wing 201 ; tail 177 mm.
2 ? ?, Ytarare.* Wing 200, 202 ; tail 17.5, 176 mm.
6, Jagnaraiba.* Wing 205 ; tail 190 mm.
2. Anthus lutescens Pncb.
Aiithus lutescens (ex Cuvier, MS.) Pucheran, Arch. Mas. Paris vii. (1855) p. .343 ["Brfsil '' — coll.
Delalande].
Mns. Paris, ad. labelled : " Farlonse jannatre. Anf/iiis lutescens (h\v. type,
du Br^sil, par M. Delalande." Wing 63| ; tail 46 ; bill 12 mm.
Tbis is tbe species called A. vuj'us iu tbe Cat. B. Brit. Mns., as already
recognised liy Count Berlepsch {Zeitschr. ges. Ornith. ii. 1885. ji. 114).
Mr. Ridgway having recently em])loyed the name A. parrus for tbe small
neotropical Pijjit, I took the series of tbe Tring Museum over to Paris in order
to settle the question definitely, and found the surmise of tbe Count fully
confirmed.
Tbe type agrees in every respect with a series from Babia and Rio, but,
owing to its having been exposed to the light for nearly a century, the colour.s
have faded and appear rather paler, especially on tbe back and chest. The
markings on tbe two outer tail-feathers are e.xactly as in other specimens from
S.E. Brazil.
With a considerable series before me, 1 can find no differences between
examples from Rio, Bahia, Para, Corrientes, and Tucuman. All have a broad
blackish strijie along tbe inner web of the outermost tail-feather, reaching almost
to tbe tip, and there are no dusky shaft-stripes on the sides of the body.
Eight skins from Chiriqui (yl. parvus Lawr.) differ very markedly in the
following points : Tbe outermost tail-feather is almost entirely white, there being
only at the extreme base a faint indication of a greyish inner margin ; tbe blackish
shaft-stripes on the lower parts are much coarser, and not confined to tbe foreueck,
but extended over the sides and flanks ; the wings are somewhat shorter.
Specimens from British Guiana and Bogotd collections agree in the coloration
of the under parts with A. lutescens of S.E. Brazil, but the outermost tail-feather
is exactly as in the Panama form. Should additional skins confirm the constancy
of their characters, they must be separated as a third race.
According to our present knowledge the following geographical forms of the
small Pipit of South America are recognisable :
a. A. lutescens lutescens Puch., Eastern Brazil from Para to S. Paulo, west to
Corrientes and Tucuman, Argentine Republic.
b. A. lutescens subsp., British Guiana (Roraima, Rio Rupununi, Annai) and
Colombia (Bogota coll.).
c. A. lutescens parvus Lawr., Panama, Veragua and Chiriqni.
d. A. lutescens permianus Nicholson, Coast region of Western Peru from
Trujillo (specimens in Mns. Tring, Baron coll.) to Islay.
* As will be senn from Natterer's itinerary, these localities are not far from I'aranagua (I'elzelii,
Orn. Brasil. iv. p. v).
( 308 )
3. Haplospiza uniformis Scl. & Salv. should be Spodiornis uniformis (Scl. & Salv.)
Haploxpiza imiformh Sclater & Salvia, Nomencl. Av. Neotr. 1873. p. 157 [Near Jalapa, Vera Cruz,
S.E. Mexico],
Mr. Kidgway was much puzzled about the systematic position of this bird,
and suggested that it might prove to be identical vi\th.Acantkidopsbairdili\Agw.
This, however, is not the case, as I satisfied myself by careful examination of the
tyjie specimen, but Mr. Ridgway was quite right in sujiposing that the species
had been placed in a; wrong genus. Messrs. Sclater and Salvin were deceived by
the superficial resemblance between H. unifofmis and H. unicolor, and cannot have
examined the structure of the wing, otherwise they would have been led to recognise
that the two species were qnite distinct, and that the nearest ally of H. unifonnis
was in reality Spodiornis jardinei. Both species agree not only in the wing-
formula, but also the bill is almost of the same form. H. unicolor, on the other
hand, has a qnite differently shaped wing, as will be seen from the following lines.
Haplospiza unicolor I'ab.* : 5 > 1 > 6 ; 3rd and 4th primaries longest, 1st
about 5 mm. < 4th.
" Haplospiza " uniformis Scl. k Salv. : 4 > 1 > 5 ; 2nd and 3rd longest,
4th scarcely shorter ; 1st about 5 mm. > 5th, and only about 1 — 2 mm. < 4th.
Spodiornis jardinei Scl.f : 4 > 1 > 5 ; 2nd and 3rd longest, 4th scarcely
shorter ; 1st about 4—5 mm. > 5th, and bat 1 — 2 mm. < 4th.
The bill in H. uniformis resembles very much that of S. jardinei, being
perhaps a little longer and less curved at the tip, and it also seems that the
tomiae at the base of the upper mandible are less turned in. These differences,
however, are very trifling. In B. unicolor the bill, although of the same general
shape, is distinctly stouter and broader at the base.
In coloration there is no difference whatever between the type of //. uniformis
and a series of S ads. of 5. jardinei. The Tring Museum possesses an adult S
from Carrillo, Costa Rica (Underwood coll.), which differs from the type in being
slightly smaller, and of a jialer, more bluish slate-grey colour everywhere.
The bill is entirely phimbeous, while the type from Xalapa has the lower
mandible whitish grey. Whether these are more than individual variations I cannot
say with certainty without series from both localities.
Additional specimens from Central America may perhaps prove //. uniformis
to be not properly separable from S. jardinei, but for the present they must
be regarded as distinct. There can be, however, not the slightest doubt that
H. uniformis has to be transferred from Haplospiza to Spodiornis. Its synonymy
is as follows :
Spodiornis uniformis (Scl. & Salv.).
Jlajilospha vnlfiirmix Sclater & Salvin, Nomem-I. Ai\ Nrntnp. 1873. p. 157 (Jalapa, Vera Cniz) ;
Salvin & Godman, Biol. Centr. Americ. i. (1881')) p. 360, pi. 27, fig. 1 (Jalapa) ; Sharpe, Cat.
Birds Bril. Mux. xii. (1888) p. 627 (Jalapa) ; llidgway, Birds Xorth and .Middle America, i.
(1901), p. 521 (Jalapa).
Hab. S.E. Mexico : Jalapa (De Oca) ; Costa Rica : Carrillo (Underwood coll.).
1. Mus. Brit., (? ad., Jalapa, S. Mexico (De Oca coll.) 1872. Type of the
species. Wings 75 ; tail 50 ; bill 14 mm.
2. Mus. Tring, J ad., "Carrillo, Costa Rica, December 4, 1898." C. V.
Underwood coll. Wing 71 ; tail 46 ; bill 14 mm.
• 20 specimens. The l.st primary is either equal to or a little longer tliaii the Gth.
t 14 specimens examined.
( 309 )
For the sake of comparison I append the measurements of Sp. jardinei.
2 c?c? ad. from Ecuador. Wing 70, 68 ; tail 49, 46 ; bill 13, 121 mm.
3 cf c? from Bogota coll. Wing 67, 69 ; tail 46, 49 ; bill 12, 13 mm,
1 S jr., from Marcapata, S.E. Peru. Wing 69 ; tail 49 ; bill 1 3 mm.
1 S jr., from Bolivia (Mus. H. v. Berlepsch). Wing 69; tail 50 ; bill 13 mm.
llaploipiza and Spodiornis are doubtless ver}' near allies, and must be placed
close together, not far from Pkrt/gilus ; the pattern of coloration in tlie females is
similar to that of P. unicolor, a further proof of their near relations. The
latter species, however, cannot be included in the same genus with Haplospiza
unicolor, as has been done by several American authors, for the shape of the wing
is widely different.
4. Crithagra hilarii Bp. = Sicalis a. arvensis (Kittl.)
Frinrjilla arvensis Kittlit/,, .l/c/K. Acad. St. Petersb. (sav. eir.) ii. (1835) p. 134 (Chili).*
Crithagra hilarii Bonaparte, Consp. Av. i. (1850) p. 521 (Mus. Paris : ex Brazil.).
Mus. Paris, skin : " No. 27. Mr. St. Hilaire, 23. mai 1821. Crith. hilarii Bonap.
(typ.)."— Wing 73 ; tail 53 ; bill 19 mm.
This is by no means .S'. a. minor (_'ab., with which Bonaparte's name has been
doubtfully associated by Sharpe,t but belongs to the large race of S. arvensis found
in South Brazil. The original specimen agrees perfectly with another from Rio
Grande do Sul (Mns. Berlepsch) ; in both there are no white markings on the tail.
Unless the South Brazilian form be separable from the typical Chilian birds,
C. hilarii is to be considered as a synonym of S. a. arvensis.
It may be added that the type came from South Brazil : S. Paulo or Parana,
where, according to the registers of the Paris Museum, M. Auguste de St. Hilaire
was collecting during the year 1821.
I hope to discuss the various forms of the S. arvensis group on a later occasion.
5. Buarremon baeri Oust, should be Compsospiza baeri (Oust.) !
Buarremou Baeri Oustalet, Bull. Mus. Paris x. (1904) p. 43 (Lagunita, Tucuman).
This bird, one of Mons. G. A. Baer's discoveries in the district of Tucumdn,
belongs by no means to the genus Buarremon, as supposed by M. Oustalet, but
represents a second species of the genus Compsospiza, proposed by Count Berlepsch*
for a remarkable Finch of the highlands of Bolivia, C. garleppi. The type of this
species having remained unique in the Count's magnificent collection, the discovery
of a second form was of great interest.
C. baeri (Oust.) agrees with the type of the genus not only in the shape of the
bill and in the form of the wings and tail, but presents also a close resemblance in
coloration. The distribution of the ferruginous aud grey colours on the top and
sides of the head is absolutely alike in both species, but C. baeri differs at once by
having only the throat aud foreneck aud the under tail-coverts ferruginous, while
the remaining underparts are pale slaty grey with a slight olive hue on the sides.
Moreover, the slate grey of the upperparts is paler, the edge of the wing slate-
grey (not ferruginous), and there is no trace of the white apical margins on the
two outer tail-feathers. C. baeri is also much smaller.
• I have not heen able to verify the above quotation,
t Cat. Birds xii. p. 384,
j /4;s 1893. p. 207 pi vi.
( 310 )
Besides the type in the Paris Maaeum, I examined two c? J in the Trinj;
Musenm and one c? ad. in Count Berlejjsch's collection. All were taken at Lagnuita,
Tucuman, at an elevation of 3000 metres, January 31, February 1, 4, 5, 19U3. They
present the following measurements: Wing 70 to 77i ; tail 74 to 75|; bill
13i to 14i mm.
0. Tachyphoniis chloricterus Vieill. should be Orthogonys chloricterus (Vieill.)
Tachi/pliuniis cJilorirtertis Vieillot, Xduv. Diet, xxxii (1819) p. ,'ifiO ["on le trouve au Bn'sil d'oii U a
ete apportt' par M. de Lalande fils"] ; Vieillot, Tahl. em: melh. Oniilh. ii.(182'2)p. 804 [" dvposd
au Museum d'histoire naturelle "].
" Twhi/phom a qnudettes bleues" Lesson, Traile d'Orn. (1831) p. 463 [part.: " Femelle : olive
jaunCitre. Du Bresil"] ; Pucheran, Arch. Mm. vii (18.'j5i p. 378.
Tamigru virirlii SpLx, Ac. Brasil. ii. (1825) p. 36 tab. XI.VJII. fig. 2 [" in provincia Rio de Janeiro"—
type in Munich Museum examined].
Orthof/oni/ft vh'idi.'< auct.
Mns. Paris ; spec, typ., labelled as follows : " Ln.mprote icterope ? , Piji-iMiija
ictei-opus Vii'ill. Tacki/phonu-i chloricfenis Vieill. (T.),ilu Bn'sil, par Delala/^i/.f"
and on the bottom of the perch: '^ du Brcsil par Delalaudc 1816. Tachyphonus
chloricterus Vieill. (type)." — Wing 92 ; tail 84 ; bill 19 mm.
This specimen agrees perfectly with an example of Orthogonys viridis (Spix)
from Parana, with which I compared it. The lower surfiice is perhaps rather paler
olive yellow and the back duller greenish, both diftereuces being without doubt due
to fading, for the specimen was exposed to light since nearly a century.
Vieillot's name, being the earliest, must be accepted for the species commonly
called 0. viridis.
In the Cat. Birds xi. p. 193 it was doubtfully referred to the ? of Cyanictcrus
cyanicterus (Vieill.).
7. Nemosia chrysopis Sd. & Salv. = Thlypopsis sordida (Lafr. k Orb.) juv. !
Nemosia sordida Lafresnaye & Orbigny, Si^ii. Ac, i. in : Mar/. Zoo!, cl. ii. (1837) p. 28 [Yuracares,
rep. Boliviana].
.V. chri/sopis Sclater & Salvin, P. Z. S. 1880. p. 155 [Sarayayu, E. Ecuador].
Spec. typ. in Mns. Brit. : " Sarayaiju, E. Ecuador.— 0. Buckley coll. Type—
Nemosia chrysopis Sol. & Salv." — Wing 66 ; tail :j7 ; bill 11 J mm.
The type is an immature bird of T. sordida in the well-known juvenile
dress, as descrilied and accurately figured by D'Orbigny,* and it is difficult to
understand liow it could be mistaken for "a very distinct species of the section
Thlypopsis." The Tring Mnseum possesses a specimen in corresponding plumage
from Altagriuiii, Orinoco, and I have seen others from Bahia in Count Berlepsch's
collection. U'Orbigny's types from Yuracares, which I examined in the Paris
Museum, are also immature, one being exactly like the typical specimen of
N. chrysopis, the other even younger, in the fluffy plumage of the nestling.
In both, the type of N. chrysopis and the older specimen of D'Orbigny's, some
of the feathers of the adult plumage are just coming out, so several orango-rnfous
ones on the jiilenm, and some creamy-buff ones on the foreneck. Tiie tail-feathers
are slightly pointed — an unmistakable sign of immaturity. The top of the head and
mantle are yellowish-green as in the juvenile jdnmage, but rump and upper tail-
coverts have already attained the olive-grey colour of tlie adults; as in the latter,
* Voij. Avier. tncr/d. OU. tab. IS, fig, 2.
( 311 )
the middle of the abdomt-n and tlie under tail-covcrts are buff}' white, but the chi'st
and sides of the body are olive j'ellow as in j'oung birds.
That the t3'pe of ]V. chrysopis is the young of T. sorclida and not of
T. s. amazonum is proved by the newly growing feathers on tlie foreneek being
creamy-buff', not jjure cinereous.
The second specimen, from Santa ( 'ruz, E. Pern, referred to T. chn/iiopis by
Sclater,* is quite a young bird of T. s. amazoinim, the types of wliich came also
from the Ucayali. The range of the two forms is as follows :
1. Thl//popsis sordida sordida (LalV. k Orb.)
East Bolivia : Ynracares (D'Orbiguy), San Mateo (G. Garlepp. — Mus.
Berlepsch and Coll. Hellmayr). Brazil: Goiaz, (Juyaba, Rio Madeira (Natterer
coll.t), Chapada (H. H. Smith coll.). — Minas Geraes : Lagoa Santa and Sete
Lagoas (Burmeister & Reinhardt coll. J). — Bahia (trade skins in Mus. Berlepsch,
Tring, etc.). — Pernambuco (Forbes coll.) — -Venezuela, Orinoco R. : Capuchin,
El Fraile, Altagracia (Clierrie coll.).§ — East Ecuador : Saraya(ju (C. Buckley coll.).
2. T. sordida amazonum Scl.
N.E. Peru : Lower Ucayali, Santa Cruz, Nanta (E. Bartlett coll.), Pebas
(Hanxwell coll.). — C. Peru: La Merced, Chanchamayo (J. Kalinowski coll.).
N.B. — The locality " Bolivia " of the Cat. B. xi. p. 220 is evidently erroneous.
s. Arremon polionotus Bp.
Arremon pollonotiin Bonaparte, Consp. Ae. i. (1850) p. 488 [Corrientes. — Mus. Paris].
An-emon callistiis Oberholser, Proe. Biol. Sue. Wash. xiv. (1901) p. 188 [Sapucay, Paraguay].
One mounted sjiecimen, in the Paris Museum, is labelled : " Arremon jiolionote ;
Arremon polionotus Pchr., Bp. (T.) ; de (Corrientes, par M. D'Orbiguy." The stand
bears the following inscription : " de Corrientes, par M. D'Orbiguy, juillet 1820.
Type de la description de M. (Jh. Bonajiarte dans le (!onspectus Avium." — Al. TOA ;
Cauda 75; rostr. 14i mm.
This specimen, an adult cJ, agrees in every respect with the original description
of Bonaparte, and is undoubtedly the type of the species. Two adult males from
Paraguay (^-1. callistus) are in no way distiuguishable, as we should have expected
from the locality. Most probably Mr. Oberholser, when describing his .1. callistus,
compared the Paraguay bird with specimens from Mattogrosso, which he seems to
have considered to be true .1. polionotus. Having always suspected the identity of the
Paraguay form with the latter, 1 took to Paris some specimens from Mattogros.so and
Paraguay, and found my surmise fully confirmed. The type in the Paris Museum
has the black jugular band just as broad as the Paraguay birds, and the back is ol'
the same dark slaty grey without any olivaceous tinge ; only the edge of the wing
is slightly more mixed with white in the tyi>e specimen. The lack of the yellowish-
green humeral patch upon which Mr. Oberholser lays so much stress does not seem
* Cat. Jiirds xi. p. 229.
t Nemosiafulvncena + X. sordida Pelzeln, Orn. Bras. iii. (ls(;<))pp. 215, 216. — I Lave examined Ihese
specimens. N.fnUrscem are the adults, N. sordida tlie young birds. They belong to T. s. sordida and
not to T. s. amazo7iuin, as supposed by Wclaler.
t Xemnsia fiilvicejis Burmeister, Si/st. Vhers. iii. p. 1511; iV. fulvcscens + K sordida Reiuhardt, \'id.
Medd. Kjlihenhavn 1S70. p. -123.
§ Nov. Zool. vs.. p. 23.
( 312 )
to be a relialile character, for it is prpseut in the Corrieutes ami Paraguay birils.
I may add that in all adult males of A. polioitotus and of its northern form the head
is uniform black with no trace of the grey median stripe which is to be seen in
females and yonng birds.*
Measurements of .4. polioiwtus :
1. S ad., Paraguay, November 10, l.SiJ3.— Bohls coll. No. oD. Al. 77 ; caud. 7(1 ;
rostr. 14 mm.
? ad., Paraguay, November 10, 1893.— Bohls coll. No. 50. Al. 70 ; caud. 02 ;
o
rostr. 141 miu.
3. (? ad., Colonia Risso, Rio Ai)a, N. Paraguay ; A. Borelli leg., No. 241.
Al. l^ ; caud. 68; rostr. 14J mm.
Specimens from Mattogrosso and Goia;6 differ from the typical form in their
decidedly narrower jugular band and in having the back of a paler grey with a slight
olivaceous suffusion. The black band across the foreneck, which in the type of
A. polionotus and in the examples from Paraguay is about 10 mm. broad, measures
only "'O mm. These differences, although slight, seem to be constant, and the
form from Central Brazil may bo regarded as a distinct subspecies. As I have
ascertained by examination of the type specimen, its ])roper name is
Arremon polionotus devillii I'es Murs.
Arremon Devillii Des Murs (ex Bonaparte MS.) ia Castelnau, Vn,j. Amir. SmJ, Oln. p. 60, tab. x.\.
fig. 2 (185o) [no locality].
A. dnillii Sclater, P. Z. X. 1856. p. 81 ['• prov. Goiaz in Brazil.''— Castelnau & Deville leg.—
AIus. Paris].
Arremon poliunvtiis (nee Bp.) Pelzeln, Zur Om. Brasil. iii. (18G9) p. 217.
Arremon (krillii + A. polionotus Sclater, Cut. Birds xi. (1886) pp. Hi, -278.
Mus. Paris ; mounted in the gallery and labelled as follows : " Arremon de
Deville, Arremon Devillii, Bp. (T.), du Brisil, par MM. de Castelnau et Deville."
The stand bears the foUowiug inscription : " 1840— Cat. gen. No. 972. Arremon
Devillei Bonap., type de I'espece ; iudividu figure dans I'atlas du voyage. Province
de Go)/as, Brt^sil, par BIM. Castelnau et Deville."— Al. 71 ; caud. Of); rostr. 1.5 mm.
This specimen is a young bird, as proved by the strong olivaceous tinge of the
back, by the presence of the greyish crown-stripe, by the pale brownish wash on the
sides of the body, and by the colour of the bill. The lower edges of the upper
mandible are dark brownish yellow, not clear yellow as in adult birds, and not so
sharply deiined against the blackish culminal stripe. The bird differs from adult
males of Chapada, Cuyaba, and Goiaz in the same way as does a female of true
A. polionotus (from Paraguay) from the adult males of the same locality, thus
jiroving that the differences are only due to age or sex. In both examples the back
and the ashy stripe along the middle of the occiimt show a distinct olivaceous
suffusion. in adult males the back is of a clearer slate-grey colour, with only
a faint greenish admixture here and there, and the crown-stripe is altogether
wanting or but slightly indicated by grey edges to some of the feathers. The sides
of the body are also pure ashy grey, whereas they are of a jnilc brownish tnit m
females and young males. The ground colour of the under surface in Des Murs'
type, however, is pure white, as in adult males. In all other respects the type
of A. (lerillii fully agrees with specimens from C. Brazil, and I have not the
slightest doubt it is a yonng male of the northern form of .1. polionotus.
* The same has been stated by Mr. AUen ilivll. Amir. Mus. iii. p. 3G2 : " A. polionotus ").
( 313 )
2 cJc? iicl. from Chiqnitos ex D'Orbigiiy in the Paris Museum belong also to tiie
form witli narrow black jugular band.
Thus we have two sub.-iiiecies, the distribution of which is as follows :
a. Arremon jjolioiiot.iis polionotus Bp.
Hab. Corrientes, Argentina (D'Orbigny). Paraguay : Sapuoay (Foster) ;
Colonia Risso, Rio Apa (Borelli).
h. An-i'mon polionotus denllii Des Murs.
Similar to the typical form, but distinguishable by its decidedly narrower
jugular band, and by having the back of a lighter, paler grey colour, with a slight
admixture of olivaceous.
Hab. Central Brazil: Goiaz (Casteluau & Deville); Cuyaba (Natterer) ;
Chapada (Smith) ; Urucum and Corumba (Borelli) ;
Eastern Bolima : Chiquitos (D'Orbigny — Mus. Paris).
9. Arremon wuchereri Scl. & Salv. = A. flavirostris Sw. S ad.
Arremon flavirostris Swaiasou, Animals in ^[(■n<lg. (1838) p. 347 [" Brazil"].
A. wuchereri Sclater & Salviu, Nomend. Ai\ neoti-op. (1873) p. 278 [Bahia] ; Iheriog, Revisl. Mns.
Paul, T. (I'JOl), p. -JliS [Jabotioabal, S. Paulo].
A. flavirostris Reinhardt, Vidensk. Medcl. Kj^henhami (1870) p. 421 [Minas Geraes].
As can be seen from Mr. Sclater's key * to the genus, A. ivuchereri differs
from A. flavirostris only in lacking the cinereous vertical stripe. This character,
however, is very variable, and evidently depends on age, as we learn from
the examination of a large series of the allied ^1. polionotus. In this series the
adult males, witli pure schistaceous ujjper parts and pure white belly, have the
head uniform black, only the young males aud females show an olivaceous grey
vertical stripe. Through the kindness of Professor von Ihering, of S. Paulo, Brazil,
I had the opportunity of comparing some specimens of A. flavirostris, collected in
the north-eastern part of S. Paulo ; and in the Tring Museum there is one example
obtained by Mr. A. Robert in the province of Araguay, Minas Geraes. This series
presents the same variation us A. polionotus. In three specimens,! two of them
marked " ? " by the collector, there is a broad olivaceous grey vertical stripe,
beginning just above the eye and confluent with the olive-grey colour of the nape,
the sides of the body are pale greyish brown or brownish, and in one specimen
the chest, below the black jugular band, shows a slight buffy tinge. Both
characters are signs of immaturity.
On the other hand, one <S (No. 4568, Bebedonro — Mus. Tring), with jiure
schistaceous nape and pure grey sides of the body, lacks the vertical stripe, some
feathers of the hind crown only showing narrow slate-grey margins. This is also the
case in the type of A. wuchereri, which differs from our specimens (ex Bebedonro).
only in its longer wings and in having the black jugular band slightly bmader
Yet, as will be seen from the measurements given below, one 6 from Jaboticubal
comes very near in the length of the wing, and I have no doubt the type of
A. wuchereri is nothing but a c? ad. of A. flavirostris.
Mus. Tring, c? ad., Bebedouro, iv. 1904. Wing 75i ; tail 75| ; bill IT, mm.
* Cat. Birds xi. p. 272.
t No. 4567, Banx'tos— Mus. Paulist.; No. 4569, Bebedouro— Mus. Berlepsch; No. 4S2, .\. Robert coll.,
Rio Jordao, Minas Geraes.
( 314 )
Mns. Tring (?), Rio Jordao, Miuas. Wing 74i ; tail 67 ; bill ITii min.
Mus. Bcrlepscb, ?, Bebedouro, iv. 1904. Wing 75 ; tail 08 ; bill i:ij mm.
Mns. Paulist, ?, Barretos, May 19U4. No. 4507. Wiug 74; tail 08 ; bill
144 mm.
Mus. Panlist., 1240, c?, Jaboticabal, S. Paulo. Wing 79^ ; tail 78 mm.
Mus. Brit. ((?) ad., Babia. (Type of ^1. ima-hereri Scl. k Salv") Wing 82 ; tail
73 ; bill 14| mm.
The distribution of A. Jiavirostris is as follows : Bnliia (Wucherer) ; Minas
Geraes : Lagoa Santa and Sette Lagoas (Roinhardt & Lund) ; Rio Jordao,
province Aragnay, 700 — 900 m. (A. Robert coll. —Mus. Tring); north-eastern
S. Paulo : Jaboticabal (Lima coll. — Mus. Paulista) ; Bebedouro aud Barretos, Rio
Grande (Garbe coll. — Mus. Paulista et Tring) ; Santa Maria (Behn coll. — Mus.
Berlin). The locality Cametii. Lower Amazons (specimen in Mus. Berlin), seems to
me a little doubtful.
A. Jiavirostris is not at all related to A. silens, but a very near ally of
A. p. polionotus and .4. p. decillii, from both of which it only differs in having the
back olive-green (instead of cinereous). A. silens, on the other hand, is characterised
by its entirely black bill, a black chin-spot, and by the lengthened snperciliaries
which commence on the front instead of above the eye.
10. Saltator azarae D'Orb. = Saltator coerulescens azarae D'Orb,
Sahator Azarae D'Orbigny, Voijage AnU-r. Mi'rki., 0/se««.c (publ. between 18,^8 and 1847) p. 287
[" dans les provinces de Moxos et Santa-Cruz de la Sierra en Boli\-ie "].
Four specimens in the Paris Museum, labelled as follows :
No. 1 (mounted) : " Saltator azarae D'Orb. (type) Mojos. D'Orbigny (1834—
D. 251— 317) male." Wing 104 ; tail 92 : bill 20 mm.
No. 2 (skin): " D. 317. AJojos. Saltator azarae D'Orb. D'Orbigny, 1834."
Wing 110 ; tail 100; bill 20 mm.
No. 3 (skin) ; " D. 317. Mojos. Saltator azarae D'Orb. 251 ; D'Orbigny, 1834."
Wing 111 ; tail 100; bill 20 mm.
No. 4 (mounted) : " Saltator azarae D'Orb. (type). Sta 0-«.;— D'Orbigny,
(1834. D. 317— 251)-femelle." Wing 102 ; tail 98 ; bill 21^ mm.
Nos. 1, 2 and 3 agree very closely with the dark-backed form inhabiting the
Upper Amazonian region. Compared with two adult birds from Peru (biuitos
and C'hanchamayo) they are a shade paler both on the upper and lower surface,
but very much darker than true 5'. c. coerulescens. From the latter they are
also distinguished by having the outer webs of the remiges dark slate-colour, in
which respect they agree exactly with Peruvian specimens.
D'Orbigny's female, No. 4, however, does not belong to this form, but is in
every way similar to typical S. c. coerulescens of Corrientes aud Paraguay. The
upper surface is olive-grey (not dark slaty olive as in Nos. 1 — 3), and the breast
much paler greyish buff. As in so many other instances, the form found in the
plains to the east of the Bolivian Andes proves to be the same as that inhabiting
the campos region of the interior of Brazil ; while farther to the north, on the
tributaries of the Rio Madeira, the Amazonian representative occurs.
The slaty-backed form has been identified in the Cat. Birds xi. with Tnnagra
superciliaris Spix. The type of this species, however, agrees perfectly with
specimens from Paraguay (typical S. c. coerulescens), as will be shown in my forth-
( 315 )
coming paper on Spix types. Although D'Orbigny's so-called female belongs to
another subspecies, his diagnosis is clearly referable to the dark-backed form, which
accordingly ought to be called S. coendescens azarac, D'Orb.
The distribution of the two subspecies is as follows :
a. Saltator coerulescens coerulescens Vieill.
Saltator coerulescens Vieillot, Nutiv. Diet. xiv. (1817), p. 105 (ex Azava No. 81 : Paraguay).
Tanugra siiperciliaris Spix, Av. Bras. ii. (1825), p. 44, tab. Ivii. ('• in campis fl. St. Frauoisci prope
pagum Joaseiro" Bahia, Brazil).
Saltator azarae (nee D'Orbigny), Pelzein, Zur Orit. Bras. iii. (1869) p. 219 (Mattogrosso).
S. coerulescens Salvador!, Boll. Mns, Torino xv. No. 378 (1900) p. 4 (Mattogrosso).
S. Azarae D'Orbigny, Voi/age, Oiseaux, p. 287 (part. : female ; Santa Cruz de la Sierra, Bolivia).
Hab. Campos region of the interior of Brazil from Joazeiro on the Rio
Francisco, Bahia to Mattogrosso (Onyaba, Villa Bella, San Vicente, Oarandasinho,
Urucum), and thence to Santa Cruz de la Sierra in East Bolivia ; southwards
through Paraguay (Bohls coll. — in Mus. Tring et Berlepsch) to Corrientes, Argentina
(D'Orbigny coll. — specim. in Mus. Paris).
b. Saltator coerulescens azarae D'Orb.
Saltator Azarae D'Orbigny, Voyage, Oiseaux, p. 287 (part : descr. (J ; Mojos, Bolivia).
S. nuperciliaris (nee Spix) Sclater, Cat. Birds xi. p. 2'Jl (part. : Ecuador and Peru).
Hab. N.E. Bolivia (Moxos); Upper Amazonia: Central and Eastern Peru,
East Ecuador, Bogota coll. ; N.W. Brazil : Rio Jurna (Garbe coll. — Mus.
Berlepsch).
A third closely allied form occurs on the Lower Amazons :
c. Saltator coerulescens mtttiis Scl.
Saltator miitus Sclater, P. Z. S. 18.'i6, p. 72 ^Para].
S. superciliaris (neo Spix) Sclater, Cat, Birds xi. p. 291 [part. : Mexiaua].
Very much like S. c. azarae, but with the anal region and the crissum
considerably paler, ochraceous buff (instead of deep ochraceous).
Hab. Vicinity of Para : Mexiaua Island (Wallace), Oamolins (Steere coll. —
Mus. Tring).
11. Saltator albicollis Vieill. = S. guadeloupensis Lafr.
Saltator albicollis Vieillot, Xonr. Diet. xiv. (1817) p. 107 ['' Cayenne " — errore !].
Specimen in Mus. Paris, labelled: " Habia albicol, Saltator albicollis, Vieill.
(T.), de Cayenne, donn^ par M. Moussier." On the bottom of the perch : " donn(5
par M. Moussier, 1808. Salt, albicollis Vieill. type. Type do la description de
M. Ch. Bonaparte." Wing 94 ; tail 90; bill l8i mm.
Mr. Sclater* says : " I have examined tlie type of S. albicollis of Vieillot at
Paris, and believe it to be an immature bird of this species " (i.e. S. striatipictus
Lafr.).
This statement is quite erroneous, and, I su])pose, the examination must have
been a hasty one, for otherwise Mr. Sclater would not have i'ailed to recognise
the true afHnities of the bird. Vieillot's type is by no means immature, but an
adult bird in very worn plumage, and belongs to the species inhabiting the islands
of Guadeloupe, Santa Lucia, etc. It agrees with specimens from the latter island in
• Cat. Birds, xi. p. 295.
( 316 )
haviu"' the whitish superciliary streak continaed to the posterior end of the ear
coverts. This character alone wonld be sufficieut to jirove that the bird lias nothing
to do with S. striatipictus (= albicollls anct.), in wliich species the snperciliavies
always end above the eye. Moreover, the lower mandible is pale yellowish,
except for the blackish basal half of the sides, just as in a series of 16 specimens of
S. ynadeloupensis, while in 71 examples of S. striatipictus (= albicollis anct.)
the whole lower mandible is deep black, only in a few skins a very small
spot at the extreme tip being yellow. In Vieillot's type specimen, also the
whole apical half of the upper mandible is yellow, as in some examples from
Gnadelonpe, Santa Lncia, etc. Most of the specimens of S. striutipictiis have the
whole ujiper mandible deep black; sometimes there is a minute yellow spot at the
very tip to be seen, but never more I
After all, there can be no doubt that Vieillot's type is a sjiecimen of S. guade-
loupensis with wrong locality. Comjiared with some examples from Gnadelonpe
and Santa Lucia, it presents but very slight diflerences. The back appears rather
greyer, since the olive-green apical portion of the feathers is mostly worn off, but
on the head the green colour is well to be seen. The same applies to the under-
parts, which are rather greyer, less greenish, with the middle of the abdomen and
the under tail-coverts paler bnfty than usual.
The nomenclature of the two species which enter into the question is as
follows :
Saltator albicollis Vieill.
Stillator albicollis Vieillot, Nouv. Did. xiv. (1817) p. 107 [■' Cayenne "— errore !].
S. guadeloiipciisis Lafresnaye, Rev. Zool. 1844. p. 167 [Guadeloupe].
S. giiadelotqii-nsis Sclater, Cat. Birds xi. p. 295.
Hab. Lesser Antilles: Gnadelonpe, Dominica, Martinique, Santa Lncia,
Nevis.
Saltator striatipictus Lafr.
Saltator striiilipictus Lafresnaye, lier. Zunl. 1847. p. 73 [Caly, W. Colombia].
Saltator albimllis Sclater (nee Vieillot !), Cat. Birds xi. p. 294.
iS. albicollis auot.
Hab. Colo?nbici, North Vevezuda, Trinidad, Western Ecuador, North Peru.
12. Polioptila lactea Sharpe sp. ojit. !
Polioptila lactea Sharpe, Cat. Birds Brit. Mas. x. (1885) p. 45:5 [Hab. " unknown ": we substitute
S.E. Brazil, vicinity of Jiio],
1. Mus. Brit, ex coll. P. L. Sclater (<?) ad. "Sonth America.— Argent" Type
of P. lactea Sharpe. [The specimen is of the well-known "/^/o-makc."]— Wing 47 ;
tail 46; bill lOf mm.
2. Mus. Brit, tj ad. Rio de Janeiro. Alex. Fry coll. No. 263.— Wing 45J ;
tail 44; bill lOJ mm.
This is a very distinct species, and easily distinguishable from P. bilineata
(Bp.), with which I united it in the Tierreich, 18. Lief. p. 27, owing to lack of
material.
P. lactea agrees with P. bilineata in having the lores, cheeks, ear-coverts
and broad snperciliaries pure white, but differs at a glance by its much darker,
slaty-blue (instead of bluish grey) ujiper surface, entirely white outer tail-feather
without any black at the base, and beautiful creamy colour of the breast and
(31T )
abdomen. The sides of the body show uo trace of the bluish grey colour so
conspicuous in P. bilineata.
No. 2, a fresh skin, has the breast and abdomen of a delicate cream j'-yello wish ;
in the type, which is slightly faded, these parts are almost white. Tliere can be
no longer any doubt that P. lactea is an excellent species. It need.s no com-
parison with the other black-headed forms, since the broad white superciliaries
serve to distinguish it at once.
13. Curruca olivacea Less. = Vireo chivi (Vieill.) !
Sylvia chivi Vieillot, Xom-. Diet. xi. (1817), p. 174 [ex Azara ; Parai/wni].
Curruca oliracea Lesson, ia Voyage Coquille, Zoologie, Tome I. part ii. (1828), p. G64 [" I'ile de
Sainte Catherine au Bresil "].
Mus. Paris, spec. typ. (skin) labelled : " Lesson et Garuot. — Duperrey.
Fauvette, Ste Catherine dn Brdsil, No. 38. Si/hia olivacea Less., Brfisil, No. 72."
Wing 74 ; tail 56 ; bill 13^ mm. (tip broken oif).
This specimen, without doubt the type, agrees in every respect with a topo-
typical skin from Paraguay (Bohls coll.). The second primary is the longest, the
third and fourth hardly shorter ; the fifth about 4 J mm. less than the third and
equal to the first. As in the Paraguay example, there is no dusky malar streak.
Several times before I tried to make oat the species from Lesson's description,
but I never expected that it would turn out to be V. chivi !
14. Dacnis salmoni Scl. should be Nemosia salmoni (ScL).
Daciiis salmoni Sclater, Cat. Birds Brit. Mm. xi. (1880) p. 27, pi. ii. fig. 2 (Remedies, Antioquia).
L Mus. Brit. ?. Remedios. T. Salmon coll. Type of species. — Wing 58;
tail (moulting) ; bill 13 mm.
Hartert* has already remarked that this bird had nothing to do with Dacnis,
but belonged to the genus Nemosia. This statement seems to have been overlooked
by all later authors, for it is not even mentioned by Ridgway f and Oberholser.J
The careful study of the type specimen leaves not tiie slightest doubt that the
species is most closely allied to Nemosia cknjsomelas (Scl. & Salv.), which, though
originally assigned to Tachyphonus (! !), has since been jiroved by Mr. Ridgway §
to belong to that section of Nemosia separated by Cabanis under the name
Hemithraupis. In fact, both species are practically identical in structure ;
perhaps the bill in N. salmoni is a very little broader at the base, but otherwise
it is of the same shape. In coloration, however, the single known ? differs widely
from that of N. chrysomelas. Whether N. salmoni is really the ¥ of iV. rosenbergi
Rothsch. cannot be decided without knowing the S , but I notice that the type of
the latter species is much larger : wing 70 ; tail 53 mm.
15. Knipolegus unicolor Kaup.
This name was exclusively based on the description of Flucicola cyanirostris
Lafr. & D'Orb. from Corrientes. The Paris Museum still possesses the skin of au
adnlt male with D'Orbigny's original label, which reads as follows :
" Ada cyanorostris Nob. Corrientes. D'Orbigny, juillet 1829. No. 37." — Al. 80 ;
caud. 72 ; rostr. 13| mm.
• Nm. Zool. V. 1898. p. 484.
t Birds Xi'rth and Middle America, ii. 1902. p. 391.
+ Proc. U.li. Mus. xxv. 1902. p. 141.
§ Birdx Ntrrth aiid Middle Amerioa ii. p. 106.
( 318 )
Tbis specimen agrees in every respect with a good series of K. o/nnirostris
from Paragua.y and Sontii Brazil. It is a perfectly adnlt male iu black jilumage,
with the inner webs of the remiges narrowly margined with white. The allied
K. aterrimus Kaup. differs at a glance by having the whole basal half of the inner
webs pure white as far as the shaft. K. unicolor Kaup sinks, therefore, as a
synonym of K. o/anirostris. The A', unicolor auct. (nee Kaup) from the upper
Amazons, however, is a very different bird, without any white at all on the ciuills, and
is much nearer A', orenocensis Berl. The synonymy of these species is as follows :
Knipolcgus cyanirostris (Vieill.)
Muscicajm ojanirustriis Vieillot, Noiir. Diet. xxi. (1818) p. 447 [ex Azara, No. IHl. — Paraguay]
[= c? ad.]
jll. rufinipilla Vieillot, /. e. p. 459 [ex Azara, No. 178. — Paraguay] [= $].
Ada ciftntirnslris D'Orbigny, Vot/. .liner, iiieritl. Ois. p. :^»40 [Corrieutes]. \
Fbivicuhi cyanirostris Lafresnaye & D'Orbigny, Hijii. Ar. i. ia May. Znul. - uudfe :
1837. cl. ii. p. 5'J [Coirientes, rep. Argentina]. J
Ciiipolegus unicolor Kaup Jouru.f. Urnith. 1853. p. 29.
Hab. Paraguay (Azara) : Sapucay (Foster coll. — Mus. Tring) ; Corrientes
(D'Orbigny) ; South Brazil : Rio grande do >Sul, S. Paulo, Espiritn Santo (Mus.
Berlepsch).
Knipolegus sclateri uom. nov.
Cnipohgus tinicolor (nee Kaup) Pelzeln, Zur Ornith. Brasil. ii. (18G8) p. 99 [Rio Madeira, Brazil].
Cnipolegua unicolor Sclater, Cat. Birds Brit. Mus. \iv. (1888) p. 47 [Pebas, N.E. Peru].
<? ad. Nearest to K. oreiwcemis Berl. and without any white on the quill-
lining, but easily distingnishable by the consiiierably darker sooty-black colour of
the upper parts and much darker, brownish black lower surface. In A', orenocensis
only the top of the head is dull blackisli olive, the back olivaceous-grey, and the
lower parts are likewise paler, slaty-olivaceous.
Type in Mus. Vindob. No. 17591, S ad. -'Rio Madeira," W. Brazil,
November 18, 1829. — coll. J. Natterer.— Al. 7" ; caud. 69 ; rostr. 14| mm.
Hab. Upper Amazonia : Rio Madeira (Natterer) ; Pebas (Mus. Brit.).
16. Muscicapa cristata Less. = Knipolegus cyanirostris (Vieill. j ? !
Muscicapa cristula Lessou, Tniile d'Orn. (1831) p. 385 [without locality ; descr. mala].
Muscicapa crislata Puclieran, Arch. Mus. Paris, vii. (1855) p. 371 (crit.)
One specimen, mounted, without label, but with the following notes on the
stand : " Bresil, Mr. de St. Hilaire, aout 1822. Muscicapa cristata Less, type." —
Al. 73 ; caud. 70^ mm.
M. Pncheran {I. c.) already stated that this bird was a female of some .species
of Knipolequs. I was enabled to compare it with the females of the four sj)ecies
which came into the (pie.stion, and found it perfectly agreeing with that of
K. cyanirostris. It differs from a female from Rio Grande do Sul (Mus. H. v. B.)
only in having the lower parts less densely striated with blackish.
17. Muscisaxicola striaticeps Lafr. & D'Orb. should be Knipolegus striaticeps
(Lafr. k D'Orb.).
There is iu the Paris Museum a mounted specimen, labelled as follows :
^' Geositta striaticeps (d'Orb. et Lafr.). D'Orbigny. — -Bolivie " ; and its
stand bears the following inscription : " Bolivie, ( 'hiipiitos. D'Orbigny 1834.
I
( 319 )
No. 3921 = 1380. Mascisaxicola striaticei^s Lafr. &, D'Orb." — Al. 50 ; cauda 53 ;
tars. 1 8 ; rostr. 1 1 i mm.
It answers very well to the original description and to the plate in D'Orbigny's
work, and is no doubt the type of the species. No attempt having as yet been
made to identify the name, Conut Berlepsch drew my attention to it, snggesting
that it might prove to be the female of K. cinereus Scl. This supposition I
found fully confirmed by the examination of the type specimen, which agrees
perfectly with a female of this species in the Tring Museum. It may be described
as follows : Forehead and crown pale cinnamon-rufous with narrow blackish
shaft-stripes ; back pale greyish brown, upper tail-coverts pale cinnamon-rufous.
Tail-feathers Ijlackish brown, the inner web of the four onter pairs except at the
tip cinnamon-rufous ; the outermost with a distinct white margin along the onter
web. Lesser wing-coverts greyish brown, like the back ; median aud greater series
dark brown with large white apical spots forming two wing-bands. Quills dark
brown, primaries very narrowly edged with greyish, the inner secondaries with
broader white margins. Sides of the head huffy whitish ; lower parts whitish,
with pale greyish-brown shaft-stripes on the breast ; under tail-coverts cinnamon-
rufous, whitish at the tips. Axillaries and under wing-coverts bnfty whitish.
Bill brownish horn-colour.
Mus. Tring ; ? ad. Tapia, Tucuman, 000 m., December 2, 1902, coll. Baer. —
Al. 55 ; caud. 58 ; rostr. 11^ mm.
This specimen differs from the type only by its slightly longer tail.
The female of A'. striatici>ps bears a certain resemblance to that of A", qiani-
rostiis, but besides being very mnch smaller, it is easily to be distinguished by the
white onter web of the outermost tail-feather,'the white (instead of rusty-whitish)
wing-bands, the much paler greyish-brown back, aud by having only the breast
very indistinctly streaked with greyish.
The type in the Paris Museum is marked as coming from ( 'hiquitos,* viz. from
the lowlands in Eastern Bolivia. This seems much more probable than the locality
La Paz (in the high Andes), first indicated by Lafresnaye and D'Orbigny, the
more so, as the type of K. cinereus was obtained not far from the Bolivian
boundary in S.W. Brazil.
The synonymy of the species is as follows :
Knipolegus striaticcps (Lafr. & D'Orb.).
Muscisaricnla strialiceps Lafresnaye & Orbigny, Sy/i. Ad. i. iu Mnij. Zu'il. 18.37. cl. ii. p. CG [" La
Paz, Bolivia." — the type is marked " Chiquitos."] (= ? ).
Muscisiij-iciila strialiceps D'Orbigny, Vnijiigc Ois. p. 350, tab. 41, fig. 1 (= ? ).
Cnqj()le(jus cinereus Sclater, P. Z. S. I87U. p. 58 [Corumba, Mattogrosso] {= i).
Hab. Eastern Bolivia : Chi(|nitos (?) (D'Orbigny) ; San Miguel (Behn).
Mattof/rosso, Brazil : Corumba (L'ajit. Pago).
Argentina: Cordova (Doering) ; Salta (Duruford) ; Tucuman (Durnfoid,
Mus. Brit. ; G. Baer coll. — Mus. Tring).
* As I learn from a MS. not«, supplied by Count Curlcp-sch, Prof. Bclin got a female at San Miguel,
in tlie lowlands of Eastern Bolivia. It was preserved in the Kiel Museum, where it was examined by the
Count, and has passed probably into the Berlin Museum — loijctlier witli the otlier specimens of Behn's
collection.
21
( 320 )
18. Anaeretes sclateri Oust. = Hapalocercus sclateri (Oust.).
Anaerctcs sclateri Oustalet, Nuio:. Arch. Mus. Paris. (3) iv. (181)2) p. 217, descr. (J ad. ["Chili" —
errore !]
Uiijialimrciis hoUaniU Sclater, Ihis 180lj. p. lilT, descr. orig. ? [SantaElena, Argentina] ; Hellmayr
Verhamtl. zool. hil. Geseilsch . Wirii 1911.3. p. 204 [crit. — Pausecco, Mattogrosso].
H.Jfariventris Pelleln (nee Lafresnaye et D'Orbigny !), Orii. Brasil. ii. (1808) p. 103 [Pansecco].
Wing. Tail. Bill. Grad. of tail.
1. Mas. Paris. (J) ad. Typn of Anacrctt-.i
sclateri Onst. "adonis en 1837, du
Chili" 44 48 11 8 miu.
2. Mas. Brit. ? Sta. Elena, Argentina,
January 15, 1895. Holland coll.
'Syi^a oi H. hollamli ^c\. . . 4U 45 lOi C miu.
3. Mus. Vindob. No. 17809. c? ad.
" Pansecco, Mattogrosso, June 27,
1826." Natterer coll. ... 41 47 11 C,\ mm.
4. Mus. Vindob. No. 17810. ? ad.
" Pansecco, Mattogrosso, June 27,
1826." Natterer coll. . . .42 45 10 4i mm.
5 — 18. Mus. Triug. S3 ad. Ocampo and
Mocovi, near Ocampo, Argentina,
November 1905 and December
1903. Venturi coll. . . . 41—44 43—48 10-11 0—10 mm.
19—24. Mus. Triug. ? ? ad. Ocampo,
Argentina, November and December,
1905. Venturi coll. . . . 40—43 42—46 10—11 C— 8 mm.
In the paper quoted above I described the curious wing structure of the S from
Natterer's specimen.
Since that time I had the opportunity of examining the type of Anaeretes sclateri
Oust, in the Paris Museum, and found it to be an adnlt S of H. hollaruH, with the
fourth and fifth primaries exactly of the same shape. Latel)', the Tring Museum has
received twenty-two skins of this species, among them fourteen adult ? ¥ , all of
which show the same peculiarity. Onstalet's type and tlie series from Argentina
agree perfectly with the i from Pansecco in having the whole bill black and no buff
superciliary stripe. The lores, cheeks and sides of the head are dull black, the
feathers of the superciliary region with faint whitish edges ; the feathers of the
jiilenm are mnch elongated and narrowed towards the tip, forming a long pointed
crest, longitudinally striped with black and oehreous yellow. The back is green,
with slight dusky shaft-strijies, more apparent in the freshly moulted than in the
W(iru jilumage ; the median and greater upper wing-coverts have distinct bully
whitish apical margins; the whole undersnrface is bright yellow.
The females diifer from the males by the presence of a broad buff superciliary
stripe anil by having the lower mandible whitish. Natterer's specimen from Matto-
grosso and the series from Ocampo are practically identical with the type of
U. hollandi. None of the females show the peculiar shaj)e of the fourth and fifth
jirimaries, which are (|uite as long as the others, though somewhat narrower.
The range of this species, which ought to be called //. sclateri (Oust.), is as
follows :
( 321 )
East Argcnthm : Santa Elena, east of La Paz (Holland), Ocampo and Mocovi
(Venturi).
C. Brazil, Mattogrosso : Pansecco (Natterer).
Ohs. The original locality " Chili " is doubtless erroneous. Oustalet's type
formed part of a collection which contained, among others, Lcptastheniira platrnsis
(Rchli.), a species only known to occur in the eastern portions of the Argentine-
Republic.
A very near ally of 11. sclateri is //. striaticcps Salv.,* of which Imt a single
specimen, tlie type, is known. This is an adult c?, with the fonrtli and filth jirimaries
of exactly the same sha^ie, as in the SS ad. of //. sclateri. It is strange that this
})eculiarity has not been noticed by Salvin, who never referred to //. hoUandi when
describing the species, but compared it with 11. flaviventris, with which it has not the
least relations ! On comparing Salvia's type with our series of //. xclateri, I find
not the slightest diflerenco in coloration : bnt the bird from British Guiana has
a smaller bill with the under mandible whitish. Perhaps wings and tail are also
rather shorter,t bnt the specimen being in very worn plnmage, I would not lay
ranch stress upon this trifling variation. As 16 (J c? of the southern If. sclateri have
the bill entirely black, I think the pale lower mandible of the Aunai sjieeimen is
sufficient to warrant the recognition of a northern form : //. sclateri striaticeps Salv.
In the hills near Tncuman another species of the same group is found, the male
of which has the same curious wing-strncture as //. s. sclateri and //. s. striaticeps.
It dilt'ers, however, from both in the following points. The eye region and ear-
coverts are not blackish, but dull greenish, and the back is of a much duller, more
brownish green, without any trace of the dusky mottling. The most striking
diifercnce consists of the form and colour of the feathers of the pilenm. AVhile
the two forms discussed above have a long jwinted crest striped with black and
ochraceous, in the Tncumau species the feathers of the pilenm are of normal shape
and length (as in II. flaviventris) and uniform earthy brown, only those of the
forehead being narrowly edged with rusty buff. The Triug Museum possesses three
3 S ad. of this bird, which has been described by M. Lillo as a new genus, Pseiulo-
coloptcryx dinellianus. Its proper name is therefore Ilapalocercus dinellianus
(Lillo).
19. Leptopogou tristis Scl. & Salv. = Pliylloscartes ventralis augustirostris
(Lafr. & D'Orb.).
Miisciciqm (Diyust/rosti-is Lafresnaye et D'Orbigny, Si/ii. Av. i. in May. ZuvK 1837. cl. ii. p. 52
(Yungas, Bolivia).
Muscicapara angnntiroatrh D'Orbigny, Voi/at/e, Oinriinx, p. 325 (Yungas, de la Paz).
Lfjitopogoii (!) irislis Sclater & Salvin, P. Z. S. 187l>. p. 25-1 (Simacu, Yungas of Bolivia).
Wing. Tail. Bill.
1. Mus. Paris. Adult. Tyiie oi M.a/i(/Hstirostris
Lafr. & D'Orb. Yungas, Bolivie, D'Orbigny,
1834 ."JT 59 in. mm.
2. Mus. I'rit. Av.juv. Simacu, Bolivia. Buckley
coll. No. 0(j. Type of Leptopogon tristis
Scl. & Salv 54 54 1]| mm.
3. Mus. Brit. Adult. Samaii)ata, Bolivia. G.
Garlepp coll 551 58 12 mm.
* Jiull. B. U. C vii. (isil-) p. xvi. (Aun.ai, Brit. (Juiana).
t Wing 41-J- ; tail 45 ; bill lu mm.
( 322 )
Wing. Tail. Bill.
4. Mils. Brit. S ad. Cliacbajioj-as, N. Pern,
7300 ft., October 4, 1894. Baron coll. . 55 54 . 11| mm.
5 — 10. Adults of both sexes, Tncnman, N.W.
Argentine. Baer & Dinelli coll., Mns. Triug 51—55 54—60 111— 12 mm.
Having always snspectcd tbat tbere mnst be some mistake about Lcptopogon
tristis, I was not in tbe least surprised to find tbat tbis species bad notbing what-
ever to do with Leptopogoii, bnt was in every respect a typical member of tbe genus
Phylloscartes.
The type is an immature bird with some of the fluffy feathers of the juvenile
plumage still retained, especially on the nape and lower tail-coverts, but otherwise
it agrees in structure and coloration with my series from Tucuman.
When lately in Paris I was so fortunate as to discover among tbe skins in the
collection of tbe Museum d'Histoire Naturelle tbe type of Mnscicapa an(jnstirostris
Lafr. & D'Orb., a species which bad never been identified, although the original
description was sufficiently clear, tbe shape of the bill and tbe characteristic markings
of the wings being very accurately described. The type, an adult bird, is exactly like
the specimens from Tncnman. The S ad. from Chachapoyas does not differ either.
PliijUoscartcs ventralis anipistirostris (Lafr. & D'Orb.), as tbis form ought to
be called, is practically identical in all structural details with a large series of true
P. V. ventralis of Eastern Brazil, and differs but slightly in coloration. The upper
parts are of a duller, paler, more greyish green ; tbe lower ones rather paler yellow ;
the throat more mixed with whitish ; the yellow wing-bands as a rule a little
broader, and the forehead always washed with greyish. Thus tbere are two very
nearly allied forms recognisable, the range of which is as follows :
1. Phylloscartes ventralis -centralis (Temm.). Wood region oi S.E. Brazil: from
S. Paulo to Rio grande do Sul.
2. Phylloscartes ventralis angustirostris (Lafr. & D'Orb.) N. W. Argentina :
Tucumiiu ; Bolivia : Yungas of La Paz (D'Orbignj'), Simacn (Buckley) ;
iV. Peru : Chachapoyas (0. T. Baron).
N.B. — Tbe bird mentioned by Lafresnaye and D'Orbigny s. n. Muscicapa
ventralis (p. c. p. 53) is Capsiempis Jiateola (Lcbt.). There are two skins from
Guarayos in the Paris Museum, fully agreeing witli Bahia specimens.
Specimen h of Leptopogon tristis (^Cat. Birds xiv. p. 118) is widely different,
and represents a new species of Pogonotriccus, which will be described by
Count Berlepscb.
20. Leptopogon godmani Scl. = Pogonotriccus ophthalmicus Tacz.
Pnrjonolrircus ophlhahiiiciiK Tacziiniiw.ski, P. Z.S. 1874. p. 135 [Amable Maria, C. Peru].
Lcptopngnn godiiianl Sclater, P. Z. S. 1887. p. 48 [Saraya^u, E. Ecuador].
Mu8. Brit. Types of L. godmani Scl. Sarayaru, E. Ecuador (Buckley coll.).
No. 1. AVing 57i ; tail 52 ; bill 10 mm. ; No. 2. Wing 54^ ; tail 51 ; bill U) mm.
Tbis " sj>ecie8 " has nothing to do with LepAopogon, and is, indeed, very
different from all members of tbat genus, but on comparing it with a series of
P. ophthalmicus (9) from Central and North Peru, Ecuador and Bogotsl, I fail to see
any dillerence. The types are both slightly immature, as manifested by their short
and stout bills. Adult speciuiens have rather longer and more slender bills.
No. 1 agrees perfeitly with a ? from Pichincha, Ecuador, in shaj)e and size of
till' bill, while No. 2 has a rather broader and stouter bill. Three other examples
( 323 )
from Pichincha and Corazon, Eciiailor, liowever, are absolutely identical with a
topotype from Central Peru, thus proving that this divergency is of no geographical
significance. L. godmani Las, hence, to ue relegated to the synonymy of P.
ophthalmicus.
I may mention that the same difference in the shape of the bill between adult
and young birds is observable in many other genera of the Ti/rannidac, f. e.
Sublegatus, Xniojisaris, Suiriri (= Kmpithtqrct), Phaeomyias, etc.
21. Capsiempis caudata Salv. should be Serpophaga caudata (Salv.).
Capsiempis mudatii Salvin, Bull. Bill. Oni. CI. vii. no. 48 (1897) p. xvi [Ourumee, Brit. Guiana].
Serpopharja oreiincemiK Berlep.sch & Hartert, Nnv. Znnl. ix. (1902) p. 40 [Altagracia on the Orinoco
R., Venezuela].
Mns. Brit. " ? " ad. Ourumee, Brit. Guiana, October 22, 1890. H. Whitely jr.
coll. Type of C. caudata Salv. Wing 49 ; tail 48 ; bill lOf mm.
This bird is totally ditFerent from Cap.vi'mpis flaxeola, with which it was
compared by its describer, but belongs to the genus Serpophaija, and has since
been redescribed as S. oreiioce/mis by Berlepsch & Hartert.
Salvin's type agrees perfectly with some specimens out of the series collected
by Cherrie on the Orinoco. The Tring Museum has lately received several skins
from the environs of Paramaribo, Surinam.
The systematic jiosition of (S'. caudata has been fully discussed by Berlepsch &
Hartert in the paper quoted above. It is certainly a very near ally of Serphophaga
sub/lava 8cl. & Salv., and a larger series from Pard might even prove their identity.
Comparing the unique type of the latter with our series of S. caudata, I notice that
it has the chin yellow like the whole under-surface (not white), no buff suffusion on
the foreneck and no white at all on the tail. The rectrices, however, are extremely
worn, and I would not lay too mucli stress upon the absence of the white markings.
It is very unfortunate that the name 8. oreuocensis has to give way to that
of Salvin, who did not recognise the true position of the species.
22. Tyrannus tuberculifer Lafr. & D'Orb. should be Myiarchus tuberculifer
(Lafr. & D'Orb.).
TyrannuK tuhercidifer Lafresnaye & D'Orbigny, Syii. Av. i. in Mag. Zool. 18.37. cl. ii. p. 4.3
[GuarayoF, East Bolivia] ; D'Orbigny, Voyage, OUcaux, p. 307.
Mijinrdius tricolor Pelzeln, Ziir Orn. Brusil. ii. (1868) pp. 117, 182 [Rio de Janeiro and .Sapitiba :
S.E. Brazil].
Mi]iarchu:t r/nnilimstfis Pelzeln, Zur Orn. Brasil. ii. (1868), pp. 117, 183 [Villa Maria, Mattogrosso].
Jhjiarchui: coa/d Ridgwa-y, Pnif.. U.S. Nat. Mus. ix. 1886 (1887), p. 520 [" Orinoco "—male].
J/, tricolor + il. nigrkeps (part.: Colombia, Venezuela, Guiana & Amazonia) Sclater, Cut. Birds
xiv. p. 258.
Wing. Tail. Bill.
1. Mus. Paris, sjiec. typ. labelled as follows: " Mgiarchus
tuberculifer (D'Orb. & Lafr.) ?. Type. D'Orbigny,
Bolivie." On the bottom of the perch : " Guarayos.
M. D'Orbigny, 1834, No. 12(). Cat. gen. 373:<-. ?
Tgrannus tub&rculi/er D'Orh. & hafr. tyyc." . . 77 731 17 J mm.
2. Mus. Vindob. No. 18382. " <S alt " Sapitiba, March 28,
1818. Natterer coll. ly^e o{ M. tricolor Fe\z. . ~U 69 17 mm.
3. Mns. Vindob. No. 18383. " ? alt " Rio de Janeiro, Decem-
lier 13, 1!SI7. Natterer coll. Type of M. tricolor Pelz. 69^ C3 17J mm.
(324)
Wing. T.iil. Bill.
4. Mus. Vindob. No. 1S3S4. " S " ail. Villii Maria, Matto-
grosso, August ^4, 1S2^). Natterer coll. Type of J/.
f/)-ar/liro.s/r>s I'cV/,. . . . . . . . .So! Tl^l Ifi mm.
5. Mus. Vindob. No. 1S392. " c? " ad. Borba, Kio Madeira,
March 1, 1830. Natterer coll 80i
C). Mns. H. V. Berlejiscli. Baliia-skiii . . . .714
7. Mus. Tring. " ? " ad. Igarape Assu, I'an'i, January "Jo,
1904. A. Robert coll. No. It) 39 . . . . 7Z
Count Bcrlepsch * ba.s already suggested tliat 7\ tuhc.rculij\:
to be the same as M. tricolor, and the comjiarison of the types uf both species
convinced me of the correctness of his determination.
I'elzelu distinguished a specimen from Mattogrosso s.u. M. gracilirostris on
account of its smaller, narrower bill, less dusky cap, and larger size. The type is
now before me, and I find a specimen from Borba in every respect identical. Both
e.Nami>les differ from the types of J/, tricolor ex Eio as well as from two skins from
Bahia and Para in their longer wings and tail, narrower bill and brighter yellow
belly. IhniiyiiC oi M. tuberctilifcr {ay: Guarayos, East Bolivia), however, has the
bill quite as broad and long, and the belly as pale as those from East Brazil, while
in the dimensions it agrees with M. ffracilirostria.
The coloration of the cap is very variable. In the male type of ^f. tricolor it
is decidedly sooty blackish, in the female type dusky, in Nos. 1, 4, 5 and 6 scarcely
darker than the back. In a series of fourteen adults from Cumana the same
variation is to be observed.
Specimens from British Guiana, Cumana, Trinidad, Orinoco, Bogota and Eastern
Ecuador agree in dimensions and colour perfectly with the tyjies of ^1/. (jracilirostris
and M. tuberculi/er, and belong certainly to the same form. There remains only
the (piestion whether the form inhabiting Eastern Brazil (from Kio to Para) is
always distingnishal)le by its slightly smaller size, in which case it ought to be
called M. tuberculifer tricolor Pelz. I notice, however, tbat one specimen from
Paramaribo, Surinam, is scarcely larger than the latter (wing 74 ; tail 074 ; bill
17 mm.).
There can be no longer any doubt that M. tuberculifer and M. gracilirostri.s
are synonymous, and have nothing to do with M. atricejjs Cab., which is a much
larger bird, with a deep black cap. It may be added that the specimens m—y
of M. " nifjricejts " apud Sclater {C'nt. B. xiv. p. 258) are absolutely indistinguish-
able from the types of M. tuberculifer and M. gracilirostris, while .1/. iiigriceps of
West Ecuador and North Pern (Cajabamba, Cutervo, etc.) is readily known by its
deep black head.
23. Xenopipo subalaris Godm. = Chloropipo unicolor Tacz.
Chlorojiipn unicolor Taozanowski, Orn. riroit ii. (1884), p. 335. descr. orig. ? [Amable Maria, Peru
ccntr.] ; Berlepsch & Stolzmann, P. Z. .S. 189(5. p. aiiS (descr. ? ; crit.) [Garita del Sol, Peru
centr.].
Xenopipo siibaUiris Godman, Bull. Biit. Orn. CI. x. no. Ixvii. (December 189'.!), p. x.wii descr. orig.
cJ ? [Guajabamba, Peru sept.].
Wing.
1. Mns. Brit. " t? " imm., Gnayabamba, N. Peru, 14. ix. 1894 76
•J. Mns. Brit. (?) imm., Gnayabamba, N. Peru, 20. viii. 1894 72
[Types of Xenopipo subalaris Godm.]
* Journ.f. Ornith. 1884. p. 304, and Ibis, 18S3. p. 141.
( 325 )
WiDg. Tail. Bill,
3. Mils, liraiiicki. " ? " ail, Gaiitiidel Sol, t!. Fern, July 8,
1891. Kaliuowski coll. ...... 75 50 13 mm.
[Compared by Dr. Stolzmauu with the typo of G. unicolor
Tacz. in the Warsaw University Mnseiim.]
Through the kindness of Dr. Stolzmann I was enabled to examine speiiintMi
No. 3, and fonnd it to agree with tiie female of Godman's Xenoiiipo nubalaris,
excejjt that the bill is rather longer and the crown decidedly darker green. These
differences, however, are very slight and well within the range of the individual
variation to lie observed in the allird species, and I have not the slightest doubt of
their identity. It is hardly to be understood iiow Mr. (lodman could associate this
bird with XcDOpipo atronitens Cab., to which the male has only a superficial
resemblance in being of the same glossy black colour. X. atronifcits is well
characterised by its broad, flattened mesorhiuium. C. unicolor, on the other hand,
has a much narrower and qnite differeutlj-shaped bill, with the culmen distinctly
ridged, like ChloropriJoflaacaiAlUt. The proportions of the wings and tail are the
same as in the latter species, with which it also shares the presence of a large tnft
of soft silky white feathers on the sides of the breast. In coloration, however, it
is altogether different from the other species of Chloropipo, showing the same
sexnal difference as Xenopipo atronitens. Mr. Sclatei * put G. unicolor as a synonym
of C. uniformis from British Guiana, but Count Berlej-sch and Mr. Stolzmann {I. c.)
have already pointed out the differences existing between these two birds. The
species has thns to stand in future as Vhlorojiipo unicolor Tacz. It apjiears to
be an inhabitant of high elevations. 0. T. Baron collected it near Guayabamha,
at an altitude of 4000 to 5500 feet ; Kalinowski obtained his sijecimen at Garita
de Sol, 11,000 feet; and Jelski found the type at Amable Maria, a little over 200IJ
feet. These localities are the only ones as yet known.
24. Chloropipo holochlora Scl.
Cat. Birds Brit. Mus. xiv. p. 287 [type ex Bogotd coll.]. <
The type in the British Mnseum is perfectly similar to specimens from Eastern
Ecuador and N.E. Pern (Chyavetas). The upjier jiarts are of a deep sinning grass-
green, the throat, chest and sides of the breast rather duller, more greyish green, and
the middle of the abdomen is clear pale yellow. Axillaries and under wing-coverts
dull greenish grey.
The specimen from I'asto f (spec, b of iSdater's list), however, is very diflerent,
and agrees in coloration with a large series from various places in Northern
Ecuador. I propose to call this form
Chloropipo holochlora litae n. snbsp.
Similar to G. It. holochlora, but easily known by having the whole upper
snrface pale olivaceous green (instead of deep shining grass-green).
Type in Mus. Tring : No. 143. " $ " ad. Lita, N. Ecuador, 3ii0n ft., September
22, 1899. Collected by Mr. Miketta, one of Mr. Rosenberg's correspondents.
Wing 70i ; tail 47 ; bill 12 mm.
I examined more than 20 specimens from the following localities, all in N.W.
Ecuador : Cachabi, 500 ft. ; Ventana ; Lita, 3000 ft. ; Paramba, 35i)(i ft.
• Cat. Birds Brit. Mus. xiv. p. 417.
t Pasto is not iu Ecuador, as (juotcd in tlie Cat. Binh, but in Colomliia.
( 326 )
There is no iliffereuce iu colour between the sexes, the females being only ii
little smaller: dS, wing 70—72; tail 47—49 mm.; ??, wing 65—67; tail
42 — 44 mm. From C. uniformis it differs chiefly in its much smaller size and in
having the axillaries greenish grey instead of yellowish white.
The various forms of C///oro/>ipo can be distinguished by the following key : —
A. Plumage glossy black, axillaries and a large tuft of
silky feathers on the sides of the breast snow-white C. unicolor <?.
B. No black whatever in the plumage.
«'. Top of the head bright yellow, breast and abdomen
jialer yellow G._flavicapilla S ? .
a-. Head green like the back.
i'. Axillaries and a large tuft of silky feathers
on the sides of the breast pure white . . C unicolor ? .
b-. No white tuft on the sides of the breast ;
axillaries not white,
c'. Larger (wing 78 — SO ; tail 55 — 58 mm.) ;
axillaries pale yellowish, greyish in
the middle C uniformis <? ? .
C-. Smaller (wing 65 — 72 ; tail 42 — 49 mm.);
axillaries greyish green
r/'. Upper surface bright grass-green . C. h. holocklora 6 f .
fP. Ujiper surface {)ale olivaceous green C. h. litae <S ? .
25. Muscicapa luteocephala Less, should be Heterocercus luteocephalus (Less.).
Muscicapa lutrocrphala Lesson, Traife d'Oni. (1831), p. 392 [no locality]; Pucheran, Arch.
Mils. Paris, vii. (1855), p. 374 [crit. — "rAmerique meridionale "].
MiTs. Paris.— etiq. : " Moucherolle, Amerique meridionale." On the bottom of
the stand I find the following notes iu Pucherau's handwriting : " Muscicapa
luteocephala Lesson (type) de I'Amc'rique meridionale." Wing 7U ; tail (moulting)
44 ; bill 111 ; graduation of tail 25 mm.
This is a very distinct species oi Heterocercus, agreeing in the form of the bill
and in the peculiar shape of the tail with the other species, from which, however, it
is easily recognisable liy its widely different coloration. That of the uuder-parts
reminds one of Seotothorus chrysoccphalus (Pelz.) [= igniceps Scl.]
Upper surfiice jiale olive-green ; top of the head and uajie ashy grey, middle
of the crown briglit Icmon-yclIow ; lores whitish, cheeks and ear-coverts pale
olive greenish. Throat and chest dirty whitish, the latter with a slight brownish
hue ; rest of under-surface very jjale suljihur yellowish. Upjier wing-coverts, quills
and tail-feathers dark brown, exteriorly edged with olive-greeuish. Bill pale
brownish, lower mandible almost whitish.
As will be seen from the above description, it differs from the other known
species in having the forehead and sides of the crown ashy grey, and the posterior
l)arts of the lower surface pale sulphur yellow. In structure it agrees perfectly with
ll.Jlnvixcvtcx Pelz. The tarsi are short and slender, the bill of exactly the same
form, and the tail, although not full-grown, shows the characteristic reduction of the
onter rcctrices, the outermost pair being short, attenuated, and entirely hidden by
the under tail-coverts. The middle pair has not yet reached its full length, hence
the tail ajjpears shorter than in the allied species.
(327 )
It is only fair to state that M. Oiistalet had already recognised the true position
of this remarkable bird, as I learn from a M8. note of his on the bottom of the
i-tand.
The iyyiG specimen has no locality, but M. Onstalet suggested that it might have
come from the interior of Cayenne, which is almost entirely unknown.
The following key to the species of Huterocefcus may be acceptable to
ornithologists :
A. With a brightly coloured crest (t?c?)-
a. Forehead and sides of the crown very different
from the olive-green colour of the back.
a'. Crown-patch fiery red, forehead and sides
of crown black ; breast chestnut . . Il.Unteatusi^incW.)
a". Crown-patch lemon-yellow, forehead and
sides of crown ashy grey ; breast and
abdomen pale sulphur yellow . . //. b.iteoceplialns (Less.)
b. Forehead and sides of the crown olive-green
like the back,
a'. Crown-patch golden yellow ; breast
chestnut-brown, decidedly darker in
colour than the abdomen . . . H. jlamvcrtex Yeiiz.
a^. Crown-patch orange, breast and abdomen
of a uniform pale ochreous reddish . H. aurantiivertex Scl.
& Salv.
B. No bright crest ( ? ? ).
c. Throat white with a slight isabelline tinge ;
upper parts dull brownish olive-green . . //. linteatus (Strickl.)
(I. Throat pale grey ; upper parts pure olive-
green ........ H.Jlavimrtex Pelz.
26. Muscicapa luteocephala Lafr. = Neopelma aurifrons (Wied).
Musc'icapa aurifrons Wied, Beilr. Xnturg. Brazil, 3. ii. (1831), p. 829 [" Camnmu and Bahiii''].
Muscica2M hiteocephalii. (nee Lesson 1831 !j Lafresnaye, Marj. Zoid. 1833. cl. ii. tab. 13.
Mus. Paris, etiq. : " Piprotnorpha luteocephala, Bresil, capt. des Mines,*
M.S. Hilaire, aodt 1822. Muscicapa luteocephala Lafresnaye." AVing 68^; tail
62 ; bill 10 mm. Not different from specimens from S. Paulo.
As pointed out by Pucheran, the " Muncicapa luteocephala " of Lafresnaye is
very diiferent from Lesson's species. Lafresnaye's name has been quite correctly
given as a synonym of ^1/. aurifrons in tlie Cat. Birds, xiv. p. 323.
27. Psaris erythrogenys Selby should be Tityra inquisitor erythrogenys (iSelb) ).
Psaris er/jtliriii/riii/.'s Selby, Znnl, Jnurii. ii. uo. viii. (January — April 1S2IJ) p. 483 [Pernjimbuco. —
Mus. Paris].
Titijra crtjthrogcmjs Berlepscli & Hartert, Nnr. Znal. i.v. (^1002) p. .55 [Orinoco ; crit.].
Mus. Paris. Spec. tyj). (?) ad. labelled: '' Titijra inquisitrix (Liclit.) ; ?.
J. Verreanx, Bresil," and on the bottom of the stand I find the note: "de Pernambuco,
fichange b. M. Verreaux, 1823. Brt^sil, no. 3593," Wing 04 ; tail 62 ; bill
2U mm.
This is undoubtedly the type of Psaris erythrogenys. Selby says expressly
that he described an example from Pernambuco in the Paris Museum, and
• Minas Geraes.
( 328 )
the above specimen agrees in every respect with liis account. Berlepscli
and Hartert have alreaily rccogiiiseil the distinctness of this form, and its characters
are fnlly discussed I.e. The type in the I'aris Museum is perfectly identical
with a female from Kio Catauapa, Orinoco (Tring Museum). The back is of
a clear jnile grey (without the luownish suft'nsion always to be seen in females
of 7'. /. inquisitor), and wings and tail are iiuite as short. Tiie males of the two
forms differ only in size, those of the soutluTU race being invariably larger. The
two subsjiccies have to stand as follows :
a. Titi/ra i/iijtii.'<itor iz/'jiiisi/or (Lcht.).
Laiiiiis iiiijiiisilor (v. Olters MS.) LichtensteiD, Ver:. Ilnhl. 1823. p. 50 (San Puuki).
Hab. S.l'^. Jlra.:il : S. Paulo, Minas, and Bahia.
MKASURE5IENTS.
S ad. Alambiiry, S. Paulo. Wing 114 ; tail 71^, mm.
c? ad. Victoria, S. Paido. Wing 112 : tail (js mm.
2 (Jc? ad. Rio Jordao, Minas Geraes. Wing 115, 112 ; tail 6b mm.
t? ad. S. Francisco, Minas Geraes. Wing 111 ; tail 73 mm.
?. Yi)anema, 8. Paulo. Wing 10(3 mm.
?. Victoria, S. Paulo. Wing InG ; tail O.s mm.
2 ? ?. Bahia. Wing 103, 104 ; tail 68 ram.
b. Tifi/ra inquisitor eri/throgenys (Selby).
Hab. JS.H Brazil : Pernamlmco (type) ; Surinam : Paramaribo (Mus. Tring) ;
Venezuela : Maipnres, Perico, and Rio Catauajia on the Orinoco River, Suapure on
the t'aura River, Lagunillas near Merida ; Colombia : Bogotd coll. ; East Ecuador :
Archidona (Goodfellow coll. — Tring Museum).
MEASUREMENTS.
5 ad. Maipnres, Orinoco. Wing 100; tail 03 mm.
(? ad. Suapure, Canra. Wing 102 ; tail 6u mm.
6 ad. Paramaribo, Surinam. Wing 98 ; tail 6U mm.
c? ad. Bogota coll. Wing lOoi ; tail 04 mm.
2 ? ?. Orinoco River. Wing OH, lUO ; tail 61, 05 mm.
? . Snapure, Caara. Wing 99 ; tail 64 mm.
? . Lagunillas, Venezuela. Wing 99 mm.
2 ? ? . Bogota coll. Wing 99, 100 ; tail OOi, 6.') mm.
2 ? ? . Paramaribo, Surinam. Wing 97, 99 ; tail 03 mm.
?. Archidona, E. Ecuador. Wing 96 ; tail OU mm.
28. Attila brasiliensis Less.
Atlilu birisiliemis Lesson, Tr<ul<< d'Orn. (1831) p. 300 [" Du Br&il "] ; Pucheran, Avrli. Mus. Pai-is
vii. (1855) p. 3G0. [Typical locality stated to be Cayenne, and not Brazil.]
Mus. Paris. Spec, typ., mounted, labelled as follows: "Attila brasiliensis
(Less.). Type. Cayenne," on the bottom of the stand : " No. 3847. Aftila
brasiliensis Less., type."— Wing 8.5i ; tail (missing) ; bill 2U mm.
I am not able to say whether Lesson's bird is really the same as ^1. iirop;/gialis
(fall.),* having not yet examined the type of the latter; but it is nndoubtedly
identical with the sjiecimens obtained by the late IL Whitely in British (Juiana,
and referred to by Jlr. Sclater as ^L vropygialis.i
* Oaxi/crj/liala iiri>i>ijghdis Ciljauis ; iu Scliomburgk, Ikise Brit. Guiana iii. (IS-lf*) p. 686.
t Sclater, Cat. ISirilx xiv. p. 300.
( 329 )
On comparing a series of seven specimens from Camacnsa anil the River
Carimang, we notice consideral)le variation in the colour of the back, wliich, as
fiir as I can see, does not in any way depend on sex, as supposed by Mr. 8clater.
In two specimens (marked " S " and " ? " by the collector) head and back are
uniform pale olive-green, only that portion of the lower back adjoining the snlphur-
yellow rump, being fiiintly washed with pale rufescent. The same coloration of
the upper parts is shown by the type of Lesson's A. brasiUensis. Another male
from the River Carimang has the entire middle portion of the back rather more
distinctly sntfnsed with rufescent-brown, while in a female from Camacnsa the whole
mantle is almost uniform dark rufescent-brown. The other extreme is reiiresented
by a female from Camacnsa, which has the back of a uniform dull greyish green
without the slightest trace of the brownish admixture. Exactly the same variation
is shown in a series from the Caura River, Venezuela. Two specimens (both
marked " S " by the collector) have not only the back, but also the head (including
cheeks and ear-coverts), strongly suffused with russet. Some peculiarities in
coloration — viz. the broader and more rufous apical margins on the wing-coverts,
the rusty suffusion on the sides of the body, rump and forehead, etc. — clearly
indicate their immaturity ; and I have, therefore, little doubt that the russet-backed
specimens are the young, the green-backed ones the adult birds.
As to Lesson's type, there may be further stated that it agrees in coloration
of the under-parts with a female from River Carimang. Throat and foreneck are
dull grey with narrow whitish longitudinal streaks, the middle of belly white, sides
and crissum pale yellowish. In other specimens, throat and chest are decidedly
olive-green, streaked with pale yellow ; but there is every possible intergradation
between these two extremes in the series before me.
In South-east Brazil occurs a closely allied form which presents the same
variation regarding the colour of the back, and differs from its northern repre-
sentative only by its considerably larger size. It ought to be called Attila
brasiUensis uropygiata (Wied). One of the specimens in the British Museum
(" Rio "-make) ex coll. Sclater is marked as having been compared with the type
of ^1. brasiUensis in the Paris Museum. As a matter of fact, however, the latter
belongs to the smaller race found in Guiana, and the locality " Bresil," assigned
to it by Lesson, had already been corrected by Pncheran, a statement that seems to
have escaped the notice of Mr. Sclater. It is very unfortunate that a bird which
does not occur in Brazil should be called " brasiUensis" but there is no way to avoid
this regrettable change of nomenclature.
These two forms have, consequently, to stand as follows :
1. Attila brasiUensis brasiUensis Less.*
Altila bnisUh-nah Lesson, TralU (TOrn. 1831. p. 36n [" Br&U "— errore ! Tbo type came from
Cayenne] ; Salvin, Ibis, 188.5. p. 30.
A. urvjnjgialis Sclater, Cat. Binln xiv. p. 360 (Brit. Guiana).
MEASUREMENTS.
3 t?c?. Brit. Guiana. Wing 85—87 ; tail ()5— 67^ mm.
3 ? ?. Brit. Guiana. Wing 8(1—81 ; tail 60—62 mm.
2 Sd (with green back). Caura River, Venezuela. Wing 84, 87 ; tail 67 mm.
* A. vii-idescem RUlgWiiy, P. U. S. Mns. x. 18S7 (18HS) p. 522 [type ex Santarem] belongs, most
probably, also to A. i, braiiliemw ; I have not been able, however, to examine a specimen from the
typical locality.
( 330 )
1 ? (l)Mck greeu). t'aura River. Wiug ^.j ; tail M mm.
2 SS (l)uck rasset). ('aura River. Wing 8;"), 80 ; tail (57 mm.
1 cJ. Salto Girao, River Madeira, Nattercr coll. (back rusf;ct). Wing 85;
tail 70 mm.
Hab. Cayc)i))C; British Guiana; Venezuela: Canra River ; \V. Brazil: Salto
do Girao, Rio Madeira (Natterer coll.).
Obs. Dasi/cephala uropygialis Cab. may perhaps be the same as Attila f^p'ulicea
(Gm.) [type ex Cayenne in Mns. Vindob. ex Mus. Leveriano].
2. Attila brasilie/isis uropygiata (Wied).
Musiiciipa nrnpiiriiafa Wied, Bc'ilr. Nalurri. Driisil. .3. ii. (1831) p. 8r,8 [Rio Doco, Espiritu Santo
S.E. Brazil].
Attila lirasilieiiKiH (nee Lesson !) Sclater, Cut. Binl.i xiv. p. 359 [S.E. Brazil].
MEASUREMENTS.
Three Bahia skins. Wing 90, 92, 97 ; tail 68, 71 i, 70 mm.
One adult, " Rio "-make. Wing 95 ; tail 74i mm.
Hab. S.E. Brazil : Rio Doce, Espiritu Santo (Wied coll.) ; Bahia. In the
British Museum is a skin of the well-known " Rio "-make.
29. Attila bolivianus Lafr.
In a paper published in the VcrhanJlungen dcr h. k. zool. hotan. Gesellsckaft
of Vienna, 1002. p. 97, 1 pointed out that A. ralidm Pelz. was obviously synonymous
with A. holicianus Lafr. Last year I had an opportunity of inspecting, in the
Paris Museum, some of Lafresnaye's original examples, and found my surmise to
be quite correct. There are three skins of D'Orbigny's in the French National
collection, labelled as " D. 367. Gnarayos, D'Orbigny 1834. Tyrannus nt/escri/s
nob." They agree in every respect with an adnlt male, collected by Natterer
in Central Brazil (typical A. validus), their identity being thus established beyond
doubt. The synonymy is given in my paper I. c, and need not be repeated here.
D'Orbigny's specimens measure as follows :
No. 1. Wing 95 ; tail 83 ; bill 22 J mm.
No. 2. Wing 98 ; tail 83 ; bill 24 mm.
No. 3. Wing 94A ; tail 80 ; bill 23 mm.
The original specimens of A. validus in the Vienna Museum, ~ S S ■■
Wiug 94-lOU ; tail S3— «9 ; bill 22—25 mm.
3(1. Lanius uninifiis Puch. = Attila thamnophiloides (Spix).
Musi-icapa tliiniimiiliiluidcs Spix, Ar. Bins. ii. (IKL'5) p. VJ. lab. xxvi. fig. 2 [" in locis sylvaticis fl.
Aniazinuaitl.
Lanius uiiirufus (Cuvier MS.) Puchcran, An /i, .Uus. Paris vii. (1855) p. 332 [Cayenne].
1. Mus. Paris. Spec, typ., labelled as follows : " Attila thamnophiloides,
Lanius unirufus (Cuv.) Type. Cayenne, 3640." Wing 88 ; tail 82 ; bill 21 mm.
2. Mus. Monac, av. jr. " Amazonas. Spix coll." Type of M. thamno-
philoides Spix. Wing 90 ; tail 89 ; bill 23 mm.
Pucheran's type differs from No. 2 iu its shorter tail and narrower, shorter bill.
A series from Borba, however, varies iu the length of the tail from 80 to 87, and
in that of the bill from 20 to 22A mm. Both types are somewhat faded, and
look, therefore, paler than fresh skins. In No. 1 tlie whole lower surface is pale
ochraceous ; in No. 2 only the breast and abdomen are of this colour, while the
(331 )
throat is decidedly darker, more ferrnginons. It may be added that a good
series of skius from Surinam, lately received in the Tring Mnsenm, is iu no way
distingaishable from Amaznnian specimens.
31. Upucerthia bridges! Scl. = U. andaecola Lafr. & Orb.
Uppucerthiii amlaerola Lafresnaye & D'Orbigny, Si/n. Av. ii. in 3fug. Ziml. cl. ii. (1838) p. 21
[" La Paz, Sicasica, rep. Boliviana "].
Vppucerlhia amUcoht D'Orbigny, Vojjarje, etc., Oiseaux, p. 371. tab. 50. fig. 2.
Upucerthia brkhjesi Solater P.Z.S. 1889. p. 32 [Bolivia (Bridges)].
a. Mns. Paris. Tyjjes of U. andaecola Lafr. & D'Orb., three specimens with
D'Orbigny's original labels, which read as follows : —
Wing. Tail. Bill.
No. 1 (mounted). " La Paz. D. lOG. D'Orbigny,
1834.— 378" 85 79i 10 mm.
No. 2 (skin). "378. D'Orbigny, 1834. D. 196. La
Paz " 70 72 26 mm.
No. 3 (skin). " D. 196. Sicaca * 378. D'Orbigny,
1834" 77 76 25i mm.
h. Mns. Brit. Types of U. hridgesi Scl.
No. 1. adnlt, ex Mus. T. C. Eyton. Int. Bolivia,
Bridges 80 72 2.5 mm.
No. 2. av. juv. Bolivia — Bridges .... 77 75 201 mm.
U. brklgesi is merely U. andaecola redescribed. By neglecting the typical
locality. Dr. Sclater considered the Peruvian U. serrana the same as U. andaecola,
while some Bolivian skins (typical andaecola) were referred to a new species 1
Through the kindness of Mons. Onstalet, I was enabled to compare two of
D'Orbigny's types (Nos. 2 and 3) with the original specimens of U. Ijrid(/csi. As
was to be expected from the localities, they are absolutely identical.
U. serrana Tacz., however, is a very distinct species, easily known from
U. andaecola by its dark earthy-browu head and mantle, without any rufous tinge,
ljull'y-white strijies on the forehead, and by the pale striatious on tiie foreneck
and breast.
The synonymy of the two species is the following :
1. Upucerthia andaecola Lafr. & Orb.
For original references see above. — U. andecola Scl. & Salv. P. Z. S. 1879. p. G44 (ex d'Orbigny).
Upitcerthut hridgesi Sclater, Cat. Dinh xv. p. 19.
Hab. Andes of Bolivia : La Paz, Sicasica (D'Orbigny) ; interior of Bolivia
(Bridges).
I have seen many specimens in Count Berlepsch's collection.
2. Upucert/tia serrana Tacz.
U. serrana Taczanowski, P.Z.S. 1874. p. 525 [C. Peru]; idem, P.Z.S. 1880. p. 20 [Cutervo,
North Peru]; idem, Urn. Pa-oii, ii. (1884) p. 107 [Peru] ; Salvin, Nm: Zwd. ii. (IH'.)u) p. 13
[North Peru : Cajamarca, nu.amachuco, Cajabumba] ; lierlepsch & Stulzmann, /'. Z, S. IS'.KJ.
p. 371 [Central Peru : Palcamayii, (Jueta].
Upucerthia (indicula (nee D'Orliigny Ai Lafresnaye !), Sclater, Cut. Birih xv. p. 19 (Cutervo,
Maraynioc : Peru].
Hab. Central Peru : Jiuiin, Acancdclia, Monterico (.lelski coll.); Palcamayo,
Quota (J. Kalinowski coll.). Xoitk Peru: C'utervo, 9i)UiJ ft. (Stolznianu coll.);
• A| part-nlly pen-slip fur Sirtisica.
( 332 )
Cajamarca, 11,000 ft.; Hnamachnco, 10,400ft.; Cajabamba, 9000 ft. (0. T.
Baron coll.).
In the Tring Mnsenni tliore is a good series of this species from North aud
Central IVru (Barou and Kaliuow.ski coll).
32. Synallaxis setaria Teram. = Leptasthenura setaria (Temni.) sp. opt. !
Sy)mlh.ri.i sihiiiit Temminck, I'l. inl. livr. 52. tab. .311 tig. 2 [November 1H24— " .lu Bri'sil dans la
capitainerie de S.iint Paul.— Miisi'e dc Paris "]. Cf. Mi'nc'gaux et Ilellmayr, J/e/«. Sue. liht.
mil. AuliiK xix. {VJM) p. G8 (crit.).
1. Mus. Paris (monnted). Adult, lalielled as follows : " Mr. St. Hilairo,
Br^sil, prfes Casto, capt. de Saint Paul. Type de I'espece, PI. col. 311. lig. ~.
Synallaxe a filets." AViug 501 ; tail lO."") ; grad. of tail 70 ; bill 12i mm.
2. Adnlt, labelled : "Mr. St. Hilaire, Mines, cajit. de Saint Paul, 1.^22. Type
de I'espece." Wing 56 ; tail 118 ; grad. of tail 90 ; bill 121 mm.
This is a very distinct sjiecies of Leptastlicnura, and, as far as I know, tlie two
sjiecimens are the only ones in existence ; at least, 1 never met with any other
e.xunijile. Temmiiick's jihite gives a fairly good idea of the bird ; but tlic tail,
although well described in the letterpress, is not correctly coloured.
Whole top of the head blacki.sh, each feather with a sharply defined whitish
shaft-streak ; back nuiform bright rnfous-brown, rump a little jialer. Upper
wing-coverts and remiges blackish brown, exteriorly broadly edged with the colour
of the back'. Bend of the wing, axillarles, and under wing-coverts pure white.
Tail-feathers brigliter than the ujijier parts, more ferruginous, the three median
jiairs with a blackish brown margin along the inner web. Lores and distinct
eyeiirow white ; ear-coverts white with dark brown shaft-lines ; sides of the neck
greyish witli indistinct whitish streaks ; throat and jngnlum white, slightly freckled
with greyish ; middle of breast and abdomen dirty whitish, sides of body pale
bufl'v brownish. Inner webs of the remiges edged with fawn-colour. Bill blackish,
basal half of lower mandible yellowish white.
The feathers of the head are somewhat elongated, so as to form a sort of
loose crest. Tail much graduated, the central pair of rectricea strongly emargiuated
ou the inner web, distinctly rounded at the tip, not pointed as in the other species
of Leptasthenura, and about 28 mm. longer than the next.
L. setaria stands quite by itself, and needs comparison only
with L. J'uliginiceps, which also has the tail rnfous. It differs,
however, from the latter in having the head blackish striped
with white (instead of uniform pale brownish rnfous), the back
bright rufous (not pale earthy brownish), a very distinct white
eyeltrow and quite (liU'erently coloured nnder-parta. The bill is
also mnch longer and the basal half of the under mandible
yellowish white (in A. fiili<iiiiice/is the whole lower jaw being
(lark horn-colour).
,..,ji . -1 r .1 The locality ^^ Casto" is evidently meant for Castro, in the
Middle tail-feallicr •' •' '
ot L. setaiia. state of Parana, a province which formed part of the State
S. Paulo at the time Aug. St. Hilaire travelled in Soath Brazil.
It is strange that this bird has never been met with since its discovery ; bnt
still more remarkable is it that Katterer obtained in the state Parana another very
distinct species, the type of wiiich has also remained unique hitherto. This is
( 333 )
33. Leptasthenura striolata (Pelz.).
Si/miUoxis siriolald, (Natterer MS.) Pelzeln, Sitziiiigsher. Ahul. Wirn xx. (I80G) p. 159 [Curytiba,
Parand] ; idem, Zm- Oniith. Bra»il. i. (1867) p. 38.
1. Mus. Vindob., No. 20,01U. c? ad. t'urytiiia, Parana, October 8, 182U.
Natterer coll. Type of Si/nalldxis striolata Pelz. Wing 54 ; tail 90 ; bill
10 mni.
Top of tlie head riifoiis brown, each featlier distinctly margined with
blackish ; whole back paler and duller rnfcscent, den.sely covered with blackish
brown lougitndiual stripes, only the rump and upper tail-coverts being imiform.
Upper wing-coverts dusky, the lesser and the median series with indistinct
rul'escent edges, the greater ones more distinctly margined with brownish white.
Remiges dusky, margined exteriorly with pale rnfcscent. Middle pair of tail-
feathers entirely dark brown, the others dark brown with pale rnfons tips ; two
outer pairs almost entirely rufous, only the basal half of the inner web being brown.
Above the eye a narrow buffy superciliary streak. Cheeks and ear-coverts dark
brown ; sides of the neck pale sandy brownish with slight dusky mottlings.
Lower surface pale buff, throat faintly dotted with pale brown ; sides of the
body washed with light brownish. Axillaries, under wing-coverts, and a distinct
margin to the inner web of the remiges fawn-colour. Bill horn-brown, base of
the lower mandible yellowish white.
This species is perhaps nearest to L. andicola ; but that bird has a striped
nnder-snrface, the back dark sepia-brown striped with white, a much broader white
eyebrow, ai]d no rnfons on the tail. L. jjileata has also white stripes on the back,
greyish white tips to the outer tail-feathers, and dillerently coloured lower parts ;
it agrees, liowever, with L. striolata m the yellowish basal half of the lower
mandible (which is wholly dusky iu L. amlicola) and in having only a narrow
superciliary stripe.
According to my views, the genus Li'ptastJienura consists of the following
species :
1. L. aiiijitlialoidi'S (Kittl.), ('Iiili ; N.W. Argentina: Tucuman ; S.W. Peru:
Islay.
2. L. platensis Reichb.* Central and South Argentina: (Jordova, Parana,
Bnenos Aires.
3. L. amlicola andicola Scl. Andes of Ecuador and Sierra Nevada de Sta
Marta, ( 'oloml)ia.
4. L. andicola certliia t (Mad.) Andes of iMerida, Venezuela.
5. L. pileata Scl. Andes of Lima, W. Peru.
6. L. striolata (Pelz.) Curytiba, Parana, South Brazil.
7. L. setaria (Temm.). Castro, Parana, S. Brazil.
8. L. fuliffiuiccps fuliijiniccps (Lafr. k D'Orb.) [type ex Sicasica, Bolivia]
(= boliviuna Allen — type ex Bolivia !). Andes of Bolivia.
9. L.fuliginiceps ]Mranensis Scl. Argentina: Parana, Cordova, Tucuman, etc.
I hope to discuss the various species of this genus on another occasion.
* This species is quite distinct from L. ai'ijitliaUndes. I have seen a large series,
t SijiiarnU ccrthia Madarisz, Ann. Mm. Iliimj. i. (IIIO.')) p. HVA [San Antonio, Merida] ; Lcjttiutlienura
mtmliraijmis Eilcy, I'm: liinl. .V»c. W'asli. x\iii. (liios) (p. L'l'J) [San Antonio, Merida].
( 334)
34. Notes on Synallaxis elegans Less, and allies.
Si/)inllaxis eleyaiis Lesson, Siiji/ili-iii. Ocurres Btiffun (ed. Leveque), Descr. ilammif. el Oin., 1847,
p. 289, descr. orig. (J ad. (Guayaquil).
S. paucalensis Tatzanow.ski, Ornilli. PcroH ii. 1884. p. 131, descr. orig. (J ad. (Paucal, N. Peru).
S. subupeciom Salvadori & Festa, B<ill. Mus. Torino, xv. no. 3C"2 (1899) p. 21, descr. orig. (J jur.
(Balzar, near Guayaquil).
Lesson's excellfiit description, which has hoen overlooked by all recent writers,
refers without any donbt to the West-Ecuadorian rei)resentative of the gronp,
comprising 5. speciosa, S. subspeciosn , and S. paucalensis Tacz. Thus, S. subspeciosa
becomes a synonym of -S. elcgans.
The bird described by Lesson was evidently a perfectly adalt male : cf. " nne
calotte d'un noir mat et profond recouvre le dessus de la tcte depuis le front jnsiiu'a
la niique," and " tout le devant da con, depuis le menton jusqn'au thorax, est
blanc lav^ de roussatre par places, ce qni forme nn large plastron de cette conlenr,
arrcte dans le bas, en travers du con, par nn cordon noir, bord^ Ini-meme par uuo
echarpe d'lrn roux-marron vif. Le ventre, le bas-ventre et les flancs, sont d'un
jaune-rouille tres pale et nniforme, le dos et le cronpion d'un cendre clair ; " while
the type of <S', subspeciosa was a young male : cfr. " pileo et cervice saturate
brnnneis, colore griseo dorsi qnoqne brunnescente," etc.
In the British Mnseum there is a specimen from Balzar (collected by
Illingworth) which diflers from both these descriptions by lacking the chestnut
pectoral area, the whole nnder-parts below the black jugular band being uniform
fulvous. This is no doubt the female of S. elegans, as in the closely allied form
S. e. speciosa of Puna Island, and in a series of the so-called S. paucalensis of
Nortliern Peru the females differ e.xactly in the same way from the males.
Judging from the material before me, there arc two distinct forms to be
recogni.sed — one confined to Puna Island, the other inhabiting S.W. Ecuador and
N.W. Peru.
Five adult males, from various places in N.W. Pern (Tembladera, Trnjillo,
Otnsco, Platanar ; coll. 0. T. Baron), agree very well with the description of
.S'. paucalensis Tacz.*, all having below the black jngnlar collar a chestnut band
well defined against the fulvons colour of the abdomen.
This tallies as well with what Lesson says about the coloration of the
nnder-parts (vide supra). Furthermore, two females from North Peru (Trnjillo
and Platanar) in the Tring Museum are certainly identical with a female from
Balzar in the British Museum. Therefore I have no hesitation in nniting
<S'. 2"'"calensis and S. elegans.
The form from the Puna Island, described by Salvin as a Formicieora'\ (! 1 I),
however, seems sullicientiy distinct to lie separated snbsiiecifieally. The male type
(in London) and an adult male in Tring have the abdomen much darker, chestnut
rufons, only a shade lighter than the chest. The females,* like those from Balzar
and North Pern without any chestnut on the lower surface, differ from these by
having the nj)per wing-coverts much paler, isabelle instead of cinnamon rnfous.
These two forms have accordingly to stand as :
• Orn. Virm ii. 1S84. p. 131 (type : ^ ad. ex Paucal, N, Peru),
f Firrmictvora ipectosa Salvin, lbi» 1H76. p. 491 (Puna Island).
\ The statement by Sclater {^Cat. liii-ds xv. p. 251) : "Female similar, but without the black gorget,"
is a mistake, it being present in the three females in the British Museum.
( 335 )
({. Sjiiallaxis eleffai/s elci/iijis hesti. S.W. Ju-tiador : Guaya'juil, lj;il/.iir ;
iV. II'. Peru: Puucal, Tnijillo, Toiuliluilera, OLusco, utc.
1 ?. Balzar. Wing 54 ; tail 55^ ; bill 15 mm.
5 SS. North Peru. Wiug 55— 5.S ; tail 58—05 ; bill 14—15 mm.
2 ? ?. North Peru. AViug 56, 50' ; tail r,l, 04 ; bill 14, 15 mm.
b. SijnaUaxis elajans speciosa (Salv.), Pnua Islaml, in the Bay ol' Giiayaqnil.
■Z Si. Wiug 58, 60; tail 63; bill 14, 15 mm.
2 ? ?. Wing 56, 57 ; tail 57, 59 ; bill 14, 15 mm.
35. Philydor cervicalis Scl. sb(juhl be Automolus infiiscatus cervicalis (Scl).
\^.\ii(ihates hiftist^atna Sclater, Ann. Marj. Nat. Hid. i^i) xvii, (1S5IJ) p. 4lj8 ('' in Puruvia oricQtali) ".]
Phili/ihir Ki-ekulis Sclater, P.Z.8. 1889. p. 33 [British Uuiaua: Bartica Grove & C'amacusa] ;
idem, Cat. Birds xv. p. 101 [Bartica Grove, Camacusa ; Surinam].
Aulomuhis sflalcri Salviu, Ibis 1885. p. i'20 [Bartica Grove, Camacusa] ; Sclater, Cut. B/rd'^ xv.
p. 'J5 [part. : spec, j — m. Camacusa, Bartica Grove] ; Berlcpsch & Ilartert, A'nw. Zonl. ix.
(l'JU2) p. Gl [Ncricagua : Orinoco ; La I'riciou, La Union, Nicarc ; Caura River, Venezuela].
Wing. Tail. Bill.
1. Mils. Brit, ex coll. P. L. Sclater : cj jr. Bartica Grove,
Jauuary 1(J, 1880 (specimen a oi Phili/dor cercicalis
Hcl, Cut. B.) 82 71 lOJ mm.
2. Mus. Brit, e.x coll. P. L. tScl.-iter : ? jr. Camacusa,
May 21), 1882. H. Whitely coil. Type of P. cervi-
calis Scl. ........
3. BIus. Triug. ? ad. Camacusa, May 27, 1882. H.
^Vhitely coll
4. Mus. Triiig. J ad. Camikii Muuut., December 11, 1889.
H. Whitely coll
5. Mus. Triug. ¥ jr. Biver Carimaug, Juue 15, 1885.
H. Whitely coll. . . . '" .
0. Mus. Berlcpsch. S ad. Bartica Grove, Jauuary 21,
1880. H. Whitely coll
7. Mus. Berlepsch. tS ad. Camacusa, July 11, 1882. H.
Whitely cull 86 74 20 „
The type oi 1'. cercicalis as well as No. 1 (of the above list) are both immature.
The "species" has nothiug to do with Phibjilor cnjthroccrcus, with which it has
been compared iu the original descrii)tioa ; bat is very closely allied to Automolus
infiiscatus {= sclateri auct.). Indeed, it is so near the latter that I only
reluctantly allow it to staud as a dilfereut form. Specimens Nos. 3 — 7 are exactly
like the typical examples. This series dillers from typical Peruvian skins only iu
having the top of tlie head decidedly rufescent, iu slight contrast with the olive-
brown back. In tiie birds from N.E. Pern the head is olive-brown like the back, only
the forehead being slightly tinged with rufescent. A good series from the Caura and
Orinoco Bivers agrees in every respect with Guiana skins, and the specimens of
Automolus scl'ttcri from British Guiana iu the British BInseum are also absolutely
identical with the type of 7-". cervicalis! Although the colour of the crown serves
to distinguish most specimens of the two subspecies, I may remark that one male
from British Guiana (No. 4) and two skins from the Caura are hardly dill'erent from
Peruvian examples. Immature birds show slight greyish edges to the feathers of
the foreneck, producing a mottled appearance,
■12
( 336 )
36. Dendrocolaptes atrirostris LhIV. ^t iruili. should be Deudi'ociucla atrirostris
(Lair, it D'Oib.).
DcudrocohqiUs (drirostrh Lafrcsuaye & D'Orbigny, Sijn. Ai: u. in .l/n;/. /C<'<il. 1K!8. cl. ii. p. 12
[Guarayos, rep. BoUviaiia] ; D'Orbigny, Voi/titjc, 0,\. p. 30'.l, tab. 54, fig. 1 (fig. pussirna!).
Dendrmimht aliinislris Ridgway, P. U. S. Mils. x. 1887. (1888), p. 4'.IU, 4;i;! ; Meuugau.v ut llullniayr,
Mem. S.ic. hist. mil. Aiiluii. .\i.K. (I'jllli), p. 1^!0 (crit.).
D. m/no;- Pelzcln, Ziir nrii'dk. Ihusil. i. (1807), p. (iO [S. Viccnlc, Matlogrosso].
D.finiiiijatu idoui, /. r. p. 42 [part. ; spucimeu e.\ 8. Vkintc].
Wing. Tail. Liill.
1. Mils. I'aris. >Siiec. Inj). (mounted), lalielled :
"Giuiriiyon, Bolivic, D'Orbigny. DciulrflcolajiU'.'i
atr/rosfr/.s, nob. (type)" '•'■' '"' -'' "i"'-
•2. Mns. Vindob. No. I.">s92. Type oi' D. minor l\-h.
? juv. " S. Vicente, December 1820." Natt. coll.
* 3. Mus. Vindob. No. 15899. ? ad. " S. Vicente,
December 182G." Natterer coll.
4. Mus. H. V. Berlepscli. No. 1257. G. Garlejip cull.
(? jnv. " S. Mateo, Bolivia, August 10, 1891 " .
5. Mus. H. V. Berlepscb. No. 1113. G. Garlepp coll.
¥ . " S. Mateo, Bolivia, July 20, 1891 "
There can be no doubt that Dendrocincla minor of Pelzeln is identical with
D. atrirostris (Lafr. & D'Orb.). Both species are based on immature specimens
with the bill entirely blackish. In the adult, the upper mandible is dark brownish
lioru-colour, the lower one, except the gonydeal angle, yellowish white (No. 3 of
the above list). No. 4 agrees with the types of atrirostris and minor in having
the bill entirely blackish. In No. 5 only the upper mandible and the basal half
of the lower jaw are blackish, the apical half of the latter being yellowish white ;
thus the coloration of the liill is intermediate between that of No. 3 and that of
Nos. 1 and 2.
The black-billed specimens (Nos. 1, 2, and 4) have a distinct oclireous patch or
sufl'usiun on the occiput, a sign of immaturity, also to be seen in young examples
of D.fuliginosa, phaeockroa, and olicacea. In No. 4 it is barely indicated, and in
No. 3 there is no trace of it.
In coloration, the skins from Mattogrosso are practically identical with the
typical birds from Bolivia. The type of D. minor has a decidedly shorter tail than
the others, but this is doubtless due to its being very young.
In all essential particulars the five skins before me are very uniform : forehead
and crown finely striated with buffy ; from behind the eye along the upjier margin
of the ear-coverts a broad ochraceous-buff jjostocular streak ; throat rather paler
than the rest of the under parts, and slightly mixed with greyish ; jugulum with
hair-like, buffy shaft-lines ; exposed portion of the remigcs clear cinnamon rufous
without any olivaceous tinge, but rather paler than tlic tail.
D. atrirostris is evidently the western representative of J>. turdimi, with which
it shares the pale striations on the crown ; it differs, however, by its broad ochraceous
postocular Btrijie, and in having the outer webs of the quills and the tertials pure
cinnamon-rufous (instead of olivaceous brown, scarcely different from the colour of
* It is interesting to observe tliat ilic collector of tiie speeiraen.s, Natlerei', correctly recognised tbeni
as belonging to the same speeies, both being marked by bim with number " 705."
( 3.V )
the back), and the throat finely nmliihited with greyish (instead of niiiform bi-ight
buff). The wings, too, are considerably shorter.
The distribution of these two allied forms is as follows :
1. Dendrodnclu turdiim (Lcht.).
Dendrocolaptcs lurduius Lichtenstein, Abhuiull. Akail. Berlin n. d. ./. lSlt>-l:) (pulil. lS:iO), p. 2il-t,
tab. ii. fig. 1 [i5«/u« ; cfr. 1. c. a. d. J. 1820-21 (publ. 1822), p. 2i;4J.
Eastern Brazil from Bahia to Santa Catharina.*
2. JJendrocincla atrirontris (Lafr. & D'Orb.).
Eastern Bolivia : (lUarayos (D'Orbigny), San llaleo (G. (iiirlepji cull.) — Central
Brazil, Western Mattogrosso : S. Vicente on the It. Guapore (Natterer coll.).
I hope that the following key will help ornithologists to recognise the species
allied to D. turdina.
A. Top of the head finely striated with bulF.
a. E.xposed portion of remiges olivaceous brown,
hardly different in colour from the back.
Thi'oat bright buff. Scarcely any indi-
cation of a postocular streak. Wing
about lU-4 — 114 mm. Hab. Eastern Brazil
(Bahia to S. Catharina) . . . . JJ. turdina (Lcht.).t
I). E."vposed portion of remiges bright cinnamon
rnf'ous, very different from the colour of the
back. A broad ochraccous postocular
streak. Throat finely undulated with
greyish. Wing 95 — 99 mm. Hab. Eastern
Bolivia and Mattogrosso .... U. atrirostris (Lafr. & Uib.).
B. Toj) of the head uniform.
c. Upper wing-coverts deep dark chestnut like
the outer web of the remiges. Anterior
portion of throat distinctly whitish. Hab.
Cayenne, Surinam, Brit. Guiana ; Orinoco
region ; Lower and Upper Amazonia . . D. morula (Lcht.).+
'/. Upper wing-coverts olive-brown, very different
from the cinnamon or cliestnut rufous
colour of the remiges. No whitish jjatch
on the throat,
a'. A broad ochraccous postocnlar streak
from behind the eye to the ])osterior
edge of the ear-coverts. Lower throat
with narrow, but distinct, buffy shaft-
lines or dots. Hab. Cayenne, Guiana,
Lower Amazonia .... D.fuliijinosa (Vieill.).§
* Specimeu in Mu3. Deilupocb (HjavupcuU.). J 13 specimens,
t 17 specimens. § 15 specimens.
( 338 )
'. No distinct jialu jiosloculur .stroiik. no pule
markings on thu lower throat.
i'. Exjiosed jiortion of the rcmigcs distiuctly
washed with olivaceous browu. Ui)i)er
mandible as well as rami and upiier
edge of lower jaw deeji black. JIab.
Costa Kica, Panama, West Colombia
(Caiica Valley : also Ibnnd in i'ogota-
coll.), West Ecuador ....
b'-. Exposed [lortiou of the remiges clear
chestunt rufous. Ujiper mandible as
well as rami and upper edge of lower
jaw horn-brown.
e'. Upi)er parts olive browu.' Throat
bufi}', distinctly paler than the
breast. Hab. Orinoco region, East
Ecuador, E. Colombia (found in
Bogota coll.), Upper Amazonia
(N.E. Peru : Ic^nitos ; llio Jnrna) .
f^. Upper parts light cinnamon brown.
Throat ochraceons-browu like the
breast. Hab. North coast (jf
Venezuela (8. Estebau to Cumana),
Trinidad, Tobago ....
IK olimcm Lawr.*
D. pliaeochroa Berl. & Ilart.f
1). meruloides (Lafi-.) J:
37. Thamuophilus tristis Scl. ct tSalv. — T. polionotus Pelz.
Thaninophitus poliomlus Pulzelii, Zur Ornith. Braail. ii. (ISi'iS), p. 147 [Marabitauas, Barcellos],
descr. orig. (J J .
T. Irisli.-. Sclatur & Salvin, Niniiciwl. Ai: Ncotrup. (1873), p. 100 [Oyapoc, Cayenne], descr. orig. J.
Thamitiiiihilus sp. iuc , Berlepsch & Harteit, Noe. Zvnl. ix. (1902), p. G'J [La Pricion and Nicare :
Caiira R., Venezuela].
Wing. Tail. Bill.
75 [damaged] 20 mm.
1. Mas. Viudob. No. lOGOU. S ad. Marabitauas, Kio
Negro, April 1S31. Natterer coll
2. Mus. Vindob. No. 1G6U7. c? imm. Marabitauas, Kio
Negro, April 0, 1831. Natterer coll.
3. Mus. Vindob. No. 1G0U8. ? ad. Barcellos, liio
Negro, November 31, 1830. Natterer coll.
[These examjiies are the type.< of 'J'.j/olioiiotits Pelz.]
-t. Mus. lirit. (c?) ad. Oyajwc, C!ayenne — ex Verdey,
1871. — Type of 7'. <;76<(s Scl. & Salv.
5. Mus. Tring. (S) ad. Nicare, Caura, Vcnez., Jan. 1,
1901.— E. Andre coll
6. Mus. Tring. ? ad. La Pricion, Caura, Jan. 31, 19ol
7. Mus. Tring. ? ad. La Pricion, Caura, Dec. 12, 1901
8. Mus. Tring. ? ad. La Pricion, Caura, Jan. 19, I'.Hil
[Nos. (J — 8 collected by Mr. E. Andre and reforifd to by Messrs. Berlepsch and
Hartert as Tkamnoplulus sp. inc. (1. c.).J
70
01
no
01
20 J.
18
20
* 2G specimens.
t 21 specimens.
X 17 specimens.
( 339 )
The type of T. triatia agrees iu every respect with the adult S of Pelzeln's
7'. polionotm, except in having the lower parts a shade paler schistaceons. Both
have the forehead and crown blackish, the bend of the wing white, and small bnt
very distinct white apical sjiots on all the nppcr wing-coverts.
No. 5 differs in having the crown less mi.xed with Ijlackish, and by its
shorter bill.
No. 2, being slightly iinmatnre, has still the brownisli rpiills of the juvenile
]iliimage, and some of the apical spots on the greater wing-coverts are tinged
with bnffy, hut otherwise it agrees with the adults.
The nearest ally of T. jwlionotiig is undoubtedly T. incertiis Pelz.* The male
of this species differs, however, at a glance by lacking the black cap and the white
apical spots on the median and greater wing-coverts, only the bend of the wing
being freckled or edged with white. The general colour, too, is of a much jialer
slaty-grey. The female of T. incertiis has the back distinctly paler rufous brown,
and the ferrnginous colour below is confined to the throat and chest, while the
abdomen is of a uinch paler tint, varying from oeiiraceous to fulvous.
Another very near ally is T. jurtiaiiKs Ihering. f This form has the black cap
of T. polionotus, bnt the uniform black wing-coverts of T. incertus.
T. aethiops Scl. belongs also to this group. The male differs from those of the
foregoing species in the deep ])lack (instead of schistaceons grey) coloration. The
wing-coverts are uniform black without white a])ical spots. The female is readily
distinguishable by its blackish tail. Moreover, the general colour above and below
is much darker, chestnut rnfons.
The range of these four forms is as follows :
T. (iPthiojis Scl. Eastern Ecuador.
T. juruanus Ihering. Rio Jnrua, W. Brazil.
T. polionotus Pelz. North Brazil : Marabitanas and Barcellos on the Rio
Negro ; E. Vcneznela : Nicare and La Pricion, tiaura R., a tributary of the Orinoco;
Cayenne : Oyapoc.
T. incertus Pelz. V^icinity of PanL
38. Thamnophilus punctuliger Pelz.
T. piiiicliiUfifv Pelzeln, Ziir (h-ii lijnxiL ii. (I8G8), p. 14C [Borlia : Uio Madeira].
Mus. Vindob. Type of species : cj ad. " Borba, December." — Natferer coll. —
Wing 77; tail 00; bill lOi mm.
This is a close ally of T. po/ionoli/s, having, like that species, the top of the
head black, and small white apical spots on the upper wing-coverts. It differs,
however, by the presence of a distinct white interscapular patch. The general
colour of the plumage, too, is even paler and purer schistaceons than in T. jtn-uanus.
So far as I know, Pelzeln's type is the only specimen in existence. Most probably
7'. poliniiotiiA, T. jiirKiinns, T. punctuli(jer, etc., are geographical representatives,
luid would [icrhaps, more properly, be designated by trinomials.
* Tn Nnr. y.iml. xii. (llin'j),p. 2S|,T placed this liirrl in Iho {;c)ius 7A/.«;//i ;7«hiw. ncai- D. xchidaopvii,'b»*-
after a careful study of the wliulc group I came to the conehisinn that there is only a supiTlicial resemblance
lictwecn the males of tlic two species, anil tliat the real affinities of T. incertus lie wil h T. potioHotiit. This
opinion is strengthencil by the coloration of the females, which are almost iilcutical.
t llerMa Mm. Paulut. vi. ^May 1905) p. 439 [Rio Junta, W. Brazil]. Types examined by nie.
( 340 )
39. On Myrmelastes exsul (f^d.) ami its allies.
M)/rmen~a exsul Scl. was describeil from a specimen, collected in Panama I)}'
Uelattre (/-'. Z. S. ISoS. p. .'540). Two years ai'torwards Mr. Sclatcr identified some
examples from EsnieraUlas, Western Ecuador (c'oll. Fraser), with this species
(P. Z. S. 1800. p. ~04), and since that time the bird fonnd on the western slopes of
the Ecuadorian Andes was commonly called ^f. exsul. In 1804 Messrs. Sclatcr
and Salvin pointed ont the differences existing between this species and a closely
allied form contained in one of MacLeannan's collections from TJon Hill Sfation,
Paridina, and described the latter as new under the name of M. immaculutn.
Having always snspected that there mnst be some error as to the original locality of
M. exsul, I asked Dr. Forbes for the loan of the typical specimen, which was
obligingly sent to me. On examining it, I was really siirjirised to find that it had
nothing to do with M. exsul auct. of Western Ecuador, but belonged apparently
to the species named M. inimaculata by Sclater and Salvin! In order to be quite
sure of my identification, I compared Delattre's specimen with the two male types
of M. immaculafa in the Rritish Museum, and found the three skins alsolutely alike.
They agree perfectly in the dimensions, and in coloration. The upper wing-coverts
are uniformly rufescent brown, only the lesser series being blackish with
very small white apical dots.* In .1/. ersiil auct. fex W. EiuadorJ all the upper
wing-coverts are marked with sharply defined, rounded white apical spots, and
the tail is considerably shorter, as will be seen fmni the measurements given below.
There can be no longer any doubt that the locality of Delattre's specimen was
quite correct, and that the name M. exsul has to be used for the Panama-form.
The one from Western Ecuador being thus without a name, I propose to call it
Myrmelastes exsul maculifer nnm. uov.t
It remains to say a few words about the proper application of the name
M. exsul. Mr. G. K. Cherrie distinguished two forms, which he calls
a. M. immaculatn Scl. k Salv., having the under wing-coverts white and the
first primary edged with white. Hab.: from Panama to Talamanca, S.E. Costa Hica;
b. M. intermedia 'Clherrie, having the under wing-coverts dark grey and the
first primary not edged with white. Hab.: from Panama along the Atlantic
lowlands of Costa Rica to Nicaragua.
The type of M. exsul Scl. and the two male types of M. immaculata Scl. & Salv.,
however, belong to M. intermedia Cherrie, having no white edge to the first
jirimary, and the under wing-coverts pale ashy grey, like the under-surface of the
quills. It follows that these three names are strictly synonymous, while the form
identified as M. immaculata by Mr. Cherrie would require to be renamed if really
distinct. From what Mr. Cherrie says, it would seem that his two forms are found
together over a considerable area. Two adult males from Cascajal, Panama, in the
Tring Museum have the basal half of the first primary distinctly edged with white,
while a third specimen from the same locality (No. 3 of the list given below) shows
no trace of this margin ; the under wing-coverts are ashy grey in all three skins.
* This lias been well describeil in St-later's origin.il description: " tilarvm tectricihns minonhi/x
7ii^r!r/r}ifihit.t ad ajnrcni albo pitjtrfaflx."
t Typo in Mus. Tring: No. 414 jj mi. I'aramba, N.W. Ecuailor, :«00 ft. C'ollecteil by K. Miketta.
( 341 )
A large series (17 siiecimens) from fjosta Rica (Atlantic side) and Eastern
Nicaragua (Cliontales and La Libertad) agrees in every respect with the two first-
named examjjles from (Jascajal. The ? ? from the Panama railroad (McLeannan
coll.: M. immaculata Scl. & Sal v.) are also absolutely identical with others of the
same sex from f!osta llica and Nicaragua. Therefore it seems to me impossible to
recognise more than one form occurring ou the Atlantic side of the mountains in
Costa Rica and Panama.
According to my view.^. the following geographical forms of this group are
distinguishable :
1. ^^!/nnclast<^■'i exsid e.rsul (Scl.)
Murmecha e.rm! Sclater, P. Z. S. 18.08. p. y40 (Panama, coll. by Delattre.— Type Xo. 45139 Mus.
Liverpool).
M!irmcci::a inimriculata Sclater & Salvin, P. Z. S. 18G4. p. .3.'J7 (Lion Hill Station, Panama).
Mip-mcriM iidermeiJia Cherrie, Prne. U. S. .Uim. xiv. 1891 (1892) p. 34.5 (Talamanca S E. Costa
Rica.)
/fa/j. Panama, Eastern Costa Rica to Nicaragua (Chontales, Escondido River).
c? ad. Upper wing-coverts uniform rufescent brown, only the bend of the
wing and the lesser wing-coverts with very small white apical dots.
? ad. Back and upper wing-coverts dark rufous brown, the latter without any
light apical spots ; tliroat very dark slate-colour, vest of under-surface dark sepia
brown, sometimes with a faint rufescent tinge.
Mus. Liverpool, No. 4030, c? ad. Panama
(Delattre coll.). Type of 31 exml Scl.
Mns. Brit., 2 SS ad. Panama (McLean-
nan coll.). T3i)es of M. immaculata
Scl. & Salv
Mns. Tring, 3 <?c? ad. Cascajal, (Jocle,
Panama (Hyde coll.) ....
Mns. Tring, 4 c? J ad. Signirres, E. Costa
Rica (Cherrie coll.). Determined by
Cherrie as J/, intermedia Cherrie .
Mus. Tring, 2 c?c? ad. Carrillo, E. Costa
Rica (Underwood coll.)
Mns. Tring, 0 ? ? ad. Signirres and Carrillo,
East (Vista Rica (Cherrie and Under-
wood coll.)
Mns. Brit., 3 ? ? ad. Panama (McLeannan
coll.)
2. Myrmdastes exsul occidcntulis (Cherrie).
Miirmrrhn liiimnaihjtu (irrldeutalis Cherrie, .^wfc viii. (1891), p. 191 (Pozo Azul, West Costa Rica).
/fidi. Western Costa Rica (Pozo Azul, Buenos Aires, Lagarto, Boruca, etc.),
and (Ihiriijui (Bogava).*
cj scarcely different from that of the foregoing lonn, but the 9 is at once
• Outiam lianus (Atilt xviii. IIIOI. p. .^07) rccnivls M. ititrrmnliiis (botli sexes) from Cbil'iqui, l>ul llu'
pair from l!oi,'.iva in the Triny Museum as well as a female from Cliiriiiui in tlie r.ritisli Museum belmif; to
iV. e. occiilenlalis.
( M2 )
recognisable bj- the (■(ildration nf the nmler jiurts, tin' roreueek being bright
ferrnginons red, the breast ami abihnneii light rnres<Tnt brciwn.
WiiifT. Tail. Rill.
1(1 3ii ad. from West Costa Rica (for exact
localities see above), coll. by Clierric and
Underwood ......
0 ? ? ad. from West Costa Kiea
1 c? ad. Hogava, Cliiri([ni (Watson cull.)
1 ? ad. I'ogava, C]iiri(|iii (Watson coll.)
3. ^fi/rmehdttrs exsiil maculij'er [Tellm.
.Ui/rmrci-.a crsii! (nee Selater \><'i9.) SM.-iter, P. Z. S. ISHO. p. 204 (EsmeraUlas) : Sclatcr & Salviii,
P. /.. N. 1S7!>. p. 520 (Xcehe, Aiitioquia) ; Berlepsch * Taezamnvski, /'. Z. .S., 188:i. p. h(A
(Chimbo) ; Hartert, Nnr. Zoi.l. v. (189S) p. 403 (Chimbo, Paramlia) ; Salvadori & Festa,
Boll. MHi>. Torino xiv. No. ?,Ci (1890) p. .".1 (Rio Peripa) ; Goo.lfellow, lh;< VMyi. p. CU (Santo
Domingo and Guanacillo).
IIiil). AV'estern Ecuador (Esmeraldas, ( Miinibo, Rio Teriiin, 8. Domingo,
Gnanaeillo, Paramba, Cacliabi, Lita, S. Javier, etc). W. Colombia: Rio Dagna
( Rosenberg coll. in Mus. Tring), Xeehe in Antioqnia (Salmon).
(?. At once distingnishabh- frmn the same sex of the two preceding forms
by its decidedly shorter tail, and by having all the npper wing-coverts spotted
with white at tlie tips.
?. Under parts as in the ? of ,1/". e.csid oendeiiliilh, bnt readily dislingnish-
able by its shorter tail, and by the fnlvons-white apical spots on all the iipjier
wing-coverts.
Wing.
20 c? S from "Western Ecuador .... 03—08
2 c? c? ad. from the Rio Dagna, Colombia . . 00
12 ? ? from Western Ecuador .... 02—00
1 ? from the Rio Dagna 00
N.P..— The specimens from Neche (Brit. Mns.) agree very well with tliose
from W. Ecuador, except in being rather paler, more olive brown, less rufescent
on the upper parts.
411. (^n Myrmeciza laemosticta Sal v. and M. stictoptera Lawr.
The Tring Musenm possesses eight specimens of a !\fiirmrci~(i, collected by
Mr. C. F. Underwood, near Carrillo, Costa Rica. Three of them, as well as a
■ipeeiraen from the same locality in the British Mnseum, agree perfectly with the
original description of Af. stictoptera Lawr. All are marked c? by the collec'tor.
Tiie live other skins at Tring, an example from Carrillo, and one from Santa F<',
Veragna (Arce), in the British ]\Insenm, all marked ? , dift'er in having the chin
and fhr(];Lt spotted with white (instead of uniform blackish), the foreneek slate-grey
like till' abdomen (not blackish like the tliroat), the white on the bond of the wing
nuich less extended, oi- even wanting, anil the pileura strongly washed with olive
lirown. This series agrees in every resjiect with the unsexed type of il/. laemosticta
Salv. from Tncurriipii, in the British Mnseum. In all other characters, snch as the
])attern of the wing-coverts, the i)resenco of a large white interscapular blotch, the
length of wing and tail, the shajie of the bill, etc., the two series agree perfectly
with each other; and as some of the specimens with a spotted throat were
( 343 )
obtained on the same day as otliers with tlie thmat nniform lilaekish, I think there
can lie no hjni^cr any donht (hat they merely represent the two sexes of one
species.
In Western Ecnadcu' a very near ally takes tiu^ phu'.c of ,1/. laemnsticta ; it has
been named ^[. iiiiiiicaiidd by Salvin and Godman. Besides tlie nnsexed type from
Intac in the British Museum, I examined three sexed ? ? of this form whieli were
collected by Mr. Rosenberg's correspondents at Paramba, Lita, and l!io Verde, in
N.W. Ecuador. Tliese four specimens agree in the markings of tlic throat witli the
/rw'wo.sY/f/ft-pInmage, but differ from the f'osta Rica series in the following details.
The white s])ots are confined to the chin and the anterior portion of the throat ; the
tail is blackish (not dark rufous browu), and the back rather dnller rnfescent brown.
In dimensions they agree with M. laemoatictn, the wing measuring 07 — 68, the tail
47—49, the bill 18—20 mm. 'W'e have thus two forms :
a. Mi/rmcciza laemosticta laemosticta Salv.
Myrmeciza laninosticta Salvin, P. Z. S. 1864. p. 582 descr. orig. ? (Tiicurriqui, Costa Rica).
Mijrmc.eh.a fikl'ipterri Lawreuce, Ann. N. York Li/c. viii. (May 1SG5) p. 132 descr. jj (Angostura,
Costa Rica).
Hal). Cosfa rtica (Tncurri(pii, (!arrillo) and Vvragiia (Rauta Fe).
4 (?c? : Wing 05-07 ; tail 45—48; bill 10 mm.
7 ? ? : Wing 02—00; tail 40-48 ; bill 18—10 mm.
b. Ari/rmeciza laemosticbt nifiricamla Salv. & Godm.
Miirmcr'izd vUjricauda Salvin and flodman, Biolng. Ccnir. Americ. ii. (18'J2) p. i.'jO (Intac).
Hal). Wc.sfcn, Eciiaflor : Intac (Bnckloy) ; Paramba, Lita, Rio Verde
(Miketta and Flemming coll. — Mus. v. Berlepsch, Monac. et Vindob.).
41. Cercomacra hypomelaena Scl. = C. serva (Scl.)
Pyrirjlriia srrra Sclater, P. Z. 8. IS.'iS. p. OlJ [Rio Napn, East Ecuador] ; Sclater, Cai. Binh xv.
p. 271 [e.xcl. dcscrijition of female wliich is referaljle to C. njipnu-imaii^ 5 , ^i™! excl, specimens
g.i].
Ccrconmrm hypmnchicna Sclater, Cat. liirih Piril. ^^u^. xv. (l.SOO) p. 208 [Cosnipata. S.E. Pern].
Wing. Tail. Bill.
I. Mus. Brit. ((?) ad. Rio Napo (e.x Verreaux), coll. P. L.
Sclater. Tyite of Pi/r/i/lriiri. serva Be].
'.'. Mns. Brit, ((?)ad. 8arayacu, E. Ecuador (0. Buckley
coll.) ex Mus. Salvin & Godman {P. scrm, specimen c).
;i. Mns. Brit, (c?) ad. Sarayaeu, E. Ecuador (C. Buckley
coll.) ex Mns. Salvin & Godman {P. serra, spec. /) .
4. Mus. Brit, "c?" ad. Cosnipata, S.E. Peru, Juno 2,
1871. — II. Whitely coll. — Type of ('. hi/pomdacna
('. Iii/jiomehii-iia has been described by a curious oversight. Mr. Sclater placed
it quite correctly in the genus Cercomacra, but having referred his P, scrfa to
another genus, he omitted to compare it with the latter species. Otherwise,
ho certainly would have found that both are the same.
Goraparing throe adult Jc? of Pi/riijlcna serva (including the type) with the
typical specimen of t '. //i///omc/nc/ia, I i'ail lo see any dilVerence between them, except
that the latter has the bill very slightly narrower. It is, however, matched in this
respect by a female from the Rio Napo.
( ■•544 )
C. serva (ScL), as the sjieoies ought to be called, is certainly not a ri/i-if/h-iiii,
Ijiit finds its nearest allies in Cercomacra approximans and C. nigrencens, with
wliicli it exactly agrees in the shnpe of the bill nnd form of the tail. The species
of rijriglfiia, on the other hand, differ very markedly in the iollowing particulars :
the bill is decidedly narrower and more slender, also rather higher and not
?o flattened; the tail-feathers are much wider; the forehead and the lores are
thickly covered with soft, erect feathers, while they are much loss densely set and
rather stiffer (more bristle-like) in Cfn'omarra.
The female of C. .'icrfa is also very similar to that of C. (ippvoximnns and
('. iiiqrrsri'iix, in having the whole lower surface and the sides of the head bright
fiM'rnginous ; it differs, however, from both in the forehead being dark olive grey
like the back (instead of ferrnginous as in the allied species). The Tring Museum
has lately received a male from Wiuchurras, near Pozuzo, province Ilnanneo, Peru.
A"i. On Dysithamnus subplumbeus, Hypocnemis schistacea, Heterocnemis
leucostigma, and H. saturata.
The careful examination of a large amount of material reveals the surprising
fact that the above four " species," placed by the latest authority ' in three different
genera, are very closely alHed and evidently but geographical representatives of the
same type. They agree perfectly with each other in all structural characters and
in style of coloration. The males differ only in the intensity of the colour of the
lower parts and in the amount of the white spotting on the upper wing-coverts,
while the females are hardly distinguishable. Altogether, they form a natural
group, and are certainly congeneric with Scdateria t argentata (Des Murs) ; but
whether these birds can be associated with <S'. vaevia, the type of the genus
Sddteria, is another question. Anyhow, they have nothing to do with Di/sitlininiius
and Ih/pocnemis. The species of the former genus have a much stronger, more
hooked bill, and a shorter tail ; those of the latter are distinguished by a much
broader and much more flattened bill, etc.
The four recognisable forms have to stand as follows :
1. Sclateria schistacea schistacea (Scl.).
Jhipneiiemis schintacca Sclater, P. Z. S. 18.'J8. p. 252 [''Rio J.avarri"], descr. orig. ^ ; ScLiter, Cut.
Birds XV. p. 287; Berlepsch & Stolzmann, P. Z. S. 18',)fi. p. ;i84 [La Merced, La flloiia,
Borgofia : Central Peru].
Wing. Tail. P.ill.
1. Mus. Brit., 6 ad., Rio .Tavarri,ex Stevens, doll. P. L.
Sclater. Type of sjiecics 03 49 17,; mm.
2. Mns. Brit., S ad., La Gloria, (Jhanchamayo, C. Peru,
January 23, 1891. Kalinowski coll CS r,l 19 „
3. Mus. Berlepsch, S ad., Borgofia, ( 'hanchamayo, C. Pern,
May 19, 1891. Kalinowski coll. . ". . .08 5i 2U „
4. Mus. Tring, <S ad., Pozuzo, lIuAunco, ('. Pern,
March 191)4. W. Hoffmanns coll 05^ :,n 17 „
.^. Mus. Berlepsch, ?, Marcapata, Cuzco, S.K. Peru,
October lU, 1899. 0. (iarlcpp coll 09 .57 29 „
C. Mus. Berlepsch, ?, Marcapata, Cuzco, S.E. Peru,
November 9, 1899. 0. Garlopp coll. ... 70 ^u 20 „
* Sclater, Cut. Birth xv.
f Sclttleria (Iberliol.-^er, I'rni: Ariiil. Philad. ISySt. p. 209 [nom. emcml. for nrtrroniemh Scl. pre-
occupied].
( 345 )
(? ad. (type at' speciesj. Al)ove .and lieneatli iinilorm dark Iduish scWstaceous.
Wings and tail sooty blackish ; upper wing-coverts with ver}' small white a]iical
dots ; remiges and rectrices exteriorly slightly edged with schistaceous. Bill black.
The specimens from Chauchamayo have rather longer bills and the lower parts
a shade darker ; the J from Poznzo, however, is absolutely identical with the type.
la all four males the under surface is uniform schistaceous and scarcely paler than
the back. The bill is entirely black in Nos. 1—3 ; while No. 4 has the lower
mandible a little paler, more brownish.
? ad. E.xactly like that of S. s. leiicostijjma and S. s. suhphimhrii, and onlv
(lifiTering in the less rufesceiit, more olive-brown tint of the upper parts and in tlic
top of the head being of the same colour a-i tlie back (instead of dark schistaceous).
Tail rather longer. Upper mandible l)lackisli, lower jaw horn-Tjrown (not whitish,
as in the following subspecies).
Nos. .5 and 0 are certainly the females of the present form, although their tail
is rather longer than that of the males (Nos. 1 — 4). I have examined a female of
S. s. subphimbca with the tail quite as long.
2. Srlatc/ia xeliistacra .vilijjhimhea (Scl. & Salv.).
D/jsHJifnmius ('.) suhphiiiiheux Sclater & .Salvin, P. Z. S. 1880. p. 158 [Saray.i^u & Zamora : East
Ecuador ; Iquitos : N.E. Peru].
DijsHhamnns plumlii'iis (not Wied !) Sclater, P. Z. S. 1858. p. 457 [Zamora : East Ecuador].
D. snhplumbeiix Sclater, Gil. Uh(h Brit. Mus. xv. p. 22(1 [East Ecuador, N.E. Peru].
ITrln-iicnemia UurDxIiijiiia Sclater, /. c. p. 270 [part. : specimens 1 — p, e.v; Bogotrt coll., examincil
by me].
1. Mus. Brit., i ad., Sarayacu, East Ecuador (C. Buckley
coll.). Type of I), stibplumbeus Scl. & Sal v. [spec, c
of Sclater's list]
2. Mus. Brit., S ad., Iquitos, August 27, 1878 (AVhitely
coll.) [specimen a of />. subphimbem']
3. Mns. Brit., S ad., Iquitos, October 10, 1878 (Whitely
coll.) [specimen b of D. subpliimbeuH]
4. Mus. Brit., c? jr., Zamora, E. Ecuador, January 1858
(Eraser coll.) [specimen e of J>. siibplinnbeHs]
5. Mus. Brit., S juv., Saraya^n, East Ecuador {V,. Buckley
coll.) [specimen d of ]>. subplumbeus]
0. Mns. Brit., S ad., Bogota coll. [specimen /« of
Sclater's list of Hctcrocncmis lewoMii/m/i^
7. Mns. Brit., S ad., Villavieencio to Me<lina, Llanos of
the B. Meta, Colombia, 800—1500 ft. (T. Wlieeler
coll.)
8. Mus. Berlepsch, c? ad., Bogota coll
>! „ <S ad., ,,,,....
„ c?jnv., „„....
''_• !> „ ? ad., „„....
1~- „ „ ? ad., „„....
Wing. T,ail. Bit
10
(? ad. Differs from that of .S'. ,s. scZ/ix/acfa only in having the lower mandililc
whitish, and the general plumage, especially the under parts, decidedly lighler
scliistaceous grey.
( 34f, )
There is absolutely no difference between the si)ecimens from Ecuailor and
N.E. I'ern and those from Bogotd, tliongh tlie latter have been referred by
Dr. Sclater to //. IciiooHtUjmd. In two exaiuj)les (Nos. :? and T of the above list)
tlie white ajiical s]iots on the npjier wing-coverts are (juite as larjje as in fi. x. h'liro-
sti<imfi, while in all the otlicr males tjiere arc only very small dots, jnst as in
6'. «. sc/a'staccd.
Nos. 4, 5, and lo arc immatnre males, showinj;- some remains of the jnvenile
]ilnma;;e. The flanks and nnder tail-coverts are still rnfons brown, and most of the
ijnills rnfcscent brown (instead of black with slaty margins), eto.
? ad. Easily distingnished from that of S. .9. scJiistacea in having the lower
mandible whitish and the top of the licnd slate-grey, sharply defineil against the
rnfons brown back : from that of the following snbsjiecies by its slightly shorter
beak and rather shorter tail.
3. Sclaterifi scl/ixt/irca leucostigma (Pelz.).*
Pereiinxl.ilu lnirn<ttir/mn Pclzeln, Ziii- Onnlli. Bmxil. ii. (ISfiS), pp. Sfi, ICiO [Barra dn Rio Negro,
Mavabitana.s, Rio Vaupe, Cayenne].
TTdrrncmmU simphj- Sclater, P. Z. S. ISGS (probably pulil. in 1809), p. 573 [Maroni R.. Surinam].
Wing. Tail. Bill.
1. Mus. Viudob., No. 15438, t? ad., Barra do Rio
Negro, October 5, 1S30. Nattcrcr coll. Tyjie of
P. leiicostigma Pelz Tij 00 2(iJ. mm.
2. Mns. Vindob., No. l.")430, S ad., same date and
locality
3. Mns. Vindob., No. loSll, S ad., 8t. George, Oyapoe,
Cayenne, March 13, 1800. C. Jelski coll.
4. Mns. Vindob., No. 1;-)43T, ? ad., Barra do Rio Negro,
•Tannary 0, 1833. Natterer coll
5. Mus. Vindob., No. 15410, ? ad., Rio Vanpo, Jnly 15,
1831. Natterer coll .' .
0. Mns. Tring, r? ad., Iponsin, Cayenne, .Tannary 5, 1903.
G. K. f'herrie coll. .......
7. Mns. Tring, S jnn., same locality, December 24, 1002.
8. „ „ ? ad., „ „ .Tannary 5, 1003
0. Mns. Berlepsch, S ad., Bartica Grove, Brit, (iniana,
September 27, ISSO. II. Whitely coll. .
in. Mns. Berlepsch, ? ad., Bartica Grove, M.iy 31, 1S80 .
11. Mus. Tring, ? ad., Mernme ]\rount., .Tnly 20, 1881.
[IT. ^Vhitely jun., coll.] OS 58^ 21 .,
I'esidcs these specimens, I have examined the series in the British IMusenui.
including the typo of 7/. simjth'.r from Surinam.
c? ad. Agrees with .S'. .9. suhpliimhca in having the lower mandible whitish,
but differs in the much paler colonr of the lower parts, which are light grey,
passing into whitish on the chin .and middle of X\w. abdomen. The blackish njjper
wing-coverts have always large ronnded white apical spots, like the two specimens
of & .?. snbpliimheit mentioned above (Nos. 3 and 7 of the list of 5. x. .vtbjditmbcti),
* Pi'lzeln"s name appears to have the priority. I'art ii. of t1\e Oriiilh. Urasil. in reviewed liy Newton
in the Jaiitiari/ niinil>er of tlic I7iis. while the l.xst part cif (he /'. Z. S., eont<iining Selater'.i de.xcription.
rertaiiily did not appear hefore Jlaioh or April ISGil.
( M7 )
Anioiiy; the eleven dS iid. ol' S. s. IciKO/stifima coiii]i:irecl by lue, there is not
a single specimen thut luis snch small, dot-like markings on the wing-cuverts as
S. s. schistacea.
? ad. Top of the head dnsky slate-colour or dark slate-grey, in strong contrast
to the dark rnfescent brown back. Upper wing-coverts a little darker brown than
the latter, terminated by large, ronnded f'nlvous spots ; (|uills dusky, dull rufescent
brown on the outer webs : tail dull blackish. Lores greyish ; cheeks, malar region
and oar-coverts slate-grey, rather lighter than the pileum. Whole lower snri'ace
deep ferruginous, nearly whitish on the chin ; flauks washed with rufescent brown.
U]iper mandible blackish, lower one whitish.
The specimens from British Guiana, Cayenne and iSurinam are practically
identical with Pelzeln's types from Barra do Rio Negro. Hence, //. simjjlex
becomes a synonym of P. Icucodiyma.
4. Sclateria schistacea saturata (Salv.).
Hctcriicneijus saturata Salvin, Ih'ts, 1885, p. 427 [Roraim.!, Brit. Guiana].
Wing. Tail. BUI.
1. Mus. Brit., S ad., lloraima, 3501) ft. January 22, 1884.
H. Whitely coll. Type of //. saturata Salv. . . UO 59 20 mm.
2. Mus. Brit., ? ad., Roraima, 3500 ft., Jane 29, 1884 . 06 50 19^ „
S ad. Agrees with S. s. schistacea in the dark schistaceous colour above and
below, and in having the bill entirely black, but dilfers in its rather deeper black
wing-coverts with much larger white apical sjwts, which are quite as large as in
& 5. Icucostigma.
? ad. Not distingui.-shable i'rom that of S. s. Ici/costit/i/ta ; bill perhaps a little
shorter.
Tlie distribution of the four subsjjccies is as follows :
1. ''Sclatcria schistacea schistacea (Scl.).
Central Peru: La Merced (2000 ft.), La Gloria (3200 ft.), Borgoiia (2(iOi) ft.)
[Kalinowskl coll.] ; Pozuzo, prov. Huanuco (2400 ft.) [\V. Hoffmanns coll.]. —
S.E. Peru: Cuzeo, Marcapata (30011 ft.) [0. Garlc]ip coll.].
Thus, this bird is an inhabitant of high elevations, and 1 doubt the correctness
of the original locality : " Rio Jacarri."
2. S. schistacea sttbplumhca (Scl. & Salv.).
East Ecuador: Sarayai;u (Buckley coll.); Zamora (Fraser coll.). — y.E. Peru:
Iquitos, on the Amazons (11. Whitely eoll.j. — Colombia: Llanos of the Rio Bleta,
between ,S0() and 1500 ft. {\\ . T. Wheeler coll.) ; Bo<iotd coll. (Mus. Berlepsch,
Brit).
3. S. schistacea leucostignm (Pelz.).
Xorth Brazil: Barra do Rio Negro, Marabitanas, Rio Vanpe (Natterer coll.),
— Caijemie : Ipousin (Cherrie coll.), St. George d'Oyajioc (Jelski coll.). — Surinam :
Maroui River (Bartlett coll.). — British Guiana : Bartica Grove, Camacusa, Takutu
River, Merume Mountains (H. Whitely coll.).
4. 6'. schistacea saturata (Salv.).
British Guiana : Roraima Mountains (3500 ft.) [H. Whitely coll.].
( •^^>^ )
,., (Terenura melauoleuca I'elz. 1 tt „• i, -i v i , ^' i
43. !, , , , .,, > = Hypocnemis nemileuca Ncl. iV: balv.
( Myrmochanes nypoleucus Allcu J
Hypociicinis hcmilcuiM Scluter & Salviu, /'. Z. S. ISGrt. p. 186. dcscr. oiig, (J [Lower Ucayali\
Terenura mclimuleuca Pelzelu, Zur Oni. liraail. ii. (18G8), p. 157. descr. orig. ^ [Borba, Rio
Kladcira].
Mi/niiovli<iiKii Iiy/iuleucKii Allcu, Bull. Anur. Mus. ii. (ISS(I), p. 05. dcscr. orig. (J [Reyes, N.E.
Bolivia].
Wing. Tail. Bill.
1. JMus. Brit, d lul., Lower Ucavali, June 4, 1805.
E. Bartlett coll. Type of s[iL'cit's . . . ,
•2. Mus. Viiidoli. c^ ad., Borba, July 1829
3. „ „ J ad., ., August 1829 .
Natterer coll. Types of T. mrhuioleuca, Vclx.
Thanks to the kindne.ss of my friend l)r. von Loriiiz, of Vienna, 1 was enaliled
to examine Pelzeln's specimens, and I found them absolutely identical witli tiie
type of//, hemih'uca in the British Museum. Perhaps they have a slightly longer
bill, but, as will be seen from the measurements given above, the difference is hardly
apparent. Mr. Allen's careful description leaves also no doubt that his iL/rmochanes
hi/jjoleucus is nothing else but the species in ijuestion.
It need not be added that this bird has nothing to do with Terenura or with
I'ljrujlena, but it agrees structurally with some species of the genus Ilijijocnemis,
except that the bill is rather flatter and rather more compressed towards the tip.
However, //. melanopoijon and //. macuUcauda are intermediate in this respect, and
therefore I do not venture to sejMrate //. hemilcuca generically from Ili/pociicmin.
In the Cat. of Birds sv. p. 291, Mr. Sclater gives the description of tlie ? of
//. hcmilciica, but there must be some mistake here, as neither Bartlett nor any
other naturalist collected specimens of this sex.
The British Museum possesses only the single S ad. upon wliich Sclater and
Salvin based their original description.
The range of //. hemileuca, thus far known, is as follows :
E. Peru : Lower Ucayali (Bartlett); y.E. Bolivia: Reyes (Rusby); -X. Brazil :
Borba (Natterer).
44. Hypocnemis stellata tScl. it iSalv. = Dichrozona cincta (Pelz.).
C'l/jiliorliiinis (Micrucerciil us) cincliix Peizela, Zur Oniilli. Bnts'd. i. (1867), pp. 47, 65 [Borba: Rio
Madeira ; S. Joaquim : Upper Rio Negro],
JIi/poc7iemis slelUita Sclater & Salvin, P. Z. .S. 1880. p. Ilil) [Saraya^u : East Ecuador].
Dkhro-oiia zoiioiuila Ridgway, Pror. U. S. Mus. x. 1887 (1888), p. 524 [Santarem ; Lower Amazons].
Cf. Hcllmayr, ./««)■«. /. Ornilh. l',)03. p. 536.
Wing. Tail. Tar.sus. Bill.
1. Bins. Viudolj. 10350, S ad., >S. Joanuim, July 28,
1831. Natterer coll. Type of C. Ci'«c<«s Pelz. . 01 25 20 19 mm.
2. Mns. Vindub. 10351, ? ad., Borba, January 4,
1830. Natterer coll. Cotype of C. cinctuts Pelz. 5S 24 20 18i „
3. BIus. Brit. av. jr., tSarayavu, E. Ecuador. C Buckley
coll. Type of //..>■?(■//'//'? 8cl. & Sal v. . . . 02 27 — 19 „
4. Mus. Brit. av. juv., Saraya<.'n, E. Ecuador. ('. Buckley
coll. Type of //. «;cZ/a<a Sol. & Salv. . . . 58 25 — \~ ,,
In the paper quoted above I pointed out the identity of Dichrozona zonoiiota
and Cyjjiwrlunus cinctun. Through the kindness of Dr. von Loreuz, of Vienna, I
( 341) )
was euiililed to comitare the fypes of tlic latter species with those of If. stellata
in the British Museum, and fouud them to be ideutical.
No. 1 is au adult male. The spots on the lesser and median upper wing-coverts,
as also the cross-baud on the uropygium, are pure white. The female (No. 2) has
a slight creamy wash on the latter, and the aiiical spots on the wing-coverts are
pale fulvous.
The typical specimens of //. stellata are both immature. No. 3, in more
advanced plumage, agrees perfectly with Natterer's female. The only differences
I cau find consist of the very slightly darker upper surface and the more fulvous
_ tinge of the rump-liand, both being evidently' signs of immaturity. No. 4 is a
very young bird in fluUy plumage, with the markings everywhere less pronounced
than in the adults, and with but few very small black spots on the foreneck.
Strange to say, neither in the original description of 11. stellata nor in the
Cat. Birds xv., is any mention made of the coloration of the tail.* All the
specimens have the two outer tail-feathers and the two basal thirds of the outer
web of the third pair pure white, as correctly described by Mr. Ridgway.
45. Myrmotlierula guayabambae Sharpe = M. atrogularis Tacz. ? !
Mijriiiii/lienila ainnjularis Taczaaowski, P. Z. S. 1874. p. 137 [Amablc Maria and Monterico, Central
Peru. — Mus. Uuiv. Warsaw],
J/. tju<ii/ab'nnbae Sharpe, Hull. Brit. Oni. CI. xi. no. l.\xiv. (I'.IOO), p. 2. descr. J [Guayabamba,
North Peru].
Specimen typ. in Mus. Brit. : ¥ ad., Guayabamba, 4U0U ft., August -JO, ls94.
0. T. Baron coll. Wing 53, tail 39, bill 15J mm.
Tliis example agrees perfectly with a female of M. atro(/iilaris from Huambo,
N. I'oru, ill the British Museum. Wing 53, tail 37, bill 15 mm.
J/, guayabambae has been described by mistake. Sharpe never referred to
M. atiogularis, but compared his bird with M. fulciventris, to which it is not <it all
related. M. fulciventris belongs to t^uite another group, in which the sexes are
nearly alike, differing only in the colour of the throat. The ? of J/, atrogidaris
bears, it is true, a su[)erficial resemblance to the S of M. fulciventris, but difi'ers
at a glance in having the back dark olive grey (not brownish), the outer margins
of the remiges olive grey (not rufescent), and the apical sjiots on the upper wing-
coverts pure white (instead of fulvous).
4r.. Picumnus macconnelli Sharpe should be P. cirratus macconnelli Sharpe.
PicumHiis maccuniicUi Sharpe, Bull. B. 0. C. xii. (Oct. I'JOl), p. 4 [Brilisli Uuiana].
I'ifiiniiiiiJi amuzvnicus Sncthlagc, Oni. Mimhcr. xiv. (l&UG), p. GO[Dibtrict o£Par;i : Marajo.aud Wonte
Alegre].
Wing. Tail. BUI.
1. Brit. Mus. (c?; ad., British Guiana. McConnell coll.
Type of 7"'. Ma«-y///rt'/// Sharpe 54 3U 14 mm.
'Z. Mus. Tring, i ad., Boavista, near Paril. Steere coll. 55J 32 13 „
3. „ „ ? ad., „ „ „ „ „ 55 32 13 „
In the original description Ur. Sharpe com]iared the species with /'. steindachneri,
' In >.u. 3 the outermost pair is wanting.
( SoO )
wiiirli lit'loiigs tu quite ii iliUVrriit group, havinv the chest spotted with white!
while ill 1'. mnccdniiclli tlic wlidle lower surface tVoiu the chin Ui tlie under tail-
fuverts is closely ijarrcil with black auil white, just, as iu P. cirratus. There are,
however, several small dillereiices whiih justify the separatiou of the uortherii birds
as a subspecies. Comiiared with a series of 20 adult birds of true P. c. cirnitns
from Kio, Espiritn Santo and S. I'aulo, the three specimens have a slightly larger
and stronger bill, the back of a darker brown, and the black bars on the lower
surface much broader and deeiicr black. The cheeks and ear-coverts, too, are
darker, more blackish. The other distinctive characters pointed out by Miss
.Snethlage do not hold good, some of my specimens from S.E. Brazil having,
the bill entirely plnmbeous, and the forehead ([uite as pale buffy whitish as the
northern form.
The birds from Para being practically identical with the type from P.rilish
Guiana, P. umazoiiicus becomes a synouym o*' Sharjie's name, and the ibrm has to
stand as P. cirratus maccoii/ieUi ISharpe.
47. Ortalis squamata (Less.)
OrlulUla sqnamtla Lesson, Dkl. Sci. Nat. Ivi. (1829) p. VJ5 [" Brusil'] ; idem, Traili d'On,. lK:il.
p. 481 ["du Cresil"] ; Berlcpsch & Ihering, Zcilsclir. ijck. Oni. ii. (1885), p. 17'J [Rio graude
do SulJ.
1. Mas. Paris, spec. typ. labelled as follows : " Ortalida squamiUa (Less.)
A. de S. llilaire, Bresil." Ou the bottom of the stand: " 2vu. \V.)Vi. I'.resil,
Ste. Catherine. M. de St. Hilaire, li<22.— Vrtali(/a sqiiainata Less., type."
Wing 209, tail 240, bill 25i mm.
2. Mus. Triug, ad., Blnmeuau, Santa (Jatharinu. \\'iug l»n, tail 220, bill
25 mm.
3. Mus. Tring, ad., Taqnara do Muiido Novo, Rio graude do Sul, July ILI, 1>>S3,
11. V. Ihering coll. Wing 18(3, tail 217, bill 24 mm
Lesson's type agrees perfectly with No. 2, dillcriug only in its rather larger
size. The determination of the species by Count Berle[isch and Mr. Ogilvie- Grant
thus proves to be correct. As clearly pointed out by the former authority, the
species is quite distinct from the Bahia form named 0. alliiveiitris in the Cat.
Birds xxii. p. 508. In the latter bird the whole breast and abdomen are uniform
pure white, while in O. S'juamata they are wood-brown witli distinct greyish apical
margins. The thighs, dirty white iu U. albicentris, are dark brown, or rnfesceut
browu, in the South Bra/.iliau form. The top of the head is but a shade more
rnfesceut olive brown than the back (instead of bright brownish chestnut as in
(). ulbiveiitris), and there is scarcely any trace of the rufous bntf forehead and
superciliary stripe to be seen in the Bahia form. The feathers of the mantle have
uo pale margins, the feathered jiortiou of the lower throat is uniform olive brown
(not at all spotted with greyish white), and the foreiieck decidedly darker, more
blackish brown with definite greyish white margins (not dark grey with whitish
tii>s as in the Bahia form). Altogether, 0. squamata resembles much more
O. (jatlnta from the Upper Amazons, but still it is distinguishable by its unspotted
(dive browu lower throat and uniform rufesccnt lirown upper part of the head
(not blackish browu or dusky with greyish spots on forehead and superciliary
region).
0. squamata is only known from Santa t'alharina (whence I received a
( 'i/anocorax caendcus (Vieill.)
S pod to mis luiifonnis (Scl. & ISalv.)
Sicalis a. ancnsis (Kittl.)
Compsospiza baeri (Oust.)
p. 30.-.
p. 307
p. 308
ji. 300
( 351 )
specimen* directly from a collector in Blumeiiuu) and liio yraude do Siil. No
Ortatis~%\i&ciidf, has as yet been recorded from Paraiiil, S. Paulo, and Uio.
In Minas Geraes, Bahia, and Pernambuco, the white-bellied species {O.ulbiccidris
auct.) occurs.
The following species are discussed in this jiaper : —
Cijanocorax inexpectatiis Elliot "|
Ci/anocorax keckelii Pelz. j
Anthiis latesceiis Puch.
Uaplospiza uiti/onHin Scl. & Salv.
CrUkaijra hilarii Bonap.
Buarremon baeri Oust.
Tachi/phonus cklorictcrus Vieill.
Orthogonys viridis (Spi.x) et auct
Ncmosia cknjsopis Scl. & Salv.
Arremon callistus Oberholser
Arremon deiillii Des Murs
Arremon wuchereri Scl. & Salv.
Saltator azarae D'Orb.
Saltator albicollis Vieill.
Folioptila lactea Sharpe
Curruca olivacea Less.
Jiacnis salmoiii Scl.
Knipolegus unicolor Kaup
Muscicapa cristata Less.
Mt/scisaxicol(i stria ticeps Lafr.
D'Orb.
('iiipolegus cinereus Scl.
Flapalocercus hollandi Scl.
Ilapaloccrcus striaticeps Salv.
Leptopogoii tvislis Scl. & Salv. 'i
Muscicapa, angustirostris Lafr. &, ,
D'Orb. i
Leptopogoti godmani Scl.
Capsiempis caudata Salv,
=; (hthogoiii/s clilorictcrus iW'KxW.) . ji.
= T/d>/popsis s. sordida(Lai'i:&'D'Ovh.) ji.
= Arremon p. jJolioHOttis Bouap. . . p.
= Arremon polionotus decillci Des Murs p.
= Arremon Jlavirostris Sws
= Saltator coeridesccns azarae D'Orb.
= Saltator guadoloupensis Lafr. .
= sp. optima ! . . . .
= Vireo chici (Vieill.) .
— 2\'emosia salmoni (Scl.)
= Knipolegus cganirostris (Vieill.)
= Knipolegus ci/anirostris (Vieill.)
^= A n/j/olegusstriaticeps(Ls,h. &I> Orh.) p.
= Hapalocercus s. sclateri (Oust.)
= Hapalocercus sclateristriaticepsHiilv.
I I'hgUosca.rtes rentralis angustirostris
'[ (Lafr. & D'Orb.)
: Pogonolriccus ophthalmicus Tacz.
300
310
310
311
ZVZ
313
314
315
31fi
317
317
317
31s
31 s
_/;
r. ; ■ T> 1 I IT t f = " Serpopkaoa caudata (Salv.)
>icrpopnagaorenocensisiJi.'Y\.^lVM{.] \^ t^ f - \ 'j
ilgiarchus tricolor Pelz.
Mgiarchus graxilirostris Pck.
Xenopipo subalnris Godm.
Chloropipo holochlora Scl.
Muscicapa lateocephala Less.
Muscicapa luteocephala Lal'r.
Psaris erijthrogenijs Selby
Attila brasilicnsis Less.
Attila bolicianus Lafr.
Lanius unirufus Pnch.
Upucerthia bridgesi Scl.
Synallaxis setaria Temm.
I .\fgiarchus tuberctdifer (Lafi'. &
^\ D'Orb.) . .' .
= Chloropipo unicolor Tacz.
= sp. optima !
= Ileterocercus luteocephalus (Less.) .
= Xeopelma aurifrons (Wied)
= Titi/ra impiisitor eri/throgeni/s {i^ol\i\ ) jj.
= Attila brasilicnsis brasilicnsis Less. ,
= Attila validus Pelz. ....
= Attila thamnophiloides (Spix) .
= Upucerthia andaecola Lafr. & D'Orl.i,
= Leptasthcnura setaria (Temm.)
Actually in the Triug Museum.
23
( ■^52 )
Si///alla:ci,'< Mriolata Pclz.
Si/nnUa.ri.s paiirali'n.-ii.sTmiY,. . "I.
Syii. subsjjt'ciosa Salvsul. & Fi'sta J
I'hilyilor certicalis IScl.
Dendrocolaptcs atrirostris Lafr. &)
D'Orb. / ■
Tlmiintfljiliiliia trlstis Scl. it Salv.
TkumiioiiliHii.s jjunctulii/cr Pelz. ")
Tltamiwphibisjuniaiius Ihei'ing' ]
Mi/rmccizn immacnlatu Scl. & 8alv.^
Mi/rmeciza intermedin Cherrie /
Mi/rmecizu stictoptcra Lawr.
M'p'mcciza nigricauda 8alv. it)
Godm. J
Ccrcoinacra hi/pomelaena Scl.
Hijpocnemis schistacca Scl.
Di/!<ithanuni!< suhplunbeus Sc
Salv.
Pcrcitostola Icacofitigma Pelz.
Hctcrocnemis saturata Salv.
Terenura melanoleuca Pelz. "|
Mi/rmochancs hi/poleucus Allen J
llijpocnemis stxllata Scl. & Salv.
Mi/rmotlterula giuiijahambac Sliarpe
PicicmntDs macconnelli Sharpe "|
Picumims amazonicus Suethlage J
Ortalida squamata Less.
p-
= Mgrmelastes exsul exsid (Scl.). . ]).
= Mgrmeciza lacmosticta Salv. . . \k
_ \ Mi/rmeciza lacmosticta iiigricnnil t
I Salv. & Godm. . . .p.
= Ccrcomacra serva (Scl.) . . . j).
= Sclateria schistacea schistacea (Scl.) . p.
& 1 [ Sclateria schistacea suhphimbca (Scl.
/ == \ & Salv.) p.
= Sclateria schistacea lcucostigmn{'PiAY..) p.
= Sclateria schistacea saturata (Salv.). ]).
= lli/pocHcmis hemileuca Scl. & Salv. . p.
= iJichrozona ciiicta (Pelz.) . . p.
= M. atrognlaris Tacz. . . .p.
= Ficumnus cirruius macconnelli Sbarpe p.
= Urtalis squamata (Less.) . . . ]i.
33'.J
3411
343
343
344
340
34i;
347
348
348
34!)
34!J
( 353 )
NOTES ON A SECOND COLLECTION OF BIRDS FROM THE
DISTRICT OF PARA, BRAZIL.
By (J. K HELLMAYE.
IN t|](' l;isl, volume of tlie Woritates Zoologicae I piililished an account of a small
collei'tion of Mrds made by Mons. A. Robert at a place called Igarape'-Assu,
east of the town of Para. Soon afterwards, Mr. W. Hoffmanns, who had been
collecting for some time in Pern, where he discovered the Leantifnl Pipia ei-(]U!sita
Hellm., came to Tring, and I suggested to him to continue the researches in the
Para district so successfully taken up by M. Uobert.
Some time ago the Tring Museum received the first consignment from this able
collector, comprising the results of two months' work. It consists of 42i) specimens,
representing 12(J species, of which no less than 21 are new to the fauna of Para.
Besides four forms new to science, Mr. Hoffmanns was so fortunate as to get large
series of the newly described Compopliaya rohcrti and Hypocnemis vidua, each of
which was represented in Europe only by a single specimen ; and he likewise
succeeded in obtaining a good number of skins of two exceedingly rare Humming-
birds, viz. Agi/rtria nitklij'ron^ and Lopltornis gouldii. Several species, only known
from Upper Amazonia, were found in the district of Para, as well as a few
others supposed to be confined to the Guianas — f. i. Avoccttula recurvirostris.
Mr. Hoffmanns being still at work on the Lower Amazons, I defer general
remarks about tlie ornithology of Paril to a later occasion, and proceed to give the
account of the various sjiecies contained in his collection.
All specimens were collected near Prata, Igarape-Assu, at an elevation of
4.'3 metres above the sea-level. The notes on the colour of the iris and soft parts,
carefully indicated on the label of every specimen, are given with (| notation marks.
We are under great obligations to Herr Hermann Schindler, director of the
railway comjiauy, and to Herr Schrader, Imperial German Consul of Para, for
many acts of kindness and assistance given to Mr. Hoffmanns.
1. Turdus phaeopygus Cab.
Turdus ph<ieoj)>jr/u.i Cabanis : in Schomljurgk, Ruise Brit. Gtiiana iii. (18-18) p. COS [Brit. Guiana] ;
Pclzclu, Zur Orn. Braul. ii. (18G8) p. 93 (Pani) ; Layard, IhU 1873. p. 376 (Para).
2 c?c? ad. and 2 ? ? ad. : 2, 28. xi., 2, 6. xii. 05. "Iris brown." Nos. 01,
218, 246, 282.
They agree perfectly in colour and size with specimens from British Guiana.
2. Turdus fumigatus Licht.
Turdus fitiiiiyaliis Lichtcnsteiu, Ver-^. Duhl. 1823. p. 38 ('■Brasilion") ; Sdater & Salviu, P. Z. S.
1867. p. 068 (Mexiaua Island) ; Layard, Ihis 1873. p. 376 (Para).
A single c? ad. from Prata, 'i. xii. U5. No. 251. " Iris brown."
It dift'ers from an adult <? from Para, collected by Prof. Steere, in having
a distinct white jiatcli in the middle of the lower abdomen. Thereby h ajiproaches
'/'. Iiatuicelli of Upper Amazonia, but in other respects it is very dissimilar. The
specimens obtained by Natterer ou the npper branches of the Kio Madeira (near
Engeuho do Gama, S. Vicente, Rio Guapore) are also typical T. fumiyatus, showing
not the slightest transition towards T. hauxwelli.
( 354 )
3. *Microcerciilus marginatus marginatus (•^(■l).
Ift'Uro'iiemis marghutt'is Suliiter, /*. X. N, 1H55, p. 1-15 [liogutii coll.- t^'pe uxainintHl].
Hetcrnrnemis bicolur Des Miirs : in Castelnau, Vai/age. Oiscaux (1855) p. 51. [No locality ; tliu type
(which I examined) came from the Peruvian Amazons.]
A single S jr., 23. xi. 05. No. 185. " Iris brown."
This is the first record for Pari'i, extending the range of the species from the
upper l{io Negro (Marabitauas), where batterer collected four specimens, to the
Lower Amazons. The skin from Para agrees perfectly with a good series from
Ujijier Amazonia and Bogota collections.
The various iiluniages of this species were not well understood hitherto, and
none of the authors who wrote upon the subject ai)i)ears to have recoguised that the
birds with dusky margins to the feathers of the pileum, throat and chest are
the youug, and those without these edges the adults of oue and the same species.
This is clearly proved by the series before me, where every possible transition
between the two extremes can be found.
In North-western Ecuador M. m. marginatus is represented by a closely allied
form, to which Mr. Sharpe erroneously applied the name M. bicolor. I i>ropose
to call it
Microcerculus marginatus occidentalis n. subsp.
Similar to M. m. inarf/inatus, but the upper parts darker and less rufoscent»
the sides of the body much darker, deep chocolate brown with scarcely any rufous
tinge, and the whole abdomen crossed by broad blackish-brown subterminal bars.
The bill, too, is slenderer, and somewhat longer.
Type in Tring Museum : <? ad., Lita, N.W. Ecuador, 3000 ft. elevation,
Oct. 4, 1899. No. 210. Wing 60, tail 23, bill ITJ mm.
I examined more than twenty specimens of this form from various places in
N.W. Ecuador : Lita, Cachjjacu (3200 ft.), Pambilar (60 ft.), and Cacbavi (500 ft.).
The skins in the British Museum, said to be from Saraya^n, East Ecuador (Buckley),
and described by Sliarpe s.n. M. bicolor,] are perfectly identical with this series.
I feel perfectly sure that they never came from East Ecuador, where only
M. m. marginatus is found. Buckley was a very careless collector, and bis
specimens had no original labels, but were labelled afterwards by Gerrard, almost
iiivariulily " Sarayafn." Bnt as so many species which have only been found on the
western slopes of the Andes by Stolzmann, Goodfellow, Festa, Eraser, etc., and by
Mr. Rosenberg's correspondents, bear this locality ; and as we know that Buckley's
men, Villagomez and lllingworth, collected in Western Ecuador, there can be no
doubt that many of the specimens labelled '" Sarayafn " really came from western
Ecuador.
Dr. Sharjie, when employing tlic name M. bicolor for (his western lurm, was
apparently misled by the superficial resemblance between his specimens from
" .Sarayacu " aud the type of M. bicolor, all being adult birds with jjlaiu white
foreneck and chest. By a singular coincidence all the specimens uf tJJC eastern furm I
before him were immature birds with dnsky margins to the feathers of the i'oreneck
and chest, aud therefore he regarded the coloration of these parts as the principal
difference between the two forms ; but, as 1 have jiointed out above, this is only due
to age, the adults of both forms having the throat and breast unbarred, jilain white.
t Cat. Birds Brit. .ViM. v,i. p. 'i'.I.H.
X Three from Biigoti (incluiling the type of II. margi)uitiis), one from liast J'eni (ux \xTrcaux). two
from I'ebas, aud one from t'Lamicuros, N.E. I'eru.
( 355 )
The type of M. Incolor (from the Peruviau Amazons) agrees in every way with
several aihilts from Bogota. The upper parts and the flanks are of the same
rnfesccnt brown colour, and there are only a few blackish and whitish bars in
the middle of the lower abdomen.
Thus we have two distinct forms, the range of which is as follows : —
(ri') M. margiiiatus mai-ffinatus (Scl.). From Eastern Colombia (Bogota coll.)
sonth through Eastern Ecuador to N.E. Pern and (ientral Peru, and even to
Northern Bolivia (Busby coll. — Amer. Mas. New York) ; east through the Bio
Negro region to Para.
(b) M. marginatiis occidentalis Hellm. Only known from N.W. Ecnador.
N.B. — M. tacniatus Sal v. (tyj)e ex Balzar, W. Ecuador) belongs to a different
section of the genus, having the entire under-surface banded with smoky grey, aud
is a very near ally of .]/. fujiiamnlafiis iScl. & Salv.
4. Thryothorus genibarbis Sw.
Cf. Xov. Zool. xii. (190,5) p. 271.
2 t?c?, 1 ? ad., and 1 young ? from Prata : 9, 17. si., 9. xii. U5. Nos. 92, 94,
149, 301. Iris marked as " brown " or as " red."
5. * Granatellus pelzelni paraensis Eothsch.
G.p.pnmensis Rothschild, Bull. Brit. Oni. CI. xvi. (April lOOH) p. 81.
No. 141. (? ad., Prata, 17. xi. 05. "Iris lirown, bill grej'-blue, feet bluish
black." Wing .52^1, tail .501, bill lOJ mm. Type of subspecies.
No. 132. ? ad., Prata, lo. xi. O.'i. "Iris brown, feet and bill grey-blue."
Wing .04, fail 53, bill 11 mm.
The <? of this interesting new form differs from that of 0. p. pelzelni by lacking
the silky white stripe which separates the grey flanks from the rosy red middle of
the belly, the whole sides of the abdomen being uniform dark grey. On the top
of the head the black does not extend over the whole crown, as in the typical form,
but is restricted to a well-defined frontal band and a narrow stripe above the eye,
while the vertex is slaty bine like the back. There is only a small white spot
at the posterior margin of the eye, whereas typical G. p. pelzelni shows a long,
broad, white postocular stripe.
In other respects the Para form does not difi'er. The sides of the head are dee]!
black, the throat is white, bordered below by a number of small black sjiots, and
tlie breast as well as the middle of the abdomen are boautifnlly rosy red.
The ? of G. p. paraensis has the sides of the breast strongly washed with grey,
which is barely indicated in a good series of ? ? of the typical form.
It appears that G. p. paraensis has a very limited range, as the specimens
collected by Mr. Andre on the Caura Biver, Venezuela, agree with the type of
G. p. pelzelni which was obtained by Natterer on the Upper Bio Madeira.
6. Pachysylvia thoracica semicinerea (Scl. & Salv.)
[iri/lojihihis ihnniciriiH Temniinck,'/'/. .7,/., livr. -J'.i (Iki;l>) pi. 173^ fig. l {" Bresil"— coll. Natterer.
Wc substitute liin as iy]i. locality).]
llyhipliiliis scmicincrrii.i Sclater & Salvin, P. Z. S. 18G7. p. 570. pi. xxx. fig. 2 (ParA) : Layard, Ihl/:
161^. p. 377 (Paril).
Two adult c?(?, 1(5, 17. xii. 05. Nos. 3.54, 307. " Iris white (No. 354), whitish
yellow (No. 367)."
( 356 )
The real affinities of this rare form have never been recognised. Mr. Sclater,
in his review of tlie genns,* included it in the section of P./lnvipes. This,
however, is not its proper place, and a thorough study of all members of the genus
proved it to be a geographical race of P. thorarka (Temm.), most nearly allied to
P. t. gnseiventris (Berl. & Hart.), t
It agrees with them in the coloration of the under-snriace of the wing and in
having the forehead and vertex tinged with yellowish olive, the occiput and nape
being pure cinereous. The abdomen is ]iale cinereous, as in P. t. i/i'iseiten/ris.
The Para form diflFers, however, from lioth by having the sides of the head and
neck ashy grey instead of yellowish olive-green, and by the lack of the olive-
yellow jugular band. Yet, in one of the two specimens (No. 307) there is a faint
yellowish tinge to be seen across the chest, thus indicating the close relations to
P. tkoracica. Therefore I have not the slightest doubt that the three forms
with olive-yellowish forehead and grey nape form a natnr;il group, consequently
they must be designated by trinomials.
As pointed out by Berlepsch and Hartert,t P. pectoi-alis (Scl.) is quite a
different species, easily recognisable by having the whole top of the head (from
the fnint to the nape) pure cinereous, the middle of the alidomeu white, and by the
yellow pectoral band being much brighter and extended over the whole chest.
This is an exclusively Amazonian species, ranging from Mattogrosso to Cayenne,
Surinam and British Guiana. In Tring there are more than twenty specimens
from the three latter countries, which I am unable to distinguish from Sclaler's
type (ex Mattogrosso — Natterer).
In another jiaper I intend to discuss the distribution of P. thoracica and its
allies.
7. Dacnis cayana cayana (Linn.)
Miil'irilhi niiiitiiii Ijinn.ieus, Sysl. Nut. xii. 1 (17(5(5) p. .'53(5 [Cayenne : ex Brisson].
Damis anjnnii Sclater & Salvin, P. Z. S. 18(57. p. 570 (P.ira) ; Lay.ar(l, Ih!^ 1S73. p. 378 (Pani)
V. q/iinnrejilitihi Pelzeln, Ziir Oni. Biiifil. i. (18(57) p. 2u (Para).
S ? ad. fnim Prata, 0, 11, xii. (15. Nos. 277, 'M2. " iris brownish red."
8. Cyauerpes cyanea (Tiinn.)
Cf. Nor. Zon!. xii. (1905) p. '27-.'.
A single <? ad. from Prata, 10. xii. (15. No. 3."iS. " Iris brown." F.ill 10 mm.
!•. Coereba chloropyg'a (< 'ab.)
Cf. Nor. Zuol. xii. (1005) p. 272.
1 c?, 3 ? ? ad. from Prata, 7. xi., 7, IG, 27. xii. 07). Nos. 81, 280, 302, 410
" Iris dark brown."
111. Euphonia violacea lichtensteinii (Cab.)
\Frhigina r'lnhie.m Linnaeus, .S'?/«/. Nat. x. p. 182 (1758. — nn locality).]
Plirmixca Lichlriislfinii Cabanis, Juiirii.f, OriiilJi. 18(50. p. :'..'!1 (Cayenne).
Eiiphotm Lkhlimhhiii Pelzeln, Zur Oni. Brasil. iii. (18(59) p. 204 (Pani).
3 c?c? and 2 ¥ ? ad., 25. xi., 5, 9, 14, 19. xii. 0.5. Nos. 205, 272, 3u4, 330,
384. " Iris brown."
Tiic wing measures 54 — 55, the tail 30 — 31 mm. Therefore the sjiecimens
• Ms 1881. p. 308. t Nor. Xool. i\. (1!»02) p. 12.
( 357 )
from Pani belong to the smaller northern form separated by Cabanis as P. licliten.
steinii.
Birds from Bahia, S. Patilo and Paraguay are much larger, the wing measuring
59 — 04 mm.
11. Euphonia cayennensis (Gm.)
Tanar/m cayemmish Gmelin, Si/«/. Nal. 1. ii. (178.^) p. 8fl4 [ex Brisson & Buffon : Cai/nme].
Eii/,Ji„>,;i( ciniuiia (nee Tiiiiaf/m mi/titia Linn.' !) Sclater & Salvin, P. Z. S. 18C7. p. 570 (Par;!).
EiiphuiiK riii/iimi Pelzeln, Zui- Oriiilh. Bruxil. iii. (I8G9) p. 204 (Pan'i).
2 (?c? ad. and 1 ? ad. from Prata, 27. x., T. xii. 05. Nos. 24, 287, 289. " Iris
brown."
Besides these specimens, we possess another c? ad. from Bemfiea, collected by
Professor Rteere. They agree perfectly with a series from British Guiana.
There is a cnrious mistake in the nomenclature of this specie.s. Rclater {Cat.
Birds .xi.p. 81) quotes as the original deserijition : " Tunagra cayana Linn., 6'. N. i.
[ed. xii. 1766] p. :515," but this account refers to Calospiza crnjana = Cidlisfe
cuyana auct. ! ! The Etiphojiin is described on the next page (p. 316) under the
same term, viz. T. caj/ana (ex Brisson : Cayenne). This second name, being
preoccupied by tliat of page 315, must give way to 7'. cayennensis of Gmelin.
12. Calospiza albertinae fPelz.)
A'ric. Z,»,/. .xii. (190u) p. 27.3.
1 cT fere ad., 2."). xi. 05. No. 204. " Iris brown." Wing 70 ; tail 48 ; bill
11 J mm.
Tills specimen is slightly )'onnger than the one sent by Robert. The orange-
rufous shonhler-spot is less extended, the blue of the rnmp and breast paler and
still mixed with pale green, and the chestnnt-rufous of the head duller.
Dr. Lorenz kindly sent me the type belonging to the Vienna Museum, and
I find that the supposed differences in coloration do not exist, Natterer's specimen
having the sides of the body and the throat green like those from Para ; but the
latter have a decidedly larger bill. The type of C. albertinae, however, being
immature, a series from the Bio Madeira is reipiired to confirm the constancy of
this slight difference or otherwise.
Dimensions of the type: S imm., Salto do Girao, Gctober 8, 1820. Wing 71 ;
tail 48 ; bill lo^ mm.
13. Ramphocelus carbo carbo (Pall.)
Laii'mn Carhu Pallas in Vroeg, Oil. nih. d'Oismu.c, Aihimhral. p. 2 [17G-t.— Surinam].
Ramphoeel IIS jactipu auct.
Cf. Ni,r. Zni'il. xii. (190S) p. 274.
A single cJ ad., 17. xii. 05. " Iris brown." No. 370.
14. Tachyphonus surinamus insignis n. subsp.
T. surin/imiix (nee Linnaeus!) Selater & Salvin, P. Z. S. 1«G7. p. S71 (Pani) ; Pelzeln, Zui- Oniilli.
Brusil, iii. (1869) p. •-'l.'. (part. : Borba, Pani) ; Hellmayr, Nor. Zooh y\\. 1905. p. 275 (Parii).
S ad. and ? ad. from Prata, 28. xi., 0. xii. 05. Nos. 219, 305. " Iris brown."
Since I recorded Robert's specimens, the Tring Museum has received several
6S ad. lif true T. s. surinamus from Surinam which agree with a large number of
( 358 )
skins from British Guiaua, Uiianoco (Orinoco delta) and the L'aura Itiver iu having,
on the sides of the chest, a small white patch slightly washed witli bufi'y towards
the inner edge.
In three 6S ;id. i'roni Para tiiis ])afcli is inucli larger, dccj) ochreous yellow
and mixed with ferruginous interiorly. Three c?cj, collected by Natterer near
Borba on the Rio Madeira, are perfectly alike. I propose, therefore, to separate
these birds as a different form.
Type in Tring Museum : d ad., Bemfica, Pan'i. Cnllccted by Professor Steerc.
AVing 87 ; tail TO.} ; bill 18 mm.
Five (?c? ad, from Barra do Rio Negro (Manaos), collected by Natterer, how-
ever, represent typical 7'. .«. sKiinamtm^ agreeing iu all resjiects with specimens
from Surinam, etc.
Two SS ad. from Marabitanas, and another from the Rio Iranna, Ujjper Kio
"Negro, are again diflercnt, and belong to T. s. napensis. I can tind no diflerenccs
between them and a series from the Rio Napo, N.E. Peru and Bogota collections.
This form differs from T. s. siirinamns in the i}atch on the sides of the chest being
much smaller and pure white like the axillaries, and iu having the uropygium of a
darker ochreous colour.
The range of these tliree forms is thus as follows :
fi. T. s. surinamus (Linn.), t'ayenne, Surinam, British Guiana, Orinoco dellii
(Gnanorii; Andre coll. — Mus. Tring), westwards to the Canra River, E. Venezuela,
south wards to Manaos on the northern ))ank of the Amazons (Natterer). \_S 6 ad.
examined : 4 from Surinam ; 0 British Guiana ; Vi, (!aura River ; 1 Orinoco delta ;
5 Barra do Rio Negro.]
b. T. s. napensis Lawr. From Eastern Colombia (Bogota coll.) south through
Eastern Ecuador (Napo) to N.E. Peru, east to Marabitanas, etc., on the Upper Rio
Negro. [(?c? examined: 3 Rio Napo; 3 Upper Kio Negro; 1 Chjavetas, N.M.
Peru ; fi Bogota coll.]
c. T. s. insignis llellm. Lower Amazons, from Para westwards to Borba on
the lower course of the R. Madeira. [c?cj ad. examined : 3 Pard ; 3 Borba.]
A\hile the females of T. s. surinajnus and T. s. insignis are not distinguishable,
that of T. s. napensis can easily be recognised by the much deeper colour of the
throat and breast, which are ochraceous buff (Ridgw. JS'amenr/. v. 7), instead of
being cream-buff (v. 11).
I compared twenty females of the eastern forms with nine of V. .s. naj/rnsis
(3 Marabitanas ; 4 Bogota; 2 (ihyavetas, N.E. Peru).
15. Arremon silens (Bodd.)
Cf. Nor. Zool. xii. 1005. p. 275.
4 6 6 ad. and 1 ? IVmn Prafa, 31. x., 3, 4, 10, 14, 31. xi. In.-.. Nos. 0, 48, W,,
74, 11)1. " Iris brown."
Like the s|)ocinien sent by I\I. Robert, tlie males liavc^ tlie black jugular band
rathrr luirrow, but one is hardly distinguishable in this respect from Guiana
examples.
10. Saltator magmis (Gm.)
Xoi-. Xnol. xii. (1905) p. 27fi.
•Z 66 ad., 7, 12. xii. 0.5. Nos. 288, 321. " Iris brown,"
( 359 )
17. Pitylus grossus (Linn.)
Ltuiii f/riKxn Linnaeus, Si/^l. Xnt. xii. I (I7(i(i) p. :107 ["America": ex Bi-isson : we substitute
Cayputii'. as typical locality].
Pitylus g,-ox>,„ii Pelzeln, }^,ir Oniilh. JSrusll. iii. (18U0) p. J20 (Pard).
d arl., 2 ? ? from Prata, 22 xi., 4, 8. xii. 1)5. Nos. ISO, 264, 2'.i8. ." Iris
brown."
Not (lift'creut from Guianan specimens.
18. Pitylus erythromelas (dm.)
Cf. A"'iii. Znol. xii. (100.5) p. 27C.
2 c?c? ad. and 1 ? , 30, .31. x., 15. xi. 0.5. Nos. 43, 49, 128. " Iris browu."
The male from 30. October is still in moult, wliile the other one has a) read}'
assumed perfect plumage.
10. Pitylus canadensis canadensis (Linn.)
Cf. iVoi'. Zudl. xii. (1905) p. 270.
2 c?':? ad. from Prata, 4, 18. xi. 05. Nos. 71, 151. '■ Iris brown."
20. Guiraca rothschildii Hartl.
Cf. AV. Zuol. xii. (190.")) p. 277.
3 c?c? and 1 ? ad. from Prata, 28. x., 7, 10. xi., I. xii. 05. Nos. 34, 88, 135,
240. " Iris brown."
These specimens agree perfectly with Bartlett's types, now in the Tring
Museum.
21. Volatinia jacarina splendens (Vieill.)
Cf. Nur. ZimI xii. (1905) p. 278.
2 <?c? imm., 2 c?<? .juv., 4, G. xii. 05. Nos. 254, 257, 280, 281. "Iris dark
brown."
22. Coryphospingus cucuUatus (P. L. S. Mull.)
Cf. N,ir. Znul. xii. (1905) p. 278.
1 (? ad. and 2 ? ? , 10. xi. 05. Nos. 131, 133, 130. " Iris brown."
The (? ad. is, like those sent by Layard and Robert, much paler, more rosy
red on the lower jiarts than examples from Paraguay and S. Brazil. Perhaps tlie
Para form conld be sei)arated, but a good series from Cayenne should be compared.
23. Cacicus cela (Linn.)
Panis Cela Linnaeus, Syd. Nat. x. (1758) p. 191 [" in Indiis "— errore ! As typical locality acceptcil
Surinam ; cf. Non. Zool. xiii. (190G) p. 20].
Cacicus 2>ersicus Sclater & Salvin, P. Z. S. 1867. p. 573 (Parii).
Cassicus persicus Layard, Ibia 1873. p. 381.
2 <?(? ad. from Prata, 10, 20. xi. 0.5. Nos. 101, 3lo. " Iris light blue."
Not clift'ereiit from Surinam and (Guianan shins.
( 3G0 )
24. Cacicus haemorrhous haemorrhous (Linn.)
Cf. Xov. Zonl xii. (1905) p. 279.
?. SS ad., 1 ? imm., 7, 9, 13. xi., 2\. xii. nf). Nos. 84, OT, 115, 308. "Iris
light, l)liu'."
2."). Cassidix oryzivora (Om.)
Orinlus or;izirn7-iis fim<'V\n, .S;/s/. Nut. 1. i. (1788) p. IWll [ex Liitliani Oaypnnp].
Cussiilir uripirnni Sclater &',Salrin. P. Z. S. 18i;7. p. IhV, (I'aiVi).
C'Mxirli.r atrr Pelzeln, Ziir Onidh. Bmxil. iii. (18i;9) p. 201 (Para).
A single ? ad., 2."). xi. 0."). No. 203. " Iris pale yellow."
26. *Colopteryx galeatus (Bodd.)
Mnlanll,, gahata lioddaert, Tah) . PI. r>il. 178.3. p. 24 [ox Daiibcnton, PI. ctil. ?m fiir. 1. -
Cayenne].
Cnluplen/T 'imrimtiiK Riilgway, P. U. S. Mns. x. (1888) p. .^119 (lesnr. orig. ? [Santavcm].
1 S ad., and 2 young <? J : 4. xii., 0, 14. xi. n.'). Nos. 93, 117, 200.
" Iris white, or whitish yellow." In addition to these, we possess an adult cJ,
collected by Professor Steere near Bemtica, Para.
The adult males differ in no way from a very large series of the same sex from
Cayenne, .Surinam, Orinoco, Caura, and British Guiana. The characters given by
]{idgway as distinguishing his C. inornntiis are sira])ly those of the female. We
have a good many sexed ? ? from the above localities, and all dilTer from the
males in their shorter, less conspicuous crest, smaller size, and in having the throat
less, sometimes not at all streaked with greyish. The back is not brighter green in
the Para specimens ; on the contrary, some of our Cayenne skins are decidedly
brighter than the latter. Natterer's specimen from Manaos (Barra do ]{io Negro),
which I examined, is also exactly similar to others from Cayenne.
Wing. Tail.
3 c?c? ad. from Para 40,40,47 40i, 42, 42 mm.
2 (JcJ jnv. „ „ 44i, 4;-) 37,38 „
10 iS ad. IVoni Siu-inam, Cayenne, and liritish
Guiana ........ 40—48 40—44 „
6 ?? and 6S .juv. from Surinam, Cayenne, and
British (Jniana ....... 42—45 35—31" „
This species has not before been recorde<l from I'ara.
27. Mionectes oleagineus oleagineus (Lcht.)
Muxricupa oleuginea Lichtenstein, Vi-i-y.. Diihl. p. 55 (1823. — Baliia).
Mioiiecles oleagineus Sclater & Salvin, /'. Z. S. 1SC7. p. 577 (Para) ; Layard, This 187.3. p. ,382
(Pard).
2 cJJ, 2 ? ? : 10, 21, 23. .\i., 14. xii. o5. Nos. 99, 171, 188, 34.5. "Iris
brown."
These specimens agree with others from British (4niana and IJaliia,
( 361 )
28. Ornithion pusillum pusillum (Ciili. and Heine).
Mijiopnti>n>ii!<'iUa Cabanis & Heine, Mas. Ut'iii. ii. (1K.09) p. .S.S [Cartagena, N. Colnmbia].
Camjilodomn fl'iriniitir Sclater & Salvin, P. Z. S. 1867. p. 577 (Mexiana Isl.— specimen in Brit.
Mus. examined).
(Jniilhioii inciiiwsmis (neo Wied ! ) Layard, Thh 1870. p, .B8li (Nazan', near Pani. — specimen in
Brit. Mus. e.xamined).
A .single t? ad. frnm Prata, IT. xii. O."). No. 309. "Iris liglit brown."—
Wing 51 ; tail 42; bill In mm.
This bird agrees with a series from Surinam, while r>ogot!'i skins liave the eap
of a darker, more brownish tinge. E\amiiles from the Orinoco valley and from
the Canra lliver present a large amonnt of variation in the colour of the crown.
lu some of them it is quite as dark as in Bogotii skins, but others are scarcely
distinguishable from those of Surinam and Para.
If the latter represent (>. napacinn Hidgw.,tl doubt the jiossibility of separating
it from O. p. pusillum.
20 *Elainea gaimardii (D'Orb.)
Miixciaipnrn Giiimanlii D'Orbigny, Vni/ni/r, (Hsrnii.r (piil)l. bctw. 1838 and 1S47) p. ?>2C) [Yuraoares,
N.E. Bolivia. — Type examined].
1 cJ ad. and 1 ? ad. : 2, 0. xii. 05. Nos. 243, 270. " Iris brown."'
These specimens are not different from others from Brit. Guiana, the Orinoco
valley, etc.
30. Myiozetetes cayanensis cayaneusis (Linn.)
Cf. Nnr. Znol. xii. (1905) p. -29:!.
A single ? ad., IS. xi. 05. No. 150. " Iris brown."'
It agrees perfectly with a large series from Cayenne, Brit. Guiana, etc.
31. Craspedoprion olivaceus (Tcmm.)
RhiliirhnnjeUis ulimceiiM Hellmayr, Nnv. Zovl. xii. (1905) p. 293.
One c? ad., 23. xi. 05. No. 186. " Iris brown."— Wing 70 ; tail 07 ; bill 15^ mm.
This specimen is fully as large as examjiles of true C. olicacrii.'i from Bahia,
and differs only in having the fulvous margins to the upper wing-coverts rather paler
and narrower. This slight diH'erence should lie confirmed by a larger series before
sej)arating the Para form.
32. Rhynchocyclus poliocephalus sclateri Ilellm.
R. p. srhileri Hellmayr, Vfrli<iiiill. '.onl. hi,lim. Gmell.-ii-h. Wien 1903. p. 207 dcsor. orig. [Rarra do
Rio Negro].
R. suljihiiresrnis (ncc Spix !) Sclater & Salvin, P. Z. S. 18G7. p. 578 (Pani).
R. mci/acrphiihis (nee Swainson !) Layard, ////.s 1873. p. 383 (Pani).
One S ad. from Prata, 21. xi. 05. No. 172. " Iris yellowish white "—Wing 54 ;
tail 44; bill 12 mm.
This example agrees perfectly with a large series from Brit. Guiana, Surinam
and the Orinoco region. Tlu! chin and throat are always strongly suft'iised with
whitish, and the belly is of a pale yellow, slightly underlaid with greyish on
the chest.
t I'ri'e. V. S. Miis. x. (1888) p. 520 (Santarem).
( 362 )
Two specimens oC 7i'. p. jinliorephalus Taez. from Xeberos and the upper
Uca^ali are mncli lirigliter yellow nnderneath and the throat is not mixed with
whitish. Tiio differences in size mentioned by me I. c. do not appear to be constant.
I examined one of Wallace's specimens, previonsly determined as R. milphii-
irscciis by Sclatcr and Salvin, and tlie S ad. collected by Layardjf and fonnd both
to belong to B. p. schiteri.
■■^3. * Myiobius barbatus barbatus (Gm.)
Miisrirapa hiirhain Gmelin, .S>/. Nal. 1. ii. (1788) p. 933 [ex Daubenton, VI. fill. 830 fig. 1 —
C.-iyenne.]
One c? ad. (with lemon-j-ellow crown-patch ) and ibnr V ? or ynnni; c?c? :
9, 20. xi., 2, 3. xii. Nos. 9(5, 160, 245, 248, 249. " Iris brown."
The series agrees witli s]ici:inicns from British Gniana iiud the Canra TJiver,
Venez., excejit that the foreneck is slightly more snffnsed with oclireons. This
colonr, however, is much dnller and jialer than in .1/. /j. mastacaUs (Wied)
from Babia.
34. Myiobius erythrunis eiythnirns Cab.
ilyiiih'nis ert/thruriis Cabanis, Arch. Nritiny. 13. i. p. 249 pi. 'i fig. I [1847 — "Guiana, Cayenne"] ;
Sclater & Salvin, P. Z. S. 1867. p. 578'(Capim River).
One S in monlt, 14. xi. 05. No. 122. "Iris brown."— Wing 5U ; tail 40;
bill S mm.
This bird evidently belongs to the typical fm-ni, and differs from a series of
the {!anra Hiver only in having the back more fidvons brown, less mixed wilh olive
greyish. Topotyi}ical (inianan sjiecimens are not available for comjiaristin.
The distingnisliiiig characters of the western form, .lA. r. fuh-ii/ii/nri.^
Salv. & Godm., have been jjointed ont by Berlepsch iV- llartert, Aor. Zoo/, ix. (1902)
p. 50.
35. Empidonomus varius (Vicill.)
Cf. Xm: Zoo!, xii. (lOO.";) p. 293.
A single ? ad. : 7. xi. 05. No. 82. " Iris brown."
30. Pipra nibrocapilla subsp.
3 S ad. 2 c? jnv. and 1 ? ad. from Praia: 24, 31. x., 12, 20, 29. xi.,
12. xii. 05. Nos. 17, 46, 100, 216, 236, 323. Iris marked as "whitish" or
" pale yellow."
The (?c? ad. sent by Mr. Hoffmanns, as well as two others collected by
Prof. Steere near Pani, differ from a large series of true /'. ;•. rubrocKpilla
(more than twenty specimens) by the mnch lighter, more orange-red colonr of the
forehead and crown, which, in the typical form, are of the same dark crimson-red
as the occipnt. The difference between the two forms is about the same as that
existing between P. m. mentnlis Sel. and P. m. if//ii/?ra Bangs.
In my pajjcr on the genus Pipra t I alluded to the pale forehead, in two
specimens from Pani ; having now received three more skins which exhibit the
same peculiarity, I feel quite confident as to the constancy of this character.
This form will shortly be described by a fellow-student of ornithology.
t Cat. Birth jflrit. ^f1ls. xiv. p. 171 : Jt. jidlioerjiJitilx.i, specimen g. | Iltit I'-'OC. p. l.'l.
( 363 )
37. Pipra leucocilla leucocilla Limi].
Pijim leiicocilln Linnaeus, ^^lls. Ail. Frid. ii. Prodr. p. 3^5 [1761. — No locality, we substitute
Sicnnam]; Sclater & Salviu, /'. Z. S. 1867. p. 580 (Pant) ; LayarJ, Jhis 1873. p. .■!84 (Parii).
3 cJ ad., 2 c? jnv., and 3 S ad.: 1:3, 31. x., 1, 3, 7, 3U. xi., 1, 3. xii. 05.
Nus. 7, 45, 49, 02, 78, 237, 241, 250.
38. Pipra opalizans Pelz.
Cf. .Vol-. Zonl. xii. (19U5) p. 2'Ji.
A siugle ? from Prata, 27. x. 05. No. 23. " Iris while, feet grey, bill
greyish white." — Wing 65; tail 21); bill 11 nam.
According to a letter, Mr. lioli'manus observed also the male, bnt unfortunately
lie was not able to get it.
39. Chiroxiphia pareola pareola (Linn.)
Pipra Pareola Linnaeus, S>/st. Nat. xii. 1. p. 339 [17(56. — ex Brisson, Edwards and Maicgrave:
we accept diycnnc (ex Brisson) as the typical locality].
aiiro.r!phia pareola Pclzehi, Zur Oni. Bras. ii. (18G8) p. 129 (Pard) ; Layard, Ibis 1873. p. 384
(Pard) ; Sclater & Salvin P. Z. S. 1867. p. 580 (Pard).
9 S ad., 3 (? jnv., and 3 ? ? from Prata : 7, 8, 10, 13, 16, 20, 20, 30. xi ;
6, 19, 20, 24. xii. Nos. 86, 89, 102, 113, 114, 140, 142, 153, 154, 175, 231, 283,
SS2, 388, 417. Iris marked as " brown " or " dark red."
The specimens agree perfectly with others from Cayenne and British Gniana.
As far as we know, there are three geographical races of C. pareola to
be distiugnished.
a. C. p. pareola (Linn.) : Guianas, Lower Ama>;ons and Eastern Brazil as far
soutli as Esjiiritu Santo (Wied).
//. ( '. p. atlantka Ualm. Island of Tobago.
c. C. p. boluiana Alien. Eastern Bolivia.
40. Chiromachaeris manacus purus (Bangs).
Maiiacus manacus purus O. Bangs, Proc. Ncio Enrjl. Znol. CI. i. (1899) p. 30 (Santarem).
Chiromachaeris manacus (nee Linne) Sclater & Salvin, P. Z. S. 1867. p. 580 (Para) ; Layard,
Ibis 1873. p. 384 (Pani).
6 t? ad. and 5 ? ¥: 11, 21, 29.xi. ; 3, 13, 18, 20, 23. xii. Nos. 118, 162,
229, 233, 234, 235, 253, 332, 368, 387, 415.—" Iris brown or dark brown."
In addition to these, we possess two males and a female, collected by
Prof. iSteere near JIarca da Legua, and at Margnary, in the district of Para.
The adnlt males fully bear out the characters indicated by Mr. Bangs. All
have the under tail-coverts white, only the sides of the abdomen being ])ale grey ;
the black area on the back is restricted to a rather narrow transverse bund.
Four d d ad. collected by Natterer near Borba (ou the lower course of
the Ivio Madeira) belong also to this well-marked form.
4 1 . Scotothorus amazonum wallacii (Scl. & Sal v.)
[I/elcro/K'lma amaziinuni Sclatur, P.Z.S. I.s6t.). p. -166 (Chamicuros, East Peru).]
IIc(eropclma laillacii Sclater & Salvin, I'.Z.S. 1867. p. 579 ("in vie. urbis Pard").
5 cJc^ ad.: 28. x. ; 12, 15, 29. xi. 19U5. Nos. 35, 107, 124, 129, 227.
" Iris brown." Wing 87, 91, 92, 92, 94 ; tail 07, 70, 70, 70i, 72; bill 15—10 mm.
( 364 )
Tliesc specimens arc particularly iiiturcslinjr, as they como from tlie ty|iical
locality.
S. a. ivallacii is a very slightly differentiated form, but may be distinguisbed
from 6'. a. amazoiuim by its purer green back and by the duller, less rufescent outer
webs of tlie (juills. The crown is scarcely washed with rnt'ous, and there is no
rufous tinge on the throat. As a rule, breast and abdomen are of a paler, rather
more greyish green colour than in S. a. amazoimm, but some specimens are hardly
different on this score.
A series from I'ritish Guiana (Caniacusa, Bartica Grove, R. ('arimai)g), and
two adult specimens from the ('aura Kiver, Veue/.., agree with those from I'an'i
in the colour of the head and wings, but the back is decidedly darker, more
brownish green, almost as dark as in S. a. amazonum from Upper Amazonia, and
the throat and foreneck are tinged with brownish. These birds are thus somewhat
intermediate between S. a. amazonnm and .S'. a. wallacii.
42. Tityra cayana (Linn.)
Cf. Not. Znol. xii. (l'J05) p. 294.
3 cJJ'ad. and 1 ? from i'rata : 7, 12, 24. xi. n.). Nos. l(i,s, loii, 2iiu, 2'.Jl.
" Iris brown."
These skins are typical in every way, and agree exactly with others from
Cayenne and Surinam. Uf. my remarks /. c.
43. Lathria cinerea (Vieill.)
Cf. Nuc. Zwil. .xii. (1005) p. -2115.
1 cJ ad. and 1 immature ? : 3. .\i., 13, xii. lOtJo. Nos. 04, oSti. '• Iris brown."
44. Lipangus simplex (Licht.)
Cf. I. c. p. 295.
One pair from Prata : 2i5. x., 4. xi. no. Nos. 38, 75. " Iris ' dark red ' in tlie
male, ' brown ' in the female."
45. *Laiiiocera hypopyrra (Vieill.)
Ampdishijpi>i>iina Vieillot, Xoui\ iJkt. viii. (1817) p. 1U4 ("ii la Guy;me" sc. Cajx-unt).
1 S from Plata, 1. xi. Uo. No. 52. " Iris brown."
This specimen has the bright patch on the sides of the chest sul|iliur-yellow,
and no orange spots in the middle of the breast. It agrees with others from British
(iuiana and the Orinoco region.
The species is new to the fauna of Parfl.
4(j. *Sclerurus rufigularis Pelz.
Sckriiri'S riijhjiilavls Pulzulu, Zar Ontitli. BrasU. ii. (18G8) p. Kil. dcscr. oriy.(15orIja ami Maiabitaiias,
Brazil sept.).
A single adult (? from Prata, 24. X. 05. No. 11. " Iris brown."
It agrees perfectly with Pekeln's types: 1 ? ad. and 2 c?c? imm. Besides
these I have examined an immature bird from tiie Takutu River, British (iuiana,
and a young ? from Ipousin, Rio Ajijirouague, Gayennc (Cherric coll.) in the
Tring Museum ; and the British Museum possesses five specimens, uoue of them
( 3G5 )
quite adnlt, oLtaiued by H. Wliitely, jr., in Briti.sli Guiana at tiie following localities:
Rio Cavimang, R. Takutu, Onnimee aud Bartica Grove. They bad been referred by
Dr. iSelater partly to S. mcxicanus, partly to S. caudacutus.
After comparing these eleven skins with some thirty specimens of <S'. mexicanus,
I have come to the conclusion that S. nijij/i.daris is a perfectly distinct, though
nearly allied, species. I at first thought it might be a geographical race of
S. mcxicanus ; but snch cannot be the case, as true .S'. mexicanus also is found in
some of the localities where jS'. nifigidaris is met with. For instance, the British
Museum has an adult S from Ournmee, British Guiana, and a specimen from the
Capim Kiver (coll. Wallace) which 1 am unable to distinguish from several ( Central
American skins. On the other hand, among the large number of skius from Mexico,
Nicaragua, Costa Rica, etc., I never found any specimen approaching the form
here called S. nijiijidaris.
All the eleven specimens dilfer from S. maxicanus in their much weaker
and shorter bills and in having the throat of a decidedly brighter and clearer
rufous. In <S'. m. mcxicanus and S. m. obscurior Hart, the young birds have the bill
(juite as long and stout as the adults.
For the sake of comparison I append the measurements of the three forms :
29 <S. m. mexicanus from Central America, bill 22 — 25J mm.
5 S. m. obscurior from N.W. Ecuador, bill 22 — 241 mm.
11 -S'. rujigularis, 17 — 19^ mm.
47. Synallaxis omissa Hart.
SynallKxU iimUsa Hartert, liiiU. Brit, Oni. CI. xi. (Juue I'JOl) p. 71 (Panl).
Sijnalla.ris ridilans (ucc Teinm.) Sclator li Salvin, P. Z. S. 18(37. p. bli (Para).
1 6 juv. from Prata, 14. si. 1005. No. 121. " Iris brown." Wing 63; tail OS;
bill 13i mm.
This specimen is a young bird, as shown by tlie llntfy structure of the feathers
on the rump, and diifers from the type in having the head and mantle of a more
brownish hue, the sides of the neck strongly, the chest and the sides of the body
slightly washed with ferruginous, and the quills outwardly margined with the same
colour. The blackish throat-])atch is rather smaller, and the under wing-coverts
are narrowly tipped with orange. The tail is uniformly dull blackish, as in the
type specimen. The specimen collected by ^V^allace and recorded s. n. S. rutilans
by Sclater & Salvin, is an immature bird of the present species.
<S'. omissa is jjeculiar to the district of Para.
48. Automolus infuscatus paraensis Hart.
Cf. iVrji). Zool. xii. (1905) p. 279.
1 ? ad. and 1 adnlt bird not sexed, from Prata, 23. x., 17. xi. 05. Nos. 2,
145. Iris marked as " greyish white "' (No. 2) and " brown " (No. 145).
These examples fully confirm the differences stated in my former article.
49. Philydor erythrocercus (Pelz.)
Cf. A'.-f. ZwA. xii. (1905) p. 280.
A single i ad. from Prata, 28. x. o5. No. 37. " Iris Ijrown."
( 306 )
on. Xenops genibarbis genibarbis 111.
Xniiips ;iniibarlji.i Itt'geT, Prudr. Si/sl. Maiiim. cl Av. p. 'll'-'i (1811 — CameUi) ; Lajuril, ISTo. p. 38u
(Panl).
A single j aJ. from Prata, 15. xi. 05. No. 127. " Iris brown."
51. Glyphorhynchus cuneatus cuneatus (Lcht.)
Cf. Nov. Zool. xii. (I90r.) p. l'Sh.
3 SS and 2 ? ?, 24, 20. x., In, 10, 23. xi. U5. Nos. 1, 18, 105, 131), l!)0-
" Iris brown."
52. Dendrornis eytoni (Scl.)
Cf. /..-. p. 283.
3 c?<? ad. from Prata, In. xi., 5, 12. xii. 05. Nos. Iii3, 273, 325. "Iris
brown."
The bill is entirely black. Wing 123, 123, 127 : tail ItiO, KiS, 112; bill
42—43 mm.
53. Dendrocolaptes certhia certhia (Bodd.),
Cf. I.e. p. 282.
A single S ad. from Prata, 23. x. 05. No. 3. " Iris brown."
This specimen agrees with others from (Aiycnne, etc., and shows no apjiroach
to 1). c. ridgwajji from Santarem.
54. Thamnophilus major semifasciatus (Cab.)
T. m. s. Hellmayr, Nov. Zvnl. xii. (UI05) p. 283.
A single <? ad. from Prata, 21. xii. 05. No. 397. " Iris red."
55. Thamnophilus amazonicus Scl.
Tliamiinjjliilus amazoniciis Scla.tei, p. Z. a. 1858. p. 214 (dcscr. J 'i ) pi. l.l'J (Upper Ama/ioiis) ;
Sclater & Salvin, P.Z.S. 1867. p. 575 (Para and Rio Capim) ; Liyaiil, Ibis 1873. p. 38G
(Pari).
Nine males, some of them not i|nite adnlt, 3, 4, !), lOi, 24, 25, 28. xi., 12, 13. xii.
1905. Nos. 6b, 70, 98, 143, 194, 205, 221, 322, 333. " Iris brown."
1 ? ad., 7. xi. 05. No. 90. " Iris brown."
These specimens do not differ from Qpper Amazonian skins.
Salvin and other authors called this bird T. rujlcollis ex Spix. The fVniales of
7'. amaso/ticus and 3'. ci/iereicrps Pelz., however, are qnite identical in coloration,
dilfering only in size, and as Spix collected also on the Rio Negro where the latter
species is met with, his description and figure could refer with equal certainty to
the female of T. cinereiceps. Unfortunately the type does not any longer exist
in the Munich Bluseum, and I sec uo alternative but to drop the name rujlcollis
altogether in favour of those proiwsed by Sclater aud Pelzelu, about the application
of which there is not the slightest doubt.
50. Thamnophilus palliatus (Lcht.)
Lanius jmllialiis LichtensteiD, Fos. Diibl. 1823. p. 46 (Bahia).
A single ?, 0. xi. 05. No. 77. " Iris whitish grey."
( 367 )
57. Thamnophilus incertus Pelz.
Thanuiiiphilus incerliia Pelzeln, Zur Orn. Brasil. ii. (ISfiS) p. 149 (Par,i,).
Dysithamnus incertus Hellmayr, Nov. Zool. xii. (1905) p. 284 (crit.)
Four adult males, 7, 22, 25. xi., 9. xii. 05; and 1 ? ad., collected 14. xi. 05.
Nos. 83, 120, 176, 207, 306. Iris marked as "browu," "reddish brown,"
" yellowish red," or " bright red."
These specimens are in every way identical with those sent by Mons. Robert
from the same district. The males have the bend of the wing and the lesser upper
wing-coverts freckled with white. The top of the head is always slate-grey like the
back, and there is no trace of a white interscapular blotch. About the systematic
position of this species cf supi-a, p. 339.
58. Dysithamnus mentalis mentalis (Temm.)
ilijnthcra menlalh Temminck, PI. cul. livr. 30. talx 179. fig. 3 = ^ ad. (1823— Bresil, sc. Curytiba,
ParanA. Type in Vienna Museum examined).
Dysithamnus mentalis Goeldi, Ibis 1903. p. 499 (Capim River).
1 (? ad., 14. X. 05, and 1 ¥ ad. 24. xi. 05. Nos. 15, 197. " Iris brown."
They agree well with a good series of South Brazilian skins, except that in the
male the yellow colour on the abdomen is less extended.
59. Pygiptila stellaris (Spix).
Thamniiphilus stellaris Spix, .\i\ Bras. ii. (1825) p. 27. tab. xxxvi. fig. 2. desor. orig. (J (Pard).
Tlianinaphilus manilipennis Sclater, Ediiih. New Philos. Journ. (new ser ) i. (185.5) p. 247 (" Quixos
in Cisandean Ecuador and Peruvian Amazons ").
Thamiviphilus stellaris and Pijqnptila niamlipennis Sclater, Cat. Birds Brit. Mas. xv. pp. 105, 217.
I S vix ad. and 1 ? ad., 24. xi., 21. xii. 05. Nos. 193, 399. " Iris greyish
brown" {S), "brown" (?).
They agree in every respect with sj)ecimens from the Upper Amazons and the
Orinoco district. The Triug Museum has lately received a pair from near Paramaribo,
Surinam, which does not diifer either.
There can be scarcely any doubt that T. stellaris is the oldest name for this
species, although Spix's type does not any longer exist in the Munich Museum.
The descrijjtion and figure suit our bird very well, and the only reason for rejecting
the name stellaris was apparently the locality, which had been considered erroneous,
but which now proves to be quite correct.
1 may mention that in the Cat. of Birds this species is described under two
different names, as I have satisfied myself by examining the specimens in the
British Museum. The two skins recorded under the head of Thamnophilus stellaris
are absolutely indistinguishable from those named Pygoptila maculipennis. It need
not be emphasised that they have nothing to do with Thamnophilus tristis Scl. &
Salv., which belongs to quite a different genus.
60. Thamnomanes caesius hoflfmannsi Hellm.
r. c. hnffmannsi Hellmayr, Bull. B. 0. C. xvi. Febr. 1906. p. 53.
T. caesius (nee Temminck), Hellmayr, Nov. Zuul. xii. (1905) p. 285 (Igarapc-AssiS, Pari).
2 c?(? ad. from Prata : 15, 20. xi. 0.5. Nos. 148, 158. " Iris brown."
This interesting new form resembles T. c. caesius from Bahia in having no
white interscapular blotch, but agrees with T. c. qlaucus in the clear slate-grey
24
( 368 )
colour of the plnmage. It differs from l)olh by its pure white (not slaty greyish)
under wiug-coverts. The throat is variegated with white (tliis lieing sometimes
slightly indicated iu T. c. ylaucug), the ear-coverts show fine whitish shaft-lines,
and the edge of the wing is white, not slate grey as in T. c. glaucus.
The S imm., sent liy M. Robert, and recorded s. n. T. caesium, exhibits the
same differences when compared with examples of T. c. qlaucus iu corn'spoudiiic
plnmage.
The female of the Para form is not distiugnishable from the same sex of
T. c. caesius.
The range of the three subspecies of Thnmnomnnes is as follows :
a. T. c. caesius (Temm.) : Eastern Brazil : Bahia and Espiritu Santo (Wied),
west to the Rio Madeira (Salto do Girao, Natterer coll.), and the Rio Jnrud (Garbe
coll. ; spec, in Bins. Panlista examined).
b. T. c. hoffmanyisi Hellm. : N.E. Brazil, vicinity of Para.
c. T. c. glane/is Cab. : Cayenne, Brit. Guiana, Surinam, thence thrnngh the
Orinoco region (Mnnduapo, Caura R.) to the Rio Negro (Marabitanas, Rio Iganna,
Barcellos, Manaos), to Eastern Ecuador, E. Pern, and N.E. Bolivia (D'Orbigny
coll. — spec, in Paris Museum examined). Occasionally also found in Bogota
collections.
61. *Myrmotherula surinamensis surinamensis (Gm.)
Sitta surinamensis Gmelin, fit/st. Xat. 1. i. (1788) p. 44'2 (ex Latham : " Surinam Nuthatch," Geii.
Syn. 1. ii. p. 654. tab. 28 (= ? )— Surinam).
A single S ad., collected at Prata, 16. xi. 05. No. 138. " Iris brown." —
Wing 52 ; tail 25 ; bill 14 mm.
It agrees perfectly with four <S d ad. from near Paramaribo (topotyjiical
M. surinamensia). All have the lower mandible whitish, and tlie whole lower
surface striped with black.
This is not only the first record of the species for Para, but also for tlie
Brazilian Avifauna.
62. Myrmotherula cinereiventris cinereiventris Sd. k Salv.
Cf. Nor. Zool. xii. (1905) p. 286.
1 S ad., 1 S juv., 1 ? from Prata, 26. s., 15, 17. xi. 05. Nos. 20, 126, 146.
" Iris brown."
The female agrees in the coloration of the underparts with others from
Cayenne, while the one sent by Robert is much darker underneath.
63. Myrmotherula axillaris axillaris (Vieill.)
Myrmothera axillaris Vieillot, Noiir. Dirt. xii. (1817) p. 113 (" La Guyane," sc. Cayenne).
Thamnophihis melatioga^ter Spix, Av. Brnx. ii. (1825) p. 31. pi. xliii. fig. 1. ( = (J) (Cnrupit, month of
the Amazons. — Type in Munich Museum examined).
3 cJc? : 14. X., 16. xi., 21. xii. 05. Nos. 10, 134, 401. Iris brown or black
(No. 10).
1 ? : 13. X. 05. No. 6. Iris brown.
These specimens are in no way different from a good series of topotypical
Cayenne skins, having the sides of the l)elly snow-white. I examined the tyi)e of
T, melanogaiiter Spix, preserved in the Munich Bluseum, and fonnd it also to belong
to typical J/, axillaris. Tlie form of Eastern Brazil (from Pernambuco to Rio),
( 3fi9 )
hitherto called J/, melanogastra, onght to bear the next available name, M. luctuosa
Pelz., of which I have likewise seen the types, and its proper appellation is,
therefore, M. axiUrnis luctuosa Pelz.
64. Myrmotherula longipennis Pelz.
Myrinotherula longij/etmis Pelzeln, Ziir Oiu. Brasil. ii. (1868) p. 153 (Marabitanas and Rio Negro).
M. brevicauda (neo Swainson !) Solater & Salviu, P. Z. S. 1867. p. 576 (Capim River, the (J only).
2 (?c? ad., 30 X., 15. xi. 0.5. Nos. 40, 130. "Iris brown."— Wing .59*, 60;
tail 33 ; bill 13i, I4| mm.
These two skins have the middle of the breast and the cheeks decidedly paler
grey than the rest of the plnmage, and the ear-coverts show distinct silvery-white
shaft-lines, which are entirely wanting in the types of the species and in one c? ad.
from Snapure, Canra River.
I have examined the specimens recorded by Sclater & kSalvin s. n. M. brevi-
cauda. The S belongs to M. longipennis, agreeing perfectly with Guianan examples
in size and coloration except the lower surface, which is mnch clearer grey, almost
as pale as in M. beidepschi Hellm. ; bnt I feel snre the colour has been altered
by some external influence.
The so-called ? , however, belongs to M. a. axillaris (Vieill.).
6.5. Myrmotherula hauxwelli hellmayri Snethlage.
M. It. hellmayri Snethlage, <Jni. ilonbrr. xv. (1906) p. 9. descr. orig. {J (no locality given, the types
came apparently from Panl).
Mi/rmntherula spec. ; Hellmayr, Nov. Zool. xii. (1905) p. 285 (Pard).
3 (?c? and 3 ? ?, collected 15, 24, 28. x., 1, 26. xi. 05. Nos. 14, 16, 33, 36,
50, 213. Iris marked as " brown" or " blackish brown."
These specimens differ from a good series of true M. hauxwelli by the complete
absence of the white interscapular patch; besides this, the females have the
upper jiarts of a warmer olive-brown. The male has been fully described by
Miss Snethlage.
M. hauxwelli does not belong to the section of M. axillaris, where it has
been placed by Mr. Sclater {Cat. Birds xv. p. 237), bnt it is a very near ally of
M. guttata (Vieill.). In fact, both species agree perfectly in the markings of the
wing-coverts, tertials, npper tail-coverts, and rectrices, and in the shortness of
the tail. The only difference between the males consists of the colour of the
abdomen, which is ferruginous red in M. guttata, and slate grey like the breast in
M. hauxwelli. The females differ only by the colour of the throat and breast.
These are pale l)uffy browni.sh in M. guttata, while in M. hauxwelli the whole
nndersurface from the chin to the crissum is bright ferruginous.
66. Formicivora grisea (Bodd.)
Tnrdus grieseus (sic) Boddaert, Tahl. Pl.etil. 1783. p. 39 (ex '• Le Grisin de Cayenne," Daubeuton,
PI. oil. 643. fig. 1 = ^.—Cayenne).
Furmirirora gnxea Sclater & Salvia, P.Z.S. 1867. p. 576 (Rio Tocantins) ; Pelzeln, Ziir Ornith,
Brasd. ii. (1868) p. 73 (Pard) ; Lay.ard, Ihia 1873. p. 387 (Pard).
3 c? <? and 2 ? ? ad. : 4, 8, 21, 23. xi., 20. xii. 05. Nos. 60, 80, 170, 182, 386.
" Iris brown."
This series agrees well with specimens from British Guiana and Bahia, the
females being bright ochraceous underneath except the whitish throat.
( 370 )
67. Cercomacra sclateri Hellm.
Cercomacra sclateri Hellmajr, Xov. Zool. xii. (1905) p. 288 (type ex Chyavetas, Peru).
A single S ad. from Prata, 14. xi. 05. No. 119. " Iris brown."
This specimen agrees in all essential characters with the type from Eastern
Pern, but is mnch paler grey both above auil below. The e.xamples collected by
Natterer on the Rio Madeira show the same pale coloration, and seem to indicate
that the Brazilian birds represent a different form. A good series of &esh skins
from Pern, however, should be compared before attempting any separation.
68. Cercomacra tyrannina ('Scl.)
Cf. Nov. Zool. xii. (1905) p. 286.
7 c?c? ad. and imm., 5 ¥ ? from Prata: 14. x., 3, 7, 8, 10, 14, 24, 28. xi.,
2, 15. xii., 05. Nos. 12, 13, 07, 79, So, Kii), 110, 190, 220, 242, 346, 347. " Iris
brown or greyish brown " (No. 196).
69. Pyriglena leuconota leuconota (Spix).
Cf. Xoc. Zool. xii. (1905) p. 290.
6 c? J ad., 1 ? ad., and 2 ? ? juv. : 27. x., 2. xi., 9. xii. Nos. 20, 28, 29, 31, 53,
54, 59, 303. Iris marked as " red."
The females present all the characters described by me /. c.
7<i. Hypocnemis poecilinota vidua Hellm.
Hypocneiuis vidua Hellmayr, Nov. Zool. xii. (I90j) p. li'jO, descr. orig. ? (Igarapt'-Assii) ; Snetblage,
Om. Monhcr. xiv. (1906) p. 29 (descr. (J).
4 c?(? ad. : 28, 30. x., 2, 27. xi. . . 1 „ .,-.,, o ,.,, ,
5 ? ? ad. and r. : 27 sV, 30. x., 16, 25. xi. """^i;,^:^^ '\ '\ '"' '''
1 c? jnv. : 20. xi. 05 . . . . .J ^'"^^^^^^'-- "I-'-wn.'
When describing this interesting addition to the fauna of Pard, I had only a
single female before me. Miss Snethlage has since published a note pointing out
that the male closely resembled that of H. poecilinota. This is quite an unexpected
discovery, as the type showed no trace of the black and fulvous markings on the
back and wings so conspicuous in the female of 11. poecilinota.
Mr. Hoffmanns has now sent a good series of both sexes, which enables me
to speak more confidently about the characters of H. vidua.
As stated by Miss Snethlage, the males resemble in general coloration those
of JI. poecilinota poecilinota. and //. p. lepidonota, having the back, the uj)per
wing-coverts, and inner secondaries marked with black and white cross-bars, but
differ at a glance by their whitish throat, which is in decided contrast witli the
cinereons colour of the remaining underparts. while in the two allied forms the throat
is quite as dark grey as the belly. The markings of the tail are the same ; each
•rectrix with a white apical margin and a roundish white spot on the inner web,
tlie outermost pair with a longitudinal white spot also about the middle of the outer
web. Sometimes there are two white spots on the inner web of the two or tliree
outer tail-feathers, but the same variation is to be observed in sjiecimens of the
allied forms.
The females agree perfectly with the tyjjical example ; some of them have a
( 371 )
little more white at the base of the dorsal feathers, but always much less than in
H. p. gi'iseiventris, of which, thanks to the kindness of my friend Dr. von Lorenz,
two of the types are now before me. In addition to the characters indicated in
the diagnosis of H. vidua, I find that the edge of the wing is cinereons (not pale
ferrnginons as in //. griseiventris), the chin whitish like the throat (not ferrugiuous),
and the upper surface of a much less reddish tint.
Considering the apparent close relation between the females of H. grisekentris
and //. vidua, I at once suspected that the male of the former was most probably
of a similar style of coloration to those of //. poecilinota and allies, and might
have been mistaken by Pelzeln for the male of H. p. lepidonota, of which Natterer
collected a fine series at Marabitanas on the upper Rio Negro. Dr. Lorenz having
kindly sent me the whole series, I found among the skins an adult male collected near
Borba, where Natterer also obtained two of the type specimens of Pitbjs gri.sei-
tentris. This skin is marked by Natterer himself as the male of the latter —
a statement whi('h seems to have entirely escajied tlie notice of Pelzeln — and, as it
agrees with them in all structural details, there can he no doubt that it represents
the hitherto unknown male sex of P. griseiventris !
On comparing it with a large series of H. p. poecilinota and II. p. lepidonota,
the general resemblance is very close indeed, but the Borba bird differs from either
in having tlie upper tail-coverts pale grey, with only a narrow black subtermiual
bar just behind the white tip, while they are deep black tipped with white in
its allies. The base of the three median pairs of rectrices is distinctly washed
with cinereous (in H. p. poecilinota, H. p. lepidonota, and H. p). vidua they are
deep black), and the throat is decidedly paler grey than the breast, but not whitish
as in //. p. vidua.
We have thus four geographical races of one t)'pe : the males not or very
slightly difl'erent, lint, the females with striking differences in coloration. The
most interesting fact, however, is that iu two of these forms — //. p. poecilinota and
H.jj. lepidonota — the females have attained the ornamental characters of the male
plumage, while in the two others — H. p. vidua and H. p. griseiventris — they are
still devoid of these, tlius apjiarently representing a more primitive state of
development.
In tlie following lines I give the characters and the range of the four forms : —
a. Hypocnemis poecilinota poecilinota Cab.
Hypoenrmis poecilinota CabanLs, Arch. f. Naturg. 13. i. (1847) p. 212. tab. 4. fig. 2 (cJ) [British
Guiana].
Hab. British Guiana : Bartica Grove, Camacusa, Merume Mts., Roraima,
Rio Atapurow. Surinam : Maroni River. Cayenne : Rio Approuagne (Cherrie
coll., Mus. Tring). Venezuela : Mnnduapo and Nericagua on the Orinoco R. ;
Canra R. : Nicare, La Pricion, La Union, Snapure.
d. Tliroat quite as dark grey as the rest of the underparts ; ujiper tail-coverts
black with white tips.
Wing 65—71 ; tail 40—4.5 ; bill 17—18 mm.
?. Forehead and sides of head, including chin, bright clear ferruginous, passing
mto rufesoeut brown on tiie pileum ; back olivaceous brown, and like the upper
wnig-coverts and inner secondaries with broad black subterminal and fulvous apical
cross-bands; upper tail-coverts olive brown witli fulvous tijis. Edge of the wing
pale ferruginous. Tail-feathers black, witli a. wiiite apical margin and a crramy
( 372 )
white spot abont the middle of the inner wel), thp outermost pair with a similar,
but more lengthened spot on the outer web. Undersurface clear cinereous.
Wing 64—07 ; tail 39—45; bill 16^- 17J mm.
b. Hypocnemis poecilinota lepidonota Scl. & Salv.
Hypocnemis Upidonota Selater & Salviu, P. Z. S. 1880. p. 160 [Saraja^u, E. Ecuador].
B. poecilonota (not of Cabanis) Pelzeln, Zwr Oni. Bras. ii. (1808), p. 88 (part. : Marabitanas and
Kio Vaupe).
Hab. East Ecuador : Sarayavn, Santiago, Rii) Napo : East Colombia : Bogota
coll. ; iV. 11'. Brazil : Marabitanas and Rio Vaupe, Upper R. Negro. Ferti : Iquitos,
Chamicuros, Huambo, Yurimagnas, Guayabamba, Loreto ; Monterico and La
Merced, Chanchamayo ; Pozuzo, Hnauuco (Hofl'manns coll. — Mus. Tring).
S. Not to be distinguished from that of the preceding form.
Specimens from Colombia, E. Ecuador and Peru : Wing 67 — 70 ; tail 42 — 47 ;
bill 16—17 mm.
Specimens from Marabitanas : Wing 69—71 ; tail 49—52 ; bill 18 mm.
? . Whole lower surface, like forehead and sides of the head, bright ferrn-
ginons; rest of upper parts warm rufescent brown ; back, uj)per wing-coverts and
secondaries marked as in the preceding form, but the apical margins white instead
of fulvous. Upper tail-coverts rufescent brown at the base, separated by a broad
black subterminal band from the white tips. Tail as in B. p. poecilinota, but the
cross-baud pure white.
Wing 65—69 ; tail 45—49 ; bill 16—18 mm.
c. Hypocnemis poecilinota griseiventris (Pelz.)
Pilhys grisdrentris Pelzeln, Zur Oni. Brasil. ii. (1868) p. 167, desor. orig. ? [VUla Maria and
Engeuho do Gama, Mattogrosso ; Borba, R. Madeira].
Hi/jjociiemis poecilonota (nee Cabanis) Pelzeln, /. o. p. 88 (part. : Borba).
Hab. West Brazil: Borba on the Rio Madeira, Villa Maria and Eugenho do
Gama on the R. Gnapore, Mattogrosso.
S ad. (Mus. Viudob. No. 15514, Borba, January 1830, Natterer coll.). Differs
from a and b in having the upper tail-coverts cinereous, tbis colour being
separated from the white tips by a sharply defined black subterminal bar. The
middle tail-feathers are washed with grey at the base, and the throat is decidedly
paler grey than the belly.
Wing 69; tail 46 ; bill 16| mm.
? . Forehead, sides of the head and a small chin-spot pale ferruginous ; rest
of pileum, back, wings, and upper tail-coverts uniform warm rnfescent brown
without any cross-bars ; a distinct white interscapular patch ; tail-feathers rufescent
olive-brown with a black subterminal band and a white apical margin, but with
no other white markings. Edge of the wing pale ferruginous. Throat whitish,
rest of undersurface pale grey, flanks washed with rufescent brown.
Wing 67—70 ; tail 45—49; bill 16—18 mm.
(/. Hypocnemis poecilinota vidua Hellm.
Uypocnemia vidua Hellmayr, Nov. ZwA. xii. (1905) p. 290 desor. orig. ? (Pard).
Hab. N.E. Brazil, vicinity of Para: Igarape-Assii (Robert), Prata
(Hofl'manns).
c?. Like //. p. poecilinota and //. p. lepidonota, with the upper tail-coverts
( 373 )
black tipped with wbite, but at once known by having the throat whitish in marked
contrast to tlie schistaceous-grey belly.
Wing 65-66; tail 40—42 ; bill 10—17 mm.
? . Like //. p. grisekentris without spots on inner web of tail-feathers and
without any markings on back, npper tail-coverts and wings, bnt the upper parts
are less rnfescent, more olive-brown, the edge of the wing and the sides of the head
pale grey (not ferruginous), the chin whitish like the throat and the forehead olive-
brown like the pileum. Moreover, the flanks are washed with olive-brownish, and
the axillaries pale grey (not brownish).]
Wing 62i— 05; tail 39—41 ; bill 15—17 mm.
71. Formicarius ruficeps (Spix).
Fnrmicarius viificejjs amazonicus Hellmayr, A'oi'. ZooL xii. (1905) p. 292.
1 ? ad. from Prata, Para, November 16, 1905. No. 137. " Iris brown."
It has the top of the head much paler rufous than the c? sent by Robert, and
agrees perfectly with specimens of true F. ruficeps of Eastern Brazil, so that I can
no longer distinguish my F. r. amazonicus. The throat is black as in the male,
only narrow shaft-lines being white, which, however, are not visible unless the
feathers are raised.
72. Conopophaga roberti Hellm.
Nov. Zool. xii. (1902) p. 292 (Igarape Assu, Pan'i) ; Snethlage, Orn. Moiiber. xiv. (1906) p. 9 descr. J .
9 <?c?: 20, 31. X. ; 2, 13, 15, 21. xi. ; 2. xii. 05. Nos. 19, 47, 65, 56, 57, 111,
125, 189, 247. " Iris brown, greyisli brown or reddish brown."
5 ? ? : 13. X. ; 9, 13, 20. xi. ; 11. xii. 05. Nos. 5, 91,-110, 156, 318. "Iris
brown."
The adult males agree exactly with the type. The young males, with the tail-
feathers slightly pointed, show more or less distinct rufcscent apical margins to the
greater wing-coverts, secondaries and rectrices.
The female has been described by Miss Snethlage. It ditfers from the male
in the following details. Forehead and pileum are of a dull russet brown, lores and
superciliary region jiale cinereous (not black), the white postocular stripe is rather
narrower and bordered below by a tine blackish line. Cheeks, malar region and
foreneck pale cinereous, throat white, ear-coverts rufescent brown ; flanks more
distinctly washed with brownish than in the male.
In all other respects the sexes are alike.
<?(?: Wing 09—72 ; tail 35-38 ; bill 15—10 mm.
? ? : Wing 05—69 ; tail 32—30; bill 15 mm.
The female of C. melanogastra Mfinitr. difi'ers from that of C. roherti by its
much larger size, much stronger and entirely black bill (in C. roberti the lower
mandible being always whitish), deeji chestnut ujiper-jiarts, dull blackish instead of
rufescent brown ear-coverts, and by the presence of a broad, light grey superciliary
stripe.
73. Corythopis torquata anthoides (Puih.)
Cf. JVoii. Zoul xii. (1902) p. 293.
A single $ ad., 14. xi. 05. No. 123. " Iris brown."
It agrees perfectly with the one sent by Robert.
(374 )
74. * Threnetes cervinicauda Gould.
Threneles cervhiicauila Gould, P. Z. S. 1854. p. llJ9 (• Quijos, in Ecuador ").
1 ? fere ad. 30. x. 05. No. 42. " Iris black."— Wiug 57 ; tail 31 ; bill 30 mm.
This is a highly interesting addition to the fauna of Panl, being not only the
first record of the species for Brazil, but e.xteuding its range from Eastern Peru and
Ecuador to the Lower Amazons.
In the Paris Museum there is a sjiccimen, collected near Para by M. P>araci|uin
in 1859, which I found to differ slightly from typical T. cervinicauda ; but the
female now sent by M. Hoffmanns does not exhibit these differences, being in every
detail exactly similar to a good series from Eastern Ecuador and Bogota collections.
It is strange to find T. cervinicauda at Para, where I should rather have exjjected
T. leucurus of Cayenne and Surinam.
75. Glaucis hirsuta (Gm.)
Trochilus hirsutiis Gmelin, Si/st N<il. 1. i. p. 490 (1788.— ex Brisson : ex Marcgrave.— " Brasilia."—
We accept Bahiu as typical locality).
Glaucis lanceolala Gould. Momiqr. Trorhil. i. (1861) pi. 8 (Par.i) descr. orig.
Glaucis hirsuta Layard, Ibis 1873. p. 388 (Pani).
1 adult, not sexed ; 1 S juv. : 14. xii. 05. Nos. 335, 340. " Iris black."—
Wing 62, 63 ; tail 38, 42 ; bill 30, 31 mm.
Not different from a series of Bahia skins. The characters upon which
Gould founded his G. lanceolata, viz. the pointed tail-feathers and the bufify
edges to the quills and upper wing-coverts, are those of the young birds. One
of Mr. Hoffmanns' specimens (No. 335) agrees with Gould's plate and description,
while the other one is exactly similar to adults from various localities.
76. * Phoethornis affinist moorei Lawr.
[Phaefm-tiis affiiiis Pelzeln, Silziuiysler. Ahul. V.'iui xx. (1800) p. 157 (Marabitanas, Barra do Rio
Negro).]
Phaethoniis mnorei Lawrence, Ami. New York Lye. vi. (1858) p. 'JuS ("Ecuador").
1 (J jr. and 1 ? ad., 28. xii., 23. x. 05. Nos. 371, 4. " Iris black, feet grey,
bill black, lower mandible red." Wing 60, 58 ; tail 68, 64; bill 38, 37 mm.
These specimens nearly agree with a series of Bogottl skins and others from
the Peruvian Amazons. The underparts are dirty greyish, only the middle of
the lower belly being faintly tinged with buffy.
In true P. a. a(finis the whole lower surface is bright butf, slightly underlaid
with greyish on the sides of the throat. Pelzelu's types from Marabitanas and
Barra do Rio Negro which lie before me are practically identical with a very large
series (some thirty specimens) from Cayenne, British Guiana and the Orinoco
River. P. guianensis Bouc. is, therefore, a pure synonym of P. q//inis.
In the Gould collection, British Museum, there are several examples of P. a.
affinis said to be from Para, but these are skins of the unmistakable Cayenne make,
and certainly never came from Pani I
It is interesting that tlie Upjier Amazonian form occurs near Pani. We should
rather have expected to meet with ]'. a. a//inis of Cayenne here.
t This is the proper specific name, I believe, for the small species witb white under tail-coverts,
called 1'. mpereiVosua by Berlepscb & Hartert. Trochilug sujicrritiosus Linnaeus appears to be referable
to P. malari.i N'ordm., since Brisson, upon whose description Linn6"s account is exclusively based, states
that the under tail-coverts like tlic rest of tlie lower parts arc " d'un hlanc roussatre." The description of
the four lateral tail-feathers also applies much better to P. vmlaris.
( 375 )
77. Phoethornis ruber ruber (Linn.)
Troehiliis ruber Linnaeus, Syat. Nat, x. p. 121 (1768. — ex Edwards, Oruilh. i. p. 32, pi. 32, fig. sup.
— Surinam).
Phaethornis rujigaster and P. pygmaeus auct.
Pygmoniis pygmaeus Layard, Ibis 1873. p. 388 (ParA).
Six specimens, three marked as (J, two as ?, and one not sexed, 2, 15, 8, 9, 22,
28. xi. 0.5. Nos. 58, 77, 87, 95, 174, 224. " Iris black."
Tiiese specimens are all immatnre, with the tail-feathers tipped or margined
with cinnamon-rnfons, only one showing the purple-black cross-band on the breast
characteristic of the adnlt plumage. In most of them the rectrices are uniformly
bronzy brown, as in sj)ecimens from Rio and Bahia ; but two examples (Nos. 58 and
174) have a slight purplish tinge just behind the rufescent tips, thereby approaching
P. ruber episcopus Gould of British Guiana and the Orinoco district. The latter
form is very closely allied, and single specimens are not always distinguishable.
Berlepsch and Hartert {Noi\ Zool. ix. (1902) p. 82) replaced the name of
P. episcopus by that of ruber Linn., which is strictly referable to the bird of
Surinam. Four specimens from near Paramaribo, however, agree with the Brazilian
form (= pygmaeus Spix), as do also skins from Cayenne, of which I examined a
good series in the Museums of Tring, London and Paris. It is, therefore,
evident that the P. pygmaeus auct. ought to be called P. r. ruber, while the form
of British Guiana has to stand as P. r. episcopus Gould.
78. Campylopterus obscurus obscurus Gould.
Campylrqjtfrua obscurus Gould, P. Z. .S'. 1K4K. p. 13 (" River Amazon ") ; Sclater & .Salvin, P. Z. S.
1867. p. 584 (Pard) ; Layard, Ibis 1873. p. 388 (Pari).
2 cJc? jr. and 5 ? ? : 4, 15, 21, 22, 24. xi. 05. Nos. 166, 168, 178, 181, 199,
262, 351. " Iris black."
This series agrees with the type of C. obscurus (Brit. Mus.) and with the
specimens collected near Para by Wallace and Layard. The British Museum
possesses also an adult c? (with the shafts of the outer primaries much dilated)
obtained by Natterer, which, although having no locality on the label, doubtless
came from Para, as it formed part of Natterer's last consignment. Altogether, I
examined eighteen skins of this form, which is characterised by having the two
outermost tail-feathers on each side tipped with smoky-grey.
In a large series from the Upper Amazons,* including the type of C. aeqiia-
torialis Gould (ex Quito), these patches are pure white or scarcely shaded with
greyish. The difference, though slight, being evidently constant, two distinct races
ought to be recognised :
a. C. obscurus obscurus Gould, from Pant.
b. C. obscurus aequatorialis Gould, ranging from Eastern Ecuador through
Eastern Peru to N.E. Bolivia.
79. Plorisug^a mellivora (Linn.)
Trijchi/us uiellivnrus Linnaeus, Syst. Nat. x. (1768) p. 121 [ex Edwards, Oruilh. i. pi. 3,5, fig. sup. —
Surinam].
Fh,risu;ia mellivora Sclater & Salvin P. Z. S. 18G7, p. 584 (Pari) ; Layard, Ibis 1873. p. 388 (Pari).
A large series of both sexes and young birds, obtained in October, November
and December.
* 4 (J (J ad., 255 I'ebas; 2 jj (J ad. East Eonador ; 1 J or J jr. from Raniya<;u, E. Ecuador;
4 skins from N.E. Peru (B.arHett coll.) ; 1 ^J ad., 1 J Mapiri, N.E. Bolivia (Buckley); 4 5 $ or ^ J jr.
from ,S. Augustin and Guanay, N.E. Bolivia (M. Stuart coll.— Mus. Tring).
( 376 )
80. Agyrtria nitidifrons (Oonlil).
Tlutumalias nilidi/rons Gould, P. Z. S. 1860, p. 308 (loc. ign.).
Agi/itria iiiikUfrons Berlepsch & Leverkuhn, Ornis vi. (1890) p. 28 (Par;!).
13 (?c? ad. aud imm., aud 3 ? ? : 26, 27, 28. xi., 2, 3, 4, 5, 8, 11, 14, 15, 18, 22.
xii. Nos. 209, 212, 222, 223, 244, 250, 263, 270, 274, 297, 313, 338, 353, 380, 407,
410. " Iris black, bill black, lower mandible at basal half red or yellow."
Of this species only two specimens were known to exist: Gould's t3'pe in the
British Museum (from unknown origin), aud a specimen in the Kiel Museum said
to be from Pari, a locality which proves to be quite correct. Mr. Hoffmanns is to
be congratulated upon having secured a good series of this rare bird.
As pointed out by Berlepsch and Leverkuhn, A. nitidifrons is a near ally of
.1. aflinis (Gould), agreeing with it in the greyish green tail, in the coloration of
the under tail-coverts, aud in the throat being spotted in the same manner. The
adult males differ, however, at a glance in having the forehead and anterior part of
the crown of a beautiful glittering bluish green instead of dull bronze green; the
glittering rounded spots on the throat and sides of tlie head have also a decided
bluish hue, while they are pure golden green in A. affinis. Another striking
difference consists of the markings of the tail, the four outer pairs of rectrices
showing a broad, well-defined steel-black subterminal band on both webs ; in
A. affinis this is but slightly indicated on the inner web of the two outermost
tail-feathers.
The females of A. nitidifrons lack the glittering patch on the head, the
forehead and crown being shining metallic green like the back, and the sjwts
on the throat are less bright and almost devoid of the bluish tinge. They are,
however, easily distinguishable from .4. affniis by the markings of the tail.
6S. Wing 47— 50| ; tail 28—31 ; bill 15-ll6i mm.
? ?. Wing 45—48 ; tail 25—27 ; bill 15—16 mm.
The type differs from our series only by the white ground colour of the throat
being almost entirely hidden by the glittering bluish green apical portion of the
feathers, and in the strong golden bronze suffusion on the rump and upper tail-
coverts.
81. *Hylocharis cyanus viridiventris Berl.
[Trochilus cyanus Vieillot, Nou>\ Dk-t. xxiii. (1818) p. 426 (" Br(5ail," coll. Delalande— sc. Rio).]
llyloi'haria cyanea, subsp. ririiliventris Berlepsch, Ibis 1880. p. ll.S ["Venezuela (Merida), Trinidad,
and the Orinoco district "].
•i Si ad., 24, 25, 27. xi. 05. Nos. 195, 202, 210. " Iris black."
They agree perfectly with specimens from Cumand, and British Guiana, being
much darker green both above and below than typical II. c. a/anus of S.E. Brazil.
New to the fauna of Para.
82. Hylocharis sapphirina (Gm.)
Trochilus sapphirhms Gmelin, Syst. Aal. 1. i. (1788) p. 4'JtJ (ex Buft'nu ; Hab. ign.).
Hylocharis sapphirina Sclater & Salvin, P. Z. S. 1867. p. 584 (Pari).
2 SS ad., 1 S juv., 1 ? : 5, 7, 8, 20. xii. 05. Nos. 207, 284, 292, 390. "Iris
black."
These specimens agree exactly with a scries from Surinam and Brit. Guiana
{11. yuiunensis Bone). If two geographical races of this species are recognised,
the name sapphirina must be retained for the northern form, Bnfifon's description
being most probably based upon Cayenne e.xamples.
( 377 )
83. Chlorestes caeruleus (Vieill.)
Trorhilus raeruleus Vieillot, Noiw. Diet. vii. (1817), p. 361 (type ex Cayenne).
Eiwephala meridea. Sclater & Salvin, P. Z. S. 1807, p. 584 (Pard) ; Layard, /i/s 1873. p. 388 (Pard).
5 (J c? ad., 3 ? ¥ : 18, 21, 22, 23, 2G. xii. 05. Nos. 376, .390, 4U2, 4U8, 409, 413,
414, 418. " Iris black."
They do not differ in any way from Cayenne sjiecimens.
84. Thalurania furcata furcatoides GouJd.
Cf. Nov. Zool. .xii. (1905) p. 297.
14 (?c? ad. and juv., 13 ? ? : 14, 26, 30. x. ; 19—22, 25, 28, 29. xi. ; 3, 5, 10,
11, 12, 14, 16, 18, 20, 21. xii. 06. " Iris black."
The characters of this form are extremely variable, and I am a little doubtful
whether it can be maintained as distinct. As a rule, the tail is less deeply forked
than in T. f. furcata of Cayenne, the green of the throat less extended, and the
blue band across the ujiper back slightly interrupted in the middle.
85. *Avocettula recurvirostris (Sw.)
Troi-lnlnsremrviroslvh Swainson, Zoulng. lllustr. ii. (1821-L'), pi. 105, descr. orig. (^ ad. ("Peru."—
errore I )
A single <S imm. 22. xii. 0.5. " Iris black." No. 404. Wing 55 ; tail 26J ;
bill 17 mm.
The three outermost tail-feathers are fiery coppery red on the under-snrface,
as in the S ad., but widely tipped with white. On each side of the breast there is
a tnft of silky snow-white feathers, apparently a peculiarity of the adult male, but
never mentioned before as far as I know.
This is the first record for Brazil, the species having been only known to occnr
in Cayenne and British Gniana.
86. Anthracothorax nigricoUis (Vieill.)
Ti-ochihis Mi/ncolliK Vieillot, Nmiv. Did. vii. (1817) p. 349 (" Br(5sil ").
Lavipornis mango (nee Linnaeus !), Sclater & Salvin, P. Z. S. 18G7. p. 584 (Mexiana).
L. violkauda (nee Boddaert) Layard, lUs 1873. p. 388 (Pari).
2Si ad., 1 c? jr. and 3 ¥ ? : 7, 14, 15, 17, 21. xii. 05. " Iris black." Nos. 285,
334, 349, 350, 366, 400.
87. Anthracothorax gramineus (Gm.)
Trochihis gramineus Gmelin, Si/sl. Nat. 1. i. (1788) p. 488 [based on the " Haussecol vert " of Buffon.
—As typical locality accepted Cayenne \ cf. Nm. Zonl. xiii. p. 35].
Lamponiis gramineus Sclater & Salvin, P. Z. S. 1867. p. 584 (Mexiana).
A single ¥ juv. collected near Prata, 20. xii. 05. No. 389. " Iris black."
Wing 72|; tail 39^; bill 30 mm.
It agrees well with specimens from Cayenne and Surinam. I examined also
the adult <J obtained by Wallace, and found it to be identical with our series.
88. Chrysolampis mosquitus (Linn.)
leus, Sijat, Nut. X. (1758
ility].
1 S in moult, 21. xii. 1905. No. 392.
Trochilus moKrjnitus Linnaeus, -S^;, .V«/. x. (1758) p. 120 [ex "Indiis.-— errore! We substitute
Surinam as typical locality].
( 378 )
89. Psilomycter theresiae theresiae (I'a Silva).
Trochilii.i thrrcsiae Da SUva, Mui. Min. Bras. 1843. p. 2 +
Polytmus Theresiae Cabanis & Heine, Mus. Hem. iii. (18GU) p h (Parii).
1 S .jr. : 18. -xii. 05. No. 377. " Iris black."
This specimen agrees perfectly with a good series from Surinam and British
Guiana, having the under tail-coverts spotted with shining green.
4 (JcJ collected by Natterer at ManAos belong also to this form, while those
obtained by him near Marabitanas on the Upper Rio Negro (3 (?(?,2 ? ?) represent
P. t. leucorrlums Scl. & Salv., which diftVr.s only in having the nnder tail-coverts
pnre white.
90. *Topaza pella (Linn.)
Trnchilua Pella Linnaeus, Syst. Nat. x. (1708) p. 119 (ex Edwards, Ornith. i. pi. 32 fig. inf.-
Stirinam).
A single ? moulting: 16. xii. 05. No. 359. " Iris black."
Though collected by Natterer near Manios, this species has not yet been
recorded from Para.
91. Heliotlirix auriculatus phainolaema (Jould.
Cf. Hellinnyr, Nor. Znol. xii. (I'.iOo) p. 297 (Igarap^-Assii).
1 S ad., 2. xi.; I ? ad., 20. xi.; ?. 3 S juv., 4, 13, 14. xi. 05. Nos. 72, 122,
155, 152, 337. " Iris black."
The (? ad. agrees perfectly with the one sent by Robert, having the whole
throat glittering golden-green without any trace of the white median stripe,
characteristic of typical //. a. auriculatus of S.E. Brazil.
The ? ad. and the young males have the underparts pure white, not spotted,
while in the corresponding plumages of the typical form there are always distinct
bronze-brownish spots_on the foreneck and chest. In the nearly allied H. auritus
(Gm.) there occur occasionally females with pure white underparts, though, as a
rule, they are spotted like those of//, auriculatus auriculatus.
Besides" the colour of the throat in the males, there is a marked difference in
size between //. a. auriculatus and H. a. phainolaema, the latter having decidedly
shorter wings.
//. a. auriculatus : U S S ad., wing 68— 71.— 10 ? ? , wing 67—70 mm.
H. a. phainolaema: 'i S S ad., wing 01, 63, 65.-4 ? ? and c?c? juv., wing
62—65 mm.
It may be added that I have examined the type of //. aequatoriali.-i Boncard, +
said to have been collected by Buckley in Ecuador, and found it to lie a sMn of the
well-known Rio-make, differing in no way from typical H. a. auriculatus. The
dillerences alluded to by Boncard are merely individual. In Natterer's series there
are from the same localities specimens with the chin and sides of the head pure
glittering green and others which show there a more golden hue (Ypanema,
Goiaz, etc.)
t I have not l)con able to verily this quotation.
J Gen. Humming liiriU p. 311 (1893.— Type now in Paris Mnscum).
( 379 )
92. * Calliphlox amethystina (Gm.)
Tnicliihis niiirthtjstiiiiis Gmelin, Syst. Nat. 1, i. p. 496 [1788.— ex Daubenton, PL oil. 672 fig. 1
aud Buiion. — Cayenne].
Three yonng (Jc? and 4 ? ? : 16, 21, 22, 23. xii. 05. Nos. 356, 393, 394, 403,
406,411,412. "Iris black."
New to the fauna of Para, but very widely distributed in South and East Brazil.
93. Lophornis gouldii (Less.)
Ornismija gouldii Lesson Hist. nat. Truchilid. 1832. p. 103 pi. 36 descr. orig. ^ ad. (hab. ign.)
Trochilus retjiiiae Schreibers, Isis 1833. p. 534 descr. orig. (J ad. ("in provinoia Mattogrosso ").
4 iS ad. and one young i : 10, 19, 22, 23. .xii. 05. Nos. 328, 360, 361, 379,
405.—" Iris black, bill red, tip blackish."
The exact range of this lovely species has been very imperfectly known till
now. The specimens in the British Museum and in the Boucard Collections (now
in the Paris Museum) were said to be from the Lower Amazons, but as an adult
male had been collected by Natterer far in the interior of Mattogrosso, the
correctness of this indication appeared to me always very doubtful, and 1 was
really pleased to find a series in Mr. Hoffmanns' collection.
It seems, therefore, that L. gouldii has a more extensive range than hitherto
supposed. Thanks to the kindness of Dr. Lorenz, the type of T. reginae Schreib.
(ex Mattogrosso) is now before me, aud it proves to be absolutely identical with the
Para specimens. The discrepancies noted by Pelzelu t are certainly only due to
inaccuracies of the figure in Lesson's work, the elongated feathers on the sides
of the head being evidently much exaggerated and the coloration of the outer
tail-feathers obviously wrong. Lesson never saw the bird himself, but based his
description upon a coloured drawing from a stuffed specimen in the Loddiges
collection, forwarded to him by Mr. Stokes.
The specimens before me present the following measurements :
4 c?t? ad. from Para : Wing 39—40 ; tail 23— 23i ; bill lOJ— lU mm.
5 ad. from Mattogrosso (type of T. reginae Schreib.) : Wing 40 ; tail 24 ;
bill 1 1 mm.
94. * Discosura longicauda (Um.)
TrucUiliis lunijicavdus Gmelin, Syst. Nat. 1. i. (17^8) p. 498 [ex Buffon : "du cabinet de Mons.
Mauduit," no locality. We substitute Cayenne].
A single ? not quite adult; 18. xii. 05. No. 375. " Iris black."
New to the fauna of Pani, but found in Cayenne and in Bahia collections.
95. Chloronerpes flavigula (Bodd.)
Of. NoiK Zool. xii. (1905) p. 301.
A single S imm. from Prata : 3. xi. 05. No. 68. " Iris brown."
96. Trogon melanurus Sw.
TrcKjiiii melamirus Swainson, Aiiim. in Mniag. p. 329 (1838. — Demerara) ; Solater & Salvin,
P. Z. S. 1867. p. 583 (Pard.) ; Pelzeln, Zur Ornith. Brazil, i. (1867) p. 19 (Pard).
1 c? ad., Prata: 14. xii. No. 342. "Iris pale brown, bill yellow."
Not different from Demerara specimens.
t Zur Ornith. BrasU. i. (lSli7) p. 32.
( 380 )
97. *Trogon atricoUis atricoUis Vicill.
Trogon atricoUis Vieillot, Noiiv. Did. viii. (1817) p. 318 (ex Levaillant, Couroucous pi. 8.—
" Guyane, Surinam et ^ la Trinitij ").
A single ? from Prata, 4. xi. (15. No. 73. " Iris brown."
The South Braziliau form, T. a. cimjsochlorus Pelz., differs only by its larger
size aud less golden upperparts.
98. Neomorphus g'eoffroyi (Temm.)
Cf. Nov. Znol. xii. (1905) p. 298.
A single adult from Prata: 30. x. 05. No. 44. " Iris yellow, feet grey-blue,
bill gre}ish blue, base of upper mandible black." Wing 160; tail 273; bill 42i mm.
Not different in any way from the ? ad. sent by Mons. Robert.
99. Ramphastos ariel Vig.
Cf. /. c. p. 300.
A single S ad. from Prata : 13. .xii. 05. No. 329. "Iris greyish white, bill
black, basal band yellowish green."
100. Pteroglossus aracari aracari (Linn.)
Cf. /. c. p. 300.
1 S ad. : 8. xii. 05. No. 300. « Iris brown."
As shown {I. c), P. aracari of Linue refers to the Brazilian I'orm, commonly
called P. wieclii.
li'l. Pteroglossus bitorquatus Vig.
Pteroglossus bitorqiiativi Vigors, Zool. Journ. ii. (1826) p. 481 (loc. igu.); Sclater & Salvin,
P. Z. S. 1867. p. 586 (Parji) ; Pelzeln, Zur Oniith. Brmil. iu. (1869) p. 237 (Pari).
1 (? ad. from Prata: 16. xii. 05. No. 352. "Iris reddish yellow; upper
mandible greenish yellow, lower one white at basal half, black at apical half."
Agrees well with an adult c? from (Jamolins, near Para, collected by
Prof Steere.
102. Galbula cyanicollis Cass.
Cf. Nor. Zonl. xii. (1905) p. 296.
4 (?c? ad., 2 ? ? from Prata : 1. xi., 1, 11, 18. xii. 05. Nos. 51, 238, 239, 315,
316, 373. " Iris brown, bill yellow, tip of upper mandible black."
The Tring Museum possesses now a fine series of this rare species, whicli is only
known to occur on the Lower Amazon and its southern affluents, the Hio Madeira
and Rio Jurna.
There is considerable variation in the colour of the upper mandible to be
observed. In one male it is entirely yellow, while all the other specimens have the
apical half more or less blackish.
103. Bucco tectus (Bodd.)
Cf. I. c. p. 296.
1 d, 1 ? ad. from Prata : 5. xii. 05. Nos. 205, 200. " Iris dark brown."
( 381 )
104. •Bucco striolatus Pelz.
Bucco striolatus Pelzeln, Sttzxwgsher. Akad. Wieii (Math. Physik. CI.) xx. (18.56) p. 500
(Engenho do Gama & No Dourado, Mattogrosso).
1 t? ad., 2 ?? ad. from Praia: 23. xi. 16, 18. xii. 05. Nos. 187, 363,
3T4. " Iris pale yellow, feet and bill bine-grey ."^^^Wing 80 — 88 ; tail 76 — 83 ;
bill 32—33 mm.
These specimens have the sides of the head, the forenock and the band across
the na])e pale buff while the same parts are deep ochraceons in a female from
No Dourado (typical J>. striolatus). Very likely the Para birds are separable as a
geographical form, but I would like to see more specimens of the typical race, since
of two examples from Saraya^n, East Ecuador, one is quite as bri^;htly coloured as
Natterer's ? from Mattogrosso, while the other is almost as pale as those from Para.
Anyhow, B. striolatus is a very interesting addition to the fauua of the district,
extending the range of the species from Mattogrosso and Eastern Ecuador to the
Lower Amazons.
105. Malacoptila rufa (Spis).
Bucco rufus Spix, Ac. Bras. i. (1824) p. 52, pi. xl. fig. 1. (''ia sylvis fl. Amazonuni").
Monasa rufa Pelzeln, Zur Omith. Brasil. i. (1867) p. 23 (Pard).
Malacoptila rufa Sclater & Salvin, P. Z. S. 1867. p. 583 (Pard).
1 ? in moult, 5. xii. 05. No. 76. " Iris brown, bill greyish black."
Differs from Upper Amazonian examples only in having the frontal band of a
paler ferruginous, and the back more olive-brown (not so rafescent). The white
shaft-lines on the head, too, are somewhat broader.
106. Monasa morphoeus morphoeus (Hahn.)
Cf. Nov. Zool. xii. (1906) p. 297.
1 ¥ ad. from Prata, 28. xi. 05. No. 217. " Iris brown, bill red."
This bird is intermediate between M. m. morphoeus and M. m. peruana Scl.,
having the narrow white frontal band of the latter, but the large chin-spot of the
former. Other specimens from Para, however, are entirely typical of J/, m. morphoeus.
107. Chelidoptera tenebrosa tenebrosa (Pall.)
Cuculus tenebrosus Pallas, Neue Nord. Beijtr. iii. (1782) p. 2 (Surinam).
Chelidoptera tembrosa Sclater & Salvin, P. Z. S. 1867. p. 583 (Parji) ; Layard, Ibis 1873. p. 392 (Pan-l).
A single i ad. from Prata, 30. xi. 05. No. 232. " Iris brown, bill black.''
Wing 108 ; tail 53 ; bill 1!J mm.
This specimen agrees with a series from Snrinam as regards size and
coloration. Skins from S.E. Brazil (C. t. brasiliensis Scl.) average rather larger,
and have the abdomen mncli lighter, more ochraceons.
108. Pyrrhura perlata (Spix).
Cf. Nov. Zool. xii. (1905) p. 301.
2 c?(?, 1 ?, and one adult without indication of the sex: 7, 9, 13. xii. 05.
Nos. 290, 302, 307, 331. " Iris brown."
One of these specimens has a few brownish red feathers on the frontal edge,
( 382 )
and the anterior i)art of the bhiish cheeks slightly mixed with yellowish green.
In tiic three others the cheeks, as well as a distinct, thongh narrow band across
the forehead, are dull greenish hlne.
This species is pectiliar to the district of Pard.
Iii9. Pionites leucogaster (Knlil).
Cf. X<n\ Zml. xii. (1905) p. 302.
A single adult, sex not stated, from Prata, 10. xii. Ii5. No. 309. "Iris golden
yellow."
In every way similar to the examples sent by Robert.
This species is also confined to the district of Pard.
III*. *Accipiter bicolor bicolor (Vieill.)
Sparviw! bicolor Tieillot, Nonr. Dirt. x. (1817) p. 325, descr. orig. juv. (Cayeune).
A single immature ? from Para, 24. xi. 05. No. 201. " Iris light yellow."
In addition to it, the Tring Museum possesses an adult <S, collected by Prof.
Steere at Bemfica, near Pard, July 25. The iris is marked as " dark yellow."
It agrees in the whiteness of the under])arts with a series from British Guiana,
Bogota coll., and Caicara, Orinoco River.
Specimens from Panama, Chiriqui, and Western Ecuador are much darker
underneath, and have been described by me as Accipifer bicolor sckistochlamys.]
This is the first record of the species for the district of Para, though it was
collected near Santarem X by Riker. The ? ad. obtained by Natterer near Barra
do Rio Negro, and determined by von Pelzeln as A. pilcatus, § belongs likewise
to this species, as I have satisfied myself by examining the specimen.
111. Asturina nitida (Lath.)
Falro nitidus Latbam, Ind. Ornilli. i. (1790) p. 41 (Cayana).
Asturina lutida Pelzeln, Zur Ornith. Brasil. i. (1867) p. 3 (Paid,).
A single ? changing from the young into the adult phimage, 29. xi. 05.
No. 230. "Iris brown."
112. • Leucopternis albicollis (Lath.)
Faleo ttlbiioUis Latham, Ind. Oni. i. (1790) p. 3(i (Cayaii.i).
1 ? ad. from Prata, 18. xii. 05. No. 378. " Iris brown."
Diifers from all other specimens before me by having no black s])ots on the
mantle and interscapular region.
113. Leucopternis kuhli Bonap.
LcucopteriiU Kuhli, Bonaparte, Cuimp. Ar. (1»4'.() p. 19 (no locility).
L. hiupi Bonaparte, Iter. Ma;/. Zoo!. 1850 (Sept.) p. 481.
L. siij)frrili<iris Sclater <t Salvin, P.Z.S. lKI'i7. p. 589 (Paril) ; Pelzeln, Znr Oniith. Briinil. i. (I8C.7)
p. 3 (Borba, Paril).
1 S ad. from Prata, 8. xii. 05. No. 299. " Iris yellowish red."
t BM. B.O.C. xvi. (1906) p. 82. % Aul.-, ISUl. p. IGl. § Orn. JlrMil. i. (ISlw) p 8.
( 883 )
J c? iul. and 1 ? ad. from Igarape-Assi'i, 21. i., 5. iii. 04. Noe. 1025, 2015.
Iris : S, " ronge f'once " ; ? , " bruu ronge." Collected by Mous. A. Robert.
This well-marked species is still verj' rare in collections, and hitherto only
known from Para and Borba, Rio Madeira. It differs from L. melanops in having
the toj) of the head and the mantle slaty black, only the bases of the feathers being
white, and in having no visible white spots on the lower back and upper wing-
coverts. The well-defined white superciliary stripe is another point of distinction.
The juvenile plumage of this species seems to be unknown.
Its jirojier name is the one used above, L. ktikli having undoubted priority over
L. Itaupi. Bonaparte's diagnosis, though very bad, cannot be referred to any other
sjjecies.
Wing 20.), 210, 210 ; tail 130, 13s, 138 mm.
IH. Columba speciosa Gm.
Culiiinha sjicriiixu Gmelin, Sysl. Xnt. 1. ii. (1788) p. 783 (ex Buffoii, ct D'Aubonton, I'l. nit. 213 —
Cayenne) ; Sclater & Salviu, /'. Z. S. IKG7, p. .5'.I0 (Pani).
Lepklociias speciuxa Pelzeln, Ziii- Orii. linisil. iii. (1800) p. "274 (Para).
1 S ad. in beautifnl plumage, 23. xi. iio. No. Iy3. Iris "brownish red"';
bill " cardinal red."
Identical with sjK'cimcus from British Guiana.
11 -J. Columba plumbea togoteusis Berl. k Leverk. (an subsp. dist. ?)
Clihirueiiaa plnmbca Vieill., subsp. uov. hoijotauis Berlcpsch and Levcrkiibn, (Jrnis vi. (1890), p. 32
(Bogota coll.)
C'ltumba rimicm (nee Temminck !) Sclater & Salvin, P.Z.S. 18("i7. p. 580 (Capim River).
C. phimhca (nee Vieillot!) Goeldi, Ibis 1903, p. 409 (Rio Gapim).
1 c? ad. from Prata, 11. xii. 05. No. 317. " Iris white." Wing 175 ; tail 135 ;
bill l(i^ mm.
It diifers from an adnlt bird from Pina, Nortln'rii Peru, which we may consider
to represent G. p. bogotensis, by having the throat almost of the same lilac tint as
the Ibreueck, while in the Peruvian skin there is a distinct vinaceous bufi' throat-
patch. This is also very well pronounced in a good series from Merida and N.AV.
Ecuador ; they differ, however, a little in coloration from the Peruvian and Para
examples.
"While it is quite jiossible that the birds from northern South America may
be divisible into several geographical races, there can be iitt the slightest doubt
that they are altogether different from (J. p. plumhca Vieill., whicli is confined in
its range to Eastern Brazil, from Bahia to Rio Grande do Sul. I have now examined
2(j specimens of the tyjiical, and about a dozen of the northern form, and found
them always easily sej)arable. In C. p. bogotensis the head and nape, as well
as the nnderparts, are always strongly washed with vinaceous or lilac, while in
C p. plumbea they are pure light grey, with an almost imperceptible lilac hue.
Berlepsch and Hartert* have already shown that C. piirpureotincta Ridgw.,
erroneously united with C. plumbea by Salvador!, had nothing to do with this
species, but lieloiiged to the grouji of ('. iiigriroilris, which is characterised by its
short, thickish bill, rufescent blown (not greyish) under wing-coverts, etc., etc.
• ^V«r. iSir./. ix. (1002) p. 117.
2i)
( 384 )
P.S. — Since the ubove was written, I exainiued tlie series in the British
Mnseum. The adnlt bird colleited b)- Wallace on the Cajiim River a<rrees perfectly
with our d from Para. A topotyiiical skin of C. />. bo(jote)isis (from Boj];ot;i) dilfers
from both in its cinereous occiput, only the forehead, sincijjut and naj)e being
vinaceons. A larger series is reipiired to confirm the constancy of tliis character
or otherwise.
110. "Geotrygon violacea violacea (Knip & Temm.)
Culiimha fhilaccii, Knip & Temminck. rii/cuiix, Fam. (jihiinhia (IS08-11) p. 137, pi. :i9 (luu. ign. —
Mus. Paris).
1 ? fere ad. from Prata, Para, 23. xi. 0.5. No. 184. " Iris light brown, bill
greyish red." Wing 145; tail 8<J (moulting); bill 15 mm.
It agrees with several adults from Victoria, >S. Paulo, but has the foreneck
rather more greyish. All Brazilian specimens examined by me have the vertex
l)ure light grey, shading into whitish on the forehead and the cheeks ; the malar
region and ear-coverts are very pale greyish.
Para is a very interesting locality, and extends the range of the species far to
the north. I have examined specimens from Rio de Janeiro (Mus, H. v. Berle])sch),
Victoria (Tring) and Ypanema, S. Paulo (Natterer ; Vienna Mus.), Rio Jordao,
jirovince Araguay, Minas Geraes (Robert ; Mus. Tring), and Bahia (Wucherer ;
Mus. Brit.).
Schlegcl t records a specimen from Surinam, which, however, cannot be
Temminck's type {ciJe supra).
Young birds liave the forehead dnll whitish, the vertex and occiput dull bronzy
greeu, the back bronze-brown, with only slight purple reflections on the mantle,
the foreneck and chest dirty brown, aiul the (piills with distinct cinnamon ajiical
margins.
The Central American form, (J. c. alljirciitcr, % though closely allied, can easily
be distinguished by having the forehead (as far as the anterior edge of the eye),
cheeks, malar region, and ear-coverts clear bulf-pink (Ridgw., Sonien<:l. iv. ^nj,
there being no trace of the bluish grey colour on the crown. C)f this form I have
compared a good series from Miravolles, Costa iiica (Underwuod coll.).
117. Coliimbigallina passerina griseola (Spix).
[Ciiliiiiihii jHissciimi Linnaeus, ■S'yx/. »Y"f. x. (176H) p. IG.'i [part. ; as typical locality accepted : Jamaica
(ex Sloauc — first quotation.)]
('nlnnihiwi (jriseola Spix, -li'. Bra/;, ii. (IH25) p. 58, pi. l.^.-cv'., fig. i ('■ in sylvia fl. Aiiav.nimm ").
Cliamae/iiiia /lasKirina (nee Linnc) Sclater & Salvin, P. Z. S. 1 8li7. p. 591 (Para) ; Layard, IbU 1873.
p. 305 (Parti).
1 c? fere ad. from I'rata, IT. xi. U5. No. 14V. " Iris rosy red."
Besides this, we received an adult cJ from Prof. Steere, collected near Bemfica,
Para, August 30. Both specimens have the bill entirely blackish.
As I have shown in my paper on Spix's types, ( '. i/riseola was based ujion a
female of the present species. Consequently this luxme must be accepted for the
continental form of C. j/a^sserimi.
t .l/w.<. l'inj»-li(iK iv. Cidnmbac, p. liiij.
{ Ccolrijiji'n alliivciitir J,awrtnce, iVw. Acoit. I'hilad. l!*G,j. p. lOS (l.lon Hill, I'anama).
( 385 )
lis. Creciscus viridis viridis (P. L. ,S. Mull.)
Of. Nne. ZwjI xii. (1905) p. .%4.
A siDgle ? ad. from Prata, 3. xii. H.j. " Iris browu, feet light red." No. 252.
119. *Crypturus cinereus (Gm.)
Tiiivo cinereus Gmelin, Sijst. Nat. 1. ii. (1788) p. 7G8 [o.'i; Buffoii : Oayeane].
A single ? imm. from Prata, 1.5. xii. Oo. No. 34.S. " Iris browu."
Besides, we received au adult S from Prof. Steere, collected ou the islaud
of Marajo, which differs only iu having the top of the head more washed witli
rufous.
The specimens from Para agree i)erfectly with a series from Britisli Guiaua,
and cannot, consequently, beloug to the pale " variety" mentioned by Schlegel and
von Pelzeln.
New to the list of Para birds.
120. Crypturus soui (Herm.) subsp.
Tinaiiim sou/ Hermaun, Ttill. .iff. Aniiii. 1783. pp. 104,235 [ex Daubenton, PI. c«?. 829— Ciiyenne.
Cryjdttrus 2)ilcati(.i Layard, Ibis 1873. p. 391! (Nazaru, Pard).
. 1 c? imm. from Prata, 21. xi. O.j. No. 163. "Iris greyish brown."
We have also one c? ad. from Igarape-Assu, 10. ii. 04, collected by Mous. A.
Robert, and inadvertently omitted from the report ou his collection. Both specimens
are much paler above and below than a series from British Guiana and Trinidad,
agreeing best with others from Pernambuco. Most probably they belong to a
different form, but whether the birds from N.E. Brazil or those found in Guiana
represent the typical C. sou/', can only be decided by a good series of Cayenne
skins.
( 38ti )
MISCELLANEA OllNITIIOLOGICA.
CRITICAL, NOMENCLATOUIAI,, AN]J OTHKR NUTHS, MOSTIA' oN
PALAEARCTIC BIRDS AM) THEIR ALLIES.
By ERNST IIARTKRT. I'n.U.
PART III.*
The genus ANTHOSCOPUS.
ICANXOT do otherwise than unite the European " rcnJuline Tits" with the
African ones in one genus. The latter have a much longer first primary, but
in A. macronyx it is also longer than in the various allie.s of A. pendulinus, and
from the great variability in the length of the first primary iu many other groups I
must decidedly refuse to separate the African Penduline Tits generically from the
Palaearctic ones. Tlierefore the name Anthoscopus Cab. 1851 (type A. cajjensis)
must be u.sed instea I of Reraiza Stejn. 1880 (type A. jjendidinus). On the other
hand, the American species (Auripariis fiavifrons) deserves to be separated
generically. Its tail is longer, its bill more curved, its first primary long (as in the
African forms), and it lays blue eggs with coppery-brown spots, while the Palaearctic
and African species lay white, unspotted eggs. Also the Himalayan " Aerjithaliis
Jldmiiticejjs" has no place in the genus Anthoscopus^ and must be called Cephulopyi-us
Jlaniniiceps. The more open nostrils, stiff, narrow feathers of the forehead, com-
paratively longer wings and longer tail, and very tiny first jiriraary, justify its generic
separation. Moreover, it does not build a purse-shaped or globular hanging nest,
but bleeds in holes of trees, and lays not white, but blue eggs. Mr. Dresser
(Manual Pal. B. p. 183) still uses the name Aegithulus Boie 1822, although it is
well known that this name was already given by Hermann to the Ix)ng-tailed Tits
in 1804. As Mr. Dresser generally accepts those names which have priority, there
seems to be no reason for a deviation from his usual custom in this case. Mr.
Dresser's treatment of the various forms of Penduline Tits in the manual is also not
very convincing. Why macronyx has been treated as a subspecies of jjendidinua
is not clear, as it occurs together with forms of pendidimts, and is more distinct
than many species acknowledged by ]Mr. Dresser; and why the jierfectly distinct
forms aColicskae and consobrinus have been placed as synonyms under pendulinns
is not explained. The only possibility seems to me to be that the author did not
compare them with pcndxdiniis, or very superficially.
The Asiatic Penduline Tits are not easy to unravel. In I lie Far East, on the
Vang-tse-Kiang, the great ornithologist Swiuhoe discovered a inw i'nrni, which he
named A. consobrinus. This is a very rare bird iu collections, but a number of
specimens were obtained by i\Ir. Ringer near Nagasaki, on Kiu-shiu Island, Jai)an.
It is a very distinct form, easily distinguished by the grey crown and black line
through the eyes, instead of extended black auricular region. Why several authors
have doubted its distinctness is an enigma. The most astonishing remark is that of
Seebohm, who (J5. Japanese Enip. p. 182) says that he has seen two males and two
females, and " that it is, however, very [irobable that it may eventually be proved to
• For I'ait I. suu Xor. Zvh liiOI. i>ii. lOG UO; for Tart II. yof. Zoitl. i:iOJ. p)). IDT-oOa.
( 387 )
be the female of the wei^tern species, or be degraded to subspecific rank." How a
male can be proved to be the female of another species is not known to me. and why
could not Seebolim compare the well-known females of " the western species," by
which he means the European pendtdimis?
The Central Asiatic Anthoscopi were first separated by Sewertzow and Hume.
Sewertzow established two forms, coronntus and tiuicronyx. hut unfortunatel}' com-
plicated matters by creating other names as well, atricupiUus for the former,
cucullatus and pectoralis, like a " Brehm redivivus," and with the same want of clear
discrimination which was Brehm's weakness. Sewertzow also made a "noinen
nudum " jaxartica, which was revived and established by diagnosis by Dr. 8uschkin
in 1904, under theiViimeot'' Remisapendidina jax.rrtensis" {Bull. B.O.C. xiv.p.45).
Hume described the East^Turkestan bird as stoiiczkae (Stray Feuth. ii. p. 521, 1874).
With this form were united the West Tnrkestan birds (Syr Darja), until Suschkin
separated them clearly as jaxartensis (t.c). Unfortunately, at the same time the
last-named author describe<l two more Penduline Tits ! One he named centralasiae,
which is the same as stoiiczkae, and the other yenisseinsis. It is possible that the
birds from the Upper Yenissei, between Tannu-ola and the Sayan mountains, may
ultimately be found to belong to a separate form, but the name yenisseinsis is
merely based on young specimens, which are hardly separable from the corresponding
plumage of Anthoscopus pendulinics jaxartensis — only our jaxartensia have a narrow
chestnut line behind the white foreliead, while this is not seen in the types of
yenisseinsis, which were kindly sent for my inspection by Dr. Suschkin. However,
one jaxartensis from Iskander Kul in the Petersburg Museum* has no trace of this
che.'^tnut line. Therefore it is very doubtful what yeiiis.'<einsis really is, until we
know the adult birds. Dr. Suschkin's diagnosis is of very little use, because he
conijiares his new form with the European pendiilinus.
At present I recognise the following forms :
A. pendidinus penduKnus : South Europe.
A. pendnlinus caspius : Caspian basin to Orenburg.
A. pendidinus stoiiczkae : E. Turke.stan to Dsungaria (figured and enumerated
by Dr. Sharpe in Sec. Yarkand Mission under the wrong name covoniUus).
A. pjendul inus jaxartensis : W. Turkestan (Svt Darja).
A. pendulinus consohnnus : Valley of the Yang-tse-kiang and Kiu-shiu, Japan.
{A. yenisseinsis, Upper Yenissei, doubtful.)
A. coro'itulns : Transcaspia, West Turkestan to Persian Baluchistan.
A. macronyx : West Turkestan to East Persia (Seistan).
The g-enus PANURUS.
The position of the Bearded Titmouse has often been (|^('^tioned, and several
ornithologists have denied its close affinity with the Tits. The nostrils are not
covered with narrow bristly feathers, which are directed forwards, 1ml covered with a
thin operculum; the oesophagus is widened in its second tliiid; the nest differs
from that of the I'arinae. These peculiarities, and the striking resemblance in
colour with the Chinese Furadoxornis lieudei, prove that the correct position of
Fiinufus is among the Faradoxoi'nithinae, which agree with the Faridne in so
many points that they are best placed as a subfamily in the family Fandae.
' Dr. Bianchi most kimllj- sent tliis and many uthcr specimens for comparison, and Dr. Susohliin lent
me his yt-ntssfiniiH.
( 388 )
The genus COLOEUS Kaup.
By many authors united with Comts. The following characters, however, may
be sutlicient excuse for its separation. Bill somewhat short, high, hardly as long as
the head, upper mandible rather straight. Feathers on crown and neck decomposed.
Tail straight, tips of rectrices not rounded, but square. Nests in holes, eggs lighter,
with fewer and larger spots than in C'orvus. Smell ])eculiar, musl<y, different from
that of the true Corvi.
Only Coloeus-monedula, with its subspecies 0. in. dauuncus, and C. ne/jledus —
the latter probably not a species, but a dark aberration — belong to this genus. The
American Corvus ossifrofjus has none of the characteristics of Coloeus, Also the
eggs of C. of<sifragus differ in no way from the type of the genus Con'iis, and might
be taken for small rooks' eggs. They do not at all resemble jackdaws' e^gs.
The genera of PALAEARCTIC MUSCICAPIDAE.
Kowhere, perhaps, do we find more unuecessarj' genera than among the
Muscieapidae. If it would only he understood that zoologists have invented the
so-called genera in order that we may find our way through the vast multitude of
species, and that we may, by this eminently practical method, group together the
most closely allied forms, thus expressing their affinities* in the names by which
we know them, and that in nature only species and subspecies have evolved, but no
such things as genera !
The generally' adopted genera of Miiscicapuhie are alleged to be based on
certain " structural " differences — i.e. length of vibrissae at gape, width and length
of lull ; but if wo examine tliese supposed characters we find it impossible to follow
the customary arrangement, and we shall soon perceive that really the genera were
separated by colour and an unlucky attemjit made afterwards to find structural
differences. Thus I cannot separate the genera Muacicapa, Hemichelidon, Ficedula
(or rather Hedymela), Sijjhin, Zanthopyi/ia (generally altered into Xanihopjygia,
but originally deliberately spelt Zanthojiyfjia), Arizelmnyia, and others. On the
other band, I .should have separated '^Hemichelidon," on account of its short first
jiriinary, if it was not for the fact that sibirica has as short a first jirimary as
i/riseisticta, while the southern representative of sibinca, i.e. Muscicapa sibirica
fulifjinosa, has a longer first primary, this being as long as, or sometimes even a
little longer than, the primary coverts.
On TCHITREA PRINCEPS ILLEX.
]Mr. Outram Bangs {Bull. Mas- Comp. Zool., Harvard Coll. xxxvi. 8, p. 264),
described as a " new species " a " Terpsiphone illex." He had only two specimens, a
male and a female, both from Ishigaki. He declares that they have a different wing-
* One o£ the objections made to mojein soieatifio trinomial nomenclature is, that its adherents
strive to express afllnities in tlicir names, and that this is against our iirincijiU-s of nomenclature, which
should only serve practical purposes, and should not be based on rescarclics of afliiiitics and development.
I do not know who first invented the dopnia that names should not express affinities, but it seems to mo
to be only the personal idea of some of our ornitholoyital friends, and by no means a generally adopted
view, though repeated parrot-like by souk; writers. Moreover, it is at) erroneous and objectionable view,
because, first of all, our r.ld uni\ersally arloptcd nomenclature already expresses most emphatically the
aflinities of species. Everybody will admit that in a genus — unless it is a wrontrly constructed one — the
nearest allies are comprised, in opposition to other groups which are more distantly related. Not only is
there no reason why we should not express affinities in trinomi.als in a similar way as wc do in binomial
names, but the more wc do so the more valuable will our nomenclature be.
( 389 )
formula, differently shaped reetrices, narrower crest -feathers, and smaller size. I have
examined six adult males and as manj' females from Ishigaki, and nearly twenty more
specimens from other islands in the Loo-Choo grouji, i.e. from Amami, Okinawa,
Iriomote, and Iheya, and I find no difference between them and a series of skins
from Hondo, i.e. "typical" princejys. The different wing-formula does not exist.
Mr. Bangs' statement must have been based on a moulting or abnormal specimen.
Nor are the reetrices differently shaped. It is true that the wing of Loo-Choo
specimens is generally about 2 — 5 mm. shorter, and therefore Tchitrea princeps ill/'X,
as this form must be called, may be regarded as a slightly differentiated sub.species.
Single sjjecimens, however, cannot always be distinguished. As a rule, Hondo males
have the wing about 89 — 93 mm., while males from the Loo-Choo Islands have wings
of 84 — 89 mm. A male shot on Okinawa, however, on April 24th, has the wing
91 mm., and some from Southern China have wings of 88 and 89, so that one can
hardly say to which race they belong.
It is difficult to believe that birds with such enormous soft-feathered tails as
Tchitrea can be migrants, and yet we find Tchitrea incei to wander to the Malayan
peninsula, and the same is the case with T. princeps pi'inceps : at least specimens
with long wings (93 and even 94 mm. !) occur there, but aiiparently only in winter.
We must, therefore, assume that they are migrants. It is difficult tounder.-tand why
Mr. Outram ]5angs named such closely allied forms as Tchitrea princeps ille.r with
two names, as he often employs trinomials. Such inconsistency is most disturbing
to students. Or does he still adhere to the exploded theory that one must see
" intergradation " to employ trinomials? Let him read ]Mr. Hart ^lerriam's lucid
article, who remarked most truly, that it depends on our material alone whether we
see intergradation or not. Therefore our scientific language cannot be based on sucli
accidental circumstances; in fact, we must u.se trinomials in all cases where allied
forms, which agree in tlieir essential characters, replace each other geographically. It
is true that more knowledge is required for using trinomials correctly than for
merely separating so-called species binomially, but it is desirable that closer studies
be made than hitherto, and if a mistake is made it will be corrected, as has been
done before.
On the African forms of the genus PYCNONOTUS.
As in so many other genera, ornithologists have hitherto for the most part
confined themselves to distinguishing as many different forms as possible as so-called
species — or occasionally to "lum]iing" them again — liut few attempts have been
made towards an understanding of their actual affinities in connection with their
distribution. Trying to study all these forms with the help of their distribution — as
far as we know it at present — I come to the following conclusions :
1. Undoubtedly all the I'l/cnonoti witli white or whitish under tail-coverts are
geographical reiiresentatives of one species. This has already been recognised by
Oscar Neumann. Thus barbalus, inornatus, gabonensis. arsinoe, somaliensis, and
schoanus are undoubtedly forms of one species.
2. These forms are connected with yellow-vented forms through f/nhonensis,
which has the under tail-coverts white, edged, and sometimes all over tinged with
yellow. P. h. gabonensis is so much like young tricolor that even Professor
Keichenow mistook the latter hr r/ahnnensis. He states that r/ahonensis extends to
the Congo, on the strength of a bird collected at Manyanga by Eohndorff; but the
specimens collected by Kohndorff at Manyanga are now before me, and they are
( 390 )
tricolor, but hv no means r/abonensis, though a young specimen has the under
tail-coverts very pale sulphur-yellow. Thus tricolor, layardi, micrits, sjmrnis,
and minor become also subspecies of the barbatas group.
3. Together with the forms of the barbatm group, however, we find others,
which have almost the same plumage, but diflFer in having a protruding, wattle-like,
mostly oranj^e or yellowish eyelid. These forms, therefore, though disguised in a
similar plumage, are specifically different. These " wattled " forms are capensis,
nif/ricanx, reichenoivi and xanthopygos.
4. Another form, Fycnonotua dodsmii Sliarp(>, appears to stand by itself.
There is evidently no wattled naked ring round the eye, though the eyelid is scantily
feathered. /'. dodaoni does not, therefore, belong to the capensis group, and I do
not think that it can be placed as a subspecies of the barbatus group because it
appears to inhabit countries in which forms of the barbatus group partly occur, and
it has also marked differences, so that there is no reason why it should not be looked
upon as a third specialised species. The size is very small, the rectrices have a wide,
sharply defined white tip, the feathers of the back and throat have whitish edges, so
that these parts have a scaly appearance.
Thus we arrive at the following table :
I. Pycnonotus barbatus : eyelid feathered.
la. P. barbatus barbatus : ^Morocco, Algiers, Tunis. Crown brown. Under tail-
coverts white. (28 specimens examined.)
1//. r. barbatus inornatus : Senegambia to the Niger. Like P. b. barbatus, hv\i
smaller. (3.5 specimens examined.)
le. P. barbatas (jabonensis : Kamerun to Gabun. ]Ako P. b. inornatus, hut under
tail-coverts white, edged or sometimes tinged with yellow. (G specimens
examined, including the type.)
As I said before, this form does not extend to the Congo, so far as we know at
present, where P. b. (jabonensis is replaced by P. b. tricolm:
\d. P. barbritus nrsinoe : N.E. Africa (Egypt to Kordofan).* Under tail-coverts
white, crown black. (10 specimens examined.) Wing about 87 — 95 mm.,
i.e. males about 91 — 95, females less.
Ic. P. barbatus schoaiius : Mountains of Abyssinia and Northern Galla-land. Very
similar to arsinoe, but much darker above. This is easily seen in fresh
specimens, but not in worn plumage. (51 examined, among them the
type.)
1/. P. barbatus somaiiensis : Zeila and Somadu. Smaller than arsinoe, but of
about the same pale coloration. (8 examined, among them tlie type.)
This form is indeed very closely allied to P. b. arsinoe, in fact so closely that
even Prof. Reichenow called it P. arsinoe somaiiensis. It differs only by its smaller
size, the wings measuring in the c? 87 — 91, in the ? about 77 to 84 mm. It is
thus evident that this form is not easily recognisable, and I should doubt its
distinctness if it were not for the slender bills which it exhibits when compared
with P. b. arsinoe. Eeichenow's statement that it is paler brown on the ui)pcr
surface is not correct, as far as one can make out from the 8 rather worn specimens
collected by I'aron von Erlanger. On the contniry, judging from a few fresh growing
• Mr. Grant (Xov. Xool. 1!)()0, p. 2")7).saTs that he procuri-il si)eciinens ne.iv Aden ; and Ycibuiy .stated
tliat this .species nests there. I have not been alile to find an Arabian siwcimcn in the ISritish Museum,
but I .should say that Aral)ian specimens cannot be tlie true iirtiiinf.
( 391 )
feathers, I am inclined to think that somaliensis is rather darker than nrsinoe, not
paler. Probably lleichenow's material of arsinoe is very poor. The measurements
have not been given by Keichenow, and no type has been marked. I fix as the
type specimen No. 5, c? (sexed), Dadab T.i January 1900, this being in the best
condition.
Ig. P. barbatus tricolor : S.W. Africa, north to Congo. Under tail-coverts yellow,
crown brown. Axillaries whitish. (26 examined.)
\h. P. barbatus layardi : Eastern Cape Colony northwards, probably to Mozamljique
and Lake Nyassa. Under tail-coverts yellow, crown black. (20 examined.)
1/. P. barbatus micriis : British East Africa and Kilimanjaro. Exact limits not
yet known. Like layardi, but upperside darker, size smaller. (.5
examined.)
IJ. P. barbatus minor : Upper White Nile to Mctoria Nyanza. Like tricolor,
but axillaries greyish brown, and slightly smaller. (9 specimens examined.)
l/i'. P. barbatus spM7'i«s Rchw. (Vui/d Afr. iii. p. 841) : P^nnia, Gallaland.
Very similar to micrus, but ear-coverts quite black, not brown, with oidy the
upper portion black. Apparently more whitish underneath, than micrus. Not
the whole throat, but only the upper part of the throat is black, as in micrus.
There is no close similarity to dodsoni at all, which is described below. Of
the three specimens one is marked as " ?," the other two as " cj." The ? fi'om
Arbadule and the one from Oda Muda have the wings 87 and 88 mm. long, the
S from Arbadule 93 mm. I fix the last as the type, and think the other two must
be both females. (3 specimens examined, including the type.)
II. Pycnonotus capensis : eyelids protruding, wattle-like.
2a. P. capensis capensis : Southern parts of Cape Colony. Eyelid mostly whitish,
underside brownish, paler in the middle of the abdomen. (10 examined.)
2h. P. capensis ni<jricans : Central parts of Cajie Colony north to the Transvaal
in the east, to Benguella in the west.
Under-surface, except the throat and jiigulum, whitish, crown black. Eyelid
as a rule bright reddish or '■ chrome orange." This can easily be seen in well-prepared
skins, and the bright colour is clearly described on each label on the specimens
collected in Benguella by Dr. Ansorge. In two specimens, both marked ? , one
from Katenge, 14. vii. 1904, the other from Huxe, 2. vii. 1904, the eyelid is — though
bare and protruding — blackish, and Dr. Ansorge described it as greenish black in
one, dark green in the other, while in other females it is bright orange as in males.
(21 .skins examined.)
'2c. P. capiensis rcichenou'l : South Arabia. L'pper surface more greyish, eyelid
brownish. (15 examined )
2d. P. capensis xantliopyi/os : Palestine to the Sinai-penin.sula. Like reichenawi,
but larger, bill stronger, ui)per surface not so greyish, black on throat less
abruptly separated. (7 examined.)
III. Pycnonotus dodsoni: eyelid not protruding and wattle-like.
ort. Pycnonotus dodsoni: Middle and Southern Somaliland to Witu and ISIakindos
River (tributary of the Athi River) in British East Africa.
Differs at a glance from all its allies by the liroader whitish tips to the
( 392 )
rectrices, very short wings and somewhat, scaly appearance of chest and back. Under
tail-coverts bright yellow. Wing : c? about 82—87-5, but seldom above 84, ¥ about
74 to 79 mm. (45 specimens measured.) Young specimens have the under tail-
coverts paler, lemon yellow, and the tips to the rectrices brownish.
With this species and the others hitherto known to occur in Somaliland, our
knowledge of the Pymionoti of Somaliland and the neighbouring countries is,
however, not yet finished. We have a specimen shot by Dr. Donaldson Smith
on June 16th, 1895, on the western shore of Lake Stephanie. It is an immature
bird, sex not determined. It is as small as P. dodsoni, the wing only measuring
78 mm. It i.s, however, not dodsoni, as the under tail-coverts are white, and the
back has no scaly apiienrance. I have little doubt that this is a new form, but the
.single specimen before me is young and in moult, moreover its sex is unknown ;
therefore I shall not create a new name for this bird.
Prof Reichenow mentions a male from Ginir as an evident hybrid between
P. arsiuoe fichoamts and P. dodsoni. This bird is an adult male, and Prof.
Keichenow's surmise is very likely correct.
Notes on the Palaearctic forms of the geuiis LANIUS.
It has long been known that the genus Lanivs is not an easy one for those who
study its forms, and that the Grey Shrikes especially are one of the most difficult
groups in ornithology, though several works have been written about them. One of
these, the eighth volume of the Catalogue of Birds, however, is far fi-om satisfactory,
partly because its author was not an experienced systematist when he wrote it, partly
because the material at his disposal was far from being sufficient for such a task.
The other most noteworthy attempt to classify the Palaearctic Shrikes, that by
Professor Modest Bogdanow, is written in Eussian, and therefore unfortunately a
sealed book to the ornithologists of Western Europe; moreover, the material in
Kussian collections is only rich in specimens from the Russian possessions and
Central Asia, and the work, which is entitled Swohopidni Russkoi {i.e. Kussian
Shrikes), professes only to discuss the Shrikes of the Kussian Fauna ; nor is it up to
date, as it appeared 25 years ago, in 1881. It was therefore hailed with delight by
Ur. Kothschild, when an experienced systematical writer, Mr. Ogilvie-Grant, resolved
to write a new review of the genus Laniiis, and the editors of Novitates Zoologicae
accepted his article for Volume IX. of the Tring .lournal, where it appeared in
December 1902.
Mr. Grant (p. 449) invited criticism, hoping "that in this way any mistakes
in the present paper may be corrected before the work ap) cars in the new edition
of the eighth volume of the Catalogue of Birds."
Unfortunately there is ami)le occasion for criticism; but it is not so much a
number of mistakes that nmst be corrected — no work being without mistakes — .as
the entire plan and scope of Mr. Grant's article which is at fault. The work is
not what we should expect in the twentieth century, but it is written with the
perceptions of bygone ornithological ages. The author takes the bull by the horns,
and selects a striking character or two, by which he endeavours to distinguish
what he calls species, without regard to their distribution. Thus " L. deidhdlus,"
according to Grant, occurs from Algiers to Turkestan; " L. jjallidiroslHs " from
Morocco to Mesopotamia and India ; while " L. excubilor " and " L. major " on the
one,"/., homeymi" and " /y. leucoplems" on the other hand, occupy about the
( 303 )
same ranges. Hybrids occur between L. leucopterns and L. fiphenocercus, " many
intermediate forms " are found between L. excvbitor and major, lietween L. decdbatus
ami assiiiiUis, between L. assimilis and pallidirostris, belwceu L. elerjans and
dealbatus, between L. elerjans and Hs^tmi^is, between L. dealbatus and pallidirostris,
between L. assimilis and lahtora, between L. algeriensis and assimilis, algeriensis
and pallidirostris. This is certainly not very reassuring for students who are
desirous of naming a Grey Shrike, and in a case like this the question so often put
by laymen : " Why do you distinguish between such and such forms?'' is certainly
more than ju'^tified.
I'ut the reason why we do not approve of Mr. Grant's method, is that it is an
artificial one. A hundred and fifty years ago, in Linnaeus' time, was the era of
artificial .systems. It was then useful, and a great step forwards. When Linnaeus
classified the plants by their sexual organs, creating families and orders on account
of the numbers of stamina and pistils, it was admired and praised ; but we have
gone past that time — we classify plants by the summary of their characters,
and we have to arrange the animals by their actual affinities, and not by one
or two artificially selected peculiarities. What was admired a hundred and fifty
years ago is not naturally looked ui)on with fiivour in our days, since we have
considerably advanced in method — and esiiecially within the last twenty-five years.
If the general appearance, and not only the markings of the wings and tails, had
been studied, such conclusions as Mr. Grant's could hardly have been arrived at.
Unless we con.sider about one-third of the specimens as hybrids, Lanius excubitor
and L. major, both inhabiting Northern Europe and Siberia to the Amur, cannot be
regarded as two different species. Mr. Grant says that L. excidiitar and L. major are
alike, except that L. niajm- has only one speculum, i.e. the bases of primaries only
are white, while L. excubitor has also the basal half of the six outer secondaries white
on both webs, and that the white terminal portion of the outer rectrices is wider.
Unfortunately both these characters are most variable, and especially that of the
greater extent of white in the tail is perfectly useless; and between the specimens
with only one speculum and those with two we find all sorts of intermediates.
Moreover, both varieties inhabit the same area, and both have been found in one nest !
It is thus perfectly clear that only one form of Grey Shrike inhabits Northern
Europe, this being L. exmbitor L.
A very closely allied form inhabits Northern Asia, but it lias never two specula,
the base of the primaries only being white; the under surface, which gets uniform
white in adult European birds, is always barred, the rump and upper tail-coverts are
more whitish, and the young are dark greyish brown. The name //. major Pallas
1827 is not available for this form, as there is already a Lanius majoi- Gmelin 1788,
which is a synonym of L. excubitor, a fact which has been overlooked by many authors.
The next oldest name for the Siberian Grey Shrike is L. mollis Eversm. 1853.
Eussian authors have persistently adhered to the idea that these dark brown Shrikes
were adult ones, and Mr. Dresser has endor.sed their opinion. Mr. Grant admitted
that the so-called L. mollis were " unquestionably immature birds," and described the
ty\>e of L. seebohmi as the adult male; but nevertheless he placed his i. 7?ioWw far
away from L. excubitor and L. borenlis, which are its nearest allies, placing such
entirely different types as the Red-headed Shrikes and a number of others between
these closely allied subspecies. To this we must object, as even the adherents of the
old binomial nomenclatorial system, as a rule, try to arrange their species so that the
more or less allied forms stand near each other — though of course the trinomial
( 394 )
system makes the relations of forms infiniteljr clearer. It must not be left
nimifntioiieil that Taozauowski correctly described the various jilumages of the
North Siberian Grey Shrike, fully recognising that the brown liirds were the yountf
of the grey ones. It is strange that Russian authors and Mr. Dresser have not
grasped the fact that their L. mollis is the young of a Grey Shrike. Not only do
they bear all the peculiarities of juvenile birds, such as the closely barred under-
surfact" and pale bill, but they are variable, and moreover the North American large
4
Fourth scconcLary from inside of L, c. 2"'-i>i-'<il>^ii-
Grey Shrike, L. boredlis Vieill.. has the same brown young, some being almost
indistinguishable from the brownest mollisl It is possible that this fact was
unknown to the Ivussian ornithologists who wrote about L. mollis, and who were
probably little acquainted with North American birds; but Mr. Dresser could have
known it as well as any one else. In fact, the North American Great Grey Shrike
is so much like the North Siberian one that it cannot easily be distinguished.
Single specimens are sometimes indistinguishable, but the upper surface seems to be
clearer grey, and t lie rump is more tinged with grey in adult birds ; moreover, the
( 395 )
wing appears to be generally shorter. ^Measurements are given in my hook. It is
strange that Dresser, who — except in one or two groups to which he took a fancy,
such as C'inclits — does not, as a rule, separate very closely allied forms, nevertheless
recognised as different " L. fv/ner&is," which is nothing but " mollis."
Under these circumstances I recognise, instead of ;\Ir. Grant's No. 1, L. borealis
(N. America) ; No. 2, L. major (N. ICm-ope and N. Asia) ; No. 3, L. excuhitor
(N. Europe and N. Asia); and No. 33, L. nwUis (N. Asia) — the following subspecies :
Lanius exciibitm' cxntbitor, N. Europe,
Lanius excubitor mollis, N. Asia,
Laniiis excuhitor borealis, N. America,
thus simplifying matters ver}' considerably.
I differ equally from ]\Ir. Grant in the case of his L. homeyeri and L. leacu-
pterus. Instead of taking into consideration the distribution and studying the
variation within a given area, he sticks to an artificial and partly erroneous diagnosis
— based on the lesser or greater extent of the white on the secondaries — and thus
makes these two geographical races occupy the same range. The accompanying
figures will show the variation of the white on the secondaries in the bird named
by Mr. Grant leumpteni^s, a name which is a " nomen nudum," and must therefore,
of course, give way to przeivcdskii Bogd., 1881. The supposed British specimen of
L. homeyeri in the British IMuseum has no exact locality and no history ! It would
be better if si>ecimens which have no better history than " England, out of
Mr. Fred. Bond's collection " — without locality, date, or name of collector — were not
considered in a scientific work.
I must equally disagree with Mr. Grant's treatment of the North African Grey
Shrikes and their allies in Western Asia, where he allows to occur together, iu
the same countries (i.e. N.^^^ Africa) L. heinileucurus, L. elegans, L. dealbatiis and
L. palliilirosiris, and in " Soulh-West Asia and North-East Africa" both L. assindlis
and L. pallidirostris. The inevitable result is, that " intermediate forms " between
the various " species " are frequent. The fact is, that the characters bj- which Mr.
Grant distinguished his species are very variable (see accompanying figures), but that,
nevertheless, there is a certain limit of variation and certain general colorations
are observed which serve well enough to separate the various forms and to cover even
more or less aberrant specimens ; these forms, however, are not species distributed
anywhere at random, but geographical forms (so-called subspecies) of Grey Shrikes,
and their areas are different during the breeding season, although occasionally (as
in Central Tunis) individuals may overstej) their boundaries and specimens may
occur which are not easy to determine, or which may actually be intermediate,
though they are naturally very rare.
In the charming book of Mr. Whitaker on the Birds of Tunis the N.E. African
Shrikes have already been discussed with perfect correctness.
The following details must be understood :
Lanius hemileucurus (Mr. Grant's species 8) is a synonym of L. degans, being
based on an individual with more white in the tail. When Dr. Gadow kept L. Iievii-
leucurus separate, in 1883, he could not be blamed, but nowadays sufficient series
have been collected to show that every intermediate occurs between the specimens
with more and less white in the tail, and that these are clearly individual aberrations.
Lii/itlus elcijd.ns (Grant's sp. 9) is the correct name of the pale Grey Shrike
of Southern Tunis and South Algeria, though originally described erroneously
as coming from " the Fur Countries " in Canada. The distribution gi\ en by
( 3&6 )
Mr. Grant {i.e. South Algeria and S. Tunis) is perfectly correct, and the recent
explorations of Tripoli have shown that it extends eastwards to Barka.
jMr. Grant's No. 10, which he calls " Lanius dealbatus," is a dreadful nii.xture.
The name L. deulbalus was given by Defilippi to a specimen fiom the White Nile,
TliiKl tail-fcatbcr of L. c. elcgajif,
collected hy Brun KoUet (cf. Rev. & Mag. Zool. 1853. p. 289). The description
as well as the critical notes by Salvador! and Finsch, in connection with the locality,
leave no doubt that the name dealbatus refers to the small white-rumped species,
which extends from Nubia to the White Nile. Moreover I have carefully compared
the type in the Turin Museum. Mr. Grant quotes as the original locality " Tripoli,''
( 3'J7 )
evidently because Defilippi adds to his diagnosis the remark "jam extabat a Tripoli,"
which, however, means " it was already there from Tripolis," and not that the type
came from Tripolis ! This specimen from Tripoli, which Dcfilippi thought to be
the same as his dealbatus, is still in the Turin Museum, where I have examined
it also. It is a sj)ecimen of L. elegans, and Defilippi was wrong in uniting it with
his dealbatus. It is, of course, very important to know that the type of dealbatus
came from the White Nile, and not from Tripoli, as in Tripoli elegans is found, and
on the White Nile not.
The name Lanins leucopygos Hemprich & Ehreuberg 1828 has been rejected,
as it has been said to be a " nomen nudum." This, however, is not correct, for, if
one takes the trouble to read the foregoing sheet, one will see that a perfectly
recognisable description is given on fol. d, though the new name of the bird
described on fol. d is only given on fol. e and fol. dd. It is true that the authors
had the silly idea that their bird was a hybrid between L. excubitor and minm; but
as they gave a binomial name to it, and diagnosed it, we must accept it. But even
if L. leii.cojji/ijos had been a " nomen nudum " the name dealbatus could not have
been accepted, because the name L. pidletis (afterwards altered to pallens) of Cassin
had more than a year's priority. Last, not least, L. grimmi Bogdanow, a name
referring to a Transcasi)ian and Turkestan form, does not refer to the young of
Irucopyrjos. but to pcdlidirostris, leucopygos being only known from Nubia to the
Blue and White Nile.
The next species in Mr. Grant's list is No. 11, /y. assimilis Brehm. How it
came that this bird was kept as a separate species is not easy to understand, still
less that it was granted such an area as from Turkestan, India and " Pamir " to
N.E. Africa. L. assimilis is merely a synonym of L. pallidirostris.
A still wider distribution is attributed to species 12, L. pjallidirostris, which
is said to range firom Morocco to India. This is, however, quite erroneous, and
jNIr. Grant's pallidirostris is again a mixture. Among the synonyms appears
correctly i. pa^iicZws Antinori, 1804, but its locality is quoted as " Kadaref, ? Blue
Nile," while Kadaref is a village on the Blue Nile. L. fallax appears wrongly
under L. pallidirostris, and so do dudsoni and hwrifi. About L. alijeriensis dodsoni
Whitaker (-/6zs 1898, p. 599), from Central and Southern Morocco, Mr. Grant had evi-
dently made up his mind that it must be made a synonym of something, as according
to his idea, the types were all intermediate between L. algeriensis, assimilis and
pallidirostris (p. 460). The fact is, that dodsoni is a pale form of alrjerlensis,
between the latter and elegans. It has been discussed in Mr. Whitaker's book on
Tunisia, and in my forthcoming treatise of the Shrikes in Viigel d. pal. Fauiui.
Mr. Grant was also convinced that my L. idgeriensis koenigi from the Canary
Islands was of no good whatever, and so he placed it as a synonym under his
L. algeriensis. We could not blame him much for that, for my koenigi — tliough
certainly not quite the same — is very much like algeriensis ; but on p. 4.59 he
states that dodsoni " are indistinguishable from the palest forms from the Canaiy
Islands." Now if dodsoni and the Canarian Shrikes are " indistinguishable," then
dodsoni should not appear as a synonym of '■^pallidirostris" (p. 459, No. 12),
while koenigi is placed under algeriensis (p. 462, No. 16), far away.
L. fallax, looked upon as the same as "pallidirostris," is a very distinct form,
with more or less grey on the breast and flanks, but unfortunately its oldest name
is auckeri ! The unfortunate decision to classify the Grey Shrikes only by the
markings of the tail and wings, disregarding more or less all other characters.
( 398 )
led Mr. Grant to stale that L. auchen Bp. 1853 was based on birds " intermediate
between L. dealbdtus and L. jM'Uidhvstris."
I have examined the cotype of L. aucheri in the British JMuseum, and Mr.
liellmayr lias confronted the two tj-pical examples in the Paris Museum with skins
from the Tring JIuseum. There is no doubt whatever that L. aucheri is the bird
J\
'\
'I'wo outei- iLctrices o£ X. c, aveheri.
afterwards so well described and figured by Dr. Finsch under the name of L. faU(U.
The following is a de.-^cription of the tyjjical examples of L. dtichcri in Paris.
1. Adult bird, mounted; on label: " L. aucheri Bp. Type, M. Auclier-Eloy.
Perse 1840. No. 10003." Rill entirely black. Narrow, sharjjly defined black
frontal line, rest of upper surface unifoi-m ash-grey (scapulars incomplete !) Upper
wing-coverts i^lack, only the least wing-coverts grey. Only one large white alar
" speculum," reaching from the second to the tenth i)riniary and extending over both
( 399 )
webs, about 40 mm. wide, thus occupying tlie basal half of the feathers. Secondaries
with wide white edges to the inner webs, and with white tips about 7 mm. wide, of
equal width, across both webs. Inner secondaries entirely black to the base, with
white tips of about 1 to 1'5 mm. Lores and usual laniine black markings on sides
of head pure black. Under-surface white, sides of breast tinged with grey. Two
outer rectrices white with black shafts. Third rectrix black with outer web and tip
of inner web white for 21 mm., the black running up along the shaft, in a narrow
line, nearly to the tip. Fourth rectrix black, with tip white for about 10 mm. Fifth
rectrix black, with tip about 2 — 3 mm. white, sixth entirely lilack. Asillaries and
under wing-coverts white, only the under primary coverts greyish. Wing about
114, tail 117, outer pair 28 mm. shorter, bill 17 mm.
2. Adult bird, mounted; on label: " L. aucheri (Bp.) Type. M. Aucher-Eloy,
Perse. 10018 D." Like No. 1, upper surface equally grey, rump as grey as back.
Narrow black frontal line only 1-5 mm. wide. Size smaller, bill larger. Wing 105,
tail 110, gradation 33, bill 17'5 mm. Scapulars complete, therefore with a distinct
white line. Tail differently marked : Outer pair with white outer web, inner web
black at base, tip white for 40 mm. Second pair similar, but tip only white for
34 mm. Third pair, outer web white at tij), towards base only with white outer edge,
inner web black, tip white for about 19 mm. Fourth pair black, tip white for 10 mm.,
fifth with narrow white tip as in No. 1, sixth entirely black. For further description,
diiferences between pnUidirostris and aucheri, and the somewhat puzzling dis-
tribution, I must refer to my forthcoming book, Part IV.
In 1905 Mr. Grant independently discovered that two different grey Shrikes
were found in South Arabia. He named the dark form Lanius arabicus (Bull.
B. 0. C. XV. p. 78, May 1905). Unfortunately, however, this same dark form was
already described as Lanius huryi by Lorenz and Hellmayr (J.f.O. 1901. p. 238).
The mistake which Mr. Grant made in renaming L. huryi — a name which he had
placed as a synonym under his L. 'pallidirostris (\\. 459) — was entirely due to his
unhappy method of separating his species principally by the markings of the tail and
wings. How variable these are is shown by the accompanying figures. If the general
appearance — in this case the dark colour of the upper and under surface — combined
with the distribution is taken into consideration, huryi (= arabicus) is easily
distinguished. We know two grey Shrikes to be common in S. Arabia ; L. exc.
auclieri {= fallax) and L. exc. huryi. Probably the latter is the breeding race,
replacing the commoner migratory aucheri as a resident. Under this assumption I
decided to regard huryi also as a subspecies of excubitor.
For my reasons for keeping separate from the excubitor group the long-tailed
.Hphenocercus and giganteus I must refer to my book. Part IV.
Under Lanius ludovicianus Mr. Grant unites not less than six forms dis-
tinguished by American authors (cf. Kidgway, B. North and Middle Anier. iii.
pp. 235-52). I have not had the opportunity of studying very good series of all of
these forms, but from what I have seen I am convinced that it is impossible to unite
them all, and I do not quite see why they should be united by Mr. Grant, when he
separates so many North African and European forms. It seems to me that mearnsi
and anthonyi are the same, but that the other four forms are fairly well separable
from ludovicianus. All these forms are considerably smaller than L. excubitw
horealis, which lives farther to the north than any of them, but otherwise they do not
differ in any essential characters. I am therefore of the opinion that they should
all be treated as subspecies of L. excubitor.
2G
( 400 )
Of L. inei'idiondlis, which I call L. excali/'lor meridionalis. Mr. (iraiit gives as
the " habitat " : " 8outh of France, Spain and Portugal," and he adds : " Records of
this species occurring in Italy, Sicily, and Sardinia, etc., are apparently erroneous,
though it has undoubtedly been obtained in Corfu and the Ionian Islands."
Unfortunately what Mr. (irant says about the occurrence outside of the regular
breeding-range {i.e., as correctly stated, "South of France, Spain and Portugal") is
not right; in fact, almost the opposite is true. L. e. tnerklionalis extends along
the Mediterranean coast at least as far as Nice, and has occurred, though irregularly
and more or less accidentally, in Liguria, Toscana, Umbria, and even as far as Rome,
specimens being jireserved in Italian museums. On the other hand, the occurrence
on the Ionian Islands is erroneous (cf. Reiser, Fauna Balcanica, iii. p. 569), and there-
is no reason to believe that Lord Lilford's statement of its occurrence on Corfu is
correct, since he evidently only " observed " it, no specimen from Corfu being preserved
in his or any other collection. In Sardinia a young L. minor has been mistaken for
tneridionalis. The same might have happened elsewhere. Nordmann's statement
of its occurrence in S. Russia is sure to be erroneous ; Kolombatovics' note of its
presence near Spalato in Dalmatia is not verified ; Giitke records it from Heligoland,
but no specimen of Tneridionalls exists in his collection : therefore it is probably
erroneous, or, let us say, not verified, like several others of Giitke's records. Under
Laniua mibicus Mr. Grant says, '• North- East Africa and South- Western Asia, ranging
west to Senegal." This is a most incautious statement, as the only reason for it
appears to be one skin in the British Museum which is labelled " Senegal." This
skin, however, was pm'chased from the "Maison Verreaux" in Paris, which is notorious
for its inaccuracy regarding localities, as I ha\e repeatedly had occasion to prove.
Altogether the practice of stating the range of a bird from single occurrences is a
very crude one, because we want to know where a species breeds and where it
migrates. So in the case of L. nubicua, which breeds in Asia Minor and Palestine
to S.W. Persia, but appears to be only a winter visitor in N.E. Africa, and never
occurs in Senegambia or on the Senegal. It would also have been interesting to
state that L. nubicus used to breed, until 1864 at least, near Athens (cf. Reiser,
Ornis Balcanica, iii. p. jJ60).
!Mr. Grant's No. 21 is called Laniics povuranus. The earliest name is L.
senator L. 1758 ; but Mr. Grant, belonging to the small band of British ornithologists
who do not follow the international code of nomenclature, and adhere to the
" Stricklandian Code," does not adopt names prior to 1706, and therefore refused
it. It has likewise been refused by Dr. Schiebel, in his interesting treatise " Die
Phylogenese der Lanius-Arten " in Joiirn. f. Oru. 1906. p. 190. Dr. Schiebel
based his opinion on the consideration that Linnaeus' diagnosis was not clear.
Every ornithologist, however, who has any exjjerience in interpreting Linnaean
names must know that we must judge Linnaean names by the source he took
them from. Now, as it is perfectly clear that Linnaeus did not know the bird,
but merely took his attempt at a diagnosis from Albin, whom he faithfully quoted, or
made it up from Albin's figure, we must look to Albin for a solution, if we are
uncertain; and Albin, though his tigure is bad, undoubtedly figures the Ked-headed
Shrike. Albin does not give a locality for his bird, merely stating that he had it out
of the collection of Sir Thomas Lowther,* but adds that Willonghby had described
a specimen from the Rhine. If is evidently merely a slip that Linnaeus said " In
* It is a thousand pities that tliis rich collection has not been preserved— at least I never lieard of it.
It contained many of the birds figured by Albin, among them the Comaiibis eremita from Switzerland.
( 401 )
Indiis," but, unfoitunatel}', such careless statements occur several times in his
Systema Naturae. V)r. Schiebel has evidently no experience in interpreting old
names, and can hardly have consulted Albin's figure, or he would have understood
that Linnaeus had no idea of describing a bird he knew, merely adding a reference to
Albin (" mit einem bloss angehiingten Hinweis auf Albin "). Also the ventilation
of his personal views about Brisson's names, which Dr. Schiebel treats us with
(p. 192), had better remained unprinted. Nor can I agree, that the next available
name is none other than pomeranus of 1786, because there are one or two before that
date : Lanixis ruficeps Pallas, Vroeg's Gat. Verzavi. Vogelen dieren, AdumWati-
uncula, p. 1, 1764, and L. auriculatus P. L. S. Miiller, Natursysievi, Suppl. p. 71
(1776). I am glad that we are not forced to adopt this latter name, as its diagnosis
is not clear, and the author mixed up with the Red-headed Shrike the common
Red-backed Shrike. On the other hand, he based his miricidati'S primarily on
Daubenton's " Pie grieche rousse," the male of which was figured on that gentleman's
PI. 9. In mixing up with it L. collurio he partly followed Daubenton himself, who
figured a female of i. collurio as that of the Red-lieaded Shrike. Therefore there
was some excuse for adopting the name auriculatus, as Dresser and Gadow ha\'e
done, and in no case can L. auriculatus be quoted as a doubtful synonym of minor,
as has been done by Grant (p. 470).
Mr. Grant's quotation of Daubenton's plates is not correct. He quotes : "Pie-
grieche Rousse de France Buff., PL Enl. i. p. 239, PI. LllL, No. 9, Fig. 2," and
similarly in the footnote. I do not know what PI. LIII. No. 9 means at all, as it is
PI. 9, Fig. 2, but not PI. 53 ; moreover it is impossible to quote "■ PI. Enl. p. 239,"
as the PI. Enl. are plates without text by Daubenton and not by Bufifon, who,
however, together with Montbeillard, wrote the great Hisl. Nat. des Ois., to which
Daubenton's plates, as far as they represented birds, were added as an atlas, though
they originally appeared quite independently.
I am glad to be able to agree with j\lr. Grant regarding the question of the races
of L. senator, of which there are three. IMr. Grant justly unites the Shrike inhabiting
N.W. Africa with the European form. I have been one of the ofi'enders who
separated a paler North African race, and who, worst of all, adopted once upon a time
{Senckenh. Cat. p. 90) for it the name rutilans, which is based on Daubenton's
I'l. 477, Fig. 2. Koenig insisted, in various articles, that the North African form had
a paler head and back, and was a good species. Erlanger declared that Koenig was
wrong, and that the head, in fresh plumage, was at least as bright as in Europe, but
that the under-surface was more strongly washed with ru.sty-buff. Kleinschmidt
(in litt.) believes that the lighter back which is seen in some specimens is peculiar to
North-M'est African specimens, while Hilgert (in litt.) doubts if the head in European
examples gets so bright as in some African birds. I have examined over a hundred
specimens from various parts of Europe and North Africa, and I must now agree with
Grant, Whitaker, and most of the older autliors, that we cannot separate the form
from North-^^'est Africa from that of Southern and Central Europe. Two other sub-
species, howeter, are fairly well distinguishable : i.e. Lanius senator niloticus, with
wide white base to the central rectrices as well as to the rest ; and Laniths senator
hadius, without white speculum, i.e. white base to the outer primaries. The latter
inhabits Corsica and Sardinia, and migrates to West .\frica ; the former breeds from
southern Persia to Palestine, and seems to winter in North-East Africa, where it has
not yet been found breeding with any certainty.
Mr. Grant calls Lanius senatw niloticus binomially Lanius i"ufus. This,
( 402 )
however, cannot be done. Gmelin's Lanius rufvs is based on Brisson's " Pie-
Griesche rousse," which is the European Eed-headed Shrike. The tail is described
by Brisson as half white, and this is the reason why Mr. Grant used his name
for niloticH,fi. But it is quite evident that Brisson described and only knew the
European bird, and he must have overlooked the fact that the base of the central
pair of rectrices was black, while the rest really are about half white. Nor is the
country where niloiiciut lives indicated by Brisson and Linnaeus, and the name nifus
is thus strictly synonymous of seniUor, ruficeps, auricidatus, ijomerunus, and
rvitilus.
Lanius cognalus Breliui, Vogdfang, \). 84 (1855), is a further synonym of
L. senator, and not of collv.rio, as stated by Grant, and, probably following Grant,
also in Keicheuow's great work on the Birds of Africa.
The name L. spinit or quits Yjech-item, a synonym of fo/inrio, dates from IT'.tl,
and not from 1805, as generally quoted, being already published in the first edition
of the Nalurr/esch. Deidschl. ii. p. 392.
A very i)uzzling group are the Asiatic Brown-tailed Shrikes. I mean " Lanius
isabellinus, phoeiiicnroides, cristatus, superciliosus " and their allies. Mr. (irant
separated about as man}' forms as I consider separable, and I even must unite his
numbers 37 and 38 (pp. 472-3); but it appears to me strange that Mr. Grant did
not grasp the affinity of his numbers 37 and 38 with the cristatas-phoenicuroides-
isabellinus grou\>, as the latter are so closely allied to phoenicuroides. Mr, Grant
calls his number 38 " Laniios elaeagni" but this must have been an oversight, as
the name hogdanowi is nine years older, and must therefore be used. Mr. Grant
remarks, on p. 474, " I am doubtful if this bird is really a species distinct from
L. raddei, or merely a stage of plumage." After a careful study of the entire
material iu the St. Petersburg Museum, of the types of L. hogdanowi, L. raddei,
L. dichroiirus, L. infuscatus, " L. phoenicuroides pseudocollurio" of authenticated
spiecimens of L. elaeagni, and some examjiles kindly lent by Dr. Suschkin, I cannot
see any possibility for keeping these separate, thus agreeing with Mr. Grant, and
even going a step further. There is no doubt that this group of Shrikes is a
difficult one, and this fact should be a reason for being careful and hesitating before
naming supposed new forms belonging to it ; instead of this, however, our Russian
colleagues treat us with new names whenever one or more individuals differ from
the usual types. Dr. Suschkin, in an article in the Bull. B. 0. Club xvi. pp. 58-61,
endeavoured to enhghten us about these Shrikes. As we poor ones in the west of
Europe cannot see these birds alive, and Dr. Suschkin collected many specimens,
his article should be of great service to us; but unfortunately it only adds to our
embarrassment. According to Dr. Suschkin, L. raddei, infitsadus, pufludocollurio,
and elaeagni are " suhs\>ecies'' of phoenicuroides, while bogdunowi (^aid to be the
same as varius) is a " variety" (" personal variation " in the text) of L. ph. phoeni-
curoides, and, to add to the number of names, a new aberration is described as
analogus. ^^'orst of all, the distribution is not explained in Suschkin's article ; but
it is evident from the original descriptions, and most kindly confirmed by Dr.
Suschkin in litt., that his idea of subspecies is not the same as ours. Advanced
ornithologists have now agreed to use trinomials for geographical representatives.
Dr. Suschkin employs them partly for forms which breed in the same area.
Oniithologists have hitherto not named aberrations, becau.se we could very well
do without them, the cases in which species or subspecies occurred in numerous
well-marked colour-variations not being very numerous, and there being no necessity
( 403 )
for loading our lists with names for the latter. We thought hitherto that we could
dispense with names for such colour-aberrations, though Naun)ann and others used to
call albinoes and other variations by Latin names; but their names were not actual
names to be considered in nomenclature, because every species in a genus had a
" varietas p((llid((, hUea, alba, variegata," etc. What is now Suschkin's (malogus ?
What is the dift'erence between Suschkin's " jiersonal variation" and his "aber-
ration " ? .Surely we shall be happier without names for aberrations — as looked
upon by entomologists — while I fail to understand what a " variety " is, if the
author has besides the latter " subspecies " and " aberrations " ! I can only recognise
geographical and non-geographical variations. The former we call by trinomials, the
latter we do not recognise in nomenclature, and I sincerely hope that we shall
be spared names for them. In cases where well-marked groups of varieties weie
known, as in many owls, some goatsuckers, a few petrels, Beiiiiefjrelta sacra, and
others, it has become customary to talk of these colour-varieties as " phases." 1
think another expression would be more suitable, as a phase, as a rule, is the term
for a changing stage of a thing, while we know that these colorations are not stages
that change gradually, but that some individuals aiijjear in one, others in the other
colour throughout life. The term "phases" should be retained for age-variations,
so that we may talk of the brown and the white " phase " of the Gannet, the former
being the juvenile, the latter the adult stage ; or of the white and the brown " phase "
of the Snow Grouse, the former being the winter, the latter the summer garment.
To return to Lanius hogihniovn : it is a very x-ariable form. The types of crtcWei
and dichrourus are much the same, but a worn so-called pseud ocollurio shows the
transition to the bogdanowi-, elaeagni-, infiiscattts-tjiye, moreover raddei and
dichrourus do not come from the .same country. I fail to see how Suschkin came to
the dictum that raddei, infiiscal.as, pseudocoUurio and elaeagni are " subspecies,"
while bogdanmvi was only a " variety " or " personal aberration." The distribution
does not seem to lead to such a view! Moreover, I fail to see that Otoinela varia
Sarudny is a synonym of bogdanotvi, as Suschkin says. It is, judging from specimens
named " varia" by Sarudny, and from the descriiition, a synonym of phoenicuroides.
The remarks bj' its author are truly humorous, for he says (according to the trans-
lation in Orn. Monatsber. 1897. p. 183) that this bird looks like a hybrid between
romanowi and karelini, e.xcept that the tail is differently marked ; that one might
consider it as a species, which accidentally straggled to the locality where it was
shot and there paired with a female of rotnaiiowi ! And, if it should be new, the
author proposes to call this energetic bird Otomela varia. A pity it was shot, and we
do not have the hybrids between varia and romanowi !
The list of what I consider synonyms of phoenicuroides is apiialling : montana,
canesrens, ruficepis, canicejjs, romanowi, karelini, varia. I am glad to agree with
Mr. Grant in my view about these, and to be able to state that Dr. Walter Rothschild
has taken great interest in these Shrikes, and that we have together fought many a
battle about them, and at last agreed perfectly about all of them. For the rest I
nuist refer my readers to Part TV . of my " ^'iigel d. pal. Fauna."
In Mr. (irant's distribution of bogdanoim there appears to be a sliii ; Astrabad
being a town in North Persia, he should not have said to Astrabad and North Persia.
In the description of L. phoenicuroides it is said that the female is similar to the
male, but this is not quite correct. The distribution is given very roughly, no
distinction being made (as in the case of isabellimis, tigrinus and several others)
between the breeding range and winter quarters.
( 404 )
Under L. isnbeUmvfi Mr. Grniit has several synonyms which are referable to
phoenicuroides, and the description of the female is not correct. In the description
of L. darwini (a doubtful species, founded on a single skin), the measure of the
tarsus should be 0'95, not 1'95 in.
In the distribution of L. hentef (p. 478) the "Moluccas" are included; but
L. bcntet has never been found on any island of the Moluccas.
Under L. schach hybrids between L.schach and L. fuscatus are quoted, following
Mr. Eickett's suggestion; but this is not correct, the birds in question being young
L. fuscatus.
Under L. vittaius Jlr. Grant describes birds in the " ftrst plumage" and "still
younger ones." This looks funny, but I think I understand it, and this leads me to
the remark that in most books the young birds are not clearly described. In most
cases the pas.serine birds have a plumage in which they leave the nest, and from this
they moult into a plumage which is generally described as the first plumage. It is
not always easy, and sometimes impossible, to distinguish between these two plumages,
our knowledge often being incomplete ; but we should try to describe both these
plumages, and distinguish between them, as the nest-plumage and the juvenile
plumage.
A very interesting article on the genus Latiitis appeared recently in the Jourrud
fur Ornilholoffie. It is a study of the "Phylognese der Lanius-.Xrten," by Dr.
(iuido Schiebel. It is most readable, and contains many interesting facts. The
conclusions arrived at are sound and sensible, though they are, of course, not all
undoubted facts, but speculations, based only on the developments of the coloration
and markings. The article is that of an advanced systematic worker. Trinomial
nomenclature is used throughout. In a few cases the author endeavoured to bring
forward his ideas about the propriety of old names, but in this he failed altogether.
If the year 1758 is taken as the commencement of our nomenclature —and this is
fortunately done by Dr. Schiebel- -the name Lanius senator must be used for the
Ked-backed Shrike, as I have explained above. The judgment of ancient names
requires practice, and Dr. Schiebel would have done well if he had consulted an older,
more experienced ornithologist. Uf tiome forms tlie author had no material, and thus
some errors were unavoidable, but they are few and far between. Very instructive
plates by the author himself accompany the text: birds arrayed side by side in series,
in the shape of skins. This excellent method, first employed by Kleinschinidt, is
most commendable : room is spared, as more forms can be figured on one plate, the
imaginary surroundings of the birds do not distract attention and eyes, and the more
or less arbitrary and often erroneous positions of birds never seen by the artist are
avoided.
I am sorry to say that the name Lnnius caudatus, given by Cabanis to an
East African Shrike in 1869, is preoccupied by Lanius cavdatus Brehm, 1855
{Vogelfanrj, p. 84), a silly synonym of Laniun iitthicus. It is therefore necessary to
rename the bird known as L. cawlatus, and I projiose to call it
Lanius cabanisi nom. nov.,
in memory of the veteran ornithologist .lean Cabanis, who died some months ago at
the age of almost 90 years.
My notes on the Shrikes have become longer than 1 intended, and a great
portion of them consists of criticism and corrections of Mr. Grant's article. My
brother ornithologists, and first of all Mr. Grant himself, will understand the spirit in
( 405 )
which they have been written. They contain, to use Mr. Grant's words, the criticism
of another ornithologist, " who may have paid special attention to these birds." But,
as I have said before, it is not so much the number of errors in quotations, geographical
statements, etc., as the scojie of the " Review of the species of the genus Lanius "
which I oppo.se. We all make mistakes, but we do not all adhere to the system of
systematic study which was reigning in former ornithological times. Our mistakes
must be corrected, for the truth for which we work mu.«t come out by mutual efforts ;
and it is important that we take a wider view and .study birds in the light of their
geographical distribution as far as possible, and that we no longer rely on single
charaeter.s, but thoroughly study the birds in all their parts, in order to understand
their actual affinities. This is less easy than the old system, and especially
geograjihical distribution is sometimes difficult to fully understand, when we do not
know the country, its aspect, geological and climatic conditions, and where maps are
scarce and bad ; but we must try to do our best, and leave it to the future to correct
our errors. And other difficulties, too, stand in our way. Valuable works are
published in Russian, a language not understood, as far as I know, by any Western
ornithologi.st ; new names are sometimes hidden in the text, not showing in
headings ; descriptions are often too short and insufficient, and sometimes taken
from single individuals. Our friends should be more careful in creating new names ;
tliey should, if forms are closely allied, not describe " new .species " from single
examples, and they should boldly show what they do, and not hide their deeds
inconspicuously in the letteiiwess. All these troubles I have felt much during the
work on the last parts of my book on the Palaearctic Birds, but nevertheless it has
been pleasant and fascinating to me, and I look forward with much pleasure to more
such work and to more criticism !
My work has been aided by the kind help of many friends and authorities of
museums. Without this it would have been impossible to come to the conclusions
at which I arrived. ]\Iany an hour has been spent in the bird room at the British
Museum, where I di.'cussed two or three questions with Mr. Grant ; and I have been
able to compare types and series in Paris, Turin, and Florence, in all of which places
I have been received with the greatest kindness — by the late Dr. Oustalet and
Mons. Mi^n^gaux, by Count Salvadori and Professor Giglioli. My warmest thanks
are due to Dr. Bianchi, who generously sent me nearly all the Shrikes in the St.
Petersburg Museum. I am much indebted to Dr. Suschkiu for the loan of some of
his types, to Professors Reichenow and Brauer in Berlin, Dr. Forbes in Liverpool,
Dr. Hoyle in Manchester, Mr. Joseph M^hitaker in Palermo, the late Baron Carlo and
bis mother, Baroness von Erlanger, in Nieder-Ingelheim, Dr. Julius von ^NladarAsz in
Budapest, Dr. von Lorenz in Vienna, Pfarrer Kleinschmidt in Volkmaritz, Profes.sor
Lampert in Stuttgart, Professor Dr. Scliauinsland in Bremen, Count Berlepsch of
Berlepsch Castle, Professor Dr. Wilhelm Blasius in Braunschweig. Herrn N. Sarudny
in Pskov, for kindly lending specimens in their possession or under their care ; to
Dr. Ijouis Bureau, llerrn N. Zarudny, Herrn Dr. llartlaub in Helgoland, Herrn Sergius
Buturlin, Dr. Bianchi, Dr. Suschkin, Herrn von Tschusi, Herrn Hilgert, Baron Loudon,
and others for kind information, translations, descriptions; and last Ijut not least to
Herrn Hellmayr for making comparisons for me in the Paris Museum.
( 406 )
SOME NEW SPHINGIDAE IN THE BRITISH MUSEUM.
By the HON. WALTER KOTHSCllI r.D and KARL JORDAN.
L Polyptychus poliades spec. nov.
6. Ill size and pattern nearest to J', ort/zof/rap/ius R. & J. (lOO:!). Pinkish
jirey. A Inown mesial stripe on head and anterior portion of thorax. .Joint of
palpns open, second segment rounded off. Tibial s])urs somewhat shorter than in
/'. ortlixiqruphus.
Wings, npperside : forewing with brown lines : a short thin subbasal line,
a more prominent line at basal fourtli, cnrving costad, an anteiuedian ilonble
line, feeble, bifle.KUOse, curved basad anteriorly and ]iiisteri(irly and in niiiUUc ;
interspace between this double line and the preceding one brownish : a small
stigma with jiale centre ; an almost straight discal line, a little more proximal
at costal margin than the corresponding line of P. orthograjjhtts, followed by a
faint erenate line and a .•^ubmarginal line, the latter being straight from costal
margin to R^, and faint and flexnose farther back ; hinder angle less prodnced
than in P. orthoyrnplnis, no distinct black-brown sjiot near this angle. Hindwing
pinkish drab, deeper in tint than forewing, grey at anal angle ; a trace of a brown
median line and of an interrnpted snlianal bar ; fringe dotted with brown.
L'luh'rside more clayish grey than npper, both wings with a curved discal
line and a row of submargiual dots, the" dots of hindwing being replaced near
anal angle by a line.
(Genitalia : tenth tergite broad, narrowed beyond middle, then widening again
a little, with an antemedian tubercle at the lateral edge, apex sinuate, the lobes
broad, obtuse, somewhat slanting downwards ; tenth sternite divided into two
obtuse lobes. Clasper sole-shaped, with large patch of small friction-scales, the
eighth tergite bearing laterally on inner surface a band of large friction-scales.
Harpe nearly as broad proximally as the valve, and two-thirds the length of tlie
same, sinuate at apex, the two lobes short and obtuse, the njiper a little longer
than the lower, both denticulate ; dorsal edge of harpe prodnced at base into a long
thin process which projects anad. Penis-sheath thin, bearing at apex a long pointed
process which lies close on the sheath, being directed proximad.
Length of forewing : 33 mm.
llab. Obuassi, Ashanti (G. E. Bergmann).
One c?.
Typhosia gen. nov.
6. Paljjus closely appressed to the head, second segment much longer tiiau
the first ; joint not open. Antenna less deeply grooved than in Lycosphinx
\i. it .L (1003); end-segment somewhat siiorter, without long bristles, the scales
of upperside iMojecting beyond apex. Posteriorly below eye a tuft of hair-scales,
projecting forward, covering half the eye. Abdominal tergites with dispersed flat
spines all over, besides the nnmerons spines at the edges. Tibiae spinose, s])urs
as short as in Lycosphinx^ one pair to hindtibia, midtibia as long as midtarsiis,
hindtibia somewhat shorter than tarsus, but more than half as long again as
tirst tarsal segment.
Neuration as in l.ycosphinx, but SC^ and R^ of hindwing on a shorter stalk,
D- shorter, M- less prdximal. Distal margin of forewing dccjily emarginate between
( 407 )
the pointed apex and M', sinuate behind M', angulate at this vein, hinder margin
•deeply emarginate proximall^- of angle, the latter therefore produced backwards.
Type : T. illustrts s])ec. nov.
Nearest to J^ycospliinx R. & J. (1003), from which it difl'ers especially in the
eyes being covered by a brush of hair-scales as in Rkodoprusina and Plnjlloxiphia
R. & J. (1903).
'Z. Typhosia illustris spec. nov.
c?. Body creamy butf, washed with olive anteriorly.
Wings, upperside : forewing creamy buff, somewhat shaded with clay, marked
•with olivaceous brown lines ; a double line crossing cell at base of M-, two parallel
lines on disc, dentate on M' and M-, the second crossing subcostals just distally of
fork; a liar R^ — M' nearly halfway to distal margin as remnant of a third line,
which is faintly indicated at costal margin, an evenly curved line from tip of wing
to R^ and continued from here straight across lower angle of cell to costal margin,
which it reaches not far from base. Hindwing rnfous-red, costal margin cream-
colour as far as covered by forewing, abdominal margin creamy buff, this border
widened forward distally and here irrorated with some brown scales ; fringe brown.
Underside buff, irrorated with brown scales. Forewing rufous red from base
to disc, a somewhat S-sbaped line from subcostal fork to hinder margin, crossing R'
about 3 mm. from cell, brown in front, reddish behind ; a second line about 3 mm.
from the first, parallel with it, but dentate, brown, a sharjily marked but thin brown
line from apex to R^ joining here the second line. Hindwing slightly washed
with red posteriorly at base, two parallel lines on disc, the first heavier than the
second, almost touching lower angle of cell, ending at costal edge in a brown spot.
Genitalia : Tenth tergite almost completely divided into two processes which
stand separate proximally but are converging, each process with almost parallel
sides and ending suddenly in a very short brown point. Clasper without friction-
scales, long, pointed, nearly symmetrical ; harpe very short, basal, consisting of
-a small curved piece of brown chitiu. Penis-sheath without armature.
Hah. Obuassi, Ashanti (G. E. Bergmann).
Two (?c?.
3. Macroglossum hirundo samoanum snbspec. nov.
cJ. Smaller than .17. It. citiense and It. Itiruiu/o ; straight black-brown median
band of forewing distinct, but the grey band absent; yellow baud of hindwing as
broad as in /(. hirundo ; yellow side-spots of abdomen also as in the Tahiti form.
llah. Samoa.
One i.
Connects ritiense with liirttmhi.
4. Macroglossum hirundo confluens subspec. nov.
? . Close to M. It. lifuense llothsch. (1894), from Lifu. Black-brown antemediau
band of forewing broader, almost completely merged together with the median
band ; yellow band of hindwing broader than black-brown basal area. A small
yellow side-spot on third alxlominal segment and a few yellow scales on the fourth
segment.
IJab. New Hebrides.
One ? .
( 408 )
SOME NEW ANTHRIBIDAE EROM THE COLLECTION OF
H. E. ANDREAVES.
By KARL JORDAN.
1. Mecocenis allectus elegans subspci-. uov.
c? ? . Similar iu jiattern to M. allectus maculatus Jord. (1894) from Teuasserlm,
Burma, and Assam. The elytra much longer in proportion and more regularly
tessellated ; the mesial vitta of the head broader, and that of the pronotum strongly
constricted before middle, the triangular apical portion of the vitta being separated
from or just touching a round spot which stands on eacli side of the disc before the
middle.
Hub. South India : Anamalai and Nilgiri Hills:.
" Flying among felled trees and on living trunks, at 3oOo to 4U0U ft., in May.
Very wary. In the Ouchterloney Valley in the Nilgiri Hills during November
and December at 3500 ft., on recently felled Ficus and other trees " (H. Leslie
Andrewes).
~. Physopterus analis spec. nov.
c??. Black, the tibiae rufescent; pubescence grey beneath, more olive above,
being obviously grey in middle of pronotum, at the base and apex of the elytra, the
grey ajjical area of the latter occupying about one-third, being rather sharply limited
in front and rounded ; third elytra! interspace with some black pustules, one of
which, situated just in front of the grey ajiical area, is more prominent ; the other
alternate interspaces also with small black spots ; a brown-black spot upon suture
at apex : tibiae ringed with brown at base, iu middle, and at apex.
In structure similar to P. tuberculatus Jord. (1894) from Ceylon. Elytra much
more convex, and the postmedian pustule much less elevate. Hindfemur of <S with
an acute tooth on the underside beyond middle.
Ilab. South India : Anamalai Hills.
"In May, at 3000 to 4nou ft." (H. Leslie Andrewes).
3. Sintor andrewesi spec. uov.
cJ¥. Brown-black. Kostrum with parallel sides, very feebly dilated at apex,
mesially carinate, somewhat impressed along the carina, at base mesially grooved,
like the head unifurmly covered with a greyish clay jmbescence. Eye small.
Antenna rufescent, cliilj black. Prothorax uneven above, depressed, subtuberculate
in centre, and somewhat elevate also laterally of depression ; carina interrupted iu
middle, evenly tiexed forward at sides. Elytra flattened at suture, especially before
ajiex, basal callosity tuberculiform, black in front, a large tubercle befjre apex
between the third and eighth stripes, third stripe ending in a small tubercle,
sutnral and alternate interspaces tessellated and pustulated with black ; a median
jiustule in lifth interspace, and a submedian one in third interspace larger than the
others and bordering a greyish area occupying the sutnral depression. Legs
rufescent, tibiae and tarsi annulated with brown ; midtibia with apical mucro (J).
Length, 4J to .5] mm,
Uab. Nilgiri Mills.
( 409 )
" At 6000 feet, by beating " (H. Leslie Audrewes).
A few specimens.
The tubercles of the elytra and the small eyes separate this species from all the-
other forms of Sintor.
4. Habrissus sellifer spec uov.
cJ?. ^imil-a.r to n. indicug Jord. (1903), but head and rostrnm with sharply
marked grey mesial vitta; eiid-segmeat of aatenna black-browu like the other
segments of the club, not luteons ; disc of pronotnm distinctly bi-imjiressed, the
grey markings ill-defined ; the grey transverse postmedian band of elytra further
back; mesosternal process more rounded; metasternnm of c? with velvety lilack
spot on each side, without combs.
Hab. Nilgiri Hills (H. Leslie Andrewes).
Several specimens.
Phloeomimus gen. nov.
(J ?. Head similar to that of Pkloeohhis Sch., but neither carinate nor sulcate.
Antenna not reaching beyond base of prothorax, segments 2 and 3 thinner than the
others, 5 slightly thicker than 4 and G, esjiecially in c?, 8 as long as broad, club
hardly longer than segments 6, 7 and 8 together. Eye as in Pkloeobius, coarsely
granulated, deeply sinuate. Antennal groove not covered, transverse. Prothoracic
carina basal, laterally extending to middle. First tarsal segment short, claw-
segment of foretarsus a little shorter than segments 1, 2 and 3 together.
Distinguished from PIdoeobius especially by the antenna. We are acquainted
with only one species, which resembles Basitropis in shape.
5. Phloeomimus griseus spec. nov.
<??. Subcylindrical ; brown-lilack, somewhat rufescent ; densely covered with
a clayish grey pubescence, the npperside being indistinctly marked with brown,
Kostrum slightly impressed in middle. Eye about four times as long transversely
as broad in centre. Prothorax broader than long, truncate, widest a little behind
middle, being rounded at sides ; puucturatiou very dense ; angle of carina a little
over l)ti , but not rounded. Elytra cylindrical, practically truncate at base, basal
callosity just vestigial ; regularly punctate-striate, liearing dispersed indistinct
brown dots. Pygidium twice as broad as long, rounded. Prosteruum depressed in
front, very short. Mesosternal j)rocess vertical, curved backwards at apex.
Length, 5^ to ~\ mm. ; breadth, 2 to 2i mm.
Hub. Tharrawaddy, Burma.
Several specimens.
( 410 )
TWO NEW XEXOCERVS (AyTHRIBIDAE) IN THE
COLLECTION OE R. VON BENNIGSEN.
By KARL JORDAN.
1. Xenocerus bennigseni spec. nov.
? . Black ; pubescence creamy white. Antennae reaching beyond middle of
^lytruui, pL'caliar in strnctnre ; segment 7, base of 8 and apex of 6 grey, 1 to G
clothed with brushes of brown hairs ; segments 1 and 3 short, 2 almost thrice as
long as 1, equalling 4, this a little shorter than ;j = 6. I'rothorax broadly and
deejily impressed longitudinally in middle, the imjiressiou occupying nearly one-third
the width of the prouotum, narrowing apically, tilled in with creamy-white
pubescence ; a slightly darker vitta on the side, partly obliterated. Elytra broadly
impressed at the suture, the impression rather sharply limited laterally ; apical
declivity irregular, each elytrum being elevate in middle at the beginning of the
declivity and impressed laterally of this elevation ; base also more strongly
impressed at shoulder than usually ; pubescence creamy white ; a large lateral
area brown, reaching from basal fourth to apical declivity, irregular, extending
upwards in middle to the elevate third interspace, and posteriorly produced into
a narrow, irregular, transverse band which runs over the suliaiiical elevation and
is dilated forward upon suture; two large round dots on elevate third interspace
{but extending to second and fourth interspaces) black, one subbasal, the other
autemedian, joined to the fuscous lateral area ; humeral and apical areas somewhat
marmorated with fuscous.
Pygidinm creamy white. Underside the same laterally, less densely pubescent
in middle ; abdomen with a row of small black lateral spots. Process of mcso-
.sternum sub-bitubercnlate. Femora and tibiae rufescent.
Length : 17 mm.
llab. Sattelberg, Hnon Gulf, German New Guinea (R. v. Bennigsen).
One 3.
The remarkable structure of the antennae and the broad impression of the
pronotum and elytra distinguish this species from all the others which are known.
2. Xenocerus lacrymans herbertus snbsjjec. nov.
(J ? . Differs from A'. /. lacnjimns iu the pattern of the elytrum : the basal
sntural stripe is thinner, the rounded cordiform sutural spot is replaced by an
anguliform spot,. the anterior lateral spot is transverse, irregular, the postmedian
lateral spot is more transverse than in /. lacnjmans, being often continuous with the
sntural anguliform spot, and the apical si>ot is smaller.
Ilab. HerbertshOhe and Baining Jits., Gazelle Peuiusnla, Neupommern
(R. v. Bennigsen).
Two jiairs.
This form is interesting inasmuch as it forms a transition from the i)attern of
X lacri/nnvts /acn/ma/is to the ordinary Xenocerus pattern.
' ?9 SEP 1906
NOYITATES ZOOLOGICAE.
Vol. XIII. AUGUST, 1906. No. 3.
A REVISION OF THE AMERICAN PAPILIOS.
By the HON. WALTER ROTHSCHILD, Ph.D., and KABL JORDAN, Ph.D.
AFTER the pnblication, in 189.5, of the Revision of the Papilios of the Eastern
Hemisphere, exclusive of Africa, we intended to coutiune the work by
revising also the Pajiilios of the Aethiopian Region and of America. However,
we soon found that the material at onr disposal from these regions was not
extensive enough, and therefore pcstponed the researches. We knew our task
to be s])ecially difficult with some groups of American Papilios, which are either
so variable individually or present such slight external specific differences that, in
the absence of breeding from the egg, a comparison of long series of specimens
appeared to us necessary to render the conclusions reliable. Since 1895 the
collection of Lepidoptera in the Tring IVIuseum has been steadily increased, so
that, after the issue of the Revision of the Sphingidae in April 1903, we considered
the material large enough to base upon it a Revision of the American Papilios.
We undertook the work the more readily, as we were assured of the kind assistance
of several friends who were willing to place the material in their collections at
our disposal for the purpose of this Revision. Many of the deficiencies in our
collection have thus been made good, and the number of specimens compared has
been rendered considerable in the case of most species. We take this opportunity
of tendering our very best thanks to all who have so kindly assisted us.
The collection of the British Museum, containing the types of the numerous
names given to Papilios by Doublcday, Gray, Hewitson, and others, and the
collection of Mr. F. Ducane Godman, comprising not only a large Central American
material, but also many South American forms which are rare in collections, have
been of the greatest help to us. We have also examined the specimens in the
large collections of Messrs. H. Grose-Smith, H. Druce, and H. J. Adams, as well
as the Hopeian collection at Oxford, all of which contain material which it was
very important for us to see. Of Continental collections we have visited those
of Monsieur Paul Dognin, the Paris Museum, and Mon.sieur Charles Oberlhiir ;
and Dr. Rebel has been kind enough to send us some of the types of Kollar and
Felder contained in the Hofmuseum at Vienna. The most valuable assistance has
been rendered to us by Monsienr Charles Oberthlir, whose magnificent collection
comprises, in addition to the numerous specimens described by Boisduval, and
others described by Lucas, a very large recent material, among which there are
a number of forms which we have not seen in other collections, several being
undescribcd.
We have further been assisted in our task of clearing up the synonymy by
photographs of some Godartiau types preserved in the Royal Scottish Museum
at Edinburgh, and of some llopflerian specimens iu the Berlin Museum, and we
27
(412 )
are also indebted to Herr G. Weymer for a sketch of P. orthosilaus. AVo liave
not had au opportuuity of visiting for the purpose of this Revision the fine colliction
of the late Dr. U. Standiuger. However, we do not think that we have made a
mistake in regard to any of the forms of I'apilio of which the name-type is
contained therein, nearly all the forms described by Dr. Standiuger being now
also in other collections.
The series of Papilios sent for inspection by Prof. Dr. Goeldi, the founder
of the well-known Goeldi Mnseum at I'ara, lias been very serviceable. We were
the more pleased to have these specimens for examination, as our own material
from the Lower Amazons is very limited.
With the exception of the North American forms, the literature on tin;
American Papilios is generally not very extensive. There are comparatively few
local lists of South and Central American Lopidoptera, collectors of neotropical
butterflies apparently not being given to publishing lists of captures, as is so
frequently done with African Lepidoptera. We are not over-sorry that this is so ;
for we have found some of the lists more a source of trouble than a help, on account
of the species being })artly enumerated under obviously wrong names.* In the
case of some difticnlt species, such as P. protesilaus and P. iphidamas, we have
been obliged to discard a number of references, as we could not possibly ascertain
which of the numerous allied forms is concealed under the name quoted in the list.
Utherwise we have endeavoured to make the bibliography fairly complete. Most
of the works referred to are in the Tring Museum, and those which are wanting
have been consulted at the British Museum, very few books being quoted in this
llevision which we have not seen ourselves. We hope, therefore, that we have not
missed any observation of importance bearing on our subject, or names proposed
for American Papilios.
However, although there is little written abont the greater percentage of
American Swallowtails, the literature jiresents nevertheless a great deal of nomen-
clatorial intricacy, which it was one of the olijects of this Revision to unravel.
Owing to a great looseness in the descriptions published by some of the old authors,
and an equally great arbitrariness in the application of names, the uomenclatorial
puzzles are numerous and partly difKcult to solve. This laborious research in
mere nomeuclatorial matters might easily have been avoided for the greater part,
if a little more preciseness had been exercised on the part of the authors of names.
We can only regret the unnecessary burden unconsciously put on the revisers by
former authors ; but we express the hope that it will serve our contemporaries,
as it has served us, as a warning not to be equally loose in matters nomeuclatorial
and vague in the introduction of new names. A name too many does not much
matter, if one knows to what it is meant to apply ; but a name which one does not
know where to place correctly is a great nuisance.
It is now close on a century and a half since tlie publication of Linne's
Si/stema jSatiirae (1758), the starting-jxiint in nomenclature. The knowledge of
American Papilios was extremely meagre at that time, only seven diiferent species
{glauiKs, jiliiknor, thoas, acneas, anckiscs, and protesilaus) being distinguished by a
* An erroiR'uu.s name for a .siiecies is distiiii^uislicd by us in the synonymy from lliu prcdccupieil
name by putting err. det. {error dttcrmiiiutionls) after ihe autbur wlio misapplied the name. For
instance, Fajiilia a^terioides Slaudinger (non Keakirt, l.sG:t, en-, det.) means tbat Staudiuger called an
insect aMrriindcs ileakirt which was not Ueakirt's insect ; while I'ajnUo mentor Boisduval (non Jialman,
1S23) means that lioisduval gave, in 1836, to a new species the name mentor which had already been
employed in lS2a for another insect.
( 413 )
naiue in tliat, ruuJiimeutal work, au eighth name (aiitilochiis) being proposed for
an exaggerated and partly fictitions figure of the male of P. glaucus. Small as
this number is, it presented a good deal of contentious matter for subsequent
authors to write about. Linne laboured under the great disadvantage that, as a
reformer of the methods in Natural History, he bad to base his work not on actual
specimens alone, but had also to include the recorded forms of animals which he
did not know authentically.
Since the pre-Linuean descrijrfious and figures are for the greater part
very poor productions, Linne was fre((ueutly misled to quote these figures and
descriptions in a wrong 2)lace. Some of his erroneous quotations, however,
are doubtless due to mere oversight or absentmindedness — as, for instance, the
citation of Edw. av. 34* under Papilio ajax in 1758, under P . protesilaus in 1764,
and again under P. ajax in 1767, Edwards's figure not agreeing iu the least with
the other quotation given in 1758 and_ 1767, or with Linne's own description.
The citations under P. protesilaus comprise several Nymplinlidar and Papilionidae,
the references under P. anchises being also wild. Now, iu dealing with such
names, it is obvious that the description given by Linne must be the primary
guide in the apj)lication of the name ; at least, so it appears to us. The three
Linnean names are good illustrations of the diflerent results one arrives at. In
the case of P. anchises the description and the reference to the Museum Ludovicae
Ulricae (M. L. U.) leave no doubt that the insect figured by Clerck in 1764 is
the true P. anchises, though post-Linnean authors have often erred iu tbe
application of the name. No such positive result can possibly be arrived at in
the case of P. protesilaus. From the descriptions given by Linnd in 1758, 1764,
and 1767, we can only conclude that P. protesilaus was one of the numerous
white neotroiiical species distantly related to P. potlalirii/s. Among the figures
referred to by Linne in 1758 there is only one which does not contradict Linne's
descriiJtion. This figure of Merian is unfortunately very incorrect. In 1 764 Linn6
gave a better description, which, taken in conjunction with Clerck's figure referred
to by Linne, applies best to that species to which we have restricted the name iu
this llevisiou. Since neither the figure nor the description is exact enough for
absolutely certain identification, it is obvious that Linno's specimen, if he had one,
may very well have belonged to one of the other white Papilios. However, it
would only be possible to disprove the correctness of our application of the name
protesilaus, if the authentic specimen from which the description was presumably
takeu were 2)reserved, and iu a sufficiently good state of preservation to exhibit
those delicate differences by which the various species allied to P. protesilaus are
distinguishable. Perhaps one might agree in this case with Mr. G. H. Verrall t that
it is fortunate there is no such Linnean specimen, since it is really very indifferent
which sj)ecies bears the name protesilaus, as long as there is no j)ossibility of our
application of the name being justly reversed. | Nevertheless, this case renders
* Kdw. av. p. 36.1. 226 is quotecl_by Linne in 1767 on p. 756 under \Aa Papilio Equcs HcUamiHs
thfillo, and again p. SOT under Sjjhiti.v j>i'ctiiiici>rnix.
t I'rusidential Address, iu J'ruo. JCiU. Stic. Land, for 19.00. p. 47.
J There are two speeimens of this gioup of species in Linne's culloctiun preserved at the
Linnean fiociety iu London. One speeimeu is similar to Clerck's (igiirc, wldlc the other is V. IdcMaus.
Unfortunately a former liljrariau, under wliose ctire (?) the collection was, tliought fit to snpitlcment the
collection by adding fresli specimens ! ! It is therefore hardly possible to say if these two I'apilios, or
cue of them, were originally in the Liuuean collection. The one which agrees fairly well with Clerck's
figure has no abdomen.
( 414 )
it quite clear that, as the distingnishing characters of the various si)ecies in
question have never been recognised, absolute certainty as to the application of the
name jjiotcsilaus might oul)' be arrived at by ourselves examining the original
specimen on which the description was based, or the " name-type," as such a
specimen may conveniently be called. In all instances where new distingnishing
characters are discovered between forms which were formerly considered the same,
it is nearly always indispensable for a conscientious reviser to examine the name-
type of each form. As long as the baptizers of new forms are not omniscient,
and hence arc liable to publish descriptions and figures which future discoveries
may render insufficient, a great deal of instability in nomenclature and of haggling
about names, and therefore of waste of time, would be avoided, if every name
introduced were made monotypical, and the name-specimen carefully preserved.
Many systematists are fortunately in the habit of doing this, thus saving future
classifiers much unnecessary labour. The habit of designating as ty])e every
specimen of the series the describer had originally before him is not to the point,
since there is no guarantee that all these specimens are the same. The history
of American Papilios offers many examples of composite species, and not only
among those described in the eighteenth century, as will be seen in the body of
this llevision. If Linne had been quite precise in the aj'plication of his names,
fixing each uame to one particular specimen or a previously published figure or
description, we should not now be in such a peculiar predicament with regard
to his Fapilio ajax as we are placed in. As said above, the descrij>tion of this
r. ajiix and the two references given beneath it contradict one another, each
ap])lying, without the slightest doubt, to a different insect. The description fits
the Papilio described later as poli/xenes by Fabricins and as asterius by Cramer,
and does not agree with the species which is generally known as P. ajax. If
we had here to do with some little-known insects, we should hardly hesitate to
apply the name ajax L. to the insect figured as such by Clerck — namely,
2>olyxenes Fabr.
However, there is an enormous literature on both these insects, and the
rejdacement of the names pob/xcncs or asterius by ajax would lead to endless
confusion. The whole mischief is occasioned by Linne's reference under 1'. ajax
to Edwards's figure. Now, this reference Linne himself removed to P. prolcsilam
in 1704. Under this same name protcsiluus we find in 1758, 17f)4, and 1707 a
reference to a figure in Catesby which represents the same insect as Edwards's.
And in 1707 Linnt^ described Papilio xuthus as being similar to P. ajax, wiiich
would have been quite ludicrous if Linne's ajax had been the insect now so called.
There is a remote possibility that Linne described a,jax from a small male of
P. i/luiicus. For this reason we have thought it advisable to overcome the difficulty
by rejecting the name ajax altogether on the ground of its being of doubtful
application.
The name ajax does not appear in Linno's ^fuscum iMciovicac Ulricae ; this
is unfortunate, since the descriptions given in that work are far superior to those
of the Si/strma Naturae of 17uS and 1707.
The most famous and, at that time, the most iuiportauL ]iust-Linnoan works
on Entomology were those of Linne's disciple Fabricins. The Si/stcma Entomologiae
of 1775, the Spixies Tmcctoram of 17S1, etc., were conceived on the same lines
as Linne's Si/stema Naturae. They gave a short, concise classification of all the
insects known to the author either from specimens or previous i>ublicatious. No
( 415 )
entomologist has ever exercised so ninth inflnence on the works of others, and for
such a long time, as Fabricins. Nowadays the Fabrician works are rather a source
of trouble. The descriptions are no better than in the Sijstema Naturae, and the
frequent changing of names indulged in is most confusing. The nomenclatorial
errors of Fabricius have, unfortunate!}', been perpetuated by subsequent authors,
many of whom, as late as the middle of the last century, followed Fabricius blindly,
giving his names preference to older ones, and accepting his identifications even
if known to be erroneous.
The habit of supplanting the name of a species by a new one is bad only
in so far as it swells the synonymy unnecessarily ; but very pernicious it is to
transfer arbitrarily a name from one species to another. This is occasionally done
even nowadays, though not in so flagrant a way as by Fabricius. Some authors
appear to be of the opinion expressed by Boisduval in 1836 — that a name which has
become a synonym on account of another name having priority, may be used again
in the same genus for another species which has no name. Boisduval named
a Brazilian species P. mentor according to this principle, though the name had
already been employed for a closely allied species but become a synonym, stating :
" Dalman a donne le nom de mentor k un autre PapiUo qui est le suivant, et
qu'flubner avait fait connaitre avaut lui sous le nom de hjcophron ; nous avons crn
pouvoir prendre sans inconvenient le nom de Dalman pour I'appliquer ii celui-ci
qui est nonveau."* As this principle leads unavoidably to confusion, we are
strenuously opposed to it. We go even farther, and maintain that a name should
not recur within the same genus even as a name for varieties.
A good many of the Fabrician descriptions were taken from specimens which
he had seen in England daring his several visits to this island, and from the
unpublished drawings of Mr. Jones, of Chelsea. These drawings are now in the
possession of Dr. Drewitt, a descendant of Jones. Dr. Drewitt has kindly allowed
us to examine the drawings and to take photographs of some of the figures.
The two volumes are in the best state of preservation, while Jones's collection
of insects had much suffered before it came into Dr. Drewitt's hands. The execution
of the drawings is admirable. There are ten names for American Papilios char-
acterised by Fabricius from Jones's drawings, respectively from specimens which
had served as originals for these drawings (^2^elaus., acamus, dardanus, tros,
zaojntlnis, dimns, idaeus, iliis, iphidamas, and komerus). The specimens were
in Drury's collection, with the exception of homenis, which was contained in the
collection of Latham. Since the Fabrician descriptions are mostly rather meagre,
and as most of his types have disappeared, the preservation of Jones's drawings
is a very fortunate circnmstance.
The butterflies described by Fabricius were revised in 1809 by Butlor ; but
this revision was not so thorough as the subject required. With the help of
Jones's drawings we have been able to identify all the Fabrician Papilios about
which there was some doubt.
In 1779 appeared the third volume of Goeze's Entomologische Beytri'ige, which
is a kind of catalogue enumerating all the Lepidojitera known at that time. A few
critical remarks are oflered by the author, often beside the mark ; and .some new
names are introduced for some of Seba's more or less bad figures, which might
with advantage have been left unnamed.
The short descriptions of Linne and Fabricius being generally insufficient for
• Sjjec. Gin. Lip. i. p. 352 (183G).
(416 )
precise identification, it was soon recognised that frood illnstrations were a necessity.
The first ironograplicr prndnciiig figures wliieli can be culled good was Itosel.
The prodnctions of the earlier authors (Petiver, Mofi'at, Merian, etc.) ,as well as
of some later ones, are mnch inferior to the plates in the Inseltten-Belustigungen.
There are only a few figures of Exotic Insects in Rosel. The first iconography of
great importance for the nomenclature of Lc])ido]itera, however, were the /roues
Iiisectontm of Olerck (1764), the figures of which were for the greater part talcen
from specimens in the museum of the Queen Ludovica Ulrica, and hence may be
considered typical.
Shortly after the appearance of the twelfth edition of the Si/stenia Ndfiirae,
the last edited by Linne himself (1767), Drury published his Illmtrations of
Natural History, which is nsnally quoted for the sake of convenience under its sub-
title as Illustrations of Exotic Insects. The three volumes contain only insects.
The plates are nearly all very good for that time. In using the work one should
bear in mind that in some cases the localities are erroneous, some North American
insects being stated to have been received from Jamaica, and the reverse. However,
in respect to localities the work was a great improvement on former authors.
If we find the localities given in the works of the earlier writers often
deplorably vague and frequently erroneous, we miist remember that the majority
of the specimens were collected by people who did not take an actual interest
in Natural History, but brought the specimens home as curiosities from foreign
countries. Our forefathers iu systematics had not the good opportunities we have
nowadays of obtaining correctly localised material. They had to be content with
what they got. The wrongly localised specimens misled them often to attribute a
much wider distribution to many tropical species than they actually have, it being
stated of many species that they occur in all tropical countries. The knowledge
of the great importance of exact localisation has come very slowly ; but we may
now fairly say that every serious student of some branch of systematics is aware
that specimens without exact locality are of little value to the scientist.
Being acquainted with many large collections of Lepidoptera, we note that
the progress made during tlie last ten years in labelling insects is very marked.
While the labels, if there were any, were formerly usually hand-written, bearing
mostly only such general localities as Brazil, Veueznela, West Africa, etc., we
find the labels of recent additions mostly printed, giving often, besides the exact
locality, the date or season of capture, altitude, name of collector, and even some
biological fact. No doubt we are on the right track, and there is hope that also
the general collecting public, which depends to a large extent on dealers for
additions to the collection, will soon follow, rejecting with disgust the specimens
olfercd with such vague localisation as East Africa, Australia, or Amazons. On
critically examining the geographical distribution of the American Papilios, we
have come across many errors. Localities mentioned in the literature on these
insects which are flagrantly wrong have been put by us between inverted commas
(" ") in the bibliography quoted in this Itevision, and have further been designated
as /a/.vc or loci error. In a few cases, however, where there is a possibility of
the insect being fonnd in the district, we have referred to the record from that
locality as being doubtful or as requiring confirmation.
How erroneous localities get into collections and hence into literature is shown
by the fullnwing instructive instance, where we can trace the error (o its source.
.Streeker figured erroneously as I'lipilio astcrioides Reak. a specimen of VitpHio
( 417 )
■polyxenes americxs Koll., giving Costa Rica as loealit}'. An itinerant German
dealer, Heyne, sold as P. astivioidrn specimens lie had received from Messrs.
Standinger and Bang-Haas without locality labels. Heyne, following a custom
of collectors and dealers, ticketed his specimens Costa Rica, the locality whence
astcrioidcs had been recorded by Strecker (such specimens with a label in
Heyne's handwriting are iu the Tring Museum). Eimer described and figured as
P. asterioides similar specimens, giving as locality Mexico, these specimens
having been received either from Heyne or direct from Standinger and Bang-Haas.
Now, all these individuals are neither the asterioidcs of Reakirt nor of Strecker,
but are nnmistakably Cuban specimens of P. pob/xeni's. The locality Costa
Rica and Mexico for them is "manufactured." We add that the error was not
made by Messrs. Standinger and Bang-Haas ; however, that firm is much to
blame in selling their specimens without locality labels on the pins.
Fabrication of localities is even nowadays going on. Wc mention the following
instance as a warning to other lejiidopterists. Two specimens of what was said
to be PajiUio oregonia were offered to us by an American dealer, who had received
them from a correspondent who apparently loves the dollar more than his honour.
On receipt of the specimens we found them to be Old World P. machaon, one being
a Sikkim individual, the other a British specimen ! The specimens are labelled
" Plumas ( 'o., California, May '03." AVe are keeping them as a memento, an
additional label giving the necessary explanation.
With the publication of (Jramer's Papilloits E.roti'pies the number of known
forms of exotic Lepidoptera increased enormously.* Though the descriptions are
useless and the figures not always good, very few of the numerous new Papilios
are not sufficiently well represented for identification. Only one of Cramer's
American Pajiilios has remained doubtful to ns. This is P. eiiristevs, the figure
of which does not agree with any specimen which we have seen. The fignre is
coarse, and may be erroneous, but it is also possible that the insect has not been
rediscovered. Some of Cramer's American Papilios are very rare in collections,
his specimens being mostly from Surinam, whence no extensive collections of
butterflies have been seut in recent years. Collecting in the French and Dutch
Guianas is difficult, we hear, owing to the extensiveTbrests without roads and
the impenetrable swamps, the only means of exploring the interior being by
means of canoes. Moreover, the butterflies, some species excepted, do not appear
in such great numbers of individuals as elsewhere, having to be searched for.
Jablonsky's Natursystem aller Tnsehten, continued by Herbst, introduces in
the volumes devoted to Lepidoptera some original matter and many copies from
Cramer, and contains also several artefacts. The figure which Esper named
later P. pclcides, and usually considered fictitious, may have been taken from
an actual specimen ; but nothing of the kind exists now iu collections.
Far more critical than any of his predecessors and contemporaries was Esper,
in whose work Die Ausliindiscl/rm Sc/imetterl/»f/e (l~84 — 1801) all the species of
Papilio which he considered distinct are dej)icted. The dates of issue of the
work are given by Anrivillius in 1882. t The lengthy text accomjiauying the
jilates is difficult to understand for a non-German, being full of provincialisms
and now antiqnated words, the meaning of which the foreigner does not find
* The copy in the Tring ^Inscum contains the original covers bearing the years of issue. We obtained
this valuable copy from the library of the late J. H. Leech.
\__Heccimo Critica, in K. Sv. Vet. Alt. Ilandl. xis. 5. p. 182 (1882).
f 418 )
ia ordinary dictionaries. There are many remarks wliich bave qnite a modern
flavonr.
The name aeneides on Esper's Plate 15 raises a point in uomenchxtnre which
we think it is necessary to discnss here, since our decision with regard to this
name is opposed to that arrived at by most other authors. Esper described under
the name of P. aeneas Linne two males of two different species, believing these
males to be S and ? of uciicns. These two specimens are figured on Plate 1">.
Though the figures are referred to as acneus everywhere iu the test, they stand
as aenifh's on the plate, a name which is nowhere mentioned in the text. The
name is doubtless due to a mistake on the part of the engraver. Bock. However
that may be, is the name aeneides to be employed for the one of the two species
called aeneas in the text which had no name at that time ? The facts jiut in a
formula are these :
■n T-i 7.nL I. T, -y-ii f P. species indcuominata c?.
P. aeneas Esper 6 ? text = P aeneides m tab. = -J „ -. . , »
'^ I P aeneas L\un4 c?.
In our opinion a new name proposed for a composite sjjecies sinks as a
synonym if a component part of this composite species had already a valid name.
In this instance, be the name aeneides a mere lapsus of the engraver of the plate,
or a name intentionally given by Esper himself, there was no justification whatever
for a new name, since the supposed female of what Esper considered to be a sjiecies
had the valid name aeneas. Similarly Swainson renamed Linne's Papilio
protesilaus, calling it Protesilaiis leilas. The description and figure given by
Swainson are, however, those of the Brazilian insect, not the Surinam form of
Linnii's sjiecies. Therefore what Swainson considered to be one species leilus
consists of two forms, of which one had already the valid name protesilaus, leilus
sinking consequently as a synonym of the latter. In general terms, if an author
wishes to deal with A and B (individaals, varieties, species, genera, families) under
one name, a new name is valid only if neither A nor B has already a valid name.*
Two (if Esper's Papilios have been said to be antedated by names given
in Martyn, Psyche. We agree now with Mr. Sherborn f that J'syelie should be
treated as non-published. There is one copy and portions of two others in the
Tring Museum, some jilates being numbered and others not. The plates have
the appearance of being nothing but printer's proofs. However that may be, in
the case of. the two American I'apilios {aristodemus and hectorides) Martyn has no
priority over Esper, the latter having published a description and figure three
years previous to Martyn in the Magazin der Neuesten Auslandischen Insecteii
(1794), a rare work which has been overlooked by recent authors. Esper himself
quotes the Magazin, and it has also been mentioned by Donovan iu Naturalist's
Repository, Ent. ii., text for plate 177 (1S27). It has not been consulted by
Sherborn.
In 179" there ajipeared what is perhaps the best lepidopterological work of
the eighteenth century. The Natural History oj the Rarer Lepidopterous Insects
of Georgia, by Abbot & Smith, deviates entirely from the other iconographies
above referred to in illustrating the life history of the insects observed. Apart
from the pictures in Merian's Inseeta Sari name nsia, often fanciful and grossly
incorrect, and Seba's worthless drawings of caterpillars and pupae in the Thesaurus,
Stoll's Supplement to Cramer, Papillons Exotiques, was practically the only work
• .See also Nov. Zool. i.t. Svppl. p, xxiv. (1903).
t Imtce Anmulhtm (lOOli).
( 419 )
containing illustrations of the earl}' stages of exotic Lepidoptera. Tlie Natural
ffistori/ of Georgia meant an enormous advance in this direction.
The series of le])idopterological works of the nineteenth centnr)', as far as
thej' concern us here, opened with Hubner's Sai/iml/nu/ Exotincher Sc/imcffl/t/ffr,
appearing from 180G onwards. As a collection of fine illustrations the S((mmlii)ii]
was a great success, while as a scientific work it was an entire failure. With
the exception of the separate volume, entitled Zutriige, there is hardly any text
accomjiiinying the plates. Sherborn * says that he does not recognise as valid
the names appearing on plates without te.\t. We wonder if he will be bold
enough to reject the majority of the new names of the Sammlung. The plates
not being numbered, and no date of publication f being given, the work is a
great trouble in compiling the bibliography of the insects figured. As every
little contribution towards fi.\ing the years of publication of the plates is of some
value, we draw attention to tlie fact that Hiibner figures on Plates 114 and 115
of the second volume the identical insects which Godart described in 1819 as
? and doubtful c? of his Papilio ]irotoi!amas, which name Hiibner emjdoyed for
the male. This can hardly be a coincidence. Hiibner doubtless knew of Godarfs
publication when he engraved the two plates — i.e. the plates appeareil after 1819.
Hiibner's Verzeichniss hekanntev Schim'ttlirige was published from 1810 to
1827 or 1828, the Papilios appearing about 1818. A few new names are proposed
for American Pajiilios in this much-abused work, which represents the first attempt
at classification uf all the known Lepidoptera.
If Hiibner erred much on the side of descriptions, Godart's treatment of the
Butterflies in the Encgdopddie Mi'thodiqui' ix. (1810) suffered from the entire lack
of figures. The work is purely descriptive, but the descriptions are admirable
for that time. It is a most careful piece of work. There are naturally a number
of mistakes, especially in the mating of the sexes of dimorphic species, which is
excusable, since hardly any reliable observations on di- or polymorphism among
exotic Lepidoptera were known. Godart's species were mostly described from
specimens contained in the Paris Museum. Very few authentic specimens are
preserved ; the majority appear to have become destroyed already during the
first half of the last century. A few of the specimens are in the Royal
Scottish Museum {P. serville, triopas, imerius). While previous authors, with
some exceptions, employed generally the Linnean formula of nomenclature for
Butterflies, interposing between the generic title rapilio and the specific name
either the sectional name Eques or the subsectional name Trojanus (or Troes),
respectively Achivus, or both these names {Eques Trojanus and Eques Aehivus),
Godart adhered to a pure binominal system of nomenclature. In the Supplement
to his work, issued in 1824, several Papilios appear with modern personal names
standing in the nominative form : Papilio serville, P. devilliers, etc. This kind
of name for species had become the fashion in Prance. We do not consider it
advisable to alter snch names into servillei, denlliersi, etc., us has been done
by most subsequent authors, servillei being no more Latin than is serville.
The custom of naming species after persons, which is nowadays almost a
mania, hails from Liune himself, who named insects Scldijferi, Bergmanniana,
Listerellu, etc.
* Index Animaliiim ji. vii (1S)02).
t Tlie new eilition o£ the Sammliin// which is now being issued by P. Wytsman is acrompaniod by a
text written by W. F. Kiiby. Here, again, no dale ot publication of this text is given 1 1
( 420 )
Many of the entomological works of Godart's period have a Fronoh nonicn-
clatnre, which cannot he considered valid. Rogers's names for Papilios pul)lished
in 1 ^^20 cannot stand, the names given being I'upiUon bias, Papillon piritlious, etc.
A few of the specimens described by Godart were subsequently fignrcd
by Lneas in his Lrpuloptcres Exo/i(/>irs (183.5), which appeared shortly before
Boisdnval's Species Gnu'ral i/es Lepidopteres i. (ls:iri). This Sjieeies (Inund
is a fundamental work for the study of Papilios. Many errors are cleared up,
and also new ones are made. The original specimens from which the new species
were described are nearly all in the collection of Monsienr (!harles Oberthiir. A
few appear to have been replaced by Boisduval by specimens which do not agree
with the descriptions. Snch a result of the habit of removing from the collection
the original specimens in favour of better-preserved individuals is very instructive,
confirming our contention (see p. 414) that the type-sj)eeimen of a new name should
be marked as snch, and be carefully jn-eserved. Barring accidents, the revision
of the nomenclature of a family is made comparatively easy by that means, and,
what is more important, the results are more reliable, and hence the nomenclature
rendered more stable than if a reviser has only the descriptions and figures to
go by. The nomenclatorial type, or " name-type " for short, is of no other
imjiortance. That should be clearly understood. Nomenclature is an extraneous
matter. It is not the natural history, bnt only a convenient method of recording
some of the results of descriptive science. The natural history types of one and
the same species or form are manifold. One niay call an individual a type, if it
represents the average. Since the same individual is not the average in all
characters, a species or form has many average-types, one individual being a
morphological type in one detail of structure or pattern, other individuals in
other details. There are, further, two kinds of jihylogenetic types. Sjiecimens
which are the most generalised in a certain character may be called ancestral
types in respect to this character, others being ancestral types in other organs.
Some individuals are more advanced in certain characters than other specimens,
and therefore represent a more advanced type, other individuals being more
advanced in other details. Since the variation of the various organs is to a
large extent independent — i.e. since retrogressive or progressive development docs
not take place in the same degree in the various organs — an individual may be
an average type in one organ, an advanced or an ancestral type in another, and
not typical in a third organ. To these morjiho- and phylotypes may be added
bionomical types, habits being also variable within a species ; and so on. It is
obvions that all these types have nothing to do with the name-typo.
Besides Drury's llli(strations of Exotic Insects (1770-82) very little of
importance on American Pajiilios was published in England during the eighteeiitii
century and the first four decades of tlie nineteenth. The Zoologicnl llliist rations
by Swainson and Donovan's Natui-alist's Repositonj * were the only works
which contain more than an occasional reference to exotic Papilios. This
became entirely altered iu the forties. With Doubleday's List of the Specimens
of Lepidopterous Insects in the Collection of the British Museum (1845)
commenced a series of catalogues which, though in the first instance meant to
be a list of the contents of the British Museum collection, became synonymic
catalogues of all tlie species and varieties described. Doubleday's List of 184r)-48
was foHowed by his Genera of Diurnal Lepiiloptcra (]84(i-5~!), by Gray's Catalo(jue
* £!ach plate of the Ih-ptrntttrtj bears the date of publication,
( 421 )
of Li'pulopfp.rou^ Tiisecfs (1852), and Gray's IJst of Lepidopferous Fnsccts (1850),
both works of Graj' dealing only with the Papjiliomdac.
The Catalogue of Gray is for onr subject the most important of these
jiublications, since it contains descriptions and figures of numerous new " species."
The work is faulty in many respects. The iigures are in some instances mislcailins;
on account of errors in coloration, the sexes are often wrongly mated, and the
localities are not always reliable.
In these English catalogues we find for the first time nomina nuda of American
Papilios, names without any attempt at description, the author of such names
considering it apparently sufficient publicity if the name was in a museum behind
the specimen. This bad example has fortunately not been followed in the case of
American Papilios by later authors, e.Kcepting certain Continental dealer-authors,
who did it for the sake of earning an additional shilling, the purchasers of
specimens paying more for a supply of named specimens than for unnamed ones.
Such an abuse of nomenclature cannot be too strongly condemned.
During the fifth and sixth decades quite a number of new American Papilios
were described by Kollar, Lucas, Menetries and others. Among the Lepidoptera
recorded from Culia by Lucas, in Sayra, Ilistoria Fisica vii. (1857), there are
several that do not occur on the island. The species published by Lucas in the
Revue de, Zoologie for 1852 were mostly based on specimens lent to him by
Boisduval, and are now contained in the Oberthiir collection.
Kollar's species were collected by Prince Sulkowsky. In dealing with the list
of captures published in 1850 one must bear in mind that Sulkowsky travelled up
the Magdalena River, and crossing the Cordillera of Bogota went down the Rio Meta
and Orinoco. The localities of the specimens have not in every case been carefully
kept, some specimens said to be found near Angostura (Ciudad Bolivar, on the
Orinoco) being doubtless of Colombian origin.
So far very little was known of the habits of the butterflies of tropical America.
The study of exotic insects was almost purely museological. The more valuable
were the essays on the Amazonian fauna published by Wallace and Bates. The
essay on the Paj)ilios of the Amazons by Bates (in Trans. Ent. Sac. Land. 1861) is
a classical work. It is full of notes on habits, distributions, and on geographical
and individual variation which were quite new at that time. Nothing equalling
this essay has since appeared on tropical American butterflies. However, in his
references to non-Amazonian forms Bates was very often at fault, nor have all his
conclusions as to variability and distribution in the Amazon valley been borne out
by recent explorations and researches. It is a great pity that the material collected
by Bates has been scattered over many collections. Variation cannot be studied
withont comparison of long series of specimens. The problems of the distribution
of the various geographical varieties in the Amazonian fauna touched upon by
Bates will remain open till adequate material (properly labelled) is available.
Everybody has Amazonian Papilios, but nobody has long series from a suflicientlv
large number of localities. The large collections made by Dr. Hahnel — the best
collector of bultorflics who ever visited South America — are also scattered.
Shortly after Bates's essay there appeared a work of quite a different character,
but of no less importance. The Species Lepidopterorum huiusqiie descriptae vel
iconi/jus erpressae, by C. &. R. Feldrr, contains, like Gray's Catalotjiie and List,
only PAriLioNiHAK. It is the first classification and revision of the entire family
which might be called thorough. The species are mostly grouped correctly, with
(422 )
the exception of some mimetic forms, which are classified with their models.
In charactei'isitig the groniis of species the Folders laid great stress upon small
differences of neuration, in which they were wrong. The series of specimens com-
pared by the authors was generally small, and therefore many small ditferences
appeared to them to be constant, while they are in fact individually variable. The
same applies to difTerences in size, wing-shape, and pattern, which served tlie Felders
as specific distinctions. The numoions " species " of Papilio described by them in
the essay mentioned, in the Rcise (lev Fregatte Norara, and elsewhere, are mostly
varietal forms, often mere individuals deviating in some character from the average.
In many cases the authors were well aware that the new " species " of Papilio was
a geographical form ; in fact, they make sometimes a statement to that effect.
However, they treated nomenclatorially all such forms as species. This curious
phenomenon is by no means uncommon among systematists of our day. To
consider all geographical representative forms as distinct species, and treat them as
Buch nomenclatorially, is quite consistent. To regard all individual, seasonal and geo-
graphical varieties of a species as not necessary to be distinguished by special names
and hence deal with all of them under the one specific name, is again methodical.
But to call a form a local race, naming it Papilio paudion, and to employ the same
formula Papilio uuchisiades for the species, is certainly ill-considered. The formula
expressing a conception should show which special conception is meant, whether
a family, a subfamily, a genus, a species, a geographical form or another kind of
variety. It will doubtless take a long time before all systematists have learned to
employ a nomenclature which is consistent witli their own ideas. The more is it
necessary to bring the matter again and again before their mind.
The Felderian types are mostly in the Tring Museum, some being in the
Hofmuseum at Vienna.
In the second half of the last century the number of works and treatises
dealing with American Lepidoptera increased very rapidly. In North America
especially entomologists became very active in the si.xties, and have ever remaiiied
so, being now in some branches well in advance of European systematists. We
mention here only those authors whose work is of special importauoe for our
Revision.
The species of Papilio described by Hopfier in 18G6 have partly been over-
looked subsequently. We hope that wo shall be found to have identified them
correctly. Kirby's Catalofiue of Diurnal Lepidoptera (1871 ; Supplement, isTT)
is too well known to require more than passing mention. The nomenclature
of varieties needs j)urifyiug. The formula " Papilio eurimedes Cram. var. a.
P. agathocles Koll. " for a variety is very cumbersome. The work has been moat
useful to us, in spite of the errors in synonymy, which are hardly avoidable
in a compilation of this kind. A second edition of the catalogue is a great
desideratum.
The Lepidoptera of the Argentine Republic are dealt with by Burmeister in
his Description Physique de la Pcpiihlique Argentine, vol. v. (1ST8) and Atlas
(1879). The figures of the early stages, though some are not correctly identified,
are very useful.
Another faunistic work of the same time is Gnndlach's Contribucion a, la
Entomologia C'uhana (1881). It is a descriptive catalogue in which former work
on Cuban Lepidojjtera by Pooy, Lncas, II(!rricli-Schiifl'er and others is revised, notes
on the life-history of many species being added. Since Gundlach had resided so
(423)
long on Cuba, nobody was better fitted to write on the subject. One of the prettiest
American Papilios bears his name {P. gundlachianm). Unfortunately we are
bound to employ for it the older name columhus, given to it previously by Gundlach
ill lift, and published by Herrich-Schiiffer. The insect was renamed gimdlncJnanua
by Felder, because the name colwmbus had already been employed for another species
in the genus Papilio. Our reasons for retaining nevertheless the name cohimbiis for
tlie Cuban species are of a general nature.
We distinguish between the name of a form (species or variety) and the
nomenclatorial formula in which this name appears. Papilio marccllus f. telamonides
is the formula for the summer variety of the North American species Papilio
■marcellus, the name of this variety being telamonides. Some authors consider it
permissible, or even advisable, to replace the name by a new one, if the supposed
variety tnrns out to be a distinct species. Among prominent Lepidopterists this
view was held by Niceville. If the principle were correct, one ought consequently
to replace a name by another also in the case of supposed species being proved
to be varieties, or a supposed geographical form to be seasonal, etc. We regard
this opinion as utterly opposed to stability in nomenclature. The first name given
to a form, if not preoccupied, is the name of this form, whatever position the name
takes in the nomenclatorial formula according to the individual opinion of an author.
Now, systematists differ as to when a name is preoccupied. Leaving aside the view
that a name is preoccupied by another which has the same root or the same meaning
{_fluciatilis, Jiiuiorum ; alhoniger, leucomelas), there are two opposite opinions
with regard to this question. Most systematists adhere to the rule that a name
is preoccupied if at the time of its publication there was already a name identical
in spelling in the genus where the new form was j^laced by the author. This rule
would be excellent, and we should adhere to it, if there were not serious drawliacks.
Opinion as to the extent of each genus is not at all unanimous, and never will be.
According to one author a certain genus contains many species ; according to another
writer only one or a few belong to it. Hence it is often impossible to decide, if at
the time of publication of a new form the name was or was not contained in the
respective genus as conceived by the author of the new form. For instance, if
somebody published to-day a new Swallow-tail, calling it Papilio ulysses, who will
decide if this name is preoccupied, since some authors restrict the generic title to
machaon and allies, others to priamtis and allies, others again to some Nymphalids,
while the majority of Lepidopterists comprise in Papilio many hundreds of
differently organised Papilionidae ? To follow the above rule consistently is the
more difficult in the case of Lepidoptera, as the genera were formerly so very
extensive that the species belonging at one time to a single genus are now
often distributed over several families. There is, moreover, the great difliculty
in Lepidoptera that one would have to decide whether Papilio E<jues Trojaims
dan/anus Fabr. (1793) is preoccupied hj Papilio Eques Achimis dardanus Brown
(1775) or by Papilio Damnis Festivits dardanus Cram. (1775); and whether Papilio
Eqiies Achims orestes Fabr. (1703) is superseded by Papilio (orestes) Meerb. (1776).
The above rule further compels ns to ask, if Papilio harmodius Donbl. (1845)
which is a swalluw-tail, is preoccupied by Papilio Ni/mplialis Fcsticus harmodius
Cram. (1779), which is a moth, or the name of the swallow-tail Papilio hesperas
Westw. (1843) by Papilio Petttieits Jiesperus Fabr. (1793), which is a Nymphalid ?
Now, there is no difficulty about these questions, and there can be no ditt'ereuce
of opinion, if one accej)ts our principle of dealing with such names. We consider
( 424 )
a name preoccupied on]}' if there is au identical older name iu the genus to which
the sjiecies or variety noio belongs, it being qnite irrelevant whether the name was
or was not preoccupied iu the genus where the author originally placed the I'orm.
rapilio bruttia Fabr. (17U3), which is a swallow-tail, does not sink as a synonym
on account of rapilio brutiis Cram. (1775), which is a Nymphalid. As in the case
of the Revision of the Fapilios of the Eastern Hemisphere (1895), we revert also
in the present Revision to the first name of each form, though this name may recur
in this so-called genus Papilio. In our proposed generic revision of the Papilionidac
the forms which appear homonymous in the present work will come under diilerent
generic titles. The number of such names is very small, and it is certainly advisable
to bring already here the names of the forms in accord with the names they will
bear in the generic revision of the family.
The list of Fapilios iu his collection published by Charles Oberthiir in l8Si)
contains many useful hints, besides a number of fine figures, and descriptions of
new forms. The collection was very small at that time as compared with what
it contains now.
Aurivillius, in 1882, gave a revision of the Lepidoptera described by Linne
in Museum Ludocicae Ulricac (1764). Though there are some points which the
author conld not satisfactorily decide for want of adequate material, the essay is
an examj)le of very painstaking work, nothing being taken for granted and every
question carefully investigated. It is a Recisio Criticn in the true sense.
The only popular work on exotic Butterflies which it is necessary to mention
here is Staudinger's Exotische Tag/alter. The book, which was destined for the
great mass of " collectors " of butterflies, was not meant to be a critical entomo-
logical work. But, in spite of numerous errors in identification, it was also from
a scientific point of view a welcome contribution towards the knowledge of tropical
butterflies. Here, and in some other i)laces, notably in the Iris, Standinger
described (piite a number of new species and viu'ieties of American Fapilios, among
which hahiidiy quadratus, tasio and (jaiicppi are the most noteworthy. We need
not dwell here on Staudinger's enormous influence on Falaearctic Lejiidopterology,
which was his chief interest ; but it is only fair to mention that no other ento-
mologist has had so great an influence on the exploration of the South American
butterfly fauna as Dr. Standinger. A great many collectors and residents were
encouraged and subsidised by him, among whom Dr. Hahuel and the Garlepps were
tiie most successful. A large proportion of the American Fapilios which one sees
in collections are Standingerian specimens. In systematics Standinger was guided
more by the general appearance of the species, or by intuition, if we may say so,
than by hard facts of morphology, and therefore was often led astray. However,
ho was far too keen an observer not to recognise some general truths in respect
to relationship. He was the first to see that there is a diflcreuce between
geographical and non-geographical varieties, and he endeavoured to distinguish
even nomenclatorially between these two grades of varieties, calling the geographical
variety va/ietas (var.) and the non-geographical variety abeinUio (ab.) The
distinction remained, however, more or less theoretical, Lepidopterists emjjloying
car. and ab. just as iu<liscriminately as before. This is one of the reasons why
we reject var. altogether as a special nomenclatorial term. In another matter we
have followed Standinger now for some years. In the Revision of the Fapilios
of the Eastern Hemisphere we altered, following precedent, all the adjective names
of species and varieties into the masculine gender, I'apilio being masculine. We
( 423 )
have siuce come to accept Standinger's view that every name should Ije treated
as a nouu, and therefore its gender be independent of that of the generic name.
For ns Fapilio oregoiiia is as correct as F. opalinus.
StandiDger was an ardent adherent of the habit of writing all names of
Lepidoptera with a capital. One of his argnments for the correctness of this purely
lepidopterological custom — in no other branch of Zoology have all the names ever
been written with capital initials — was that Liun6 had employed capitals for all
names of butterflies. In this Standinger was wrong, Linne having written with a
small initial the few adjective names bestowed on butterflies (jlissimilis, assimilis).
We consider the writing of all specific and varietal names with small initials, and
of generic names with capitals, as by far the most convenient method, generic and
non-generic titles being at once recognisable as such. In contradistinction to the
habit of capitalising all names (Paj>ilio Priamus), there was early in the nineteenth
century the method in vogne, especially among French authors, of writing both the
generic and specific names with small initials — papilio priamtis Such matters are
purely conventional. One ought to select that method which is the least confusing.
Among the literature on Nearctic Lepidoptera no works are so prominent as the
Butter/lies of North Ameiica,hj'\N . H. Edwards (1808-97), and Scuddcr's BKttcrflies
of the Eastern United States and Canada (1880). The plates issued by Edwards
are nearly all of a quality hardly ever reached on this side of the Atlantic, nor have
we any work iu which the life history of the butterflies is so well illustrated.
His greatest discovery among Papilios was the demonstration by breeding of the
polymorphism of I'apilio hairdi and of the seasonal variation of F. mancellus.
Though in other places Edwards rather ridicules the idea of frequent occurrence of
hybrids in nature, he explains nevertheless this polymorphism of F. bairdi by
assuuiing that the insect is a product of hybridisation.
iScudder's Butterjiies is the most intrinsic work written on Diurnals. No other
work on Butterflies can be comjjared with it. The mass of morphological detail
which was new is enormous, and, what is more, the facts were well digested, and
not merely comjjiled and put together anyhow. But it was perhaps just this
abundance of small characters which obscured the great distinctions in Papilio so
much that Scudder did not clearly perceive the three main divisions of this so-called
genus. It was leil to Erich Haase to rediscover the three natural sections into
which the Papilios of all regions are separated.
l]i his Untcrstwhuiiijcn iiber Mimicnj (1803) Haase gives a classification of the
Papilios which is in the main quite correct, starting from the three main divisions
which Horsiield had defined in 1857. Many obscure points in relationship which
had defied every other author were successfully solved. He was the first and has
remained the only author who saw the close connection that exists between Papilio
ariarathes, harmodius, eiinjlcon, etc., on the one hand, and Papilio protcsilaus and
allies on the other. Those mimetic Papilios are placed everywhere in books and
collections with P. unchisiades, or even P. acneas, instead of with P. protesilaus,
marcellus, etc. As a student of Mimicry Haase was aware that models and mimics
are usually not nearly related, and this general knowledge may have guided him iu
the right direction.
The morphological distinctions advanced by Haase for the three main divisions
oi Papilio are only sliglit, and do not ai)i)ly to all the species. We have endeavoured
to give the classification a better morphological basis. The only serious mistake
which Haase made in rusjject to American Papilios was the position he assigned to
( 42fi ;
Papilio hellanichus, placing this insect with P. machaon, as all otiier authors • had
done, instead of close to P. scaniander.
In Scndder's work, mentioned above, the genitalia of both sexes have to a
certain extent been taken into consideration in distinguishing the species. Messrs.
Godman and Salvin followed that line of research, at least with the S genitalia,
details of which are figured of nearly all the Central American Papilios in their
fiimous work, the Diologia Ccntrali-Amtricaiia. The female genitalia were only
referred to cnrsorily in a few instances, inclusive of the bursa copulatrix, which was
often fonnd to be different in different species. It was the first fannistic work on
tropical insects in which such researches were carried out. These reseaiches have
greatly advanced our comprehension of the true taxonomical value of these organs.
The iiuthdrs fonnd the genitalia to be excellent guides, in many instances the sole
trustworthy guides ; but they recognised also that there are groujis in which the
genitalia do not present any tangible differences between the species, while in other
instances the organs were observed to be variable. We commenced to work at this
problem when the Revision of the Papilios of the Eastern Hemisphere was in
preparation (1894). The main results, which we have since repeatedly verified in
many groups of Lepidoptera, were published in 1896. t They may be epitomised
as follows :
(1) The majority of species are different in the genitalia of both sexes, a small
percentage only showing no distinctions in these organs. Among American Pajjilios
P. ariarathes and allies cannot be separated by these organs with certainty, and
P. bairdi and pohjxfncs appear to be identical in the genitalia.
(2) About half the number of geographical forms are more or less distinctly
different in the genitalia, at least in the males, the differences being often found to
be entirely bridged over by intergradations.
(3) There is always a certain amount of individual variability in the genitalia.
Specimens abnormal in these organs also occur. It requires, therefore, often a
series of dissections to arrive at a correct estimate of the distinctive characters
presented by the genitalia. There is no individual dimorphism in these organs
connected with the dimorphism in jiattern or colour. The only seasonal dimorphism
we have come across, though a great number of seasonally dimorjihic species liave
been examined, is fonnd in P. xuthus, the spring specimens (from hibernated pupae)
differing slightly and not quite constantly in the harjie from the summer sj)ecimens.
No such difference obtains in the seasonally dimorphic P. marcelltis.
It will be observed that the differences in the genitalia, though generally less
variable than those of colour and pattern, and often much more striking than
wing-difl'erences, require in each case the same careful investigation as colour and
pattern, before their true taxonomic value can be pronounced upon. A difference
in the genitalia may be specific or varietal, just like wing-differences, t
A number of American Papilios have been dealt with by Eimer in his
treatises on ArthiUlKiKj und VencamUschift bei Schmctterliiigeii (1889 and IS'.to).
These essays are of a philosophical nature. As contributions towards Papilio-
systematics they are a failure, the researches uot having been intrinsic enough to
* Burmeistcr, in 1S7S, pliiccd hdlaitichKS iic;ir scamandvr. II uppear.s that be a.ssigned the right
pusitiun to it only bceausc lie identified it erroneously with clcotM Gray (1832), wbieh was known to be
a near relative tif icamttiuh'r,
t " Jlechanical Selection," in Xov. Zmil. iii. pp. 420 — 525 (IsutJ).
X r. Born lias arrived at similar results from a study of the forceps of Cartihui (see Jiuckicit-lior;c
xvii. (1900) and «.).
(427 )
Warrant the majority of the conclusions. The essays suffer also greatly from
verbosity. Nevertheless we may say (with some poet) that mistakes are often more
instructive than facts. The name mediocauda introduced by Eimer for a specimen
of F. polyxenes without locality has been overlooked by subsequent authors, not
being mentioned in the catalogues of Nearctic Lejiidoptera. The specimen without
home might better have been left also without a name.
It has been the object of the present Revision to correct to the best of our
ability the mistakes contained in the literature on the American Papilios, and to
broaden the morphological basis of the systematics of these insects. We have
widened the scope of research as far as the material permitted, and therelbre we
have arrived in many cases at a clearer insight into the relationship of the Papilios
with one another than if we had followed the customary methods of investigation.
We hope to be pardoned for not having solved every knotty jioiut.
The most interesting general result of our researches is perhaps the demonstra-
tion of geographical variability in secondary sexual characters apart from the
genitalia. The occurrence of such variability is of great bearing on systematics,
since many authors consider secondary se.xual differences to be of generic value.
The remarkable difference obtaining in the scent-scales of some species which are
otherwise very similar (see P. prolesilms and allies) is also noteworthy. Since we
shall have to discuss the general questions as to distribution, relationship and
evolution of the Pupilionidae in our proposed essay on the classification of the
family, we abstain here from entering on such problems. For the same reason
we have restricted the illustrations of details of structure as far as it was possible
without serious injury to the lucidity of the descriptions. We have further
abstained from describing in detail the various known species and varieties, but
have given at least some principal feature of each form in order to enable the
reader who is not in possession of the literature quoted to determine his specimens
from this Revision. The keys to the groups and species will, we venture to hope,
also be found useful. The groujis of species have been defined only from the
American species which they contain. These groups are not all of generic value.
We hope nobody will find it necessary to propose generic names for them. The
e.xtent of some of the groups maybe considerably altered in our proposed generic
revision by the inclusion of Old- World forms. Besides, Hiibner and Moore have
already supjilied a great number of generic names for Papilios, the diagnoses given
with the names being of the most superficial kind, and those of Moore, moreover,
often very faulty. It is common knowledge that the delimitation of genera in
cosmopolitan families cannot well be based on the limited nnmber of species
occurring in a single faunistic district. Classification has always suffered from
the habit of systematists of studying the systematics of a district rather than
concentrating their labours on certain families, taking into account all the species
of the globe.
The treatment of the matter embodied in this Revision requires a few more
remarks. Wc are in favour of simjililicatiou of nomenclature. Every simplificatiou
which is consistent with the object of nomenclature should be welcome to the
systematists, whose labour suffers from unnecessary uomeuclatorial complications.
One such simplification is to write in the te.xt, headings, and in the synonymy all
specific and varietal names with small initials, and the names of higher classificatory
categories (subgenus, genus, subfamily, etc.) with capitals, no matter whether they
were thus written or not by the authors quoted. We consider it utterly indifferent,
28
( ^28 )
whether Lncas wrote Papilio orbignijanus or papilio orhigmjanus or Papilio
Orbignyanus. Snch ontside matter does not in the least affect the natural history
of the insect thus designated. Papilio oi-ljigivjanus is the most convenient form of
spelling, and is therefore here adopted in every case. We have also simplified, as in
former essays, the spelling of dedication-names standing in the genitive form ending
in i. The authors of such names are very inconsistent in the spelling of the names.
We find birchdllii and dunali, blumci and latrcillii, tcallacei and tcallacii, buinlii and
brucii, lorquiiti and lorquinii, kirhii and kirbi/i, etc. One cannot possibh' remember
what in each case the original spelling of such a name was. If one has to write
the name, one has to look up the original description. But is it really necessary
to stick to this inconsistency and this burdensome variety in spelling ? We think
not, since uniformity can be arrived at without the slightest difficulty. However,
what is an unnecessary burden in nomenclature should certainly be dropped. We
write these dedication-names with one i added to the name of the person, wallacii,
WaUarii, icallacci and Wallacci lieing all reduced to one form, ivallacri. That
snch a simplifying principle is really opportune is best shown by the fact that, in
consequence of the geueral arbitrariness in s]>elling the ending of dedication-names,
the name of a new form spelt dnicei will invariably be written by some later author
drucii ; or if the name was written originally drucii, the spelling drucei will surely
also crop up. As we treat names like androgeos and androgeus, pobjdnmns and
polidamas as being different, brucei and brucii, or icestwoodi and tcesttcoodii, or
Idrbii and kirhji would be rather embarrassing without the foregoing principle of
simijlification which renders such names uniform. For similar reasons the German
a, (V, and il (which were originally ae = a, oe, tic,) have been changed into ae, or, and
tee in all names.
About the naming of forms below species there are many different opinions.
All agree that what an author considers to be a " species " should bear a name.
But one may very well ask, is it necessary to give names also to the various categories
of varieties ? The answer depends on what is the object of naming. Liune invented
his binominal formula for the species with the purpose of reducing chaos to order.
However, if we agree that for the sake of lucidity in studying the species of each
genus it is necessary to have a special name for each species contained therein, it
follows that in researches on the varieties which compose each species names are
likewise required for these varieties. Now, is the study of these varieties essential
enough for systematics to warrant the introduction of names for the enormous host
of varieties ? With many authors systematics have been and are essentially a
description of the differences of " species." The knowledge of these differences
is certainly in each case essential ; one cannot do without it. But it is not the
final aim of systematics as part of the science of life. A collector learns to
know the various "species" by handling them, just as a child learns to know a
language by practice. AV'lien once a candidate who sjiuke and wrote French and
English fluently, having resided in these countries for a number of years, presented
himself for examination pro facultate docendi at a German University, the professor
of modern languages gave him the advice to become foreign correspondent in a
mercantile house, since he had no philological kiidwledgc at all. And similarly
a professor of zoology once said to a candidate for the degree of rii.IJ. who could
and did boast of knowing by heart practically all the vertebrates and a large
proportion of the invertebrates of (.entral Europe : " That is very good ; if you now
study Zoology for a couple of years I shall be pleased to accept you as a candidate."
( 429 )
Both profussors were perhaps a little sarcastic. Nevertheless there is a good deal
of truth in what they said. The describing and cataloguing of " species " are
certainly the basis of systematics, but also the lowest degree in this science. After
that comes classification, or in other words, research in relationship. To have a
sound basis in this research one has to start from the individuals wliich are blood-
related, and work upwards to the species. The individuals composing a species
have each some peculiarity. This individual variability, however, is not everywhere
indiscriminate. The individuals are in many instances found to fall into different
groups characterised by some corporeal distinction. These are the varieties of
which a species is composed. If the species are the product of evolution, the
commencement of the splitting up of one species into more must be found among
the varieties. The study of the varieties is, therefore, a study of the origin oi
species, or the relationship of species with one another ; from which follows that
the classification of species according to their relationship depends on the study of
varieties. However, if the study of varieties is essential for the classifier, varieties
have as much a claim to a precise nomenclature as the species. If we speak, for
instance, of Paj/ilio f/ioas from Cuba, /-'. t/ioas from the Gnianas, P. t/ioas from
Brazil, etc., all we gather is that Papilio thoas occurs in these different districts.
On the other hand, if we write of Papilio thoas oviedo from (!uba, P. thoas thoas
from the Gnianas, P. thoas thoantiades from Argentina, etc., we perceive at once
that the species P. thoas has developed into a number of different varieties, and we
are able to discuss these varieties and their bearing on the general questions of
evolution without having constantly to repeat the localities where each variety
occurs, P. thoas cinyras being a decidedly more convenient term than " the variety
of P. thoas from the Upper Amazons, Pern, and Bolivia."
The varieties fall into three categories : the geograi)liical, the seasonal, and
the individual variety ; the last two being the lower grade variety, and the first
the higher grade variety. This distinction between a lower and a higher grade
of varieties has been habitual with most entomologists for over a century. It
was Esper who first made the distinction. He dealt with variability in a far
more philosophic spirit than any contemijorary systematist. In his essay De
varietatibiis (1781) he says, p. 18 :
" In pluribus generibus species iterum subdividi jubet copia et proxima earnm
affinitas. Essentiales quibusdam insunt characteres, diversitatem in ipsa specie
constitnentes, quos in aliis pro accidentibus habere debes. Ulas subspecies, has
meras tarietates appellandas censeo, de tpiibus nunc uberius quid constet est
dicendum.
§ ^iv-
" Subspecies (Untergattungen, Rages *) quae vulgo annumerantur varietatibus,
[plane ab his sunt separandae. Originem e.\ speciebus du.xisse, i)erfectissima in iis
declarat partium essentialium similitudo. . . ."
AV^e have accepted Esper's term subspecies for the essential variety — namely,
that kind of variety which is an incipient species. For an incipient species no
better term could have been coined than subspecies. According to our researches
the incipient species is represented by the geographical race. As no other variety
forms the basis of the development of a species into several species, the term
subspecies is employed by us for nothing else but the geographical variety. Since
* Esper's " Gattung " means, of course, what is now called a species, " was sicb gattet,"
( 430 )
tlic contrast existing between the geograpbical variety (= subspecies) and tlie
u<)n-geograi)liieal variety (= individual and seasonal forms) has been demonstrated
recently in Lejiidoptera in another jilaec,* we mention here merely that the
combination of characters in a subspecies is essentially the same as in a species,
the difference being one of degree more than of kind. An overlapping of characters
often takes jilace in subspecies, showing that these races have not attained to
that kind of complete separation which exists between synpatric species. t
In dealing uomenclatorially with the varieties it appears to us highly
advisable to emphasize also in the nomenclatorial formula the contrast which
exists between the essential variety, or the subsjiccies, and the lower-grade varieties.
This, we think, is best attained by the formula first employed for the geographical
race by another Icpidopterist, Drury, in 1773, and adopted during the last twenty
years by a great many systematists. In this formula the name of the subspecies
follows directly after the name of the species, jnst as this comes immediately
after the name of the genus, no explanatory term, subsp. (= subspecies), or var.
geogr. (= varietas geographica), or anything of that kind being put in between
the specific and snbspecific names : Papilio polyxenes americm meaning Papilio
jiol'/re/ies subsp. (= var. geogr.) amcricus. Entomologists appear to be rather
reluctant to adopt this simple Drnryan formula.
However, we repeat that the main point is not the nomenclatorial formula
by which species and varieties are recorded, but the recognition of the existence
in nature of species contrasting with a higher grade of varfety (subspecies =
geograjihical race), and this contrasting with a lower grade of variety (seasonal and
individual forms). Linne rendered chaos into order ; let it be the duty of tiie
modern systematist to follow him by bringing order into the chaos of varieties.
A geograjihically variable species consists of at least two subspecies. For
instance, the Colombian specimens of I'apiUo backus, which species occurs from
Colombia to Bolivia, are different from the more southern individuals. We have
therefore a northern and a southern subspecies. The opinion still held by many
collectors and describers that the (Colombian form is the "species" (the " Stammart "
of German describers), because Felder gave a name to it some forty years ago,
while the more southern form is the " variety of it," on account of its name being
of a later date, should be abandoned as utterly unscientific. All the subspecies,
inclusive of the first described one, are co-ordinate ; the entire series of (two or
more) subspecies is the species, t
As regards the nomenclature of subs])ecies we have first to repeat that, if the
stability of names is one of the iirincipal aims of nomenclatorial rules, the first name
given to any member of a species must be adopted as the name of the entire species.
For instance, though Linno described the black female of a Nearctic Swallowtail as a
* ''Der Gcgcns.atz zwischon j^cop'apluschcr und nichtgcogr.apliischcr Variation, "in yCfituchr, Wins. Zi>i>l,
Ixxxiii. pp. 151—210 (1!I05).
t Tlic principal criterion of Ibc conception "species" is tliat ."siToeics can exist together without
fusing, no other barrier keeping them apart tlian their own organisation.
t Lorenz, in isy2, called the series of sulispecics constituting a si)ecies the Formciilircis of thi
species. A l''ormvnhrt'is is Llierefore the same as our species. 'I'lie term tWincnkreis is very convenient.
Unfortunately il has later hecji employed by llerr Kleinsclimiill in a .slightly widened sense, closely
.■dlied species being sometiuK's ineluded in the lhrmcn/:rci-<. In this sen.'-e the Fvrmiiitinie is a kind
of half-caste between species and .subgenus, ami the Linnean binomiiud specilic formula bcuig employed
for it by Kleinschmidt obscures the distinction between .species and uon-species. The older delinitiun
of the term by I.orenz was preciso, correct, and has priority ; there is therefore no reason for modifying
the moaning of the term.
(431 )
species difterent from the male, the wAme. glaucus, given to that kind of female only,
is tiic name of the entire Ri)ecies. Sirailarl}', thongii only the male of an Amboina
Papilio was named priamus by Linnt', the female being described by liim as a
different species, the name of the species is piiamus. In neither case is a new name
necessary or permissible. In a great many instances different individnals (sexes
or otherwise) liavo been described as separate sjiecies. The first name given to
any specimen is the name for all, however restricted the original ajiplication of
the name may have been. It appears to us further self-evident that the philosophic
conception which an author may have of " species " or " variety " cannot be
permitted to affect the name of the forms, which are realities in nature. Whether
we believe that Paiiilio machaoii is the product of a special act of creation or the
product of evolution ; whether we believe that the various varieties from the
Old and New World constituting the species machaon are evolved from an ancestral
homomorphic created species, or that each variety has been created as such ; wliether
we believe that the species is the product of evolution by slow degrees, or per
saltum, by Natural Selection, or by the direct influence of external conditions,
etc., etc. ; all such differences of opinion cannot be allowed to overthrow the name
maclmon for this species of iSwallowtail, unless one wishes nomenclature to become
chaotic. In the same way, the subsjiecies (= geographical race) takes that name
which is the first given to a member of this subspecies, whatever conception the
author of the name may have had of the individuals so named. Since (jlmicus
was the first name for a specimen of a Nearctic Swallowtail, it is the name for the
southern Subspecies to wliich that specimen belonged, as well as the name for
the entire species. The formula for this subspecies is therefore Papilio glaucus
fflaucus.* This formula is precise, showing at one and the same time that the
species is geographically variable, and that the particular subspecies thus designated
was the first one of that species of which a sjiecimen, or sjiecimens, received
a name.
The number of systematists who object to having sjiecial names for subspecies
appears to be very small as compared with those who deal with subspecies, at least
* In the Revision of the l^apilios of the .Eastern Hemispliere, tlie first describeri subspecies was
terraeil/*'r;»rt fi/jnca, as it was noraenclatorially the typical form, giving tlie name to tlie species. At the
eml of the Introduction to that essay I stated tliat it is wrong to call the first-named form the " species "
and the later-named forms subspecies of it, but that one ougiit to treat the first-described forni as a
subspecies like the others, "so that one could speak of P. euryptjliis L., meaning the entire species with
all its subspecies, and of P. curyptjlus mri/pylm L., P. eitri/jtyhis lyeatm Feld., P. eitri/jif/hts pamphyUtK
Feld., P. eurypylii^ niihadfl Leech, etc., meaning the local races." The proposal did not meet at that
time with the approval of the co-editors of Xoi: Zool. Subsequently, when I worked out the idea, I
found that Dr. Loreuz had already given expression to the same view some years previously (1S92) in
very lucid language. lu 1895 I had no knowledge that I had been anticipated by Lorenz ; but it has
since dawned upon ime that I have nevertlicless little claim of having invented the formula P. mryjiyhis
eurypylus independently of former authors. One is apt to forget where one's ideas originally came
from. In the Cataloguti Coh-iy/tefori/m by Stein and Wcise, whicli was one of my treasures wlien a
schoolboy, I found a sample of nomenclature which was at first very puzzling to me. Under the sjKicies
Carahus schi'iiUcri (of course written with a capital S in tlie Catahnjiin) tliere was a whole string of
varieties, one of which was named var. scheUlU'ri. When I came to understand the meaning of this
formula Cartibits Sfheidlrri var. scheidlcri I was much impressed with the wisdom of thus designating
the first-described variety in contrast to the whole species to which it lends the name. This impression,
I think, expressed itself in 1895 in the formula P. eiirypyhis evryjiyhix.
\Vc have some hope that entomologists will sooner or hater all follow the Cataloyvs of Stein and
Weise, not only in this particular instance, but carry out consistently the nomenclature exemplified
by Carahm sdickHnl var. selieidU-ri, at the same time dropping the misleading .and unnecessary " var."
in the case of subspecies (= geographical races), calling the English Carabus arveiixis by the concise
formula Varahm arrensis aiiyticun.— K.J,
(432)
nomenclatoriall}-, as if they had the rank of species. Nor can the former view
be justified. There would be much more justification iu rejecting names for the
lower grade varieties. However, entomologists on the whole apjjear to be inclined
to multiply names for individuals characterised by some striking peculiarity. Now,
it seems to us obvious that it is impossible to provide consistently a name for every
peculiarity and combination of special characters observed in the specimens, since
every individual differs to some extent from every other, and as, further, an
individual may agree with a second iu some peculiarity and with a third in some
other character, and would have to receive two names. The number of individuals
is endless. The number of names cannot possibly be allowed to be so. Therefore,
restriction in naming individuals is absolutely necessary. We believe the most
sensible way of dealing with the range of individual variability, apart from marked
dimorphism, is that of employing, instead of names, descriptive morphological
terms which would cover the corresjjonding individual varieties of all the nearly
and distantly related species. One might have, for instance, one term for all
individuals of I'apilio which have yellow spots instead of the normally red ones,
and another term for the aberrant iudividuals which have these spots white. This
method has been advocated by various authors, and we think will ultimately be
adopted. For a study of variation this method is certainly better adapted than
that of giving an unlimited number of names to individuals. For a specimen
can bear only one name, though the individual may have many peculiarities iu
pattern, colour, and structure ; while by the other method it would be possible
to refer to each peculiarity by a special morphological term if necessary. For
instance, one and the same specimen may be diminutive, tailless, diffuse in
markings, xanthochromatic, heterographic right and left, etc. ; these peculiarities
could not all find expression in a name given to the specimen. However,
the method requires carefiil working out before it can be successfully applied
iu all groups.
We have considered it sufficient in this Revision to have special names for
seasonal forms, and for the forms of conspicuously di- or trimorphic species
respectively subspecies, also in the case of this di- or trimorphism obtaining in
one sex only.
Bearing these explanations in mind, the reader will not find any difficulty in
understanding the system of nomenclature emjiloyed.
A fairly large number of subspecies and a few species are here described for
the first time, and we feel sure that there are a good many new ones yet undis-
covered in those parts of South and Central America and the West Indies which
are not exhaustively explored. The species and subspecies which are represented in
collections by uniques or by very few specimens is suspiciously large, always a sign
of the incompleteness of our knowledge of the fauna of the respective districts.
The interior of Brazil, especially the province of Goyaz, is still a good field for a
collector. The districts north of the mouth of the Amazons are also practically
untouched by entomological collectors, not to speak of the mountains at the
boundary of the Gnianas. "West and North Peru, the north coast of (Colombia,
the Atlantic side of the Volcano de Chiriqui, the mountains of Costa Uica, West
Mexico, and especially the island of Haiti (and S. Domingo) will doubtless yield
interesting results to a competent explorer ; and the swamjts of the Amazonian
region may still harbour some unknown species allied to I'apilio triopas, aeneaa
or (crtumnus.
( 433 )
The Papilios, inclusive of the so-called genns Troides (= Oinithoptera), {a,\l into
three natural groups, which arc sharply separateil in the larval, pnpal and imaginal
stages. This classification was given by Horsfiehl in 1857 for the Indo-Malayan
Papilios (in Horsf. & Bloore, Cat. Lep. Ins. Mas. E. I. Comp.), being based almost
exclusively on larval characters. Haase, in 1893, recognising the sonndness of
the classification, applied it to all Papilios, separating them correctly into the three
Horsfieldian groups. Investigation in the classification of the Papilios must start
from these primary divisions, as we have repeatedly insisted upon. The recent
attempts by Moore * and Kirby t to divide up the mass of Papilios into small genera
have proved to be more or less abortive, the three large natural sections not having
been recognised by these authors. We find consequently united in one genus models
and mimics which belong to different main groups, in Kirby's genus Itliobalus all
three main groups being represented.!
A detailed description of these primary Sections will be given in our proposed
generic revision of the Papilioiiidae. The following short synopsis, we think, will
suffice for the present :
I. A?v's/o/of7«'«-Swallowtails ; p. 435. The larvae feed on Ariatolochia,
occasionally on allied plants. They are densely covered with minute hairs, which
give them a velvety appearance, the head, prothoracic plate and the thoracic legs
remaining glossy ; each segment bears a belt of tubercles, which vary in length
according to species, but are always fleshy, being covered with fine hairs like the
body, never with heavy spines, one of the tubercles standing beneath the stigma and
another above the legs. The proximal abdominal segments of the pnpa are
depressed dorsally, and, like the wing-cases, dilated laterally, the pnpa being much
more broadened in the centre than in the other two Sections of Papilio ; on each
side of the abdomen there is dorsally a row of ttibercles or flaps, sometimes forming
a nearly continuous crest. The antennae of the imago is not scaled, and appears
to the naked eye less distinctly segmented than in the other Sections of Pajtilio, on
account of the segments not being much compressed or constricted at the base ;
each segment bears a sensory groove ventrally at each side, the grooves being in
most species of this Section deep and ovate (reminding one of the Nymphalid
antennae) ; the sensory pores on the dorsal side of the segments are rather large.
The arrangement of the spines on the tarsi is also characteristic for this Section ;
the outer ventral row of spines (there are normally four ventral rows in Lepidoptera)
is not separated by a sharply defined, spineless, impressed space from the spines of
the dorsal surface, as is the case in the other two Sections.
The American species are distinguished from the Old- World forms by the sinns
of the fifth tarsal segment in which the claws are inserted being much less extended.
This Section is not represented on the African Continent, only one species occurring
on Madagascar, while the species are numerous in the Oriental Region and in
America.
II. Fluted Swallowtails ; p. 537. The larvae are without tubercles, or the
tubercles are hard and bear spines (for instance, in the Oriental species aegeus,
anactus and ch/tia) ; the third thoracical segment is enlarged, the larva therefore
• Lej>i(loj)trra Intlira v. (1901-3).
t Hiibncr, Samvil. Exut. Schmitt. ed. ii. (1110 -; year;).
J Ithohalus as conceived hy Hiibner in ISIX (I) contains only siwcics which are really closely allied
with one another. The exponents of Jlimicry will doubtless be glad to see that mimics have managed to
deceive such old hands at Lepidoptera as Kirby and Moore.
( -134 )
tapering in fmnt and also becomiu<r almnst gradnally thinner backwards. Tho
chrysalis is more or less strongly rngato, often resembling a piece of wood ; the head
and thorax are nsnally curved upwards as in the preceding section, but not so
strongly, being almost straight iu certain species (for instance, cbjtia). The
antenna! segments are more or less narrowed at the bases and somewhat compressed;
the fine sensory hairs are either vcntrally concentrated in a patch on each side,
there being no distinct grooves as in the j)re('ediug Section, or the hairs cover nearly
the whole ventral surface. There is no scaling on the antenna, except at the extreme
base. The tibiae are never incrassate iu the (?, as they often are in the Aristolochia-
Swallowtails ; the ventral spines of the tarsi are separated from the dorsal spines
by a regular, somewhat impressed, sjjineless interspace. The abdominal margin of
the hiudwiug is always curved downwards, having the appearance of being fluted
beneath, the two sexes resembling each other in this respect, while in the males of
the two other Sections the abdominal margin is usually modified, bearing nearly
always a distinct scent-organ.
This Section comprises the majority of the Papilios. It is less homoeomorphic
than the preceding Section. Many species are mimetic.
III. Kite-Swallowtails ; ji- 'j^-i- The most characteristic forms of this
Section somewhat resemble a paper kite (for instance, iloi-cus, cmtlwus, pro/cxi/aiis).
• The third thoracic segment (jf the larva is enlarged, as in the preceding section;
the thoracic segments and the anal one bear often spinelike tubercles, the anal
spines standing close together : in other forms the tubercles are absent or vestigial,
traces of tubercles being usually found on all segments. There are no eye-spots or
oblique bands, the pattern consisting of small dots, or several transverse lines
(belts) on each segment, or more or less irregular longitudinal bands. The
chrysalis is more smooth than in the other Sections ; its head and thorax are hardly
at all curved upwards. The mesonotum bears a pyramidal projection which is
carinate in front and behind and at the sides. The lateral carina is continuous
with the carinate edge of the sheath of the hindwing. The al)domeu bears dorsally
two carinae which converge in front and behind, the anal segment being longer than
broad and almost regularly pyramid-shaped. The antenna of the imago has a
more distinct club than in the previous Sections (which is noticeable already in the
chrysalis). The upperside of the antenna and the tibiae and tarsi are scaled, but
the scales fall off easily in most s])ecies. The arrangement of the tarsal spines is as
in the previous Section. The tibiae are never incrassate in the males. The abdominal
margin of the hindwing is widened in the males and bears usually a distinct scent-
organ. The scalingof the wing is often less dense than iu the previous Section, the
wings becoming transparent distally. In a large proportion of the species the first,
or the first and second subcostal veins of the forevving are anastomosed with the
costa, which does not occur in the other Sections, and the cell of the hindwing is
narrow in most cases, the cross-vein D' (in the third cellule) being more or less
strongly incurved.*
The Section is cosmopolitan, like the preceding, but goes less far north and
south, being essentially tropical. The mimetic American species are all chara<'terised
by red spots situated at the base of the wings on the underside, either on both wings
or on the hindwing only.
* For exi)lan.'ition of ncuration sec diagram after Species No. Ifi'.i,
( 435 )
SECTION I.— ARISTOLOCHIASWALLOWTAILS.
Tlie following gonoric or siibgotieric names have American species as types :
Parides Hiibner (1818 ?) ; type echelus.
Ttliolialus Hiibuer (1818 ?) ; type jioli/damas.
F.iulopofioii Lacordaire (1833); type si'.watn's.
Blahea (Jrote (1875) ; type columhxs (= giiinllricliiaintfi).
Lacordaire (^Ann. Soc. Ent. France ii. p. 384) gives Esclis(-holtz as autlior of tlie
name EmlojiO(/on. We do not know when and where Eschscholtz proposed the name.
The American Aristoloehia-Swallowtails fall into two very distinct subsections.
Subsection A.*
Antenna long ; club slender ; sensory grooves more or less large, sharply
defined ; end segment conical, almost as long as it is liroad. Claws asymmetrical.
Markings of body red. Hindwing usnally witli red band or row of red spots on disc,
these markings seldom white or yellowish white. Forevving of ? bearing often white
or 3'ellowish white patches on disc and in cell, being sometimes all black. Snbbasal
cellule long, widening distally ; PC curved near its base. Cross-veins of forowing
oblique ; upper angle of cell obtuse. Cell of hindwing more or less acuminate, D'
more or less leaning basad anteriorly, the cell-angle D^ — D' being smaller than the
angle D^ — D^, or vein D^ reduced to a point, rarely transverse, never leaning distad.
(?. Scent-organ woolly or densely scaled, no naked streak at its discal side.
Tentli abdominal sternite not reaching to the ape.x of the long and slender tergite.
Tibiae often incrassate and hairy.
? . Anal segment with numerous hairs and bristles which are mostly tapering
to a fine point, others ending abruptly, being somewhat thicker at the tip than at
the base ; in many species there are some bristles whicli are distinctly club-shaped.
The American species which come here can be conveniently placed into three
groups.
Key to the groups :
Fringe-spots white. Hindwing with submarginal spots and
usnally also diseal spots or dots, or a discal band ;
mostly with tail Ascanius Gronp.
Fringe-spots white. Palpus black or red. Hindwing with
discal band or row of spots, but without submarginal
spots Aeneas Group.
Fringe-spots red. Palpus always black. Hindwing marked
as before ......... Lysamler Group.
I. Ascanius Group.
In the preservation of a row of submarginal spots on the hindwing tliis gronp
is more ancestral than the other American Aristolochia-Swallowtails. The forewing
of some species, especially c'ohtmbm, shows also ancestral characters in pattern. The
tail of the hindwing is a third generalised peculiarity, whicii is more strongly and
more generally developed in this grouj) than in the other two groups of red-
spotted species.
Key to the species :
II. Forewing with green-bine band on forewing . . . Species No. 1.
Forewing with white band on forewing ... b.
* Hubscction B. fallows after Species No. 45.
(436 )
Forewing withont band (or onl\- a trace of it)
b. Band of hiiidwing partly red ......
Band of hiudwing entirely white .....
c. Band of forewing angnlatc at lower angle of cell, red anal
spot of hiudwing very large
Band of forewing practically straight ....
(I. Palpns red
Palpns black ........
e. Snbmargiual spots of hindwing sandglass-shaped
Snbmarginal spots of hindwing transverse, oblong or Inni
form
/. One row of spots on hindwing .....
Two rows of spots on hindwing .....
Fringe of forewing completely white or very slightly inter-
rupted at the veins ........
Fringe of forewing uneven, spotted with white .
Forewing with a white dot on disc ; central snbmarginal
spots of hindwing slightly curved
Forewing withont white discal dot
spots of hiudwing strongly arched
As before, but discal spots of hindwing small, more or less
white ..........
9-
I,.
central snbmarginal
discal spots red
/
Species No. 2.
c.
Species No. 3.
(I.
e.
Species No. 7.
Species No. h.
Species No. 4.
9- .
It.
Species No. 0.
Species No. 11.
Species No. 10.
Species No. 8.
Species No. 0.
1. Papilio Columbus H.-Sch. (1862).
Papilin cohimbiis Herrich-Sch., Ou-resphl. Zonl. Min, Ver. lirf/eiish. xvi. p. 141 (1802) (Cuba) ; id.,
I.e. xviii, p. 17.3 (1864).
Papilin i/nmnaihuniiis Felder, ]'erh. ZuuL Bui. Gi-x. Wini xiv. p. 294. n. 75 (1804) (nom. nov. loco
" columlms H.-S.") ; id., lieixe Nornra, Lep. p. 137. D. 101. t. 27. fig. 1. 2 S (1*''5) ; Herr.-Sch.,
Pi-Mh: SijxI. Lep. ii. p. 20(aepar.) (1807) ; Kirby, Cat Diiini. Lep. p. .IHO. n. 120 (1871) ; C.undl.,
Papilio i. p. 113 (1881) (Cuba) ; id., Co?ilr. Ent. Cuhaim p. 124 (1881) (Eiistern Cuba) ; Honr.,
Sitzher. Bill. Ent. Zeit. xxx. p. 4 (1886) ; id., ll,H. Ent. Zeit. xxx. p. 131. t. h. fig. h. ? (188i;) ;
Bonzon, Trans. Amer. Ent. Soc. xv. p. 293 (1888) (Uirva); Haase, Vnlerxurli. ilimirry p. 77 (189.3).
Papilio grolfi Blake, Proc. Ent. Soc. Pliilad. iv. p. 313 (180&) (Cuba).
Blaleea ynndlarliianus Grote, Trans. Ainer. Ent. Soc. v. p. 118 (1875).
As this P. Columbus of Herrich-Schaeffer belongs to quite a different division
of the subfamily Fapilioninae than P. columbiis of Kollar and P. columbiis of
Hewitson, we employ that name for the present species, instead of the later name
gundlachianiis, according to our rules of nomenclature.*
The species comes close to the Brazilian P. ascanhis and agavus. The tibiae
of the male are hardly at all incrassate, bearing bristles and numerous small hairs.
The blue scales of the wings are entire, while the scales in the costal area on
the npperside of tlie hindwing and in the posterior area on the underside of the
forewing are dentate as in the allied species. The anal submarginal spot M- — SM-
of the hindwing is absent from the npperside, being small and short on the
underside. As a remnant of a discal band there are on the underside of the
hindwing three or four white bans proximally of the red spots. The distal edge
of the forewing is dotted with white, as in the Aeneas Group.
Scent-orgau as in P. agavus and allies.
Genitalia iu general structure as in P. agavus and allies, but characteristically
modified, <J. Clasper short, rounded ventrally, the inner surface deeply concave,
' .See p. 423,
( 437 )
the hairy ventral margin very broad distally, being at the apical third nearly
half the v?idth of the entire clasper ; this hairy convex area gradually widening
apicad ; dorsal margin of clasj)er emarginate, the apex acuminate, slightly pointing
npwards. Harj)e elongate, narrowest in middle, lying flat on the clasper, being
cnrved npwards distally ; ventral edge denticulate proximally ; this proximal
])ortion dilated into a rounded or acuminate lobe, which is directed obliquely basad
and ventrad ; apical lobe rounded off, denticulate. Tenth tergite very slender,
slightly incurved in middle. Penis-sheath acuminate, a disc-like piece of chitiu
projecting ventrally from the orifice as in P. agavus. ? . In non-virgin
specimens the vaginal area covered with a hardened substance, which is whitish
and has a spongy appearance ; this coital substance has no snch definite shape
as in P. proncus, but it is always constricted in the middle and there are also
several holes or grooves, which are more or less in the same place in different
specimens. In virgin individuals a broad central process is visible without
dissection ; this process stands behind the vaginal orifice, being somewhat cnrved,
subaeuminate, convex on proximal side, hollowed out on hinder side. In front of
the vaginal orifice there is a heart-shaped lobe covered with minute hairs.
Larva described by Bonzou, I.e.
Hah. Eastern districts of Cuba, especially plentiful in the hills near Santiago
de Cuba.
In the Tring Museum 14 c?c?, 0 ¥ ? , from : Sardinero, Santiago, January 1904
(Wirt Eobinson) ; Gibara (Tollin).
2. Papilio ascanius C!ram. (1775).
FapiWo Equcs Trojanue ascanius Cramer, Pap. Exot. i. p. 20. t. 14. fig. A (1775) (Rio de Janeiro) ;
Goeze, Enl. Biylr. iii. 1. p. 42. n. 14 (1779) ; Fabr., Spec. Ins. ii. p. 2. n. G (1781) (Brazil) ;
Drury, Ilhistr. Exot. his. iii. p. 11. t. 9. fig. 1 & Index (1782) (Rio de Janeiro) ; Jabl. &
Herbst, Naturs. Schmelt. ii. p. 148. n. 3G. t. 13. fig. 3 (1784) ; Fabr., .]fciii!. Ins. ii. p. 2. n. 7
(1787) ; Gmelin, Syst. Nat. i. 5. p. 222G. n. 274 (1790) ; Jung, Alplmb. Vcrz. Schmett. p. 57
(1791) ; Fabr., Ent. Sysl. iii. 1. p. 3. n. 8 (1793).
Menelaides ascanius, Hiibaer, Verz. heh. Sclunell. p. 85. n. 871 (1818 ?) ; id., Samml., Exot. Schmett.
ii. t. 105(1822?).
Papilio a.'icanius, Godart, Enc. Melli. ix. p. 73. n. 138 (1819) (Brazil) ; Lucas, Up. Exot. p. 31. 1. 16.
fig. 1 (1835) (Brazil) ; Boisd., Spec. Gin. Lip. i. p. 30G. n. 141 (18.iC) (Rio de Janeiro ; North
Brazil) ; Drury, ed. Westw., Illustr. End. Ins. iii. p. 11. t. 9. fig. 1 (1837) (Rio de Janeiro) ;
Dune, in Jard., Xal. Libi: xxxvi. p. 101. t. 3. fig. 1 (1843) ; Doubl., List Lep. Ins. Brit. Mus.
i. p. 13 (1845) (Brazil) ; Lucas, Lip. Exot. ed. ii. p. 31. t. IG. fig. 1 (1845) ; Doubt, Westw. &
Hew., Gen. Diurn. Lcp. i. p. 18. n. 190 (184G) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 42.
n. 217 (1852) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 58. n. 2;S0 (1856) (Brazil) ; M.'niStr.,
Enum. Corp. Anim. Mux. Pelrop., Lip. i. p. 5. n. 73 (1857) (Brazil) ; Felder, Verli. Zool. Bot.
Ges. Wten xiv. p. 294. n. 73 (18G4) (S. Brazil) ; Butler, Cat. Diurn. Lep. descr. Fabric, p. 236.
n. 9 (18G9) (Brazil) ; Kirby, Cat. Diurn. Lep. p. 530. n. 117 (1871) ; Capronn., Ann. Soc. Ent.
Bclg, xvii. p. 9. n. 5 (1874) (Icarahy, Aug. ; Botafogo, Nov.) ; Burm., Discr. Rip. .iri/eut. v.
Lip., Atlas p. 8. n. 20 (1879) (descr. of egg, larva and pupa; Rio de Janeiro); Oberth.,
Et. d'Ent. iv. p. 77. n. 249 (1880) (Brazil) ; Stand., Exot. Tagf. p. 14 (1884) (Brazil) ; Haa.se,
Untenwh. Mimicry i. p. 77 (1893) ; Bonningh., Vcrh. Ver. Nat. Unterh. ix. p. 27 (189G) (Rio
de Janeiro).
Ileclurides ascanius, Hiibner, Sanml. Exot. Schmett. ii. t. 105. $ (1822?); Kirby, in Allen's Nat.
Libr., Lep. Butt. ii. p. 270. t. 65. fig. 1 (189G) ; id., in Hiibn,, Samml. Exot. Schmelt. ed. ii.
p. 89. t. 318, fig. 3, 4 (1905?*).
<??. Median band very broad on both wings, wider in female than in male,
washed with red on hindwing, traversing apex of cell on forewing, but usually not
* The new edition of Hiibner is being issued in parts, no year of publication being given, even with
the new text written by Kirby.
( 438 )
reaching (|nite across cell. The hand of the forewiug nearly the same in position
as the hlno-green hand of the (Iiibiui P. cobimbus {= guiullueliianus); some males
have obscnro white spots distally of apex of cell, while in some females these
spots, as well as a triangular streak sitnated before cell, are qnite distinct ; in one
of onr females the band extends basad to point of origin of M-, being very broad
also on hindwing. The anal submarginal spot is comi>k'tely merged together
with the anal spot of the discal biuid in most specimens, hnt in some individnals
there is a separate snbmarginal anal bar distally of the last patch of the band.
Genitalia scarcely diflereut from those of P. cliamissonia and jKrrhehm ; harpe
])roximally a little wider.
Early stages described by Burmeister, he.
Ilab. llio de Janeiro.
Jn the Tring Musenm 13 c?c?, 4 ? ¥.
3. Papilio agavus Urury (1782).
Pajiiliri Rjiies TrnjuiiuK h/snmhr Fabricius (non Cramer, 1775, err. dot.), Gen. Ins. p. 251. n. •2:5-4
(177l'i) (" India," Dr. Fothergill.— Mutilated specimen of this species ?) : Ooeze, I-:nt. Bnjtr. iii.
1. p. 4fi. n. 29 (177'.!) ; Fabr., Sim-. Ins. ii. 9. n. 3?> (17H1) ; id., .Iliinl. liii. ii. p. 3. n. 20(1787) ;
Gmelin, .S><. Nat. i. 5. p. 222',l. n. 285 (1790) ; Jung, Alphab.Verz. Schmetl. p. S.W (1791) ;
Fabr., Ent. Syst. iii. 1. p. 9. n. 25 (179.^).
rnpilin liiines Achivus (iijarus Drury, IHuxtr. Exot. Ins. iii. p. 11. t. 9. Eg. 4 & Index (1782) (Uio de
Janeiro) ; StoU, in Cram., Fap. E.rot., Snppl. p. 144. t. 32. fig. 1 (1790) (Rio de Janeiro).
Princqis hcroicns agavus, Hiibner, tiamniL Ej-nt. Srhniflt. i. t. 113. fig. I. 2. (J (ISOfi— ).
Menelimlrs ognrus, id., Verz. brk. ScJimell. p. 85. n. 872 (1818 ?) (= li/mnili'i- Fabr.).
Pupilhi agavus, Godart, Enc. Mel!), ix. p. 73. n. 137 (1819) (Brazil); Boisd., S/)cc. Gin. Up. i.
p. 30i;. n. 142 (1831'.) (Brazil) ; Drury, ed. Westw., Illnstr. E.n.t. Ins. iii. p. 12. t. 9. fig. 4
(18.37) (Brazil) ; Doubl., List Lep. Ins. Brit. Mm. i. p. 13 (1845) (Brazil) ; id., Westw. it-
Hew., Gen. Uiurn. Lep. i. p. 17. n. 189 (184G) ; Gray, Cat. Lep. Im. Brit. Mns. i. Pap. p. 42.
n. 216 (1852) ; id.. List Lep. Ins. Ihil. Muk i. Pap. p. 58. n. 229 (1S56) (Brazil) ; M.'n.'tr.,
Ennm. Corp. Anim. Mns. Petrop., Lip), i. p. 5. n. 72 (1857) (Brazil) ; Felder, Ve)-h. Zuul. Bot.
Ges. Wien xiv. p. 294. n. G8 (1804) (Brazil) ; Prillw., Stelt. Enl. Zeit. xxvi. p. 130 (18(15)
(Corcovado) ; Kirby, Cat. Binrn. Lei), p. 53G. n. 112 (1871) ; Burm., Deser. Ri-p. Argent, v.
Lip. p. C6. n. 8 (1878) (Rio de Janeiro ; Paraguay ; Corrientes) ; id., I.e., Atl»s p. 9. n. 21
(1879) (Rio de Janeiro) ; Capronn., Ann. Snc. Ent. Belg. xvii. p. 9. n. 6 (1874) (Botafogo.
Aug. ; Rio, Oct. ; Copa Cabana, Oct. ; common) ; Oberth., /■."/. i/'Eii/. iv. p. 77. n. 247 (1880)
(Brazil) ; Stand., E.i-nt. Tayf. p. 14, t. 9. ? (1884) (Rio de Janeiro ; Sao Paulo) ; Haa.sc,
Unte)-s),eh. Mi)nicr)j i. p. 77 '(1893) ; Weym., Stett. Ent. Zeit. \v. p. 315. n. 8. (1895) (Rio
Grande do Sul) ; Mabilde, Guiu P)-aet. Borhol Rio G)-ande do S)d p. 48 (189(1) ; Biinningb.,
Ve)h. Ver. Nat. Unlei-h. ix. p. 28 (189(1) (Rio de Janeiro); Eimer, Orthogen. p. 137. fig. til
(1897) ; Peters, Jlliislr. Zeilselir. Ent. ii. p. 52 (1897) (Nova Friburgo, common ; pa)iin)).
Papdh) lysandfr, Butler, Cat. Diia-n. Lep. descr. Fabric, p. 238. D. 17 (1869) (Fabr.'s " lysamkr is
evidently harrisianns," false).
nectorides agavus, Kirby, in Hiibner, .'ianind. E.rot. .?«■/,/»«//. ed. ii. p. 89. t. 113. fig. 1. 2 (1905 ?)
d ? . Anal snbmarginal spot large on npperside, in male completely merged
together with the preceding spot, the edge of the wing at anal angle being
very narrowly black ; in female the two spots connected, the anal si)ot being
produced distad along M=, the two spots forming a Z-shaped mark, which is well
separated from the anal sinns. There is often a minute dot in cell of forewing
in male.' The white band of the hindwing is nsnally composed of three spots,
which vary in size, the second spot being sometimes missing, while in other
individuals there are one, two or three small additional spots, sitnated one in front
of and two behind the apex of the cell. Vein M" of hindwing has a more distal
position than in the allied species,
(439)
Genitalia: d. Tenth tergite slenderer than in P. jjroneus. Ilarpe long,
kuife-like, the distal half denticulate ventrally ; apex rounded dorsally, being
veutrally acuminate and somewhat twisted. Penis-sheath acuminate ; a flat,
rounded disc of chitin jirojecting ventrally from the orifice of the sheath, the
disc being continuous with the membranaceous ventral portion of the sheath.
?. In front and at the sides of the vaginal orifice an irregular ridge, much folded,
being semi-membranous, forming a ring which is open distally in the middle ;
within this ring and just behind the vaginal orifice a short process, curved anad,
being convex ventrally, hollowed out on hinder (or upper) side ; the membrane
connecting the ridge with the seventh sternite den.sely folded, there being a more
strongly chitinised, smooth, rounded lobe laterally where this membrane joins the
seventh sternal sclerite ; in a non-virgin female the central process is envelo})ed by
a hardened substance blocking up the vaginal orifice. Anal segment with numerous
short stout bristles.
Early stages described by Burmeister, I.e.
Hah. Brazil : Minas Geraes to Rio Grande do Sul ; Paraguay ; Argentina :
(.'orrientes.
In the Tring Museum 34 cJt?, KJ ? ?, from : Minas Geraes, March and April
(A. Kennedy) ; Tijuca and Petropolis, March ; Castro, Parana, March and October
(E. D. Jones) ; Sapucay, Paraguay, January, August, October, December (W. Foster) ;
Yhu, Paraguay, September to December (Audeer).
4. Papilio proneus Hubu. (1825).
Becturides praneus HUbner, Summl. Exot. Schmett., Zutrdge p. 25. n. 249. fig. 497-8 (1825) (Brazil).
Papiliu prmeus, Boisduval, Sjicc. Gen. Lip. i. p. 307. n. 143 (183(5) (Brazil) ; Doubl, Westw. and
Hew., Gen. Diiirn. Lep. i. p. 17. n. 188 (184(5) ; Doubl., Lht Lrp. ]„>:. Brit. .1/h.s. i. Apprwl p. 3.
(1848) (Rio de Janeiro) ; Gray, Cat. Lep. Ins. Brit. Mns. i. Pap. p. 42. n. 214(1852) ; id., List
Lep. Ins. Brit. Miis. i. Pap. p. 58. n. 22(5 (185(5) (Brazil) ; Me'ne'tr., Knuni. Corp. Anim. Mus.
Petriip., Lep. i. Suppl. p. 68. n. 1125 (1857) (Brazil) ; Felder, Verh. Zuol. Pot. Ges. Wien xiv.
p. 294. n. 70 (18(54) (.S. Brazil) ; Kirby, G(t. Diiirn. Lep. p. 536. n. 114 (1871) ; Burm., Descr.
Rep. Arijcnl. v. Lep. Atlas p. 9. n. 22 (1879) (Petropolis ; Nova Friburgo) ; Oberth., Et. d'Ent.
iv. p. 77. n. 246 (1880) (Brazil) ; Staud., E.rat. Tag/, p. 14 (1884) (Rio de Janeiro ; Sao
Paulo) ; Biinningh., Verli. Ver. Nat. Unterh. Hamburg ix. p. 28 (1896) (Organ Mts.).
Papilin phrijneus Lucas, Rev. Znol. (2) iv. p. 13G (18.52) (" Cayenne " loci error) ; Gray, Cat. Lep.
Ins. Brit. Mils. i. Pap. p. 42. n. 215 (1S52) (Brazil ; "Cayenne"') ; Kirby, Cat, Dinrn. Lep.
p. 536. n. 113 (1871) (" Cayenne " loci error).
(? ? . Red submarginal spots of hindwing transverse, oblong or Inniform, not
sandglass-shaped as in F. chamissonia ; anal spot not V-^^li^-peJ, there being uo
discal spot proximally of the anal submarginal one. Width of central band and
the number of spots composing it on hindwing variable. Foretibia and first
foretarsal segment more finely hairy, less spinose, than in P. ugams.
Genitalia different from those of the allied species, the female bearing after
copulation a kind of pouch, externally visible and homologous to the pouch of
Euri/ar/c.s, Pafncissiii.'<, and Acraea. S- Tenth segment as in the allied species,
the tergite being very long and slender. No separate harpe on clasper, the harpe
being rej(resented by a polished central space which extends from base to middle,
being rounded distally ; this space depressed, a triangular distal portion of it being
slightly elevate ; ventral edge of clasper with a short stout conical tooth. Peiiis-
shcatU strongly chitinised at apex, ending in a sharp point. -?. In a virgin
specimen there is at each side of the vagina a large flap, rounded, asymmetrical,
bearing distally several carinae, the two flaps inclining towards each other ; from
( 440 )
the slit between them, which wiileus distally, tliere projects a curved process
pointing anad ; these organs are distinct without dissection, projecting free, not
being covered by the scaling. On dissection tlie central process is (bund to be situated
immediately behind the vaginal orifice, being convex ventrally, and channelled
on the posterior or dorsal side ; the apex of this process is slightly incrassate,
giving the process a feebly ladle-shaped outline in lateral aspect. The lateral
flaps are continuous proximally of the vaginal orifice, being the lateral portions
of a ridge which is almost entirely sejiarated into two halves by a deep mesial
sinus. In a female which has copulated the central process is enveloped by a
hardened substance which forms a large irregular cone effectnally blocking up the
vaginal orifice, the lateral flaps remaining free, at least distally, the hardened
substance covering the flaps only proximally.
Early stages not known.
Hah. Brazil : Minas Geraes to Parana, perhaps farther south.
lu the Tring Museum 42 c? c?, 14 ? ? , from : Minas Geraes, April (A. Kennedy) ;
Petropolis, October— February (A. Foetterle) ; Bahurn, Sao Paulo (Dr. Hemjjel) ;
Castro, Parana, November (E. D. Jones).
5. Papilio chamissonia Eschsch. (1821).
rupilio iliiuitissonia Eschscholtz, in Kotzebue, /iViV iii. p. 203. n. 3. t. 2. fig. 3. jf (1821) (Brazil).
Meiielaides huuirhu.t Hiibner, Siimml. Exnl. Sclimrll. ii. t. 1(13 (1822 ?)
Papilio ascahis Godart, Kite. Meth. ix. Siippl p. 812. n. 137-8 (1824) (Brazil).
c? ?. Upperside of wings with a slight metallic green tint ; white .spot in cell
of hindwing not extending farther basad than point of origin of M=, white
sjmt M' — M- reaching at least as fiir distad as spot IP — BI'; red submargiual spots
constricted, sandglass-shaped ; a red (discul) bar iiroximally of anal bar, connected
with the latter to form a V-shaped spot ; harpe of c? a little narrower, and central
process of ? a little longer.
Genitalia as in P. perrhchus.
For early stages described by E. D. Jones, see below.
Hab. Brazil.
Two subspecies.
a. P. c/tatnissonia diodorus Hopff. (18(56).
Papilio diodorus HopfiEer, Stett. Eiil. Zcit. xxvii. p. 23. n. 2 (186G) (Brazil) ; Kirby, Cat. lynirii. Lcp.
p. 567. n. .333 (1871).
Papilla echcdiiriis, Oberthiir (nan Boisduval, 1836, err. det.), Et cFEiit. iv. p. 77 n. 244 (1880).
Papilio raiiipciiv Foetterle, Rei\ Mus. Paulista v. p. 622. t. 15. fig. 2. $ (1902) (Minas Geraes).
c? ? . Fringe of both wings entirely white. Band variable in width, straight
on hindwing, diodorus being based on narrow-banded individuals, and campeiro on
narrow- and broad-banded ones, the difference not being gcograpiiical.
There are two Buisduvalian specimens in coll. Oberthiir which belong to this
form. In 1880 Oberthiir considered them to be typical sjiecimens of eclicdorus.
However, according to the description and the locality — Sta. C'atharina — Boisduval's
echedonis is the next form. Probably Boisduval replaced his original specimens
later by some better preserved ones wliicli happened to belong to another
geographical race.
Hub. Minas (jeraiis, interior of Sao Paulo ; (ioyaz ; San Antonio de Barra,
Bahia.
In the Tring Museum ASS from Minas Geraes, February (A. Kennedy).
( 441 )
b. P. chamissonia ckamissonia Eschsch. (1821).
Piipilio chamisnoitki Eschscholtz,?.!'. ; Gray, Cat. Lfp. Ins. Brit. Mus. i.Pap. p. 41. n. 212(1852) (Brazil);
id., List Lep. Jn>:. lirit. .l/««. i. Pap. p. 57. n. ■224 (1856) (Brazil) ; Felder, Verh. Ziml. Bnl. Ges.
^Vic>l xiv. p. 294. n. 72 (1864) ( = echedorus = ciiri/dorus) ; Kirby, Cut. Diiirn. Lrp, p. 536.
D. 115 (1871) ; Oberth,, Et. d'Ent. iv. p. 77. n. 24.3 (1880) (Brazil) ; Staud., Emt. Tagf. p. 14
(1884) (S. Catharina).
Meiielaules hunichux Hiibner, I.e. ,
Papilio asailits Godart, I.e.
Papilin eihedorns Boisduval, Spee. Gen. Lep. i. p. 308. n. 144 (1836) (S. Catharina) ; Doubl., Westw.
and Hew., Gen. Di'irn. Lep. i. p. 17. n. 187 (1846) ; Gray, Cat. Lep. Ins. Brit. Mux. i. Pnp.
p. 42. n. 213 (1852) (Brazil) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 57. n. 225 (1856) (Brazil V) ;
M('nutr., Eniini. Corii. Anini. Mas. Pelmp,, Lep. i. p. 4. n. 71 (1857) (haec subsp. ?).
Papilin hiiniehus, Boisduval, I.e. n. 145 (1836) (Brazil) ; Doubl., List Lep. Ins. Brit. .Mas. i. p. 13
(184.5) (Brazil) ; id., Westw. and Hew., Gen. Diurn. Lep. i. p. 17. n. 186 (1846) ; Gray, Cat. Lep.
Ins. Brit. Mns. i. Pap. p. 41. n. 211 (1852) (Brazil) ; id. List Lep. Ins. Brit. Mas. i. Pap. p. 57.
n. 223 (1856) (= aseahis ; Brazil) ; Mi'netr., I.e. n. 70 (1857) ; Felder, I.e. n. 71 (1864) (Brazil) ;
Kirby, I.e. n. 116 (1871); Burm., Deser. Rep. Argent, v. Atlas p. 9. sub n. 21 (1879) (S.
Catharina; Pt. Alegre ; "not at Rio de Janeiro"); Oberth., I.e. n. 245 (1880) (Brazil);
Jones, Prve. Lit. Philos. Sue. Liverpool p. 15 (1883) (metamorphosis) ; Stand., Ea-ot. Tagf.
p. 14 (1884) (Rio de Janeiro : Sao Paulo) ; Haase, Untersueh. Mimienj i. p. 77. t. 10. fig. 69. ?
(1893) ; Mabilde, Gtiia Praet. Bnrhul. Rio Grande do Sul p. 48 (1896).
Papilio eurydorus Luca.s, Rev. Zoul. (2). \v. p. 135 (1852) (S. Catharina) ; Doubl., Westw. and Hew.,
Gen. Diurn. Lep. ii. p. 529 (1852) ; Gray, Cat. Lep. Ins. Brit. .Mus. i. Pap. p. 42. n. 216 (1852)
(Brazil) ; id., List Lep. Lis. Brit. Mns. i. Pap. p. 58. n. 228 (1856) (Brazil) ; Lucas, in Casteln.,
Voy. Amer. Sud^ iii. Lej,. p. 198. t. 2. fig. 1. J (18.57).
Hectorides hunichus, Kirby, in Hiibn., Samud. Exot. Schrnelt. ed. ii. p. 89. t. 316. fig. 3. 4 (190 — ?).
c? ? . Fringe of forewing less e.xtended white than in the preceding form,
sometimes all black ; fringe of hindwing black at t,he apex of the teeth. White
band variable in width, on forewing often very thin and anteriorly obsolete ; on
hindwing on the whole narrower and posteriorly less widened in specimens from
Santa Catharina {chamissonia = echedorus = euri/dorus) than in the individuals
from Sao Panlo and Rio de Janeiro (bunichus) ; however, the diiference does not
hold good.
Hab. Petropolis ; Sao Paulo ; Parana ; Santa Catharina.
In the Tring Museum 24 <?c?, IS ¥ ? , from : Petropolis (Foetterle) ; Sao Panlo,
August to January (E. D. Jones) ; Jnndiahy ; Theresopolis, S. Catharina
(J. Michaelis) ; Castro, Parana, October and November (E. D. Jones).
6. Papilio perrhebus Boisd. (1830).
Papilio perrhebus Boisduval, Sjjcc. Gin. Lep. i. p. 305. n. 140 (1836) (Paraguay ; Rio Grande).
Papilin perrhaebus (!), Burmeister, Dcser. Rtp. Argent, v. Lep. p. 65. n. 7 (1878) ; id., I.e. Atlas p. 8
t. 2. fig. 8, t. 3. fig. 10 (1889) (larva ; g).
cJ ? . No discal band on wings. Submarginal spots of hindwing transverse,
constricted or interrupted in middle ; a spot proximally of anal one representing
the discal row, this spot distinct only below, often joined to the anal submarginal
bar, the two spots forming a kind of V ; some specimens have a small white discal
spot W — M-. Cell of hindwing broader distally than in the .allied species, and
the sulibasal cellule longer.
tJenitalia: cJ. Tenth tcrgite slender, tapering, much longer than the sternite.
Harjie reaching close to apex of clasper, snblinear, slightly tajiering at apex,
rounded at base ventrally, dorsally subangulate a short di.stance from apex, the
whole ventral edge and the dorsal edge from apex to the before-mentioned angle
denticulate. ? . Behind vaginal orilice a curved, ladle-shaped process, convex
( 442 )
Vpiitrall}', excavate on tipper or distal side ; carved aiiad ; iiroximally of orifice
a low folded ridge extcndiug ou each side bcyoud tlie central process, formiug a
ring which opens behind ; this ridge widest behind ; at each side of the ridge
the membrane densely plicate and fnrthcr latcrad again raised into a smootli,
somewhat ronnded ridge.
Early stages described by Bnrmeister, l.r.
Ilab. Sao Panlo to Buenos Aires and northwards to Paraguay.
Two subspecies.
(I. P. perr/tcbus perrhebus Boisd. (183G).
I'ajiilio ]>en-hi>l/ii.i Boisdiival, I.e. ; Doubl., Wesfcw. and Hew., Gen. Diiirn. Lej>. i. p. 19. n. 226
(I84U) ; Gray, Cal. Le/,. I„s. liril. Mnx. i. Pap. p. 41. n. 20i; (1852) ; id., Lhl Lep. Ins. Biit.
.Mils. i. Piiji. p. hi\. n. 217 (18.^11) (Paraguay) ; Felder, Verh. ZouJ. But. Grs. Wini xiv. p. 2y4.
n. 74(1864) (Paraguay ; Rio Grande) ; Kirliy, Cut.Dinni. Lfp.^i. 536. n. ll'J(1871) (Paraguay);
Oberth., Et.il'Ent. iv. p. 97. n. 248 (1880) (Paraguay); Gosse, Eiitnm. .\iii. p. 194(1S8(I)
(Assuncion, not scarce, Nov. Dec.) ; Staud., Etnl. Tiiij/'. p. 15 (1884) (Paraguay ; Rio Grande
do Sul teste Boisd.) ; Ha.ise, Viilermirh. Miiuk-nj i. p. 77 (1893) ; Weym., Strtl. EnL Zuil. Iv.
p. 315. n. 7 (1895) (Rio Grande do Sul).
Ptqiilio pcrrhiiibitii var., Mabilde, (jtiia Prart. Borhol. Rin Grande do Sul p. 48 (1896).
c? ? . Frons and palpus red. Submarginal spots of hindwing bright red on
both sides. Fringe of hindwing black at the ends of the veins.
I/iib. Brazil : Sao Paulo to Ilio Grande do Sul ; Paraguay and neighbouring
districts of Argentina.
In the Tring Museum 12 c?c?, 6 ? ?, from : Sao Panlo; Itio Grande do Sul ;
Sapucay, Paragnay, August to October (W. Foster).
b. P. perrhebus damocrates Guen. (1872).
Piipiliii damocrales Guenee, Mem. Soc. Fhijs. Hist. Nal. Geneve xxii. p. 371. n. 4. fig. 2. ^f (1872)
(Buenos Aires); Staud., E.ivl. Ta(if. p. 15 (1884) (Argentina).
Piipilii, pnrliibiis^ Guenee, Pel. Niiiiv. Enl. p. 201 (1872) (= dainorralcs) ; Kirby, thid. p. 239 (1872)
(= ihiiiioa-iiles) ; Guen., /.<;. p. 244 (1872) ; Kirby, Oil. IJInni. Lip. \\ 810. n. 119 (1877)
(partim ; dainocmtes = perrhebus) ; Staud., l.r. t. 9. ^ (1884) (the figure too blauk).
PujiiUo perrliaebii-i, Burmeister, U.it. (Buenos Aires ; 1. on Aristuhichin riliiita).
Piijiiliu perrhebus var. duniocrute", Oberthiir, Et. d'Ent. iv. p. 77 and 115. sub n. 248 (1880)
(Buenos Aires).
S ¥. Paler than the preceding, es])ecially the female ; red colour on liody more
restricted, palpus and frons usually quite black. White border of forewing wider
anil that of hindwing not interrupted ; submarginal spots less distinct, paler on
both sides.
J fab. Buenos Aires and Entre Rios, probably also in Uruguay.
In the Tring Museum 31 Jc?, 24 ? ? , and a series of larvae and jiujiac, from :
Buenos Aires, December to February (Ruscheweyh) ; Paysaudu ; La Soledad, Entre
Rios, end of November to January (('has. Britton and Miss E. A. Britton).
7. Papilio phalaecus Hew. (1861).
Piijiilio phiiltieriis Hewitson, Triiiia. Enl.. Sue. Lund. p. 32 (18159) (Ecuador) ; id., Ejul. Bull. iv. t. 11.
fig. 37 (1869) ; Kirby, Cut. Dinrn. Lep. p. 536. n. 1 18 ( 1871) ; Stand., Ej:ol. Taijf. p. 14 (18S4);
Dogiiin, Emini: Lfp. I.nj,i i. p. 14 (1887) ; Maass. i>i Wtyni., in Stiibel, lieiscn S. Aimr., Lrp.
p. 61. n. 87 (1890) (Muamboya) ; Dognin, I.e. ii. p. 37 (1891) ; Uaase, Untersiich. .Uiinkrij p. 77
(1893).
Piipilio phalueiliiis (!), Hewitson, Exnt. Bull. iv. Jiidr.r (1872).
c??. Body woolly ; head, prothurax and palpns Idack, a few red scales beliiiid
eye. Mid- and hindtibiae minutely hairy, and armed with dispersed spines, as in
( 44:i )
P. agartis. A white band from costal margin of forewing to anal angle of liiiidwlng,
jiiirallel to distal margin of forewing, shaded with black scaling on forewing and
distally on hindwiiig, interrnjited by the black veins ; the band close to cell on both
wings, wider in ? than in c? ; ? with white spot in cell of forewing ; a row of red snb-
marginal spots on hind wing, denselj' shaded with black on nj)perside, especially in 6.
Scent-organ : fold rather narrower than in the allied species.
Genitalia : c?. Clasper rounded at apex or slightly eniarginato ; harpe longi-
tudinal, mnch shorter than tlie clasper, the apici-ventral marginal area of the
latter being broad, apes of harpe feebly acnminate, a little cnrved ujiwards,
ventral and apical edges dentate, at base of harpe two heavy conical teeth, both
vertical on the plane of the liarpe, one standing at the ventral edge, the other
farther dorsad. ? not dissected.
Early stages not known.
Ilab. Eastern Ecuador.
This is the only American species of ,4r?.??o/oc/«'a-feeders bearing a spatidate
tail which occurs in a central district of the Nootropic region, all tiie other tailed
yl;7.s<ote/(/a-Swallowtails being found either in Brazil and the Rio La Plata
(R. Paragnay and H. Parana) districts, or in Central America from Co.sta Rica
northwards, and the West Indian Islands Cuba and Haiti.
In the Tring Museum, 2 S i from : Loja, July 18s0 ; Zamora (0. T. Barorj).
8. Papilio photinus (Donbl.) (1844).
Papilin phnliiius Doubleday. Ann. .Uxf/. X. II. xiv. p. 415 (1844) (W. Mexico?) ; id., Lid Lip. Ins.
ISi-U. Mas. i. p. \i (184.5) (W. Mexico ?) ; id., Westw. & Hew., Gen. Dhirii. Lep. i. p. 17. n. •-'■J9
(1847) (Mexico) ; Gray, OU. Lep. Im. Brit. .Uns. i. Pap. p. 65. n- 287. t. 11. fig. 2. ? (18,52)
(West coast of America) ; id., List Lep. Ins. Brit. Mas. i. Pap. p. 75. n. 304 (185(5) (West coast
of Am. ; Mexico) ; Weidem., Proc. Eiit. Stir: Plillail. ii. p. 147 (18C3) (.Mexico) ; Felder, Vfrli.
Znal. Hot. Ges. Wieii xiv. p. 294. n. 77 (1804) (Mexico) ; Kirby, Cat. Uim-n. Lep. p. 53Ci. n. 122
(1871) (Mexico) ; Guen., Mem. Sue. Phi/s. Jlist. Nat. Geiiiire p. ,379 (1872) (Mexico, t? ? ) ;
Kirby, Pet. Noiiv. Ent. p. 23'.) (1872); Butl. & Druce, Pnc. Zanl. Sor. Land. p. Srt.l n. 31)4
(1874) (Costa Pica) ; Oberth., Et. d'Ent. iv. p. 80. n. 262 (1880) (Mexico) ; Staud., Exnt. Taaf-
p, 15 (1H84) (Mexico) .; Godm. & Salv., Bivl. Cnitr. Anier., Rhup. ii. p. 19G. n. t). t. 65. fi^. 7,
7.^, fold, genit. (18',)0) (Mexico to Costa Rica) ; Haase, Untersueh. Mindery p. 77. t. 9. fig. 62. J
(1893) ; Godm. & Salv., I.e. p. 728 (1901) (Guadalajara).
J?. Upperside metallic, especially tlie hindwing, which has a strong blue
gloss in male, being less strongly glossy in female ; scales of ujiperside feebly
dentate, the njiper scales partly entire in male, the same being the ca-^e in the
jiosterior area of the underside of the forewing ; no markings on forewing, e.xcept
the distinct white fringe-spots ; bnt one of our females has a row of indistinct
jiinkish grey jiatches about 5 mm. distant from distal margin, the row curving
costad in front, the tirst spot standing behind R' and the last before SM-, this
last sjiot being present in several other specimens ; this row of jiatclu s corresjionds
to the blue band of F.colunibus {= guiullachiuiius) ; posterior fringe-sj)ots souje-
times pink. Hindwing : two jiarallel rows of red spots, submarginal spots
strongly arched e.xcept njiper two or three and anal one ; the latter distinct on
uj>perside only in i'emale.
Genitalia : iS . Tenth tergite only a little longer than the sternite ; clasper
somewhat variable in outline, short, broad, obliquely emarginate-truncate, both
angles strongly rounded, dorsal edge somewhat incurved ; harpe mnch shorter
than the clasper, divided into two lobes, both rounded at ajie.x, one a[)ical, the
other basal and ventral, the latter smaller thnii the former, wliicli is somewhat
:-'J
( -144 )
c.nrvcd ventrad ; penis-sheath pointed, hut the apex a little twisted and cnrved
over the oritice, less strongl}' chitinised than in P. montezuma, atjacm, etc.
? . Anteyaginal ridge small, low ; postvaginal process short, broad, continued
laterad as a low carina ; anal sograeat with sliort stout blunt bristles veutrally.
Early stages not known.
Hab. East and West Mexi('o, southwards to Costa Rica.
In the Tring Museum, 54 S 6 , K) ? ?, from : Jiilai)a and Espinal, Vera Cruz
(W. Schaus) ; Patzcuaro, Michoacan ; Cuernavaca, July, 4000 ft. (A. Hall) ;
Clnantla, Morelos, June, 3800 ft. (A. Hall); Colima ; Guerrero (0. T. Baron) ;
Amatitlan, W. Guatemala, August, 4000 ft. (A. Hall) ; La Antigua, W. Guatemala,
August, 5200 ft. (A. Hall) ; Pozo Aznl, June, and Volcan de Miravalles, Costa Kic.a,
(Underwood).
9. Papilio alopius Godm. & Salv. (1800).
riijiiliii filnpiuit Gray, List Lep. Inx. Ilrit. Mna. i. Pop. p. 58. n. 231 (185(!) (Mexico ; nnm. imil.) ;
Godm. and Salv., Biol. Ceiilr. Amer., Rhnp. ii. p. 197. n. 12. t. r,.5. fig. 10. 11.? (1890) (Chiliualiua
and Durangn, Mexico ; J ? only) ; Haa.se, V idei-sw-h. Mimicry p. 78 (18'J3) ; Godm. and Salv.,
l.r. p. 728 (HiriJ) (Nicaragua).
c?S. Larger than P. p/totiniis, less glossy; spots of hindwing much smaller,
partly white, those of pro.ximal row very small, white or jiiukish wliite, the iqijier
ones obliterated, sometimes the whole row missing, tail sj)atulati'.
Genitalia: S- Claspor rounded at ape.x ; ventral lobe of harpc narrow,
]iointed. ? not dissected.
Hab. West Mexico ; Nicaragna.
lu the Tring Museum, 10 c?(?, 2 ? ?, from : Guadalajara, August and Cic.tober
(W. Hchans) ; Guerrero ((). T. Baron).
10. Papilio dares Hew. (IsfiT).
Papilio iliiiT.1 llewiUon, Triinx. Ent. Soc. Ltiiul. (.S). v. p. .501. n. 2 (18fi7) ( ? , Nicar.igiia) ; id.,
AVo(. liiitl. iv. Pi:p. t. 11. fig. 34. 9 (186'.l) ; Kirby, Cat. Diurn. Lip. p. WMi. n. 123 (1871)
(Nicaragua) ; Haase, Untei:':nrh. .Miniirri/ p. 77 (1893) ; Godm. and Salv., Biol. Ceiitr. Amfi:,
liliop. ii. p. 190. n. 10 (1890) (Nicaragua).
?. Only one specimen known. Resembling pbotit//i.i ; forewing with a white
dot R' — R^ ; iip]ier two sulimarginal spots of hindwing not curved, the foiiowing
three very slightly luuifurm, anal sjiot large. Femora scaled.
Ilab. Nicaragua.
11. Papilio montezuma Westw. (1842).
I'ajiiliii Nioiilnziimii Westwood, Arc. Ent. i. p. 07. t. 18. fig. 3. ^ (18-12) (Mi'xico) ; Douljl., List I.,rp.
Lis. Brit. Mils. i. p. 12 (1845) (VV. Mexico) ; id., Westw. and Hew., (leii. Diiini. Lip. i. p. 19.
n. 227(1847); Gray, Oi?. Lpp. Ins. Brit. .Mils. i. Pap. p. G5. n. 280 (1852) (West Caa5<t of
Ameriia) ; id., List Lrp. Ins. Brit. .Mus. i. Pap. p. 75. n. 803 (1850) (Yucatan ; Nicaragua ;
We.st Coast of Amer.) ; Weidem., Pror. Knt. Soc. Pliilad. ii. p. 147. (180!) (Mexico; Ceutr.
Amer.) ; Felder, Verb. Zool. Bot. Ges. Wien xiv. p. 294. n. 70 (18ii4) (Mexico; Chiapa* ;
= tulawi) ; Kirby, Cat. Diiirn. Lep. p. 5.30. n. 121 (1871) (Mexico) ; Obertb., /•,'/. iV Ent. iv.
p. 77. D. 250 (1880) (Mexico ; Yucatan) ; Godm. aud «alv., Biol. Ceiitr. Aiiiir., Rhop. ii. p. 197.
n. 11. t. 05. fig. 8. 8a. fold, genit. (1890) (Mexico to Nicaragua) ; Haase, Untcrsucli. .Mimicry
p. 78 (1893) ; Hoag, Eiit. News xiv. p. 320 (1903) (Altamira, Mex.).
Papilio perrliehus, Menetrie.s {non Boisduval, 1830, err. det.), Eiuiin. Corji. Aiiiin. .Mas. Pctrop.,
Lip. i. p. 5. n. 89 (1857) (Nicaragua).
Papilio tiilana Reakirt, i'/oc. Ent. Soc. Pliilad. ii. p. 140. n. 12 (1803) (Chiapas); Strecker, I^ep.
Shop. Uet., Suppl. iii. p. 17 (1900) (jj, type ; = montezuma).
c??. No markings on forewing, e.xcejjt the white fringe-spols, which are
often indistin<t. On liindwing a row of red sul/marginal sjmts, which are larger
( 445 )
in tlie ferartle than in the male ; on npperside the first three spots often absent
or vestigial ; the spots larger in sonthern specimens than in most individuals
from the more northern districts of the range, the tail being somewhat reduced
in the former. Legs characteristic. Femora naked, rough with dispersed con-
spicnons grannies, mostly beariug a bristle or thin hair. The bristles of the
tibiae and tarsi also situated on grannies, which are smaller than on the femora.
Spur of foretibia proximally of middle.
Genitalia : <S. (Jlasjier rounded at apex ; hari)e gradually tapering, ending in
a sharp point, almost reaching apical edge of clasper, the free apical half longi-
tudinally impressed. Penis-sheath essentially as in ayarxs, colinnhns, etc. Tentli
tergite about one-third longer than the lobes of the sternites, slender, pointed,
basally snbcarinate above. ?. Behind the vaginal orifice a short, broad process
about half as long again as it is l)road in middle, slightly sinuate at apex, convex
on proximal, concave on distal side ; a large lobe proximally of orifice, emarginate
in middle, rounded, finely hairy, its distal surface concave ; in non-virgin females
these organs concealed under a hardened coital substance.
Early stages not known.
Ilab. Mexico to Nicaragua.
In the Tring Museum, lUU cJc?, 48 ? ?, from : Colima ; Guadalajara, October
(W. Schaus) ; Fatzcuaro ; fluernavaca, end of August, September (Dr. Gadow);
Gnerrero (0. T. Baron) ; S. Jose, Guatemala, Sejitember (A. Ilall) ; 8. Pedro Sula,
Honduras.
II. Aeneas Group.
Marginal spots of hindwiug white (in klwfcsi vestigial). Palpus black or red.
Tibiae of c? incrassate and hairy, or non-incrassate and spinose as in ? . Scent-
organ woolly, or the scales at least elongate, the wool being white, brown-black or
tawny.
This is the largest gronp of Aristolochia-Swallowtails. Though some of the
species differ much from one another, the gronp does not appear to fall into sharply
defined divisions. We have not seen P. kahneli, but the figures published Liy
Standinger show it to be a near relative of P. triopas, which itself is very close to
chabrias. The great gap between kakiieli and aeneas is overbridged by a series of
more or less close allies : triopns, chabrias, quadratus, pizarro, coeUis, and
steiiibaclii.
The species of this group are i)artly very difficult to distinguish, there being
often very little in the outward appearance by wliich one can differentiate thein.
However, if some attention is paid to the structure of the tibiae of the males, the
colour of the palpi of both sexes, the extent of red at the apex of the abdomen of
the females, and the shape of the apex of the cell of the hiudwing, the reader will
generally be able to identify the species and mate the sexes correctly.
Key to tlie species : —
A. Palpus bia(;k (in drucei sometimes red).
a. Hindwing with yellowish white area which occuiiies at
least half the cell b.
Hindwing witii yellowish white area which stands outside
the cell or occiijiies only the apex of tlie same; no
white patch on forewing . .... c.
( 44C )
Hindwing with red patch or baud on npperside, or with
white patch in cell of fore wing ..... d.
b. Forewing with large basal patch of the same colour as the
median and subajiical bands ..... Species No. 12.
Forewing without basal jiatch ...... Species No. 13.
c. Hindwing below with red anal spot ..... Species No. 16.
No red anal spot on nnder.side of hindwing; forewing with
a row of postdiscal spots, sometimes absent in ? ,
yellowish patch of hindwing entering cell . . . Si)ecie8 No. 14.
No red anal spot on hindwing ; {brewing without post-
discal spots ; cell of hindwing entirely black, or only a
minnte creamy spot at apex Species No. 1 7.
<L Hindwing below with a red anal spot which is much more
distal than the dTscal spot which stands in front of it . e.
Last spot of underside of hindwing ou a level (or nearly)
with the next spot, or forewing with green ])atch. . /.
e. Forewing with diffuse white patch at apex of cell . . Species No. 15.
Forewing with sharply defined white patch posteriorly on
disc Species No. 18.
/. Tooth R^ of hindwing prolonged to a tail .... Species No. 21.
Tooth R' of hindwing not or very little more prominent
than the other teeth ....... g.
g. Fringe-sjiots of hindwing absent or vestigial ; abdomen
without red spots (? ; J not known) .... Species No. 19.
Fringe-spots of hindwing distinct ..... h.
h. Males .......... I.
Females .......... m.
i. Red area of npperside of hindwing l.irge, occupying at least
the apical fourth of the cell ; no green patch on forewing Species No. 22.
The red area occupying more than half the cell ; a green
patch on forewing ....... S()ecie8 No. 20.
A row of rounded or ovate spots on hindwing ; forewing
without green patch ....... Species No. 26.
Cell of hindwing black, a green patch on forewing . . j.
j. Abdominal fold black at base ; green patch of forewing
very large, glossy ; hindwing black above, with or
withont a red spot near anal angle .... k.
Alxlominal fold not black at base ; hindwing with a band
of red patches ; red spots of hindwing strongly
opalescent ......... I.
k. Green area of forewing entering cell Species No. 24.
Green area of forewing not entering cell .... Species No. 23.
I. Green patch of forewing reaching forward to R^, or narrow,
being at least twice as long as broad ; longest red spot
of hindwing three or four times as long as broad,
nsnally a red streak behind M- .... Species No. 27.
Green patch more or less triangular, about as broad
jKisteriorly as long, ofter broader; longest red spot of
hindwing al'ouL twice as Iwig as broad . . . Species No. 25.
( 447 )
m. Forewing without white patch, or the edges of the patch
washed out and the fringe-spots of the forewing indis-
tinct or absent n.
Forewing with sharply defined patches and distinct fringe-
spots q.
n. Hindwing with a row of widely separated ovate spots
which stand nearer the margin than cell . . . Species No. 26.
Red spots of hindwing merged together, or at least some
separated only by the black veins .... o.
0. Apex of cell of hindwing acute, vein D' being oblique and
longer than D* Species No. 22.
Apex of cell of hindwing truncate, vein D' being trans-
verse and shorter than D' . . . . . . p.
p. FriDge-sjwts of forewing distinct, no white on disc and
in cell .......... Species No. 27.
Fringe-spots absent, indistinct, or distinct bnt small, in
the last case always a white patch in middle of wing . Species No. 20.
q. White spot M'— IP of forewing smaller than spot R' — M',
often absent ; cell-spot large ..... Species No. 25.
White spot M' — M- of forewing larger than spot R' — M',
the latter often absent ; cell-spot small or absent,
usually a spot behind M- . . . . . . r.
r. A row of small white spots outside apex of cell . . . Species No. 24.
No such spots Species No. 23.
J3. Palpus red (often black in P. drucei).
a. Males b.
Females (the kc;/ should be consulted in connection with the
descriptions) ........ i.
b. Forewing without any trace of a green patch ; hindwing
with more than three red spots on upperside . . Species No. 33.
Forewing with green ])atch, or trace of it, rarely absent ; in
the latter case there is always a white dot on forewing c.
c. Tibiae and first tarsal segment somewhat incrassate, hairy . d.
Tibiae and first tarsal segment not incrassate, spinose as
in ? c.
d. Tooth H^ of hindwing mneli more prominent than the
others; forewing with large white patch; hindwing
with three rather small red spots, there being often a
minute fourth spot standing near the ai)ifal angle . Spscies No. 37.
Tooth R' of hindwing not or only a little more ])roiuinent
than the otlier teeth ; the red sj)ot behind M- on upper-
side of hindwing large, the red band triangular, con-
sisting of three spots which stand close together; no
white spots on forewing, or only a minute one ; green
patch not extending forward beyond M' . . . Species No. 30.
Tooth R' of hindwing not or slightly more prominent than
the others ; green patch of forewing narrow, or reaching
forward beyond M', or forewing with a large white spot
M' — M° or several spots, or red patch of bind wing small,
( 448 )
consisting of three spots and occasional!)' a small
fourth one which stands behind M- .
Species Nos. 34
and 3.7.
,Si)ecies No. 28.
/
Species No. 33.
9-
Species No. 31.
e. Bed patch on npperside of hiudwing small, elliptical, con-
sisting of two contignons spots ; spots on underside of
liindwing glancons cream-colour ....
Ilindwiug with at least three red spots on upjierside .
/. Fatcli of hindwing triangular, s]iot M- — (SM') large .
Spot M- — (SM') of hiudwing absent or small .
g. Hindwing with a red spot behind M- .
Hindwing without a red spot behind M-, or this spo'
minute ........
h. Spots on underside of hindwing almost nniforml}' red in
colour Sjiecies No. 29,
Spots on underside of hindwing partly pale jiink ; forewing
often with prominent white sjiot before M' or before
R', sometimes without green patch ....
i. Forewing without white patch
Forewing with white patch ......
Forewing with broad white band ; underside of hindwing
for the greater jiart white and pink ....
_;'. Band of hiudwing creamy white ......
Band of hindwing red ........
k. Hiudwing with a band of six red spots which stand nearer
the distal margin than the cell : ajiex of forewing very
little j)aler than base .......
Forewing semi-transparent in a]iical half; hindwing feebly
dentate .........
Forewing semi-transpareut in apical half; hindwing
strongly dentate ........
/. Forewing with greyish green patch before hindmargin
Forewing without greyish green patch ....
111. AVhite s]iot 11- — W of forewing larger than patch W — 51' .
'\Vhite spot R' — W smaller than R^ — M', often absent
n. Hindwing with tooth R' prominent, red spot C — SC^
usually jtresent, while spot SC" — R' is absent
Toolh R'^ of hindwing nut much more prominent tlian the
other teeth ; if spot C — S(J- is present, spot SC- — R' is
also marked .........
o. Spot M' — M- of forewing present, usually large .
Sj)ot M' — M" of forewing absent .....
p. Cell-iiatch of forewing very large, occupying at least one-
third of the cell
Cell-patch smaller ........
q. Red band of hindwing narrow and short, nfit extending
forward beyond R-, consisting of four or five spots only
which are more or less completely merged together
Red band of hindwing consisting of at least six spots
Species No.
( 449 )
>■. Cell-spot of forewing al)scut or small Species No. 30.
CelI-sj)ot of forewing large, extending across the cell (or
Dearly) g.
s. Cell-spot of forewing transverse, narrow, about twice as
long as broad Subspec.No.34«.
Cell-spot of forewing broad, nearly square, sometimes
triangular t,
t. Forewing semi-transparent in apical half .... Species Nos. 32
and 35.
Forewing not semi-transparent in apical half . . . Species No. 31.
12. Papilio hahneli Stand. (1882).
cJ ? . riqiHio luihiidi Staudlnger, Proc. Zoo/. Soc. Lnml p. 3flC. t. 24. fig. 1. (J (1882) (jrassauaiy) ;
id., Exot. T,t;,f. p. U). t. 1.3. ? (1884) ; Hahnel, Iris iii. p. 257 (18'JO) ^Massauary, below
Maue's) ; Haase, Unlersuch. Mimkrij i. p. 79 (1893).
S ? . Seses similar ; three broad bands of yellowish grey patches on forewing,
one subbasal, the second central, and the third subapical. Hindwing tailed ;
yellowish area occupying the greater part of the wing ; anal angle strongly
produced in male.
Hab. Manes, Amazons.
In coll. Staudinger.
13. Papilio triopas Godt. (1810).
Papilh, IrapuH Godart, Enc. Melh. ix. p. 33. n. 23 (1S19) ( ? , hab. ?) ; Grimsb., Trans. Ruy. Soc.
EiViih. xxxi.x. 1. No. 1. p. 7 (1807) (type in Mus. Edinburgh).
S ? . Palpus black. End-segment of antenna hardly as long as broad.
Forewing elongate, lower angle of cell very obtuse. Hindwing reduced in size,
tooth R^ prominent, cell acuminate, veins R- and R^ standing close together, D'
being very short. Pattern similar in the sexes, but tlie yellowish markings larger
in female than in male ; the subapical and central patches of the forewing
reminding one of the spots oi Papilio childrenae and the blue band of P. cohunhux.
Tibiae of mid- and hindlegs of male somewhat incrassate, hairy, and anal angle
of hindwing strongly produced. Abdomen of female with a red spot in front of
vaginal cavity and another behind it.
Scent-organ : fold large ; a line of broad buff scales on SM^, at the abdominal
side of which there is a broad clayish streak consisting of very small scales. These
scales very densely jiaeked together, erect, elongate-triangular, widest at apex, which
is somewhat rounded, being centrally produced into a thin thread : the base of each
scale also threadlike. This scent-organ, as regards the shape of the scales, represents
doubtless a less advanced type than the woolly scent-organ of other species.
Genitalia: S. Ninth tergite bearing at each side of base of tenth a conical,
spinelike tooth. Harpe narrow, curved, distally dilated, the apical portion
triangular, short, ventral edge denticulate. ?. Anal segment ventrally with
numerous thin gradually tapering bristles ; edge of vaginal aperture raised, the
proximal wall of this short funnel tawny, smooth ; behind the ai)ertuie and
covering the vaginal cavity a rounded plate wliieh is concave on its distal surface ;
the edge ofthi-; plate dilated to a short rounded process which curves distud.
Early stages not known.
llab. The Guianas and Lower Amanons.
Two subspecies.
( 450 )
a. P. triopas triopas Godt. (1S19).
Papilin IrinjHin Godart, I.e.; Boisd., Spec. Ghi. Up. i. p. 313. n. 151 (1830) (Cayenne) ; Lucas, in
Gu^r., Dirt. Pill, flist. Xal. vii. p. 48 (1838) ; Doubl., Westw. and Hew., Geti. Diiirn. L,:p. ii.
p. .Wil (1852) ; Gray, Oit. Lep. Ins. Brit. ilus. i. Pap. p. Ci'J. n. 314 (1852) {pnrlim ; Amazons) ;
Wall., Traim. Enl. Sor. Lmul. (2). ii. p. 255 (18'>4) (Para, and all over the Amazons) ; Lucas, in
Chenu, Enc. Hist. A'a(., Pap. i. p. 38. t. 4. fig. 1. ^ (1851-53) (Cayenne) ; Gray, /.;»( Lfp. I„>.
Bill. Mux. i. Pap. p. 80. n. 331 (1856) (partiiit \ Para ; Amazons) ; Bates, Trans. Ent. Sue.
Lond. (2). V. p. .343, .3.58 (18C1) (Guiana ; Lower Amazons) ; id., Juurn. Enlum. i. p. 226. n. 1(3
(18i'>2) (Lower Amazons and Pard, forest) ; Felder, Vi'rii. Zool. But. Gen. Wini xiv. p. 2D2.
n. 3G (18i;4) ; Kirby, Cat. niia-n. Lep. p. 528. n. fiG (1871) (Guiana; Lower Amazons);
Miischl., Vn-h. Znul. But. Gr.i. Wirn xxvii. p. 2i»5 (187G) (Surinam) ; Oberth., Et. il' Enl. iv.
p. 117. n. 2115 (1880) (.Guyane ; Para) ; Stand., Exot. Tatff. p. 14. t. 9. ^ (1884) (Guiana ;
Lower Amazons) ; Haase, Untersiioh. .)IiinkTi/ i. p. 79 (1893).
Asmiii<le-< li-iapiis, Hiibner, Samml. Exot. Schmetl., ZntrUqe v. p. 32. n. 4i;5. fig. 929. 930. J (1837) ;
Kirby, in Allen's Nut. Libr., Lep. Butt. ii. p. 2G9 (189G) (Guiana ; Lower Amazons).
Pajii/io triiipiis (!), Munetries, Enum. Corp. An'un. .Mm. Petrop., Lep. ii. p. 69. n. 1134 (1857)
(Cayenne).
cj?. Tlie markings of fore- and hindwing variable in size and number.
Forewing of male nsnally with one large and two small snbapical spots, the small
ones being often vestigial on npperside, the one in subcostal fork more often (juite
obliterated; male mostly without cell-spot, while all the females have an elongate
cell-spot.
llab. French and Dutch Guiana ; Amazons, from Para to Obidos.
In the Tring Museum 10 SS, 9 ¥?, from: Cayeune; Igarape, Para
(W. Hoffmanns); Obidos, October and November 1004 (M. de Mathan).
In coll. Oberthiir a small series from Mai'oui, Guiana, and Para.
b. P. triopas mWd-as Grose-Smith (1902).
Papilin trinjxis. Gray, II. ec. (Demerara).
Fapilio mithras Grose-Smith, Rluip. Exot. iii. P((p. t. 23. fig. 1. ^J (1902) (British Guiana).
t? ? . Sp(Tt8 paler and smaller than in the j)receding, the last spot of the
hindwing, above, especially smaller. The ditfereuces may not be constant. We
have seen only a few sjiecimeus.
Nab. British Guiana.
In the Tring Mnsenra 1 ? from Bartica, British Guiana, 22. v. 1004
(R. Haiusch).
14. Papilio chabrias Hew. (1852).
Papitio chahriati Hewitson, Tranfi. Ent. Soc. Land. (2). ii. p. 2.3. t. G. fig. l.$(1852) (Amazons) ;
Doubl., Westw. & Hew., Gen. Diuni. Lep. i. p. 20. n. 247 (1847) (Cayenne ; N. Brazil) ; Gray,
Cat. Lep. //<.-■. Brit. .l/«.v. i. Pap. p. 69. u. 315 (1852) (Ega) ; Wall., Tra?is. Ent. Sor. Lnnd. (2).
ii. p. 255 (1H54) (Upper Amazons ; forest) ; Gray, List Lep Jn«. Brit. Mnx.'x. Pap. p. 80. n. 332
(ISoG) (Ega) ; Bate.s, Trana. Ent. Soc. Lnnd. (2). v. p. .343. 358 (ISGI) (Upper Amazons, J' ? ;
subspecies of trioiiax) ; id., Juurn. Entnni. i. p. 226. n. 17 (1862) (Upper Amazons; "local
niodili(^iti<m " of trinpnx) ; Felder, Vrrh. Zool. Bot. Gex. Wien .xiv. p. 292. n. 37(1864) ; Oberth.,
Kl. d'Eiit. iv. p. 9G. n. 294 (1880) (Teffe ; " Para " fal.se) ; Kirby, Cat. Diurn. Lep. p. 528.
n. GGa (1871) (Upper Amazons) ; Hopff., Stitl. Ent. Zeit. xl. p. 51. n. 10 (1879) (Amazons,
Peru) ; Staud.. Ejeot. Tag_f. p. 14 (1884) (Upper Amazons) ; Hahnel, L-is iii. p. 307 (1890)
(Juriniitgua-s) ; Haase, Uiiterxmh. Mimiciij i. p 79 (1893) ; Michael, Iris vii. p. 214 (1894) (Sao
Paulo de Oliven^a).
Papilio n;/niplia.i Grose-Smith, Riiop. E.rat. iii. Paj). t. 23. fig. 2. 5(1902) (Ecuador ; "Para" false).
cj ¥ . In structure like triopas. Forewing of both sexes with a submarginal
row of spots, wliicli are often missing in female inde])endently of locality. Patch of
hindwing more distal tban in trioija.i, therefore the s])ots around ape.x of cell larger,
while ihe cell-spot is smaller.
(451 )
Hab. Upper Amazons, from Ega to the eastern slopes of Ecuador and Pern.
In the Tring Museum 5 6 S, 6 ? ? , from : R. Chuchuras, affluent of
R. Palcazu, Ilnunuco, Pern, 320 m. (W. Hoifmanns) ; R. Cachyaco, affluent of
R. Huallaga, Peru (Maxwell Stuart) ; Iquitos ; S. Paulo de Olivenoa.
In coll. Oberthiir a series from Mo3'obamba ; S. Paulo de 01iven(;a ; Tocantins ;
Teffe.
15. PapiUo coelus Boisd. (1836). (PL IV. fig. 2).
?. PapUin r.nelus'Bo\i'Au\a.\, Sjuv. Gen. Lei), i- P- 289. n. 117 (1836) (Cayenne) ; Donbl., Westnr.
and Hew., Geii.Diurn. Lep. i. p. 18. n. 209 (1846) ; Gray, Cat. Lq). Ins. Brit. Mus. i. Pnp. p. 53
n. 2o3 (1852) ; id , Lht Lq>. Inx. Bril. Mns. i. Pup. p. 66. n. 268 (1856).
5 . Papilin vertummis var., Bates, Trans. Eut. Sue. Loud. (2). v. p. 340, 355 (1861).
j. Papilw verUwmus, Felder, Verh. Zool. Bot. Ges. Wieii xW. p. 292. n. 42 (1864) (partini).
9 . PapiUo verliim/ms var. d. P. roeliis, Kirby, Cut. Diurii. Lep. p. 525. sub n. 61 (1871).
(J. PapiUo verciwjetorij- Oberthiir, El. d'Eiit. xii. p. 5. n. 10. t. 7. fig. 51 (1888) (French Guiana).
S ? . Palpus black. Abdomen in male with a tiny red dot before claspers,
in female with a red spot around vaginal cavity, no red scales dorsally at apex of
eighth tergite. Sexes similar in pattern ; forewing with a wliite patch in apex
of cell extending on to disc, larger in female than in male, washed out at the
edges. A red patch or band on hindwing close to cell, consisting in male of
fonr spots, in female of six, in this sex the band being continued to abdominal
edge ; tooth R' prominent, almost pointed, the following two teeth also somewhat
pointed.
On umleniide the white patch of forewing a little larger than above. Red
band of hindwing paler red than above and the spots in female smaller, in male
two additional spots between M^ and anal angle.
Scent-urgan white.
Hab. French Guiana.
Only two specimens known, the type (?) of coelus being in Mus. Paris, and
the type (c?) of vercingetorix in coll. Charles Oberthiir.
16. PapiUo quadratus Stand. (189U).
Papilin quadratus Staudinger, h-is iii. p. 337. t. .3. fig. 1. (J (1890) (Manicore'); id.. I.e. iv. p. 61 (1891).
cJ ? . Sexes similar. Palpus black. Mid- and hindtibiae of male slightly
dilated, hairy, but also bearing many bristles. , Abdomen beneath with minute red
sjiot before claspers in male, and with large spot around vaginal cavity in female,
no red scales dorsiiJly at aj)ic!il edge of eighth segment. Forewing very long,
costal and distal margins almost parallel in male. Hindwing with a creamy patch
consisting of five discal spots and (in S) a, small cell spot ; tooth R^ somewhat
projecting in female, while the wing is strongly rounded at apex, being produced
anally.
On uiulersitle, the hindwing boars a red spot close to anal angle.
Scent-organ with a very broad stripe of white wool ; this stripe not quite
extending to base, where there is a brush of white spreading hairs.
Genitalia : (?. Ninth tergite with a conical spinelike jn-ocess at each side of
base of tenth tergite. Harpe half the length of the clasj)er, bearing four teeth,
all projecting ventrad and curved inward, the apical one very long, the next two
shorter, the most proximal one being minute.
Early stages not known.
Jltih. Up{ier Amazons.
Two subsj)ecies.
( 452 )
a. r. quadratm quadratus Stand. (1891).
jj. PnpiVio qiKulratiiit Staudinger, U.cr.
Forewing with yellow quadrate spot M' — M^ ia male. The female not kuown.
This may be only an aberrant individnai.
Uab. Mauicore, Rio Madeira.
b. I', quadratus spoUatus Stand. (1898).
(J J . Papilio qtiadratiia var. spoUatus id., l.c. xi. p. 138 (1898) (Sao Paulo de Oliven^a ; Pebas ;
Iquitos) ; Groae-Smith, Rhop. Exut. Ui. P'ip. t. 17. fig. 1 (J, 2. ? (189'J) ; SUud., l.c. xi. p. 370
(i8ini).
c??. Forewing withont markings, or the spot M' — M- vestigial above and
distinct thongh small lielow.
JJiili. U{)per Amazons.
In the Tring Mnseum 2 (?(?, 1 ? , from : R. Jurna; S. Paulo de Olivenva.
In coll. H. J. Adams 6 c?<?, 4 ? ?, from the Ui>iier Amazons.
17. Papilio pizarro Stand. (1884).
(J ?. Papilio pizmro Staudinger, Ej-nl. Tnr/f. p. 18. t. 13. (J (1884) (Sao Paulo de 01iven(,-a, Pcbas,
Jurimaguas; $ parti m) ; Hahnel, Iris iii. p. 286 (189U) (Pebas); id., i.e. p. 307 (1890)
(Jurimiiguas) ; Haase, Unlersuch. ilimicry i. p. 79 (1893) ; SUud., Iriis xi. p. 141 (1898).
(J ? . Palpns black. Abdomen of male withont red scales before claspers, of
female with red spot behind vaginal cavity and at ape.\' of seventh sternite, bat
no (or very few) red scales dorsally and laterally at apex of eighth segment, in
the female of P. cutorina this segment being edged with red. Forewing without
white fringe-spots, no markings on disc. Hindwing with creamy patch consisting
of three or fonr spots in male, of three to six in female ; D' short, transverse, much
shorter than D^.
Mid- and hindtibiae of male slightly incrassate, hairy ; foretibia thinner,
spinose, with comparatively few short hairs.
Scent-organ grey, with a streak of small erect scales as in P. rliabrias and
steinbachi.
Ilab. Upper Amazons.
In coll. Oberthiir from : f 'hamliiriyaeu ; Tarapoto ; Pebas.
In coll, H. Grose-Smith from Ynrinnignas ; and in cull. II. J. Adams from the
Upper Amazons.
As Dr. Staudinger, Lis .xi. p. 141 (1808), when cnrapariiig the females of
I'. jAzarro, bolivar, and cutorina with one another, did not tiiid any constant
character, except size, by which to separate bolirar and cutorina, and mentions
as a distinguishing character between cutorina. and pizarro, besides the greater
expanse of cutorina, otily a creamy spot situated in cutorina on the underside of
the hindwing at the abdominal margin, which character is not constant, we append
here a short exposition of those external charat;ters by which the females of the
three insects can be recognised. In the Godman collection there is a female
received from Staudinger as jnzarro, being labelled " original." Thi.s female is
bolivar.
1. Palpus black ; a red spot behind vaginal cavity and in front of it, no red
scales at apex of eighth segment dorsally and laterally ; fringe of forewing quite
black or with a white scale here and there, not bearing distinct spots ; cell of
hindwing narrow, vein D^ transverse, mucii shorter than D^ ; s[iot M- — (!SM')
absent or small : Female oi pizarro.
( ^5:5 )
2. Paljins and apex ofabdomen as before, or the latter more extended red ;
fringe of forewing with more or less distinct white spots ; lower angle of cell of
hindwing acute, D' obliqne, as long as D\ being usually longer, rarely shorter ; spot
M-— (SM') about half the size of sjxit M'— M- : Female of bolimr.
3. Palpns red ; ajiex of eighth abdominal segment red all round, the red
scaling forming a ring around the anal segment ; fringe of forewing with sharply
marked white spots ; D' of hindwing shorter than D^ : Female oi cutoiiiM.
18. Papilio steinbachi Rothsch. (1905). (Plate IV. fig. 5. c?, G. ?).
$ ? . Papilio iteinliac/u Rothschild, Enlum. p. 125 (1905).
S ? . Palpus black. Abdomen of male without red scaling before claspers ;
of female with red scaling around vaginal cavity, but not dorsally at apex of eighth
segment. Tibiae of male somewhat dilated and hairy. Forewing in male with
white or huffish double spot M" — (SM') ; in female with a larger white patch
consisting of a minute cell-streak, a very large spot M' — M^, a triangular spot
before M' and a streak behind M^. Hindwing with red patch in both sexes, the
last spots being more or less white in male, this colour only indicated in female.
On underside both sexes with a red spot close to anal angle, separated from
the discal spots.
Scent-organ : fold grey inside ; a broad stripe of small scales as in P. triopas,
the scales of the same type as in that species, but larger.
Genitalia : S . Ninth tergite with conical lateral process as in the allied species.
Harpe about two-thirds the length of the clasper, curved, narrow, with short but
strong teeth from base to apex.
Early stages not known.
Hab. Eastern Bolivia.
In the Tring Museum b S <S , 1 ? from : Sara, S. Cruz de la Sierra, end of
February to June 1U04 (.J. Steinbach); Mapiri.
19. Papilio klagesi Ehrm. (1904). (PI. V. fig. 20).
?. Pajjilio kliigesi Ehrman, Ent. Xncs xv. p. 215 (1904) (Suapure, Venezuela).
The male is not known.
?. Paljius and apex of abdomen black, no red spot behind or in front of vaginal
cavity. Fringe of forewing (juite black, lower angle of cell completely rounded
oft'; a white band from M' to SM-. Hindwing: fringe-spots white, vestigial or
entirely absent, this being the only case in the j)resent group of Ariisfoloc/i/a-
Papilios of the fringe of the hindwing not being distinctly spotted ; tooth R'
slightly prominent, while the others are very obtuse and short ; band of discal
spots somewhat jiinkish, njiper two spots small, more distal than the last four,
wiiiih are contiguous ; M- originating almost on a level with R', D' shorter than
1)*, transverse, D^ nearly as long as upper partition of M.
Anal segment ventrally with very numerous short stiflf yellowish bristles. In
front of vaginal cavity a low ridge continued distad on each side of the cavity; the
Jiroximal j)ortiL,n of this ridge densely clothed with very short stift" luxirs ; behind
the vaginal aperture a long smooth process, convex on proximal side, excavate on
distal side, rounded at apex.
■Hab. Caura River, Orinoco.
In the Tring Museum 3 ? ? from Suapure, Caura R., February and March
1S90 (S. M. Klages).
( 454 )
20. Papilio aeneas L. (1758).
(J. Pnpilin Eqiies Trrjaiiiin neyimx Linne, Si/nt. Nat. ed. x. p. 461. n. 15 (1758) ("Asia";.
5 . Prinr.ppK domhianf: marcius Hiibner, Samml.' Exot. Sckmetl. i. t. 122 (180fi — ?).
(J ? . rajiilin aeneas, Erichson, in Schomb., F. F. Brit. Guiana p. 593 (1848) ( ? = nwrrlus).
c??. Palpns black; posterior abdominal segments of male withont red
markings, there being in the female a restricted red spot situated behind vaginal
cavity, neither the seventh nor eighth segment as a rnle bearing any red scales
at the edge. Fringe of forewing black, showing rarely vestiges of white spots in
female. Mid- and hiudtibiae of male densely hairy, except base, incrassate, especially
the hiudtibia, foretibia and tarsi simply spinose. Scales of red patch of hindwing
of male above entire ; in female the scales also rounded at apex, but seldom entire,
bearing from three to five teeth, very few scales being only bidentate ; on the
underside the white scales covering the red ones nearly all sharply tridentate
in female, only a few having two teeth, while in male the majority is bidentate.
No opalescent gloss on upper side of hindwiug. White patch on forewing of
female washed out at edges.
Neuratiou : D^ of hindwing about as long as D^, often considerably longer,
however in one of our males much shorter.
Scent-organ : wool blackish hair-brown.
Genitalia : <S. Tenth tergite a very little longer than the sternite ; aj)ical edge
of ninth tergite somewhat dilated near the tenth tergite and denticulate. Harpe
reaching close to apex of clasper, bearing at the apex from two to six heavy teeth,
and at the ventral edge beyond the middle one more tooth, the distal portion of
the ventral edge being sometimes minutely denticulate. ¥. No strongly elevate
ridge in front of the vaginal cavity ; posterior edge of cavity rounded, slightly
incrassate, partly covering the cavity, rest of postvagiual area membranous,
transversely wrinkled.
Early stages not known.
Ilab. The Guianas ; Amazons from Para upwards ; eastern slopes of Andes of
Pern and Bolivia ; Upper Orinoco.
a. P. aeneas aeneas L. (1758).
Roesel, Jns.-Behisl. iv. B. p. 24. t. 2. fig. 2. c? (1755).
(J. rajiUio Eques Trojamis aeneas Linn^, I.e.. ("Asia"; citat. jKirtiin) ; Kleem., in Roe-iul, l.c.
(ed. ii., 1761) (Surinam) ; Linne, J/»w. Lml. Vlr. p. 197. n. 16 (I7(U) ; Houtt, .\,itiirl. Hint. i.
11. p. 198. n. 15 (17ri7) ; id., Sy^l. Nat. ed. xii. p. 747. n. Iti (1767) ; Fabr., Si/sl. Eui. p. 448.
n. 23 (1775) ('India") ; Goeze, Ent. Beylr. iii. 1. p. 36. n. 16 (1779)(cit. Sebac excl.) ; Cramer,
Fnp. Emt. iii. p. 1,56. t. 279. fig. C. D (1780) (Surinam) ; Fabr., S^pec. Fm. ii. p. 8. n. 32 (1781)
(citat. Cram, excl.) ; Jabl. and Herbst, Natiirx. ffrhniett. ii. p. 53. n. 19. t. 9. fig. 5, 6 (1784)
(jMrlim) ; Fabr., Mmil. Iim. ii. p. 6. n. 35 (1787) ; Esper, Ami. Srhmi-U. p. 40. n. 15, p. 60(1788)
(parlim, ? ) ; Gmelin, Syt. Nal. i. 5. p. 2233 n. 16 (1790) (pnrlim) ; Jung, Aljilial,. Vcrz. p. 11
(1791) (synon.;rar^m) ; Fabr., Ent. Si/xt. iii. 1. p. 17. n. 50 (1793) (,eha.t. jKirlim).
FapiUo Eqiies aenms Linnn, Syst. Nat. ed. Lange p. 461. n. 15 (1760).
Papilin (Tros) ae.nens, Miiller, Natut-s. v. 1. p. 570. n. 16 (1774).
Fapilin (neneng), Meerburgh, Afb. Zelch. Gen: t. 19. ^ (1775).
Papilin Eqiies Trojanun aew.ides Esper, AuhI. Schmell. t. 15. fig. 4 (1788).
Paridex gargasus Hubner, Vers, bel: Silnnetl. p. 87. n. 909 (1818?) (partim ; nmn. novum hic.
"aeneas Cram. 279. A— D'').
Papilin apneas, Godart, En<: Melh. ix. p. 33. n. 24 (1819) {partim ; " ? " only) ; Boisd., f^pee. Gen.
Up. i. p. 286. n. 112 (IS.^fi) (partim; " $ "only) ; Constable, Miseell. Bnlt. p. 140. t. 13. $
(1832) ("Cochin China") ; Doubl., List Lep. hm. Brit. Mns. i. p. 12 (1845) (Demerara ; »yn.
partim) ; id., Westw. and Hew., Gen. Diurn. hep. i. p. 18. n. 202 (1846) (Guiana ; cit. Cram.
partim); Erichs, in Schomb., /". B\ Brit. Guiana p. 593(1848); Guy. List Lep. Ins. Brit.
( 455 )
jVms. i. Pnp. p. (i5. n. 2G4 (185fi) (jmrtim. ; Demerara) ; Bates, Trans. Ent. Snc. Lnml. (2). v.
p. 342, 357 (1861) (partim ; Guiana) ; Felder, Ve.rh. Zool. Bot. Ges. Wieii x\v. p. 295. n. 80
(1864) {partini ; Surinam ; Guiana) ; Guenee, Ann. Soc. Ent. France p. 305 (1867) (synonjmy);
Bntler, Cat. Diuni. Lep. descr. Fabric, p. 236. n. 11 (1869) (Demerara ; synon. par tim) \ Kirby,
Cnl.Diurn. Lep. p. 528. n. 63 (1871) (partim ; Guiana) ; Moschl., Verh. Zool. Bot. Ges. Wienxx-vu.
p. 295 (1876) (Surinam) ; Auriv., K. Si'. Vet. Ak. Handl. xix. 6. p. 21. n. 16 (1882) (Recrnsio
crllica) ; Mijschl., I.e. xxxii. p. 303 (1883) (Surinam) ; Haase, Untcrsiifh. Mimicnj i. p. 79 (1893).
Papilio marciiK, Doubleday, Lv<t Lep. //«. Brit. Mus. i. p. 12 (1845) (Demerara).
(J. Papilio hochus Lucas, Rev. Zool. (2). iv. p. 191 (1852) (Cayenne).
(J. Green spot distant from cell, usually shortest in basi-distal direction,
mostly shorter behind SM^ than before this vein. Spots on underside of hindwing
red, separate, spot M^— (SM') usually the largest.
? . Dichromatic, the forewiiig being sometimes without the usual white patch.
Red band of hindwing distant from cell, the two spots M' — SM^ confluent, or at
lea.st standing close together.
a. ? -f. specularis nov. Forewing : A large white spot W — M', a smaller
one in cell, and usually a small spot before W and another behind M'. The
ordinary female.
b'. ?-f. f//(/(? nov. Forewing: No white patch. A rare form.
IJab. of P. aeneas aeneas : The Guianas.
In the Tring Museum IG S S, 15 ? ? from : Cayenne ; Surinam ; Rio Demerara,
Essequibo K., and Camaria (January 1904, R. Haensch), Brit. Guiana.
h. P. ae/ieas marciits Hiibn. (1806 — ?).
$ . Princeps doiniiiaits marriiis Hiibner, Saiiiinl. Erot. Srhiiieft. i. t. 122 (1806 — ?).
J. Priamides marciits id., Verz. belc. Schinett. p. 87. n. 900 (1818?) (syn. excl.).
J. Pajiilio mariiuK, Boisduval, Spec. Gen. Lep. i. p. 288. n. 115 (1836) ; Gray, Cat. Lep. Ins. Brit.
Mas. i. Pap. p. 55. n. 259 (1856) (Para) ; Wallace, Trans. Ent. Soc. Land. (2). ii. p. 255 (1854)
(Amazons ; forest) ; Gray, List Lep. Ins. Brit. Mits. \. Pap. p. 67. n. 274 (1856) (Para).
} . Papilio echelus, Doubleday, Westw. & Hew. (k»« Hiibner, 1806 — ?, err. det.). Gen. Diurn. Lep.
i. p. 18. n. 210 (1H46) (partim).
$ . Papilio parsodes Gray, Cat. Lep. Ins. Brit. Miis. i. Pup. p. 54. n. 256. t. 8. fig. 4. $ (1852)
(partim, ? only ; Pari).
£j. Papilio aeneas, id., I.e. p. 52. n. 249 (1852) (syn. excl. ; Para) ; Wall., Trans. Ent. Soc. Land. (2).
ii. p. 2.56 (1854) (Pari; forest) ; Gray, List Lep. Ins. Brit. Mus. i. Pap. p. 65. n. 264 (1856)
{partim ; Para) ; Bates, Trans. Ent. Soc. Land. (2). v. p. 342, 357 (1861) {partim ; Pari) ; id.,
Joiirn. Entnm. i. p. 226. n. 14 (1802) (partim ; Para) ; Felder, Verh. Zool. Bot. Ges. Wien xiv.
p. 295. n. 80 (1864) (partim; Para); Bates, Natural. Rir. Amaz. p. 26 (1864) (Pari, cj in
swampy shades, $ iu more open places) ; Guen., Ann. Soc. Ent. France p. 307 (1867) ; Kirby,
Cut. Diurn. Lep. p. 528. n. 63 (1871) (partim ; Pari) ; Oberth., Et. d'Ent. it. p. 95. n. 291
(1880)(c}= bochtis, J = imirt/us — jiarsodes ; Para; Amazons, //(jriim) ; Hahnel, /lisiii. p. 212
(1899) (Pari).
Pajiilio hjsander local var. parsmles, Bates, Trans. Ent. Soc. Land. (2). v. p. 344 (1861) (5 sub
synon.).
Parides aeneas, Kirby, in Hiibn., Samml. Exot. Schinett. ed. ii. p. 90. t. 122. fig. 3, 4 (190 — V).
c?. As in P. a. aeneas, last but one spot on underside of hindwing rather
smaller.
?. White jiatch of forewing usually reaching backwards to M-, the posterior
spot being larger tliau in the preceding. The central spots of band of hindwing
larger and closer together, spot C — SC^ present.
J/ab. Lower Amazons : Para district, probably found farther south on the
Tocantius.
Ill the Triug Museum o cJcJ, 3 ??, from: Para (Stuart) ; Igarape (VV.
llollmiiiiiisj.
( 45C )
c. p. a/jneas linvs snbsp. nov. (PI. V. fig. 28).
(J. Similar to the preceding ; spots 11^ — M^ of nnderside of Liudwiiig jialer,
close together, nearer to cell, spot R'— M' longer than the distance of this spot
from cell.
?. White patch of forewing .is in P. a. aeneas, consisting of a large spot
R^ — M', a smaller cell-sjjot, a minute spot R^ — W, and a small streak behind
M'. Red band of hindwing broad, extending from R' to abdominal margin,
touching cell or nearl}', spot R' — M' the longest, about four times as long as broad,
the spots close together above, the last one standing a little separate ; on underside
the first spot minute, sejiarate, the last small, also separate, the four others nearly
as large as above.
Hah. Middle Amazons : Santarem ; Obidos (Ji/pe) ; Massauarv.
Bates did not meet with aeneas on the Middle Amazons.
In the Tring Museum ](?,!? from Obidos and Santarem.
In coll. Oberthiir from Massauary.
d. P. aeneas damis subsp. nov.
fapillo aeneas, Druce, Proc. Zool. Sdc. Lrmd. p. 24.'). n. 4 (187G) (Ucayali).
(?. Green patch of forewing larger than in the three preceding forms,
touching cell in most specimens, extending basad to SM' or even beyond. Spots
R^ — M^ of underside of hindwing pinkish white, nearer the cell than in /'. a. aeneas
and marcius and closer together, spot M' — M'' sometimes obsolescent.
?. Dichromatic.
«'. ?-f. ji>/romelas nov. (PI. VI. fig. 37). — This is the usnal form. Forewing
without white patch, deeper black than in the other subspecies, with vestiges of white
fringe-spots. Hindwing : red baud broad, much broader than the black distal
area, the sjiots close together, tlie central ones touching cell as in female of litws, or
the band 3 or 4 mm. distant from cell, in this case its inner edge being concave ;
no spot before R' ; spot at abdominal margin (the si.xth) separate, distinct or
vestigial or absent; scales of red band either mostly entire or finely denticulate;
the band narrower on underside, sixth (last) spot small or .absent. Tliis female
diil'ers from that of 7'. dnieei especially in cro'ss-voin D-' of iiindvving being at least
as long as D^, and in the apex of the abdomen bearing a red spot only behind
vaginal cavity, there being no red scales at the edges of segments 7 and 8 ; tlie
forewing is also more opaque.
b'. ?-f. eucltaria nov. (PI. YI. fig. 35). Apparently a very rare form,
which we thougiit first to represent a distinct species. Besides tlie spccitnen
figured (from the Peren6 River) we have seen only one other, which is in the
collection of Mr. H. J. Adams, being also found in the Perene valley (5000 ft.,
May — June 1902, W. Watkins). Forewing with large white patcii. Eiglith
abdominal segment more extended red than in other females of /'. aeneas.
Hab. of P. a. damis : East Peru.
In the Tring Museum : 5 J J, 9 ? S, from : Poznzu, Hndnuco, 8U0— lOOO m.
(W. IloU'nianns) ; R. Chuchuras, Hnaimco, 320 m. (\V. Holi'miinns) ; R. Perene,
March 1900 (Simons) ; La Merced, Clinnclianuiyo, May and dune 1903, 2500 ft;.
(Watkins & Tomlinson) ; La Union, R. Huacamayo, C'arabaya, 2000 ft., November
1904 to January 1905 (O. Ockenden) ; Paehilea.
( 457 )
e. P. aeiieas locris subsp. nov. (PI. VI. fig. 20).
<S. Like (/amis, red area of hindwiug rather larger, and spot M' — M'' of
underside of hindwing mostly obliterated.
?. Similar to the Gniana ?-f. specularis. Forewing deeper black, the
apical area being less transparent ; cell-patch larger, extending from M' to lower
aiigle, reaching ncarlj- across cell, the spots rather more sharply- defined and spot
R' — M' longer; fringe with or without minute white spots. A row of six
spots on hindwing, standing nearer cell tlian onter margin, bnt being ijnite remote
from cell, all separate, spot li^ — M' tlie largest ; fifth spot smaller than first,
and the sixth vestigial, both small on underside and separate.
Hub. Bolivia.
In the Tring Mnsenm : 'i i S , 2 ? ?, from : Mapiri ; Province Sara, S. Cruz
de la Sierra, April — June lOiU (J. Steinbach).
/. P. aeneas bolivar Hew. (I85i>).
PapiHo hiVirnr Hewitson, Tram. Ent. Sue. Loud. (2). i. p. 97. t. 10. fig. 2. J (1850) (.imazons) ;
Doiibl., Wastw. and Hew., Gen. D'mrii. Up. ii. p. .529 (1852) ; Gray, Cat. Lep. Im. Brit. Mas. i.
P«p. p. 57. n. 265. t. 10. fig. 7. ? (1S52) (E^a) ; Wall., Trans. Ent. Soc. Lund. (2). p. 256.
(1854) (Upper Amazons; forest) ; Gray, List Lep. Ins. Brit Mas. i. Pap. p. 69. n. 280 (1856)
(Ega) ; Bates, Trans. Ent. So/;. Land. (2). v. p. 342, 357 (1861) (Upper Amazons) ; id., Juarn.
A'«/";/i. I. p. 226. n. 15 (1862) (Upper Amazons, abundant, "local modification " of a«ncn«) ;
Felder, Vtrh.Z„<A. B„t. Ges. )!>« xiv. p. 29.5. n. 81 (1864) (Ega) ; Kirby, Cat. Diurn. Lep.
p. 528. n. G3a (1871) (U|iper Amazons); Hopff., Stett. Ent. Zcit. xl. p. 50. n. 7 (1879;
(" Brazil '•) ; Oberth., El. tV Ent. iv. p. 96, 117. n. 2:i3 (1880) (Teffe ; Sao Paulo de Olivenva ;
Pebiis ; Iquitos) ; Staud., Ej-nt. Tagf. p. 19 (1884) ; Hahnel, Iris iii. p. 275 (1890) (S. Paulo
deOlivenyi) ; id., I.e. p. .307 (1889) (Jurimaguas) ; Haase, Untersueh. Mimicry i. p. 79 (1893) ;
Michael, Iris vii. p. 214 (1894) (Sao Paulo de Olivensa).
Hates, I.e., considered this to be the Upper Amazonian form of P. aeneas, in
which he was doubtless right. The differences, thongh consjiicnous enough, are
not very trenchant.
S. Green ])ateli of forewing as in the Peruvian subspecies, spot M- — SM^
proxiiually rather more reduced than in most Peruvian and Bolivian males. lied
area of hindwing deeper red and smaller, the spots around the cell being reduced ;
these spfits without bright red tijis, or oidy spot R^ — M' with small red dot at
apex; fringe with small wi}it,e sjiots. On underside all the spots of hindwing
cream-colour, one or the other with a few red scales at the edge, spot M- — SM^
much the largest.
?. Forewing without wliite patcli ; fringe with white spots. Hindwing
with tlie baud cream-colour injstead of red, last sjiot separate, often absent ; width
of band vaiiiilile.
Hub. Upper Amazons ; Orinoco.
Iii the Tring Museum : 8 c?c?, 5 ? ? , from : Maipures, Orinoco, December 1S98
(("henie) ; R. (^achyaco, affluent of R. lluallaga (Stuart); S. Paulo de 01iven(,'a ;
Tefle, January 1'.I(I4 (M. de Mathau).
The fenuile resemliies that sex of the Peruvian subsj)ecies e.\cept in the
colour of the band. One of our males of the Bolivian subspecies has all the spots
of the underside of the hindwiug creain-coldur and cm the uji]ierside only the two
middle spots tipped with bright red.
( 4.58 )
21. Papilio dardamis Falir. (ITO^).
(^. PaplIio Kqufn Trnjanvs darthinuii Fabricius, Knf, Si/yt. iii. 1. p. H). n. 21) (17'.).'i) (Brazil).
$. Papilio Eqw.'t Ti-ojaitu.i Imx id., I.e. n. 30 (ITM) (Brazil).
(J. Papilio opleus Godart, Enc. Milh. ix. p. 33. n. 22 (ISl'.l) (Brazil ; — mutilated specimen).
cJ. Papilio ihmkmm^iA. J. c.f.lZ.-a. 1.34 (1810) (Brazil) ; V.MC%!i, Up. E.rot. p. 2(1. t. 13. fig. 2(18.3.5) ;
Boisd., Spec. Gin. Up. i. p. 304. n. 139 (183(5) (Brazil) ; Lucas, in Gu^r., Du:l. Pitt. IJiM. Nat.
vii. p. 47 (1838) ; Doubl., List Up. /ii."). Brit. .l^w. i. p. 13 (184,5) (Brazil).
?. Papilio Iros, Godart, I.e. 0. 1.3.5 (1811)) (Brazil) ; Donov., .\,it. Hepox. ii. t. 29 (18:23) (Brazil);
Boisd., I.e. p. 304. n. 139 (1836) ; Doubl., I.e. p. 13 (1844) ; Prillw., Sitlt. Eni. Zeit. xxvi. p. 129
(1865) (Corooyado).
(J ? . Papilio danlaiiu.i, Doubleday, Westw. & Hew., Gen. Diarn. Uji. i. p. 18. n. 194 (184t)) ; Gray,
Cat. Up. U.I. Brit. .1/w.v. i. Pap. p. 44. n. 224 (1852) ; id., Ust Up. /».v. Brit .I/?/,5. i. Pap. p. 60.
n. 237 (1856) (Brazil) ; Ment'tr., Eniim. Corp. Anim. .Uiis. Pelrn/i., Up. i. p. 5. n. 75 (1857) ;
Felder, Verh. Zool. Bol. Ge«. Il';™xiv. p. 294. n. 78 (1804) (S. Brazil): Butler, Cat. Dinrn. I.rp.
descr. Fabrie. p. 236. n. 10 (1869) ; Kirby, Cat. Dinrn. Up. p. 531. n. 77 (1871) (Brjizil) ; Burm.,
Deser. Pep. Argent, v. Lep., Atla.s p. 8. n. 18 (1879) (Corcovado) ; Oberth., El. dEnt. iv. p. 91.
n. 283 (1880) (Brazil) ; Haase, Uutersuch. Mimicry i. p. 79 (1893).
cJ ¥ . Palpiis black ; posterior abdominal segment not spotted red in male,
there being in female a spot only behind vaginal cavity ; vein R' of hiudwing more
distal than M- ; tail long, spatulate, tooth M' also somewhat prolonged. Red
spots of hindwing not opalescent. Scales behind SIP of underside of forewing
elongate, entire in both sexes ; scales of bnffish-green patch of forewing and most
scales of the red patch on upjierside of hindwing of male also entire. Mid- and
hindtibiae of male incrassate, densely covered with minute hairs, the s])ines being
small and almost restricted to the underside ; foretibia spinose. Fore- and midtarsi
in male about half as long again as tibiae ; not quite so long in female. In
male sometimes a greyish band on underside of forewing across apex of cell,
corresponding to the band of female.
Scent-organ : fold large, containing a very narrow strij)e of long grey wool.
Genitalia : c?. Tenth tergite a little longer than the sternite, bearing proximaily
on each side some irregular prominent teeth. Harpe very broad, rounded and
strongly dent.ite at apex, one large conical tooth veutrally in middle. Penis-sheath
acuminate, the tip a little curved over the oriKce. ?. Some distance proximaily
of vaginal oritice on both sides a large, triangular, somewhat rounded lobe, convex
on outer side ; postvaginal area transversely wrinkled, mesially impressed ; proximal
edge of this impression somewhat raised, especially mesially. Bristles of anal
segment gradually tapering to a fine point.
Early stages not known.
Jlal/. Brazil : Province of Rio de Janeiro.
In the Tring Museum: 21 <?(? and 16 ? ?.
22. Papilio orellana Hew. (1852).
Papilio o)f//'i)»r Hewitson, Trans:. Enl. Soc. Load. (2). ii. p. 2.3. t. 5. fig. 2. cj (1852) (.Amazon);
Uoubl., Westw. and Hew., Cen.Diurn. Up. ii. p. 529 (18,52) ; Gray, Cut. Up. Ins. Hril. .\liis. i.
Pap. p. 51. D. 246 (1852) ; Wall., Trans. Ent. Soc. Und. (2). ii. p. 250 (1H54) (Upper Amazons ;
forest) ; Gray, Ust Up. Ins. Brit. .Mus.i. Pap. p. 05. n. 201 (185ii) (Ega) ; Bates, Tran.t. Enl.
Soc. Land. (2). V. p. 34,3, .358 (1801) ((J, Ega) ; id., Joitrn. Entom. I. p. 226 n. 18 (1802) (Ega) ;
Felder, Verli. Zool. Bol. Gcv. W'lrn xiv. p. 295. n. 79 (1804) (Ega); Kirby, Cat. Dinni. Up.
p. 528. n. 04 (1871) (Ega) ; Oberth., Et. d'Eut. iv. p. 116. n. 270'"- (1880) (Iquito.-) ; .StauJ.,
E.rot. Tagf. p. v.). t. 1.3. ? (1884) (Iquitos) ; Halmel, Iris iii. p. 275 (18;i(l) (Sao I'anlo de
OliveDfa).
Palpus black ; M- of hindwing more distal than R' ; tooth R^ prominent.
6- Forewing witiiout markings, excejit the wiiite fringe-dots, whicli are
( 459 )
sometimes vestigial ; Llue-blacl!:, scales feebly deutate, those of upper layer narrow
in apical area ; a large purplish red discal area ou hindwiug, entering cell,
extending from SC^ be3'oud M-, non-opalesceut, the red spots of the underside
shining through ; scales of this patch entire. On underside scales behind SM- of
forewing and some before SM^ elongate, entire. Tibiae simple, spinose. Abdomen
with red spot only at base.
? . Forewing similar to that of male, less blue, scales dentate. A red band
ou hindwiug consisting of six spots, first and second smallest and usually isolated,
third, fourtli and fifth much longer than broad, scales dentate ; the spots smaller
below, paler, and all separated.
Scent-organ : white wool from base to apex of fold, the wool short, even, as if
shorn. In one of our specimens there is a large cinnamon streak on fold (SM'_).
Genitalia: cj. Tenth tergite with small tubercle on each side dorsally at base,
the lateral edge being somewhat dilated proximally of the tubercle. Clasper
rounded ; harpe of a similar type as in /-'. hfsunder and allies, long, reaching close
to apex of clasper, rounded-truncate, with about six large apical [teeth, no tooth
at ventral margin proximally of middle. Fenis-sheath acuminate.
llab. Ujiper Amazons.
In the Tring Museum : 'i S 6 , 'i ? ? , from : S. Paulo de Olivent-a ; Iquitos
(Stuart).
In coll. Oberthilr from : Ljuitos, November ; Cavallo Cocho, May — July.
In coll. H. J. Adams a series of 4 <?<?, .5 ? ?, from the Upper Amazons.
Ti. Papilio sesostris (!ram. (1779).
cJ. Papil'w Equen Trojauvs sesostris Cramer, PaiJ.Exot, iii. p. 34. t. 211. fig. F. G (1779) (Surinam).
?. Papilio Eques Trojanus tullus, id., I.e. iii. p. 153. t. 277. fig. C. D (1780) (Surinam).
Palpus black in both sexes, M- of hindwiug on a level with W or more distal.
S. Forewing : a large green patch from M^ to hinder margin, contiguous
with cell, consisting of three partitions, the first and second or all three acuminate
distally, the scales composing the patch broad, rounded ; rarely a streak of
dispersed green scales in cell ; rest of wing velvety black, upper scales narrower,
lanceolate, nearly all entire ; under scales dentate, broad. Hindwiug with or
without red spot behind BI-, the spot not being opalescent.
On underside the upjjer scales in posterior area of forewing entire and
lanceolate, lieing bidentate on rest of wing ; occasionally small creamy patches
posteriorly on disc. Iliudwing always with three large rod contiguous spots
li^ — (SM'), more or less shaded with white, and a small spot ou abdominal fold,
there being usually also a red dot R' — R- near margin and often a small spot
11^ — 11' in front of the patch of three ; a submarginal dot SO^ — R' does not seem
to be ever present. Tibiae not incrassate, spinose.
?. Forewing opa(|ue, being very little jialer distally than basally ; there are
always at least two creamy or bufiish-white patches on forewing ; upper scales
of forewing partly lanceolate, acuminate. Hindwiug : red band broad and
euutiuuous either from R- or R' to abdominal margin, rarely represented by only
two isolated spots ; spots SC- — R- usually present, isolated, small as com2)ared
with the posterior spots ; red upper scales nearly all entire, except near abdominal
margin. Scales on underside dentate, a very few in the white patch of the
forewing lieing acuminate.
Scent-organ : white wool not extending to base, the basal third of the fold
30
( 460 )
being covered with long, half-erect, rounded-trnncate or feebly sinuate, purplish
black scales, there being also a brush of long hairs ; the greyish white scales at
the discal side of the patch of wool nearly all entire.
Genitalia: c?. Tenth tergite a little longer than the long acuminate lobes
of the sternite ; edge of ninth tergite minutely serrate. Harpe long, gradually
narrowed distally, longitudinally impressed ; apex rounded, either entire or
denticulate, the most ventral tooth being usually somewhat enlarged ; ventral
edge with a solitary small conical tooth proximally ; a compressed, tuberculiform
ridge on clasper close to ajiex of harpe. Penis-sheath oblicpiely truncate, the
proximal (or ventral) edge produced into an abrupt short-jiointed lobe. ? .
Vaginal orifice surrounded by a ridge which is widened behind into a broad lobe ;
this lobe rounded, concave on distal side, somewhat like an oyster-shell in outline,
bent forward, lying over the vaginal orifice ; the edge of this lobe a little iucrassate
centrally ; proximally of the vaginal ridge there is on each side a large, irregularly
triangular lobe which is finely hairy.
Early stages not known.
Ilab. Mexico to Bolivia and Goyaz.
a. P. sesostris zestos Gray (1852).
/"(jy^^V/o .scso»7;v.<, Doubleday, Westw. & Hew., Gen. Diurti. Lrp. i. p. 19. n. 213 (1S17) (/«(rt!Hi) ;
Reak., Pmi:. Ent. Soc, Philatl. ii. p. 13'.). n. 9 (1863) (synon. jmrliiii ; Honduras) ; Weidem.,
ibid. p. 148 (1863) (Central America ; parl'im) ; Dist., Pror. Enl. Sue. J.nnd. p. xiv. (1876)
(Costa Rica) ; Hopff., Slelt. Eut. Zeil. xl. p. 50. n. 9(1879) {partim ■ Honduras).
Papilio sesios Gray, Cat. Lej). Ins. Brit. Mus. i. Pup. p. 47. n. 23,5. t. x*. fig. 5. ? (18.52) (Honduras) ;
id., List Lep. Ins. Brit. Mus. i. Pap. p. 62. n. 248 (1856) (Honduras) ; Bates, Trans. Ent.
Soe. Loud. (2). v. p. 340 (1861) (Honduras) ; Felder, Vrrk Zool. Bot. Grs. Wicn xiv. p. 292.
n. 39 (1864) (Mexico; Honduras); Stand., Exnt. Tngf. p. 13 (1884) {partim; Central
America); Godm. & Salv., Binl. Centr. Amcr., Rhop. ii. 190. n. 1 (1890) (Guatemala; Brit.
Honduras ; Nicaragua ; Honduras).
Pajiilio sesoslris, local var. zestos, Bates, Trans. Ent. Sue. Lnnd. (2). v. p. 355 (1861) (2""'l''>^t
Honduras).
Papilio sesostris var, a. P. zestos, Kirby, Cat. Diurn. Lep. p. 525. sub n. 60 (1871) (Honduras).
S. Hindwing always with red spot on npperside ; on underside the band of
spots K'— (SM') almost at right angles to the veins, the last of these three spots
ellijjtiual, not being produced distad, spot on abdominal fold rather large.
? . Patch R' — M^ of forewing contiguous with cell, a small spot in cell.
Band of hindwing bright red on xippcrside, not shaded with buff ; on underside
spots R- — M- well separate from cell.
Ilab. youth Mexico to Gosta Rica.
IntheTriug Museum 5 SS, 4 ¥ ¥ , from : Coatzalcualcos, S. Mexico, July rjU4
(A. Hall) ; Volcan de Miravalles, Costa Rica (Underwood) ; Juan Vinas, 2500 ft.,
and Carillo, 30110 ft., Costa Rica, October 1904 (A. Hall).
b. F. sesostris tarqtdnius Boisd. (lS3G).
Papilio tarquinius Boisduval, Spec. Gin. Lip. i. p. 296. n. 127 (1836) (¥, Colombia) ; Gray, Col.
Lep. Int. Brit. Mus. i. Pap. p. 47. u. 234 (1852) (Colombia) ; id., List Lep. Ins. Brit. .Mus. i.
Pap. p. 62. n. 247 (1856) (Colombia) ; Bates, Trans. Ent. Soc. Land. (2). v. p. 358 (1861);
Felder, Verh. Zool. Bot. Ges. Wieu xiv. p. 292. n. 40 (1864) (Bogota) ; Oberth., Et. d'Eul. iv.
p. 88. n. 280 (1880) (Colombia ; Panama ; partim, ^ excepted) ; Godm. & Salv., Biol. Centr.
Amer., B/iop. ii. p. 191. n. 2 (1890) (Panama ; Bugaba, Chiriqui, Lion Hill).
Papilio sesostris local var. zestos. Bates, Trans. Ent. Snc. Loud. (2). V. p. 355 (1861) (jntrtim ;
N. Granada).
Papilio sesostris var. b. P tar uiuiiis, Kirby, Cat. Diurn. Lep. p. 525. sub n. 60 (1871) (N. Granada).
(461 )
Papilio sesostris, Pelder, VerJi. Znnl. But. Ges. Wiett xiv. p. 292. n. 38 (1864) (partlm ; N. Granada ;
Ecuador) ; Hopffer, Slett. Eiit. Zeit. xl. p. 50. n. 9 (1879) {pariim ; N. Granada) ; Godm. &
Salv., Trans. Enl. Soc. Loud. p. 126. n. 2.TO (1880) (Sta. Marta) ; Hahnel, 7,-;.-.- iii. p. 147 (1890)
(San Esteban) ; id., I.ii. p. 194 (1890) (M.'rida) ; id., I.e. p. 203 (1890) (Valera) ; Haeusch, Bcrl.
Enl. Zeihchr. xlviii. p. 1.54 (1903) (Archidona, 640 m,).
Fapilio upso-^lrii var. [J sc.sfo.v, Obertbiir, Et. d'Ent. iv. p. 90, sub n. 281 (1880) ( partim ; N. Granada).
PapiUo 2«.s(o.«, Staudinger, E.mt. Ta/jf. p. 13 (1884) (partim ; Colombia).
iS. Most specimens with red spot on upperside of liiiidwing ; baud
R' — (SM') on underside more oblique than in the preceding, the posterior one
of these three spots being more distal and the anterior a little more proximal
than the central one : spot on abdominal fold often small or absent.
? . Patch M' — M- of forewing on the whole longer than in the preceding ;
band of hindwing a little nearer the cell, more oblique, the distal marginal area
therefore wider, which is especially noticeable on underside.
llab. Panama ; Colombia ; Ecuador ; North Venezuela.
In the Tring Museum 30 dS, 15 ? ? , from : Boquete, Chiriqui, 3500 ft.
(Watson); Brava I., Sevilla I., and Parida I., January 1902 (J. A. Batty) ; Cohjn,
Rio Dagua, West Coast of Colombia (Rosenberg) ; Onaca, Sta. Marta, 2200 ft.,
September — October 1901 (Chas. Engelke) ; Cananche, Cundinamarca, July 1903
(Mathan) ; Villaviceiicio to R. Ocoor, Colombia, January 1897 (Dr. Blirger) ;
Cachabi, West Ecuador, November 1896 and January 1807 (Rosenberg); Cachabi
to Paramba, February 1897 (Rosenberg).
c. P. .sesostris sesostris Cram. (1779).
Seba, ThcMiiir. iv. p. 32. t. 26. fig. 19. 20. ^ (1764).
(J. Pupilio Eqiies Trojiiims xesnstris Cramer, I.e. (Surinam) ; Jabl. & Herbst, Naliirs. Schmetl. ii.
p. 70. n. 21. t. 10. fig. 1. S (1784) ; Esper, Ami. Srhmll. p. 51. n. 20. t. 12. fig. 2 (1788).
5. Ptijiilio Eque.^ Triijanus tnllus Cramer, I.e. (Surinam) ; Esper, Aiis!. Sehmctt. p. 52. n. 21. t. 12.
fig. 3 (1788) (artefact ; forewing of tidlus Cram, and hindwing of lysaiider Cram.).
Pajiiliu Eques Trojunns aenens e, Paj}ilio scsoslrix, Gmelin, Si/st. Nal. i. 5. p. 2233. sub n. 16 (1790).
Priamides tullus, Hiibner, Verz. hek. Schmett. p. 87. n. 901 (1818 ?)
Parldes sessostris (!), id.. I.e. n. 912 (1818 ?).
Papilio tulluK, Godart, Enc. Meth. ix. p. 37. n. 37 (1819) (Guyane) ; Boisd, Spec. Gen. Up. i. p. 295.
n. 126 (1836) (Cayenne ; Surinam) ; Doubl., Lid Lcp. Ins. Brit. Mns. i. p. 12 (1845) (Brazil) ;
id., Westw. & Hew., Gen. Diurn. Lep. i. p. 18. n. 201 (1846) (partim) ; Hewits., Trans. Ent.
Soc. Land. (2). i. p. 97 (1851) ( ? of sesodrix) ; Doubl., Westw. & Hew., I.e. ii. p. 529 (1852) ( ?
of sesostris).
Papilio sesostris, Godart, I.e. ix. p. 38. n. 40 (1819) (Guyane) ; Lucas, Lej). E.r.ol. p. 28. t. 14. fig. 1.
cJ (1835) (Gujane) ; Boisd., Spec. Gen. Lep. i. p. 299. n. 131 (1836) (Surinam ; Cayenne) ;
Doubl., List Lep. Ins. Brit. Mus. i. p. 13 (1845) (Brazil) ; Erichs., in Schomb., F. F. Brit.
Guiana p. 593 (1848) ( ? = tnllns Cram.) ; Doubl., Westw. & Hew., Gen. Diurn. Lep. i. p. 19.
n. 213 (1847) ; Gray, Cat. Lep. Ins. Brit. Mas. i. Pap. p. 58. n. 267 (1852) (partim. ; Brazil) ;
Wall., Trans. Ent. Soe. Land. (2). ii. p. 255 (1854) (Amazons ; forest) ; Gray, List Lep. Ins.
Brit. Mus. i. Pap. p. 69. n. 282 (1856) (jjartim) ; Menetr., Enum. Corp. Auim. Mns. Pelrap., Lep.
i. p. 5. n. 84 (1857) (Brazil) ; Bates, Trans. Ent. Sae. Loud. (2). v. p. 339, .355 (1861) (Paril to
Tabatinga) ; id., Jou™. Entum. I. p. 225. n. 11 (1862) (throughout the Amazons, in the forest) ;
Felder, ]'er/i. Zool. Bnt Ges. Wien xiv. p. 292. n. 38. (1864) (partim ; Surinam ; Cayenne ;
Guiana ; Amazonia) ; Bates, Natural. liiv. Amaz. p. 25. (18IJ4) (P,arii, (J in swampy shades, ?
in more opeu places); Kirby, Cat. Diurn. Lep. p. 525. n. 60(1871); Druce, Prac. Zool. Soe. Land.
p. 244. n. 1 (1876) (Peru) ; Moschl., Verh. Zool. Bot. Ges. Wien xxvii. p. 295 (1876) (Surinam) ;
Hopff., Slett. Ent. Zrit. .\1. p. 50. n. 9 (1879) (partim \ Surinam ; Peru ; Brazil) : Oberth.,
Et. d'Ent. iv. p. 90. n. 281 (1880) (Guyane ; Par^ ; Amazons) ; Staud., E.rot. Taijf. p. 13. t. 8.
$ ? (1884) (Amazons) ; Sharpe, Pme. Zool. Soc. Land. p. 555. n. 2 (1890) (Prov. of Goyaz) ;
Hahnel, /™ iii. p. 240 (1890) ( ViUabella, Amaz.) ; id., /.'■. p. 253 (1890) (Mbup.=) ; Haase,
( 4fi2 )
riilersuch. itimkrij i. p. 79 (1893) ; Michael, Ins vii. p. 213 (1S94) (Sao Paulo de Olivenc^a) ;
Weeks, lUuHti: niuni. Lq,. p. 20 (190')) (Clmlumani).
Priiicrps ihim/naiis scsy.<fr/s, Hiibner, Sminnl. Exnt. Schmctl. i. t. 12S. (J. (180G — .') (Ii:ib. V ; " ? "
false).
rojiilin ctitnra Gray, C'<(/. Lrp. Iiis. Brit. JI/«.--. i. Paj>. p. ;>H. n. 2("i('). t. 5. fig. 2. $ (1852) {jiitrliiii ;
Hon (J).
Papilio lijcmnes id., I.e. p. 52. n. 251 (1852) (nom. nov. pro Esper. t. 12. fig. 3 ; — artefact) ; id., Lht
Lep. Ins. Brit. Mus. i. Pap. p. GG. n. 2r)6 (1851'.).
Papilio scHostris var. (J zestos, Oberthiir, Et. d'Eiit. iv. p. 00. sub n. 281 (1880) {/utrlim ; Amazons).
Entlnpogim se^iostris, Kirby, in Allen's Nat. Libr., Lep. Butt. ii. p. 270 (IH'.IG) ; id., in Hiibn., Sdiiiml.
EjcoI. Schmett. ed. ii. p. 88. t. 128. fig. 3. 4. (190—?).
c?. Usnall}' no red spot on upperside of hiiulwing ; spots of underside rather
more distal than in the previous, in Bolivian specimens sometimes minute.
?. Two patches M' — SM- on forewing, remote from cell. One of our two
females- from East Bolivia (Santa Crnz de la Sierra) has only two small spots
on upperside of hindwing, bearing on underside an additional dot R' — 11^ ; the
spots of upper- and underside of hindwing are ])ink in both specimens.
Hal). Orinoco ; the Guianas ; Panl to Peru ; Bolivia ; Goyaz.
In the Tring Mnseam TO S<S, 20 ? ?, from: Snapnre, Caura It., Orinoco,
Oct. I'J02 (S. M. Klages) ; Britisli Gniana; Surinam ; Sautareiu ; Obidos ; R. Jnrna,
June 1897 (Dr. Bach); Iquitos (Stuart); R. Ucayali and R. Oachyaco (Stuart);
R. Mi.xiollOjLoreto (Baer) ; Pozuzo,Hu;innco (Hoffmanns); ('hanchamayo(Schnnkc);
R. ("huchuras, Hnannco (Iloifmanns) ; La Union, Carabaya, ~tJOU ft., Nov. I'.dl-t
(Ockenden) ; Salinas, Bolivia, July 1895, and Reyes, August 1895 (Stuart);
Mapiri ; Province Sara, S. Oruz de la Sierra (Steinbach).
24. Papilio childrenae Gray (1832).
(J. Papilio childrenae Gray, in Oriff., .1«(»(. KimjJ. xv. p. (573. t. 38. fig. 1 (1832) (" Brazil,'' error
loci).
? . Papilio childrenae, Felder, Wien. Eut. Mun. v. p. 73. n. (j (1801) (Bogota).
(? ? . Close to P. nesostr/s. In male, green area of forewing much larger,
entering cell, occupying from one-third to two-thirds of the cell, one or more
white spots beyond apex of cell, either on both sides, or only below, or the
spots absent; red spot of hindwing always present, obliijue, contiguous with
cell, cxjianded at cell from before M' to M^, the red scaling before M' being
restricted to the very base of the cellule. On underside of liiiidwiiig there is a
row of spots from SO" to anal angle, but the upper spots, which are small, are
always shaded with black, being in most specimens absent ; spot R' — M- larger
than the others, spot M' — M- often the only one preserved, besides a more or less
linear spot M- — SM-, which is often interrnjited. In female, two huffish white
patches M'— SM" on forewing, separate from cell, the first sometimes minute, the
second always large, often a minute spot behind SM" and another before M' ;
several spots beyond ajiex of cell, but no spot in cell ; hindwing resembling that
of P. sesosfiis.
Scent-organ as in P. sesostris.
Genitalia as in P. sesostris; bari)e more curved, dentition at apex a little more
ventral, the solitary ventral tooth larger.
Early stages not known.
llab. Guatemala to Ecuador.
Two subspecies.
( 463 )
a. P. chilclrciKK' ohiUlrcnac Gray (1832).
Papdin ihndrnme Gray, I.e. (1832) ; Rtaud., Ej-oi. Tnrjf. p. 13 (1HS4) (Central America) ; Gotlm. &
Salv., Biul. Ceiitr. Amer., Ithip. ii. p. 192. n. 3. t. G.5. fig. 1. (J, 2. ?, 3. genit. (18'.l0)
(Guatemala ; Nicaragua; Costa Eica ; Panama) ; Haase, Uidersuch. Mimierij i. p. 79 (1K9:!).
ri(j>ilio se.tostrii var., Gray, Cut. Lep. Ins. Brit. Mus. i. Pap. p. 58. sub n. 2G7 (1852) (Brazil) ; id.,
Lixt Lep. Tiis. Vril. Miik. i. Pap. p. 70. sub n. 282 (1852) (Brazil).
S . Forewing always with a rather large white spot, which is nsnaliy preceileil
by a smaller oue.
?. Band of hiudwing on the whole brighter red than in P. ch. childrenae.
Hab. Panama to Guatemala.
lu the Tring Musenm 74 (^(S*, 8 ? ?, from : Carillo, Costa Rica, June— July
19U3 (Underwood); Carillo, 3000 ft., Oct. 1904 (A. Hall); Chiriqui; Colon.
h. P. childrenae oedippus Lucas (1857).
(J. Pfipilin nedippus (!) Lucas, in Casteln., Vnij. Amer. Siul, Enl. p. 197 (1857) ("Inter, of Brazil ");
Staud., E.rnt. Tmjf. p. 13 (188-i) (Colombia) ; Godm. & Salv., Biol. Ceidr. Amer., Rhop. ii.
p. 192. sub D. 3 (1890) ; Skinn., Eid. News xv. t. 1. fig. 3. t? (1901).
Pajiilio ocdypiiH (!) Lucas, I.e. Lip. t. 2. fig. 4. $ (1857).
Papilio semstris, local var. childrenae. Bates, Trans. Ent. Soe. Loud. (2). v. p. 355 (1861) (New
Granada).
Papilii) cliildifnae, Felder, Wien. Ent. Mmi. v. p. 73. n. 0 (IS(jl) ; id., Verh. Zool. Bnt. Ges. Wien xiv.
p. 292. n. 41 (1804) (Bogota ; R. Napo ; Interior of Brazil) ; \A.,RriseNorara,Lep. p. 21. n. 11
(18G5) (Bogota) ; Kirby, Cat. Diiirn. Lep. p. 525. n. fiOa (1871) (N. Granada; Ecuador) ;
Oberth., Et. d'Enl. iv. p. 90. n. 282 (1880) (N. Granada).
Pajiilio oedipus, Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 292. sub n. 41 (1864).
(?. No white spot on upperside of forewing, or (rarely) only one small dot*
Underside of hindwiug mostly without red spots SC- — R-.
?. Band of hindwing pale, soraewhat buffish proximally on upperside.
Ilah. Colombia and Ecuador.
An Ecnador specimen (c?) in coll. Hewitson (British Museum) has a small red
spot on upperside of hindwiug and two small sjiots R' — M- on underside ; the green
scaling in cell of forewing is mnch reduced in extent.
lu the Tring Museum 40 Si, 7 ??, from: Rio Dagua, W. Colombia
(Rosenberg) ; " Bogota."
25. Papilio erlaces Gray (1852).
cJ. Papilio rrlaees Gray, Cat. Lep. Ins. Beit. .Uns. i. Pap. p. 49. n. 240. t. 8. fig. 9. J (1852) (^ only
? .another species ; Bolivia).
? . Papilio rijplwiex id., I.e. p. 49. n. 241 (1852) (" S. America, coll. Hewitson ").
Palpus always black in both sexes. Fringe of both wings dotted with white.
c?. Tibiae spinose, not incrassate, resembling those of female. Eighth
abdominal segment edged with red veutrally in front of clasper. Forewing : a
large olivaceous green patch from inner margin forward, sometimes entering cell,
the scales composing it elongate, tongue-shaped ; a creamy white spot M' — M-,
usually large, very often preceded by a smaller spot R^ — M', occasionally followed
liy a small dot standing behind M-, these spots wanting in nearly all Ecuadorian
specimens and in a small projwrtion of the individuals from Peru and Bolivia.
Hindwing: three red spots R- — M', abont half-way between cell and distal margin,
strongly opalescent, the first spot sometimes minute, occasionally absent, rarely
a fourth dot R' — R= marked ; on underside the spots much paler, in the northern
forms more or less creamy, the row being usually prolonged forward by one or
two spots, there being also two spots present on the abdominal fold.
(464)
?. Ventral edge of seventh al)(loiuiiial segment and apex of eighth red.-
Forewing with large white patch, consisting of two or three discal spots and a
large cell-spot, there being often some small dots distally of ajjex of cell.
Hiudwing : a band of red spots, distant from cell, rarel}- tonching cell at W.
Nenration : cell of hindwing acuminate, D' being transverse and shorter than
D"*, usnally mnch shorter.
Scent-organ with white wool.
Genitalia : c?. Tenth tergite abont one-third longer than the sternite, lateral
edge at base sharp, somewhat projecting, irregularl)' sinnons, minntelj- denticulate
like the edge of the ninth tergite. Harpe of almost even width from base to ajiex,
somewhat curved dorsad, its dorsal edge being concave, apex rounded dorsally ;
a large tooth at ventral margin in or beyond middle ; from this tooth to apex
a number of smaller conical teeth. — — ?. In front of vaginal orifice a long lobe,
of nearly even width, feebly sinnate at apex ; on each side but more jiroximally
another lobe, rounded at apex, the internal edges of these lateral lobes extending
on to the proximal surface of the median lobe ; behind the vaginal orifice a large
rounded lobe covering the vaginal cavity. Anal segment with numerous short,
stont, club-shaped or pointed bristles.
Early stages not known.
Ila/j. Ecuador to North Argentina.
No representative known from the Amazons, Brazil, the Guiauas, Venezuela,
and Colombia.
a. P. erlaces laci/des Hew. (18C9).
J'. Papilio lacydes Hewitson, Equal. Lej). i. p. 1. n. 1. (18C9) (Ecuador) ; Kirby, Cat. Diurn. Lep.
\\ 527. n. 62c (1871) ; Maas. & Weym., in Stiibel, R(^iseu S. Amei:, Lep. p. G4. n. 80 (1890)
(Huamboya).
5 . Papilio erithuliaii $ ab. ? eqiieslris Oberthiir, Et. il'Eiit. iv. p. 88. sub n. 270. t. 5. fig. 2 (1880)
("Quito"),
fj. Papilio erithalion var. laci/iles Oberthiir, I.e. p. IIG. n. 276 (1880) (Ecuador).
(J?. Papilio laeyiks, Kirby, Trans. Eut. Soc. Land. p. 351 (1881) (dcscr. of J ; Saraya9U ;
Chiquinda); Grose-Smith & Kirby, RIiojj. ExuI. ii. p. 39. n. 2G. Pap. t. 10. fig. 1, 2. ^ (1897)
(Sta. Ines ; Saraya^u ; Chiquiuda).
Papilio erithalion var. equeslris, Maassen & Weym., in Stiibel, Piisen S. Amer.. Lep. p. 04. sub n. 88
(1890) (Huamboya).
1^. Forewing rarely with a white spot. Spots of hiudwing below small,
creamy white, partly pinkish.
? . Forewing : white cell-patch large, reaching across cell, there being always
a triangular streak in front of cell, most specimens with some small spots distally
of apex of cell, the cross-veins alone being black ; white spot U- — W large, often
larger than spot R^ — M', which is mostly much reduced iiosteriorly, extending to
M' only distally ; no sjwt behind M'. 13aud of hindwing white.
Hai. Eastern Ecuador.
In the Tring Museum, Go c?c?, 26 ? ?, from : Loja ; Zamora, 3000—4000 ft.
(0. T. Baron) ; Santa Inez (R. Haensch).
In coll. Oberthiir a series from : Ambato ; Saraya^u ; Banos to Canelos.
b. F. erlaces xanthias subsp. uov. (PI. V. fig. 24).
(J. Papilio erlaces, Druce, Proc. Zool. Soc. Lotid. p. 245. n. 3 (1876) (I'ozuzo).
c?. Not constantly different from : the following ; forewing nearly always with
one or two white spots, there being sometimes a minute third spot behind M".
( ^65 )
Spots on nuderskle of hindwiag either cream-colonr as in lacydes, or red as in
erlaces, or intermediate.
? . Forewing : two white spots R- — M', the second mnch the largest, its
posterior proximal corner cnt off ; cell-spot more or less triangnlar, jnst reaching
across cell, rarel}- a small streak in front of cell. Hiudwing : band j-ellowish
cream-colonr, broader than in lacijdes, sjiot C — SC^ small or absent.
Hub. North-East Pern, sonthward to Hnanuco. Type of name : ? from Poznzo.
In the Tring Mnseum, '~;i c? c?, 5 ? ?, from: Poznzo, 800— lUOO m. (W.
Hoffmanns) ; Cnshi, 1820 m. (W. Hoffmanns).
In coll. Oberthttr a series of males from Uhachapo3-as and Moyobamba.
c. P. erlaces erlaces Gray (1852).
(J. PapHio erlaces Gray, I.e. (1852) (Bolivia); \i.. List Lep. Lis. Brit. Miis. i. Pup. p. 63. d. 253
(18.'>6) (pmiim) ; Bates, Trans. Enl. Soo. Land. (2). v. p. 341, 356. n. 8 (1861) (Bolivia ;
Eastern Peru) ; Felder, Verh. Zool. Bid. Ges. Wien xiv. p. 293. n. 46 (1864) (Bolivia ; Peru) ;
Kirby, Cat. Diurii. Lep. p. 628. n. 62g (1871) (Bolivia) ; Hopff., StM. Eiit. Zeit. xl. p. 49. n. 5
(1879) (= luctiiosa ; Bolivia ; Chanchamayo ; descr. of $ ) ; Weeks, lUiistr. Diiirn. Lep. p. 28
(1905) (Bolivia : Cochabamba).
?. PapiUii hlerodes, Bates, I.e. p. 341 (1861) (parthn).
?. Pa2>!lio cjjphotes Gray, I.e. p. 49. n. 241 (1852) ("S. America") ; Kirby, l.r. p. 527. n. 60d (1871)
(pnrtim).
(J. PapUio lucluosa Butler, Cist. Ent. i. p. 12 (1869) (Peru) ; Kirby, Cat. Diurn. Lep. p. 527. n. 62 b
(1871).
S. AVhite spots of forewing very variable in size, rarely absent. On nnder-
side of hiudwing from five to seven red spots, the central ones the largest and more
or less extended pinkish white.
? . Forewiug : most specimens with three white spots behind cell, there being
a streak behind M' in addition to spots R^ — M' ; spot R- — R' sometimes minute ;
cell-spot very varialile, in most individuals triangnlar, not reaching across cell, in
some specimens as large as in lacydes, there being in this case a prominent streak
in front of cell ; some individuals with minute spot before R^. Hindwing : band
red, on the whole paler and spots R^ — M^ longer in the specimens from East
Central Peru than in those from South-East Peru and Bolivia ; the band consisting
usually of seven spots, the last of which are more or less merged together, the
upper two being occasionally absent.
Hub. Pern to North Argentina, from the Rio Peren(5 (Chanchamayo) southward
to Tucuman.
One of the (!hanchamayo females which we have seen has the band of the
hiudwing orange-red (coll. Charles Oberthiir), being a transition to xantkias.
In the Tring Museum, 70 SS, 23 ? ?,from : Chanchamayo (W. Hoffmanns) ;
R. Toro, Chanchamayo, August— September 1001 (Simons) ; R. Perene, March 1900
(Simons) ; R. Slucuri, S.E. Peru, June 1901 (Simons) ; various places in Carabaya,
S.E. Peru, apparently all through the year (G. Ockenden) ; Vilcanota, Cnzco,
3000 m. (Garlepp ; from Stand. & Bang-Haas, altitude correct ? ?) ; Cajon, Cnzco
(Garlepp) ; Huancabamba, N.E. of Cerro de Pasco (Boettger) ; R. Cachyaco, affluent
of R. Hnallaga (Stuiirt) ; Marcapata ; Mapiri ; Yungas de la Paz ; R. Burmcjo to
R. Pilcomayo, Bolivia, December 1903, and Province Sara, S. Cruz de la Sierra
(J. Steinbach); R. Grande, Province Cordillera, December 1903 (J. Steinbach);
Tucuman (G. A. Baer).
( 406 )
26. Papilio burchellanus Wostw. (1872). (PI. IV. fig. 1).
?. Piipilin hiiirliilhiiiiis Westwood, Tmnx. Kill. So,: Lnml. p. 101. t. :!. fig. .') (1.S72) (Tcncnte,
Farinhapodre, Brazil) : Kirby, Cut. Diiirii. Lep. p. 812. n. 3.51 (1877).
(J. Piipilio soaiina Schaus, Proc. U.S. Xut. iliis. \x\v. p. 424 (1002) (" Bolivia ").
(? ? . Palpns black. Mid- and hindtibiae of male slightly iucrassate and liairy,
foretiiiia spinose. Forewing without markings, except the more or less distinct
white fringe-spots. Rindwing with a row of widely separate rounded spots
nearer margin than cell, almost parallel to margin, tiie spots rather smaller and
paler beneath.
Scent-organ : a streak of huffish white wool in fold, the streak narrower than
in P. vertumnus.
Genitalia : S . Harpe long, straight, reaching to apical margin of clasper, and
bearing at the apex about half a dozen teeth ; no tooth in middle of ventral margin.
Ilab. Interior of Brazil. The occurrence in Bolivia requires confirmation ;
Mr. Schaus did not receive the specimen described by him direct from Bolivia,
but from a correspondent who has been a resident of Ilio de Janeiro. The species
is very interesting, being an exact counterpart of P. paiitlionit.i iiiiimi, which also
occurs in Brazil.
In coll. Charles Oberthur one male from Goyaz.
27. Papilio drucei Butl. (1874).
? . Papilio ciitora, Bates (iimi Gray, 1852, err. det.), Trans. Eiil. Snc. Lnml. (2). v. p. .111, .355 (ISill)
( ? only ; Ega).
cJ. Piqiilio (h-iicci Butler, TmiiK. Eiit. Soc. Lnml. p. 434. t. G. fig. 2. ^ (1874) (Ecuador) ; Kirby, Cat.
Diiini. Lep. p. 812. n. 357 (1877); Obertb., Et. tl'Ent. iv. p. 111!, n. 276"" (1880) (Pebas.
$ = fi/iuliiius) ; Kirby, Trails. Eiit. Soc. Lowl. p. 353 (1881) (Canelo.', jj) ; Michael, Iris vii.
p. 214 (1894) (Sao Paulo de 01iven9a).
?. Papilio opaHiiiis Butler, Trans. Ent. Soc. Loml. p. 145. n. 225. t. 3. fig. 5 (1877) (Rio Purus).
S. Palpus black, rarely red. Eighth abdominal segment edged with red
beneath. Tibiae and first tarsal segment hairy above, very slightly iucrassate.
Forewing : a sage-green patch from inner margin to M' or R', separated from
cell, often reduced to a narrow band; fringe dotted with white; sometimes two
small creamy white spots M' — (SM'), on underside occasionally four spots.
liindwing : three opalescent red spots R- — M- close to cell, often preceded by a
dot or streak R' — R- and followed by a narrow streak M- — (SM').
¥ . Palpns black or red. Apex of eightli abdominal segment and of seventh
sternite red. Forewing without white patch ; fringe dotted with white. Baud of
hindwing red, sometimes rather strongly opalescent, the red scales being in this
case nearly all entire, while the red non-opalescent scales are obtusely bidentate ;
the number of spots varying from five to seven, their size being also variable, the
band not touching cell, but standing sometimes close to it ; last two spots usually
merged together.
Ncnration, scent-organ and genitalia essentially as in P. laci/r/cs ; in female the
postvaginal plate more triangular, the iucrassate edge being mesially dilated into a
rather prominent tubercle which is a little curved backwards.
Early stages not known.
Ifaf). Ecuador to Bolivia, Upper Amazons.
In the Tring Museum 12 <Jc?, 15 ? ?, from : Archidona (R. Haensch) ; Coca,
Upiier Bio Napo, May— July 1899 (W. Goodfellow) ; Zamora (0. T. Baron);
Jnluity, Amazons, April I'm.") (Mathan) ; R. Jurua; Ljuitos ; K. Cachyaco, adlnent
( 467 )
of R. Hnallaga (Stuart); Poznzo, Hndnnco (W. Hoffmanns); R. (Ilmchuras
(W. IIiiiFraaiins) ; Paohitoa ; Cuzco ; Mapiri, Bolivia.
In coll. Obcrthiir a series of both sexes from : Tarajjoto ; ('avallo Coclio ;
Sarayaou ; Pebas.
28. Papilio cutorina Stand. (1808).
(J. Piipilio reiiiiminis local var. nilorii, Bates {iinii Gray, 18.')2, err. det.), Trans. Ent. Soc. Loml. (2).
V. p. Ml, .355 (ISr.l) (/mrlhn ; ^ only).
(J. riijiilin riilnra, Felder, Vcrh. Zonl. Bat. Ges. Wieit xiv. p. 202. n. 43 (18(;4) {parlim) ; Dnice,
Proc. Znol. Sue. Land. p. 245. n. 2 (1876) (tJcayali and Huallaga) ; Kirby, Trans. Eiit. Soc.
p. .353 (1881) (Ecuador, common !— this insect? ?) ; Hahnel, Iris iii. p. 275 (1800) (Sao Paulo
de Olivenfa) ; Michael, Iris vii. p. 214 (1804) (Sao Paulo de Olivenja).
Papilio vertumnus var. g. P. ciitura, Kirby, C'ul. Diurn. Lep. p. 625. sub n. Gl (1871) (Upper
Amazons).
S ? . Papitio cutorina Staudinger, Iris xi. p. 139 (1898) (Sao Paulo de Olivenga ; Pebas ; Iquitos) ;
id., l.c. p. 37l"i (1800) {mazeppa ii ? of cutorina) ; Grose-Smith, Rhnp. Exot. iii. p. 51. n. 37.
Pap. t. 22. fig. 1. 2. S (1902).
?. Papilio maspppa Grose-Smith, l.c. p. 42. n. 29. Pap. t. 17. fig. 3 (1809) (Liuitos).
Palpns red. Fringe of forewing spotted. Cell of hindwiug narrow at apex,
D' being transverse and short.
(J. Tibiae spiuose, not incrassate. Eighth abdominal segment edged with
red ventrally. Glaucons green patch of forewing from inner margin to M' or a
little beyond, proportionally longer than in P. eertumnti.i in basi-distal direction.
Only two red spots on njiperside of hiudwing situated between _M' and (SM').
Spots on underside of hindwing cream-colour, somewhat glaucous buff at edges,
resembling the spots of P. aeneas holivar.
? . Porewing without white patches. Hindwing with creamy baud consistiug
usually of five contiguous spots.
Neuration, scent-organ and genitalia essentially as in P. rcrtuirmus.
Early stages not known.
Ilab. Upper Amazons, East Peru, and East Ecuador.
In the Tririg Museum : 3 J <?, 3 ¥ ? , from : Coca, R. Napo, Ecuador (R. Haensch) ;
Rio Chuchuras, Hminuco (W. Hoffmanns); R. C'achyaco, affluent of R. Huallaga
(Stuart) ; Sao Paulo de Oliveuf a ; Itaituba ; Pelias.
In coll. Oberthiir : 2 cJc?, 7 ? ? from Icpiitos and Sao Paulo de 01iven(;a.
29. Papilio phosphorus Bates (1861) (PI. IV. lig. i). in. II).
(J. Papilio phosjihorii.'i Bates, Trans. Ent. Soc. Loml. (2). v. p. 342 note (1800) (R. Demerara).
? . Papilio ijraliauHs HewitsoD, Exot. Bull. ii. Pap. t. 5. fig. 13 (1801) (New Granada ; "^J" false).
Having received a female agreeing almost in every detail with Hewitson's type
(which is a female with a male abdomen stuck on), we have no longer any doubt
\\\a,t jilmsiphonis and gratianu.'i are the sexes of the same species.
(?. Tibiae spinose, not incrassate. Palpus red. Vppersidc : forewing semi-
transparent in apical area; a glaucons or huffish green patch from hinder margin
forward to M' or beyond ; one or two white spots behind M^ often a tliird before
M% minute or large, usually elongate, often absent. Hiudwing rather strongly
dentate ; a row of three, four, or five spots, gradually decreasing in size from M-
liirward, the last two more or less contiguous, the others separate, distance of spot
K' — M' from cell at least equal to half the length of the spot.
Untkrsii/e : forewing always (?) with iit least one minute white dasli behind or
before M-, usually with two spots, often three, two of which stand between M" and
(468)
SM^ Hindwing with six or seven spots, the row in middle abont lialfwaj-
between cell and distal margin, spot behind M- beyond middle of this vein.
? . This is the only instance in which the female bears on the forewiug a patch
somewhat similar to that of the male. Uppersidc : forewiug with slaty grey
patch from inner margin to M- or beyond, the patch narrower than in male, consisting
of huffish white, narrow, entire scales lying on black ones ; three or four white
spots within the patch, two standing between M^ and SM-. Hindwing : a pale
red band from abdominal margin forward, gradually decreasing in width from M-j
curved, situated about halfway between cell and distal margin, the posterior spots
contiguous, the anterior ones separate.
Underside : white spots of forewing as above, but no slaty scaling. Hind-
wing : band narrower than above, the spots all or nearly all separate.
Genitalia : c?. Harpe shorter than in the allied forms, the apical half more
acuminate-triangular, ventral tooth large.
Hail. British Guiana ; Colombia ; Eastern Peru ; Lower Amazons.
May be expected to occur also on the Middle and Upper Amazons.
a. P. ji/io,y)/i(VU!i p/w.yj/wriis Bates (1861) (PI. IV. tig. 9. S. type, 10. ?).
(J. Pa/iilio phosphorus Hales, I.I. (Rio Tlemeran) ; Felder, Verh. Zonl. Bol. Ges, H'/jh xiv. p. 29."!.
n. 47 (1864) ; Kirby, Cat. Diurn. Lep. p. 528. n. 62e (1871).
(?. Upperside : forewing, a patch from inner margin to M', being in that
direction much longer than it is broad, distant from cell, with two or three minute
white linear spots, or one, or no spot. Hindwing with an evenly curved row of
four or five spots.
Underside : on forewing at least one minute white spot.
?. Upperside : forewiug with a row of four white spots ; spot M' — M- much
larger than in Hewitson's figure. Band of hindwing consisting of six spots, the
first minute, the last four much smaller than in Hewitson's specimen.
Ilnh. British Guiana ; Lower Amazons.
Li the Tring Museum : 0 Jc?, 1 ?, from : British Guiana (H. Haensch) ; British
Gniana ; Igarape, Para (\V. Hoifraanns).
*. P. phosphorus gratianus Hew. (1861) (PI. IV. fig. 11).
? . Pupilin rjratiaiiiis Hewitson, I.e. (New Granada) : Felder, I.e. p. 295. n. 82 (1864) ; Kirbj-, l.i\
p. 528. n. 65(1871).
c?. Upperside : forewing, green patch much broader than in the preceding,
touching cell or nearly ; with or without white spots. Hindwing : three red spots
11- — M- and sometimes a minute dot R' — R-, the row less curved than in the
2>revious subspecies, the last spot not reaching further pruximad than the last
but one.
Underside : at least one white dot on forewing. On hindwing five or six red
spots, the spots R' — M- smaller than in P. ph. phosphorus.
$. Forewing with two white spots; last three spots of hindwing very large.
Ilab. Colombia ( ? ) ; Peru (cf <J).
"VVe have not seen Peruvian females, nor Colombian males, and therefore do
not know if there are two races in these countries or one.
In the Tring Museum 'i 6 S from Pachitea and Hillapani, Peru (received from
Messrs. Staudinger and Bang-Haas).
In coll. Oberthiir from Tarapoto, Peru.
(469 )
30. Papilio vertumnus Cram. (1T7',»).
(J. Piipaio Eques Troja/ius vertumnus Cramer, Paj). Ej^ot. iii. p. 32. t. 211. fig. A. B (177'.l)
(Surinam ; fig. C = P. anchises (J).
? . Pajnlio clxins Gray, Cat. Lep. Ins. Brit. Miis. i. Pap. p. 48. n. 2.37. t. 8. fig. 0 (1852) (Ega).
As the tibiae are in a number of species of the present gronp simply spinnse
and nou-iucrassate, resembling the tibiae of the females, while in other species the
tibiae are incrassate and densely hairy, bearing only a limited number of spines, it
is of great interest to observe that in P. vertumnus this specific difference breaks
down.
Though the tibiae of vertumnus are never much incrassate, they are densely
hairy in one geographical form, and almost normally spinose in the other forms.
c?. Palpus red, eighth abdominal sternitc edged with red. Forewing with
large glaucous green patch from inner margin forward beyond M-, sometimes
extending a little across M', the streak at inner margin sometimes reduced, seldom
absent, the patch variable in width, touching cell at least at base of M^, rarely so
much reduced as to be 1 or 2 mm. distant from cell. Red patch of hindwiiig
close to cell, triangnlar, consisting of three or four spots, the last (rarely the last
but one) being the longest, this last spot standing behind M- ; the spots small on
underside, spot R' — M' being placed about midway between cell and distal margin.
¥. Palpns and apex of eighth abdominal segment and of seventh sternite red.
Forewing with white or creamy patch. Red band of hindwing broad, consisting of
five to six spots, there being rarely a small seventh spot in front of R', the last four
spots more or less completely merged together, the scaling on veins M' and M^
being red, at least jiroximally, within the red band ; the band paler beneath, the
veins traversing it black, the last two spots alone confluent, rarely sejmrated like
the other spots.
Neuratiou : D'^ of hindwing transverse, short, being shorter than D^
Scent-organ : fold with broad streak of white wool ; scales at discal side of
wool grey, elongate, entire.
Genitalia : S. Tenth tergite one-fourth longer than the sternite, its lateral
edge dilated basally into a small ridge. Harpe as in P. lacydes, apex a little
more regularly rounded. ? . As in P. ei-laces, but autevaginal lateral ridge
membranaceous, wrinkled, not raised to a distant lobe ; edge of postvaginal plate
tuberculiibrm in middle.
Early stages not known.
Hub. Eastern slopes of the Andes from Colombia to Bolivia, extending
eastwards to Para and the Guianas ; not yet found in Venezuela.
a. P. vertumnus yuracares subsp. nov.
(J. Green patch of forewing touching cell between M' and M-, its proximal
edge between M- and inner margin slightly obliciue, somewhat concave, tlie patch
narrower in basi-distal direction than across veins; a white s])ot M' — M^, rounded,
in one specimen reaching from JI' to M-, a minute dot behind it iu two of our three
individuals; these white spots repeated on uudersurface. Hindiving with four
red spots, the upper two separate, the second rounded off proximally and distally ;
five small pale red spots on underside.
? not known.
( I'O )
Ihib. Eastern Bolivia.
In the Triug Museum 3 c?c?, from: Eucorado, January l'.)04, and Province
Sara, S. Cruz de la Sierra, March — April 1904 (J. Steinbach) ; Majiiri.
h. P. rertumnus autumnus Stand. (1S98) (PI. IV. fig. 3).
(J. rapiliii rcrliimiDis Cr. var. initumii)!>t Staudiuger, /)■/'.•.■ xi. p. 142 (1898) (Chanchamayo).
c?. Green jiatch of forewing larger than in the preceding, nsnally touching cell
also behind M-, often extended along cell as far as halfway between M^ and base ;
most specimens with one or two white elongate spots. Hindwing : red patch
consisting of three spots, large, the third spot reaching close to base of M-,
sometimes extending a short distance along cell ; four small red spots on underside,
sometimes a minute tifth before R', spot M' — M^ often obsolete.
?. Very different from the well-known females from the Amazons and Guiana.
Forewing : patch cream-colonr, very large, consisting of four spots ; a very large
cell-spot, occupying about half the cell, proximaliy edged with glaucous buff, a small
spot before II', a large spot, R^ — M', truncate, and another large spot M' — M-, as
long as the one before it, but not so broad, not quite extended to M-. Hindwing :
red band broad, consisting of six sjiots, one sjiecimen bearing a minute seventh
spot before R' ; on underside some of the spots shaded with black proximaliy.
Hab. Central East Pern : Ghanchamayo district, probably also farther south.
In the Tring Museum : 20 cJt?, 4 ? ¥, from: Chanchamayo (W. Hoffmanns ;
ScliunkeJ ; R. Toro, August — September 1901, and R. Perene, March I'.HJO (Simons).
c. P. vcrtumnus bogotainis Feld. (1864).
^. Papilio rertumnus var. logotanus Felder, Verh. Zool. Bot. Ges. W'len xiv. p. 292. sub n. 42 (18G4)
(Bogota) ; Maass. & Weym., in Stiibel, Reimi S. Amer., Lep. p. 31. n. 127 (1890) (Colombia).
S- Connecting the preceding subspecies with the following, (ireen patch of
forewing contiguous with cell between M' and M- ; no white spot. Red patch
of hindwing large, consisting sometimes of four spots ; the spots small on underside
as in the jjrcceding form.
Hab. Rio Palcazu and Pachitea in Peru, northward to Bogota, eastern slopes
of the Andes.
In the Triug Museum 9 c?c? from: Pozuzo, Huilnuco, and Rio ('hucliuras,
affluent of R. Palcazu (W. Hoffmanns) ; R. Mixiollo, Lorcto (Baer) ; Pachitea ;
Rio Caohyaco, affluent of R. Huallaga (Stuart); Archidona (H. llaensch);
Bogota.
d. P. rertumnus diceros Gray (1852).
?. PapiUo (Vrerm Gray, Cat. Lep. Ins. lirit. Mus. i. rap. p. 48. n. 23f.. t. 11. fig. 4 (18.52) (Para) ;
id., List Lep. Ins. Ilril. Mns. i. Pap. p. G3. n. 249(185(1) (Pani) ; Wall., Trans. Enl. Soc. Lonil.
(2). ii. p. 250 (1854) (Paril ; forest).
? . Pnpihn cirhis (Jray, Cat. Lep. Ins. Brit. Miis. i. Pap. p. 48. n. 237. t. 8. fig. C (18.52) ; id.,
List Lep. /ns. Brit. Mus. i. Pap. p. G3. n. 250 (185fi) (Ega ; vars. from R. Tapajos and Villa
Nova) ; Wall., Trans. Ent. Soc. Land. (2). ii. p. 25Ci (1854) (Upper Amazons ; forest),
fj. Papilir, rertumnus, Gray, Cat. Lep. Ins. lirit. Mus. i. Pap. p. 57. n. 20G (1852) ; Wall., Tram. Ent.
Sue. Luml. (2). ii. p. 255 (1854) (Pant ; forest) ; Gray, List Lep. Ins. Brit. Mus. i. Pap. p. (>9.
n. 281 (185G) (partim ; Panl) ; Mdnrtr., Ennui. Corp. Anim. .Mus. Pelrop., Lip. i. p. 5. n. 78
(18.57) (" Brazil").
(J. I'apilio rertumnus var. c. Pajiilio cutora Gray, Cat. Lep. Ins. Brit. .Mus. i. Pap. p. 58. sub n. 2Gl).
t. 10*. fig. G. cJ (1852) (J only ; <f alia spec.) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. fl9.
sub n. 281 (185G) (Ega; Villa Nova; "Brazil"); Bates, Trans. Kut. Soc. Lnml. (2). T.
( 471 )
p. 340, 355 (1861) {tiyn.jnrtim) ; id., Jmini. Bnlom. i. p. 225. D. 12 (18i;2) (partim) ; Felder,
Verli. Zuul. But. Ges. Wkn xiv. p. 292. n. 42 (1804) {pdrtini ; Amazons) ; Bates, Xahinil. Rh,
Ainaz. p. 26 (1864) (Parti, g in swampy shades, 5 in more open places) ; Butler, Oii. Diurn.
hep. descr. Fabric, p. 235. n. 6 (ISGD) (Panl) ; Oberth., El. d'J-Jiii. iv. p. 83. n. 274 (1880) (Pant ;
Obydos) ; Stand., E.rot. Tar/f. p. 13 (1884) (purlim; Amazons); Hahnel, Iris iii. p. 212
(18911) (Pani) ; id.,/.c. p. 240(1890) (Villa Bella, Amaz.) ; Michael, Iris vii. p. 213 (1894) (Sao
Paulo de Olivenfa).
Piipilio vrrtummis var., Gray, Cat. Lep. Ins. Brit. Utts. i. Pap. p. 58. sub n. 266 (1852) (Para) ; id.,
List Lep. Lis. Brit. Mus. i. Bap. p. 69. sub n. 281 (1856) (Para).
Papilio nttoraf Wallace, Trans. Knt. Soc. Loud. (2). ii. p. 255 (1854) (Amazons ; forest) ; Felder,
Vcrh. Znol. But. Ges. Wkn xiv. p. 292. n. 43 (1864) {partim).
S. Green patch touching cell between M' and M^, often a little separated from
cell, in many specimens not reaching M' ; white spots transverse, obliqne, n.snally
absent. Red patch of hindwing smaller than in tlie preceding forms, especially
the last spot; spots of underside usually larger than in the previous forms.
? . Forewing with chalky-white {)atch, consisting of two, three or four spots,
most specimens having a small spot in cell, a large spot R' — M', a small one before
W and a streak behind M'. A specimen from S. Paulo de Olivenca, in coll.
Oberthilr, has only a small double spot, divided by vein M', extending forward
and backward only to the middle of the cellules. Gray figures as cixitis a female
with a single, square spot R^ — M^ The only specimen which we have seen of
this form, besides the type, came from Surinam. Gray's sjjecimen was said to
be from Ega. It was bought from Stevens, the locality being perhaps erroneous.
Ilab. Amazons : from Para to Iqnitos.
In the Tring Museum 14 SS, 10 ? ?, from : Sao Paulo de Olivenpa ; Jnhuty,
April 1905, and Obidos, October — November 1904 (Mathan); Itaituba ; Manicore.
e. P. vertumntis vertumnus Gram. (1779).
PapUin Eqncs Trojanus rcrtumnns Cramer, I.e. ; Jabl. & Herbst, Naturs. Schmctt. ii. p. 61. n. 20.
t. 11. fig. B (1784) ; Esper, Ausl. Schmett. p 58. n. 25. t. 15. fig. 1 (1788) ; Fabr., Ent. Si/sl.
iii. 1. p. 16. D. 49 (1793) (Surinam).
Papiliii Eques Itomanus vertumnus, Jablonsky & llerhst, Naturs. Sehnielt. ii. p. 64 (1784).
Papilii) Eques Trajanus aeneas, Fabricins, Sjjec. Ins. ii. p. 8. n. 32 (1781) (partim) ; Gmelin, Si/st.
Nat. i. 5. p. 2233. n. 16 (1790) {partim).
Puridcs vertumnus, Hiibuer, Vers. bek. Schmett, p. 87. n. 911 (1818 ?).
Papilio vertumnus, Godart, E}ic. Meth. ix. p. 37. n. 38 (1819) {partim) ; Boisd., Spec. Gen. Lep. i.
p. 298. n. 129 (1836) (Cayenne ; Surinam); Lucas, in Gu(5r., Diet. Pitt. Hist. Nat. vii. p. 47
(1838) ; Doubl., Westw. & Hew., Gen. Diurn. Lep. i. p. 18. n. 200 (1846) (Guiana) ; Felder,
Vcrh. Znol. But. Gcs. ]Vien xiv. p. 292. n. 42 (1864) {partim- ; Surinam ; Cayenne) ; Kirby, Cat.
Diurn. Lep. p. 525. n.Ol (1871) (partim) ■ Staud., E.cnt. Ta.jf. p. 13 (1884) (partim ; Surinam) ;
Haase, Untcrsuch. Mimicry i. p. 79 (1893).
Papilio vertumnus var.. Gray, Cat. Lep. Ins. Brit. .Uus i. Pap. p. 57. sub u. 266 (1852) (Surinam) ;
id., List Lep. Ins. Brit. .Mus. i. Pap. p. 69. sub n. 281 (1856) (Surinam).
Papilio rertumnus var. dicervs, Mosohler, Verh. Znol. Bnt. Ges. Wien xxxii. p. 303 (1883) (Surinam).
S . As in the Amazonian form, but the mid- and hindtibiae short-hairy ; this
character not quite constant.
?. Forewing with the white patch as in the Amazonian form, or the patch
red need lo a single sjiot R' — BF, which is sometimes vestigial only.
Iliil). The (Jnianas.
In the Tring Museum 0 cjcj, 4 ? ?, from : Gamaria, British Guiana, January
1904 (R. Haensch) ; Essequibo R. ; Surinam.
A series of both sexes iu coll. Oberthiir from Maroui, French Guiana.
(472)
31. Papilio lycimenes Boisd. (187o) (PI. VI. fig. 31. 33. 34).
(J. Pajiilio li/rimciics BoiaduTal, Coiisid. Lep. Guatemala p. 7 (1870) (Costa Rica ; synon. excl.).
A near relative of P. vertummis. The ranges of the two species overlap in
Colombia. The forewing is shorter and proportionately broader, the green jiatch of
the forewing and the red patch of the hindwing of the male are ditfercntly shaped,
and the red spots of the underside of the hindwing, instead of being small, as in
the Colomliian form of vertumnus, are large, having also a different position. In
the female the wings are less deep black than in vertumnus, opaque, the sjwts of
the forewing are j-ellowish white, the cell-spot is transverse, reaching across the
cell, and the band of the hindwing is much paler both above and below. There
is hardly anything in structure by which P. bjcimenes could be distinguished from
all forms of P. vertumnus.
Besides a subspecies of /'. li/cimenes there are in L'olombia three more
Papilios with almost the same pattern. They occur in the same localities, and
are apparently quite distinct from one another, being independent forms — i.e. true
species. Leaving P. rertumnus apart, this insect being easily recognised, there
are fonr species generally mixed np in collections. At first sight one is inclined
to lake these insects for mere individual varieties of one species, the sjiecies
resembling each other so much, and each species being in itself so variable, that
only by a careful study of long series of specimens are we now enabled to draw
the lines of separation. If one has once understood that there are four species
ill Colombia occurring apparently everywhere together in suitable places (perhaps
with the exception of P. ancJdses serapis, which has not been found in West
('olombia), and if one has moreover grasped the distinctions between the species
in Colombia, it will be comparatively easy to separate into species also the material
from Central America, where the same problem obtains. Therefore we confine
our remarks in this place to the Colombian forms of the four insects in question.
Three of these insects are common in " Bogota " collections.
The males of the Colombian subspecies of these species are separable by
comparing the size of the red patch or baud of the hindwing and the structure
of the tibiae.
a. Tibiae simply spinose as in female.
a. Ked patch of hindwing large, there being always a
streak behind M^ .... A subspecies of P. bjcimcncs.
b'. Hindwing with three small spots, no streak behind
M-, or only a minute dot . . .A subspecies of P. eritkalion.
b. Tibiae and first tarsal segments dilated, densely hairy.
c. Hindwing below with a band of six or seven spots,
the band close to apex of cell . . Two subspecies of P. anc/iises.
d'. Hindwing usually with four or five spots on under-
side, spot M- (SM') much nearer to the distal
margin than to the ceil . . .A subsi)ecies of P. iphitlamas.
The females are best distinguished by the different relative size of the spots
of the forewing and the colour or width of the band of the hindwing.
c. Spot IV" — M' smaller than the spot in front of it ; cell-
patch large A subspecies of P. eritkalion.
( 473 )
d. Spot R' — M' larger than the spot in front of it.
e. Forewing semitransparent distally ; cell-spot large ;
band of hindwing entering cell, pinkish cream
proximally . . P.ancltises serapis s.nil i\. ^.vib'i]}. o? P. iphidamas
/'. Forewing opaqne, cell-spot large ; band of hindwing
less pale proximally . . . .A snbspecies of P. bjcimenes.
(/. Forewing opaqne, cell-spot narrower ; band of hind-
wing broad, entering cell, lieueath more rosy, and
black distal marginal area wider than in the
other females P. ancliises ah/attes.
For some other distinctions see the note nnder the respective heading of each
species. The geographical variability of the varions sjiecies is not the same.
Scent-organ and genitalia of P. bjcimenes essentially as in P. vertuinnus.
Early stages not known.
Ilah. Gnatemala to Ecuador.
a. P. li/cimenes bjcimenes Boisd. (1830).*
(J. Papilin lycimenes Boisduval, I.e. (Costa Rica).
(J ? . Papiliu qiliidiuiKis, Godman & Salvin (iinn Fabriciua, 179.3, err. det.), Biol. Centr. Aiiier., lihop.
ii. p. I!i2. n. 4. t. f.5. fig. 5, 5a. (J-fold, genit. (1890) {pnHim).
cJ. PaiiiVw ahjotlrs, iid. {iion Felder, 18G1, err. det.), U. p. 194. n. 5 (1800) (Panama).
S . Uppersiilc. — Forewing : olive-green patch limited behind by SM^, always
reaching this vein, often a small streak behind this vein, many specimens with
green scaling in cell ; a creamy spot R^ — M' present in almost every specimen,
being absent only in a few of the individuals from the southern limit of the range
((.'hiri(jui) ; a second creamy spot occasionally behind M', and often also a creamy
spot in cell. Hindwing : a baud of four, tive or six spots, narrowing costad, the
last sjjot standing behind M-, narrower than the last but one, but nearly as long.
Underside. — Forewing : white spots as above, many specimens with a dot
in cell and some ill-defined creamy scaling in front of cell. Hindwing with a
band of five to seven spots, the upper two small, sometimes ves(;igial, the first
occasionally absent.
? . Forewing : creamy white spot R^ — M' much larger than spot R- — -Rl
Hindwing : band broad, uniformly red or proximally a very little paler than
distally, its proximal edge slightly convex or straight in most specimens.
Hah. Gnatemala to Panama ; islands off the west coast of Panama.
Occurs together with /'. iphidamas, which it resembles. The male differs
from iphidamas in the green patch of the forewing reaching down to SM^ or
beyond, in the spot behind M- of the hindwing being much larger, and in the
tibiae being sjiinose. The female is distinguishable from the female of iphidamas
by the second discal spot of forewing being larger than the first and by the band
of the hindwing being less evenly curved.
In the Tring Museum: 42 c?cJ, 22 ¥ ? , from : Polochic Valley, Guatemala
(Salvin) ; San Jose, Costa Rica, 4000 ft., September 1004 (A. Hall) ; Carillo, Gosta
Hlca, 3000 ft., October 10o4 (A. Hall) ; Juan Vinas, Costa Rica, 2500 ft., October
1004 (A. Hall); Carillo, June— Jnly 10ii3 (Underwood): Carthago and Careblanco
In the bibliograpliy of this an<l the allied forms we have quoted only .such references as are
accompanied by a descriptiun ur figure, and a few others which we could verify by the examination of the
specimen.s referred to.
(474 )
de Siirajiiiini, Costa Rica (Umlorwouil) ; VuK;ui lU; Mirav;.IIi-n, Cosla Rica
(Uiidi'iwDnil) ; Cubacd 1., Brava 1., and Scvilla I., Jauiiary VM'Z (1. 11. Batty);
Bogava, C'hirinui, nOii ft. (Watsou) ; Colon.
/;. 2'. li/cimencs eri/thriis sub.sp. iiov. (1*]. VI. fig. ;53. 34).
$. Pajiilio zciu-is. Gray (iinn Lucas, 1852, err. det.), Cut. Lfp. Ins, Brit. Mas. i. Puji. p. Ii'i. ii. '2'M.
t <J. fig. 7 (18.52) (cJ alia specie.s).
$. Pa/)ilio crytUnlion (!), Wood (non Boisduval, 183G, err. det.), lux. Abmnd p. U'sl. fig. 302 (1883)
(Bogota ;— this species V)
This common Bogota insect has alwa3's been confounded with the Colombian
forms of 1'. iphidamas and P. anchises. The main differences from tliese insects
liave been mentioned above (}). 472).
(?. Upperside. — Forewing : olive-green patch mostly broader than in tlie
preceding form, and always reaching close to inner margin, bnt never entering cell ;
a small white spot present in some specimens, standing behind M', there being
occasionally also a tiny dot in front of M', this latter spot more often marked
on the underside. Hindwing : four spots, contiguous, the first more or less
rounded.
Underside: five, rarely six, spots on hindwing, paler than in the preceding;
sj)ot R- — R^ farther away from cell.
¥ . Upperside.— FoKvivag : discal spot R' — M' much larger than spot ll'^ — 11^,
and also larger than in the previous subspecies. Hindwing : band paler red than
in Central American females, touching cell or entering it ; spot SC- — K' small or
absent.
Underside : white upper scales of band of hindwing in most specimens
tridentate.
Ilab. Colombia : Magdalena and t'auca valleys ; Sta. Marta ; Northern
Venezuela.
Type : c? from Cnndinamarca.
Most Venezuelan specimens have two white spots on forewing, separated l)y
vein M>.
In one of our males, probably from Bogota, the patch on the upperside of the
hindwing is orange.
In the Tring Museum 90 S <S , 8 ¥?, from: Pereira, Canca valley; Mnzo,
December IS'.Hj ; Pnruio, Magdalena valley, October — November ItS'Jti (Dr. I'iirger);
Cananche, Cnndinamarca, July l'JU3 (Mathan); Pacho ; Tachira and Merida,
Venezuela (Briceno).
c. P. lijcimenes paraUus subsp. nov. (Pi. VI. fig. 31).
A very distinct, small form.
(J. Upperside. — Forewing : green patch as in crythras ; a round creamy sjiot
M' — M-. Hindwing : red patch smaller than in the preceding, consisting of
four spots ; first spot minute, sometimes absent.
Underside: wliite spot of forewing large, tonching both M' and M- ; four red
spots on hindwing, first sometimes absent ; no sjjot on abdominal fold.
?. Upperside: forewing less opaque than in the previous forms; sjiots purer
white; cell-spot reduced, triangular, being the smallest of tlie three spots jjreseiit,
rarely reaching halfway across cell; no spot between R- and R^ ; spot It'' — M'
triangular, being obliquely truncate distally, separated from the cell-spot by the
( 475 )
black vein ; spot M' — IP the largest, either oblong or proximally narrowed, being
reduced behind. Hiudwing : band rather more rosy than in eri/tJirus, mnch
narrower than the brown-black distal area, not entering cell, almost straight,
extending from abdominal edge to R^ or a little beyond.
U7iderside : white patches sometimes larger than above, and band of hindwing
paler, usually consisting of four spots R- — (SM'), the abdominal spot (SM')— SM^
being rarely present ; some specimens with a minute dot before W.
llab. West Ecuador : Guayaquil ; Chimbo ; La Chima.
In the Triug Museum 11 (?(?, 5 ? ?, from: Chimbo, 1000 ft., August ls97
(Rosenberg); Naranjas, Guayaquil (0. T. Baron).
32. Papilio erithalion Boisd. (1836).
J. Pa/)i7<o eW(/io//o« Boisduval, Spec. Gen. Lep. i. p. 295. n. 125 (183G) (Colombia ; "Jamaica"
false).
(J. Papilio pyrochhs Doubleday, A)m. Mag. N. H. xiv. p. 416 (1884) (Colombia).
(J. Tibiae spinose. No distinct red streak behind SP of hiudwing, usually
DO red scales whatever behind this vein.
?. Spot R^ — M' of forewing smaller than spot R^ — R^, often minute or
absent, or larger, but then band of hindwing wider than in the respective form
of lycimenes and proximally paler ; fringe-spots generally large.
Genitalia : S. Harpe less dentate than in P. lycimenes, more curved.
Early stages not known.
Hab. Costa Rica to Colombia and Northern Venezuela.
a. P. erithalion zeuxis Lucas (1852).
(J. PapiUu rham'ises Doubladay, List Lep. Ins. Brit. Mas. i. p. 147 (1845) (Venezuela ; iionicn
milium ; haeo species ?).
(J. Papiliu rhesus Kollar, Deiil.sckr. K. Ale. Wiss. Wien, Hath. Nat. CI. i. p. 35.3. sub n. 7 (1850)
(Klug iu litt. ; iudescr. ; haec species?).
(J. Papilio scnxis Lucas, Reo. Zool. (2). iv. p. 190 (1852) (Venezuela ; — coll. Oberthiir) ; Gray, Cat.
Lep. Ins. Brit. ilus. i. Pup. p. 46. n. 231. t. 9. fig. G. (1852) ((J only, ? alia species) ; Lucas, in
Ca.steln., Voi/. Ame'r. Siiil, Ent. p. 198, Lfp. t. 2. fig. 3 (18.57) (upper white dot e.xaggerated).
? . Papil'O erithalion, Gray {non Boisduval, lS:i6, err. det.), Cat. Lep. Ins. Brit. Mas. i. Pap. p. 46
n. 230. t. 10*. fig. 4 (1852) (Venezuela ; J alia species).
cJ. Papilio rhainascs Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 293. n. 58 (1801) (type : Gray, I.e.
t. 9. fig. 6 ; no description).
(J. Papilio rhesus Felder, /.<■. (sub synon.).
(J. Papilio abiliiis Felder, I.e. (sub synon.).
(J. Papilio rhamses (!), BoisJuval, Consitl. Lep. Gaatem. p. 7 (1870).
(?. Upperside. — Forewing : green patch strongly narrowing discally, rarely
extending forward a little beyond M' ; a rather large rounded creamy white spot
M' — M'-, often followed by a second spot and not rarely preceded by a third, the
■white spots being seldom missing. Hindwing : three, seldom two, small red
spots, sometimes a vestige of a fourth spot behind M^.
UndiTsidi' : forewing always with two or three white spots, one of tliem
usually grc) or vestigial. Hindwing usually with five spots, the central discal
ones often touching cell ; the spot on abdominal fold sometimes absent, a few
specimens bearing a sixth spot before R^.
?. AVe are not sure that the female figured by Gray really belongs to this
species. We have several specimens agreeing fairly well with the figure. The
second discal spot of the forewing larger than the first, as is the case also in
31
(476 )
the female of P. li/ciiiwncs eiyt/uits ; the band of the bindwiiig entering cell, being
jiroximally mucli paler than distaliy.
Ila/j. Nortbei'u Veneznela, and Colombia east of the (Jordillera of Bogota.
In the Tring Museum 23 JcJ, 5 9 ?, from : Cuca, Valeucia, Caracas, Puerto
Cabello, M^rida, and Cumana, in Venezuela ; Peperital to Bnenavista, Eastern
Colombia, January 189", 400— 13uO m., dry (Dr. Biirger).
b. P. eritkalion eritkalion Boisd. (183G).
?. PapiUo erithiilion Boisduval, I.e. ; Felder, Reise Novnro, Lep. p. 25. n. 15. t. G. fig. d (1SG5).
cJ. PapiUo 2>yrnrhles Doubleday, Ann. .Vaff. N. H. xiv. p. 4I() (1840) (Colombia) ; Gray, Cat. Lep.
Ins. Brit. Mus. i. Pap. p. 46. n. 229. t. 9. fig. 2 (1852) (partini ; ^ only),
cj. Papilio jiliaciion Kollar, Denkschr. K. Ah. Wiss. Wien, Math. Nat. CI. i. p. 353. n. 7. t. 42.
fig. 5. G (1850) (Cundinamarca).
? . PapiUo ahjattes Felder, Wim. Ent. Hon. t. p. 73. n. 7 (18G1) ( $ only).
cJ. Upperside. — Forewiug : olive-green patch extending from inner margin
to M\ seldom beyond, very often reduced and ill-defiued ; white spot rarely
present, standing between R^ and M' (not M'— M^ as in the Venezuela form),
usually narrow, oblique, and separate from the green patch. Hiudwing : three
red spots R- — M-, occasionally a vestige of a spot behind M-, sometimes a dot
S(J- — R', rarely another dot R' — R'-, and stiil more rarely a sixth dot before SC^
(a specimen in the Vienna Museum).
Underside : Forewing often with a greenish grey s2iot R' — M' in specimens
which have no white dot on upperside, preceded sometimes by a second dot; the
spot R' — M^ white in specimens with white dot on upperside ; sometimes a cloud
of whitish scales in front of cell, rarely condensed to a white elongate-triangular
spot. Hindwing nsually with five spots, which are on the whole farther away
from cell than in the previous subspecies, often seven spots, very rarely eight.
? . Uppjerside. — Forewing : spot R' — M' smaller than R- — R', many specimens
with small spots distaliy of apex of cell ; cell-spot large, close to cross-veins.
Band of hindwing very variable, spot R' — M^ longer than its distance from distal
edge of wing.
Hab. Colombia : Magdalena valley ; Cordillera of Bogota.
In the Tring Museum: 90 c?(?, 40 ? ? , from : Valdivia, July 1897 (Pratt);
La Palma, August, Guadalite, September, Cananche, July and August, Pizarra,
August 1903, (kindinamarca (Mathau); Muzo, January 1898 and December 1890;
Puniio, October — November It<9(3 (Dr. Biirger) ; La Vega, cast of Bogota, 19U0 m.,
January 1897, dry (Dr. Biirger) ; Pacho ; Villaviceucio.
c. P. eritkalion cauca Oberth. (18S0).
cj ? . Papilio eritkalion cauca Oberthiir, Et. d'Ent. iv. p. 84. sub n. 276 (1880) (Cauca).
J j . Papilio caiiea, Staudinger, Exof. Tag.f. p. 13. t. 9. i (1884) (Cauca).
While in P. bjcimenea and P. iphidamas the males from the Canca valley
have the green patch of the forewing as large as it is in the specimens from the
Magdalena valley, the Cauca males of P. eritkalion have the green patch absent
(jr vestigial. Tliis fact proves conclusively that P. eritkalion is specifically distinct
froni I lie other two insects.
(J. Uppcrgide. — Forewing : green patch absent or vestigial ; no white spot. •
Hindwing with three spots, mostly separated from one another, rarely preceded by
one or two minute dots.
Undermle. — I'orewiug not seldom with glaucous grey spot R' — M' and an
( ^<7 )
elongate-triaugular spot before cell. Hiiulwiiig nsnally with five spots, sometimes
with six or seven, distant from cell.
? . Upperside. — Forewing : cell-sjwt close to cross-veins, an elongate-triangular
spot in front of cell and several small ones on distal side of cross-veins ; discal
spots R- — B? larger than spot R' — M', the latter rarely touching M^ Hindwing:
band narrow, evenly curved, situated about halfway between cell and distal margiu.
Underside like ujiper, band of hindwiug paler.
Hab. Colombia : Cauca valley.
In the Tring Museum 30 cJcJ, 10 ? ?, mostly from Pereira.
In coll. Oberthiir a long series of both sexes from Manzales and Pereira.
d. P. erithalion sadifattes Druce (1874).
(J. Papilio sadi/aiies Druce, Eiil. Mo. Mag, xi. p. 36 (1874) (Costa Rica) ; Kirby, Cut. Diurn. Lrp.
p. 814. n. 389 (1877) ; Godm. & Salv., Biol. C'enti: Amei:, Rhop. ii. p. 195. n. 7. t. Cih. fig. 4
(1890) (Costa Rica) ; iid., I.e. p. 7-.'8 (1901) (Costa Rica).
Papilio iphidainas, iid,, I.e. p. l'J2. n. 4 (1890) (partim).
(S. Upper.side. — Forewing: olive-green patch very variable, either large,
extending from inner margin to R^ or reduced in length and width, or altogether
absent, our series showing all intergradations between the extremes ; black
specimens being known from Costa Rica and Chiriqui, in which localities occur
also sijecimeus with large olive-green patch ; all individuals with at least one
small creamy spot, most specimens with a large spot, the spot standing either
before or behind R^, there being often a spot at both sides of the vein ; sometimes
an olive-green streak in cell, but no creamy spot. Hindwing : three spots,
often preceded by one or two dots, there being occasionally also a minute dot
behind M^.
Underside. — Forewing : always one or two white or huffish white spots on disc,
and the majority of specimens with a sharply defined elongate-triangular spot in
front of cell.
? . Upperside.—YoxQvrwg : spot R^ — M' minute or absent ; in our only Costa
Rica specimen which we place here, spot R^ — R' small, oblique, a very little larger
than in the figure of d in Biol. Centr. Amer., I.e., the cell-spot of this specimen also
reduced, narrower, transverse, not reaching across cell, no other spots. Hindwing :
in the Costa Rica specimen the band evenly curved, separate from cell, half as
wide again as black distal border ; iu the specimens from the more southern
localities the band entering cell, twice as broad as the distal marginal border,
almost uniformly orange-red.
Hub. Islands off West t!oast of Panama; Chiriqui; Costa Rica.
The males with large olive-green patch on forewing are distinguished from
P. li/cimenes lijcimencs by the absence of a red streak behind M^ on the upperside
of the hindwing.
In the type of sadyattes the white dot on the upperside of the forewing stands
behind R'.
The males without olive-green patcli and the above-mentioned female with
reduced spots on forewing lead over to the uc.\t species, which has in both se.xes an
entirely black forewing, very few specimens bearing a creamy white dot R' — R^ .
In the Tring Museum 23 cjcj, 4 ? ?, from: Brava and Sebaco Is., January
l'JU2 (J. H. Batty) ; Boquete, Chiriqui, 2500 ft. (Watson) ; Carillo, Costa Rica,
3000 ft., October lt)04 (A. Hall) ; Limou, Costa Rica, October 1904 (A. Hall).
( 4T8 )
33. Papilio polyzelus Fekl. (1805).
(J. Pupilin ani-hhcs, Doubleday {iian Linnu, 1758, err. det.), List Lep. Jus. Brit. Mm. i. p. 12
(1845) (partim ; Honduras).
IPajiilii) (declur Bates, Tran.'i. Enl. Sm: Loud. (-2). v. p. .'i-ll, 357 (ISGl) (iioh,. iiiul. ; baec spec.?).
(J ?. PiipiVu puhjzeliis Felder, Verli. Zonl. Dot. Ge.i. Wien xiv. p. 29.3. n. 54 (18G4) (iiom. nuil;
Mexico ; Honduras) ; id., Mcise Nomra, Lcji. p. 24. n. 14. t. 6. fig. a. ^ (1805) (Mexico).
The differences between tliis insect and black specimens of F. erithalion are
very slight. A more exhaustive e.xploration of Nicaragua and Honduras may
possibly furnish material of specimens completely connecting P. e. sai/'/atfes with
F. poh/zelus.
The differential characters of the two subspecies of F. polyzelus are highly
interesting. "We have mentioned under P. tertumnus that one of the subspecies
of that species has the tibiae of the male hairy, while they are simply spinose in
the other subspecies. In F.poli/zelus we meet with the same phenomenon. The
males from Western Mexico have hairy and somewhat iucrassate tibiae, while
those from Eastern and South Mexico, Guatemala and Honduras have spiny tibiae,
both forms differing also somewhat in pattern. In a classification which is based
strictly on the quantity of difference, the forms of F. vertumnus and P. jwh/zdus
with hairy male tibiae would have to be treated as specifically distinct from the
forms with spinose male tibiae. But in a system based on true relationship (as far
as we are able to make out relationship from the morphological and biological
characters known) all the circumstances have to be taken into account. As the
spiny-legged P. hjcimenes and P, erithalion are very closely related to hairy-legged
P. ipliidamas, it is quite natural that the difference which is constant and specific
in these insects should appear in other species as a difference between geographical
forms, the development of a species into geograi)hical forms being the first step
towards the splitting up of one species into more species which can exist side by
side, no longer being separated geographically, as are the geographical forms.
(J ? . Sexes similar, female a little paler than male ; forewing rarely with
white dot R- — II' (some Honduras males), on underside occasionally grey scaling
in front of cell. Hindwing with red band from SO'- to abdominal margin, standing
much nearer distal margin than in the allied species.
Early stages not known.
Hah. Mexico to Honduras.
Two subspecies.
a. P. polyzelus polyzelus Feld. (1865).
jj. PapiUo anehiscs, Doubleday {non Linn<5, 1758, err. det.). I.e. (1845) {partim ; Honduras) ; id.,
Weslw. & Hew., Geii. Diurn. Lep. i. p. 19 n. 224 (1847) (partim) ; Gray, Cat. Lcp. Ins. Brit.
Mus. i. Pap. p. 64. n. 283 (1852) (syn, exol. ; Honduras) ; id., List Lcp. Ins. Brit. Mus. i. Pap.
p. 74. n. 299 (185G) (syn. excl. ; Honduras) ; Weidem., Proc. Ent. Sue. Philad. ii. p. 14G(18G3).
(?) Papilio alcctor Bates, I.e.
(J ? . Papilio poli/zHus Felder, U.rc. ; Kirby, Cat. Diurn. Lep. p. 527. n. Gig. (1871) (Mexico) ;
Oberth., El. d'Ent. iv. p. 82. n. 270(1880) (Mexico) ; Godm. & Salv., Biol. Centr. Amer., liliup. ii.
p. 195. n. 8. t. 05. fig. 0. gouit. (1890) (partim).
Papilio candezei Borre (Uelval ined.), C. H. Sue. Ent. Belt/, xxviii. p. 120 (1884) (= 2'oli/ielus).
<J ? . Tibiae of male spinose, non-incrassate.
Genitalia: <J. Harpe acuminate, no teeth between ventral conical tooth and
apex.
Uab. Eastern Mexico, southwards to Honduras.
(479 )
In the Tring Museum 44 SS, 10 ??, from: S. Pedro Sula, Honduras;
Escuintia, W. Guatemala, 1100 ft., August 1904 (A. Hall); Mazateiiaiiga, W.
Guatemala, 1000 ft., September 1904 (A. Hall); Salia, Vera Paz, Guatemala
(Champion) ; " Mexico" ; Coatzalcoalcos, July 1004 (A. Hall) ; Orizaba.
b. P. poli/zelus trichopus subsp. nov.
(J ?. Pap'iUo pohjzelus, Godm & Salv., I.e. (West Mexico).
t? ? . Til)iae and first tarsal segments hairy and somewhat incrassate. Spots
of hindwing on the whole larger and nearer to the margin than in the preceding,
the baud being in one of the females only 2i mm. distant from cell, in male usually
a small spot behind M-. The posterior marginal spots of fore- and hindwing often
edged with red.
Genitalia : S . Harpe distally broader than in the preceding, denticulate.
Hub. West Mexico : Guerrero ; Michoacan ; Jalisco.
In the Tring Museum 04 (Jc?, 2o ? ¥, from : Guerrero, type (0. T. Baron) ;
Patzcnaro, Michoacan ; S. Sebastian (Dr. Buller).
34. Papilio ipMdamas Fabr. (1793). (PI. IV. fig. 7.)
$. Papilio Equi's Trojanua ijihidntna^ Fabrieius, Eiil. Stfsf. iii. 1. p. 17. n. 52 (1793) (type: Jones's
drawing).
c?. Tibiae and first tarsal segments somewhat incrassate and densely hairy.
?. Forewing : spot R^ — RMarger than R' — M' (Central American form), or
smaller (South American forms). For differences between the Colombian females
of this species and the females of the allied insects see p. 472.
Genitalia: c?. Harpe more curved than in P. lijcimenes and erithalio7i, besides
the large ventral tooth with several small teeth, which are variable in size and
number.
Hab. Mexico to Ecuador anil North Venezuela.
The drawing of Jones, upon which the name iphidamas is based, represents
in our opinion a Central American female of the present species. The band of the
hindwing is too uniformly red for a South American female of this or any allied
species. The P. idaeus of Fabrieins, likewise described from Jones's drawing, is
also a Central American form.
As only P. iphidamas is known to extend into Mexico as f;ir north as Vera
Cruz, there can hardly be any doubt that the Central American males are mated
correctly with the females described below. It is interesting to observe that the
proportional size of the two discal sppts R^ — M' of the forewing (?) is reversed in
the forms from Colombia and Ecuador, as is the case also in several other Papilios.
a. P. iphidamas iphidamas Fabr. (1793).
? . Papilio Eques Trojnnus ipliidumas Fabricius, I.e. (no locality given).
?. Piij/ilio ipliidamuii, Godart, Ew. Miil/i.ix. p. 37. n. 34(1819) (copied from Fabricius) ; Boiad. , ,5y>cc.
Gin. Lip. i. p. 292. n. 121 (1830) (copied from Fabricius) ; Doubl., Westw. & Hew., Gen. Diiirn.
Lrp. i. p. 19. D. 220 (1847) (" S. America " false) ; Gray, Cat. Lep. Inn. Brit. Mas. i. Pap. p. 44.
n. 225. t. 8. fig. 1. cj, 2. ? (18.52) (Honduras.; Mexico) ; id., List Lep. Jus. Bill. Man. i. Pap.
p. liO.n. 238(1850) (Mexico; Honduras); Bates, Tmii<!. Ent. Son. Lond. (2). v. p. 341, .357
(1801) (Mexico; Honduras); Weidem., P,-uc. Eat. Soc. Philail. ii. p. 147 (1803) (= a;r«s,
fa!se) ; Felder, Vrrh. Zoul. Dot. Gck. Wieii xiv. p. 293. n. 53 (1864) (Mexico; Nicaragua;
Honduras ;/«)•(,))!.?) ; Butler, C'««. £)('«)■«. Lrp. dcscr. Faliric. p. 23G. n. 8 (18G9) ("Bogota
specimen agreeing with Joues's figure," false) ; Godm. & Salv., Biol. Ceiitr. Aiim-., Rtinp. ii.
p. 102. n. 4. (1890)(yinrt(«().
( 480 )
{J. PapiVio areas, Doubleday (mo?; Cramer, 17f'2, err. det), Lht Lep. hm. Brit. Mtis. i. p. 12 (1845)
(" S. America '' false).
? . PapiUit paimres Gray, Cat. Lep. Ins. Dril. Miis. i. Pup. p. 45. n. 226. t. 10. 6g. 4 (1852) (Mexico) ;
id., List Lep. ln.«. Bril. ilim. i. Pap. p. fiO. n. S.^O (ISSC) : BatPS, Trams. E,d. See. Lmul. (2). v.
p. 341, 357 (1861) {parlim ; (J alia spec.) ; Weidem., Pmc. Ent. Soc. Philad ii. p. 147 (1863);
Felder, I.e. p. 294. n. 67 (1864).
Papilio serajiis, Mi'>netries (ho« Boisduval, 1836, err. det.), Eniim. Corp. Anim. ilus. Pelntp,. Lep. i.
p. 5. n. 76 (1857) (syn excl. ; haec spec. V).
(J ? . PapiUo eehehis, Reakirt {71011 Hiibner, 1806—, err. det.), Proc. Ent. Soe. Phihul. ii. p. 138. n. 7
(1863) (Nicaragua ; syn. excl. ; haec spec. ?).
tj. Papilio aeheloiis Hopffer, Slelt. Ent. Znt. xxvii. p. 22 (1866) (Central America).
? . Papilio vcrliimnus var, h. P. ijjhidamas, Kirby, Cat. Diurn. Lep. p. 526. sub n. 61 (1871) (" New
Granada " false).
(J. Papilio a-vlfpiu.i, id.. I.e. p. 537. n. 126 (1871) {partim ; achelous Hopff. sub syn.).
(J. Papilio ineandescens Butler, Trans. Ent. Soc. Loml. p. 433. t. 6. fig. 1 (1874) (■' Para " false ;
this subsp. according to type in coll. Godman).
jj J . Papilio lyeimenes, id. & Druce {non Boisduval, 1870, err. det.), Pror. Zool. Soc. Loml. p. 363.
n. 362 (1874) (Costa Rica).
(J ? . Papilio abjattes, Staudinger {non Felder, 1861, err. det.), Ejcot. Tagf. p. 13. t. 8. J ? (1884)
(this locality ? this species ?).
S. Upperside. — Forewing : apex not semitransparent ; green patch usually
ranch reduced, seldom extending to hindmargin, being on the whole longer in
the southern than in the northern specimens ; always one or two creamy white
spots, often a white spot in cell. Hiudwiug : band gradually widening behind,
extending usually from SC" to M-, with a narrow streak behind M-, the first spot
often small, rarely absent.
Underside. — Forewing always with white creamy spot R^ — M', mostly also
with spot R- — R^ and a cell-spot, the latter often reaching across cell. Band
of hindwing from SO- to anal angle, widest between R- and M-.
?. Upperside. — Forewing: cell-spot close to cross-veins, square or oblong;
a triangular streak in front of cell; discal spot R' — M^ smaller than R- — R^,
often absent, rarely larger, but in this case gradually tapering proximally, its
oblique hinder edge being continuous (or almost) with the proximal edge of the
cell-spot ; usually one or more dots at distal side of cross-veins. Hindwing :
band nearly uniformly red, variable in width, but always nearly evenly curved.
Ilab. Southern Mexico to Panama.
Some of the southern individuals leading over to the next form.
In the Tring Museum 73 S S, 30 ? ?, from : Coatzacoalcos, Mexico, at sea
level, July 1904 (A. Hall); Escuintla and Mazatenanga, W. Guatemala, 1100
and 1000 ft., August and September 1904 (A. Hall); S. Pedro Sula, Honduras;
Nicaragua; Jnan Vinas, 2500 ft., August, Carillo, 3000 ft., October 1904, Costa
Rica (A. Hall) ; Juan Viuas, Escazu, Limon, San Jose, and Tarbaca, Costa Rica
(Underwood); Bogava, 800 ft., and Boqnete, 3500 ft., Chiriqui (Watson); Brava,
Cebaco, and Sevilla Is., Jannary 1902 (J. H. Batty).
b. P. iphidamas plwlins stibsp. nov.
$. Upperside. — Forewing: olivaceous green patch from hinder margin to M'
or beyond, widest behind, the streak along hinder margin being rarely somewhat
reduced in length and width ; no green scaling in cell ; no white sjiot, except in a
very small percentage of specimens, the spot standing usually behiud M'.
Hindwing : throe red s])ots, separate from cell, often a minute streak or dot behind
M-, rarely a dot in front of R-, sometimes only two spots present.
(481 )
Underside. — Forewiug often with a greyish white spot across M' or in front
of M-. Hindwing with three spots 11^ — M', a smaller one behind M' and usually
a minute dot on abdominal fold.
? . Uppcrsidc. — Forewing : apical area slightl)' transparent, being visibl}' less
opaque than in the Colombian subspecies of P. lycimenes and P. erithalion ;
cell-spot verj' large, longer than broad, often somewhat reduced in width costall_y,
in this case not quite reaching across cell ; subcostal streak often absent, discal spot
R'— M^ much larger than spot R- — R^ Hindwing: band very broad, entering
cell, very pale proximally, spot SO' — R' small or absent in most specimens.
Underside : spots of forewing purer white, and band of hindwing distally
brighter red than above.
I/ak Colombia : Blagdaleua valley, and Cordillera of Bogota.
In the Tring Museum 30 c?c?, 13 ??, from: Cananche and La Palma,
Cundinamarca, July, August and September 1903 (Mathan) ; Muzo, December
1896; Pacho ; Peperital to Buenavista, 400 — 1300 m., January 1897, dry season
(Dr. Biirgcr) ; Villavicencio to Monte Redondo, March — April 1897, beginning of
wet season (Burger).
c. P. ipJddamas elatos subsp. nov.
S. Like the preceding ; green patch of forewing more olive, duller in tint,
posteriorly reduced, the streak behind SM^ small, the patch being rhombiform, with
the upper proximal and posterior distal angles strongly rounded. Three small
spots on hindwing.
On underside five spots R^ — SM^ on hindwing, all well separated from one
another, spot R^ — M' the largest, being about half as long again as broad.
Hab. Cauca valley.
In the Tring Museum 1 S from Popayan.
d. P. iphidamas calogyna subsp. nov. (PI. IV. fig. 7).
(?. Upperside : forewing a little less opaque in apical area than in phalias ;
olive-green spot abont the same in size, on the whole somewhat smaller, duller
green ; many specimens with a white spot M' — BI-, which is often large, touching
both veins, being sometimes preceded by a small dot. Hindwing : three small
red spots, contiguous, often a minute dot behind M-.
Underside : forewing with white or greenish buff, distinct or vestigial, spot
M' — M*, or R''— M', or a double-spot across M', or the spot larger, extending
from M- forward to R-. Hindwing : five spots from R- to anal angle, last one
often absent, the spots on the whole less pale and smaller than in P. i. pkalias, the
upper three close together.
? . Upperside .• forewing : apical area slightly transparent ; cell-spot large,
but often reduced costally, in this case not reaching across cell ; subcostal streak
present or absent ; two discal spots R'^ — M', the second the largest. Hindwing :
band from R- to near abdominal edge, often a detached dot before R', the band
bright red, usually almost white proximally, the two colours contrasting strongly,
inner edge of band almost straight, but more or less curved distad before abdominal
margin, sometimes also incurved at apex of cell, in many specimens band entering
apex of cell; width of band variable, but not exceeding (or very little) the width
of the distal marginal area.
( 482 )
Underside: spots of forewing a little larger than above, cell-spot reacbing
across cell in all specimens, subcostal streak present. Hindwing : band paler
than above, narrower posteriorly, spot SC-— K' nearly always indicated.
Hub. West Ecuador and West Colomliia.
Tbe West Colombian males have rarely a white spot on the forewing, being
hardly distinguishable from jjlialias, while the females agree well with Ecuadorian
ones.
In the Tring Museum 20 S S , 10 ? ?, from: Paramba, 3500 ft., Febrnary,
March and April 1S97, dry (Rosenberg); Chimbo, luoii ft., August I8'.)7 (Rosen-
berg) ; Cachabi, low country, January 1894 (Rosenberg) ; Zaruma, Jnne 1899,
1000 m., wet (Simons). 13 JcT from Rio Dagua, W. Colombia (Rosenberg).
e. F. iphidamas tene.ates subsp. nov.
(J. rupilio osi/iv's, Godmau & Salv. (»oh Felder, 1861, err. det.), Trmis. Eiil. Sac. Loin], p. 121). n. 231
(1880) (Sta. Marta).
<S. Upperside, forewing: apical area usually rather more transparent than
in P. i. phalias ; green patch narrow, separate from cell ; most specimens with one
or two white spots. Hindwing with three red spots, separate from cell, often a
small spot behind M', sometimes an additional dot behind R'.
Undeiside : forewing with one, two or three white spots. Hindwing with four
to seven spots in an almost straight row, somewhat resembling the band of ahjattes.
? not known.
Hah. North Venezuela, and Santa Marta, Colombia ; name-type from Cucnta,
Venezuela.
Resembling alyattes in being rather more glossy blue on the hindwing than
in P. i. phalias, in the green patch of the forewing being reduced in width, the
streak along inner margin being often obsolescent (t'/pt'), and in the forewing
bearing often rather large white spots. The hindwing, however, is not so glossy
as in alj/attes, the spots of the underside are less close to cell, and the spot behind
M* of hindwing is smaller. The harpe is also somewhat diiferent, being narrower
and more curved, agreeing with that of 7'. /. phalias.
In the Triug Museum 9 c?c? from : Cncuta and Porto Cabello, Venezuela;
Onaca, Santa Marta, 2200 ft. (Chas. Engelke) ; R. Hacho, Santa Marta, March 1898
■^Wilmot Brown).
35. Papilio anchises L. (1758) (PI. IV. fig. 8. 12.; VII. fig. 44—47).
5. Papilio EquPi Tnijii/iiis aiirhisex Linm', Sysl. Xul. ed. x. p. 4G0. n. 10 (1758) (cit. exceptis) ;
Clerck, Icon. Ins. ii. t. 2d. fig. 1 (17(U).
(J ? . Papilio anchises, Boisduval, Spec. Gen. Lip. i. p. 201. n. 119 (18I5G).
cJ. Tibiae, and first tarsal segments somewhat incrassate and densely hairy.
Apical area of forewing semitransparent. Hindwing strongly glossy Idue on
disc ; red sj)ots rather closer to cell than in P. i. phalias, variable in number
and in size, often forming a continnous baud, many specimens bearing a small .-ipot
behind M-. Spot M*— (SM') of underside of hindwing rather larger and more
proximal than in P. iphidamas, the band of spots extending usually forward to SCI
? . Apex of forewing somewhat transparent ; discal spot R- — R' smaller than
spot R'— M'.
Early stages only known of P. a. orbigiiijanus.
Hab. South America, from Colombia to Par.'i, Sao Paulo, Bolivia, and Paraguay.
Not known from Pern,
( 483 )
We arc not snre that the forms here united under P. anchises are specifically
distinct from tliose treated as subspecies of P. iphidamas. It is very well possible
that all these forms of which the males have hairy tibiae are only geographical
varieties of one single species. Onr knowledge of the distribution of these PapiUos
in Colombia is very imperfect, and the material examined too scanty for the purpose
of deciding the question.
a. P. anchises ah/aftes Feld. (1861).
(J ? . PapiUo alyaites Felder, Wim. Ent. Mon. v. p. 73. n. 7 (18G1) (parlim, ^ only ; Bogota) ; id.,
Verh. Zool Bot. Ges. Wicn xiv. p. 29,3. n. 57 (1864) ; id., Rase Novam, Lep. p. 2G. n. 16. t. 0.
fig. e. (?, f. ? (1865).
Pajnlin vnytiiimiiis Tar. k. P. alyaites, Kirby, Cat. Diiirn. Lep. p. 52G sub n. 61 (1871) (New Granada).
(?. Upperside : forewing distally somewhat transparent, bat less than in the
following forms ; green patch reduced, not touching cell, streak behind SM- often
ve.stigial ; always at least one white spot, sitnated between M' and M-, often a
second spot before M', some specimens having also a dot behind M°. H.indwing
much more strongly glossy bine than in all the allies, the scales on the disc being
entire ; three red spots, the last usually the largest, often a minute spot behind M^
and another before W.
Underside : red baud of hindwing extending forward to SC°, spot M^ — (SM')
large as compared with the respective spot in P. iphidamus phalias.
? . Upperside : forewing less opaque distally than in P. iphidamas ; cell-spot
transversely longer than broad, narrower than in P. i. phalias ; two discal sjjots,
the second larger than the first, separated from cell or touching it behind
R^ Hindwing : band crossing apex of cell, almost gradually widened behind,
reaching forward to SC^ its distal edge farther away from margin of wing than in
P. iphidamas.
Genitalia : <S. Harpe decidedly broader and less curved than in P. iphidamas,
the ventral median tooth smaller, being sometimes not larger than the other teeth.
Hab. Colombia : Magdalena valley, probably on both sides of the Cordillera of
Bogota.
In the Tring Museum : 13 J J, 5 ? ? , from " Bogota."
h. P. anchises serapis Boisd. (1S36).
(J. PapUio serapis Boisduval, Spec. Gen. Lep. i. p. 298. n. 130 (1836) (Colombia) ; Doubl., Westw. &
Hew., Gen. Ditirn. Lep. i. p. 18. d. 196 (1840) ; Gray, Cat. Lep. Lis. Brit. Miis. i. Paji. p. 45.
n. 227 (1852) (partiin ; (J only ; $ alia species) ; id.. List Lep. Lis. Brit. .^U3. i. Pap. p. 01.
n. 240 (1856) (^w)-/;™) ; Feld., Verh. Zool. Bnt. Ges. Wieu .\iv. p. 249. n. 65 (1864) (y;«rt;/») ;
Kirby, Cat. Diiirn. Lep. p. 526. n. 61 d (1871) (partim) ; Godm. & Salv., Trans. Ent. Sac. Land.
p. 126. n. 282 (1880) (Sta. Marta) ; Prinz. Theresa, Berl. Ent. Zeilschr. xlvi. p. 241. u. 4 (1901)
(La Popa, Cartagena, August, 2 (J ^J ;— we have seen one of them).
c?. Upperside: forewing rather more transparent distally than in a('/«/^e« ; green
band very narrow, usually extending forward to R^ a little wider behind than in
Iront. Hindwing : band consisting of five spots, there being usually a sixtli
minute spot or narrow streak behind M".
Underside : the band of the hindwing as in abjattes.
?. Upperside: forewing more transparent distally than in abjattes; spots
more yellow, as are also the fringe-spots on both wings ; cell-spot snbtriangiilar,
not reaching across cell, no spot in front of cell ; discal spot U^ — M' very much
larger than spot R^ — R\ but much smaller than the cell-spot. Hindwing : baud
( 484 )
very ])roa(l, Imff or yellow proximally, strongly imlmate, its inner edge crossing
cell proximally of base of IM'-, spot SC-— R' larger than in ah/attes ; one of our two
specimens with a dot before SC.
Underside similar to upper, band of hiudwing paler red.
The two specimens here described have the appearance of being killeil too soon
after emergence, which may account for tlie yellowisli colonr of the markings.
Ilab. Northern Colombia : Santa Marta ; Cartagena.
In the Triug Mnsenm 2 JcJ, 2 ? ? , from " Bogota."
c. r. iiiic/ii.scs osi/ris Feld. (1861).
?. Papilio eritlmVon, Kollar (iim Boisduval, 18.%, err. det.), Denkxrhr. K. Al. Wiss. Wieii, ilatli.
Nat. CI. i. p. ."iSS. n. G (1850) (Angostura).
Papilio protem, Hewitson (hoh Boisduval, 1836, err. det.), Tnnis. Ent. Sue. Loud. (2). i. p. 97 (1851)
( (J of areas, Venezuela !).
(J. PajiUio erUhalion, Gray, Cat. Up. Ins. Brit. Mus. i. Pap. p. 46. n. 2.30. t. 10*. fig. 3. ^ (18.')-i)
(Venezuela ; synon. excl.) ; id., List Lep. Tits. Brit. Miis. i. Pap. p. 61. u. 243 (18,56) (partim).
(J ?. Papilio nsyris Felder, Wifii. Ent. Man. v. p. 74. n. 8 (1861) (Caracas) ; id., Verh. Znol. Bol.
Ges. Wieii xiv. p. 294. n. 66 (1864) ; id., lieise Novara, Lep. p. .31). t. 9. fig. b. <?, o. d. ? (1865).
$ . Papilio xenares id., Verh. Zool. Bot. Ges. Wim xiv. p. 294. n. 59 (1864) (Orinoco ; nom. nov. loc.
•' erithnlion KoU.") ; id., Reise Novara, Lep. p. 28. n. 17. t. 8. fig. a (1865) (Angostura).
(J. Papilio to.raris id., Verh. Zool. Bol. Ges. Wieii xiv. p. 294. n. 61 (18G4) (nom. nov. loco erUhalinn
Gray, I.e. t. 10*. fig. 3 ;-"f. 4" laps. cal.).
(J ? . Papilio sererus id , Verh. Zool. Bot. Ges. Wien xiv. p. 291. sub n. 66 (1864) ('■ Moritz in litt." ;
= osi/ris).
Papilio areas var. a. P. xeiiarrs, Kirby, Cal. Diiirn. Lep. p. 526. sub n. I'.la (1871).
Papilio toxaris, id., I.e. n. 61b (1871).
Papilio serapis var. a. P. osyris, id., I.e. sub n. 61d (1871).
Papilio areas, Hahnel (no,, StoU, 1782, err. det.). Iris iii. p. 138(1889) (San Eate'ban, in forest).
(?. Similar to .serapis, bands of forewing and hindwing broader ; white sjmts
of forewing usually large, sometimes absent, occasionally a white spot in cell.
?. (!ell-spot of forewing reaching usually across cell, many specimens with a
subcostal streak, two discal spots, R-— M', the second much the larger. Band of
hindwing only very little paler proximally than distally, usually entering apex of
cell, variable in width, spot SG^— 11' rarely small.
Haft. Venezuela.
One single female from Ciudad B(divar has the spots of the forewing pure
white, while in the type of xcnares from the same place they are only a little less
huffish than in the more northern individuals. It is possible that these two
specimens represent another subspecies ; however, more material is wanted to
decide the point. The sjiecimens from Caicara and the Caura River belong to the
next form.
In the Tring Museum 0 c?c?, 0 ? ? , from : Cucuta ; Merida ; Puerto Cabcllo ;
Ciudad Bolivar.
In coll. Oberthiir fl S 6 , A ? ?, from San Esteban.
d. P. anchises cymochles Donbl. (1844).
(J. Papilio cymochles Doubleday, Ann. Mag. N. H. xiv. p. 416 (1844) (Trinidad) ; id., Lv>l Lep. Im.
Brit. Miis. i. p. 12 (1845) ; id., Westw. & Hew., Gen. Dinrn. Lep. i. p. 18. n. 205 (1846) ; Gray,
Cat. Lep. Ins. Brit. Mas. i. Pap. p. 50. n. 245. t. 10. fig. 8 (1852) (Trinidad) ; id., Li.it Lep. Ins.
Brit. .Mas. i. Pap. p. 64. n. 260 (1856) (Trinidad ?) ; Bates, Trans. Ent. Soc. I.ond. (2). v. p. 341,
357 (1861 ) (Trinidad ?) ; Feld., 1 \rh. Z,;,l. Pot. Ges. Wien xiv. p. 294. n. 63 (1864) (Trinidad ?) ;
Kirby, Cat. Diurn. Lep. p. .526. n. 61e (1871) (Trinidad).
(J. Papilio nuaeharsis Felder, I.e. n. 62 (1864) (hab.1 ; nom. nutl.) ; id,, Heise Novara, Lep. p. 29.
n. 18. t. 7. fig. d. (186.5) (hab. ?).
(485)
tJ. Piijiiliu to.mris var. a. P. aiiacharsis, Kirby, l.r.
Papilio xciixk (!), Kaye, Proc. Eiit. Soc. Laml. p. 19 (1901) (= nli/ulle.n, both bred from one ^ ;
error of identification).
Papilhi zeti;ris, id., Trans. Ent. Soc. Land. p. 206. n. 194 (1904) (Trinidad ; " larvae on orange ! ! "
fal-se. — These larvae belonged probably to P. anchisiades).
PapUh cymochs (!), id., I.e. n. 195 (1904) (Trinidad).
S. Similar to os'/ris ; band of forewitig on the whole rather narrower, at least
in many specimens, with one, two or three white spots ; hiiidwing nsually with
three red spots, often preceded by a small sjiot and followed by a minute streak.
On underside most specimens with fonr spots on hindwing, often preceded by a
small fifth, bnt one of onr sjiecimens with seven spots.
?. Spots of forewing pnrer white than in the previous forms, the discal ones
differently placed ; cell-spot small, triangular, not reaching halfway across the cell,
often a mere streali ; two discal spots 11^ — M-, the second extending farther distad
than the first, occasionally a minute dot before W. Hindwing : band almost
evenly red, widest in middle, no spot before R'.
On underside the last spot of the band of the hindwing much reduced, some-
times minute, usually isolated.
Ilab. Trinidad ; Peninsula of Paria ; Orinoco.
In the Triug Museum 24 c?c?, 17 ? ?,from: Caicara, Orinoco, March 1897,
May and July 1898 (Cherrie) ; Snapure, February 1899, La Vuelta, May 1904,
t'orosito, June 19U4, Caura R. (S. M. Klages); Patao, Guiria, August 1891 ; various
places on Trinidad, December, January and February.
e. P. anchises anchises L. (1758) (PI. IV. fig. 8. VS).
% . Papilio Eques Trojaniis anchises h\nu6, Syst. JVat. ed. x. p. 460. n. 10(1858) (cit. excl.) ; Clerck,
Iron. Lis. ii. t. 29. fig. 1 (1764) ; Linn^, Miis. Lvd. Ulr. p. 191. n. 10 (1764) ; id., Syst. Nat.
ed. xii. p. 746. n. 11 (1767) ; Fabr., Spec. Ins. ii. p. 7. n. 26 (1775) (cit. excl.) ; Goeze, Ent.
Beytr. iii. 1. p. 34. n. 11 (1779) (cit. excl.) ; Fabr., Spec. Ins. ii. p. 7. n. 26 (1781) (cit. excl.) ;
Gmelin, Syst. Xut. i. 5. p 2230. n. II (1790) (cit. excl.) ; Fabr., Ent. Syst. iii. 1. p. 13. n. 40
(1793) (cit. excl.).
Papilio Eques anchises Linne, Syst. Nat. ed. Lange, p. 460. n. 10 (1760).
Papilio (Troes) anchises, Miiller, Nutiirs. v. 1. p. 669. n. 11 (1761) (larva exclusa).
(J. Papilio Eques Trojanus vertummis Cramer, Pap. Exot. iii. p. 32. t. 211. fig. C (1779) (Surinam ;
non fig. A. B.).
Priamides vertumnus, Hiibner, Verz. hek. Schmelt. p. 87. n. 911 (1818 ?) (partini).
Pajiilio vertiiinniis, Godiitt, Enc. Meth. ix. p. 37. n. .38 (1819) (jiartim) ; Boisd., Sjiec. Gen. Lip.\.
p. 298. n. 129 (1836) (parOm).
cj ?. Fupilio anchises, Boisduval, I.e. p. 291. n. 119 (1836) (Surinam) ; FelH., Vei-h. Zool. Dot. Ges.
Wien xiv. p. 293. n. 44 (1864) (Surinam) ; Guen., Ann. Soc. Ent. France p. 309 (1867) (descr.
of $ ) ; Auriv., Kongl. Sv. Vet, Ah. Ilandl. xix. 5. p. 15. n. 10 (1882) (recensio critica ; descr.
of $ ; probably type specimen).
Papilio vertumnns var. a., Gray, Cat. Lep. Ins. Brit. .Mus. i. Pa2>. p. 57. sub n. 266 (1852) ; id.. List
Lep. Ins. Brit. Mus. i. Pap. p. 69. sub n. 281 (1856).
(J. Papilio telmosis Bates. Trans. Ent. Soc. Land. (2). v. p. 340, 366 (1861) (Surinam ; type : Cramer's
fig. C, I.e. ; " extends into Columbia " false) ; Feld., Ve7-h. Zool. Bot. Ges. Wien xiv. p. 293. n. 45
(1864) (Surinam) ; Kirby, Cat. Diurn. Lep. p. 528. n. 62 F (1871) ; Moschl, Verh. Znol. Bot.
Ges. Wien xxvii. p. 295 (1876) (Surinam) ; Oberth., Et. d'Enl. iv. p. 82. n. 273(1880) (Gnyane ;
variability ; = eteocles).
(J. Papilio eteocles Felder, Verh. Zool. Dot. Ges. Wien xiv. p. 293. n. 49 (1864) (hab. ? ; nom. nud.) ;
id., Rcise Nomra, Lep. p. 22. n. 12. t. 7. fig. e. ^ (1865) (bab.?).
Papilio cyphotes var. a. P. eteocles Kirby, I.e. p. 527. sub n. 62 d (1871).
cJ. Upper.vdL'. — Fovc-<N\-Qg: band very variable, extending from M' to inner
margin, often being reduced to a narrow strgak, not rarely altogether absent, all
( 486 )
intergradations occnrring ; it varies also in colour, being sometimes bluish, sometimes
greenish ; a white spot M' — M- in some specimens. Hindwing strongly dentate,
mostly with three spots, which are nsnally well separated from one another and
from cell, there being often a streak behind M- and occasionally a dot before R- ;
the spots sometimes yellowish.
On ?/;ir/«'.s/V/e of hindwing, a row of five to seven spots, nsnally j)laced rather
nearer the distal margin than the cell.
?. U//persi(/e.— Forev/iug: no spot in cell or only a small streak, two spots
R' — M- on disc, the upper touching cell, or only one spot R^ — IP, touching cell or
separate from it, or this spot vestigial, or the wing without any spots, except the
fringe-dots. Hindwing with a row of six or seven evenly red spots, all separate
from each other and from cell, last two usually merged togetiier.
JIab. Dutch and French Guiana.
In the Tring Museum 3 (J(J, 2 ? ¥, from Surinam.
In coll. Oberthiir a long series of both sexes from Maroni and Cayenne.
/. P. anchises thelios Gray (1852).
?. P<ipn;o thelios Gray, Cat. Lpp. Ins. Brit. 3Tuk. i. Pap. p. 52. n. 250. t. 10*. fig. 7 (18.52) (Par.i);
Wall., Trans. Euf. Soc. Loml. (2). ii. p. 256 (1854)( Parii; forest) ; Gray, List Lep. Ins. Brit.
Mas. i. Pap. p. 6.5. n. 265(1856).
(J ? . PapiVw hierochs Gray, Cat. Lep. Ins. Brit. Miis. i. Pap. p. 55. d. 258. t. 10. fig. 2. (J, t. 9. fig. 9.
? (1852) (Pard) ; Wall., Traiw. Enl. Soc. Loml. (2). ii. p. 255 (1854) (Amazons ; forest) ; Gray,
List Lep. Ins. Brit. Mus. i. Pap. p. 67. n. 273 (1856) (Para) ; Bates, Trans. Enl. Son Land. (2).
V. p. 341, 356 (1861) (synonymy ; " cyphotes " excl. ; Para) ; id., Journ. Enlom. I. p. 225.
n. 13. (1862) ; Felder, Verli. Zool. But. Ges. Wien xiv. p. 203. n. 48 (1864) (Parit) ; Oberth.,
Et. (TEnt. iv. p. 88. n. 277 (1880) (Pard) ; Wood, Ins. Abroad p. 551. fig. 301 (1883).
?. PapiUo aglaope Gray, Cat. Lep. Ins. Brit. .Vits. i. Pap. p. 56. n. 260. t. 10. fig. 6 (1852) ( ? only,
(J alia spec. ; Pari); id., List Lep. Ins. Brit. Mus. i. Pap. p. 67. d. 275 (1856) ( ? ; ParA).
Papilio cyphotes, Kirby (nnn Gray, 1852, err. det.). Cat. Diurn. Lep. p. 527. n. 62 a (1871) {partim) ;
Maass. & Weym., in Stiibel, Reisen S. Amer., Lep. p. 89. n. 42 (1890) (Baiilo, Lower Amazons).
c?. Upperside. — Forewing with triangular green patch and one or two white
spots, seldom a minute third spot being present. Hindwing nearly as strongly
dentate as in P. a. anchises, with three or four separate red spots and often a
narrow line behind M-, s])ot M' — M- the longest.
On underside of hindwing a row of six se])arate ])ale red spots, of which the
central ones stand nearer the cell than outer margin.
?. Upperside. — Forewing with or without white cell-streak; two larger
distal spots R^— M-, often i)receded by a small s])ot, spot M' — M= the largest,
oblong. Hindwing : a row of seven or eight separate red sj)ots, the last two
merged together.
On underside the spots of the hindwing somewhat paler than above.
Hah. Lower Amazons : Para to Santarem.
In the Tring Museum 2 iS,2, ? ? , from Panl.
(J. P. anchises ctias subsp. nov. (PI. VII. tig. 40. 47).
<S . Like ]'. a. orbigmjanus, but spot SC^ — R' of hindwing absent, being rarely
represented by a minute dot, the others on the whole somewhat shorter and
narrower. Palpns sometimes almost entirely black.
? . Spots of forewing pure white ; cell-spot minute ; a small spot before R' an4
( 487 )
two large spots R^ — M^, not touching M-. Spots of bindwing smaller, being
shorter and narrower, than in orbignijanus; spot SO- — li' sometimes missing.
Hah. Eastern Bolivia.
In the Tring Mnsenm 11 c?(?, 2 ? ?, from : Santa Crnz de la Sierra, Jannary,
April — May 1!.H)4 (J. Steinbach) ; E. Grande, Province Cordillera, December 1903
(J. Steinbach).
In coll. Gudraan from Rio San Mateo.
h. P. anchises orhignyanus Lucas (1852).
(J. Painlio nrhirjnijiinns Lucas, Reo. Zooh ("2). iv. p. 192. t. 10. fig. 3 (I85'2) (Corrientes) ; Doubl.,
Westw. & Hew., Gen. Diiirii. Lep. ii. p. 530 (1852) ; Gray, Lht Lep. Ins. Brit. Mun. i. rup.
p. 64. n. 25G (1856); Bates, Trans. Ent. Sue. Loml. (2). v. p. 311, 357 (1861) ; Felder, Vcrh.
Znol. Bot. Ges. Wien xiv. p. 294. n. 64 (1864).
P(ipilii) cyiiuicldcs var. a. P. orhiijmjaniis, Kirby, Cat. Diurii. Lep. p. 526. sub n. 61 e. (1871)
(Corrientes).
(^. Papilio serapis, Burmeister {non Boisd., 1836, err. det.), Descr. Rep. Argent, v. Lip. p. 64.
n. 6 (1878) {parlhn).
(S. Green patch of forewing rarely much widening posteriorly, its pro.ximal
and distal edges being ]iarallel in most individuals ; always a buffish white spot
M' — M-, often very large, usually also a second spot before M' and often a third
behind M-. Hindwing with an evenly curved band which gradually widens
posteriorly, consisting of five spots and a line behind M'-, the spots close together,
but separate, the veins being black, rarely a dot in cell (type-specimen).
On u)>dcrsi(le of hindwing a row of seven spots, the last two being usually
merged together.
? . Forewing : a sjiot of variable size in cell, rarely absent, reaching sometimes
halfway across cell ; three spots R- — M- on disc, the first minute, often absent, the
second the largest, the third mostly rounded off behind and then not touching vein Ml
Hindwing : band from SO- to abdominal edge, separate from cell.
Larva reddish-brown ; a spot at sides of the thoracic segments and of the last
two abdominal rings, a dorsal spot on each side of mesial line on second and tenth
rings, and an oblique, sometimes macular, side-band on si.xth and seventh ;
tubercles of prothoras not longer than those on the following segments. Dorsal
tubercle of thorax of pupa small, divided ; abdomen with two pyramid-shaped dorsal
tubercles and a smaller one in front of them.
Hab. Paraguay; adjacent district of Argentina; Blatto Grosso ; Province
Goyaz in Brazil.
In the Tring Museum IGJcJ, 8 ??,6 larvae, 1 pupa, from: Formosa,
Argentina; Patino cue, Paraguay, February 1894 (Montforts) ; Villa Maria to
Diamantino, January 1897 (Andeer).
i, P. anchises foetterlei subsp. nov. (PI. VII. fig. 44. 3", 45. ?).
i. V i>periiiik. — Forewing : two very large white patches R' — M-, often
followed by a smaller spot and usually accompanied by a minute streak in cell ;
bluish grey scaling between the white patches and inner margin, partly edging the
patches, especially on distal side. Hindwing : red band rather paler than in
orhignyanus, broader, entering cell (always ?), streak behind M- very distinct, inner
edge of band almost straight from SC^ to base of M".
Underside paler than npper ; white spots of forewing somewhat larger,
( 488 )
especially tbe oell-sjmt. Banil of liiiulwing almost elongate-rhombiform, extending
from SO^ to near anal angle ; a dot in cell.
?. I'pperside : s])ots of forewing wLite, cell-spot large, but not reaching across
cell ; two large patches W — M-, a minute dot before R' and sometimes a small
streak behind M-. Hiudwing : red band almost as in male, anteriorly wider than
in that sex, crossing apex of cell, the veins only partly black, the central ones
almost entirely red within band ; a small spot before 80- present or absent.
Underside: band paler than above, the veins for the greater part black.
Ilah. Brazil : Sao Paulo.
We have much pleasure in naming this form after Herr J. Foetterle, from
whom we have received some useful material of Papilio.
In the Tring Museum 3 c? c?, 3 ? ¥ from Sao Paulo (received from Messrs.
Rtaudinger & Bang-Haas).
36. Papilio hedae Foett. (1902).
? . Papilw hakie Foetterle, Rev. J/(is. Paulista v. p. 620. t. 15. fig. 1 (1902) (S. Paulo).
The figure gives one the impression of the unique specimen being an abnormal
individual of P. anckises foetteiiei. However, it is impossible to be certain on this
point without comparing the specimen with a series of females oi foetterlei. If it
is, as we suppose, an aberration, the iudividiial will doubtless remain unique for a
long time.
The forewing is bufiieh white from inner margin forward to near 11°, the inner
edge of this broad band entering cell just proximally of M^ a large semicircnlar
cell-spot forming part of the band ; on the underside the band wider and reaching
close to costal margin, its distal edge irregular. A pale red band ou hiudwing
gradually shading utf into grey, the grey scaling nearly extending to base on
underside.
Hub. Interior of Sao Paulo.
37. Papilio nephalion Godt. (1819).
$. PupUiij tirphal'wii Godart, Enc. Melh. ix. p. 37. n. 3G (1819) (Brazil) ; Lucas, Lej'- T^-tnt. p. 29.
t. 14. fig. 3 (183.5) (Brazil); Boisd., Spec. Gihi. Up. i. p. 294. n. 124 (18:i(;) (Brazil) ; Lucas,
in Guiif., Diet. Pitt. Hixt. Nat. vii. p. 47 (1838) ; Doubl., Westw. & Hew., den. Diiini. Lep. i.
p. 18. n. 199 (184IJ) (Brazil) ; id., List Lep. Ins. Brit. Mus. i. App. p. 147 (1848) ; Mune'tr., Enum.
Corp, Aidni. Mus. Petrop., Lep. i. p. 5. n. 77 (1857) (Brazil).
(J. Papiliu vertunmiis, Godart (mm Cramer, 1782, err. det.). I.e. p. 37. n. 38 (1819) (parlim ;
Brazil) ; Luca.<i, Lep. Eu-vt. p. 13. t. 7. fig. 2 (1835) (Brazil) ; M^n($tr., I.e. n. 78 (1857) (Brazil).
^. Pruiiiiittes osyinintdims Hiibner, Samml. Exot. Schtnett. iii. t. 27. (1834':") (Brazil).
(J. Papilio prnteus Boisduval, Spec. (lin. Lip. i. p. 297. n. 128 (1830) (Brazil) ; Doubl., List Lep.
Ins. Brit. Mus. i. p. 12 (1845) (Brazil).
5 , Papiliu tiillus, Doubleday («oti Cramer, 1780, err. det.), List Lep. his. Brit. JJus. i. p. 12 (1845)
(Brazil) ; id., Westw. & Hew., Gen. Dium. Lep. i. p. 18. n. 201 (184G) {i>artim ; Brazil ;
cj = pruteifs).
cJ. Papiliu stilhiin Kollar, Ann. Wien. .Musiuni ii. p. 215. t. 12. fig. 1 (1839) (Brazil) ; Bates, Trans.
Enl. Sue. Land. (2). v. p. 357 (1861).
J ? . Papilla pruteiis, Giay, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 46. n. 233 (1852) (Brazil ; stilbun is
var.) ; id., List Lep. Ins. Brit. .]fiis. i. Pap. p. 62. n. 246 (1856) (Brazil ; Rio Grande do Sul) ;
Kates, /.r. p. 341,356(1861) (Rio); Prillw., Stett. Eitt. Zeit. x.xvi. p. 130 (1865) (Corcovado) ;
Capronij., Ann. Sue. Ent, Belt), xvii. p. 8. n. 4 (1874) (Gavia, August).
?. Papiliu protens var. b. Papilio nephalion^ Gray, Cat. Lep. Ins. Brit. Mus. \. I'liji. p. 47. sub n.233.
t. 10*. fig. 8 (1852) (Brazil).
( 489 )
c? 9. Pajii!ionep1wlioii,¥e\ieT,1'erh. Zool. Bot. Ges. ir/mxiv.p. 293.D. 51 (18G4) (Southern Brazil);
Kirby, CkI. Diiirii. Lep. p. 527. n. 62 (1871) (Brazil) ; Weym., ^Mt. Ent. Zdt. Iv. p. 315. n. it
(IK'Jy) (Rio Grande do Sul) ; Mabilde, Triim Pravt Bui-bol. liio Grande do Sid p. 44 (1890) ;
Bonningh., Vei-!>. Ver. Nut. Unterh. ix. p. 27 (189G) (Rio de Janeiro ; rather common).
cJ. Piipiliu ncplmlion ab. 1 ^ P. haemon Felder, Verh. Zool. But. Ges. Wien xiv. p. 293. sub n. 51.
p. ;«(■,( 1864) (Mus. "Vienna).
cJ 5 . Pdp'din veftumniis, Burmeister, Desci: Ri'p. Argent, v. Lej)., Atlas p. 7. n. IG (1879) (larva,
pupa; Rio de Janeiro to Santa Catharina ; synon. j'cii'tim') ; Seitz, Stett. But. Zcit. liv. p. 18
(1893) (Santos).
cJ. Pop'diiiosniandnjas (.'), Burmeister, I.e. (1879) (sub syn.).
?. Papilio nephuhon (.'), Bonninghausen, I.e.
Endoporjnn nephal'ion, Kirby, in Hiibn., Samtnl. Exot. Schnetl. ed. ii. p. 88. t. 4G5. fig. 3. 4 (190— V).
In structure practically the same as P. anchises, but very different in colour
from the Brazilian form of that insect.
S . The creamy patch of the forewing large, consisting of two or three spots,
spot M' — M^ being the largest, the patch extending occasionally to near SM^,
sometimes only spot M^ — M- distinct ; some specimens with a minute dot in cell.
Hiudwing usually with three spots R- — M-, rarely with two only, often an
additional s^Mt SC- — R' marked, but apparently never a spot R' — R- ; tooth R^
prominent.
On underside the three spots R- — M- of hindwing pinkish white, their distal
portions remaining red ; two red spots between M- and anal angle, usually confluent,
and a spot SC- — R\ the latter often vestigial.
? . Forewing with or without cell-spot ; a patch of two or three discal spots,
spot M' — M- the largest. Hindwing : red band from R^ to abdominal margin,
consisting of five spots, the last two being usually confluent ; most specimens with
an additional spot SO- — R', while in some others there is also a dot R' — R- ; rarely
spot W — R- present and SC- — R' absent.
On underside there is apparently always a cell-spot on forewing. The
hindwing bears always a sjiot SC- — R', while spot R' — R- is rarely marked ; spots
R- — M- are rather large, sometimes spot R^ — M' touching cell, but there is never
a dot in cell ; colour of these spots as in c?.
Early stages described by Burmeister, I.e.
Hub. Brazil ; Matto Grosso ; Paraguay and adjacent districts of Argentina.
In the Tring Museum 40 <S i , 32 ¥ ?, from: Minas Geraiis (A. Kennedy);
Riu de Janeiro ; Organ Mts., Sarapuhy, and Corcovado ; Sao Paulo ; Castro, Parana
(E. D. Jones); Hajahy, S. Catharina, February 1897; Yhu, Paraguay, September
— December 1896 (Audeer) ; Sapucay, Paraguay, October, December, and January
(W. Foster).
III. Lysander Group.
Marginal spots of hindwing red. Palpus always black. Hindtibia of male
incrassate, hairy, foretibia not enlarged, spinose as in female. Harpe long, reaching
close to ajiex of clasper, truncate, with two to five apical spinelike teeth, ventral
edge non-dentate or minutely denticulate, never with a prominent conical tooth as
in most species of the Aeneas Group. Anal segment of female without the short
Btout spinelike bristles of the Belus Group ; in the vaginal cavity, on the proximal
side, two broadly triangular lobes close together, these lobes jjointing proximad
when the walls of the cavity are pushed outside.
( 490 )
Key to the species : —
A. Vein M- of hindwing branching off from cell far beyond iSC-
iisymmetrical.
a. Forewing without markings in both sexes, except the red
or pinkish white marijinal dots ; an evenly cnrved row
of red spots on hindwing about halfway between cell
and distal margin
Forewing with bine or bluish green band in male ; in
female with or without wliite patches, no distinct
fringe-spots, or the fringe not indented between the
veins ; red band of hindwing not regularly arched, the
spots closer together and less regularly ovate than in
Species No. 38
b. S without white wool in fold of hindwing ; ? , distal
margin of forewing straight, last two spots on hindwing
on a level with one another, usually confluent
(J with white wool in fold of hindwing ; ? , distal margin
of forewing rounded, last two spots of hindwing not on
a level .........
c. (f , band of forewing oblique, strongly tapering, patch
M^- — SM- obliquely truncate proximally ; ?, apical
half of forewing semi-transparent, distal margin feebly
rounded, tooth R^ of hindwing usually prominent.
6, band of forewing more straight, patch M- — SM^
square, one or two white spots on disc ; ? , apex of
forewing more opaque than in No. 40 ; distal margin
more convex, hindwing more rounded ....
B. Veins SC- and M- of hindwing almost at the same distance
from base, the cell being nearly symmetrical.
d. c?, midtibia densely covered with small hairs ; ?,
forewing with broad white subapical cell-patch, or
with two rounded spots on disc, in the latter case the
red band of hindwing always entering cell ,
S, midtibia spinose like foretibia, cell of hindwing red
on npj)erside (base excepted), or the forewing with
large white patches ; ? , no spot in cell, or only a
small one, band of hindwing outside cell
e. cJ, forewing with two white spots on disc; ¥, forewing
with a small or no spot ......
<?, forewing with one or no white spot on disc ; ? , fore-
wing with large cell-patch .
f. i, forewing with large white spots on disc, band blue, cell
of hindwing black; ?,i'on'wing o]iaque, or a straiglit
row di' three white spots on disc, the upi)er one being
more or less obscure, but no spot in cell
S, band of forewing green, cell of hindwing red on
npperside ; ? , apical half of forewing semitrausjiarent,
no spots on disc, or small ones, or the spots are large,
usually also a streak in cell ......
the cell bein"
Species No. 38.
b.
Species No. 41.
Species No. 40.
Species No. 39.
/•
Species No. 45.
.Species No. 44.
Species No. 43.
Species No. 42.
(491 )
38. Papilio panthonus Cram. (1780).
Papilio Eques Trnjanus jHuilhoims Cramer, I.e. iii. p. 154. t. 278. fig. C. D. jj (1780) (Surinam) ;
Esper, Aitsl. Schmell. p. 67. n. 30. t. IG. fig. 4 (1789).
Papilio Eques Trojaims pninqjeius Fabricius, Spec. Ins. ii. Apipcnd. p. 502 (1781) (nom. nov. loco
panihoiius^.
<? ? . Sexes similar, but the female ]>aler tliaa the male; fringe of forewiiig
spotted with pale red ; hiudwing with regularly curved row of red spots situated
about halfway between cell aud distal margin.
Scent-organ : fold with white wool as in P. lijsander.
Genitalia : <?, harpe truncate at apical edge, with about six teeth of nearly
ecpial size.
Early stages not known.
liab. The Guianas and Brazil.
Two subspecies.
a. P. panthonus ni.ima Boisd. (1876).
? . Papilio lumm Boisduva), Spec. Gin. Lip. i. p. 280. n. 116 (1836) ( ? ; hab. ?) ; Doubl., Westw.
& Hew., Gen. D'lm-n. Lep. i. p. 18. n. 208 (1846) ; Gray, Cat. Lep. Ins. Brit. .Mas. i. Pap. p. 64.
n. 285 (1852) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 74. n. 301 (1856) (po.rtiin) ; Bates,
Trans. Kid. Soc. Land. (2). v. p. 361 (1861) ; Feld., Verh. Zool. Bot. Ges. Wien xiv. p. 295. n. 85
(1864) (Surinam ; Demerara) ; Butler, Cat. Diurn. Lep. descr. Fabric, p. 237. sub n. 11 (1869)
\^ panthonus 'i); Kirby, Cat. Diurn. Lep. p. 529. n. 70 (1871) ; Oberth., Et. d'Ent. iv. p. 82.
n. 271 (1880) (diS'erences from panthonus).
(J. Papilio jai/uarae Toetteile, Rev. Mas. Paulista v. p. 619. t. 15. fig. 8 (1902) (Minas Geraes ;
Sao Paulo).
c? ? . Differs from the Guiana form in the forewing being rather paler and in
the red spots of the hindwing being smaller, and therefore farther apart.
Hab. Interior of Sao Paulo and Minas Geraes, Brazil.
Type (?) of numa in coll. Oberthiir.
b. P. panthonus pa?ithoni(S Cram. (1780).
Papilio Eques Trnjanus panthonus Cramer, I.e. (Surinam) ; Esper, I.e.
Papilio Eques Trojanus pompeius Fabricius, I.e. ; id.. Maid. Ins. ii. p. 5. n. 37 (1787).
Papilio Eques Trojanus pompejus, Jablousky & Herbst, Naturs. Sehmett. ii. p. 48. n. 19 (1784) d^) ;
Gmelin, Sjjst. Xa i. 5. p. 2233 n. 295 (1790) ; Fabr., Ent. Si/sf. iii. 1. p. 18. n. 64 (1793).
Priamidcs ponqiejiis, Htibner, Verz. hek. Schtnett. p. 87. n. 904 (1818 I).
Papilio pompeius, Godart, Ene. Meth. ix. p. 36. n. 32 (1819) (partim)
Papilio arbates, Boisduval (non Cramer, 1782, err. det.). Spec. Gen. Lep. i. p. 291). n. 118 (1836)
(jiartint).
Pajiilio jmnthouus Gray, Cat. Leq>. Ins. Brit. Mus.'i. Pap. p. 64. n. 284 (1802) (synon. ;«(Wi7«) ; id., Lint
Lep. Ins. Brit. Mus. i. Pap. p. 74. n. 302 (1856) ; Bates, Trans. Ent. Soc. Lond. (2). v. p. 358
(1861) ; Feld., Verh. Zool. Bot. Ges. Wien xiv. p. 295 n. 87 (1864) ; Kirby, Cat. Diurn. Lep.
p. 528. n. 69 (1871) (Guiana ; synon. partim); Oberth., Et. d'Ent. iv. p. 82. n. 272 (1880)
(Guyane).
Papilio aeneas, Butler {non Linnu, 1758, err. det.). Cat. Diurn. Lep. descr. Fabric, p. 236. n. 11 (1869)
{partim ; $ ).
Papilio santhonus (!), M.'pschler, Verh. Zool. Bot. Ges. Wien xxvii. p. 295(1876) (Surinam).
Papilio plnjlarehus Hopll'er, Stett. Ent. Zeit. xxvii. p. 24. n. 3 (1866) (Cayenne).
i ? . The red spots of the hindwing variable in size, but apparently never so
small as in the Brazilian form. The fringe-spots of the furewiug are sometimes
nearly pure white ; iu other sijecimens they are represented only by a very few
rosy scales.
Hab. The Guianas.
32
( 492 )
in the Triug Museum 7 tJrf, 5 ? ?, from : Baitica, Britisli Guiana, February
1904 ; Paramaribo, February 1892 (Ellacombe) ; New Amsterdam.
39. Papilio aglaope Gray (1852).
(2) Papilio Eqiies Trojanus eurisleus Cramer, Phjj. ExuI. i. p. 47. t. 29. fig. F (177;'j) (f^urinam ;
this species ?).
(?) Papilio eiiristeiis, Boisduval, Spec. Geii. Lip. i. p. 282. n. 107 (183G) ; Kirby, Cat. Diurn. Lcp.
p. 528. n. CS (1871).
cJ. Papilio aglaope Gray, Cat. Lep. Lis. Brit. .Uiis. i. Pap. p. 55. n. 2C0. t. 10. fig. 5. ^ (1852) (Pani ;
partim, J alia species) ; Wall., Trans. Eiit. Sac. Loud. (2). ii. p. 255 (1854) (Amazona, forest) ;
Gray, List Lcp. Lis. Brit. .Vus. i. Pap. 07. n. 275 (1850) {jiartim) ; Bates, Trans. Ent. Soc. Loud.
(2). V. p. 343, 358 (1801) (Pari ; only two ^ ^ known) ; id., Journ. Enlom. i. p. 220. n. 19
(1802) (Para, rare) ; Feld., Verh. Zool. Bot. Ges. Wieii xiv. p. 2'.I5. n. 83 (1804) (Pari) ; Kirby,
Cat. Diurn. Lep. p. 528. n. 07 (1871).
? . Papilio crlaces Gray, Cat. Lep. Lis. Brit. Mas. i. Pap. p. 49. n. 2(0. t. 8. fig. 10 (1852) (Bolivia ;
cj alia sjiecies') ; id., .s Lrj>. Lis. Brit. J/«s. i. Paj). p. 03. n. 253 (1850) (partim).
¥ . Papilio callicles Bates, Tram. Ent. Soc. Loud. (2). v. p. 301 (1801) (nom. nov. pro erlaces J ) ;
Feld., I.e. p. 295. n. 80 (1804) ; Kirby, Z.c.p. 629. n. 70 (1871) ; Hopff., Stctt. Ent. Zeit. xl.p. 50.
n. e (1879) (Bolivia).
cJ ?. Papilio hjsimachus Honratb, Bcrl. Ent. Zeitschr. xxxii. p. 251. t. 5. fig. 5. (J, fig. 0. ? (1888)
(Soathera Peru).
We believe Cramer's figure of euristeus to represent a specimen of tliis species.
But as the figure is very rough, and as, further, the species is not known from
Surinam, we are not certain that our identification is correct. For this reason we
treat the name euristeus as of doubtful application, and accept Gray's name aglaope
for the present insect.
cJ. Forewing shorter, its distal margin more convex than in P. lysandcr;
bluish green band more straight, patch M^ — SM- square ; a large white spot
M' — M^ in band, generally a second spot R^ — M' ; fringe usually with small red
spots. Hindwing : a row of four red spots, which are much shorter than in
P. hjsander, being separate from cell ; sometimes a minute fifth spot before R'.
? . Resembling closely the white-spotted female of li/sander. Apical area of
forewing less transparent, apex broader, distal margin more convex ; hindwing
more evenly rounded.
There are two forms, one which resembles the male in the position of the spots
of the forewing, and the other resembling P. lysander ? -f. parsodes.
a'. ?-f. lysimachus Hour., I.e. — Forewing with three spots in a straight row,
spot M^ — SM- greenish, small, spot M' — M- white, ronuiled, a little longer than
broad, spot R' — M' white, shaded with brown, narrow, situated along M'.
b^. ¥-f. callicles Bates, I.e. — Forewing with large white patch M' — M-, a
smaller patch W — M' and a streak in cell.
Scent-organ as in P. li/sander.
Hub. of P. aglaope : Lower Amazons ; Southern Pern ; East Bolivia.
In the Tring Museum 3 c?c?, 2 ? ¥, from : Igarape (W. Hoffmanns) ; Panl
(Stuart); Province Sara, Santa Cruz de la Sierra, Bolivia, February—Aiiril 1904
(J. Steiubach).
411. Papilio lysander Cram. (177.5).
Seba, Tliesaur. iv. p. 12. t. 7. fig. 27. 28. ? (1704).
(J. Pajjilio Equcs Trocs lysander Cramer, Pap. Exot. i. p. 46. t. 29. fig. C. D. (1775) (Surinam) ;
Goeze, Ent. Beytr. iii. I. p. 30. note (1779) (var. of aeneas) ; Esper, .li(.s/. Schmett. p. 62. n. 27.
t. 16. fig. 1 (1788) (fringe-spots of hindwing white in figure !).
(403 )
$. Popil/o Eqties Trojanus ancJiises, Fabricius (mvt Linue, 1758, err. det.), Si/st. Ent. p. 440. n. 19
(1775) (pavtim); Sulzer, Grgrli. SchniPlt. i. p. 141. ii. p. 24. n. 4. t. 12. fig. 4 (1776) (forewing with
white patches) ; Goeze, I.e. p. 34. n. 11 (1779) (parlini) ; Fabr., Spec. Ins. ii. p. 7. n. 2G (1781)
(partiiii) ; Gronov., Zonphtjktc. p. 188. n. 727 (1781) (partim) ; Esper, I.e. p. 13. n. 11. t. G.
fig. 1 (1785) ; Fabr., Mniit. /«,?. ii. p. 4 n. 28 (1787) (partim) ; Roem., Gni. Inn. p. G8. t. 12.
fig. 4 (1789) ; Gmelin, %.</;. Nat. i. 5. p. 2230. n. 11 (1790) (partim) ; Fabr., Enl. S.i/xt. iii. 1.
p. 13. D. 40 (17S3) (partim).
?. Papi/iu Eques Trnjanns arhates Stoll, in Cram., Pap. Exnt. iv. p. 198. t. 38G. fig. C. D (1782)
(.Surinam) ; Jabl. & Herbst., Naturn. Sehmetl. ii. p. 41, n. 17. t. 9. fig. 4 (1784) ; Esper, And.
Schmett. p. 55. n. 23. t. 14. fig. 1 (1788).
(^. Papilio Equps Tnijaiar.^ meleander Jablonsky & Herbst, Nature. ScJnnett ii. p. 75. n. 23. t. 10.
fig. 2 (1784) (nom. nov. loco hjsandcr Cram.).
(^. Papilio Eqiies Trnjanna acncas, Esper (iimi Linn^, 1758, err. det.), I.e. p. 40. n. 15. t. 9. fi^'. 1
(1786) (syn. excl.) ; Gmelin, Sijxt. Nat. i. 5. p. 22,33. n. IG (1790) (partim).
P. anbates (!), Goelt. Gelehrte ^«-.'40. Stiiok p. 400 (1790).
J . Prineepis dominans arhates, Hubner, Samjnl. Erat. Schmett. i. t. 123. fig. 3. 4 (180G — ).
5 . Priucepa dominans pnmprjus, id. (umi Fabricius, 1770, err. det.), I.e. i. t. 124. fig. 3. 4 (1806 — )
(fringe-spots of hindwing white in fig. 4, error in colouring).
(J. PrineejJS dominans lijsandcr, id., /.(•. t. 127. fig. 1. 2 (1806 — ).
?. Priamides anehises, id. (non Linne, 1758, err. det.). Vers. hek.Sehmetl. p. 87. n. 902 (1818?)
(partim).
? . Priamides hrissonius id.. I.e. p. 87. n. 903 (1818 ?) (nom. nov. Xoco pompejns Iliibn.).
j. Parides lysamler, id., he. p. 87. 910 (1818 ()•
(J. Papilio euri/mas Godart, Eiie. Melh. ix. p. 34. n. 27 (1819) (nom, nov. loco hjsandir Cram. ;
Guyane) ; Boisd., Spec. Gen. Lip. i. p. 284. n. 110 (1836) (" var." discoloured specim. ;
Cayenne; Surinam); Kollar, Denkschr. K. Ak. Wiss. Wien, Math. Nat. CI. i. p. 352. n. 3
(1850) (Las Palmas, Nova Granada).
Papilio anehiscs, Godart, Ene. ilHk. ix. p. 36. n. 31 (1819) (p irtim) ; Guea,, Ann. Soc. Ent. France
p. 309 (1867) (Sulzer's fig. of anchises represents dimas = ? of zaeynthnx, error) ; id.. I.e. p. 309
(1867) (Esper's fig. of am;hiscs represent? arhates, = ? panthoniis) ; Butl., Cat. Diiirn. Lep.
descr.Eahr. p. 235. n. 7 (1869) (partim) ; Kirby, Cat. Diurn. Lep. p. 529. n. 74 (1871)(^)fn'(ini);
Moschl., Verh. Zool. Bot. Ges. Wien xxvii. p. 295. (1876) (partim. ; Surinam ; = lysander =
arhates = eurymas); Stand., E.rot. Tnijf. i. p. 14. t. 9. (J ? (1884) ; Hahnel, /)■/»■ iii. p. 275
(1890) (Sao Paulo de 01iven9a).
Papilio lycander (!), Swainson, Zoo/. Illustr. iii. text of t. 92 (1823).
?. Papilio arhates, Boisduval. .S/«e. Gen. Lep. i. p. 290. n. 118 (183G)(;jai;/»i ; his (J is ? ); Kollar,
Denksehr. K. Ak. Wiss. Wien, Math. Nat. CI. i. p. 353. n. 5 (1850) (Venezuela).
Papilio panthonus, Doubleday (non Cramer, 1780, err. det.). List Lep. Lis. Brit. Mas. i. p. 12 (1845)
(partim) ; id.. Westw. & Hew., Gen. Diurn. Lep. i. p. 19. n. 225 (1846) (partim).
cJ ? . Pujnlio lysamler, Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 53. n. 254 (1852) (Demerara) ; id..
List Lep. Ins. Brit. Mus. i. Pap. p. GG. n. 2G9 (185G): Me'uetr., Enum. Corp. Anim. Mils. Petrop.,
Lip. i. p. 5. n. 79. 83 (1857) ("Brazil") ; Bates, Trans. Ent. Soc. Lond. (2.). v. p. 343 (1861)
(variabihty) ; id., Joum. Entom. i. p. 22G. n. 20 (1862) (typ. form in Guiana and Upper
Amazons as far down as Villa Nova ; local form parsodes at Par^) ; id.. Natural. Rir. Amaz.
p. 26 (1864) (Parit, J in swampy shades, J in more open places); id., I c. p. 151! (1864)
(lysamler replaces parsodes on the Upper Amazons) ; Fold., Verh. Zool. Bol. Ges. Wien xiv.
p. 295 n. 93 (1864) (Surinam ; Guiana; Cayenne ; Amazonia inf.; "Brasilia?") ; Oberth.,
El. d'Ent. iv. p. 91. n. 285 (1880) (Guyane).
(J ? . Papilio hrissonius, Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 53. n. 255. t. 8. fig. 7. ^ (1852)
(Ega); Wall., Trans. Ent. Soc. Lond. (2). ii. p. 256 (1854) (Upper Amazons ; forest); Gray, List
Lep. Ins. Brit. Mus. i. Pap. p. GC). d, 270 (1856) (Ega ; Villa Nova) ; Feld,, Verh. Zool. Bot.
Ges. Wien xiv. p. 295. n. 92 (1864) (Ega ; Orinoco).
(J ? . PapUio parsodes Gray, Cat. Lep. Ins. Brit. Mus. i. Pup. p. 54. n. 256. t. 8. fig. 3. J (1852) (Pard ;
partim ; $ alia spec.) ; Wall., Trans. Ent. Soc. Lond. (2). ii. p. 255 (1854) (Par;! ; forest) ;
(iray, List Lep. Ins. Brit. .Mas. i. Paji. p. 66. n. 271 (1856) (Para ; Villa Nova) ; Bates, Natural.
Pie. Amaz. p. 156 (18G4) (replaces lysamler in the Delta region) ; Feld,, Verh. Zool. Bot. Ges.
Wien xiv. p. 295. n. 91 (1864) (Par;! ; Tocantins ; " Mexico (") ; Oberth., El. d'Ent. iv. p. 92.
u. 286 (1880) (Para ; geogr. form of lysamler) ; Sharpe, Proe. Zool. Soc. Lond. p. 555. n. 3(1890)
(Prov. Goyaz).
? . Papilio sonoria Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 57. n. 263 (1852) (nom. nov. loc. "anchises
Sulz." ; Pard) ; Wall., Trans. Ent. Soe. Lond. (2). ii. p. 256 (1854) (Pard ; forest) ; Gray, List
( 494 )
Lep. TiiK. brit. M,i>:. i. Ptip. p. G8. n. 278 (I80O) (Para) ; Butl., Trims. Eiit. Soc. Loml. p. 145.
n. 224 (1877) (^, R. Tnpajos, March).
? . Pajiilio sonoria var. a., Gray, Cot. Lfji. Ins. lirit. Mus. i. Puj). p. 57. sub n. 2ti:i (1852) (Pari).
?. Pupilio aimxiiiiander Felder, Vcrli. Zool. B'll. Ges. Wien .\iv. p. 2;i5. n. 89 (ISO!) (h'ihi. nnd.,
hab. ?) ; id., Ri-isc Novara, Lep. p. 'M. n. 21. t. 18. fi^'. b (18l!5) (hab. V— Mus. Tring) ; Kirby,
Cat. Diuni. Lep. p. 52il. n. 73 (1871).
(J. J'li/iilin jiliri/nichus Felder, V'oh. Ziml. Bui. Gen. Wim xiv. p. 295. n. 90 (1804) {iioni. mid. ; Nova
Granada) ; id., Rcisc Xoram, Lep. p. 33. n. 22. t. 8. fig. e (1805) (Las Palmas, N. Granada ; —
Mus. Vienna).
(J. Papilin bjnander var. hari Oberthiir, Et. d'Eiit. iv. p. 91. sub n. 285 (1880) (patch of bindwing
orange).
Piipiliii uiichises y^T. parsodes, Moschler, Virh. Zool. Boi. Gtn. Wien xxxii. p. 304. (1883) (Surinam).
? . Paridcs arbates, Kirby, in Hiibn., Saniiul. Ecut. Schmetl. ed. ii. p. 90. t. 12.1. fig. 3. 4 (190— ?).
cj ? . Parides brissonius, id., I.e. p. 90. t. 124. fig. 3. 4, t. 127. fig. 1. 2 (190— ?).
The considerable series of specimens we have compared appears to prove that
the sjiecies is not split np into vf ell-defined geograjihical races, in spite of the large
area it occupies. The males at least do not show any reliable differences iu the
various districts. However, there is something geographical in the variability of
the other sex, in one j>lace the one female form being prevalent or occurring alone,
■while in another district another form is more commonly met with. According
to the females the area inhabited by P. li/sander can be divided into three
districts :
(a) Eastern jjortions of Colombia, Ecuador and Peru, and the Uiiper and Middle
Amazons. In this district the females have no white spots on the forewing or only
traces of them.
{h) The Lower Amazons, south side of the river. Here the females have always
a large white patch consisting of several spots.
(c) The Guianas (and ])robably the north side of the Lower Amazons). The
females agree either with (</) or with (/;), or are intermediate. In Surinam the
larger proportion of the females are without white patch or have only a round spot
M' — M-, while in British Gniana the majority of females have as large a white
patch as the Para specimens.
Gray, I.e., and also Bates, I.e., said that the Para males have a larger green
band on the forewing than those from other i)lace8. This distinction does not
hold good.
c? ? . Intermediate between P. echemoti and aylaope, the main differences being
stated under these species. Linn(5's Papilio anehises is quite a different insect. The
earlier writers treated all the males of the species of this group marked green and
red as being specifically the same.
P. h/sander is the first name given to the present species. Gray correctly
assigned the name of hjsander to Guiana specimens of this insect. But Kirby
in his Catalogue enumerated the species again as anehise.i L. In a S m coll.
Oberthiir the patch of the hindwing is orange instead of red (ab. bari).
In order to facilitate reference the females may be grouped in three individual
forms :
a. H-L jmrsodes Gray, I.e. ; .\oiioria id., /.c— Forewing with large white jiatch
consisting of several spots ; besides a large spot M' — M" there being a spot Ii'' — 11',
another behind M-, and often a small spot in cell.
b'. ?-f. arbates Stoll, I.e.; anaximenes Feld., I.e. — Forewing with a single,
more or less njunded, spot M' — M'-'. In type of anaximenes the red spots of the
hindwing are faded, except the upper two.
( 493 )
c'. ?-f. hrissoniiis Hiibner, I.e.; pompejus id., I.e. (iwn Fabricins, 1T87, err.
det.). — Forewiug without distinct white spot.
Early stages not known.
Hab. of F. lijmnder : The Guianas ; Amazons from Para to Peru and Ecuador ;
" Bogota."
In the Tring Museum 38 cJc?, 4.5 ? ?, from : Bartica, Brit. Guiana, Febrnar}'
1902; Rio Demerara ; Fort Aka3'raa; New Amsterdam; Upper Real Berbico R. ;
Berg-en-Daal, May 1892 (Ellaoombe) ; Sariuain ; Teffe, January 1905 (Mathan) ;
Juhuty, April 1905 (Mathan); Maues ; Itaitnba ; Iquitos (Stuart); R. Jurua ;
Igaraji^ (W. Hoffmanns) ; Para (Stuart) ; Archidona, E. Ecuador (R. Haeusch).
41. Papilio echemon Hubn. (1806—).
Prhiceps dominans echemon Hiibner, Samnil. Exoi. Schmett. i. t. 121. fig. 3. 4. J (180G — ).
Princepes (!) dominans echelus id., I.e. t. 126. fig. 1. 2. ^J (1806 — ).
The male of this species is easily distinguished from that of P. I'/sander.
The females of the two insects come often very near one another, but can be separated
by the differences hereafter mentioned.
cJ. Apex of eighth abdominal segment less extended red than in P. hjsander.
Forewing narrower, apex more acute, distal margin concave, the blue-green spot
situated behind SM- small, not produced basad into a point ; underside with
cyaneous gloss posteriorly ; scales of upper layer in apical area of upperside
bidentate, those of under layer trideutate, the teeth being short ; on underside the
upper scales bi-, the under scales tridentate in apical half of wing, the teeth,
though long, being shorter than in P. lijmnder. Hindwing with cyaneous gloss
above, especially along abdominal fold ; scales of upperside in distal area nearly
all denticulate, the larger portion of the red patch included ; tooth R^ longer
than in P, hjsander.
? . Apex of seventh and eighth abdominal segments less extended red than
in P. hjsnnder female, the red scaling usually restricted to a postvagiual spot.
Forewing narrower than in P. hjsander ?, distal margin less convex, being
straight from SC' to SM-. Hindwing : tooth R^ prominent, anal angle produced,
last two red spots M- — SM- on a level with one another, usually not separate,
forming a transverse bar either above or below or on both sides, while in
P. li/saiidfr the two spots are separate, the posterior one being more distal than
spot M- — (SM^), the oblique position towards each other being especially obvious
on underside.
Nenration : Apex of cell of forewing narrower than in P. bjmnder, cross-veins
D' and D- less oblique ; D^ of hindwing usually much shorter than D*.
Scent-organ : edge of fold with a fringe of long hairs ; no wool in the fold,
the surface of the fold being occupied by densely packed, lanceolate scales, which
have a cyaneous gloss.
Genitalia : cj. Harpe as in P. li/sander with several teeth at apex, the tooth
situated at the ventral angle being the longest.
Early stages not known.
Hab. The Guianas and Lower and Middle Amazons.
Two subspecies ;
( 490)
a. P. echemon echemon lliibn. (180G — ).
$ . Princepi dominans echemon Hiibner, I.e.
(J. Prhiceps domirutHs echelus id., I.e.
5. Priatnides echemun id., Verz. belc. Schmett. p. 87. n. 898 (1818?).
(j. Pcirides echelus id., !..■. n. 907 (1818 ?).
(J. Papilio echebi.1, Boisduval, Spec. Gen. Lfp. i. p. :287. n. 113 (183r,) (" Amer. mtr.", descr. from
Hilbner's fig.).
?. Ptijillin pohjmetus, id. {iion Godart, 1819, eiT. dft.), I.e. p. 28:!. sub n. 108 (183(1) (j>arllm ■ =
erhetnoii Hiibn.).
?. Paiiilio fpartacun Doubleday, Westw. & Hew., Gen. Diiiru. Lrp. i. p. 18. n. 200 (1846) (num.
mid. ; Brazil ; "cit. Doubl." erroneous).
J . Papillo echelus var. a. Papilio spariacus Gray, Cut. Lep. Ins. Brit. .1/».<. i. Pap. p. 55. sub n. 257.
t. x*. fig. 1 (1852) (Brazil); id., List Lep. Ins. Brit. Mus. i. Pap. p. (!7. sub n. 272 (185C): Kirby,
Cat. Diuru. Lep. p. 530. sub n. 75 (1871) (Brazil).
(J ¥ . Papilio eclielus Doubleday, Westw. & Hew., Gen. Diurn. Lep. i. p. 18. n. 120 (184fi) ; Gray, Cut.
Lep. Ins. Brit. Mus. i. Pap. p. 54. n. 257 (1852) (syn. i)artim ; Pant) ; Wall., Trans. Eiit. Sue.
Loiul. (2), ii. p. 255 (1854) (Amazons ; forest) ; id., List Lep. Ins. Brit. Mus. i. Pop. p. G7. n. 272
(1856) (syn. 2>artim ; Pani ; Santarem ; Brazil); Mi'netr., Emim. Corp. Anim. Mus. Pelrop.,
Lep. i. Suppl. p. G8. n. 1130 (1857) (Parii) ; Bates, Trans. KnI. Sne. L„u,l. (2). v. p. 344 (IHOl)
(Lower Amazons) ; id., J.mrn. Enl. i. p. 227. n. 21 (1802) (Par.-l and south shore of Amazon
as f.ir as Santarem, nowhere else) ; Reak., Proc. Ent. Soc. Philad. ii. p. 138. n. 7 (1803)
("Nicaragua" false); Feld., Verh. Zonl. Bot. Ges. Wien xiv. p. 290. n. 94 (1804) (I'ani;
" Nicaragua" error) ; Bates, Natural. Rir. Amaz. p. 100 (I804)(only on south side of Amazon).
S ? . Papilio echelus Kirby, Cat. Diurn. Lep. p. 530. n. 75(1W71)(" var. c." excl.) ; Oberth., /•;/. d'Ent.
iv. p. 93. n. 287 (1880) (Pari) ; Staud., Erot. Ta<jf. i. p. 14 (1884) (Lower Amazons) ; Maass.
& Weym., in Stiibel, Rcisen S. Amir., Lep. p. 89. n. 43 (1890) (Baiao, Lower Amazons) ; Uaase,
Untersuch. Mimicry i. p. 79 (1893).
Papilio anchises ? var. parsodes, id., I.e. ii. p. GO. t. 10 fig. 72. ? (1893).
(J ? . Parides echemon, Kirby, in Hiibn., Samml. Exot. Schmrlt. ed. ii. p 91 t I'^l fi" 3 4 t 120
fig. 1.2(190— ?). " ' '
c?. Bluish green baud of forewiiig narrow, tapering, extending from hind-
margin to W, separated from cell.
?. Forewing with band of white patches, separated from cell, tapering in
front, reaching from SM- forward to R^, sometimes being vestigial as far as R= or
even beyond ; patch M=— SM- as a rule square or almost, larger than patch
M'— iP; often a bluish grey streak behind SM-\ Posterior red (iloublc) patch of
hindwiiig, above, large.
Ilab. Lower Amazons, southern side as far upwards as Santarem.
In the Tring Museum 10 cJcJ, 5 ? ?, from: Igarape (W. HoflFmanns) ;
Sautarem.
b. F. echemon ergetelcs Gray (1852).
(J ?. Papilio ergetelcs Gray, Cat. Lep. Ins. Brit. Mus. i. Paj>. p. 52. n. 252. t. 8. fig. 5. J (1852)
(J" ? ; Amazons) ; id.. List Lep. Ins. Brit. Mus. i. Pup. p. GO. n. 207 (185G) (Amazons, (J);
Wall., Trans. Ent. Soc. Land. (2). ii. p. 255 (1854) (Upper Amazons ; forest) ; Bates, I.e. y.
p. 344, 359 (18G1) (descr. of ?, north shore of Amazon, westward apparently not beyond
R. Negro) ; id., Jouru. Entom. i. p. 227. n. 22 (18G2) (north side of Amazon from Obydos to
Rio Negro) ; Feld., Verh. Zool. Bol. Ges. Wien xiv. p. 290. n. 99 (1804) (Amazons) ; Bates,
Natural. Riv. Amaz. p. IGO (1804) (only on north side of Amazon from Obydos to Rio Negro) ;
Oberth., Et. d'Eut. iv. p. 91. n. 284 (1880) (Guyane) ; Wood, /,.*. Abroad p. 550. fig. 300
(1883).
cJ. Papilio echephrou Bates, Trans. Enl. Sr>c. Loud. (2). v. p. 345 note(18i;i) (French Guiana); Feld.,
Verh. Zool. Bot. Ges. Wieu xiv. p. 290. n. m (1804) (Cayenne).
0 . Papilio echion id.. I.e. xiv. p. 290. n. 95 (1804) (uom. nud. ; h.ib. V). ; id., Tteise Nuvara, Lep. p. 33.
n. 23. t. 8. fig. d (180,5) (hub. V— Mus. Vienna).
cJ. Paiiilio poli/phron Fclder, Verh. Zool. Bot. Ges. ^yien xiv. ]i 290. n. 9G(18G4) (worn. nud. ; Surinam);
id., Rcisc Not-ara, Lcj). p. 33. n. 24. t. 8. fig. c (18G5) (Surinam).
(497 )
? . Papilio pisauihr Felder, Verh. Zool. But. Ges. Wien xiv. p. 295. n. 88 (I8G4) (iwm. nud. ; hab. ?) ; id.,
Reise Novam, Lep. p. 31. n. 20. t. 8. fig. f (186,5) (hab. ?— Mus. Tring) ; Kirby, Git. Diiirn. Lep.
p. 529. n. 72 (1871) ; Miiscbl., Verh. Zuul. Bui. Ges. Wien xxxii. p. 304 (1883) (Surinam).
Papilio echelus var. a. P. erhioii, Kirby, Cat. Dium. Lep. p. 530, sub n. 75 (1871).
Papilio echelus var. b. P. polijphron, id.. I.e.
Pajiilio echelim var. e. P. echcphron, id., l.c
Papilio echelus var. f. ergeteles, id., I.e.
Papilio echelus, Moschler, l.c. xxxii. p. 304 (1883) (Surinam).
(?. Baud of fore wing wider tbau in the preceding, nsnally extending to cell at
M-, its inner edge being elbowed at this vein, spot R' — M' mostl}' absent; white dots
absent from underside in most specimens, being occasionally present in specimens
from the Amazons. Hindwing : red spots larger than in echemon, contiguous,
tiie two middle ones touching cell or nearl}', sometimes all four reaching cell, the
latter bearing in one such individual (from Anteirim, north shore of Lower Amazon)
a red dot at apex. Some Amazonian specimens intermediate between the present
subspecies and the preceding one.
? . Dichromatic, the white patches disappearing sometimes.
a'. ?-f. ergeteles Gray, I.e.; Bates, l.c. Forewiag : a large white patch
M^— M^, longer than broad, touching cell, preceded by a smaller patch R^ — W,
wliich is usually reduced to a streak or a shadowy spot standing before M', some-
times barely vestigial, a third white sjMt behind M-, narrowed behind, rarel}'
extended to M-, having usually the same shape as in Gray's figure 1 on PI. X*
{spartacus, see above under subspec. echemon). This is the ordinary form of
the female, commonly met with.
b'. ? -f. pisander Feld., l.c. Forewing : white patches represented by a few
white scales. There are two females of this form in coll. Felder, no locality
being given ; we have not seen it in other collections.
Hab. Lower and Middle Amazons, north shore; the Guianas.
The type of echioii, a male without locality, takes a somewhat intermediate
position between typical echemon and typical ergeteles ; the baud of the forewing
is narrower than in the latter, the red spots of the hindwing, above, are rather
shorter, and tliere are vestiges of white spots on the underside of the forewing.
In the Tring Museum 8 cjt?, 7 ¥ ¥, from : Camaria, British Gniana, January
l'Ji.)4 (R. Haensch) ; Cayenne ; Surinam ; Ouoribo, March 1893 (Ellacombe).
42. Papilio neophilus Hiibn. (1837).
Seba, Themur. iv. p. 38. t. 30. fig. 25. 26. (J (1864) (marginal spots descr. as being white !).
(J. Papilio Eques Trojanns aeiieas, Cramer (^no» Linne, 1758, err. det.), I'ap. Exot. iii. p. 155.
t. 279. fig. A. B ((1780) (Surinam) ; Esper, Ausl. Schmelt. p. 40. n. 15. p. 60 (1788).
(J. Papilio Eques Trojanus acncides Esper,' ?.c. t. 15. fig. 3. ^ (1788) (ho« text, noii fig. 4).
S- Pa rides yargasus UiihneT, Verz. bek. Schmett. p. 87. n. 909 (1818?) (partim).
? . Priamides neophilus id., \Samml. Exot. Schmett., Zutruge p. 46. n. 499. fig. 997. 998 (1837)
(Surinam).
Papilio aeneides, Gray, Cat. Lep. Ins. Brit. Mus. i. Pa^y. p. 51. n. 247. t. 9. fig. 8. ? (1852) ; Guenee,
Ami. Soc. Ent. Fr. p. 307 (1867) ; Obertb., Et. d'Ent. iv. p. 94. n. 290 (1880) (Trinidad ;
Guyane ; Pard) ; Stand., Exot. Tagf. p. 14 (1884) (Trinidad; Guiana : Amazons).
The early authors considered the male of the present species and the male
of r. aeneas to be the sexes of one species. Esper is quite emphatic on this
pomt. Like Cramer, he describes thjs composite species as aeneas L. On the
( 498)
plate, however, the name aeiieides appears instead. No mention of this new name
being made anywhere in the text, we think Espc'r was not responsible for it, bnt
the engraver of the plate, aeneides being perhaps a missjielling of aeneas. Anyhow,
as the name aeneides was proposed for a supposed species of which the "female"
had already a name {aeneas L.), aeneides is a synonym of this older name
aeneas.*
Hiibncr introduced for the same two insects the name of (jn n/as,/s. This
name, covering exactly the same species as aeneides, is a pure synonym of the
latter.
The first name given to a specimen of the present species alone is neophiliis.
We employ it accordingly for the entire species. We add that the name aeneides
on Esper's plate was entirely overlooked or perhaps sujipressed by tlie older
authors. Gray introduced it again, erroneously referring it to the Para form of
the present insect.
S. Cell of hindwing red, except extreme base. The cell-patch and more or
less also the bases of the spots around the cell have a purplish appearance, owing
to the presence of black scales among the red ones. On the underside, the red
area is reduced to a row of spots standing distally of the cell ; these spots are
pale, the upper scales being white, transparent.
? . Resembles that sex of P. li/sander and aylaope, but is easily distinguished
by the different position of M- of the hindwing, this vein originating from cell as
much proximally (or nearly as much) as vein SC-, the cell therefore being almost
symmetrical in P. neophilus. From P. eurimedes, which has practically the same
nenration as neophilus, the latter is distinguished by the forewing bearing two
or three white patches on disc and a streak in cell, or being devoid of white
patches, or being intermediate between these extremes ; there is never a patch
across the cell, as in areas.
Genitalia: S. Harpe truncate or oblicjuely rounded, usually with three long
apical teeth, sometimes with four, many specimens bearing one or two small
additional teeth ; sometimes, especially often in Peruvian specimens, there is a
row of minute teeth at the ventral edge.
Early stages not known.
Eah. Colombia to Bolivia and Paraguay, the range extending eastwards to
Southern Brazil, the Lower Amazons, Trinidad, and to the Guiauas ; not found
in the western districts of Ecuador and Colombia ; also not, occurring in Brazil from
Rio de Janeiro to Pernambuco, being here replaced by P. zat'i/nthus.
The subspecies are not very sharply defiued in characters. Unlike P. li/sandei;
the males from the various fauuistic districts are fairly well separated, while the
females of some of the geograjihical races come very close to each other, occasionally
overlapping in characters. In the Guianas the female has usually small or no
white patches, rarely fairly large ones. On tlie Upper Amazons and on the
eastern slopes of the Andes from Peru to Colombia, the forewing never bears
distinctly marked spots, while in Venezuela and Trinidad, on the Lower Amazons,
and in Bolivia, Paraguay, Matto Grosso, and Southern Brazil, the white spots are
always large. Bates did not meet with the s])ecies at the Middle Amazons, bnt it
is hardly likely that it is entirely absent from that district. We have it from the
Rio Jnrn^ and Mandos (received from dealers), and Felder descrilied a specimen
from the Rio Negro.
• See r. 418.
( 499 )
a. P. neophilus eurybates Gray (1852).
?. Papilin dimas, Doubleday (nan Fabricius, 1793, err. det.), List Lep. Tun. Brit. Mux. i. p. 12
(1845) (jiarlim).
cJ Papilio ciri/hati:^ Gray, Oif. Lep. Lis. Brit. Mus. i. Pap. p. 51. n. 248. t. 9. fig. 1 (1852)(" Bolivia "
laps. cal. ; Brazil on label of type) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 65. n. 203 (185G) ;
Feld., Vcrh. Zuol. Bol. Ges. Wieu xiv. p. 29G. n. 101 (1864) (" Bolivia" error loci).
9 . r(ii>ilio eiipales Gray, Cut. Lrp. Tns. Brit. Mus. i. Pap. p. 56. n. 262. t. x*. fig. 2 (1852) (Brazil) ;
id.. List Lep. Ins. Brit. .I/«8. i. Pap. p. 68. n. 277 (1856) (Brazil).
$. Pa/iiliii neophilus^ Gray (noii Hiibner, 1837, err. det.), Cut. Lip. Ins. Brit. Mus. i. Pap. p. 56.
n. 262 (1852) (sub synon.).
S- Papilin afiiiiiles local var. riiri/hates, Bates, Trans. Ent. Soc. Loml. (2). v. p. .360 (1861) (" Bolivia ").
? . Papilin sacijiithus var. ? , Bates, Trans. Ent. Six: Loml. (2). v. p. 360. (1861).
? . Papilia zaajnthus ab., Felder, Verh. Zool. But. Ges. Wien xiv. p. 296. sub n. 105 (1864).
(J. Papilia aeneides var. e. P. eurybates, Kirby, Cat. Diiirn. Lep. p. 531. sub n. 76 a (1871) (" Bolivia,"
error loci).
? . Pajiilio zaeyntJius var. b. P. eupales, id., I.e. p. 531. sub n. 76 b. (1871).
d. Forewing rather narrower than in the other subspecies, white spots large.
Hindwing : red spots around apex of cell not shaded with black proximally,
except the first and last ; at least the two middle ones touching cell on underside,
the sj)ots being less reduced than in tlie other forms. Tiie type of eiiri/bafe.'i agrees
with Brazilian specimens, and bears the locality label Brazil, not Bolivia, as said
by Gray.
? . Forewing slightly narrower than in the other races, usually with two
large patches W — M- and a cell-streak, but the patches occasionally vestigial.
Hindwing : red band usually wider than in the other forms, touching cell, there
being sometimes a minute spot in cell ; the spots of underside paler and usually
longer than in the other subspecies.
Had. Sao Paulo ; Matto Grosso.
One single s])ecimen (a female) from Sapncay, Paraguay, 60 miles east of
AssuDcion, has the narrow forewing of this form, but agrees in the red band of
the hindwing with the next.
In the Tring Museum 7 cJc?, 12 ? ? , from : Bahuru, Sao Paulo (Dr. Hempel) ;
R. Bitalha, Sao Paulo; Araras, Matto Grosso; Cuyaba (Andeer) ; Sapucay, Paraguay
(W. Foster).
b. P. neophilus comus subsp. nov.
Papilio eurybates, Hopffer (noH Gray, 1852, err. det.), Stett. Ent. Zelt. xl. p. 50. n. 8 (1879) (Bolivia).
Larger on an average than the preceding.
c?. Forewing : green patch BI-— SM- longer than broad, often bearing a wliite
dot in the u]ii)er distal corner; white spot M' — M= smaller than in ei(ri/ljafi'g,\iXi.t
always very distinct, rounded, sjwt W — M' usually larger than sjjot M' — M-, but
shaded with black ; most specimens with a vestige of a spot R- — R'. Hindwing :
bases of red discal spots shaded with black like cell-patch, spot R' — M' often
excepted — in one of our numerous specimens hardly any black scales in the red
markings, the cell-patch being nearly as pure red as the discal spots ; the s))ots on
underside much shorter than in the preceding, standing separate from cell.
? . Forewing : a large white patch M' — M^ preceded by another large patch,
which is usually somewhat shaded with brown distally ; a more or less distinct
cell-streak, which is in one of our Mapiri specimens enlarged to a triangular patch
which nearly reaches across the cell ; most specimens with a small dot behind M*,
there being also often a small spot marked before R^ Hindwing : band usually
( 500 )
narrower than in tlie preceding, mostly not touching cell, the spots often well
sejiarated from one another.
Ilab. Bolivia : Santa Cruz de la Sierra northward to tlie Beni River.
The individuals from Reyes approach a little the next form.
In the Triug Museum : 27 c? J,2U ? ?, from : Santa (Jruz de la Sierra, .January
to April 1004 (.1. Stcinbach) ; Rio Grande, December 19o3 (.J. Steinbach) ; Blapiri ;
Salinas, Beni K., July 1803 (Stn.irt); Reyes, Beni li., August 1903 (Stuart).
c. P. neophiliis olivencius Bates (1801).
(J $ . Pitpilio ulkvnciiis Bates, Trmis. Ent. Sac. Limd. (2). v. p. 345 (1861) (S. Paulo de Olivenoa ;
Bogota) ; Feld.. Verh. Zonl. Bot. Gets. Wicn xiv. p. 'JOG. n. 102 (18fi4) (Upp. Amazons ; Bogota);
Hahnel, Iris iii. p. 275 (1800) (Sao Paulo de Olivenoa).
Papilla aeneiilex local var. oUveiuiiis Bates, I.e. p. ."iGO (18G1) ; id., Jiiurn. Eiitinn. i. p. 227. sub
n. 23 (18C2) (S. Paulo de Olivenoa).
? . PapiUiianaxiiiitupsV{i\A<iT,Wieii.Eiit. Mun. vi. p. G5. n. 1 (1862) (Upper R.Negro) ; id.,lV)7i. Zuul.
Bot. Ges. Wv'.n xiv. p. 296. n. 100 (1864) ; id , Reise Novnra, Lrp. p. 3G. n. 25. t. 7. fig. b. (1865)
(Upper R. Negi-o).
Pajiiliiiiieveidet! var, b. P. iilivcncius,Kivhy, Cnt.Diiini. Lep. p. 531. .sub n.76a. (1871) (Upp. Amazons).
Pupilio aeiieidea var. d. P. aitaximenes, id., he. p. 531. sub n. 76 a (1871) (R. Negro).
Po/iHio nnieidex, Oberthur, El. d'Ent. iv.p. 116. n. 290 (1880) (Sao Paulo de Olivenva, November) ;
Michael, Iris vii. p. 214 (1894) (Sao Paulo de 01iven(;a).
Papilio nriieides var. etirylales, Maassea & Weym. {nun Gray, 1852, err. det.), in Stiibel, liriscii
S. Amer., Lep. p. 24. n. 105 (1890) (West side of Cordillera of Bogota) ; iid., I.e. p. 79. n. 29
(1890) (N. Peru) ; Haase, Uiitersuch. Mimicnj ii. p. CO. t. 9. fig. 65. ^ (1893).
S. Forewing : spot M' — M^ not distinctly white, being much shaded with
green and brown; spot R'— M' absent or just vestigial. — Hindwing : red discal
spots long, shaded with black proximally, small on underside, being separated
from cell.
?. Forewing with vestigial white spot M'— M' or without trace of su<;h a
spot. In the specimen described as aiiaximenes the red spots of the hindwing
exceptionally long.
Ilah. Up])cr Amazons, from Sao Paulo de Olivenea and Upper R. Negro
westwards ; eastern slopes of the Andes of Peru and Ecuador, as far north as
the Cordillera of Bogota.
In the Tring Museum 40 6 3, 32 ? ? , from : "Bogota"; Villavicencio to
Rio Ocoor, January 1897, 350—400 m., dry season (Dr. Biirger) ; Chanchamayo
(W. Hoffmanns ; Schunke) ; La Union, R. Huacamayo, Carabaya, 2000 ft., December
1904, dry season (G. Ockenden) ; Per(5n(5, March 1900 (Simons); Rio Toro, La Merced,
August— September 19ni (Simons); Cumbure ; Hualliiga; P. Cachyaco, R. Huallaga
(Stuart) ; R. Jurua ; Manaos ; R. Negro.
d. P. neophilus ecbolius subsp. nov.
Papilio aeneides, Gray (n«H Esper, 1788, err,^det.), Cat. Lep. Ins. Brit. Mas. i. Pap. p. 51. n. 247.
t. 9. fig. 8. $ (1852) (Para; syn. excl.) ; Wall., Tram. Ent. Soc. Land. (2). ii. p. 256 (1854)
(Par.'i ; forest) ; Gray, List Lep. Ins. Brit. Miis. i. Pup. p. 65. n. 262 (1856) (parlim ; Parii ;
ViUa Nova); Bates, Trans. Ent. Soc. Lond. (2). v. p. 345, 360 (1861) ( parti m ; Pani to
Obydos) ; id., Juurn. Eiitom. i. p. 227. n. 23 (1862) (Lower Amazons, Tocantins, Guiana);
Kirby, Cat. Diiirn. Lep. p, 530. u. 76 a (1871) {/lurliin ; Lower Amazons) ; Ilahnel, Jris iii.
p. 240 (1890) (VilLabella, Amaz.).
Papilio garijasus, Wallace (mm Hiibner, 1818 ?, err. det.). Trans. Ent. Soc. Lond. (2). iL p. 255
(1854) (Amazons; forest); Felder, IVWi. Znol. Bot. Ges. Wien xiv. p. 298. n. 103 (1864)
(partini ; Para ; Lower Amazons) ; Butler, Trans. Ent. Soc. Lond. p. 145. D. 223 (1877) (Rio
Tapajos, Marcb).
( 501 )
c?. Forewing : green patch ir — SIVP abont as long as broad, rather shorter
than in the previous forms ; white spot M' — M' very distinct, oblong or elliptical,
transverse, spot R^ — M' also distinct, transverse, spot R^ — R^ vestigial. Hindwing;
Ted discal spots shorter than in the preceding forms, the two middle ones nsnally
not shaded with black i)roximally, small on underside, and spot M'— M'^ mostly
closer to cell than in the other forms.
?. Forewing: white patch M' — M- large, touching cell behind base of M', spot
R' — M' more or less shaded with browo, projecting as much distad as spot M' — M^
(a line touching both spots would be parallel to distal margin of wing), a small
streak in cell, often vestigial, a dot behind M- (never absent ?), and often a patch
before R', its distal edge being in a line with the edges of the two patches R^ — M' ;
the arrangements of the distal spots reminding one of P. echemon echemon ?,
P. zacynthus poh/metus ?, and of the S of P. aylaope. Hindwing: red band
much narrower than the black distal area between R* and M-, the middle sjwts
touching cell, on underside at least spot R^ — M' close to cell.
Hab. Lower Amazons, from the Tocantins to Obidos.
In the Tring Museum 5 cJc?, 7 ? ?,from: Igarape (W. Hoffmanns); Santarem ;
Obidos, October— November 1904 (M. de Mathan) ; Juhuty, April lOOo (Mathan).
e. P. neophiliis neopkUaa Hiibu. (1837).
$. PapHio Eqiies Trnjaims aeiieas, Cramer {non Liiine, 1758, err. det.), l.i: (partim ; Surinam) ;
Esper, I.e. (purtiin).
(J. Papilin Equea Trojaii'c^ aenf'nlea Esper, I.e. {partim).
^. PriamitUs f/argasus Hiibner, I.e. (^ parti m).
? . Parides neophiliis id., >iamml. Exol. Srhmell., Zutriige fig. 997. 998 (1837).
(J. Papilio aeiimx, Godart, Eue. Melli. ix. p. .33. n. 24 (1819) (partim, ^ ; Guyane) ; Lucas, Lep.
E.rot. p. 27. t. 13. fig. 3. ^ (1*^35) (Guyane) ; Boisd., Spec. Gen. Up. i. p. 28C. n. 113 (1836)
(jmrtim, ^ ; Surinam ; Cayenne) ; Doubl., List Lep. Ins. Brit. Miis. i. p. 12 (1845) (partim) ;
id., Westw. & Hew., Gen. Diurn. Lep. i. p. 18. n. 202 (1846) (partim).
? . Papili.i nenjihUiis, Gray, Cat. Lep. Ins. Brit. Mas. i. Pap. p. 49. n. 239 (1852) ('■ (J " ! , Surinam)
(J. Papilio aeneides, id., I.e. p. 51. n. 247 (1852) (desor. and fig. exol.).
(J ?. Papilio aeneides, Bates, Trans. Ent. Sac. Land. (2). v. p. 345, 3l'i0 (1861) (partim ; Guiana) ;
Kirby, Git. Diurn. Lep. p. 530. n. 76 a (1871) (partim ; Guiana) ; Staud., E.cof. Tagf. t. 9. cJ ?
(1884).
Papilin r/iirgasiis, Felder, I'tr/i. Znnl. Bot. Ges. Wien xiv. p. 296. n. 103 (1864) (partim ; Surinam ;
Guiana).
Papilio aeneides var. a. Priam, neophilus, Kirby, I.e. p. .^31. sub n. 76 .\ (1871).
Papilio eiiristeas?, Moschler, Virli.Znol. Bot. Ges. Wien xxxii. p. 304 (1883) (Paramaribo, ? ? ).
Papilio eiirimedcs var. eur.i/hates, Muscbler (non Gray, 1852, err. det.). I.e.
Parides aeneas, Kirby (nun Linne, 1758, err. det.), in Allen's Nat. Libr., Lep. Butt. ii. p. 271. t. 66.
fig. 2. cJ (1896).
c?. Similar to ccholiiis, green patch wider ; white spots not quite so distinct ;
red spots of underside of hindwing rather smaller, spot M' — M- farther away
from cell.
?. Forewing: white s})Ots absent, or vestigial, most specimens having two
small round spots R^— M'-, rarely both patches large (occasionally in British
Guiana). Hindwing : band often distant from cell, third spot the longest ;
black distal area of uj)pcrside wider than in oUteitcitis.
llab. The Guiunas.
In the Tring Museum 10 (?<?, Ill ? ?, from: (laycnm.' ; Surinam ; Bartica,
Brit. Guiana, March— April 1901,
( 502 )
f. P. neophilns pariamis snbsp. nov.
PapHin aeneidpK, Oberthiir {iMii Esper, 1788, err. det.), FJ. iVEiil. iv. p. 94. n. 20O (1880) {imrllm ;
Trinidad) ; Staud., Exnt. Tmxf- P- 1* (1884) {jmrlim ; Trinidad).
(?) Paiiilio (jaryaniii,, Kaye (;m« Hiibner, 1818?, err. det.), Tnnix. Ent. Soc. Lnml. p. 200, n. 19G
(1904) (Trinidad ;— this insect?).
cJ. Forewing : green patch M- — SM" longer than Ijroail ; white sj)o(; M' — W
large, spot R^— M' nsnally larger than M'— JP, a more or less distinct spot II'-— R' ;
green streak at inner margin broad. Hindwing : red pateh less ])ahnate tlnin
in the Bolivian form, sjwt R'— 11' rarely pnrple at base, the spots paler beneath
than in consus, W — M' close to cell.
S . Forewing : a large jiatch M'— M-, a somewhat smaller one R'— M', often
a spot R-— R' and a dot behind W-, cell-streak distinct in most specimens.
Hindwing : band more S-shaped than in the other forms, narrow middle spots close
to cell above and below, jialer beneath than in the Bolivian subspecies, with which
this subspecies agrees best.
Hab. Trinidad ; Venezuela : Cumana and Orinoco.
In the Tring Museum 55 6<S, 50 ? ? , from : I'aparo valley, Trinidad, December
1896 and January and February 1897 (Dr. P. Rendall) ; Maraval, Trinidad, .July
1891 ; (lampo Alegre, Cumana, 1500 ft., April 1899 (Andre) ; Patao, Guiria, Augnst
1891 ; Maipures, Orinoco, December 1898 (Cherrie) ; Suapure, Caura R., February
and March 1899, October 1900 (S. M. Klages) ; La Vuelta, May 1904, and Corosito,
June 1904, Caura R. (S. M. Klages).
43. Papilio zacynthus Fabr. (1793).
(J. Papilio Equei Trojatms zacynthus Fabricius, Eiil. St/st. iii. 1. p. 15. n. 4G (179S) (Brazil).
? . Paiiilio Eques Trojamix dimas id., I.c. p. 16. n. 47 (1793) (Brazil ; cit. Cram, e.xceptis).
(J ? . Pajiilio zacynthus, Doubleday, Westw. & Hew., Gen. Diuni. Lfji. i. p. 18. a. 211 (1846) {ilimas
? of zacynthns) ; Felder, Vcrh. Zool Dot. Ges. ]Vien xiv. p. 296. n. 103 (1864) (Brazil^ J =
puli/melus ; 9 aberr. = enjniles, error!) ; Burm., Descr. lii'p. Arr/eiil. v. Lep., Alias p. 7. n. 17
(1879) (Rio ; larva mentioned) ; Stand., Exot. Tagf. i. p. 14 (1884) (Bahia ; Rio de Janeiro).
(J. Foiewing with greenish bine patch and at least two white spots, lied baud
of hindwing sei)arate from cell or contiguous with it, no red spot in cell or only
a minute one. Midtibia spinose as in P. neoj>/iilus.
? . Very close to certain females of P. neojMus ; either the white spots of
forewing more or less rounded and no spot in cell, or spot M'— M" oblong, large,
and a sharply defined sjiot in cell ; band of underside of hindwing paler pink than
in neophilus, there being hardly any black scales in the band.
Scent-organ and genitalia as in P. neophilus.
Early stages mentioned by Bnrmeister, l.r.
Ilab. Brazil, from Rio de Janeiro to Pernambuco.
In the district where xacjjiithua occnrs P. ncoplrilm is not found. Tiio two
insects are so closely related that it is quite possible there exists an intermediate
form, perhaps in Goyaz, where the Brazilian and Amazonian faunae meet. Of
the two subspecies of /'. zaajntlms the more northern one agrees better with
P. neophilus than the sonthern one.
a. P. zaci/nthus pohjmelus Godt. (1819).
Painlio pohimetus Godart, Enc. MM. i.x. p. 35. n. 28 (1819) (i, Brazil ; " Peru," error) ; Swains.,
Zoul. iUuslr. iii. t. 92. S ? (1823) (Bahia; var. excl.) ; Lucas, Up. Exol. p. 10 t. 6 tig.
(18:!5) (fig. mala? hacc subsp. ?) ; Boisd., I.e. p. 283. n. 108 (1836) (;;«rt/m); Doubl., Westw.
& Hew., Gm. Viurn. Lrp. i. p. 18. n. 212 (1846) (cil. Hiibn. excl.); Oberth., Et. il' hid. iv. p. ii-
n. 288 (1880) (Brazil).
{ 503 )
Papilio cliiiias, Godart, I.e. p. 36. n. 33 (1819) {partim).
Pupilio zaryuthus var. a. Papilio jyoli/iiietiis, Gis.y, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 5G. sub n. 201
(1852) (Brazil) ; id., list Lep. Ins. Brit. Mils. i. Pap. p. 68. sub n. 276 (1856) (Brazil ; " Pard,"
error loci) ; Kirby, Cat. Diurn. Lep. p. 531. sub n. 76 B (1871).
Papilio ursilius Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 57. n. '-'64 (1852) (Pernambuco) ; id., List
Lep. Ins. Brit. Mus. i. Pap. p. 68. n. 279 (1856) (Pernambuco; Tapajos) ; Bates, Trans. Ent. Soc.
Land. (2). v. p. 346, 360 (1861) ; id., Juurn. Entum. i. p. 227. n. 24 (1862) (Pernambuco ;
Tapajos, not in the Amazonian plains) ; Feld., I.e. n. 104 (1864) (Bahia ; Pernambuco ; R.
Tapajos).
Papilio sacijnilais Fabr. var. polymelus, Bates, Trans. Ent. Soc. Land. (2). v. p. 346, 360 (1861) ;
id., Jomm. Enloni. i. p. 227. n. 24 (1802) (" Pari," locality doubtful !).
Papilio pohjmatus (!), Guenee, .\nn. Soc. Ent. France p. 308. note (1867) (= zacynthus ; ? dimas).
Papilio zacynthus var. c. P. orsillus, Kirby, I.e.
Papilio zacytithns, Grimshaw, Trans. Roy. Soc. Edinh. xxxix. i. No. 1. p. 7 (1897) (in Edinburgh
Museum, one of Godart's types).
cj?. Apex of forewing semitransparent ; in male spot M' — M- a little smaller,
but purer white, tban spot R^— M', separate from cell, rarely a white dot behind M^;
in female spot M'— M- a little larger, or at least broader, than sjjot R^ — M',
usually a spot R^— R', but not cell-spot.
Ilab. Pernambuco ; Bahia ; R. Tapajos.
In the Triug Museum ~ S <S and 3 ? ¥ from Pernambuco and Bahia.
b. P. zacynthus zaci/nthus Fabr. (1793).
cJ. Papilio Eques Trojanus zacynthus Fabricius, I.e. (Brazil).
? . Papilio Eques Trojemus (liinas id.. I.e. (Brazil),
$ . Prianiales hijjpasan, Hiibner, Verz. hek. Schmett. p. 87. n. 906 (1818?) (partim).
(J. Papilio zaeynlhus, Goda.Ti, Ene. Metli. ix. p. 34. n. 25 (1819) ; Douov., Nat. Repos., Ent.n.
t. 26, 27. fig. 1 (1823) (Brazil) ; Boisd., Spec. Gin. Lep. i. p. 284. n. 109 (1836) ; Doubl., List
Lep. Ins. Brit. Mus. i. p. 11 (1845) (cit. Hiibn, excl. ; Brazil).
?. Papilio rf/Hin.s, Godart, /.c. p. 36. n. 33 (1819) (Brazil, ^iro^'m) ; Donov., I.e. fig. 2 (1823) (Brazil);
Lucas, Lep. E.eol. p. 30. t. 14. fig. 2 (1835) ; Boisd., I.e. p. 292. n. 120 (1836) (Rio de Janeiro ;
" $ var. in coll. Roger," probably J neophilus) ; Doubl., List Lep. Ins. Brit. Mus. i. p. 12
(1845) (Brazil).
Papilio pvlymetus, Swainson, I.e. (1823) (var., Rio de Janeiro).
(J ?. Papilio zacynthus, Doubleday, Westw. & Hew., Gen. Diurn. Lep. i. p. 18. n. 211 (1846) (dimas
5 of zacynthus): Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 56. n. 261 (1852) (Brazil); id., List
Lep. Ins. Brit. Mus. i. Pciji. p. 68. n. 276 (1850) (Brazil) ; Butl., Cat. Diurn. Leji. descr. Fabric.
p. 237. n. 12 (1869) (Brazil) ; Kirby, Cat. Diurn. Lep. p. 531. n. 76 b (1871) (BraziUa) ;
Capronn., Ann. Soc. Ent. Belg. xvii. p. 8. n. 3 (1874) (Jacarepagua, August) ; Haase, (/nfersucA.
Mimicry i. p. 79 (1893).
Papilio demas (!), Doubleday, Westw. & Hew., I.e. ii. p. 529 (1852).
cJ ? . Papilio dimas, Oberthiir, Et. d'Ent. iv. p. 93. n. 280 (1 880) (Brazil; difference irom jiolymetus)
Papilio zacinthua (!), id., I.e. iv. p. 9.3. sub n. 289 (1880) (err. cal.).
S ? . Apex of forewing much more opaque than in polijmetus, the wing
altogether deeper black, the spots purer white, spot M' — M" much larger than
in poljjmctus, lowgev than broad, spot R' — M' smaller, in both sexes a white spot
behind M", in female a spot in cell, small but sharply defined. Baud of hindwing
narrower than in polymetus.
Hub. Province of Rio de Janeiro.
In the Tring Jliiseum : 3 J J and 3 ¥ ? I'runj Rio de Janeiro (E. May).
44. Papilio areas Cram. (17S1).
{•) Papilio Eques Trojanus viridimaculatus Goeze, Ent. Beylr. iii. 1. p. 44. n. 24 (1779) (this
species? or P. aeneas ?— Type : Seba, Thes. t. 30. fig. 25. 26. ^).
? . Papilio Eques Trojanus areas Cramer, Pap. Exot. iv. p. 174. t. 378. fig. C (1781) ("Brazil").
( 504 )
(J. PapiVii Eques Tinjanus eitri ineiles id., /..■. iv. p. 199. t. 38(1. fig. E. F (1782) (Berbices).
(J 5 . Pdjiilio luri/metli'S (!). ErichsoD, in Schomb., F. F. Brit. Guiana p. 59.^ (1848) ( ? = arriphus).
^ J. Papilio eiirinmUs, Kirby, Cat. Diurii. L-p. p. 530. n. 76 (1871) (" var. b. P. limiuH" excl.) ;
Obertb., El. d'Enl. iv. p 95. n. 292 (1880) (synon.;OTr((Hi)-
J?. Antenna deeper lilack than m P. neophil)is,ihe sensory grooves, thongh
sharply defined, not so distinct, being bhxck like the rest of the antenna. Mid-
tibia mnch more densely hairy than in P. neophilus. Apical half of forewing
opaque in both sexes. In male the cell of hindwing red from centre to apex on
npperside, the cell-patch being, however, often reduced, sometimes absent,
occasionally occupying about three-fifths of cell. In female forewing with a broad
white (slightly yellow) patch across cell, a large patch R^ — M' on disc, a smaller one
R- — R', and sometimes a small spot behind M'. Abdominal fold as iu P. neophilus.
Genitalia: i . Harpe truncate, with two long spinelike teeth at apex,
sometimes accompanied by one or two small ones.
Early stages not known.
Hub. Mexico to Colombia, Venezuela and the Gniauas.
Seba's figures, on which the name viridmaculatus was based by Goeze, are
rough. The underside (fig. 26) does not agree with any Papilio known. The
marginal spots are described as being white. The figure may have been meant
for P. aeneas, areas, or even neophilus.
a. P. areas mi/lotes Bates (1861).
(J 9 • Piiji'l'" iinjlfiles Gray, List Lep. Iiis. Brit. Miis. i. Pap. p. 64. n. 258 (1856) («om. niid. ;
Mexico ; Nicaragua) ; Bates, Trans. Ent. Soc. Lniid. (2). v. p. .346. note (1861) (Nicaragua ;
descr. of (J J ) ; Weidem., Pror. Enl. So,:. P/iilad. ii. p. 147 (1863) (Mexico ; Central America) ;
Felder, Verh. Zuol. Bot. Ges. Wie.n xiv. p. 296. n. 109 (1864) (Nicaragua ; Mexico ? ; ihjciinus J
of miilotes); Butl. & Druce, Pro,-. Znol. Soc. Loud. p. 364. n. 365 (1874) (Costa Rica); Strecker,
Bull. ^Toths N. Amer. p. 68. n. 4 (1878) (" S. California " false ; Mexico, Nicaragua, Panama) ;
Godm. & Salv., Biol. Ceiitr. Amer., Rhop. ii. p. 198. n. 13. t. 65. fig. 9. 9a. genit. (1890) (Mexico
to Costa Rica).
PapiU'i docimus Gray, List Lfp. Ins. Brit. J/h.s. i. Pap. p. 64. n. 259 (1856) {iioni. mul. ; Nicaragua) ;
Weidem., I.e. p. 147 (1863) (Mexico).
cJ. Pupilio caleli Reakirt, Proc. But. Sor. Philad. ii. p. 138. n. 8 (1863) (Guatemela); Felder, Verh.
Zoo!. Bot. Ges. nVewxiv. p. 296. n. 107 (1864) (Guatemala; tonila ? otraMi?); Strecker,
Lep. Rhop. Iht. p. 15. (1873) (" = polymetus Godt.") ; id.. I.e. Suppl. iii. p. 17 (1900) (type in
coll. Strecker).
? . Papilio tonila Reakirt, /.<■. p. 140. n. 10 (1863) (Guatemala); Streck., /.-;. iii. p. 17 (1900) (type
in coll. Strecker).
(J. Papilio almincdfs Felder, Verk. Zool. Bot. Oe$. Wicii xiv. p. 286. n. lOG (1864) {uom. mul.;
Nova Granada ''.) ; id., Reise Novara, Lep. p. 36. n. 26. t. 7. fig. e (1865).
?. Papilio aristuiuenes Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 296. n. 108 (1864) [mim. mul.;
Mexico) ; id., Reise Xovara, Lep. p. 33. n. 27. t. 7. fig. a (1865).
Papilio euriiiiedis, Boisduval, Consid. Lip. Guatem. p. 6 (1870) (Honduras; Nicaragua; Costa
Rica; " Venezuela " alia subspec. ; alcamedes = etiriinedes).
Papilio eiirimedes var. c. P. mylotes, Kirby, Cal. Diiini. I^ip. p. 530. sub n. 76 (1871) (Nicaragua).
Pap>ilio eurimcdes var. d. P. tonila, id., /.<•. (Mexico).
Papilio enrimeiles var. e. P. caleli, id., I.e.
Pajiilio zaeynlhus var. P. polymetus, id. («tw Godart, 1810, err. det.), /.•■. p. Sii'.i. n. 76b (1.177)
(caleli = polymetus, error).
S. Forewing: green patch R' — M'. long, rarely without white spot; cell
usually with green and white spot; no streak at inner margin, patches ]M' — SM-
large.— — Hindwing : red spots large, all separate from cell on underside.
?. Band df hindwing sejiarate from cell above and below, bright red, uo spot
SC*— R'.
( 505 )
la coll. CJharles Oberthur there is a curious aberration of the male, from
San Pedro Sula, Honduras. The specimen has a large pale pink patch on the
underside of the forewing, the band on the underside of the hindwing being
also pale pink.
Ilah. Mexico to Costa Rica.
In the Tring Museum : 57 SS, 52 ? ¥ , from : Motzorougo, Mexico ; Maza-
tenauga and Retalhuleu, W. Guatemala, 1000 ft., September 1904- (A. Hall);
Escuintla, W. Guatemala, 1100 ft., August 1904 (A. Hall); San Pedro Sula,
Honduras ; Carillo, Costa Rica, 3000 ft., October 1904 (A. Hall) ; Carillo, June —
July 1903 (Underwood); Carthago, Costa Rica (Underwood); San Jose, (_'ost«i
Rica, September 1904 (A. Hall) ; Pozo Azul, Costa Rica, June 1902 (Underwood) ;
Guatil Pirris, Costa Rica, January 1902 (Underwood).
b. P. areas mijcale Godm. & Salv. (1890).
PapiUo niijcah Godman & Salv., B'idI. Cailr. Amci:, Rhoji. ii. p. 19'J. n. 14. t. 65. fig. 12. (J, 13. ?
(1890) (Panama : Chiriqui, Bugaba, Lion Hill, Veraguas).
c?. Forewing : green spot R^ — M' always present, often with white dot ;
patch M' — M- large, usually extended close to cell, filling in base of cellule M' — M- ;
patch M' — SM- sometimes reduced posteriorly ; streak at inner margin present
or absent ; occasionally a white dot in cell on underside. Hindwing : cell-spot
and bases of discal spots more or less shaded with brown.
?. Hindwing with or without a small cell-spot, the band deeper red above
and below than in the Colombian forms, sometimes separate from cell on underside.
Ilab. Northern Panama and the islands off the south coast.
Completely connecting P. a. m>/lotes with P. a. arripkus.
In the Tring Museum : 24 tJc?, 11 ? ?, from : Chiriqui ; Bogava, 800 ft., and
Boquete, 3500 ft. (Watson) ; Brava I., January 1902, and Cebaco I., February 1902
(J. H. Batty) ; Colon.
c. P. areas arripkus Boisd. (1836).
? . Pajiilio arrijihm Boisduval, Sjiec. Gen. Lip. i. p. 393. n. 123 (1836) (Colombia).
? . Pajtil/o seraph, Doubleday (iion Boisduval, 1836, err. det.), in Doubleday, Westw. & Hew., Gen.
Diurn. hep. i. p. 18. n. 196 (1840) (parthn ; Colombia) ; Weidem., Proc. Eut. Soc. Philud. ii.
p. 148 (1863) {amphiis J of s^crapis, false).
S- PaplVo agallwklm Kollar, Deiihsrhr. K. Ah. Wlss. Wien, Math. Nat. CI. i. p. 352. n. 4 (1850)
(" R. Orinoko ") ; Doubl., Westw. & Hew., Gen. Diurn. Lep. ii. p. .529 (1852).
Pupilio eurimedes, Gray, Cat. Lep. his. Brit. Mus. i. Pap. p. 50. n. 244(1852) (partim ; arriphus ? of
eurimedes) ; Felder, Verli. Zm,l. But. Ges. Wien xiv. p. 296. n. Ill (1864) (partim) ; Kirby, Cat.
Diurn. Lep. p. 530. n. 70 (1871) {partim ; N. Granada).
PapiUo arrhipus (!), Weidemeyer, I.e.
PapiUo arripus (!), id., I.e.
PapiUo eurimedes var. a. P, agatkocles (!), Kirby, Cat. Diurn. Lep. p. 530. sub n. 76 (1871).
PapiUo eurimedes var. mi/lotes, Staudinger («y« Bates, 1861, err. det.), Exut. Tatjf. p. U. t. 9. (J ?
(I8.S4).
S. Forewiug : spot R^ — M' nearly always distinct, usually isolated, often
centred with white. Hindwing with cell-spot on upperside, not on underside.
?. Band of hindwing entering cell, pale, its inner edge crossing cell in most
specimens just distally of point of origin of M-.
Hah. Colombia : Magdalena valley, and R. Meta, east side of Cordillera of
Bogota.
( 506 )
Tbe locality given b_y Kollar for his agiitholdi's is doubtless erroneous. The
specimen agrees with certain individuals from Colombia iu which tlie patch of
the hindwing is very jiale. We are the more convinced that agathoklcs came
irom Colombia, as Kollar records also Papilio americus from the Orinoco, where
it is hardly likely to occnr, being in Colombia and Venezuela a species of high
altitudes. Prince Sulkowsky, who brought these specimens home, went up the
iiio Slagdalena in Colombia, crossed the Cordillera of Bogota and came down
the Rio Meta and Orinoco, which accounts for errors in localisation.
In the Triug Museum 40 Si, 30 ? ? , from: Valdivia, July 1897 (Pratt);
Pacho, November IS'JS ; Muzo, November 1896 ; Purnio, October — November 1896
(Dr. Burger) ; Villavicencio.
Very common in " Bogota " collections.
d. P. areas antheas snbsp. nov.
Papilio agathol-leSjfila.a.sseD & Weym. {non Kollar, 1850, err. det.), in Stiibel, Reiscii in S. Amer.,
Lep. p. 3li. n. 37 (18y0) (La Plata, Cauca).
c?. Forewing : green band more or less reduced, no spot before M' either
above or below. Hindwing : no spot in cell or only a minute one, the band
pale in most individuals and very narrow : first spot of underside, R' — R-, much
smaller than the last.
?. Forewing : patch R- — R^ on the whole larger than in an-iphus, sometimes
nearly as long as patch W — M' ; most specimens with a white streak in front
of apex of cell ; cell-patch often completely filling in apex of cell. Hindwing :
band very pale, narrow, standing ontside cell or just entering apex ; spot SO- — R'
(the first of the Bogota form) absent from most specimens.
Hab. Cauca valley.
This is one more instance of the reduction of the markings observed among
the Cauca Papilios. As in the case of P. eritkalion cauca and P. euryleon pithonius,
we find also here all intergradations between the Cauca and Magdaleua or " Bogota"
forms. It appears to us that intergradations are especially often oliscrved iu the
lower as well as the upper districts of the Cauca valley, while the typical (_'auca
forms come from the middle portion of the valley.
In the Tring Mnseum 40 c? c? and 34 ? ? from Pereira and Popayan.
e. P. areas areas Cram. (1781).
(?) Seba, Thesaur. iv. p. 3fi. t. 30. fig. 25. 26 (1764) (fig. malae).
(?). cJ. Papilio Eques Tmjdmis viridimacuhilus Goeze, Eiit. Beytr. iii. 1. p. 44. n. 2-1 (1770) (this
species?).
? . Papilio Eqms Trojanus areas Cramer, I.e. iv. p. 174. t. 378. fig. C. (1,781) (" Brazil) "; Jabl. & Ilerbst,
Nalura. Schmell. ii. p. 83. n. 24. t. 10. fig. 3 (1785) ; Jung, Alphab. Ver:. Schmctl. p. 46 (17U1) ;
Espcr, U. p. 146. n. 68. t. 38. fig. '2. (1703).
? . Papilio Equcs Trojaiius curiiiiales Stoll, /.(,-. ; Esper, Ausl. Schmctl. p. 60. n. 26. t. 15. fig. 2 (1788).
Pariden aeimis, Hiibner (mm Liunu, 1758, err. dot.), Verz. hrk. Schmctl. p. 87. n. 008 (1818 ?).
Papiiliii CHivmerfM, Godart, Eiic. .Mclh. ix. p. 34. n. 26 (1810) ((J, Guyane) ; Boisd., Spec. Hin. Liji. i.
p. 285. n. Ill (183i;) ; Doubl., Westw. & Hew., den. Diitni. Lqj. i. p. 18. n. 204 (1846)
(Guiana ; Venezuela) ; Gr.ay, Cat. Lep. Ins. Brit, .\ttis, i. Pap. p. 50. n. 244 (1852) (Venezuela ;
sjnon. partim) ; id., List Lep. Ins. Brit. Mus. i. Pap p. 64. n. 257 (1856) {partim ; Venezuela) ;
Felder, Verh. Zool. But. Ges. Wieu xiv. p. 296. n. Ill (1864) (partim); Hahnel, Jris iii.
p. 138 (1800) (San Est.''ban, in forest); Poujade, Bull. Snc. Eiit. Frame p. 140. n. I (1895)
(Venezuela).
( 507 )
?. Papillo areas, Godart, I.e. p. 37. n. 35 (1819) (-'Brazil") ; Boisd., I.e. p. 293. n. 122 (183G)
("Brazil") ; Doubl., Westw. & Hew., I.e. i. p. 18. n. 195 (18iG) (" Brazil") ; Gray, Cat. Lep.
Inn. Brit. Miis. i. Pap. p. 40. n. 228 (1852) (" Brazil ") ; id., List Lep. Ins. Brit. .V«.«. i. I'ap
p. 61. n. 241 (1850) ; FeWer, Verh. Zool. Bnt. Gen. Wien xiv. p. 295. n. 60 (1864) (Brazil ?) ;
Kirby, Cat. Diurn. Lep. p. 527. n. Ola (1871) (Brazil ?)•
Fajidio eurymeeUs (!), Erichson, in Schomb., F. F. Brit. Guiana p. 593 (1848).
(?. Forewing with white spot R^ — M' above and below, seldom vestigi.al. ■
Hindwing : red patch large, entering cell on underside as well as upper, but the
cell-spot usually much shaded with black on underside.
?. Red band of hindwing large, brighter red than in the Colomljian forms.
Hab. The Guianas ; Venezuela.
The insect figured by Cramer as areas is without donlit the female of the
present subspecies. The fringe-spots of the hindwing are white in the figure ; but
that is surely a mere error in coloration, since similar mistakes occur in other
figures — for instance, in fig. F of PI. 386, which represents the male of areas
(= eurimedes).
In the Tring Museum 16 c?c?, 14 ? ?, from : Cayenne ; San Esteban, October,
1896 (Dr. Biirger) ; M6rida and Tachira (Briceuo) ; Valencia.
45. Papilio timias Gray (1852).
(^. Papilio timias Doubleday, Westw. & Hew., Gen. Diurn. Lep. i. p. 18. n. 203 (1846) (uoin. nud. ;
Guayaquil) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pa/), p. 50. n. 242. t. 9. fig. 3 (1852) ; id., Li.it
Lep. In». Brit. ilus. i. Pap. p. 64. n. 255 (1856) ; Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 296.
n. 110 (1864).
?. Papilio bimaeulatus Hewitson, E.tut. Butt. v. Pap. t. 14. fig. 47 (1875) (Ecuador).
cJ ? . Papilio eurimedes var. b. P. timias, Kirby, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 530. sub n. 76
(1871) ; id., (.c. p. 809. n. 76 (1877) {bimacukitus ? of timias).
This insect takes a similar position towards P. areas as does P. zaci/nthus
towards P. neophilus (= aeneides auct.) ; but, while in zacyntkus it is the male
which differs more essentially from P. neophilus, it is the female of timias which
differs conspicuously from P. areas.
There is apparently no structural diiference between timias and areas. Male
with a large green patch on forewing from inner margin beyond R' and two
conspicuous white spots R^ — M' ; apical half or third of cell of hindwing red. In
female the forewing bears two rounded spots, R-— BI', white, more or less shaded
with brown, the upper one being the larger; red baud of hindwing across apex of
cell, its proximal edge nearly straight. Cell of forewing with or without small
subapical spot, never with broad patch as in ? P. areas.
Hah. West Ecuador.
Two subspecies.
a. P. timias timias Doubl. (1840).
(J. Papilio timias Gray, I.e. (Guayaquil).
? . Papilio bimaeuliilus newitson, I.e. (Ecuador).
P'j/iilio eurimedes var. timias, Maas. & Weym., iu Slubel, Rcisen S. Amer., Lep. p. CG. u. 23 (1890)
(Guayaquil).
S ? . l\:pdi<t bimaeulatus, Haeusch, Berl. But. Zeitselir. xlviii. p. 150 (1903) (Palmar, W. Ecuador,
Jnb).
c?. Forewing : no green spot in cell, or only a few grecu scales ; green patch
M- — fcjM" usually not extended to base of M^. Hindwing : red spot R^— R- of
anderside (first spot) longer than, or at least as long as, the last spot;
33
( 508 )
? . Forowing : no spot in cell or only a vestige ; white discal spots more or
less washed with brown, often small. Hindwiug : cell-spot usnall)' shaded with
brown proximally, spot R' — R- at least as long as spot M' — M^, the band somewhat
narrowing behind and dee]ily incised distally at the veins ; on underside the band
more strongly narrowing than above, spot R' — R- being much longer than spot
M- — SM- ; no spot at abdominal edge or only a vestige of a spot.
Hab, Guayaquil and neighbouring districts at a low elevation : Chimbo, La
Chima, Los Rios, Bahahoyo, Arenillas, Palmar, etc.
In the Tring Museum 22 SS, 13 ??, from: Cachabi, .January 1897 and
Chimbo, 1000 ft., August 1897 (Rosenberg) ; Palmar (R. Haensch) ; Naranjas,
Guayaquil (0. T. Baron) ; Quevedo (v. Bnchwald).
b. P. timias potone snbsp. nov.
S. Forewing : a more or less distinct cell-spot ; green patch wider than in
P. t. timias, white spots generally larger. Hindwing : spot R'— R- not longer
than M' — M-, streak M- — (SM') usually long ; on underside spot R'— R- smaller
than spot M-— SM-.
¥ . Forewing : mostly with conspicuous cell-sjiot, patch R'' — M' usually larger
than in P. t. timias. Hindwiug : band widening behind ; on underside spot
R' — R- shorter than M- — SM- ; a distiact spot at abdominal edge, the cell-spot
and the middle ones larger than in the preceding, the veins separating them less
extended black.
Ilab. AVest Ecuador : Paramba ; also Ambato. At higher elevations than the
preceding subspecies.
In the Tring Museum 6 cJ^j, 10 ? ?, from : Paramba, 3.500 ft., February and
March 1897 (Rosenberg) ; Paramba (Flemming & Miketta).
Subsection B.
Antenna without distinct grooves ; end-.segment very short, more or less
completely fused with the preceding segment. Claws more or less symmetrical, at
least of hindleg. Markings of body not bright red, being white or yellow, rarely
rufous, in this case the hi[idwing with a red dot at base ; dots on abdominal
sternites (if present) always white ; no red spots or band in centre of hindwing;
submarginal sjiots always present, often also admarginal ones. Subbasal cell of
hindwing not widening apicad ; cell rounded at apex, cross-vein D' leaning distad
anteriorly, the cell-angle D^ — D' being larger than the cell-angle U- — D^ or at
least as large.
S. Scent-organ never woolly, consisting of a stripe of short scales, there being
a naked streak along the fold on the discal side. Tenth abdominal tergite shorter
than the sternite. Clasjier short ; harpe very short, not reaching to centre of
clasper, usually ending in two teeth, of which one is lost or modified in many forms,
there being often some small additional teeth. Tibiae not iucrassate, similar in
the sexes.
?. Vaginal cavity large, covered on hinder side by a. smootli convex sclerite,
which bears distally a small groove of which the proximal edge is raised, somewiiat
resembling the lip of a jug. Proximally within this cavity two large membranaceous
hairy flaps, which are either si'parated or are almost merged together. Ana
( 509 )
segment proximally with some short stont spinelike bristles. The lateral tubercle
of the prothorax of the larva is mnch prolonged.
The species are miicli less numerous than in Subsection A. Though some of
the species are more nearly related with one another than with the other members
of the Subsection, the differences in structure are very slight. It appears to us,
therefore, advisable to keep all the species united in one grou]), by which means
the close relationship between them will be best emphasised. In the key the
species are grouped according to relationship.
IV. Polydamas Group.
Key to the species : —
a. Tailed ; tail sometimes absent, such specimens recognisable
as belonging here by tlie strong blue-green gloss in the
distal half of the underside of the hindwing ; white
discal spots on underside of hindwing, besides the
snbmarginal row of orange or red spots ... b.
No tail ; spots of body rufous red, or underside of hindwing
with white central band ...... c.
No tail ; spots of body yellow ....... d.
No tail ; breast and sides and undersurface of abdomen
greenish yellow, long-hairy ..... i\
b. Hindwing below strongly glossy bluish green in outer half. Species No. 46.
Hindwing below not glossy green, distally with several
small white spots in middle of wing behind cell and in
apex of same ........ Species No. 47.
Hindwing below not glossy green distally, with a band of
large white patches in middle ..... Species No. 48.
c. Forewing without band of spots on npperside . . . Species No. 49.
Forewing with band of sjiots on npperside ; hindwing
below for the greater part creamy buff . . . Species No. 50.
Forewing with band of spots on npperside ; hindwing
below brown Species No. 51.
d. Hindwing below with red spots Species No. 52.
Hindwing below without red spots ..... Species No. 53.
e. Hindwing above with a regnlarly curved discal row of
seven greenish white spots parallel to distal margin,
separate from cell, the last spot, which is double, being
at least as large as the second ; snbmarginal spots
also marked ; forewing with a row of spots . . /.
Discal band of hindwing absent, or touching cell, or
entering it, or the spots decreasing in size, the last ones
being minute, while the first is very large ... g.
Hindwing above with white streak along abdominal fold . S[tecies No. 58.
f. lied snbmarginal spots on underside of hindwing transverse,
contiguous with creamy white spots .... Species No. 54.
lied snbmarginal spots of hindwing arched ; no white
admarginal spots Si)ecies No. 55.
( 510 )
White costal patch on iipperside of liindwing of male very
large, extending to base ; forewing with white streak
in cell and some large patches oa disc (females and
most males) ........ Species No. bi).
White costal patch of hindwing not reaching base, creamy
white ; no streak in cell of forewing and no large
patches on disc close to cell ; or there is a large patch
in apex of cell . ........ h.
Hindwing below with small white adiiiarginal dots . . Species No. 56.
Hindwing below without white admargiual dots ; red snb-
marginal spots thin Species No. 57.
40. Papilio philenor L. (1771).
Papilio Eques Trnjanus philenor Linne, Mant. Plant, p. 535 (1771) (America).
Pajjilio Eques Trnjanus astinous Drury, Ilhislr. Exut. Ins. i. p. 21. t. 11. fig. 1. 4. (J and Index (1773)
(New York ; Maryland ; Virginia).
Larrtias phitemir, Hubner, Verz. be!.: Schmett. p. 84. n. 858 (1818?); Scudd., Bull. E. U.S. tfc Canada
ii. p. 1-241 (1889).
cJ ? . A close ally of P. poli/damas, being more generalised than that species
in the preservation of a tail (in most specimens), and of dots on the abdominal
sternites, and more specialised in the development of metallic colours and in tlie
reduction of the markings of the upperside.
The presence of white discal dots on the underside of the hindwing is also
a generalised character, wliich is still more in evidence in P. devilliers and zetes.
Comparing the forewings of pohjdamas and philenor a close agreement in the
phyletic development of the spots will be observed. lu both species the spots
in the aj)ical region are the first to disappear, being longer preserved on the upper
than on the underside, the apical area of tlie underside assuming a faded tint. The
posterior spots of the upperside, however, which in pohjdamas are always present,
while they are often absent like the anterior ones in philenor, are always smaller
than the respective spots of the underside, the latter spots being preserved when
those of the ui)iierside have disappeared.
The develojiment in the markings of the hindwing is o]>posite in the two
species. In poli/damas the band of spots of the upperside remains comparatively
broad, in connection with the development of the spots of the forewing, the spots
being rarely reduced to narrow lunules. The corresponding spots of the underside
have been shifted towards the distal margin, and are reduced to more or less
narrow bars. In philenor, on the contrary, the spots of the upperside arc much
reduced, being sometimes absent, those of the underside being much larger.
The preservation of a tail in P. philenor and the West Indian zetes (Haiti)
and decilliers (C'uba), and the occasional absence of the tail from Mexican
specimens of philenor, are significant facts worthy of special mention. If the
tail is a nseful appendage developed to give protection to the individual, the
projecting tail being taken hold of by an insect-feeder and breaking off, altbrding
the insect a chance of escape, as is the opinion of adherents of this variety of
Natural Selection, we must naturally conclude that in countries where there is
a greater number of tailed species the persecution of the insects must be more
vigorous than in districts wliich are inhabited by comparatively few tailed species.
I 611 )
Now, with the exception of two species {kahneli from the Amazons, and pltalaecus
from Ecuador) all the tailed species of the vast gronp of American Aristolochia-
Swallowtails occur in the coantries from Costa Eica northwards, and in the
Brazilian fannistic snbregion (inclusive of Paraguay and Argentina). Is it at
all likely that in the Andesian and Amazonian provinces, from Bolivia to Costa
Rica and eastwards to the Guianas and Para, the tail is an unnecessary appendage
in these insects and has therefore been dropped, while it is a useful passive means of
defence in the other districts ? Considering further that these Aristolochia-feeders
are supposed to be practically immune, their nasty smell protecting them from
insect-feeders (perhaps not against inexperienced young individuals), it appears to us
certain that the presence of a tail in nearly all the Aristolochia-Papilios occurring
from Nicaragua northwards, as well as in almost every species found in the
Brazilian province, has nothing to do with a supposed usefulness of the tail as a
passive means of protection, and that accordingly the occasional absence of the
tail from Mexican specimens of phUenor is due to a physiological change of the
species in the southern districts of its range. P. philenor was originally doubtless
an inhabitant of the Atlantic district of the Nearctic Region ; the close morphological
connection between the Continental philenor, the Cuban devilliers and the Haitian
x:etes pointing decidedly in this direction. From this original home the species
spread southward and westward into Mexico, and at the Pacific coast again
northward as far as Northern California, Californian, Arizonan and Mexican
specimens differing on the whole slightly from eastern ones.
The scales of the upperside of the forewing are denticulate in P. philenor,
except towards the hinder angle ; on the hind wing, on the contrary, the scales are
entire, except in the region of the tail and anal angle. On the underside the scales
are dentate on both wings, the inner area of the forewing excepted. The white
spots attached to the orange spots of the hindwing are homologous to the white
spots of F. polydamas, which stand in the same connection with the red spots
in this species. The basal dot of the hindwing of P. philenor, on the underside,
is found in all the near allies of P. poli/damas, the corresponding dot of the
forewing being also present in P. polydamas. The scent-organ of the male and
the genitalia of both sexes are essentially as in P. polydamas, being only slightly
different.
The spring specimens of P. philenor appear to be on the whole smaller than
the later individuals, and bear always a row of spots on the upperside of the
forewing, this row being often absent from the males of the summer brood
(or broods). The hairiness of the body varies considerably, early Californian
specimens having quite a shaggy appearance.
Scent-organ : the scales are smaller than in P. polydamas, those of the streak
of small scales situated along the naked streak being moreover less triangular.
Genitalia: S. The lateral edges of the tenth tergitc are elevate in proximal
half, the distal half of the process appearing depressed in a lateral view. Harpe
with two processes as in P. polydamas, both short, pointed, the ventral one com-
pressed and broader than the dorsal process. ?. Hairy flaps in front of cavity
large, connected with one another at base, acuminate, each bearing on the distal
side a carina which extends on to the membranous proximal wall of the cavity.
For early stages, see literature under P. ph. philenor.
Hah. United States, except the central district from Colorado northwards .
Mexicp, '
( 512 )
a. P. philenor philenor L. (1'71).
PaiulU, Eqties Trojanus philenor Linn^, Maiit. Plant, p. 535 (1771) (America) ; Fabr., .Si/s/. Enl.
p. 445. n. 12 (1775) (America ; = aslenom) ; Goeze, Ent. Beijli: iii. 1. p. 30. n. 2 (177'J) ; Fabr.,
Spec. Ins. ii. p. 4. d. 15 (17S1) ; id., Miwt. Ins. ii. p. 2. d. 15 (1787) ; Jabl. & Herbst, Xalurs.
Srhmelt ii. p. 271. n. 52. 1. 19. fig. 2. 3 (1784) ; Esper, And. Sohinett. p. 49. n. 19. 1. 11. fig. 3 n7K5) ;
Panz., nnir,/s Ahhihl. p. 54. t. 11. fig. 1. 4 (178.5) ; Gmelin, Syst. Nat. i. 5. p. 2228. n. 282
(1790) ; Fabr., Eni. Sijst. iii. 1. p. G. n. 18 (1793).
PapiUo Eques Trojanus astinous Drury, lllu.'ilr. Exoi. his.l p. 21. t. 11. fig. 1.4. <? (1773) (New Y.>rk ;
Maryland ; Virginia) ; Cram., Pa/,. Exot. iii. p. 26. t.208. fig. A. B. (J (1779) (New York) ; Jung,
Alplmb. Verz. Sdimrll. i. p. 59 (1791) {= pluknor).
Papilio Eques Achiviis astinnous (!), StoU, in Cram., Pap. E.rot. iv., Ordre Si/sl. p. 3. note 2 (1782)
(= philenor).
Frinreps tlomi nans philenor, Hubner, Samml. Exot. SchmtU. i. t. 128 (1800— ?).
Laerlias philenor, id., Verz. bek. Schniett. p. 84. n. 858 (1818?) ; Scudd., S,/st. Rer. Amer. Butt. p. 43
(1872) • id , Geol. New Ilumpdi. i. p. 359. t. \. fig. 1.5. 17 (1874) ; id., Ihitt. E. U.S. & Canada ii.
p 1241 t 16. fig. 3, t. 2G. fig. 6, t. 3.-;. fig. 24. 25, t. 40. fig. 8, t. 43. fig. 19, t. 45. fig. 4. 5, t. 46
fig. 42, t. 56. fig. 8, t. 66. fig. 4. 8, t. 72. fig. 7, t. 76. fig. 13. 20. 21, t. 80. fig. 1-5, t. 85, fig. 14
(fsHg) ; id., Psijrhe viii. p. 207. t. 5. fig. 1 c. larva juv. (1898) ; Dyar, Bull. U.S. Nal. Mus.
Iii. p. 4. n. 23 (1902).
Papilio 2>hilenor, Jung, xilphah. Verz. Schmett. u. p. 102 (1792) (India !) ; Abbot & Smith, Lep. fns.
Georgia i. p. 5. t. 3 (1797) (1., p., (?,?); Say, Amcr. Entom. No. 1. plate i:i817) ; Godt, Em:
MHh. ix. p. 40. n. 47 (1819) ; Boisd. & Lee, Hist. Gin. Lip. Amir. Sept. p. 29. t. 11 (1833)
(1., p., S) ; Luca.s, Lip. E.zot. p. 15. t. 8. fig. 2 (1835) ; Boisd., Spec. Gin. Lip. i. p. 324. n. 167
(1836) ; Lucas, Vade-mecum p. 50 (1838); id., in Gue'r., Diet. Pill. Hist. Nal. vii. p. 48 (1838) ;
Drury, ed. Westw., ///«s/r. £j-o(. /».«. i. p. 20. t. 11. fig. 1. 4(1837) ; Harris, Eh^om. i. p. GO (1841)
(Massachusetts, 1. on Aristolochia sipho., August) ; Doubl., List Lep. Ins. Brit. Mus. i. p. 15
(1845) ; id., in Westw., Arc. Ent. i. p. 68 (1845) (habits) ; Lucas, Lip. Exot. p. 15. t. 8. fig. 2
(1845) ; Doubl., Westw. & Hew., Gen. Diurn. Lcp. i. p. 19. n. 230 (1846) (U.S.A. ; Slexico) ;
Kirtl., Proc. Ent. Soc. Land. (2). i. p. 101 (1851) (1. on Aristolochia) ; Gray, Cat. L/j,. Ins.
Brit. Mus. i. Pap. p. 66. n. 291 (1852) (California) ; Boisd., Ann. Soc. Ent. France p. 282. n. 4
(1852) (California); Gray, List Lep. Ins. Brit. J/«.s. i. Pap. p. 75. n. 308 (1856) (Ohio ; Florida;
California ; " Nicaragua ") ; Me'netr., Enum. Corp. Anim. .Mus. Pctrop., Lip. i. p. 6. n. 90 (1857)
(Mexico) ; Urban, Can. Nat. Geol. iii. p. 400. fig. a. b. (1858) ; Gosse, Letters from Alahama
pp. 77, 148, 272 (1859) ; Newm., Proc. Enl. Soc. Philad. 26 (1861) (N. Jersey ; on Srrpcntaria) ;
Morris, Syn. Lep. N. Amer. p. 6. n. 8 (1862) ; Reak., Proc. Ent. Soc. Philad. ii. p. 141. n. 13
(18i;3) ("Chiapas," error loci) ; Weidem., ibid. p. 147 (1863) ("Mexico, West Indies, Cent.
America") ; Kirkp., ibid. iii. p. 328 (1864) (Cleveland, Ohio, common) ; Jaeger, Life N. Amer.
Ins. p. 209 (1864) ; Feld., Verh. Zool. Bot. Ges. Wien xiv. p. 297. n. 113 (18G4) (U.S.A., Calif.,
Mexico) ; Behr, Sletl. Ent. Zeit. xxvii. p. 216 (1866) (Calif.) ; Pack., Guide Sludi/ Ins. p. 248.
fig. 181 (1868) ; Butl., Cat. Diurn. Lep. de.m: Fabric, p. 237. n. 13 (1869) (Ohio) ; Harris, ed
Flint, Ent. Corrcsp. p. 147, 273. fig. .37. 38 (1809) ; Beth., Camid. Nal. iii. p. 320 (1871) (habits);
Riley, 2nd Missouri Repl. p. 110. fig. 84. 85. 86 (1870) (life history) ; Kirby, Cat. Diurn. Lep.
p. 521. n. 20 (1871) ; Scudd., Canad. Ent. iv. p. 74. 0872) (Abbot's MS. in Brit. Mu-.) ; Edw.
Proc. Cal. Ac. Sc. v. p. 162 (1873) (Larva, pupa ; occurrence in Calif.) ; Butl. & Druce, Proc
Zool. Soc. Loud. p. 364. n. 367 (1874) (" Costa Rica," error) ; Edw., Proc. Calif. Ac. N. Sc. v
p. 102 (1875) (pupa) ; Ison, Kept. Ent. Soc. Ontario p. 15 (1870) (rare, Cleveland) ; H. W.
Edw., Trans. Amer. Ent. Soc. vi. p. 9. n. 3 (1877) (Atlantic to Pacific, Canada to Gulf of
Mexico) ; H.Edw., Proc. Cal. Ac. Sc. v. vii. p. 19 (1877) (pupa) ; French, Tran-i. Dept. Aijric.
Illin. XV. p. 136 (1877) (larva) ; Aaron, Canad. Enl. ix. p. 200 (1877) ; Grote, I.e. p. 220(1877)
(pollen on eye) ; White, I.e. x. p. 20 (1878) (pollen on eye) ; Street, Cal. Butt. Moths N. Amer.
p. 67. n. 1 (1878); Gerh., Macro-Lep. N. Amer. p. 25. n. 430 (1878); Dury, Cincinnati Soc. Nat
Uist. i. p. 12 (1878) (Cine, common) ; Beth., Canad. Ent. xi. p. 203 (1879) (Hamilton. Ont. ;
one year in abundance) ; Oberth., El. d'Enl. iv. p. 98. n. .304 (1880) (Mexico ; Texas ; Florida);
Saund., Rept. Ent. Soc. Ontario p. 37. fig. 16. 17. 18 (1880) ; Moffat, ibid. p. 10 (1881) (Long
Point and Ridgeway) ; Saund., ibid. p. 39. fig. 16. 17. 18 (1881) (i., 1., p.) ; Edw., Canad. Ent.
xiii. p. 9. (1881) (life history) ; Hagen, ibid. p. 37 (1881) (1. on A. sipho) ; Middl., Trans. Dept.
Agric. Illin. xvu\. Aj'pend. p. 73. fig. 0 (1881); Coquill., ibid, xviii. Append, p. 104 (1881)
(larva) ; Riley, Amer. Natural. ],. 327. fig. 1. 2. 3 (1881) (life hist.) ; Edw., Canad. Enl. xiv.
p. 21 (1882) (oviposition) ; Butl,, Journ. Linn, Sue. Land. xvi. p. 472. n. 58 (1883) (Mendocino
( 513 )
and Lake Co., June) ; Behr, Bull. C'al. Ac. Sc. i. p. 64 (1884) (Calif., common, 1. on Ar'.slo-
lochia); Edw., Oimid. Eiit. xvi. p. 109, 112 (1884) (egg, larvaj ; Gruber, ./»'*/;scAp Zeitschr.
Nalin-ir. xvii. p. 474. t. 7. fig. 20— 24 (1884); \A.,rapiUo iv. p. 88. t. 2. f. 20— 24 (1884)(transf.);
Lintn., ibkl. iv. p. 136. n. 1 (1884) (Rio Grande) ; Aaron, ;hkl. iv. p. 172 (1884) (S. Texas) ;
Mayn., Butt. N. Engl. p. 49. n. 67. t. 5. fig. 67. 67a. ^ (1886) ; French, Bull. E. O. States p. 88
(1886) ; Mayn., Butt. New Engl. p. 49. t. 5. fig. 67. 67.\ (1886) ; Royst. & Pigott, Jouni. Quel:
Club (■>). iii. p. 205 (1888) (scaling) ; Edw., Si/ii. N. Aiuer. Butt., in Butt. N. Amer. i. p. 1. n. 1
(1888) ; id., Bull. U.S. Nat. Mus. xxxv. p. 9 (1889) (liter, on metaraorph.) ; Skinn. & Aaron,
Canad. Ent. xxi. p. 126 (1889) (larva on Ipumoea !) ; Godm. & Salv., Biol. Centr. Amer., liliop.
ii. p. 204. n. 21. t. 65. fig. 18. 18a. fold and genit. (1890) (North Amer. ; Mexico) ; Mayn., Man.
N. Amer. Butt. p. 4. n. 2 (1891) ; Staley, Cciuacl. Eiil. xxiv. p. 204 (1892) (Marshall, Mifsouri,
common, iv. — x.) ; Kunze, ibid. xxv. p. 17 (1893) (Long I., I. eating I. for want of food) ;
Haase, Uutersuch. Mimicry \. p. 74(1893); Davis, Journ. N. Yorh Eut. Soc. I. p. 47(1893)
(Staten I., N.Y., May to Sept.) ; Skinn., E?,!. News iv. p. 82 (1893) (N. Carolina) ; Jones, ibid.
iv. p. 190 (1893) (Richmond Co., N.C.) ; HoU., Caiiad. Eut. xxv. p. 311 (1893) (Florida);
Beutenm., Bull. Amer. Mus. N. H. v. p. 242 (1893) (N. York ; descr. of 1., p., i.) ; Cockerel!,
Traus. Amer. Ent. Sue. xx. p. 353. n. 648 (1893) (Rosita, Colorado) ; Denton, E?il. News v.
p. 41 (1894) (Cambridge, Mass.) ; Moore, ibid. p. 77 (1894) (Bridgeport, Conn., larvae on
Arislol. serpentariti) ; White, ibid. v. p. 175 (1894) (Brooklyn) ; Soule, Psgrlir vii. p. 155(1894)
Nonqnitt, Mass., August) ; Weed, ibid. vii. p. 130. n. 39 (1894) (N.E. Miss.) ; Beth., Rej,t.
Ent. Soc. Ontario xxiv. p. 6 (1894) (Toronto & Hamilton) ; Holl., ibid. p. 53. fig. 27 (1894) ;
Osburn, Ent. News vi. p. 282. n. 48 (1805) (Tennessee, abundant, iv. to x., probably three
broods); Long]., ibid. vi. p. 314 (1895) (Chicago) ; Blatchl., Carnal. Ent. xxviii. p. 266 (1896)
(Indiana ; I. on Asarum !) ; Truman, Ent. News viii. p. 29 (1897) (Volga, S. Dakota, travel-
worn) ; Bubna, ibid. viii. p. 98 (1897) (Cleveland, Ohio ; plentiful on May 2nd and 3rd, a few
in July) ; Britton, ibid. ix. p. 173 (1898) (Newhaven, Ct., common) ; Duzee, Bull. Buffido Soc.
N.Sc. V. p. 107. n. 2. (1897) (Buffalo, very scarce) ; Christ, Mitt. Schweiz. Ent. (ies. ix. p 273
(1897) ; Moffat, Rept. Eut. Soc. Ontario xxvii. p. 79 (1897) (London, Ontario) ; Gibson, iljid.
p. 106. fig. cJ (1897) (Toronto, said to be seen 1) ; Moffat, I.e. p. 109. n. 85. (1897) (Ft. Hope,
August) ; Dent., Moths Butt. U.S. p. 354. t. 19 (1898—1900) ; id., Eut. News xi. p. .580 (1900)
(Wellesley) ; id., I.e. p. 643 (1900) (ex. with five red submarg. spots on hindwing) ; Beutenm.,
Butt. N. 'York City p. 8. n. 6. fig. (1902) ; Wasman, Ent. News xiii. p. 28 (1902) (aberr.,
corresp. to calrerleyi) ; Comst., ibid. xiii. p. 76 (1902) (L. Josephine, Fla.) ; Foster, ibid. xiii.
p. 326 (1902) (Claremont,N.H.); Hoag, ibid. xiv. p. 321 (1903) (S. Louis Potosi, Mex.) ; Baker,
Proc. U.S. Nat. .Mus. xxLx. p. 128 (1905).
Papilio asleuous (!), Doubleday, List Lep. Ins. Brit. .Mus. i. p. 15 (1845) (sub synon.).
Pachliopta {">) philenor, Reakirt, Proc. Ent. Soc. Philnd. iii. p. 504 (1865).
Pajnlio pidlenor var. acauda Oberthur, Et. d'Ent. iv. p. 98. sub n. 304 (1880) (hab ?).
Papilin jihilinor (!), Edwards, List Diurn. Lep., in BtM. N. Amer. ii. No. 2 (1884).
PapUio luzuhualcoyotl Strecker, Proc. Ac. Nat. Sci. Philad. xxxvii. p. 174 (1885) (Mexico) ; id.,
Lep. Rliop. Iltt , Suppl. iii. p. 17 (1900) (= acaudu = corbis = orsua).
Papilio corbis Godman & Salv., Ann. Mag. N. H. (6). iii. p. 357. n. 16 (1889) ; iid., Biid. Ce.utr.
Amer., Rhop. ii. p. 205. n. 22. t. 66. fig. 7. 8. (J (1890) (Valladolid, Yucatan).
Papilio pthilenor, Linn., var. obsoleta, (J, Ehrman, Canad. Ent. xx.xii. p. 348 (190(1) (S.W. Penn.).
Pai>ilio phileuor Linn., aberr, wasmuthi Weeks, Jouru. N. York Ent. Soc. ix. p. 82. t. 6 (1901)
(Brooklyn).
Ithobalusacaudt!, Dyar, Bull. U.S. Nat. Mus. Iii. p. 4. n. 24 (1902) (New Mexico ; Mexico: philenor
standing in another genus !).
Pterurns (!) phileuor, Kirby, in Hiibner, Samml. Exot. Schmetl. ed. ii. p. 100. 1. 129. fig. 1. 2 (190—?).
Papilio jihilenofT var. wasmuthi, Franck, Eut. News xv. p. 47 (1904) (Flatbush).
Papilio philenor wasmuthi. Skinner, ibid., Index p. 6 (1904).
Papilio (Laertias) philenor, Floersheim, Eut. Rec. xviii. p. 104 (1906) (protective taste of pupa).
(? ?. The iudividnals from the Atlantic States have generally a more elongate
hindwing than the majority of the southern and western specimens ; the marginal
spots of both wings are often smaller, the spots on the npperside of the forewing
are more often absent or vestigial in the males, and the distal margin of this wing
is rather more concave ; the metallic distal area of the underside of the hindwing is
less often green, and the body is rarely so shaggy as in some western specimens.
( 514 )
However, these distinctions are by no means reliaMe, the species being apparently
in a process of separating in an Atlantic and in a simtliern and Pacific form.
Two kinds of striking aberrations from the ordinary type are known :
a', ab. u-asmuthi Weeks, I.e. ? . Marginal sjjots of both wings very mnch
enlarged, forming large patches; on the underside of the hindwing these patches
are luero-cd together with the orange spots into a complete band, the veins remaining
very thinly black. One specimen in tlie Tring Mnsenm.
b'. ab. mex. acauda Obertli., I.e. ; P. nezahualeoyotl Strccker, I.e. ; /'. rorhis
Godm. & Salv., I.e. $. Tail of hindwing reduced to a tooth ; the metallic distal
area of the underside of the hindwing reduced in width (always ?), its inner edge
(and the white discal dots) being far separate from cell. This form is known only
from Mexico (Yacatan ; Guadalajara).
Hah. of F. ph. ■philenor : Atlantic States and Southern Canada, but in the
latter country and New England only a straggler, appearing sometimes to breed
where Arisfolochia sip/to is found (as a cultivated plant) ; its range being originally
less extended on account of the true food-plant {A. serpentaria) not occurring in
N. England and Canada ; Mexico, from Vera Cruz to the Pacific Coast ; Colorado ;
Arizona ; California.
In the Tring Mnsenm some larvae and pupae, and 193 c?c?, 95 ? ? , from :
Brooklyn ; Ealeigh, N. Carolina (Brimley) ; Nelson Co., W. Virginia (Wirt
Robinson) ; Sanford, Florida ; Makanda and Evanston, Illinois (Snyder) ; Nash-
ville, Tennessee (W. Osburn) ; Jefferson Co., Kentucky (Troxler) ; Monterey and
San Luis Potosi, Mexico; Jalapa, Vera Cruz, February 1894 (W. Schaus) ;
Iguala, Guerrero, 2500 ft., June 1904 (A. Hall) ; Guadalajara, July and October
189G (W. Schans) ; Verde E. and Nogales, Arizona (Oslar) ; Huachuca Mts.
and Phoenix, Arizona (Dr. Kunze) ; Benson, Arizona (0. T. Baron) ; Clarion I.,
December 1900 (Beck); San Lnis Obispo, California; McCloud P., Schasta,
and Siskiyou Co., California (0. T. Baron) ; Bntte Co., California, April 1898
(Mrs. Austin).
b. r. philenor oi'siia.
Papilio orsiia Godman & Salv., Ami. Mag. N. H. (6). iii. p. 358. n. 17 (1889) ; iid., Biol. Ceutr.
Amer., Rlop. ii. p. 205. n. 23. t. G6. fig. P. 10. S (ISHO) (Tres Marias Is.).
cJ?. A small form, with short rounded hindwing bearing a tooth instead of
a tail. Upperside of hindwing more strongly glossy than in P. ph. philenor,
ispecially in female. Glossy area of underside of hindwing touching cell, the
apex of which is also somewhat metallic.
llab. Tres Marias Islands.
47. Papilio devilliers Godt. (1824).
Papillon devilliers Godart, Mini. Soc. Linn. Paris ii. icy;, t. 1. fig. 3. 4, (J (18-2-2) (Cuba).
Pa2>ilio ilmlliers id., Eiic. MM. ix. Supi>l. p. 810. n. 47-8 (1824) (Cuba) ; Poey, Mem. Ii. Soc. Econ.
Ifuhnmi p. 235(1846).
Papilio rilliei-fi (!). Boihduval& Lee, Uist. Gen Up. Amir. Sept. p. 36. t. 14 (1833) (Florida; Cuba) ;
Boisd., Spec. Gi-n. Lq>. i. p. 325. n. 1G8 (1836) ; Doubl., Westw. & Hew., Gen. Diurn. Lep. i.
p. 19. n. 231 (1846) (Cuba : Florida) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 66. n. 292
(1852) C N. America ") ; id., List Lep. Im. Brit. Mus. i. Pap. p. 76. n. .^09 (1856) (" N. America ");
Lucas, in Sagra, llixt. Fis. Cuba vii. p. 207 (1857) ; Morris, Si/ii. Lep. N. Amer. p. 12. n. 17
(1862) ('■ Southern States ") ; Weidem., Proc. Enl. Soc Phihal ii. p. 148 (1863) (U. St. ? ; West
Indies) ; Feld., Verh. Zoo!. Bot. Ges. Wien xiv. p. 2'.I7. n. 112 (1864) (Cuba ; Florida) ; Kdw.,
Tn»is. Amer. Ent. Soc.vi.p. 9. n. 4(1877) (Florida ; Cuba); Oberth., Et. d'Ent. iv. p. 98. n. 305
( 513 )
(1880) (Cuba) ; Gundl., PiipHw i. p. 113 (1881) (Cuba) ; Edw., .Si/m. N. Amcr. Bull., in Buti.
N. Amer. i. p. 1. n. 2 (1888) (Florida ; Cuba) ; Haase, Vntfrsii.-h. Mimkryl p. 75 (180:!) (Cuba ;
Floiida).
Papilio demlliersi, Herrlch-Scb., Corr.-Bl. Zoul. Mhi. Ver. liegenxb. p. 173. n. 7 (18lU) (Cuba ; not
common) ; Kirby, Cat. Diurn. Lep. p. 520. n. 19(1871) ; Streck., Cal. Butt. Moths N. Amer.
p. 67. n. 2 (1878) (? Florida ; Cuba) ; Gerh., Macro-Lrp. N. Amer. p. 25. n. 435 (1878)
(Florida) ; Gundl., Cotitr. Ent. Cubaiia p. 123 (1881) (Cuba ; Florida).
S ?. Sexes similar. More specialised in pattern than in P. zete.'i, which comes
in some respects near the ancestral form of P. poli/damas, as do P. archidamas and
P. sfnv/icr/aniis. On the forewing there are some discal dots preserved, sitnated
beyond the apex of the cell, either on both sides (?) or at least on the underside
(c?), the spots being larger on the underside than on the upper. The pattern
of the underside of the hindwing is not unlike that of P. polijdamas ; the basal dot
and costal streak of P. poli/damas are in der/'lliers represented by a heavy basi-costal
streak ; behind C there is in most specimens a vestige of a creamy subbasal spot
and a silvery antemedian spot, which are homologous to the two creamy spots
C — SC^ found in some subspecies of P. polydamas ; sometimes there is a complete
row of silvery dots on the disc, but most specimens have only three or two or one
silvery spots behind the cell and usually one in the apex of the cell, these spots
being found again in P. philenor.
Scent-organ as in P. philenor.
Genitalia : S . Harpe triaiignlar, produced apically into a single j)rocess, which
bears some conical 'teeth. ? . Hairy flaps narrowed proximally as well as apically,
being ovate-lanceolate, sejiarate at base, each bearing a ridge on hinderside as in
P. philenor.
Early stages not described.
Ilab. Cuba.
In the Tring Museum 9 c?(? and 9 ? ? from noh|uin and Gibara (Tollin).
48. Papilio zetes Westw. (1847).
PainUo zetrs Westwood, Trans. Ent. Soc. Land. v. p. 36. t. 3. fig. 1. 1*. ? (1847) (Haiti) ; Doubl.,
Westw. & Hew., Gen. Diurn. Lep. ii. Append, p. 529 (1852) ; Gray, Cat. Lep. Ins. Brit. .Uim. i.
Pap. p. 66. n. 293 (1852) ; id., List Lep. Ins. Brit. Mu.<:. i. Pap. p. 76. n. 310 (1856) (Haiti) ;
Weidem., Proc. Ent. Soc. Philad. ii. p. 148 (1863) (We-t Indie?) ; Feld., Verh. Zool. Bot. Ges.
Wien xiv. p. 309. n. 283 (1864) ; Kirby, Cat. Diurn. Lrp. p. 543. n. 165 (1871) ; Haase,
Untersuih. Mimicnj i. p. 77 (1893) (near pliilenar).
S ? . Sexes similar. In shape resembling P. devilUers. A yellowish band
across u]>perside of both wings, somewhat as in P. polydamas ; some dots in and
beyond apex of cell of forewing, yellowish above, larger and paler on underside.
Hindwing with a broad l)and of white patclies on underside proximally of black
discal band, these i)atches being all contiguous with cell except the last ones ; the
white spots attached to the red ones large, the pattern of hindwing reminding one
of til at of/', streclierianus.
Hah. Haiti.
In coll. F. D. Godman and also in that of H. J. Adams.
The three following species of this group are very closely allied, the near
phyletic connection between P. polydamas., P. archidamas, and P. streckeriatii/s
being evident in every detail of pattern and structure. In fact, there would be some
reason for considering ail three as geographical forms of one species.
<S ? . Abdomen greenish black above in both sexes ; sternites not dotted.
Forewing below with a reddish or greenish yellow dot at base proximally of
( 516 )
praecostal spin-; SC" originating I'ruui roll more proxiuially than in P. lirhts and
P. madijcs.
Scent-organ : a streak of minute triangular scales along the naked streak ;
these scales not very close together, smaller than the scales situated along
nlidominal edge ; striation of scales heavy.
Genitalia : S. Harpe short, sinuate at apex, each angle ])rodnced into a curved
horn-like process, the upper process being absent from some forms of /'. poh/-
damas. ? . The two membranous lobes situated in front of the vaginal cavity
standing close together, forming one large lobe which is deeply cleft mesially.
49. Papilio streckerianus Hour. (1884).
Papilin stvcrherianus Honrath, Bci-l. Eiil. Ziitsvhr. x.xviii. p. 395. t. 8. fig. t. l.\. (J (1884)
(Guahango.o, Peru).
Papilio mathani Oberthur, El. (VEnl. xiv. p. 1. t. 2. fig. 8. J, 12.? (1891) (Chachapoyas).
S. A close relative of P. arr.hidamm. lilarkings of body pale greenish yellow,
not orange or reddish. Forewiug withont discal band above and below, but there
are in nearly every specimen vestiges of greenish streaks in the basal third of the
upperside, the streak along inner margin reaching sometimes as far as the greenish
spot situated in many sjiecimens at distal fourth ; the greenish yellow scaling is more
e.xtended on underside, covering here usually the greater half of the cell, often
extending anteriorly beyond the apex of the cell; in many specimens there are
s')me greenish yellow spots posteriorly on disc. The discal band of the hindwing,
on upperside, is variable in width ; it usually crosses the a])ex of the cell, but
in some specimens the cell-spot is just vestigial ; the marginal spots are always
large, being in some specimens twice as large as in others. The black discal baud
of the underside is always broad ; the silvery white submarginal spots SC^ — M' are
sharply defined ; there is sometimes a vestige of a red spot at the proximal side
of each white spot ; the red anal bar is always i)resent, while the red subcostal
bar is often replaced by white.
?. Much paler than the male, the discal band of the upperside of the hindwing
strongly oj)alescent, this opalescent gloss being vestigial also in the female of
P. arc/iidainas.
Genitalia as in P. archidamas.
Early stages not known.
Hal. Northern Peru, in the dry districts of the Maranon.
It is quite possible that in a more southern district of AVest Peru a form exists
which connects P. streckeriamis with P. archidamas.
In the Tring Museum 39 cJcJ from: Upper Maranon, east of Iluamachuco,
North Peru, dry country (0. T. Baron).
50. Papilio archidamas I'oisd. (183G).
(?) rapilio psillaciis tHoXwia, Sagg. Slor. Nal. Chili p. 211, 347 (1781).
Papillon bias Roger, Bull. Snc. Linn. Bordeaux i. p. 159 (182('>) (Chili).
Papilio archiihiuias Boisduval, Spec. Gen. Up. i. p. 321. r. 1(!3. (18.36) (Chili) ; Feisth., .1/";/. Zoi'l-
(2). i. Ins. Up. p. 1 (1839) (Chili); Doubl., Usl hep. Im. Bril. .l/«.t. i. p. 14 (1845) (Chili) ;
id., Westw. & Hew., Gen. Diurn. hep. i. p. 20. n. 242 (184(1) (Chili) ; Blanch., in Gay, Ilia. Fis.
Chile, y.ool. vii. p. 8. Alias t. 1. fig. la. b. (1852) ; Gray, Cat. Lep. Ins. Brit. Mns. i. Pa/i. p. GG.
11. 295 (18.52) (Chili) ; id., List Lep. Ins. Bril. Miis. i. Pa),, p. 70, n. 312 (185G) (Chili) ; Feld.,
Verh.Znnl But. Ges. Wien xiv. p. 297. n. IIG (1864) (Chili) ; Kit-by, Cut. Diurn. Lep. p. .121.
n. 25c (187!) ; Mathew, Ent. Mo. Mag. xiv. p. 152 (1877) (Val|>ariLiso, Oct. to Jan. ; habit?,
descr. of larval stages) ; Oberth., El. d'Enl. iv. p. 98. n. 302 (1880) (ChUi) ; Walk., Enl. Mo,
( •'51V )
Mad. xviii. p. 83 (1881) (Coquimbo, larva on Aristnlochia ; "common during our stay from
Jan. 21st to March lith") ; id., !.r. xx. p. 223 (1884) (Coquimbo, July and August, common,
fresh ; succession of broods all the year) ; id., l.r. xxi. p. 118 (1884) (Coquimbo, common,
March) ; Stand., E.:-ot.. Tu,jf. i. p. 12 (1884) (Chili) ; Haase, Untermch,. Minucr,/ i. p. 76 (1803).
Papllio bias, Kirby, Cat. Diitrn. Lep. p. 521. n. 2ob (1871); id., Trans. Roy. Soc. Dublin (2). ii. p. 324
{ISSO) (archidamas = bias) ; Butl. & Edm., Trans. F.nl. Soc. Land. p. 474. t. 21. fig. 1 (1881)
(larva) ; Elwes, ibid. p. 29,3. n. 53 (1003) (Santiago, "seen").
Pajiilio arcliemas (!), Mathew, Entomol. vii. p. (J2. n. 92 (1874) (Valparaiso, Nov., common, fast
flight).
Molina's description ajipe.ars to nppl}' to a certain extent to this species.
The French name " Papillon bias " of Roger is nomenclatorially not valid.
t? ?. The species varies a good deal in the amount of brown in the marginal
area of the forewing and on the disc of the hindwing, on the nnderside. The
yellowish marginal spots of the fore- and hiudwing are sometimes enlarged.
Genitalia : c?. Harpe with two jirocesses, the npper one smaller than the lower
one.
Early stages described by Mathew and again by Walker, 11. cc.
IM. Chili.
In the Tring Mnsenm 14 c?(?, 6 ¥ ? .
51. Papilio polydamas L. (1758).
Papilio Eqiies Trojanuspohjdamas hinni, Syst. N'al. ed. x. p. 4G0. n. 11 (1758) (citat. Merianae
excepta ; America).
One of the most interesting features of this species is the jieculiar distribution
of its geographical varieties. While the American continents, from Buenos Aires
to the southern Atlantic states of the United States, are inhabited by one single
subspecies, the West Indian islands appear to have each a special subspecies. On
the continents and the Greater Antilles the species is very common in open ground,
while it is decidedly rare on the Lesser Antilles, having perhaps become rare in
consequence of extensive cultivation of the soil. There are quite a number of
islands from which the species has not been recorded, though it doubtless e.xists there,
probably in special forms, for instance on Barbuda, Grenada, St. Christopher, etc.
In the position of the band of the hindwing the subspecies from Martinique is the
most difierent from the ordinary continental form ; the Sta. Lucia subspecies is
characterised by an exceptionally broad band on both wings ; the Guadeloupe form
is extreme in the reduction of the number of spots on the forewing, the Haiti and
Jamaica forms deviate from all the others in the absence of the njqjer process of the
harpe.
In all the subspecies the band of the upperside of fore- and hindwing is on the
whole rather wider in the female than in the male.
For literature on the early stages see /'. polyil. jioh/dainas.
llab. Southern Atlantic states southward to Buenos Aires ; West Indies.
ft. P. polydamas vincentius subsp. nov. (PI. VII. tig. 30).
cJ. Halfway between P. pohjd. lucianus and P. poli/d. polydamas.
Upperside. — Forewing : a band of spots as in P. polyd. polydamas, spots SC' — R-
a little farther away from margin. Hindwing : band curved, about five mm.
from cell, not essentially different from that of polyd. polydamas ; creamy white
marginal spots distinctly enlarged, resembling those of P. archidamas ; marginal
teeth broader than in polyd. polydamas.
( 518 )
Underside. — Forewing : apical area paler than basi-discal area, but uut so pale
brown as in P. pohjd. pohjdamas ; spots SC'^ — 11' small, while spots R- — SM- are
ranch larger than above. Hindwing almost uniformly brownish black, a grey
costal streak as in lucianiis, .renodamas, etc., and also a vestige of grey spot
proximally of the brick-red sjiot (_'— SC^ ; snlmiarginal spots as large as in
ar>iodamas,Y>{i]eT red, nearly as close to margin as in P. poli/d. poli/damas ; marginal
spots larger than in the other subspecies ; a thin grey bar M- — SM- proximally ot
red anal spot, preceded before M- by a thin obliipie grey bar.
Genitalia as in poh/d. poli/dnmas ; the carina which rnus from tlic tip of the
Tipper hook of the harpe proximad is oblicpie.
Ilab. St. Vincent, March 1897 (Dr. Percy Rendall).
One cJ in the Tring Musenm.
6. P. pohjdamas lucianns subsp. nov. (PI. VII. fig. 37).
Pdjiilin xenodamas, Sharpe (ho« Hiibner, 1822?, err. det.), Pmr. Znnl. Soc. Lniid. p. 22.'? (1901)
(Sta. Lucia) ; Bull., ibid. p. 713 (1901) (Sta. Lucia).
<? ? . Uppei-side.—FoTewiDg : a complete band of large spots from SC^ to inner
margin, spot SC— R' being the smallest, a streak in front of SCS somewhat
shadowy bnt always distinct, some yellowish scaling between SC'-^ and R' close to
cell ; the band much nearer the margin than in neodamas and xenodamas.
Hindwing : a distinct spot behind C ; band curved, about two mm. distant from
cell, patches SC- — M- about equal in size, or the second patch a little longer than
the others.
[';?r/(V«iV/t'.— Forewing : spots SC^— R' much smaller and the others rather
larger than above, no streak before SC^ or only a vestige of it. Hindwing : a
more or less distinct grey costal streak at base outside praecostal spur, and usually
a distinct grey spot behind C a little beyond middle ; brick-red snbraarginal spots
as in xenodamas, bnt much nearer the margin ; a distinct grey bar jiroximally of red
anal spot, the bar usually extended to M' ; creamy white marginal spots rather
larger than in xenodamas.
Genitalia as in P. polyd. pohjdamas.
Ilah. Santa Lucia.
In the Tring JIuscum 6 S 3.
In the British Musenm several pairs.
c. P. pwljdamas xenodamas Hubn. (1822?) (PL VII. fig. 3b).
hhobalus xenodamas Hiibner, Siimml. Exot. Schmetl. ii. t. 113. fig. 1. 2. ^ (1822 ?).
Papillon eurydamas, Roger, Bull. Soc. Linn. Bordeaux i. p. luS (1820) (Martiuique).
Papilin xenodamas, Boisduval, Spec. Gen. Lep. i. p. 320. n. ICl (183lj) (■' Brazil ") ; Doubl., AVe..tw.
& Ilew., Gen. Diurn. Lep. i. p. 20. n. 240 (184G) (•' Brazil "); Doubl., List Lep. Lis. Brit. .Urn. i.
Append, p. 3 (1848) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 67. n, 301 (18;j2) (" Brazil ") ;
id., List Lep. Ins. Brit. Mus. i. Pap. p. 77. n. 318(1850) (•' Brazil ") ; Feld., Verlu Zool. But. Ces.
Wim xiv. p. 297. n. 1 17 (18C4) (" Bras, austr.") ; Kirby, Cat. Diurn. Lep. p. 521. n. 23 (I87I)
("Brazil"); Oberth., Et. d'Enl. W. p. 97. n. 300(1880) (•' Brazil"); Staud., £:x«(. Ta/j/.t.
p. 12 (1884) ("Brazil").
Papilio cehrimes Dalman, .\nal. Entom. p. 38. n. 3 (1823) (hab. ?).
Fapilio eurydamas, Kirby, I.e. p. 521. n. 25 a (1871) (Martinico).
Itlwhalus xenndam'is, id, in Hubn,, Samml. Exot. Selimrlt. ed. ii. p. 91. t. 32G. fig. 3. 4 (190—?)
(" Brazil," false).
There are apparently only old specimens in collections. The insect inhabits
Martinique, not Brazil, llWhnar's xenodamas being the same as Roger's "Papillon
eurydamas." It is hardly possible to decide with certainty if xenodamas was
( 519 )
published after or before cebrmies. We accept, tlie view of all previous authors,
giving xe/Wf/amas priority.
t? ?. Upperside. — Forewing : band broad, upper three spots elongate, the third
the smallest, sometimes an additional spot before subcostal fork. Hiudwiug :
band broad, bluish, close to cell, spot M-— SM- large.
Underside uniformly black, forewing brownish, band creamy white, broader
than above, except upper three spots. Hiudwing deeper brown than forewing, a
subbasal costal streak and a large subbasal patch before cell bluish grey, recalling
the pale underside of P. streckerianus ; red snbmarginal spots twice as large and
twice as far away from margin as in average specimens of P. pohjd. polijdamas.
Genitalia : S . Harpe with two processes as in P. pohjd. polijdamas, both slender
and acutely pointed, the lower one bearing proximally a regular row of teeth.
Ilab. Martinique.
In the Tring Museum 1 S.
Two pairs in coll. Charles Oberthilr.
d. P. polijdamas dominicus subsp. nov. (PL VII. fig. 41).
Pupilio neudainas, Godman >i Salvia {non Lucas, 1852, err. (let.), Proc. Zuul. Soc. Loud. p. 318. u. 17
(1884) (Dominica).
c? ? . Intermediate between P. pohjd. neodamas and P. pobjd . xemdamas.
Upperside. — Forewing : spots sitnated as in xenodamas, standing nearer the
margin than in neodamas, the six spots from R- to hinder margin smaller than in
xenodamas, bnt rather larger than in neodamas, especially the uppermost one ;
three small dots SC — R- and a vestige of dot SC^— SC^ Hindwijig : band
distinctly curved, standing two or three mm. from cell, very slightly narrowing
backwards ; a vestigial spot C — SC- in male, a distinct spot in female.
Underside. — Forewing : creamy white spots intermediate in size betsveen those
oi xenodamas smCl neodamas, spot R- — R^ arrowhead-shaped like the others, spots
between costal margin and R- absent or vestigial. Hindwing slightly paler
brown from base to disc than in xenodamas ; a vestige of a grey subbasal streak,
no grey subbasal patch C— SC- ; red submarginal spot a little smaller than in
xenodamas and a little nearer the margin ; no grey bar proximally of red anal spot,
or only a vestige of it.
Genitalia not markedly different from those of P. pohjd. pohjdamas, but the
carina which runs from the upper hook proximad is parallel with the carina of the
lower hook, the two carinae being proximally connected with one another by a
transverse ridge which stands at right angles to them.
Hab. Dominica.
In the Tring Museum 2 c?c?, 3 ? 9 (E. A. Agar, and Elliott).
In the British Museum 2, S$,\ ? .
e. P. polijdamas neodamas Lucas (1852) (PI. VII. fig. 42).
Pupilio i>olydamas, Boisduval & Lee, Hi.tt. Gen. Lip. Amer. Sejit. t. 15. fig. 1. 2. (J (1833).
Papdio neudaniiis Lucas, Itco. Zuul. (2). iv. p. l'J3. 1. 10. tig. 5. (J(1852) (" Autilles "); Gray, Cut. Lei<.
Ins. Brit. Mils. i. Pap. p. 67. n. 300 (1852) ("Antilles") ; Lucas, in Sagra, Hist. Fis. Cuba vii.
p. 208. t. 14. fig. 3. 4. 1. & p. (1857) (Guadeloupe ; "Yucatan " error loci) ; Gray, List Lep.
Im.Bril. Mns. i. Pup. p. 77. n. 317 (1850) (" Antilles") ; Feld., Vcrh. Zuol. Bot. Ges. Wien xiv.
p. 297. n. 118(1804) (Guadeloupe ; " Cuba, Yucatan " error loci) ; Oberth., Et. d'Ent. iv. p. 97.
n. 301 (1880) (Guadeloupe).
Pupilio xenodamas var., Gueue'e, Mem. Soc. Phi/s. Hist, Nat. Geneve xxii. p. 370. u. 2 (1872) (hab. ''.).
S ? . Similar to P. pohjd. xenodamas, but the upper three spots of the forewing
( 520 )
absent (c?) or vestigial (?), the remaining spots fiirtlier away from margin.
Hindwing : discal band of itjipcrsiJe almost straight, no spot behind C, middle
patches smaller than anterior ones and nsnally also smaller than posterior ones, the
band therefore appearing narrowed in centre. On tindersidc no grey snbbasal
patch in front of cell.
Genitalia : S . Harpe as in xcnofhimas ; the carinae extending from the tip of
each process jiroximad almost parallel.
Ilab. Gnadeloupe.
In the Tring Museum 1 S,2 ? ?.
In coll. Charles Oberthiir 3 c?c?, 1 ?.
f. P. pohjdamas polydamas Linne (1758).
Seba, Thesaur. iv. p. 53. t. 44. fig. 14. 15 (1765).
Pajnlio Eques Trojaims jmlydanuis Linn^, Syst. Nat. ed. x. p. 460. n. 11 (1758) (citat. Merianae
excepta) ; id., Mtis. Ltid. Ulr. p. 192. n. 11 (17G4) (in India occidentali) ; Houtt., Naluii. Uist.
i. 11. p. 195. u. II (1767) ; Linne, Syst. Nat. ed. xii. p. 747. n. 12 (1767) ; Fabr., Si/xt. Ent.
p. 447. n. 22 (1775) ; Cram., Pap. Exnt. iii. p. 3.3. t. 21 1. fig. D. E (1779) (Surinam) ; Goeze, Ent.
Beyti: iii. 1. p. 34. n. 12 (1779) (cit. Sebae et Druryi excepta) ; Fabr., Spec. Ins. ii. p. 8. n. 29
(1781) ; Jabl. & Herbst, Nntiirs. Sehinelt. ii. p. 91. n. 26. t. 10. fig. 6. 7 (1784) ; Esper, Aitsl.
Schmell. p. 33. n. 12. t. 7. fig. 1. 2 (1785) (an fig. 1. ad banc subsp. pertin. ?) ; Fabr., .Mant. Ins.
ii. p. 4. n. 31 (1787) ; Gmel., Syst. Nat. i. 5. p. 2231. n. 12 (17901 ; Fabr., Ent. Syst. iii. 1. p. 14.
n. 42(1793).
Papilio Etpies piilydamas, Linii6, ed. Lange, Syst. Nat. p. 460. n. 11 (1760).
Papilio (Troes) polydamas, Miiller, Natiirs. v. 1. p. 569. n. 12 (1774).
Papilio (polydamas), Meerburgh, Afb. Zeldz. Gew. t. 2. 3. (1775).
Princejis duminans polydamas, Hiibner, Saniml. E.cot. Schmett. i. t. 130. fig. 1. 2 (1806— ).
Ithobaliis polydamas, JlahneT, Verz. beh. Schmett. p. 88. n. 913 (1818?); Scudd. , Pn"/. .Imc;-. Jc.
Arts <£• Sci. X. p. 198 (1875) ; Kirby, in Allen's Nat. Lihr., Lep. Butt. ii. p. 272 (1896); Dyar,
Bull. U.S. Nat. litis. Iii. p. 4. n. 25 (1902) (Florida ; "Antilles") ; Kirby, in Hiibner, Samml
Exot. Schmett. ed. ii. p. 91. t. 130. fig. 1. 2 (190—?).
Papilio 2)olydamas, Godart, Enc. ilith. ix. p. 39. n. 44(1819) (citat. Druryi et Sebae excepta) ;
Lacord., Ann. Sac. Ent. Fr. ii. p. 384 (1833) (Guyane) ; Boisd. & Lee, Hist. Gen. Lip. Amei:
Sept. p. 37 (1833) (sjnon. 2>artim ; t. 15. fig. 1. 2 ad subspeciem insulae Guadeloupe pertinet) ;
Lucas, L(p. Exot. p. 33. t. 17. fig. 2 (1835) ; Boisd., Spec. Gen. Lep. i. p. 321. n. 162 (1836)
(citat. " Boisd. & Leconte " fals.a) ; Doubl., List Lep. Ithi. Brit. Mus. i. p. 14 (1845) ; id., Westw.
& Hew., Gen. Diurn. Lep. i. p. 20. n. 241 (1846) (partim) ; Poey, Mem. R. Soc. Ec. Ilahana
p. 236 (1846) ; Erichs., in Schomb., F. F. Brit. Guiana p. 593 (1848) ; KoUar, Daikchr. K.
Ak. Wiss. Wien, Math. Naturw. CI. i. p. 354. n. 9 (1850) (Venezuela) ; Lucas, in Cbenu, Enc.
Hist. Nat., Pap. i. p. 38. t. 20. f. 2 (1851-53) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 67.
n. 299 (1852) (" var., Jamaica " alia subsp.) ; Wall., Trans. Ent. Soc. Lond. (2). ii. p. 2.55(1854)
(Amazons ; everywhere, common) ; Gray, List Lep. Ins. Brit. Mus. i. Pap. p. 77. n. 316 (1856)
(" var." excepta) ; Mcni'tr., Ennm. Corp. Anim. Mus. Petrop., Lep. i. p. 6. n, 95 (1S.')7) ; Lucas,
in Sagra, Hist. Cuba p. 486 (1857) ; Bates, Trans. Ent. Soc. Lond. (2). v. p. 228 (1861) (open
ground) ; id., Journ. Entom. i. p. 224. n. 6 (1862) (throughout the Amazons, in cultivated
places) ; Morris, Synops. Lep. N. Amer. i. p. 12. n. 18 (1862) (Georgia) ; Weidem., Proc. Ent.
Soc. Philad. ii. p. 148 (1863) {partim ; U.S.A., Centr. Amer.) ; Feld., Verh. Zool. Bot. Ges.
Wicn xiv. p. 297. n. 119 (1864) ("var." aliae subsp.) ; Herr.-Sch., Con: Bl. Hryrnsb. p. 174.
n. 14(1864) (Cuba, commonj ; Jaeger, Life N. Am. Ins. p. 210 (1864) ; Kirby, Cat. Diurn.
Lep. p. 521. n. 25 (1871) ; Butl. & Druce,' Proc. Zool. Soc. Lond. p. 364. n. 3i;7 (1874) (Costa
Rica); Cipronn., /l«)i. Soc. Ent. Belrj. xvii. p. 9. n. 11 (1874) (Petropolis, Nov., common) ;
Druce, Proc. Zool. Soc. Lond. p. 245. n. 6 (1876) (Upper Ucayali) ; Moschl., Verh. Zofil. Hot.
Ges. Wien xxvi. p. 295 (1876) (Surinam) ; Edw., Trans. Amer. Ent. Soc. vi. p. II. n. 22 (1877)
(Florida ; Cuba) ; Butl., Trans. Ent. Soc. Lond. p. 145. n. 226 (1877) (Obydos, January ; B.
Tapajos, March) ; Gerh., Marro-Lep. N. Amer. p. 25. n. 437 (1878) ; Streok., Butt. <D Moths
N. .1>«. p. Ii7. n. 3(1878); Dewitz, Arch. Natury. xliv. 1. p. 2. t. 1. fig. 1 (1878) (larva) ;
Hopff., Stett. Eut. Zeit. xj. p. 51. n. 12 (1879) (Brazil, Surinam, Venezuela, N. Granada, Peru.
Honduras, Mexico, Cuba) ; Burm., Descr. Rep. Argent, v. Li-p., Atlas p. C. n. 13. t. 2. fig. 7. 7a
( 521 )
(1879) (larva, pupa ; Buenos Aires) ; Oberth., Et. d'Eiit. iv. p. 98. n. 303 (1880) (Mexico ;
Guyane ; Para); Godm. & Salv., Trans. Eiit. Soc. Load. p. I'M. n. 244 (1880) (Sta. Marta) ;
Gosse, Eiil'im, xiii. p. 193 (1880) (Assuncion, Dec. to Marcli, not uncommon) ; Guudl., Pajjilin i.
p. 11.3 (1881) (Cuba) ; id., Enl. Cuhuua p. 121 (18.S1) {pm-tjm ; Cuba, larva, pupa); Edw.,
Pai>ilio ii. p. 122 (1882) (Florida) ; id., Gwml. Eiit. xiv. p. 120 (1882) (Florida) ; Walk., Eiit.
Mo. ilaij. xix. p. 2G (1882) (Panama, not r.are) ; Auriv., K. Sv. Vet. Ak. Handl. xix. b. p. 16.
n. 11 (1882) (recensio critica) ; Miiller, Kosnios xii. p. 448 (1883) (metam. : pupa brown or
green, no intergradations) ; Meldola, Proc. Ent. Soc. Lond. p. 23 (1883) (colour of pupae and
larvae); Jones, Proc. Lit. PhUox. Soc. Livcrp. p. 16. n. 77 (1883) (raetamorph.) ; Heczko,
Proc. Ent. Snc. Lond. p. 24 (1884) (drinking) ; Staud., E.rot. Tafif. i. p. 12. t. 8. ^ (1884) ;
Edw., Ball. U.S. Nat. Miis. xxxv. p. 13 (1880) (liter, of transf. ; partim) ; Godm. & Salv.,
Biol. Centr. Amer., Ehop. ii. p. 200. n. 15. t. 65. fig. 14. genit. (1890) (Mexico to Panama ;
'' S. Domingo, Jamaica, St. Thomas " aliae subsp. ) ; Sharpe, Proc. Zool. Soc. Lond. p. 555. n. 1
(1890) (Prov. of Goyaz) ; Hahnel, Iris iii. p. 203 (1890) (Valera) ; Maass. & Weym., in Stiibel,
ReKen S. Amer., Lep. p. 11. n. 37 (1890) (Colombia) ; iid.. I.e. p. 34. n. 29 (1890) (Pitol,
Colombia) ; Seitz, Stett. Ent. Zeit. Ii. p. 98 (1890) (Corcovado) ; Mayn., .1/on. iV. Amer. Bull.
p. 15. n. 22. fig. 9a (1891) (Cuba ; Mexico ; occasionally Florida) ; Haase, Unlersuch. iliniicry
p. 75 (1893) ; Michael, 7m vii. p. 214 (1894) (Sao Paulo de Oliven^a) ; Bonningh., Verh. Ver.
Nat. Unterli. Ilamhnrc] ix. p. 28 (1895) (Rio de Janeiro, very common) ; Weym., Slelt. Ent.
Zeit. Iv. p. 312. n. 1 (1895) (Rio Grande do Sul) ; Mabilde, (r'«;o pract. Borbolet. Rio Grande
do Sul p. 43. t. 1. fig. 2.\. n. c (1896) (larva, pupa, imago) ; Peters, llluslr. Zeitschr. Ent. ii.
p. 52 (1897) (Nova Friburgo, larva, pupa) ; Christ, .Uill. Schweis. Ent. Gen. ix. p. 273 (1897)
(Florida) ; Holland, Bull. Book p. 316. n. 21. t. 41. fig. 4 (1899) ; Denton, Moths Butt. U.S.
p. 355. fig. (1898-1900) ; Prinz. Therese, Berl. Ent. Zeitschr. xxxv. p. 240. n. 1 (1901) (Palmer,
W. Ecuador, 100 m., Aug. 19 ; R. Negro, Juli) ; Kaye, Trans. Ent. ,'oc. Lond. p. 206. n. 193
(1904) (Trinidad) ; Weeks, Illu.itr. Diurn. Lep. p. 20 & 28 (1905) (Bolivia).
c? ? . There is considerable individual variation in the size of the spots
cotniiosiug the band of the upperside of fore- and hiudwing. The range of this
subspecies is enormous ; in fact, P. pohjd. polydamo.s is the most widely distributed
American Papilio. The insect reminds one in this resjiect of the African P.
demodociis demodocus, which occnrs as such all over the African continent south
of the Sahara. Both species, which belong to widely different groups of Papilio,
have further in common that their distribution depends apparently to a great
extent on the ground being cleared of forest for cultivation, both species being
essentially inhabitants of open country.
The only ajiproach to a division of this subspecies into two geographical races
which we can find is the frequent reduction of the red submarginal spots on the
underside of the hiudwing in Cuban specimens, these spots being jaartly shaded
over with black.
Genitalia : Harpe bifurcate, the two processes of about the same size and
shape, pointed, curved, horn-shaped, the ventral one bearing usually some minute
teeth pro.ximally.
Ilah. Cuba ; Georgia southward to Buenos Aires.
In the Tring Museum several larvae and pupae, and 230 odd specimens from :
Cuba ; many places in Central and South America ; from East and West Mexico
southwards to Argentina.
In a crippled female from Barbados in the British Museum the red submarginal
spots on the underside of the hiudwing are rather larger than they are in average
specimens of _/'. poli/d. poli/damas. There may be a si)ecial form on Barbados ; but
more and better sj)ecimens are required for comparison with P. pohjd. polydamas.
(J. P. pohjdamas lucayus subs]), nov.
(? ? . Band of uppcr.nide liroad. Underside of hiudwing paler than in
P. pohjd. imhjdamas, yellowish white anal bar larger, usually extending forward
( 522 )
to M\ reacliiog at least beyond M- ; red submarginal spots of the same irregular
shape as in the coutiueutal form, but larger.
Ilab. Bahamas : Nassau aud New Providence.
Ill the British Museum (uame-type) and in coll. F. D. Godmau.
/(. r. poh/damas polycrates Hopff. (1806).
Papilio Equea Trojamis jjohjdamas var., Esper, Aual. Schmetl. p. ,33. n. 12. t. 7. lig. 1 (1784).
FapHio pill ij(hi mas. M^aetrios, Bull. Mvsruii ii. p. 293. n. 1 (1832) (Haiti ; common); Dewitz,S/?W. Eiil.
Zeit. xxxviii. p. 284. n. 3 (1877) (Porto Eico, common) ; Ilahnel, Ins iii. p. 134 (1890) (Porto
Rico) ; Moschl., Ahh. Seid-enh. Nat. Ges. xvi. p. 91. n. 3 (1891) (Porto Rico, common) ; Gundl.,
A71, Hist. Xut. iladrUl xx. p. 114. 11. 3 (1891) (Porto Rico ; syDOi\. parti in).
Papiliij poli/ilainas var.. Gray, List Lep. Ins. Brit. Mus. i. Pap. p. 77. sub n. 316 (1856) (/Mrtaii ; S.
Domingo) ; Feld., Verh. Zoo!. But, Gcs. Wieu xiv. p. 297. n. 119 (1804) (^partiiii ; S. Domingo).
Papilio polycrates Hopffer, Stetl. Eiit. Zrit. xxvii. p. 24. n. 4 (18i56) (" Par.-i '' error loci).
Papilio pohjdamas var. P. poh/crates, Kirby, Git. Diiirn. Lep. p. 521. sub n. 25 (1871) (" Antilles").
Papilio hi/pmlainas Guenoe, .Mem. Sor. Phijs. Hist. Nat. xxii. p. 371. n. 3 (1872) (Haiti) ; Kirby, Pet.
Noiiv. Enl. p. 239 (1872) ( = ^;«;^cr(i(es Hopff.) ; id., I.e. p. 809. n. 25 (1877) {= pvli/crates).
c? ? . Upper- and underside of both wings deeper in tint than iu P. pohjd.
polydamas. Band of forewing, npperside, a little more proximal, and that of
hiudwing less curved, than in P. pohjd. poli/dam.as, the spot M' — M- of this band
being more distad than in pohjdamas ; the anal sjjot standing proximall}' of the
anal marginal sinus is always small, often vestigial. iSize of spots of both wings
somewhat variable, the spots of hindwing being in some specimens only half as
broad as in others. Ked submarginal spots of hindwing below usually larger than
in P. pohjd. pob/damus, the silvery white spots attached to spots S(J- — R' large ;
black discal area mostly touching cell at R', much deeper black than iu P. polj/d.
polydamas \ spots of forewing paler than in P. polyd. polydamas, creamy white.
Genitalia : Harpe with one curved process only, the upper process being
absent; the process bears one or two teeth on the hinderside.
Hab. Haiti. Porto Rico (this form ?).
We have not seen specimens from Porto Rico.
In the Tring Museum 8 Jc?, 1 ?.
A series from Port au Prince in coll. (Jharles Oberthiir.
/. /'. polydamas jamaicensis subsji. nov. (PI. VII. hg. 40).
Sloaiie, Voy. Junmica ii. p. 216. t. 239. fig. 19. 20 (1725).
Papilio polydamas var., Doubleday, List Lep. Ins. Brit. .Mus. i. p. 14 (1845) (Jamaica) ; id., Westw.
& Hew., Oeii. Diurn. Lep. i. p. 20. n. 241 (184G) (jiartim ; Jamaica) ; Gray, Cat. Lep. Ins. Brit.
.Mus. i. Pap. p. 67. sub n. 299 (1852) ; id.. List Lep. Iiis. Brit. Mus. i. Pap. p. 77. sub n. 310
(1856) ; Butl., Cat. Diurn. Lep. deser. Fiihr. p. 237. n. 14 (1869) (Jamaica) ; id., Proc. Zool.
Soc. Lond. p. 481. n. 33 (1878) (Jamaica).
Papilio polydamas var. polyrrales, Cockerell (««« Hopffer, 1806, err. det.), Jamaica Instil, i. p. 27
(1891) (larva) ; Fox & Johns., Ent. News iv. p. 3 (1893) (Jamaica).
Papilio polycrates, Robinson, ibid, xiv. p. 18 (19U3) (Jamaica).
c? ?. Close to P. polyd. polycrates from Haiti, differing in the rufous brick-red
spots of the underside of the hindwing being larger and the white spots attached
to them smaller, the fourth red spot being usually without a vestige of white at
its hinder end, and the iifth and sixth spots mostly without any white scales
at both ends.
Ilfth. Jamaica.
In the Triug Museum 4 larvae, 1 yupa, 10 cJcJ, 0 ? ?.
( 523 )
j. P. pohjdamas thjamus snbsp. nov.
rapiVio Eques Trnjimits polydamas, Esper, Aud. Schmell. p. 33. n. 12. t. 6. fig. 2 (1784).
Papilii} pohjdamas var,, Gray, List Lep. Ins. Brit. Mus. i. Pap. p. 77. sub D. 31G (1856) (parlim ;
St. Thomas).
This form stands mnch nearer P. pohid. polydamas than do the subspecies from
Haiti and Jamaica, though geographically imhjd. thtjamus is farther away from
polyd. pol'/dama.i. Esper's figure, I.e., agrees better with this subspecies than
with any other we know.
(? ? . Upperside as in pohjd. polydamas ; distal margin of forewing a little
more strongly scalloped, tooth R' of hindwing rather prominent.
Undersiae, hiudwing : submarginal spots rufous brick-red, much paler and
much larger than in polyd. pohjdamas and not quite so close to the margin, these
spots even paler than in the Jamaica subspecies, and as irregular in shape as in
polyd. polydamas ; the white spots attached to second, third, and fourth spot not
quite so large as in the Haiti subspecies ; the black band situated at the proximal
side of the red s^iots ill-defined, much narrower than in the Jamaica and Haiti forms,
and mnch deeper black than in polyd. pohjdamas ; a long yellowish white costal
streak or (instead) dispersed yellowish white scales situated along costal margin
from praecostal spur to thrce-fonrths ; a heavy creamy white bar proximally of red
anal spot.
Genitalia as in pohjd. polydamas ; the two jirocesses of the harpe short and
stout.
Ilah. St. Thomas.
In the Tring Museum 2 c? <? (E. Hartert). A pair in the British Museum; a
male in coll. F. D. Godman.
k. Papilio polydamas antiquus subsp. nov.
PiipHio Eques Triijanus jmli/rJamas, Driny, Ilhistr. E.ciU. Ins. i. p. 32. t. 17. fig. 1. 2 (1770)
(Antigua).
Known to us only from Drury's figure, which comes near the form from
St. Thomas, but does not agree with it.
S . Upperside, forewing : a baud of well separated spots from SC* to inner
margin, the upper four spots small, the sixth, which is the largest, shorter than
its distance from margin. Hindwing : band narrower than in the St. Thomas
form, the black marginal area being one-third broader ; anal spot as large as in
t/iyamus.
Underside, forewing : three small dots SC^ — R-, the other spots larger than
above. Hindwing : snbmarginal spots rufous brick-red as in the St. Thomas
form, large, the white silvery spots attached to the second to fourth spots smaller
than in our St. Thomas specimens.
Hab. Antigua.
In the following two species the abdomen is yellowisli white above in the male,
the first segment and the claspors excepted, the scales being tongue-shaped, entire,
while in the female the u^iperside is blackish green, the scales being dentate. The
head, thorax, and underside of the abdomen are long-hairy, the sterna, coxae, and
femora being greenish yellow like the sides of the abdomen. The abdominal
sternites are dotted with white like the palpus, but these dots are often indistinct
on account of the hairiness of the body. The cell of the hindwing is rather shorter
34
( 524)
and more ronndeJ than in P. belus and allies. The underside of the hindwing is
greenish yellow from the base to the black discal band, the upper scales in this
basi-discal area being greenish yellow and the tinderscales blackish brown. The
claws of the hindtarsns are nearly symmetrical.
Scent-scales ochraceons yellow.
52. Papilio philetas Hew. (1869).
PopUio philetas Hewitson, Trains. Knt. Sue. LnwL p. 31 (ISCi'.lj (Ecuador) ; id., Exnt. Butt. iv. Pap.
t. 11. fig. 35. 36. c? (ISr.O) ; Kirby, Cat. Dliini. Lep. p. 522. n. 32 (1871) (Ecuador) ; Haenscb,
Berl. Eiit. Zeilschr. xlviii. p. 152 (11)03) (Baiios, R. Pastaza).
c? ? . Sexes similar, apart from abdomen. There is little variability in pattern ;
the spots in the costal region on the npperside of the forewing are sometimes
minnte. The species is easily recognised by the row of red spots on the disc of
the hindwing below.
Scent-organ : Scales of densely scaled streak three or fonr times as long as
broad, those situated between this streak and the abdominal edge being much less
elongate.
Genitalia: c?. Harpe broader than long, strongly but gradually narrowed
apicad, apex bifid, each angle being produced into a long curved liorulike tooth,
dorsal edge of harpe with some small teeth proximally. ? not dissected.
Early stages not known.
Had. Eastern Ecuador ; North Peru.
In the Tring Museum 22 (?c?, 1 ?, from: Loja; Zamora (0. T. Baron);
Baiios (R. Haensch).
In coll. Charles Oberthiir a series from Ambato, Ecuador, and from Chacha-
poyas, Peru.
lu coll. Paul Dognin both sexes from Loja. In the British Museum from
Nauta, Loreto, Peru.
53. Papilio madyes Doubl. (1846).
Papilio matli/es Doubleday, Ann. Mag. N. H. xviii. p. 375 (184(;) ( ? , Bolivia) ; Gray, Cat. Lep.
Ins. Brit. Mus. i. Pap. p. GG. d. 29G. t. 6. fig. 4. ? (1852) (fig. of type).
c? ? . Sexes similar, apart from abdomen ; female a little less metallic than
male. The red spots on the underside of tlie hindwing of P. philetas are in
P. 7natli/es replaced by greenish yellow ones, which are more or less completely
merged together with the submarginal greenish yellow patches. The scales of
the forewing, above, are more heavily denticulate than in P. philetas. There is
considerable geographical variation in this insect.
Scent-organ : Scales ovate, or broadly ovate, or asymmetrical, one side being
reduced.
Genitalia: S. Harpe triangular, the apex produced into an acute, curved,
hornlike tooth, a similar but much smaller tooth at the ventral [iroxi.ual corner,
occasionally accompanied by a minute tooth. ?. Vaginal lobes longer than
broad, separate, not carinate on hiuderside ; spines situated proximally on anal
segment heavy.
Early stages not known.
llab. Peru and Boliviii.
Five subspecies.
( 525 )
a. P. maffi/es plinius Weym. (1890).
Pajiilio j)li>iiiis Weymer, in Stiibel, RK/xeit S. Amer., Lep. p. 73. n. 14, p. 125. n. 42. t. 1. fig. 1. ^
(1890) (North Peru, between Tambo Almirante and Pueatambo).
S. Onl}' one s])eciraeii known ; ajiparently somewhat discoloured. Ujiperside
brigbter green than in the other subspecies ; forewiug with ouly four small white
spots from R- to SM^ ; spots of liiudwing about the same size as in the uext form,
but more sharply defined (?, according to figure).
U lu/ersicle, forewing: no indication of the distal spots SC — R-. Hindwing
ochre-yellow ; snbmarginal spots small, curved, the second to fourtli iy>-shaped.
Hub. North Peru : Chachapoj'as-Moyobamba district.
1^. P. madyes Morodamas Guen. (1872) (PI. VII., fig. 39).
PujiUio ililorndamas Guenfe, Mem. Soc. Phijs. Hist. Nat. Geneve xxii. p. 369. n. 1. fig. I. ^ (1872)
(Peru ; " $ ," errore).
Papilin madyas (!), Kirby, Prt. Noui:. Eiit. p. 239 (1872) (Morodamas — maili/as (!), errore).
Papilio mudyci, id., Cat. Dinrn. Lep. p. 809. n. 22 (1877) ; Druce, Proc. Zool. Soc. Lond. p. 24.5. n. 5
(1876) (Peru: Huiro, Santana) ; Hopff., Stetl. Ent. ZeH. xl. p. 51. n. 11 (1879) (^parlJm ;
Chanchamayo ; " chloruJamas is (J of madi/ex ; Chanchamayo specimens smaller than Bolivian
ones '■) ; Oberth., Et. d'Eiit. iv. p. 97. n. 299 (1880) (Peru).
Papilio madyes Tar. mursyas Staurlinger, Iris vii. p. 59 (1894) (Chanchamayo).
c??. Sjiots on upp)erside of fore- and hindwing large, those on forewing either
yellowish (most c?c?) or white (c?, ?).
Scent-scales ovate.
Iltd. Eastern Central Peru : Dejiartments of Huannco and Jnnin.
In the Tring Bluseum : 47 6S,2 ? ? , from : Huancabamba, Cerro de Pasco
(Bnttger) ; Pozuzo, ('iishi, and Chanchamayo (W. Hoffmanns); Rio Toro (Simons).
c. P. madijes crispus subsp. nov. (PI. VII. fig. 43).
Papilio madyes, Staudinger (iton Doubl., 1846, err. det.), Exot. 'Tayf. i. p. 12 (1884) (partim ;
S. Peru).
S. Upperside, forewing: spots much reduced, those at apex of cell vestigial ;
of the small distal spots SC^ — R- only the one in subcostal fork distinct, spots
R- — SM- less than half the size of the spots of chlorodamas. Hindwing :
submarginal spots very small as compared with those of chlorodamas.
Underside : apical area of forewing densely powdered with yellowish scales,
the yellow and the brown scales almost regularly alternating ; greenish yellow
snbmarginal spots on the whole somewhat smaller than in chlorodamas.
Hab. South-East Peru : Cnzco.
Resembles on the upperside the following subspecies, the spots of the forewing,
however, standing a little farther away from the margin.
In the Tring Museum : "i S S from Callanga, 1500 m. (Garlepp).
lu coll. F. Ducane Godmau a male from Santa Ana, Cuzco (Whitely).
(/. J', madijcs madijcs Doubl. (1846).
-/'"^;/7/o »iotZ)/cs Douhleday, ? f. (1346) (Bolivia) ; id.,Westw. & Hew., Geii. Diiirii. Le/i. i. p. 20.
n. 243 (1846) ; Doubl., List Lrp. Ins. Brit. Mus. i. Appeml. p. 4 (1848) ; Gray, Cat. Lep. Ins.
Brit. Mils. i. Pap. p. 66. n, 296. t. 6. fig. 4. ? (1852) (Bolivia) ; id.. List Lep. Ins. Brit. .Mus. i.
Pap. p. 76. n. 313 (1856) (Bolivia) ; Feld., Verh. Zool. Bot. Ges. Wien xiv. p. 297. n. 115(1864)
(Bolivia) ; Kirby, Cat. Diuru. Lep. p. 521. n. 22 (1871) (Bolivia) ; Hop£E., Stetl. Eid. Znt. xl.
p. 51. n. 11 (1879) (partim ; Moxos in Bolivia) ; Staud., Exot. Tayf. i. p. 12 (1884) (parlim ;
Bolivia) ; Haase, Unlersiieh. Mimicry i. p. 76 (1893) ; Staud., Iris vii. p. 59 (1894) (Uoc.ipata,
Bolivia) ; Weeks, IHustr. Dinrn. Lep. p. 20 (1905) (Chulumaui, Bolivia).
(526 )
li $. Veius ou vmierside of hindwing streaked with lilack.
IScent-organ : most of the scaJes asymmetrical, one side beiug reduced iu width.
Hab. Bolivia.
In the Tring Museum ~ SS from: R. Undnawe, 2000 m., February 1901
(Simous) ; Chuhiiuaui, :iiiOU m., January lOol, wet season (Simons) ; Ynugas de
La Paz, 1000 m., November 1899 (Garlepp).
In coll. F. U. Godman from Coroico, Bolivia, 050U ft. (Garlep(i).
e. P. madijes tucumanus subsji. nov.
c??. Ly^jivtfr.svV/c, forewiug : spots R' — SM'on'the whole a little larger and a
little more sharply defined than in P. m. madyes.
Underside, forewing : the band of spots sharply defined, forked, the apical
area not being suffused with pale primrose-yellow, the interspace between the
proximal and distal costal bands and the distal margin of the wing being brown,
the brown border anteriorly wider than the submarginal spots, gradually becoming
narrower jwsteriorly. Hindwing : basi-discal area smoky, appearing washed with
fawn-colour ; veins black as in P. m. madi/es ; black discal band wider than in the
Bolivian subspecies ; greenish yellow submarginal sjiots smaller.
Ilab. Tucuman, Argentina.
In the Tring Museum 3 <?(J, 1 ? , from : Tucuman (J. Steinbach).
A c? in coll. F. D. Godman from Bueyes, Bolivia (Garlepp).
A ? from Tucuman (Kinkelin coll.) in coll. Charles Oberthiir.
54. Papilio polystictus Butl. (1874).
Pa/tilio jmilodaytuis (?) Godart, Enc. MM. ix. p. 40. sub n. 4G. cJ(1810) (" pourrait bien utre female
de protodamas " ! ; Brazil).
Uhobalus prolodamas, Hilbner \nnn Godart 1819, err. det.), Samml. Ej:oL Schmett. ii. t. 115.
fig. 1. 2. S (1822 ?).
PapU'w polystidus Butler, Trans. Ent. Soc. Lond. p. 435 (1874).
Fiqiilia pnitudamas, Burmeister Descr. Rep. Argent, v. Lep., Atlas p. G. n. 12. t. 2. fig. 6. Oa (1879)
(larva, pupa).
Godart described P. protodamas from a female, adding the description of a
male Papilio, which, he says, might perhaps be the male of that female. As these
two sjiecimeus belong to different species, there can be no doubt whatever that the
name protodamas has to be applied to the species to which that female belongs.
Following the lead of Hiibner, who erroneously restricted the name protodamas to
the male of Godart and renamed the other species hyperion, all authors have
treated the present insect as beiug the true protodamas of Godart. The species
is geographically variable in the size of the markings on the upperside of the wings.
Butler, iu 1874, separated the small-spotted form as a distinct species, calling
it polystictus, from the large-spotted form, which he considered to be typical
protodamas. Though Butler was wrong in creating a new " species," his name
]wl>jsticti(s is nevertheless valid for the species, being the first name originally
given to individuals of the present species only. The so-called protodamas, namely
the large-spotted form, requires a new name, having remained without a valid name
of its own.
cj ? . Forewing with three to seven grej'-green submarginal spots on upperside,
and occasionally a streak behind SM'', spot M-— SM- double, all more or less
angle-shaped ; the corresponding spots of the underside large, greyish white, apex
( 527)
of cell and the disc beyond being sufifused with greyish white ; scales of npperside
entire, somewhat fan-shaped, being almost gradually widened, with the ajiex
ronuded as in P. laodamas, belus, etc. Hindwing : scales of npperside nearly
all entire, those in anal area being denticulate ; two rows of grey-green or greenish
yellow spots; the discal spots more or less ovate, often mere dots, sometimes
vestigial, the middle ones nsnally of almost even size, the first, second and last
being generally smaller than the others ; the snbmargiual spots angle-shaped, the
npper ones being reduced to dots in many specimens ; red submarginal spots of
underside resembling those of P. laodamas and j^olydamas ; no white dots distally
of them.
Scent-organ : scales twice or 2i times as long as broad, strongly striate, mostly
somewhat asymmetrical.
Genitalia of the same type as in P. belus.
Early stages described and figured by Burmeister, I.e.
Hab. Brazil ; Paraguay ; and Argentina.
The report of an occi;rrence of this species near Buenos Aires (Burmeister) is
confirmed by a female in coll. Charles Oberthlir collected by Kinkelin at Zarate,
Two subspecies :
a. P. 2'olijstictus janira subsp. nov.
Papilio protodamns (?) Godart, I.e. S (1819).
Ithobalus protodamas, Hilbner (iwn Godart, 1819, err. det.), I.e. (1822 ?) ; Kirby, ibid. ed. ii. p. 01.
t. 328. fig. 1. 2 (190—?).
Ithobafus (l) jjroliichmas, Charpentier, in Esper, Auxl. Schmett. i. Zimatze p. 13 (1831).
Papilio jirotodanias, Boisduval, Spec. Gcii. Lip. i. p. 322. n. llj-t (I83C) (Rio de Janeiro) ; Doubl.,
List Lep. /hk. Bril. Mun. i. p. 14 (1845) (Brazil) ; id., Westw. & Hew., &',■«. Diuni. Lep.i. p. 20.
n. 244 (1846) (Brazil) ; Gray, Cat. Lep. Ins. Brit. Mm. i. Pap. p. G7. n. 302 (1852) (Brazil);
id., List Lep. Ins. Brit. J/».5. i. Pap. p. 78. n. 319 (185G) (Rio de Janeiro ; " var." excl.) ;
Mi'netr., Enum. Corp. Anim. Mus. Petrop., Lep. i. p. 6. n. 96 (1857) (Brazil) ; Feld., Verh. Zool.
Bot. Ges. Wien xiv.p, 2S/7. n. 120 (1804) (Brazil ; " var." excl.) ; Capronn., Ann. .Soc. Ent. Bclrj.
xfii. p. 9. n. 10 (1874) (Gavia,Aug. ; Entre Rios, Sept. ; Botafogo, Nov.) ; Oberth., Et. d'Ent.
iv. p. 98. n. 306 (1880) (Brazil) ; Bonningh., Vcrli. Ver. Not. Unlerh. Ilambnr;/ ix. p. 28 (1895)
(Petropolis, common).
c? ? . Submarginal spots of npper- and nnderside of forewing and discal ones
of npperside of hindwing larger than in the following form.
lu some individuals the discal spots of the hindwing are so large that the middle
ones extend close to cell. Most males have seven distinct submarginal spots on the
npperside of the forewiug, while in the females the upper two or three spots are
nsnally missing.
JIab. Province of Rio de Janeiro ; Minas Geraes.
In the Tring Museum 25 J c?, 10 ? ? , from : Minas Geraes (R. Haensch); Nova
Fribnrgo; Petropolis, November 1897, January 1898 (Foetterle) ; Rio de Janeiro.
b. P. pohjstidus polijstictus Butl. (1874).
Papilio protodamas var. b.. Gray, List Lep. Ins. Bril. Mns. i. Pap. p. 78. sub n. 319 (1856).
Papilio polystii:tus Butler, Trans. Ent. Sue. Loud. p. 435 (1874) (Rio Grande ; Espirito Santo) ;
Kirby, Cat. Diiirn. Lep. p. 814 (1877) ; Weym., Stell. Ent. Zeit. Iv. p. 312. n. 2 (1895) (Rio
Grande do Sul).
Papilio 2irolodainas,Baimei!iteT, De.ier. Rep. Argent, v. Lep., Atlas p. G. n. 12 (1879) {partim ; (Buenos
Aires) ; Jones, Proc. Lit. Pliilos. Soc. L'lrerp. p. 17. n.'_83 (1883) (larva, pupa ; forest) ; Haase,
Untersuch. Mimicry i. p. 76 (1893) (South Brazil).
Papilio neodamas, Mabilde (non Lucas, 1852, err. det.), Gnia pract, Borbol. Rio Grande do Sul p. 43
(1896).
( 528 )
(S ? . Spots on npperside of wings reduced in site, especially the disoal series
of hinJwinf^ ; the njiper two or three sjiots of the forewiug vestigial or missing in
most individuals. Specimens with exceptionally large spots scarcely distinguishable
from Ja/iira specimens with exceptionally small markings.
IJab. Brazil : Sao Paulo southward to Rio Grande do Sul ; Paraguay ; Bnonos
Aires.
In the Tring Museum 1 pupa, 21 cJcT, 11 ? ?, from: Castro, Parami (E. D.
Jones) ; S. Cutharina ; Rio Grande do Sul ; Yhu, Paraguay, December 1890
(Andeer) ; Sapucay, Paraguay, October to February, June (W. Foster).
55. Papilio eracon Godm. & Salv. (IS'JT).
Papilio eracon Godman & Salvin, Trans. Ent. Soc. Lonil. p. 248 (1897) (Colima, Mexico) ; iid., Biol.
Ceiitr. Amer., Rlmp. ii. p. 729. n. 18 (n). t. 111. fig. 11. 12. ^ (1901) (Colim.i).
(??. Wings, upperside: forewiug with a submarginal row of four to seven
spots from hindmargin forward, the spots gradually decreasing in size. Hind-
wing : an evenly curved row of spots placed (in centre) halfway between cell and
distal margin, the upper two spots the smallest, the others of about eipial size,
the band broader in female than in male ; a row of admarginal spots, being the
remnants of transverse bars.
On underside a row of heavy transverse red bars on hindwing, the u]>per bars
bearing at costal side a silvery spot, at the distal side of which there is a rather large
creamy admarginal dot, in cellule R^ — M' two silvery dots and two creamy ones,
in cellule M' — M^ the silvery and the creamy dot standing at posterior end of red
bar — i.e. at vein M'^, not at M'.
Scent-organ : a streak of dense erect scaling as in the allied species, the .scales
twice as long as broad or less, multistriate, nearly as broad at base as at apex.
Genitalia as in P. belus, tenth tergite a little slenderer.
Early stages not known.
Hab. AVest Mexico : Colima and Guerrero.
In the Tring Museum 2 <?<? from Guerrero (0. T. Baron).
In the British Museum a pair from " Mexico " (coll. Crowley).
In coll. Adams 3 cJ c? without special locality.
56. Papilio belus Cram. (1777).
Papilio Eques Acltimis beltis Cramer, Pap. Exot. ii. p. 23. t. 112. fi^. A. B. (^ (1777) (Surinam).
Papilio Eqiiis Achiviis numitor id., I.e. p. 25. t. 113. fig. B(1777) (Surinam).
Papilio Eqiies Trojanus anndius Esper, .4ms/. Schmetl. p. 113. D. 48. t. 27. fig. 1. $ (1792) ; Martyn,
Psi/i-he t. 1. fig. 1. t. 2. fig. 1 (1797) (iiied.).
Papilio varus Kollar, Deiiksclir. K. Aeail. Wiss. TIVch, ilaOi.Xat. CI. i. p. 354. u. 8. t. 42. fig. 5. G. ?
(1850) (Cundinamarca, Colombia).
We have not seen a si)ecimen agreeing with Esjier's or Martyu's figures of
amulius, in which the submarginal spots on the underside of the hindwing are
yellow instead of red. We believe this insect to le a xantliochromati(^ female of
belus. Martyn's figure is better than that of Esper. Tiie yellow spots on the
up|ierside of the hindwing as shown in Martyn's figure are rather often limiid in
females of belus, but are not so pure yellow.
<?. Scaling of body and wings practically as in /'. li/c/das. No white streak
along abdominal fold of hindwing, red sjiots on underside farther away from
margin than in J', ii/cidas, accomjiauicd by white admarginal dots, which are
( 529 )
rarely vestigial ; tlie number of white discal patches ou npperside variable from
one to seven, the first always large, often occupying half the cellnle C — S(;-, but
never extending to base as in P. crassus, the others smaller and usually more
greenish, very variable in size, the scales composing them lying on top of the
metallic scales (as in P. lycidas), less acuminate than in /-". lycidas ; SC'^ ou the
whole branching off from cell a little more proximally than in P. lycidas.
?. Dichromatic; the ordinary form similar to the male except in the abdomen
being metallic green above and the forewing bearing often some submarginal spots
on uj)perside ; the rarer form bearing a large huffish yellow patch on forewing
occupying apex of cell and the adjacent portions of the disc.
Scent-organ : scales geographically variable, very close together, erect, either
about four times as long as broad, slightly curved, obtuse at apex and faintly or not
at all denticulate, somewhat cucumber-shaped, but compressed, though not ijuite
flat as ordinary scales ; or even shorter than in P. lycidas ; or of intermediate size.
Genitalia : 3 • Harpe small, short, triangular, with the apex strongly rounded ;
ventral margin dilated into an acuminate lobe which stands vertically on the plane
of the harpe ; tliis lobe cnrved basail, denticulate at distal edge. ?. Hairy lobes
of vaginal cavity larger than in P. lycidas and the postvagiiial tubercle higher.
Early stages not known.
IJab. Mexico to Bolivia, eastwards to Parii ; not in Brazil ])roper, but may be
expected to occur in the province of Goyaz.
Five subspecies.
If the figure of P. niimitor Cram., I.e., does not represent a specimen of
P. lycidas, it represents a specimen of P. belus belus with spots on the disc of the
hindwing. There is no white streak on the hindwing of the figure, as there is
in P. lycidas, but the artist may have forgotten to put that streak in.
a. P. belus chalccus subspec. nov.
c?. Wings, npperside: hindwing deeper green at distal margin than on disc;
within this marginal area a row of angle-shaped submarginal spots of the same
green colour as the disc ; discal row of creamy white spots similar in position to
that of the Bolivian form, P. belus cochabamba, the row being straight or only
faintly curved from SC- to abdominal margin, spots from U' backwards small, spot
R' — R- farther away from cell than spot R- — R'.
Underside, forewing: white fringe-spots as distinct as above. Hindwing:
red submarginal bar C — SO- sharply marked (absent from the other forms or
vestigial) ; white admarginal dots very distinct.
? not known.
Scent-organ : scales very short, being partly broader than long, cuj)-shaped,
j)roximally strongly rounded, striatiun heavy, the apical edge appearing multidentate.
Ilab. Guerrero, West Mexico (0. T. Baron).
Four (?(J.
b. P. belus varus Koll. (185U).
VupiUn varus KoUar, Denksriu: K. .ilc. Whs. ^Yifn, Math. Nat. CI. i. p. .354. n 8. t. 42. fig. 3. 4. ?
(1850) (Cundinamarca, Colombia) ; Doubl., Westw. &'Hew., Gen. Ditirn. Lrp. ii. p. ,12'.) (1852) i
Gray, Cat. hep. Ina. Brit. .Mux. i. Pap. p. G8. n. 308 (1852) ; id., List Up. Jus. Brit. Mus. i. Pai>.
p 79.11.325(1856) ; Feld., Verh. Zool. Bot. Ges. Wien xiv. p. 297. n. 124 (WCA) {parti m
N. Granada).
Papilio numitor, Gray (non Cramer, 1777, err. det.), Cat. hep. Ins. Brit. ifus. i. Pap. p. 68. n. 305
(1852) (Venezuela) | id,, List Lcp. Ins. Brit. Mas. i.i'a/).p.78.n.322(185C) (Venezuela) ; Bates
( 530 )
Ent Mil. m,,). i. p. 2 (18G4) (Guatemala) ; Godm. & Salv., Biul. Ccntr. Amer., Ehop. ii. p. 202.
n. 17. t. GG. fig. 1. ?, 2. 3. £? (180n) (Guatemala ; Nicaragua ; Costa Rica) ; iid., I.e. p. 728
(1901) (Honduras).
PapiUn laliimx Felder, Wien. Ent. Mon.y. p.72n. 2(1861) (MiSrida, ? ;— Mus. Tring) ; id., Verh. Zoo!.
Bol. Ges. iVit'ii xiv. p. 298. n. 126 (1864) (Venezuela ; Bogotd) ; id., licise Novara, Lep. p. 39.
n. 28. t. 10. fig. b. cJ (18G.5) (Bogota ; M^ridi) ; Kirby, Cut. Dhmi. Lep. p. 522. n. 29 (1871)
(Amer. Centr. ; N. Granada) ; Oberth., Et. d'Eid. iv. p. 98. n. ?m (1880) (Panama ; Colombia ;
San Esti'ban) ; Godm. & Salv., Trans. Ent. Sa,: Loud. p. 12G. n. 243 (1880) (Sta. Marta) ;
Hahnel, his iii. p. 203 (1S90) (Valera) : Maass. & Weym., in Stiibel, lieism S. Amer., Lep. p. 1 1.
n. 3K, and p. 31. n. 124 (1890) (Colombia) ; Haensch, Berl. Ent. Zeitschr. xlviii. p. 154 (1903)
(Arcbidona, E. Ecuador, G40 m.).
c?. Hindwiiip, upperside : distal margin of the same colour as the disc, or,
if darker, not bearing distinct paler green snbmarginal angle-shaped sjiots ; au
evenly carved row of discal spots, variable in size, but always gradually diminishing
in size from spot SC^ — R' backwards, the number being usually four or five,
seldom seven, rather often two, inclusive of the large costal patch. Six red
subraarginal spots on underside, there being no spot between C and SC- or only
a vestige of it.
? . Dimorphic.
a. ?-f. latinus Feld., I.e. Forewing with three or four submargiual spots,
enlarged on underside, where the upper ones are usually elongate arrowhead-
shaped. Hindwing : distal edge darker than disc, this border pro.ximally creuate,
limited by some creamy scales forming more or less distinct dots or curved bars ;
an evenly curved band of five or six spots, gradually diminishing in size, the
first and second of about the same size, the last minute.
h'. ?-f. varus Kollar, I.e. Forewing with large yellowish patch occupying
ajjex of cell and adjacent portions of disc.
Scent-organ : scales elongate, gradually widened, the larger proportion of
them about three or four times as long as broad, somewhat curved, not or very
feebly denticulate, the scales from the distal part of the fold being especially long.
Hub. Guatemala southward to North-East Ecuador and North Venezuela.
In the Tring Museum 27 c?c?, 5 j ?, from: San Pedro Sula, Honduras;
Cbiri(|ui ; "Bogota"; Purnio, II. Magdalena, November 1890 (Dr. Burger);
Archidona (Goodfellow) ; Merida and Tachira (Briceno) ; M6rida (type of latiiiii.s,
ex coll. Felder).
A specimen of ¥-f. vartt.'i in Brit. Mus. from " Bogota " (Crowley betpiest).
c. P. bclus helus Cram. (1777).
Pap'dio Eques Achlmis beliis Cramer, I.e. (1777) (Surinam) ; Goeze, Ent. Bcijlr. iii. 1. p. 8;). n. 67
(1779).
PapiUo Eques Acliirus numitor id., I.e. {1111) (Surinam) ; Goeze, I.e. p. 82. n. 45 (1777).
Piipilin Eques Trnjmms Mus, Fabricius, Spec. his. ii. p. 9. n. 34 (1781) {parti'm) ; id.. .Vuut. Jns.
ii. p. 5. D. 30 (1787) (parthn) ; Jabl. & Heibst, Nutnrs. SchmHt. ii. p. 95. n. 27. t. 11. fig. 1. ^
{n 8i)(_ parti m); GmeVm, Syst. Xal. i. 5. p. 2233. n. 294 (1790) (partim); Fa.hr., Ent. Syst.ui. 1.
p. 17. n. 53 (1793) (partim).
Papitio Eques Tnijauus numitor Esper, Au.il. Scliuiett. p. 81. sub n. 35, p. 114. n. 49. t. 27. fig. 2
(1792).
Pajiilio Eques Tmjunus amidius id., I.e. p. 113. n. 48. t. 27. fig. 1. ? (1792).
Jthulndus beliis, Hiibner, Ve.rz. hek. Schmelt. p. 88. n. 915 (1818?).
Jthobcilus numilur, id.. I.e. p. 88. n. 918 (1818?).
Papilio helus, Godart, Euc. Math. ix. p. 38. n. 42 (1819) (partim) ; Lacord., A?iu. Soe. Ent. Fr. ii.
p. 384 (1833) (Godart's Ijelus = brlus, emssiis and rri/mauthus) ■ Boisd., Sjiee. G<!n. L/p. i.
p. 315. n. 154 (183G) (Surinam) ; Doubl., Westw. & Hew., Geu. Diuru. Lep. i. p. 20. n. 233
(1846) (Surinam) ; Gray, Cat. Lep. Ins. Brit. J/us. i. Pap. p. 68. n. 303 (1852) (Ega ; cit, Lucas
( 531 )
excl.) ; Wall., Trans. Ent. Soc. Lonl. (2). ii. p. 255 (1854) (Amazons) ; Gray, List Lep. Ins.
Brit. Miis. i. Pap. 78. n. .320 (I85G) (Ega) ; Mt-netr., Eman. Corp. Anim. Miw. Petrnp., Lep. i.
p. G. n. 92 (1857) (" Brazil"); Bates, Trans. Ent. Soc. Lond. (2). v. p. 227 (1861); id., Journ.
Entom. i. p. 223. n. 2 (18G2) (Upp. Amazons; Guiana) ; Feld., Verli. ZooL Bnt. Ges. Wien xiv.
p. 298. n. 128 (18G4) (Surinam ; Upp. Amazons) ; But!., Cat. Diurn. Lep. desr.r. Fabric, p. 2,38.
n. 15 (18G'.I) (Peru) ; Kirby, Cat. Diurn. T^ep.\i. .522. n. 30 (1871) (Surinam ; Upper Amazons) ;
Druce, Proc. Znol. Soc. Lond. p. 245. n. 8 (187G) (Peru) ; Butl., Trani>. Ent. Sue. Lund. p. 145.
n. 227(1877) (R. Jutahi,Feb.) ; B.opS.,Stctt. Ent.Zeit. xl. p. 51. n. 13(1879) (Brazil, Surinam,
Peru) ; Obertb., Et. d'Ent. iv. p. 98. n. 309 (1880) (Guyane ; Toltc) ; Muschl., Verh. Zool.
But. Ges. Wiin xxxii. p. 3U3 (1883) (hindwing with macular band, Surinam) ; Stand., Ej-ot.
Ta;if. i. p. 12 (18S4)( ? dimorphic ; Amazons) ; Haase, Untersuch. Mimicry i. p. 7G (1893)(jJ ? );
Michael, Iris vii. p. 213 (1894) (Sao Paulo de 01iven9a).
Papilio niimitor, Boisduval, Spec. Gen. Lip. i. p. 317. n. 157 (183G) ; Bates, Journ. Entom. i. p. 223.
n. 4 (18G2) (partim ; Guiana ; " Venezuela " alia subsp.) ; Kirby, Cat. Diurn. Lep. p, 522. n. 31
(1871) {jiartim ; "Guatemala" alia subsp.).
9. PapiUo varus, Bates, Trans. Ent. Soc. Lond. (2). v. p. 228 (18G1) ( ? of Indus ; Ega).
?. Papilio caburi Kaye, Enlom. xxxix. p. 51. t. 2. fig. 1 (190G) (Brit. Guiana).
? . Hiudwiog, upperside. There is usually only the costal patch present ;
many specimens have a' small ill-defined patch behind the costal one, while very
few specimens have a row of small sjjots across the disc (jiumitor').
? . Dimorphic as in the preceding form.
a. ?-f. belus Cram., I.e. ; caburi Kaye, I.e. (19(.i0).— Similar to the male.
Hindwing, upperside, with a costal patch as in male, bnt nsually smaller, followed
either by one single spot or by a row of spots extending straight across disc, the
row being not or little curved, the middle spots standing close to cell and being
the smallest of all ; the spots more or less yellowish ; distal marginal area more
or less blackish green, with brighter green halfnioous ; red submarginal sjiot
C— SC^ of underside present ; in ab. amuliiis the snbmarginal spots of the underside
of the hindwing yellow according to Esper and Martyn, ll.cc.
l). ?-f. ammonis nov. Like varus ?-f. varus; the yellow spots beyond
apex of cell smaller ; the costal patch of the hindwing larger, the dark green distal
border less well defined and the bine-green angle-shaped markings within it larger.
Scent-organ : scales much shorter than in P. belus varus, but longer than in
chalceus and cocliabamha.
Hah. Guiana ; Amazons ; Peru.
In the Tring Museum 25 Si, 5 ¥ ?, from : Surinam; R. Negro; Iquitos,
R. Ucayali, and R. Cachyaco (Stuart) ; R. Chuchuras and Chanchamayo
(Hoffmanns) ; R. Perene (Simons) ; type of ¥-f. amazonis from Iquitos.
In coll. Godman a S from (Jhapada, Brazil (H. H. Smith).
In coll. Oberthiir a ? of ?-f. amazonis from Massanary ; three specimens of
^ -belus from Cayenne, two of which have a complete band of spots across
hindwing, a band of smaller spots being present in the third.
In coll. F. I). Godman two specimens of ?-f. amazonis from Ega and
Massanary.
d. P. belus belemus Bates (1864).
Pai>iUo numitor, BKtes (non Cramer, 1777, err. det.), Trans. Ent. Soc. Lond. (2). v. p. 228 (ISGl)
(Pard.) ; id., Journ. Entom. i. p. 223. n. 4 (18G2) (jparlim ; Pard ; local form of belus).
(J. PapUio belemus id., Ent. Mo. Mag. i. p. 2. note (1864) (Pard ; —coll. Godman); Kirby, Cat.
Diurn. Lep. p. 522. n. 31a (1871) ; Hahnel, Iris ui. p. 212 (1890) (Pari,).
(J?. Hindwing with straight band of spots across disc close to cell, nearly as
in cochabamba ; green submarginal half-crescents distinct.
Ilab. Para, district, a])parently only on the south side of the Amazon,
( 532 )
e. P. belus cochahamba Weeks (1901).
$. Piijiilio cochahamha Weeks, Canail. Ent. xxxiii. p. 2G5 (1901) (Bolivia, some 200 miles north of
Cochabamba) ; id., A«(. Xews t. 5 (1902) ; id , //te(r. Diurii. Ltp. p. 39. t. 7. i (1905)
(reprint of orig. descript. !).
S. Hindwing on upperside with a row of greenish wliite patches straight
across disc, the middle patches, if large, standing near cell, there being sometimps
an additional spot in apex of cell ; the second to seventh spots nsnally of abont
equal size, occasionally spot 3 to 0 minute and 7 absent, or 2 to 6 large and 7
vestigial, or 2 and 4 to 0 large, 3 small and 7 absent ; size of spots very variable,
sometimes minute, very rarely absent, except the upper two (a male in coll. Godman
from S. Mateo, Bolivia).
? not known to us.
Scent-organ : scales as in P. h. chalceus, short, mostly as broad as long.
Hah. S.E. Peru ; Bolivia.
In the Tring Museum 18 tjt? from: La Union, R. Huacamayo, Carabaya,
2000 ft., December 1904, and La Pampa, R. Huacamayo, 2500 ft., November 1904,
wet season (G Ockenden) ; Cbirimayo, Carabaya, 1000 ft., July 1901, dry season
(Ockenden); Montanas, R. Madre de Dies, Seiitember 1901 (Ockenden); Oroya,
R. Inambari, 3500 ft., November 1901 (Ockenden) ; R. Slucuri, Carabaya, June
1901 (Ockenden) ; Salinas, R. Beni, Bolivia, July 1895 (Stuart); Province Sara,
S. Cruz de la Sierra, March— April 1004 (J. Steinbach).
57. Papilio laodamas Feld. (1859).
PapiUo laodamas Felder, Wicn. Ent. Mm,, iii. p. 393. n. 33. t. 8. fig. 1. ^ (18.^9) (Bogota).
S ? . Close to P. belus varus ; red submarginal spots on underside of hindwing
much thinner and nearer the margin, transverse, angulate, spot C— SC= always
present, being at least vestigial ; no wliite dots distally of these spots. The female
resembling the male, but the band of the hindwing different, the first patch being
much reduced, often to a mere dot, and patches R-— M' being at least as long
as the preceding patches, sometimes patch M'— M= even longer than R'— M' ;
abdomen as in the allied species.
Scent-organ : scales about as long as in J', belus belus, being longer than in
West Mexican P. belus chalceus and shorter than in P. belus varus, occurring from
Guatemala to North Venezuela.
Genitalia : S essentially the same as in P. belus.
Early stages not known.
Ilab. Mexico to Colombia.
Four subspecies.
a. P. laodamas procas Godm. k Salv. (1890).
Papilio procas Godman & Salvin, Biol. Ceiiir. Amer., Rhnp. ii. p. 203. n. 19. t. 65. fig. 15. 16. $ (1890)
(San Bias, Jalisco ; 1 (J).
PapiUa !npas iid., Tram. Ent. Sac. Lm„l. p. 248 (1897) (Colima) ; iid., Binl. Cent:: Amer., lihop. ii.
p. 728. t. Ill, fig. 9. 10. tJ (1903) (Colima, 1 (J).
cJ. First patch of hindwing somewhat smaller than in the other three subspecies ;
the band usually close to cell, there being an additional spot in the apex of the
cell ; sometimes the band separate from cell {iopas) ; vestiges of white submarginal
spots. White spots on underside of forewiug on the whole smaller than in
( 533 )
copanae, especially the upper ones, and the orange-red snbmarginal spots of the
hindwing slightly larger.
? . Greenish band of upperside of hindwing widest in centre, nsnally entering
cell, the patches larger than in the following form, especially patches R^ — M-.
Scent-organ : scales shorter than in the other subspecies.
Ilah. West Blexico : Jalisco, Guerrero, Blichoacan.
In the Triug Museum G ^j^j, 5 ? ¥, from : Colima ; Guerrero (0. T. Baron);
Patzcnaro, Michoacan.
In our two females from Patzcnaro the band of the hindwing does not enter
the cell ; the spots of the forewing are smaller than in the next form.
b. P. laodamas copanae Reak. (1863).
Papiito copanae Reakirt, Proc. Ent. Soc. Philad. ii. p. 141. n. 16 (18G3) (J, Copdn, Guatemala) ;
Kirby, Cut. Diiirii. Lep. p. 521. n. 21 (1871) ; Streck., Lep. Rhop. Net. p. 61. t. 8. fig. 1. ? (1874)
(fig. of type) ; Kirby, l.'c. p. 809. n. 21 (1877) ; Godm. & Salv., Biul. Centr. Amer., Rhup. ii.
p. 202. n. 18. t. 66. fig. 4. ? , 5. 6. (J (1890) (Yucatan; Brit. Honduras; Guatemala; Honduras);
Strecker, I.e., Siippl. iii. p. 17 (1900) (type, ? , in coll. Strecker).
Papilio ehri/sodamas Bates, Ent. Mu. Maij. i. p. 1. n. 2 (1864) (Guatemala) ; Feld., Verh. Zool. Bot.
Ges. Wien xiv. p. 297. n. 122 (1864) (cit. erroneous) ; Kirby, !.e. p. 522. n. 30c (1871) ; Oberth.,
Et. d'Ent. iv. p. 117. n. 307"" (1880) (Mexico).
c?. Band of hindwing always outside cell ; the first patch narrowed proximally,
not touching the cell. Forewing with a row of arrowhead-shaped snbmarginal
spots, spot R^ — R' produced basad ; these spots much more distinct on underside,
there being also a spot in cell near lower apical angle. Red snbmarginal spots
of underside of hindwing thin ; there are often vestiges of white snbmarginal spots
on upperside.
¥. Forewing as in male, spot R- — R^ occasionally vestigial. Band of
liindwing placed outside cell, there being no spot in cell or only a very few greenish
white scales.
Scent-organ : scales about two or three times as long as broad,
Hab. East Mexico southward to Honduras. May be expected to occur in
Nicaragua.
In the Tring Museum 5 c?c?, 4 ? ? , from : Songolica, Espiual, and Jalapa,
Vera f'ruz, April l.s'.)0 and June 1899 (W. Schans) : Coazualcos, Vera Cruz,
July 19U4 (A. Hall) ; San Pedro Sula, Honduras.
r. P. laodamas rkipidius subsp. nov. (PI. IV. fig. 4).
? . Forewing shorter and its distal edge less emarginate than in the two
preceding sulfspecios ; vestiges of snbmarginal spots on upperside ; on underside
four long creamy spots, the npper one reaching cell, and a spot in lower angle
of cell. Hindwing, upperside : a large central area of a greenish straw-colour,
consisting of a patch which occupies about three-fourths of the cell, and six patches
around cell, these patches acuminate distally, contiguous proximally, patch M' — M-
tbe longest, the tips of the patches almost eqnally distant from outer margin;
area between cell and abdominal edge washed with greenish straw-colour ; outside
this area two minute dots R- — M' of the same colour situated near the central
area ; red snbmarginal spots of underside larger than in the next snbspecies, the
first spot distinct.
cJ. A Costa Rica male in coll. H. Druce agrees with the above-described
( 534 )
female. The white area of the hindwing extends almost to the base in and before
the cell ; spots R- — M- become gradually longer, the last being the longest. Spots
on underside rather smaller than in female.
Hah. Costa Rica.
In the Tring Museum 1 ? from Carillo, June— July 10ii3 (Underwood).
(I. P. hiO(l(i7n<ts laoihtmas Feld. (1869).
rajiUio Jaodamas Felder, l.r. (lfi.^9)i (Hogota) ; id., I.e. v. p. 72. n. 3 (1861) ; id., Verh. Zonl. Bol.
Ges. Wkn xiv. p. 297. n. 121 (lf<G4) (Bogota) ; Kirby, Cat. Diiirn. Lep. p. 522. n. 30b. (1871) ;
Obcrth., El. d'Eul. iv. p. 9S. n. 307 (1880) (Toquiza, Llanos de San Martin, Colombia) ;
Maass. & Weym., in Stiibel, Ueiseii S. .imer., Lep. p. 31. n. 125 (1890) (Colombia).
S. Uppevside : forewing without submargiual spots. Hindwing : a broad
deep green distal border, at the edge of which there are sometimes some white
dots ; first patch of greenish white baud large, touching cell, only a few mm.
short of subbasal cellule, followed by four, seldom five, more spots, whicli gradually
decrease in size, touching the cell, except the last one or two ; usually a spot in
cell, oblique, often produced to a streak situated in anterior part of cell along S(J-
and the cross-veins U' to D'.
Umlerside much deeper brown-black than in the Mexican races. Forewing :
three grey snbmarginal spots R' — SM^ and some grey scales before R' and in lower
angle of cell ; these spots not jirolouged. Hindwing : red submargiual spots
more regularly arched than in procas and cojjanae, deeper red, more heavily
bordered with black, the upper ones often partly shaded over with black, the first
being mostly vestigial.
Scent-organ : scales more triangular than in the preceding, being apically
broader, difForing much in outline from P. heliis varus with which the present
insect occurs together.
? not known to us.
JJab. Colombia.
In the Tring Museum ~6 S S from: Cananche, Cnndinamarca, Jnly 1903
(Mathan) ; Mnzo, September 1903 (Blathan), and December 1890; "Bogota";
R. Dagua (W. Rosenberg).
A common insect in Bogota collections.
58. Papilio lycidas Cram. (1777).
Papilio Eqites Achivus lycidas Cramer, Pap. Exot. ii. p. 25. t. 113. fig. A. ? (1777) (Surinam);
Goeze, Eiif. Beylr. iii. 1. p. 81. n. 43 (1779).
Pajiilio Eqties Acliirits enjmanthus id., I.e. p. 25. t. 113. fig. C. J" (1777) (Surin.am) ; Ooeze, fii(.
Beylr. iii. 1. p. 79. n. 35 (1779).
Papilio Eqtien Trojnmis hdns, Fabricius (hoh Cramer, 1777, err. det.), tipiec. Ins. ii. p. 9. n. 34 (1781)
(jiarlim) ; Jabl. & Herbst, XuUirn. S,:l,melt. ii. p. 95. n. 27 (1784) (^partim); Fabr., .Mant. Im.
ii. p. 5. n. 3G (1787) (parlim); Cmel., Syst. Nat. i. 5. p. 2233. n. 294 (1790) (_partim); Fabr.,
Ent. Syst. iii. 1. p. 17. n. 53 (1793) (parlim).
Papilio erimanthus (!), Jabl. & Herbst, ynliirs. Sehmett. ii. p. 97 (1784).
Papilin b/eid,is,'EspeT, A,(.'il. Se/iinelf. p. 80 (1792) ; Boisd., Spec. Gin. L6p. i. p. 317. n. 15G(183G)
(Cayenne : Surinam) ; Doub].,'Westw. & Hew., Gen. Diimt.Lep. i. p. 20. n. 235 (184G); Gray, Oit.
Lep. Ins. Brit. Mux. i. I'ap. p.OS. n. 304 (1852) (Pard); Wall., Trans. Ent. Soc. Land. (2). ii. p. 255
(1854) (Amazons ; banks of rivers); Gray, List Lep. Ins. Brit. .Viis. i. Pap. p. 78. n. 321 {185G)
(Pari!) ; Mene'tr., Enum. Corp. Anim. Miis. Pttr., Lfp. i. p. 6. n. 93 (1857) (" Brazil") ; Bates,
Trans. Ent. Soc. Land. (2). v. p. 228 (18G1) (Par.'i ; Ega) ; id., Journ. Entnm. i. p. 223. n. 5
(18G2) (Upper Amazons & Pari ; Guiana ; habits) ; Feld., Verh. Zwl. Bot. Ges. Wien xiv.
p. 297. n. 123 (18G4) (Surinam ; Par.'i ; Upp. Amazons; Bogota); Kirby, Cat. Diimi. Lep.
p. ,V22. n. 27 (1871) (Guiana; Pari); Distent, rroc. Ent. Soc. Land. p. xiv. (187G) (Costn
( 535 )
Rica) ; Druoe, Proc. Zool. Soc. Land. p. 245. d. 7 (1876) (Haallaga ; Ucayali) ; Obertli.,
Et. d'Ent. iv. p. 99. n. 310 (1880) (TefEi-); Godm. & Salr., Trans. Ent. Soc. Loml. p. 120. n. ■_'42
(1880) (Sta. Marta) ; Staud., Exot. Ta^. i. p. 12. t. 8 J (1884) (Amazons ; Peru ; Venezuela ;
Chirifiui) ; Godm, & Salv., BvA. Cenlr. Amer., Rhnji. ii. p. 201. n. IG. t. Go. fig. 17. genit. (1890)
(Guatemala; Nicaragua; Costa Rica; Panama); Hahnel, /m- iii. pp. 149, 203, 297(1890)
S. Estoban ; Yalera ; Iquitos) ; Maas.s. & Weym.,in Stiibel, Hehen S. Ainer.^ Lf/i. p. 31. n. 123
(1890) (Colombia) ; Haase, n,ikrsurli. Mimicnj i. p. 7G (IS',13) ; Micbael, Iris vii. p. 213 (1894)
(Sao Paulo de 01iven9a) ; Godm. & Salv., I.e. p. 728 (1901) (Honduras) ; Haeusoh, DerJ. Ent.
ZelUchr. xlviii. p. 154 (1903) (Archidona, G40 m.).
Khohalus li/cklas, Hilbner, Verz. bek. ScluneU. p. 88. n. 91G (1818?).
Ilhobalus eri/muiitJiKs, id., I.e. D. 917 (1818 ?).
Papiliu lieliis var. hjcidas, Godart, Ene. ileth. ix. p. 38. n. 42 (1819).
Pupilii) hehta var. erymantlnis id.. I.e.
Papilio erymanlhu.-i, Lacordaire, Ann. Soc. Ent. Fr. ii. p. 384 (1833) (Guyane).
c?. Upperside of abdomen (except claspers) whitish primrose-yellow, as in
the allied species ; a streak of similar colour on biudwing along abdominal fold.
The scales of the nj)perside of both wings nearly all entire, being rounded at apex,
except the scaling in the cell of the forewing and along the distal edge of
the hindwing. The striation of the scales is strongly marked. The scales of the
yellowish white markings of the hindwing are entire, acuminate, like those of
the npperside of the abdomen. There is usually only one spot on the disc of the
hindwing, situated between C and SC-, but many specimens bear a complete row
of spots. The red submargiual spots on the underside of the hindwing are much
nearer the distal margin than in P. belus, there being also no white admargiual
dots between the veins. The forewing bears two or three creamy white submargiual
spots from hinder angle forward.
? . Abdomen green above. White streak along abdominal fold of hindwing
shorter than in male, not reaching base ; there is usually a complete row of spots
on the disc.
Scent-organ : the scales short, some broader than long, multidentate, others
about half as long again as broad, tri- or quadridentate, less often bidentate,
yellowish.
Genitalia: S. Harpe a slightly concave, irregularly square piece of chitin,
produced at the ventral and at the distal corner each in a carved tooth, a
smaller tooth at the oblique upper edge, sometimes accompanied by a second
small tooth. ? . Hairy lobes situated in the vaginal cavity somewhat acuminate,
rather close together ; postvaginal sclerite, which forms the roof of the cavity,
with a thin mesial carina.
Early stages not known.
Hab. Guatemala southwards to Bolivia, and eastwards to Pard ; may be
expected to occur in the province of Goyaz, Brazil.
The species does not seem to be separated into geographical races.
In the Tring Museum 56 Jc?, 4 ¥ ?, from : Moran, W. Guatemala, 45U0 ft.,
August 1904 (A. Hall) ; San Jos5, Costa Rica (Underwood) ; G'hiriciui ; Muzo,
Colombia, December 1890 ; K. Dagua (W. F. H. Rosenberg) ; Coca and Archidona,
Ecuador (W. Goodfellow) ; R. Ucayali, and R, Cachyaco (iStuart) ; Palca/.u, Juuin
(Hollmauns) ; R. Mixiollo, Loreto (Baer) ; La Union, R. Huacamayo, Carabaya,
2000 ft., November and December 1904 (G. Ockcnden) ; Province Sara, S. Cruz de
la Sierra (J. Steinbach) ; Blerida, Venezuela (Briceno) ; Palma Sola, N. Venezuela ;
Snapure, September 1899, La Union, September 1901, and La Vuelta, May 1903,
Canra R., Orinoco (S. M. Klages) ; Paramaribo, Surinam.
( 536 )
59. Papilio crassns Cram. (1777).
Papilio Eques Trojanus crassus Cramer, Ay>. ExnI. ii. p. 23. t. 112. fig. C. (? (1777) (Surinam) ;
Goeze, Ent. Beytr. iii. 1. p. 85. n. 58 (177U)-
rujiiliii Eques Trojanns beluf, Fabricius (nan Cramer, 1777, err. det.), Sjirc. Inx. ii. p. 9. n. 31 (1781)
(parlim) ; Jabl. & Herbst, Natitrs. Schmelt. ii. p. !I5. n. 27 (1784) (parlim) ; Fabr., Mnnt.
Ins. ii. p. 5. n. 3G (1787) {parlim) ; Gmelin, Si/sl. Xiil. i. 5. p. 2233. n. 29-1 (17'.>0) (parlim) ;
Esper, Aiisl. Schmett. p. 80 sub n. .3.5, and p. 11(5. n. 5f). t. 27. fig. 3. ^ (17112) : Fabr., Knl.
%< iii. 1. p. 17 n. 53 (1793) (;OTWim).
Princeps dominans crassus, Hiibner, Snniml. E.rot. Si'hmell. i. t. 131 (1806— ?).
Ilhohalus crassus, id., Vers. bth. Srhmell. p. 88. n. 914 (1818?); Kirby, in Hiibner, .S„m»i/. £j:«^
ScJimHt. ed. ii. p. 92. t. 131. fig. 1. 2 (190—?).
Papilio belus var. crassus, Godart, Enc. Melh. ix. p. 38. n. 42 (1819).
Papilio crasnis, Lacordaire, Ann. Sue. Ent. Fr. ii. p. 385 (1833) (Gayane ; descr. of larva and pupa) ;
Lucas, Up. Exot. p. .33. t. 17. fig. 1. ^ (1*'^5) ; Boisd., Spec. Gen. Up. i. p. 314. n. l.'.S (183G)
(Brazil ; Gu.iane ; larva and pupa) ; Lucas, in Gu^rin, Diet. Pilt. Nat. vii. p. 48 (1838) ;
Doubl.. List Up. Ins. Brit. .Uns. i. p. 14(1845) (Brazil) ; id., West. & Hew., Gen. Dinrn. Up.\,
p. 19. n 232 (1846) (Brazil ; Cayenne) ; Gray, Git. Up. Ins. Brit. Mus. i. Pap. p. 68. n. 307
(1852) (Brazil) ; id.. List Lep. Ins. Brit. Mus. i. Pap. p. 79. n. 324 (1.856) (Brazil ; Parji) ;
Mi'ni'tr., Enuni. Corp. Anim. Mm. Petrup., Lep.K. p. 6. n. 91 (1857) (Brazil) ; Bate.", Trans. Ent.
Sue. Land. (2). v. p. 227 (1861) (Pari ; S. Paulo, Upper Amazons) ; id., Juurn. Entom. i. p. 223.
n. 1 (1862) (Upp. Amazons ; Pani) : Feld., I>)7). Zonl. Bat. Ges. Wien xiv. p. 298. n. 130
(1864) (Surinam : Amazons ; Brazil) ; Kirby, Cat. Dinrn. Lrp. p. 522. n. 33 (1871) ; Capronn.,
Ann. Soc. Ent. Belg. xvii. p. 9. n. 9 (1874) (Rio, Sept. ; Botafogo, Nov.) ; Druce, Pmr. Zool.
Soe. Land. p. 245. n. 9. (1876) (Peru) ; Butl., Ann. Mar/. X. 11. (4). xx. p. 127. n. 59 (1877)
(R. Mairo, Peru) ; Burm., Descr. Rep. Argent, v. Lep., Atlas p. i>. n. 11 (1879) (larva, pupa ;
Rio de Janeiro) ; Oberth., El. dEnt. iv. p. 99. n. 311 (1880) (Ecuador ; Brazil) ; Stand., E.mt.
Tu(if. i. p. 12. t. 8. S (1884) (S. Brazil ; S. Peru to Venezuela) ; Maass. .and Weym., in
Stiibel, Reisen S. Amer., Lep. p. 91. n. 37 (1890) (S. Catharina 0 ; Haase, Untcrswh. Mimicr>i i.
p. 76 (1893) ; Michael, Iris vii. p. 213 (1894) (Sao Paulo de Olivenfa) ; Bonningb., Verh. Ver.
Nat. Unlerh. Uamhiiry ix. p. 28 (1895) (Rio de Janeiro ; rather common); Peters, lilustr.
Zeitschr. Ent. ii. p. 52 (1897) (Nova Friburgo, larva, pupa) ; Haensch, Berl. Ent. Zeitsclir. xlviii.
p. 154 (1903) (Archidona, 640 m.) ; Weeks, IU„str. Diurn. Lep. p. 20 (1905) (Chulumani,
Bolivia).
Papilio Irpidus Felder, Wien. Ent. Man. v. p. 72. n. 1. (1861) (Caracas; coll. Kaden, var. in coll.
Godman) ; Bates, /o»i7(. &((. i. p. 223. sub n. 1 (1.S62) (Venezuela, local form of crassus);
Feld., Verh. ZuuL Bat. Ges. Wien xiv. p. 208. n. 129 (1864) (Venezuela ; BogoU) ; id., Reise
Novara, Lep. p. 40. n. 29. t. 10. fig. a. (J (1865) (Venezuela ; Bogota) ; Staud., Exot. Taaf. i.
p. 12 (1884) (Colombia ; Venezuela) : Godm. & Salv., Bial Centr. Amer., Rlinp. ii. p. 203.
n. 20 (1890) (Panama) ; Weeks, llhixtr. Dinrn. Lep. p. 20 (1905) (Chulumani, Bolivia).
Papilio crassus var. a. P. lepidus, Kirby, Cat. Dinrn. Lep. p. 622. sub n. .33 (1871) (Venezuela).
<?. White scales on npperside of abdomen acuminate ; the proximal segments
more or less extended green-black. Scalin.fc of npperside of forewins dcniticulate ;
I wo large patches R'— M-, a long streak in cell and a streak behind M- occnpying
the hinder angle behind M'^ and cell, yellowish cream-colonr ; these patches often
reduced or entirely absent ; the same patches on the underside, but paler, here
often preserved in specimens in which they are absent from the upperside.
Scaling of upperside of hindwing non-dentate ; a large greenish white ])atch
between C and SCI- reaching from sabbasal cellule to near apex of C, limited in
front by this vein, many specimens with a short streak behind SC- near distal
margin ; snbmarginal spots of underside brick-red, occasionally slightly nifoiis,
usually accompanied by more or less distinct bnt miunte white admargiual dots
which stand at the veins (as in the allied species), not in the centre between the
veins, being remnants of transverse bars ; there is rarely a red spot between
C and SC%
? . Wings similar to those of male, but the white subcostal patch of the
hindwing reduced, being represented by an ill-defined snbmarginal spot.
( 537 )
Scent-organ similar to that of P. lycidas, the scales being short and broad,
deuticnlate.
Genitalia: S- Harpe small, short, triangular, rather broader than long, apex
rounded, ventral edge dilated into a flat triangular process as in P. hclus, laodamas,
etc., this process standing almost vertically on the plane of the harpe, leaning
dorsad, being curved basad, denticulate at distal edge, the dentition extending
to apex of harpe. ?. The two hair}' flaps situated proximall}- in the vaginal
cavity rounded, widely separated from each other.
Early stages described by Lacordaire, I.e. (1833), and again shortly noticed by
Burmeister, Lc. (1879), and Peters, I.e. (1897).
There are no geographical forms, but the male is dichromatic in the forewing,
only one form of the female being known.
a'. cJ-f. crassus Cram., l.r. Forewing with yellowish creamy patches.
b'. <S-{. lepidtis Feld., I.e. No creamy patch on npperside of forewing.
This male form alone occurs in Venezuela, Colombia, Panama, and Costa Rica, the
females from these countries not being devoid of the patch. Males with hardly
any creamy scaling on the ujiperaide of the forewing occur also in other districts,
for instance Bolivia.
Ilah. of P. erassus : from Costa Rica southwards to Rio de Janeiro.
In the Tring Museum 60 cJc?, 10 ? ?, from: Carillo, Costa Rica, 300U ft.,
October 1904 (A. Hall) ; Chiriqui ; Muzo, Colombia, December 1896 ; Pereira,
Popayan and Call, Canca ; Zamora, Ecuador (0. T. Baron) ; Loja ; Palcazu, Junin
(Hofl'manns) ; Chanchamayo ; Pozuzo ; R. Cachyaco, affluent of R. Huallaga
(Stnart) ; La Union, R. Huacamayo, Carabaya, 2n00 ft., November and December
1904, wet season (Ockenden) ; R. Sougo to R. Suapi, Bolivia, 1100 m., March-
April 1S96 (Garlepp) ; Mapiri, 1000 ft., September 1895 (Stnart) ; Villa Maria to
Diamantino, Matto Grosso, January 1897 (Andeer) ; Tijnca ; Petropolis ; Amazons :
Iquitos and Raituba ; Surinam ; British Guiana ; Caura R., Orinoco, May 19o0
(S. M. Klages) ; Valencia, Venezuela.
SECTION II.— FLUTED SWALLOWTAILS.
(For differential characters see p. 433.)
This section contains the largest number of species. It is the most widely
distributed Section, occurring in all regions, except the Antarctic, going in one
species {P. machaoii) as far north as Alaska and Kamtchatka. The variety in
structure and pattern among the species is very great. Owing to the preservation
of many intermediate species, the division of this Section into minor groups is not
quite so well marked as in Sections I. and III.
The American si)ecies fall into two Subsections, which, taken each as a whole,
are characterised, the one by the prevalence of yellow colour and the softness of the
costal edge of the forewing, the other by the prevalence of black on the body and
wings and the hardness of the costal margin of the forewing, the insects of this
second Subsection being more strong-winged than those of the first Subsection.
However, in the second Subsection there is a mimetic group of soft-winged species
{Zagreus Group).! which appears to have acquired secondarily the soft costal edge
and the great amount of yellow on the wings and body. This mimetic group
exhibits a most curious feature in the coloration of the head. While in all Papilios
( 538 )
which have the frons striped with yellow, a yellow stripe is situated on each side
along the eye, P. zagrcus and allies have a yellow line in the middle of the frons.
The obtnse apex of the forewiug and convex distal margin, the position of SC' of
the forcwing proximally of the upper angle of the cell, the great width of the cell
and the long slender antennae are further peculiarities which separate P. zagreus
and allies from the other American Papilios. The hard dentate costal edge of the
species of the second Subsection (apart from P. zagreus and allies) has doubtless
been evolved in connection with a strong flight. We find the same feature in a
group of Symphalidae, namely, the prionopterous genera Charaxes, Eulepis, and
allies, which are all strong fliers, and also in the males of some Pieridae. The
great robustness of the thorax (as compared with the size of the wings) observed
in those Nymphalids is again met with iu the Papilios with serrated costal edge
to the forewing.
The following generic names have American species for name-types, nomina
nuda not being here mentioned :
Euphoeades Hiibner, Verz. bek. Schmett. p. 83 (1818 ?) (typo : glauaix).
IleracUdes id.. I.e. (type : crcsphontes).
Calaides id.. I.e. p. 86 (1818 ?) (type : androgeus).
Priamides id.. I.e. p. 87 (1818 ?) (type : tonjuatus).
Pgrrhosticta Butler, Cist. Ent. i. p. 86 (1872) (type : eleotas).
Troilides Kirby (ex Htibner, indescr.), in Allen's Nat. Lihr., Butt. ii. p. 283
(1896) (type : torquatus).
Subsection U.*
Palpus yellow at side. Frons yellow, or with black mesial stripe, rarely all
black. Abdomen at least with yellow dots at sides, usually with yellow stripe or
for the greater part yellow. Scaling at the costal edge of forewing very dense and
tough, difflcult to rub off, the edge non-dentate.
We divide the American species into six groups :
a. Hindwing on underside with a subbasal and a submedian
band, which unite near anal angle, forming a large
black V Glaucits Group.
No such bands b.
b. Pronotum and underside of thorax with red or orange
dots; no metallic blue spots on underside of hindwing Aneldsiades Group.
No such spots ........ c.
c. No metallic blue spots on underside of hindwing ;
abdomen yellow beneath, with black line along
upper edges of sternites ; if this line absent, then
tail with yellow marginal spot at apex; or abdomen
black, with yellow line along lower edges of tergites,
hindwing without regular row of yellow submarginal
.spots ......... '/'onp/iif/i.-i (jroHj).
With UR'tallic blue spots on hiiidwiDg .... d.
d. Abdomen striped with black and yellow beneath, or
black dotted with yellow, there being two rows of
yellow dots on each side of the tergites ; cross-veins
of forewing with yellow bar at least on underside . Maekaon Group.
* For Subsection D see after .Species No. 101.
( 539 )
Abdomen striped with lilack and yellow, antenna pale
tawny ; or abdomen black, dotted with yellow, only
one row of dots on each side of the tergites, antenna
black Troilus Group.
Abdomen entirely yellow beneath and at sides ; or all
black, except a row of yellow dots situated laterally
on the sternites, there being no dots on the tergites ;
or the abdomen black (? ¥), with a yellow line
at the lower edges of the tergites, the hindwing
of these females bearing a regular row of yellow
snbmargiual spots at least on underside . . Thoas Group.
V. Machaon Group.
Antenna reaching to two-thirds of cell of forewing ; club thick, obtuse, the
last segment very short, the preceding three or four more than twice as broad
as long. Abdomen either with yellow longitudinal bauds, or black, with rows of
yellow spots, two rows on each side of the tergites. Tail nou-spatnlate. Basal
half of hindwing yellow or black, without the large V formed by black bands in
the Glaucus Group. Cell of hindwing not widened. Harpe of male saw-like, tiie
distal portion being a denticulate ridge and the pro.ximal portion being less elevate,
non-dentate, subeylindrical. In female on each side of vaginal orifice a long
three-cornered flap which is dentate at the edges ; proximally of these flaps
a continuous, slightly elevate, ridge from side to side ; behind the orifice a
membranaceous tubercle clothed with extremely small hairs.
The anal eye-spot, composed of the marginal and submarginal spots, is better
developed in this group than in the Glaucus Group. The origin of the ocellus
from the two sjjots mentioned is well illustrated by P. daunus and allies ; also
iu the present group the component submarginal and marginal sjjots remain
occasionally sejiarate.
With the exception of the Asiatic P. xuthu.s, which stands apart, the species
of the Machaon Group are all very closely allied to one another. P. indra is the
only one which is recognisable by the genital armature, all the others being identical
in structure, as far as we know. The larvae also not presenting any very trenchant
characters, one might well ask if P. machaon, the dimorphic P. bairdi, the variable
P. polyxenes, the comparatively constant P. ::cUcaoii and P. nitra are really
specifically distinct from one another. Judging from the structure and pattern
aloue, one might be inclined to regard all these insects as individual and
geographical forms of one species. However, one important fact is known which
speaks entirely against this assumption. Papilio bairdi and P. pobjxen.cs asterius
occur in the same canons in Colorado, but keep perfectly separate, the one living
as caterpillar on Umbclliferae, the other on a Composite plant {Artemisia dracunca-
loidu.i). If these two insects exist iudej)endently side by side — i.e. are true species —
there is no reason to treat P. zclicaon otherwise than as a species by itself. And
P, machaon aliaska, which occurs as far south as Oregon according to the two
Edwardses, must also be considered independent of P. bairdi f. oregonia in spite
of the close general resemblance in pattern. We doubt if P. nitra is more than
a northern form of P. bairdi.
35
( 540 )
Papilio xutkus and the Pacific Palaearctic P. mackaon hippoo-ates are
seasonally dimorphic. In the American species dimorphism appears to be purely
individual, the two varieties being produced by one female independent of the
season, as is the case in some African Precis. However, besides the dimorphic
P. bairdi from Colorado, no other dimorphic form has been actually reared.
Breeding from the eggs of one female of P. nitra, of an Oregon female of
P. bairdi f. oregonia, of a Mexican female of P. pnli/xenes asterius, and of a
female of P. polyxeiies americus, is a great desideratum.
Key to the American species :
a. Yellow patch IP — SM- of forewing close to cell . . b.
Yellow patch M^ — SM- of forewing widely separated from
cell, sometimes absent . ...... d.
b. Abdominal sternites yellow, with two black lines . . c.
Abdominal sternites black, with or without indication of
yellow lines Species No. 63.
c. Anal ocellus with pupil ....... Species No. 61. c'.
Anal ocellus without pupil Species No. 65. a-
d. Palpus and breast entirely black ..... Species No. 64.
Palpus yellow ......... e.
e. Abdomen dotted with yellow f.
Abdomen with broad yellow lateral stripe . . . Species No. 61. i'.
f. Discal spots of hindwing below reddish orange, more or
less edged with yellowish cream ; or the yellow band
gradually shading off proximally, often extending to
base ; or the discal spots of upperside small, shaded
over with black, the baud being distal of cell on
hindwing Species No. 60.
Orange colour of discal band of underside of hindwing more
gradually shading off, the band always crossing apex of
cell, never extended to base . . Species No. 61. a and No. 62.
60. Papilio polyxenes Fabr. (1775).
Papilic Equea Achivus ajax Linnu, Si/st. Nat. ed. x. p. 462. n. 2G (1758) (imrtini) ; Clerck, Icon. his.
ii. t. 33. fig. 3. J (1704).
Papilio Eques Trnjanus pobjxenes Fabricius, Syst, Ent. p. 444. n. 10 (1775).
Linn^'s description of ajax applies, we think, to the present insect. The two
references given after the description count for nothing, a])plying to totally different
insects, the second being quoted later by Linnd himself under P. protcsilaus. In
the twelfth edition of Si/st. Nat. Linnd described P. xuthus as being very similar
to P. ajax. Esper (179{S) tried to explain this comparison by assuming that Linne'
meant Clerck's ajax, not his own. The three facts: (1) that Linne characterised
the anal ocellus of ajax and machaon by the same words {angulo ani falvo), (2) that
Clerck's work was executed under Linne's eyes — one might almost say under his
supervision, and (3) that in 1767 Liun6 describes P. xuthus as being very similar
to ajax, leave little doubt that the true ajax of Linn^ was the insect which Cramer
named asterius (corrupted by Fabricius into asterias), which is the same species
as polyxenes Fabr., but a different geographical race. However, since Linne
himself misled later authors by the erroneous quotation of Kaj. iii. ~ and Edw.
(541 )
av. 34Jnnder ajax, wo think the wisest coarse to follow is to siijipross the name
ajax altogether.*
The subspecies of P. poli/xenes, which name comes next in priority after ajax,
do not all corapletel_v grade into one another. The differences are, liowever, so
slight that the close connection between the various forms becomes at once evident
on comparing a long series.
a. P. polijxems amcricus Kollar (185iJ).
PapiUo amcrii/tix Kollar, Denkschv. K. Ak. II'/'ss. Wien, Math. Nat. CI. i. p. 354. n. 10 (18."j0)
(" N. Granada, ad ripas fluminis Orinoco") ; Doubl., Westw. & Hew., I.e. (1852) ; Gray, Cat.
Lep. Ins. Brit Mus. i. Pap. p. rtS. n. 294 (1852) ; id.. List Up. Tns. Brit. Mu.i. i. Pap. 76. n. .311
(1856); Felder, Verb. Zopl. Bol. Ges. Wicii xiv. p. 315. n. 359 (1864) {= saAdus ; Bogota;
Venezuela ; Ecuador ; " Mexico ? " false) ; Kirby, Cat. Diuru. Lep. p. 566. n. 322 (1871) ; Edw.,
Tram. Amer. Eiit. Sor. vi. p. 10. n. 13 (1877) (" S. Calif., Arizona" false ; = sarlaliis) ; Staud.,
Exot. Tagf. p. 18. t. 12. J (1884) (iiartlm ; Colombia ; Ecuador ; Venezuela) ; OllifP, Prnr.
Ent. Soc. Loud. p. 22 (1881) (monstr. in neuration) ; Maass. & Weym., in Stiibel, Rcisen S. Amer.,
Lep. p. 14. n. 12 (18110) (Bogofci, 2600—3000 m) ; iid., I.e. p. 34. n. 30 (1890) (Oparapo,
Colombia, 1200 m.) ; iid., /.c. p. 41. n. 5 (1890) (Ales, near Pasto, 2100 m.) ; iid., /.-•. p. 48. n. 1 1
(1890) (near Ibarra, 2370 m.); iid., /..-. p. 56. n. 80 (1890) (Bano.^, 1800 m.) ; iid., /..-. p. 58. n. 13
(1890) (Pululagua, 2.500 m.) ; Hahnel, Iris iii. p. 185 (1890) (Mi'-rida) ; Godm. & Salv., Binl.
Ceulr. Aiwr., Lap. Rhiip. ii. p. 243. n. 82 (1890) [pa,rtim\ Colombia; Venezuela; Ecuador;
"Angostura" false); Mayu., Man. N. Amfr. Bidt. p. 6. n. 6 (1891) (Arizona); Edw., Butt. X. Amer.
i\i. Pap. iii. fig. 1. 2. (J, 3. 9 (1891) ('■ Arizona" error loci; Bogota; "Southern Mexico to
Ecuador "yjart/m) ; Godm. & Salv.. in Whymper, Andes of Equator, App. p. 109. n. 95 (1891)
(Macbachi, 10,000 ft.) ; Haase, Uiitersuch. Mimicry i. p. 92 (1893) ; Eimer, Arthild. Verwaiultseh.
Sclimetl. ii. p. 125 (1895) {partim ; — t. 7. fig. 3. represents a Chiriqui specimen).
Papilio sardahis Becker, Bull. Soc. Ent. France p. 33 (1851) Qwm. nudum ! Quito).
Papilio sadalus Lucas, in Gue'r., Rer. Zool. (2). iv. p. 133. t. 10. fig. 4 (1852) (Quito) ; Doubl., Westw.
& Hew., I.e. ii. p. 529 (1852); Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p .39. n. 185 (185^) (Quito) ;
id.. List Lep. Ins. Brit. Mus. i. Paj). p. 51. n. 191 (1856) (Quito; Colombia; Venezuela);
M^ne'tr., Enum. Corp. Auim. Mus. Petrop., Lep. i. p. 4. n. 59 (1857) (Quito) ; Godm. & Salv.,
Trans. Ent. Soc. Land. p. 126. u, 236 (1880) (Sta. Marta) ; Oberth., Et. d'Ent. iv. p. 68. n. 195
(1880) {pnrtim; Ecuador).
Papilio asterins var. a, Gray, Cat. Lep. 7«-i. Brit. .Mus. i. Pap. p. 38. sub n. 184 (1852) (Venezuela).
Papilio asterioides, Strecker (non Reakirt, 1866, err. det.), Lep. Rhop. Het. p. 47. t. 6. fig. 4. J
(1873) (■' Costa Rica " false).
Papilio asierius, Dewitz, in Wiegm., Arch. Xaturg. xxxxiv. 1. p. 4. t. 1. fig. 2. 3. 4 (1878) (larva,
pupa ; Venezuela).
Painllo 2foli/.reucs, Staudinger (nou Fabr., 1775, err. det.), E.rot. Tagf. p. 18 (1884) {partim ;
Venezuela ; Colombia).
Pajiilio sadatus (!), Haensch, Berl. Ent. Zeitschr. xlviii. p. 152 (1903) (Banos, R. Pastaza, 1800 m.).
Papiilio polgxencs var. asterioides, Maassen & Weym. (non Reakirt, 1866, err. det.), in Stiibel, Reisen
S. Amer., Lep. p. 38. n. 34 (1890) (Popayan).
c? ? . Specimens which resemble the following subspecies in the width of the
yellow discal band can be distinguished by the last spot of this band on the
forewing being produced basad and by the black basal area of the hiudwing
below being much less sharply defined, gradually shading off.
The subspecies is remarkably dichromatic, in one extreme the yellow colour
of the liindwing extending to the very base, while in the otlier extreme the yellow
discal baud is much reduced. There arc all iutergnidations. The black specimens
were erroneonsly called pobjxenes Fabr. by Staudinger, and asterioides Reak. by
Maassen & Weymer and by Strecker.
a', f. amcricus Kollar, I.e. The discal baud of the forewing, above, is often
washed with orange, as is also the case in some specimens of P. poli/xeiies stabilis,
* 8ee p. 414 ; also P. marcellus.
( 542 )
while one of onr females (M(5rida) is almost as pale as P. imh-a ; tlie last sjiot of
the band is uearly always mnch longer than the jiatch M- — SM-. The base of the
hindwing is in most individuals black as far as SC- or a little beyond, but in
many specimens the black basal area is much restricted, having practically
disappeared iu a few individuals. On the underside the basal area of the hind-
wing is more or less washed out distally, the black scaling being in a largo
percentage of the specimens restricted to the extreme base or being altogether
absent ; sometimes the forewing is also much shaded with cream-colour. The
abdomen bears often a broad yellow lateral stripe, with which the dorso-lateral
dots are more or less merged together. The postmedian cell-bar of the forewing
is often more or less distinct on the underside.
The figures in Edwards, Butt. N. Amer., I.e., represent average specimens of
this form ; figs. 1 and 2 agree perfectly with typical Bogota specimens. We do
not believe the original of these figures came from Arizona. If the specimen had
been a straggler from the south, one would expect it to belong to the Central
American form, which it does not. Some mistake in labelling is, we think, the
true explanation of the record of americus from Arizona.
b'. f. iiiclasina nov. Yellow discal baud of forewing reduced on both wings
to a row of spots which are shading off proximally ; bar on cross-veins of forewing
present. The band rather broader on underside, but never entering cell of hindwing,
more sharply defined on both wings than above, but occasionally gradually shading
off jiroximally ; the basal half of the hindwing rarely washed with buft'.
These black individuals resemble the dark Mexican form asterioides of
P. poltjxenes asterius ; however, the forewing is proportionally broader, the discal
sjiots are powdered over with black proximally, the discocellnlar bar is present,
the tail is shorter, etc.
This form appears to occur more frequently in the C'anca valley (probably at
high altitudes) than anywhere else. Type of name from Pereira, Cauca valley.
Strecker, I.e., figured as P. asterioides Eeak. a female of this form, said to be from
Costa Rica. It is not impossible that a melanotic form occurs in that country,
but we cannot accept Strecker's statement without further evidence. Mistakes as
to locality occur easily, and we know for certain that at least in one case among
Papilios Strecker gave a wrong locality, his I', cleombrotus being erroneously stated
to have come from the Amazons.
Hab. of P. jjobjxenes americus : North Peru to Colombia and Venezuela.
The type of americus came doubtless from the Cordillera of Bogota, which
Sulkowsky traversed on his way from the R. Magdalena to the Orinoco, going by
the R. Meta.
In the Tring Museum 110 cJcJ, 37 ? ?, and 4 pupae, from: Cayambe, N.W.
Ecuador, Jnne — July 1897, 9000 ft. (W. Rosenberg) ; Banos (R. Haensch) ;
Pereira, Cauca; "Bogota"; Villavicencio to Monte Redondo, March 1897
(Dr. Burger); Valdivia, Colombia, July 1897 (Pratt); Bogota to Coachi, 2800
down to 1700 m., January 1807 (Ur. Burger); Merida, Venezuela, L'OHO— 300n m.,
October to January (Briceno).
b. I', jjoli/xenes stabilis subsp. nov.
I'tipilio siiththiK, Butler & Druce {non Lucas, 18o'i, err. (let.), Proc. Zool, Sot; Lviul. p. 365. n. 383
(1874) (Costa Rica) : Obertb., Et. d'E/il. iv. p. 08. n. 195 (1880) {parlhn ; Chiriqui).
( 543 )
Papilio americim, Edwards, Papilin iii. p. 55 (1883) (Panama ; Cbiriqui) ; Stand., E.rut. Tmif. p. 18
(1884) (jiarlhn ; Honduras to Chiriqui) ; Godm. & Salv., Bin/. Centr. Amer., Leji. Rhnp. ii.
p. 243. n. 82. t. 72. fig. 8. genit. (1890) [partim \ Costa Rica ; Chiriqui ; not farther north) ;
Eimer, Arthild. Verimndtsih. Srhmell. ii. p. 125. t. 7. fig. 3. ? (1895) (partim).
S ? . The Central Ainericaa specimens do not vary so much inter se as do the
South American ones, the present subspecies being monochromatic. The yellow band
across the disc of both wings is always broad, not varying very much in width ;
the last spot of this band on the forewing is about as long as the patch before it,
not projecting basad. There is sometimes a postmedian cell-bar present above and
below, being larger on underside. On the hindwing the inner edge of the band
crosses the cell at point of origin of SC- or proxiraally of it.
On the niidcrside the' band is proximally much more sharply defined than in
the South American form, the hindwing being jnirer black from base to band.
Hab. Costa Rica; Chiriqui; Sevilla I., Pacific side of Panama; Honduras
{teste Staudinger, I.e.) ; name-type from Costa Rica.
The only specimen (a ?) from Sevilla Island which we have is of interest, the
anal ocellus lacking the black dot above and below.
Eimer's figure, I.e., is undoubtedly taken from a female of the present subspecies
(probably from a Chiriqui specimen received from Messrs. Staudinger & Bang-Haas),
though the habitat is given beneath the figure as being Colombia, Venezuela, and
Ecuador, which countries are inhabited by the preceding subspecies.
The dot in the yellow spot SC — SC is absent from one of our Chiriqui females.
In the Tring Museum 46 c? cj, 14 ? ? , from : Sevilla I., January 1902 (Batty) ;
Chiriqui ; Boquete, Chiriqui, 3500 ft. (Watson) ; San .Juan, Costa Rica, 4000 ft.,
September 1904 (A. Hall) ; Carillo, Costa Rica, 3000 ft., October 1904 (A. Hall) ;
S. Jos^, 4000 ft., October 1904 (A. Hall); Escazu, Costa Rica, October 1903
(Underwood); Puriscal, October 1903 (Underwood) ; Cartago (Underwood) ; Guatil
Piris, December 1901 (Underwood); Alahuela, 4000 ft., September 1904 (A. Hall).
e. Papilio pohjxenes asterius Cram. (1782).
Petiver, Gas,q,h. t. G. fig. 12 (1709).
Pdpllio Eques Achivus ajax Liane, Syst. Nat. ed. x. p. 462. n. 26 (1758) (^)fi)((m) i Clerck, 7™n.
Lis. ii. t. 33. fig. 3. ^ (1764).
Pajnlio Eqiies Trojcimis trnilus, Drury {non Lian^, 1758, err. det.), lllustr. Exot. Ins.i. p. 22. t. 11.
fig. 2. (J, 3. ? , and Index (1770-71) (N. York ; Maryland ; Virginia) ; Fabr.,Sys^ Ent. p. 444.
n. 7 (1775) (parlim) ; Cramer, Pap. Exot. iii. p. 25. t. 207. fig. A. ? (1779) (N. York ;
"Jamaica" false ; ocellus blind) ; Goeze, Ent. Beyli: iii. 1. p. 31. n. 6 (1779) (partim) ; Jabl.
& Herbst, Naturs. Schmelt. ii. p. 242. n. 58. t. 17. fig. 3. 4 (1784) (parlim).
Papilio Eques AehiruH asterius Cramer, Pap. E.rot. iv. p. 194. t. 385. fig. C. D. ^ (1782) (N. York ;
Carolina ; Virginia) ; Esper, Aiisl. Sclimett. p. 47. n. 18. t. 11. fig. 1. 2 (1786) (N. York).
Papilio Eques Trojanus poli/xeiies, Jabl. & Herbst, Nalia-s. Srhmelt. ii. p. 253. t. 18. fig. 1. g (1784)
(= asterius = ajax Clerck ; puirlitn).
Papilio Eques Trojanus asterias (!), Fabricius, Mant. Ins. ii. p. 2. n. 13 (1787) (ixirtim) ; Gmelin,
Si/st. Nat. i. 5, p. 2228. n. 280 (1790) (partim) ; Fabr., Ent. Sijst. iii. 1. p. 6. n. 16 (1793).
Papilio troilus, Abbot & Smith, Ins. Geortjia ii. p. 1. t. 1. (J. ? . 1. p. (1797); Butler, Cut. Diurn. Lep.
ilescr. Fabric, p. 249. n. 52 (1869) (" S. Domingo" perhaps erroneous, the ? in Brit. Mns.
being like ordinary North American ? ? ).
Papilio asterias, Esper, I.e. p. 248. t. 40ii. fig. 6. ? (1798) ; Gray, Cat. Lep. Ins. Brit, il/us. i. Pap.
p. 37. n. 184 (1852) ; id., List Lep. Ins. Brit. Mns. i. p. 51. n. 193 (1856) (partim) ; Gosse,
Letters from Alabama p. 78 (1859); Rrak., Proc. Ent. Sor. Philad. vi. p. 123 (1867) (Colorado) ,
Strecker, Butt. Moths N. Amer. p. 71. n. 17 (1878) ; Haase, Untersiie/i. Mimicr;/ i. p. 92 (1893) ;
Grant, Canad. Ent. xxviii. p. 273 (1890) (Orillia, Ont., formerly common, now rare) ; Eimer,
Orthogen. p. 37. fig. 17 (1897) ; Bubna, Ent. News viii. p. 98 (1897) (Cleveland, Ohio ; common
( 544 )
as Dsual); Duzee, n,iU. Buffalo Soc. N. Sri. v. p. 107, n. 3 (1897) (Buffalo) : Christ, Mitt.
Sclnoeiz. Ent. Ges. ix. p. 271 (18'J7) ; Thorns., Caiuul. Ent. xxix. p. 2lj.'i (18',i7) (larva on liuta
graveolensl); Beutenm., Bull. Anier. ^flli:. N. H. x. p. 310 (1898) (Highland Falls, N.Y.) ;
Fyles, Rejil. Eiil. Soc. Ontario xxix. p. 44 (1899) (on parsnip, carrot, etc.) ; Holland, Bull. Book
p. 314. t. -2. fig. 17. 24. 27. larva, t. 6. fig. 13. 18. 19. pupa, t. 40. fig. 1. cJ (1899) ; Webst., E„t.
iVws xi. p. 577 (10nO)(larva on Cosmos); Beat.. .Uolhs Bail. T.S. p. 340. fig. jj. ? (1898—1900);
Beutenm., Bull. X Yoii- Cilij p. 4. fig. (J (1902) ; Wasm., Ent. Neicsxm. p. 29 (1902) (a larva
feeding in January) ; Ellsw., /Wd. p. 104 (1902) (aberr., similar to indra, Lestershire, N.Y.,
June 17, 1899) ; Comst., ihiil. xiv. p. 197 (1903) (Adirondack Mts., rare in Aug.) ; Laur., il/id.
p. 290 (1903) (Miami, Fla., common).
Euphoeades asterius, Hiibner, Verz. be!.: Schiiiett. p. 83. n. 849 (1818 ?).
Papilin a.tleria.'t, Godart, Enc. Meth. ix. p. 58. n. 91 (1819) ; Boisd. & Lee, Hint. (!iii. Lep. Amh:
Sept. p. 14. t. 4. cj. $ . 1. p. (1833) (parliin ; Virginia ; Georgia) ; Luca.a, Pap. Ej-vI. p. 38.
t. 20. fig. 1 (1835) ; Boisd., Spec. Gen. Lep. i. p. .3.32. n. 175 (mK){parlin, ; U.S.A. ; Mexico) ;
Drury, ed. Westw., Illuslr. Exot. Ins. i. p. 21. t. 11. fig. 2. 3. 5 (1837) ; Harris, his. Inj. Veget.
p. 212 (1841) ; Doubl., List Lep. Ins. Brit. Mm. i. p. 15 (1845) ; id., Wcstw. & Hew., Gen.
Diurn. Lep. i. p. IG. n. 161 (1846) ; Karsten, Arch. Anal. Phi/s. & Med. p. 375. t. 11 and 12
(1848) (thoracical gland of larva) ; Kirtl., Aw. Ent. Soc. Loml. (2). i. p. 101 (1851) (larva on
Vmleltiferue, incl. of C'iinta rirvsa \) ; Harris, I.e. ed. ii. p. 231 (1852); Urban, Camid. Nat.
Geol ii. p. 220. fig. a. b. t. 3. fig. 2. 3 (1857) ; Mi'n^tr., Emim. Corp. Auim. .Uu.s. Petrop., Lip.
p. 4. n. GO (1857) (Mexico) ; Vollenh., Tijdschr. Ent. iii. p. 85. n. 125 (1860) ; Morris, Si/n.Lep.
N. Amer. p. 5. n. 7 (1862) ; Harr., ed. Flint, 7ns. Inj. Veg. p. 265. t. 4. f. 4. (J, 5. ? , 6. 1, 7. p.
(1862) (life history) ; Reak., Proc. Ent. Soc. PhiJad. ii. p. 137. n. 4 (1863) (Chiapas ; syn.
excl.) ; Weidem, ibid. p. 146 (1863) {jMrUm) ; Lintn., ihid. iii. p. 51 (1864) (Eastern N. York,
eggs, larva, pupa) ; Kirkp., /iiV/. p. 329 (1864) (Cleveland, Ohio, common) ; FeUlcr, Yerh. Znol.
Bol. Ges. Wien xiv. p. 315. n. 361 (18G4) (partim ; Canada to Nicaragua) : Edw., Proc. Ent.
Soc. Philad. iv. p. 390 (1865) (hermaphrodite) ; Tenney, Man. Zool. fig. 281. 282 (1867) ; Reed,
Ginnd. Ent. i. p. 19 (1868) (London, Ont.) ; Riley, Amer. Enlom. i. p. 58 (1868) ; Harris, Ent.
Corr. p. 270 (1869) ; Bethune, Canad. Enl. ii. p. 8 (1870) (Toronto, July) ; Parker, Amer.
Eiitoin. i\. p. lib (1870) (Iowa); Riley, Canad. Ent. iv. p. 37 (1872) (Peterboro Co., Ont.,
May) ; Scudd., ibid. iv. pp. 74, 84 (1872) (Abbot's MS.) ; Edw., ibid. v. p. 8 (1873) (name to be
retained!); id., Proc. Cat. Ac. Sc. v. p. 163 (1873) (larva descr. after Boisd.; Marin Co.,
Oakland) ; Treat, Atner. Natural, p. 129 (1873) (controlling sex, experiment) ; Bean, Eul. Mo.
Mat;. X. p. 248 (1874) (Galena, III., common, June and again midsummer) ; Mend, in Wheeler,
Sept. Ej-pl. Sun: V. Zool. 8. p. 740 (1875) (Colorado ; N. Mexico ; " California,'' error) ; Moore,
Canad. Ent. vii. p. 60 (1875) (Fulton Co., Ohio, July, larva) ; French, Trans. Dcpt. Ar/ric
Jllln. XV. p. 137 (1877) ; Saund., Jiepl. Ent. Soc. Ontario p. 37. fig. 8 (1877) ; Pagenat., Verh.
Nat. M(d. Ver. Heidelh. (2). i. p. 87 (1874) ; Edw., Trans. Amer. Ent. Soc. vi. p. 10. n. 12 (1877)
(Atlantic to Pacific ; Canada to Gulf of Mexico ; Arizona) ; Worth., Canad. Ent. x. p. 17 (1878)
(?,with "fungus" pollen!); Drury, Cincinnati Soc Nat. Hist. i. p. 12 (1878) (Cine,
common) ; Edw., Canad- Ent. xi. p. 86 (1879) ("Costa Rica " false ; black ^ g) ; BalUrd,
lus. Lives p. 321. fig. (1879) ; Oberth., Et. d'Ent. iv. p. 69. n. 197 (1880) (Florida ; Mexico) ;
M.ddl., Trans. Dept. Agrie. Illin. xviii. App. p. 74 (1880) ; Coq., ibid. p. 173 (1880) ; Olliff,
Proc. Ent. Soc. Land. p. 28 (1881) (abnormal neuration) ; Newm., Proc. Ent. Soc. Philad. \.
p. 26 (1881) (N. Jersey ; on carrot) ; Edw., Papdio iii. pp. 54, 60. t. 1. fig. 1. 2. 3. ocellus (188,3)
("Panama" false) ; Fern., Butt. Maine p. 23 (1884) ; Edw., Canad. Ent. xvi. p. 115 (1884)
(habits of larva) ; Gruber, Jena. Zeilschr. Nut. xvii. p. 467. t. 7. fig. 1-5 (1884) (metam.) ; id.,
Papilioiv. p. 84. t. 1. fig. 1-5 (1884) (transf.) ; Lintn., ibid. p. 136. n. 2 (1884) (Rio Grande) ;
Aaron, ibid. p. 172 (1884) (S. Texa^ ; Hagen, Ent. Mo. .Mag. xx. p. 169 (1884) (hunted by
Auar longipes); Tepper, Ent. Amer. i. pp. 159, 186 (1885) (variability of ocellus); Christy, Ent.
Mo. Mag. v. p. 278 (1885) (attr. by faded leaves) ; Mayn., Bull. N. Eng. p. 51. t. 6. fig. 71. 1\.K
(1886) ; French, Butt. East. U.S. p. 89. fig. 14. 15. 10 (1886); Riley, Insect Life i. p. 161 (1888)
(parasites. Tragus obsidianator and c.resorius) ; Weed, Psijche v. p. 52 (I8>'8) (larvae in July,
Champaign, Ohio) ; Hagen, ibid. p. 305 (1888) (caught by Anax) ; Skinn. & Aar., Canad. Ent.
xxi. p. 126 (1889) (Philadelphia, common); Edw., Bull. U.S. Nal. .\Ius. xxxv. p. 10(1889) (liter,
rel. to metam.) ; Mayn., Man. N. Amer. Bull. p. 8. n. 12. fig. 6. b (1891) ; Kent, Insect Life iii.
p. 338 (1891) (Roxie, Mi.'ss.); Edw., Cauad. Enl. xxiv. p. 49 (1892) (Colorado, black (J); Foster,
ibid. p. 192 (1892) (Marshall Pass, Colorado, 10,000—13,000 ft.) ; Staley, ibid. p. 204 (1892)
(Marshall, Missouri, common) ; Weed, ibid. p. 277 (1892) (I-s.iquena Co., Mis.sissippi) ; Davis,
Journ. N. York Ent. Soc. i. p. 47 (1893) (Staten I., N.Y., May to Oct.) ; Skinn., Ent. Neu.-s iv.
p. 82 (1893) (N. Carolina) ; Jones, ibid. p. 190 (1893) (Richmond Co., N.C.) ; Cockerel!, Trans.
( 545 )
Amer. Ent. Soc. xx. p. 353. d. Cii) (1803) (Colorado) ; Beutenm., Bull. Amer. Mus. N. II. v.
p. 242. t. 2. f. 1. cJ (1893) (N. York ; descr. of 1., p.. i.) ; White, Eni. News v. p. 175 (1894)
(Brooklyn) ; Riley, Inncct Life vi. p. 211 (1894) (larva on celery) ; Ehrm., Camd. Enl. xxvi.
p. 292 (1894) (specimen without pupil in ocellus ; larva variable) ; Eimer, Arth. Vunaiiullsrh.
Schmett. ii. p. 127. t. 7. fig. 10. ? (1895) ; Clevel., Ent. Neics vii. p. 73 (1896) (Oneonta, N.Y.) ;
Fiske, ibid. p. 241 (1896) (Webster, N.H., scarce, formerly common, two or three broods) ;
Trum., ibid. p. 298 (1896) (Volga, S. Dakota, common) ; Wright, Butt. West Coast ed. ii. p. 89.
n. 29. t. 29. S, 29b. 29 ? (1906) (East. States ; Arizona).
Pajnlin aslerius var. ampliuta Mcnetrii's, Enum. Corp. Anini. ilus. Petrop., Lip. ii. p. 99. sub n. 60
(1857) ("Amer. sept, par Motschulsky" ;— doubtless from Mexico, where Motsch. had made
collections).
PapiUo astcrioides Ueakivt, Proc. Ac. Nat. Sci. Pliiliid. p. 331. n. 27 (1866) (Mexico); Kirbv, /.c.
p. 567. n. 325a (1871).
Papilio cah-erleyi Grote, Proc. Ent. Soc. Philad. ii. p. 441. t. 10. J (1864) (New Lots, Queen's Co.,
Long I., Aug.).
PapiUo (var. ?) calvcrlcyi, Mead, Amer. A\itural. p. 332 (1869) ( 9 , Florida, April ; abdom. with six
rows of yellow spots).
Papilio pohjxenes, Kirby, Cat. Diuni. Lrp. p. 566. n. 325 (1871) (partim) ; Grote, Bull. Buffalo Soc.
N. Sc. i. p. 185 (1873) (in the Southern States the (^ pohjxenes seems to approach the ordinnry
? type);Gerh.,.l/acTO-i(|;). N. Amer. p.25.n.451 (1878); Auriv., K. Sr. Vet. Ak. IIaiidl.x]x.5.
p. 178. n. 23 (1882) (recensio critica ; — cit. ex parte ad formam insularem refer.); Stand.,
Exot. Tag/, p. 18. t. 12. ^ (1884) diarlim) ; Scudder, Butt. East. U.S. ii. p. 13.53. t. 8. fig. 2. 3,
t. 27. fig. 3, t. 35. fig. 30, t. 40. fig. 1, t. 57. fig. 2, t. 01. fig. 15. 16, t. 66. fig. 2, t. 72. fig. 11.
t. 76. fig. 17. 24. 27, t. 79. fig. 56—60, t. 85. fig. 13. 18. 19 (1889) ; Godm. & Salv., Biol. Centr.
Amer., Lep. Wiop. ii. p. 242 (1890) (partim ; Mexico to Nicaragua) ; Soule, Psijclie viii. p. 435
(1899) (colour var. in larvae).
Aniaryssus poli/.reiies, Scudder, Proc. Boston Soc. N. II. xvii. p. 90. D. 18 (1874) (Heart R. Crossing,
Yellowstone) ; Sprague, Psyche ii. p. 257 (1879) (WoUaston, Mass., May 18.) ; id., I.e. p. 259
(1879) (Mass., May 18 , June 19.).
Papilio asterias var. calrerteyi, Edwards, Trans. Amer. Ent. Soc. vi. p. 10. sub n. 12 (1877) (occas..
Long I., Florida) ; Eimer, Arth. Vericandtsch. Schmett. ii. p. 131. t. 8. fig. 5. (J, 6. ? (1895)
(cop. from Edwards); id., Orlhogen. p. 31. fig. 9 (1897) ; Holland, Butt, Book p. 314. t. 41.
fig. 6. ? (1899).
Papilio asterias var. asteroides (!), Edwards, Trans. Amer. Ent. Soc. vi. p. 10. sub n. 12 (1877)
(S. States ; Arizona ; occas. in Northern States).
Papilio polyxenes var. ampliata, Gerhard, I.e.
Papilio 2'oly.venes var. calverleyi, id.. I.e.
Pajiilio asterius ab. calverleyi, Strecker, Butt. Moths N. Amer. p. 72 sub n. 17 (1878).
Papilio asterius var. astcrioides, id., I.e.
Papilio asteroides (I), Edwards, Canad. Ent. xi. p. 85 (1879) (Reakirt's insect not the same as
Strecker's ; black cj (J, " Costa Rica" false) ; Skinner, Ent. News xiii. p. 183 (1902) (type " so
marked "in coll. of Amer. Ent. Soc. \% jiobjxenes, not the insect fig. by Strecker as asteroides) ;
Wright, Butt. West Coast ed. ii. p. 89. n. 30. t. 4. fig. 30. 30b. ^J. ? (1906) (Mt. Shasta;
Mogave Desert).
Papilio polyxenes caloerleyi, Scudder, Butt. East. U.S. ii. p. 1355 (1889).
Papilio asterias astcrioides Maynard, Man. N. Amer. Butt. p. 8. n. 12a (1891).
Papilio asterias var. ? , alunnta Skinner & Aaron, Canad. Ent. xxi. p. 126 (1889) (Philadelphia ? ;
submarginal spots of hindwing vestigial above, olive-buff).
Papilio astyunax, Scudder (non Fabr., 1793, err. det.) Psyche, viii. p. 210. t. 5. f. 6, /. juv. (1898).
Papilio asterias ab. calverleyi, Spengel, ZtoL Jahrb. Aht. Syst. xii. p. 356. fig. C. D. (1899) ; id.. I.e.
xiii. p. 205 (1900).
Papilio polyxenes Fabr, var. curcifascia Skinner, Enl. News xiii. p. 183 (1902) (Rincon, N. Me; ioo).
Papilio asterias, Fabr. var. serni-alba Ehrmann, Canad. Ent. xxxii. p. 348 (1900) (J' , S. W. Penn.).
There is an interesting difference in the degree of variability between the
specimens from the Nearctic Region jirojier and tlioso from the Central American
countries. While the North American males, with rare exceptions, conform more
or less closely to one type, there are three distinct-looking tyi)es in the southern
districts of the range, two, or in some [daces all three, occurring promiscuously
together, one of them not being distinguishable from the ordinary North American
( 546 )
type. The tliree forms intergrade com])lctel}-. The females from Central America
are on the whole the same as North American ones. The development in Central
America evidently tends towards a black type; the insect is still nnstable, the
causes which are producing the black type not taking effect in all individnals.
It is one of the numerous cases where an insect is on the point of splitting up
into some well-marked geographical varieties. We emphasize nomenclatorially the
occnrrence of the kind of variability mentioned by recording the three types of
males nnder three names.
a!. c?-f. asterius Cram., I.e. Both wings with a yellow discal band of spots
which enters the cell on hindwing. This is the most widely distributed form
occurring from South Canada to South Me.xico. The black dot in the anal ocellus
is rarely absent. Ehrmann, I.e., records as semialba 2 Si from Pennsylvania
in which the spots of the forewing are pure white, while the markings of the
hindwing are deep golden yellow. '
6'. <S-i. cia-i-i/aseia Skinu., I.e. Shorter winged and shorter tailed than the
preceding ; discal band broad, but not entering the cell of the hindwing, or there
being only a small spot in the cell. This is a form occurring in New Mexico,
Vera Cruz and Guatemala, being in the first and last mentioned countries the
prevailing if not the only form of the male. Some North American males
come very close to it, and there are also West Mexican specimens which arc
scarcely distinguishable. Standinger's figure of P. pob/xenes, I.e., represents
the present form.
c'. cJ-f. ampliata Men(5tr., I.e.; asterioides Reak., I.e. Discal spots of
fore- and hindwing, above, strongly reduced, partly obsolete, often all absent from
forewing. This black form is more common in Guerrero than the eJ-f a.iteriu^,
intermediate specimens being about as plentiful as S-l ampliata. Reakirt's
description of asterioides applies to specimens with small spots on the forewing.
The type sj)ecimeu of P. asterioides preserved in the American Museum is said by
Skinner to be "simply an inconstant and accidental variation, and a collector could
take a dozen equally as aberrant wherever the species is common." We have not
seen a single specimen from North America which agrees with Reakirt's description.
The only instance of the occurrence of a black male similar to the female within
the United States we know of is recorded by Edwards, Canad. Ent. xxiv. p. 49
(1892), who bred a male of that form from a Colorado chrysalis.
The female does not vary so much as the male ; the majority of specimens have
small yellow discal spots on the upperside, in many individuals these spots are
nearly all missing, while in others again the spots are large. The females received
from Rincon with the specimens of cJ-f. curvifascia are described by Skinner as
being like the males, but having the spots on the forewing lighter in colour.
We have a female from the same place (received from Mr. G. Franck, of Brooklyn) in
which the discal spots of the forewing are obsolete except the upper two or three,
the middle spots of the hindwing being also much smaller than in the males
from that place. A bred female from Iowa, July 1st, LSOT, killed too soon after
emergence from the chrysalis, the wings being somewhat crinkly, agrees with ordinary
specimens of S-t asterius, but the spots are paler and are irregularly stained with
black, the band having the appearance of being smeared over with black here and
there. A specimen in which the submargiual spots of the hindwing are reduced and
of a bluish colour (ab. alunata) has been named by Skinner & Aaron, I.e. A rare
ttberration common to both sexes is ab. cahcrleyi Grote, I.e., which is well known
( 547 )
from Edwards' figures. We have a transitional specimen (?) caught at Passaic,
New Jersey (acquired for ourselves by Mr. G. Frauck). The forewing is nearly the
same above as below, the spots being smaller and the discocellular bar being
vestigial. The upperside of the right hiudwing is normal, while on the left wing
the first submargiual spot and the anal one are enlarged, and the other submarginal
spots produced discad, this additional yellow scaling, however, not being conspicuons.
On the nnderside the S2)ecimen agrees fairly well with calverleyi.
Hah. of P. pohjxenes asterius : Honduras to Arizona and Canada, in North
America from Arizona and the Mississippi basin to the Atlantic ; a black male
recorded liy Wright from North California.
In the Tring Museum 170 SS, 110 ? ?, and a series of larvae and pupae from
Guatemala northwards.
In coll. Oberthiir from Honduras.
((. P. polyxenes polyxenes Fabr. (1775).
Papilio Eqiies Trojamm poli/.cenes Fabricius, Syst. Ent. p. 44-i. n. 10 (1775) (America) ; Goeze, Eiit.
Beytr. iii. 1. p. 41. n. 7 (1779); Fabr., Sjm: Iiis. ii. p. 4. n. 13 (1781) (ia Araericae meridionalis
insulia).
Pajillin Eqiies Trojanua askrms (!), Fabricius, Mant. Ins. ii. p. 2. n. 13 (1787) {jvtrtiin \=jmJyxenes) ;
Gmelin, Siixt. Nat. i. 5. p. 2228. n. 280 (1790) {jiarlim ; in insulis Americae meridionali oppositis).
Papilio askrias, Boisduval & Lee, Hist. Gen. Lip. Amir. Sejtt. p. 14 (1833) {pin-tim\ Antilles); Boisd.,
Spec, Gill. Lip. i. p. 332. n. 175 (183li) {partim ; Antilles) ; Poey, Mem. Soc. R. Econ. nobitiia
p. 235 (1846) : Lucas, in Sagra, Hist. Cuba vii. p. 205 (1857) (partim) ; Weidem., Proc. Ent.
Soc. Philad. ii. p. 146 (18C3) (partim) ; Felder, Verh. Znol. Bot. Ges. Wien xiv. p. 315. n. 361
(1864) (partim ; insular specimens have wider band).
Papilio arteritis, Herricli-Sch., Corr. Bl. Zonl. Mm. Vcr. Reyensb. p. 172. n. 4 (1864) (Habana) ; Dew.,
Zeitschr. Ge.i. Naturu\ Iii. p. 158 (1879) (Cuba, larva) : iieum. , Eiit. Amer. i. p. 160 (1885)
(Cuba, ? resembling (J) ; Smith, ibid. (1885) (relationship between asteriux, asterioides, and
2)olyxeiies).
Papilio piohjxeiiex, Kirby, Cat. Diiirn. Lep. p. 566. n. 325 (1871) (partim) ; Gundl., Papilio i. p. 113
(1881) (Cuba) ; id., Contr. Ent. Cuba p. 136 (1881) (piartiin ; Western Cuba); Auriv., K. Sv. Vet.
A/.: Handl. xix. 5. p. 178. n. 23 (1882) (partim) ; Godm. & Sa.lv., Biol. Cenlr. Ainer., Lep. Shop.
ii. p. 242. n. 81 (1890) (partim ; Cuba) ; Christ, Mitt. Schweiz. Ent. Ges. ix. p. 271 (1897) (diff.
from asterias).
Papilio asterioides, Eimer {non Reakirt, 1866, err. det.), Arlb. Verwundtsch. Schnieit. ii. p. 121.
t. 7. fig. 6. cj, 7. ? (1895) (" Mexico " false) ; id., (Mhogeii. p. 36. 42. fig. 16 (1897) (" S. Amer.").
There is nothing in the description given by Fabricius in 1775 which points to
the insect described being the present insular form ; but ia 1871 it is stated by him
that the insect came from South American islands. As in the case of his Sphinx
lusca, where the same locality is given, we apply the name to the Cuban form of
the species.
£??. Very similar to ordinary North American specimens of P. p. poli/xencs,
the discal band of the hiudwing broader; the sexes less dissimilar, the female
bearing a discal band almost like the male, the band being narrower and somewhat
paler ; submarginal spots of underside rather larger than in P. p. polyxenes.
Elmer's figures are a good representation of the two sexes of this insular form.
"We suspect that Eimer, as in the case of P. p. amerieus, trusted implicitly in the
correctness of the names under which he received the specimens from dealers.*
Ilab. Cuba.
The specimens labelled S. Domingo (Tweedie) in the British Museum and
* See p. 417.
(548)
in coll. F. D. Godman agree with tlie North American form ; we have not seen
fresh material from this island.
In the Tring Mnsenm, 4 c? J, 2 ? ? . from t!iiba.
(I. P. polyxenes brevicauda i-^auuil. (1868).
Papllioaiileriiix,Gosse (//wi Cramer, 1782. err. det.), Camil. Naliii: p. 184 (1840) (Newfoundland).
PiipiVio hrerkauiUi Saunders, in Packard, Guide I/ix. p. 278 note (1868) (Newfoundlind) ; Ivirby,
C'ul. Diurii. Lep. p. 507. n. 32."jb (1871) ; Saund., Caii-id. Eiit. v. p. 117 (1871!) (reply to Strecker's
attack) ; Grote, Bull. liaffalo Sue. N. Scl. i. p. 185 (187:J) (Anticosti) ; Edwards, Ciiiiiiil. Ent.
ti. p. 20 (1874) (larva); Couper, iliil. vi. p. .3.3(1874) (localitiea) ; Cow^ev, ibid. vii. p. 18
(1875) (Perci', distr. of Gaspi', north shore of Gulf of St. Lawrence) ; Bates, Ent. Mn. May.
xi. p. 244 (1875) (Belts Cove and Terra Nova River, Newfoundland, evidently local form
of aslerias) ; Edw., Butt. X. Aiiirt: ii. Puji. t. 8 (1875) ; id., Tninii. A)nei: Kill. Snr. vi. p. 10. n. 11
(1877) (Anticosti ; Newfoundland ; Quebec ; = ant!coxtieiiiii>^) ; Kirby, I.e. p. 812. n. 325b
(1877) ; Gerh., Mncm-Lep. N. Amer. p. 25. n. 452 (1878) ; Edw., Butt. N. Amer. ii. Pap. t. 8b
(1880) (transf.); Gosse, C'««orf. Enl. xv. p. 44 (1883) (Newfoundland, transformation); Gruber,
Zeitschr. Ges. Naturu: xvii. p. 4(18 (1881) ; id., Papilio iv. p. 85 (1884) (transf., after Edw.) ;
Scudd., Biitl. East. U.S.A. iii. p. 1851 (1889) ; Edw., Bull. d.S. Xat. .I/u,s-. xxxv. p. 10 (1889)
(liter, rel. to transform.) ; Mayn., Man. N. Amer. Bull. p. 8. n. 10. fig. 6c (18'J1) (Newfound-
land ; Anticosti; Labrador; Quebec, June); Elmer, AM. Verwandtseh. Schmetl. ii. p. 13fl.
t. 7. fig. 2. (J (1895) (copy from Edwards) ; Christ, Milt. Srhtveiz. Ent. Ges. ix. p. 272 (1897)
{■= piilijreiieis); Winn, Camid. Ent. xxx. p. 304 (1898) (Kamouraska, 85 miles south of Quebec);
id., Bepl. Enl. Sue. Ontario xxix. p. 36 (1899) (range, life hist. ; 85 miles south of Quebec, on
south shore of St. Lawrence; Bic, Rimou,-ki Co.; Orleans I., P.Q. ; Metis; Kamouraska;
all stag, s in August, on .ircluitiiielica; probably two broods); Fyles, iljid. p. 45 (1899) (pupation);
Bethune, ibid. xxx. p. 104 (1900) (first specim. May 23 in breeding cage); Dyar, Bntl. U.S. N<il.
Miis. Iii. p. 3. n. 21 (1902) (N.E. Coast) ; Lyman, Canad. Ent. xxxv. p. 340 (1903) (larva on
parsley and other Umbellif .) ; Brain., ibid, xxxvi. p. 52 (1904) (rearing) ; Ljman, liipt. Ent.
Soe. Ontario xxxvi. p. 96. n. 21 (1905) (N.-W.-River Post, Hudson Bay; Lake Melville, Ungava).
Papilio polygenes var. brcricauda, Couper, Caniid. Enl. iv. p. 202 (1872) (Anticosti).
Papilio antirostien.ils Strecker, Lep. Bhop. llrl. p. 10. t. 2. fig. 2 (1873) ; id., I.e. p. 41 and 49 (1873);
id., /.c. 68. t. 8. fig. 13. larva (1874) ; Gerh., Macro-Lep. N. Amtr. -p. iii.n. ih^ {\%1%)\ Streck. l.e.
Siippl. iii. p. 17 (1900) (2cJ (J, 2 ¥ ? ).
Papilio asterius var. brevicauda, Strecker, Bull. .Uolhs N. Amer. p. 71. sub n. 17 (1878).
Papilio asterius var. antieostien'^is id., I.e.
Papilio medioeanda (!) Eimer, I.e. ii. p. 119 (1895).
Papilio medioeanda Eimer, I.e. ii. p. 138 (1895) (hab. V, this form or aslcriiin (J-f. eurvifaseia'^).
c? ?. Sexes similar. Distal margin of forewing rounded; yellow bar on eross-
veins always present ; no cloudy yellow spot in front of subcostal fork ; discal spots
often all of about the same length. Discal band of hindwing curved, usually
no spot in cell, the spots often somewhat orange distally ; tail shorter than in
average specimens of P. p. asterius.
For early stages see Edwards, I.e. (1880), and Strecker, I.e. (1874).
Ilab. Newfoundland : Anticosti ; Gulf of St. Lawrence ; and neighbouring
districts, the exact range not being known.
In the Tring Museum 11 c?c?, 8 ? ?, from: St. John's, July 1898 (James);
Baie St. Claire, Anticosti.
01. Papilio bairdi Edw. (1869).
PapUio haiydiEd\ya.rds, Proc. Enl. Soe. P/nlad. vi. p. 200 (1869) (Arizona) ; Kirhy, Cat. Diurii.
Lep. p. 567. n. 331 (1871) (" Mexico " false) ; Mead, in Wheeler, liepl. ICrpl. Surv. v. Zool. 8.
p. 740 (1875) (New Mexico ; Arizona) ; Edw., Trans. Amer. Ent. Soe. vi. p. 10. n. 10 (1877)
(Arizona) ; Gerh., Maero-Lep. N. Amer. p. 25. n. 447 (1878) (N. Mexico) ; Strecker, Butt.
Moths N. Amer. p. 72. n. 17a (1878) (Arizona); Edw., Canad. Ent. xi. p. 83 (1879)
( ¥ ; variability ; = var. ulahensis) ; id., Bull. N. Amer. ii. Pap. t. 10. (J ¥ ■ (1880) (Arizona) ;
Mayn,, Man. N. Amer. Butt. p. 8. n. 11. fig. 6a (1891) (Arizona) ; Edw., Canad. Enl. xxiv. p. 50
( 549 )
(1892) (W. Colorado) ; id., I.e. xxv. p. 253 (1893) (nregmiia bred from bnirJi and the reverse) ;
Haase, fJiilpisurh. Mimicry i. p. 92 (1893) ; Edw., Caiiail. Eiil. xxvii. p. 229 (1895) {oregonia
bred from eggs of bairdi, and the reverse) ; id., I.e. p. 241 (1895) (Sioux Co., Nebraska) ; Eimer,
Arlb. Vei-Lm.iidtsch. SchinHt. ii. p. 118. t. 7. fig. 1. (J, 9. ? (1895) (copies from Edw.); id.,
0)-W(o,-/(». p. 3G. fig. 15. (J, 18. ? (1897); Christ, Mitt. Srhweis. Enl. Gcs. ix. p. 272(1897);
Beutenm., Joiirn. N. York. Ent. Stic. T. p. 101 (1897) (var. of nreijonia, not of aslcrias) ;
Edw., I.e. XXX. p. 11 (1898) (? bairdi produced urrgonia (= brucei) and bairdi); Holland,
Butt. Book p. 313. n. 14. t. 40. fig. 2. (^ (1899) (Arizona northwards) ; Brown., Ent. News xii.
p. 301 (1901) (Salt Lake City, usually rare) ; Dyar, Bidl. U.S. Nut. Mus. Hi. p. 3. n. 17 (1902).
Papilid luppocraies var. oregonia Edwards, Trans. Anier. Ent. Soe. v. p. 208 (1871!) (Colombia R., ? ).
Papilio (asleriiis) var. idahensis Strecker, Lep. Rhop. Uet. p. 128 (1878) (Utah).
Papitio asterius'^AV. utnbensis id. ^ Jltdt. Jloths N. Ainer. p. 72. sub n. 17(1878) ; id., Lep. Rhop. Ilet.j
Sujiji!. iii. p. 17 (1900) (Utah, 2 ^ ^, I '} ; var. of bainli , one (^ with abdomen spotted, the
other as in mnchaon).
Papilio oregonia 'Edwavdn, Putt. N. A nier. ii. Pap.t.7. J J (1880) (Oregcm); Stretch, PapHio ii. p. 119
(1882) (Washington Terr., larva on Artemisia, descr.) ; Edw., ibid. iii. p. 5(5. t. 1. fig. 6. 7. 8 .
ocellus (1883) (distinct species); id., Butt. N. Ainer. ii. Snppl. p. 1 (1884); Mayn., .l/iitt. N. Amer.
Butt. p. G. n. 4 (1891) (Oregon ; Wasliington ; Vancouver's I.) ; Edw., Canad. Ent. xxiv. p. 52
(1892) (Utah ; West Colorado) ; id., I.e. xxvii. p. 241 (1895) (Sioux Co., Nebraska; S.E.
Wyoming ; Pullman, Washington ; Idaho) ; Christ, I.e. ix. p. 273 (1897) ; Elwes, Pi-oc. Ent.
Soe. Loud. p. 11. (1897) (= machaon) ; Wright, Butt. West Coast ed. ii. p. 87. u. 26. t. 3. fig. 26. ?
(1906) (WaJiington).
Papilio machaon, Hagen, Canad. Ent. xiv. p. 178 (1882) (Washington Terr.).
Pajiilio oregmiius (!), id., Papilio ii. p. 150 (1882) (cannot be separated from zolicaon) ; Haase,
Untersach. Mimicry i. p. 92 (1893).
Papilio oregomis (!), Hagen, Psyche iii. p. 415 (1882) (Washington Terr., " = zolicaon ").
Papilio hfjUandi Edwards, I.e. xxiv. p. 50 (1892) (W. Colorado) ; Holland, Butt. Bonk p. 314.
n. 16. t. 40. fig. 3. J (1899) (Arizona; Colorado).
Papilio brucei Edwards, Canad. Ent. xxv. p. 253 (1893) (name for oregonia from Colorado) ; id., I.e.
xxvii. p. 239 (1895) (Colorado ; '■ nov. spec") ; id., Butt. N. Amer., Pup. iv. (1897) (ic«(;«i is
result of hybridism between P. oregonia and bairdi ! ; life history, results of breeding) ; Elwes,
I.e. p. 11 (1897) (grad.ition from oregonia through brucei to zolicaon) ; Fletch., Rept. Ent, Soe.
Ontario xxxi. p. 56 (1900) (Regina, Canada ; aho in the Kootenay Mts. at Kaslo) ; Burr., Enl.
News xii. p. 244 (1901) (Yellowstone Nat. Park).
Papilio machaon oregonia, Eimer, Artb. Verwandtsch. Schmett. ii. p. 109. t. 6. fig. 2 (1895) (copy from
Edw.).
Papilio bairdi oreejonia, Dyar, I.e. (1902).
Papilio bairdi brucei, id.. I.e.
Papilio bairdi hnllandi, id., I.e.
d ? . Tricbroinatic in both sexes, at least in certain districts.
«'. f. bairdi Edw., I.e. Similar to P. polijxencs astcrius, the sexes differing
in a similar way as iu the ordinary form of that insect. The discal spots are paler
on the underside than m astffiu.s, and gradually shade off proximally ; size of these
spots very variable. Many specimens with vestige of yellow lateral stripe
posteriorly on abdomen.
b'. f. hollandi Edw.,/.(,'. Like the preceding, but the abdomen yellow, striped
with black, as in the next form.
c'. L oregonia Edw., I.e.; hrucci id., I.e. Similar in appearance to
P . machaon \ anal ocellus pnpilled as in the preceding forms. Colorado specimens
{brucei) are said by Edwards to be different from Oregon individuals (oregonia), but
we fail to find any constant distinction, the differences given by Edwards not at all
holding good.
This yellow form doe.s not occur in Arizona, the black f bairdi being the only
one found there, while the black form has not been met with in Oregon. Polj--
chromatism of a species in one district and nionochromatism of the same species in
another country is a phenomenon often met with among insects. Edwards had
( 550 )
probably overlooked this fact when he advanced the hypothesis that the variable
Colorado insect was the product of a cross between a black (southern) species,
P. bain/i, and a yellow (northern) species, P. oregonia. A parallel case is P. chjtia
of the Oriental Region. The variability of P. poh/xencs is also similar to that of
/'. bain/i, that species being strongly di- or trichromatic in Mexico, Guatemala, and
South America, practically monomorphic (apart from occasional aberrations) in the
other districts of the range.
For early stages see Edwards, /..:•. (1807).
Ilab. Arizona and New Me.xico (Jii7e Mead) northwards, the yellow form
extending to Uaunda (Regina) and British Columbia (Kootenay Mts.), the black
form not being known so far north (or is it represented by P. nitra ?).
In the Triug Museum 20 c?c?, 3 ¥ ?, from: Glenwood Springs, Colorado,
June 1901 (Oslar); Thumb Bntte, June 1901 (Oslar) ; Garfield Co.; Beaver, Utah,
July ; Wickenbnrg, Arizona, May 1898 (Dr. Kunze) ; Ozoyoos, British Columbia
(Reynolds).
62. Papilio nitra Edw. (1883).
Papilio nilm Edwards, Papilio iii. p. 158. 162 (1883) (Judith Mti., Montana) ; Fletcher, Camd. Ent.
xix. p. 22.i (1887) (Rocky Mts. ; Regina, N.W.T.) ; id., Repl. Ent. Soc. Ontario xviii. p. 25
(1888) (Regina, N.W.T. ; Rocky Mts.); Edw., Bull. N. .imei: iii. Pap. t. 1. (J? (1880)
(Canmore, June, on the summit ; Regina ; Montana, July) ; Mayn., .Man. N. Amer. Bull. p. (5.
D. 7 (ISiO) (Montana) ; Christ, MittSrhweiz. Ent. Ges. ix. p. 27.S (1807) ; Holland, Bull. Bool:
p. 312. n. 11. t. 41. fig. 2. (J (1809); Dod, Catiad. Eiil.x%xm. p. 171. n. 80 (1901) (Alberta,
June) ; Baird, Jiepl. Ent. Soc. Ontario xxxiii. p. 93 (1903) (High River, Alta).
Papilio intira nitra, Dyar, Bull. U.S. JVat. Mm. Iii. p. 3. n. 20a (1902).
This insect resembles P. bairdi f. bairdi, but is shorter winged. It is restricted
to Montana and Western Canada, and is said to occur there together with
P. zelicaon and P. bairdi f. oregonia. We believe these oregonia to be the
yellow form of nitra. We have not seen Canadian oregonia, but have two females
from Ozoyoos, British Columbia. These females are shorter winged than our
Colorado specimens and than Edwards's figures of Oregon individuals, and have
the abdominal margin of the hindwing more extended black. (Janadian oregonia
may be similar to these Ozoyoos specimens, which would render it probable that
there is really such a connection between the black and the yellow Canadian
specimens as here suggested. Breeding will decide the question. P. indra is
qnite distinct from nitra.
The sexes are similar, the yellow markings being rather paler in the female
than in the male (they are too pale in Edwards's figure). The size of the
markings is variable, also the extent and intensity of orange on the underside of
the hindwing.
Early stages and food-plant not known.
Hab. West Canada ; Montana.
In the Tring Museum 4 c?^, 1 ?, from: Red Deer, Allicrta, June 1003;
Didsbury, Alberta, Jnne 1904.
63. Papilio zelicaon Lucas (1852).
Papilio zelicaon Lucas, in Gui'r., Rev. Zool. (2). iv. p. 13G (1852) (California); Doubl., Wcstw. & Hew.^
Gen. Diurn. Lep. ii. p. 520 (1852) ; Gray, Cat. Lcp. Ins. Brit. Muk. i. Pap. p. 37. n. 182 (1852)
(Calif.) ; id., List Lep. Ins. Brit. ihis. i. Pap. p. 51. n. 190 (185G); Lucas, Bull. Soc. Ent. France
p. G7 (185(>) (distinct from rnachaon) ; Felder, Verh. Zool. Dot. Ges. Wien xiv. p. 315. n. 358
(1804) (California ;— " Labrador ; Missuri ? " .ad aliam spec, refer.).
Papilio zolicaon (!) ISoisduval, Ann. Soc. Ent. France p. 281. n. 3 (1852) (California) ; Vollenh.,
( .351 )
tijdschr. Ent. iii. p. 84. n. 122 (1S60) (S. Fraacisco) ; Morris, Si/n.. Lep. N. Anier. p. 4. n. 5 (1882)
(California); Weidem., Proc. Ent. Soc. Philad. ii. p. 148 (1863) (" Labrador" false ; U. States ;
hardly more than a variety of machaoii) ; Kirby, Cat. Dinrn. Lep. p. 5GG. n. 321a (1871)
(California) ; Couper, Ginad. Ent. vi. p. 34 (1874) ; Strecker, Lep. Rlmp. net. p. 4(3. t. C.
fig. 3. ? (1873) ; Edw., Proc. Cal. Ac. Sc. v. p. IGS (187.-5) (larva, pupa) ; Mead, in Wheeler,
Mept. Expl. Suit. v. Zo,,I. 8. p. 740 (1875) (Colorado ; S. Utah) ; Edw., Butt. N. Amei: ii. Paji.
t. 6. (J. ?. 1. p. (18751 (Vancouver's I. to Lower California ; Arizona ; Colorado ; Montana ;
Idaho) ; Kirby, /.'■. p. 812. n. 321a (1877) ; Edw., Trans. Amer. Ent. Soc. vi. p. 10. n. 7 (1877)
(Oregon to Arizona ; Montana ; Colorado) ; Gerh., ihxcro-Lep. N. Amer. p. 25. n. 44!) (1878);
Streclfer, Butt. Motlia N. Amer. p, 71. n. 14 (1878) ; Oberth., Et. d'Ent. \v. p. G8. n. 194 (1880)
(Calif. : Canada) ; Hagen, Psyche iii. p. 415 (1882) (Washington Terr., " var. of machaoii ") ;
^dvi., Papilio iii. p. 48. t. 1. fig. 4. 5. ocellus (1883) (distinct from machuon, oreyonia, etc.);
Lyman, PapHio iii. p. 109 (1883) (var. 9 , Nevada) ; Butl., Journ. Linn. Soc. Loud. xvi. p. 472. n. 59
(1883) (Mendocino) ; Edw., PiqiiUo iv. p. 1G2 (1884) (early stages ; mostly one brood only) ;
Behr, Bull. Cal. Ac. Sc. i. p. G4 (1884) (Calif., common, 1. on Ocnanthi, Angelica, Carum); Edw.,
Butt. N. Amer. ii. Stipjd. p. 1 (1884) (S. Bernardino; near asterias) ; id., Bull. U.S. Nut.
Mus. XXXV. p. 9 (1889) (liter, relating to metam.) ; id., Butt. N. Amer. iii. Pap. t. 3. fig. a-g
(1891) (transform.); Eiley, /hsctJ ij/c iii. p. 412 (1891) (parasite: Apnnteles); Mayn., J/«;/.
A". Amer. Butt. p. G. n. 5 (1891); Foster, Canad. Ent. xxiv. p. 192 (1892) (MarshaU Pass, Colorado,
10-13,000 ft.) ; Haase, Untersuch. Mimicry i. p. 92 (1893) ; Oslar, Ent. News iv. p. 22r, (189.3)
(Los Angeles, Febr.) ; Cockerell, Trans. Amer. Ent. Soc. xx. p. 353. n. 647 (1893) (Rosita,
Colorado); Danby, Jo»ra. N. York Ent. Soc. ii. p. 33 (1894) (Vancouver I., scarce) ; Wiley,
Ent. News v. p. 38 (1894) (Miles City, Montana, rare) ; Snyder, ibid. v. p. 1G7 (1894) (Park
City, Utah) ; Jordan, Cannd. Ent. xxvi. p. 257 (1894) (Napa, Calif. ; metam.) ; Cunningh.,
Ent. News vi. p. 251 (1895) (Ft. Klamath, Oregon) ; Eimer, Artb. Verwandtsch. Schmetl. ii.
p. 109. t. 6. fig. 5(1895) (California) ; Walk., Proc. Ent. Soc. Lond. p. 11 (1897) (Vancouver I.,
larva and pupa not distinguishable from those of machaon); Twog., Ent. News viii. p. 31 (1897)
(Riverside, Calif., rare, late Febr. & March) ; Snyder, ibid. viii. p. 1G4 (1897) (Utah) ; Christ,
Mitl. Schwcin. Ent. Ges. ix. p. 270 (1897) ; Holland, Butt. Book p. 312. n. 10. t. 38. fig. 1. ^
(1899) (Vancouver I. to Arizona and Colorado) ; Denton, Maths Butt. N. Amer. ii. p. 350.
fig. S (1898—1900); Dod, Cannd. Ent. .xx.xiii. p. 171. n. 79 (1901) (Alberta, June) ; Brown.,
Ent. News xii. p. 301 (1901) (Salt Lake City, usually scarce, up to 9500 ft.) ; Dyar, Proc. U.S.
Nat. Mtts. xxvii. p. 782 (1904) (Kootenai) ; Dennis, Repl. Ent. Soc. Ontario xxxiv. p. 90 (1904)
(Beulah, Manitoba) ; Wright, Butt. West Coast ed. ii. p. 8G. n. 24. t. 3. fig. 24 (1906) (as far north
as Wrangel, Alaska).
Papllio machnnn, Men^trits, Enuni. Corp. Anim. Mus. Petrop., Lip. i. p. 4. n. 58 (1857) {parlim ;
California).
Papilio machaoii var. californica, id.. I.e., Lep. iii. p. G9. n. 58 (18G3) (" = xelicaon Lucas").
Papitio dolicaon (!), Behr, Stett. Ent. Zeit. xxvii. p. 21G (ISGG) (Calif.).
Ainarijsxus zolicaon, Scudder, Proc. Boston N. H. Soc. xvii. p. 90. n. 19 (1874) (Yellowstone, July 18).
Papilio zdicayn (!), Dyar, I.e. (1902) (sub syn.).
Papilio coUro Wright, I.e. p. 86. n. 25. t. 3. fig. 25. $ (1906) (Colorado Desert, S.E. Calif.).
Tliere may be a closer connection between the present insect and Papilio
nitra than we suspect. But until positive proof by breeding is forthcoming,
l\ zelicaon should be treated as a separate species. Lucas's name zelicaon has
priority over zolicaon of Boisdnval.
The species is more constant than any of the allied forms. However, there
occur promiscuously in a series some inconspicuous but significant deviations from
the ordinary type. The pattern of the abdomen is by no means so constant as
Edwards, I.e. (1883), stated it to be. The broad black dorsal stripe bears
occasionally at its lateral edges on segments 4, 5 and 0 a yellow dot partly
separated from the yellow side-stripe, these dots being homologous of the respective
dots found in both Papilio nitra and P. hairdi f. bairdi. The unilerside of the
abdomen is usually quite black, many individuals, however, bearing posteriorly
on eacli side a vestige of a yellow stripe. These stripes are sometimes quite
distinct, extending almost to the base of the abdomen in some females, there being
occasionally also a thin yellow mesial line ou the posterior segments.
( 552 )
The variabilit)' in size is not inconsiderable, the forewing measuring from
34 to 46 mm. in length in onr males, and from 35 to 50 mm. in onr females.
The clondy sj)ot in front of the .snbcostal fork of the forcwing assumes sometimes
a distinct blue tint ; the black dot within the fork is occasionally very small,
rarely absent ; the yellow cell-bar varies much in size, being vestigial in one of our
numerous specimens from Mount Shasta, a male, the posterior discal patches being
in this individual also rather smaller than usual. The amount of black at the
base and behind the cell of the hindwing is variable ; tlie yellow colour extends
usually to the base of the cell, but the basal fourth or third of the cell is sometimes
black ; the width of the black distal border is quite inconstant. The cell of the
forewing is, in some individuals, distinctly striped with pale yellow; some specimens
have hardly any orange colour on the disc of the hindwing, while others bear
conspicuous orange patches. Deep yellow individuals are ab. coloro Wright.
For early stages see Edwards, l.i'.
Hah. Alaska, British Columbia, Alberta, southwards to Arizona and Colorado.
In the Triug Museum 95 iS, 27 ? ?, from: Qu'Apjielle, Assiniboia, June
19U1 ; Ozoyoos, British Columbia (Reynolds) ; Gold Hill, Oregon, May — July 1901
(Biedermann) ; Quincy, California, 3400 ft., May 1890 (Watson); Trucky, California,
6000 ft. ; McCloud E., Shasta Co., June 1884 (0. T. Baron) ; Butte Creek,
Butte Co., May 1898 (Mrs. Austin) ; Davis Creek, Madoc Co., 4500 ft., July 1898
(Mrs. Austin); N. Tulare R, California, July 1897 (Purpus); Siskiyou Co.,
California (0. T. Baron) ; Reno, Nevada ; Chimney Gulch, Colorado, May 1901
(Oslar) ; Garfield Co. and Park Co., Colorado.
In coll. H. J. Adams from Calgary.
64. Papilio indra Reak. (1867).
? . Papilio indra Reakirt, Proc. Ent. Soc. Philad. vi. p. 123 (1867) (Colorado, Pike's Peak).
S ? . Sexes similar. Body black ; a line bordering the mesothoracic tegula
and extending forward, ending behind antennae, creamy, often somewhat
ochraceous ; abdomen either wholly black, or the last segments laterally creamy,
or there is a creamy stripe of variable width from base of abdomen to claspers,
the stripe being situated laterally on the tergites, the claspers remaining always
black.
The colour of the creamy markings of the wings darkens by e.^posure to
light, as in other pale yellow species.
Forewing. There are usually two creamy bars across the cell, either of
which or both may be wanting ; the discal band is very variable in width, the
submarginal spots also varying much in size ; in fresh specimens the basal third
of the wing is powdered with creamy scales. The discal band of the hindwing
stands either outside the cell, being narrow, or enters the cell, occupying sometimes
as much as the apical third of the cell ; the submarginal spot M' — M- is often
absent ; the anal orange spot (submarginal and marginal spots M' — SM- merged
together) is always centred with black ; the orange ring is usually complete, but
in a small percentage of specimens it is interrupted on the abdominal side, being
occasionally open also on the discal side, in which case the orange submarginal sjiot
stands separate from the marginal one like the other submarginal spots ; the
discal spot R^— M' is acuminate, often also the spot in front of it, both being
somewhat prolonged on the underside, and in many specimens stained with orange
distally.
( 553 )
Genitalia: c?. Saw of harpe longer than, or at least as long as, the proximal
non-serrate portion of tlie harpe ; this non-serrate portion compressed, elevate,
forming a sharp ridge which is highest before joining the saw, being here somewhat
curved ventrad ; the saw itself somewhat cnrved dorsad jiroximally ; the teeth small
and close together. ? not dissected.
Early stages noticed by Edwards, I.e. (1897).
a. P. indra imlra Reak. (1867).
Papilio inlra Reakii-t, I.e. ; Kii'by, Cat Diuni Lep. p. 5G7. d. 3)!7 (1871) ; Strecker, Lrp. Rh'ip. Ilet.
p. 9. t. 2. f . 1 (1873) (Pike's Peak) ; id., Proc. Ac. N. Sc. Philad. xxviii. p. 150 (1876) ( ? , Clear
Creek, Colo., July 1) ; Putn., Proc. Daoeiip. Ac. Nal. Sci.i. t. 35. fig. 5. ? (1876) ; Kirby, I.e. p. 812
(1877) ; Edw., Traiis. Aiuer. Ent. So,: vi. p. 10. n. 8 (1877) (Colorado ; Nevada) ; id. Ball.
N. Amer. ii. Pap. t. 9. (J. ? (1878) ; Gerh., Macro-Lep. N. Anier. p. 25. n. 454 (1878) ;
Strecker, BiM. Moths N. Amer. p. 71. n. 15 (1878) (Colorado) ; Edw., PapiUo iii. p. 2 (1883)
(Boulder, Colo.) ; Butl., Jouru. Linn. Soc. Loud. xvi. p. 472. n. 62 (1883) (Siskiyou Co.) ;
Mayn., JJaii. N. Amer. Butt. p. 7. n. 8. fig. 6. d (1891) (Colorado ; Nevada ; California); Bruce,
aiiiad. Ent. xxiii. p. 110 (18'J1) (Colorado, June, 7000 ft.); id., Ent. News viii. p. 1.34
(1897) (Denver, Colo., May 4); Christ, Mitt. Schweiz. Ent. Ges. ix. p. 272 (1897); Edw.,
Butt. N. Amer. iii. Suppl. p. i. (1897) (egg and larva noticed ; food plant, Artemixiu) ;
Holland, Butt. Book p. 312. n. 12, t. 41. fig. 3. ? (1899) (mountains of Colorajo, Nevada,
California) ; Denton, Moth>< Butt. U.S. Amer. ii. p. 350(1898—1900) ; Snyder, Eid. News xi.
p. 365 (19U0) {Silver Lake, Utah) ; Strecker, Lep. Rhop. Net., Suppl. iii. p. 17 (1900) ( J , Clear
Creek Caiion) ; Brown., Ent. News xii. p. 301 (1901) (Salt Lake City, quite rare, 6000 ft.).
Dyar, Bull. U.S. Nat. Mus. Iii. p. 3. n. 20 (1902) (parliui) ; Wright, Butl. West Coast ed. ii.
p. 87. n. 27. t. 4. fig. 27. S, ^^b. ? (1906).
(??. Tail short.
Hab. California ; Nevada ; Utah ; Colorado.
In the Tring Mnseum 122 SS from: Siskiyou Co., California (0. T. Baron);
McCloud R., Shasta Co., June 1884 (0. T. Baron); Chimney Gulch, Colorado,
May and June 1900 and 1901 (Oslar).
A ? in coll. H. J. Adams.
b. F. indra pci-gamus Edw. (1875).
Papilio pergamus Edwards, Proc. Calif. Ac. Sci. v. p. 423 (1875) ; Kirby, Cat. Diurn. Lep. p.813.
n. 383 (1877) ; Edw., Traus. Amer. Ent. Soc. vi. p. 10. n. 9 (1877) (S. California) ; Strecker,
Butl. Moths N. Amer. p. 71. n. 16 (1878) (California) ; Mayn., Man. N. Amer. Butt. p. 7. n. 9
(1891) (S. California) ; Beutenm., Bull. Amei: Mus. N.H. iv. p. 167 (1892) (type in Amer.
Mus., (J, S. Barbara) ; Christ, Mitt. Schweiz. Ent. Ges. ix. p. 273 (1897) (= indra) ; Wright,
Butt. West Coast ed. ii. p. 88. n. 28. t. 4. fig. 28. ^, 29b. ? (1906) (S. California, 2000—3000 ft.).
t?. Tail longer than in the preceding form.
Bab. South California, coast range.
In the Tring Museum 2 S6,Jnae and July.
05. Papilio machaon L. (1758).
Papilio Eques Ackivus machaon Linnii, Si/sl. Nat. ed. x. p. 462. n. 27 (1758).
This species is represented in America by the following subspecies :
a. Fajjilio machaon alias/ia Scndd. (18G9).
Papilio mut:huou, Edwards, Canad. Ent. i. p. 22 (1868) (Rupert House, Hudson Bay) ; .Strecker,
Butt. Moths N. Amer. p. 70. n. 13 (1878) ; id.. Butt. N. Amer. ii. Pap , in test for t. 7 (1880)
(Dalles).
Papilio aliaska Scudder, Pror. Boston N. 11. Sor. xii. p. 407 (1869) (Nulato, May 20— June 14 ; also
E. coast of Hudson Bay) ; Kirby, Cat. Diurn. Lep. p. 566. n. 321b (1871) ; id.. I.e. p. 812.
n. 321b (1877) ; Gerh., Macro-Lep. N. Amer. p. 25. n. 450 (1878) ; Holland, Butt. Book p. 312.
n. 9. t. 41. fig. 1 (^ (1899) (only in Alaska).
( •«-! )
PajiiUo machaon var. aliaska, Edwards, Papilio ii. p. 75 (1882) (Hudson Bay; Alaska; common at St.
Michael's on coast above outlet of Yukon R. ; Dalles, Columbia E.) ; id., I.e. iii. pp. 58. 60
(1883) ( = HimaIaTan form, false); Webster, Caiiad.Ent. xxvi. p. 117(1894) (Alaska, eastw. to
Hudson Bay) ; Christ, Mitt. Sdnveiz. Ent. Gex. ix. p. 270 (1897) ; Lyman, Oinml. Ent. xxxii.
p. 119 (1900) (Dawson, Yukon) ; Stand. & Reb., Cat. ed. iii. p. 2. n. 4h (19(11) (Alaska).
PajiiUo machann aliasha, Maynard, ^fan. N. Amer. Butt. p. 6. n. 3a (1891) (Oregon northwards) ;
Dyar, Bull. U.S. Nat. Mus. Iii. p. 3. n. IGa (1902) (Alaska ; North Pacific States) ; Wilson, Repf.
Ent. Soc. Ontario xxxiv. p. 90 (1904) (Nagagami R., Hudson Bay slope) ; Keelo, ibid. xxxv.
p. 16 (1904) (common along the shores of Mayo Lake, and valley of Mayo R,, Yukon Terr.,
July & August).
The black band of the hindwing is broader than in P. machaon kamtschadalus
Alph^r. (1897). The anal ocellns is alwajs blind as in the other forms of
P. machaon, the black admarginal sjiot standing at the distal side of the orange
spot, not within it.
The insect is rare in European collections. Sometimes one finds as aliaska
and oi-egonia Old-World specimens of P. machaon in collections. We have received
ourselves from America two machaon, of which one is undoubtedly a Sikhim
individual and the other a British one.
Hub. Alaska ; Oregon ; Hudson Bay.
In the Tring Museum one bad S.
VI. Thoas Group.
Underside of thorax and abdomen not striped with black, being either all
yellow, or black dotted with yellow laterally or bearing a yellow lateral line.
Hindwing beneath from base to disc yellow or black, no black bands Ibrming a
large V ; PC at two-thirds or nearer apex of basal cellule, the latter not much
produced.
Young larvae with dorsal tubercles which are replaced by small circular spots
in later stages ; adult larvae with yellowish side-stripe on thoracical segments, a
very large pale dorsal or dorso-lateral patch occupying the central segments
(V-shaped or mesially divided) and a large side-patch on the last segments tapering
in front, these markings rather ill-defined.
Key to the species :
a. Tail with yellow spot in centre of widened apical portion . b.
Tail without yellow spot in centre on upperside ... e.
b. Forewing beneath without black band across cell . . c.
Forewing beneath with black band across cell continued to
hindmargin ......... <l.
c. Discal patch R-— K' of forewing projecting much lieyond
patch R' — W ; dilated part of tail rounded ; tenth
tergite of $ bifurcate, the projections curving laterad . Species No. 67.
As before ; spatule of tail more elongate ; tenth tergite of
$ short, simple Species No. 08.
Discal patch U-— R^ of Ibiewing very little imiji'cting
beyond patch R' — M' ; cell olten with yellow spot,
striped with yellow and black beneath ; spatule of tail
elongate ; tenth tergite of S long Species No. 00.
il. Yellow band of forewing parallel to distal margin . . Species No. 7:t.
Yellow band of forewing interrn]iteil, both portions oblique,
the posterior portion continuous with the broad cell-bar. Species No. 74.
( 555 )
e. Wings black, with a row of sharply defined snhmarginal
spots, first spot on forewing standing in front of SC^,
being about 5 mm. distant from distal margin ; cell all
black ; liindwing with an orange-red spot R^ — M' close
to cell, sharply defined .
Wings at least partly yellow, or the disc mnch paler than
the base, or hindwing more or less bine on upperside ;
snbmarginal spots of forewing absent from black speci-
mens (which are all ? ?), or tlie spot SO' — SC^ absent or
close to margin ; or rufons-orange spots on underside of
hindwing in a row which is separate from cell
/. A regularly curved row of blue lialfmoons on disc of hind-
wing beneath, preceded by a row of rufous or orange
halfmoons which are distant from cell ....
Hindwing beneath with a row of sharply defined rufous or
orange spots around apex of cell ; no yellow spot in cell
of forewing on upperside ......
Hindwing beneath with au irregular, but complete, row of
metallic blue spots on disc, spot R^ — M' being more
distal than the others ; the rufous red spots preceding
the blue ones ill-defined, often vestigial ; no spot in cell
of forewing on upperside ......
ff. Hindwing with sharply marked yellow submargiual spots
on upperside, at least in <? ; in ? disc not greenish
blue, without large greenish patches ....
Submargiual halfmoons of upperside of hindwing thin,
always washed over with black in c?: these spots blue
or green in ? ; disc also blue or green or huffish green .
L Cell-patch of S on forewing aliove produced basad Ijeyond
point of origin of M-, occupying about one-third of the
cell ; no submargiual spots on upj>erside of forewiug ;
? with carved yellow band from costal to hinder margin
on upperside of forewing ......
S without patch in cell on upperside of forewiug ; on
underside of forewing a row of small spots between
discal band and submargiual spots ; submargiual spots
of upperside of hindwing vestigial in ? , being washed
over with brown ; forewing with yellow markings in
costal area, but no pale shadowy band on disc
cJ with patch in cell of forewing on upperside ; ? with
pale band on forewing or the disc mnch paler than the
area from base to apex of cell, especially ou underside .
00,
Papilio thoas L. (1771).
Species No. 70.
/•
.'/•
Species No. 09.
Species No. 72.
Species No. 78.
Species No. 7.5.
Species No. 76.
Species No. 77.
Seba, Thrsaur. iv. p. 4G. t. .'JS. fig. G. 7. (1764).
Papilio Equcs Achiinis IhoaslAnnv, Mant. Pliint. p. ,'J36 (1771) {paiiim).
Papilii, thoas, Godart, Eiic. Mrth. ix. p. ^i. n. 103 (1819) (partini) ; Boisd., Spec. Gin. Lip. i. p. 355.
D. 197 (183(1) (piirtim) ; Kirby, Cat. Diuni. Lcp. p. 541. n. 155 (1871) (partiw) ; Haase,
Uiilemich. Mimicry p. 90 (1893).
HeracUdes thoas, Kirby, in AUeu's A'(((. Lihr.. Lep. Butt, ii, p. 'liB'i (1896).
36
( 556 )
The varioas subspecies vary so much inter se, and each is again individnally so
variable, that there is no single character in the pattern found in all forms by which
to distiugiiish the entire species J', tlioas from its near relative P. cresphontes. The
genitalia are more uniformly developed in all subspecies of P. thoas. The tenth
tergite of the 3 is always long, spatnlate ; the tenth sternite has on each side a
long, pointed, thornlike process, which bears prosimally at the base a ridge or
broad tooth ; the clasjier is sliglitly acuminate, being much less rounded than in
P. cresphontes and dorsally less emarginate ; the harpe, with the exception of the
Jamaica form, gradually narrows into a long point, being subtruncate and denticulate
in the Jamaican subspecies. The vaginal armature of the female is geogra{)hically
somewhat variable like the genitalia of the male ; the organs are similar to those
of P. cresp/io/ztes, but there is proximally of the vaginal orifice on each side a
broad and rather strongly chitinised depression, the edge of which projects as an
irregular ridge.
Early stages essentially as in P. cresphontes, the tubercles of the head and
thorax of the chrysalis shorter.
Ilab. Texas to Buenos Aires ; Cuba; Jamaica; not yet known from Haiti and
Porto llico, where the species may be expected to occur.
a. P. thoas melonius subsp. nov. (PI. VIII. fig. 59).
Papilio Eqiies Acliirns crexphonlfs Cramer, Pap. E,rnt. ii. p. lOG (1777) (partim ; Jamaica).
Papilio cresphniilen, Doubleday, Westw. & Hew., Gen. Dinni. L'p. i. p. 17. n. 16'J (1846) (purliin ;
Jamaica) ; Gray, Lkl Lep. Lis. Brit. Mus. i. Pap. p. 53. n. 2114 (1856) (partim) ; Felder, Verh.
Zonl. Dot. Ges. Wicn xiv. p. 310. n. 294 (1864) (partim ; Jamaica) ; Bull., Proc. Zool. Snc.
Lo7ul p. 481. n. 37 (1878) (Jamaica).
c? ? . A remarkably distinct form, usually of small size, resembling small
specimens of P. cresphontes. Yellow markings of iippersicle pale ; forewing :
discal patch R- — R^ as in P. cresphontes much longer than the following patch ;
patch M' — M- acnmiuate distally ; no spot in cell ; three snbmarginal spots,
occasionally preceded by one or two small dots ; i)atch SC — R' entire or nearly, but
sometimes deeply sinuate. Hindwiug : yellow marginal abdominal spot large,
continuous with the orange-red halfmoon, at the proximal side of which there is a
blue crescent; spot on tail small in male.
Underside, forewing : snbmarginal spots R^ — M- much larger than the others,
spot M^ — M^ being the largest, spot R- — R' about the same size as spot M*- — SM-
or smaller. Hindwing : three large orange patches R^ — M', sharply defined, the
third being the longest, being more than half the length of the pale yellow sub-
marginal patch R^ — M' ; a comj)lete series of j)ale blue sjiots, all the same pure
colour ; occasionally some orange scaling in apex of cell and behind SC- ; orange
anal halfring larger than in the other forms of P. thoas.
Genitalia : c?. Tenth tergite long, much slenderer than in the other subspecies,
strongly spatnlate, longitudinally grooved beneath, not carinate ; long pointed
process of sternite straight, the hairy tooth at base of this process small ; harpe
quite difl'ereut from that of all other subs])ecies, being short, subtruncate, with the
apex denticulate. ?. Anterior edge of vaginal orifice not tubercnliform, on each
side of the orifice a ridge, and behind the orifice two donble ridges.
Uah. Jamaica.
In the Tring Museum G <?<?, 3 ? ?.
Also in coll. Grose-Smith and coll. Adams.
( 557 )
b. P. thoas omedo Gundl. (1800).
Papilio thmis, Lucas, in Sagra, BiH. Cuba vii. p. 200 (1857) {parUm ?).
Pajnlio cresiihontes, Herrich-Suh., Corrrsp. Bl. Znol. Miii. Ver. Regensb. p. 173 sub n. 5 (ll^!>4)
{crespJioiites = oriedo Gundl, i. litt., false) ; Felder, Ver/i. Zoul. Bot. Ges. Wieii xiv. p. SIO-
n. 294 (1804) (pnrtim ; Cuba).
PapiUo oviedo Gundlach, in Poey, Rep. Fin. Nat. Cuba. i. p. 279. t. 5. fig. 1 (1860) ; id., Cnntr. Eitl. Cuba.
p. 133 (1881).
rapiliii thoas var. c. P. oviedo, Kirby, Cat Diurn. Lep. p. 541, subn. 155 (1871) (Cuba).
Papilio thoas var. oviedo Gundlach, Papilio i. p, 113 (1881),
Papilio erespluintes var, oriedo id,, Berl. Ent. Zeit. xxx. p. 132 (1881)).
PapiUo epitlwas Oberthiir, Bull. Soc. Ent. France p, 179, fig, 5 (18117) (" Me.xico?"j,
Papilio thoas var. P. epitlwas, Godman & Salv. , Biol. Centr. Amer., Lep. Rhop. ii. p, 729 (1901),
c? ? . Upperside : markings deeper yellow tlian in the preceding; discal band
broad, especially in male ; forewiug : patches 11- — M- of nearly equal length ; four
large snbmarginal spots preceded by one, two, or three smaller ones. Hindwing :
yellow anal spot large, continnons with red halfmoon, a blue spot at the proximal
side of the latter.
Underside much deeper yellow than in all other forms of P. thoas ; black
colour on forewing much reduced. Hindwing : two orange-red spots R- — M',
sharply defined, not so large as in the Jamaica form, but larger than ia continental
specimens of P. thoas ; a row of large pale blue discal spots, the middle ones partly
yellow, especially spots Ii- — M' ; anal crescent only slightly reddish, sometimes the
same colour as the other submarginal sjJots ; sjiot on tail large above and below.
Genitalia : S. Tenth tergite spatulate, being constricted before apex, shorter
than in continental specimens, carinate beneath, broader than in P. thoas melonius ;
acute process of sternite comparatively short, curved ; harpe long, gradually tapering
to a point.
Hab. Cuba.
The deep maize-yellow underside and the large blue patches of the hindwing
are the most distinctive external features of this insect, which cannot easily be
confounded with /-". cresphontes.
In the Tring Museum 2 cJc?, 2 ? ? , from : Cuba (Gundlach); Gibara (Tolliu).
c. P. thoas autocles subsp. nov.
Papilio thoas. Gray, List Lep. Ins. Brit. Mus. i. Pap. p. 54. n, 200 (1850) (partlm ; Mexico ; Yucatan;
Nicaragua) ; Reak,, Proc. Ent. Soc. Philad. ii, p, 138, n, G (1803) (Nicaragua ; syn. jiartini) ;
Weidem,, ibid, ii, p, 148 (1803) {partim); Felder, Verh. Zool. Bot. Ges. Wien xiv, p. 310, n, 295
(1804) (partini ; Mexico ; Yucatan : Nicaragua?) ; Oberth., Et. d'Etd. iv, p. 70. n. 210 (1880)
{parlini ; Mexico) ; Godm. & Salv., Biol. Ctntr. Amer., Lep. Rhvp. ii. p. 223. n. 53. t. 09. fig. 4.
genit. (1890) (purtim) ; Mayn., Man. N. Amer, Butt. p. 14. D. 21. fig. 10b (1891) (S. Arizona ;
Texas; "probably Florida"; partim'i) ; Holland, Butt. Book p. 311. n. 7 (189;i) (partim ■
Texas ;— t. 42. fig. 4. alia subsp.) ; Denton, .Voths Butt. U.S.A. ii. p, 345 (1898—1900) [partim ;
occasionally in Texas ;— fig. ad sequent, subsp. referendae) ; Godm. cS: Salv,, I.e. p. 729 (1901)
(Honduras).
Papilio cresphontes, Felder, Verh. Zool. Bot. Ges. Wien .xiv. p, 310, n, 294(1804) {partim ; Mexico ;
Honduras),
Papilio orni/thion, Staudinger (mm Boisd,, 1830, err. det.), E.rot. Tayf. p. 10 (1884) (Mexico ; " if my
specimen is true orui/thion, the latter is var. of thoas "}.
c? ?. Paler than /'. cresphontes, with which it occurs together, being also paler
than the other continental forms, of P. thoas. Upperside, forewing: no spot in
cell or only a vestigial one ; patch SO* — R' deeply excised, rarely without black
sinus or spot; spots before upper angle of cell small ; four submarginal spots,
( 558 )
rarely three, sometimes a complete series, the njiper ones being small ; last spot
nsnally distinctly smaller than the third (from behind).
On umlergide the last snlnuarginal spot of the forewing smaller than the fourth
sjiot (counted from behind), the third sjjot being the largest ; cell almost entirely
j)ale primrose-colour, the black streaks being either short or indistinct.
Genitalia: S. Tenth tergite broad, narrowing apicad, constricted before apex ;
jiointed process of sternite curved, the hairy ridge standing at its base shortened to
a tooth ; harpe long, gradually narrowed to a point.
Hab. Texas to Nicaragua; name-type from Guerrero.
In the Tring Museum 00 c?t?, 7 ? ?, from: Houston, Texas; Jalapa, July
1897 (W. Schaus) ; Cordoba, February 1896 (W. Schans) ; Songolica, July 1S9G
(W. Schans) ; Cuesta de Misantla, June 1896 (W. Schans) ; Guerrero (0. T. Baron) ;
Mexico City (ex coll. FeJder) ; Escuintla, W. Guatemala, llOD ft., September 1904
(A. Hall) ; San Pedro Sula, Honduras.
In coll. F. D. Godman 6 c? c?, 2 ? ? from Nicaragua, and a long series from
more northern localities.
d. P. thoas nealces subsp. nov.
Pajiiliu (Acliirtis) ajax, Miiller (nun Linn^, 1758, err. det.), Nalurx. v. p. 575. n. 32. t. 17. fig. :i
(1774).
Pupilio «Aoo.s, Kollar, Denkschi: K. Ak. Wis^. ]Vie)i, Math. Xnt. CI. i. p. 355. n. 11 (lfi50) (KioMeta,
smaller than Brazilian) ; Gray, Cut. Lep. Ins. Brit. Muk. i. Pup. p. 39. n. 1!M5 (1852) (/mrlini ;
Venezuela); Bull. & Druce, Proc. Zool. Soc. Lund. p. 3(;5. n. 377 (187-1) (Costa Rica); Oberth.,
Et. d'Ent. iv. p. 70. n. 210 (1880) iimrtim ; Muzo ; Carare ; Car.icas) ; Godm. & Salv., Tmnx.
Ent. Sue. Loud. p. Hi], n. 245 (1880) (Sta. Marta) ; Walk., Enf. .!/<.. .1%. xi.x. p. 26 (1882)
(Panama, common, a fast flyer) ; Godm. & Salv., Biol. Ciinti: Amer., Lep. Rhnp. ii. p. 223.
n. 53 (1890) (partini) ; Maas. * Weym., in SlUbel, Reisvi S. Amer., Lep. p. 11. n. 41 (1890) ;
iid.. I.e. p. 18. n. 31 (1890) ; iid.. I.e. p. 24. n. 100 (1890) (west side of Cordillera of Bogota) ;
iid., /.'■. p. 32. n. 135 (1890) (Colombia); iid., /.c. p. I! 6. n. 24 (1890) (Guayaquil); Hahnel,
Iris iii. p. 149 (1890) (San Este'ban) ; id., I.e. p. 201. 205 (1890) (Valera, Tenez.) ; Eimer,
Orthogen. Schmetl. p. 138. fig. 63 (1897) (Neu-Grauada) ; Dent., Moths Butt. U.S.A. ii. p. 345.
figs. (1898 — 1900) {/inyfim; — the figures apparently taken from Costa Rica specimens) ; Kaye,
Trans. Ent. Sne. Lmid. p. 207. n. 198 (1904) (Trinidad).
cJ ? . In colour intermediate between the preceding and the following form,
beiug deejier yellow than P. t. autocles and paler than P. f. t/ioa.s. — Uppin-.^tit/e,
forewing : cell-spot conspicuous in nearly all si)ecimens ; small spots in front of
apex of cell larger than in the preceding form, but smaller than in the following
one ; four submarginal spots, rarely three only, the row sometimes continued by
one, two, or three small spots, last spot about as large as the third (from behind),
or larger.
Underside : submarginal spots of forewing larger than in P. t. thoas, especially
the uj)per ones, last spot usually about the same size as the fourth (from behind),
often larger; cell with distinct black streaks.
Genitalia : S . Pointed process of tenth tergite long, the ridge at its base only a
little lower ju'oximally than distally, with feeble indication of being siunate ; harpe
short, pointed, usually denticulate.
Hab. Nicaragua to West Ecuador, eastwards to Trinidad and the Lower
Orinoco, Nicaragua specimens leading over to the jireceding form ; name-type
from Muzo, Colombia.
In the Tring Museum 210 c? cJ, 28 ? ? , from : Azaiiar de Carthago, Costa Rica,
February liS09 (J. Underwood); Carillo, Custa Pica, 300(i ft., October 1904
( 559 )
(A. Hall); Volran de Miravalles (Uuderwood) ; Limon, October 1004 (A. Hall);
Bogava, 800 ft, and Boqnete, 3500 ft, Chiri(|ni (Watson) ; Parida I., Sevilla I.,
Cebaco I., and Brava 1., January 1902 (Batty) ; K. Dagua, W. Colombia (Rosenberg);
Pereira, Cauoa; Muzo, Ducember 1896; Peperital to Buenavista, January 1897
(Dr. Burger); Villavicencio to R. Ocoor, January, and Villavicencio to Monte Redondo,
March 1897 (Dr. Biirgcr) ; Canancbe, Oundinamarca, July 1903 (Matban) ; Oiiaca,
S. Marta, 2000 ft. (Eiigelke) ; . Mocotone and Tacbira, Venezuela (Briceilo) ;
Cnmana, 1300 ft. (Andr^) ; Trinidad; Caparo valley (Dr. T. Rendall) ; La Vuelta
and Suapure, Caura R., Orinoco, February and May (S. M. Klages); Paramba,
N.W. Ecuador, February — June 1897 (Rosenberg); Cachabi, January 1897,
Cbimbo, August 1897 (Rosenberg) ; Rita and R. Cayapas, N.W. Ecuador (Flemming
and Miketta) ; Quevedo (v. Buchwald).
e. P. thoas tlioas L. (1771).
Seba, T/iesaiir. iv. p. 46. t. 38. fig. 6. 7. J (1764) ; Drury, Illustr. Exol. Lik. i. p. 44. t. 22. fig. 1. 2. ^J
(1770) (Surinam).
Piijillio Eqiies AchimK thnas Linn^, }tant. Plant, p. 5.3G (1771) (partim) ; Drury, l.r. Inrlex (1772) ;
Fabr., Syxl. Ent. p. 454. n. 48 (1775) (partim); Cramer, Pap. Exot. ii. p. 108. t. 167. fig. A. B.
(1777) (Surinam) ; Goeze, Ent. Beytr. iii. 1. p. 71. n. 4 (1779) (ijarthn) ; Fabr., Sprc. Jjis. ii.
p. 19. n. 70 (1781) {partim) ; id., Maiit. Ins. ii. p. 10. d. 87 (1787) {partim) ; Jabl. & Herbst,
Naturs. Schmett. iii. p. 127. n. 90. t. 40. fig. 3. 4 (1788) (Surinam) ; Gmelin, Stjst. Nat. i. 5.
p. 2240. n. 321 (1790) [partim) ; Jung, Aljihab. Verz. Srhm. p. 23G (1792) ; Fabr., Ent. Syat.
iii. 1. p. 32. D. 94 (1793) {partim) ; Esper, Ansl. Schmett. p. 198. u. 90. t. 49. fig. 1 (1797).
Papilio {thoas), Meerburgh, Afh. Zelik. Gew. t. 21 (1775).
Princeps heroicus thoaa, HUbner, Samml. E.tot. Schmett. i. t. 114. fig. 1. 2 (1806 — ?).
Iferaclkles thoas^ Hiibner, Ver^. bek. Schmett. p. 83. n. 852 (1818 ?) ; Kirby, in Hubner, Samml.. Exot.
Schmetl. ed. ii. p. 96. t. 114. fig. 1. 2 (190—?) {Viterat. partim).
Papilio thoas, Godart, Enc. MHh. ix. p. 02. n. 103 (1819) {partim) ; Lacord., Ann. Soc. Ent. Fr. ii.
p. .383 (1833) (Guyane) ; Lucas, in Guur., Diet. Pitt. Hist. Nat. vii. p. 50. (1838) {parlim) ;
Wallace, Tran.'<. Ent. Sue. Luiid. (2). ii. p. 255 (1854) (Amazons, gardens) ; Bates, ibid. (2). v.
p. 347 (1801) (Pari) ; id., Journ. Entom. I. p. 228. n. 27 (1862) ; id.. Natural. Riv. Amaz. p. 52
(1864) (Paril.in street) ; Fel.ler, Verh. Zool. Bot. Ges. Wien xiv. p. 310. n. 290 (1864) (Surin,-im ;
Paiit); MoscLl., ibid, xxvi, p. 296 (1876) (Suiinam) ; Butler, Trans. Ent. Soe. Loud. p. Ull.
n. 231 (1877) (Serpa, April) ; Sh.arpe, Proe. Zool. Soc. Lond. p. 555. n. 4 (1890) (E. Araguaya);
Eimer, Orthogrn. Schmett. p. 138. fig. 62 (1897) (Surinam).
Papilio thoas var. thoas, Obertbur. Et. d'Ent. iv. p. 70. sub n. 210 (1880) (Guyane).
J ?. As yellow in tint as tbe large subspecies P. thoas cint/ra.'i. Uppcrside,
forewing: ajiical spot usually small, sometimes a mere dot; spots before upper
angle of cell larger than in tbe other forms, e.x'cept cinijras, there being often a
small additional spot within the subcostal fork; discal patch R' — If' smaller than
in the other forms ; patch R" — R' as a rule the same in size as patch R-^ — M', rarely
a little longer, the veins R- and backwards more narrowly black between the patches
than in P. thoas nealces and P. thoas thoantiades ; j)atch SC'' — ^R^ usually without
black spot or sinus ; four submarginal spots, sometimes three, small ; cell-patch
rarely absent.
Underside : submarginal spots of forewing corajjaratively small, especially the
upper ones; black discal area of hindwing usually broad and the bine halfmoons
\mva in colour as a rule, black marginal line wider and submarginal patch S(J- — R'
on the whole less projecting liasad than in /'. thoas nealces.
Ilab. The Gniauas ; Lower Amazons.
In the Tring Museum 12 J'c?, 0 ? ?, from: Demerara, Esscquibo R., and
Berbice R., British Gniana; Surinam.
( 560 )
/. P. thoas cinyras M6adtr. (1857).
Papil/ii himpednnGny, List Lip. Ins. Brit. Miis. i. Pd/j. p. 54. n. '206(1850) (Villa Nova; tiom.mtdum !).
Papilid cinynis MenetriiSs, Eniim. Corp. Anim. .!/««. Pi-trop., Lep. i. Suji/il. p. G8. n. 1124. t. 7. fig. 3.
(1857) (" Bahia" error loci); id., I.e., Descr. p. 111. n. 1124 (1863); Gerst., Stflt. Ent. Zi'it. six. p. 302
(1858) ("is aberrat. of /'. tlwax") ; Felder, Wieii. Ent. .Uun. iii. p. .393. note (1859) (" distinct
from thoas ") ; Bates, Trims. Ent. Sm: Lond. (2). v. p. 347 (18(51) (from Villa Nova upwards) ;
id., Journ. E/ttom. i. p. 228. n. 29 (I8C2) (Upper Amazons ; ■' interior of province of Bahia "
error of local, or of identif.) ; Felder, I'erh. Zuul. Rot. Gex. Wien -xiv. p. 310. n. 29U (18G4)
{pnrlim ; Ega ; Villa Nova) ; Butl., Ann. Mug. N. II. (4). xx. p. 127. n. fi2 (1877) (R. Mairo,
Peru) ; Hopff., .S7e». Ent. ZHt. xl. p. U% n. 19 (1879) (Peru) : Oberth,, /■;/. d'Ent. iv. p. 7(i.
n. 209 (1880) (Teffe, Obydos) ; Mil., Nat. Sci. v. p. 243 (188(5) (Monaco !) ; Hahnel, Iris iii.
p. 240 (1890) (Villabella, Amaz.) ; id., I.e. p. 283 (1890) (Pebas).
rapilio llioas L. var. eimjras, Staudinger, Exol. Tagf. p. 16. t. 11. ^J (1884) ; Michael, Iris vii. p. 213
(1894) (Sao Paulo de Oliven^a).
Piipilio thoas, Maassen & Weym., in Stiibel, Reisen 8. Amer., Lep. p. 82. n. 50 (1890) (Upper Amazons);
Dognin, Lep. Loja p. 37 (1891); Holland, Bntt. Book t. 42. fig. 4 (1899); Weeks, lUiislr. Dium.
Lep. p. 20 (1905) (Chulumani).
Papilio eyniras (!), Eimer, Orlhogen. Sehmett. p. 137 (1897).
In the south iatergradiug with the next subspecies.
(??. Vjtperside, forewiug : cell-spot pre.sent, sometimes small ; spots in front
of apex of cell usnallj' large, sometimes minute, patch SO' — R^ entire, sometimes
bearing a black spot, seldom deeply sinuate ; submargiual spots absent or small,
usually present in Bolivian specimens. Hindwing: yellow band broad, upper
bubmarginal spots usually much smaller ;than the others ; red anal crescent absent
or (in southern specimens) small, blue halfmoon distinct.
Hah. Eastern Ecuador, Amazons (except delta district), southward to Bolivia.
In the Tring Museum 79 c? c?, 9 ? ? , from : Obidos ; R. Uaupes, Upper R. Negro ;
Iquitos (Stuart) ; R. Cachyaco, affl. of R. Huallaga (Stuart); R. Napo, E. Ecuador
(R. Haensch) ; Zamora (0. T. Baron) ; R. Chuchnras, affl. of R. Palcazu, 320 m.
(W. Hoffmanns) ; Chanchamayo (Schnuke) ; Palcazu (Sedlmayr) ; Per(jn6 R„
March 1900 (Simons) ; R. Mixiollo, Loreto (Baer) ; R. Toro, Ija Merced, August —
September 1901 (Simons); Pozuzo, Huauuco (W. Hoftmanns) ; La Union, R.
Huacamayo, Carabaya, 2000 ft, November and December 1904 (G. Ockenden) ;
Chirimayo, Carabaya, July 1901, 1000 ft. (Ockenden) ; Oroya, R. Inambari, 3500 ft.,
November 1901 (Ockenden); Caradoc, Marcapata, 4000 ft., February 1901
(Ockenden); R. Slucuri, S.E. Peru, 25(Mi ft., June 190l (Ockenden); Salampioni,
Bolivia, September 1900 (Simons) ; Charuidaya, 13iX) m., June 1901 (Simons) ;
R. Songo (Garlepp); Sorata (Guenther) ; Mapiri ; Salinas, R. Beni, July 1895
(Stuart) ; Prov. Sara, S. Cruz de la Sierra (J. Steiubach).
g. P. thoas bra.siliensis subsp. nov.
Papilio thoas, Mt'ndtriKS, Mini. Sor. Imp. Moseou vii. p. 188. n. 3 (1829) (Brazil, larva) ; Doubl.,
Westw. & Hew., Gen. Dinni. Lep. i. p. 17. n. 170 (184(5) (Brazil) ; Gray, Cat. Lep. Ins. Brit.
Mux. i. Pap. p. ,39. n. 196 (1852) {partiin ; Brazil) ; id., List Lep. Ins. Brit. Mas. i. Pap. p. 54.
n. 2(M'i (185(5) (jiarliin ; Brazil) ; Me'nutr., Ennm. Corp. Anim. .Uns. Petrop., Lep. i. p. 4. n. 63
(lS.^i7) (Brazil); Felder, Verh. Zool. Hot. Ges. Wieti xiv. p. 310. n. 295 (1864) (partim; Brazil);
Piillw., Stetl. Ent. Zeit. xxvi. p. 129 (18G5) (Corcovado); Butler, Cat. Dium. Lep. deser. Fabric.
p. 24C.. n. 44 (1869) (Brazil) ; Capronn., Ann. Sor. Ent. Behj. xvii. p. 9. n. 12 (1874) (partim ;
Botafogo, August; Rio, Entre Rios ; very common); Burm., Deser. Rep. Argent. V. Lfp.,
Atlas p. 3. t. 2. fig. 1. 2. larva (1879) (Bio do Jan.); Oberth., Et. d'Ent. iv. p. 70. n. 210 (1880)
(partim; Brazil); Jones, Proe. Lit. Phil. Sue. Lirerp. p. 41 (1883) (metamorph.) ; Seitz,
Stett. Ent. Zeit. li. p. 98 (1890) (Corcovado) ; Weym., ihid. Iv. p. 315. n. 14 (189.'.) ; Mabilde,
Guia Prart. Borbol. liio Grande do Sid p. 49 (1896) ; Peters, Illnstr. Zeitsehr. Ent. ii. p. 51
(1897) (Nova Friburgo ; larva).
Papilio eresphontes, Felder, Verh. Zool. But. Ges. Wien xiv. p. 310. n. 294 (1864) (partim ; Brazil).
( 561 )
(? ¥ . A large form, nearl}' as bright yellow as P. thoas cimjras ; the femule
very little paler than tlie male. Upperside : fore wing somewhat falcate ; cell-spot
absent or small ; snbapical spot large in nearly all specimens, usually produced into
a point ; patch ^C' — R' di^cply sinuate, sometimes completely divided ; submarginal
spots always present, usually four. Hindwing : red anal spot present ; distal
edge of yellow band crossing cell at base of M- or proximally of it ; upper sub-
marginal spots usually more or less rotnndate.
Underside : snbapical spot of forewing large, the third and fourth snbmarginal
spots somewhat transverse, often also the second ; black marginal band of both
wings broad in the majority of specimens.
Genitalia essentially as in P. thoas mitodes ; S, lateral ridge of tenth tergite
not toothlike, not subsinuate ; harpe somewhat curved, ventrally denticulate before
apex. ¥ , edge of vaginal orifice proximally raised into a tubercle or an obtuse
process.
Hab. Bahia southwards, extending westwards to South-East Bolivia, inter-
grading completely with the next form, and in S.B. Bolivia with the preceding
subs])ecies, there being neither sharply defined morphological nor geographical
limits. Type of name from Petropolis.
It is probable that P. thoas is in these districts a wanderer like the Nearctic
/'. cresphontes, which would explain the absence of a strict line of separation
between the Brazilian and Argentinian forms.
The large Brazilian specimens are usually regarded as being the same as the
form from Surinam, which is the nomenclatorially typical subspecies of P. thoas.
In the Tring Museum 50(Jcf, 26 ?¥ and a larva from: Minas Geraes,
February 1897 and 1901 (A. Kennedy) ; Petropolis, December and January
(J. Foetterle) ; S. Paulo; Castro, Parana (E. D. Jones) ; Yhu, Paraguay, September
— December 1890 (Andeer) ; Sapucay, Paraguay (W. Foster); Patino Cud, Paraguay,
February (Montforts), Tucuman (J. Steiubach); Salta (J. Steinbach); S. Jose' de
Chiqnitos, E. Bolivia, July 22. 1904 (J. Steiubach).
h. P. thoas thoantiades Burm. (1878).
Piipilio thoantiaik's Burmeister, Dcm:): Rej>. Argent, v. Leji. p. 59. n. 6 (1878) (var. of Uinas) ; id., l.r.
Atlnx t. 2. fig. 3. larva, 3a. pupa, t. 4. fig. 9. ? (1879) ; Oberth., Et. d'Ent. W. p. 70. n. 210
(1880) {partim ; Bueno.s Aires) ; Gosse, Entum. xiii. p. 194 (1880) (Corrientes).
Pajiilio thoaiitUks (!), Staudinger, Exot. Tag/, p. IC (1884) (Argentina).
c? ? . Smaller than the preceding; discal band of fore- and hindwing very
variiible in width, usually paler than in J\ thoas brasiliensis ; di.scal and marginal
black bands on underside of hindwing broad.
Genitalia : t?, harpe more curved than in brasiliensis.
Hab. Province of Buenos Aires and northward.
In the Tring Museum: 30 S6, 20 ? ? and some larvae and pupae from:
Buenos Aires, January, February and March (Ruscheweyh) ; Cordoba, Paysaudu,
and Rosario, March (Ruscheweyh) ; La Soledad, Entre Rios, February 1899
(Chas. Britton).
67. Papilio homothoas s])cc. nuv. (PI. V., fig. 13).
Ptipilln crrsphonles, Felder (non Cramer, 1777, err. det.), \'erh. Ziiol. But. Ges. Wien xiv. p. 310.
n. 294 (18G4) {parti m ; Bogota).
Resembling P. thoas ; forewing shorter ; bases of distal segments of antenna
of male broadly yellow beneath ; tail shorter and more strongly spatulate, being
( 562 )
narrower proximally ; no spot in cell of forewing on upperside ; patch SC— R' of
forewing entire or with a small siuns only, patch R- — R^ projecting beyond the
next as in P. crespkontes, submargiual spot R' — M' being more proximal than in
P. tkoas ; genitalia quite iliiferent.
S. "Wings, upprrgidc : markings chrome-yellow, deeper in tint than in
P. thoas. Forewing : no spot in cell ; spots before npper angle of cell as
large as in P. thoas thoas ; a small additional spot at base of subcostal fork ;
patch SC* — R' entire or with black spot or sinus, 2)atches R=— SM- contiguous,
patch R- — R^ truncate distally, projecting beyond the next one, its outer edge being
in a line with that of the preceding patch ; three submarginal spots of almost
equal size standing in an oblique row. Hindwing rather shorter posteriorly than
in P. thoas ; tail more evenly s]iatnlate, broader at apex and narrower proximally
than in P. thoas.
Underside deeper yellow than in South American P. thoas, but not so deep as
in P. thoas oeiedo from Cuba. Forewing : cell striped with black ; submarginal
spot R' — M' much larger than the others and extending much more proximad.
Hindwing : cell yellow, except a narrow ajjical crescent ; black discal band narrow,
each patch bearing a blue or a yellowish spot; two orange-red si)0ts R- — M' at cell;
orange-red anal spot connected with the marginal spot.
?. Somewhat paler than male, markings of upperside smaller, patch St'-^ — R'
of forewing with small black sinus.
Genitalia: S . Tenth tergite bifurcate, the lobes projecting laterad, being narrow
and pointed ; tenth steruite short, on each side with an apically sinuate lobe, the
lower one of which is acute and somewhat produced. Harpe a broad concave plate,
obtuse at ajiex, which is a little curved inward; ventral edge parallel to edge of
clasper, distally denticulate. ? . Edge of vaginal orifice slightly elevate all round ;
from this low ring extends on each side backwards a ridge which borders a large
postvaginal depression, the ridge being raised close to the vaginal orifice into a
pointed, triangular process ; no strongly chitinised ridge or dej)ression proximally
of the orifice.
Hal). (Jiudad Bolivar, Lower Orinoco, June 1801, 1 S, type; Maripa,
('aura R., October 1903 (S. M. Klages), 1 c? ; Lower Orinoco, November 1897
(Cherrie), 1 ? ; Colombia, 3 c?c?.
In coll. Godman from Colombia ; in coll. Adams from Marguerita L ; in the
British Museum from Venezuela and Bogota.
08. Papilio cresphontes Cram. (1777).
Aubent., Planch. Eiilum. t. i;;t. $ (1765) ("Guadeloupe" false).
Papilh Eques Arhnux tlwas Linnc', Maid. Plant, p. 536 (1771) (partim) ; Fabr., Syst. Ent. p. 454.
D. 48 (1775) {jmitim) ; id., Spec. Im. ii. p. 19, n. 76 (1781) (partim) ; id., .]f(wl. Ins. ii. p. 10.
n. 87 (1787) {parlhn) ; id., Ent. Si/st. iii. 1. p. 32. n. 94 (1793) (pni-lhn).
Pupiliu Ecpifs .Iclnnix cresphontes Cramer, Pap. Exot. ii. p. 106. 107. t. 16,5. fig. A. ? , t. 166. fig. B.
cJ (1777) (N. York ; Carolina— "Jamaica" alia subsp.) ; Goeze, Ent. Peyti: iii. 1. p. 86. n. 64
(1779) ; Jabl. c*i Herbst., Natnrs. Sehmett. iii. p. 121. n. 89. t. 39. fig. 1. 3. ? , 2. (J (1788).
Papilio E(jues .Ichlriis tlwas fi) Papilio cresphontes, Gms:\m,S:ist. Nat. \. 5. p. 2240. sub n. 321 (1790).
Papilio ci-esphontfs, Esper, .'l«.s7. Schmelt. p. 199. subn. 90(1797) ; Doubl.,Westw. & Hew., <,'en. Dinrn.
Lep. i. p. 17. n. 169 (1846) (partim ; United Sts., Mexico) ; Kirtl., Proc. Ent. Sue. Loml. (2).
i. p. 101) (1851) (south shore of L. Erie ; recent arrival ; = thoas, Boisd. & Lee.) ; Gray, Cat.
Lep. Jns. Brit.Mus. i. Pap. p. 39. n. 194 (1852) (jmrlim) ; id., List Lep. Itts. Brit. Mns. i. Pap.
p. 53. n. 204 (1856) (partim) ■ MeniHr., Enum. Corp. Anim. J/«,s. Petrop., Lip. i. p. 68. n. 1123
(1857) (Mexico) ; id.,/.f., Lip. Descr. p. lll.n. 1123(1863) (.litliuct from thx,as) : Reak., Proc.
( 563 )
Ent Snc. PliHatl. ii. p. 137. n. 5 (18G3) (Honduras ; syn. excl.) ; Kirkp., ilitl. iii. p. 320 (1864)
(Cleveland, Ohio, rarely); Felder, Verh. Zi'ol. But. Gcx. Wkn xiv. p. 310. n. 294 (18(14)
Iparlim) ; Herr.-Sch., Conesp. Bl. Zonl. Mm. Ver. Brgensh. p. 172. n. .5 (18fi4) (Cuba) ; Edw.,
Trans. Amer. Ent. Sue. vi. p. 11. n. 21 (1877) (S. & W. States j occas. in Wisconsin, Michigan,
and Ontario) ; Colem., Amer. Natural, p. C88 (1877) (Berlin, Conn., Sept. G) ; Gerh., Macro-
Lcp. N. Am. p. 25. n. 438 (1878) ; Strecker, Biilt. Mulhs N. Am. p. G8. n. 7 (1878) ; Burm.,
Descr. Rfp. Ardent, v. Lep. p. 58. n. 1 (1878) (var. of thons); Bean, Otimil. Ent. x. p. 35 (1878)
(correct, of note in Amer. Nat. 1877'. p. 688) ; Saund., ihhl p. 48 (1878) (life hist.) ; Peck,
ibid. p. 60 (1878) (Fairfield Co., Conn.) ; Bo\],ihl(l. p. 1.54 (1878) (Dallas, Texa.s, larva and
pupa, on Zan till IX 1)1 urn raroliniuimm) ; French, /6/V/. p. 204 (1878) (position of pupa) ; Saund.,
ibid. p. 223 (1878) ; French, Trans. Drpt. Agric. lUiii. xv. p. 139 (1878) ; Saund., I.e. xi. p. 203
(1879) (Ontario) ; Moflfat, ibid. p. 240 (1879) (Hamilton, Out.) ; Murray, \hid. p. 240 (1879)
(Hamilton) ; Saund., T(e2>t. Ent. Soc. Ontario p. 60. fig. 38 (1879) (distrib. ; food-plants) ; id..
I.e. p. 41. fig. 19 (1880) ; French, Bept. S. J II. Norm. Uuir. vi. p. 43 (1880) ; Murray, Ca.tal.
Ent. xii. p. 120 (1880) (Hamilton) ; Saund., ibid. p. 120 (1880) (larva June, imago July, 1 irva
autumn, pupa hybernating) ; Oberth., 77(. </'&;/. iv. p. 70. n. 210 (1880) {parlim; Texas);
Comst., Rept. Dep. Ar/rie. p. 246 (1880) (metam.) ; Moffat, RejU. Ent. Sue. Ontario p. 10 (1881)
(Long Point and Ridgeway) ; Saund., ibid. p. 41. fig. 19 (1881) ; Moffat, Cunad. Ent. xiii.
p. 115 (1881) (Hamilton); French, ihid. p. 177 (1881) (life history); Gundl., Coidr. Ent.
Cuba. p. 131 (1881) ;iGoodall, Pupdio ii. p. 188 (1882) (Amherst, Mass., Sept. 5) ; Bruce, ibid.
p. 188 (1882) (Monroe Co., N.Y., larva on Ruta i;raeeolrns) ; Moffat, Rept. Ent. Soc. Ontario
p. 30 (1882) (larva in October; Hamilton, Ont.) ; Reed, Canad. Ent. xiv. p. 181 (1882)
(London) ; Jack, ibid. p. 219 (1882) (15 miles south of Montreal, August) ; Saund., ibid. xv.
p. 204 (1883 (larva on Zanthoxylnm amerieannm & Ptelea trifoVnita) ; id., I.e. p. 234 (1883) (egg,
JounglaTYa, on Zanthnxyhim fra.rineum) ; id., Ins. Inj. /''/■«;/ p. 377. fig. .389-91 (1883); Edw.,
Papilio iii. p. 26 (1883) (Wise, Northern N.Y., Me., Mass., Conn.) ; Colem., ibid. p. 43 (1883)
Conn.) ; Dimmock, Psyche iv. p. 99 (1883) (Cambridge, Mass., August) ; Saund., Canad. Ent.
xvi. p. 50 (1884) (Lake Erie) ; Edw., iliid. p. 109 (1884) (egg) ; Liutn., Pajiilio iv. p. 136. d. 3
(1884) (Rio Grande) ; Saund., Rept. Ent. Sue. Ontario p. 16 (1884) (larva on Pldea and Zanth-
oiyhini) ; Jack, ihid. p. 37 (1884) (Prov. Quebec, end of August) ; Ne.al, null. Dept. Agric, Ent.
iv. p. 87 (1884) (larva destr. by ants and .}fi,tilla !) ; Saund., Rept. Ent. Soc. Ontario xv. p. 29. 30
(1885) (Point Pelee, L. Erie) ; Hubb., r?is. Affiel. Orange p. 137. fig. 66. t. 10. 11 (1885) ; Bates,
Canad. Ent. xviii. p. 80 (1886) (Massachusetts) ; French, Butt. East. U.S.A. p. 103. fig. 20—22
(1886) ; Mayn., Butt. N. Engl. p. 50. n. 69. t. 5. fig. 69. 69.V (1886) ; Shann., Canad. Ent. xix.
p. 180 (1887) (Indiana ; larva on Pnpulus dilatata) ; Geddes, Rept. Ent. Sue. Ontario xviii. p. 23.
fig. 5 (1888) (Oxford, Ont.) ; Skinn., Canad. Ent. xxi. p. 127 (1889) (Philadelphia, occasionally);
Riley, 7)!,s-ec( Life ii. p. 2 (1889) (larva on orange); Edw., Bull. tf.S. Nat. i[us. xxxv. p. 12
(1889) (Hter. on metam.) ; Riley, Insect Life iii. p. 32 (1890) ; Dyar, Psyche v. p. 421 (1890)
(number of moults) ; Pack., Fifth Rept. U.S. Ent. Cunun. p. 472, 661 (1890) (egg, various larval
stages, pupa ; food-plants); Mayn., Man. N. Amer. Butt. p. 14. n. 20. fig. 10c (1891) ; Edw.
& Ell., Bull. Amer. Mus. N. H. iv. p. 75 (1892) (egg and first three stagts of larva);
Staley, Canad. Ent. xxiv. p. 204 (1892) (Marshall, Missouri, iv. — x., common) ; Davis, Journ.
N. York Ent. Soc. i. p. 47 (1893) (Staten L, N.Y., Aug., Sept. 1882) ; Beutenm., Bull. Amer.
Mus. N. H. v. p. 245 (1893) (N. York ; descr. of 1., p., i. ; two broods, June and Aug.) ;
Beth., CanwI. Ent. xxv. p. 260 (1893) (Lake Sincoe, Aug. 28) ; Holl., ibid. p. 311 (1893)
(Pittsburgh, larva on Zanthoxylnm and Ptelea, in Florida on orange and lemon) ; Soule,
Psyche vi. p. 530 (1893) (Brookline, Mass.) ; Haase, Untersuch. Mimicry i. p. 96 (1893) ;
Moffat. Canad. Ent. xxvi. p. 54 (1894) (London, larva in Oct. from being nearly full ■
fed to little ones) ; id., I.e. p. 123 (1894) (London, July and Aug.) ; EUi?, ibid. p. 176 (1894)
(Sparrow Lake, 110 miles north of L. Ontario); Davis, Ent. News v. p. 109 (1894) (Little
Rock, Ark.); White, ibid. p. 175 (1894) (Brooklyn); Ehrm., ibid. v. p. 212 (1894) (Pitts-
burgh, Pa.) ; Weed, Psyche vii. p. 130. n. 37 (1894) (N.E. Miss.) ; Moffat & Saund., Rept.
Ent. Soc. Ontario xxiv. p. 6 (1894) (London) ; Beth., ihid. (1894) (Lake Simcoe) ; Holl.,
ibid. p. 53. fig. 26 (1894) (Pittsburgh ; larva on Zanlhoxyluni and Ptelea) ; Moffat, Canad. Ent.
xxvii. p. 147 (1895) (London, June, full-fed larva in July) ; Denton, Psyche vii. p. 263 (1895)
(Wellesley, Mass., June 2.) ; Osburn, Ent. Neics vi. p. 282. n. 45 (1895) (Tennessee, rare, vi. to
ix., two broods) ; Grant, Canad. Ent. xxviii. p. 273 (1896) (Orillia, Ont.) ; Truman, EiU. Nem
viii. p. 29 (1897) (Volga, S. Dakota, travel-worn) ; Beutenm., Journ. N. York Ent. Soc. v.
p. 101 (1897) (distinct from thoa.i) ; Duzee, Bull. Buffalo Soc. N. Sc. v. p. 107. n. 6 (1897)
(Buffalo, occasional) ; Thorns., Canad. Ent. xxix. p. 263(1897) (larva on Rutaceae, Zanlhoxyluni
and Ptelea) ; Rowley, Ent. Ncn-s ix. p. 37 (1898) (Louisiana, Mo., larva on hop tree and prickly
( 564)
asb) ; Britton, ihitl. p. 173 (1898) (Newhaven, Conn., June ly) ; Ashm. & Schwarz, Proc. Eiit.
Soc. U'ashiiKjI. iv. p. 50 (18;W) (change of food) ; Beutenm., Bull. Amer. Mux. N. If. s. p. 310
(1898) (Highland Falls, X.Y.) ; HoUand, llntl. Boo/.- p. 311. n. 8. t. 2. fig. Kl, t. 4. Eg. 8—10,
t. 42. fig. 3 (IS'.l'.i) ; Beutenm., Butt. X. Ynrk City p. (1. n. 4. fig. J (1902) ; Moffat, Reid. Knt.
Sor. Oiitari., xx.\iii. p. 51 (1902) (Trenton, Aug. ; London, Ont.) ; Evans, Ibid. p. 82 (1902)
(Trenton); Walk., ibid. p. 85 (1902) (Point Pelee, Leamington; Walpole I.); Clark, Enl.
Xeu-s xiii. p. 27 (1902) (Xewtonville, Mass., Sept. 13) ; Field, ibid. xiii. p. 331 (1902) (East
Alstead, N.H., June 12); Hoag, ibid. .\iv. p. .320, 321 (1903) (Altamira & S.Louis Potosi,
Mex.) ; Mofltat, Rej}t. Eiit. Soc. Ontario xxxiii. p. 58. fig. BH (1903) (London, Ont.).
Ilcradides nxihis Hiibner, Verz. hek. Schmett. p. 83. n. 850 (1818 ?) (nom. nov. loco cresplionte-i).
rupilin tlioas var., Godart, &ic. .Vetli. ix. p. G2. n. 103 (1819) (2iariim) ; Boisd., Spec. Oeii. Lip. i.
p. 355. n. 197 (183C) {jMrlim) ; Doubl., List Lep. Im. Brit. ilus. i. p. 17 (1845) (jmrtim).
Papilio tlioo.t, Boisduval & Leconte, /7/.s(. Gcii. Lip. Amn: Sijit. p. 32. t. 12. fig. 1. ? , t. 13. fig. 1. (J,
2. larva, 3. pupa (1833) (Georgia ; Florida) ; Lucas, in Gui'r., Diet. Pitt. Hist. Nat. vii. p. 50
(1838) (partim) ; Doubl., in Westw., Arcana Eiit. i. p. 144 (1845) (habits) ; Poey, Mem. R.
Soc. Econ. Uabana p. 234 (1846) ; Lucas, in Sagra, llixt. Cuba vii. p. 206 (1857) (partim) ;
Gosse, Letters from Alabama p. 170 (1859) ; Morris, Sijn. Lep. N. Am. p. 7. n. 10 (1802) ;
Weidem., Prrjc. Ent. Soc. Philad. ii. p. 148 (1863) (partim) ; Reed, Canml. Ent. i. p. 19 (1868)
(London, Ont.) ; Butler, Cat. Diuru. Lep. descr. Fabric, p. 24G. n 44 (1869) (partim) ; Parker,
Amer. Entom. ii. p. 175 (1870) (Iowa) ; Bean, Ent. Mo. Mag. x. p. 248 (1874) (Galena, 111. ;
quite uncommon, Aug. Sept.) ; Carey, Rept. Ent. Sor. Ontario p. 5 (1875) (Amherstburg) ;
Saund., ihid. p. 15 (1876) (Northbridge, Out.); Cook, iliid. (1876) (Lansing); Dent., Caiiad. Ent.
ix. p. 160 (1877) (Ontario) ; Murray, ibid. x. p. 120 (1878) (Hamilton, Ont.) ; Godm. & Salv.,
Biol. Ceiitr. Amer., Lep. Rliop. ii. p. 223. D. 53 (1890) (partim) ; Christ, Mill. Scbwei:. Eiit. Ges.
ix. p. 273 (1897) (= cresplumtex) ; Dyar, Bull. U.S. Nat. Mux. lii. p. 3. n. 14 (1902) (Atl.
States; Arizona; Mexico; partim 'i); Laur., Ent. Neici xiv. p. 296 (1903) (Miami, Fla.,
common) ; Sherm., Rept. Ent. Soc. Ontario xxxvi. p. 26. fig. 11 (1905) (N. Carolina).
Papilio Ihoas var. b. P. crcsphontes, Kirby, Cat. Diurn. Lep. p. 541. sub n. 155 (1871) (Univ. Amer.).
Papilio Ihoas var. cre.y.Iiontes, Gundlach, Papilio i. p. 113 (1881) (Cuba) ; Aaron, Papilio iv. p. 172
(1884) (S. Texas, common).
Papiliii thoas (crexphontes), Dwight, Psyche, iii. p. 327 (1882) (Dutchess Co., N.Y.) ; Bunker, Canad.
Ent. XV. p. 100 (1883) (N. York, two broods) ; Perrin, Rejit. Enl. Soc. Ontario xxxiv. p. 90
(1904) (15 miles from Halifax, Aug. 1901).
neraclichs crexphnntes, Scudder, Belt. East. U.S.A. ii. p. 1.334. t. 14. fig. 12, t. 27. fig. 2, t. .35. fig.
21-3, t. 41, fig. 4, t. 57. fig. 3, t. 66. fig. 3, t. G8. fig. 19, t. 73. fig. 1, t. 76. fig. 16, t. 79. fig. 02-6,
t. 85. fig. 8, 9, 10 (1889) (morphol., metara., etc. \—\\\.eT. partim ad aliam spec, ref.) ; Kirby, in
Allen's iYo<. Libr., Lep. Bnlt. ii. p. 282 (1896) ; Scudder, Psyche viii. p. 210. t, 5. f. 5, I. juv.
(1898); Mayer, ;6»/. p. 299 (1898) (Dry Tortuga) ; Kirby, in Buhn., Samml. E.eol. Schmett.
ed. ii. p. 96. t. 314. fig. 3, 4 (190—?).
Papilio chresphontes ('.), Dury, Cincinnati Soc. Nat. Hi.it. i. p. 12 (1878) (Cine, not common) ;
Bubua, Enl. News viii. p. 98 (1897) (Cleveland, Ohio ; rare).
Specifically distinct from P. tlioas, with which it occnrs together in Central
America, Texas, and Cuba. Dr. Holland separated it correctly from P. thoas, but
unfortunately figured as thoas not the nortberu form of that species, but a South
American specimen (doubtless from the Amazons). The difference in pattern
between P. thoas and cresphontes being but slight, many authors have considered
the two insects to be one species. However, that opinion is erroneous. The true
distinction between the two insects is not on the surface. The sexual organs, as
sjiecified below, exhibit a divergency which is quite astonishing, and it is no less
surprising that this difference has never been noticed, although one of the most
peculiar features of the organs is visible without dissection.
The only constant difference in ])attern between P. cre.iphontes and the northern
forms of P. thoas is the larger size of the spot R-— R' (the fifth) of the discal
row of the forewing. This spot is larger in crcsphoiit.es than the nest sj)ot, the
snbmarginal spot W — M' being consequently more proximal above and below than
the snbmarginal spot R- — R'.
( 565 )
Individually variable. The jiatch SC' — R' on tlie npperside of the forewing
is nsnally excised, seldom completely divided, the black siuns being often rednced
to a spot, which is sometimes altogether absent, as is always the case on the
underside. Some specimens have a complete row of submargiual spots. One of
the most interesting variations is represented by specimens in which there is a
second yellow spot before the subcostal fork standing at the proximal side of
the subapical spot, this additional spot proving the suba])ical spot to belong to the
submarginal series, not to the discal band. The size of the discal patches is very
variable, spots R' — M" being more or less pointed distally. There is apparently
never a yellow spot in the cell, as is so often the case in P. thoas. The width of
the yellow antemedian band of the hindwiug is variable.
On the underside considerable variability obtains in the amount of black on
both wings and in the size and number of the orange spots on the hindwing ; these
spots are occasionally of a rufous tawny tint.
There is apparently no marked variation according to season and locality. The
few Cuban males which we have examined show, however, a more or less distinct
orange-red spot in the cell of the hindwing below, which spot is hardly ever
indicated in continental specimens ; our Cuban females do not possess the spot.
The specimens from the southern Atlantic States are on the whole the largest,
the Central American ones being, on the contrary, of inferior size.
The distribution is somewhat fluctuating, the occurrence in the northern
districts of the range not being regular. We suspect the species to be a wanderer
also in the southern districts.
P. cresphontes was originally doubtless a northern form of P. thoas. The
ranges now overlap to a considerable extent, the insects having become so difi'erent
that they are independent of one another and can exist in the same locality without
fusion (compare definition of " species " in Nov. Zool. 1896, p. 438, and 1903
Suppl. p. xli ; also Poulton, Pres. Address, Etit. Soc. Lond. 1903).
Genitalia : S. Tenth tergite with a very short process instead of the long
spatulate process of P. thoas, there being in consequence an open space dorsally
between the claspers ; the process of cresphontes can be felt by gently moving
the finger over the upper edge of the claspers from behind ; the tenth sternite,
which in thoas bears on each side a long curved pointed process which is
proximally dilated into a hairy ridge, has in cresphontes a long proximal
process which is somewhat irregular at the edges, being narrowed to a point ;
from beneath this process, i.e. standing distally of it, projects an acute and
somewhat curved thornlike process. Clasper shorter and much more rounded than
in P. thoas ; harpe broad, hollowed out, narrowest at apex, which is rounded,
the apical and ventral edges being minutely denticulate, no apical process as in
P. thoas. ?. Edge of vaginal orifice anteriorly raised into a small, smooth
trtbercle, and posteriorly into a transverse ridge ; from this ridge extends on each
side backwards a longitudinal ridge bordering a large mesial depression ; just
in front of the orifice a fold which is laterally raised into a small triangular Libe.
Anal segment on inner surface with tliree bristles on each side.
Early stages similar to those of P. thoas.
Hah. Canada to Costa Rica ; Cuba.
In the Tring Museum 66 $$, 54 ?9 and some larvae and pujiao from:
Buffalo; Evanston, and St. Augustine, Illinois (Snyder); Nashville, Tennessee
(W. Osburn); Iowa ; Jefferson Co., Kentucky (Troxler); Louisiana; Florida ; Cuba;
( 506 )
Houston, Texas; Orizaba and Huatnxco, Vera Criiz ; Espiual, Vera Crnz, July
1896 (W. Schaus) ; Guadalajara, September — October (Dr. Butler); Gnadalajara,
Jnly 1896 (W. Scbaus) ; Hermosillo, Sonora, March 1903 (Oslar) ; Guerrero
(O.T. Baron); S. Pedro Sula, Honduras; ('arillo, Costa Rica, 3000 ft., October
1904 (A. Hall) ; Azahar de fartago, Costa Bica, February ls9i) (Underwood).
09. Papilio paeon Boisd. (1830).
Papillon pdon Roger, Bull. Sue. Lviii. B"nleiiii.r i. p. 101 (182li) (Chili).
Pupilio iwtmt Boisduval, f^pcc. Gen. Up. i. p. 350. n. U)« (IR.'IO) (Chili) ; Walk., Ent. .1/". Mag. xi.\.
p. 53 (1882) (Callao ; larva and pupa).
(J ? . Sexes similar, the female being lari;i'r and paler tliau the male. Forewing
resembling that of P. tlioas. Hindwing, below, with a row of rufous red spots
around cell ; apex of cell bearing a rufous red spot bordered distally by a black
crescent, which is often double, the interspace between the two black lines and
the tip of cell being buffisb-yellow ; tail without yellow spot in centre ; marginal
spot behind tail large.
Neuration : SC^ of forewing strongly curved at base, stalk of SC'* shorter than
the two cross-veins D- and D^ together, D^ very oblique, sometimes almost in the
same direction as D^, lower angle of cell very obtuse.
Genitalia: cJ. Tenth tergite long, narrowing apicad, apex rounded; sternite
laterally incrassate in middle, the incrassation divided transversely into an anterior
and a posterior toothlike ridge. Clasper rounded ; harpe broad, rounded distally,
bearing several long thornlike teeth at the edge. ?. Edge of vaginal orifice
anteriorly in middle raised into a tubercle which is carinate on the anterior and
convex on the posterior side ; behind the orifice a rounded tnbercle which is densely
covered with minute hairs ; behind this tubercle two folds extending forward into
the orifice ; in front of the orifice on each side a broad shell-like ridge, with
irregular edge, and laterally of this another smaller ridge ; on the posterior surface
of the large ridge stands laterally of the orifice a long, slender, pointed, thorn-
like process. Anal segment on inner surface with ibnr bristles on each side (tliis
number constant ?).
Early stages described by Walker, I.e.
Ilab. Columbia and Northern Venezuela southward to Bolivia.
There is no reliable evidence that the species occurs in Chili.
Two subspecies :
a. P. paeon thruson Feld. (1805).
Papilio paeon, Gray, Cat. Lep. Ins. Brit. Mas. i. Pap. p. 39. n. 197 (1852) (parlim ; Bogota) ; id.,
Li-st Lep. Itis. Brit. Miis. i. Pap. p. 54. n. 207 (1856) (^partim) ; VoUenh.. Tijilsriir. Ent. iii.
p. 8fl. n. 140 (18G0) (Bogota) ; Obertb., Et. d'Ent. iv. p. 70. n. 208 (18S0) {jmrtin, ; Muzo ;
Carare ; R. Magdalena).
Papilio thrasson Felder, Vcrh. Zool. Bot. Ges. Wkn xiv. p. 309. n. 292 (18(;4) (num. mid. ; Bogota ;
Venezuela) ; id., Reise Norara, Lep. p. 74. n. 57 (1805) (Venezuela ; Bogota) ; Hahnel, Iris iii.
p. 201 (1890) (Valera, Venez.) ; Godm. and Salv., Biol. Ceidr. Amcr., Lep. liliop. ii. p. 224.
n. 54 (1890) (Costa Rica) ; iid., in Whymper, Amlex of Equator, App. p. 109. D. 96 (1891)
(Nauegal).
Papilio piirou var. a. /'. Ihrason, Kirby, Cat. Dinru. Lep. p. 542. sub n. 1.57 (1871) (New Gran-ada).
Papilio proii \aT. lliraso!i,M.aass. AWeym., in SlUbel, liiisen S. .Imer , Lep. p. 14. n. 14(1890)
(Honda to Bogota); iid., I.e. p. 38. n. 33 (1890) (Pop-iyau).
cJ?. Upperside : submarginal spots of forewing usually linear, thin.
Hindwing rather longer posteriorly than in the following form ; yellow marginal
(567 )
spot beliind tail reaching to tliree-fourtbs of tail. Orange-red markings on
underside of bindwing smaller than in P. p. paeon ; the two black crescents
M' — M^ more widely separated ; black marginal line not so broad, not so strongly
arched between the veins, and not interrupted before tail ; the latter longer. — —
Harpe usually with fewer teeth.
Ilab. Colombia and Northern Venezuela ; Costa Rica (von Patten).
As the specimens in von Patten's Costa Rica collection were not all from
Costa Rica, it is possible that also the present insect got into the collection by
some mistake.
In the Tring Museum 28 c? c?, 0 ? ? , from : R. Dagua, W. (Colombia (Rosenberg) ;
" Bogota" ; Pereira, Cauca ; Merida, Venezuela (Briceilo).
h. P. paeon paeon Boisd. (1836).
PapiUo piieuii Boisduval, I.e. ; Doubl., Westw. & Hew., Gen. Diurn. Lep. i. p. 17. n. 172 (I84G)
(Chili) ; Blanch., in Gay, Hist. Fis. Chile, Zool. vii. p. 8. d. 2 (1852) (Chile ?) ; Gray, Cat. Lep.
Ins. Brit. Mus. i. Pap. p. 39. n. 197 (1852) (^partiin ; "Chili") ; id., List Lep. Ins. Brit. .!/«». i.
Paj). p. 54. n. 207 (1856) {partim) ; Felder, Verli. Zool. But. Gcs. Wien xiv. p. 309. n. 291 (1H64)
(Peru ; Chile ?) ; Kirby, Cat. Diurn. Lep. p. 542. n. 157 (1871) (var. excl. ; " Chili " ; Pern) ;
Burm. , Z)esc)-. Rep. Argent, v. Lep. p. 58. n. 4 (1878) (var. of thoas\); Ropd. , Steft. Ent.
Zeit. xl. p. 49. n. 4 (1879) (Peru) ; Obertb., Et. d'Eiit. iv. p. 70. n. 208 (1880) {partim ; Peru) ;
Walker, Ent. Mn. ^[ag. xix. p. 53 (1882) (Callao ; larva on parsnip, descript. of various stages
and pupa) ; Bartl.-Calv., Cat. Lep. Chile p. 5. n. 2 (1881)) (doubtful as Chilian).
Papiliopeon, Staudinger, Erut. Tagf. p. 16 (1884) ; Maass. & Weym., I.e. p. 72. n. 7 (1890) (Tambo
de Chillo, 3000—3600 m.) ; Dognin, Lep. Loja p. 15 (1887) ; id., I.e. p. 37 (1891).
Papiliii Ihrastin, Weeks {non Felder, 1865, err. det.), IlUtstr. Diurn. Lep. p. 20 (1905) (Chuluinani).
c??. Yellow marginal spot behind tail extending about half-way to ape.K of
tail. Snbmarginal spots of underside of forewing more or less rounded, small ;
black marginal line of hindwing broad, strongly arched between the veins ;
orange-red spots large. Harpe usually with four teeth or more.
Ilab. Ecuador to Bolivia ; the locality " Chile " of the older writers doubtless
erroneous.
In the Tring Museum 2'i <S <S , 3 ? S, from: Chimbo, N.W. Ecuador, July
1897, 20UO ft. (Rosenberg) ; Ibarra, Blay 1897 (Rosenberg) ; Ambato : Zamora
(0. T. Baron); Chosica, Peru, 850 m., January 1900 (Simons); Hnancabamba,
Cerro de Pasco (E. Boettger) ; R. Mi.\iollo, Loreto (Baer); Chanchamayo (Schunke);
Carobas, Peru, 2-500 ft., December 1899 (Simons); La Merced, Clianchamayo
1000 m., April 1900 (Simons); Chulnmani, Bolivia, 2000 m., January 1901
(Simons) ; R. Tauampaya (Garlepp) ; Yungas de La Paz (Garlepp) ; R. Unduawe,
Bolivia, 2000 m., February 1901 (Simons).
TO. Papilio caiguanabus Poey (1851).
Papillon perithous Roger, Bull. Soc. Linn. Bordeaux i. p. 159 (1826) (Cuba).
Papilio caigmumhus Poey, Mem. Hist. Nat. Cuba i. p. 442. t. 15. fig 1. 2. $ , 3. 4. (J (1851) ; Gray,
Cat. Lep. Ins. Brit. Mus. i. Pap. p. 38. n. 186 (1852) (Cuba) ; id., List Lep. Ins. Ilril. Jliis. i.
Ay), p. 52. n. 194 (1856) (Cuba); Lucas, in Sagra, Hist. Cuba vii. p. 207 (18,57) ; Weidem.,
Proc. Eut. Soc. Philad. ii. p. 146 (1863) ("West Indies") ; Felder, Vcrh. Zool. Hot. Ges. Wien
xiv. p. 309. n. 286 (1864) (Cuba ; = niimicus) ; Herr.-Sch., Corresp. Bl. Zoul. Min. Ver. liegcusb.
p. 173. n. 8 (1864) (Cuba) ; Kirby, Cat. Diurn. Lep. p. 542. n. 162 (1871) (Cuba) ; Oberth.,
Et. d'Ent. iv. p. 69. n. 202 (1880) (Cuba) ; Gundl., Cuntr. Ent. Cuba p. 127 (1881) ; id., Pajnlio i.
p. 113. n. 162 (1881) (Cuba) ; Haase, Uiitersuch. Mimicryi. p. 98(1893) (Cuba ; " near relative
of P. erostratus " false).
Papilio nuiniius Hopffer, Neue Hchnult. p. 1. n. 2. t. 1. fig. 3. 4 (1856) (Cuba) ; Hew., Ej:ot. Hull. iii.
Paj). t. C. fig. 17 (1864).
( 568 )
Roger's nomenclatnre being French, not Latin, his name cannot be accepted
for this species. Boisduval (1830) erroneonsly applied the name " pirithous
Roger" to the female of a very different insect (/'. hjrophron).
S ? . Nearest to P. thoas and aristor ; tlie discal markings of the uppcrside
have nearly all disa])peared, while the submarginal ones are large, the latter being
opalescent white on the hindwing of the female. On the underside of the hindwiug
there arc two red spots at cell as in P. tkoas, and a row of blue discal spots as in
r. aristoJemus and tlwns.
■ Genitalia : 6- Tenth tergite long, narrowing apicad, slightly constricted just
before apex ; sternite armed at each side with a large, pointed, dentate j)rocess ;
harpe short, broad, rounded, ending in a short point. ? . Anterior edge of
vaginal orifice raised into a smooth tubercle ; laterally of the orifice a large
depression, the lateral edge of which is elevated, standing posteriorly in connection
with the postvaginal ridges ; behind the orifice a quadrangular groove with
elevated edges, laterally of this strongly chitiuised groove a ridge which ends
anteriorjy near the hinder edge of the orifice in a truncate denticulate process.
Early stages not known.
Hab. Cuba.
In the Tring Museum 7 JcJ, 7 ? ? , from : Gibara and Hok^nin (ToUiu).
71. Papilio aristor Godt. (1810) (PI. V., fig. 21).
Papilio ai-islor Godmt, Eur. MM. ix. p. GO. n. 95 (18111) (hab.?) : Boisd.. Sppc. Gin. Lip. i. p. 51.
n. 192 (183C) (type-specimen with three wings in Mus. Paris) ; Doubl., West. & Hew., Gen.
Diurn. Lfp. i. p. IG. n. IGO (184G) ; Gray, Cat. Lcp. Ins. Biil. .ir»s. i. Pap. p. 37. n. 183 (1852);
id.. List Lep. Ins. Bnl. .Mus. i. Pap. p. 51. n. 192 (185G) ; Felder, Verh. Zoul. Bid. Gcs. Wini,
xiv. p. 315. n. 360 (18G4) ; Kirby, Cat. Diurn. Lrp. p. 5G6. n. 324 (1871) ("Mexico? Antilles?").
The type-Specimen of this species appears to have been destroyed. According
to Boisduval, I.e., the specimen e.xisted in the Paris Museum when he wrote the
Species General, but it is no longer in that collection — at least we have not found
it there. The only specimen known to us is in the magnificent collection of Mons,
Charles Oberthiir, who kindly lent it to us for figuring.
(?. Abdomen black, with a subdorsal and a snbveutral row of buff-yellow spots,
the claspers and the eighth segment being more extended yellow.
Underside of wings similar to upper, spots paler, anterior discal ones of
forewing and submarginal ones of hindwing larger ; forewing with buff subapical
cell-patch, which does not extend across the cell ; hindwing : the discal spots
C — R' yellowish, vestigial in the sjiccimen, the first indicated also on upperside,
traces of two orange-red spots R- — M' at cell, which were ajaparently mure distinct
in Godart's specimens ; a row of small blue spots proximally of the submarginal
spots.
Nenration : cell of forewing as in /'. euii/uanadtis, asymmetrical, the lower
angle being very obtuse.
Genitalia : c?. Tenth tergite long, curved downwards at ajjcx, spatnlate :
sternite at each side with a long, pointed tooth bearing proximally a rounded
hairy projection ; harpe broad, excavated, suddenly narrowed to a point, dentate
distaliy at ventral edge.
Female and early stages not known.
Hub. Port-aii-Piince, Haiti (F. Odile Josej)h), 1 <S in coll. Oberthiir.
( 569 )
72. Papilio aristodemus Esp. (1794).
Papilio Eques Acliivui: aristodemus Esper, Macja:. Neucst. Aiisl. Iiis. p. 8. t. 2 (1794); id., Ansl.
Schmett. p. 240. n. 113. t. .59. fig. 2 (1798) (" Cuba" false).
Papilio aristoilemus, Boisduval, .S/jcc. Gen. Lep.i. p. 357. n. 199 (1830); Vollenh., Tijdschr. Ent. iii.
p. 86. n. 141 (1860) (Antilles).
Papilio daphnis Gray (ex Martyn, Psi/clte, errore), Cat. Lep. Ins. Brit. JUiis. i. Pap. p. 39. n. 198
(1852) (partim) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 54. n. 209 (185G) (nynon. 2>artint ;
S. Domingo); Weidem., Proc. Ent. Snr. Philad. ii. p. 147 (18(53) (West Indies, partim ;
" Mexico " false).
Papilio crcsj)honfi?nis Martyn, Psiiche (ined.), t. 3. fig. 8, t. 4. fig. 10. (1797) ; Kirby, Cat. Biiirii.
Lep. p. 542. D. 168 (1871); Burm., Dcscr. i.V>. Arrjent. v. Lep. p. 58. n. 2 (1878) (var. of */toa.s !) ;
Gundl., Contr. Eut. Cuba. p. 130 (1881) (Cuba ; S. Domingo ; Porto Rico ; "Mexico" false).
S ? . Antenna yellow at base of club in most specimens.
Underside of both wings more extended yellow tban in P. thoas ; cell of
forewing not striped with black ; cell of hiudwing either all yellow or bearing
a small black spot at apex; yellow submarginal spots of both wings merged
together to a band which is anteriorly on forewing not separated from the basi-
discal area ; black discal band of hindwing narrow, bearing a row of large bine
spots, and being bordered proximally by dirty rufous red scaling, which is some-
times absent or vestigial. Tail black on upperside, edged behind with yellow
from base to near apex, beneath more or less yellow in middle.
Genitalia : S. Tenth tergite spatulate ; sternite with small, obtuse, double
ridge at the sides ; harpe long, acuminate, nearly straight, flat, denticulate at
apex, armed with a very long, thorulike process ventrally before the middle,
homologous to the process of P. lycopkron. ? . Armature of the same type as
in P. lycophron ; edge of orifice anteriorly raised into a compressed tubercle which
extends into the orifice, lateral edge elevate, thin, abruptly ending posteriorly ;
behind orifice a membranaceous tubercle densely covered with minute hairs ;
laterally of the orifice a large lobe bearing numerous long, thornlike teeth at
the edge, and further laterad a small chitinised depression, the free external
edge of which is rounded.
Early stages not known.
Hah. Cuba ; Haiti.
Two subspecies.
The sjjecies connects P. thoas with P. li/cophron and allies.
a. P. aristodemus temenes Godt. (1819).
Papilio temenes Godart, Euc. Mith. ix. p. 63. n. 104 (1819) (" Antilles and North America") ;
Oberth., Bull. Sue. Ent. France p. 176. fig. 4 (1897).
Papilio tecmenes (!), Lacordaire, A7vi. Soe. Ent. Fr. ii. p. 384 (1833).
Papilio aristodemus, Boisduval, I.e. (1836) (= temenes) ; Poey, Mem. R. Soc. Econ. Hubana p. 235
(1846) ; Lucas, in Sagra, Hist. Cuba vii. p. 206. t. 16. fig. 2. 2a (1857) (jmrtim ■ Cuba) ;
Herr.-Sch., Corre-yi. Bl. Zool. Uin. Ver. Rerjensb. p. 174. n. 12 (1864) (Cuba).
Papilio daphnis Gray, /..-. [partim) ; Felder, I.e. xiv. p. 309. n. 290 (1864) (partim ; Cuba).
Papilio rresjdionli/iiis, Doubl., Westw. it Hew., Gen. Diurn. Lep. i. p. 1-7. n. 173 (1846) (= Imtenes) ;
Kirby, Cat. Diurn. Lep. p. 542. n. 158 (1871) (partim) ; Gundl., Contr. Ent. Cuba. p. 130(ISK1)
(partim ; Cuba) ; id., Papilio i. p. 113 (1881) (Cuba).
This form has always been treated as being identical with the Haiti form until
Mons. Charles Oberthiir, I.e., pointed out the differences.
(J ¥ . Forewing : discal band broad, of nearly even width from II- to hiud-
( 570)
margin, about as bn ad as tbe cell is wide at M- ; a row of five to seven snbinarginal
sjiots.
Ilab. Cuba.
lu tbe Triug Museum 1 <?, 2 ? ?, iVom : Gibara (Tollin) ; Cuba (Gundlacb).
h. r. aristodenms aristodemus Esp. (1794).
I'ltpii'ii) Eqiws Achirits anstodcinds Espcr, l.c-
I'tijiiliii arislihlennis, Boisduval, /.(■. (piirlim) ; Doubl., List Lep. Int. Brit. Miia. i. Ajipniil. p. 4(1S4S)
(Haiti?) ; Dewitz, Sictl. Ent. Zeit. xxxviii. p. 234. n. 2 (1877) (Porto Rico, common, Aug.
Sept.) ; Oberth., El. d'Eiit. iv. p. 70. n. 207 (1880) (Haiti).
rapilio claplims. Gray, ll.cc. (jiarlim) ; Felder, Vrrh. Zuul. But. Gi's. Wkn xiv. p. 30'J. n. 290 (1864)
{partiiii ; Haiti).
Papilio cresphonliiiiis, Martyn, l.c. (inedit.) ; Kirby, Oil. Dinrti. Lep. p. 542. n. 158 (1871) (parlim) ;
Mfischl., Ahh. Snikeiib. Nut. (?cs. xvi. p. 91. n. 2 (188G) (= arixtuiletni/s ; Porto Rico ; common);
Gundl., A)i. Hht. Xal. Madrid xx. p. 114. n. 2 (1891) (Porto Rico).
d ? . Band of forewiug mncb narrower tbau iu tbe previous subspecies,
interrupted at tbe veins ; a row of four submargiual spots, the row much more
cnrved tban in temenes, s])ots U- — (SM') standing farther away from the edge of
the wing tban in tbe Cuban form.
Ilab. Haiti.
In tbe Tring Museum 5 Jc?, 1 ¥ from Haiti.
73. Papilio andraemon Hiibu. (IslS— ?).
Ilcmdidi'S umlmemuii Hubner, Samml. Exot. Sehmett. ii. t. 98. (J, 99. ? (1818— ?).
Papilio aiidmemoii, Boisduviil, Spec. Gen. Lep. i. p. 343. n. 183 (183l>)(Cuba) ; Doubl., Lixt Lej>. [m.
Brit. Mils. i. p. IG (1845) (" Honduras " error luci) ; Gray, Cat. Lep. Ins. Brit. Mas. i. Pap.
p. 24. D. 108 (1852) (" Honduras" false) ; id., Lixl Lep. Lis. Brit. Mus. i. Pap. p. 32. n. 115
(1856) ("Honduras" false); Weidem., Pruc.. Ent. Soc. Philad. ii. p. 146 (1863) ("West
Indies," " C'entr. Amer.") ; Godm. & Salv., Biol. Ccntr. Amer., Lep. Rhop.M. p. 243 (1893)
(occurrence in Mexico and Honduras requires confirmation).
c? ¥ . Club of antenna for the greater part yellow beneath in male, also in
female partly yellow or at least tawny. On upperside of wiugs a nearly straight
yellow band from apex of forewing to abdominal margin of hindwing, the cross-
veins of the hindwing being within the baud ; a bar across cell of forewing at
point of origin of R' and a short band from base of subcostal fork to costal margin
also yellow, contiguous with the discal band. Hindwing very strongly toothed,
tooth M' long ; tail spatulate, bearing a yellow central spot near apex.
Underside of hindwing washed with yellow from base to yellow discal band,
this band deeply crenate distally, being followed by a broad black band bearing
a row of blue spots ; a large tawny patch R' — M' proximally of tbe black band
often preceded and followed by a small tawny spot.
Genitalia : c?. Tenth tergite narrow, spatulate ; sternite feebly chitinised,
without distinct lateral process, bearing only an obtuse double ridge ; harpe broad,
somewhat scythe-shaped, denticulate distally, produced ventrally into a long acnte
conical process directed downwards and inclining a little basad. ¥ . Edge of
vaginal orifice raised into a transverse rounded lobe ; behind the orifice a small
rounded tubercle covered with extremely small hairs ; at each side a very largo
non-dentate lobe which is anteriorly connected with the vaginal lobe by means of
a low folded ridge.
Early stages described by Gundlach, I.e. : see /'. a. andraemon.
Hab. Cuba ; Bahamas ; Great Cayman.
Three subspecies.
( 071 )
a. P. a.ndraemon andraemon Hiibn. (1818 ? — ).
Eeraclides andraemon Hiibner, I.e. ; Kirby, in Hiiba., Samml. E.rnt. Sehme/l. eJ. ii, p. ilC. t. 3ll.
6g. 1. 2. t. 312. fig. 3. 4 (190—?) (Cuba ; " Mexico, Guatemala," false).
rapiVio (i/«/)'apnio«, Boisduval, /.f. ; Poey, Mem. R. Soc. Econ. ffahniia p. 23.") (184ri) ; Doubl., We.stw.
& Hew., Geii. Dhirii. Lep. i, p. 13. n. 87 (184G) (Cuba ; " Hondura.s " false) ; Lucas, in Sagra,
Uisl. Ciiha vii. p. 203 (1857) ; Minitr., Enum. Corp. Aiiim. Mus. Pelrnp., Lip. i. p. Cs. n. 1115
(1857) (" Honduras " false) ; Vollenh., njdsehr. Eul. iii. p. 85. n. 1.32 (1860) (Cuba) ; Felder,
Verh. Zoo}. Bot. Ges. Wieii xiv. p. 309. n. 287 (18G4) (Cuba ; "Mexico and Honduras" false) ;
Herr.-Scb., Corresp. Bl. Zool. Miii. Ver. lierjeush. p. 172. n. 1 (1864) (Cuba) ; Kirby, Cat. niurn.
Lep. p 542, D. IGl (1871) (Cuba ; "Mexico" false) ; Oberth., Et. d'Ent. iv. p. 69. n. 205(1880)
(Cuba) ; Goudl., Coiitr. Eiil. Cuha. p. 128 (1881) (larva and pupa descr.) ; id., Papilio i. p. 11 !
(1881) (Cuba) ; Haase, Unters. Mimicry i. p. 95 (1893) (Cuba ; " Mexico and Honduras " false)
c? ? . Forewing without submarginal spot?, or the last one or two vestigial ;
yellow discal band as broad (or nearly) as the black marginal area.
Hab. Cuba.
Ill the Tring Museum 14 Jc?, 9 ? ? , from : Hohiuin (ToUin); Cuba (CJundlach).
b. P. andraemon bonhotei Sharpe.
Papilio honhnfci Sharpe. Proc. Zool. Soc. Loud. p. 201. t. 19. (J ? (1900) (Nassau).
c? ?. Yellow markings paler than in the previous ; discal band much narrower;
forewing with a variable number of thin submarginal spots on nppersido.
Hal). Bahamas : Nassau.
In the Tring Museum 5 cJc?, 3 ? ? from Nassau.
c. P. andraemon tailori snbsp. nov.
S ?. Discal band as broad as in the Cuban form ; bar across cell of forewing
vestigial above, while it is wider below than in the Cuban form ; black antemediau
band of underside of forewing more curved, and black discal patches of hiudwing
smaller than in P. a. andraemon.
Genitalia : $ . Harpe much narrower than in Cuba specimens.
Hab. Great Cayman Island.
One pair in the Tring Museum, collected in April 1S96 by Mr. Taylor.
74. Papilio machaonides Esp. (1796).
Papilio Eques Aeliivm machaonides Esper, And. Schmetl. p. 191. n. 86. t. 46. fig. 2 (1796) (Port au
Prince).
Papilio li/coraeus Godart, Enc. Melh. ix. p. 63. n. 105 (1819) (America) ; Mdne'tr., Nonv. Mem. Soc.
Imp. Moscoti. iii. p. IIG. n. 2. (18.32) (Haiti).
Papilio lycoroeus (I), Lucas, Lep. Exot. p. 34. t. 18. fig. 1. (1835).
Papilio machaonides, Boisduval, Spec. Gen. Lip. i. p. 344. n. 184 (1836) (Haiti ; = h/coraeus) ;
Doubl, Westw. & Hew., Gen. Diuru. Lep. i. p. 13. n. 86 (1846) (Haiti) ; Doubl., List Lep. Ins.
Brit. Mus. i. Append, p. 4 (1848) (Haiti) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 24. n. 107
(1852) ; id.. List Lep. Ins. Brit. Mus. i. Pup. p. 31. n. 114 (1856) (Haiti) ; Lucas, in Sagra, Hist.
Cuba vii. p. 206 (1.S57) (" Habana'' false); Me'netr., Enum. Corp. Anini. J/i/.t. Petrnp., Lep. i.
p. 2. n. 29 (1857) (Haiti) ; Vollenh., 'Tijdsrhr. Ent. iii. p. 85. n. 134 (1860) (Haiti) ; Weidem.,
Proc. Ent. Soc. Philad. ii. p. 147 (1863) (" West Indies ") ; Felder, Verh. Zo,d. Bot. Ges. !('» „
xiv. p. 309. n. 289 (1864) (S. Domingo ; " Cuba" false) ; Kirby, Cat. Diurn. Lep. p. 542. n. 159
(1871) ("Antilles") ; Oberth., Et. d'Ent. iv. p. 70. n. 206 (1880) (ftiiti) ; Haase, Untersu h.
Mimicry i. p. 94 (1893).
(??. An interrupted discal baud on forewing ; the posterior portion of tiic
baud continuous with a broad cell-bar ; upper portion of band extending from
costal margin to R- ; a complete series of submarginal spots, the first spot
37
( 572 )
corres])OD(ling to the first spot of the so-called discal baiul of P. thoan aud
P. andraemon.
Underside, of hindwing yellow from base to middle of disc ; no tawny or red
spots at apex of cell.
Genitalia : c?. Tenth tergite narrow, long, spatnlatc ; sternite feebly chitinised,
with a small double ridge on each side ; harps narrow, ending in a long point,
angnlate dorsally before middle, bearing a few teeth at the edge distally of this
angle, there being also a small tooth at the ventral edge near base. ?. Anterior
edge of vaginal oritice raised into a longitndinal tubercle accompanied on each side
by a fold ; these folds diverging laterad, each bearing on the inner side a long,
narrow process armed with a few small thornlike teeth ; behind the orifice a
glossy, rounded tubercle clothed with extremely small hairs.
Early stages not known.
Ilab. Haiti.
So far this fine species has not been fonnd on the other West Indian islands,
where it appears to be replaced by P. andraemon, which does not occur on Haiti.
In the Triug Museum 4 c?(?, 2 ? ? , from Haiti.
75. Papilio thersites Fabr. (1775).
(J. PiijiilioEqiies Aefiivus (AcrsiVesFabricias, Sjs<. £«/. p. 453. n. 43 (1775) (America ; mus. Hunter):
Gotze, Eiii. Bei/lr. iii. 1. p. 73. n. 9 (1779) ; Fabr., Spec. Jns. ii. p. 18. n. G8 (1781) ; id., Mant.
Lis. ii. p. 9. d. 78 (1787) ; GmeUn, Sijst. Nat. i. 5. p. 2238. n. 313 (1790) ; Fabr., E»t. Sijst. iii.
1. p. 30. n. 88 (1793).
J . I'miiliii Equea Afhiuis jialaiiiedes Fabricius {unit Drury, 1770), Syst. Ent. p. 454. n. 45 (1776)
(America ; mus. Hanter) ; Goeze, Ent. Bci/lr. iii. 1. p. 73. n. 11 (1779) (partim) ; Fabr., Spec.
/)M. ii. p. 18. n. 73 (1781) ; id., Maid. Im. ii. p. 10. n. 84 (1787) ; Jabl. & Herbst, Nalurs.
Schmett. iii. p. 141. sub n. 94 (1788) ; Gmelin, Syst. Nat. i. 6. p. 22.39. n. 320 (1790).
5 . Piqiilin Eijues Trojaiius acama.'i Fabriciu.s, Ent. Sij-it. iii. 1. p. 8. n. 22 (1793) (Jamaica ; Drury).
? . Papiliii NiimphaliiiptalamciJes id., Ent. Sysl. iii. 1. p. G8. n. 213 (1793).
5 . Papiliii animas, Godart, Em; Aleth. ix. p. 42. n. 60 (1819) ; Donov.. Xat. Repos., Ent. ii. t. 18
(1823) ; Boisd., Spec. Gen. Up. i. p. 360. n. 203 (18.30) (Jamaica) ; Doubl., List Lep. Ins. Brit.
Mas. i. 2. p. 17 (1815) (Jamaica?) ; id., Westw. & Hew., I.e. p. 17. n. 170 (1840) (Jamaica).
? . Papiliii palaniedcs, Godart, I.e. Stippl. p. 812. n. 133-4 (1824) ; Boisd., I.e. p. 359. n. 202 (1836
(=j,irillwus?) ■ Doubl,, Westw. & Hew., I.e. p. 21 (1847) ; Gray, Cat. Lep. Lis. Brit. Mas. i.
Piiji. p. 72 (1852) (doubtful species) ; id., Li.it Lep. Lis. Brit. .Uns. i. Pap. p. 85 (1850).
(J. J'apilia lliersites, Donovan, Nat. Repms., Ent. ii. t. 24 (1823) ; Boisd., /.'•. p. 353. n. 195 (183G) ;
Doubl., Westw. & Hew., I.e., p. 17. n. lOG (184G) ; Doubl., List Lep. Lis. Brit. Mns. i. Append.
p. 4 (1848) (Jamaica).
<J ? . Papiliu thersites, Hewitson, Trans. Ent. Sue. Loud. (2). i. p. 97 (1851) (cj of ncama-i) : Doubl.,
Westw. & Hew., I.e. ii. p. 529 (1852); Gray, Cat. Lep. Ins. Brit. .Mus. i. Pap. p.SS. d. 191
(1852) ; id., List Lep. Lis. Brit. Mus. i. Pap. p. 52. n. 200 (1856) (Jamaica) ; Lucas, in Sagra,
m.'it. Cuba vii. p. 205 (1857) (Jamaica; "Cuba" false) ; Weidem., Pruc. Ent. Sue. P/iilad.
ii. p. 148 (1863) (" West Indies") ; Kirby, Cat. Diiirn. Lep. p. 639. n. 149 (1871)("Antilles") ;
Butl., Pruc. Zwil. Sue. Loud. p. 481. n. 38 (1879) (Jamaica) ; Oberth., El. d'Ent. iv. p. 71. n. 213
(1880) (Jamaica); Haa.se, Untersuch. Mimien/ i. p. 97 (1893) ("Antilles"); Towns., Joimi.
Inst. Jamaica i. p. 370 (1893) (larva) ; Fox & Johns., Ent. Neirs iv. p. 3 (1893) (Jamaica) ;
Robins., ibid. xiv. p. 20 (1903) (Jamaica).
J?. Papilio aeamas, Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 311. n. 310 (1804) {partim;
Jamaica) ; Butler, Cat. Diurn. Lep. descr. Fabric, p. 24G. n. 45 (1H69) (types of 2>'tlaniedes
Fabr. nun Drury, and tliersilcs in the Hnnterian coll. at Glasgow).
S ? . Close to /'. li/cophron. The male is easily recognised by the great width
of the yellow band of the forewing, the ])rulongation of the cell-patch halfway down
to base and the absence of submarginal spots from the upperside of the forewing.
In the female there is a curved yellow baud on the forewing ; the submarginal
(5?3)
spots of the bindwing are large on the nnderside, being usaally merged togethei'
with the marginal spots.
Genitalia : similar to those of P. lycojphron. Rasplike ridge of harpe raised,
forming a narrow process which is ronnded at the apex.
The caterpillar closely resembles that of P. androgens.
Ilab. Jamaica.
In the Tring Museum 8 c?(?, 2 ? ? and 1 larva.
70. Papilio ornythion Boisd. (1830).
(J. PiijiHin orm/lhloii Boisduval, Sper. Gen. Lqi. i. p. 354. n. 197 (1830) (Yucatan) ; Doubl., Westw.
& Hew., Gen. D'mrn. Lep. i p. 17. n. 171 (1846) ; Gray, Cat. Lep. Inx. Brit, Mm. i. Pup. p. 3'.).
n. 195 (1852) ; id., List Lep. Ins. Brit. Mm. i. Pap. p. 54. n. 205 (1856) ; Weidem., Proc. Ent.
Sue. Phihul. ii. p. 147 (18C3) ; Felder, Verh. Zonl. Bot. Ges. Wien xiv. p. 310. n. 293 (1864) ;
Kirby, Cat. Diurn. Lep. p. 541. n. 156 (1S71) ; Burm., Descr. Rip. Argent, v. Lep. p. 58. n. 3
(1878) (var. of thoas) ; Godm. & Salv., Biol. Cenlr. Amer., Lep. Rhop. ii. p. 227. n. 57. t. 09.
fig. 7. 8. cJ (1890) (fig. of type) ; iid., I.e. p. 729 (1901) (Coatepec ; Guatemala).
(?. Close to P. hjcophron pallas. Yellow band narrow, no spot in cell of
forewing ; on underside of forewing an additional row of linear spots between
the discal band and the submarginal spots ; bine spots on nnderside of hindwing
large.
¥. Yellow markings of upperside vestigial. Underside, forewing: a row
of submarginal sjwts ; ill-defined streaks between costal margin and R-, from which
two postdiscal spots SC^ — SC° are separated ; a series of minute postdiscal spots
from SC^ to M^ ; yellow cell-streaks vestigial. Hindwing : submarginal jjatches
large, blue crescents larger than in P. I. pallas ; tail as long as in male, slightly
spatulate.
Genitalia of male as in P. I. pallas.
Hah. Yucatan ; West Mexico ; Guatemala.
In the Tring Bluseum 1 cJ, 1 ?,from Guadalajara.
77. Papilio lycophron Hiibn. (1818—?).
Seba, Thcsaur. p. 13. t. 8. fig. 17. 18. ? (1764) (" India ").
(J. Heraclides lycophron Hiibner, Samml. Exol. Schmett. ii. t. 100 (1818 — ?).
(J. Papilio astyaliis Godart, Euc. MUh. ix. p. 62. n. 102 (1819) (Brazil).
$ . Papilio pirithous Boisduval, S2)ec. Gen. Lep. i. p. 358. n. 201 (1836) (Uruguay ; " Cuba " error
loci).
(??. Closely agreeing with P. androgens in structure. Submarginal spots
on underside of forewing larger, those of hindwing much larger above and below ;
tail usually somewhat spatulate, teeth R^ and M' short. ? dicliromatic in
some districts.
Genitalia nearly the same as in P. androgens, geographically variable, while
they are not geographically variable in P. androgens ; rasplike ridge of harpe more
dorso-ventral in direction than in P. androgens, sometimes developed to a process.
? . Lateral dentate lobe of vaginal cavity narrower at base and wider at apex
than in P. androgens.
Larva similar to that of P. thoas, more strongly marmorated, the patches
yellow.
Pupa slenderer than in P. thoas, the thoracic tubercle longer. On Citrus.
Ilab. Mexico southward to Argentina and Uruguay ; Santa Lucia.
(574)
We are Dot sure tbat Hiibner's name has priority over that of Godart.
The Santa Lncia sjieciiueus recorded by Miss Sbarpe (Proc. Zool. Soc. Loud.
1901. p. 223), if lieloDging to this species, represent doubtless a distinct subspecies.
Boisdnval, I.e., records a form of this species from Cuba under the name of
P. pirithous Eoger, several other authors (Doubleday, Lucas, Felder, Kirby, etc.)
following suit. Gnndlach (1881) rejects the species as Cuban. There is indeed
no evidence that a form of/', h/cophron occurs on that island, though Lucas (185T)
records not only P. t/tersites, but also two forms of the present species from there —
namely, pirithous and li/rophron, the latter being stated to be common on the
island. The mistake began with Boisduval, who mistook some South American
females of P. hjcophron for Eoger's " Papillon pirithous." This " Pajiillon piri-
thous " being from Cuba, the locality of specimens erroneously identified as
pirithous was accepted to be also Cuba. The description given by Roger has
doubtless never been carefully read, since nobody appears to have noticed that it
is a descriiitiou of the male of P. caiguanabus !
It is quite possible that a form allied to P. thcrsites or P. Ujcophrou will be
discovered on Cuba or Haiti.
Seba's figure. I.e., represents doubtless a form of the present species. However,
we have not seen a specimen agreeing with the figure. Seba's type came possibly
from Surinam.
a. P. hjcophron pallas Gray (1852).
J V • Papilio oehalus, Gray (iion Boisd., 1836, err. det.), Cai. Lep. Ins. Brit. Mus. i. Pujy. p. 39.
n. 193. t. 6. fig. 1. (? (1852) (Mexico) ; id., List Lep. Ins. Brit. Mm. i. Pap. p. 53. n. 203 (I85G) ;
Weidem., Pm: Ent. So,: Ph'dad. ii. p. 147 (18G3) ; Felder, Verh. Znnl. Hot. Gcs. ]Vieii xiv.
p. 311. n. 311 (181)4) (Mexico) ; Kirby, Cat. Diuni. Lrp. p. 539. n. 148 (1871) (Mexico ; cit.
Boisd. excl.) ; Obcrtb., Et. (J'Ent. iv. p. 71. n. 212 (1880) ; Staud., Exot. Tugf. p. Ifi (1884) ;
Godm. & Salv.. Biol. Centr. Amer.. Lep. Rhop. u. p. 225. n. 5fi. t. 69. fi','. 5. G. '? (1890) (Mexico
to Costa Rica) ; Winkle, Caiiad. Ent. xxv. p. 212 (1893) (only in Mexico).
(J. Pap'diii piilliis Doubleday, List Lep. Ins. Brit. Miis. i. p. IT (1845) (num. mid. ; Oajaca) ; id.,
Westw. & Hew , Gen. Diiirn. Lep. i. p. 17. n. 1G8 (1846) (uom. nwl. ; Mexico) ; Hewits., Trims.
Ent. Soc. Land. (2). i. p. 97 (IH'ol) (parti in) ; Gray, Cat. Lep. Ins. Brit. .l/«s-. i. Pap. p. 39. n. 193.
t. 6. fig. 1. (J (1852) (subsynon.)
Papilio lycopjhron, Butler & Druce, Proc. Zool. Soc. Loml. p. 365. n. 379 (1874) (Costa Rica).
<?. Forewiug with a row of submarginal spots on upperside, marginal spots
also distinct, band more cut up than in the South American forms, the veins
S(.;^, R', R-, R^ being more broadly black, the central patches of the baud more
or less rounded distally. Tail variable, ai)parently more obtuse in Easteru
Me.xico than in other localities, and in the same district the black distal area of
the hindwiug more extended both above and below.
¥ . Forewing more uniformly brown-black than in South American specimens ;
some butTdots distally of ape.x of cell ; submarginal spots SC — SC!''^, or at least the
first, absent. Hiudwing : tail short, sometimes only a little more projecting than
the other teeth ; three rows of spots, two inner rows often more or less confluent,
spots of proximal row either all red or at least the last spot, second row more
distinct than in South American females, and third row nearer the margin.
On underside a buff' band on forewing from costal margin to M' or SM",
consisting of rather well-defined spots.
Genitalia: (?. Rasplike ridge of harjie at right angles to the dorsal edge of
the harpe.
( 575 )
Hab. Mexico to f'osta Rica.
In the Tring Museum 18 cJc? from : Mexico ; Sau Pedro Sula, Honduras.
A series of 8 ? ? , besides males, from San Pedro Sula in coll. Oberthiir.
h. P. lycophron hippomndoii Feld. (1859).
cJ. rapiVio hippnmedon Felder, Wie7i. Ent. Man. iii. p. 393. n. 34 (1859) (hab. ?) ; id., Verli. Zool.
Bnl. Ges. Wicn xiv. p. 311. n. 309 (1864) (Venezuela).
(J. PapUio prjlycaon var., or nov. spec, Vollenhoven, Tijflschi: Ent. iii. p. 80. n. 14'2a (18C0)
(Bogota).
cJ. Papilla thenpih-on Felder, Verh. Zool. Bot, Ges. Wien xiv. p. 311. n. 308 (1864) (wom. imiescr. ;
Bogota) ; id., Brise Norara, Lep. p. 76. n. 59 (1865) ; Godm. & Salv., Tram. Ent. Sar. Loud.
p. 126. n. 237 (1880) (Sta. Marta) ; Hahnel, Iris iii. p. 201 (1890) (Valera, Venez.).
(J. Papilio lycophron var b. P. theopliron Kirby, Cat. Diiint. Lep. p. 540. sub n. 150 (W7i) (N.jw
Granada).
cJ. Papilio lyi'opliron var. c. P. hijipomedoii, id., I.e. (Venezuela).
(J. Papilio lycophron, Hahnel, I.e. p. 201 (1890) (Valera) ; Maass. & Weym., in Stiibel, Iln.-!/in
S. Aiiier., Lep. p. 11. n. 40 (1890) (Colombia).
c?. A small form, differing from the Brazilian subspecies especially in the
hind wing being more rounded, in the snbmarginal spots of the same being smaller
and standing much nearer the edge of the wing, and in the tail being short and
slender.
Genitalia : c?. Kasplike ridge of harpe extending in the same direction as the
upper edge of the harpe, being longitudinal, not transverse in direction, and being
moreover somewhat angulate.
? not known.
Hal). Colombia and Northern Venezuela.
In the Tring Museum S 6 S from ; " N. Granada " ; Venezuela.
c. P. hjcophron phania.^ subspec. nov.
cJ. Papilio hjcophron, Bates, Trans. Ent. Soe. Land. (2). T. p. 347 (1861) (Cametit, To^anlins) ; id.,
Joiirn. Entom. i. p. 228. n. 26 (1862) ; Dognin, Lep. Loja p. 15 (1867) ; id., I.e. p. 37 (1891) ;
Weeks, Ilhist. Diurn. Lep. p. 20 (1905) (Ghulumani).
Papilio theophron, Hahnel, Iris iii. p. 283 (189(1) (Pebas) ; Michael, ihld. vii. p. 213 (1894) (Sao Paulo
de Olivenja).
<?. Larger than P. I. hippomedon. Forewing : band more broken anteriorly
than in P. I. lycophron, and the cell-spot on the whole smaller ; spot SC — SC
absent from nearly all specimens. Hiudwing more strongly dentate than in
P. I. hippomedon, rather shorter posteriorly than in P. I. li/cophron ; submarginal
spots almost in the same jiosition as in I. hippomedon, being smaller and more distal
than in P. I. hjcophron.
Underside, forewing : submarginal spots linear, smaller than in /. lycophron,
costal margin at apex and veins SC to R^ more extended black-brown.
Hindwing : black discal area bearing the rufous red and the blue spots on the
whole larger than in hippomedon and hjcophron ; yellow submarginal spots all
separated, the last three or four smaller than in the other South American forms;
black marginal border a little wider than in hippomedon, but narrower than in the
specimens of lycophron, in which the submarginal spots are not strongly enlarged.
?. Only one specimen known to us, resembling the dark form of ? P. I.
lycophron.
Upperside : forewing llack-brown from base to apex of cell, then paler, some
Hnall ill-defined yellow spots around ajiex of cell; vestiges of yellow submarginal
( 576 )
Bpots in posterior half. Hindwing strongly dentate, tail slender, non-spatnlate,
the specimen resembling in this respect P. (uulrogeus ; submarginal spots much
closer to margin than in I. lijcophron, bnff, posterior ones slightly greenish, all
shaded with black.
Underside. — Fore wing : no distinct cell-patch ; a row of small creamy bnff
spots around apex of cell, the row continued on disc by a triangular spot R' — M',
this being the largest of all, being about as long as broad, upon which follows a
small indistinct double-spot M' — M- ; a row of submarginal spots from SC" to SM",
proximally of which there is a thin interrupted line disappearing anteriorly ;
between discal spots and apex of wing there are hardly any buff' scales.
Hindwing : submarginal spots less strongly curved than in /. li/ccphron, mncli nearer
the margin, the black marginal border being between the veins half the width
of the spots or less ; interspace between submarginal and rufous red discal spots
about three times the width of the submarginal spots, first spot excepted ; white
fringe nearly continuous, being narrowly interrupted at the veins.
Genitalia as in P. I. hippomedon.
Ilab. Eastern Ecuador (type) to Bolivia and Matto Grosso, eastwards to Pani
and the Orinoco.
In the Tring Museum 48 c?<?, 1 ?, from : Suapnre, Canra E., Orinoco, March
1899 (S.M. Klages) ; La Union, Caura E., August Kml (Klages); Itaitnba; Zamora
(0. T. Baron); R. Cachyaco, affluent of E. Huallaga (Stuart); E. Chuchuras,
affluent of E. Palcazn, 320 m. (W. Hoffmanns) ; Per6n(5, 3000 ft., October— November
1903 (Watkins) ; Mapiri ; Eeyes, August 1895 (Stuart) ; Villa Maria to Diamantino,
Matto Grosso, January 1897 (Andeer).
d. P. hjcophron bjcophron Hiibn. (1818 — ?)
(J. neraclides li/rophron Hiibner, Samml. Exot. Sehmelt. ii. t. 100 (1818-?).
J. Papilh: aMyalm Goi&ri, Enc. MM. ix. p. G2. n. 102 (1819) (Brazil); ViouU., Lkl Lep. Ins.
Brit. Has. i. p. 17 (1845) (= thersitesi ; Brazil); Menfitr., Enniii. Corp. Aiiim. Mus. Pelrop., Lip.
i. p. 4. n. 62 (1857) (Brazil),
cj. PapWo mentor Dalman, Amd. Enl. p. 37. n. 2 (182,B) (Brazil ?).
(J. rapitin li/rophron, Boisduval, Spec. Gen. Lep. i. p. 362. n. VJi (1836) (Brazil ; " J resembles [J "
false ; var. A. alia species = himeros) ; Doubl., Westw. & Hew., Gen. D'mrn. Lep. i. p. 17.
n. 167 (1846).
(J $. Ptipilio hjcophron Hewitson, Trans. Enl. Soc. Lonil. (2). i. p. 1)7 (1851) (^J of "perilhous") ;
Dotibl , Westw. & Hew., l.c. ii. p. 529 (1852) ; Gray, Cat. Lep. Lis. Brit. .!/«.«. i. Pap. p. 38.
n. 192 (1852) (Brazil) ; id., List Lep. Lis. Brit. Mns. i. Pap. p. 53. n. 201 (1856) (Brazil) ;
Lucas, in Sagra, Bist. Cuba vii. p. 206 (1857) (" Cuba" error loci) ; Feldor, Vcrh. Zonl. But.
Ges. Wien xiv. p. 311. n. 306 (1864) (Brazil ; Uruguay. — " Amazonia inf." alia subsp.) ; Kirby,
Cat. D'nirn. Lep. p. 540. n. 150 (1871) (var. b and c excl.) ; Burm., Descr. Rep. Argent, v. Lep.
p. 60. n. 3 (1878) (partim ; Corricntes ; Buenos Aires) ; id., l.c. Atlas p. 5. n. 5 (1879) ; Gosse,
Entoni. xiii. p. 194 (1880) (Corrientes) ; Oberth., Et. iV Ent. iv. p. 70. n. 211 (1880) (Brazil) ;
Honr., Berl. Ent. Zeit. xxxii. p. 5(J0. fig. B (1888) (gynandromorphous specimen ; Sao Paulo) ;
Weym., Stett. Ent. Zeit. Iv. p. 315. n. 12 (1895) (Rio Grande do Sul) ; Blinningh., Verh. Ver.
Nat. Vnterh, IJamliurg ix. p. 20 (1896) (Sa. Theresa) ; Lathy, Trans. Enl. Sue. Land. p. 69.
n. 38 ( 1904) (a ^ without submarg. spots on upperside of hindwiug).
?. Paj/ilio pirithous Boisduval^ l.c. p. 358. n. 201 (1836) (Uruguay; " Cuba '' false ; — "Papillon
pirithous Roger" alia spec. = caigiianahus) ; Doubl., Westw. it Hew., l.c. i. p. 17. n. 175(1846)
(Uruguay ; " Cuba ? " error loci) ; Meni'tr., I.e. p. 4. n. 64 (1857 (" Cuba" false).
?. I'ujiilin nebaliis Boisduval, l.c. p. 360. n. 204 (1836) (hab. ?) ; Doubl., Westw. & Hew., l.e. i.
p. 17. n. 177 (1846) (S. America); Hewits., Trans. Ent. Soc. Land. (2). i. p. 97 (1851)
(= ? of pallas ?) ; Kirby, l.c. p. ,539. n. 148 (1871) (partim).
Pnpilio drepanon Gray, Li.it Lep. Lis. Brit. iTus. i. Pap. p. 53. n. 202 (1856) (jiow. nud. ; Rio Grande
do Sul— doubtless a $ oi lycophrmi).
( 577 )
Fapilio Vicophron (!), Hewitson, Jx. p. 97 (1851) (Brazil) ; Mabilde, Guia Pract. Bnrlnl. Rio Granile
do Snip. 40 (1891'.)
Papilio perithotii^ (!), Hewitson, l,€. (sub synon.).
(J ? . Papilin jii/rithoiis (!), Lucas, in Sagra, Hist. Cuba vii. p. 267 (1857) (•' Cuba " false) ; Felder,
Verb. Zool. Bot. Ges. Wien xiv. p. 311. n. 307 (1864) (" Cuba" false).
$ . Papilio lyrophron var. a. P. pii-ithoiis, Kirby, Cal. Diurii. Lep. p. 540. sub n 150 (1871) ("Cuba "
error loci).
(J. Calaiiles hjtopliroii, Kirby, in Hiibn., Samml. Exot. Sckmelt. ed. ii. p. 97. t. 313. fig. 1. 2
(190-?).
(?. Upperside, forewing : siibmarginal spots often present ; snbapical spots
SC^ — SC^ seldom absent; veins SC to R- narrowly black. flindwing: sub-
marginal spots variable in size, bat their centres always farther away from margin
than in the other South American subspecies.
Underside, forewing : snbmarginal spots usually large, often joined to the
marginal spots. Hindwing : extent of black discal area very variable, often
hardly any black scaling left ; submargiual spots large, contiguous, sometimes
joined to the marginal spots, but in most specimens the black marginal border
continuous and broader than in 7'. I. Idppomedon and phanias \ occasionally some
yellow scaling in centre of tail.
¥ . Dichromatic.
«'. ?-!'. oebalus Boisd., ^.<^. Hindwing from near base to black discal band
pale creamy buff, this colour extending on to forewing as far as cell, there being
.1 spot of the same colour in the apex of the cell ; in the palest specimens the
biiffish band reaching to costal margin.
h'. ?-f. pirithous id.. I.e. Both wings almost uniformly brown above, with
a row of pale buff snbmarginal spots, which are often much shaded with brown.
liab. Brazil ; Paraguay ; Argentina.
A common insect.
In the Tring Museum: 100 SS, 34 ? ? and some larvae and pupae from:
Tucuman (J. Steinbach ; P. Girard) ; Tapia, Tucuman (Baer) : Sapucay, Paraguay
(W. Foster) ; Patino Cu6, February (Montforts) ; Yhu, Paraguay, September —
December 1896 (Andeer) ; Corricntes ; Blnmenau ; S. Catharina ; Castro, Parana
(E. D. Jones) ; S. Paulo ; Rio de Janeiro ; Espiritu Santo; Minas Geraes ; Bahia.
T8. Papilio androgeus Cram. (1775).
Merian, Tin. Surinam, t. 31 (1705) (cj, ?, larva, pupa).
cJ. Papilio Eques Trujauus uiidrngeux Cramer, Pap. E.rot. i. p. 24. t. ICi. fig. C. D (1775) (Surinam).
Papilio {orestes), Meerburgb, Afb. Zeklz. Gew. t. 20. 30. ^ (1775).
(J. Fapilio Eques Achirus pohjcaoii Cramer, I.e. iii. p. 17. t. 203. fig. A. B (1779) (Surinam).
$ . Papilio Eques Achimis piranthus Cramer, l.r. iii. p. 18. t. 204. fig. A. B (177'j) (Surinam).
Papilio Eques Trojanus ainosis, Stoll, in Cramer, l.i: Siippl. p. 1. t. 1. fig. 1 .\. v.. larva, pupa (1787)
(this species ?).
{J ¥ . Papilio pohjcaoii^ Boisduval, Spee. Gen. Lip. i. p. 361. n. 205 (1836) ( = ajidro/jeus = j'l/ranthus !
= laodocu.s) ; Ilaase, Untevsuch. Miiniery i. p. 97 (1893).
Papilio piranthous (!), Weidemeyer, Proc. Ent. Soc. Philad. ii. p. 148 (1863) (sub synon.).
g ? . Papilio androgeus, Felder, Verh. Zool. Bol. Ges. Wien xiv. p. 311. D. 312 (1864) (West Indies ;
Central and South America) ; Hopfl., Stett. Ent. Zeit. xl. p. 52. n. 21 (1879).
cJ ? . Papdio androgens (!), Kirby, Cat. Diurn. Lep. p. 539. n. 147"" (1871) (South and Central
America); Staud., Exot. Tagf. p. 15. t. 10. (J J (1884)(Chiriqui ; Surinam ; Amazons ; Brazil).
(J ? . Sexes dissimilar in colour, the females being dichromatic in certain
districts. Snbmarginal spots absent from upperside of forewing or vestigial,
thill and linear on underside. Hindwing : tail narrow, non-spatulate, teeth R-
( 578 )
and M' nsnally somewliat produced, especiall)- in female ; submarginal spots thin
crescents on npperside ; on nnderside, pvoximally of the submarginal spots, a row
of bine Innnles, and proximally of these and parallel with them a row of rufous red
crescents.
Genitalia: 6. Tenth tergite spatulate. Clasper and harpe broad; the latter
about two-thirds the length of the former, rounded at apex, bearing ventrally
a long pointed conical process directed distad and reaching a little beyond the
apex ; at dorsal margin of harpe an oblique ridge bearing numerous curved,
conical, pointed teeth, forming a kind of rasp. ?. Edge of vaginal orifice raised
right and left into a rounded lobe, and proximally into a kind of smooth ridge
which extends into the orifice, being narrowed posteriorly ; behind the orifice a
membranaceous lobe or tubercle densely clothed with minute hairs ; laterally of
the orifice a broad brown flap bearing several acute teeth ; anal segment witli a
few short stout bristles on innerside.
Larva and pupa first figured by Merian, and later by Burmeister, Sepp, and
others (see literature below). Both Sepp and Burmeister considered the cateriiillar
represented by Stoll, in Cramer, Pap. Exut. SuppL, t. 1. fig. 1, to be that of the
present species, not of P. ainosis = hi/ppason as stated by Stoll. We are not sure
that those authors were right. Stoll's figure bears distinct tubercles, such as are
found in P. epinetes and allies, to which group P. hi/ppason belongs, while the adult
caterpillar of P. androgcus has no distinct tubercles.
Ilab. West Indies, Mexico southward to Paraguay and Parana, Brazil.
Three subspecies, which differ constantly only in the females.
a. P. androgeus epidaurus Godm. & Salv. (1890).
cj ?. PapiVm polycacn var. b, Gray, Cat. Lep. Jns. Brit. Mus. i. Pap. p. 36. sub n. 173 (185'2)
(Guatemala) ; id., List Lep. his. Brit. Mus. i. I'ap. p. 48. sub n. 181 (1856) {parlim ; S. Domingo ;
Guatemala).
tJ $ . Pupilin aiiiho;ieus, Felder, I.e. ; Herr.-Sch., Corresp. Bl. Zoul. iliii. Vei: Hetjemh. p. 172. r. 3
(18G4) (Cuba) : HopfE., I.e. ; Gundl., Papilio i. p. 113 (1881) (Cuba) ; Miischl., Abh. Seiikenb.
Nat. Ges. xvi. p. 91 . n. 1 (18111) (Porto Rico, ? jiiraiMus).
(J ?. Papilio poUjcaoii, Poey, Mem. R. Soc. Earn. Habana p. 234G (1846) (Cuba) ; Lucas, in Sasra,
Hist. Cuba vii. p. 204 (1857) ; Bates, Proc. Zool. Soc. Loud. p. 242. d. 3 (1863) (Panama) ;
Reakirt, Proc. Ent. Snc. Fhilad. ii. p. 137. n. 3 (1863) (syn. excl. ; Honduras) ; Butl. & Druce,
Proc. Zu(A. Sor. Land. p. 365. D. 380 (1874) (Costa Rica) ; Dewitz, StM. Ent. Zeit. xxxviii.
p. 233. n. 1 (1877) (Porto Rico, one $ -form) ; Gundl., Coiilr. Ent. Cuba. p. 134 (1881) ; id.,
An. Hist. Nat. Madrid xx. p. 113. n. 1 (1891) (Porto Rico).
(J ? . Paiiilio polijcaon var, Weidemeyer, Proc. Ent. Soc. Philad. ii. p. 147 (18l")3) (syn. excl. ;
West Indies ; Central America).
Papilio pir.udhus, Butler, Pror. Zool. Soc. Loud. p. 714 (1901) (Sta. Lucia).
Pajjilio androgeos (!), Staud., Eirot. Tarjf. p. 15 (1884) (jmrtiin ; Chiriqui).
g ?. Papilio ppidaiirns Godman * Salvin, liinl. Centr. Amer., Lep. lihnp. ii. p. 2-'4. n. 55. t. 69.
fig. 1.2. ? , 3. genit. (1890) (Mexico to Panama ; S. Domingo).
S. Yellow area of both wings on the whole more extended than in the Soutli
American forms; tlie small patch in front of the subcostal fork of the forewing
usually larger.
?. Only one form known. Forewing with small yellow patches or vestiges
of them around upper angle of cell, the patches larger below than above, forming
a curved band ; no patch in coll or on disc, but there are sometimes some yellow
scales in the apex of cell on underside. Greenish blue scaling on disc of hind-
wing denser than in the other forms, the blue area being rather sharply defined
proximally, entering cell.
( 579 )
Hab. Mexico to Panama ; Cuba ; Haiti ; Santa Lncia.
The only S. Lncia specimen which we have seen {S in Brit. Mns.) has no
black dorsal line on abdomen ; the submarginal crescents of the hindwing are
rather large.
In the Triug Museum 2U c?c?, 6 ? ? , from : Cuba ; Haiti ; Songolica, Mexico,
June 1899 (W. Schaus); Orizaba; Guatemala (Salvin); San Pedro Sula, Honduras ;
Costa Rica; Chiriqui ; Bogava, Chiri(iui, SOO ft. (Watson); Brava I., ("ebaco I.,
and Parida I., Jaouary 11)02 (Batty).
b. P. (tndrogeus androgens Cram. (1775).
Merian, Ins. Sarhiam. t. 31 (170.5) ((J ?, larva, pupa) ; id., J.r. t. G7. ? (1719) ; Kleemann, Beylr.
Nal. Ins. Gesch. i. p. C3. t. 8. fig. 1. 2. J,t. 9. fig. 1. 2. ? (175G) ; Seba, Thesam: iv. p. 40. t. 38.
fig. 13. U. ? , p. 47. t. 39. fig. 2. 3. S (1764).
Papilio Eques Trojanus polydamas Linn^, Sysl. Nat. ed. x. p. 460. n. 11 (1758) {parlhn\ citat.
Merian t. 31) ; Fabr., .S'.yrf. Eut. p. 447. n. 22 (1775) {partim) ; id., Ent. Sijst. iii. 1. p. 14.
D. 42 (1793) (yjttrtiw)-
J . Papilio Eques Achivus glaums, Fabricius {uon Linne, 1758, err. det.), Sjst. Eiil. p. 445. n. 14
(1775) i/xirtim) ; Goeze, Naturf. ix. p. 70 (1776).
?. Papilio Eques Trojanus androgens Cramer, I.e.; Fabr., Gen. Ins. p. 251. n. 22-3 (1776); Goeze,
Ent. Beylr. iii. 1. p. 43. d. 15 (1779) ; Fabr., S2>ec. Ins. ii. p. 8. n. 30 (1781) ; Jabl. & Herbst,
Natnr.^. Schmell. ii. p. 105. n. 29. t. 11. fig. 2. 3, t. 12. fig. 1 (1784) ("Merian's t. 31. erroneously
quoted by Liune and Fabricius under polydamas ") ; Fabr., Mant. Ins. ii. p. 4. n. 32 (1787) ;
Gmelin, Sysl. Nat. i. 5. p. 2231. n. 290 (1790) ; Jung, Alphih. Ven. Sehmett. p. 35 (1791) ;
Fabr., Enl. Sysl. iii. 1. p. 15. n.43 (1793) ; Esper, Ausl. Sehmett. p. 138. n. 03. t. 36. fig. 1 (1798).
(J. Papilio Eques Achious polymon Cramer, i.e. iii. p. 17. t. 203. fig. A, B (1779) (Surinam) ; Fabr.,
Spec. Ins. ii. p. 19. n. 78 (1781) ; id., Mant. Ins. ii. p. 10 n. 89 (1787) ; Jabl. & Herb-st, I.e. iii.
p. 133. n. 92. t. 41. fig. 1. 2 (1788) (larva excl.) ; Gmelin, I.e. p.. 2236. n. 307 (1790) ; Fabr.,
Enl. Sysl. iii. 1. p. 33. n. 96 (1793) ; Esper, Ausl. Schmell. p. 200. n. 91. t. 49. fig. 2 (1798).
?. Papilio Eques Aehivus p/ranlJius Cramer, I.e. iii. p. 18. t. 204. fig. A. B (1779) (Surinam).
$ . Papilio Eques Acliirus peranthns (!), Jablonsky ^^- Herbsfc, Nat'trs. SrluneU. ii. p. 111. n. 30. t. 12.
fig. 2 (1784) ; Esper, Ausl. Schmell p. 81. n. 30. t. 20. fig. 1. 2 (1788) (Surinam ; Cayenne).
^.Papilio Eques Trojanus androijeus ^) Papilio acanthus Gme\ia, Sysl. Nat. i. 5. p. 2231. sub
n. 290 (1790) {laps. enl. ; piranihus'i).
(J. Calaides polyeaon, Hilbner, Vers. bek. Schmell. p. 80. n. 891 (1818 ?).
? . Cedaides androgens, id.. I.e. n. 892 (1818 ?).
? . Calaides piranlhus, id.. I.e. n. 893 (1818 ?) ; id., Samml. E.eol. Sehmett. ii. t. 110 (1822 ?) ; Kirby,
ihlel., ed. ii. p. 98. t. .323. fig. 3. 4 (190—?) (liter, and habitat ^Mj-i/m).
(J. Papilio polyeaon, Godart, Enc. Meth. ix. p. 41. n. 48 (1819) (partim ; Guyane).
2 . Papilio androgens, id., I.e. ix. p. 41. n. 49 (1819) {partim ; Guyane ; = laodocus = pyranlhus !
= 2^^f'^^*fhus ~ glaucus).
(J$. Papilio polyeaon, Boisduval, Spec. Gen. Lep. i. p. 301. n. 205 (1830) {partim; Guyane;
= androgeus = pyranlhus ! = laodocus) ; Lucas, in Gue'r., Diet. Pitt. Hist. Nat. vii. p. 5 (1838)
( ? = androgens) ; Doubl., Westw. & Hew., Gen. Dinrn. Lep. i. p. 16. n. 152 (1840) {p.irlim ;
Guiana) ; Sepp, Vlind. Surinam iii. p. 147. 148 (1853) (larva, pupa, (J ? ) ; Wall., Trans. Enl.
Soe. Land. (2). ii. p. 255 (1854) (Amazons) ; Bates, ibid. (2). v. p. 346 (1801) (common in
open places throughout the Amazon region) ; id., Journ. Eiitoni. i. p. 228. n. 25 (1862) (common
throughout the Amazons) ; id., Natural. Riv. Aniaz. p. .'/2 (1864) (Pard, in street) ; Oberth.,
Et. d'Ent. iv. p. 72. n. 216 (1880) (Carare, Colombia ; Cayenne ; Sautarem) ; Michael, Iris vii.
p. 213 (1894) (Sao Paulo de Oliven^a).
tT ¥ , Papilio androgeus, Felder, I.e. ; Godm. & Salv., Trans. Ent. Soc. Land. p. 126. n. 238 (1880)
(Sta. Marta).
(J?. Papilio androejcos (!), Kirby, Cat. Diurn. Lep. p. 539. n. 147''" (1871) {partim); Druce,
Proe. Zool. Soe. Loud. p. 246. n. 14 (1876) (Peru : Huasampilla, 10,000 ft.) ; Stand., Exol.
Tagf. p. 15. t. 10. ^ ? (1884) {partim ; Amazon.s) ; Hahnel, Iris iii. p. 194 (1890) (Merida) ;
id., I.e. p. 201 (1.890) (A'alera, Venez.) ; Maass. & Weym., in Stiibel, Reiscn S. Amer., Lep. p. IS.
n. 30 (1890) (Magdalena valley, 800—1500 m.) ; Kaye, Trans. Ent. Soc. Loud. p. 207. n. 197
(1904) (Trinidad).
J'apilio androgeas (!), MOschler, \'crh. Zool. Bol. Gcs. Ween xxxii. p. 3u4 (1883) (Surinam).
( 580 )
c?. Usually deeper yellow than the Brazilian form ; the spots in front of cell of
forewing, vppeiside, rather larger in the majority of specimens, and the snbmarginal
bars of the nmlerside mostly thinner : black dorsal line of abdomen often vestigial,
sometimes absent.
? . Dichromatic.
a. ?-f. androgens Cram.. I.e. Forewing: yellow patches R= — M' of about
equal length, occasionally a sjiot before 11', patch ]{■'— M' often shaded with black ;
greenish or greyish bine discal area of hindwing entering cell.
b'. ? -f. pirantkus id., I.e. Forewing without yellow patches on upperside,
sometimes vestiges of the patches on underside, rarely also on upper ; greenish or
greyish blue discal scaling of hindwing rather diffuse, entering cell.
I/al/. Colombia to Trinidad, tlie (Juianas, Amazons, southward to Bolivia and
western Matto Grosso.
In the Tring Museum 58 c?c?, 25 ? ?, from : R. Dagua, W. Colombia (Rosen-
berg); Pereira, Cauca;" Bogota"; Marawal, Trinidad, August 1891 ; Lower Orinoco,
November 1897 (Cherrie) ; Temblador, Blaripa, La Vuelta and La Union, Caura R.,
Orinoco, May, September and October (S. M. Klages) ; Esseqnibo R. ; Surinam;
Obidos (Mathan) ; R. Cachyaco, affluent of R. Huallaga (Stuart) ; Paramba, N.W.
Ecuador, 3500 ft. (Flemming & Mikctta) ; Cachabi, N.W. Ecuador (Roseid)erg) ;
H. Chuchuras, affluent of R. Palcazu, 3:3(» m. (W. Hoffmanns); Perc^ni^ R.,
JIarch lOnu (Simons) ; Pozuzo (W. Hoffmanns) ; t^hanchamayo (Schunke) ; Cnzco,
February 1901 (Garlepp) ; Caradoc, Marcapata R., February 1902, 4000 ft.
(Ockenden) ; Mapiri R. ; Guanay, Mapiri, 1500 ft., August 1805 (Stuart) ; Ynngas
de la Paz, 1000 ra., September 1800 ((iarlejq)) ; Prov. Sara, S. Cruz de la Sierra
(J. Steinbach) ; Villa Maria to Diamantiuo, Malto Grosso, January 1897 (Audeer).
c. F. atidrogeus laodocm Fabr. (1703).
J. Papilin Eqiies Trojnmm midrngeiis Stoll (iioii Cramer, 1779, err. det.), in Cram., Ptip. Exol.
iv, p. 317. t. 350. fig. A. B (1781) (Brazil) ; Esper, .liisl. SrJimell. p. 138. n. 63.' t 3t! fio- 2
(1798) (jMi-tim).
? . Pajiilio Eqiies Trojaiius liiodocus Fabricius, EiU. Synl. iii. 1. p. 8. n. 23 (1793) (jtartim).
(J. rapilio jmlyciioii, Godart, Enc. Jleth. ix. p. 41. n. 48 (1819) (piirt/m ; Brazil).
$ . Pupilio audrogeiiK, id., l.r. p. 41. n. 49 (1819) {partim ; Brazil).
5 . Pajiilio laodocus, Donovan, Nat. Ripos., Ent. ii. t. 130 (1823) (Brazil).
J ? . Pupilio jwhjcuoii, Me'n^tr., Mini. Sue. Imp. Moncou vii. p. 188. n. 4 (1829) (Brazil, larva);
Boisduval, I.e. ; Doubl., List Lep. Im. Brit. .Vus. i. p. 17 (1845) (Brazil); id., Westw. & Hew.,
Geii. Diurii. Lep. i. p. 16. n. 152 (184G) {parlim ; Brazil) ; Gray, Cat. Lrp. Ins. Brit. Mas. i.
Pap. p. 35. n. 173 (1852) (Brazil) ; Lucas, in Chenu, Ene. Hist. Aat., Pap. p. 38. t. 6. fig. 2. ^
(1851-3) (Brazil); Gray, Li.ft Lep. Ins. Brit. Muk i. Pap. p. 48. u. 181 (1856) (/<n;7i;« ; BraziP ;
Lucas, Bull. Soe. Ent. Prance p. 25 (1857) ( ?, aberration, Rio de Jan.) ; Burm., Drmr. Rip.
Argent, v. Lep., Atlas p. 4. n. 4. t. 2. fig. 2. larva (1879) ("the same as StoU's t. 1. fig. 1 ") ;
Gosse, Entom. xiii. p. 194 (1880) (Corrientes & Paraguay) ; Seitz, Stett. Ent. Zeit. 11. p. 97
(1890) (Corcovado) ; Buuningh,, Verb. Vcr. Nat. Vnterhalt. Uamhnnj ix. p. 26 (1896) (Rio de
Janeiro).
(J. Calaides poh/mon, Geyer, in Hiibner, Sunnul. E.riit. Sriinielt. iii. t. 26. fig. 1. 2 (1834) (Brazil).
cJ $ . Pupilio androgens, Felder, I.e. ; Butler, Cat. Diurn. Lip. deser. Fahrie. p. 247. n. 46 (1869)
(Brazil).
(J?. Papilio androgens (!), Kirby, Cat. Diurn. Lep. p. 539. n. 147'"" (1871) {parlini); Stand.,
Exot. Tagf. p. 15 (1884) (parlim ; Brazil) ; Maas. & Weym., in Stubel, lieiseu .S. Amer., Lep.
p. 91. n. 39 (1890) (S. CatharinaV); Peters, Illn.'<tr. Zeilsehr. Ent. ii. p. 51 (1H97) (Nova
Friburgo, larva on Citrus).
Calaides androgens, Kirliy, in Alien, Nat. Lib., Butt. ii. p. 283 (1897).
Calaides androgros (!), id., in Hiibn., Sarnml. Exot. Sehmelt. ed. ii. p. 98. t. 464. fig. I. 2 (190- ?)
(liter, et hab. partim).
( 581 )
<S. Yellow baiid pale ; the small spots standing in front of npper angle of
cell of forewing rednced, being often absent. Length of streak in snbcostaj fork
as variable as in the other forms ; apex of cell of hindwiug sometimes black. We
have a curious S (Sapncay, Paraguay) bearing patches of the colour of the ? ; the
left clasper of this specimen is somewhat reduced.
?. Monochromatic, the only form corresponding to P. a. androffeus ?-f.
androgens. Forewing : yellow patch R' — M' much larger than patch R- — R^, the
latter being rednced distally ; often a small patch M' — M^ present, standing behind
the distal portion of patch R^ — MK Hindwing : greenish blue scaling forming
a band of patches which stands always separate from cell.
Hub. Brazil ; Paraguay.
In the Tring Museum 48 cj(?, 14 ? ?, from: Sapncay, Paraguay, Se])tember
to February (W. Foster) ; Yhu, Paraguay, September to December IS'JO (Aiideer) ;
Castro, Parana (E. D. Jones) ; Bahuru, S. Panlo (Dr. Hempel) ; Rio de Janeiro ;
Petropolis (Foetterle) ; Nova Fribnrgo ; Tijuca.
VI. Glaucus Group.
Cell of the hindwing (except P. pilumnus) enlarged, strongly asymmetrical,
mnch broader between SC- and M- than in any other American species. Basal
cellule of hindwing long. Fifth black band of forewing connected at costal margin
with distal marginal border, forming a kind of C filong the costal edge ; a black
median band on hindwiug joining posteriorly the black abdominal border, the two
bands forming a large V- Abdomen beneath striped with black and yellow. P.
pilumnus stands somewhat isolated in this group. The other five species which
belong here (P. glaucus, rutulus, alexiares, eunjmedon. and daunus) are very closely
allied with one another. If these five insects inhabited separate districts there
would be good reasons for treating them as subspecies of one species. But as
P. daunus, ewymedon and rutulus occur together, aud are known to be independent
of each other, each breeding true, and P. daunus and alexiares are also found
together (in Eastern Mexico at least, alexiares having a restricted range), these
Papilios are certainly quite distinct. As further the morphological differences
between these species are not more trenchant than those between any of these
insects and P. glaucus, we must consequently treat P. glaucus also as a species
independent of the others. That these Papilios are develoiiments from the same
ancestral form there can be no doubt.
The sexual armature is of the same type in those five species — resembling to a
certain extent the genitalia of P. lycophron and aristodemus. The tenth tergite of
the (J is long, being slightly spatulate ; the sternite bears a small oblique double
ridge at each side. The harpe is broad, being produced ventrally at the apex into a
long conical pointed process ; the oblique dorso-apical edge of the harpe is more or
less dentate, being dorsally produced into a simple or a dentate hook which is curved
auad. ?. The edge of vaginal orifice is proximally raised into a long jirocess
which is more or less lanceolate ; at each side of tlie orifice there is a large
dentate flap and proximally of this flap two folds ; behind the orifice there is a
membranaceous tubercle or projection which is densely covered with minute hairs.
The larva bears an eye spot on each side of the tliird thoracic segment snb-
dorsally, and a bluek transverse dorsal line on the fourth, this line not being present
in /'. pilumnus.
( 582 )
Kej- to the species.
a. Cell of hindwing nearly symmetrical Species No. 84.
Cell of hindwiiig asymmetrical , b.
h. The snbapica! costal o-shaped spot of forewing below with
sharply defined buff or yellow centre, the edges re-
maining pure black ....... c.
The c^-mark without sharply defined bntf or yellow centre e.
c. Tooth M' of hindwing long ; fourth black band of forewing
narrower than the yellow apical cell-space (<?), or at
least not wider (?) . • • . . . . Species No. 81.
Tooth M' short d.
(I. Submarginal spots of underside of hindwing nearly all
stained with orange, at least in centre .... Species No. 79.
The last two spots orange, the other buff, or the first one
also orange and the middle ones with a trace of orange Species No. 83.
e. Ground-colour huffish white Species No. 82.
Ground-cciliiur yellow ....... Species No. 80.
79. Papilio glaucus L. (1758).
Mouffet, 7ns. Theatr. p. 98. fig. (1634) ; Petiver, .lA«s. p. 50. n. 50.') (1703) ; Rajas, 11,^1. In^. p. 111.
n. '1 (1710) ; Cutesby, Nat. IJht. Car., Flo., Bahama /s.ii. t. 83 (1743).
$ . Papilio Eques Trojaiius glaiicus Liniii', Si/sl. Xat. ed. x. p. 4C0. n. 9 (1758) (Amer. sept.) ;
Clerck, Icon. Ins. ii. t. 24. fig. 1. ? (1764).
Papilio Eques Arhivu-'i ajnx Linni?, Syst. Nat. ed. x. p. 462. n. 26 (1758) (jjnW/'m ; cit. 7i"J. ins. iii.
n. 2).
5 . Papilio Eqncs Achiinis antilocJius id., Stjat. Nat. ed. x. p. 463. n. 28 (1758).
(J. Papilio Eqnes Achivus turnus id., Mant. Plant, p. 536 (1771) (}>artim).
cJ ?. Papilio (/lav: us, Kirby, Cat. Diurn. Lep. p. 505. n. 316 (1870) (= anlilochus = turnus =
ahi(lama--<).
(J?. Papilio tnrnn.i, Edwards, Trans. Amer. Ent. Sm: vi. p. 11. n. 17 (1877) (Atliintic States to
Rocky Mts.; Brit. America to Mackenzie R. ; Alaska ; Canada ; Nova Scotia ; Newfoundland).
S. Upjjerside.—FoTewmg : yellow subbasal interspace strongly narrowing
costad, triangular ; subapical e->-mark centred with a yellow streak, the pro.xiiual
and posterior edges of the spot remaining pare black ; the yellow spot SC — SC*
situated immediately behind the n-mark smaller than spot SC^ — SC'^ ; subapical
submarginal yellow dot small, rounded. Hindwing: first submarginal spot
usually orange ; tail more or less strongly spatulate, asymmetrical.
Underside. — Forewing : e->-mark broadly centred with buff", the black border
.around the bull' spot SC^ — SC^ remaining pure black, the butf-scaling of the c-»-inark
continuous with the similarly coloured powdery line situated on the black postdiscal
band; the buff scaling absent from the c-»-mark in one of our British Columbian
specimens, the "-mark being much reduced in the individual. Hindwing: the
last two submarginal spots orange, some of the other submarginal spots being also
more or less washed with orange ; disc with some more or less large orange patches
H2_SM- situated prosimally of the black postdiscal line. Several melanistic males
have been found.
? . Dichromatic in the southern districts. One form similar to male, the black
bands more extended and the blue patches of the hindwing orange, more or less
rounded. The second form of female blackish brown, the black bands more or
less vestigial, being more distinct beneath than above, the submarginal spots
remaining yellow resj ectivel} orange. Intermediate specimens rare,
( 583 )
Genitalia : <S. Process of barpe situated proxiiuuUy of m ist distal point of
apex, the apical edge of the harpe being rotnndate-angulate ; oblique dovsal edge of
barpe bearing at least two strong hooks at the upper corner, and several prominent
teeth near these hooks, there being occasionally also some teeth at the ventral
edge ; individual variability considerable. ?. Process in front of vaginal orifice
lanceolate, pointed, rarely bidentate at tip; dentate lateral flaps standing farther
anad than the orifice.
For early stages see Edwards and others (see literature below).
One brood in the north, two or more in the south ; the spring specimens
(from hibernated pupae) of the southern districts are smaller than the summer
specimens.
Hdb. Alaska, British Columbia, eastwards to Newfoundland and southwards
to Florida and Texas, not in the Pacific (and Rocky Bits.) district of the United
States.
Two geographical forms, southern spring specimens somewhat resembling the
small northern form, but being easily distinguishable by the much narrower black
.abdominal border to the hindwing. The geographical boundary line between the
northern and sonthern forms is in the lake district, and it is probable that the two
forms completely intergrade in Southern Canada. Observations on this point are
wanting. As far as we could, we have kept separate the literature on the two forms.
It is to be hoped that in future local records of this species it will be expressly
stated by the authors which of the two forms occurs, this being especially desirable
in records from southern Canada and the northern districts of the United States.
a. P. glaucus glaucus L. (1758).
$. Papilio Eques Trojanus glaums Linne, I.e. (1758) ; Clerck, I.e. (1704) ; Linne, itus. Liid. Ulr.
p. 190. n. 9 (1764) ; Houtt., Naturl. Hist. i. 11. p. 104. n. 9 (1767) ; Fabr., Syst. Ent. p. 445.
n. 14 (1775) (cit. Kleem. excl. ; cit. Linn. "74G" falsa) ; Cramer, Pap. Exot. ii. p. 64. t. 139.
fig. A. B (1777) ; Goeze, Ent. Bei/tr. iii. p. 33. n. 9 (1779) ; Fabr., Sjiec. Ins. ii. p. 5. n. 18
(1781) ; Jabl. & Herbst, Natur.s. Schmett. ii. p. 229. n. 47. t. 17. fig. 1. 2 (1784) ; Esper, Aiisl
Schmctt. p. 27. n. 9. t. 5. fig. 1 (1785) ; Fabr., Maiit. Ins. ii. p. 3. d. 18 (1787) ; Gmelin, Si/st.
Nat. i. 5. p. 2229. n. 9 (1790) ; Fabr., Ent. Sy.-it.m. 1. p. 4. n. 11 (1793).
Papilio Eques Aehivus aja.v Linn6, I.e. (^partiin).
(J. Papilio Eques Aehii'us antilochus id., I.e.; id., Mui. Lnd. Ulr. p. 207. n. 26 (1764) (Amer. sept.);
Houtt., iVa(«i-;. Hist. i. 11. p. 209. n. 28 (1767) ; Fabr., Syst. Ent. p. 4.01. u. .37 (177.0) ; Goeze,
Ent. Beyti: iii. 1. p. 61. u. 35 (1779) ; Fabr., Spse. Ins. ii. p. 15. n. 57 (1781) ; id., Mant. Ins.
ii. p. 8. n. 63 (1787); Gmelin, Sy.it, Nat. i. 5. p. 2241. u. 35 (1790); Fabr., Ent.Syst. iii. 1. p. 24.
n. 70 (1793).
Papilio Eques glaucus, Lange, in Linne, Syst. Nat. p. 460. n. 9 (1760).
Papilio Eques antilochus, id.. I.e. p. 403. n. 28 (1760).
Papilio Eques Aehivus antkiluehus (!) Linne, Syst. Nat. ed. xii. p. 751. n. 35 (1767).
jj. Papilio Eques Aehivus tui-uus Linne', ic. (1771) ; Miiller, Naturs. Sujjjil. p. 284. n. 496 (1774) ;
Fabr., Syst. Ent. p. 452. n. 41 (1775) ; Goeze, Ent. Beytr. iii. 1. p. 71. n. 5 (1779) ; Fabr., Siiec.
Ins. ii. p. 16. n. 66 (1781) ; id., Mant. Ins. ii. p. 9. n. 76 (1787) ; Jabl. & Herbst, Naturs.
Schmett. iii. p. 136. n. 93, t. 41. fig. 3. 4 (1788) (= aleiflumas) ; Gmelin, Syst. Nat. i. 5. p. 2243.
n. 338 (1790) ; Jung, Alphah. Verz. Sehmelt. p. 253 (1792) ; Fabr., Ent. Syst. iii. 1. p. 29. n. 86
(1793) ; Esper, Aushind. Sehinetl. p. 195. n. 88. t. 48. fig. 1 (1797).
Papilio (^Troes) glaucus, Miiller, Naturs. v. 1. p. 568. n. 9 (1774).
Papilio lAehivus) antilochus, id.. I.e. p. 570. u. 35 (1774) (New York ; " Surinam " errors).
Papilio Eques Aehivus (dddamas Cramer, Pap. Exot. i. p. 62. t. 28. fig. A. B (1775) ("Jamaica"
error loci; New York, Carolina); Goeze, Ent. Beytr. iii. 1. p. 77. u. 27 (1779); Stoll, in
Cramer, I.e. iv., Ordre Syst. p. 3. n. 3 (1782) (= turnus = antilochus).
Papilio (rinthdoehus), Meerburgh, .4/6. ZeUh. Geiv. t. 40 (1775).
( 584 )
Pairlh i,l,iiH-u«, Palisot, Ins. A/,: Amer. p. 99. Up. i. 1. b. fig. a. b. ? (1805-21) ; Godart, Enc.
MM. ix. p. 60. n. % (1819) (N. Amer. ; "Jamaica" false) ; Boisd. & Lee, Hist. Gin. Lip.
Amir. Sept. p. 22. t. 8. 9. J , larva, pupa (18:-13) (Georgia ; Virginia) ; Boisd., Spec. Gin. Lip.
!. p. 335. n. 177 (18.36) (Georgia ; Virginia ; Carolina ; "Jamaica" false); Doubl., in Westw.,
Arc. Ent. i. p. 143 (1845) ; Lucas, in Clienu, Enc. Hist. Nat., Pap. 1. p. 38. t. 10. fig. 1. ?
(1851-53) ; Meni''tr., Eniiw. Corp. Anim. Mm. Petrnp., Lip. i. p. 3. n. .32 (1857) ; Gosse, Letters
from Alabama p. 122 (1859) ; Vollenh., Tijihchr. Ent. iii. p. 85. n. 128 (1800) ; Morris, I.e. n. 2
"(1860); Scudd., Canad. Ent. iv. p. 74 (1872) (Abbot's MSS. in Brit. Mus.) ; id., Rept. Geol.
N. Hampsh. i. t. A, fig. 16 (1874) ; Auriv., K. Sv. Vet. Ak. Handl. xix. .5. p. 14. n. 9 (1882)
(rccensio critica) ; Smyth, Ent. News vi. p. 244 (1895) (Montgomery Co., Virginia ; this the
only ? form of tnrnus I have seen here); Dyar, Bull. U.S. Nat. Mus. Iii. p. 2. n. 11 (1902)
{parthn: Atlantic States); Laur., Ent. News xvi. p. 312 (1905) (?, "large admixture of
yellow ").
Papilio turn,,.-, Palisot, I.e. p. 119. Lip. t. 2. b. fig. 1. J (1805-21) ; Godart, I.e. ix. p. 55. n. 87
(1819) ; Say, Amer. Ent. iii. p. 86. t. 40 (1828) ; Boisd. & Lee, I.e. p. 19. t. 6. fig. 1. t. 7.
fig. 1—3 (18:!3) ; Lucas, Lip. Exot. p. 35. t. 18. fig. 2 (1835) ; Boisd., fipec. Gin. Lip. i. p. 338.
n. 179 (1836); Harris, Enlom. i. p. 61 (1840) (larva solitary, covers leaf with coating of silk
and binds up the sides to form a kind of trough); Doubl., in Westw., Arc. Ent. i. p. 143 (1845);
id., List Lep. Ins. Brit. Mus. i. p. 16 (1845) ; Gray, Cat. Lcp. /«.?. Brit. Mus. i. Pap. p. 24. n. 112
(1852) ; Emm.. Ai/ric. N. York v. p. 201. t. 38. fig. 3 (1854) ; Gray, List Lcp. his. Brit. Mus. i.
Pap. p. 32. n. 119 (18.56) ; Fitch, Bcpt. fns. N. York iii. p. .341 (1856) ; Durban, Can. Nat. Geol.
ii. p. 223. fig. c. d. t. 3. fig. 1 (1857) (larva, pupa) ; Mdne'tr., Enum. Corp. Anim. Mus. Petrop.,
Lip. i. p. 2. n. 31 (18.57); Gosse, Letters from Alabama p. 202 (1859); Morris, Si/n. Lep.
N. Amer.p. 2. n. 1 (18,;0); Noneah..rijilsclir. Ent.nl p. 85. n. 129 (1860) (Indiana ; Tennessee);
Durban, /..-. v. p. 87 (1860) ; Newm.,' Pror. Ent. Soc. Philad. i. p. 26 (1861) (N. Jersey ; on
tulip- p<'plar) ; StaufF., ibid. i. p. 265 (1862) (larvae of rjaucus and turnus different !— They were
found on different food-plants) ; Rid., ibid. i. p. 260 (1862) ((,Uurus is ? of turmis, as found
by Dyson already in 1843); Walsh, ibid. p. 349 (1863) (in southern Illinois all ? ? black iu
summer 1861, on Atlantic coast all ? ? black perhaps up to 36° lat., in Mississippi v.alley up
to 38°, north of 41° on Atlantic coast, and 4.3° in Mississippi valley perhaps all yellow) ; H.arr.,
ed. Flint, Ins. hij. Veg. p. 268. fig. 97. ? , 98. 1. (1862) (life history) ; Weidem., Proc. Ent. Hoc.
Philad. ii. p. 148 (1863) ; Lintn., ibid. iii. p. 50 (1864) (Eastern N. York, larva, descr. of pupa;
$-f. S(Z(rac»s not occurring); Kirkp., ibid. iii. p. 329 (1864) (Cleveland, Ohio, common, ?-f.
glavcus does not occur) ; Fclder, Verb. Zool. Bot. Gcs. ]Vicn xiv. p. 314. n. 351 (1864) (partim;
"Jamaica" false) ; Tenney, .Uan. Znnl. fig. 281. 282 (1867) ; Reak,, Proc. Ent. Soc. Philad.
vi. p. 124 (1867) (Pike's Peak, Colo. ; also aberr.) ; Edw., Trans. Amer. Ent. Soc. ii. p. 207
(1868) ( ? one side yellow, the other black ; aberr. ? ? mottled with black) ; Butler, Cat.
Diurn. Lep. descr. Fabric, p. 248. n. 49 (1869) ; Riley, Amer. Entnm. i. p. 99 (1869) ; Harris,
ed. Flint, Ent. Corresp. p. 270 (1869) ; Parker, Amer. Entom. ii. p. 175 (1870) (Iowa) ; Morris,
Psyche i. p. 35 (1874) (White Mts.) ; Scudd., Canad. Ent. iv. p. 84 (1874) (Abbot's MSS.
in Brit. Mus.) ; Bean, Ent. Mo. Mag. x. p. 248 (1874) (Galena, III., scarce, sometimes not
uncommon in June) ; Pagenst., Verb. Nat. Med. Ver. Heidelb. (2). i. p. 101 (1874) ; Mead, in
Wheeler, Rept. E.rpl. Surr. v. Zool. 8. p. 741 (1875) (occas. in Ricky Mts.) ; Boll., Tafjcbt. Ver.
Nat. Hamburg. Beil. 49. p. 176 (1870) ; Perk., Rept. Vermont Board Agric. ii. p. 589 (1870) ;
Bruner, Canad. Ent. ix. p. 20 (1877) (Omaha, Nebraska, black ? more frequent than yellow ;
Niobrora R., both ? ?) ; Edw., Butt. N. Amer. ii. Pap. t. 3. 4. 5 (1877 and 1884) ; Pack.,
Half-hours p. 180 (1877) ; Dury, Cincinnati .Soc. Nat. Hlsl. i. p. 12 (1878) (Cine, abundant) ;
Gerh., Macro-Lep. N. Amer. p. 25. n. 448 (1878) ; Strecker, Butt. Moths N. Amer. p. 09. D. 10.
(1878) {partim) ; Oberth., Et. d' Ent. iv. p. 67. n. 188 (1880) (partim) ; Middl., Trans. Dept.
Agric. lllin. xviii. p. 74 (1880) ; Coquill., Ibid. p. 173 (1880) ; Skinn., Proc. Ac. N. Sci. Philad.
p. 239 (1882) (scent-glands of larva) ; Saund., Ins. Lij. Fruits p. 83. fig. 80. 81. 82 (1883) ;
Edw., Canad. Ent. xv. p. 109 (1883) (larvae of turnus and rutulus different) ; Gruber, Jena.
Zeitschr. Ges. Nat. xvii. p. 470. t. 7. fig. 9 (1884) ; id., Papilio iv. p. 86. t. 1. f. 7—11 (1884)
(transf.); Fern., Butt. Maine p. 24. fig. 1. 2 (1884) ; Edw., Canad. Ent. xvi. p. 115 (1884) ; id.,
l.c. xvii. p. 113 (1885) (larva refused willow) ; Hoy, Rept. Ent. Soc. Ontario xv. p. 12 (1885)
(Racine, Wise.) ; Merr., ibid. (1885) (Central Adirondacks, common, black J ? occurring) ;
Fern., Canad. Ent. xviii. p. 50 (1886) (al>errat., Maine) ; Edw., Ibid, xviii. p. 1.39 (1886)
(food-plants, also willow); Mayn., Butt. E. IK S. p. 51. n. 51. t. 6. fig. 70. 70a (1886) ;
Morton, Canad. Ent. xx. p. 228 (1888) aberrat. ; N. Windsor, N.Y.) ; Riloy, L(se,-t Life i.
p. 161 (1888) (parasite: Tragus e.resori u.i) : Skiun., Canad. Ent. xxi. p. 127 (I88'.l) (Phila-
delphia, black and yellow ? ? in about equil numbers; vars. of $ ) ; Fletcher, ;//"/. xxi.
( 585 )
p. 20). fig. 9, 11 (1880) (aberrat. ; life hist.) ; Edw., Bull. U.S. Nat. Mus. xxxv. p. 11 (1889)
(literal, refer, to raetam.) ; Pack., Fifth Rept. U.S. Ent. Conim. p. 217. 472. 480. 529. 531. 53i;.
555. 6G9 (1890) (food-plants) ; id., I.e. p. 480 (1890) (early stages) ; Mayn., Man. N. Amer.
Butt. p. 11. n. 15 (1891) {partiin ; fig. alia subsp.) ; Fletcher, Insert Life v. p. 126 (1892)
(parasite of egg: Trichngmmma); Staley, OdkuI. Ent. xxiv. p. 204 (1892) (Marshall, Missouri,
commou) ; Haase, Untersuch. Mimierij i. p. 90 (1893) ; Brodn., fn.feel Life vi. p. 40 (1893)
(larva on Camphora nfficinalis) ; Daggett, Ent. News iv. p. 15 (1893) (decoying) ; Skinn., ibid.
iv. p. 82 (1893) (N. Carolina) ; Meeske, ibid. iv. p. 117 (1893) (Long I., reared $ ylaucus) ;
Jones, ibid. iv. p. 190 (1893) (Richmond Co., N.C.) ; Winkle, Canad. Ent. xxv. p. 212 (1893)
(black (J ! !) ; Davis, Jouni. N. York Ent. Soe. i. p. 47 (1893) (Staten I., N.Y., May to Sept.) ;
Beutenm., Bull. Anicr. Mus. N. II. v. p. 244. t. 2. f. 2. ^ (1893) (N. York ; descr. of I., p., i.);
Davis, Ent. News v. p. 109 (1894) (Little Rock, Ark.) ; White, ibid. v. p. 175 (1894) (Brooklyn) ;
Weed, P.'ttjrhe vii. p. 130. n. Sfi (1894) (N.E. Miss.) ; Ehrm., Canad. Ent. xxvi. p. 292 (1894)
(aberrat. of J , right forewing yellow, left black) ; How., Ins. Life vii. p. 44. fig. 15. 10
(1894) (melan. ^ ; Kansas); yVeith, Ent. News vi. p. 158 (1895) (Vermont, larva 3 weeks
without food before dying) ; Osburn, ibid. vi. p. 282. n. 44 (1895) (Tennessee, common, iv.
to ix,, two broods ; ? glancns frequently in Aug. & Sept.) ; Long!., ibid. vi. p. 314 (1895)
(Chicago) ; Eimer, Artb. Verwandlseh. Schmett. ii. p. 79. t. 5. fig. 1. 2 (1895) ; Clevel., Ent.
News vii. p. 73 (1890) (Oneonta.N.Y.); Fi.ske, ibid. vii. p. 241 (1890) (Webster, N.H., common.
May, June); Soule, Psijehe vii. p. 398 (1896) (Brookline, Mass., ovipositing on higher branches
of young aah-trees) ; Truman, Ent. News viii. p. 29 (1897) (Volga, S. Dakota) ; Bubua, ibid.
viii. p. 98 (1897) (Cleveland, Ohio ; scarce this year); Eimer, Ortlwgen. p. 32. fig. 10. ? (1897);
Christ, .Willi. Srhweiz. Ent. Ges. ix. p. 270 (1897) ; Duzee, Bull. Buffalo Soc. N. Se. v. p. 107.
n. 5 (1897) (Buffalo, common); Rowley, Ent. News ix. p. 37 (1898) (Louisiana, Mo.); Beutenm.,
Bull. Amer. Mns. N. II. x. p. 310 (1898) (Highland Falls, N.Y.) ; Holland, Butt. Book p. 309.
n. 4. t. 2. fig. 15. 26. 28. larva, t. 6. fig. 1—4. pupa, t. 43. fig. 1. ^, 2. ? (1899) {partini) ;
Denton, Moths Butt. ii. p. 335. fig. ? (1898—1900) ; Beutenm., Bult. N. York Citg p. 3. n. 1.
fig. ? (1902); Macgill. & Hought., Ent. News xiv. p. 205 (1903) (Adirondack Mts.); Comst.,
ibid. xiv. p. 197 (1903) (Adirondack Mts., very common in June) ; Briml. & Sherra., ibid.
xiv. p. 230 (1903) (Raleigh, N.C, also glaurus, March 31) ; Heink, ibid. xiv. p. 335 (1903)
(Meramec Highlands, St. Louis Co., April 12) ; Pass., ibid. xvi. p. 328 (1905) (colour of larva
harmonising with that of leaf).
Jasoniades turnus, Hiibner, Verz. bek. Selinietl.p. 83. n. 843 (1818?).
Eujihoeades (jlauens^ id., I.e. p. 83. n. 846 (1818) ; Sprague, Pstjche ii. p. 2.07 (1879) (Wollaston,
Ma.ss., May 24) ; id., I.e. p. 259 (1879) (Mass., May 25 to July 15) ; Morse, Pstjche vii. p. 155
(1894) (Stamford, Conn., Aug. 22) ; Kirby, in Allen, Nat. Libr., Bull. ii. p. 284 (1897);
id., in Hubn., Samml. Exot. Schmett. ed. ii. p 99. t. 308. fig. 1. 2 (190—?).
Papilio antiloehus, Boisduval, Spec. Gen. Lep. i. p. 340. n. 180 (1836) (turnus with artificial tail) ;
Auriv., K. Sv. Vet. Ak. Handl. xix. 5. p. 28. n. 26 (1882) (recensio critica ; '' spec. fict.").
Papilio troilus, Jaeger, Life N. Amur. Ins. fig. 53. ? (1864).
Papilio tttrnus var. glaucus, Edwards, Canad. Ent. v. p. 9 (1873) (name for the black var. only) ;
Dury, Cincinnati Soc. Nat. Hist. i. p. 12 (1878) (Cine., abundant) ; Stevens., I'.ii/che iv. p. 233
(1885)(Poughkee[sie, N.Y., August); Eimer, Arth. Vrrwandsch. Schmett. ii. p. 142. t. 8. fig. 1. ?
(1895) ; Jeheb., Ent. News .xvi. p. 1 1 1 (1905) (dark J produced by diseased food !).
Papilio turnus dim. var. $ glaucus, Edwards, Trans. Amer. Ent. Soc. vi. p. 11. sub n. 17 (1877)
(Southern New York, and Wisconsin to Uulf of Mexico ; Kansas to Texas).
Jasoniades r/laucus, Scudder, Bull. East. U.S. ii. p. 1288. t. 8. fig. 1, t. 13. fig. 10, t. 26. fig. 8, t. 35.
fig. 31—34, t, 40. fig. 10, t. 57. fig. 4, t. 01. fig. 13, t. 60. fig. 1, t. 08. fig. 18, t. 76. fig. 15. 26. 28,
t. 80. fig. 7—10, t. 85. fig. 1—4 (1889) (morph., metam., habits, etc.) ; id., Psyche viii. p. 208.
t. 5. f. 3, 1. juv. (1898).
Papilio turnus dim. form, ijloiicus^ Edwards, Bull. U.S. Nat. Mus. xxxv. p. 11 (1889) (liter, relat. to
metamorphosis).
Papilio turnus glaucus, Skinner, Eid. News iv. p. 82 (1893) (N. Carolina) ; Jones, ibid. iv. p. 190
(1893) (Richmond Co., N.C.) ; Bubua, ibid. viii. p. 98 (1897) (Cleveland, Ohio ; three specimens) ;
Eimer, Orthogen. p. 37. fig. 19. J (1897).
Papilio turnus a.h. Jietcheri Kemp, Ent. Neios xi. p. 481 (1900) (N. Jersey).
Papilio turnus J rjlaucus, Grote, Canad. Ent. xxxiv. p. 94 (1902) (J glaucus represents the original
colour of the insect ! ! ; P. turnus allied to troilus).
Papilio turnus uu.itralis Maynard, Man. N.Amer. Butt. p. 215, n. 15a (1891) (Florida).
Papilio glaucus turnus, Dyar, I.e. lii. p. 2. sub n. 11 (1902).
Papilio glaucus australis, id,, I.e.
( 586 )
£?. Snbmarginal bnflF spots of underside of forewiug separate, bnt in small
spring specimens more or less continaous. Black abdominal border of hiiidwing
narrower than the 3-ellow interspace between it and the cell.
The third black band of the forewing, above, varies from being restricted to
the cell to being extended a little beyond M-, in most specimens not ijnite reaching
M-. The band situated on or beyond the cross-veins bears occasionally several
yellow spots. The first snbmarginal spot is rarely absent from the uppersidc.
The black markings of the iipperside are occasionally so much extended as to
occupy the greater part of the wing. The melanism of these males, of which
several have been described, is not the same as that of the females, in the black
female the grotind-colour having assumed a blackish or brown tint, the bands
remaining normal, while in these black males the bands are extended.
¥ . The form resembling the male is the ordinary one in the northern districts
of the range, while in the southern districts the dark form is the more frequent of
the two. Intermediate specimens are comparatively rare. Specimens have been
recorded in which the left side resembles the one female, while the right side
resembles the other.
We have four females which are intermediate between tlie two female forms.
In one of them (New Jersey, July 1898), a black female, the postdiscal area of the
hindwing is more or less ochraceous in posterior half of wing, both alwve and
below. The second black female (Staten Island, August 1901) has traces of the
yellow ground-colour on both wings, especially on the underside. The third
specimen (.Jeiferson Co., Kentucky, September 1897, C. R. Troxler, senr.) is much
more extended yellow than the second, especially on the disc of the hindwing.
The fourth female has the yellow parts merely shaded with black (Baltimore,
August 1894). The two forms of the female may conveniently be referred to as
?-f. ylaucus and ¥-f. turnus.
Hah. Atlantic district, from Florida to New England, westward to the
Mississippi basin.
In the Triug Museum 100 cJcJ, 32 ? ? , and some larvae and pupae from:
Texas; Florida; Carolina; Georgia; Tennessee; Kentucky; Staten I.; New Jersey ;
Buffalo ; Illinois.
h. r. glaucus canadensis snbspec. nov.
Pajtilio ttirims,Kiihj (mm Linne, 1771, err. det.), Fauna Bor. Amer. iv. p. 286. n. -10! (18.37)
(Canada) ; Gosse, Caiiad. Nairn: p. 183. fig. (1840) ; id., I.e. p. 194. 223. 2;t3 (1840) (habits) ;
Felder, I.e. (18G4) {part'nu) ; Reed, Cwiach Eiil. i. p. 19 (1868) (London, Ont.) ; S.iijiiJ., ibid.
i. p. 22. 74 (18G9) (London, Ont. ; early stages ; imago in May and later, perhaps two broods) ;
Beth., ibid. ii. p. 8 (1869) (Toronto, July) ; Riley, ibid. iv. p. 37 (1872) (Peterboro Co., Ont,
May) ; Grote, Bidl. linffah Sm: Nal. Sci. i. p. 185 (1873) (Anticosti ; small) ; Saund., Cumid.
Enl. vi. p. 2. fig. 1 (1874) ; id., I.e. p. 140 (1874) (Essex Co.) ; Lym., ibid. p. 158 (1874); Saund.,
Repl. Ent. Sue. OiUari,, p. 20. fig. 13. 14 (1874) (life hist.); Bates, Eid. J/o. Mag. xi. p. 244 (1875)
(Newfoundland ; small, pale, with narrow black border to hindwing) ; Edw., Butt. N. Amer. ii.
Pap. t. 5. fig. 1. (J (1877) ; Beth,, Caiiad. Enl. x. p. 217 (1878) (Canada) ; Saund., Jiepl. Ent.
Sac. Oiilario p. 73. fig. 40. 41 (1880) ; Fletcher, ibid. p. 62. fig. 45 (1881) ; Gosse, Cawid. Enl. xv.
p. 48 (1883) (Newfoundland) ; Saund., ibid. p. 204 (1883) (larva on Magnolia acnminala,
exceptional) ; id., Jiept. Ent. Soc Ontario p. 16 (1884) (larva on Magnolia) ; Fyles, ibid. p. 63.
fig. .32 (1884) ; id., ibid. p. 44 (1889) (on apple, etc.) ; Moffat, ibid. p. 101 (1899) (London,
Ont.) ; Fletcher, ibid. p. 79. 83 (1889) (Nepigon, L. Superior; eggs laid on aspen, early
in July) ; Mayn., Man. N. Amer. Bnlt. p. 11. n. 16. t. 1. fig. 1. ^ (1891) (.partim) ; Fletcher,
Kept. Ent. (ioc. Ontario xx. p. 38. fig. 12. 13. 14 (1890) (life hist., melanic cj) ; Fyles, ibid.
xxiv. p. 39. fig. 19 (1S94) (Quebec, larva on Anielanelrer eanadensix) ; Danby, Joxirn. N. York
Ent. Soc. ii. p. 33 (1894) (Vancouver I., common) : Grant, Canad. Enl. xsviii, p. 273.
( 587 )
fig. 23. aberr. (189G) (Orillia, Ont., usually common, v. vi. vii.) ; Holland, BiM. Book p. 309
n. 4 (1899) (parlhn ; Sitka; Canada); Lyman, Caiiad. Ent. xxxii. p. 119 (1900) (Dawson,
Yukon) ; Bethune, liept. En/. Sor. Ontario xxx. p. 101 (1900) (parasites : Tragus fulviprs rare,
T. exesorius common) ; id., I.e. p. 104(1900) (first speoim. of lurmis May 28) ; Dod, Camul. Ent.
xxxiii. p. 171. n. 81 (1901) CAlberta, June, fairly common) ; Moffat, Repl. Ent. Sor. Ontario
xxxii. p. 51. fig. .30 (1902) (middle of May to middle of Aug. this year) ; Gibson, ibid, xxxiii.
p. 7C. fig. 53 (1903) (larva) ; Dyar, Proc. U. S. Nat. Mus. xxvii. p. 782 (1904) (Kootenai).
PajiiUo thoas, Heust. (nou Linne, 1771, err. det.), Canad. Ent. xi. p. 239 (1879) (St. John's, N.B.).
Papilio glaums, Linn., var. Iiirnus, Linn., Weir, Entnm. xiv. p. 99 (1881) (Hudson Bay, July).
PapiUo glaucus, Dyar, Bull. U.S. Nat. Mus. lii. p. 2. n. 11 (1902) {partha ; Canada ; Ala-ska).
Papilio glaucus L. a. turmis, Cockle, Rept. Ent. Sac. Ontario xxxiv. p. 90 (1904) (Kaslo, B.C.).
c? ? . A small form. On the upperside the third band (from base) of the
forewing reaches nearly always down to M" in male, to SM- in female; marginal
sjiots thinner and longer than in P. glaucus glaucus. The black abdominal
border of the hiudwing broader than the yellow interspace between it and the cell.
On the luulerside the submarginal spots of the forewing form a continnous
band, only the last one or two spots being separated. The abdominal border
of the hindwing as broad as above ; snbmarginal spots on the whole less curved
than in P. g. glaucus ; the blue spots larger, and the black proximal borders to
them on the whole more straight,
//(///. Newfoundland ; Auticosti ; New Brunswick ; Canada ; northern districts
of British Columbia ; Alaska ; name-type from Newfoundland.
In the Tring Museum SO c?(?, 2 ? ? , from : Newfoundland, June 1898; Bale
St. Claire, Auticosti; Rainy Lake, June 1S92 (Daggett); Ottawa; Didsbury,
Alberta, June 1904 ; British Columbia.
80. Papilio rutulus Lucas (18.52).
Papilio rutulus Lucas, in Guer., Per. Zoul. (2). iv. p. 158 (1852) (March ; California) ; Boisd., Ann.
Soc. Ent. France p. 279. n. 1 (1852) (California) ; Doubl., Westw. & Hew., G(?n. Diurn. Lrp. ii.
p. 529 (1852) (" rutulus Lucas" ; Boisd. not mentioned, his description beiug of a later date) ;
Gray, Cat. Lep. Jus. Brit. Mus. i. Pup. p. 24. n. Ill (1852) ("var." excl.) ; id., List Lep. Ins.
Brit. Mus. i. Pap. p. 32. n. 118 (1856) ('-var." excl.) ; Morris, Syn. Lep. N. Amer. p. 3. n. 3
(1862) ; Weidem., Proc. Ent. Soc. Philad. ii. p. 148 (1863) ; Felder, Verh. Zool. Bot. Ges. Wien
xiv. p. 314. n. 350 (1864) (California ; Sonora) ; Behr, Stett. Ent. Zeit. xxii. p. 215 (1866)
(Calif.) ; Beak., Proc. Ent. Soc. Philad. vi. p. 125 (1867) (Pike's Peak, Colo. ; diff. from turmts
and daunus) ; Kirby, Cat. Diurn. Lep. p. 565. n. 315 (1871) (" var." excl.) ; H. Edw., Proc. Cal.
Ac. Sc. V. p. 165 (1873) (chrysalis) ; Mead, in Wheeler, i?c^)<. Expl. Surv. v. Zuol. 8. p. 741
(1875) (S. Utah ; Colorado, June, one brood only) ; Edw., Trans, Amer. Ent. Soc. vi. p. 11. n. 16
(1877) (Pacific States ; Rocky Mts.) ; Gerh., Macro-Lep. N. Anwr. p. 25. n. 445 (1878) ; Strecker,
Butt. Moths N. Amer. p. 70. n. 11 (1878) (Cahf. ; Oregon, etc.) ; Edw., Papilio ii. p. 112 (1882)
(early stages) ; Hagen, ibid. ii. p. 160 (1882) (charact. and distrib. ; " Kamscbatka" loci error);
id.. Psyche iii, p. 415 (1882) (Washington Terr., western form .and turnus eastern form ;
rutilus found as far east as Salt Lake, Utah, and Fort Bridger, Wyom.) ; Edw., Pajiilio iii.
p. 4 (1883) (dist. spec.) ; id., C'<i«a(;. Ent. xv. p. 169 (1883) (larva diff. from that of turnus) ;
Butl., Journ. Linn. Soc. Lnnd. xvi. p. 472. n. 60a (1883) (Lake and Tehama Cos.) ; Behr, Bull.
Cal. Ac. Sc. I. p. 64 (1884) (Calif, common ; 1. on Amygdalaceae) ; Edw., Butt. N. Amer. ii.
Pap. t. 12. 13. (1884) (cJ ? metliam.) ; id., Canad. Ent. xvii. p. 112 (1885) (larva on willow) ;
Denton, ibid. xxi. p. Ill (1889) (Nevada) ; Skinn., ibid. xxi. p. 238 (1889) (Ft. Qu'Appelle,
N.W. Terr.) ; Edw., Bull. U.S. Nat. Mus. xxxv. p. 12 (1889) (liter, relat. to metamorph.) ;
Mayn., Man. N. Amer. Butt. p. 12. n. 17. fig. 9 b (1891) ; Pack., Fifth Rept. U.S. Ent. Comm.
p. 625 (1891) (egg and larval stages) ; Wright, Canad. Ent. xxiv. p. 73 (1892) (how to get
eggs from ?) ; Haase, Unlersuch. Mimicry i. p. 89 (1893) ; Oslar, Ent. News iv. p. 226 (1893)
(Los Angeles, Feb.) ; Cockerell, Trans. Amer. Ent. Soc. xx. p. 353. n. 645 (1893) (Colorado) ;
Snyder, Ent. News v. p. 133 (1894) (Park City, Utah) ; Cunningh., Ent. News vi. p. 251
(1895) (Ft. Klamath, Oregon) ; Twog., ibid. viii. p. 31 (1897) (Riverside, Calif, commop,
Feb, to Oct.) ; Christ, Mittlt. Schweiz. Ent. Ges. ix. p. 278 (1897) ; Holland, Butt. Baokp. 309 •
38
( 588 )
n. 3. t. 45. fig. 1 cJ (1899) ; Denton, .W.ths n«tt. r..9..1. ii. p. 337. fig. (1898—1900) ; Brown,
Eut. Neics xii. p. 301 (1901) (Salt Lake City, commou) ; Dyar, Bull. U.S. Nat. .\Ins. lii. p. 2.
n. 10 (190-') (Pacific States, Rocky Jits.) ; id., Proc. U.S. Nat. Mux. xxvii. p. 782 (l'.H)4)
(Kootenai ; larva on birch, etc.) ; Wright, liiitl. West Coast ed. ii. p. 84. n. 19. t. 3. fig. 19
(1906) (lowland species).
Papilio nilulu.i, var. or ab. ?, Strecker, Leji. Itlmp. Hit. p. 128 (1877) (Arizona).
Papilio tiirniis var. (geogr.) rntulu,'!, Oberthiir, Et. dEiit. iv. p. G8. sub n. 188 (1880).
Pajiilio ridtihis var. arhimeiisis Edwards, Piii)iliu iii. p. 4 (1883) (Arizona) ; id.. Butt. N. Amcr. ii.
Paj). t. 13. cJ ? (1884) ; Winkle, Camid. Eiit. xxv. p. 212 (1893).
Papiliu turmis, Butler, Jnurti. Linii. Sue. Laud. xvi. p. 472. n. 61 ( 1883) (Tehama and Mendocino Co.).
Papilio ruttdus var. ammoiii Behrens, Cauad. Eut. xix. p. 199 (1H87) (orange colour) ; Winkle, l.c.
xxv. p. 212(1893).
Papilio nitidus (!), Cockerel), Trans. Anier. Eut. So,: xx. p. 353 (1803) (lajs. rah).
Papilio turiius rutuliis, Elmer, Arlh. Verwaiidlsch. Srhmrtt. p. 83 (1895).
Papilio rutulus arizo/wiisi.i, Blaynard, Man. N. Amer. Butt. p. 13. n. 17a. fig. 10a (1891) (Arizona;
New Mexico ; Colorado) ; Dyar, Bull. U.S. Nat. Mas. lii. p. 2. sub n. 10 (1902).
Papiliu rutulus ammou', Maynard, I.e. (Nevada) ; Dyar, l.r.
Papilio arizomnsis, Wright, Butt. West Coast ed. ii. p. 85. n. 20. t. 3. fig. 20 (1906) (S. Arizona).
Papilio ammoiii, id., l.c. n. 21 (1906) (" not seen ").
(??. Sexes similar. Forewiug on the whole rather more poiated than in
J', glauciis, the subapical e->-mark rarely centred with yellow, or at least the
yellow colour within the mark rarely so e.Ktt^nded as to leave only the edges of
the spot j)nre black, the c»-mark proximally often separate from SC, the proximal
portion being often almost isolated, especially on underside ; first submarginal spot
linear like the others, usually larger than the second ; the subbasal yellow band
less narrowed costally than in P. glaiicus, the third black band on the whole longer
than in specimens of P. glaucus of the same size, reaching usually beyond M- ;
yellow fringe-spots very thin. — —Black abdominal border of hiudwing always wider
than the yellow interspace between it and cell ; first submarginal spot small or
absent, seldom nearly as large as the second, never orange in either sex.
The buff submarginal spots of the underside of the forewiug merged together
to a continuous line. No orange patches on disc of hiudwing, or only traces of
them ; submarginal spots less orange than in P. glaucus, usually only the last two
being of this colour.
We do not tind any fairly constant differences between specimens from Arizona
and California.
Early stages see Wright, l.c.
Genitalia: 6. Harpe truncate, apex not produced beyond base of process;
dorsal hook longer than in P. glaucus, simple, non-dentate, there being never
two hooks, as is always the case in P. glaucus. ? . Antevaginal process obtuse ;
lateral dentate flaps larger than in P. glaucus, extending farther frontad than in
that species.
Hab. British Columbia to Arizona.
One of our specimens from the Frazer K. (June I'JOl) is worthy of being
specially mentioned. The upper submarginal spots on the upperside of the forewing
are rounded, being connected with the margin by means of dispersed yellow scales.
The submarginal spots of the underside of both wings are extended to tlie margin,
forming a nearly continuous marginal band, a very little of the black marginal line
being left at the apex of most veins.
In the Tring Museum 27U c?(?, 40 ? ? , from : Senator, Arizona, July 1898
(Dr. Kunze) ; South Park, August 10(11, Cliimncy (4ulcii, June lOiiu, Grand
Junction, July 19ul, Colorado (Usjar); Garfield Co.; Reno, Nevada; North Tulare Ii.,
( 58d )
Califoruia, July 1897 (Purpus) ; McClou-l R., Shasta, June 1884 (0. T. Baron) ;
Siskiyon Co. (0. T. Baron) ; Hoopa valley, July 1896 (Dougherty) ; Quincy,
California, 3400 ft., June and July 1897 (Watson) ; Davis Creek, Modoc Co.,
June 1898 (Mrs. Austin) ; Butte Creek, Butte Co., April 1898 (Mrs. Austin) ; Pine
Creek, Oregon, June 1808 (Mrs. Austin) ; San Reno, California, June 1897 ; Sonora
(Lorqnin ; coll. Felder) ; Gold Hill, Oregon, June and July 1901 (Biedermann) ;
Nicomin I., Frazer R., May and June 1900; Qu'Appelle, Assiniboia, June 1901 ;
Vancouver (A. H. Bush) ; Ozoyoos (Raynolds).
81. Papilio daunus Boisd. (1836).
Papilio (kimttis Boisduval, Sjiee. Gin. Li'p. i. p. 342. n. 182 (IS.'iO) (Mexico) ; Doubl., List Lep. Lis.
Brit. Mus. i. p. IG (1«45) (Oajaca); id.. Westw. & Hew., den. Diurn. Lep. i. p. 13. n. 88 (18+0);
Gray, Cat. Lep. Lis. Brit. .!/«,<. i. Pap. p. 24. n. 109 (1852) ; id., List Lep. Lis. Brit. .Utis. i.
Pap. p. 32. n. IIG (18oG) (Oajaca) ; MOncjtr., Eimm. Corp. Aiiim. Mus. Petrop., Lep. i. p. 2.
n. 30 (18.07) (Mexico) ; Vollenh., Tijthclir. Enl. iii. p. 85. n. 131 (18G0) (Mexico) ; Rid., Pruc.
Ent. Soc. Philad. i. p. 278. fig. 2 (i8G2) (Kansas) ; Felder, Verh. Zool. Bat. Gcs. Wien. xiv.
p. 31.'). n. 346 (18G4) ; Beak., Proc. Ent. Soc. Philad. vi. p. 124 (18G7) (Colorado) ; Kirby,
Cat. Diurn. Lip. p. 5fi4. D. 312 (1871); Strecker, Lep. Pliop. Het. p. 45. t. G. fig. 1. (J, 2. ?
(1873) (Rocky Mts. ; Vera Cruz) ; H. Edw., Proc. Cat. Ac. Sc. v. p. 325 (1874) (pupa, larva
noticed) ; id.. Bull. N. Amer. ii. Pap. t. 2. (J (1875) ; Mead, in Wheeler, Pept. E.rpl. Surr. v.
Zool. 8. p. 741 (1875) (Colorado, S. Utah) ; Kirby, I.e. p. 811 (1877) ; Edw., 'frans. Amrr. En'.
Soc. vi. p. 11. n. 18 (1877) (Arizona to Montana ; Oregon) ; Uhler, in Hayd., Bull. U.S. Geol.
Geor/r. Surv. iii. p. 3oG (1877) (Clear Creek, Colo.) ; id., I.e. p. 7G5 (1877) (Clear Creek and
Ute Pas.s, Aug. G— 13) ; Gerh., Macro-Lep. N, Amer. p. 25. n. 440 (1878) ; Strecker, Butt. Moths
N. Amer. p. 08 (1878) (Colorado ; N. Mexico ; Mexico ; Central America) ; Oberth., El. d'Ent.
iv. p. 08. n. 191 (1880) (Mexico ; type) : Hagen, Psijche iii. p. 415 (1882) (probably the same as
riUulns) ; id., Papilio ii. p. 163 (1882) (doubtfully distinct from riituln.'<); Edw., ibid, iii p. 2.
(1883) (Arizona, larva & pupa ; Boulder, Colo.) ; id., I.e. iii. p. 158 (1883) (Montana) ; id., Butt.
N. Aiacr. ii. Siippl. p. 1 (1884) (larva) ; Schaus, ibid. iv. p. 100 (1884) (adult larva descr.) ;
Behr, Bull. Cal. Ae. Sc. I. p. 64 (1884) (Calif., local, 1. on Prunus demissa) ; Denton, Canad.
Ent. xxi. p. Ill (1889) (Nevada) ; Edw., Bull. U.S. Nat. Mas. xxxv. p. 12 (1889) (liter, relat. to
metam.); Mayn., Man. N. Amer. Butt. p. 13. n. 18. fig. 9. g (1891) (Arizona to Montana;
Utah ; Nevada ; Oregon ; Mexico) ; Godm. & Salv., Biol. Centr. Amer., Lep. Rliop. ii. p. 240.
n. 78. t. 70. fig. 9. genit. (1893) (Oregon and Montana southward to Guatemala ; open grassy
tracts in the higl)Iands) ; Haase, Untersttch. Mimicri/ i. p. 89. fig. 9 (1893) ; Snyder, Ent. News
V. p. 166 (1894) (Park City, Utah, larva, eggs) ; Eimer, Artb. Vertcandtsch. Sclimett. p. 87. t. 5.
fig. 6. (J, 7. ? (1895) ; Christ, Milth. Schwciz. Ent. Ges. ix. p. 278 (1897) ; Eimer, Orthoi/cii.
p. 30. fig. 8 (1897); Edwards, Butt. N. Aincr. in. Suppl. p. 1 (1897) (larva); Holland, £«(/.
Booh p. 310. t. 38. fig. 2. (J (1899) (eastern valleys of Rocky Mts., Arizona, Mexico) ; Denton,
3toths Bull. U.S. A . ii. p. 338. fig. (1898—1900) (Colorado ; New Mexico ; Mexico ; occas. in Kansas
and Texas); Brown, Eiil. N'ews xii. p. 301 (1901) (Salt Lake City, common, June, up to
7000 ft. ; Dyar, Bull. U.S. Nat. ,!/«.<. Iii. p. 2. n. 8 (1902) (Rocky Mts.) ; Hoag, Ent. News xiv.
p. 321 (1903) (S. Luis Potosi, Mcx.); Honeym., Kept. Ent. Soc. Ontario xxxv. p. 61. n. 8 (1904)
(Regina) ; Wright, Butt. West Coast ed. ii. p. 82. n. 16. t. 3. fig. 16 (1906) (rapid flight ; food
plants).
Papilio nmlticaudata Kirby (ex Peale, ined.), Papilio iv. p. 104 (1884) (= daunus).
c? ?. Sexes similar, the black bands and blue spots being larger in the female
than in the male. The hindwing is as a rule much more strongly dentate than
in the allied species, the tail being very long and tooth M' being prolonged to
a second tail. However, sometimes the dentition is hardly more prominent than
in P. ridultis. The blaciv bauds are always narrower than in P. nitulus ; the
e-j-sliaped subapical mark of the forewing is always centred with yellow ; the fourth
black band is narrower than the yellow apical cell-space ; second black band seldom
as broad as the yellow cell-space situated at its distal side; the third band la
the male often stopping short at the median vein, while in some other males
( 590 )
and ill aluidst all females it reaches to near SM-; the tifth band also very variable
in length. The median band of the hindwing is often vfry thin, sometimes
vestigial ; the black abdominal border is narrower than in /-*. rutuliis, while the
distal border is usually broader. The ground-colour becomes deep yellnw (also in
the allied species) in consequence of exposure to light and damp.
In many specimens there are large orange patches on the underside of the
hindwing, the disc being sometimes washed with orange at the black distal border
also on the ujiperside.
Mexican specimens are on the whole larger than those from Oregon, Washington,
and British Columbia, the females esjiecially being often very large. However, we
cannot find any fairly constant character by which to distinguish northern and
southern specimens, the species being strongly variable individually in all districts.
Genitalia : <S . Harpe broad, either truncate, or the dorsal edge slanting down
to the base of the process ; this edge with two or more teeth ; one dorsal hook,
usually bearing a few teeth. ?. Autevaginal lube triangular; lateral ridge very
large, strongly dentate.
For early stages see Edwards, Wright, etc.
Hab. British Columbia and Alberta southwards to Guatemala, eastwards to
Colorado and Vera Cruz.
In the Tring Museum 170 c?(?, 05 ??,from: Ozoyoos, British Columbia
(Raynolds) ; Gold Hill, Oregon, July 1901 (Biedermanu) ; Modoc Co., July 1S'J7,
and Butte Co., April 1898 (Mrs. Austin) ; Verdi, Nevada, 7000 ft., June 1896 ;
McCloud R., Shasta, June 1884 (0. T. Baron) ; Siskiyou Co. (0. T. Baron) ; Tuckes,
California, GOOO ft. ; North Tulare R., California, July 1S97 (Purpns) ; Quincy,
California, June (Watson) ; Prescott and Senator, Arizona, June, July and August
(Dr. Knuze) ; Verde R., Copper Basin, Thumb Butte and Huachuca Mts., Arizona,
July— September 1903 (Oslar); Denver, Colorado (Mason); Grand Junction,
South Park and Chimney Gulch, Colorado, June — August 1900 and 1901 (Oslar);
Las Vigas, May 1S90, Jalapa, July 1896, Cholula, March 1896, Orizaba, March
1896 (W. Schans) ; Oaxaca, July 1896 (W. Schans) ; Guadalajara, August 1890
(W. Schans) ; Cuernavaca, 4000 ft, July 1904 (A. Hall) ; Guerrero (0. T. Baron) ;
Salvaticrra, Guanajuato.
82. Papilio eui-ymedon Lucas (1852).
Pajtilio eurijiiiechn Lucas, in Guurin, Rev. Zonl. (2). iv. p. 140 (1852, March) (California) ; Boisd.,
Aim. Hoc. Kill. France p. 2iiO. n. 2 (l»iu2) (California) ; Doubl., Westw. H Hew., Geii. Diiiiii.
Lep. ii. p. 529 (1852) (quote " eurijmedon Lucas," not mentioning Boisd., whose description came
out later) ; Morris, Sijii. Lep. N. Ainer. p. 4. n. 4 (1862) ; Weidem., Pmc. Eiit. Soc. Pldltid. ii.
p. U7 (18G3) ; Behr, .S7«'((. Ent. Zeit. xxvii. p. 215 (ISliO) (Calif.) ; Reak., Proc. Eiit. Soc.
Phikid. vi. p. 120 (18G7) (Pike's Peak, Colo. ; Washington Terr.) ; Kirby, Cat. Dhnii. Lep.
p. 5fi5. n. 314 (1871); Strecker, Lep. Rhop. JJel. p. 25. t. 4. fig. 1 (1873) (California;
"Washington ; Vancouver L) ; H. Edw., Proc. Cul. Ac. Sc. v. p. 1(14 (1S73) (larva, pupa) ;
Edw., Pull. y. Ainei: ii. Pup. t. 1 (1874) (metara.) ; Mead, in Wheeler, Rcpl. Expl. Sm-r. v.
Zool. 8. p. 742 (1875) (Colorado) ; Edw., Trans. Aiiier. Eid. Soc. vi. p. 11. n. 15 (1877) (Calif,
to Brit. Col. ; Arizona to Montana) ; Cierh., Macro-Lep. X. Am. p. 25. n. 444 (1878) ; Strecker,
Diilt. Moths N. Am. p. 70. n. 12 (1878) (Calif. ; Oregon ; Vancouver I.) ; Oberth., El. d'Ent. iv.
p. 68. n. 18y (1880) ; Edw., Papilio iii. p. 158 (1883) (Montana) ; Behr, Unll. Cal. Ac. Sc. i.
p. 64(18H4) (Calif., common, I. on Hhamims califoniim); Denton, Canud. Eiil. xxi. p. Ill (1889)
(Nevada); }hdw., Bull. U.S. Xal. Mas. xxxv. p. 12 (188',i) (literat. rel. to metamorphosis)':
Majn., .Uati. N. Amer. Bull. p. 12 n. 16 (18'J1); Haase, Uiilersuch. Mimicri/ i. p. 89 (1893) ;
Dyar, Eid. News iv. p. 243 (1893) (life history ; on Rliainmis cali/ornica) ; (Junningh., ibid. vi.
p. 251 (189.5) (Ft. Klamath, Oregon) ; Eimer, Arlb. VerwKiidhch.Schmell. ii. p. 90. t. 5. fig. 6
( 591 )
(1895) (California) ; id., Orthoffen. p. 28. fig. 2 (1897) ; Christ, Mill. ScJnoeiz. Enl. Gex. ix. p. 278
(1807) ; Twog., Enl. News viii. p. ai (1897) (Riverside, Calif., one ex., Aug.) ; Holland, BiUI.
&)oA- p. 308. n. 2. t. 44. fig. 5. cJ (1899) (Mexico to Alaska, eastwards to Colorado); Denton,
Molhs Bull. U.S.A. ii. p. 341) (1898—1900) ; Brown, Enl. News xii. p. 301 (1901) (Salt Lake
City, common from 4500 to 6000 ft.) ; Dyar, Bull. U.S. Nal. Mm. lii. p. 2. n. 9 (1902) (Pacifio
States ; Colorado) ; id., Proi: U.S. Nal. Mux. xxvii. p. 782 (1904) (Kootenai, Juno, larva on
Cewmlhus) ; Wright, Bull. WesI Coast ed. ii. p. 83. t. 3. fig. 17 (190G) (mountain species, up to
8000 ft.).
PapUiu ruluhis var. a. Pajiilio eurymedon, Gray, Cat. Lep. Ins. Brit 3{us. i. Paji. p. 24. sub n. Ill
(1852) (California) ; id., List Lep. I71.S. Brit. Mus. i. Pap. p. 32. sub n. 118 (185G).
Papilio rululiis var. euri/medou, VoUenhoven, Tijdschr. Eul. iii. p. 85. n. 133 (1860).
Papilio euriinodou (!), Felder, Verh. ZmA. But. Ges. Wieii xiv. p. 313. n. 348 (1864).
Papilio alhanus Felder, l.r. xiv. p. 314. n. 349 (1864) (nom, inde.sor. ; California) ; id., Reise Norrira,
Lep. p. 93. n. 71. (1865) ; Edw., Papilio ii. p. 122 (1882) (J (J in the mountains or at 2000 ft.
are nearly always this form) ; Butler, Jaurn. Liuu. Soc. Loud. xvi. p. 472. n. 60 (1883)
( = ? eurijmeduii ; Mendocino) ; Wright, Butt. West Coast ed. ii. p. 84. n. 18 (1906) (characters
do not hold good).
Papilio rutiilus var, a. P. albanus, Kirby, Cat Diurii. Lep. p. 565. sub n. 315 (1871).
Papilio lewisi Kirby (exPeale, ined.), Papilio iv. p. 104 (1884) (= eurijmcdon).
Papilio euryinedon var. ulhauus, Winkle, Canad. Enl. xxv. p. 212 (1893).
Papilio rulahis, Danby, Joiirn. N. York Eiit. Soc. ii. p. 33 (1894) (Vancouver I., common ; =
eurymedon, as stated on p. 141).
Papilio eurymedon alhanus, Dyar, I.e. (1902).
S ? . The black bands on both sides of the wings much heavier than in either
P. rutulus or dauniis, bnt their width very variable. At higher altitudes there
occur often specimens in which the blaclv distal marginal border is mnch reduced,
being about the same width on the forewing between R- and M'- as the yellowish
white discal band. This is Felder's P. albaims, I.e., which may nomenclatorially
be distinguished from the ordinary form as —
f. mont. albanus Feld. (1865).
Besides two specimens from Felder's collection we have this form from Colorado
and California.
The costal cn-mark of the forewing is not centred with creamy buff in
P. eurijmedon either above or below ; the submarginal spots of the forewing
above are more or less linear; the powdery huffish line on the black postdiscal band
of the underside of the forewing is narrow or absent. In Colorado specimens the
black bands are on the whole a little narrower than in Califoruiau ones.
Genitalia : cJ. Ilarpe dorsally less elevate in the allied species, the hook being
shorter, bearing nsnally a few teeth, there being generally no other prominent
teeth at the oblique dorsal edge of the harpe ; but this edge occasionally minutely
denticulate, the ventral edge being also often provided with one or two teeth ;
apex of harpe as in P. riUuhis, not produced, the process standing at the ventral
apical corner. ? . Antevaginal process pointed, lateral flaps rather larger than
in P. rutulus.
For early stages see Dyar, I.e. (1893).
Ilah. British Colnmbia to Southern California and Colorado.
Holland, in Butt. Book, records it from Alaska and Mexico, which re piires
confirmation.
In the Tring Museum 180 cJc?, 32 ? ? , from : Glen wood Springs and Chimney
Gulch, Colorado, June 1900 and 1901 (Oslar) ; Mendocino, California, August
(0. T. Baron) ; McCloud II., Shasta, June 1884 (0. T. Baron) ; Quiucy, California,
May, June and July 1807 (Watson) ; Butte Co., Lake Co., and Modoc Co., June
( 592 )
and July 1897 and 1898 (Mrs. Austin); Hoopa Valley, Jnly 1890 (Dotighei-ty) ; Gold
Hill, Oregon (Biedermann) ; Ozoyoos, British Columbia (Raynolds) ; Nicoiuiu I.,
Frazer R., May and June 1900; Kaslo; New Westminster (A. D. Jones); Qii'Appelle,
Assiniboia, July 1 9i 1 1 .
83. Papilio alexiares Hopflf. (18GC).
PiijilVn ule:rhirc!i Hopffer, Stelt. Ent. Ziil. xxvii. p. 31. n. 12 (ISOO) (Mexico).
(? ? . In sbape of wings intermediate between P. rutxlHs and /'. gluucus.
Forewing : distal margin straight or feebly concave ; snbbasal yellow interspace
acutely triangular, more pointed anteriorly than in P. glaitcus ; third band reaching
to IP or beyond. Hindwing : black abdominal border a little narrower than or
as broad as the yellow interspace between it and cell at base of M'- ; black median
band proximal of SC-.
Undersidf : costal cs-mark of forewing centred with yellow, the posterior edge
or the proximal dilated portion remaining jiure black ; the yellow scaling within
this mark continuous with the greyish or huffish yellow band situated on the black
jiostdiscal band, the pnre black edges of the latter being sharply defined both
prdximally and distally ; yellow submarginal spots linear, forming a continuous (or
nearly) band which porcej)tibly widens costally. Hindwing with large or small
orange patches on disc from R' backwards.
Genitalia : c? . Apex of harpe produced beyond base of ventral process, acuminate,
a long, curved, simple, conical tooth at dorsal angle, a few small teeth between it and
the apex nf the harpe. ? not dissected.
Early stages not known.
Ilab. Eastern Mexico.
Two subspecies.
a. P. alexiares garcia subsp. nov.
(?. Upperside : black bands of both wings much narrower than in P. ale.r.
alexiares. Forewing : third black band not extending beyond M" ; yellow apical
cell-baud about as wide as the black baud outside it ; black distal border not
wider between R- and M^ than the yellow discal band ; a row of eight distinct
submarginal spots, which are larger than in P. alex. alexiares ; yellow discal
spots SC'^R' also larger than in that form. Hindwing : black distal burder
narrower than in P. alex. alexiares, especially behind, the yellow interuervular
patches around apex of cell therefore larger ; submarginal spots and orange anal
marginal spot much larger, tail slenderer and less curved than in the next form ;
submarginal spot M' — M- orange, small.
Underside : yellow submarginal line of forewing more or less distinctly
interrupted at the veins. Hindwing : on disc between R' and SM- large
elongate-triangular orange patches ; iirst and sixth submarginal spots and anal
marginal spot totally or for the greater part orange, the other submarginal spots
slightly or not at all touched with orange.
Hab. Monterrey, San Luis Potosi.
In the Tring Museum 3 c?c?.
fj. J', alexiares alexiares HopfF. (18GG).
PaijiVw alexiares Hopffer, I.e. ; Kirby, Cat. Dimii. Lep. p. 567. n. 328 (1871) ; Godm. & Salv.,
Biol. C'enlr. Amer., Lep. Rhop. ii. p. 241. n. 80. t. 72. fig. 6. 7. ^ (1893) (Cuosta de Misantia).
(? ? . Upperside. — Forewing : the yellow jiortions often more or less shaded
( 5'J3 )
over with black ; third black band extending beyond M- ; yellow apical cell-band
narrower than the black band standing at its distal side ; black distal border wider
thronghout than the yellow discal band ; submarginal spots thin, posterior ones
absent or vestigial. Hindwing: anal marginal spot small, slightly orange; black
distal border sometimes almost tonching cell.
Underside : yellow submarginal line of forewing broad, continuous, only the
last one or two spots standing separate. Hindwing : orange discal spots small.
Ilab. Vera Cruz : Cnesta de Misantla.
In the Tring Museum 2 c?c?, 2 ? ? .
84. Papilio pilumnus Boisd. (1830).
J'ajiHiii pihiiiiniis Boisduval, Spei-. Gen. Lep. i. p. 340. n. 181 (1836) (Mexico) ; Doubl., List Lfji.
Ins. Brit. Mils. i. p. 1(5 (1845) (Mexico) ; id., Westw. & Hew., Gen. Diiirn. Lep. i. p. 13. n. 89
(1846) ; Gray, Cat. Lep. L,^. Brit. .1/»s. i. Fuji. p. 24. n. IIU (1802) ; id., List Lep. Inn. Brit.
Mus. i. Pap. p. .^2. n. 117 (1850) ; Mc'nt'tr., Enurn. Corp. Anini. Miis. Pelmp.^ Lep. i. Supjil.
p. 68. n. 1116. t. 7. fig. 2. ? (1857) (Mexico) ; id., I.e. ii. p. 110. n. 1116 (1863) ; Vollenh.,
Tijdsclir. Ent. iii. p. 85. n. 130 (1860) (Mexico) ; Weidem., Pror. Enl. Soc. Pliihid. ii. p. 149
(1863) (" probably ? of P. danmi.H") ; Felder, Ver/i. ZooL Bot. Gci. Wien xiv. p. 313. n. 345
(1864) (■' uot ? of ihimni.^ ") ; Realc, Proc. Enl. Soc. Philad. vi. p. 127 (1867) (New Mexico ;
?also Texas and Mexico); Kiiby, Cat. Diurn. Lep. p. 564. n. 311 (1871) ; Stvecker, Lep. Rhnp.
Hel. p. 13. t. 2. fig. 3 (1873) (New Mexico ; Vera Cruz) ; Mead, in Wheeler, Rept. E.rpl. Surv.
Y. Zool. 8. p. 741 (1875) (New Mexico) ; Edw., Trans. Amer. Ent. Snr. vi. p. 11. n. 19 (1877)
(Arizona ; New Mexico) ; Gerb., Macrn-Lep. N. Amer. p. 25. n. 450 (1878) (New Mexico) ;
Strecker, Biilt. Maths N. Amer. p. 08. n. 8 (1878) (New Mexico ; Mexico ; Central America) ;
Oberth., El. d'Enl. iv. p. 68. n. 190 (1880) (Mexico ; 2 (J (J, typ. specim.) ; Schau?, Papilio
iv. p. 100 (1884) (descr. of larva & pupa) ; Edw., Bull. U.S. Nat. Mus. xxxv. p. 12 (1889)
(liteiat relat. to metamorpb.) ; id., Bntt. N. Amer. iii. Pap. t. 2. J. $ (1889) ; Mayn,, Man.
N. Amer. Butt. p. 14. n. 19. fig. 9. c (1891) (Arizona ; Mexico) ; Haase, Dntersuch. Mimicry i.
p. 90 (1893) ; Godm. & Salv., Biol. Cenlr. Amer., Lep. Rhap. ii. p. 241. n. 79. t. 72. fig. 10.
genit. (1893) (New Mexico ; Mexico ; Guatemula ; open grassy plains ; " Colorado " errore V) ;
Eimer, Arlh. Verwandtsrh. Sclimelt. ii. p. 84. t. 5. fig. 3 (1895) (Mexico) ; Christ, Miltli. ScJiweiz.
Ent. Ges. ix. p. 278 (1897) (southern form of dtinnns'>) ; Edw., Bull. N. Anter. iii. Suppl. p. 8
(1897); Holland, Bull. Book p. 310. t. 38. fig. 3. J (1899) (Mexico ; occasionally in Arizona);
Denton, Moths Bull. U.S.A. ii. p. 339 (1898—1900) ; Dyar, Bidl. U.S. Nat. Mus. Iii. p. 2. n. 7
(1902) (Texas ; Arizona ; Mexico).
In pattern a primitive form, being of all species the nearest approach to the
ancestral form from which the groups of tlioas, hjcophron, glaucus and troilus
originated. It agrees in pattern best with the Glaucus Group of species, but dilfers
in having in the hindwing the nearly symmetrical cell of P. lycopkron. The tibiae
and tarsi are as pale as in P. (/was, but less green. The two sharply defined orange
spots M' — SM- on the iipj/cr.-</i/e oi ihn hindwing and the broad black antemedian
band of the underside centred with drab are the most characteristic features iu the
pattern of P. pilumnus. These orange spots are sometimes preceded by two more
spots, which are of the same colour, but not sharply defined. The discal portion of
the apical cell-band of the forewing is rather variable, being usually continuous,
while in a few specimens the last veiu-spot is separated from the band, being minute
or vestigial. The yellow or yellowish line distally of the yellow tripartite band on
the upperside of the forewing is sometimes absent, while it is rather broad in other
specimens, especially in Guatemalan individuals.
Genitalia: <?. Tenth tergite long, feebly spatulate ; sternite on each side with
an obliquely transverse double ridge, both low, but rather sharply cariuiform ; harpe
broad, produced at apex into a sharp tooth, ventral edge nearly straight, the short.
( 594 )
oblique, dorsal edge dentate, tlie number and size of the teeth variable. ? no
dissected.
Early stages described by Schaus, I.e., nearest to those of P. Iroilus.
Hah. Arizona to Gnatemala.
In the Tring Museum 28 cJc?, 1 ?, from : Arizona ; Monterey, Mexico ; Las
Vigas, Mexico, June 1896 (W. Schans); Orizaba, May 1896 (W. Schaus) ; Jalapa,
February 1896 (VV. Schans); Oaxaca; Guerrero (0. T. Baron) ; Palin, AV. Guate-
mala, 2500 ft., August— September 1904 (A. Hall) ; La Antigua, W. Guatemala,
5000 ft., August 1904 (A. Hall).
VII. Troilus Group.
Closely allied to the Anchisiades Group.
SO- of forewing in middle or at two-fifths between SC and SO' ; lower angle
of cell not much more obtuse than upper angle ; D- shorter than D' ; basal cellule
of hindwing longer than in the Ancliiaiados Group, PC more strongly curved. Two
widely separated rows of spots on underside of forewing; hindwing below with
metallic blue patches.
Genitalia : S- Harpe with ventral tooth and apical process, resembling the
harpe of the Anchisiades Group. ?. Vaginal armature also of the type of
P. anchisiades.
Larva with eye-spot on each side of thorax.
Two species.
Antenna and legs black Species No. 85.
Antenna tawny, tibiae and tarsi pale yellowish green . . Species No. 86.
85. Papilio troilus L. (1758).
PajiiVo Efjiies Trnjamis IroUns Linne, Syst. Nat. ed. x. p. 459. n. 6 (1758).
The close relationship between P. troilus and P. palamedes becomes at once
evident on comparing the pattern and structure of the two insects. The pale
snbbasal band on the underside of the hindwing so characteristic of P. palamedes
is sometimes rejireseuted in southern sjiecimens of P. troilus by a line which is
continued across the forewing, there being a ditfase yellow costal streak at tbe base
of the forewing in all specimens. The subapical cell-spot, usually double, on the
underside of the forewing corresponds to the bar oi P. palamedes ; it is occasionally
missing. While most specimens have on the disc of the forewing above and below
only one row of spots, which are usually small on the npperside, being often absent,
some females bear two rows. These rows are in some males represented anteriorly
by a dot in the subcostal fork and a more proximal dot before the fork. Besides
the orange costal spots the npperside of the hindwing bears sometimes in the female
a small orange spot behind SC-. One of the most interesting features in the pattern
of P. troilus is the disappearance of the orange spot W — M' on the underside of
the hindwing. Most siiecimens bear a few orange scales where the spot has been,
sometimes there is even a distinct bnt small orange spot, but in the majority of
specimens the spot is suppressed, the scales having assumed a glaucous buff tint.
First protarsal segment longer than the other four together.
The variability in the dentition of the scaling of the underside is of some
interest. The scales have in most females one tooth less than in the males, being
on the anterior area of the forewing tri- or (jnadridentate and in the 'posterior area
( 595 )
bidentate or entire. On the hindwing the scales of the central orange spots of the
discal row have also on the whole one tooth less than the upper and i30sterior discal
spots and the submarginal spots.
Nenration : SC- of forewiug about halfway between SC and SO', the latter
often a little jirosimal of apex of cell ; D' before middle of cell ; subbasal cellnle of
hindwing narrow, long, PC angnlate.
Genitalia : c?. Tenth tergite spatnlate ; sternite with a lateral lobe which is
more strongly chitinised than the rest of the steruite, divided by an obliipie
transverse depression or groove into two ridges. Clasper elongate-triangnlar, the tip
being rounded off; harpe ending in a long and slender, smooth, acute process as iu
P. anchisiades, the nearly straight ventral margin bearing in the middle a prominent
tooth ; the short dorso-apical edge oblique, dentate. ? . Armature of the same
type as in P. anchisiades ; edge of orifice proximally produced into a long lobe
■which widens apically, the apex being truncate and bearing some teeth ; anterior
surface of lobe longitudinally impressed, posterior surface carinate, except at apex ;
sides of orifice raised into a smaller lobe or ridge ; behind orifice a membranous
tubercle densely covered with minute hairs ; a large lateral ridge, dentate, gradually
becoming lower towards the mesial line of the body, curving towards the middle
lobe on the anterior surface of which it disappears ; between this ridge and the
orifice on each side of the latter a large dentate lobe which stands iu connection
with the lateral ridge as well as with the elevate edge of the orifice. Bristles on
inner surface of anal segment numerous but short.
Early stages first described by Abbot and Smith.
Ilab. Atlantic district of the Nearctic Region.
Two subspecies.
a. P. troilus troilus L. (1758).
F(qnlio Eqiies Trojamm Iroihis Linne, Sysl. Nal. ed. x. p. 459. n. 6 (1758); id., Mus. Lud. Vb: p. 187.
n. C (1764) ; Houtt., Naturl. Hist. i. 11. p. 192. n. 5 (1767) ; Linng, S;/st. Nat. ed. xli. p. 746.
n. 6 (1767) ; Fabr., Syst. Eiit. p. 444. n. 7 (1775) {partim) ; Goeze, Ent. Beijtr. iii. 1. p. 31. n. 6
(1779) ; Cramer, Pap. E.wt. iii. p. 25. t. 207. fig. B. C (1779) ; Fabr., Spec. Iiis. ii. p. 3. n. 9
(1781) {parti III) ; Jabl. & Herbst, Xaturs. Schmett. ii. p. 242. n. 58 (1784) (purdm ; nee fig.) ;
iid.. I.e. p. 291. t. 20. fig. 2 (1784) ; Esper, Ausl. Schmett. p. 21. n. 6. t. 3. fig. 2. (J (1784) (cit,
jxirtim) ; Panzer, Drur>/'.'< Abhikl p. 55. t. 11. fig. 2. 3. 5 (1785) ; Fabr., Mant. Inn. ii. p. 2. n. 9
(1787) {partim) ; Gmelin, Syst. Nat. i. 5. p. 2225. n. 6 (1790) {partint) ; Fabr., Ent. Sijst. iii.
1. p! 4.n. 10 (1793) (partim).
Papilio Eques troilus, Lange, in Linne', Sy-it. Nat. p. 459. n. 5 (17liO) (■' in ludiis ").
Papilin (Troes) troilus, Mtiller, Natui'f. v. 1. p. 567. n. 6 (1774).
Papilio (troilus), Meerburgh, Afb. ZeUh. Gew. t. 4. 7 (1775).
Pterouriis troilus, Scopoli, Intr. Hist. Nal. p. 433 (1777) ; Scudder, Syst. Rev. Aner. Bull. p. 44
(1872;; Sprague, Psyche ii. p. 257 (1879) (Wollaston. Mass., May 25.) ; id., l.e. p. 259 (1879)
(Mass., May 25 to Aug. 5).
Papilio ilioneus Abbot & Smith, Ins. Georgia i. p. 3. t. 2. fig. 1 (1797) (larva, pupa, (J, ?);
Felder, Vcrh. Zool. Bot. Ges. Wien xiv. p. 315. n. 362 (1864) ; Kirby, Cat. Diuru. Lcp. p. 567
n. 326 (1871).
Euphoeades troilus Hubner, Verz. belc. Schmett. p. 83. n. 847 (1818 ?) ; id., Samml. Exot. Schmett. ii.
t. 96 (1822 ?) ; .Scudd., Butterfl. p. 304. 309. fig. 50. 51. 63, 173 (1881) ; id., Butt. East. U.S. ii.
p. 1313. t. 8. fig. 4. 5, t. 27. fig. 1, t. 35. fig. 19. 20, t. 40. fig. 3, t. 57. fig. 6, t. 61. fig. 48. 56, t. 66.
fig. 13, t. 68. fig. 20, t. 72. fig. 8. 9, t. 76, fig. 18. 19. 22, t. 79. fig. 69-73, t. 82. fig. 4-6, t. 85.
fig. 5-7, t. 86. fig. 20-23. 34. 35. 78-80, t. 87. fig. 6. 15. 24 (1889) (metam., morpbol., etc.) ; id.,
Psyche viii. p. 209. t. 5. f. 4, 1. juv. (1898) ; Durand, ibid. ix. p. 87 (1900 (N. Toronto, one-
brooded).
Pajiilio troilus, Goda.tt, Enc. MM. ix. p. 60. n. 97 (1819) (cit. Drury excl. ; " Jamaica " false) ;
Charpent., in Espcr, Ausl. Schmett., Appeml. p. 5 (1830) ; Boisd. & Lee, Uist. Gin. Lip. Amir.
( 596 )
Sept. p. 26. t. 10. fig. 1-4 (1833) (larva, pupa, ^ ; Georgia ; Virginia) ; Boisd., Spec. Gen. Lep.i.
p. 334. n. 176 (1830) ; Harris, Enlnm. i. p. 61 (1840) (larva solitary, covers leaf with coating
of silk and binds up the sides to form a kind of trough) ; Doubl., List Lep. Ins. Bill. Mux. i.
p. 15(184:)) ; id, Westw. & Hew., G™. Diiini. Lep. i. p. 13. n. 85 (1846); Gray, Cat. Lep.
/«.«. Bril. Mim. i. Pap. p. 23. n. 106 (1852) ; id.. Lint Lep. Inn. Brit. Mus. i. Pap. p. 31. n. 113
(1856); Moni-tr., Emiiii. Corp. Anim. Uiis. Petrop., Lep. i. p. 2. n. 28 (1857) ; D'Urb., Cnnad.
Xat. GenL ii. p. 312. fig. a. b., t. 4. fig. 1 (1857) ; Gosse, Lett. Alahnma p. 78 (1859) ; Newm.,
Prnc. Eiil. Sor. Philiid. i. p. 26 (1861) (N. Jersey ; on sassafras) ; Morris, Si/ii. Lep. N. Aiiier.
p. 5. n. 0 (1862) ; Harr., ed. Flint, Ins. Inj. Vecj. p. 266 (1862) (life history) ; Reak., Proe. Enl.
Sor. Pliilatl. ii. p. 135. n. 1 (1863) (" Chiapas " false) ; Weidem., ibid. ii. p. 148 (1863) (Canada
to Mexico ; " West Indies" false) ; Lintn., Hid. iii. p. 51 (1864) (Eastern X. York, very rare) ;
Kirkp., ibid. iii. p. 329 (1864) (Cleveland, Ohio, common) ; Felder, Verli. Zn.d. Bnl. Ges.
^yien xlv. p. 316. n. 363 (1864) ; Saund., Cauad. Eiit. i. 73 (1868) (early stages ; London ;
imago appearing in June) ; Riley, Anier. Eiitnm. i. p. 60 (1868) ; Pack., Guide St'idij Ins. p. ^47
(1868) ; Bethuue, Cannd. Ent. ii. p. 8 (1869) (Toronto, July) ; Harris, Eid. Vorresp. p. 271.
t. 2. fig. 1, t. 4. fig. 16 (1869) ; Scudd , Cnnad. Ent. iv. p. 84 (1872) (Abbot's MSS.) : Saund.,
ibid. vi. p. 140 (1874) (Essex Co.) ; Bean, Enl. Mo. Mag. x. p. 248 (1874) (Galena, 111., scarce,
late Aug.) ; Pagenst., Verb. Nat. Med. Ver. Ueidelb. (2). i. p. 89 (1874) ; Edw., Trans. Amer.
Ent. Sac. vi. p. 10. n. 14 (1877) (Atlantic States ; Mississippi valley ; = ilioneus) ; Dury,
Cincinnati Soc. Nal. Hist. i. p. 12 (1878) (Cine, common) ; Gerh., ifaero-Lep. N. Amer. p. 25.
n. 455 (1878) ; Streck., Butt. Moths N. Amer. p. 72. n. 18 (1878) ; French. Trans. Dept. Aijric
Illin.xv.p. 138 (1878); Oberth., Et. d'Ent. iv. p. 69. n. 198 (1880) ; Middl., Trans. Dept.
Aciric. Illin. xviii. p. 74 (1881) ; Skinn., Proc. Ac. N. Sci. Philad. p. 239 (1882) (scent-organ of
larva); Auriv., A'. Se. Vet. Ak. llandl. xix. 5. p. 12. n. 6 (1882) (Recensio critica) ; Edw.,
Canad. Ent. xvi. p. 115 (1884) (habits of larva) ; French, Butt. East. t"..S'.p.93 (1884) ; Gruber,
Papilio iv. p. 87. t. 2. f. 12-15 (1884) (transf.) ; Saund., Kept. Ent. Soc. Ontario xv. p. 30 (1885)
(Point Pelee, L. Erie) ; Mayn., Bult. X. Ene/I. p. 49. n. 68. t. 7. fig. 68. 68a. ^ (1^*6) ' M''-
Nat. Sicil. V. p. 243 (1886) (Monaco !) ; Riley, Insect- Life i. p. 161 (1888) (parasite : Tror/us
exesorius) ; Skin. & Aaron, Canad. Enl. xxi. p. 127 (1889) (Philadelphia, common) ; Edw.,
Bull. U.S. Nat. Mas. xxxv. p. 10 (1889) (literat. on metam. ; cit. Gundl. ad speciem
P. poli/xenes dictam referenda); Pack., Fifib Kept. U.S. Ent. Comm. p. 650 (1890) (early
stages) ; id.. I.e. p. 063. 669. 909 (1890) (food-plants) ; Riley, Insect Life iii. p. 462 (1890)
(parasite : Pintpla notanda) ; Mayn., .l/«(i. X Amer. Butt. p. 10. n. 13. fig. 8. d (1891) ; Staley,
Canad. Ent. xxiv. p. 204 (1892) (Marshall, Missouri, iv.— x. not very common) ; Haase,
Untersuch. Mimirri/ i. p. 91 (1893) ; Skinn., Enl. Neit-s iv. p. 82 (1893) (N. Carolina) ; Meeske,
ibid. iv. p. 117 (1893) (Long L, common) ; Jones, ibid. iv. p. 190 (1893) (Richmond Co.,N.C.) ;
Davis, Journ. N. York Ent. Soc. i. p. 47 (1893) (Staten I., N.Y., May to Sept.) ; Riley, Insect
Life V. p. 207 (1893) (Falls Church, Va., larva abundant, November) ; Beutenm., Bull. Amer.
.l/«.f. N. H. V. p. 242 (1893) (N. York ; descr. of 1., p., i.) ; White, Enl. News v. p. 175 (1894)
(Brooklyn); Wi'eed. Psyche viii. p. 130. n. .38 (1894) (N.E. Miss.); Osburn, &/^ News vi.
p. 282. n. 47 (1895) (Tennessee, common, vii. to ix., two broods) ; Longl., ibid. vi. p. 314 (1895)
(Chicago); Eimer, .4rti. Vcrwandtsch. Sehmell. ii. p. 143. t. 8. fig. 2. ^ (1895) (•' ne.arest to
Pap. asterias") ; Clevel., Ent. News vii. p. 73 (1896) (Oneonta, N.Y.) ; Fiske, ibid. vii. p. 241
(1896) (Webster, N.H., not so common as turnus, June, July, second brood rare in Aug.) ;
Bubaa, ibid. viii. p. 98 (1897) (Cleveland, Ohio, May 2 and 3) ; Daze', Bdl Biff do Soc
A'. Sc. V. p. 107. n. 4 (1897) (Buffalo); Christ, Mitt. Schwei:. Enl. Ges. ix. p. 273 (1897);
Rowley, Enl. News ix. p. 37 (1898) (Louisiana, Mo., larva making silken c.ise liy drawing the
edges of the leaf together) ; Holland, Bull. Book p. 315, n. 18. t. 2. fig. 18. 19. 20. larva, t. 6.
fig. 5. 6. 7. pupa, t. 41. fig. 5. J (1899) ; Denton, .Moths Bull. U.S. p. 351. fig. larva and pupa,
plate ? (1898-190(1) ; Walk., liepi. Ent.. Soc. Ontario xxxii. p. 85 (1902) (Point Pelee,
Leamington) ; Beutenm., Bull. N. Vorlc Cit;/ p. 5. n. 3. fig. ? (1902) ; Dyar, Bull. U.S. Nat.
Mas. Iii. p. 3. n. 13 (1902) (Atlantic States ; N.W. Territory) ; Heink. Enl. News xiv. p. .335
(1903) (Meramec Highlands, St. Louis Co.. April 12) ; Franck. Enl. News xv. p. Ill (1904)
(aberration) ; id.. I.e. xvi. p. 91 (1905) (Passaic, N. Jersey, aberration).
Papilio Irolius (!), Edwards, Pajiilio ii. p. 76 (1882).
Papilio troilus var. ilioneus^ Haase, Untersuch. Mimicry i. p. 91 (1893).
Pterums (!) troilus, Kirby, in Allen's Nat. Libr., Lep. Bull. ii. p. 289 (1896) ; id., in Hiibn., Samnd.
Exot. Schmett. ed. ii. p. 100. t. 309. fig. 1. 2 (190—?).
Ptcrurus ilioneus, Kirby, in Allen's A'(//. Libr., I.e.
Papilio troilus yar.radialns Strecker, Lep. libop. Ilet. Suppl. iii. p. 17 (1900) (Washington, D.C. ;
Allegheny Co., Pa.).
(597 )
(J ? . Snbmarginal spots of both wiugs smaller than in sontherQ specimens,
those of hiadwiug more or less bluish. We have received from the American
Ent. Co. (Mr. G. Franck, who has shortly described it in 1905, I.e.) a curious
specimen which deserves special description, as it corresponds in pattern to the
ab. calverleyi of P. polijxenes aste/'lu.s. It was captured in June at W'estfiud, New
Jersej'. Upperside. — Forewing : snbmarginal sjjots enlarged, extended to edge
of wing, where they are conflaent; costal edge thinly creamy white at apex ; distal
margin slightly nndnlate. Hindwing : orange costal spot small, discal Imiid
olive-bnfF, rather well defined and narrow, not entering cell ; snbmarginal spots
extended to margin, bluish, fringe entirely creamy white ; a yellow anal sub-
marginal streak, a yellow marginal spot M' — M-, and another behind M^ ; dentition
of wing feeble ; the colour of the snbmarginal spots pervading the tail, except a
black central streak. On tiiulerside the marginal bands as above, but the spots
composing that of the hindwing a little more distinctly separate and all orange
mesially ; the orange discal spots all present, inclusive of spot R' — M' ; tail black,
except at edges. The specimen has also a very peculiar neuration. Forewing :
costal vein forked before middle of wing, the fork being on a level witli the point of
origin of SC, the short additional (anterior) branch extending a little beyond apex
of cell : SC before angle of cell ; M- forked in middle, the two branches remaining
separate in the left wing, while in the right wing the branches unite again before
reaching the edge of the wing ; on both wings there is an additional discal spot
between the two branches, and on the left wing also an additional spot at the
margin ; sjjur of median nervure continued distad for one-third the way to distal
margin — this additional vein being the so-called first submedian, of which the spur
of Papilioiiidae is the remnant. On the hindwing M- forked on the left wing
beyond the orange discal spots of the underside, on the right wing before these
spots, there being on the latter wing an additional discal spot and also a spot at
the margin.
llab. Georgia northward to Canada, westward to Texas and the Mississippi
jilains, in the north-west as far as the North-West Territory of Canada.
In the Tring Museum 70 6i, 50 ? ? and some larvae from : Ilayleigh, N.
Carolina (Brimley) ; Nelson Co., Virginia (Wirt Robinson) ; Jefferson Co., Kentucky
(Troxler); Nashville, Tennessee (Osburn) ; Makauda, lUinciis (Snyder) ; Texas.
b. r. troilus texanits Ehrm. (lOtH)).
rnpiliii troilus var. te.raims Ehrmann, C'aiiad. Ent. xxxii. p. .348 (ItlOO) (" Houston, Texas").
Ptipilhi troilus te.vanus, Dyar, I.e. p. .S. n. ]3a (190:2) ; Skinn., Ejit. Xews xiv. p. 275 (190.S)
(C'hokoloskee, Fla.).
Papilio trnitiis, Laurent, ibiil. p. 296 (1903) (Miami, Fla., common).
cJ ? . Submarginal spots of both wings large. On underside rather often a
pale subbasal baud on hindwing or on both wings, and a streak behind SM-.
IM. Florida.
Described by Ehrmann from a single specimen said to be from " Houston,
Texas." The Texas specimens which we have seen agree much better with the
previous form than with Florida individuals. The locality given by Mr. Ehrmann
is perhaps erroneous.
In the Tring Museum 14 c?c?, 2 ¥ ? , from : Chokoloskee, June 1903 ; Titnsville,
August 1894 ; Florida, August— September 1895 (Pridday).
( 598 )
86. Papilio palamedes Drury (1770).
Piip'ilin Equrs Arliiviix jialameiks Drury, JUustr. E.O't. Ins. i. p. 37. t. ID. fig. 1. 2. & Index (1770)
(Carolina).
S ? . Antenua tawnw Tibiae and tarsi pale greenisli, as in P. fkoas and allies.
In neuratiou and genitalia similar to 1'. troiliis.
Discal band of hindwing, above, xisnally distal of cell, often touching cell, many
sjiecimens bearing a small pale yellow cell-spot. Cell-bar of forewing sometimes
vestigial on uj)perside ; yellow subbasal band of underside of hindwing often con-
tinued across cell of forewing.
Genitalia : <?. Tenth tergite longer and narrower than in P. troilus ; harpe as
in that S])ecies, but the ventral tooth shifted towards the apex, standing close to the
apical process. ?. Mesial process standing at vaginal orifice sinuate apically,
Bon-dentate, lateral lobe less strongly dentate than in P. troiliiis, lateral ridge non-
dentate, but the edge irregular. Bristles on inner snrfiice of anal segment as in
P. troilus.
For early stages see Boisd. & Lee. (1833).
Hab. Southern Atlantic district of the Nearctic Region, extending into
Mexico.
Two subspecies.
a. P. jialamedes palamedes Drury (1770).
Seba, Thesaur. iv. p. 51. t. 43. fig. 3. 4 (1764).
PapiJin Eques Achirus palamedes Drury, I.e. ; Cramer, Pap. Exot. i. p. 14C. t. 03. fig. A. B (177G)
(N. York) ; Goeze, Ent. Beijir. iii. 1. p. 73. n. 11. (1779) {parlim) ; Jung, Alphah. Verz. p. 77
(17',l-2) (= cUalcus).
Papilio Eqiies Achinis clialcas Fabricius, Si/st. Ent. p. 453. n. 44 (1775) ; Goeze, Eiil. Bei/lr. iii. 1.
p. 73. n. 10 (1779) ; Fabr., Si/sf. Enl. iii. 1. p. 31. n. 90 (1793).
Papilio Eqiies AcJi i rns Jlaroiitaculatus Goeze, Ent. Beijtr. iii. 1. p. 87. n. 72 (1770) (nom. pro Sebae
t. 43. fig. 3. 4).
Papilio Eques Achirus clialncs Fabricius, Sjiec. Ins. ii. p. 18. n. 70 (1781) ; id., Mnnl. Ins. ii. p. 9.
D. 80 (1787); JaU. & Herbst, Nat. Schmett. iii. p. 1.39. n. 94. t. 42. fig. 1. 2 (1788) {=■- jmlamedes
Drury) ; Gmelin, Syst. Xat. i. 5. p. 2230. n. 315 (1790).
Papilio Eques Achirus calchas Esper, Aiml. Schmett. p. 229. n. 10(1. t. 5G. fig. 3 (1798) (nom. nov.
loco chalcas).
Papilio oiM«.v, Godart, Eiic. MM. ix. p. 59. n. 92 (1819) ; Boisd. & Lee, Fliaf. Gen. Lip. Amh.
Sejil. p. 17. t. 5. fig. 1—4 (1833) (larva, pupa. (J) ; Boisd,, Spec. Gen. Lep. i. p. 337. n. 178
(1836): Doubl., List Lep. Ins. Brit. JIus. i. p. 16 (1845) (E. Florida; Georgia); id..Westw. & Hew.,
Gen. Diurn. Lep. i. p. 13. n. 91 (1846) (U.S. ; Mexico ; " Jamaica'' false) ; Gosse. Letters from
.■!M-anm p. 169. 272 (1859).
Papilio palamedes, Drury, ed. West;!'., Illuatr. E.rnt. Ent. i. p. 36. t. 19. fig. 1. 2 (18:i7) ; Poey, .Mem.
Hist. Xat. Cuha i. p. 197. n. 12 (1851) (Cuba? ; palamedes has priority) ; Gray. Cat. Lep. Ins.
Brit. Mm. i. Pap. p. 25. n. 113 (1852) ; id.. List Lep. Ins. Brit. Mus. i. Pap. p. 33 n. 120 (1856)
(E. Florida ; Georgia); Lucas, in Sagra, Ilisff. Cuba vii. p. 203 (18r)7) (" Cuba" false); Weidem.,
Proc. Ent. Sue. Pliilad. ii. p. 147 (18C3) (U.S. : " West Ind." false) ; Felder, Verh. Zool. But.
Ges. Wien xiv. p. 316. n. 364 (1864) ; Jaeg., Life N. Am. Ins. p. 210 (18C1) ; Butler, Cat.
Diurn. Lep. flescr. Fabr. p. 260. n. 53 (18C9) ; Kirby, Cat. Diurn. Lep. p. 543. n. 166 (1871) ;
Edw., Trans. Amer. Ent. Soc. vi. p. 11. n. 20 (1877) (Florida to Virginia; Gulf States;
= calchas) ; Gerh., Macro-Lep. N. Amer. p. 25. n. 439 (1878) ; Streck., Butt. Moths N. Amer.
p. 73. n. 19 (1878) (Virginia southward, south-west to Louisiana) ; Oberth., Et. d'Ent. ir.
p. 69. n. 199 (1880) (Florida) ; Edw., Canad. Ent. xiii. p. 119 (1881) (life hist.) ; id.. I.e. xvi.
p. 115 (1884) (habits of Larva) ; Aaron, Papilio iv. p. 172 (1884) (S. Texas) ; Edw., I.e. xviii.
p. 15 (1886) (Glencoe, Nebraska) ; French, Butt. East. U.S. p. 95 (1886) ; Edw., Bull. U.S.
Nat. .Mas. x.kxv. p. 11 (1889) (liter, on metam.) ; Skinn., Ent. A'eu-s i. p. 110 (1890) (Phila-
'V- delihia, one fpecimpn); Mayn.. ^fan. N. Amer. Butt. p. 10. n. 14. fig. 8. a (1891); Jones, Ent.
Kius iv. p. 1£0 (181 3 (EichmoLi. Co., N.C.) : Haasc, Unlenuch. Mimicry i. p. 90 (1803) (■' the
( 599 )
Palamades Group shows close affinities with the Dauiitis Group") ; Eimer, Arlb. Vcrivauiludi.
Schmelt. ii. p. 145. t. 8. fig. 3. jj, fig. 4. ? (1895) ("closely related to aalci-iokks, axlei-iax and
hrevimuda on one side and to halrdi $ on the other") ; id., Orthogcn. p. 34. fig. 13 (1897) ;
Christ, Mill. Sfhwei::. Eiil. Gr.i ix. p. 270 (1897) ("near relative of zolk-ann") ; Holland, Ball.
Book p. 315. n. 19. t. 42. fig. 1. ? (1899) ; Dent., Mollis Bull. U.S. p. 353. 354. fig. (1898—
1900) ; Comst., Eiit. News .xiii. p. 75 & 77 (1902) (L. Josephine, Fla., quite common, Feb. 20
to March 25) ; Dyar, Bull. U.S. Nat. Mus. Hi. p. 3. n. 12 (1902) (S. Atl. States).
Papilio chalcas, Morris, Syii. Lep. N. A met: p. 7. n. 9 (18C2).
Pterurus palamedes^ Kirby, in Hiibn., Snminl. Exot. Schmelt. ed. ii. p. 101. t. 115. fig. 3. 4 (190- ?)
t? ? . The discal spots on the underside of the bindwing on the whole Itss
extended orange in Florida specimens than iu individnals from other places, and the
orange portion of the spot R^ — M^ often reduced.
IJab. Florida to Philadelphia, westwards to the Mississipi plains.
In the Tring Museum 34 <?(?, 14 ? ?, from : Te.xas ; New Orleans, April H)02
(A. Hall) ; Titnsville, Florida, August— September 1894 and 1895.
b. P. pala?nedes leontis subsp. nov.
J ? . A small form. Upperside. — Forewing : discal spots smaller than iu the
previous, in the female being smaller than the submarginal ones ; streak in front of
SC absent or vestigial ; cell-bar absent or faintly ve.stigial. Hindwiug : discal
band narrower than in P. p. palamedes, the spots R^ — M^ better defined.
Underside. — Forewing : cell-bar smaller than in the preceding form, often
reduced to two small dots ; discal s])ots also smaller ; no subbasal baud across cell.
Hiudwing : discal baud narrower and on the whole more extended orange, and
the glossy blue spots larger, than in P.j). palamedes.
Hah. Monterey, Mexico.
In the Tring Museum G <?<?, 1 ?.
VIII. Ancliisiades Group.
Pronotum, or underside of thorax, or both, spotted with red or orange ; abdomen
black, at base with a lateral dot or short streak which is of the same colour as the
thoracical spots, no yellow or buff lateral line on abdomen. Cell of forewing
beneath not streaked with yellow. Hiudwing without blue spots.
Nenration : SC- of forewing very much nearer SC than SC^ ; D- longer than
D^ ; lower angle of cell very obtuse ; PC of hiudwing long and gradually carved.
Genitalia : cJ. Tenth tergite spatulate ; sternite laterally in middle incrassate to
form a small double ridge or two teeth, of which the proximal one is hairy. Harpe
long and (with the excej)tion of P. bjppason) fiat, being apically produced into a
long and nearly straight point. ?. Edge of vaginal orifice proximally raised into
a prominent curved process, which is bicarinatc or channelled iu front and deeply
grooved on posterior side ; behind the orifice a membranous tubercle clothed with
minute hairs ; laterally a spatulate and dentate lobe or a pointed process, and farther
proximad a ridge which is shell-like laterally.
The larvae are gregarious, feediug chiefly on Citrus.
A. Sexes dissimilar ; SC- of hindwiug much more proximal than
M^ ; a red basal patch on underside of bindwing behind cell ;
spots of pronotum rufous red, those on underside of thorax
and at base of abdomen pale buff Species No. 87.
( eoo )
B. S-exes similar, with long spatulate tail ; a cream)' white band
across disc (No. 68) ; or tliis band vestigial, in the latter
case the marginal spots of both wings large (No. 89) . Species No. 88. 89.
C. Sexes similar ; no sharply marked band across forewing ;
marginal spots of forewing small or absent, or the tail absent
a. Hiudwiug without tail, marginal spots all large, cream-
colour Species No. '.lO.
Hindwing with or without tail, marginal spots small, e.\cej)t
the tirst spot on the hindwing, which is often somewhat
enlarged ......... b.
b. On upperside of hindwing two parallel rows of spots, well
separated from one another, the proximal or the distal
row often missing ....... c.
The submarginal spots of hindwing close to the discal ones,
touching them or being merged together with them,
the snbmargiual spots R' — M- enlarged ... d.
c. Submarginal spots of hindwing above cream-colour in S ;
marginal sjiots of hindwing [large in ?, almost
entirely orange-red above and below .... Species No. 93.
Submarginal spots of hindwing red in both sexe.? ; mar-
ginal spots of ? moderately large, more white than
orange-red below Species No. 92.
Submarginal spots absent or vestigial, only one row on
npperside, consisting of a few small spots . . . Species No. 94.
(I. Forewing above with yellowish white patch from hind-
margin forward to M-, vestigial Iielow . . . Species No. 91.
No such patch ......... e.
e. Hiudwiug with sliort acute tail ; harpe uon-dentatc . . f.
Hindwing without tail, or tooth R^ projecting, broad ;
harpe dentate ........ Species No. 9.').
/. Forewing below with white jiatch across apex of cell . . Species No. 97.
Forewing below witliout white patch across apex of cell . Species No. 90.
87. Papilio hyppason Cram. (177.)).
(J. Papilio EqiieH Trojaiins lii/ppasO)i Cramer, Pap. Eu-at. i. p. 4(5. t. 20. fig. E {\~b) (Surinam) ;
Goeze, Ent. Beijir. iii. 1. p. 37. note (1779) (" var. of /'. aencaii ") ; Esper, Aiisl. Schindt. p. 03.
n. 28 (1788).
$ . Papilio Eijiies Tiojanux umasis Cramer, l.r. iii. p. 139. t. •2G9. fig. A. B (1780) (Surinam) ; Stol.,
ibid, Siippl. p. 1. t. 1. fig. 1a. Id (1787) (larva, pupa ; this species ? or 1'. andrujeiis V).
(^ . Papilio Eqite& Trojaiiits hippnaon, Esper, I.e. t. 10. fig. 2 (1788).
9 . Papilio Eques Trojanun amosix, id., I.e. p. 04. n. 29. t. 70. fig. 3 (1788).
Papilio Eques Trojauus aeneax, Fabricius, Spec. Ins. ii. p. 8. n. 32 (1781) (jiarlim).
Papilio Eques Trojaims aencas y) P. lii//ijiasoii, Gmelin, Si/st. Nal. i. 5. p. 2233. sub n. 10 (1790).
Papilio Eques Tmjaims ilimas Fabricius, Eiil. iSi/sl, iii. 1. p. 10. n. 47 (1793) (paiiim).
Prinreps (lomimiiis hyjqiuson, Hiibner, Samml. E.rol. Sclntiell. i. t. 124 (1800 — ? ).
Priamitks amosis, id., Vcrz. bek. Sclimetl. p. 87. n. 905 (1818 ?).
PriaiHides hippasoii, id., I.e. p. 87. n. 900 (1818?) (parliiii).
cJ ?. Papilio amosis, Godart, Eur. MM. ix. p. 35. n. 29 (1819) (Guyane).
1^. Papilio Iiippasoii, id., I.e. ix. p. 35. n. 30 (1819) (Guyane ; "euristeiis " e.\cl.).
(J $ . Papilio liippasoti, Boisduval, Spec. Gtn,Lep. i. p. 281. n. 100 (1830) (amosis = J of hippasoii;
Surinam) ; Doubl., List Lcp. Lis. Brit. Mas. i. p. 11 (184.''i) (Brazil) : id., Westw. & Hew.,
Gen. Diurii. Lep. i. p. 19. n. 214 (1847) (Guiana); Erich.i., in Schomb., /■". /•'. Brit. Guiana
( 601 )
p. 693 (1848) (? probably = nw»/s/s Cram.); Gray, Cal. Lei>. Inn. Brit. Mux. i. Pap. p. 58.
n. 268. t. 10. fig. 3. ? (1852) (Demerara ; Surinam ; Pard) ; Wall., Tmm. Ent. Sue. Loud. (2).
ii. p. 256 (1854) (Pari ; forest) ; Gray, List Lep. /«.s. Brit. Mm. i. Fiip. p. 70. n. 284 (185t;)
(Demerara; Surinam; Para); Me'nc'tr., Eiium. Corp. Anim. Mus. Petrop , Lep. i. p. (J8.
n. 1127 (1857) (Guiana) ; Bates, Trans. Ent. Soc. Loud. (2). v. p. 337 (18G1) (Guiana ; Pard) ;
Felder, Verh. Zool. Bol. Gen. Wieii xiv. p. 208. n. 131 (I8G4) (Surinam ; Pard) ; Kirby, C<it.
Diiirn. Lep. p. 523. n. 34 (1871) (Guiana); Oberth., Et. d'Ent. iv. p. 80. n. 203 (18S0)
(Surinam) ; Staud., Exot. Tmjf. i. p. 13 (1884) (Surinam ; Amazons ; Peru) ; Haase, Unl-.rsneli.
3Hm>er<j i. p. 'J9. t. 10. fig. 73. ? (1893) (Surinam ; Para on plate ; figure not correct).
Paj)ilio hipjiaso?/, Cram. Local var. jxiraeiinis, Bates, Jourii. Entom. i. p. 225. n. 9 (1862) (Para") ;
Oberth., Et. d'Ent. Iv. p. 110. n. 203 (1880) (Pard).
Papilio hippason var. a. P. hipp. var. paraensia, Kirby, Cat. Diurn. Lep. p. 523. sub n. 34 (1871)
(Pard).
Pajtilin hyppason, Moschler, Verh. Zool. Bui. Ges. ^Yiell xxvi. p. 295 (1870) ( ? , Surinam).
Papilio amasis (!), Burmeister, Deser. Pfp. Aryent. v. Lqi. p. 4. sub n. 4 (1879) (StoU's t. 1 fig. 1.
" is larva of P. pulijeaon ").
(J. Papilio hippasonides Grose-Smith, Rhop. E.cot. iii. Puji. t. 22. fig. 3. 4 (1902) (Yungas, Bolivia).
lilwbalus amosis, Kirby, in Hiibn., Samml. Exot. Sohmett. ed. ii. p. 92. t. 125. fig. 3. 4 (190- ?) ( 'not
the ? of P. hippason Cram.," errore).
"While Bates, Felder, Kirby (1871) and Standinger, deceived b}' the iiatterii
cil the iusect, put P. hyppason in the Ariarathes Group, Haase placed it with
P. pharnaces and anchisiades. That is indeed the correct position, as is borne out
by the structure of the imago. In the new edition of Hilbner the species is placed
by Kirby in Ithobalus, the type of which is an Aristolochia-Swallowtail. The
insect has many peculiarities sejjarating it from the other members of the
Anchisiades Group. It is individually rather variable, but does not appear to have
split up into geographical forms. The sj)ecies can easily be recognised by the
presence of a large red spot ou the imderside of the hindwing behind cell close
to base. Kirby, I.e., makes the erroneous statement that the marginal spots of the
hindwing of P. ki/ppason are red.
(J ¥ . Antennae in male a little extending beyond apex of cell of forewiiig, in
female not reaching apex of cell ; club gradual and long, end-segment conical,
basally broader than in the allied species. Palpus black. Spots of breast grey
or buff, those on pronotum rufous red. Scales of upperside of forewing obtusely
((?) or more acutely (?) tridentate, the white ones partly bide ntate, those of the
upper layer of the last patch in male entire ; on underside the scales strongly
bidentate, those of the upper layer entire from the middle of the hindmargiu
forward, especially the white ones. The spots of tbe hindwing consist of rufous red
and white scales ; the red scales narrow, long, entire or feebly sinuate, opalescent,
many being yellow at base ; some of the dark scales at the edge of these spots or
within them bright metallic blue under the lens.
Nenration : M' and M- of forewing closer together tha\i R-' and M' ; PC of
hindwing evenly curved, SC- more proximal than in the allied species, standing
closer to base than to R'.
Genitalia : <?. Tenth tergite spatulate, beneath carinate, except at apex ; tenth
sternite laterally with two teeth projecting upwards, the second tooth pointed,
the first mucli shorter, hairy ; clasper short, broad, rounded ; harpe broad, the
ventral margin raised and distally produced into a tapering process ; beyond this
process the har])e narrowed to a slightly spatulate lobe, which is curved vcntrad
and bears a variable number of teeth at the rounded apical edge. ? Edge of
vaginal aperture raised, denticulate laterally, bisinuate in front, the mesial frontal
lobe triangular ; in front of the orifice a high ridge, strongly chitinised ; between
( 602 )
this ridge and the orifice on each side of the ktter a rather long and slender
pointed process.
Each sex dichromatic, with iutergradation. These forms are : —
a. J-f. hjppason Cram. I.e.; ///j/paso/iuh's Gvose-Simth, I.e. Baud of fore-
wing broader than the interspace between the band and cell, its inner edge crossing
vein SW.
b'. (?-f. ptilion nov.— Band of forewing narrow, farther away from cell than
in the preceding form, and usually continued farther forward, the spots separate ;
hindwing above with fonr or five red spots, seldom less. Type of name from
Iquitos.
c'. ?-f. amosis Cramer, I.e. Forewing with hardly a trace of a white
patch.
d'. ¥ -f. paraeiisis Bates, I.e. Forewing with large white or huffish patch.
variable in extent, often entering cell.
Early stages not known with certainty. Stoll's figures may belong here, though
Burmeister refers them to P. androgens (= pohjcaoii).
Hah. The Guianas ; Amazons ; Peru ; Bolivia.
In the Tring Museum 2o c?cJ, 13 5 ?, from : E. Demerara, British Gniana;
Paramaribo, Surinam ; Parii, (Stuart, Bates) ; E. Jurua ; Iquitos ; Salinas, E. Beni,
July 1896 (Stuart); B. Songo to B. Suapi, Bolivia, 1100 m., March— June 1896
(Garlepp) ; Province Sarra, S. ('rnz de la Sierra, February — April 1904 (J.
Steinbach.)
88. Papilio pelaus Fabr. (1775).
Papilio Eques Trojanus pelaus Fabricius, Syst. Enl. p. 444. n. 9 (1775) (" ludia," Diury).
Papilio ornofagus Weidemeyer, I.e. p. 147 (1863) (sub syn.).
In Jones's unpublished drawings P. pelaus is figured from Drury's collection.
As Fabricius described the insect from this collection, Jones's fignre may be taken
as representing the type of pelaus. The figure agrees best with specimens from
Jamaica, from which island Drnry possessed many insects. Westwood's figure
(1842) was apparently taken from the same specimen.
(??. Sexes similar, but the markings rather larger and the submarginal sjiots
of the upperside usually more numerous in female than in male, and larger beneath
than above. Forewing with a creamy band from costal margin to hinder angle
standing just outside cell or entering it. A row of rufous red submarginal
spots on hindwing parallel to distal margin, the upper spots absent from male ou
upperside, the last spot more or less creamy ; a discal row of small creamy sjiots
on underside, sometimes rufous, rarely absent, in female occasionally marked also
on upperside.
Genitalia : i. Tenth tergite spatulate ; harpe of the same type as in F.
a«e/«"s/«f/«s, long, non-dentate, aj)ex ])roduced into an acute process. ?. Proximal
ridge of vaginal cavity membranaceous in middle and here hardly raised, laterally
dilated, rounded ; lateral dentate lobe spatulate, rounded at apex ; bearing two or
three teeth ; the process standing at orifice short, broad, widened laterad, triangular ;
the mesial ridge extending from this process forward bicarinate, the edges being
raised.
Larva described by Cockerell (1893).
Hab. Jamaica ; Cuba ; Haiti : Porto Eico.
( 603 )
a. P. pelaus pelaus Fabr. (1775).
Papilio Eques Trojanus pelaus Fabricius, I.e. ; Uoeze, Ent. Beijlr. iii. 1. p. 41. n. 6 (1779) ; Fabr.
Spec. Ins. ii. p. 4. n, 12 (1781) ; id., Mant. Ins. ii. p. 2. n. 12 (1787) ; id., Ent. Sijsl. iii. 1. p. 5.
n. 15 (1793) (cit. Cram. excl.).
Papilio Eques TrojaMts pelem (!), Gmelin, Sijsl. Nal. i. 5. p. 2228. n. 279 (1790) (partim).
Heraclides pelaus, Htibner, Verz. bek. Svlimell. p. 83. n. 853 (1818 ?) (partim) ; Godart, Ene. MM. ix.
p. 61. n. 99 (1819) (de.scr. from Fabric.) ; Boisd., Spec. Ghi. Lip. i. p. 367 n. 210 (183C)(descr.
from Fabric); Westw., Arc. Ent. i. p. 107. t. 18. fig. 1. 2 (1842) ; Doubl., List Lep. Ins. Brit.
Mus. i. p. 17 (1845) (Jamaica) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 40. n. 202 (1852)
(Jamaica) ; id., List Lep. Ins. Brit. Miis. i. Pap. p. 55. n. 213 (1850) (.Jamaica) ; Herr.-Sch.,
Corresp. Bl. Zool. Min. Ver. Rerjensb. xvi. p. 141 (18G2) (Cuba) ; Weidem., Proc. Ent. Soc.
Philad. ii. p. m (imS) (partim) ; Hcrr.-Scb., I.e. xviii. p. 173. n. 9 (1864); Felder, Verh.
Zool. Bot. Ges. Wien xiv. p. 3(19. n. 284 (1864) (partim ; Cuba, Jamaica) ; But!., Cat. Diiirn.
Lep. (User. Fabric, p. 246. n. 42 (1809) (Jamaica) ; Kirby, Oit. Diurn. Lep. p. 542. d. 164(1871)
(partim) ; Dewitz, Stelt. Ent. Zeit. xxxviii. p. 234. n. 4 (1877) (Porto Rico) ; Bull., Proc. Zool.
See. Land. p. 481. n. 35 (1878) (Jamaica) ; Gundl., Papilio i. p. 113 (1881) (Cuba) ; id., Contr.
Ent. Cuba. p. 126 (1881) (imrtim ; Eastern Cuba, .Jamaica, Porto Rico) ; Moaclil., Abh.
Senkenb. Nat. Ges. xvi. p. 91. n. 4 (1891) (Porto Rico) ; Gundl., An. Hist. Nat. Madrid xx.
p. 114. n. 4 (1891) (Porto Rico) ; Haase, Untersueh. Mimicri/ i. p. 98 (1893) [partim ; Jamaica,
Cuba) ; Fox & Johns., Ent. News iv. p. 3 (1893) (Jamaica) ; Cocker., Psi/cbe vi. p. 4,50 (1893)
(full-grown larva, July); id., Jourit. Jamaica Inst. i. p. 179 (1893) (larva); Ehrm., Ent.
News vi. p. ,303 (1895) ( ? , Jamaica) ; Swain.s., Journ. N. York Ent. Soc. ix. p. 78 (1901) (larva) ;
Robins., ibid. xiv. p. 19 (1903) (Jamnica).
cJ ? . Band of forevring close to cell, in female always entering it, sometimes
also in male, especially often beneath, the last spot of the band nsually ill-defined.
On the hindwing there are in male three or more submarginal spots, the female
having six, the spots variable in size, larger and on an average more numerous in
Cuban specimens than in Jamaican individuals.
Ilab. Jamaica ; Cuba ; Porto Rico, this form ?
In the Triug Museum 9 S6, 5 ? ¥ , from : Bath, S. Thomas, Jamaica (Taylor) ;
Cuba (Gundlach).
b. P. pelaus imerius Godt. (1.819).
Papilio imerius Godnvt, Enc. MM. ix. p. 69. n. 121 (1819) (" Ind. orient.") ; BoisA., Spec. Gen.
Lep. i. p. 312. D. 150 (1830) (Haiti) ; Grimsh., Trans. Roy. Soc. Edinh. xxxix. 1. No. 1. p. 8
(1897) ( = "sctes Westw." err. det.).
Papilio auf/ias Menijtri^s, Bull. Moseou ii. p. 293. n. 3 (1832) (Haiti).
Papilio pelaus, Doubleday, West. & Hew., Gen. Diurn. Lep. i. p. 17. n. 179(1846) (partim ; Haiti) ;
Miniti., Eniim. Corp. Anim. Mus. Petrop., Lep. i. p. 4. n. 65 (1857) (Haiti) ; Weidem., Proc.
Ent. Soe. Philad. ii. p. 147 (1863) (partim) ; Felder, Verb. Zool. Bot. Ges. Wien xiv. p. 309
n. 284 (1864) (partim ; S. Domingo) ; Kirby, Cat. Diurn. Lep. p. 542. n. 164 (1871) (partim) ;
Oberth., Et. d'Enl. iv. p. 69. n. 203 (1880) (Haiti) ; Haase, Untersuch. Mimicry i. p 98 (1893)
(partim) ; Haiti).
Papilio pelaus var., Gray, List Lep. Ins. Brit. Mus. i. Pap. p. 55. sub n. 213 (1856) (Haiti).
c? ? . Band of forewing anteriorly narrower than in the preceding, more straight,
not narrowing behind, the last sjiot being proportionally larger than in the preceding
subspecies, spot SC-— SC^ reduced in cj or absent, the band not entering cell either
above or below. Submarginal spots of hindwing smaller.
Ilab. Haiti.
In the Tring Museum 2 c? c?.
89. Papilio oxynius Hubn. (1834?).
Laertias oxynius Hiibner, Samnd. Exot. Sclunett. iii. t. 5 (1834 ?) (Cuba).
Papilio augustus Boisduval, Spec. Gin. Lip. i. p. 358. n. 20O (1836) (Cuba) : Lucas, in Gu.T., nicl.
Pitt. Hist. Nat. vi- p. 50 (1838) (Cuba).
39
( 604 )
Papilio oxyn'ms, Doubleday, Westw. & Hew., Gen. Diurii. Le/). i. p. 17. n. 174 (1846); Poey,
Mem. R. Soc. Emu. Hahtim p. 230 (1846) ; Gray, Cal. Lep. Inn. Brit. .Uii^. i. Pup. p. 39. n. I'.t'j
(1852) ; id., List Lep. Ins. Brit. Miis. i. Pap. p. 55. n. 210 (1856) (Cuba) ; Weidem., Proc. E„t.
Soc. PhitwI. ii. p. 147 (1863) ; Felder, Verh. Z«,A. Bui. Ge-i. iVie:/ xiv. p. 30'.l. n. 285 (1864)
(Cuba) ; Herr.-Sch., Cnrreap. Bl. Ziiol. Miii. Ver. Rejinnili. p. 173. n. 6 (1804) (common) ; Kirby,
Cal. Ditini. Lep. p. 542. n. 163 (1871) (Cuba) ; Oberth., El. d'Eiil. iv. p. 69. n. 204 (1880) ;
Gundl., Papilio i. p. 113 (1881) (Cuba) ; id., Co„l,: E„l. Cuba. p. 127 (1881) ; id., Beil. E„l.
Zeit^chy. x.x.\T. p. 131 (1890) (descr. of larva) ; Honr., Silzber. Brrl. Eiil. Zeiticlir. xxxiii. 1889.
p. 8 (1890) (larva social, on Zaiil/i<ij-i/biiii) ; Riley, Insect Life in. p. 32 (1890) (larvae soi.ial
ace. to Gundlach) ; Haase, Unlersiirh. Mimienj i. p. 99 (1893).
Papilio (Laertias) oxi/niiis, Lucas, in Sagra, Hist. Cuba vii. p. 207 (1857).
Plenirus (!) o.ri/niii.'s, Kirby, in Hiibn., Samml. Exul. Sclunetl. ed. ii. p. 100. t. 459. fig. 1 2 (190-?):
c? ? . Similar to P. pelaua ; marginal spots of both wings larger ; band of
forewing more or less vestigial, in male nsnally absent from npperside.
Genitalia : cT. Harpe broad, denticulate.
Larva gregarious at night, resting together on the trunk of the tree of which
the leaves serve as food.
Hab. Cuba.
In the Tring Museum 4 c? c?, 1 ? .
90. Papilio epenetus Hew. (1861).
Papilio rpenelii.i Hewitson, Exot. Butt. ii. Paj). t. 5. fig. 14. 15. J (1861) (Cinchona, Ecuador);
Felder, Verl,. Zool. Hot. Ges. WIen xiv. p. 312. n. 319 (1864) (hab. ?) ; Kirby, a/^ Diiini. Lep.
p. 539. D. 146 (1871) ; Ilaase, Untnsiirh. Mimicry i. p. 99 (1893) ; Haensch, Berl Eiil. Zeilsclu:
xlviii. p. 151 (1903) (Balzapamba, W. Ecuador, larva on orange-trees in June, short descr. of
larva and pupa).
c? ? . Sexes similar, female a little paler than male. Forewing with a row of
huffish yellow jmtches on underside. Hindwing, on upperside, with or without some
red dots on disc ; on underside a complete row of red discal spots, and rather close
to it a row of bufiSsh yellow submarginal spots, of which the posterior ones are
somewhat larger than the anterior ones ; marginal spots large above and below,
huffish yellow.
Genitalia : S. Tenth tergite elongate, slightly sjiatulate. Clasper ventrally
somewhat emarginate before apex ; harpe of the same type as in /'. anchisiades,
ending in an acute process which is slightly curved towards the clasper ; proximally
of the process there is or is not a small marginal tooth. ? not dissected.
Larva and pupa described by Haensch, I.e., closely agreeing with those of
P. anchixiades.
llab. Western Ecuador.
In the Tring Museum 3 cJc?, 2 ? ?, and several larvae and pujjae, from:
Balsapamba (R. Haensch); Celica to Sapatillo, July 1600 (Simons).
In coll. Oberthiir from Chimbo and Balsapamba.
91. Papilio chiansiades Westw. (1872).
Papilio chiansiades Westwood, Trans. Ent. Soc. Lond. p. 101. t. 3. fig. 4. 5. (J (1872) (R. Topo,
Ecuador) ; Oberth., Et. d'Enl. iv. p. 116. n. 258"" (1880) (TefEc ; Pebas).
Papilio chinsiudes (!), Kiiby, Oil. Diiirn. Lep. p. 812. n. 353 (1877) ; Kirby, Trans. Ent. Soc. Lond.
p. 353 (1881) (Sar.iyacu) ; Stand., ICrol. Tai/f. p. 16. t. 11. ? (1884) (Amazons; Ecuador);
Haaae, UntersiicU. Mimii-ry i. p. 99 (1893) ; Michael, Iris vii. p. 213 (1894) (Sao Paulo de
01iven9a) ; Eiiner, Ortlioijeu. Sc/imell. p. 327 (1897) {ah/ulles group ! !).
cJ Forewing, above, with large yellowish white patch fmni hinder margin to
near M", <he patch being vestigial below. Hindwing, on npperside, with two
( 605 )
small discal and three larger sulimarginal spots, all red, often much shaded with
black, the last snbmarginal spot distal of the one before it ; on underside two
discal spots M' — SM- and a complete row of submargiual spots, those between
R^ and M- being the largest and partly white.
? and earl)' stages not known.
Hab. Eastern slopes of Ecuador aod Pern ; Upper Amazons.
In the Tring Blusenm 11 <?<? from : Pebas ; S. Paolo de Olivenr'a ; Iquitos ;
R. Cachyaco, affl. of R. Hnallaga (Stnart) ; Coca, Ecuador (R. Haeiisch); Archidona
(W. Goodfellow); R. Chuchuras, affl. of R. Palcazu, 320 m. (W. Hoffmanns);
La Union, R. Huacamayo, Carabaya, 2000 ft., January 190.5, wet season (G.
Ockenden).
02. Papilio pharnaces Doubl. (1846).
Papilii) 2>Jior!iaccs Doubleday, Ann. JJay. X II. xviii. p. 374 (184fi) ("America merid.") ; id.,
Westw. & Hew., Gen. Diurii. Lep. 1. p. 19. n. Slfi (1847) (" Bolivia ") ; Gray, Cat. Lip. hm.
Biit. Mils. i. Pap. p. 04. n. 282. t. 5. fig. 1. ^ (1852) (Guatemala?) ; id., List Lep. Ins. Bril.
iliis. i. Pap. p. 73. n. 298 (18.')6) (Jlexico) ; Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 312. n. 321
(18G4) (Mexico ; '"Guatemala") ; Kirby, Cat. Diurn. Lep. p. 539. n. 145 (1871) (Mexico) ;
Godm. & Salv., Biul. Centr. Amer., Lep. lUiap. ii. p. 231. n. 04 (1890) (Mexico : Oaxaoa,
Putla) ; iid., I.e. p. 730 (1901) (Guanajuato) ; Haase, Untersiich. Jlfimicn/ i. p. 99. t. 9. fig. 03
(1893).
cJ. Pvpilh phanostraius Godman & Salv., I.e. p. 232. d. 05 (1890) (Jalapa).
^. Papilio polycharmus iid., I.e. n. 00. t. 70. fig. 10 (1890) (Mazatlan).
S ? . Hindwing with two rows of spots, the distal row about halfway between
cell and distal margin, red, the pro-\imal spots variable in number on upperside.
Tail very variable in length ; in one of our West Mexican females, from Colima,
little more prominent than the other teeth, while it is long in another female
from the same place. We do not find any fairly constant difference between
specimens from East, West, and South Mexico.
Genitalia: c?. Tenth tergite slightly spatulate ; sternite laterally with obtuse
ridge which is transversely impressed, bearing hairs proximally. Harpe dentate at
apex dorsally and ventrally. ?. Vaginal armature as in P. aiichisiade.s ; the
curved process standing at the orifice shorter, the lateral dentate lobes with fewer
teeth.
JJab. East, South, and West Mexico.
In the Tring Museum 56 ,Si,2» ? ?, from: Orizaba, April and May 1890
(W. Schaus) ; Huatuxco ; Guadalajara, September — October 1889 (Dr. Buller) ;
Guadalajara, July, August and October, 1896 (W. Schaus); Oaxaca, 6000 ft,
June 1904 (A. Hallj ; Guernavaca, end of August 1904 (Dr. Gadow).
93. Papilio erostratus Westw. (1847).
Papilio e,-o.itratus \Yeatv!Ooi, Trans. Ent. Soc. Land. v. p. 30. t. 3. fig. 2. 2* (1847) (Guatemala);
Doubl., Westw. & Hew., Gen. Diurn. Lep. ii. p. 529 (1852) ; Boisd., Consid. Lep. Gnatem. p. 8
(1870) (Mexico ; Honduras ; rlietiis = J of erostratus teste Godman & S:ilvin) ; Kirby, Cat.
Diurn. Lep. p. 540. n. 153 (1871) (Guatemala) ; Butl. & Druce, Pruc. Zuol. Soc. Land. p. 305.
n. 378 (1874) (Costa Rica, teste van Patten) ; Oberth., El. d'Eut. iv. p. 80. n. 259 (1880)
(Guatemala ; Mexico) ; Godm. & Salv., Biol. Centr. Anter., Lep. Rhop. ii. p. 01. t. 09. fig. 9.
genit. (1890) (Brit. Honduras; Guatemala; ''Costa Rica" teste van Patten); Haase,
Untersueh. Mimirri/ i. p. 99. t. 9. fig. 00. J". 01, ? (1893) (Guatemala).
?. Pa/iilio rlietus Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 05. n. 288. t. 11. fig. 5 (1852) (Guate-
malaj ; id., List Lep. Ins. Brit. Mm. i. p. 75. n. 305 (1850); Felder, Verh. Zuol. But. (_!es.
Wien xiv. p. 299. n. 140 (1804) (Guatemala) ; Kirby, Cat. Diurn. Lep. p. 523. n. 38'"' (1871) ;
id., l.c. p. 810. n. 153 (1877) (= !? of erostratus).
Papilio Iieroslratus, Felder, Verli. Zool. Bol. Ges. Wienxiv. p. 310. n. 302 (1804) (Guatemala),
( 606 )
This aud che iirecediug insect (P. pkar/iace.i) occur together in "Western Mexico,
remaining perfectly distinct. There can be no doubt that the}' are sijcciticaily
distinct, though they are closely allied.
Sexes dissimilar.
J. Snbmargiual sjjots of npperside of hindwing cream-colonr, the last one
being nsnally red ; discal spots red or creamy red, never all present, always small,
sometimes all absent, most specimens bearing three small spots ; on underside botli
rows red. On tlio forcwiiig there are posteriorly some creamy submarginal spots,
which appear occasionally also on upperside. Fringe-spots of hindwiug creamy.
¥. Creamy marginal spots of forewing rather large, the submarginal ones of
underside often joined to them. Discal and submarginal spots of hindwing red
above and below, larger than in female of P. pharnnccs ; marginal spots red, also
large, the extreme fringe being white, last submarginal spot of upperside connected
anteriorly with the anal marginal one ; tail very slender, being also in male slenderer
than in F. pkarnaces.
Genitalia : S. Harpe angulate dorsally, bearing dorsally one or more teeth, and
vcntrally no teeth or only one.
Early stages not known.
llah. Guatemala aud British Honduras, the record from Costa Uica (van Patten)
being very doubtful.
In the Tring Museum, \) i S,\ ?, from : Cimlad de Guatemala (Hodrignez) ;
Palin, "W. Gnatemala, August— September 1004, :i;")Ua ft. (A. Hall) ; Guatemala
(Salvin).
94, Papilio rogeri Boisd. (1836).
Pajillio rugeri Boisduval, Spec. Gin^Lej). i. p. 278. n. 102 (1836) (Yucatan) , Doubl., Westw. & Hew.,
Geii. Dhmi. Lep. i. p. 19. n. 221 (1847) ; Gray, Cat. Lrp. Li.i. Brit. Mus. i. Pap. p. 62. n. 278
(1852); id., Lia Lep. Iiis.Iirit. Mus. i. Pap. p. 72. n. 294 (1866) ; Felder, Verb. Zool. Dnt. Ges.
Wien xiv. p. ,312. n. 318 (1KG4); Godm. &- Salv,, Biol. Cciitr. .Imei:. Lep. Rhnp. ii. p. 231. n. 6,3.
t. 70. fig. 8. 9. cJ (1890) (Yucatan ; Brit. Honduras).
Papiliu piimpeius var. d. P. rogeri, Kirby, Oil. Diurii. Lep. p. 5.39. sub n 144 (1H71) (Yucatan).
S. Disc of forewing, abore., pale from costal to hinder margin, this area
widening costad, its inner edge almost evenly curved ; on uniler.^u/c, a row of grey
spots at distal side of black basi-discal area. Hindwing with short tooth R', the
red spots on disc arranged in two rows as in the preceding insects, the discal row
being represented by only a few spots, usually R- — M^ on upper, and R- — SM- on
under side, some black dots continuing the series to costal vein ; submarginal series
not marked on upperside, or the first as well as the last spot are vestigial ; these
two spots usually distinct on underside, while the other submarginal spots are
either absent, or are indicated by shadowy blackish dots, bearing rarely a few red
scales ; marginal spots creamy, anal one red.
? . Hindwing with two rows of red spots.
Genitalia : cJ. Harpe more strongly rounded ventrally before apex than dorsally,
bearing a few teeth at both edges.
Early stages not known.
Ilab. Yucatan ; British Honduras.
In the Tring Museum 1 6 from Yucatan (received from Messrs. Standioger
and Bf;,ng-Haas).
( 607 )
95. Papilio anchisiades Esp. (1788).
PapiVio Eques Trojaiius aiicliiniittlef: Esper, Au.4. SchifieU. p. 53. n. 22. t. 1.3. fig. 1. (J, 2. $ (1788).
Papilio 2>ompelus, Kirby (noii Fabricius, 1781, err. det.), Cat. Diurn. Lfp. p. 538. n. 144 (1871)
{partini).
c? ? . Hindwing comparatively shorter iu the costal region and longer
abdominally than in P. isidorus and P. rhodostictus ; tail absent or short, always
broader than in the species mentioned, bnt in one of our Bolivian females fully
as long as in those insects. Mai'kinga very variable individually and geographically.
The white markings of underside of forewing form occasionally a discal ami
a submarginal band.
Genitalia : S . Harpe always denticulate, the number of teeth quite variable
individually. ?. A long curved channelled process at proximal side t)f vaginal
orifice ; laterally a broad mnltidentate lobe, and further proximad a high ridge, the
posterior surface of which is concave, shell-like.
Early stages several times described and figured: see below.
Hab. Mexico to Southern Brazil.
Three subspecies.
The name pompeius, proposed by Fabricius for paiit/ioniis Cramer, was meant
to supersede this earlier name. Why Kirby has applied this Fabrician name to the
present insect we do not know ; it was perhaps a mere oversight.
a. P. anchisiades idaeus Fabr. (1793).
Papilio Eques Trojaiius idaeus Fabricius, Eiit. Syst. iii. 1. p. IG. n. 48 (17'J3) ("Madras," Drury ;
Jones'3 fig.).
Papilio idaeus, DonoTan, ]iis. hid. t. 18. fig. 2 (1800) ("Madras") ; Godart, Eiic. ilith. ix. p. 32.
n. 20 (1819) ; Donov., ed. Westw., Ins. Ind. p. 32. t. 1'.). fig. 2 (1842) (" Madras") ; Gray, Cat.
Lep. Ins. Brit. ilus. i. Pap. p. G3. n. 279 (1852) [partini ; Mexico ; Honduras) ; Felder, Verli.
Zool. Bot. Ges. Wien xiv. p. 312. n. 31() (1804) (bab. ?) ; Butler, Cat. Diurn. Lep. descr. Fabric.
p. 247. n. 48 (18C9) (Honduras); id. & Druce, Prac. .^oo/. Sor.Loiid. p. 3G5. n. 382 (1874)
(Costa Rica).
Papilio ideas (!), Boisduval, .V- Oin. Lip. i. p. 299. n. 132 (1836) (South America?).
Papilio anchisiades var. a, Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. C3. sub n. 280 (1852) (Honduras);
id., List Lep. Iim. Brit. Mus. i. Pap. p. 73. sub n. 290 (185G) (Honduras).
Papilio idaeus var. a, id., I.e. p. 72. sub n, 295 (185G) [parlim, ; Mexico).
Papilio paiidion Bates, Trans. Eiit. Soc. Loud. (2). v. p. 338 (1801) (Mexico ; Honduras ; nom.
indescr.) ; Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 311. n. 315 (1864) (nom. iiidescr. ; Mexico ;
Honduras) ; id., Reise Xovara, Lep. p. 79. n. 61 (18G5) (Mexico) ; Butl. & Druce, I.e. p. 365.
n. 381 (1874) (Costa Rica); Staud., Exot. Tartf. p. 16 (1884) (Central America); Godm. & Salr.,
hiol. Centr. Amer., Lep. Rhop. ii. p. 230. n. 62. t. 70. fig. 0. 7. $, II. genit. (1890) (Mexico to
Panama).
Papdio anchisiadfs \SiT. jiandion Bates, Pror. Zool. Sor. Loud. p. 242. n. 5 (1863) (Panama).
Papilio anchisiades, Weidemeyer, Proc. Ent. Soc. Philad. ii. p. 146 (1863) (Central America).
Papilio ecander, Oberthiir, Et. d'Ent. iv. p. 80. n. 257 (I88lJ) {partini ; Mexico).
Papilio capys var. paiidion, id.. I.e. iv. p. 80. sub n. 258 (1880) (Mexico).
Papilio pompeius vnr. puiidion, Schaus, Papilio iii. p. 187 (1883) (descr. of adult larva, pupa ; on
Orange and Japote Blanco).
Papilio pandonius Staudiuger, Iris vii. p. 104. note (1894) (nom. nov. loc. jiandinn Feld. iion Wallace).
tS ? . Forewing : a huffish white jjatch in apex of cell, either above and below
or only below, often vestigial ; on icnderside the disc in male usually huffish white
from M' forward, no spot behind M' or only a trace of it, rarely a distinct spot
M' — M-, this spot being sometimes found in specimens which bear a cell-patch on
njiperside ; the huffish white patches larger in female, the row often extending
backwards to SM-, there being submarginal bars distally of the patches in many
( COS )
individuals. Hindwiug somewhat variable in shape, being proportionally shorter
in some specimens than in others ; the two rows of red spots on underside are
sometimes widely separated and small, such individuals resembling rather closely
non-tailed specimens of P. pharnaces, but are recognisable by the more proximal
position of the anterior row of spots, by the paler colour of the spots R-^M- of the
distal row, and by the marginal spots being entirely white or bnftish white, not
being marked with rufons-tawny ; the cell frequently with a red apical dot, especially
often in females.
Jones's fignre represents a female of the present subspecies.
Hub. Mexico to Panama.
In the Tring Museum 80 3S, 00 ? ? , from : Mexico (Salld) ; Jalapa, April
and June 1S9(3 (W. Schaus) ; Huatuxco ; Cordoba, June 1904, 2800 ft. (A. Hall) ;
Guatemala (Salvin) ; Mazatenauga, W. Guatemala, 1000 ft., September 1904
(A. Hall) ; Amatitlan, W. Guatemala, 4S00 ft., August 1904 (A. Hall) ; San
Pedro Snla, Honduras ; San Jose, Costa Rica, 4000 ft. September 1904 (A. Hall);
( Jarreblanco, Costa Rica (Lankester) ; Escazu, Costa Rica, August — September 1903
(Underwood) ; Alahuela, Costa Rica, 4100 ft., September 101)4 (A. Hall) ; Bagava,
Chiriqni, 800 ft. (Watson); Bocj[uete, 2500 It. (Watson); Parida I., January 1901
(Beck).
h. P. ancMsiades anchisiades Esp. (1788).
Merian, Simn. his. t. 17 (1705).
rupiUo Eques Trojanus anclihex Linn^, Syst. Nal. ed. x. p. 4G0. n. 10 (1758) (parti in ; sub citat.) ;
Cram., Pap. Exot. iv. p. 58. t. 318. fig. A. B. C (1780) (Surinam, <? ¥ ) ; Jabl. & Herbst,
Naliirs. Schmetl. ii. p. 24. n. 15. t. 9. fig. 1. 2. 3 (1784) (synon. partiin) ; Stoll, in Cram., l.r.
Suppl. p. 3. t. 1. fig. 2. larva (1787) ; Fabr., Maul. Im. ii. p. 4. n. 28 (1787) {partiin ; sub citat.);
Gmelin, Syst. Nat. 1. 5. p. 2230. n, 11 (1790) (parlim) ; Fabr., Eiil. Sijsl. iii. 1. p. 13. n. 40 (17'.I3)
(jiartiin).
Papilio Eqites Trojanus anchisiades Esper, Ausl. Schmctt. p. 53. n. 22. t. 13. fig. 1. (J, 2. J (1788).
Papilio dumiiiam aticliises, Hiibner, Samml. Exot. Sclimett. i. t. 119. fig. 1. 2. J' (180G— V ).
Priantides liipjmiious id., Verz. hek. Srhmett. p. 87. n. 896 (1818 ?) (nom. nov. loco aiicliises Cram.).
Papilio archelaus Godart, Eiic. Meth. ix. p. 32. n. 19 (1819) (= aiicliixfs, Cram.; Guyane ; "Brazil"
alia subsp.) ; Lacord., Ami. Soc. Eiit. Fr. ii. p. 385 (1833) (larva on orange, social, large
numbers ; Stoll's fig. exact).
PapUio aiicliises, Constable, Miscell. Bull. p. 141. t. 14 (1832) (Surinam).
J'lipilio anchisiades, Boisduval, Spec. Gen. Leji. i. p. 279. n. 103 (1836) (= archelaus = anchises,
Cram. ; Guyane) ; Wall., Trans. Eiit. Soc. Lnnd. (2). ii, p. 255 (1854) (Parii ; forest) ; Gray,
Lisl Lep. Ins. Brit. Jfus. i. Pap. p. 73. n. 296 (1850) (partiin ; Santarem) ; Bates, Trans.
Ent. Soc. Loud. (2). v. p. 338 (18C1) (Amazons) ; Bates, Journ. Entom. i. p. 225. n. 10 (lsfi2)
(common throughout the Amazons ; larva on the [imported] orange-tree ; approach to " var.
isiWora.v" at Ega) ; Felder, Verh. Ziuil. But. Ges. ]\'irn xiv. p. 311. n. 313 (1S04) (Surinam ;
Cayenne ; Amazons) ; Guen^e, Ann. Soc. Ent. France p. 308 (1867) (fig. of Merian quoted by
Linnc' under anchises is anchisiades) ; Druce, Proc. Zool. Sue. Lnnd. p. 240. u. 15 (1870) (Peru) ;
Dew., Arcli. Natiirg. xliv. 1. p. 1 (separ.) (1878) (larva, pupa) ; Hopff., Stetl. Ent. Zeit.-xl.
p. 52. n. 22 (1879) (parlim; Surinam, Peru); Ernst, Ent. Nuchr. xii. p. 79 (1880) (pupa) ;
Caracc, Ent. News ii. p. 52 (1891) (Larva) ; Lathy, Trans. Ent. Soc. Lund. p. 69. n. 37 (1904)
(Cayenne, (J with the patches of the hindwiug blue instead of red).
Papilio anchisiades var. isidoriis, Bates, Trans. Ent. Soc. Loud. (2). V. p. 338 (1861) (Ega).
Papilio Iheramenes Felder, Wieii. Ent. .Man. v. p. 74. n. 9 (1861) (Caracas, Venezuela, ?); id.,
Verh. Zool. Bot. Ges. Wien xiv. p. 311. n. 314 (1864) (Venezuela ; Bogota) ; id.. Prise Nnrara,
Lep. p. 78. n. 60 (1865) (cJ ? ; Colombia ; Venezuela) ; Butl., Ann. May. N.IJ. (4). xx. p. 127
n. 61 (1877) (Cayaria, Peru) ; Godm. & Salv., Trans. Ent. Soc. Loud. p. 120. n. 2,39 (1880)
(Sta. Marta) ; Hahnel, Jris iii. p. 194 (ISUO) (M.'rida) ; id.. I.e. p. 203 (1890) (Valera) ;
Micha»l, ihid. vii. p. 213 (1894) (Sao P.aulo de Oliven^a).
Pnpilin pompeins var. a. P. anchisiades, Kirby, Cat. Oiurn. Lep. p. 5118. sub n. 144 (1871) (Gui*ina ;
Amazons).
( 609 )
Papilio pnmpeim var. b. P. tlieramriies, Kirby, I.e. p. 539. sub n. 144 (1871) (Venezuela ; Colombia).
Papiliopompejits var. aiicJiisiades, Moachler, Verh. Zuol. Bo/. Ges. Wien xxvi, p. 29G (1877) (Surinam).
Pupiliii jtompfitix var. theraiiieiies, Staudinger, Exot. Tarjf.p. llj. t. 11. (J (1884) (Amazons; northern
S. America).
Pap'din /ilaeiis, Maass. & Weym., in Stiibel, Eeiseu S. Amer., Lep. p. 31. n. 133 (1890) (Colombia) ;
iid., I.e. p. 38. n. 32 (1890) (Popayan).
Priamides pompeius, Kirby, in Allen's Nut. Libr., Lep. Butt. ii. p. 284 (1896).
Priamides anchisiades, ii., in Hiihn., SammL Exot. Sdimetl. ed. ii. p. 98. t. 119. fig. 1. 2 (190- ?)
(" a/ic/utjties !, archalaus ! " laps. typ.).
Papilin pompeius, Kaye, Tram. Ent. Soc. Loud. p. 207. n. 197 (1904) (Trinidad ; "anehisiades and
theramems bred from the same batch of eggs ").
c? ?. Both sexes dichromatic, the differences being neither local nor seasonal,
the forms completely intergrading, and each varying again considerably in the
details of the pattern.
In the specimens named anehisiades by Esper and tkeramenes by Felder the
forewing bears two white patches M'— SM- on the disc, either on both sides or on
nnderside only. These patches are very variable in size. There is on underside
often a patch in cell and also freqnently a spot before M' and a second before R',
the spot R^ — M' being occasionally also present on upperside. The posterior
fringe-spots of the hiudwing are usually not much reduced.
In a smaller proportion of specimens the basi-discal area of the forewing is
much deeper brown-black than the apex of the cell and the disc beyond, the pale
area widening costad, bearing on the underside a row of white patches, of which
the discal ones are either in the same position as in the Brazilian P. a. capys,
which these specimens closely resemble, or are more proximal. The posterior
marginal spots of the hindwing are usually very small in these specimens.
Hah. Colombia to Para, southward to Bolivia; common everywhere.
In South-Eastern Bolivia the majority of specimens agree with the following
subspecies.
In the Tring Museum ICO c? c?, 70 ? ? , and a series of larvae and pupae.
c. P. anehisiades capijs Hiibn. (1S06 — ?).
Papilio dominans capys Hiibner, Samml. Exot. Schmett. i. t. 120. fig. 3. 4. J (1806 — ?).
Priamides capt/s id., Vers. belc. Sch7Hett. p. 87. n. 897 (1818?); Kirby, in Hiibn., Snmm?. Exot. Schmett.
ed. ii. p. 98. t. 120. fig. 3. 4, t. 3-25. fig. 1. 2 (190- ?).
Papilio eraiider Godart, Eiic. ileth. ix. p. 32. n. 18 (1819) (Brazil) ; Boisd., Spec. Geii. Lep. i. p. 277.
n. 101 (183G) (Brazil ; = capys) ; Swains., Zool lUiistr. iii. t. 101 (1822) (Brazil) ; Lucas, in
Gu(T., Diet. Pitt. Hid. Nat. vii. p. 47 (1838) ; Felder, Verli. Zool. Bot. Ges. Wieii xiv. p. 312.
n. 317 (18G4) (Brazil) ; Jones, Proc. Lit. Philos. Sue. Limp. p. 43. n. 38 (1883) (larva, pupa) ;
Meldola, Proc. Eid. Soc. Loud. p. 24 (1883) (larva gregarious).
Priamides evander, Hiibner, Samml. Exot. Schmett. ii. t. 112 (1822 ?).
Papilio hipjioiioiis, MeniStries, Mem. Soi\ Imp. Moseuu vii. p. 188. n. 5 (1829) (Brazil, larva).
Papilio idacus, Doubleday {non Fabricius, 1793, err. det.). List Lep. Ins. Brit. Mus. i. p. 11 (1845)
(Brazil; synon. jiartim); id., Westw. & Hew., Gen. Diiirii. Lrp. i. p. 19. n. 219 (1847) (Brazil;
= evander = capys) ; Gray, CcU. Lep. Ins. Brit. Mils. i. Pap. p. G3. n. 279 (1852) (partim;
Brazil) ; id.. List Lep. Ins. Brit. Mus. i. Pap. p. 72. n. 295 (1852) {partim ; Brazil) ; Mon^tr.,
Emim. Corp. Anim. Mus. Petrop., Lep. i. p. 5. n. 85 (1857) (Brazil) ; Maass, & Weym., in
Stubel, Reiseu S. Amer., Lep. p. 91. n. 38 (1890) ; Bonningh., Verh. Ver. Nat. Vnlerh. Hamburg
ix, p. 27 (1896) (Rio de Janeiro, common).
Papilio idaeus var. a. Papilio pompeius. Gray (««)» Fabricius 1781, err. det.). Cat. Lep. Ins. Brit. Mus.
i. Pap. p. 63. Bub n. 279 (1852) (Brazil) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 73. sub n, 295
(1856) (partim ; Brazil).
Papilio anehisiades, id., Cat. Lep. Ins. Brit. Mus. i. Pap. p. 03. n. 280 (18.52) (Brazil "var. a.'' alia
subsp.) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 73. n, 296 (1856) litarlim ; Brazil).
( 610 )
Papilio pompejun. Capronnier, Aim. Soc. Ent. Bch/. xvii. p. 8. n. 2 (1874) (Paquetd, Aug.) ; StanH.,
Exoi. Tatif. p. 16 (1884) (Brazil) ; Seitz, Slell. Ent. Zeit. li. p. 98 (18011) (Corcovado) ; Weym.,
Slett.Enl. Zeit. Iv. p. 315. n. 11 (1895) (Rio Grande do Sul): Mabilde, Guia Pmct. Dorbol. Wo
Grande do Sul p. 45 (189G).
(? ?. Forewing withont white iiatclies on upperside, bearing always a pale band
which widens costad, its inner edge crossing cell at or just beyond base of R'" ; this
band emphasized on umlerside by a row of wliite patches, one of which occni)ies
apex of cell in all specimens. Tooth W of hindwing often projecting.
Hah. Plains of Eastern Bolivia ; North Argentina ; Paraguay ; and Brazil.
In the Tring Museum 64 <?cJ, 26 ? ?,from: Sapacay, Paraguay (W. Fostei) ;
Yhu, E. Paraguay (Andeer) ; La Soledad, Entro Rios (Chas. Britton) ; R. Grande
do Sul ; Castro, Parana (E. D. Jones) ; Bahurn, 8. Paulo (Dr. Hempel); Petropolis,
Tijuca, Rio de Janeiro (Foetterle ; E. May) ; Minas Geraijs (Kennedy) ; ( "eara.
90. Papilio isidorus Doubl. (1846).
rapilio isidorus Doubleday, An/i. Hay. X. H. xviii. p. 375 (1846) (BoliTia).
c??. Forewing below withont a bufKsh white patch across the cell, the discal
patch just entering cell in most males, being absent from other males and from the
female (only one seen) ; the male bearing usually a huffish white discal patch also
on upperside, the patch being here much smaller than below and much shaded with
black. The last red spot of the submarginal row of the hindwing stands distally
of the large spot which is at its discal side. Hindwing much shorter posteriorly
than in F. anchisiades, tail more or less distinct, narrow.
Genitalia : S. Harpe non-dentate, its dorsal edge more strongly rounded near
the apical process than the ventral edge
Early stages not known.
Hab. Chiriqui to Bolivia.
The geographical forms are not quite constant.
As previous authors did not strictly distinguish P. isidorus from P. rhoJoMctus,
we have discarded all references which appear doubtful.
a. P. isidorus c/tiroins subsp. nov.
S. Forewing, npperside, with bufiish white spots R-— M-\ the first one minute,
the third not quite reaching M=, a small si)ot in cell ; on underside, these jJatches
all large, the cell-sjwt being the smallest, the triangular spot R-— R' coming next
in size,^ while patch R^— M' is the largest, being about twice as long as broad.
Hindwing, above, with the two red spots R^— R^ merged together; discal spots
R^— M- of underside well marked, not separate from the submarginal ones.
Hab. Chiriqui.
In the Tring Museum 1 tj (received from Messrs. Staudinger & Bang-Haas).
b. P. isidorus briscs subs]), nov.
S. Forewing, underside, with greyish white spots R^— M= or R'-— M=, upper
spot small ; no s])ot in cell. Hindwing, above, with two red spots R'— M',
the pro.ximal (= discal) portion of spot R-— M' about as large as the distal
(= submarginal) portion ; on underside the discal portions of spots R^— M- standing
separate from the submarginal ones, small, vestigial.
II lb. Bogota; probably the Magdalena valley.
In ihe Tring Museum 2 c?(?.
( fill )
c. p. isidorus f/avescens Oberth. (1880).
Papilio isidorus var., Gray, Cat. Lep. Ins. Brit. Mus. i. Pup. p. 04. sub n. 281 (18.')2) ; id., List Lep.
Ins. Brit. Mus. i. Pap. p. 73. sub n. 207 (I85fi) (hab. ?).
Papilio isidorus y3.T.J!urescf>is Oberthiir, Et. d'Enl. iv. p. 79. sub n. 2.')4 (1880) (Colombia).
Papilio isidorus var. leiicostictiis Honrath, Berl. Ent. Zeitschr. xxix. p. 276 (188o) (Colombia) ;
Maass. & Weym., in Stiibel, Reisen S. Amer., Lep. p. 31. sub n. 134 (1890) ("Bogota");
Dognin, Lep. Loja p. 37 (1891).
(?) Papilio ceus " Boisd.," Dognin, I.e. i. p. IS ( 1887) (itom. mid.).
Papilio isodorus, Maassen & Weym., I.e. p. 77, n. 33 (1800) (Rioja to Moyobamba).
cJ. Forewing, iinder.fidc, the white patch reduced. — — Hindwiug usually with
one spot io cellale R- — R' on npperside ; this spot as well as the ne.xt one mostly
white, either on both sides or only on one side of the wing ; discal portion of
spots R' — M' of underside separate from the distal (= submarginal) portion, or
obliterated.
?. Forewiug above and below paler brown distally in cell and beyond
than in basi-discal area ; on underside a row of ill-defined white submarginal
spots from R' to SM-. Hindwing above and below with the basi-discal area
paler than in male ; submarginal patches R^ — M- much enlarged, white ; four
submarginal spots C — W', the fourth white, the others more or less reddish,
almost entirely rufous red on underside, a rufous red anal submarginal sjwt ;
five discal spots, small or vestigial, spots R^ — M- being merged together with
the large white patches ; tail short, acute.
Hab. (Colombia (probably the south-east) ; Eastern Ecuador ; North Pern.
In the Tring Museum 8 Si, 1 ?,from: Arcliidona, April 1899 (W. Goodfellow);
Coca, R. Napo, May— July 1899 (W. Goodfellow) ; Mirador, February 1899 ;
Zamora (0. T. Baron) ; Loja.
In coll. Oberthiir from :" Bogota " ; Ambato ; Archidona; Moyobamba.
d. P. isidorus isidorus Doubl. (1840) (PI. VIII. fig. 56).
Papilio isidorus Doubleday, I.e. (1846) (Bolivia) ; id., List Lep. Ins. Brit. Mils. i. Append, p. 3(1848);
Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 63. n. 281. t. 7. fig. 4. ^ (1852) (Bolivia) ; id., List
Lep. Ins. Brit. Mus. i. Pap. p. 73. n. 207 (1856) ; Felder, Verh. Zool. Bol. Ges. Wieu xiv. p. 312.
D. ,320 (1804) (partim) ; Druee, Proc. Zool. Sor. Lond. p. 246. n. 16 (1876) (Pozuzo) ; Hnpff.,
Sletl. Ent. Zeil. xl. p. 53. n. 23 (1879) (distinct from auchisiades ; partim; '' Brazil," Peru,
Bolivia) ; Haase, Untersuch. Mimicry i. p. 99 (1803) ; Weeks, lUustr. Diiim. Lep. p. 20 (19ii5)
(Chulumani).
Papilio jjimpeius var. e. P. isidorus, Kirby, Cat. Diurii. Lep. p. 530. sub n. 144 (1871) {jmrlim).
(?. Forev/ing, on underside, with white patch which nearly always enters the
cell and is often vestigial also on upperside, some specimens, however, being
without trace of this patch. Red spots R^ — M- of hindwing, above, on the
whole larger than in the previous form, two spots in cellule R- — R^ separate
from one another ; on underside the small discal spots separate from the
submarginal ones.
Hab. Eastern slopes of Bolivia and Peru, as far north as Huannco.
In the Tring Museum 21 c?c? from: Poziizo, Huanuco, 8ii0— ItiOO m.
(W. Hoffmanns) ; C'hanchamayo (Schunke) ; Caradoc, Marcapata, 4000 ft., February
1001 (Ockenden); Perdnfi R., March 1900 (Simons) ; Palcazu (Sedlraayr) ; Cuzco,
March 1901 (Garlepp) ; Mapiii.
( 612 )
97. Papilio rhodostictus Bntl. & Drnce (1874).
cJ. PapUio rhoilnsticlus Butler & Driice, Proc. Zool. Sr'C. Loml. p. 3G4. n. 370 (1874) (Costa Rica).
c?. In shape of the wings agreeing with F. isidorug. Forewing, either on
both sides or only below, with a white patch across the cell near apex ; on disc two
white patches R=— M', always present below, but on upperside either both present,
or the first, or the second, or both absent ; on underside only an additional white
streak behind M'. Hindwing: spots U- — M- larger than in P. isidoriis, the discal
and snbmarginal spots being confluent forming three large patches on upperside ;
seldom the two spots in cellule R- — R' slightly separate, the line of separation,
however, being often indicated ; on underside the two spots R- — R' mostly separated,
nearer the cell than in P. isidor/ts, and therefore the snbmarginal spot R^ — R-
much more distal than the spot R- — R^
? . Similar to male, wings broader, markings larger.
Genitalia : c?. Harpe more symmetrical than in P. isidorus, the dorsal and
ventral edges being almost equally rounded near apical process.
Early stages not known.
Ilab. Costa Rica to Western Ecuador.
a. P. rkodostictus rhodostictus Butl. & Drnce (1874).
PapHin rhodcisfuius Butler & Druce, I.e. (1874) (Costa Rica) ; Kirby, Cal. DUmi. Lep. p. 814. n. .'ISS
(1877) ; Godm. & Salv., Bh.l Cent,-. Amer., Lip. lihap. n. p. 232. n. 67 (1890) ($ ? ; Costa
Rica ; Chiriqui ;— "Ecuador" alia subsp.).
^?. Cell-patch of forewing narrow, discal spot R-— R^ larger than spot
j{,3 — ]yji^ the latter sometimes absent from upperside, rarely larger than the
preceding spot but then more or less shaded over with black.
Eab. Costa Rica ; Chiriqui.
In the Tring Museum 'i S i from : Boqnete, Chiriqui, 3500 ft. (Watson) ;
Chiriqui.
b. P. rhodostictus pacificus subsp. nov. (PI. VIII. fig. 49).
Papilio rhoihstldus, Godman & Salv., Biol. Cenli: Amcr., Lep. Rhop. ii. p. 232. n. G7 (1890) l^partim;
Ecuador).
P»y<,7;o ;jaH(/;o« ?, iid., in Whymper, ^«rf<;s of Equator, Ajip. p. 109. n. 97 (1891) (west of Quito,
with spot at end of cell of forewing).
cJ ? . Forewing : cell-patch larger than in the preceding, discal spot R=— R'
smaller than R' — M', sometimes vestigial on ttpperside.
Hab. West Colombia and West Ecuador.
In the Tring Museum 24 c?c?, 1 ? , from : R. Dagna (Rosenberg), type;
Paramba, W. Ecuador, 3.300 ft., May— June 1897, dry season (Rosenberg) ; Lita,
3000 ft. (Flemming).
In coll. Oberthi'ir a series of males from Juntas, R. Dagna.
c. P. rhodostictus nymphius subspec. nov. (PL VIII. fig. 48).
S. Forewing without white spot on upper.^ide ; the markings of underside
as in P. r. pacijicus, but smaller. The three patches of hindwing sometimes
creamy white above and below, very often creamy white bordered with red, normally
red above.
( 613 )
Ilab. Central and Eastern Colombia, rather frequently found in Bogota
collections.
In the Tring Museum 40 c?c? from : BIuzo, July 1903 (M. do Matlian) ; Mnzo
(Lindig, coll. Felder) ; " Bogota."
IX. Torquatus Group.
Palpus yellow ; breast yellow or spotted with yellow ; abdomen for the greater
part yellow, or at least with a yellow lateral line ; both sexes tailed : the sexes
different in colour ; red discal spots of underside of hindwing round, not luniform,
or the row represented by one or a few spots only ; in male, cell of forewing below
striped with yellow or entirely yellow, no bar across it.
Neuration : SC^ of forewing at about one-third from SC^ to SC-''; lower
angle of cell much more obtuse than upper ; D- longer than D' ; basal cellule of
hindwing very little prolonged beyond PC, almost truncate ; PC strongly arched.
Larva similar in colour to that of P. amifogeiis, with four rows of rather
prominent tubercles. Thoracical projection of pupa long.
Key to the species ;
A. Males.
a. Band of forewing not broadly interrupted, the upjiermost
spot standing before SC^ or in fork, separate from base
of fork b.
Band of forewing broadly interrupted, or continued to
costal margin around apex of cell, uppermost spot at
base of subcostal fork small, often absent, or the band
quite short d.
b. Patch SC'"' — R' of band of forewing (second or third from
apex) long, reaching beyond base of subcostal fork , c.
This patch smaller than the next, widely separate from
base of fork Species No. 100.
c. Tail with yellow spot at apex Species No. 98.
Tail without yellow spot at apex Si^ecies No. 99.
d. Band of forewing complete, extending from inner to costal
margin, or interrupted e.
Band abbreviated, extending from inner margin to about
middle Species No. 103.
e. Band of forewing complete, consisting of a series of spots
which are all separated from one another . . . Species No. 104.
Band either broadly interrupted at R- or continuous, the
veins not black /.
/. Hindwing below with two orange-red spots behind M-,
band of forewing always broadly interrupted . . Species No. 102.
Hindwing below with one red spot behind M-, band of
forewing often complete, tooth M' of hindwing as
prominent as tooth M= Species No. 101.
B. Females (not known of several species).
g. Hindwing with a row of strongly arched submarginal spots,
either on one side or on both h.
( 614 )
Hindwing without curved snlimarginal spots, but with
two or three large patches R- — M- which touch cell . i.
Ii. Tail with 3-ellow spot at apex Sjiecies No. 98.
Tail without yellow spot at apex Species No. 100.
i. Hindwing crossed by a white baiul which extends to
abdominal margin ....... Species No. liJ3.
Hindwing without such a band ...... Species No. 1 1)2.
98. Papilio himeros Hopff. (1866).
cJ. Papilio mentor Boisduval («o» Dalman, 1823), Spec. Gen. Up. i. p. 351. n. 193 (18.3G) (Brazil).
? . rapilio himeros Hopffer, SMI. Enl. Zeil. xxvii. p. 2G. n. 7 (18Gt;) (Brazil).
Both sexes bear a yellow spot at the ajiex of tlie tail. Cell of forewing beneath
yellow, at least not merely striped.
(?. Band of wings very broad, on hindwing extending beyond apex of cell ;
OD forewing a yellow spot distally of upper angle of cell.
?. Band narrower than in male, and the patch SC* — R' of forewing much
smaller ; no spot before upper angle of cell. Snbmarginal sjjots of hindwing
small, red, except upper two, which are yellow.
Genitalia : <S . Tenth tergite non-spatulate. Harpe long, reaching close to
apex of clasper, gradually tapering, being elongate-triangular, acute. ? not
dissected.
Early stages not known.
Hab. Brazil.
Two subspecies.
a. P. himeros baia sabsp. nov.
cJ. Yellow band of wings a little narrower thau in the following subsjjecies,
the black distal area of hindwing nearly touching cell ; the additional spot before
ujijier angle of cell of forewing small ; cell of forewing below brown, striped and
washed with yellow ; yellow snbmarginal band of forewing below about half the
width of the brown band situated at its ]>roximal side ; snbmarginal spots of
hindwing a little smaller than in P. It. himeros.
?. Yellow band very much narrower than in the following form, on forewing
much narrower than the brown distal border, not touching cell, on hindwing
extending to apex of cell, the extreme tip of cell remaining brown, proximal edge
of baud crossing cell at point of origin of SC- and just proximally of M-.
Hab. Bahia.
In tlie Tring Museum 1 i,l ?.
b. P. himeros himeros Hopff. (1866).
Papilin mentor Boisduval, I.e. ; Doubl., Li.sl Lep. Ins. Brit. Mus. i. p. 17 (1845) (Brazil) ; id. ,
Westw. & Hew.. Gen. Diurn. Lep. i. p. 17. n. 165 (1846) ; Gray, Cat. Lep. Ins. Brit. ilus.x.Pap.
p. 38. n. 190 (1852) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 52. n. 199 (1856) (Brazil) ; Felder,
Verh. Zool. Bot. Ges. Wien xiv. p. 310. n. 305 (1864) (Brazil) ; Kirby, Cat. Diurn. Lep. p. 540.
n. 151 (1871 ) (Brazil) ; Burm., Descr. Rep. Argent. V. Lep., Atlas p. 5. n. 6 (1879) (Corcovado) ;
id., I.e. p. 61 (1879) (descr. of $ ) ; Oberth., Et. d'Enl. iv. p. 71. n. 214 (1880) (Brazil ; liii'es
of mentor) ; Staud., Exol. Tagf. p. 16 (1884) ; Haase, Vnttrsueh. Mimicry i. p. 9(i, 100 (1893) ;
Bonningh., Verh. Ver. Nat. Unterh. Hamburg ix. p. 27 (1896) (Santa Tlieresa, Rio de Janeiro,
rare).
?. Papilio hjcophron var. A,, Boisduval, I.e. p. 352. n. 194 (1836) (partim ; "tail with yellow spot
at apex ").
( 615 )
?. Papilio hhiieros Hopffer, Stell. Ent. Zeit. xxvii. p. 20. n. 7 (18GG) (Brazil) ; Kirby, Cat. Diurn.
Lei>. p. 5G7. n. 3Sfi (1871).
?. PapiVtn herodnlus C&'pTonmev, Aim. Soc. Ent. Belij. xvii. p. 10. n. 14(1874) (nom. indescr. ;
Batofogo, August) ; Oberth., Et. d'Enl. iv. p. 71. n. '215. t. 4. fig. 2 (1880) (Brazil).
(^. Papilio hjcoiiliron var. minor; P. mentor^ Burmeister, Descr. Rip. Artje/if. v. L/p. p. CO. sub n. 2
(1878).
" Dalman a doiine le nom de MeHtor" .says Boisduval, Lc, "a nn autre Papilio
qui est le snivaiit, et qu'Hubuer avait fait counaitre avant lui sous le uom de
Lycophron ; nous avons cru pouvoir prendre sans inconvenient le nom de Dalman
pour I'appliqucr a celui-ci qui est nouveau." This is a mischievous practice to
which also Fabricius adhered, causing much confusion.
The 3-ellow band of the upperside touching cell on forewing and extending
beyond tip of cell on hiudwing, broader on forewing of both sexes than black
marginal border. Some males with a vestige of a yellow spot in cell of forewing.
Hub. Brazil : Minas Geraes ; Rio de Janeiro.
In the Tring Museum 5 c?c?, 2 ? ? , from liio de Janeiro.
99. Papilio lamarchei Stand. (1892).
Pupdio lamarchei Staudinger, //■;'.< v. p. 428 (18'J2) (Bueyes, Bolivia ; tonnmliitux Esp. var. V).
c?. Upperside. — Forewing : yellow band narrower than in P. himeros, the apical
spot absent, the next sjiot larger than in P. himeros, the black marginal border
narrowing apicad ; spot SC* — R' eijlarged as in P. himeros, being much larger
than in P. hectorides ; no additional spot distally of upper angle of cell.
Hindwing even more strongly dentate than in P. hectorides, tail fringed with
yellow, not bearing a yellow spot at apex, yellow band not extending to apex
of cell.
Underside resembling that of P. hectorides, but more extended yellow.
? not known.
Genitalia: 3. Harpe short, apex ronnded and dentate.
Hub. Northern Argentina and Bolivia.
Nearest to P. hectorides, bnt approaching also a little P. himeros.
In the Tring Museum 11 c?c? from : Tucumau (J. Steinbach) ; Tapia, Tucuman
(Baer) ; Bueyes ; II. Tanampaya (Garlepp).
100. Papilio hectorides Esp. (1794).
?. Papilio Eques Trnjaiiiis hedoridex Esper, Magaz. Ansl. Iiis. i. p. 5. t. 1. fig. 1 (1794) ; id., Au.il.
Schmelt. p. 249. n. 115. t. 40c fig. 1 (1798) (" Ost-Indien ").
(J. Papilio Eques Achirus lorquatinwt id,, p. 206. n. 94. I.e. t, 51. fig. 2 (1798) (" Surinam ").
(J. Papilio pandromsGoia.\t, Eiu: Milh.\%. p. G2. n. 101 (1810) (" Guyane " ; Brazil); Swains.,
Zool. Illuntr., Eut. ii. t. 93 (1822) (Rio de Janeiro).
? . Papilio li/sitlioux Godart, Eiic AJelli. ix. p. 73. n. 13G (1819) (Brazil) ; Lucas, Lep. Kxol. p. 32.
t. IG. fig 2 (1835) ; id , in Chenu, Em: Hist. Nat., Pap. i. t. 10. fig. 2. ? (1851-53).
? . Thuas hj.'iillious. Swainson, I.,: ii. t. 121 (1822) (Brazil).
'i. Meiielaide.i chirodaiuas Hiibner, Saiiiml. Exol. Sriimcll. ii. t. 103 (1822?).
?. Pajiillo he.:loride>i, Donovan, Nat. Repos., Ent. t. 177 (1827) : Boisd.. Spec. Gen. Up. i. p. 303.
n. 137 (183C) (Brazil) ; Doiibl., Lid Lep. Ins. Brit. Mus. i. p. 13 (1845) (Brazil) ; id., Westw.
& Hew., Geu. Diurn. Lep i. p. 17. n. 180 (1840) (Brazil).
cJ. Papiliii torqualinus, Boi.sduval, Spec. Gen. Liji. i. p. 368. n. 212 (1836) (= pandrosus ; Brazil) ;
Doubl., List Lep. Ins. Brit. .Uuf. i. p. 13 (1845) (Brazil); id., Westw. & Hew., Gen. Diurn. Lep.
i. p. 16. n. 143 (1846) (Brazil) ; Gray, Cat. Lep. Ins. Brit. Mus. i. p. 34. n. 1G5 (1852) ; id., LiM
Lep. Ins. Brit. Mus. i. Pap. p. 46. n. 173 (18JG) (Brazil) ; Menetr., Enum. Corp. Aniut. Mus.
Petr., Lep. i. p. 4. n. 52 (1857) (Brazil).
( fllfi )
$. Papilio mezenthis Doableday, Ann. Maq. N. H. xiv. p. 417 (1844) (West Coast of America) ;
a.. List Lep. Ins. Brit. Mug. i. p. 13 (1845); id., Westw. & Hew., Gen. Diitni. Lei), i- P- 17.
n. 181 (184C) ; Gray, Cat. Lep. Ins,. Brit. .V»«. 1. Pap. p. 40. n. 205. t. 3. fig. 4 (1852) ; id., List
Lep. Ins. Brit. .!/«.«. i. Paj). p. 50. e. 21G (1856) ; Weidem., Proc. Ent. Sue. Philad. ii. p. 147
(18G3) ("Mexico," errore) ; Felder, Verli. Zuol. Bat. Ges. Wien xiv. p. .310. n. 304 (ISlU) ;
Kirby, Cat. Diiini. Lep. p. 540. n. 152a (1871) (" Amer. occ") ; Wood, fns. Abruad p. 549. fig.
299 (1883) ; Staud., A'xo/. Tag/, p. 10 (1884) (-perhaps a ? form from western (?) America)."
? . Papiliu arijentiis Martyn, Psi/che t. 14. fig. 3. 4 (1797) (ined.) ; Gray, Cal. Lep. Ins. lirit. Mas. i.
Pap. p. 40. n. 204 (1852) (j = li/silhous Godt., ? = argenliis Martyn— error, both ? ? ) ; id.,
List Lep. Ins. Brit. Mus. i. Pap. p. 50. d. 215 (1856) (BrazU).
cJ?. Papilio argent IIS. Y&Mer, I.e. i^.ZXO. n. 303 (18G4) ((J = torquatinns) ; HopfEer, Sirll. Ent.
Zeit. xl. p. 52. n. 20 (1879) ( ? = hcctoriiles, ^ = torquatinns ; Brazil, Maxos in Bolivia).
(J ? . Papilin hectorideg, Kirby, Cat. Diurn. Lep. p. 540. n. 152 (1871) ( ? = argentus = cliirmlamas
= lysithous ; (J = torquatinns = j'amlrosus) ; Capronn., Ann. Soe. Ent. Belg. xvii. p. 10. n. 15
(1874) (Itaipu, October) ; Staud., E.vot. Tagf. p. 10. t. 11. <? ? (1884) (Brazil) ; Seitz, SW^
Ent. Zeit. li. p. 97 (1890) (Corcavado, common) ; Haase, Untersucli. Mimlrry i. p. 97. t. 10.
fig. 07. cj, 08. J (1893) ; Weym., Stelt. Ent. Zeit. Iv. p. 315. n. 13 (1895) (Rio Grande do Sul) ;
Mabilde, Guia Prael. Borbnl. Rio Grande do Sul p. 40. t. 1. fig. 3. ^ (1890) ; Peters, Illustr.
Zeitschr. Ent. ii. p. 52 (1897) (Nova Friburgo, both sexes black with white band, errore ; larva,
onAnagyrisfoetida?) ; Schroder, ibid. p. 485. fig. 1. 2 (1897) (1., p.) ; id., I.e. p. 497. fig. 3. 4
(<J, ? ) (1897).
(J ?. Papilio torquatinns, Burmeister, Descr. Rep. Argent, v. Lip., Atlas p. 5. n. 8 (1879) (larva and
pupa ; on a species of Piperaeeae) ; Oberth., Et. d'Enl. iv. p. 78. n. 251 (1880) (Brazil) ; Gosse,
Entoui. xiii. p. 194 (1880) (Paraguay, Dec. to March) ; Bijnningb., Verli. Ver. Naturio. Unlerh.
Hamburg ix. p. 26 (1890) (common ; larva on Citrus and Piperaeeae).
¥ . Papilio torquatinns aberr. $ , melania Oberthiir, Et. d'Enl. iv. p. 78. sub n. 251. t. 3. fig. 3 (1880)
(Brazil) ; Weym., Stetl. Ent. Zeit. Iv. p. 315. sub n. 13 (1895) (R. Grande do Sul).
9 . Papilio hestorides (!), Peters, I.e. ii. p. 61 (1897) (Nova Friburgo ; larva ; " (J of hectorides"
false).
Troilides heetorides, Kirby, in Hubn., Sanunl. Exot. Schmett. ed. ii. p. 97. t. 317. fig. 3. 4 (190- ?)
(Esper '■ 1785 " false).
The sexes dissimilar, tlie females being again diclaromatic. The hind wing
strongly dentate.
c?. Upperside: band almost gradnally widening from apex of fore wing to
abdominal margin of hindwing, narrower than in /'. lamarchei, varying somewhat
in width and also in tint ; many specimens with a sjjot before SC'\ others being
without it. Hind wing : from one to six red spots on disc, the anal submarginal
spot being also red. — Cell of forewing striped beneath.
? . Band of wings, if present, white, usually not extending to abdominal margin
of hindwing, stopping at M or at M' or IP ; on forewing somewhat different in
position from the band of male, being nearly parallel to distal margin, slightly
curving away from it anteriorly, crossing the subcostals just outside fork. Sub-
marginal spots of hindwing red, thin. Three colour-forms :
a. ?-f. heetorides Esp., I.e.; mecentius Doubl., I.e.; argentus Gray (ex
Martyn), I.e. — Both wings with white baud ; the band in one of our specimens
continued to abdominal margin of hindwing by means of two white spots which are
merged together with the last two red spots of the discal series ; the band very
variable in width.
Ij. ¥ -f. catamelas nov. — Band vestigial on forewing, distinct on hindwing, but
rather narrower than in ?-f. heetorides.
c' . ?-f. melania Oberth., I.e. — Band vestigial on both wings or absent.
Genitalia : J. Tenth tergite long, spatulate ; sternite laterally with a ridge
which is transversely divided into two short tubercles or teeth, the proximal one being
long- hairy. Clasper emarginate ventral ly at the apex ; harpe long, flat, its ventral
( 617)
margin nearl}' straight ; apex truncate, bearing several long teeth. ?. Edge of
the circular orifice raised in front to a very short glossy lip, the long process found
in P. torquatus being absent ; on each side a little way from the orifice a large,
variable, dentate lobe; behind vaginal orifice rather strongly chitinised brown sclerite,
this large jilato posteriorly incised in the middle ; anal segment on innerside without
the usual curved bristles or bearing only two rather thin ones (accessories in
ovipositing).
Early stages described by Burmeister, l.c.^ and figared by Schroder, I.e.
Ilab. Brazil ; Paraguay.
In the Tring Museum 06 cjc?, 45 ??, from: Sapucay, Paraguay, all months
from July to February (W. Foster) ; Yhu, Paraguay, September — December 1897
(Andeer) ; Bahia ; Minas Geraes, December 1898, February 1901 (A. Kennedy);
Espiritu Santo ; Rio de Janeiro ; Gorcovado ; Nova Friburgo ; Petropolis ; Sao
Paulo ; Castro, Parana (E. D. Jones) ; S. Catharina ; Blumenau ; Porto Alegre.
101. Papilio garleppi Stand. (1892).
$. Papilio garleppi Staudinger, Iris v. p. 427 (1892) (S. Mateo, R. Juntas, Chapare).
(?. Upperside : yellow band much broader than in P. torquatus, extending
beyond apex of cell on hindwing. Forewing, the band interrupted or not, the
spots composing the costal portion large. Hindwing : tooth M' prominent, acute,
longer than in P. torquatus; the red anal spot preceded by two or three spots
which are composed of dispersed yellowish butf and bluish white scales ; these
spots on a level with the red anal one, corresponding to the bluish spots of the
underside ; no red spots proximally of them.
On underside the hindwing bears a row of rufous red discal spots as in
P. torquatus, but the spot behind M- is replaced by a tiny bluish dot, and spot
M^ — M^ is the largest of the series ; the yellow submarginal spots are more or
less luniform.
? not known. Staudinger, when describing in 1892 a subspecies of P. garleppi,
referred to the yellow-spotted female of P. torquatus figured by Gray (P. patros
var.) as being possibly the female of P. garleppi. However, Gray's specimen is
undoubtedly a female of P. torquatus.
Genitalia: cj. Very different from those of P. torquatus; tenth tergite very
slender, long, curved ; sternite on each side with a curved, smooth ridge which is
posteriorly produced into a tapering point, the ridges of the two sides curved
towards each other, being hairy on innerside. Clasper emargiuate ventrally at apex ;
harpe ending in a long, sharp point, and bearing ventrally beyond middle a long
pointed process.
Ilab. Bolivia to the Upper Amazons.
Two subspecies.
a. P. garleppi interruptus Stand. (1892).
Papilio torquatus, Staudinger (ho//. Cram., 1777, err. det.), Exot. Tug/, t. 11. ^ (1884).
(J. Papilio garleppi var. interrupt ux id., Iria v. p. 427 (1892) (.S. Paulo de Olivenfa, Shanusi,
Chanchamayo ; ^^ patros Gray perhaps J of this insect ') ; Michael, ibid. vii. p. 209 (1894).
Baud of forewing interrupted between R- and R^ as in P. torquatus.
Hah. Upper Amazons ; Eastern Peru, as far south as Carabaya.
In the Tring Museum 3 cJcJ from : La Union, R. Huacamayo, Carabaya, S.E.
Peru, 2000 ft., December 1904 and January 1905, wet season (Ockenden).
( 618 )
b. P. garleppi qarlcppi Stand. (1892).
(J. Papilio garleppi Staudinger, I.e.
Band of forewing not, interrupted, but there is nsnally a more or less distinct
vestige of an interruption.
J I'll). Bolivia.
Jn the Tring Museum (i J c^ from Ma]iiri.
II '2. Papilio torquatus.
cj. Papilio Eques Achirtis tdrfptatiix Cramer, Pap. Exot. ii. p. 123. t. 177. fig. A. B (1777) (Surinam).
?. Prinreps dominans caiidiii.': Hiibner, Snmml. Exot. Schmctt. i. t. 117 (180G — ?).
Sexes very dissimilar, the female resembling certain females of Aristolochia
Papilios, occurring like the.se in forests, while the male frequents more open
localities. The female bears also a close resemblance to P. ?s>(/orus and
P. rhodostictus, being, however, easily recognised by the colour of the spots on the
thorax and the huffish line on each side of the abdomen.
S ■ The band of the forewing is interrupted between R- and 11' ; however,
there is occasionally a spot in front of W which is sometimes so large as to bridge
over the gap, though not entirely filling it up. On the underside of the hindwing
there is a discal row of five or sis rufous red spots, variable in size, the last
standing before abdominal margin ])roximally of the anal submarginal spot; of this
row the fifth spot counted from behind is the largest in nearly every specimen.
? . Wings brown-black. Forewing with or without white patches, in our
only Bolivian specimen on underside a creamy buff, submarginal, ill-defined
band. Hindwing, on yppernidc, with a complete row of red submarginal sjiots
and an incomplete row of red discal ones, the discal row extending from SM-
forward to R' or R^ ; the discal and submarginal spots R' — M^ in the South
American forms always merged together to large elongate patches, often also
spots R- — R' forming together a third patch, these jmtches occasionally creamy
on the Amazons, the development being on the same lines as in P. isidorus and
P. rhodostictus ; the rows of sirots remaining separated in the Mexican form.
Genitalia : S . Tenth tergite spatulate, sternite on each side with a distally
truncate ridge which is more strongly chitinised than the rest of the segment, the
ridges of the two sides forming in dorsal view a pair of ( ). Clasper somewhat
acuminate ; harpe broad, widening distally, being widest a short distance from
ajiex, the ventral edge almost straight, the dorsal edge slightly angnlate, apex
obtuse, the surfixce feebly concave, distally practically flat and longitudinally
wrinkled, the edges ajiically densely denticulate, dispersed minute teeth also along
the ventral edge. ?. Edge of vaginal orifice raised proximally into a large
obtnse process which is curved distad, being longitudinally impressed on the
proximal surface, slightly tapering apically, the apex being truncate ; behind
the orifice a small rounded lobe densely covered with minute hairs ; laterally
of the vaginal process several low ridges connected with one another, no large
dentate lobe as in P. anchisiades and P. hectorides.
Early stages described by Bnrmcister, Dcscr. Rep. Argent, v. Ldp., Atlas p. 5.
n. : (1S79).
JJab. Mexico to Soath-East Bolivia, Paraguay, and Rio de Janeiro.
Six subspecies.
( 619 )
a. P. torquatm tolus Godm. & Sal v. (1890).
PapiUo tolus Godman & Salvin, Biol. C'etdr. A met:, Lep. Rhop. ii. p. 228. n. 59. t. 71 1. fig. 1.2. cJ,
3. 4. ? (1890) (Tampico ; Mexico) ; Haase, Untersuch. Mimicrij i. p. 98 (1893).
5. Band of forewing narrow, streak SC° — R^ long, streak R' — R^ short, a
large spot R^ — R', the gap in the band being narrow, some small spots at upper
angle of cell. Tail broad, submargiual spots large.
? . No spots on Ibrewing. Hiudwiiig with two separate rows of red spots,
only spots R^ — M' connected with one another by some diffuse red scaling ; tail
long, spatnlate.
Ilab. Mexico.
h. P. tovquatus tolmidcs Godm. & Salv. (1890).
Papll'io tolmides Godman & Salvin, I.e. p. 229. n. CO. t. 70. tig. 5. ^ (1890) (Panama : Bugaba,
Chiriqui, Veragua).
(?. Forewing : band broader than in the preceding form, no spots before upper
angle of cell or only minute ones ; streak SC* — R' longer than streak R' — R",
patch R- — R^ present, the gap in the band being sometimes almost completely
filled up. Tail narrower than in P. t. tolus, very feebly spatulate.
Harpe slightly sinuate or incised dorsally.
? not known.
Hub. Panama : from Chiriqui southwards ; Sevilla I.
May be expected to occur in Costa Rica.
In the Tring Museum 'i 6 <S from : Chiriqui ; Sevilla I., January 1902 (Batty).
c. P. torquatus orchamus Boisd. (1836).
?. Papilio orchamus Boisduval, S^iec. Gen. Lep. i. p. 300. n. 133(1836) (Colombia) ; Doubl., Westw.
& Hew., Geii. Diuni. Lep. i. p. 18. n. 192 (1810) (Colombia) ; Gray, Cat. Lep. Ins. Brit. Mus. i.
Pap. p. 44. n. 222. t. 7. fig. li (1852) (Venezuela) ; id., List Lep. Ins. Brit. Mas. i. Pap. p. 59.
n. 235 (1856) (Venezuela) ; Felder, Verh. Zool. Bat. Ges. Wien xiv. p. 310. n. 297 (1861)
(Venezuela) ; Oberth., Et. d'Ent. iv. p. 79. n. 253 (1880) (Colombia, type).
?. Papilio torquatus var. c. P. orchamus, Kirby, Cat. Diurn. Lep. p. 541. sub n. 151 (1871)
(Venezuela).
(J ?. Papilio ore/<am«.s, Oberthiir, ;.<_■. p. 116. n. 253. t. 6. fig. 3. ^ (1880) (Colombia) ; Godm. &
Salv., Biol. Centr. Amer., Lep. Rliop. ii. p. 228. sub n. 59 (1890) (Colombia, Venezuela),
fj. Papilio torquatus, Hahnel, Iris iii. p. 201 (1S90) (Valera, Venezuela).
? . Papilio torquatus var. orchantus, Haase, Untersuch. Jliuiicry i. p. 98 (1893) (Venezuela).
6. Uppcfside. — Forewing : yellow sjiots before upper angle of cell small, the
spot before SC'''' especially smaller than in most specimens from Guiana and
the Amazons ; patch SC"' — R' a little longer than the next spot or distally on a
level with it, some specimens approaching the preceding form, bearing a spot before
R^ like tolmides. Hindwing : submargiual spots usually clearly marked.
?. Forewing: a patch across apex of cell; a discal patch R^ — M', usually
accompanied by a patch R^ — R^ and a vestigial patch M' — M-. Four to six
large red patches on hindwing, the last standing behind M- ; a rather large spot
in cell ; tail short, acute, uon-spatulate.
llab. Colombia ; Northern Venezuela.
In'the Tring Museum 28 <? (?, 2 ? ? , from : " Bogota" ; Canauclic, Cundinamarca,
July 1903 (Mathan) ; Muzo, December 1896.
In coll. Oberthiir a series of cj c^ from Juntas, West Coast of Colombia ; in one
of these the upper streak of the subapical patch on the forewing is longer than the
second, and there is also a spot before R^, the specimen resembling the preceding
■subspecies.
40
( 620 )
d. P. torquatus kptalea subsp. uov. (PI. V. fig. 18).
tJ. Upperslcle. — Forewing : spots at iijiper angle of cell miuiite ; streak SC^ — B'
shorter than the next, taj)ering, both narrow, the second streak not extended to
R^ ; band narrower than in all the other forms, being narrower than the black
marginal area. Hindwing : snbmarginal spots clearly marked ; tail spatulate.
Underside. — Forewing : snbmarginal spots rather large, especially the double
spot M" — SJI-. Hindwing : apex of cell black at least as far as base of
R' and M'.
? . Like the female of P. t. orchamm ; spot R^ — R' of forewing and the cell-patch
smaller ; discal and snbmarginal spots R- — R' of hindwing separate, discal jiatch
M-'— (.SM') less enlarged.
Ilab. Western Ecuador.
In the Triug Musenm 1 c?, 1 ? from Naranjas (0. T. Baron) ; Zaruma, lOitU m.
vii. 1899 (Simons).
In coll. Oberthilr several males from Chimbo and Balsapamba, a S from the
latter place being here figured (type of name).
e. P. torquatus torquatus Cram. (1777).
Sfcba, Thesaur. iv. p. 12. t. 7. fig. 21. 22. ^ (17G4).
(J. Papilio Eques Achirus torquatus Cramer, I.e. ; Goeze, Ent. Beytr. iii. 1. p. 8G. n. 65 (1779); Jabl. &
Herbst, Naturs. Schmett. ii. p. 270 (1784) ; ud., he. iii. p. 175. n. 104. t. 45. fig. 5. 6 (1788) ;
Esper, Ausl. Schmelt. p. 148. D. 09. t. 39. fig. 1 (1793).
Papilio Eques Achivutt pelaui Fabricius, Spec. Ins. ii. p. 4. n. \2{\18\) {parti m); Jang, .llphah.
Verz. Schmett. p. 91 (1792) {partim) ; Fabr., Ent. Syst. iii. 1. p. 5. n. 15 (1793) \torquatus quoted
as syn. with ?).
Papilio Eques Trojanus peleiis, Gmelin, Syst. Nat. i. 5. p. 2228. D. 279 (1790) (_partim).
J. Princeps domiaans «i!/rf/«s Hubner, Samml. Exot. Schmett. i. t. 117 (1806 — ?).
g. Ileraclides pelaiis, id., Vcrz. bek. Schmett. p. 83. n. 853 (1818 ?) (partim).
? . Priamides caudius, id., I.e. p. 87. n. 895 (1818 ?).
cj. Papilio torquatus, Godart, Etic. Meth. ix. p. 62. n. 100 (1819) (Guyane ;— " Brazil" alia subsp.) ;
Lucas, in Guer., Diet. Pitt. Hist. Nat. vii. p. 51 (1838).
?. Papilio caudius, Boisduval, Spec. Gen. Lep. i. p. 301. n. 135 (1836) ("Brazil") ; Doubl., Westw.
& Hew., Gen. Diurn. Lep. \. p. 18. n. 191 (1846) ("Brazil ") ; Gray, Cat. Lep. Ins. Brit. Mus. i.
Pap. p. 42. n. 220 (1852) (Para) ; Wall., Trans. Enl. Soc. Land. (2). ii. p. 256(18.')4) (Amazons ;
gardens) ; Gray, List Lep. Ins. Brit. Mus. i. Pap. 59. n. 233 (1856) (Para; Villa Nova) ; Wood,
Ins. Abroad p. 546. 547. fig. 296. 297 (1883) (''jj" false ; "Java" false).
?. Papilio patros Gray, Cat. Lep. Ins. Brit. .Vus. i. Pap. p. 43. n. 221. t. 7. fig. 7 (1852) (Ega) ;
Wall., /,(■. p. 256 (1854) (Upper Amazons ; forest) ; Gray, List Lep. Ins. Brit. Mus. i. Pap. p. 59.
n. 2311(1^56) (Ega) ; Felder, Verh. Zool. Bot. Ges. ll'/cn xiv. p. 310. n. 297. p 358. n. 173 (1864)
(Ega) ; Wood, Ins. Abroad p. 545. fig. 295 (1883).
?. Papilio patros var. a. Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 43. sub n. 221. t. 7. fig. 8 (1852)
(Ega).
? . Papilio jmtros var. b, Gray, I.e. p. 43. sub n. 221. t. 7. fig. 5 (1852) (Ega).
(J. Papilio to7-quatus, Wallace, I.e. p. 255 (1854) (Amazons ; gardens) ; Bates, Natural. Rif. .inixxz.
p. bi (1864) (Pari, in street).
J ? . Papilio caudius, id.. Trans. Ent. Soc. Land. (2). v. p. 1 (1859) (this is the ? of P. torquatus ;
found a pair in copula) ; Butler, ibid. p. 146. n. 230 (1877) (Serpa, Amazons, April).
(J ?. Papilio torquatus, Bates, Trans. Ent. Soc. Lond. (2). v. 347(1861) (caudius and patros ? ? of
torquatus; tony, and «("(/. found in copula); id., Journ. Entoiu. i. p. 22.S. n. 30 (1862) (Pari
and Lower Amazons, abundant ; J var. jinlros Upper Amazons) ; Felder, Verh. Zool. Hot.
Ges. Wien xiv. p. 310. n. 297 (1864) (Surinam, Amazonsi) ; Kirby, Cat. Diurn. Lep. p. 541.
n. 154 (1871) (Suriuain ; Amazons); dberth., Et. d' Ent. iv. p. 78. n. 252 (1880) {partim;
Cayenne; Amazons); Moschl., Verh. Zool. Bot. Ges. Wien xxxii. p. 304 (1883) (Surinam);
Stand., £xot. Ta^. p. 16. t. 11. (J (1884) (Surinam; Cayenne; Amazons); Godm. & Salv.,
Biol. Centr. Amer., Lep. Rhop. ii. p. 228. sub n. 59 (1890) (Guiana ; Lower Amazons) ; Hahnel,
( 621 )
Irh iii. p. 283 (1890) (Pebas) ; Haase, Unlfi-swh. MimJcry i. p. 97 (1893) ; Michael, Iris vii.
p. 213 (1894) (Sao Paulo de OUveiifa).
J . Papllio lorquntus var. a. P. patron, Kirby, I.e. (Upper Amazons).
?. Papilio torquatus V3.T. flavida Oberthiir, El. d'Eiit. iv. p. U5. n. 252 (1880) (Teffie).
cj ?. Papilio patros, Godman & Salv., Biol. Centr. Amer., Lep. Rhop. ii. p. 228. sub n. 59 (1890)
(Upper Amazons).
$ . Papilio toi-quatus var.Jlara, Haase, Untersuch. iliinicry i. p. 98 (1893) (laps. cal. ; "Para" false).
J . Papilio torquatus Tar. caudius, id.. I.e. (Para).
? . Papilin torquatus yht. patros, id., I.e.
There is iu the Tring- Museum from the old collection of Leiinep a male which
agrees with Cramer's figure, and is probably the type specimen. This figure has
several characteristic features which are also found iu that specimen. The band
of the forewing is rather uarrower than usually in the present subspecies, the spots
at the upper angle of the cell are small ; the submarginal spots of the hindwing, on
npperside, are comparatively large, and the apex of the cell is more extended black
on underside than is usually the case.
Since Bates's classical paper in 18(51 the specimens from the Guianas and the
Lower Amazons have generally been regarded as different from the individuals
found on the Upper Amazons. The material examined by us does not bear out
this opinion. However, we find that in the males from Bolivia and Peru there are
usually only five red discal spots on the underside of the hindwing, while there
are mostly six in the specimens from the Amazons and Guiana. Having
unfortunately only one female from the former districts, we do not know whetlier
its peculiarities are merely individual or geographical.
(S. The spots at the upper angle of cell of forewing are mostly larger than in
the other subspecies ; streak SG^^l' shorter than the next, both being broad; the
band broader than in F. t. leptalea and orchamus, varying from being half as
wide again to twice the width of the black distal border. Submarginal spots
of hindwing usually much shaded with black,
Submarginal spots of forewing below generally thin.
? . Tail slender, pointed, rarely somewhat spatulate. There are five principal
forms ; the markings of the hindwing individually variable in number and size.
a'. ?-f. theras nov. Forewing with cell-spot, which does not reach
across cell ; one or more white patches on disc, usually only patch R^ — M' well
developed. Surinam, Upper Amazons.
b'. 2 -f. caudius Hiibn. ,/,<■. — —Forewing without cell-spot; one to three patches
on disc, spot M' — M- being usually the best developed, often alone present, sometimes
no other patches than M' — SM^. The Guianas ; Lower and Upper Amazons.
c'. ?-f. patros Gray, I.e. No white patch on forewing; patches of hindwing
red. Ujjper Amazons ; Cayenne (coll. Oberthiir).
d'. ? -i.Jlatida Oberth. ; patros var. h Gray, I.e. ; Jlava, Haase, I.e. Forewing
without white spots ; patches of hindwing creamy. — —Upper Amazons : Ega.
e. ?-f. cleolas nov. Foi owing without white spots, but with a kind of
submarginal buffish band on underside, consisting of short double streaks R' — R'
and a row of small spots R^ — SM-. Bolivia (Mus. Tring).
IJiib. Orinoco ; Patao, Gairia ; the Guianas ; Amazons ; Eastern slopes of
Ecuador, Peru and Bolivia.
In the Tring Museum 34 cJc?, 13 ¥ ?,from: Patao, Gniria, August 1891 ;
La Vuelta, Caura R., May 1903 (Klages) ; Surinam ; Teffe, January 190.5 (Mathan) ;
Obidos (Mathan) ; .Tuhuty, April 190o (Mathau) ; Itaituba ; Liuitos ; R. Cachyueo,
( (12-i )
affluent of R. Huallag-a (Stuart); ArchiJoua, April 1899 (W. Goodfellow) ; i^amora
(0. T. Baron) ; Pereni R., 3UUU ft. (Watkins) ; Perene K., March 190U (Simons) ;
Upper R. Tore, La Merced (Simons) ; R. Colorado, 2500 ft., September— October
1903 (Watkins & Tomlinson) ; Guanay, Mapiri, 1.500 ft., August 189.5 (Stuart);
Salinas, R. Beni, .July 1895 (Stuart) ; Mapiri ; Eucnentra Grande, mouth of
La Paz R., August 1895 (Stuart); Prov. Sara, S, Cruz de la Sierra (.J. Steinbach).
/. P. torquatus polybius Swains. (1823).
(J. PapHio torqualiix, Godart, Eiic. Meth. i.\.p. 62. n. 100 (1819) (Brazil—" Guyane " alia subsp.) ;
Boisd., Spec. Gin. Lip. i. p. 367. n. 211 (]8:i6) (Brazil) ; Doubl., Lhl Lep. Inx. Brit. .l/(/.<. i.
p. 17 (1845) (Brazil) ; id., Westw. & Hew., Geii. Diurn. Lep. i. p. 16. n. 142 (1846) (Brazil) ;
Gray, Cat. Lep. Ins. Brit. Mus. i. Pup. p. 34. n. 164 (1852) (Brazil) ; id., LUt Lep. Im. Brit.
ilus. i. Pap. p. 46. n. 172 (1856) (Brazil) ; M^aetr., Ennm. On-p. Anim. Miis, Pelrnp.^ Lep. i.
p. 3. n. 51 (1857) (Brazil) ; Prillw., Stetl. Enl. xxvi. p. 130 (1865) (Corcovado).
$. Papilio pohjhius Swainson, Zuol. Illiistr., Enl. ii. t. 94 (1823) (Minas Geraes) ; Doubl., List Lep.
Ins. Brit. Mus. i. p. 13 (1845) (Brazil) ; id., Westw. & Hew., Gen. Diiirn. Lep. i. p. 18. n. 193
(1846) (Brazil) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 44. n. 223 (1852) (Brazil— var. a. alia
spec.) \ id., List Lep. Iits. Brit. Mus. i. Pa}>. p. 59. n. 236 (1856) (Brazil— var. a. excl.) ; M($aL'tr.,
U. p. 5. n. 74 (1857) (Brazil).
? . Troilides tros Hiibner, Samml Exot. Schmett. ii. t. HI (1822 ?).
?. Papilio trojamis Boisduval, Sjiec. Gen. Lep. i. p. 301. n. 134 (18.36) (nom. nov. loco tms Hiibn.
non Fabr. ; Brazil),
?. Papilio tros, Doubleday, List Lep. Ins. Brit. Mus. i. p. 13 (1845) (Brazil) ; Prillw., SirtI.Enl.
Zeit. xxvi. p. 129 (1865) (Corcovado).
cJ ?. Papilio polyhi us, Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 310. n. 297, p. 358. n. 172 (1864)
(Brazil) ; Godm. & Salv., Biol. Centr. Amer., Lep. Rhnp. ii. p. 228. sub n. 59 (1890) (South-
Eastern Brazil ; $ = tros = trojanus).
(J ?. Papilio torquatus, Lucas, in Chenu, Enc. Hist. Nat., Pap. i. p. 38. t. 6. fig. 1. cJ (1851-53)
(Brazil) ; Capronn., Ann. Soc. Ent. Bely. xvii. p. 9. n. 13 (1874) (Entre Rios, Sept.) ; Burm.,
Descr. Rep. Ari/ent. v. Lep., Atlas p. 5. n. 7 (1879) (Rio de Janeiro ; larva, pupa) ; Oberth., Et.
tl'Ent. iv. p. 78. n. 252 (1880) {partim ; Brazil, $ ) ; Maass. & Weym., in Stubel, Reiscn S. A uier.,
Lep. p. 91. n. 40 (1890) ; Bonningh., Verh. Ver. Nat. Untcrli. Ilamhurgix. p. 26 (1896) (common).
(? ? . Papilio torquatus, var. b. P. poh/bius, Kirby, Cat. Diurn. Lep. p. 541. sub n. 154 (1871).
Papilio torquatus ^ va.T. 2>ol!/bi us, Haase, Unlcrsucli. Mimicry i. p. 98 (1893).
(J ? . Troilides torquatus, Kirby, in Allen, Nat. Lihr., Butt. ii. p. 283 (1897) ; id., in Hiibn., Samml.
E.cot. Schmett. ed. ii. p. 97. t. 118. fig. 1. 2, t. 324. fig. 1. 2 (190—?).
c?. Porcwing : the spots before ujjjier angle of cell small ; submarginal spots
of underside rather large. Hiadwing : submarginal spots usually distinct, not
or little shaded with black ; cell below entirely or almost entirely yellow, the black
spot in ape.x being on an average smaller than in P. t. torquatus ; tail broad.
?. Wonomorphic. Forewing : a cell-patch and a large discal patch M' — M-,
no spot R^ — R''', but often a spot R' — M' and a streak behind M- ; tail spatulate,
rounded at tip.
Ilab. Brazil ; Paraguay ; Matte Grosso.
In the Tring Museum 36 c?c?, 10 ? ?,from: Bahia; Minas Geraes (R.Haensch) ;
Tijnca ; Petropolis ; Rio de Janeiro ; Villa Maria to Diamantino, Matto Grosso,
January 1897 (Audeer).
103. Papilio tasso Stand. (1884).
$ . Papilio pohjlius var. a, Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 44. sub n. 223 (1852) (bab. '!)
{J?. Papilio tasso Staudinger, Emt. Ta(xf. p. 19. t. 13. $ (1884) (hab. ?, probably Brazil;
? Brazil).
S . Forewing : band abbreviated, the snbapical si)ots SC* — R- of /'. torquatus
being absent. Hindwing : three yellow submarginal spots on upjjerside, fartli(r
away from margin than in P. torquatus.
( 623 )
? . A broad white band from near R' of forewing to abdominal margin of
hiiidwing ; tail narrow, not spatnlate ; cell of forewing below with some yellow
streaks.
Ilah. Brazil.
104. Papilio peleides Esp. (1793).
Papilio Eqiirs Trojnnus pelaus ?, Jablonsky & Herbst, Naturs. Schmett. ii. p. 205. n. 51. t. 10. fig. 1
(1784) (^>ari/m).
Papilio Eques Trnjanus peleides Esper, Atisl. Schmett. p. 150. n. 70. t. 39. fig. 2 (1703) (copy of
Jabl.'s figure).
Pupil,,, peleiiUs, Boisduval, Spec. Gin. Lip. i. p. 306. n. 200 (1836) (artefact?) ; Doubl., Westw. &
Hew., Gen. Diurn. Lep. i. p. 16. n. 144 (1846) ; Gray, Cat. Lep. Jim. Brit. Mns. i. Pap. p. 34.
n. 106 (1852) ; id., Lixt Lep. In,<. Brit. Miis. i. Pap. p. 47. n. 174 (1856) (S. America) ; Felder,
Verh. Zool. Bot. Ges. Wien xiv. p. 310. n. 301 (1864) (bab. ?) ; Kirby, C,it. Diurn. Lep. p. 541.
n. 154a (1871) (spec. fict. ?) ; Godra. & Salv., Bi(jl. Ceiitr. Amer., Lep. Rhnp. ii. p. 228. sub n, 50
(1800).
Besides Jablonsky's figure and description there is no evidence of the existence
of this insect. Jablonsky expressly states that the figure was carefully drawn from
a specimen. The individual may have been an artefact, as suggested by Boisduval,
but the figure does not give us that impression. Considering that of many American
I'apilios only very few specimens are known, it is quite conceivalile that P. pdeides
has not been rediscovered, as is the case also with F. tasso, which was described
from an old pair without precise indication of locality.
Ilab. Presumably South America.
Subsection D.
Palpus with a white (rarely yellow) dot, sometimes quite black. Frons black,
or with a yellow central line, never yellow along eyes. Costal margin of forewing
somewhat incrassate, more or less dentate, the scaling of the edge easily rnbbed
off, with the exception of the Zagreiis Group.
The Zagreus Group is a mimetic offshoot of this subsection, having acquired
several peculiarities which render it difficult to give a general characterisation of the
whole Subsection inclusive of the Zagreus Group. Apart from these mimics, the
species of the present Subsection are generally strong-winged species with heavy
nenration in the forewing. Black is the prevailing colour on the body and wings.
The abdominal margin of the hindwing is clothed with long hairs in the males.
Key to the gronjis :
A. Costal margin non-serrate ; frons with yellow mesial line ;
abdomen for the greater part orange-yellow . . . Zagreus Group.
B. Costal margin almost smooth, the serration being vestigial . Scamander Group-
C. Costal margin .serrate Ilomerus Group.
X. Zagreus Group.
Frons witii yellow mesial line. Antenna long, very slender, with thin club,
yellow, except proximally. Breast obliquely striped with yellow ; abdomen orange-
bufF, with a black mesial stripe above and below, or only on one side. Distal
margin of forewing convex, farthest point about R', costal margin non-serrate ;
cell very broad, upper and lower angles obtuse ; SC' usually before angle of cell ;
stalk SC'-" very variable in length. Hindwing ovate, longest centrally ; long-
( 624)
hairy at iibdominal margin in male ; basal cellule long and narrow, spur near its
apex, SC near its base ;; SC" from before middle of cell. Tenth abdominal tergite
of male bifurcate ; harpe long, gradually tapering to a point, somewhat flexuose,
similar in the three species. Spines on upperside of tibiae and tarsi few iu number
and short in both sexes.
Key to the species :
a. Hindwiug black for the greater part Species No. 107.
Hindwing with large central orange area .... d.
b. Hindwing with a black spot iu cell and several around cell . Species No. 10.5.
No such spots on hindwing Species No. 106.
105. Papilio zagreus Doubl. (1S4T).
Pajjilio sai7)-f»sDoubleday, Ann. May. N. II. xix. p. 174 (1847) (Venezuela) ; id., Westw. & Hew.,
Gen. bliim. Lep. i. t. 1. fig. 1. $' (1847) ; iid., I.e. ii. p. 529 (1852) ; Gray, Cut. Lep. Ins. Brit.
Mtis. i. Pap. p. 8. n. 30 (1852) (Venezuela ; " Quito " alia species) ; id., Li.-^i Lep. Ins. Brit.
Mu.^.i.Pap. p. 9. n. 33 (185t;) ; Bates, Trans. Ent. Soc. Land. (2). v. p. 349 (1861) (Ega,
November, one example) ; id., Joiirn. Ent. i. p. 229. n. 35 (1862) (Upper Amazous) : Felder,
Verh. Ziml. Bot. Ges. Wien .\iv. p. 312. n. 322 (1864) (Bogota ; Venezuela ; Ega) ; Kirby, Cat.
Diurn. Lep. p. 538. n. 142 (1871) ; Druce, Proc. Zonl. Soc. Land. p. 246. D. 17 (1876) (Pozuzo) ;
Hopff., Stett. Ent. Zeit. xl. p. 53. n. 24 (1879) (N. Granada, Venezuela, Amazons, Bolivia) ;
Obertb., Et. d'Ent. iv. p. 99. n. 312 (1880) (Guayaquil) ; Stand., E.t<it. Tagf. p. 15. t. 10. ^
(1884) (South Peru to Venezuela) ; Dognm, Lip. Loja p. 15 (1887); Hahnel, Iris iii. p. 149
(1890) (San Estt^ban) ; Dognin, I.e. p. 37 (1891); Haase, Untermch. J/Zmwy i. p. 94 (1893) ;
Kirby, in Allen's Nat. Libr., Lep. Butt. ii. p. 284 (1896) ; Eimer, Orthogen. p. 209 (1897).
(? ? . In spite of the large area inhabited by this species there is no decided
geographical variation. The individual variability in the size of the creamy and
orange markings of the forewiug and in the black spots of the hindwing being
considerable, all the differences between specimens from northern and southern
localities which may be discovered in a small series of individuals disappear when
a larger number of specimens is compared. The only difl'erence which ajiproaohes
constancy is iu the colour of the cell-bar of the upperside of the forewiug, this bar
being pure cream-colour in Colombian and Venezuelan males, rarely showing a
trace of orange, wliile it is more or less orange at the discal end in the majority
of the specimens from Ecuador, Pern and Bolivia. The black dorsal stripe of the
abdomen is absent from the female ; in one of our Ecuadorian males the black
ventral stripe of the abdomen is wanting.
Genitalia : i. The processes of the tenth tergite long, similar to those of
P. backus, being much longer than in P. ascolius. Harpe long, very narrow, the
apex being curved inwards. ?. Lobe situated in front of the vaginal orifice
short, rotundafe-f ruucafe ; lateral flap very large ; transverse fold or low ridge
situated behind vaginal orifice continued laterad, disappearing on the inner side
of the lateral flaps.
Early stages not known.
IJa/j. Venezuela, Colombia, southward to Bolivia.
In the Tring Museum 47 c?c?, 1 ? , from : Onaca, S. Marta, 2000 ft. (Chas.
Engelke) ; Villavicencio to Monte Eedondo, March— Ajiril 1807 (Dr. Burger) ;
Viliaviccncio to R. Ocoor, January 1897 (Dr. Biirger) ; "Bogota"; Zamora
(0. T. Baron); Chanchamayo, Peru (Schunke) ; Pozuzo, Hudnuco, 800— lOUO m.
(\V. Hoflmanus) ; La Union, 11. Huacamayo, Carabaya, 2000 ft., November 1904
(G. Ockenden) ; Reyes, E. Beni, August 1895 (Stuart) ; Guanay, Mupiri R.,
August 189.5, 1500 ft. (Stuart).
( 025 )
106. Papilio ascolius Feld. (1865).
Papilio znrjreus, Gray {nnn Doubleday, 1847, err. det.), Cal. Lep. Ins. Brll. .l/«s. i. Pnp. p. H. ii..30
(1852) (pnrtrm ; Quito, in coll. Hewitson).
Papilln aHndiiix Felder, Verli. Zool. Bot. Gfs. Wimi.'xiv. p. 312. n. 323 (1SC4) (nom. iudescr. ; Quito ;
Bogota) ; id., Pelsc Nuvam, Lep. p. 82. n. G3 (1865) (Bogota, ^) ; .Staud., Exot. Tag/, p. 15
(1884) (Ecuador ; Colombia; Chiriqui).
cJ ? . Differs from P. zagreits especially in the Iiintlwing, the basi-discal area
bearing no black patches in the cell and between W and abdominal mar<;-in ;
besides the black subcostal streak, which is always present, the wing bears only
one black discal spot SC-— R' either on both sides or only below, this spot being
absent from many specimens; the black distal border of the hindwing is usually
broader than in P. zagreus, often touching cell, but in Panama individuals the
border does not surpass in width that of P. zagreu.'i. The abdomen of the female
is on the back either entirely ochraceous or blackish brown, there being no sharply
defined dorsal black mesial stripe as in the male.
Genitalia : S. The processes of the tenth tergite short, the dorsal lateral edge
of the tergite irregnlarly dentate and sinuate, the two sides not being exactly
identical ; harpe flattened beyond middle and here broader than in P. zagreus, the
apex straight, not curved inward. — ? . Lobe at vaginal orifice slightly acuminate,
narrower and longer than in P. zagreus ; lateral flap not so large as in that
species, its posterior edge continuous with the low transverse ridge standing
behind the vaginal orifice.
Early stages not known.
Hab. Chiriqui to Western Ecuador.
The four snbspecies differ only in pattern, each form varying considerably.
There are in a long series of species all intergradations from one snbspecies to the
other.
a. P. ascolius zalates Godm. & Salv. (1890).
Papilio zalates Godman & Salvin, Biol. Ceiilr. Amer., Lep. Pimp. ii. p. 233. n. 68. t. 71. fig. 1. 2. cj,
3. genit. (1800) (Chiriqui).
<?. Upperside. — Forewing : cell-patch more or less densely shaded with black ;
subapical cell-spot narrow ; discal spots short, spot R' — R- as large as the next
one or larger ; no spots R' — M- at cell or only traces of them. Hindwing :
distal marginal border narrower than in the other forms ; no black discal spot
SC2— Ri.
Underside. — Hindwing, deeper orange than in the other forms ; snbmargiual
spots smaller ; no black spot SC^ — R'.
? not known.
Hab. Panama; Chiriqui; Bugaba ; Veragna.
In the Tring Museum 3 c? c? from Chiriqui.
b. P. ascolius daguanus subsp. nov.
c?. Upperside. — Forewing : basal cell-patch as in zalates, subapical cell-jiatch
as in zalates or larger ; discal spots as in ascolius, spot R' — R- large, but much
shorter than the next one ; slight traces of spots R= — M- at cell. Hindwing :
basi-discal area of the same colour as the spots of forewing, slightly washed with
orange distally, but much less so than in the palest specimens of ascolius, agreeing
in this character best with pale individuals of rosenbergi from West Ecuador ;
black subcostal streak broader than in zalates and ascolius, invading cell a little ;
( 626 )
a large black discal spot SC- — R', almost tonching the subcostal streak ; creamy
spots around apex of cell nearly as large as in zalatcs.
Underside. — Basi-discal area of liindwing nearly as pale as above, orange-tawny
at costal margin and distally ; black subcostal streak broader than in the other
forms, washed out at the edges, entering cell ; black discal spot SC- — R' triangular,
larger than in the other subspecies.
Hub. West Colombia : Bio Dagua.
In the Tring Museum Z S S-
A long series of males in coll. Oberthi'ir from Jnntas, R. Dagna.
c. P. ascolius ascoliiis Feld. (1865).
PiipiVio nariiliiis Feider, l.c. {partim ; Bogota); Kirby, Cal. Diurn. Lep. p. ."iSS. n. 14-'b (1871);
Oberth., El. iTEnl. iv. p. 99. n. 313 (1880) (Colombia) ; Staud., l.c. (18H4) (jmrllm) ; Eimer,
Orlhngen. p. 209 (1897).
<?. Basal cell-patch of forewing always clearly rnarked, snbapical cell-patch
narrow or broad, occasionally continuous with the basal patch posteriorly ; discal
spot R' — R-' usually as large as the last discal spot, rarely a mere dot ; the discal
spots very variable in size, in some specimens only one-third the size as in others ;
the spots R^ — M- at cell never ab.sent, but often small, sometimes joined to the
discal spots. Hindwing always orange in cell and beyond ; the orange spots
around apex of cell very variable, spot R- — R^ often absent, sometimes both
spots R- — M' mere specks ; black discal spot SC- — R' often vestigial, diffuse.
Underside. — The discal spots situated distally of cross-veins on forewing very
rarely reduced to dots. Basi-discal area of hindwing usually pale in middle, but
sometimes the pale colour reduced to some traces situated at the black subcostal
streak ; black discal spot SO- — R' present in most specimens.
? . A specimen in Mr. Godman's collection from Mnzo. Abdomen black-brown
above ; proximal cell-patch of forewing and patches between cell and hiiidmargin
orange; patches R^ — (SM') extended to cell; a broad streak along hindmargin.
Black subcostal streak of hindwing reduced.
Ilah. Colombia : Magdalena valley ; Cordillera of Bogota.
In the Tring Museum 27 cJc? from : "Bogota"; Muzo, December 1896.
d. J', nscoliits rosenben/i Druce (10O3).
PapiV.o ziigreiis. Gray, I.,-, {paiiiin ; " Quito ").
PapiVio iiH-iiVius Feider, I.e. (partim ; Quito).
I'dpiliu roxenliergi Druce, A)i>i. Mag. ]V. [I. (7). xii. p. 221 (1903) (Paramba).
i. Ujiperside. — Forewing, subapical cell-patch large, more or less rounded
distally ; s])ots R^ — M'- situated at cell absent or well marked; discal spot \V — R-,
which is nearly always large in the other forms, usually minute, sometimes absent,
seldom as large as the last spot of the discal row; discal spots R^ — M' usually longer
than in the other subspecies; submargiual spots R' — SJl- as a rule smaller than
in the preceding form, often vestigial, the posterior ones being sometimes absent.
Hindwing, basi-discal area paler than in ascolius, usually hardly at all washed
with orange, but sometimes more extended orange than in the palest specimens of
P. a. ascolius ; width of black distal border as in ascolius, variable ; black discal
spot SC^ — R' mostly absent.
Underside. — Subajjical cell-patch of forewing rounded distally. Basi-discal
area of hindwing either pale in middle, or the pale colour marked only at the black
subcostal streak; discal spot SC-— R' varying from black to tstwny, often vestigial,
( 627 )
never nbseiit ; the Lliick streak usnally washed out at the edges ; submarginal spots
small.
¥. Abdomen ocliraceons above. Discal patches W — M° of forewing more
proximal than in male ; spots M' — (SM') situated at cell large but ill-defined.
JJa/j. "West Ecuador.
In the Tring Mnsenm 11 cJtJ, 1 ?, from : Paramba, 3500 ft., March, April ami
May 1897 ; and (Jhimbo, lOoO ft., August 1897 (W. Rosenberg).
107. Papilio bachus Feld. (1865).
PapiUo bachus Felder, Verli. Znoh But. Ges. Wieii xiv. p. 312. n. .^24(1864) (nom. indescr, ; Bogota);
id., Rehe Nomra, Lep. p. 80. n. 62. t. 14. fig. a. b. (J (1865) (Bogota).
cJ. The two cell-patches of the forewiiig are usnally confluent behind, the
black space at the costal side of the proximal patch being often reduced to a
small snbbasal streak ; the discal sj)ots SC — R- are always present, the upper
two being sometimes very small, the third reaching occasionally from cell halfway
to outer margin ; three long patches from cell to near outer margin, the second and
third often imperfectly divided into a proximal portion corresponding to the spots
R^ — M- situated in P. ascolius and zagreus close to the cell, and a distal portion
homologous to the discal spots of the allied species ; the submarginal series of
spots usually absent from upperside, but occasionally the sj)ots partly distinct
partly vestigial, the last three spots R^ — SM- often rej)resented by minnte dots.
The hindwing is quite black, except a narrow band separating the basi-discal area
from the marginal border, the latter being narrower than in the other species ; that
orange band often reduced to some small spots ; creamy submarginal spots distinct;
the veins partly creamy in some specimens.
The underside is somewhat paler than the upper ; the hindwing is more
extended tawny-orange, the black area being often divided np into patches by
the veins being bordered with tawny. In one of onr specimens of the southern
subspecies (Pozuzo, Peru) the marginal baud of the hindwing is creamy for the
greater part, sharply defined sj)ots situated in the middle of the cellules between
the blue dots being black and therefore very conspicuous on the pale ground ; the
same specimen bears on the forewing, above and below, ill-defined creamy streaks
at the distal margin from SC to R', the first being the longest and most indistinct.
? not known.
Genitalia : S. Tenth tergite similar to that of P. zagreus ; the two apical
projections long ; harjie also as in that species, a little broader beyond middle.
Ilah. Colombia, Peru and Bolivia.
Two subspecies.
a. P. bachus bachus Feld. (1,865).
PapiUo bachus Felder, I.e. ; Kirby, Cat. Dhu-n. Lep. p. 538. n. 142a (1871).
(?. Markings of forewing pale above and below; proximal cell-patch and
posterior discal patches washed with orange ; discal streaks R- — M- narrowed
distally on upper- and underside ; vestiges of three submarginal spots SC^ — R',
the first spot standing distally of the fork. Veins of hindwing partly creamy.
Apparently rare, at least in collections.
Ilab. Colombia.
In the Tring Museum 3 tJJ from : " Bogota" (Lindig, type) ; Villavicencio to
Monte Redondo, March— April 1897, beginning of rainy season (Dr. Burger).
( 628 )
b. P. backus chrysomelus subsp. nov.
rupUio harlnis, Staudinger, Exot. Tagf. p. 15 (1S84) (Peru) ; Weeks, Ilhislr. Diurii. Lep. p. 20
(1905) (Chulumani, Bolivia).
? . Basi-iliscal area of forewing orarifre above and below, more or less creamy
yellow at costal margin ; discal streaks R- — M- distally less acuminate on underside
than in the preceding subspecies.
The individual variability is considerable. Some specimens have hardly any
cream-colour on the upperside, while in others the costal half of the subapical
cell-patch, the patches outside the short cross-veins and the distal portion of the
streak R^ — R^ are creamy. The amount of tawny orange on the hindwing is
very variable.
Hab. Pern ; Bolivia.
In the Tring Museum 24 cjc? from: Poznzo, Hn^nnco, 8U0— lUOO m.
(W. HoflFmanns) ; Chanchamayo (Schunke ; Hoffmanns) ; Montanas, R. Madre
de Dios, September 1901 (Oekenden) ; R. Slucuri, S.E. Pern, June 1901, 2500 ft.,
dry season (Oekenden) ; La Union, R. Huacamayo, S.E. Pern, 2000 ft., November
1904, wet season (Oekenden) ; R. Songo to R. Suapi, Bolivia, 1100 m., March — June
(Garlepp) ; Salinas, R. Beni, June 1895 (Stuart); Mushay, R. Beni, August 1895
(Stuart) ; S. Augnstin, R. Mapiri, 3500 ft., September 1895 (Staart) ; Charuplaya,
May 1901, 1300 m., some rain (Simons).
XI. Scamander Group.
Dentition of costal margin of forewing vestigial.
Of all the species of Subsection D a member of the presnt group, namely
P. hellanichus, appears to us to stand in pattern nearest the ancestral form.
The mimetic Zagreus Group is probably an ofiPshoot from some such insect as
P. hellanichus.
The dentition of the costal edge of the forewing is very feebly developed, but
on denuding the margin small teeth become visible, being especially distinct
towards the base of the wing.
Key to the species :
a. A yellow band of spots across the upperside of both wings b.
No yellow band on upperside ...... r.
b. Hindwing, upperside, with large yellow spot in cell, discal
spots more or less orange distally .... Sjiecies No. 108.
Band of hindwing, ujiperside, not entering cell, or cell-spot
very small Species No. 109.
c. Abdomen with large creamy side-band or patch . . Species No. 111.
Abdomen black Species No. 110.
108. Papilio hellanichus Hew. (1868).
Papilio hellanichus Hewitson, Exol. Butt. iv. Paj). t. 9. fig. 27. 28 (1868) (Uruguay) ; Kirby, Col.
Dlurn. Lep. p. 5C6. n. 323 (1871) (Uruguay) ; Oberth., El. d'Eiit. iv. p. tj'.l. n. I'JG (1W80)
(Brazil); Haase, Untcmuch. Mimicry i. p. 92 (18y3) (belongs to the •' Machaon Group");
Eimcr, Artb. Vcrxi-dmllseh. Schmett. ii. p. 138. t. 7. fig. 5. (J (1896) (near /'. aii,ericu.'<) ; id.,
Orthogen. p. 37 (1897) ; Christ, Milt. Schweiz. Ent. G«.i. ix. p. 272 (1897).
Papilio hellauicus (!) Hewitson, l.r. Itulex (1871).
Papilio cleotas, Burmeister (n™ Gray, 1832, err. det.), Dc^ci: Rep. Ardent, v. p. 61. n. 3 (1878)
(mouth of R. Paranii ; Entre Rios ; Rrsario ; Gualeguaychu ; Cordova ; Las Couclias ;
P. hellanirux ! sub sjn.).
( 629 )
(? ?. With the exception of Burineister, all <atithors have regarded this insect
as a near relative of P. americus and allies. However, it has nothing to do with
the Machaon Group, the resemblance to those insects being quite superficial. We
are astonished to find that the pattern of the upperside deceived even Haase, who
in many other cases has shown such a keen insight in the true relationship of man}-
species of Papilio over which other authors had blundered. F. hellanickus is in
structure practically identical with P. scamander, and the close agreement in pattern
will also become at once evident to every one who compares the underside of
P. Iiellanichus with that of P. scamander scama)tder. In fact, P. kellaiuelius is
nothing else but the most southern development of P. scamander ; the two insects
may be specifically distinct, as we believe they are, but are nevertheless very
closely related. The patch in the cell of both wings gives the species an
americiis-Wlie appearance ; but even these patches are nothing new in the species,
as the cell-spot of the forewing occurs on the underside in P. scamander, while the
hind wing of P. scamander often bears a small cell-spot on both sides.
Early stages not known.
ITab. Uruguay and the adjacent districts of Brazil and Argentina.
In the Tring Museum 10 SS, 3 ? ?, from: S. Isidor, north of Buenos Aires
(Rnscheweyh) ; La Soledad, Entre Rios, border of Uruguay, October, December,
January (Chas. Britton) ; La Soledad, March and December (Miss E. A. Britton).
In coll. Oberthiir from Zarate, January and February 1881 (Kinkelin), and
a ? labelled " Brdzil."
109. Papilio scamander Boisd. (1836).
Papilio scamander Boisduval, S^wc. G/n. Ltp. i. p. 363. n. 20G (1836) (Brazil).
S ? . Upperside. — Forewing : costal edge practically non-serrate ; no spot in
cell ; a curved discal row of rounded spots from costal to hinder margin ; a
submarginal row of much smaller spots from costal margin, not reaching to hinder
margin. Hindwing : a curved discal band of spots parallel to distal rnargin
situated outside cell, often touching it, there being sometimes a small spot in cell,
cream-colour or buff-yellow like the bands of the forewing ; a row of sabmarginal
spots, red or pale orange-buflf, the first ones being sometimes cream-colour ; tail
thin, tooth M' 2)rodnced, sometimes also tooth M-.
Underside of forewing similar to upper, the cell bearing sometimes a spot
at apex, and the submarginal spots being enlarged in one of the sub.^pecies.
Hindwing : ground-colour either brown-black, or yellowish buff, or intermediate, the
three subspecies being different in the colour and pattern of the wing.
Neuration : D- of forewing half the length of D'^ or less ; SC- of hindwing
midway between base and R' or beyond middle.
Genitalia: S. Tenth tergite long, spatulate, the tip being rounded; upper
edge of sternite strongly chitinised, smooth, bearing anally two processes, the
anterior one being conical, pointed and proximally hairy, the posterior one being
subcylindrical, feebly increasing in width apically. Harpe gradually widening
from base, somewhat twisted, suddenly narrowed at apical third of clasper from
the ventral side, ending in a slightly curved acnte process, the ventral edge bearing
one or more small teeth proximally of the apical process. ? . In front of vaginal
orifice a lanceolate, flat process, slightly irregular at the edges, carinate on the
anterior surface ; lateral edges of orifice feebly elevate, forming posteriorly together
( G30 )
a short, flattened, longitudinal ridge whicli bears a rounded tubercle behind the
orifice, the tubercle being covered with extremely small hairs ; the whole area
at each side of the orifice brown, chitinised, the lateral edge being free, projecting ;
between this ridge and the postvaginal mesial ridge a deep groove in front of which
there is a long, pointed, curved process.
Early stages several times described and figured ; see literature under graiji.
Hab. Brazil.
Three subspecies.
a. P. scaniandei- grayi Boisd. (1836).
Vuiuho f,rm/i Boisduval, /.-■. p. 365. D. 208 (183(5) (Brazil) ; Felder, Verh. Zool. Bot. Ges. Wien xiv.
p. 313. n. 3,S8 (lS3i)) (Bras, austr.) ; Lucas, in Gui'r., Dirt. Pitt. Hist. Nat. vii. p. 51 (1838) ;
Doubl, List Lep. /«.«.-. Ihil. Miis. i. p. 18 (184,o) ; id., Westw. & Hew., Gen. Diiini. Lep. i. p. IG.
n. 151 (184G) ; Gray, Cat. Lep. Ins. Brit. M,:s. i. Pap. p. 35. n. 171 (1852) ; Lucas, iu Chenu,
Ehc. ma. Nat , Pap. i. t. 16. fig. 2. (J (1853) ; Gray, List Lep. Ins. Brit. Miis. i. Pap. p. 47.
n. 1711 (1856) (Brazil) ; MentHr., Eniiiii. Corp. Anim. Miis. Petrnp., Lep. i. p. 4. n. 53 (1857)
(Brazil) ; VoUenh., Tijdsch: Ent. iii. p. 87. n. 145 (1860) (Brazil) ; Kirby, Cat. Diurn. Lep.
p. .5.37. n. 130 (1871); Burm., Descr. Pep. .Irpent. v. Leji., Atlas p. 5. n. 9. t. 2. fig. 4. 4a (187>.0
(larva, pupa; Nova Friburgo ; Petropolis) ; Oberth., Et. iVEnt. iv. p. 74. n. 223 (18S0)
(jmrlim ; type) ; .Tones, Pnn: Lit. Phil. Snc. Liverpool xxxvi. p. 42. n. 45. t. 4. fig. 10 (IK83)
(larva ; pupa ; Sao Paulo) ; Biinningh., Verli. Ver. Nat. Unterh. ffamhiirg ix. p. 28 (18%) (not
at Rio ; common at Petropolis ; larva on Canelhi and Magnolia) ; Peters, Illiislr. Zeitsehr.
Ent. ii. p. 52 (1807) (Bahia, ratber common ; larva, on Laurus).
(J ? . The row of submarginal spots of forewing not angulate ; posterior discal
spots larger than anterior ones. Submarginal spots of hindwing red, sometimes
several red spots on disc outside the yellowish buff band.
Basal area of i/m/erside of hindwing a little paler than forewing, but never .
washed with buff; a row of red spots distally of and separate from the creamy
white discal band.
IM. Bahia to Parana, southern specimens often similar to the next form.
In the Tring Museum 22(?(?, 10 ? ? , from : Espiritu Santo; Petropolis,
October 1897 and January 1898 (Foetterle) ; Sao Paulo; Castro, Parana (E. U.
Jones).
b. P. scamander mnjmander Ilopff. (1866).
PapiUo eurymander Hopffer, Stelt. Ent. ZeU. xxvii. p. 29 n. 10 (1866) (Brazil) ; Kirby, Cat. Diurn.
Lep. p. 567. n. 335 (1871).
Papilio gra;ii,Mu\\eT. Ko.mos p. 187 (1878) (agreeable odour in $) \ id.. Trans. Ent. Soe. Lonil.
p. 219 (1878) (variability) ; Oberth., I.e. (1880) (jiarliw ; Rio Grande do Sul).
S ? . Discal band of forewing wider than in the preceding, of about even width
throughout, or the anterior spots larger than the posterior ones ; submarginal row
of spots angulate, the upper two or three spots being more proximal than the
others. Band of hindwing on the whole more widely separate from cell; at
least the upper submarginal spots creamy-yellow.
Forewing occasionally with spot in eel! on underside. Basal area of hindwmg
paler than in the preceding subspecies, often washed with buff; red discal spots
smaller, closer to the band, often obsolete ; the blue bars beyond them more distuict ;
submarginal spots paler. Abdomen sometimes spotted with buff at the sides.
Hab. Santa Catharina ; northern districts of Rio Grande do Sul.
Intergrading with the next.
In the Tring Museum 6 cJc?, 4 ??, from: S. Catharina: Theresopolis,
800—1000 ft., November 1904 to February 19(15 (J. Michaelis).
( fi3l )
C, p. scamander scamamler Boisd. (1830) (PI. V. fig. 22).
Papilio scamamler Boisduval, I.e. (Brazil) ; Doubl., Westw. & Hew., Geii. Diiirn. Lcp. i. p. 10.
n. UG (1846) ; Gray, Cat. Lcp. Lis. Brit. Mas. i. Pap. p. 35. n. 168 (185i!) ; Lucas, ia Chenu,
Enc. Hist. Nat. Pap. i. t. 16. fig. 1. ^ (18.51-3) ; Gray, List Lep. Lis. Jhil. .)liis. Pap. i. p. 47.
E. 176 (1856) : Lucas, in Casteln., Vay. Amir. Sud, EiU. p. 109. t. 1. fi;,'. 2. ^ (1S57) (Interior
of Brazil) ; Felder, Yrrh. Zn„l. But. Ges. Wien xiv. p. 313. n. 339 (1864) ; Kirby, Cat. Diurn.
Lep. p. 537. n. 129 (1871) ; Oberth., El. d'Ent. iv. p. 73. n. 222 (1880) (type).
Papilio grayi var. scaimiiider, Mabilde, Gaia Pract. Borbol. Rio Grande do Sul p. 45 (1896).
S ¥ . Discal band of uppersidc more yellow than in the other forms ; on
forewing wider in front than behind, spots SC'" — R- being the longest, the band
posteriorly more distal than in the other two subspecies ; submarginal row
angulate. Hindwing : snbmarginal spots buff-yellow, the posterior ones often
washed with red.
Underside very different from that of tlie other forms. Forewing : anterior
submarginal spots enlarged to large patches which extend close to margin.
Hindwing : basal area the same creamy yellow colour as discal band, the veins
remaining black ; no red discal spots ; blue crescents conspicuous ; submarginal
spots varying from being mili^y white to being red.
Hab. Eio Grande do Sul, some specimens agreeing closely with extreme
specimens of eurymander.
In the Tring Museum 2 c? <^, 1 ? , from Rio Grande do Sul.
110. Papilio birchalli Hew. (1803).
Papilio birchalli Hewitson, Trans. Eiit. Soc. Lnnd. (3). i. p. 517 (1863) (Bogota).
(?. Abdomen black, claspers more or less creamy at sides. Costal edge of
forewing practically non-serrate ; some indistinct or small spots around apex of
cell and a postdiscal and a submarginal row of spots creamy, the upper jiostdiscal
spots more or less shadowy. Hindwing: a broad creamy discal band, slightly
greenish as all the markings of the upperside, narrowing behind, entering cell,
the cell-spot varying in size, being sometimes very small ; a snbmarginal row of
olive-buff sjjots, between which and discal band there is occasionally another row
of small spots ; no tail, tooth R^ being very little longer than the other teeth.
On underside of forewing a greenish creamy cell-patch, and from lower angle
of cell to hindmargin a row of greenish creamy patches ; four submarginal spots
from R' })ackwards, sometimes a fifth vestigial. Discal band of hindwing
consisting of a rather large costal creamy spot and a similar subanal spot con-
nected with one another by a row of minute red spots, some of which are often
vestigial, spot M' — M- being creamy ; submarginal spots red, halfmoon-shaped,
the first alone being usually straight, spot M'— BI- being the largest.
? . Similar to male, but the markings of the upperside more bluish green and
the discal baud of the hindwing much wider. On the underside the costal spot
of the discal band of the hindwing is red and smaller than in male, while the small
intermediate spots are larger than in male.
Neuration : SC- of hindwing at one-third (or a little beyond) from subbasal
cellule to R'.
Genitalia: S. Tenth tergite long and broad, spatulate ; double process of
sternite not deeply divided, the two projections short, the anterior one slightly
more acuminate than the second ; harpe very long, tapering, acute, dorsally
augulate, and bearing between this angle and tlie apex many small teeth, the
( 632 )
iijiical portion of the barpe somewhat resembling a shark's tail. ■?. Edge of
vagiual orifice anteriorlj- raised into a rather broad, deeply sinuate flap, each lobe
of which is pointed ; lateral edges of orifice continned beyond the orifice as a mesial
di)ul)le fold; behind the orifice a ronnded, densely folded, tubercle, which is glabrous,
bearing minute hairs only posteriorly at the base ; in front of this tubercle, within
the orifice, there is a transverse ridge ; behind the orifice there is on each side
of the double mesial fold a deep impression, at the side of which there stands a
long, pointed, somewhat scythe-shaped process ; the whole lateral area of the
vaginal depression strongly chitiuised, the edge projecting free, being continuous
with the anterior surface of the sinuate vaginal flaj).
Early stages not known.
Eab. Panama ; Colombia ; Argentina.
The last locality requires confirmation.
Two subspecies.
a. P. birchalU godmani subspec. nov.
rapilio hirchalll, Godman & Salvin, Bio!. Ceulr. Amer., Lep. lilinji. ii. p. 2,37. n. 74. t. 71. fig. 8. 9. <^,
10. geiiit. (1890) (parltm ; Chiriqui ; Bugaba).
This form stands in some collections as godmani ; wc fail to find the description,
but accept the name.
tj. Discal band of upperside of hindwiug posteriorly obsolete, anteriorly broader
than the black distal marginal area.
? . Uiscal baud of hindwiug very broad, including apical third of cell, irosteriorly
irrorated with black, the greenish scaling not being dense.
Genitalia: c?. Harpe angulate at apical two-fifths. ?. As described
above.
Hab. Panama : Chiriqui ; Bugaba.
In the Tring Museum 2 t?c?, 1 ?, from Chiriqui (received from Messrs.
Staudinger <& Bang-Haas).
b. P. birchalU birchalU Hew. (1863).
Papilio birchalU Hewitson, I.e. (Bogota); Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 312, n. 327
(18G4) ; Kirby, Gil. Dinni. Lep. p. 538. n. 13fl (1871) (paiiim) ; Staud., Exol. Tagf. p. 15 (1884) ;
Obeith., Fa. (XEni. xii. y. ?,. t. 2. fig. Ci.$ (1888) (Muzo) ; Godm. & Salv., Binl. Centi: Amer., Lep.
Ulinji. ii. p. 237. n. 74 (1S'.I0) [partiiu ; Colombia ; Cauca valley ; R. Quarto, Cordova,
Argentina) ; Maass. & Weym., in Stiibel. Reiseii S. Amer., Lep. p. 11. n. 39 (189U) (Colombia) ;
Haase, Unterstich. Mimirri) i. p. 94 (1893).
c?. Discal band of hindwiug narrower than in the preceding, extending to
abdominal margin ; cell-spot smaller.
? not known.
Genitalia : Harpe somewhat shorter than in the preceding, dorsally angulate
at ajiical fourth.
Ilah. Colombia : Magdalena and Cauca valleys.
In the Tring Museum 7 c? c? from " Bogota."
111. Papilio xanthopleura Godm. & .Salv. (18U8).
Papilio xanthopleura Godman & Salv., Ann. .Vag. N. II. (4). ii. p. 150. n. 25 (18G8) (R. Huallagn) ;
Hew., Exot. Butt. iv. Pap. t. 10. fig. 32. ^ (1869); Kirby. Cat. Diurn. Lep. p. 538. n. 140(1871);
Druce, Proc. Zool. Soc. Land. p. 246. n. 19 (187G) (Lower Huallaga) ; HopfiE., Sictt. Ent. Zeit. xl.
p. 53. n. 25 (1879) (Peru) ; Godm. A Salv.. Binl. Centr. Amer.. Lep. Rhop. ii, p. 238. sub n. 74
( 633 )
(1890) (smooth costa) ; Hahnel, fm- iii. p. 21)7 (180O) (Iquitos) ; Staud., ibid. iv. p. 63(1891)
(Iquitos ; S. Thomar, R. Negro ; S. Paulo de Oliv. ; ^J, ? ); Haase, Untersuch. Mimicry i. p. 94
(1893).
J. Papilio xanthojAeura Salv. & Godm. var. tl/njiJium Staudinger, I.e. (1891) (Miinicoru ; also Sao
Paulo de 0Iiven9a ace. to Michael).
iS ?. Thorax with short yellow mesial stripe, correspoudiug to the mesial stripe
of F. zagreus and allies. Abdomen with broad creamy yellow lateral strii)e.
Costal edge of forewing practically non-serrate. Disc of hindwing abode with
greyish green or greyish blue patches. No discal band on underside of hindwing ;
a creamy spot near anal angle being all that is left of the band ; red submarginal
spots large. The female is dimorphic :
a'. ?-f. xanthopleura is similar to the male, the bluish area of the hindwing
being larger.
b'. ? -f. diaphora Stand., I.e., bears a large creamy yellow central area on the
forewing occupying the greater part of the cell and the disc between the median
and lower radial veins, the bluish scaling of the hindwing being reduced to a
narrow band.
Nenration : SC- of hindwing at one-third from base to R^
Genitalia: c?. Tenth tergite long, strongly narrowing distad, rounded-dilated at
the apex ; sternite on each side with two processes, the anterior one triangular,
multidentate, the posterior one rather longer, narrower, slightly acuminate, concave
on proximal surface ; harpe broad, gradually narrowed from middle to base, dorsal
edge of apical half somewhat elevate, densely denticulate, apex rotundate, longi-
tudinally grooved, bearing a great number of small conical teeth at and near the
edge; no process. ? not examined.
Early stages not known.
Ilab. Upper Amazons and eastern slopes of Peru ; may be expected to occur
in Eastern Ecuador and Eastern Bolivia.
In the Tring Museum 2 c? c? from Iquitos (Stuart).
XII. Homerus Group.
Costal margin of forewing dentate, the teeth especially prominent in male.
Abdomen without spots, entirely black, or the underside tawny-olive. Frons much
wider than the eye is high (frontal aspect). Antenna short ; club rather strong.
Ujiper cross-vt'in of forewing (D-) longer than the second (D').
A. Hindwing, on underside, with a band of spots across disc,
the spots rounded on distal side, at least the central
ones :
a. Males .......... b.
Females d,
b. Discal patch R^ — M' of forewing about three times as
long as liroad, extended to cell ; cell-patch very large,
at least on luiderside ..... Sjjecies Nos. ll.j and 110.
Discal jiatch R^ — M' widely separated from cell . . c.
c. Discal band on underside of hindwing consisting of separate
red spots Species No. 112.
This band continuous, more or less creamy white, at least
along its proximal edge . . . .' Species Nos. 113 and 114.
( 634)
(1. Discal spots of underside of hindwing small, red, edged
with black, last one entirely' red or only a small
portion of it cream}' white
Last discal spot of underside of hindwing for the greater
part creamy white .......
e. Underside of forewing with a large shadowy patch in cell
near its middle ........
Cell-patch, if present, situated close to ape.K ol'cell .
B. Hindwing lielow with coutinnons pale hand (sometimes
washed over with brown) which is dentate upon the
veins, being sinnate between the veins. Hindwing
above with broad pale yellow continuous band across
middle, or the posterior submargiual spots of the
forewing beneath orange, or there is a discal row
of orange-red lunules, or a dentate orange-red band on
hindwing .
Hindwing without such a band on tipperside ; no orange
spots ..........
e. Last one or two submargiual spots on underside of fore-
wing orange
No orange spots on forewing ; submargiual spots of
underside of hindwing orange or red ....
J\ Pale band of underside of hindwing washed over with
brown .........
This band very sharply defined, strongly dentate on distal
side ..........
C. Hindwing rounded, without band ; forewing with large
orange-red patches .......
Species No. J 12.
(Species No. 1 1 ."i.
Species No. 114.
Species No. 119.
Species No. l~!i).
Species No. 118.
Species No. 117.
Species No. 121.
112. Papilio victorinus Doubl. (ls44).
Pajulio victorinus Doublecky, Ann. Mag. N. H. xiv. p. 418 (1844) ( $ , west coast of Amer.) : Gray,
Cat. Lei,, -^ns- ^>'>>- -^J""- '• ^"i'- P- 35- n. CO. t. 6. fig. 3. ? (1852).
cj. i'ppei-side. — Forewing : a row of creamy buff submargiual spots, and a discal
row of similar spots, this row curving costad, the upper spots usually absent or
vestigial, sometimes the whole row absent ; some specimens with an additional
row of indistinct olivaceous buflf blotches anteriorly between the discal and sub-
marginal series ; costal margin serrate. Hindwing : a discal row of more or less
rounded or ovate spots, widely separated from one another, and a row of submargiual
crescent^ all creamy buff, in between the two rows a series of olivaceous buff
blotches, which are absent from the Costa llica form (of which only one specimen
is known !) ; no tail, tooth B? usually a little more prominent than the others.
Underside. — Forewing : discal row of spots complete ; atiterior subniarginal spots
vestigial, brown. Hindwing : a discal row of red ronuded spots, all nearly the
same size, the last one creamy posteriorly ; submargiual hall'moous red ; some
specimens with bufBsh blotches at the outer side of the discal spots.
? . Dimorphic, at least in certain districts, perhaps everywhere. One form
resembling the male, bearing larger postdiscal olivaceous buff blotches on the
upperside of the hindwing, the second form bearing instead of these blotches and
( 635 )
the diseal spots a broad olive band, which narrows anteriorly ; snbmarginal spots
sometimes orange on npperside.
Neuration : lower angle of cell of forewing obtuse ; SO- of hindwing midway
between base and R', or more proximal.
Genitalia: (J. Tenth tergite long, spatnlate, rather narrow before apex; sternite
laterally with a transversely divided ridge, the proximal portion acuminate, the
distal portion shorter, rounded, its anal edge irregular; beneath this second lobe,
on the anal side, a small but distinct tooth ; harpe long, slender, of almost even
width from base to near apex, dentate distally, ending in a long tapering process
which is curved upwards. ?. In front of the vaginal orifice an acuminate or
sinuate flap ; lateral edges of orifice converging posteriorly, forming behind the
orifice a channelled mesial ridge at each side of which there is a deep impression ;
laterally of this groove and somewhat more frontad there is a long, sharply pointed,
curved, twisted process, which stands on the inner side of a long and strongly
chitinised plate, the free edge of which is continuous with the anterior surface of
the vaginal flap ; this process often dentate on the posterior side.
Early stages described by Schaus (see literature below, under P. v. victorinus).
Hah. Mexico to Costa Rica.
Three subspecies.
a. P. uctorinus morelius subsp. nov.
c? ? . Only one form of female known. Upperside : distal spots of forewing
absent or small, those of hindwing also smaller than in the next form.
Uiidcrside : no spot in cell of forewing ; submarginal spots of hindwing larger
and more strongly arched than in the next.
Genitalia : S. Apical process of harpe longer and more strongly curved than
in the next subspecies ; teeth less numerous but larger, one or more long teeth on
dorsal side.
llab. West Mexico : Guerrero ; Oaxaca.
In the Triug Museum 16 c?<?, 5 ? ?, from: Guerrero (0. T. Baron), type; Los
Cojones, 17. August 1904, and Balsas R., near Iguala, Guerrero, 26. August 19U4
(Dr. Gadow).
In the Hope collection at Oxford from Oaxaca.
b. P. victorinus victorinus Donbl. (1844).
PainUo vidorhms Doubleday, I.e.; id., List Lep. Lis. Brit. Mas. i. p. 18 (1845); id., Westw. & Hew.,
Gen. Diurn. Lep. i. p. 16. n. 147 (184G) ; Gray, I.e. ; id., List Lep. Ins. Brit. ilus. i. Pap. p. 47.
n. 177 (1856) (West Coast of America) ; ToUenh., Tijdschr. Ent. iii. p. 87. n. 144 (1800) (Vera
Cruz) ; Felder, Verh. Zool. Bot. Ges. Wieii xiv. p. 313. n. 337 (1864) ; Kirby, Cut. Diiirii.
Lep. p. 537. n. 131 (1871); Oberth., Et. d'Ent. iv. p. 73. n. 221 (1880) (" Ecuador" error loci);
Godm. & Salv., Biol Ceiilr. Amer., Lep. Rhop. ii. p. 235. n. 71 (18y0) (Vera Cruz, dxaca ,
Guatemala ; Honduras ; Nicaragua ; = ainphissus = helleri) ; Haase, Untersuch. .Mimicnj i.
p. 94 (1803).
? . Pajiiliu helleri Felder, I.e. n. 336 (1864) (Mexico ; iioin. indeser.) ; id., Reise Novara, Lep. p. 91.
n. 70. t. 13. 6g. c. d. ? (1865) (Mexico) ; Schaus, Papilio iv. p. 101 (1884) (descr. of larva and
pupa).
? . Papiliv ampltissus Hopffer, .S7e</. Enl. Zeit. xxvii. p. 27. n. 8 (1866) (Mexico); Kirby, I.e. p. 567
n. 329 (1871).
Papilio victorinus var. a. P. helleri, Kirby, l,c.
6- Upperside: diseal row of spots on forewing represented at least by three
spots ; most specimens with a row of postdiscal olivaceous buff or bluish blotches.
41
( 636 )
riiu Iwiiig : discal spots larger th;m submarginal oucs, outside tbeiu a row of
olivaceous bull' spots, which are sometimes verj' small.
On underside a sjiot of variable size iu cell of forewing.
?. Dichromatic. One form resembling the male, but the postdiscal spots
usually larger, sometimes confluent with the discal ones (?-i. vtctorinus). In the
type-specimen the olivaceous or bluish blotches are very small (they are not
indicated in Gray's figure of the type). In the second form ( ? -f. amp/iissi/s) tlie
markings of the upperside are bluish or greenish, and the discal and postdiscal
spots of the hind wing are replaced by a broad band.
Genitalia : c?. Harpe more densely dentate distally than in the preceding, the
teeth being also smaller.
Ilab. Eastern McyIco to Nicaragua.
In the Tring Museum 18 cJc^, 8 ? ?, from: Cuesta de Misantla, June l>>9(i,
Jalapa, July 1897, Espinal, June 1896, Orizaba, March 1896, Vera Cruz (W.
Schaus) ; Cordoba, 2800 ft., July 1904 (A. Hall) ; S. Pedro Sula, Honduras.
c. P. nctorinus vulneratus Butl. (1872).
Papilio vulneratus Butler, Cist. Eiit. i. p. 85 (1872) (Costa Rica) ; Kirby, Gtl. Diiini. Lep. p. 814.
n 394 (1877) ; Godm. & Salv., Biol. Centr. Amer., Lejt. Rhnp. ii. p. 230. d. TJ. (1890) (Costa
Rica).
ryrrhosticta vuliienita Butler, Lcp. Erot. p. 1G5. t. 58. fig. 3. g (1874) ; id. & Druce, Pruc. ZwA.
Soc. Loud. p. 3G6. n. 385 (1874) (Costa Rica).
cJ. Only one imperfect specimen known. Discal sj)ots of both wings, on
upperside, larger, and the submarginal ones smaller, than in the preceding forms ;
no bluish postdiscal blotches on upperside of hiudwing.
Had. Costa Rica.
One specimen (type) in coll. F. D. Godman.
113. Papilio cephalus Godm. & Salv. (1890).
Pajiilio cephalus Godman & Salvjn, Biol. Centr. Amer., Lep. Wiop. ii. p. 235. n. 70. t. 71. fig. 4. 5. (J
(18'JU) (Chiriqui, iu coU. Staudinger).
cJ. One specimen only is known. Perhaps an aberration of P. cleotas
arcliytas, which we think could easily be proved or disproved by an e.\.amination
of the iunersidc of the clasper.
The row of discal spots on the upperside of the forewing extends to SM-, spots
B? — SM- not reaching to cell ; submarginal row of spots evenly curved. Underside
of hindwing with a row of blue crescents distally of the discal band ; tail long.
Head and pronotum are said to be without dots.
Hab. Chiriqui.
One specimen in coll. Staudinger. We have not examined the insect.
114. Papilio cleotas Gray (1832).
Pap'lio cleotas Gray, in Griffith, Aiiim. King,/,,,,, xv. p. CuS. t. RG. J (1832) (Brazil).
The evidence that this and the next species (P. aristeus) arc both repre-
sented iu some districts rests on the record of two single specimens bclmiging
to P. aristeus iitias. Messrs. Godman k Salvin (1S90) mention under the name of
hitias a specimen of P. aristeus from Chiriqui (Staudinger, collected by Trotsch).
The second specimen is recorded by Felder from Bogota, and described as P. ctesias.
This latter s])ecimen has the body much compressed, looking as if it hiid been
carried in a pocket-book. It may have been brought to Bogota from tlic casteru
( 637 )
side of Ecnador. If Standinger's Cbiriqui example is not authentic, the viiriciiis
forms of P. cleotas and of P. aristeus represent each other geograjihically and
should accordingly all be regarded as forms of one species. This would be a rational
systematic treatment of the insects, considering that the distinguishing characters
are individually variable, that some forms differ constantly and others do not, that
some differ only in pattern or colour, others in structure, and others again in
strncture and pattern. The geographical distribution of the forms of P. cleotas
(Costa Rica, Panama, Colombia, Venezuela, and S.E. Brazil), and of P. arist'us
(Ecuador to Bolivia, Upper Amazons, Guiana, S.E. Brazil ; and " Bogota,"
" Chiriqui," as mentioned above), renders it highly probable that we have here to do
with one species only, the occurrence of P. aristeus bit/as in " Chiriqui and Bogota,"
tcing accidental or the records being erroneous. This conclusion is corroborated by
the fact that the Brazilian form of P. cleotas agrees in structure better with the
Andesian forms of P. aristeus than with the nortliern forms of P. cleotas.
However, as we may hope to get soon further evidence of the actual distribution
of these insects, we deem it sufHcient for the present to have drawn attention to
P. cleotas and P. aristeus being only doubtfully distinct from one another.
(J. Costal margin of forewing serrate ; a submarginal row of spots nearly
parallel to distal margin, curving costad in front ; a discal row of large spots
from lower angle of cell to hindmargin, oblique or curved, approaching the
submarginal row posteriorly ; the two rows very strongly diverging anteriorly,
there being usually a row of olivaceous buff or greyish blue blotches between
them ; cell-patch large, small, or absent ; often some spots distally of cross-
veins. Hindwing : tail acnte or absent ; a creamy discal band of variable
width ; a submarginal row of spots.
Underside always with a patch iu cell of forewing ; hindwing always with a
discal band of spots, these spots red, proximally more or less creamy, sometimes
more extended creamy than red, last spot always creamy yellow, often also the first.
? . Dimorphic ; one form resembling the male, the other having the markings
of the upperside more or less bluish or olive-buff.
Genitalia : S. Tenth tergite long, spatulate ; sternite geograjihically and
individually variable, the usual double ridge on each side, the first projection
being usually pointed and often dentate, the second tooth more obtuse, bearing
often on anal side a tooth ; at apex of sternite mesially several teeth one behind
the other, or one tooth, or a tooth right and left, or no teeth. Harpe long, of nearly
even width, slightly curved, apical portion geographically and individually variable,
mostly produced into a fork, fishtail-shaped, often the dorsal prong reduced and the
ventral one curved upwards, sometimes three processes, in most forms a number of
teeth in between the prongs and proximally of them. ? . Similar to the armature
of P. victorinus ; lateral process dentate ; proximal flap variable, dentate or simple,
long or short.
Early stages not known.
Mab. Costa Rica; Panama; Colombia; North Venezuela; South-East Brazil.
a. P. cleotas arclnjtas Ilopff. (1806).
Papilio archylns HopSer, Sleti. Enl. Zeit. xxvii. p. 2S. n. 0 (ISGl".) ((J, Central America); Kirby,
Cat. Diurn. Lep. p. .'i67. n. 330 (1871) (Amer. contr.).
Papilio laetitia Butler, Cist. Eni. i. p. 84 (187^) (Costa Rica) ; Kirby, Cut. Dium. Lep. p. 813<
n. 370 (1877); Staud., Exot. Tag/, p. 15. t. 10. (J (1884) (Costa Rica ; Chiriqui, 2U00— 2o00 m,,
$ similar to (J).
( 638 )
PyrrhoxHrla hietHia Butler, Lrp. Exol. p. 16i. t. 58. fig. 4 (1874) ; id. & Druce, Proc. Zool. Soc.
Lnml. p. 366. n. .884 (1874) (Costa Rica).
Papilir, phafinn var. laetifia, Oberthiir, Et. d'Eiit. iv. p. 7-3. sub n. 220 (1880) (Chiriqui).
Papilio phaeton, Godman & Salvin, Biul. Centr. Amer., Lep. Rhop. ii. p. 2,34. n. 6'.1. t. 71. fig. 6. genit.
(1890) (variab. of ? ; Costa Rica ; Panama ; 4000—5000 ft. ;— " Colombia ' alia subsp.).
c?. The middle discal spots and the cell-patch of the forewing on the whole
larger and the submargiual spots smaller than in the Colombian form ; hindwing
obtusel}- dentate. The cell-patch is very variable, being sometimes almost srpiare
and sometimes nearly obsolete; the discal spots also vary mnch in size. Tlie npper
submarginal dots of the forewing are nsnally smaller than the posterior ones, the
first one being occasionally absent ; the spots around apex of cell mentioned iu
Ho2)fter's description are seldom all present.
?. The two forms are each individually very variable, some specimens standing
intermediate between the two forms :
ci}. ? -f. arclnjtas similar to the male, sjiots of forewing on the whole smaller,
if sharply defined, or large and ill-defined.
i'. ?-f. pantliias nov., markings of upperside more or less greenish or bluish
(type of name from Chiriqui).
Genitalia: 3. Harpe with three apical prongs, there being besides the iisiial
two prongs one on the surface proximally of the ventral one of the usual pair ; this
additional prong is a prolonged tooth, there being mostly several other but smaller
teeth between the prongs. ? . Vaginal flap acuminate, dentate, long ; lateral
processes with several prominent teeth.
Hab. Costa Rica ; Panama ; Brava I.
In the Tring Museum 12 cJc?, 5 ? ?, from: Carillo, Costa Rica, 3000 ft.,
October 1904 (A. Hall); Guatil Piris, Costa Rica, December 1901 (Underwood);
Chiriqui (Gounelle) ; Volcano de Chiriqui, 5000 — 9000 ft. (Watson) ; Boqnete,
3.500 ft. (Watson) ; Brava I., January 1902 (Batty).
b. r. cleotas phaeton Lucas (1857).
PapiUo phaeton Lucas, in Casteln., Voy. Amir. Sud, Lep. p. 197. t. 2. fig. 1. i^ (1857) ("Bre'zil
interieur" false); Felder, Verh. Zool. Bat. Ges. Wieii xiv. p. :-il2. n. 332 (1864) ( parti ni ;
Bogota) ; Kirby, Cat. Diurn. Lep. p. 537. n. 136 (1871) (Amer. mer.) ; Oberth., Et. d Ent. iv.
p. 73. n. 220 (188U) (var. excl. ; N. Granada, type) ; Godm. & Salv., Biol Centr. Amer., Lep.
Rhop. ii. p. 234. n. 69 (1890) (parlim ; Colombia) ; Maass. & Weym., in Stiibel, Reiaen
S. Amer., Lep. p. 17. n. 15 (1890) (east side of Bogota Cordillera, 1300—1500 m.).
Papilio phaeton, Doubleday, List Lep. Ins. Brit. Mus. i. p. 18 (1845) (Colombia ; nom. nudum) ;
id., Westw. & Hew., Gen. Diurn. Lep. i. p. 16. D. 149 (1846) (nom. nud.) ; Gray, Cat. Lep. /«.<.
Brit. Mus. i. Pap. p. 35. n. 170 (1852) (uom. nud.) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 47.
n. 178 (1856) (nom. nud.) ; Hopff., Stett. Ent. Zeit. xl. p. 54. n. 28 (1879) {partim) ; Staud.,
Exot. Tagf. p. 15 (1884).
Papilio clearchus Felder, I.e. p. 313. n. 332 (1864) (nom. indescr. ; Bogota) ; id., Reise Novara, Lep.
p. 88. n. 67 (1865) (Bogota) ; Kirby, Cat. Diurn. Lep. p. 537. n. 134 (1871) (Bogota).
(J. Size of cell-patch and of discal spots on npperside of forewing very variable,
the cell-patch sometimes absent, usually more oblique than in the preceding
subspecies ; upper submarginal spots larger than the posterior ones. Width
of band of hindwing likewise variable, the spots composing it sometimes separated
from one another ; tail longer than in 1'. c. arclnjtas.
On underside the iwsterior submarginal spots of forewing usually close to
the discal ones, often joined to them.
Felder's name clearchus is based on specimens without cell-patch on upperside
of forewing, with small discal sjiots, and with the band of tlie hindwing, above, cut
( 639 )
np into spots. Such specimens are hardly distingnishable from the next form,
except by the genitalia. '
?. Only that form of this sex is known to us which is similar to the male.
Genitalia : S ■ Tenth sternite usually with a small mesial tootli at apex as in
/'. aristeus ; harpe ending in a hook which is strongly curved internad, the dorsal
prong of the fork reduced to a short dentate projection which is more ])roximal than
the dorsal prong in P. c. archytas, the oblique edge between the dentate projection
and the apical hook more or less dentate. ? not dissected.
Hah. Colombia : Magdalena and Cauca valleys, and Cordillera of Bogota.
No representative is known from the West Coast of Colombia.
In the Tring Museum 90 c?(?, 1 ? , from : " Bogota " ; Muzo, November 1890 ;
Pereira, Canca ; Cauca (Child) ; Guadalite, Cundinamarca July 1903 (M. de Mathan).
c. P. cleotas coroehtis Feld. (1860).
(J PapiUo cleolax, Kollar (iion Gray, 1832, err. det.), Silz. Bcr. K. Ak. Whs. Wien, Ulath. Nat. CI.
i. p. 355. n. 12 (1850) (Angostura, Venezuela ; — locality correct?).
$. Papilio curocbiis Felder, Wien. Enl. Mon. v. p. 75. n. 11 (1860) (MiSrida, coll. Kadeu) ; id.,
Verh. Zool. Bot. Gen. Wien xiv. p. 312. n. 326 (1864); id., Reise Novara, Lep. p. 84. n. 64. t. 13.
fig. a. b. (1865) (Merida) ; Staud., Exot. Tagf. p. 15 (1884).
(J. PnpiVin pliilocleon Felder, Verh. Znnl. Bot. Ges. Wien xiv. p. 313. n. 333 (1864) (nom. nud. ;
Colombia); id., Eeise Novara, Lep. p. 89. n. 68 (1865) (hab. ?); Kirby, Cat. Diurn. Lep. p. 537.
n. 135 (1871) (Colombia).
5 Papilio corehiiK (!), Hewitson, Exot. Butt. iv. Pap}, x. text sub runthupleiira (1869).
? Papilio birehalli var. a. P. roroebus, Kirby, I.e. p. 538. sub n. 139 (1871) (Venezuela — "Yucatan"
error loci).
J. PapiUti 2>liaeton, Hopffer, Stelt. Ent. Zeit. xl. p. 54. n. 28 (1879 {partiin).
^ Papilio lijrortax Felder. ll.ec. ( parti m ■ R. Negro) ; Stand., I.e. p. 15. t. 10. ^ (1884) (Venezuela) ;
Habnel, Iris iii. p. 194 (1890) (Mrrida) ; Eimer, Orthogeii. p. 221. fig. 169 (1897) (diagr. copy
of Staud. 's fig. ; neur. of bindw. erroneous).
(J. Papilio cleotas var. lycortas, Oberthiir, El. d'Eiii. iv. p. 73. sub n. 219 (1880) (Merida).
In pattern not sharply separated from the preceding form.
<S. Disoal spot R^— IP of upperside of forewing more proximal than in
P. c. phaeton, the last two spots of this row comparatively larger, the submarginal
spots on the whole smaller, cell-patch vestigial or absent, never so sharply marked
as it is in many phaeton. Hindwing : third and fourth spots of discal band
smaller than in phaeton, all the spots separated, the third occasionally obsolescent ;
submarginal spots small in most individuals.
? . Dichromatic.
a'. ?-f. (Hone nov. ; similar to the male, usually with bufHsh blue scaling
proximally of discal band of upperside of hindwing.
b'. ?-f. coroebus Felder, I.e. ; markings of upperside nearly olive-buff or
bluish, the spots of forewing reduced or ill-defined ; discal band of hindwing,
above, widening behind ; discal band of underside of hindwing narrower than in
?-f. dione, more extended red.
Genitalia : S- Harpe similar to that of phaeton, broader before apex, the
proximal dorsal prong rather longer, pointed, the teeth between this prong and
the apical hook longer and more regular in position. ?. Vaginal flap broad
and long, dentate, lateral process armed with several long teeth.
Hah. Eastern side of the Cordillera of Bogota ; Venezuela.
In the Tring Museum 33 SS, 10 ? ?, from: Peperital to Buenavista, East
Colombia, 400 — 1200 m., January 1897, dry season, forest (Dr. Biirger) ; Merida,
Tachira and Mocotond, Venezuela (Briceno).
( 640 )
(I. P. cleotas cleotas Gray (1832).
cJ. Popillo c!e;l>i.i Gray, in Griffith, Anim. Khifld. xv. p. 073. t. 86 (1832) (Brazil) ; Boisd., Sper.
Gen. Lip. i. p. BiU. n. 207 (183(5 (Brazil ; Uruguay : ^, ? ) ; Doubl., Lhl Lep. Ins. Brit. Mux.
i. p. 18 (1845) (Brazil) ; id., Westw. & Hew., Gen. Diurn. Lep. i. p. IG. n. 50 (1846) ; Gray,
Cfa*. Lep. Ins. Brit. Mils. i. Pop. p. 35. n. 172 (1852) (partim) ; id., List Lep. In.<<. Brit. Mm.
i. Pop. p. 47. n. 180 (1856) (pnrtim) ; FeHer, Vn-I,. Ztml. Hot. Ge«. H>M xiv. p. SCO. n. 184
(1864) ; Kirby, Cat. Diurn. Lep. p. 537. n. 1.33 (1871) (Brazil ; Uruguay) ; Burm., /Jrsrc. Pep.
Argent. \. Lip. p. 61. n. 3 (1878) (/xirtim ;— descriptio ad spec. P. hellmiirhux dictamreferend.) ;
Oberth., Et. d'Ent. iv. p. 73. n. 219 (1880) (Uruguay) ; Haase. Cntvrsncli. .Mimicnj i. p. 'J4
(1893) ; Weym., Stett. Ent. Zeit. Iv. p. 315. n. 10 (istl.-,) (Bio Graude do Sul) ; BOnningh.,
Verh. Ver. Nat. Unterh. Ihtmhurg ix. p. 28 (18'J6) (Petropolis).
Papilio bjcortas Felder, Wien. Ent. Mnn. v. p. 75. n. 10 (1860) (" Caracas " error loci) ; id., Verh.
Zool. But. Ges. Wien xiv. p. 313. d. 3.35 (1864) {partim ; "Caracas") ; id., Rrixe Nnrara, Lep.
p. 90. n. 69 (1865) (_p,irtint ; " Caracas ") ; Kirby, Cat. Diurn. Lep. p. 537. n. 132 (1871).
Papilio oleutas (!), Felder, I.e. p. 313. n. 334 (18i;4) (Bras, austr. ; Uruguay).
Papilio pliaHtun, Hopffer (non Lucas, 1857, err. det.), Slelt. Enl. Zeit. xl. p. 54. n. 28 (1879) [partim).
Papilio eleothas (!), Mabilde, Guia Pract. Bnrhul. Rio Grande do Sul p. 46 (1896).
Papilio cleotas var. od. n. sp. ?, Bonningh., I.e. (''markings green " — doubtless ? ).
Felder's lycortas, which is based on specimens supposed to be from Caracas
(Kaden), is the same as cleotas ; Kaden's specimens were doubtless Brazilian.
$. Discal band of forewing, upperside, more oblique than in the other
subspecies, spot M- — S5P small or vestigial; coU-patch small or absent; a row
of large olivaceous buff blotches from costal margin backwards bet\Yeen the discal
and submarginal rows of spots. Band of hindwing always broken uji into spots ;
the last four or five submarginal spots red in most specimens.
? . Dichromatic.
«'• ?-f- cleotas similar to the male, easily distinguished from P. cleotas coroebus
?-f. clioiie by the row of olivaceous buff blotches on the forewing and the smallness
of the discal patch M-— SM-.
b'. ?-f. adaea nov. Name-type from Blumenau. An oblique discal row
of bluish blotches on upperside of forewing, the row gradually disappearing in
front, usually not extending forward beyond lower angle of cell ; postdiscal row
of similar blotches from costal margin backwards; sometimes the posterior post-
discal spots merged together with the discal ones ; a submarginal row of spots as
in male, the upper ones being more or less lijuish. Hindwing : a discal and a
postdiscal band of spots huffish blue, the discal band broader than the jjostdiscal
one, the first and the last discal s])ot often partly creamy ; submarginal spots
huffish blue, anterior ones more or less creamy, last one usually red. Discal
band of underside of hindwing more extended red than in ?-f cleotas ; last spot
cnamy on abdominal side.
Genitalia : cJ. Tenth sternite with or without anal mesial tooth ; harpe similar
to that of P. ansteus bitias, with two straight prongs at apex of nearly the same
length, both directed anad, there being often a very few teeth between the jirongs,
some specimens bearing some minute teeth on the hinder side. ?. Lobe in front
of viigiual orifice short, simple or dentate, lateral process slender, very sharjdy
pointed, bearing a tooth on the posterior side.
Ilab. Brazil ; Uruguay (teste Boisduval).
In the Triiig Museum 9 c? <?, 6 ? ? , from : Petropolis, March 1898 (J. Foetterle) ;
E^piritu Santo; TheresopoJis, S. Catharina, November 1894— February 1895
(J. Michaelis) ; Blumenau.
( 641 )
115. Papilio aristeus Gram. (1781).
? . PapiUo aristeus Cramer, Pap. Exot. iv. p. 139. t. 361. fig. A. B (1781) (Surinam).
Since the /'. aristeus of Cramer's plate 318 belongs to the Kite-Swallowtails,
and therefore will stand in another genns when the Swallowtails are genericall/
revised, there is no objection against retaining the name aristeus also for the present
insect. There is no reference to this species in the Inde.K of Cramer's volnme.
We have explained above, nnder P. cleotas, onr reasons for regarding P. aristeus
as being doubtfnlly distinct as a species from P. cleotas.
(S. Cell-patch of forewing very large ; discal patches W — M- very long,
contignoiis with the cell-patch, sometimes patch M' — M- obsolescent. Hiiidwing
with or without broad creamy discal band.
? . Dichromatic. One form resembling the male. In the second form the
markings of npperside bluish, snbmarginal dots the same colour or partly creamy;
forewing with band of ill-defined patches on disc ; hindwing with broad discal band,
widening behind, entering cell.
Cell-patch of forewing below more or less distinct, large.
Genitalia similar to those of P. cleotas ; tenth sternite of male always with
mesial apical tooth or several teeth standing one behind the other ; harpe with two
apical processes and some teeth. • ? . Nearly the same as in P. cleotas cleotas ;
lobe in front of vaginal orifice short, broad, dentate, carinate on hinder surface.
Early stages not known.
Hah. Dutch and French Guiana; Upper Aiiiazous ; Ecuador to Bolivia; Sao
Paulo (S.E. Brazil) ; Bogota and Chiriqui.
We doubt the correctness of the record from the last two localities.
a. P. aristeus aristeus Cram. (1781).
$ . PapiUo aristeus Cramer, I.e. (Surinam).
$ . CalaiJes menalius Hiibner, Verz. bete. Sc/imelt. p. 8G. n. 804 (1818 ?) (nom. nov. loco ari.'itevs).
? . Papilio hitias a. Papilio aristeus. Godart, En-, .\fetli. ix. p. 39. sub n. 43 (1819).
$ . Papilio coristlieus Boisduval, Spec. Gen. Lip. i. p. 323. n. 166 (183G) (nom. hov. loco aristeus) ;
Doubl, Westw. & Hew., Geu. Diurn. Lep. i. p. 20. n. 246 (1847) ; Gray, Cat. Lep. Ins. Brit.
Mus. i. Pup. p. 67 D. 298 (1852) ; id.. List Lep. Ins. Brit. Mas. i. Pap. p. 77. n. 315 (18o6) ;
Felder, Yerh. Zool. But. Ges. Wicn xiv. p. 312. n. 325 (1864).
5 Papilio aristiieus (!), Boisduval, I.e. (sub syn.).
?. PapiUo memitiiis, Kirby, Cat. Diurn. Lep. p. 538. n. 141 (1871) (Surinam) ; Haase, Untersueli.
Mimicry i. p. 94 (1893).
cj. Papilio ctesias var. (geographica ? An species distincta?), Imri Oberthur, Et. il'Ent. iv. p. 72.
sub n. 217. t. 5. fig. 3 (1880) (Guyane frangaise).
(?. Upperside. — Forewing : cell-patch rliombiform ; discal spot R- — R' long,
patch R^ — M' nearly reaching snbmarginal spots, ill-defined distally, patch M' — M-
rcpresented by a narrow shadowy streak situated along M'. Hindwing : discal
band represented by two creamy patches M' — SM- and vestiges of the other spots.
Discal patch M' — M^ of underside of furcwiug much larger than above,
extending to M-.
? . The figure of Cramer appears to be all that is known of this sex. A row
of long, ill-defined, greyish blue streaks on disc of forewing from costal to inner
margin ; blue band of hindwing very broad.
Hab. French and Dntch Guiana.
One (J in coll. Oberthtir. We have not seen this insect in other collections.
( G42)
b. P. aristeus ctesiades snbsp. nov.
cj. Papilio clesias Hahnel (nnn Felder, 1865, err. det.), Iris iii. p. 297 (1890) (Iquitos).
fj. Papilio ctesias var., Staudinger, Iris iv. p. 64 (1891) (Upper Amazons).
This form has been distributed by Messrs. Staadinger and Bang-Haas nnder
the name ctesiades, which we retain.
The name has apparently not been published by the late Dr. Standinger.
<?. Cell-patch of forewins:, ?^/)/.<^r.s;V/(', placed along median vein, nearly reaching
to base, touching SO near base ; distal patches W — M- large, there being also a spot
behind M- occnpying the angle formed by this vein and M. No discal band on
hindwing, but occasionally traces of the last two spots of this band.
Discal spots of underside of hindwing red, small, the first one or two and the
last one or two creamy white.
? not known.
Hab. Upper Amazons.
In the Tring Museum 2 SS from Iquitos.
In coll. Oberthiir from : Iquitos ; Cavallo Cocho, Peru, May — July (Mathan).
c. P. aristeus di/smias subsp. nov.
Papilio bitias, Godman & Salv. (iion Godart, 1819, err. det.), Biol. Centr. Amcr., Lep. Rhup. ii. p. 237.
sub n. 73 (1890) (S.E. Brazil).
(J. ^\rm\&x to P. a. ctesiades. Upperside, (orewing : cell-patch smaller, being
narrower, not extending to the base ; a small spot at apex of cell and a dot
beyond cross-vein D^ ; two large patches R^ — M- on disc, the first not quite reaching
W, tlie second distally somewhat narrowed, joining the subcostal spot, a small patch
behind base of M-, as in ctesiades, hnt smaller; submarginal spots SC — R' elongate.
Hindwing as iu bitias, the discal band being represented by a single sj)ot
C — SO- and some blnish dispersed scales posteriorly on disc ; submarginal spots
mnch larger than in ctesiades.
Underside : cell of forewing with a large blotch at npper angle ; discal jmtches
as above, the first being rather wider, jiatch behind base of M- vestigial ; three
distinct submarginal spots. Hindwing as in bitias.
Bab. Province Sao Paulo (Rogers), probably from the interior ; may be
expected to occur in Goyaz and Matto Grosso.
1 cJ in coll. F. D. Godman.
Connects ctesiades with bitias.
d. P. aristeus bitias Godt. (1819).
(J. Papilio hitias Godart, Enc. Meth. ix. p. 39. n. 43 (1819) (South America.— quot. Cram, excl.) ;
Boisd., Spec. Gen. Lep. i. p. 323. n. 165 (1836) ; Doubl., Westw. & Hew., Gm. Diurii. Lep. i. p. 20.
n. 245 (1847) ; Gray, Cat. Lep. Ins. Brit. Mnn. i. Pup. p. 67. n. 297 (1852) ; id., List Lep. Ins.
Brit. Mus. i. Pop. p. 76. n. 314 (1856) (South America) ; Felder, Verh. Zool. Bot. Ges. Wien xiv.
p. 312. n. 329 (18C4) (hab.?); Kirby, Cat. Diurn. Lep. p. 537. n. 138 (1871); Staud., Exnt.
Tagf. p. 15. t. 10. $ (1884)(Chanchamayo) ; Maass. & Weym., in Stiibel, Reisen S. Amer., Lep.
p. 77. n. 32 (1890) (Rio Mayo, N. Peru); Haase, Untersm-h. Mimicry i. p. 94 (1893) (" New
Granada") ; Haen.sch, Bn-l. Ent. Zeitschr. xlviii. p. 153 (1903) (Sa. Inez. R. Pastaza, 1250 m.).
(J. Papilio curolas Felder, Wien. Ent. .Uon. vi. p. 66. n. 2 (1862) (R. Negro) ; id., Verh. Zool. Bot.
Ges. Wien xiv. p. 312. n. 328 (1864) ; id., lieise A\wara, Lep. p. 85. n. 65 (1865) (Upper Rio
Negro) ; Hopff., Stett. Ent. Zeit. xl. p. 53. n. 26 (1879) (Chanchamayo ; = ? hitias) ; Staud., I.e.
p. 15 (1884) ; Godm. & Salv., Biol. Centr. Amer., Lep. Mhvp. ii. p. 237. n. 73. t. 71. fig. 7. genit.
(18yO)(Chiriqui).
( 643 )
cJ. Papilio ctesias Felder, Verh. Zool. Bol. Ges. Wien xiv. p. 312. n. 330 (18G4) (nom. nnd. ; New
Granada) ; id., Reise Novara, Lep. p. 86. n. GG. t. 14. fig. c. d (18G.5) (Bogota) ; Druce, Ptoc
Zool. Soc. Loii/l. p. 240. n. 18 (187G) (Pozuzo, Cosnipata, Ucayali, Huasatnpilla) ; Oberth., Et.
d'EiU. iv. p. 72. n. 217 (1880) (var. excl. ; Ecuador).
Papilio hitias var. a. P. euro/as, Kirby, Cat. Diurn. Lcp. p. 538. sub n. 138 (1871).
Papilio bitias var. b. P. ctesias, id., I.e.
Pajjilio lacordairei Borre (Belval ined.), C. R. Soc. Eut. Belrj. xxviii. p. 12G (1884) (= cteaias).
Papilio hitias var. ctesias, Dognin, Lep. Lnja p. 14 (1887).
The characters bj- which Felder distiuguished eurotas and ctesias are purely
individual. The body of the type-specimen of ctesias is much compressed ; it may
have been brought to Bogota from Eastern Ecuador in a pocket-book. The specimens
named eurotas are said to be from the Upper Rio Negro, where no recent collections
have been made. The species probably extends eastwards between the Amazons
and Rio Meta.
(?. Cell-patch of forewing more transverse than in the preceding form, very
variable in size like the discal spots. Discal band of hindwiug, above, obsolete
or at least widely interrupted.
?. Dimorphic.
a'. ?-f. bitias resembling the male ; a specimen from Sarayafu in coll. F. D.
Godman.
b'. ?-f. therapes nov. Bluish discal patches of forewing and discal area of
hindwing less extended than in female of P. a. aristeus ; cell-patch of forewing
below partly creamy yellow.
Ilab. Eastern Ecuador and Eastern Peru ; Upper Rio Negro ; " Bogota " ;
" Chiriqni " ; the last two localities requiring confirmation.
In the Tring Museum 22.5 c? c?, 1 ? , from : " Bogota " and " Upper R. Negro "
(coll. Felder) ; Zamora, Ecuador (0. T. Baron) ; Loja ; R. Mixiollo, Loreto (Baer) ;
Pozuzo, Huilnuco (W. Hofi'manns) ; Chanchamayo (Schunke, Hoffmanns) ; Perdn^ R.,
March 1900 (Simons) ; Huayabamba ; La Merced (Watkins & Tomlinson) ; R. Toro,
La Merced, August — September 1901 (Simons) ; Caradoc, Marcapata, 4000 ft.,
February 1901 (Ockenden); Cajon, Cuzco, October 1900 ((larlepp) ; Oroya, R.
Inambari, 3500 ft., November 1901 (Ockenden); La Union, R. Huacamayo, Carabaya,
January 1901, 2000 ft., wet season (Ockenden).
A female from Ecuador in coll. Hewitson.
e. P. aristeus vilcanotus subsp. nov.
S. We know only three specimens, two of which are in the collection of Mons.
Charles Oberthiir, who received them from Staudinger under the above name, which
we accept, a third specimen from the same source being in coll. Adams.
They agree with the preceding form, but differ in the cell-patch of the forewing
and the costal spots of the hindwing being ochraceous on the upperside, the discal
patches of the forewing above being washed with buff proximally ; cell-patch below
clayish.
Hab. Vilcauota, South Peru.
./. P. aristeus coelebs subsp. nov. (PI. V. fig. 14).
(?. Papilio lenaeus,OheTi\i\ir (mm Doubleday, 184G, err. det.), Et. d'Enl. iv. p. 72. n. 218 (1880)
(Tambillo, Peru).
(?. Upperside. — Forewing: cell-patch narrow, often a mere bar; discal jiatch
R^— M' reduced distally and on costal side, patch M'— M- narrow, triangular.
( 644 )
not reaching cell, widest distally ; a row of large olivaceous bnff blotches from W
to costal margin, a more or less distinct similar blotch in apex of cell ; submarginal
spots large, a spot at hinder margin close to angle. Hindwing : band complete,
entering cell or not, the veins traversing it less extended black than in lenaeus ;
a row of rather large olivaceous buff spots at outerside of band ; submarginal spots
creamy yellow, large.
Underside \ia\&\ than in /tvwcwsand bitias. Forewing : cell-patch larger than
above, but much smaller than in bitias and lenaeus ; a blotch in upper angle of cell,
as in most bitias ; submarginal spots as in bitias, four or five clearly marked, the
others vestigial. Discal band of hindwing creamy white, each spot except the
last bearing a rufous red spot distally, the veins narrowly black, the band not being
broken up into separate spots.
? not known.
Hab. North-AVest and North-Central Peru.
In coll. Oberthiir a small series from Tambillo and Chachapoyas.
In coll. Dognin from Loja (probably western side).
g. P. aristeus lenaeus Doubl. (1846).
cJ. Pupilio Icnaciis Doubleday, in Doubt, Westw. & Hew., Gen. Diuni. Lej>. i. p. 16. n. 148. t. 4.
fig. 2. (^ (184G) (Bolivia) ; id., LikI Lep. Ins. Bnl. Mux. i. Ajijienrl. p. 4 (1848) ; Felder, Verli.
Zool. Bui. Ges. Wien xiv. p. 312. n. 331 (18G4) (Bolivia) ; Kirby, Cut. Diuni. Lep. p. 537. n. 137
(1871) ; Hopff., Stelt. Evt. Zeit. xl. p. 54. n. 27 (1'57'J) (Bolivia) ; Haase, Untersiich. .Uimicri/
i. p. 1)4 (1893) ; Weeks, Tllustr. Diuni. Lep. p. 20 {VM:>) (Cbulumani).
Papilio cleolax. Gray, Cat. Lep. Ins. Brit. ilus. i. Pap. p. 35. n. 172 (1852) (partim ; " lenaeus = $ of
cleotas " false) ; id., List Lep. Ins. BHt. Mus. i. Pap. p. 47. n. 180 (1856) (partim).
Papilio phaeton, Hopffer, I.e. xl. p. 54. n. 28 (1879) {jiartim) ; Weeks, I.e. (1905) (Cbulumani).
<?. Cell- patch of forewing on the whole less oblique than in bitias, discal spot
R- — W always present, patch M' — W strongly narrowing proximally. Band of
hindwing complete or narrowly interrupted, often entering cell.
? not known.
Bab. Eastern Bolivia, and S.E. Peru.
In the Triug Museum IT JcJ from : E. Songo to R. Snajii, llnO in., March —
June IS'JO (Gark'])}.) ; Gnanay, Majiiri E., 1500 ft., March 1895 (Stuart); S.
Augnstin, Mapiii R., 3500 ft., and Majiiri, 1800 ft., September 1895 (Stuart); Reyes
and Muscliay, R. Beui, Augnst 1S95 (Stuart) ; R. Tanampaya (Garlepp) ; Province
Sara, S. Cruz de la Sierra, February — April 1004 (J. Steinbach).
110. Papilio judicael Oberth. (1888).
cJ. Pupilio juclieaei Oberthiir, Et. d'Ent. xii. p. 3. n. G. t. 2. fig. 4 (1888) (Huambo, N. Peru).
(?. Upperside. — Forewing : a narrow patch across cell and a band of four
discal spots from lower angle of cell to SM- dirty ochraceous ; a row of small
submarginal spots of the same colour. Hindwing resembling in outline that of
P. warscicicxi; tooth W prolonged to a tail which is less acute than in P. aristeus,
tooth M' also long, and tooth M" long, broad and obtuse ; a row of vestigial
ochraceous buff discal blotches, the first and the last spot being more distinct, a:i
incomjjlete row of post discal blotches, and a submarginal row of small spots.
Underside. — Forewing: cell-patch and discal {)atches lai-go, clayish ochraceous ;
submarginal spots small. Hindwing : a discal band as in the allied sjjecies, dirty
white, distally slightly washed with orangc-rcd ; submarginal spots rufous-red, the
upper ones shaded with olive-buff'.
( M5 )
? not known.
Genitalia as in P. aristeus, but tenth sternite anally in middle with two teeth
side by side, not one behind the other. Harpe with two simple prongs as in
P. aristeus and P. cleotas cleotas, the upper proug a little longer than the lower,
not lying close on the clasper, the dorsal apical edge of the harpe being a little
twisted away from the latter.
Ilab. Huamba, Dept. Amazonas, North Pern ; in coll. Oberthilr.
Only one specimen is known. The insect stands intermediate between P. aristeus
and P. cleotas, bnt has longer tails than these species. The band in the middle
of the forewing being very narrow above and very broad below is an interesting
character.
IIT. Papilio garamas Hiibn. (1834?).
?. Eupliomdes gammas Hiibner, Samwl. Ejul. Srhmelt. iii. t. 19 (1834 ?) (Mexico),
(j. Heradides asdejiius id., I.e. iii. t. 20 (1834 ?) (Mexico).
This purely Central American species consists of five subspecies, of which the
differences in pattern, though not very striking, are almost constant in a series,
being moreover accomjJanied by interesting differences in the genitalia.
<?. Costal edge of forewing serrate ; a pale j'ellow baud from costal margin
pro.Kimally of apes of cell to distal third of hindmargin ; a row of submarginal
spots of the same colour, starting at distal fourth of costal margin, consisting of
four or more spots, the fourth and following spots standing near the distal margin.
Hindwiug strongly dentate, teeth M' and M- prolonged, tail long, asymmetrically
spatulate ; a continuous discal band as on forewing, of about even width, strongly
and regularly dentate on distal side ; a row of submarginal bars, sometimes absent ;
a row of blue postdiscal blotches, varying in number, often small or vestigial.
Underside like upper, bands paler, especially that of hindwing ; submarginal
row of spots of forewing usually continued to hinder angle. — -Band of hindwiug
edged with orange-tawny distally ; submarginal crescents longer than above, more
or less orange or tawn3--orauge.
?. Dichromatic, at least in Mexico. One form is devoid of the yellow discal
band on both wings. The forewing bears a row of olivaceous buff postdiscal
blotches and a row of small submarginal spots, there being on the hindwing a row
of strongly arched discal crescents which are bright brick-red or orange-rufous,
followed by a row of blue patches and a row of submarginal spots ; on the underside
of the forewing there is sometimes a trace of the discal band of the male.
The other form resembles the c?. There are occasionally some small spots at
the distal side of the cross-veins of the forewing, corresponding to the distal branch
of the forked baud of P. homerus.
Genitalia of the same type as in P. aristeus, victorinus, etc. <S . Tenth tergite
long, strongly compressed beneath, jjrismatical, except at apex ; sternite with a
broad lateral lobe, which is divided by a groove and a sinus into a proximal
and a distal projection, there being anally in the middle sometimes a distinct
conical tooth as in P. aristeus. Harpe long, fiat, similar to that of P. aristeus and
P. homerus, geographically variable. ?. Almost exactly as in P. /ioi?ierus, the
vaginal tubercle and the lateral process somewhat dentate, the posterior lateral
ridge visible without dissection.
Early stages not known.
Hab. Mexico to Panama.
( 04fi )
a. P. garamas abderus Hopff. (1856).
cJ. rap'iVio ahilmts UopSer, Neue ScJimHI. ii. p. 1. n. 1. t. 1. fig. 1. 2 (185G) (Mexico); Felder,
VerJi. Zoo!. ISol. Ges. Wini xiv. p. 313. n. 343 (18G4) ; Kirby, Cal. Viiirn. Lep. p. 537 n I'^o
(1871).
(J ?. Papilio asdepius, Godman & Salvia {iinii Hubner, 1834 V, err. det.), Blnl Coili: Ainei:, Lep.
Rlwp. ii. p. 238. n. 75. t. 72. fig. 3. genit. (1800) {parlJm ; Jalapa ; Orizaba).
PtipiUo electrion (!), Barrett (mm Bates, 1864, err. det.), Eiit. Nrwsxi. p. 428 (1900) (Orizaba, de.scr.
of $ , similar to ^ ; distinct from P. am-lepidt:).
?. Upperside, forewing : fonr snbmarginal spots, fiftli often vestigial.
Hindwing : band always entering cell ; no snbraavginal spots.
I'm/t'rsii/t' : snbmarginal row of spots of forewing continued to SM-, more or
less distinctly angnlate at R-, usnally some traces of spots on disc between the
median band and the snbmarginal series. Hindwing : orange-red jirojections of
median band shorter than in the preceding, of the same length on both sides of
each vein.
?. Dichromatic.
a'. ?-f. abderus. — Similar to male. Forewing, above, occasionally with more
postdiscal spots than fonr and with blotches of huffish scales on disc ; band of
hindwing not always entering cell, the distal projections more or less orange-red
above, and the snbmarginal spots often vestigial, red on npperside.
b' . ?-f. amerias nov. — L'pperside, forewing: a row of obscure olivaceous buff
blotches on disc ; a snbmarginal row of spots from costal margin to SM-, the spots
small, not sharply defined. Hindwing : orange-red crescents very large (cor-
responding to the distill portion of the median band of male), strongly arched,
except the first one, crescents R' — M^ being the largest ; submarginal sjjots large,
pale tawny-orange, upper ones creamy proximally. Underside : forewing with
some creamy scaling in cell representing the cell-jjatch of male ; blotches on disc
as above, but larger ; jtroximally of them some other blotches, which are a remnant
of the median band ; snbmarginal spots small. Hindwing : all the discal crescents
strongly arched, paler than above, and the posterior ones not so broad ; submarginal
spots only slightly curved.
Genitalia : Harpe straight, apical processes also straight, lower one shorter
than the upper.
llab. Eastern Mexico : Vera Cruz.
In the Tring Museum 9 c?<?, ft ? ?, from: Orizaba, February and March,
Jalajia, July, Espinal and Cuesta de Misautla, June (W. Schans); Jalapa, August ;
Monterey ; Huatuxco.
b. P. garamas garamas Hiibn. (1834 ?).
5 . Euphoeades garamas Hiibner, J.c.
tJ. Ileraclicles asrlijjiiix id., I.e. ; Kirby, iu Suhn., Sam ml. Exot. Schmcll. ed. ii. p. 99. t. 458. fig. 1. 2,
t. 460. fig. 1. 2 (190-?) (Mexico ; synoD. puriim ; amchinatus (!) laps. typ.).
^. Papilio citiciviiatus Boisduval, Spec. Gen. Lip. i. p. 346. n. 18G (183li) (Me.\ico) ; Lucas, in
Gm'r., Did. Pill. HiM. Nat. vii. p. 50 (1838).
(J. Papilio asclepiiis, Doubleday, List Lep. Jtis. Ilrit. Miis. i. p. 17 (1845) (parliin) ; id., Westw. &
Hew., Gen. Diuni. Lep. i. p. 13. n. 84 (1840) (partim ; Mexico) ; Gray, Cut Lep. Ins. liril. Mas.
i. Pap. p. 23. n. 105 (1852) (partim ; id.. List Lep. Ins. Brit. Mus. i. Pap. p. 31. n. 112 (1856^
(partim ; Mexico) ; Weidera., Pror. Eiit. Soc. Philad. ii. p. 140 (1863) ; Felder, Verb. Zool.
But. Ges. Wien xiv. p. 313. n. 342 (1864) {partim ; Mexico) ; Kirby, Qit. Ditmi. Lep. p. 537.
D. 126 (1871) (^<«h;»0-
( 647 )
f. Papilto ijaraiims, Doubleday, Westw. & Hew., Gen. Diurn. Lep. i. p. 17. n. 178 (1846) ; Gray,
Cat. Lep. Ins. Brit. ilus. i. Pap. p. 40. n. 201 (18.')'2) ; id., List Lep. Ins. Brit. Mm. i. Pap. p. 55.
n. 212 (185G) ; Weidem., Prnc. Enl. Soc. Philad. ii. p. 147 (1863) ; Felder, Verh. Zonl. But. Ges.
Wieii xiv. p. 31.S. n. 344 (1804) ; Kirby, Cat. Diurn. Lep. p. 537. n. 124 (1871).
Pupilio concinnatus (!), Gray, ll.cc. (sub syn.).
Papilio asclepins, Oberthiir, Et. d'Ent. iv. p. 74. n. 225 (1880) (Cuernavaca) ; Godm. & Salv., Biul.
Cenir. Anier., Lep. Rhop. ii. p. 238. n. 75 (18'J0) (partim ; Oaxaca) ; Haase, Uidersuch. Mimicry
i. p. 93 (1893) ; Barrett, Ent. News xi. p. 428 (1900) (Central and Western Mexico).
Papilio asclepuis (!), id., I.e.
Messrs. Godmau and Salvin were wrong in considering Hiibner's figures as
representing two females ; the second figure is that of a male of the present sub-
species. The Mexican material at the disposal of these authors was insufficient,
which explains that the West and East Mexican forms were treated by them as
being identical, while the forms from Guatemala and farther south are described
as belonging to two separate species.
The West Mexican specimens are larger than East Mexican individuals, the
dentition of the hindwing is stronger, the tail is more spatulate, and the pattern is
different.
c?. Uppersicle. — Forewing : an evenly curved row of five or six snbraarginal
spots, often a seventh or even an eighth spot indicated ; no olivaceous buff blotches
on disc ; proximal edge of median band more even than in P. g. abderus.
Hindwing : median band not entering cell, or only just the extremity ; submarginal
spots distinct.
Underside : no blotches on disc of forewing. Hindwing : distal orange
projections of band shorter than in abderus, the anterior portion of each projection
shorter than the portion behind the vein.
? . Two forms.
a'. ?-f. amisa uov. — Similar to male; forewing often with some small spots
at upper angle of cell and distally of the cross-veins, the spots being larger below.
b'. ?-f. garamas Hiibn., I.e. — Resembling abderus ?-f. amerias. Postdiscal
olivaceous buff spots of forewing arched ; crescents of hindwing much smaller than
in ¥ -f. amerias, their horns being much shorter ; blue spots larger ; submarginal
spots thinner ; no vestiges of median band, or only a very few buff scales in cell.
Genitalia : <?. Proximal lobe of tenth sternite broad, denticulate, sometimes
sinuate ; mesially at apex of the sternite a tooth as in P. aristeus.
Ilab. West Mexico : Guadalajara, Cuernavaca, Oaxaca.
In the Tring Museum 25 JcJ, 7 ? ?, from : (Juernavaca, 4000 ft., July l'J04
CA. Hall) ; Cuernavaca, end of August 1904 (Dr. Gadow) ; Coantla, Morelos,
June 1004, 3000 ft. (A. Hall) ; Salvatierra ; Uruapan ; Mexico CUty (coll. Felder).
c. P. garamas baroni subsp. nov.
(?. Upperside. — Forewing : a row of five submarginal spots, the fifth spot
small, standing several mm. nearer the margin than the fourth. ^Ilindwing :
median baud entering cell, its proximal margin crossing cell halfway between
\\? and M' ; blue spots small ; submarginal spots vestigial.
Underside : apex of forewing and basal area of hindwing paler than in the
previous forms. Forewing : submarginal spots SC — R- continuous, this narrow
band followed from R- to SM^ by a much thinner line which begins at R- distally
of the bands. Hindwing : projections of median band as short as in P. g. garamas,
almost the same length at both sides of tiie veins ; four upper submarginal spots
( 648 )
connected with one another, as is often the case in P. (j. nbderus; dentition
prominent, tail long, spatnlate.
? not known.
Genitalia : Tenth sternite as in ff. ganams ; barpe qnite different, the upper
prong being very much shorter than the lower.
Hah. Sierra Madre do Snl, Guerrero (0. T. Baron).
1 c? in the Triug Mnseum.
(I. P. gammas electryon Bates (1864).
Papilio anclepius Doubleday (iion Hubner, 1834 ?, err. det.), List Lep. Inx. Br'U. Mux. i. p. 17 (1845)
(syn. excl. ; Honduras) ; id., Westw. & Hew., Gen. Ditirii. Lrp. i. p. 13. n. 81 (184(i) (jmrtim ;
Honduras) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 23. n. 105 (IHo'i) (syn. excl. ; Honduras) :
id., List Lep. Ins. Brit. Mus. i. Pap. p. 31. n. 112 (185G) {jiartini ; Honduras) ; Felder, Verli.
Zool. Bot. Gcs. Wien xiv. p. 313. n. 342 (18G4) (parliui ; Honduras) ; K;irby, Cat. Diurn. Lep.
p. 537. n. 1-2G (1871) (partim ; Honduras ; '-P. achehjiis" alia species).
(J. Papilio eleclryon Bates, Eitt. Mo. Mag. i. p. 3. n. 4 (18f;4) (Guatemala) ; Felder, I.e. p. 313.
n. 341 (18G4) (cit. falsa) ; Kirby, I.e. p. 537. n. 12Ga (1871) ; Godm. A Sah'., Biol. Centr. Amer..
Lep. Rhup. ii. p. 239. D. 76. t. 72. fig. 1. 2. c? (1890) (Guatemala).
i. Upperside. — Forewing : cell-patch narrowing costally ; fonr or five sub-
marginal spots, the second smaller than the third, sometimes a sixth spot indicated ;
I he first four larger than in Mexican males. Hindwing : median band entering coll
as in P. g. ahderm ; blue spots small, often vestigial ; snbmarginal spots absent ;
tail spatnlate ; red anal marginal spot rarely present.
Underside : band of forewing strongly dentate on disc ; snbmarginal line
angnlate at R- ; no blotches on disc. Projections of band of hindwing about
as long as in P. g. garamas, nearly the same length on both sides of each vein ;
snbmarginal spots linear, red.
? not known.
Genitalia : ProxitDal lobe of tenth steruite narrower than in P. g. garamas,
denticulate, mesial ai)ical tooth jircsent or vestigial ; iiarpe quite different from that
of tbe other furms, curved, denticulate, strongly widened at apex, the ujipcr prong
curved downwards.
Hah. Guatemala ; Honduras {fide Doubleday, but there are no Honduras
specimens in the British Museum, only West Mexican ones !).
In the Triug Museum 9 c? c? from: Cindad de Guatemala (Rodriguez);
Guatemala.
e. P. garamas sijedra Godm. & Salv. (1878).
Papilio syerlra Godman & Salvio, Proe. Zool. Soe. p. 271. n. 19 (1878) (Chiriqui) ; iid., Biol. Ceiitr.
Amer., Lep. lihnp. ii. p. 239. n. 77. t. 72. fig. 4. (J, 5. $ (1890) (Costa Rica ; Chiriqui).
c?. Upper.^ide. — ¥ore\ving: cell-patch of even width; five to seven snbmarginal
spots, third the largest, fifth and following thin.— Hindwing : median band entering
cell, more strongly and regularly dentate than in electryon ; blue patches large ; no
submargiual spots ; tail long, spatnlate, teeth M' and M" also long.
Underside: apex of forewing and basal area of hindwing as pale brown as in
haroni ; dentition of median band of hindwing as in clectrtjon, the orange-red border
deeper in colour, broader in between the veins.
? like <? : seven snbmarginal spots on forewing ; median band of hindwing
partly edged with orange-red above ; no snbmarginal spots.
( 649 )
Genitalia : Tenth tergite shorter than in 1'. (j. elfctri/on ; harpe peculiar, twisted,
lower process long, gradnally broadening proximally, the np[)er ])roces.s, which is
short, appearing as a tooth at the dorsal edge of tjie harpe.
Hab. Costa Rica ; Chiriqui.
In the Triug Maseum 10 cJcJ, 1 ?, from: Azahar de Cartago, Costa Rica
(Underwood) ; San Jose, May — Jnne 1899, Guatil Piris, December 1901 and
February 1903, Costa Rica (Underwood); Chiriqui (Gounelle) ; Chiriqui.
118. Papilio homeras Fabr. (1793).
PapiUii Eqiiei Arjilnix humerus Fabricius, Eiit. St/sl. iii. 1. p. 29. n. 85 (1793) (America ; "Latham
coll." false, teste Donovan) ; Esper, Ausl. Schmett. p. 190. n. 85. t. 40. fig. 1 (17'J6).
Papilio humerus, Donovan, Xal. Rejios. ii. Kitt. t. 19 (1823) (Jamaica) ; Godart, Em'. Melh. ix.
Sujipl. p. 811. n. 105-G (1824) ; Boisd., Spec. Gen. Lip. i. p. 345. n. 185 (1836) ; Doubl., Westw.
& Hew., Gen. Diurn. Lep. i. p. 13. n 8.3. t. 4. fig. 1. ? (1846) (.Jamaica) ; Gray, Cat. Lep. Ins.
Brit. .Urn. i. Pap. p. 23. n. 104 (18.52) ; id., Ll.'it Lep. Ins. Brit. Mus. i. Pup. p. 31. n. Ill
(1856) ; Weidem., Proc. EiU. Sue. Phihul ii. p. 147 (1863) (West Indies) ; Felder, Verh. Zaol.
But. Ges. Wieii xiv. p. 309. n. 288 (1864) (Jamaica) ; Kirby, Cat. Viimi. Lep. p. 542. n. 160
(1871) ; Rutherf., Enl. J/o. Mag. xv. p. 28 (1878) (Jamaica, habits) ; Butler, Prne. Zuol. Soc.
LumJ. p. 481. n. 3C (1878) (Jamaica) ; Gosse, Pror. Enl. Soc. Land. p. 55 (1879) (larva ; habits);
Oberth., Et.d'Ent. iv. p. 74. n. 226 (1880) (Jamaica); Lucas, Bull. Soc. Enl. Fram-e p. Gi
(I88H) (Jamaica) ; Aaron, Camid. Ent. xxv. p. 258 (1893) {S. Domingo, Cibas range ; Jamaica,
Sulphur R. and Devil's R. ; Ijmmoea food-plant of larva) ; Panton, Journ. Ins. Jamaica 1.
p. 375 (1893) ; Pox Oc Johns., Ent. News iv. p. 3 (1893) (Pt. Antonio, Jamaica) ; Haase,
Untersuch. Mim.icri/ i. p. i) J {lSd3) ; Taylor, Ent. News v. p. 101 (1894) (de.scr. of larva and
pupa; on "Water Wood," prob. Chimarrhis ci/niosa, Nov. 27) ; id.. Trans. Ent. Soc. Land.
p. 409 (1894) (larva and pupa) ; Kirby, in Allen'.s Nat. Libr., Lep. Butt. ii. p. 282 (1896)
(Jamaica) ; Swains., Journ. N. York Ent. Sue. ix. p. 77 (1901) (larva, on Thespesea populnea) ;
Robins., Ent. Neics xiv. p. 19 (1903) (capture of a series) ; Swains., Proc. Ent. Soc. Lond. p. 55
(1904).
Felder, and also Haase, regarded this species as a near ally of P.
machaonides. However, it is closely related to P. garamas (= asclcp/m), near
which it was placed by Oberthlir. The shape of the hindwing is practically the
same in P. Iiomeriis, garamas, wrirscewiczi , and judicaiil. The serration of the
costal margin of the forewing is not quite so strong in P. komerus and garamas as
in the other species. The agreement in pattern between P. homerus, toarscewiczi,
cacicus, garamas, etc., is easily perceived in spite of the specific differences. The
white spot on the palpus so characteristic for P. aristeus, P. cleotas, and allies is
also preserved in P. homerus. The cell of the hindwing of P. komerus is short and
broad, being widest near the apex, as in many specimens of P. garamas syerha.
Genitalia : S . Tenth tergite broad proximally, strongly narrowing towards
apex ; sternite on each side with short projections, the anterior one being somewhat
acuminate, anally in middle a conical tooth as in P. aristeus ; harpe similar to that
of P. aristeus, nearly as long as the clasper, linear, flat, truncate at apex, the two
angles produced into a straight thornlike process, the lower one being sometimes
very short. ?. Edge of vaginal orifice anteriorly raised into a short, broad,
smooth, rounded tubercle, which is concave on the posterior side; on each side far
back a high ridge which separates further i'routad into two ridges, the inner one
ending in a long acute jirocess as iu P. aristeus and allies.
Early stages described by Taylor, I.e.
Hab. Jamaica ; Santo Domingo (interior of, teste Aaron, I.e.).
In the Tring Museum 3 cJJ, 6 S ? and 1 pupa.
In coll. H. J. Adams a fine series of 7 66,H ? ?.
( 650 )
119. Papilio warscewiczi.
(J. Papilin warscexciczi Hopffer, Stetl. Eiit. Zeil. xxvii. p. 29. n. 11 (1866) (Bolivia).
(?. Underside of abdomen densely covered with tawny-olive hair-soales,
similar hairs on the breast. Costal margin of forewing serrate ; two rows of sjiots
on 7/ppersidc, no cell-patch, discal row sometimes partly obsolete, the two rows
almost parallel, the snbmarginal row augulate at SC^, the first two spots being more
distal than the third spot, the snbmarginal spots larger than the discal ones, except
the two npper ones. Hind wing with long obtuse tail, teeth M' and M- being
also i^rolonged ; snbmarginal spots large, creamy or yellowish, shaded with black, a
discal band or a row of spots of the same colour, distant from cell, the spots often
obsolete, a postdiscal row of blue spots.
Underside resembling P. cleotas in so far as there is the same kind of
markings, thongh differently developed ; forewing black from base to hinder angle
and anteriorly beyond cell (as far as the wing is covered by the hiudwing, when
at rest), apical area cinnamon like the hindwing ; a creamy or yellow cell-patch ;
discal and snbmarginal spots larger than above, the npper snbmarginal spots often
enlarged to broad patches. Hindwing : a broad continuous discal band, its inner
edge even, slightly curved, the outer edge crenate, the band creamy white, often
washed with cinnamon ; snbmarginal spots the same colour, large, ujjper ones often
much enlarged, extending from edge of wing to discal band, being either merged
together with the latter or separated from it by their cinnamon crescents ; hairs
in posterior area tawny-olive ; shoulder often creamy.
? not known.
Genitalia : Tenth tergite very long, snbijrismatical, the underside being some-
what roof-shaped e.xcept at apex, which is flat above and below, appearing sinuate
V)eneath in lateral view ; sternite strongly chitinised laterally, separated by an
oblique groove into a larger anterior portion which is produced into a proximal and
a distal tooth, both more or less denticnlated, and into a smaller posterior portion
which is obtuse. Harpe similar to that of P. aristeus, flat, with two apical
jjrocesses.
Early stages not known.
Hab. Ecuador to Bolivia.
Three subspecies.
a. P. warcsceiciczi jelskii Oberth. (1881).
Pcqjillojrhkli OberthUr, Ef. d'Eiit. vi. p. 113. n. 1. t. 20. fig. 6 (1881) (Tatnbillo, Peru) ; Dognin,
Lep. Lojti p. 14 (1887) ; id., If. p. 37 (1801).
cj. Similar to the next form ; discal spots of forewing ill-defined ; the under-
scales being all black, except in the spots at costal margin. Discal band of
hindwing rather strongly curved ; ill-defined ; bine spots narrow, curved ; last
snbmarginal spot orange-red.
Discal spots on underside of forewing large and brown, distal marginal area
much widened before middle. Posterior brown postdiscal spot W — M' of hindwing
smaller than in the other forms ; snbmarginal s{)ot M' — M- orange in middle.
Genitalia (Loja specimen) : Tenth tergite broad, nou-spatulate, apex less curved
downward than in the Bolivian form ; first and second tooth of sternite close
together, the interspace being much smaller than that between the second and third,
the first tooth long, strongly denticulate. Upper prong of harpe much longer than
the lower one, both straight.
( 651 )
Hah. Loja, Ecuador ; North and North-West Peru : Chachapoyas, Tambillo.
lu the Tring Museum 1 S from Loja.
In coll. Oberthiir several males from Tambillo and Chacha2)oyas.
In coll. Doguin a series from Loja.
b. P. icarscewiczi mcrcedes subsp. nov.
Papilio ivarscewirzi, Hopffer {tion id., 18G6, err. det.), Stett. Ent. Zeit. xl. p. 54. n. 30 (1879)
(Chanchamayo) ; Druce, Proc. Zool. Soc. Land. p. 246. n. 20 (1876) (Cosnipata).
(J. Closely agreeing with the preceding form ; discal spots of forewing abore
on the whole larger ; blue spots of hiudwiiig broader ; brown distal marginal border
of ((//(/(VAvWe of forewing narrower before middle, and discal spots R-' — M- smaller;
brown postdiscal band of hiudwing on the whole broader, especially spot R — M',
submarginal spot M' — M^ without orange scales.
Genitalia: Tenth segment similar to that of the ue.xt form, the second tooth of
the sternite standing close to the third ; both prongs of the harpe straight, usually
of the same length as the lower one, a little shorter than the upper.
Hab. Central Eastern Peru : Huanuco.
In the Tring Mnsenm U SS from: Pozuzo, Hnanuco, 800—1000 m. (W.
Hotfmanns), ti/pe ; Chanchamayo (AV. Hoffmanns) ; Cushi, Huanuco, 182U m.
(W. Hoffmanns) ; Huancabamba, Junin (Bottger).
c. P. tvarscewiczi tmrscewiczi Hopfif. (18(56).
Papilio loarscemczi Hopffer, Stett. Ent. Zeit. xxvii. p. 29. n. 11 (1866) (Bolivia) ; Hewits., Exot.
Butt. IV. Pap. t. 10. fig. 30. (J (1869) (Apolobamba, type of somteiisif) ; Kirby, Cat. Diurii. Lep.
p. 537. n. 128 (1871).
Papilio soratensis Godman & Salvin, Aim. Mag. N.ll. (4). ii. p. 152. n. 26 (1868) (Apolobamba).
Papilio wardscevjicsi (!), Weeks, Illuntr. Diurn. Lep. p. 20 (1905) (Chulumani).
(?. Markings oiupperskh on the whole paler than in the previous forms, less
yellow. Discal spots of forewing much smaller than the submarginal ones, often
minute and some of them absent. Discal baud of hindwiug broken up into spots,
sometimes vestigial.
Discal band of hindwing below usually entering cell, in many specimens this
band and the submarginal spots washed with brown.
Genitalia : Apex of tenth tergite more strongly curved downwards than in the
other forms ; sternite as in mercedes, dentition individually variable. Upper prong
of harpe strongly curved, often several minute teeth on and in between the
two prongs.
Hab. Bolivia and South-East Peru.
In the Tring Museum 22 S S from: R. Inambari, S.E. Peru, lOOO m.,
July 1900 (Simons); Chirimayo, 1000 ft., July 1901, R. Slncnri, June 1901,
2500 ft., S. Domingo, 0000 ft., January 1001, and Limbani, April 1904, Carabaya,
S.E. Peru (G. Ockenden) ; Charuplaya, Bolivia, 1300 m., May 1901 (Simons);
S. Antonio, Bolivia, 1800 m., March— June 1896 (Garlepp) ; R. Tanampaya
(Garlepp) ; R. Songo, Prov. Yungas (Garlc])p) ; Bneyes.
120. Papilio cacicus Lucas (1852).
Papilio cacicus Lucas, in Guerin, Reo. Zool. (2). iv. p. 132 (1852) (Colombia).
(J. Closely allied to P. tvarsceivkzi ; forewing with three bands on upperside :
a creamy yellow median band interrupted at lower angle of cell, the posterior
( 652 )
portion being continuous witii a variable cell-patch, which is rarely absent, a
postdiscal row of glaucous bnif spots, and a row of snbmarginal spots of which
the posterior one, two, or three are orange, cell-patch sometimes washed with
orange. Hind wing with a broad creamy or creamy yellow median band, tapering
behind, always entering cell ; a postdiscal row of bine spots ; and a snbmarginal
row of creamy ones, sometimes shaded with black ; last spot orange as a rule,
usually merged together with the marginal one.
Underside : apical area of forewing and the hindwing cinnamon-brown.
Forewing : discal band and cell-patch broader than above, npper snbmarginal spots
absent or vestigial. Discal band of hindwing milky white or creamy, wider
posteriorly than above, externally bordered by black bars or crescents, except
towards costa ; snbmarginal spots vestigial.
?. Trichromatic ; one form like male, bnt wings broader, band of hindwing
wider. The other forms with large cell-patch and complete discal band on fore-
wing which are tawny-orange or white ; hindwing without discal band on upperaide,
the band vestigial on underside.
Genitalia : c?. Tenth tergite long, curved, transversely dilated beneath at the
base, apex sharply pointed ; sternite divided at each side by a transverse groove
into two i)rocesses, the first being strongly chitinised and pointed, and the second
being paler and obtuse. Harpe vestigial, being represented by a flat piece of
chitin of about half the length of the clasper, not separated from the latter, tip
acuminate. ?. Edge of vaginal orifice and the area around strongly chitinised,
the cavity in which the orifice proper is situated ajipearing heart-shaped, the edges
of the cavity meeting behind the orifice, forming here a brown tubercle; within this
cavity several transverse folds, and in front of the cavity also some transverse folds ;
no processes.
Early stages not known.
a. P. cacicus cacicus Lucas (1852).
Papilio cacicus Luca'i, I.e. ; Doubl., Weatw. & Hew., Gen. Diiirn. Lej>. ii. p. 529 (1852) ; Gray,
Cat. Lep. Ins. Brit. Mus. i. Pap. p. 40. n. 20.5 (1852) (Quito) ; id., List Lep. Ins. Brit Miis. i.
Pap. p. 55. n. 214 (185G) ; Lucas, in Casteln., I'oy. Ania: Siul, Lep. p. 199. t. 1. fig. 3. ^ (1857)
(Colombia); Felder, Verli. Zoo!. But. Ges. Wien xiv. p. 313. n. 340 (1804) (Ecuador ; Colombia) ;
Kirby, Cut. Diuru. Lep. p. 537. n. 127 (1871) (Amer. mer.) ; Oberth,, /:.■(. d'Enl. iv. p. 74. n. 224
(1880) (" Brazil" error loci) ; id., I.e. vi. p. 4. n. 8 (1881) (Muzo, J ) ; Honr., Deutsche Ent. Zeit.
p. 223 (1889) [zntUlaehi = ? of cacicus) ; Dewitz, iliid. p. 224 (1889) ; Hahnel, Iris iii. p. 194
(1890) (M.'rida) ; Maass. & Weym., in Stubel, Reisen S. Amer., Lep. p. 31. n. 132 (1890)
(Colombia) ; Haase, Cntersueli. Mimicry i. p. 94 (1893) ; Haensch, Berl. Ent. Zeiischr. xlviii.
p. 151 (1903) (Balzapamba, W. Ecuador).
Pajnlio zaddachi Dewitz, .Mitth. Miinch. Ent. Ver. i. p. 85. t. 2. fig. 1 (1877) (Colombia); id., Deutsche
Ent. Zeit. p. 332 (1888) (distinct from cacicus ; against Honratb).
Papilio cacicus ab. J zuddachi, Honrath, Berl. Ent. Zeit. xxxii. p. 253 (1888) (a second specimen
of tbis form of $ from Colombia ; not specifically distinct from cacicus).
(?. There is considerable variability in the size of the markings. Forewing :
the cell-patch, which is occasionally washed with orange, is sometimes a broad baud
reaching across the cell, while it is reduced in other specimens to a small sjwt,
being altogether absent in one of our specimens from Bogota. In this Bogota
individual the band of the hindwing is more distal in position than usually, jnst
entering apex of cell.
? . Trichromatic.
a. ? -f. cacicus similar to the male.
( 653 )
h'. ? -f. zaddaclu Dewitz, I.e. Discal band of forewing continnons, not being
interrupted at apex of cell, and like tlie large patch in cell tawny-orange ; hindwing
without discal band on iipperside, the band being vestigial below.
c'. ?-f. vais nov. Like the preceding, but the discal band and cell-patch
of forewing white. Coll. Oberthiir.
Hab. Merida ; Colombia ; Ecuador.
On comparison of a larger material than we have examined it will perhaps
be found that there are several subspecies in the districts mentioned. In M6rida
specimens the last snbmarginal spot of the forewing, on upperside, appears to be
always (?) absent or vestigial ; in our two individuals from the Cauca valley and the
Eio Dagua the first spot of that row is obsolescent and the last small, the band
of the hindwing being also more evenly tapering than in Bogota specimens.
In the Tring Museum 16 c? (?, 1 ¥ , from : Merida, Venezuela, June 1898 (Briceno);
"Bogota"; Bogota to Pandi, December 1896, dry season (Dr. Burger); Pereira,
Cauca ; R. Dagua, west coast of Colombia (W. Rosenberg) ; Ecuador.
In coll. Oberthtir from Ambato and Balsapamba, Ecuador.
b. P. cacicus inca subsp. nov.
Papilln cacicus, Hopffer, Slett. Ent. Zed. xl. p. 54. n. 29 (1879) (Peru ; this form ?).
(?. Upperside. — Forewing : cell-patch much more oblique than in the preceding,
forming a very acute angle with M on basal side ; last three sjjots of postdiscal
row close to the submargiual spots ; first submargiual spot vestigial, second small,
the others also smaller than in c. cacicus, partly shaded with black, last one
orange-red. Hindwing : discal band more convex distally than in the preceding
subspecies, broad anteriorly, strongly tapering ; blue postdiscal spots large ;
submargiual spots narrow, shaded with black, no orange anal sjiot ; black discal
area wider centrally than in the preceding, the wing being longer in middle ; tail
narrow, non-spatulate.
Underside. — Forewing : discal band extending to base of M-', its proximal
edge almost on a level with the edge of the cell-patch, which is more proximal than
in the previous subspecies ; black discal area much wider in middle than in front
and behind ; upper jjostdiscal spots large, contiguous, forming an oblique band,
posterior spots of that row thin, close to the snbmarginal spots. Hindwing :
black discal bars sharply defined, luniform, separated from the discal band by a
narrow interspace of the brown ground-colour; a minute orange-red anal snbmarginal
spot, separated from fringe.
Genitalia: Anterior process of tenth tergite shorter than posterior one ; apex of
harpe not raised into a small tubercle or point.
Hab. Eastern Peru : Upper Rio Toro, Chanchamayo district, August — September
1901 (Simons).
1 c? in the Tring Museum ; not seen in other collections, but the specimen
recorded by Hopffer, I.e., may belong here.
121. Papilio euterpinus Godm. & Salv. (1868).
Pap'dh eulerjihms Godman X- Sulvin, Ann. Maij. .V. //. (4). ii. p. 150. n. 24 (18G8) (GuaJalquiza,
Ecuador) ; Hewits., Ejcot. Jlnlt. iv. Pap. t. 10. f. 31. J (18G9) (type in G. & S. coll.) ; Kii-by,
Cat. Ditirn. Lrp. p, 538. n. 143 (1871) ; id., Trans. Ent. 8oc. Land. p. 351 (1881) (Chiquinda ;
moist places at a high elevation ; deacr. of worn $ ) ; Staud., Exot. Tnijf. p. 15 (18S4) ; Hiase,
Untersuck. Miniicry i. p. 94 t. 11. fig. 83. (J (1893) (Ecuador.— neuration erroneous in fig.) ;
Haensch, Berl. Ent. Zeitschr. xlviii. p. 153 (1903) (Sa. Inez, R. Pastaza, 1250 m.).
( 654 )
This jiciuliar sjiecius Las generally been associated with P. zagreus. However,
its true position is near P. cacicus and P. loarscewiczi, with which it agrees in the
short antenna, serrate costal margin of the forewing, the short cell of the hindwing,
the origin of the subcostal vein of the hindwiug from close to the base of the
subbasal cellule, the hairs of the underside of the body and hindwing, and in
the genitalia (of 6, ? not examined). The hindwing has lost the tails and the
markings, only the submarginal row being represented by two or three orange
spots situated from anal angle forwards, the row heing sometimes continued by
some olive-bnfF spots. The cell-patch of the forewing and the three discal spots
W — M- correspond to the respective spots of P. cacicus, the two posterior discal
spots being prolonged as in P. aristeus bitias, nsnally touching the submarginal
dots, which are homologous to the posterior orange submarginal spots of P. cacicus.
On the UDclerside the pattern of the forewing agrees often rather closely with that
of P. cacicus cacicus ?-f zaddachi in tlie discal row of patches being continued
to costal margin by some orange and creamy spots situated at the distal side of
the cross-veins.
Genitalia : tj. Tenth tergite very broad, rounded at ape.x, beneath at base
transversely dilated as in P. cacicus ; sternite on each side with only one long
acute process, which is somewhat angulate on the anal side ; this process is
homologous to first and second process of P. cacicus, the first process being
shifted nuad in P. euterpinus and the two processes having become merged together,
the small projection on the hiuderside of the long process of P. euterpinus being
homologous to the second process of P. cacicus. Harpe as in P. cacicus, but a
little longer, not so completely fused with the clasper, th« edges and especially
the acuminate apex being free.
Early stages not known.
Hab. West Colombia ; Ecuador ; North Peru.
In the Tring Museum 2 Si from: Sante Inez (R. Haensch) ; Zamora
(0. T. Baron).
In coll. Oberthiir 7 c? <? from : Honda, AVest Colombia ; Moyobamba and
Chachapoyas, Peru.
A ? in coll. H. Grose-Smith from Chiqninda.
SECTION III.— KITE-SWALLOWTAILS.
(For characters see p. 434.)
There is considerable divergency among the New- World Kite-Swallowtails, the
American forms presenting a greater variety in structure, shape, and pattern than
the Old- World species. The two types of hindwing, the tailed triangular type
and the non-tailed rounded type, which we find in the Eastern Hemisphere, obtaia
also in America, but here the two types are connected by a third type, a more or
less rounded hindwing bearing a thin tail {dolicaon and allies). The development
of the subcostal veins of the forewing exhibits interesting features in this Section of
Papilios. In one of the American species the first subcostal is lost {bellerophon) —
the only instance among true Swallowtails of a reduction in the number of veins.
We take as the most generalised state that venation in which the five subcostals
are all present, and end free in the margin of the wing. This state obtains in a
large percentage of American species, while only two Old- World species have the
same neuration — namely, the Palaearctic poclalirius and the Australian leostkenes,
( 655 )
none of the African Kite-Swallowtails having all the subcostals free. The next
phyletic stage is represented by those sjiecies in which the first subcostal is
anastomosed with the costa, and in the third stage both the first and second
snbcostals aro joined to the costa. Both the less and the more advanced specialisa-
tions are observed among the American Papilios, some of the species exhibiting
a remarkable flnctnation in this respect among the individuals.
The scent-organ in the abdominal fold of the hiudwing of the males is another
structure which exhibits interesting variety among these Papilios, and striking
variation within many of the species. The scent-scales differ in some of the
species enormously, while in others the differences are less marked, and in others
again the scent-scales are apparently identical. The geographical variation of the
scent-organ is very marked in several species of this Section. In quite a number
of American and Old-AYorld species the scent-organ is vestigial or absent, though
the fold remains always slightly indicated at the base of the wing.*
The genitalia of the American Ivite-Swallowtails are ia some groups of great
nuiformity, while they are different in every species, and sometimes even subspecies,
in other groups. One organ, the tenth abdominal segment of the male, is however
of remarkable constancy all through the American species of this Section, with
the exception of one species (P. celfidori), which one would not expect to deviate
so much from the normal, considering that the aberrant-looking mimetic forms have
the tenth tergite of the same trifid type as P. protesilaus, dolicaon, marcellus, etc.
This exception from the normal is instructive. It is one of the instances we meet
with so frequently among Lepidoptera of the simplification of an organ by reduction,
the tenth tergite of P. celadon having become simple in consequence of the loss of
the side-lobes.
The edge of the eighth abdominal tergite of the male is very characteristically
modified in the American Kite-Swallowtails, the smooth erect scaling at this edge
not being met with in other American Papilios.
We know deplorably little about the larvae and pupae of the various species.
There are some points in the variability of the species, especially the mimetic
ones, which breeding from a female would put beyond dispute. Under P. phaon
and P. lysithous we have united quite a number of differently coloured forms which
other authors have considered distinct species. Though we have no doubt about
these insects being polymorphic, it is nevertheless desirable that this polymorphism
be investigated by breeding from the eggs.
The following two generic names have American species as types :
Iphiclides Hiibner, Verz. belt. Schmett. p. 82 (1818 ?) (type : dolicaon).
Eurytides Kirby (ex Hiibner, indescr.), in Allen, JS'at. Libr., Butt. ii. p. 272
(1896) (type : dolicaon).
Since some Old- World species are apparently close allies of American species,
and may perhaps come into one or the other American Group when we classify
all the Papilios, we abstain from giving a detailed characterisation of the two
Subsections in which the American Kite-Swallowtails fall.
Subsection E.
Underside of wings with red spots at the base, or hiudwing with a red line
parallel to abdominal margin, extending from costal margin before middle towards
the anal angle. Snbcostals of forewing always free.
* The result of our investigation into the variation of this organ will be the subject of a special pafer.
( 656 )
Here come three gronps :
a. Underside with red spots at base, either on both wings
or only on hindwing Li/sit/ious Group.
b. Underside of hindwing with red line parallel to
abdominal margin, extending from costal edge
proximally of middle in the direction of the anal
angle, being bordered with black on both sides at
least at costal margin Marcellus Group.
c. Bed line on underside of hindwing as before, but
bordered with blade only on one side . . . Protesilaiis Group.
Subsection F.
Underside of hindwing with a red or tawny band on disc parallel (or nearly)
with distal margin, commencing beyond middle of costa, or no red or tawny band
or spots on nnderside.
cl. Hindwing below with red or tawny line (or row of
spots) parallel with distal margin ; first subcostal
of forewing free Thyastes Group.
e. Hindwing lielow without red line ; SC^ of forewing
anastomosed with C .... Dolicaon Group.
XIII, Lysithous Group.
With the exception of P. asiiis, which stands a little apart, the species of
the present group are all closely related with one another. They resemble in
pattern various members of the Aristolochia-Papilios, with which they have been
associated by nearly every author, Haase alone having recognised their close
relationship witii the Marci'llus Grouii. The red basal spots on the underside of
the wings are a distinguishing character peculiar to these mimics. The species
are modifications of an ancestral form which, in the shajje of the hindwing, the
number of red basal and subbasal spots on the same, and the development of the
scent-organ in the abdominal fold, resembled P. asius. This insect has preserved
the triangular shape of the hindwing jjeculiar to the Marcellus, ProtesHaus, and
Leucaspis Groups, while in most other species of the Mimetic Group the hindwing
has assumed a more or less rounded shape. Of the five red markings in the basal
area of the underside of the hindwing of P. asius the otlier mimics have preserved
only four, three or two, the sjwt between C and SO not being marked in any other
species. One should exj^ect this spot to be occasionally vestigial in the species
with four red sjiots {P. ilus, branclius, tlii/mbraeus, etc.), but we have not come
across a specimen in which such a spot is indicated.
The scent-organ of P. asius is far less reduced than that of the other species.
With the exception of P. asius, the abdominal edge of the hindwing {S) is not
curved upwards, and the scent-scales are restricted to the basal third or half, forming
a narrow greyish stripe. In several species (P. protodamas, pausanias, and xt/nias)
the scent-organ is altogether lost, while in P. curi/leon it is lost only in the
subspecies from East Ecuador. The scent-scales of P. asius are similar to those
of P. agesilauSjhcmg sjiindle-shaped, with both poles produced into a threadlike
( 657 )
process. In the other species the scent-scales resemble narrow ordinary non-dentate
scales.
The genitalia are of great sameness, the differences between the species, if there
are any constant difterences, being very slight. They agree also closely with the
genitalia of P. p/n'lolai/s.
The larva is bright-colonred, bearing nnmerons longitudinal bands and behind
the middle a V-shaped dorsal patch, the thorax being dotted ; no spiuelii;e
tnbercles. The pupa is short ; the abdomen is widest in middle, being somewhat
barrel-shaped.
Sixteen species :
a. Forewiug below without red spots
Forewing below with one or two red spots
b. Hindwiug below with two red spots at costal margin
Hindwing below with one red spot at costal margin .
c. Hindwiug below with five red sjwts at base, there being a
spot between C and SC ..... .
Hindwing below with four red spots at base, there being
no spot before St! ...... .
d. Hindwing with red band on disc .....
Hindwing with straw-coloured band across both wings
e. Forewing all black, or a white spot or patch on disc at
and near lower angle of cell .....
Forewing with large grey patch M-— SM^, besides some
other greyish or huffish patches
y. Red band of hindwing distant from cell ....
Red band of hindwing close to cell
ff. Hindwing without markings on ui)perside, except a row
of small white snbmarginal lunules ....
Hindwing with red, or huffish green, or straw-colonred
band or patch
//. Posterior segments of abdomen and sterna with red spots
Thorax and abdomen with bnffish markings
I. Forewing with a snbmarginal row of huffish straw-colonred
spots, discal patches absent or vestigial
Forewing with large patches on disc, either in centre or
before inner margin ; hindwing with red discal patch
or band Species No. 120.
j. Hindwing with pale pink band on npperside from R' to
abdominal margin
Hindwing with greyish green or greyish blue discal band,
or the whole central area this colour, or the band
carmine red and extended to costal margin
L Forewing with two red spots at base|
Forewing with one red spot at base standing at costal
margin, the cell spot being absent ....
/. Hindwing with tail ; posterior basal spot of hindwing
produced to form a streak
Hindwing without tail ; posterior basal spot of hindwing
a dot ......•■..
i.
h.
c.
9-
Species No. 137.
(1.
e.
Species No. 123.
/■
Species No. 132.
Species No. 134.
Sjiecies No. 133.
Species No. 122.
h.
I.
Species No. 124.
J-
Species No. 127.
Species No. 125.
I.
Species No. 130.
Species No. 13i.
( 658 )
m. Hindwing witli four basal spots on underside, there being
two spots at costal edge Species No. 135.
Hiadwing with two or three basal spots, one only at costal
edge .......... n.
n. Patch of uppersidc of forewing pale green (c? ; ? uot
known) Species No. 130.
Patch of upperside of forewing grey or white, or absent . o.
0. Hindwing rounded, red spot R^ — M' of upperside usually
larger than spot M'— IP Species No. 128.
Anal angle of hindwing more produced, red spot W — M'
of upperside not larger than spot W — M- . . . Species No. 129.
122. Papilio pausanias Hew. (1852).
Pajiilio pausaidas Hewitson, Tr. Eni. Soc. Land. (2). ii. p. ii. t. G. fig. 2 (1852) (British Guiana ;
Ega).
c??. Abdomen with broad huffish lateral streak. Forewiug proximaily and
nearly the whole upper surface of the hindwing metallic greenish blue ; a straw-
or primrose-yellow area on forewing, consisting of a large cell-patch and two large
discal patches R' — M-, there being often a small additional spot before R^ and
another behind M-. Hindwing without discal markings, bnt with a row of more
or less distinct white curved submarginal bars ; shape of hindwing variable, distal
margin (apart from dentition) often straight, especially in female.
Underside brown, without distinct black cell-streaks ; hindwing with three red
basal spots which are often developed to streaks ; a row of red postdiscal lunules,
also often produced basad, forming more or less distinct streaks.
Scent-organ absent, vein SM- being clothed with metallic blue scales, only a
few of the scales being greyish.
Genitalia not essentially different from those of P. protodamas ; the dorso-
ventral dentate ridge of the harjjc ventrally a little more curved, apical lobe a little
larger, and the teeth beneath this lobe rather more numerous.
Early stages not known.
Hub. Costa Rica southward to Bolivia and the Brazilian province of Goyaz.
Three subspecies.
a. P. jjausaniiis prashuis subsp. nov.
c?. Similar to clcomhrotiis. I'pperside of wings more green in tint, this
metallic colour also more extended, the costal area of the hindwing being hardly
k'ss metallic green tlian the disc. Forewing rather shorter ; discal patches
longer, reaching three-fourths the way from cell to distal margin. Hindwing :
white submarginal spots minute, upper ones vestigial, practically absent both above
and below ; red basal sjwts of underside smaller than in cleombrotus.
Hal). Carillo, Costa Rica, June — July 1003 (C. Underwood); one S in the
Tring Mnseum.
h. /'. pausanias cleombrotus Streck. (1885).
Pa/iiliii rleomlrotim Strecker, Proc. Ac. Xiil. Se. Philtid. xxxvii. p. 175 (1885) (" Upper Amazons"
error loci) : id., Lep. Rliap. Het. Su/i/,l.. iii. p. 17 (1900) ('Pebaa ' false).
The locality given for tins form by Strecker is erroneous. We know the
subsiiecics only from the West Coast of -Colombia, but it occurs presumably also irj
( f.69 )
Panama, whence Strecker had received Lepidoptera. The description fits our
specimens from the Rio Dagna.
(?. Pale apical j)atcli of forewing absent; posterior area of forewing and distal
area of hindwing more extended metallic green-blue, costal area of hindwing
practical]}- black. Underside deeper black-brown than in P. p. pausanias ; red
basal spots of hindwing not prolonged to streaks, no pale or reddish streaks on
disc; red postdiscal spots, especially the last one, larger than in P. p. pausanias.
Forewing occasionally with vestiges of creamy submarginal spots on upper as well
as on underside.
? not known.
Ilalj. Rio Dagua, West Colombia.
In the Tring Mnsenm 0 c? <? (W. F. H. Rosenberg).
2 S <S from Juntas, R. Dagua, in coll. Oberthilr.
c. P. 2iausanias pausanias Hew. (1852).
PapUio pausanias Hewitson, I.e. (1852) ; Doubl., Westw. & Hew., Gen. Diurn. Lep. ii. p. 529 (1852);
Gray, Cat. Lep. Ins. Brit. Mns. i. Pap. p. 09. n. 313 (1852) (Ega ; var. a. " Honduras and
Guatemala," no such specimens in Brit. Mus.) ; id.. List Lep. Ins. Bnt. Mm. i. Pap. p. 80. n. 330
(1856) (Ega ; Demerara ; " var. a " e.xcl.) ; Wall., Trans. Ent. Soi: Land. (2). ii. p. 255 (1854)
(Amazons ; banks of river.s) ; M('netr.,£H«;H. Corp. Anim. Mus. Petrojt., Lep. i. p. 6. n. 94 (1857)
(Brazil) ; Bates, Trans. Enl. Snc.Loml. (2). v. p. 335 (18G1) (Villa Nova ; Ega ; habits different
from those of its model, //e//VnH;«s rli/tia); id., Journ. Entnm. i. p. 224. n. T (18tJ2) (Upp.
Amazons as far down as Villa Nova) ; Felder, Verh. Zoo!. Bot. Ges. Wien. xiv. p. 299. n. 153
(1864) (Demerara; Amazons; Venezuela; Bogota; ■' Hondura*!, Guatemala" — error loci);
Kirby, Cat. Diurn. Lep. p. .524. n. 52 (1871j (Amer. mer. ) ; Druce, Proc. Zool. Soc. Land.
p. 245. n. 10 (187G) (Peru ; Chamicuras) ; Hopff., Slelf. Enl. Zeit. xl. p. 52. n. 17 (1879)
("Brazil," Surinam, Brit. Guiana, Venezuela, N. Granada, Bolivia ;—" Honduras, Guatemala"
error loci) ; Oberth., Et. d'Ent. iv. p. 97. n. 290 (1880) (Ecuador ; Santarem) ; Staud., Exot.
Tiidf. i. p. 12. t. 8 (1884) (Para to Peru ; Colombia) ; Maass. & Weym., in Stubel, Reisen
S. Amer., Lep. p. 24. n. 104 (1890) ( Villavicencio, 450 m.) ; iid., I.e. p. 31. n. 120 (1890)
(Colombia) ; Hahnel, Iris iii. p. 268 (1890) (Teffe) ; id., I.e. p. 285 (1890) (Pebas) ; Haase,
Untersueh. Miniiery i. p. 87 (1893) ; Michael, Iris vii. p. 213 (1894) (Sao Paulo de 01ivenv> ;
? seen).
Papilio herrnolaus Guende, Mem. Soc. Phijs. Hist. Nat. Gcuere xxii. p. 379 (1872) ( ? , Poi'to Cabello;—
now in coll. Obertliur) ; Hew., Pet. Nouv. Ent. p. 213 (1872) (= pausanias) ; Kirby, Utid. p. 809.
n. 52 (1877) (= pausanias) ; id., Cat. Diurn. Lep. p. 239 (1872) (= pausanias).
Papilio pausainas (!), Weeks, Illnstr. Diurn. Lep. p. 20 (1905) (Chulumani).
S ?. Apex ef forewing with large greyish patch, the black upper scales being
mostly absent from this patch. Underside of hindwing with pale discal streaks
which join the red postdiscal spots; red basal spots variable in length ; outline
of hindwing variable, in our Bogota female the hindwing short, being sharply
truncate, in other females hindwing not shorter than in male, or even more ronndcd
than in that sex.
Hab. Central Colombia to the Gnianas, southward to Bolivia and the province
of Goyaz in Brazil.
In the Tring Mu.senm 30 (?c?, 2 ? ?, from : " Bogota"; Ecuador ; La Vaelta,
Canra R., Orinoco, May 1904 (S. M. Klages) ; Rio Demerara; Surinam; Ega;
Pebas ; R. Uaupes, Upper R. Negro ; Iquitos (Stuart) ; Rioja, near Moyobamba ;
R. Chuchuras, affl. of R. Palcazu, 320 m. (W. Hoffmanns) ; Gnanay and Musbay,
R. Mapiri, Bolivia (Stuart) ; Jatahy, Goyaz,
( 660)
123. Papilio microdamas Bnrm. (1878).
PapiVio microdamas Burmeister, Z)fsr)'. Rip. Arf/enl. v. Li'p. p. G3. n. 5 (1878) (Corrientes) ; id., I.e.
Atlas p. 19. t. 5. fig. 8. ? (1879) , Oberth., Et. d'Eut. xii. p. 2. n. 5. t. i. fig. 3. S (1888)
(Cara^a, Brazil).
(? ? . Sexes similar. A band of }-ellowish IjiifF patches from costal marg-in of
forewiug to aual angle of hindwing, above and below, straight on forewing, curved
on hindwing, variable in tint, somewhat paler on hind- thau on forewing ; spot
within subcostal fork of forewing nsnallj' vestigial. Hindwing with a red anal
spot, which is larger above than below, and four red basal spots on iinderside, the
last continnons with a red line on abdominal fold ; forewiug without red spots.
Scent-organ : numerous narrow grey scales on SM%
Genitalia: <?. Apical lobe of harjje rather small, dorso-ventral ridge irrega-
larly and rather heavily dentate, central process dentate ; ventral process short.
Early stages not known.
Hub. Paraguay and adjacent districts of Argentina ; Caraca, Brazil.
lu the Tring Museum 5 c?(?, 2 ? ?, from : Patino Cu(5, Paraguay, February
(Montforts) ; Sapucay, Paraguay, September and December (W. Foster) ; Entre
Rios.
124. Papilio protodamas Godt. (1819).
Papilin 2>rotodumas Godart, E/ic. Melh. ix. p, 40. n. 45 (1819) (Brazil ; " (J " excl.).
Itliohahis hyperion Hiibner, Samnd. E.cot. Schm. ii. t. 114 (1822?) ; Burm., Descr. Rep. Argent, v-
Lep,, Atlas p. 6. t. 2. fig. 5 (1879) (larva and pupa).
Godart described as P. protodamas a specimen of the present insect, adding the
description of another insect which he believed to be probably the i of the former.
The first-described insect was, in our opinion, also a cj, but that point is of no great
importance. The name protodamas must be applied to the first insect, not to the
second, of which Godart says : " Nous avous actuellement ii parler d'un individu qui
ponrrait bien etre le male de celni-ci."
Htibner, I.e., figures on t. 114 the first iusect, to which he gives the new name
hi/perion, and on t. 115 the second insect, to which he applies Godart's na.mB proto-
damas. From the fact that Hiibner figures these two insects in the same order as
they are described by Godart, and that to one of them the name protodamas is given,
we conclude (1) that Hiibner knew of Godart's descriptions when he put the names
on his plates 114 and 115, (2) that Hiibner's plates 114 and 115 appeared after 1819,
the year of issue of Godart's work*), and (3) that hijperion sinks as a synonym of
protodamas, both names being proposed for the same iusect. In any case, we give
precedence to Godart's names, of which tlie year of publication is known, since there
is no means of proving that Hiibner's names have priority.
cJ ? . Dichromatic in both sexes, the one form possessing on the forewing a large
straw-coloured patch in the cell and two large patches on the disc, while the other
form has only a vestigial cell-jiatch and small discal patches ; intergradations
between the two forms are rare. Scales of discal spots of hindwing either entire or
more or less denticulate. Forewing without red spots at base on underside, and
hindwing with three red spots.
Scent-organ absent.
Genitalia: $. Dorso-ventral dentate ridge of harpe slightly curved towards
* Godart's name erander for the Brazilian form of P. ancltisiades appears on UUbncr's pi. 112, which
confirms the above statement.
( 661 )
base ventrally ; rounded apical lobe rather small, bearing only very few teeth on its
nnder surface ; apex of central process obliquely truncate, its ventral apical angle
acuniiuate.
Early stages figured by Burmeister, I.e.
a.' P. protodamas f. protodamas Godt. (1819).
Papilio protodamiH Godart, I.e. (1819) (Brazil ; "(J" excl.).
Ithobnhis hyperion Hiibner, Ir. (after 18UI) ; Kirby, ibid. ed. ii. p. 92. t. 327. fig. 1. 2 (I'JO— 1).
Papilioii hi/pei'ion. Hoisduva], Spec. Geii. Lep. i. p. 319. n. 159 (183G) (Brazil); Doubl., Lint Lep.
Ins. BrH. Mus. i. p. 14. (1845) (S. Amer.) ; id., Westw. & Hew., Gen. Diiirii. Lep. i. p. 20.
n. 236 (1846) (Brazil) ; Gray, Cat. Lep. Lis. Brit. .Mus. i. Pap. p. 69. n. 310 (18.'i2) ; id., List
Lep. Lis. Brit. .Mus. i. Pap. p. 79. n. 327 (1856) (Rio de Jan.) ; Felder, Verh. ZanL Bol. Ges.
Wien xiv. p. 299. n. 150 (1864) (Brazil) ; Kirby, Cat. Diui-n. Lep. p. 521. n. 26 (1871) (Brazil) ;
Staud., Exul. Tarjf. i. p. 12 (1884) (Brazil) ; Haase, Vntersucli. .Mimicry i. p. 87, and ii. p. 69
(1893) ; Biinningh., Ver/i. Ver. Xal. Unterli. B'amburij ix. p. 28 (1895) (Rio de Janeiro ;
S. Theresa and Nictheroy) ; Mabilde, Guia Pract. Borhol. R. Grande do Sul p. 44 (1896).
Pajnlio zonaras, Perty, Del. .\niw. Artie, p. 152. t. 29. fig. 3. 3b (1830-34) (Amazon, false?).
Papilio zmiaros{l) Kirby, I.e. (1871) (sub syn.).
c?. Forewing with two rows of spots, the upper oues merged together into
streaks ; an ill-defined patch in cell, more or less vestigial.
?. Streaks of forewing less distinct than in S, cell-patch absent from npper-
side (always ?).
Known from Bio Grande do Sul northward to Minas Geraes.
b'. P. protodamas f choridamas Boisd. (1836).
Papilio cJioridanias BokdaT^l, Sjjec. Gen. Lep. i. p. 318. n. 158 (1836) (Rio de Janeiro); Doubl.,
List Lep. Ills. Brit. .Mus. i. p. 14 (1845) (" West Indies, Demerara," error loci) ; id., Westw. &
Hew., Gen. Diurn. Lep. i. p. 20. n. 237 (1846) (Brazil ; " Guiana " error loci) ; Gray, Cat. Lep.
Ins. Brit. Mus. i. Pap. p. 68. n. 309 (1852) ; id., List Lep. Ins. Brit. .Mus. i. Pap. p. 79. n. 326
(1856) ("Demerara, West Indies," error loci) ; Felder, Verh. Zool. But. Ges. Wien xiv. p. 299.
n, 152 (1864) (Brazil ; "Demerara, Ind. occid.," error loci); Kirby, Cat. Diurn. Lep. p. 5^4.
n. 51 (1871) (Amer. mer.) : Capronn., Ann. Soe. Ent. Belij. xvii. p. 9. n. 8 (1874) (Copa Cabana,
Sept. ; Botafogo. Sept.) ; Burm., Deser. Rep. Argent, v. Lgp. p. 7. sub n. 14 (1879) (var. of
hjperinn) ■ Uberth., El. dEnt. p. 97. n. 297 (1880) (Brazil) ; Stand., Exot. Tag/, i. p. 12 (1884)
(N. Brazil ; Surinam) ; Haase, Untersuch. Mimicry i. p. 87, and ii. p. 69 (1893) ; B6nningh., I.e.
p. 28 (1895) (Corcovado).
S ?. Forewing with a large patch in cell and two on disc ; snbmarginal dots
variable in size and number, occasionally absent ; the streaks found in ajiical third
of wing of the preceding form usually absent from choridama.s or only vestigial.
This form is known to us from the provinces of Rio de Janeiro and Minas Geraes.
Hab. of P. protodamas : Minas Geraes southward to Rio Grande do Sul.
In the Triug Museum 11 SS, 1 ? , of f }jrotodamas, from : Minas Geraes;
S. Catharina ; Castro, Parana (C. D. Jones); Blumenan. 8 S S , '2 ? ? , of
f. choridamas from : Minas Geraes ; Copa Cabana, September ; Alto de Theresopolis ;
Rio de Janeiro (E. May).
125. Papilio phaon Boisd. (1836).
Papilio phuon BoisduvaJ, Spec. Gen. Lep. i. p. 319. n. 160 (1836) (Mexico ; Peru ;— coll. Oberthiir)
S ?. Sexes similar. Spots of breast and abdomen red, posterior segments of
abdomen with red lateral patches ; spot on palpus creamy. Pattern of wings very
variable; markings of uppergide of forewing usnally straw-colour or creamy, the
( 662 )
discal ones ofhiudwing gvey-blue, occasionall.v replaced by ;i red band. C/u/rr-
side brownish black ; no distinct black cell-streaks ; furewiiig without red basal
spot ; hind wing with three red sjjots at base, but the one in cell often vestigial.
Scent-organ : numerous small greyish scales on SM^
Genitalia : S. Apical lobe of harpe irregularly rounded ; dorso-ventral ridge
somewhat curved, dentate veutrally, the lower ventral angle of this ridge not
jirodiiced busad.
Early stages not known.
Two principal forms, each variable;
«'. P. ijhaon f. loc. xenarchus Hew. (1801).
Papilio xetianhus Hewitson, Exot. Butt. ii. Pap. t. 5. fig. 12. ? (1861) (Mexico) ; Felder, Verli.
Zofil. Hot. Gcs. Wioi xiv. p. 2'J<J. n. 147 (18G4) (Mexico) ; Kirby, Cat. Diiirti. Lcp. p. 524. n. 4«
(1871) ; Staud., Ei-ot. Tayf. i. p. 12 (1884) (Mexico) ; Godm. & Salv., Biol. Centr. Amer.,
Ehop. ii. p. 210. n. 33 (1800) (Jalapa) ; Haase, Vnlersuch. Mhnicnj i. p. 87 (18'J3).
Papilio cridamas Eeakirt, Proc. Ac. Xat. Sc. Philatl. xviii. p. 248. n. 25 (1866) (Mexico) ; Kirby,
/.'■. p. 524. n. 47 (1871) ; Godm. & Salv., I.e. p. 210. n. 34. t. 67. fig. 10. ^ (1890) (Mexico ;
Atoyac, Cordova, Omeaica).
cJ ?. A red band on liindwing, the patches composing the band either large and
contignous {xenarclii(>i) or smaller and separate {erklani'is).
There is a broad-banded male in coll. Godman and another in coll. Charles
Oberthiir, and two females in coll. Hewitson (Brit. Mus.). Narrow banded speci-
mens appear to occur more often. In a male in coll. Hewitson the red spots are
pro.ximally more or less extended grey-blue ; a similar transition to the next form
is in coll. Standinger.
The red-banded form is known only from Eastern Mexico, where it occurs
together with the next, being much the rarer of the two.
b'. P.phaon i. phaon Boisd. (1830).
Papilio pliann Boisduval, I.e. (1836) (Mexico ;— coll. Oberthiir) ; Doubl., List Lip. In«. Brit. Mus.
i. p. 14 (1845) (Honduras; Oaxaca) ; id., Weatw. & Hew., Gen. Diurn. Lep. i. p. 20. n. 239
(1846) (Me.vico; Honduras); Gray, Cat. Lep. Inn. Brit. Mus. i. Pap. p. 69. n. 311 (1852)
(Mexico ; Honduras ; Venezuela) ; id., List Lrp. Lis. Brit. Mus. i. Pap. p. 80. n. 328 (1856) ;
Reak., Proe. Put. Sue. Phihul ii. p. 141. n. 14. (1803) (Honduras) ; Felder, Vtrh. Zuot. Bol. Ges.
in™ xiv. p. 290. n. 148 (1864) ; Boisd., Cousicl. Lfp. Guatem. p. 6 (1870) (Me.xico ; Honduras ;
Costa Rica); Kirby, Cat. Diurn. Lep. p. 524. n. 48 (1871) (Mexico; Honduras); Obertb.,
Et. d'Eut. iv. p. 97. n. 298 (1880) (Mexico) ; Godm. & Salv., Trans. Eut. Soc. Loud. p. 126.
n. 241 (1880) (Sta. Marta) ; iid., Biol. Centr. Amer., Jihop. ii. p. 211. d. .S5. t. 67. fig. 9. genit.
(1890) (Mexico; Brit. Honduras; Guatemala; Honduras); Slaud., Exot. Taijf. i. p. 12. t. 8
(1884) (Mexico ; Guatemala ; Honduras ; Venezuela) ; Haase, Untersueh. Mimirnj ii. p. 69
(1803).
Papilio idopos Gray, Cat. Lrp. Lis. Brit. .Vus. i. Pa/i. p. 69. n. 312 (18.'.2) (Mexico ;— Mus. Brit.) ;
id., List Lep. Ins. Brit. Mus. i. Pap. p. 80. n. 329 (1856) ; Feld., Verh. Zool. Bot. Ges. Wien
xiv. p. 299. n. 151 (1864) (Mexico); Kirby, I.e. p. 524. n. 50 (1871) (Mexico); Obertb., Et. d'Eut.
iv. p. 117. n. 298''" (1880) (Polochic Valley ; = im}minjinatus)\ Staud., E.col. Taijf. i. p. 12
(1884) (Mexico) ; Godm. & Salv., I.e. p. 212. n. 38 (1890) (Mexico ; Brit. Honduras ; Guatemala ;
Honduras).
Papilio ulopas (!), Weidemeyer, Proe. Ent. Sor. PJiilad. ii. p. 148 (1863) (Mexico).
Papilio therodamns Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 299. n. 149 (18l>4) (Bogota; nom. mid.);
id., Reise Novara, Lep. p. 45. n. 34. t. 10. fig. e. $ (1865) (Bogota ;— Mus. Tring) ; Kirby, /.<■.
p. 524. n. 40 (1871) (X. Granada) ; Haase, I.e. i. p. 87 (1893).
Papilio metaphaon Butler, Trans. Ent. Sor. Loud. p. 434. n. 3 (1874) (Mexico ; coll. Kaden, now in
coll. Godman) ; Kirby, Cat. Diurn. Lep. p. 813. n. 370 (1877) ; Godm. & Salv., Biol. Centr.
( 663 )
Amer., Rhop. ii. p. 212. n. 37 (1890) (Mexico? ; Colombia) ; iid., !.r. p. 720. t. 111. p. 13. 14. ^
(im) {l,,i,r).
Pupilio phnonvax. iinmarginatusOheTihvx, Et. d'Ent. iv. p. 97. sub n. 298 (1880) (Mexico ; — coll.
Oberthiir).
Papillo pharaxGoAm3,n & Salv., I.e. p. 211. n. 3G. t. 67. fig. 8. g (1890) (Brit. Honduras ;— coll.
Godman) ; id., I.e. p. 729 (1901) (Mexico ; Coatepec).
S ? . Markings (band or large patch) on disc of hindwing greenish or bluish
grey.
Very variable, but not separable into more or less sharjjly defined forms. If
specimens agreeing with the types of ulopos, therodamas, metapkaon, phaon, and
pharax were treated under separate names as f. ulopos, f. therodamas, etc., a host
of other names would have to be proposed for the numerous individual varieties
which are not covered by those names. However, we do not think it necessary to
deal with the present assemblage of individuals under more than one name.
The forewing bears usually a row of creamy or straw-coloured submarginal
spots, the last ones being the largest ; sometimes there are, proximally of this row,
some large discal patches, while in other individuals the forewing is practically
devoid of markings. The hindwing has occasionally some small red dots distally of
the discal band in specimens from Mexico and Honduras, such individuals occurring
presumably in all the countries from Nicaragua to Blexico ; the discal band has
either develoj)ed to a large central patch which enters the cell to a more or less
great extent, or it is more baud-like, standing distally of the cell. The name
pharax is based on an individual from British Honduras in which the hindwing has
the band sitnated outside the cell and bears a red anal spot; ulopos { = immargi natus)
was proposed for specimens which have only small or vestigial spots on the forewing,
the band of the hindwing entering the cell ; Felder's individuals which he called
therodamas have the spots on the forewing well marked, and the band of the
hindwing placed distally of cell ; in the type of phaon there are submarginal spots
on the forewing, but no discal patches, and the band of the hindwing just enters the
cell ; in the tyjie of mctapihaon the central area of the hindwing is large, occupying
a good part of the cell.
In a male in coll. Hewitson the spots on the breast and abdomen are butfish
pink, not carmine red.
Hab. of F. phaon : Mexico to West Ecuador and Venezuela.
In the Tring Museum 42 c? c? of i. phaon from : Orizaba (Bilimek) ; Polochic,
Guatemala ; San Pedro Sula, Honduras ; Rio Dagua, W. Colombia (Rosenberg) ;
Bogota ; Tachira and Mocotone, Venezuela (Briceiio) ; Paramba, W. Ecuador.
126. Papilio euryleon Hew. (1855).
Papilio eui-ijleon HewitsoD, Exot. Bull. i. Pap. t. 2. fig. 6. ^ (1855) (New Granada— Mus. Brit.) ;
Felder, Rehe Novara, Lep. p. 44. n. 38. t. 6. fig. 4. <f (1865) (Bogota).
tf. Spots of breast and abdomen red ; posterior segments of abdomen with red
side-patches. Forewing, above, with buflfish grey area on disc behind cell,
variable in extent and position, either reaching to hinder margin or being separate
from it; sometimes a patch in cell ; occasionally a row of submarginal spots.
Hindwing : a discal band of red spots, more or less continuous, either restricted to
centre of wing or continued to abdominal margin, entering cell or separate from it,
the band often reduced to two spots, occasionally pinkish grey ; a row of admarginal
spots, upper one large, the others usually minute or vestigial, all grey or the
( 664 )
posterior ones reddish, these spots absent from one of onr specimens of P. eurijleon
clusoculis.
Underside TpaXer than npper ; no distinct black cell-folds. Forewing withoiif
red basal spots ; either entirely greenish brown, with basal and jiosterior areas
deeper in tint, or marked with one or two grey discal patches. Hindwiug : three
red basal spots, but the one in cell usually absent or vestigial, the costal one being
also often very small ; discal band of spots much more restricted than above and
pale j)ink in colour, sometimes vestigial or absent ; a row of red submarginal spots,
often vestigial ; a row of grey admarginal dots as above.
?. Forewing : a patch across cell and two patches R- — M' on disc white.
Band of hindwing much broader than in male, not broken uji into spots or incised
distally on the veins, all red, or jiinkish buff washed with red distally. In
P. c. clusocidis the female similar to male.
Scent-organ : basal third of vein SM- either densely covered with numerous
greyish scales, which are smaller than the metallic scales situated before and
behind this vein, or the scales on SM^ also metallic and not reduced in size, The
absence of the scent-organ in the subspecies inhabiting Eastern Ecuador is a
very remarkable character. The length of the jiortion of SM^ covered with scent-
scales in the other subspecies is not constant.
Genitalia: c?. Not essentially different from those of P. /mrmod/us and
P. pkion ; apical lobe of harpe large, asymmetrically rounded ; dorso-ventral ridge
dentate only ventrally ; central process at apex irregnlarly ronnded and denticulate ;
ventral process individually variable in width and length, sometimes triangular,
obtuse.
Early stages not known.
Hab. Costa Rica, southwards to Ecuador.
Five subspecies.
a. P. euryleon clusocuUs Bntl. (1872).
Papilio clusocuUs Butler, Cist. Ent. i. p. 85 (1872) (Costa Rica) ; id., Lep. Exot. p. 163. t. 58. fig. 2. jj
(1874) ; Kirby, Cat. Lep. Rhop. p. 812. n. 354 (1877) ; Butl. & Druce, Proc. Zual. .Sw. Loud.
p. 3G4. D. 3G8 (1874) (Costa Rica) ; Staud., E:cot. T<i(jf. i. p. 19 (1884) (Chiriqui) ; Godm. &
Salv., Biol. Cenli: Amer., Rhop. ii. p. 209. n. 32 (1890) (Costa Rica ; Panama).
6. Upper side : patches of forewing purer grey than in the other subspecies ;
discal patches extending in extreme specimens from SM^ forward beyond R', while
in the other extremes only one patch M' — M- and a small streak before M' are
present ; cell with or without patch. Hindwing : red band broad, always entering
cell, last two spots sometimes vestigial only ; spot SC-— R' very variable, sometimes
large, but usually small or absent ; rarely a dot before SC^ ; in one of our si)ecimens
a row of red submarginal spots confluent with the discal patches, no grey spots
in this specimen.
Underside : forewing with two or three grey patches.
? . Similar to male ; on forewing a creamy jiatch R'— M', and a smaller one
M' — M-, no spot in cell ; red patch of hindwing a little larger than in male.
Scent-organ ])resent.
Hab. Costa Rica ; (_!hirii]ui.
In the Tring Museum \b i 6 from : Carillo, Costa Rica, 3U00 ft., October 1901
(A. Hall) ; Carillo, June— July 1903, and San Jos6, Costa Rica (Underwood),
A female in coll. H. Grose-Smith.
(665 )
b. P. earyleon pithonius snbsp. uov. (PI. VIII. fig. .55. 58).
Papilio curylcon, Maassen & Weym., in Stiibel, Reisen S. Amer., Lep. p. 36. n. 34 (1890) (La Plata,
Cauca, 1000 m.).
S. Upperside : patch of fore wing very much reduced (tj/pe), or as large a.? in
the following subspecies. Hindwing with three to five red spots which stand
usuall}' well away from cell, seldom touching it, rarely a dot iu apex of cell.
Underside : red snbmarginal spots of hindwing minute or absent.
? . Forewing : cell-patch anteriorly broader than in ? P. e. eun/lco/?. — —
Hindwing : discal band narrower ; red submarginal spots of underside smaller.
Scent-organ present.
Bab. Cauca valley, and Rio Dagua, West Colombia.
In the Tring Museum 14 c?c?, 4 ? ¥, from: Canca Valley; Rio Dagua
(Rosenberg).
c. P. eunjleon euryleon Hew. (1855) (PI. V. fig. 23).
Papilio euryleon Hewitson, I.e. (New Granada) ; Gray, List Lep. Ins. Brit. Mus. i. Pap. p. 70. n. :i83
(1856) ; Felder, Wien. E,it. M>m. v, p. 73. n. 4 (1860) (? descr.) ; id., Verh. Zool. But. Ges.
Wicu xiv. p. -Idil n. 146 (1864) (Bogota) ; id., Peixe Novnra, Lep. p. 44. n. 33. t. 6. fig. 4. $ (1865)
(Bogota) ; Kirby, Cat. Dinni. Lrp. p. 524. n. 44 (1871) (New Granada) ; Obertb., Et. iVEnl. iv.
p. 82. n. 269 (1880) (Carare, and from Bogota to Buenaventura, Colombia) ; Stand., Exot.
Tagf. i. p. 19 (1884) (Colombia).
S . Patch of forewing variable in extent, twice the size in some specimens as
in others, mostly reaching from M' to inner margin, but the streak behind SM^
often missing, there being on the other hand often a streak in front of M'.
The band of the hindwing consists usually of four spots standing closely around
apex of cell, and two more or less distinct spots before abdominal margin, there
being also a spot or some red scales in the ajjex of the cell in about half the
specimens.
On the underside the forewing bears occasionally a vestigial patch M' — M-.
The discal spots of the hindwing are pale pink, spots R^ — M^ are always present, spot
R^ — R^ is often vestigial, spot M- — SM' standing proximally of red anal submarginal
spot either present or absent ; anal spot with or without white proximal border.
?. Cell-patch of forewing more or less narrowed anteriorly ; band of hindwing
entering cell, red or pinkish buff.
Scent-organ present, but variable in extent.
Hab. Colombia : Magdalena valley and Cordillera of Bogota.
The male common in Bogota collections.
In the Tring Museum 84 <?(?, 3 ? ?, from : " Bogota" ; El Lumbo, Cundina-
marca, July 1903 (Mathan) ; Muzo, November and December 1896.
d. P. euryleon haenschi subsp. nov. (Plate VIII. fig. 54).
S. I'pperside. — Forewing : patches purer grey than in P. e. euryleon, different
in shape and position, extending from W to SM-, there being often a few white
scales before M' and sometimes a streak behind SM", the patch M' — W i)rojccting
beyond the second patch. Hindwing : a small spot in apex of cell, seldom
absent, and three to five spots around iipex of cell, the last one being vestigial ; a
trace of a sixth spot at abdominal margin.
Underside, — Forewing : a large white patch M' — M-, both the upper and under
scales of the patch being white, or a portion of the upper scales brown, a greyish
( 666 )
patch M- — SM- corresponding to the patch of npjierside ; on tins greyish patch
the under scales are white, the upper layer being brown. Hiudwing : three
pinkish discal spots, the first and second touching cell ; often a trace of a fourth
spot behind M".
?. i'ppcrsifle. — Forewing : cell-patch not reaching across cell, being
abbreviated anteriorly ; patch W — M' followed by a smaller patch shaded with
black, and in one (of the three specimens seen) preceded by a small whitish spot.
Hindwing : a red band from R^ to abdominal margin, entering cell, proximally
whitish behind cell, more or less shaded with black towards abdominal margin, pink
on underside.
Scent-organ jiresent.
Ilab. Western Ecuador.
In tlie Tring Blnseum 4 c? (? from : Balsapamba (R. Haensch), type ; Paramba
(W. Rosenberg).
4 c? c? and 3 ? ? in coll. Charles Oberthiir from Balsapamba.
3 cJ(J in the British Museum from the valley of Chimborazo and Porvenir.
e. P. earyUon anatmus subsp. iiov. (PI. Vlll. fig. 53).
Pajniio eiiryleoii, Dognin, Lep. Loja p. 14 (1887); id., I.e. p. 37 (1891); Haensch, Deri. Ent.
Zeitschr. xlviii. p. 153 (1903) (Sa. Inez, R. Pastaza, 1-250 m.).
(?. Upperside. — Forewing : grey patch more speckled with black scales than
in 1'. e. euri/leon, on the whole larger, extending from inner margin beyond M-,
touching cell, there being often a streak in cell proximally of origin of M-, streak in
front of M- elongate-triangular, narrowing towards cell ; some specimens with
submarginal spots. Hindwing ; three red spots E^ — M' on disc, well separated
from cell, occasionally pinkish grey, the second the longest, first occasionally absent,
sometimes the second alone distinct.
Underside : forewing without a trace of white patches. Hindwing : discal
spots R2— M- usually grey, rarely pinkish, usually a little nearer cell than distal
margin, often vestigial; red submarginal spots R'— SM' distinct, last one (anal)
without white border, or the border very thin.
¥ not known.
Scent-organ absent.
Hab. East Ecuador.
In the Tring Museum So 6 S from: Archidona (W. Goodfellow) ; Zamora
(0. T. Baron), name-type ; Loja.
127. Papilio hipparchus Stand. (1884).
Papilio hippmrlms Staudinger, Exol. Tagf. i. p. 20. t. 13. S (1^84) (Cauoa valley).
Not known to us. The hiudwing is figured as having only one rod sjiot at the
base, but that may be an error of the colourist, the colouring of the figure not being
quite correct according to Staudinger, I.e.
S. Forewing black, with a row of grey submarginal spots. Hindwing with
pale pink distal band from R' to abdominal margin, the band absent (or vestigial ?)
on underside, except the last s])ot, submarginal and admarginal spots as in
P. eunjleon.
? not known.
Hah. Cauca valley, Colombia.
( 607 )
128. Papilio harmodius Donbl. (184G) (PL V. fig;, ifi. 19 ; PI. VIH. fig. 52).
Papilio harmodius Doubleday, Ann. ilaij. N.H. xviii. p. .374 (1840) (Bolivia) ; Gray, Cat. Lap. Im.
Brit. Mus. i. Pap. p. 511. n, 272. t. 7. fig. 2 (1852) (Bolivia).
Sexes differeiit ; female dimorphic, one form somewhat resembling the male,
the other form qnite different. Geographically variable.
cJ. Dots on head, palpns, collar and coxae bnffish. Forewing semitransparent
in apical half, the scales being reduced in size; a white patch of variable size before
inner margin, sometimes reduced to a narrow streak situated behind SM-, never
reaching forward as far as R^, but occasionally extending across M' ; rarely a vestige
of a band across cell.- Hindwing : a row of red spots on disc, variable in size and
number ; a row of white arched submarginal spots, which are vestigial or absent in
most specimens.
Underside much paler than upper, cell of both wings with black folds.
Forewing with a red costal spot, Hindwing with three red spots at base, the one
in cell being 'small and often vestigial or even absent ; discal spots more or less
pinkish white proximally.
? ,. Forewing with white band across cell continuous with two large discal
patches R^ — M' ; spots of hindwing white or yellowish, or red centred with white.
?2. Forewing with trace of the white markings of female; spots of hindwing
red.
Scent-organ : numerous greyish scales on SM" of hindwing, these scales much
smaller than the scales situated before and behind this vein.
Genitalia : S ■ Distal lobe of harpe large, rounded, proximal angle of ventral
dentate edge not produced. ? not dissected.
Early stages not known.
Hab. Colombia to Bolivia.
The coloration of the first kind of female agrees with that of the females of
P. erlaces, the Ecuador females of both insects being white-banded on the hindwing,
the females from Northern Peru yellowish-banded, and the females from Southern
Peru and Bolivia red-banded.
a. P. harmodius isus Oberth. (1880).
Papilio xeniadcs var. isus Oberlhiir, Et. d' Eiit. iv. p. 81. sub n. 2G8 (18811) (Colombia).
Papilio aristogiton Staudinger, Exot. Tagf. i. p. 19. t. 1.S (1884) (Cauca).
S ■ Forewing : white patch large, reaching from hinder margin to M- or beyond,
being large also on the underside. Hindwing : a band of six or seven red spots
which are paler in the centre than at the edges, being occasionally pinkish white
edged with red ; white submarginal biirs usually absent.
¥ not known.
Hab. Cauca valley, Colombia.
5 c?(J in coll. Oberthiir, and 1 (? in coll. Godtnan.
b. P. harmodius halex sabsp. nov. (PI. VIII. fig. 52).
Papilio xeniadcs, Maassen & Weym. (non Hewitson, 18G8, err. det.), in Stubel, Reiscn S. Amer.,
Lep. p. 24. n. 103 (1890) (Villavicencio).
S. Forewing as in the preceding. Hindwing above with five red spots
which are not paler in centre, the upj)er two spots vestigial or minute.
?. Upper side. — Forewing with white patch from W bej-ond M-, just entering
cell. Hindwing with five pinkish red spots ; tooth R' prominent.
43
( 668 )
llah. « Bogota," Colombia.
In the Triiig Museum b S i from " Bogota."
A female in coll. Adams from " Colombia."
c. r. Iiannodius xeniades Hew. (1868).
Papilh xeniades Hewitson, Tram. Enl. Soc. Lnml (.3). v. p iifil. n. 1 (1868) (Ecuador ; ^ & '■ ? " ;
this " ? " is (J) ; id., Exot. Bull. iv. Pap. t. 9. fig. 26 (1868) ; Kirby, Cat. Viiini. Lep. p. 524.
n. 45 (1871) (Ecuador) ; Oberth., El. d'Ent. iv. p. 81. n. L'llK (1880) (Ecuador) ; Kirby, Trans.
Ent. Soc. Loud. p. 353 (1881) (Rio Topo, Ecuador ; "hardly distinct from gai/i Luc. ") ; Staud.,
Exol. Tagf. i. p. 20 (1884).
Papilio harmodiiis, Kirby, I.e. p. 524. n. 44 (1871) {purlhn ; Ecuador) ; Hopff., StfJI. Ent. Zeit.
x]. p. 52. n. 16 (187;!) (/lartim ; Ecuador); Staud., Exot. Tag/, i. p. 19 (1884) (/«/■(»«; Ecuador);
Dogn., Lep. Loja p. 15 (1887) ; id., /.<.-. p. 37 (1891).
<?. White patch of forewing very variable in size, often reduced to a streak
situated at inner margin, in other specimens the patch being extended beyond M-,
in most individuals e.xcised in front or obliquely truncate, its distal edge extending
beyond M-, while proximally the patch docs not reach M- ; on underside the patch
always small. Hindwing : three to five red spots, variable in size, occasionally
centred with pinkish white, being on underside pinkish white edged with red
distally ; white submarginal bars more often absent than present ; white marginal
spots very variable in size ; tooth R' in some sjiecimens more j)rominent than
in others.
? . Dichromatic :
a}. ?-f. nndrona nov. (PI. V. fig. 19). Forewing with some grey scales on
disc between lower angle of cell and M- and in cell. Hindwing : a row of five red
spots, spot W — M' the longest, almost three times as long as broad, ujipermost spot
small ; the spots pink-white on underside, slightly edged with red distally. One
specimen in the Tring Museum from Zamora (0. T. Baron).
d^. ?-f. Virginia Kirby (ISSl).
Papilio viryima Kirby, Tran.^. Ent. Sue. Loud. p. 352 (1881) (Rio Copataza, affluent of Rio Pastaza ;
"allied to P. larydes" '.) ; Grose-Smith & Kirby, Phop. E.rvl. ii. Pap. t. 16. fig. 3. 4 (18;i7)
(fig. of type). Resembling the female of /'. laci/dex. In coll. Oberthiir from Ambato ; in
coll. Urose-Smith from the Rio Capataza ; in Mus. Tring from the neighbourhood of Zamora.
Forewing : a large patch across apex of cell, two large patches R- — M' and a
streak behind M' white. Hindwing : a complete band of white patches, some of
them slightly edged with red both above and below.
Mab. of P. harm, xeniades : West coast of Colombia (Rio Dagua), and
Ecuador ; apparently common in Eastern Ecuador.
The females from West Ecuador and West Colombia may possibly be different
from the East Ecuadorian females. The long series of males in coll. Oberthiir from
Honda, Pacific slope of Colombia, agree on the whole with the specimens from
Eastern Ecuador, but the range of variation is somewhat different. One of the
Honda individuals has a vestigial band across the cell.
In the Tring Museum 1:20 <J<^, 3 ? ? , from : Zamora (0. T. Baron) ; Ambato ;
Santa Inez (11. Haensch) ; Archidona (W. Goodfellow).
d. P. Iiannodius imaus subsp. nov. (PI. V. fig. 16).
Papilio harmodina, Hopffer, 6'M(. Ent. Zeit. xl. p. 52. n. 16 (1879) (partim ; Peru); Staud., £i;o<.
Tarjf. i. p. 19 (1884) {jiartim ; Peru).
Papilio xeniades var. harnwdius, Oberthiir, Et. d' Enl. iv. p. 81. sub n. 268 (1880) (Peru).
S, Like the following; white patch of underside of forewing often reduced, in
most specimens a white streak behind SM-.
( 669 )
?. Forewing similar to that of xeniades ?-f. Virginia, but the cell-patcK
costally more or less reduced, aud the ii])per discal pateh smaller than iu ?-t'.
Virginia. Band of hindwing yellowish bnff on upperside, almost white on
underside, spots R^ — M^ long, spot R' — W- being nearly four times as long as
broad, extending close to cell, last (double) spot and one or two of the others
somewhat edged with red both on upper and under surface.
Ilab. North-East aud East Central Pern ; type : ? from Cnshi.
In the Tring Museum 27 JcJ, 3 ? ?, from : Fozuzo (W. Hoffmanns); Cushi,
Huanuco, 1800 m. (W. Hoffmanns); R. Chuchuras, affl. of R. Palcazu, 320 m.
(Hoffmanns) ; Huayabamba R., S.E. of Chachapoyas, 3500 ft. (0. T. Baron).
e. P. harmodius harmodius Doubl. (1840).
Pap'dio harnwdtiis Doubleday, I.e. (184<j) (Bolivii) ; Doubl., Westw. & Hew., Gen. Diurn. Lep.
i. p. ID. n. 2l'3 (184ti) (Bolivia) ; Doubl., Lhl Lep. Ins. lint, ^fl,s. i. App. p. 3 (1848) (Bolivia) ;
Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 59. n. 2T2. t. 7. fig. 2. fj (1852) (Bolivia) ; id.,
List Lep. Ins. Brit. Mus. i. Pap. p. 71. n. 288 (18o(3) (Bolivia) ; Felder, Verh. Zoul. Bot.
Ges. Wieii xiv. p. 299. n. 145 (1864) (Bolivia; "Ecuador" loci error, teste specim. in coll.
Felder) ; Kirby, Cat. Diurn. Lep. p. 524. n. 43 (1871) {partim ; Bolivia) ; Hopff., Stett. Eul.
Zeit. xl. p. 52. n. 16 (1870) {partim; Bolivia); Staud., A'xo(. Tag/, i. p. 19 (ISSi) { parti m ;
Bolivia) ; Haase, Untersuch. Mimicry i. p. 86 (1893) ; Weeks, Illustr. Diurn. Lep. p. 20 (190."))
(Chulumani).
c?. Forewing always with a large patch M' — SM-, consisting of two large spots,
both above and below, there being in most specimens a small white streak behind
SM-, and occasionally another streak in front of M' ; the white spots often larger
on underside than on upper. Hiudwiug with five red spots, the second being the
smallest, the third and fourth occasionally i)ale in centre, all the spots more or
less extended pinkish white on the underside of the wing ; often a series of white
submarginal bars present, the bars being usually more distinct above than below.
¥ . Forewing : white cell-patch not reaching across cell ; white discal area
consisting of three patches R- — M'-, the upper patch small, the third as long as the
middle one. Spots of hindwing larger than in male, red, sj)ots R^ — M' being
whitish in centre ; marginal tooth R' obtuse, not longer than tooth R^.
Hal). Bolivia aud South-East Pern ; the only female seen being in coll. Charles
Oberthiir, from Cochabamba.
A common insect.
In the Tring Museum 66 (?(? from various places, from Chanchamayo to the
Mapiri River.
129. Papilio trapeza spec. nov. (PI. V. fig. 15),
c?. Body as in 1'. harmodius xeniades, but the hairs of the frons shorter.
Wings, upperside. — Forewing proportionally narrower than in P. harmodius,
the hinder margin being shorter; a creamy, faintly greenish, patch, from hinder
margin towards M'-, which it does not reach, the second partition of this patch
broader and longer than in P. harmodius xeniades, reaching pro.ximally to vein
SM.'. Hindwing more triangular than in P. harmodius, more acutely dentate,
vein R- somewhat shorter tlian cell, while in P. harmodius this vein is as long as,
or longer than, cell ; white fringe-spots thinner ; three red spots from R^ to
abdominal margin, the first shaded over with black, the second elliptical, in one of
the specimens examined also shaded with black, the third spot much the largest,
being the same length as the second, but twice the width.
( fi70 )
VnJerside. — Forewing : jiatcb creamy wliitc, smaller than above, but the streak
along hinder margin broader and longer than in any form of P. harmodius, the
tipper patch not reaching beyond the fold (SM'). Hiudwing with fonr red spots
on disc, there being here an additional spot R- — R^, which is not present on the
npperside ; first and fourth spots small, second the largest, about twice as long as
broad, white jjroximally or in centre, or white with red distal border ; third sj)ot
one-third shorter than second, with trace of white proximally or in centre, or
similarly coloured as second spot ; three white curved snbraarginal bars R- — M-,
which are not present above.
Scent-organ : vein SM- densely covered with greyish brown scales as in
P. harmodius, but these scales rather broader than in that species.
Genitalia as in P. harmodius, the central process of the har))e being a little
broader than in most specimens of P. harmodius which we have examined.
Hab. East Ecuador.
In the Tring Museum 2 i i from : Rio Napo (R. Haensch), fi/pe ; Rio Curarai
(Simson).
In coll. F. D. Godman 'i S 3 from: Santa Inez and Aguamo. In coll.
Oberthiir from Saraya(,'« (Buckley).
This insect cannot easily be confounded with P. harmodius xeniades from the
same district, the sha])e and markings of the hindwing and the different development
of the patch of the forewing distinguishing P. trapcza at a glance.
In coll. H. Grose-Smith there is a male from "Ecuador" which is doubtless
an individual of the present si)ecies, though it differs from the specimens described
above in several points, the most remarkable difference being the total absence of a
patch from the upper and underside of the forewing. The hindwing bears on the
npperside three distinct red spots besides the vestige of a fourth, there being on the
underside four distinct spots and a vestigial fifth. The white submarginal spots of
the hindwing described above are absent. This specimen may be a geographical
form or an individual aberration of P. trapeza.
130. Papilio xynias Hew. (1875).
Papillo xyttias Hewitson, Ent. .\fo. Mug. xii. p. 153 (1875) (Bolivia) ; id., _B.ro/. Buli. v. Pap. t. 15.
fig. 48. (J (1877) ; Kirby, Cat. Diurii. Lep. p. 8U (1877) (Bolivia) ; Haa?e, Unta-sueh. Mimicry i.
p. 80 (1893) C'N. Granada" error loci) ; id., I.e. ii. p. 69. t. 9, fig. Gfi (1893).
cJ. Allied to P. Imrmodius ; hairs of frons as short as in P. trapeza. -
Wings, npperside : scaling of forewing non-dentate or feebly dentate ; a pale green
patch from M- to inner margin, variable in length, apparently never extending
beyond M-, always touching hindmargin, the scales entire, while in P. harmodius
and P. trapeza the scales of the patch are dentate. Hindwing : acutely dentate,
tooth R^ developed to a more or less distinct tail ; three red spots R'"' — SM'-, the last
large, often reaching halfway down to base, the first and second elongate, occasion-
ally reduced, rarely absent ; scales of these spots entire or a few of them slightly
dentate ; three to five white admarginal bars.
Underside much paler than upper ; cell-fold and red basal spots as in P.
harmodius. Forewing : patch of forewing jmler and rather smaller than above,
but very distinct. Hindwing: four red spots, the second and third longer than
broad, more or less pinkish white proximally, the last one double, about half the
( 671 )
length of the third, which is longer than the second; white submarginal bars rather
larger than above ; cell a little longer than vein R^.
? not known.
Scent-organ : no modified scales on SM- of hindwiug, the scales upon this vein
being the same size and colour as those in front of and behind the vein, P. xynias
differing in this respect remarkably from all its allies.
Eiirly stages not known.
Hub. Bolivia and Peru, eastern slopes.
In the Tring Museum 15 (?c? from: Pozuzo, Huannco, 800—1000 m. (W.
Hoffmanns) ; Chanchamayo (Schnnke) ; La Union, R. Huacamayo, Carabaya,
2000 ft., December 1904, wet season (G. Ockenden) ; Mapiri, Bolivia.
In coll. Oberthiir from Hillapani, Tarapoto and Chanchamayo.
131. Papilio ariarathes Esper (1788) (PI. V. fig. IT, VIII. fig. 57).
Painlio Eqiics Trnjanus ariarathes Esper, Ausl. Schmelt. p. 57. n. 24. t. 14. fig. 2. ? (1788)(S. Amer.;
" var." fig. 3 alia species).
Pajjilio ariarathes, Bates, Trans. Ent. Soe. Land. (2). v. p. 336 (1861) (variability) ; id., .Tourn.
Entom. i. p. 224. n. 8 (1862) (geogr. variability ; cyanon, gaiji, eragoras are local vars.) ; Kirby,
Cat. Diiau. Ly,. p. 523. n. 35 (1871).
c??. Buffish grey dots on occiput, pronotum, palpus and coxae. Spination of
legs rather heavy. Wings, beneath, with two red spots at base of forewing, one in
cell and the other before it, and three basal spots on hindwing, the second situated
in cell being often vestigial or absent.
Very variable in both se.xes ; forewing of male with a whitish or buffish band
or patch on upperside e.\ tending from inner margin forward, being seldom vestigial,
Hindwing of male with a discal series of red patches in most specimens,
sometimes only one or two distinct jjatches present on upperside, situated near
abdominal margin ; cell rather narrower in middle than in the allied species.
In tlie female the forewing has usually one or more white patches on disc and
often a small patch in cell, but is sometimes all black. A red discal band on
hindwing, variable in width, often entering cell.
Scent-organ : basal third of vein SM^ densely covered with small greyish
scales.
Genitalia : c?. Apical lobe of harpe short ; ventral lobe of dorso-ventral ridge
slightly curved upwards in a view from the base of the clasper, obliquely rounded ;
central process spatulate, dentate, usually rounded at ape.x, but sometimes obliquely
truncate.
Early stages not known.
Hal). Colombia to Bolivia, and eastwards to the Guianas,Par;C, and the Brazilian
province of Goyaz.
AVith the excei)tion of the form from Goyaz, the subspecies do not appear to be
sharjily defined. In the case of species individually so variable as /*. ariarathes it
requires a large material from many districts to obtain a correct view of the
geographical variation. The material 'of P. ariarathes which we have seen in
various collections is not quite sufficient for this purpose. Therefore we are not
sure that our division of P. ariarathes into six subspecies is quite true to nature.
The species varies in different localities according to the models mimicked, the
females from different districts being generally more obviously different than the
males.
( 672 )
a. P. arkirathcii ariarathex Esjier (1788).
Pnplllo Eqncs TrojnnKS nriamthcs'EspeT, l.r. J (ITSS).
rajiiUo ilii.i, Godart («o« Fabr., 17'J3, err. det.), Eiic. Melh. ix. p. 33. n. 21 (l«l',l) (syn. excl. ; ^,
"America"); Boisd., Kpfc. Ghi. Lep. i. p. 280. n. 104 (183()) (-'America"); Oberth., Et.
(VEiil. iv. p. 81. p. 264 (1880) (partim ; Cayenne).
Papilio ariaratlies, Boisduval, Spec. Gen. Lep. i. p. 287. n. 14 (1836) (Surinam ; ? ) ; Doubl., Westw.
& Hew., Gci. Diimi. Lep. i. p. 18. n. 207 (1846) (Guiana) ; Gray, Gil. Lep. 7».s-. DHl. Mux.
i. Pajj. p. Gl. n. 275 (1852) (descr. of (J ; '' N. Granada '■"' error) ; Felder, Verh. Zool. Bol.
Ge.i. Wien xiv. p. 298. n. 136 (18G4) (^parlin,) ; Kirby, Gil. Dium. Lep. p. 523. n. 35 (1871).
Papilio acesles Boisduval, I.e. p. 288. sub n. 14 (1836) (nom. maxime superfluum).
<J. Forewing : a ratlier narrow patch from inner margin to M", nearl}- as large
lelow as above. Hindwing : fonr to six red sjiots, separated from one another,
distant from cell.
? . Forewing with white patches, or these patches vestigial. Five or six red
spots on hindwing, spots R-— M- being long, touching cell or nearlj-.
a'. ?-f. ai-iarathes Esp., I.e. — Forewing with one to three white patclics on disc
and often a narrow patch posteriori}* in cell.
h'. ?-f. eumelea uov. — White markings of forewing vestigial (name-type from
Surinam).
Hub. French and Dutch Gniana.
In the Tring Museum 4 ¥ 5? from Surinam.
b P. ariarathes menes sabsp. nov. (Plate VIII. tig. 57).
Pupilio ariarathes, Erichson, in Schomb., F. F. Brit. Guiana p. 593 (1848) (descr. of ^) ; Bates,
Trans. Ent. Sue. Land. (2). v. p. 336 (1861) (partim) ; Brit. Guiana).
(J. Forewing, above, with band from inner margin to M' or M-, tiie jiosterior
s];ot often much reduced, patch i\l- — SIM- sometimes alone distinct, 4 to G mm.
wide ; this patch alwaj's present on underside, where it is a little smaller than
above, the spot behind SM- and the one before M- also present below in the
si)ecimens which have them clearly marked on the npperside. Hindwing rather
strongly dentate, tooth W prominent; three or four red spots on npperside, standing
about halfway between cell and distal margin ; five, seldom four, spots on underside,
spot R^ — M' being pale proximally.
?. Forewing with a large patch W — M', a smaller one M' — M'-, often reduced
to a streak, a vestigial spot in cell and occasionally another vestigial one before
R^ {type). Hindwing : five or six red spots, all well separate from cell, spots
II'— M- longer than the others.
llah. British Guiana ; type : ? .
In the Tring Museum 5 (?cJ, o ? ?, from: C'hristianburg, R. Demerara ;
liartica, 26. February 19U4 (R. Haensch) ; Upper Ileal Berbice R.
c. P. ariarathes evagoras Gray (1852).
Pupilio iltis, Doubleday (mm Fabr., 1793, err. det.), List Lep. Ins. Brit. .l/«s. i. Ajip. p. 3 (1848)
(partim ; Venezuela) ; Oberth., El. d'Eut. iv. p. 81. n. 264 (1880) (partini ; Paril).
P.ijilio evarjoras Gray, Git. Lep. Tint. Bril. Miis. i. Pap. p. 61. n. 276. t. 9. fig. 3. cj, 4. ? (1852)
(Venezuela) ; id.. List Lep. Ins. Brit. Mus. i. /'u/a p. 72. n. 292 (1856) ; Feld., Verh. Zoul. Bol.
Ges. Wien xiv. p. 298, n. 132 (1864) (parlim ; Venezuela) ; Oberth., El. d'Enl. iv. p. 81. n. 265
(1880) (Caracas); Stand., E.ti,l. Taijf.l p. 13 (1884) (Venezuela); Haase, Unlersiuh. .Uimicri/
i. p. 97 (1893) (" Xew (iranada" errore).
Papilio ariarathes local var. evagoras, Bates, Trans. Ent. Sm: Land. (2). v. p. 336 (H61) { parti in ;
Venezuela).
P.ipilio ariarathes var. P. evagoras, Kirby, Gil. Dium. Lep. p. 523. sub n. 35 (1871).
( 673 )
(?. Forewing : a narrow band, extending from inner margin to M' or beyond,
situate halfway between cell and distal margin at W and M' or nearer the cell ; the
band represented on u/idersi(/e by two or three clearly marked spots. Hindwing :
four or five red spots, spots R- — M- close to cell as a rale, spot R' — R- less distant
from cell than in the other forms, especially on underside, the band therefore
appearing more curved ; spots R- — M- pinkish white beneath with red distal border,
sometimes partly bnffish white on upperside.
?. Three white or bnffish discal patches on forewing and a rather large ccll-
jjatch. Hindwing : six large patches around cell (sometimes huffish) and apical
half or two-fifths of cell red; abdominal margin at least partly red.
Ilah. Venezuela, from Caracas to the Orinoco.
In the Tring Museum 5 S S , 1 ?, from: "Venezuela" (Moritz) ; Porto
Cabello ; Caicara, Orinoco, 1898 (Cherrie) ; Ciudad Bolivar, 5. September 1898
(S. M. Klages).
d. P. ariarathes metagenes subsp. nov.
Papilla ariarathes, Wallace, Trans. Ent. Soc. Loud. (2). ii. p. 256 (1854) (Pard) ; Bates, I.e. (18'j1)
(partim ; FarA) ; Feld., Verh. Zonl. Bot. Ges. Wieit xiv. p. 298. n. 136 (1864) {purtim ; Pari).
Papilio cyamon, Haase {iioit Gray, 1852, err. det.), Untcrsuch. Mimienj i. p. 87. ^ . t. 10. fig. 71. 5
(1893) (Pard).
cJ. Forewing : usually with a baud from hinder margin beyond R^, gradually
disappearing, vestigial on underside. Hindwing : four or five red spots, separate
from cell, ujiper one or two small, spots R^ — M- elongate, longer on upper than
underside ; tooth R' prominent.
? . Forewing : a white band along hinder side of cell consisting of two large
patches R^ — M^, a smaller patch R- — R', and a fourth (much variable in size)
behind M", the white scaling sometimes nearly reaching inner margin of wing ; cell
with or without white streak. Ilindwiug : si.\ red spots, R- — M- touching cell,
at least on upperside.
Hab. Pard.
The female resembles the females of some Aristolochia->Swallowtails(P. anckiscs
thelios, P. uglaope).
Name-ty23e iu coll. Obertliiir.
e. P. ariarathes gayi Lucas (18.52).
Papilio rjaiji Lucas, Rrr. Zool. p. 193 (1852) (Ciizco, Peru) ; Gray, Cat. Lep. Lis. Brit. Mas. i. P<ij>.
p. 60. n. 273 (1852) ; ifl., List Lep. Ins. Brit. .Miis. i. Pap. p. 71 n. 289 (1850) ; Fel.l., Verb. Zool.
Bot. Ges. Wien xiv. p. 298. n. 134 (1864) (Cazco ; Ega) ; Hopff., Stett. Eiit. Zeit. xl. p. 51. n. 1 1
(1879) (Peru) ; Staud., E.cot. Tayf. i. p. 13 (1884).
Papilio ajamoii Gray, Cut. Lep. Itis. Brit. Jfu.i. i. Pap. p. 60. n. 274. t. 7. fig. 1. d, i. 11. fig. 3. ?
(1852) (Ega, cj ; ? hab. ? ; var. g " Brazil ") ; id.. List Lep. Ins. Brit. Mas. i. Po/;. p. 71. n. 290
(1856) (Ega ; Villa Nova ; Rio Negro) ; Wall., Trans. Ent. Soc. Land. (2). ii. p. 256 (1854)
(Upper Amazons ; forest) ; Felder, l.r. p. 298. n. 135 (1864) (Ega ; Villa Nova ; Rio Negro) ;
HopfP., I.e. p. 51. n. 15 (1879) (Brazil, Peru); Staud., I.e. p. 13 (1884)(distiDct species ; Amazons).
Papilio eragoras Gray, Cat. Lep. Ins. Brit. Mas. i. Pap. p. 62. sub n. 277 (1852) ( partim ; J , Burra) ;
Wall., Trans. Ent. Soc. Loml. (2). ii. p. 256 (1854) (Upper Amazons ; forest) ; Felder, Verb.
Zoul. Bot. Ges. Wien xiv. p. 298. n. 132 (1864) {partim ; Ega ; R. Negro).
Papilio ariarathes local var. cijamon, Bate.'), Trans. Ent. Soc. Land. (2). v. p. 336 (1861) (Upper
Amazons).
Papilio ariarathes local var. yai/i, id.. I.e. (1861) (Ega).
Papilio ariarathes local var. eoar/oras, id., I.e. p. .337 (1861) (Ega).
Papilio ari.itarjoras Felder, I.e. p. 298. d. 133 (1864) (Bogota) ; id., Ileise Norara, Lep. p. 41. n. 30. t. i .
fig. e. J' ab., f. ? (1865) (Bogota).
( 674 )
Papilio ariarathes var. P. cyamon, Kirby, Cat. Ditirn. Lep. p. 523. sub n. 35 (1871).
Pai>Uio ariarathes var. P. gayi, id., I.e.
Pajiilio ariarathes var. P. aristagoras, id., I.e.
Papilio Has, Oberthur {iwn Fabr., 17113, err. det.), Et. d'Eiit. iv. p. 81. n. 264 (1880) {parlim ;
Obydos ; variability of " ihis ").
Papilio charnba Kirby, Trans. Eiit. Soe. Loud. p. 352 (1881) (R. Pastazza).
Papilio arianus Staudinger, Exot. Tagf. i. p. 12 (1884) (Amazons : R. Maut's to E. Huallaga ;
Esper's fig. of aWa)-a(/iM erroneously consid. (^); Michael, Iris vii. p. 213 (1804) (Sao Paulo
de 01iven{,a).
Papilio ariarathes, Staudinger, I.e. t. 8. (J (1884).
(J?. There are three principal individual form.s, connected b}' intorgradations.
Most specimens of the male have no sharplj- marked white spots on the underside
of the forewing or only small spots.
a'. P. a. gayi f. anargus nov. ; P. a. var. ojamon ?, Bates {non Gray, err.
det.), I.e. — Forewing without white band or patches ; in male a few grey scales
indicating the band ; hindwing of female with a vestige of a red dot in apex of cell.
Middle and Upper Amazons ; name-type: ? , from Iqnitos.
U. P. a. gayi f. cyamon Gray, I.e. ; P. eharoha Kirby, I.e. — S. Similar to male
of P. a. evagoras ; forewing with a band which is on the whole a little more distal
than in evagoras. Hindwing : four or five red spots, middle ones either reaching
cell or separate from it. ?. With narrow vestigial band on forewing; hindwing
with red cell-spot ; sometimes nearly all the sjiots of the hindwing creamy.
Middle and Upper Amazons ; Ecuador ; Pern ; Bolivia.
e'. P. a. gayi f. gayi Lucas I.e. ; P. aristagoras Feld., I.e. ; P. arianus Stand.,
I.e. ; P. eyamon Gray, l.c ., ? non S. — $. Forewing: a large huffish patch before
inner margin ; seldom white, varying much in size, sometimes not reaching M-,
in other specimens externally produced forwards, this projection corresponding to
the band of the next form (cJ-f. cyamon). Hindwing : two to five red spots
(occasionally creamy), sometimes only the last spot distinct, spots variable in size,
sometimes ajiproaching cell. ?. Forewing : one or two large white patches on
disc, usually a small third spot and often a cell-spot. Hindwing mostly with
cell-spot. Middle and Upper Amazons ; Colombia; Pern; Bolivia.
Hah. of P. a. gayi : Middle Amazons to Colombia and southward to Bolivia.
In the Tring Museum 48 c?c?, 5 ? ?,from: Bogota; Archidona and Coca,
E.Ecuador (W. Goodfellow) ; Aguamo, R. Napo (R. Haensch) ; Mau;ios ; Juhuty,
April 1905 (Mathan) ; Yurimaguas ; Thomar ; Iqnitos : Sao Paulo de Oliveufa ;
R. Cachyaco, affl. of R. Huallaga (Stuart) ; Chanchamayo (Schunke) ; La Merced
(Watkius) ; Reyes, August 1895 (Stuart) ; Prov. Sara, S. Cruz de la Sierra,
February 1904 (J. Steinbach).
/. P. ariarathes leiwtra subsp. nov. (PL V. fig. 17).
S. Forewing : a pure white patch from inner margin beyond M', of almost
even width, 6 mm. broad behind M', hardly narrower below than above, the spot
behind SM- sometimes absent. Hindwing : a band of six long red spots, the
last double, spots R- — M-' touching cell, i-ome grey and red scales between C and
SC- representing a seventh spot ; in one of the sjiecimeus some grey scales in apex
of cell on underside of hindwing.
Ilah. Goyaz, Brazil.
4 c?cJ in coll. Charles Oberthur; also iu the Hope collection, Oxford, collected
by Burchell.
( 675 )
132. Papilio ilus Fabr. (1793) (PL VIII. fig. 50).
PapiUo Eques Tros Hks Fabricius, Eiit. Syst. iii. 1. p. 17. n. 51 (17y3) (Amer., "Jon. fig. pict. i.
tab. 29").
Papilio ilus, Gray, Cat. Lep. Ins. Brit. Miis. i. Pap. p. 59. n. 271 (1852) {»yaon. partini) ; id., List
Lep. Ins. Brit. Mus. i. Paj). p. 71. n. 287 (1856) (synon. partini) ; Bates, Trans. Enl. Soc. Land.
(2). V. p. 335 (18G1) ; id., Jouni. Entom. i. p. 224. sub n. 8 (1802) (prob. = hostilius Feld.) ;
Feld., Verh. Zonl. Bot. Ges. Wieii xiv. p. 299. n. 143(1864); Butl., Cat. Diurn. Lep. descr. Fabr.
p. 238. n. IC (1869) ; Kifby, Cat. Diurn. Lep. p. 623. n. 41 (1871) ; Godm. & Salv., Trans. Enl.
Soc. Land. p. 126. n. 240(1880) (Sta. Marta) ; iid., Biol. Centr. Amer., Rhup. ii. p. 208. n. 30
(1890) (cj ¥ , Panama ; Colombia ; Venezuela ; = hostilius).
Papilio hostiliuH Felder, Wicn. Enl. Munatschr. v. p. 73. n. 5 (1861) (Prov. M^rida) ; id., Verh. Zool.
Bot. Ges. Wien. xiv. p. 299. n. 144 (1864) (Caracas) ; id., Peise Nornra, Lep. p. 43. n. 32. t. 9.
fig. a (1865) (Venezuela, coll. Kaden ; — type now in coll. Godman) ; Kirby, Cat. Diurn. Lep.
p. 523. n. 42 (1871).
Paj)iHo gwieo Staudinger, Verh. Zool. Bot. Ges. Wien xxv. p. 91. n. 1. (1876) (Chiriqui) ; id., Exot.
Tagf. i. p. 13 (1884) (Cbiriqui) ; Godm. & Salv., Biol. Centr. Amer., Phop. ii. p. 209. n. 31.
t. 67. fig. 1. cJ (1890) (Chiriqui).
Jndging from a small series of specimens of this apparently rare insect — rare
in collections at least — we come to the conclusion that the differences between the
types of ilus (Jones's figure), hostilius and guaco are individual, not specific or
geographical.
3 ? . Close to P. bfcinchus, of which it is perhaps a southern form, the patches
of the forewiug being, however, in a different position. Forewing with or without
bufiish white spot in cell ; two or three patches of the same colour behind cell,
patch M- — SM- being the largest; distal margin with small white spots, except at
apex. Hindwing : a row of red spots from R- or U' to abdominal edge, the spots
variable in size, separate from cell or close to it, larger in female than in male.
Underside : forewing black-brown, patch M- — SM- vestigial or distinct, but
always smaller than above. Hindwing with four basal spots as in P. belesis and
braiichus ; discal spots much paler than above, being more or less whitish pink, at
least proximally.
Early stages not known.
Hab. Northern Venezuela; Colombia: Sta. Marta and Valdivia ; Panama:
Lion Hill and Chiriqui.
In the British Museum, coll. Godman, and coll. Grose-Smith.
133. Papilio branchus Uoubl. (184(3).
Papilio branchus Doubleday, Ann. May. N. II. xviii. p. 373 (1846) (Honduras).
J ? . Some spots on occiput and collar red ; frons usually black, often with
two red vittae ; a white dot laterally on forecoxa, a jiink or red dot each on meso-
and metasternite and at base of first abdominal segment. Va\\A usually with
white dot or dots. Wings black, opaque, paler below than above. Forewing
with white patch situated on disc and in cell, separated into two, three, or four
spots by the black veins, or this patch absent ; fringe white between veins, but
these spots often very small, usually absent from apex of wing. Hindwing : a
broad band of six, seldom seven red spots; spots II- — M- long, situated near cell.
Underside : white patch of forewing usually a little larger than above, no red
spot at base. Hindwing : four red spots at base, the posterior one continuous, with
a line situated on abdominal fold ; red discal spots much paler than above, pinkish
white, shading into red distally.
( 676 )
Genitalia: c?. Apical lobe ol haqte slightly triaugnlar, rounded off; dentate
ridge rather strongly elevate ventrally and heavily dentate.
Early stages not known.
There are two individual forms :
a'. F. hrancliKs f. biani-hux Uoiibl. ('Is4(i).
PapiUo braitchus Doubleday, I.e. (Honduras) ; id., Westw. & Hew., Gen. Dinrii. Lep. i. p. 19. n. 217
(1846) ; Doubl., Li>'l Lep. Inn. Biil. Miis. i. Appeml p. 3 (1848) ; Gray, C»l. Lrp. Ins. Brit.
Miis. i. Pap. p. 62. n. 277. t. 7. fig. 3. ? (1852) (Honduras ; Guatemala) ; id., List Lep. Ins.
Brit. Mus. i. Pap. p. 72. n. 293 (185f)) ; Weidem., Proc. Ent. tine. Pliilml. ii. p. 146 (1803) ;
Feld., Verh. Zool. Hot. Ges. Wieii xiv. p. 298. n. 137 (1864) (Honduras ; Guatemala; Mexico):
Boisd., Consid. Lep. Guatem. p. 7 (1870) (Honduras ; Mexico ; = he/ilutestiim errore) ; Kirby,
Cat. Ditirn. Lep. p. 5'.'3. n. 36 (1871) (Amer. centr. ; Mexico) ; Butl. & Druce, Proc. Zool. Sue.
Land. p. 364. n. 369 (1874) (Costa Rica) ; Obertb., Et. d Ent. iv. p. 81. n. 266 (1880)
(Honduras) ; Godm. & Salv., Biol. Centr. A mer., Rlioji. ii. p. 207. n. 26. t 67. fig. 7. genit. (18'.i0)
(Mexico : Cordova, Omealca, Atoyac, Oaxaca ; Guatemala ; Honduras ; Nicaragua ; Costa Rica).
c? ? . Forewing with white central patch. This patch variable in size. It
consists in the male of a large cell-spot, a large sj)ot W — M', and a minute spot
R- — W, there being occasionally also a spot M' — M' ; spot R- — R' rarely absent.
In the female, spot R^ — R^ much larger than in the male.
This form occnrs from Vera Crnz in Mexico to Costa Rica.
I)', r. I /•(Die //lis f. l/elepAantes Godm. & Salv.
Papilio belephantes Godman & Salv., /.<•. p. 208. n. 29. t. 67. fig. 0. J (1890) (Mexico: Atoyac;
Guatemala ; Honduras).
(?. Forewing without white patch.
This form is much rarer than the preceding one. It has apjiarcntly the same
distribution, but is so far known only from Southern Mexico, Guatemala, and
Honduras.
IlaL of F. bra)wl/us : Mexico to Costa Rica.
In the Tring Museum 26 (? cj and 2 ? ? of f. branchiis from : Cordoba, Vera Cruz,
2SU0 ft., July 1904 (A. Hall) : Songolica, June 1899 (W. Schaus) ; Mazatenauga,
W. Guatemala, lUOO ft., September 1904 (A. Hall; ; Guatemala, San Pedro Sula,
Honduras; Cartago and Guatil Piris, Costa Rica (Underwood). 1 c? of
f. belephantcs, no locality.
134. Papilio belesis Bates (1804).
Papilio helesis Gray, List Lep. In.i. Brit. Mus. i. Pap. p. 74. n. 300 (1856) (Mexico ; «om. nurf.) ;
Weidem., Proc.E/it. Soc. Philad. ii. p. 146 (1863) (Mexico ; no/n.nud.) ; Bates, i'/if. Mo. Mag.\.
p. 1. n. 1 (1864) (Guatemala).
(??. Clo^Q io P. bianchus. Forewing usually quite black, but sometimes with
one or two white spots on disc ; fringe-spots nearly always distinct, except at apex
of wing, rarely vestigial. Hindwing variable in length ; a row of six or seven red
spots, situated parallel to distal margin and nearer to the margin than to the cell ;
the middle s])0ts not so much larger than the others as in P. branch/is ; size of the
spots very variable.
Genitalia : S. Apical lobe of hurpe shorter and more evenly rounded than in
r. bi-anchtis, dentate ridge more narrowly elevate ventrally, dentition less heavy.
Early stages not known.
Two individual forms :
( 677 )
a'. P. belesis f. hephaestion Feld. (18G5).
Papilio hejihaestioii Felder, Verh. Zoal. Bot. Ges. Wicn xiv. p. 298. n. 139 (1864) (Mexico ; nom.
nud.) ; id., Efiae Novum, Lep. p. 42. n. 31. t. 6. fig. b (18(j.5) (Mexico— Mus. Tring) ; Kirby,
Cat. Dhirn. Lrp.-p. 523. n. 38 (1871) (Mexico) ; Obertb., Et. d'Ent. iv. p. 81. n. 267 (1880)
(Mexico); Godm. & Salv., Biul. Cent. Amer., Rhnp. ii. p. 208. n. 28 (1830) (Mexico; Guatemala;
Honduras).
Papilio branchus, Boisduval, Consul. Lep. Guatein. p. 7 (1870) (hejihaention = branchus ! false).
cJ. Forewing with a white spot R^ — R^ and sometimes a trace of a spot R' — M' ;
there are also some bufiish scales behind R^ and near the hinder angle iu our
specimens. ? not known.
This form is known to us from : Mexico, Guatemala, and Hondaras (San Pedro
Sula, in coll. Charles Oberthlir).
b'. P. belesis f. bele.vs Bates (1864).
Pajiilio belesis Bates, I.e. (Guatemala) ; Feld., Verb. Zool. Bot. Ges. Wieu xiv. p. 2118 n. 138 (1864)
(cit. falsa ; Guatemala) ; Kirby, Cat. Diurn. Lep. p. 523. n. 37 (1871) ; Godm. & Salv., l.f.
p. 207. n. 27. t. 67. fig. 3. 4. J", 5. genit. (1890) (Mexico : Atoyac ; Guatemala ; Nicaragua) ;
iid.. I.e. p. 729 (1901) (Saa Pedro Sula, Honduras).
c? ? . Forewing without white spot on disc and in cell. In one of four males from
Guerrero, Mexico, the red submargiual spots of the hindwing are minute, while iu
the only female we have from that province they are as large as in P. branchus, but
have a position as distant from the cell as in ordinary specimens of P. belesis.
We know this form from Mexico : Atoyac and Guerrero, Guatemala, Honduras,
and Nicaragua.
Hab. of P. belesis : Mexico to Nicaragua.
While in P. branchus the form with white patch on the forewing is the common
one, in P. belesis the white-spotted form is the rarer one.
In the Triug Museum 2 S S of f hephaestion from Orizaba and " Mexico."
12 cJd'aud 1 ¥ of f belesis from: Orizaba; Guerrero (0. T. Baron); Escuintla,
W. Guatemala, 1100 ft., August 1904 (A. Hall) ; Polochic valley ; San Pedro Sula,
Honduras.
135. Papilio thymbraeus Boisd. (1836).
Papilio Ihymhraeus Boisduval, Spec. Gen. Lep. i. p. 3ii2. n. 136 (1836) (Tlatlecope, Mexico) ; Schaus,
Papilio iii. p. 186 (188.S) (descr. of adult larva & pupa ; on Chirimoija).
c? ? . Some dots on head, a dot on palpus, on pro-, meso- and metasternum and
first abdominal segment, and some speckles on the other abdominal segments, on the
sternites as well as the tergitcs, huffish, often somewhat pinkish.
Wings, npperside, olivaceous black, distinctly metallic blue or green. Fore-
wing without markings, fringe white, often brown at apex of wing. Hindwing
with a slender nou-spatulate tail, and one or two rows of red s])ots.
Underside iKile olive, somewhat metallic; no black cell-streaks. Forewing
with a red costal basal spot. Hindwing with four basal spots as in P. branchus and
allies, the jiosterior one contiiuious with a line situated on abdominal fold ; red
submarginal spots edged with bullish white proximally ; a row of huffish white
curved admarginal bars.
Genitalia: S. Ajiical lobe of harpe finely dentate ; dorso-ventral ridge highest
dorsally, becoming gradually lower ventraily, with feeble dentition ; central process
Jiracticaliy non-dentate; ventral process broad, obtuse.
( 678 )
Mature larva and pnpa described by Schaus, I.e.
ffab. Mexico to Honduras.
Two subspecies.
a. P. t!ii/mbraeus th>/mbraeus Boisd. (1836).
ritpilio tliijinhmeus Boisduval, I.e. (TIatlecope) ; Doubl , Westw. & Hew., Get\^ Diurn. Lip. i. p. 19.
n. 'i-iS. t. 4. fig. 3 (1846) ; Doub)., Lhl Lep. Iiis. Brit. Mus. i. App. p. 3 (1848) (Mexico) ; Gray,
Cat. Lep. Lis. Brit. .l/«s. i. Pap. p. 65. n. 289 (185'j) (Mexico) ; id., List Lep. Lis. Brit. J/w.s. i.
Pap. p. T5. n. 306 (1856) (Guatemala; Mexico) ; MeniT-tr., Eniim. Corp. Anim. Mas. Petrnp.,
Lip. i. Stippl. p. 68. n. 1129 (1857) (Mexico) ; Ueak., Proc. Enl. Soc. Philad. ii. p. 140. n. 11
(1863) (Chiapas) ; Weidem., ibid. p. 148 (18(;:i (Mexico : Centr. Amer.) ; Feld , Verli. Z,ml Bot.
Ge.i. ^Vie)l xiv. p. 299. n. 142 (1864) (Mexico; Guatem.ala) ; Kirby, Cat. Diurn. Lep. p. 523.
n. 40 (1871) (Mexico; Guatemala); Obertb., El. d'Eiil. iv. p. 80. n. 260 (1880) (Mexico) ; Schaus,
Papiliii iii. p. 186 (1883) (all the year ; open country) ; Stand., Exol. Tarif. i. p. 15. t. 9 (1884)
(Mexico; Guatemala); Godm. & S.alv., Binl. Ceiilr. Amer.. Rlmp. ii. p. 206, n. 24. t. 67. fig. 2.
genit. (1890) (Mexico: Vera Cruz, Oaxaca ; Brit. Honduras ; Guatemala); Haase, Uiitersurh.
Mimiiry i. p. 80. t. 9. fig. 64. ? (1893).
Papilio thymbraus (!), Guen^e, Mem. Sor. Phys. Jlist. Nat. Genive p. 379 (1872).
cJ 5 . Hindwiug with two jows of red spots.
Hub. Eiistern Mexico : Vera Cruz, southward to Honduras.
One of our males from Jalapa has a red dot on the underside of the left forewing
between IV and E-.
In the Tring Museum 28 J (?, 9 ? ?, from: Jalapa, February 1896 and July
1897, and Espinal, June 1896, Vera Cruz (W. Schaus) ; Orizaba, Vera Crnz, June
1904 (A. Hall) ; Palin, W. Guatemala, 2500 ft., Aug., Sept. 1904 (A. Hall) ;
Guatemala (Salviu).
b. P. tki/mbraeus aconophos Gray (1852).
Papilio aeoiwphos Gray, Cat. Lep. Ins. Brit. Mits. i. Pap. p. 65. n. 290. t. 9. fig. 1. ? (1852) (Pnebla,
Mexico) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 75. n. 307 (1856) (Puebla) ; Feld., Verh.
Zool. Bot. Ges. Wien xiv. p. 299. n. 141 (1804) (Mexico) ; Kirby, Cat. Diurn. Lep. p. 523. n. 39
(1871) (Mexico); Oberth., Et. d'Enl. W. p. 80. n. 261 (1880) (Me.icico) ; Wood, Ins. Abroad
p. 548. fig. 298 (1883) ; Stand., Emt. Tag/, i. p. 15 (1884) (Mexico) ; Godm. & Salv., Biol.
Centr. Amer., Rhop. ii. p. 200. n. 25 (1890) (Mexico: Puebla, Oaxaca); Haase, I.e. (1893);
Godm. & Salv., I.e. p. 729 (1901) (Guanajuato).
Papilio aconophas (!), Wcidemeyer, Proc. Ent. Soc. Philad. ii. p. 146 (1863) (Mexico).
c? ?. Hindwing with one row of red spots, the discal spots being absent.
Hub. Central and Western Mexico : Puebla, Guanajuato, Jalisco, Guerrero,
Oaxaca.
In the Tring Museum, 33 <? J, 5 ? ¥, from : Guadalajara, October 1896 (W.
Schaus): Guadalajara (Dr. Butler); Coantla, Morelos, 3800 ft., June 1904 (A.
Hall); Cuernavaca (Bilimet) ; Cuernavaca, end of August 1904 (Dr. Gadow) ;
Ayutla, Guerrero, 5. August 1904 (Dr. Gadow) ; Guerrero (0. T. Baron).
136. Papilio lysithous Hiibn. (1822?).
Hectorides lysitliaiis Hiibner, Samml. E.rot. !<r!,m. ii. t. 100 (1822?).
Pajtilio Imrrisiamis, Godart, Enc. Melh. ix. Supjil. p. 812. n. 138-9 (1824).
Pa2Jilio Claudius Boisduval, Spec. Gen. Lip. i. p. 311. n. 149 (1836) (Rio de Jan. ; harri.fiaims, laiiis
and Claudius perhaps one species).
Papilio lysithous, Burmeister, Deser. Rep. Argent, v. Li'p., .i(/((8 p. 9. n. 24 (1879) (Petropolis ;
N. Friburgo ; larva & pupa).
Papilio , Jones, Proc. Lit. Philos. Soc. Lirerpool xxxiv. t. 65. fig. 1 (1880) (larva, pupa).
Papilio lysilhout, id.. I.e. xxxvi. p. 44. n. 42 (1882) (larva, pupa).
( 679 )
We unite under this heading all the tailed specimens from I^razil belonging to
the present group of iVj(;i7/o, namel}' the forms described as h/sitlioiis, Iiarrisianus,
athous, sebast/d/iiis, rinik, pompoinm, and eupatorion, which have hitherto been
considered distinct species. Intergradations between the various forms are rather
rare. There are no structural differences between them. Fortunately, the absence
of structural differences and the occasional occurrence of intermediate examples are
not the only reasons which induce us to treat all these different-looking forms as
specifically identical. Three of the forms have been bred from one female (by a
corresjjondent of Mr. H. Wernicke, of Blasewitz). These forms are pomponius,
rarik, and h/slthous — ie.,the forms with comparatively small red subraarginal spots
on the hindwing. The various varieties are imitations of species of Papilio feeding
on Aristolockia (P. ascanius, agavus, etc.).
c? ? . Body black ; a grey spot on palpus ; a grey or huffish line ventro-laterally
on abdomen in many specimens. Wings densely scaled above ; forewing varying
from being all black to being crossed by a broad white band, the band crossing cell,
or standing outside cell, or being reduced to a large patch ; a red spot at base,
often absent. Hindwing with or without white central area ; a series of red
submarginal spots, variable in size, the last four larger than the upper three, these
three often absent ; tail variable in length and width, usually spatulate.
Underside : forewing paler than above, scales smaller in distal area; two red
spots at base, one standing in cell and the other before it. Hindwing as black as
above, no distinct cell-lines, three red basal spots, the posterior one produced into a
long line on abdominal fold.
Genitalia: S. Apical edge of harpe very little produced, broadly rounded;
dorso-ventral ridge ending abruptly ventrally, the ventral corner not more elevate
than the rest of the ridge ; central process widened at apex, dentate apically and
dorso-apically, ventral apical angle marked by a rather prominent tooth, proximally
of which there are no other teeth.
Early stages described by Mr. E. D. Jones, I.e., and Burmeister, I.e.
Hah. Brazil ; Eastern Paraguay.
We arrange the varieties in two groups :
A. Red submarginal spots of hindwing large, longer than broad above.
a . P. h/sifhous f. platydesma nov.
Papilio hurrisHianus (V), Boisduval {noii Swainaon 1822, err. det.), Spec. Gin. Lep. i. p. 310. n. 147
(1836) (Brazil).
Papiliu Iiarrisianus, Doubleday, Westw. & Hew., Gen. Diurn. Lep. i. p. 17. n. 184 (1846) (partim ;
Brazil) ; Gray, Cat. Lei,. Ins. Brit. JIus. i. Pap. p. 41. n. 210 (1852) {partim ; Brazil); id..
List Lep. Ins. Brit. Mas. i. Pup. p. 57. n. 222 (1856) (partijn, ; Brazil) ; Menetr., Emim. Corp.
Anim. Mus. Petrop., Lep. I p. 4. n. 68 (1857) (Brazil) ; Felder, ]'erli. Zool. Bat. Ges. Wien xiv.
p. 299. n. 154 (1864) (Brazil) ; Kirby, Cat. Diurn. Lep. p. 524. n. 53 (1871) (partim ; Brazil) ;
Capronn., Ann. Sue. Knt. Belg. xvii. p. 9. n. 7 (1874) (Gavia, August) ; Burm., Descr. Rip.
Argent, v. Lop., Atlas p. 9. n. 23 (1879) (Rio de Jan. ; larva & pupa); Oberth., El. d'Ent. iv. p. 76.
n. 240 (1880) (Brazil) ; Staud., E.cot. Ta,jf. i. p. 15 (1884) (Brazil) ; Haase, Untersuch. Mimicry
i. p. 85 (1893) (Brazil) ; id.. I.e. ii. p. 92 (1893) ; Bonningh., Verh. Yer. Nat. Unterli. ix. p. 27
(1895) (Rio).
c??. White band of forewing broad, continued across cell. On hindwing the
band usually reaching to M-, the last partition of the band merged together with the
red submarginal patch, the band mostly rather narrower in male than in female,
being in the male reduced proximally, the white cell-patch occasionally absent in
( 680 )
this sex, and the white discal patches small and mnch shaded with black ; a row of
seven red submars:inal spots, the last double.
We know this form from the Province of Rio de Janeiro and from Matte Grosso ;
it is doubtless more widely distributed.
In the Tring Museum 7 c?c?, 2 ? ?, from : Tijuca, Rio de Janeiro.
A specimen from Matto Grosso in coll. Adams.
//. /'. lynithous f. harrimaiius Swains. (1822).
PapiUo harrlslamis Swainson, ZooL Jlhtslr. iii. Ent. ii. t. lOO (1822) (hab, ? prob. S. Amer.) ;
Doubl., List Lep. Lis. Brit. Mu». i. p. U (1845) (Brazil) ; id., Westw. & Hew. Gen. Diurn.
iep. i. p. 17. n. 184 (184ti) {partim: Brazil); Gray, Cut. Lep. Ins. Brit. Mm. i. Pa;), p. 41.
n. 210 (1852) (pmiim ; Brazil) ; Kirby, Cat. Dinru. Lep. p. 524. n. 53 (1871) (parlini).
Papilio chn/ilins Boisduval, Spec. Gen. L/'p. i. p. ,311. n. 149 (18311) (partim).
PapiUo allimis Felder, VerJi. Zool. Bot. Ge.t. Wien xiv. p. 2!)'.l. n. 155 (1864) (iioin. mill.) ; id., liei.te
JVovara, Lep. p. 40. n. 35 (18G5) (Brazil) ; Kirby, I.e. p. .524. n. 54 (1871) ; Staad., Exot.
Tag/, i. p. 15 (1884) (Brazil).
A specimen in the British Museum agreeing with Swainson's figure is
presumably the type of harrisianm. Boisduval (183(5) described tlie preceding
form as Iiarrisianus (erroneously spelling the name /larrissiamis), since when that
broad-banded form has always been treated as being the true harrisianus.
Felder's athous is nothing but Swainson's harri.^ianus.
S ?. White band of forevving in male from inner margin to SC, the fir.st and
last spot often absent, the band standing outside cell, but usually touching it at
lower angle, where the band is more or less angulate or interrupted ; occasionally
a vestige of a white patch in apex of cell ; in female the band vestigial fmm lower
angle of cell forward. White discal area of hindwing not extending beyond M',
much narrower in female than in male, the patch occupying in male more than halt
the cell, in female restricted to apex of cell and proximally ill-defined ; upper
three red submarginal spots absent or vestigial in male, more or less distinct in
female.
We know this form from Rio de Janeiro and Espiritn Santo.
In the Tring Museum 4 cJcJ, 2 ??, from: Rio de Janeiro; Leopoldina,
Espiritn Santo.
e'. P. h/sithous f. oedipus Feld. (1865).
Papilio harrisianm, Godart, ICtic. Metli. i.x. Suppl. p. 812. n. 138-9 (1824) (partim).
Pajiilio oedipus Felder, Verb. Zool. Bot. Ge.i. Wien xiv. p. 299. n. 15() (18G4) (nom. nutl.) ; id., Reise
Xorara, Lep. p. 47. n. 30 (1865) (Brazil) ; Kirby, Cut. Diurn. Lip. p. 524. n. 56 (1871) (Brazil);
Staud., Exot. Tiuxf. i. p. 15 (1884) (Brazil).
Papilio sebantianus Oberthur, Et. d'Ent. iv. p. 71'. n. 241. t. 2. fig. 4 (1880) (Brazil).
c? ?. White band of forewing reduced to a large double patch M' — SM-, there
being usually also a small streak behind SW and another in front of M'. the latter
streak occasionally developing to a triangular jiatcli. In one of our males from
Bahia the white double patch is rejilaced by a narrow yellowish one which, anteriorly,
does not reach M'. Hindwing without white band or patch ; the four posterior
submarginal spots large, the anterior ones small and often absent.
Known to us from Bahia, Miuas Geraes, Espiritn Santo.
In the Tring Museum I'i S S from : Bahia; Minas Geraes ; Espiritn Santo.
( "81 )
(l. P. bjsithous f. h/sitJious Hiibii. (1822 ?).
Uerlorirlfs tyxllhoiis Hiibner, Sainml. Exid. Scliw. ii. t. lOf, (1822?) ; Kirby, HihI. ed. ii. p. 90, t. 319.
fig. 3. 4. (190—?).
Papilio claudiux Boisduval, Spec. Gim Lep. i. p. 311. n. 1-19 (1830) (pnrlim).
PapiUo lysilltouK, Doubleday, List Lep. Inn. Brit. Mas. i. p. 14 (1844) (Brazil) ; id., AVestvv. & Hew.,
Geii. Diurii. Lep. i. p. 17. n. 182 (184ij) (partim ; Brazil) : Gray, Cut. Lep. Im. Bril. Mus. i. Pup.
p. 41. D. 209 (1852) (Brazil) ; iJ., List Lep. Ins. Brit, .^fns. i. Pup. p. 57. n. 221 (ISSli) (Brazil) ;
M^niUr., Emim. C'lirp. .Anhu. Miis. Petrop., Lep.K. p. 4. n. G6(I8o7) (Brazil) ; Oberth., Et. d'Ent.iv.
p. 77. n. 242 (1880) (Brazil) ; Staud., E.rut. Tagf. i. p. 14. t. 9 (1884) (Brazil) ; Haase, Untersuch.
Mimicrij i. p. 85 (1893) (Brazil); id., I.e. ii. p. 92. t. 10. fig. 70 (1893); Bonuingh., Verh.
Ver. Natiirw. Unterii. Ilaiiiliuri] ix. p. 27 (1895) (Organ Mts. ; not at Rio) ; Eimer, Orthogen.
p. 137 (1897).
Papiliu hjsitfiuus var. breelfasemtus Weymer, Stett. Eiil. Zeit. Iv. p. 312 (1895) (Rio Grande do Sul).
PapiUo eietendaliis id., I.e. p. 313 (1895) (Rio Grande do Sul).
Papilio agavus, Peters, llhtslr. Zeilxhr. Ent. ii. p. 52 (1897) (Nova Friburgo, ^wiV/'w).
cJ?. Forewing : wliito Viand narrow (rarely huffish on both wings), usually
reaching SC, bnt often not extending forward beyond lower angle of cell {hred-
fasciatus), in most specimens tapering, bnt sometimes of almost even width, variable
in breadth, being posteriorly in some individuals twice as wide as in others ; a row
of distinct white marginal spots in some specimens, such spots being often indicated
in the preceding forms. White band of hindwing usually stopping short at M',
bnt occasionally reaching to M- (e.rteiu/atus), in the latter case the band mostly a
little more distal in position, the cell-patch being smaller and the spots around apex
of cell larger than in ordinary specimens ; red submarginal spots variable in size
and number, the last four smaller than in the preceding forms ; white marginal
spots sometimes a little enlarged : tail sometimes with red spot on underside.
In a male from Minas Geraes in coll. Hewitson (Brit. Mus.), with broad band
on fore- and hindwing and white fringe-spots to forewing, there is some white
scaling proximally of the red submarginal spots of the hindwing.
We know this form from the provinces of Minas Geraes, Rio de Janeiro, Santa
Catharina, Parana, Sao Paulo, Rio Grande do Sul.
In the Tring Museum 35 SS, 6 ?¥, from: Petropolis, January 1898
(J. Foetterle) ; Nova Fribnrgo (Peters) ; Jundiahy ; Sao Paulo ; Castro, Parana
(E. D. Jones) ; Rio Grande do Sul.
e'. P. bjsithous f. vurik Eschsch. (1821).
Papilio rurik- Eschscholtz, in Kotzeb., Reise iii. p. 202. n. 1 (1821) (S. Catharina).
Pajiilio rurikia id.. I.e. t. 1. fig. la. lb (1821) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 41. n. 207
(1852) (Brazil) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 56. n. 218 (18.^6) (Rio Grande do Sul) ;
Felder, Verii. Zool. Bot. Oes. WIen xiv. p. 304. n. 159 (1864) (S. Catharina ; Rio Grande do
Sul) ; Kirby, Cat. Dinrn. Lep. p. 524. n. 58 (1871) (Brazil) ; Stand., Eeol. Tagf. i. p. 15 (1884)
(Brazil); Haase, Untersuch. Mimirrg i. p. 86 (1893) ; Weym., Stett. Ent. Zeit. It. p. 312. n. 3
(1895) (= lains).
Papilio Imrrisianns, Godart, Enc. MM. ix. Snppl. p. 812. n. 138-9 (1824) (parilni).
Papillon laius Roger, Bull. Sue. Linn. Btinleaii.c i. p. 160 (1826) (Brazil).
Papilio laius Boi-sduval, S^)«-. Ght. Lyi. i. p. 311. n. 148 (1836) (Brazil); Doubl., List Lep. Ins.
Brit. Mus. i. p. 14 (184.5) (Brazil) ; id., Westw. A Hew., Gen. Diurn. Lep. i. p. 17. n. 183 (1846)
(Brazil) ; (Jray, Cat. Lep. his. Brit. Mus. i. Pap. p. 41. n. 208 (1852) (BrazU) ; id.. List Lep.
Ins. Brill. Mus. i. Pap. p. 57. n. 220 (1856) (Brazil) ; Ment'tr., Eunm. Corp. Anim. Mus. Petrop.,
Lip. i. p. 4. n. 67 (1857) (Brazil) ; Felder, I.e. p. 304. n. 158 (1864) (var. of rurikia ?) ; Kirby,
l.c. p. 524. n. 57 (1871) ; St.aud., I.e. p. 15 (1881) (Brazil) ; Oberth., Et. iVEnt. xii. p. 5. n. 9. t. 7.
fig. 47 (1888) (Paraguay) ; Haase, l.c. p. 86 (1893) ; Bonningh., Verh. Ver. Nat. Unterh.
Hamhurg ix. p. 27 (1895) (Icaraby) ; Mabilde, Guia Pract. Borbol. Rio Graiuie do Sul p. 47
(1896).
( 682 )
S ? . White band of forewing as in f. oedipus, very variable in width, sometimes
not wider than in broad-banded specimens of f. bjsithous, never continued to costal
mnrgin, occasionally rcdncod to a patch ]\1-— SM- and a streak behind SJI-,
occasionally bnffish ; wliite marginal dots sometimes very distinct. Xo white
band or patch on hindwing ; usually seven red submarginal spots, upper three not
rarely vestigial or absent (from upperside), last four sometimes nearly as large as
in f Of'i/ip>is ; wiiite marginal sjiots often somewhat enlarged ; tail with red spot on
underside in a smiill percentage of specimens (as in Esclischoltz's figure).
We know this form from Paraguay, Santa Catarina and Rio Grande do Sul.
In the Tring Museum 32 $$, 19 ??, from: Sapncay, Paraguay, August
1900 and October 1004 (W. Foster) ; Yhu, Paraguay, September — December bs'-nJ
(Andeer) ; Theresopolis, S. Catharina, November 1904 — February 1905 (J.
Michaelis) ; Rio Grande do Sol.
f. P. lijsithous f. pomponius.
Papilio pomponius HopfFer, Ste/t. Eiit. Zdl. xxvii. p. 25. n. 5 (18lJG) ("'Mexico" error loci) ; Kirhy,
Cat. Dhmi. Lep. p. 607. n. 341 (1871) (-Mexico") ; Honr., BerL Ent. Zeitschi: xxx. p. 29C. t. 6.
fig. 4. ? (1886) (Rio Grande do Sul ; J ? ) ; 'VVejm., Slell. Ent. Zeil. Iv. p. 314. n. C (18115)
(Rio Grande do Sul) : Bbnningh., Verh. Ver. N<it. Uiiterh. JIamhuri/ ix. p. 27 (1805) (one
specimen, 8 miles from Nova Friburgo) ; Mabildo, Guia Pract. Borhol. Rio Grande do Sul
p. 48(1896).
Papilio lysithous, Lathy, Trans. Ent. Soc. Loud. p. 69. n. 36 (1'.104).
(??. Forewing black, usually with wliite marginal spots; white band absent,
often vestigial, rarely represented by a distinct line from near costal to inner margin,
or by a short band extending from BI' to inner margin. Hindwing without white
band or patch, fringe-spots large as a rule ; four to seven red submarginal spots,
the three upper ones usually vestigial ; proximally of these spots sometimes some
white scales.
We know this form from Santa Catharina and Rio Grande do Sul (where its
model, Papilio perr/iebus, is also found) ; BOnninghausen records it from Nova
Friburgo.
In the Tring Museum 8 cJc?, 2 ? ?, from: S. Catharina and Rio Grande
do Sul.
ff'. P. lysitJious f. eupatorion Lucas (1857).
Papilio eupatorion Lucas, in Casteln., Voy. Amer. Sud, Zool. iii. Ent. t. 2. fig. 2 (1857 — coll.
Oberthiir) ; Kirby, Cat. Dinrn. Lep. p. 567. n. 334 (1871) (Am. mer.) ; Oberth., Et. d'Eut. iv.
p. 76. n. 239 (1880) (Brazil ; type).
S. Like f. jMinponius ; but forewing with a broad huffish white marginal
band, marginal spots of hindwing enlarged and white dots at ])roximal side of red
submarginal s])ots distinct, the up])er ones larger than the respective red spots.
Only one specimen known, in coll. Oberthiir, from Brazil.
137. Papilio asius Falir. (1781).
Papilio Eqiies Trnjanus asius Fabricius, .S/jf c. liLs. ii. p. 5. n. 17 (1781) (S. Amer.) ; Fabr., .Ifant. Im.
ii. p. 3. n. 17 (1787) ; Gmelin, .S>«. Nat. i. 5. p. 2229. n. 283 (1790) (S. Amer.) ; Fabr., Ent.
Syet. iii. 1. p. 8. n. 21 (1793).
Papilio Ejues Tnijanne astyagas Drury, lllustr. Ej-ol. Lis. iii. p. 47. t. 35. fig. 4. & Imk-x (1782)
(Rio de Janeiro).
( 683 )
IphicUdes amis, Htibner, SammJ. Exot. Seltni. ii. t. 92 (tSlS?) ; Kirby, ihrd. ed. ii. p. 03. t. 306.
fig. 1. 2 (190- ?).
riipilin asius, Godart, Eric. Meth. ix. p. 55. n. 84 (1819) (Brazil) ; Boisd., Spec. Oin. Lip. i. p. :'O0,
n. 146 (1836) (Rio de Janeiro); Lucas, in Gui^r., Diet. Pitt. Hist. Nat. vii. p. 48 (1838) ; Doubl.,
List Lep. Ins. Brit. Miis. i. p. 14 (1845) (Brazil) ; Butler, Cat. Diurii. Lep. ilescr. Fabr. p. 239.
n. 18 (1869) (Rio de Janeiro) ; Oberth., Ef. d'Eiit. iv. p. 76. n. 2.38 (1880) (Brazil) ; Staud.,
Exot. Tarjf. i. p. 14 (1884); Haase, Untersuch. Jliinicrt/ i. p. 85 (1893) (Brazil) ; Bonningh.,
Verh. Ver. Nat. Unterh. Hamhurrj ix, p. 27 (1895) (Niotheroy, Rio de Janeiro, rare) ; Eimer,
OrthncjPH. p. 137 (1897).
rapilio manlius Perty, Dfhet. Anini. Art. p. 151. t. 29. fig. 1. lb (1830-34) (Minas Geraes).
I'apilio asti/iigiis, Doubleday, Westw. & Hew., Gen. Diuin. Lep. i. p. 17. n. 185 (1840) (Brazil) ;
Gray, Cut. Lep. Ins. Brit. Miis. p. 42. n. 219 (1852) ; id., List Lep. Ins. Brit. Miis. i. Pfip. p. 59.
D. 232 (1850) (Brazil) ; Munetr., Entim. Corp. Auim. Mus. Petrop., Lep. i. p. 4. n. 69 (1857)
(Brazil) ; Felder, Verh. Zoo!. Bot. Ges. Wien xiv. p. 300. n. 100 (1804) (Brazil).
PapiUo asins (!), Kirby, Cut. Diurn. Lep. p. 524. n. 59 (1871) (Brazil).
Pajiilio assius (!), Mabilde, Guia Pract. Burholet. Rio Grande do Sul p. 47 (1896).
A very interesting species, connecting the lijsithous group with the marccllus
group.
S . The band of the wings is variable in width, often entering the cell of the
forewing, this cell-spot occasionally extending down to M' ; scales of forewing
somewhat reduced in width. Besides a spot on the abdominal fold and an anal
spot there are never more than two red spots situated between R' and JP, the ujiper
one being often missing ; these spots are repeated on the underside, the abdominal
one being i)roduced proximad to near base, four more red spots being present in the
basal area, namely two at costal margin, one behind C and the fourth in the cell.
In shape of the hindwing asius resembles more the marcellus group than the
members of the present group.
Scent-organ better developed than in any other member of the present group ;
the abdominal edge bent upwards, forming a narrow pocket ; vein SM- and a narrow
stripe in front of SM- and another behind it covered with small scent-scales, the
scaling having partly a somewhat woolly appearance.
Genitalia: i. Apical lobe of harpe asymmetrical, large, long, denticulate
ventrally ; dorso-ventral ridge with some prominent teeth veutrally ; central process
sjiatulate, entire or feebly denticulate ; ventral process small.
Early stages not known.
Ilah. Brazil and Eastern Paraguay.
In the Tring Museum 52 c? c? from : Bahia ; Minas Geraes ; Espiritu Santo ;
Petropolis ; (G. Foetterle) ; Ilio de Janeiro (E. May) ; Bahuru, Sao Paulo (Dr.
Hempel) ; Castro, Parana (E. D. Jones) ; Yhu, Paraguay (Andeer).
A ? in coll. H. J. Adams.
XIV. Marcellus Group.
For characters see p. 055.
Ten species :
. SC of forewing present.
a. Forewing black from ajjex of cell to distal margin, with a
pale snbmarginal line or row of spots and one or two short
pale costal bands within tbis black area.
o! . Bed line of underside of hindwing edged with white
proximally Species No. 138.
44
( 684)
h'. Red line witliont white border.
a-. Hindwiog, above, with complete black median
band.
a}. Green snbapical cell-bar of forewing
widely separate from green discal
band ......
}?. White or greenish snbapical cell-liar
of forewing continnons with the
discal band.
a*. Fourth and fifth green cell-bars
of forewing merged together .
Zi'. These bars widely separate.
«°. Submargiual spots of fore-
wing more or less rounded.
5\ These sjiots transverse,
linear ....
Jr. Median band of npperside of hindwing widely
interrupted or marked only at costal margin.
el Black subbasal band of upperside of
hindwiug very thin beyond cell
d}. This band broad throughout.
c*. Eed line of underside of hind-
wing stojiping short at cell
d^. Red line continued to brown
distal border ....
h. Forewing with very broad transparent submarginal area
II. SC of forewing absent
Species No. 140.
Species No. 139.
Species No. 141.
Species No. 142.
Species No. 143.
Species No. 144.
Species No. 1 45.
Species No. 146.
Species No. 147.
138. Papilio marcellus Cram. (1777).
Petiver, ihis. Pefw. Cent. p. 50. n. 5112 (1G99) ; Catesby, Nat. Hist. Carol, ii. p. inO. t. 100 (174.'?) ;
Edw., Nat. Hist. Birds i. p. 34. t. 34 (1743).
Papilio Eques Ackivus ajax LinD(5, Sy.it. Nat. ed. x. p. 4G2. n. 2G (1758) (partim, only cit. " Edw.
av. 34 " ; descript. and cit. "Raj. ins. iii. n. 2 " do not apply to this species) ; id., Syst. Nut.
ed. xii. p. 750. n. 32 (1767) (jjartim) ; Fabr., Syst. Ent. p. 455. n. 49 (1775) (partim) ; Esper,
Schm. i. Forts, p. 1. t. 51. fig. 1 (1780) (" S. France " errore) ; Fabr., Spec. Lis. ii. p. 20. n. 79
(17Sl)(;)ar(im)'.; ''^■, ^""'- ^i"- "■ P- 10- °- ''0 (l'?*^^) : Schneid., Europ. Schm. p. 54. n. 3 (1788);
Villiei-s, Car. Linn. Ent. ii. p. 2. n. 1 (1789) (Amer. bor., " Helvetia and Germania '' errore) ;
Jabl. and Uerhat, Naturs. Schm. iii. p. 144. n. 96. t. 42. fig. 5. 6 (1790)(synon. yvur/Zm); Gmelin,
Syst. Nat. i. 5. p. 2238. n. 32 (1790) (partim) ; Fabr., Ent. Syst. iii. 1. p. 33. n. 97 (1793).
Papilio Eques Achivus prolesilaus Linne, Syst. Nat. ed. x. p. 463. n. 29 (1758) (partim ; citat.
Catesb.) ; id., Mus. Lud. Ulr. p. 209. n. 28 (1764) (/«i)7;»i ; v.ar. /3., cit. " Seb. mus. i. t. ii. f. 2 "
excepta) ; Gmelin, Syst. Nat. i. 5. p. 2243. n. 39 (1790) (sub citat.).
Papilio Eques aiax, Lange, in Linne, Syst. Nat. p. 462. n. 26 (1760) (partim).
Papilio Eques Achivus manvllus Cramer, Paj>. Ecot. ii. p. 4. t. 98. fig. F. G. (1777) (spring form).
Papilio aja.r, Seligmann, transd. Houtt., Verz. Ultl. ZeUh. Vogel. ii. p. 54. t. 67 (1772) (Maryland) ;
liorkh., Nat. Eur. Schm. i. p. 112. n. 3 (1788) (" S. France " ; N. Amer.) ; id., /..-. p. 249. n. 3
(1788) (synon. partim) ; Abbot .^- Smitb, Lis. Gcoryia i. p. 7. t. 4 (1797) (metam.) ; Fabr.,
Epit. Ent. Nomencl. p. 12H (1797) (i>arHm ; Am. bor.) ; Paliaot, his. Afr. Amer. p. 71. Le/i. t. 2.
fig. 2 (1805) ; Ochsenh., Schm. Enr. i. 2. p. 117. n. 1 (1808) ("Italy and S. France," errore) ;
Laspeyr., Jen. Ally. Lit. Zeil. p. 98 (1809) (LiuniS's ajax is not the iij'ax of Esper !) ; Ochsenh.,
Schm. Eur. iv. p. 149 (1K16); Godart, Enc. MHh. ix. p. 52. n. 79 (1819) [synon. partim); Lucas,
Lep. Enr. p. 9. t. 14. fig. 2 (1834) ("Greek Archip." ; fig. mala) ; Doubl., List Lep. Ins. Ilrit.
Mus. i. p. 9 (1845) ; id., in Westw., Arc. Ent. i. p. 61 (1845) (Florida, habits) ; id., Westw. &
( 685 )
Hew., Gen. Dhirn. Up. i. p. 15. n. 133 (1846) ; Kirtl., Proc. Eiit. Soc. Land. (2). i. p. 101 (1851)
(south shore of L. Erie ; Ohio; larva on Atin/ia triloba) ; Gray, List Lej>. Ins. Brit. Mim.i.
p. 33. n. 155 (185'J) {imircellus = var. of ajax) ; Lucas, in Chenu, Em. Hist. Nut. Pap. i. fig. 1
(1851-53); Dutreux, Stett. Ent. Ze.it. xv. p. 142 (1854) (Faro, Portugal ;— imported, or local,
crioneous); Keferst., ihid. p. 330 (1854) (Dutreux's ujaj: from Portugal is lyViJ- Boisd.); Lucas,
/;((//. Sue. Ent. France p. 9 (1855) (" Portugal " teste Keferstein, " Greece " teste Lucas); Gray,
Li.it Lep. Ins. Brit. Miis. i. p. 44. n. 1G3 (185(5) ; Men^tr. Enum. Corp. Anim. Petrop., Lip. i.,
p. 3. n. 47 (1857) ; Gosse, Letters from Alabama p. 51. figs. (185'.)) (larva, pupa) ; id., I.e. p. 148
(1859) ; Morris, Si/n. Lep. \. Am. i. p. 8. n. 11 (1862) (ajaj: = marccllus) ; Weidem., Proc.
Ent. Soc. Philad. ii. p. 146 (186.3) (marcellus ?); Kirkp., ibid. iii. p. 328 (1864) (Cleveland, Ohio,
common) ; Butl., Cot. Ditirn. Lep. descr. Fabr. p. 241. u. 2(; (1869) ; Parker, Amer. Entom. ii.
p. 175 (1870) (Iowa) ; Edw., Canad. Ent. ii. p. 115. 133. 162 (1870) (-yVix and marcellus oue
species, proved by breeding) ; id.. I.e. iii. p. 70 (1871) (bred) ; Scudder, ibid. iv. p. 74. 84
(1872) (Georgia, Abbot's MS.) ; Meldola, Ann. Mag. N. U. (4). xii. p. 301 (1873) (.substance
waste in pupal state) ; Pagenst., Verh. Nat. Med. Ver. Heidelb. i. p. 108 (1874) ; Gerh., Maero-
Lep. N. Amer. p. 25. n. 443 (1878) ; Dury, Cincinnati Soc. Nat. Uist. i. p. 12 (1878) (Cine,
common) ; French, Rep. Ins. Illin. vii. p. 135 (1878) ; Morris, Canad. Ent. xi. p. 203 (1879)
(Ontario, local) ; Oberth., Et. d'Ent. iv. p. 65. n. 176 (1880) ; Clayp., Canaxl. Ent. xii. p. 120
(1880) (Ohio, April) ; Middl., Rep. Ins. lUin. x. p. 74 (1881) ; Edwards, Canad. Ent. xiv. p. 27
(1882) (connection between ajai^, wahlii and telamonides ; life history) ; Auriv., Kongl. Sr. Vet.
AI-. Handl. xix. 5. p. 30. n. 28a (1882) (recensio critica) ; Gruber, Jena. Zeitseh. Naturw. xvii.
p. 473. t. 7, f. 16-19. 1. (1884) ; id., I.e. xviii. p. 881 (1884) ; id., Papilio iv. p. 88. t. 2. f. 16-19
(1884) (transf.) ; id.. I.e. p. 188 (1884) (correction).; Edw., Ent. Amer. i. p. 213 (1885) (larvae
will not eat sassafras and spice-wood) ; id., Canad. Ent. xviii. p. 15 (1886) (larva on Paupaic,
refusing spice-wood and sassafras in W. Va.) ; French, Butt. Ea.it. U. St. p. 84 (188i!) ;
Hancock, Amer. Nat. xx. p. 970 (1886) (Chicago, migrating northward) ; Hulst, Ent. Amer. ii.
p. 182 (1886) (Long L, end June '86, one specira.) ; Riley, Insect Life i. p. 161 (1888) (parasite :
Tragus cxesoritts. Pimpla aiundipcs) ; Edw., Bttll. U. St. Nat. Mas. xxxv. p. 9 (1889) (liter, on
transf.); Skinner it Aaron, Cnwarf. £.'«(. xxi. p. 126 (1889) (Philadelphia, rare) ; Eimer, Arlb.
Veneandtseh. Selimett. p. 195 (1889) ; Pack., Fifth Rept. U.S. Ent. Comm. p. 669 (1890) (larva
on Asimina triluha) ; Mayn., Man. N. Amer. Butt. p. 3. n. 1 (1891) ; Staley, Canad. Ent. xxiv.
p. 203 (1892) (Marshall, Missouri, f. telamonides rare, marcellus not uncommon) ; Davis, Journ.
N. Vorl: Ent. Soc. i. p. 47 (1893) (Staten I., N.Y., 2 ex.) ; Jones, Ent. News iv. p. 190 (1833)
(Wilmington, N.C.): Beutenm,, Bidl. Amer. Mus. N. II. v. p. 241 (1893) (N. York ; descr. of
1., p., i.) ; Weed, Psi/clie vii. p. 130 (1894) (N.E. Miss.) ; Davis, Journ. N. York Ent. Soc. iii.
p. 141 (1895) (Staten 1., N.Y., seen June 30) ; Osburn, Ent. News vi. p. 282. n. 43 (1895)
(Tennessee, common, iv. to x., two broods) ; Deard., ibid. vi. p. 296 (1895) (Lonsdale, R.I.,
July 19, one ex.) ; Langl., ibid. vi. p. 314 (1895) (Chicago) ; Hills, Canad. Ent. xxviii. p. 190
(1896) (Toronto, June); Anonym., ibid, xxviii. p. 190 (1896) (Port Hope, Ontario ; never before
observed so far east) ; Gibson, ihid. xxviii. p. 294 (1896) (Toronto, June) ; Bubua, Ent.
Neu;s viii. p. 98 (1897) (Cleveland, Ohio ; plentiful, three forms) : Edw., Butt. N. Amer. iii.
Pajj. V. (1897) (results of breeding); Gibson, Rtjil. Ent. Soc. Ontario xxvii. p. 105 (1897)
(Toronto, June and July) ; Moffat, ibid. p. 109. n. 79(1897) (Pt. Hope, end of May and June ;
Toronto); Dnzce, Bull.' Buffalo Soc. N. Sc. V. p. 107. n. 1 (1897) (Buffalo) ; Snyder, Ouiad.
Ent. xxix. p. 119 (1897) (Evanston, 111., very rare) ; Christ, Mitt. Schweiz. Ent. Gcs. ix. p. 279
(1897) ; Eimer, Orthogen. p. 399 (1897) ; Tro-xler, Canad. Ent. xxx. p. 257 (1898) (Louisville,
Ky., ? with yellow markings. May) ; Holl., Bait. Boole p. 3u7. n. 1. t. 2. f. 14, t. 6. fig. 11. 12,
t. 44. fig. 1-4(1899); Wenzel, Ent. News xi. p. 643 (I'.IOO) (Wildwood.N.J., two ex.); Beutenm.,
Butt. N. York City p. 7. n. 5. fig. ^ (1902) ; Comst., Ent. Neais xiii. p. 76 (1902) (L. Jo.sephine,
Fla.) ; Walk., Re/Jl. Ent. Soc. Ontario xxxii. p. 85 (1902) (Point Pelee, Leamington); Briml.
& Sherm., Ent. News xiv. p. 230 (1903) (Raleigh, N.C.).
Papilio jrrotesiluus, Fabricius, Ej). Ent. Nomeucl. p. 128 (1797) (partim ; Am. Sept.).
Princeps heroicus ajax Hiibner, Samml. Exot. Sehm. i. t. 106 (1806 — ?).
Iphirlides ajax, id,, Vcrz. bek. Srhm.. p. 82. u. 83i; (1818*); Soudd., Proe. Boston Soc. Nil. xvi.
p. 117 (1873) (substance waste ; u-ahhi and telamonides from winteriug pupae, marcellus
offspring of both these forms ; id., Amer. Nat. viii. p. 257 (1874) ; Meldola, Ann. Mag. N. II.
(4). xiv. p. 239 (1874) (substance waste) ; Scudder, Butt. Eaxt. U. St. & Can. ii. p. 1264. t. 15.
fig. 11. t. 35. fig. 26—29. genit., t. 56. fig. 9. bead, etc. (1889) ; id.. Psyche viii. p. 208. t. 5. f. 2. 1.
]'uv. (1898) ; Dyar, Bull. U. St. Nat. Mus. Iii. p. 2. n. 5 (1902) ; Kirby, in Hilbn., Samml.
Exot. Sckmett. ed. ii. p. 99. t. 106. fig. 3. 4 (190- ?).
Papilio marcellus, Boisduval & Leconte, Hist. Gin. Lip. Amer. Sept. i. p. 8. t. 2. fig. 1 — 4. 1., p., i.
( 686 )
(1835) (summer form) ; Lucas, in Gui5r., Diet. PHI. Hid. Nal. vii. p. 4') (1838) ; Doubl., [J^l
Lep. Jus. Brit. ilus. i. p. 8 (1845) ; id., ia Westw.. .1 re. Ent. i. p. Gl (1845) (Virginia, Uhio. etc.) ;
Doubl., Westw. .t Hew., Geii. Diimi. Lep. i. p. 15. n. 13-2 (ISlCi) ; Kirtl., Proe. Ent. Sor. L,mtl.
(2). i. p. 101 (1851) (differ.s from ajnx in flight) ; Mem'tr., Eitum. Corp. Aniin. Petrop., Lip, i.
p. 3. n. 46 (1857) ; Newm., Prne. Ent. Soe. Phihd. i. p. 2G (ISGI) (N. Jersey ; on Paicpan-) ;
Morris, Si/n. Lep. X. Am. i. p. a. n. 12 (1862) ; Weidem., Proc. Ent. Soc. Philml. iii. p. 14G (180:!)
{^ajax'i); Kirkp., Prvc. Ent. Soc. Phihid. iii. p. 328 (18G4) (Cleveland, Ohio, common);
'Eifi., Amer. Entom. ii. p. 305 (1870) (ovipositing): id., C'««a(/. Ent. iii. p. 70 (1870) (bred);
Saund., ibid. vi. p. 140 (1874) (Essex Co.) ; Ison, R'-pl. Ent. Soc. Ontario p. 15 (187G) (Cleveland,
rather common) ; Moffat, //</(/. p. 10 (1881) (Long Point & Ridgeway) ; Mundt, Cnnad. Enl.
XV. p. 80 (1883) (Pontiac, lUin., May and later) ; Saund., ibid. xvi. p. 50 (1881) (Lake Erie) ;
id., Rppt. Enl. Soc. Ontario xv. p. 20 (1885) (Point Pelee, L. Erie) ; Pack., Fifth lirpt. I'.S.
Ent. Coinm. p. &&'i (1800) (larva on Asimina triloba) ; Moffat, Rept. Ent. So_'. IJnIario xxvii.
p. 70 (1807) (London, Ont.) ; Bethune, ibid, xxviii. p. 33 (1808) (Pt. Hope); id., ibid. xxx.
p. 101 (1000) (Pt. Hope).
Pupllio protesilau.i " Drury," Reitzenstein, Cal. Lep. N. Orleans (18G3) (Greville, rare ; —doubtless
an error of identification, probably form of marcellus).
Pathysa marcellus, Reakirt, Proc. Ent. Soc. Philad. iii. p. 504 (18G4).
Papilio iijax var. marcellus, Fletcher, Canad. Ent. xxxi. p. 8 (1800) (Cowichan Rd., Vancouver I.,
strange occurrence !).
Owing to Linnti's short and vague descriptions, and his freqnent quotation of
figures and previous descriptions which have nothing to do with the animal
descrilied, the uomenclature of many Linnean species is much involved. Tbe
nomenclatorial difficulties created by Linn^ have often been enhanced by post-
Linnean authors, many of whom were quite arbitrary in the application of names
and non-critical to an amazing degree. However, the difficnlties would have long
disappeared to a great extent, if the authors of the second third of the nineteenth
century, when it was still early enough to mend matters without much inconvenience,
had had the courage of being thorough in nomenclatorial matters. The Asimina
Swallowtail of North America, usually called ajax Linne, is a striking instance.
The Swallowtails found in the Atlantic States are all common insects. Tbere
occur five species, not counting the southern species F. palamedcs and polydaimm.
As three of the five are recognisably described by Linn^ in Syst. Nat. ed. x. and
later, there remain two — namely, the A.simina Swallowtail allied to the European
jjodalirius and the Parsnip Swallowtail allied to machaon. For which of the two
species did Linn^ propose the name ajax f
Linne's description in St/st. Nat. ed. x. p. 462 (1758) is as follows :
Ajax. 26. P.E. alis obtuse caudatis concoloribns fuscis ; fasciis flavcsceutibus,
angulo ani fulvo.
Raj. ins. III. /(. 2. Edii\ ai\ 34.
Habitat iu America boreali.
This description certainly does not fit the Asimina Swallowtail, which has a
red anal angle ; but applies very well to the Parsnip Papilio, Linne describing tlie
similar, but more extended yellow machaon as follows :
Machaon. 27. P.E. alis caudatis concoloribns flavis ; fasciis fnscis : angulo
ani fulvo.
AVe draw attention to avyulo nni fnleo appearing in both dcscrij)tions.
Linne quotes two j)revious authors under ajax. H<iwever, the insects described
by Eay and the one described and figured by Edwards are not only widely different
from one another, but Linne's description fits neither the one nor the other. The
insect of Ray is doubtless the yellow female or the male of Papilio glaucus. Kay's
description is as follows :
( 687 )
2. Papilio alis' ampUssimis, Jlavicante et nigra coloiihus pulcherrime
vai-iegatis, interiorihus caudatis, major Virginiana. Diurnarum prima, omnium
maxima., Monffet. Theat. Insect, ji. 98.
Haec praecedenti,* excepta magnitudine, qna earn excellit, simillima est, ut
dnbitem an accidentaliter potius qnam specifice, lit vocant, alj ea differat :
ideoque qnamvis exotica sit, a praecedeute minime separandam censemus.
Mouflfet's ligure referred to by Ray is a bad representation of a yellow P. glaucus.
The figure is very large, the apex of the forewing being produced into a long acute
hook, and the tail being long, curved and pointed. Liiin6 says of ajax : alis obtuse (!)
caudatis.
The second reference under Linne's ajax is Edwards, Nat. Hist. Birds i. t. 34.
The insect here represented is the Asimina Swallowtail. The anal spot is
expressly described in the text as being bright red, while Licnd calls that spot of
ajax fulvous, as in the case of machaon. How is it that Linne' quotes this figure
under ajax, althongh it does not at all conform to the description ? We think it was
a mere oversight ; he did not mean to put the reference to Edwards's figure under
ajax but under protesilaus, as appears to us proved by the description of protesilaus
and the various references given by Linnd under that heading. The description of
protesilaus and the references are in Sg.st. Nat. ed. x. p. 463, as follows :
Protesilaus. 29. P.E. alis caudatis snbconcoloribus albidis : fasciis fnscis :
unica subtus sanguinea, angulo ani rubro.
Pet. Mus. 50. n. 502.
Sloan, jam. 2. p. 218. t. 239./. I. 2.
Mer. surin. 43. f. 43.
Sed. mus. 1. t. U ./. 2.
Catesb. car. 2. t. "luO.
Habitat in America septentrionali.
Sinillimus Podalirio Europae australis </• Africae ; an satis dicersus'^
Now, the first citation under protesilaus (Petiver) and the last quotation
(Catesby) refer both to recognisable descriptions and figures of the North American
Asimina Swallowtail ! 1 Meriau's figure rej)resents the white South American insect
usually known by the name oi protesilaus. Seba's and Catesby's figures represent
some species of the Nymphalid genus 2l/i"_^«/«ra. Since Linne considered all these
insects as being one species — which, moreover, was in his opinion only doubtfully
distinct from the European podalirius — is it likely that he believed Edwards's
figure to represent a diflerent species ? An unambiguous answer is given by Linne
in Mus. Lud. Ulr. p. 209 (1764). In this work, in which a page is devoted to each
species of Lepidoptera, the descriptions being fsir superior to those in Sg.<st. Nat.,
the figure of Edwards is quoted under protesilaus I, where it ought to have been
quoted also in Sgst. Nat.
Perhaps the most striking evidence that Linne himself treated the Parsnip
Swallowtail as being ajax is offered in the 12th edition of Syst. Nat. Here the
Eastern Palaearctic P. xutlius is described on p. 751, being placed after P. machaon,
while P. ajax is placed before P. machaon. This P. xuthus Linne describes as
being very similar to p. ajax (" simillimns P. ajaci "). Now, P. xuthus is utterly
different from the Asimina Swallowtail, while it resembles P. machaon as well as
the Parsnip Swallowtail.
Considering all this evidence without bias, it appears to us to be beyond doubt
* Namelv, machaon.
( 688 )
that the name ajax "Linnt^" caiuidt jiossilily be eiujjloyi'il for the Asimina
Swallowtail. lu our opinion the descriiition nndcr the heading (ijnx was meant for
the Swallowtail now called poI>/xi'nes or asfcn'^s. Bnt as the descrii)tion is quite
insufficient for precise recognition, and as, further, by the reference to Ray yellow
specimens oi Fapilio f/lauciis are inclnded in the " species ajax Linne?," we deem it
correct to treat the name ajax as a synonym of glaucus* which name is described
before ajax, and to quote it again as a doubtful synonym under poh/xenes. A
change in the names of the North American Swallowtails is thus rendered
unnecessary, except that the name ajax is dropped altogether, the name marcellus,
which comes nest in priority, and with which everybody is familiar, being employed
instead for the species. The wings of profesilaiis being described by Liuue as white,
we restrict this name to the white insect figured by Merian and Glerck.
(? ? . Antenna tawny, carinate beneath (except club), the two patches of
sensory hairs of each segment being impressed; scaling of upperside usually
fallen oflF, scales of distal segments brown or black, of proximal segments white.
Tibiae and tarsi pale green, the former bearing scales in fresh specimens (the
scaling may be present also on the tarsi when the specimen emerges from
the chrysalis, but we have no information on this point) ; mid- and hindtibial
spurs longer than the tibia is broad.
Forewing with eight pale bands, the third very narrow, not extending beyond
cell, fourth and fifth continnous with the discal band, sixth distally of apex of cell,
reaching down to R-, seventh represented by a spot situated before SC-* ^ eighth
submarginal, more or less separated into sjiots, of which the posterior ones are
Inniform.
Red line of underside of hindwing bordered with white proximally, distally
of the red line and pro.ximally of white one a black band, the distal black band
being the thinner one, the white line situated just outside cell, being contiguous
with the cross-veins D-. Scales of upper surface nearly all dentate.
Neuratiou : Praecostal spur of hindwing at three-fourths of basal cellule.
Scent-organ resembling that of F. protesilaiis, being very diti'erent from that of
P. philokms; vein SM^ covered with white scales of the ordinary shape but obliquely
truncate at apex, being either more or less dentate or entire ; between this vein
and abdominal edge of fold a dense covering of thin, white, long hairs, intermixed
with a small proportion of much longer and thicker ones ; these latter slightly but
distinctly widening towards apex, which is rather abruptly narrowed to a point.
Genitalia : S. Tenth tergite narrow, slightly-widened distally, divided into
three lobes. Dorso-vcntral ridge of harpe not extending ventrad beyond the central
process, being almost continuous with the non-dentate dorsal edge of the apical
lobe of the harpe ; central process long, curved ventrad ; ventral process triangular.
? . Walls of vaginal cavity very strongly wrinkled, the wall elevate in front of
the vaginal aperture, but there is no sharidy separate process or ridge.
Early stages well known. Except in first stage, the segments of the caterpillar
bear each six black or brown transverse lines, besides a short line in front and
another behind which are connected with one amther; the intersj)aces between
lines 2 and 3 and between 4 and 5 very narrow, these lines being usually more
or less merged together. Frontal jiromiuences of chrysalis divergent, carinate
above, fruns broadly concave between them in dorsal asjject. Thoracic prominence
vertical, slightly concave behind, lateral carina coutinueil to cremaster without
* See page'4H.
( 689 )
Dreak ; punctnratiou of abdomen rather dense, segments 2 to 8 with transverse
carina before apex, tenth segment hardly one-third longer than broad ; hooks of
cremaster pale. Food-plant : Asimina triloba and other species of Asimina,
rarely Ei'tcaceae and Lauraceae.
A seasonally variable species. Two principal forms : a variable spring form
emerging from hibernated pupae, and a summer form emerging from pupae which
have not wintered. As the size and markings of the specimens appear to depend
on the temperature of the period critical for the chrysalis, the late spring specimens
approach the summer and autumn specimens, and the early Spring individuals from
the most southern districts of the range of the species resemble in certain characters
the late spring specimens of the more northern districts. The time of appearance
in spring varies also with the latitude of the locality,
I. Spring forms. Hairs on frons long. Pupa hibernated.
a. P. ?narcellus f. hib. marcellus Cram. (1777).
Fap'din Eqiies Achivus marcellus Cramer, Paj). Exot. ii. p. 4. t. 98. fig. F. G (1777) (Virginia) ; Stoll,
in Cram., Pap. Exot. iv., Onlre Si/st. p. 3. n. 4 (nS2) (= ajax, false).
Papilio marcellus Stoll, I.e. iv. p. 195 (1782) (= ajax L., false).
Papilio ajax, Abbot & Smith, I.e.; Boisd. & Lee, Hht. Gen. Lep. Amh: Sept. i. p. 4. t. 1.
fig. l_4, l.,p., i (1833) ; Boisd., Spec. Gen. Lip. i. p. 258. n. 82 (1836) ; Felder, Verh. Zool. Bot.
Ges. Wicn xiv. p. 303. n. 20G (1864).
Papilio ajax var. u-alshi Edwards, Butt. N. Am. i. Pap. i. fig. 1—5 (18G8) ; id., Caiiad. Eut. iii. p. 70
(1871) ; Dury, Ciurimiati Soc. Nat. Uisl. i. p. 12 (1878) (Cine, common).
Papilio ajax var. walslii subvar. abboti Edwards, Butt. N. Am. i. Pap. 1. fig. 6 (1868).
Papilio ajax dim. var. walslii id.. Trans. Anier. Ent. Soc. vi. p. 9 (1877)(= marcellus).
Papilio ajax sub- var. ubl/oti id., I.e. (1877).
Papilio ajax var. abbotii, Gerhard, Macro-Lep. N. Amer. p. 25. n. 443c (1878).
Papilio icalshi, Mundt, Canad. Eut. xv. \t. 87 (1883) (Pontiac, Illin., March).
Papilio abboti, id., l.c. xv. p. 87 (1883) (Pontiac, Illin., March).
Papilio ajax iralshi sub-rar. abbotii, Scudder, Biitl. E. U. St. and Cau. ii. p. 1269 (1889).
Papilio ajax walslii, Eimer, Artb. Veru-andtsck. Schmitt. p. 195. t. 3. fig. 12 (1889); Heink, £««.
News xiv. p. 335 (1903) (Meramec Highlands. St. Louis Co., April 12).
Papdio ajax var. abbotii, Holland, Butt. Book p. 3U7 (1899).
The early spring form.
cJ?. Pale bands broad ; usually no distinct pale band along abdominal fold
of hindwing ; only tip of tail white ; red anal spot large, not divided into spots.
Specimens with some of the red discal spots of the hindwing more or less
distinct on upperside are ab. abboti.
b'. P. marcellus f. loc. ]i\h. Jioridensis Holl. (1899).
Papilio ajax winter iotm flnridaisis Holland, Bull. Book p. 307. t. 44. fig. 2 (1899).
The early spring form of Florida.
S ?. The black bands broader than in f. hib. marcellus.
c. P. marcellus gen. hib. telamonides Feld. (18G4).
Papdio telamonides Felder, Verh. Zool. Bot. Ges. Wicn xiv. p. 303. n. 205 (1804) ; id., Reise Noraru,
Lep. p. 60. n. 46 (1805) ; Mundt, Canad. Ent. xv. p. 88 (1883) (Pontiac, Illin., later than toalshi) ;
Pack., Fifth Rcpt. U.S. Ent. Comm. p. 669 (1890) (larva on Asimiiia triloba).
Papilio ajax var. telamonides, Edwards, Butt. N. Am. i. Paj). ii. fig. 1—8 (1868) ; Dury, Cincinnati
Soc. Nat. Hist. i. p. 12 (1878) (Cine, common).
Papilio ajax dim. var. telamonides, Eil wards, Trans. Amer. Ent. Soc. vi. p. 9 (1877).
Pajnlio ajax telamonides Eimer, Artb. Verirandtsch. Schm. p. 195 (1889).
Papilio ajax f. telamonides, Moffat, Canad. Eut. xxxiv. p. 170 (1902) (Kingsville, Lake Erie, May).
( 690 )
The late spring form.
cj?. Wings rather longer than in gen. hib. marccUus; black bands on the
whole a little wider ; white colour of tail more extended laterally. Transition to
the summer form.
II. Summer and autumn form. Large ; hairs on frons short ; wings longer
and forewing more falcate than in the sj)ring forms ; white colour of tail extending
laterally' at least to middle. Only one form, the various broods not aiipearing to
diifer from one another.
d'. P. marcellus f. aest. lecontei nom. nov.
Pajnlio marcellus, Boisduval & Leconte {nnn Cramer, 1777, err. det.) ; Flixt. Gen. Lip. Amir. Sept.
i. p. 8. t. 2. fig. 1-4. 1., p., i (1833) ; Boisd., .V- Gm. Up. i. p. 257. n. 81 (1836) ; Felder,
Verh. Znol. Hot. Ges. Wicn xiv. p. 303. n. 204 (1804).
Pripilio ajax var. marcellus, Edwards, Butt. N. Am. i. Pap. iii. fig. 1 — G (18(58).
Papilio ajax dim. var. marcellus, id., Trails. Amer. Eiit. Snc. vi. p. 0 (1877).
Papil/o ajax var. marcellus, Dury, Cinchinati Soc. Nat. Hist. i. p. 12 (1878) (Cine, common).
Papilio ajax marcellus Eimer, Arlb. Vcrwandtuch. Schm. p. 195. t. 4. fig. 5 (188'J).
/pliiclides ajax ajax, Scudder, Butt. East U. St. <t Can. ii. p. 1264. t. 15. fig. 11 (1889).
There is no valid name available for this form, which was erroneonsly figured
by Boisduval and Leconte as the true marcel/us of Cramer. We do not know what
uomenclatorial considerations induced W. H. Edwards to rename the early spring
form as walshi, sinking marcellus Cram, as a synonym of iralsJn, and to employ
marcellus Boisd. & Lee. (1834 ; 7ion Cramer, 1779) for the summer form. Scudder
correctly referred to the early spring form as marcellus Cram.
(??. Black bands of body and wings broader than in the spring forms;
hindwing usually with distinct pale band along abdominal fold, second red sjjot
of upperside much reduced, in male very often, in female always, absent.
Name-type of lecontei from Nashville.
Hah. P. marcellus occurs from Florida and Texas to southern Canada, west-
wards extending to the Mississippi plains ; in the northern districts it is more a
visitor than a resident.
One specimen found on Vancouver Island ; chrysalis imported (?).
In the Tring Museum 120-odd specimens, and several larvae and pupae from :
Nashville, Tennessee (W. Osbnru) ; Jetferson Co., Kentucky (C. Troxler) ; Nelson Co.,
W. Virginia (Wirt Robinson) ; Sanford, Florida.
139. Papilio marcellinus Doubl. (1845).
Sloane, Voy. Jamaica ii. p. 218. t. 239. fig. 17. 18 (1725).
Papilio Eques Achirus prote.Hilaus Linni^, Mas. Lad. Vlr. p. 209. n. 28 (1764) (sub citat. : "Sloan,
jam. 2. p. 218. t. 2.39. f. 17, 18") ; Drury, Illuslr. E.rot. Ins. i. p. 45. (and Index) t. 22. fig. 3. 4
(1770) (Jamaica).
Pajnlio Eques Achivus stnon Fabricius {non Poda, 1761), Sysl. Ent. p. 452. n. 39 (1775) (partim;
" India ") ; id., Spec. Ins. ii. p. 15. n. 59 (1781) (partiin) ; Goeze, Ent. Beijtr. iii. 1. p. 72. n. 7
(1779) (partim); Cram., Pap. Exot. iv. p. 57. t. 317. fig. C. D (1780) ("Jamaica"; syn.
excl.) ; Fabr., 2Iant. Ins. ii. p. 8. n. 67 (1787) (parlim) ; Gmel., Si/st. Nat. i. 5. p. 2241.
n. 329 (1790) (partim) ; Fabr, Ent. %.?/. iii. 1. p. 26. n. 75 (1793) (par'tim ; " India ").
Papilio sinon Fabricius, Nomencl. p. 128 (1797) ("India"); Godart, Enc. Meth. \x. p. 53. n. 80
(1819) (partim) ; Boisd. & Lee, Hist. Gin. Lip. Amir. Sept. p. 11 (1833) (partim) ; Boisd.,
Spec. Gen. Lip. i. p. 260. n. 83 (1836) (partim) ; Morris, Si/n. Lep. N. Am. p. 9. n. 13 (1862)
(partim ; sub synon.) ; Felder, Verh. Zool. Bot. Ges. Wien .\iv. p. 302. u. 200 (1864) (/mrlim) ;
Bull., Cat. Diurn. Lep. descr. Fabr. p. 240. n. 25 (1869) (Jamaica ; type in coll. Banks) ;
Kirby, Cat. Diurn. Lep. p. 557. n. 269 (1871) (Jamaica) ; Bull., P roc. Zool. Soc. Land. p. 481.
( 691 )
n. 34 (1878) (Jamaica) ; Mosohl., Ahh. Senl-piib. Nat. Ges. xiv. iSrIini. Jamaica p. 2(i. n. 1
(1888) ; Eimer, Arth. VerwandUch. Srjim. p. 183. t. 3. fig. 11 (1889) (Jamaica); id., Orlhnyen.
pp. 44. 131. 1.-59. 305 (1897) (Jamaica).
Iphidides mnon, Hiibner, Ven. helc. Sfhm. p. 82. n. 838 (1818 ?) (imrtim).
I'apiUo marcelliniis Doubleday, List Lep. Ins. Brit. Mas. i. p. 8 (1845) (nam. nor. loco prutcaHaus
Drury ; Jamaica) ; id., Westw. & Hew., Gen. Diurn. Lep. i. p. ly. n. 131 (1846) (Jamaica) ;
Gray, Cat. Lep. Lis. Drit. Mux. i. p. §2. n. 152 (1852) (Jamaica) ; id., Lid Lep. Ins. Brit.
Mus. i. p. 44. n. 100 (1851')) (Jamaica) ; Feld., Verb. Zmil. lint. Ges. Wien xiv. p. 303. n. 20;i
(18G4) (Jamaica) ; Oberlb., E/. d'Ent. iv. p. 65. n. 172 (1880) (Jamaica).
S ? . Abdominal tergites edged with white, the white edge incomplete above.
Tibiae and tarsi pale green, tarsal segments somewhat ochraceons at apices ; no
scaling ; external spur of mid- and hiudtibiae a little longer than internal.
Wings, ttpperside, with pale green bands ; scales nearly all denticulate, those
of the pale markings small, comparatively few in number, easily falling off, leaving
the bands naked. Forewing : a basal and a subbasal band from costal to inner
margin, a very thin line across cell just proximally of M-, a broad band beyond
middle of cell, divided costally by a black line or spot, the band continuous with
a broad discal band ; a short costal band SO' — R' proximally of subcostal fork ; a
row of seven submarginal spots, there being no spot or only a trace of one behind
M-. Hiudwing: black submedian band complete, reaching to black distal border.
Pale bands of underside scaled, the scales smaller than the brown ones on
forewing. Red hne of hindwing very broad, reaching to brown distal border of
wing, the line edged with black distally, especially in costal region, thinly edged
with black jiroximally.
Scent-organ : Fold small, scent-scales long, similar to those of agesilaus, but
longer and more slender.
Genitalia : i. Tenth tergite elongate but broad, divided aincally by two narrow
incisions into three lobes. Dorso-ventral ridge of harjie reaching ventral edge
about middle, ending dorsally in an acute, somewhat conical process or tooth ;
ventral process short but distinct. ? not dissected.
Early stages and food-plant not known.
As Poda gave the name sinon in 1761 to the European P. podaliriu.i, which
belongs also to the present section of Pajjilio, the same name cannot be em]iloyed
for the Jamaican insect. Doubleday was quite right in renaming the species.
Ilab. Jamaica.
In the Triug Museum (i iS from : S. Thomas, .Jamaica, May 1892 (Taylor) ;
"Jamaica."
Several pairs in coll. Oberthlir.
UO. Papilio celadon Lucas (1852).
Seba, Thesniir. iv. p. 45. t. 37. fig. 13. 14 (17G4).
Papilio Erjucs Achivus sinon Cramer (non Poda, 17G1 ; non Cramer, fig. C. D.), Pup. Exoi. iv.
p. 57. t. 317. fig. E, r (1780) ; Fabricius, Maiit. Ins. ii. p. 8. n. 67 (1787) {partim) ; Jabl. &
Herbst, Naturs. Schm. iii. p. 159. n. 101. t. 44. fig. 5 (1788) {partim ; fig. copy of Cramer's fig. E);
Gmelin, fiyst. Nat. i. 5. p. 2241. n. 329 (1790) (partim ; " India " !) ; Fabr., Ent. Si/st. iii. 1. p. 26.
n. 75 (1793) (partim ; "India" !).
Papilio siiwn, Godart, Enc. Mitli.ix. p. 53. n. 80(1819) (2)artim) ; Boisd. & Lee, Hist. Gen. Lij>.
Am(r. Sept. p. 1 1 (IS'AS) (partim) ; Boisd., Sjm: Gen. Lep. i. p. 260. n. 83 (1836) (partim) ; Duncan,
in Jard., Nat. Lihr., Ent. v. p. 106. t. 4. fig. 2 (1843) (syn. partim) ; Poey, Mem. Real Soe. Eron.
Hahana p. 236 (1846); Menetr., Enum. Corp. Anim. Mus. Pclr., Lip. i. p. 3. n. 45 (1857) ( partim) ;
Herr.-Sch., Corresp. Bl. Zool. Min. Ver. Regensh. p. 172. n. 2 (1864) (Cuba) ; Felder, Verh. Zool.
Bot Ges. Wien xiv. p. 302. n. 200 (1864) (partim) ; Edw., Trans. Amer. Ent. Soc. vi. p. 9. n. 2
( 692 )
(1877) (Florida, occasionally Cuba) ; Gerh., Macio-Lep. N. Amer. p. 25. n. 442 (1878) (Florida) ;
Strecker, Hull, ^folhs N. Amer. p. 68. n. G (1878) (Florida? ; Antilles ; jxirliiii).
Il,hkUil,'H simii. Hiibner. Ver-. heh. Srhm. p. 82. n. 8.38 (1818 ?) (parOm).
PapiUo celadon Lunas, Bee. Zool. (-'). iv. p. 130 (1852) (" Amer. Sept.") ; Doubl, Westw. & Hew.,
Gen. Dluni. Lep. ii. p. 529 (1852) ('• N. America ") ; Gray, Cut. L,p. /«.«. Jlril. .l/«.s-. i. p. 3 1 . n. 1 53
(1852) ("N. America") ; id., L!^l Lep. Ins. Ilril. .!/«<. i. p. 44. n. IHI (1851)) ("N. America ") ;
Lucas, in Sagra, fll.< F/.s-. Cuba vii. p. 204 (18.57) (Cuba); Morri?, Si/n. Lep. N. Am. p. 10. n. 14
(1862) (" California— Oregon ?," error); Weidem., Pme. Enl. Snc. Phihid. ii. p. 146 (ISin!)
(= sinoii = marerllimis. error) ; Felder, I.e. xiv. p. 303. n. 202 (1864) (partim) ; Kirby, Cut.
/j;«ni. ifyj. p. 557. n. 268 (1871) ("Antilles"); Oberth., Et. il'Enl. iv. p. 65. n. 173 (1»80)
(Cuba) ; Gundl., Papilio i. p. 113 (1881) (Cuba) ; id., Conlr. Ent. Cuhana p. 125 (1881)(Cuba);
Eimer, Artb. Venv. Schmetl. p. 182. t. 3. fig. 10 (1889) (Cuba); id., (Mlwgen. p. 44. 131.
305 (1897) (Cuba).
rapilio serion (!), Edwards, Canad. Ent. xiv. p. 120 (1882) (Cuba, perhaps also Florida).
Ijihirlides celadim, Kirby, in Allen's Nat. Libr., Lep. Butt. ii. p. 274. t. 68. fig. 2. (1896) (Cuba ;
" Jamaica," errore).
c? f . Though in general aspect not unlike J'. marceUinus, there are very
important diiferences, of which we mention the following : On the forewing it is
the third and fourth pale green cell-bands (instead of the fourth and fifth) which
are continuous with the discal band ; these two cell-bands narrowing costad,
separated by a black baud which usually extends to M ; a pale green cell-band
close to apex of cell, widely separate from the discal baud ; fifth submargiual spot
enlarged and, like the second, somewhat more proximal than the others. Red line
of hindwing below thin, bordered .with black on both sides, the red scaling not
extending beyond cell, often restricted to costal region, the line and its bonlers
continued to the brown distal area as a brown band.
Neuration : Cell of hindwing broader ai)ically than in P. marceUinus, veins R-,
W and M' being less close together and the apical angle less acute.
Genitalia : S. Tenth tergite long, narrow, slightly compressed, rodlike in dorsal
aspect, a little curved downwards, tip acute, with a very feeble notch at each side
indicating the lateral lobes of the allied species. Apical lobe of harpe much larger
than in P. marcellimis ; ventral process absent ; dorso-ventral ridge not ending
dorsally in a tooth or process. ?. Within vaginal cavity, at the proximal side
of the orifice, a tubercle which is mesially channelled ; at each side of the vaginal
orifice, but a little farther back, a deep impression, the two grooves being separated
in consequence of the mesial portion of the segment being non-depressed ; there
are some long bristles on this mesial part, doubtless of sensory function. Anal
segment with some short stout spines.
Early stages and food-plant not known.
Fresh specimens are pale bluish green, which colour fades, by exposure, into
greenish white or yellowish green.
Hah. Cuba ; probably also in Florida.
There is no authentic record of this insect from Florida ; but the species may
be expected to occur in southern Florida, like so many Cuban insects.
In the Tring Museum 8 c?c?, 7 ? ?, from : Hohjuin and Gibara, January and
April— May 1904 (Tollin) ; Cotorro.
141. Papilio zonaria Bntl. (1869).
Aubenton, rianch. Enlum. i. t. 18. fig. 1. 2 (1765).
Papilio Eques Achivus sinnn Fabricius {non Poda, 1761), S,/sl. Enl. p. 452. D. 39 (1775) (partim ;
" India") ; id., Spec. Ins. ii. p. 15. n. 59 (1781) (partim) ; Goeze, Enl. Beijlr. iii. 1. p. 72. n. 7
(1779) (parlim) ; Fabr., Mant. Ins. ii. p. 8. u. 67 (1787) (partim) ; Jabl. & Herbst, Naturs.
( C93 )
ScJim. iii. p. 159. n. 101. t. 44. fig. fi (1788) (ixirtim. ; fig. is copy of Aubent.'s fig. 2) ; Gmelin,
Syst. Nal. i. 6. p. 2-241. n. 329 (1790) {partim ; "India") ; Fabr., Ent. SijhI. iii. 1. p. 26. n. 75
(1793) (jMirtiiii ; "India").
Papilio sinon, Godarfc, E/w. MM. ix. p. 53. n. 80 (1819) {partim) ; Boisd. & Lee, Uht. Gen. Lip.
Amir. Sept. p. 11. t. 3. fi^. 1. 2 (1833) (partim; " Florida," error ??) : Boisd., Spec (li„.
i. p. 260. n. 83 (183G) (partim) ; Doubl., in Westw., Arc. Ent. i. p. 60 (1845) (not the same as
Drury's^«-o(es.); Gray, Cat. Lep. /«.«. Brit. Mu.'t. i. p. 32. n. 151 (1852) (" N. Amer. & Jamaici");
id., List Lep. Ins. Brit. Miis. i. p. 43. n. 159 (1856) (St. Domingo ; synon. partim) ; Lucas, iu
Sagra, Hist. Fis. Cuba vii. p. 204 (1857) (" Cuba, Jamaica, Florida," error) ; Menctr., Enum.
Corp. Aii-m. Mus. Petr., Lep. i. p. 3. n. 45 (18'i7) (jiarlim) ; Morri«, Si/ii. Lep. N. Am. p. 9. u. 13
(1862) (synoD. 7;aw;»( ; "Southern States"); Felder, Verh. Zoot. Bot. Ges. Wien xiv. p. 302.
n. 200 (1864) (partim) ; Oberth., Et. d'Eiit. iv. p. 65. n. 171 (1880) (S. Domingo).
Iphidides simin, Hiibner, Tci-s. hek. Schm. p. 82. n. 838 (1818?) (partim) ; Dyar, Bull. U. St. A\it.
Mus. Iii. p. 2. n. 6 (1902) (jyartim).
Papilio zonaria Butler, Cat. Diurii. Lep. descr. Fabr. p. 240. n. 24 (1869) (San Domingo) ; id.,
Ent. Mo. Mag. v. p. 271. n. 3 (1869) (S. Domingo) ; Kirby, Cat. Diurn. Lep. p. 557. n. 207
(1871) (partim); Gerh., Macro-Lep. N. Amer. p. 25. n. 441 (1878) ("Union!") ; Eimer,
Artb. Verwandtsrh. Sehmett. p. 186 (1889) ("Jamaica," errore).
Papilio celadon, Weidemeyer, Proc. Ent. Soc. Philad. ii. p. 146 (1863) ( = siuoii = marcellinus, errore).
Papilio sonaria = serion (!), Edwards, Camul. Ent. xiv. p. 120 (1882) (" Cuba," errore).
c??. Closely related to P. marcellinus; in aspect similar to P. philokms in
conseqnence of the reductiou in width of the pale bands. On forewing the fourth
and fifth pale green cell-bands continuous with the narrow discal band, third
cell-baud thin, as in P. marcelUmis, but a little more distal in position, fourth
and fifth bands much thinner than iu P. marcellinus, much more distal and
comjiletel}- separate from one anotiier ; a small pale dot distallj' of short si.xth
baud. Red line of underside of hindwing bordered with black on both sides,
reaching to brown distal area or nearly.
Scent-organ and neuration essentially as in P. marcellinus.
Genitalia as in P. marcellinus, but apical lobe of harpe longer and more
evenly rounded.
Early stages not known.
Ilab. Haiti.
The early records of this insect from Florida, Cuba, or Jamaica are not
authentic; marcellinus and cel'idon were doubtless confounded with zonaria. All
the specimens oi zonaria which we have seen were from the island of San Domingo
(Haiti).
If a representative of this group occurs on Porto Uico, it is most likely zonaria
or a form closely allied to it.
In the Tring Museum I S.
In coll. Oberthiir 5 cJc?, 1 ? ; also a small series in coll. F. D. Godmau.
142. Papilio philolaus Boisd. (1836).
Papilio philolava Boisduval, Spec. Gen. Lep. i. p. 256. n. 80 (1S36) (Mexico) ; Doubl., List Lep. Ins.
Brit. Mhs. i. p. 8 (1845) (Oaxaca) ; id., Westw. & Hew., Gen. Diurn. Lep. i. p. 15. n. 129 (1816)
(Mexico) ; Gray, Cat. Lep. Ins. Brit. .Uus. i. Pap. p. 33. n. 154 (1852) (Mexico) ; id., List Lep.
Ins. Brit. Mies. i. p. 44. n. 1G2 (1856) (Mexico ; Nicaragua) ; Mum'tr., Enum. Corp. Anim. Miis.
Petr., Lep. i. Suppl. p. 68. n. 1121 (1857) (Mexico) ; id.. I.e. iii. p. 110. n. 1121. t. 7. fig. 1 (1863)
("Amer. Sept.," errort-) ; Felder, Verh. Zool. Bol. Ges. Wien xiv. p. 303. n. 201 (1804) (Mexico ;
Nicaragua ; " Amer. Sept.," errore) ; Boisd., Cons. Lip. Gnat. p. 6 (1870) (Honduras ; Mexico);
Kirby, Cut. Diurn. Lip. p. 557. n. 266 (1871) (Amer. centr. ; "Amer. bor.," errore) ; Gerh.,
Macro-Lep. N. Amer. p. 25. n. 440 (1878) ; ObtTtb., Et. d'Ent. iv. p. 65. n. 174 (1880) (Mexico);
Staud., Exot. Tayf. i. p. 18. t. 12. J (1884) (Guatemala ; Mexico) ; Eimer, Artb. ['erwandtsch _
( 694 )
Schmelt. p. -.'lO. t. 4. fig. 1. 7 (18H'J) ; Godm. i- Salv., Biol. Cenlr. Aiurr.. Rhqi. p. 'liO. n. 4!).
t. 68. fig. 12. harpe (18'.)0) (Me.\ico : Guatemala; Brit. Honduras; Honduras; Nicaragua):
Eimer, Orllwflen. p. 401) (I8'.I7) (" Nord Amer.," errore ; Mittel Amer.).
Pujiilio xuitticles, Rogenhofer, in Staud., Ej-ul. Tufjf. i. p. 305 (18HH).
riijiiliojihilolaiis aja.c Eimer (mn Linm', 1758), Aiib. Verwiimlhili. Schmell. p. '212. t. 4. fig. 1 (1889)
(Me.xico).
Papilio philolam nlgrescens id. (non id., 1880, jioilnHriKs iiii/resiens). I.e. p. 213 (1889) (Hondura.«i).
PapilUi philolaiis tilger id.. I.e. 214 (1889) (Honduras).
Papilio philolaus ah. feliris Fruhstorfer, Sue. Enl. p. 2,1 (1904) (Honduras).
(? ?. Aiitetiua bl;u-k, more or less feebly tawny at base. Tibiae and tarsi jiale
green, tarsal segments slightly ochraceons at ajjcx ; mid- and hindtibial s])nrs about
as long as the tibia is broad, the sj)nrs being sliorter than in F. inarcellm^ ; claw
also shorter than in that species. Seventh jiale band of forewing represented by a
spot SC — SC"'''', there being often some pale scales costally of this spot ; bindwing
abore with two red s]iots between M' and abdominal margin, black median band
running across apex of cell, being contignons with the oblique cross-vein D-, very
seldom slightly separate from this vein, this band not interrupted, always extending
to the black distal area.
On underside the red median line of hindwing is bordered with black on both
sides, at least costally ; it is undulate costally and is always ccmtignous with cross-
vein D^. The praecostal vein stands close to the apex of the basal cellule; the
ajiical angle of the cell of the hindwing is acute, D- being very oblique ; D' is short,
being about one-fourth the length of I)- ; D-' and D'' are together shorter than D-.
The scales of the upperside, apart from the anal area and the tail, are nearly all
entire.
Scent-orgau : scent-scales short, with numerous long filaments, the shape of
the scales and the number of filaments being variable.
Genitalia : S . Dorsal ridge of harpe reaching ventral edge nearly in middle-
?. A deeply sinuate lobe in front of vaginal cavity, the lobe plicate ; laterally
of cavity a larger lobe of which the edge is simply convex.
Early stages not known.
The species does not vary geographically, but there is considerable imlividuiil
variability, especially in the width of the bands. The red markings of the hindwing
are occasionally pinkish yellow (a male from San Pedro Sula in Mus. Tring).
The third pale line of the forewing is often washed over with black, the line being
sometimes distinct only near the costal margin ; the second pale line is also
reduced in length in some specimens. The sixth pale band is rarely so much
prolonged as to nearly extend to the discal band. The latter is usually very much
narrower than the black band situated between it arid the submarginal row of
greenish spots, but there occur also specimens in which the greenish white band is
wider between M' and hinder margin than the blai'k band situated di.stally of it.
The submarginal spots of both the fore- and hindwing are largest in the specimens
with wide discal baud, the last of the forewing and the first and last of the hindwing
being often vestigial in individuals with narrow pale bands. The specimens with
broad pale bands are a little more short-winged than the darker specimens, the
forewing being on the whole also less falcate. As these differences are somewhat
similar to those observed between the seasonal varieties of the North American
P. marcellii.i, it is possible that the dark and the pale specimens of /'. philolaus
belong to difi'ereut broods ; but no observations have been made towards this ])oint.
The pale band along tlie abdominal margin of the underside of the hindwing is
( 695 )
often somewhat washed with red, reminding one of the coi'resi)onding red line of
r. asius.
The female is dichromatic.
a'. ?-f. /^/«7o/a«s Boisd. (1836) is similar to the male, differing especially in
the underside being paler. The vestige of a pale postdisoal band on the nnder-
surface of the forewing is on the whole more distinctlj' marked than in the male.
b'. ?-f. nii/er Eimer (1889) (=i/el/cis Fruhst., I.e.) has lost all the pale bands,
the row of snbmarginal spots being however more or less distinct. The red
markings of the hindwing are also preserved. The black surface of the wings has
not been attained by a widening of the black bands, but by the pale bands being
washed over with black, these bands remaining as faint shadows. It is the same
development which we observe in the African P. illi/ris and kirbi/i, and the North
American P. glaucus glaucus ¥ -f. glaucus.
Hah. of P. philolaus : Mexico to Nicaragna and Honduras.
In the Tring Mnsenm 55 cJcJ, 7 ? ?, from: Sangolica, June 1897, Espinal,
June 1896, Vera Cruz (W. Schaus) ; Guerrero (0. T. Baron) ; Guatemala (Salvin);
San Pedro Sula, Honduras.
Men^tries, when describing and figuring this species, I.e., gave North America
as the country where Motschoulsky had obtained the specimens, and since then
several authors (Felder, Kirby, Eimer) have included " Amer. Sept." in the range
of the species. However, the sjjecies does not occur north of Mexico. In the list
of 1857 Men^tri6s correctly gave Mexico as habitat.
143. Papilio xanticles Bates (1863).
Papilio xaiiticlc.'i Bates, Proc. Zool. Soe. Land. p. 241. n. 1. t. 29. fig. 3 (1863) (Panama) ; Pelder,
Verli. Zool. Bat. Ges. Wien xiv. p. 302. n. 1<J9 (1861) ("Guatemala," false); Kirby, Cat.
Diurn. Lei>. p. 557. n. 265 (1871) (Panama) ; Oberth., El. d'Eitl. iv. p. 67. u. 186 (1880)
(Panama); Godm. & Salv., Trans. Eid. Soc. Loud. p. 126. n. 234 (1880) (Manaure, .Sta. Marta);
Eimer, Artb. VerwamUscli. Schmett. p. 178. fig. U (1889) ; Godm. & Salv., Biol. CtiUr. Ainer.,
Rhop. p. 221. n. 50. t. 68. fig. 10. ? , 11. cJ (1890) (Lion Hill, Panama ; Colombia).
Papilio plaesiolaus Staudinger, Emt. Tagf. i. p. 17 (1884) (Sta. Martha) ; Eimer, he. p. 182 (1889)
(var. of arcf.'iilaii.^, errore).
<S ? . Pale bands of upperside of wings huffish straw-yellow ; the cell-bands
of forewing rather wider than in P. philolaus, sixth pale band extending to R-
where it joins the discal band, the latter continued costad to S(/'', the single spot
SC" — SC* of P. philolaus being replaced in P. xanticles by a band which is con-
tinuous with the broad discal band, being the direct ])rolongation of the same.
Black subbasal band of hindwing much thinner than in P. philolaus ; black median
band widely interrupted ; red anal spots smaller than in P. jthilolaus; snbmarginal
spots larger.
On underside, position of red line of hindwing as in P. philolaus, central
portion of line thin. Pale streaks on abdomen broader than in P. philolaus.
The species does not vary much. The ? , however, appears to be dimorphic as
in philolaus :
a'. ?-f. xanticles Bates (1863) similar to the <S . This form is not known,
but doubtless exists.
b'. ?-f. snheha nov., wings black, except a row of yellowish snbmarginal spots
and two red anal spots.
Though somewhat resembling in colour P. arcesilaus, of which it has generally
been considered a near ally, P. xanticles is in fact hardly more than a southern
( 696 )
form of P. philolaus. There is no strncturul differpiice between P. rrniticles and
jiliilolaus, xanticles having all the jipiuliarities in tlie structure of the scent-
organ, the genitalia and legs which we have mentioned nnder P. ]>liilolaii.<<. In
pattern the two insects are well separated. However, we must bear in mind that,
/'. p/iilolaiis being known to occur as far south as Nicaragua, and xanticles having
been found only in Colombia and Panama, there is a geographical gap between the
two Pajiilios, where possible an intermediate form exists which has as yet escaped
observation.
The fact that the extended-black P. philolaus and the much less extended-
black P. xanticles are two closely related species, if they are not geographical
varieties of one species, appears to us to upset one of those "laws " of development
established on insufficient evidence by Eimer and accepted by others. In Ortho-
genesis, p. 401, Eimer comparing the characters of philolaus with those of the
various seasonal forms of marcellus {ajax, auct. non Linn6) comes to the conclusion
that the black bands of philolaus are strongly developed because philolaus is a
more southern insect than marcellus. However, if high temperature and moistnre,
as Eimer says, were the real causes which have turned the less extended-black
ancestral philolaus into the extended-black present-day philolaus, then the ally
of philolaus which lives in a hotter and more moist climate than philolaus itself,
ought to be more extended-black than philolaus. We find, on the contrary, that
P. xanticles from the coast of Panama and the north coast of Colombia is far less
black than philolaus from Nicaragua and the countries northwards, inclusive of
Mexico, and even than the North American marcellus.
Eimer makes another mistake when talking of the lines of development of
P. philolaus and P. marcellus. He calls tlie black form of philolaus — Eimer did
not know that this black form is restricted to the female sex — as being arrived at by
extension of the black bands. That is not correct. The pale bands have not become
narrower and narrower until they finally disappeared, but the i)ale bands have
become suppressed by the scaling turning black. This assumption of black colour
on the part of the scales of the pale bands is a new kind of development (see
P. ph. ?-f. niger), the black wing of these females being not at all the final result
of Orthogenesis, i.e. of a gradual widening of the black bauds.
The Papilio plaesiolaus Stand. (liSS4), which is the same as xanticles, is treated
by Eimer, I.e., as being a variety of arcesilaus, a very different species. However,
Eimer knew xanticles only from Bates's figure and description, and plaesiolaus
from Staudinger's description.
In tlie Tring Museum 8 cJ cJ from Panama.
In coll. F. D. Godman a series of males and the only known female. One of
the males, from Manaure, S. Martha, has the yellow discal areas of both wings and
the snbmarginal spots of the forewing enlarged.
144. Papilio oberthueri spec. nov. (PI. VI. fig. 25).
i. Body, antenna, and legs essentially as in /'. jthilolaus.
Wings, upperside : less deej) black than in 1'. jihiloluu.s, (lie scales nearly all
mii- or bisinuiite. Forewing seniitran.sparent distally, jr.ile bands white, proximal
ones greenish, snbmarginal spots also greenish ; these bands broader than in
P. philolaus ; second pale band about two-thirds the width of the black band
situated distally of it, sixth band extending to R-, being separated from the white
discal area only by the black vein R-, costally a little more distal in position than in
( 697 )
philolaus ; spot SC — SC* as in pkilolaus, white discal area about half as wide again
at RP as the bhxck distal area ; third black cell-baud extending a very little beyond
M, there being only a small black dot at the base of cellule M' — M' ; upper sub-
marginal spots larger than in philolaus, the posterior ones vestigial. Hiudwing
narrower than in pkilolaus ; black median band stopping at cell, not reaching across
apex of cell as is the case in pkilolaus ; black distal area a little paler than in
pkilolaus, narrower, veins R^, R', and M' thinly black ; upper submarginal spot
vestigial, the others more or less washed with black.
Underside : black bands somewhat paler than in pkilolaus. Forewing : last
submarginal spot more or less vestigial ; black dot at base of cellule M' — M^ very
small or absent. Hindwing : black snbbasal band narrower than in pkilolaus ;
red line not undulate, crossing cell between M' and R^ or at M', the posterior
portion of the line more or less vestigial, 2 or 3 mm. distant from apex of cell ; the
line bordered with black distally and very feebly or not at all proximally ; two red
anal spots as in pkilolaus ; submarginal spots less distinct than in pkilolaus,
especially the upper ones.
Neuration : cell of hindwing much less acute at apex than in pkilolaus, M' less
close to R^, and D" less oblique ; praecostal spur more proximal.
Scent-organ : scales longer than in philolaus, the lateral filaments shorter.
Genitalia similar to those of P. philolaus.
Hab. San Pedro Sula, Honduras (VVittkugel) ; 3 cJ c? in coll. Charles Oberthlir.
145. Papilio arcesilaus Lucas (1852).
Papilln arcesilaus Lucas, Rer. Zmil. p. 131. t. 10. fig. 2 (1852) (" Ami'r. du Nord " errnre ; — coll.
Oberthilr) ; Doubl, Westw. & Hew., Gen. Diurn. Lep. ii. p. 52y (1852) ("North America,"
errore) ; Gray, Cat. Lep. Ins. Brit. Miis. i. p. 33. n. 15ri (1852) ("N. America") ; id., List Lep.
Jns. Brit. Mus. i. Pap. p. 45. n. 164 (1856) ("N. America ") ; Morris, Syn. Lep. N. Am. p. 11.
n. 15 (18G2) (" California— Oregon ? ", errore); Weidem., Proi: Ent. Soc. PhUatl. ii. p. 146
(18G.3) (" U.S.A." errore) ; Felder, Verh. Zool. Bot. Ges. Wien xiv. p. .'i02. n. 197 (1864) (Caracas);
id., Reise Norara, Lep. p. GO. sub n. 45 (1865) (Venezuela) ; Kirby, Cat. Diurn. Lep. p. 557.
n. 2G4 (1871) (Venezuela) ; Oberth., Et. (VEnt. iv. p. G5. n. 175 (1880) (Colombia ; Valera,
Venezuela); Staud., E.mt. Taijf. i. p. 16. t. 11. (J (1884) (Valera, Venezuela; ?, Caracas);
Eimer, Artb. Venrandtsch. Schmett. p. 179. t. 3. fig. 9 (1880) (Venezuela ; Bogota) ; Hahnel,
Iris iii. p. 200. 203 (1890) (VaMra) ; Eimer, Orthogen. p. 44 (1897).
Papiliu anaxilans Felder, Verh. Zont. But. Ges. Wien xiv. p. 302. u. 198 (1864) (Bogota) ; Godm.
& Salv., Trans. Ent. Soc. Land. p. 12G. n. 233 (1880) (Sta. Marta) ; Staud., E.rot. Taijf. i. p. 17
(1884) (Antioquia, Colombia).
Papilio arcesilaus var. a. P. auaxilaus, Kirby, I.e. (1871) (Bogota).
Papilio arcesilaus-anaxilaus, Eimer, Artb. Verwandtxh. Schmett. p. 181 (1889) (Colombia).
(J ? . Antenna black, beneath slightly tawny proximally ; scaling of upperside
black. Tibiae and tarsi pale green, not scaled, or only a very few scales present ;
mid- and hindtibial spurs as long as the tibia is broad, outer spnr a little shorter
than inner.
Wings, upperside: scales nearly all entire, apart from tail and anal area.
Forewing : six greenish white bands and a row of submarginal spots, sixth band
short, reaching only to R' or R', being separate from the greenish white disciil ;ir(;i.
Hindwing : black median line marked only costally, sometimes vestigial.
Red median line of underside of iiindwing reachirjg to blackish brown distal
band, contiguous with cross-veins D' and D-, there being no white spot in apex of
cell outside the red line or only a trace of such a spot ; the line bordered with black
on both sides costally, especially on the proximal side.
( 698 )
Scent-organ : scales similar to those of P. agesilaus.
Genitalia : <?. Harpe reserablinj!; that of P. philolatts, dentition slightly
different, proximal edge of central process subdeutate, ventral process vestigial.
The three lobes of tenth tergite a little longer. ? not dissected.
Early stages not known.
The species does not vary mnch. The fourth black bandof the forewing reaches
usually to the median vein, but is often narrowed behind or abbreviated. The
submarginal spots of the forewing vary in size and distinctness, the whole series
being sometimes more or less washed with black. In a male from Venezuela, in
coll. Godman, the cell-bands of the forewing are washed over with black. The
single Colombian specimen in the Felder collection differs rather obviously from
the only Venezuelan individual which Dr. Felder had for comparison when he
described the former specimen as belonging to another species, which he named
anaxila.us. The differences are, however, not constant. The small series of
Colombian specimens which we have seen proves that the individuals vary inter se.
Hab. Venezuela and Colombia.
In the Tring Museum Q SS from : V^alera ; " Venezuela" (Moritz) ; Bogota.
140. Papilio epidaus Doubl. (1840).
Papilio epidaus Doubleday, ia Doubleday, Westw. & Hew., Gen. Diurn. L'p. i. p. lu. n. 138. t. 3.
fig. 1 (1846) (Mexico ; Honduras).
c? ? . Antenna black, occasionally brown at the apex of the segments ; scaling
black, usually fallen off'. Tibae and tarsi pale green ; mid- and hindtibial spurs
about as long as the tibia is broad, inner one a little longer than outer.
Scaling of wings peculiar. Fvrevfiug, !/jipersic/e : black bands densely scaled,
scales dentate, those of marginal band somewhat narrower than the scales of the
other bands ; the white scales all very narrow, sinuate, distant from each other,
therefore the wing more or less transparent, the hinder portion, however, appearing
o])aque owing to the denser white scaling of the undersuriiice shining through.
If the wing is looked at in a slanting position, the eye being between light and
specimen, a broad elongate-triangular band situated between distal margin and cell
appears transparent, while the rest of the wing is more or less white, apart from
the black liands ; this transparent band is covered with minute black scales on the
upperside, being quite naked on the underside. The homology of this band is easy
to perceive, the band corresponding to the postdiscal band of P. agesilaus, which
is a double one ; the pale line which divides this postdiscal band of /■■. agesilaus
(most distinctly in P. agesilaus aiitosilaus) is represented in P. epidaus by a white
costal si)ot. The external edge of the transparent band of P. epidaus is usually
more densely scaled than the centre, at least at the costal margin, forming a black
proximal border to the white submarginal band, the border extending often down
to M-. The jiroximal portion of the postdiscal band of P. agesilaus is in P. epidaus
represented by a black costal spot situated proximally of the subcostal fork, aad
by a black baud which runs from the lower angle of the cell towards the hinder
angle of the wing, being a direct continuation of the discocellnlar band. The
subapical cell-band is represented by a costal spot in must specimens, there being
from this spot across the cell a faint band of dispersed minute black scales ;
occasionally the band is distinct as far as middle of cell. Most of the scales in
the costal and central area of the hindwing are entire.
( 699 )
The transparent spaces of the underside of the forewing are practically devoid
of scales ; tlie white scales of the posterior area are entire. The scales of the
liiiidwiug are dentate, except at the abdominal margin and between the subbasal
and median bands.
Markings very characteristic. Second band of forewing reaching hinder margin
beyond middle, discocellnlar band continued to M' or hinder angle, as explained
above. Abdominal edge of hiudwiug black ; subbasal band heavy, continued to
anal spot ; median band complete or posteriorly vestigial on upper-side, bordering
cell, on underside the band bordered with black proximally down to anal spots,
its distal black border either being restricted to the costal region, or being thinner
than the proximal border, at least from SG- to 11'^
Neuration : lower angle of cell of forewing obtuse ; cell of hiudwing broad,
widest at origin of nervule SO" ; D' and D" variable, sometimes nearly the same in
length, sometimes D' almost twice the length of D- ; D^ always very short ; D' about
half or two-thirds the length of \f' ; praecostal spur elbowed, not evenly curved.
Scent-organ : fold small ; scent-scales nearly as in P. agcsilaus.
Genitalia : c?. Tenth tergite trilobate as in P. agesilaus ; harpe strongly
elevate where the dorso-ventral ridge meets the ventral edge, apical lobe broadly
rounded, short. ? . A small, feebly chitinised tubercule at the jiroximal side of
the vaginal orifice ; behind the orifice, laterally on each side, a deep large groove,
the walls of which are rather strongly chitinised ; anal segment with numerous very
short spines.
Early stages not known.
Hab. Central America : Mexico to Honduras ; Nicaragua (according to
M6ndtrids).
Butler and Druce record it from Costa Rica. The record is certainly erroneous.
This specimen is in the Godman collection. It agrees with P. epidaus epidam, which
would hardly be the case if the specimen came from Costa Rica. The record from
Nicaragua, though requiring confirmation, may be correct, since Nicaragua belongs
to the northern faunistic district of Central America.
Three subspecies.
a. P. epidaus epidaus Donbl. (1846).
Papilio epidaus Doubleday, in Doubl., Westw. & Hew., I.e. p. 15. n. 138. t. 3. fig. 1 (1840) (Mexico ;
Honduras) ; id., List Lep. Lis. Brit. Mus. i. A]>p. p. 2 (1848) (Honduras; Yucatan); Gray,
Cat. Lrp. Lis. Brit. Mus. i. p. 34. n. 161 (1852) (Honduras ; Yucatan, ace. to Becker); id., List
Lcp. Ills. Brit. Mus. i. p. 4li. n. 169 (185G) (Nicaragua : Honduras ; — ? Becker coll.) ; Mene'tr.,
Eniim. Cmp. Auim. Mus. Petrop., Lip. i p. 3. n. 50 (1857) (Nicaragua); Reak., Proc. Ent. Soc.
P/iilud. ii. p. 135. n. 2 (1803) (Honduras ; good description) ; Weidem., ibid. p. 147 (1863)
(Mexico ; Central America) ; Felder, Vcrli. Zool. But. Ges. Wicit xiv. p. 302. ii. 196 (1864)
(Mexico ; Honduras) ; Boisd., C'niis. Lip. Guatem. p. 6. (1870) (Mexico ; Honduras ; Nicaragua;
— " Colombia," error) ; Kirby, Cat. Ditirn. Lrp. p. 557. n. ilii (1871) (Amer. centr.) ; Butl. &
Druce, Pi'oc. Zuul. Soc. Land. p. 305. n. 370 (1874) (" Costa Rica," errore) ; Oberth., Et. d'Ent.
iv. p. 65. n. 177 (18S0) (Mexico) ; Eimer, Arth. Verwamltsch. Schiii. p. 51. 111. t. 1. fig. 7 (1889)
(Central Amer.) ; Godra. & Salv., Biul. Ceutr. Amer., Rhnp. p. 221. n. 51. t. 68. fig. 5. genit.
(1890) (Mexico : Vera Cruz, Yucatan ; Brit. Honduras ; Guatemala ; Honduras ; Nicaragua. —
"Costa Kica " error loci; "San Bias'' alia subsp.) ; Eimer, Orthogen. p. 47. 109. 311.
397 (1897).
6 ? . Upperside : second black baud of forewing half the width or less of the
interspace between it and third band ; filtli band not joining the marginal band
behind, stopping short at M" or at least not reaching this vein. Median band of
45
( 700 )
liiiiJwiiig- vestigial lii'vond cell or verj' thin ; red anal spots LonkM-L-J with white
ill IVoiit ; black distal band not broader between tS(_'- and 1{' than the greenish white
snbmarginal hall'inoon.
In one of our females (Hondnras) the fifth band of the forcwing reaches well
beyond SM-, nearly touching the marginal band.
Ilab. Eastern Mexico: Vera Cruz, Yncatan ; Guatemala; British Honduras;
Honduras ; " Nicaragna " (accurding to Menetries).
The black snbmarginal line of the forewing is often clearly marked down to M-,
while in other specimens it is restricted to a costal spot.
In the Tring Museum lo SS, 4 ? ?, from : Esjunal, Vera Cruz, June 1890
(W. Schaus) ; Guatemala (Salvin) ; San Pedro Siila, Honduras.
b. P. epidaus tepicus subsp. nov.
rapilio qmhtiis Godman & Salv., JS/o/. Ceiitr. Amer., Rhop. p. 221. n. 51 (1890) (p<(r/!m ; San Bias).
J. Wings. Upperside. — Forewing : first and second black bands broader than
in ep. cjiidaus, fifth band continued to hinder angle where it joins the marginal
band. Hindwing longer than in ep. cpidaus and costally narrower, median band
broad in front, more distinct beyond cell than in cp. epidaiis ; snbbasal band thinner
from cell backwards than in front ; white border of red spot M' — M- thinner;
black patch R' — M' larger, greenish white submarginal spots larger; tail more
extended silvery white.
Underside : the same difterences as above, but white border of red spot M' — M-
of hindwing as large as in ep. epidaus.
ITah. West Mexico : Jalisco.
In the Tring Museum 1 S from Jalisco ; several specimens in coll. F. D.
Godman from San Bias.
c. P. epidaus fenochionis Godm. & Salv. (1808).
Papiliofeiinchiouh Godman !.<• Salvin, Ann. Mag. N. H. (4). ii. p. 1.^0 (1868) (Oaxaca) ; iid., Biol.
Ceiilr. Amer., Rhop. p. 222. n. 52. t. G8. fig. 13. 14. S (180O) (Oaxaca).
S ?. Fifth liand of forewing extended to hinder angle. Snbbasal and median
bands of hindwing broad above and below, median band always reaching to the black
distal area, which is mucli more extended than in the other two subspecies ; red
anal spots above without distinct white border, or the border very thin, the red spots
being larger than in the other forms. Width of black bands of hindwing very
variable ; intersj)ace between subbasal band and abdominal border often almost
completely filled in with black-brown scaling.
Ilab. South-Western Mexico : Oaxaca ; Guerrero.
In the Tring Museum 64 c?<?, 2 ? ?, from : Guerrero (0. T. Baron) ; Salina
Cruz, Tehuantepec, July l'.»04 (A. Hall) ; Oaxaca, July 1890 (W. Schans).
147. Papilio bellerophon P.ilin. (1823).
Papilio bellernjiliijii Dalman, .l/»i/. ICnt. p. ST. n. 1 (1H2:5) ; Boisd., Sjiec. Gen. Lip. i. p. 204. n. 87.
(IS.Ifi) (Bahia ; Pernambuco) ; Doubl., Westw. & Hew., O'ch. DUmi. Lep. i. p. 15. n. i::!5 (1S4C) ;
Doubl., List Lcp. [lis. Ihit. .Mux. i. Append, p. 3 (1848) (Brazil) ; Gray, Cut. Lep. Jus. i. Pup;
p. 33. n. 157 (1852) (Brazil) ; id., List Lep. J/is. Bril. .Uus. i. Paj}. p. 45. n. 105 (1850) (Brazil).
Menutr., Emtm. Corp. Ani7u. Mux. Petrop., Lep. i. p. 3. n. 48 (1857) (Brazil) ; Felder, Vcrli
Zool. Bot. Gcs. Wien .xiv. p. 301. n. 175 (1804) (Bras, austr.) ; Kirby, Cat. Diiirii. Lep. p. 565.
n. 245 (1871) (Brazil) ; Oberth., Et. d'En/. iv. p. 07. n. 184 (1880) (Brazil) ; Eimer, Artb.
(701 )
Vcrwmultscli. Sclnn. p. 53. 112. fig. N. t. i. fig. 12 (1889) ; Fickert, in Eimer, t.c. ii. p. r,2. fig. E
(\9>K)) (neuration).
Papilio coresilaus Godart, Enc. Mith. ix. Suppl. p. 810. n. 61-2 (1824).
I'rolesilaHK sioainsonius Swainson, Zool. Illnstr. ii. t. 104 (1833).
c? ? . Antenna black, scaling of upper.siJe lilack, usually fallen off. Black
Lairs of frons long. Tibiae ami tarsi pale green, scales preserved in fresh specimens
only ; inner tibial spnr longer than outer.
Wings yellowish white. Forewing semitransparcnt ; four black bands, one
across cell, stopping short in front of M-, a second on cross-veins, joined at lower
angle of cell to the third, which e.xtends from costal margin to hinder angle or close
to the angle, a fourth being marginal ; scales of yellowish white areas very narrow,
separate, sinuate, in posterior and basal areas alternately hair-shaped and triangnlar;
on underside as above, but scales large and entire in posterior area of wing.
Hindwing much denser scaled than forewing, scales entire in costal area, abdominal
and central areas from base to median band rather densely hairy ; black median
band thin, vestigial in front, slightly curved, touching apex of cell ; the band
complete on underside, centred with a more or less incomplete red line ; scales of
yellowish areas much narrower than on upperside, interspersed with fine hairs from
base to median band.
The female larger than male ; wings broader, distal margin of forewing more
convex, red line of median band of hindwing vestigial on upperside ; the median
band more or less dilated in centre of wing on distal side, reminding one of P. salcini.
Neuration : SC of forewing absent ; C ending farther distad tiian in other
species; PC of hindwing elbowed or nearly evenly curved; D^ of hindwing about
twice the length of D-, D' as long as or a little shorter than D'', angle of cell a little
less than 90°.
Scent-organ : fold woolly, scales before and behind SM^' entire, large, some
scales of this kind also beneath the wool, whicli consists of thin long hairs, widest
in middle, and of longer and thicker stiff hairs vfcich end in an abrupt point.
Genitalia : S . Tenth tergite long, compressed, trilobate at apex; dorso-ventral
ridge of harpe nearly continuous with dorsal edge of apical lobe, extending close to
the rounded apex of this lobe, ventral margin of harpe dentate, the harjie produced
proximally into a rounded lobe which reaches close to the ventral process, central
process strongly compressed, abruptly i^ointed. • ? not dissected.
Early stages not known.
IJnh. Brazil.
In the Tring Museum 1.3 c? (J, 1 ?, from: Minas Geraes, February 1901 (A.
Kennedy) ; Castro, Parana (E. D. Jones) ; Theresopolis, S. Catharina (J. Michaelis).
XV. Protesilaus Group.
The species of this group resemble one another so closely in pattern that many
authors luxve treated them as mere individual varieties of one single species, with
the exception of J', ((ijesilaus, the distinctness of which the more recent authors at
least have not doubted. We have studied this group carefully, the result being
that we recognise no less than nine distinct species (as opposed to geographical and
individual forms). These species are distinguished in the genitalia, in jiattern and,
at least partly, also in the scent-organ of the hindwing. Five of these species occur
bide by side over the greater part of tropical South America, while the others have
( 702 )
a more restricteil range, Brazil being inhabited by no less than eight ont of the nine
species, of the ninth so far only a few Ecuadorian specimens being known. Two of
these insects extend northwards to Mexico {F. wjesilaus and P. protesilaus), a
third species {glaucolaus) being found as far north as tlie isthmus of Panama.
The various species as they occur in the same district are in most cases easily
distinguislied from one another, but there is often hardly any difference in pattern
between the individuals of one species from one district and the individuals of
another species from another district, or two species are very different in one locality
and almost identical in pattern in another. This renders it practically impossible
to give a workable key to the species based on pattern only. Therefore we only
grouji the species together according to i)attern, and then give a key based on the
genitalia of the males, the females being so rare in this group that for want of
material we cannot take them into account in this key.
a. Inner edgeof black postdiscal band of hindwing, upperside,
quite straight down to M', the partitions R- — M^ of
this band not luuiform
A line of black spots in middle of upperside of hindwing . Species No. 153.
No black spots in middle of upperside of hindwing . . Species No. 152.
b. Inner edge of black postdiscal band of hindwing incised
on veins R-, R' and M', the partitions R- — M' of the
band being more or less Inniform
Ad- and submarginal interspaces of hindwing yellow . Species Nos. 154
and 15G.
Ad- and submarginal interspaces of hindwing white or
slightly huffish Species Nos.149,
150, 151, 155.
c. Red lino on underside of hindwing bordered with black
on outerside, not on innerside as it is in all the other
species Species No. 148.
Key based on the c?-genitalia and scent-organ ; Sjiecies No. 148 not
included, as it can easily be recognised by the pattern. See fig. 1, 2, and 3
on p. 708.
a. Dorso-apical ridge of harpe only slightly deflexed, being
vertical on the plane of the clasper; the ridge con-
tinuous from dorsal edge of clasper to apex of harpe . Species No. 149.
Dorso-apical ridge of harpe strongly deflexed, lying
almost flat on the main body of the harpe ... b.
b. This ridge dilated into a rounded lobe or a large tooth . c.
The ridge very narrow, hardly at all widened ... d.
c. The ridge rounded-dilated, not produced into a prominent
tooth Species No. 150.*
The ridge dilated into a large trianguhir tuotii . . . Species No. 151.
(I. Hair-.scales of scent-organ very thin .... e.
Hair-scales of scent-organ shorter anil l)roador than in
1'. jirotcsilaus and the other species excejit ai/ctsihiiis . Si)ecies No. 156.
e. Ventral process of harpe reaching to ventral edge of
clasper /.
• See also the subspecies a, b, and c of Species No. 151.
( 703 )
Ventral process of liarpe not reaching to ventral edge of
clasper g.
f. Doflexed edge of dorso-apical ridjie of Iiarpe uon-deutate . Species No. 154.
Defiexed edge of dorso-apical ridge of barpe serrate
proximal]}' Species No. 152.
(j. Central process of harpe short, broad, strongly dentate . Species No. 155.
Central process of harpe slender, its teeth vestigial . . Species No. 153.
148. Papilio agesilaus Gner. (1835).
Pcipilio Eques Achivus protesilaus, Esper {nnn Linno, 1758, err. det.), Ansl. Schmrll. p. 2U7. n. 95.
t. 52. fig. 1 (180.3?) (iMu-lim).
Papilio protesilaus, Godart, Enc. Metli. ix. p. 50. n. 73 (1819) (partitn).
Papilio agesilam Guerin & Percheron, Gen. Ins. Lip. t. 1. fig. 1 (18.35) (R. Magdalena, Colombia) ;
Boisd., Spec. Gin. i. p. 2G3. n. 86 (1836) (Mexico ; Colombia) ; Doubl., Westw. & Hew., Gen.
Diiini. Le/i. i. p. 15. n. 136 (1846) (Mexico; Colombia) ; Eimer, Arih. Verwamllsch. Sclim.
p- 98 (1889) ; id., Orthogen. pp. 44. 47. 111. 139. 217. 497 (1897).
(? ? . Antenna hrownish black, scaled black on npperside in fresh specimens.
Tibiae and tarsi pale green, scaled white, scales easily falling off, tips of tarsn!
segments ochraceous ; mid- and hindtibial spurs a little shorter than the tiliia
is briiad.
Wings greenish white ; a few scales on disc of forewing, iippcrs/df, the scales
from SC- of hiiidwing to costal margin and a large percentage of the scales between
SC° and II' of hindwing, entire ; on underside the scales all denticulate, excejit
between SM- of hindwing and abdominal margin ; scales in apical area of forewing
reduced, those of the transparent snbmarginal band very narrow, as are those in
the costal area jiroximally of the black jsostdiscal band. Forewing with seven
black bands, subapical cell-band the most variable one, sometimes vestigial.
Hindwing, below, with black subbasal band which is almost parallel to abdominal
margin, a black median band converging with the former, beginning at costal
margin proximally of middle, meeting the subbasal band distally, or being
abbreviated, bordered with red on proximal side ; a red costal spot at distal side
of subbasal band, another distally at proximal side of the band, besides the two
red transverse spots situated near anal angle : these anal spots present also above,
bordered with white in front, at least on the underside. Tlie female is like the
male, but has rather larger pale subraargiual spots on the hindwing.
Neuration : D= of hindwing shorter than D' + D', seldom as long as these
cross-veins together.
Scent-organ : scent-scales forming a layer of black-brown wool ; long hairs
black-brown, thin, ending in a long fine point, not ending abruptly as in P. telesilaiis
or P. tnarcellus ; no scales underneath the wool, except on SC.
Genitalia: S. Tenth tergite trilobate; dorso-ventral ridge of harpe reaching
ventral edge proximally of middle, the apical lobe of harpe being long, ventral
process present.
Early stages not known.
Ilab. From Goyaz, Brazil, and Bolivia to Panama, and again from Honduras
to Mexico ; not yet known from Costa Eica and Nicaragua.
Four very distinct geograjjliical races.
There are some peculiarities in the variability of this species which are worth
special notice. The subbasal band of the hindwing is present on the npperside
( 704 )
iu the two Central American subspecies and in the Sonth American snlispecies,
but is nsuiilly absent or only vestigial in tlie subspecies inhabiting Colombia,
Panama, and North Venezuela. In this geographically intermediate sidisjiecies
the submedian band of the underside of the hindwing crosses the cell at or
liroximally of M\ while in the northern races and in the southern one this band
is much more distal. On the other hand, the Central American and Colombian
races differ from the southern subsj)ecies in the scales of the black marginal and
postdiscal bands of the forewing being broader and the bands therefore more deeply
black, the postdiscal band being moreover not divided, and the black distal band
of the hindwing, above, not bearing a distinct white sjjot R- — R' proximally of the
white submarginal spot which stands in front of the tail.
Another interesting feature of agesilaus are the opposite lines of devclojiment
obtaining in the subbasal and submedian bands of the hindwing. The subbasal
band, if not complete, is on the upperside usually more or less distinct from the
cell baclvwards, the costal portion being missing. The submedian band of
the upperside, if not altogether absent, is present only in the costal region, the
posterior portion being missing.
a. P. agesilaus fortis snbsp. nov.
Papilio iieosilmis, Godm. & Salv. (iinn Hopffer, 1860, err. det.), Biol. Ceiitr. Amer., Bhop. p. 219.
n. 4R {\m)) (Oaxaca ; Atoyac).
S- Black bands broad. Forewing: first and second bhick bands abont
two-thirds the width of the interspace between them, both extending to inner
margin, or the second at least beyond SM-; pale submarginal band not wider, or
even narrower, than black postdiscal band ; the latter not includiug a distinct i)ale
line ; the scales of this band and of the marginal one rather broad. Hindwing :
abdominal edge black from base to anal angle, black subbasal band complete ;
submedian band usually distinct from costal edge to cell, or at least vestigial ;
red aual spots separate from each otlier, each comidetely surrounded by black
scaling, their white anterior borders vestigial or very narrow.
On itndcysi(h the red submedian line of the hindwing extends to hinder edge
of cell, its black border being heavy.
? not known.
Ilab. Guerrero, South-west Mexico ; Oaxaca ; and Atoyac.
The Atoyac specimens (in coll. F. D. Godman) are a transition to the next
form, the Oaxaca specimens also partly inclining towards the next.
In tlie Tring Museum 4 (J(? from Guerrero (0. T. Baron) ; name-type.
b. P. agesilaus neosilatis Hojiff. (18G6).
VaijiUu uiji'silaus, Bois-duval, Spec. Gen. Lep. i. p. 203. n. 80 (1830) (jiarllm ; ■'Mexico ") ; 'Wfidem.,
Proc. Ent. Sor: Pliihid. ii. p. 140 (1803).
Pajiilio mosilam HopfEer, Slelt. Ent. Zril. xxvii. p. i(,. n. 0 (1800) (" Mexico,'' coll. Deppe) ; Kirby,
Cat. Dhmi. hep. p. 556. n. 248a (1871) (Mexico) ; Godm. & Salv., U)ul. Cent,: Amer., li'hcp.
p. 219. n. 48. t. 8. 9. c? (1890) (Guatemala ; Brit. Honduras ; Honduras).
Papilio coiwii, Oberthiir («(/« Hew., 1854, err. det.), El. ifEiit. iv. p. 60. n. 179 (1880) (^jiarlim ;
Mexico).
Piipilio nr,esilaiis ueoHilmis, Eimer, Aril. Verivamllsch. Sclim. p. 100. 101 (1889) (Honduras ; Mexico).
cJ. Black bands of wings narrower than in /'. a. foiiis; first and second bands
of forewing abont half the width (or less) of the interspace between them at M;
transjiarent submarginal band broader than tlic lilack band stauditig at its proximal
(705 )
side. Abdominal edge of liiiidwing partly white, submedian band absent from
tippersiile or vestigial ; red anal spots with broad white border in front, black
scaling in front and behind the red spots mnch more restricted than in fortis,
and black border to red submedian line of underside of Mndwing narrower.
Subapical cell-band of forewing vestigial in one of onr specimens from Espiual,
Vera Cruz.
Hub. Vera Crnz, East Mexico ; Guatemala ; British Honduras ; Honduras.
In the Tring Museum 10 cJ (J from : Espiual, Vera Crnz, June 1800 (W. Schaus) ;
S. Pedro Sula, Honduras.
c. P. agcsilaus eimeri subsp. iiov.
Pujiilio arjeaihiuK, Godman & Salv., Biol. Cenlr. Amer., Rhop. p. 219. n. 47 (1890) (Panama).
Papilio ar/nsifuiis agrsihws, Eimer, Arlli. VenrniiiJtsrh. Srlim, p. 99. t. 1. fig. 10 (1889) (S. Juan,
West Colombia).
tj ? . Transimrent submarginal band of forewing as narrow between SC^ and
SC* as the black band standing at its proximal side, or even narrower.
Hab. Rio Dagua, West Colombia ; Upper Cauca valley.
In the Tring Museum 5 tJcJ from : R. Dagua (\V. F. H. Rosenberg) ; Popayan
(Lehmann).
The Panama specimens stand intermediate between this form and ordinary
Bogota s])ecimens. Both sexes are from this locality in coll. F. D. Godman.
d. P. agesilaiis agcsilaus Gudr. (183.j).
Papillo Equcs Achivus protf.-<fhfu>!^ Esper {non Linn^, 1758, err. det.), /.';.
Papilio cigesilaiis Gurrin & Percheron, Gen. Iiix., Lip. t. 1. fig. 1 (1835) (R. Magdalena, Colombia);
Boisd., Spec. Gin. Lep.\. p. 2G:i n. 8G {\&ii'y) (imrlim ; Colombia) ; Felder, Verh. Zool. Bot. Gen.
117™ xiv. p. 301. n. 17G (18G4) (N. Granada; Venezuela) ; Kirby, Cat. Diurii. Lep. p. 555.
n. 247 (1871) (N. Granada; Venezuela); Oberth., Et. d'Enl. iv. p. 66. n. 178 (1880)
(N. Granada ; Venezuela) ; Staud., Exot. Taijf. i. p. 16 (1884) ; Hahnel, Irix iii. p. 149. 156
(1890) (Sin Esteban) ; id., I.e. p. 203. 205 (1890) (Val«a).
Papilio eoiion Gray, Cat. Lep. Ins. Brit. Mux. i. Pap. p. 33. n. 159 (1852) {miin. nml. ; Bogota) ;
Hew., Traua. Ent. Soc. Loud. (2). ii. p. 246. t. 22. fig. 3 (1854) (N. Granada) ; Gray, List Lep.
Ihs. Brit. ilux. i. Pap. p. 45. n. 167 (1856) (Bogota).
Papilio ageKilaiis agesihitis, Eimer, Artb. Veriraiidlscli. Schin. p. 99. t. 1. fig. 11 (1889) (Colombia ;
Venezuela).
Papilio agesilaus agesihws septemlineatus id., I.e. p. 100 (1889) (N. Granada).
Papilio ngesilans Boisd. ab. septemlineatus id., I.e.. t. 1. fig. 11 (1889).
Papilio agesilaus var. eonon, JIaassen & Wcym., iu Stiibel, Jicisen S. Anur., Lep. p. 24. n. lO'J (1890)
(west side of Cordillera of Bogota).
Papilio agesilaus septemliuealus Eimer, Ortliogen. p. 47 (1897).
c?. Black postdiscal band of forewing, above, not centred by a pale line, the
scales of this and the marginal band rather broad ; anterior half of transparent
submarginal band broader than black postdiscal band. Subbasal black baud of
hindwing absent from upperside, except a thin line on M- ; submedian band absent,
merely showing through ; no white spot R^ — R^ in black postdiscal baud.
Undei-side. — Submedian band of hindwing crossing cell proximally of or at M',
the band complete as a rule, joining the subbasal band at an acute angle.
(Genitalia : Distal lobe of harpe broad, ventral edge ending proximally in a
rather heavy tooth.
This subspecies varies a great deal in size, some specimens being almost twice
as large as ntliers. We do not know if the variation is seasonal. Dr. Biirgei
( "ofi )
obtained only large specimens dnring the dry season and at the beginning of the
wet season on the eastern side of the Andes of Bogota. We have also a large
S])ecimen from Mnzo, besides some small ones. The subniediau band of the
underside of the hindwing is occasionally vestigial beyond cell. In a specimen
from the Felder collection, bearing no locality label, bnt coming jirobably from
Bogota, there are two whitish halfmoons 11- — M' within the black distal band, the
black scaling situated jiroximally of those halfmoons being ill-dctiiiod.
Ilah. Magdalena valley, C^olombia, eastwards to North Venezuela.
In the Tring Museum 94 tJc? from: "Bogota"; Valdivia, t'olombia, July
1897 (Pratt); Muzo, December 1890; Villaviceucio to R. Ocoor, January 1897,
350 — 400 m., dry season (Dr. Biirger) ; Villavicencio to Monte Rcdundo, end of
March — early April 1897, 400 — 1300 m., beginning of rainy season (Dr. Burger) ;
Peperital, Buenavista, Jannary 1897 (Dr. Burger) ; Mocotou^, Venezuela (Briceuo).
e. P. agesilaus autosilaus Bates (1861).
Papilio affesihuis, Doubleday, Li-^t Lep. Inn. Brit. Mtis. i. p. 9 (1845) (S. Amer.) ; Gray, List Lep.
Im. Brit. Mus. i. Pap. p. 4.5. n. 1615 (1856) (S. Amer. ; Brazil) ; Wall., Trann. Eiit. Soc. Loud.
(2). ii. p. 254 (1854) (Amazons); Sharpe, Proc. Zool. Snc. Loml. p. 555. n. 6 (181)0) (R. Araguaya).
Papilio autosilaus Bates, Trans. Ent. Soc. Loud. (2). v. p. 348 (18G1) (Ega) ; id., Jour/i. Ent. i.
p. 229. n. 34 (1862) (Upper Amazons) ; Felder, Verh. Zool. Bot. Gfs. WIeu xiv. p. 301. n. 177
(1864) (Ega; "Mexico," error) ; Kirby, Cat. Diurn. Lep. p. 555. n. 246 (1871) (Ega) ; Druce,
Proc. Zool. Soc. Loml. p. 245. n. 12 (1876) (Ucayali) : Oberth., Kt. d'Ent. iv. p. 66. & 115.
n. 180 (1880) (French Guiana ; TefEe ; " Colombia," errore) ; M^^cbI., Vcrh. Zool. Bot. Ges.
TF/fii xxxii. p. 304 (1883) (Surinam) ; Staud., E.cot. Tar,f. i. p. 16. t. 11 (1884) (Amazons) ;
Habnel, Iris iii. p. 250 (1890) (JlauiJs) ; id., I.e. p. 283 (1890) (Pebas) ; Michael, ihid. v. p. 214
(1894) (Sao Paulo de Olivenca, only during the dry months) ; Haensch, Birl. Ent. Ztitichr.
xlvii. p. 154 (19113) (Archidona, C40 m.).
Papilio conon, Obertbur, Et. d'Ent. iv. p. 66. n. 179 (1880) (parlim ; Peru).
Papilio agesildtis autosilaus, Eimer, Artb. Verwandtsch. Schm. p. 100. 101. t. 1. fig. 9 (1889)
(Amazons).
<?. Scales of black marginal border and jiostdiscal band of forewing narrower
than in the other forms, these bands therefore less deep black ; postdiscal band
divided longitudinally by a usually distinct pale line ; transparent submarginal
band of forewing narrower than in the other subspecies, always narrower between
SC* and SC^ than the black postdiscal band. Subbasal band of hindwing present
on upperside, snbmedian baud of undeiside of hindwing crossing cell at M\ curved
behind, often feebly developed in and beyond cell, sometimes vestigial beyond cell ;
red subanal spot M' — M' vestigial or absent, apparently never so well developed
as it is in most specimens of the preceding races ; a more or less distinct white
spot within black band proximally of white spot 11- — W of hindwing above, seldom
vestigial only, often a second spot between W and M', and the vestige of a third
between R' and R-, rarely also a trace of the spot SC- — R'.
The black subajncal cell-band of the forewing varies very much ; it is vestigial
in a Felder specimen (without locality).
Genitalia: Apical lobe of harpe narrower than in the otlier races, the dentate
ventral edge proximally not produced into a ]irominpnt tooth.
The 8i)ecimens from British Guiana have rather heavier black bands ; the
second band of the forewing is nearly as broad in the cell as the interspace between
the tirst and second bands ; the marginal and postdiscal bands are deeper black
than in atito.vlaus from the Amazons and Andes, the j)ale line within the postdisc.il
band being jiractically absent from I he n]ip('rside. The interspace between the post-
( 707 )
discal band and the discocudlnlar l>and is uarrowfr thau the postdiscal band, while
in nearly every specimen from other districts tliis interspace is wider at and before
lower angle of cell than the black postdiscal band. On the hindwing, the black
snbmedian line of the underside is rather heavy beyond cell, ending in a black spot
M' — M- which is larger than in ordinary autosilaus.
The black bands are on the whole rather narrower in tlie specimens from
Bolivia, Sonth-Eastern Pern and Goyaz (Brazil) than in the individuals from the
more northern localities (Pern, Ecuador, Amazons).
Eimer's fig. 9, I.e., said to be taken from an Amazonian specimen, has a very
short hindwing, the anal area being far less prolonged than is the case in all our
specimens. The individual which served as model may have been an imperfectly
developed specimen.
JJaL Bolivia to Eastern Ecuador ; Amazons; Goyaz; Guiana; Orinoco.
In the Tring Museum 140 cj c? from: Suapure, Caure R., Orinoco, February
and June 1899 (S. M. Klages) ; Essequibo R., Brit. Guiana; British Guiana;
Manicore ; Ljuitos ; R. Cachyaco, affl. of R. Huallaga (Stuart) ; Zamora, Ecuador
(0. T. Baron) ; Loja ; Archidona (W. Goodfellow); R. Ghuchuras, affl. of R. Palcazn,
320 m. (W. Hoffmanns); Palcazu (Sedlmayr) ; Chanchamayo (Scbnnke) ; Perene R.,
3000 ft., October -November 1902 (Watkins & Tomlinson) ; Per(^n6 R., March 1900
(.Simniis) : Caradoc, Marcapata, Eebruary 19U1, 4O0i) ft. (Ockeuden) ; Montanas,
R. Madre de Dios, September 1901 (Ockeuden); R. Slucuri, Carabaya, June 1901,
2000 ft., dry season (Ockeuden); Chirimayo, Carabaya, lOoO ft., July 1901
(Ockenden) ; Callanga, (Juzco, l-oOO m. (Garle])p) ; Cajon, Cuzco, September 1900
(Garlepp); Mapiri ; Reyes, R. Beui, August 1895 (Stuart); R. Tanampaya (Garlepp);
Yungas de La Paz, December 1899 (Garlepp) ; Province Sara, S. Cruz de la Sierra,
February — April 1904 (J. Steinbach) ; Sapucay, Paraguay, July 1902 (W. Foster) ;
Jatahy, Goyaz.
149. Papilio glaucolaus Bates (1804) (PI. IX. fig. 03. 64. 05).
Papilh, ijlaucolaus Bates, Eiil. Mi. Mag. i. p. 4. n. 7 (18G4) (Pauama).
The species has not been recognised by any of the authors who liave dealt
with this group of Papilios. Staudinger, in Exot. Tag/, i. p. 18 (1884), speaks of
glaucolaus from Panama as being larger thau macrosiilaus from Honduras ; his
glaucolaus was doubtless protesilaus, since the true glaucolaus from Panama is
smaller thau the average specimens of maeros/'l'ius. In Eimer, Arfl). Venvamltsch.
Schm. p. 102 F /.8S9), the various species and snbspecies are all muddled up. In
this work glaucolaus is called the largest protesilaus, while in fact P. glaucolaus
from Panama and Colombia is decidedly smaller than the forms of P. protc.vlaus
with which it occurs together.
We recognise three geographical forms o? glaucolaus. While the two northern
forms (Pauama and Colombia) are easily recognised by the distinctions in pattern,
the third subspecies almost exactly resembles P. protesilaus protesilaus, being
distinguishable with certainty only by comparison of the genitalia.
S. Postdiscal Ijand of Ibrcwing more widely separate from lower angle of cell
than in P. p. protesilaus, or the hindwing more obtnsely dentate at R' and R' and
the posterior submarginal sjiots slightly huffish, the wings not being so irare white
as in P. p. protesilaus.
Scent-organ as in P. prote.vlaus.
( 708 )
Genitalia: Harpe shorter than in P. protesilmts ; deflexed dorso-apical ridge
(r/r) snbvertiral on tlie harpe, contiuned to apex, enlarsred into a prominent dentate
tooth {() ; ventral process {rp) short, reaching halfway (or less) to ventral ed?e of
clasper ; central process {cp) also short (Fig. 1).
? not known.
Hab. Panama to Matto Grosso, not known to us from the Brazilian siihreo-Jon.
Fig. 1.
Fig. 2.
Fig. 3.
a. p. ghrticoUnis glaucohois Bates (1864) (PI. IX. fig. 04).
Piiplllo (/hiicohms Bates I.e. (Panama).
Piq/iVin prntesilaus var. e. P. gUuicolmis, Kirby, Cul. Dhirii. Lep. p. 556. sub. D. 248 (1R7I) (Pan.ima).
Papilio prol, silaus, Obertbiir, Ef. d'Ent. iv. p. CC. sub n. ISl (1880) (Carare, greenish, tliis insect teste
Jordan).
PapiUo prntesiJaus, Godman & Salv., Biol. Ccnlr. Amer., Pliop. ii. p. SI.*!, n. .30 (1890) (paiiim).
c?. Both wings pervaded with a greenish tint, which becomes very distinct if
a series of P. gl. glaucolaus are put side by side with P. protesilaus. Forewing
similar in appearance to that of P. a^. agesilaus; bands 1 and 2 stopinng at SM-,
2 often not reaching this vein ; fifth band narrowed to a point, mostly not quite
reaching lower angle of cell ; sixth band mure di.stal in position than in P. protesilaHs,
being IJ to 3 mm. distant from lower angle of cell, gradually widening costad, not
narrowed before R- ; transparent submarginal interspace at SC* at least twice the
width of the black marginal band. Hiudwing : red anal spot often much reduced,
being overpowdered with black scales at the abdominal side ; black sfiot situated in
front of red one usually oblong, often continued abdominad beyond M-.
Underside : fifth baud of forewing mostly reaching across cell, extended at
least to third cell-fold in our .series. Subbasal band of hiudwing touching apex
of basal cellule ; median band a little distal of base of M', the band usually not
touching the point of origin of this vein ; cross-veins partly bordered with red,
sometimes the red scaling forming a complete ring in apex of cell.
Genitalia : Harpe tapering ; tooth of dorso-apical ridge broad ; central process
slightly spatnlate.
Length of forewing : 40 to 47 mm.
Hah. Panama ; Colombia, except S.W. coast ; jjp
Venezuela.
In the Tring Museum 20 $$ from: Panama; Mnzo,
"Bogota;" La Vega de San .Tnaii.
ablv also Northern
December 1890;
( 709 )
b. 1' iiliiiicolaiis mclitciius subsp. nov. (PI. IX. fig. 03).
Papilin spec. ?, Staudinger, E.riit. Taijf. i. p. 18 (1S84) (R. San Juan).
S. Black bands of'tlie forewing on tbe whole broailev tlian in the jireceding ;
the submarginal transparent interspace narrower, being nsually onl}' a little broader
at SC^ than the black marginal band, or as broad as this band, specimens in
which the submargiiial interspace is nnusnally wide being recognizable as belonging
to P. glaiK'olaus melacnus by a corresponding reduction of the interspace between
bauds 5 and G.
In this as well as the preceding form the black jjostdiscal band of the forewing
stands often distally of the point of bifurcation of the subcostals SC^ and SC°, which
is rarely the case in tbe following form.
Length of forewing : 40 to 50 mm.
Hah. Rio Dagua, West Colombia ; Upper Cauca valley.
In the Tring Museum 22 ? ? from: E. Dagua (W. F. H. Rosenberg), type ;
Popayan (Lehmann).
c. P. glaucolaus leucas subsp. nov. (PI. IX. fig. 65).
(?. Wings not pervaded with green, except base, which is distinctly green np
to second band on forewing ; postdiscal band of forewing in the same position as in
P . p. protesilaus, being much nearer the lower angle of cell than in the preceding
forms of P. glaucolaus, and being, moreover, narrowed before R- in most specimens ;
transparent snbmarginal interspace wider than in P. gl. glaucolaus; white inter-
S])aces of cell covered with narrower scales, therefore appearing less densely scaled
than in the preceding forms. Subbasal band of underside of hindwing entering
basal cellule.
Genitalia : Harpe shorter and more obtuse at apex than in P. gl. glaucolaus
and melaenuSi the tooth of the dorso-apical ridge narrower, and the central process
longer, pointed (Fig. 1, p. 708).
Resembling in pattern more closely P. p. proh'silaus than the Colombian
forms of P. glaucolaus; ditfers from P. p. pi-otesilaus in the hindwing being morn
obtusely dentate, in the ad- and snbmarginal spots being ])ervaded with buff, which
colour becomes rather distinct if specimens of P. p. protesilaus and P. gl. leucai
are compared side by side; the second band of the forewing is on the npperside
nsually much narrower from the cell backwards than on the underside, this jiortion
of the band appearing on the npperside much blurred in consequence of the band of
the underside shining through, and being in some individuals very thin or even
absent ; the fourth cell-band is shorter on the whole than in Guiana specimens of
P. protesilaus. The frons has always clearly defined white lateral bands, never
being all fuscous as it is in many specimens of P. p. protesilaus. The scales of the
pale bine admarginal spots of the hindwing are dentate, while in P. p. protesilaus
they are mostly entire.
Length of forewing : 41 to 48 mm.
Ilab. Orinoc(j, Caura R. ; British Guiana ; Amazons ; Eastern Ecuador ; Peru ;
Slatto Grosso. Name-type irom Rio Chnchuras.
In the Tring Museum 60 (? <? from : Suapure, Caura R., Orinoco, apparently
all the year (S. M. Klages) ; La Vuelta, Caura H., May 19(.)3 (S. M. Klages) :
R. Demerara, August 1897; Upper Real Berbice R. ; R. Negro; R. L^anpes,
B.Negro; Coca and Archidona (W. Goodfellow) ; Upper Amazons ; R. Chnchuras,
( 710 )
affl. of R. Palcazn, 320 m. (W. Hoffmanns); Rio Cachyaco, affl. of R. Hnallaga
(Stuart) ; R. Ucayali (Stnart) : Iqnitos (Stnart) ; Chancliamayo (Sclmnlce) ; Pereii(5 ll.,
4000 ft., Angust— September 1002 (Watldus & Tomlinson) ; Villa Blaria to
Diamantino, Matto Grosso, January IsOT (Andeer).
150. Papilio molops spec. uov. (I'l. IX. fig. On. 01. 02).
As in P. (/huicnlaus there are also in the present species no striking characters
in the pattern by which one could recognise P. molops with absolute certainty, except
the molops form from the coast of Ecuador and Colombia and the form from
Brasilia, which differ very conspicuously from the forms of P. protesilaus occurring
in the same districts. However, as the genitalia of mo/ojJS (of the male, the
female being unknown) are constantly different from those of the various South
American subspecies of protesilaus, it is not very difficult to distinguish also the
molops specimens from Surinam, the Amazons and the Andes from protesilaus
as well as from glaucolaus.
S. Frons always greyish white at the sides, inclusive of tl\e Brazilian form
with black antenna. Both the first and second black bands of the forewing
extending to hinder margin, or the second slightly abbreviated, or the first
abbreviated ; in the last case the postdiscal band very broad, being at lower angle
of cell as broad as the transparent submarginal band ; fourth band reaching
nearly across cell, seldom stopping at third cell-fold.^ Hindwiug less strongly
dentate than in P. protesilaus archesilmis, but more strongly than in P.glaxeolrois;
black admarginal bar R^-R^ as slender as in P. protesilaus, while the admarginal
bars R3— M- are usually broader than in that species.
On the underside, the snbbasal and median bands of the hindwing on the
whole closer together than in P. protesilaus, the interspace less widening costad
and that portion which lies behind the cell longer, the black patch M'— M-
which connects posteriorly the subbasal and median bands being smaller ; red
anal bar usually quite close to the black anal marginal spot, the white spot
separating them from one anotiier being reduced to a small dot in nearly every
specimen.
Scent-organ as in P. protesilaus.
Genitalia : Dorso-apical deflexed ridge of harpe rounded-dilated, denticulate,
but not produced into a prominent triangular tooth (Fig. 2, p. 708).
Early stages not known.
Jlab. South America.
Three subspecies.
a. P. molops molojis subsj). nov. (PL IX. fig. G2).
c?. Antenna tawny, club black, at least on niiperside ; black lateral streak
of abdomen as broad as the bnffish white streak situated above "it.
Wings. — Upperside : first band of forewing stopping at SM^ second band
strongly tapering, reaching hinder margin, broad in cell, third and fifth bands
also broad, interspace between fifth and sixth bands not wider than the sixth
band, which is broader than in all the other forms of the species allied to
P. protesilaus, the white costal dot of tliis interspace minnte, the sixth band
very close to lower angle of cell, therefore differing consideralily in jiosition from
( 711 )
the baud of P. (/laitcolaus ; margiuul band aLso broad, the siibmargiual intersjiace
tlierefore narrower than in the other molops forms, being at R- about as wide as
the marginal bands. Hindwing : red anal spot large, stopping halfway between
Ml and IP, obliquely truncate, the black spot M' — M- in front of it large, trapezi-
form, somewhat ])rodnced basad at M', no black bar M- — SM- proximally of red
spot ; postdisco-margiual band broad, partition II- — R' wider in middle than the
white admarginal lanule R- — R^ ; partition R^ — M' also broad, its edges not
sharply defined, the white (slightly bnfifish) submarginal lunule which stands at its
distal side reduced, somewhat powdered with black, as is the white submarginal
spot M' — M- ; the postdiscal black bar bordering this latter spot proximally joining
the black anal sjiot ; white marginal ^pot before R- small, the one before tail
less extending distad than in the next form ; first admarginal Inunle vestigial,
submarginal lunule R^ — R^ also densely shaded with black.
Underside : — fourth band of forewiug extending across cell or very nearly.
iSnbbasal band of hindwing distally of basal cell, not entering it; red scaling of
median band extending along cross-veius, forming a more or less complete ring
in apex of cell ; red transverse bar W- — SM- heavy ; red bar M' — M^ Pj-shaped ;
black arrowhead-shaped patch on abdominal fold large, white lunules W — M-
small.
Genitalia : Deflexed dorso-apical ridge of harpe more or less abru23tly dilated,
the lobe sinuate in one of our two Ecuadorian specimens, no teeth beyond the lobe ;
teeth on ventral edge of harpe irregularly placed and of different sizes ; central
process rather broadly spatulate, denticulate on dorsal side, the teeth somewhat
curved in the direction of the base of the process ; ventral process acute, non-
dentate, not quite reaching the ventral edge of the elasper, its ajjex somewhat
curving distad and away from the elasper.
Ilab. N.W. Ecuador : R. Cayapas (Flemming and Miketta), t>/pe ; Cachabi ;
low country, January 1897 (W. F. H. Rosenberg).
2 c? c? in Mus. Tring. In coll. Charles Oberthiir, from Juntas, R. Uagua,
West Colombia (M. de Mathan).
li. P. molopss ketaerius snbsp. nov. (Pi. IX. fig. 61).
(?. Very closely agreeing in pattern with F. protesHaus protesilaus ; the
Colombian specimens smaller than the Colombian P. p. archesilaus, the hindwing
less strongly dentate. Frons never entirely brown-black, as is so often the case
in Andesian specimens of P. p. jx-otesilaus. First band of forewing always
extending to hinder margin, second band very broad in cell, reaching to hinder
margin in most specimens, or at least close to it. Hindwing with a white
submarginal lunule R' — R-, which is thinner than the admarginal one, being
occasionally vestigial ; black admarginal spots R' — M^ rather larger than in
P. p. protesilaus, especially M^ — M-.
Underside : intersjsace between subbasal and median bands of hindwing longer
than in protesilaus.
Genitalia : Deflexed dorso-apical ridge of harpe (Fig. 2, p. 706) usually
dentate beyond the proximal dentate lobe ; ventral edge of harpe more densely
dentate than in P. m. molops, central process more slender and its teeth shorter,
ventral process longer, reaching edge of elasper, carved towards this edge, not
away from it, usually with some teeth on the distal side or at the apex.
( 712 )
Iliih. Guiana; Amazons ; Colombia ami Ecuador (Pacific side excepted) ; Peru;
]'()liviu ; fi/pc froTa Saramacca K., Snriiiam.
In the Triug Museum 17 cTcf from : " Bogota" ; Villavicencio to Kio Ocoor,
East Colombia, January 1807 (Dr. Burger) ; R. Demevara ; Saramacca B., Surinam,
May 1893; Iqnitos ; "Amazons"; Archidona, N.E. Ecnador, April 1899 (W.
Goodfellow) ; Coca, R. Napo (W. Goodfellow) ; R. Chncliuras, affl. of R. Palcazu,
320 m. (AV. HotTmanns) ; Mapiri, Bolivia.
c. P. mnlops megalnriDi subsji. nov. (PI. IX. fig. fin).
S. Antenna black, as in P. jirotcsilaati idyricornis, bat sides of frons white;
white dorso-lateral stripe of abdomen narrow. First and second band of forewing,
upperside, reaching binder margin, but second strongly tapering, interspace a little
wider than in the preceding form ; sixth band separate from lower angle of cell ;
transparent snbmarginal interspace narrower than in P. in. //etacriiis, not being
wider within subcostal fork than the interspace between fourth and fifth bauds
measured at third cell-fold. Black markings a little more extended than in
hetaerius, first white admargiual lunule hardly traceable, second very thin, or
also vestigial, no white snbmarginal lunule R' — B', the admargiual one being alone
present, the other white ad- and submarginal spots also smaller than in ketaerius ;
red anal spot longer, the white spot M' — M- standing at its discal side
correspondingly smaller, the black spot M' — M^ in front of the red spot larger,
continued abdominad and produced basad in the abdominal fold, the spot some-
what widened also discally at M', there being some black scales between the
spot and the cell corresponding to the outer edge of the median band of the
underside ; tail longer and broader than in the other two forms.
Interspace between subbasal and median bands of underside of hindwing
rather wider than in the two preceding forms, the median band being distinctly
more distal, crossing cell at base of M' ; white lunule C — SC- hardly vestigial,
only one white lunule R' — R-, the other white lunules also reduced.
Genitalia : Distal lobe of harpe narrow, teeth small ; deflexed dorso-apical
ridge less dilated than in the other forms of molops ; central process curved
proximad at tip, teeth vestigial ; ventral process not quite reaching edge of clasper,
curved towards this eilge, non-dentate.
Ilab. Brazil : Leopoldina.
In the Triug Museum 2 cJcJ.
lol. Papilio protesilaus L. (175n) (PI. IX. tig. OO. 07).
Merian, Iiis. Surinam t. 43 (1705); Gronov., Zoophyl. ii. p. 188. n. 726 (1764); Seba, Thesaur.
iv. p. 44. t. 36. fig. 11. 12 (1764) ; Aubent., Planch. Eiilum. t. 44. fig. 1. 2 (1765).
Papilin Eqiies Achiviis jirotpxilaus hmnv, Si/st. Xtil. ed. x. p. 46H. n. 29 (1758) (parlini); Clerck,
Icon. Inn. ii. t. 27. fig. 2 (1764) (fig. sat mala) ; Liunt-, Mux. Lud. Ulr. p. 209. n. 28 (1764)
(partini) ; id., Syst. Nal. ed. xii. p. 752. n. 3'.l (1767) (paiiim) ; Fabr., Si/kI. Ent. p. 450. n. 36
(177.5) (/«(Wj«i) ; Sulzer, 6'f.iv-/(. /«.■!. i. p. 143; ii. t. 14. fig. 5 (1776) (synon.ywrt'm) ; Qoeze,
Ent. liei/li: iii. 1. p. 64. n. 39 (1779) (cit. " Pet. Mus.'' e.\cl.) ; Cram., Pap. E:rnl. iii. p. 16. t. 202.
fig. A. B (1779) ('North & South America ') ; Fabr., Spec. Ins. ii. p. 14. n. 56 (1781) (paiiiin);
id., Manl. Ins. ii. p. 7. n. 62 (1787) (jHirlim); Jabl. & Herbst, Xalurs. Schm. iii. p. 147. n. 97
(1788) (partirii ; Sulzer, ed. Roem., Gesch. Ins. p. 17. t. 14. fig. 5 (1789) (" Am. sept." errore) ;
Gmelin, Sijst. Nat. i. 5. p. 2243. n. 39 (1790) (yj.(W;«.) ; Fabr., Ent. 8i/st. iii. 1. p. 23. n. 69 (1793)
(pailim) ; Turton, Syst. af Nat. iii. 2. p. 16 (180()) (partim).
(713 )
PaplUo Eques prolesUaus, Lange, in Linne, Sysl. Nat. p. 41)3. n. 29 (1760) (parlim).
Pa/iilio (Achiriis) prolesilaiis, Miiller, Xaliirs. v. 1. p. 577. n. 39 (1774) ("N. Am." error).
Pupilio Eques Achhma proteHileiis (!), Meuscbsn, in Gronov., Zouplvjl., Index (1781).
Papilio protesilaus Godart, Enc. J/clIi. ix. p. 50. n. 73 (181'J) (parti m) ; Lucas, i^^j. Exot. p. 41
(1835) ( jxiiiim ; Brazil ; iiecfi;/.); BoisJ., Spec. Gen. Lep. i. p. '262. n. 85 (1830) (French Guiana ;
Brazil) ; Lucas, in Gucr., Diet. Pitt. Hist. Nat. vii. p. 4G (1838) {pai-tim) ; Duncan, in Jard.,
Nat. Libr. xx.xvi. p. 104. t. 4. fi;^. 2 (1843) (partim) ; Doubl, List Lep. Ins. Brit. Mils. i. p. 9
(1845) (Brazil ; Brit. Guiana) ; id., Westw. & Hew., Gen. Diurn. Lep. i. p. 15. n. 137 (1846)
(syn. jiartim ; Honduras ; Guiana ; Brazil) ; Erichs,, in Schomb., F. F. Brit. Guiana p. 593
(1848) ; Gray, Cat. Lep. Ins. Brit. Mus. i. Pap. p. 33. n. 160 (1852) {partim) ; Burm., Ahh.
Naturf. Ges. Halle p. G3 (1854) (Meriau'.s plate 43) ; Gray, List Lep. Lis. Brit. Miis. i. Pap.
p. 45. n. 168 (1856) (Brazil ; R. Demerara) ; Bates, Trans. Ent. Soe. Limd. (2). v. p. 348 (1861)
(partim) ; id., Juurn. Ent. i. p. 229. n. 33 (1862) (partim) ; Weidem., Proc. Ent. Soc. Pliilad.
ii. p. 148 (1863) (partim) ; Bates, Pror. Zoul. Sac. Land. p. 241. n. 2 (1863) ; Felder, Verli.
Zool. Bot. Ges. Wieu xiv. p. 301. n. 178 (1864) ; Kirby, Cat. Diurn. Lep. p. 555. n. 248 (1871)
(partim) ; id., /.■". p. 811. n. 248 (1877) ; Burm., Descr. Rep. Argent, v. Lep., Atlas p. 3. n. 1
(1879) (mixture of many .species) ; HopfF., Slett. Ent. Zeit. xl. p. 52. n. 18 (1879) (Surinam,
Brazil, Peru) ; Auriv., A'. Sv. Vet. Akad. IlamH. xix. 5. p. 29. n. 28 (1882) (recensio critica ;
cit. " Herbst t. 43" et "Lucas t. 21 " exceptae) ; Eimer, Artb. Verwandtsch. Sc/im. p. 50. 103.
108. 1. 1. fig. 5. 6 (1889) ( partim) ; Haase, Untersiicli. Mimicry i. p. 82 (1893) ; Eimer, Orthogen.
p. 33. 44. 47. 48. 217. 397 (1897) (partim).
Protcsilaus leiliis Swainson, Zool. lUiistr. ii. t. 93 (18 52) (nom. nov. loco " protesilai").
Cosmodfsmus protesilaus, Kirby, in Allen, Nat. Libr., Lep. i. p. 273 (1896).
Iphiclides protcsilaus, id.. I.e. t. 68. fig. 2 (1896)
Uuder the uame of protesilau-'t Liune described a mixture of severtal bauded
species of Papiliouids and N\-mphalids. As the wings are said in his diagnosis
to be white, we restrict the name to a black-banded wliite Pajiilio, as has been
done by every author since Linne's time. However, which of the various si)ecies
of the present group the white Papilio of Linn6 was, nobody can possibly tell
with certainty from the description given by Linne and the figures quoted by him.
In Clerck's figure the hindwing is obviously too obtusely dentate. There is a
specimen (without abdomen) in the collection of the Linnean Society of London
which agrees fairly well with Clerck's figure.* Aubenton's figure, which is very
good for that time, represents doubtless the most common one of the different
white species, to which insect we apply the name protesilaus. As Linnd's
Neotropical Lepidoptera were i)ractically all from Surinam, we treat also in this
case the Surinam form as nomenclatorially typical.
S 9 . Submarginal interspaces of hindvviug white, rarely slightly washed with
yellow ; dentition of hindwing stronger than in P. glaucolaus. The first and
second band of the forewiug on the whole shorter and narrower than in P. molops.
Frons often all black or slightly whitish at sides.
Scent-organ : wool long and dense, consisting of long and thin hairs which
are widest iu centre, being much thinner than in P. telesilaus.
Genitalia different in the various subspecies ; dorso-apical ridge of harpe
. slightly or strongly dilated, always strongly dentate, in the South American
forms widened to a large triangular tooth (Fig. 3 ; p. 708) ; the ridge strongly
deflexed, lying almost flat upon the main body of the harpe.
Early stages not known. The larva figured by Slerian, I.e., is not that of
this sjiecies, but of a Heliconiiis. It is much to be desired that the early stages
of this and the allied species be carefully observed.
Hah. Mexico to Paraguay and Kio Grande do Snl.
♦ See p. 4i:i.
( 714 )
a. P. protesilaus penthesilaus Feld. (1805).
Papilio peiithesllaus Felder, Vcrh. Zoal. Bot. Ges. Wieu xiv. p. 301. n. 181 (18G4) (Mexico; 7iom.
mid.) ; id., Reiw. Novara, Lep. p. 52. n. 40. t. II. fig. C (18G5) ; Staud., Exot. Tagf. i. p. 18
(1884) (Mexico): Godm. & Salv,, i?/.-/. Ceiitr. Amer., lihop. p. 214. n. 40 (1890) (Mexico;
Atoyac, Oaxaca, Yucatan).
Piijiilio protesHmis var. c. P. penthesilaus, Kirby, Cat. Viiirii. Lep. p. 55G. sub n. 248 (1871) (Mexico).
Papilio archesilaiis va.T. peiil!iesiluu,9, Obertliiir, Et. d'Eiil. iv. p. G7. u. 182 (1880) (Mexico).
S. A large form. Dorsal stripe of abdomen very narrow. First band of
forewiijg stopping at SM-, second band reaching a little beyond SM-, strongly
tapering behind, fourth band short, triangular, rarely reaching to second cell-fold,
sixth band separate from lower angle of cell, slightly narrower from R^ to SC^
than behind R-, posteriorly almost separated from the marginal band in most
specimens, there being a semitransparent space behind M- between the two bands ;
submarginal transparent interspace bearing very narrow scales which are widest
at their apex, the apical sinus being more or less distinct, these scales easily falling
off. Red anal spot of hindwing large ; black postdiscal lunules R- — M' more
or less separate from each other ; black admarginal lunules reduced ; edge of
wing white, except the very tips of veins C, SO-, R' and M- ; dentition very
prominent, especially at Rl
Vmlerside : second band of forewing broader than above between M and SM'.
Subbasal band of hindwing about half the widtli of the median band ; red scaling
of the latter more or less extended along cross-veins, often forming a ring in
apex of cell ; red bar R^ — M' not much continued along M'.
? not known.
Genitalia : S . Apical lobe of harpe acuminate, the apical and ventral margins
dentate, dorso-apical deflexed ridge rounded, widened distally of the central
process, serrate ; central process long, slender, slightly spatnlate, somewhat
denticulate at the apex ; ventral process reaching a little beyond the edge of
the valve, non-denticnlate, pointed, applied to the valve, somewhat S-shaped, its
curvature corresponding to that of the inner surface of the valve.
Early stages not known.
Hab. Mexico : Vera Cruz, Yucatan, Atoyac, Oaxaca.
Felder's specimens were from Oaxaca. The individuals from East and South
Mexico are not always different from the next form.
In the Tring Museum, 4 cJc? from: Oaxaca (ex coll. Felder); Motzorougo ;
Orizalia.
b. P. protesilaus macrosilaus Gray (1852).
Papilio proleailaiis var. b Papilio macrusilaus Gray, Cat. Lep. Iiis. Brit. Miis. i. p. 34. sub n. IGO
(1852) (Honduras) ; id , Li.it Lep. Ins. Brit. Uus. i. Pap. p. 4G. sub n. 168 (185G) (Honduras).
Papilio protesilaus, Weidemeyer, Proc. Ent. Soc. Philad. ii. p. 148 (1863) {partiui) ; Boi.sd., Cons.
Lip. Guatem. p. 6 (1870) {piarlim ; Guatemala) ; Butl. & Druce, Proc. Zool. Soc. Land. p. 365.
n. 375 (1874) (Costa Rica) ; Godm. & Salv., Biol. Centr. Amer., Rhop. p. 214. n. 40. t. 68. fig. 7.
genit. (1890) {parlim; Guatemala: Vera Paz, Cubilguitz, Polochic valley, Cahabon ; Brit.
Honduras : Corosal, R. Sarstoon ; Honduras ; Nicaragua : Chontales).
Papilio protesilaus var.? a. uiaerosilaus, Felder, Vcrli. Zool. Bot. Ges. M'iui xiv. p. 301. sub n. 178
(1864) (Honduras).
Papilio protesilaus var. a. P. macrosilaus, Kirby, Cat. Diurn. Lep. p. 556. sub u. 248 (1871)
(Honduras).
Papilio macrosilaus, Staudinger, Exot. Tarjf. i. p. 18 (1884) (Honduras).
Papilio protesilaus rubrocinclus macrosilaus, Eimer, Artb. Verwaiidtseh . ISJim. p. 107 (1889)
(Honduras).
( 715 )
S ? • Similar to penthesilaus, paler in appearance. Sabbasal band of forewing
tliiuner, second band also narrower, stojiping short at SM', crossing M one mm.
]iro.\imally of M", the interspace between bands 1 and 2 being rather narrower
at M than that between bands 2 and 3; band 4 absent, or vestigial at costal
margin, or represented by a distinct costal spot ; band 5 subangnlate at R^, narrow
from R'^ costad, merged together with the marginal band at M° or close behind
it, the pale insterstitial spot behind M", if at all present, being smaller than
in penthesilaus ; marginal band less sharply defined and a little paler than in
penthesilaus, the scales composing it being narrower ; submarginal transparent
interspace bearing very narrow hairlike scales which do not fall off easily, being
present even in worn specimens, while the respective scales of penthesilaus, which
are much broader, are nsnally lost even in comparatively fresh specimens.
Red anal spot of hindwing narrower than in the Mexican form ; black postdiscal
Innules R" — M' on the whole more reduced, while the costal portion of the black
postdiscal band is a little wider than in that subspecies.
On the mtderside the red line of the median band coutinnes along M',
reaching usnally the red transverse bar M' — M'^, this red bar more or less widely
interrupted, being, like the anal red bar, thinner than in penthesilaus.
Genitalia: c?. Apex of harpe subtriiucate, more obtuse than in penthesilaus;
deflexed dorso-apical ridge less rounded-dilated than in penthesilaus ; ■ central
process much shorter, spatnlate, strongly dentate at apex ; ventral process also
much shorter, not reaching ventral edge of valve, curving distad. The clasper
figured by Godman & Salvin I.e., is that of this form.
Ilah. Guatemala ; British Honduras ; Honduras ; Nicaragua.
In the Tring Museum 12 6 S , 1?, from : Vera Paz, Guatemala ; San Pedro
Sula, Honduras.
c. P. protesilaus leucones subsp. nov.
Papilio protesilaus, Godman & Salv., Trans. Ent. Soc. Land. p. 126. n. 235 (1880) (Sta. Marta).
S. Resembling P. p. macrosilaus. Black dorsal line of abdomen absent (type),
or very narrow. Wings, upperside. — Forewing : first band thin, stopping short
at SM-, second band continued to hinder margin or nearly (type), interspace
between second and third band of the same width at M as between second and
first band ; fourth band short ; sixth band narrow from R^ forward ; transparent
submarginal space with hair-scales as in Honduras specimens of macrosilaus, the
wing in costal region less densely scaled than in archesilaus and dariensis.
Hindwing : red spot large, black postdiscal lunules R- — M' ill-defined, shaded with
white, white ad- and submarginal interspaces large, especially the submarginal spot
R^ — R' ; edge of wing white, except at tip of veins, but the fringes partly black
between C and R= ; dentition of wing much less prominent than in the other
Colombian forms and the Central American ones.
Underside. — Forewing : postdiscal band more straight from M' to SM- than
in the allied forms of P. protesilaus. Hindwing : median band half as broad
again as snbbasal band ; black admarginal bars C — R^ thin ; white submarginal si)ot
R' — R- nearly as wide as the admarginal lunule ; white submarginal spot R- — W
twice as broad as the admarginal spot.
Genitalia resembling those of mao'osilaus, but differ in one or more of the
teeth at the ventral edge of the harpe being prolonged, and vertical on the plane
( 7ir, )
of the clasper. Ventral process of harpe simple, pointed, not reaching ventral edge
of clasper ; central process spatulate, strongly dentate at apex, tiie teeth mure or
less curving dorsad.
Ilab. Sta. Marta, North Colombia : 2 S S \a coll. Godmau ; type from
Manaiire (F. Simons).
(L P. protesilaus dariensis siihsp. nov.
Pajiil/u prnti'xilnux var, macrosilaiis, Boisduval, Cons. Le.p. Guatrm. p. G (1870) (Costa Rica ;
= aychesihuis ~ Flantbe du Perou ex errore).
Pnjiilin archrsilauK var. macnisilmis, Oberthur, Et. d'Eiit. iv. p. 07. sub n. 182 (1880) (Costa Rica).
Pajillio pcnthexilanx, Godman & Salvin {noii Felder, 18(15, err. det.), Biol. Ceiilr. Amci:, Ithnp.
p. 214. n. 40 (1890) l^parlim ; Chiriqui ; Panama).
Nearest to archi'silaus, with which it agrees in the hhiolv dorsal stripe of the
abdomen being broad, in the distal edge of the liindwing being entirely or almost
entirely black between C and R', iu the ventral process of the harpe being
denticulate, etc.
Wings less pnre white than in an-Jiesilaus, a little more extended green at
base, slightly broader ; bands of forewing on the whole thinner, first baud always
stoj)ping short at SM-, second band tapering behind, seldom reaching a little beyond
SM-, often not extending to this vein, fourth band short, triangular, rarely attaining
second cell-fold, sixth band usually well separate from lower angle of cell, evenly
curved, not narrowed before R- or only a little, white scaling of cell denser than iu
macrosilaus, the scales being larger both on ujiper- and underside, hair-scales in
submarginal transparent space also broader. Tooth R- of hindwiug on the whole
less prominent than in arc/iesilaus, the greyish blue admarginal lunnles smaller ;
black admarginal spots C — R- not separated from edge of wing, there beiug only a
small white spot in front of SC- and R' appearing as an anterior prolongation of
the white lunnles standing behind these veius, the fringe remaining usually black
also at these small white spots.
On the viulersiile, the subbasal band of the forewing narrower than iu
archesilaus ; red median line forming a more or less complete ring in apex of cell,
as in archesilaus, the line continued along M\ joining the red transverse halfring
W—W.
?. Wings pervaded with yellow, es])ecially iu anal region of hindwing.
Genitalia: i . Harpe on the whole a little shorter than in archesilaus; deflexed
dorso-apical edge widened proximally, the dilated j)ortion ending distally in a
triangular tooth, which is more distal in position than the respective tooth of
archesilaus ; ventral process more or less denticulate on the distal side from
base to apex. ? not dissected.
Length of forewing : c?, 43 to 53 mm. ; ¥ , 47 to 56 mm.
Ilab. Costa Rica ; Panama : Chiriijni, type ; islands near the west coast of the
Isthmus of Darien : Gobernador, Brava, Jicaron, Cebaco.
The specimens from those islands are jiartly a transition to archesilaus, while
the Costa Rica individuals approach marrosilaux.
In the Tring Museum 29 JcJ, 2 ? ?, from: Carillo, Costa Rica, 3UU0 ft.,
October 19U4 (A. Hall) ; Chiriqui ; Boquete, Chiriqui, 3500 ft. (Watson);
Gobernador I., Jicarou I., Brava I., and Cebaco I., January and February 1902
(J. H. Batty).
( 717 )
e. P. profesilaiis archesilaus Fold. (1S67) (PI. IX. fi;,'. 66).
Piiji'ilio archesilaus Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 301. n. 180, p. 345. n. 89 (18r,4)
(Bogota) ; id., Reise Novara, Lep. p. 51. n. 39. t. 11. fig. a. b (1865) (Bogota) ; Oberth.,
Et. d'Enl. iv. p. 67. n. 182 (1880) (parlim ; Muzo, October) ; Hahnel, Iris iii. p. 203 (18'J0)
(Valera ; large insect).
Papilio proteailaus var. d. P. archesilaus, Kirby, Cat. Diurn. Lep. p. 556. sub n. 2-18 (1871).
Papilio protesilaus, Eimer, Artb. Verwandtsch. Schiii. t. 1. fig. 5 (1889).
Papilio podalirius var. archesilaus, Staudinger, Exot. Tagf. i. p. 17 (1884) (Venezuela).
Papilio penthesilaus, Godman & Salvia (no7i Felder, 1865, err. det.), Biol Cenlr. Amer., Rhop. p. 214.
n. 40 (1890) {parlim ; Colombia).
Papilio jnotesilaus ruhrocinclus archesilaus, Eimer, I.e. p. 106 (1889) (Colombia).
Papilio glaucolaus, Eimer {non Bates, 18G4, err. det.). I.e. p. 107 note (1889).
Papilio protesilaus ruhrociucius archesilaus (jlaueoUms, id.. I.e. p. 108 (1889) (this form ? Panama
wrong locality ? ; on p. 107 " largest ^)rotes(7nw.« ").
(?. Usually larger than p. protesilaus, hindwing more strongly dentate, median
band of underside of hindwing mostly more distal ; the apex of the cell more or less
bordered with red inside, ventral process of harpe on the whole more densely
denticulate. In the specimens from the Rio Dagna (West Coast) and from the
Canca valley the red line R^ — M' of the underside of the hindwing is shorter than
in individuals from other places, being only a little produced along M^ The
individual variability in the bands appears to be considerable ; band 4 of forewing
rarely absent, sometimes reaching almost across the cell. ? not known to us.
Length of forewing : 6, 50 to 60 mm.
Hab. Colombia, Northern Venezuela, Western Ecuador.
Eimer, I.e., says under what he terms " protesilaus rubrocinctus mihi " : " Here
belong the large forms which live in the neighbourhood of the Equator, from
Colombia to Mexico." The purport of this innocent-looking statement will be under-
stood if one remembers that according to Eimer there is a " law " that the forms of
a species are the smaller the farther away from the Equator (non-tropical countries)
they live. Now, in the case of protesilaus this statement is very unfortuiinte,
inasmuch as the form of protesilaus living under or near the Equator (namely in
the Amazon valley, East Ecuador and Peru) is smaller than the more northern
forms.
In the Tring Museum 114 $$ from: R. Dagua (Rosenberg); "Bogota";
Peperital to Buenavista, January 1897, dry season (Dr. Burger) ; Guayaquil,
Colombia, January ; Paramba, N.W. Ecuador, 35U0 ft., March 1897, dry season
(Rosenberg) ; Mocotone and Merida, Venezuela (Briceno) ; Campo Alegre, Cumana,
January 1899 (Andre).
/. F. protesilaus protesilaus L. (1758) (PI. IX. fig. 07).
Papilio Eques Achirus protesilaus Linn^, I.e. (1758) (partim).
Papilio protesilaus, Godart, I.e. (1819) (iiartim); Boisd., I.e. (1836) (parlim); Lacord., Ann. Soe.
Eut. Fr. ii. p. 383 (1833) (Guyane) ; Doubl., List Lep. Ins. Brit. .\Ius. i. p. 9 (1845) (partim ;
Brit. Guiana) ; Wall., Trans. Eut. Soo. Loud. (2). ii. p. 254 (1854) (.Vmazons, habits) ; Gray,
List Lep. Ins. Brit. Mas. i. Pap. p. 45. n. 168 (1856) {partim ; R. Demerara) ; Butler, Cat.
Diurn. Lep. deser. Fahr. p. 239. n. 21 (1869) (Demerara) ; Moschl., Verh. Zool. Bot. Ges. Wien
xxvi. p. 296 (1876) (Surinam ; partim) ; Butler, Auu. .Mag. N. II. (4). xx. p. 127. n. 60 (1877)
(Uoayali) ; id., Trans. Eut. Soc. Lond. p. 146. n. 228 (1877) (R. Maui's, May ; Uraria, May ;
R. Negro, near Manaos, June) ; Obertli., Et. d!Ent. iv. p. 66. n. 181 (1880) (partim ; Obydos ;
Cayenne) ; Hahuel, Iris iii. p. 250 (1890) (Mau(>s) ; id., I.e. p. 283 (1890) (Pebas) ; Sharpe, Proc.
( 718 )
>.ool. Sue. Loud. p. 555. n. 5 (1800) (R. Araguaya) ; Godm. & Salv., Biol. Cmtr. Amcr., /?/-/).
p. 213. n. 39 (18110) {/larlliii); iid., iu Whymper, Amiex of Eqnalor, App. p. 109. n. 'J3 (1891)
(Nanegal) ; Michael, Irin v. p. -lU (1894) (Sao Paulo de Olivenva).
PapiUo arrhe^ilatiK, Staudinger («<;« Feldcr, 18(i.i, err. det.), Kj-ot. Tiiyif. i. t. 12 (1884) (Amazons).
Pajiilio /irotesilaiis var., Staud., I.e. p. 17 (1884).
Fajiilio prole.Hilaus protesilaiis, Eimer, A/ib. Vencaiullsch. Sclini. p. 104 (1889) (Amazons; Peru;
"Brazil ' alia s'lb-p ).
Pajiilio pciilhfsilaus, Godm. & Salv. {non Felder, 1805, err. det.), Riol. Ceiilr. Amer., lihop. p. 214.
n. 40 (1890) { parti o, ; Peru).
Papilio matro.<ii!aux, Weeks, Ilhistr. Diurii. Lep. p. 20 (1905) (Chulumaui).
This form is imlividuall}' so variable in every locality that none of the slight
differences found in specimens from diflferent districts appear to hold good. On the
whole the median band of the bindwing below is more pro.ximal in the individuals
from the Guianas and the Lower and Middle Amazons than iu the Andesiaii
sj)cc'imcus. In many of the Andesian individuals the frons is nearly or ipiitd us
extended brown as in the Brazilian subspecies.
The first and second bands of the forewing reach nsually beyond SM-, some-
times extending to the hindmargin, bnt there occur also specimens in which both
l)!inils stoj) short at SM-. The fourth band extends mostly across two-thirds the cell,
often nearly attaining the hindmargin of the cell, sometimes being reduced to a
small costal spot. The cell-bauds are usually rather heavy in Guiana specimens, the
fonrth reaching often nearly or entirely across cell. The transparent submarginal
intersp.ace bears in costal third, sometimes nearly all over, brown scales instead of
white ones, these brown scales occnpying on the whole a larger portion of the
transparent space than in the preceding geographical forms. The position of the
median band of the bindwing is somewhat variable, the Andes specimens agreeing
in the position of the band and the extent of the red scaling with archcsilaus.
The apex of the cell of one of our numerous Zamora (Ecuador) examples has the
apex of the cell of the hindwing filled in with red scales. In an individual from
Rio Demerara, Britisli Guiana, July 1897, the sixth band (postdiscal) of the
forewing is much shaded with white behind, not joining tiie marginal band.
Genitalia: The har[ie exhibits also considerable variability. In nearly all the
specimens the dorso-apical deflexed ridge is produced into a more or less denticulate
triangular tooth of variable dimensions (fig. 3, p. 708) ; this tooth is occasionally
absent, the ridge being ronnded-dilated proximally and simply serrate. This
reduction of the tooth obtains in our four specimens from Archidona, N.E. Ecnailor,
and in one of our individuals from the Rio Chuchuras, Huanuco, Peru.
Hab. Orinoco ; the Guianas ; Amazons, from Pard to the Andes ; Eastern
Ecuador ; Peru ; Bolivia.
In the Triug Museum 220 Si from: ('aura II., various places, February and
September to November (8. M. Klages) ; R. Demerara, August 1897 ; Aroewarwa
Creek, Surinam, July 19u5 (S. M. Klages) ; R. Uaupes, R. Negro ; Poznzo, Huanuco,
800-1000 m.(\V. HofTmanns); R. Chuchuras, affl. of R. Palcazu, 320 m. (W.
Holl'maniis) ; ( 'hanchaniayo (\V. Hofl'mauns ; Schunke); Palcazu (Sedlmayr);
Cajon, Cuzco, October 1000 (Garlepp) ; Cuzco, March lOOl ((.iarlepp) ; Chirimayo,
S.E. Peru, 1000 ft, July 1901, dry season (G. Ockenden) ; Montanas, Madre de Dios,
September 1901 (Ockenden) ; R. Slucuri, 2r)ll() ft., June 1901 (Ockenden); Salinas,
R. Brni, Bolivia, July ISOO (Stuart) ; Salampioni, Bolivia,, S(mi m., September 19o0
(Simons) ; ( 'liavujilaya, 1300 m., June 1901 (Simons) ; Mapiri ; S. Jose de Chiquitos,
East Bolivia, July 22. 1904 (J. Steinbach).
( 719 )
y. P. protesilaus nigricornis Stand. (1884).
rapiUit protesilaiis, Godart, I.e. (1819) {partim ; Brazil) ; Boisd., I.e. (1836) (partim ; Brazil) ;
Doubl., List Lep. Ins. Brit. Mus. i. p. 9 (1845) (partim; Brazil); Gray, List Lep. Ins. Brit.
.Mus. 1. Pup. p. 45. n. 168 (1856) (partim ; Brazil, Rio de Jan.) ; Mt-nL'tr., £«»;»,. Cor/j. Anim.
J/hs. Primp., Lep. i. p. 3. n. 49 (IS.")?) (Brazil) ; Capronn., .lim. Soc. Enl. Brhj. xvii. p. 8. n 1
(1874) (Botafogo, Oct.) ; Weym., SlHt. Ent. Zeit. I7. p. 315. n. 15 (1895) (partim ?) ; Boiiningh.,
Verh. Ver. Nat. Vntcrh. Ilamhurg ix. p. 27 (1895) (Rio, rare; Petropoli.s more common);
Peters, lUustr. Zeitschr. Ent. ii. p. 52 (1897) (Nova Frilurgu).
Priilrsilnns leitiis Swainson, Zunl. Ilhistr. ii. t. 93 (1832) (fiij. .iiiper. ; new name for jinilrsilans L.).
Papilio protesitaus var. nigricornis Staudinger, Exot. Tag/, i. p. 17 (18.H4) (S. Paulo Brazil).
Papilin 2'rntesihms protesilaus, Eimer, .Irtb. Verwandtsrh. Hrhm. p. 104 (1889) (/«o7/«! ; Brazil).
Swaiiisoii's fignro was taken, we think, from a Brazilian specimen, judging
from the cohinr of the antenna and frons. Hi.s name leilus, however, was proposed
to replace Linnd's name pi-otesilatts, this latter term being employed by Swainson for
what he called a subgenus. As leilus Swains, is, therefore, nothing more than another
term for jivotesilaus, it is a synonym pnre and simple of protesilaiis, and cannot be
accepted as a name for the particular Brazilian form which Swainson figured.*
S ? . Antenna brownish black, seldom tawny. Frons brownish black, the
sides not being creamy white ; this character often met with also among P. prot.
protesilaus from the Andes.
Genitalia : c?. Deflexed dorso-apical ridge of harpc dilated into a large
denticnlate tooth, which is much larger than in the other subspecies ; central
process sjiatnlate, denticnlate at apex ; ventral jirocess more or less denticulate.
IJab. East Paraguay ; Brazil.
In the Tring Museum 30 <?c?, 5 ? ?, from : Yhu, East Paraguay, December
1896 (Andeer) ; Minas Geraes, February 1901 (A. Kennedy) ; Tijuco ; Petropolis ;
Eio de Janeiro (E. May); Nova Friburgo; Leme, Sao Paulo; Parana; Leopoldina,
• S. Catharina ; Blnmenaii.
Together with P. protesilaus 7iigricornis in the same districts and at the same
season there occur specimens which differ very remarkably in several jioints.
Two of these specimens from the Province of Parana agree with P. protesilaus
nigricornis, except in the hindwing being somewhat dift'ascd with j'ellow, the
admarginal and submarginal spots R- — M- being more or less distinctly yellow.
In several other individuals from Parana the yellow diffusion is present in
both wings, being especially distinct on the underside of the hindwing, the forewing
is practically naked from M- forward, the postdiscal band of the hindwing is
broad, etc.
We thought at first that these yellowish individuals were the product of
crossing between P. protesilaus nigricornis and /'. telesilaus or stenodes/nus. But
as they do not combine the characters of these species, standing for instance in
the broad, straight, black, postdiscal band of the hindwing quite outside the limits
of variation of these species, and as they differ also in the genitalia, we can but
treat them as belonging to a distinct species, described below. That there are
several closely allied species in Brazil occurring in the same district is nothing
unusual ; but the question of specific distinctness is in this case rendered very
difficult to decide from a small series of specimens, since there occur individuals
which stand just intermediate between P. jirotesilaus ni//ricornis and the new
species. These individuals have the forewing practically naked from M" forward,
have yellow ad- and submarginal spots on the hindwing, etc., as in the new
* ,See p. 418.
( -20 )
species, bnt differ from the uew species in the harpe being practically like that
of ni(jriconus. Moreover, in some of the characters — for instance, the development
of the black postdiscal band of the uppersidc of the hiudwing, and the black line
along the abdominal edge on the underside of this wing — the specimens incline
decidedly towards nigricornis. The teeth of the hindwing are, however, on the
whole rather more acute than in either nigricornis or the new species. In one of
these individuals, from !Minas Gcraes, February, the antennae are tawny, not black ;
a second specimen from Minas Geraes, August, has unfortunately only a remnant
of one antenna, which is black. What are these specimens ? Their characters
are such that the individuals may be hjbrids between the new species and
nigricornis. Besides tlie two Minas Geraes specimens we have one from Castro,
Parana, September, and two labelled simply " Brazil," all males. Is it likely tliat
we should have received five hybrids, while we got only sis specimens of one of
the supposed parent species ? Moreover, the tawny antenna of one of the Minas
Geraes specimens could hardly be explained by assuming the specimen to be the
product of two parent-species which have both black antennae. It would be
necessary to assume further that in this individual there was also the blood of
P. telesilaus, which has a tawny antenna. We are inclined to believe that the
specimens belong to a third species ; they may even be the Atlantic representatives
of the insect figured PI. VI. fig. 29, tliough they resemble in colour much more
PI. VI. fig. 30. We purposely abstain from giving a name to these individuals,
believing it to be much wiser to wait till a larger material has been examined.
One necessary desideratum is also the knowledge of the extent of variation of
P. protesilaus tugricor/tis during the dry and wet seasons. After our discovery
of a slight but distinct difference in the genitalia of the seasonal forms of Papilio
xuthus,* it would not be very astonishing if some such variability should be proved
by breeding to obtain also in P. protesilaus. The difference in the density of the
scaling of the forewing is a character known to vary with the season in P. podalirius
and allies.
152. Papilio helios spec. nov. (PL VI. fig. 30).
(? ? . Antenna and frons black as in P. protesilatis nigricornis. Wings more
or less pervaded with yellow. Forewing transparent, the scaling of the upperside
dense only from hindmargin to M-, but even in this area the scales somewhat
reduced in size, not covering one another as in P. protesilaus, either rounded at
apex, entire, or obtusely bidentate ; wliite scaling between M- and costal margin
more or less completely fallen oif, the scales narrow, those in the distal intersjiaces
between the black bands rather broader than in P. protesilaus ; first band stojiping
short at SM^, there being only a very few black scales behind this vein ; second
band mostly reaching beyond SM", but not attaining hindmargin, interspace between
these bands wider than that between second and third band ; fourth band short
or vestigial ; fifth band not narrowing to a point at lower angle of cell, postdiscal
band closely approaching lower angle of cell, posteriorly almost separated from
tbe marginal band, the white interspace M-— (SM') being obscurely continued to
SM-. Hindwing more sharply dentate than in P. protesilaus nigricorni.'<.
especially at C, SC^ and W ; black postdiscal band with more or less straiglit
jiroximal edge, the partitions E- — M' of this band not separate from one another,
larger and proximally more sharjily defined than in P. protesilaus, red anal spot
ZeitscJir. ]Viss. Zool. Ixsxiii. p. 17'J (1905).
( 721 )
reaching to M', the black sput M' — M- in front of it more or less distinctly
connected with the black postdiscal band, and the white spot M' — M- behind the red
one rcdnced, often to a minnte dot; snbraarginal and marginal sjiots from R- back-
wards yellow, marginal spots somewhat paler, yellow spot at anal angle not divided,
the vein M^ traversing it being distinctly black only proximally ; black snbmarginal
bar R- — R' straight as a rnle, the yellow ad- and snbmarginal sjiots R' — M' not
regnlarly crescent-shaped, the snbmarginal one irrognlarly triangnlar or trapeziforra ;
most specimens with a black bar behind M^ in front of the red anal spot.
Underside more yellow than npper. Forewing practically naked from M-
forwards ; the small costal spot between discocellnlar band and postdiscal one
separated from the latter by yellowish scaling, the extreme costal edge remaining
black ; scales of hinder area entire, not touching one another. Hindwing :
subbasal and median bands a little more widely apart at costal edge and the
black postdiscal band broader, therefore the discal area proportionately narrower
costally than in P. p. iiigriconiis ; red line with distinct white border on distal
side ; thin black snbmarginal bar W — M' incomplete, not reaching R^, yellow
anal spot not separated from sjjot M' — M^, or incompletely, M- being only partially
black between these spots or not at all ; black arrowhead-shaped spot which stands
proximally of red anal bar very little produced basad, being smaller than in
P. protesilaus; a distinct black line on abdominal fold of male extending from
base at least two-thirds to anal angle, this line absent or vestigial in P. protesilaus.
Scent-organ as in P. protesilaus.
Genitalia : S. Detlexed dorso-apical ridge of harpe serrate proximally, not
dilated into a large tooth, apex of harpe tapering almost to a point, tip of central
process curved proximad in dorsal aspect, teeth vestigial ; ventral process reaching
edge of clasper, non-dentate.
Length of forewing : c?, 42 mm. : ? , 4.5 mm.
Hab. Brazil.
In the Tring Museum 5 S <S , 1 ?, from: Castro, Parana, December 1898
(E. D. Jones) ; Parana, name-type.
The white distal border to the red median line of the hindwing below is
a remarkable character, which is vestigial in the allied species P. protesilaus,
P. telesilaus, and P. stenodesmus, being more distinct in P. stenodesmus than in
the others. The white border is best noticed in these species if one looks at
the hindwing from the upperside, with the eye between specimen and light.
153. Papilio orthosilaus Weym. (1899) (PI. YI. fig. 29).
Papilio nrlhosilaus Weymer, Eiit. Nachr. xxv. p. UI5 (1899) (Paraguay).
c?. Similar to P. kelios, larger. Antenna tawny, not black. Frons broadly
buiF at eyes, not all black.
Wings somewhat narrower than in P. helios. Upperside : forewing trans-
parent as in P. helios ; first and second band continued to hindmargin, third
band extending just beyond M, the vein itself being black and there being some
black scales distally of the vein, fourth band vestigial, discocellnlar band as in
P. helios, postdiscal baud not so close to lower angle of cell as in P. helios,
posteriorly a little wider than in that species. Hindwing : black median band
incompletely represented also above from near costal margin to M', a black half-
crescent inside the apex of the cell connected with this band ; postdiscal band
( 722 )
broad, much broailer tban in P. helios, aJmargiual and snbmarginal spots yellowisb,
but paler than in P. helios, all small, j'ellow aiial marginal spot separated from
snbmarginal spot M' — M-, the vein M'- being black ; red anal spot ver)' large,
yellowish spot M' — M- behind it reduced to a minute bar, black spot M'— M'
contiguous with the red patch ; blue spots R' — M- rather large, but not sharply
defined i)ro.\imally ; marginal teeth C — ]{-' sharp, distal edge of wing black from
C to tail, there being only a minute yellowish spot close to the margin in front
of tail.
Underside more green at base than in P. helios ; anal and subanal pinkish
spots of hindwing large ; black line along abdominal edge extending to anal
angle, postdiscal band broad, yellowish ad- and snbmarginal spots narrow.
? not known.
Scent-organ as in P. protesilaus.
Genitalia as in P. helios, but apical lobe of harpe much shorter and broader,
rounded at apex ; ventral process also shorter.
Ilab. Paraguay ; Brazil.
A c? in coll. Weymer, from Paraguay. The type-specimen, also from
Paraguay, is no longer in coll. Fruhstorfer; we have been unable to find out in
which collection it is now contained ; the bands, especially the marginal one of
the forewing, are narrower than in our iigiirc, according to a sketch which
Herr G. Weymer has kindly sent us.
The 6 here figured is from Goyaz, Brazil, in coll. Obertliilr.
154. Papilio stenodesmus sjiec. nov. (PI. IX. fig. (58).
cJ ? . Antenna Idack. Frons wliite at sides. Transparent area of forewing
much more extended than in P. protesilaus and P. telesilaus, the opaque (densely
white-scaled) area reaching from hinder margin to M-, not entering cell ; the scales
of this opaque space narrow, sinuate, becoming more and more narrow towards
cell and in basal half of cell, assuming a hairlike shape in the apical half of
the wing ; the white scales at the costal edge between the black bands about the
same size as those in the posterior area of the wing ; the hairlike scales of
the transparent spaces usually fallen off to a great extent; the scales of the
black bands about twice or three times as long as broad, denticulate, some
narrow; scaling of rmderside as above, but transparent spaces practically denuded
of scales, the sockets of the scales, however, being present ; scales larger than
above and entire from M^ to inner margin, except distally ; seven comparatively
thin black bands, the first not quite reaching inner margin, stoi)ping at vSM',
the second extending usually beyond SM-, but never reaching inner margin, a little
more oblique and therefore posteriorly more distal than in telesilaus ; interspace
between these two bands about half as wide again as that between second and
third band, the latter stopping at M, fourth band reduced to a costal spot, usually
triangular, rarely reaching halfway across the cell, fifth and sixth bauds usually
touching each other at lower angle of cell, seldom here li mm. distant, sixth
always reaching to hinder angle, here joining the marginal band; transparent
snbmarginal band reaching costally beyond SO' and posteriorly beyon<l M", there
being in black costal border a thin transparent streak before SO''. Hindwing
more elongate than in P. protesilaus and P. telesilaus, often slightly yellow ; the
marginal teeth R-, M' and M" longer, snbmarginal spots R- — M' pale yellow, paler
( 723 )
than in P. telesilaus ; medium band of underside a little curved, crossing cell
closer to apex than in P. protesilaus and P. telesilaus.
Nenration and scent-organ as in P. prutesilaiis.
Genitalia : S . Tenth tergite narrow, trilobate at apex, a little longer than in
P. pi-otesilaus ; apical lobe of harpe broad, obtuse or pointed, about two and a half
or three times as long as broad in middle, finely denticulate at convex apical edge
and in middle of ventral margin, deflexed dorso-apical edge not dilated ; ventral
process reactiing ventral margin of clasper, central process pointed in dorsal view,
the tip being carved proximad. — — ? not dissected.
Early stages not known.
Ilab. Paraguay ; Brazil.
A combiuation of black antenna, white (or greyish) sides of the frons, broad
interspace between the first and second black bands of the forewing, and a strongly
dentate hindwing is not met with in the allied species. The frons of the Brazilian
P. protesilaus, which has black antennae, is nearly as black at the sides as in the
centre ; and P. telesilaus, small specimens of which resemble P. stenodesmus, has
always tawny antennae.
We have seen specimens said to be from outside the range as given above.
But as they were obtained from a Continental dealer, we do not believe that the
localities are authentic, and therefore abstain from publishing them.
In the Tring Museum 68 c?c?, 1 ? , from : Sapucay, Paraguay, August to
October 1901 and 1904 (W. Foster), tijpe ; Yhu, Paraguay, September — December
1896 (Andeer) ; Rio de Janeiro (E. May); Petropolis, December 1897 (Foetterle) ;
Lenie, S. Paulo, February and March 1898 ; Bahuru, S. Paulo (Dr. Hempel) ;
S. Paulo, August 1884 (E. D. Jones) ; Castro, Parana, September 1898
(E. D. Jones) ; Espiritu Santo, September 1896 (Foetterle) ; S. Catharina ;
Blumenau.
155. Papilio earis spec. nov. (PI. VI. fig. 32).
c?. Frons huffish white at sides. Antenna dark tawny. Abdomen beneatl'
more or less ochraceous. Wings pervaded with buff, especially on the underside,
which is mostly washed with pink.
Forewing : first band stopping at SM", second prolonged a little beyond that
vein or also stopping short at it ; interspace between these bands a little narrower
than the interspace between second and third bands ; fourth band reduced to a
costal dot ; sixth close to lower angle of cell, narrower from R- forwards than
backwards. Hindwing : red anal spot elongate, extending to near M' proximally,
black bar in front of it restricted to cellule M'— M"; black postdiscal spots R- — R'^
luniform, narrow, more or less distinctly separate from spot 11^ — M* ; ad- and
submarginal interstitial spots huffish, narrow ; dentition of wing rather more sharp
than in P, prot. protesilaus.
Underside : ad- and submarginal huffish spots of hindwing smaller than in
P. p. protesilaus, upper two more or less washed with brown, submarginal huffish
bar 11' — R- very thin ; white distal border of red anal bar wider than in the form
just mentioned, the black arrowhead spot on abdominal fold not reduced as in
that form.
Scent-organ as in P. protesilaus.
Genitalia: Deflexed dorso-apical ridge of harpe .^lightly widened, with some
teeth ; ventral process simple, not leaching ventral edge of clasper, somewhat
(724)
curving apicad ; central process broad, short, spatnlate, heavily dentate, the teeth
somewhat curving basad, the process bearing one or two teeth on the ventral
side near the base.
? and early stages not known.
Ilnb. Ecuador : Zamora, 3o00— 4000 ft. (0. T. Baron), ti/pe ; Zarnma,
1000 m., June 1899, wet season (Simons); three S3 in the Tring Museum.
Judging from the genitalia, it appears to us jiossiblethat P.cnris, P. orthosilaus
and P. helios may turn out to be geographical races of one sjiccics. However, the
materia] examined is as yet quite insufficient to decide the question.
150. Papilio telesilaus Feld. (18G4).
Pnpilio Eques Acliivtis protesilmis, Jablonsky & Herbst (iioii Linn^, 1758, err. det.), Xnttirs. Schm.
iii. p. 147. n. 97. t. 43. fig. .3. 4 (1788) (" Carolina & Jamaica," errore).
Priiirejjs heroicus jn'rilesilaun, Hiibner, Samntl. E.rol. Srhm. i. t. 108 (180G — ?).
Iplih-lkkx prolesUauK, id., Verz. hel: Sehm. p. 82. n. 834 (1818?)
Papilio protesilaus, Godart, Em: AtHh. ix. p. 50. n. 73 (1819) (partini).
Papilio telesilaus ¥e\der, Verh. Zool. Bot. Ges. M'ioi xiv. p. .'iOl. n. 179 (18(54) (Amazonia; Nova
Granada ; Brasilia austral.).
(?¥. Body and wings more pervaded with yellow than in P. protesilans.
Antenna always tawny, though varying in depth of colour. Frons always
3-ellowish white at sides. Forewing less densely scaled in costal area than in
South American P. protesilaus, the small scales falling easily otf, tlie anterior
aud discal portions of the wing appearing naked, glossy ; first and second bands
closer together than second and third in nearly every specimen ; fourth band
sometimes reaching as far as third cell-fold, usually very short, often reduced to
a dot, not rarely absent ; sixth band close to lower angle of cell, which it touches
in many specimens, more or less reduced in width from this angle of cell forwards,
this costal portion sometimes practically separate from the posterior ]>ortiou, the
latter then being continuous with the discocellular baud. Hindwing : submar-
ginal spots R- — M- huffish yellow, often the whole anal region strongly pervaded
with yellow above and below ; blue admarginal scales entire, or some with small
teeth.
Scent-organ : Lair-scales long, but decidedly shorter aud broader than in
P. protesilaus.
Genitalia : $. Deflexed dorso-apical ridge of harpe not or very little dilated,
not enlarged into a tooth ; ventral process reaching ventral edge of clasper ;
central jirocess either simple or bearing at the base on the ventral side a dentate
ridge, the apex being in this case also strongly dentate. ? not dissected.
Early stages not known.
Ilah. Panama to Southern Brazil and Paraguay.
a. P. telesilaus dolius subsp. nov.
Papilio protesilans var. macrosiluus, Bates (inin Gray, 1852, err. det.), Proc. Zool. Soc. Loiicl. p. 241.
n. 2 (18G3) (Panama).
Papilio telesilaus, Godm. & Salv., Biol. Ceutr. Amei:, liliop. ii. p. 215. n. 41 (1890) (Panama;
" S. America " alia sub.«p.).
<J. First band of forewing not continued along SIP to hinder margin, stoj>ping
short at SM-. Friuge of hindwing more extended white between C and \i" than
in the following form, black postdiscal Innules 11^ — M' on the whole smaller, the
blfick submarginal bar K- — 1\' reaching H'' halfway between the black postdiscal
( 725 )
and inarginul liumles R' — BI' ; sulibasa! li;uid (if underside not untei'ing apex of
basal cullule.
Genitalia : Deflcxed dorso-apical ridge of liarpe very narrow, bardly at all
dentate ; central process slender, without basal dentate ridge, apex often entire,
non-dentate, never so strongly dentate as in the following form.
Hab. Panama and west coast of Colombia ; name-type from the Rio Dagiia,
West Colombia.
In the Tring Museum 14 cj<? from : Panama (Salviu) ; R. Dagua (W. F. H.
Rosenberg).
b. P. teleslhms telesikms Feld. (1864).
Papilio Eqtics Acliirus pi-otcsilaus, Jablonskj & Herbst (iimi Linn«, 1758, err. det.), I.r.
Prhiceps heroicus jjrotesllaus, Hubner, Ix.
Papilio protesilaus, Godart, l.C- (1819) (partim) ; Kollar, Denksrhr. K. Ak. Wiss. Wien, Math. Nut.
CI. i. p. 5b'2. n. 2 (1850) (syn. partim) ; Moschl., I'erh. Zoul. Bol. Ges. Wien xxvi. p. 296 (187G)
(Surinam ; partim); Auriv., K. Si: Vet. Akail. Ilnndl. xix. 5. p. 29. n. 28 (1882) (Recensio
critica ; sub synon.).
Papilio jirolesilaiis var. a., Gray, Cat. Lep. Ins. Brit. Miis. i. p. 34. sub. n. IGO (1852) (Brazil).
Papilio telesilaus Felder, I.e. (18(54) (Amazonia ; Nova Granada ; Brasilia austral.) ; Druce, Pror.
Zool. Soc. Loiifl. p. 245. n. 13 (187C) (Ucayali) : Staud., Jixot. Ta(jf. i. p. 17 (1884) (Amazons) ;
Elmer, Arth. Verwandtscfi. Sclim. p. 104. t. 1. fig. 0 (1889) ; Hahuel, Iris iii. p. 2.50. 253
(1890) (Maues) ; id., I.e. p. 283 (1890) (Pebas) ; Michael, Iris v. p. 214 (1894) (Sao Paulo
de 01iven9a) ; Eimer, Urthngeii. p. 21 (1897).
Papilio protesilaus, Linn., var. b. P. telesilaiis, Kirby, Cat. Diurii. Lep. p. 556. sub n. 218 (1871).
Papilio protesilaus var. (etevVau.^, Oberthur, K. iVEiit. iv. p. 6G. sub n. 181 (1880) (Pard ; Cayenne);
Miischl., Verh. Zool. Bot. Ges. II'/c;; xxxii. p. 304 (1883) (Surinam ; in one specim. red anal
spot absent from upperside); Maass. & Weym., in Stubel, Reisen S. Amer., Lep. p. 24. sub
D. 110 (1890) (west side of Cordillera of Bogota); lid., l.c. p. 82. n. 51 (1890) (tipper Amazons).
Papilio protesilaus telesilaus, Eimer, I.e. p. 47. 48 (1897).
Cosmodesmus telesilaus, Kirby.in Huhv., Saiiiml. E.cot.8chneU. ed. ii. p. 93. t. 108. fig. 1. 2 (190— ?).
S ? . First band of forewing continued along SM^ ; subbasal band of hindwing
below entering basal cellule.
The sjaecimens from Brazil have on the whole thinner black bauds than the
individuals from the Andes, Amazons and Gnianas, the median band of the
hindwing is often a little more distal, and both wings are more frequently con-
spicuously pervaded with yellow.
There is considerable variability in the width of the wings as well as in
the size of the markings in the distal area of the hindwing.
Genitalia : Defiexed dorso-apical ridge of harpe distinctly widened, dentate ;
central process S[)atulate, dentate, the teeth standing on the dorsal side, and being
more or less curved, a conspicuous dentate ridge on the central j^rocess at its base.
Ilab. Colombia : Magdalena valley eastwards ; Venezuela ; the Guianas ;
Amazons ; Eastern Ecuador to Bolivia ; Paraguay ; Brazil as far south as Rio
Grande do Sul.
In the Tring Museum 110 Jt?, 2 ¥ ?, from : " Bogota " ; Villavicencio to Rio
Ocoor, January 1897 (Dr. Biirger) ; Temblador, Suapure, and La Union, Caura R.,
Orinoco, February, June, September and October (S. M. Klages) ; R. Demerara,
August 1898 ; Esscquibo R. ; Aroewarwa Creek, Maroewyni valley, Surinam, June
and July 1905 (S. M. Klages) ; Manicore ; Thomar ; Obidos; R. Uaupes, Upper
R. Negro ; R. Napo (W. Goodfellow) ; Archidona, N.E. Ecuador (W. Goodfellow) ;
R. Cachyaco, affl. of R. Huallaga (Stuart) ; R. Chuchuras, affl. of R. Palcazu,
320 m. (W. Hoffmanns) ; Palcazu (Sedlmayr) ; Pereue R. ; Prov. Sara, S. Cruz
de la Sierra, February — April 1904 (J. Steinbach); S. Jose de Chiquitos, East
( 726 )
Bolivia, Jnly 1004 (J. Steinbach) ; Sapncay, Paraguay, September 1902 (W. Foster) ;
Yhii, East Paragnay, December 1890 (Andeer) ; Villa Maria to Piamaiitino,
]\Iatto Grosso, Jannary 1897 (Andeer) ; Minas Geracis, February 1899 and 19ul
(A. Kennedy) ; Petropolis ; Rio de Janeiro (E. May) ; Bahuru, S. Panlo (Dr.
Heiup.'l) ; S. Catliariiia ; Bhimeuau.
XVI. Tliyastes Group.
This gronp is very closely related to the following one, the two together con-
trasting rather strongly with the previons gronp in pattern as well as structure.
The genitalia of the Thi/astes and Dolicaon Groups are practically the same, the
differences between the species being very slight in these organs. The females
are even rarer in collections than those of the I'rotesilaus Group.
Six species :
a. Hindwing below with a tawny band on disc . . . Species No. 157.
Hindwing lielow with a line of red spots on disc . . b.
b. Forewiug with row of sharply marked subraarginal spots
in apical area ........ c. '
Forewing without sharply marked snbmarginal spots in
apical area . , Species No. 162.
c. Hindwing, on npperside, with a yellnw spot i\l' — M-
beside the marginal anal spot Species No. 158.
Only the marginal spot present d.
d. Pale discal area of forewing not reaching to lower angle
of cell Species No. 159.
Pale discal area of forewing reaching to lower angle
of cell e.
e. Snbmarginal spot R'^— R^ of forewing distant from discal
area, or absent Species No. 100.
Snbmarginal spot R- — R^ of forewing contiguous with
spot R^ — R' of discal area, always pre.-;ent . . . Species No. 161.
157. Papilio marchandi Boisd. (1830).
Paj>ilio mavchiinili Boisduval, Spec. Gen. Up. i. p. 350. u. 192 (1836) (Mexico) ; Strecker, Lep.
Rhop. Uet. p. -25. t. 4. fig. 2 (1873) (Panama ; Costa Rica ; Hondura-) ; Godni. * Salv., Biol
Centi: Amrr., lilinp. ii. p. 218. n. 4i;. t. G8. fij. 5. liaipe (IH'.lO) (Ea«t and Soutli Mexico;
Guatemala; Brit. Honduras; Costa Rica; Panama; Colombia); Haase, Uiitn-siich. Mimicry
i. p. 83 (1893) (Colombia ; Guatemala).
S ?. Instead of the discal row of red spots on the underside of the hindwing
of the allied species P. marchandi bears a tawny band, the wing being more or
less washed with the same colour dislally of the black subbasal band and along
the abdominal fold. The tail is broadly bordered with yellow from base to tij) on
hindside. Individual variability obtains on the forewing especially in the size of
the yellow snbmarginal spots and in the upper discal spots, of which spot R'— R-
is usually absent or vestigial, and on the hindwing in the size of the black bands
of the underside. Some of the specimens from Ecuador and Colombia e.vhibit an
interesting character on the upjierside of the hindwing. The ujiper three yellow
bars belong to the admarginal series, the fourth spot, which stands on a level
( 727 )
with the third, belongiug to the submargiual series, lu the specimens ri'fL'r?-oi:l to
there are from one to three snbmarginal bars between C and R-.
Scent-organ : wool brown ; scales beneath the wool somewhat elongate,
narrowed towards apex, mostly trnncate or feebly bidentate, partly entire.
Genitalia : c?. No ventral process on valve ; central ridge of harpe widest
above, obliquely trnncate, without a row of toctli across its lateral surface.
Early stages not known.
Ilah. Ble.xico to West Ecuador.
Two sulispecies.
a. P. marchaiidi mtirchandi Boisd. (1836).
Papllin mnri-hamU Boisduval, /.c. (Mexico); Doubl., List Lep. Ins. Brit. ifus. i. p. 16 (1S45)
(" Brazil," errore) ; id., Westw. & Hew., Gen. D'nirii. Lep. i. p. 17. n. 164 (1840) (Mexico) ; Gray,
Cat. Lep. Ins. Brit. Mu.s. i. Pap. p. 38. u. 18f) (1862) (Guatemala ; " Brazil," errore) ; id., List
Lep. IiiK. Brit. Mk-i. i. Pap. p. 52. n. 197 (1856) (Guatemala) ; Weidem., Proc. Ent. Soi: Phdad.
ii. p. 147 (1863) (Mexico; Centr. Amer.) ; Felder, Verh. ZooJ. But. Ges. Wien xiv. p. 300.
n. 172 (1864) (Guatemala) ; Lucas, Aim. Sne. Kiit. France, p. 532 (1860) (Cordoba, Orizaba;
Guatemala); Boisd., Gmsiil. Lep. G-i item. p. 5 (1870) (Honduras; Mexico); Kirby, Ou.
Diurn. Lep. p. 555. n. 242 (1871) (Guatemala) ; Strecker, I.e. (jiarlim ; Horduras) ; Kirby, I.e.
p. 811. n. 242 (1877) ; Oberth , El. (lEnl. iv. p. 75. n. 236 (1880) (Mexico) ; Staud., Exot. Tugf.
i. p. 18 (1884) (Centr. Amer.) ; Godm. & Salv., I.e. (peirtim ; East & West Mexico ; Guatemala;
Brit. Hondura.s) ; Haase, I.e. {pariim ; Guatemala).
(? ? . The tawny discal band of the underside of the hiudwing is usually
somewhat curved anteriorly ; its black or brown proximal border is narrow as a
rule, bat is sometimes as broad as, or even broader tlian, the lawny band between
SC'' and R^ ; the i)ale central area is usually rounded off. The submarginal spot
W — R'on the upperside of the hindwing is small in most specimens, the admarginal
bar at its outer side is vestigial or absent, and the submargiual spot R' — M' is small
or absent.
Genitalia : Apical edge of harpe with a few more teeth than in the following
form.
Hab. West and East Mexico; Giiatemahi ; Brit. Honduras; "Hondnras"
{teste Boisduval) ; presumably also in Nicaragua.
In the Tring Museum \\ S S from : Orizaba (Bilimct) ; Guatemala (Salvin).
h. P. inarcliandi pauamensis Oberth. (lS8u).
Papiliii marehandi ■va.v. jiananiennis Oberthiir, Et. d'Ent. iv. p. 75. sub n. 236 (1880) (Panama).
Papilin ntarehamli, Strecker, I.e. [parti m ; Costa Rica ; Panama) ; Butl. & Druce, Proe. Zool. Soc.
Loud. p. 365. n. .374 (1874) (Costa Rica) ; Godm. & Salv., I.e. (partim ; Costa Rica ; Panama ;
Colombia); HviSise, I.e. { partim ; Colombia).
cJ. The yellow markings of the upperside ou the whole paler, but sometimes
deeper yellow, than in certain Central American specimens (does the colour darken
with age?); the yellow submarginal spots R'^ — M' of the hindwing larger, spot
R' — M' being always distinct and the admarginal bar R- — R^ never (?) absent;
the yellow discal band of the forewing dtejier incised ou the veins above and below ;
the black bands of the underside of the hindwing rather wider, the tawny discal
band more straight, the pale central area less rounded distally, and the wing
narrower between R' and R-, the distance from apex of cell to tip of R' being
somewhat shorter in P. in. panameiisi.'i than in /'. m. marcliamIL None of these
characters are constant, the two forms completely iutergrading.
Hab. Costa Bica to West Ecuador.
( 728 )
In the Tring Museum 20 c? t? from : (!arillo, Costa Rica, 3000 ft., October
1904 (A. Hall); Chiri.ini; R. Dagna, West Colomliia (Rosenl)erg) ; "Bogota";
Pambelar and Paramba, West Ecuador ; Cachabi, W. Ecuador, low country,
January 1897 (Rosenberg).
158. Papilio thyastes Drury (1782).
Papilio Eques Achiims thyantes Drury, IHiistr. ExnI. Ins. iii. p. 47. t. 35. aud In<!i:r (1782) (Rio de
Janeiro) ; Fabr., Eiil. Si/sf. iii. 1. p. 2C.. n. 77 (1793).
Iphiclhks diajjhorus Hiibner, Samml. ExnI. S'-lun. ii. t. 93 (1820?).
Papilio thi/astc.'<, Godart, Em: Meth. ix. p. 54. n. 83 (1819) (Brazil).
c?. Similar to P. calliste ; ground-colour of npperside varying from maize- to
chrome-yellow ; one red anal spot on hiudwing, a spot M' — M- occasionally
vestigial ; yellow submarginal spot R^ — M' much smaller than spot M' — M-.
Red bars R^ — M- of underside of hindwing broken up into dots. Hairs on frons
short. Yellow dorso-lateral line of abdomen broader than the blaclv lateral line.
Scent-organ : wool buffish, scales beneath it denticulate, mostly broad, those in
front of fold mostly entire, acuminate.
Gen ita lia : . No ventral ijrocess on clasper ; central ridge of harpe almost
square, its ventral angle not produced basad, the tooth at this angle large, conical ;
no row of teeth across the lateral surface.
? and early stages not known.
Hab. Ecuador to Bolivia ; Brazil.
Three subspecies.
The most striking feature in the pattern of P. thi/aMes and P. calliste appears
to us to be the development in different directions of the yellowish ad- and sub-
marginal spots R' — M= in the hindwings of the two species. In P. calliste there
are on the upperside of the hindwing no ad- and submarginal spots M' — M' or only
traces of them, while P. thyastes has a large submarginal spot M' — M^. On the
other hand, in cellnle R' — M' there is in P. calliste a large submarginal spot and
a shar])ly defined admarginal curved bar, P. thyastes bearing in this cellule only
a reduced submarginal spot. A similar contrast is observed on the underside of the
two insects. We find in P. tliyastes between the discal area of the forewing and
the so-called submarginal line a yellowish line which represents the true submarginal
interspaces ; on the hindwing there is a corres2)onding row of spots between 0 and
R-. These markings are entirely lost in P. calliste, while in this species the
admarginal spots R- — M^, which are vestigial in P. thyastes, are large.
a. P. thyastes thijastini(S Oberth. (1880).
Papilio thyastiiiiis Oberthilr, Et. d'Eiit. iv. p. 75. n. 235. t. 2. fig. 3 (1880) (Ecuador) ; Hahnel, Iri.i
iii. p. 282 (1890) (Pebas) ; Michael, ibid. v. p. 214 (1894) (Sao Paulo de Olivenga).
Papilio tlnjasles, Staudinger, Exol. Ta(jf. i. p. 18. t. 12 (1884) (Pebas, tln/tistiiiiis = ihi/itstes) ; Haenscb,
Serl. Ent. Zeitsclir. xlvii. p. 154 (1903) (Arcbidona, 640 m.).
(?. Submarginal spot R2 — R^ of forewing separate from the discal spot; discal
band usually broadly interrupted at R^ ; first submarginal spot absent from n])i)erside
or ill-defined. Ape.x of cell of hindwing black.
In some of our Peruvian specimens the snliapical cell-bar of the forewing
above, and the discal costal bar arc interrupted. The position of the central red
spots on the underside of the liindwing is variable ; many specimens have an
additional red spot situated in the cell. One Peru specimen has the spots buffish
pink instead of liright red.
( 729 )
Hab. North-East Ecuador to East Central Peru ; Upper Amaiioas.
la tke Tring Mnseuna 30 SS from: Coca, R. Napo, May— August 1800
(VV. Goodfellow); E. Chuchuras, affl. of R. Palcazu, 320 m (W. Hoffuiauns) ;
Falcazu (Sedlmayr) ; Pozuzo (W. Hoffmanns) ; Icpitos ; Pebas.
b. P. thyastes zoros subsp. nov.
$ . Like the preceding ; but first submarginal spot of upperside of forewing
sharply defined. In the majority of specimens the upper outer angle of discal
patch R^ — M' of forewing not rounded off, the apex of the cell of hindwiug a little
less extended black than in I', th. tlii/astiims, and the yellow patch at base of cellule
B,3 — M' longer. The individuals of thijastiiuis from the Upper Amazons and the
eastern slopes of the Peruvian Andes completely connect the Ecuador form with
the Bolivian one.
Jlab. S.E. Pern ; Bolivia {name-ti/pe from Mnshav).
In the Tring Museum 10 c?c? from: S. Domingo, Carabaya, 4500 ft.
(G. Ockenden) ; Mushay, R. Beni, March 1895 (Stuart) ; Mapiri.
c. P. thyastes thi/astes Drnry (1782).
PapUlo Equcs Achivus thyasles Drury, I.e. (Rio de Janeiro).
Iphiclidcs diuphorus Hiibner, I.e.
Papilio tJnjastes, Godart, I.e. ; Boisd., Spn-. Geii. Lip. i. p. 349. n. 191 (183(5) (Brazil) ; Doubl., Lhl
Lep. Ins. Brit. Mus. i. p. 17 (1845) (Brazil) ; id., Westw. & Hew., Gen. Diurn. Lip. i. p. IG.
n. 163 (1846) (Brazil) ; Gray, Cat. Lep. Lix. Brit. .!/»». i. Pap. p. 38. n. 188 (1852) (Brazil) ; id.,
List. Lep. Ins. Brit. Mus. i. Pap. p. 52. n. 19G (1856) (Brazil); Mc'niitr., Enum. Corp. Anim.Mus.
Petr., Lep. i. p. 4. n. 61 (1857) (Brazil) ; Felder, Vcrh. Z,ml. But. Ges. Wien xiv. p. 300. n. 173
(1864) (Bras, austr.) ; Butl., Cat. Diurn. Lep. deser. Fair. p. 239. n. 20 (1869) (BrazU) ; Kirby,
Cut. ninrn. Lep. p. 555. n. 243 (1871) (Brazil) ; Oberth., Et. d'Ent. iv. p. 75. n. 234 (1880)
(Brazil) ; Staud., Exot. Tagf. i. p. 18 (1884) (Southern Brazil) ; Haase, Untersuch. Mimlery i.
p. 83(1893).
Iphielides tlnjustes, Kirliy, in Hiibn., Sanunl. Exot. Sehnielt. ed. ii. p. 94. L 307. 6g. 1. 2 (190-?).
S ■ Paler than tlie Amlesian forms, with a sliglitly greenish tint in the yellow
colour ; discal baud of forewing not interrui)ted at R^, or only the vein itself black,
submarginal spot R- — R^ merged together with the respective discal spot, or
(rarely) the two spots separated by a very narrow black line. Black distal border
of hiudwing not entering cell ; most specimens with a minute red dot M' — M- on
npperside, which is rarely marked in the Andesian forms.
Genitalia : Central dentate ridge of harpe smaller than in the other forms ;
dorso-apical ridge more densely hairy on jjroximal side.
Hab. Brazil.
No representative of P. thyaste.i is known from Paraguay, Matto Grosso, the
Middle and Lower Amazons and the Guianas.
In the Tring Museum \Q S S from: Bitalha, S. Paulo; Castro, Parana
(E. D. Jones) ; S. Catharina ; Blumenau.
150. Papilio dioxippus Hew. (Is55).
Papilio dioj-ijijiux Hewitson, Exot. Butt. i. Pap. t. 2. fig. 3. 4. J (1855) (N. Granada) ; Gray, Lji.it
Lep. Lis. Brit. .I/k,?. i. Pap. p. 52. n. 198 (1856) ; Felder, Vcrli. Zool. Bot. Ges. Wien xiv. p. 300.
D. 170 (1864) (Bogota) ; Kirby, Cat. Diimi. Lep. p. 555. n. 240 (1871) (•' var." excl.) ; Hopff.,
Stelt. Ent. Zeit. xl. p. 47. n. 2 (1879) {partin ; N. Granada) ; Oberth., Et. d'Ent. iv. p. 75.
n. 233 (1880) (Muzo ; Carare) ; Stand., Exot. Tayf. p. 17 (1884) ("Poru bis Columbien'
errore) ; Haase, Untersuch. Miinierij i. p. 83 (1893).
( 730 )
Oue of the commonest Papilios in Bogota collections. It is known from the
('auca and Magdalena valleys, and occnrs also on the eastern side of the ('ordilh'va
of Bogota. The insect comes very close to P. lacandones. As we know as yet
nothing of the occurrence of P. lacandones in Colombia, and as dioxippus is
a]iparently restricted to that country, it is not impossible that dioxippus may turn
out to be the Colombian representative of lacandones. However, as the lacandotu's
form which occnrs from Ecuador to Bolivia agrees both in structure and pattern
closely with the Central American form, it is hardly probable that the Colombian
dioxippus, which differs in structure and pattern from both lacandones forms, is a
third form of the same species.
S. Forewiug with two to five submargiual spots ; discal area often reaching
beyond W, but never up to R^ ; the two distal posterior spots in cell usually merged
together, rarely quite separated from one another; the brown excision, at the base of
M', of the yellowish discal area is seldom absent (for instance, in type specimen).
The purplish white submargiual spots R- — M" on the underside of the hiudwing are
always present, the posterior one being, however, often much reduced.
(Scent-organ : wool creamy ; scales beneath it mostly entire, truncate, apically
brcjader than in P. marchandi.
Genitalia : Upper angle of dentate ridge of harpe rounded.
? and early stages not known.
Hab. Colombia : Magdalena and Cauca valleys ; Llanos of Rio Meta.
In the Tring Museum 140 S$ from: '"Bogota"; Muzo, December 1896;
Pereira, Cauca.
In coll. Godman from: Cauca valley (Ribbe); Quiudia, 4000 ft. (Wheeler);
Muzo ; Llanos de R. Meta (Child).
160. Papilio lacandones Bates (1864).
Papilio lacanchnes Bates, Ent. Mo. Mag. i. p. 4. n. G (1804) (Guatemala).
S. Near P. dioxippits Hew. (1855), with which it apparently occurs together
in Colombia. The yellowish discal area of the forewing extends anteriorly to R' or
beyond ; the remnant of the yellowish subapical cell-band stands distally of base
of R^, beiug widely separate from the (tell-spot which precedes it at hinder edge
of cell. The hindwing is proportionately longer than in P. dioxippus, the pale
submarginal spot M'— M- of the underside absent or just vestigial, and the cell
narrower.
Scent-organ as in P. dioxippus ; the white scales beneath the wool and discally
of the fold rather broader and mostly rounded at apex.
Genitalia as in P. dioxippus, but the upper angle of the dentate ridge
acuminate, not rounded.
? and early stages not known.
Hab. Guatemala to Bolivia; two subspecies.
a. P. lacandones lacandones Bates (1855).
Papilio lacamloiiea Bates, (.<■. (Guatemala) ; Godm. & Si\v.,Bio!. Ceiilr. Amcr., IVwp. ii. p. -'If!, n. 43.
t. 67. fig. 11. 12. (J, la. harpe (1890) (Guatemala ; Pauama; ''Ecuador ; Peru" alia subspecies).
Papilio lacandores (!), Felder, Verh. Znol. B„t. Ge.s. Wien xiv. p. 301), n. 16;i (1864) (Guatemala;
cit. falsa).
Papilio dioxippus var. a. P. lacandonei, Kirby, Cat. Diurii. Lep. p. 555. sub n. 240 (1871) (Guatemala).
Papilio dioxippus, HopfEer, Stell. Ent. Zeit. xl. p. 47. n. 2 (1879) (_ parti m ; Guatemala).
( 731 )
Fonr snbmarginal spots on forewing, thin, the first nsnally vestigial ; yellowish
discal area extended beyond R-, spot R" — R^ trnncate at R- ; snbapical posterior
cell-spot about 5 mm. long, reaching from R' nearly to angle of cell ; distal marginal
border narrower at M' than yellowish discal area. Yellowish ad- and snbmarginal
spots of hindwing shaded with brown, small; whitish snbmarginal spots R^ — M' of
underside vestigial or very small.
Hab. Guatemala ; Panama.
In the Tring Museum : 2 c? c? without precise locality.
In coll. F. D. Godman a series of males.
rapilio lacamloMs, Godman & Salv., Biul. C'eiitr. Amer., lilmji. ii. p. 216. n. 43 (1890) (partim ;
Cururai, Ecuador, and Cosnipata, Peru).
F. lacandones diores subsp. nov.
Salv., Biul. C'eiitr. Amer., lihnp. ii. p. 2:
lipata, Peru).
c?. Wings a little broader than in I. lacandones. Forewing : five snbmarginal
spots, larger than in the preceding form, sjiot SC — SC^ distinct, spot R- — R^ seldom
vestigial ; yellowish discal area a little straighter distally, slightly incised on tlic
veins, stopping at R-, the spot R^ — R' triangular, not trnncate at R^ ; snbapical
posterior cell-spot small, about as large as snbmarginal spot R' — R-, usually less
than half the respective cell-spot of /. lacandones. Ad- and snbmarginal yellowish
spots of hiudwing larger above and below, especially the whitish snbmarginal spots
R- — M' of underside, upper bars nearer edge of wing than in I. lacandones.
Hab. Ecuador ; Pern ; Bolivia {name-ti/pc from R. Slucnri).
In the Tring Museum \A S S from: C'hanchamayo (W. Hoffmanns); R. Slncnri,
S.E. Pern, 250U ft., June 1901, dry season (G. Ockenden) ; Ghirimayo, S.E. Peru,
Jnly 1901, dry season, 1000 ft. (G. Ockenden); >S. Domingo, S.E. Pern, 4500—
6000 ft., Jul}'— August 1901 (G. Ockenden) ; La Pampa, R. Huacamayo, 2500 ft.,
November 190-1 (G. Ockenden) ; Mapiri, Bolivia.
101. Papilio calliste Bates (18(54).
Papilio mlliste Bates, Ent. Mo. Mag. i. p. 3. n. 5 (1864) (Guatemala).
Papilio lorzae Boisduval, Insect. Agrie. p. 103 (1869) (Guatemala).
c?. Nearest to P. dioxippus and P. lacandones. Hairs of frons long. Pale
dorso-lateral line of abdomen narrow. Pale bands and spots of wings greenish
yellow or primrose-colour. Snbmarginal line of spots of forewing curved, spot
R- — 11^ of this line confluent with tlie respective spot of the discal band. Hindwing,
above, with two red spots, sometimes with traces of a third dot R' — 51' ; while the
ad- and snbmarginal greenish yellow sjiots R- — BI' are strongly developed, there are
only slight traces of such spots between M' and M^.
On the underside of the hindwing the yellowish and whitish spots at and near
the distal edge are merged together, forming a jiale border to the wing ; red spot
M' — M- bordered with white in front like the anal one, not divided into dots.
Sceut-organ as in P. lacandones, a portion of the scales beneath the wool rather
strongly bidentate.
Genitalia : cJ. No ventral process on valve ; central ridge of harpe widest above,
with large teeth at edge, ventral edge not produced basad, no row of teeth across
lateral surface.
Female and early stages not known.
Uah. Mexico to Costa Rica.
Two subspecies.
47
( 732 )
a. P. calliste calliste Bates (1864).
Papilin caUisle Bates, I.e. (Guatemala) ; Felder, Verh. Znol. Bid. Ges. Wieii xiv. p. 300. n. 171
(1864) (Guatemala ; cit. falsa) ; Kirby, Cul. Diurn. Lep. p. 555. n. 241 (1871) (Guatemala) ;
Stand. Exot. Tarjf. i. p. 18 (1884) ("northern Centr. Amer.'') ; Godm. & Salv., Biul. C'eiiti:
Amer., Rhop. ii. p. 217. n. 45. t. 68. fig. 1. 2 (1890) (Atoyae, Mexico; Guatemala ; Brit.
Honduras; — "Costa Rica" alia subsp.) ; Haase, Untersuch. Mimicri/ i. p. 83 (1893)
(" N. Granada " errore ; Guatemala).
Papilio lorzae Boisduval, /.<■. (Guatemala).
(?. Forewing, above : apical cell-band ami the following band clearly marked ;
postmedian cell-band also present, but usually more or less shaded with brown in
middle, while the submedian cell-band is vestigial, being rarely distinct ; discal area
e.Kcised on cell-side between M^ and M-, the sinus rounded, an indication of a sinus
also between R' and M' ; outer edge of discal area slightly irregular ; a distinct
submarginal spot E' — M'. Black distal area of hindwing entering or touching
cell ; yellowish marginal spot M' — M- distinct.
On underside, the submarginal line of forewing continued to SM^.
Ilab. Mexico : Jalisco ; Guatemala ; British Honduras.
In the Tring Museum 15 cJ<? from: Motzorougo, Mexico; Polochic valley,
Guatemala ; " Guatemala."
b. P. calliste olbius spec. nov.
PapH'w raUiste. Butler & Druce, Proc. Zool. Soe. Lond. p. 365. n. 373 (1874) (Costa Rica ; this
form according to specimen in coll. Godman) ; Oberth., Et. d'Ent. iv. p. 75. n. 237 (1880)
(Coita Rica) ; Godm. & Salv., I.e. (partim ; Costa Rica).
S. A little larger than the preceding. Forewing, upperside : submedian cell-band
absent, postmedian one much shaded with brown, only an anterior and a posterior spot
being clearly marked, subaincal cell-band also shaded witli brown in middle ; discal
costal band very thin except costally ; discal area wider than in the preceding
subspecies, its distal edge more straight, the sinus M' — M- on costal side reduced ;
submarginal spot W — M' vestigial. Hindwing : black distal area narrower than
in calliste in front, not widening costad, not touching cell ; black basi-abdominal
band also a little narrower ; yellowish marginal spot M'— M- vestigial.
['lidcrside. — Submarginal line of forewing vestigial from M' to hindangle ; the
bands less distinctly washed with sulphur-yellow than in the preceding.
Hindwing : brown discal band of almost even width from costal margin to M',
not triangular.
Ilab. Costa Rica.
In the Tring Museum 4 c? (^ from : Oachi, Costa Rica, May 1901 (Underwood),
ti/pc ; Carillo, Costa Rica, 3000 ft., October 1904 (A. Hall).
102. Papilio leucaspis Godt. (1819).
Papitio leucfi.y>;s Godart, Eii.: Meth. ix. p. 55. n. 85 (1819) (Peru ?) ; Grimsh., Trans. Roy. Soc.
Edinb. xxxi.\. 1. p. 8 (1897) (" type" from Dufresne collection).
t?. Frons all brown-black, hairs long. Abdomen black above, pale lateral line
Vestigial, buff-yellow or clay-colour beneath. Brown area of wings paler than in
P. dioui'ppu.i, thi/astes, etc. Cell-folds of forewing distinct ; greenish straw-yellow
cell-bars represented by spots situated at costal margin and at hinder side of cell ;
discal area triangular, stopping short at R-, but there is usually a small dot in front
of this vein ; a spot in front of subcostal fork as in P. dioxippus ; brown marginal
area with deep brown or (beneath) black lines which are more distinct beneath than
( 733 )
above, fonr in nnmber, the two middle oues confluent costally. Greenish straw-
yellow area of hiudwiiig triangular, extending beyond apex of cell ; abdominal fold
brown ; a red line from R- to abdominal edge, interrupted at the veins, the line of
spots continued to costal margin on nnderside.
Scent-organ : Wool huffish, scales beneath it denticulate, mostly asymmetrical,
those in front of fold mostly short, truncate, feebly denticulate.
Genitalia : Central dentate ridge of harpe narrowed at base, widest above,
longer than broad, the dentate edge incrassate, bearing conical teeth, no row of teeth
across its lateral surface ; no ventral process on clasper.
? and early stages not known.
Ilab. Colombia to Bolivia.
Two subspecies.
a. P. leucaspis lamis subsp. nov.
Papilio leucaxpis, Hewitson, Exot. Butt. i. Paj). t. 2. fig. 5 (1855) (Colombia) ; Feld., Verh. Zoul. But.
Gets. Wien xiv. p. 300. n. 1(58 (1864) {imrtim ; Bogota) ; Kirby, Cat. Diurn. Lep. p. 555. n. 239
(1871) {partiii, ; Colombia); Oberth., Et. d'Eiit. iv. p. 75. n. 232 (1880) (Toquiza and Llanos de
San Martin, Colombia) ; Stand., Exot. Tagf. i. p. 17 (1884) (partim ; Colombia).
cJ. Forewing somewhat distinctly falcate ; posterior spots of cell large, very
distinct also below, the subapical one at least as long as the interspace between the
two spots. The red spots of the npperside of the hindwing and the yellow sub-
marginal spot R- — R' on the whole somewhat larger than in the following
subspecies.
Hub. Colombia : Magdalena valley, presumably also Cauca valley.
A specimen from Frontino, Antioquia (T. K. Salmon), in coll. Godman,
approaches in the size of the cell-spots Ecuadorian examples of P. I. leucaspis.
In the Tring Museum 12 cJ c? from : " Bogota " ; Antioriuia.
b. P. leucaspis leucaspis.
Papllio leucasjih Godart, I.e. (Pern?) ; Boisd., Spec. Gen. Lep. i. p. 349. n. 190 (1836) (Peru?) ;
Lucas, in Guer. Diet. Pitt. Uist. Nat. vii. p. 50 (1838) (Peru) ; Doubl., Westw. & Hew., Gen.
Dinrn. Lep. i. p. 16. n. 162 (1846) (Peru?); Gray, Cat. Lep. Inn. Brit. Miis. i. Pap. p. 38.
n. 187 (1852) (Peru ?) ; id., List Lep. Ins. Brit. Mus. i. Pap. p. 52. n. 195 (1856) (Peru ?) ;
Felder, I.e. {partim; Ecuador) ; Kirby, I.e. {partim ; Ecuador) ; HopfE., Stctt. Ent. Zeil. xl.
p. 47. n. 1 (1879) (Peru); Stand., I.e. {partim; Peru) ; Maass. & Weym., in Stubel, Rcisen
S. Amer., Lep. p. 64. n. 91 (1890) (Huamboya) ; iid., I.e. p. 80. n. 14 (1890) (N. Pern) ;
Haensch, Berl. Ent. Zeitselir. xWii. p. 154 (1903) (Archidona, 640 m.) ; Week.s, Illustr. Diurn.
Lep. p. 20 (1905) (Chulumani).
cJ. The spots in the cell of the forewing small, generally absent from the
underside or vestigial, the interspace between the two posterior spots much wider
than in the preceding subspecies, the subapical spot standing about halfway between
lower angle of cell and R^ ; this spot on the whole larger in Ecuadorian specimens
than in Peruvian and Bolivian ones. Blany individuals have in front of R- a small
discal dot, which does not appear to be ever present in P. I. lamis.
The specimen (name-type) in the Royal Scottish Museum belongs to this form.
Hab. Eastern Ecuador to Bolivia.
In the Tring Museum 100 SS from: Zamora (Ganjon, and 0. T. Baron);
Pozuzo, Huanuco, 800 — 1000 m. (W. Hoffmanns); Chanchamayo (Schunke) ;
Huancabamba, Cerro de Pasco (E. Bottger) ; Caradoc, Marcapata, 4000 ft., February
1901 (G. Ockenden) ; R. Inambari, S.E. Peru, July 1900, 1000 m. (Simons) ;
R. Slucuri, S.E. Peru, June lOol, 25(J0 ft. (G. Ockenden) ; Oroya to Limbani
(734)
Carabaya, S.E. Peru, January 1901 (G. Ockenden) ; S. Domingo, Carabaya, 4500—
6000 ft., January 1901 (G. Ockenden); Charnplava, Bolivia, Vim m., June 1901
(Simons).
XVII. Dolicaon Group.
One of tbe most interesting features of this group is the anastomosis according
to species of one or of two subcostal veins of the forewing with the costal vein. The
character appears to be constant within most species.
The scent-organ is practically the same in all the species. The wool consists
of long and very thin hairs which are widest in the centre. The fold is densely
scaled beneath the wool.
Six species :
a. Forewing with broad black band across cell joining the
black distal border b.
The band across cell not reaching to distal border . . c.
b. Underside of hiudwiug with two pale postdiscal spots
W — M' separated from the greenish white basi-discal
area by a brown band Species No. 163.
These spots absent Species No. 164.
c. Veins SC and SO" of forewing confluent with C . . d.
Only vein SC^ of forewing confluent with C . . . e.
d. Black distal border of hindwing narrow ; a narrow discal
baud on underside from costal margin to anal spot . Species No. 106.
Black distal border of hindwing very broad ; a short
discal band ou underside from costal margin to distal
border, which it reaches before middle . . . Species No. 165.
e. A red anal spot on hindwing Species No. 167.
No red anal spot on hindwing /.
/. Wings greenish white above Species No. 168.
Wings creamy buff above Species No. 169.
163. Papilio serville Godt. (1824).
Papillo serville Godart, Eiic. Meth. ix. Suppl. p. 809. n. 4G-7 (1824) (Atner. merid.).
Papiliu servilkl (!), Boisduval, Spec. Gen. Lep. i. p. 34G. n. ls7 (1831!) (America).
(?. The only reliable difference between this insect and P. columbus Kollar
which we can find is the presence, on the underside of the hindwing in P. senille,
of two pale spots R=— M', which are. separated from the pale basi-discal area by a
black-bruwn band.
If the whole range of P. serville is taken into consideration, the characters by
which the two insects are usually distinguished in books do not hold good. The
clayish lateral streak on the abdomen, the brown streaks in the cell of the forewing,
the" dots ou the underside of the head and on the breast, which are said to charac-
.terise P. serville, are not always present in the specimens of this species i'rom
Venezuela and Colombia. More constant in all localities, but by no means (iuite
constant, is the slightly diflferent position of the short greenish costal band of the
forewing, situated proximally of the subcostal fork, this band being more obli()ue in
P. columbus than in serville ; and further, the colour of the submarginal area of the
underside of the forewing, which is more purplish white in columbus than in serville,
the black streaks in this area, moreover, being more prominent in serville than in
( 735 )
columbus. The clayish streak on tlie nnderside of the abdominal fold is more
distinct in scrrillc than in cohimbns.
The clasping organs are individnall}' somewhat variable in both insects. We
liave not found any distinction between the two species in these organs.
The specimens of P. serville from Venezuela are the most interesting. They
look exactly like a cross between Colombian serville and Colombian colnmbus.
However, as P. columbus is not known from Venezuela, there can be no question of
hybridisation, to which describers so often resort in order to get easily over
difficulties.
Godart did not know from which locality the type came. Hitherto Colombian
specimens have usually been regarded as typical. This is apparently erroneous;
though Godart's description, being taken from a single specimen, — probably lost :
Boisduval, I.e., says that he had seen the type, but does not add anything to Godart's
description, — is naturally not so precise as to enable us to say with absolute certainty
from which country the type came. However, there is one point in his description
which bears on this question. The abdomen is said to be like that of P. dolicaon.
Now, in P. dolicaon the pale stripes of the abdomen are always broad. This is the
case only in the southern subspecies of P. serville. Therefore we have to accept as
typical the southern form of serville, not the northern. The Colomliian insects were
not yet known at Godart's time ; all the very common Colombian forms have been
described later. The South American species of the Enc. Meth. were from Brazil,
Surinam (and Guyane), and from Peru. We may therefore safely assume that the
sjiecimen of P. serville also came from Peru. The " Peru " of that time, however,
included what became Bolivia in 1825. Therefore it is impossible to say whether
the Peruvian specimens came from what is now termed Peru, or whether they came
from some part of the present-day Bolivia.
Neuration : SC and SC" of fore wing confluent with C, SC- seldom free.
Genitalia : Ventral process of clasper minute ; central ridge of harpe a half-
crescent, acuminate ventrally and dorsally, the ventral angle being produced basad.
? and early stages not known.
Hab. Northern Venezuela, Colombia to Bolivia.
Two subspecies.
a. P. serville acritus subsp nov.
rap'dio servilhi, Gray {nan Godart, 1824, err. det.), Cat. Lep. Lis. Brit. .l/«s. i. Pap. p. 30. D. 174
(1852) (Colombia) ; id., List Lep. Lis. Brit. Miis. i. Pap. p. 48. n. 182 (185t;) (parlim) ; Felder,
Verh. Zoo!. Bol. Ges. Wieii xiv. p. .BOO. n. 166 (1864) (Bogota ; Venezuela) ; id., Relse Nurara,
Lep. p. 49. n. 38 (1865) (Bogota) ; Kirby, Cat. Diurii. Lep. p. 555. n. 238 (1871) (Colombia ;
"var." excl.); Butl. & Druce, Proc. Zool. Soc. Loud. p. 364. n. 371 (1874) ("Costa Rica"
eiTore) ; Oberth., Et. d'Eiil. iv. p. 74. n. 230 (1880) (Colombia) ; Maass. & Weym., in Stiibel,
Behen S. Amer., Lep. p. 24. n. 108 (1890) (west side of Cordillera of Bogota) ; iid., I.e. p. 30.
n. 38 (1890) (Guayabo, Cauca) ; Haase, Uiitersiich. .Mimicry i. p. 84 (1893).
S. Dots on head and breast usually small, sometimes absent; abdomen often
all black, the clayish lateral stripe absent or thin. The cell-streaks on forewing
often absent, seldom heavy. The pale submarginal spot C — SC- on underside of
bindwing standing usually in middle between distal margin and pale basi-discal area.
Hab. Northern Venezuela ; Eastern and Central Colombia ; name-type from
Venezuela.
In the Tring Museum 28 SS from : Tachira, Venezuela (Bricefio); Mocotonfi,
Venezuela (Bricefio) ; "Bogota" ; Muzo, December 1890 ; Casanare, October 1898,
( 730 )
b. r. sercille scrcille Godt. (1824).
Papilio servlUe Godart, I.e.
Papilio serviUei, Boisduval, I.e. ; id., Bull. Soc. Eiit. France p. 153 (1874) (distinct from hippodamus ;
Ecaador) ; Hopff., Slelt. Ent. Zeit. xl. p. 47. n. 3 (1879) (Chanchamayo ; dist. iromhippodaimni);
Staud., Exot. Tarif. i. p. 17 (1884) (Peru ; Ecuador) ; Haensch, Berl Eiit. Zeilsrhi: xlvii. p. 154
(1903) (Archidona, G40 m.).
rapilio sen-llUiei C). Lucas, iu Guer., Dkl. Pill. HUl. X,:t. vii. p. 50 (1838) (liab?).
Papilin boliriana "Weeks, Jllusli: Diurn. Lep. p. 20 (1905) (Chulumani ; this insect ? nomeii nudum !
dealer's name ?).
g. Dots on head and breast always j}reseiit ; abdomen always with two pale
Stripes on each side, the npper one being broad. Cell-streaks of forewing heavier
than in P. s. acritns \ the snbapical cell-patch always washed with brown. The
pale patches in the distal area of the underside of the hindwing on the whole more
distinct, and the npper one of them nearer the basi-discal area than in acritns; the
hindwing on an average less elongate.
JIab. Western Colombia : Cauca ; Ecaador to Bolivia.
In the Tring Mnsenm 110 SS from: Archidona (W. Goodfellow) ; Loja ;
Zamora (0. T. Baron) ; Chanchamayo (W. Hoffmanns ; Schnnke) ; Marcapata,
4500 ft. ; Poznzo, Huilnnco (W. Hoflmanns) ; Perene R. (Simons ; Watkins &
Tomlinson) ; S. Domingo, La Union, and Oroya, Carabaya (G. Ockendeu) ;
Hnaucabamba, Cerro de Pasco, Jnnin (E. Bottger) ; Chulnmani, Bolivia, January
1901 (Simons) ; R. Tanampayo, Bolivia (Garlej)])).
164. Papilio columbus Kollar (1850).
Papilio columhus Kollar, Denkschr. K. Ak. Wiss. Wirn, Malli. Ifalum: CI. i. p. 351. n. 1, t. 42. fig. 1. 2
(1850) (Rio Meta).
Papilio hippodamiLi Doubleday, i/8< Lep. Ins. Brit. Mus. i. p. 9 (1845) {nom. nnd., Colombia);
id., Westw. & Hew., Gen. Diurn. Lrp. i. p. 15. n. 134 (1846) ; iid.. I.e. ii. p. 529 (1852)
(hippodamus synon. of serrillei — eolumljus Koll.) ; Feld., Verh. Zool. Bol. Ges. Wien xiv. p. 300.
n. 167 (1864) (Bogota ; "Venezuela," errors); Boisd., Bull. Soe. Enl. Franee p. 153 (1874)
(distinct from .^lerriUei = rohtmbus) ; Obcrth., El. d' Ent. iv. p. 74. n. 231 (1880) (Muzo ;
Carare); Maass. & Weym., in Stiibel, lieisen S. Anter., Lep. p. 32. n. 136 (1890) (Colombia) ;
Prinz. Theresa, Berl. Ent. Zeilsehr. xlvi. p. 241. n. 5 (1901) (Bogota, bought).
Papilio servillei, Gray, Cat. Lep. Lis. Brit. Mns., Pup. i. p. 36. n. 174 (1K52) (/jarlim).
Papilio hurtoniVieakvit, Proe. Acad. Nat. Sci. Philad. p. 89. n. 55 (1868) (Issagasuga, Colombia) ;
Strecker, Lep. Rhop. Het. p. 15 (1873) (= "coji/mfow Hew." errore); id., I.e. Suppl. iii. p. 17
(1900) (type ; = hippodamus).
Pajnlio serrillei var. a. P. hijipodamus, Kirby, Cat. Diurn. Lep. p. 555. sub n. 2.38 (1871) (Colombia ;
Venezuela).
Papilio hippodamus var. /«/«( Oberthiir, Et. d'Ent. iv. p. 74. sub n. 231 (1880) (Muzo).
Papilio servillei Godt. var. hijipodamus, Staudiiiger, Emt. Taijf. i. p. 17. t. 12 (1884) (North
Colombia ; Venezuela) ; Haase, Vntersuch. Mimieri/ i. p. 84 (1893).
Tlie name hippodamu.H can unfortnnately not stand for this species. The
name appears first in Doubluday's List of 1845, where it is a mere nomcn nudum,
placed between P. agcsilaus and P. pliiloxenus, not the shghtest indic.ntion beiog
given as to what kind of Papilio the name was meant to designate. In Doubleday's
"Genera" of 1840 the name again ajipears as a nomen nudum, hare standing between
{aja.v ■=) marcellus and belle ropho ii ; but we gather from a note by Oberthiir, /.tf.
(1880) and a reference given by Folder, I.e. (1864), to a fignre in Doubleday's
" Genera," that some copies of the first number of the "Genera" — advance copies no
donbt — were distributed which contained a plate without number on which were
figured P. evan, hippodamus and pohjcuctes. We do not think that this plate can
( 737 )
be considered " published." In the Appendix to the "Genera" (1852) Westwood
says that hippodamus is the same as scrviUe, and that also colitmhts Kollar is a
synonym of the same. This is the first published statement of what hippodanius
is like, though the statement is in so far erroneous, as hippodamus cannot be
identical with sercille as well as with colambus, these two names applying to
two different species. The first description of hippodamus was given by Felder in
1864, Kirby, in his Catalogue, enumerating for this reason the insect as hippodamus
Felder. As the nomen nudum of 1845 and 184G {hippodamus) cannot supersede the
name columbus of 1850, we have to employ the latter name for the present insect.
Kirby, I.e., ajipears to have sunk colum'ius Kollar as a synonym of hippodamus not
because hippodamus had in his opinion a claim to priority, but because there is
another Papilio called columbus by Hewitson, which name Kirby dates also 1850.
However, Hewitson's name was published in 1851 — i.e. after Kollar's columbus.
This sjjecies has a very restricted range, being found only in Colombia and
North west Ecuador. Kollar, and after him Felder, ll.cc, gave Venezuelix as
habitat. However, the type came from the llio Meta, eastern side of the Cordillera
of Bogota, no specimens from Venezuela proper being known.
The black distal area of the hiudwing is nsnally separated from the cell, but
occasionally touches it, though it never enters it, as the band so often does in
P. sercille. The size of the small greenish costal band of the forewing and of
the subapical cell-patch is very variable. In one of our specimens the anal spot
of the hindwing is not red, but nearly as pale as the ground-colour of the wing.
Oberthi'ir, I.e., mentions a similar colour-aberration as ah.J'ulva.
The female resembles the male, the black bands being a little less extended.
Bab. Colombia : Cordillera of Bogota to west coast ; North-west Ecuador.
In the Tring Museum 140 Jc? from : R. Dagua (Rosenberg) ; Cauca (Child) ;
Pereira, Cauca; "Bogota"; Muzo, December 1S9G ; Lita, W. Ecuador, 3000 ft.
(Flemming & Miketta) ; Paramba, W. Ecuador, 3500 ft., March 1897 (Rosenberg) ;
Paramba, January — May 1898 (Flemming & Miketta).
A female in coll. Adams from the Cauca valley, the only one which we
have seen.
In coll. Godman from : Muzo ; Llanos de Rio Meta ; Froutiuo, Autioquia.
165. Papilio orabilis Butl. (1872).
PtipiVo orahiVs Butler, Cht. Ent. i. p. 84 (1872) (Costa Rica).
c?. Similar in appearance to P. serville and columbus Kollar (1850, non Hew.
1851), but easily recognised by the club of the antenna being black, seldom slightly
tawny, by the black median band of the forewing stopping short at or before M",
not joining the distal black border, the cell of the forewing being proportionately
longer, the red spot W — M^ of the hindwing being well marked above and
below, etc.
Genitalia: i. Ventral process of clasper much larger than in P. seniUc and
columbus, central ridge of harpe wider, proximal ventral edge of harpe not
produced basad.
? and early stages not known.
Ilab. Guatemala to West, Colombia.
The occurrence in Guatemala requires confirmation.
Two subspecies.
( 738)
a. P. orabilis orahilis Bntl. (1872).
Papilio orabilis Butler, l.r. ; id., Lep. Exot. p. 103. t. 58. Eg. 1 (1874) ; id. & Druce, Pmr. Zaol.
Soc. Loml. p. 3G5. n. 372 (1874) (Costa Rica) ; Kirby, Cul. Diurn. Lrp. p. 813. n. 380 (1877)
(Costa Rica) ; StauJ., Exnt. Taijf. i. p. 17 (1884) : Godra. .t Salv., Uiol. Ceiitr. Ainer., Rhop.
ii. p. -210. D. 42. t. 07. fig. 14. g, 15. genit. (IS'.til) (Guatemala; Costa Rica; Panama.—
" Colombia " alia subsp.).
S . The black obliqne discal band of the hindvving not marked on the nii])erside,
only shining thronsrh from below. Discal spot R' — R-of forewing, upperside, usually
absent, obliiine black median band often not reaching M-, the j)ortion between
cell and M- being shaded with greenish white, especially often in Costa Rica
specimens.
riab. Guatemala (a specimen in the Berlin Mnscnui, doubtful ?) ; Costa Rica ;
Bugaba and Chiriqui, Panama.
In the Tring Museum S Si from: Carillo, Costa Rica, June — July 1903
(Underwood) ; Carillo, 3000 ft., October 1904 (A. Hall) ; Volcan de Chiriqni,
5000—8000 ft. (Watson).
b. P. orabilis isocharis subsp. nov.
Papilio orabilis, Godman & Salv., I.e. (partim ; Colombia).
(?. Upperside. — Forewing : black median baud always extending to M- ;
distal border a little wider than in P. o. orabilis ; greenish white discal spot
R' — R- always present. Hindwing : black distal border wider than in the
preceding form, sending out obliquely forward a spur corresponding to the
oblique band of the underside, the spur not reaching beyond SC' (a similar spur
found in P. iphitas, doUcaon, etc.).
Underside. — The pale postdiscal band situated in the brown-black distal border
narrower than in the Central American subsi)ecies. Hindwing : submarginal
Innules R- — M' rather larger.
Clavish streaks of abdomen on the whole narrower than in P. o. orabilis.
Hob. West Colombia.
In the Tring Museum S c? c? from R. Dagua (W. F. H. Rosenberg).
A long series in coll. Charles Oberthiir from Juntas, R. Dagua.
10(5. Papilio salvini Bates (1804).
Papilin salvini Bates, Ent. Mn. Mag. i. p. 4. n. 8 (1864) (Guatemala) ; Felder, Verli. Zoo/. DnI. Gcs.
Wieii xiv. p. 301. n. 174 (1864) (cit. falsa) ; Hew., Exol. Bull. iii. Pap. t. 8. fig. 23 (1805) ;
Boisd., C'oiisiiJ. Lep. Giialfm. p. 5 (1870) (Yucatan; Costa Rica); Kirby, Cat. Dinrn. Lc/i.
p. 555. n. 244 (1871) ; Oberth., El. irEnl. iv. p. 67. n. 183 (1880) (Guatemala ; " Mexico") ;
Godm. & Salv., Biol. Cenlr. Amer., lihop. ii. p. 217. t. 68. fig. 3. S, 4- genit. (1890) (Yucatiiu ;
Guatemala ; Brit. Honduras) ; Haase, Vnlersuch. Mimicry i. p. 85 (1893) (near lellerophuii,
errore).
Papilio eacus Hewitson, I.e. (Boisd. MS.).
In appearance reminding one of P. bellerophoii, of which sahini has
generally been considered a near ally ; but the species is in point of fact a relative
of P. orabilis and callias.
cJ. Body as in P. callias and P. dolicaon. Antenna tawny. Cell of forewing
long, as in P. columbus Kollar (1850, non Hewitson 1851) and orabilis; the same
kind of jiattfrn, but the black coll-biir narrow, not extending beyond cell, at right
( 739)
angles to costal edge, greenish white discal band much wider and longer than in
P. callias and orabilis. Black distal marsjinal border of hindwing narrower than
in the allied species ; yellowish white admargiual hmules R- — M- regnlarlj- cnrved,
not interrupted in middle. Scales of upper layer of fore- and hindwing (excejit
costal area of the latter) narrow.
Underside glossy, excepting posterior area of forewing, where the scales are
entire and have an oi)aqnc appearance. Forewing : brown discocellnlar band
continued to M' or beyond, remaining separate from the narrow brown distal
border, the interspace having become white (secondarily). Hindwing : brown
discal band from costal margin nearly straight to red anal spots, corresponding
to the costal portion of the forked band of P. callias and the inner arm of the
fork, the outer arm of the fork being indicated in P. saicini by a slight projection
from the band before R"; brown distal border about as broad as the discal band.
Neuration : SO* and SC' of forewing confluent with C, the tip of C curving
towards SC'^.
Genitalia : A broad denticulate ventral process on clasper ; central ridge of
harpe narrowed apicad, without a row of teeth transversely across the lobe ; ventral
proximal edge of harpe not produced basad.
¥ and early stages not known.
Ilab. Mexico, Yucatan ; Guatemala ; British Honduras.
The locality Yucatan reijnires confirmation.
In the Tring Museum 'J S S from : Vera Paz, Guatemala ; " Guatemala."
KiT. Papilio callias uom. nov.
rupUio cahuiihus Hewitson {wm Kollar, 1850), Tows. Eni. Soc. Loud. (2). i. p. 80. t. 10. fig. 1 (IS.jI)
(Villa Nova, Amazons) ; Doubl., Westw. & Hew., Gen. Diurn. Lep. ii. p. 529 (1852) (Amazons) ;
Gray, Cut. Lep. Im. Brit. itnx. i. Pap. p. Sll. n. 175 (1852) (Amazons) ; Wall., Trans. Enl. Son.
Lone!. (2). ii. p. 255 (1854) (Amazons) ; Gray, List Lep. Lis. Brit. Jfiis. i. Bap. p. 49. n. 183
(1856) (Amazons) ; Bates, Trans. Ent. Sor. Lond. (2). v. p. .348 (18(;i) (Villa Nova, November ;
Ega ; R. Japura) ; id., Joiirn. Eiit. i. p. 229. n. 32 (18G2) (Upper Amazons) ; id., Xaliiriil. Riv.
Amaz. p. I5G (18G4) : Felder, VerJi. Zool. Bot. Ges. Wien xiv. p. .300. n. 1(31 (18(;4) (Villa Nova ;
Ega ; R. Japura) ; Kirby, Cat. Diurn. Lep. p. 554. n. 236 (1871) (Amaz. sup.) ; Oberth., Et.
d'Eiit. iv. p. 74. n. 229 (1880) (Tefii) ; Stand., Exol. Toijf. i. p. 16. t. II (1884) (Rio Maue's to
Ecuador and Peru); Habnel, //•« iii. p. 250 (1800) (Manes); id., /.c. p. 283 (1890) (Pebas) ;
Haase, I'ldersuch. ijiinicri/ i. p. 84 (1893) (Amazons) ; Michael, Iris vii. p. 214 (1894) Sao
Paulo de Olivenfa).
Though Westwood as well as Gray and Felder, ll.cc, give Kollar's columbus
priority over Hewitson's columbus, they accept nevertheless the name columbus for
the later discovered insect. We think they were guided in doing this by the
Fabriciun habit in nomenclature of employing again in the same genns for another
species a name which is a synonym, columbus Kollar being treated by those authors
as a synonym of hippodamus. Kirby, I.e., dates both columbus Hew. and columbus
Kollar from 1850, in which he is wrong, the former being published in 1851. The
present species, therefore, has no valid name.
S. Body clayish white beneath, with the usual black lines on abdomen ; clayish
dorso-lateral stripe of abdomen broad. Antennae brownish black (Haase, I.e.,
erroneously describes the club as tawny).
Wings, upperside : forewing subtransparent apically, the scales being reduced
in width ; pattern nearest to that of P. dolirno/i, greenish white area distaliy almost
(740)
evenly ronnded ; short costal discal band nsnally stopping short at R', never
extending beyond R-, varialile in width from 1 to 4 mm. Hindwing shorter than
in P. salfiiii, oruhilis, dioxijijius, etc., Imt distinctly triangnlar, dentate, tooth li'
nsnally prolonged to a short tail, tail R^ thin ; black distal marginal area produced
discad on veins R- and following, usually a black bar on disc in front of and again
behind S(.'- corresponding to the discal band of underside ; red anal spot varying
from being large and rounded to being transverse and thin.
On underside the most interesting character in pattern is the black forked discal
band of the hindwing ; the proximal branch nsnally touches apex of cell, being
seldom so far proximal that the extremity of the cell is occupied by a spot of the
ground-colour. The onter branch is short, joining the distal marginal area at R'^
The greenish white interspaces between the two branches vary much in width, being
sometimes shaded over with brown. The development of the two branches in the
allied species is peculiar : in P. sahini the outer branch has disappeared, in
P. orabilis, serrille, and columbus Kollar (1850) the inner branch has become
obliterated, being often indicated by a thin line between R- and R'', and in
P, dolicaon and iphitas both branches are preserved, but the proximal one has
been shifted basad, standing at right angles (or nearly) to the abdominal margin.
Now, in the female of callias the whole band is shifted distad, crossing R- about
3 mm. from cell ; the costal interspace between the band and the distal border is
practically absent, there being just a pale shadow between C and SC'', and the spots
of the gronnd-colonr between the two branches of the band are reduced to small
spots of a huffish tint.
? . Besides the difference in the discal band of the hindwing as just mentioned,
the female differs from the male in the black cell-band of the forewing being
narrowed to a point. The red anal spot of the npperside is large ; the greenish
white spot M' — M'- standing near the red spot is surrounded with black, being
separated from the greenish white basi-discal area, which is occasionally the case
also in male.
Nenration : SC of forewing joining C ; SC^ free.
Genitalia : S. No ventral process on clasper ; ventral proximal angle of
harpe produced basad ; central ridge with transverse dentate ridge. ¥ not
dissected.
Early stages not known.
Hab. East Ecuador ; Pern, eastwards to Oyapock R., Lower Amazons.
In the Tring Museum 30 tJcJfrom: Coca, R. Napo, May— July 1809 (W.
Goodfellow) ; R. Chuclniras, affl. of R. Palcazii, 320 m. (W. Hoffmanns) ; Palcazn
(Sedlmayr) ; Poznzo (W. Hoffmanns) ; R. Uanpes, Upper R. Negro.
Two ? ? from R. Oyapock in Mns. Giildi, Para, one of which has been kindly
transferred to the Tring Museum by the owner.
168. Papilio dolicaon Cram. (1775).
Papilio Eqnes Adiiviis dolicann Cramer, Pup. E.nit. i. p. 25. t. 17. fig. C. D. (1775) (East Indies !).
Though there are considerable differences between P. dolicaon and serville, the
two species have much in common.
6. Head and breast black, dotted with white; abdomen huffish clay-colonr,
black lines usually broader basally than ajiically. Apex of antenna fawny ochrareous.
( '^1 )
Forewiiig : black patcli across cell oliliqne, continued anteriorly to base ;
greenish white subapical interspace of cell often washed with black on npperside, as
in P. serviUe ; a row of three to six spots distall}' of cell from costal margin to R',
rarely a spot behind E^, the spots more or less elongate, separate ; on underside this
row of spots replaced by a very broad, glossy, purplish bnif band, which usually
extends to M^ or beyond. Hindwing broader and shorter than in P. senille,
costal margin not much shorter than distal margin ; black distal border broad,
usually dentate at some of the veins, dilated between SC'-* and Ri in most specimens,
a spot of the ground-colour being isolated between these veins ; a row of pale bine
admarginal dots, two in each cellule, those between C and R^ usually absent or
vestigial ; two rows of such spots on underside ; no red anal spot ; black discal band
crossing cell at M", the branch across cell standing at right angles (or nearly) to the
costal portion, which is about parallel to abdominal margin ; tail thin, apex buff
yellow.
? like male, larger, black distal areas of both wings and cell-band of forewing,
as well as the greenish white discal costal band of sjwts on furewing, reduced.
Neuratiou : SC of forewing conflaent with C ; SC" free.
Genitalia: S. Clasper without ventral process, or the process vestigial;
proximal ventral angle of harpe somewhat produced basad ; central ridge regularly
dentate, the teeth conical, especially those along the proximal edge, a short trans-
verse row near ventral angle. ? not dissected.
Early stages not known.
IJab. Colombia to Paraguay and Brazil.
a. P. doUcaoii luhriis snbsp. nov.
Pajjilio dolieaon, Staudinger, E.rot. Tagf. i. p. 18 (1884) {partim ; Antioquia, Colombia).
c?. Black ventral mesial line of abdomen broad. AVings, abore : forewing
more obtuse at apex than in the other forms ; white subapical cell-band narrow,
narrowing costad, sometimes reduced to a point at costal margin or abbreviated ;
discal spots long, spot R' — R' always present, rarely a dot behind R' ; veins M' and
M- more extended black than in the following forms. Hindwing more rounded,
resembling the wing of P. iphitas ; white spot SC^ — R' within the black distal
border always distinct, the black spot or line separating it from the basi-discal area
usually smaller than the white spot.
Undcrsiilr. — Pale distal area of forewing and the hindwing more purple than
in the other subspecies. Forewing : pale distal area more restricted, the veins
R' to M' traversing it very heavily black, separating the band into patches, the
patches M' — M^ being small or vestigial. Hindwing more or less obviously
washed with ochreous, especially in anal region ; black discal band broad, especially
the costal portion and the cell-liand; pale blue admarginal dots large, bnffish
patches C — R* outside Hack discal band rather sharply defined.
Ilab. Colombia ; known to us from the Magdalena valley, and the Llanos of
the Rio Meta, east side of the Cordillera of Bogota.
The form found in " Bogota " collections, usually confounded in collections witii
P. (I. deilcon.
In the Tring Museum 10 c?c? from " Bogota."
In coll. F. U. (Jodnian from the Llanos of R. Meta.
( 742 )
b. P. dolicaon deilcon Feld. (1865).
Piipilio dflleon Felder, Verb. Zool. Bot. Qes. Wien xiv. p. 300. n. 1G3 (18C4) (Bogota ; worn, niiil.) ;
id., Reisii Xorara, Lej,. p. 48. n. 37 (18C5) (Bogota); Staud., Exot. Tagf. i. p. 18 (1884)
(S.E. Peru).
Papilio (lolifiioii, Wallace, Trans. Eiit. Soc. Loud. (2). ii. p. 255 (1854) (Amazons) ; Bates, ih'nl. (2),
V. p. 348 (1861) (Paril to Peru) ; id.,/o«ra. Entom. i. p. 228. n. .31 (18(;2) (sparingly throughout
the Amazons) ; Druce, Proc. Zool. Soc. Loml. p. 245. n. 11 (1876) (Ucayali) ; Butl., Trana. Ent.
Soc. Ln>i,!. p. 146. n. 22'.l (1877) (R. Jutahi, February); Stand., E.ml. Tiiaf. i. p. 18. t. 12 (1884)
(partini ; Amazons) ; Hahnel, Iris iii. p. 250 (1890) (Maufo) ; id., I.e. p. 183 (1890) (Pebaa).
Piipilio dolicaon var. a. P. deilcon, Kirby, Cat. Diurn. Lep. p. 555. sub n. 237 (1871) (Colombia).
S. Black distal border of hindwing, underside, thin from R^ backwards, 1 to
2 mm. broad at R-; black discal band of hindwiug crossing cell at base of M',
the point of origin of this vein situated within the band, the patch of ground-
colour in apex of cell much larger than tlie patch sitnatcd at base of cellule
SC^— R'.
The subapical greenish white cell-baud of the forewing, upperside, usually
strongly shaded with black in the specimens from East Central Pern, and the
discal spots SC^ — R^ reduced in size ; these spots largest in Bolivian examples,
which have nearly always a distinct spot in the subcostal fork. The black distal area
nf the hindwing produced to cell at veins R- and R', widest in Peruvian examples,
ill which the white spots R' — R' situated at cell are often very small ; patch
SC^ — R' of ground-colour situated within black distal border small, often vestigial,
always smaller than the black spot standing at its proximal side ; pale postdiscal
patch SC* — R' of underside of hindwing larger and less sharply defined in
Amazonian, Peruvian and Bolivian specimens than in individuals from more
northern localities. Width of cell of forewiug and length of hindwing somewhat
variable.
Ilah. Colombia: probably "terra caliente " on eastern side of Cordillera of
Bogota ; from East Ecuador eastwards to Pard, and southwards to South-East
Bolivia and Matto Grosso.
Felder's specimens are said to be from "Bogota." As the "Bogota"
collections, which nowadays come on the market, contain apparently only the
preceding subspecies, it is possible that Felder's specimens are wrongly labelled,
being perhaps obtained in Eastern Ecuador.
In the Tring Museum 65 c?c? from : " Bogota" ; Coca, Rio Napo ("\V. Good-
fellow) ; R. Cacbyaco, affl. of R. Huallaga (Stuart) ; Chanchamayo (Schunkc) ;
Palcazn (Sedlmayr) ; Pozuzo (W. Hoffmanns); R. Cbuchuras, afB. of R. Palcazn
(W. Hoffmanns); Perene R. (AVatkins) ; Slontanas, R. Bladre de Dios (Ockenden) ;
La Union, R. Hnacamayo, :iOOO it. (Ockenden) ; JIapiri ; JIushay, Beni R. August
1895 (Stuart); mouth" of La Paz R. (Stuart); Yungas de "la Paz (Garlcpp) ;
Prov. Sara, Santa Cruz de la Sierra, February— A]iril 19(i4 (J. Steiuliach) ; Villa
Maria to Dinmantino, Matto Grosso, January 1807 (Andeer).
c. P. dolicaon tromcs subsp. nov.
Papilio dolicaon, Staudinger, E:rol. Toijf. i. p. 18 (1884) (jiarlim ; Venezuela) ; Hahnel, Iris iii
p. 203 (1890) (V.ilera).
S. Similar to the preceding subspecies. Subapical greenish white cell-band
of forewing, abore, very slightly shaded with black ; distal border of hindwing
( "43 )
broad, spots R' — R^ of gronnd-colonr situated around apex of cell small. Black
distal border of forewing, below, narrow, about 1 mm. wide from R' backwards ;
the pale distal area contiDued be_yond M" ; black discal band of hindwing crossing
cell a little distally of point of origin of M'-, there being a small spot of the
ground-colour at the base of cellule M' — M" : spot SC' — R' of ground-colour longer
than, or as long as, the spot situated in apex of cell.
Hah. Northern Venezuela.
In the Tring Museum 2 S <S from Palma iSola.
d. P. dolicaoii dolicaon Cram. (1775).
Pajnlio Eqiies Achictis dolicaon Cramer, I.e. ; Goeze, Enl. Bojtr. iii. 1. p. 79. n. 34 (177'J) ; Fabr.,
Spec. Iiu. ii. p. 13. n. 51 (1781) (Amer. merid.) ; id., Mant. Ins. ii. p. 7. n. 57 (1787) ; Jabl. &
Herbst, Nnturs. Schm. iii. p. 142. n. 95. t. 42. fig. 3. 4. (1788) ; Fabr., Eiit. Sy-sl. iii. 1. p. 23.
D. 60 (1793) (Amer. merid.) ; Gmelin, Syaf. Nal. i. 5. p. 2237. n. 312 (1790) (Amer. merid.).
Papilio dolicaon, Godart, Enc. Mclh. i.x. p. 40. n. 46 (1819) (partim) ; Feld., Verli. Zool. lint. Ges.
Wien xiv. p. 800. n. 162 (1864) (partim ; Surinam) ; Kirby, Cat. Diurn. Lep. p. 555. d. 237
(1871) ; Mbschl., Verh. Zool. But. Gcs. Wicii xxxii. p. 304 (1883) (Surinam) ; Haase, I'idersuch.
Miiniciu i. p. 84 (1893).
Cramer's figure represents this form.
(? ? . Greenish white subapical cell-band of forewing, above, not, or very little,
shaded with black ; distal marginal border of underside broader than in the otlier
forms, 3 to 5 mm. wide at R^, narrower in female. Distal border of upj)erside of
hindwing distant from cell ; on underside the discal band crossing cell be3'ond point
of origin of M-, spot of ground-colour at base of cellnle'SC' — R^ as long as, or longer
than, the spot situated in apex of cell ; buffisb postdiscal spot SC^ — R' transverse ;
marginal tooth R^ often prominent, sometimes with yellow dot on iipperside at tip
(corresponding to yellow apex of tail).
Hah. Orinoco ; the Guianas.
In the Tring Bluseum lU JcJ, 1 ?, from : Suapure, June 1901, and La Yuelta,
May 1904, Caura R. (S. M. Klages) ; R. Demerara, July 1897 ; Surinam.
e. P. dolicaon deicoon Feld. (1864).
Iphiclides dolicaon, Hilbner, Verz. bek. Schm. p. 82. n. 831 (1818?).
Eurytides dolicaon id., Samml. Exot. Schm. ii. t. 91 (1820?) ; Kirby, in Allen, Nat. Lilii:, Lrji. i. 2.
p. 272 (1896).
Papilio dolicaon, Boisduval, Spec. Gin. Lip. i. p. 347. n. 188 (1836) (Rio do Janeiro) ; Doubl., Li>il
Lrp. Ins. Brit. Mus. i. p. 16 (1845) (Brazil) ; id., Westw. & Hew., Gen. Diurn. Lep. i, p. 16. n. 154
(1846) (Brazil) ; Gray, Cat. Lep. Lis. Brit. Mas. i.Pap. p. 36. n. 176 (1852) (Brazil) ; Lucas, in
Chenu, Enc. Hid.. Nat., Pap. p. 38. t. 9. fig. 2. J (1851-53) (Brazil); Gray, List Lep. InslSrit. ilns.
i. Pap. p. 49. n. 184 (1856) (Brazil) ; Meaiir., Enum. Corp. Anim. Mas. Petr., Lep. i. p. 4. n. 55
(1857) (Brazil) ; Bull., Cat. Diurn. Lep. descr. Fabr. p. 239. n. 19 (1869) (Brazil) ; Burm.,
Descr. Rep. Arr/ent. v. Lep., Atlas p. 3. n. 2(1879) (Corcovado ; Tijuca) ; Oberth., Et. d'Ent. iv.
p. 74. n. 227(1880) (Brazil) ; Staud., Exot. Tag/, i. p. 18 (1884) (partim ; Sta. CatliariD.a) ;
Bijnningb., Verh. Ver. Xat. Unterh. Ilamhiirij ix. p. 27 (1895) (Corcovado).
Papilio deicoon Felder, Verh. Zool. Bot. Ges. Wien xiv. p. 300. n. 164 (1864) (Bras, austral.) ;
Staud., I.e. p. 18 (1884) (var. of dolicaon).
Papilio dolicaon var. P. deicoon, Kirby, Cat. Diurn. Lep. p. 555. sub n. 237 (1871) (Brazil).
Euri/tides deicaon (!), id., in Hiibn., Samml. Exot. Schmett. ed. ii. p. 92, t. 304. fig. 3. 4 (19:)-?).
3 ? . White spots in black distal area of forewing large ; subapical cell-patch
suhloiii a little washed witli black ; black border to hind win"' narrower than in
( ^44 )
the preceding forms ; white spot SO" — R' within the black border sharply defined,
the black spot separating it from the basi-discal area not always complete. Discal
band of hindwing, on ^inderside, crossing cell at M-, the point of origin of this
vein being within the band, the costal portion of the band a little more obliqne
than in the other forms, the greenish white basal area therefore more oblong ; the
yellowish white patches distally of the curved enter discal band sharply defined
as a rnle ; veins in proximal half of wing more extended black than in the other
subspecies.
Ilab. Brazil : Kio Grande do Sul northwards ; Paraguay.
In the Tring Museum 40 t?c?, I ?, from: Minas Geraes, February ISO'J
(Kennedy) ; Espiritu Santo ; Petropolis and Rio de Janeiro ; Bahuru, Sao Paulo
(Dr. Hemjiel) ; Castro, Parana (E. D. Jones) ; Sapucay, Paraguay, August and
October 19U1 and December 19U3 (W. Foster) ; Jataliy, Goyaz.
169. Papilio iphitas Uiibn. (1820?).
Papilio dolicaoii, Godart (nou Cramer, 1775, err. det.), E»c. .Velli. ix. p. 40. n. 41") (181(1) ; Donov.,
Nal. Repos. ii. t. 65 (1824) ; Prillw., Slett. Eiit. Zeh. xxvi. p. 120 (1865) (Corcovado : "light
yellow," perhaps ipJiHas?).
EurytiiUs iplutas Hubner, Samiul. Exot. Srl,m. ii. t. 02 (1820?) ; Kirby, ibi'l ed. ii. p. 92. t. 305.
fig..3. 4(190-?).
PapiUo !plilla.^,Boisduv3.\, Spec. Gen. Leji. i. p. 348. n. 180(1836)(Biode Janeiro; var. of rfo?;cao« ?) ;
Lucas, itv>. EmI. p. 14. t. 8. fig. 1 (1835) ; DouW., /./•«' Lep. Ins. Bril. Mas. i. p. 16 (1845)
(Brazil?) ; id., Westw. & Hew., Gen. Diurn. Lep. i. p. 16. n. 155 (1846) ; Gray, Cat. Le/i. Tn-i.
Bril. Mils. i. Pap. p. 36. n. 177 (1852) (Brazil) ; Lnras, in Chenu, E/ic. Uisl. Xiil., Pup. i. t. 11.
fig. 2. jj (1851-53) ; Gray, Lhl Lep. Ins. Bril. .!/«». i. Par- p. 40. n. 185 (1856) (Brazil) ;
Mdnutr., Ennm. Corp. Anim. JIiis. Pelrop., Lep. i. Siipjil. p. 68. n. 1122 (1857) (Brazil) ; Felder,
Verli. Zuul. Bui. Ges. Wien xiv. p. 300. n. 165 (1864) (Bras, austral.) : Kirby, Cal. Diurn. Lep.
p. 554. n. 237a (1871) (Brazil) ; Burm., Descr. Rep. Argent, v. Lep. p. 3. sub n. 2 (1879) (var. of
iloUcaou) ; Oberth., El. d'Enl. iv. p. 74. n. 228 (1880) (Brazil) ; Staud., E.rnl. Tagf. i. p. 18
(1884) (var. of ilolieaon ? ; Pernambuco) ; Haase, Unkr!<iii-li. .Vimii-r;/ i. p. 84 (1893) ; Bonningh.,
Verh. Ver. Nal. Unlerli. Hamburg ix. p. 27 (1895) (Organ Mts. ; not near Rio).
d. Similar to P. dolicaoii ; ground-colour creamy buif ; distal border of
forewing narrower behind, both above and below, than in P. dolicaon, veins
M'— SM- heavier black distally ; yellow discal spot R'— R° longer than the others.
Hindwing more rounded than in dolicaon ; black discal baud marked on
upperside between SC- and R', usually entering cell ; extreme tip of tail yellow.
Black discal band of forewing below shifted to margin between M' and SM".
Nearly all the veins of hindwing black ; discal band crossing cell distally of M*,
spot of ground-colour in apex of cell smaller than the spot situated at base of
cellule SC- — R' ; one row of jmle blue spots.
Neuration ; SC of forewing confluent with 0 ; SC'^ free.
Genitalia: No ventral process on valve; proximal ventral angle of harpe nut
produced; central ridge without teeth at proximal edge, a long tooth near ventral
angle corresponding to the transverse row of teeth of P. dolicaon, tip of ridge
pointed, no teeth near it.
? and early stages not known.
Ilab. Brazil.
In the Tring Museum 8 c?c? from : Fazenda Jerusalem, Espiritu Santo; Rio
de Janeiro (E. May ; doubtless from the Organ Mts.).
743 )
EXPL^UlfATION OF NEPRATION.
Fig. 4.
( 746)
INDEX
abboti, r.89.
abbotti, 689.
abderus, 646, 047, 048.
abilius, 47.i.
acamas, 415. ."iT'J.
acanthus, ."i79.
acauda, 513, 514.
acestes, 672.
achelous, 480.
aconopbas, 678.
aconophos, 678.
Acraea, 439.
acritjs, 735.
adaea, 640.
aegeus, 4.S3.
aeneas. 412. 417, 425, 432. 435, 454-7,' 461, 471,
489, 491, 493, 497, 498, 501, 506, 600.
aeneides, 417, 454, 497-502.'
agavus, 430, 438, 439, 443, 445, 079, 681.
agathokles, 505, 506.
agesilaus, 656, 098-703-705-708, 736.
aglaope, 480, 492, 494, 498, 501 .
aiax, 084.
ajax, 413, 414, 540, 543, 558, 582, 583, 684-90,
694, 690, 736.
albanus, 591.
alcamedes, 504.
Elector, 478.
alexiarcs, 581, 592.
aliaska, 553, .o54.
alopiu.i, 444.
alyattcs, 473, 470, 480, 482, 483.
amasis, ODl.
amazonis, 531.
amciias, 646, 647.
americus, 417, 430, 500, 543, 547, 029.
arnica, 647.
ammoni, ,588.
amosis, 577, 578, 600, 002.
ampbissus, 035, G30.
ampliat.a, 546.
amulius, .'•,28, .WO, .531.
anacharsis, 484.
anactus, 433.
anargua, 674.
anatmus, 666.
anaxilau.s, 097.
anaximander, 494.
anaximeiies, 494, 500.
anchises, 412, 413, 472, 474, 478, 482, 483, 485
486, 489, 493, 494, 496, 008.
iiiicliisiade.s 422, 425, .'(94, 595, 599, 002, GO),
607. 608, 009, G18, 000.
anrhisses, 0U9.
andraemou, 570, 571.
■inlrogeas, 579.
androgeos, 428, 577, 578, 580.
androgens, 428, 538, 573, 576, 577, 578, 579, 5«o,
002, 613.
androni, 668.
antheas, 506-
antliens, 434.
anthilochus, 583.
anticostiensis, 548.
antilochus, 413, 582.
antiquus, 523.
arbates, 491, 491. 494.
areas, 480, 484, 40.'<, 503, 506, 507.
arcesilaus, 697.
archalaus, 009.
archelaus, 608.
archemas, 517.
archesilaus, 710, 714-6, 717, 718.
archidamas, 515, 516, 517.
arcbytas, 637, 038, 639.
argentus, 016.
arianus, 674.
ariarathes, 425, 420, 671, 672-4.
aristaeus, 641.
aristeus, 636, 637, 039, 641, IU:<, 047, 049, 660.
aristodemus, 418, 508, 569, 570, 581.
aristogiton, 607.
ari^tomenes, 504.
aristor, 568.
ari/.ouensis, 588.
arrhipus, 505.
arripbus, 505.
arripus, 505.
ascalus, 440, 44 1 .
ascanius. 435, 4:')0, 437, 079.
asclepins, 48ii, li4.'i-8.
a.scolius, 024, 625, 626, 027.
asins, 683.
asiua, 650, 682, 083.
( ^4? )
assius, G83.
astenous, 513.
asterias, 543-5, 547.
asterioides, 416, 417, 541, 542, 545-7.
asterius, 414, 539-41, 543, 545-9, 597, 688.
asteroides, 545.
astinous, 510, 512.
astionous, 512.
astyagas, 682, 683.
astyalus, 573, 576.
astyanax, 545.
athous, 679, G80.
augias, 603.
augustus, 60.S.
australis, 585.
autocles, 557, 558, 561.
autosilaus, 098, 706.
autumnus, 470.
bachus, 430, 624, 627.
baia, 614.
bairdi, 425, 42i!, 539, 543, .'i51.
bairdii, 428.
bari, 641.
baroni, 647, 648.
belemus, 531.
belepbautes, 676.
belesis, 676, 677.
bellerophou, 654, 700, 736, 7,-!S.
belus, 489, 516, 5-^4-528-530-537
Bergmanniana, 419,
bias, 516, 517.
bimacuhitus, 5U7.
birchalli, 631, 632, 639.
bitias, 636, 637, 640, 642, 614, 651
Blakea, 435.
bochus, 455.
bogotanus, 470.
bolivar, 452, 453. 457, 467,
boliviana, 736.
bonhotei, 571.
brancbus, 656, 675, 676, 677.
brasilienais, 560.
brevicauda, 548.
bre-i-ifasciatus, 681.
brises, 610.
brissonius, 493-5.
bruoei, 549,
brutus, 424.
bunichus, 440, 441,
burchellanu.s, 466.
burtoni, 736.
caburi, 531.
cacicus, 651, 652- 1
caiguanabus, 567, 568,
Calaides, 538.
calchaa, 598.
oaleli, 504.
callias, 738, 739, 740.
callicleg, 492.
calliste, 728, 731, 732.
calogyna, 481.
calverleyi, 545 -6-7, 697.
campeiro, 440.
canadensis, 583.
candezei, 478.
capys, 607, 60).
catamelas, 616.
cauca. 476, 506,
caudius, 618, 620. 621.
cebriones, 518, 519,
celadon, 654, 691 3.
cephalus, 636.
ceus, 611.
chabrias, 445, 450, 452,
chalcas, 598.
chalceus, 529, 531, 532.
chalcus, 598.
cbamissonia, 438, 439. 440, 441.
Charaxes, 538.
charoba, 674.
chiansiades, 604.
childrenae, 449, 462, 463.
chinsiades, 604,
chii'odamas, 615.
chironis, 610.
chlorodamas, 525.
choridamas, 661.
cbresphontes, 564.
chrysodamas, 533.
chrysomelus, 628.
cincinnatus, 646.
cinyras, 429, 559, 560.
cixius, 469-71.
Claudius, 678, 680, 681.
clearchus, 638.
cleolas, 621.
cleombrotus, 542. 658.
cleotas, 538, 628, 636, 637, 639, 640, 641. 644,
645, 649, 650.
cleothas, 040.
clusoculis, 664.
clytia, 433, 434, 550.
cochabamba, 529, 531, 532.
coelebs, 643.
coelus, 445, 451.
coloro, 551, 552.
Columbus, 423, 435, 436, 438, 443. 445, 449, 735,
736-40.
concinnatus, 647.
conon, 704-6.
census, 499.
copanae, 533, 534.
corbis, 513.
corebus, 639.
coresilaus, 701 .
coristbeus, 641.
coroebus, 639.
48
( ?48)
crassus, 529, 536, 537.
cresphoutes. 538. 550, 557, 560, 561_, 562.
cresphontinus, 56'.l. 571).
crispus, 525.
ctesiades, 642.
ctesias. G3l!, 041-3.
cutora. 462, 466. 471.
cutorina, 452, 453, 467.
curvifascia, 545. 546.
cyamoD, 073, 674.
cymochles, 484, 4^7.
cymocles, 485.
cyniras, 560.
cyphotes, 463, 485, 486.
daguanus, 625.
damis, 456.
damocrates, 442.
daphnis, 569, 570.
dardanus, 415, 423, 458.
dares, 444.
dariensis, 715, 716.
daunus, 539, 581, 589.
dcicaon, 743.
deicoon, 743.
dcileon, 742.
demas, 503.
demodocus, 521.
devilliers, 419, 510, 511, 514, 515.
devilliersi, 419, 515.
diaphora, 633.
diaphorus, 728, 729.
diceros, 470.
dimas, 415, 502, 503, 600.
diodorus, 440.
dione, 639.
diores, 731.
dioxippus, 729, 730, 732,
docimus, 504,
dolicaon, 551, 654, 655, 735, 738, 740, 741,
743, 744.
dolius, 724.
dominicus, 519.
dorcus, 434.
drepanOD, 576.
drucei, 428, 445, 447, 456, 466.
drucii, 428.
dysmias, 642.
eari.'<, 723.
ecbolius, 500, 501,
echedorus, 440, 441,
echelus, 435, 455, 48u, 495-7.
echemoD, 494, 495, 496, 497, 501.
echephron, 496.
echion, 496, 497,
eimeri, 705.
elatos, 481.
electrion, 046.
742,
electi-yon, 648, 049.
Endopogon, 435,
epenetus, 604,
cpidaurii.i, 578.
epidaus, 698, 699, 70i).
epithoas, 557,
equestris, 464,
cracon, 528.
orgeteles, 496, 497,
eridamas, 662,
erimanthus, 5')4,
erithalion, 464, 472, 474, 475, 476, 478, 470, 481,
484, 500,
erlace.s, 463, 404, 465, 469, 492.
erostratus, 605.
erymanthus, 531, 535.
erythrus, 474, 470,
eteocles, 485.
etias, 486.
eucharia, 456.
Eulepis, 538.
eumelea, 672.
eupales, 499.
eupatorion, 679, 682.
Euphoeades, 538,
eurimedes, 422, 498, 501, 504-7,
eurimodon, 591,
euristeus, 417, 492, 501.
eurotas, 642, 643,
Euryadea, 439,
eurybate8, 499.
eurydamas, 518,
eurydorus, 441,
euryleon, 425, li56, 663, 665, 606.
eurymander, 630, 631.
eurjmas, 493,
eurymedes, 504, ,51)7,
eurymedon, 581, 590.
Eurytides, 655.
euterpinus, 653, 054.
evagoras, 672, 673.
evan, 730.
evander, 007, 009, 660.
extendatus, 681.
felicis, 694.
fenochionis, 700.
flava, 621.
flavescens, 611.
liavida, 621.
flavomaculatus, 598.
fietcheri, 585.
tloridensis, 689.
foetterlei, 487, 488.
fortis, 704.
fulva, 730, 737.
garamas, 645, 646 9,
garcia, 592.
f 749 )
gargasus, 454, 497, 498, 500-2.
garleppi, 424, 617, 618.
gayi, 673, G74.
glaucolaus, 702, 707, 708-11, 713, 717.
glaucus, 412-4, 431, 538, 579, 581, 582, 583-5
588, 592, 593, 687, 088, C95.
godmani, 633.
gratianus, 4(J7, 468.
grayi, 630, 631.
grotei, 436.
guaco, 1175.
gundlachianus, 423, 43.'), 430, 443.
baenschi, 665.
habneli, 424, 445, 449, 511.
halex, 667.
hai-modiu.s, 423, 425, 604. 667, 608, 669, 070.
barrisianu,s, 678, 079, 680, 081.
harris.sianus, 079.
hebnis. 741.
hectorkics, 418, 615, 010, 018.
bedae, 488.
Heliconiiis, 713.
helios, 720-2.
bellanichus, 420, 628, 029.
bellanicus, 028.
belleri, 035.
hephaestion, 677.
Heraclides, 538.
hermolaus, 059.
herodotus, 015.
herostratus, 005.
hesperus, 423.
hestorides, 016.
betaerius, 711, 712.
bierocles, 465, 486.
himeros, 614, 615.
bipparchus, 666.
bippason, 503, 000, 001.
bippasonides, 002.
bippodamus, 7.;0, 737, V.39.
bippomedon, 575, 577.
bipponous, 008, 009.
bollandi, 549.
homerus, 415, 02.3, 033, 045, 649.
homothoas, 581.
hostilius, 075.
byperion, 520, 000, 001.
hypodamas, .522.
hyppason, 578, 599, 600-2.
idaeus, 415, 479, 607, 009.
ideus, 007.
ilioneus, 595.
illyria, 695.
ilus, 415, 050, 072, 074, 675.
imaus, 668.
imerius, 419, 603.
immarginatup, 003.
inca, 653.
incandescens, 480.
indra, 539, 542, 552, 553.
interruptus, 617.
iopas, 532.
Ipbiclides, 055.
iphidama.s, 412, 415, 472-4, 476-479, 482, 483.
ipbitas, 740, 741, 744.
isidorus, 007, 008, 610, 611, 012,018.
isocharis, 738.
isus, 667.
Ithobalus, 433, 435.
jaguarae, 491.
jamaicensis,522.
janira, 527, 528.
jelskii, 650.
judicael, 644, 649.
kamtschadalus, 544.
kirbyi, 095.
klagesi, 445, 453.
lacandones, 730, 731.
lacandorea, 730.
lacordairei, 643.
lacydes, 464, 405, 460, 409.
laetitia, 037, 038.
laius, 681.
lamarcbei, 615.
lamis, 733.
lampedon, 500.
laodamas, 527, 532, 534, 537.
laodocus, 580.
latiQUs. 5.W.
lecontei, 690.
leilup, 418, 71.3,719.
lenaeus, 043, 644.
leontis. 599.
leosthenes, 654.
lepidus, 5 !0, 537.
leptale:i,629, 021.
leucas, 709.
leucaspi.. 732, 733.
leucone^, 715.
leucostictus. Oil.
leuctra, 674.
lewisi, 591.
licophroD, 577.
linus, 456.
Listerella, 419.
locris, 457.
lorzae, 731, 732.
lucayus, 52l.
lucianus, 517, 518.
luctuosa, 405.
lycander, 493.
( 750 )
lycidas. 528, 529, 534, 535, 537.
lycimenes, 472, 473, 475-81.
lycomes, ■402.
lycophron, 41.i, 568, 569, 572, 573, 574, 575, 576,
581, 59.3, 014.
lycoraeus, 571.
lycoroi'Us, 571.
lycortas, 639, 040.
lysander, 438, 459, 489, 491, 492-5, 498.
lysimachus, 492.
lysithous, 615, 616, 655, 678, 079-681, 682.
machaon,417, 423, 420, 431, 537, 539, 540, 549,
550, 553, 654, 686, 087.
machaonides, 571, 649.
macrosilaus, 707, 714-7.
madyas, 525.
madyes, 516, 524, 525, 526.
manllus, 083.
marcellinus, 690-3.
marcellus, 423-0, 055. 684-9, 090, 094, 090, 736.
marchandi. 726, 727, 730.
marcius, 454, 455,
mathaDi, 516.
mazeppa, 467.
mediocauda, 427, 548.
megalurus, 712.
melaenus, 709.
melania, 616.
melasina, 542.
meleander, 493.
melonius, 556,
menatius, 641,
menes, 672.
mentor, 415, 576, 614.
mercedes, 651.
metagenes, 673.
metaphaon, 602, 663.
mezentius, 616.
microdamas, 660.
mithras, 450.
molopa, 710-2.
montezunia, 444.
morelius, 635.
mullicaudata, ."i89.
mycale, 505.
mylotes, £04, 505.
nais, 653.
nealces, 558, 559.
neodama.s, 518, 519, 527.
neophilus, 497, 498, 499, 501, 502, 504, 507.
neosilaus, 704.
nephaleon, 489. ■■
nephalion, 488, 489.
nezahualcoyotl, 513, 514.
niger, 694, 695.
nigrescens, 094. «
nigricornis, 712, 719-21.
nitra, 539, 550, 551.
nitulus, 588,
numa, 400, 491.
numicus, 567.
numitor, 528-31.
Nymphalidae, 413.
nymphas, 450.
nympbius, 612.
obertbueri, 696.
oebalus, 574-7.
oedippus, 463.
oedipus, 403, 680, 082
oedypus, 403.
olbius, 732.
oleotas, 040.
olivencius, 500.
opalinus, 425, 466.
opleus. 458.
orabilia, 737, 738-40.
orbignyanus, 428, 486, 487.
orcbamup. 619, 620, 621. '
oregoDia. 417. 425, 549, 550.
oregonius, 549.
oregonu.i, 549.
orellana. 458.
Orestes. 423. 577.
Ornitboptera, 433.
ornofagus, 602.
omytbion, 557, 573t
orsilius, 503.
orsua, 514,
ortbosilaus, 412. 721.
osmandryas, 489,
osymanduas, 488.
osyris, 482. 484, 485.
oviedo, 557, 562.
oxilus, 5i'p4.
oxjnius, 603, 004.
paciflcus, 612.
paeon, 566, 567.
palamedes. 572, 594, 598, 599, 086.
pallas, 573, 574.
panaiuensis, 727.
panares, 480.
pandiou, 422, 007, 012.
pandonius, 007.
pandrosus, 615, 616.
panthias, 638.
pantbonus, 491, 493, 607.
paraensis, 602.
paraliu?, 474.
parianus. 502,
Paridcs, 435.
Parnassius, 439.
parsodes, 455. 492-4.
patros, 617, 020, 621.
pausainas, 659.
pausanias, 656. 658, 659.
pelaus, 415, 602, 603, 004, G20.
peleides, 417. 623.
peleus, 603, 020.
penthesilaus, 714-(;, 718.
peon, 567.
peranthus, 579.
pergamus, 553.
perithou.s, 577.
perrhaebus, 441, 442.
perrhebus, 4.38, 440, 441, 44-? 444^ 682.
phaenoD, 476.
phaaton, 638-40, 644.
pbaetoD, 638.
phalaecus, 442, 511.
phalias, 480-3.
phanias, 575, 577.
phanostratus, 605.
phaon, 661, 662-4.
pharax, 663.
pharnaces, 601, 605, 606, 608.
philenor, 412, 510-2-6.
philetas, 524.
philinor, 513.
philocleon, 631).
philolaus, 657, 688, 693, 695-8.
philo.\-enu.s, 730.
phosphorus, 467, 468.
photinus, 443, 444.
phryneus, 439.
phrynicbus, 494.
phylarcbus, 491.
pilumnus, 581, 593.
piranthous, 577.
piranthus, 577-9.
pirithous, 568, 573, 574, 576.
pisander, 497.
pithonius, 506, 665.
pizarro, 445, 452.
plaesiolaus, 695, 696.
platydesma, 679.
plinius, 525.
podalirius, 413, 654, 687, 720.
polidamas, 428.
polybius, 622.
polycaon, 577-80, 602.
polyoharmus, 605.
polycrates, 522.
polydamas, 428, 510, 511, 515, 517-20-3
579, 086.
polyeuctes, 736.
polyinatus, .50.3.
polymetus, 496, 501, 502, 503.;
polypbroD, 49(;, 497.
polystictus, 526, 527.
polyxenes, 414, 417, 426, 427, 430, 539, 540
543, 545, 547, 548. 5.50, 688.
polyzelus, 478, 479.
pompeius, 491, 0O0-9, 611.
pompejus, 491, 493, 495, 609, 610.
pomponius, 679, 682.
pot'. ue, 508.
( 751 )
prasinus, 658.
Priamides, 538.
priamas, 423.
procas, 532, 534.
proneus, 437, 439.
protesilaus, 412-4, 418, 425, 427, 434 655
684-90, 702-712-7-25.
protesileus, 713.
proteus, 484, 488.
protodamas, 419-35, 526, 527, 656, 658, 660,
661.
psittaous, 516.
ptilion, 602.
pyrocbles, 475, 476.
pyromelas, 456.
Pyrrhosticta, 538.
quadratus, 424, 445, 451, 452.
radiatus, 596.
rhamases, 475.
rhameses, 475.
rhamses, 475.
rhesus, 475.
rhetus, 605.
rhipidius, 533.
rbodostictus, 607, 610, 612, 618.
rogeri, 606.
rosenbergi, 625, 626.
rubroeinctus, 714, 717.
rurik, 679, 681.
rurikia, 681.
rutulus, 581, 587-90, 592.
sadalus, 541, 542.
sadyattes, 477, 478.
salvini, 738-40.
santhonus, 491.
sardalus, 541.
scamander, 426, 623, 628, 629, 631,
Schafferi, 419.
scheba, 695.
sebastianus, 679, 680.
septemlineatus, 705.
serapis, 472, 480, 483, 484, 487, 505.
serion, 692.
527, servillaei, 736.
serville, 419, 734, 735, 736, 737, 740.
servillei, 419, 734, 736.
sesostris, 435, 459, 460, 461-3.
sererus, 484.
sinon, 690-3.
socama, 466.
541, sonoria, 493, 494.
soratensis, 651.
spartacus, 496.
specularis, 457-
spoliatus, 452.
stabilis, 642,
steinbachi, 445, 452, 453,
( 752 )
stenodesmus. 719, 720. 728.
stilbon, 488.
streckeriai.us, 515, 516, 519.
swainsonius, 701.
syedra, 648, •")49.
tailori, 571.
tarquinivis, 460.
tasso, 424, 622, 62.3.
tecmenes, 569.
telamonides, 4'2.S, 689.
teletilaus,7I.S, 719-724, 725.
telmosis, 485.
temenes, 569.
teneates, 482.
tepicua, 700.
tc.xanus, 597.
thelios, 486.
theophron, 575.
therameue.=, 608, 609.
therapes, 643.
theras, 621.
therodamas, 662, 663.
thersites, 572, 574.
thoantiades, 429, 559, 561.
thoantides. 561.
Ihoas. 412, 429, 554, 555, 557, 558, 659, 560,
562, 564, 568, 569, 57.3, 587, 593.
thrason, 566, 567.
thyamu.s, 523.
thyastes, 728, 729, 7.32.
thyastinus, 728, 729.
thymbraeu.><, (']56, 677, 678.
thymbraus, 678.
timia.1, 507, 508.
toImide,«, 619.
tolus, 619.
tonila, 504
torquatiuus, 61.5, 616.
torqiiatiH, .538, 613, 617, 618, 620 2.
to.xaris, 484, 485.
trapez.T, 669, 670.
trichopus, 479.
triopa.-!, 419, 432, 445, 449, 450, 453.
triopus, 450.
Troides, 433.
Troilides, 538.
troilus, 543, 585, 593, 594, 595-8.
trojanus, 622.
tromes, 742.
tros, 415, 4,58, 622.
tucamanus, 526.
tulana, 444.
tullus, 459, 461,488.
turnus, 582, 584, 58G, 587.
ulopas, 662.
ulopos, 662, 663.
ulysses, 423.
utahensis, 549.
varus, 528, 529-32, 534.
vercingetori.'i, 451,
vertumnus, 432, 451, 467, 469, 470, 471-3, 478,
480, 485, 488, 4S9.
victorinus, 634, 635, 630, 645
vilcanotua, 643.
villiersi, 514.
vincentius, 517.
Virginia, 668, 669.
viridimaculatus, 503, 506.
vulnerata, 636.
vulneratu.a, 636.
w.ilshi, 689.
wardscewicsi, 651.
warscewiczi, 644, 649, 650, 651, 654.
wasmuthi, 513, 514.
xantliia.s, 464.
xanthopleura, 632, 6.33,
xanticles, 694, 695, 696.
xenarchus, 662.
xenares, 484.
xeniades, 667, 668, 669.
xenodamas, 518-20,
xuthus, 414, 426, 539, 540, 687, 720,
xynias, 656, 670.
yuracares, 469.
zacinthus, 503,
zacynthus, 415, 498, 499, 502, 503, 504.
zaddachi, C52-4.
zagreus, 538, 623, 624, 625, 627, 654
zalate.a, 625, il26.
zelicaon, S.i'.i, 550, 551.
zelicayn, 551.
zestos, 460-2.
zetes, 510, 511, 515.
zeuxis, 474, 475, 485.
zolicaoD, 550, 551.
zonaria, 692, 693.
zoros, 729.
•Privttd bit Rattll, WaUon tt Yiney, Ld., London ami Ayleabmy.
( 753)
^{i\\:v^^
T
A NEW SPECIES OP GIANT TORTOISE.
By the Hon. WALTER ROTHSCHILD, Ph.D.
Testudo gouff^i.
HIS tortoise is at once distinguishable from (ji(janten, and its subspecies,
(hmdiiii and soumeirei, which form this group of the genus Testudo, by its
much more elevated carapace, which is also more declivous in front, mnch larger
scutes, and squarer front edge to the carapace.
But the most prominent and easily perceived distinctive character is the
develojiment of the scutes on the forelegs and hind part of head. On the legs
the scutes are raised and j)rotrnde separately, almost as prominently as in T. calcarata
of Abyssinia. On the head the scutes, instead of being sunk in the skin, stand
out like flattened knobs. These characters were even more ap])arent when the
animal was alive. There is a double nuchal plate. Caudal and marginal jilates
behind somewhat recurved, thus approaching those of T. daudini.
Skull considerably smaller than in an individual of T. giyantea of similar size.
Central alveola on ridges of upper jaw much closer at the front of jaw, 12 mm.
apart, as ojiposed to 15 mm. The front margin of intermaxillaries does not project
as far as in T. gigantea, so that the nasal opening does not slope so obliquely.
Palatal region less concave than in gigantea, and the central ridge more pronounced.
Edges of pterygoid much contracted posteriorly, while in T. gigantea they run
almost parallel. The inner edges of the alveolar surface do not meet at the anterior
extremity of the vomer, and form a short broad isosceles triangle between that and
the foramina palatina, while in T. gigantea these edges meet and form a long
narrow triangle. In this new species the basi-occipital is much raised above the
ex-occipital owing to the deeji concavity of the latter, while in T. gigantea it is
almost in the same plane. The posterior lateral jwrtions of the pterygoid which
join the quadrate are strongly concave, and sunk much below the posterior portion
(if the sharp edge of the jtterygoid, while in T. gigantea these portions are almost
flat, and are level with the sharp edge of the pterygoid.
Skull :
Greatest width, T. gouffci : \\ inches = 114 mm.
„ ,, 7'. gigantea : \\ inches = 114 mm.
Length to end of liasi-occipital, T. goujfii : 5i inches = 140 mm.
„ ., ,, 7'. gigantea : ()\ inches = 15S mm.
3| inches = 90 mm.
4 A inches = 103 mm.
: 4| inches = 104 mm.
4 inches = 102 mm.
49
( '54)
Carapace :
Length over curve, 02 indies = 1550 mm.
„ iu straigbt line, 491 inches = 1238 mm.
Breadth over curve, 6ii indies = loiii) mm.
Ila//. Therese Island, St. Anue's Channel, Seychelles Islands.
The carapace of the T. (/igantca used for comjiarison measured 40 inches in a
straight line, = 122;") mm. It is true T. gigantea of Schweiger, of which 7'. hololissa
Giinther is a synonym. This form inhabited the smallest of the three Aldabra
islands, while T. gigantea elephantina inhabited the northern large island, and
T. (laitdiiii is still found on the southern large island. T. soiimeirei inhabited one
of the smaller islands either of the Seychelles or neighbouring groups.
I
ADDITIONAL NOTES ON BIRDS FROM N.W. AUSTRALIA.
By ERNST HAETERT, Pii.D.
(Plate I.)
Amytornis woodward! Hart.
(Plate I. fig. 1, 6 ad.)
IN Nov. Zool. 1905, p. 225, I mentioned a series of a bird under the name
Amytornis Itousei. Mr. Bernard Woodward, curator of the Perth Museum,
has now kindly sent me a specimen of the true A. homei, and I find it quite
distinct from our series. I therefore described the birds from the Alligator River,
Aruhem Land, under the name of Amijtornis icoodicardi, iu honour of Mr. Woodward
{Bull. D. 0. Club xvi. p. 30, November 1905). In Amyt. housei, originally described
as Amytis housri (Rep. Kimberley Exped., Ajip. B, 1902) the tail is much shorter
(in the sj)ecimen before me 01 mm., against 106 to 114 in A. icood/rardi), the
throat is striated with black and white, each feather being white with black
margins, the malar region similarly marked, the chest, breast and abdomen are
light chestnut, the former only being slightly marked with buff shaft-lines, the
vent and under tail-coverts blackish with rufous shafts, and the iijiper wing-coverts
chestnut with whitish shaft-lines. In other resjiects A. Iwusvi is much like
A. icoodivardi (cf Bull. B. 0. Club xvi. pp. 30, 31).
Amytis housei is well figured on PI. XIII, Emu iv.
The generic name of these jieculiar little birds has been rather unlucky.
They were named Amytis by Lesson in 18:;l, but as this name had been
preoccupied by Savigny in 1820 for a genus of worms, Stejneger in 1885
created the name Amytornis, which fact being overlooked by Uberholser, this
author made another substitute f(U' Amytis, namely Difipliorillas, in 1S09.
In r.lOl Mr. Milligaii described a new sjiecies under the name gigiintura.
This name was changed first for megalura and then for ineyalurus by Sharpe
{Zoological Record for 1901, ji. 08, and Hand-list iv. j). 240) under the belief that
giyantura was a " vox hybrida." Fortunately, however, the original name is
comjiosed of two genuine jiure Greek words, and " megalura " as well as
" megalarus " are therefore under any circumstances useless synonyms. The fear
of Mr. Milligau (cf. Victorian Naturalist xx. p. 138) that Dr. Sharpe " forestalled"
him with his uamo megalurus is of course unfounded !
( 755 )
Colluricincla woodwardi Hart.
(Plate I. fig. 2.)
This interesting new species has been described in JVor. Zool. 190."i, p. 228,
and the male is figured on PI. I.
Gymnorhina tibicen longirostris Milligan.
In Nov. Zool. 1905, p. 230, I described as a new subspecies from the 8onth
Alligator River a G. t. longirostris. Shortly afterwards I found that Mr. Blilligan
had already named a Gymnorhina longirostris {Emu iii. p. 90) as a new species
from North-western Australia. From his description it is clear that he described
the same bird, which must therefore be called G. t. longirostris Milligan. It is
fortunate that I chose the same name, and thus avoided making a new synonym
iu spite of my overlooking the prior publication.
Mirafra javanica pallida Hall.
(Cf Noc. Zool. 19o:j, p. 237.)
In the text occurs unfortunately a slip. I have said : " We have also a male
from Swan Hill, Victoria, which is ])aler and greyer than typical korsfiehli, very
near to grisescens, but not qnite like it." Instead of f/ri.sc.scen.s I meant to say
pallida, as there is no such name as ffri.sescens.
NOTES ON BIRDS FROM THE PHILIPPINE ISLANDS.
By ERNST HARTERT, Ph.D.
(Plate II.)
PART I.
WE have in the Tring Museum considerable numbers of birdskins from the
Pliiiil)pine Islands, which are not yet worked out. The late William
Doherty collected industriously on Basilau and Palawaa ; the first collection made
by Mr. Goodfellow on Mount Apo in South Mindanao is preserved in Tring, and also
the first set of those collected iu the same districts by Mr. John Waterstradt. We
also have some skins collected at Ayala (Mindanao) by an orchid hunter.
I should have written an account of tliese birds — es])ccially those from
Mt. Apo — if I had not been waiting for a second and larger collection from there.
Mr. Goodfellow has made such a collection, but it has been sold to the British
Museum, and Mr. Ogilvie-Grant has published a full account in the //;/,s for 190(3.
I can therefore content myself with short notes on most of these forms, and
herewith publish the first of them. The rest will follow in A'oc. Zool. 1907.
(PSITTACI.)
Trichoglossus johnstoniae Hart.
Trichoglossus johnstoniae Hartert, Bull. B. 0. Club, xiv. p. 10 (l'J03— Mt. Apo) ; Acicullural
Magazine 1906, plate.
This wonderful new parrot was discovered at an elevation of 8000 ft., and it
was at similar altitudes that Mr. John Waterstradt and Mr. Goodfellow on his
second visit to Mt. Apo collected specimens.
( 756 )
Mrs. Johnstone's parrakeet is of special interest, because it seems to connect the
so-called jrcnera rsitfcuteles and Ptilosrlcra, while TricIwijlossuK is in ray opinion
not sejiarable from Psiftcuteles. Therefore I call Jolinstone's j)arrakeet simply
Trichoglossus, though I have little doubt that some ornithologist afflicted with
"furor genericns " will one day create a new generic term fcir it.
Cacatua haematui'opygia (P. L. S. Miill.).
Not a mountain bird, being found at Davao, Daliaon, and other places of the
plains. This species is widely distributed, being known from twenty-five islands
and islets of the Philippine Archipelago. Local forms have not been distinguished,
and it seems to me that there are no differences between birds from the varions
islands.
Doherty obtained it on Basilan, and described the iris as scarlet, feet dirty
purplish, bill whitish.
Prioniturus waterstradti Rothsch.
We have the three specimens collected by Mr. Waterstradt on Mt. Apo, it is
said at an elevation of 3000 ft., while Mr. Goodfellow sent several examjiles from
a height of 8000 ft.
There can be no doubt that this is the form of the mountains, where it seems
to take the place of P. (Uscurus. Three thousand feet, however, is not far from the
lowlands, and we should await further investigations before deciding if P. waterstradti
everywhere replaces discurus, or whether the two occur together and may be of
different stock.
Prioniturus discurus Vieill.
William Doherty obtained four specimens on Basilan, Mr, Goodfellow shot a
number at Davao, Piso, and Martina, in South Mindanao, and we have also received
one from Ayala. Doherty described the iris as dull chestnut, the bill dull whitish,
feet dull slaty blue.
1 have before me the type of " Prioniturus discurus (Vieill.) var. nov. sulu'-zisis "
(sic), described in Journ. f. Orn. 1890, p. 140. I am sorry to say tliat I must agree
with Mr. Worcester {Hand-list B. Philippine Is., p. 49, footnote) that tlie Snlu form
caunf)t be separated.
Tanygnathus lucionensis (L.).
PiiUacua luciotietisis Linnaeus, .S'(/.>i(. Nat. Ed. xii. 1. p. 140 (17GC — ex Brisson).
Doherty obtained a series of skins on Basilan, and Mr. Goodfellow others on
Mindanao, at Daliaon, Davao, and Piso. Mr. Waterstradt sent it from iSanta (h-nz
in S. Mindanao.
We have in the Tring Museum a considerable series from most of the islands
of the Philippine Archipelago. Mr. Ogilvie-Grant {Ibis 1896, p. 561) gave some
verv interesting notes on Tant/ynathus lucionensis, distinguishing between certain
groups, but fortunately without naming them. I have before me thirty-six
specimens from Luzon, Mindoro, Mariuduque, Negros, Panay, Cebu, Basilan,
Mindanao, Bongao, iSibutn, Samar, Leyte, Sulu Islands, Palawan, and the small
group of islets called Mantanani, near Borneo. I cannot see any constancy in the
differences mentioned by Mr. Grant for specimens from various islands. It is true
( 757 )
that our ('xani]il('s fVom Fjiizon liave very pale lilue Iiemls, lint sjiecinieiis from the
suutlR'ni islauils are souietiiues ([uitc similar. Tlu^ shade of blue on the crown varies,
ai)j)arently according to age, and it seeius to be lighter in freshly monlteil birds,
darker in worn birds. The length of wing is also variable and not constant on any
island. We have, however, a very remarkable specimen from Negros, in which the
bine on the crown is entirely bine from the nape, leaving only a narrow frontal line
green, and in which the cheeks too are bine.
Mr. Ogilvie-Grant (Ibia 189(i, p. 5<W) sejiarated two birds from the Mantanaui
islets, near Borneo, nnder the name of Tanygnathus salvadorii. Count Salvadori
had correctly mentioned these birds {Cat. B. xx. p. 426) as yonng birds. Mr. Grant
"thinks this is a mistake," because the two birds have about as much blue on the
crown as old birds ; but we very often find that in birds certain portions of the
plumage moult irregularly before others, and evidently Mr. Grant is mistaken
rather than Count Salvadori, as two adult birds from the Mantanani islets in the
Tnng Museum show, which have the lesser upper wing-coverts black and the rest
exactly as in specimens from other islands.
With regard to Mantanani it is very interesting to read Everett's letter in the
Ibis, 1888, p. 282.
The amount of blue on the rump also varies considerably, but I have not been
able to localise these variations.
The occurrence of T. lucionensis on the Sangi Islands is in my opinion most
unlikely, and evidently due to an error in labelling in the Darmstadt Museum. I
do not hesitate to regard falaute/isis as a subspecies of lucionensis and to name it
trinomially (cf. Meyer & Wiglesworth, B. Celebes, i. p. 14.5).
Bolbopsittacus lunulatus mindanensis (Steere).
This form is evidently a bird of the lowland regions, for it was found at
Davao, Daliaon, Piso, Surigao. Butuan, Zamboanga, Ayala, and on the small
island of Panaon, liut not on Mt. Apo.
The three Bolbopsittacus from Luzon (B. lunulatus lunalatus), Leyte and
Samar {B. lunulatus intermedius), and Mindanao {B. lunulatus mindanensis) are
as typical subspecies as can be, representing each other on three different
islands, and being so much alike that they can only be distinguished if they
are closely examined.
B. I. mindaiiriisis was scjiiiriitcd as long ago as ISOO, and duly described in
the Catalogue of Birds, xx. pp. 504 and 500. In 1905 (Bull. B. 0. Club xvi.
p. IT), Mr. Ogilvie-Grant redescrilied this subspecies as a " new species," fortu-
nately choosing the same name as Mr. Steere. He thus had similar luck as I had
with my (Ji/miwr/iina tibicen longirostris (anteJi., p. 755).
Loriculus philippensis dohertyi subsp. nov.
The late William Doherty sent 3 c? c? and 3 ? ? from Basilan, and I have
also examined some specimens collected there by Professor Steere.
This series shows at a glance a much stronger and more reddish golden-yellow
wash on the back than in the specimens from Mindanao. In the old males the
whole back appears to be golden orange, about the apical half of the feathers
being golden orange. In the females it is as strong, or nearly so, as in the males,
or in some specimens less developed, though always evident.
( 758 )
Type of L. philippensis dohert>/i : No. llsOS, Basilaii, .Tiimiary IHIIS, William
Uoherty coll.
It is somewhat dillicult to decide how many ol' the Loricnli should be
considered as snbspecifically related. There can be little donbt that the smaller
forms — i.e. exilig, ainahilis, auranfiifro//s, tener, and others — are not subspecies
of the larger ones, because they stand apart by coloration and size, and one
form, L. cxilis, occurs together with a large one, L. stiymatus, on Celebes.
/,. i/i(//c/i.<, of C'eylon, with its high beak, stands by itself, and so does galgulus,
with its dri-p bine patch on the crown, tenmlin with a blue throat-jjatch, and
pusiUits from .Java. On the other hand, there is good reason to link the
Philippine forms together into one group of representative forms, at least those that
I am acquainted with, and to call them L. phiUppenKis philippensis, L. philippensis
mindore/isis, L. p/iil/ppi'/isis (ipicnlis, L. philipp:Vtsis tvor'ccsti'ri, I,, pliilippensis
regulus, L. p/iilippcnsis (/oke/i>/i, etc.
I cannot see that the black bill of bonapartci and stigmatiix is anything
more than a subspecific character, according to my treatment, and I should
even link sclateri and ruber to this series of forms. Meyer & Wiglesworth's
considerations in their Birds of Celebes, i. p, 160, fF., are most interesting,
though I cannot agree to their wide separation of a " northern " and " southern "
group.
Loriculus philippensis apicalis Souance.
Lnrinihis apicalis Souance, Rci\ and Mnij. Zunl. 18.5G. pp. "220, 221 (Mindanao).
I have before me in the Tring Museum 9 adult specimens from Mindanao.
Mr. Goodfellow obtained it at Daliaon, Dovao, Piso, and Tandaya, and on
Mt. Apo at 8000 feet. Mr. Waterstradt sent specimens from Santa Cruz,
from 2000 feet on Mt. Apo. Steere obtained a series near Ayala. All these
specimens show some golden-yellow tinge on the back, very faint in most
females, but distinct in the males.
Rhipidura nigTOcinnamomea Hart.
(Plate II. fig. 1.)
IhiU. /}. 0. Ch,h xiv. p. 12 flilO.'?).
This fine and ]>erfectly new Uhipiilnra was obtained by Mr. Goodfellow at
an elevation of <S0O0 feet, and Jlr. Waterstradt sent a specimen from a similar
height. Its wing measures only 73-5 mm., while the type had a wing of
80 mm.
Goodfellowia miranda Hart.
(Plate II., fig. 2.)
am. IS. 0. ciiiii .xiv. p. 11 (100.^).
Mr. Goodfellow obtained several fine specimens during his first c.xpeilition, and
a large series on his second trip; while Mr. Waterstradt did not j)rocure it,
probably because he depended mostly on native hunters and did not ascend to
such elevations as Mr. Goodfellow.
(To he continued in Nov. Zool.. 1907.)
( 759 )
NOTES TO PLATES III. AND X.
By KARL JORDAN.
NYMPHALIDAE.
1. Morphopsis meeki It. & J., -Vot'. Xool. xii. p. 456. u. 2 (1905) (Augalmiig:i,
Brit. N. Guinea).
(n. 111. fig. 2. c?, 3, ?)
Mr. A. S. Jlcek discovered this species in the Owen Stanley Range at the
head-waters of tlie St. Josepli River. He lias recently met witli it again on the
north-east side of the range at the Upper Mambare River.
2. Morphopsis ula //'(/., I.e. p. 454. n. 1 (IDO.")) (Angahnnga).
(PI. III. fig. 13. (J, 14. ?)
This fine species has also been fonnd by the same explorer at both places
where mecki was obtained.
PAPILIONIDAE.
3. Troides chimaera Rothsch., I.e. si. p, 311. n. 1. t. 3. fig. 25. 5 (1904)
(Owgarra); i>l. & Jord., I.e. xii. p. 460. n. 9 (1905) (Angabunga, cJ ?).
(PI. III. fig. 1. S)
This species does not appear to be rare at higher altitudes, though it is difficult
to obtain in good condition on account of its high flight. It visits flowering trees,
like the allied species. The female varies considerably in the amount of black on
the abdomen and in the size of the white markings of the wings. In some females
the spots of the discal row on the forewing are all of nearly the same size, while
in the majority of specimens some of the anterior spots have disappeared. This
species occurs also at the Upper Mambare River, north-east side of the Owen
Stanley Range.
LYCAENIDAE.
4. Deudorix grandis R. k, J., I.e. xii. j). 465. n. Is (l(i(i5) (Angabunga).
(PI. III. fig. 9. 10. c?)
Nearest to 1). dohertiji Obcrth., Et. (VEnt. xix. p. 13. t. 3. fig. 10. i (Isut)
(Andaij, but easily distinguished by the reduced metallic area of the npperside and
the deep green band of the nnderside.
5. Hypochrysops meeki R. & J., I.e. p. 464. u. 17 (1905) (Angabunga).
(PI. III. fig. 16. i)
Mr. Meek met neither with this nor the previous insect at the Upper Mambare
River.
( vr.o )
CASTNllDAE.
i'>. Castnia icarus Cram., I'ap. Kxot. i. \^. 20. t. 18. fig. a. b (1775) (Snrinnm).
(PI. X. fig. 1-4)
These linii' t'orras, which orciir together, at least in Paraguay, belong doubtless
to one variable species. There appears to be nothing that distingnishos them
from one another but the amonnt of white on the hind wing. Tiie short subapical
band of fig. 1 is also no constant difference. The white form is eiulelechia Dmce.
UBANIIDAE.
7. Cyphura pardata Warr., Nov. Zool xii. p. (i(i. n. 14 (1906) (Angabunga).
(PI. X. fig. 15. S)
GEOMETRIDAE.
8. Milionia pericallis R. & J., I.e. p. 407. n. 21 (1905) (Angabnnga).
(PI. III. fig. 5. S)
Mr. Meek fonud a long series of this prett^v insect at Angabnnga, but did not
meet with it at the Mambare River.
9. Craspedosis desmiata iid., I.e. p. 468. n. 25 (1905) (Angabunga).
(PI. III. p. 6. (?)
M. Dicyclodes hieroglyphica Warr., I.e. xiii. p. 76. n. 34 (1906) (Angabunga).
(PI. X. fig. 13. c?)
11. Hypochroma purpurissa id., I.e. p. 77. n. 36 (1906) (Angabnnga).
(PI. X. fig. 14. c?)
This and the other Geometrides of PI. X. are reproduced by way of experiment :
we wanted to know if the three-colour process is sufficiently accurate for the
reproduction of insects of that size.
12. Anisogamia commaculata ?>/., I.e. p. 78. n. 40 (1906) (Angabnnga).
(V\. X. fig. 11. (J)
13. Chlorochroma latistriga id. I.e. p. 84. n. 53 (1906) (Angabnnga).
(P). X. fig. 6. c?)
14. Xanthorhoe cerasina iiL, I.e. ji. 104. n. 107 (1906) (Angabnnga).
(PI. X. fig. 5. 3)
15. Xanthorhoe interrufata id., I.e. p. 105. n. 110 (1906) (Angabnnga).
(PI. X. fig. 10. c?)
1'''. Arycanda fulviradiata id., I.e. p. 138. n. 179 (1906) (Angabunga).
(PI. X. fig. 9. J)
17. Moneta plenicolor id.. I.e. p. 159. n. 222 (1906) (Angabunga).
(PI. X. tig. 8. (?)
(761)
ARGTIIDAE.
18. Asura rhodina R. & J., l.c. p. 460. n. 27 (1905) (Angabnnga).
(PI. III. fig. 12. 6)
A series of specimens similar to the type were fomid by A. S. Meek at the
Upper Mamijare River in March and April 10()(i.
19. Caprimima metallica //>/., l.c. p. 47(J. n. 28 (1905) (Angabunga).
(PI. III. fig. 17. ?)
21 ». Caprimima aenea //'/., l.c. n. 29 (1905) (Aogabunga).
(PI. III. fig. 11. ?)
21. Neoscaptia albata Hrl., I.e. n. .30 (1905) (Angabunga).
(PL III. fig. 8. ?)
22. Neoscaptia aequalis ii<l., l.c. p. 471. n. 31 (lOOu) (Angabunga).
(PL III. fig. 7. ?)
The figures of the last two species and of C. aenea are much too pale blue.
Plate III. is reproduced by another process than Plate X. ; the result is not
satisfactory.
AGARISTIDAE.
23. Argyrolepidia palaea Ud., l.c. p. 473. u. 35 (1905) (Angabunga).
(PL IIL fig. 15. ?)
24. Xanthospilopteryx zeodita Niepelt (ubi ?).
(PL X. fig. 7. c?)
We have both sexes from Lulnabonrg, Kassai River, Congo Free State. A
German dealer lately submitted the species to us under the above name, which we
adopt to avoid confusion. The insect is nearest to A', niyridorsa Mab. (1890).
25. Andrhippuris caudaequina Karsch, Ent. Nac/ir. .\.\i. p. 353. t. 1. fig. 1. S
(1895) (Lower Guinea).
(PL X. fig. 12. (?)
We have several specimens from the Kassai River, Congo Free State.
NOCTUIDAE.
20. Buzara calodesma R. & J., l.c. \\ 474. n. 38 (10(i5) (Angabunga).
(762)
ERKLARUNG.
In seiner Arbeit " On the African Forms of the Genns Fycnonotm " {Soc. Zool.
xiii. p. 389) schreibt Dr. Hartert : " P. b. gabonensis is so much like yonng tricolor
that even Professor lieicheuow mistook the hitter for ffabo/u'/isis. He states that
gabonertsis extends to the Congo, on the strengtli of a bird collected at Slanyanga
by Bohndorff : but the specimens collected by ]5(ilindorff at Mauyanga are now
before me, and they are tricolor, but by no means gaboiit'iisis, though a young
specimen has the under tail-coverts very pale suli>hnr-yellow."
Alle drei in dem vorstehendcn Satzc von Dr. Hartert anfgestellten Behaup-
tungen sind unzutreffend. Weder ist der von Bohndoril' in JIanyanga gesammelte
Vogel, woranf meine Angabe sich bezieht, Pgcnonotus tricolor, noch hat Dr. Hartert
den Vogel vor sich gehabt, wie der Leser aus dem Inhalt des Satzes annehmen
muss, und ferner ist es fur den erfahrenen Ornithologen unmciglich, den iun-'en
Vogel von Pi/eiioiwtiis tricolor mit P. gabonensis zu verwechseln.*
leh wiederhole daher meine im Journal fiir Ornithologic 1887, s. 3Ul, und in
meinem Buche Z^/e Vogel Afrikas, iii. S. 419, gemachte Angabe, A9.t.% Pycnonotaa
gabonensis am Kongo vorkommt, wo er von Bohndorff in Manyanga gesammelt ist.
Das Belegstikk befindet sich im Berliner Museum. Ubrigens gibt Prof. Barboza
de Bocage das Vorkommen von P. gabonensis am Quillu und sogar in Kakonda in
Benguella an.
Dr. Reichenow.
* Mir liegen im Museum zu Tring ein ganz alter unil ein junger von Bobndorff zu Manyanga
gesammelter Vogel vor. Dafs der alte Vogel nicLt gahmnsis sondern tricolor ist, liegt auf der Hand.
Dass der jiinge dazu gehijrt unterliegt meiner Ansiubt nacli keinem Zweifel, obwol er gabonensis so
ahnlicb siebt, dass aucb ein erfabrener Oinithologe— selbst I'rof. Eeicbenow vor 29 Jabren— ihii leicbt
damit verwechseln kiinnte. Dr. Walter Eotbscbild kaufte vor langer Zeit, ebe iob micb in Tring befand,
von dem verstorbenen Naturalienh,andler Schneider in Basel eine Sammlung, von der er damals glaubte'
es wiire die von Bohndorff gesammelte Vogel.^ammlung. Nun wusste ich allerdings sehr wohl, dass
sich im Berliner Museum viele von diesen Vogein befinden, der Zabl nacb aber augenscheinlieh we'niger,
als in Tring, natiiilicb .aber glaubte icb, d.ass I'rof. Eeicbenow damals die ganze Sammlung vorgelegen
babe. Wie niir I'rof. Eeicbenow freundlichst mitteilt hat er nicbt alle Exeniplare geseben, was er aber
damals auch nicbt wusste. Es ist leicbt erklarlicli, dass ich die voti Bobndorff zu Manyanga gesammelten
und als fi/nionotiix guhmnim bestimmten Stiickc fiir die von Eeicjienow crwiihnten hielt. Icb bedauere,
dass ich versiiumte, Hcrrn I'rof. Eeicbenow zu fragen, ob sich dicselben noch in Berlin befaudcn.
Auf der anderen Seite bin ich auch heute noch nicbt davon Uberzeugt, dass P. r/ahoninsis und
tricolor rcgelmiissig nebeneinander wohncn. Eino Nachpriifung der von Barboza de Bocage crwiihnten
Stucke erscbcint mir nothwendig. Nach der 1 c. p. L'42 gcmachten Bemcrkung bin ich geneigt,
anzuncbmcn, dass dem Autor von Cacouda ebcn auch ein alter und ein junger tricolor (das letztercr gibt
er selbst zu) vorlagen, und dass auch das Exemplar von Quillu ein junger tricolor ist. Ich hoffe spiiter
auf die Formen der Gattung Pycnonolus und ihre Vcrbreitung zuriickzukommcn und die verschiedcnen
Ansicbten dariiber wenigstens teilweise zu kliiron.
B. Hartert.
( 763 )
IN J) EX,
[For Index to "A Rovision of the American Papilios," pp. 411 to 745, see pp. lit) (o 7.">2
abbreviiitus (Buteo), 40.
Acanthidops, 308.
Acantholipes, 264-6.
Acanthylis, 36, 37.
Accipiter, 382.
Acolutha, 106, 107.
Acontia, 210.
Acronycta, 198.
acuminatus (Hctcropygia), 303.
acuticauda (Hylophilus), 3.
— (Paohysylvia), 12.
acutipennis (Chordeiles). 60.
acutissima (Ophiusa), 257.
Ada, 317, 318.
Addaea, 63.
Adeta, 113.
adumbrata (Myrioblopliara), 142.
ad vena (Orthnocichla), 298, 299.
Aedia, 2.54.
Aegialitis, 53.
Aegithalmus, 386.
aegithaloide.s (Leptasthenura), 333.
Aegocera, 161.
aenea (Caprimima), 761.
— (Neoscaptia), 761.
acqualis (Stelgidoptery.x), 13.
aeiiuatorialis (Campylopterus), 375.
— (Heliothrix). 378.
aeqiunoctialis (Gcothlypis), 7.
— (Motacilla), 7.
aestiva (Dendroioa), 6.
— (Motacilla), 6.
— (Pyranga), 16.
— (Tanagra), 16.
aefhiops (Thamnophilu.s), 339.
affinis (Agyrtria), 376.
— (Dysithamnus), 2, 31.
— (Phaetornis), 374.
— (Phoethomis), 374.
agami (Agamia), 51.
— (Ardea), 51.
Agamia, ,'51.
Agolaius, 21.
agenoris (Malacopsylla), 172, 173.
— (Megapsylla), 171.
agilis (Lanius), 11.
— (Vireo), 11.
Aglaia, 15.
Agyrtria, 34, 59, 353, 37G.
Ajaja, 49.
ajaja (Ajaja), 49.
— (Platalea), 49.
alba (Lanius), 403.
albata (Neoscaptia), 761.
alberta (Diradopsis), 69.
albertinae (Calospiza), 357.
albescens (Synallaxis), 28.
albiapicata (Xanthorhoe), 103.
albibasali.s (Tolmera), 152, 153.
albibasis (Dirades), 68.
— (Syntaracta), 134, 135.
albibisecta (Tephroclystia), 129.
albicaudatus (Buteo), 46.
albicincta (Chaetura), 36.
albiclausa (Paralcis), 148.
albicollis (Caprimulgus), 38.
— (Falco), 46, 382.
— (Lcgatus), 23.
— (Leucoptcrnis), 382.
— (Nyctidromiis), 38.
— (Saltator), 17, 315, 316, 351.
— (TyranniLs), 23.
— (Unibitinga), 46.
albicomma (Eutelia), 229.
albicosta (Capot«na). 240.
— (Pseudacidalia), 217.
albicrissus (Tliamnophilus), 30.
albicristata (Euoymatoge), 115.
albifimbria (Comostola), 86.
albifusa (Anisogamia), 77.
albigularis (Sclerunis), 28.
— (Synallaxis), 28.
albimedia (Eucymatoge), 115
albimixta (Tripterida), 131.
albinata (Anisogamia), 80.
albinigra (lulocera), 155.
( :^4 )
albirostris (Tanagra). '25.
albisoriata (Anisogamia), 78.
albisigillata (Cultripalpa). 275.
— (Zethes), 275.
albisparsa (Gonodcla), 154, 155.
albispcc'ularis (Tachyphonus), 17.
albi.-^fiuaina (Nasica). 20, 30.
albivcnter (Cicotrygon), 384.
— (Hirumlo), 12.
— (Tachycincta), 12.
albivcntris (Agyrtria), 59.
— (Astur), 292.
— (Myrmcciza), 33.
— (Orchilus), 22.
— (Ortalis), 350, 351.
— (Ramphocaenus), 32.
albivertex (Trochistis), 153.
albiviridis (Myrioblephara), 144.
alboarcuata (Eugorna), 269.
albogriseus (Pachj-rhamphus), 58.
albolineata (Corgatha). 214.
albolineatiis ( Dendrocolaptes), 30.
— (Picolaptes), 30.
albonotatus (Caprimulgus), 296.
albo-serrata (Acantholipes), 266.
alboplagiata (Catephia), 252.
albopunctata (Cultripalpa), 276.
— (Gonopteronia). 239.
— (Seneratia), 272.
— (Zethes), 276.
albostigmata (Euryglottis), 178.
albostriata (Paragarista), 255.
albotessellata (Capotena), 241.
Alcedo, 41.
Alois, 139, 140.
Alcyone, 294.
algeriensis (Lanius), 393, 397.
Alibama, 221.
allectus (Mecoeerus), 408.
alterata (Paralcis), 148.
altemata (Propithex), 96.
altirostris (Dendrocolaptes), 59.
aluensis (Dissolophus), 200.
amadis (Xylophanes), 183, 184.
amaura (Azelinopsis), 156.
amaurochalinus (Turdus), 6.
Amazona, 2, 44.
amazonica (Amazona), 2, 44.
— (Sporophila), 58.
amazonicus (Fnrmic.arius), 373.
— (Picumnus), 349, 350, 3.52.
— (Psittaous), 44.
— (Thamnophihis), 366.
amazonum (Hetcropclma), 303.
— (Scotothonis), 363, 364.
— (Thlypopsis), 311.
ambiguus (Caprimulgus), 296.
americana (Alcedo), 41.
— (Ccrylc), 41.
— (Sporophila), 3.
amethyst ina (Calliphlox), 60, 379.
amethystinus (Trochilus), 379.
AmpeHs, 28. 364.
Amphipyra, 191.
Amytis, 754.
Amytornis, 7.54.
Anabatcs, 335.
Anacretes, 320.
anahs (Pliysopterus). 408.
analogu.s (Laniua), 402, 403.
Anas, 54.
anatum (Falco), 46.
Ancaroides, 196, 197.
andaccola (Upucerthia), 331, 351.
andicola (Leptasthenura), 333.
andinus (Empidonax), 25.
andosa (Polyptychus), 179, 180.
andrei (Dysithamnus), 2, 31, 32.
andrewesi (Sintor), 408.
Andrhippuris, 761.
androcli (Malacopsylla), 172, 173.
angolensis (Loxia), 19.
— (Oryzoborus), 19.
angulata (Cosmophila), 236.
— (Plasmaticus), 268.
angulatiUnea (Episparis), 268.
angulilinea (Sypna), 247.
angulosa (Temnora), 182.
angustipennis (Coenooalpe), 96.
angustirostris (Muscicapa), 321, 322, 351.
— (Museicapara), 321.
— (Phylloscartes), 321. 322, 351.
anhinga (Plotus), 51.
ani (Crotophaga), 43.
Aniserpetes, 114.
Anisocolpia, 109.
Ani.sogamia, 77-82, 88. 760.
annabcllae (Myzomela), 301.
annchus (Polyptychus), 179.
antcnnata (Chactolopha), 93.
Anthela, 191.
Anthierax, 109.
anthoides (Corythopis), 373.
anthonyi (Lanius), 399.
Anthoscaenus, 36.
Anthoscopus, 386, 387.
Anthracothorax, 35, 377.
Anthus, 307, 351.
Anticlea, 93.
aplianta ( Dondrocincla), 29.
apicalis (Loriculus, 758.
apicebrunnea (Zethes), 275.
apifa.scia (Eutolia), 229.
approximans (Cercomacra), 343, 344.
( 765 )
aquilus (Fregata), 303.
— (Pelecanus), 303.
arabicus (Lanius), 399.
aracari (Pteroglossus), 380.
Aramides, 52.
Arbelorhina, 8, 10.
arctitorquis (Pachycephala), 300.
arcuosa (Sarrothripa), 225.
Ardea, 49-51, 303.
Ardetta, 51.
Ardonis, 114.
arenaceus (Empidochanes), 56.
Arenaria, 53.
argentata (Scateria), 344.
argentea {Westermannia), 218.
argentipuncta (.-Vsthena), 107.
Argopsylla, 170.
argus (Eurostopus), 296.
Argyrolepidia, 161, 761.
ariel (Ramphastos), 380.
Arizelomyia, 388.
aroa (Acantholipes), 265.
— (Acontia), 210.
— (Ceromacra), 285.
— (Cosmophila), 236.
— (Ilattia), 199.
— (Oraza), 215.
— (Pangrapta), 277.
— (PaniUa), 251.
— (Pseudodeltoida), 287.
— (Rivula), 211.
— (Sarrothripa), 224.
— (Thyria), 199.
— (Zethes), 273.
Aroana, 280, 281.
aroana (Acantholipes), 265.
— (Perigea), 194.
aroensis (Anisocolpia), 109.
— (Anthierax), 109.
— (Tathodelta), 212.
Arremon, 311-4, 351, 358.
arsinoe (Pycnonotus), 389-92.
Artamus, 302.
arthuri (Pachycephala), 299.
Arundinicola, 21.
arvensis (Fringilla), 309.
— (Sicalis), 309, 351.
Arycanda, 138, 760.
Asarcia, 53.
asinina (Trochi-stis), 153.
assimilis (Lanius), 393, 395, 397.
— (Mionectis), 22.
• — (Rhynchocyclus), 24.
Astheoa, 107.
Astur, 292.
Asturina, 382.
astygonua (Protambulyx), 179.
Asura, 761.
ater (Cassidix), 360.
aterrimus (Knipotegus), 318.
atlantica (Chiroxiphia), 3, 363.
Atmoceras, 140.
atricapillus (Anthoscopus), 387.
— (Orchilus), 22.
— (Tachyplionus), 17.
atriceps (Myiarchus), 324.
atricilla (Lams), 55.
atricollis (Trogon), 380.
atrilineata (Sauris), 112.
atripennis (Caprimulgus), 296.
atriplena (Syntaracta), 134, 135.
atrirostris (Dendrocincla), 336, 337, 352.
— (Dendrocolaptes), 330, 352.
atrogularis (Myrmotherula), 349, 352.
atronitens (Molothrus), 2, 20.
— (Xenopipo). 1, 325.
atroserieeus (Ramphocelus), 15, 16
atrostrigata (Hydrelia), 108.
Attila, 328-30, 351.
aucheri (Lanius), 397-9.
audacis (Geoeichla), 298.
— (Muscicapula), 296.
audax (Muscieapa), 24.
— (Myiodynastes), 24, 25.
aurantiifrons (Hylophilus), 12.
— (Pachysylvia), 12.
aurantiivertex (Heterocercus), 32V.
auratisquama (Steirophora), 113.
aureiplaga (Tibiocillaria), 232.
auricapilla (.Setophaga), 7.
auricapillus (Basileuterus), 7.
auricrissa (Tanagra), 15.
auricularis (Orchikis), 22.
auriculatus (Heliothrix), 378.
— (Lanius), 401, 402.
aurifrons (Muscieapa), 327.
— (Neopelma), .327, 351.
Auriparus, 386.
auritus (Heliothrix), .378.
aurocapilla (Pipra). 27.
australasia (Halcyon), 294.
austraHs (Euryst4:)mus), 303,
— (Rhopalopsyllus), 174.
Automolus, 335, 3.52, 365.
Avocettula, 353, 377.
avola (Erygia). 246, 247
— (Risoba). 234.
— (Sarrothripa). 225.
axillaris (Aramides), 52.
— (Myrmothera), 32, 368.
— (Myrmotherula), 32, 368, 369.
Axiocteta, 278, 279.
azarae (Canis), 175.
— (Saltator), 314, 315, 351.
( 766 )
Azelinopsis, 156.
azurca (Alcyone), 294.
babooni (Axiocteta), 278.
— (Blenina), 223.
— (C«romacra), 284.
— (Dipterygia), 197.
— (Plusia), 270.
badius (Lanius), 401.
baeri (Buarremon), 309, 351.
— (Compsospiza), 309, 351.
Bagada, 208. 209.
bahamensis (Momotus), 3, 40, 41.
— (Prionites), 40.
bainbridgei (Libyoclanis), 180.
bairdi (Acanthidops), 308.
Baniana. 264.
Banisia, 63, 64.
Barasa, 221, 222.
barbata (Muscicapa), 362.
— (Prosthetopteryx), 123, 124.
barbatus (Myiobius), 362.
— (Pycnonotus), 389-91.
Bartramia. 54.
basalis (Euryglottis), 178.
— (Pharambara), 65.
Basileuterus, 7.
Basitropis, 409.
bassetti (Zosterops), 300, 304.
batavica (Psittaca), 45.
— (Urochroma), 45.
batis (Anisogamia), 78.
bella (Cultripalpa), 276.
— (Pseudozalissa), 250.
— (Sypna), 247.
— (Zethes), 276.
bcnnigscni (Xenoccrus), 410.
bcntet (Lanius), 404.
bergii (Sterna), 289, 302.
berlepschi (Myrmotherula), 369.
— (Tanagra), 3, 15.
Berrcsa, 199.
bicolor (Accipiter), 382.
— (Amphipyra), 191. ,
— (Clanis), 180.
— (Dacnis), 11.
— (Eiiethcia), 3.
— (Heterocnemis), 354.
— (lodis), 88.
— (Microcereulus), 354, 355.
— (Sparvius), 382.
— (Sylvia), 11.
bicolora (Zagira), 213.
bidentatus (Falco), 46.
— (Harpagus), 46.
bifasciatus (Ceratopsyllus), 176.
bifvilvata (Xanthorhoe). 103.
bilincata (Acantholipes), 264.
— (Polioptila), 316, 317.
bipartita (Ptycbopoda), 92.
biplagiata (Callyna), 217.
bipunctata (Corgatha), 213.
birchalli (Catharus), 1.
biseeta (Pcrixera), 90.
bistrigata (Pereiana), 206.
bitorquatus (Pteroglossus), 380.
Blaeicus, 25.
Blenina, 222, 223.
bogdanowi (Lanius), 402. 403.
bogotensis (Columba), 383, 384.
boiei (Myzomela), 301.
Bolbopsittacm, 757.
boliviana (Cbiroxiphia), 363.
— (Leptasthenura), 333.
bolivianus (Attila), .3:!0, 351.
boliviensis (Herodon), 174.
boobook (Ninox), 293.
Borbacha, 134.
Borbotana, 207, 208.
borcalLs (Lanius), 393-5, 399.
boreas ( Dysdaemonia), 189.
Botaurus, 51.
bouvronides (Pyrrhula), 58.
• — (Sporophila), 58.
bouFFonoides (Spermophila), 58
bracbytarsus (Blaeicus), 25.
— (Empidonax), 25.
bracbyiira (Acanthylis), 37.
— (Chaetura), 37.
brasiliana (Strix), 45.
brasilianum (Glaucidium), 45.
brasiliensis (Alcedo), 41.
— (Attila), 328-30, .351.
— (Chelidoptera), 3H1.
brevicauda (Myrmotherula), 369.
brovipennis (Myiarcluis), 26.
brevipes (Cyanerpes), 10.
— (Heteractitis), 290.
brevirostris (Arltelorliina). 8.
bridgesi (Upueerthia). 331, 351.
Brixia, 64.
brunnea (Focillodes). 282.
brunneata (Eucymatoge). 116.
brunneicosta (Sarrothripa), 225.
brunneistriga (Polydcsnia), 251.
bninneotincta (Eueymatoge). 115, 110.
Buarremon, 351.
Bueeo, 380, 381.
Bunaea, 189.
Bursada, 138, 139.
buryi (Lanius), 397, ,399.
Buteo, 46.
(767 )
Butorides, 51.
Buzara, 761.
cabanisi (Empidochanes), 56.
— (Lanius), 404.
Cacatua, 756.
Cacicus, 20, 359, 360.
caelonota (Deva). 271.
caerulea (Arbelorhina), 8.
— (Ardea), 50.
— (C«rthia), 8.
— (Coereba), 9.
— (Cyanerpes), 2, 8-10.
— (Eucephala), 377.
— (Florida), ,50.
— (Pica), 305.
eaerulescens (Saltator). 315, 351.
eaeruleus (Chlorestes), 377.
— (Cyanocorax), 305-7, 351.
— (Trocliihis), 377.
cassia (Hypochroma), 77.
ca«siata (Gonodela), 154, 155.
— (Prosthetopteryx), 123, 124, 126.
caesius (Thamnomanes), 367, 368.
cajanea (Aramides), 52.
— (Fuliea), 52.
caledonica (Nycticorax), 291, 303.
Callidrepana, 61.
Callingura, 227.
calliope (Pachycephala), 299.
CalUphlox, 60, 379.
CalUspiza, 14.
Calliste, 14, 357.
callistus (Arremon), 311, 351.
Callyna, 217, 218.
calodesma (Buzara), 761.
Calospiza, 2, 14, 357.
Calothorax, 59.
Camptostoma, 361.
CampylopteriLS, 3, 59, 375.
cana (Tanagra), 15.
canadensis (Pitylus), 359.
— (Thamnophilus), 31.
Cancroma, 51.
canescens (Empidochanes), 56.
— (Lanius), 403.
canibrunnea (Epiplema), 69.
canicep.s (Lanius), 403.
canicosta (Acolutha), 106.
canigularis (Sclerurus), 28.
Canis, 173, 175.
canis (Ctenocephalus), 175.
canticua (Cyclarhis), 11.
capensis (Anthoscopus), 386.
— (Pycnonotus), 390, 391.
capitalia (Ramphocelus), 15.
Capnodeg, 284.
Capotena, 240, 241.
eaprata (Pratincola), 298
Caprimima, 761.
Caprimulgus, 37, 38, 296.
Capsiempis, 322, 323, 351
Caradrina, 201, 202.
carbo (Lanius), 357.
— (Ramphocelus), 357.
Carduelis, 58.
Carea, 242, 243.
Careades, 243-5.
carens (Bursada), 138.
caribaeus (Chlorostilbon), 35.
caripensis (Steatornis), 138.
cameata (Borbacha), 134.
Carpophaga, 289, 302.
carri (Synallaxis), 2, 29.
casbiata (Epiplema), 69.
casius (Tardus). 4.
easpius (Anthoscopus), 387.
Cassicus, 20, 359.
Cassidix, 21, 360.
castanea (Heterodisca), 157.
castaneata (Comostolodes), 87.
castanotis (Taeniopygia), 304.
Castnia, 760.
Catephia, 251^.
catephioides (Targalla), 227.
Catharus, 1.
Catocala, 246.
caudacutus (Sclerurus), 305.
caudaequina (Andrhippuris), 761.
caudata (Capsiempis), 323, 351.
— (Serpophaga). 323, 351.
caudatus (Lanius), 404.
cavicola (Pulex), 170, 174.
cavilinea (Prosthetopteryx), 123, 125.
Cavis, 174.
cayana (Calliste), 357.
— (Calospiza), 357.
— (Dacnis). 10. 356.
— (Euphonia), 357.
— (Motacilla), 10. 356.
— (Piaya). 2, 43. 44.
— (Tanagra), 357.
— (Tityra), 27, 364.
cayanensis (Myiozetetes), 361.
cayanus (Cueulus), 43.
— (Lanius), 27.
cayenncnsis (Euphonia), 357.
— (Hirundo), 37.
— (Panyptila), 37.
— (Tanagra), 357.
cela (Cacicus), 20, 359.
— (Parus), 20, 359.
celebensis (Caprimulgus), 296.
( '68)
celebensis (Compsogene), 179.
Celeopicus, 40.
Celeus, 2. 39, 40.
cenchroidcs (Falco), 292.
— (Tinnunculus), 292.
centralasiae (Anthoscopu.s), 387.
C'eophloeus, 40.
Cephalopyrus, 380.
cera-sina (Xantliorhoe), 104, 105, 760.
Ceratophyllus, 170.
Cferatopsylla, 176, 186.
Ceratopsyllus, 175-7, 186, 187.
Cercomacra, 59, 343, 344, 352, 370
Ceromacra, 284, 285.
certa (Ercheia), 248.
C'erthia, 8, 9, 28.
certhia (Dcndrocolaptes), 366.
— (Leptasthenura), 333.
— (Siptornis), 333.
Certhiola, 7.
certior (Ophiuaa), 256.
cervicalis (.\utomolus), 335, 352.
— (Philydor), 335, 352.
cervinieauda (Tlirenetes), 374.
Cervus, 175.
Ceryle, 2, 41.
cestonii (Nyctinomus), l8s.
Cha«tocercus, 59.
Chaetolopha, 93-6.
Chaetura, 1, 36, 37, 60.
Chalcophapa, 289.
chalybea (Hirundo), 13.
— (Progne), 13.
Chamaepelia, 384.
C'haradrius, 53, 290, 303.
charon (Ercheia), 249.
Cha,sraorhynchus, 28.
Chelidoptera, 381.
cherriei (Cyancrpes), 9.
chimaera (Troides), 759.
chionopectus (Agyrtria), 2, 34.
— (Thaumatias), 34.
Chiromachaeris, 27, 363.
chiron (Xylophancs), 184, 185.
Chiroxiphia, 3, 363.
chivi (Sylvia), 11, 317.
— (Vireo), 11, 317, 351.
Chlorestes, 35, 377.
chloricterus (Orthogonys), 310, 351.
— (Tachyphonus), 310, 351.
chloris (Halcyon), 295.
Chlorochroma, 83-6, 760.
Chloroclystis, 114, 115, 123, 127, 128.
Chloroenas, 383.
Chloronerpes, 38, 379.
Chlorophanes, 1, 10.
Chloropicus, 39.
Chloropipo, 324-6, 351.
chloroplaga (Egnasia), 279.
chloropyga (Cocreba), 7, 356.
Chlorospingus, 17.
Chlorostilbon, 35.
Chogada, 140, 141.
choliba (PiHorhina), 45.
— (Strix), 45.
Chordeiles, 60.
chry.iobronchos (Polytmus), 36.
— (Trochilus), 36.
chrysocephaliis (Scotothorus), 326.
chryeochlora (Chaleoiihapa), 289.
chrysochlorus (Trogon), 380.
Chrysococcyx, 295.
chrysogaster (Chlorospingus), 17.
Chryaolampis, 36, 377.
chrysomelas (Nemosia), 317.
chrysopis (Nemosia), 310, 311, 351.
Chrysoptilus, 60.
Chrysuronia, 59.
Ciccaba, 45.
Cinclus, 395.
cincta (Dichrozona), 348, 352.
— (Ptilinopus), 288, 302.
cinctus (Cyphorhinus), 348.
— (Microcerculus), 348.
cinerascens (Rynchops), 55.
cinerea (Lathria), 364.
— (Terekia), 290.
cinereella (Epiplema), 70.
cinereiceps (Thamnophilu.s), 366.
cinereiventris (Chaetura), 1, 37.
— (Myrmotherula), 368.
— (Pachyrhamphus), 27, 28.
cinereus (Cnipolegus), 319, 351.
— (Crypturus), 385.
— (Knipolegus), 319.
— (Paehyrhaniphus), 58.
— (Tetrao), 385.
cingulata (Urochronia), 45.
cinnamomea (Certhia), 28.
• — (S3Tialla.xi.s), 28.
einnamomeus (Picus), 39.
einnamomina (Nino.x), 293.
cirratus (Picumnus), 349, 3.50, 3.52.
eirrhatus (Thaiunophilus), 31.
Cisticola, 298.
citrinella (Zosterops), 3CX).
Clanis, 180.
Claravis, 47.
clarus (Troglodytes), 3, 6.
elathrata (Syntaraeta), 134, 135.
cleophontis (Rhopalopsyllus), 174.
Cletthara, 224.
Cnipolegus, 318, 319.
coalei (Myiarchus), 323.
( 769 )
Coccyzus, 60.
cochlearia (Caneroma), 51.
cocyti (Rhopalopsyllus), 174.
coeca (Letchena), 64.
Coenocalpe, 96-8.
Coereba, 7-9, 356.
coeruleata (Xanthorhoe), 1(>4.
coerulescens (Paralcis), 147.
— (Saltator), 314, 315.
— (Tephroclystia). 129.
coeruleus (Chlorestes), 35.
— (Trochilus), 35.
cognatus (Lanius), 402.
Colibri, 1, 35.
collaris (Acanthylis), 36.
— (Trogon), 41.
Collix, 98.
Colluricincla, 755.
collurio (Lanius), 401, 402.
Collusa, 191.
Coloeus, 388.
Colopteryx, 360.
colorata (Lobophysa), 158, 159.
— (Syncosmia), 128.
Columba, 2, 46-8, 383, 384.
Columbiana (Piaya), 2, 44.
eolumbianus (Pyrrhococcyx), 44.
Columbigallina, 46, 384.
Columbina, 384.
Colymbus, 52.
C'omatibis, 400.
commaculata (Anisogamia), 78, 760.
commixta (Arycanda), 138.
— (lodis), 89.
C'omostola, 86.
Comostolodes, 87.
conipar (Poljrptyohus), 180.
conipectiiiata (Chogada). 141.
eomplicata (Myrioblephara), 142.
Compsogene, 179.
Compsospiza, 309, 351.
Compsothlypis, 6.
compunctalis (Microbelia), 64.
coneinna (Alois), 139.
— (Carpophaga), 289.
concitus (Pulex), 174.
concolor (CVenularia), 280.
— (Felis), 175.
concolora (Axiocteta), 279.
concoloris (Ctenocephalus), 176.
— (Pulex), 170, 175.
configurata (Epiplema), 70.
confluens (Macroglossum), 407.
confundens (Perigea), 194.
confusa (Adeta), 113.
conjunctiva (Problepsiodes), 92.
connectens (Ramphocelus), 16.
connexa (Plutodes), 137.
Conopophaga, 353, 373.
consimilis (Polyptychus), 180.
consobrinus (.Anthoscopus), 386, 337.
— (Dendrornis), 29, 30.
con.spicillatus (Pelecanus), 292.
constellata (Eucymatoge), 117.
contaminata (Eucymatoge), 117.
continen talis ( Veniliornis), 3, 39.
Contortivena, 283.
conurus (Dasypus), 173.
Corgatha, 213, 214.
coronatus (Anthoscopus), 387.
corrasa (Diradis), 68.
Corvus, 302, 388.
C'oryphospingus, 359.
Corythopis, 373.
Cosmia, 210.
cosmius (Xylophanes), 183.
Cosmophila, 236, 237.
Cosmorhoe, 122.
costilinea (Reticulana), 205.
costimacula (Craspedosis), 139.
costipallens (Gadirtha), 220.
coetiplaga (Aedia), 254.
— (Baniana), 264.
— (Capnodes). 284.
costistrigata (Ingura), 226.
couchi (Tyrannus), 26.
Crasilogia, 99.
Craspedoprion, 361.
Craspedosis, 139, 760.
crassifemur (Gymnoscelis), 122.
crassirostris (Chrysococcyx), 295.
— (Loxia), 19.
— (Oryzoborus), 19.
— (Rallus), 52.
— (Tringa), 291.
crassistriga (Stesichora), 67, 68.
Crax, 48.
Creciscus, 385.
Crenularia, 280.
crenulata (Acronycta), 198.
— (Caradi-ina), 202.
— (Labanda), 221.
— (Parepisparis), 269.
Crex, 53.
crissalis (Formicarius), 33.
cristata (Muscicapa). 318, 351.
cristatus (Lanius), 402.
cristigera (Polysphalia), 123.
th-ithagra, 309. 351.
crosbyi (Ceratopsylla), 187.
Crotophaga, 42, 43.
crucigera (Pisoriiina), 45.
— (Strix), 45, 46.
Crypturus, 49, 385.
50
( 770 )
Ctenocephalus, 175, 176.
cubanus (Xylophanes), 185.
cucuUatus (Anthoscopus), 387.
— (Coryphospingus), 359.
— (Spinus), 56.
tHiculus, 43. 295. 304, 381.
eujubi (Pipile), 48.
Cultripalpa, 275, 276.
cumanensis (Penelope), 48.
— (Pipile), 2, 49.
euneatus (Glyphorliynchus), 366.
cimeigera (Madagara). 160.
cuneilinea (Chloroclystis), 114.
cura^oensis (Xanthornus), 20.
Curruca, 317, 351.
curucui (Trogon), 41.
eurvilinea (Paralci.s), 147.
curvimacula (Tephroclystia), 129.
curvinota (Microbelia), 64, 65.
cyanea (Certhia), 9.
— (Cyanerpes). 9, 10, 356.
— (Hyloeharis), 376.
— (Pseudelydna), 242.
Cyanerpes, 2, 8-10, 356.
eyanescens (Ingura), 226.
cyanicollis (Galbula), 380.
Cyanicterus, 310.
*cyanicterus (Cyanicterus), 310.
cyanirostris (Ada), 318.
— (Fluvicola), 317, 318.
— (Knipolegus), 318, 319, 351.
— (Muscicapa), 318, 351.
cyanocephala (Aglaia), 15.
— (Dacnis), 356.
— (Tanagra), 1, 15.
Cyanocorax, 305-7, 351.
cyanopus (Numcnius), 303.
cyanorostris (Ada), 317.
cyanurus (Trogon), 42.
cyanus (Hyloeharis), 376.
— (Trochilus), 376.
Cyclarhis, 11.
Cyclodes, 254.
Cyclorhis, 11.
cypereti (Rallus), 52.
Cyphorhinns, 348.
Cyphura, 66, 760
Cypseloides, 60.
C'ypselus, 37.
Daenis, 10, 11, 317, 351, 356.
Dammeria, 297.
dammeriana (Halcyon), 294.
— (Pachycephala), 299.
darwini (Lanius), 404.
Dasycephala, 328, 330.
Dasypus, 173.
dauuricus (Coloeus), 388.
dealbata (Epiplema), 71.
dealbatus (Lanius), 392, 393, 395-8.
debilis (Turdus), 5.
decens (Tephroelystia), 130.
deeorata (Anisoganiia), 79.
decunianus (Ostinops), 19.
— (Xanthornus), 19.
decussata (Pisorhina). 46.
deformis (Paralcis), 148.
defulvata (Myrioblephara), 143.
Deilephila, 181.
delectans (Xenoclystia), 132.
deletoides (Eutelia), 230.
delicata (Egnasia), 279.
— (Episteira), 1()9.
— (Risoba), 233.
— (Xenoclystia), 133.
delphinae (Colibri), 1, 35.
— (Orni3m3'a), 35.
Demiegretta, 291, 303, 403.
Dendrocincla, 29, 336-8, 352.
Dendrocolaptes, 29, 30, 59, 336, 337, 352, 366.
Dendrocops, 29.
Dendroica, 6.
Dendroplex, 59.
Dendrornis, 3, 29, 30, 59, 366.
Dendrotliripa, 226.
densinotata (Gonodela), 154, 155.
dentifascia (Nyctipao), 250.
dentifera (Ardonis), 114.
dentilinea (Ophiusa), 257.
Dermatophilus, 170.
desmaresti (Calliste), 14.
— (Calospiza), 14.
dcsmiata (Craspedosis), 760.
despecta (Epiplema), 71, 72.
detecta (Epiplema), 72.
Deudorix, 759.
Deva, 271.
devillii (Arrcmon), 312, 313, 351.
Diactinia, 99.
Diallactes, 30.
Diaphorillas, 754.
Dieaeum, 300, 301.
dichrourus (Lanius), 402, 403.
Dichrozona, 348, 352.
Dictyophora, 162, 169.
Dicyclodes, 75, 76, 760.
Dicma, 286.
dilataria (Ochyria), 100.
dinawa (Acantholipes), 265.
■ — (Aeronycta), 198.
— (Bagada), 208.
— (Borbotana), 208.
— (Catephia), 252.
( 771 )
dinawa (Catocala), 246.
— (Ceromacra), 285
— (Egnasia), 280
— (Elusa), 200
— (Eublemmoides), 212
— (Euplexia), 193.
— (Eurois), 196.
— (Eutelia), 229.
— (E.\yra), 226.
— (Focillodes), 282.
— (Ophideres), 269.
— (Oruza), 215.
— (Penicillaria), 231.
— (Perigea), 194
— (Phanaspa), 216.
— (Plusia), 270.
— (Xanthoptera), 210.
dinawaensis (Tibiocillaria), 232.
dinellianus (Hapalocercus), 321.
— (Pseudocolopteryx), 321.
Dinumma, 217.
Dipterygia, 197, 198.
Dipthera, 203.
Diptheroides, 203, 201.
Dirades, 68.
Diradopsia, 69.
discata (C'hiorochroma), 83.
— (Paralcis), 148.
discinota (Eucymatoge), 117, 119.
discipunctata (Callidrepana), 61.
diseora (Anas), 54.
— (Querquedula), 54.
Discosura, 379.
discriminans (Ophiusa), 259.
discuru.s (Prioniturus), 756.
dispar (Crasilogia), 99.
disrupta (Eucymatoge), 118.
Dissolophus, 200.
Disticta, 277, 278.
distincta (Hypotliripa), 224.
distorta (Eucymatoge), 118.
doddi (EpipjTops), 162.
dodsoni (Lauiua), 397.
dohertyi (Loriculus), 757.
— (Pycnonotus), 390-2.
doliatus (Lanius), 30.
— (Thamnophilus), 30.
dolius (Xylophanes), 183.
dominica (Anas), 54.
dominicua (Charadriua), 303.
— (Nomonyx), 54.
dorsalia (Gerygone), 297.
dorsinotata (Hydrelia), 108.
Drymophila, 33.
duperreyii (Megapodius), 288.
Dysdaemonia, 189.
Dysithamnua, 2, 3, 31, 32, 339, 344, 352, 367.
Earias, 219.
ebusa (Iscliyja), 267.
ecauJatum (Todiroitruin), 22.
ecaudatus (Orchilus), 22.
Echidiiophaga, 170.
Egnasia, 279, 280.
Egretta, 50.
egretta (Ardea), 49.
— (Herodias), 49.
ekeikei (Acronycta), 198.
— (Anthela), 191.
— (Borbotana), 207.
— (Careadea) 244.
— (Ceromacra), 284.
— (CoUusa), 191.
— (Coamophila), 237.
— (Cultripalpa), 275.
— (Ercheia), 248.
— (Ophiuaa), 256.
— (Plusia), 270.
— (Stictoptera), 235.
— (TargaUa), 227.
— (Thoracolophotoa), 2G1.
— (ZaUssa), 211.
— (Zethes), 275.
elaeagni (Lanius), 402, 403.
elaina (Capotena), 240.
Elainea, 23, 361.
Elania, 58.
Elanoides, 46.
elegans (Celeua), 2, 39, 40.
— (Laniua), 393, 395, 397.
— (Mecocerus), 408.
— (Picus), 39.
— (Synallaxia), 334, 335, 352.
elegantula (Rhipidura), 297, 298.
elongatus (Ceratopsyllus), 176.
Elusa, 200.
Emberiza, 21.
emboloacia (Acontia), 210.
Empidagra, 323.
Empidochancs, 56.
Empidonax, 25.
Empidonomua, 362.
enicurus (Calothorax), 59.
enaipennis (Campylopterus), 3, 59.
Eoa, 293.
Epa, 192, 193.
Epiplema, 69-75.
Epipyrops, 162.
episcopus (PUoethornia), 375.
— (Tanagra), 3, 14.
Episparis, 268.
Epiateira, 109.
Epitherapis, 156, 157.
Ercheia, 248, 249.
eremita (Comatibia), 400.
( 772 )
Ereunetes, 54.
crnesti (Faico), 292, 293.
errabunda (Paralcidia), 145.
Erygan.sa, 246.
Erygia, 240, 247.
erythrocephala (Pipra), 27.
erythrocephalus (Parus), 27.
eiythrocercus (Muscicapa), 26.
— (Philydor), 335, 365.
erythrogenys (Psaris), 327, 351.
— (Tityra), .327, 328, 351.
erythromelas (.Ardea), 51.
'- (.\rdetta). 51.
— (Pitylus). 359.
erythronotos (Ornismya), 34.
erythronotus (Saucerottea), 2, 3, 34.
Erythrura, 301.
erythrurus (Myiobius), 362.
Esacus, 291, 303.
Eublemma. 212.
Eublemmoides, 212.
Eucephala, 377.
Eucymatoge, 94, 115-21.
Eudocimus, 49.
Euetheia, 3, 18, 19.
Eugoma, 269.
eupeplodes (Epiplema), 72.
Euphonia, 13, 14, 57, 356, 357.
Euplexia, 195.
Eurois, 196.
Eurostopu.s, 296.
euryalus (Protambuly.x), 179.
eurycles (Protambuly.x), 179.
Euryglottis, 178.
Eurystomus, 303.
euteles (Trichoglossus), 293, 303.
Eutolia, 228, 231.
examplata (Colli.x), 98.
excubitor (Lanius). 392, 393, 395, 397, 399, 400.
exempta (Ptychopoda), 92.
exililinea (Rhomborista), 89.
exilis (Cisticola), 298.
eximia (Arbelorhina). 10.
— (Cyanerpes), 10.
exquisita (Pipra), 353.
cxsul (Myrmeciza), 340, 341.
— (Myrmelastes), 340-2, 352.
Exyra, 226.
eytoni (Dendrornia), 360.
faceta (Piranga), 57.
falcataria (Careades), 244.
falcigera (Epiplema), 73.
Faloo, 46, 292, 293, 382.
fallax (Lanius), 397-9.
fasciata (Aniserpetes), 114.
fa-soiata (Muscicapa), 25.
— (Syntaracta), 135, 130.
fasciatus (Myiobius). 25.
— (Sublegatus), 56.
feUciae (Saucerottea). 2, 3, 34.
Felis, 175.
felis (Ctenocephalu.s), 175, 176.
— (Pulex), 175.
festiva (Perixera), 90.
Ficedula, 388.
flammiceps (Aegithalus), 386.
— (C<?phalopjTus), 386.
Flatinae, 162, 168.
flava (Axiocteta), 279.
— (Carea), 242.
flaveola (Capsiempis), 322, 323.
flavicapilla (Chloropipo), 325, 326.
flavicorpus (Tephroclystia). 130.
flav'ifimbria (Comostola), 86.
flavifrons (Auriparus), 386.
— (Bursada), 138.
flavigula (Chloronerpes), 379.
flavilimes (Chlorochroma), 83.
flavilinea (Anisogamia), 80.
tlavipectu.s (Cyclarhis), 11.
flavipes (Hylophilus), 3.
— (Pachysylvia), 12, 350.
— (Seolopax), 54.
— (Totanus), 54.
— (Turdus), 57.
flavirostris (.\iTemon), 313, 314, 351.
flaviventre (Camptostoma), 361.
flaviventris (Hapalocerous), 320, 321.
— (Muscipeta), 23.
— (Ochtlioeea), 25.
— (Rhynchocydus), 23.
flavivertex (Heterocercus), 326, 327.
flexifascia (Epiplema), 73.
flexilinea (C'haetolopha), 94.
fiorestan (Protoparce), 178.
Florida, 50.
Horis (.Spizaetua), 303.
Florisuga, 34, 375.
fluriatilis (Sterna), 55.
Fluvicola, 21, 317, 318.
Focillodes, 281-3.
Fodina, 266.
Formicariua, 33, 373.
Formicivora, 3, 56, 334, 369.
fosteri ( Dysdaemonia), 189.
— (Protoparce), 178.
— (Xylophanes), 182, 183.
fragilis (lodis). 88, 89.
fraterculus (Tbamnopliilus), 30.
Fregata, 303.
frenata (Peristera), 48.
Fringilla, 14, 16, 18, 309, 356.
( 773 )
frontinus (Ophiusa), 259.
fulgurata (Chaetolopha), 94.
Fulica. .52. .'iS.
fuliginiceps (Leptasthenura), .332, 333.
fuliginosa (Dendrocincla), 33G, 337.
— (Euctheia). 18, 19.
— (Fringilla), 18.
— (Miiscicapa). 388.
fulminans (Gonopteronia), 239.
fulva (Careades), 244.
— (Focillodes), 283.
fulvata (Syntaracta), 136.
fulvescens (Gerygone), 297.
— (Nemosia), 311.
fulviceps (Nemosia), 311.
fulvigularis (Myiobius), 362.
fuh'inotata (Xanthorhoe), 104.
fulvipuncta (Micromia). 122, 130.
fulviradiata (Arycanda), 138, 760.
fulvisecta (Paralcis), 149.
fulvistriga (Ochyria), 100.
fulvivena (Myrioblephara), 143, 144.
fulviventris (Myrmotherula), 349.
fulvus (Charadrius), 303.
fumiccps (Siculodes), 6.5.
fumigata (Dendrocolaptes), 336.
fumigatus (Turdus), 4, 353.
fumipennis (Cra.silogia), 99.
fumosa (Phonipara), 18, 19.
funerea (Caradrina), 201.
funereus (Lanius), 395.
furcata (Thalurania), 377.
furcatoides (Tlialurania), 377.
furcatus (Elanoides), 46.
— (Falco), 46.
fuscantaria (Callyna), 218.
fuscatus (Empidochanes), 56.
— (Lanius), 404.
— (Pholus), 181.
fuscicapilla (Pachysylvia), 12.
fiiscofulvus (Picus), 39.
fu.scoinarginata (Zethes), 272.
fusconifa (Rliipidura), 297.
fuscostrigata (Bagada), 209.
gabonensis (Pycnonotus), 389, 390, 762.
Gadirtha, 220.
gairaardii (El.ainea), 23, 361.
— (Muscicapara), 23, 361.
Galbula. 42, 380.
galeata (Crex), 53.
— (Gallinula), 53.
— (Motacilla). 360.
galeatus (Colopteryx), 360.
Gallinula, 53
Garaeus, 157.
garleppi (Compsospiza), 309.
garraani (Gallinula), 53.
Garzetta. 303.
Gelasma, 87, 88.
gemini (Caradrina), 202.
geminipuncta (Clilorochroma), 83.
gcnibarbis (Thryothorus), 355.
— (Xenops), 366.
Geocichla, 298.
geoffroyi (Ochthodromus), 290, 303.
Geopelia, 289, 302.
Geositta, 318.
Geothlypis, 7.
Geotrygon, 3, 48, 384.
Gerygone. 297.
gestroi (Isehnopsyllus), 187.
gibbosa (Petrodava), 156.
gigantea (TeUulo), 753, 751.
giganteus (Lanius). 399.
gigantura (Araytomis), 754.
gilvus (Mimus), 3.
glaber (Sublegatus), 56.
glareola (Totanus), 303.
Glaucidium, 45.
Glaucis, 33, 374.
glaucus (Thamnomanes), 367, .368.
Glottis, 290. 303.
Glyphorhynchus, 366.
godmani (Leptopogon), 322, 323, .351.
Gonodela, 154.
Gonophaga, 158.
Gonopteronia. 239.
gouffei (Te.studo), 7-53.
gouldi (Lophornis), 353, 379.
— (Ornismya), 379.
gracilirostris (Myiarchus), 324, 351.
— (Tyrannus), 323.
gracilis (Canis), 175.
Gracula, 304.
Grallina, ,304.
gramineus (.Anthracothorax), 3.5.
— (Lamjjornis), 35, 377.
— (Trochilus), 35, 377.
gr.anadensis (Carduelis). 58.
Granatcllus, 355.
grandis (Deudorix), 759.
Graucalus, 298, 304.
grimrai (Lanius). 397.
grisea (Formicivora), 369.
— (Hyalospectra), 62.
— (Loxia). 18.
— (Risoba), 234.
— (Sarrotliripa), 225.
— (Sporophila), 18.
griseata (Encymatoge), 119.
— (Focillodes), 282.
griseipeotus (Myrmeciza), 33,
( "4 )
grispisticta (Mus-i'apa), 388.
griseistriga (Zet!ies). *273.
griseiventris (Hypocncmis), 371-3.
— (Pachysylvia). 350.
— (Pithys)", 371, 372.
griscola (Ancaroidcs), 197.
— (Barasa). 221.
— (Columbigallina), 384.
— (Columbina), 384.
griseolauta (Sauris), 1 12.
griseonigra (Labanda). 221.
griseonotata (Anisogamia), 80.
griseus (Canis). 173.
— (Caprimulgus), 37.
— (Xyctibius). 37, 38.
— (Phloeomimus), 409.
— (Sittasomus), 3.
— (Turdus), 369.
Grison, 175.
grossa (Loxia), 359.
grossiventris (Malacopsylla), 171-3.
— (MegapysUa), 171.
— (Pulex), 170. 171, 173.
— (Sarcopsylla), 173.
grossus (Pitylus), 359.
guadeloupensis (Saltator), 315, 316, 351.
guatemalensis (Chlorophanes), 10.
guayabambae (Myrmotherula), 349, 352.
guianensis (Chaetura), 1, 37.
— (Coereba), 7.
— (Hylocharis), 376.
— (Leistes), 21.
— (Oriolus), 21.
— (Phoethornis), 374.
— (Piaya), 43, 44.
— (Pyrrhococcyx), 44.
Guiraca, 359.
guttata (C'allispiza), 14.
— (Calospiza), 14.
— (Myrmotherula), 369.
— (Ortalis), 3.")0.
gutturali.s (Fringilla), 18.
— (Sporophila), 18.
guy guy (Phaethornis), 2.
guy (Phoethornis), 34.
— (Troc-liihis), 34.
gymnophtliahnus (Turdus), 5.
Gymnorliina, 755.
GymnosceHs, 122.
gyrola (Caloapiza), 14.
Habrissus, 409.
hades (Dinumma), 217.
haemalea (Piranga), 57.
Haematopus, 303.
haematuropygia (Cacatua), 756.
hacmorrhous (Cacicus). 360.
Halcyon, 294, 295.
Halia.stur, 292.
hampsoni (Acantholipes), 265.
— (Disticta), 277.
— (Eut<'lia), 228.
hampsonia (Catephia). 253.
Hapalocercus, 320, 321, 351.
Haplospiza, 308, 309, 351.
Harpagus, 46.
Hastina, 107.
hau.xwelli (Myrmotherula), 369.
— (Turdus), 353.
heckelii (Cyanocorax), 306, 351.
Hectopsylla, 170, 171.
Hedymela, 388.
Hehothrix, 378.
hellmayri (Myrmotherula), 369.
Hemichehdon, 388.
hemileuca (Hypocnemis), 348, 352.
hemileucurus (Lanius), 395.
Hemiprocne, 36.
Hemithraupis, 317.
henrici (Dammeria), 297.
hepatica (Pyranga), 57.
herbertus (Xenocerus), 410.
Herodias, 49.
Herodon, 174.
Heteractitis, 290.
Heteranax, 298.
Heterocercus, 326, 351.
Heterocnemis, 33, 344, 345-7, 352, 354.
Heterodisca, 157.
Heteropelma, 363.
Het«ropygia, 290, 303.
heudei (Paradoxornis), 387.
hieroglyphica (Dicyclodes), 76, 760.
hilarii (Crithagra), 309, 351.
hirsuta (Glaucis), 33, 374.
Hirsutipes, 263, 264.
hir.sutus (Trochilus), 33, 374.
Iiirtivena (Coenocalpe). 97.
hirundinis (C<"ratopsyllus), 177.
Hirundo, 12, 13, .36, 37, 296.
hiruntlo (Macroglossum), 407.
liolTjuanni (Formicarius), 33.
— (Myrmornirt), 33.
holTniannsi (Tliamnomanes). 367.
hollandi (Hapalocercus), 320, 321, ,351.
holochlora (Chloropipo), 325, 326, 351.
homeyeri (Lanius), 392, 395.
Homodes, 218.
housei (Amytis), 754.
hueti (Urochroma), 60.
humerata (Xenoclystia), 133,
Hyalospectra, 62.
Hydranassa, 2, 50.
( "5 )
hydrafa (Xylophancs), 183.
llydrclia, UI8.
Hylocharis, 37l».
Hylonympha, 1.
Hylophila, 219.
Hylophilus, 3, 12, 3.55.
Hypaetra, 262, 264.
hyperythra (Muscicapula), 296.
hyperythrus (Thryothoru.s), 6.
Hypocala, 249.
Hypochroma, 77, 760.
Hypochrysops, 759.
Hypocnemis, 344, .348, 349, 3.52, 353, 370-3.
hypoleucos (Totanus), 290, 303.
— (Tringa), 290.
hypoleueus (Graucalus), 304.
— (Myrmochanes), 348, 352.
hypomelaena (Cercomacra), 343, 352.
hypopyra (Ampelis), 364.
hypopyrra (Laniocera), 364.
hypospodius (Thryothorus), 6.
H3q5othripa, 224.
hypoxantha (Pachysylvia), 12.
Ibis, 291.
icarus (Castnia), 760.
icterocephalus (Agelaiu.s), 21.
— (Oriolus), 21.
icterope (Lamprote), 310.
icteropus (Pyranga), 310.
Ictinia, 46.
igniceps (Sootothorus), 326.
ignifera (Pipra), 362.
ignobilis (Turdus), .5.
Ilattia, 199.
illex (Tchitrea), 388, .389.
— (Terpsiphone), 388.
illibata (Ophiusa), 257.
illustris (Typhosia), 407.
imitans (Gelasma), 87.
immaculata (Myrmeoiza), 340, 352.
— (Myrmelastes), 340-2.
incanesoens (Ornithion), 361.
incei (Tchitrea), 389.
incertans (Elusa), 200.
ineertissima (Bagada), 209.
incertus (Dysithamnu.s), 367.
— (Thamnophilus), 339, 307.
incomta (Phaeomyias), 56.
inconspicua (Cosmia), 210.
— (Gadirtha), 220.
indicus (Habrissu.s), 409.
indigens (Perixera), 91.
indus (Haliastur), 292.
incrmis (Megapsylla), 172. 173.
inexpectatus (Cyanocorax), 305, 306, 35).
infusoatus (Anabates), 335.
infuscatus (.Vutomoliis), 335, 352, 365.
— (Lanius), 402, 403.
Ingura, 226.
innocens (Epiplema), 74.
inornatus (Colopteryx), 360.
— (Pyciionotus), 389, 390.
inquinata (.\Iyriol>lephara), 143.
inquisitor (Lanius), .328.
— (Tityia), 327, 328, 351.
inquisitrix (Tityra), 327.
insignifica (Caradrina), 201.
— (Opliiufsa), 258.
insignis (Ceratopsylla), 187.
— (Tachyphonus), 357, 358.
insulana (Piaya), 243.
insularis (Hylophilus), 3, 12.
— (Lcptotila), 47.
— (Ostinops). 19.
— (Platyrhynchus), 22.
— (Taeniopygia), 304.
intermedia (Formicivora), 3, 56.
— (Myrmcciza), 340, 352.
— (Myrmelast€s), 340, 341.
intermedins (Haliastur), 292.
— (Myrmelastes), 341.
interposita (Halcyon), 294.
interpres (Arenaria), 53.
— (Morinella), 303.
— (Tringa), 53.
interrufata (Xanthorhoi;), 10.5, 760.
intimalis (Microbelia), 64.
intromissa (Diactinia), 99.
invidens (Gelasma), 88.
lodis, 88, 89.
iridescens (Anisogamia), 81,
irritans (Pulex), 174.
irrufata (Aids), 140.
Isabella (Stiltia), 291, 303.
isabej'inus (Lanius). 402—4.
Ischnopsyllus, 176, 177. 186-8.
Ischyja, 206, 207.
isidori (Geratophyllus), 170, 177.
— (Ischnopsylhis), 177.
— (Vespertilio), 177.
Isoura, 261.
lulocera, 155.
jacana (Parra). 53.
jacapa (Raniplioccliis). 2, 15, 16, 357.
— (Tanagra), 1.5.
jacarina (T.anagra), 18.
— (Volatinia). 18, 359.
jacquini (Pipilc), 48.
jacutinga (Pipile), 48, 49.
jamaicensis (Nyctibius), 37, 38.
jardinei (Dendrornis), 3. 30.
— (Spodiomis), 308, 309.
(776)
javanica (Hirim(ii)), 296.
jaxartonsis (Antlioscopua), "87.
— (Remiza), 387.
jaxartica (Anthoscopus), 387.
johrLStoniae (Trichoglossus), 755.
jourdani (Chaetocercus), 59.
jubata (Ceratopsylla), 186.
junctiUnea (Paralcis), 149.
junianus (ThamDophilus), 339, 352.
karelini (Lanius). 403.
kaupi (Leucoptcniis), 382, 383.
kebea (Alibama), 221.
— (Ancaroides), 196, 197.
— (Dipterygia), 197, 198.
— (Disticta). 278.
— (Ereheia), 249.
— (Erygansa), 246.
— (Fodina), 266.
— (Ophiusa), 255.
— (Plusia), 270.
— (Risoba), 233.
— (Sarrothripa), 225.
kebeae (Borbotana), 207.
— (Callingura), 227.
— (Dipthera), 203.
— (Eutelia), 229.
— (Hypocala), 249.
— (Ischyja), 266.
— (Tarache), 208.
kebeana (Polychrisia), 271.
kebeensis (Cosmophila), 236.
— (Erygia), 247.
— (Zalissa), 211.
kebirensis (Pachycephala), 300.
— (Stigmatops), 301.
kenricki (Diptheroidcs), 203
— (Ophiusa), 2,59.
kirki (VeniliortiLs), 3, 39.
kirkii (Chloropicu.s), 39.
kisscrcnsis (Gcrygone), 297.
Knipolegus, 317-9, 351.
kopnigi (Lanius), 397.
kuhli (Leucopternis), 382, 383.
kiihni (Gerygone), 297.
Labanda, 220, 221.
lacrymans (Xenoccrus), 410.
lactea (Polioptila), 316, 317, 351.
laemo-sticta (Myrmeciza), 342, 343, 352.
Lagoptera, 260.
lahtora (Lanius), 393.
Lalage, 298, 304.
Lampornis, 35, 377.
Lamprote, 310.
lanccolata (Glauris). 374.
Lauio, 17.
Laniocera, 364.
Lanius. 11, 24, 25. 27, 30, 31, 328, 351, 357, Siili,
392-404.
Laphyctes, 26.
Larus, 55.
Lasiopoderes, 263.
Lasiosceles, 204.
Lathria, 364.
laticostata (Chlorochroma), 84.
latimedia (Paralcis). 150.
latirostris (Corrus), 302.
latistriga (Chlorochroma), 84, 760.
— (Prosthetopteryx), 123, 125.
latizona (Ophiusa), 259.
lawTencoi (Chaetura), 1, 37.
— (Empidonax), 25.
lawrencii (Lanio), 17.
layardi (Pycnonotus), 390, 391.
Legatus, 23.
leighi (.\egoccra), 161.
Leistes, 21.
leotaudi (Celeus), 2, 39, 40.
— (Chlorospingus), 17.
Lepidoenas. 383.
lepidonota (Hypocnemis), 370-2.
Leptasthenura, 321-33, 351, 352.
Leptopogon, 321-3, 351.
Leptotila, 47, 48.
lessoni (C!hlorostilbon), 35.
— (Spermophi)a), 58.
— (Sporophila), 58.
Letchena, 64.
lettiensis (PtiUnopus). 302.
— (Zosterops). 300.
leucocephala (Arundinicola), 21.
— (Pipra), 21.
Icucocilla (Pipra), ,363.
leucocyma (Megaloba), 110.
leucoga.ster (Ardea), .50.
— (Pionitcs), 382.
leuconota (Pyriglena), 370.
Icucopliragtna (Chaetolopha), 94.
leucops (Tardus), 3, 57.
Leucopternis. 382, 383.
k'ucoptcrus (Lanius), 392. 393, 395.
Icucopyga (Cavis), 174.
leucopygos (Lanius), ,397.
Icucorhynchus (Artamua), .302.
Icucorrhous (Psilomycter), 37.8.
leucostigma (Heterocnemis), 344, 34.5.
— (Percnostola), 346, 347.
— (Selateria), 345-7.
leucurus (Tlirenetcs), 374.
Libyoclanis. 180.
lichenca (Protoparce), 178.
( 777 )
lichenosa (Eucymatoge), 119.
lichtensteinii (Euphonia). 14, 356, 357.
— (Phonasca), 14. 3r,G.
lifuense (Macroglossum), 407.
lignigera (Bagada), 209.
lilacea (Dierna), 28fi.
— (Ophiui3a). 2.56.
— (Zethes), 272.
limnaetus (Spizaotua), 303.
Limnopardalu.s, 52.
Limonites, 290, 303.
Limosa, 290, 303.
limosa (Limosa), 290, 303.
— (Scolopax), 290.
linearis (Geotrygon), 3.
— (Peristera), 48.
lineata (Sporophila), 3.
llneatus (Ceophloeus), 40.
— (Picus), 40.
lineola (Sporophila), ,58.
linteatus (Heterocercus), 327.
Lipangus, 364.
litae (Chloropipo), 325, 326.
lobata (Remodes), 111, 112.
Lobivanellus, 291.
Lobophy.sa, 158, 159.
LocBstella, 304.
longicauda (Bartramia), 54.
— (Discosura). 379.
— (Tringa), 54.
longicaudus (Trochilus), 379.
longipennis (M3Tmotherula), 369.
longipes (Drymophila), 33.
— (MjTmeciza), 33.
longiplaga (Westermannia), 218.
longirostri.s (Anthoscaenus), 36.
— (Gymnorhina), 755.
— (Haematopus), 303.
— (Rallus), 2, .52.
— (Trochilus), 36.
longuemareus (Phaethornis), 2.
— (Phocthornis), 34.
— (Trochilus), 34.
Lophornis, 36. 353, 379.
Loriculus, 757, 7.58.
Lo.xia, 18, 19, 301, 359.
lucianus (Xylophancs), 184.
luoidata (Myrioblcphara), 144.
lucioncnsis (Tanygiiatlius). 756.
luctuosa (Myrmotherula), 309.
luctuosus (Tachyphomis), 17.
ludovicianus (Lanius). 399.
lugubris (Quiscalus), 21.
Lurocalis, 38.
luteocephala (Muscicapa), 326, 327, 351.
— (Pipromorpha), 327.
luteocephalus (Heterocercus), 326, 327, 351.
luteola (Certhiola), 7.
— (Cocreba), 7.
lutcsccns (Anthus), 307, 351.
lutzi (Pulcx), 175.
— (Rhopalopsyllus), 174.
Lycosphinx, 406, 407.
macconnelli (Picumnus), 349, 3.50, 3.52.
Maceda, 232.
mackloti (Dicaeum), 300, 301.
macrocerca (Hylonympha), 1.
Macroglossum, 407.
macronyx (Anthoscopus), 386, 387.
maerorhynchus (Corvus), 302.
macrothyris (Bunaea), 189.
macrourus (Caprimulgus), 296.
macularia (Tringa), 54.
macularius (Totanus), 54.
maculata (Addaea), 63.
maculatus (Limnopardalus), 52.
— (Mecocerus), 408.
— (Rallus), 52.
maculicauda (Hypocnemis), 348.
maculicaudis (Cercomacra), 59.
maculifer (.Myrmelastes), 340, 342.
maculipoiinis (Pygiptila), 367.
— (Thamnophilus), 367.
maculirostris (Turdus), 5.
maculosus (Turnix), 288.
mafalui (Cosmophila), 237.
magna (Problepsis), 92.
magnirostris (Esacus), 291, 303.
— (Phaetusa), 55.
— (Rampliocelus), 2, 15, 16.
— (Sterna), 5.5.
magnus (Saltator), 358.
Magulaba, 212.
major (C'rotophaga). 43.
— (Lanius), 392, 393, 395.
— (Tliamnoj)hi!us), 30, 366.
Malacopsylla, 171-3.
Malacoptila, 381.
malaris (Phocthornis), 374.
Manacus, 363.
manacus (C'liiromachaeris), 27, 363.
— (Manacus), 363.
— (Pipra), 27.
mango (Lampornis), 377.
manilensis (Caprimulgus), 296.
marcescens (Tolmera), 152.
Marcipa, 271.
marginata (Paralcidia), 146.
marginatus (Heterocnemis). 354.
— (Microcerculus), 354, 355.
martialis (Ischnopsyllus), 187, 188.
martinica (Fulica), 53,
( 778)
martinica (PorphjTiola), 53.
maugcus (Geoiielia), 289, 302.
meamsi (Lanius), 399.
Mecocerus, 408.
Mecodina, 272.
media (Calospiza), 14.
mcdioalbata {Opliiusa), 2."i9.
mcdiobrunnea (Dinumma), 217.
mcdiofusca (Eutelia). 228.
mediogriseata (Catephia), 252.
medionigra (Focillodes), 283.
mediopallens (t'orgatha), 214.
mediotaeniata (Eucymatoge), 120.
mediotincta (Chlorochroma), 86.
meeki (Berresa), 199.
— (Hypoclirysops), 759.
— (Morphopsis), 759.
— (Penicillaria). 231.
— (Perciana), 206.
— (Rectangulipalpus), 286, 287.
— (Rivula), 211.
— (Stictoptera), 235.
— (Zethes), 273.
megacephalus (RhjTichoc}'clus). 361.
Megaloba, 109-11.
megalorhynchos (Tanygnathus), 294.
megalura (Amytornis), 754.
Megapodius, 288.
Megapsylla, 171-3.
Megarhynchus, 25.
megaspila (Zethes), 273.
melaleueus (Taehyphonus), 17.
melanoholiciia (Tyrannus), 26.
Melanerpes, 3.
melanogaster (Thamnophilus), 368.
nielanogastra (Conopophaga), 373.
— (Myrmotherula), 369.
melanogenys (Faloo), 292.
melanoleuca (Tcrenura), 348, 352.
molanoleucos (Phalacrocorax), 303.
melanopleura (Morula), 57.
inelamipogon (Hypocnemis), 348.
nielanops (Graucalu.s), 298, 304.
— (Leucopternis), ,383.
melanoptera (Tanagra), 15.
mclanoptcrus (Minuis), 3.
melanopygia (Parra), 53.
melaniira (Paf'liyeoj)hala). 299.
— (Rynchops), 55.
melanuroidcs (Limosa), 290, 303.
melanurus (Ramphocaenus), 32.
— (Trogon), 379.
melas (Aedia), 254.
melicerte (Ophiuaa), 257.
mellivora (Florisuga), 34, 375.
mellivorus (Trochilus), 34, 375.
melpomene (Catharus), 1.
nienstnius (Pinnns), 44.
— (Psittacus), 44.
mcntalis (I)ysitharanus), 3, 31, 32, 367.
— (.Myothera), 367.
— (Pipra), 362.
meridiuiialis (Lanius), 400.
— (Trogon), 42.
Merula, 57.
merula (Dcndrocincla), 29, 337, 338.
meruloides (Dendrocops). 29.
mesoplaga (Sarrothripa), 224.
metallica (Caprimima), 701.
mexicana (Calospiza), 2, 14.
— (Tanagra), 14.
mcxioanus (8clerurus), 365.
Microbelia, 64. 65.
Microcerculus, 348, 354, 355.
Microeschus, 217.
Micromia, 122, 130.
microrhyncha (Coereba), 9.
— (Cyanerpcs), 9.
miorus (Pyononotus), 390, 391.
miles (Lobivane!lu.s), 291.
milionia, 760.
militaris (Emberiza), 21.
— (Leistes), 21.
— (Tanagra), 21.
Mimus, 3.
mindancnsis (Bolbopsittacus), 757.
minimus (Molothrus), 20.
minor (C'hlorochroma), 85.
— (Dendrocincla), 336.
— (Dendrocolaptes), 336.
— (Halcyon), 294.
— (Lanius), 397, 400, 401.
— (Pyononotus), 390, 391.
— (Sicalis), 309.
minuta (Capnodes), 284.
— (Loxia), 18.
— (Piaya), 43.
— (Sporophila), 18, 58.
minutilla (Tringa), .54.
Mionectes, 22, 58, 360.
mirabilis (Parelydna), 241, 242.
Mirafra, 755.
miranda (Goodfellowia), 758.
Misocalivis, 295, 303.
mollis (Lanius), 393-5.
Molothrus, 2, 20, 21.
molucea (Itjis), 291.
moluccensis (Tinnunculus), 292.
Momotus, 3, 40, 41.
Monarcha, 298.
Mona,sa, 381.
monaatica (Xanthorhoi?), 105.
monedula (Corvus), 388.
Moneta, 159, 760.
( "9 )
inontana (Lanius). 403.
montivagans (Lpptasthenura). 333.
moorei (Phaethornis), 374.
— (Phoethornis), 374.
Morinella, .303.
morphoeus (Monasa), 381.
Morphopsis, 759.
mosquitus (Chry.solampis), 36, 377.
— (Trochilus), .36, 377.
Motacilla, 6, 7, 10, 356, 360.
multicolora (Deva), 271.
multilinea (Cletthara), 224.
— (Zethes), 275.
mundus (Heteranax), 298.
— (Monarcha), 298.
Mania, 301.
Muscicapa, 21-7, 318-22, 326, 327, 330, 351,
360, 362, 388.
Muscicapara, 23, 321, 861.
Muscicapula, 296, 297.
Muscipeta, 23.
Muscisaxicola, 318, 319, 351.
Muscivora, 27.
musculi (Pulex), 176.
muaculu.s (Troglorlytes), 3, 6.
mutu.s (Saltator), 315.
Myiarchus, 26, 323, 32), 351.
Myiobius, 25, 362.
Myiodynastes, 24, 25.
Myiopatis, 23, 56, 361.
Myiozetetes, 23, 361.
Myothera, 367.
M3rrioblephara, 142-4.
Myrmeciza, 33, 340-3, 352.
Myrmelastes, 340-2, 352.
Jlyrmochanes, 348, 352.
Myrmornis, 33.
Myrmothera, 32, 368.
Myrmotherula, 32, 349, 352, 368, 369.
mystaceua (Platyrhynchus), 22.
nacunda (Podager), 00.
Nadagara, 160.
naevia (Ardea), 51.
— (.Muscicapa), 2.5.
— (Sclateria). 2, 33, 344.
— (Sitta), 33.
— (Tapera), 43.
naevius (Cuculus), 43.
— (Myiobiu.s), 25.
— (Nycticorax), 51.
napaeum (Ornithion), 361.
napensis (Tachyphonus), 358.
Nasica, 29, 30.
Nasua, 175.
nasuae (Ctenocepiialus), 175.
nasuae (Pulex), 170, 175.
nattereri (Lurocalis), 38.
nebularius (Glottis), 290, 303.
— (Scolopax), 290.
nechu.s (Xylophane.s), 18.5.
neglcctus (Coloeu.s), 388.
Nemosia, 310, 311, 317, 351.
Neopelma, .327, 351.
nieuwenhuisi (C'hrysococcyx), 295
niger (Pachyrliamphu.s), 27, 28.
— (Pachyrynchus), 27.
— (Tornosinus), 286.
nigra (Magulaba), 212.
— (Rynchops), 55.
— (Zethes), 274.
nigrans (Blenina), 222, 223.
nigrellata (Syntaracta), 136.
nigrescens (Barasa), 222.
— (Cercomacra), 344.
nigribasis (Eucymatoge), 120.
nigrican.s (Cercomacra), 59.
— (Petrochelidon), 296.
— (Pyennnotus), 390, .391.
nigrioauda (.Myrmeciza), 343, 352.
nigriceps (Myiarchus), 26, 323.
nigricollis (Anthracothorax), 35, 377
— (Euphonia), 57.
— (Lampornis), 35.
— (Trochilus), 35, 377.
nigrierissa (Piaya), 44.
nigrilinea (Zethes), 274.
nigripes (Ardea), 303.
— (Garzetta), 303.
nigriplaga (Acantholipes), 266.
nigrirostris (Coluraba), 383.
nigrithorax (Siculodes), 66.
nigroeinnamomea (Rhipidura), 75S.
nigropunctata (Corgatha). 214.
nigrostrigata (Bagada), 209.
nilgirica (Plutodes), 137.
niloticus (Lanius), 401, 402.
Ninox, 293.
nipalensis (Caprimulgus), 296.
nisoria (.Munia), 301.
nitida (Asturina), 382.
nitidifrons (Agj-rtria), 353, 376,
— (Tliaumati.as), 376.
nitidus (Falco), 382.
niveicosta (Dirades), 68.
nivifascia (Borbotana), 207.
Nomonyx, 54.
Noreia, 76.
Notophoyx, 291, 303.
novaeguineae (Yula), 193.
novaehoUandiae (Notophoyx), 21'. -303
— (Plotus), 291.
— (Scythrops), 295,
( '80 )
novagiiinensis (Prota^otis), 192.
noveboracensis (Jlotacilla), 7.
— (Seiurus), 7.
novogiiincana (Earias), 219.
— (Hypaetra), 262.
— (Mecodina), 272.
niibicus (Lanius), 400, 404.
nudigenis (Turdiis), 5.
Numenius, 290, 303.
Nyctanassa, 51.
Nycteribia, 176.
Nyctibius, 37, 38.
Nycticorax, 291.
nycticorax (Ardea), .51.
— (Nycticorax), 51.
Nyctidromus, 38.
Nyctinomus, 188.
Nyctipao, 2.50.
oberi (Myiarchus), 26.
obliqua (Axiocteta), 278.
— (Careades), 245.
obscura (Nyctipao), 250.
obscurior (Sclerunis), 365.
obscurus (Campylopterus), 375.
— (Ctenocephalus), 175.
— (Pulex), 170, 175.
obsoleta (Chlorochroma), 84.
— (Lobophysa), 158, 159.
occidentalis (Ereunetes), 54.
— (Microcerculus), 354. 355.
— (Myrmelastes), 341, .342.
— (Tinniinculus), 292.
occipitalis (Synallaxis), 28.
ocellata (Ninox), 293.
• — (Sporophila), 58.
ochotensis (Locustella), 304.
— (Sylvia), 304.
ochracea (Phanjuspa), 215.
ochraceus (Dissolophus), 2(X).
ochrciplaga (Eutelia), 231.
ochreistriga (Zcthcs), 274.
ochreovenata (Acantholipcs), 264.
ochrimixta (Gonodela), 154, 155.
Ochthodromus, 53, 290, 303.
Ochthoeca, 25.
Ochthophora, 220.
Ochyria, KJO, 101.
octactena (Ceratopsylla), ISfi.
octactcnus (Ceratopsyllus), 186.
— (Ischnopsj'llus), 186.
— (Vespenigo), 186.
ocularis (Eutelia), 231.
Oecophylla, 164.
oenone (Chryauronia), 59,
ocnoplcx (.Axiocteta), 278.
oleaginea (Muscicapa), 360.
oleagincus (llioncctes), 22, 360.
— (Muscicapa), 22.
oliva (Empidonax), 25.
olivacca (Aroana), 281.
— (Curruca), 317, 351.
— (Dendrocincla). 336, 338.
— (Myrioblephara), 144.
— (Sylvia), 317.
olivaceiplaga (Eutelia), 228.
olivaccus (C'raapedoprion), 361.
— (Dysithamnus), 32.
— (Mionectes), 58.
— (Rhynchocyclus), 361.
oliva,scpns (Basileutcrus), 7.
— (Saltator), 18.
olivens (Risoba), 234.
Olonia, 162.
ombiranus (Argyrolepidia), 161.
omissa (Euethcia), 3.
— (Synallaxis), 365.
ommatopis (Ri\nila), 211.
opalizans (Pipra), 363,
Ophideres, 269.
Ophiusa, 255-60.
ophthalmicus (Pogonotriecus), 322, 351.
Ophyx, 260.
Orchilus, 22.
orenocensis (Knigolegus), 318.
— (Serpophaga), 323, 351.
Orcta, 62.
orientalis (Eurystomus), 303.
— (Hylophila), 219.
Oriolus, 20, 21, .304, 360.
ornatipennis (Tephroclvstia), 131.
ornatus ( Lophornis), 36.
— (Trochilus), 36.
Omismya, .34, 35. 379.
Omithion. 23, 361.
Ortalida, 3.50.
Ortalis, 3, 3.50-2.
Orthnocichla, 298, 299.
Orthogouys, 310, 351.
orthographus (Polyptychus), 406.
Oruza, 215.
oryzivora (Cassidix), 21, 360.
— (Oriolus), 21.
oryzivorus (Oriolu.s), 360.
Oryzoborus, 19.
cssifragus (Corvus), 388.
Ostinops, 19.
Otomela, 403.
ottonia (Ptilinopus), 283, 302.
owgarra (Blcnina), 222.
— (Sinariola), 238.
oxyura (Acanthylis), 37.
( 781 )
Pachycephala, 299, 300.
Pachyrhamphus, 27, 28, 58.
Pachyrynchus, 27.
Pachysylvia, 12, 355, 350.
pagana (Elainea), 23.
— (Museicapa), 23.
palaea (Argyrolepidia), 761.
pallens (Lanius), 397.
palliatu.s (Lanius), 300.
— (Thamnophilus), 300.
pallida (Lanius), 403.
— (Mirafra), 7.55.
— (Munia), 301.
— (Ophiusa), 260.
pallidifrons (Hylophilus), 3.
pallidimargo (Paralcis), 150.
pallidiplaga (Remodes), HI.
paUidirostrLs (Lanius), 392, 393, 395-C.
pallidiventris (Mionectes), 22.
pallidivirens (Ptychotheea), 127.
pallidus (Lanius), 397.
paUiolatus (MisooaUus), 295, 303.
palmarum (Tanagra), 15.
palposus (Ceratopsylla), 187.
pamphila (Argyrolepidia), 161.
Pangrapta, 276, 277.
Panilla, 251.
panopus (Compsogene), 179.
Panurus, 387.
Panyptila, 37.
Papilio, 411-745.
papuensis (Garaeus), 157.
— (Gonodela), 154.
— (Hydrelia), 108.
Paracarea, 245, 246.
Paradoxomis, 387.
paraensis (Automolus), 365.
— (Granatellus), 355.
Paragarista, 255.
Paragonitis, 238, 239.
Paralcidia, 145-7.
Paralcis, 140, 147-51.
paraUelaria (Carea), 242.
paranensis (Leptasthenura), 333.
Parapsyllus, 174.
parens (Mionectes), 22.
pardaria (Burbacha), 134.
pardata (Cyphura), 00, 760.
Parelydna, 241, 242.
pareola (Chiroxiphia), 3. 303.
— (Pipra), 363.
Parepisparis, 269.
Parra, 53.
partita (Caradrina), 201.
Paras, 20, 27, 359.
parva (Microeschus), 217.
parviceps (Pulex), 175.
parvipennata (Prosthetopteryx), 123, 126.
parviplaga (Remodes), 111.
parvus (Anthus), 307.
passerina (Cliamaepelia), 384.
— (Columba), 384.
— (Columbigallina), 47, 384.
patiens (Oreta), 62.
paucalensis (Synallaxis), 334, 352.
paueilinea (Poecilasthena), 108.
pectoralis (Antlioscopus), 387.
— (Pachysylvia), 356.
pelagica (Chaetura), 36, 37.
pelecanoides (Sterna), 289, 302.
Pelecanus, 292, 303.
pella (Topaza), 378.
— (Trochilus), 378.
pelzelni (Granatellus), 355.
pendulina (Remiza), 387.
pendulinus (Anthoscopus), 386, 387.
Penelope, 48.
penetrans (Pulex), 170.
— (Rhynchoprion), 170.
Peuicillaria, 231.
Perciana, 206.
Percnostola, 346, 347.
perdricipennia (Catephia), 252.
peregrinus (Falco), 46, 292.
pericaUis (Milionia), 760.
Perigea, 194.
Peristera, 47, 48.
Perixera, 90, 91.
Perizoma, 95.
perlata (Pyirliura), 381.
— (Strix), 171.
peronii (Geocichla), 298.
persieus (Cacicus), 359.
— (Cassicus), 20, 359.
— (Oriolus), 20.
persitrigata (Lobophysa), 158, 159.
perspicillata (Pulsatrix), 45.
— (Strix), 45.
peruana (Monasa), 381.
peruvianus (Anthus), 307.
Petrochelidon, 290.
Petrodava, 156.
phaeochroa (Dendrocincla), 29, 336, 338.
Phaeomyias, 56, 323.
phacopygoides (Turdus), 2, 4.
phaeopus (Numenius), 290.
phaeopygus (Turdus), 2, 4, 353.
Pha«thornis, 2, 374.
Phaetornis, 374, 375.
Phaetusa, 55.
phalnolaema (Heliothrix), 378.
Phalacrocorax, 303.
phalaenoides (Glaucidium), 45.
— (Strix), 45.
(782)
Phanaspa, 215, 216.
Pharambara, 65.
phelpsi (Sitta-somus), 3.
Philydor, 335, 352, 36.5.
Phloeobius, 409.
I'hloeomimuR, 4()!).
phocniceus (Carduelis), 58.
Phoenicothraupi.s, 2, 17.
phoenicuroitles (Laniiia), 402-4.
phoeopigus (Turclus), 4.
Phocthomis, 34, 374, 375.
Pholus, 181,
Phonasca, 14, 356.
Phonipara, 18, 19.
Phrygilus, 309.
Phyllomyias, 56.
Phylloscartcs, 321, 322, 351.
Phylloxiphia, 407.
Physopterua, 408.
Piaya, 2, 43, 44.
Pica, 305.
pica (Fluvicola), 21.
— (Muscicapa), 21.
picata (Gracula), 304.
— (Grallina), 304.
Picolapt&s, 30.
pictaria (Acolutha), 107.
pictipennis (Chaetolopha), 95.
Picumnus, 349, 350, 352.
Picus, 38-40.
picus (Dendropk'x), 59.
pileata (Empidona.x), 25.
— (I.«ptasthenura), 333.
pileatus (Accipiter), 382.
— (Tinamus), 49, 385.
piimatus (Ardea), 51.
— (Botaurus), 51.
Pionites, 382.
PioniKS, 44.
Pipile, 2, 48, 49.
pipile (Ch-ax), 48.
— (Pipile), 2, 48.
pipistrellus (Vesperugo), 186.
Pipra, 21, 27, 353, 362, 363.
Pipromorpha, 327.
Piranga, 16, 57.
Pisorliina, 45, 46.
pitangua (Lanius), 25.
— (Megarliynchus), 25.
Pitangus, 2, 24.
Pithys, 371, 372.
pitiayiimi (Compsothlypis), 6.
— (Sylvia), 6.
Pitta, 296.
Pitylus, 359.
placens (Eucymatoge), 120.
placida (Deilephila), 181.
plagiostola (Corgatha), 214.
planimargo (Epiplema), 74.
Pliismaticus, 208.
Platalea, 49, 291.
platcnsis (Leptasthenura), 321, 333.
Platyja, 267, 268.
PlatjThynehus, 22, 23.
plenicolor (Moneta), 159, 760.
Plotus, 51, 291.
plumbea (Chloroenas), 383.
— (Columba), 383.
— (Dacnis), 11.
— (Ictinia), 40.
— (Sylvia), 11.
— (Syntaracta), 134, 135.
plumbeus (Dy.sithamnus). 345.
— (Falco), 46.
pliirabilinea (Xanthorhoe), 106.
plumipes (Zethes), 273.
plumosa (Atmoceras), 140.
Plusia, 270.
Plutodes, 137.
Podager, 60.
podiceps (Colymbus), 52.
— (Podilymbus), 52.
Podilymbus, 52.
Poecilasthena, 108.
poecilinota (Hypocncmis), 370-2.
Poecilodryas, 297.
Pogonotriccus, 322, 323, 351.
poliades (Polyptycbus), 400.
poliocephalus (Rhynchocyclus), 361, 302.
polionotus (Arremon), 311-4, 351.
— (Astur), 292.
— (Thamnophilus), 338, 339, 352.
— (Urospizias), 292.
Polioptila, 316, 317. 351.
poliourus (Acantbylis), 37.
— (Cyijselus). 37.
poliura (Chaetura), 37.
Polyclirisia, 271.
Polydesma, 250, 251.
Polyptycbus, 179, 180, 406.
Polysphalia, 123.
Polytmus, 36, 378.
pomeranus (Lanius), 400-2.
Porpliyriola, 53.
porphyrodes (Platyja), 267.
praeferrata (Dictyopbora), 102, 169.
Pratincola, 298.
pratti (Caradrina), 202.
— (Carea), 243.
— (Elusa), 200.
— (Epa), 193.
— (Hypaetra), 262.
— (Hypocala). 249.
— (Ingura), 226.
( 783 )
pratti (Isoura), 261.
— (Lagoptera), 260.
■ — (Lasiopoderes), 263.
— ( Lasiosceles), 205.
— (Pangrapta), 276.
— (Platyja), 267.
— (Polydesma), 2.50.
— (Pseudogonitis), 238.
— (Pseudophyx), 260.
— (Risoba), 234.
— (Sphingiforma), 241.
— (Targalla), 227.
— (Tibiocillaria), 231.
— (Zagira), 213.
pretiosa (Claravis), 47.
— (Peristera), 47.
princeps (Tehitrea), 388, 389.
Prionia, 160.
Prion ites, 40.
Prioniturus, 7.56.
Problepsiodes, 92.
Problepsis, 92.
Progne, 13.
Propithex, 96.
Prosthetopteryx, 123-7.
Protagrotis, 191, 192.
Protambulyx, 179.
Protoparce, 178.
przewalskii (Lanius), 394, 395.
Psaris, 327, 351.
Pseudacidalia, 216, 217.
Pseudelydna, 241, 242.
Pseudoclanis, 180.
pseudocollurio (Lanius), 402, 403.
Pseudocolopteryx, 321.
Pseudodeltoida, 287.
Pseudogonitis, 238.
Pseudophyx, 260.
Pseudozalissa, 250.
Psilomycter, 378.
Psittaca, 45.
Psittaci, 755.
psittaci (Hectopsylla), 171.
Psittacus, 44, 171, 293.
Pteroglossus, 380.
Ptilinopus, 288, 289, 302.
Ptychopoda, 92.
Ptychotlieca, 127.
pulchella (Och}Tia), 100.
pulchra (Corgatha), 213.
— (Cyelodes), 254.
pulchricolor (Prionia), 160.
pulchristrigata (Catephia), 253, 254.
Pulex, 170-6, 186.
pulex (Rhynchopsylla), 171.
— (Rhynchopsyllus), 171.
pullens (Lanius), 397.
Pulsatrix, 45.
pulverosa (Noreia), 76.
pumilus (Coccyzus), 60.
punctata (Paialcis), 149, 150.
— (Phanaspa), 216.
punotatissima (Steirophora), 113.
punctigula (Chrysoptilus), 60.
punctilligera (Chlorochroma), 84, 85.
punctipectus (Cbrysoptilus), 61).
punctistriga (Homodes), 218.
punctulata (Munia), 301.
punctuliger (Tliamnophilus), 339, 352.
pura (Barasa), 222.
— (Stesichora), 67.
purpurascens (Acantholipes), 265.
— (Chlorophanes), 1.
— (Sadarsa), 235.
pui'purea (Ceromaera), 285.
purpureonigra (Eutelia), 230.
purpureotincta (Columba), 383.
purpurissa (Hypocliroma), 77. 760.
purus (Chiromachaeris), 363.
— (Manacus), 363.
pusilla (Myiopatis), 23, 361.
— (Tringa), 54.
pusiUum (Ornithion), 23, 361.
pusillus (Ereunetes), 54.
Pycnoloma, 128.
Pycnonotus, 389-92, 762.
Pygiptila, 367.
pygmaeus (Phaetliornis), 375.
— (Phoetliornis), 375.
— (Pygmornis), 375.
Pygmornis, 375.
Pyranga, 57, 310.
Pyriglena, 343, 344. 348, 370.
Pyrrhoeoccyx, 44.
PjTrhoracliis, 89.
Pyrrhula, 58.
Pyrrhura, 381.
pyrrhus (Xylophanes), 185.
quartaria (Brixia), 64.
Querquedula, 54.
quinquelineata (Banisia), 63.
quintieolor (Loxia), 301.
— (Munia), 301.
Quiscalus, 21.
raalteni (Synoicus), 302.
raddei (Lanius), 402, 403
radicata (Bursada), 138.
Rallus, 2, 52.
ramonianus (Trogon), 42.
Rampliastos, 2, 42, 380.
( T84 )
Rampliocaenus, 32.
Ramphocanus, 32.
Ramphocelus. 2, If), 10, 357.
Rectangulipalpus, 286, 287.
rectilinea (Aegocera), 161.
rectilineata (Chaetolopha), 95.
— (Perizoma), 95.
recurvirostris (Avocettula), 353, 377.
— (Trochilus), 377.
refulgens {Tlialurania), 59.
regia (Platalca), 291.
reginae (Trochilus), 379.
reichenbachi (Celeopicus), 40.
— (Celeus), 2, 39, 40.
reichenowi (Pycnonotus), 390, 391.
— (Rhipidura), 297. 298.
Remiza, 386, 387.
Remodes, HI, 112.
renalis (Ilattia), 199.
Reticulana, 205.
reticulatus (Eos), 293.
— (Psittacus), 293.
Rhipidura, 297, 298, 758.
rhodina (Asura), 761.
rhododactyla (Megaloba), 110.
rhodophila (Carea), 243.
Rhodoprasina, 407.
Rhomborista, 89.
Rhopalopsyllus, 173, 174.
Rhynchocyclus, 23, 24, 361, 362.
Rhynclioprion, 170.
Rhynchopsylla, 171.
Rhynchopsyllus, 170, 171.
ridgwayi ( Dendrocolaptes), 306.
Risoba, 233-5.
Rivula, 211.
roberti (Conopophaga), 353, 373.
robinsoni (Xylophanes), 183.
romae (Dicacum), 3(K), 301.
romaaowi (Lanius), 403.
rosacea (Carpophaga), 289, 302.
roscipileum (Ptilinopus), 288.
rosenbergi (Nemosia), 317.
rostratus (Tyrannus), 27.
rothschildii (Guiraoa), 3.59.
rotundata (Prosthctopteryx), 123, 120
rubecula (Ochyria), 101.
ruber (Eudocimus), 49.
• — (Phoethornis), 375.
— (Tacliyphonus), 17.
— (Tantalus), 49.
— (Trochilus), 375.
rubiginea (Paracarea), 245, 246.
rubigiuosus (Chloronerpes), 38.
— (Picus), 38.
rubra (Aroana), 281.
— (Fringilla), 16.
rubra (Phanaspa), 216.
— (Plioenicothraupis), 2, 17
— (Pirauga). 16.
rubraria (C'ultripalpa), 270.
— (Zcthes), 276.
rubricosa (Careades), 245.
rubripicta (Megaloba), 110, 111.
rubrocapilla (Pipra), 362.
rubromarginata (Hylophila), 219
rufa (A.xioeteta), 278.
— (Berresa), 199.
— (Malacoptila), 381.
— (Monasa), 381.
— (Sinariola), 238.
— (Spectrobasis), 102.
— (Stictoptera), 235.
— (Tanagra), 17.
rufalia (C'ultripalpa), 275.
— (Zethes), 275.
rufalis (Protagrotis), 191.
rufaxilla (C'olumba), 48.
— (Leptotila), 48.
rufescens (Maceda), 232.
— (Tyrannus), 330.
rufibasalis (Pycnoloma), 128.
ruficapilla (Muscicapa), 318.
ruficauda (Galbula), 42.
— (Ortalis), 3.
ruficeps (Formicarius), 373.
— (Lanius), 401-3.
— (Pyrrhorachis), 89.
ruficoUis (Egretta), 50.
— (Hirundo), 13.
— (Hydrauassa), 50.
— (Limonites), 290, 303.
— (Stelgidopteryx), 13.
— (Thamnophilus), 300.
ruficosta (Chloroehroma), 85.
rufigastcr (Phaethornis), 375.
ruligularis (Selerurus), 3(>4, 305
rufilunata (Eueymatoge), 117, 120, 121.
rulimentum (Hydranassa), 2, 50.
rulina (Columba), 2, 47.
rulinueha (Ochthodromua), 5.3.
rufipemiis (C'ulumbigallina), 47.
— (Pitangus), 2, 24.
— (Sa\irophagus), 24.
— (Talpaootia), 47.
ruliplaga (V'ittappressa), 200.
rulistriga (Chloroehroma). 85.
rufivenata (Paraleidia). 146.
rufoflava (Pseudclydna). 242.
rufoliva (Trochistis), 153.
rufopunctata (Earias), 219.
rufostrigata (Catephia), 251.
rufulus (C'eratophyllus), 170, 175.
— (Ctenocephalus), 175.
( "85 )
nifulus (Troglodytes), 6.
rufus (Anthus), 307.
— (Bucco), 381.
— (Cervus), 175.
— (Lanius), 401, 402.
— {Pacliyrhamplius), 58.
— (Tachyphonus), 17.
ruptilinca (Paralcis), 151.
ruptistriga (Chaetolopha), 95.
ruticilla (Motacilla), 7.
— (Setophaga), 7.
rutila (Piaya), 43.
rufilans (Lanius), 401, 402.
— (Synallaxis), 365.
rutilus (Cuculus), 43.
— (Cypseloides), 60.
— (Thryothorus), 6.
— (Xenops), 29.
Rynchops, 55.
sacra (Demiegretta), 291, 303.
salmoni (Dacnis), 317, 351.
— (Nemosia), 317, 351.
salomonis (Deilephila), 181.
Saltator, 17, 18, 314-16, 351, 358.
salvadorii (Dicaeum), 300, 301.
— (Stigmatops), 301.
— (Tanygnathus), 757.
samoanum (Macroglossum), 407.
sancta (Halcyon), 295.
sanguinea (Careades), 243.
sapphirina (Hylocharis), 376.
sappliirinus (Trochilus), 370.
Sarcinodes. 76.
Sarcopsylla, 170, 173.
Sarrothripa, 224, 225.
satelles (Letchena), 64.
satrapa (Laphyctcs), 26.
— (Tyrannus), 26.
saturata (Heterocnemis), 344, 347, 352.
— (Pachysylvia), 12.
— (Sclateria), 347, 352.
saturataria (Hypochroraa), 77.
saturatus (Cuculus), 304.
— (Formicarius), 33.
Saucerottea, 2, 3, 34.
Sauris, 112.
Saurophagus, 24.
schach (Lanius), 404.
schistacea (Hypoenemis), 344, 352.
— (Sclateria), 344-7, 352.
schistaceus (Dysithamnus), 339.
schist ochlamys (Accipjter), 382.
schoanus (Pycnonotus), 389, 390, 392.
scintillans (Anisogamia), 81.
sclateri (Anaeretes), 320.
sclateri (Automolus), 335.
— (Cercomacra), 370.
— (Hapalocercus), 320, 321, 351.
— (Knipolegus), 318.
— (RhjTichocyclus), 361, 362.
— (Tanagra). 3, 14. 15.
Sclateria, 2. 33, 344-7, 352.
Sclerurus, 28, 364, 365.
Scolopax, 54, 290.
Scotothorus, 326, 363. 364.
scutigera (Syntaracta), 136, 137.
Scythrops, 295.
seebohmi (Lanius), 39.3.
Seiurus, 7.
sellifer (Habrissus). 409.
semicinerea (Pachysylvia), 355.
semicinereus (Hj-Iophilus), 355.
semidiscata (Chogada), 140, 141.
semifascia (Sjmtaracta), 136, 137.
semifasciatus (Diallactes), 30.
— (Thamnophilus), 30, 306.
semiflava (Anticlea), 93.
semifusca (ilyiopatis), 5G.
— (Phyllomyias), 56.
seminivea (Anisogamia), 82.
semipalmata (Scolopax), 54.
— (Symphemia), 54.
semipalmatus (Aegiahtis), .53.
— (Charadrius), 53.
semirufata (Coenocalpe), 97.
semiscripta (Chloroclystis), 115.
seniitorquatus (Caprimulgus), 38.
— (Lurocalis), 38.
senator (Lanius), 40<J-2, 404.
Seneratia, 272.
sequens (Gerygone), 297.
sericina (Oclithophora), 220.
Serpophaga. 323, 351.
serrana (L'pucerthia). 331.
serrata (Diptheroides), 204.
serratilinea (Ophiusa), 257.
serva (Cercomacra), 343, 344, 352.
— (Pyriglena), 343.
setaria (Leptasthenura), 332, 333, 351.
— (Synallaxis), 332, 351.
Setophaga, 7.
sexcinctus (Dasypus), 173.
shivula (Zethes), 273.
sibirica (Muscicapa), 388.
Sicalis, 309, 351.
Siculodes, 65, 66.
silens (Arremon). 314, 358.
simplex (Crasilogia), 99.
— (Heterocnemis), 346, 347.
— (Ischnopsyllus), 186.
— (Lipangus), 304.
Sinariola, 238.
51
( 786 )
Sintor, -108, 400.
Siphia, 388.
Siptorni.s. 333.
Sitta, 33, 308.
Sittasomus, .3.
smaragdina (Blenina), 223.
— (Euplexia), 19.5.
socialis (Na.sua), 175.
solitarius (Myiodynastes). 24, 25.
somaliensis (Pycnonotus), 389-91.
sordida (Xcmo.sia), 311.
— (Thlypopsis), 310, 311, 351.
soui (Crypturus), 49, 385.
— (Tinamus), 49, 385.
spadicea (.Attila), 330.
Sparvius, 382.
speciosa (Columba), 47, 383.
— (Formicivora), 334.
— (Lepidoenas), 383.
— (SynaUaxis). 334, 335.
Spectrobasis, 102.
Spermophila. 58.
sphenocercus (Lanius), 393, 399.
Sphingiforma. 241.
spicea (Bagada), 208.
spinicauda (Chaetura), 60.
spinitorqims (Lanius), 402.
Spinus, 56.
spiza (Chlorophanes), 10.
— (MotaciUa), 10.
Spizaetus, 303.
splendens (Catephia), 253.
— (Chaetolopha), 93, 95.
— (Fringilla), 18.
— (Volatinia), 18, 3.59.
spodionotus (Dysithamnus), 2, 31, 32.
Spodiomis, 308, 309, 351.
Sporophila. 3, 18, 58.
spumata (Gelasma), 88.
spurius (Pycnonotus), 390, 391.
squamata (Ortalida), 350.
— (Ortalis), 350, 352.
— (.Stigraatops), 301, 304.
squamulatus (Microcerculus), 355.
squatarola (Charadrius), 290.
stabilata (Perixcra), 91.
stabilis (Tephroclystia), 131.
Steatomis, 1, 38.
steindachneri (Picumnus), 349.
Steirophora, 113.
Stelgidopteryx, 13.
stellaris (Pygiptila), 367.
— (Thamnophilus), 367.
stellata (Hypocncmis), 348, 349, 352.
Sterna, 55, 289, 302.
Ste-sichora, 67.
Stictoptera, 235.
stictoptera (Myrmeciza), 342, 343, 352.
Stigmatops, .301, 304.
Stiltia, 291. 303.
stoliczkae (.Anthoscopus), 386, 387.
straminea (Gonophaga), 158.
stramineata (Epitherapis), 156, 157.
striaticeps (Geositta), 318.
— (Hapalocercus). 321, 351.
— (Knipolegus), 318, 319, 351.
— (Muscisaxicola), 318, 319, 351.
striaticollis (Elania), 58.
striatipectus (SynaUaxis), 2, 3, 29.
striatipictus (Saltator), 17, 315, 316.
strigifera (Stesichora), 67.
strigocrenulata (Paragonitis), 238, 239.
striolata (Leptasthenura), 333, 352.
— (SynaUaxis). 333, 352.
striolatus (Bucco), 381.
Strix, 45, 46, 171.
Styx (Ercheia). 248.
subacuta (Ophiusa), 258.
subaffinis (Tanygnathus). 294.
subalaris (Xenopipo). 324, 325, 351.
subauratalis (Siculodes), 06.
subcaesia (Ochyria), 101.
subcinerea (Tanagra), 1, 15.
subditaria (Asthena), 107.
subflava (Capotena), 240.
— (Serphophaga), 323.
subfla\'Tam (Ornithion), 23.
subidaria (Xanthorhoe), 106.
Sublegatus, 56, 323.
SLiblutea (Ophiusa), 259.
submarginata (Myrioblephara), 144.
subpartita (Caradrina), 201.
subplaga (Ophiusa), 260.
subplumbea (Sclateria), 345, 347, 352.
subplumbeus (Dysithamnus), 344, 345.
subpunctata (Hypaetra), 262.
subpurpurea (Cosmophila), 237.
subradiata (Petrodava), 156.
subrosea (Perixera), 91.
subrabescens (Eucymatoge), 121.
subrubra (Carcades), 245.
subrufescens (Momotus), 41.
subspeciosa (Spiallaxis), 334.
subulata (Orthnocichla), 298, 299.
subumbr.a (Ophiusa), 250.
Suiriri, 323.
sulphuratus (Lanius), 24.
— (Pitangus), 2, 24.
sulphurea (Muscicapa), 23.
— (Protambulyx), 179.
sulphuresccns (PlatjThynehus), 23.
— (Rhynchocyelus), 23, 361, 362.
sulphureus (Myiozetetes), 23.
supcrba (Callidrepana), 61.
(787)
superciliaris (Leucopternis), 3S2.
— (Saltator), 315.
— (Tanagra), 314, 315.
superciliosa (Alcedo), 41.
— (CCTyle), 2, 41.
superciliosus (Lanius), 402.
— (Phoethornis), 374.
— (Trochilus), 374.
surinamensis (Myrmotherula), 36S.
— (Sitta), 368.
Burinamus (Tachyphonus), 357, 358.
susurrans (Dendrocolaptes), 29.
— (Dendrornis), 3, 29, 30, 59.
swainsoni (Momotus), 40.
— (Myrmeciza), 33.
sylvestris (C'oluml)a). 47.
Sylvia, 6, 11, 304, 317.
Symphemia, 54.
Synallaxi.s, 2, 3, 28, 29, 332-5, 351, 352, 365.
Syneosmia, 128.
Synoicus, 302,
Syntaracta, 134-7.
Sypna, 247.
S3Tnium, 45.
Tachycineta, 12.
Tachyphone, 310.
Tachyphonus, 17, 310, 317, 351. 357, 358.
taeniatus (Microcerculus), 355.
Taeniopygia, 304.
talpacoti (ColumbigalUna), 47.
Talpacotia, 47.
taminata (Epiplema), 74.
Tanagra, 1, 3, 14-21, 310, 314, 315, 357.
Tantalus, 49.
Tanygnathus, 294, 756.
Tapera, 42.
Tarache, 208.
Targalla, 227.
Tathodelta, 212.
Tchitrea, 388, 389.
tectus (Bucco), 380.
Teninora, 182. '
tenebrosa (CheUdoptcra), 381.
tenebrosua (Cuculus), 381.
tepa (Pachycephala), 299.
Tephroclystia, 122, 123, 129-32.
tephrospila (Syntaracta), 13G.
Terekia, 290.
Terenura, 348, 352.
Terpsiphone, 388.
terrestris (Synallaxis), 2, 3, 29.
terricolor (llelanerpes), 3.
tessellata (Zethes), 274.
testacea (Piranga), 57.
Testudo, 753.
teatudo (Hectopsylla), 170, 171.
— (Pulex), 170, 171.
Tetrao, 385.
thalassias (Poecilasthena), 103.
Thalassodes, 90.
Thalurania, 59, 377.
Thamnomanes, 367, 368.
thamnophiloides (Attila), 330, 351.
— (Muscicapa), 330.
Thamnophilus, 30, 31, 338, 339, 352, 36S-8.
thaumantias (Polytmus), 36.
— (Trochilus), 36.
Thaumatias, 34, 376.
theresiae (Polytmus), 378.
— (Psilomycter), 378.
— (Trochilus). 378.
Thlypopsis, 310, 311, 351.
thoracica (Pachysylvia), 355, 356.
thoracicus (Hylophilus), 355.
Thoracolophotos, 261.
Threnetes, 374.
Thryothorus, 0, 355.
thyelia ( Xj-lophanes), 185.
Th3Tia, 199.
Tibiocillaria, 231, 232.
tigrina (Turtur), 289.
tigrinus (Lanius), 403.
timorensis (Chalcophaps), 289.
timoriensis (Lalage), 298, 304.
Tinamus, 385.
Tinnunculus, 292.
Tityra, 27, 327, 328, 351, 364.
tobagensis (Formicivora), 56.
— (Mimus), 3.
— (Troglodj-tes), 3, 6.
Todirostrum, 22.
Tolmera, 152, 153.
tolypeutis (ilalacopsylla), 173,
Topaza, 378.
tomopunctata (Disticta), 277.
Tornosinus, 286.
torquata (Corythopis), 373.
torquatus (Astur), 292.
torrida (Loxia), 19.
Tetanus, 54, 290, 303.
Trichoglossus, 293, 303, 755.
tricolor (Ardea), 50.
— (Erythrura), 301.
— (Hydranassa), 2, 50.
— (Myiarehus), 26, 323, 324, 351.
— (Pycnonotus), 389-91, 762.
— (Tyrannus), 323.
trifasciata (Hirsutipes), 263, 264.
Tringa, 53, 54, 290, 291.
trinitatis (Coereba), 8.
— (Cyanerpes), 2, 8-10.
— (Cyelarhis), 11.
( -88)
408.
323, 324, 351.
trinitatis (Euphonia), 13.
— (Pitangus), 2, 24.
— (Ramphocaenus), 32.
— (Spermophila), 58.
— (Sporophila), 58.
— (Thamnophilus), 31.
Tripteridia, 124, 131.
tripuncta (Jlarcipa), 271.
triseriata (Anisoganiia), 82.
tristis (Leptopogon), 321, 322, 351.
— (Thamnophilus), 338, 339, 352, 367
tristriata (Chaetolopha), 96.
Trochilus, 33-6, 374-9.
Trochistis, 153, 134.
Troglodytes, 3, 6.
Trogon, 2, 41, 42, 379, 380.
Troides, 759.
tuberculatus (Physopterus),
tubercuUfer (Mjnarchus), 26
— (TjTannus), 26, 323.
turbata (Xylophanes), 183.
tiirdina (Dendrocincla), 336, 337.
turdinus (Dendrocolaptes), 337.
Turdus, 2-6, 57, 353, 369.
turneri (Ochthophora), 220.
Turnix, 288.
Tiirtur, 289.
Tympanota, 110.
Typhosia, 406, 407.
t^Tannina (Cercomacra), 370.
tyrarmiilus (Muscicapa), 26, 27.
— • (Myiarchus), 26.
TvTannus, 23, 26, 27, 323, 330.
t3Tannus (Jluscivora), 27.
ula (Morphopsis), 759.
ulterior (MierobeUa), 64.
umbrosa (Contortivena), 283.
— (Ercheia), 248.
unicolor (Chloropipo), 324r-6, 351.
— (Cniiiolegus), 318.
— (Halospiza), 308, 309.
— (Knipolegus), 317, 318, 351.
— (Phrygihis), 309.
— (Ramphocelus), 16.
uniformis (Chloropipo), 325, 326.
— (Haplospiza), 308, 351.
— (Spodiornis), 308, 351.
unilineata (Pseudacidalia), 216.
uniplaga (Earias), 219.
unipuncta (Xenoclystia), 133.
unipunctata (Caradrina), 201.
— (Carea), 242.
— (Oruza), 215.
unirufus (Lanius), 330, 351.
unitaeniata (Ochyria), 101.
unoplaga (Earias), 219.
Upucerthia, 331. 351.
urbana (Xantliorhoe), 106.
Urochroraa, 44, 60.
uropygialis (.\ttila), 328, 329.
— (Dasycephala), 328. 330.
- (Stelgidopteryx), 13.
uropygiata (Attila), 329, 330.
— (Muscicapa), 330.
Urospizias, 292.
Urubitinga, 46.
ustimacula (C'oenocalpe), 97.
ustula (Phanaspa), 216.
vagilinea (Lobophysa), 158.
vahdus (Attila), 330, 351.
varia (Lanius), 403.
— (Otoraela). 403.
variabilis (Parra), 53.
varians (Blenina), 223.
variegata (Ampelis), 28.
— (Lanius), 403.
— (Oruza), 215.
rariegatus (Chasmorhynchus), 28.
— (Cueulus), 295.
— (Numenius). 290.
varius (Empidonomus), 362.
— (Lanius), 402.
vauxi (C'haetura), 37.
velata (Bursada), 139.
Tenalis (Dendrothripa), 226.
venezuelae (ilomotus), 3, 41.
venezuelensis (Geotrygon), 3, 48.
— (Jlerula), 57.
— (Molothrus), 20, 21.
— (Ramphocelus), 16.
— (Turdus), 57.
VeniUornis. 3, 39.
ventraUs (Muscicapa), 322.
— (Phylloscartes), 321, 322, 351.
venusta (Paralcis), 151.
vermivorus (Basileuterus), 7.
verreauxi (Leptotila), 47.
V'espertilio, 177.
vespertilionis (Ceratopsyllus), 176.
— (Pulex), 186.
Vespenigo, 186.
vestigiata (Chlorochroma), 86.
vialactea (Epiplema), 75.
vidua (Hypocnemis), 353, 370-2.
vieilloti (CaUiste), 14.
— (Calospiza), 2, 14.
vigors! (Pitta), 296.
vinacea (Columba), 383.
violacea (.\rdea), 51.
— (Columba), 384.
( 789)
violacea (Euphonia), 14, 356.
— (Fringilla), 14, 35fi.
— (Geotrygon), 384.
— (Nyotanassa), 51.
— (Steirophora), 113.
violaeeus (Trogon), 2, 42.
violicauda (Lampornis), 377.
Vireo, 11, 317, 351.
vireouinus (Empidochanes), 56.
virescens (Ardea), 51.
— (Butorides), 51.
virgata (Ciccaba), 45.
virgatum (Symium), 45.
virginaUs (Trogon), 41.
viridacea (Euplexia), 195.
viridata (Blenina), 223.
— (Eutelia), 230.
— (Hastina), 107.
— (Risoba), 233.
viridescens (Attila), 329.
— (Labanda), 220.
viridifimbria (Comostolodes), 87.
viridis (Creciscus), 385.
— (Gracula), 304.
— (Oriolus), 304.
— (Orthogonys), 310, 351.
— (Spectrobasis), 103.
— (Tanagra), 310.
— (Trogon), 42.
viridisecta (Prosthetopterjrx), 123, 127.
viridissima (Anisogamia), 82.
— (Agyrtria), 59.
viridiventris (Hylocharis), 376.
vitellinus (Ramphastos), 2, 42.
vitiense (Macroglossum), 407.
vitiensis (Cosmophila), 236.
Titis (Pholus), 181.
Vittappressa, 206.
vittata (Grison), 175.
vittatus (Lanius), 404.
Volatinia, 18, 359.
vulgaris (Ophiusa), 258.
vulpina (Trochistis), 154.
wallacei (Ptilinopus), 289.
wallacii (Heteropelma), 363.
— (Scotothorus), 363.
waterstradti (Prioniturus), 756.
welisi (Saucerottea), 34.
Westermannia, 218.
whitelyi (Agyrtria), 34.
wiedii (Petroglossii.s), 380.
wilsonia (Charadrius), 53.
— (Ochthodromus), 53.
wilsonius (Aegialitis), 53.
wolffsohni (Ischnopsyllus), 177.
woodwardi (Amytornis), 754.
— (Colluricincla), 755.
wuchereri (Arremon), 313, 314, 351.
xanthogaster (Ptilinopus), 2->8, 302.
Xanthoptera, 210.
Xanthopygia, 388.
xanthopygns (Pycnonotus), 390, 391.
Xanthorhoe, 103-6, 760.
Xanthornus, 19, 20.
xanthornus (Xanthornus), 20.
xanthoscelus (Turdus), 3, 57.
Xanthospilopt«ryx, 761.
xanthus (Protambulyx), 179.
Xenocerus, 410.
Xenoclystia, 132, 133.
Xenopipo, 1, 324, 325, 351.
Xenops, 29, 366.
Xenopsaris, 323.
Xestopsylla, 170.
xylinata (Coenocalpe), 98.
Xylophanes, 182-5.
yamdenae (Alcyone), 294.
yenisseensis (Anthoscopus), 387.
Yula, 193.
zagira, 213.
zalissa, 211.
zanthopygia, 388.
zebrata (Thalassodes), 90.
zeodita (Xanthospilopteryx), 761.
Zethes, 272-6.
zonaris (Chaetura), 36.
— (Hirundo), 36.
zonocercus (Buteo), 46.
zononota (Dechrozona), 348.
Zosterops, 300, 304.
52
NoVITATES ZoOLOGIC^.VoL.XIII.1906.
PI. I.
J.G.Keulemans del.et lith-
1. AMYTOHNIS WOODWARDI. <S .
2. COLLURICINCLA WOODWARDI. (^.
Wast^ewma-n imp.
^J,_ (J.C
NoviTATEs.ZooiiOGic^ Vol, XIII. 1906.
Pl.II.
JC.Keuiemans del et lith,.
1. RHIPIDURA NIGROCINNAMOMEA.9.
2. GOODFELLOWIA MIRANDA, d".
Vfe St, Newman amp
D
EXPLANATION OF PLATE IIL
Fig.
1.
2.
3.
4.
5.
6.
9.
10.
11.
12.
13.
14.
15.
16.
17.
Troii/es chimaera S .
Morphopsis meeki S .
Buzara caloclesma c?
Milionia pericallis i
Craspedosis desmtata t?
Ncoscaptia aequalis ?
yeoscaptia albata ?
Deudorix grandis S
)> i>
Caprimhna aenea ?
Asura rhodina S
Morphopsis ula $
)) J) ^
Argyrolepidia palaea ?
Hijpochnjsops meeki <?
Gaprimima metallica %
I). 759
J)
)>
I). 761
J). 760
p. 761
p. 759
I)
p. 761
p. 759
))
p. 761
p. 759
p. 761
NoVITATES ZoOLOGICt VoL. XIII. 1906.
PL. III.
ttirtlD Runmivil & Co., Honiunstatt«(«U. Stuttgart.
EXPLANATION OF PLATE IV.
Fig. 1 . Pu/Jilio burchellaniis c?
„ 2. „ coelus ? , type
„ 3. „ vertumnus autumnus ? .
4. ,, laodamas rhipidius ? , type
,, 0. ,, stehidacki (^ , type .
., «. „ „ ? .' . .
,, T. „ ipliulamas ealog>/na ? .
„ 8. „ anchises anchises c?
„ 9. „ p/wsp/iorus phosphorus S , tyite
., II'- ., „ „ ?
,,11. ,, „ gratianus <3 .
„ lii. „ anchises anchises 'i
p. 406
ji. 451
J). 470
p. 533
p. 453
p. 481
p. 485
]). 4fi8
p. 485
NoviTATics Z()()I.(h;ic.k. V'or,. XIII. i()o6.
Pi.. \V.
1. liURCHRl.I-.ANUS cf
2. COELUS ?
3. AUTUMNUS 5
4. RHIPIDIUS' S
These RcpraiitfLtiuns iifc four-Jijths Natiiml Size
5. STKINBACHI J
6. STEINBACHI J
7. CALOGYNA ?
8. ANCHFSES 3
y. PHOSPHORUS <^
10. PHOSPHORUS $
I I. GRATIANUS 3
12. ANCHISKS 5
■\ C lowlir. Imp.. MoorfloldB, E.C.
EXPLANATION OF PLATE V.
Fig. 13.
U.
].■>.
10.
17.
18.
19.
23.
24.
Papilio homolhoaa S , type .
aristeus coelebs S , t_vpe .
trapeza cJ, type
hariHodius imaus S, type
ariarathes leuetra S , type
tonjuatus leptalea <?, type
harm, xeniades ?-!'. amlrona,
klagesi ? . . .
aristor S . . .
scamander scamander S
euryleon eun/leon ?
erlaces xanthias ? , type
type
p. 501
p. 043
p. 669
p. 668
p. 674
ji. 020
!>. 668
p. 453
p. 508
p. 031
p. 665
p. 464
NoviTATEs ZooLOGiC/E. Vol.. .\ni. igo6.
Pi.. V.
13. HOMOTHOAS S
14. COIil.EBS <J
Ij. TRAPEZA 3
16. IMAUS 9
These Rt'pruUitefions arc four-fifths Nutitnil Size.
17. LEUCTRA <?
18. LEPTIS (?
19. J -f. ANDRONA
20. KLAGESI J
21. ARISTOR S
22. SCAMANDER (J
23. EUKYLEON J
24. XANTHIAS ?
EXPLANATION OF PLATE VL
type
pe
p. (;s)()
p. 457
p. 45(i
p. 72]
p. 72U
p. 474
p. 723
p. 474
\). 4r)t;
NoviTATES ZooLOGic^. VoL. XIII. igo6.
Pi.. \'I.
25. OHF-KTHLKHI ^
26. LOCRIS J
27. J -f. pyiiO.MI-LAS.
28. LINUS J
lowlcr. Imp,. ^lourllvldH, K.C.
These Reproductions irre fottr-Ji/lhs Nutiirul Size
2g. ORTHOSII.AIS J
30. HEl.IOS J
31. PARAI.ILS 9
32. EAHIS df
33. ERVTHRUS cf
34. ERVTHRLS 5
35. J -f. EUCHARIA.
EXPLANATION OF PLATE VIL
Fig. 36. Papilio polydamas vincentius S, type.
37. „ „ lucianus i, type .
38. ,, ,, xenodamas S
39. „ madyes chlorodamas S ■
40. ,, polydamas jamaicensis <S, type
41. „ „ dominiciis (5 , type
42. „ „ neodamas S .
43. „ madyes crispus S, type .
44. ,, anchises foetterlei 3, type
45. „ „ „ ? .
46. „ „ etias ?
47. „ „ „ S, type
p. .517
]). 518
n
p. 525
p. 522
p. 519
p. 525
p. 487
ij
p. 486
N'cn'ITATKS Zl)OI,(l(.IC K. Vol.. Xlll, igofi.
Pl. \'II.
EXPLANATION OF PLATE VIII.
ji. 012
NoviTATKs Z()oi.cKiic,+:. \'i)i.. XI 11. i<;o6.
Pi.. \'III.
EXPLANATION OF PLATE IX.
. 60. Papilio molops megalurus S , type
6L „ „ /letaerius S , tyTpe
62. ,, „ molops S, type .
63. „ glaucolaus mlaenus S , type
64. „ „ glaucolaus S
65. „ „ leiicas S, type
66. ,, protfsilaus archesilaus S , tyjie
67. „ „ protesilaus S
68. ,, Mgnodesmus S , type
p. .1'^
p. Til
p. 711)
p. 709
p. 708
p. 709
p. 717
p. 722
NoviTATKs Zoni,()(;ic,+:. Vol.. XIII. 1906.
1^1.. IX.
EXPLANATION OF PLATE X.
Fis
1.
2.
3.
4.
5.
6.
7.
8.
9.
10.
11.
12.
13.
14.
15.
Castnia icnius
Xanthorhoi' cerasiim S
Chlorochrotna latistriga S .
Xa7ithospilopteryx zeodita i
Moneta jAenicolor <S .
Arycanda fuhiradiata S .
Xanthorho'e interrufata i .
Anisogamia commaculata S
Andrhippnris caudaequina S
Dici/clodfs hieroghjpkica S
Hypochroma purpurissa c?
Cyphura par-data S ■
760
p. 761
p. 760
p. 761
p. 760
»
NoVlTATKS ZOOLOGIC.F.. \'or.. Xni. IC)06.
Pi,. X.
A, 0. I-owlor. Imp. MoorfloldH, K.C
NOVITATES ZOOLOGICAE.
H journal of ZooioQ^,
EDITED BY
The Hon. WALTEE ROTHSCHILD, Ph.D.,
Dk. ERNST HARTERT, and Dr. K. JORDAN.
Vol. XIII.
No. 1.
Pages 1—190.
IssDED, February 24™, at the Zoological Museum, Trino.
PRINTED BY HAZELL, VVAISON i; VlNEi", Ld., I.ONDOK AND AVLESBURY.
1906.
Vol. XIII.
NOVITATES ZOOLOGICAE.
EDITED BT
WAITER ROTHSCHUD, ERNST HARTERT, and KARL JORDAN.
CONTENTS OF NO. I.
pAoes
1. ON THE BIRDS OF THE ISLAND OF
TRINIDAD C. E. Bellmayr . 1—60
2. NEW DBEPANULIDAE, THYRIDIDAE,
URAKIIDAE, AND GEOMETRIDAE, FROM
BRITISH NEW GUINEA ....
3. TWO NEW. AGARISTIDAE ....
4. ON A NEW PARASITIC TINEID MOTH
FROM QUEENSLAND ....
5. NOTES ON THE SIPHONATTERA FROM
THE ARGENTINE DESCRIBED BY THE Karl Jordan and
LATE PROFESSOR DR. WEYENBERGH . N. C. Rothschild 170—177
6. NEW SPHINGIDAE Walter Rothsckild and
Karl Jordan . 178—185
7. NOTES ON BAT FLEAS iX. C. Rotfischild . 186—188
8. TWO NEW SATURNIIDAE .... n'alter Rothschild . 189—190
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R. TANCRE,
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receives every year from his collectors Large
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TRAL ASIA (Turkestan, Kuldscha.Northern
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?8Jl)L,190B
NOVITATES ZflOLOGlCAE,
H journal of Zooloo^.
EDITED BY
The Hon. WALTEE EOTHSCHILD, Ph.D.,
De. ERNST HAETERT, and Dk. K. JORDAN.
Vol. XIII.
Pages 191—410.
Issued, July 10th, at the Zoological Museum, Thing.
PRINTED BY HAZELL, WAT.SON & VINEV, Ld., LONDON AND AYLESBITRY.
1906.
Vol. XIII.
NOVITATES ZOOWaiGAE,
EDITED BY
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CONTENTS OF NO. II.
PA0E3
1. NEW KOCTCIDAE FROM BRITISH NEW
GUINEA G. T. JHhicae-Iiaker 191—287
2. ON THE BIRDS OF THE ISLAND OF
BABBER Eniat Ha/rterl . 288—302
3. ON THE BIRDS OF LUANG .... Eiiist Hartert . 302—304
4. CRITICAL NOTES ON THE TYPES OF
LITTLE-KNOWN SPECIES OF NEO-
TROPICAL BIRDS . . . Part I. C. E. Uellmayr . 305—352
5. NOTES ON A SECOND COLLECTION OF
BIRDS FROM THE DISTRICT OF PARA,
BRAZIL C. E. Uellmayr . 353 — 385
«. MISCELLANEA ORNITHOLOGICA— Part III. Ernst Hartert . 386—105
7. SOME SPHINGIDAE IN THE BRITISH
MUSEUM . . • Walter liolhschild and
Karl Jordan . 40C — 407
S. SOME NEW ANTHRIBIDAE FROM THE
COLLECTION OF H. E. ANDREWES . Karl Jordan . . 408—403
9. TWO NEW XENOGERUS (ANTHRIBIDAE)
IN THE COLLECTION OF R. VON
BENNIGSEN Karl Jordan . . 410
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AND
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set or in Papers, from all parts of the world, in
finest condition; 1,400 kinds of PREPAEED
LARVAE ; numerous LIVING PUPAE, etc. SEPA-
RATE Price Lists, Nos. XX. and XXII., for
COLEOPTEEA (21,000 Species).
Lists V. and VI., for HYMENOPTERA (2,800
Species). DIPTERA (1,850). HEMIPTERA (1,5.50),
NEUROPTERA (550), ORTHOPTERA (950).
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LARGEST STOCK OF
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Collectors In many parts of the World.
I^iti-yet*t EHtahliHttmcnt on fhe i'ttiiti iient.
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57, HAVERSTOCK HILL, LONDON, N.W.
Fresh consignments are constantly arriving-, in-
cluding MAMMALS, BIRDSKINS, BIRDS' EGGS,
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poKt free on apjtl'tcatinn.
RECENT ARRIVALS:
A fine collection !of pinned SYNTOMIDAE and
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including many extraordinary mimics.
JUST PUBLISHED. PRICE LIST No. 5: «
Reptiles, Batrachians, and Fishes, containing
over 300 Species, and including a large
number of rare and recently-described forms.
For particulars applu to—
W. F. H. ROSENBERG,
57, Haverstock Hill, London, N.W.
ROWLAND WARD, Ltd.,
"The Jungrle,"
166, PICCADILLY, LONDON, W.
MAMMALS, BIRDS,
ETHNOGRAPHICAL SPECIMENS
for Museums.
R. TANCRE,
ANKLAM, POMMERN, GERMANY,
receives every year from his collectors Larg&
Collections of LEPIDOPTERA from CEN-
TRAL ASIA (Turkestan, Kuldscha,Northern
Thibet) and EASTERN SIBERIA (Amoor),
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rBI.N'TED BV HAZELl,, WATSON AND VI>'EV, LD., LONDON AND AYLESBURY.
29 SEP. 1906
NOVITATES ZOOLOGICAE.
H Journal of Zooloo^.
EDITED BY
The Hon. WALTEE EOTHSCHILD, Ph.D.,
De. ERNST HAETERT. and Db. K. JORDAN.
Vol. XIII.
No. 3.
Pages 411—752. Plates IV.— IX.
Issued, August 30th, at the Zoological Museum, Tring.
PRINTED BY HAZELL, WATSON & VlNEiT, Ld., LONDON AND AYLESBURY.
1906.
Advertisements of Zoological Objects and Zoological Books only accepted.
Subscriptions for the present Volume are due NOW.
Dr. STAUDINGER & BANG-HAAS,
BLAS£:^VITZ-DRESDEN,
In their New Tbice List, No. XLVI., offer more
than 16,000 Species of well-named LEPIDOPTEEA,
set or in Papers, from all parts of the world, in
finest condition ; 1,400 kinds of PREPARED
LARVAE ; numerous LIVING PUPAE, etc. Sepa-
rate Price Lists, Nos. XX. and XXII., for
COLEOPTERA (21,000 Species).
Lists V. and VI., for HYMENOPTERA (2,800
Species), DIPTERA (1,850), HEMIPTERA (1,550),
KEDROPTERA (550), ORTHOPTERA (960).
All Lists have a-^onvenient index of genera.
Lihi^ritl Diacovnt fur Caah Orders. Prit'es line.
W. F. H. ROSENBERG,
Importer of EXOTIC ZOOLOGICAL COLLECTIONS,
57, HAVERSTOCK HILL, LONDON, N.W.
Fresh consignments are constantly arriving, in-
eluding UAUMALS, BIRDSKINS, BIBCS' EGGS,
EEPTILES, AMPHIBIANS, FISHES, INSECTS
OF ALL OBDEBS, SHELLS, etc., etc., from all
parts of the world.
Localities Guaranteed.
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post free on application.
WiLHELM SCHLUETER,
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tiARGEST STOCK OF
European and Exotic BIRD SKINS.
European and Exotic BIRDS' EGGS.
In stock : nearly 20,000 Bird Skins and 60.000 Birds' Eggs
LATEST PRICE-LISTS POST FRKE ON APPLICATIO.N.
A new Catalogue of Exotic Birdx, rontaining more
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Skins and Eggs sent on approval if deaired.
Collectors In many parts of the World.
Ljitrgeftt Establishnifnt on the fonltlient.
RECENT ARRIVALS:
A fine collection of pinned SYNTOMIDAE and
ARCTl/DAE from the Caura Valley, Venezuela,
including many extraordinary mimics.
JUST PUBLISHED. PRICE LIST No. S :
Reptiles, Batrachians, and Fishes, containing
over 300 Species, and including a large
number of rare and recently-described forms.
For particulars apply to—
W. F. H. ROSENBERG,
57, Haverstock Hill, London, N.W.
ROWLAND WARD, Ltd..
"The Jungrle,"
166, PICCADILLY, LONDON, W.
MAMMALS, BIRDS,
ETHNOGRAPHICAL SPECIMENS
for Museums.
R. TAN C RE,
ANKLAM, POMMERN, GERMANY,
receives every year from his collectors Large
Collections of LEPIDOPTEEA from CEN-
TRAL ASIA (Turkestan, Kuld.scha,Northern
Thibet) and EASTERN SIBERIA (Amoor),
and sells all his Duplicates at very Moderate
Prices. Price Lists post free to any address
on application.
R. TANCRE.
Annual Subscrijtlion to '' Xovitates Zoologicae" £1 Is.
Price of Yearly Volume, mhen completed, £1 10s. (Commission for Booksellers ort
covipleted volumes only.)
Communicationst etc.. may be addressed to
THE EDITORS OF " NOVITATES ZOOLOGICAE,"
ZOOLOGICAL MUSEUM,
TRINO.
rUlSTED BY HAZELL, WATSON AND VINEY, LD. , LONDON AND AVI.ESUUUY.
/t;^
NOVITATES ZOOLOGICAE,
H Journal of ZooIoq^^,
EDITKD BY
The Hon. WALTEE EOTHSCHILD, Ph.D.,
Dk. EENST HAETEET, and Db. K. JOEDAN.
Vol. XIII.
No. 4.
Pages 753—789. ^lates I., II., III., X.
Issued, December 22nd, at the Zoological Museum, Tring.
PRINTED BY HAZELL, ,WATSON Si VINEY, Ld., LONDON AND AYLESBURY.
1906.
Vol. XIII.
NOVITATES ZOOLOGICAK
EDITED BY
WAITER ROTHSCHILD, ERNST HARTERT, and KARL JORDAN.
CONTENTS OF NO. IV.
1. A NEW SPECIES OF GIANT TORTOISE . WulUr Rothschild
■2. ADDITIONAL NOTES ON BIRDS FROM N.W.
AUSTRALIA (Plate I.)
Ernst Hartert
3. NOTES ON BIRDS FROM THE PHILIPPINE
ISLANDS (Plate II.) Enist Hartert
4. NOTES TO PLATES III. AND X
5. ERKLARUNG ....
INDEX
Karl Jordan
Anton Reichenotv
PAGES
753—754
754—755
755—758
759—761
. 762
763—789
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A REVISION OF THE LEPIDOPTEROUS FAMILY
SPHINGIDAE.
BY THE
Hon. WALTER ROTHSCHILD, Ph.D.
and
KARL JORDAN, M.A.L., Ph.D.
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cxxxv and 972 pages, TOth 67 Plates.
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