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L161— H41
ORCHIDS OF PERU
Photograph by Louis O. Williams
CHARLES SCHWEINFURTH
FIELDIANA: BOTANY
A Continuation of the
BOTANICAL SERIES
of
FIELD MUSEUM OF NATURAL HISTORY
VOLUME 30
CHICAGO, U.S.A.
1958-1961
Library of Congress Catalog Card Number: 58-105^6
PRINTED IN THE UNITED STATES OF AMERICA
BY CHICAGO NATURAL HISTORY MUSEUM PRESS
ORCHIDS OF PERU
CHARLES SCHWEINFURTH
FIELDIANA: BOTANY
VOLUME 30, NUMBER 4
Published by
CHICAGO NATURAL HISTORY MUSEUM
MARCH 31, 1961
ORCHIDS OF PERU
ORCHIDS OF PERU
CHARLES SCHWEINFURTH
Research Fellow, Ames Orchid Herbarium
Botanical Museum of Harvard University
FIELDIANA: BOTANY
VOLUME 30, NUMBER 4
Published by
CHICAGO NATURAL HISTORY MUSEUM
MARCH 31, 1961
Library of Congress Catalog Card Number: 58-105^6
PRINTED IN THE UNITED STATES OF AMERICA
BY CHICAGO NATURAL HISTORY MUSEUM PRESS
FB
v/. 3
CONTENTS
Genera Included in Volume 30, Number 4
PAGE
Trichopilia 787 Hofmeisterella 923
Cochlioda 793 Ornithocephalus 924
Neodryas 797 Notylia 9
_ , , 7qq Cryptarrhena 941
Odontoglossum 7W _ . .
Q00 Macradema »44
Symphyglossum odd R
Tehpogon »4o
Brassia * Trichoceros 958
Miltonia 847 Stellilabium 961
Solenidium 850 Dichaea 962
Oncidium 851 Pachyphyllum 973
Sigmatostalix 916 Centropetalum 981
Lockhartia. . 921 Campylocentrum 984
LIST OF ILLUSTRATIONS
TEXT FIGURES PAGE
Trichopilia gracilis 790
Trichopilia juninensis 792
Odontoglossum brevilabium 809
Odontoglossum digitatum 813
Odontoglossum graminifolium 817
Odontoglossum pusillum 827
Odontoglossum tigroides 831
Brassia caudata 838
Brassia cauliformis 839
Brassia juninensis 843
Miltonia parva 849
Oncidium ampliatum 859
Oncidium luridum 887
Oncidium ovatilabium 895
Oncidium pusillum 900
Sigmatostalix bicallosa 918
Sigmatostalix crescentilabia 919
Ornithocephalus patentilobus 928
Notylia carnosiflora, N. mirabilis 933
Notylia conduplicans 935
Notylia parvilabia 937
Notylia rhombilabia 939
Cryptarrhena lunata 943
Macradenia tridentata 945
Telipogon rhombipetalus 953
Telipogon urceolatus 956
Telipogon Vargasii 957
Dichaea ancoraelabia 964
Dichaea campanulata 967
Dichaea panamensis 971
Dichaea tenuis 972
Pachyphyllum gracillimum 977
Campylocentrum micranthum 987
Campylocentrum minutum 989
Orchids of Peru
TRICHOPILIA Lindl.
A rather small genus of American epiphytes extending from
Mexico through Central America and the West Indies to Brazil
and Bolivia.
Rhizome abbreviated. Pseudobulbs commonly conspicuous and caespitose,
more or less ancipitous, 1-leaved. Leaves fleshy or coriaceous, linear and subterete
or more commonly rather broad and flat. Scapes lateral, basal, usually short,
few-sheathed below, loosely 1- to (rarely) 7-flowered above. Flowers showy and
often large, with spreading, often twisted, sepals and petals. Sepals subequal,
narrow, usually free, the lateral ones rarely somewhat connate. Petals very sim-
ilar to the dorsal sepal. Lip varying from sessile to deeply adnate to the column,
simple to more or less distinctly 3- or 4-lobed, concave and commonly surrounding
the column below; disc smooth, carinate or rarely callose. Column more or less
elongate, commonly dilated above, terminated by an ample, membranaceous, en-
tire to 3-lobed and fimbriate clinandrium. Anther opercular, incumbent, 1-celled.
Pollinia 2.
Al. Leaves fleshy, linear-ligulate, commonly subterete; pseudobulbs inconspicu-
ous to small, up to 2.5 cm. long; flowers relatively small, sepals 2.6 cm. or
less long T. subulata (T. hymenantha)
A2. Leaves chartaceous to coriaceous, always flat; pseudobulbs conspicuous,
4.5 cm. or more long I
la. Flowers small, sepals 2.5 cm. or less long; lip entire, obtusely angled in front,
bicarinate below T. peruviana
Ib. Flowers relatively large, sepals about 3 cm. or more long; lip commonly more
or less markedly 3-lobed, not obtusely angled in front 1
la. Leaves definitely petioled; sepals broadly lanceolate, about 15 mm. wide,
greenish mottled with chocolate; lip 3-carinate T. brevis
Ib. Leaves sessile, at most with a narrowed, conduplicate base; sepals linear-
lanceolate or linear, 7.5 mm. or less wide, not mottled 2
2a. Pseudobulbs linear-cylindric, about 10 mm. or less in diameter in the dried
specimen; leaf narrow, about 2 cm. wide 3
2b. Pseudobulbs oblong-cylindric to ellipsoid or ovoid, 25 mm. or more in diam-
eter; leaf relatively broad, 2.6 cm. or more wide; base of the lip without semi-
elliptic calli, merely unicarinate 4
3a. Flower large, sepals 4.3 cm. or more long; lateral sepals connate below; lip
distinctly surpassing the sepals, disc with a pair of semielliptic, fleshy calli
near the base T. juninensis
3b. Flower relatively small, sepals 3.3 cm. or less long; lateral sepals entirely free;
lip a little shorter than the lateral sepals, disc without a pair of semielliptic
calli below T. gracilis
787
788 FIELDIANA: BOTANY, VOLUME 30
4a. Median keel uniform, not dilated above T. fragrans
4b. Median keel abruptly dilated and cut off in front.
T, fragrans var. grata (T. grata}
Trichopilia brevis Rolfe, Lindenia 7: 91, t. 332. 1892.
Pseudobulbs aggregated, elongate-conic, 8-12 cm. long, unifoliate. Leaf peti-
oled; lamina lanceolate-elliptic or oblong-elliptic, acute, cuneate at the base,
10-16 cm. or more long, 4-5 cm. wide, coriaceous. Scapes lateral, basal, pendent,
loosely 2- to 3-flowered above. Flowers large and showy, with spreading segments.
Sepals and petals yellowish green blotched with chocolate. Sepals broadly lanceo-
late or elliptic-lanceolate, acute, with somewhat revolute sides, about 4 cm. long
and 1.5 cm. wide. Petals rather similar to the sepals, lightly oblique. Lip white,
subtrilobed above, very broad, nearly equaling the sepals in length, up to 4.8 cm.
wide; lateral lobes erect and surrounding the column, broadly rounded, undulate;
mid-lobe very broadly rounded, retuse, crisped-undulate, about 2.5 cm. wide; disc
with 3 yellow keels in the center of the lower half. Column short, subclavate, about
1.3 cm. long, biauriculate above, with a membranaceous, serrulate clinandrium.
Peru: Habitat and collector not recorded; introduced by L'Horti-
culture Internationale at Brussels.
Trichopilia fragrans (Lindl.) Reichb. f. Hamb. Gartenz. 14:
229. 1858; Reichb. f. Saund. Refug. Bot. 2: t. 127. 1882; Cogn. &
Goos. Diet. Icon. Orch. Trichopilia t. 3. 1898. Pilumna fragrans
Lindl. Bot. Reg. 30, Misc. 74. 1844.
Plant variable. Pseudobulbs often aggregated, strongly complanate, oblong-
cylindric to ellipsoid or ovoid, unifoliate, 4.5-12 cm. long, invested below by sev-
eral distichous, deeply imbricating, evanescent sheaths. Leaf oval to elliptic or
elliptic-oblong, rarely oblong-ligulate, acute, more or less narrowed to a condupli-
cate base, chartaceous, 11-30 cm. long, 2.7-7.6 cm. wide. Scape lateral, basal,
shorter than the leaf, erect to pendent; peduncle bearing 2-4 close, tubular sheaths
which are remote above; raceme very loose, bearing 1-4 (commonly 2 or 3) flowers.
Flowers rather large, fragrant, with spreading segments. Sepals green, white or
whitish green, with more or less undulate margins. Dorsal sepal linear-lanceolate
or linear-elliptic, acuminate, up to 4.5 (6) cm. long and 7.5 mm. wide. Lateral
sepals similar but slightly smaller, oblique, entirely free. Petals very similar to
the dorsal sepal but commonly a little broader. Lip rhombic-obovate in outline,
lightly 3-lobed above the middle, long-cuneate at the base, with the center strongly
adnate to the column, white with a yellow spot near the base, 2.5-4 cm. long,
1.6-3 cm. wide where broadest; lateral lobes shallowly semiobovate and broadly
rounded in front, erect, undulate; mid-lobe semiorbicular, retuse and lightly apicu-
late; disc with a more or less conspicuous, low, uniform, central keel on the basal
half. Column straight, more or less dilated above and round-auriculate on each
side, 1.5-2.1 cm. long, with a conspicuous, 3-lobulate, lacerate clinandrium.
Cuzco: Prov. of Calca, Lares Valley above Huallhuayoj, 2200-
2400 meters, on rocks, Weberbauer 7903—Junm: Chanchamayo
Valley, 1500 meters, Schunke 1294- Also Hispaniola, Colombia
• (type of Pilumna fragrans), and Venezuela.
SCHWEINFURTH: ORCHIDS OF PERU 789
Trichopilia fragrans (Lindl.) Reichb. f. var. grata (Reichb. f.)
C. Schweinf. Bot. Mus. Leafl. Harvard Univ. 12: 193. 1946. Tricho-
pilia grata Reichb. f. Card. Chron. (1868) 1338.
This concept differs from the type in having the keel on the lip dilated and
abruptly cut off in front. Like the type, it has fragrant flowers, the sepals and
petals green and the lip white with yellow blotches on either side of the basal keel.
Peru(?): Habitat and collector not recorded; introduced by
Messrs. Veitch (type of T. grata) .— Cuzco : Prov. of Quispicanchis,
Marcapata Valley, "evergreen hard-leafed bushwood consisting of
shrubs and trees," 1800 meters, Weberbauer 7840— Junin: Rio Man-
taro, Weberbauer 6582.
Trichopilia gracilis C. Schweinf. Bot. Mus. Leafl. Harvard
Univ. 15: 168, t. 56. 1952. Figure 161.
Plant epiphytic, medium-sized for the genus, about 29.5 cm. or less high.
Rhizome abbreviated. Pseudobulbs caespitose, numerous, linear-cylindric, appar-
ently compressed, unifoliate at the apex, 5-11 cm. high, up to 9 mm. in diameter,
striate-sulcate, concealed at the base by a few firm, evanescent sheaths. Leaf
linear-oblong to elliptic-oblong, subcoriaceous, acute, shortly narrowed below to
a sessile or shortly petioled, complicate or conduplicate base, 11.7-18 cm. long,
1.4-2.1 cm. wide. Inflorescence basal, suberect, very loosely 2- to 3-flowered
above, shorter than the leaf or rarely (including the terminal flower) subequaling
the leaf; peduncle slender, about 6.5-15 cm. long, with 3 close, tubular, separated
sheaths; floral bracts closely clasping and much shorter than the pedicellate ovary.
Pedicellate ovary up to 4.5 cm. long. Flowers rather small for the genus, cam-
panulate, white with orange on the throat of the lip. Sepals rather fleshy, longi-
tudinally concave. Dorsal sepal linear or lanceolate-linear, acuminate, about 3 cm.
long and 4 mm. wide when expanded. Lateral sepals free, obliquely linear or lan-
ceolate-linear, lightly decurved, long-acuminate with a conduplicate apex, about
equaling the dorsal sepal but narrower. Petals obliquely lanceolate-linear, more
or less lightly decurved, acuminate with a subconduplicate apex, slightly shorter
than the dorsal sepal. Lip a little shorter than the sepals, with the lower portion
convolute around the column in natural position, when expanded oblong-obovate,
broadly rounded in front, lightly contracted on each side above the middle (thus
subtrilobed), cuneate below, with the central line below adnate to the lower half
of the column, about 2.5-2.8 cm. long and 1.6-1.8 cm. wide near the apex; disc
with a short, more or less distinct, fleshy keel below the middle. Column short,
about 9 mm. high at the back, stout, slightly dilated upward, terminating in an
erect, somewhat bilobed, denticulate clinandrium.
Junin: Prov. of Tarma, Agua Dulce, 1800 meters, epiphyte in
forest, pseudobulbs, leaves and peduncle dark green, Woytkowski
35473 (type). Same locality, 1900 meters, epiphyte in shady forest,
mid-nerve of the leaf yellowish, lip with 7 orange spots in the center,
Woytkowski 35428.
TRICHOPILIA
qracilis
c
FIG. 161. Trichopilia gracilis C. Schweinf. 1, plant; X ^. 2, flower ex-
panded; X 1. 3, lip and column from side; X 1.
790
SCHWEINFURTH: ORCHIDS OF PERU 791
Trichopilia juninensis C. Schweinf. Am. Orch. Soc. Bull. 13:
424, t., 1945. Figure 162.
Plant slender for the genus. Rhizome apparently abbreviated. Pseudobulb
strongly ancipitous, linear-cylindric, unifoliate, more or less arcuate in the dried
specimen, about 7.5 cm. long, up to 10 mm. in diameter, clothed below with sev-
eral distichous, imbricating, evanescent sheaths. Leaf oblong, acute, shortly
narrowed to a petiole-like, conduplicate base, about 15 cm. long and 1.9 cm.
wide, chartaceous. Scape lateral, basal, short, arcuate or spreading, very loosely
2-flowered above; peduncle about 6 cm. long, provided with 3 separated, closely
clasping, tubular sheaths. Flowers large, white, with spreading, somewhat twisted
segments. Dorsal sepal linear, acute or short-acuminate, about 4.3 cm. long and
4.8 mm. wide in the middle. Lateral sepals narrowly linear, complicate-acute,
slightly longer than the dorsal sepal, carinate without, connate below for about
1.8 cm. Petals similar to the dorsal sepal but slightly wider, linear, lightly curved
near base, complicate-acute. Lip obovate in outline when expanded, distinctly
exceeding the sepals, retuse in front, adnate to the column below for about 8.5 mm.,
about 5.7 cm. in greatest length, lightly 4-lobed above the middle, about 3-3.5 cm.
wide across the lateral lobes; lateral lobes semiobovate from a long-narrowed base;
mid-lobe transversely oval or reniform, about 1.2 cm. long on each side of the retuse
apex and about 2.7 cm. wide; disc below with a low central keel which is surrounded
by a pair of narrow, semielliptic, fleshy calli. Column rather small, abruptly
dilated above, about 2 cm. long, including the deeply trilobulate, lacerate wing.
Junin: Chanchamayo Valley, 1500 meters, Schunke 1299 (type).
Same locality and altitude, Schunke 523.
Trichopilia peruviana Kranzl. Oesterr. Gartenz. 10: 11. 1915.
Rhizome short. Pseudobulbs ancipitous, linear-cylindric or linear-oblong, uni-
foliate, up to 10 cm. long and 2 cm. wide. Leaves lanceolate, acute or acuminate,
shortly complicate at the rounded base, up to 15 cm. long and 4 cm. wide, coria-
ceous. Inflorescences 2- or more-flowered above; peduncle clothed with small,
short, acute, scarious sheaths. Flowers white. Sepals lanceolate, acuminate,
straight or very slightly twisted, 2.5 cm. long, 4-5 mm. wide. Petals oblong-
lanceolate, acute, equally long but nearly twice as wide as the sepals. Lip simple,
cuneate-obovate, obtuse-angled in front, lightly emarginate in the middle [?of each
side], equally as long as the other segments, 1.5 cm. wide in front; disc very mi-
nutely velutinous, with 2 rather obscure yellowish calli extending from the base
to beyond the middle. Column about half as long as the lip, the margin of the
clinandrium 3-partite and minutely denticulate.
Peru: Habitat and collector not recorded; imported by P. Wolter
of Magdeburg (type) .—Junin, fide Schlechter, Fedde Repert. Beih.
9: 169. 1921.
No material of this concept was seen.
Trichopilia subulata (Sw.) Reichb. f. Flora 48: 278. 1865;
Cogn. Urban Symb. Antill. 6: 628. 1910. Epidendrum subulatum
TRICHOPIUIA
/uninensis C/. (5 c Pi we in
/I
FIG. 162. Trichopilia juninensis C. Schweinf. 1, plant; X 4A- 2, dorsal
sepal; XI. 3, petal; XI. 4, lateral sepals; XI. 5, lip expanded; XI. 6, col-
umn from side; X 1.
792
SCHWEINFURTH: ORCHIDS OF PERU 793
Sw. Prodr. Veg. Ind. Occ. 123. 1788. Cymbidium subulatum Sw. Fl.
Ind. Occ. 3: 1473. 1806. Trichopilia hymenantha Reichb. f. Bonpl.
2: 90. 1854; Reichb. f. Xen. Orch. 1: 15, t. 7, figs. I, 1-11. 1854;
Hook. f. Bot. Mag. 98: t. 5949. 1872. Leucohyle Warscewiczii Kl.
Ind. Sem. Hort. Berol. App. 1854.
Plant slender, variable. Rhizome abbreviated. Pseudobulbs aggregated,
small and inconspicuous, linear-cylindric, lightly compressed, 8-25 mm. long,
unifoliate. Leaves fleshy, linear or subterete, channelled, acuminate and often
mucronate, 10.8-22.9 cm. long, up to 10 mm. wide. Scapes lateral, basal, short,
lax to pendent, loosely several-flowered nearly to the base, up to 15 cm. long.
Flowers small, with spreading segments. Sepals and petals white to pale yellow,
lip white with red or purplish spots. Dorsal sepal lanceolate, linear-lanceolate or
elliptic-linear, acuminate, 1.6-2.6 cm. long, 3-4 mm. wide near the middle. Lateral
sepals entirely free, obliquely linear-lanceolate, acuminate, dorsally carinate, with
a more or less prominent subapical mucro, about as long and broad as the dorsal
sepal or narrower. Petals similar to the dorsal sepal, more or less oblique. Lip
suborbicular and obscurely 4-lobed or round-obovate, concave with the sides of
the lower part surrounding the column, abruptly acute or obtuse and apiculate,
the margins irregularly denticulate to lacerate, 1.5-2 cm. long, 1.4-1.8 cm. wide;
disc provided at the base with a small, subquadrate, depressed callus or with a pair
of small, converging calli. Column short, stout, clavate, lightly recurved above,
6-9 mm. long including the wing, terminating in a relatively large, cucullate, sub-
orbicular-obovate, denticulate wing with a small subulate process on each side
below.
Peru: Habitat not recorded, Warscewicz s.n. (type of Leucohyle
Warscewiczii). Also the West Indies (type of Epidendrum subu-
latum), Panama and Colombia (T. hymenantha).
COCHLIODA Lindl.
Small or medium-sized South American epiphytes apparently
restricted to Ecuador, Peru and Bolivia.
Rhizome abbreviated. Stems very short, thickened into compressed, 1- or 2-
leaved pseudobulbs. Leaves linear or oblong to lorate. Scapes 1 or 2, lateral, basal,
erect, arcuate or pendent, loosely racemose (sometimes paniculate) above. Floral
bracts large or more commonly small and inconspicuous. Flowers small or me-
dium-sized, scarlet to rose-red, with spreading segments. Sepals subequal, free.
Petals broader than the sepals, somewhat oblique. Lip with an erect claw which
is more or less adnate to the column and with a spreading lamina; lamina 3-lobed,
with the lateral lobes rounded or oblong and often reflexed; mid-lobe porrect, en-
tire or dilated above and retuse to bilobed in front. Column erect and more or less
incurved, slender, often auricled above, with 2 stigmatic orifices. Anther terminal,
opercular, incumbent. Pollinia 2.
Al. Mid-lobe of the lip oblong or cuneate, not markedly dilated in front. . C. rosea
A2. Mid-lobe of the lip markedly dilated above and retuse to bilobed in front. . . I
794 FIELDIANA: BOTANY, VOLUME 30
la. Floral bracts conspicuous, about equaling the pedicellate ovary.
C. densiflora
Ib. Floral bracts small and inconspicuous, much shorter than the pedicellate
ovary 1
la. Pseudobulbs 2-leaved; flowers rose-purple C. vulcanica
Ib. Pseudobulbs 1-leaved; flowers scarlet 2
2a. Lateral sepals elliptic-lanceolate; petals broad, oval C. Noezliana
2b. Lateral sepals oblanceolate-oblong; petals oblong-elliptic C. Beyrodtiana
Cochlioda Beyrodtiana Schltr. Orchis 13: 5. 1919.
Plant medium-sized. Pseudobulbs aggregated, ellipsoid to broadly ovoid,
compressed, unifoliate, 3-4 cm. high. Leaves narrowly lorate, subobtuse, 10-
15 cm. long, 1.7-2 cm. wide. Scape erect, up to 20 cm. high; peduncle slender,
with several (5-7) small, tubular, remote sheaths; raceme about 8 cm. long, very
loose, 7- to 10-flowered. Floral bracts small, ovate-lanceolate, acuminate, con-
cave, about one third as long as the pedicellate ovary. Flowers bright scarlet, with
widely spreading segments. Dorsal sepal lanceolate or oblanceolate, acute, about
1.5 cm. long. Lateral sepals about equally long but apparently more or less nar-
rower, oblique. Petals obliquely oblong-elliptic, acute, about as long as the sepals,
wider than the dorsal sepal. Lip about 1.5 cm. long, with a short cuneate claw,
3-lobed in the middle where about 1.2 cm. wide; lateral lobes obliquely oblong,
obtuse; mid-lobe reniform, shortly clawed below, bilobulate in front; disc with 4
parallel, minutely puberulent keels extending to the base of the midlobe. Column
slender for the genus, about 1.5 cm. or less high, with a pair of semiquadrate auricles.
Peru: Habitat and collector not recorded. Plant appeared in the
nursery of Herr Beyrodt in Marienfeld near Berlin.
Cochlioda densiflora Lindl. Fol. Orch. Cochlioda 1853; Schltr.
Orchis 13: 3. 1919. Mesospinidium densiflorum Reichb. f. Card.
Chron. (1872) 393.
Plant about 21 cm. high, stout. Pseudobulb complanate-oval or oblong-oval,
about 5 cm. high, unifoliate, surrounded at the base by 2 pairs of distichous sheaths
of which the longer upper ones are leaf-bearing. Leaf on the pseudobulb oblong,
obtuse, narrowed below to an indistinct, conduplicate petiole, about 14 cm. long
and 1.8 cm. wide, coriaceous, with the mid-nerve prominent beneath. Scape sub-
erect, lateral, basal in the axil of a leaf-bearing sheath, slightly surpassing the leaf;
peduncle about 12.7 cm. long, entirely concealed by several close, tubular sheaths;
raceme subdense, about 9-flowered. Floral bracts large and prominent, spreading,
ovate-lanceolate, cucullate, about as long as the pedicellate ovary. Flowers rather
large, apparently white to cream-color, with widely spreading segments. Dorsal
sepal ovate-elliptic or "roundish," acute, nearly 1.5 cm. long. Lateral sepals much
narrower, spatulate-oblong or oblanceolate, oblique, acute, "nearly parallel," ap-
parently a little longer than the dorsal sepal. Petals very similar to the dorsal
sepal, oblique. Lip adnate to the column up to the middle of the latter, deeply
3-lobed, ovate in outline, distinctly shorter than the lateral sepals; lateral lobes
"roundish" or semioval, spreading; mid-lobe "linear-oblong" or subquadrate,
abruptly dilated and bilobed in front; disc at the base with a pair of linear, fleshy
SCHWEINFURTH: ORCHIDS OF PERU 795
calli which terminate in a "bivalve gland." Column elongate, arcuate, biauricu-
late at the apex.
Peru: Habitat unrecorded, Mathews s.n.
Cochlioda Noezliana (Hort. ex Card. Chron.) Rolfe, Lindenia
6: 55, t. 266. 1891; Hook. f. Bot. Mag. 122: t. 7474. 1896; Cogn.
& Goos. Diet. Icon. Orch. Cochlioda t. 2. 1899; Schltr. Orchis 13: 4.
1919. Odontoglossum Noezlianum Hort. ex Card. Chron. ser. 3, 8: 570,
602. 1890.
Plant rather large and showy, with an abbreviated rhizome. Pseudobulbs
clustered, complanate-ovoid to broadly ellipsoid, unifoliate, 3-5 cm. high, sur-
rounded at the base by 1 or 2 pairs of distichous, leaf-bearing sheaths. Leaves
oblong to linear-oblong, obtuse to acuminate, cuneate, narrowed to a subpetiolate,
channelled base, 7.6-25 cm. long, 2-2.5 cm. wide. Scape erect to arcuate or pend-
ent, up to 40 cm. long; peduncle with numerous short, clasping, subremote, tubular
sheaths; flower cluster racemose or somewhat paniculate, lax, few- to many-flowered.
Floral bracts oblong-lanceolate or ovate, acute or acuminate, commonly equaling
one third or one fourth of the pedicellate ovary. Flowers bright scarlet with
yellow on the lip, with spreading segments, 2.5-4 cm. across. Dorsal sepal ob-
long-elliptic, abruptly acute, about 1.6-2 cm. long, 5-8 mm. wide. Lateral sepals
lanceolate-elliptic, acute and apiculate at the tip, slightly longer but markedly
narrower than the dorsal sepal, somewhat oblique. Petals ovate-elliptic or oval,
acute, slightly oblique, about as long as the dorsal sepal but somewhat broader,
with irregular crenulate margins. Lip adnate to the column nearly to the apex of
the latter, deeply 3-lobed above the middle, cuneate at the base, about 1.7-2 cm.
long and broad when expanded, rotund-ovate in outline; lateral lobes obliquely
quadrate-suborbicular, spreading; mid-lobe subequal, decurved, obcordate, from
a broad cuneate base dilated above with a lightly retuse apex and undulate upper
margins; disc below with a broad, transverse, 4-lobulate callus or with 4 short,
parallel, fleshy keels. Column long and slender, with a trilobed apex, 1.3-1.5 cm.
long at the back.
Peru: Habitat and collector not recorded; introduced by M.
Noezli (type). — Cuzco, Puno, fide Schlechter, Fedde Repert. Beih.
9: 168. 1921. Also Bolivia.
Cochlioda rosea (Lindl.) Benth. Journ. Linn. Soc. 18: 327. 1881;
Cogn. & Goos. Diet. Icon. Orch. Cochlioda t. 3. 1899; Schltr. Orchis
13: 7. 1919. Odontoglossum roseum Lindl. Benth. PL Hartw. 151. 1844;
Hook. f. Bot. Mag. 100: t. 6084. 1874. Mesospinidium roseum
Reichb. f. Gard. Chron. (1872) 393.
Plant small to medium-sized. Rhizome abbreviated. Pseudobulbs aggre-
gated, ovoid to broadly ellipsoid, strongly complanate, unifoliate or bifoliate,
3-5 cm. high, surrounded at the base by 2 pairs of distichous sheaths of which the
larger upper ones are leaf-bearing. Leaves variable, elliptic-oblong to linear, acute
to obtuse, cuneate-narrowed to a conduplicate, subpetiolate base, 5.8-20 cm. long,
796 FIELDIANA: BOTANY, VOLUME 30
1-2.5 cm. wide, with the mid-nerve conspicuous beneath. Scapes lateral, basal in
the axil of a sheath, erect to flexuous or arcuate, 13-40 cm. long; peduncle with
several more or less remote, short, tubular, evanescent sheaths. Flower cluster
racemose to sometimes paniculate, loosely 5- to 20-flowered. Floral bracts ovate
to lanceolate, acute, concave, half as long as the slender pedicellate ovary or less.
Flowers rose-red, with spreading segments, 2-3.5 cm. in diameter. Dorsal sepal
narrowly elliptic-lanceolate, acute, prominently apiculate, about 11 mm. long,
up to 4 mm. wide. Lateral sepals similar, obliquely elliptic-lanceolate, slightly
longer and markedly narrower than the dorsal sepal. Petals lanceolate-elliptic,
acute, about as long as the dorsal sepal but more or less broader. Lip about as
long as the petals, adnate to the lower third of the column, about 9.5-16 mm. long,
sharply 3-lobed below or near the middle, ovate in outline; lateral lobes short,
very obliquely ovate or rhombic from a cuneate base; mid-lobe much larger, por-
rect, linear-oblong, linear-lanceolate or cuneate, obtuse or truncate and not or very
slightly dilated (but not bilobed) at the apex; disc with 1 or 2 pairs of short fleshy
keels near the base. Column stout, with a 3-lobed apex, about 6.5 mm. high at
the back.
Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 168. 1921.
Ecuador (type of Odontoglossum roseum) .
Cochlioda vulcanica (Reichb. f.) Benth. & Hook. f. ex Veitch
Man. Orch. PI. 9: 189. 1893 j1 Cogn. & Goos. Diet. Icon. Orch. Coch-
lioda 1. 1. 1898; Cogn. Martius Fl. Bras. 3, pt. 6: 209, t. 44, fig. 2. 1905;
Schltr. Orchis 13: 6. 1919. Mesospinidium vulcanicum Reichb. f.
Card. Chron. (1872) 393; Hook. f. Bot. Mag. 98: t. 6001. 1872.
Plant medium-sized, with an abbreviated rhizome. Pseudobulbs aggregated,
narrowly ovoid, strongly compressed, 2.5-6 cm. long, bifoliate, surrounded in
youth by several pairs of distichous, leaf-bearing sheaths. Leaves elliptic-oblong
to narrowly oblong-ligulate, abruptly acute or subobtuse, attenuate below to a
subpetioled conduplicate base, 7.6-15 cm. long, 1.3-3.5 cm. wide, coriaceous, with
the mid-nerve carinate beneath. Scape lateral, basal in the axil of a sheath, sub-
erect to nodding, 20-40 cm. long; peduncle provided with several clasping, short,
subremote sheaths; raceme rather laxly several- to many-flowered. Floral bracts
narrowly triangular, concave, short-acuminate, much shorter than the pedicellate
ovary. Flowers rose-purple, with spreading segments. Dorsal sepal elliptic-lance-
olate, acute, about 1.6 cm. long and 5 mm. wide. Lateral sepals markedly longer
and narrower than the dorsal sepal, obliquely oblong-lanceolate or oblong-oblance-
olate, dorsally carinate. Petals similar to and as long as the dorsal sepal but
markedly broader, lightly oblique, acute. Lip adnate to the column up to the
middle of the latter, about 1.8-2.2 cm. long when expanded, rather deeply 3-lobed
near the middle, rhombic-ovate in outline; lateral lobes recurved, triangular-
semiobovate, spreading from a cuneate base; mid-lobe reflexed, much longer, with
an oblong claw, dilated and retuse in front; disc at the base of the mid-lobe with
2 pairs of short, oblong, fleshy keels of which the middle ones are longer. Column
rather slender, lightly clavate above, 1.2-1.5 cm. long, denticulate at the apex.
1 This orchid was not included by Schlechter among the Peruvian species, but
it was designated from that country by Cogniaux in Martius Fl. Bras, 3, pt. 6:
212. 1905.
SCHWEINFURTH: ORCHIDS OF PERU
Peru: Eastern part, source of the Maranon River, Spruce s.n.
Also Ecuador (type).
NEODRYAS Reichb. f.
A small genus of American epiphytes apparently confined to the
Andean regions of Bolivia and Peru.
A1 Lio definitely 3-lobed, with a 7-lobed callus at the base ....... N. densiflora
A2* L D slml without a 7-lobed callus at the base; disc of the lip with a more or
tess ZmS cauus which is dilated and lobulate above ..... . . . . . ...... I
la JSSTSlf- the lip broader than long, truncate to lightly retuse and
sometimes *»™^n irovt^ ^ ^^ ^ ^.^ N. reniformis)
Ib. Anterior part of the lip commonly longer than broad,
Neodryas densiflora Reichb. f. Gard. Chron. n. ser. 3: 492. 1875.
rect, auriculate wings above.
798 FIELDIANA: BOTANY, VOLUME 30
Peru: Habitat not recorded, Davis s.n.
Description drawn from a record of the type from the Reichen-
bach Herbarium. No example of the species was available.
Neodryas rhodoneura Reichb. f. Bot. Zeit. 10: 835. 1852; Xen
f ± n '.V69i^ 12~25' 1854' Neodryas MaMi Reich"'
Xen. Orch. 3 : 2L 1878. Neodryas latilabia Smith & Harris, Con-
ib Gray Herb Harvard Univ. 117: 39, figs. a-c. 1937. Neodryas
39 m '
Rhiz™n, V6getatively and in flor*l details, growing on trees or rocks
Rhizome more or less creeping. Pseudobulbs complanate, ovoid (rarely) to nar
rowly ellipsoid or pyriform-cylindric, unifoliate, 2.5-6 cm. long, invited beL by
2 or more pairs of distichous, imbricating sheaths of which the upper ones areleaf
°n a
obacut f ae oe .
>blong acute or subacute, more or less long-cuneate at the base, up to 28 cm long
net/ T' ™def(°ft,envmuch smaller); Petiole to about 7 cm. iong, d^TcS
nelled. Scapes lateral, basal, in the axil of an upper sheath, more or less exceeding
the leaves, 13-40 cm. or more long; peduncle slender, with 2 to 4 remote short
tubular, appressed and evanescent sheaths; inflorescence '
T'?1S C°mTnIy 5~15 Cm" IOng' F1°Wers small« eampanu ate puple
and white or rarely red-orange. Dorsal sepal strongly concave-cucullate
mTone sTm ^th«^»™ —what carinate on thTbtck about
mm. long, 3.8 mm. wide when expanded. Lateral sepals about equally long
connate into a deeply concave, oval lamina which is bicarinate without somewhat
broader than the dorsal sepal and bidentate at the apex, forming a pair rfSSSd
st'L t^thaeP1CUlar 10HbeS' /!falS suborb-ular-ovate, ap'icuLe, somewhat
than the sepals and markedly broader than the dorsal sepal. Lip slightly
shorter than the sepals, simple, abruptly deflexed above, somewhat sigmoid when
viewed from the side; basal portion incurved, claw-like, short, sometiTs moTe Or
Ivate ±deer0tha°nT " ^^ ""**" ^^ ^^ eXPanded ™<*™-
ovate, broader than long, more or less cordate at the base, truncate or lightly retuse
and sometimes apiculate in front, with the sides more or less undulate aL thus
apparently obulate; disc below (through the claw-like base) ^than oblong
SSSTSSjfr wl;:ch is abruptly dilated (at the ba- °f *» ^0"^
l?n, at tL h I I" CaI1US' C°IUmn V6ry Sh°rt and stout' about 3 mm. or less
long at the back with a pair of porrect, oblong-ovate, sometimes bidentate wings
above and just below, with a pair of short, triangular lobes which areToinTfn
)Q
1993.-
C: ^bancai' Quebrada de Quera, 3050 meters, Vargas
.co: Bosques de Choque-Chusco to Mollepata, 3400 me-
ters, Vargas 1163. Prov. of Paucartambo, slopes of Pillahuata, 3000
neters epiphyte in "ceja de la montana," Vargas 3663. Also Bo-
livia (types of Neodryas rhodoneura, N. Mandonii, N. latilabia and
zv. remformis).
SCHWEINFURTH: ORCHIDS OF PERU
799
Neodryas Weberbaueriana (Kranzl.) Schltr Orchis 12: 90.
1918. Cochlioda Weberbaueriana Kranzl. Engler Bot. Janrb. t>4,
Beibl. 117: 31. 1916.
Plant variable, epiphytic. Rhizome creeping, more or less abbreviated Pseu-
flo^duS£™£ or loosely paniculate with 2 or 3 J"^-J£j£jj
many-flowered, commonly loose below, more or less d^ '^JP^^j
Tampanulate, orange to dark orange-red, on rather long, J^jf^^
«Pnal strongly concave, lanceolate-elliptic to oval-ovate ( oblong ), acute to
M about 7 8^9 mm bng, 4-5.2 mm. wide when expanded. Lateral sepals con-
nate' f orm ng an empi or suborbicular lamina which is bicarinate without, nearly
edly shorter than the other segments in natural position, obovate-oblong and pan
front) just below the apical pair.
Huancavelica: Prov. of Tayacaja, Ampurco woods, on trail be-
tween Salcabamba and Surcubamba, 2900 meters, common in fog
zone forest, flowers waxy, Stork & Horton 10428. Montepungo,
5 km. east of Surcubamba, 3000 meters, flowers waxy, Stork & Hor-
ton 10370,-Huanuco: Yanano, about 1800 meters, on trees, Macbnde
3666 (a smaller and stouter form than typical);-Junm: Prov c
Huancayo, on the left side of the valley of Rio Mantaro above
Huachicna, 3100 meters, in sclerophyllous vegetation, Weberbauer
6549 (type of Cochlioda Weberbaueriana).
ODONTOGLOSSUM HBK.
A large genus of tropical American orchids, chiefly epiphytic,
extending from Mexico through Central America, locally in the West
800 FIELDIANA: BOTANY, VOLUME 30
Indies and south to Peru and Bolivia, the center of distribution
being Colombia.
Plants variable, small and inconspicuous to very large and showy with an
abbreviated or creeping rhizome. Pseudobulbs commonly conspicuous, 1- to 3-
leaved at the apex, surrounded at the base by one to several pairs of distichous,
imbricating, commonly leaf-bearing sheaths. Leaves usually narrow and elongate.
Scapes lateral, basal, stout, commonly equaling or surpassing the leaves, racemose,
simple-paniculate or compound-paniculate. Floral bracts minute to conspicuous
and subequaling the pedicellate ovary, concave. Flowers very small to large and
showy, with spreading segments. Sepals subequal, commonly lanceolate or oblong
to broadly obovate, free or the lateral sepals rarely connate below. Petals similar
to the sepals, commonly shorter and either narrower or broader. Lip simple or
3-lobed, with the basal portion (often narrow and claw-like) erect and commonly
parallel with the column and the anterior part more or less sharply reflexed, the
margins entire to lacerate or fimbriate; lateral lobes short, erect; mid-lobe entire
or emarginate, narrow or very broad; disc below commonly carinate, callose or
crested, sometimes with conspicuous tubercles or radiate keels. Column very
short to rather elongate, naked or with a pair of simple to lobed or lacerate wings
above. Anther terminal, opercular, incumbent, 1-celled or imperfectly 2-celled.
Pollinia 2, ovoid, affixed to a narrow stipe.
Al. Inflorescence racemose or rarely with a short branch at the base I
A2. Inflorescence paniculate, rarely with but a single short branch (in the speci-
mens seen) 21
la. Sepals and petals strongly crisped-undulate 1
Ib. Sepals and petals not strongly crisped-undulate, much longer than broad ... 3
la. Lip with a prominent, free, auricled claw at the base; sepals and petals sub-
acute to acute O. Harryanum1
Ib. Lip without a prominent, free claw at the base; sepals and petals obtuse to
lightly retuse 2
2a. Dorsal sepal oblong, much longer than broad; pseudobulb "linear."
O. brevifolium var. Weberbauerianum (0. Weberbauerianum)
2b. Dorsal sepal not oblong (obovate or oblong-obovate), a little longer than
broad O. brevifolium (0. chiriquense, O. coronarium, 0. brachypterum)
3a. Margins of the anterior portion of the lamina of the lip entire or nearly so ... 4
3b. Margins of the anterior portion of the lamina of the lip denticulate to lacerate-
dentate (rarely undulate-crenate; cf. 0. epidendroides HBK.) 14
4a. Lamina of the lip long-narrowed above (very acuminate) 5
4b. Lamina of the lip not long-narrowed above, obtuse or retuse to apiculate or
acute 7
5a. Disc of the lip with 2 sigmoid, upcurved horns 0. cirrhosum
5b. Disc of the lip with 4 crests or numerous filaments 6
6a. Lip with 4 dentate-lacerate crests near the base; wings of the column more
or less lacerate 0. praestans
6b. Lip with numerous slender, more or less branched filaments; wings of the
column subentire (subdenticulate above) O. Hennisii
1 In O. Harryanum the sepals and petals are merely more or less undulate, but,
in order to avoid possible misunderstanding, this concept is included in both
branches of the key.
SCHWEINFURTH: ORCHIDS OF PERU 801
7a. Lip distinctly 3-lobed (not pandurate), with the mid-lobe markedly narrower
than the basal portion 8
7b. Lip simple or pandurate, if 3-lobulate the apical portion (mid-lobe) nearly as
broad as or broader than the basal portion 10
8a. Flowers large, sepals about 2.5 cm. or more long; sepals lanceolate or oblong-
elliptic, more or less acuminate; disc of the lip with a digitately lobed callus.
O. digitatum
8b. Flowers small; sepals and petals oblong-cuneate to broadly obovate, rounded
to abruptly acute at the apex; disc of the lip with a pair of entire keels or
subentire calli 9
9a. Raceme few-flowered; flowers sulphur-colored; lateral sepals carinate below;
disc of the lip with a pair of elongate, fleshy calli O. anthoxanthum
9b. Raceme many- (10- or more-) flowered; flowers with 2 broad, brown bands
on a yellowish ground; lateral sepals not carinate below; disc of the lip with
a pair of fleshy, ellipsoid calli O. tigroides
lOa. Flowers relatively large, sepals about 3.3 cm. or more long 11
lOb. Flowers medium-sized to small, sepals about 2 cm. or less long; lamina of the
lip not pandurate 12
lla. Lamina of the lip pandurate O. Harnjanum
lib. Lamina of the lip oblong or elliptic-oblong, not pandurate ... 0. epidendroides
12a. Lower, erect portion of the lip broader than the anterior portion (thus no
evident claw); disc with a sulcate callus or a pair of simple elongate calli
which are free at the apex O. graminifolium
12b. Lower, erect portion of the lip much narrower than the anterior part (thus
markedly clawed); disc without a pair of simple calli 13
13a. Flowers small, dorsal sepal 7.5 mm. or less long; petals narrowed below.
O. pusillum
13b. Flowers medium-sized, dorsal sepal about 15 mm. or more long; petals much
broader below O. crocatum
14a. Wings on the column lacerate or pectinate 15
14b. Wings on the column entire or crenulate, at least not lacerate-dentate ... 17
15a. Flowers relatively large, sepals 4 cm. or more long (rarely smaller); basal
callus consisting of a large multipectinate crest O. Hallii
15b. Flowers smaller, sepals 3.5 cm. or less long; basal callus consisting of radiating
processes 16
16a. Lip rounded and apiculate in front; basal callus with the 4 middle keels much
larger and longer than the others and spiny-produced O. tripudians
16b. Lip acute or acuminate; basal callus consisting of similar keels. .O. cristatum
17a. Lamina of the lip ovate-lanceolate, lacerate-dentate O. lacerum
17b. Lamina of the lip oblong or subquadrate-ovate to pandurate, sometimes
broader near the apex, denticulate or crenate 18
18a. Margins of the lip undulate-crenate; lamina obtuse; disc with a 3-tuberculate
callus at the base O. epidendroide?
18b. Margins of the lip denticulate; lamina subacute to acute; disc without a
3-tuberculate callus 19
19a. Lamina of the lip "oval-oblong" or subquadrate-ovate, not contracted on
each side O. juninense
19b. Lamina of the lip pandurate or dilated at the apex (i.e., contracted on each
side) . . 20
802 FIELDIANA: BOTANY, VOLUME 30
20a. Sepals and petals acuminate; basal crest consisting of numerous filiform or
setaceous calli; column-wings rhombic O. subuligerum
20b. Sepals and petals acute; basal crest bi dentate in front with a tooth or apicule
on each side; column wings obsolete, crenulate O. Hrubyanum
2 la. Wings of the column very prominent and spreading, setose or elongate-
falcate; sepals and petals long-acuminate 22
21b. Wings of the column not long and prominent, often deflexed or wanting ... 24
22a. Lip abruptly 3-lobed in the middle; column with a trifid hood at the apex.
O. multistellare
22b. Lip simple or lightly 3-lobulate 23
23a. Sepals and petals white spotted with brown; lip with a pair of diverging
horns; column without angles below O. cirrhosum
23b. Sepals and petals yellow spotted with brown or purple; lip with four dentate
keels; column with a pair of angles below O. praestans
24a. Lip pandurate, or with the anterior portion broader than the basal part ... 25
24b. Lip not pandurate, the anterior portion not broader than the basal part ... 29
25a. Lamina of the lip broadly obovate; column with apical wings.
O. mystacinum (Cyrtochilum mystacinum, Odontoglossum rigidum)
25b. Lamina of the lip not broadly obovate; column without noticeable apical
wings 26
26a. Flowers relatively large, sepals 1.6 cm. or more long 27
26b. Flowers small, sepals about 8.8 mm. or less long 28
27a. Lip sharply acute or acuminate; basal crests dentate below, 2-toothed above.
O. ringens
27b. Lip very shortly acute; basal crests divided into 3 subequal teeth above.
O. fracliflexum1
28a. Sepals and petals spatulate or oblong-spatulate; lip very slightly constricted
in the middle and very little broader above the middle O. obscurum
28b. Sepals and petals oblong-oblanceolate; lip much constricted in the middle
and markedly broader above the middle O. pusillum
29a. Outline of the lip 3-lobed, often indistinctly so 30
29b. Outline of the lip simple 42
30a. Base of the lip with a linear, sulcate callus or 2 prominent keels 31
30b. Base of the lip without a linear, sulcate callus or 2 prominent keels 36
31a. Lip obscurely lobulate 32
31b. Lip distinctly 3-lobed or with a conspicuous linear-oblong or lanceolate an-
terior lobule 34
32a. Flowers small, sepals about 8-10 mm. long; disc of the lip with one sulcate
keel or 2 approximate keels ending in diverging lamellae O. longipes
32b. Flowers medium-sized to large, sepals 15 mm. or more long 33
33a. Leaves flat in the dried specimen, acute or acuminate. .O. aureo-purpureum
33b. Leaves strongly revolute in the dried specimen, obtuse O. revolutum
34a. Flowers large, sepals and petals about 15 mm. or more long; lip contracted
into a linear-oblong or lanceolate lobule above O. revoluium
34b. Flowers small, sepals and petals about 13 mm. or less long; lip not contracted
into a linear-oblong lobule above 35
1 When specimens of this concept are available, it may not prove to be spe-
cifically separable from O. ringens.
SCHWEINFURTH: ORCHIDS OF PERU 803
Lateral lobes of the lip semiorbicular; mid-lobe lanceolate-triangular.
O. myrianthum
Lateral lobes of the lip rhombic, antrorse; mid-lobe of the lip trapezoid.
0. melanthes
Flowers very small, lateral sepals about 5 mm. or less long 37
Flowers larger, lateral sepals 10 mm. or more long 38
Leaves oblong or elliptic-oblong, 6 cm. or less long; panicle short, few-
branched, about 5 cm. or less high; dorsal sepal suborbicular. .O. microthyrsus
Leaves linear or oblong-linear, 14.5 cm. or more long; panicle elongate,
many-branched, about 57 cm. long; dorsal sepal obovate-oblong.
0. depauperatum
Apex of the lip broad, retuse or subtruncate and apiculate 39
Apex of the lip narrow, oblong, linear-lanceolate to triangular or triangular-
cuspidate 40
Pseudobulb unifoliate; flowers relatively small, dorsal sepal 1.7 cm. or less
long, orange-scarlet; lip bicallose below O. retusum
Pseudobulb bifoliate; flower large, dorsal sepal about 2.5 cm. long, pale yel-
low lined with brown; lip with a single digitately divided callus at the base.
0. digitatum
Flowers small, sepals 12.8 mm. or less long; lip less than half as long as the
sepals 0. brevilabium
Flowers relatively large, sepals about 23 mm. or more long; lip nearly as long
as the sepals 41
Sepals and petals acuminate; lip 3-lobed in the middle; mid-lobe triangular-
cuspidate, long-acuminate O. multistellare
Sepals and petals acute; lip 3-lobed below the middle; mid-lobe linear-
lanceolate, obtuse to subacute O. trilobum
Apex of the lip broad, truncate or retuse and apiculate O. retusum
Apex of the lip narrow, at most minutely bilobulate at the tip 43
Flowers very small, sepals 10 mm. or less long; lip simply unicarinate or
bicarinate at the base 44
Flowers relatively large, sepals 2 cm. or more long 47
Petals sessile (not distinctly clawed), much broader than the sepals; keels of
the lip high O. megalophium
Petals clawed or distinctly narrowed below, narrower than the dorsal
sepal 45
Inflorescence with a few simple branches (subpaniculate) ; sepals 6-10 mm.
long 46
Inflorescence with numerous, compound branches (compound paniculate);
sepals minute, apparently 5 mm. long O. myanthum
Leaves over 2 cm. wide; lip bicarinate, with entire margins; column with a
pair of small, falcate-oblong wings above O. longifolium
Leaves 1.2 cm. or less wide; lip unicarinate, lightly retuse in the middle of
each side; column terminating in a pair of blunt, triangular teeth above.
O. obscurum
Inflorescence very slender, subpaniculate; lip lacerate; column with a pair
of subtruncate wings O. lacerum
Inflorescence broad, spreading-paniculate; lip not lacerate; column wing-
less . . 48
804 FIELDIANA: BOTANY, VOLUME 30
48a. Base of the lip with a pair of membranaceous, serrate to dentate keels and
with a long central keel which is flanked in front by a pair of sharp teeth or
horns 0. angnstatum (O. bellum, O. Loesenerianum, O. telraplasium)
48b. Base of the lip with a pair of fleshy, entire keels with a successive series of
31 and 2 fleshy calli 0. aureo-purpureum (0. compactum, O. Koehleri)
Odontoglossum angustatum Lindl. Bot. Reg. 23: sub t. 1992.
1837; Fol. Orch. Odontoglossum 17, no. 48. 1852. Odontoglossum
tetraplasium Reichb. f. Card. Chron. n. ser. 3: 558. 1875. Odonto-
glossum bellum Schltr. Fedde Repert. Beih. 9: 108. 1921; Mansf.
Fedde Repert. Beih. 57: t. 127, nr. 499. 1929. Odontoglossum Loese-
nerianum Schltr. Fedde Repert. Beih. 9: 110. 1921; Mansf. Fedde
Repert. Beih. 57: t. 128, nr. 502. 1929.
Plant stout, very variable. Pseudobulbs strongly complanate, cylindric to
ellipsoid or ovoid, commonly bifoliate, about 8 cm. or less long, surrounded by
1-5 pairs of distichous, imbricating, conduplicate sheaths of which the upper ones
are leaf-bearing. Leaves oblanceolate or oblanceolate-oblong, acute, sessile at the
gradually narrowed, conduplicate base, very variable in size, up to 42 cm. long
and 4.5 cm. wide (the outer ones on each side successively smaller). Scapes lat-
eral, basal, stout, commonly much surpassing the leaves, many-flowered, with a
narrow (short-branched) or widely spreading (long-branched) panicle above.
Flowers rather large with widely spreading segments, yellow (sometimes greenish
or brownish) commonly with brown spots. Sepals and petals more or less long-
acuminate with undulate margins. Dorsal sepal lanceolate, elliptic-lanceolate or
narrowly lanceolate, concave, up to 3.5 cm. long and 8 mm. wide below, often
with involute margins. Lateral sepals linear-lanceolate, more or less conduplicate,
carinate, subequally long but markedly narrower than the dorsal sepal. Petals
lanceolate or ovate-lanceolate, slightly oblique, markedly shorter than the dorsal
sepal but wider. Lip variable, ovate to ovate-oblong or oblong (sometimes slightly
narrowed on each side in the middle), short-clawed at the base, with the basal
portion subparallel to the column and the anterior portion spreading, acute (rarely)
to long-acuminate, about 2.7 cm. or less long and 1.1 cm. or less wide below (sub-
equaling the petals), provided near the base with a crest consisting of a pair of
relatively large, high, irregularly serrate-dentate or lacerate keels separated by a
pair of smaller, fleshy, narrow calli and in front of these by an elongate, central,
laterally flattened callus on either side of which are single (sometimes two) tuber-
cles or triangular horns. Column small, straight, dilated above, wingless, about
1.2 cm. or less high at the back.
Peru: Habitat not recorded, Davis s.n. (type of 0. tetraplasium).
— Ayacucho: Prov. of Huanta, Choimacota Valley, 2900-3000 me-
ters, in evergreen forest, Weberbauer 7547. — Cuzco: Prov. of Con-
vencion, from Quellomayo to Lucumayo, 2800 meters, Vargas 4498.
Prov. of Paucartambo, slopes of Pillahuata, 2900 meters, "ceja de
montana," epiphytic in rain-forest, Vargas 3664- "Pillahuata,"
Cerro de Cusilluyoc, 3000-3300 meters, epiphyte in thicket, Pen-
1 Often this series of three calli has a lateral tooth on each side.
SCHWEINFURTH: ORCHIDS OF PERU 805
nell 14137. Prov. of Urubamba, on trail from Puyupata to Sayac-
marca, 3600 meters, terrestrial, Vargas 2888. Prov. of Quispican-
chis, Marcapata, 2900 meters, in "bushwood consisting of high shrubs
and small trees, with many hard-leafed types," Weberbauer 7797
(this collection is rather abnormal in lacking apparent pseudobulbs
and in having somewhat dissimilar details of the basal crest).—
Huanuco: Huallaga, near Chaglla, 3200 meters, Weberbauer 6693.
— Junin: Wall of the valley of Rio Mantaro above Huachica, 3100
meters, Weberbauer 6550 (type of 0. Loesenerianum).—Loreto: Moy-
obamba, Filomeno s.n. (type of 0. bettum). Also Colombia and
Ecuador (type of 0. angustatum).
Odontoglossum anthoxanthum Reichb. f. Gard. Chron.
(1869) 388.
Vegetative portions lacking. Raceme fractiflex, few-flowered. Floral bracts
triangular, acute, half as long as the pedicellate ovary. Flowers small, sulphur-
colored. Dorsal sepal obovate-oblong or cuneate-oblong, abruptly acute. Lateral
sepals longer, narrower, dorsally carinate. Petals obovate ("cuneato-ovatis"),
apiculate, apparently similar to the dorsal sepal. Lip ovate when expanded, with
the lower part parallel to the column and the anterior part spreading with an in-
curved apex, sharply 3-lobed in the middle, subcordate at the base; lateral lobes
narrow, with an abbreviated, rounded free apex; mid-lobe triangular-ovate ("semi-
ligulato"), apiculate; disc with a pair of stout, parallel keels diverging in front,
extending from the base of the lip to the base of the mid-lobe. Column short,
stout, with a pair of ligulate, upcurved, erect and acute wings above.
Peru (probably) : Habitat and collector not recorded, introduced
by Messrs. Backhouse & Son of York, England.
In the Ames Herbarium is a floral analysis of this species from
the Reichenbach Herbarium.
Odontoglossum aureo-purpureum (as auropurpureum)
Reichb. f. Linnaea 22: 848. 1849; Lindl. Fol. Orch. Odontoglossum
15, nr. 44. 1852; C. Schweinf. Bot. Mus. Leafl. Harvard Univ. 14:
63. 1949. Odontoglossum compactum Reichb. f. Gard. Chron. n.s. 3:
492. 1875. Odontoglossum Koehleri Schltr. Fedde Repert. Beih. 9:
109. 1921; Mansf. Fedde Repert. Beih. 57: t. 128, nr. 501. 1929.
Plant very large and stout, about 2.5 meters or less tall, terrestrial or epiphytic.
Pseudobulbs oblong-ovoid or cylindric, complanate, about 6 cm. or more long,
bifoliate, surrounded at the base by one or more pairs of conduplicate, leaf-bearing
sheaths. Leaves oblong or linear-oblong to lorate, flat, narrowed above to an acute
or acuminate apex, more or less narrowed below to a conduplicate base, about
28-70 cm. long, up to 3.2 (rarely 4.2) cm. wide (some blades very much smaller),
rigid and coriaceous. Inflorescence lateral, basal, very large and stout, much sur-
passing the leaves, terminating in a many-flowered, much-branched panicle which
806 FIELDIANA: BOTANY, VOLUME 30
is simple or compound, loose or dense. Flowers rather large, with wide-spreading
segments, yellow commonly with purple, red or brown spots. Sepals and petals
more or less recurved and circinate above, with undulate margins. Dorsal sepal
lanceolate (rarely ovate-lanceolate), conduplicate at the narrowed (clawed) base
and the upper portion, long-acuminate, about 2-3.6 cm. long and 6-8 mm. wide
when expanded. Lateral sepals linear-lanceolate (or lanceolate), acuminate, dor-
sally carinate, connate at the very base, about as long as the dorsal sepal (or
slightly longer) but narrower. Petals obliquely lanceolate, acuminate, unguiculate-
narrowed with a flat and broad base, markedly shorter and broader than the dorsal
sepal. Lip lanceolate, sessile, with the lower portion erect and parallel to the col-
umn, the middle portion abruptly recurved and the acuminate upper portion more
sharply recurved, the lower margins being lobulate-crenate and undulate, nearly
equaling the petals when expanded, about 1.8-2.8 cm. long, up to 1 cm. wide below;
disc with a prominent basal crest which consists of a pair of high, fleshy keels or a
sulcate callus separated by a lower linear callus, in front of which is a series of 3 or
5 subequal, more or less lobulate calli and in front of these 2 larger, complanate-
ovoid or ellipsoid calli. Column relatively small, slightly dilated above and below,
about 1 cm. or less high at the back, wingless.
Peru: Habitat not recorded, Lobb, Pearce & Davis s.n. (type of
0. compactum). — Cuzco: Prov. of Urubamba, Puyupatamarca, 3600
meters, close to ancient ruins, Vargas 2879. Same locality, near
Wenner Gren ruins, 3400-3600 meters, in very wet places in moss
and on trees, Metcalf 30742. — Junin: Chanchamayo, Kohler s.n. (type
of 0. Koehleri). Also Colombia (type of 0. aureo-purpureum) and
Venezuela.
Odontoglossum brevifolium Lindl. Benth. PI. Hartw. 152.
1844; Fol. Orch. Odontoglossum 21, no. 61. 1852; Hook. f. Bot. Mag.
125: t. 7687. 1899. Odontoglossum chiriquense Reichb. f. Bot. Zeit.
10: 692. 1852; P. H. Allen, Ann. Mo. Bot. Gard. 36: 158. 1949.
Odontoglossum coronarium Lindl. Fol. Orch. Odontoglossum 21, no.
60. 1852; Batem. Monog. Odontoglossum, t. 27. 1874. Odontoglos-
sum brachypterum Reichb. f. Linnaea 41: 26. 1876. Odontoglossum
coronarium var. chiriquense Veitch, Man. Orch. PL 1: 23. 1886.
Odontoglossum coronarium var. miniatum Veitch, I.e. Odontoglossum
miniatum Hort. ex Veitch, I.e. Odontoglossum candelabrum Hort.
Linden ex Veitch, I.e.
Plant large, stout, very variable, with a robust, creeping rhizome (often not
present in the dried material) concealed by close, imbricating, scarious sheaths.
Pseudobulbs more or less distant, complanate, ovoid to cylindric-ovoid (rarely
suborbicular), unifoliate, about 4-11 cm. high, surrounded at the base by 2 or more
pairs of distichous, imbricating sheaths of which the upper pair is commonly leaf-
bearing. Leaves oval to elliptic-oblong (sometimes conduplicate in the dried
specimen), retuse to subacute at the apex, more or less cuneate-narrowed into a
conduplicate petiole, about 10-30 cm. long, up to 9 cm. wide. Inflorescences stout,
SCHWEINFURTH: ORCHIDS OF PERU 807
commonly erect but sometimes nodding above, several- to many-flowered, usually
more or less surpassing the leaves, up to 60 cm. high. Flowers large and conspic-
uous, brown edged with yellow or yellow blotched with reddish brown; column
white spotted with purple. Sepals with widely spreading segments, having crisped-
undulate margins. Dorsal sepal obovate or oblong-obovate ("orbicular and
clawed"), obtuse or generally retuse, about 1.6-3 cm. long, 1.6-2 cm. wide above.
Lateral sepals obovate-oblong, slightly oblique, somewhat longer and narrower
than the dorsal sepal. Petals obovate to elliptic-oblong, retuse or obtuse to sub-
acute, subequal to the sepals (sometimes broader or narrower), rarely slightly in-
dented on each side above (and thus subpandurate). Lip somewhat shorter than
the other segments, 3-lobed or pandurate in outline, about 1.7-2.5 cm. long, with
the basal part suberect and parallel to the column and usually conspicuously nar-
rower than the apical part; lateral lobes (at the base) small, erect, semiovate (often
subtriangular), rounded at the apex; mid-lobe relatively large, cuneate to obovate,
broadly rounded, retuse or prominently bilobed at the apex, 0.5-1.5 cm. wide;
disc with a small, fleshy keel, a smaller, transverse, retuse to subacute callus imme-
diately in front and a pair of fleshy calli at the inner base of the lateral lobes.
Column short, stout, about 8 mm. high at the back, with a conspicuous wing above
which is more or less trilobed and denticulate (especially toward the apex).
Southern Peru: Habitat not recorded, Pearce 48 (type of 0. bra-
chypterum). — Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 170.
1921. — Huanuco: Prov. of Huanuco, Carpish Pass, about 3000 me-
ters, Hodge 6283. Same locality, terrestrial in boggy clearing, Hodge
6276 (the flowers of this number are too old for examination).—
Junin: Chanchamayo Valley, 1800 meters, Schunke s.n. (Herb.
CNHM 571635; this collection is abnormal in having a broader
pseudobulb and a rather dissimilar lip). Also Costa Rica, Panama
(type of 0. chiriquense) , Colombia (type of 0. coronarium), Ecua-
dor (type of 0. brevifolium) , and Venezuela.
Odontoglossum brevifolium Lindl. var. Weberbauerianum
(Kranzl.) C. Schweinf. Bot. Mus. Leafl. Harvard Univ. 14: 64. 1949.
Oncidium Weberbauerianum Kranzl. Engler Bot. Jahrb. 37: 389. 1906.
Cyrtochilum Weberbauerianum Kranzl. Notizbl. Bot. Gart. Berlin 7:
95. 1917, nomen nudum; Engler, Pflanzenr. IV. 50, pt. 2 (Heft 80),
60, fig. 5H, a-c. 1922. Odontoglossum Weberbauerianum Schltr. Fedde
Repert. Beih. 27: 109. 1924.
This concept differs from the species in having narrower ("linear") pseudo-
bulbs, in having oblong (not oval or oblong-elliptic) leaves, and in having oblong
sepals which appear to be much longer than wide.
Amazonas: Molinopampa, east of Chachapoyas, 2000-2300 me-
ters, apparently epiphytic in open places in sclerophyllous vegeta-
tion, Weberbauer 4358 (type of Oncidium Weberbauerianum).
No record of this taxon has been seen.
808 FIELDIANA: BOTANY, VOLUME 30
Odontoglossum brevilabium C. Schweinf. Am. Orch. Soc.
Bull. 18: 578, t., 1949. Figure 163.
Plant large, epiphytic, with a stout rhizome. Pseudobulb cylindric, compla-
nate, slender, about 14 cm. high, bearing three leaves above, surrounded on either
side of the base by three spreading, imbricating, conduplicate sheaths which are
successively larger upward and mostly leaf-bearing. Leaves spreading, lanceolate-
oblong or ligulate, sharply acute, slightly narrowed below with a conduplicate base,
with the mid-nerve sulcate above and carinate beneath, those on the pseudobulb
larger, up to 42.6 cm. long and 2.5 cm. wide. Inflorescence basal in the axil of an
inner sheath, paniculate above, about 57 cm. high; peduncle lightly complanate,
about 32 cm. high, provided with three remote, short, infundibuliform sheaths;
panicle virgate, loosely many-branched with the branches loosely forking; bracts
broadly ovate, acute or acuminate, up to 12 mm. long at the base of the panicle.
Flowers small, numerous, yellow, very loosely arranged on a more or less fractiflex
rachis. Perianth segments widely spreading, fleshy. Sepals acuminate, longitudi-
nally concave. Dorsal sepal narrowly oblong-oblanceolate, about 10-12 mm. long
and 2.2-3.2 mm. wide. Lateral sepals narrowly and obliquely elliptic-lanceolate,
about as large as the dorsal sepal. Petals obliquely oblong-lanceolate, short-
acuminate, distinctly shorter and somewhat narrower than the sepals. Lip strongly
adnate to the column below; lamina abruptly recurved or reflexed, sharply 3-lobed
above the middle, about 4 mm. long and slightly broader when expanded; lateral
lobes relatively large, semiorbicular, lightly recurved; mid-lobe short, triangular,
fleshy, porrect or lightly reflexed; disc in the center with a large, indistinct, papil-
lose thickening which is slightly depressed across the middle. Column very short
and stout, terminating in a large, cucullate, irregularly dentate wing, 4.9-5.5 mm.
high at the back.
Junin: Prov. of Tarma, Agua Dulce, 2000 meters, on tree in low
highland forest, Woytkowski 35489.
Odontoglossum cirrhosum Lindl. Gen. & Sp. Orch. 211. 1833;
Fol. Orch. Odontoglossum 3, no. 5. 1852; Hook. f. Bot. Mag. 103:
t. 6317. 1877; Cogn. & Goos. Diet. Icon. Orch. Odontoglossum t. 9.
1897. Oncidium cirrhosum Beer, Prakt. Stud. Orch. 284. 1854.
Plant large, with an apparently abbreviated rhizome. Pseudobulb oblong-
ovoid, strongly compressed, unifoliate, 5-8 cm. high, surrounded below by about 3
pairs of distichous, imbricated, conduplicate, mostly leaf-bearing sheaths. Leaves
linear-oblong to elliptic-oblong, acute, narrowed to a conduplicate base, 10-30 cm.
long, about 2.8 cm. wide. Scape lateral, basal, much surpassing the leaves; pedun-
cle with several short, clasping, infundibuliform-cylindric sheaths; inflorescence
rather slender, simple or more commonly a many-branched panicle, many-flowered.
Flowers large, with widely spreading segments, milk-white with brown blotches on
the sepals, petals and mid-lobe of the lip, and with the basal part of the lip yellow
lined with red. Dorsal sepal narrowly lanceolate, long-acuminate, terminating in
a recurved, cirrhose point, about 4 cm. long and 7 mm. wide, with the margins
somewhat undulate. Lateral sepals very similar, slightly longer, somewhat oblique.
Petals rhombic-lanceolate, long-acuminate, markedly shorter than the sepals and
almost twice as broad. Lip about two thirds as long as the petals, about 3 cm.
ODONTOGLOSSUM
FIG. 163. Odontoglossum brerilabium C. Schweinf. 1, plant; X ]4- 2, portion
of the inflorescence; X Y*. 3, flower from side; X 1M- 4, dorsal sepal; X 2.
5, petal; X 2. 6, lip from above; X 2.
809
810 FIELDIANA: BOTANY, VOLUME 30
long when expanded, with the convolute basal portion erect and clasping the
column, 3-lobed near the base; lateral lobes broadly rounded, erect-spreading,
denticulate; mid-lobe recurved, elongate, narrowly triangular-lanceolate, long-
acuminate; disc with a pair of sigmoid-ascending horns between the lateral lobes.
Column straight, dilated upward, about 1 cm. long at the back, with a pair of tri-
angular-linear, ascending tendrils at the apex.
Ecuador (type). Peru(?), fide Schlechter, Fedde Repert. Beih. 9:
170. 1921.
Odontoglossum cristatum Lindl. Benth. PI. Hartw. 152. 1844;
Fol. Orch. Odontoglossum 18, no. 53. 1852; Rolfe, Bot. Mag. 145:
t. 8809. 1919. Oncidium cristatum Beer, Prakt. Stud. Orch. 283. 1854.
Plant rather large, epiphytic, apparently with an abbreviated rhizome. Pseu-
dobulb oblong-ellipsoid to ovoid-cylindric, rarely ovoid, compressed, 3-8 cm. long,
bifoliate, surrounded below by 2 pairs of distichous, imbricated, conduplicate, leaf-
bearing sheaths. Leaves lorate (rarely oblong-elliptic), more or less acute, slightly
narrowed toward the conduplicate base, gently arcuate, commonly 15-22 cm. long,
1.8-2 cm. wide. Scapes basal, axillary, arcuate, exceeding the leaves, 25-33 cm.
long, terminating in a laxly many-flowered raceme. Flowers rather large and
showy, with widely spreading segments, yellow spotted and blotched with red.
Sepals and petals very similar, elliptic-lanceolate, acuminate, 2.5-3 cm. long, about
9 mm. wide, the lateral sepals and petals slightly oblique. Lip with the claw-like
basal portion erect and parallel to the column, cuneate, about 8 mm. long; lamina
geniculate-reflexed, ovate or broadly pandurate-oblong, acute or acuminate, irreg-
ularly fimbriate-dentate and about 1.5 cm. long; disc at the base with a palmately
many-lobed crest of which the anterior divisions are broader. Column arcuate,
about 1.5 cm. high, membranaceous-margined below, with a pair of rounded,
irregularly fimbriate wings above.
Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 170. 1921.
Ecuador (type) and Colombia.
Odontoglossum crocatum Linden & Reichb. f.1 Gard. Chron.
(1867) 404.
Plant apparently large (only the pseudobulbs and the flowering raceme are
illustrated in the Reichenbach Herbarium and only the flower is described).
Pseudobulbs aggregated, strongly compressed, ovoid, ellipsoid or ovoid-oblong,
2-5.5 cm. long. Peduncles slender, with few, very small, tubular-infundibuliform,
remote sheaths. Raceme loosely several- (6- to 7-) flowered, with very small bracts
and elongate, arcuate, pedicellate ovaries. Flowers rather small, bright yellow,
with widely spreading segments. Dorsal sepal lanceolate or ovate-lanceolate
("oblong"), "acute" or acuminate, about 1.7 cm. long and 7 mm. wide. Lateral
sepals similar, semiconnate, slightly oblique. Petals very similar to the sepals
1 This species is very close to O. mystacinum (Lindl.) Lindl., and when actual
material is available, it may prove to be conspecific. For the present, O. crocatum
appears to be separable by having a simple, not paniculate inflorescence; by having
the lateral sepals semiconnate, not mostly free; and by having the wings of the
column linear and serrate on the lower margin, rather than broad and multifid.
SCHWEINFURTH: ORCHIDS OF PERU 811
and subequal, rounded ("subcordate") at the base. Lip much larger than the
other segments, nearly 2 cm. long and 1.5 cm. wide; claw short, narrow, sub-
parallel to the column; lamina broadly obovate with an abrupt, subquadrate basal
portion or "cordate-pandurate," prominently apiculate; disc at the base with
abrupt keels in front of which are 2 series of papillae ("apicules") on each side.
Column very small, about 6 mm. high, with a pair of wings which are linear and
serrulate below.
"Peru," fide Schlechter, Fedde Repert. Beih. 9: 170. 1921. In-
troduced by M. Linden.
In the Ames Herbarium is a drawing of this species from the
Reichenbach Herbarium.
Odontoglossum depauperatum Kranzl. Engler, Bot. Jahrb.
37: 391. 1906. Cyrtochilum depauperatum Kranzl. Notizbl. Bot.
Gart. Berl. 7: 98. 1917, nomen nudum, in key; Kranzl. Engler,
Pflanzenr. IV. 50 (Heft 80), 75, fig. 8D, a-c. 1922.
Plant large, slender, epiphytic, apparently with an abbreviated rhizome. Pseu-
dobulbs aggregated, ellipsoid-cylindric ("linear"), unifoliate, up to 4 cm. long,
surrounded below by 4 or 5 pairs of distichous, imbricating, conduplicate, mostly
leaf-bearing sheaths. Leaves linear or oblong-linear, acute or short-acuminate,
narrowed below to a conduplicate base (the lamina on the pseudobulb with an
elongate, narrow, basal portion), up to 20 cm. long and 1.5 cm. wide. Scape lateral,
axillary, very long, much surpassing the leaves, over 94 cm. in length; peduncle
with few, very short, tubular-infundibuliform sheaths; panicle elongate, about
57 cm. long, very loosely paniculate, with many short, distichous, simple or com-
pound branches which are several- to many-flowered, the floral axes being lightly
fractiflex. Flowers very small, with widely spreading or reflexed segments, pale
dull yellowish spotted with brown. Sepals obovate-oblong, short-clawed, shortly
acute, about 4.5 mm. long and 1.5 mm. wide, the lateral sepals more or less oblique,
carinate without. Petals about equally long with the sepals but markedly broader,
subspatulate, about 2.5 mm. wide. Lip sessile, 3-lobed, about 3.5 mm. long and
scarcely 2 mm. wide at the base; lateral lobes semiorbicular, membranaceous, re-
flexed; mid-lobe linear, excavated (cymbiform), obtuse; disc having a fleshy, some-
what wrinkled callus at the base with 2 globose warts in front. Column very short,
with rather broad, acute wings.
Amazonas: East of Chachapoyas, between Tambo Ragazan and
Almirante, 2700-2800 meters, in dense woods consisting of high trees
and low shrubs, Weberbauer 4443-
In the Ames Herbarium there is a photograph of the type.
Odontoglossum digitatum C. Schweinf. Am. Orch. Soc. Bull.
14: 208, t., 1945. Figure 164.
Plant epiphytic, medium-sized, apparently with an abbreviated rhizome. Pseu-
dobulb complanate-ovoid or pyriform, bifoliate, about 4.6 cm. high, surrounded
below by 3 pairs of distichous, imbricated, conduplicate sheaths of which the two
„ nt ILLl
812 FIELDI ANA: BOTANY, VOLUME 30
uppermost bear relatively small, oblong, sessile leaf-blades. Leaves (on the pseudo-
bulb) oblong or oblong-oblanceolate, acute, more or less long-narrowed to a slender,
conduplicate base, 16-17.8 cm. long, up to 2 cm. wide. Scape lateral, basal in the
axil of the uppermost sheath, arcuate above; peduncle stout, apparently compla-
nate, about 17 cm. high, with a single close, tubular sheath above; flower-cluster
very loosely 8-flowered, simply racemose except for a short, lateral, 2-flowered
branch at the base, lightly fractiflex. Flowers medium-sized for the genus, pale
yellow with brown lines. Perianth segments spreading. Dorsal sepal oblong-
elliptic or elliptic-lanceolate, concave, acute or subacuminate, about 2.6 cm. long
and 9 mm. wide. Lateral sepals mostly free (connate only near the base), elliptic-
oblong, lightly oblique, acute or subacuminate, slightly longer and narrower than
the dorsal sepal. Petals lanceolate-elliptic, acuminate, mucronate, lightly oblique,
a little shorter and sometimes narrower than the dorsal sepal. Lip somewhat
shorter than the sepals, with the sides of the anterior part incurved in natural
position, rhombic-obovate in outline from a slender, cuneate base, about 2.4 cm.
long and 1.8 cm. wide, sharply 3-lobed (with overlapping lobules) near the apex,
truncate-retuse and sharply mucronate at the tip; anterior portion (mid-lobe)
transversely oval-oblong, about 6.2 mm. long and nearly 1.3 cm. wide when ex-
panded; disc with a fleshy callus (extending from the base nearly to the middle),
4-lobed or 3-lobed near the middle with the mid-lobe deeply bilobed, each of the
4 lobules digitately 2-divided. Column short, stout, lightly recurved, terminating
in a large, 3-lobed clinandrium of which the middle lobule is broad and lacerate-
dentate and the lateral lobules triangular-lanceolate, about 11.8 mm. high at the
back including the deep clinandrium.
Cuzco: Prov. of Paucartambo, Pillahuata, 3200 meters, in rain-
forest, Vargas 3040.
Odontoglossum epidendroides HBK. Nov. Gen. et Sp. 1: 351,
t. 85. 1816; Reichb. f. Xen. Orch. 1: 59, t. 22, fig. II, 1, 2. 1854;
Cogn. Martius Fl. Bras. 3, pt. 6: 254. 1905. Oncidium epidendroides
Beer, Prakt. Stud. Orch. 285. 1854.
Plant large, epiphytic. Pseudobulb oblong-ovoid or ellipsoid, strongly com-
pressed, bifoliate, 5-6 cm. long, surrounded below by one or several pairs of dis-
tichous, commonly leaf-bearing sheaths. Leaves oblong-elliptic to narrowly
oblanceolate ("lanceolate"), more or less acute, long-narrowed below, 17.8-30 cm.
long, 2-3 cm. wide. Scape lateral, basal, in the axil of a sheath, stout, about
45 cm. or less high, erect below, rather flexuous above, terminated by a very
loose, several-flowered raceme. Pedicellate ovary elongate, arcuate, 3.5-5 cm.
long, much surpassing the bract. Flowers large, with widely spreading segments.
Sepals and petals yellow with 3 purplish spots, of which the middle one is very
large. Sepals similar, narrowly elliptic-lanceolate ("lanceolate"), short-acuminate,
lightly undulate, the lateral sepals slightly oblique, 3.5-3.7 cm. long, 6-7 mm.
wide. Petals similar to the sepals, acute, subequally long but slightly wider than
the sepals, suboblique. Lip somewhat shorter than the sepals, white; claw linear,
erect and parallel to the column, 9-11 mm. long; lamina abruptly reflexed, oblong
or elliptic-oblong, obtuse or subobtuse, crenate-undulate or serrulate, 13-16 mm.
long, 6-7 mm. wide; disc through the lower half with a linear-oblong callus produced
into 2 acute, flattened teeth separated by a much shorter tooth, and with 1-3 mi-
ODONTOGL^CXSSUM
FIG 164. Odontoglossum digitatum C, Schweinf. 1, plant; X %. 2, dorsal
sepal; X 1M- 3, petal; X 11A- 4, lateral sepal; X 11A> 5, lip and column from
side; X \1A- 6, lip, spread out; X 1M-
813
814 FIELDIANA: BOTANY, VOLUME 30
nute, dentiform calli on each side below. Column slender, clavate, incurved above,
with a pair of semiorbicular, sometimes bilobed, auricles above, 18-19 mm. long!
Prov. of Bracamoros: Between the Amazon River and the city of
Jaen, 480 meters, in rather warm locality, Humboldt & Bonpland s.n.
— Amazonas, Junin, fide Schlechter, Fedde Repert. Beih 9- 170
1921.
No example of this species was available.
Odontoglossum flavescens Rolfe, Orch. Rev. 12: 92. 1904.
This is an obscure species, since there is no description of the
vegetative parts and a very indefinite description of the floral seg-
ments. Accordingly, it is not entered in the key.
A photograph of the type in the Ames Herbarium furnishes, to-
gether with the original description, the following skeleton diagnosis:
Plant large. Pseudobulbs apparently fusiform, bifoliate, about 8 cm. long.
Leaf linear-lorate, acute, about 52 cm. long and 2.4 cm. wide. Inflorescence panic-
ulate, that in the specimen illustrated showing 2 divaricate branches about 5 cm.
apart, each one loosely 5-flowered. Flowers secund, with the slender pedicellate
ovary up to 2 cm. long. Floral segments small, elliptic-oblong, subconnivent,
about 1 cm. long, clear yellow. "Lip rounded below, with an oblong obtuse apex,
and a very large and prominent two-lobed crest, which is appressed to the column
at the base."
Peru and Ecuador: Habitat not cited, introduced by Messrs.
F. Sander & Company.
Odontoglossum fractiflexum Kranzl. Engler, Bot. Jahrb 37-
390. 1906.
Plant large, apparently with an abbreviated rhizome. Pseudobulbs narrowly
cylindric, subcompressed, unifoliate, up to 8 cm. high, entirely concealed in youth
by 5 or 6 pairs of distichous, imbricating, conduplicate sheaths of which the upper
ones are successively longer and bear larger leaf-blades. Leaves linear-oblanceolate,
acute or short-acuminate, long-narrowed below with a conduplieate sessile base,
up to 40 cm. long and 2 cm. wide (those on the sheaths shorter and less narrowed
below). Scape lateral, basal, in the axil of a sheath, somewhat longer than the
leaves, up to 50 cm. in length, bearing through the upper half a loose panicle which
is about 26 cm. long in natural position, the rachis (especially of the few-flowered
branches) very strongly fractiflex. Pedicellate ovary slender, elongate, much ex-
ceeding the minute bracts. Flowers yellow, with widely spreading segments, the
sepals and petals lightly undulate on the margins, with a recurved apex. Dorsal
sepal elliptic-lanceolate or oblong-lanceolate, acute or acuminate, concave, 1.6-
2 cm. long. Lateral sepals similar, appearing somewhat longer. Petals lanceolate-
elliptic, flat, similar to the sepals but slightly shorter and wider, apparently
dark-lined. Lip oblong-pandurate, with the narrower basal portion subparallel
to the column, shortly acute, about 1.8 cm. or less long and 8 mm. wide near the
SCHWEINFURTH: ORCHIDS OF PERU 815
front- disc below with a pair of rather high keels passing into 3 subequal teeth
toward the apex and with a short keel or large tooth between the keels. Column
angled slightly above the middle, wingless.
Puno: Between Sandia and Cuyocuyo, 2500-2800 meters, on cliffs
and on trees, Weberbauer 871 .
In the Ames Herbarium there is a photograph of the type.
Odontoglossum graminifolium C. Schweinf. Am. Orch. Soc.
Bull. 15: 30, t., 1946. Figure 165.
Plant slender, epiphytic, about 28 cm. high. Pseudobulbs rudimentary, uni-
foliate, entirely concealed by several pairs of distichous, conduphcate, leaf-bearing
sheaths of which the inner ones are much larger with larger blades. Leaves linear,
acuminate, gradually narrowed below to a more or less elongate, conduphcate base,
up to 25 5 cm. long and 9 mm. wide (leaf on the pseudobulb), the outer blades very
short Scapes lateral, in the axils of sheaths, gently arcuate, about 25 cm. or less
long- peduncle slender, with about 4 very small, tubular or infundibuliform sheaths;
raceme loosely 7-flowered, about 6 cm. long. Flowers very small for the genus,
yellow with spreading segments. Dorsal sepal oblong-oblanceolate, acute, mucro-
nate concave below, 7-7.6 mm. long, about 2 mm. wide. Lateral sepals linear-
oblanceolate, subparallel and connate near the base, acute, mucronate, longitudi-
nally concave, carinate, somewhat longer and slightly narrower than the dorsal
sepal Petals broadly and obliquely oblong, acute, distinctly shorter and wider
than the dorsal sepal. Lip with the lower portion (to above the middle) erect and
parallel to the column and the anterior part abruptly reflexed, ovate-lanceolate in
outline concave at the base, rounded or subtruncate and minutely apiculate at
the apex, about 8 mm. long and 4.5-5.6 mm. wide near the base when expanded;
disc with a large, median, sulcate callus extending to above the middle and termi-
nating in a pair of free, ovoid apices. Column very short and stout, clavate, about
3.5 mm. high at the back, with a pair of large porrect, semiovate wings which are
indented near the apex.
Cuzco: Prov. of Convention, "alturas de Pintobamba," 2700
meters, Vargas 3257.
Odontoglossum Hallii Lindl. Bot. Reg. 23: sub t. 1992. 1837;
Fol Orch. Odontoglossum 3, no. 6. 1852; Batem. Monog. Odonto-
glossum t. 21. 1874; Hook. f. Bot. Mag. 102: t. 6237. 1876.
Plant large, variable, epiphytic, with an abbreviated rhizome. Pseudobulbs
ovoid or oblong-ovoid, strongly compressed, unifoliate or bifoliate, 5-10 cm. long,
surrounded below by 2 or 3 pairs of distichous, leaf-bearing sheaths. Leaves ellip-
tic-oblong or oblong-oblanceolate, 13.7 to over 30 cm. long, 1.9-4.5 cm. wide,
acute, more or less narrowed to a conduplicate, often petiole-like base. Scape
lateral in the axil of a sheath, more or less surpassing the leaves; peduncle with a
few remote, short, tubular-infundibuliform sheaths; flower-cluster commonly ra-
cemose, rarely somewhat branched, loosely 4- to 20-flowered, 30-90 cm. tall, with
the rachis more or less fractiflex. Flowers large, with widely spreading segments.
Sepals and petals yellow with spots and blotches of purplish brown; lip white with a
816 FIELDIANA: BOTANY, VOLUME 30
few dark spots and a large yellow crest. Dorsal sepal lanceolate, elliptic-lanceolate
or "ovate-lanceolate," long-acuminate, concave, 3.6-5.5 cm. long, about 1.2 cm. or
less wide. Lateral sepals similar, often somewhat larger, oblique. Petals similar
to the lateral sepals, but shorter and often broader, obliquely ovate-lanceolate.
Lip shorter than the petals in natural position, with the short, narrow claw erect
and parallel to the column; lamina pandurate-oblong, the anterior portion lacerate-
dentate, abruptly mucronate and markedly broader than the crenulate basal por-
tion; disc at the base with a large crest consisting of several multifimbriate keels.
Column strongly arcuate, about 1.6-1.9 cm. long at the back, 2-winged above with
the wings divided into narrow teeth or tendrils.
Peru[?] : Valley of Lloa, about 2400 meters, Hall s.n. Also Colom-
bia and Ecuador (type).
Odontoglossum Harryanum Reichb. f. Gard. Chron. n. ser.,
26: 486. 1886; Warn. & Will. Orch. Alb. 8: t. 366. 1889; Cogn. &
Goos. Diet. Icon. Orch. Odontoglossum t. 11. 1897.
Plant large. Pseudobulbs approximate, oblong-ovoid to pyriform, 6-8 cm.
long, bifoliate, clothed at the base on each side by 2 or 3 sheaths of which the inner
ones are leaf-bearing. Leaves narrowly oblong to oblong-elliptic, acute to obtuse,
gradually narrowed below to a more or less distinct, channelled petiole, those
on the pseudobulb (which are the largest) up to 44 cm. long (including the peti-
ole), 2.5-4 cm. wide. Scape up to 1 m. high, racemose above, several- to many-
flowered. Flowers rather large, variable in size and color, pale brown to chestnut-
brown, petals and lip at the base with purple lines on a white ground and the large
apical part of the lip white, all parts more or less undulate. Dorsal sepal elliptic-
oblong to subelliptic, more or less acute, about 4.5 cm. long, 1.3- nearly 2.5 cm.
wide. Lateral sepals similar, lightly oblique. Petals elliptic-oblong to narrowly
elliptic, acute, lightly oblique, more or less smaller than the sepals. Lip about
equaling or slightly exceeding the sepals, from a stout, basally auriculate claw
abruptly dilated into an ovate-pandurate or lightly 3-lobed lamina, the sides in-
curved and the anterior portion subcordate to suborbicular and abruptly acute to
obtuse; disc with a prominent yellow crest which is lacerate or fimbriate in front.
Column relatively small, 1.2-1.7 cm. long, with a pair of small dentate wings above.
Junfn: Prov. of Tarma, Utcuyacu, 1900 meters, on a rotten tree
trunk in low forest, Woytkowski 35352. Also Colombia (type).
Odontoglossum Hennisii Rolfe, Gard. Chron. ser. 3, 10: 158.
1891.
Pseudobulbs densely aggregated, oblong-ovoid, subcompressed, about 4 cm.
long. Leaves oblanceolate-linear, acute, 10.2-15.2 cm. long, up to 2.1 cm. wide.
Scape about 20 cm. long, loosely 6-flowered. Pedicellate ovary about 2.5 cm. long,
five times as long as the subtending bract. Flowers rather large, with spreading
segments. Sepals similar, lanceolate, acuminate, about 2.6 cm. long, yellow with
a large brown area near the base and also about the middle. Petals somewhat
smaller, yellow with brown spots and lines. Lip free, white with red-brown spots
and blotches; claw about 4 mm. long; lamina spreading, 3-lobed, about 2.1 cm.
OD ONTTOGLCtS.SUM
or am i
FIG 165. Odontoglossum graminifolium C. Schweinf . 1, plant; X %. 2, flower
from side; X 4. 3, dorsal sepal; X 4. 4, petal; X 4. 5, lateral sepal; X 4. 6, lip
and column from side; X 4. 7, lip from above, spread out; X 5H-
817
818 FIELDIANA: BOTANY, VOLUME 30
long and 1.7 cm. broad; lateral lobes rounded, denticulate; mid-lobe triangular-
ovate, long-acuminate; disc with numerous slender, somewhat branched filaments
from 2 to 5 mm. long. Column clavate, about 1.5 cm. high, yellowish-white with
brown wings, denticulate at the triangular, acute apex.
Peru or Ecuador, Hennis s.n. (sent to Messrs. Charlesworth,
Shuttleworth & Co. of Heaton, Bradford and Clapham).
No record of this species was available.
Odontoglossum Hrubyanum Reichb. f. Gard. Chron. ser. 3,
4: 234. 1888.
Plant medium-sized. Pseudobulb pyriform, compressed, bifoliate, about 5 cm.
high, surrounded below by one or more pairs of leaf-bearing sheaths. Leaves ligu-
late, acute, about 22.7 cm. or less long. Inflorescence racemose above, laxly
5-flowered. Sepals and petals similar, cuneate-lanceolate, acute, apparently de-
flexed, dark sepia-brown or horse-chestnut color with a yellow apex. Lip clawed;
claw slender, adnate to the column at the base; lamina pandurate, subacute, dentic-
ulate on the deflexed sides, light ochre blotched with chestnut-brown near the base;
disc with a conspicuous basal, white, apically bidentate callus with a tooth ("api-
cule") on each side below and a swelling on each side at the very base. Column
arcuate, trigonous, with obscure, very narrow, crenulate wings.
Peru: Habitat not recorded, Hubsch s.n. Introduced by Messrs.
F. Sander & Sons.
The description was taken from the original diagnosis.
Odontoglossum j immense Schltr. Fedde Repert. Beih. 9: 109.
, 1921; Mansf. Fedde Repert. Beih. 57: t. 128, nr. 500. 1929.
Plant epiphytic, rather large, about 40 cm. high, with an abbreviated rhizome.
Pseudobulbs narrowly ovoid, strongly compressed, unifoliate or bifoliate, 4.5-
5.5 cm. long, surrounded on each side at the base by a leaf-bearing sheath. Leaves
linear-ligulate, acute, 22-30 cm. long, about 1-1.3 cm. wide in the middle, slightly
and gradually narrowed toward the base. Scape erect, arcuate above, about 40 cm.
long; peduncle slender, few-sheathed, 20-25 cm. long; raceme loosely several-
(about 8-) flowered. Flowers rather large, greenish yellow with large brown spots.
Parts of the perianth widely spreading. Sepals similar, narrowly lanceolate, acumi-
nate, narrowed near the base, with the lateral ones oblique, about 3.5 cm. long and
7.5 mm. wide. Petals obliquely oblong-lanceolate or elliptic-lanceolate, acuminate,
markedly shorter and a little broader than the sepals. Lip clawed, about 1.6 cm.
long and 9 mm. wide above the middle; claw short, linear, erect and parallel to the
column, nearly 5 mm. long; lamina reflexed, from an oblong base abruptly dilated
into a subquadrate-ovate ("oval-oblong") lamina which is acute or short-acumi-
nate, with a serrulate, lightly undulate margin; disc below with a linear, depressed
callus which is thickened and 3-dentate in front and extended into a pair of short
bidentate lamellae, and with a small bidentate callus on each side near the base.
Column lightly arcuate, about 1.4 cm. high, lightly auriculate-lobulate on the sides
of the stigma.
SCHWEINFURTH: ORCHIDS OF PERU 819
Junin: On the mountains of Janangu, east of Huacapistana, 2300-
2400 meters, in damp mossy woods, Weberbauer 2120.
No material of this species was seen.
Odontoglossum lacerum Lindl. Sert. Orch. sub t. 25. 1838;
Fol. Orch. Odontoglossum 19, no. 54. 1852.
Plant rather large. Pseudobulb ellipsoid, complanate, unif oliate about 3.2 cm.
high, surrounded below by about 3 pairs of distichous, leaf-bearing sheaths. Leaves
linear-oblong to elliptic-oblong, acute, narrowed below to a slender, channelled
petiole; lamina about 12 cm. or less long and 1.4-1.8 cm. wide; P^e about 3 cm.
Jong Scape lateral, basal in the axil of a sheath, much surpassing the leaves, terrm
nated by a slender, very loosely several- (about 10-) flowered raceme , whic * ha
a single short branch at the base of the flower-cluster. Flowers rather large, bright
"emon-yellow with one or two large brown blotches on the spreading segment^
Se7a*s and petals similar, lanceolate or elliptic-lanceolate, acuminate with incurved
tips, about 2.9 cm. long and 8.5 mm. wide, the lateral sepals and £*«*"?
oblique Lip markedly shorter than the other segments in natural position, claw
narrow, erecLnd parallel to the column; lamina reflexed-spreading ovate or ova^e-
lanceolate, concave, irregularly lacerate-dentate, cus P'date in f ront , disc at the
base with a crest consisting of fimbriate lamellae of which ^e \wo front one are
subulate. Column slender, more or less arcuate above, about 1.3 cm. long, wit!
pair of short, porrect, subtruncate auricles near the apex.
Huanuco: Cassapi (Casapi), Mathews 1867 (type).-Loreto, fide
Schlechter, Fedde Repert. Beih. 9: 170. 1921.
No example of this species was accessible, but a drawing from the
Lindley Herbarium was examined.
Odontoglossum longifolium Lindl. Benth. PI. Hartw. 152.
1844' Fol Orch. Odontoglossum 14, no. 39. 1852. Cyrtochilum longi-
folium Kranzl. Notizbl. Bot. Gart. Berl. 7: 98. 1917, nomen nndum,
in key; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 79, fig. 8J, a-c.
1922
Plant large, slender. Pseudobulb pyriform-cylindric, compressed, unif oliate,
about 6 cm. high, surrounded below by two or more pairs of distichous, imbricat-
ing, conduplicate, leaf-bearing sheaths. Leaves linear-lanceolate, acute or shor
acuminate, gradually long-narrowed below with a conduphcate, petiole-like base
up to about 35 cm. long and 2.3 cm. wide. Scape lateral, basal in the axil of a
sheath slender, much surpassing the leaves, about 56 cm. tall; peduncle arcuate,
with several very short, close sheaths, about 30 cm. long; flower-cluster an erect,
very loose, few-branched panicle, with the short, fractiflex branches erect-ascend-
ing Bracts minute, squamiform, much exceeded by the slender pedicellate ovaries
Dorsal sepal oblanceolate-spatulate, acute, with a slender base (thus clawed), about
9 mm long and 3 mm. wide. Lateral sepals similar but somewhat longer and
narrower with a longer claw, oblique. Petals similar to the dorsal sepal but some-
what shorter. Lip geniculate in the middle, with the lower half erect and paralle
820 FIELDIANA: BOTANY, VOLUME 30
to the column and the anterior half reflexed, oblong-lanceolate when expanded,
acute, bilamellate below with the tips of the lamellae free, diverging and ear-like.
Column small, abruptly clavate above, with a pair of very small, falcate-oblong
wings.
Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 171. 1921.
Ecuador (type).
In the Ames Herbarium there is a photograph of the type in the
Reichenbach Herbarium.
Odontoglossum longipes Reichb. f. & Warsc. Bonpl. 2: 100.
1854; Reichb. f. Walp. Ann. 6: 837. 1864. Cyrtochilum longipes
Kranzl. Notizbl. Bot. Gart. Berl. 7: 99. 1917, nomen nudum, in key;
Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 84, fig. 9B, a, b. 1922.
Plant rather large. Pseudobulbs broadly ellipsoid and unifoliate (according
to Kranzlin), surrounded on each side by several (5-7) conduplicate, imbricating,
leaf-bearing sheaths. Leaves not described. Scape (incomplete) undoubtedly
elongate, stout, paniculate above, with the uppermost internodes very short;
branches slender, abbreviated or up to 20 cm. long (Kranzlin), strongly fracti-
flex. Bracts minute, triangular-cucullate. Pedicellate ovary up to 2.7 cm. long,
nearly ten times as long as the subtending bract. Flowers small, yellowish, carti-
laginous. Sepals carinate on the back. Dorsal sepal cuneate-oblong or obovate,
concave, apiculate, 8-9 mm. long, 3-4 mm. wide. Lateral sepals a little longer
and narrower, elliptic-oblanceolate, acute, longer-clawed than the dorsal sepal.
Petals oblong-rhombic or rhombic-obovate, obtuse, slightly smaller than the dor-
sal sepal. Lip rhombic to ovate-lanceolate, subentire to very obscurely 3-lobed
near the middle with rounded lateral lobes, obtuse, about 8-9 mm. long, up to
5.2 mm. wide when expanded; disc through the lower half with one sulcate keel
or two approximate keels ending in thick, diverging lamellae with a small tooth
between them. Column dwarf, broadly margined on each side throughout-
Peru: Habitat not recorded, Warscewicz s.n. (type). — Piura: Ca-
mino Piura to Huancabamba, El Tambo, about 3000 meters, terres-
trial, Scolnik 1414>
Odontoglossum megalophium Lindl. Orch. Linden. 15, no. 83.
1846; Fol. Orch. Odontoglossum 14, no. 41. 1852. Oncidium mega-
lophium Beer, Prakt. Stud. Orch. 289. 1854. Cyrtochilum megalo-
phium Kranzl. Notizbl. Bot. Gart. Berl. 7: 98. 1917, nomen nudum,
in key; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 78. 1922.
Plant rather large, epiphytic. Pseudobulbs broadly ellipsoid, ribbed on the
sides, surrounded on each side by several distichous, imbricating, conduplicate,
leaf-bearing sheaths. Leaves long-petioled; lamina oblong-lanceolate, 35 cm. or
more long, about 3 cm. wide, acute or acuminate, long-narrowed below; petiole
about 16 cm. long, very slender, channelled. Scape lateral in the axil of a sheath,
elongate, much surpassing the leaves, terminated by an elongate panicle having the
branches remote (the lower branches again branched), with a fractiflex rachis.
SCHWEINFURTH: ORCHIDS OF PERU 821
Bracts very small, triangular, spreading. Pedicellate ovary slender, up to 1.2 cm.
long. Flowers numerous, very small, yellow. Dorsal sepal spatulate, obtuse or
rounded at the apex, often apiculate, concave, 4-5 mm. long, about 2.5 mm. wide.
Lateral sepals similar, longer and longer-clawed than the dorsal sepal. Petals obo-
vate-spatulate, rounded and apiculate at the apex, not clawed, a little shorter than
the dorsal sepal and markedly broader. Lip shorter than the lateral sepals, re-
curved in the middle, ovate or subcordate-ovate, obtuse or subacute, keeled beneath
near the apex, 4-5 mm. long; disc through the lower half with a pair of apically con-
verging, high keels which are lobed above. Column very short, clavate, wingless.
"Peru," fide Schlechter, Fedde Repert. Beih. 9: 171. 1921.
Colombia and Venezuela (type).
Odontoglossum melanthes Reichb. f. & Warsc. Bonpl. 2: 100.
1854. Cyrtochilum melcmthus Kranzl. Notizbl. Bot. Gart. Berl. 7:
98. 1917, nomen nudum, in key; Kranzl. Engler, Pflanzenr. IV. 50
(Heft 80), 80. 1922.
Pseudobulbs and leaves unknown. Scape stout, paniculate above with short,
few- (about 4-) flowered branches, the rachis flexuous. Bracts very small, ochreate,
acute, equaling about a ninth of the slender pedicellate ovary. Flowers small, dark
purple with a whitish yellow lip. Dorsal sepal obovate-spatulate ("elliptic and
clawed"), abruptly acute or apiculate. Lateral sepals subsimilar, longer-clawed.
Petals obliquely oblong-cuneate, acute. Lip [lanceolate or ovate-lanceolate],
3-lobed in the middle; lateral lobes antrorsely rhombic; mid-lobe from a rather
broad base, slightly dilated, then triangular (thus trapezoid); disc with 2 con-
tiguous keels slightly diverging at the apex. Column very short, broadly winged
on each side [throughout].
Peru: Habitat not recorded, Warscewicz s.n.
This diagnosis is a translation of the original description. The
only available material of this concept, which bears the designation
"Odontgl. melanthes Rchb. f. & Wzw." in Reichenbach's handwrit-
ing, shows a fragment of a panicle with compound branches about
8 cm. distant, and flowers inseparable from those of Odontoglossum
longipes Reichb. f. & Warsc. Suspicion points to the synonymy of
these species.
Odontoglossum microthyrsus Kranzl. Engler, Bot. Jahrb. 37:
390. 1906.
Plant relatively small, with an abbreviated rhizome, about 13 cm. high. Roots
very stout for the plant. Pseudobulbs approximate, oblong-cylindric ("oblong-
linear"), slender or stout, compressed, unifoliate, up to 2 cm. long, surrounded
below by 3 pairs of distichous, imbricating, conduplicate sheaths which are com-
monly leaf-bearing. Leaves all similar, oblong or elliptic-oblong, acute, narrowed
to a sessile (on the pseudobulb shortly petioled), conduplicate base, 5-6 cm. long,
l-1.2"cm. wide. Scapes lateral, basal, in the axil of sheaths, commonly about twice
as long as the leaves, about 12 cm. or less long; peduncle slender, about 7.5 cm. or
822 FIELDIANA: BOTANY, VOLUME 30
less long, with one or more short, close sheaths; flower-cluster commonly loosely
paniculate (when well-developed) with a single short branch at the base, loosely
several- (up to 10-) flowered. Flowers small, with undulate segments, brown-
violet with the lateral lobes of the lip and column white. Dorsal sepal suborbicular,
clawed below, concave, about 5 mm. long and broad. Lateral sepals oblanceolate-
oblong, subacute, a little longer and much narrower than the dorsal sepal. Petals
suborbicular, clawed below, very similar to the dorsal sepal. Lip 3-lobed, about
4.5 mm. long and 3.5 mm. wide near the base; lateral lobes rounded, membrana-
ceous; mid-lobe broadly linear, lightly bilobed at the apex; disc at the base with
3 calli of which the middle one is oblong and is surrounded on each side by a shin-
ing, excavated tubercle. Column very short.
Junin: Prov. of Tarma, mountains east of Palca, 2700-3000 me-
ters, on steppe with scattered shrubs and numerous grasses, Weber-
bauer 2447.
Description amplified by a photograph of the type in the Ames
Herbarium.
Odontoglossum mul tistellare Reichb. f . Linnaea 41 : 25. 1876.
Plant large. Vegetative parts not recorded. Scape tall, stout, terminated by
a rather narrow panicle about 44 cm. long, with subremote, ascending, few- (6- or
less) flowered branches. Flowers rather large, numerous, with spreading segments,
straw-color adorned with brown lines. Pedicellate ovary slender, 2-2.7 cm. long,
many times exceeding the small, lanceolate, cucullate bracts. Sepals and petals
very similar, lanceolate, acuminate or long-acuminate, about 2.5-3 cm. long and
6-8 mm. wide. Lip slightly shorter than the other segments; claw broad, thick-
ened, subparallel to the column; lamina 3-lobed, hastate-triangular or very broadly
ovate in outline, abruptly narrowed to form a long-cuspidate or triangular-lobulate,
crenulate-serrulate termination; disc at the base with 4 keels of which the outer
are 2-divided (into filiform tubercles) and the inner keels 3- to 4-divided (into sim-
ilar tubercles). Column less than half as long as the sepals, terminated by a trifid,
cucullate apex, the lateral lobes falcate and the middle one broad and serrulate at
the apex, with an arm on each side of the stigmatic cavity.
Peru: Habitat not recorded, Pearce s.n.
There is a drawing of this species from the Reichenbach Her-
barium in the Ames Herbarium.
Odontoglossum myanthum Lindl. Benth. PL Hartw. 152.
1844; Fol. Orch. Odontoglossum 13, no. 38. 1852. Cyrtochilum myan-
thum Kranzl. Notizbl. Bot. Gart. Berl. 7: 98. 1917, nomen nudum,
in key; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 77, fig. 8F, a-e.
1922.
Plant large, rather slender. Pseudobulb obscure, elongate-cylindric, unifoliate,
up to 10 cm. long according to Kranzlin, entirely concealed by 2 or 3 pairs of dis-
tichous, imbricating, conduplicate, leaf-bearing sheaths. Leaves oblong-lanceolate
SCHWEINFURTH: ORCHIDS OF PERU 823
or narrowly elliptic-lanceolate, acuminate, more or less narrowed below to a con-
duplicate base (that on the pseudobulb very long-narrowed below, about 45 cm.
long and 2.7 cm. wide), the blades on the sheaths successively smaller outward.
Inflorescences lateral, in the axil of a sheath, nearly twice as long as the leaves,
about 66 cm. or more long, very loosely and narrowly panicled nearly to the base;
branches horizontal, commonly branched and fractiflex in age, about 5 cm. or less
distant. Bracts minute, triangular-ovate, much shorter than the slender pedicel-
late ovary. Flowers very small, brownish or dull purple, with spreading segments.
Dorsal sepal oblong-oblanceolate, acute, concave, about 5-6 mm. long. Lateral
sepals narrowly oblanceolate, acute, apparently distinctly longer and narrower
than the dorsal sepal. Petals oblanceolate, acute, somewhat shorter and nar-
rower than the dorsal sepal. Lip strongly reflexed near the middle, with the basal
portion erect and parallel to the column, elliptic-lanceolate and scarcely 5 mm. long
when expanded, acute; disc below with 2 lines or crests which are slightly thick-
ened in front. Column short, very stout, with a pair of minute, lanceolate-
subulate, erect wings near the apex.
Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 171. 1921.
Ecuador (type).
In the Ames Herbarium there is a photograph of the type.
Odontoglossum myrianthum Reichb. f. Xen. Orch. 1: 189,
t. 68, fig. II, 4, 5. 1856. Cyrtochilum myrianthum Kranzl. Notizbl.
Bot. Gart. Berl. 7: 99. 1917, nomen nudum, in key; Kranzl. Engler,
Pflanzenr. IV. 50 (Heft 80), 82. 1922.
Vegetative parts lacking. Inflorescence several- to many-flowered, loosely
paniculate above, with the lower branches flexuous and few-flowered and the upper
portion racemose. Bracts minute, triangular, many times shorter than the slen-
der pedicellate ovary. Parts of the flower spreading. Sepals obovate or spatulate-
obovate, rounded to subacute at the apex, 4.5-5 mm. long, about 1.75 mm. wide.
Petals oblong-obovate, rounded at the apex, with a broad (sessile) base, a little
wider than the sepals. Lip with the basal portion appressed to the column, re-
flexed above, triangular-ovate in outline, lightly 3-lobed below the middle, about
3.5 mm. long; lateral lobes basal, semiorbicular; mid-lobe much larger, porrect,
ligulate- triangular, subacute; disc with a fleshy, carinate callus (between the lat-
eral lobes) which is sulcate above and has a semiorbicular papilla on each side at
its apex. Column very short, stout, wingless.
Peru: Palca (Pavon Herbarium).
No example of this species was available.
Odontoglossum mystacinum (Lindl.) Lindl. Fol. Orch. Odon-
toglossum 6, no. 14. 1852. Cyrtochilum mystacinum Lindl. Bot. Reg.
24, Misc. 30, no. 38. 1838; Bot. Reg. 25: t. 62. 1839. Odontoglossum
rigidum Lindl. Benth. PI. Hartw. 152. 1844; Fol. Orch. Odontoglos-
sum 7, no. 17. 1852. Oncidium rigidum Beer, Prakt. Stud. Orch. 290.
1854.
824 FIELDIANA: BOTANY, VOLUME 30
Plant large. Pseudobulbs pyriform or "oval," compressed, unifoliate, about
3.5 cm. high, surrounded by several distichous, conduplicate, leaf -bearing sheaths.
Leaves ligulate (rarely elliptic-oblong), acute, carinate, apparently about 15 cm.
or less long and up to 1.6 cm. wide. Inflorescence much surpassing the leaves,
loosely paniculate above. Floral bracts lanceolate, navicular, acuminate, sphace-
late, up to 1.5 cm. long, about half as long as the pedicellate ovary. Flowers
2.5-3.5 cm. distant, yellow, medium-sized. Sepals widely spreading. Dorsal sepal
ovate-lanceolate, acuminate, concave, about 1.5 cm. long and 5 mm. wide. Lateral
sepals lanceolate or narrowly elliptic-lanceolate, slightly longer and narrower than
the dorsal sepal, apparently free or nearly so. Petals very similar to the dorsal
sepal but somewhat broader. Lip with the base applied to the column, distinctly
clawed below; claw linear; lamina obovate from a cordate base, slightly pandurate,
with a prominent, recurved, apiculate apex, about 2 cm. long when expanded and
1.2 cm. wide where broadest; disc bilamellate on the claw, with several teeth in
front of the keels. Column small, clavate, with a pair of irregularly lacerate wings
at the apex.
Peru: Habitat not recorded, Harrison s.n. (type of Cyrtochilum
mystacinum). — Amazonas: Chachapoyas, Mathews 168, sub Odonto-
glossum rigidum. Also Ecuador (type of 0. rigidum).
The description was chiefly prepared from the plate of Cyrtochi-
lum mystacinum cited, no material being available.
This taxon appears to be very similar to Oncidium aureum Lindl.,
but has a lip with a deflexed apex and a prominent apicule.
Odontoglossum obscurum C. Schweinf. Bot. Mus. Leafl.
Harvard Univ. 14: 67. 1949. Mesospinidium Wallisii Reichb. f.
Linnaea 41 : 106. 1876, not Odontoglossum Wallisii Linden & Reichb.
f. (1870).
Plant slender, delicate, growing on rocks, about 25 cm. high. Rhizome appar-
ently abbreviated. Pseudobulbs aggregated, ovoid to cylindric, complanate, uni-
foliate, 1-2.5 cm. high, surrounded and often surpassed by 2 or 3 pairs of distichous,
leaf-bearing sheaths. Leaves linear, subobtuse to acute, more or less narrowed to
a conduplicate base, 10-20.4 cm. long, 5.5-12 mm. wide. Scape lateral, basal in
the axil of a sheath, distinctly surpassing the leaves, arcuate-spreading; peduncle
slender, about 21-50 cm. long, with 3 to 6 very small and close, tubular-infundibuli-
form sheaths; panicle either short, 9- to 10-flowered, about 5 cm. long with few
branches above or relatively elongate, loosely many-flowered and up to 25 cm.
long; rachis lightly fractiflex. Flowers small, rather fleshy, with spreading seg-
ments. Dorsal sepal obovate-spatulate from a narrow base, abruptly acute or
apiculate, about 6.7 mm. or less long and 3.8 mm. wide. Lateral sepals obliquely
spatulate or oblong-spatulate, acute, carinate, distinctly longer and narrower than
the dorsal sepal. Petals variable, obliquely oblanceolate to oblong-obovate, apicu-
late, shorter and narrower than the dorsal sepal. Lip with the basal half erect and
parallel to the column in natural position with the anterior half abruptly reflexed,
obtuse or slightly retuse at the apex, ovate-oblong, oblong-elliptic or "oblong-
ligulate" when expanded, lightly retuse in the middle of each side, about 6 mm.
long; disc with a linear, sulcate callus extending through the lower hah" and ending
SCHWEINFURTH: ORCHIDS OF PERU 825
in 2 free teeth or lobules. Column very short and stout, broadly winged on each
side throughout, the wings ending in a blunt triangular tooth.
Peru: Habitat not recorded, Wallis s.n. (type of Mesospinidium
Wallisii).—Cuzco: Prov. of Urubamba, Machu-Picchu, 2040 meters,
on rocks, Vargas 3373.
Odontoglossum praestans Reichb. f. & Warsc. Bonpl. 2: 99.
1854; Rolfe, Lindenia 7: 71, t. 322. 1892; Orch. Rev. 23: 156, fig. 14.
1915.
Plant showy, variable. Rhizome apparently abbreviated. Pseudobulbs ovoid
or pyriform-cylindric, often aggregated, subcompressed, about 6 cm. or less long,
bifoliate, mottled with purplish brown, surrounded by 3 or 4 pairs of distichous,
imbricating, conduplicate sheaths of which the upper ones bear successively larger
leaves. Leaves linear or linear-oblong, abruptly acute and apiculate, more or less
narrowed to a sessile or shortly petioled, conduplicate base, up to 23 cm. long,
1.4 cm. or more wide. Scapes lateral in the axils of sheaths, equaling or exceeding
the leaves, above racemose (or with a large-branched panicle according to Reichen-
bach); peduncle (in the collection seen) 10-11 cm. high, with about 2 close, very
small, tubular sheaths; raceme short to long, rather densely few- (6-) to many-
(15-) flowered, about 10-30 cm. long. Flowers rather large, with widely spreading
segments, greenish yellow spotted with cinnamon-brown or purple. Dorsal sepal
lanceolate or linear-lanceolate, long-acuminate, 2.9-3.6 (rarely 4.5) cm. long,
6-9 mm. wide. Lateral sepals similar, very slightly longer and narrower, some-
what falcate-incurved, dorsally carinate. Petals similar to the lateral sepals, lan-
ceolate or ovate-lanceolate, long-acuminate, lightly oblique, shorter and slightly
wider than the dorsal sepal. Lip simple, lanceolate or narrowly lanceolate, long-
acuminate, 2.5-2.7 cm. long; claw short, erect and parallel to the column, deeply
tubular-concave; lamina with a broadly cuneate or subtruncate base which is
9-10 mm. wide, gently recurved above; disc at the base (of the lamina) with 4
short, stout, longitudinal, irregularly dentate keels of which the inner pair is much
the larger. Column stout, 9-11 mm. high at the back, prominently angled in front
below the middle, with a pair of porrect, more or less elongate, decurved, irregu-
larly lacerate wings above, the most conspicuous portion being at the summit.
Peru: Amazonas [?] sources of the Maranon River, Warscewicz
s.n. (type).— Cuzco: Prov. of Urubamba, Machu-Picchu to Sta. Rita,
2040 meters, on rocks, Vargas 3363.
In the Ames Herbarium are several records of the type.
Odontoglossum pusillum C. Schweinf. Am. Orch. Soc. Bull.
14: 340, t., 1946. Figure 166.
Plant small and slender for the genus, variable, growing on rocks. Rhizome
stout, creeping. Pseudobulbs usually aggregated, oblong-cylindric or rarely oblong-
ellipsoid, strongly compressed, unifoliate, about 2-6 cm. tall, surrounded below by
1 to 3 pairs of distichous, conduplicate, imbricating sheaths which are successively
larger above and commonly leaf-bearing. Leaves linear to linear-oblong, abruptly
826 FIELDIANA: BOTANY, VOLUME 30
rounded with a more or less obliquely bilobed apex, more or less narrowed to a con-
duplicate (often petiole-like) base, 2-19.2 cm. long, 4-10 mm. wide. Scapes lat-
eral, in the axils of sheaths surrounding the pseudobulbs, slightly shorter than the
leaves or much longer, 12-29 cm. long; peduncle complanate, up to 13 cm. long,
with a single, small, close, tubular sheath above the middle; inflorescence racemose
or paniculate with 1 or 2 short, strict branches near the base, loosely few- to many-
flowered, the rachis 1.5-18 cm. long. Floral bracts minute, many times shorter
than the slender pedicellate ovary. Flowers very small for the genus, with widely
spreading segments, yellow orange or greenish brown with whitish, yellow-dotted
lip. Dorsal sepal lightly recurved and tubular-involute in natural position, ob-
long-oblanceolate when expanded, acute at the thickened apex, 7-7.5 mm. long,
2.1-2.4 mm. wide. Lateral sepals connate into a tubular-concave, bilobed lamina
which is curved or lightly sigmoid when viewed from the side, markedly longer and
wider than the dorsal sepal, the terminal lobes ovate and acute or mucronate.
Petals obliquely oblanceolate or oblong-spatulate, falcate-incurved, acute, about
as large as the dorsal sepal. Lip divided into 2 parts; basal portion upright and
parallel to the column, oblong, angled or round-dilated and erect on each side at
the base, about 3 mm. or more long, almost entirely occupied by an oblong, fleshy
callus which is slightly dilated toward the 3-lobed apex, the middle lobe deeply
emarginate; anterior portion abruptly reflexed in course of development, reniform-
ovate or cordate-ovate when expanded, broadly rounded and often obscurely lob-
ulate in front with a minute apicule, truncate or subcordate at the base, 4.2-
4.8 mm. long, 5.2-6 mm. wide. Column stout, short, recurved, clavate, wingless,
3-3.5 mm. long at the back.
Cuzco: Prov. of Calca, Lares Valley above Man toe, 2500 meters,
Weberbauer 7916 (type). Prov. of Urubamba, Machu-Picchu, 2060
meters, Vargas 3338.
Odontoglossum retusum Lindl. Benth. PI. Hartw. 152. 1844;
Fol. Orch. Odontoglossum 13, no. 36. 1852; Hook. f. Bot. Mag. 123:
t. 7569. 1897. Oncidium retusum Beer, Prakt. Stud. Orch. 290. 1854.
Cyrtochilum retusum Kranzl. Notizbl. Bot. Gart. Berl. 7: 99. 1917,
nomen, in key; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 86. 1922.
Plant rather large, epiphytic or growing on rocks. Pseudobulbs ovoid to nar-
rowly ellipsoid, sometimes aggregated, subcompressed, unifoliate, about 2.5-5 cm.
long, surrounded below by 3 or 4 pairs of distichous, imbricating, conduplicate
sheaths which are successively larger and leaf-bearing upward. Leaves (the largest
on the pseudobulb) elliptic-linear to linear-oblanceolate, obtuse to subacute, grad-
ually narrowed below to a channelled, petioled base, up to 29 cm. long and about
2 cm. wide (those on the sheaths less narrowed below). Scape lateral, basal, in
the axil of a sheath, more or less exceeding the leaves; peduncle slender, about
23.5 cm. or more long, provided with about 3 remote, small, close, tubular sheaths;
inflorescence slender, loosely many-flowered, with few to several short, few-flowered
branches, the branches sometimes compound; rachis 12-30 cm. or more long.
Flowers rather small for the genus with spreading segments, orange-scarlet with a
golden yellow lip. Dorsal sepal broadly oblanceolate or narrowly obovate, acute,
deeply concave or subconduplicate, about 1.3-1.6 cm. long and 5.4 mm. wide when
expanded. Lateral sepals oblanceolate, lightly oblique, acute and apiculate, dor-
ODONTOGLO55UM
pusi£Curn G.uc/iweinJ.
FIG 166. Odontoglossum pusillum C. Schweinf. 1, plant; X %. 2, flower in
natural position, oblique view; X 3^. 3, lip and column from side; X 4. 4, lip
spread out, from above; X 4.
827
828 FIELDIANA: BOTANY, VOLUME 30
sally carinate, somewhat longer and narrower than the dorsal sepal. Petals nar-
rowly oblong-obovate, acute, lightly oblique, distinctly shorter and narrower than
the dorsal sepal. Lip oblong-subquadrate, sessile, with the base suberect and
parallel to the column, recurved-spreading above, simple to obscurely 3-lobulate
with rounded basal lobules, subtruncate to retuse in front, apiculate, about 1 cm.
long and 7.6 mm. wide near the base; disc below with a sulcate callus or 2 approxi-
mate calli extending about to the middle. Column dwarf, very stout, about 3 mm.
high at the back, prominently margined on each side throughout.
Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 171. 1921.
Ecuador (type).
Odontoglossum revolutum Lindl. Ann. & Mag. Nat. Hist. 15:
256. 1845; Fol. Orch. Odontoglossum 16, no. 45. 1852. Oncidium
revolutum Beer, Prakt. Stud. Orch. 290. 1854.
Plant large and robust, variable, with a stout, apparently creeping rhizome.
Pseudobulbs congested, apparently unifoliate, entirely concealed by about 4 pairs
of distichous, imbricated, conduplicate sheaths which are successively larger and
leaf -bearing upward; thus is formed a very stout and robust basal portion. Leaves
linear, rigid, abruptly obtuse, sessile, slightly narrowed at the conduplicate base,
up to 30 cm. or more long and 2.4 cm. wide when the strongly revolute sides are
expanded. Scape stout, lateral, in the axil of sheaths, much surpassing the leaves,
about 100 cm. tall in the type; peduncle about 79 cm. tall, with several (about 5 or
more) short, appressed, tubular sheaths. Inflorescence a compound, pyramidal
panicle about 30 cm. or less long, the branches more or less remote on the axis,
densely flowered. Bracts prominent, ovate or lanceolate, cucullate, spathaceous.
Flowers medium-sized, with spreading, recurved segments, apparently yellow.
Sepals conduplicate, with undulate margins, dorsally carinate especially the lat-
eral ones. Dorsal sepal lanceolate-elliptic when expanded, acuminate, clawed,
1.5-3 cm. long, 5.8-11 mm. wide. Lateral sepals very similar, slightly longer and
narrower. Petals lanceolate or elliptic-lanceolate, acuminate, much shorter than
the dorsal sepal and slightly narrower than the lateral sepals. Lip recurved, with
the base erect and parallel to the column and the sides of the anterior portion erect
in natural position, lightly 3-lobed with a suborbicular basal portion and a lanceo-
late or oblong terminal portion, acute or acuminate, with a prominent mucro,
1.4-2 cm. long when expanded, 5.6-7 mm. wide below; disc with a conspicuous
pair of high, thin keels near the base, a much smaller keel in the center, and a group
of 2 or 3 filiform papillae on either side in front. Column small, wingless, about
5.5-8 mm. long at the back.
Junin, fide Schlechter, Fedde Repert. Beih. 9: 171. 1921. Colom-
bia (type).
Odontoglossum ringens Reichb. f. Saund. Refug. Bot. 2: t. 99.
1878.
Plant large. Pseudobulb oblong-ovoid ("oblong"), complanate, bifoliate, ap-
parently about 12 cm. long, surrounded at the base by one or more pairs of short,
distichous, leaf-bearing sheaths. Leaves narrowly oblong or "ligulate," elongate,
SCHWEINFURTH: ORCHIDS OF PERU 829
acute, apparently over 40 cm. long, about 2.5 cm. wide. Scape lateral, basal, ter-
minated by a long, loosely branched panicle of which the branches are short, f racti-
flex and rather densely few- (3- to 8-) flowered. Floral bracts minute many times
shorter than the pedicellate ovary. Flowers medium-sized, with widely spreading
segments pale yellow with brown lines on the sepals and petals and violet lines on
the basal part of the lip. Dorsal sepal oblong-lanceolate, acute, about 2.3 cm. long
and 5 mm. wide. Lateral sepals similar, oblique, often with reflexed tips. Petals
elliptic-lanceolate ("oblong"), acute, oblique, undulate-margined, shorter and
wider than the sepals. Lip rhombic-pandurate, about as long as the petals, sub-
erect at the very base and recurved in front, quadrate below, then abruptly dilated
near the middle to form an ovate-triangular, acute or subacuminate apical portion;
disc at the base with a saddle-like callosity consisting of a pair of high, thin keels
which are dentate below and prolonged in front into 2 or 3 lobes or teeth and with
a small callose keel between them. Column small, clavate, about 8 mm. high,
with a pair of nearly obsolete, angulate wings above and 2 tooth-like blunt lo
in front of the stigmatic cavity.
Peru: Habitat not recorded, Pearce (Veitch's collector) s.n.
No material of this species was available.
Odontoglossum subuligerum Reichb. f. Linnaea 41: 27. 1876.
Plant large. Leaves lanceolate, acute. Scape slender, apparently over 35 cm.
long with several (4 or more) short, close, tubular, very distant sheaths. Inflores-
cence short, racemose, distantly few- (2- or 3-) flowered. Floral bracts much
shorter than the pedicellate ovary. Sepals similar, lanceolate or oblong-lanceolate,
acuminate, about 3-3.5 cm. long and 8 mm. wide. Petals similar to the sepals,
slightly shorter. Lip shorter than the other segments, about 2.4 cm. long, with a
small linear claw which is adnate to the column at the base; lamina "oblong-
ligula'te" or obovate-oblong, dilated above, acute, plicate-denticulate in front; disc
at the base with 7 calli in 2 series on each side, the upper 4 filiform, the anterior ones
triangular-setaceous and runcinate at the back. Column arcuate, trigonous, about
1.5 cm. high, angled above the base, with a pair of wings which are rhombic at
the apex.
Peru: Habitat and collector not recorded.
No material of this taxon was available, but there is a record of
the type in the Ames Herbarium.
Odontoglossum tigroides C. Schweinf. Am. Orch. Soc. Bull. 14:
22, t., 1945. Figure 167.
Plant medium-sized, epiphytic, with an apparently abbreviated rhizome.
Pseudobulb narrowly ovoid to cylindric-pyriform, strongly complanate, bifoliate
or trifoliate, 3.5-5 cm. high, surrounded below by about 3 pairs of distichous,
deeply imbricating sheaths of which the upper pair bears small, conduphcate
blades. Leaves oblong to oblong-spatulate, abruptly rounded at the apex, grad-
ually narrowed below to a conduplicate, petiole-like base, 9-13 cm. long, 1.3-
1.7 cm. wide. Scapes lateral, basal in the axils of the inner sheaths, much exceeding
the leaves, 20.5-32 cm. long, suberect to spreading in youth, becoming arcuate or
830 FIELDIANA: BOTANY, VOLUME 30
pendent in age, one on either side of the pseudobulb; peduncle rather stout, com-
planate, 3-10 cm. long, with 1-3 remote, stout, infundibuliform sheaths; raceme
subdensely or loosely many-flowered, somewhat looser below, sometimes inter-
rupted in the middle by several sterile, imbricated, infundibuliform bracts; rachis
about 20 cm. or less long, becoming fractiflex in age. Floral bracts conspicuous,
broadly ovate, amplexicaul, equaling or slightly surpassing the pedicellate ovary.
Flowers racemose, small for the genus, fragrant, strongly distichous, with widely
spreading, fleshy segments; sepals and petals yellowish white with 2 broad, trans-
verse, brown-red bands, lip white with 2 brownish red dots. Sepals and petals with
the lower portions revolute-convex and the anterior parts recurved and undulate.
Dorsal sepal broadly obovate or suborbicular-obovate when expanded, broadly
rounded above with a minute apicule, up to 12.8 mm. long and 8.6 mm. wide when
expanded. Lateral sepals similar but a little longer and narrower, obovate-spatu-
late, broadly obtuse or subacute, suboblique. Petals obliquely oblong-obovate,
obtuse to rounded above, shorter than the dorsal sepal and narrower than either
sepal. Lip much smaller than the other segments, with the lower portion erect
and parallel to the column and the larger anterior part reflexed-spreading, lightly
3-lobed above the middle, about 8.8 mm. long in natural position; lateral lobes
semiorbicular, convex-re volute; mid-lobe much smaller, convex, broadly ovate
when expanded, abruptly narrowed near the middle to an acute tip; disc between
the lateral lobes with a pair of fleshy, ellipsoid, more or less indented, diverging
calli. Column short, stout, straight, about 5 mm. high at the back, with a pair of
conspicuous, falcate-semiovate wings at the summit.
Huanuco: Yanano, about 1800 meters, on mossy tree, Macbride
3840.
Odontoglossum trilobum Schltr. Fedde Repert. Beih. 9: 111.
1921; Mansf. Fedde Repert. Beih. 57: t. 128, nr. 503. 1929.
Plant large, epiphytic, with a short rhizome. Pseudobulbs compressed, ovoid
or oblong-ovoid, bifoliate, 9-12 cm. tall, surrounded on each side by one or more
pairs of conduplicate, leaf-bearing sheaths. Leaves narrowly oblong or lorate,
subobtuse, gradually narrowed below to a conduplicate, petiole-like base, up to
41 cm. long and 3.5 cm. wide. Scape lateral, basal in the axil of a sheath, much
surpassing the leaves, gently arcuate, loosely paniculate above, up to 200 cm. tall;
peduncle about 30-60 cm. long, with several (6 or more) short, remote, tubular,
evanescent sheaths; panicle narrow, elongate, many-flowered, with rather remote,
loosely 2- to 4-flowered, fractiflex branches. Bracts very small, deltoid, concave,
much shorter than the slender, pedicellate ovary. Flowers medium-sized, with
widely spreading segments, light green spotted with brown and white. Sepals
and petals with lightly undulate margins. Dorsal sepal oblong-lanceolate or ellip-
tic-lanceolate, about 2.5 cm. long and 7 mm. wide, acute, dorsally keeled at the
apex. Lateral sepals very similar, slightly oblique, as long as the dorsal sepal and
very little narrower. Petals obliquely lanceolate-oblong, abruptly acute and apic-
ulate, very slightly shorter than the sepals, about as wide as the lateral sepals.
Lip rhombic-lanceolate in outline, sharply 3-lobed below the middle, about 1.8-
2.4 cm. long and 1-1.4 cm. wide across the lateral lobes, spreading from an erect,
narrowly cuneate, claw-like base; lateral lobes spreading, obliquely dolabriform or
semiobovate-rounded, with irregular anterior margins; mid-lobe oblong-lanceolate,
FIG 167. Odontoglossum tigroides C. Schweinf. 1, plant; X ^. 2, flower,
natural position, front view; X 2. 3, lip and column, obliquely from side; >
4, lip expanded, from above; X 4. 5, capsule; X %•
831
832 FIELDIANA: BOTANY, VOLUME 30
obtuse to subacute, apiculate, about 1-1.4 cm. long and 4 mm. wide; disc from
the base of the lip through the lower third with an oblong, fleshy callus which is
slightly dilated above with a pluridentate upper portion and a bilobed apex.
Column relatively short and straight, about 7 mm. high at the back, with a pair
of minute, obliquely ovate-lanceolate, falcate wings above.
Junin: Near La Merced, in the Chanchamayo Valley, Kohler
brothers s.n. (type). Schunke Hacienda, above San Ramon, 1300-
1700 meters, in dense forest, Schunke A64 (this specimen is very poor
and fragmentary; hence questionable). — Loreto: Pumayacu, between
Balsapuerto and Moyobamba, 600-1200 meters, in forest, King 0.16.
Odontoglossum tripudians Reichb. f. & Warsc. Bonpl. 2: 100.
1854; Hook. f. Bot. Mag. 99: t. 6029. 1873; Cogn. & Goos. Diet. Icon.
Orch. Odontoglossum t. 27. 1902.
A rather large and showy plant. Pseudobulbs oblong-ovoid or ellipsoid, com-
pressed, bifoliate, 6-10 cm. high, apparently approximate, surrounded below by 1
or 2 pairs of distichous, conduplicate sheaths which are commonly leaf-bearing.
Leaves oblong-oblanceolate, acute or acuminate, long-narrowed below with a con-
duplicate base, 17.7-25 cm. long, about 2.5 cm. wide. Scape lateral, basal in the
axil of a sheath, exceeding the leaves, arcuate, about 33 cm. long, terminating in a
loosely 8- to 12-flowered raceme which is 18.5 cm. or more long. Bracts very small,
triangular-lanceolate, much shorter than the pedicellate ovary. Flowers rather
large, with widely spreading segments, dull yellow-green on the back. Dorsal
sepal lanceolate-elliptic, acute or acuminate, 2.4-3.5 cm. long, 6-13 mm. wide,
maroon with a yellow tip. Lateral sepals*similar but slightly narrower, elliptic-
lanceolate, suboblique, acuminate. Petals similar to the lateral sepals but a little
smaller, oblique, acuminate, sometimes with a few spreading teeth on the sides,
golden yellow with very broad, maroon blotches. Lip with a short, narrow, erect
claw parallel to the column, 2.3-2.7 cm. long when expanded, white or yellowish
with rose or violet blotches; lamina subquadrate-pandurate, the basal part cordate-
ovate with erose or crenulate margins and the anterior part suborbicular and apic-
ulate, with irregularly laciniate margins; disc near the base with about 10 radiating
keels, the inner 4 larger and produced into spinous processes. Column stout,
arcuate, 1.3-1.5 cm. long, with a pair of irregularly lacerate or deeply 3-toothed
wings above.
Peru: Habitat not recorded, Warscewicz s.n. (type). Also Co-
lombia.
Odontoglossum Wyattianum G. Wilson, Orch. Rev. 36: 47.
1928.
This is an obscure species, since no vegetative features were de-
scribed, and even the origin is uncertain. Consequently it was not
entered in the key.
Flowers said to be similar to those of O. Harryanum but smaller and not so
highly colored, 5-7.6 cm. in vertical diameter. Sepals up to 3.8 cm. long, golden
brown with a triangular white blotch at the base. Petals about equal to the sepals
SCHWEINFURTH: ORCHIDS OF PERU 833
but more pointed. Lip about 3.8 cm. long, purple and white, broadly ventricose,
trumpet-like in profile, contracted at the base, 3-lobed; lateral lobes large, rounded,
erect; mid-lobe spreading, bilobed. Column white lined with red-brown, having
a pair of yellow pointed wings above the stigma.
Peru?
SYMPHYGLOSSUM Schltr.
A very small genus of American epiphytes, at present limited to
two species growing in Ecuador and Peru.
Plants with quite the habit and general appearance of Cochlioda, to which they
have been generally united, but they appear to be distinct by reason of certain
floral characters. Lateral sepals connate through the lower half (free or nearly
so in Cochlioda); petals adnate to the column below (free in Cochlioda); lip simple
(3-lobed in Cochlioda); column bearing a simple stigmatic orifice (2 separate ones
in Cochlioda).
Symphyglossum sanguineum (Reichb. f.) Schltr. Orchis 13:
9. 1919. Mesospinidium sanguineum Reichb. f. Walp. Ann. 6: 858.
1864; Batem. Bot. Mag. 93: t. 5627. 1867. Cochlioda sanguinea
Benth. & Hook. f. Gen. PI. 3: 560. 1883; Cogn. Martius Fl. Bras. 3,
pt. 6: 208, t. 44, fig. 1. 1905.
Plant medium-sized to large, epiphytic. Rhizome abbreviated. Pseudobulbs
aggregated (rarely solitary), oblong-ovoid or ellipsoid, strongly compressed, 3-7 cm.
long, usually bifoliate, surrounded at the base by several pairs of distichous, deeply
imbricating sheaths of which the longer uppermost ones are leaf-bearing. Leaves
variable, elliptic-oblong to linear, acute, long-narrowed below to a conduplicate,
petiole-like base, 6.3-25 cm. long, 7-20 mm. wide, subcoriaceous, with the mid-
nerve carinate beneath. Scape lateral, basal in the axil of an inner sheath, erect
to nodding, sometimes loosely branched, 5-50 cm. long, often much surpassing the
leaves; peduncle with several remote, closely clasping, short sheaths; flower cluster
usually racemose, rather laxly few- to many-flowered. Floral bracts small, trian-
gular-ovate or lanceolate, acute or acuminate, much shorter than the slender
pedicellate ovary. Flowers rather small, rose-purple with a whitish rose lip, sub-
secund, the subspreading segments rather fleshy. Dorsal sepal elliptic to oval-
obovate, deeply concave, broadly rounded and minutely apiculate at the apex,
10-15 mm. long, about 7 mm. wide when expanded. Lateral sepals connate to the
middle or above forming a broadly obovate, concave lamina which is about as long
as the dorsal sepal but considerably broader, with rounded, subacute lobes. Petals
similar to the dorsal sepal but a little smaller, with the central basal part adnate
to the column, oval-obovate, abruptly subacute. Lip fleshy, shorter than the sepals,
the center adnate to the column to the upper part of the latter with the margins
free, the lower portion parallel to the column, the anterior portion deflexed, entire,
ovate, concave and narrowed to a subacute point; disc with a pair of high, semi-
orbicular plates at the base of the free lamina. Column short, gradually much
dilated upward, about 4.7-9 mm. long at the back.
834 FIELDIANA: BOTANY, VOLUME 30
Peru: Near the source of the Maranon River, Warscewicz s.n.
(type of Mesospinidium sanguineum). — Cajamarca, fide Schlechter,
Fedde Repert. Beih. 9: 169. 1921. Also Ecuador.
BRASSIA R. Br.
A rather small but confusing genus of American epiphytes extend-
ing from southern Florida (U.S.A.), the West Indies and Mexico,
through Central America and South America to Brazil, Bolivia and
Peru.
Plants small to very large, but usually medium-sized, often with a stout rhi-
zome. Pseudobulb subglobose-ovoid or oblong-ovoid to oblong-cylindric (rarely
elongate-cauliform), 1- to 2-leaved (rarely 3- to 5-leaved), surrounded on each side
below by 2 or more pairs of imbricating, often leaf-bearing sheaths. Leaves elliptic-
oblong to oblong-oblanceolate or rarely linear. Scapes lateral, basal, axillary,
shorter to much longer than the leaves, terminated by a lax to dense raceme or
rarely a panicle, few- to many-flowered. Floral bracts commonly much shorter
than the pedicellate ovary. Flowers small or usually large, with more or less spread-
ing segments. Sepals similar, free, narrowly lanceolate or linear-lanceolate, acumi-
nate or with an elongate, caudate apex. Petals similar to the sepals but more or
less smaller. Lip commonly sessile, spreading from the base of the column, simple
or pandurate, shorter and much wider than the other segments, commonly with 2
or more keels or calli at the base. Column very short, stout, footless, wingless.
Anther terminal, opercular, incumbent. Pollinia 2, obovoid, unappendaged.
Al. Inflorescence paniculate, with several spreading-ascending branches.
B. thyrsodes
A2. Inflorescence simply racemose, few- to several-flowered I
la. Lip pandurate in outline 1
Ib. Lip not pandurate 4
la. Anterior part of the lip strongly dilated, gradually acuminate above; calli in
front of the basal pair very short and broad B. cryptophihalma
Ib. Anterior part of the lip not strongly dilated, abruptly pointed or aristate ... 2
2a. Basal keels of the lip confluent into a broad apex; flowers numerous (about 16) ;
sepals and petals transversely chocolate-barred near the base . B. Forgetiana
2b. Basal keels of the lip separate, not confluent above; flowers few (about 10 or
less) ; sepals and petals apparently not transversely barred 3
3a. Anterior part of the calli of the lip not terminating in a pair of arcuate keels;
flowers relatively large, lateral sepals over 5 cm. long B. signata
3b. Anterior part of the calli of the lip terminating in a pair of long, arcuate keels;
flowers relatively small, lateral sepals 4 cm. or less long; column dilated at
the base B. arcuigera
4a. Lamina of the lip with a subquadrate-oblong lower third, then abruptly
rhombic-dilated above; basal callus bicuspidate in front B. Filomenoi
4b. Lamina of the lip without a subquadrate-oblong lower third, not abruptly
dilated above 5
5a. Base of the lip with 6 distinct calli; flowers fleshy; sepals 2-2.6 cm. long.
B. villosa (B. hexodonta)
5b. Base of the lip with 2 or 4 distinct calli 6
SCHWEINFURTH: ORCHIDS OF PERU 835
6a. Flowers small, lateral sepals 3.7 cm. or less long 7
6b Flowers large, lateral sepals 4.5 (rarely 4) cm. or more long 9
7a. Leaf linear-ligulate, 6-10 cm. long and 1.3 cm. or less wide; lip with 2 simple,
longitudinal keels at the base B- elegantula
7b. Leaf oblong or oblanceolate, 16-24 cm. long, 4 cm. or more wide; lip witl
2 pairs of 2 calli at the base
8a Sepals 3.4-3.7 cm. long; lip obovate-oval, abruptly long-acuminate, about
' 2.5 cm. long B.jumnensis
8b Sepals about 2 cm. long; lip elliptic-lanceolate or oblong-ovate, shortly acu-
minate, about 1.3 cm. long B- KoeMe
9a Pseudobulbs elongate-cauliform, about 35 cm. or more tall, remotely 3- to
' 5-leaved B' «*«K/omw
9b. Pseudobulbs oblong-cylindric to ovoid-subglobose, 18 cm. or less tall, sub-
approximately 1- or 2-leaved (very rarely 3-leaved) 1"
lOa Lip more or less verrucose above; anterior pair of calli prominent and
divergent E' blde
lOb. Lip not verrucose above; anterior pair of calli consisting of erect, often
minute teeth
11 a. Petals about as long as the dorsal sepal or somewhat shorter 1*
lib. Petals about half as long as the dorsal sepal or less
12a Pseudobulbs oblong-cylindric to oblong-ovoid, 2-leaved; leaves subobtuse;
sepals 5 mm. wide B- buolor
12b Pseudobulbs ovoid-subglobose, 1-leaved; leaves short-acuminate; sepals
' 2.5 mm. or less wide B. perunana
13a Pseudobulbs broadly ellipsoid to ovoid-oblong, commonly 1-leaved; basal
callus of the lip low, consisting of 1 low, sulcate callus or 2 approximate calli,
usually terminating in an erect tooth -B. longissima
13b Pseudobulb oblong-cylindric, 2- or (very rarely) 3-leaved; basal calli of the
lip separate and commonly high, with 2 conspicuous, usually separate teeth
in front B. caudata
Brassia arcuigera Reichb. f. Gard. Chron. (1869) 388; Rolfe,
Orch. Rev. 25: 222. 1917.
Vegetative parts not described. Raceme (incomplete in the drawing of the
type from the Reichenbach Herbarium) loosely several-flowered; rachis about
8 cm. long, gently arcuate, about 7-flowered. Floral bracts triangular-lanceolate,
acute or acuminate, amplexicaul and concave, much shorter than the pedicellate
ovary. Flowers relatively small for the genus, with spreading segments. Sepals
subequal, "linear" or linear-lanceolate, acuminate, 3-4 cm. long. Petals very sim-
ilar, but markedly shorter. Lip about as long as the petals, lightly pandurate,
"oblong" or ovate-oblong, "aristate" or very abruptly acuminate, sessile; disc at
the base with a pair of "semioblong" or obliquely oblong, fleshy calli which are
produced in front into conspicuous arched or arcuate-diverging keels. Column
short, dilated at the base.
Peru: Habitat and collector not recorded; introduced by Messrs.
Veitch.
No specimen of this species has been seen, but in the Ames Her-
barium are drawings of the type from the Reichenbach Herbarium.
836 FIELDIANA: BOTANY, VOLUME 30
Brassia bicolor Rolfe, Lindenia 8: 87, t. 378. 1893.
Plant rather large and showy. Pseudobulbs aggregated, oblong-cylindric or
oblong-ovoid, subcompressed, bifoliate, up to about 12 cm. high, surrounded at
the base by 1 or 2 pairs of distichous, long-acuminate, imbricating sheaths of which
the larger inner one is often leaf-bearing. Leaves "linear-oblong" or elliptic-
oblanceolate, subobtuse, about 22 cm. or less long and 6 cm. or less wide. Scapes
erect, with a subdensely flowered raceme. Flowers about 8, rather large, with
widely spreading segments, greenish yellow spotted or deeply banded below with
blackish brown. Dorsal sepal linear-lanceolate, long-acuminate, about 5 cm. long
and 5 mm. wide. Lateral sepals similar to the dorsal sepal but somewhat longer,
oblique. Petals similar to the lateral sepals but shorter and slightly narrower than
the dorsal sepal. Lip "lanceolate-oblong" or rhombic-lanceolate, rather long-
acuminate, about 3.5 cm. long and 1 cm. wide, with slightly undulate margins;
disc at the base with a pair of small keels which are parallel, fleshy, erect, glabrous,
and terminate in an erect tooth at the apex. Column clavate, about 7 mm. long.
Peru: Habitat and collector not recorded; introduced by Messrs.
Linden.
No material of this species was available.
Brassia bidens Lindl. Bot. Reg. 30, Misc. 6, no. 6. 1844; Fol.
Orch. Brassia 3, no. 4. 1853; Cogn. Martius Fl. Bras. 3, pt. 6: 266.
1905. Brassia Lanceana Lindl. var. viridiflora Hook. Bot. Mag. 67:
t. 3794. 1841. Oncidium bidens Reichb. f. Walp. Ann. 6: 769. 1863.
Pseudobulb cylindric-oblong or rarely narrowly cylindric, compressed, 2- or
(very rarely) 3-leaved, 6-13 cm. high, surrounded below by 2 or 3 pairs of dis-
tichous, imbricating sheaths which are successively larger upward and occasionally
leaf-bearing. Leaves oblong to oblong-elliptic or oblong-oblanceolate to oblanceo-
late-elliptic, acute or subacute, more or less cuneate below, 14-29.5 cm. long,
2.2-7 cm. wide, subcoriaceous. Scape lateral, basal, in the axil of a sheath, com-
monly equaling or exceeding the leaves, suberect to lax; peduncle 23 cm. or less
long, with 3 or more remote, short, infundibuliform sheaths; raceme subdensely
5- to 15-flowered, 11-23.5 cm. long. Floral bracts very small, infundibuliform.
Flowers rather large, with widely spreading segments, green or yellow-green, with
brown or blackish spots. Dorsal sepal linear-lanceolate, long-acuminate or caudate,
up to 7 cm. long and 6 mm. wide, concave below, tubular-involute above. Lateral
sepals very similar, but longer and slightly narrower, dorsally carinate. Petals
similar to the sepals, but markedly shorter than either, obliquely linear-lanceolate,
long-acuminate. Lip much shorter and broader than the other segments, rhombic-
obovate or rhombic-oblanceolate, more or less abruptly acuminate, sessile at the
rounded base, up to 4 cm. long and 2.3 cm. wide above the middle, undulate; disc
usually prominently verruculose through the center, at the base with a pair of
approximate, low, fleshy keels which terminate in a pair of conspicuous, erect-
spreading, triangular teeth. Column very small, about 5 mm. or less high at the
back.
Amazonas: Prov. of Bagua, canyon of Rio Maranon, between
km. 247 and 250 along Mesons-Muro Highway, Hutchison 1532.—
SCHWEINFURTH: ORCHIDS OF PERU 837
Loreto: Mishuyacu, near Iquitos, 100 meters, in forest, Klug 1199.
Vicinity of Iquitos, 100 meters, epiphyte, Klug 10008. — San Martin:
San Roque, 1350-1500 meters, Williams 7717. Also Brazil (type),
British Guiana, and Venezuela.
Brassia caudata (L.) Lindl. Bot. Reg. 10: t. 832. 1824; Fol. Orch.
Brassia 3, no. 5. 1853; Hook. Bot. Mag. 62: t. 3451. 1835; Cogn.
Martius Fl. Bras. 3, pt. 6: 257, t. 58. 1905. Epidendrum caudatum
L. Sp. PL ed. 2: 1349. 1763. Malaxis caudata Willd. Sp. PL 4: 93.
1805. Oncidium caudatum Reichb. f. Walp. Ann. 6: 766. 1863.
Brassia Lewisii Rolfe, Orch. Rev. 1: 199. 1893. Figure 168.
Plant epiphytic, very variable. Pseudobulbs often aggregated, oblong-cylin-
dric (commonly narrowly so), strongly compressed and ancipitous, 6-15 cm. long,
bifoliate (very rarely trifoliate), surrounded below by 3 or 4 pairs of distichous,
imbricated sheaths which are successively larger upward and occasionally leaf-
bearing. Leaves elliptic-oblong to oblong-oblanceolate, abruptly acute, more or
less long-cuneate below, 12.5-35.5 cm. long, 2.1-5 (rarely 5.7) cm. wide. Scapes
shorter or longer than the leaves, suberect to nodding; peduncle with 3 or more,
mostly remote, short, close, tubular-infundibuliform sheaths; raceme loosely to
subdensely 4- to 15-flowered, with the rachis up to 16.5 cm. (rarely 28 cm.) long.
Floral bracts triangular-ovate or triangular-lanceolate, deeply concave, amplexi-
caul, much shorter than or subequaling the ovary. Flowers rather large, with
widely spreading segments, greenish yellow with brown or reddish spots. Dorsal
sepal narrowly lanceolate, long-caudate, concave below, tubular-involute above,
up to 8 cm. long and 6 mm. wide. Lateral sepals lanceolate-linear, oblique, much
longer than the dorsal sepal (often twice as long) and somewhat narrower, navicu-
lar. Petals falcate-lanceolate and incurved toward the dorsal sepal, shortly cau-
date, about half as long as the dorsal sepal or less. Lip somewhat shorter or longer
than the petals, ovate-oblong, more or less abruptly long-acuminate or aristate,
rounded and sessile at the base, about 3.5 cm. long and 1.5 cm. wide or smaller;
disc at the base with a pair of short, approximate, usually high, pubescent keels and
a pair of small, erect teeth in front. Column very short and stout, about 5 mm.
high at the back.
Loreto: Mishuyacu, near Iquitos, 100 meters, in forest, Klug 1047,
1284- Vicinity of Iquitos, 100 meters, on tree in dense forest, Klug
10007. Also Florida (U.S.A.), Mexico, through Central America, the
West Indies (type of Epidendrum caudatum) and northern South
America to Surinam, Ecuador, Bolivia and Peru.
Brassia cauliformis C. Schweinf. Am. Orch. Soc. Bull. 15: 132,
t., 1946. Figure 169.
Plant tall, robust. Pseudobulb elongate-cauliform, up to 35 cm. or more tall,
remotely 3- to 5-leaved (the interval between the lowest leaves 3.5-10 cm.), sur-
rounded below by two pairs of imbricating, conduplicate, usually leaf-bearing
sheaths. Leaves of two forms: blades on the upper sheaths oblong-oblanceolate
or elliptic-lanceolate, short-acuminate, long-narrowed to a sessile, conduplicate
FIG. 168. Brassia caudata (L.) Lindl. 1, plant; X %. 2, base of lip, showing
calli and column, front-side view; X 2}4. 3, base of lip, showing calli from above,
with column removed; X 2J^.
838
Cif
cau Ci fformis
FIG. 169. Brassia cauliformis C. Schweinf. 1, plant; X l/w. 2, inflorescence;
X Vi. 3, lip and column, obliquely from side; X 1. 4, lip from above; X 1.
839
840 FIELDIANA: BOTANY, VOLUME 30
base, 19-43 cm. long, up to 4.7 cm. wide; blades on the pseudobulb linear-oblan-
ceolate, short- or long-acuminate, very long-narrowed to a channelled base, up to
59.5 cm. long and 4 cm. wide. Scapes lateral, basal in the axil of a sheath, some-
times about 1.5 m. tall (according to the collector's notes), robust, with one or
more remote, small, close, amp lexi caul sheaths; raceme loosely 12- to 14-flowered,
rachis about 28.5 cm. or less long. Floral bracts ovate-lanceolate, acuminate, am-
plexicaul, up to 1.6 cm. long, much shorter than the pedicellate ovary. Flowers
variable in size and often rather large, with widely spreading segments, greenish
yellow with a brown-spotted lip or yellow. Dorsal sepal linear-lanceolate, acumi-
nate, 3.8-7.4 cm. long, 5-6 mm. wide. Lateral sepals similar but very slightly
larger, lightly oblique and navicular. Petals lanceolate-linear, acuminate, dis-
tinctly shorter than the sepals. Lip ovate-lanceolate or rhombic-lanceolate, acumi-
nate, 3-5.6 cm. long, 1.3-2.9 cm. wide, with the margins (except near the base)
crenulate and undulate; disc at the base with a pair of short, glabrous keels which
are low and approximate below, then diverging and semielliptic-dilated above
(when viewed from the side). Column very short and stout, about 5 mm. high
at the back.
Cuzco: Prov. of Convention, Hda. Potrero, Sapan-Sachayocc,
2200 meters, on open grass slope, Vargas 2528. Prov. of Quispican-
chis, Chaupichaca, Marcapata Valley, 1600-1700 meters, in "shrub-
wood," Weberbauer 7823 (type).
Brassia cryptophthalma Reichb. f. Gard. Chron. n. ser. 12:
554. 1879.
Vegetative portion and lower part of the inflorescence lacking. Raceme short,
arcuate, subdensely several- (about 7-) flowered, with a purple-violet rachis.
Floral bracts small, lanceolate-ovate, amplexicaul, much shorter than the pedicel-
late ovary. Flowers rather large, with widely spreading segments, pale yellow
with dark violet lower portions to the sepals and the lip deeper yellow with a few
dark violet spots. Sepals linear-lanceolate, long-acuminate or "aristate," 6-7 cm.
long, the lateral ones somewhat longer than the dorsal and slightly oblique. Petals
similar to the lateral sepals, but shorter than the dorsal. Lip pandurate, oblong-
obovate in outline and strongly dilated above, acuminate, rounded at the sessile
base, about 4.5 cm. or more long and 2.1 cm. wide above; disc at the base with 4
smooth keels in two series; posterior keels approximate, oblong becoming lower in
front and ending in a sharp, upcurved tooth; anterior keels broadly oblong, short
and diverging. Column very short and stout.
Peru: Habitat and collector not recorded; introduced by Messrs.
Mayer of Karlsruhe.
No example of this species was available, but in the Ames Her-
barium there are drawings of this plant from the Reichenbach
Herbarium.
Brassia elegantula Reichb. f. Gard. Chron. n. ser. 24: 616.
1885; Rolfe, Orch. Rev. 13 : 200. 1905. Oncidium elegantulum Reichb.
f. Gard. Chron. n. ser. 24: 616. 1885.
SCHWEINFURTH: ORCHIDS OF PERU 841
Plant very small for the genus, tufted, about 15 cm. high including the inflores-
cence Rhizome abbreviated. Pseudobulbs "rather small, glaucous, ' otherwise
not described. Leaves linear or "cuneate-ligulate, rigid, acute, glaucous, 6.3-
10 2 cm long, 8-13 mm. wide. Inflorescence slender; raceme nearly 8 cm. long,
few- (2- 'to 5-) flowered, loose to subdense. Floral bracts relatively large, ovate or
"triangular," acute, deeply concave and amplexicaul, shorter than the pedicellate
ovary Flowers small, with widely spreading segments, green with brown trans-
verse bars and a white lip having a few madder-purple dots; about 2.4 cm. in
diameter Dorsal sepal lanceolate or oblong-lanceolate, acute, about 1.2 cm. long.
Lateral sepals about equal, deflexed, falcate-incurved. Petals similar to the dorsal
sepal but distinctly shorter, erect. Lip oblong-ovate or "oblong," rather abruptly
acute or "apiculate," with the upper half reflexed; disc at the base with a pair of
longitudinal keels which are hairy inside and abruptly inflexed in front. Column
very short, wingless, green and brown.
"Peru," fide Schlechter, Fedde Repert. Beih. 9: 172. 1921. Hab-
itat uncertain; said to be imported from Mexico by F. Sander.
No example of this species was available, but in the Ames Her-
barium are drawings of the type from the Reichenbach Herbarium.
Brassia Filomenoi Schltr. Fedde Repert. Beih. 9: 111. 1921;
Mansf. Fedde Repert. Beih. 57: t. 129, nr. 504. 1929.
Plant epiphytic, probably large, vegetative parts lacking. Scape up to 40 cm.
long- peduncle distantly few-sheathed, raceme loosely 10- to 15-flowered, erect-
spreading, secund, up to 20 cm. long. Floral bracts oblong-deltoid, apiculate,
much shorter than the ovary. Flowers showy, rather large; sepals and petals
greenish yellow with dark brown at the base; lip sparingly spotted with dark brown.
Sepals lanceolate-linear or "linear," long-acuminate, diverging, 6.3-6.5 cm long,
the lateral ones oblique. Petals very similar to the sepals but a little shorter,
oblique. Lip obovate-oblong in outline, about 4 cm. long and 1.5 cm. wide near
the middle; claw very short; lamina quadrate-oblong through the lower third,
rounded at the base, then rhombic-dilated and narrowed above into an elongate,
gradually acuminate point; disc at the base with a pair of approximate, fleshy,
subglabrous calli which are abruptly lobulate-dilated above and very shortly
cuspidate in front on each side. Column short, about 6 or 7 mm. high, glabrous.
Loreto: Near Moyobamba, Filomeno s.n.
No specimen of this species was accessible.
Brassia Forgetiana Hort. ex Gard. Chron. ser. 3, 48: 434, 463,
471, fig. 209. 1910, nomen tantum; C. Schweinf. Bot. Mus. Leafl.
Harvard Univ. 12: 196. 1946.
Plant large, allied to B. maculate R. Br., but heretofore without substantial,
definite diagnosis. Description drawn from the detailed illustration cited. Leaf
large, oblong or elliptic-oblong, abruptly subacute or rounded at the apex, appar-
ently over 30 cm. long and 6-7 cm. wide. Raceme arcuate, densely many-flowered,
the rachis nearly 20 cm. long. Floral bracts ovate or ovate-lanceolate, acuminate,
clasping, about one third as long as the pedicellate ovary. Flowers medium-sized
842 FIELDIANA: BOTANY, VOLUME 30
for the genus, with horizontally spreading segments, greenish white with trans-
verse chocolate or chocolate-purple bars near the base of the sepals and petals.
Dorsal sepal narrowly lanceolate, long-acuminate, concave, about 6 cm. long and
6 or 6.5 mm. wide below the middle. Lateral sepals linear-lanceolate, long-acumi-
nate, oblique, slightly falcate-incurved, a little longer than the dorsal sepal. Petals
obliquely linear-lanceolate, long-acuminate, falcate-incurved, somewhat more than
half as long as the dorsal sepal and slightly narrower at the base where broadest.
Lip pandurate-oblong, rounded and very abruptly long-cuspidate at the apex, ses-
sile at the rounded base, nearly 4.5 cm. long, about 1 cm. wide near the base and
0.9 cm. wide near the apex; disc at the base with a pair of approximate, longitu-
dinal, fleshy calli which are fused above into a somewhat dilated, abruptly truncate
and 3-lobulate apex. Column very short and stout, about 6 mm. high.
Peru: Habitat not recorded, Forget s.n.; introduced by Messrs.
Sander.
No example of this species was available.
Brassia juninensis C. Schweinf. Am. Orch. Soc. Bull. 14: 248,
t., 1945. Figure 170.
Plant medium-sized for the genus, epiphytic, with a stout rhizome. Pseudo-
bulbs oblong-cylindric, strongly compressed, obliquely bifoliate at the apex, 8-
12 cm. high, surrounded below by 2 or 3 pairs of imbricating sheaths which are
progressively larger upward and rarely leaf-bearing. Leaves elliptic-oblong to
broadly oblanceolate or oblong-oblanceolate, more or less abruptly acute, cuneate
or long-cuneate at the complicate base, 16-20.5 cm. long, up to 4.4 cm. wide.
Scape lateral, basal in the axil of a sheath, shorter than the leaves or rarely
slightly surpassing them, suberect or rarely with a reflexed raceme; peduncle
7-14 cm. long, commonly with several short sheaths, which are remote above;
raceme loosely 6- to 12-flowered. Floral bracts small, triangular-ovate, amplexi-
caul, acute or acuminate, about one half as long as the pedicellate ovary or less.
Flowers rather small, subfleshy, with spreading segments, pale greenish cream-
color with the petals brown-purple below and the lip brown-spotted. Dorsal sepal
linear-lanceolate or elliptic-linear, long-acuminate, concave below, about 3.4 cm.
long, 3.5-6.5 mm. wide. Lateral sepals obliquely linear-lanceolate, acuminate,
dorsally carinate with the keel produced into a prominent mucro, a little longer and
very slightly narrower than the dorsal sepal. Petals linear or linear-lanceolate,
long-acuminate, lightly oblique, markedly shorter and narrower than the sepals.
Lip sessile, obovate-oval or oblong-obovate when expanded, with the apical por-
tion recurved in natural position, abruptly long-acuminate, about 2.5 cm. long and
1.4 cm. wide, with crenulate margins especially above; disc at the base with a pair
of very small, approximate, fleshy, pubescent calli which become lower beyond the
middle and with 2 fleshy ovoid-dentiform, spreading calli just in front. Column
very short and stout, about 5 mm. high at the back, with a large stigmatic
cavity.
Junin : La Merced, Hacienda Schunke, about 1230 meters, in large
clumps, flowers fragrant in the morning, Macbride 5660 (type).
Chanchamayo Valley, 1200 meters, Schunke 531. Same place and
altitude as the last, Schunke 562, 1807.
FIG. 170. Brassia juninensis C. Schweinf. 1, plant; X %. 2, flower from
front; X 1. 3, lip and column, oblique view; X 2.
843
844 FIELDIANA: BOTANY, VOLUME 30
Brassia Koehlerorum Schltr. Fedde Repert. Beih. 9: 112. 1921;
Mansf. Fedde Repert. Beih. 57: t. 129, nr. 505. 1929.
Plant epiphytic, erect, about 40 cm. high, with a short rhizome. Pseudobulbs
narrowly oblong-cylindric, subcompressed, bifoliate, 15-17 cm. high, surrounded
at the base by short, non-leaf-bearing sheaths. Leaves obliquely oblong-ligulate,
subacute, gradually somewhat narrowed toward the base, 21-24 cm. long, nearly
4-4.5 cm. wide in the middle. Scapes erect-spreading, 20-25 cm. long; peduncle
up to 10 cm. long; raceme rather loosely 10- to 18-flowered, secund. Floral bracts
rather small, deltoid. Flowers small for the genus, with slightly spreading seg-
ments. Dorsal sepal narrowly lanceolate, subacute to subacuminate, about 2 cm.
long and 4.3 mm. wide. Lateral sepals similar, but slightly longer, suboblique,
lightly carinate. Petals obliquely linear-lanceolate, subacute, slightly shorter and
markedly narrower than the sepals. Lip from a subunguiculate-narrowed base
elliptic-lanceolate or "oblong-elliptic," shortly acuminate to a subobtuse tip, about
1.3 cm. long and nearly 5 mm. wide in the middle; disc at the base with a pair of
short, fleshy, minute-papillose keels which are retuse in the middle of each side or
coarsely bicrenate with a pair of rounded, spreading, papillose calli just in front.
Column very short and stout, about 4 mm. high.
Junin: Near La Merced, in the Chanchamayo Valley, Bros.
Koehler s.n.
No specimen of this species was seen.
Brassia longissima (Reichb. f.) Nash, Bailey Stand. Cyclop.
Hort. 1: 541. Mar. 1914. Brassia longissima Schltr. Die Orchid. 496.
Oct. 1914, in part. Brassia Lawrenceana Lindl. var. longissima
Reichb. f. Gard. Chron. (1868) 1313; Hook. f. Bot. Mag. 95: t. 5748.
1869.
Plant large, variable, with a stout creeping rhizome. Pseudobulb strongly
ancipitous, oval or ellipsoid to pyriform-cylindric, unifoliate or less commonly
bifoliate, 5-18 cm. long, surrounded below on each side by 2 or 3 imbricating
sheaths which are successively larger upward, the uppermost one leaf-bearing.
Leaves (on the pseudobulb) elliptic-oblong to oblong, 15-56 cm. long, 4-7 cm. wide,
acute, shortly cuneate at the complicate base. Scape lateral, basal in the axil of
sheaths, shorter than or longer than the leaves, 30-60 cm. high; peduncle with 2
or more remote, short, close, tubular sheaths; raceme rather loosely 4- to 15-flow-
ered, with the rachis 9-24 cm. long. Floral bracts small, triangular-ovate and
amplexicaul or infundibuliform, much shorter than the pedicellate ovary. Flowers
very large, with horizontally spreading segments, yellow-green to orange-yellow
with dark brown spots and lip whitish or cream-color with a few brown spots.
Dorsal sepal linear-lanceolate, elongate-setaceous, 7-12 cm. long, 5-7 mm. wide
at the concave base. Lateral sepals similar but about twice as long or more. Petals
obliquely linear-lanceolate, falcate-incurved, long-acuminate, about half as long
as the dorsal sepal. Lip oblong-lanceolate, long-acuminate, 4-7.6 cm. long, up to
1.5 cm. wide; disc at the base with a pair of approximate, narrow, fleshy, pubes-
cent ridges which commonly terminate in erect tubercles. Column very short and
stout, about 6 mm. high.
SCHWEINFURTH: ORCHIDS OF PERU 845
Amazonas: Habitat not recorded, Woytkowski 23. — Loreto: Pu-
mayacu, between Balsapuerto and Moyobamba, 600-1200 meters,
epiphytic in forest, King 0.1. Also Costa Rica (type) and Panama.
Brassia peruviana Poepp. & Endl. Nov. Gen. ac Sp. 2: 12, t.
117. 1838; Cogn. Martius Fl. Bras. 3, pt. 6: 263. 1905. Oncidium
peruvianum Reichb. f. Walp. Ann. 6: 767. 1863.
Plant large, with a short rhizome. Pseudobulb rather small, subglobose-ovoid,
compressed, unifoliate, 4-5 cm. long, glaucous, sheathed below by 1 or 2 pairs of
conduplicate, imbricating, leaf-bearing sheaths. Leaves large, linear-oblanceolate,
shortly acuminate, long-narrowed below to a more or less petioled, channelled base,
40-60 cm. long, 3-5 cm. wide. Scape lateral, basal, robust, erect, a little shorter
than the leaves, 40-50 cm. tall; peduncle with a few, very remote, rather short,
acute sheaths; raceme loosely several- (about 9-) flowered. Floral bracts trian-
gular-ovate, acute, somewhat ventricose, 2 or 3 times shorter than the pedicellate
ovary. Flowers medium-sized for the genus, subfleshy, with somewhat spreading
segments which are yellowish green spotted with purple. Dorsal sepal linear or
lanceolate-linear, gradually narrowed to an acute apex, 4-4.5 cm. long, 2-2.5 mm.
wide, slightly concave. Lateral sepals similar but somewhat longer, subfalcate.
Petals linear or lanceolate-linear, long-acuminate, subfalcate, about equaling the
dorsal sepal. Lip spreading with the apex lightly recurved, oblong-ovate, acute,
somewhat attenuate below, about 2.5 cm. long, 9-12 mm. wide, with the margin
crisped and minutely crenulate; disc at the base with a pair of short, fleshy, smooth
calli which are approximate below and rather diverging above and 2 minute, re-
mote teeth in front. Column very short and stout, slightly attenuate below,
3-4 mm. high.
Huanuco: District of Chihuamcala, near Cuchero (Cochero), in
rather dry thickets, Poeppig 1589.
No example of this species was available.
Brassia signata Reichb. f. Gard. Chron. n. ser. 16: 6. 1881;
Veitch, Man. Orch. PI. pt. 7, Brassia 126. 1891. Oncidium signatum
Reichb. f. I.e.
Plant medium-sized for the genus, with an abbreviated rhizome. Pseudobulbs
caespitose, oblong-ovoid to ellipsoid, compressed, 7.6-12.7 cm. long, bifoliate or
trifoliate, surrounded at the base by one or more pairs of imbricating, leaf-bearing
sheaths. Leaves oblanceolate-oblong or "cuneate-ligulate," acute or subacute,
slightly and gradually narrowed below, 22.8-30.4 cm. long, about 3.6 cm. wide.
Scapes subequaling or exceeding the leaves, suberect, loosely racemose above, with
2-10 flowers. Flowers rather small for the genus, with widely spreading segments,
the sepals and petals green turning to yellow and lined with brown below, and
the lip white with 2 or 4 purplish spots. Dorsal sepal linear-lanceolate, long-
acuminate, about 5 cm. long and 5 mm. wide. Lateral sepals similar but somewhat
longer, lightly oblique. Petals similar to the lateral sepals but smaller than either
sepal. Lip broadly pandurate, oval-ovate in outline, abruptly long-acuminate,
about 2.5 cm. long; disc at the base with 2 approximate yellow keels which are
846 FIELDI AN A: BOTANY, VOLUME 30
emarginate in the middle of each side. Column very short and stout, about
6 mm. high.
Peru: Habitat not recorded; introduced by Messrs. J. Backhouse
and Sons of York, England.
No specimen of this species was available, but the Ames Her-
barium contains a drawing with floral analysis from the Reichenbach
Herbarium.
Brassia thyrsodes Reichb. f. Gard. Chron. (1868) 842.
Plant large and stout. Pseudobulb pyriform-cylindric, with two remote con-
strictions (one near the middle and one above), apparently bifoliate or trifoliate,
about 12 cm. high. Leaves not described or illustrated. Scape lateral, basal,
robust, suberect, apparently about 53 cm. tall; peduncle with a few small, remote,
appressed sheaths; inflorescence a stout panicle with several spreading-ascending
branches, densely or subdensely many-flowered. Floral bracts triangular-ovate,
acute or acuminate, concave and amplexicaul, widely spreading, about half as long
as the pedicellate ovary or more. Flowers rather small for the genus, yellow with
greenish spots, with widely spreading segments. Dorsal sepal linear-lanceolate,
acuminate, 3.5-4.5 cm. long, about 4 mm. wide. Lateral sepals similar but some-
what longer, oblique. Petals similar to the lateral sepals but shorter than either
sepal. Lip elliptic-oblong or rhombic-ovate, acuminate, angled and often narrowed
at the base, crisped, recurved above, about 3.5 cm. or less long when expanded,
nearly 1.1 cm. wide near the middle; disc at the base with a pair of low lamellae
which are angled at the apex and with a pair of obtusely angled lamellae just in
front. Column very small, about 5 mm. high.
Peru: Habitat not recorded; introduced by Messrs. Veitch and
Sons.
No example of this species was accessible, but the Ames Herbar-
ium contains drawings of the type from the Reichenbach Herbarium.
Brassia villosa Lindl. Fol. Orch. Brassia 4, no. 7. 1853; Cogn.
Martius Fl. Bras. 3, pt. 6: 264. 1905. Brassia hexodonta Reichb. f.
& Warsc. Bonpl. 2: 98. 1854. Oncidium villosum Reichb. f. Walp.
Ann. 6: 767. 1863.
Pseudobulbs, leaves and lower parts of scape lacking. Raceme densely or sub-
densely many-flowered, secund, about 23 cm. or more long. Floral bracts broadly
triangular-ovate, acute, amplexicaul, appressed, 3 or 4 times shorter than the pe-
dicellate ovary. Flowers small, fragrant, with rather fleshy, spreading segments.
Dorsal sepal oblong-lanceolate, long-acuminate, 2 cm. long, 3 mm. wide. Lateral
sepals similar but considerably longer, subfalcate. Petals narrowly lanceolate,
acuminate, strongly arcuate-falcate, markedly shorter and slightly wider than the
dorsal sepal. Lip sessile, spreading-reflexed, ovate-oval, abruptly acute or apicu-
late, undulate on the margin and sometimes lightly lobulate, subrounded at the
base, about 1.5 cm. long, 7-10 mm. wide; disc at the base with 6 short, pubescent
calli, the 2 posterior calli oblique and somewhat confluent above, the 2 median
SCHWEINFURTH: ORCHIDS OF PERU 847
calli subapproximate and central, and the 2 anterior calli spreading and semiorbic-
ular. Column short, stout, clavate, acutely angled on each side at the apex, about
5 mm. high.
Peru: Habitat not designated (type of B. hexodontaJ—Hubnuco:
Source of the Maranon River, Warscewicz s.n. (type of B. mllosa).
Also British Guiana.
No specimens of this concept were available, but the Ames Her-
barium contains a photograph with floral analysis of typical B. ml-
losa from the Lindley Herbarium as well as a drawing of B. hexodonta
from the Reichenbach Herbarium.
MILTONIA Lindl.
A rather small genus of tropical American epiphytes, extending
from Costa Rica and Panama through Colombia, Ecuador and Peru
to Paraguay and Argentina, and centering in Brazil.
Plant commonly large, rarely small. Rhizome abbreviated or long-creeping.
Pseudobulbs often aggregated, 1- to 2-leaved, surrounded below by one or more
pairs of distichous, often leaf-bearing sheaths. Leaves linear or oblong, narrowed
below. Scapes lateral, basal in the axil of sheaths, shorter or often longer than the
leaves, loosely 1- to many-flowered above, racemose or rarely paniculate. Flowers
small, medium-sized or very large, with widely spreading segments. Sepals sub-
equal, free or the lateral ones shortly connate at the base. Petals similar to the
sepals or often broader. Lip spreading from the base of the column, sessile or
shortly clawed, simple or pandurate; disc near the base 1- to several-lamellate.
Column short or very short, footless, often with a cucullate clinandrium. Anther
terminal, opercular, incumbent. Pollinia 2, ovoid, unappendaged.
Al Scape much shorter than the leaves; flowers small for the genus, lip 1.5 cm.
or less long M- Parva
A2. Scape as long as the leaves or longer; flowers relatively large; lip about 2.7 cm.
or more long *
la. Inflorescence commonly branched; sepals and petals crisped-undulate, spatu-
late-oblong, 7.5 mm. or less wide; lip not auricled at the base.
M. Warscewiczii
Ib. Inflorescence always simple; sepals and petals flat, obovate-oblong, 10 mm.
or more wide; lip with a pair of triangular-ovate auricles at the base.
M. vexillaria
Miltonia parva C. Schweinf. Am. Orch. Soc. Bull. 14: 294, t.,
1945. Figure 171.
Plant small and slender for the genus. Rhizome creeping, slender, concealed
by loose, concave, amplexicaul sheaths. Pseudobulbs cylindric, strongly com-
pressed, unifoliate, 3.5-4.5 cm. long, surrounded at the base by 2 distichous, imbri-
cating sheaths, the larger upper one commonly bearing a small leaf-blade. Leaf
(on the pseudobulb) oblong-linear to elliptic-linear, sharply acute, gradually nar-
848 FIELDIANA: BOTANY, VOLUME 30
rowed below to a conduplicate, petiole-like base, 17-23.5 cm. long, 1-1.5 cm. wide.
Scape lateral, basal, very short, with a loosely 2-flowered raceme, about 5 cm. or
less tall. Flowers small for the genus, with widely spreading segments. Dorsal
sepal elliptic-oblong, abruptly acute, dorsally subcarinate, 1.68-1.8 cm. long, 5.2-
7 mm. wide. Lateral sepals very similar in shape and size, dorsally carinate.
Petals similar to the sepals but markedly smaller. Lip spreading from the base,
simple, ovate-suborbicular or suborbicular, lightly retuse to broadly rounded in
front, crenulate-erose and undulate except near the base, about 1.2-1.5 cm. long
and 1.7-1.8 cm. wide; disc with a single more or less distinct, oblong, fleshy callus
in the center of the base. Column small, lightly arcuate, terminating in a cucullate
clinandrium, 8.9-9.5 mm. high at the back, with a pair of porrect, triangular teeth
near the base.
Junin: Chanchamayo Valley, 1200 meters, Schunke N. (type).
Same locality, 1500 meters, Schunke 1272, 1295.
Miltonia vexillaria (Reichb. f.) Nichols. Diet. Gard. 2: 369
(1884-1888); Veitch, Man. Orch. PI. 8: 110. 1892; Cogn. & Goos.
Diet. Icon. Orch. Miltonia t. 1. 1896. Odontoglossum vexillarium
Reichb. f. Gard. Chron. (1867) 901; I.e. (1872) 667, fig. 163; Hook. f.
Bot. Mag. 99: t. 6037. 1873.
Plant medium-sized to large, apparently with an abbreviated rhizome. Pseudo-
bulbs oblong-ellipsoid, strongly compressed, about 4-6.3 cm. high, 1- or 2-leaved
at the summit, surrounded at the base by 2 or 3 distichous, imbricating, leaf-bear-
ing sheaths. Leaves narrowly elliptic-lanceolate to linear-lanceolate, acute, about
15-30 cm. long, 2.5-3.7 cm. wide. Scapes lateral, basal in the axil of sheaths,
usually 2 or 3 (rarely 6) to each plant, slender, arcuate, commonly longer than the
leaves, up to 50 cm. long, very loosely racemose and 2- to 7-flowered. Flowers
large and showy, very variable in size and color, usually pale rose-color (less com-
monly white or rose-carmine). Sepals similar, ovate-oblong, subacute to truncate,
recurved above, white on the margins, about 2-3 cm. long and 1-1.4 cm. wide, the
lateral sepals lightly oblique. Petals similar to the sepals but commonly broader.
Lip much larger than the other segments, round-obovate to suborbicular, promi-
nently bilobed in front, extended at the base into a pair of small, erect, triangular-
ovate auricles, about 2.7-5.4 cm. long (on each side of the middle) and 2.4-6.4 cm.
wide; disc at the yellow base with a minute, bilobed callus and 3 obscure teeth in
front. Column abbreviated, clavate, wingless.
Cuzco: Prov. of Paucartambo, from Sta. Isabel to Asuncion, 1800
meters, epiphyte, Vargas 5525. Also Colombia (type) and Ecuador.
Miltonia Warscewiczii Reichb. f. Xen. Orch. 1: 129, 132. 1856;
Hook. f. Bot. Mag. 96: t. 5843. 1870; Cogn. & Goos. Diet. Icon.
Orch. Miltonia t. 8. 1900; Cogn. Martius Fl. Bras. 3, pt. 6: 283. 1905.
Oncidium fuscat um Reichb. f. Walp. Ann. 6: 763. 1863. Odontoglos-
sum Weltonii Hort. ex Gard. Chron. (1870) 1: 417. Oncidium Wel-
tonii Hort. ex Gartenfl. 25: 339. 1876.
MILXONIA
jo ar i)a
C. Ocfiiweinji.
FIG. 171. Miltonia parva C. Schweinf. 1, plant; X 1. 2, dorsal sepal; X 2.
3, petal; X 2. 4, lateral sepal; X 2. 5, column from side; X 3. 6, lip spread out;
X 2.
849
850 FIELDIANA: BOTANY, VOLUME 30
Plant large, epiphytic. Rhizome apparently abbreviated. Pseudobulbs con-
gested, strongly compressed, oblong-cylindric (sometimes slightly narrowed up-
ward), unifoliate, 8-12.7 cm. long, surrounded below by 1 or 2 pairs of conduplicate,
leaf-bearing sheaths. Leaves oblong, rounded or abruptly acute at the apex, grad-
ually narrowed below to a conduplicate base, up to 33 cm. long (on the pseudobulb),
about 3 cm. or more wide. Scape lateral, basal in the axil of a sheath, surpassing
the leaf, suberect to arcuate, 25-54 cm. long; peduncle about 41 cm. or less long,
entirely or mostly concealed by numerous, close, tubular sheaths; inflorescence
simple or more commonly loosely branched, laxly to subdensely many-flowered.
Floral bracts lanceolate, concave, appressed, about half as long as the slender
pedicellate ovary. Flowers several to numerous, with widely spreading segments;
sepals and petals crisped-undulate on the margins, reddish brown with white or
yellowish apex; lip rose-purple, variegated with white, yellowish and reddish-
brown. Dorsal sepal oblanceolate-oblong or spatulate-oblong, obtuse or retuse
and mucronate, 1.7-2.1 cm. long, up to 7.5 mm. wide. Lateral sepals similar but
a little longer and narrower, obtuse, oblique and lightly recurved, dorsally carinate.
Petals very similar to the dorsal sepal but less narrowed below, oblanceolate-oblong,
obtuse or retuse. Lip variable, much larger than the other segments, spreading
from the base, oval-suborbicular or suborbicular-obovate (sometimes slightly quad-
rangular), deeply bilobed at the rounded or subtruncate apex, up to 2.8 cm. long
(from the base to the apex of a lobule) and 2.5 cm. wide; disc naked or with an
obscure, fleshy, bilobed thickening at the very base. Column very short and stout,
4-5 mm. high at the back, with a pair of rounded, inconspicuous wings.
Huanuco: Cuchero (Cochero), on trees, Poeppig 1765, Warsce-
wicz s.n. (type). Also Colombia, and Ecuador, fide Schlechter, Fedde
Repert. Beih. 8: 155. 1921.
SOLENIDIUM Lindl.
A very small genus rather closely allied to Oncidium, at present
limited to four species, confined to the northern half of South Amer-
ica. A single species is known in Peru.
Plants epiphytic, medium-sized to rather large, with an abbreviated rhizome.
Pseudobulbs aggregated, ovoid or oblong-ovoid to ellipsoid, 1- to 2-leaved at the
apex. Leaves oblong or linear-oblong. Inflorescence lateral, basal, laxly or rarely
subdensely racemose above. Flowers medium-sized, long-pedicelled. Sepals sub-
equal, free, wide-spreading. Petals similar to the sepals. Lip spreading from the
base of the column, simple, with a narrow claw-like base, dilated above; disc with a
pair of calli or lamellae. Column erect, footless, broadly 2-winged, the wings dilated
into auricles above. Anther opercular, incumbent, galeate, 1-celled. Pollinia 2,
ovoid, unappendaged.
Solenidium peruvianum Schltr. Fedde Repert. Beih. 9: 113.
1921; Mansf. Fedde Repert. Beih. 57: t. 129, nr. 507. 1929.
Plant medium-sized, erect, about 20 cm. high. Pseudobulbs aggregated, [ellip-
soid], compressed, unifoliate, about 2 cm. long, below with a leaf-bearing sheath on
SCHWEINFURTH: ORCHIDS OF PERU 851
each side. Leaf erect, ligulate-linear, subobtuse, up to 15 cm. long and 1 cm. wide.
Inflorescence erect, about 18 cm. high; peduncle subcompressed, about 12 cm. long;
raceme about 7 cm. long, loosely several- (about 6-) flowered. Flowers rather
small and subfleshy, with wide-spreading segments. Dorsal sepal oblong-obovate,
broadly obtuse or rounded at the apex, about 11 mm. long. Lateral sepals sim-
ilar but a little larger, oblique, acute or apiculate. Petals similar to the sepals but
a little shorter, elliptic-obovate, oblique, obtuse. Lip somewhat shorter than the
other segments, broadly obovate in outline, from a quadrate-unguiculate basal
portion with erect sides, suborbicular-dilated, apiculate, about 9 mm. long and
6 mm. wide near the middle; disc from the base to beyond the middle with a pair
of approximate, fleshy, minute-papillose, linear-oblong calli. Column short, stout,
about 6 mm. high, terminating in a pair of erect, falcate-oblong, obtuse auricles.
Loreto: Near Moyobamba, Filomeno s.n.
No example of this species has been available.
ONCIDIUM Sw.
A very large genus of New World epiphytes extending from south-
ern Florida (U.S.A.), the West Indies and Mexico through Central
America and South America to Brazil and Argentina.
Plants very variable, small and insignificant to large and showy, with a creep-
ing or abbreviated rhizome, either terrestrial or epiphytic. Pseudobulbs short and
suborbicular to elongate and narrowly cylindric (sometimes minute or lacking),
more or less compressed, usually 1- to 3-leaved, commonly surrounded below by
one or more pairs of frequently leaf-bearing sheaths. Leaves variable, rather flat,
more rarely ancipitous and equitant or terete, commonly elongate. Inflorescence
lateral, basal in the axil of sheaths, above racemose or paniculate, loose or dense,
few- to many-flowered, erect, frequently long-climbing or twining. Flowers com-
monly showy and usually yellow, often spotted with brown, with spreading or
reflexed segments. Sepals commonly subequal, the lateral sepals free or more or
less connate. Petals similar to the dorsal sepal, commonly smaller or rarely larger.
Lip spreading from the base of the column, sessile or with a short claw, usually
pandurate or 3-lobed, the mid-lobe being often much larger, transverse and emargi-
nate, rarely small, narrow and entire; disc with a commonly tuberculate callus at
the base. Column short, stout, footless, usually with a petaloid wing at the apex
on each side of the stigmatic orifice, sometimes wingless, often with a conspicuous
swelling in front. Anther terminal, opercular, incumbent. Pollinia 2, obovoid,
unappendaged, affixed to a stipe.
Al. Lip distinctly narrower toward the apex, very rarely subpandurate above ... I
A2. Lip distinctly broader toward the apex, or pandurate 25
la. Flowers relatively small, lateral sepals 1.5 cm. or less long 1
Ib. Flowers medium-sized to large, lateral sepals 2 cm. or more long 9
la. Column with conspicuous, well-developed wings above 2
Ib. Column without conspicuous or well-developed wings 5
2a. Lip cuneate-obovate or cuneate-rhombic 0. acinaceum
2b. Lip broadly triangular (i.e., much broader near the base) 3
852 FIELDIANA: BOTANY, VOLUME 30
3a. Crest of the lip with a pair of high keels, higher in front; lateral sepals rounded
or retuse at the apex O. xanihornis
3b. Crest of the lip without a pair of prominent carinate keels, but consisting of
numerous short papillae or tubercles 4
4a. Panicle not climbing, at most nodding above, with many-flowered branches;
flowers of one form; column- wings semiovate, entire, acute O. deltoideum
4b. Panicle climbing, with few-flowered, short branches; flowers of two kinds;
column-wings semiorbicular, lobulate, obtuse O. ionopterum
5a. Floral bracts prominent, although much shorter than the ovary 6
5b. Floral bracts minute 7
6a. Scape lax, climbing, with very loosely flowered branches; lip triangular in
outline O. cimiciferum
6b. Scape erect or suberect with densely flowered branches; lip ovate in outline.
O. ovatilabium
7a. Lip ovate in outline, bicarinate below with the keels attenuated in front;
ultimate branchlets of the panicle rather densely flowered 0. mentigerum
7b. Lip triangular in outline, with numerous warts which are more or less fused
at the base; ultimate branchlets of the panicle loose or very loose 8
8a. Petals distinctly broader than the dorsal sepal; lip with more or less protu-
berant basal angles; column with prominent angles below O. trulla
8b. Petals scarcely broader than the sepals; lip without protuberant basal angles;
column without prominent angles below O. cocciferum
9a. Petals usually nearly or quite as large as the dorsal sepal, broadly rounded
or at least strongly obtuse at the apex 10
9b. Petals definitely smaller than the dorsal sepal, commonly narrowed toward
the apex 12
lOa. Lip acute to subacute; lateral lobes or auricles not retrorse.
0. superbiens (O. aemulum)
lOb. Lip long-acuminate; lateral lobes strongly retrorse 11
11 a. Crest of the lip consisting of 3 equal, retrorse, ancipitous lamellae with 3
smaller lamellae in front; wings of the column broad, rounded.
O. macranthum
lib. Crest of the lip consisting of 1 large median keel, flanked by a smaller pros-
trate lamella on each side; wings of the column relatively narrow, ligulate.
O. macranthum var. hastiferum
12a. Lip with linear or linear-oblanceolate, erect lateral lobes 0. minax
12b. Lip with broadly triangular or auriculate lateral lobes, more or less distinct
dilations or altogether simple 13
13a. Dorsal sepal obtuse to broadly rounded above 14
13b. Dorsal sepal commonly narrowed above 18
14a. Floral segments finely serrate-denticulate to fimbriate; lamina of the dorsal
sepal much broader than long O. serratum
14b. Floral segments not denticulate 15
15a. Dorsal sepal long-clawed, spatulate-oblong; lip simple O. incarum1
15b. Dorsal sepal shortly and abruptly clawed; lip more or less 3-lobed 16
1 No material of this species was seen, and so its position in the key is somewhat
uncertain.
SCHWEINFURTH: ORCHIDS OF PERU 853
Claws of the sepals without basal auricles O. Leopoldianum
Claws of the sepals auricled at the base 17
Lateral lobes at the base of the lip acute; petals commonly conduplicate and
oblique or falcate O. faldpetalum (O. ionodon, O. Davisii )
Lateral lobes at the base of the lip rounded; petals flat, not falcate.
O. superbiens (0. aemulum)
Lip with a pair of narrow, lanceolate, retrorse, basal lobes having a filiform
tooth at their base; crest consisting of 5 short lamellae, the central 3 bilobed.
O. cordatum
Lip without narrow, retrorse, basal lobes with a filiform tooth at their base ;
crest not consisting of 5 short lamellae 19
Mid-lobe of the lip conspicuously crisped-undulate; lateral lobes prominently
subquadrate-auriculate and often coarsely toothed 20
Mid-lobe of the lip flat or with involute margins; lateral lobes rounded, tri-
angular, bilobed or sometimes indistinct 21
Bracts abbreviated, not spathaceous; basal callus of the lip consisting of 3
oblong keels, the central one higher O. Kienastianum
Bracts large, spathaceous; anterior part of the callus consisting of 3 ovate or
ovate-oblong lamellae, the central one largest 0. trilingue
Lateral lobes of the lip rounded; wings of the column rounded; sepals and
petals white with a large, purple central portion O. Leopoldianum
Lateral lobes of the lip (if distinct) triangular or hastate (rarely bilobed);
wings of the column linear or dentiform 22
Claws of the sepals without basal auricles 23
Claws of the sepals with basal auricles, the dorsal sepal prominently au-
ricled 24
Lamina of the sepals elliptic-lanceolate, long-narrowed below; sepals and
petals white with transverse, red- violet or brownish bars 0. zebrinum
Lamina of the sepals ovate or oval-ovate to suborbicular, at least the dorsal
sepal abruptly narrowed below; sepals and petals cinnamon-brown with a
yellow upper half to the petals O. ventilabrum (O, undulatum)
Lateral lobes of the lip obtuse; mid-lobe obtuse, dilated above; disc with 2
central keels O. gargantua
Lateral lobes of the lip more or less acute; mid-lobe acute, not much (if at
all) dilated above; disc with a 3-ridged, central keel.
O. faldpetalum (0. ionodon, O. Davisii)
Leaves equitant, imbricating, entirely concealing the stem; plants very small,
flabelliform 26
Leaves flat, not equitant (occasionally the leaves are conduplicate or they
are unknown) ; plants commonly medium-sized to large 27
Crest at the base of the lip with 3 or 4 laciniate or fimbriate lobes.
O. glossomystax
26b. Crest at the base of the lip with entire lobes 0. pusillum
27a. Flowers of two kinds, one fertile and relatively large, the other abortive and
small 28
27b. Flowers uniform, fertile 33
28a. Anterior part of the lip much larger and commonly much broader than the
basal part 29
28b. Anterior part of the lip subequaling the basal part or smaller 30
854 FIELDIANA: BOTANY, VOLUME 30
29a. Lip acute at the broadly rounded apex; sepals and petals with the basal half
purple and the upper half yellow; petals acute or subacute, nearly four times
as large as the sepals O.formosissimum1
29b. Lip prominently bilobed; sepals and petals wholly yellow; petals obtuse,
somewhat larger than the sepals O. calanthum
30a. Claw between the basal and the anterior portions of the lip very short or
wanting 31
30b. Claw between the basal and the anterior portions of the lip rather prom-
inent 32
31a. Basal part of the lip semiorbicular, with rounded angles on each side.
O. heteranthum
31b. Basal part of the lip rhombic, with acute angles on each side.
O. tigratum1
32a. Petals obovate-oblong or spatulate-oblong, more or less retuse at the broad
apex; lobes at the base of the lip spreading O. retusum
32b. Petals lanceolate-oblong, commonly, .acute; lobes at the base of the lip
retrorse 0. pentadactylon
33a. Lateral sepals connate up to the middle or above 34
33b. Lateral sepals free or only slightly connate at the base 37
34a. Lip with a definite claw at the base; wings of the column fimbriate.
O. aureum (Odontoglossum festatum, Odontoglossum hemichrysum,
Oncidium hemichrysum)
34b. Lip sessile (without a claw) 35
35a. Sepals and petals clawed, obtuse; lip with a 5-lamellate callus at the base.
O. Lucasianum*
35b. Sepals and petals sessile (not clawed), acute or subacuminate 36
36a. Apical lobe of the lip reniform, much wider than the basal portion; lobes of
the lateral sepals not diverging; flowers brown, white and purple.
0. nubigenum (O. cucullatum)
36b. Apical lobe of the lip broadly obovate or obovate-reniform, not much wider
than the basal portion; lobes of the lateral sepals diverging like a swallow's
tail; flowers yellow and brown O. tripterygium
37a. Stem (rhizome) and inflorescence long-creeping or climbing 38
37b. Stem or rhizome (if present) not long-creeping or climbing; inflorescence erect
to arcuate or deflexed, not long-creeping or climbing 47
38a. Inflorescence or scape abbreviated, 1- to 3-flowered 39
38b. Inflorescence or scape elongate, racemose or usually paniculate, commonly
several- (10) to many-flowered 40
39a. Lip simple, oblong-pandurate, with the anterior part obcordate and scarcely
wider than the basal part; wings of the column wanting O. serpens
39b. Lip truly 3-lobed, with the anterior part reniform and much wider than the
basal part; wings of the column evident, dolabriform.
O. globuliferum (0. convolvulaceum)
1 The original descriptions, as well as the records of these species from the
Reichenbach Herbarium, fail to show flowers of two kinds, but Kranzlin mentions
such in Engler, Pflanzenr. IV. 50 (Heft 80), 1922, so they are entered in both
branches of the key.
1 O. Lucasianum sometimes has the lateral sepals free, and so is keyed on the
other branch also.
SCHWEINFURTH: ORCHIDS OF PERU 855
Sepals oblong (the lamina with subparallel sides); dorsal sepal sessile (only
narrowed below); lip equaling or exceeding the sepals in length 41
Sepals not oblong (at least without parallel sides), the lamina commonly
ovate-elliptic to suborbicular or reniform; dorsal sepal long-clawed or
abruptly clawed; lip markedly smaller than the sepals 42
Pseudobulbs 1-leaved, remote (up to 20 cm. apart) ; inflorescence not climb-
ing, subdensely racemose above; sepals and petals obtuse.
O. Weberbauerianum
Pseudobulbs 2-leaved, only subremote; inflorescence climbing, very loosely
paniculate; sepals acute (or apiculate) to acuminate 0. Schillerianum
Apex of the lip broadly rounded to bilobed 43
Apex of the lip acute to apiculate or obtuse 45
Claw of the dorsal sepal not auricled at the base; lip broadly rounded in
front; column-wings none O. volubile (0. corynephorum)
Claw of the dorsal sepal auriculate at the base; lip more or less bilobed;
column- wings present 44
Lamina of the dorsal sepal oval-suborbicular, broadly rounded above, flat
or nearly so; column- wings minute, cornute or falcate-ascending.
0. diceratum
Lamina of the dorsal sepal deltoid-ovate, acute, prominently crisped-undu-
late; column- wings cuneate and retuse O. cryptocopis
Lamina of the dorsal sepal reniform or semiorbicular, much broader than
long, broadly rounded above O. serratum
Lamina of the dorsal sepal "oblong" or broadly ovate, longer than broad,
obtuse or acute 46
Petals without a creamy white or yellow upper half, rounded or subhastate
at the base; mid-lobe of the lip cuneate-spatulate, often denticulate-crisped
toward the apex O. xanthodon1
Petals with a creamy white or yellow upper half, cuneate at the base; mid-
lobe of the lip oblong-lanceolate or elliptic-lanceolate, not denticulate-crisped
toward the apex O. ventilabrum (O. undulatum)
Flowers very large; dorsal sepal and petals oblanceolate-linear, 5.5 cm. or
more long 48
Flowers medium-sized to small; dorsal sepal and petals 2 cm. or less long. . . 49
Column- wings on each side bilobed, the upper lobe linear-subulate and
glandular, and the lower lobe broad and somewhat lacerate 0. Papilio
Column-wings on each side uniformly pectinate with the cilia glandular-
capitate O. Sanderae
Plant very small, about 10 cm. or less tall; scapes 1- to few-flowered; mid-
lobe of the lip conspicuously 4-lobed O. crista-galli
Plant medium-sized to very large, about 13 (rarely 9) cm. or more tall; scape
usually many-flowered; mid-lobe of the lip simple (rarely) to retuse or bi-
lobed 50
Mid-lobe of the lip broadly rounded in front and apiculate in the middle ... 51
Mid-lobe of the lip lightly retuse to bilobed in front 52
Sepals shorter than the lip; wings of the column very large. .O.formosissimum
Sepals longer than the lip; wings of the column very small or wanting.
O. ochmatochilum
1 This species, of which no material has been seen, may prove to be referable
to O. ventilabrum.
856 FIELDIANA: BOTANY, VOLUME 30
52a. Petals definitely broader than the lateral sepals, often twice as broad or
more 53
52b. Petals at most slightly broader than the lateral sepals or narrower 66
53a. Basal half of the lip orbicular in outline, with numerous, free teeth or tuber-
cles surrounding the central callus O. polyadenium
53b. Basal half of the lip not orbicular in outline, commonly without numerous,
free teeth surrounding the central callus 54
54a. Anterior portion of the lip (mid-lobe) sessile, i.e., without a distinct claw. . . 55
54b. Anterior portion of the lip (mid-lobe) with a claw, even if short 56
55a. Leaves small (commonly conduplicate), up to 12.7 cm. long and 2.5 cm. wide
(when expanded) ; basal portion of the lip transversely oblong, equally wide
with the anterior part O. onustum
55b. Leaves large, up to 40 cm. long and 4 cm. wide; basal portion of the lip sub-
quadrate, much narrower than the anterior part O. cajamarcae
56a. Basal half of the lip rhombic, often acute on each side; claw between the
basal and anterior portions of the lip short or very short 57
56b. Basal half of the lip not rhombic, more or less rounded or auriculate on
each side 58
57a. Column-wings semilunate or bifid, forming 2 ligulate segments. .O. tigratum
57b. Column-wings simple, dolabrif orm-semicordate O. fasciferum
58a. Column without wings; claw between the basal and anterior portions of the
lip oblong-linear; sepals and petals strongly repand O. tipuloides
58b. Column winged on each side; claw between the basal and anterior portions
of the lip not oblong-linear 59
59a. Lateral (basal) lobes of the lip minute, auriculate; anterior portion of the
lip many times broader than the basal portion; sepals cucullate above.
O. ampliatum
59b. Lateral lobes of the lip rather prominent; anterior portion of the lip not
many times broader than the basal portion; sepals not cucullate 60
60a. Lip about twice as long as the lateral sepals; lamina with one or more free
tubercles surrounding the middle callus; flower-cluster a short, rather densely
flowered, pyramidal panicle O. Batemanianum
60b. Lip subequaling or somewhat surpassing the lateral sepals; flower-cluster an
elongate, commonly loosely flowered panicle 61
61a. Rostellum long-beaked; wings of the column from a narrow base divergently
bilobed O. pyramidale (O. chrysopyramis)
61b. Rostellum not long-beaked; wings of the column with a broad base 62
62a. Basal portion of the lip when expanded noticeably broader than the anterior
portion (mid-lobe); mid-lobe cuneate, lightly retuse O. Mathieuanum
62b. Basal portion of the lip equaling or narrower than the anterior portion; mid-
lobe transverse, renif orm, emarginate to bilobed 63
63a. Sepals oblong, elliptic-lanceolate or oblong-ovate, narrowed above 64
63b. Sepals narrowly obovate or cuneate-spatulate, i.e., broader above 65
64a. Petals elliptic-oblong, sessile, acute; column- wings small, unequally bilobed.
O. hieroglyphicum
64b. Petals oblong-ovate, shortly clawed, broadly obtuse and apiculate; column-
wings large, lightly 3-lobed, falcate above , O. leopardinum
65a. Petals not retuse; claw between the basal and apical portions of the lip more
or less elongate; column- wings long-acute above, falcate O. obryzatum
SCHWEINFURTH: ORCHIDS OF PERU 857
65b Petals often prominently retuse; claw between the basal and apical portions
of the lip very short; column-wings subquadrate-rounded or subdolabnform,
often retuse in the middle 0. excavatum
66a. Column-wings narrow, dentiform to subulate 67
66b. Column-wings relatively broad, neither dentiform nor subulate 68
67a Plant small, pseudobulbs 7 mm. or less long, unifoliate; panicle dense; sepals
about 1 cm. or less long, broadly obtuse to rounded above O. nanum
67b. Plant large, pseudobulbs 7.5 cm. or more long, bifoliate; panicle loose; sepals
about 1.3 cm. long, acute or subacute O. piclum
68a Basal part of the lip (across the outspread lateral lobes) distinctly wider than
' the anterior part (mid-lobe) O. granlhmum
68b. Basal part of the lip (across the lateral lobes) distinctly narrower than the
anterior part (rarely subequal) 6a
69a. Sepals obtuse to rounded at the apex (sometimes obscurely apiculate) .... 70
69b. Sepals acute to acuminate "4
70a Lip oblong, obovate-oblong or subquadrate-oblong in outline, commonly
about twice as long as broad; lamina with an upcurved, 4-fingered callus at
the base °- hians
70b Lip not oblong in outline, shorter than broad (rarely slightly longer, m
0. sessile and O. pardothyrsus); lamina without an upcurved, 4-fingered
callus 71
71a. Disc of the lip with a 5-lamellate callus surrounded by warts . .0. Lucasianum
71b. Disc of the lip without a 5-lamellate callus surrounded by warts 72
72a Sepals and petals sessile (without any claws), oblong or elliptic-oblong.
O. sessile
72b. Sepals definitely clawed, the lamina broadly ovate or oval to rounded. . .73
73a Lip distinctly shorter than the sepals; anterior portion of the lip but little
wider than the basal portion; sepals and petals more or less undulate.
O. pardothyrsus
73b Lip equaling or surpassing the sepals; anterior portion of the lip more than
twice as wide as the basal portion; sepals and petals strongly undulate.
0. lundum
74a Lip produced on each side at the base into a conspicuous, rounded or oblong-
rounded, spreading or retrorse auricle or lobe; claw between the basal and
anterior portions prominent and elongate 75
74b Lip not produced into conspicuous auricles at the base, but with a more or
less rounded or rhombic basal portion; claw between the basal and anterior
portions short or wanting ' '
75a Basal central part of the lip with a series of sulcate lines below the callus;
column-wings simply and unequally bilobed O. hneohgerum
75b. Basal central part of the lip without a series of sulcate lines; column-wings
not bilobed 76
76a Callus at the base of the lip cruciform with broadly ligulate, retuse-dentate
lateral lobes and an elongate, linear-cuneate anterior lobe. . .0. cruciferum1
76b. Callus at the base of the lip oval or subquadrate in outline, with subequal
fleshy tubercles in three series 0. Bauen
i This species, which was described from a portion of an inflorescence only,
appears to be very close to the widespread O. Bauen Lmdl., and subsequent col-
lections may prove that these species are synonymous.
858 FIELDIANA: BOTANY, VOLUME 30
77a. Lateral sepals nearly twice as long as the lip; inflorescence a narrow, open,
virgate panicle O. hirundo
77b. Lateral sepals shorter than the lip (rarely subequaling the lip) ; inflorescence
a pyramidal, often dense panicle 78
78a. Anterior portion of the lip scarcely wider than the basal portion; callus with
2 tubercles on each side 0. discobulbon
78b. Anterior portion of the lip markedly wider than the basal portion; callus with
4 or more free papillae on each side O. armillare (O. foveatum)
Oncidium acinaceum Lindl. Sert. Orch. sub t. 48, no. 13. 1842;
Fol. Orch. Oncidium 18, no. 52. 1855; Kranzl. Engler, Pflanzenr. IV.
50 (Heft 80), 180, fig. 15G, a-d. 1922.
Plant large, up to 60 cm. high when extended. Pseudobulbs oval or ovoid,
densely crowded according to Kranzlin, I.e., subcompressed, about 6 cm. high,
2- or 3-leaved at the summit, surrounded at the base by 2 or 3 distichous, leaf-
bearing sheaths. Leaves oblanceolate or oblong with a narrowed, complicate base,
acute to obtuse, 8-13 cm. long, 1.5-3 cm. wide. Scape lateral, in the axil of an
upper sheath, up to over 60 cm. in length, climbing above, loosely flowered, ra-
cemose, or paniculate with short, fractiflex branches (according to Kranzlin).
Flowers uniform (according to Lindley), dimorphic (according to Kranzlin). Sterile
flowers very minute, with lanceolate sepals, oblong petals and lip about 5 mm.
long, very short-pedicelled. Fertile flowers relatively large, rose-lilac, long-pedi-
celled; dorsal sepal oblong or oblong-linear, obtuse, about 1.5 cm. long, 3-4 mm.
wide. Lateral sepals spatulate-linear, connate through the lower fourth, as long
as the dorsal sepal. Petals obovate, obtuse, as long as the sepals but about three
times broader. Lip obovate-rhombic, broadly obtuse to retuse, indistinctly tri-
lobulate above, about 1.4 cm. long and 1 cm. wide above; disc with 4 approximate,
longitudinal keels (hamate at the apex), the outer pair being shorter. Column
much shorter than the other segments, lightly recurved, 8-9 mm. high, with a pair
of large acinaciform wings above.
Amazonas: Chachapoyas, on trees, Mathews 3186 (type). — Caja-
marca: Prov. of Chota, mountains west of Huambos, 2900-3000
meters, frequent in sclerophyllous vegetation, Weberbauer 1^173. Also
Colombia (Antioquia, fide Kranzl.).
No example of this species was available, but the Ames Her-
barium contains a photograph of the Mathews type in the Lindley
Herbarium.
Oncidium ampliatum Lindl. Gen. & Sp. Orch. 202. 1833; Bot.
Reg. 20: 1. 1699. 1835; Fol. Orch. Oncidium 28, no. 90. 1835; Kranzl.
Engler, Pflanzenr. IV. 50 (Heft 80), 230. 1922. Oncidium Bernoul-
lianum Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 231. 1922.
Figure 172.
Plant usually large, variable. Rhizome abbreviated. Pseudobulbs orbicular
to broadly ovoid or pyriform, strongly compressed and often subtetragonous, fre-
quently crowded, bearing 1 or 2 leaves, 3-7.5 (10 fide Kranzl.) cm. long, surrounded
SCHWEINFURTH: ORCHIDS OF PERU
859
FIG. 172. Oncidium ampliatum Lindl. Plant; X 1.
at the base by 1 or more pairs of evanescent sheaths. Leaves elliptic-oblong, oblan-
ceolate-oblong or obovate-oblong (more rarely elliptic), obtuse to broadly rounded
(rarely subacute) at the tip, more or less narrowed to a complicate or conduphcate
base 8 (rarely) -51 cm. long, 2.8-9.2 cm. wide. Scape much exceeding the leaves,
suberect or spreading from a decumbent base, up to 83.5 cm. or more tall; peduncle
with several (11 or less) remote, short sheaths which are close and tubular; inflores-
cence racemose or more commonly paniculate, usually subdensely many-flowered.
Floral bracts very small, squamiform. Flowers long-pedicelled with spreading seg-
ments yellow with more or less minute brown spots within; whitish without. Sepals
more or less distinctly clawed. Dorsal sepal spatulate-obovate, concave above,
broadly rounded at the apex, 7-8.8 mm. long, 5.6-8.3 mm. wide when expanded.
860 FIELDIANA: BOTANY, VOLUME 30
Lateral sepals free, similar to the dorsal sepal but a little longer and narrower,
oblique. Petals obovate-suborbicular, more or less truncate above, clawed below,
somewhat longer than the lateral sepals and much broader than the dorsal sepal.
Lip deeply 3-lobed, 13.3-23.5 mm. long from the base to either side of the retuse
apex; lateral lobes minute, auriculiform; mid-lobe (separated by a short claw) sub-
quadrate-reniform, retuse or bilobed in front, cordate at the base, about 16-33 mm.
wide, with irregular or lobulate sides; disc at the base with a small, fleshy callus of
2 series, the basal part of 2 larger, spreading lobes and the anterior part of 3 small
lobes. Column very short, stout, 2.3-3.9 mm. high at the back, with a pair of
small aliform, denticulate or lobulate wings.
Peru: Without locality, Warscewicz s.n., fide Kranzl. Also Cen-
tral America from Guatemala (type of 0. Bernoullianum) to Panama
and Colombia (type of 0. ampliatum), Venezuela and Trinidad.
Oncidium armillare Lindl. Fol. Orch. Oncidium 51, no. 182.
1855; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 246, fig. 20F, a-c.
1922. Oncidium foveatum Lindl. Fol. Orch. Oncidium 52, no. 183.
1855; Cogn. Martius Fl. Bras. 3, pt. 6: 387. 1906; Kranzl. Engler,
Pflanzenr. IV. 50 (Heft 80), 247. 1922.
Plant medium-sized to large; pseudobulbs oblong-ovoid, compressed, unifoli-
ate, yellowish, densely crowded (according to Kranzlin), 3-4.5 cm. long. Leaf
linear or oblong-linear, obtuse or subacute with an apicule, narrowed at the com-
plicate base, 6.5-23 cm. long, 1-1.5 cm. wide. Scape slender, lateral, basal in the
axil of a sheath, exceeding the leaf, paniculate above with ascending, lightly fracti-
flex branches which are rather loosely few-flowered and 8.5 cm. or less long. Floral
bracts very small and slender. Pedicellate ovary long and slender, many times
exceeding the bracts. Flowers with spreading segments, yellow barred with brown.
Dorsal sepal lanceolate or oblong-lanceolate, acute or acuminate, about 8-9 mm.
long, 2.5-3.5 mm. wide. Lateral sepals similar, free, oblique. Petals oblong, acute
or apiculate, almost as long as the sepals but somewhat broader. Lip much longer
than the sepals, prominently 3-lobed, about 12-13 mm. long from the base to either
side of the retuse apex, 7-11 mm. wide near the base; lateral lobes semiorbicular
with crenulate margins; mid-lobe reniform, sessile or with a very short isthmus,
deeply bilobed, subcordate at the base, about 12-15 mm. wide; disc at the base
with a variable callus (sometimes foveate below) consisting of 3 undulate, callose
ridges terminating in the middle in an acute tooth, and 2 similar teeth in front of
the lateral lines and several free teeth in a semicircle on each side. Column short,
with a pair of prominent, bilobulate, dolabriform or semiovate wings.
Peru: Habitat not recorded, W. Lobb s.n. (type of 0. armillare).
Sources of the Maranon, Warscewicz s.n. (type of 0. foveatum). Also
Colombia and Ecuador.
Oncidium aureuni Lindl. Sert. Orch. sub t. 25, no. 5. 1838;
Fol. Orch. Oncidium 22, no. 67. 1855; Rolfe, Orch. Rev. 3: 69. 1895;
Cogn. Martius Fl. Bras. 3, pt. 6: 306. 1905; Kranzl. Engler, Pflanzenr.
IV. 50 (Heft 80), 209. 1922. Odontoglossum bicolor Lindl. Bot.
SCHWEINFURTH: ORCHIDS OF PERU 861
Reg. 31, Misc. 59, no. 38. 1845. Odontoglossum festatum Reichb.
f. Bonpl. 2: 100. 1854. Odontoglossum hemichrysum Reichb. f. &
Warsc. Bonpl. 2: 100. 1854. Oncidium aureum (B) stenochilum Lindl.
Fol. Orch. Oncidium 22, sub no. 67. 1855. Ornithidium bicolor
Lindl. ex Reichb. f. Walp. Ann. 6: 489. 1863. Oncidium festatum
Reichb. f. & Warsc. ex Reichb. f. Walp. Ann. 6: 733. 1863, in synon.
Oncidium hemichrysum Reichb. f. Walp. Ann. 6: 733. 1863. Oncidium
dichromum Rolfe, Orch. Rev. 3: 363. 1895.
Plant medium-sized to large, variable throughout. Pseudobulb ovoid to pyri-
form (sometimes cylindric in the dried specimens), 2-5 (rarely 13) cm. long, 1- or
2- (sometimes 3-leaved), crowded (often only one present in the dried specimen),
surrounded at the base by 2 or 3 pairs of imbricating, leaf-bearing sheaths. Leaves
elliptic-oblong to linear, linear-oblanceolate or oblong-oblanceolate, subacute to
acuminate, narrowed to a conduplicate base, rigid, up to 40.7 cm. or more long,
1.2-3.4 cm. wide (commonly largest on the pseudobulbs). Scape more or less sur-
passing the leaves, 23 to over 144 cm. long, with a strict, remotely sheathed pedun-
cle, racemose to paniculate, with several ascending or spreading branches, few to
many-flowered; inflorescence or its apical portion commonly nodding; rachis more
or less fractiflex. Floral bracts conspicuous, spreading, ovate, concave. Pedicel-
late ovary much surpassing the bracts, about 3 cm. or less long. Flowers very
variable, yellow throughout, or the sepals and petals commonly brown, wine-red
or purplish, and the lip yellow. Dorsal sepal ovate, concave, acute or acuminate,
9.7-15 mm. long, 5-7 mm. wide. Lateral sepals lanceolate-ovate to oblong-lanceo-
late, acute or acuminate, concave, connate below the middle, somewhat longer and
narrower than the dorsal sepal, more or less oblique. Petals oblong-ovate, acute or
acuminate, shorter than the dorsal sepal, and either narrower or broader. Lip
much larger than the other segments, obovate, oblong-obovate or reniform-obo-
vate, distinctly clawed, 15-22 mm. long; claw short, occupied by a fleshy, sulcate
callus, the erect sides of which are often toothed near the apex and sometimes en-
close several approximate papillae which extend onto the disc; lamina broadly
rounded to subtruncate in front with an abrupt apicule, cordate at the base, about
10-32 mm. wide in front. Column short, stout, clavate, 4-6 mm. high at the back,
with a pair of small, porrect, decurved, toothed or lacerate wings near the apex.
Peru : Near Palca, Ruiz & Pav. 1 794 (type of Odontoglossum fes-
tatum). Source of Rio Maranon, Warscewicz s.n. (0. aureum (B)
stenochilum). — Amazonas: East of Chachapoyas, between Tambo
Ventillas and Piscohuanuma, on grassy plains, 2700-2800 meters,
Weberbauer 4420. — Apurimac: Trail from Abancay to Soccllaccasa
Pass, about 3500 meters, in moss and leaf-mold, overhanging edges
of cliffs, in full exposure to sun, West 3817. — Cuzco: "Valle de Pilla-
huata, montanas de Paucartambo," Herrera 3333. "Pillahuata,"
Cerro de Cusilluyoc, 2200-2400 meters, epiphyte in forest, Pennell
13964- Prov. of Paucartambo, between Ananacu and Pillahuata.
about 2800-3300 meters, on rock outcrops and among shrubs in open
brushland, West 7062. Prov. of Paucartambo, District of Marca-
862 FIELDIANA: BOTANY, VOLUME 30
chea, Achirani, 2750 meters, Vargas 11115.— Huanuco: Cani, "pueblo
7 miles n.e. of Mito," about 2600 meters, on steep grassy slopes,
"Remi Remi," Macbride 3482. "6 miles south of Mito," about 3070
meters, in soil on "dense bunch grass slope," Macbride & Featherstone
1839. Pampayaco (Pampayacu), Kanehira 305. Prov. of Pachitea,
above Molina near Panao, Asplund 13707. Panao, about 2700 me-
ters, on shrubby slopes, Macbride 3609. Playapampa, about 2700
meters, on sphagnum banks, Macbride 4483.— Lima: Near Lima
(type of Odontoglossum bicolor).—Junin: Carpapata, about Hua-
capistana, almost 2400 meters, terrestrial on open hillside, Killip &
Smith 24341. Same locality, 2700-3200 meters, terrestrial on edge
of forest, Killip & Smith 24482. Huassa-huassi, 2900 meters, "in
moss reaches height of 4 feet," Woytkowski 41. Near Andinamarca,
in high mountains, Mathews 1068 (type of Oncidium aureum).—
Puno: Prov. of Sandia, near Limbani, 3200-3450 meters, "buried in
wet moss sheets, in shade," Metcalf 30521 . Also Bolivia and Ecuador.
Oncidium Batemanianum Farm, ex Knowles & Westc. Fl.
Cab. 3: 183, t. 137. 1840, as 0. Batemannianum; Cogn. Martius Fl
Bras. 3, pt. 6: 385, t. 86. 1906; Kranzl. Engler, Pflanzenr. IV 50
(Heft 80), 247. 1922.
Plant large, showy, variable. Pseudobulbs broadly ovoid to subconic-cylindric
more or less compressed, 2- or 3-leaved, 3-10 (rarely 13) cm. long, provided at the
base with 1 or 2 leaf-bearing sheaths. Leaves oblong or linear, acute or acuminate
gradually long-narrowed to a conduplicate base, 18-40 (rarely 60) cm. long, up to
2 (rarely 3) cm. wide, blades on the pseudobulbs much the largest, rather strict.
Scape much surpassing the leaves, paniculate above; peduncle stout, about 103 5
cm. or less tall, with 5-8 remote, short, close, tubular sheaths; panicle rather
short, with several arcuate-recurved or spreading branches, rather densely many-
flowered, about 15-27 cm. long, the rachis and branches more or less fractiflex.
Floral bracts small, narrowly triangular, spreading, much shorter than the slender
pedicellate ovary which is 1.5 cm. or less long. Sepals and petals yellow spotted
with chestnut brown, more or less distinctly clawed; lip yellow. Dorsal sepal ellip-
tic to ovate, acute or acuminate, concave, 6-8 mm. long, 2.5-3.8 mm. wide. Lateral
sepals free, obliquely oblong-elliptic or elliptic-lanceolate, cymbiform, somewhat
longer and much narrower than the dorsal sepal. Petals obliquely oval or ovate,
rounded to slightly retuse at the apex, apiculate, commonly about as long as the
dorsal sepal but much wider (sometimes nearly twice as wide as the lateral sepals).
Lip much larger than the other segments, deeply pandurate or 3-lobed, bilobed in
front, 14 (rarely 11)-17 mm. long from the rounded base to either side of the retuse
apex, 10-11 mm. wide near the base and 13 (rarely 11)-16 mm. wide near the apex;
lateral lobes relatively small, retrorse, triangular-auriculate, rounded at the apex!
mid-lobe obovate-reniform, shortly clawed, often with irregularly crenulate mar-
gins; disc with a multituberculate callus between the lateral lobes and one or more
free teeth on each side. Column very short and stout, clavate, 2.5-3 mm. high at
the back, with a large, fleshy, laterally flattened, longitudinally sulcate protuber-
SCHWEINFURTH: ORCHIDS OF PERU 863
ance in front, provided with a pair of small, transversely oval, often lobulate
wings above.
Cuzco: Altura de Quellow . . ., 1,000 meters, terrestrial, Bues s.n.
Prov. of Convention, Hda. Potrero, Sapan-Sachayocc, 2200 meters,
terrestrial on open grassy slope, Vargas 2530. Prov. of Quispican-
chis, Marcapata Valley, near Chilechile, 2500-2600 meters, in grassy
plains, mixed with shrubs alternating with "shrubwoods," Weber-
bauer 7869.— Huancavelica: Prov. of Tayacaja, northwest of Surcu-
bamba, on trail to Mantaro Bridge, 2200 meters, loam in shrubwood,
'Sacapa," Stork & Horton 10410. Also Brazil (type) and Bolivia.
Oncidium Baueri Lindl. Bauer, 111. Orch. (Genera), t. 7. 1833;
Cogn. Martius Fl. Bras. 3, pt. 6: 398, t. 85. 1906; Urban, Symb.
Antill. 6: 645. 1910; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80),
257. 1922. Oncidium altissimum Lindl. Bot. Reg. 19: t. 1651. 1833,
not Sw. Oncidium bolivianum Schltr. Fedde Repert. 10: 459. 1912;
Mansf. Fedde Repert. Beih. 58: t. 57, nr. 226. 1930.
Plant large, variable, with a stout and long-creeping rhizome. Pseudobulbs
oblong-ovoid to cylindric, strongly compressed, bifoliate (rarely unifoliate), 5-
15 cm. long, apparently subremote, surrounded below by 2 or more pairs of evanes-
cent, imbricating sheaths that bear successively larger leaves upward. Leaves
(much larger on the pseudobulbs), linear-oblong to lorate, obtuse to short-acumi-
nate, gradually narrowed below to a conduplicate base, 20-78 cm. long, commonly
2-3 (rarely 6) cm. broad. Scape much surpassing the leaves, sometimes 3 m. or
more high, erect or nodding; peduncle about 22 cm. or more long, provided with
several short, remote, close, tubular sheaths; inflorescence consisting of an elongate
panicle with numerous short to elongate, simple, few- to many-flowered branches.
Floral bracts triangular-ovate, scarious, commonly much shorter than the slender,
pedicellate ovary which is 30 mm. or less long. Flowers rather small, with spread-
ing, undulate-margined segments, greenish-yellow or sulphur-yellow, barred with
brown. Dorsal sepal elliptic-lanceolate, acute or acuminate, 12-15 (rarely 8.3) mm.
long, 3.5-5 mm. wide. Lateral sepals free, a little longer and often narrower than
the dorsal sepal, oblique, elliptic-lanceolate or oblanceolate-oblong, lightly navicu-
lar, acute, with a carinate, recurved apex. Petals obliquely elliptic-oblong, some-
what sigmoid, acute with a recurved apex, similar to the dorsal sepal but commonly
slightly shorter and narrower. Lip pandurate-trilobed, about as long as the lateral
sepals but much broader, 12-17 mm. long from the base to either side of the retuse
apex, 10 (rarely 7)-12 mm. wide across the lateral lobes; lateral lobes small, spread-
ing, rounded, auriculiform; claw (between the lobes) abrupt, long and broad, sub-
quadrate, slightly narrowed above; mid-lobe reniform, emarginate, 11.5 (rarely
10)-15 mm. wide; disc at the base with a prominent, subquadrate, multitubercu-
late callus in 3 series, terminating in 3 subequal teeth. Column rather short and
stout, arcuate, up to 6 mm. high, 2-angled in front, and above with a pair of short
but broad, subdolabriform wings which are lobulate on the margins.
Loreto: Mishuyacu, near Iquitos, 100 meters, in forest, King 498.
Vicinity of Iquitos, 100 meters, on living tree by river, King 10002.
864 FIELDIANA: BOTANY, VOLUME 30
—San Martin: Alto Rio Huallaga, Rio Mayo, Tarapoto, 360-900
meters, in forest, Williams 6240 (flowers poor). Zepelacio, near
Moyobamba, in forest, 1100-1200 meters, Klug 0.19. Also West
Indies (type of 0. Baueri), Colombia, Venezuela, British Guiana
(type of 0. altissimum Lindl.), Brazil, Ecuador, and Bolivia (type of
0. bolivianum).
Oncidium cajamarcae Schltr. Fedde Repert. Beih. 9: 113.
1921; Mansf. Fedde Repert. Beih. 57: t. 129, nr. 506. 1929; Kranzl.
Engler, Pflanzenr. IV. 50 (Heft 80), 323. 1922.
Plant large, stout, erect, growing on rocks, with an abbreviated rhizome.
Pseudobulbs oval, compressed, unifoliate, about 6.5 cm. high, surrounded on each
side by 1 or 2 leaf-bearing sheaths. Leaves broadly ligulate, subobtuse, up to
40 cm. long, about 4 cm. wide in the middle. Scape tall, slender, erect, paniculate
above, 120-130 cm. high; peduncle few-sheathed, becoming glaucous, 90-100 cm.
high; panicle few-branched, loosely many-flowered. Floral bracts spreading, ob-
long, acuminate, much shorter than the pedicellate ovary. Flowers rather large
for the genus, golden yellow with brown spots in the center. Dorsal sepal broadly
obovate, rounded at the apex, about 15 mm. long, nearly 9 mm. wide. Lateral
sepals free, oblong-obovate, oblique, apiculate at the rounded apex, slightly longer
and distinctly narrower than the dorsal sepal. Petals obovate-subquadrate, retuse
with a minute apicule, oblique, slightly longer and much wider than the dorsal
sepal. Lip much larger than the sepals, subquadrate-obovate in outline, deeply
constricted (and thus 3-lobed) below the middle, cordate at the base, deeply retuse
or bilobed at the apex, about 18 mm. or more long, 13 mm. wide near the base,
20 mm. wide across the anterior portion; lower portion ovate-subquadrate, with
rounded, spreading, auriculiform basal lobes; anterior portion (mid-lobe) reniform,
retuse, obscurely 4-lobed at the broadly rounded apex; disc at the base largely
occupied by an obovate callus which is 3-dentate below and multiverrucose or
tuberculate above with numerous surrounding warts. Column short, fleshy-dilated
on each side below, about 7.5 mm. high, above with a pair of obliquely semiovate
wings, which are obtusely acute or apiculate at the apex.
Cajamarca: Near Hualgayoc, on cliffs below Hacienda La Ta-
hona, 2600 meters, Weberbauer 4041 (type).
No material of this species was available. The type number was
cited under 0. Batemanianum by Kranzl. Engler, Pflanzenr. IV. 50
(Heft 80), 248. 1922.
Oncidium calanthum Reichb. f. Gard. Chron. (1870) 39;
Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 177. 1922.
Plant with a very long-creeping (flagelliform) or freely hanging rhizome.
Pseudobulbs very distant from one another, ovoid, bifoliate. Leaves oblong-
lanceolate or elliptic-lanceolate, acute or acuminate, complicate below. Scape
large, climbing, 2 or 3 times surpassing the pseudobulb, racemose or paniculate
above, twining, with the short, fractiflex branches loosely several-flowered, often
SCHWEINFURTH: ORCHIDS OF PERU 865
with flowers of 2 kinds (heteranthous). Floral bracts very small, triangular, scari-
ous spreading, much shorter than the slender pedicellate ovary which is 2.5 cm or
less' long Flowers golden yellow. Sterile flowers relatively small, with similar
spatulate-oblong or obovate-oblong segments. Fertile flowers several times larger.
Dorsal sepal obovate-oblong, subacute, about 12-13 mm. long and 4.5 mm wide
(according to Kranzlin). Lateral sepals similar but apparently somewhat longer
and narrower, oblique. Petals oblong or obovate-oblong, hastate-clawed, obtuse,
about as long as the dorsal sepal but markedly wider. Lip about as long as the
sepals but much broader, pandurate, three-lobed, the basal portion (across the
lateral lobes) being about 10 mm. wide and the anterior portion (mid-lobe) about
20 mm wide; lateral lobes small, obovate-auriculate, rounded at the apex, con-
tracted in front into a short, broad claw; mid-lobe cordate-reniform, emargmate
at the broadly rounded apex; disc at the base and on the claw with a large, verru-
cose callus consisting of a cluster of warts in 5-7 longitudinal series. Column short,
with a pair of rounded or semirhombic dilations below and with a pair of obliquely
triangular or semiovate, lobulate wings above.
Peru or Ecuador; imported by Messrs. Backhouse.
No exact dimensions of the vegetative parts are given; the floral
description is compiled from a reduced drawing in the Ames Her-
barium.
Oncidium cimiciferum Reichb. f. ex Lindl. Fol. Orch. Oncid-
ium 9, no. 22. 1855. Cyrlochilum flexuosum HBK. Nov. Gen. et Sp.
1- 350 1816; Kranzl. Notizbl. Bot. Gart. Berl. 7: 96, in key. 1917,
nomen; Engler, Pflanzenr. IV. 50 (Heft 80), 63, fig. 6C, a-c. 1922.
Odontoglossum cimiciferum Reichb. f. Linnaea 22: 849. 1849. On-
cidium flexuosum Lindl. Sert. Orch. sub t. 48. 1842, not Sims.
Plant large, straggling. Pseudobulbs oblong-cylindric, up to 7.5 cm. high,
1- or 2-leaved, surrounded at the base by 3 or 4 pairs of imbricating, conduplicate,
leaf-bearing sheaths which bear successively larger blades upward. Leaves linear-
oblong to narrowly lanceolate (rarely elliptic-ligulate), acute or acuminate, nar-
rowed at the conduplicate base, 23 to over 65 cm. long, 2-5 cm. wide. Scape very
long sometimes 3 m. in length, more or less climbing above, with numerous short,
remote, flowering branches in the upper portion, the rachis spreading or reflated,
loosely few-flowered, fractiflex, and often compound (according to Kranzlin).
Floral bracts conspicuous, ovate-oblong, acute, concave, much shorter than the
pedicellate ovary which is commonly 2-3 cm. long. Flowers small, fleshy, brown
or greenish-yellow with brown bands, the perianth segments reflexed and undulate.
Dorsal sepal elliptic-oblanceolate to spatulate, acute or acuminate, 9.5-11.5 mm.
long, 4-4.2 mm. wide. Lateral sepals obliquely oblong-oblanceolate, a little longer
and narrower than the dorsal sepal. Petals obliquely obovate-elliptic, acute, dis-
tinctly shorter and broader than the dorsal sepal. Lip shorter and broader than
the sepals, simple, sessile, ovate-triangular, strongly convex, acute or acuminate,
9-9.5 mm. long, 7-8 mm. wide across the subtruncate base; disc on the central part
of the basal half with a fleshy, multilobulate callus. Column very short and stout,
about 4 mm. high at the back, with a broad, fleshy, semirhombic thickening on
866 FIELDIANA: BOTANY, VOLUME 30
each side below and with a pair of very obscure, broad wings above. Kranzl. (I.e.)
gives smaller floral measurements.
Huanuco: Muiia, about 2150 meters, epiphyte, Macbride 3974.—
Junin: Prov. of Tarma, Agua Dulce, 2300 meters, in low highland
wood about the grass-steppe, Woytkowski 35453. Also Colombia
(type of Cyrtochilum flexuosum), Venezuela (type of Odontoglossum
cimiciferum) and Ecuador.
Oncidium cocciferum Reichb. f. & Warsc. Bonpl. 2: 101, 279.
1854; Lindl. Fol. Orch. Oncidium 9, no. 23. 1855. Cyrtochilum cocci-
ferum Kranzl. Notizbl. Bot. Gart. Berl. 7: 96, in key. 1917, nomen;
Engler, Pflanzenr. IV. 50 (Heft 80), 64, fig. 6E, a-c. 1922.
Plant large, epiphytic, with a very stout, creeping rhizome. Pseudobulbs ap-
proximate, oblong-ovoid, flattened or ancipitous, unifoliate, obliquely inserted on
the rhizome, up to 8 cm. long (according to Kranzl.), invested at the base by 3 or
more pairs of conduplicate, imbricating, leaf-bearing sheaths, the blades successively
larger upward. Leaves similar, oblong-elliptic (linear, according to Kranzlin),
acuminate, gradually narrowed to a conduplicate base, 24-60 cm. long (the latter
measurement according to Kranzl.), 1.5-4 cm. wide. Scape up to more than
1.5 m. long (according to Kranzl.), paniculate above, with remote, strongly fracti-
flex, loosely few- (3- to 5-) flowered branches of which the lower are often com-
pound. Floral bracts very small, convolute, oblong-ovate, acute, many times
shorter than the slender pedicellate ovary that is up to 2.5 cm. long. Flowers
small, greenish brown, obscurely banded with yellow, with widely spreading seg-
ments. Dorsal sepal obovate or spatulate, clawed, acute, up to 9 mm. long and
5 mm. wide. Lateral sepals similar but somewhat longer, oblique. Petals broadly
obovate, scarcely clawed, apiculate, shorter than the dorsal sepal but equally wide.
Lip triangular or triangular-ovate, subacute, subcordate at the base, strongly con-
vex, about 8 mm. long and 5 mm. wide below; disc with a large ovate-subquadrate,
verrucose callus extending from the base to beyond the middle or about to the
middle and with several smaller tubercles on each side. Column short, stout,
dilated on each side near the base, the apical wings scarcely produced.
Peru: Sources of the Maranon River, Warscewicz s.n., fide Lind-
ley.— Huanuco, fide Schlechter, Fedde Repert. Beih. 9: 173. 1921.—
Junin : Prov. of Tarma, mountains of Yanangu, east of Huacapistana,
2200-2300 meters, thickets with scattering trees, Weberbauer 2127.
Also Colombia (type).
No material of this species was available, but the Ames Her-
barium contains several drawings of the vegetative and floral parts
of the type, from the Reichenbach Herbarium.
Oncidium cordatum Lindl. Sert. Orch. sub t. 25, no. 2. 1838;
Fol. Orch. Oncidium 4, no. 2. 1855. Cyrtochilum cordatum Kranzl.
Notizbl. Bot. Gart. Berl. 7: 92, in key. 1917, nomen; Engler, Pflan-
zenr. IV. 50 (Heft 80), 41, fig. 2G a-b. 1922.
SCHWEINFURTH: ORCHIDS OF PERU 867
Plant large, with a stout, long-creeping rhizome. Pseudobulbs more or less
distant, narrowly ovoid or conic, bifoliate, about 9 cm. long, invested and almost
concealed by several pairs of conduplicate, leaf-bearing sheaths. Leaves on the
sheaths larger, elliptic-lanceolate (linear-lanceolate according to Kranzl.), acute or
acuminate, narrowed to a complicate base, 20-44 cm. long, up to 3.8 cm. wide.
Scape about 1.5 m. long, twining, paniculate above with branches (rarely com-
pound) about 15 cm. long. Floral bracts large, deeply concave, suborbicular-ovate
when expanded, obtuse, 1.5 cm. long, much shorter than the slender, pedicellate
ovary which is about 3.5 cm. or less long. Flowers large, thin, with spreading seg-
ments, about 5.7 cm. across, apparently brown bordered with yellow. Dorsal sepal
long-clawed; lamina round-ovate, acute, about 19 mm. long and 13-15 mm. wide;
claw slender, about 6 mm. long. Lateral sepals similar but a little longer than the
dorsal sepal; claw longer, auriculate at the base, gradually cuneate above; lamina
rhombic-ovate, subacuminate, about equally long with the dorsal lamina but nar-
rower. Petals shorter than the dorsal sepal with a short, broad claw; lamina
broadly triangular-ovate, acuminate, lightly oblique, about as broad as the dorsal
sepal or slightly narrower. Lip strongly reflexed, clawed, about 25 mm. long when
expanded and more than 10 mm. wide near the base; claw short, gradually dilated
above; lamina hastate-lanceolate, acuminate, with short, retrorse, lanceolate auri-
cles or lobules which are acute or acuminate and have a filiform tooth at the base;
disc with 5 oblique, transverse lamellae, of which the inner 3 are asymmetrically
bilobed and the smaller outer ones irregularly lacerate. Column small, almost
wingless, but with a rather conspicuous linear-lanceolate tooth on either side
below.
Peru: Near Pangoa, on rocks near banks of river, Mathews 1067
(type).— Junin: Prov. of Tarma, mountains of Yanangu, east of Hua-
capistana, 2300-2400 meters, Weberbauer 2119—~Loreto, fide Schlech-
ter, Fedde Repert. Beih. 9: 173. 1921.
No specimen of this species was available, but the Ames Herbar-
ium contains a photograph of the type from the Lindley Herbarium.
Oncidium crista-galli Reichb. f. Bot. Zeit. 10: 697. 1852;
Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 165. 1922. Oncidium
iridifolium Lindl. Bot. Reg. 22: 1. 1911. 1836, not H.B.K. Oncidium
decipiens Lindl. Fol. Orch. Oncidium 22, no. 68. 1855. Odontoglossum
crista-galli Kranzl. ex Weberb. Pflanzen. Peru. And. 290. 1910.
Plant small, caespitose, rarely up to 10 cm. high. Pseudobulbs suborbicular
or broadly ellipsoid, strongly compressed, apparently aggregated, about 10 mm.
or less high, bearing one small, immature leaf at the summit, invested and mostly
concealed by 2 or more pairs of the dilated bases of distichous leaves. Leaves flat
(but often conduplicate in the dried specimen), elliptic-lanceolate to linear-lanceo-
late, acute or acuminate, membranaceous, up to 7.5 cm. long excluding the sheath-
ing base, up to (rarely) 1 cm. wide. Inflorescence about equaling the leaves, fili-
form; peduncle bearing one or more close, tubular sheaths; raceme loosely 1- to
5-flowered. Floral bracts prominent, spreading, ovate to oblong-lanceolate, am-
plexicaul, more or less shorter than the pedicellate ovary. Flowers large for the
868 FIELDIANA: BOTANY, VOLUME 30
plant, successive, yellow often with brown markings, the segments wide-spreading
and very variable in size. Dorsal sepal elliptic, elliptic-lanceolate or obovate-
elliptic, acute or apiculate, 5.7-6 mm. long, 3.2-3.6 mm. wide. Lateral sepals free,
obliquely elliptic-lanceolate, more or less semilunate, acute, about as long as the
dorsal sepal but about half as wide. Petals ovate-oblong or subquadrate-oblong,
obtuse, distinctly longer than the sepals and slightly broader than the dorsal sepal,
lightly oblique. Lip much larger than the other segments, 3- or 6-lobed, about
13.5-22 mm. long from the base to the apex of an anterior lobule, 12-22 mm. wide
across the lateral lobes; lateral lobes obliquely round-obovate, horizontally spread-
ing from a short claw, more or less irregularly crenate on the outer margins; mid-
lobe much larger, shortly clawed, semiorbicular in outline, deeply bilobed, with a
sharp indentation in the middle of each side (thus 4-lobed), with the apical lobules
broadly oblong or ovate and parallel, and the lateral lobules broadly rounded or
semiorbicular; disc at the base with a large, flat, lobulate callus, surrounded on
each side by 1 or more papillae. Column very short and stout, with a prominent,
fleshy, ovate protuberance in front below and with a pair of short, broad, sub-
dolabriform wings above.
Peru: Habitat not recorded, Jurgensen 646, Warscewicz s.n. (type
of 0. crista-galli) , fide Kranzlin. — Loreto, fide Schlechter, Fedde Re-
pert. Beih. 9: 174. 1921. Also Mexico (type of 0. decipiens), through
Central America to Panama, and Colombia.
Oncidium cruciferum Reichb. f. & Warsc. Bonpl. 2: 109. 1854;
Lindl. Fol. Orch. Oncidium 50, no. 178. 1855; Kranzl. Engler, Pflan-
zenr. IV. 50 (Heft 80), 242, fig. 20H, a-e. 1922.
Pseudobulbs and leaves not recorded. Scape very long, the portion existing
about 1.2 m. long, branching throughout, the branches short, widely spreading,
lightly fractiflex, loosely few- (4- to 5-) flowered and 5-6 cm. apart. Floral bracts
ovate, concave, paleaceous, spreading, about a fourth as long as the slender, pedi-
cellate ovary which is up to 2.5 cm. long. Perianth segments spreading, yellow
with red or brown spots, according to Kranzlin. Sepals narrowly elliptic-lanceolate,
acute or acuminate, undulate, about 15 mm. long and 3 mm. wide, the free lateral
sepals somewhat longer. Petals similar to the sepals but slightly shorter and
broader. Lip nearly as long as the sepals but much broader, pandurate-trilobed,
about 14 mm. long, 8-9 mm. wide across the basal lobes, 12-15 mm. wide in front;
lateral lobes relatively small, retrorse-spreading, auriculiform, subquadrate-rounded;
mid-lobe cordate-reniform from a long, gradually narrowed isthmus, lightly emar-
ginate in front; disc at the base with a conspicuous, velutinous, 3-lobed or cruci-
form callus, of which the lateral arms are broadly ligulate, arcuate-incurved and
dentate at the apex, and the middle arm elongate, retuse, gradually dilated above,
and dentate at the apex. Column small, sigmoid when viewed from the side, below
in front a suborbicular plate and aboveja pair of very short, broad, tricrenate
wings.
Peru: Habitat unrecorded, Warscewicz s.n.
The Ames Herbarium contains a drawing of the type from the
Reichenbach Herbarium.
SCHWEINFURTH: ORCHIDS OF PERU 869
Oncidium cryptocopis Reichb. f. Card. Chron. (1870) 827;
Hook. f. Bot. Mag. 96: t. 5858. 1870. Cyrtochilum cryptocopis
Kranzl. Notizbl. Bot. Gart. Berl. 7: 92, in key. 1917, nomen; Engler,
Pflanzenr. IV. 50 (Heft 80), 36. 1922.
Plant large and showy throughout. Pseudobulbs very narrowly ovoid to
cylindric, apparently aggregated, much compressed, bifoliate, 10-12.7 cm. long,
clothed at the base by several (up to 3) pairs of leaf-bearing sheaths. Leaves ob-
long-linear or lorate, slightly broader above the middle, acute, about 30 cm. long
and 3 cm. wide. Scape very long, twining above, about 1-1.5 m. or more long,
with remote branches which are short, twining and few- (often 1-) flowered.
Floral bracts cucullate, scarious, acute, much shorter than the slender pedicellate
ovary which appears to be 8 cm. or less long. Flowers large, about 7.6 cm. wide
from the apex of the dorsal sepal to the apices of the lateral sepals, pale chestnut-
brown with the margins of the sepals and petals golden yellow and the mid-lobe of
the lip yellow. Dorsal sepal shortly clawed; lamina broadly deltoid-ovate, acute or
short-acuminate, with an undulate-crisped margin, about 22 mm. long, 22-28 mm.
wide; claw auricled at the base. Lateral sepals subparallel, much longer than the
dorsal sepal, long-clawed; lamina obliquely elliptic-ovate or oblong-ovate, acute or
acuminate, with recurved apex; claw slender, gradually dilated above, about
20 mm. long. Petals nearly as long as the dorsal sepal but much narrower, with
undulate-crisped margins; lamina ovate-lanceolate or rhombic-ovate, obtuse, re-
curved above; claw short, broad. Lip about half as large as the petals, with the
basal portion spreading from the base of the column and the anterior part abruptly
reflexed, pandurate-trilobed when expanded, about 12 mm. long (according to
Kranzlin); lateral lobes relatively small, falcate-triangular, reflexed, acute; mid-
lobe much larger, from a long stout claw abruptly dilated into a transversely ob-
long, more or less retuse lamina which is about 11 mm. wide; disc at the base
(between the lateral lobes) with a callus which is carinate in the middle and multi-
papillose on each side. Column very short, with a pair of spreading, obliquely
rhombic-cuneate wings and with a pair of flattened-subulate, parallel, decurved
horns in front.
Peru: Habitat not recorded; imported by Mr. Bull.
No example of this species was available, but the Ames Herbar-
ium contains sketches of the flower from the Reichenbach Herbarium.
Oncidium deltoideum Lindl. Bot. Reg. 23: t. 2006. 1837; Fol.
Orch. Oncidium 31, no. 103. 1855. Cyrtochilum deltoideum Kranzl.
Notizbl. Bot. Gart. Berl. 7: 97, in key. 1917, nomen; Engler, Pflan-
zenr. IV. 50 (Heft 80), 70, fig. 7A. 1922.
Plant medium-sized, rather showy. Pseudobulbs oblong-ovoid, compressed, 2-
or 3-leaved, up to 6 cm. high, approximate, clothed at the base by 1 or 2 pairs of
sheaths of which the larger upper one is commonly leaf-bearing. Leaves oblong-
lanceolate, acute, complicate at the base, 12-20 cm. long (according to Kranzlin),
2-3 cm. wide. Scape about 42 cm. to almost 1 m. high (according to Kranzlin), ter-
minating in a large, often nodding panicle of which the branches are spreading, com-
pound, loosely many-flowered, with a fractiflex rachis. Floral bracts minute, ovate,
870 FIELDIANA: BOTANY, VOLUME 30
acute, much shorter than the slender, pedicellate ovary which is up to 2 cm. long.
Flowers small for the genus, pale yellow, sometimes spotted with red, the sepals and
petals horizontally spreading and recurved above. Dorsal sepal spatulate from a
narrow claw-like base, rounded-obtuse, about 8-12 mm. long, 3-6 mm. wide.
Lateral sepals oblong-oblanceolate, oblique, acute, somewhat longer (with a longer
claw) and narrower than the dorsal sepal. Petals obovate-oblong to subspatulate,
rounded to subacute at the apex, apparently somewhat shorter and broader than
the dorsal sepal, with more or less undulate margins. Lip much shorter and broader
than the sepals, simple or subsimple, broadly deltoid-ovate, subacute or bluntly
apiculate, cordate at the base with minute, cucullate lobules next to the column,
convex, with rounded outer sides, 7-8 mm. long, 10-13 mm. wide below; disc with
the central portion adorned to above the middle with a crest consisting of 2 series
of short oblong papillae (the anterior series broader with longer members). Column
short, about 6 mm. long, above with a pair of spreading, broadly semiovate wings
which are entire to minutely denticulate, and below with a pair of subquadrate
plates.
Amazonas: Chachapoyas, Mathews s.n. (type). — La Libertad:
Prov. of Otuzco, Hda. Mombrillar, 1670 meters, epiphyte, Ldpez
0582.
The Lopez collection has a flower that coincides well with Kranz-
lin's figure of Cyrtochilum deltoideum, although it appears to be some-
what different from Lindley's plate of the type of Oncidium deltoideum.
Oncidium diceratum Lindl. Fol. Orch. Oncidium 6, no. 11.
1855. Cyrtochilum diceratum Kranzl. Notizbl. Bot. Gart. Berl. 7:
95, in key. 1917, nomen; Engler, Pflanzenr. IV. 50 (Heft 80), 60,
fig. 5G, a-d. 1922.
Plant large, without any description of the vegetative portions. Scape very
long (according to Kranzlin), twining, paniculate above. Flowering portion
(?branch) shown by the type about 34 cm. long, loosely several-flowered, the lower
flowers on short, 2-flowered branches, the upper ones solitary; branches (according
to Kranzlin) strict, up to 12 cm. long and 12 cm. distant. Floral bracts apparently
prominent, ovate, concave, about half as long as the pedicellate ovary which is up
to 3.2 cm. long. Flowers rather large, with horizontally spreading segments, about
4.7 cm. long from the apex of the dorsal sepal to the apices of the lateral sepals,
apparently dark brown with the apex of the lip yellow. Dorsal sepal abruptly
clawed, up to 21 mm. long; lamina ovate-suborbicular, rounded at the apex, up to
15 mm. wide; claw short, auriculate at the base. Lateral sepals free, slightly longer
and markedly narrower than the dorsal sepal, the claw dilated above and the lam-
ina obliquely elliptic-ovate, broadly obtuse. Petals markedly shorter and narrower
than the dorsal sepal with a very short, broad claw; lamina ovate, obtuse. Lip
much smaller than the sepals, 15-18 mm. long, pandurate-trilobed, 7-8 mm. wide
across the dilated basal portion and about 10 mm. wide in front (according to
Kranzlin), subcuneate at the very base, retuse at the apex, spreading below and
reflexed below the middle; basal portion suborbicular with reflexed, semiorbicular
lateral lobes; anterior portion (mid-lobe) much larger, obcordate from an indistinct
SCHWEINFURTH: ORCHIDS OF PERU 871
claw; disc at the base (between the lateral lobes) with 4 approximate warts (the 2
lateral ones parallel to the central ones), and with several smaller warts in front.
Column short, stout, subclavate, about 7.5 mm. high, with a pair of small, fleshy,
falcate-ascending horns above.
Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 174. 1922.
Ecuador (type) and Colombia.
No example of this species was available, but the Ames Herbarium
contains a photograph from the Lindley Herbarium with a drawing
of the lip and column.
Oncidium discobulbon Kranzl. Engler, Bot. Jahrb. 54, Beibl.
117: 33. 1916; Engler, Pflanzenr. IV. 50 (Heft 80), 196. 1922.
Plant medium-sized. Pseudobulbs aggregated, oblong-ovoid, very strongly
compressed, up to 6 cm. long, unifoliate, yellow and shining in the dried specimen,
surrounded at the base by 1 or 2 pairs of leaf-bearing sheaths. Leaves linear,
abruptly acute, up to 36 cm. long and 1.5 cm. wide, chartaceous, paler beneath.
Scape about 57 cm. high; peduncle rather slender, about 22 cm. high, provided with
a very few remote, close, short, tubular sheaths; panicle with numerous, ascending,
compound branches, the rachis loosely many-flowered and f ractiflex. Floral bracts
very small, triangular-lanceolate, much shorter than the slender pedicellate ovary
which is 1.5 cm. or less long. Flowers small, yellow spotted with brown. Dorsal
sepal lanceolate, acute, about 7 mm. long and 1 mm. wide. Lateral sepals free,
similar to the dorsal sepal but longer. Petals "ovate" or ovate-oblong, acute,
equally long with the dorsal sepal but somewhat broader. Lip slightly longer and
much broader than the rest of the perianth, pandurate-trilobed, lightly bilobed at
the apex, subcordate at the base, about 9.5 mm. long from the base to either side
of the retuse apex, about 7 mm. wide near the base and 11 mm. wide near the apex;
lateral lobes semiovate, reflexed; mid-lobe from a very short, broad isthmus
abruptly reniform, bilobed; disc at the base having a small, tripartite crest with 2
tubercles on each side and some small papillae in the middle. Column very small,
sigmoid when viewed from the side, with a pair of erect wings divided into 2 falcate
arms.
Cajamarca: Prov. of Chota, near Huambos, 2400-2500 meters,
Weberbauer 4198.
No material of this species was available, but the Ames Her-
barium contains a photograph of the type.
Oncidium excavatum Lindl. Sert. Orch. sub t. 25, no. 3. 1838;
Fol. Orch. Oncidium 27, no. 85. 1855; Hook. Bot. Mag. 88: t. 5293.
1862; Cogn. & Goos. Diet. Icon. Orch. Oncidium t. 20. 1899;
Cogn. Martius Fl. Bras. 3, pt. 6: 363. 1905; Kranzl. Engler, Pflan-
zenr. IV. 50 (Heft 80), 241. 1922. Oncidium aurosum Reichb. f.
Bonpl. 2: 108. 1854. Oncidium excavatum Lindl. var. Dawsonii Wil-
liams, Orch.-Grow. Man. ed. 6: 483. 1885.
872 FIELDIANA: BOTANY, VOLUME 30
Plant large, showy. Pseudobulbs numerous, aggregated, oblong-ovoid, some-
what compressed, 7-12 (rarely 18) cm. long, 1- or 2-leaved, clothed at the base on
each side with a few evanescent, leaf-bearing sheaths. Leaves linear-oblong or
lorate, acute, long-attenuate to a conduplicate, subpetiolate base, 30-50 cm. long,
2.5-4 cm. wide. Scape stout, erect or subnodding, 60-150 cm. high; peduncle pro-
vided with a few short, remote, acuminate sheaths; inflorescence paniculate with
a few spreading-ascending, simple, elongate branches. Floral bracts small, spread-
ing, narrowly triangular, much shorter than the slender, pedicellate ovary which
is 1.5-3 cm. long. Flowers loose to subdense, variable in size, with wide-spreading
segments, yellow barred and spotted with red-brown through the lower half.
Sepals somewhat undulate on the margin. Dorsal sepal narrowly obovate or ob-
lanceolate-spatulate, rounded and abruptly acute at the apex, 12-16 mm. long,
7-9 (rarely 12) mm. wide, concave. Lateral sepals free, oblong-oblanceolate,
abruptly acute, lightly oblique, a little longer and markedly narrower than the
dorsal sepal. Petals oblong-obovate or triangular-obovate, apiculate at the often
retuse apex, somewhat longer than the dorsal sepal and nearly twice as wide,
strongly undulate on the margin. Lip slightly longer and much broader than the
other segments, pandurate-trilobed, 12-22 (rarely 25) mm. long, up to 10 mm.
wide across the basal portion and 20 mm. or less wide in front, sessile at the sub-
truncate base, more or less deeply emarginate at the apex; lateral lobes small,
rotund-semiovate; mid-lobe much larger, from a short, broad claw abruptly cor-
date-reniform; disc at the base (between the lateral lobes) with a convex plurisul-
cate callus having about 4 verrucose lines and extended into 2 carinate ridges in
front, surrounded on each side by several minute, free tubercles. Column very
short and stout, about 5 mm. high, with a pair of porrect, subdolabriform wings
which are truncate or lightly retuse without.
Peru: Habitat not recorded (type of 0. aurosum). — Amazonas:
Near Chachapoyas, Mathews s.n. (type of 0. excavatum). — Near the
source of the Maranon River, Warscewicz s.n., Wallis s.n. — Caja-
marca, Huanuco, fide Schlechter, Fedde Repert. Beih. 9: 174. 1921.
Also Ecuador.
No example of this species was available, but the Ames Herbar-
ium contains a photograph of the type from the Lindley Herbarium.
Oncidium falciferum Reichb. f. & Warsc. Bonpl. 2: 108. 1854,
fide Schlechter, Fedde Repert. Beih. 9: 174. 1921, sphalm.
Oncidium falcipetalum Lindl. Orch. Linden 14, no. 76. 1846;
Fol. Orch. Oncidium 5, no. 7. 1855; Cogn. & Goos. Diet. Icon.
Orch. Oncidium t. 21. 1899. Oncidium falcipetalum Lindl. var. Pa-
vonii Reichb. f. ex Lindl. Fol. Orch. Oncidium 5, sub no. 7. 1855,
nomen. Oncidium ionodon Reichb. f. Linnaea 41: 23. 1876. On-
cidium Davisii Reichb. f . Linnaea 41 : 24. 1876. Cyrtochilum falci-
petalum Kranzl. Notizbl. Bot. Gart. Berl. 7: 92, in key. 1917, nomen;
Engler, Pflanzenr. IV. 50 (Heft 80), 37, fig. 1G, a-c. 1922. Cyrto-
chilum Pavonii Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 38. 1922.
SCHWEINFURTH: ORCHIDS OF PERU 873
Plant large, sprawling, from a stout, creeping rhizome, both terrestrial and
epiphytic. Pseudobulbs subremote, oval or oblong-ovoid to subcylindric, com-
pressed, 5-10 (rarely 16) cm. long, 1- or 2-leaved, surrounded below by about 3
pairs of conduplicate, leaf-bearing sheaths producing successively larger blades up-
ward. Leaves usually variable in size, those on the pseudobulb commonly the
largest, oblanceolate to narrowly elliptic-lanceolate, acute or short-acuminate, long-
narrowed below to a conduplicate, petiole-like base, up to 52 cm. long and 5.5 cm.
wide. Scape much longer than the leaves, stout, twining, up to 6 meters long
(according to Lindley), bearing at remote, irregular intervals flowering branches
which are commonly simple, loosely several- (3- to 6-) flowered, the rachis (when
mature) fractiflex and about 16 cm. or less long. Floral bracts conspicuous, con-
cave, broadly ovate to suborbicular when expanded, equaling about one half or
one third of the pedicellate ovary which is up to 4.5 cm. long. Flowers large, with
widely spreading segments, commonly 6-7 cm. broad. Sepals chestnut-brown with
a narrow yellow border, strongly undulate on the margins. Dorsal sepal 25-30 mm.
long, clawed; lamina broadly ovate or suborbicular-ovate, obtuse, 17-26 mm. wide
below; claw short, auricled at the base, abruptly passing into the lamina. Lateral
sepals free, clawed, longer and commonly narrower than the dorsal sepal; lamina
obliquely ovate or elliptic-ovate, subacute; claw longer than that of the dorsal
sepal, rather gradually dilated into the lamina, auricled on the outer side at the
base. Petals from a very short, broad claw ovate-lanceolate, oblique, acuminate,
very strongly undulate and often twisted or conduplicate, somewhat shorter than
the dorsal sepal and more or less narrower than the lateral sepals, yellow spotted
with brown. Lip much smaller than the other segments, lanceolate in outline,
more or less 3-lobed, spreading from the base of the column, abruptly reflexed be-
low the middle, about 20-23 (rarely 15) mm. long, about 10-11 mm. wide near the
basal portion when expanded; the side angles or lateral lobes obliquely triangular,
reflexed, acute; the apical portion or terminal lobe lanceolate to oblong-obovate,
acute or acuminate; disc at the base with a high, fleshy, tricarinate or bisulcate
callus having several irregular fleshy tubercles in front and an irregular, fleshy ridge
extending from the callus to the lateral angle of the lip. Column short, stout, about
8.8-10 mm. high at the back, with a pair of fleshy, porrect angles in front below,
and a pair of ascending, linear-lanceolate to cuneate arms (often toothed at the
oblique apex) on each side in the middle.
Peru: Habitat and collector not recorded (types of Oncidium iono-
don and 0. Davisii) .— Huanuco : Pillao, 2700 meters on thick tree
trunks and rarely on the ground, Woytkowski 37019.— Puno: Prov.
of Sandia, 2-6 km. from Oconeque, on trail, 1800-2100 meters, in
moist places in much fog and rain, Metcalf 30589. Also Venezuela
(type of 0. falcipetalum) and Colombia.
Oncidium fasciferum Reichb. f. & Warsc. Bonpl. 2: 108. 1854;
Reichb. f. Walp. Ann. 6: 787. 1863; Cogn. Martius Fl. Bras. 3, pt. 6:
393. 1906; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 243, fig. 20D,
a-e. 1922.
Plant large, showy. Pseudobulbs large, broadly ovoid, unifoliate, clothed at
the base on each side by a leaf-bearing sheath. Leaves linear or lorate, shortly
874 FIELDIANA: BOTANY, VOLUME 30
acute, up to 50 cm. long and 4 cm. wide. Scape strict, 1 m. high, terminating in a
densely many-flowered panicle consisting of more than 30 remote branches which
are up to 5 cm. long and 4- to 8-flowered. Floral bracts ovate, acute, concave,
paleaceous, much shorter than the pedicellate ovary which is about 1.5 cm. long.
Flowers rather small, golden yellow, with wide-spreading segments, purple-spotted
on the lower half. Dorsal sepal clawed, elliptic-lanceolate or linear-lanceolate,
acute or acuminate, with a reflexed apex, up to 11 mm. long, 3 mm. wide. Lateral
sepals similar but rather longer, free. Petals oblong, about as long as the sepals
but markedly wider, oblique, subobtuse. Lip slightly longer and much broader
than the other segments, broadly obovate in outline, pandurate-trilobed, about
13 mm. long from the subtruncate base to either side of the retuse apex and
about 8 mm. wide near the base; lateral lobes semiovate-triangular or semirhom-
bic, spreading, obtuse without; mid-lobe from a short, broad claw abruptly dilated
into a cordate-reniform, bilobed lamina which is 10-11 mm. wide; disc at the base
with a crest consisting of 5 longitudinal series, the middle one shorter and linear,
with larger tubercles on each side of the middle. Column very short and stout,
dilated upward, with a pair of porrect, short, semiovate or dolabriform wings above.
E. Peru: Near the source of the Maranon River, Warscewicz 25
(type).— Huanuco, fide Schlechter, Fedde Repert. Beih. 9: 174. 1921.
No material of this species was available, but the Ames Herbar-
ium contains a floral analysis from the Reichenbach Herbarium.
Oncidium formosissimum Reichb. f. Bot. Zeit. 10: 694. 1852;
Lindl. Fol. Orch. Oncidium 28, no. 92. 1855; Reichb. f. Walp. Ann. 6:
745. 1863; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 174, fig. 15C,
a-c. 1922.
Plant rather large and showy. Pseudobulbs oblong-cylindric, ancipitous, uni-
foliate, 2.5-5 cm. high, clothed below by several leaf-bearing sheaths. Leaves
elliptic-oblong to "oblong-lanceolate," acute, shortly narrowed to a complicate
base, 9 cm. or more long, about 2 cm. wide. Scape much surpassing the leaves,
stout, up to 48 cm. or more tall, terminating in a many-flowered panicle; peduncle
suberect, provided with several remote, close, evanescent sheaths; panicle spread-
ing, consisting of several subremote, short branches the lower of which are again
branched; rachis fractiflex. Floral bracts very short, triangular, acute, much
shorter than the slender, pedicellate ovary. Flowers medium-sized, with wide-
spreading segments. Sepals and petals dark brown or purple below and yellow
above, with a yellow lip. Dorsal sepal elliptic-lanceolate or "oblong," subobtuse,
about 10 mm. long, 3.5 mm. wide. Lateral sepals free, similar to the dorsal sepal
but longer and slightly narrower, oblique. Petals "oblong" to elliptic-oblong,
apparently as long as the lateral sepals and wider than either sepal (sometimes four
times larger, fide Reichenbach), acute or short-acuminate. Lip much larger than
the other segments, about 16 mm. long, pandurate-trilobed, about 12 mm. wide
across the base and nearly 23 mm. wide in front; lateral lobes small, retrorse, ob-
long-ovate (sometimes bilobed), rounded at the apex; mid-lobe much larger, from
an indistinct, broad isthmus, reniform-cordate, broadly rounded in front and
abruptly acute or very obscurely bilobed; disc at the base with a small, low crest
which is tridentate in front and altogether covered with minute tubercles. Col-
umn short, about 7 mm. high, with a pair of very large, transversely oblong or
SCHWEINFURTH: ORCHIDS OF PERU 875
dolabriform wings. Sometimes, according to Kranzlin, there occur forms with
small, sterile, imperfectly 5-divided flowers which are more numerous than the
normal fertile flowers.
Peru(?), fide Schlechter, Fedde Repert. Beih. 9: 174. 1921. Co-
lombia and Ecuador (type).
No specimens of this species were available, but the Ames Her-
barium contains sketches of vegetative and floral parts from the
Reichenbach Herbarium.
Oncidium gargantua Reichb. f. Linnaea 41: 24. 1876. Cyrto-
chilum gargantua Kranzl. Notizbl. Bot. Gart. Berl. 7: 91, in key. 1917,
nomen; Engler, Pflanzenr. IV. 50 (Heft 80), 32, fig. 1A, a-c. 1922.
Pseudobulbs and leaves unknown. Scapes surely very long, stout, climbing,
apparently many-branched at intervals of 12-15 cm. (according to Kranzlin), the
branches fractiflex, loosely up to 7-flowered, and 21 cm. or less long. Floral bracts
conspicuous, triangular-ovate, concave, about half as long as the slender, pedicel-
late ovary which is up to 4 cm. long. Flowers large with wide-spreading segments,
about 6.5 cm. long from the apex of the dorsal sepal to the tip of a lateral sepal,
apparently brown, the petals paler with large spots, similar to those of O. falci-
petalum Lindl. Sepals long-clawed. Dorsal sepal about 28 mm. long; lamina
broadly ovate or triangular-ovate, subacute, rounded or subcordate at the base,
with undulate margins, about 18 mm. or more wide; claw about 5 mm. long, auri-
cled at the base. Lateral sepals similar but distinctly larger (with a longer claw
and a longer and broader lamina), the lamina obliquely suborbicular-ovate, sub-
acute to acute. Petals much smaller than the sepals, shortly clawed, obliquely
ovate-lanceolate or oblong-triangular, acuminate, with margins strongly undulate,
recurved above. Lip much smaller than the other segments, shortly clawed, the
lower portion (nearly to the middle) spreading from the column, then abruptly
reflexed and recurved (forming a semicircle when viewed from the side), triangular-
lanceolate and about 15 mm. long when expanded; lateral lobes small, triangular,
obtuse, deflexed, about 5 mm. long; mid-lobe linear, somewhat dilated above, ob-
tuse or subacute; disc at the base (between the lateral lobes) with a crest consisting
of 2 keels (the rear one bicarinate and papillose in front), the forward one consist-
ing of 2 short lamellae with 1 or 2 small papillae on each side. Column rather
small, apparently about 8 mm. high, dilated below with a pair of semiorbicular
processes, near the middle with a pair of linear, spreading wings which are abruptly
angled near the apex with an ascending tip.
Southern Peru: Habitat not recorded, Pearce 835, 598 (according
to Kranzlin).
No example of this species was accessible, but the Ames Her-
barium contains a drawing of the inflorescence from the Reichenbach
Herbarium.
Oncidium globuliferum HBK. Nov. Gen. et Sp. 1: 347. 1816;
Reichb. f. Gard. Chron. (1871) 1678; Kranzl. Engler, Pflanzenr. IV.
876 FIELDIANA: BOTANY, VOLUME 30
50 (Heft 80), 167. 1922. Oncidium globuliferum /3 costaricense
Reichb. f. Gard. Chron. (1871) 1678. Oncidium scansor Reichb.
f. Linnaea 22: 844. 1849; Reichb. f. Walp. Ann. 6: 742. 1863. On-
cidium convolvulaceum Lindl. & Paxt. Paxt. Fl. Gard. 1: 102, sub t.
21. 1850-51. Oncidium Wercklei Schltr. Fedde Repert. Beih. 19:
68. 1923.
Plant extremely variable, with a very long-creeping, twining stem which is
smooth, often branching and up to 5 m. long (according to Kranzlin). Pseudo-
bulbs small, commonly more or less distant (rarely approximate), up to 27 cm. or
more apart, apparently ancipitous, ellipsoid (rarely obovoid) to suborbicular, up
to 3 cm. high, unifoliate, invested below by 2 pairs of conduplicate sheaths of which
the upper ones are leaf-bearing. Leaves oblong or elliptic-oblong to oval, rounded
or subacute with a minute apicule at the apex, shortly cuneate or subrounded at
the complicate base, up to 8 cm. long and 2.5 cm. wide (usually smaller); those of
the pseudobulb commonly the largest and most persistent. Scapes very short,
arising both from the base of the pseudobulbs (in the axils of sheaths) and from
the stem (at remote intervals), 1- to 3-flowered, apparently up to 3 cm. high, the
peduncle wholly concealed by imbricating, scarious sheaths and the rachis strongly
fractiflex. Floral bracts ovate-lanceolate, concave, much shorter than the slender
pedicellate ovary which is 6 cm. or less long. Flowers very variable in size (medium-
sized to large), with wide-spreading segments, yellow usually barred or spotted
with red-brown at the base of the segments. Dorsal sepal oblong-elliptic or obo-
vate-oblong, obtuse to acute, 11.5-17 mm. long, 5-9 mm. wide. Lateral sepals
distinctly longer and narrower, obliquely oblanceolate-oblong, shortly clawed be-
low, acute to obtuse. Petals obovate-oblong, oblique and somewhat sigmoid,
rounded and minutely apiculate at the apex, about as long as the dorsal sepal
or a little longer, commonly somewhat broader. Lip much larger than the other
segments, quadrate-obovate to suborbicular in outline, pandurate-trilobed, cor-
date at the base, retuse to bilobed (sometimes with a pair of small supplementary
lobules) at the apex, 22-32 mm. long from the base to either side of the bilobed
apex, 10-13.5 mm. wide across the lateral lobes and 23-41 mm. wide across the
mid-lobe; lateral lobes small, auriculate-spreading, suborbicular, passing into a
broad, subquadrate claw; mid-lobe many times larger, cordate-reniform, bilobed;
disc between the lateral lobes with a more or less plurituberculate callus which
terminates in a triangular point flanked on each side by a small, semiobovate
lamella. Column small, stout, about 5.5-6.5 mm. high, above with a pair of short,
broad, subdolabriform wings which are more or less crenulate.
Amazonas, fide Schlechter, Fedde Repert. Beih. 9: 175. 1921.—
Cuzco: Prov. of Calca, Lares Valley, between Huallhuayoj and Calca,
1900 meters, in "shrubwood," Weberbauer 7922. Prov. of Urubamba,
Machu-Picchu, 2000 meters, on cliffs, Vargas 1892. Also Colombia
(type of 0. globuliferum), Venezuela (types of 0. scansor and 0. con-
volvulaceum), Bolivia, fide Schlechter, Fedde Repert. Beih. 10: 78.
1922, Costa Rica (types of 0. globuliferum var. costaricense and
0. Wercklei Schltr.), and Panama.
SCHWEINFURTH: ORCHIDS OF PERU 877
Oncidium glossomystax Reichb. f. Bot. Zeit. 10: 696. 1852;
Walp. Ann. 6: 714. 1863; Cogn. Martius Fl. Bras. 3, pt. 6: 432, t. 62,
fig. 2. 1906; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 98. 1922.
Plant very small, epiphytic, with an abbreviated rhizome, consisting of a sub-
orbicular-flabelliform cluster of leaves whose bases are imbricating. Leaves more
or less numerous, ensiform, equitant, linear or oblong, often lightly oblique, acute or
subacute, 1 to rarely 4 cm. long, 2 to rarely 6 mm. wide. Scapes usually somewhat
exceeding the leaves, simple, commonly very slender, about 2-3 (rarely 5 cm. long),
erect to arcuate, 1- to 3-flowered above. Floral bracts minute, ovate-triangular,
concave, much shorter than the pedicellate ovary which is 8 mm. or less long.
Flowers large for the plant, yellow, with wide-spreading, membranaceous segments.
Dorsal sepal suborbicular-obovate or broadly elliptic, rounded at the apex, dor-
sally carinate with the keel produced into a mucro, 2.3-4 mm. long, 1.6-2.3 mm.
wide. Lateral sepals free, obliquely elliptic-lanceolate, acute, with the dorsal keel
extended into a mucro, about as long as the dorsal sepal and nearly half as wide.
Petals very broadly oblong or narrowly obovate, obliquely rounded or truncate at
the apex, about as long as the dorsal sepal and slightly narrower (often a little
broader). Lip much larger than the other segments, suborbicular in outline,
sharply 3-lobed, deeply bilobed in front, about 9.5-13 mm. long from the base to
the apex of an apical lobule, about equally broad across the mid-lobe in front and
across the lateral lobes at the base; lateral lobes relatively small, wide-spreading,
obliquely spatulate, abruptly round-dilated from a narrow claw, irregularly crenate
without; mid-lobe much larger, abruptly dilated from a broad subquadrate claw,
cordate-reniform, deeply bilobed in front and sharply indented on each side (thus
4-lobed) with the apical lobules oblong-suborbicular and the outer lobules much
larger and obliquely semiorbicular; disc near the base with 2 pairs of broadly
oblong lamellae terminating in deep fringes (one pair in front of the other) the
apical pair more or less coalescent. Column minute, porrectly dilated on each side
below, with a pair of broadly dolabriform, crenulate wings above.
Peru: Habitat not recorded, Mathews s.n., fide Cogn. Tocache, in
dry places, Poeppig 857.— Cuzco: Prov. of Paucartambo, Victoria,
Asuncion, 700-800 meters, Vargas 5151.— Junin: Puerto Bermudez,
about 395 meters, epiphyte in dense forest, Kittip & Smith 2651 9a.
— Loreto: Near Moyobamba, 800-900 meters, Weberbauer 4511 —
Madre de Dios: Iberia, Rio Tahuamana, 330 meters, Seibert 1947.
—San Martin: Near Moyobamba, 860 meters, Scolnik 1206. Also
Mexico, Guatemala, Nicaragua, Colombia, Venezuela, French Gui-
ana, Brazil, and Ecuador (type).
Oncidium gracillimum Rolfe, Kew Bull. (1898) 197.
Plant large, but not showy. Pseudobulb oblong-ovoid, subcompressed, 7.6 cm.
long, bifoliate. Leaves linear-oblanceolate, acute, gradually long-narrowed below
to a slender petioled base, up to 20.3 cm. long, about 2 cm. wide. Scape large,
terminating in a spreading, loosely many-branched panicle, about 90 cm. high, the
slender fractiflex branches 15-22.5 cm. long. Bracts oblong-lanceolate, concave,
much shorter than the slender pedicellate ovary which is 1.1 cm. or less long.
878 FIELDIANA: BOTANY, VOLUME 30
Flowers small, having wide-spreading segments, yellow with a few large brown
spots at the base of the perianth-members. Dorsal sepal cuneate-spatulate or
oblanceolate, subobtuse, about 6 mm. long, 2 mm. wide. Lateral sepals similar
but a little longer and narrower, lightly oblique. Petals "spatulate-oblong" or
broadly oblong from a clawed base, obtuse, apparently a little shorter and wider
than the dorsal sepal. Lip larger than the other segments, deeply 3-lobed with the
mid-lobe bilobed, spreading from the base of the column, lightly reflexed above,
about 8 mm. long from the base to the apex of a terminal lobule, nearly 7.5 mm.
wide at the base across the lateral lobes; lateral lobes divaricate or lightly retrorse,
broadly oblong, rounded at the apex; mid-lobe larger, oblong or more usually obo-
vate-oblong, somewhat larger than the lateral lobes, bifid with ovate-rounded,
slightly spreading lobules; disc at the base with a 6-lobed crest. Column sigmoid
in lateral view, about 4 mm. high, with an elongate beaked rostellum, bearing a
pair of oblong, unequally 3-denticulate wings, the posterior tooth falcate-recurved.
Peru: Habitat not recorded; imported by Messrs. F. Sander &
Company.
No example of this species was available, but the Ames Her-
barium contains a photograph of the type.
Oncidium heteranthum Poepp. & Endl. Nov. Gen. ac Sp. 1:
34, t. 60. 1836; Cogn. Martius Fl. Bras. 3, pt. 6: 378, t. 87. 1905;
Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 175. 1922. Oncidium
bryolophotum Reichb. f. Card. Chron. (1871) 738. Oncidium inops
Cogn. & Rolfe, Journ. des Orch. 3: 346. 1893 and 4: 74. 1893. On-
cidium megalous Schltr. Fedde Repert. 9: 30. 1911.
Plant small to large, extremely variable throughout, with an abbreviated rhi-
zome. Pseudobulbs aggregated, ovoid or ellipsoid to pyriform or cylindric in the
dried specimen, 1- or more- (commonly 2-) leaved, up to 6.5 (rarely 8.5) cm. long,
angulate, complanate or ancipitous, surrounded at the base by 1 to 3 pairs of
evanescent, conduplicate sheaths of which the upper ones are leaf-bearing. Leaves
elliptic or elliptic-oblong to linear-oblong (rarely obovate-oblong), acute or acumi-
nate, more or less narrowed to a complicate, petioled base, up to 34.5 cm. long
(commonly much shorter), 8 mm.-3.4 cm. wide, the largest and most persistent
on the pseudobulb. Scape usually much exceeding the leaves, up to 120 cm. long,
often flexuous or arcuate above; peduncle (often very short) up to 25 cm. long,
with a very few close, remote, short sheaths; inflorescence loosely paniculate with
numerous short or rather short, compound branches having a fractiflex rachis.
Floral bracts minute, triangular-lanceolate, much shorter than the pedicellate
ovary of the fertile flowers which is 1.5 cm. or less long. Flowers of 2 kinds; sterile
flowers much more numerous, smaller (often much so), of 3-6 linear to oblanceo-
late segments, one of which is sometimes partially developed into a lip; fertile
flowers medium-sized for the genus, with wide-spreading segments, yellow with
brown blotches or bars on the lower portion of the perianth parts. Dorsal sepal
narrowly oblanceolate or spatulate-oblanceolate, rounded or obtuse and often
apiculate at the apex, 7.2-8.8 (rarely 10) mm. long, 2-2.6 (rarely 3) mm. wide.
Lateral sepals free, sometimes a little shorter or longer than the dorsal sepal but
about equally broad, very obliquely oblanceolate-spatulate, subacute to apiculate.
SCHWEINFURTH: ORCHIDS OF PERU 879
Petals obovate-oblong, distinctly broader than the sepals, oblique especially at the
rounded to subtruncate-retuse apex. Lip much larger than the other segments,
sessile, pandurate-trilobed, with a bilobed apex, 9.5-12.5 mm. long from the base
to the tip of an apical lobule, 9-10 (rarely 11) mm. wide near the base and 9 (rarely
7) -13 mm wide near the apex, rather abruptly narrowed near the middle or above
the middle; basal part more or less semiorbicular with the dorsal margin cordate
and often tabulate, about equaling or distinctly larger than the antenor part (mid-
lobe); mid-lobe sessile to very shortly clawed, reniform, deeply bilobed; disc at the
base with a multilobulate callus terminating in 3 usually recurved lobules and
often surrounded by 1 or more separate papillae. Column very short, 5-6 mm.
high at the back including the erect wings, somewhat sigmoid when viewed from
the side, dilated upward, provided with a pair of spreading, more or less retuse,
fleshy dilations below and with a pair of transversely oblong wings above which
are commonly recurved and rounded at the apex, but sometimes angular and acu
or acuminate above.
Eastern Peru: Habitat not recorded, Mathews 1917—Cuzco:
Near Rio Yanamayo, below Pillahuata, 2000-2300 meters, epiphyte
in forest, Pennell 14051, 14078. Prov. of Convention, Hda. Potrero,
Sapan-Sachayocc, 2200 meters, epiphyte in forest, Vargas 2531.—
Huanuco: Near Cuchero (Cochero) and Pampayaco (Pampayacu),
Poeppig 1619 (type of 0. heteranthum) , Kanehira 333.— Junin:
Chanchamayo Valley, 1500 meters, Schunke s.n. Prov. of Tarma,
Utcuyacu, 1900 meters, common on trees, Woytkowski 35359. Also
Colombia (type of 0. inops), Venezuela, and Bolivia in South Amer-
ica and Costa Rica (type of 0. bryolophotum, 0. megalous) and Pan-
ama in Central America.
Oncidium hians Lindl. Bot. Reg. 24, Misc. 65, no. 124. 1838;
Fol Orch. Oncidium 36, no. 122. 1855; Reichb. f. Walp. Ann. 6: 775.
1863; Regel, Gartenfl. 36: 345, 1. 1250, figs, a, b. 1887; Cogn. Martius
Fl. Bras. 3, pt. 6: 416. 1906; Kranzl. Engler, Pflanzenr. IV. 50
(Heft 80), 155. 1922. Oncidium leucostomum Hoffmgg. Verz. Orch.
58. 1843.' Oncidium quadricorne Kl. Allg. Gartenz. 20: 249. 1852.
Oncidium maxilligerum Lem. Illustr. Hortic. 3, Misc. 43, fig., 1856.
Plant small, inconspicuous. Pseudobulbs very small, ovoid, more or less com-
pressed, unifoliate, 1-1.5 cm. high. Leaf small, oblong or linear-oblong, subobtuse
to abruptly acute or acuminate, long-attenuate to a complicate base, 3-7 cm. long,
8-13 mm. wide. Scape much exceeding the leaves, very slender, erect or ascend-
ing, 12-25 cm. long; peduncle provided with a few very remote, short sheaths;
inflorescence simple or slightly branched, with the branches erect-spreading, short,
filiform, loosely 2- to 4-flowered. Floral bracts very small, ovate-triangular, much
shorter than the filiform pedicellate ovary which is 8-13 mm. long. Flowers very
small, with wide-spreading segments, dull yellowish brown. Dorsal sepal oval
("obovate"), rounded at the apex, 5-5.5 mm. long, 3.2 mm. or less wide. Lateral
sepals free, very similar to the dorsal sepal but slightly smaller, oblique. Petals
880 FIELDIANA: BOTANY, VOLUME 30
broadly oblong or obovate, oblique especially at the rounded apex, distinctly
smaller than the dorsal sepal. Lip much surpassing the sepals and petals, pan-
durate, obovate-subquadrate in outline, sessile at the subcordate base, more or
less retuse at the apex, 6-8 (sometimes 10) mm. long, 2.5-3.5 mm. wide near the
base, 4-4.5 mm. wide near the apex; disc at the base with a large, fleshy, erect callus
which is bilobed at the apex and prominently dentate on each side (thus 4-lobed).
Column very short and stout, about 2 mm. high, with a pair of conspicuous, semi-
ovate, falcate-ascending wings.
Peru(?), fide Schlechter, Fedde Repert. Beih. 9: 175. 1921. Brazil
(type).
Oncidium hieroglyphicum Reichb. f. Allg. Gartenz. 22: 274.
1854; Lindl. Fol. Orch. Oncidium 29, no. 94. 1855; Reichb. f. Walp.
Ann. 6: 745. 1863; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 240.
1922.
Plant apparently large. Pseudobulbs caespitose, oval or ovoid to disc-shaped,
sharply ancipitous, unifoliate. Leaf oblong or ligulate, obtusely acute, up to
11.5 cm. long and 5 cm. wide, undulate. Raceme simple or with a few very short
(apparently 2-flowered) branches below, very loosely several-flowered. Floral
bracts minute, lanceolate, acuminate, much shorter than the pedicellate ovary
which is apparently about 20 mm. long. Flowers medium-sized for the genus,
with wide-spreading segments, the sepals and petals greenish-yellow with dull
brown spots except near the apex, and the lip pale yellow with a large brown spot
near the base. Sepals free, about equal, elliptic-lanceolate ("oblong"), acuminate
("acute"), undulate on the margins, about 12 mm. or more long, 5-6 mm. wide.
Petals similar but definitely broader. Lip longer and much broader than the sepals
and petals, from a short claw abruptly obovate-subquadrate in outline, sharply
pandurate-trilobed, cordate at the base, bilobed in front, about 15 mm. long from
the base to the apex of an apical lobule and slightly broader in front; lateral lobes
horizontally spreading, oblong-obovate or saddle-shaped, rounded without; mid-
lobe abruptly spreading from a conspicuous narrow claw, cordate-reniform, bi-
lobed; disc at the base with an oval, downy, 7-lobed crest. Column very short,
rhombic-dilated on each side below, having 3 pits below the stigma, with an apical
pair of semiovate, retuse wings, the upper lobe much smaller than the lower.
Peru: Habitat not recorded, Warscewicz s.n. (living original in
Herr Jenisch's garden).
No specimen of this species was available, but the Ames Her-
barium contains several sketches of the plant and flower from the
Reichenbach Herbarium.
Oncidium hirundo Reichb. f. Bot. Zeit. 15: 158. 1857; Kranzl.
Engler, Pflanzenr. IV. 50 (Heft 80), 216, fig. 19B, a-b. 1922.
Plant large. Pseudobulbs and leaves not recorded. Scape high, more than
65 cm. in length, loosely paniculate above; panicle about 30 cm. or more long, very
loosely several-flowered; floral branches 5 to 6 cm. apart, short, laxly 2- to 4-flow-
ered almost to the base, with a fractiflex rachis. Floral bracts ovate-lanceolate,
SCHWEINFURTH: ORCHIDS OF PERU 881
acute or acuminate, much shorter than the slender pedicellate ovary which is ap-
parently 20-25 mm. long. Flowers medium-sized, with wide-spreading segments,
sepals and petals yellow marked with chestnut-brown throughout, lip yellow with
chestnut spots below. Dorsal sepal elliptic-oblanceolate, relatively short-clawed,
acute, 16-20 mm. long, about 5-7 mm. wide. Lateral sepals free, obliquely oblan-
ceolate, acuminate, diverging like a swallow's tail, apparently much longer (with
longer claw) and somewhat narrower than the dorsal sepal. Petals oblong-lanceo-
late ("oblong"), obtuse, slightly subhastate below, markedly shorter and some-
what wider than the dorsal sepal. Lip much shorter than the sepals, subquadrate-
obovate in outline, pandurate-trilobed near the middle, sessile, emarginate in front,
about 13 mm. or more long from the base to an apical lobule and 10 mm. or more
wide in front (across the mid-lobe) ; basal portion rhombic, with rounded-triangular,
spreading lateral lobes, about 8 mm. wide; mid-lobe abruptly dilated from a short
claw into a reniform lamina; disc at the base with a suborbicular, pluridentate crest.
Column very small, with a dentiform process in front below and with a pair of
retuse wings above.
Peru: Habitat not recorded, Warscewicz s.n., fide Kranzl. (I.e.).
The Ames Herbarium contains a drawing of the scape with a
floral analysis from the Reichenbach Herbarium, but no material
was available.
Oncidium incarum (Kranzl.) C. Schweinf. Bot. Mus. Lean1.
Harvard Univ. 14: 70. 1949. Cyrtochilum incarum Kranzl. Engler,
Pflanzenr. IV. 50 (Heft 80), 58. 1922.
Plant large. Rhizome almost 2 m. long, with very long roots. Pseudobulbs
very distant, narrowly ovoid, compressed, bifoliate, up to 15 cm. long, clothed be-
low by several long, leaf-bearing sheaths. Leaves linear, acuminate, shortly com-
plicate below, plicate, 20-60 cm. long, 3 cm. or more wide. Scape up to 4 m. long,
climbing; peduncle 2.5 m. long; panicle about 1.5 m. long, with elongate branches
which are slender, flexuous and several- to many-flowered. Bracts broadly cymbi-
form, ovate, subacute, about a third as long as the pedicellate ovary which is
almost 3 cm. long. Flowers large for the genus. Sepals yellowish-brown. Dor-
sal sepal spatulate-oblong, long-clawed, 3 cm. long; lamina oblong, obtuse, about
2 cm. long and 1.5 cm. wide; claw 1 cm. long. Lateral sepals connate at the very
base, rather shortly clawed, about as long as the dorsal sepal; lamina oblong, sub-
acute, as wide as the dorsal sepal. Petals shortly and broadly clawed, markedly
shorter than the sepals; lamina ovate, subcordate-retuse at the base, acute, slightly
narrower than the sepals, the basal third yellow-brown, the upper part first white,
then violet. Lip deflexed, much shorter than the other segments, about 1.6 cm.
long and 1.3 cm. wide, from a broad base suborbicular or very broadly rhombic
with obtuse tip, apiculate in front, brown- violet; disc at the base with a crest con-
sisting of irregularly placed tubercles having 5 divergent teeth in front. Column
slender, linear, strict, 6-7 mm. high, capitate above, with a pair of very narrow
wings.
Peru: Huacapistana, Weberbauer 2136.
No example of this species was available.
882 FIELDIANA: BOTANY, VOLUME 30
Oncidium ionopterum Reichb. f . Linnaea 41 : 20. 1876. Cyrto-
chilum ionopterum Kranzl. Notizbl. Bot. Gart. Berl. 7: 97, in key.
1917, nomen; Engler, Pflanzenr. IV. 50 (Heft 80), 70, fig. 7C, a-d.
1922.
Plant delicate, probably elongate. Pseudobulbs and leaves not described, the
former said to be small. Peduncle and rachis of the inflorescence lacking. Panicle
slender, climbing, the existing part about 40 cm. long; branches 4-8 cm. distant,
flaccid, up to 4 cm. long, provided near the base with abortive flowers and at the
apex with 2 or 3 normal fertile flowers, the rachis becoming fractiflex. Bracts
minute. Pedicellate ovary of the normal flowers up to 10 mm. long. Abortive
flowers composed of 5 linear segments about 2.5 mm. long. Color of normal
flowers: sepals yellow-green, petals brown with a white apex, lip violet on each
side, brown in the center and with a white tip in front. Dorsal sepal oblong-cuneate
or oblong-oblanceolate with a long claw, acute, 7 mm. long, 2 mm. wide. Lateral
sepals free, oblong-oblanceolate, with a long claw, oblique, acute, distinctly longer
than the dorsal sepal but equally wide. Petals oblong-obovate from a narrow-
cuneate base, acute, oblique, a little shorter and notably broader than the dorsal
sepal. Lip simple, sessile, broadly triangular-ovate, abruptly acute in front, con-
vex, with the rounded sides denticulate or crenulate, apparently subtruncate at
the base, about 5 mm. long and 8 mm. wide; disc velutinous, with a large central
crest consisting of several finger-like tubercles, terminating in a pair of larger,
elongate, outcurved tubercles in front and with a separate, bilobed papilla on
either side. Column small, with an angular plate on either side below and a pair
of large transversely oblong or semiorbicular, lobulate wings above.
Piura: Near Huancabamba, 3076 meters, Wallis s.n. (type).—
Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 175. 1921.
No material of this species was available, but the Ames Herbar-
ium contains drawings of the type from the Reichenbach Herbarium.
Oncidium Kienastianum Reichb. f. Card. Chron. n.s. 9: 558.
1878. Oncidium trilingue Sander, Orch. Guide 193. 1901, not Lindl.
Cyrtochilum Kienastianum Kranzl. Notizbl. Bot. Gart. Berl. 7: 91,
in key. 1917, nomen; Engler, Pflanzenr. IV. 50 (Heft 80), 33, fig. ID,
0-6. 1922.
Plant large. Pseudobulbs aggregated according to Kranzlin, oblong-ovoid or
ellipsoid, more or less ancipitous, bifoliate, about 8 cm. high, clothed at the base
by several pairs of conduplicate, leaf-bearing sheaths. Leaves oblanceolate-oblong
("linear"), acute or acuminate, more than 50 cm. long and 2.5 cm. wide according
to Kranzlin. Scape very long, twining, paniculate above, the short, few-flowered
branches described as 3-4 cm. long. Bracts small, oblong-ovate, acute, more than
a third as long as the pedicellate ovary which is about 3.5 cm. long. Flowers rather
large, with wide-spreading segments. Sepals brown with pale yellow margins, at
length becoming dark purple, crisped. Dorsal sepal clawed, 1.5 cm. long; lamina
triangular-ovate, acute, about 1 cm. wide; claw relatively short. Lateral sepals
longer-clawed, distinctly longer than the dorsal sepal and about as wide, cuneate-
oblong, acute. Petals shortly and broadly clawed, as long as the dorsal sepal and
SCHWEINFURTH: ORCHIDS OF PERU 883
nearly as wide; lamina ovate, acuminate ("acute"), with crisped margins and a
reflexed apex, yellowish with brown spots or bars, at length purple. Lip triangular-
lanceolate in outline, lightly 3-lobed, about 1.5 cm. long and 1.2 cm. wide across the
lateral lobes near the base, yellowish with brown or purple spots at the base and
on the margins; lateral lobes relatively small, subquadrate-auriculate, spreading,
sometimes divided in the middle; mid-lobe much larger, "triangular" or triangular-
ovate, acute or acuminate, recurved above, with the margins strongly crisped-
undulate; disc at the base with a 3-carinate crest, the central keel highest, the lat-
eral ones semilunate-curved with 3 lamellae on each side. Column short and stout,
sigmoid when viewed from the side, purple, with a pair of angular, fleshy processes
below and a pair of minute, ascending, curved wings above.
Northern Peru or Colombia, habitat not recorded, Roezl s.n.
No example of this species was available, but in the Ames Her-
barium there are sketches and floral analyses from the Reichenbach
Herbarium.
Oncidium leopardinum Lindl. Fol. Orch. Oncidium 28, no. 93.
1855; Williams, Orch.-Grow. Man. ed. 7: 618. 1894.
Plant apparently large, with no record of the pseudobulbs, leaves or complete
scape. Panicle very loose, about 90 cm. high (according to Williams), the branches
often very remote, apparently short, compound and with a strongly fractiflex ra-
chis. Floral bracts minute, much shorter than the pedicellate ovary which is about
20 mm. or less long. Flowers rather small, with wide-spreading or reflexed seg-
ments, yellow with distinct, dark brown bands on the sepals, petals and base of the
lip. Sepals apparently oblong and free, the lateral ones (which alone show clearly
in the photograph of the type) narrowly oblong, acute or obtuse, about 9 mm. long
and 2 mm. wide. Petals markedly broader than the lateral sepals, ovate-oblong
from a short, subunguiculate base, apiculate at the broadly obtuse to retuse apex.
Lip nearly equaling the lateral sepals, about 9 mm. long, subequally wide in front
and slightly narrower near the base, pandurate-trilobed, sessile; lateral (basal)
lobes small, auriculate-rounded; mid-lobe cordate-reniform, bilobed in front, sep-
arated from the basal portion by a rather long, abrupt claw; disc at the base with
a multilobulate callus which terminates in 5 distinct tubercles the three middle
ones longer, with a longer outcurved tubercle on each side in front. Column very
short and stout, with a terminal pair of large wings which are broadly dolabriform,
lightly 3-lobulate and falcate at the summit.
Peru : Sources of the Maranon River, Warscewicz s.n.
No specimens of this species were available, but the Ames Her-
barium contains sketches from the Reichenbach Herbarium.
Oncidium Leopoldianum Rolfe, Gard. Chron. ser. 3, 8: 556.
1890; Lindenia 6: 71, t. 274. 1891; Williams, Orch.-Grow. Man. ed.
7: 618. 1894. Cyrtochilum Leopoldianum Kranzl. Notizbl. Bot. Gart.
Berl. 7: 94, in key. 1917, nomen; Engler, Pflanzenr. IV. 50 (Heft 80),
55, fig. 5B, a-6. 1922.
884 FIELDIANA: BOTANY, VOLUME 30
Plant large and showy. Rhizome long-creeping, stout according to Kranzlin.
Pseudobulbs more or less distant, about 12.7 cm. or less long, narrowly oblong-
ovoid to ovoid-cylindric, 1- or 2-leaved, clothed at the base by several pairs of
sheaths which are mostly leaf-bearing. Leaves on the sheaths larger than those
on the pseudobulbs (according to Kranzlin), oblanceolate, gradually narrowed be-
low, acute or acuminate, up to 20 cm. long and 3 cm. wide, those on the pseudo-
bulbs oblanceolate ("lanceolate"), up to 16 cm. long and 2 cm. wide according to
Kranzlin. Scapes elongate, climbing, panicled above, many-branched, the existing
portion about 2.5 m. long, clothed at the base by several leaf -bearing sheaths, the
branches about equally long and up to 12 cm. in length. Bracts small, broadly
ovate, about one fourth as long as the pedicellate ovary which is 2.5 cm. long.
Flowers rather large, with wide-spreading segments, about 4.5 cm. across; sepals
and petals white with a crimson-purple lower central portion, lip violet-purple with
the basal crest and part of the column yellow. Dorsal sepal very shortly clawed,
exauriculate at the base, about 2.3 cm. long; lamina broadly elliptical (or oval),
obtuse, about 1.1 cm. wide. Lateral sepals very similar, somewhat broader than
the dorsal sepal, or narrower according to Kranzlin. Petals elliptic-ovate, sub-
sessile, subacute, shorter and narrower than the sepals, or broader according to
Kranzlin. Lip much smaller than the sepals and petals, triangular-lanceolate,
lightly 3-lobed near the base, fleshy, about 1.3-1.5 cm. long, 7-10 mm. wide across
the lateral (basal) lobes, subacute to acuminate; lateral lobes rounded; disc at the
base with a crest consisting of 3-5 short, fleshy, parallel ridges, the central one
highest. Column short, with a pair of short, rounded, fleshy wings.
South America: Habitat and collector not recorded; introduced
by Messrs. Linden of Brussels (type). — Loreto: Mountains east of
Moyobamba, 1300-1400 meters, transitional country between the
savanna woods and the upland evergreen woods, Weberbauer 4732.
No material of this species was available.
Oncidium lineoligerum Reichb. f. & Warsc. Bonpl. 2: 109.
1854; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 280. 1922.
Plant large. Pseudobulbs and leaves not recorded. Scape incomplete in the
type specimen, loosely panicled above and then racemose near the apex; panicle
stout, about 73 cm. long including the terminal racemose portion; branches 4-6 cm.
apart, loosely 2- to 3-flowered, with a fractiflex rachis. Bracts lanceolate-ovate,
cucullate, much shorter than the slender pedicellate ovary which appears to be
about 3 cm. long. Flowers rather small, with wide-spreading or reflexed segments,
yellow or orange with an irregular brown band across the middle of the lip. Dorsal
sepal oblong or ligulate, obtusely acute, cuneate at the base, undulate, about
16 mm. long and 5 mm. wide according to Kranzlin. Lateral sepals similar but
slightly larger, free. Petals about as long as the dorsal sepal, elliptic, obtuse, very
shortly clawed below, slightly wider than the lateral sepals according to Kranzlin.
Lip subquadrate in outline, pandurate-trilobed, about 18 mm. long, sessile at the
subtruncate base, bilobed in front; lateral (basal) lobes relatively small, triangular-
auriculiform, spreading, rounded without, rather abruptly narrowed into a prom-
inent, subquadrate isthmus; mid-lobe cordate-reniform, retuse to bilobed, about
22 mm. wide; disc with a basal crest which is divided into a depressed, plurisulcate
SCHWEINFURTH: ORCHIDS OF PERU 885
lower portion and an anterior rhombic-ovate callus which is rostrate in the center,
with 3 to 5 sharp teeth on each side. Column small, above with a pair of short,
broad, repand or unequally bilobed wings.
Peru: Habitat not recorded, Warscewicz s.n. (type). — Amazonas:
Near Chachapoyas, Oversluys s.n.
A sketch of this species from the Reichenbach Herbarium is in
the Ames Herbarium.
Oncidium Lucasianum Rolfe, Kew Bull. (1894) 185; Sander,
Orch. Guide 185. 1901; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80),
141. 1922.
Plant large. Pseudobulbs ovoid, sub compressed, bifoliate, 2.5-5 cm. high.
Leaves linear-lanceolate, acute, 5-8 cm. long. Scape arcuate, several-jointed with
swollen joints according to Kranzlin, few-flowered and 15-20 cm. long, or many-
flowered, paniculate and 65-120 cm. long according to Sander. Bracts small, tri-
angular, acute, much shorter than the pedicellate ovary which is up to 2 cm. long.
Flowers of variable structure, rather large, about 3 cm. across, entirely yellow.
Dorsal sepal obovate-oblong from a very narrow base, obtuse, dorsally carinate,
undulate, about 17 mm. or more long, 8 mm. or less wide. Lateral sepals similar,
subfree or connate to the middle. Petals broadly oblong from a shorter claw, ob-
tuse, about as long as the sepals but somewhat wider. Lip pandurate-trilobed,
about 20 mm. long and 21 mm. or more wide; lateral lobes small, obovate-oblong,
auriculiform; mid-lobe large, suborbicular-reniform, bilobed, cordate at the base,
about 25 mm. wide; disc with a 5-lamellate callus of which the lamellae are verru-
cose and are surrounded by warts. Column short, with a pair of rather large, semi-
ovate, erose wings.
Habitat originally uncertain. Peru : Without locality, fide Sander
and Kranzlin.
No example of this species was accessible.
Oncidium luridum Lindl. Bot. Reg. 9: t. 727. 1823; Hook. Bot.
Mag. 64 : t. 3603. 1837. Oncidium luridum var. guttatum Lindl. Bot.
Reg. 25: t. 16. 1839, not Epidendrum guttatum L. Oncidium gut-
tatum Reichb. f. Walp. Ann. 6: 782. 1863; Cogn. Martius Fl. Bras. 3,
pt. 6: 411, t. 83. 1906. Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80),
110. 1922. Figure 173.
Plant large, extremely variable and widespread. Rhizome very short. Pseudo-
bulbs minute or small, ovoid, unifoliate, more or less compressed, rarely up to
3.5 cm. high, entirely concealed and surpassed by several imbricating, scarious
sheaths. Leaf very variable, oval to elliptic-oblong or oblong, acute, more or less
narrowed to a complicate base, very thick and fleshy, 15 (rarely smaller) -77 cm.
long, 2.2 (rarely) -13 cm. wide. Scape much longer than the leaves, erect or nod-
ding, loosely paniculate and many-flowered above, 30-180 cm. long; peduncle stout,
about 9 cm. (rarely) or more long, provided with remote, short, close, tubular
sheaths; panicle large, with remote branches which are short to elongate, simple
886 FIELDIANA: BOTANY, VOLUME 30
or compound, with a more or less fractiflex rachis. Floral bracts very short, ovate-
triangular, acute, much shorter than the slender, pedicellate ovary which is up to
3.5 cm. long. Flowers medium-sized, with wide-spreading segments having undu-
late-crisped margins, usually dull yellowish or greenish, thickly spotted with brown.
Dorsal sepal broadly spatulate or obovate-spatulate, rounded at the apex, grad-
ually unguiculate-narrowed below, 12-19 mm. long, 7.5-12 mm. wide. Lateral
sepals free, similar to the dorsal sepal but a little narrower and often longer. Petals
oblong-obovate with a short or very short broad claw, a little shorter and narrower
than the dorsal sepal. Lip slightly longer and much broader than the other seg-
ments, suborbicular-obovate in outline, pandurate-trilobed, sessile and subcordate
at the base, retuse to bilobed above, 14.6-21 mm. long from the base to the tip
of an apical lobule, about 7.8-10 mm. wide near the base and 15-21 mm. wide in
front (across the mid-lobe); basal portion ovate-subquadrate with an auriculate,
spreading base; anterior portion (mid-lobe) cordate-reniform, abruptly dilated
from a scarcely perceptible claw; disc at the base with a large plurituberculate
callus having 2 larger, more or less oblong, irregular tubercles below and 3 smaller
ones in front. Column very short and stout, 4-5 mm. high, with a pair of trans-
versely oblong-subquadrate, unequally bilobed wings above.
Peru: Habitat not recorded, Pavdn s.n. Also Florida (U.S.A.),
the West Indies, Mexico and Central America, South America (type),
Colombia, Venezuela, British Guiana and Brazil.
Oncidium macranthum Lindl. Gen. & Sp. Orch. 205. 1833;
Fol. Orch. Oncidium 4, no. 1. 1855; Hook. f. Bot. Mag. 94: t. 5743.
1868. Cyrtochilum macranthum Kranzl. Notizbl. Bot. Gart. Berl. 7:
95, in key. 1917, nomen; Engler, Pflanzenr. IV. 50 (Heft 80), 55,
fig. 5C, a-c. 1922.
Plant large and showy. Pseudobulb ovoid to conic-cylindric, more or less com-
pressed, 7-15 cm. tall, bifoliate, clothed at the base with 3 or 4 pairs of imbricating,
conduplicate sheaths (progressively larger upward) of which the upper ones are leaf-
bearing. Leaves very variable in size but apparently larger on the pseudobulb,
oblanceolate or linear-oblanceolate to oblong, acute or short-acuminate, gradually
narrowed below (often much so) to a complicate base, mostly 25-52.5 cm. long
and 2.5-5 cm. wide above. Scape elongate, climbing, 60-300 cm. long, loosely
branching (paniculate) above; peduncle stout, with very remote, short, close, tubu-
lar sheaths; branches of the panicle very remote, about 33 cm. or less long, mostly
concealed below by tubular sheaths, loosely 2- to 5-flowered above. Floral bracts
large, cymbiform, obtuse to acute, about half as long as the slender pedicellate
ovary which is up to 7 cm. or more long. Flowers very large for the genus, with
wide-spreading segments, about 7-10 cm. across from the apex of the dorsal sepal
to the tip of a lateral sepal. Sepals dull yellowish brown, somewhat undulate on
the margins. Dorsal sepal about 3.5-4 cm. long, prominently clawed; lamina sub-
orbicular, rounded and slightly retuse at the apex, subcordate at the base, about
2.5-3 cm. long and 2.4-3.1 cm. wide, concave; claw narrow, auriculate on each side
at the base, about 7-10 mm. long. Lateral sepals distinctly longer, prominently
clawed; lamina oval or subquadrate-oval, broadly rounded or retuse at the apex,
narrowed to the claw, distinctly narrower than the dorsal sepal; claw similar to
that of the dorsal sepal, auriculate at the base without. Petals suborbicular-ovate,
FIG. 173. Oncidium luridum Lindl. Flowers from a many-flowered panicle; X 1.
887
888 FIELDIANA: BOTANY, VOLUME 30
very shortly and broadly clawed, rounded at the apex, strongly crisped-undulate,
distinctly shorter than the dorsal sepal but about equally wide, golden yellow.
Lip much smaller than the other segments, triangular in outline, spreading from
the base of the column and sharply reflexed near the deeply cordate base, acumi-
nate toward the recurved apex, subsimple with small, rounded and subacute, re-
trorse, basal (lateral) lobules, nearly 2.5 cm. long when expanded and 1.9 cm. wide
at the base, brownish purple on the sides and cream-color in the center; disc with
a stout, retrorse tubercle consisting of 3 basally confluent subequal lamellae and
in front with 3 smaller tubercles. Column short, stout, constricted in the middle,
about 1 cm. high, with a pair of small, fleshy wings which are flabellate-semiorbic-
ular and irregularly bilobed or retuse above.
Peru: Habitat not recorded, Pavon s.n., Mathews 1919. — Ama-
zonas: East of Chachapoyas between Tambos Bagazon and Almi-
rante, 2400 meters, Weberbauer 4441- Sources of the Maranon,
Warscewicz 131. Chuquiribamba, Andre 4498. Also Colombia and
Ecuador (type).
Oncidium macranthum Lindl. var. hastiferum (Reichb. f. &
Warsc.) C. Schweinf., comb. nov. Oncidium hastiferum Reichb. f.
& Warsc. Bonpl. 2: 102. 1854.
This concept, of which no example was available, differs from the species in
having at the base of the lip 1 large median keel flanked on each side by a smaller
prostrate lamella and in having a pair of narrow, ligulate wings on the column.
Peru(?), fide Schlechter, Fedde Repert. Beih. 9: 175. 1921.
Ecuador (type).
Oncidium Mathieuanum Reichb. f. & Warsc. Allg. Gartenz.
24: 321. 1856; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 260,
fig. 21E, 0-6. 1922.
Plant large. Pseudobulb oblong-ovoid, ancipitous, deeply sulcate in the dried
specimen, about 8 cm. high, obliquely bifoliate, adorned with violet lines and spots.
Leaves unknown. Scape about 90 cm. long, slender, flexuous to arcuate, paniculate
above; branches numerous, about 4.5-6 cm. distant, short, 1- to 2-flowered, about
3 cm. or less long. Floral bracts very minute. Pedicellate ovary up to 2 cm. long.
Flowers rather small, with wide-spreading segments having a cinnamon central and
lower portion with yellow margins. Sepals free, obovate, obtusely acute, cuneate
below, about 10 mm. long, 4.5-5 mm. wide. Petals markedly larger, undulate,
oblong and obtuse according to Kranzlin. Lip prominently 3-lobed, about 13 mm.
long and 8-9 mm. broad near the base (according to Kranzlin), rounded at the ses-
sile base; lateral lobes small, auriculiform, round-obovate; mid-lobe broadly cuneate
from a short, broad, semiorbicular claw, lightly retuse at the truncate apex; disc
between the lateral (basal) lobes with a high callus which is "ligulate," diver-
gently angled on each side at the base and tridentate in front with the lateral teeth
decurrent. Column medium-sized, with a pair of broadly dolabriform ("subtri-
angular") wings above.
SCHWEINFURTH: ORCHIDS OF PERU 889
Northern Peru: Habitat not recorded, Warscewicz s.n.
In the Ames Herbarium there is a drawing of this orchid with
floral analysis from the Reichenbach Herbarium.
Oncidium mentigerum (Reichb. f. & Warsc.) Lindl. Fol. Orch.
Oncidium 34, no. 115. 1855; Cogn. Martius Fl. Bras. 3, pt. 6: 371.
1905. Odontoglossum mentigerum Reichb. f. & Warsc. Bonpl. 2: 100.
1854. Cyrtochilum mentigerum Kranzl. Notizbl. Bot. Gart. Berl. 7 : 98,
in key. 1917, nomen; Kranzl. Engler, Pflanzenr. IV, 50 (Heft 80),
78. 1922.
Plant large. Pseudobulbs and leaves unknown in the type. According to
Kranzlin (I.e.), the pseudobulbs are numerous, aggregated and obliquely ovoid, and
the leaves are narrowly lanceolate or oblanceolate, acuminate, revolute on the
margins and up to 20 cm. long and 1.8 cm. wide. Scape robust, elongate, termi-
nated by an elongate panicle. Panicle loosely many-branched, pyramidal, about
54 cm. high; branches wide-spreading with a fractiflex rachis, about 5-15 cm.
long, the lower branches several-branched. Floral bracts minute, triangular-
lanceolate, very acute. Pedicellate ovary filiform-clavate, up to 12 mm. long.
Flowers very small, with wide-spreading membranaceous segments. Sepals and
petals ochraceous. Dorsal sepal oblanceolate, short-acuminate, long-narrowed be-
low, about 7 mm. long, 2-2.5 mm. wide. Lateral sepals free, very similar to the
dorsal sepal but somewhat longer and narrower, slightly oblique. Petals oblanceo-
late-obovate, acute, long-attenuate below, shorter than the dorsal sepal but about
as wide. Lip about half as long as the lateral sepals, simple, narrowly ovate, acute
or short-acuminate, rounded at the sessile base, lightly concave, yellow, 4.5-5 mm.
long, 2.5 mm. wide; disc through the lower half with 2 approximate, erect crests or
keels which are contiguous below and attenuate and slightly diverging above.
Column short, stout, about 3 mm. high, with a pair of short ("obsolete"), trian-
gular wings.
Peru: Sources of the Maranon River, Warscewicz s.n. (type of
Odontoglossum mentigerum). Also Ecuador and Colombia.
In the Ames Herbarium there is a photograph of the type from
the Lindley Herbarium.
Oncidium minax Reichb. f. Linnaea 41: 22. 1876. Cyrtochilum
minax Kranzl. Notizbl. Bot. Gart. Berl. 7: 91, in key. 1917, nomen;
Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 33, fig. 1C, a-d. 1922.
Plant apparently large. Pseudobulbs, leaves and scape not recorded. Scape
probably elongate and climbing. Branches (only 2 separate ones present in the
type) stout, 13-16 cm. long, apparently once-branched below, loosely several-
flowered. Floral bracts conspicuous, ovate, acute or acuminate, apiculate, deeply
concave. Pedicellate ovary slender, spreading, 3-3.5 cm. long, about twice as long
as the subtending bracts. Flowers rather large with spreading segments, appar-
ently brown with the paler petals barred. Dorsal sepal clawed, about 24 mm. long;
lamina broadly triangular-ovate, obtuse, cordate at the base, about 16 mm.
long and equally wide; claw slender, auriculate at the base. Lateral sepals mark-
890 FIELDIANA: BOTANY, VOLUME 30
edly longer than the dorsal sepal, longer-clawed; lamina oblong-ovate, subacute,
cuneate at the base, a little longer and narrower than that of the dorsal sepal.
Petals ovate-triangular, abruptly dilated from a short broad claw, crisped on the
margin, apparently definitely smaller than the sepals. Lip about 13 mm. long
according to Kranzlin, with a prominent narrow claw, deeply 3-partite; lateral
lobes erect, obliquely linear or oblanceolate-linear, acute; mid-lobe larger, oblong-
linear ("ligulate"), acute; disc at the base of the mid-lobe with 3 rhombic, crenate
keels. Column small, angled on each side at the base, with a pair of minute tri-
angular wings above.
Southern Peru: Habitat not recorded, Pearce s.n.
The Ames Herbarium contains a drawing of the type from the
Reichenbach Herbarium.
Oncidium nanum Lindl. Bot. Reg. 28, Misc. 37, no. 30. 1842;
Lindl. & Paxt. Paxt. Fl. Gard. 2: 8, fig. 130. 1851-52; Cogn. Martius
Fl. Bras. 3, pt. 6: 375. 1905; Kranzl. Engler, Pflanzenr. IV. 50
(Heft 80), 114. 1922. Oncidium patulum Schltr. Orchis 8: 18, t., 1914.
Plant small to medium-sized. Pseudobulbs caespitose, very small, thick-
cylindric, up to 7 mm. high, unifoliate, concealed by 1 or 2 pairs of small, imbri-
cating, evanescent sheaths. Leaf broadly elliptic to elliptic-oblong or oblong,
acute, more or less long-narrowed to a complicate, channelled, petiole-like base,
7-23.5 cm. or more long, 2- (rarely) 7 cm. wide. Scape shorter or longer than the
leaves, reflexed to arcuate-suberect, about 9-24 cm. long, loosely branched nearly
to the base; peduncle up to 8 cm. long, with about 3 very short, amplexicaul, con-
cave sheaths; panicle up to 14 cm. long, with several remote, short, simple branches,
the latter densely several- to many-flowered. Floral bracts ovate or ovate-lanceo-
late, concave, distinctly shorter than the slender pedicellate ovary which is 10 mm.
or less long. Flowers very small, with wide-spreading segments. Sepals and petals
yellow or orange, spotted with purplish brown or violet. Dorsal sepal spatulate
to obovate, broadly rounded at the apex, concave above, about 7-9 mm. long,
5-6.5 mm. wide. Lateral sepals similar but a little longer and narrower, free,
oblique, broadly obtuse. Petals obovate-oblong, oblique, broadly rounded and
retuse at the apex, slightly shorter than the dorsal sepal and more or less narrower
than either sepal. Lip when expanded slightly longer and much wider than the
sepals, suborbicular-obovate in outline, pandurate-trilobed, lightly retuse at the
apex, sessile at the base, about 8.6-9.5 mm. long from the base to the apex on either
side of the retuse middle; basal portion oblate-suborbicular when expanded with
revolute, semiorbicular sides (lateral lobes), 4-7.6 mm. wide when expanded; mid-
lobe transversely oblong or reniform, retuse, sessile or nearly so, about 6-10.5 mm.
wide, yellow; disc between the lateral lobes with a fleshy callus consisting of 2
approximate, suborbicular or triangular-ovoid lobes at the base and an oval-ovate,
bisulcate (tricarinate) lobe in front. Column very short and stout, about 2.5-
4 mm. high at the back, with a pair of small, obliquely triangular-dentiform wings.
Loreto: Mishuyacu, near Iquitos, 100 meters, epiphyte in forest,
King 1296, 10006. — San Martin: Juan Jui, Alto Rio Huallaga, about
400 meters, epiphyte in forest, Klug 3871. Also British Guiana
(type) and Brazil.
SCHWEINFURTH: ORCHIDS OF PERU 891
Oncidium nubigenum Lindl. Gen. & Sp. Orch. 197. 1833;
Gard. Chron. (1867) 376; Cogn. & Goos. Diet. Icon. Orch. Oncidium
t. 30. 1900; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 130. 1922.
Oncidium cucullatum Lindl. Sert. Orch. sub t. 21. 1838; Lindl. &
Paxt. Paxt. Fl. Gard. 3: t. 87. 1852-53; Lindl. Fol. Orch. Oncidium
22, no. 69. 1855; Cogn. &'Goos. Diet. Icon. Orch. Oncidium t. 23.
1899. Oncidium cucullatum var. nubigenum Lindl. Fol. Orch. On-
cidium 22, no. 69. 1855; Hook. f. Bot. Mag. 94: t. 5708. 1868.
Plant medium-sized to large, variable. Rhizome abbreviated. Pseudobulbs
aggregated, narrowly ovoid to pyriform-cylindric, somewhat compressed, 1- or
commonly 2-leaved, 3-10 cm. high, clothed below by about 2 pairs of imbricating
sheaths of which the upper ones are sometimes leaf-bearing. Leaves linear-elliptic
to narrowly oblong or oblanceolate, acute or acuminate, 5-26 (rarely 41) cm. long,
up to 1.5 (rarely 4) cm. wide. Scape slender, suberect or nodding, up to 25 (rarely
60 or more) cm. long, loosely racemose (rarely paniculate) above; peduncle up to
about 22.5 (rarely 53) cm. long, with a few (up to 6) remote, tubular, evanescent
sheaths; raceme (rarely compound or paniculate) loosely 3- to 10-flowered or each
branch few-flowered. Floral bracts minute, triangular-infundibuliform. Pedicellate
ovary 2-2.5 cm. long. Flowers rather small to medium-sized, with wide-spreading
segments. Sepals and petals of variable color, either white and purple spotted or
brownish or olive with yellow margins. Dorsal sepal oblong-oval or obovate-oblong
("oblong"), acute to truncate and abruptly acute at the apex, cucullate, about
7.2-15 mm. long, up to 8 mm. wide. Lateral sepals semiconnate or usually con-
nate into a suborbicular-obovate lamina which is slightly longer and distinctly
broader than the dorsal sepal, the free apices semiorbicular and apiculate. Petals
broadly or narrowly obovate, acute or obtuse at the apex, a little shorter than the
dorsal sepal but about as broad or slightly broader. Lip much larger, especially
wider, than the other segments, obovate-suborbicular in outline, pandurate-trilobed,
retuse in front, cordate at the sessile base, 16-18 mm. long from the base to the tip
of an apical lobule, yellowish white more or less spotted with rose, or crimson
spotted with purple; basal portion transversely oblong, 9-16 mm. wide, on each
side produced into a small, spreading, auriculiform lateral lobe; mid-lobe much
larger, broadly flabellate to reniform, retuse and more or less indented on each
side in front, cordate or subcuneate below, about 22 (rarely 28) mm. or less wide;
disc with a small, 3- or 5-lobulate callus at the base. Column very short and stout,
wingless above, about 3 mm. or less high at the back.
Peru: Habitat not recorded, Lobb s.n., according to Lindley. Also
Ecuador (type of 0. nubigenum and 0. cucullatum) and Colombia.
Oncidium obryzatum Reichb. f. & Warsc. Bonpl. 2: 108. 1854;
Lindl. Fol. Orch. Oncidium 29, no. 95. 1855; Regel, Gartenfl. 27:
t. 925. 1878; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 239, fig.
20B, a-d. 1922.
Plant medium-sized to large, variable, apparently without a long-creeping
rhizome. Pseudobulbs ovoid to cylindric-pyriform, strongly compressed, 1- or
2-leaved, about 4-8 cm. high, clothed at the base by 3 or 4 pairs of imbricating
892 FIELDIANA: BOTANY, VOLUME 30
sheaths which are progressively larger upward, the upper ones leaf-bearing. Leaves
very variable in size, linear-elliptic to linear-oblong, the largest ones 28-52.5 cm.
long and 2-2.8 cm. wide, acute, narrowed below to a conduplicate base. Scape
commonly elongate, about 24-150 cm. or more long, paniculate above; peduncle
about 16-27 cm. or much more in length, commonly stout, provided with several
remote, short, close sheaths; panicle about 8.5-100 cm. long, with 4 to numerous
branches which are 2-6.5 cm. distant, mostly short and loosely few-flowered with a
fractiflex rachis. Floral bracts rather small, ovate, concave, scarious. Flowers
small or medium-sized, with wide-spreading segments, yellow with reddish brown
spots or bars on the lower half. Dorsal sepal spatulate-cuneate or obovate-oblong,
abruptly acute, broadly rounded or subtruncate and retuse at the apex, about
11-14 mm. long, 4.2-6 mm. wide above. Lateral sepals similar but longer and
more or less narrower, oblique, free, obtuse to truncate-retuse at the apex. Petals
obovate-oblong, very similar to the dorsal sepal but more or less wider, and not so
long-attenuate below, oblique, undulate. Lip longer and much wider than the other
segments, subquadrate-obovate in outline, pandurate-trilobed, sessile at the sub-
cordate base, prominently bilobed in front, about 12.2-20 mm. long from the base
to the tip of an apical lobule, about 8-10 mm. wide at the base (across the
lateral lobes) and 11.9-18 mm. wide in front across the anterior lobe; basal portion
triangular-subquadrate, from the auriculate, spreading lateral lobes gradually
narrowed (clawed) to the mid-lobe; mid-lobe reniform, commonly much wider
than the basal part; disc with a small, many-lobed callus at the base. Column
small, sigmoid in side view, 4.1-6 mm. high with a pair of prominent, porrect, angu-
late processes below, and a pair of dolabriform to obliquely subquadrate (often
long-acuminate) wings above.
Peru: Habitat not recorded, Warscewicz s.n. (type). — Cajamarca,
Prov. of Cutervo, above Socota, on Socota River, 2800 meters, epi-
phyte, Stork & Horton 10152. Also Colombia, Venezuela, and Ecuador.
Oncidium ochmatochilum Reichb. f. Bot. Zeit. 10: 698. 1852;
Lindl. Fol. Orch. Oncidium 55, no. 193. 1855; Kranzl. Engler, Pflan-
zenr. IV. 50 (Heft 80), 219, fig. 19D, a-c. 1922. Oncidium cardio-
chilum Lindl. Fol. Orch. Oncidium 27, no. 87. 1855. Odontoglossum
cardiochilum Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 286, 288.
1922.
Plant large, stout, up to 2 m. or more tall, variable, without a long-creeping
rhizome. Pseudobulb complanate-cylindric or ovoid, bifoliate, about 6-12.5 cm.
tall, mostly enveloped by several (up to 5) pairs of imbricating, conduplicate
sheaths which are successively larger upward and mostly leaf-bearing. Leaves
oblong-linear or lorate to oblanceolate-oblong, acute or acuminate, more or less
narrowed below, up to 67.5 cm. long and 5.6 cm. wide (often much smaller).
Scape commonly large, stout, 45-240 cm. long, more or less erect and usually much
surpassing the leaves, paniculate above; peduncle 8-41 cm. or more long, com-
monly with remote, very short, close, tubular sheaths; panicle 37 cm. or much
more in length, loosely many-branched, with the branches short or elongate,
simple or compound, several- to many-flowered, with a very fractiflex rachis.
Floral bracts minute, concave, acute. Pedicellate ovary about 3.5 cm. or less
SCHWEINFURTH: ORCHIDS OF PERU 893
long. Flowers small, variable, with wide-spreading, subfleshy segments, fragrant,
green mottled with reddish brown, with white on the lip. Sepals and petals
recurved above. Dorsal sepal narrowly elliptic-lanceolate, acuminate, about
13-20 mm. long and 3-5 mm. wide. Lateral sepals obliquely linear-lanceolate or
linear-elliptic, acute or acuminate, dorsally carinate, free, a little longer and
narrower than the dorsal sepal. Petals ovate-lanceolate or elliptic-lanceolate,
acuminate, distinctly shorter than the dorsal sepal but nearly twice as wide.
Lip shorter than the sepals, suborbicular-ovate in outline, deeply pandurate-
trilobed, sessile at the subcordate base, abruptly apiculate at the rounded or
sometimes lightly retuse apex, about 9.8-17 mm. long, about 9.6-15.6 mm. across
near the base and 6-14.6 mm. wide in front; lateral lobes small, spreading, auriculi-
form, rounded, curved forward, abruptly contracted into a broad triangular-sub-
quadrate claw or isthmus; mid-lobe cordate-semiorbicular to reniform, broadly
rounded and commonly apiculate at the apex; disc at the base with a large, fleshy,
plurituberculate callus. Column short and stout, about 5-6.2 mm. high at the
back, with a pair of low, fleshy, porrect angles below and a pair of very incon-
spicuous, short, broad wings above.
Huanuco: 6 km. south of Tingo Maria, near highway, about
738 meters, epiphyte in moist forest valley, flowers very fragrant,
Seibert 2259. — San Martin: Zepelacio, near Moyobamba, 1100-1200
meters, epiphyte in forest, "fls. white, golden-yellow, violet and
brown-spotted," Klug 0.24- Also Mexico and Guatemala to Panama
(type of 0. ochmatochilum) and Colombia (type of 0. cardiochilum).
Oucidium onustum Lindl. Gen. & Sp. Orch. 203. 1833; Fol.
Orch. Oncidium 12, no. 35. 1855; L. Linden in Lindenia 11: 39, t. 498.
1895; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 208. 1922. On-
cidium holochrysum Reichb. f. Hamb. Gartenz. 18: 33. 1862; Wil-
liams, Orch.-Grow. Man. ed. 7: 612. 1894.
Plant medium-sized. Pseudobulbs aggregated, ovoid or oblong-ovoid, com-
pressed, 1- or sometimes 2-leaved, commonly green mottled with brown, 2-4 cm.
high, clothed below by 2 or 3 pairs of imbricating sheaths which are successively
larger upward, the uppermost leaf-bearing. Leaves oblong or linear-oblong, often
conduplicate in the dried specimen, acute, very slightly narrowed toward the base,
fleshy, very variable in size, 5-12.7 cm. long (according to collector's notes)
and 1-1.8 cm. (2.5 cm. according to Kranzlin) wide. Scape much surpassing the
leaves, about 18 to more than 40 cm. long, racemose or rarely paniculate above,
commonly nodding toward the summit; peduncle up to 22 cm. or more long,
provided with several short, close, tubular sheaths; inflorescence densely several-
to many-flowered, often secund, up to 20 cm. long. Floral bracts very minute.
Pedicellate ovary slender, commonly 1.5-2 cm. long. Flowers medium-sized,
with wide-spreading segments, entirely yellow. Dorsal sepal strongly cucullate,
ovate or ovate-elliptic, acute or acuminate, 7-7.4 mm. long, nearly 6 mm. wide
when forcibly expanded. Lateral sepals free, very similar to the dorsal sepal but
slightly longer and narrower, oblique, acute and apiculate. Petals nearly twice
as large as the sepals, suborbicular, broadly rounded or lightly retuse and apiculate
894 FIELDIANA: BOTANY, VOLUME 30
at the apex, with an undulate margin. Lip larger than the petals, suborbicular
in outline, deeply 3-Iobed near the base, sessile at the broadly cuneate base,
emarginate in front, about 19 mm. or less long from the base to the tip of an apical
lobule, about 20 mm. wide near the base and across the middle; lateral (basal)
lobes relatively small, spreading, obliquely obovate-oblong, broadly rounded and
often subcrenate at the apex; mid-lobe much larger, nearly sessile, reniform-
semiorbicular; disc at the base with a 3-lobulate callus of which the lateral lobules
are large, semioval and flat or cochleate and the mid-lobule small and ancipitous.
Column very short and stout, about 2 mm. high at the back, with a pair of large,
obliquely semilunate, obtuse wings.
Piura: Negritos, Cerro Negro, Cerro Prieto, 12-20 miles east of
Cape Parifias, about 730 meters or more, on crevices of cliffs, on trees
or cacti or at the base of tree trunks, H aught 123, 184, 0466. Also
Panama and western Colombia (type), and Ecuador.
Oncidium ovatilabium C. Schweinf. Am. Orch. Soc. Bull. 14,
no. 9: 384, t., 1946. Figure 174.
Plant epiphytic, medium-sized. Rhizome stout, concealed by sheaths. Pseudo-
bulbs apparently rather remote, very slender, cylindric, gradually narrowed up-
ward, complanate, 1- or 2-leaved, about 7 cm. or more high, surrounded on each
side by 4 or 5 distichous sheaths which are imbricating, conduplicate, successively
longer upward and mostly leaf-bearing. Leaves linear-oblanceolate to narrowly
elliptic-oblong, acute or short-acuminate, more or less narrowed below (those on
the pseudobulb very gradually narrowed to a conduplicate, petiole-like base), up
to 27 cm. long and 2 cm. wide. Scapes in the axil of a sheath, stout, suberect,
more or less surpassing the leaves; peduncle 19-33 cm. long, with 2 or 3 remote,
very short and close, tubular sheaths; inflorescence a virgate panicle 16-49 cm.
long, consisting of 6-14 short, densely flowered, wide-spreading branches, the
lower ones 4.5-9.2 cm. distant; main rachis above and that of branches strongly
fractiflex. Floral bracts conspicuous, round-ovate, deeply concave, obtuse. Flow-
ers numerous, very small for the genus, fragrant, with wide-spreading or reflexed
segments, brownish yellow with the central part [lip] dark red or reddish brown.
Dorsal sepal spatulate or oblong-spatulate, rounded at the apex, adnate to the
column below, the free part 7-7.4 mm. long, about 3 mm. wide. Lateral sepals
oblanceolate, more or less oblique, subacute, dorsally carinate above, distinctly
longer and commonly narrower than the dorsal sepal. Petals obliquely oval or
obovate-oval, broadly obtuse or rounded at the apex, obliquely adnate to the
column, about equally long with the dorsal sepal but markedly wider. Lip simple,
strongly convex, ovate, sessile at the rounded base, obtuse at the concave apex,
about 8 mm. long, 6.4-7.8 mm. wide when forcibly expanded; disc through the
lower half with a broadly oblong, lightly bisulcate (i.e., tricarinate) thickening.
Column relatively large, stout, acute at the tip, without distinct wings, about
3-3.9 mm. high at the back.
Hudnuco: Yanano, about 1800 meters, on mossy tree, Macbride
3844 (type). Huacachi, near Muna, about 2000 meters, in "mon-
tana," Macbride 4102.
ONCIDIUM
odaiiCafrium
FIG. 174. Oncidium ovatilabium C. Schweinf. 1, plant; X %. 2, flower from
front, natural position; X 3. 3, column and lip, oblique view; X 4. 4, lip some-
what expanded, from above; X 4.
895
896 FIELDIANA: BOTANY, VOLUME 30
Oncidium Papilio Lindl. Bot. Reg. 11: t. 910. 1825; Hook. Bot.
Mag. 55: t. 2795. 1828; Cogn. & Goos. Diet. Icon. Orch. Oncidium
t. 3. 1896; Cogn. Martius Fl. Bras. 3, pt. 6: 424, t. 91. 1906; Kranzl.
Engler, Pflanzenr. IV. 50 (Heft 80), 170. 1922.
Plant large, showy. Pseudobulbs aggregated, rather small, suborbicular,
strongly compressed, about 3-5 cm. high, unifoliate, clothed at the base by several
pairs of evanescent, ovate-triangular, scarious sheaths. Leaf oval to elliptic-
oblong, coriaceous, acute, rather long-narrowed to a complicate base, about
12-22 cm. long, 4-7 cm. wide, green mottled with brown. Scape many times
longer than the leaves, ascending or spreading, 60-120 cm. long; peduncle elongate,
not thickened at the joints but terete and clothed below with numerous remote,
short, close, evanescent, tubular sheaths, above dilated, strongly compressed
and often bialate; inflorescence simple or somewhat branched, each branch densely
few-flowered with 1 or 2 flowers opening at once. Floral bracts short, broadly
triangular, strongly concave. Pedicellate ovary abbreviated. Flower very large,
with wide-spreading, membranaceous segments. Dorsal sepal oblanceolate-linear,
acute, with slightly undulate margins, purplish-brown, sparsely mottled with
greenish yellow, about 8-13 cm. long and 5-7 mm. wide. Lateral sepals free,
oblong-lanceolate, falcate-decurved, obtuse to short-acuminate, strongly crisped-
undulate on the margins, much shorter and about three or more times as wide
as the dorsal sepal, brown and yellow in transverse bands. Petals very similar
to the dorsal sepal but apparently slightly shorter and narrower. Lip slightly
shorter and much broader than the lateral sepals, broadly obovate in outline,
pandurate-trilobed below the middle, truncate-subcordate at the sessile base,
commonly lightly emarginate in front, 3.5-6 cm. long from the base to either side
of the apex, 1.8-2.7 cm. wide across the lateral lobes at the base and 3-6 cm.
wide across the mid-lobe in front; lateral lobes relatively small, spreading, semi-
orbicular to semiovate, more or less abruptly narrowed in front to the short broad
claw, somewhat undulate on the margins; mid-lobe much larger, suborbicular to
transversely suborbicular-oval, with the margins strongly crisped-undulate, yellow
and reddish-brown more or less mottled with yellow and the center yellow; disc
at the base with a fleshy ovate or ovate-lanceolate, obscurely 3-lobed callus.
Column small, slender, 7-10 mm. high, with a pair of prominent, porrect wings
which have the posterior lobe linear-subulate and capitate at the tip and the
anterior lobe relatively large, obliquely triangular-ovate and laciniate in front.
Loreto: Tarapoto, 400 meters, epiphyte, Vie 6607. Also Pan-
ama(?), Trinidad (type), Venezuela, and northern Brazil.
Oncidium pardothyrsus Reichb. f. & Warsc. Bot. Zeit. 15: 158.
1857; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 273, fig. 23A,
ar-b. 1922.
Plant large, showy. Pseudobulbs (according to Kranzlin) large, flat, ridged,
resembling those of O. suavis Lindl. Leaves unrecorded. Scapes probably 90 cm.
or more long, not climbing, paniculate above; peduncle provided with distant,
scarious, short sheaths (according to Kranzlin); panicle narrow and elongate,
many-flowered, with numerous short, remote branches which are 6 cm. or less
SCHWEINFURTH: ORCHIDS OF PERU 897
distant, about 4 cm. long (up to 10 cm. long, according to Kranzlin), simple,
apparently 2- to 4-flowered. Floral bracts rather conspicuous, ovate, acute or
acuminate. Pedicellate ovary 2.5 cm. or less long. Flowers rather small, with
wide-spreading segments. Dorsal sepal with a short narrow claw, apparently
about 10 mm. long; lamina broadly oblong-ovate, obtuse, about 5 mm. wide,
strongly undulate, chestnut-brown with a yellow border near and at the apex.
Lateral sepals free, similar but somewhat shorter and longer-clawed according to
Kranzlin. Petals similar to the lateral sepals. Lip much shorter than the sepals,
obovate-suborbicular in outline, pandurate-trilobed near the middle, 7 mm. long,
5.5 mm. wide near the base and 6 mm. wide in front according to Kranzlin, sessile
with a subtruncate base, retuse in front, yellow with a broad chestnut band
across the basal half; lateral lobes short, semiovate- triangular, broadly obtuse
without, narrowed above to a short broad isthmus; mid-lobe cordate-reniform
("transversely oblong"); disc at the base with a large, velvety, multituberculate
callus which is 3-lobulate in front. Column very short and stout, with a pair of
short, fleshy angles below and a pair of short, subquadrate wings above.
Peru: Habitat not recorded, Warscewicz s.n. (type). Also Wallis
s.n., according to Kranzlin.
The Ames Herbarium contains a sketch of this species, with
several floral analyses.
Oncidium pentadactylon Lindl. Ann. & Mag. Nat. Hist. 15:
383. 1845; Fol. Orch. Oncidium 42, no. 136. 1855; Kranzl. Engler,
Pflanzenr. IV. 50 (Heft 80), 181. 1922.
Plant large, epiphytic, forming large clumps. Pseudobulbs ovoid to narrowly
ovoid, more or less compressed, bifoliate, up to 7 cm. high, clothed below by 1 or
2 pairs of slightly longer, scarious sheaths of which the upper ones are occasionally
leaf-bearing. Leaves lanceolate-elliptic, up to 17.6 cm. long and 3.4 cm. wide
(up to 30 cm. long and 2 cm. wide according to Kranzlin), acute or short-acuminate,
gradually narrowed to a complicate base. Scape elongate, up to 79 cm. long
(said to reach 3 m. in length), much surpassing the leaves, suberect to arcuate,
loosely paniculate above; peduncle about 21 cm. or more long, provided with 2 or
3 remote, short, close tubular sheaths; panicle very long with branches more or
less remote (up to 8.5 cm. distant), loosely few- to many-flowered, the lower ones
compound and the rachis of all fractiflex. Floral bracts lanceolate-triangular,
concave, scarious. Pedicellate ovary 1.7 cm. or less long, much surpassing the
subtending bract. Flowers of two kinds, all with wide-spreading or reflexed
segments, yellow blotched with brownish purple in the lower part. Sterile flowers
very small, with 5 oblong-oblanceolate, subequal segments. Fertile flowers much
larger, medium-sized for the genus. Dorsal sepal oblanceolate-oblong, abruptly
acute, about 10.2-11.5 mm. long (7 mm. long according to Kranzlin) and 3-3.8 mm.
wide. Lateral sepals obliquely linear-oblanceolate, abruptly acute at the apex,
commonly lightly falcate, markedly longer (rarely shorter) and somewhat narrower
than the dorsal sepal. Petals lanceolate-oblong or pandurate-oblong, slightly
shorter (rarely longer) and broader than the dorsal sepal, oblique. Lip commonly
shorter than the lateral sepals, pandurate-trilobed, sessile at the minutely auriculate
base, retuse or lightly emarginate and often apiculate in front, about 11.6-14 mm.
898 FIELDIANA: BOTANY, VOLUME 30
long (9-10 mm. long and 6 mm. wide according to Kranzlin) from the base to the
tip of an apical lobule, about 8.6-14 mm. wide across the basal portion (lateral
lobes) and 10.4-16 mm. wide across the anterior part (mid-lobe); lateral lobes
oblong-auriculate or round-auriculate, retrorse, unguiculate-narrowed to the mid-
lobe; mid-lobe much larger than the lateral lobes, transversely oval-oblong to
cordate-reniform; disc at the base with a multituberculate callus, usually having
3 tubercles in front. Column very small, sigmoid when viewed from the side,
about 3.2 mm. high at the back, below with a pair of porrect, fleshy angles and
above with a pair of conspicuous, dolabriform or transversely oblong, more or
less bilobed wings.
Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 176. 1921.
Ecuador (type), Colombia and Venezuela.
Oncidium pictum HBK. Nov. Gen. et Sp. 1: 346, t. 81. 1816;
Lindl. Gen. & Sp. Orch. 201. 1833; Fol. Orch. Oncidium 28, no. 89.
1855; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 193. 1922. On-
cidium falcicorne Reichb. f. in sched., fide Kranzl. I.e.
Plant large. Pseudobulb oblong-ovoid, subcompressed, bifoliate, about 7.5-8
cm. high, clothed below by several pairs of imbricating, conduplicate sheaths which
are progressively larger and leaf-bearing upward. Leaves linear or linear-oblong,
obtuse to acute, gradually narrowed below to a complicate base, about 25-30 cm.
long and 2.5 cm. wide. Scape erect, about twice as long as the leaves, paniculate
above, about 60 cm. long; peduncle with several remote, close, short sheaths;
panicle many-flowered, with several subremote branches which are spreading,
short, simple and few- (2- to 3-) flowered, with a fractiflex rachis. Bracts triangular
or ovate-lanceolate, acuminate, concave, much shorter than the pedicellate ovary
which is about 2 cm. long. Flowers medium-sized, with wide-spreading segments,
yellow with red or purplish spots. Dorsal sepal oblanceolate, acute, lightly un-
dulate on the margins, 1.3 cm. long and about 3 mm. wide (according to Kranz-
lin). Lateral sepals similar but apparently a little narrower, free, oblique. Petals
oblong or ligulate, obtuse, about equally long but somewhat wider than the sepals.
Lip slightly exceeding the sepals, pandurate-trilobed, sessile at the cordate base,
retuse at the apex, subquadrate in outline, about 1.5 cm. long, 6-7 mm. wide
across the lateral lobes at the base and 1.5 cm. wide in front according to Kranz-
lin; lateral lobes small, triangular-semiovate, apparently abruptly narrowed to
a rather long isthmus; mid-lobe much larger, cordate-reniform; disc at the base
with a large crest consisting of 7-8 minute tubercles. Column small, sigmoid
when viewed from the side, below with a pair of prominent, porrect, semiobovate
plates and above with a pair of erect-ascending, narrowly triangular or subulate
wings.
Peru: Habitat not recorded, Mathews 1920, fide Lindley. Also
Colombia (type) and Ecuador.
Oncidium polyadenium Lindl. Fol. Orch. Oncidium 27, no. 86.
1855. Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 241. 1922.
SCHWEINFURTH: ORCHIDS OF PERU 899
Plant large. Pseudobulbs, leaves and lower part of the scape unknown in
the type. Scape stout, adorned with several short, close, remote sheaths, the
portion present up to about 15 cm. high; inflorescence a large pyramidal panicle,
many-flowered, about 24 cm. high; branches spreading, simple, up to 10 cm.
long, few-flowered above, with a fractiflex rachis. Floral bracts small, triangular,
acuminate, much shorter than the slender pedicellate ovary which is up to 3 cm.
long. Flowers rather large, with wide-spreading segments, apparently whole-
colored, yellow. Dorsal sepal cuneate-obovate, retuse or broadly obtuse, about
12.5 mm. long and 7 mm. wide. Lateral sepals similar but a little longer, free,
oblique. Petals similar to the dorsal sepal but much longer and broader than
either sepal, oblong-obovate, retuse. Lip about as long as the lateral sepals or
longer, pandurate-trilobed, sharply indented at about the middle of each side,
subquadrate or oblong-obovate in outline, sessile at the cordate base, retuse at
the apex, about 14-17 mm. long from the base to the tip of an apical lobule,
about 9-12 mm. wide across the basal portion and 11-21 mm. wide in front;
lateral lobes indistinct, spreading, semiorbicular with undulate-crenate margins;
mid-lobe broader, reniform or transversely oval; disc at the base with a large,
multipapillose callus (papillae in 5 rows) extended in front into several crenate-
dentate keels, and with several separate warts on each side. Column small,
with a pair of spreading semiobovate dilations below and with a pair of semiovate-
dolabriform, retuse or crenate wings above.
Two collections from Ecuador (Espinosa 1607, 1798), which have been referred
to this species, furnish the following vegetative characters: Plant rather large,
forming rosettes, 50-80 cm. tall. Pseudobulbs small, entirely concealed by several
pairs of imbricating, conduplicate sheaths which are mostly leaf-bearing and are
progressively larger upward. Leaves oblong or elliptic-oblong, up to 55 cm. long
and 5 cm. wide, acute or subacuminate.
Peru: Habitat not recorded, Lobb s.n. (type). — Amazonas: Cha-
chapoyas, Cerro Puma Urco, Soukup 4094- Also Ecuador.
The Ames Herbarium contains a photograph of the type, with
an analysis of the lip and column.
Oncidium pusillum (L.) Reichb. f. Walp. Ann. 6: 714. 1863;
Cogn. Martius Fl. Bras. 3, pt. 6: 430, t. 62, fig. 1. 1906; Kranzl.
Engler, Pflanzenr. IV. 50 (Heft 80), 99. 1922. Epidendrum pusillum
L. Spec. PI. ed. 2. 1352. 1763. Cymbidium pusillum Sw. Nov. Act.
Ups. 6: 74. 1799. Oncidium iridifolium HBK. Nov. Gen. et Sp. 1:
344. 1816. Figure 175.
Plant small, caespitose, very variable, up to 8 cm. high. Stems abbreviated,
entirely concealed by a flabelliform or suborbicular cluster of leaves which are
imbricated at the base. Leaves equitant, more or less oblique and falcate, linear,
linear-oblong to oblong-elliptic or rarely elliptic (often slightly broader above the
middle), acute or rarely obtuse, up to 7 cm. long and (rarely) 1.2 cm. wide. Scapes
subequaling the leaves, axillary and often several to a plant, more or less flattened
and narrowly bialate above, terminating in an abbreviated 1- to 6-flowered raceme.
Floral bracts short, conduplicate-triangular, dorsally carinate, shorter than the
900
FIELDIANA: BOTANY, VOLUME 30
FIG. 175. Oncidium pusillum (L.) Reichb. f . Plant; X 1.
pedicellate ovary which is 6- winged and 8 mm. or less long. Flower variable, large
for the plant, membranaceous, with wide-spreading segments, yellow often with
orange, red-brown or purplish spots. Dorsal sepal relatively small, elliptic to
obovate or suborbicular, acute, apiculate to broadly rounded at the tip, dorsally
carinate, 5-6 mm. long, 2.2-4.5 mm. wide. Lateral sepals obliquely lanceolate,
acute or acuminate, apiculate, concave, dorsally carinate, a little shorter (longer
according to Cogniaux) and much narrower than the dorsal sepal. Petals much
longer and broader than the dorsal sepal, elliptic-oblong to obliquely round-
obovate, acute, apiculate or rounded at the apex, undulate. Lip much larger
than the other segments, pandurate-trilobed, sessile with an auriculate base,
retuse to bilobed at the apex, about 13.2-18.5 mm. long from the base to the
tip of an apical lobe, about 12-18 mm. wide across the lateral lobes near the
base and 10.8-22 mm. wide across the mid-lobe; lateral lobes relatively small,
spreading, suborbicular or obovate-suborbicular; mid-lobe much larger, from a
short broad claw semiorbicular, more or less indented in the middle of each side
(and thus 4-lobed) ; disc at the base with a prominent, 3-lobed callus of which the
lateral lobes are broad, lobulate and flat and the mid-lobe longer and convex.
Column very short and stout, about 1.5-2 mm. or more high at the back, semi-
SCHWEINFURTH: ORCHIDS OF PERU 901
orbicular-dilated on each side below, with a pair of prominent, semiorbicular-
flabellate, crenulate or deeply dentate wings above.
Peru: Habitat not recorded, MacLean s.n. — Huanuco, fide Schlech-
ter, Fedde Repert. Beih. 9: 176. 1921.— Junin: Along Rio Perene,
near "Hacienda 3," Colonia Perene, about 600 meters, in forest,
Killip & Smith 25171. Colonia Perene, 590 meters, Hitchcock 22105.
Chanchamayo Valley, 1000 meters, Schunke 556; 700 meters, Schunke
1665. Hacienda Schunke, La Merced, about 1230 meters, Macbride
5600. Puerto Bermudez, about 375 meters, epiphyte in dense forest,
Killip & Smith 26519. — Loreto: Bajo Amazonas, Gamitanacocha, Rio
Mazan, 100-125 meters, Schunke 135; Vargas 4673. Caballo-Cocha on
the Amazon River, Williams 2028. Lower Rio Huallaga, Fortaleza, Yu-
rimaguas, 155-210 meters, Williams 4326. Mishuyacu, near Iquitos,
100 meters, epiphyte in dense forest, Killip & Smith 29896; Klug 893;
Pebas, Rand s.n. Vicinity of Iquitos, 100 meters, Klug 10102. Vicin-
ity of Iquitos, Rancho Indiana, 110 meters, Mexia 6436. — San Mar-
tin: Prov. of Lamas, Trocha Agua Blanca-Desquite, on road to
Tarapoto, 600-800 meters, in tropical forest, Ferreyra 7973. Prov.
of Mariscal Caceres, "Frente a Puerto Huicte, entre Tingo Maria y
Juanjui," in tropical forest, 500-550 meters, Ferreyra 4409. Also
Mexico, through Central America and the West Indies to Surinam
(type of Epidendrum pusillum), Colombia, Brazil, Bolivia and Peru.
Oncidium pyramidale Lindl. Ann. & Mag. Nat. Hist. 15: 384.
1845; Fol. Orch. Oncidium 29, no. 98. 1855; Veitch, Man. Orch. PI. 8:
73. 1892; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 195, fig. 16J,
a-c. 1922. Oncidium chrysopyramis Reichb. f. & Warsc. Bonpl. 2:
108. 1854; Lindl. Fol. Orch. Oncidium 29, no. 97. 1855; Veitch, Man.
Orch. PL 8: 25. 1892; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80),
196, fig. 16F, a-d. 1922.
Plant medium-sized. Pseudobulbs aggregated (according to Kranzlin), ovoid
to narrowly ovoid-cylindric, compressed, 1- to 3-leaved, 2.5-7.3 cm. high, clothed
at the base with 2 or 3 pairs of scarious, imbricating, conduplicate sheaths (often
evanescent), which are successively larger upward, the uppermost leaf-bearing.
Leaves elliptic-oblong to oblanceolate, acute or short-acuminate, more or less
long-narrowed to a complicate base, 11-21.5 cm. long, 1.5-3 cm. wide, those
on the pseudobulb largest. Scape from the axil of a basal sheath, much longer
than the leaves, 32-45 (sometimes 60) cm. long, suberect, often somewhat arcuate
above, terminating in a rather narrow, loose, many-flowered panicle; peduncle
16-24.5 cm. long, provided with a few remote, short, close sheaths; panicle about
16-32 cm. long, with numerous branches which are short or very short and either
simple and few-flowered or branched and many-flowered, with a fractiflex rachis.
Floral bracts minute, triangular, much shorter than the slender pedicellate ovary
902 FIELDI ANA: BOTANY, VOLUME 30
which is 10 mm. or less long. Flowers small, yellow, fragrant, with wide-spreading
or reflexed segments. Dorsal sepal oblanceolate-oblong, "oval," broadly obtuse,
about 7.5 mm. long (5-6.5 mm. long according to Kranzlin) and 2.7 mm. wide.
Lateral sepals free, linear-oblanceolate, oblique, acute or subacute, narrower
and slightly longer than the dorsal sepal. Petals oblong-ovate, commonly abruptly
acute or apiculate, about as long as the dorsal sepal but nearly twice as broad.
Lip much larger than the other segments, pandurate-trilobed, sessile at the
abruptly auriculate base, bilobed in front, about 10.5-12.5 mm. long (8 mm.
long according to Kranzlin) from the base to the apex of an anterior lobule, about
9.8-11.3 mm. wide across the lateral lobes near the base and about 7.5-8.5 mm.
wide across the mid-lobe; lateral lobes suborbicular-auriculate, reflexed, gradually
narrowed to the claw in front; mid-lobe reniform or obovate-reniform, rather
abruptly dilated from the claw, bilobed in front; disc at the base with a large,
plurituberculate callus having 3 prominent tubercles in front. Column small,
strongly sigmoid when viewed from the side, about 3.4 mm. high at the back,
below with a pair of porrect, semiorbicular plates, above with a pair of narrowly
cuneate, more or less divergently bilobed wings and with a prominent, recurved,
linear-triangular, rostellar process.
Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 176. 1921.
Also Colombia (type of 0. pyramidale and 0. chrysopyramis) and
Ecuador.
The Ames Herbarium contains a photograph of the type of 0.
pyramidale and drawings of the type of 0. chrysopyramis.
Oncidium retusum Lindl. Bot. Reg. 23: sub t. 1920, no. 6.
1837; Lindl. & Paxt. Paxt. Flow. Card. 2: 9, no. 265, fig. 136. 1851-
2; Lindl. Fol. Orch. Oncidium 42, no. 138. 1855; Kranzl. Engler,
Pflanzenr. IV. 50 (Heft 80), 185, fig. 16D. 1922.
Plant variable, medium-sized to large. Pseudobulbs often crowded, ellipsoid
to fusiform-cylindric or pyriform-cylindric, commonly 2-leaved (rarely 1-leaved),
strongly compressed, 3-8 cm. high, concealed below by several pairs of imbricating,
conduplicate, scarious sheaths (later evanescent) which are successively larger
upward, and frequently leaf-bearing above. Leaves oblanceolate to oblong-
oblanceolate (rarely elliptic), 8-50 cm. long, 2.1-4.5 cm. wide, (much larger on the
pseudobulb), acute to short-acuminate, commonly gradually narrowed below to a
conduplicate, often petiole-like base. Scape much exceeding the leaves, about
34-94.5 (up to 120, fide collector's notes) cm. long, suberect, paniculate and many-
flowered above; peduncle about 18-32 cm. long, with 2-4 remote, short, close
sheaths; panicle more or less elongate, about 20-68 cm. long, with remote, short
branches which are simple or compound, few- to many-flowered, with the rachis
strongly fractiflex. Floral bracts very small, triangular or triangular-lanceolate,
concave, much shorter than the pedicellate ovary which is 1.7 cm. or less long.
Flowers of two kinds; sterile flowers more or less numerous, relatively small,
with 5 subequal, spatula te-cuneate segments, yellow with purplish-red blotches;
fertile flowers much larger, with wide-spreading or reflexed segments, yellow or
greenish yellow mottled or blotched with purplish red. Dorsal sepal cuneate-
spatulate, broadly rounded or lightly retuse at the apex, about 11.3-12.6 mm.
SCHWEINFURTH: ORCHIDS OF PERU 903
long, 3.9-4.4 mm. wide. Lateral sepals similar, slightly shorter and narrower,
broadly rounded or slightly retuse and dorsally mucronate at the apex, free,
suboblique. Petals obovate-oblong, lightly clawed, retuse and apiculate or dorsally
mucronate at the apex, oblique, slightly shorter and distinctly wider than the
dorsal sepal, with the margins lightly undulate. Lip much larger than the other
segments, pandurate-trilobed, sessile at the subtruncate base, bilobed in front
with lightly overlapping lobules, about 12-13.2 mm. long from the base to the
tip of an apical lobule, 11-11.6 mm. wide across the lateral lobes at the base,
12-12 2 mm. wide across the terminal lobe; lateral lobes semitriangular, broadly
rounded and spreading without, much narrowed in front forming a claw-like
isthmus; mid-lobe cordate-reniform, deeply bilobed; disc at the base with a
large, plurituberculate callus terminating in 3 prominent tubercles. Column
very 'short, dilated upward, more or less sigmoid when viewed from the side,
about 4 mm. high at the back, below with a pair of porrect, bluntly angular
plates and above with a pair of broadly dolabriform, lacerate-dentate, more or
less bilobed wings.
Amazonas: Near Mt. Pariahuanca, Mathews 1066 (type).— Caja-
marca: Prov. of Chota, near Huambos, 2200 meters, in low woods,
Ferreyra 8408—Cuzco: Prov. of Urubamba, between Sta. Rita and
Tuncapata, 2200 meters, on open grassy slope, Vargas 2639.— Hua-
nuco: Cani, "7 miles N.E. of Mito," about 2460 meters, on tree
trunk, Macbride 3434 — Junin: Huacapistana, about 1800 meters,
epiphyte in densely forested valleys, Kittip & Smith 24287. Also
Bolivia.
Oncidium Sanderae Rolfe, Card. Chron. ser. 3, 48: 67. 1910;
Bot. Mag. 137: t. 8374. 1911; Kranzl. Engler, Pflanzenr. IV. 50
(Heft 80), 171. 1922.
Plant medium-sized to large. Pseudobulbs often aggregated, orbicular-ovoid
to oblong-ovoid, more or less compressed, 2.5-6 cm. high, unifoliate, clothed
when young by several pairs of evanescent, imbricating sheaths. Leaf elliptic
to oblong-elliptic or oblong, acute to rounded at the apex, complicate at the sessile
base, brown-reticulated, 10-45 cm. long, 3.1-8 cm. wide. Scape suberect to
arcuate, much exceeding the leaves, about 25-80 cm. long, densely few- to several-
flowered at the apex, terete below, compressed and sometimes dilated above,
with 5 or more, not swollen joints producing short, close, evanescent sheaths.
Floral bracts oblong-lanceolate, acuminate, concave, much shorter than the
pedicellate ovary which is about 2.5-5 cm. long. Flowers large and showy, with
wide-spreading segments, successive. Dorsal sepal linear-oblanceolate, acute or
short-acuminate, about 5.5-10 cm. long, 5.5-8 mm. wide above, reddish brown.
Lateral sepals decurved, free, lanceolate-oblong, acute, oblique, craped-undulate
on the margins, yellow with red spots, 4-7 cm. long, about 1-2 cm. wide. Petals
very similar to the dorsal sepal but somewhat shorter and narrower, oblique,
acute or short-acuminate, reddish brown. Lip much shorter and broader than the
other segments, pandurate-trilobed, sessile at the subtruncate base, broadly
rounded in front, about 3-4 cm. long, about 1.5 cm. or more broad across the
lateral lobes, and 2-3 cm. broad across the mid-lobe; lateral lobes semiorbicular,
904 FIELDIANA: BOTANY, VOLUME 30
spreading, crisped-undulate, yellow with red-brown spots; mid-lobe suborbicular
with a pronounced claw, crisped-undulate, yellow with reddish brown spots
near the margin; disc at the base with a small, 5-lobed callus. Column very short,
clavate, 8-10 mm. high, above with a pair of broad wings which are long-pectinate
with glandular tips.
Junin: Chanchamayo Valley, 1200 meters, Schunke s.n., 1304-—
Loreto: Moyobamba district, Forget s.n. (type). Pongo de Man-
seriche, 160 meters, in rain-forest, Tessmann 4159. Puerto Mele"ndez,
below Pongo de Manseriche, 155 meters, in rain-forest, Tessmann
4752.
Oncidium Schillerianum Reichb. f. Bonpl. 2: 91. 1854; Cogn.
Martius Fl. Bras. 3, pt. 6: 400. 1906; Kranzl. Engler, Pflanzenr. IV.
50 (Heft 80), 280, fig. 23J, a-b. 1922.
Plant large, climbing. Pseudobulbs rather distant on an obliquely creeping
rhizome, subtetragonous (according to Kranzlin), oblong-ovoid, bifoliate, about
5 cm. high, clothed at the base by 2 pairs of imbricating sheaths of which the
upper one is leaf-bearing. Leaves oblong, subacute and obliquely bilobed at the
apex, narrowed at the complicate base, up to 15 cm. long, 3.3 cm. wide. Scape
very long, climbing, much exceeding the leaves, 60-120 cm. long, terminating in
an elongate narrow, very loose panicle which becomes flexuous above; branches
lax, short, mostly 2- to 4-flowered, often twice or three times branching. Floral
bracts minute, triangular, acute, much shorter than the slender, pedicellate ovary
which is about 2 cm. long. Flowers numerous, small, with wide-spreading or
reflexed segments, greenish yellow transversely mottled with brown. Dorsal sepal
oblong or oblanceolate-oblong, subacute or apiculate, narrowed below, about
15 mm. long and 4-5 mm. wide. Lateral sepals similar but a little longer, narrower
and more acute, free. Petals similar to the dorsal sepal but a little wider, oblong
or oblanceolate-oblong, acute or obtuse, undulate on the margin. Lip longer and
much broader than the other segments, pandurate-trilobed, sessile at the broad
base, emarginate to bilobed in front, about 18 mm. long and more than 20 mm.
wide near the front (according to Kranzlin); lateral lobes semitriangular or semi-
ovate, rounded at the subretrorse outer margin, retuse on the posterior margin,
narrowed in front to form a short, broad claw; mid-lobe much larger, cordate-
reniform, retuse; disc at the base with a large, multituberculate callus with 3
tubercles in front. Column small, lightly sigmoid when viewed from the side,
dilated on each side near the base, above with a pair of broadly dolabriform,
lobulate or dentate wings which are acute at each end.
Cajamarca: Prov. of Chota, between Huambos and Montan,
2300-2600 meters, epiphytic on (and between) bushes, Weberbauer
4210, fide Kranzlin, I.e. Habitat of type uncertain; introduced by
Consul Schiller.
No material of this species was available, but the Ames Her-
barium contains sketches of the plant with floral analyses from the
Reichenbach Herbarium.
SCHWEINFURTH: ORCHIDS OF PERU 905
Oncidium serpens Lindl. Gen. & Sp. Orch. 204. 1833; Lindl. &
Paxt. Paxt. Flow. Card. 2: 8, no. 254, fig. 125. 1851-52; Lindl. Fol.
Orch. Oncidium 36, no. 121. 1855; Kranzl. Engler, Pflanzenr. IV. 50
(Heft 80), 168. 1922.
Plant small, slender, with an elongate, filiform, creeping and flexuous stem
or rhizome. Pseudobulbs about 8-12 cm. distant, unifoliate, oblong-ovoid, appar-
ently 2 cm. or less high (2.5-3 cm. high, according to Kranzlin), clothed below
by several relatively large, unequal, sometimes leaf-bearing sheaths. Leaves
elliptic or oblong-elliptic, acute, narrowed below to a complicate, often petiole-
like base, up to 5 cm. long, 1.3-1.7 cm. wide (those on the pseudobulbs much the
largest). Scapes a little exceeding the leaves, diffuse, loosely 2- to 3-flowered above.
Floral bracts very minute, concave, much shorter than the slender pedicellate
ovary which is about 1.5 cm. long. Flowers small, with wide-spreading segments,
yellow, mottled with chestnut-brown. Dorsal sepal oblong-obovate, obtuse, 10-15
mm. long, about 5 mm. wide above. Lateral sepals obliquely oblanceolate-oblong,
a little longer and distinctly narrower than the dorsal sepal. Petals very similar
to the dorsal sepal, about equally long and wide or wider, oblique. Lip simple,
pandurate-oblong, truncate-obtuse or lightly retuse in front, about 12 mm.
long and 7 mm. wide near the apex; disc at the base with 2 parallel keels which
are obtuse-angled when viewed from the side. Column small, marginate, clavate,
about 4 mm. high, apparently wingless.
Peru(?): Patacocha, on the road to Esmeraldas, about 1800 me-
ters, on tree trunks, Hall s.n.
The Ames Herbarium contains a photograph and pen and ink
drawing of this species from the Lindley Herbarium.
Oncidium serratum Lindl. Sert. Orch. sub t. 48, no. 17. 1842;
Lindl. & Paxt. Paxt. Flow. Card. 1: 28, no. 42, fig. 15. 1850-51;
Batem. Bot. Mag. 93: t. 5632. 1867; Veitch, Man. Orch. PI. On-
cidium 8: 77. 1892. Oncidium Sanderianum Rolfe, Kew Bull. (1893)
337. Cyrtochilum serratum Kranzl. Notizbl. Bot. Gart. Berl. 7: 93,
in key. 1917, nomen; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 45.
1922.
Plant very large. Pseudobulbs often aggregated and set obliquely on a stout
rhizome, oblong-ovoid or pyriform-cylindric (more rarely oval), more or less
compressed, bifoliate (rarely unifoliate), 8-13 cm. high, clothed at the base by
2 or 3 pairs of imbricating sheaths which are successively larger upward, the upper
ones leaf-bearing. Leaves linear-oblanceolate, acute or short-acuminate, long-
narrowed to a channelled, petioled base, about 40-57 cm. long and 4 cm. wide.
Scape elongate, twining, up to 4 m. long, very loosely paniculate above, the
branches 13-25 cm. apart, loosely few- (3- to 6-) flowered, with a fractiflex rachis.
Floral bracts ovate-oblong, strongly concave, more or less obtuse, much shorter
than the slender pedicellate ovary which is 4-6 cm. long. Flowers large, with
wide-spreading or reflexed segments. Sepals brown with yellow tips and margins.
Dorsal sepal abruptly clawed, about 1.6-2.7 cm. long; lamina cordate-reniform
or semiorbicular, rounded at the apex, denticulate on the strongly crisped-undulate
906 FIELDIANA: BOTANY, VOLUME 30
margin, about 1.3-2.2 cm. long, 2.4-3.3 cm. wide; claw rather short, auricled on
each side at the base. Lateral sepals much longer than the dorsal sepal and longer-
clawed; lamina obliquely ovate or rhombic-ovate, obtuse, denticulate on the
strongly crisped-undulate margins, much longer and narrower than the lamina of
the dorsal sepal; claw rather gradually dilated into the lamina, auricled at the
base on the outer side. Petals much smaller than the sepals, connivent in natural
position, from a short, broad claw oblong-ovate, broadly obtuse or rounded at the
apex, shortly fimbriate on the strongly undulate margins, brown below with the up-
per portion yellow. Lip relatively very small, 3-lobed, broadly lanceolate in outline,
about 1.5 cm. long and 1 cm. wide across the lateral lobes, purplish-brown with a
white margin; lateral lobes small, obliquely triangular, acute, wide-spreading, re-
flexed, in front narrowed with bulging sides and then constricted to form the mid-
lobe; mid-lobe much larger, oblanceolate, acute or obtuse; disc at the base with a
large, rectangular crest which is outwardly denticulate, consisting of 3-5 more or
less distinct parallel keels and with a fleshy callus below on the median line. Col-
umn very small, 5-8 mm. high, with a pair of fleshy, rounded projections below
and with a pair of narrowly cuneate or falcate-linear, apically denticulate wings
above.
Peru: Habitat not recorded, Ruiz & Pavdn s.n. (type of 0.
serratum). Habitat unknown, imported by F. Sander & Co. (type
of 0. Sanderianuni)— Cajamarca, Huanuco, fide Schlechter, Fedde
Repert. Beih. 9: 177. 1921. Also Ecuador.
The Ames Herbarium contains a drawing of the type with floral
analyses from the Lindley Herbarium.
Oncidium sessile Lindl. & Paxt. Paxt. Flow. Card. 1: 101, t. 21.
1850-51; Warner & Williams, Orch. Alb. 5: t. 228. 1886; Veitch,
Man. Orch. PI. Oncidium 8: 78. 1892; Kranzl. Engler, Pflanzenr.
IV. 50 (Heft 80), 256. 1922.
Plant large and showy. Pseudobulbs more or less aggregated, oblong-ovoid,
compressed, bifoliate, 5-10 cm. high, clothed at the base with 2 or 3 pairs of im-
bricating sheaths which are successively larger upward, the upper ones leaf-bearing.
Leaves linear-oblong or lorate, subacute to acute (acuminate, according to Kranz-
lin), slightly narrowed to a complicate base, about 20-37.5 cm. long, up to 2.5 cm.
wide, those on the pseudobulb much the larger. Scape erect, stout, surpassing the
leaves, about 30-60 cm. tall, paniculate through the upper half; panicle loose, with
short, spreading, loosely few- (3- to 5-) flowered branches. Floral bracts short,
triangular-ovate, concave, acute, much shorter than pedicellate ovary which is
3-4 cm. long. Flowers rather small, with wide-spreading segments, yellow with
numerous reddish brown spots through the lower half. Sepals oblong or obovate-
oblong, obtuse, with lightly undulate margins, about 13-17 mm. long and 4-5 mm.
wide, the dorsal sepal slightly broader than the free lateral sepals. Petals very
similar to the sepals, but slightly shorter and broader according to Kranzlin. Lip
much larger than the sepals and petals, pandurate-trilobed, sessile at the base,
lightly retuse in front, nearly 20 mm. long, 14 mm. wide across the lateral lobes
at the base and 25 mm. wide in front according to Kranzlin; lateral lobes relatively
small, auriculiform-rounded, rather abruptly narrowed in front to form a short
SCHWEINFURTH: ORCHIDS OF PERU 907
broad claw; mid-lobe much larger, reniform-semiorbicular, retuse to shallowly
bilobed; disc at the base with an excavated, 3-lobed callus which is bilamellate in
front. Column short and stout, from the middle dilated toward the base, above
with a pair of short, broad wings of which the apical margin is truncate and den-
ticulate.
Peru(?), fide Schlechter, Fedde Repert. Beih. 9: 177. 1921.
Colombia (type) and Venezuela.
No material of this species was available.
Oncidium superbiens Reichb. f. Linnaea 22: 843. 1849; Hook,
f. Bot. Mag. 98: t. 5980. 1872; Warn. & Will. Orch. Alb. 6: t. 276.
1887. Oncidium aemulum Reichb. f. & Warsc. Bonpl. 2: 102. 1854;
Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 47. 1922. Oncidium in-
ferlobum Hort. ex Card. Chron. (1872) 904, in synon. Oncidium
undulatum Warn. & Will. Orch. Alb. 8: t. 368. 1889, notO. undulatum
(HBK.) Lindl. Cyrtochilum aemulum Kranzl. Notizbl. Bot. Gart.
Berl. 7: 93, in key. 1917, nomen; Kranzl. Engler, Pflanzenr. IV. 50
(Heft 80), 47, fig. 3C, a-e. 1922. Cyrtochilum superbiens Kranzl.
Notizbl. Bot. Gart. Berl. 7: 93, in key. 1917, nomen; Kranzl. Engler,
Pflanzenr. IV. 50 (Heft 80), 49, fig. 3E, a-/. 1922.
Plant very large, variable, with an elongate rhizome. Pseudobulbs commonly
aggregated, elongate-ovoid, subcompressed, about 6-10 cm. high, commonly 2-
leaved, clothed below by several imbricating sheaths which are successively larger
upward and mostly leaf-bearing. Leaves linear-oblanceolate, acute or short-
acuminate, up to 68 cm. long and 6 cm. wide, long-narrowed below to a compli-
cate, petiole-like base. Scape very elongate, twining, about 3-6 m. long, loosely
paniculate above with the branches remote in varying degrees (13 cm. or much
more apart) ; branches commonly short (up to 20 cm. long) and very loosely few-
flowered above, with a flexuous rachis. Floral bracts broadly ovate, obtuse or
subacute, cymbiform, much shorter than the pedicellate ovary which is about
3-4.5 cm. long. Flower large, with wide-spreading segments. Sepals brown with
the apex and the margin partly yellow. Dorsal sepal clawed, about 1.6-3 cm. long;
lamina ovate-suborbicular, broadly rounded and sometimes lightly retuse at the
apex, subcordate at the base, with the margins crisped-undulate, about 1.6-2 cm.
long and 1.8-2.8 cm. wide; claw short, with a large auricle on each side at the base.
Lateral sepals much longer, long-clawed, minutely connate at the base; lamina
obliquely suborbicular-ovate, broadly rounded or subtruncate above, very similar
to the lamina of the dorsal sepal but somewhat longer and either narrower or
broader; claw auricled at the base without, more or less cuneate-dilated into the
lamina above. Petals commonly smaller than the dorsal sepal, oblong-ovate to
suborbicular-ovate from a very short broad claw, rounded at the apex, with the
margins strongly crisped-undulate, yellow (rarely white) with transverse brown
bars on the lower half. Lip much shorter than the dorsal sepal and a little shorter
than the petals, making a right angle with the column at the base, then abruptly
recurved, auricled or rarely 3-lobed at the base, about 2 cm. long when forcibly
expanded, purple; auricles or lateral lobes short, obliquely ovate-triangular, broadly
908 FIELDIANA: BOTANY, VOLUME 30
obtuse, not retrorse, reflexed, commonly about 4.5-7 mm. long; mid-lobe much
longer, oblong-lanceolate or oblanceolate, commonly more or less acute and con-
cave at the apex, 1-1.5 cm. long, usually 5-6 mm. wide where broadest; disc at the
base with a long, fleshy ridge which is dilated in front into a plurituberculate crest
and has a transverse bifurcate ridge on the lateral lobes. Column very short and
stout, 7-10 mm. high at the back, with a pair of porrect, semielliptic dilations be-
low, and a pair of falcate-ligulate to cuneate-dolabriform wings above.
Peru: Paramo de la Vuelta, 4000 meters, Wallis s.n., fide Kranzl.
— Cajamarca: Prov. of Chota, near Huambos, Weberbauer 4196.—
Huanuco: Yanano, about 1800 meters, on mossy tree, Macbride 3850.
— Junin: Prov. of Tarma, mountains west of Huacapistana, Weber-
bauer 2103. Also Colombia (type of 0. superbiens), Venezuela and
Ecuador.
Oncidium tigratum Reichb. f. & Warsc. Bonpl. 2: 109. 1854;
Lindl. Fol. Orch. Oncidium 40, no. 129. 1855.
Plant apparently medium-sized. Pseudobulbs and leaves not described. Scape
slender, loosely paniculate above; peduncle more than 12 cm. long, with several
remote, close, short, evanescent sheaths; branches of the panicle remote, com-
pound, loosely several-flowered, with a strongly fractiflex rachis, apparently about
9.5 cm. or less long. Floral bracts minute, linear-triangular, much shorter than
the slender pedicellate ovary which is about 1.5 cm. long. Flowers small, with wide-
spreading or reflexed segments, deep yellow, with crimson blotches except on the
anterior part of the lip. Dorsal sepal cuneate-oblong, apiculate, apparently about
8 mm. long. Lateral sepals obliquely cuneate-oblong, apparently free, a little
longer and narrower than the dorsal sepal. Petals flabellate from a cuneate base,
rounded, lightly retuse and often apiculate at the apex, twice as wide as the sepals.
Lip much wider than the sepals and petals, pandurate-trilobed, sessile at the sub-
cordate base, lightly emarginate in front; basal half transversely rhombic, acute
on each side, abruptly narrowed to a very short narrow claw; mid-lobe cordate-
reniform, retuse, a little broader than the basal portion; disc at the base with a
verrucose callus having 3 short keels in front. Column very short, recurved, cla-
vate, below with a pair of bilobulate, porrect dilations and above with a pair of
broad, deeply bilobed column-wings which appear to be sublunate when the lobes
are taken together.
Peru: Habitat not recorded, Warscewicz s.n. (type). — Huanuco,
fide Schlechter, Fedde Repert. Beih. 9: 177. 1921.
The Ames Herbarium contains sketches of this species with floral
analysis from the Reichenbach Herbarium.
Oncidium tipuloides Reichb. f. Bot. Zeit. 10: 856. 1852; Lindl.
Fol. Orch. Oncidium 43, no. 142. 1855; Cogn. Martius Fl. Bras. 3,
pt. 6: 391. 1906; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 269,
fig. 22E, a-c. 1922.
SCHWEINFURTH: ORCHIDS OF PERU 909
Plant large. Pseudobulbs not described. Leaves (but one recorded) long-
petioled; lamina oblong or elliptic-oblong, acute or short-acuminate, cuneate be-
low, about 20-27 cm. long, 3-4 cm. wide; petiole elongate, stout, conduplicate,
10-12 cm. long. Scape much exceeding the leaves, erect or ascending, rather stout,
loosely paniculate, many-branched and many-flowered above; branches rather
short, widely spreading and often decurved, simple or sometimes compound,
loosely few-flowered, 3-4 cm. distant, with a more or less fractiflex rachis. Floral
bracts minute, narrowly triangular-ovate, subulate at the apex, much shorter than
the slender, pedicellate ovary which is 1-1.5 cm. long. Flowers medium-sized, with
spreading or reflexed segments, yellow, sparingly spotted with red-brown. Dorsal
sepal cuneate-spatulate, obtuse to'subacute, more or less undulate or repand on
the margins, 13-14 mm. long, about 3 mm. wide. Lateral sepals similar but a little
narrower, oblanceolate, free, suboblique, subacute. Petals oblong-obovate or spat-
ulate-obovate, subobtuse to acute, somewhat undulate on the margin, a little
shorter and broader than the dorsal sepal, suboblique. Lip longer and much
wider than the other segments, broadly obovate in outline, pandurate-tnlobed,
subtruncate at the base, emarginate at the apex, 17-18 mm. long, about 9 mm.
wide across the lateral lobes at the base and 16-18 mm. wide across the mid-lobe
in front; lateral lobes small, semitriangular-auriculiform, rounded without, con-
tracted in front into a long, narrow claw; mid-lobe much larger, abrupt, cordate-
reniform, strongly emarginate; disc at the base with a small, transverse, semilunate
ridge having an obtuse lobule on each side and a narrow central keel in front.
Column small, about 4 mm. high, sigmoid when viewed from the side, with a pair
of prominent, obliquely subquadrate, retuse plates below, wingless above.
Huanuco: Near Cuchero (Cochero), on trees, Poeppig 1635.
The Ames Herbarium contains a sketch of this species from the
Reichenbach Herbarium.
Oncidium trilingue Lindl. & Paxt. Paxt. Flow. Gard. 1: 42.
1850-51; I.e. 2: 125, t. 63. 1851-52; Lindl. Fol. Orch. Oncidium 4,
no. 5. 1855. Cyrtochilum trilingue Kranzl. Notizbl. Bot. Gart. Berl.
7: 92, in key. 1917, nomen; Kranzl. Engler, Pflanzenr. IV. 50 (Heft
80), 38, fig. 24, a-c. 1922.
Plant apparently large. Pseudobulbs and leaves not recorded. Scape very
long, climbing, the upper portion (which alone exists in the type) about 80 cm.
long (according to Kranzlin), paniculate in the lower part with short, few- (3- to 6-)
flowered branches having a fractiflex rachis. Floral bracts broadly ovate, deeply
concave, obtuse to acute, shorter than the pedicellate ovary which is 2.5-3 cm.
long. Flowers large, loose, with wide-spreading or reflexed segments, about 1.5-
5.3 cm. apart, dark or light brown with yellow margins on the sepals and petals
and a yellow crest on the lip. Dorsal sepal clawed, about 17 mm. long; lamina
broadly ovate or triangular-ovate, obtuse and apiculate, with the margins strongly
crisped-undulate, the upper portion recurved in natural position, about 12 mm.
long and 10.5 mm. wide when expanded; claw short, channelled, with a rounded,
fleshy auricle on each side at the base, rather abruptly dilated into the lamina.
Lateral sepals much longer and slightly narrower above, obliquely oblanceolate,
910 FIELDIANA: BOTANY, VOLUME 30
obtuse (acuminate, according to Kranzlin), with the claws (which are auriculate
at the base) connate below and gradually dilated into the lamina, dorsally carinate
above, somewhat undulate on the margins. Petals obliquely elliptic-lanceolate
with a short, indistinct claw, acute or acuminate, strongly crisped-undulate on the
margins, with the apex recurved, about as long as the dorsal sepal when expanded,
but somewhat narrower. Lip spreading from the column, with the upper part cir-
cinate-recurved, about 18 mm. long when expanded, auriculate-dilated on each
side at the base, with the subtriangular, lobulate dilations reflexed; terminal por-
tion longer, elliptic-lanceolate, undulate-crisped on the margins, obtuse and apicu-
late (acuminate, according to Kranzlin); disc at the base with a stout callus termi-
nating in 3 fleshy, ovoid teeth (the middle tooth largest) and in front with 3 flat
teeth of which the lateral ones are smaller and oblong-lanceolate and the middle
one ovate, convex and often lobulate above. Column very short and stout, dilated
above, about 6 mm. high, with a pair of porrect, angular dilations below and a pair
of small, obliquely oblong, falcate, spreading wings in the middle.
Peru: Near the source of the Maranon River, Warscewicz s.n.
(type).— Huanuco, fide Schlechter, Fedde Repert. Beih. 9: 177. 1921.
Also Bolivia, fide Kranzlin.
A flower of apparently typical material has been examined.
Oncidium tripterygium Reichb. f. Bot. Zeit. 10: 694. 1852;
Xen. Orch. 1: 192, t. 69, fig. Ill, 8-9. 1856; Kranzl. Engler, Pflan-
zenr. IV. 50 (Heft 80), 128. 1922.
Pseudobulbs, leaves and inflorescence not described. Flower (a single one
present) medium-sized, with wide-spreading segments, brown with a yellow lip.
Dorsal sepal ovate-lanceolate or oblong-lanceolate ("ovate"), acute or acuminate,
about 16 mm. long and 5-6 mm. wide (according to Kranzlin). Lateral sepals
connate to above the middle, forming a lamina which is obovate-oblong in out-
line, a little longer and broader than the dorsal sepal and deeply bilobed above
with acute, diverging lobules which have undulate inner margins. Petals elliptic-
lanceolate ("oblong-lanceolate"), about as long as the dorsal sepal but a little
broader. Lip pandurate-trilobed, obovate-oblong in outline, about 20 mm. long,
10 mm. wide near the base and 14 mm. wide in front (according to Kranzlin),
cordate at the base, emarginate at the apex, with lightly crenulate margins; lateral
lobes small, semiorbicular; mid-lobe much larger, sessile or nearly so, obovate-
reniform; disc at the base with a short denticulate keel divided in front and sur-
rounded on each side by a solitary callus. Column very short and stout, without
wings above.
Amazonas: fide Schlechter, Fedde Repert. Beih. 9: 177. 1921.
Also Ecuador (type).
The Ames Herbarium contains sketches of the type from the
Reichenbach Herbarium.
Oncidium trulla Reichb. f. & Warsc. Bonpl. 2: 101. 1854; Lindl.
Fol. Orch. Oncidium 9, no. 24. 1855. Cyrtochilum trulla Kranzl.
SCHWEINFURTH: ORCHIDS OF PERU 911
Notizbl. Bot. Gart. Berl. 7: 96, in key. 1917, nomen; Kranzl. Engler,
Pflanzenr. IV. 50 (Heft 80), 66, fig. 6F, a-b. 1922.
Plant large, slender, forming dense clusters. Pseudobulbs aggregated, elongate-
ovoid to narrowly fusiform-cylindric, one- or two-leaved, commonly 3.5-5 cm.
high (6-8 cm. high, according to Kranzlin), clothed below with several pairs
of imbricating, evanescent, long sheaths (successively larger upward), which
are leaf-bearing. Leaves linear-oblong to linear or linear-oblanceolate, acute
or acuminate, up to 26 cm. long (40-45 cm. long, according to Kranzlin), about
0.8-1.5 cm. wide, very variable in size. Scape very long, 1 m. or more in length,
often climbing, loosely paniculate above; peduncle slender, up to 47 cm. long,
with several remote, very short, close sheaths; panicle with more or less remote,
short branches which are loosely few-flowered, the rachis strongly fractiflex
in age. Floral bracts minute, triangular, concave, much shorter than the slender,
pedicellate ovary which is 1.5-2.2 cm. long. Flowers very small, with widely
spreading segments, cinnamon-color or old rose with white tips. Dorsal sepal
oblanceolate, acute, with a recurved tip, 6-9.5 mm. long when expanded, 1.5-3 mm.
wide. Lateral sepals similar but rather longer and sometimes a little narrower,
connate near the base, dorsally carinate near the apex. Petals elliptic-ovate or
obovate-oblong, acute, shorter and broader than the dorsal sepal, lightly oblique.
Lip subequaling the petals, convex, simple, triangular in outline, subtruncate
at the broad base, abruptly acute above with an obtuse tip, having the outer
sub-basal angles more or less prominent and obtuse, about 5-7.6 mm. long and
nearly as wide at the base; disc at the base with a plurituberculate callus which is
tridentate in front, the central tubercle longer than the lateral ones. Column
very short, about 4 mm. high at the back, without prominent wings but with a
pair of small, fleshy angles below.
Huanuco: Yanano, about 1800 meters, on mossy trees, Macbride
SSUl. — Piura: Prov. of Huancabamba, near Huancabamba, 1800-
2000 meters, on rocky banks of stream between bushes, Weberbauer
2 ser. 6062. Also Colombia (type).
Oncidium ventilabrum Reichb. f. & Warsc. Bonpl. 2: 101.
1854; Lindl. Fol. Orch. Oncidium 6, no. 12. 1855; C. Schweinf. Bot.
Mus. Leafl. Harvard Univ. 14: 73. 1949. Cyrtochilum undulatum
HBK. Nov. Gen. et Sp. 1: 349, t. 84. 1816; Kranzl. Notizbl. Bot.
Gart. Berl. 7: 95, in key. 1917, nomen; Kranzl. Engler, Pflanzenr.
IV. 50 (Heft 80), 57. 1922. Oncidium undulatum Lindl. Sert. Orch. sub
t. 48, no. 1. 1842; Fol. Orch. Oncidium 7, no. 15. 1855; Reichb. f. &
Warsc. Bonpl. 2: 108. 1854, not 0. undulatum Salisb. 1812. Cyrto-
chilum ventilabrum Kranzl. Notizbl. Bot. Gart. Berl. 7: 92, in key.
1917, nomen; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 39, fig. 2C,
o-c. 1922.
Plant large, variable, with a stout, creeping rhizome which reaches 2 m.
in length. Pseudobulbs 5-30 cm. distant (according to Kranzlin), oblong-cylindric
912 FIELDIANA: BOTANY, VOLUME 30
to ovoid or ellipsoid, subcompressed, up to 8 cm. high, 1- or 2-leaved, clothed
below by 2 or 3 pairs of stout, imbricating, leaf-bearing sheaths which are suc-
cessively larger upward. Leaves very variable in shape and size, oblong to oblong-
oblanceolate (rarely the small outer ones subelliptic), commonly 19.5-60 cm.
long and 3-6 cm. wide, acute, usually long-narrowed to a conduplicate base.
Scape elongate, 1 m. or more long (4 m. long according to Kranzlin), twining,
loosely paniculate above, the branches (often compound in the lower portion)
very distant (up to 12 cm. or more apart), about 32 cm. or less long, loosely few-
(about 5-) to many-flowered, with the rachis lightly fractiflex. Floral bracts
relatively small, broadly triangular-ovate, deeply concave, much shorter than
the pedicellate ovary which is 2.5 cm. long. Flowers rather large, with wide-
spreading segments, brownish often with a narrow yellow border, the petals dull
yellow above. Dorsal sepal clawed, about 21-25 mm. long; lamina elliptic-
ovate to suborbicular, abruptly acute, about 16-18 mm. long, 12-16 mm. wide,
with the margins undulate; claw narrow and channelled below, exauriculate
at the base, rather gradually dilated above and passing into the lamina. Lateral
sepals similar, a little longer and narrower, subacute, slightly oblique. Petals
markedly smaller than the sepals, obliquely elliptic-ovate from a short, broad,
indistinct claw, acute or short-acuminate. Lip relatively small, prominently
recurved above, triangular-ligulate in outline, about 12-14 mm. long (8-9 mm.
long, according to Kranzlin) when expanded, rose outside, yellow with red and
white spots within, sessile at the subtruncate base, subacute at the complicate
apex, triangular-dilated or subquadrate-dilated (sometimes bilobed) and reflexed
on either side of the base, the anterior portion oblong-lanceolate or elliptic-
lanceolate; disc near the base with a large, plurituberculate callus terminating
in 3 narrow plates, with an undulate ridge curving onto the basal dilations.
Column small, sigmoid when viewed from the side, clavate, about 6 mm. high
at the back, adnate to the base of the lip, with a pair of small, decurved, falcate-
linear or lanceolate wings near the middle.
Peru: Habitat not recorded, Warscewicz s.n. (type of 0. ventila-
brum). — Junin, ex Kranzlin, fide Schlechter, Fedde Repert. Beih. 9:
177. 1921. Also Colombia (type of CyrtochUum undulatum).
The Ames Herbarium contains drawings of Oncidium ventilabrum
with floral analyses from the Reichenbach Herbarium.
Oncidium volubile (Poepp. & Endl.) Cogn. Martius Fl. Bras.
3, pt. 6: 289. 1905. CyrtochUum volubile Poepp. & Endl. Nov. Gen.
et Sp. 1: 35, t. 61. 1836; Kranzl. Notizbl. Bot. Gart. Berl. 7: 95, in
key. 1917, nomen; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 59.
1922. Oncidium corynephorum Lindl. Sert. Orch. sub t. 25. 1838;
Fol. Orch. Oncidium 8, no. 19. 1855.
Plant large, with an elongate, stout, creeping, simple rhizome. Pseudobulbs
commonly aggregated in scattering clusters, narrowly oblong-cylindric, attenuate
above, bifoliate, subcompressed, 4-5 cm. high, clothed below by a few pairs of
evanescent, long, leaf-bearing sheaths. Leaves linear-oblanceolate, acute or short-
SCHWEINFURTH: ORCHIDS OF PERU 913
acuminate above, more or less long-narrowed below to an often petiole-like,
conduplicate base, about 30-45 cm. long, 3-4 cm. wide. Scape elongate, climbing
or twining, stout, about 5-7 m. long, loosely paniculate above; branches remote
(about 19 cm. or less apart), simple, loosely few- (3- to 5-) flowered, with the
fractiflex rachis about 5-12 cm. long. Floral bracts small, triangular-ovate,
clasping, obtuse, much shorter than the pedicellate ovary which is up to about
2 cm. long. Flowers large, with wide-spreading segments. Sepals about equally
long, long- and narrow-clawed, free, about 20-26 mm. long, cinnamon-brown;
lamina of the dorsal sepal suborbicular-oval, obtuse, about 12-14 mm. wide;
lamina of the lateral sepals a little longer than that of the dorsal sepal, subacute,
lightly oblique. Petals distinctly shorter than the sepals, shortly and broadly
clawed; lamina obliquely ovate, acute, markedly narrower than the sepals, pale
cinnamon-brown with snow-white above the middle. Lip about half as long as
the sepals, pandurate-obovate, sessile, broadly rounded in front, 12-14 mm.
long, 12-13 mm. wide near the apex, rose-purple with the basal third white or
cinnamon-brown; disc at the base with a small, apically 3-dentate callus surrounded
by several free, short tubercles. Column small, slender, clavate, wingless, about
5 mm. high.
Huanuco: In the mountains of Cassapillo near Cuchero (Co-
chero), on stout trees, Poeppig 1636 (type of Cyrtochilum volubile).—
Loreto: Near Moyobamba, Mathews 1918 (type of 0. corynephorum).
The Ames Herbarium contains photographs of the type of
Oncidium corynephorum.
Oncidium Weber bauerianum Kranzl. Engler, Bot. Jahrb. 37:
389. 1906. Cyrtochilum Weberbauerianum Kranzl. Notizbl. Bot.
Gart. Berl. 7: 95, in key. 1917, nomen; Kranzl. Engler, Pflanzenr.
IV. 50 (Heft 80), 60, fig. 5H, a-c. 1922.
Plant large, with a stout, long-creeping rhizome. Pseudobulbs linear-cylindric,
compressed-ancipitous, up to 10 cm. high, unifoliate, up to 20 cm. distant, clothed
below by numerous, densely imbricating, long, leaf-bearing sheaths. Leaves on
the pseudobulbs oblong or oblanceolate-oblong, obtusely acute, long-complicate
below, up to 20 cm. long and 4.5 cm. wide. Scape (incomplete in the type) appar-
ently simple, not twining, below clothed with a few very remote, small sheaths,
terminating in a subdensely few- (about 12-) flowered raceme. Floral bracts
rather long, sheathing, about half as long as the pedicellate ovary which is 2.5-2.8
cm. in length. Flowers large, brown with yellow spots in the center. Dorsal
sepal elliptic-oblong ("oblong"), obtuse, sessile, lightly undulate on the margin,
15 mm. long, 10-12 mm. wide. Lateral sepals free, clawed, much (nearly twice)
longer; lamina similar to that of the dorsal sepal. Petals a little smaller than
the sepals, obovate-oblong, obtuse, lightly undulate. Lip about equaling the
dorsal sepal, pandurate-trilobed, 15 mm. long, about 5-7 mm. wide across the
basal and apical portions, yellow above the middle; lateral lobes relatively minute,
rounded, reflexed; mid-lobe cuneate, obscurely lobulate in front; disc at the base
with a prominent callus of which the posterior part is verrucose and obscurely
5-tuberculate and the anterior part dilated and lightly complicate with a reflexed
914 FIELDIANA: BOTANY, VOLUME 30
apex. Column very short, stout, above with broad, membranaceous wings con-
nate into a cup and below with a pair of small, oblong, porrect wings.
Amazonas: Molinopampa, east of Chachapoyas, 2000-2300 me-
ters, among sclerophyllous vegetation in open places, Weberbauer
4358 (type). Also Colombia.
No record of this species has been seen.
Oncidium xanthodon Reichb. f. Card. Chron. (1868) 1338;
Hook. f. Bot. Mag. 95: t. 5756. 1869. Cyrtochilum xanthodon Kranzl.
Notizbl. Bot. Gart. Berl. 7: 94, in key. 1917, nomen; Kranzl. Engler,
Pflanzenr. IV. 50 (Heft 80), 54. 1922.
Plant large, with a stout, creeping rhizome. Pseudobulbs separated on the
rhizome, oblong-ovoid to ellipsoid, compressed, unifoliate (bifoliate, according
to Kranzlin), about 10-12.7 cm. long, clothed at the base by several pairs of
densely imbricating, leaf-bearing sheaths which are successively longer upward.
Leaves oblanceolate, acute (acuminate, according to Kranzlin), long-narrowed
below to a complicate base, about 40-60 cm. long, 5-6.4 cm. wide. Scape elongate,
climbing, 2-2.7 m. or more long (5 m. long, according to Kranzlin), with distant,
appressed sheaths below, terminating in an elongate, loose panicle; branches
(lower ones compound) at least 11.5 cm. distant, loosely few- (about 3- to 5-)
flowered, with the rachis lightly fractiflex. Floral bracts ovate-lanceolate, acute
or acuminate, concave, much shorter than the pedicellate ovary which reaches
nearly 5 cm. in length. Flowers large, with wide-spreading segments which are
recurved above, chocolate-brown with yellow, crisped-undulate borders. Dorsal
sepal conspicuously clawed; lamina broadly ovate ("oblong"), acute, about 15-
20 mm. long and almost as wide near the broadly rounded or subhastate base;
claw stout, linear, about 5 mm. long. Lateral sepals similar but markedly longer,
with a somewhat longer claw, free. Petals similar to the dorsal sepal but markedly
smaller, with a short broad claw, acute. Lip smaller and narrower than the
petals, strongly recurved above, pandurate-trilobed or oblong-pandurate, about
12 mm. long, 5 or 6 mm. broad at the base when forcibly expanded, brown with
a yellow callus; basal portion (with reflexed lateral lobes) subquadrate or sub-
hastate, sessile, bilobed above; anterior portion (mid-lobe) cuneate-spatulate,
acute, recurved; disc at the base with a large, fleshy crest consisting of a central
rostrate ridge and several surrounding tubercles. Column small, sigmoid when
viewed from the side, adnate to the base of the lip, with a pair of minute wings
above.
Junin (ex Kranzlin), fide Schlechter, Fedde Repert. Beih. 9: 178.
1921. Ecuador (type) and Colombia.
No material of this species has been accessible.
Oncidium xanthornis Reichb. f. ex Linden Catal. no. 21: 16.
1867; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 201. 1922. Cyrto-
chilum xanthornis Kranzl. Notizbl. Bot. Gart. Berl. 7: 97, in key.
1917, nomen; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 72, fig.
7F, a-c. 1922.
SCHWEINFURTH: ORCHIDS OF PERU 915
Plant apparently medium-sized. Pseudobulbs, leaves, and the lower part
of the scape not described. Inflorescence paniculate, up to 25 cm. tall; branches
(sometimes compound) irregularly diverging, partly springing from the lower
part of the peduncle, fractiflex in age. Floral bracts very short, much exceeded
by the pedicellate ovary which is about 8 mm. long. Flowers small. Sepals
similar, cuneate-spatulate, retuse at the rounded apex, about 7 mm. long and
2.5 mm. wide above. Petals suborbicular, short-clawed, a little longer than the
sepals and more than twice as broad. Lip short-clawed, very broadly triangular
in outline, sublunate at the base, about 6 mm. long and 10 mm. wide below,
obscurely 3-lobed; lateral lobes large, rounded, recurved at the back; mid-lobe
shortly triangular, obtuse; disc with a crest consisting of 2 bipartite keels (the
anterior portion higher) with an interjected cluster of warts (the middle ones
longer). Column sigmoid, with a pair of large, dolabriform wings and a prominent
infrastigmatic plate.
Piura(?) : Huancabamba, Linden 4, 7. Ecuador, fide Kranzlin.
No record of this species was accessible.
Oncidium zebrinum (Reichb. f.) Reichb. f. Bonpl. 2: 12. 1854;
Gard. Chron. (1872) 1355, fig. 304; Hook. f. Bot. Mag. 100: t. 6138.
1874; Cogn. & Goos. Diet. Icon. Orch. Oncidium t. 9. 1897. Odon-
toglossum zebrinum Reichb. f. Linnaea 22: 849. 1849. Cyrtochilum
zebrinum Kranzl. Notizbl. Bot. Gart. Berl. 7: 94, in key. 1917, nomen;
Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80), 54. 1922.
Plant large, with a stout, long-creeping rhizome. Pseudobulbs more or less
large, ovoid, compressed, up to 12 cm. high according to Kranzlin, bifoliate, about
7-15 cm. apart, clothed at the base and often entirely concealed by several (about 5)
pairs of closely imbricating, leaf-bearing sheaths which are successively longer up-
ward. Leaves oblong or elliptic-lanceolate, acute or acuminate, apparently long-
narrowed to a complicate base, very variable in size, up to 30 or 40 cm. long,
2.5-4 cm. wide. Scape elongate, climbing, up to 4 m. long, trailing over bushes,
very loosely paniculate above; branches simple above and often branched near the
base, loosely few- to several-flowered, about 26 cm. or less long, about 14 cm. or
less apart, with a more or less fractiflex rachis. Floral bracts small, lanceolate-
ovate or ovate, acute or obtuse, deeply concave, much shorter than the pedicellate
ovary which is 2-4 cm. long. Flowers small for the section, with wide-spreading
segments. Sepals and petals similar, white with numerous transverse, violet or
brownish bars. Dorsal sepal elliptic-lanceolate or oblanceolate, acute or acumi-
nate, gradually unguiculate-narrowed below, strongly crisped-undulate on the
margins, 19-20 mm. long and 8-9 mm. wide (28 mm. long and 15 mm. wide,
according to Kranzlin). Lateral sepals similar (or oblanceolate- elliptic), slightly
longer and narrower than the dorsal sepal, free, lightly oblique. Petals oblong-
ovate, acuminate, oblique, crisped-undulate on the margin, markedly shorter than
the dorsal sepal and slightly narrower. Lip spreading from the base of the column,
then abruptly reflexed, triangular-lanceolate in outline, with the convex basal por-
tion irregularly triangular-dilated on each side, 13-15 (rarely 10) mm. long when
expanded, the anterior part oblong-lanceolate or ovate-lanceolate and acute or
acuminate; disc at the base with a yellow crest which is 3-lobed above and con-
916 FIELDIANA: BOTANY, VOLUME 30
sists of numerous blunt warts and a central, commonly high, thin, apically dentate
keel. Column small, arcuate or sigmoid, clavate, about 6 mm. high, often with a
pair of minute lanceolate wings near the middle.
Loreto (ex Kranzlin), fide Schlechter, Fedde Repert. Beih. 9: 178.
1921. Venezuela (type).
SIGMATOSTALIX Reichb. f.
(including Petalocentrum Schltr.)
Small to medium-sized epiphytes extending from Guatemala
south to Colombia, Brazil, Bolivia, Peru and Argentina.
Stems very short, rarely from a shortly creeping rhizome, pseudobulbous,
commonly aggregated, 1- or 2-leaved at the summit, surrounded below by several
pairs of distichous, leaf-bearing sheaths. Leaves narrow, elliptic-oblong to linear.
Inflorescences lateral, basal in the axil of the sheaths, shorter or more commonly
longer than the leaves, racemose or often paniculate with abbreviated branches.
Flowers very small to medium-sized, membranaceous, distant. Floral bracts small,
infundibuliform. Sepals wide-spreading or reflexed, free, subequal, subacute to
acuminate. Petals similar to the sepals, either broader or narrower. Lip sessile to
long-clawed, commonly simple or subsimple, either longer than broad or trans-
verse, callose near the base. Column elongate, very slender, dilated and often
auricled near the apex. Anther terminal, opercular, incumbent, 1-celled. Pol-
linia 2, complanate-ovoid, unappendaged, affixed to a linear-triangular stipe.
Al. Lip sessile or nearly so, transverse (i.e., broader than long); petals with a
spur-like horn near the base.
S. peruviana (S. pusilla, S. bicornuta, Petalocentrum spp.)
A2. Lip clawed or else distinctly narrowed toward the base; petals without a
horn I
la. Pseudobulbs bifoliate; anterior part of the lip multidenticulate on each side;
petals broader than the sepals S. aurosanguinea
Ib. Pseudobulbs unifoliate; anterior part of the lip not multidenticulate on each
side 1
la. Claw of the lip very narrow and elongate; lamina of the lip crescentiform ... 2
Ib. Claw of the lip broad and short or the base of the lamina long-narrowed;
lamina of the lip not crescentiform '. 3
2a. Dorsal sepal fornicate, concave-conduplicate; lip abruptly acute or apiculate,
unicallose at the base S. crescentilabia
2b. Dorsal sepal scarcely cucullate; lip acuminate, bicallose with approximate
calli at the base S. bicallosa
3a. Leaves narrowly linear, about 2 mm. wide; flowers very small, sepals about
2.5 mm. long; lip abruptly short-clawed, base of the lamina truncate.
S. graminea
3b. Leaves linear to elliptic-oblong, 4.5 (rarely) -14 mm. wide; flowers larger,
sepals about 5 mm. or more long; lip long-narrowed toward the base.
S. amazonica
Sigmatostalix amazonica Schltr. Beih. Bot. Centralbl. 42,
Abt. 2: 148. 1925.
SCHWEINFURTH: ORCHIDS OF PERU 917
Plant small, slender, variable, 7-25 cm. tall. Pseudobulbs aggregated, 1.4-
3.5 cm. high, strongly compressed, narrowly or broadly ellipsoid, unifoliate at the
apex, clothed on each side of the base by 1 or 2 pairs of imbricating, conduplicate,
leaf-bearing sheaths. Leaves linear to narrowly elliptic-oblong, acute or subacute,
more or less narrowed below to a complicate, petiole-like base, up to 15 cm. long
(commonly much less), 4.5-14 mm. wide. Inflorescence lateral, basal in the axil
of a sheath, erect or spreading, shorter than to much exceeding the leaf, about
5-18 cm. long; peduncle about 3-7 cm. long, with few, remote, close, tubular
sheaths; flower-cluster narrowly paniculate with subremote, abbreviated, 1- or
2-flowered branches. Flowers small, membranaceous, with wide-spreading seg-
ments, greenish white, olive or yellow spotted or streaked with brown, pink or lilac.
Dorsal sepal lanceolate-oblong, acute, 5-7 mm. long, 1.2-2 mm. wide. Lateral
sepals similar but a little larger, oblique. Petals similar to the dorsal sepal but
slightly shorter. Lip cuneate-obovate, broadly rounded at the apex with an irreg-
ular margin, 6-7.2 mm. long, 4-5.9 mm. wide near the apex, long-narrowed toward
the base which is occupied by a relatively large, ligulate-subquadrate (or elliptic),
apically truncate callus which is free above the base. Column long, very slender,
abruptly dilated near the apex, 4.9-7.2 mm. high.
Loreto: Middle Maranon River, at the mouth of the Apaya River,
about 145 m., Tessmann 3849. Soledad, on Rio Itaya, about 110 m.,
epiphyte in dense forest, Killip & Smith 29743. Vicinity of Iquitos,
100 m., Klug 10101. Also Brazil (Amazonas), type, and Surinam.
Sigma tostalix aurosanguinea Reichb. f. Linnaea 41: 68. 1876.
Plant very small, densely caespitose, apparently about 3.5 cm. high. Pseudo-
bulbs aggregated, ellipsoid, strongly compressed, bifoliate at the apex, apparently
about 1 cm. high. Leaves linear, acute, spreading or arcuate-recurved, appar-
ently about 3 cm. long. Inflorescence few-flowered. Floral bracts shorter than
the pedicellate ovary. Flowers yellow and blood-red. Sepals lanceolate ("linear")f
acuminate ("acute"). Petals broader, lanceolate-ovate. Lip about equally long
with the other segments but much wider, broadly clawed; lamina from a broadly
cuneate basal portion (claw) abruptly dilated into a larger, reniform anterior por-
tion which is retuse in front with a prominent apicule in the center, and with the
outer margins multi denticulate; disc apparently callose below. Column slender,
with an uncinate-decurved wing on each side of the apex.
Peru: On the summit of Mt. Guayraparina, on trees, Spruce s.n.
No example of this species has been available, but the Ames Her-
barium contains a drawing from the Reichenbach Herbarium bearing
the above data.
Sigmatostalix bicallosa Garay, Arquiv. Jard. Bot. 11: 57, t. 2.
1951. Figure 176.
Plant rather large for the genus, erect, slender, up to 30 cm. high. Rhizome
abbreviated. Pseudobulb narrowly ovoid, unifoliate, 2.5-3 cm. high, surrounded
at the base by 2 pairs of distichous, leaf-bearing, conduplicate sheaths. Leaves
linear-lanceolate, obliquely obtuse, apiculate, 8-12 cm. long, 6-7 mm. wide. Scapes
918
FIELDIANA: BOTANY, VOLUME 30
lateral as in the genus, erect; peduncle with a few imbricating sheaths; raceme dis-
tantly many-flowered. Floral segments reflexed-spreading. Flower rather large,
membranaceous. Sepals and petals pale yellow spotted with dark purple, lip
yellow with purple cilia. Dorsal sepal ovate-lanceolate or oblong-lanceolate,
scarcely cucullate, acuminate, 10 mm. long, 2 mm. wide. Lateral sepals obliquely
ovate-lanceolate, as long as the dorsal sepal but apparently somewhat broader.
Petals obliquely lanceolate-oblong, falcate, subacuminate, slightly smaller than the
FIG. 176. Sigmatostalix bicallosa Garay. 1, flower expanded; X 2J^. 2, lam-
ina of lip, showing calli; X 5.
dorsal sepal. Lip about as long as the sepals but much broader, long-clawed; claw
narrowly linear, subterete, about 4 mm. long; lamina cordate-reniform, deeply
cordate at the base, abruptly acuminate at the apex, with the margins (except on
the anterior third) ciliolate; disc near the base with 2 approximate, semirhombic
calli. Column erect, very slender, abruptly inflexed near the apex, about 10 mm.
high.
Peru: Chanchamayo, Rio Blanco, about 1500 meters, epiphytic
on coffee trees, Koehler s.n.
Sigmatostalix crescentilabia C. Schweinf. Am. Orch. Soc. Bull.
16: 162, t., 1947. Figure 177.
Plant large for the genus, very variable in size. Pseudobulbs oblong-ellipsoid
(sometimes narrowly so), strongly compressed, unifoliate, about 3-4 cm. high,
clothed below by about 3 pairs of imbricating, conduplicate, leaf-bearing sheaths
which are successively longer upward. Leaves oblong-linear to (rarely) elliptic-
oblong, subacute, more or less long-narrowed below with a conduplicate base, up
to 15.7 cm. long and 1 cm. wide, the blades on the outer sheaths often very small.
Scapes lateral, basal in the axils of sheaths, more or less surpassing the leaves,
slender, arcuate or spreading, about 17.5-37 cm. long; peduncle 6-12 cm. long,
with very few remote, close, tubular sheaths; inflorescence loosely several- to
many-flowered, up to 23.5 cm. long, racemose with each flower arising singly from
^ I GM ATQSTALIX
crescentifafiia G.
FIG. 177. Sigmatostalix crescentilabia C. Schweinf. 1, plant; X Vs. 2, flower
from side, natural position; X 3. 3, dorsal sepal, natural position; X 3. 4, petal;
X 3. 5, lateral sepal; X 3. 6, lip from above, expanded; X 5.
919
920 FIELDIANA: BOTANY, VOLUME 30
an abbreviated, bracted branch. Flowers large for the genus, membranaceous,
with reflexed sepals, petals and lamina of the lip. Dorsal sepal fornicate and con-
cave-conduplicate in natural position, when expanded elliptic-lanceolate (com-
monly narrowly so), acute or acuminate, up to 12 mm. long and 4 mm. wide.
Lateral sepals obliquely elliptic-lanceolate, acute or acuminate, a little shorter than
the dorsal sepal but wider. Petals obliquely linear-lanceolate, lightly falcate, acute,
a little shorter and markedly narrower than the sepals. Lip long-clawed, about
7.2-11 mm. long; claw very narrow and elongate, filiform-linear, channelled, 3.2-
5.9 mm. long; lamina semiorbicular-crescentiform, abruptly acute or mucronate at
the rounded apex, 7-9 mm. wide; disc at the base with a large, fleshy, ascending,
subcylindric (rarely subglobose) protuberance. Column very slender, arcuate,
somewhat dilated near the apex, with a large stigmatic orifice and a prominent
linear rostellum which is divergently bifid at the tip.
Junin: Chanchamayo Valley, 1200-1600 meters, Schunke 526,
1192, 1286, M (type).
Sigmatostalix graminea (Poepp. & Endl.) Reichb. f. Bot. Zeit.
10: 769. 1852; Xen. Orch. 1: 19, t. 8, fig. Ill, 11-17. 1854; Cogn.
Martius Fl. Bras. 3, pt. 6: 458. 1906. Specklinia graminea Poepp. &
Endl. Nov. Gen. ac Sp. 1: 51, t. 89 B. 1836.
Plant small, with a rather short, creeping rhizome which is much branched
and concealed by sheaths. Pseudobulbs numerous, approximate, very small, ob-
long-cylindric to cylindric-ellipsoid, strongly compressed, unifoliate, 7-12 mm.
high, clothed below by several pairs of imbricating, leaf-bearing sheaths. Leaves
small, narrowly linear, acute or subobtuse and obliquely emarginate at the apex,
lightly narrowed toward the concave base, mostly strongly arcuate-recurved,
3-5 cm. long, about 2 mm. wide. Scape basal in the axils of sheaths, about equal-
ing the leaves, erect or spreading, 3-6 cm. long, below with 1 or 2 short, acute,
appressed sheaths, above laxly racemose and few-flowered. Flowers short-pedi-
celled, very small, membranaceous, pale yellow with purple spots. Sepals sub-
equal, oblong or elliptic-lanceolate, acute, concave, 2-2.5 mm. long, about 0.66 mm.
wide, the lateral sepals subfalcate. Petals narrowly ovate, acute, oblique, about
as long as the dorsal sepal but slightly wider. Lip about 2.66 mm. long, shortly
and broadly clawed; claw about 0.66 mm. long and 1 mm. wide; lamina broadly
trapezoid or ovate-subquadrate, retuse in front, truncate with rounded angles at
the base, about 2 mm. long and 3 mm. wide; disc at the base with a small, sub-
orbicular, excavated callus which is longitudinally carinate through the middle.
Column short, very slender, clavate, up to 1.25 mm. high, with a pair of rounded-
oblong auricles above, the rostellum short-triangular and minutely bidentate.
Huanuco: Near Pampayaco (Pampayacu) and Cuchero (Cochero),
on trees, Poeppig 1007, 1514- Cerro de Ponasa, 1300 meters, Ule G.
I have seen no record of this concept.
Sigmatostalix peruviana Rolfe, Kew Bull. (1910) 371; C.
Schweinf. Bot. Mus. Leafl. Harvard Univ. 14: 76. 1949. Sigmato-
SCHWEINFURTH: ORCHIDS OF PERU 921
stalix pusilla Schltr. Fedde Repert. 10: 392. 1912. Sigmatostalix bi-
cornuta Rolfe, Kew Bull. (1913) 342. Petalocentrum pusillum Schltr.
Fedde Repert. 15: 145. 1918; Kranzl. Engler, Pflanzenr. IV. 50
(Heft 80), 312. 1922; Mansf. Fedde Repert. Beih. 58: t. 59, nr. 235.
1930. Petalocentrum angustifolium Schltr. Fedde Repert. 15: 145.
1918; Mansf. Fedde Repert. Beih. 58: t. 59, nr. 234. 1930. Petalo-
centrum bicornutum Schltr. Fedde Repert. Beih. 9: 179. 1921.
Plant very small, caespitose, epiphytic, commonly 5-7 cm. tall. Rhizome
short, sometimes branched. Pseudobulbs approximate, ovoid or ovoid-cylindric,
more or less compressed, about 1-1.5 cm. high, unifoliate, clothed below with 2
pairs of imbricating, leaf-bearing sheaths. Leaves linear or linear-oblong, sub-
obtuse or minutely bilobed at the apex, conduplicate at the scarcely narrowed
base, usually 3-5 cm. long, 1.5-3.5 mm. wide. Scapes lateral, basal in the axils
of sheaths, slender, loosely few-flowered above (often with abbreviated branches),
about 4-5 cm. high. Flowers very small, thin, with wide-spreading segments;
sepals and petals light yellow, dorsal sepal and petals with a deep purple band or
stripe. Dorsal sepal "oblong" to "linear-lanceolate" or oblong-lanceolate, acute
to subobtuse, 3-3.5 mm. long, about 1.2 mm. wide, with a recurved apex. Lateral
sepals similar, lightly oblique. Petals oblong or oblong-lanceolate, about equally
long with the sepals, more or less acute, near the base with a short, conical tooth.
Lip sessile or subsessile, deltoid-reniform or orbicular-reniform, lightly retuse in
the center of the broadly rounded, undulate, anterior margin, about 3-4 mm. long
and 4-5 mm. wide; disc at the base with a large, lobulate, often shining callus.
Column slender, clavate, 2-2.5 mm. high, with a pair of short, decurved auricles
at the apex.
Peru: Habitat not recorded, Forget s.n. (type of S. peruviana and
S. bicornuta). — Cuzco: Prov. of Quispicanchis, Cadena, 1020 meters,
epiphyte, Vargas 6141- — Junin: Chanchamayo Valley, Rio Blanco,
Hacienda St. Teresa, about 1400 meters, on coffee-trees, Kohler s.n.
(type of S. pusilla and Petalocentrum angustifolium).
LOCKHARTIA Hook.
Small to medium-sized epiphytes extending from Mexico and
Central America southward to Brazil, Bolivia and Peru.
Plant caespitose, without pseudobulbs. Stems simple, leafy. Leaves numer-
ous, distichous, equitant, imbricating, rather short, spreading to suberect, entirely
concealing the stem. Peduncles lateral in the axils of the leaves, commonly very
short and 1- or 2-flowered, racemose or more rarely loosely paniculate. Flowers
long-pedicelled, small or medium-sized, membranaceous with spreading or reflexed
segments^ Sepals free, subequal. Petals similar to the sepals or larger. Lip sim-
ple or deeply 3- or 4-lobed, equal to the sepals or longer, with the disc callose at
the base. Column very short and stout, broadly bialate or auricled above. Anther
terminal, opercular, incumbent, imperfectly 2-celled. Pollinia 2, ovoid, rather
long-acuminate.
922 FIELDIANA: BOTANY, VOLUME 30
Al. Lip simple to obscurely 4-lobulate; disc with an incurved horn in the middle.
L. hologlossa (L. unicornis)
A2. Lip deeply 3-lobed with linear or linear-lanceolate lateral lobes; disc without
an incurved horn in the middle L. amoena
Lockhartia amoena Endres & Reichb. f. Card. Chron. (1872)
666. Lockhartia costaricensis Schltr. Fedde Repert. 3: 81. 1906.
Plant small to large, epiphytic, caespitose, up to 50 cm. tall. Stems erect or
spreading from a more or less decumbent base, entirely concealed by the very
numerous, densely imbricating, distichous leaves. Leaves equitant, triangular to
triangular-lanceolate, commonly ascending (rarely slightly incurved), broadly ob-
tuse to subacute, 1.5-4 cm. long (measured on the lower margin), up to 1 cm. wide
across the base. Inflorescences racemose or more commonly loosely paniculate,
axillary, rather short. Floral bracts conspicuous, orbicular-cordate, usually apic-
ulate at the rounded apex. Flowers loose, few, small, yellow, lip with brown or red
markings. Dorsal sepal oval, more or less rounded at the apex, up to about 6 mm.
long and 4 mm. wide. Lateral sepals very similar. Petals ovate-oblong, usually
retuse or truncate at the apex, commonly rather larger than the sepals. Lip deeply
3-lobed, very variable, up to 9 mm. long in greatest length and 13 mm. wide across
the lateral lobes at the base; lateral lobes oblong-linear or lanceolate-linear, gently
recurved, more or less obtuse; mid-lobe much larger, more or less deeply 4-lobed,
with the lateral lobules spreading, semiorbicular to triangular and the narrower
anterior portion deeply bilobed forming a pair of obovate or broadly oblong lobules,
the entire margins more or less irregularly dentate; disc in the center of the base
with a subquadrate, excavated, fleshy callus produced into an oblong-subquadrate
mass of more or less distinct, verrucose lines. Column minute, stout, with a pair
of rounded-dolabriform wings.
Huanuco: Tingo Maria, 670 meters, epiphytic on trees, Carpenter
106. Also Mexico and Central America (Costa Rica, type).
Lockhartia hologlossa Schltr. Fedde Repert. Beih. 7: 199. 1920;
Mansf. Fedde Repert. Beih. 57: t. 71, nr. 274. 1929; C. Schweinf.
Bot. Mus. Lean1. Harvard Univ. 17: 57. 1955. Lockhartia unicornis
Schltr. Fedde Repert. Beih. 7: 200. 1920; Mansf. Fedde Repert.
Beih. 57: t. 71, nr. 275. 1929.
Plant epiphytic, or terrestrial on rocks, up to 53.5 cm. high. Stems caespitose,
erect-spreading from a decumbent base, entirely concealed by the numerous densely
imbricating leaves. Leaves equitant, narrowly triangular-lanceolate, erect-spread-
ing and often slightly incurved above, narrowed to an acute tip, the largest blades
(near the middle of the stems) about 3-6.5 cm. long (measured on the lower mar-
gin) and 5-8 mm. wide across the base. Inflorescence abbreviated, racemose, in
the upper or uppermost axils of the leaves, few- (1- to 6-) flowered. Flowers small,
yellow. Sepals concave, apiculate. Dorsal sepal ovate-elliptic, about 5 mm. long
and 3 mm. wide. Lateral sepals similar, elliptic-ovate, slightly larger throughout.
Petals broadly oval to oblong-oval, considerably larger than the sepals, broadly
rounded at the apex with a more or less distinct apicule. Lip suborbicular-ovate,
reduplicate in natural position, retuse in front, usually constricted on each side and
SCHWEINFURTH: ORCHIDS OF PERU 923
thus 3- or 4-lobulate (sometimes simple and merely retuse), broadly rounded at
the base, about 6.3-7.5 mm. long from the base to the apex of an anterior lobule
and 6.3-7 mm. wide near the base when expanded; disc through the center of the
lower half with a prominent, concave, oblong-ovate, fleshy callus which terminates
in an upcurved horn. Column very short and stout, with a pair of obliquely semi-
ovate, more or less lobulate wings. (The flowers of the typical forms from Colom-
bia were noted as either a little smaller or larger than the Peruvian specimens
described.)
Cuzco: Prov. of Quispicanchis, Ttio to Murayaca, Marcapata, on
rocks, 1960 meters, Vargas 3138. Prov. of Urubamba, Machu-
Picchu, on rocks, 2020 meters, Vargas 5546. — Junin: Huacapistana,
about 1800 meters, epiphyte in densely forested valley, Killip &
Smith 24%96 (plants in fruit, and therefore the determination is open
to some doubt). Also Colombia (type).
HOFMEISTERELLA Reichb. f.
A monotypic genus, confined to northern South America.
Plant small to medium-sized, epiphytic. Stems abbreviated, not pseudo-
bulbous, leafy. Leaves clustered, imbricating, distichous, linear. Scapes much
(often many times) exceeding the leaves, laxly racemose, ancipitous and narrowly
bialate, arcuate, more or less fractiflex. Floral bracts distant, conduplicate, erect-
spreading or incurved. Flowers relatively large for the plant, with long slender
pedicels. Sepals and petals similar, free, lanceolate-linear, strongly reflexed. Lip
triangular-lanceolate, simple, long-acuminate, sessile at the cordate or subcordate
base, erect-spreading, somewhat longer and much wider than the other segments.
Column footless, broadly 2-winged at the base, with a long, rigid, inflexed or de-
curved rostellum and an oblong clinandrium. Anther terminal, opercular, incum-
bent, 1-celled, produced into a linear-triangular appendage. Pollinia 4, complanate,
oblong-ovoid, affixed in pairs to the dilated apex of the long, slender stipe.
Hofmeisterella eumicroscopica (Reichb. f.) Reichb. f. Walp.
Ann. 3: 563. 1852; Xen. Orch. 1: 41, t. 18, figs. I, II, 1-8. 1854;
C. Schweinf. Bot. Mus. Leafl. Harvard Univ. 8: 50. 1940. Hof-
meistera eumicroscopica Reichb. f. De Poll. Orchid. Gen. ac Struct.
... 30. 1852.
Plant inconspicuous, epiphytic, with an abbreviated rhizome and large, stout,
fibrous roots. Stem very short, entirely concealed by the basal cluster of leaves.
Leaves several to numerous, deeply imbricating below, linear, equitant, acute, up
to 3.5 cm. long and 2 mm. wide in natural position. Scapes lateral in the axils of
the leaves, twice to many times longer than the leaves, up to about 27 cm. long,
suberect to arcuate, complanate and narrowly bialate, usually simple, few- to
many-flowered, laxly racemose, more or less fractiflex. Floral bracts distant, con-
duplicate, dorsally winged, spreading or incurved. Flowers rather small, mem-
branaceous, the segments greenish yellow or lemon-yellow with purple at the base
of the lip, the sepals and petals strongly reflexed. Dorsal sepal linear, condupli-
924 FIELDIANA: BOTANY, VOLUME 30
cate or with involute margins, acute or short-acuminate, 8.8-13.6 mm. long, 0.6-
0.8 mm. wide. Lateral sepals linear from a recurved base, slightly longer and
wider than the dorsal sepal. Petals very similar to the lateral sepals, a little shorter
than the dorsal sepal but somewhat wider than the lateral sepals. Lip triangular-
lanceolate, sessile, cordate or subcordate at the base, 10.8-14.4 mm. long, 4.4-
5.8 mm. wide near the base. Column small, recurved, with a pair of broad, semi-
ovate dilations at the base adnate to the lamina of the lip, about 4-5.7 mm. high
at the back, with an oblique clinandrium and a prominent, elongate, decurved
rostellum.
Cuzco: Prov. of Convention, Valle de Santa Ana, Herrera 3007. —
Huanuco: Yanano, about 1840 meters, on mossy tree, Macbride 3774.
Also Venezuela and Ecuador (type).
ORNITHOCEPHALUS Hook.
Epiphytic plants extending from Mexico through Central Amer-
ica, some of the West Indies and northern South America.
Plants small to medium-sized. Stems abbreviated, non-pseudobulbous in the
Peruvian species, entirely concealed by the deeply imbricating, conduplicate leaf-
sheaths. Leaves equitant, elliptic-oblong to linear, fleshy or coriaceous. Scapes
lateral in the axils of leaf-sheaths, simple, racemose above, commonly equaling or
exceeding the leaves. Flowers small, with wide-spreading segments. Sepals sub-
equal, free, concave. Petals commonly larger than the sepals. Lip sessile, simple
(and frequently inflexed above) or 3-lobed with the lateral lobes relatively small
and the mid-lobe (in the Peruvian species) commonly large. Column short, stout,
wingless, footless, and with a very long, subhorizontal rostellum. Anther terminal,
opercular, incumbent. Pollinia 4, affixed in pairs to the slightly dilated apex of
the linear stipe.
Al. Lip distinctly 3-lobed, with horizontally spreading lateral lobes; flowers large
for the genus I
A2. Lip simple, at most with a basal callus which sometimes extends beyond the
basal margins; flowers commonly small 1
la. Lateral lobes of the lip many times smaller than the mid-lobe, acute or acu-
minate, ecallose, paucidentate on the posterior margin near the apex.
O. patentilobus
Ib. Lateral lobes of the lip a little smaller than the mid-lobe, obtuse, entire, with
a pluri dentate callus extending across the disc of the lip O. polyodon
la. Outline of the lip ovate, with a conspicuous lobe-like apicule at the apex; disc
of the lip apparently ecallose at the base O. apiculatus
Ib. Outline of the lip oblong, cuneate or ovate-subquadrate, without a lobe-like
apicule at the apex; disc of the lip with a conspicuous basal callus 2
2a. Sepals smooth on the back 0. gladiolus
2b. Sepals setose on the back 3
3a. Lip more than twice as long as the sepals; basal callus many times shorter
than the lip O. longilabris
3b. Lip only slightly longer than the sepals; basal callus equaling about half of
the lip O. avicula
SCHWEINFURTH: ORCHIDS OF PERU 925
Ornithocephalus apiculatus Lindl. Ann. Nat. Hist. 4: 383.
1840.
Plant small, up to 7.6 cm. tall. Stem abbreviated, wholly concealed by sev-
eral dense, imbricating leaf-sheaths. Leaves obliquely linear-oblong. Scape much
exceeding the leaves (about three times as long), erect, densely several- to many-
flowered above. Flowers apparently deep yellow. Sepals not described. Petals
orbicular-obovate ("oblong"), broadly rounded above, coarsely dentate above the
middle. Lip ovate, entire, concave, with a conspicuous lobule-like, acute apicule
in front, sagittate at the base, markedly longer and narrower than the petals.
Column minute, with an elongate, recurved rostellum.
Peru: Habitat not recorded, Pavon s.n. (type). — Huanuco(?), fide
Schlechter, Fedde Repert. Beih. 9: 179. 1921.
The Ames Herbarium contains a photograph of this species with
an analysis of the petals and lip from the Lindley Herbarium.
Ornithocephalus avicula Reichb. f. & Wullschl. ex Reichb. f.
Walp. Ann. 6: 495. 1863; Cogn. Martius Fl. Bras. 3, pt. 6: 222, t. 45,
fig. 3. 1905.
Plant small for the genus, epiphytic. Stem abbreviated, entirely concealed by
numerous, conduplicate leaf-sheaths which are imbricating below and have a scari-
ous band on the upper margin. Leaves spreading, elliptic-lanceolate to oblong,
acute, often more or less oblique, 2- (rarely) 5 cm. long, 8-15 mm. wide, obliquely
articulated below, about 9 to 13 in each plant. Scape lateral in a leaf-axil, shorter
than or somewhat exceeding the leaves, laxly flexuous to arcuate-recurved, about
4-10 cm. long, racemose above, entirely glandular-setulose; peduncle short, with
1 or few spreading, amplexicaul sheaths; raceme loosely several- (7-) to many-
flowered, with a fractiflex rachis. Floral bracts semiamplexicaul, concave, ciliate-
dentate. Flowers very small, white and green (often various other shades), with
wide-spreading, membranaceous segments. Dorsal sepal cuneate-spatulate, con-
cave, rounded at the apex, ciliate-dentate above, subdensely setose without, about
3 mm. long and 2 mm. wide when expanded, 3-nerved. Lateral sepals similar but
slightly shorter, oblong-obovate, concave, broadly rounded at the apex, dorsally
setose, 1-nerved. Sepals sometimes smaller. Petals suborbicular-obovate, very
broadly rounded to subtruncate at the apex, irregularly dentate, 2-nerved with the
mid-nerve once branched, about 2.5 mm. long and slightly wider above. Lip
somewhat exceeding the rest of the perianth, broadly oblong-pandurate or pan-
durately ovate-subquadrate, slightly wider at the base, abruptly acute in the
middle of the broadly rounded or lightly retuse apex, subcordate at the sessile
base, 5- to 7-nerved, with the basal half entirely concealed by a large, fleshy, ovate-
suborbicular callus which is concave and 3- to 5-lobulate within, about 3.5-4.5 mm.
long and 2-3 mm. wide near the base. Column very small, fleshy-dilated below
and at the apex in front, about 1 mm. high at the back, with an elongate, recurved
rostellum.
Loreto: Vicinity of Iquitos, 100 meters, King 10022, 10023, 10108.
Also Surinam (type) and Brazil.
926 FIELDIANA: BOTANY, VOLUME 30
Ornithocephalus gladiatus Hook. Exot. Fl. 2: t. 127. 1824;
Cogn. Urban Symb. Antill. 6: 632. 1910. Ornithocephalus gladiatus
Hook. var. peruvianus C. Schweinf. Bot. Mus. Leafl. Harvard Univ.
14: 77. 1949.
Plant small for the genus, epiphytic. Stems very short, entirely concealed by
a cluster of leaf-sheaths which are deeply imbricating, conduplicate and have a
broad scarious border on the upper margin. Leaves oblanceolate or linear-oblan-
ceolate to oblong (rarely elliptic-lanceolate), abruptly acute, more or less oblique,
up to 6 cm. long and 7 mm. wide (the outer blades much smaller), about 9 to 12 to
a plant. Scapes lateral in the axils of leaf-sheaths, shorter than or markedly ex-
ceeding the leaves, glabrous and narrowly bialate, about 3-6 cm. long, suberect to
arcuate-recurved; peduncle short, with several amplexicaul, spreading sheaths;
raceme loosely few- (4- or 5-) flowered. Floral bracts amplexicaul, ovate, concave,
ciliolate. Flowers small, with membranaceous wide-spreading segments, white
with green markings. Dorsal sepal suborbicular, deeply concave, dorsally cari-
nate with the keel extended into a prominent mucro, glabrous without, about
3 mm. long (including the mucro) and 2.5 mm. wide. Lateral sepals very similar
but slightly shorter and narrower, oblique. Petals much larger than the sepals,
cuneate-obovate, abruptly dilated above, very broadly rounded at the apex, about
3 mm. long and broader above, 2- to 3-nerved with the mid-nerve carinate. Lip
simple, sharply inflexed in natural position, navicular-concave, linear-oblong when
expanded, slightly narrowed below, abruptly acute (rarely obtuse), about 5.7-
6 mm. long, up to 2 mm. wide near the apex, 3-nerved, the entire base concealed
(the margins more or less exceeded) by a large, suborbicular-ovate callus which
is cordate at the base and depressed-foveate in the center. Column very small,
dilated at the base, about 1 mm. high at the back, with a conspicuous, elongate,
decurved rostellum.
Hudnuco: Prov. of Hudnuco, Tingo Maria, on gravelly bank of
Rio Huallaga, epiphyte, Asplund 12372. — Junin: La Merced, about
610 meters, epiphyte in montana, Macbride 5480. Also Trinidad
(type), Grenada and Tobago.
Ornithocephalus longilabris Schltr. Fedde Repert. Beih. 9:
114. 1921; Mansf. Fedde Repert. 57: t. 130, nr. 508. 1929.
Plant epiphytic, rather large for the genus, about 10-12 cm. high. Leaves
8-10, erect-spreading, equitant, linear-elliptic ("linear-ligulate"), acute, sometimes
obliquely subfalcate, up to 10 cm. long and nearly 7 mm. wide in the middle.
Scapes in the axils of the leaf-sheaths, erect-spreading to arcuate, up to 9 cm.
long, entirely muriculate-villous; peduncle 4-5 cm. long; raceme rather loosely
6- to 12-flowered, up to 4.5 cm. long. Bracts broadly ovate, apiculate, ciliate-
dentate. Flowers medium-sized for the genus, thin, with wide-spreading segments.
Dorsal sepal concave, oblong-elliptic, subacute ("obtuse"), serrate-dentate ("sub-
crenulate"), sparsely setose on the back, 1-nerved, about 3.25 mm. long and
2.1 mm. wide. Lateral sepals similar but narrower, oblique. Petals obliquely
suborbicular-flabellate, about equally long and broad, slightly surpassing the sepals,
dentate on the broadly rounded apical margin, sparsely muriculate-papillose and
SCHWEINFURTH: ORCHIDS OF PERU 927
with an obtuse horn above the middle on the back. Lip narrowly cuneate-oblong
(lingulate), slightly dilated above and contracted below, abruptly narrowed to an
obtuse apex, 3-nerved, about 8.5 mm. long and 3 mm. wide above; disc at the base
with a short, broad, trapezoid-rhombic callus protruding beyond the lateral mar-
gins. Column very short, with an elongate, narrow rostellum.
Cajamarca: Habitat not recorded, Lehmann 3370.
No material of this species was available.
Ornithocephalus patentilobus C. Schweinf. Am. Orch. Soc.
Bull. 16: 510, t., 1947. Figure 178.
Plant large for the genus, epiphytic, up to about 14 cm. high. Stems abbre-
viated as in the genus, entirely concealed by the crowded, spreading leaf-sheaths
which are conduplicate and scarious-margined on the upper edge. Leaves 8-10,
equitant, spreading in a suborbicular cluster, narrowly elliptic-lanceolate to
linear-oblong or elliptic-oblong, acute (apiculate when young), more or less
narrowed to an oblique base, often lightly oblique above, 3.7-10.1 cm. long,
0.8-1.2 cm. wide. Scapes lateral in the axils of leaf -sheaths, spreading or arcuate,
subequaling or slightly exceeding the leaves, about 8.2-15 cm. long, setulose
throughout (very sparingly so on the basal portion), racemose above; peduncle
with 3-5 remote, small, amplexicaul, spreading sheaths; raceme about 4-13 cm.
long, loosely 6- to 18-flowered. Floral bracts small, spreading, amplexicaul,
broadly ovate, glandular-ciliate. Flowers rather large for the genus, with mem-
branaceous, wide-spreading or reflexed segments, "white and green, striped with
dark green (hairy)." Dorsal sepal strongly concave, oblong-obovate, rounded
at the apex, irregularly glandular-ciliate or ciliate-dentate, densely glandular-
setulose on the outer surface, about 5 mm. long in natural position and sub-
equally wide above. Lateral sepals similar, slightly oblique, as long as the dorsal
sepal but slightly narrower. Petals spatulate-obovate or oblong-obovate, broadly
rounded at the apex, irregularly dentate, longer and usually slightly wider above
than the dorsal sepal. Lip distinctly larger than the other segments, deeply
3-lobed near the base, about 8 mm. long and 9.8 mm. wide across the lateral
lobes when expanded; lateral lobes very small, erect in natural position, wide-
spreading, triangular-oblong, oblique and lightly retrorse, acute or acuminate,
with a few irregular teeth on the posterior margin toward the apex; mid-lobe
many times larger, broadly pandurate-oblong, broadly rounded in front, about
7 mm. long and 5 mm. wide near the apex; disc with the entire central basal
portion callose-thickened and pluridentate in front. Column small, about 2 mm.
high at the back, abruptly dilated at the apex in front, with an elongated, deflexed,
linear rostellum.
Loreto: Vicinity of Iquitos, 100 meters, epiphyte in dense forest,
King 10038.
Ornithocephalus polyodon Reichb. f. Linnaea 41: 33. 1876;
C. Schweinf. Bot. Mus. Leafl. Harvard Univ. 13: 151. 1948.
Plant rather large and stout for the genus, epiphytic, about 11-19 cm. tall.
Stems abbreviated, entirely concealed by 9-11 crowded sheaths of the leaves.
ORNITHOC^PHALUS
6cotomnandlip;
XV.. 2, flower
: X 4'
928
SCHWEINFURTH: ORCHIDS OF PERU 929
Leaf-sheaths imbricating, conduplicate, with a scarious border on the upper
edge. Leaves elliptic-lanceolate to linear-lanceolate, more or less acuminate,
commonly lightly oblique above, somewhat narrowed to the obliquely articulated
base, about 3 (outer blades) -16 cm. long, 0.85-1.4 cm. wide. Scapes lateral
in the axils of leaf-sheaths, usually 2 to each plant, suberect or flexuous to arcuate,
about 9.5-19.5 cm. long, entirely glandular-setulose, racemose above; peduncle
short, 4 cm. or more long, with 3 or more remote, spreading, semiamplexicaul
sheaths; raceme up to 14.5 cm. long, loose to subdense, several- (8-) to many-
flowered. Floral bracts spreading, suborbicular-ovate, semiamplexicaul, glandular-
ciliolate. Flowers rather large for the genus, with membranaceous, wide-spreading
segments. Dorsal sepal deeply concave, suborbicular-obovate, broadly rounded
at the apex, irregularly glandular-ciliate, rather densely glandular-setose on
the outer surface, about 6 mm. long and 4 mm. wide. Lateral sepals oblong-
obovate, broadly rounded above, lightly concave, sparingly serrate, rather densely
glandular-setose without, dorsally keeled, a little shorter and narrower than the
dorsal sepal. Petals cuneate-flabellate with recurved sides in natural position,
oblong-obovate when expanded, broadly rounded at the apex, shorter than the
sepals but subequally wide above. Lip deeply 3-lobed, shorter than the other
segments in natural position with a saccate base, erect lateral lobes and a recurved
mid-lobe, when expanded about 6.5 mm. long and slightly broader across the
base; lateral lobes small, divaricate, triangular-oblong, with a recurved, convex
apex; mid-lobe larger, subquadrate-ovate when expanded, rather abruptly narrowed
above to an acute tip; disc with a transverse, subelliptical, pluridentate callus
which extends as a toothed ridge from the base of the lateral lobes, continues
in a curved line in front of the lateral lobes and is extended across the base of
the mid-lobe. Column very small, clavate, with a reflexed, linear rostellum.
Loreto: Rio Mazan, 100-125 meters, Schunke 391. Also Ecuador
(type).
NOTYLIA Lindl.
A rather small but perplexing genus of very small to medium-
sized American epiphytes ranging from Mexico through Central
America and the West Indies to Brazil and Argentina.
Rhizome abbreviated. Stems very short, usually developed into pseudobulbs.
Pseudobulbs often clustered, small, unifoliate. Leaves commonly flat, linear to
oblong or elliptic, or rarely equitant. Scapes lateral, basal, usually arcuate
or recurved, racemose or rarely paniculate above. Flowers small, dense or lax.
Dorsal sepal free, lanceolate, oblong or ovate-oblong, strongly concave, usually
fornicate. Lateral sepals subequal, free or commonly more or less connate,
lightly oblique. Petals similar to the sepals, oblique. Lip commonly unguiculate;
lamina simple, ovate, lanceolate, pandurate or rarely obovate. Column stout or
usually slender, wingless, footless. Anther dorsal, deeply concave. Pollinia 2.
Al. Leaves equitant I
A2. Leaves flat, not equitant 2
la. Leaf-blades articulated to a sheath; lamina of the lip triangular-hastate
above N. Wullschlaegeliana
Ib. Leaf-blades not articulated to a sheath; lamina of the lip not triangular-
hastate 1
930 FIELDIANA: BOTANY, VOLUME 30
la. Leaves numerous, erect or incurved; sepals and petals setaceous-acuminate.
N. coffeicola
Ib. Leaves few, more or less recurved; sepals and petals acute or short-acuminate.
N. mirabilis
2a. Lamina of the lateral sepals deeply bidentate with sharply recurved apices;
leaves linear or nearly so 3
2b. Lamina of the lateral sepals minutely bidentate, without recurving tips;
leaves oblong (rarely linear-oblong) to elliptic-oblong or oval 5
3a. Lip sessile, without a distinct claw, long-acuminate N. rhombilabia
3b. Lip distinctly short-clawed, abruptly acute or apiculate 4
4a. Raceme dense; sepals about 4 mm. long; petals acute or short-acuminate;
lip subquadrate-ovate N. boliviensis
4b. Raceme loose; sepals about 3.5 mm. long; petals obtuse; lip ovate-suborbic-
ular N. peruviana (Dipteranthus peruvianus)
5a. Lip shorter than the column (usually distinctly so) 6
5b. Lip longer than the column (usually distinctly so) 7
6a. Leaf 1.4 cm. or less wide; lamina of the lip not sigmoid, cuneate at the base,
often gradually merging into the claw N. parvilabia
6b. Leaf 2.7 cm. or more wide; lamina of the lip strongly sigmoid when viewed
from the side, with an abrupt or subcordate base 2V. conduplicans
7a. Petals equally long with the lip; lip obtuse N. Koehleri
7b. Petals longer than the lip (often much longer) ; lip acute to long-acuminate . . 8
8a. Flowers fleshy; lip acute or short-acuminate; column stout, short, minutely
pubescent N. carnosiflora
8b. Flowers membranaceous; lip long-acuminate; column slender, glabrous.
N. Buchtienii (N. moyobambae)
Notylia boliviensis Schltr. Fedde Repert. Beih. 10: 53. 1922;
Mansf. Fedde Repert. Beih. 57: t. 141, nr. 553. 1929.
Plant slender, epiphytic, 10.5-12 cm. high. Rhizome abbreviated. Pseudo-
bulbs caespitose, slender, cylindric, compressed, unifoliate, 1-1.5 cm. high, mostly
concealed by several pairs of distichous, closely imbricating sheaths of which the
upper pair is leaf-bearing. Leaves oblong-linear to elliptic-linear, obtuse and
obliquely bilobed with an apicule, cuneate-narrowed to a conduplicate base,
6.3-11 cm. long, 0.7-1.7 cm. wide. Inflorescence lateral, basal, spreading, occa-
sionally once-forked; peduncle slender, 2-4 cm. long, with a few remote sheaths;
raceme (or panicle) about 8 cm. long, densely many-flowered. Bracts spreading,
linear-lanceolate, about equaling the pedicellate ovary. Flowers small, mem-
branaceous, light yellow, entirely glabrous. Dorsal sepal strongly fornicate,
concave, when expanded lanceolate-oblong, obtuse, dorsally conic-apiculate, nearly
4 mm. long, about 1 mm. wide. Lateral sepals connate into an oblong-elliptic,
concave lamina which is about as long as the dorsal sepal but twice as wide and
is sharply bidentate (often as far as the middle), with recurved apices. Petals
obliquely elliptic-lanceolate ("ligulate"), acute ("acuminate"), distinctly shorter
and narrower than the sepals. Lip very shortly clawed, about 3.5 mm. long;
lamina subquadrate-ovate, abruptly acute or apiculate, truncate at the base,
2.25-3 mm. wide; disc with a low carinate keel extending nearly to the middle
(the type description says the lip is ecarinate). Column cylindric, about 2 mm.
long.
SCHWEINFURTH: ORCHIDS OF PERU 931
Loreto: Vicinity of Iquitos, 100 meters, epiphyte beside the river,
King 10027. Also Bolivia (type).
Notylia Buchtienii Schltr. Fedde Repert. 10: 458. 1912; Mansf.
Fedde Repert. Beih. 58: t. 55, nr. 217. 1930; C. Schweinf. Bot. Mus.
Leafl. Harvard Univ. 12: 198. 1946. Notylia moyobambae Schltr.
Fedde Repert. Beih. 9: 108. 1921; Mansf. Fedde Repert. Beih. 57:
t. 127, nr. 498. 1929.
Plant epiphytic, very variable. Rhizome abbreviated. Pseudobulbs solitary
or often approximate, cylindric, compressed, unifoliate, 1.5-2.9 cm. long, invested
by several closely imbricating, distichous sheaths of which the upper 1 to 3 pairs
are leaf-bearing. Leaves elliptic to elliptic-oblong or oval (rarely oblong-oblance-
olate), broadly rounded and sometimes oblique at the apex, cuneate below to a
sessile or short-petioled, channelled base, 3-13 cm. long, 1.1-4 cm. wide, coriaceous.
Inflorescence spreading, arcuate or recurved, lateral, basal, 12-30 cm. long;
peduncle with several short, loose sheaths which are remote above and imbricating
at the base; raceme densely many-flowered, 7-19 cm. long. Floral bracts small,
linear-lanceolate, spreading or reflexed, much shorter than the pedicellate ovary.
Flowers small, membranaceous, pale yellow, or deep yellow and white dotted
with orange, entirely glabrous. Dorsal sepal strongly fornicate and concave,
when expanded lanceolate or oblong-lanceolate, acute to subacute, 5.5-7 mm.
long, 2 mm. wide. Lateral sepals connate into a deeply concave, oblong-lanceolate
or elliptic-lanceolate lamina which is shortly bidentate at the apex and is subequal
to the dorsal sepal or slightly longer and wider. Petals oblong-lanceolate, acute or
acuminate, slightly smaller than the dorsal sepal, more or less oblique and falcate.
Lip prominently clawed, about 5 mm. or less long; claw up to 1.6 mm. long,
linear, carinate-thickened; lamina triangular-lanceolate, long-acuminate, fleshy-
thickened, up to 2.4 mm. wide across the cordate to truncate or rounded base;
disc with a keel (extending from the claw) onto the lower part. Column cylindric,
slender, glabrous, more or less dilated at the base, reflexed near the apex, up to
3.1 mm. long.
Amazonas: Prov. of Bagua, Rio Utcabamba Valley, Cerro Tapur,
Hacienda Misqui, 1200 meters, Hutchison 151 7. — Loreto : Near Moyo-
bamba, Filomeno s.n. (type of N. moyobambae). Vicinity of Iquitos,
100 meters, in dense forest, King 10051, 10059. — San Martin: Tara-
poto, Alto Rio Huallaga, 360-900 meters, Williams 6183. Also Bo-
livia (type of N. Buchtienii).
Notylia carnosiflora C. Schweinf. Bot. Mus. Leafl. Harvard
Univ. 12: 199, t. 28, figs. 1-7. 1946. Figure 179.
Plant small. Rhizome abbreviated. Pseudobulb short, complanate-cylindric,
unifoliate, about 1 cm. high, invested by several distichous, closely imbricating
sheaths of which the upper pair is leaf-bearing. Leaves elliptic-oblong, rounded
at the apex, shortly cuneate-narrowed below with a complicate base, 4.6-6.2 cm.
long, 1.3-1.9 cm. wide, thickly coriaceous. Inflorescence lateral, basal, arcuate-
recurved, about equaling the leaves or shorter; peduncle about 3.5 cm. long,
932 FIELDI ANA: BOTANY, VOLUME 30
with a few sheaths which are remote above; raceme about 3 cm. long, subdensely
several-flowered, with small, reflexed bracts. Flowers small, reflexed, with spread-
ing fleshy segments. Dorsal sepal strongly fornicate and concave, oblong, acute,
about 7.5 mm. or less long in natural position, about 2.6 mm. wide when expanded.
Lateral sepals connate into an elliptic-oblong, tubular-concave lamina which is
minutely bidentate at the apex, about as long as the dorsal sepal but slightly
wider. Petals obliquely elliptic-lanceolate or oblong-oblanceolate, falcate-incurved,
acute, distinctly shorter and narrower than the dorsal sepal. Lip spreading from
the column at a right angle, very shortly clawed, about 3.5 mm. long; claw abbre-
viated, very fleshy, about 0.75 mm. long; lamina triangular^ovate, acute or short-
acuminate, subcordate at the base, about 2.9 mm. long, up to 2 mm. wide at the
base; disc with a fleshy, minutely pubescent callus (continued from the claw)
through the lower portion. Column short, stout, commonly dilated at the base,
about 2.9 mm. high, minutely pubescent.
Junin: Chanchamayo Valley, 1800 meters, Schunke s.n. (type in
Herb. Chicago Nat. Hist. Mus. No. 571652).
Notylia coffeicola Schltr. Fedde Repert. 10: 393. 1912; Mansf.
Fedde Repert. Beih. 58: t. 55, nr. 218. 1930.
Plant small, epiphytic, 3-8 cm. tall. Rhizome and stem much abbreviated,
entirely concealed by a suborbicular cluster of leaves. Leaves equitant, closely
and radially imbricating, numerous (11 or more), oblong-lanceolate, more or
less distinctly falcate-incurved, acute or short-acuminate with a minute mucro,
6-40 mm. long, up to 6.5 mm. wide, coriaceous. Inflorescences lateral, axillary,
spreading or recurved, commonly surpassing the leaves; peduncle filiform, with
several remote, small, loose sheaths; flower-cluster either an abbreviated, few-
to several-flowered, subcapitate raceme or a panicle consisting of a few congested,
appressed, few-flowered branches. Flowers large for the plant, spreading, with
membranaceous, glabrous, spreading segments. Dorsal sepal fornicate and deeply
concave, when expanded linear-lanceolate, setaceous-acuminate, about 9.5 mm.
or less long and 1.3 mm. wide. Lateral sepals linear, setaceous-acuminate, connate
near the base, a little longer (1.3 cm. or less) and narrower than the dorsal sepal.
Petals obliquely linear-lanceolate, setaceous-acuminate, very slightly longer than
the dorsal sepal and equally wide. Lip narrowly elliptic-lanceolate in outline,
8.2-9 mm. long, setaceous-acuminate, the basal portion narrowly unguiculate
with a pair of minute, fleshy auricles below the middle, the central portion rhombic-
lanceolate or ovate and about 2 mm. wide near the middle. Column slender,
cylindric, clavate above, about 4 mm. high.
Junin: Chanchamayo Valley, Rio Blanco, Hacienda St. Teresa,
about 1400 meters, on coffee trees, Koehler s.n. (type). Chancha-
mayo Valley, 1800 meters, Schunke 539. Same locality, 1500 meters,
Schunke 1213, 1214- Schunke Hacienda, above San Ramon, 1300-
1700 meters, in dense forest, Schunke A 51 .
Notylia conduplicans C. Schweinf. Bot. Mus. Leafl. Harvard
Univ. 12: 200, t. 25. 1946. Notylia Amesii Smith & Harris, Contrib.
N. mirafrids
C/
NOXYLJA
iji fo
carnos ora
FIG. 179. Notylia carnosiflora C. Schweinf. 1, plant; X 1. 2, flower; X 2.
3, dorsal sepal; X 4. 4, petal; X 4. 5, lateral sepals; X 4. 6, lip and column from
side; X 4. 7, lip from above, expanded; X 6. Notylia mirabilis C. Schweinf.
8, plant; X 1. 9, flower; X 5. 10, dorsal sepal; X 6. 11, petal; X 6. 12, lateral
sepal; X 6. 13, lip from above; X 6. 14, lip from side; X 6.
933
934 FIELDIANA: BOTANY, VOLUME 30
Gray Herb. Harvard Univ. 117 (1937), 40, in part (as to color and
Peruvian specimens). Figure 180.
Plant medium-sized, 28-33 cm. high, epiphytic. Rhizome abbreviated.
Pseudobulbs small, complanate-cylindric, unifoliate, about 1.5 cm. high, sur-
rounded and mostly concealed by several pairs of distichous, closely imbricating
sheaths of which the upper one or two sheaths are leaf-bearing. Leaves oblong
or elliptic-oblong, broadly rounded above (often with slightly unequal lobes),
more or less narrowed to a cuneate, conduplicate base, 8.2-18.2 cm. long, 2.7-3.1
cm. wide. Inflorescence lateral, basal, gently arcuate-spreading or deflexed;
peduncle about 9.5 cm. or less long, bearing several remote, short, tubular sheaths;
raceme 21-22 cm. long, densely many-flowered. Flowers reflexed, small, bright
green and white, glabrous throughout, with widely spreading, membranaceous
segments. Dorsal sepal strongly fornicate and concave, lanceolate-oblong or
elliptic-oblong, about 5.6 mm. or less long in natural position and 2.2 mm. wide
when expanded, obtuse, carinate above the middle. Lateral sepals connate
into a lanceolate-oblong, tubular-concave lamina which is very shortly bidentate
at the apex, a little shorter and narrower than the dorsal sepal. Petals obliquely
oblong, acute, somewhat shorter than the dorsal sepal and half as wide. Lip
adnate to the base of the column, about 3 mm. or less long, prominently clawed;
claw narrow, about 1 mm. long, with a high central keel extending onto the
middle of the lamina; lamina with conduplicate sides, recurved and sigmoid
when viewed from the side, when expanded ovate-cordate, up to 2 mm. in greatest
length, about 1.6 mm. wide. Column cylindric, abruptly reflexed near the apex,
3.1-3.9 mm. high at the back.
Loreto: Mishuyacu, near Iquitos, 100 meters, in forest, Klug 999.
Vicinity of Iquitos, 100 meters, in dense forest, Klug 10031 (type).
Also Venezuela.
Notylia Koehleri Schltr. Fedde Repert. 10: 393. 1912; Mansf.
Fedde Repert. Beih. 58: t. 55, nr. 220. 1930.
Plant low, epiphytic, about 6 cm. high. Rhizome abbreviated. Pseudobulbs
aggregated, oblong-cylindric, unifoliate, 1-1.5 cm. high, surrounded and more
or less concealed by several pairs of distichous, closely imbricating sheaths of
which the upper ones are leaf-bearing. Leaves oblong-elliptic or oblong, obtuse
to rounded and often unequally bilobed at the apex, cuneate below with a condu-
plicate base, 4-5 cm. long, about 8-12 mm. wide. Inflorescence lateral, basal,
arcuate, distinctly exceeding the leaves, about 10 cm. long; peduncle about 3.5 cm.
long, with several short, remote sheaths; raceme densely many-flowered, about
6.4 cm. long. Flowers small, reflexed, entirely glabrous, with spreading segments.
Dorsal sepal strongly fornicate and concave in natural position, when expanded
oblong-lanceolate ("narrowly oblong"), obtuse, about 6 mm. or less long and
1.6 mm. wide. Lateral sepals connate into a concave, elliptic-oblong lamina
which is lightly retuse at the apex, about as long as the dorsal sepal but broader.
Petals obliquely elliptic-lanceolate ("subfalcate-ligulate"), subobtuse, slightly
shorter and narrower than the dorsal sepal. Lip shortly clawed, about 5 mm.
long and equaling the petals; claw small, mostly occupied by a fleshy carinate
callus which extends onto the lower part of the lamina; lamina triangular-lanceolate
NO1 "YJ^I A.
cone) up fi cans &. Ocn we in ft.
FIG. 180. Notylia conduplicans C. Schweinf. 1, plant; X 4A- 2, flower from
side; X 4. 3, dorsal sepal; X 8. 4, petal; X 8. 5, lateral sepals; X 8. 6, column
and lip from side; X 8.
935
936 FIELDIANA: BOTANY, VOLUME ,30
or ovate-lanceolate, obtuse, rounded or subtruncate at the base, about 2.2 mm.
wide. Column cylindric, about 3 mm. high.
Junin: Chanchamayo Valley, Rio Blanco, Hacienda St. Teresa,
about 1400 meters, on coffee trees, Koehler s.n.
Notylia mirabilis C. Schweinf. Bot. Mus. Leafl. Harvard Univ.
12: 202, t. 28, figs. 8-14. 1946. Figure 179.
Plant very small, epiphytic, caespitose, about 2.5-4 cm. high. Rhizome
abbreviated. Stems abbreviated, entirely concealed by the flabelliform clusters
of leaves. Leaves equitant, closely imbricated, about 6 (3 on each side), obliquely
linear-oblong, linear-lanceolate or elliptic-linear, commonly strongly falcate-
recurved, acute to subobtuse, unjointed to sheaths at the base, about 1.4 cm.
or less long (from the apex to the base on the lower side), 1.5-3.5 mm. wide.
Inflorescence filiform, erect to slightly arcuate, much surpassing the clusters
of leaves, apparently 2.5-3.5 cm. long, provided near the apex with several ascend-
ing to strict, 1- to few-flowered branches. Floral bracts minute, ovate, concave.
Flowers small, thinly membranaceous, light lilac and dark violet, with widely
spreading segments. Dorsal sepal ovate-oblong or elliptic-ovate, acute, deeply
concave, dorsally carinate below, about 3.6 mm. or less long and 2 mm. or less
wide. Lateral sepals connate at the very base, obliquely elliptic-lanceolate,
acuminate, carinate near the base, longer and much narrower than the dorsal
sepal. Petals lanceolate, narrowed to an acute apex, about as long as the lateral
sepals and slightly wider. Lip about equaling the lateral sepals, prominently
clawed; claw relatively long and slender, fleshy, sub terete, somewhat dilated
above, about 1.3 mm. or less long; lamina concave, simple, obovate when expanded,
broadly rounded in front with a prominent apicule, auriculate at the base, about
2.8 mm. long and 2 mm. wide. Column slender below, very abruptly dilated
at the apex, about 2 mm. high.
Loreto: Vicinity of Iquitos, 100 meters, on dead tree, Klug 10112.
Notylia parvilabia C. Schweinf. Bot. Mus. Leafl. Harvard Univ.
12: 203, t. 26. 1946. Figure 181.
Plant small, caespitose. Rhizome abbreviated. Pseudobulbs small, aggregated,
complanate-cylindric, unifoliate, about 1.3 cm. or less high, invested and partially
concealed by several distichous, closely imbricating sheaths of which the upper
one or two on each side are leaf-bearing. Leaves oblong or oblanceolate-oblong,
bilobed at the apex with slightly unequal lobes, more or less narrowed below with
a conduplicate base, 4.8-11.3 cm. long, about 1.4 cm. or less wide, thickly cori-
aceous. Inflorescence lateral, basal, solitary, arcuate-recurved, more or less
surpassing the leaves; peduncle up to 7.5 cm. long, provided with a few short,
erect-spreading sheaths which are remote above and imbricating at the base.
Raceme densely many-flowered, incomplete in the specimens at hand. Floral
bracts short, linear-lanceolate, reflexed. Flowers small, reflexed, entirely glabrous,
with widely spreading, subfleshy segments. Dorsal sepal strongly fornicate and
concave, when expanded lanceolate-oblong, acute, about 5 mm. or less long
and 2 mm. wide. Lateral sepals connate into an oblong-lanceolate or lanceolate-
elliptic, deeply concave lamina which is minutely bidentate at the apex, and is
NOXYLJA
pari)i£a(sia. U. OcfiuveinJ.
FIG. 181. Notylia parvilabia C. Schweinf. 1, plant; X 1. 2, flower from side;
X 5. 3, dorsal sepal; X 8. 4, petal; X 8. 5, lateral sepals; X 8. 6, lip and column
from side; X 8. 7, lip from above, expanded; X 16.
937
938 FIELDI ANA: BOTANY, VOLUME 30
about as long and wide as the dorsal sepal. Petals lanceolate-oblong or oblance-
olate-oblong, subacute, more or less oblique, shorter and noticeably narrower
than the sepals. Lip small, adnate to the base of the column, distinctly shorter
than the column, conspicuously clawed, up to 2.2 mm. long; claw about 1 mm.
or less long, gradually merging into the lamina, with a high central keel which
extends to about the middle of the lamina; lamina rhombic-ovate with incurved
sides, narrowed to a subacute apex, about 1.4 mm. wide when expanded. Column
long, cylindric, abruptly recurved at the apex, about 3.5 mm. or less long, more
or less clavate.
Junin: Chanchamayo Valley, 1200 meters, Schunke 2054 (type in
Herb. Chicago Nat. Hist. Mus.). Same locality, 1500 meters,
Schunke 1106.
Notylia peruviana (Schltr.) C. Schweinf. Bot. Mus. Leafl. Har-
vard Univ. 12: 205. 1946. Dipteranthus peruvianus Schltr. Orchis 10:
187, t. 44, figs. 8-16. 1916.
Plant epiphytic, slender, up to 12 cm. high, with an abbreviated rhizome.
Pseudobulbs cylindric, slightly compressed, unifoliate, 1.3-1.5 cm. high. Leaf
erect, ligulate, subobtuse, slightly narrowed below, about 8.5 cm. long, nearly
1 cm. wide. Inflorescence pendent, 10-13 cm. long; peduncle slender, with a
few remote, clasping sheaths; raceme laxly many-flowered. Flowers small, pale
yellow with a yellow lip, glabrous. Dorsal sepal deeply concave, lanceolate-
oblong ("oblong-ligulate"), subobtuse, about 3.5 mm. long and 1.1 mm. wide.
Lateral sepals connate into a subequally long, oblong-elliptic, concave lamina
which is deeply bidentate with falcate-diverging, acute lobes. Petals obliquely
elliptic-oblong ("oblong-ligulate"), obtuse, cuneate below, about as long as the
sepals. Lip very shortly clawed, somewhat shorter than the sepals; lamina ovate-
suborbicular ("suborbicular"), lightly apiculate, subtruncate at the base, about
3.25 mm. long and 3 mm. wide below; claw and center of the lip below occupied
by a fleshy, carinate callus. Column slender for the genus, glabrous, lightly re-
curved, about 2 mm. high.
Peru: Habitat not recorded, Forget s.n. (sent to Sander & Sons,
St. Albans).
Notylia rhombilabia C. Schweinf. Bot. Mus. Leafl. Harvard
Univ. 12: 206, t. 27. 1946. Figure 182.
Plant medium-sized, slender. Rhizome apparently abbreviated. Pseudobulbs
narrowly complanate-cylindric, unifoliate, up to 1.8 cm. high, invested and
partly concealed by 2 or 3 pairs of distichous, deeply imbricating sheaths of which
the upper one or two on each side are leaf-bearing. Leaves linear or elliptic-
linear, subacute with a conspicuous caducous mucro, slightly cuneate-narrowed
below to a conduplicate base, up to 13.2 cm. long and 1.2 cm. wide (sometimes
very small). Inflorescence lateral, basal, sharply deflexed near the base, usually
much surpassing the leaves, subdensely racemose above, rarely once branched
below; peduncle about 11.9 cm. or less long, provided with several small, erect-
spreading sheaths which are remote above and imbricating at the base; raceme
NOT^YKIA
FIG. 182. NotyliarhombilabiaC.SchwemL 1, plant; X 4A- 2, flower, oblique
view; X 4. 3, dorsal sepal; X 4. 4, petal; X 4. 5, lateral sepals; X 4. 6, lip from
above, expanded; X 4. 7, lip and column from side; X 4. 8, anther; X 4.
939
940 FIELDIANA: BOTANY, VOLUME 30
10-13.6 cm. long, straight or flexuous. Floral bracts linear-lanceolate, the longest
ones (at the base of the raceme) slightly shorter than the pedicellate ovary.
Flowers reflexed, light green and white, entirely glabrous, with widely spreading
segments. Sepals subfleshy, lip fleshy. Dorsal sepal strongly fornicate and
concave, when expanded lanceolate-oblong, acute, about 8 mm. long, up to 3 mm.
wide. Lateral sepals connate into an oblong-elliptic, deeply concave lamina
which is sharply bidentate to above the middle with the lobules recurved at the
apex, a little shorter and broader than the dorsal sepal when expanded. Petals
linear, falcate-incurved, acute, about as long as the lateral sepals but much
narrower. Lip adnate to the column at the base, rhombic or ovate-rhombic
with a cuneate, sessile base and rounded, spreading sides, long-acuminate above
(often slightly incurved on each side), about 6 mm. long, up to 4.1 mm. wide;
disc with a low, callose keel extending from the base to about the middle. Column
cylindric, slightly tapering above, about 4.8 mm. high at the back, somewhat
sigmoid-recurved near the apex.
Loreto: Vicinity of Iquitos, 100 meters, epiphyte in dense forest,
Klug 10028.
Notylia Wullschlaegeliana Focke, Bot. Zeit. 11: 343. 1853;
Reichb. f. Xen. Orch. 1: 46. 1854; Cogn. Martius Fl. Bras. 3, pt. 6:
135, t. 38, fig. 3. 1904.
Plant very small, caespitose, epiphytic, about 5 cm. or less high. Rhizome
abbreviated. Pseudobulb very small, strongly complanate, oblong-ellipsoid to sub-
orbicular, unifoliate, about 4.5 mm. high, more or less concealed by 1 to several
pairs of distichous, deeply imbricated, leaf-bearing sheaths. Leaves in a semi-
orbicular-flabellate cluster, equitant, obliquely elliptic-lanceolate, sometimes lightly
falcate, acute to subacute, cuneate below, apparently very fleshy, up to 2 cm.
long and 5 mm. wide, all articulated to sheaths. Inflorescences lateral, basal,
filiform, suberect or arcuate, a little longer than the leaves; peduncle up to 4 cm.
long, provided with a very few small, remote sheaths; flower-cluster consisting
of a subumbellate, few-flowered raceme which is frequently at the apex of 2 or
3 strict, unequal branches. Floral bracts minute, ovate, much shorter than^the
widely spreading, filiform, pedicellate ovary. Flowers small but large for the
plant, greenish or whitish with purple spots, the floral segments widely spreading
and translucent. Dorsal sepal lightly fornicate and deeply concave, when ex-
panded linear-lanceolate, long-acuminate, about 7 mm. or less long and 1.8 mm.
or less wide. Lateral sepals free to the base, obliquely elliptic-linear, setaceous-
acuminate, about as long as the dorsal sepal but much narrower. Petals obliquely
linear-lanceolate, setaceous-acuminate, slightly shorter than the sepals and about
as wide as the lateral sepals. Lip adnate to the column above the base of the
latter, 5.5-6 mm. long, with a short, linear or narrowly cuneate, fleshy claw;
lamina about 5 mm. long, the lower third fleshy-auriculate at the base and oblong-
linear above, the middle portion abruptly semiorbicular- or ovate-dilated with a
cordate base and erose-denticulate sides, and the anterior part narrowly triangular
and setaceous-acuminate. Column filiform, abruptly recurved at the dilated apex,
5-6 mm. high.
Loreto: Vicinity of Iquitos, 100 meters, flowers silver-white, light
and dark lilac, Klug 10122. Also Surinam (type), and Costa Rica.
SCHWEINFURTH: ORCHIDS OF PERU 941
CRYPTARRHENA R. Br.
Small or medium-sized American epiphytes extending from Mex-
ico through Central America and the West Indies to Surinam,
Colombia, Ecuador and Peru. A single variable species now posi-
tively known in Peru.
Stems abbreviated (sometimes thickened into a unifoliate pseudobulb), often
clustered, mostly or entirely concealed by several distichous, deeply imbricated,
leaf-bearing sheaths. Leaves several to subnumerous, spreading, narrow, articu-
lated to sheaths. Inflorescences lateral, axillary, commonly more or less surpassing
the leaves, erect or arcuate, bearing several- to many-flowered racemes above.
Flowers small. Sepals subequal, free, spreading, the lateral ones oblique. Petals
similar to the sepals, often narrowed toward the base. Lip continuous with the
base of the column, spreading, 3- or 4-lobed, with a distinct, usually long, carinate-
thickened claw. Column short, thickened above, wingless, footless. Anther
opercular, incumbent. Pollinia 4, attached in pairs to the slightly dilated apex
of the linear stipe.
Al. All the lobes of the lip linear-falcate and subequal; petals linear-lanceolate.
C. acrensis
A2. Lobes of the lip consisting of 2 unequal pairs, the anterior lobes relatively
short and broad; petals obovate-oblong C. lunata
Cryptarrhena acrensis Schltr. Notizbl. Bot. Gart. Berl. 6: 126.
1914.
Plant rather small, stemless, epiphytic. Leaves ligulate, obliquely acute,
4-6 to a plant, up to 8 cm. long and 1.5 cm. wide. Scapes surpassing the leaves,
loosely racemose and many-flowered above, up to 12 cm. long. Flowers small,
whitish, with spreading or recurved segments. Sepals and petals linear-lanceolate,
acute or acuminate, glabrous, about 5 mm. long. Lip clawed; claw rather long,
with a rather high fleshy keel; lamina 4-lobed, produced in front into a mucro a
little shorter than the lobes; lobes linear-falcate, recurved, acuminate, about
equal. Column short, clavate, glabrous, footless, with a rather high clinandrium
and a lanceolate, acute, erect rostellum.
"Peru": Alto Acre Territory near the Auristella Rubber Planta-
tion, about 350 meters, Ule 89. This locality appears at present to
be within the boundaries of Brazil.
No material of this species was available.
Cryptarrhena lunata R. Br. Bot. Reg. 2: t. 153. 1816; Cogn.
Urban Symb. Antill. 6: 633. 1910; Kranzl. Engler, Pflanzenr. IV. 50
(Heft 80), 313. 1922; C. Schweinf. Bot. Mus. Lean. Harvard Univ.
8: 52. 1940. Clinhymenia pallidiflora A. Rich. & Gal. Compt. Rend.
Acad. Sci. Paris 18: 512. 1844, nomen. Orchidofunckia pallidiflora
942 FIELDIANA: BOTANY, VOLUME 30
A. Rich. & Gal. Ann. Sci. Nat. ser. 3, 3: 24. 1845. Cryptarrhena
Kegelii Reichb. f. Bot. Zeit. 10: 766. 1852; Cogn. Martius Fl. Bras.
3, pt. 6: 228, t. 94, fig. 1. 1905. Cryptarrhena pallidiflora Reichb. f.
Bot. Zeit. 10: 766. 1852; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 80),
314. 1922. Cryptarrhena unguiculata Schltr. Fedde Repert. Beih. 8:
103. 1921; Mansf. Fedde Repert. Beih. 57: t. 99, nr. 389. 1929.
Figure 183.
Plants small to rather large, caespitose, about 15-35 cm. tall. Stems very
short, entirely concealed by deeply imbricating, distichous sheaths which are
successively larger upward and commonly leaf-bearing. Leaves approximate,
about 3-7 to a plant, elliptic-linear to oblong-linear or oblanceolate-linear, acute
to subacute at the more or less oblique and sometimes bilobed apex, narrowed
to a complicate base, wide-spreading in flabelliform clusters, about 5-25 cm.
long, 0.5-2.5 cm. wide, the outermost blade usually much smaller. Scapes lateral
in the axils of the leaf-bearing sheaths, erect to arcuate, more or less surpassing
the leaves, subdensely racemose above, about 31.5 cm. or less long; peduncle
about 5-13 cm. long, with several remote, narrow, acute or acuminate sheaths;
raceme many-flowered, more or less elongate, about 8-23 cm. long. Flowers
very small, with wide-spreading segments, green (rarely whitish), yellow or
orange. Dorsal sepal oblong to elliptic-lanceolate, rounded to acute and apiculate
at the apex, concave, 4.1-5 mm. long, 1.6-2 mm. wide. Lateral sepals obliquely
elliptic-lanceolate, acuminate, subequally long with the dorsal sepal or a little
longer and slightly wider. Petals obovate-oblong, acute to rounded and apiculate
at the apex, distinctly shorter than the sepals but about as wide as the lateral
sepals or wider, with the upper margins irregularly and often minutely denticulate.
Lip long-clawed, somewhat exceeding the sepals, about 4.6-5.2 mm. long from
the base of the column to the apex of a spreading anterior lobule; claw linear,
wholly occupied by a high, fleshy callus, about 2-2.2 mm. long; lamina trans-
versely subquadrate or trapeziform in outline, deeply 3- or 4-lobed, sessile at the
broadly cuneate base, retuse to bilobed in front; lateral lobes from a broadly
triangular base extended into a linear, falcate-recurved lamina which is often
abruptly reflexed above; mid-lobe divided into 2, more or less spreading, relatively
short, subquadrate, triangular or lanceolate lobules which are more or less den-
ticulate on the apical margin. Column short, stout, somewhat dilated upward,
about 2.8-3 mm. high at the back, terminating in a large, erect, concave, denticu-
late clinandrium. Sometimes the flowers are recorded with markedly smaller
measurements.
Junin : Rio Pinedo, north of La Merced, 700-900 meters, epiphyte
in dense woods, Killip & Smith 23651. — Loreto: San Isidro, at the
mouth of the Pastaza River, 135 meters, Tessmann 4983. Vicinity
of Iquitos, 100 meters, epiphyte in dense forest, Klug 10086. A wide-
spread and variable species extending from Mexico (type of Orchido-
funckia pallidiflora) to Costa Rica, the West Indies (Jamaica, type
of Cryptarrhena lunata, and Trinidad) and to most of northern South
America (types of Cryptarrhena Kegelii and C. unguiculata} .
FIG. 183. Cryptarrhena lunata R. Br. 1, plant; X H- 2, flower, front-side
view; X 3. 3, lip, side view; X 4. Drawn by D. E. Tibbits. Adapted in part
from Fawcett and Rendle, Flora of Jamaica 1: pi. 30. 1910.
943
944 FIELDIANA: BOTANY, VOLUME 30
MACRADENIA R. Br.
A small genus of tropical American epiphytes extending from
Florida (U.S.A.) and Guatemala through Central America and in
South America southward to Brazil and Paraguay. But one species
appears at present to occur in Peru.
Plants medium-sized, with the habit and in the alliance of Notylia. Rhizome
abbreviated. Pseudobulbs aggregated, mostly narrow, 1-leaved. Leaves linear
to elliptic, more or less coriaceous. Scapes lateral, basal, suberect to nodding,
shorter than or surpassing the leaves, racemose above. Racemes laxly few-
to many-flowered. Sepals nearly equal, free, spreading. Petals similar, com-
monly a little smaller. Lip sessile and continuous with the column, prominently
3-lobed; lateral lobes erect, broad and embracing the column, relatively large,
entire or dentate; mid-lobe relatively small, lanceolate or linear, spreading.
Column stout, dilated upward, footless, provided with a conspicuous cup-like,
commonly lacerate-margined clinandrium and with a prominent rostellum. Anther
erect in the bottom of the clinandrium, oblong-cylindric, more or less decurved.
Pollinia 2, on a linear or triangular stipe.
Macradenia tridentata C. Schweinf. Am. Orch. Soc. Bull. 14:
62, t., 1945. Figure 184.
Plant medium-sized, with an abbreviated rhizome, up to 22.5 cm. high.
Pseudobulbs approximate, cauliform to very narrowly pyriform-cylindric, uni-
foliate, 1.5-4 cm. high. Leaves very variable, linear, oblong or narrowly elliptic-
oblong, acute or acuminate, cuneate-narrowed below to a short sulcate petiole,
9-15.7 cm. long, 3.5-18 mm. wide, subcoriaceous. Scapes lateral, basal, sub-
erect to arcuate-spreading, shorter than or somewhat exceeding the leaves,
11-24 cm. long, racemose above; peduncle with several scarious, infundibuliform
sheaths which are remote above and imbricated at the base; raceme loosely
several- to many-flowered, 4.7-15 cm. long. Floral bracts horizontally spreading,
triangular-lanceolate, acuminate, much shorter than the pedicellate ovary. Flow-
ers submembranaceous, with spreading segments. Dorsal sepal strongly con-
cave, when expanded linear-lanceolate or linear-elliptic, acute, 13.3-15 mm. long,
3.2-3.8 mm. wide. Lateral sepals similar but narrower. Petals similar to the
sepals, distinctly smaller, oblique. Lip shorter than the other segments, deeply
concave below, sharply 3-lobed near the middle, 8.3-9.3 mm. long, about 7 mm.
wide across the lateral lobes when expanded; lateral lobes erect, relatively large,
obliquely ovate or rhombic-ovate, acuminate, porrect; mid-lobe relatively small,
linear, rather fleshy, slightly protruding beyond the lateral lobes, 4.2-4.9 mm.
long, about 0.9 mm. wide; disc with a central linear (often indistinct) keel through
the lower half and with a pair of linear, flat intramarginal keels near the base.
Column stout, dilated upward, about 6.1 mm. high, with a cup-like, lacerate-
margined clinandrium which is produced in front into a pair of prominent, sub-
quadrate wings and with a prominent linear rostellum. Anther erect, protuberant,
oblong-cylindric, deeply sulcate, decurved.
Loreto: Above Pongo de Manseriche, Creek Torompisa, 220 me-
ters, flowers white, maroon-spotted, Mexia 6358a (type). Vicinity
MACRAD^NIA.
trie) en iaia G Ocfi we in ft.
~FlG, 184. Macradenia tridentata C. Schweinf. 1, plant; X %• 2, column and
lip, natural position; X 4. 3, lip expanded; X 4. 4, dorsal sepal; X 2. 5, petal;
X 2. 6, lateral sepal; X 2.
945
946 FIELDIANA: BOTANY, VOLUME 30
of Iquitos, 100 meters, "fls. greenish red-brown, white and dark vio-
let," King 10089.
TELIPOGON HBK.
Plants small to medium-sized, epiphytic or terrestrial, with a creeping rhizome
or decumbent lower portion of the stem. Stems very short or sometimes well-
developed, without pseudobulbs, sometimes branching above. Leaves few to
numerous, distichous, commonly narrow, coriaceous or fleshy. Peduncles terminal,
erect, often elongate, racemose above. Flowers one to several, medium-sized
or more commonly large and showy, sometimes very small. Sepals subequal,
free, wide-spreading, narrow. Petals commonly much larger and often transverse,
usually prominently veined and often reticulated. Lip very similar to the petals
and often broader (more rarely smaller), commonly simple, sessile. Column
usually abbreviated, footless, wingless, long-setose or hispid. Anther erect,
distinctly 2-celled. Pollinia 4, 2 separate ones in each cell, unappendaged, affixed
to the slightly dilated apex of the linear stipe.
Plants frequent in Costa Rica and in northern South America
(Venezuela, Colombia, Ecuador, Peru and Bolivia).
Al. Stems (if known) elongate; peduncles many times longer than the leaves. . .1
A2. Stems (if known) more or less abbreviated (at least not markedly elongate) ;
peduncles at most twice as long as the leaves 1
la. Petals 5-nerved; lip 7- to 9-nerved; flowers small, lip about 9 mm. long.
T. Benedicli
Ib. Petals 9- or 10-nerved; lip 11- to 13-nerved; flowers relatively large, lip about
2 cm. long T. Boissierianus
la. Lip distinctly trilobulate with a fold on each side of the apex, much smaller
than the petals T. Fritillum
Ib. Lip simple, commonly larger (at least broader) than the petals 2
2a. Petals and lip oblong-elliptic, much (nearly twice) longer than broad.
T. gymnostele
2b. Petals and lip not oblong-elliptic, usually little longer than broad or broader
than long 3
3a. Lip about 49-nerved; petals urceolate, abruptly angle-dilated above.
T. urceolatus
3b. Lip about 25-nerved or less; petals not urceolate nor abruptly dilated above. . . 4
4a. Anterior portion of the lip distinctly narrowed toward the apex 5
4b. Anterior portion of the lip not distinctly narrowed toward the apex 8
5a. Lip reticulate-veined throughout; column relatively long T. Vargasii
5b. Lip not reticulate-veined (i.e., without cross-reticulation) ; column very short. . . 6
6a. Petals 15- or 16-nerved; lip 1 9-nerved T. rhombipetalus
6b. Petals 7- to 11-nerved; lip 15- (rarely 17-) nerved 7
7a. Lip suborbicular-ovate, distinctly narrowed in front, subequally long and
broad; petals 9- to 11-nerved T. latifolius
7b. Lip transversely oblong-oval, only slightly narrowed in front, much broader
than long; petals 7- to 9-nerved T. gnomus
8a. Petals and lip wholly conspicuously reticulated (with prominent veins and
horizontal reticulations) T. tessellatus
SCHWEINFURTH: ORCHIDS OF PERU 947
8b. Petals and lip without cross-reticulations, or rarely (T. Hercules) with occa-
sional obscure, oblique reticulations 9
9a. Flowers relatively large, lip about 3.3 cm. long and 4 cm. wide . . .T. Hercules
9b. Flowers smaller, lip 2.6 cm. or less long and 3.7 cm. or less wide 10
lOa. Lip 21- to 25-nerved 11
lOb. Lip 13- to 17-nerved 12
lla. Petals 13- to 15-nerved; lip about 25-nerved T. Papilio
lib. Petals 9- to 11-nerved, about 1.9-2.2 cm. long; lip 21-nerved . . . . T. radiatus1
12a. Petals much longer than wide; lip very broadly obovate or suborbicular,
nearly as long as wide T. obovatus
12b. Petals little longer than wide, about equally wide, or wider than long; lip
transversely oblong-oval, much wider than long 13
13a. Flower large, petals 2.2-2.7 cm. long; lip 2.8 cm. or more wide. . .T. gnomus
13b. Flower small, petals 1.5-1.7 cm. long; lip 2 cm. or less wide . . . .T. semipictus
Telipogon Benedict! Reichb. f. Linnaea 41: 3. 1876; Kranzl.
Ann. Naturh. Hofmus. Wien 33: 29. 1919.
Stems elongate, leafy, sometimes branched with short strict branches, mostly
concealed by the sheathing bases of the leaves, bearing more or less numerous,
scattered simple roots which are stout, fibrous and glabrous. Leaves small, very
numerous, distichous, spreading-ascending, elliptic-oblong or ovate-oblong ("lig-
ulate"), acute (with a recurved apex), amplexicaul and sheathing the stem through
the lower part, up to 3.7 cm. long and 1 cm. wide. Inflorescence ascending, loosely
racemose above; peduncle up to 10 cm. long, with 1 or 2 tubular-amplexicaul
sheaths below; raceme few- (4- to 6-) flowered, up to about 6 cm. long, with a
flexuous or fractiflex rachis. Floral bracts spreading, amplexicaul, conduplicate,
triangular-ovate, obtuse, dorsally carinate. Pedicellate ovary trialate. Flowers
relatively very small, with wide-spreading segments; sepals green, petals and lip
green with dark brown nerves or the "perianth orange-yellow, purple ribbed."
Dorsal sepal oblong-ovate ("oblong-triangular"), acute, short-apiculate, strongly
concave, 3-nerved, 7-10 mm. long, 3-5.6 mm. wide when forcibly expanded, ecari-
nate. Lateral sepals similar, but somewhat narrower. Petals suborbicular-ovate
or oval-ovate ("elliptic"), acute, apiculate, 5-nerved with an additional small nerve
on each side below, about as long as the dorsal sepal and 6-8 mm. wide. Lip very
similar to the petals, subacute, 7- to 9-nerved, about 8-10 mm. long and wide.
Column very short, triangular-rostrate and short-pubescent in front, densely setu-
lose in the rear.
Cuzco: Prov. of Convention, "alturas de Pintobamba," 2700 me-
ters, epiphytic in forest, Vargas 3544- Prov. of Paucartambo, Pilla-
huata, 2800 meters, epiphyte, Vargas 6107. Also Bolivia (type).
Telipogon Boissierianus Reichb. f. Bonpl. 4: 213. 1856; Xen.
Orch. 1: 229, t. 97, fig. I. 1858; Kranzl. Ann. Naturh. Hofmus. Wien
33:32. 1919.
'This species was described from only a single flower, and no authentic record
was available.
948 FIELDIANA: BOTANY, VOLUME 30
Stem elongate, slightly decumbent below, leafy, with a few short, strict branches,
more or less concealed by the sheathing bases of the leaves, producing in the lower
portion occasional roots which are fibrous, glabrous, simple, sulcate and finely
verruculose when dry. Leaves very numerous, distichous, spreading-ascending,
elliptic-oblong, acute with a recurved tip, amplexicaul by a long-sheathing, mem-
branaceous, basal portion, up to 2.5 cm. long and 7 mm. wide, the free portion
(up to 1.4 cm.) cartilaginous, ovate to oblong-ovate. Inflorescences strict to spread-
ing-ascending, racemose above, about 6^9.5 cm. long; peduncle with a single close,
tubular sheath near the middle or below the middle; raceme short, 3-flowered;
rachis flexuous, about 2-3.5 cm. long. Floral bracts spreading, amplexicaul, lan-
ceolate-ovate, concave. Flower small for the genus, with spreading segments, "dull
yellow-brown dotted and veined," with a purple pubescent center. Dorsal sepal
oblong-lanceolate or ovate-oblong, strongly concave below, abruptly acute, mucro-
nate, 3-nerved, about 1.6 cm. long and 5.6 mm. wide, dorsally subcarinate. Lateral
sepals similar, oblong-lanceolate, slightly shorter and narrower than the dorsal
sepal. Petals much larger, rhombic-ovate, acute, slightly oblique, about 10-nerved,
about 1.8 cm. long and 1.5 cm. wide. Lip very similar to the petals but larger,
suborbicular-ovate, acute, about 11- to 13-nerved, about 2 cm. long and 1.8 cm.
wide. Column very short and stout, finely pubescent in front, densely setose in
the rear. Sometimes the flower has smaller measurements.
Huanuco: Playapampa, about 2700 meters, on wet rocky sphag-
num slope, Macbride 4888. — De la Cunta (Cuerta?) de Saria, Herb.
Pavon (type).
Telipogon Fritillum Reichb. f. & Warsc. Bonpl. 2: 101. 1854;
Xen. Orch. 1: 230, t. 97, fig. III. 1858; Kranzl. Ann. Naturh. Hofmus.
Wien33:16. 1919.
Plant low, 7.6-12.7 cm. high. Stem very short, provided with a few congested
leaves. Leaves oblanceolate, apiculate, minutely undulate, up to 5 cm. long and
6 mm. wide (according to Kranzlin). Inflorescence straight or curved, up to 10 cm.
long, racemose above; raceme 1- to 3-flowered. Floral bracts triangular-ovate,
acute or acuminate, dorsally carinate. Pedicellate ovary rather broadly tripter-
ous, up to 4 cm. long, much surpassing the bracts. Sepals triangular-ovate or
triangular-lanceolate, acute or acuminate, 3-nerved, the lateral sepals lightly
oblique, about 1.5 cm. long and 6 mm. wide. Petals much larger, obliquely
rhombic-ovate or suborbicular-ovate, very shortly acute or apiculate, 11- to 13-
nerved, about 1.8 cm. long and broad. Lip much smaller than the petals, trans-
versely oval or suborbicular in outline, with a fold on each side of the apex (thus
3-lobulate), broadly rounded in front in the middle, more or less ciliate, 21- to
23-nerved, 1-1.2 cm. long and broad (according to Kranzlin). Column probably
abbreviated, very densely long-setose. (The measurements were mostly taken
from Kranzlin, I.e.)
Peru: Habitat not recorded, Warscewicz s.n.
No example of this species was available.
Telipogon gnomus Schltr. Fedde Repert. Beih. 9: 114. 1921;
Mansf. Fedde Repert. Beih. 57: t. 130, nr. 509. 1929.
SCHWEINFURTH: ORCHIDS OF PERU 949
Plant very variable, from about 6 to 22 cm. high. Stems rather short, pro-
cumbent or flexuous, commonly shortly branched, entirely concealed by the imbri-
cating, often marcescent sheaths of the leaves and bearing numerous, scattered
roots which are fibrous, glabrous, with a more or less enlarged, shining velamen.
Leaves several, congested on each short branch, oblong-elliptic to broadly or nar-
rowly oblanceolate, acute and short-apiculate ("obtuse"), more or less narrowed
to a complicate, sessile or petiole-like base, 2.5-10 cm. long, about 5 mm.-2 cm.
wide. Scapes slightly shorter than the leaves or somewhat exceeding them, 2-
17 cm. long; peduncle naked, subcompressed and very narrowly bialate above;
raceme 1- to 7-flowered, up to about 6.5 cm. long, lightly fractiflex. Floral bracts
spreading, conduplicate, triangular or ovate-triangular, dorsally carinate, subacute
to acuminate, much shorter than the trialate pedicellate ovary. Flowers variable,
medium-sized to large for the genus, with wide-spreading, membranaceous seg-
ments, pale yellow with brown-red or purple lines. Dorsal sepal lanceolate or
oblong-lanceolate, acute or acuminate, dorsally strongly carinate with the keel
produced as a mucro, navicular, 3-nerved, about 1.7-2.3 cm. long and 4.5-7 mm.
wide. Lateral sepals similar, slightly narrower, oblique. Petals much larger, obo-
vate-rhombic to suborbicular-rhombic, commonly rather sharply acute and apicu-
late, 7- to 9-nerved, 2-2.7 cm. long, 1.8-2.4 cm. wide. Lip transversely oblong-oval
("subreniform-flabellate"), minutely apiculate at the broadly rounded or slightly
narrowed apex, cuneate-rounded at the base, 15- to 17-nerved, 2.2-2.6 cm. long,
2.75-3.2 cm. wide. Column very short and stout, with a tuft of numerous decid-
uous bristles (often 2- or 3-forked) in the rear.
Cajamarca: Prov. of Chota, pass south of Conchan, 2500 meters,
terrestrial in loose humus, Stork & Horton 10082. — Huanuco: Hua-
cachi (near Muna), in montana, about 2000 meters, Macbride 4114-
Panao, 3100 meters, Zimmer s.n. 12 miles south of Panao, about
3000 meters, epiphyte on shrub in shade, Macbride & Featherstone
2224- — Loreto: Near Moyobamba, Filomeno s.n. (type).
Telipogon gymnostele Reichb. f. Linnaea 41: 70. 1876; Kranzl.
Ann. Naturh. Hofmus. Wien 33: 16. 1919.
Plant small, about 6 cm. high including the flowers. Stem short, about 1 cm.
high, apparently somewhat branched (judged from Reichenbach's record). Leaves
rather numerous, congested, elliptic-oblong ("oblong"), articulated to sheaths,
acute or obtuse, about 2.5-3.5 cm. long and 7 mm. wide. Inflorescence including
the flowers slightly surpassing the leaves; peduncle very short, much shorter than
the leaves; raceme 2-flowered. Floral bracts triangular or ovate, amplexicaul, dor-
sally carinate, much shorter than the pedicellate ovary which is slender, trialate
and about 2 cm. long. Sepals elliptic-lanceolate or oblong-lanceolate ("lanceo-
late"), acute, lightly carinate, 1-nerved throughout or 3-nerved below the middle,
1-1.5 cm. long, 3-4 mm. wide. Petals oblong-elliptic ("oblong"), acute, 5-nerved
with a few oblique reticulations, about 1.2-1.7 cm. long and nearly twice as wide
as the sepals. Lip oblong-elliptic ("oblong"), more or less acute, 7-nerved, with
one of the lateral nerves branching on each side and the nerves velutinous near the
base, about 1.2-1.5 cm. long, 7-8 mm. wide. Column very short and stout, gla-
brous, with a conspicuous curved rostellum.^ The measurements were taken in
part from Kranzlin (I.e.) and in part from a drawing of the type.
950 FIELDIANA: BOTANY, VOLUME 30
Peru: Pustaco(?), 3300 meters, Warscewicz s.n.
No material of this species was available, but there is a drawing
of the type with a floral analysis in the Ames Herbarium.
Telipogon Hercules Reichb. f. ex Kranzl. Ann. Naturh. Hof-
mus. Wien 33: 27. 1919; C. Schweinf. Bot. Mus. Leafl. Harvard
Univ. 17: 57. 1955.
Plant medium-sized, about 20 cm. or more high. Stems abbreviated. Leaves
several, congested, obovate-oblong to oblanceolate, acute or subacute, more or less
long-narrowed to a complicate base, 4.5-10 cm. long, up to 2 cm. wide. Inflores-
cence (excluding the flowers) about 15-33 cm. long; peduncle flexuous, naked,
apparently angulate in section; raceme very loosely few- (about 3-) to several-
flowered, the rachis fractiflex and about 5 cm. high. Floral bracts conspicuous,
spreading, conduplicate, dorsally carinate, triangular-ovate when expanded, up to
1.7 cm. long. Flowers very large, with spreading segments, golden yellow with
brown nerves. Dorsal sepal oblong-lanceolate, acute or acuminate, dorsally cari-
nate, 3-nerved with a short outer nerve on each side, strongly concave or navicular,
about 2.6 cm. long and 9.6 mm. wide. Lateral sepals similar but a little narrower,
lightly oblique. Petals much larger, rhombic-suborbicular, broadly acute and
apiculate, 13-nerved with oblique reticulations, about 3.3 cm. long and equally
broad. Lip transversely suborbicular-oval, minutely apiculate at the broadly
rounded apex, about 3.3 cm. long and 4 cm. wide, 17-nerved. Column very short
and stout.
Northern Peru and Ecuador: Habitat not recorded, Roezl s.n.
(type). Also Colombia.
Telipogon latifolius HBK. Nov. Gen. et Sp. 1: 336. 1816; Cogn.
Martius Fl. Bras. 3, pt. 6: 109, t. 37, fig. 1. 1904; Kranzl. Ann.
Naturh. Hofmus. Wien 33: 18. 1919.
Plant extremely variable in size, epiphytic, about 2.5-16 cm. high (excluding
the flowers). Stems short, decumbent, simple or with abbreviated, erect branches,
concealed by the imbricating leaf-sheaths, the lower portion (producing the roots
and leaves) about 1-4 cm. long. Leaves (present) commonly 4 or 5, congested,
linear-oblanceolate or obovate-oblong to elliptic-oblong, up to 9 cm. long and 2 cm.
wide (often minute), abruptly acute, more or less narrowed below to the articula-
tion with the conduplicate leaf-sheaths. Inflorescence (excluding the flowers)
about 1-15 cm. long, erect to ascending or arcuate, commonly about twice longer
than the leaves; peduncle about 10.5 cm. or less long, naked except for the sheathed
base, stout, narrowly bialate above; raceme 1- to 7-flowered, loose; rachis about
4 cm. or less long. Floral bracts spreading, triangular, conduplicate, dorsally cari-
nate, acute or acuminate, much shorter than the trialate pedicellate ovary. Flowers
medium-sized for the genus, with wide-spreading membranaceous segments, yellow
with dark nerves. Dorsal sepal lanceolate or oblong-lanceolate, acuminate, dor-
sally carinate, 3-nerved, navicular, 2.1-2.3 cm. long, 6.8 mm. wide. Lateral sepals
similar but narrower, lightly oblique. Petals much larger, suborbicular-ovate or
rhombic-ovate, acute or short-acuminate, 9- to 11-nerved, about 2.6-2.7 cm. long,
SCHWEINFURTH: ORCHIDS OF PERU 951
1.8-2 cm. wide. Lip ovate-suborbicular, narrowed to an acute or apiculate apex,
about 2.6 cm. long and equally broad, 15- to 17-nerved. Column very short and
stout, protruded onto the lip, surrounded (except in front) by a dense cluster of
dark bristles.
Cajamarca: Prov. of Jaen de Bracamoros, between the districts
of Chores and Tomependa, near the confluence of the Chinchipes
and Amazon rivers, in very hot places, Humboldt & Bonpland s.n.
(type). Also Colombia and Ecuador.
Telipogon obovatus Lindl. Bot. Reg. 33: sub t. 27. 1847;
Kranzl. Ann. Naturh. Hofmus. Wien 33: 25. 1919.
Plant medium-sized. Stem abbreviated. Leaves oblong or oblong-oblanceo-
late, acute, up to 7.6 cm. long and 1.8 cm. wide, coriaceous. Inflorescence stout,
loosely racemose above, about 18 cm. or less long; peduncle naked, compressed
and enlarged above, about 13 cm. high; raceme arcuate, about 6-flowered. Floral
bracts conspicuous, spreading, ovate, concave, acute or acuminate. Flowers large,
with membranaceous wide-spreading segments, bright yellow. Sepals similar, lan-
ceolate or ovate-lanceolate, acuminate, dorsally more or less carinate, about 1.8 cm.
long and 7 mm. wide. Petals much larger, oblong-subrhombic ("oblong"), acute,
11-nerved, about 3 cm. long and 1.8 cm. wide. Lip very broadly obovate or sub-
orbicular, rounded and not apiculate in front, 13- to 15-nerved, about 2.5 cm. long
and nearly 3 cm. wide. Column apparently short and stout, adorned with very
many long, brown bristles.
Peru: Habitat unrecorded, Lobb s.n. (type). Also Ecuador and
perhaps Colombia, fide Kranzl. I.e.
In the Ames Herbarium is a photograph of the type in the Lind-
ley Herbarium, as well as one Ecuadorian specimen referred to this
species.
Telipogon Papilio Reichb. f. & Warsc. Bonpl. 2: 101. 1854;
Kranzl. Ann. Naturh. Hofmus. Wien 33. 26. 1919.
Plant small, epiphytic. Stem very short, concealed by imbricating, condupli-
cate leaf-sheaths and by the numerous long, fibrous, glabrous roots which have a
very stout velamen. Leaves several, congested, oblanceolate to oblanceolate-
oblong, acute with an apiculate tip, cuneate-narrowed to an articulated base,
3.5-6 cm. long, 1.2-1.7 cm. wide (up to 8 cm. long and 6-8 mm. wide, according
to Kranzlin). Inflorescence strict, shorter than or surpassing the leaves, about
4-17.7 cm. tall, subterete to narrowly bialate and angulate, racemose above; ra-
ceme 1- to 7-flowered, with a more or less fractiflex rachis. Floral bracts conspicu-
ous, spreading, triangular-ovate, conduplicate, dorsally carinate, acute. Pedicellate
ovary narrowly trialate, much longer than the bracts. Flowers medium-sized for
the genus, with wide-spreading membranaceous segments, orange-yellow or primu-
line with red nerves. Dorsal sepal lanceolate-ovate or narrowly triangular, acumi-
nate ("acute"), concave and navicular, dorsally strongly carinate, up to about
1.8 cm. long and 7.4 mm. wide when expanded, 3-nerved. Lateral sepals similar
952 FIELDI ANA: BOTANY, VOLUME 30
but shorter and narrower, oblique. Petals much larger, broadly rhombic-obovate,
oblique, abruptly acute, commonly 13- to 15-nerved, about 1.9 cm. long and very
slightly narrower (2.2 cm. long and equally wide, according to Kranzlin). Lip
transversely suborbicular-oval, minutely apiculate at the very broadly rounded
and plicate apex, about 25-nerved, about 1.85 cm. long, 2.5 cm. or more wide,
densely setose near the base. Column abbreviated, very stout, surrounded (except
in front) by a tuft of numerous, long bristles.
Peru: Habitat not recorded, Warscewicz s.n. (type). — Cuzco:
Prov. of Urubamba, "Huinaihuaina" Inca ruins, 2550 meters, epi-
phyte in forest, Vargas 4140a.
Telipogon radiatus Reichb. f. Linnaea 41: 70. 1876; Kranzl.
Ann. Naturh. Hofmus. Wien 33: 24. 1919.
Vegetative parts lacking in the type (and apparently the only collection) and
the description was drawn from a single flower. Pedicellate ovary sharply trialate.
Sepals triangular or triangular-ligulate, similar, dorsally carinate, acuminate, 3-
nerved, 1.6 cm. long, 5-6 mm. wide. Petals much larger, rhombic ("cuneate-
elliptic"), acute, 9- to 11-nerved, 2.2 cm. long and 2 cm. wide. Lip intermediate
between transversely oblong-oval and very broadly obovate, without any apicule
at the broadly rounded apex, 21-nerved, 2.5 cm. long and 3.7 cm. wide, the basal
portion densely pilose. Column with violet bristles.
Peru: Habitat not recorded, Davis s.n.
No authentic material of this species was available. The floral
measurements were derived from Kranzlin's description (I.e.).
Telipogon rhombipetalus C. Schweinf. Am. Orch. Soc. Bull.
15: 180, t., 1946. Figure 185.
Plant rather small, epiphytic. Stem decumbent, leafy, somewhat branched,
apparently entirely concealed by the leaf-sheaths. Leaves lanceolate-oblong to
oblong-elliptic, acute, short-apiculate, more or less narrowed to a sessile, compli-
cate and articulated base, about 2.9-4 cm. long, up to 1 cm. wide. Inflorescence
racemose above; peduncle strict, narrowly bialate, subequaling or somewhat ex-
ceeding the leaves, about 4 cm. or less high; racemes very short, 3-flowered, with
the fractiflex rachis about 1.2-1.5 cm. long. Floral bracts spreading, strongly
conduplicate, dorsally carinate, acute, apparently ovate when expanded. Flowers
medium-sized for the genus, with wide-spreading membranaceous segments, appar-
ently yellowish with purple lines near the base. Pedicellate ovary trialate. Dorsal
sepal elliptic-lanceolate, concave, subacute, with a prominent dorsal keel (broader
at the base) produced into an apicule, 3-nerved with 2 short additional nerves near
the base, about 1.9 cm. long and 7.6 mm. wide. Lateral sepals similar but slightly
narrower, oblong-lanceolate, somewhat oblique. Petals a little longer and about
three times broader than the sepals, suborbicular-rhombic, more or less acute,
15- to 16-nerved. Lip very similar to the petals but less rhombic, ovate-suborbicu-
lar, minutely apiculate, 19-nerved with the lateral nerve branching on each side
without, densely setulosejat the base, about 1.95 cm. long and 2.1 cm. wide.
Column very short and stout, with very numerous long, purple bristles.
TEJ-JPOGON
ijo
FIG 185. Telipogon rhombipetalus C. Schweinf. 1, plant; X 1. 2, dorsal
sepal; X 2. 3, lateral sepal; X 2. 4, petal; X 2. 5, lip; X 2.
953
954 FIELDIANA: BOTANY, VOLUME 30
Cajamarca: Celendin, 2625 meters, in moss on a thick branch of
a tree over a shady brook in a canyon, "F.W.D." s.n. (type in Herb.
Chicago Nat. Hist. Mus.).
Telipogon semipictus Reichb. f. ex Kranzl. Ann. Naturh. Hof-
mus. Wien33:25. 1919.
Plant small, apparently epiphytic, with very long roots. Stem abbreviated,
somewhat short-branching, entirely concealed by imbricating leaf-sheaths. Leaves
several, aggregated on each branch, oblanceolate or elliptic-oblong, acute, short-
apiculate, narrowed below to a sessile or petiole-like, articulated base, very variable
in size, up to 8 cm. long (rarely) and 1.2 cm. wide. Inflorescence surpassing the
leaves, racemose and arcuate above; peduncle slender, more or less dilated above,
naked, about 6.7 cm. long; raceme short, loosely 3- to 5-flowered; rachis fractiflex,
about 5 cm. or less long. Floral bracts conspicuous, spreading, conduplicate, dor-
sally carinate, ovate when expanded. Flowers small for the genus, with wide-
spreading membranaceous segments, "greenish yellow with broken cross marks of
red-brown, center purple." Pedicellate ovary trialate. Dorsal sepal oblong-ovate
or lanceolate-ovate, acute, strongly concave, dorsally carinate with the keel pro-
duced in a mucro, 3-nerved, about 1.3-1.65 cm. long and 4-7 mm. wide. Lateral
sepals similar but shorter and narrower, lanceolate, acuminate, lightly oblique.
Petals much larger than the sepals, suborbicular-rhombic or broadly rhombic-
ovate, acute, minutely apiculate, about 1.5-1.7 cm. long and equally wide or
slightly narrower, 8- to 11-nerved. Lip transversely oblong-oval, broadly rounded
in front with a minute apicule, 15- to 17-nerved, about 1-1.5 cm. long and 2 cm.
wide, densely setulose at the base. Column very short and stout, produced onto
the lip, densely long-setose (especially on the sides and rear).
Huanuco: Huacachi, near Muna, in montana, about 2000 meters,
Macbride J^.166. Also Colombia (type) and Ecuador.
Telipogon tessellatus Lindl. Benth. PI. Hartw. 150. 1844;
Kranzl. Ann. Naturh. Hofmus. Wien 33: 21. 1919.
Plant small, about 6.5-12 cm. high (exclusive of the flowers). Stem abbrevi-
ated, often with a few short, suberect branches. Leaves several, congested, variable
in size, oblanceolate ("oblong"), acute, apiculate, narrowed below to a sessile,
articulated base, rarely up to 10 cm. long and 1.8 cm. wide. Inflorescence suberect,
racemose above, becoming a little longer than the leaves; peduncle naked, grad-
ually enlarged and bialate above, about 3.8-8.1 cm. long; raceme abbreviated,
2-flowered; rachis about 6.5 mm. or more long. Floral bracts conspicuously spread-
ing, conduplicate, triangular-ovate when expanded, dorsally carinate, acuminate.
Flowers large for the plant, with wide-spreading segments. Sepals narrowly lan-
ceolate or oblong-lanceolate, acuminate, concave, dorsally carinate, about 1.3-
1.6 cm. long and 5 mm. broad. Petals much larger than the sepals, suborbicular-
rhombic, acute, 7- to 9-nerved, with the nerves conspicuously cross-reticulated,
about 1.5-2 cm. long and 1.35-1.8 cm. wide. Lip transversely oblong-oval, about
15-nerved with dark nerves and conspicuous cross-reticulations, broadly rounded
SCHWEINFURTH: ORCHIDS OF PERU 955
and apparently lightly retuse in front, about 1.4-1.8 cm. long and 2-2.5 cm. wide.
Column abbreviated, adorned with a few bristles.
Cajamarca, fide Schlechter, Fedde Repert. Beih. 9: 179. 1921.
Also Colombia and Ecuador (type).
The type, as recorded by a photograph in the Ames Herbarium,
appears to be rather smaller than one plant of an Ecuadorian col-
lection (Espinosa 761).
Telipogon urceolatus C. Schweinf. Am. Orch. Soc. Bull. 16:
292, t., 1947. Figure 186.
Plant epiphytic, small but with a large flower, about 15 cm. or less high. Roots
numerous, recurved, with a stout, shining velamen. Stems decumbent, with an
ascending, branching apex. Branches (including the inflorescence) up to 12 cm.
long; lower portion relatively short, entirely concealed by 2-4 imbricating, con-
duplicate leaf -sheaths; upper portion consisting of a relatively long, naked peduncle
terminated by a short raceme. Leaves elliptic-oblong (rarely) to oblanceolate,
acute, short-apiculate, more or less long-narrowed below to a sessile, often petiole-
like, articulated base, 5-10 cm. long, about 1-1.5 cm. wide (sometimes very small).
Peduncle about 7 cm. or less long. Raceme short, loosely 2- to 5-flowered, the
fractiflex rachis about 3 cm. or less long. Floral bracts spreading, conduplicate,
dorsally carinate, broadly ovate when expanded, acute or acuminate. Pedicellate
ovary much surpassing the bracts, trialate, about 3.5 cm. long. Flower large for
the genus, with wide-spreading segments, pale greenish yellow lined with dark red.
Dorsal sepal elliptic-lanceolate, navicular, sharply complicate-acute, dorsally cari-
nate, 5-nerved, about 2.6 cm. long and 9 mm. wide when expanded. Lateral sepals
similar, obliquely oblong-lanceolate, somewhat shorter and narrower than the dor-
sal sepal. Petals much larger, oblong-urceolate in outline, abruptly dilated above
with a very broadly triangular-acute apex having the outer angles rounded, about
3.2 cm. long and 2.7 cm. wide above, 13-nerved with 2 short additional nerves at
the base on each side. Lip transversely oval-suborbicular, very broadly rounded
at the apex, subcordate at the base, about 3.1 cm. long and 4.2 cm. wide, about
49-nerved. Column very short and stout as in the genus, with a surrounding tuft
of deciduous, purple hairs.
Huanuco: Mufia, about 2150 meters, in dry woods, Macbride 4193.
Telipogon Vargasii C. Schweinf. Am. Orch. Soc. Bull. 15: 290,
t., 1946. Figure 187.
Plant epiphytic, medium-sized. Stem decumbent, the lower portion leafy and
apparently wholly concealed by the sheathing bases of the leaves. Leaves numer-
ous, oval to oblong-elliptic, acute, short-apiculate, sheathing the stem toward the
base, up to about 5.5 cm. long and 2 cm. wide. Peduncle slender, naked, strict,
about 9 cm. long. Raceme short, 2- to 3-flowered, about 3 cm. long. Floral bracts
small, triangular-lanceolate, amplexicaul, concave. Pedicellate ovary much sur-
passing the bracts, about 3.5 cm. long. Flower large for the genus, yellow, with
wide-spreading segments. Dorsal sepal lanceolate, acuminate and complicate
above, concave, 5-nerved near the base, dorsally subcarinate, about 2 cm. long
T^LIPOGON
FIG. 186. Telipogon urceolatus C. Schweinf. 1, plant; X 4A- 2, dorsal sepal;
XI. 3, petal; XI. 4, lateral sepal; XI. 5, lip; X 1.
956
FIG 187. Telipogon Vargasii C. Schweinf. 1, plant; X %• 2, dorsal sepal,
expanded; X 1. 3, petal, expanded; X 1. 4, lateral sepal, expanded; >
5, lip, expanded; X 1.
957
958 FIELDIANA: BOTANY, VOLUME 30
and 7.6 mm. wide. Lateral sepals similar but slightly narrower, oblique, dorsally
carinate. Petals much larger, very broadly ovate, short-acuminate, 11-nerved
with cross-reticulations throughout, about 3.7 cm. long and 2.7 cm. wide. Lip
transversely broad-ovate, acute and minutely apiculate, subtruncate at the base,
17-nerved with cross-reticulations throughout, about 3.3 cm. long and 4.2 cm. wide.
Column long for the genus, stout, densely setose throughout, about 8.9 mm. high
at the back.
Cuzco: Prov. of Convention, Hda. Potrero, Sapan-Sachayocc, in
forest, Vargas 2551.
Telipogon Warscewiczii Reichb. f.
This concept, which was described only from two flowers, was
apparently found in Ecuador (near Loxa), although it was cited by
Kranzlin (Ann. Naturh. Hofmus. Wien 33: 26. 1919) from Peru.
TRICHOCEROS HBK.
Small to medium-sized epiphytes or terrestrials, recorded only
from Colombia, Ecuador, Peru and Bolivia.
Plants with a more or less elongate rhizome, bearing very short pseudobulbous
stems. Leaves 2 or more, distichously imbricating and sheathing the pseudobulb
(which commonly bears 1 smaller blade), small, fleshy or coriaceous, unjointed.
Inflorescences lateral, 1 or 2 at the base of each pseudobulb, simple, racemose
above, commonly much exceeding the leaves; racemes loosely few- to several-
flowered. Flowers more or less small, with spreading segments. Sepals subequal,
free, spreading. Petals similar to the sepals (often narrower or broader). Lip
usually 3-lobed (very rarely simple); lateral lobes spreading or erect, commonly
narrow and retrorse; mid-lobe much larger, usually more or less ovate. Column
very short and stout, wingless, footless, with a fimbriate clinandrium, and often
long-setose in the rear. Anther attached to the erect rostellum, 2-celled. Pol-
linia 4, affixed to the slightly dilated apex of the linear stipe.
Al. Lip simple, without lateral lobes, broadly obovate; inflorescences very short,
little exceeding the leaves T. Tupaipi
A2. Lip deeply 3-lobed at the base, with oblong-ovate or obovate to linear lateral
lobes; inflorescences commonly many times exceeding the leaves I
la. Lateral lobes of the lip linear or antenna-like.
T, parviflorus (T. armillatus, T. muscifer)
Ib. Lateral lobes of the lip obliquely semiovate to round-obovate.
T. muralis var. platyceros (T. platyceros)
Trichoceros muralis Lindl. var. platyceros (Reichb. f.) C.
Schweinf. Bot. Mus. Leafl. Harvard Univ. 17: 58. 1955. Trichoceros
platyceros Reichb. f. Xen. Orch. 1: 22, t. 9, fig. III. 1854.
Plant small to medium-sized, epiphytic. Roots stout, glabrous. Rhizome
creeping, more or less concealed by close, tubular sheaths. Pseudobulbs small,
SCHWEINFURTH: ORCHIDS OF PERU 959
broadly ovoid, mostly concealed by sheaths or the bases of the imbricating leaves.
Leaves 2 or 3, clustered at the base of the stems, suborbicular to elliptic-oblong,
sessile at the clasping base, apiculate, up to about 4 cm. long and 1.4 cm. wide,
coriaceous. Inflorescences up to 15.5 cm. high; peduncle slender, provided with
2 or 3 short, infundibuliform sheaths; raceme short, loosely up to 7-flowered.
Flowers rather small, with spreading segments. Dorsal sepal cucullate, ovate-
elliptic, acute, apiculate, 8-16 mm. long, up to 9 mm. wide. Lateral sepals similar,
sometimes slightly longer. Petals similar to the sepals but often slightly broader,
ciliolate throughout. Lip about equaling the other segments, 3-lobed at the base,
up to 15 mm. long; lateral lobes relatively small, obliquely semiovate to round-
obovate, rounded-obtuse; mid-lobe relatively large, obovate or oblong-obovate,
rounded at the apex, sometimes with a minute apicule; disc more or less pilose
throughout. Column minute, densely pilose.
Ancash : Prov. of Bolognesi, Chupa, mountain near Aquia, 3200-
3250 meters, on stony soil, "rojo oscuras," called "michimichi,"
Ferreyra 7546. — Cajamarca, fide Schlechter, Fedde Repert. Beih. 9:
180. 1921. Also Ecuador (type).
Trichoceros parviflorus HBK. Nov. Gen. et Sp. 1: 337, t. 76.
1816; Lindl. Gen. & Sp. Orch. 174. 1833; Reichb. f. Xen. Orch. 1: 21,
t. 9, figs. I-II, 1-6. 1854; C. Schweinf. Bot. Mus. Leafl. Harvard
Univ. 17: 60. 1955. Trichoceros armillatus Reichb. f. Bonpl. 4: 212.
1856; Xen. Orch. 1: 231, t. 97, fig. V. 1858. Trichoceros muscifer (as
muscifera) Kranzl. Engler, Bot. Jahrb. 37: 387. 1906.
Plants extremely variable, commonly terrestrial in humus or on rocks (rarely
epiphytic). Stem more or less elongate, ascending, sometimes branched, mostly
concealed by close, tubular, not imbricating, evanescent sheaths, bearing (at
more or less remote intervals) clusters of leaves surrounding a pseudobulb beneath
which are 1 to several elongate, stout, fibrous roots. Leaves 2 to several, distichous,
imbricating, unjointed, ovate to elliptic-oblong or oblong, sharply acute or acum-
inate, narrowed below to a sheathing base, spreading, commonly thickly coriaceous,
often condu plicate in the dried specimen, up to 7 (rarely 9) cm. long and 2 cm.
wide, supported below by 2 or 3 concave, ovate to lanceolate sheaths. Pseudo-
bulbs small, more or less concealed by the leaves, broadly ellipsoid to oblong-
cylindric, about 1-1.5 cm. high, commonly bearing a single rudimentary or small
leaf at the summit, rarely 1-jointed near the summit and bearing (at the lower
articulation) a well-developed blade. Inflorescences lateral from beneath the
pseudobulb, 1 or 2 to each pseudobulb, racemose above, much surpassing the
leaves, more or less erect to arcuate; peduncle slender, about 5 to more than 25 cm.
long, with 2-4 remote, small, close, tubular sheaths; raceme few- (3-) to sev-
eral- (18-) flowered, more or less loose below, the rachis up to 6 cm. long. Floral
bracts small, ovate to ovate-lanceolate, acute or acuminate, much surpassed by
the pedicellate ovary in the course of development. Flowers small, with spreading
segments, the color varying from greenish or whitish green to pale yellow brown-
lined, with the central portion and often the lip with its appendages dark red
(the latter often barred). Dorsal sepal lanceolate-ovate or elliptic-ovate, concave,
960 FIELDIANA: BOTANY, VOLUME 30
acute or acuminate, 3- to 5-nerved, about 10-13 mm. long and 5-7 mm. wide.
Lateral sepals similar but commonly slightly wider, dorsally more or less carinate.
Petals elliptic, acute, 5- to 7-nerved, a little shorter than the sepals and equally
wide (or somewhat narrower), with the margins minutely ciliolate. Lip deeply
3-lobed at the base, about 9-13.1 mm. long; lateral lobes linear or oblong-linear,
obtuse, ciliate (especially above), 1-nerved, strongly retrorse in course of develop-
ment; mid-lobe much larger, variable, elliptic-ovate to suborbicular-oval, acute
to rounded and apiculate at the apex, usually 7-nerved, often with a fleshy, longi-
tudinal thickening below in the middle, more or less pubescent or setose toward
the base, about 6-9 mm. wide. Column very short and stout, protruding onto
the lip, with a cluster of long bristles at the rear.
Peru: Habitat not recorded, Pavdn s.n. (type of T. armillatus).—
Cajamarca: Near Hualgayoc, beyond Hacienda La Tatoma, 2600
meters, on cliffs, Weberbauer 4061. — Cuzco: Prov. of Calca, Hda.
"Urco," 2900 meters, on rocks, Vargas 3051. Prov. of Calca, Vilca-
bamba (along Paucartambo River), 2600 meters, in humus of forest
slopes, Vargas 4009. — Huanuco: Cani, 7 miles northeast of Mito,
about 2600 meters, "humus floor of thicket," Macbride 3480. Yanano,
about 1800 meters, "floor of dense wood," Macbride 3833. — Junin:
Prov. of Tarma, Agua Dulce, 2300 meters, on tree in low highland
wood, Woytkowski 35460. Prov. of Tarma, above Huacapistana,
Weberbauer 2048. Prov. of Tarma, between Palca and Carpapata,
2900 meters, "over boulders that have accumulated some humus,"
Stork 10955. Carpapata, above Huacapistana, about 2400 meters,
on open hillside, Kittip & Smith 24345. Huacapistana, 1800-2400
meters, epiphyte in thickets and open woods, Killip & Smith 24220.
Same locality and altitude as the last, terrestrial, Killip & Smith
24124- Yanahuanca, about 3000 meters, on canyon ledge, "Miche-
huagte" or "Flor de Gato," Macbride & Featherstone 1239. — Puno:
Sandia, 2100-2300 meters, on cliffs, Weberbauer 542 (type of T. mus-
cifer}. Also Colombia (type of T. parviflorus), Ecuador and Bolivia.
Trichoceros Tupaipi Reichb. f. Linnaea 41: 33. 1876.
Plant small, apparently about 10 cm. or less high. Rhizome creeping, short,
with numerous stout, elongate roots. Pseudobulbs apparently subapproximate,
small, suborbicular-conic, unifoliate, less than 1 cm. high, surrounded below by
1 or 2 pairs of leaves. Leaves imbricating at the sheathing base, recurved and
fleshy-coriaceous in the dried specimen, apparently lanceolate to elliptic-oblong
("ligulate-triangular"), commonly more or less conduplicate in the dried speci-
men, acuminate, apparently about 3 cm. or less long. Inflorescences short,
about 7.6 cm. or less high, racemose above; raceme loose, up to 3-flowered.
Floral bracts small, ovate, apiculate, amplexicaul. Sepals similar, oblong-ovate
("oblong- triangular"), acute, 5-nerved, the dorsal sepal apparently a little shorter
and broader than the laterals. Petals ovate-elliptic, acute, 7-nerved, apparently
slightly longer than the lateral sepals and markedly broader than the dorsal
SCHWEINFURTH: ORCHIDS OF PERU 961
sepal. Lip suborbicular-obovate ("cuneate-oblong"), abruptly apiculate at the
broad apex, 9-nerved. Column short, stout, with the clinandrium setose-fimbriate.
Peru: Habitat not recorded, F arris s.n. Huansaquasi, Dombey
s.n. (without flower) .
This description was largely compiled by an examination of a
drawing with floral analysis from the Reichenbach Herbarium, as no
other record of this species was available.
STELLILABIUM Schltr.
A very small, perhaps monotypic genus, confined, in South Amer-
ica, to Peru.
Plant small, epiphytic, with an abbreviated rhizome and a very short stem.
Leaves several, approximate, articulated, narrow, their sheaths entirely con-
cealing the stem. Inflorescence subequaling or somewhat surpassing the leaves,
racemose at the apex; peduncle very slender, relatively elongate; raceme con-
tracted, densely several-flowered. Flowers very small, not wide open. Sepals
ovate or oval-ovate, acute. Petals smaller, oblong ("ligulate"), rounded at the
apex, minutely serrulate. Lip 3-lobed near the base, triangular in outline, entirely
velvety-papillose, with 3 clusters of stellate hairs; lateral lobes retrorse-spreading,
relatively small, ovate-oblong; mid-lobe relatively large, oblong-ovate ("oblong
from a broader base")» apiculate at the rounded apex. Column very short and
stout, dilated upward, with a cluster of elongate papillae or bristles on either
side at the apex. Anther very slender, cordiform. Pollinia 4, compressed, oblong-
cylindric (the 2 outer smaller), inserted at the dilated apex of the linear stipe.
Stellilabium astroglossum (Reichb. f.) Schltr. Die Orchideen
530. 1914. Telipogon astroglossus Reichb. f. Xen. Orch. 1: 16, t. 7,
fig. II, 12-19. 1854; Kranzl. Ann. Naturh. Hofmus. Wien 33: 15.
1919. Trichoceros astroglossus Lindl. ex Benth. & Hook. f. Gen. PL
3: 588. 1883.
Plant slender, delicate, about 5-10 cm. high. Stem abbreviated, entirely
concealed by the short, approximate leaf-sheaths. Leaves 3 or 4, linear-lanceolate,
acute, slightly narrowed at the complicate base, up to 5 cm. long and 6 mm. wide.
Inflorescence terminal, slightly exceeding the leaves, racemose above, about 6 cm.
tall; peduncle very slender, with a single small sheath in the middle; raceme
abbreviated, densely few-flowered. Floral bracts ovate-triangular, acute, about
a fourth as long as the filiform pedicellate ovary. Flowers very small, little
spreading, hyaline. Dorsal sepal ovate or oblong-ovate, acute, about 3-4 mm.
long and 2.5 mm. wide, 1-nerved. Lateral sepals similar, broadly ovate, oblique,
apparently slightly shorter and broader than the dorsal sepal. Petals broadly
oblong ("ligulate"). obtuse or rounded ("retuse") at the apex, minutely serrulate
or ciliolate, 1-nerved, slightly shorter than the sepals and much narrower. Lip
triangular in outline when expanded, with a very short and broadly cuneate base,
lightly 3-lobed near the base, about 2.5 mm. long and equally broad, according
to Kranzlin; lateral lobes relatively small, erect in natural position, retrorse-
962 FIELDIANA: BOTANY, VOLUME 30
spreading when expanded, ovate-oblong ("ligulate"), rounded at the apex; mid-
lobe much larger, oblong-ovate, rounded and apiculate at the tip; disc entirely
pilose on the upper surface, with 3 clusters of stellate radiating bristles (the
anterior central one largest). Column very short and stout, protruded onto the
lip, dilated upward, with a cluster of long bristles or setae on each side at the apex.
Amazonas: Near Chachapoyas, Mathews s.n.
No material of this concept was available.
DICHAEA Lindl.
Inconspicuous epiphytes extending from Mexico and the West
Indies through Central America, and in South America as far as
Brazil, Bolivia and Peru.
Plants small to medium-sized, with more or less elongate, leafy stems. Stems
often pendent, concealed by the leaf-sheaths or the sheathing bases of the leaves.
Leaves numerous, distichous, spreading or reflexed, articulated or not articulated,
very short to relatively long. Peduncles axillary, 1-flowered, solitary, commonly
much shorter than the leaves. Flowers very small to (rarely) medium-sized.
Perianth segments more or less spreading. Sepals subequal, free, the lateral
ones oblique and forming an inconspicuous mentum with the short column-foot.
Petals similar to the sepals or often smaller. Lip rigidly joined to the column-
foot, sessile to long-clawed, concave, varying from ovate to broadly triangular or
anchor-shaped and often produced on each side into a more or less prominent lobe,
angle or tooth. Column erect, short, commonly exalate, produced at the base
into a short or minute foot, often with a prominent ligule in front. Anther terminal,
opercular, incumbent. Pollinia 4, ovoid, attached in pairs to a small stipe.
Al. Lamina of the leaves not articulated to a sheath, i.e., not deciduous.
D. muricata var. maculata (D. maculata')
A2. Lamina of the leaves articulated to a sheath, i.e., deciduous I
la. Ovary pilose, capsule echinate 1
Ib. Ovary and capsule smooth 3
la. Plant robust; leaves oblong to elliptic-oblong, commonly1 10 mm. or more
wide; sepals 11 mm. or more long D. Morrisii
Ib. Plant relatively slender; leaves linear-lanceolate to linear, 6 mm. or less wide;
sepals 6 mm. or less long 2
2a. Stems very short, 3-7 cm. long; leaves narrowly linear, 2.5 mm. or less wide.
D. humilis
2b. Stems relatively elongate, 10-50 cm. long; leaves lanceolate-linear, 4 mm. or
more wide D. laxa
3a. Leaf-blades very narrowly linear or acicular, many times longer than broad;
lip more or less emarginate and apiculate 4
3b. Leaf-blades linear-oblong or broader, 4.4 cm. or less long; lip acute to rounded
at the apex, not distinctly emarginate 5
4a. Plant relatively large, stems 20 cm. or more long; leaves 5-12.5 cm. long.
D. calyculata
1 Rarely a leaf may be found in D. Morrisii which is only 5 mm. wide in the
dried specimen, but such a blade is unusual in any collection.
SCHWEINFURTH: ORCHIDS OF PERU 963
4b. Plant very small, stems about 7 cm. or less long; leaves about 1.1 cm. or less
long D. tennis
5a. Lower half of the lip with a keel or a longitudinal callus 6
5b. Lower half of the lip without any distinct keel or longitudinal callus 7
6a. Lip with the lower half broadly cuneate, narrowed toward the base.
D. campanulata
6b. Lip with the lower half broadly auriculate-rounded on each side.
D. graminoides
7a. Anterior portion of the lip broadly rounded, with a truncate, subretuse apex.
D. ancoraelabia
7b. Anterior portion of the lip with an acute or acuminate apex . .D. panamensis
Dichaea ancoraelabia C. Schweinf. Am. Orch. Soc. Bull. 16:
614, t., 1947. Figure 188.
Plant medium-sized for the genus, recalling D. graminoides (Sw.) Lindl.
Stems simple or branched, erect to flexuous or arcuate, up to 24 cm. or more
long, entirely concealed by imbricating leaf-sheaths. Leaves numerous, distichous,
rather loose, articulated, elliptic-oblong, acute, minutely apiculate, up to 2.4 cm.
long and 6 mm. wide, smooth on the margin. Inflorescences axillary, 1-flowered,
the free portion of the peduncle less than one half as long as the leaf-blades. Flower
rather small, campanulate, white with pale purple markings. Ovary smooth.
Dorsal sepal ovate, acute, concave, 3-nerved, about 5.1 mm. long and 4 mm. wide.
Lateral sepals obliquely ovate, acute or short-acuminate, concave, markedly
longer than the dorsal sepal and equally wide. Petals ovate-elliptic, acute, slightly
longer than the dorsal sepal but narrower. Lip anchor-shaped, concave, from
a long-cuneate base abruptly dilated above, broadly rounded in front with a
subtruncate, minutely apiculate apex, the anterior part terminating on each
side in an obliquely triangular, retrorse lobule, about 7 mm. long and broad
across the lobules. Column short, stout, about 2.3 mm. high at the back.
Loreto : Right bank of Rio Santiago, above Pongo de Manseriche,
200 meters, common, Mexia 6221.
Dichaea calyculata Poepp. & Endl. Nov. Gen. ac Sp. 2: 4, 1. 106.
1838; Cogn. Martius Fl. Bras. 3, pt. 6: 498. 1906. Epithecia calycu-
lata Schltr. Orchis 9: 26. 1915.
Plant medium-sized for the genus, epiphytic. Stems solitary or two, erect or
ascending, simple or very rarely bifurcate, spreading, arcuate or deflexed above,
entirely concealed by imbricating leaf-sheaths, 20-45 cm. long. Leaves numerous,
distichous, articulated, very narrowly linear, more or less recurved, acute or
acuminate and apiculate, 5-12.5 cm. long, about 2-3 mm. wide, loose to rather
dense. Inflorescences axillary, 1-flowered, much shorter than the leaves; peduncle
capillary, erect or spreading, below with several acuminate, scarious sheaths, and
above with 1 bract which loosely envelops the smooth ovary. Flowers small,
sessile, white with purple nerves or yellowish, the segments strongly concave,
erect and subconnivent. Dorsal sepal obovate-oblong, rounded and minutely
apiculate at the apex, about 5-7.8 mm. long and 2.75-4 mm. wide. Lateral
sepals very similar to the dorsal sepal but a little broader or narrower, suboblique.
DICHABA
an cor a e£a fria
C/. Oc&ureinfi.
FIG. 188. Dichaea ancoraelabia C. Schweinf. 1, plant; X %• 2, flower, nat-
ural position; X 4. 3, dorsal sepal; X 4. 4, petal; X 4. 5, lateral sepal; X 4.
6, lip, spread out; X 4. 7, column, oblique view; X 8.
964
SCHWEINFURTH: ORCHIDS OF PERU 965
Petals narrowly obovate-oblong, subrounded and minutely apiculate at the apex,
a little shorter and narrower than the lateral sepals, minutely ciliolate. Lip
fleshy, sessile, broadly obovate-deltoid, concave, abruptly dilated above with a
retrorse, uncinate angle on each side, broadly rounded-subtruncate in front with
an incurved, emarginate and minutely apiculate apex, about 4.5-6 mm. long
and 4-8.2 mm. wide above when expanded. Column rather long, stout, sub-
clavate, 3-4.2 mm. high, with an elongate linear-subulate rostellum.
Huanuco: Near Cuchero (Cochero) and Pampayaco (Pampayacu),
epiphyte, Poeppig 1092 (type). — San Martin: In jungle east of Tingo
Maria, 625-1100 meters, Attard 20739, 20742. On ridge east of
Tingo Maria, Allard 20940.
Dichaea campanulata C. Schweinf. Am. Orch. Soc. Bull. 16:
346, t., 1947. Figure 189.
Plant epiphytic, small to medium-sized for the genus, lax. Stems spreading,
suberect, flexuous or arcuate, simple or branched, rarely up to 30 cm. or more long,
entirely concealed by loose, imbricating leaf-sheaths. Leaves numerous, small,
distichous, articulated, rather loose, oblong to elliptic-oblong, commonly acute or
apiculate, up to 2 cm. long (rarely) and 5 mm. wide. Inflorescences short, in the
axils of upper leaves, much shorter than the leaves; peduncle filiform, 1-flowered,
with 3 small, loose, imbricating, infundibulif orm sheaths at the base. Ovary sessile,
smooth, surrounded by a loose, infundibuliform sheath. Flower very small, with
spreading, membranaceous segments, snow-white and carmine, pinkish or pinkish
white. Dorsal sepal oblong-ovate or elliptic-ovate, acute, apiculate, concave, about
4.2-4.5 mm. long and 2-2.5 mm. wide. Lateral sepals similar to the dorsal sepal
and sometimes slightly broader, suboblique. Petals oblong-elliptic, acute, com-
monly slightly smaller than the sepals, often somewhat oblique. Lip deeply con-
cave, a little shorter than the sepals but much wider, broadly rhombic-obovate,
rounded in front with a more or less prominent apicule, cuneate below to a sessile
base, with the outer angle on each side subacute (or lobulate) and spreading, about
3.5-4 mm. long and 5.1-6 mm. wide when expanded; disc with the mid-nerve cari-
nate-thickened about to the middle. Column very short and stout, with a large
stigmatic cavity below and a cucullate clinandrium above.
Ayacucho: Rio Apurimac Valley, near Kimpitiriki, 400 meters,
at edge of dense forest along beach, Killip & Smith 22978. — Junin:
Chanchamayo Valley, 1500 meters, Schunke 524- East of Quimiri
Bridge, near La Merced, 800-1300 meters, in dense forest, Killip &
Smith 23955 (fruit). Tarma, Vitoc, "Eyebrow of the jungle," 1800
meters, Woytkowski 3. — Loreto: Vicinity of Iquitos, 100 meters, on
living tree by river, King 10014 (type).
Dichaea graminoides (Sw.) Lindl. Gen. & Sp. Orch. 209. 1833;
Fawc. & Rendle, Fl. Jam. 1: 138, t. 30, figs. 8-12. 1910. (Not D.
graminoides Cogn. Martius Fl. Bras. 3, pt. 6: 492, 1. 102, fig. 2. 1906.)
Epidendrum graminoides Sw. Prod. Veg. Ind. Occ. 125. 1788. Cym-
966 FIELDIANA: BOTANY, VOLUME 30
bidium graminoides Sw. Nov. Act. Upsal. 6: 71. 1799; Fl. Ind. Occ.
1459. ?1806. Epithecia graminoides Schltr. Orchis 8: 101. 1914.
Plant small to medium-sized, erect to lax or arcuate, epiphytic or on rocks.
Stems up to 30 cm. or more long, simple or somewhat branched below, entirely
concealed by leaf-sheaths, the lower portion destitute of leaf-blades. Leaves dis-
tichous, linear-oblong to elliptic-oblong, acute and prominently apiculate, articu-
lated, wide-spreading, minutely ciliolate above, up to 4.4 cm. long, about 4-6 mm.
wide, rather loose. Inflorescences axillary, 1-flowered, about half as long as the
leaves or less; peduncle capillary, straight or arcuate. Ovary smooth. Flower
medium-sized for the genus, whitish. Segments campanulate. Dorsal sepal ellip-
tic-lanceolate, acute or short-acuminate, 5.9-7.9 mm. long, 2.2-3 mm. wide, 3- or
indistinctly 5-nerved. Lateral sepals similar to the dorsal sepal but slightly longer
and either narrower or more or less wider, lightly oblique. Petals similar to the
sepals and either slightly smaller or larger. Lip shorter than the other segments
but broader, 4.5-5.8 mm. long, 3.2-5.8 mm. wide above, divided by a constriction
near the base into a broadly deltoid-ovate, acute to acuminate anterior part and
a shorter, narrower basal part which is auriculate-rounded on each side; disc with a
prominent central, longitudinal callus through the lower part. Column small,
stout, dilated upward.
Peru, probably, Poeppig 2024, fide Kranzlin, Engler, Pflanzenr.
IV. 50 (Heft 83), 47. 1923. Also the West Indies (type), Mexico,
Guatemala and Nicaragua.
Dichaea humilis Cogn. Martius Fl. Bras. 3, pt. 6: 502. 1906.
Epithecia humilis Schltr. Orchis 9: 27. 1915.
Plant small, epiphytic. Stems numerous, erect or ascending to arcuate, short,
3-7 cm. long, with erect, densely fastigiate branches. Leaves numerous, densely
distichous, articulated, narrowly Jinear, sharply acute and short-apiculate, minutely
ciliate above, more or less falcate, usually conduplicate, 1.5-2.5 cm. long, 1.5-
2.5 mm. wide. Inflorescences subterminal; peduncle very short, slender, erect,
3-5 mm. long. Ovary shortly and very densely muricate. Flowers small, erect or
nodding, purplish brown. Sepals about equally long, oblong, short-acuminate,
somewhat concave, 5.5-6 mm. long, 1.75-2 mm. wide, submembranaceous, the
lateral sepals lightly oblique. Petals narrowly lanceolate, acute, a little shorter
and narrower than the sepals. Lip shorter and broader than the other segments,
broadly rhomboid from a rather long and broadly clawed base, apex broadly tri-
angular-subrounded and minutely apiculate, the lateral angle on each side sharply
acute, about 4 mm. long and equally wide when expanded. Column short, stout,
dilated above, about 2 mm. high.
Loreto: Near Cerro de Ponasa, 1200 meters, on trees, Ule 6666.
No material of this species was available.
Dichaea laxa Poepp. & Endl. Nov. Gen. ac Sp. 2: 4, t. 105,
figs. a-d. 1838; Cogn. Martius Fl. Bras. 3, pt. 6: 500. 1906. Epithecia
laxa Schltr. Orchis 9: 26. 1915. Fernandezia laxa Ruiz & Pav. Syst.
Veg. Fl. Peruv. et Chil. 1: 240. 1798, fide Schltr. I.e.
camoanu^aia (3. OcAuse i n ft.
FIG. 189. Dichaea campanulata C. Schweinf. 1, plant; X %. 2, flower; X 4.
3, dorsal sepal; X 8. 4, petal; X 8. 5, lateral sepal; X 8. 6, lip; X 8.
967
968 FIELDIANA: BOTANY, VOLUME 30
Plant small to medium-sized, lax, epiphytic. Stems solitary or few, ascending
to pendent, flexuous or arcuate, simple or sometimes slightly branched, 10-50 cm.
long. Leaves numerous, distichous, articulated, linear-ligulate or lanceolate-linear,
acute and rather long-aristate, minutely ciliate above, 2-3.5 cm. long, 4-6 mm.
wide, rather loose or distant. Inflorescences axillary, filiform, 1-flowered, straight
or arcuate, 10-13 mm. long, a little longer than the leaf-sheaths. Ovary shortly
and densely echinate. Flowers small, pale violet, sessile with incurved, rather con-
nivent, submembranaceous segments. Dorsal sepal oblong-lanceolate, acuminate,
6 mm. long, 2 mm. wide, rather concave. Lateral sepals similar to the dorsal sepal
but shorter and slightly narrower, suboblique. Petals narrowly ligulate, acumi-
nate, as long as the lateral sepals but slightly narrower. Lip as long as the lateral
sepals and petals, about 5 mm. long, from a short, filiform claw abruptly dilated
into a broadly ovate, cordate-sagittate lamina (about 3 mm. wide) which is
obliquely mucronate in front, with the sides terminating in a subulate, reflexed-
curved lobule. Column short, stout, clavate, lightly arcuate, about 2 mm. high.
Huanuco: Near Pampayaco (Pampayacu) and Cuchero (Cochero),
Poeppig 1009.
No material of this species was accessible.
Dichaea Morrisii Fawc. & Rendle, Journ. Bot. 48: 107. 1910;
Fl. Jam. 1: 139, t. 30, figs. 31-36. 1910. Epithecia Morrisii Schltr.
Orchis 9: 26. 1915.
Plant very stout for the genus, epiphytic, medium-sized to large. Stems soli-
tary or several and caespitose, up to 45 cm. long, erect to arcuate or flexuous,
entirely concealed by imbricating leaf-sheaths from which the leaf-blades have
commonly fallen in the lower portion, the sheathed stem of the upper portion (in
the dried specimen) about 10-15 mm. wide. Leaves numerous, distichous, articu-
lated, wide-spreading, oblong to elliptic-oblong (rarely linear-oblong), subdistant,
about 2.7-7.5 cm. long and 5 (very rarely) -15 mm. wide, rounded and mucro-
nate at the sometimes oblique apex with minutely ciliate upper margins. Inflores-
cences short, axillary, 1-flowered, the peduncle about 8 mm. long. Ovary and
capsule densely echinate. Flower large for the genus, rather fleshy, pale or yellow-
green often striped with lavender and lip lavender, or greenish white with brown
dots and rays. Dorsal sepal oblong-lanceolate, sharply acute or acuminate, con-
cave, minutely ciliate, 11-17 mm. long, about 5 mm. wide. Lateral sepals obliquely
ovate or lanceolate-ovate, acuminate, as long as the dorsal sepal but broader.
Petals obliquely elliptic-lanceolate or oblong-lanceolate, acuminate, a little shorter
and narrower than the dorsal sepal. Lip concave with a reflexed apex, anchor-
shaped, about 9 mm. or more long, from a linear-oblong claw abruptly dilated
above, abruptly acute or apiculate in front, the lateral lobe on each side retrorse,
acute or obtuse and more or less elongate. Column short, stout, arcuate.
Cuzco: Prov. of Convention, Hda. Potrero, Sapan-Sachayocc,
2200 meters, in forest, Vargas 2533. Also Jamaica (type), Sto. Do-
mingo, Costa Rica, Panama and Colombia.
Dichaea muricata (Sw.) Lindl. var. maculata (Poepp. & Endl.)
C. Schweinf. Bot. Mus. Leafl. Harvard Univ. 16: 17. 1953. Dichaea
SCHWEINFURTH: ORCHIDS OF PERU 969
maculata Poepp. & Endl. Nov. Gen. ac Sp. 2: 3, 1. 105, figs. 1-6. 1838;
Cogn. Martius Fl. Bras. 3, pt. 6: 489. 1906; Kranzl. Engler, Pflan-
zenr. IV. 50 (Heft 83), 40. 1923.
Plant medium-sized to large, epiphytic, lax, variable throughout. Stems pend-
ent, simple or more rarely branched, straight, lightly arcuate or flexuous, 7-120 cm.
long, entirely concealed by the imbricating, sheathing bases of the leaves, ancip-
itous and more or less robust. Leaves very numerous, distichous, small, commonly
very dense, either horizontally spreading or somewhat deflexed, not articulated;
lamina oblong or elliptic-oblong, obtuse to rounded at the apex, apiculate, promi-
nently ciliolate above, about 9-22 mm. long and 4-8 mm. wide. Inflorescences
axillary, 1-flowered, suberect to arcuate-recurved, much shorter than the leaves;
peduncle subfiliform, about 1 cm. long, clothed at the base with 2 or 3 short imbri-
cated sheaths. Ovary and capsule densely echinate. Flowers rather small to
medium-sized, variable in color (pale blue with large violet spots, dark purple, or
cream-color spotted with lilac), campanulate, more or less fleshy. Dorsal sepal
oblong-lanceolate, acute, concave, about 9-10.2 mm. long and 2.5-4 mm. wide.
Lateral sepals obliquely lanceolate, acuminate, a little larger or smaller than the
dorsal sepal. Petals subelliptic, acute or shortly acuminate, markedly shorter than
the dorsal sepal and as wide or nearly as wide as the lateral sepals. Lip from a long
cuneate base or a short, oblong, fleshy claw (attached to and continuous with the
column) dilated into an obovate-suborbicular lamina which is rounded above with
a more or less prominent mucro, 7-11 mm. long and 8.8-10 mm. wide when ex-
panded, on each side terminating in a narrowly triangular, acuminate, spreading or
retrorse lobule. Column short, stout, with a prominent lanceolate or subquadrate-
triangular ligule below.
Peru: Habitat not indicated, Pavdn s.n. — Cuzco: Prov. of Con-
vention, "alturas de Pintobamba," 2700 meters, pendent epiphyte
in forest, Vargas 3244- — Huanuco: Near Pampayaco (Pampayacu)
and Cuchero (Cochero), in dark humid woods, Poeppig 11+79 (type).
— Junin: Pichis Trail, Yapas, 1350-1600 meters, in dense forest,
Killip & Smith 25490. Rio Paucartambo Valley, near Perene Bridge,
700 meters, in dense forest, Killip & Smith 25249. Tarma, Vitoc,
1400 meters, Woytkowski 3 A. — Loreto: Left bank of Rio Santiago,
above Pongo de Manseriche, 200 meters, Mexia 6237. Near Tara-
poto, Spruce 4601 . Puerto Melendez, below Pongo de Manseriche,
on banks of the Maranon River, 155 meters, Tessmann 4766. — San
Martin: San Roque, 1350-1500 meters, Williams 7202. Also Costa
Rica, fide Kranzlin (I.e.).
Dichaea panamensis Lindl. Gen. & Sp. Orch. 209. 1833; Kranzl.
Engler, Pflanzenr. IV. 50 (Heft 83), 51. 1923. Epithecia panamensis
Schltr. Orchis 9: 25. 1915. Dichaeopsis panamensis Schltr. Beih. Bot.
Centralbl. 36, Abt. 2: 519. 1918. Figure 190.
Plant small to medium-sized, epiphytic. Stems simple or rarely branched,
commonly several and caespitose, erect or less commonly arcuate or flexuous, en-
970 FIELDIANA: BOTANY, VOLUME 30
tirely concealed by imbricating leaf-sheaths, rarely up to 27 cm. long. Leaves
numerous, distichous, articulated, rather loose, linear or oblong-linear (rarely
elliptic-oblong), acute, apiculate, spreading, commonly glaucous, up to 4 cm. long
(rarely) and 5 mm. wide, commonly in zones of more or less different lengths.
Inflorescences axillary, 1-flowered, usually much shorter than the leaves; peduncles
filiform. Flowers small to medium-sized, campanulate, variable in color (whitish,
greenish or yellowish, often with red or purple spots). Ovary and capsule glabrous.
Dorsal sepal oblong or oblong-lanceolate, acute or acuminate, concave, about 4-
7.3 mm. long and 2-2.8 mm. wide. Lateral sepals triangular-lanceolate, very
oblique, acuminate, deeply concave below, somewhat longer and broader than the
dorsal sepal. Petals obliquely ovate or oblong-ovate, acute, shorter and much
wider than either sepal. Lip obovate-anchoriform, about 4.3-9 mm. long and
nearly as wide or narrower across the lateral lobules, from a broadly oblong claw
abruptly dilated near the middle or below into a broadly ovate, acute or acuminate
lamina with a sharp, spreading-retrorse, more or less prominent angle or lobule on
each side. Column short, stout, with a small recurved ligule below.
Junin : East of Quimiri Bridge, near La Merced, 800-1300 meters,
in dense forest, Killip & Smith 25412. Also Mexico and Central
America to Panama (type), Colombia and Venezuela.
Dichaea tenuis C. Schweinf. Am. Orch. Soc. Bull. 21: 510, t.,
1952. Figure 191.
Plant small and slender, epiphytic, up to about 7 cm. or more high. Stems
apparently caespitose, slender, commonly arcuate, up to 7 cm. in length, entirely
concealed by tubular, leaf-bearing sheaths which waste into fibres at the base.
Leaves numerous, distichous, spreading, very fleshy, acicular, semiterete or sub-
quadrate in section with a deep groove on the upper surface, articulated, acumi-
nate, more or less recurved above, about 9-11 mm. long. Inflorescences axillary
on the upper part of the stem; peduncle filiform, distinctly shorter than the sub-
tending leaf. Flower subglobose in natural position, with the subfleshy perianth-
segments seemingly connivent. Ovary smooth. Sepals and lip concave in natural
position. Dorsal sepal oblong-ovate, acute, about 3 mm. long and 1.6 mm. wide.
Lateral sepals obliquely elliptic-ovate, acute, somewhat longer and broader than
the dorsal sepal. Petals ovate-elliptic, acute, slightly shorter and broader than the
dorsal sepal. Lip anchor-shaped, from a cuneate-unguiculate base dilated into an
abruptly crescentiform anterior portion, very broadly rounded above and abruptly
apiculate in the center, about 3.1 mm. long and 3.8 mm. wide across the disc when
expanded. Column very short and stout, about 1.4 mm. high at the back.
San Martin: In a deep ravine near the top of a ridge east of Tingo
Maria, 525-1100 meters, on fallen tree, Allard 21577.
FIG. 190. Dichaea panamensis Lindl. Plant; X 1. 1, flower, spread open, side
view; X 3. 2, lateral sepal; X 3. 3, dorsal sepal; X 3. 4, petal; X 3. 5, column,
side view; X 3. 6, pollinia and stipe; much enlarged.
971
FIG. 191. Dichaea tennis C. Schweinf. 1, plant; X 11A- 2, flower from side,
natural position; X 9. 3, lip expanded; X 7. 4, petal; X 7. 5, dorsal sepal
expanded; X 7. 6, lateral sepal expanded; X 7. 7, column from side; X 7.
972
SCHWEINFURTH: ORCHIDS OF PERU 973
PACHYPHYLLUM HBK.
Inconspicuous, minute to medium-sized epiphytes without pseudo-
bulbs, extending from Costa Rica and Panama to Colombia, Ecuador,
Peru and Bolivia.
Stems short or long, simple to more or less branched, erect to arcuate and
pendent, often caespitose, entirely concealed by densely imbricating leaf-sheaths.
Leaves distichous, articulated, numerous, short, fleshy, spreading, often condupli-
cate and recurved. Inflorescences axillary, short, racemose, few-flowered or re-
duced to one flower. Flowers small to minute, shortly pedicelled. Sepals free or
more or less connate, equal or nearly so. Petals similar to the sepals, often broader.
Lip erect, simple or obscurely 3-lobed, longitudinally concave, more or less adnate
to the sepaline tube, the disc usually provided in the middle or below with a pair of
short, broad keels or calli. Column very short, stout, dilated into a large petaloid
clinandrium. Anther opercular, incumbent. Pollinia 2, obovoid-oblong, sessile on
a small gland.
Al. Petals and anterior portion of the lip minutely denticulate; anterior portion
of the lip distinctly narrower than the basal portion and often recurved.
P. Pastii
A2. Petals and anterior portion of the lip entire; anterior portion of the lip only
slightly narrower than the basal portion or broader I
la. Leaves elliptic or oval to oblong-elliptic, never equitant or conduplicate ... 1
Ib. Leaves not elliptic (sometimes broadly oblong) in outline, often conduplicate
or equitant 3
la. Disc of the lip naked, i.e., destitute of lamellae or tubercles. .P. parvifolium
Ib. Disc of the lip with a pair of prominent lamellae or tubercles 2
2a. Plant relatively small and slender, leaves 8 mm. or less long; lip broadly
rounded at the apex P. gracillimum
2b. Plant relatively large and stout, leaves 11 mm. or more (commonly 15 mm.)
long; lip strongly acute or acuminate P. breviconnatum
3a. Sepals, petals and lip strongly adnate into a tube below 4
3b. Sepals, petals and lip free or adnate only at the very base 5
4a. Leaves fimbriate-ciliate; lip broadly ovate, acute or subacute, with 2 small
basal calli P. muscoides
4b. Leaves entire, smooth on the margin; lip broadly obovate, strongly obtuse or
rounded at the apex, with 2 large triangular lamellae near the middle.
P. tenue
5a. Leaves triangular when seen from the side; lip obscurely 3-lobed with a fold
on each side P. capitatum
5b. Leaves oblong, ligulate, or semilunate when seen from the side; lip strictly
simple 6
6a. Lip rhombic-ovate or rhombic-ligulate, with a pair of short, fleshy calli; leaves
commonly conduplicate P. distichum
6b. Lip oblong, obovate-oblong or pandurate 7
7a. Leaves apparently not conduplicate; lip oblong, obtuse P. lycopodioides1
7b. Leaves distinctly conduplicate; lip strongly acute or mucronate 8
1 This species, of which we have not seen any authentic material, is rather
obscure and may be referable to P. distichum.
974 FIELDIANA: BOTANY, VOLUME 30
8a. Plant relatively tall, 15 cm. or more high; leaf-sheaths shining, transversely
rugose; leaves about 5 mm. wide in natural position P. serra
8b. Plant relatively short, about 9 cm. or less high; leaf-sheaths not transversely
rugose; leaves about 2-3 mm. wide in natural position P. pectinatum
Pachyphyllum breviconnatum Schltr. Fedde Repert. Beih. 9:
115. 1921; Mansf. Fedde Repert. Beih. 57: t. 130, nr. 510. 1929.
Plant large for the genus, pendent, up to 53 cm. or more long. Stems flexuous
or arcuate, more or less branched, rather slender, entirely or mostly concealed by
loose, infundibuliform leaf-sheaths. Leaves small, very numerous, distichous, artic-
ulated, wide-spreading, elliptic to elliptic-oblong ("obliquely oblong"), acute or
subacute and prominently mucronate, minutely ciliolate near the apex, fleshy,
1.1 (rarely) -2.3 cm. long, up to 7.5 mm. wide, rather loose. Inflorescences numer-
ous, axillary, shorter than the leaves, commonly 3- or 4-flowered above; floral
bracts spreading, amplexicaul, triangular-lanceolate to ovate-triangular, acumi-
nate, the lower ones successively larger. Flowers rather large for the genus, white
or primrose yellow, fleshy, with wide-spreading or recurved segments, slightly con-
nate below. Dorsal sepal elliptic-lanceolate or oblong-lanceolate, acute, mucro-
nate, dorsally strongly carinate, about 6 mm. long and 1.5-1.9 mm. wide. Lateral
sepals obliquely oblong-lanceolate, more or less lightly falcate, acute, mucronate,
dorsally carinate, much longer and slightly wider than the dorsal sepal, consider-
ably connate below. Petals obliquely oblong-lanceolate, acute, mucronate, sim-
ilar to the dorsal sepal but shorter and either slightly narrower or broader. Lip
with the claw adnate to the sepaline tube; free portion lanceolate-elliptic, sharply
acute, about 5-5.5 mm. long and 2-2.2 mm. wide; disc below the middle with a
pair of spreading, fleshy, semiorbicular-conic calli. Column rather short and stout,
about 3-4 mm. high at the back, with a large, concave, retuse clinandrium. Ovary
trigonous, trialate.
Cuzco: Prov. of Paucartambo, below Tres Cruces Inn, 3600-
3700 meters, Weberbauer 6976 (type). "Pillahuata," Cerro de Cusil-
luyoc, 2800-3100 meters, epiphytic in thickets, Pennell 14129. The
latter collection, which is somewhat smaller throughout than the
type, has "primrose-yellow" flowers.
Pachyphyllum capita turn Kranzl. Engler, Bot. Jahrb. 37: 386.
1906; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 83), 21. 1923.
Plant medium-sized. Stems rather stout, straight or scarcely curved, 10-1 5 cm.
tall, densely leafy. Leaves equitant, triangular as seen from the side, obtuse,
thickly fleshy, about 10 mm. long and 5-6 mm. wide. Inflorescences axillary on
the upper portion of the plant, a little surpassing the leaves, densely 3- to 4-flow-
ered and subcapitate; floral bracts minute. Flowers small, white, shining without.
Dorsal sepal oblong, acute, 4-5 mm. long, 2 mm. wide. Lateral sepals equally long,
ovate-oblong, acute. Petals oblong, apiculate, rather broader than the sepals.
Lip oblong-obovate, obscurely 3-lobed above with a fold on each side, rounded,
retuse and apiculate at the apex; disc through the lower portion with a pair of
lamellae (higher in front). Column very broad and foliaceous.
SCHWEINFURTH: ORCHIDS OF PERU 975
Amazonas: East of Chachapoyas between Tambo Ventillas and
Piscohuanuna, 2600-2700 meters, in grass-steppe with scattering
shrubs, Weberbauer 4421.
No example of this species was available.
Pachyphyllum denticulatum (Ruiz & Pav.) Schltr. Fedde Re-
pert. Beih. 9: 180. 1921. Fernandezia denticulata Ruiz & Pav. Syst.
Veg. Fl. Peruv. et Chil. 1 : 240. 1798. = an obscure concept, perhaps
referable to P. distichum HBK.
Huanuco: Pillao, on trees and rocks, Pavdn s.n.
Pachyphyllum distichum HBK. Nov. Gen. et Sp. 1: 339, t.
77. 1816; Reichb. f. Bonpl. 3: 219. 1855; Xen. Orch. 1: 165, t. 58,
fig. II, 5-7. 1856; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 83), 25.
1923.
Plant epiphytic or rarely terrestrial, commonly rather large and stout. Stems
usually arcuate, simple or more commonly branched, about 52 cm. or less long,
entirely or nearly enveloped by distichous, loose leaf-sheaths. Leaves very numer-
ous, rather dense, broadly to narrowly oblong, articulated, usually strongly com-
pressed and conduplicate in the dried specimen, more or less falcately recurved,
apiculate at the rounded apex, minutely serrulate-ciliate throughout, up to 3.1 cm.
long and 1 cm. wide when expanded. Inflorescences numerous, axillary, shorter
than the leaves, racemose, recurved, densely about 6- to 8-flowered; floral bracts
amplexicaul, conduplicate, triangular-ovate, ciliolate. Perianth segments free.
Flowers very small, green or greenish white, with the floral parts fleshy and cam-
panulate. Dorsal sepal oblong, navicular, acute, dorsally carinate, up to 3.5 mm.
long and 1.4 mm. wide. Lateral sepals oblong-lanceolate, acute, navicular, lightly
oblique, somewhat longer and slightly narrower than the dorsal sepal. Petals
obliquely oblong, sometimes broader above, acute, slightly shorter and narrower
than the dorsal sepal. Lip oblong-lanceolate, rhombic-lanceolate or rhombic-ovate,
longitudinally concave, obtuse, up to about 4 mm. long and 2.2 mm. wide; disc
above the middle with a pair of fleshy, finger-like calli. Column relatively large
and stout, dilated upward, trilobulate at the apex, up to about 2.6 mm. high at the
back. Ovary trigonous, narrowly trialate.
Cuzco: Habitat not recorded, Bues 2158. Prov. of Convention,
Tanamanche-Quellomayo, 3750 meters, Vargas 4445. "Pillahuata,"
Cerro de Cusilluyoc, 2800-3100 meters, epiphyte in thicket, Pennell
14128. Prov. of Urubamba, near Wenner Gren ruins, 3400-3600
meters, in dense wet dark forest with much fog and rain, Metcalf
30767 (fl. old on fruit). — Huanuco: Cani, 7 miles northeast of Mito,
about 2600 meters, Macbride 3462. Prov. of Pachitea, above La
Molina near Panao, on bushy slope, Asplund 13643. Also Colombia
and Ecuador (type).
976 FIELDIANA: BOTANY, VOLUME 30
Pachyphyllum gracillimum C. Schweinf. Am. Orch. Soc. Bull.
16: 564, t, 1947. Figure 192.
Plant epiphytic, very slender and lax. Stems short to relatively elongate, lax,
usually remotely branched, about 37 cm. or less long, entirely (or mostly) con-
cealed by loose spreading leaf-sheaths. Leaves numerous, distichous, very
small, articulated, elliptic to lanceolate-elliptic or oblong-elliptic, never equitant,
acute and prominently mucronate, flat and never conduplicate, wide-spread-
ing, 6-8 mm. long, up to 3.3 mm. wide, minutely ciliolate on the upper margins.
Inflorescences very numerous, axillary, shorter than the leaves, racemose, densely
3- to 6-flowered, nodding in the upper half. Floral bracts semiamplexicaul, spread-
ing, triangular, conduplicate, larger toward the base of the raceme. Flowers very
small, yellow-greenish, campanulate, fleshy, the sepals, petals and lip connate be-
low to form a short tube. Dorsal sepal (including the connate basal portion)
oblong or ovate-oblong, subacute at the rounded apex, about 3 mm. long and
1.2 mm. wide. Lateral sepals similar to the dorsal sepal but somewhat narrower,
oblong, navicular, slightly oblique. Petals very similar to the dorsal sepal but
shorter and slightly narrower, ovate-oblong, acute. Lip small, adnate to the sepa-
line tube, oblong-oval, broadly rounded at the apex, about 1.8 mm. long (free part)
and 1.1 mm. wide; disc below the middle with a pair of erect-spreading, obliquely
triangular lamellae which are truncated on the outer angle. Column very short
and stout, about 1.7 mm. high, with a large concave clinandrium. Ovary trigo-
nous and trialate.
Cuzco: Prov. of Urubamba, Puyupata-Tuncapata, 3200 meters,
Vargas 2932 (type). Prov. of Urubamba, near Wenner Gren ruins,
3400-3600 meters, in dense wet forest with much fog and rain,
Metcalf 30770.
Pachyphyllum haemathodes (as "haematodes") (Ruiz & Pav.)
Schltr. Fedde Repert. Beih. 9: 180. 1921. Fernandezia haemathodes
Ruiz & Pav. Syst. Veg. Fl. Peruv. et Chil. 1: 240. 1798.= obscure
concept, probably referable to Maxillaria crassicaulis C. Schweinf.,
a new name for Ornithidium Weberbauerianum Kranzl.
Huanuco: Mufia, on trees and rocks, Pavdn s.n.
Pachyphyllum lycopodioides Schltr. Fedde Repert. Beih. 9:
116. 1921; Mansf. Fedde Repert. Beih. 57: t. 130, nr. 511. 1929.
Pachyphyllum Pastii Kranzl. ex Weberb. Pflanzenw. Peru. And. 254.
1911, not Reichb. f.
Plant epiphytic, erect, about 25 cm. high. Stems strict, rather stout, densely
leafy, entirely concealed by close, imbricating, denticulate leaf-sheaths. Leaves
recurved-spreading, oblong-ligulate, apiculate, fleshy, sparingly papillose-serrulate
on the margin, up to 9 mm. long, about 3 mm. wide. Inflorescences lateral, 3- to
4-flowered, erect-spreading but nodding above; peduncle subcompressed, up to
5 mm. long; floral bracts ovate, short-acuminate. Flowers small, white, glabrous.
Sepals shortly connate at the base. Dorsal sepal ovate-oblong ("oblong-ligulate").
PACHYPHXi-L-UM
C/.
FIG. 192. Pachyphyllum gradllimum C. Schweinf. 1, plant; X 1. 2, flower
from side; X 8. 3, flower opened out; X 12. 4, lip from side, natural position;
X 12.
977
978 FIELDIANA: BOTANY, VOLUME 30
mucronulate, sharply carinate on the back, about 3.75 mm. long and 1.25 mm.
wide. Lateral sepals oblong-lanceolate ("oblong-ligulate"), oblique, carinate, ap-
parently longer than the dorsal sepal. Petals obliquely and narrowly elliptic-
oblong, subacute, a little shorter than the dorsal sepal, entire. Lip suberect and
parallel to the column, shortly connate at the base with the sepaline tube, oblong,
obtuse, equally long with the petals, slightly narrowed in the middle of each side;
disc in the middle with a pair of obliquely subulate, subobtuse calli. Column very
short and stout, about 2 mm. high, with a broad, concave clinandrium which is
obscurely trilobulate at the truncate apex. Ovary clavate, trigonous.
Huanuco: Southwest of Monzon, on the mountains, 3400-3500
meters, Weberbauer 3327.
No example of this species was available.
Pachyphyllum muscoides (Kranzl.) Schltr. Fedde Repert. 15:
216. 1918. Orchidotypus muscoides Kranzl. Engler, Bot. Jahrb. 37:
383. 1906; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 83), 59, fig. 3 C,
a-e. 1923.
Plant minute, epiphytic. Stems suberect to arcuate or lax, simple or very
rarely branched, slender, about 7.5 cm. or less long, entirely concealed by leaf-
sheaths. Leaves very small, distichous, more or less numerous, articulated, ellip-
tic-oblong to oblong or linear-oblong, acute, mucronate, often conduplicate and
falcate-recurved in the dried specimen, conspicuously ciliate throughout, about
7 mm. or less long and 1-1.5 mm. wide. Inflorescences axillary, abbreviated, much
shorter than the leaves, apparently 1- or 2-flowered. Flowers pale green to yellow-
ish white, very small, campanulate, about 2 mm. long. Parts of the perianth all
connate into a tube below the middle. Dorsal sepal (free portion) oblong-ovate,
acute or obtuse, about 1 mm. long and 0.6 mm. wide. Lateral sepals (free portion)
obliquely lanceolate-ovate, acute, little longer than the dorsal sepal. Petals (free
portion) ovate, subacute, overlapping the sepals at the sides, similar to the dorsal
sepal but a little shorter and wider. Lip with narrow claw adnate to the sepaline
tube; free portion rotund-ovate, subacute, about 1-1.2 mm. long and 1 mm. broad;
disc at the base on each side with a small, erect, obliquely subquadrate lamella.
Column relatively large, stout, about 1.4 mm. high, mostly occupied by a large,
concave clinandrium.
Cajamarca: Prov. of Chota, mountains west of Huambos, 2700-
3000 meters, in sclerophyllous vegetation, especially shrubs with
scattering small trees, Weberbauer 4189 (type, as Orchidotypus).
Also Colombia and Central America (Costa Rica, Panama).
Pachyphyllum parvifolium Lindl. Hook. Ic. PI. 2: 1. 117. 1837;
Kranzl. Engler, Pflanzenr. IV. 50 (Heft 83), 23. 1923.
Plant slender. Stems few, slender, commonly several-branched with remote
ascending branches, up to about 18 cm. long, lightly flexuous, entirely concealed
by leaf-sheaths. Leaves very numerous, distichous, rather loose, articulated, ob-
long-elliptic to oval, obtuse, never conduplicate, wide-spreading, about 13 mm. or
SCHWEINFURTH: ORCHIDS OF PERU 979
less long and 4 mm. or less wide. Inflorescences numerous, axillary, commonly
densely 3-flowered (rarely 4-flowered), about equaling or a little shorter than the
leaves. Flowers small, campanulate, about 4 mm. long. Dorsal sepal oblong to
elliptic-oblong, acute, concave, navicular, dorsally carinate. Lateral sepals some-
what longer, obliquely oblong-lanceolate, acute, navicular, dorsally carinate.
Petals similar to the dorsal sepal, obtuse, ecarinate. Lip very similar to the
petals, apparently elliptic-oblong, entirely naked (without lamellae or tubercles).
Column a little shorter than the petals, broadly winged, elliptic-obovate in out-
line, concave, lightly retuse at the apex.
Amazonas: Chachapoyas, Mathews 1836.
No material of this species was available, but there is a photo-
graph of the type in the Ames Herbarium.
Pachyphyllum Pastii Reichb. f. Bonpl. 3: 239. 1855; Xen. Orch.
1: 165, t. 58, fig. I, 1-4. 1856; Kranzl. Engler, Pflanzenr. IV. 50
(Heft. 83), 25. 1923. Pachyphyllum Stuebelii Schltr. Fedde Repert.
Beih. 7: 204. 1920; Mansf. Fedde Repert. Beih. 57: t. 73, nr. 283.
1929.
Plant epiphytic, medium-sized for the genus. Stems rather stout, often aggre-
gated, simple or more commonly branched, up to 65 cm. or more long, entirely or
mostly concealed by veiny, ciliate-margined sheaths. Leaves very numerous, dis-
tichous, articulated, wide-spreading, rotund-ovate, oblong-oval or elliptic-oblong,
often conduplicate and falcate-recurved in the dried specimen, obtuse and mucro-
nate, ciliate-denticulate throughout, up to 14 mm. long and 7 mm. wide. Inflores-
cences more or less numerous, axillary, densely few- (2- to 6-) flowered and nodding
above, the peduncle much shorter than the leaves. Floral bracts semiamplexicaul,
conduplicate, triangular-ovate, acute or acuminate. Flowers very small, more or
less fleshy, campanulate, commonly yellowish green with a white lip. Sepals
shortly connate at the base. Dorsal sepal oblong to ovate-oblong (rarely ovate),
acute, mucronate, concave or navicular, dorsally carinate, about 3.2 (rarely 2) mm.
long, 1.5-1 .8 mm. wide. Lateral sepals similar to the dorsal sepal but a little longer,
obliquely oblong-lanceolate (rarely obliquely ovate). Petals oblong or lanceolate-
oblong (rarely oval-oblong), acute to subacute, more or less conspicuously ciliolate-
denticulate, a little shorter and more or less narrower than the dorsal sepal. Lip
elliptic-oblong or oblong-rhombic in outline and about 4 (rarely 3) mm. long when
expanded, more or less distinctly trilobulate above the concave base, with the an-
terior lobule more or less reflexed, subacute to obtuse, with the anterior margins
ciliolate-denticulate, about 2 mm. or more (rarely 1.8 mm.) wide near the middle;
disc in the middle with a pair of small, approximate digitiform calli. Column rela-
tively large, oblong-cylindric to oblong-ovoid, lightly trilobulate at the truncate
apex, about 2.1-2.7 (rarely 1.7) mm. high at the back. Ovary trigonous, nar-
rowly trialate.
Cuzco: Prov. of Convention, Santa Ana Valley, Alturas del
Chaco, Bues (Herrera 2124ty. Prov. of Urubamba, on trail from
Puyupata to Sayacmarca, 3600 meters, Vargas 2895. Prov. of Uru-
bamba, near Wenner Gren ruins, 3400-3600 meters, in wet shady
980 FIELDIANA: BOTANY, VOLUME 30
sphagnum bog with much fog and rain, Metcalf 30756. Same local-
ity as the last, in dense wet forest, Metcalf 30775. — Huanuco: Prov.
of Huamalies, mountains southwest of Monzon, 3400-3500 meters,
Weberbauer 3327. Also Colombia (type of P. Stuebelii) and Ecuador
(type of P. Pastii).
Pachyphyllum pectinatum Reichb. f. Xen. Orch. 3: 22. 1878;
Kranzl. Engler, Pflanzenr. IV. 50 (Heft 83), 22. 1923.
Plant epiphytic, small, simple or sparingly branched, about 10 cm. or less tall.
Stems erect or more or less arcuate, densely to loosely many-leaved, rather slender,
entirely concealed by leaf-sheaths which are longitudinally nervose. Leaves con-
duplicate, recurved, oblong-semilunate when viewed from the side, apiculate, with
the anterior margin minutely denticulate, about 5-8 (rarely 14) mm. long and
2-3 mm. wide in natural position. Inflorescences racemose, few-flowered, when
flowering a little shorter than the leaves. Floral bracts triangular or triangular-
lanceolate, much shorter than the pedicellate ovary. Flowers very small, campan-
ulate, white or ivory-color. Dorsal sepal oblong-lanceolate ("ligulate"), acute,
navicular, subcarinate, 2.3-3.2 mm. long, 0.8-1.1 mm. wide. Lateral sepals
obliquely oblong-lanceolate, acute, mucronate, carinate, slightly longer and
broader than the dorsal sepal. Petals oblanceolate or oblong-lanceolate, shortly
acuminate, slightly shorter than the dorsal sepal. Lip oblong-oblanceolate ("lig-
ulate-obovate"), acute, mucronate, longitudinally concave, 2-3.1 mm. long, 0.75-
1.25 mm. wide; disc with a pair of approximate keels extending from near the base
to above the middle where more or less dilated. Column stout, 1.6-2 mm. high
at the back, with a large suborbicular, concave wing throughout. Ovary trigonous.
Cuzco: Region of Vilcabamba, "Buena Cuesta," toward Co-
chayoc, about 2460 meters, Bues 1615. Same locality, Vargas 417.
— Puno: Prov. of Carabaya, "Ackopampa a Juro-Juro," 3500 meters,
on rocks, Vargas 7038. Also Bolivia (type).
Pachyphyllum serra Reichb. f. Bonpl. 3: 219. 1855; Reichb. f.
Xen. Orch. 1: 167, t. 58, fig. IV, 13-17. 1856; Kranzl. Engler, Pflan-
zenr. IV. 50 (Heft 83), 26. 1923.
Plant probably epiphytic, apparently simple, about 15 cm. or more tall. Stem
stout, strict, entirely concealed by leaf-sheaths which are longitudinally and trans-
versely rugose and shining. Leaves very numerous and dense, spreading, con-
duplicate, oblong-semilunate, recurved, acute, about 1.5 cm. long and 5 mm. wide
in natural position, with the anterior margin minutely denticulate. Inflorescences
very short, racemose, few-flowered, much shorter than the leaves. Floral bracts
triangular, equal to the pedicellate ovary. Flowers very small, with subparallel
segments. Sepals ligulate, acute, carinate, about 3.5 mm. long and scarcely 1 mm.
wide (according to Kranzlin). Petals ligulate-oblanceolate, acute or shortly acu-
minate, slightly shorter and narrower than the sepals. Lip ligulate-pandurate,
acute or acuminate, rounded at the base; disc in front of the constriction with a
pair of elongate keels which are angled in front. Column 3-lobed at the apex with
the mid-lobe longer and retuse. Ovary trigonous, slender.
SCHWEINFURTH: ORCHIDS OF PERU 981
Amazonas: Chachapoyas, Mathews s.n.
No material or other record of this species has been examined.
Pachyphyllum subbiflorum (Ruiz & Pav.) Schltr. Fedde Re-
pert. Beih. 9: 181. 1921. Fernandezia subbiflora Ruiz & Pav. Syst.
Veg. Fl. Peruv. et Chil. 1: 240. 1798.= obscure species.
Huanuco: Muna, on trees and rocks, Pavon s.n.
Pachyphyllum tenue Schltr. Fedde Repert. Beih. 9: 116. 1921;
Mansf. Fedde Repert. Beih. 57: t. 131, nr. 512. 1929.
Plant epiphytic, small, slender, about 8 cm. tall. Stem simple, arcuate, very
slender, entirely concealed by leaf-sheaths. Leaves rather loose, spreading, ligu-
late, apiculate, entire, 4.5-6 mm. long, scarcely exceeding 1 mm. wide in the middle.
Inflorescences solitary, densely 2- to 3-flowered, erect-spreading, slightly exceeding
the leaves, the peduncle a little shorter than the leaves. Floral bracts ovate-
deltoid, acute, much shorter than the pedicellate ovary. Flowers minute, campan-
ulate, erect-spreading, with sepals, petals and lip shortly connate below. Dorsal
sepal oblong or ovate-oblong, obtuse, dorsally carinate, about 1.5 mm. long. Lat-
eral sepals apparently a little longer and much wider than the dorsal sepal, obliquely
oblong-ovate. Petals oblong-elliptic ("oblong"), obtuse, a little shorter than the
sepals, with the lower sides overlapping the sepals. Lip broadly obovate in out-
line, rounded in front, adnate to the sepaline tube by the narrowed, clawed base,
as long as the petals, broader than the sepals in the upper part; disc below the
middle with a pair of large, obliquely triangular, wing-like lamellae (highest at the
truncate apex). Column short, scarcely 1 mm. high, terminated by a large con-
cave-cucullate, trilobulate clinandrium. Ovary trigonous.
Junin: Chanchamayo Valley, Koehler s.n.
No example of this species was available.
CENTROPETALUM Lindl. (including Nasonia Lindl.)
A small group of South American epiphytes apparently limited
to Colombia, Ecuador and Peru.
Plants very small to medium-sized, without pseudobulbs. Stems simple or
rarely branched, erect or commonly arcuate, entirely concealed by leaf-bearing
sheaths. Leaves numerous, distichous, small, flat to conduplicate or equitant,
articulated, commonly recurved or falcate, fleshy, caducous. Inflorescences lat-
eral, axillary, 1- or rarely 2-flowered. Flowers small to medium-sized, often large
in relation to the plant, yellow, orange, red or purple. Sepals subequal, narrow,
free or the laterals partly connate, more or less carinate (especially the laterals).
Petals subequally long but commonly much wider than the sepals. Lip similar
to the rest of the perianth or much larger, sessile or short-clawed, entire or rarely
subtrilobed. Column short, with a large petaloid clinandrium. Pollinia 4, on two
free or partly free stipes.
982 FIELDIANA: BOTANY, VOLUME 30
Al. Lip about equaling the other perianth segments, elliptic-lanceolate with a
saccate base C. Hartwegii
A2. Lip larger (especially broader) than the other segments (commonly much
larger), suborbicular, triangular-suborbicular or rhombic-obovate, without a
saccate base I
la. Lamina of the lip triangular-cordate or suborbicular-cordate, with a strongly
cordate base; base of the disc with a long keel terminated by a bilobed callus.
C. ionantherum
Ib. Lamina of the lip not triangular-cordate, commonly with a cuneate base;
base of the disc without a long keel 1
la. Lip suborbicular, slightly longer than the sepals, with a bilobed callus above
the base C. nigro-signatum1
Ib. Lip rhombic-obovate to transversely oval, much longer than the sepals, with
a small concave callus (rarely bilobed) at the base C. distichum
Centropetalum distichum Lindl. Sert. Orch. sub t. 21. 71838;
Kranzl. Engler, Pflanzenr. IV. 50 (Heft 83), 30. 1923; C. Schweinf.
Bot. Mus. Leafl. Harvard Univ. 17: 62. 1955. Centropetalum Wars-
cewiczii Reichb. f. Bot. Zeit. 10: 706. 1852; Xen. Orch. 1: 96, t. 38,
fig. 1, 1-5. 1855. Fernandezia disticha Schltr. Fedde Repert. 16: 349.
1920. Fernandezia Warscewiczii Schltr. Fedde Repert. 16: 349. 1920.
Fernandezia pulchra Schltr. Fedde Repert. Beih. 9: 117. 1921; Mansf.
Fedde Repert. Beih. 57: t. 131, nr. 513. 1929.
Plant small, suberect or commonly more or less arcuate. Stems simple, up
to about 12 cm. long, entirely concealed by striate-sulcate leaf-bearing sheaths.
Leaves numerous, subdensely distichous, linear, articulated, caducous, condupli-
cate or subterete, lightly recurved and falcate, acute, fleshy, about 7-15 mm. long,
about 2 mm. wide. Inflorescences very short, 1-flowered, in the axils of the upper
leaves. Pedicels rather long, with a trigonous ovary. Flower large for the plant,
yellow or orange-yellow to scarlet or purple. Sepals lanceolate or oblong-lanceolate,
acute or short-acuminate, dorsally carinate, about 7 mm. or less long, 2-2.7 mm.
wide, lateral sepals oblique and connate at the base. Petals about as long as the
sepals but often nearly twice as wide, ovate-elliptic, acute, apiculate. Lip conspic-
uously longer and much wider than the other segments, rhombic-obovate, cuneate-
flabellate or transversely oval, broadly cuneate toward the base, very broadly
rounded (and rarely retuse) at the apex, about 10 mm. long and 10-14 mm. wide;
disc with a very small ovate-oblong, concave (rarely bilobulate) callus at the base.
Column short, with a large cucullate and petaloid, denticulate, often trilobulate
clinandrium.
Amazonas: Chachapoyas, among lichens, Mathews s.n. (type of
C. distichum), Hubsch ex Reichb. f., fide Kranzlin (I.e.). — Cajamarca:
East side of the Cordillera, east of Huancabamba, epiphytic, in
sclerophyllous vegetation, Weberbauer 6141 (type of Fernandezia
pulchra). Also Ecuador (type of C. Warscewiczii) .
1 This concept was considered referable to C. ionantherum by Schlechter (Fedde
Repert. Beih. 9 [1921], 181), but it seems preferable to follow Kranzlin in consider-
ing it distinct.
SCHWEINFURTH: ORCHIDS OF PERU 983
Centropetalum Hartwegii (Reichb. f.) Kranzl. Engler, Pflan-
zenr. IV. 50 (Heft 83), 30. 1923. Nasonia Hartwegii Reichb. f. Xen.
Orch. 1: 97. 1855. Nasonia punctata Lindl. in sched., fide Kranz-
lin, I.e.
Plant small, stout, more or less arcuate. Stems commonly strongly curved,
stout, simple, about 22 cm. or less long, entirely concealed by loose, infundibuli-
form, striate, denticulate leaf-sheaths. Leaves numerous, densely distichous, flat,
lanceolate, lanceolate-oblong or elliptic-oblong, acute, strongly rugose in the dried
specimen, up to 2.3 cm. long, about 5-6 mm. wide. Inflorescences very short,
1- or 2-flowered, in the axils of the upper leaves. Pedicels long-exserted, terminat-
ing in a trigonous ovary. Flowers small but rather large for the plant, campanu-
late. Sepals similar, oblong-lanceolate ("triangular"), acute or acuminate, dorsally
carinate, about 1.2 cm. long. Petals similar to the sepals but somewhat broader,
acute. Lip narrowly elliptic-lanceolate from a minutely saccate base, subacute,
lightly sigmoid when seen from the side, about as long as the other floral segments,
about 2 mm. wide; disc with a pair of short lamellae in the middle. Column nearly
as long as the lip, with 3 subequal, round-ovate lobes.
Cajamarca(?), fide Schlechter, Fedde Repert. Beih. 9: 181. 1921.
Colombia (type).
No material of this species was available, but there are records
of the type of Nasonia Hartwegii in the Ames Herbarium.
Centropetalum ionantherum (Reichb. f. & Warsc.) Kranzl.
Engler, Pflanzenr. IV. 50 (Heft 83), 32. 1923. Nasonia ionanthera
Reichb. f. & Warsc. Bonpl. 2: 99. 1854; Xen. Orch. 1: 97, t. 38, fig. II,
6-14. 1855. Fernandezia ionanthera Schltr. Fedde Repert. 16: 349.
1920.
Plant small, erect to ascending or arcuate. Stems usually simple, rarely some-
what branching, up to 20 cm. long, entirely concealed by numerous loose, imbri-
cating, striate leaf-sheaths. Leaves several to numerous, densely or subdensely
distichous, conduplicate, falcate-recurved, semielliptic (rarely) to linear-oblong,
acute, caducous, fleshy, 2 cm. or less long, 2-3 mm. thick. Inflorescences very
short, 1-flowered, in the axils of the upper leaves. Pedicels long-exserted, terminat-
ing in a trigonous ovary. Flowers medium-sized but large for the plant, red-orange
or cinnabar-red, sometimes with a black spot. Dorsal sepal oblong-lanceolate
("triangular"), complicate-acute, dorsally carinate, about 1 cm. or less long and
2.3 mm. wide. Lateral sepals obliquely lanceolate ("ovate-triangular"), short-
acuminate, dorsally carinate, a little longer and broader than the dorsal sepal.
Petals oblong-elliptic ("ovate"), acute, slightly shorter than the sepals but nearly
twice as wide as the dorsal sepal. Lip much larger than the sepals and petals,
suborbicular-cordate or triangular-cordate, strongly cordate at the base, subacute
to broadly obtuse at the apex, about 1.5 cm. or less long and equally wide; disc
below with a long, thickened or carinate mid-nerve which terminates in a bilobed
callus. Column short and stout, with a large, cucullate, petaloid clinandrium which
is subentire or obscurely lobulate above.
984 FIELDIANA: BOTANY, VOLUME 30
Peru: Habitat unrecorded, in high mountains, Warscewicz s,n.
(type of Nasonia ionanthera). — Cajamarca: Prov. of Chota, pass
south of Conchan, 2500 meters, epiphyte on moss-covered branches,
Stork & Horton 10061. — Huanuco: Between Huanuco and Pampay-
aco (Pampayacu), Kanehira 327. Pillao, abundant on tree trunks
in rain-forest, 2700 meters, Woytkowski 37018. — Junin, fide Schlech-
ter, Fedde Repert. Beih. 9: 181. 1921.— Piura: Huancabamba, El
Tambo, Camino Piura, about 3000 meters, Scolnik 1403.
Centropetalum nigro-signatum Kranzl. Engler, Bot. Jahrb.
37: 388. 1906; Kranzl. Engler, Pflanzenr. IV. 50 (Heft 83), 31. 1923.
Plant small. Stems simple, erect or lightly arcuate, up to 10 cm. long, con-
cealed by short, transversely rugulose leaf-sheaths. Leaves more or less dense,
distichous, oblong-lanceolate, often conduplicate and recurved, acute and apicu-
late, 1-1.8 cm. long, up to 5 mm. wide, articulated, strongly rugulose in the dried
plant. Inflorescences very short, 1- or 2-flowered, on the upper portion of the stems.
Pedicels slender, long-exserted, terminating in a trigonous ovary. Flowers rather
small, scarlet, lip marked with a black spot. Sepals lanceolate, acute, dorsally
carinate especially the lateral sepals which are lightly dilated at the base, 8-9 mm.
long, 2.5 mm. wide. Petals oblong, acute, almost twice as wide as the sepals. Lip
much larger than the other segments, apparently suborbicular, minutely bilobed
in front, about 10 mm. long and broad; disc without thickened basal nerves but
with a very small bilobed callus a little above the base. Column with a very broad
cucullate clinandrium.
Cajamarca: Prov. of Chota, mountains west of Huambos, 2700-
3000 meters, epiphytic in sclerophyllous vegetation, especially shrubs
with scattering small trees, Weberbauer 41 78.
No material of this species was available, but the Ames Her-
barium contains a photograph of the type.
GAMPYLOGENTRUM Benth.
Inconspicuous and widespread epiphytes extending from Florida
(U.S.A.) through Mexico, Central America, the West Indies and
South America to Brazil and Argentina.
Plants small to medium-sized, non-pseudobulbous. Stems commonly more or
less elongate, more rarely strongly abbreviated or wanting and leafless. Leaves
(when present) more or less numerous, distichous, fleshy, lanceolate or oblong to
oblong-elliptic or oval. Inflorescences lateral, several- to many-flowered, often
very densely flowered. Floral bracts minute, persistent. Flowers very small to
minute, campanulate. Sepals and petals free, subequal, broad or narrow, often
somewhat spreading above. Lip sessile, simple or 3-lobed, ovate or triangular-
ovate to lanceolate, produced into a more or less elongate spur which is often
strongly recurved or arcuate. Column very short, wingless, footless. Anther
SCHWEINFURTH: ORCHIDS OF PERU 985
terminal, opercular, incumbent, 2-celled. Pollinia 2, globose, with 2 short, fili-
form stipes.
Al. Plants with more or less elongate, leafy stems I
A2. Plants with abbreviated, leafless stems 1
la. Small plants, with racemes much exceeding the leaves.
C. Ulaei var. peruvianum
Ib. Large plants, with racemes more or less shorter than the leaves.
C. micranthum
la. Spur of the lip about equaling or little exceeding the sepals; lip definitely
3-lobulate C. fasciola (C. Sullivanii, C. loretoense)
Ib. Spur of the lip twice as long as the sepals; lip nearly or quite simple.
C. minutum
Campylocentrum fasciola (Lindl.) Cogn. Martius Fl. Bras. 3,
pt. 6: 520, t. 106, fig. 1. 1906. Angraecum fasciola Lindl. Bot. Reg.
26: sub t. 68. 1840. Campylocentrum Sullivanii Fawc. & Rendle,
Journ. Bot. 47: 128. 1909; Fl. Jam. 1: 142, t. 31, figs. 1-4. 1910.
Campylocentrum loretoense Schltr. Fedde Repert. Beih. 9: 118. 1921;
Mansf. Fedde Repert. Beih. 57: t. 131, nr. 514. 1929. Campylocen-
trum Lankesteri Ames, Sched. Orch. 4: 57. 1923. Campylocentrum
multiflorum Schltr. Fedde Repert. Beih. 19: 156. 1923.
Plant epiphytic, small or very small, with conspicuous, elongate, fibrous roots
which are up to 30 cm. long. Stem abbreviated or none. Leaves wanting. Pe-
duncles caespitose, several to numerous (up to 40), up to 15 cm. long, subdensely
many-flowered with distichous (rarely secund) flowers, puberulent. Flowers mi-
nute, sessile, with membranaceous, campanulate segments, white, greenish white
or pale yellow (rarely brownish white). Sepals and petals 1-nerved (3-nerved
according to Cogniaux). Dorsal sepal elliptic-ovate or ovate-lanceolate, acute,
about 1.3-1.8 mm. long and 0.8 mm. wide. Lateral sepals obliquely oblong-ovate
or ovate-lanceolate, acute, apiculate, a little longer and narrower than the dorsal
sepal. Petals similar to the dorsal sepal but slightly narrower or wider. Lip prom-
inently spurred; lamina broadly triangular-ovate in outline, lightly 3-lobulate,
about 1-1.5 mm. long and equally wide or wider below; lateral lobes semiobovate-
rounded; mid-lobe triangular and acute, or broadly ovate and obtuse when ex-
panded, distinctly larger than the lateral lobes; spur short, saccate to cylindric-
clavate, straight to lightly arcuate, about 1.1-2 mm. long, shorter or slightly longer
than the lateral sepals. Column very short and stout.
Loreto: Near Moyobamba, Filomeno s.n. (type of C. loretoense).
Santa Rosa, Yurimaguas, lower Rio Huallaga, 155-210 meters, epi-
phyte on trunk of "Castafia," Williams 1+936. Near Yurimaguas,
Ule 58p. Also British Honduras to Costa Rica (types of C. Lankes-
teri and C. multiflorum), the West Indies (type of C. Sullivanii),
British Guiana (type of Angraecum fasciola) , Surinam and Brazil.
Campylocentrum micranthum (Lindl.) Rolfe, Orch. Rev. 11:
245. 1903; Cogn. Martius Fl. Bras. 3, pt. 6: 506, t. 103, fig. 2. 1906.
986 FIELDIANA: BOTANY, VOLUME 30
Angraecum micranthum Lindl. Bot. Reg. 21: t. 1772. 1835. Fig-
ure 193.
Plant medium-sized to large, very variable. Stem stout, more or less arcuate
and sometimes fractiflex, up to 52 cm. or more long, entirely concealed by close,
tubular, leaf-bearing sheaths which are commonly pierced by long, flexuous, ad-
ventitious roots. Leaves numerous, distichous, widely spreading, oblong, oblong-
elliptic to oval (rarely narrowly oblong), commonly very obliquely bilobed at the
acute to rounded apex, shortly cuneate at the sessile base, about 2-11 (rarely 12.5)
cm. long, up to 3 cm. wide. Inflorescences lateral, numerous, racemose, more or
less shorter than the leaves, densely many-flowered to the base, rarely up to 7 cm.
long. Flowers small, white or cream-color, more rarely greenish or lilac and rose-
color, with subparallel segments which are recurved above. Dorsal sepal linear-
oblong, acute or short-acuminate, 3-nerved, up to 6.1 mm. long and 1.7 mm. wide
near the base. Lateral sepals similar to the dorsal sepal but slightly longer,
abruptly oblique at the base. Petals similar to the dorsal sepal but markedly
smaller. Lip lanceolate in outline, lightly recurved, distinctly 3-lobed near the
base, tubular-involute, up to 5.8 mm. long, produced below into a distinct spur;
lateral lobes relatively small, semiobovate, rounded or dentiform above; mid-lobe
much longer, linear-lanceolate, acute or acuminate; spur slightly shorter than the
lip or subequaling it, strongly arcuate, narrowly cylindric-clavate, broadly rounded
at the apex. Column very short and stout.
Junin: Schunke Hacienda above San Ramon, 1300-1700 meters,
in dense forest, Schunke A 52 (flowers poor and abnormal) . — Loreto :
Vicinity of Iquitos, 100 meters, Klug 10132. — San Martin: Juan Jui,
Alto Rio Huallaga, 400-800 meters, in forest, Klug 431 7. San Roque,
1350-1500 meters, epiphyte on edge of water, Williams 7702. Also
Mexico through Central America to Panama, the West Indies and
South America (type, perhaps Surinam) to Brazil and Peru.
Campylocentrum minutum C. Schweinf. Am. Orch. Soc. Bull.
17: 108, t., 1948. Figure 194.
Plant very small, epiphytic, stemless, leafless. Roots elongate, fibrous, very
slender, climbing along slender branches. Inflorescences fascicled, about 6 or less,
short, about 2.3 cm. long including the flowers, subdensely racemose nearly to the
base, puberulent. Flowers minute, yellow, membranaceous, 14 or less. Dorsal
sepal ovate, acute, about 1 mm. long and 0.5 mm. wide. Lateral sepals similar to
the dorsal sepal but slightly longer, obliquely ovate, conspicuously apiculate.
Petals ovate-elliptic, acute, slightly smaller than the dorsal sepal. Lip promi-
nently spurred; lamina simple or nearly so, very broadly triangular-ovate, acute,
nearly 1 mm. long, 1.5 mm. wide when expanded; spur about twice as long as the
sepals, distinctly arcuate, thick-cylindric, about 2 mm. or more long, rounded at
the apex. Column minute, stout. Ovary glabrous.
FIG. 193. Campylocentrum micranthum (Lindl.) Rolfe. Plant; X 1. 1, in-
florescence; X 4. 2, lip and column from above; about X 10. 3, column from
side; about X 10.
IT
987
988 FIELDIANA: BOTANY, VOLUME 30
Loreto: Mishuyacu, near Iquitos, 100 meters, climbing along a
slender branch of a forest tree, Klug 923.
Campylocentrum Ulaei Cogn. var. peruvianum C. Schweinf.
Bot. Mus. Leafl. Harvard Univ. 16: 19. 1953.
Plant small, slender, epiphytic. Stem slender, about 4.5 cm. long in the type
specimen, lightly arcuate, below producing elongate fibrous roots, entirely con-
cealed by loose, conduplicate leaf-sheaths. Leaves several, distichous, wide-
spreading, small, narrowly lanceolate to narrowly elliptic-oblong, narrowed to a
minutely and commonly obliquely bilobed apex, slightly contracted to a sessile
base, up to 2.1 cm. long and 5 mm. wide. Inflorescences lateral, racemose, much
exceeding the leaves, puncturing the middle of the leaf-sheaths, densely many-
flowered, up to about 6.5 cm. long, glabrous; peduncle filiform, up to 1.6 cm. long.
Flowers very small, with membranaceous, subparallel segments. Dorsal sepal
linear-lanceolate, subacute, 1-nerved, about 2.3 mm. long and 0.8 mm. wide.
Lateral sepals subequaling the dorsal sepal, narrowly and obliquely triangular-
lanceolate, acute or acuminate. Petals triangular-lanceolate, acute or acuminate,
slightly shorter than the sepals. Lip conspicuously spurred; lamina tubular-invo-
lute in natural position, ovate-lanceolate in outline when expanded, about 2 mm.
long, deeply 3-lobed below the middle; lateral lobes relatively short, obliquely sub-
quadrate and slightly dilated above, with an obtuse outer tip; mid-lobe much pro-
tuberant, narrowly lanceolate-triangular, long-acuminate; spur relatively large,
U-shaped when viewed from the side, slender and cylindric below, laterally flat-
tened and dilated above, about 1.3 mm. long from the rounded apex to the base of
the curve. Column minute. Ovary glabrous.
Junin: Chanchamayo Valley, 1800 meters, Schunke 520.
The following collections were indeterminable due to lack of the
requisite characters:
Genus uncertain. Flower poor.
Loreto: On right bank of Rio Santiago, above Pongo de Man-
seriche, 200 meters, epiphytic on trunk of tall forest tree, in small
colonies, fruit pale green, Mexia 6165.
Genus uncertain. No flowers.
Junin: Chanchamayo Valley, 1500 meters, Schunke 1671.
Genus uncertain, perhaps Masdevallia. No flowers.
Junin: Schunke Hacienda, above San Ramon, 1400-1700 meters,
epiphyte in dense forest, perianth yellow, Killip & Smith 24525.
Genus uncertain, perhaps Spiranthes. Only floral remnants.
FIG. 194. Campylocenirum minutum C. Schweinf . 1, plant; X 2. 2, flower from
side, natural position; X 16. 3, lip and spur from above; X 16. 4, dorsal sepal;
X 32. 5, petal; X 32. 6, lateral sepal; X 32.
989
990 FIELDIANA: BOTANY, VOLUME 30
Junin: Pichis Trail, Enefias, 1600-1900 meters, terrestrial in dense
forest, Kittip & Smith 25652.
Genus uncertain, perhaps Spiranthes. Very immature.
Junin: Puerto Bermudez, about 375 meters, epiphyte(?) in dense
forest, bracts pink, Killip & Smith 26587.
Genus uncertain, perhaps Lycaste. Old fruit.
Junin : La Merced, about 700 meters, terrestrial in wooded valley,
fruit green, Killip & Smith 23745.
Genus uncertain. Old fruit.
Junin: Pichis Trail, Enenas, 1600-1900 meters, epiphyte in dense
forest, Killip & Smith 25724.
Genus uncertain, perhaps Erythrodes. Flowers very old.
Junin: Pichis Trail, Dos de Mayo, 1700-1900 meters, terrestrial
herb 3-4 feet high, in dense forest, Killip & Smith 25810.
Genus uncertain, perhaps Lycomormium. Young bud only.
Junin: Colonia Perene, about 680 meters, epiphyte in dense forest,
Killip & Smith 24946.
Genus uncertain, perhaps Scelochilus. Flower poor.
Junin: Chanchamayo Valley, Schunke 570.
Genus uncertain. Flower resembles Epidendrum ibaguense HBK.
Huanuco: Pampayacu, Hacienda at mouth of Rio Chinchao,
about 1070 meters, on grassy cliff, Macbride 5018.
Genus uncertain. No flowers.
Huanuco: Llata, about 2100 meters, in rock crevices of cliff,
Macbride & Featherstone 2257.
Genus uncertain. Only vegetative parts present.
Huanuco: Yanano, about 1800 meters, pendent on high trees,
Macbride 3851 .
Genus uncertain, perhaps Stenoptera. No flowers; only vegetative
parts.
Cuzco: Anta, environs of the town of Huaracondo, 3200 meters,
Herrera 3628.
SCHWEINFURTH: ORCHIDS OF PERU 991
Genus uncertain. No flowers.
Loreto: Wooded banks of Rio Itaya above Iquitos, about 110
meters, epiphyte, Kittip & Smith 29292.
Genus uncertain, perhaps Brassia. No flowers present.
San Martin: San Roque, 1350-1500 meters, Williams 7346.
Habenaria sp. Aff. H. Guentheriana Kranzl. Flowers too im-
perfect for certainty.
Huayaquil. Ex Herb. Ruiz & Pav. s.n.
Habenaria sp. Aff. H. caldensis Kranzl.
Cuzco: Hipayoc, 950 meters, Bues s.n.
Habenaria sp. Aff. H. distans Griseb. Raceme in fruit.
Ayacucho: Ccarrapa, between Huanta and Rio Apurimac, about
1500 meters, terrestrial in densely forested valley, perianth green,
Kittip & Smith 22389.
Vanilla sp. No flowers.
Ayacucho: Rio Apurimac Valley, near Kimpitiriki, 400 meters,
vine climbing over high trees in dense forest, Kittip & Smith 23050.
Vanilla sp. No flowers.
Junin: Rio Pinedo, north of La Merced, 700-900 meters, herba-
ceous vine in dense woods, Kittip & Smith 23637.
Vanilla sp. No flowers.
Loreto: Iquitos, about 100 meters, vine in woods, Killip & Smith
27430.
Vanilla sp. No flowers.
Loreto: Lower Rio Huallaga, Santa Rosa, 155-210 meters, in
forest, Williams 4893.
Vanilla appendiculata Rolfe aff. Flower not found.
Loreto: Mishuyacu, near Iquitos, 100 meters, epiphyte in forest,
flowers cream color, called "Bellaco Vainill," Klug 1338.
Vanilla bicolor Lindl. aff. Flower poor.
Loreto: Alto Rio Itaya, San Antonio, 145 meters, Williams 3494-
Epistephium Duckei Huber? Flowers poor.
Cuzco: Prov. of Convention, Potrero, 1300 meters, epiphyte,
Vargas 1847.
992 FIELDIANA: BOTANY, VOLUME 30
Elleanthus aurantiacus (Lindl.) Reichb. f. aff. No flowers.
Huanuco: Playapampa, about 2700 meters, terrestrial in clumps
on sphagnum slopes, Macbride 4497-
Elleanthus aurantiacus (Lindl.) Reichb. f. aff. No flowers.
Huanuco: Vilcabamba, Hacienda on Rio Chinchao, about 1800
meters, on clay banks, flowers orange, Macbride 51 71 .
Elleanthus capitatus (R. Br.) Reichb. f. aff. No flower seen.
Cuzco: Prov. of Convention, Machu-Picchu, 2400 meters, Vargas
835.
Elleanthus capitatus (R. Br.) Reichb. f. aff. No flower seen.
Cuzco: Urubamba Basin, 2000-2300 meters, Herrera 2033 in part.
Elleanthus casapensis Reichb. f. aff. No flowers.
Junin : Schunke Hacienda, above San Ramon, 1300-1700 meters,
in dense forest, Schunke A65.
Elleanthus conifer (Reichb. f. & Warsc.) Reichb. f. aff. Fruit-
ing raceme.
Huanuco: Mito, about 2700 meters, patch at grassy summit, edge
of gulch, Macbride & Feather stone 1580.
Elleanthus conifer (Reichb. f. & Warsc.) Reichb. f. aff. Fruit-
ing raceme.
Huanuco: Yanano, about 1700 meters, in patches on steep, sunny,
sparsely shrubby, grassy slopes, Macbride 3834-
Sobralia sp. Fruit.
Junin: Pichis Trail, Dos de Mayo, 1700-1900 meters, epiphyte in
dense forest, buds greenish white, pink-tinged, Killip & Smith 25830.
Sobralia sp. Flowers agglutinated.
San Martin: San Roque, 1350-1500 meters, upland, Williams
7761.
Sobralia rosea Poepp. & Endl.
Habitat? between Bafios and Rio Verde, terrestrial, Pachano 231 .
Gomphichis sp.? No inflorescence.
Peru: Abragillo, (on loose ticket).
SCHWEINFURTH: ORCHIDS OF PERU 993
Altensteinia sp. Flowers old.
Junin: Mount La Juntay, near Huancayo, about 4700 meters, in
puna, terrestrial, Kittip & Smith 22074-
Pterichis sp. Flowers poor.
Huanuco: Mito, about 2700 meters, on grassy western slope,
Macbride & Featherstone 1794.
Pterichis triloba (Lindl.) Schltr. aff . Petals ciliate and lip poor.
Junin: Huacapistana, 1800-2400 meters, terrestrial in thickets
and open woods, Kittip & Smith 24123.
Stelis sp. Flowers agglutinated.
Habitat uncertain: On the western slopes of the equatorial Andes,
flowers dark violet, about 3000 meters, Mission Andre 4482.
Stelis parviflora (Ruiz & Pav.) Pers.
Habitat uncertain. Ex Herb. Ruiz & Pav.
Stelis sp. Flowers poor.
Ayacucho: Ccarrapa, between Huanta and Rio Apunmac, about
1500 meters, epiphyte in densely forested valley, perianth greenish
white, Kittip & Smith 23217.
Stelis sp Plant sterile or too advanced for determination.
Ayacucho: Ccarrapa, between Huanta and Rio Apurimac, 2500
meters, epiphyte on wooded hillside, KiUip & Smith 22294-
Stelis sp. Flowers rotten.
Cuzco: Ccochapampa, "altura de Ccochayoc," 101 • meters,
Bues s.n.
Stelis sp. Lip variable.
Huancavelica: Prov. of Tayacaja, Montepungo, 5 km. east of
Surcubamba, 3000 meters, abundant epiphyte on mossy tree trunks,
flowers varying from blood red to pale greenish white, Stork &
ton 10385.
Stelis sp. Flowers too old for diagnosis.
Huanuco: Chaglla, in patches in thickets, Macbride 3642.
Stelis sp No good flowers.
Junin: East of Quimiri Bridge, near La Merced, 800-1300 meters,
epiphyte in dense forest, Ktilip & Smith 23964.
994 FIELDIANA: BOTANY, VOLUME 30
Stelis sp. No good flowers.
Junin: Huacapistana, 1800-2400 meters, epiphyte in thickets and
open woods, perianth yellowish green, Killip & Smith 24497.
Stelis sp. No flowers.
Junin: Pichis Trail, Yapas, 1350-1600 meters, epiphyte in dense
forest, Killip & Smith 25524.
Stelis sp. No flowers.
Puno: Ollachea, Soukup 476.
Stelis sp. No good flowers.
Puno: Ollachea, Soukup 477.
Masdevallia sp. Fruit only.
Huanuco: Rio Huallaga Canon, below Rio Santo Domingo, in
montana, about 1200 meters, Macbride 4263.
Masdevallia sp. No flowers.
Junin: Chanchamayo Valley, 1800 meters, Schunke 522.
Lepanthes sp. Aff. L. blepharistes Reichb. f.
Junin: Carpapata, above Huacapistana, 2700-3200 meters, epi-
phyte at edge of forest, segments pink, Killip & Smith 24444.
Lepanthes sp. Fruit only, allied to L. monoptera Lindl.
Junin: Carpapata, above Huacapistana, 2700-3200 meters, epi-
phyte at edge of forest, Killip & Smith 30696.
Pleurothallis sp. Aff. P. amygdalodora Kranzl. No flowers.
"Hohe von Liere," 2400 meters, Bues 1734.
Pleurothallis sp. Aff. P. velaticaulis Reichb. f. Buds only.
Apurimac: Prov. of Andahuaylas, Pincos, 2700 meters, on clay
bank, "rainy green shrubland," Stork & Horton 10664.
Pleurothallis sp. Aff. P. cordata (Ruiz & Pav.) Lindl. Flowers
poor.
Ayacucho: Aina, between Huanta and Rio Apurimac, 750-1000
meters, epiphyte in dense forest, Killip & Smith 22778.
Pleurothallis sp. Aff. P. xanthochlora Reichb. f. No flowers.
Ayacucho: Aina, between Huanta and Rio Apurimac, 750-1000
meters, on base of tree in open woods, Killip & Smith 23166.
SCHWEINFURTH: ORCHIDS OF PERU 995
Pleurothallis sp. Aff. P. velaticaulis Reichb. f. No flowers.
Ayacucho: Aina, between Huanta and Rio Apurimac, at base
of tree in open woods, Kittip & Smith 23167.
Pleurothallis sp. Aff. P. cedralensis Ames. No flowers.
Ayacucho: Aina, between Huanta and Rio Apurimac, 750-1000
meters, epiphyte in open woods, Killip & Smith 23160.
Pleurothallis sp. Aff. P. serripetala Kranzl. Flower passe".
Cuzco: Ollantaytambo, about 3000 meters, on rocks, Cook &
Gilbert 309.
Pleurothallis sp. Aff. P. sicaria Lindl.
Cuzco: Hda. Ccochayoc, 1000 meters, Bues s.n.
Pleurothallis sp. Aff. P. cordata (Ruiz & Pav.) Lindl. No
flowers.
Cuzco: Ccochapampa, Altura de Ccochayoc, 1000 meters, Bues s.n.
Pleurothallis sp. Aft. P. Reymondii (Karst.) Eeichb. f . Flowers
poor. . , ,
Cuzco: Prov. of Convencion, Sahuayaco, 1600 meters, epiphyte,
Vargas 6301.
Pleurothallis sp. Aff. P. macrorhiza Lindl. Fruit.
Cuzco- Prov. of Paucartambo, between Acanacu and Pillahuata,
2800 meters in open woods of "ceja de la montana," eprphyte,
West 1061).
Pleiirothallis sp. Aff. P. Vargasii C. Schweinf. Buds only.
Cuzco- Prov. of Convenci6n, Hda. Potrero, Sapan-Sachayocc,
2200 meters forest epiphyte, "flowers white," Vargas 2556.
PleurothalHs sp. Aff. P. frutex Schltr. No flowers.
Cuzco: Alturas de Ccochayoc, "Ccochapampa," 1000 meters,
Bues s.n.
Pleurothallis sp. Aff. P. nigrohirsuta Kranzl.
Cuzco- Prov of Convencion, Hda. Potrero, Sapan-Sachayocc,
2200 meters epiphyte in dense forest, flowers pale yellow, Vargas
2539.
996 FIELDIANA: BOTANY, VOLUME 30
Pleurothallis sp. Aff. P. genychila Schltr. Plant imperfect.
Huanuco: Muna, on trail to Tambo de Vaca, about 2400 meters,
Macbride 4323.
Pleurothallis sp. Aff. P. Tonduzii Schltr. Flower loose in
pocket.
Huanuco: Between Huanuco and Pampayacu, Kanehira 319.
Pleurothallis sp. No flowers.
Junin : Chanchamayo Valley, 1800 meters, Schunke 541 .
Pleurothallis sp. Aff. P. crocodiliceps Reichb. f. No flowers.
Junin: Schunke Hacienda, above San Ramon, 1400-1700 meters,
epiphyte in dense forest, "perianth greenish yellow," Killip & Smith
24852.
Pleurothallis sp. Aff. P. muricata Schltr. Flower poor.
Junin : Schunke Hacienda, above San Ramon, 1300-1700 meters,
in dense forest, Schunke A71.
Pleurothallis sp. Aff. P. ciliata Knowl. & Westc. Flowers poor.
Junin: Pichis Trail, Yapas, 1350-1600 meters, epiphyte in dense
forest, Killip & Smith 25466.
Pleurothallis sp. (probably). No flowers.
Junin: East of Quimiri Bridge, near La Merced, 800-1300 meters,
epiphyte in dense forest, Killip & Smith 23963.
Pleurothallis sp.? No flowers.
Junin: Pichis Trail, Yapas, 1350-1600 meters, epiphyte in dense
forest, Killip & Smith 25523.
Pleurothallis sp.? No flowers.
Junin : Schunke Hacienda, above San Ramon, 1400-1700 meters,
epiphyte in dense forest, Killip & Smith 24619.
Pleurothallis sp. Aff. P. tridentata Kl. No flower.
Junin: Pichis Trail, San Nicolas, about 1100 meters, epiphyte in
dense forest, Killip & Smith 25977.
Pleurothallis sp. Aff. P. brevis Schltr. Flowers passe".
Junin: Chanchamayo Valley, 1800 meters, Schunke 1198.
SCHWEINFURTH: ORCHIDS OF PERU 997
Pleurothallis sp. Aff. P. rigidifolia Reichb. f. Flower poor.
Junin: Schunke Hacienda, above San Ramon, 1300-1700 meters,
in dense forest, Schunke A53.
Pleurothallis sp. Aff. P. tridentata Kl. Material inadequate.
Junin: Rio Pinedo, north of La Merced, 700-900 meters, epiphyte,
"perianth greenish white, pink- tinged," Kittip & Smith 23636.
Pleurothallis sp. Aff. P. juninensis Schltr. No flower.
Junin : Schunke Hacienda, above San Ramon, 1400-1700 meters,
epiphyte in dense forest, Killip & Smith 24586.
Pleurothallis sp. Aff. P. juninensis Schltr. No flower.
Junin: Pichis Trail, Yapas, 1350-1600 meters, epiphyte in dense
forest, "perianth reddish purple," Killip & Smith 25542.
Pleurothallis sp. Aff. P. crocodiliceps Reichb. f. Flower im-
perfect.
Junin: Pichis Trail, Dos de Mayo, 1700-1900 meters, epiphyte in
dense forest, Killip & Smith 25828.
Pleurothallis sp. Buds only.
Loreto: Puerto Arturo, lower Rio Huallaga, below Yurimaguas,
about 135 meters, "perianth rose-red," epiphyte in dense forest,
Killip & Smith 27890.
Pleurothallis sp.? No flowers.
Loreto: Alto Rio Itaya, San Antonio, 145 meters, on tree trunk,
Williams 3452.
Pleurothallis sp. No flowers.
San Martin: San Roque, 1350-1500 meters, "on trunk of shrub,"
Williams 7436.
Pleurothallis sp. Mixed collection.
San Martin: San Roque, 1350-1500 meters, on tree trunk in
forest, Williams 7737.
Octomeria sp. Aff. 0. tennis Schltr. Whole plant imperfect.
Junin: Chanchamayo Valley, 1600 meters, Schunke 1722.
Malaxis sp. Aff. M. Parthonii Morr. Fruit.
Huanuco: Cani, 7 miles northeast of Mito, about 2600 meters,
"in deep humus of dense stream-wood," Macbride 3392.
998 FIELDIANA: BOTANY, VOLUME 30
Malaxis sp. Aff. M. termensis (Kranzl.) C. Schweinf. Fruit.
Huanuco: Mito, about 2700 meters, on northeastern slope, "in
deep grasses," Macbride & Featherstone 1497.
Liparis sp. Aff. L. ramosa Poepp. & Endl. No good flowers.
Junin: La Merced, Hacienda Schunke, about 1200 meters, on
floor of "montana," Macbride 5643.
Epidendrum sp. Flowers agglutinated.
Amazonian slope of the Andes, 3000-3300 meters, Ehle 1927.
Epidendrum sp. Aff. E. nephroglossum Schltr. Flower poor.
Apurimac: Prov. of Andahuaylas, west of Pincos, 2600 meters,
epiphyte in forest, flower dull waxy yellow, Stork & Horton 10706.
Epidendrum sp. Aff. E. nocturnum Jacq. Flowers wilted.
Ayacucho: Aina, between Huanta and Rio Apurimac, 750-1000
meters, terrestrial in open woods, Killip & Smith 23169.
Epidendrum sp. No flowers.
Ayacucho: Ccarrapa, between Huanta and Rio Apurimac, about
1500 meters, epiphyte in densely forested valley, "segments grass-
green," Killip & Smith 23220.
Epidendrum sp. No good flowers.
Cuzco: Valle de San Miguel, slopes of Machu-Picchu, 2400
meters, Hen era 3188.
Epidendrum sp. Aff. E. syringothyrsis Reichb. f . ex Hook, f .
Huancavelica: Prov. of Tayacaja, Ampurco woods, on trail be-
tween Salcabamba and Surcubamba, 2900 meters, epiphyte on
mossy trunk, sepals and petals dark red, lip pink to white, Stork &
Horton 10427.
Epidendrum sp. Aff. E. Funkii Reichb. f . Plant poor.
Huanuco: Between Huanuco and Pampayacu, Kanehira 337.
Epidendrum sp. Aff. E. Schlimii Reichb. f. var. gracillimum
(Reichb. f. & Warsc.) C. Schweinf.
Huanuco: Between Huanuco and Pampayacu, Kanehira 26.
Epidendrum sp. Aff. E. Haenkeanum Presl. Small flowers.
Huanuco: Between Huanuco and Pampayacu, Kanehira 338.
SCHWEINFURTH: ORCHIDS OF PERU 999
Epidendrum sp. Old flower on fruit.
Huanuco: Chaglla, about 2700 meters, epiphyte in patches, Mac-
bride 3636.
Epidendrum sp. Old flower on fruit.
Huanuco: Huacachi, near Muna, about 2000 meters, pendent
epiphyte, Macbride 1+179 .
Epidendrum (Section Amphiglottium) sp. Flowers only.
Huanuco: Mito, about 2700 meters, on very steep, grassy,
sparsely shrubby, northeastern slope, flower dull brick-red with
yellow center, Macbride & Feather stone 1481.
Epidendrum sp. Flowers passe*.
Huanuco: Vilcabamba, Hacienda on Rio Chinchao, about 1800
meters, epiphyte, Macbride 5013.
Epidendrum sp. Aff. E. inamoenum Kranzl. Flowers old.
Huanuco: Yanahuanca, about 3000 meters, on steep, densely
shrubby bank, Macbride & Feather stone 1188.
Epidendrum sp. Flowers passe".
Huanuco: Muna, about 2100 meters, epiphyte, Macbride 4044-
Epidendrum sp. Fruit.
Huanuco: Muna, about 2100 meters, epiphyte, Macbride 4043-
Epidendrum sp. Flowers poor.
Huanuco: Between Huanuco and Pampayacu, Kanehira 313.
Epidendrum sp. Flowers poor.
Huanuco: Between Huanuco and Pampayacu, Kanehira 309.
Epidendrum sp. Aff. E. polystachyum HBK.
Junin: Chanchamayo Valley, Schunke s.n.
Epidendrum sp. Aff. E. Schlimii Reichb. f. var. gracillimum
(Reichb. f. & Warsc.) C. Schweinf. No flowers.
Junin: Huacapistana, 1800-2400 meters, epiphyte on open hill-
side, Killip & Smith 24225.
Epidendrum sp. E. inamoenum Kranzl.? Fruit.
Junin: Carpapata, above Huacapistana, 2700-3200 meters, ter-
restrial at edge of forest, Killip & Smith 24418.
1000 FIELDIANA: BOTANY, VOLUME 30
Epidendrum sp. Fruit.
Junin: Pichis Trail, Yapas, 1350-1600 meters, epiphyte in dense
forest, Killip & Smith 25585.
Epidendrum sp. Aff. E. paniculatum Ruiz & Pav. Fruit.
Junin: Rio Paucartambo Valley, near Perene Bridge, 700 meters,
epiphyte in dense forest, Killip & Smith 25338.
Epidendrum sp. Aff. E. crassilabium Poepp. & Endl. Fruit.
Junin: Huacapistana, 1800-2400 meters, in thickets and open
woods, Killip & Smith 24157.
Epidendrum sp. Aff. E. blepharichilum Kranzl. Fruit.
Junin: Carpapata, above Huacapistana, 2700-3200 meters, ter-
restrial in forest, Killip & Smith 24495.
Epidendrum sp. E. anceps Jacq. (probably). No flowers.
Loreto: Puerto Arturo, lower Rio Huallaga below Yurimaguas,
about 135 meters, epiphyte in dense forest, "segments greenish
yellow," Killip & Smith 27746.
Epidendrum sp. Flowers are E. fruticulus Schltr. (probably) ;
plant apparently a Maxillaria.
Puno: Prov. of Sandia, near Limbani, 3200-3450 meters, on
very wet, shaded mountain-side in sphagnum, "flowers red and
orange-yellow," Metcalf 30540.
Epidendrum sp. Aff. E. scabrum Ruiz & Pav. Flower old.
Puno: Prov. of Sandia, near Limbani, 3200-3450 meters, in dry
sphagnum on rocks, in fog belt, "perianth segments green-brown,"
Metcalf 30534-
Epidendrum sp. Flower imperfect.
San Martin: San Roque, 1350-1500 meters, Williams 7313.
Scaphyglottis sp. Aff. S. Huebneri Schltr. No flowers.
Ayacucho: Rio Apurimac Valley, near Kimpitiriki, 400 meters,
epiphyte at edge of dense forest along beach, Killip & Smith 22977.
Scaphyglottis sp. No good flowers.
Junin: La Merced, Hacienda Schunke, about 1200 meters, on
fallen log, "fls. greenish white," Macbride 5598.
SCHWEINFURTH: ORCHIDS OF PERU 1001
Scaphyglottis sp. Flower loose in pocket; perhaps a mixture.
Junin: Chanchamayo Valley, 1500 meters, Schunke s.n. Herb.
Chicago Nat. Hist. Mus. 571663.
Scaphyglottis sp. Flower loose in pocket. Same as last.
Junin: Chanchamayo Valley, 1500 meters, Schunke 536.
Polystachya sp. Fruit.
Junin : Rio Paucartambo Valley, near Perene Bridge, 700 meters,
epiphyte in dense forest, Killip & Smith 25361.
Polystachya sp. Buds only.
Loreto: Rio Huallaga Canyon, below Rio Santo Domingo, about
1200 meters, "dry woods, fls. green," Macbride 4253.
Galeandra sp. No flowers.
Loreto: Balsapuerto (lower Rio Huallaga basin), 150-350 meters,
epiphyte in dense forest, "flowers yellow," Killip & Smith 28402.
Govenia sp. Aff. G. tingens Poepp. & Endl. Fruit.
Huanuco: Muna, about 2100 meters, in dry rocky woods, Mac-
bride 4042.
Catasetum sp. (probably). Buds only.
Junin: Colonia Perene, about 680 meters, epiphyte in dense for-
est, "young perianth segments purple," Killip & Smith 24951.
Catasetum sp. Raceme of female flowers.
Junin: Chanchamayo Valley, 1000 meters, Schunke 1653.
Catasetum sp. Abnormal growth and flowers.
Loreto: Vicinity of Iquitos, epiphyte in dense forest, "fls. migno-
nette, with small red-brown spots," Klug 10033.
Lycaste sp. Fruit.
Huanuco: Mito, about 2700 meters, "on soil ledge of shrubby
southwestern slope," Macbride & Feather stone 1480.
Lycaste sp. (probably). Immature bud only.
Junin: East of Quimiri Bridge, near La Merced, 800-1300 meters,
epiphyte in dense forest, "perianth green, pink-tinged within, lip and
column white," Killip & Smith 23840.
1002 FIELDIANA: BOTANY, VOLUME 30
Batemania sp. (probably). No good flowers.
Junin: Colonia Perene, about 680 meters, epiphyte in dense for-
est, "perianth greenish white," Killip & Smith 24960.
Zygopetalum sp.? Old fruit.
Junin: Pichis Trail, Enenas, 1700 meters, terrestrial in open
sphagnum swamp, Killip & Smith 25696.
Zygopetalum (Section Warscewiczella) sp. Fruit.
Loreto: Balsapuerto (lower Rio Huallaga basin), 150-350 meters,
epiphyte in dense forest, Killip & Smith 28624-
Zygopetalum (Section Warscewiczella) sp. Fruit.
San Martin: San Roque, 1350-1500 meters, on trunk of petrified
tree, Williams 7345.
Maxillaria sp. (probably). No flowers.
Ayacucho: Aina, between Huanta and Rio Apurimac, 750-1000
meters, epiphyte in open woods, "segments light green, column red-
dotted," Killip & Smith 23158.
Maxillaria sp. Aff. M. xantholeuca Schltr. var. peruviana C.
Schweinf. Flowers imperfect.
Cuzco: Prov. of Convencion, Sahuayaco, 1800 meters, epiphyte,
"flowers yellow," Vargas 6296.
Maxillaria sp. No flowers.
Huanuco: Pampayacu, Hacienda at mouth of Rio Chinchao,
about 1070 meters, epiphyte in dense woods, "fls. bright yellow,"
Macbride 5107.
Maxillaria sp. Aff. M. purpurea (Spreng.) Ames & Correll.
No good flowers.
Huanuco: Pozuzo, about 600 meters, epiphyte, Macbride 4579.
Maxillaria sp. Aff. M. striata Rolfe. No pseudobulb present.
San Martin: Near Tingo Maria, 625-1100 meters, "flowers pure
yellow with orange lip," Allard 21986.
Scelochilus sp. Aff. S. saccatus (Poepp. & Endl.) Reichb. f.
Flower imperfect.
Cuzco: Prov. of Urubamba, "Huinahuina," 2550 meters, epi-
phyte, "flower pale yellow," Vargas 4126.
SCHWEINFURTH: ORCHIDS OF PERU 1003
Odontoglossum sp. (probably). Flowers old.
Ayacucho: Ccarrapa, between Huanta and Rio Apurimac, about
1500 meters, epiphyte in densely forested valley, Killip & Smith
22405.
Brassia sp. Flower imperfect.
Huanuco: Confluence of Huallaga and Monzon Rivers, near Tingo
Maria, 700 meters, epiphyte on tall tree, "sepals greenish yellow
mottled with dark purple, lip yellow mottled with dark purple,"
Stork & Horton 9499.
Oncidium sp. 0. aureum Lindl.? Old fruit.
Huanuco: Muiia, about 2100 meters, epiphyte, Macbride 3955.
Lockhartia sp. Aff . L. hologlossa Schltr. Fruit.
Junin: Huacapistana, about 1800 meters, epiphyte in densely
forested valley, Killip & Smith 24296.
Ornithocephalus sp. No flowers.
Cuzco: Cochapampa, 1000 meters, "banda derecha del Uru-
bamba," Bues 2.
Ornithocephalus sp. Aff. 0. pygmaeus Reichb. f. & Warm.
Material scanty.
Cuzco : Valle de Santa Ana, Herrera 3046.
Ornithocephalus sp. Immature buds only.
Junin: Along Rio Perene, near "Hacienda 3," Colonia Perene,
about 600 meters, epiphyte in forest, "perianth segments white,"
Killip & Smith 25179.
Ornithocephalus sp. Buds only.
Junin: Colonia Perene, about 680 meters, epiphyte in dense forest,
Killip & Smith 25094*
Ornithocephalus sp. No flowers.
Junin: San Ramon, 900-1300 meters, epiphyte in dry woods,
Killip & Smith 24759.
Notylia sp. Buds only.
Junin: Along Rio Perene, near "Hacienda 3," Colonia Perene,
about 600 meters, epiphyte in forest, Killip & Smith 251 77.
1004 FIELDIANA: BOTANY, VOLUME 30
Notylia sp. N. yauaperyensis Barb. Rodr., fide Mansf. Photo-
graph only.
Loreto: Upper Rio Maranon, at mouth of Rio Santiago, 160 me-
ters, in rain-forest, "flower yellow with a white lip," Tessmann 4309.
Notylia sp. Buds only.
San Martin: Tarapoto, 750 meters, epiphyte on shrub, Williams
6148.
Trichoceros sp. Aff. T. parviflorus HBK. No flowers.
Junin: Huacapistana, 1800-2400 meters, terrestrial in thickets
and open woods, "sepals yellow-green mottled with pink, column
and lip deep pink marked with yellow," Killip & Smith 24124-
Dichaea sp. Aff. D. muricata (Sw.) Lindl. var. maculata (Poepp.
& Endl.) C. Schweinf. No flowers.
Ayacucho: Aina, between Huanta and Rio Apurimac, 750-1000
meters, epiphyte in open woods, "perianth greenish white, purple-
dotted," Killip & Smith 22592.
Dichaea sp. No flowers.
Junin: East of Quimiri Bridge, near La Merced, 800-1300 meters,
epiphyte in dense forest, "fruit green," Killip & Smith 23828.
Dichaea sp. Aff. D. calyculata Poepp. & Endl. Old fruit.
Junin: Schunke Hacienda, above San Ramon, 1400-1700 meters,
epiphyte in dense forest, Killip & Smith 24861 .
Dichaea sp. Flower poor.
Junin: La Merced, Hacienda Schunke, about 1200 meters, epi-
phyte in "montana," Macbride 5766.
Dichaea sp. Old fruit.
Junin: Pichis Trail, San Nicolas, about 1100 meters, epiphyte in
dense forest, Killip & Smith 25979, 25980.
Dichaea sp. Aff. D. campanulata C. Schweinf. Sterile.
Junin : San Luis de Shuaro, Soukup 2535.
Dichaea sp. Sterile.
Junin: Pichis Trail, San Nicolas, about 1100 meters, epiphyte in
dense forest, "buds white," Killip & Smith 25991.
SCHWEINFURTH: ORCHIDS OF PERU 1005
Dichaea sp. Aff. D. campanulata C. Schweinf. Old fruit.
Junin: Wooded banks of Rio Pichis, between Puerto Bermudez
and Cahuapanas, 340-375 meters, epiphyte, Killip & Smith 26691.
Dichaea sp. Aff. D. campanulata C. Schweinf. Sterile.
Loreto: Lower Rio Huallaga, San Ramon, Yurimaguas, 155-210
meters, Williams 4565.
Dichaea sp. Flower poor.
San Martin: Juan Jui, Alto Rio Huallaga, about 400 meters, epi-
phyte in forest, "flower white and violet," Klug 3796.
Dichaea sp. Aff. D. muricata (Sw.) Lindl. var. maculata (Poepp.
& Endl.) C. Schweinf. Sterile.
San Martin: Lamas, near Tarapoto, 840 meters, epiphyte on
shrub, Williams 6402.
Dichaea sp. Aff. D. muricata (Sw.) Lindl. var. maculata (Poepp.
& Endl.) C. Schweinf. Sterile.
San Martin: San Roque, 1350-1500 meters, epiphyte on forest
tree, Williams 7709.
Pachyphyllum sp. No flower seen.
Peru: Habitat not recorded, Mathews 1862.
Pachyphyllum sp. Buds only.
Cuzco: Bues s.n. (Herrera 2153).
Pachyphyllum sp. Flowers poor.
Cuzco: Prov. of Urubamba, Puyupata-Tuncapata, 3200 meters,
epiphyte, Vargas 2929.
Campylocentrum sp. Aff. C. micranthum (Lindl.) Rolfe.
Flowers poor.
Junin: Schunke Hacienda, above San Ramon, 1300-1700 meters,
in dense forest, Schunke A 52, A66.
Campylocentrum sp. Aff. C. micranthum (Lindl.) Rolfe.
No good flowers.
San Martin: San Roque, 1350-1500 meters, epiphytic, Williams
7511.
INDEX
Synonyms in italics; main references to names of genera and species in bold-
face numerals.
Aa92
brevis 102
calceata 94
chiogena 94
erosa 96
inaequalis 97
Lechleri 100, 101
Mandonii 98
Matthewsti 100
nubigena 101
paleacea 101
paludosa 102
pumilio 102
Eosei 103
Weberbaueri 103
Weddefo'ana 104
AcacaMt's 653
coerwZea 653
Acraea triloba 109, 110
Acrom'a phalangifera 325, 326
Adeneleuterophora graminifolia 62
Aganisia 653
coerulea 653, 654
fimbriata 653, 654
graminea 651
ionoptera 652
Altensteinia 92
boliviensis 96, 97
calceata 94
chiogena 94
elliptica 94
erosa 96
fimbriata 96, 97
inaequalis 97
longispicata 97
Mandonii 98
marginata 98, 100
Matthewsii 100, 101
nubigena 101
paleacea 101
paludosa 102
paludosa 103
pilifera 91, 92
pterostyloides 98
Rosei 103
sceptrum 96
Weberbaueri 103
Weddeliana 104
Amblostoma 384
cernua 385, 387
385, 387
gracile var. robustum 385
holochilum 472, 473
tridactylum 385
Angraecum fasciola 985
maculatum 571
micranthum 986
Anguloa 649
eburnea 649, 650
f squalida 602
uniflora 649
virginalis 649, 650
Aspe0re?w'a scirpoidea 364
Barbosella cucullata 330
longiflora 237
rhynchantha 330
Baskervilla 117
assurgens 117
Batemania peruviana 662, 663
Batemannia 662
Colleyi 662, 663
Wolteriana 663
Beloglottis costaricensis 132
Bifrenaria 638
longicornis 638, 639
saftwZosa 638, 639
saxicola 639
Bletia 562
capitata 53, 55
catenulata 562, 564
ensiformis 431, 565
Mandonii 565
nodosa 538
parviflora 566
repanda 566
Rodriguesii 562, 565
sanguinea 562, 564
Sherrattiana 562, 565
uniflora 566
penosa 538
Wageneri 565
Watsoni 562
Bonatea pauciflora 32
Brachionidium 360
serratum 360
Brachystele 128
Lechleri 140
148
1007
1008
FIELDIANA: BOTANY, VOLUME 30
Brassavola 537
grandi 'flora 538
nodosa 538
ovaliformis 538
venosa 538
Brassia 834
arcuigera 835
bicolor 836
bidens 836
caudata 837
eauliformis 837
cryptophthalma 840
elegantula 840
Filomenoi 841
Forgetiana 841
hexodonta 846, 847
juninensis 842
Koehlerorum 844
Lanceana var. viridiflora 836
Lawrenceana var. longissima 844
Lewisii 837
longissima 844
maculata 841
peru viana 845
signata 845
thyrsodes 846
villosa 846, 847
Broughtonia grandi flora 700
maculata 714
Buchtienia 127
boliviensis 128
Buesiella 569
pusilla 569
Bulbophyllum 567
incarum 567
meridense 568
Weberbauerianum 568
Burlingtonia Batemanii 771
rubescens 771
Camaridium 668
affine 685
aureum 680
var. giganteum 681
cassapense 712
densifolium 692
exaltatum 695
graminifolium 699
grandiflorum 710
longibracteatum 709, 710
luteo-rubrum 712
ochroleucum 684, 685
purpureum 724, 725
vandiforme 740
Campylocentrum 984
fasciola 985
Lankesteri 985
loretoense 985
micranthum 985
minutum 986
multiflorum 985
Sullivanii 985
Ulaei var. peruvianum 988
Catasetum 583
barbatum 584
callosum 585
var. carunculatum 584, 585
var. grandiflorum 585
carunculatum 584, 585
Charlesworthii 586
chloranthum 588
coniforme 585
cruciatum 590
fuliginosum 586
incur vum 590
microglossum 586
naso var. Charlesworthii 586
planiceps 586, 588
pusillum 588
recurvatum 588
Russellianum 591
saccatum var. incurvum 590
stupendum 590
tenebrosum 591
violascens 591
Cattleya 533
dichotoma 71
epidendroides 533
flavida 533
luteola 533, 534
var. Roezlii 533, 534
Malouana 534
maxima 534, 535
Meyeri 533
rex 535
Schomburgkii 535
sulphur ea 533
superba 535, 536
violacea 535
Centroglossa peruviana 760
Centropetalum 981
distichum 982
Hartwegii 983
ionantherum 983
nigro-signatum 984
Warscewiczii 982
Chaenanthe 766
micrantha 766
Chloraea 36
densipapillosa 36, 37
multilineolata 37
peruviana 38
reticulata 38
Chondrorhyncha 664
fimbriata 665
lojae 665
pusilla 666
Chysis 566
Bruennowiana 566
Cleiates 39
rosea 40
Clinhymenia pallidi flora 941
Coccineorchis 128
corymbosa 131
INDEX
1009
Cochlioda 793, 833
Beyrodtiana 794
densiflora 794
Noezliana 795
rosea 795
sanguinea 833
vulcanica 796
Weberbaueriana 799
Coelogyne triptera 498, 500
Coilostylis obtusifolia 466
Comparettia 780
falcata 781
peruviana 781
saccata 780
Coryanthes 619
alborosea 620
bicalcarata 620
leucocorys 622
macrantha 623
macrocorys 623
maculata 624
var. punctata 624
punctata 624
trifoliata 625
Cranichis 110
calva 111
var. Vargasii 112
ciliata 112
foliosa 115
'Koehleri 115, 116
longipetiolata 113
longiscapa 80, 81
luteola 558, 560
Mandonii 112, 113
multiflora 112, 115
muscosa 115
pilosa 116, 117
pycnantha 115, 116
stachyodes 104, 105
Viereckii 117
Wageneri 116, 117
Crocodeilanthe xiphizusa 290
Cryptarrhena 941
acrensis 941
Kegelii 942
lunata 941, 942
pallidiflora 942
unguiculata 942
Cryptocentrum 757
inaequisepalum 757
minus 760
peruvianum 760
pseudobulbosum 760
Cryptophoranthus 223
Dayanus 223
oblongifolius 224
Cyathoglottis Candida 70
crocea 71
Cyclopogon 128
alpestris 136
densiflorus 136, 138
elatus 136
inaequilaterus 139
micrantha 141
minutiflorus 136
moyobambae 136, 138
olivaceus 142
ovalifolium 134, 136
peruvianus 136
plantagineus 136
Rimbachii 145
stictophyllus 136
Cycnoches 592
Cooperi 592
Haagii 593
muscifera 605, 606
peruviana 593
Cymbidium graminoides 965
nodosum 538
ochroleucum 685
proliferum 544
pusillum 899
subulatum 793
vestitum 724
vexilliferum 380
triolaceum 535, 536
Cypripedium Boissierianum 17
caridnum 18
caudalum 19
Czerwiakowianum 19
grandiflorum 17
Hartwegii 21
Pearcei 18
Cyrtochilum aemulum 907
cocciferum 866
cordatum 866
cryptocopis 869
deltoideum 869, 870
depauperatum 811
diceratum 870
falcipetalum 872
flexuosum 865, 866
gargantua 875
incarum 881
ionopterum 882
Kienastianum 882
Leopoldianum 883
longifolium 819
longipes 820
macranthum 886
megalophium 820
mdanthus 821
rafiwfe'gerwm 889
minaa; 889
myanthum 822
myrianthum 823
mystacinum 823, 824
Pavonii 872
retusum 826
serratum 905
superbiens 907
trilingue 909
910
1010
FIELDIANA: BOTANY, VOLUME 30
undulatum 911, 912
ventilabrum 911
volubile 912, 913
Weberbauerianum 807, 913
xanthodon 914
xanthornis 914
zebrinum 915
Cyrtoglottis 750, 752
gracilipes 752, 753
peruviana 753
Cyrtopera longifolia 572
scabrilinguis 634, 635
Woodfordii 572
Cyrtopodium 574
Buchtienii 575
eburneum 650, 651
punctatum 575
Woodfordii 572, 574
Dendrobium acuminatum 270
alatum 676
bicolor 682
carnosww 634, 635
grandiflorum 700
hastatum 702
ligulatum 708
longifolium 572, 574
longipetalum 713
macwtottm 714
paniculatum 720
platypetalum 723
plicatum 720
polypetalum 723
proliferum 724
pusillum 327, 328
ramosum 725
ruscifolium 335
squalens 634, 636
tricolor 735
triphyllum 738
undatiflorum 739
undulatum 636
variegatum 740
Diadenium 766
micranthum 766
Dichaea 962
ancoraelabia 963
arbuscula 730
calyculata 963
campanulata 965
graminoides 963, 965
humilis 966
laxa 966
maculata 968
Morrisii 968
muricata yar. maculata 968
panamensis 969
tenuis 970
Dichaeopsis panamensis 969
Dicrypta bicolor 682
densifolia 692
longifolia 712
vt'Mosa 742, 744
Didactyle meridensis 568
Dienia calycina 368, 369
Diothonea 380, 447
exasperata 381
gratissima 383
heterothonaea 381
imbricata 387, 388
lloensis 383
nutans 383
oppositifolia 387
suavis 384
Dipteranthus peruvianus 938
Elleanthus 47
amethystinus 50
aurantiacus 50, 53, 58
aureus 51
bambusaceus 52
Bonplandii 52
bractescens 53
cajamarcae 50, 51
capitatus 53, 55, 56
Carolii 55
casapensis 55
cephalophorus 55
conchochilus 56
conifer 56
discolor 59
ensatus 57
flavescens 58
furfuraceus 58
furfurascens 58
galipanensis 50, 51
gastroglottis 59
Hallii 59
hymenophorus 59, 60, 66
igneus 60
kermesinus 60
Koehleri 61
laxifoliatus 61
linifolius 62
longibracteatus 63
lupulinus 63
Myrosmatis 64
oliganthus 64
pallidiflavus 50, 51
pallidiflorus 50
porphyrocephalus 65
rhodolepis 65
robustus 65, 66
Ruizii 66
Spruceanus 66
strobilifer 57, 67
virgatus 68
Weberbauerianus 56, 57
xanthocomos 68
Encyclia macrostachya 410, 411
nana 555, 558
polystachya 555, 558
Epidendrum 380, 384, 390
INDEX
1011
acuminatum 405, 425
adenoglossum 405
aemulum 442, 443
aggregatum 405
alternans 441, 442
amazonicum 479, 480
ampliracemum 406
anceps 406, 408, 469
anthoceros 425, 426
apaganum 435
aquaticoides 408
var. pusillum 486, 487
ardens 410
armeniacum 410, 411
aromaticum 524
arundinaceum 411
atacazoicum 488, 490
avicula 388, 389
bambusaceum 411
bambusiforme 412
Bangii 507
Baumannianum 456, 457
benignum 479
bifalce 489, 490
biflorum 412
birostratum 412
blepharichilum 413
var. ma jus 415
brachycladium 446, 447
/3 crassipes 446
brachyglossum 415
brachyphylluin 433, 435
breyicaule 415
brevicaule 511, 512
breviracemum 416
Bulkeleyi 460
caespitosum 498, 500
cajamarcae 448
calanthum 457, 458
caloglossum 489, 490
capitellatum 418
capricornu 418
cardiophyllum 510, 511
carnosiflorum 419
catillus 419, 421
caucae 457, 458
caudatum 837
chrysostomum 456, 457
ciliare 421
cinnabarinum 422
cochleatum 442, 443
cochlidium 422
cordatum 424
coriifolium 424
cornutum 405, 425, 426
coronatum 426
corymbosum 426
crassilabium 426, 427
crassipes 446, 447
crassum 427
criniferum 429
cristatum 429, 430
croceum 430
cuzcoense 433, 434
cylindraceum 430
cylindrostachys 431
cyperifolium 431
decipiens 456, 457
densifolium 432
dermatanthum 432
dichotomum 426, 433, 434, 525
difforme 435
discolor 482, 484, 511
elatum 437
ellipticum var. flavum 528, 529
emarginatum 437
Englerianum 488, 490
equitans 437
euspathum 437
exaltatum 439
excisum 440
var. grandiflorum 440
Feddeanum 440
ferrugineum 441
Filomenoi 457
fimbriatum 441
var. rhomboglossum 442
floribundum 488, 490
Forgetianum 442
fragrans 442, 443
fraternum 456, 457
Friderici-Guilielmi 443
frigidum 444
var. stenophyton 444
frons bovis 488, 490
frutex 445
fruticulum 445
fulgens 457, 458
Funkii 446, 447
fuscatum 406, 408
fuscum 454, 455
gastrochilum 447
ejminiflorum 448
iroudianum 448
gladiatum 449
glossoceras 449
gnomus 495, 496
Goodspeedianum 480
gracilicaule 450
gracillimum 513
gramineum 450
graminoides 965
grammatoglossum 451
grandiflorum 452
gratiosum 490
var. linearifolium 491
gravidum 485
guttatum 885
haematanthum 452
Haenkeanum 453
Harmsianum 454
Hartwegii 454, 455
Hemiscleria 383
Herreranum 455
1012
FIELDIANA: BOTANY, VOLUME 30
huacapistanae 443
huanucoense 456, 457
ibaguense 456, 457, 458
var. confluens 457
var. Schomburgkii 458
imatophyllum 458
inamoenum 459
var. robustum 460
incisum 460, 530
inconstans 433
integrilabium 442
ionodesme 488, 490
Islhmi 489, 490
Juergensenii 449
juninense 453, 454
laceratum 461
laetum 456, 457
laeve 488, 490
lanipes 461
latilabre 435
latisegmentum 463
laxum 530, 531
Lechleri 463
liguliferum 464
Lima 464, 466
Lindenii 433, 435, 525
lineare 466
lineare 553, 554
lineatum 442, 443
longicolle 484
longicrure 488, 490
longirepens 466
loxense 510, 511
luteolum 533
luteoroseum 466
Macbridei 467
macroceras 488, 490
macrocyphum 467
macrodonax 444
macrogastrium 468
macropodum 468
macrothyrsis 530, 531
maculatum 469
magnicallosum 469
mancum 471
Mathewsii 495, 496
maximum 534
megagastrium 471
melinoacron 425, 426
mesomicron 472
micranthum 472
micro-Cattleya 473
microphyllum 389, 390
microtos 474
var. grandiflorum 431
minutidentatum 474
minutiflorum 476
modestum 476, 478
monzonense 478
moyobambae 426, 479
muricatum 480
nanum 480
nephroglossum 481
nocturnum 482, 484
var. latifolium 482
var. minus 484
var. tridens 482
nodosum 538
nutans 484, 507, 508
oliganthum 484
oligophyllum 484
oncidioides
var. grayidum 485
oppositifolium 387, 388
orbiculatum 486
Ottonis 486
pachychilum 486, 487
pachygastrium 526
panchrysum 487
panicoides 488
paniculatum 488, 490, 491
(C) cuspidatum 488
var. linearifolium 490
(B) longicrure 488
var. unguiculatum 491
parviflorum 490, 491
parvilabre 491
patulipetalum 491
Pavonianum 425, 426
paytense 457
Penlandii 508, 509
peruvianum 412
Philippii 492
physodes 496
physopus 492
pilosum 492
platyoon 513, 514
pleurobotrys 493
polystachyum 493
Porpax 495, 496
pristes 457
proliferum 544
prostratum 496
pubiflorum 496, 497
pulcherrimum 422
punctatum 576
purum 497
pusillum 899, 901
pygmaeum 498, 500
radicans 456, 457
ramosum 500
raniferum 429, 430
raphidophorum 500
rectopedunculatum 501
reflexilobum 502, 503
refractoides 502, 503
var. humile 503
rhomboglossum 442
rhopalorhachis 383, 384
rigidiflorum 504
rigidum 505
Rondosianum 505
rostrigerum 507
Ruizianum 507
INDEX
1013
rupestre 508, 509
ruscifolium 335
saccharatum 426, 427
Sanderi 452
sarcostalix 509
saxicola 509
scabrum 510
var. parviflorum 510, 511
Schlechterianum 511
var. longirepens 466
Schlimii 512, 513
var. gracillimum 513
Schomburgkii 458
var. confluens 457
sclerocladium 513
Scutella 513, 514
serpens 514
serricardium 527
smaragdinum 514
Smithii 456, 457
soratae 515
sororium 456, 457
spathaceum 508
splendens 458
Spruceanum 482, 484
stenophyton 444
strictiforme 516
strobiliferum 516
subliberum 519
subpatens 479, 480
subreniforme 519
var. ramosum 521
subulatum 791, 793
superbum 535
syringaeflorum 488
tarmense 433, 434
tenellum 512, 513
tenuispathum 521
Tessmannii 523
Teuscherianum 469
thrombodes 524
tricarinatum 524
tricrure 524, 525
tridactylum 385, 387
tridens 482, 484
triflorum 525
tripterum 498
turialvae 488, 490
unifoliatum 525
utricularioides 768, 770
validum 429, 430
vandifolium 449
variegatum 426, 427
var. crassilabium 426
ventricosum 526
vernixium 526
vesicicaule 527
veslitum^ 724, 725
vincentinum 527
vinosum 419, 421
violaceum 535
viride 528
volubile 528
Weberbauerianum 528
Woytkowskianum 437
xanthinum 460, 468, 528, 529, 530
xylostachyum 529
xytriophorum 530
yatapuense 530, 531
Epistephium 44
amabile 45
amplexicaule 46
Duckei 46
datum 45, 46
macrophyllum 46
monanthum 47
Epithecia calyculata 963
graminoides 966
humilis 966
laxa 966
Morrisii 968
panamensis 969
Eriopsis 595
biloba 596
colombiana 596
Helenae 597
sceptrum 597, 598
Schomburgkii 596
Sprucei 597
Wercklei 596
Erythrpdes 150
arietina 152
bifalcis 152
clavigera 153
foliosa 153
lobatocalcar 154
major 154
marmorata 156
minor 156
multifoliata 157
mystacina 157
ovata 159
querceticola 159
repens 160
santensis 160
simplex 162
stenocentron 163
valida 163
Eulophia 572
alta 572
longifolia 572
Mackaiana 656, 658
maculata 571
Woodfordti 572
Eulophidium 571
maculatum 571
Euothonaea imbricata 387
oppositifolia 387
Evelyna 47
amethystina 50
aurantiaca 50, 51
aurea 51
Bonplandii 52
bractescens 53
1014
FIELDIANA: BOTANY, VOLUME 30
capitata 53
casapensis 55
cephalophora 55, 56
conifer a 56, 57
cynarocephala 53
discolor 59, 60
ensata 57
flavescens 58
furfuracea 58
graminifolia 62
tfaHz'i 59
hymenophora 59, 60
kermesina 60, 61
longibracteata 63
lupulina 63, 64
Myrosmatis 64
oligantha 64
rhodolepis 65
robttsta 65, 66
Ruizii 66
strobilifera 67
Fernandezia conferta 543
denticulate, 975
disticha 982
graminifolia 545
haemathodes 976
ionanthera 983
iaxa 966
pttZcfcra 982
punctata 678
subbiflora 981
Warscewiczz'f 982
Galeandra 560
Baueri var. piloso-columna 560
juncea 561
pubicentrum 561
Gomphichis 78
goodyeroides 79
Koehleri 79
var. minor 80
longiscapa 80
Macbridei 81
plantaginea 81, 83
plantaginea 83
plantaginifolia 83
viscosa 83
Gongora 612
bufonia 617
var. leucochila 617
615
var. vitellina 615
histrionica 615
Incarum 618
leucochila 615
longipes 614
macrantha 623
maculata 614, 617
var. bufonia 617
var. leucochila 617
var. tricolor 615
nigro punctata 617
quinquenervis 614, 615
quinquevulneris 615
retrorsa 615
Sanderiana 617
scaphephorus 618
Tracyana 618
tricolor 615
Govenia 579, 580
6arfcata 572, 574
tingens 579
utriculata 580
Gyrostachys costaricensis 132
Haenkeana 136
Mrto 139
inaequilatera 139
macrostachya 91
ovalifolia 136
peruviana 136
picto 131
vaginata 148
Habenaria 21
abortiens 23
autumnalis 24
avicula var. peruviana 24
caldensis 24
chloroceras 30, 31
corydophora 26
dentifera 26
dilatata 29
dolichosceras 34
fastor 34
floribunda 27
Guentheriana 27
hexaptera 27, 32
laxiflora 28
ligulata 28, 32
Lindenii 30
linifolia 29
maculosa 29
maxillaris 33
monorrhiza 29
obtusa 30
Paivaeana 30, 31
parvicalcarata 31
parvidens 31, 32
pauciflora 32
pumiloides 32
repens 33
rupestris 33
Sartor 34
setacea 29
setifera 32
speciosa 29
uncatiloba 34
Hemiscleria 380
nutons 383, 384
Hexadesmia 541
Reedii 550
Hexisea 387
bidentata var. imbricata 387
INDEX
1015
imbricata 387
oppositifolia 387
Hofmeistera eumicroscopica 923
Hofmeisterella 923
eumicroscopica 923
Hormidium pseudopygmaeum 498
tripterum 498
Houlletia 603
odoratissima 604
Sanderi 604
vittata 606
Humboldtia 163, 261
acutiflora 170, 171, 271
affinis 271
antennifera 274
arbuscula 274
Arc/w'rfonae 275
aspera 174
asperilinguis 275
owrea 275
bicornis 276
casapensis 279
Cassidis 281
caulescens 282
cauliflora 282
ckamaelepanthes 283
chamensis 283
ciliata 285
contorta 180
cordate 180, 287
declivis 292
decurrens 292
efcjfttsa 293
dinotherii 295
diptera 295
dura 297
emarpinata 223
fimbriata 301
flexuosa 302
/ons florum 304
geftda 305
gigantea 306
grandiflora 307
Gro&i/i 309
hemirhoda 309
lanceolate 199, 314
Lindenii 339
macrorhiza 318
Mathewsii 325
misera 320
oblonga 173, 205
oborata 322
parviflora 205
parmfolia 324
picto 309
plantaginea 242
Poeppigii 326
polystachya 206
pruinosa 327
purpurea 207, 209
pusilla 327
restrepicrides 328
revoluta 210
rigidifolia 332
rubens 333
ruberrima 335
ruscifolia 335
scabrata 336
scabrilinguis 338
secunda 339
semipellucida 293
Smithiana 341
spiralis 215, 344
stenopetala 344
swceosa 335
tentaculata 346
tricarinata 350
tridentata 351
trilineata 351
irttMa 352
undulata 353
velaticaulis 354
xanthochlora 357
lonopsis 768
utricularioides 768
Isochilus 553
brasiliensis 553, 554
graminifolius 699, 700
grandiflorum 710
linearis 553
linifolium 62
peruvianus 553, 554
prolifer 544
proliferum 544
Kefersteinia 664
Zo/ae 665
Koellensteinia 650
eburnea 650
graminea 651
ionoptera 652
peruviana 650, 651
Kraenzlinella rufescens 320, 321
Laelia 536
moyobambae 536
Weberbaueriana 537
Lanium 388
avicula 388
microphyllum 389
peruvianum 389, 390
Lepanthes 248
alticola 249
cassidea 249
caudatisepala 250
juninensis 250
Koehleri 252
longipedicellata 252
mesochlora 254
minutipetala 255
pubicaulis 255
pumila 258
secunda 358, 360
1016
FIELDIANA: BOTANY, VOLUME 30
tracheia 258, 260
trachysepala 260
tricarinata 350
Lepanthopsis 358
floripecten 358
Leptorchis crispifolia 374
Leucohyle Warscewiczii 793
Limodorum altum 572, 574
maculatum 571
Lindleyella 638
saxicola 639
Liparis 373
crispifolia 374
elata 374
elegantula 376
laticuneata 377
Millet 376
neuroglossa 379
ramosa 377
var. Rusbyi 379
retusa 379
Rusbyi 379
vexillifera 380
Lockhartia 921
amoena 922
costaricensis 922
holpglossa 922
unicornis 922
Lueddemannia 594
triloba 594
Vyvereana 595
Lycaste 639
acuminata 676
barbifrons 641
Barringtoniae var. grandiflora 642
cinnabarina 641
Cobbiana 642
costata 642
Dyeriana 642
Filomenoi 646, 648
fimbriata 642
var. peruviana 643
gigantea 643, 648
grandiflora 700
lanipes 642, 643
lata 644
linguella 645
locusta 645
Sanders var. 645
longisepala 646
macrophylla 646
mesochlaena 642
peruviana 643
plana 646, 648
Reichenbachii 648
Lycomormium 600
elatum 600
squalidum 602
Macradenia 944
tridentata 944
Malaxis 367
calycina 368
carnosa 368
caudata 837
caulescens 371
excavata 369
fastigiata 369
hastUabia 369
liparidioides 370
Parthonii 371
rupestris 371
termensis 372
var. elata 372
ventricosa 373
Marsupiaria vandiformis 740
Masdevallia 224
abbreviata 228
aequilpba 232
amabilis 228
angulata 231
aspera 229
atropurpurea 229
aureo-rosea 229
auropurpurea 229
Barlaeana 230, 244
bicolor 230
biflora 230, 232
Bonplandii 231
burfordiensis 231
caloptera 232
cinnamomea 232
civilis 232
coccinea 233
constricta 234
culex 316
cyathogastra 239, 240
Davisii 234
Dayana 223
Forgetiana 236
fulvescens 244
grandiflora 234
Harryana 233
Herzogii 229, 230
hymenantha 236
infracta 236
ionocharis 237
Lindenii 233
longiflora 237
maculata 238
melanopus 238
minuta 239
nidifica 239, 240
pandurilabia 240
perpusilla 240
peruviana 242
plantaginea 242
polysticta 243
pulex 316
pumila 243
purpurina 244
rufolutea 232
Schroederiana 244
Simula 240
INDEX
1017
splendida 244
tenuicauda 239, 240
umflora 231
uniflora 245
Vargasii 245
Veitchiana 244, 247
venusta 233
Weberbaueri 247
xanthodactyla 238
xanthura 229, 230
Maxillaria 627, 668
Abelei 727, 728
acuminata 676
alata 676
alba 685
alticola 676
angustibulbosa 677
arbuscula 678
argyrophylla 678
attenuata 680
aurea 680
var. gigantea 681
Batemanii 682
bicolor 682
brachypetala 682
bractescens 628
brevipedunculata 719
breviscapa 683
brunnea 727
caespitosa 683
calantha 684
Camaridii 684
cassapensis 712
christobalensis 685
ciliata 642
cinnabarina 641, 642
colorata 687
concern 629, 631
cornuta 687
costata 642
crassicaulis 688, 976
crocea 690
cuneiformis 690
cuzcoensis 690
densifolia 692
dichroma 694
disticha 693
divaricata 693
dolichophylla 683
eburnea 700
elegantula 694
elongate, 629
exaltata 695
fallax 693, 694
fasciculata 695
fimbriata 642, 643
Fletcheriana 696
floribunda 696
foveata 629, 631
Fuerstenbergiana 698
funicaulis 699
graminea 651, 652
graminifolia 699
grandiflora 700
grandimentum 702
hastata 702
heteroclita 660, 662, 702, 704
huancabambae 704
var. cuzcoensis 704
infausta 705
insignis 705
Johniana 705
Klugii 706
Koehleri 706
laevilabris 708
laricina 749, 750
Lehmannii 700
ligulala 708
Lindeniae 708
linearis 709
longibracteata 709, 710, 712
var. grandiflora 710
var. luteorubra 710
longifolia 712
longipes 712
longipetala 713
loretoensis 713
luteorubra 712
Macleei 738, 739
macrophylla 646, 648
macropoda 714
maculata 714
Mathewsii 716
Matthewsii 699
meridensis 716
multicaulis 717
nanegalensis 717
nardoides 718
nasuta 719
nervosa 693, 694
nubigena 719
nutans 720
ochroleuca 685
ornate 632
pallidiflora 634
paniculata 720
parvibulbosa 720
parviloba 722
pendula 546, 722
platyloba 723
platypetala 723
Poeppigiana 690
polypetala 723
prolifera 724
purpurea 724
var. parviflora 546, 725
ramosa 725
ramosissima 725
ringens 726
rotundilabia 727
rufescens 727, 728
Sanderiana 728
sanguineomaculata 730
saxatilis 730
1018
FIELDIANA: BOTANY, VOLUME 30
scabrilinguis 634
serrulata 676
spathulata 731
splendens 731
squalens 634
Stachyobiorum 629, 631
stenobulbon 634
striata 732
supina 634, 635
Tafallae 734
tenuis 735
tricolor 735
trigona 735
triphylla 738
uncata 738, 739
undatiflora 739
undulata 636
unicarinata 739
vandiformis 740
variegata 740
verrucifera 740
vestita 724
villosa 742
violaceopunctata 744
Warreana 578, 579
Weberbaueri 688, 745
Woytkowskii 745
xantholeuca
var. peruviana 746
xanthorhoda 746
xylobiiflora 748
Mesadenella esmeraldae 138
Mesospinidium densiflorum 794
roseum 795
sanguineum 833, 834
vulcanicum 796
Wattirii 824, 825
Microchilus bifalcis 152
foliosus 153
major 154, 156
minor 156
ovatus 159
querceticola 159
repens 160
Microstylis 367
calycina 368
excawzta 369
fastigiata 369
hastilabia 369
humilis 498, 500
liparidioides 370
Parthonii 371
rupestris 371
termensis 372
centn'cosa 373
Miltonia 847
parya 847
vexillaria 848
Warscewiczii 848
Mormodes 581
revolutum 581, 582
Rolfeanum 582
Wolterianum 582
Mormolyca 750
peruviana 750
Myanthus barbatus 584
Myoxanthus monophyllus 326
Myrosmodes nubigenum 101, 102
N anodes discolor 511, 512
Nasonia 981
Hartwegii 983
ionanthera 983, 984
punctala 983
Neodryas 797
densiflora 797
latilabia 798
Mandonii 798
reniformis 798
rhodoneura 798
Weberbaueriana 799
Neokoehleria 773
equitans 774
Markgrafii 774
paniculata 775
peruviana 775
Neolehmannia Micro-Cattleya 473, 474
Neottia acaulis 131
orchioides 142
picto 131
speciosa 146
vaginata 148, 149
Notylia 929, 944
Awem 932
boliviensis 930
Buchtienii 931
carnosiflora 931
coffeicola 932
conduplicans 932
Koehleri 934
mirabilis 936
moyobambae 931
parvilabia 936
peruviana 938
rhombilabia 938
Wullschlaegeliana 940
Nyctosma nocturna 482, 484
Octomeria 361
boliviensis 361
var. grandifolia 362
complanata 362
erpsilabia 364
scirpoidea 364
semiconnata 365
surinamensis 365
Odontoglossum 799
angustatum 804, 805
anthoxanthum 805
aureo-purpureum 805, 806
bellum 804, 805
bicolor 860, 862
brachypterum 806, 807
INDEX
1019
brevifolium 806, 807
var. Weberbauerianum 807
brevilabium 808
candelabrum 806
cardiochilum 892
chiriquense 806, 807
cimiciferum 865, 866
cirrhosum 808
compactum 805, 806
coronarium 806, 807
var. chiriquense 806
var. miniatum 806
crista-galli 867
cristatum 810
crocatum 810
depauperatum 811
digitatum 811
epidendroides 812
festatum 861
flavescens 814
fractiflexum 814
graminifolium 815
Hallii 815
Harryanum 816, 832
hemichrysum 861
Hennisii 816
Hrubyanum 818
juninense 818
Koehleri 805, 806
lacerum 819
Loesenerianum 804, 805
longifolium 819
longipes 820, 821
megalophium 820
melanthes 821
mentigerum 889
microthyrsus 821
miniatum 806
multistellare 822
myanthum 822
myrianthum 823
mystacinum 810, 823
Noezlianum 795
obscurum 824
praestans 825
pusillum 825
retusum 826
revolutum 828
rigidum 823, 824
ringens 828
roseum 795, 796
subuligerum 829
tetraplasium 804
tigroides 829
trilobum 830
tripudians 832
vexillarium 848
Wallisii 824
Weberbauerianum 807
Weltonii 848
Wyattianum 832
zebrinum 915
Oncidium 850, 851
acinaceum 858
aemulum 907
altissimum 863, 864
ampliatum 858, 859, 860
armillare 860
aureum 824, 860, 862
(B) stenochilum 861
aurosum 871, 872
Batemanianum 862, 864
Baueri 863, 864
Bernoullianum 858, 860
bidens 836
bolivianum 863, 864
bryolophotum 878, 879
cajamarcae 864
calanthum 864
cardiochilum 892, 893
caudatum 837
chrysopyramis 901, 902
cimiciferum 865
cirrhosum 808
cocciferum 866
convolvulaceum 876
cordatum 866
corynephorum 912, 913
crista-galli 867, 868
cristatum 810
cruciferum 868
cryptocopis 869
cucullatum 891
var. nubigenum 891
Davisii 872, 873
decipiens 867, 868
deltoideum 869, 870
diceratum 870
dichromum 861
discobulbon 871
elegantulum 840
epidendroides 812
excavatum 871, 872
var. Dawsonii 871
falcicorne 898
falciferum 872
falcipetalum 872, 873
var. Pavonii 872
fasciferum 873
festatum 861
flexuosum 865
formosissimum 874
foveatum 860
fuscatum 848
gargantua 875
globuliferum 875, 876
/3 costaricense 876
glossomystax 877
gracillimum 877
guttatum 885
hastiferum 888
hemichrysum 861
heteranthum 878, 879
1020
FIELDIANA: BOTANY, VOLUME 30
hians 879
hieroglyph! cum 880
hirundo 880
holochrysum 893
incarum 881
inferlobum 907
inops 878, 879
ionodon 872, 873
ionopterum 882
iridtfolium 867, 899
Kienastianum 882
leopardinum 883
Leopoldianum 883
leucostomum 879
lineoligerum 884
Lucasianum 885
luridum 885
var. guttatum 885
macranthum 886
var. hastiferum 888
Mathieuanum 888
maxilligerum 879
megalophium 820
megalous 878, 879
mentigerum 889
minax 889
nanum 890
nubigenum 891
obryzatum 891
ochmatochilum 892, 893
onustum 893
ovatilabium 894
Papilio 896
pardo thyrsus 896
patulum 890
pentadactylon 897
peruvianum 845
pictum 898
polyadenium 898
pusillum 899, 900
pyramidale 901, 902
quadricorne 879
retusum 826
retusum 902
revolutum 828
rigidum 823
Sanderae 903
Sanderianum 905, 906
scansor 876
Schillerianum 904
serpens 905
serratum 905, 906
sessile 906
signatum 845
suave (as suavis) 896
superbiens 907, 908
tigratum 908
tipuloides 908
trilingue 882
trilingue 909
tripterygium 910
trulla 910
undulatum 907, 911
ventilabrum 911, 912
villosum 846
volubile 912
Weberbauerianum 807, 913
Weltonii 848
Wercklei 876
xanthodon 914
xanthornis 914
zebrinum 915
Ophrys
ciliata 112, 113
ensifolia 370
monophyllos 368
paleacea 101
Orchidofunckia pallidi flora 941, 942
Orchidotypus muscoides 978
Orchis monorrhiza 29
setacea 29
Orleanesia 551
peruviana 551
Ornithidium 668
album 685
aureum 680, 681
bicolor 861
bolivianum 681
dichotomum 734, 735
distichum 693
divaricatum 693, 694
dolichophyllum 683, 684
giganteum 681
huancabambae 704
multicaule 111
nubigenum 719, 720
pendulum 722
serrulatum 676
simulans 724, 725
Tafallae 734
vestitum 724, 725
Weberbauerianum 688, 976
Ornithocephalus 924
apiculatus 925
avicula 925
gladiatus 926
var. peruvianus 926
longilabris 926
patentilobus 927
polyodon 927
Pachyphyllum 973
breyiconnatum 974
capitatum 974
denticulatum 975
distichum 975
gracillimum 976
haemathodes 976
lycopodioides 976
muscoides 978
parvifolium 978
Pastii 976
Pastii 979, 980
pectinatum 980
INDEX
1021
serra 980
Stuebelii 979, 980
subbiflorum 981
tenue 981
Paphiopedilum
Boissierianum 17
caricinum 18
caudatum 19
Czerwiakowianum 19
Hartwegii 21
Pelexia 129
corymbosa 147
foliosa 153
/wrta 139
laxa 140
maculata 141
matucanensis 144
repews 160
saltensis 146
Weberbattenana 149
Peristeria 602
fuscata 602
maculata 603
pendula 603
Petalocentrum 916
angustifolium 921
bicornutum 921
pusillum 921
Phajus Hookerianus 578
Phragmipedium 17
Boissierianum 17
caricinum 18
caudatum 19
Czerwiakowianum 19
Hartwegii 21
Phragmopedilum Boissierianum 17
cajarnarcae 17, 18
caricinum 18
caudatum 19
Czerwiakowianum 19
Hartwegii 21
Physinga prostrata 496
Physosiphon 222
emarginatus 223
Physurus 150
arietinus 152
&t/a(a's 152, 153
claviger 153
foliosus 153
major 154
minor 156
mystacinus 157
ovatus 159
PresZei 154, 156
querceticola 159
repens 160
santensis 160
stenocentron 163
validus 163
Pilumna fragrans 788
Pityphyllum 749
laricinum 749
Plantanthera foliosa 33
Pleurothallis 261
acuminata 270
acutiflora 271
affinis 271
aZfoda 322
amygdalodora 271
angustilabia 272
angustipetala 272
angustisegmenta 330
antennifera 274
araguensis 339
arbuscula 274
Archidpnae 274
asperilinguis 275, 276
attenuata 275
aurea 275, 276
bicornis 276
bivalvis 345
bogotensis 325
brachyblephara 287, 288
brevispicata 277
Cajamarcae 277
carinata 277
carnosifolia 279
casapensis 279, 281
Cassidis 281
caudatipetala 281
caulescens 282
cauliflora 282
chamaelepanthes 283
chamensis 283
var. tenuis 284
chanchamayoensis 284
Chuquiribambae 119
ciliaris 285
ciliata 285
var. abbreviata 286
var. elongata 287
citrina 287
var. elliptica 272
coccinea 772, 773
complicata 293, 294
cordata 180, 287, 349
crateriformis 288
Crocodilanthe 290
cucullata 330
cyathiflora 290
declivis 292
decurrens 292
dichotoma 293
diffusa 293, 294, 295
var. grandiflora 294
dilatata 295
dinotherii 295, 296
diptera 295, 296
diuturna 320, 321
divaricans 296
dura 296, 297
echinocarpa 297
ecuadorensis 301
emarginata 223
1022
FIELDIANA: BOTANY, VOLUME 30
excisa 298
fimbriata 301
fimbrilabia 301
flexuosa 302
floribunda 304
floripecten 358, 360
fons florum 304
frutex 304
var. robusta 305
gelida 305
genychila 306
gigantea 306
gracilis 338
graciliscapa 307
graminea 282
grandiflora 307
Grobyi 309
hemirhoda 309
hirsutissima 310
huanucoensis 357, 358
inaequisepala 310
juninensis 312
var. angustifplia 314
var. subaequisepala 314
lanceolata 199, 210, 314
var. gracilis 338
lancilabris var. oxyglossa 315
Lindenii 339
var. gigantea 339
var. longiracema 340
var. pumila 339
linearis 315, 316
longiserpens 316
macroblepharis 316
macrophylla 304
macrorhiza 318, 344
magnipetala 318
Mathewsii 325, 326
misera 320
monstrabilis 237
multicaulis 335, 336
muricata 320, 321
myoxanthus 326
nigro-hirsuta 321
obovata 321
octomeriaeformis 322
oxyglossa 315
pantasmoides 322
parvifolia 322
pedunculata var. peruviana 324
penduliflora 324
phalangifera 325
phyllostachys 326
picta 309
Pittieri 354, 356
plantaginea 242
Poeppigii 326
pruinosa 315, 327
pusilla 327
quadrata 328
remotiflora 302
restrepioides 328
retusiloba 330
rhizomatosa 318
rhynchantha 330
rigidifolia 271, 332
ringens 333
rubens 333
ruberrima 335
rufescens 320
ruscifolia 335
scabrata 336
scabrilinguis 338
Schulzeana 315
Schweinfurthii 338
secunda 339
var. longiracema 340
semipellucida 293, 294
var. grandiflora 294
serripetala 340
simplicilabia 341
Smithiana 341
sororia 320
spathata 342
spathulifolia 342
spiralis 215, 344
stenopetala 344
succosa 335, 336
syringifolia 344
Talpinaria 345
tentaculata 346
tenuifolia 346
var. longisepala 348
tenuis 348
trachychlamys 349
trachysepala 349
trialata 350
tricarinata 350
tricaudata 296
tridentata 351
trilineata 351
trimeroglossa 345, 346
Trulla 352
tunguraguae 352
umbraticola 353, 354
undulata 353
unilateraUs 358, 360
vaginata 353, 354
Vargasii 295, 354
velaticaulis 354, 356
verruculosa 357
vittata 309
xanthochlora 357, 358
xiphizusa 290
Pogonia 39
monantha 47
Moritzii 40
rosea 40
Vargasii 40
Polycycnis 605
muscifera 605
vittata 606
INDEX
1023
Polystachya 554
altilamellata 555, 556, 558
amazonica 555, 558
boliviensis 554
caracasana 555, 558
cerea 555, 558
ecuadorensis 555, 558
foliosa 555
Huebneri 555, 558
luteola 558
minor 555, 558
nana 555
Poeppigii 555
stenophylla 555, 558
Ponera affinis 542
conferta 543
graminifolia 545
mapiriensis 544, 545
punctulata 548
violacea 550
Ponthieva 118
bicornuta 119
caZoa 111, 112
chuquiribambae 119
diptera 121
inaudita 121, 122
Koehleri 121
lilacina 122
Mandonii 122
microglossa 121, 122
montana 124, 125
multiflora 115
oligpneura 125
similis 125
Sprucei 126
venusta 124
villosa 126
Weberbaueri 127
Porphyrostachys pilifera 91
Fresco ttia 104
barbifrons 107, 108
colorans 104, 105
longipetiolata 104, 105
petiolaris 104, 105
pteristyloides 98, 100
stachyodes 104
tenuis 105
Promenaea graminea 651
Pseuderiopsis Schomburgkii 596
Pseudocentrum 106
bursarium 106
macrostachyum 106
Pterichis 107
acuminata 107, 108
barbifrons 108
galeata 107, 108
leucoptera 108
macrpptera 108
seleniglossa 109, 110
silvestris 109
triloba 109
Weberbaueriana 110
Regnellia purpurea 562, 565
Restrepia cucullata 330, 332
ecuadorensis 301
rhynchantha 330, 332
tentaculata 346
vittata 309
Rodriguezia 770
Batemanii 771, 772
var. speciosa 772
bicolor 783
Candida
var. Batemannii 771
ensiformis 772
lanceolata 772
secunda 772, 773
Sarcoglottis 129
olivacea 142
Pavonii 144
picta 131
speciosa 131
Satyrium elatum 134, 138
orchioides 142, 144
Scaphyglottis 541
afRnis 542
Antonii 542, 543
conferta 543
cuneata 544, 545
graminifolia 545
Huebneri 545
loretoensis 542, 543
parviflora 546, 725
pendula 546, 722
prolifera 544
propinqua 546
punctulata 548
var. Summersii 548
Reedii 550
Summersii 548
Tafallae 734
violacea 543, 550
Scelochilus 776
brevis 776
latipetalus 778
saccatus 780
Schomburgkia 536
moyobambae 536
Weberbaueriana 537
Selenipedium Boissierianum 17
caricinum 18
caudatum 19
Czerwiakounanum 19
Hartwegii 21
Pearcei 18
Serapias parasitica 368
speciosa 146, 147
Sertifera virgata 68
Siagonanthus multicaulis 717
Sievekingia 598
peruviana 598
Sigmatostalix 916
amazonica 916
1024
FIELDIANA: BOTANY, VOLUME 30
aurosanguinea 917
bicallosa 917, 918
bicornuta 921
crescentilabia 918
graminea 920
peruviana 920, 921
pusilla 920, 921
Sobralia 68
alstroemerioides 75, 76
amplexicaulis 46, 69
biflora 69
Candida 70
crocea 71
dichotoma 71 , 72, 75
D'Orbignyana 72
fimbriata 70, 73
Klotzscheana 73, 74
leucoxantha 74
Mandonii 71, 72
rosea 74, 75
Ruckeri 74
scopulorum 75, 76
sessilis 70
setigera 76
violacea 74, 75, 76
alba 77
Weberbaueriana 77
Solenidium 850
peruvianum 850
Specklinia ciliaris 285
dichotoma 293
flexuosa 302
graminea 920
linearis 315
obovata 321
plantaginea 242
pusilla 327
Spiranthes 128, 129
acaulis 131
alpestris 136, 138
colorata 146, 147
var. maculata 146
corymbosa 131
costaricensis 132
curvicalcarata 134
elata 134, 145
Esmeralda 138
hirta 139
inaequilatera 139
laxa 140
Lechleri 140
macrostachya 91
maculata 141
matucanensis 144, 145
micrantha 141
minutiflora 136
olivacea 142
orchioides 142
pachyrhiza 144, 145
Pavonii 144, 145
peruviana 134, 136
picta 131
plantaginea 134, 138
Preslii 134, 136
pumila 145
Rimbachii 145
saltensis 146
speciosa 131
speciosa 146
subumbellata 147
Ulaei 148
vaginata 148
variegata 136, 138
Weberbaueri 149
var. aurantiaca 149
Weberbaueriana 149
Stanhopea 606
anfracta 611, 612
aurata 608
connata 607
convoluta 611
graveolens 607
graveolens 608
Haseloviana 609
inodora 608
Moliana 609
peruviana 611, 612
Randii 610
tricornis 611
Wardii 611, 612, 786
var. venusta 611
xytriophora 612
Stelis 163
acutiflora 170
acutissima 171
affinis 171
alba 171
apiculata 173
apiculata 192
ascendens 173
ascensor 173
aspera 174
attenuata 174
bicallosa 176
breyiracema 176
capillaris 178
capillipes 179, 191
carnosa 368, 369
chachapoyensis 178
concaviflora 179
concinna 179
connata 179
contorta 180
cordata 180, 287
cordibractea 207, 209
cupuligera 180
curvicarina 182
densiflora 182
diffusa 184
discolor 184
dispar 186
disticha 186, 314
dupliciformis 187
elatior 187
INDEX
102{
Endresii 189, 190
eublepharis 190
var. glabriflora 190
Filomenoi 191
flacca 191
flexuosa 191
floribunda 173, 192
foliosa 555, 558
Gonzaleziana 193
gracilispica 193
grandibracteata 193
Hallii 194
var. minor 194
Herzogii 196
hirta 196
Huancdbambae 207, 209
Huebneri 189, 190
hylophila 196
insignis 192
intermedia 197
in versa 197
juninensis 198
Koehleri 198
lancea 199
lanceolala 199, 314
leucopogon 199, 200
Lindenii 201
Lindleyana 201
var. carnosior 174
lobata 202
macra 202
megistantha 203
melicoides 203
minuta 203
mononeura 204
nephropetala 204
oblonga 205
Ottonis 205
pardipes 219
parviflora 205
phaeantha 207, 209
phaeomelana 205
piestopus 205
polycarpa 206
polystachya 205, 206
punoensis 207
purpurea 207, 215
quinquenervia 209
recur vula 210
revoluta 210
rhizomatosa 210
rhombilabia 211
rhomboglossa 211
santiagoensis 211
scansor 213
serra 213
simacoensis 214
spathulata 214
spiralis 215, 344
stenophylla 215
striolata 215
Tessmannii 216
thecoglossa 216
triangulisepala 217
tricardium 217
triseta 217
var. pardipes 219
truncata 207, 209
uninervia 219
velutina 220
viridipurpurea 220
Weberbaueri 222
Stellilabium 961
astroglossum 961
Stenia 664
fimbriata 665
guttata 664
pallida 664
Stenocoryne longicornis 638
Stenoptera 84
acuta 85
ananassocomos 85
cardinalis 91, 92
ciliaris 86
data 85
goodyeroides 79
laxiflora 86
macrostachya 91
montana 88
peruviana 91
pilifera 91
var. parviflora 92
plantaginea 83
viscosa 83, 84
Stenorrhynchus 129, 132
Esmeraldae 138
laxus 140
orchioides 142
saltensis 146
speciosus 146
vaginatus 148
Sturmia elliptica 380
Sutrina 783
bicolor 783
Symphyglossum 833
sanguineum 833
Synassa 128
corymbosa 147
Talpinaria bivalvis 345, 346
Telipogon 946
astroglossus 961
Benedicti 947
Boissierianus 947
Fritillum 948
gnomus 948
gymnostele 949
Hercules 950
latifolius 950
obovatus 951
Papilio 951
radiatus 952
rhombipe talus 952
semipictus 954
1026
FIELDIANA: BOTANY, VOLUME 30
tessellatus 954
urceolatus 955
Vargasii 955
Warscewiczii 958
Tetragamestus gracilis 544, 545
Trichocentrum 761
albopurpureum 762
brevicalcaratum 762
panduratum 764
pulchrum 764
triquetrum 766
Trichoceros 958
armillatus 959, 960
astroglossus 961
muralis var. platyceros 958
muscifer 959, 960
parviflorus 959, 960
platyceros 958
Tupaipi 960
Trichopilia 787
brevis 788
fragrans 788
var. grata 789
gracilis 789
grata 789
hymenantha 793
juninensis 791
peruviana 791
subulata 791
Trigonidium 754
loretoense 754
obtusum 756
peruvianum 756
tenue 756
Trizeuxis 784
andina 784, 785
falcata 784, 785
Vanilla 42
appendiculata 42
bicolor 42
grandiflora 43
guianensis 43
hamata 42
lutescens 43
odorata 42, 43
pompona 43
Ruiziana 42, 44
surinamensis 43
Weberbaueriana 44
Vargasiella 367
peruviana 367
Warrea 576
Hookeriana 578
speciosa 578, 579
tricolor 578, 579
Warreana 578
Warscewiczella 654, 704
amazonica 655, 656
heteroclita 660
Wullschlaegelia 78
calcarata 78
Xylobium 627
bractescens 628
carnosum 635
concavum 629
elatum 628
elongatum 629
Filomenoi 631
foveatum 629
Houtei 634
latilabium 631
latifolium 634
ornatum 632
pallidiflorum 634
scabrilingue 635
squalens 634, 638
stachyobiorum 631
subintegrum 636
supinum 635
undulatum 636
Zygopetalon Mackaii 656, 658
Zygopetalum 654, 704
acuminatum 655
amazonicum 655
bolivianum 656, 658
intermedium 656, 658
var. peruvianum 656, 658
Klugii 658
Lindenii 655
Mackayi var. intermedium 656
Prainianum 658
rhombilabium 660, 662, 704
Publication 913