Vol. XX
January, 1944
No. 1
THE
Pan -Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
KEIFER, APPLIED ENTOMOLOGICAL TAXONOMY 1
COOLEY, IXODES NEOTOMAE, A NEW SPECIES FROM CALIFORNIA 7
NUNENMACHER, A NEW SPECIES OF ZACOTUS 12
PRINCE, DESCRIPTIONS OF THREE NEW SPECIES OF THRASSIS AND
THE FEMALES OF T. BACCHI AND T. PANSUS 13
REES, A NEW MOSQUITO RECORD FROM UTAH 19
FENDER, NOTES ON CANTHARIS, II 20
LEECH, LIST OF SIX SPECIES OF GYRINID WATER BEETLES COL-
LECTED BY J. L. GRESSITT IN CHINA AND JAPAN 22
MALKIN, NEW SPECIES OF BOLETOBIUS FROM THE NORTHWEST
WITH A DISTRIBUTIONAL NOTE ON COPROPHORUS 23
LEECH, SPRING FLIGHT OF ATIMIA DORSALIS...- 30
POTTS, SYSTEMATIC NOTES CONCERNING AMERICAN ACRAEINAE, II 31
PROCEEDINGS OF THE PACIFIC COAST ENTOMOLOGICAL SOCIETY.... 35
San Francisco, California
1944
THE PAN-PACIFIC ENTOMOLOGIST
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The Pan-Pacific Entomologist
VOL. XX, No. 1 January, 1944
APPLIED ENTOMOLOGICAL TAXONOMY*
BYH. H. KEIFER
Bureau of Entomology and Plant Quarantine
State Department of Agriculture, Sacramento, California
Governmental, that is tax-supported, institutions are assuming
increasing responsibilities for the welfare of the general public.
Tax-supported economic entomology is no exception to this, and
governmental organizations are the principal places where eco-
nomic entomologists find employment. When tax money is spent
for arthropod taxonomy in connection with economic organiza-
tions, that taxonomy must be the kind that can be directly ap-
plied to pest control, because pest control is the chief entomo-
logical interest of the tax payer. Thus, most tax-supported tax-
onomists work with arthropods as they relate to legislative pest
control, surveys, insect eradication, chemical control, and biol-
ogical control. He must furnish identifications of species, and
bionomical and distributional data. I have been engaged in this
type of work for the past fifteen years, and this has given me
a considerable opportunity to gain experience with, and to ob-
serve, what sort of taxonomic work is required, how adequate
the existing taxonomic literature is, how specialists in this subject
meet their problems, what attitude these taxonomists take toward
their work, and how the institutions that need this work support
their taxonomists.
From this experience I have developed the conviction that
applied taxonomy can have more definite objectives than it now
seems to show, with resulting benefits to economic entomology.
But the responsibility and vision for maintaining such objectives
must rest largely in the organizations hiring taxonomists.
We may define applied taxonomy as that portion of arthropod
classification dealing with the specific identification in and the
bionomical coordination of, anthropod groups containing spe-
cies in competition with or dangerous to the human animal; or
in groups which contain species that help control effectively the
* Presidential address read before the Pacific Coast Entomological Society on
DecembtFr 18, 1943.
2
THE PAN-PACIFIC ENTOMOLOGIST [yoL. XX, NO. 1
harmful kinds. In other words, this form of taxonomy is a re-
stricted activity. Arthropod groups having the greatest number
of important species receive the most attention, with those having
fewer important species receiving proportionately less.
Taxonomy in its broadest and most fundamental aspect is
principally the search for facts bearing on the origin of, and
interrelationships and differences between, organisms and groups
of organisms. Any restriction of taxonomy such as that here
treated, must rest on the fundamentals.
Our gauge, as to the direction in which the energies of ap-
plied taxonomy are expended, rests on evaluating each group of
arthropods according to the number of economic species it con-
tains. I use the word group because taxonomy is a science of
comparison, and the taxonomist knows important species thor-
oughly only after he has carefully compared them with all similar
species. Thus, a group may be anything from a portion of an
order to part of a family. For example, a worker who compares
and describes species in the scale insect group, continually
sharpens his wits on identifying a considerable number of eco-
nomic species. Opposed to this, a person studying dragonflies
could not, by that activity, identify any economic species. This
does not mean that the taxonomy of dragonflies, mayflies, and
other groups we might mention, is worthless. Such studies ob-
viously have a definite place in entomology, but are inappro-
priate when supported by economic organizations.
The most important activity of the applied taxonomist is the
identification of species. In taxonomy which seeks the funda-
mentals of arthropod correlation, the identification of species
may well be of secondary significance. But in economic entom-
ology the identification of an important species is the key to the
situation. It is the first step of an economic operation. It gives
the control man access to the past literature on the pest, and
enables the orderly recording of new data. Our most notable
recent example of the role of taxonomy in economic operations
is the identification of a single caterpillar from Orange County
(California) in October, 1942, as the larva of the oriental fruit
moth. There followed many meetings of fruit growers and en-
tomologists consuming considerable time and traveling expense.
The State Legislature appropriated over $850,000 for various
phases of investigation and control. Between 500,000 and 600,-
000 moth specimens that were caught in the oriental fruit moth
JANUARY, 1944]
KEIFER— APPLIED TAXONOMY
3
traps were identified in order to segregate the actual oriental
fruit moths and learn the range of that species in California.
While the moth was discovered here too late to allow eradica-
tion, taxonomy has opened the way for control procedures and
investigations before the pest has overrun commercal fruit
orchards.
With the principal activity of applied taxonomy in mind,
namely the dispensing of specific determinations, it follows that
the next most important thing is that of seeking new and better
methods for differentating species, and for the recording of new
species — a large task. Increased knowledge on species differen-
tiation has in recent years had a profound effect on the control
of screw-worm flies, red, yellow, and related scales, tomato pin-
worm, and others. As little planning was done looking toward
such results, we can only wonder how many more such things
remain to be discovered by the chance of personal initiative.
The application of taxonomy to economic entomology has
been, and is being, practiced in a very limited fashion. Not many
tax-supported entomological organizations give it a definite place
in their set-ups. It enters into the planning of almost no eco-
nomic projects. Commercial insecticide organizations freely
consult governmental taxonomists but assume no responsibility
themselves for the development of knowledge on insect identifi-
cation. Applied taxonomy is not taught anywhere as such. Aside
from what research the man on the job can do himself, the most
he usually obtains from outside sources is an occasional chance
article which is partly useful.
The main reason our systematic data so often fails the applied
taxonomist is that he must examine material and use techniques
not popular with the usual taxonomist. This material consists
mainly of species in the order Homoptera and of holometabolous
larvae. While the source material in the Homoptera is in most
respects fragmentary or preliminary from the applied viewpoint,
one family of scale insects, the Diaspididae, has recently been
well clarified. Let me point out that scales have perhaps more
economic species than any other group of comparable size, and
that the worker who has thus helped us is not employed by an
economic organization.
It is in the research on and knowledge of immature forms
that we find the most discrepancies in data available to the ap-
plied taxonomist. We have some valuable leads toward identi-
4
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 1
fying holometabolous larvae but most of them are barely intro-
ductory. Keys for identification of caterpillars by Fracker,
for coleoptera larvae by Boving and Carigbead, and for part of
tbe more primitive maggots by Mallocb, are among tbe best ex-
amples of comprebensive literature on holometabolous larvae.
In many particulars tbe identification features of important
immature stages are at best in tbe minds of but a few people.
Tbe medical side of applied taxonomy is probably in tbe best
position, with considerable tborougbly-developed literature, es-
pecally on mosquitoes and tbeir larvae. In contrast to tbis, in
spite of all tbe years that tbe cotton boll weevil and relatives
bave been serious pests, we still bave no careful analysis of tbe
larvae of species in tbe antbonomine group. We know very little
about immature Homoptera and Hemiptera. Immature stages,
especally bolometabolous larvae, are usually tbe ones found
doing tbe damage, and tbeir identification is the key to the situa-
tion, as pointed out in connection with the oriental fruit moth.
This is vital to economic entomology, but there is no clearly-
defined effort to do anything about it.
Research on new and better methods of species differentiation
includes other activities as well as critical laboratory analytical
methods. The collection and study of arthropods in the field by
the taxonomist not only enables him to get adequate material, but
gives him invaluable perspective on his work. This is true in re-
gard to quarantined insects that do not occur in the area in which
the applied taxonomist works, but the identification of which gives
a meaning to legislative control. And, in order for the taxonomist
to correlate the visble structures he studies with the habits or
activity of the species, and to disclose the true nature of poly-
morphism, applied taxonomic studies should be conducted in
connection with greenhouses and rearing rooms.
If we stopped our consideration of applied taxonomy right
here, this branch of classification would be abundantly worth-
while. But there is more to it than that. The simplest way tax-
onomy aids insect control beyond species identification is by
enabling the economic entomologist to borrow methods from the
control of species similar to the one under suppression. An
example would be the use of established scale control methods
on new scale pests. Thus, taxonomy serves as an orderly basis
for thinking, and correlates control methods. The spectacle of
an economic entomologist fastening his attention entirely on one
JANUARY, 1944]
KEIFER— APPLIED TAXONOMY
5
\
species gives the impression that he will sooner or later be
operating with a poverty of new entomological ideas. The cod-
ling moth control fraternity is perhaps the best example in this
case.
Another way taxonomy can go farther than species naming is
that of defining genera whenever possible, so that they not only
have a structural significance, but also a bionomical or geo-
graphical aspect. Geographical considerations are of prime im-
portance in parasite explorations. In this connection, biological
control organizations seem to have neglected what we might term
the “geo-taxonomy” of the insect groups that contain species
they seek to control.
Taxonomy is primarily based on the comparison of arthro-
pod structures. These structures are present because of some
mechanical value to the individual arthropod. Structures are
visible mechanisms, and the taxonomist, by comparing them,
makes logical arrangements of these visible mechanisms. Impor-
tant control methods are based on such things as oral mechan-
isms, skin structures, and gland secretions. These have definite
places in taxonomy, which has a wealth of comparative data on
such structures. It is of profound significance that serological
studies indicate that the chemical similarities of arthropods
closely follow their mechanical aflEnities.
Since taxonomy proceeds largely by the process of thorough
comparison, therefore comparison is at the basis of taxonomic
reasoning. With this fact in mind, let us momentarily analyze
present practices in arthropod-control research. We immediately
see a strong tendency to treat arthropod pest species as unrelated
entities. For example, there are many articles in existence de-
scribing the reactions of one species of insect to numerous control
chemicals. That is perfectly correct as far as it goes, but taxo-
nomic reasoning indicates that if the reverse aspect of this is
not also investigated the whole thing is lop-sided. In other words,
the reactions of numerous arthropod species to single chemicals
will give interspecific variations in reaction, and these variations
can then be correlated with visible mechanisms and biochem-
istry. This does not imply that such a thing ought to be done in
connection with each control project, but as a standard practice
to give valuable data to control projects. The significance of
the reciprocal toxicity of sulfur and cyanide to arthropods lack-
ing or possessing spiracles would be a good subject for an in-
6
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 1
vestigation of this type. The chemical side of the problem is the
“what” side; the arthropod side the “why” side. I am not re-
ferring in this case to attempts by insecticide companies to sell
their products as cure-alls.
Remarks have already been made leading up to the statement
that applied taxonomy needs some special literature. There is
no better origin for such literature than with the applied taxo-
nomists themselves who owe it to their profession to publish the
identification problems they solve. The type of literature needed
is not that which is illustrated by a photograph or general sketch,
but by figures drawn on the basis of interspecific and inter-
generic comparison, analyzing the body comprehensively. There
is enough information in existence scattered through books and
periodicals, preserved in collections, and in the minds of a few
people, to form a valuable manual or manuals, on preparatory
methods and characters for the identification of economic arthro-
pods and near relatives. With this as a basis we could build
further knowledge in a really effective form.
Let me conclude by repeating that the application of arthropod
taxonomy to economic operations means restriction to study of
pertinent arthropod groups. This taxonomy, as well as being
restricted, has certain definite procedures and objectives, the most
important objective being species identification. Applied tax-
onomy depends for its future development squarely on the intel-
ligent support of the organizations that need it. While the eco-
nomic value of taxonomy is conceded, there is a general passive
policy, both expressed and implied, that the solution of pertinent
taxonomic problems must be largely based on the chance co-
incidence of opportunity and ability or inclination. Amateur
taxonomists have done important work, but economic entomology
is a large and growing profession, and the amateur or passive
slant on applied taxonomic development cannot assure such
progress as is needed. Economic entomological organizations
will find that the only effective applied taxonomy is that for
which they are willing to be actively responsible.
JANUARY, 1944]
COOLEY— IXODES NEOTOMAE
7
IXODES NEOTOMAE, A NEW SPECIES FROM CALIFORNIA
( ACARINA : IXODIDAE ) "
BY R. A. COOLEY
Senior Entomologist, United States Public Health Service
A detailed study of numerous specimens of Ixodes which
have the auricjulae in the shape of a pair of long, curved horns
shows that the form in California occurring mainly on the wood
rat {Neotoma) differs from /. diversifossus Neumann, 1899
taken from the raccoon [Procyon lotor) in New Mexico, and /.
spinipalpis Nuttall, 1916 which is found mainly on rabbits in
the Western States. The California species is described below.
Ixodes neotomae Cooley, new species
Female
Body. No unfed specimens are available. Engorged females
may reach at least 6.5 x 4.0^ and are oval, wider and thicker
(dorso-ventrally) behind.
Capitulum. Length (from tips of cornua to tip of hypostome)
from 0.66 to 0.78; width of basis, 0.36 to 0.40. Lateral margins
straight or a little curved, mildly converging posteriorly. Poste-
rior margin straight, salient, and at about the same level as the
distinct, small cornua. Surface mildly and evenly convex, faintly
shagreened, impunctate, and without hairs. Porose areas well
separated, superficial or faintly depressed. Palpi long, with lat-
eral profile lines straight, curved on median side, widest at about
the middle. Surface nearly smooth, shining; hairs few and short
excepting a few long ones on the median side at the base. Com-
bined length of articles 2 and 3 about 0.60, greatest width 0.158.
In ventral view basis is slightly waisted at the middle, poste-
rior corners rounded and the posterior edge nearly straight and
salient. Transverse suture faint or absent. Auriculae as long,
curved horns which are directed downward and backward. Sur-
face smooth, shagreened and shining. Palpi with their inner faces
flattened. Article 1 with a distinct pointed ventral spur and
usually with two fine hairs. Several long, fine hairs arising from
the ventral edge of the palpi.
Hypostome. Long and narrow with the denticles 4|4, 3|3, and
at the base 2|2; denticles long, narrow, and sharp, progressively
smaller toward the middle line. Length about 0.405.
^ From the Rocky Mountain Laboratory (Hamilton, Montana) of the Division
of Infectious Diseases of the National Institute of Health.
- All measurements in this paper are in millimeters.
8
THE PAN-PACIFIC ENTOMOLOGIST [vOL. XX, NO. 1
Scutum. Len^h from 1.05 to 1.17, width 0.78 to 0.87. Oval,
very broadly rounded behind. Lateral carinae as rounded elevations
which may be nearly neg^ligible in some specimens. Cervical
grooves divergent, moderate or faint, and usually absent in the
anterior area. Punctations distinct, larger, and more numerous
in the median posterior area, few or absent outside of the lateral
carinae. Surface shagreened and shining. Hairs long and fine.
Legs. Moderate in length and size, and with the ventral hairs
long. Length of tarsus I, 0.48; metatarsus, 0.24. Length of tarsus
IV, 0.39; metatarsus, 0.30.
Coxae. Flattened, shagreened, and with long fine hairs. Poste-
rior margins salient posteriorly. Internal spur on I long; absent
on II, III, and IV.' External spur on I distinct but short, those
on II, III, and IV progressively smaller and that on IV very small.
Spiracular plate. Nearly circular with the slightly longer axis
trasnverse. Marginal goblets rectangular, medians circular and
larger. Length 0.17, width 0.147.
Sexual aperture. At the level of the posterior edges of coxae III.
Male
Body. Length 1.02, width 0.72. Oval, widest at about the
middle.
Capitulum. Length 0.326; width of basis 0.20. Sides nearly
straight, converging posteriorly. Cornua distinct, small and at the
level of the straight, salient posterior edge. Surface nearly fiat,
shagreened, impunctate, and without hairs. Palpi with their lat-
eral profile lines nearly straight, median sides curved, widest
well back of the middle. Surface smooth and faintly shining, de-
pressed near their median margins. Hairs few and short excepting
a few long ones on the median side at the base. Combined length
of 2 and 3, 0.22; width 0.105. In ventral view basis is broad,
rounded and salient behind; auriculae as short, pointed or rounded
lateral extensions at the same level as the ventral surface. Palpi
fiattened on their inner faces. Article 1 with sharp ventral edge
which is continuous with the ventral edge of articles 2 and 3. A
few long hairs present on ventral edge of articles 2 and 3.
Hypostome. Large, mildly notched apically. Lateral denticles
large and sharp, about four in number on each side, with one large,
rounded tooth directed behind. Median denticles as diagonal crenu-
lations or very faint teeth. Length about 0.216.
Scutum. Oval, convex, about equally declivitous in all sub-
marginal areas. Lateral carinae absent. Cervical groves distinct
but mild, divergent behind and reaching about half the length of
the scutum. Pseudoscutum distinguishable mainly by its darker
color; reaching about half the scutal length. Punctations distinct,
smaller on the pseudoscutum. Surface shagreened, shining. Hairs
long and fine.
JANUARY, 1944]
COOLEY— IXODES NEOTOMAE
9
Legs. Essentially as in the female. Len^h of tarsus I, 0.29;
metatarsus, 0.144. Length of tarsus IV, 0.27; metatarsus 0.144.
Coxae. Essentially as in the female.
Fig. 1. Ixodes neotomae Cooley. A. Capitulum and scutum,
male, dorsal view. B. Capitulum, coxae and ventral plates, ven-
tral view. C. Hypostome, male. D. Spiracular plate, male. E. Ca-
pitulum and scutum of the nymph, dorsal view. F. Capitulum and
coxae of nymph, ventral view. G. Hypostome, nymph. H. Spiracu-
lar plate, nymph.
Ventral plates. Median plate about three times as long as the
anal plate. Adanals much wider in front. Anal plate much wider
behind. Punctations on median and adanals small but distinct;
absent or few and small on anal plate. Hairs few, fine and of
moderate length.
Spiracular plate. Small, oval with the longer axis longitudinal.
10
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 1
Goblets about equal in the marginal and median fields, those in
the median field few in number. Length 0.125, width 0.96.
Sexual aperture. At the level of the intervals between coxae
II and III.
Nymph
Capitulum. Length (from tips of cornua to tip of hypostome),
0.34; width of basis, 0.29. Basis short and broad, fiattened on top.
Cornua short but distinct, a little elevated over the level of the
posterior edge which is nearly straight, salient. Surface faintly
shagreened, shining, faintly punctate and without hairs. Palpi
short, with the lateral profile lines straight, median profile lines
evenly curved. Surface smooth and shining; hairs few, short and
fine. Combined length of articles 2 and 3, 0.15.
In ventral view basis is broad in front, narrow, evenly rounded
behind, and slightly waisted at the middle. Transverse suture not
visible. Auriculae only suggested as mild extensions at the same
level as the ventral wall of the capitulum. Surface smooth, shin-
ing, impunctate and without hairs. Palpi set far away from the
mouth parts, flattened medially. Article I with a conspicuous
retrograde spur.
Hypostome. Bluntly rounded apically. Dentition 2|2 for most
of the length. Laterals larger and only a little pointed; medians
smaller and rounded. Length about 0.187.
Scutum. Length 0.60, width 0.63. Lateral carinae very dis-
tinct, more declivitous on the median side, the two limiting a
median oval depressed area. Cervical grooves deep, long, nearly
parallel, and reaching nearly to the anterior and posterior ends
of the scutum. Surface faintly shagreened, shining, and with a
few small punctations. Hairs negligible.
Legs. Essentially as in the adult.
Coxae. Much as with the adults but with internal spur on coxae
I shorter.
Spiracular plate. Oval, with the marginal and median goblets
of about equal size. Length 0.096, width 0.067.
Holotype. Two females (19430) from Neotoma f. microtis,
November 5, 1941, Los Angeles County, California.
Allotype. One male (16831) from Sylvilagus hachmani cine-
rascens, October 7, 1922, La Jolla, San Diego County, Cali-
fornia.
Paratypes. One female (15970), Sylvilagus hachmani cine-
rascens, September 17, 1922, Angels Camp, Calaveras County;
one female (15811), Neotoma fuscipes, April 17, 1939, Monte-
rey County, California; one nymph (17948), Neotoma fusci-
pes, July 13, 1940, Hastings Natural History Reservation,
California; one female (12943), deer mouse, June 30, 1936,
Oakland Hills, Oakland, California; three females (13064),
JANUARY, 1944]
COOLEY— IXODES NEOTOMAE
11
Sylvilagus sp., February 7, 1937, Gopher Canyon, Vista, Cali-
fornia.
Holotype, allotype, and paratypes 17948 deposited in the col-
lections of the Rocky Mountain Laboratory, Hamilton, Montana.
Paratype 13064 placed in the United States National Museum;
paratype 15811 in the Division of Entomology and Parasitology,
University of California, Berkeley, California, paratype 15970 in
Fig. 2. Ixodes neotomae Cooley. A. Capitulum and scutum,
female, dorsal view. B. Capitulum and coxae, female, ventral
view. C. Hypostome, female. D. Spiracular plate. E. Leg I,
female. F. Leg IV, female. G. Engorged female.
the Museum of Comparative Zoology, Harvard University, Cam-
bridge, Massachussetts.
Females of neotomae are close to those of spinipalpis but can
be separated by the narrower hypostome in neotomae which has
longer, sharper denticles; spinipalpis has the sexual aperture
between coxae IV, while in neotomae it is at the level of the
12
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 1
intervals between coxae III and IV. Adults, especially the males,
are larger in spinipalpis.
The nymphs are very distinct, those of spinipalpis having the
hypostome wider with the lateral teeth more pointed, and with
the dentition 3|3 for nearly half the length and then 2|2, while
in neotomae the hypostome is more rounded apically, with denti-
cles more rounded; dentition 2|2 from near the tip to the base.
The nymphal scutum in neotoma is nearly circular and has dis-
tinct curved lateral carinae, while in spinipalpis it is a little
longer than wide and has shorter, nearly straight lateral carinae.
Both neotomae and spinipalpis are close to diversifossus of
which no nymphs are known. 7. diversifossus is distinctly larger
and has the larger and heavier auriculae placed farther apart
and has no definite ventral horn on palpal article 1.
A NEW SPECIES OF ZACOTUS
(Coleoptera, Carabidae)
BY F. W. NUNENMACHER
Piedmont, California
Zacotus fredericki Nunenmacher, new species
Elongate narrow and convex; metallic green, shining, under
surface and legs shining black. Head a little smaller than pro-
thorax, surface rugose, the longitudinal impressions clearly out-
lined; epistoma truncate, suture distinct; mandibles black; max-
illary palpi stout, last segment truncate at apex; antennae thick,
black. Pronotum almost as wide as long, sides arcuate and finely
refiexed, surface with transverse rugulae; median stria deep;
scutellum short and broad. Elytra twice as long as broad, one-
fourth wider than pronotum, sides not quite parallel, a little wider
at two-thirds of their length; apex evenly rounded, margins very
finely refiexed; fianks with four or five impressed submarginal
punctures ; stria fine, not punctate ; intervals flat, unevenly creased,
without dorsal punctures ; subscutellar punctures distinct, the
short, feeble, and slightly oblique scutellar stria not strongly
marked. Anterior tarsi of male dilated spongiose beneath, inner
tibial tooth beyond the middle strong. Length of male, 15 mm.,
width at two-thirds its length, 4.5 mm.
Holotype male, allotype female, and two paratypes, collected
by the writer in Lincoln County, Oregon, on June 8, 1941.
The holotype, allotype, and a female paratype remain in the
author’s collection. A paratype male has been deposited in the
collection of Mr. E. R. Leach.
This fine species is dedicated to my son. It may be recog-
nized at once by its metallic green color.
JANUARY, 1944]
PRINCE— THRASSIS
13
DESCRIPTIONS OF THREE NEW SPECIES OF THRASSIS
JORDAN AND THE FEMALES OF T. BACCHI
(ROTHS.) AND T. PANSUS (JORDAN)^
BY FRANK M. PRINCE
Assistant Entomologist, United States Public Health Service
Among the fleas collected by the field forces of the Plague
Investigation Station, United States Public Health Service, in
the course of investigating the distribution of Plague in the
Western States, specimens of the genus Thrassis were obtained
which are considered representative of new species. Females of
T. hacchi and T. pansus which have not been described previ-
ously were also collected.
Thrassis setosis Prince, new species
Male and female. Frons with ocular row of 3 bristles, median
bristle small; one small bristle above ocular row in the female and
2 in the male. The postantennal region of the head has a row of
minute bristles along the border of the antennal groove in addition
to 1 large bristle and a posterior row of 1 large and 2 small
bristles. Labial palpi five-jointed and extending well beyond apex
of trochanter as in T. g. gladiolis (Jordan).
Pronotal ctenidium with 16 or 17 spines. Apical spines on
metanotum 2; abdominal tergites II, III and IV, 1 or 2.
Hind tarsal segment I equal to II, III and IV together.
Modified segments — Female. Antepygidial bristles 3, median
one longest. Head of receptaculum seminis globular; tail short,
broadest in distal half; similar to T. g. gladiolis. Sternite VII
without sinus, there are 8 or 9 bristles on each side (fig. 6). Stylet
one-half as broad as long and bears an apical bristle in addition
to 2 lateral bristles.
Modified segments — Male. Antepygidial bristles 1. Tergite
VIII with 8 or 9 large bristles along posterio-ventral border in
addition to 5 large dorsal bristles. Manubrium dilated distally
and truncated. Immovable process (P) of clasper dome-shaped,
shorter than movable finger (F). Movable finger convex on poste-
rior border and armed with 1 long bristle distally and 3 shorter
pale bristles more proximally; apex with straight cut along an-
terior face and concave near base of finger on anterior border.
Sternite' VIII broad with sclerified line near ventral edge; armed
with 4 or 5 apical bristles, 2 long subapical and 2 or 3 smaller
bristles. Internal portion of sternite IX broadest near apex or
dorsal end; proximal lobe broad and rounded with 3 large, flat-
tened, dark spines near apex in addition to 1 or 2 pale bristles;
^ From the Plague Investigation Station, San Francisco, California.
14
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 1
upper lobe broad with numerous pale bristles along posterior
border and minute bristles scattered over entire lobe on anterior
portion (fig. 5).
Length. Female, 2.4 to 2.7 mm., average 2.55 mm.; male,
1.96 to 2.07 mm., average 1.98 mm.
Type locality. Yuma County, Arizona.
Type host. Citellus harrisii saxicola (Meams) .
The holotype and allotype were collected 11 miles east of
Yuma, Yuma County, Arizona, April 6, 1938.
Paratypes. Yuma County, Arizona, 5 females and 1 male
from the type host. Yavapai County, Arizona, 42 females and
27 males from the type host and 2 females from Citellus vari-
gatus grammurus (Say), 2 females and 1 male from Neotoma
lepida stephensi (Goldman), 1 female from Peromyscus leuco-
pus arizonae (Allen), 2 females from Zapus. sp.
The holotype and allotype are deposited in the collection of
the LFnited States Public Health Service Plague Investigation
Station, San Francisco, California. Paratypes have been de-
posited at the University of California, Berkeley, California, the
California Academy of Sciences, San Francisco, California, and
the United States National Museum, Washington, D. C.
Thrassis aridis Prince, new species
Male and female. Frons with an ocular row of 3 bristles, the
median one small; two small bristles dorsal to ocular row. The
occiput bears along the border of the antennal groove a row of
minute bristles and a large bristle opposite about the center of
the groove and a posterior row of 4 bristles, the ventral one of
which is large. Labial palpi five- jointed, extending to apex of
forecoxae or slightly beyond.
The pronotal ctenidium has 15 to 17 spines. Apical spines on
each side of metanotum, 2; abdominal tergites vary but usually
I and III, 1, II, 2 in male; female I and II, 1. Tergites II to VII
with two rows of bristles.
Hind tarsal segment I shorter than II to IV together.
Modified segments — Female. Head of receptaculum seminis
longer than broad. Tail of receptaculum seminis long with tip well
sclerified. Sternite VII with sinus, upper lobe small and attenu-
ated (fig. 2). There are 7 to 10 bristles on each side of sternite VII.
Stylet approximately three times as long, as broad and has an
apical bristle in addition to 2 lateral bristles.
Modified segments — Male. Manubrium of process gradually
tapering from base to apex, curving slightly upward, resembling
T. fotus (Jordan), but not as pointed. Immovable process (P)
of clasper shorter than movable finger (F) with the apex pointed
JANUARY, 1944]
PRINCE— THRASSIS
15
posteriorly. The movable process nearly symmetrical and has 4
bristles on posterior border, basal 2 largest. Proximal lobe of
sternite IX with 4 pale bristles. Distal lobe with numerous minute
bristles. Parameres boot-shaped (fig. 1). Two long acetabular
bristles.
Length. Female 2.02 to 2.52 mm., average 2.17 mm.; male
1.35 to 1.87 mm., average 1.69 mm.
Type locality. Pima County, Arizona, and Santa Cruz County,
Arizona.
Type host — Dipodomys merriami. Holotype male was col-
lected 15 miles northwest of Nogales, Santa Cruz County,
Arizona. Allotype female was collected 13 miles north of
Tucson, Pima County, Arizona, February, 1939.
Paratypes. PiMA County, Arizona, 2 females; Cochise
County, Arizona, 1 male and 1 female; Pinal County, Ari-
zona, 3 females; Santa Cruz: County, Arizona, 2 male®;
Greenlee County, Arizona, 1 male; all taken on the type host.
In addition to those collected on the type host 2 males were
collected in Greenlee County, Arizona, from Neotoma lepida;
1 male was taken in Mohave County, Arizona, from Onychomys
leucoguster; 2 males were taken in Hidalgo County, New
Mexico, 1 from Onychomys leucogaster and 1 from Dipodomys
ordii.
The holotype and allotype are deposited in the United States
Public Health Service Plague Investigation Station, San Fran-
cisco, California. Paratypes are deposited in the United States
National Museum, Washington, D. C., and the University of
California, Berkeley, California.
Thrassis campestris Prince, new species
This species is closely related to Thrasssi aridis, except for
the modified segments.
Modified segments — Female. Tail of receptaculum seminis
without sclerified tip; shorter and broader than T. aridis. Sternite
VII with deeper and pigmented sinus (fig. 4). Stylet short, ap-
proximately one-half as broad as long, with an apical and 3
lateral bristles.
Modified segments — Male. Sternite VIII with 3 long and 4
short bristles. Proximal and distal lobes of sternite IX longer and
broader than in T. aridis. Movable process of clasper same gen-
eral shape as T. aridis but bears 7 bristles on posterior border.
Only 1 long acetabular bristle (fig. 4).
16
THE PAN-PACIFIC ENTOMOLOGIST
[VOL, XX, NO. 1
Length: Female, 2.40 to 2.85 mm., average 2.60 mm.; male,
1.95 to 2.47 mm., average 2.16 mm.
Type locality. Hooker and Grant Counties, Nebraska.
Type host — Dipodomys ordii. Holotype male was collected 7
miles west of MuLLEN, Hooker County, Nebraska. Allotype
female was collected 6 miles north of Ashby, Grant County,
Nebraska.
Paratypes. Grant County, Nebraska, 2 females and 5 males
from Dipodomys ordii and 1 male from Onychomys leucogaster;
Hooker County, Nebraska, 2 males from Dipodomys ordii and
2 females and 3 males from Onychomys leucogaster ; Cherry
County, Nebraska, 2 males from Dipodomys ordii; Garden
County, Nebraska, 1 female and 1 male from Dipodomys ordii
and 1 male from Peromyscus maniculatus.
In addition to the types and paratypes examined Mr. Glenn
M. Kohls of the Rocky Mountain Spotted Fever Laboratory,
Hamilton, Montana, sent me a large series collected by Mr.
Frank B. McMurry, United States Fish and Wild Life Service,
from Dipodomys in Harper County, Oklahoma.
The holotype and allotype are deposited in the United States
Public Health Service Plague Investigation Station, San Fran-
cisco, California. Paratypes are deposited in the United States
National Museum, Washington, D. C., and the Rocky Mountain
Spotted Fever Laboratory, Hamilton, Montana.
Explanation of Plate I
Fig. 1. Modified segments of T. aridis male. Fig. 2. Sternite
VII and receptaculum seminis of T. aridis female. Fig. 3. Sternife
VII and receptaculum seminis of T. campestris female. Fig. 4.
Modified segments of T. oampestris male. Fig. 5. Modified seg-
ments of T. setosis male. Fig. 6. Sternite VII and receptaculum
seminis of T. setosis female. Fig. 7. Sternite VII and receptacu-
lum seminis of T. hacchi female. Fig. 8. Sternite VII and recepta-
culum seminis of T. hacchi female.
Abbreviations: F, movable finger of clasper; P, immovable
process of clasper; Par, paramere; Rs, receptaculum seminis;
st. sternite; M, manubrium.
JANUARY, 1944]
PRINCE— THRASSIS
PLATE I
18
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 1
Thrassis bacchi (Rothschild) Jordan
Cerato'phyllus bacchi Rothschild, 1905, Nov. Zool. 12:159-160.
Thrassis bacchi (Roths.) Jordan, 1933, Nov. Zool. 39:73.
Female — Head. Oh the preantennal reg’ion there are three
bristles in the ocular row, median one small, in addition to 1 bristle
near the oral edge. Postantennal region has a row of minute
bristles along the antennal groove in addition to 1 large bristle.
Thorax. Pronotal ctenidium of 17 spines. Apical spines on
metanotum, 1 on each side.
Abdomen. Apical spines on tergites II, III and IV, 1 on each
side.
Modified segments. Receptaculum seminis and sternite VII as
illustrated (fig. 7). Tergite VIII armed with 17 to 21 bristles.
Stylet one-half as broad as long and bears an apical bristle in
addition to 4 lateral bristles.
Length. 1.50 to 1.84 mm., average 1.58 mm.
Allotype female collected 5 miles south of Gettesburg, Pot-
ter County, South Dakota, from Citellus tridecemlineatus
(Mitchill) . Additional specimens were taken in Potter and
Brown Counties, South Dakota, from Citellus richardsonii
richardsonii (Sabine) and in Spink County, South Dakota,
from Citellus franklinii (Sabine) ; in Rolette County, North
Dakota, from C. r. richardsonii; in Blaine, Phillips, Valley,
Daniels, Roosevelt and Sherman Counties, Montana, from
C. richardsonii.
Thrassis pansus (Jordan) Jordan
Ceratophyllus pansus Jordan, 1925, Nov. Zool. 32:109.
Thrassis pansus (Jordan) Jordan, 1933, Nov. Zool. 39:73.
Female — Head. Preantennal region with ocular row of 3 bris-
tles and 1 bristle dorsal to ocular row near oral edge. Postanten-
nal region wth a single row of minute bristles along border of
antennal groove in addition to 1 or 2 large bristles. Labial palpi
five- jointed, extending well beyond apex of trochanter.
Thorax. Pronotal ctenidium of 17 spines. Apical spines on
metanotum, 1 on each side.
Abdomen. Tergites II, III and IV 1 apical spine on each side,
occasionally 2 on III. Tergites II to VII with two rows of bristles.
Modified segments. Head of receptaculum seminis globular and
pigmented. Tail of receptaculum seminis broadest in distal half,
sclerified tip. Sternite VII as illustrated (fig. 8). Tergite VIII
bears 13 to 15 bristles. Stylet short, broad, with apical and 3
lateral bristles.
Length. 2.28 to 2.92 mm., average 2.66 mm.
JANUARY, 1944]
REES— MANSONIA PERTURBANS
19
Allotype female collected 14 miles northwest of Lordsburg,
Hidalgo County, New Mexico, from Onychomys leucogaster.
Additional specimens were taken in eastern Arizona, numerous
locations in New Mexico, and as far east as Brewster County,
Texas. The type hosts are evidently of the genus Citellus and
the subgenera Ictidomys and Ammospermophilus, whose range
includes the area outlined. Specimens were also collected occa-
sionally from Peromyscus, Dipodomys, Neotoma and Cynomys.
The writer wishes to express his appreciation to Dr. M. A.
Stewart, University of California, Berkeley, California, for his
suggestions and aid.
A NEW MOSQUITO RECORD FROM UTAH
(Diptera: Culicidae)
BY DON M. REES
University of Utah, Salt Lake City
In October, 1934, and April, 1942, the available mosquito
records for Utah were published by the author in this Jour-
nah. 2 . The following constitutes a new genus and species record
for the state.
Mansonia PERTURBANS (Walker)
Utah: Ogden, Weber County, July 8, 9, 10, 16, August 11, 27,
1943 (Rees).
Adults of this mosquito, both males and females, were taken
on the above dates on the Weber River directly south of Ogden
in what is known as Riverdale. No larvae were collected, but
conditions in the vicinity were suitable for their development.
Adults were collected at this locality in a light-trap, by hand
while feeding, and in rabbit hutches and other out-buildings. The
adults were fairly abundant but were more numerous in this
locality during July than in August.
The author was unsuccessful in finding this species in other
parts of Ogden or at several other localities along the Weber
River that were inspected for this purpose; but undoubtedly
this genus will be found to occur in other parts of the state as
this work is extended.
^ Rees, Don M. 1934. Mosquito records from Utah. Pan-Pac. Ent., 10 :161-165.
“ Rees, Don M. 1942. Supplementary list of mosquito records from Utah. Pan-
Pac. Ent., 18 :77-82.
20
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 1
NOTES ON CANTHARIS, 11
(Coleoptera, Cantharidae)
BY DOROTHY McKEY FENDER
McMinnville, Oregon
Cantharis dentatus Fender, new species
Black; thorax, head before the eyes, basal antennal joints be-
neath, and sides of abdomen testaceous; vestiture cinerous. Head
finely, closely punctate posteriorly, virtually impunctate in the
testaceous portion before the eyes; a median impressed line on the
occiput. Eyes moderately small; about one-fourth the length of the
head in the female, a little less than one-third the length of the
head in the male. Mandibles dentate; armed with a single strong,
rather blunt tooth on the inner margin of the apical third of each
mandible. Antennae slender, subcylindrical, those of the female
relatively shorter and stouter than those of the male; second joint
about two-fifths the length of the third in the male, about one-
half the length of the third in the female. Clypeus pyramidal, apex
acutely emarginate. Thorax subquadrate; anterior margin evenly
arcuate or with a very slight sinuation at apex (some females) ;
anterior angles not prominent, obtuse; posterior angles subacute
but blunt; disk nearly impunctate, with only a few fine scattered
hairs, these more abundant anteriorly; side margins bordered by
a definite, heavier marginal pubescence, blunt, especially ante-
riorly where the marginal pubescence is most abundant; rufo-
testaceous, a little darker on disk than at sides; thorax of female
relatively shorter than that of male and broadest anteriorly, mar-
ginal pubescence less evident. Elytra rugose, with a soft, cinere-
ous, primary pubescence, and stiffer, longer, black secondary
pubescence. Tarsi moderately stout; anterior tarsi of male short
and expanded. Anterior claw of protarsi of male cleft at tip,
toothed at base, anterior meso-and metatarsal claws of male cleft
at tip and with a blunt basal projection; anterior claws of all
tarsi of female cleft at tip, toothed at base. Male genital arma-
ture; (figs. 1 to 3). Last abdominal sternite of female swollen
either side of median line. Measurements: Length: Male overall
8 mm.; elytra 6 mm.; antennae 5 mm. Female overall 11 mm.;
elytra 7.5 mm; antennae 5 mm. Width: Male elytra 2.5 mm.;
thorax 1.5 mm. Female elytra 3.5 mm.; thorax 2.25 mm.
Holotype: male (Calif. Acad. Soi., Ent. No. 5404), Califor-
nia, C. W. Leng collection. Allotype: female (No. 5405), Ka-
weah, Tulare County, California, April 12, 1931, collected by
Roy Wagner. Specimens examined, five males and four females.
California: Ctto Lugger collection; three specimens Kaweah,
Tulare County, April 12, 1931, collected by Roy Wagner; Kern-
JANUARY, 1944]
FENDER— CANTHARIS
21
ville, May 7, 1931, collected by A. T. McClay; two specimens,
Sequoia National Park, June, 1937, collected by F. T. Scott.
This species has long been confused with Cantharis lautus Lee.
because of the fact that LeConte’s type series of C. lautus is a
species complex. Lautus was described in 1851, from a unique
female with yellow legs. Later, LeConte associated a black-legged
male with the type of lautus and subsequent references to C.
lautus were rather to this male. Thus, for many years, C. dentatus
has been determined as lautus, which is apparently a much rarer
insect.
Mr. C. A. Frost very kindly compared the types of C. dentatus
with LeConte’s type series of lautus and concluded that it is “the
same as the male associated with the type lautus” {in litt.) . The
Male genital armature of C. dentatus Fender. 1, dorsal;
2, lateral, and 3, ventral view.
female type of lautus does not have the last abdominal sternite
modified as in this species.
Lautus belongs with the notatus, perpallens, ingenuus, and
ochropus group, whereas dentatus belongs with divisus, which
also has the mandibles dentate. From divisus it may be easily
distinguished by the immaculate thorax, and cinereous vestiture
(golden in divisus), and from all other species by the ungual
formation. The blunt basal projection of the anterior claws of
the meso- and metatarsi of the males may, in some lights, almost
appear to be a definite tooth (as is found in divisus), but in no
case does it appear to be as acute as the basal tooth of the other
ungues. I have seen a few females of divisus having the thorax
imniaculate, but from these, dentatus females may be distin-
guished by the peculiar formation of the last visible abdominal
22
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 1
stemite. The pronotal pubescence in these species consists of
long, fine hairs each set in a very fine puncture.
Grateful acknowledgment is made to Mr. C. A. Frost for his
comparison of this species with the type of lautus Lee. and to
Mr. J. W. Green for the loan of the specimen from Kernville,
California, and for his having called the author’s attention to the
fact that this species has the mandibles dentate, as in divisus.
LIST OF SIX SPECIES OF GYRINID WATER BEETLES
COLLECTED BY J. L. GRESSITT IN CHINA AND JAPAN.^
(Coleoptera, Gyrinidae)
BY HUGH B. LEECH
Vernon, British Columbia
In 1938 some gyrinids were obtained from Mr. J. Linsley
Gressitt of the University of California, Berkeley. The beetles
were later submitted to Dr. G. Ochs, authority on the Gyrinidae
of the world, and his identifications are as follows:
Dineutus landaisi latilimbus Ochs.
Ta Han, Hainan Island, June 2, 1935.
Dwa Bi, Hainan Island, July 20, 1935.
Dinetus orientalis Modeer { = marginatus Sharp).
Takayama, Japan, August 5, 1930.
Iriomote Island, Loochoo Islands, August 20, 1934.
Dineutus mellyi Regimbart.
Mizuho, Formosa, April 22, 1932.
Tsin Leong San, E. Kwantung, S. China, June 3, 1936.
Yim Na San, E. Kwantung, S. China, June 16, 1936.
Ta Au Hong, S. Kiangsi Prov., S. China, July 5, 1936.
Gyrinus orientalis Regimbart.
Tsin Leong San, E. Kwantung, S. China, June 2, 1936.
Foochow, S. E. China, July 30, 1934.
Hong San, S. E. Kiangsi Prov., China, June 26, 1936.
Orectochilus nigroaeneus Regimbart.
Yim Na San, E. Kwantung, S. China, June 14, 1936.
Orectochilus melli Ochs.
Tsin Leong San, E. Kwantung Prov., S. China, June 5, 1936.
^ Contribution No. 2264, Division of Entomology, Science Service, Department
of Agriculture, Ottawa, Ont.
JANUARY, 1944]
MALKIN— STAPHYLINIDAE
23
NEW SPECIES OF BOLETOBIUS FROM THE NORTHWEST
WITH A DISTRIBUTIONAL NOTE ON COPROPORUS
(Coleoptera, Staphylinidae) *
BY BORYS MALKIN
Eugene, Oregon
During the course of an attempted monographic study of the
Northwestern forms of the Staphylinid genus of Boletohius, sev-
eral new species came to the writer’s attention. While the whole
work as originally conceived may not be completed for some
time, due to present conditions, it seemis worthwhile to present
at least the descriptions of these new forms.
Boletohius kremeri Malkin, new species
Dark, piceous beneath, femora and tarsi yellow; head black,
highly polished glabrous; maxillary and labial palpi pale yellow;
third joint of the former short, pubescent. Thorax black; elytra
red; abdomen piceous except last segment and posterior portion of
the penultimate one which are red. Antennae: two basal joints
pale yellow, 7-11 dark piceous. Joints 8-10 transverse. Thorax
highly polished with setae bearing punctures on each side. Elytra :
with three irregular dorsal rows of punctures with seta in each.
The lateral row with 10 punctures, the middle with 9, some of
which are obscured, the inner one with 7. Lateral elytral margins
with a row of small spinules. Abdomen: black. First two seg-
ments polished on disc punctured on sides; the remainder of the
segments with a long hairs. Punctures of the sixth segment very
coarse, prolonged; fifth segment nearly twice as long as sixth
anterior half distinctly darker. Margins of the segments with a
long spinules increasing in size toward apex. The broad, lateral
margins of the segments decreasing inward toward the apex, com-
pletely absent on the last segment. Ventral surface black. Margins
of the epipleura castaneous. Legs dark yellow slightly paler to-
ward the tarsi. Posterior half of the fifth segment and the entire
sixth chestnut. Punctures very coarse prolonged. Trochi of the
first pair of legs piceous. Femora with a dark cloud. Trochi,
femora, tibia sparely punctured and muricate. Abdomen with the
long fimbria along the sides and on the lateral portion of the seg-
ments, the largest at the posterior angle. Length, 8.2 mm.; width,
2.3 mm.
Male. Fifth segment with a small granulated area, feebly
emarginate. Sixth, deeply, longitudinally impressed in the middle.
Female. Last segment simple.
Holotype, male, Eugene, Lane County, Oregon, March 29,
1942 (B. Malkin & John S. Sadler). Allotype, female, BoYER,
* A contribution from the Zooloeical laboratory of the Unirersity of Oregon.
24
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 1
Lincoln County, Oregon, April 25, 1936 (K. M & D. M. Fen-
der). Paratypes: 1 male, Glenada, Lane County, Oregon,
September 14, 1941 (collected by the author from Boletus sp.).
One male, BoYER, Lincoln County, Oregon (K. M. & D. M.
Fender), 1 male, April 11, 1936, same locality (K. M. & D. M.
Fender). One female, labeled; Seattle, Washington, without
any other data, from Hatch collection.
The type material is distributed as follows: holotype, allotype
and Glenada paratype in author’s collection. Two paratypes in
Fender’s collection. The Seattle paratype in Dr. Hatch’s collec-
tion.
This species runs in Horn’s key to intrusus from which it
differs having only 8-10 antennal joints transverse, 3 discal rows
of punctures instead of one, coarser puctulation on the dorsal
side of the abdomen and different coloration of the last two
segments. It is also considerably larger than Horn’s species.
This species is named after Mr. Jean H. Kremer of New York
City, with whom the writer has spent many days in the field.
Boletobius ruber Malkin, new species
Reddish, shining, legs testaceous. Head: black, moderately
elongate, highly polished, widest at base. Maxillary palpi testa-
ceous; fourth joint short, conical, shorter than third; the first
three sparsely pubescent. Antennae, as long as thorax and head.
Basal joints testaceous, 5-11 piceous. Joints 6-10 very strongly
transverse. Thorax highly polished, wider than long, with sides
gently rounded. Disc of thorax reddish-piceous, margins reddish-
testaceous, median line feeble. Elytra yellow-reddish slightly
longer than wide, polished, very minutely, transversely alutaceous.
The black spot in the posterior half not attaining the suture or the
apical margin of the elytron. Elytral rows of punctures distrib-
uted thus: discal with 5 distinct, sutural with 7 somewhat less im-
pressed, reaching the sides of the scutellum, lateral with 8-10.
Abdomen red, except last two segments which are darker. Apical
portion of all segments paler, all deeply impressed near the ante-
rior margin. Surface polished very minutely alutaceous, deeply
and distinctly punctured, with yellow short hairs throughout and
longer dark spinules in the lateral portion of the last two seg-
ments. Undersurface reddish-piceous. Episterna sparsely, mi-
nutely punctured, with a tiny hair in each puncture. Abdominal
segments, coarsely, unequally punctured. Apical portion of tibia
with short, unequal spinules. Length, 5 mm.; width, 1.3 mm.
Male. The penultimate segments slightly, narrowly impressed
in the posterior half. Last segment with granulated area divided
in apical half.
Female. Last segment simple.
JANUARY, 1944]
MALKIN— STAPHYLINIDAE
25
The coloration of the last two segments varies in the speci-
mens on hand from reddish to piceous, but the two last are always
darker than the others. The punctulation also varies slightly in
the intensity, usually and stronger on the last two segments.
Figure 1
Boletobius kremeri Malkin, left; B. ruber Malkin, right.
Holotype, male and allotype, female. No. 5267, Aspen Grove,
B. C., 22. V. 1934 (collected by H. B. Leech under bark of Pinus
ponderosa ) , in the Canadian National Collection. Paratypes: 2
males and 1 female the same data; 1 male, 3.V.1934, 1 female
7.V.1934, 1 female 14.V.1934, all from the same locality (H. B.
26
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 1
Leech, collector). One male, Penticon, B. C., 4.V.1936, (Mc-
Donald, collector) . One female, Winnipeg, Manitoba, 4. VI.
1912 “from under bark of poplar tree at night” (J. B. Wallis,
collector) .
Paratypes deposited as follows: 1 in the collection of the Cal-
fomia Academy of Sciences, 1 in J. B. Wallis’ collection. The
remainder equally distributed between the collections of Mr.
H. B. Leech and that of the author.
This species seems to have its closest relative in dimidiatus
Er. from which it can be easily separated by reddish color of the
entire body, deeper punctulation of the abdominal segments,
joints 6-10 strongly transverse, more distinctly impressed sutural,
elytral punctures and thorax more broadly margined with paler
band. In Horn’s key it should be placed between dimidiatus and
intrusus.
Boletobius kelleyi Malkin, new species
Color in general reddish -testaceous. Head black, twice as long
as wide, highly polished with sparse, minute punctures. Maxillary
palpi yellow, strongly prolonged. Fourth joint as long as the third.
First minutely pubescent, second and third with tiny hairs in
their apical portion. Antennae: four basal joints testaceous, 5-11
piceous, all longer than wide. Thorax wider than long, regularly
curved on sides, highly polished reddish-testaceous in color with
disc somewhat clouded and paler margins. Median line short,
feeble. In the front near the anterior margin two small, triangular
spots on each side. Elytra as long as wide. Two spots and trans-
verse area around scutellum black. Dorsal series of elytral punc-
tures with 6, sutural with 7 less distinct. Lateral row with 10
punctures. Abdomen reddish-castaneous except for the anterior
portion of the fifth segment which is black. Segments feebly im-
pressed and very sparsely punctured. Pubescence sparse, yellow.
Hairs longer on the lateral portion of the segments. Lateral
spinules short. Ventral surface: legs, femora, tarsi, tibia testa-
ceous. Epipleura black, similarly punctured as in canadensis.
Episterna alutaceous with large, shallow punctures. Abdominal
segments similarly colored as on the dorsal side, but strongly and
closely punctured. The punctures feebler and sparser on the last
two. Length, 6.5 mm.; width, 1.9 mm.
Type No. 5266, Trinity Valley, B. C., 26.VL1937 (H. B.
Leech, collector), and 1 paratype, Langley, B. C., 26.VL1932
(K. Broham, collector), both of undetermined sex in the Cana-
dian National Collection.
In the Horn’s key B. kelleyi runs to longiceps Lee. It can be
distinguished from the latter species by: much paler coloration
of the entire body with only small portion in the apex black.
JANUARY, 1944]
MALKIN— STAPHYLINIDAE
27
presence of the two dark spots on thorax and the presence of the
distinct black area around scutellum.
Variation: In the paratype the abdominal segments are slightly
darker, while the scutellar area forms a triangle, whose lateral
sides do not reach the margins of the elytra.
Figure 2
Boletobius cascadensis Malkin, left; B. kelleyi Malkin, right.
The species is named after Mr. Donald Greame-Kelley of New
York City.
Boletobius cascadensis Malkin, new species
Head oval, highly polished, slightly longer than wide. Maxillary
palpi yellow. Second joint as long as the third. The first three
28
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 1
pubescent. Antennae nearly as long as head and thorax. Basal
joints testaceous, 5-11 piceous. Joints 6-10 slightly longer than
wide, cup shaped. Thorax wider than long, regularly rounded,
yellow, with a median dark line, and a well defined dark spot on
each side of it. Surface polished, very minutely punctured, and
almost invisibly, transversely alutaeceous. Elytra as long as wide,
with a large variable black spot. The black area not attaining the
sutural line or the posterior angles. Scutellum surrounded by the
triangular black area. Disc with a series of feeble impressions.
Dorsal series with 9 feeble, setae bearing punctures. Sutural with
10 still weaker, lateral width 7-9. Abdomen piceous with tips of
segments paler. The yellow area on the fifth segment broader than
on the rest, occupying almost four-fifths of the entire segment.
Surface polished, alutaceous, sparsely punctured. Pubescence yel-
low, short. Lateral spinules short. Ventral side: mesosternum and
episterna pitchy black. Epipleura black, with a row of regular,
deep punctures in its anterior portion. Side pieces of metasternum
regularly reticulate with a delicate, dark pubescence. Abdomen
similarly colored as on the dorsal side, closely regularly punctured.
The punctures prolonged, deep and distinct. Length, 6 mm.; width,
1.7 mm.
No secondary sexual characteristics determined.
Type and 3 paratypes, Scott Lake, one mile north of Mc-
Kenzie Pass, Lane County, Oregon, altitude 4,800 feet, August
17, 1941, taken by the author from decaying Boletus sp. 1 para-
type, Granite, Baker County, Oregon, June 15, 1941, K. M. &
D. M. Fender, collectors.
Type and 2 paratypes in author’s collection, 1 paratype in
the collection of the California Academy of Sciences, 1 paratype
in Fenders collection.
This species is very closely related to B. anticus Horn and it
has probedily been mistaken for that species in collections. It
can be separated from the latter by having the thoracic spots
posteriorly located instead of anteriorly, much broader yellow
area on the fifth abdominal segment, and a very coarse punctu-
lation on the ventral side of the abdominal segments.
The specimens on hand exhibit very little variation except for
the extent of the elytral spot and the number of punctures in the
lateral row. The dorsal row is quite uniform as well as the ex-
tent of the thoracic spots.
Boletobius cascadensis hatchi Malkin, new subspecies
Head oblong oval, black, shining. Maxillary palpi yellow all
joints of nearly equal length. First two delicately pubescent.
Thorax slightly wider than long; red, with disc indefinitely
JANUARY, 1944]
MALKIN— STAPHYLINIDAE
29
clouded, polished, very delicately, transversely alutaceous. Elytra
as long as wide, polished. The black spot variable in size some-
what posteriorly located, not attaining the suture. Dorsal series
irregular, usually of 9 punctures, sutural of 10, lateral 8-10, irreg-
ular but distinct. Abdomen piceous, segments broadly yellow in
the apical, posterior portion. Yellow area of fifth segment occu-
pying the entire apical half. Punctures sparse but distinct with
the golden pubescence. Surface transversely alutaceous. Lateral
spinules short. Ventral side: femora, tarsi, tibia red-testaceous.
Epipleura black, except in the apical portion where they are
reddish-testaceous. Mesosternum and metasternum as in kelleyi.
Color the same as on the dorsal side. Segments very coarsely punc-
tured, thickly pubescent. The punctures on the last two segments
feebler toward the apex. Length, 3.5 mm.; width, 1.25-1.6 mm.
Type and 26 paratypes of undetermined sex collected by M. H.
Hatch on Mr. Bonaparte, Washington, July 12, 1931.
Type and 17 paratypes in Hatch collection. Four paratypes in
author’s collection. One paratype each in the collections of :
California Academy of Sciences, Oregon State College, K. M. &
D. M. Fender, C. A. Frost, and H. B. Leech.
The series studied vary greatly in size and in coloration, par-
ticularly in the extent of the elytral spot and in coloration of
thorax. In some specimens the latter is almost piceous on the
disc, but the margins are usually more pale in all specimens.
This form differs from cascadensis by having the epipleura
black in the posterior half, more conspicuous pubescence on
maxillary palpi, absence of a definite pattern on thorax and con-
siderably smaller in size and more robust body.
B. cascadensis subsp. hatchi is undoubtedly, like the original
form, mistaken in the collections for B. anticus. In Horn’s key
it should be placed between cascadensis and pygmeus Fab. At
present time the writer considers hatchi as a subspecies but it is
quite possible that in the future it will be separated as a distinct
species.
Boletobius intrusus wallisi Malkin, new variety
This form resembles the original form very closely except for
the following differences: Smaller size, abdomen of much paler
color and more coarsely punctured surface of the abdominal
segments. Length, 5 mm.; width, 1.5 mm.
Holotype, male and allotype, female. No. 5268, WINNIPEG,
Manitoba, 27. VI. 1915, in the Canadian National Collection, both
collected by Mr. J. B. Wallis.
30
THE PAN-PACIFIC ENTOMOLOGIST [vOL. XX, NO. 1
COPROPORUS VENTRICULUS (Say)
In Dr. Hatch’s collection of Tachyporini a specimen of Copro-
porus ventriculus (Say) has been found taken at Pierce, Idaho,
on July 27, 1939. This is apparenty a new record for the state.
Blackwelder in his revision of the genus* lists Oregon, Nevada,
Washington and California and several Eastern and middle-
western states as habitat of the species, but does not mention any
Idaho records. There are also in the same collection 16 old
specimens labeled “Seattle, fungus” which confirms its presence
in that state. In spite of the literature records, this only represen-
tative of the genus seems to be not common in the Northwest.
The present writer has never been able to find any in Oregon,
nor was he able to discover any in the Oregon collections as well
as in other collections from the Northwest which he examined.
Acknowledgments: In addition to writer’s own collection the
following individuals contributed material for study through
loan: Mr. H. B. Leech, Mr. J. B. Wallis, Dr. Melville H. Hatch,
Professor James N. Macnab, Mr. and Mrs. Kenneth Fender, and
Mr. L. G. Centner. Thanks are also due to Mr. R. L. Post, cura-
tor of the entomological collection at Oregon State College, for
the loan of certain material in his custody and to Mr. J. L. Shuh
for the loan of the Boletohius material from his collection. The
writer is also indebted to Dr. W. J. Gertsch of the American
Museum of Natural History, to Dr. D. C. Mote of the Oregon
State College, and to Mr. H. B. Leech for the help with certain
literature, and to Miss Edith Onthank, a student at the University
of Oregon, for drawings and assistance.
Spring Flight of Atimia dorsalis (Coleoptera,
Cerambycidae)
Atimia dorsalis LeConte is common in southern British Co-
lumbia, where it breeds in western red cedar. Thuja plicata. It
is the first longhorn to appear on the wing each spring ,and
many individuals are abroad by mid-April. My earliest record
is for March 20, 1932, on which date my brother, Geof. B. Leech,
took a specimen flying, at Salmon Arm. — Hugh B. Leech,
Vernon, B. C.
* R. E. Blackwelder. 19S8. ‘'Reviaion of the N. A. Coproporaa.” Proc.
U.S.N.M. 86, 1-10.
JANUARY, 1944]
POTTS— ACRAEINAE
31
SYSTEMATIC NOTES CONCERNING AMERICAN
ACRAEINAE, II
( Lepidoptera : Nymphalidae)
BY ROBERT W. L. POTTS
University of California, Berkeley
This article, with a previous one^ makes available the princi-
pal changes and additions to the genus Altinote resulting from
preliminary studies for a proposed revision of the American
Acraeinae.
I should like at this time to express my grateful appreciation
to E. Gorton Linsley; to Wm. T. M. Forbes for the loan of ma-
terial from the Cornell University collection; to Lawrence S.
Dillon for the loan of material from the collection of the Read-
ing Public Museum; and to Charles D. Michener for the loan of
material from the collection of the American Museum of Natural
History.
Altinote (Abananote) carmentis Potts, new species
Male. Dull black with faint whitish gloss. Forewing above
marked only by red-orange streak in subcostal cell similar to that
in erinoTYie, and by a series of five light buff spots beyond cell be-
tween veins, traces of buff near anal angle and above radius; first
three spots nearly equal-sized, ovate; fourth much larger, sub-
triangular with long point toward outer margin; fifth smaller,
elliptical. Hindwing unmarked.
Underneath buffy-grey with first four buffy spots coalesced in
a band, fifth spot slightly separated; orange subcostal streak re-
peated; apex with faint, even intervenal stripes. Hindwing evenly
striped, slightly darker at outer margin.
Forewing (base to apex) 23 mm., total expanse 43 mm.
Holotype male from Moyobamba region, Peru, December 15,
1925, from the Bassler collection in the American Museum of
Natural History.
Very similar in appearance to the female of euryleuca, but
the band of spots entirely beyond the discal cell by about one-
half the length of the first of the buff spots. Shape and size, as
well as the red-orange stripe, are suggestive of erinome.
^ Potts, Robert W. L., Systematic Notes Concerning American Acraeinae, Pan-
Pac. Ent., 19(1) :31-32, January, 1943.
32
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO, 1
Altinote (Altinote) alcione mengeli Potts, new subspecies
Male. Brown-black, more to black; similar to sodalis except
forewing with basal area extended outwardly, orange, the orange
apical band very narrow and not above radials, of three or four
obsolescent spots; dark discal band widened, directed toward point
slightly above anal angle.
Similar underneath; hindwing with color striping much re-
duced, yellow-orange at base ; greyish band barely apparent on disc ;
striping between dark venal-intervenal stripes mainly brownish.
Forewing 26.0 mm., expanse 49 mm.
Holotype male from Jepelacis, Peru, without further data,
in the Mengel collection of the Reading Public Museum. One
paratype from Jepelacis and another from RiO HuALLAGA, AcHl-
NAMIZA, Peru, also in the Reading collection. Two paratypes
in the author’s collection, one from Jepelacis and one from Inca
Pechetea, Peru.
Similar to sodalis, but distinguished from it by the obsolete
apical band, the heavier discal band, and the greater extent of
the basal area. Variation of forewing in the paratype series is
from 22.5 mm. to 26.0 mm.
Altinote (Altinote) alcione bassleri Potts, new subspecies
Male. Dark brownish. Forewing above with basal area pale
orange, almost without reddish tone, extended to include lower
angle of cell and spot in cell M 3 , with less than normal basal in-
fuscation; apical band scarcely indicated; discal band not black,
but dark brownish, only barely perceptible. Hindwing without
marking.
Forewing beneath similar to upperside, weakly striped in costal
area, hence this more a dark stripe than normal, color striping in
apex nearly absent; apical band very narrow, entirely between Rs
and M2 or M3. Hindwing beneath very weakly striped, orange
virtually absent, greyish band barely, if at all, apparent.
Forewing 22.5 mm., expanse 42 mm.
Holotype male and three paratypes from Moyobamba REGION,
Peru, with no additional data; holotype and one paratype in
the Bassler collection of the American Museum of Natural His-
tory, the other two paratypes in the collection of the author.
Similar in appearance to mengeli, but conspicously smaller,
paler, and with somewhat more striped appearance beneath. Size
variation of forewing from 20.5 to 22.5 mm. in the short series
available for study.
JANUARY, 1944]
POTTS— ACRAEINAE
33
Altinote (Altinote) alcione moneta Potts, new subspecies
Male. Dark brownish-black; forewing above closely similar to
mengeli, but basal area red-orange, not so extended and scal-
loped at veins at outer margin; apical band absent; discal band
directed to well above anal angle.
Beneath similar; discal band almost parallel to inner border in
first half thence sharply angled toward anal angle, barely exceed-
ing Cu 2 , not extending into normal dark outer border ; apical band
very narrow, yellowish at outer edge; apex weakly striped, ground
tone blackish. Hindwing more regularly striped than other alcione
forms, yellow at base not strong, orange tinted, nebulous dark
area at end of cell barely apparent, greyish discal band weak,
tinted orange; dark outer border not strong; ground tone strongly
blackish.
Forewing 26.0 mm., expanse 45 mm.
Holotype male, in the collection of the American Museum of
Natural History, from 2-4° S, 78° W., Ecuador. Paratype with
similar data in the author’s collection.
Altinote (Altinote) alcione hermosa Potts, new subspecies
Male. Brown-black, closely similar to sodalis, with normal
basal area, medium discal band, comparatively extensive apical
band, but color of basal and apical bands rose-orange.
Forewing 24.0 mm., expanse 45.0 mm.
Holotype male from Hermosa, 1600 M., Peru, May 6, 1935,
in the author’s collection.
Easily distinguished from all but the following subspecies
and incarum by the rose color; distinguishable from incarum
by the presence of the apical band, and from the next subspecies
by the opacity of the colored areas.
Altinote (Altinote) alcione leucothoe Potts, new subspecies
Male. Similar to hermosa, except apical band longer, discal
band somewhat more curved, directed more toward anal angle;
basal area and apical band rose-orange, more rose than hermosa,
and basal area in particular semi-transparent.
Forewing 24.0 mm., expanse 45.0 mm.
Holotype male and seven paratypes from region of Iquitos,
Peru, according to data received. 1 should not be surprised,
however, if the locality were farther to the west.
Paratype variation exhibits a widening of the dark discal
band, some variation in transparency from a specimen in which
the pin label may be read through the basal area to translucence.
34
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 1
The specimens are not rubbed, but fresh and in excellent condi-
tion. Easily distinguished from hermosa by this transparency.
In my opinion this series of alcione subspecies are more nearly
genetic varieties, but without a considerably greater amount of
material to study it seems better to conform to the usual prac-
tice and give names to the recognizable forms. It is my belief
that it will be found that virtually every range and valley from
Colombia to Bolivia will exhibit a slightly different alcione
form readily distinguishable by the student from series collec-
tions, but difficult to describe or key out.
Altinote (Altinote) ozomene alicunde Potts, new subspecies
Male. Black with green irridescence over entire ground surface;
larger than typical ozomene, otherwise similar.
Forewing 28.0 mm., expanse 54.0 mm.
Holotype male and two paratypes in the author’s collection,
from Colombia, probably taken in February or early March,
locality unknown, but believed to be near Queremal or Buena-
ventura, altitude about 3000 to 4000 meters.
The principal differences between this and the typical are in
the definitely greenish irridescence, whereas the typical usually
runs to purplish blue, never more than slightly greenish blue,
and in the considerably larger average size, one paratype being
slightly smaller, the other slightly larger than the type.
Altinote (Altinote) stratonice albunea Potts, new subspecies
Male. Similar to aereta, but with dark discal spots heavier
above and forming an entire band both above and beneath; basal
colored area almost eliminated by dark suffusion from base. Basal
area beneath without suffusion. Disco-apical band light orange
above, huffy underneath.
Forewing 24.0 mm., expanse 44.0 mm.
Female. Similar but larger, grey-black, with colored area prox-
imal to transverse dark band clearly indicated, distally buffy-
yellow. Beneath, forewing proximal area buffy brown-orange,
distally buffy-yellow; costal area uniformly striped.
Forewing 28.5 mm., expanse 54.0 mm.
Holotype male and allotype female from RiO JONDACHI, near
Archidona, 800 M., Ecuador, collected by F. Martin and Hazel
Brown in November, 1939. In the author’s collection.
This is the most extremely marked of all the stratonice sub-
species known to me.
PACIFIC COAST ENT. SOCIETY
35
PACIFIC COAST ENTOMOLOGICAL SOCIETY
M. A. Stewart
V ice^President
H. H. Keifer
President
E. G. Linsley
Secretary
Proceedings
One Hundred and Seventh-seventh Meetim
The one hundredth and seventy-seventh meeting of the Pacific
Coast Entomological Society was held at 2:30 p.m. on February
27, 1943, in the entomological laboratories of the California Acad-
emy of Sciences, San Francisco. President Keifer in the chair.
The following members were present: H. H. Keifer, E. G. Linsley,
R. C. Miller, A. E. Michelbacher, C. D. Duncan, M. A. Stewart,
E. 0. Essig, T. F. Kelley, R. W. L. Potts, R. F. Smith, and W. Y.
Chong. Visitors were present as follows: F. M. Prince, G. L.
Smith, 0. G. Bacon, L. L. McRae, S. Vrana, F. M. Frost, R. King,
T. Algard, J. Guttseburg, C. Madsen, H. Madsen, P. H. Arnand,
W. K. Brainerd, M. Gutterman, B. Prendergast.
The minutes of the previous meeting were read and approved.
The secretary reported recent action of the Executive Board
in amending Article VI, Sec. 4, of the By-laws to permit the sale
of 20 life memberships in 1943 on a budget payment plan extend-
ing over not more than five years and including payments of not
less than $10.00 per annum.
Professor Essig reported that the Publication Committee had
appointed Dr. Linsley as editor of the Pan-Pacific Entomologist to
fill the vacancy left by Dr. Usinger, now in the armed forces.
The membership committee proposed the following names: Carl
J. Drake, F. M. Prince, and Harold Madsen. They were duly
elected to membership.
Dr. Duncan exhibited photographs of the larviform females of
a phengodid beetle, as well as a slant-faced locust and lined June
beetle (Polyphylla) . He also announced the completion of a key
to the butterflies and skippers of Santa Cruz County.
Dr. Linsley discussed retarded development and delayed emer-
gence of Buprestis aurulenta Linn, in structural timbers. He sum-
marized 12 cases ranging from 9 to 26 years, O’bserved in Califor-
nia over a five-year period. He pointed out that the location of
the infestations in the buildings, geographical areas involved, and
general habits of the species all argued that the infestations orig-
inated in the lumber yard (or possibly the forest) and existed
prior to construction.
The president then presented Dr. Linsley, who read a paper
entitled “Natural habitats, sources, and reservoirs of insects asso-
ciated with stored food products.” Dr. Linsley stated that these
36
THE PAN-PACIFIC ENTOMOLOGIST [yOL. XX, NO. 1
insects shared a number of characteristics which combined to make
them pests. (1) They have a wide and more or less cosmopolitan
distribution. (2) Most of the species reproduce more or less con-
tinuously when conditions are favorable, a fact which renders them
independent of seasonal and daily cycles. (3) Their small size
makes them easy to overlook, making" it possible to transport them
unnoticed from place to place and to fail to discover their presence
until a population reaches injurious numbers. (4) Some species
are able to survive relatively long periods without food enabling
them to endure unfavorable food conditions as well as increasing
their ability to survive transportation. (5) Most stored food prod-
ucts insects have a wide range of food habits and many include the
majority of dry food and drug products of plant and animal
origin in their diet. This food range is greater than that generally
attributed to most other insects. (6) Many of the species appear
to have a wide range of tolerance for physical factors in the en-
vironment, especially humidity. Because in any given environment
the extremes may be more significant than the means, it is likely
that these insects can tolerate more extreme conditions of tempera-
ture and humidity than many less widely distributed forms. This
is particularly evident with regard to the moisture content of food
materials and most of the important species can survive in food-
stuffs with a moisture content far below the optimum. (7) Finally,
although these insects have a broad tolerance for extremes of
humidity (and frequently temperature), man maintains suitable
(probably optimum) conditions for most of the species in many
buildings, warehouses, and food storage units.
Dr. Linsley then discussed the original habitats and natural
reservoirs of several hundred species associated with stored food
products, from which the following conclusions were drawn:
(1) Stored food products insects are widely distributed in
nature as well as in the warehouses, granaries, and households of
mankind. (2) Several of the species involved (bruchids, Sitophi-
lus, Sitotroga, etc.) are seed infesting species which infest these
same materials in nature. These insects have become pests with-
out having to undergo further evolution of food habits. (3) A
large number of insects (cryptophagids, mycetophagids, lathri-
diids, acarids, etc.) associated with stored food products are actu-
ally fungus feeders and their presence is due to molds and fungi
existing in the foodstuffs. In nature these species are found under
bark, in nests of birds, mammals and insects, in decomposing plant
materials, any place where molds and fungi are present. (4) Many
food products pests are general scavengers on dead plant mate-
rials. The majority of these are moths (phycitids, pyralids, etc.)
but a few other groups (Anobiids, nitidulids, etc.) are involved.
(5) A similar source is found in the scavengers on dead animal
materials. Included are Dermestes, Piophila, Tineola, etc. (6) A
very important natural source of stored products pests is among
the scavengers or semi-predators living under bark. To this source
JANUARY, 1944]
PACIFIC COAST ENT. SOCIETY
37
we must credit almost all of the tenebrionids (flour beetles, meal-
worms, etc.), cucujids, ostomatids, etc., and to a wood boring
origin, the ptinids, anobiids, bostrichids, and scolytids. (7) Insect
nests have also offered a number of pests. Several of the gal-
leriids, phycitids, ptinids, and dermestids, in particular, may have
come from this source. The pollen stored by bees may have had
an important role in the evolution of the food habits of some of
the species by serving as a bridge between zoophagy and phyto-
phagy. (8) Many of the insects in stored food products are preda-
tors and parasites of other insects present. A few of these, espe-
cially histerids, clerids, anthocorids, etc., are probably attracted to
the products and habitat directly rather than by any specific in-
sects present. (9) Primary natural reservoirs are to be found in
the following habitats: (a) bark and decomposing wood, (6) nests
and food caches of other insects, especially bees, wasps, and ants,
and (c) nests of birds and mammals.
After a discussion of this paper, the meeting adjourned. — E. G.
Linsley, Secretary.
One Hundred and Seventy-eighth Meeting.
The one hundred and seventy-eighth meeting of the Pacific
Coast Entomological Society was held at 2:30 p.m. on May 27,
1943, in the entomological laboratories of the California Academy
of Sciences, San Francisco. President Keifer in the chair. The fol-
lowing members were present: H. H. Keifer, E. G. Linsley, E. C.
Van Dyke, E. 0. Essig, M. A. Stewart, A. E. Michelbacher,
R. W. L. Potts, G. L. Smith, R. F. Smith, F. M. Prince, R. C.
Miller, P. Moorhead, and P. A. Harvey.
The minutes of the previous meeting were read and accepted.
President Keifer announced the appointment of Dr. M. A. Stew-
art to the publication committee and Dr. A. E. Michelbacher to the
program committee.
Mr. R. F. Smith reported that the alfalfa butterfly, Colias
eurytheme Bvd., and lygus bugs, Lygus spp., were abundant on
alfalfa in the San Joaquin Valley, indicating the possibility of
damaging populations in that area later in the season.
President Keifer then presented Dr. Van Dyke, who addressed
the Society upon “Entomological Type Localities on the Pacific
Coast.” Dr. Van Dyke confined his discussion largely to those
localities which have had their names changed or which have been
obliterated from various causes. He suggested that a list of such
places should be compiled and published in order that future
workers might know where they were. Hetch Hetchy Valley was
destroyed as a collecting grounds when it was turned into a reser-
voir; so also was Crystal Springs in San Mateo County, a type
locality for many of G. R. Crotch’s species. Other places, like
some of the old frontier forts vanished when the government
abandoned them. Many of those near the larger communities like
Los Angeles and San Francisco were destroyed by advancing cities.
38
THE PAN-PACIFIC ENTOMOLOGIST [vOL. XX, NO. 1
The localities listed by the early Russians in Alaska and Cali-
fornia are mainly easily placed since they are either existing com-
munities like Unalaska, Sitka, and San Francisco, places like Fort
Ross (Colony Ross) and Sutter’s Fort (New Helvetia) which have
been preserved as historical monuments, or stable landmarks like
Point Reyes and Mt. St. Helena. Many of the J. L. LeConte locali-
ties of the eighteen fifties are cities like San Diego or San Jose,
others like Isabella are small places which, however, still retain
their names. The famous G. H. Horn localities of Civil War days
are Fort Independence in Owens Valley, with the fort destroyed
but the site preserved by the town of Independence which has
grown up around it, and Fort Tejon, now an historical monument.
A few of the other noted localities mentioned by Dr. Van Dyke
were Clarks’ Corners, now the famous resort Wawona; Tallac, at
the lower end of Lake Tahoe, now no longer a resort; Havilah,
long past its zenith but still maintaining a post office; Sissons,
now called Shasta City, and Soda Springs, now called Upper Soda
Springs, in the upper Sacramento Valley. Smaller suburban col-
lecting sites have suffered greatly. Redwood Canyon in Marin
County has become Muir Woods, Laguinitas Canyon is a water
district and closed to the public. The main San Francisco collec-
ting ground of Henry Edwards and Behr is now the heart of
Chinatown; Lone Mountain is occupied by the Catholic Women’s
College; Mountain Lake is in the Presidio and closed to the public,
and Lake Merced scarcely remains as a collecting ground. The
famous Sylvania was Ricksecker’s ranch, near what is now Camp
Meeker, and Camp Taylor on Paper Mill Creek in Marin County,
is preserved as a picnic ground.
Following a discussion of Dr. Van Dyke’s paper, the meeting
adjourned. — E. G. Linsley, Secretary.
One Hundred and Seventy-ninth Meeting.
The one hundred and seventy-ninth meeting of the Pacific Coast
Entomological Society was held at 2:15 p.m. on September 25,
1943, in the entomological laboratories of the California Academy
of Sciences, San Francisco. President Keifer in the chair. The
following members were present: H. H. Keifer, E. G. Linsley,
C. D. Duncan, E. C. Van Dyke, M. A. Stewart, F. M. Prince, F. J.
Driver, R. W. L. Potts, and R. C. Miller. Visitors were present as
follows: Oscar G. Bacon, A. M. Adamson, Barbara Prendergast,
and Joseph E. Ryus.
The minutes of the previous meeting were read and approved.
President Keifer announced the appointment of a nominating
committee consisting of Mr. Potts, Dr. Van Dyke, and Prof. Essig
(chairman), and asked that they report at the next meeting.
Dr. Van Dyke and Dr. Miller reported recent visits from ento-
mologists in the service who were on their way overseas. Their
remarks were supplemented by various others who had had recent
JANUARY, 1944]
PACIFIC COAST ENT. SOCIETY
39
visits or communications from members in the service. The secre-
tary requested that he be kept informed of the most recent ad-
dresses, ranks and branches of the service of Society members.
Mr, Prince reported the results of recent plague surveys by the
U. S. Public Health Service in western United States. He reported
the collection of infected fleas from Neotoma and Citellus in New
Mexico, prairie dogs in eastern Colorado and parts of Montana.
His remarks were followed by considerable discussion of the geog-
raphy of plague.
Dr. Duncan called attention to the interesting paper on Rosalia
funehris Mots, by Prof. Essig in the last number of the Pan-Paciflc
Entomologist. In view of the apparent rarity of this species, he
suggested that it might be of interest to know that Rex and Norma
Bartges had collected 20 or 30 specimens on California laurel
{Umbellularia calif omica) . Dr. Van Dyke commented that Mr.
Ricksecker had collected several hundred from a skid row made of
laurel, but that he, in 50 years of collecting, had taken only three
live specimens.
Mrs. Prendergast spoke of her studies on the life history and
habits of an as yet unidentified species of Megachile in San Fran-
cisco. The bees line their cells with petals from the California
poppy (Eschscholtzia calif omica) and appear to be rather com-
monly parasitized by a species of Coelioxys.
The president then presented Dr. A. M. Adamson, Professor
of Entomology, Imperial College of Agriculture, Trinidad, who
addressed the Society on “The Biology of Trinidad Termites.” Dr.
Adamson briefly discussed the geographical features of the island
Trinidad, situated off the coast of Venezuela near the mouth of
the Orinoco River. He also remarked on the richness of its fauna
and flora, pointing out that biologists wishing to spend a short
visit in the tropics will find Trinidad an exceedingly favorable
location, with tropical rain-forests, large swamps of fresh and
brackish water, savannas and other varied types of collecting
grounds within easy reach of the capital, Port-of-Spain. In the
neighboring island of Tobago are superb barrier reefs of coral.
There is probably no region as near to the United States where so
great a variety of animals and plants can readily be seen within a
small area.
The termites of Trinidad number over 70 species in some 20
genera. Two features of their biology were selected by Dr. Adam-
son for special consideration, their systems of defense and their
feeding habits.
The most remarkable and highly developed defense practiced
by termites is probably that of the genus Nasutitermes, in which
the head of the soldiers is drawn out like a spray nozzle for the
ejection of poisonous liquids. These “mobile units of chemical war-
fare” are very numerous, swift, and aggressive. In action they
appear to be very effective against ants, the principle natural
enemies of termites.
40
THE PAN-PACIFIC ENTOMOLOGIST [yOL. XX, NO. 1
Large amounts of soil were found by Dr. Adamson in the ali-
mentary canal of a large number of trinidad termites, especially
in numerous species of Anoplotermes which live almost exclusively
in the ground. However, other genera were also involved. It ap-
pears that these termites swallow soil after the manner of earth-
worms, in order to digest organic matter which it contains. This
habit is probably common among tropical termites, although the
literature contains few references to it. It is conceivable that
termites may profoundly affect the soil in areas where abundant.
In at least some of the genera studied. Dr. Adamson found soil
in the alimentary tract of soldiers, as well as workers, but not in
nymphs or members of the reproductive caste. Further observa-
tions along this line should shed some light on the diet of the vari-
ous castes, a subject difficult to investigate by other methods and
about which almost nothing is known.
After a general discussion of Dr. Adamson’s paper, the meet-
ing was adjourned. — E. G. Linsley, Secretary.
One Hundred and Eightieth Meeting.
The one hundred and eightieth meeting of the Pacific Coast
Entomological Society was held at 2:30 p.m. on December 18, 1943,
in the entomological laboratories of the California Academy of
Sciences, San Francisco. President Keifer in the chair. The fol-
lowing members were present: H. H. Keifer, E. G. Linsley, E. 0.
Essig, A. E. Michelbacher, R. F. Smith, R. C. Miller, W. H. Lange,
E. C. Van Dyke, M. A. Stewart, F. M. Prince, W. M. Pearce, P.
Moorehead. One visitor was present, Mr. Joseph Ryus.
The minutes of the previous meeting were read and approved.
Prof. Essig reported for the nominating committee the follow-
ing nominations for officers for 1944: president, M. A. Stewart;
vice-president, F. P. Keen; secretary, E. G. Linsley, and treasurer,
R. C. Miller. It was moved, seconded, and passed that the secre-
tary cast a unanimous ballot for the candidates.
Mr. Keifer then turned the gavel over to the new president.
Dr. M. A. Stewart. Dr. Stewart paid tribute to the services of the
retiring president and called attention to the increasing obligation
of the Society to help maintain scientific values, support funda-
mental work, and maintain continuity of effort during the present
world crisis. He urged that all members make a special effort in
attendance at meetings and in helping to make the meetings inter-
esting and worthwhile.
President Stewart then presented the retiring president, Mr.
Keifer, who addressed the Society upon “Applied Entomological
Taxonomy” (see p. 1). Following a general discussion of the sub-
ject, Mr. Keifer reported upon his investigations of eriophyid
mites, as an example of applied taxonomy. Considerable discussion
followed Mr. Keifer’s paper, after which the meeting adjourned. —
E. G. Linsley, Secretary.
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tropical fauna.
The volumes already published since 1931 comprise thousands of
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Edwards, W. Horn, E. Lindner, J. W. S. Macfie, E. Martini, A. da Costa
Lima, F. Silvestri, C. Menozzi, A. Reichensperger, F. Santschi, J. D.
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ARCTIC LEPIDOPTERA
A large collection of Arctic Lepidoptera, especially Erebia,
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Collectors who desire such material please communicate
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Canaaa.
ENTOMOLOGICAL NEWS
An illustrated magazine, published monthly — except August and
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Vol. XX
April, 1944
No. 2
THE
Pan -Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
DE LONG, A NEW GENUS— HEBENARUS— AND THIRTEEN NEW
SPECIES OF LEAFHOPPERS RELATED TO CLOANTHANUS 41
LINSLEY, LARVAL LONGEVITY IN BUPRESTIS AURULENTA LINN.... 53
WOLCOTT, THE AMERICAN SPECIES OF TRICHODES 54
POTTS AND SMITH, HIBERNATION OF DASYMUTILLA AUREOLA 60
RAY, NOTES AND DESCRIPTIONS OF MORDELLIDAE FROM THE SOL-
OMON ISLANDS 61
LINSLEY, PREY OF THE ROBBER FLY CALLINICUS CALCANEUS 67
BOHART, STENODYNERUS FUNDATUS AND RELATED SPECIES IN
NORTH AMERICA 69
ROTGER, A NEW SPECIES OF CICINDELA AND TWO NEW RECORDS
OF COLEOPTERA 76
USINGER, HOST PLANT RECORDS OF WESTERN LYGUS 78
CHAMBERLAIN, SOME RECORDS OF MYRIOPODS COLLECTED BY
W. M. PEARCE IN CALIFORNIA 79
SMITH, A NOTE ON THE HIBERNATION OF MISCHOCYTT ARUS
FLAVITARSUS IN A NEST OF DOLICHOVESPULA ARENARIA 80
San Francisco, California
1944
/
THE PAN-PACIFIC ENTOMOLOGIST
EDITORIAL BOARD
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Associate Editor Editor Assistant Editor
R. L. Usinger* G, F. Ferris E. S. Ross*
R. C. Miller, Treasurer
* On military leave
Published quarterly in January, April, July, and October with Society
Proceedings appearing in the January number. Papers on the systematic
and biological phases of entomology are favored, including articles up
to ten printed pages on insect taxonomy, morphology, life history, and
distribution.
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The Pan-Pacific Entomologist
Vol. XX, No. 2 April, 1944
A NEW GENUS — HEBENARUS — AND THIRTEEN NEW
SPECIES OF LEAFHOPPERS CLOSELY RELATED
TO CLOANTHANUS
(Homoptera, Cicadellidae)
BY DWIGHT M. DeLONG
Department of Zoology and Entomology, Ohio State University
In working over the species of leafhoppers collected in Mex-
ico, which belong to the Platymetopius groups, some 50 species
have been placed in the genus Cloanthanus, one in Scaphytopius,
while others did not seem to belong to any of the described
genera. A series of rather closely related species have, there-
fore, been separated out and a new genus Hebenariis is being
erected to include these thirteen species. Scaphytopius ritanus
Ball is being placed in this genus and new species are described
in the present treatment. Unless otherwise indicated all the holo-
type, allotype and paratype specimens are located in the author’s
collection.
Hebenarus DeLong, new genus
Related to Cloanthanus Ball but differing' from it by a flat
vertex which is usually bluntly angled and as a rule is about as
wide or slightly wider than long. The sides of vertex are usually
rather convexly rounded to the blunt apex. The vertex is also
characteristically marked by a pale margin bordered above and
below by a black line. This marking on the vertex margin is more
like Scaphoideus. The face is not angled above as in Cloanthanus
and is usually banded just beneath the apex. The venation is simi-
lar to that of Cloanthanus with numerous short costal veinlets.
The species of this group are not marked with peppered or stip-
pled pigment spots as is so frequently found in the color patterns
of species of Cloanthanus.
The types of genitalic structures of both male and female,
although differing among the species of this genus, are distinct
from those types found in the species of Cloanthanus.
Genotype Hebenarus pallidus DeLong.
42
PAN-PACIFIC ENTOMOLOGIST
[VOL.XX,NO. 2
Hebenarus pallidus DeLong, new species
Resembling ritanus in general appearance but with shorter
head, pale vertex and scutellum and with distinct male genitalia.
Length 4.5-5 mm.
Vertex short, bluntly angled, distinctly wider between eyes than
median length.
Color: Vertex and scutellum white tinged with yellow, margin
of vertex pale bordered above and below with a brown line. Face
with a narrow white band just beneath the lower black line, below
this is a darker band between the eyes, and the remainder of the
face is white. Pronotum white tinged with yellow, the disc appear-
ing brown. Elytra subhyaline with dark brown veins.
Genitalia: Female last ventral segment with posterior margin
almost truncate, punctured either side of a broad, shallow, U-
shaped notch on median third. A broad black stripe extends from
this portion of margin to base of segment. Male plates short,
triangular. Style with a long finger-like process arising on inner
margin and extending outwardly. Aedeagus composed of an L-
shaped dorsal portion the base of which is bifid and about as long
as the stem. This is attached to the base of the ventral portion
which is composed of a pair of proximal pieces which in ventral
view are tapered from the base to slender pointed tips. In lateral
view each piece has a prominent spine on the dorsal side formed
at the base of the arrowhead tip.
Holotype male and allotype female from Mexico City, D. F.,
Mexico, elevation 7,500 feet, collected September 1, 1939, by
the author. Paratype male collected at same locality September
13, 1939, by the author. Paratype females collected at Carapajv,
Mich., Mexico, elevation 6,000 feet, October 2, 1941, and at
Zitacuara, Mich., Mexico, elevation 6,700 feet, September 29,
1941, by C. C. Plummer, E. E. Good, J. S. Caldwell and the
author. A male and female paratype which were collected by
Ball and Stone at Km. 66 on the Mexico City-Cuemavaca Road,
August 17, 1936, are in the Ball collection in the U. S. National
Museum.
Hebenarus ritanus (Baix)
Scaphytopius ritanus Ball Can. Ent. 63:221, 1931.
An orange yellow species with white areolar spots on the
elytra. Length 4.5-5 mm.
Vertex bluntly angled, a little longer on middle than basal
width between the eyes in female, about one-fourth longer than
basal width in the male.
APRIL, 1944]
Delong— HEBENARUS
43
Color: Yellow tinged with orange, margin of vertex paler,
bordered above and below with a slender brown line. Face dull
orange brown a conspicuous pale arc just below margin. Elytra
tinged with orange on basal portion with numerous pale spots and
areas, posterior portion almost pale hyaline, veins orange.
Genitalia: Female last ventral segment broadly, slightly pro-
duced. Male plates short, triangular, apices blunt. Style with a
long finger-like apical process arising on inner margin which
curves outwardly. Aedeagus composed of a dorsal and a ventral
portion. The dorsal portion in lateral view is L-shaped with the
base of the “L” extending vertically in the basal portion of seg-
ment. This is attached to the ventral portion by a narrow basal
strip. Ventral portion composed of a single piece. When viewed
laterally it is broad at base and tapers to a slender apex which
bears a pair of long spine-like processes extending laterally.
This species was originally described from specimens col-
lected at Tucson, Santa Rita Mountains and Pima County, Ari-
zona. A pair of specimens were collected in the Santa Rita
Mountains and compared with the types by Dr. Oman and
through his kindness have been available for study and dissec-
tion. This species has not been collected in Mexico, but un-
doubtedly occurs in some of the northern mountain ranges of
that country.
Hebenarus ritanus var. rubralineus DeLong, new variety
Resembling ritanus in form and general appearance but
marked with longitudinal red stripes on vertex, pronotum and
scutellum. Length, female 5 mm.
Vertex produced and rather sharply angled, a little longer at
middle than basal width between the eyes.
Color: Vertex, pronotum and scutellum pale brown. A pair of
broad red stripes arise at margin of vertex and extend across
vertex, pronotum and terminate in basal angles of scutellum.
These form a narrow brownish margin next either eye and a broad
median stripe which is narrow at apex and at base, and is broader
on median half. Pronotum with a pair of longitudinal red stripes
behind each eye. Margin of vertex reddish with a black line just
above and just below. Face brown with a broad white band just
beneath the black line and indications of several pale arcs. Elytra
pale brown, with dark brown markings on disc, apex smoky and
apical costal veinlets dark.
Genitalia: Female last ventral segment sloping from base to
posterior margin which is slightly trilobate.
44
PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 2
Holotype female collected at ORIZABA, V. C., Mexico, October
17, 1941, by E. E. Good, J. S. Caldwell, C. C. Plummer and the
author.
Hebenarus abruptus DeLong, new species
Resembling ritanus in general form and appearance but with
a shorter vertex and a darker coloration. Male genitalia dis-
tinct. Length, male 4.5 mm.
Vertex bluntly angled, length at middle equaling the basal
width.
Color dull yellow, tinted with brown. Vertex dirty yellow with
a broad, pale brown stripe either side of middle, a slender black
line just above pale margin and a black line dividing and another
just below the pale marginal band on face. Main portion of face
dark brown, the genae paler. Pronotum and scutellum brown.
Elytra subhyaline, tinged with brown, veins dark brown.
Genitalia: Female last ventral segment broadly roundedly pro-
duced with a slight notch at middle. The central third of segment
black from margin to base. Male valve large, triangular. Plates
short, triangular with bluntly pointed apices exceeding valve by
about one- half its length. Aedeagus with a large ventral and a
small dorsal portion. The ventral process is long with the apical
third bent at right angles and extending ventrally. This bent por-
tion is enlarged at middle. The dorsal portion which consists of
a rather small curved ring enlarged on the ventral side is connected
to the ventral portion by a rather long and broad process.
Holotype male collected near Cuernavaca, Mor., Mexico,
elevation 7,000 feet, from shrubs, October 21, 1941, by E. E.
Good and the author. Allotype female and paratype females
collected at Carapan, Mich., Mexico, elevation 6,000 feet from
Oak, October 2, 1941. Paratype male from Uruapan, Mich.,
Mexico, elevation 5,300 feet, October 1, 1941, and paratype
female collected at Zacapu, Mich., Mexico, elevation 6,500 feet,
October 4, 1941. All the paratypes and the allotype were col-
lected by C. C. Plummer, E. E. Good, J. S. Caldwell and the
author.
Hebenarus altus DeLong, new species
A blunt headed species resembling abruptus but with shorter
vertex and paler coloration. Length, female 4.5 mm.
Vertex short, bluntly angled, slightly shorter at middle than
basal width between the eyes.
APRIL, 1944]
Delong— HEBENARUS
45
PILATE I
Lateral views of male genital structures as labeled,
ventral view of aedeagus of altus (center), ventral
view of male genital structure of spinosus (bottom
center) .
46
PAN-PACIFIC ENTOMOLOGIST
[VOL.XX,NO. 2
Color dirty white to pale yellow, with few definite markings. A
faint brown line just above and one just below the pale margin.
Pronotum darker brown. Scutellum pale with rust colored basal
angles. Elytra subhyaline, veins except on claval area, dark brown.
Disc and apex of clavus slightly tinged with brown. Face white
slightly infuscated.
Genitalia: Female last ventral segment almost truncate with a
slight U-shaped notch at middle. The central third of segment is
broadly black margined and slightly, almost abruptly produced
beyond the outer portions of posterior margin. Male plates short,
broad at base, triangular, with blunt apices. Style with a curved
finger-like process arising on inner margin at apex and directed
outwardly. The aedeagus in general type resembles that of rita-
nus but the ventral portion is longer and the dorsal portion has a
thicker, more curved basal structure. The ventral portion is a
single structure, tapered to a slender apex with a pair of rather
long slender spine-like processes which extend laterally. The
dorsal portion is L-shaped with the base of the “L” thick and
curved.
Holotype male from Saltillo, Coah., Mexico, elevation
7,500 feet, September 23, 1941. Allotype female collected near
Mexico City, D. F., Mexico, elevation 8,000 feet, September 1,
1939, and paratype females from both localities collected by
C. C. Plummer, J. S. Caldwell, E. E. Good and the author.
Hebenarus rubranotus DeLong, new species
Resembling altus in general form with pronotum, scutellum
and clavus heavily irrorate with red. Length, female 5 mm.
Vertex bluntly angled, slightly longer at middle than basal
width between eyes.
Color pale yellow, a faint pencil line just above and another
just below pale margin. Pronotum, scutellum and clavus stippled
with red, except the basal angles and apex of the scutellum and
the apical fourth of clavus. Veins of elytra,, except clavus, pale
brown, and a brown spot at apex of clavus. Face pale yellow,
unmarked.
Genitalia: Female last ventral segment appearing almost trun-
cate on posterior margin with the median third slightly abruptly
produced, truncate at apex and black margined.
Holotype female and paratype female collected from Oak on
the mountains at 6,500 feet elevation just west of Saltillo,
Coah., Mexico, September 23, 1941, by the author.
When the males of this brilliantly colored species are ob-
tained and examined it may prove to be only a color variety of
OltUSm
APRIL, 1944]
D eLONG— HEB EN ARUS
47
Hebenarus curtus DeLong, new species
A blunt headed species resembling altus but with a shorter
head, diflferent coloration, and distinct female genitalia. Length
4,5 mm.
Vertex short, almost one- third wider between eyes at base than
median length.
Color: Vertex dull yellowish with a white longitudinal stripe
either side of median line which does not extend to apex and a
short white longitudinal stripe next either eye. Vertex margin
white bordered above and below with a brown line. Another white
band on face touches marginal band at middle but is narrower
and more distant from marginal band at the ends. Face dark
brown. Pronotum dark brown with a few paler spots. Scutellum
dull orange with a darker oblique broad stripe either side of mid-
dle. Elytra pale subhyaline with dark brown veins. The portion
covering body appears very dark.
Genitalia: Female last ventral segment concavely roundedly
produced from near base to form a produced rounded lobe either
side of a deep median, narrow V-shaped incision, reaching more
than halfway to the base. Each lobe is black colored.
Holotype female collected at Zacapu, Mich., Mexico, eleva-
tion 6,500 feet, October 4, 1941, by E. E. Good, C. C. Plummer,
J. S. Caldwell and the author. Paratype female collected at Km.
57, elevation 8,500 feet, on the Cuernavaca Road in Morelia,
Mexico, Ocober 21, 1941, by E. E. Good and the author. A
female paratype collected by Ball and Stone at Km. 66 on the
Mexico Gty-Cuernavaca Road, August 17, 1936, is in the Ball
collection in the U. S. National Musem, Washington, D. C.
Hebenarus comutus DeLong, new species
Resembling curtus in form and general appearance but with
distinct genitalia. Length 4-4.5 mm.
Vertex bluntly angled. About as long as width between eyes at
base.
Color: Vertex brown, margin white with a narrow black line
above. A white elongate spot just back of apex which is replaced
on the basal three-fourths by a pair of white parallel lines extend-
ing to pronotum, one either side of middle. Pronotum brown,
mottled with pale. Scutellum reddish brown, a white spot about
the middle on either side. Elytra hyaline with brown veins, a
brown spot on middle and a smaller one on apex of clavus. Face
reddish brown a complete pale arc a little below marginal band
and indications of three other pairs below.
48
PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 2
Genitalia: Female last ventral segment with short lateral mar-
gins from which the posterior margin is gradually produced to
form a broadly rounded lobe either side of a short narrow V-
shaped median notch. Male plates rather short, thick and blunt,
almost rounded at apex. A conspicuous lateral rounded lobe on
outer margin of each plate at base. Aedeagus composed of a pair
of rather long proximal pieces which are slender at base and are
widened on apical half to form a broad funnel-like tip. A small
rather square-shaped tooth is present on the ventral side of each
process at about its middle. There is also a small curved dorsal
portion which is just above the paired ventral processes.
Holotype male and allotype female from San Cristobal,
Guatemala, 7,200 feet elevation, March 30, 1942. Paratype male
and female from Panajachal, Guatemala, elevation 5,000 feet,
March 5, 1942, all collected by Dr. C. C. Plummer.
Hebenarus marginellus DeLong, new species
Resembling ritanus in general form but with different colora-
tion. Length, female 5 mm.
Vertex produced and bluntly angled, length at middle equalling
basal width between the eyes.
Color: Dull orange to brown. Vertex with a pale longitudinal
stripe either side of median line. Pronotum and scutellum with
faint, longitudinal, paler stripes. Elytra brownish subhyaline with
darker brown veins. Margin of vertex with a double white band
separated by a narrow brown line and margined above and below
by a brown line. Face pale brown.
Genitalia: Female last ventral segment produced on lateral
margins from near base, posterior margin truncate or slightly
concave.
Holotype female from Orizaba, Vera Cruz, Mexico, 3,500
feet elevation, collected October 17, 1941, and paratype females
from Carapan, Mich., Mexico, elevation 6,000 feet, October 2,
1941, and Zitacura, Mich., Mexico, elevation 6,700 feet, Sep-
tember 29, 1941. All collected from oak by E. E. Good, J. S.
Caldwell, C. C. Plummer and the author.
Hebenarus arcuatus DeLong, new species
Closely related to marginellus but with a shorter vertex and
different coloration. Length, female 5.5 mm.
Vertex slightly longer at middle than basal width between the
eyes.
APRIL, 1944]
Belong— HEBENARUS
49
Color: Vertex pale brown, margin paler. An elongated pale
spot at apex and two oblique pale stripes either side extending
from margin to base, one next each eye and one either side and
not far from median line. Pronotum dark brown, paler along an-
terior margin. Scutellum pale brown, basal angles orange. Elytra
subhyaline, veins, nervures and ramose pigment lines on disc and
clavus, brown. Face pale washed with light brown, a narrow
brown transverse line just beneath pale margin and a pale “sharks
mouth” mark between eyes not far below margin.
Genitalia: Female last ventral segment with posterior margin
almost truncate, broadly, slightly emarginate. The color markings
cause it to appear deeply squarely notched because of a dark arc
shaped spot between the ends of which and the posterior margin
the segment is brown.
Holotype female collected at Jalap A, Guatemala, elevation
4,854 feet, March 24, 1942, by Dr, C. C. Plummer.
Hebenarus huachucus DeLong, new species
Resembling ritanus in general form but with a shorter vertex
and distinct male genitalia. Length 5 mm.
Vertex rather broad and bluntly angled, length at middle equal-
ing width between the eyes.
Color yellow, marked with brown. Vertex yellowish, with a
wedge-shaped white mark at apex and a pair of white median long-
itudinal stripes which are slightly separated extending from apex
of wedge-shaped spot to the base. Margin white with a narrow
dark line just above and another just below. Face with a brown
band just below marginal white band and another narrow white
band just beneath the brown. Between this and the clypeus the
face is pale brown distinctly tinted with red. In color it is strik-
ingly contrasted with the yellowish clypeus, lorae and genae. Pro-
notum pale brown. Scutellum orange. Elytra white with dark
brown veins and numerous pigment marks which appear as cross
veins.
Genitalia: Female last ventral segment with posterior margin
gently sloping from either side to form a pair of rounded lobes
between which is a median broad V-shaped notch which extends
more than halfway to base of segment and is brown bordered.
Male plates short, triangular, apices bluntly angled. Style with a
short, tapered, sharply pointed finger-like apical process which is
directed outwardly. Aedeagus consisting of a ventral portion which
is bifid at about its middle, so that the apical half is composed of
a pair of proximal structures. In lateral view the basal half is
broad with a curved ventral bulge at middle bearing a short square
tooth. The apical half is tapered from middle to form a ventrally
50
PAN-PACIFIC ENTOMOLOGIST
[VOL.XX, NO. 2
curved broadened tip. The dorsal process is short with a bifid
basal portion and a short apically curved and pointed ventral or
apical portion.
ABRUPTUS
PLATE II
Ventral views of last ventral seg'ments of females as labeled.
Holotype male, allotype female and male and female para-
types collected in the Huachuca Mountains, Arizona, July 20,
1936, and June 9, 1935, by Prof. J. N. Knull.
Hebenarus monticolus DeLong, new species
In general form resembling ritanus but with brown banded
elytra. Length, female 5 mm.
Vertex produced and bluntly angled, long^er at middle than basal
width between eyes.
APRIL, 1944]
Delong— HEBENARUS
51
Color: Vertex bright yellow with white longitudinal markings,
in the form of an elongated spot at apex and a longitudinal stripe
each side of middle not extending to the white margin which is
bordered above and below by a black line. Pronotum brown with
five pale longitudinal stripes. Scutellum yellow. Elytra white,
spotted with yellow and with two prominent brown bands. One
crosses the middle of clavus and one crosses apex of clavus. Apex
of elytra brown bordered, veins brown. Face brown with pale
arcs above.
Genitalia: Female last ventral segment sloping from base to
posterior margin which is broadly rounded either side of a broad
shallow, median notch.
Holotype female collected at Taxco, Guerrero, Mexico, Sep-
tember 9, 1939, by C. C. Plummer and the author. This was
swept from a thick growth of shrubs on the mountainside at an
elevation of 6,000 feet.
Hebenarus maculosus DeLong, new species
Resembling ritanus in general form but with different colora-
tion and male genitalia distinct. Length 4.5-5.5 mm.
Vertex bluntly angled, length about equaling width.
Color: Vertex pale with longitudinal pale orange markings. A
prominent black line above and another below the white margin.
Face with a white band just below the lower black line. Upper
portion of face pale brown, lower portion lighter. Pronotum brown-
ish with paler longitudinal stripes. Scutellum with basal angles
pale brownish, median portion paler. Elytra whitish subhyaline,
veins brown, a brownish spot at apex of clavus and apical portion
of elytra infuscated.
Genitalia: Female last ventral segment with posterior margin
concavely rounded either side of a median roundedly produced lobe
which extends beyond the lateral angles, male plates of medium
length and triangular. Style rather long and with a prominent
apical finger-like process on inner margin which is directed caud-
ally and outwardly. Aedeagus composed of a pair of long slender
ventral processes which are pointed at apex and bear a prominent
pointed tooth on inner margin just before apex. These are con-
nected by a long slender process from the base to a small dorsal
portion composed of a single ventro-apical process and a bifid
dorsal process.
Holotype male collected at Carapan, Mich., Mexico, eleva-
tion 6,000 feet, October 2, 1941, from Oak. Allotype female
taken at Puebla, Puebla, Mexico, elevation 8,500 feet, October
18, 1941, both collected by J. S. Caldwell, E. E. Good, C. C.
52
PAN-PACIFIC ENTOMOLOGIST
[V0L.XX,N0. 2
Plummer and the author. Paratype female collected from shrubs
by Dr. Dampf at Bella Vista, D. F., Mexico, 8,000 feet eleva-
tion, September 17, 1933 (M. B. 366) .
Hebenarus luteus DeLong, new species
Resembling ritanus in general form. A sharp headed species
with different coloration and different female genitalia. Length,
female 5 mm.
Vertex produced, apex bluntly angled, distinctly longer on
middle than basal width between the eyes.
Color: Vertex orange yellow with a paler longitudinal stripe
either side of median line. Margin white, bordered above and
below by a black line. Face dark brown with a broad white are
just below margin and traces of four pairs of arcs on upper por-
tion. Pronotum dark brown, scutellum pale brown. Elytra pale
subhyaline, washed with pale brown, veins dark brown. The body
gives a dark brown appearance to the elytra as seen from above.
Genitalia : Female last ventral segment strongly sloping to pos-
terior margin which is convexly rounded from either side to form
a broad, conspicuaus V-shaped notch which extends about one-
fifth the distance to the base.
Holotype female anl paratype female collected at Orizaba,
Vera Cruz, Mexico, Rm. 280, elevation 7,700 feet, October 17,
1941, by C. C. Plummer, E. E. Good, J. S. Caldwell and the
author. These were collected from a growth of shrubs on one of
the higher mountains several miles west of the city of Orizaba.
Tenuarus DeLong, new subgenus
This subgenus is distinguished from the other species placed in
this genus by a thin edged vertex margin which is slightly turned
up and the sides of vertex distinctly convexly rounded. The costal
margin has very few costal veinlets and the male genitalia are en-
tirely different in general form. The pygofer bears a pair of long
spines on the ventro-caudal margin and the styles are long and
very slender.
Genotype Hebenarus {Tenuarus) spinosa DeLong.
Hebenarus (Tenarus) spinosus DeLong, new species
Resembling ritanus in general form but with vertex, pronotum
and scutellum longitudinally striped and with distinct male
genitalia. Length, male 5 mm.
Vertex rather strongly produced but bluntly angled, longer at
middle than basal width between the eyes, sides convexly rounded.
APRIL, 1944]
LINSLEY— BUPRESTIS
53
Color: Orange yellow, vertex with a pale median longitudinal
spot at apex, a broad white longitudinal stripe either side of mid-
dle and a pale longitudinal spot along the anterior portion of each
eye. Margin pale bordered above and below by a conspicuous
brown line. Pronotum with seven longitudinal stripes. Scutellum
orange yellow with a paler median longitudinal stripe terminating
in the apical angle which is also paler. A white longitudinal stripe
on inner margin of either basal angle. Elytra pale brown with
dark brown veins, and with round, white, brown bordered spots in
anteapical and apical cells just before and just posterior to the
cross veins. Face rather uniform pale brown except a conspicuous
angled white stripe just beneath margin which is bordered beneath
by a dark brown line.
Genitalia: Male plates long, rather broad at base, convexly
rounded on basal two-thirds then tapered to pointed apices. Style
long and narrow with pointed apex which is slightly curved in-
wardly. Aedeagus consisting of a single portion which curves
dorsally at base and is more slender. In ventral view the aedeagus
is enlarged at apex forming two rounded lobes. Each pygofer
bears a long slender spine on inner ventral portion which extends
decidedly beyond the pygofer.
Holotype male collected at Vergel, Chiapas, Mexico, valley
of the Rio Huixtla, elevation 3,000 feet, June 6, 1935, by Dr. A.
Dampf (M. F. No. 4458) .
LARVAL LONGEVITY IN BUPRESTIS AURULENTA LINN
The writer has recently reviewed^ available information on
larval longevity and delayed emergence of Buprestis aurulenta
Linn, from structural timbers. Subsequently an interesting record
from Guam was noted in an article by Swezey^. Swezey states,
“One specimen of this American species was obtained in our
residence at Piti, July 9, [1936]. It had apparently issued from a
porch floor board as there was an exit hole which appeared fairly
fresh. Two other similar holes had an older appearance. The
boards were of Douglas fir which had been shipped from the
Puget Sound Region. The house was constructed prior to 1915.”
Although Swezey expresses the doubt that the larva could
have existed for that length of time before maturing, I think
there is now little question but that this was actually the case. —
E. Gorton Linsley.
^ Linsley, E. G. 1943. Delayed emergence of Buprestis aurulenta from struc-
tural timbers. Jour. Econ. Ent., 36 :348-349.
- Swezey, O. H. 1942. Miscellaneous families of Guam Coleoptera. B. P.
Bishop Mus., Bull. 172:150-171.
54
PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 2
THE AMERICAN SPECIES OF TRICHODES
(Coleoptera, Cleridae)
BY A. B. WOLCOTT
Downer’s Grove, Illinois
A third of a century has elapsed since the writer prepared his
synopsis of the Trichodes^. At that time ten species were known
from our fauna, later two other species were described, the first
of these being bicinctus Green, the other hasalis Van Dyke, thus
bringing the number of known species to an even dozen. As it
is likely that few, if any, remain to be discovered the present
time appears to be opportune for the presentation of an amended
and augmented table of species.
It is very probable that eventually the name Trichodes Herbst,
will have to be dropped in favor of Clerus Fab. John Curtis^ in
1824 designated Attelahus apiarius Linn., as the genotype of
Clerus, in which move he was followed by a number of other
writers. In 1840 F. W. Hope^ also designated T. apiarius Lirin.,
as type of Clerus Latr., and synonymized it with Trichodes Fab.
Dr. E. A. Chapin (m litt) who has thoroughly investigated the
status of these two generic names finds that they are isotypical
and that Clerus Fab. (1775) has priority over Trichodes Herbst
(1792). Thus in conformity with the rules of nomenclatural
procedure it will be necessary to substitute Clerus Fab. for the
name Trichodes of Herbst.
The present writer retains the name Trichodes merely as a
matter of convenience.
Key to the American Species of Trichodes
1. Form slender; elytra at base two-fifths as wide as long 2
Form moderately robust; elytra at base subequal to one-half
their length 10
2. Antennal clava broadly triangular, about one-half longer than
broad 3
Antennal clava elongate-triangular, twice as long as broad..l5
3. Elytral apex truncate (rounded in bicinctus and hibalteatus.A
Elytral apex rounded 10
4. Abdomen blue black, in part red oresterus Wolcott
Abdomen and legs yellowish-brown 5
^Wolcott, A. B., 1910, Notes on some Cleridae of Middle and North America,
Field Mus. Nat. Hist., Zool. Ser., 7 (10) :339-401, Pis. 6-6.
^ Curtis, John, 1824, The Genera of Insects, British Ent., 1 :44.
® Hope, F. W., 1840, The Coleopterist’s Manual, 3 :137.
APRIL, 1944]
WOLCOTT— TRICH ODES
55
5. Pronotum metallic green, coarsely, not cribrately punctate,
disk shining basalis Van Dyke
Pronotum black with greenish lustre 6
6. Elytra fulvous or reddish, fasciae black bicinctus Green
Elytra ferrugineous 7
7. Legs usually wholly or in part ferrugineous; funicle of an-
tennae rufous, clava darker; flanks of elytra and yellow fas-
ciae broadly bordered with dull black.. /lomi Wolcott and Chapin
Legs and antenna black 8
8. Elytra dull black, abdomen shining olive green
peninsularis Horn
Elytra yellow or orange red 9
9. Two elytral fasciae and apex blue, truncate siTnulcutor Horn
Two elytral fasciae black, apex rounded, immaculate; abdomen
dark blue, apical segment always red ....bibalteatus LeConte
10. Elytra feebly shining black, with faint cu^eous lustre; ab-
domen dark green, in part red nexus Wolcott
Elytra shining blue, green, yellow or red 11
11. Prothorax with sides compressed at apex, strongly compressed
at base 12
Prothorax subquadrate, sides feebly compressed at apex and
base 13
12. Pronotum blue or green, moderately sparsely, rather coarsely
punctate, clothed with rather long dense yellow pubescence
ornatus Say
Pronotum dark aeneous, moderately densely, rather coarsely
and shallowly punctate, clothed with rather long, dark fulvous
or tawny pubescence 14
13. Robust; elytral fasciae scarcely discernible, only evident by
reason of the more sparse but coarse, confluent punctuation
of the areas normally occupied by the yellow fasciae of the
elytra omatus var. obsoletus Wolcott
Slender, small, otherwise as in typical form
ornatus var. tenellus LeConte
14. Elytra dark blue with a more or less rounded red or yellow
maculation at middle, contiguous to lateral margins
bimaculatus LeConte
Elytra blue, trifasciate with yellow or red 15
15. Pronotum blue, rather flnely, moderately densely punctate,
clothed with rather long dense grayish pubescence
nutalli Kirby
Pronotum dark brown, clothed with rather long dense brown-
ish pubescence. Elytra red or yellow, two fasciae and apices
blue or purple black apivorus Germar
Anteirior fascia reduced to a large irregularly rounded
maculation on each elytron, postmedian fascia broadly in-
terrupted at the suture.. ..var. borealis Wolcott and Chapin
56
PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 2
Notes on Some Species of Trichodes
Trichodes basalis Van Dyke
Trichodes basalis Van Dyke, 1943, Pan-Pac. Ent., 19:41.
Dr. E. G. Linsley has very kindly supplied the writer with an
excellent sketch of the elytral markings of the type specimen of
this species, a copy of which is presented (Fig. 1). Dr. Linsley
also sent some helpful and valuable notes pertaining to this
species, which are quoted in their entirety “In form it resembles
a slender specimen of oresterus Wolcott, the basic elytral pattern
is also similar to some examples of oresterus. However, the pale
areas of the elytra are clear yellow without any reddish or orange
tint. The pronotal pubescence is yellow and a little longer (pos-
sibly also denser) than in my series of oresterus. The punctures
of the pronotum and elytra are coarse but less deep than those
of oresterus. Likewise the legs and entire under surface is
brownish-yellow in contrast to the blue color and red abdomen
of oresterus.^’ (Linsley, in litt.)
Type locality: Imperial County, California.
Trichodes bicinctus Green
Trichodes bicinctus Green, 1917, Ent. News., 28:367.
Dr. J. W. Green very generously gave the writer one of the
two males mentioned in his original description of this species.
The specimen is from Green Valley, Brewster County, Texas, July
14, collected by J. W. Green. It agrees with the description in
every detail. A figure of the elytral pattern is shown (Fig. 2).
Dr. Linsley also presented the author with a pair of this spe-
cies, male and female, taken on flowers at Laredo, Texas, June 12,
1932. In both of these specimens the body beneath is green, the
abdomen greenish black. The black elytral markings reduced to
two transverse maculations on each elytron and the apex with a
narrow lunate maculation. This form is represented by the fig-
ure given (Fig. 3). The elytral apex is rounded in both sexes.
Trichodes horni Wolcott and Chapin
Trichodes homi Wolcott and Chapin, 1918, Bull. Brooklyn Ent.
Soc., 13:107.
Trichodes illustris Horn, 1876, Trans. Amer. Ent. Soc., 5:231.
APRIL, 1944]
WOLCOTT— TRICHODES
57
As pointed out by Wolcott and Chapin illustris is preoccupied
and the name horni has been substituted in its stead. Horn in
his description states that “the elytra . . . conjointly rounded 6,
truncate and sinuate at tip ? The elytra are always more or
less truncate in both sexes, never conjointly rounded, the trun-
Gature being merely of less extent in the 5 than in the 9 . A
rather small specimen of this species from the Santa Rita Moun-
I
Elytral Pattern in Trichodes
Fig. 1. Trichodes basalts Van Dyke. 2. T. bicinctus
Green. 3. T. bicinctus var.
tains, Arizona, September 29, 1932, has the elytra, with the ex-
ception of the yellow fasciae, entirely dull black with a small
rounded maculation on each midway between the median and
anteapical fasciae, red.
In the text of the writer’s descriptive notes® the fourth line
under T. illustris Horn, “The more finely punctured headf^
should read “The more coarsely punctured head!’
1 Wolcott, A. B., 1910, 1. c., 7(10) :369
58
PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 2
TrICHODES SIMULATOR HoRN
Trichodes simulator Horn, 1880, Trans. Amer. Ent. Soc., 8:149,
1880.
Trichodes simulator flavescens Cockerell, 1898, New Mex. Agr.
Exp. Sta. Bull. 28:155.
Trichodes simulator Wolcott, 1910, Field Mus. Nat. Hist., Zool,
Ser., 7(10) :170.
Abdomen shining blue, terminal segment deep blue black.
Body beneath densely clothed with long semi-recumbent yellowish-
white pilosity, the abdomen with short, sparse pile. Legs dark
blue with greenish lustre, tarsi obscurely testaceous. Elytral
apices in both sexes truncate. The following localities may be
added to those already recorded: Bandelier, New Mexico; Cedar
Qty, Utah; Grand Junction and Craig, Colorado.
Trichodes bibalteatus LeConte
Trichodes bibalteatus LeConte, 1858, Jour. Acad. Nat. Sci. PhiL,
(2)4:18; Horn, 1876, Trans. Amer. Ent. Soc., 5:231; Wolcott,
1910, Field Mus. Nat. Hist., Zool. Ser., 7(10) :371.
The blue abdomen has the terminal segment always red, often
obscurely so in the female, rarely the entire abdomen red, with
the exception of the basal and second segment, which are blue.
The elytral apices are rounded in both sexes. To the recorded
localities may be added: Fedor, Lee County, Texas (Birkmann),
and Oklahoma.
Trichodes ornatus Say
Trichodes omatus Say, 1825, Jour. Acad. Nat. Sci. Phila., 3:189;
Klug, 1842, Abh. Berl. Akad., p. 340; Spinola, 1844, Mon. Cler.,
1:327, PI. 31, fig. 5; LeConte, 1849, Ann. Lyc. Nat. Hist. N. Y.,
5:18; Say’s Comp. Writ., Lee. ed., 1859, 2:120; Horn, 1876,
Trans. Amer. Ent. Soc., 5:231; Horn, 1891, Ent. News, 2:6;
Cockerell, 1893, Trans. Amer. Ent. Soc., 20: 329; Wickham,
1893, Can. Ent. 27:249; Wolcott, 1910, Field Mus. Nat. Hist.,
Zool. Ser., 7(10) :172.
Trichodes Hartwegianus White, 1849, Cat. Cler. Brit. Mus., 4:60;
Cockerell, 1893, Trans. Amer. Ent. Soc., 20:329.
Trichodes Douglasianus White, 1849, Cat. Cler. Brit. Mus. 4:60;
Cockerell, 1893, Trans. Amer. Ent. Soc., 20:329.
A large example of this species from Banff, Alberta, has quite
broad elytral markings, but the basal fascia is broadly inter-
APRIL, 1944]
WOLCOTT— TRICHODES
59
rupted at the suture, from the inner end of the remaining portion
of this fascia a backward extending hroad vitta is united near
suture with the median fascia. In a specimen from Parma, Idaho,
the basal three-fifths of the elytra red, the posterior margin of
this portion truncate, the red portion has the humeral umbones
and a very small oblique maculation at the middle blue, the
apical two-fifths blue, with the usual oblique yellow fascia of this
part. In another specimen from the same locality the elytra are
entirely red, with a small postmedian marginal maculation and
the apices narrowly blue. In a specimen from Aspen Lake,
Klamath County, Oregon, the elytra are entirely blue, with a very
small transverse yellow maculation distant from lateral margin,
at middle of each elytron.
To the localities in which T. ornatus and T. nutalli occur in
common Pike’s Peak, Colorado, may be added.
Trichodes omatus var. nov. obsoletus Wolcott
Form rather robust, Elytra at base nearly one-half as broad as
long, sides straight and parallel, scutellar area deeply and broadly
depressed, base midway between the scutellar depression and the
humeri depressed, thus causing the humeral umbones to appear
very prominent, each elytron with a deep, rather broad groove
parallel to the suture, extends from the scutellar depression nearly
to the elytral apex. Head dark green, rather shining with slight
aeneous lustre, coarsely roughly sculptured, antennal clava broad,
short. The antennae, palpi, tibiae and tarsi fulvous, clothed with
short, rather dense fulvous pile. Pronotum green, with distinct
aeneous lustre, coarsely, rather sparsely, irregularly punctate.
Elytra cyaneus with cupreous lustre, moderately shining, coarsely,
densely punctate, sutural bead very dark blue, elytral markings
arranged as in T. ornatus, but practically obsolete, faintly badius
and this covered by a faintly sordid whitish surface gloss. The
head, pronotum and elytra clothed with fulvous pile, quite dense
and longest on head, pronotum and base and apex of the elytra.
Body beneath and abdomen green with cupreous lustre. Length
10 mm.; breadth (across humeri) 3 mm.
Holotype (^), Sierra Ancha Mts., Arizona. (D. K. Dun-
can) in collection of the author.
This form would appear to be suflSciently distinct to be
classed as a valid species, but the pattern of the scarcely dis-
cernible elytral markings, which, by the way, are very difl&cult
to describe, show it to be a mere variety of T. ornatus Say.
60
PAN-PACIFIC ENTOMOLOGIST
[V0L.XX,N0. 2
Trichodes nutalli Kirby
Trichodes nutalli Wolcott, 1910, Field Mus. Nat. Hist., Zool. Ser.,
7(10) :174.
Under this species the synonym Trichodes nutalli of Say
should be transferred to the synonymy of T. apivorus Germar.
Trichodes apivorus var. borealis Wolcott and Chapin
This name (which appears in the key) was proposed by the
authors to replace the preoccupied name interruptus LeConte.
HIBERNATION OF DASYMUTILLA AUREOLA PACIFICA
On January 21, 1944, while collecting under rocks near Pat-
terson Pass, Alameda County, Calfomia, elevation 1150 feet,
two unworn female Dasjmutilla aureola pacifica (Cresson) were
found in hibernation. Under the same rock was a broken 3-cell
nest of Sceliphron servillei (Lepeletier) which apparently had
not produced adult wasps.
MickeP cites common records of aureola pacifica from March
through November, while a single December and a single Janu-
ary record are also given for Berkeley. Mickel also notes that a
very great size variation is known in the female, indicating the
probability of a variety of hosts. In his general summary of the
life history of a Mutillid it appears that hibernation is normal
for the prepupal stage. Bohart and MacSwain^ give the activity
record for another species {D. sackenii (Cresson) as June to
November and state that pupation occurs in spring or early sum-
mer, after hibernation in a prepupal stage.
It would appear possible that in this case Sceliphron had
served as the host and that the life cycle was such as to produce
fresh adult females in the late fall or winter months. Such a
cycle is at variance with those reported by Mickel and by Bohart
and MacSwain, although Mickel lists specimens taken in the
winter months. It is possible that these also were hibernating.
Determination of the mutillids was made by Mickel; of the
wasp nest by E. Gorton Linsley. — R.W.L. Potts and R.F. Smith.
^ Mickel, C. E., 1928, Biological and Taxonomic Investigations on the Mutillid
Wasps, U. S. Nat. Mus. Bull. 14S.
^ Bohart, G. E., and J. W. MacSwain, 1939, The Life History of the Sand Wasp,
Bembix occidentalis beutenmuelleri Fox and its Parasites. Bull. So. Calif. Acad.
Sci., 38(2) :84-98.
APRIL, 1944]
RAY— MORDELLIDAE
61
NOTES AND DESCRIPTIONS OF MORDELLIDAE FROM
THE SOLOMON ISLANDS
(Coleoptera)
BY EUGENE RAY
Chicago, Illinois
The following notes and descriptions were made from thirty-
three specimens collected in the Solomon Islands by H. T.
Padgen and R. A. Lever and sent to the writer by Sir Guy A. K.
Marshall, Director of the Imperial Institute of Entomology,
British Museum (Natural History). Seven species, including an
unidentifiable one, are represented in this material. Of these
two are known forms and four are described as new. The types
of the new species are temporarily placed in the Field Museum
of Natural History, Chicago, Illinois, while paratypes remain in
the collection of Eugene Ray.
Mordella lottini Boisduval
(Fig. 1)
Mordella lottini, Boisduval, 1835. Voy. Astrolabe 2 :291.
One specimen: Tulagi, November 25, 1934 (H. T. Padgen).
This species was originally described from the island of New
Guinea and has since not been recorded elsewhere.
Mordella salomonensis Pic
(Fig. 2)
Mordella salomonensis Pic, 1929. Mel. Exot.-Ent. 54:22.
Two specimens: British Solomons, February, 1932, and Solo-
mon Islands, 1934 (R. A. Lever) . This is the first record subse-
quent to publication of the original description.
Mordella subaenea Ray, new species
(Figs. 3, 5, 6, 9)
Form slender, with the facies of a Mordellistena, sides distinctly
curved throughout. Derm black, except for labrum, proximal seg-
ments of antennae, maxillary palpi, anterior femora and spurs of
posterior tibiae, which are castaneous. Body densely covered
62
PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 2
with fine, recumbent pubescence, partaking of ground color, ex-
cept for the following albo-pubescent areas: head completely, pro-
notum wtih a thick line at base, broken at middle, scutellum com-
pletely, elytra with a pre-median rounded spot on disc of each,
which in some specimens tends to extend obliquely toward humeri,
a postmedian discal spot which varies from a rounded condition
to a thick oblique one, meso- and metasternum and legs completely
but less densely than on head, and basal margins of abdominal
segments.
Antennae .9-1.1 mm. long, segment 2 shorter than 1, 2-3 equal
in length, the former broader, 4-10 each two-thirds longer than 3,
feebly serrate, 11 longer than 10 but narrower, sides and apex
rounded. Terminal segment of maxillary palpi enlarged, with the
form of a scalene triangle.
Pronotum broader than long (.76-1.25 x .6-.95 mm.), anterior
margin and sides rounded, broadest before base, arcuate, midbasal
lobe short, broad, subtruncate. Scutellum large, broadly triangu-
lar, sides and apex rounded.
Elytra but twice as long as broad (1.6-2.43 x .76-1.25 mm.),
broadest near base, sides curved from the broadest point to apex,
apices individually rounded. Anterior and intermediate tibiae
equal to their tarsi in length, penultimate segments of the latter
expanded, distal margins straight. Anal style robust, abruptly
narrowed before middle, thence attenuate to apex, between 2.3 and
2.6 times length of apical ventral segment (.8-1.27 x .35-.48 mm.).
Posterior tarsal claws with three teeth, two equal in size, one
before, the other after the middle, the third one terminal.
Length: to apices of elytra, 2.2-3.38 mm.; to tip of anal style,
3-4.65 mm.
Ten specimens: Type, male, TuLAGi Is., March 31, 1934,
taken from low herbage on ridge (H. T. Padgen) ; a male para-
type, type locality, November 25, 1934 (H. T. Padgen and R. A.
Lever) ; three male and one female paratypes, Guadalcanal
Lunga, February 27 and March 4, 1935 (R. A. Lever) ; one
female paratype, Malaita Is., August 16, 1934, “Su’u. banana
leaf” (H. T. Padgen) ; one female paratype, Vella Lavella Is.,
September 24, 1933, “Suantali-taro” (H. T. Padgen) ; one female
paratype, Isabel, January 28, 1936, “Gojururu” (R. A. Lever).
In Blair’s key to the Mordella of the Fiji Islands (Ann. Mag.
Nat. Hist. (9) 9, 1922, pp. 562-3) this species leads to veitchi
Blair, but may easily be separated by the black anterior tibiae
and tarsi, the reduction in the albo-pubescent areas on the pro-
notum and elytra, the longer third antennal segment, the shorter
intermediate tarsi and the longer anal style.
63
APRIL, 1944]
RAY— MORDELLIDAE
Explanation of Plate
Fig. 1, Dorsum of Mordella lottini Boisduval. Fig. 2. Dorsum of
Mordella salomonensis Pic. Fig. 3, Dorsum of Mordella subaenea
Ray. Fig. 4, Dorsum of Mordellistena oceanica Ray. Fig. 5, Max-
illary palpus of Mordella subaenea Ray (male). Fig. 6, Maxillary
palpus of Mordella subaenea Ray (female). Fig. 7, Maxillary pal-
pus of Mordellistena oceanica Ray (male). Fig. 8, Maxillary pal-
pus of Mordellistena oceanica Ray (female). Fig 9, Antenna of
Mordella subaenea Ray. Fig. 10, Antenna of Mordellistena abrupta
Ray. Fig. 11, Posterior tibia and tarsus of Mordellistena abrupta
Ray. Fig. 12, Posterior tibia and tarsus of Mordellistena abrupta
Ray. Fig. 13, Maxillary palpus of Mordellistena abrupta Ray.
Fig. 14, Maxillary palpus of Mordellistena maxillaris Ray (male).
Fig. 15, Maxillary palpus of Mordellistena m.aa;tZZaHs Ray (female).
64
PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 2
Mordellistena maxillaris Ray, new species
(Figs. 12, 14, 15)
Form elongate-ovate, narrow, sides slightly curved, derm rufo-
castaneous, legs and antennae lighter, maxillary palpi flavous,
ridges of posterior legs black. Body densely covered with fine,
recumbent golden pubescence, somewhat coarser on the venter than
elsewhere.
Antennae 1.9 mm. long, slender, filiform, reaching beyond base
of abdomen, segments 1-3 short, 2 and 3 each successively shorter
than the preceding, 4 as long as 2 and 3 together, 4-10 elongate,
each somewhat broader at apex than at base, 11 distinctly longer
than 10, broadest before apex, sides strongly curved. Three distal
segments of maxillary palpi enlarged and fiattened in both sexes,
terminal segment subquadrate in male and smaller than the
isosceles-triangular segment in female; second segment greatly
enlarged and bell-shaped in male, of normal size in female; third
segment large in both sexes, ovate in male, quadrangular in
female.
Pronotum broader than long (1.13 x .94 mm.), anterior margin
and sides rounded, broadest before base, base arcuate, midbasal
lobe short, broad, subtruncate. Scutellum large, broad, triangular,
base concave, sides slightly convex.
Elytra more than three and one-half times as long as broad
(2.7X.7 mm.), broadest near base, curved from a postmedian
point to apex, apices individually rounded. Anterior and inter-
mediate tibiae longer than their tarsi, penultimate segment of the
latter broadened at apex, its apical margin excised. Posterior
tibiae with four equal, oblique, parallel ridges (excluding sub-
apical one) ; basitarsi with four, second and third segments each
with two ridges. Anal style slender, slightly curved ventrad, at-
tennuate to apex, but twice length of apical ventral segment
(1.4X.65 mm.). Posterior tarsal claws with six teeth, the basal
one before middle, others successively larger distad.
Length: to apices of elytra, 3.64 mm.; to tip of anal style,
5.04 mm.
Nine specimens: Type, male, two male and two female para-
types, Tulagi Is., October 29, 1934, on Hibiscus flower (R. A.
Lever) ; one male paratype, Guadalcanal, March 4, 1935 (R. J.
Lever) ; two female paratypes, Guadalcanal Lung a, March 4,
1935 (R. A. Lever) ; one female paratype, Tulagi, December 10,
1934, in home (H. T. Padgen).
This species is closely allied to the group that includes! the
Fijian Mordellistenu dodoneae (Montoruzier) and M. consimilis
Blair. It is similar in general appearance, but differs in its larger
APRIL, 1944]
RAY— MORDELLIDAE
65
size, the shorter second and third segments of the antennae and
the additional oblique ridge of the posterior ba&itarsi. No men-
tion is made of peculiarities of the maxillary palpi in the liter-
erature pertaining to these allied species and it is thus assumed
that the palpi are of normal form. The writer has received word
that, of the species sent for study, none can be identified with
forms described by Blair and others in the collection of the
British Museum and it is therefore assumed that additional, un-
deseribed differences exist.
Mordellistena abrupta Ray, new speeies
(Figs. 10, 11, 13)
Form elongate-ovate, narrow, slightly but distinctly curved.
Derm fusco-castaneous, legs, antennae and mouth parts castane-
ous, ridges of posterior legs black. Body densely covered with fine,
recumbent pubescence, partaking of ground color and distributed
equally finely over entire surface.
Antennae 1.6 mm. long, slender, filiform, reaching distal ab-
dominal segment, segments 2-3 short, together equal in length to
4, 4-10 equal in length, but becoming successively narrower distad,
11 distinctly longer than 10, broadest beyond middle, sides strongly
curved. Three distal segments of maxillary palpi enlarged, flat-
tened, terminal and penultimate segments subquadrate, the former
much the larger of the two, antepenultimate segment almost as
large as terminal one, subtriangular.
Pronotum distinctly broader than long (1.13x.94 mm.), ante-
rior margin and sides rounded, broadest before base, base arcuate,
midbasal lobe short, broad, subtruncate. Scutellum large, broad,
base concave, sides slightly convex.
Elytra narrow, more than three times as long as broad (2.2 x
.7 mm.), broadest near base, curved from a postmedian point to
apex, apices individually rounded. Anterior and interfiiediate
tibiae longer than their tarsi, penultimate segment of the latter
broadened at apex, its apical margin excised. Posterior tibiae
with three, equal, oblique, parallel ridges (excluding subapical
one) ; basitarsi with three, second and third segments each with
two ridges. Anal style slender, slightly curved ventrad, attenuate
to apex, almost three times length of apical ventral segment (.95 x
.35 mm.) ; posterior tarsal claws with four teeth, the basal one
postmedian, others successively large distad.
Length, to apices of elytra, 3.14 mm.; to tip of anal style,
4.09 mm.
One specimen: Type, male, Guadalcanal Lunga, March 4,
1935 (R. A. Lever).
66
PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 2
This species is most closely allied to maxillaris, new species,
described above, but may be separated by its darker color,
smaller size, and differences in the ridges of the posterior legs,
the peculiar maxillary palpi, and in the tarsal claws with a
smaller number of teeth. From consimilis Blair it may be sep-
arated by its larger size, longer antennae and the shorter tibial
ridges, in addition to the peculiar maxillary palpi. When the
female of this species is found it seems probable that sexual
differences in the shape of the maxillary palpi will be found to
exist.
Mordellistena oceanica Ray, new species
(Figs. 4, 7, 8)
Form elongate, sides subparallel, derm generally black, with
the following exceptions: front and clypeus castaneous, sides of
pronotum broadly rufo-castaneous, connected by a narrow, sub-
apical, transverse band (only in male!); legs flavocastaneous;
tibiae and femora of hind pair darker. Body densely covered with
fine, recumbent pubescence, generally partaking of ground color,
except on venter, where it is whitish, and on pronotum and elytra,
where it is condensed to form whitish bands, as follows, on pro-
notum, a thick, premedian line curved anteriorly to meet the an-
terior angles and extending along sides to base, a line along base
with three short vittae extending into disc; scutellum entirely;
elytra with a curved band surrounding humeri, a short, subbasal,
subsutural line curving inward to meet scutellar area; a median
area formed like an M, but broken at suture; a small, circular
area lateral to the previous one; a postmedian, inverted, M-shaped
area, reaching lateral margins, but broken at suture; and a short,
transverse, apical area.
Antennae 1 mm. long, elongate, narrow, filiform, reaching meso-
sternum; 4 one-third longer than 3; 5-10 each as long as 3 and 4
together; 11 distinctly longer than 10, sides curved, one distinctly
convex, the other almost straight, broadest premedially. Terminal
segment of maxillary palpi enlarged, with the form of an isosceles
triangle, distal margin transverse, straight.
Pronotum distinctly broader than long (.8-.65 mm.), anterior
margin and sides rounded, base arcuate, midbasal lobe short, broad,
rounded. Scutellum broad, rounded-triangular.
Elytra two and two-third times as long as broad (2.35 x .8 mm.) ,
sides gently curved from base to postmedian point, thence
strongly curved to apex, apices individually rounded. Anterior and
intermediate tarsi longer than their tibiae. Posterior tibiae with
three strong, equal, parallel ridges (excluding subapical one);
APRIL, 1944]
LINSLEY— PREY OF CALLINICUS
67
basitarsi with three, second and third segments with two ridges
each. Anal style almost three times length of apical ventral seg-
ment (1.2 X .45 mm.), attenuate to apex, slightly curved ventrad.
Length : to apices of elytra, 3 mm. ; to tip of anal style, 4.2 mm.
Five specimens: Type, male, a male paratype and two female
paratypes, Guadalcanal Berande, August 27, 1934 (H. T. Pad-
gen) ; one female paratype, same locality, August 26, 1934, on
wild fig (H. T. Padgen) .
This species belongs to the same group as the previous two
species, but may easily be separated by the difference in the rela-
tive lengths of the antennal segments, the striking pattern of
pubescent markings on the pronotum and elytra and the smaller
size.
PREY OF THE ROBBER FLY CALLINICUS CALCANEUS
LOEW
(Diptera: Asilidae)
BY E. GORTON LINSLEY
University of California, Berkeley
In May, 1942, near Oakhurst, Madera County, California, the
writer had several opportunities to examine the insects captured
by Callinicus calcaneus Loew. In this area the species was fly-
ing about over a hillside covered with mountain mizery, Cha-
maebatia foliolosa Benth., and was apparently preying almost
entirely upon bees which were visiting the blossoms of this plant.
Twenty-two specimens of the robber fly were captured with their
prey. The prey consisted entirely of megachilid and andrenid
bees as follows:
Species Specimens
Osmia brevis Cresson .... 9
Osmia cara Cockerell .... 5
Osmia densa pogonigera Cockerell . 1
Osmia lignaria Say .... 1
Andrena auricoma Smith ... 3
Andrena angustitarsata Viereck . . 1
Andrena carliniformis Cockerell . . 1
Andrena saccata Viereck ... 1
68
PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 2
Although the above sample is probably too small to be in-
terpreted as significant of the food preference of Callinicus cal-
caneus Loew, it is interesting to note that the genus O'smia repre-
sented nearly three-fourths of the prey examined, Andrena only
about one-fourth, although the seventeen species of Andrena
present individually outnumbered the eight or nine species of
Osnda by about five to one, as determined by analysis of 536
specimens captured by indiscriminate sweeping of the blossoms.
It might be pointed out, however, that all of the Osmia captured
are brilliant metallic green or blue, whereas the andrenids, with
the exception of the golden Andrena awricoma, are dull, incon-
spicuous species.
It is well known (Banks, 1913; Bromley, 1914-36; and Mc-
Atee & Banks, 1920) that many of our North American robber
flies prey primarily upon aculeate Hymenoptera, although most
of the records involve the honey bee and other social bees
{Bombus) and wasps [Vespa, etc.). Among the species which
do appear to prey to some extent on solitary bees may be men-
tioned Saropogon dispar Coq. (Halictididae, Andrenidae), Diog-
mites texanus Bromley (Halictidae) , D. Symmachus Loew (An-
thoporidae, Andrenidae), D. angustipennis Loew (Halictidae),
D. umbrina Loew (Andrenidae, Halictidae, Megachildae) , Proc-
tacanthus philadelphicus Macq. (Aifthophoridae, Andrenidae,
Halictidae), Promachus bastardi Macq. (Halictidae), P. fitchii
0. S. (Andrenidae, Megachilidae, Halictidae), and P. rufipes
Wied. (“solitary bees”) .
Literature C!ited
Banks, N. 1913. Asilids catching Hymenoptera. Proc. Ent. Soc.
Wash., 15:51.
Bromley, S. W. 1914. Asilids and their prey. Psyche, 21:192-198.
1923. Observations on the feeding habits of robber flies. Part
I. Proctacanthus rufus Will, and P. hrevipennis Wied. Psyche,
30:41-45.
1930. Bee-killing robber flies. Jour. N. Y. Ent. Soc., 38:159-175.
1934. The robber flies of Texas. Ann. Ent. Soc. Amer., 27:74-
110 .
1936. Asilidae feeding on bumblebees in New England. Psyche,
43:14.
McAtee, W. L. and N. Banks. 1920. District of Columbia Diptera:
Asillidae. Proc. Ent. Soc. Wash., 22:13-33.
APRIL, 1944]
BOHART— STENODYNERUS
69
STENODYNERUS FUNDATUS AND RELATED SPECIES IN
NORTH AMERICA
( Hymenoptera, Vespidae)
BY RICHARD M. BOHART
University of California, Los Angeles
The various species of the fundatus group of Stenodynerus
are among the commonest solitary wasps. Their range covers
most of North America as far south as northern Mexico. They
are all moderate in size, fairly slender, and possess the two pits
on the front face of the pronotum which are characteristic of the
genus Stenodynerus. They are more particularly distinguished
by having no acarinarium under the short first abdominal tergite,
by having a fine abundant silvery pubescence on the sides of the
thorax and by the fact that the third abdominal tergite has a
complete apical band.
Holotypes and allotypes of the new species and subspecies
have been deposited in the California Academy of Sciences.
Key to the Stenodynerus fundatus Group in America
North of Mexico
1. Posterior margin of pronotum not margined with yellow or
whitish along its entire length; pits on front face of pronotum
separated by less than a pit diameter 2
Posterior margin of pronotum margined at least narrowly
with yellow or whitish along its entire length; pits on front
face of pronotum separated by more than a pit diameter.... 10
2. Second tergite unspotted; abdominal markings deep yellow;
second and third tergites apically refiexed, particularly in
male, reflex of second tergite 2 to 3 times as high as that of
third; external surface of swollen apex of male hind tibia
with a distinct ridge near the middle foxensis
Second tergite spotted; male hind tibia not as above 3
3. Apical margin of third tergite hardly thicker medially than
that of fourth 4
Apical margin of third tergite much thicker medially than
that of fourth 6
4. Reflexed margin of second tergite about as thick as an ocellus
diameter; pronotal spots large and quadrate; markings whit-
ish, legs mostly reddish; female clypeus with apex about as
wide as length of fourth antennal segment; apical swelling
of hind tibia in male with a broad pubescent horizontal sur-
70
PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 2
face; terminal antennal segment in male strongly curved in
profile taos
Refiexed margin of second tergite not as thick as an ocellus
diameter; female clypeal apex about as wide as length of
third antennal segment; apical swelling of male hind tibia
slanting toward the inner leg surface; terminal antennal seg-
ment in male weakly curved in profile 5
5. Second sternite very finely and sparsely punctured latero-
basally; female with vertex pit usually indistinct, often hardly
larger than surrounding punctures ; female clypeus with
coarse striatiform punctures and often mostly yellow; apex of
hind tibia in male moderately and not abruptly swollen
claremontensis
Second sternite with moderately large, distinct, scattered
punctures latero-basally; female vertex pit large and distinct;
clypeus usually with fairly fine punctures and mostly or en-
tirely black in female; body markings often reddish; apex of
male hind tibia usually abruptly swollen Tnicrostictus
6. Refiexed margin of second and third tergites thickened about
equally; antennal fiagellum partly fulvous; markings whitish;
apex of male hind tibia swollen; female vertex pit larger
than an ocellus 7
Refiexed margin of second tergite two or more times as thick
as third; female vertex pit smaller than an ocellus and
relatively indistinct 8
7. Dorsum of thorax nearly impunctate xantianus
Dorsum of thorax well punctured phoenixensis
8. Antennal fiagellum partly fulvous; markings usually yellow;
male hind tibia distinctly swollen apically anatropus
Antennal fiagellum entirely dark; male hind tibia not swollen
apically 9
9. Markings whitish papagorum papagorum
Markings yellowish papagorum tinctifer
10. Second tergite without free spots, markings mainly yellow;
hind margin of second tergite 2 or more times as thick as an
ocellus diameter; male middle femur with a distinct tooth at
the end of a ridge; last antennal segment in male slightly bent
in profile ochrogonius
Without above combination of characters 11
11. Transverse carina of first sternite angled medially; male mid-
dle femur with a tooth or sharp angle at the end of a ridge;
last antennal segment in male distinctly bent in profile
fundatus
Transverse carina of first sternite forming a rounded or trun-
cate projection medially; male middle femur angled medially
but without a definite tooth; male last antennal segment
curved in profile , 12
APRIL, 1944]
BOHART— STENODYNERUS
71
12, With extensive reddish markings../t<.ndati/orm'is fundatiformis
With reddish markings restricted mainly to legs
fundatiformis gonosceles
Stenodynerus papagorum (Viereck)
Odynerus papagorum Viereck, 1908. Trans, Amer. Ent, Soc, 33:
394 (holotype, U. K.),
Distribution: Arizona (type locale), New Mexico, Colorado.
Stenodynerus papagorum tinctifer Bohart, new subspecies
Similar to typical papagorum except that markings are dis-
tinctly deep yellow instead of whitish yellow. Also, the second
and third tergites are somewhat less strongly reflexed,
Holotype male, allotype female, 10 male paratypes, and 6
female paratypes. Bear Valley, Santa Cruz County, Califor-
nia, July, 1913 (F. C. Clark). Other specimens apparently re-
ferable to this subspecies are from Salt Lake City, Utah;
Logan, Utah, and Jenny Lake, Wyoming.
Stenodynerus anatropus Bohart, new species
Male. Black, marked with yellow as follows: Mandible mostly,
clypeus, first antennal segment in front, interantennal spot, orbital
line, postocular spot, two spots on pronotum anteriorly, tegula
mostly, spot beneath, mesonotal spot, postscutellum mostly, pro-
podeum laterally, legs mostly, apical margin of first six tergites
and second to fifth sternites, lateral spots on first and second
tergites and second sternite. Inner surface of flagellum pale ful-
vous, hook black. Mandible apically, legs partly, tegula medially,
wing veins basally, reddish. Pubescence very fine, relatively in-
conspicuous, silvery. Puncturation moderate, sparse on clypeus,
very coarse toward apex of second tergite. Last antennal segment
slender; clypeus sharply and angularly incised apically; middle
femur with a prominent tooth, middle and hind femora apically
swollen. Second and third tergites reflexed apically, reflexed por-
tion of second about three times as high as that of third. Length
to apex of second tergite, 7 mm.
Female. Markings as in male except that mandible is mostly
blackish to red, clypeus has a dark margin and central spot, and
tibiae are reddish. Clypeus well punctured and obtusely incised
apically, the apical breadth about as great as length of third
antennal segment. Reflexion of second and third tergites weaker
than in male. Length of apex of second tergite 8.5 mm.
72
PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 2
Holotype male, allotype female, and 4 male paratypes, Davis
Creek, Modoc County, California, July, 1922 (C. L. Fox).
Other paratypes (all from California), 2 males and 1 female,
Buck Creek, Modoc County (C. L. Fox) ; 2 pair, Sequoia
National Park; 4 males and 1 female, Sunset Valley, Santa
Barbara County (M. Cazier) ; 2 males and 1 female, Altadena
(C. D. Michener) ; 6 males and 1 female. La Crescenta (C. D.
Michener and R. M. Bohart) ; 3 males and 6 females, San
Jacinto Mountains, May and June.
S. anatropus occurs commonly in California in the foothills
along the coast from Trinity County to Los Angeles County in-
cluding Santa Cruz Island and Mt. Diablo, along the foothills of
the Sierras, and in most of the southern California mountain
ranges from 2,000 to 5,000 feet elevation. I have also seen
specimens from Oregon, Washington, and Nevada.
Paratypes in collections of U. S. National Museum, American
Museum of Natural History, Oregon State College, California
Academy of Sciences, and J. Bequaert.
Stenodynerus claremontensis (Cameron)
Odynerus claremontensis Cameron, 1905. Invertebrata Pacifica
1 : 122 .
Distribution: Common throughout California (type locality,
mountains near Claremont). I have also seen specimens from
Nevada and Oregon.
Stenodynerus microstictus (Viereck)
Odynerus microstictus Viereck, 1906. Trans. Amer. Ent. Soc. 32:
199 (holotype U. K.).
OdynertLS gulielmi Viereck, 1908. Trans. Amer. Ent. Soc. 33:408.
Distribution: Kansas (type locality, Clark County), Texas,
Colorado, Nevada, Arizona, California (mostly in the southern
deserts) , northern Mexico, Lower California.
Stenodynerus phoenixensis (Cameron)
Ancistrocerus (Nortonia?) phoenixensis Cameron, 1908. Trans.
Amer. Ent. Soc. 34:225.
Distribution: Arizona (type locality, Phoenix), Colorado,
Utah, Nevada.
APRIL, 1944]
BOHART— STENODYNERUS
73
Stenodynerus TAOS (Cresson)
Odynerus toas Cresson, 1868. Trans. Amer. Ent. Soc. 1:381 (err.
typ.) (holotype, A.N.S.P.).
Odynerus taos Cresson, 1872. Trans. Amer. Ent. Soc. 4:243.
Odynerus cressoni Cameron, 1908. Trans. Amer. Ent. Soc. 34:198.
Distribution: New Mexico (type locality), Texas, South Da-
kota, Nebraska, Colorado, Arizona, Wyoming.
Stenodynerus foxensis Bohart, new species
Male. Black, clypeus, first antennal segment in front, inter-
antennal spot, mandible partly, orbital line, postocular spot, tegula
anteriorly, hind mesonotal angle, whitish yellow; front margin of
pronotum, legs partly, line across postscutellum, apical margin of
tergites 1 to 6 and sternites 2 to 6 (bands on 4-6 broken), yellow;
inner margin of flagellum, legs partly, wing veins basally, reddish;
wings lightly brown stained. Pubescence minute, dense, silvery.
Puncturation moderate, sparse on dlypeus, coarse on second tergite
subapically and on third sternite. Breadth of clypeus at apex as
great as 3/6 the length of antennal segment 3; last antennal seg-
ment large, stout, fitting into a groove in segment 10; interocellar
area slightly swollen; middle femur with a distinct tooth; middle
and hind tibiae apically sv/ollen, that of hind tibia with a sharp
external tooth-like ridge. Apical margins of second and third
tergites refiexed, that of second about 3 times as high as that of
third. Length to apex of second tergite 7.5 mm.
Female. Markings, pubescence, and puncturation about as in
male except as follows: Flagellum black, clypeus black save for a
basal spot, apical bands on sternites 4 to 6 complete, clypeus mod-
erately punctured. Depression of vertex about as large as an
occellus. Apical breadth of clypeus about 3/5 as great as length
of third antennal segment. Length to apex of second tergite 8 mm.
Holotype male, allotype female, 10 male paratypes, and 1
female paratype, Baboquivari Mountains, Arizona, July to Au-
gust, 1924 (collection of C. L. Fox, 0. C. Poling, collector) ; 1
male paratype, 14 miles east of Oracle, Arizona, July 27, 1924
(E. P. Van Duzee) ; 1 male paratype, Lowell Ranger Station,
Pima County, Arizona, July 20, 1916; 1 male paratype, Pata-
gonia, Arizona, August 8, 1940 (W. L. Swisher) ; 1 female para-
type, Willow Creek Mountains, New Mexico, July, 1933.
Paratypes in collections of U. S. National Museum, California
Academy of Sciences, and J. Bequaert.
74
PAN-PACIFIC ENTOMOLOGIST
[V0L.XX,N0. 2
Stenodynerus xantianus (Saussure)
Odynerus xantianus Saussure, 1870. Rev. Mag. Zool. 22:103.
Odynerus xanthianus Saussure, 1875. Smithsn. Misc. Publ. 254:
324.
Distribution: Common in many parts of Lower California
(type locality, Cape St. Lucas), also occurring uncommonly in
the southern deserts of California.
Stenodynerus fundatiformis (Robertson)
Odynerus fundatiformis Robertson, 1901. Trans. Amer. Ent. Soc.
27:200.
Distribution: Florida (type locality, Inverness) Georgia,
South Carolina, Mississippi, Arkansas, Texas.
Stenodynerus fundatiformis gonosceles Bohart, new subspecies
Black, marked with whitish yellow. Reddish markings re-
stricted mainly to legs. Clypeus well punctured; interantennal
tubercles present; posterior margin of pronotum yellowish; male
last antennal segment small and slender ; male middle femur angled
but hardly toothed medially; male hind tibia scarcely swollen;
apical margins of tergites 2 and 3 reflexed, that of two about as
high as an ocellus diameter, that of three less than half so high.
Wings brown stained, violaceous.
Holotype male, allotype, female, and 1 female paratype, Baby-
lon, Long Island, New York, June, 1936 (F. Blanton). Other
paratypes, Minnesota: 1 male, St. Anthony Peak (H. H. Knight) ;
1 male. Pine City (H. S. Telford). New Jersey: 1 male. Mill-
town; 1 pair, Lakehurst; 2 females. Browns Mills. Virginia: 1
male, Petersburg; 1 female. Cape Henry (L. Lacey). North
Carolina: 1 male, Bryson City (J. C. Crawford) ; 3 females.
Black Mountains (W. Beutenmuller) ; 2 females. Southern Pines.
Missouri: 1 female, Brandon (E. C. Van Dyke) ; 1 female,
Ozark Lake (E. C. Van Dyke). Alabama: 1 female, Tuskeegee
(R. H. Beamer). Arkansas: 1 female. Eureka Springs (E. C.
Van Dyke). Florida: 1 female, De Funiak Springs.
Paratypes in collections of U. S. National Museum, Univer-
sity of Kansas, California Academy of Sciences, and J. Bequaert.
Stenodynerus ochrogonius Bohart, new species
Male. Black, marked with whitish yellow as follows : mandible
basally, clypeus, scape in front, interantennal spot, orbital line.
APRIL, 1944]
BOHART— STENODYNERUS
75
postocular spot, front marg'in of pronotum broadly and hind mar-
g’in narrowly, tegula partly, spot beneath, postero-lateral angle of
mesonotum, line across postscutellum, legs partly, apical margins
of tergites 2 to 6 and sternites 2 to 4. Mandible apically, inner
surface of flagellum faintly, tegula and legs partly, wing veins
basally reddish. Wing membrane nearly transparent. Punctura-
tion moderate to coarse, moderate on clypeus, coarse toward apical
margins of abdominal segments 2 and 3. Pubescence fine and
silvery. Clypeus sharply two-pointed, moderately incised apically,
apical breadth about half the length of the third antennal segment.
Last antennal segment broad, flattened, somewhat bent in pro-
file. Weak interocular swellings present. Middle femur with a
strong tooth, hind tibia abruptly swollen apically. Second and
third tergites reflexed apically, reflex of second almost 3 times
that of third. Length to apex of second tergite 8.5 mm.
Female. About as in male with following exceptions: Clypeus
black save for a pale basal crescent, orbits with a spot only,
flagellum black, propodeum with a lateral yellow spot. Vertex pit
considerably larger than an ocellus. Apical breadth of clypeus
about as great as length of third antennal segment. Length to
apex of second tergite 9 mm.
Holotype male, allotype female, 31 male paratypes, and 17
female paratypes, Baboquivari Mountains, Arizona, June to
October, 1924 (C. L. Fox collection, 0. C. Poling, collector).
Other paratypes, 7 males and 11 females, Santa Rita Mountains,
Arizona (F. H. Snow, B, E. White) ; 5 males and 1 female. Oak
Creek Canyon, Arizona (G. E. and R. M. Bohart). I have also
seen specimens from PATAGONIA, SoNORiTA, Oracle, Chiricahua
Mountains, Santa Catalina Mountains, and Huachuca Moun-
tains, Arizona; Willow C^ieek Mountains, New Mexico; and
Brownsville, Texas.
S. ochrogonius is closely related to S. huasteca Saussure from
Mexico. The former is a more robust species, however, with the
second and third tergites more reflexed and the male last antennal
segment larger and less distinctly bent.
Stenodynerus fundatus (Cresson)
Odynerus fundatus Cresson, 1872. Trans. Amer. Ent. Soc. 4:242
(holotype, A.N.S.P.).
Distribution: Texas (type locality), Oklahoma, Missouri,
Alabama.
76
PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 2
A NEW SPECIES OF CICINDELA AND TWO NEW RECORDS
OF COLEOPTERA
BY BERNARD ROTGER
San Luis, Colorado
Cicindela theatina Rotger, new species
Head and thorax shining' green tinged frequently with coppery
lustre, anterior and basal impressions of thorax purplish. Elytra
dark reddish cupreous, opaque; at times greenish toward the base,
with basal reflection purplish or green. Legs and underside of
body metallic green. Labrum short, white, and more than three
times wider than long, with a small tooth. Head hairy between
the eyes, with many interocular striae; palpi pale, with last joint
green. Prothorax sparsely covered with long white hairs, sub-
quadrate, scarcely wider than long and convex. Elytra granulate-
punctate, punctures rather distant generally, and green, but more
closely spaced toward base; interspaces reddish cupreous; elytral
markings consist of humeral lunule almost reaching, and in sev-
eral instances actually reaching middle band and merging into it,
middle band obliquely bent with long extremity not hooked at tip,
but sometimes dilated, and apical lunule; all markings widely
united at margin. Palpi, four front trochanters, femora, sides of
abdomen as well as of thorax very hairy. Elytra more parallel in
male, wider and subparallel in female, apex being flnely serru-
late. Tarsal claws long. Length, including labrum, 12-14.50 mm.
Holotype male, allotype female, and topoparatypes in the
Rotger collection.
I believe that this species might be placed between Cicindela
lengi and C. limhata Say.
This species inhabits the edge of the Great Sand Dunes in the
San Luis Valley, Colorado, at an altitudef of about 8,200 feet,
where it was found by the author in three field trips, on the 26th
of June, the 28th of August, 1942, and on the 19th of May, 1943.
It was discovered on the side of Mosca Pass where the road ap-
proaches the Sand Dunes, in spots of scanty vegetation and where
the shfting sands begin. It was not found farther in the sand
where there was no vegetation. The beetle runs rapidly upon
the sand and it emits an orange liquid from its moiith. This
tiger-beetle emerges in August, when the most perfect specimens
can be obtained, and probably hibernates in the sand until
spring. By the reason of frequent sand storms to which this
species is subjected, the specimens caught in June lack most of
APRIL, 1944 ]
ROTGER— COLEOPTERA
77
the hair on the front and on the upper side of the thorax. During
the month of August a few specimens were taken belonging to
the preceding generation, and are of a dull green bronze.
On July 5, 1937, Mr. R. P. Allen discovered at Tuba City,
Arizona, Cicindela nevadica tubensis Cazier. Two years after,
on August o, 1 found four specimens of the same subspecies, at
Farmington, New Mexico; the following year on July 13, at
Canon Largo about eight miles from Blanco, New Mexico, I
Fig. 1. Elytral pattern of Cicindela theatina Rotger.
caught eleven specimens of the same. Mr. Mont A. Cazier who
identified this form informed me that it had never before been
taken in New Mexico.
Looking after butterflies, I was forced by the cloudy day to
pay more attention to Coleoptera, and luckily so, for I found
among small ants a few specimens of Fustiger knausi Schfr.
This is the first time that the Clavigeridae family has been col-
lected in Colorado. The species was found as I was descending
from Mesa Reservoir, near Durango, on April 30, 1941. Mr.
Alan S. Nicolay made the identification.
78
PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 2
HOST PLANT RECORDS OF WESTERN LYGUS
(Hemiptera, Miridae)
BY ROBERT L. USINGER
University of California, Davis
The pratensis group of the genus Lygus is rich in both speci-
mens and species in western North America. For several years I
have kept careful host records in an attempt to define the limits
of variation within species in this difficult group. The following
records appear to be new:
1. Lygus distinguendus Reuter. Three specimens of this species
were beaten from willow, Salix sp., Malahat Drive, Van-
couver Id., B. C., June 26, 1940 (R. L. Usinger). The variety
tahoensis Knight was taken on the same host on the trail to
Glen Aulen, Yosemite National Park, California, July 27,
1939 (R. L. Usinger).
2. Lygus hradleyi Knight. A long series was collected on Yel-
low lupine along the beach at Areata, California, July 8,
1940 (R. L. Usinger).
3. Lygus stratus Knight. Common on lupine at Miguel Mead-
ows, Yosemite National Park, California, July 11, 1941 (R. L.
Usinger) .
4. Lygus per plexus Stanger. Common on forest anemone. Ane-
mone sp., near timberline (5500 ft.). Paradise, Mount Rain-
ier, Washington, June 22, 1940 (R. L. Usinger).
5. Lygus convexicollis Reuter. This and the variety coloratus
Knight were common on Madia at Miguel Meadows, Yosemite
National Park, California, July 7, 1941 (R. L. Usinger).
6. Lygus usingeri Stanger. The type series was collected on
cream bush, Holodiscus discolor, on the trail to Nevada Falls,
August 14, 1938 (R. L. Usinger). No specimens have been
found earlier in the season. This and the following species
belong to the nubilus-nubilatus group. Nubilus Van Duzee,
as noted by Van Duzee, occurs on elderberry, Sambucus. I
have found it common on Sambucus in the Oakland Hills and
in the San Jacinto Mountains.
7. Lygus nubilatus Knight? Specimens which I have tentatively
placed here were collected sparingly on Spiraea douglasii in
the Blue Mountains, Oregon, July 4, 1940 (R. L. Usinger)
and along the Malahat Drive, Vancouver Id., B. C., June 26,
1940 (R. L. Usinger).
APRIL, 1944]
CHAMBERLIN— MYRIOPODS
79
SOME RECORDS OF MYRIOPODS COLLECTED BY
W. M. PEARCE IN CALIFORNIA
BY RALPH V. CHAMBERLIN
University of Utah, Salt Lake City
Following is a list of the species represented in a collection
of centipeds and millipeds made by Mr. W. M. Pearce from June
to August, 1941, in Mono, Inyo, Siskiyou and Shasta counties,
California.
Chilopoda
Otocryptops sexspinosus (Say).
Mono County: Monmouth Lakes, August 18, 1941.
Shasta County: Dickson’s Flat, July 21, 1941.
A species common throughout the state.
Tomotaenia laevipes (Wood).
Shasta County; Dickson’s Flat, July 31, 1941.
This large geophiloid, in life conspicuous because of its
bright red color, is also a form widespread in California and
Oregon.
Etho polys pusio (Stuxberg).
Mono County: Monmouth Lakes, June 25 and August 18.
Siskiyou County: July 21.
A species previously known only from the region about San
Francisco Bay.
Gosihius benespinosus Chamberlin.
Inyo County: Bishop Creek, South Fork, August 17, at ele-
vation, 10,000-11,000 feet.
Mono County: Montgomery Canyon, July 13.
Described originally from Bishop and Benton Creek, the
only previous records.
Pokahius pitophilus (Chamberlin).
Inyo County: Bishop Creek, South Fork, August 17.
Mono County: Convict Creek, June 20.
A species common in the Sierras about Truckee, from where
it was originally described.
Lo phobias castellopes (Chamberlin).
Siskiyou County: Bartle, July 21.
80
PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 2
Originally described from Mt. Shasta, the only record previ-
ously published.
Diplopoda
Calif orniidws vicinus Chamberlin.
Shasta County: Dickson’s Flat, July 21.
Five specimens, representing both sexes. These are the types
of the species.
Caliulus pearcei Chamberlin.
Inyo County: Bishop Creek, South Lake, August 17. Ele-
vation, 10,000 to 11,000 feet.
Nineteen specimens constituting the types of the species.
Caliulus furcifer (Harger)
Siskiyou County: Six miles south of McCloud, July 21.
This species is rather common over much of California and
Oregon.
A NOTE ON THE HIBERNATION OF MISCHOCYTTARUS
FLAVITARSUS IN A NEST OF DOLICHOVESPULA
ARENARIA.
On about February 1, 1944, a nest of Dolichovespula arenaria
(Fab.) was brought into the Division of Entomology and Para-
sitology, University of California at Berkeley. The nest contained
no live D. arenaria. This was to be expected at that time of year
since the colonies commonly die out from midsummer to early
fall and the nests are deserted through the winter. Oddly enough,
however, the comb contained a large amount of brood including
many individuals which had transformed to the adult stage but
had died before emerging. The cause of their death was un-
certain. They did not appear to have died of any disease. They
may have been killed by cold, or more likely, they were unable
to emerge due to lack of nourishment in the late larval stages
when the colony was declining. Another very interesting fea-
ture of this nest was the fact that it contained nearly a hundred
adults of Mischocyttarus flavitarsius (Sauss.). This unusual
number of M. flavitarsus no doubt represented females bibem-
ating in the abandoned nest. However, it is not uncommon to
find small groups of the overwintering females collected in pro-
tected places. — Ray F. Smith.
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Vol. XX
July, 1944
No. 3
THE
Pan -Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
BAILEY, REVIEW OF D. L. CRAWFORD’S THYSANOPTERA 81
ALEXANDER, UNDESCRIBED SPECIES OF TIPULIDAE FROM THE
WESTERN UNITED STATES. PART I 91
Db long, a NEW GENUS AND SPECIES OF MEXICAN LEAFHOPPER 98
HATCH, STUDIES ON NORTHWESTERN STAPHYLINIDAE I: ANTHO-
BIUM STEPH 101
BEER, NOTES ON OREGON ACMAEODERA 106
VAN DYKE, A NEW DOBSONFLY FROM CALIFORNIA 110
LINSLEY AND USINGER, NOTES ON HABITS OF FLAT BUGS WITH
A DESCRIPTION OF THE MALE OF ARADUS PATIBULUS Ill
TILDEN AND MANSFIELD, NOTES ON SPECIES OF CEONONYCHA.... 116
TILDEN, PHOEBIS SENNAE MARCELLINA IN SAN JOSE, CALIF 115
LINSLEY, HABITS OF AMOBIA (PACHYOPHTHALMUS) FLORIDENSIS 118
HEMMING, RECENT WORK BY THE INTERNATIONAL COMMISSION
ON ZOOLOGICAL NOMENCLATURE 119
SMITH AND MICHELBACHER, CLOVER LEAP WEEVIL 120
BOOK NOTICE 100
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1944
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The Pan-Pacific Entomologist
VOL. XX, No. 3
July, 1944
A REVIEW OF D. L. CRAWFORD’S SPECIES OF
THYSANOPTERA
BY STANLEY F. BAILEY
University of California, Davis
As a part of a study of California thrips it has been neces-
sary to carefully review all the previous work done in order to
accurately identify valid species and to avoid creating additional
synonymy when describing new forms. In former papers the
writer has reviewed other published works on western Thysanop-
tera. The present report covers the species described by D. L.
Crawford^.
The two genera, the sixteen species, and five varieties described
by Crawford are listed below in the order they appeared and un-
der the names assigned them. Ankothrips n^ gen., A. rohustus,
Aeolothrips longiceps, Euthrips minutus var. setosus, Phyllo-
thrips fasciculata, P. fasciculata var. stenoceps, Aeolothrips vespi-
formis, Heterothrips decacornis, Chirothrips mexicana, Euthrips
insularis var. reticulata, Rhaptothrips n. gen., R. peculiaris,
Anaphothrips longipennis, Euthrips cephalicus, E. cephalicus
var. reticulata, Dictyothrips reticulata, Thrips abdominalis,
Phloeothrips raptor, Liothrips umhripennis mexicana, Liothrips
bakeri, Liothrips mcconnelli, Anthrothrips variablis, Idolothrips
angusticeps.
Hood has studied types and cotypes and reported {in litt.)
upon the following species: Phyllothrips fasciculatus, P. fascicu-
latus var. stenoceps, Chirothrips mexicanus, Euthrips insularis
var. reticulata, E. cephalicus, E. cephalicus var. reticulata, Lid-
thrips bakeri, L. mcconnelli and Anthothrips variabilis. The syn-
onymy of these species is briefly given below:
^ Crawford, D. L., 1909, Some new Thysanoptera from Southern California,
Pomona Coll. Jour. Ent., 1 (4) :100-108 ; Some Thysanoptera of Mexico and the
South, ibid., 1(4) :109-119 ; Notes on California Thysanoptera, ibid., 1(4) :120-121,
1910, Thysanoptera of Southern California, ibid., 2 (1) :149-152 ; Thysanoptera of
Mexico and the South, ibid., 2(1) :153-170.
82
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX NO. 3
Haplothrips fasciculatus (D. L. Crawford)
1909. Phyllothrips fasciculata D. L. Crawford, Pomona Coll. Jour.
Ent. 1(4) : 105-108, fig. 48, A-H.
1909. Phyllothrips fasciculata var. stenoceps, idem, p. 108.
The complete synonymy has been given by Hood (Pan-Pac.
Ent. 3:174-175, 1927) and by Bailey (Pan-Pac. Ent. 13:90, 1937).
Chirothrips mexicanus D. L. Crawford
1907. Chirothrips crassus Hooker (nec Hinds), U.S.D.A. Bur.
Ent., Bull. 65:13.
1909. Chirothrips mexicana D. L. Crawford, Pomona Coll. Jour.
Ent., 1:114-115, fig. 51, A-H.
1920. Chirothrips fioridensis Watson, Fla. Ent. 4(2) : 2 1-22.
1923. Chirothrips fioridensis var. catchingsi Watson, Fla. Agr.
Exp. Sta. Bull. 168-76.
1927. Chirothrips mexicanus, Hood, Ent. News 38:112.
1928. Chirothrips mexicanus, Moulton, Proc. Haw. Ent. Soc. 7:
106-107.
1939. Chirothrips mexicanus, Andre, Proc. Ent. Soc. Wash. 41(6) :
197.
1939. Chirothrips mexicanus. Hood, Rev. de Entom. 10(2): 464*
466 (Brazil).
Frankliniella insularis (Franklin)
1908. Euthrips insularis Franklin, Proc. U. S. Nat. Mus. 33:715,
PI. 63, figs. 1, 2, 3, 5, 6, 7, and PI. 65, figs. 19, 24.
1909. Euthrips insularis var. reticulata D. L. Crawford, Pomona
Coll. Jour. Ent. 1:116.
1914. Frankliniella insularis. Hood, Proc. Ent. Soc. Wash. 16:38.
Frankliniella cephalica (D. L. Crawford)
1910. Euthrips cephalicus D. L. Crawford, Pomona Coll. Jour.
Ent. 2:153-155, fig. 63, A-H.
1910. Euthrips cephalicus var. reticulata D. L. Crawford, Pomona
Coll. Jour. Ent., 2:155.
1913. Euthrips tritici var. hispinosus Morgan, Proc. U. S. Nat.
Mus., 46(2008) : 10-11, figs. 17-18.
1914. Frankliniella cephalica. Hood, Proc. Ent. Soc., Wash., 16:
38, 42.
1915. Euthrips tritici var. projectus W'atson, Ent. News 26:51-52.
1919. Frankliniella hispinosus Watson, Fla. Buggist 3(l):4-7.
1923. Frankliniella tritici bispinosa, Watson, Fla. Ent. 7(1) :9-ll.
1923. Frankliniella tritici bispinosa, Watson, Fla. Agr. Exp. Sta.
Bull. 168:39.
1925. Frankliniella cephalica. Hood, Bull. Brooklyn Ent. Soc. 20:
72-73.
JULY, 1944]
BAILEY— THYSANOPTERA
83
Gynaikothrips uzeli (Zimmerman)
1900. Mesothrips uzeli Zimmerman, Bull. Inst. Bot. Buitenzorg,
7:12, fig. 4, i-iv, fig. 5, i, ii.
1908. Phlaeothrips ficorum Marchal, Bull. Soc. Ent. France, 14:
252.
1910. Liothrips hakeri D. L. Crawford, Pomona Coll. Jour. Ent.,
2(1) :161-163, fig. 67, A-D.
1913. Gynaikothrips uzeli, Hood, Ins. Ins. Mens. 1 (12) : 153-154,
Complete synon 3 rmy.
LePTOTHRIPS MALI (FiTCH)
1855. Phlaeothrips mali Fitch, Trans. N. Y. State Agr. Soc. 14:
806-808.
1902. Cryptothrips aspersus Hinds, Proc. U. S. Nat. Mus. 26:205-
206, 209.
1904. Criptothrips califomicus Daniel, Ent. News. 15:293-294.
1908. Phyllothrips aspersus, Hood, Canad. Ent. 40:305.
1909. Leptothrips aspersus. Hood, Ent. News 20:249-250.
1910. Liothrips mcconnelli D. L. Crawford, Pomona Coll. Jour.
Ent. 2:163-166, fig. 68, A-G.
The complete synonymy is given by the writer (Bailey, 1940,
Jour. Econ. Ent. 33(3) :543.
Haplothrips gowdeyi (Franklin)
1908. Anthothrips gowdeyi Franklin, Proc. U. S. Nat. Mus.
33(1590) :724-725, figs. 8, 15, 16, 23.
1910. Anthothrips variabilis D. L. Crawford, Pomona Coll. Jour.
Ent. 2:166-167, fig. 69, A-E.
1912. Haplothrips gowdeyi. Hood, Proc. Biol. Soc. Wash. 25:62.
Additional synonymy has been given by Hood (1918-21, Mem.
Queensland Mus. 6-7:121-130) and Moulton (1933, Rev. de Ent.
3(3) :389, Brazil).
The writer has previously studied the types of Ankothrips
rohustus D. L. Crawford, 1909 (Bailey, 1940 Pan-Pac. Ent.
16(3) :97-106) and Thrips abdominalis D. L Crawford, 1910
(Bailey, 1937, Canad. Ent. 69:121-126). The synonymy of valid
species has been completely reviewed in the foregoing references.
Crawford himself (1941, Pomona Coll. Jour. Ent. 2:149) sank
the variety setosus of Frankliniella minuta, Moulton), 1907.
Rhaptothrips peculiaris D. L. Crawford (Pomona Coll. Jour. Ent.
1:116-119, fig. 52, A-H) was early recognized as invalid by Bag-
nail (1910, Ann. Ent. Soc. Belg. 54:462) since the description was
84
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX NO. 3
founded on a larval form (see also Hood, 1912, Proc. Biol. Soc.
Wash. 25:61) Aeolothrips vespiformis D. L. Crawford (Pomona
Coll. Jour. Ent. 1:109-110, fig. 49, A-D) was placed in Frankli-
nothrips by Back (1912, Ent. News 23:73-77) and stands as the
type of this distinct genus (see also Hood, 1915, Ent. News
26:162-164).
The remaining species, Aeolothrips longiceps, Heterothrips
decacornis, Anaphothrips longipennis, Dictyothrips reticulatus,
Hoplandrothrips raptor, Liothrips mexicana, and Dicaiothrips
angusticeps, have not to the writer’s knowledge been previously
situdied and reported upon. Through the kindness of the Domin-
ion of Canada, Department of Agriculture at Ottawa these types
have been studied, illustrated, and redescribed as deemed neces-
sary to their clarification. ,
Aeolothrips kuwanaii (Moulton)
1907. Aeolothrips kuwanaii Moulton, U. S. D. A., Bur. Ent., Tech.
Ser. 12(3) : 47-48, PI. 1, figs. 5-8.
1909. Aeolothrips longiceps D. L. Crawford, Pomona Coll. Jour.
Ent. 1:101-103, fig. 46, A-G. New synonymy.
1913. Franklinothrips longiceps, Bagnall. 2nd. Int. Congr. Ent.
(Oxford, 1912), p. 397.
1913. Franklinothrips longiceps, Bagnall. Jour. Ec. Biol. 8:157-158.
1915. Aeolothrips kuwanaii. Hood, Ent. News 26:162.
1935. Aeolothrips longiceps, Bailey, Pan-Pac. Ent. 11(4) :163.
The male holotype when studied with carefully controlled
transmitted light exhibits the tracings of cross veins in the fore
wings. It has all the characteristics of Aeolothrips kuwanaii and
is definitely synonymous with Moulton’s species.
Heterothrips decacornis D. L. Crawford
1909. Heterothrips decacorni D. L. Crawford, Pomona Coll. Jour.
Ent. 1:110-114, fig. 50, A-I.
This species, represented on the type slide by two females, is
distinct from all other species in this genus in that it has the
scales on the posterior margin of the dorsum of the abdominal
segments fused together (see fig. 4).
The head and pronotum (fig. 7), and the third antennal seg-
ment (fig. 5) also have been illustrated from the dorsally
mounted specimen for further separation from other species.
The writer has not seen pectinifer Hood or mexicanus Watson
JULY, 1944]
BAILEY— THYSANOPTERA
85
1
6
PLATE I
Fig. 1. Head and pronotum of Anaphothrips longipennis D. L.
Crawford. 2. Dorsum of last abdominal segment of Ctenothrips
reticulatus (D. L. Crawford). 3. Antennal segments III and IV of
left antenna of Anaphothrips longipennis. 4. Left half of dorsum
of abdominal segment VI of Heterothrips decacornis D. L. Craw-
ford. 5. Antennal segment III of H. decacornis. 6. Terminal ab-
dominal segments of Anaphothrips longipennis. 7. Head and pro-
notum of H. decacornis.
Scale: Figure 3, 0.01 mm.; figure 6, 0.1 mm. Remaining figures
at same magnification as figure 6.
86
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX NO. 3
which are apparently the closest relatives of decacornis but,
based on published descriptions, the very short terminal bristles
on the scales of the comb separate it from mexicanus and lyoniae
Hood (see also Hood, 1916, Proc. Biol. Soc. Wash. 29:110-113).
Anaphothrips longipennis D. L. Crawford
1910. Anaphothrips longipennis D. L. Crawford, Pomona Coll.
Jour. Ent. 2:150-152, fig. 62, A-D.
1911. Anaphothrips zeae Moulton, U. S. D. A., Bur. Ent., Tech.
Ser. 21:41, PI. 5, figs. 31-34. New synonymy.
Two female specimens are present on Crawford’s type slide;
one severely treated in eaustic, is mounted dorsally and the other
ventrally. As is shown in figure 6 the cleared specimen shows an
incomplete comb on the posterior dorsal margin of the eigthth
abdominal segment. The ventrally mounted specimen has a fully
developed comb as is the usual case in the species. Crawford’s
original description did not mention this comb nor did he
illustrate it. Moulton in his review of the North American species
omitted longipennis (although other species of Crawford are
listed) and did not compare zeae with it. The writer has studied
a long series of this common species (ineluding macropterous
males and brachypterous forms of both sexes) and notes that in
some specimens the comb is very sparse and the number of
bristles of the vein of the fore wing is variable (as are all the
minor bristles on the head and pronotum) . One female speci-
men has forked sense cones on the third antennal segment! Fig-
ure 1 illustrates the dorsally mounted specimen of longipennis
as it actually appears on the type slide and figure 3 the third
and fourth antennal segments. There is no doubt that the two
species are synonymous when normal variation is considered.
Ctenothrips reticltlatus (D. L. Crawford)
1910. Dictyothrips reticulata D. L. Crawford, Pomona Coll. Jour.
Ent. 2:155-157, fig. 64, A-D.
1911. Dictothrips reticulatus, Moulton, U. S. D. A., Bur. Ent.,
Tech. Ser. 21:24.
1923. Ctenothrips reticulatus, Watson, Fla. Agr. Exp. Sta. Bull.
168:36.
This species is represented by the holotype female (which is
accompanied by several specimens of Frankliniella cephalica on
the same slide). The lack of reticulation on the last abdominal
JULY, 1944 ]
BAILEY— THYSANOPTERA
87
PLATE II
Fig. 8. Head and pronotum of male of Elaphrothrips angusti-
ceps D. L. Crawford. 9. Antennal segments III, IV, and V of left
antenna of Liothrips mexicanus D. L. Crawford. 10. Head and
pronotum of Hoplandrothrips raptor D. L. Crawford. 11. Head
and pronotum of L. mexicanus.
Scale: Length of lines at right of figures equals 0.1 mm. Fig-
ure 11 is at same magnification as figure 10.
88
THE PAN-PACIFIC ENTOMOLOGIST
[VOL.XX NO. 3
segment (see fig. 2) separates reticidatus from bridwelli Frank-
lin (1907, Ent. News 18:247-250). C. reticidatus shows afl&nities
with both Taeniothrips and Odontothrips ; the reticulations on
the pronotum and particularly the dorsal ridge anterior to the
ocelli, however, readily separate it from group II, D. of Taenio-
thrips in Steinweden’s key. (1933, Trans. Amer. Ent. Soc. 59:
269-293) .
Hoplandrothrips raptor (D. L. Crawford)
1910. Phloeothrips raptor D. L. Crawford, Pomona Coll. Jour.
Ent. 2:159-161, fig. 66, A-E.
1915. Hoplandrothrips raptor, Hood, Entom. 48:102-107.
Crawford’s illustration is of the head of the male of H. raptor
and not of the female. The latter sex is illustrated in this paper
(fig. 10) from the type slide which has on it one specimen of
each sex, both ventrally mounted. The species in the genus
Hoplandrothrips are most difl&cult to separate and Hood (1927,
Entom. Amer. 7(n.s.) :230) states that “for some years I have
busied myself in studying the species alive in the laboratory,
and in rearing them; but the results accomplished lie more in
the number of slides available for study than in any definiteness
of concept regarding specific limitation.”
From the limited number of species of this group in the
writer’s collection, and the published descriptions of the others,
raptor appears to be valid (Hood, 1941, Rev. de Ent. 12(3) :556,
Brazil ) . It seems desirable, however, to supplement the original
description as follows: The eyes are prolonged much more dors-
ally than venfrally (see fig. 10) . The number of auxiliary bristles
on the posterior margin of the fore wing is nine on the female
and twelve on the male. At the base of the fore wing, on the
anterior margin there are three strong bristles, all dilated at the
tip on the female and on the male only the inner two dilated.
As in most thrips of this group, the second antennal segment is
darker than the first. The sense cone formula is as follows: HI,
1+2; IV, 2+; V, 1+1; VI, 1 + 1 (+1?). Faint reticulation can
be seen in the center of the lateral margins of abdominal seg-
ments one to six. The measurement of the principal bristles on
the female are (in mm.) po, 0.0561; on the pronotum aa, 0.0594,
nl, 0.0561, pa, 0.0726.
July, 1944 ]
BAILEY— THYSANOPTERA
89
Liothrips mexicanus D. L Crawford
1910. Liothrips umbripennis var. mexicana D. L. Crawford, Po-
mona Coll. Jour. Ent. 2:161.
1918-21. Liothrips mexicanus, Hood, Mem. Queensland Mus. 6-7 :
132.
Hood raised mexicanus to specific rank in the above reference
but did not expand its description or give any comparison with
umbripennis or other species. Since Crawford devoted only a
few lines to the original description (without illustration) it
seems desirable to more fully describe and illustrate (figs. 9 and
11) mexicanus from the holotype female (ventrally mounted and
partly cleared).
Female (macropterous) . Length 2.288 mm. Color uniform dark
brown. Antennal segments I and II concolorous with the body, III
and basal half of IV clear yellow, remaining segments brown. All
legs and tarsi brown. Wings brown in the basal half becoming
gradually lighter toward the tip, marginal bristles light brown.
Head about 0.247 mm. in length and 0.221 mm. in width behind
the eyes. Mouth cone long and pointed extending beyond the pro-
sternum. Cheeks nearly straight and faintly notched at margin.
Dorsum of head with fine lateral striations near the posterior.
Three ocelli present and compound eyes extending posteriorly
somewhat more on ventral side than on dorsum. One postocular
bristle, blunt at the tip. Sense cones on antennae as follows:
III, 0-M; IV, 1-^2; V, 1 + 1 (-^1).
Pronotum 0.104 mm. in length at center and 0.312 mm. in width
at posterior angles; anastomosing lateral striations present on
posterior. Four bristles on anterior margin, one mid-lateral, one
at each posterior-lateral angle, and two somewhat forward of
posterior margin. All bristles blunt at the tips. Fore femora en-
larged and an appressed claw present on fore tarsus. Three strong,
blunt bristles equal in length present on anterior margin of fore
wing near its base. Fourteen interlocated or auxiliary bristles on
posterior margin of fore wing.
Measurements (in mm.) : Length of abdomen, 1.417 (including
tube); length of tube, 0.195; postocular bristle, 0.079; midlateral
bristles, 0.062; posterior-angular bristles, 0.112; antennal seg-
ment I, 0.049; II, 0.066; III, 0.095; IV, 0.089; V, 0.085; VI, 0.072;
VII, 0.066; VIII, 0.033; total length, 0.557.
This expanded description is based on the type, slide No. 376,
of the Canadian National Museum collected at Guadalajara,
Mexico, by D. L. Crawford.
Because of the size of the genus Liothrips and the difficulty of
90
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX NO. 3
determining species from some of the too brief, older descrip-
tions (often inadequately illustrated), the writer is at present
unable to give any synonymy for L. mexicanus. Moulton’s
dumosa (also from Oak) appears closely related but this species
likewise has never been clarified.
Elaphrothrips angusticeps (D. L. Crawford)
1910. Idolothrips angtisticeps D. L. Crawford, Pomona Coll. Jour.
Ent. 2:168-170, fig. 70, A-G.
1912. Idolothrips angusticeps, Hood, Proc. Biol. Soc. Wash. 25:62.
1923. Dicaiothrips angusticeps, Watson, Fla. Agr. Exp. Sta. Bull.
168:72.
1929. Elaphrothrips angusticeps, Moulton, Pan-Pac. Ent. 6(1): 12.
This species has all the characteristics of the genus Elaphro-
thrips Andre (1940, Proc. Ent. Soc. Wash. 42(4) :75-76) and is
apparently distinct (based on the descriptions) from the three
other Mexican species in this group: alhospinosus Moulton and
teller Pr. are much smaller in size; dampfi Hood has much
shorter interocular and postocular bristles; Crawford, however,
did not state upon which sex his measurements were based.
The type slide has mounted upon it two females and one
male, all of which are badly broken up. The most intact speci-
men (male) has been illustrated in fig. 8 as it actually appears.
Crawford based his description on fifteen females and seventeen
males from Belize, Havana, Cuba, San Marcos and Chinandega,
Nicaragua, and Guadalajara, Mexico. Not all of this material
has been available for study and it is quite possible that other
species are among them. A detailed study of a series of the
Central American species of this group should readily establish
the validity of angusticeps.
Of the two genera described by D. L. Crawford, Ankothrips is
a well-established, valid genus. Of the twenty-one species and
varieties described by him twelve are, as far as can be ascer-
tained at present, valid. It is hoped that the new illustrations
and expanded descriptions of those species in the more difiicult
genera will enable other workers to more accurately place these
and other related North American forms. With the exception of
the type of Chirothrips mexicanus (Hood, 1927, Ent. News 38:
122; 1939, Rev. de Ent. 10(2) :464-466 Brazil), all of D. L.
Crawford’s type material is at present deposited at Ottawa,
Canada.
JULY, 1944]
ALEXANDER— WESTERN TIPULIDAE
91
UNDESCRIBED SPECIES OF TIPULIDAE FROM THE
WESTERN UNITED STATES
(Diptera)
PART I
BY CHARLES P. ALEXANDER
MassachAisetts State College, Amherst, Mass.
The novelties discussed herewith are from Oregon and Wash-
ington, the former having been taken chiefly by Professor James
A. Macnab, of Linfield College, McMinnville, Oregon, and by
students of Professor Scullen and Mr. Post, of the Oregon State
College. The Washington specimens were generously given to me
by the collector, Mr. B. Brookman. Where not indicated to the
contrary, the types are preserved in my collection of World
Tipulidae.
Limonia (Dicranomyia) illustris Alexander, new species
General coloration bufFy yellow, the praescutum without distinct
darker stripes; rostrum light yellow; antennae with scape obscure
yellow, the remaining segments brown; posterior sclerites of
notum darkened, more or less pruinose; halteres with stem whit-
ened, knob dark brown; legs long and slender, yellow, the terminal
tarsal segments black; wings light yellow, the oval stigma dark
brown, very conspicuous; Rs a little less than twice the basal
section of R4+5; abdominal tergites brown, paling to yellow on sides.
Female. Length about 8 mm.; wing 8.8 mm.
Rostrum light yellow, nearly one-half the remainder of head;
palpi with basal segment yellow, the remainder dark brown. An-
tennae with scape obscure yellow, remainder brown; flagellar seg-
ments oval, rather strongly constricted at the sutures; verticils
shorter than the segments; terminal segment pointed, only a trifle
longer than the penultimate. Head with front and anterior vertex
yellow; posterior vertex darkened and heavily pruinose; anterior
vertex (female) nearly three times the diameter of scape.
Pronotum brownish yellow. Mesonotum brownish yellow, the
central portion of praescutum weakly darkened but not forming
distinct stripes; posterior sclerites of notum similarly darkened.
Pleura chiefly buffy, the posterior sclerites, and the pleurotergite
more pruinose. Halteres with stem whitened, the knob abruptly
dark brown. Legs with coxae and trochanters yellow; remainder
of legs long and slender, yellow, the terminal tarsal segments
passing into black. Wings strongly tinged with light yellow, the
prearcular and costal fields clearer yellow; a large and very con-
92
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX NO. 3
spicuous, oval, dark brown stigma; veins brown, more yellowish
toward the wing base and in costal field. Venation: Sc moderately
long, Sci ending shortly beyond origin of Rs, Sc 2 opposite origin of
Rs, the latter a little less than twice as long as basal section of
Ri+n; free tip of Sc 2 and 2^2 in transverse alignment; cell 1st Mz
about as long as vein Mz beyond it; m-cu long, gently sinuous, at
or just beyond the fork of M; anal veins nearly parallel at their
origins.
Abdominal tergites brown, paling to yellow on sides; stemites
fulvus yellow. Ovipositor with genital shield yellow; valves deep
reddish horn color; cerci slender, rather strongly upcurved to the
acute tips; hypovalvae with a black spot at base.
Habitat. Oregon (Yamhill County).
Holotjpe, 2, Carlton, Meadow Lake Road, June 5, 1942
(Fender) ; Alexander collection, through Professor Macnab.
By Doane’s key to the Nearctic species of the subgenus (Ent.
News, 19:5-7; 1908), this fly runs to Limonia {Dicranomyin)
citrina (Doane), an entirely different species. In its general
appearance, especially the conspicuous wing stigma, the fly is
more like L. {D.) ctenopyga Alexander or L. (D.) melleicauda
(Alexander), but is quite distinct. Both of these latter forms
have the male hypopygium unusually complex and it is possible
that the male of the present fly will show a comparable condition.
Besides the type, three further females from the same date and
place were sent by Professor Macnab. These differ sufi&ciently
from the type to make me unwilling to consider them as being
conspecific, while at the same time agreeing in most details. In
these specimens, three praescutal stripes are more or less devel-
oped and vein 5ci is definitely longer.
Oxydiscus (Oxydiscus) pacificus Alexander, new species
Size large (wing, male, over 5 mm.) ; general coloration of tho-
rax yellow, virtually unpatterned; antennae dark brown through-
out; tibiae spurred; wings subhyaline, the stigmal area vaguely
darkened; relatively abundant macrotrichia in outer cells of wing;
cell Ml present; male hypopygium with the basistyle produced
beyond the point of insertion of the dististyles as a long, obtusely-
pointed lobe that is provided with unusually strong black setae;
blades of gonapophyses relatively narrow; spines of ventral fork
of aedeagus relatively short and straight, exceeding the short
straight aedeagus.
Male. Length about 4.5 mm.; wing 5.4 mm.
Rostrum light brown; palpi darker. Antennae dark brown
throughout, 16-segmented ; basal three flagellar segments enlarged,
JULY, 1944]
ALEXANDER— WESTERN TIPULIDAE
93
the second and third more or less fused, possibly indicating the
beginnings of a fusion segTnent; succeeding segments passing
through oval to elongate-oval, with verticils that much exceed the
segments. Head dark brown; anterior vertex about three times
the diameter of scape.
Thorax almost uniform polished yellow, the central portion of
praescutum a trifle more darkened but not forming an evident
stripe. Halteres elongate, yellow, the knobs darkened. Legs with
the coxae and trochanters pale yellow; remainder of legs brown-
ish yellow, the terminal tarsal segments blackened; tibial spurs of
middle and hind legs long and distinct (fore legs broken) ; claws
simple. Wings whitish subhyaline, the stigmal area vaguely dark-
ened; veins obscure yellow; trichia dark. Macrotrichia of outer
cells relatively abundant, from cells to Cu, inclusive. Venation:
Sci ending just before level of fork of Rs, Scz about one-third as
long as Sci', Rs long; i? 2 + 3+4 about two and one-half times the basal
section of Rs; R2 faint, apparently placed a little more than its own
length beyond fork of i?2+3+4; cell Mi present, about one-third its
petiole; m-cu beyond mid-length of cell 1 st M2,
Abdomen dark brown, the posterior borders of the more proxi-
mal sternites paler; hypopygium yellow. Male hypopygium with
the basistyle produced beyond point of insertion of the dististyles
as a strong, stout lobe, much longer and stouter than in ameri-
caniis, obtuse at apex, provided with unusually strong and power-
ful blackened setae. Outer dististyle unusually slender, at apex
with a major axial spine, with a smaller subapical outer spine and
a series of microscopic denticles on inner or lower margin before
apex. Inner dististyle very obtuse at apex. Gonapophyses appear-
ing as relatively narrow, subcultriform blades, narrower than in
americamis. Ventral fork of aedeagus with the spine widely sep-
arated at base, nearly straight, simple and unusually short. Aede-
agus shorter and stouter than either spine, in americamis longer
than the spine and gently sinuous.
Habitat. Oregon (Yamhill County).
Holotype, $ , Carlton, Meadow Lake Road, June 5, 1942
(Fender) ; Alexander Collection, through Professor Macnab.
The present species is the first member of the genus to be
discovered in the western Nearctic region. It is most similar to
the eastern Nearctic Oxydiscas {Oxy discus) americanus (Alex-
ander), differing especially in the larger size, body coloration,
and, especially, the structure of the male hypopygium, as com-
pared above. All of the known American species show strong
hypopygial characters. Besides the four known at present from
the eastern Nearctic, a few others are now known from Mexico,
southward to Ecuador.
94
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX NO. 3
Dactylolabis postiana Alexander, new species
General coloration brownish black, the praescutum yellow polli-
nose, with four very distinct blackened stripes; halteres yellow;
wing:s pale yellow, with a very heavy brown pattern, the total
amount of dark being: approximately equal to the pale ground;
dark areas in part ocelliform; marginal areas at ends of longitu-
dinal veins, lacking only on veins Rs and Mi, becoming very large
in the cubital and anal fields; a supernumerary crossvein in cell
Ri, in transverse alignment with R^', 222 + 3+4 longer than m^cn.
Female. Length about 8 mm.; wing 8 mm.
Rostrum and palpi black, the former sparsely pruinose. An-
tennae black; flagellar segments oval, the two or three proximal
flagellar segments with short pale apical necks; longest verticils a
little shorter than the segments. Head dark brown, the front,
posterior orbits and genae light gray.
Pronotum brown, with a narrow median blackish line, the poste-
rior border of sclerite more extensively blackened. Mesonotal
praescutum yellow pollinose, with four very distinct black stripes,
the intermediate pair narrow; scutal lobes extensively blackened;
scutellum and postnotum pruinose, the ground color of the former
dark brown. Pleura and pleurotergite heavily gray pruinose;
dorsopleural membrane brown behind the spiracle, with a yellow-
ish spot immediately ventrad of the latter. Halteres pale yellow.
Legs with coxae dark brown, heavily pruinose; trochanters obscure
yellow, their apices narrowly ringed with black; femora, tibiae
and basitarsi obscure brownish yellow, narrowly blackened at tips;
remainder of tarsi black. Wings pale yellow, with a very heavy
brown pattern, the total amount of dark being approximately equal
to the ground; dark areas in part ocelliform, dark with paler
centers, especially those at cord, outer end of cell 1st Mi and at
wing-margin ; dark areas distributed as follows : A series of five
or six in cell C; major marks at arculus, origin of Rs, before outer
end of Rs, tip of Sc and over the fork of Ri+s+i, over Ri+i, Ri and
over the supernumerary crossvein in cell Ra', major marginal areas
at ends of all longitudinal veins excepting Rs and Mi, from vein
Ml backward becoming progressively larger, the largest of all in
the axillary field, including more than the proximal half of cell
2nd A, extending into cell 1st A; other areas along cord, outer end
of cell 1st Ml, fork of Mi+i, on distal third of vein M and along vein
1st A at near mid-length; veins brownish yellow, darker in the
patterned areas. Venation: Sci ending about opposite the fork
of 222+3 4, Sci at its tip ; 222 + 3 ++ relatively long, about one-third
longer than m-cu‘, a supernumerary crossvein in cell Ra, in trans-
verse alignment with vein Ri’, cell 1st Mi moderately long, sub-
equal to vein M^ beyond it; cell Mi about one-third longer than its
petiole; m-cu just beyond the fork of M.
Abdominal tergites black, their discal portions slightly paler;
genital shield dark castaneous; sternites brownish black, the cen-
JULY, 1944]
ALEXANDER— WESTERN TIPULIDAE
95
tral portions of the intermediate segments broadly obscure yellow.
Ovipositor with cerci compressed-flattened, relatively wide, the
tips subacute.
Habitat. Oregon (Columbia County).
Holotype, ?, Vernonia, April 1, 1938, on willow (K. Gray
and J. Schuh) ; type in collection of Oregon State College.
I am pleased to name this species for Mr. R. L. Post, to whom
I am indebted for numerous favors in the past. This is one of
the most distinct species of the genus so far made known. It is
readily distinguished from all other Nearctic species by the
very heavy dark wing pattern that is in part ocelliform, with
pale centers.
Cladura macnabi Alexander, new species
General coloration of thorax yellow, variegated with brown,
including a single median praescutal stripe; dorsal mesopleura
darkened ; fusion-segment of antennae short, about equal in length
to the two succeeding segments combined; halteres unifo-rmly yel-
low; wings whitish subhyaline, vein Cu darker; vein very
faintly indicated; cell 1st Mi large, about as long as vein Ms be-
yond it; cell Ml deep, about three times its petiole; male hypo-
pygium with a single dististyle, this provided at apex with short
blackened pegs; gonapophyses narrowed outwardly, produced into
a very delicate, hyaline, spinous point.
Male. Length about 6 mm.; wing 6.8 mm.
Rostrum brown, palpi dark brown. Antennae with scape yel-
low; remainder of organ dark brown; fusion-segment cylindrical,
about as long as the succeeding two segments combined and evi-
dently formed of two such segments; succeeding segments oval to
long-oval, with very long verticils. Head above clear gray, paling
to brown on the genae and beneath; anterior vertex relatively
broad, nearly four times the diameter of scape; eyes with course
ommatidia.
Pronotum brown above, paler on sides, provided with a few
long conspicuous setae. Mesonotal praescutum yellow, with a
single dark brown stripe, this broad and conspicuous, not reaching
the suture behind; scutum obscure yellow, each lobe with two
brown areas, the more cephalic one larger; scutellum and medio-
tergite chiefly dark brown; pleurotergite and lateral portions of
mediotergite obscure yellow. Pleura yellow, including the dorso-
pleural membrane; conspicuous brown spots on dorsal anepi-
sternum and dorsal pteropleurite. Halteres relatively long, entirely
pale yellow. Legs with the coxae and trochanters yellow; re-
mainder of legs yellow, the tarsi infuscated. Wings whitish sub-
hyaline, virtually unpatterned, variegated only by the more dark-
ened cubital vein and faintly darkened adjoining portions of the
membrane; no stigmal darkening; remaining veins chiefly pale.
96
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX NO. 3
those beyond cord a trifle darker. Venation: very faint, at near
mid-length of upper branch of Rs, R2+3 thus slightly less than twice
the arcuated i22+s+4; Sc2 near tip of Sci; cell 1st M2 large, about as
long as vein M3 beyond it; basal section of Ms oblique, about twice
m; cell Mi deep, about three times its petiole; m-cio just beyond
fork of M; vein 2nd A short, ending before level of origin of Rs.
Abdominal tergites dark brown; basal sternites yellow, the
outer segments slightly more infuscated, with paler caudal bor-
ders; eighth and ninth segments blackened to form a conspicuous
subterminal ring. Male hypopygium with the basistyle relatively
slender, at apex produced into a lobe that is about one-fifth as
long as the dististyle. Dististyle single, long and gently curved to
the blunt tip that is set with short blackened pegs. Gonapophyses
appearing as flattened blades, narrowed apically, the tip narrowed
into an unusually delicate hyaline spinous point.
Habitat. Oregon (Yamhill County) .
Holotype, S , McMinnville, October 3, 1936, shrub associa-
tion (Macnab) ; Collector’s No. 18; Alexander Collection,
through Professor Macnab.
This distinct species is dedicated to the collector. Professor
James A. Macnab, to whom 1 am much indebted for many inter-
esting species and records of Tipulidae from Oregon. The fly
is very different from the other regional members of the genus,
especially in the venation and the structure of the male hypo-
pygium. The wing venation is much as in Pterochionea Alex-
ander but the fusion-segment of the antennae is short, as in
Cladura. In the light of this new discovery, it seems possible
that Pterochionea will best be placed as a subgenus of Cladura
Osten Sacken.
Gonomyia (Idiocera) brookmani Alexander, new species
Belongs to the blanda group; allied to calif ornica; rostrum
darkened; antennae with scape blackened on lower face, yellow
above; sublateral praecsutal stripes present; thoracic pleura con-
spicuously striped longitudinally with yellowish white on a brown-
ish gray ground; wings patterned with brown, as in the group;
Ri +2 and R3 very closely approximated to confluent at wing margin;
abdomen brownish black, the segments conspicuously ringed caud-
ally with pale yellow; male hypopygium with the outer lobe of
basistyle pale throughout, terminating in an acute pale spine;
outer and intermediate dististyles blackened, both obtuse at tips.
Male. Length about 5.2 mm.; wing 6 mm.
Female. Length about 5.5 mm.; wing 5.8-6 mm.
Rostrum blackish gray; palpi black. Antennae with scape
blackened beneath, yellow above; pedicel and flagellum brownish
black to black. Head above obscure yellow, with a restricted dark
area on vertex.
JULY, 1944]
ALEXANDER— WESTERN TIPULIDAE
97
Pronotum dark brown above, with an obscure yellow median
spot behind, the lateral borders lig'ht yellow. Mesonotal praescu-
tum clear g’ray, with elongate intermediate dark brown stripes,
the sublateral pair narrow and poorly indicated; humeral and
lateral portions of sclerite light yellow; pseudosutural fovea cas-
taneous; posterior sclerites of notuni gray, the scutal lobes with a
darkened area on their mesal portion; posterior callosities of
scutum yellow; postnotum gray, the dorsal portion of pleuro-
tergite pale whitish yellow, crossing the suture onto the cephalo-
lateral portion of the mediotergite. Pleura dark brown, pruinose,
conspicuously patterned with yellowish white, including a conspic-
uous ventral stripe that expands behind onto the metapleura; dorso-
pleural area variegated with yellow. Halteres yellow, knobs black-
ened. Legs with the fore and middle coxae pale yellow, the poste-
rior pair more darkened basally; trochanters obscure yellow;
femora obscure yellow, with a nearly terminal dark brown ring;
tibiae obscure yellow, the tips narrowly blackened; tarsi passing
into black. Wings subhyaline, the prearcular and costal fields
somewhat more whitened; a conspicuous brown pattern, arranged
as in the group, including marks at h, arculus, origin of Rs, fork
of Sc, stigma, cord, fork of Mm and in outer end of cell Rz', in
more heavily patterned specimens, the margin of cell Rt is dark-
ened; veins brown, yellow in the brightened fields. Venation: So
relatively long, Sci ending about opposite mid-length of Rs, Sc2
near its tip; Rm and R3 very closely approximated to confluent at
margin; m-cw from one to one and one-half times its length before
fork of M.
Abdomen brownish black, sparsely pruinose, the segments con-
spicuously ringed caudally with pale yellow. Male hypopygium
with the outer lobe of basistyle terminating in a short acute pale
point, much shorter than in calif o'i'nica. Outer and intermediate
dististyles both heavily blackened and blunt at their tips, the lat-
ter ones abruptly though slightly dilated at apex; inner style
broadly flattened, its tip truncated. Aedeagus relatively narrow,
entirely pale, the tip a short recurved point.
Habitat. Washington (Yakima County) .
Holotype, $ Naches, June 5, 1941 (Brookman). Allotopo-
type, 9, pinned with type. Paratopotypes, 1 ^ , 3 9 9, with the
types.
The species is named for Mr. B. Brookman, to whom I am
indebted for many interesting Tipulidae from the Yakima Valley.
The nearest relative of the fly is Gonomyia {Idiocera) californica
Alexander, of California, which differs most evidently in the
structure of the male hypopygium especially the long terminal
spine of the outer lobe of the basistyle, and the acutely pointed
intermediate dististyle.
98
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX NO. 3
A NEW GENUS— PREMANUS— AND SPECIES OF MEXICAN
LEAFHOPPER RELATED TO OMANANA
(Homoptera, Cicadellidae)
BY DWIGHT M. DeLONG
Department Zoology and Entomology, Ohio State University
Premanus new genus
Related to Omanana. The vertex is short and broad, round-
edly produced but appearing parallel margined. The vertex is
distinctly depressed just above margin, the margin being em-
phasized by the depressed line. The first anteapical cell is quite
short, the second is long. The male plates are long and tapered
at apex as in OshorTiellus. The aedeagus is simple and not com-
posed of paired structures as in Omanana.
Genotype Premanus hebatus DeLong.
This genus seems to be somewhat intermediate between Os-
bornellus and Omanana, but in general appearance it resembles
the latter more closely.
Premanus hebatus DeLong, new species
In general form and appearance resembling Omanana but
with distinct vertex, color pattern and genitalia. Length 6-7 mm.
Vertex produced, rather broadly rounded, more than twice as
wide between eyes at base as median length.
Color pale brown with dark brown markings. The ocelli are
white between which in the female is a pale broken line. In the
male there are only faint traces of this line. Just above ocelli is
a black transverse band interrupted by a pale depressed transverse
line. On basal half of vertex there is a round black spot next
either eye and a broad longitudinal spot either side of middle. Pro-
notum irrorate with dark brown. Scutellum pale brown with a white
dash surrounded with dark brown in each basal angle, the apex,
midway on each side between these, and one either side of middle
on anterior margin equidistant from the spots in basal angle and
from each other. Elytra smoky, veins dark brown and with a few
white areolar spots. Spots along costa and apical margin dark
brown. Face black with traces of pale arcs and a white band
below ocelli in female.
Genitalia: Female last ventral segment gradually produced
from each basal angle to form a rather broad, produced medium
tooth at center which is slightly notched at apex. Male plates long
JULY, 1944]
Belong— PREMAN us
99
PLATE I
Premanus hebatus DeLong
Fig. 1. Dorsal view of head and pronotum. 2. Lateral view of
head and pronotum. 3. Ventral view of male genitalia. 4. Lateral
view of male genitalia.
100
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX NO. 3
with apical half slender and tapered. Style long, apical fifth
slender, finger-like, curved outwardly with pointed apex. Aedeagus
long, slender, with apex slightly bifid. In lateral view the aedea-
gus form an S-curve.
Holotype male and allotype female collected at Tamaz-
UNCHALE, S. L. P., November 16, 1938, by J. S. Caldwell.
UNBIDDEN HOUSE GUESTS
Unbidden House Guests. By Hugo Hartnack. Hartnack Publish-
ing Co., Tacoma, Wash. 226+62+112+160 pp., illus., 1943.
Price $12.00, plus 20c postage.
This is Volume I of a comprehensive two- volume handbook on
household pests. It is written in four parts, the first of which
considers the more general features of the biology and ecology of
household pests and their control. Part two takes up plants, both
as pests themselves like the wood-rotting fungi, and in their rela-
tion to animal pests. Parts three and four treat respectively of
the invertebrates and the vertebrates, but exclusive of the insects,
which are reserved for the second volume. Insects, however, loom
very large in this first volume in connection with the introductory
section on general principles and in relation with household plants
and vertebrates; moreover, 92 pages in the section on invertebrates
are devoted to crustaceans, myriapods, and arachnids, which come
under the perview of the practical if not of the theoretical ento-
mologist !
Dr. Hartnack’s ideal is that of a unified science of the plant and
animal ecology of the household. He insists that one’s first step
should be an understanding of the habitat relationships involved,
and that only then is one in a position to effect control measures
that are more than palliative and temporary in effect.
Among some of the more unusual topics treated in the book
may be mentioned sections on the origin of housepests, the climate
of the house, the identification of pests by means of the traces
they leave behind them of themselves and their activities, the
decay of cadavers, and keratin or hair -eating pests. In addition
there are, of course, the extensive materials on life-habits and
methods of prevention and control that one expects such a book
to provide. Like its predecessor, “202 Common Household Pests,”
the present work is amply illustrated and indexed and printed on
the best enameled paper. This attractive book should be available
to all who are interested in or have occasion to advise others on
problems related to the flora and fauna of the household. May the
reception accorded it warrant its author in completing his second
volume! — Melville H. Hatch.
JULY, 1944]
HATCH— STAPHLINIDAE
101
STUDIES ON NORTHWESTERN STAPHYLINIDAE I:
ANTHOBIUM STEPH.
BY. MELVILLE H. HATCH
University of Washington, Seattle
This is the ftrst of a series of papers in which I hope to make
known some of the apparently undescribed species of Staphyli-
nidae represented in my collection mostly from the State of
Washington.
Anthobium (Eusphalerum) californicum Fauvel
Anthohinm {Eusphalerum) californicum Fauvel, Bull. Soc. Linn.
Normand (3) II, 1878, p. 198, 199.
Blewett, Cle Elum, Swank Creek (Kittitas County). Also a
specimen from Golden, Oregon.
Anthobium (Eusphalerum) orientale Bernhauer
Anthobium {Eusphale7'um) orientale Bernhauer, Pomona Coll.
Jour. Ent. IV, 1912, p. 678.
I identify with this species originally described from New
Hampshire and Massachusetts an extensive series from Western
Washington. I suspect it is the species identified as horni Fauv.
by Blatchley, Col. Ind. 1910, p. 481, being distinguished from the
true horni by its finely punctate pronotum.
Anthobium (Eusphalerum) farrarae Hatch, new species^
Color testaceous, except for the eyes and the four basal seg-
ments of the abdomen in the male which are black; head, prono-
tum, and elytra coarsely, evidently, relatively densely punctate,
the punctures separated by about their own diameter or less, those
on the pronotum somewhat the densest and most clear cut; head
and pronotum alutaceous; pronotum about two-thirds as long as
wide, the apex about three-fourths as long as the base, the front
angles broadly, the hind angles more narrowly rounded, the sides
explanate, more strongly so behind, the side margins arcuate or
just appreciably oblique before the hind angles, the median line
^ l propose to name some of the new species in this series of papers after
members of the Scarabs Society, an informal semi-social organization composed of
some of my former students and others interested in beetles. Since its founding
in June, 1937, meetings have been held at the rate of ten or eleven per year.
The members are: Elizabeth Farrar Kinney, Barbara Gray Bruhns, Daniel E.
Bonnell, Gertrude Minsk, Robert Y. Pratt, Joseph Bruzas, Jr., Clifford J. Burner,
Frank M. Beer, Frances Bjorkman, Helen Gellerman Houk, Ervin F. Dailey,
Gordon K. Patterson, Robert W. Rogers, Mary Porter Freeman, Warren Ewing,
Kenneth M. Fender, Harriet Exline Frizzell.
102
THE PAN-PACIFIC ENTOMOLOGIST [vOL. XXNO. 3
not grooved; elytra about 2 V 3 times as long as the pronotum, cov-
ering the three or four basal abdominal segments, the apices in
the male arcuo-truncate with the sutural angles narrowly and rec-
tangularly rounded, in the female feebly obliquely truncate with
the sutural angles more broadly rounded; the fifth and sixth ven-
tral segments of the abdomen not sexually modified; hind tarsi
with the first two segments longer than the third and fourth, the
first four longer than the fifth; length 2. 7-2. 9 mm.
Type and two puratype males: Fall City, Wash. May 29,
1932, M. H. Hatch. Three additional paratype males from
Seattle and Bothell, Wash. Allotype and one paratype fe-
males: Index, Wash. May 25, 1930, M. H. Hatch. Three addi-
tional paratype females from Seattle and “Stillaguamish,”
near Harrington, Wash.
From allied species, farrarae is distinguished by its testaceous
color, the first four abdominal segments of the male black, its
densely punctate, alutaceous, ungrooved pronotum with arcuate
sides, its unmodified sixth male abdominal segment, and its
feebly dimorphic elytral apices. It is somewhat related to pothos
Mann., which is the commonest species in western Washington, in
which the pronotum is less densely punctate and grooved along
the meridian line, the male with the first four and one half
abodminal segments black, the fifth emarginate along its poste-
rior margin, and the elytral apices scarcely at all sexually
dimorphic.
Anthobium (Eusphalerum) grayae Hatch, new species
Color testaceous, the median portions of the head and pro-
notum and the outer portions of the antennae dusky, the meso- and
metathorax and abdomen piceous black, the latter slightly paler
at the extreme apex in the male; head and pronotum punctate
and alutaceous, the latter densely and strongly so, the elytral
punctures larger and dense but not so clear cut; pronotum about
three-fourths as long as wide, the apex about four -fifths as wide
as the base, the front angles broadly rounded, the side margins
broadly rounded in front, oblique or subsinuate in front of the
narrowly rounded hind angles, the side feebly explanate toward
the hind angles, the middle line more or less evidently grooved;
elytra about two and one-half times as long as the pronotum, cov-
ering four or more abdominal segments; the elytral apices in the
male arcuately truncate, the outer angles broadly, the inner angles
less broadly arcuate, in the female obliquely truncate, the sutural
angles slightly divergent and narrowly arcuate, the apical margin
just laterad to the sutural angles feebly sinuate; male with the
sixth ventral abdominal segment emarginate along the posterior
margin, the seventh ventral consisting of a broad median lobe and
JULY, 1944]
HATCH— STAPHLINIDAE
103
a narrower longer lateral lobe on either side; hind tarsi with the
first four segments subequally short, together shorter than the
fifth; length 2.1-2.25 mm.
Type male, allotype female, and 48 paratypes; Seattle,
Wash., lV-30-1930, M. H. Hatch. 30 additional paratypes:
Seattle, Duval, Evans Cr. (King Co.), Longmire (Mt. Rain-
ier), North Bend (Maloney’s Grove), all in western Washing-
ton. One paratype: Otis, Oregon.
This species is distinguished from other American species of
the subgenus Eusphalerum by the slightly sinuate apices of the
female elytra. From farrarae and pothos it is furthermore dis-
tinguished by its dusky head and pronotum and black female
abdomen.
Anthobium (Anthobium) bonnelli Hatch, new species
Piceous black, the elytra and appendages testaceous; head and
pronotum finely distinctly punctate, the pronotum more evidently
alutaceous than the head, the elytra more coarsely but less dis-
tinctly punctate than the head and pronotum; pronotum about
seven-tenths as long as wide, the apex seven-tenths as wide as
base, the sides evenly arcuate, the fore and hind angles broadly
rounded, the side margin feebly explanate in front of the hind
angles; elytra (male) two and one-half or more times as long as
the pronotum, covering the three or four basal abdominal seg-
ments, the apices broadly arcuate into the more narrowly arcuate
sutural angles; elytra (female) more than three times as long as
the pronotum, longer than the abdomen, the apices broadly curving
into the acutely produced slightly divergent sutural angles, the
apical margin just la ter ad to the sutural angle strongly sinuate;
male with fifth abdominal segment unmodified, the sixth abruptly
and semi-circularly thickened at the middle, the semi-circular area
being separated from the posterior edge by a narrow membranous
margin; hind tarsi with the first four segments subequally short,
together shorter than the fifth; length 1.8-2.25 mm.
Type male, allotype female, and two paratypes’, Sultan,
Wash. April 14, 1931 M. H. Hatch. Two additional paratypes
same locality: May 10, 1933, May 13, 1931.
By its dimorphic elytra and the structure of the sixth abdom-
inal segment in the male, this species is related to tihiale Csy.
from Arizona, from which it differs by its more strongly arcuate
pronotal side margins (subparallel, very feebly arcuate in tihi-
ale) and longer male elytra (about twice as long as the pro-
notum in tihiale, exposing four abdominal segments, as opposed
to only two or three in honnelli ) .
104
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX NO. 3
Anthobium (Anthobium) minskae Hatch, new species
Testaceous, the male with the first four segments of the ab-
domen black; head and pronotum alutaceous, moderately finely
punctate, the pronotum more conspicuously so, elytra with coarser
but less clear-cut punctures; pronotum about five-eighths as long
as wide, the apex about two-thirds as wide as the base, the median
line not grooved, the front angles broadly rounded, the side mar-
gins arcuate in front, behind oblique or broadly subsinuate before
the more or less obtuse hind angles, the side margins somewhat
explanate, more broadly so behind, elytra three (male) or four
(female) times as long as the pronotum, the apices (male) broadly
rounded into the narrowly rectangularly rounded sutural angles,
exposing the apex of the abdomen, or (female) obliquely produced
into the narrowly acutely rounded sutural angles, extending beyond
the abdomen: male with the posterior margin of the fifth ventral
abdominal broadly triangularly incised, the apex of the incision ex-
tending as a parallel-sided fissure about four times as long as deep,
the length of the fissure being slightly greater than the depth of the
triangular incision, the sixth ventral segment broadly triangularly
incised alongi its posterior margin, the median lobe of the seventh
segment triangular; male protibiae straight; last tarsal segment
shorter than the four basal segments taken together, that of the
hind tarsus being about three-fifths as long, the first and second
segments elongate; length 2.2-2. 7 mm.
Type male, allotype and one paratype female: Olympic Hot
Springs, Wash. May 31, 1931, M. H. Hatch. Ten additional
paratypes; Cooks, Lake Crescent, Lewis and Clark State
Park, Seattle, and Swauk Creek (Kititas Co.), all in Wash-
ington.
Related to fraternum Casey from northern California by the
fissure of the posterior margin of the fifth ventral abdominal
segment in the male, but distinguished therefrom by the black
color of the four basal segments of the male abdomen, which is
pale in fraternum. Casey makes no mention of the fissure in his
species being itself at the apex of a triangular incision, as in
minskae.
In farrarae and minskae described above and in such previ-
ously described species as convexum Fauv. and fraternum and
punctatum of Casey the structure of the tarsi is such as to raise
the propriety of retaining them in Anthobium. In these species
the first two segments of the tarsus are longer than the next two,
the first four together longer than the fifth. I leave the problem
to future students of the group.
JULY, 1944]
BEER— OREGON ACMAEODERA
105
NOTES ON THE HOSTS, HABITS AND DISTRIBUTION
OF OREGON ACMAEODERA
(Coleoptera, Buprestidae)
BY FRANK M. BEER
Salem, Oregon
This paper is a record of the Acmaeodera of Oregon, mainly
collected by the writer during the past seven years. A majority
of the species have been taken in the Rogue River Valley where
they inhabit their usual hosts which encroach from the Sierra
Nevada Range and adjoining foothills. The earlier records for
the state are woefully incomplete, the species in reality occurring
far more commonly and abundantly than previous reports would
indicate. Dr. Chamberlin’s^ paper in 1929 was the last to report
this genus and list new material actually taken by an Oregon col-
lector. In his catalogue, Chamberlin^ lists four species found in
Oregon, and an additional one, flavosticta, as doubtful, while
Barr^ lists five in his recent excellent resume.
After intensively collecting through the state the number of
species has been increased to fourteen and two varieties, and the
range of several materially extended. Since a large part of the
collecting was done during the winter and early spring by cutting
the adults from their pupal cells and a few having been reared
from larvae, exact host plants can be given for the majority of
the species. Though host plants may not in general be consid-
ered to have a great deal of significance, however, after much
careful observation, the species have been consistently found in-
habiting the same host plant or plants.
Acmaeodera acuta LeConte
This species is by far the most common collected in south-
western Oregon. It is generally found feeding on the flowers of
Eriophyllum lanatum (Pursh) Forbes, a yellow composite oc-
curring in the foothills of the Rogue River Valley and southward
across the Siskiyou summit into California. The insect also oc-
’ Chamberlin, W. J., 1929, Remarks of the Buprestidae of the North Pacific
Coast Region with descriptions of new species, Pan-Pac. Ent., 5:111.
- Chamberlin, W. J., 1926, Catalogue of Buprestidae of North America.
® Barr, W. F., 1941, Distribution of Acmeodera, Pan-Pac. Ent., 17 :62-69.
106
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX NO. 3
curs in the Willamette Valley as occasional specimens have been
taken on the blossoms of wild rose {Rosa pisocarpa Gary) . The
larvae mine the smaller dead branches and stems of the Garry
oak {Quercus garryana Dough) from which adult specimens
have been cut.
AcMAEODERA ANGELICA FaLL
Numerous specimens of this little Buprestid have been taken
in southwestern Oregon, most of them beaten from the foliage or
dug from dead wood scars of buckbrush {Ceanothus cuneatus
Nutt.), though a few have been cut from dead branches of Cali-
fornia black oak {Quercus californica [Torr.] Coop.)
The variety nexa Fall occurs through the same region and has
been taken from buckbrush. Recently a specimen was reared
from mountan mahogany {Cercocarpus betuloides Hook.).
Acmaeodera bishopiana Fall
The only record we have of this species is of three specimens
taken by Mr. M. C. Lane at Adrain, Oregon, on May 28, and re-
ported by Dr. W. J. Chamberlin It apparently occurs through-
out the Great Basin region, as the original specimens came from
Owen’s Valley, California, and the writer has specimens reared
from the Walker Lake region in Nevada. The host in its southern
range is the wingscale {Atriplex canescens [Pursh] Nutt.). This
plant extends northward into Oregon.
Acmaeodera connexa Fall
This species occurs commonly through the southwestern part
of the state wherever the host plant is found. Many adult speci-
mens and numerous larvae have been dug from the scars and
dead trunks of the Garry oak {Quercus garryana Dough). Dur-
ing June, it can be taken on one of the willow herbs {Epilohium
sp.) and occasionally on the composite, Eriophyllum lanatum
(Pursh) Forbes.
\
Acmaeodera fenyesi Fall
This uncommon form has not previously been recorded from
the state and to my knowledge the host plant has not been re-
ported. The species is apparently quite rare in our range as
^ Chamberlin, W. J., 1929, Pan-Pac. Ent., 5 :111.
JULY, 1944]
BEER— OREGON ACMAEODERA
107
literally days of cutting thruogh manzanita {Arctostaphylos
viscida Parry) has produced only seven specimens. These are
somewhat larger than specimens from southern California, and
one female is noticeably more maculate. All specimens have been
taken from the foothills of the Rogue River Valley.
Acmaeodera gemina Horn
Though this species is never taken in abundance, it is occa-
sionally found feeding on several varieties of flowers or is swept
from grass. Careful search over several years has not revealed
its host. The species has been taken from the floor of the Rogue
River Valley to the high mountains in the vicinity of Crater Lake.
The variety nehulosa Horn is apparently rare in this region,
as only one specimen has been taken, that from near Crater Lake.
Acmaeodera hepburni LeConte
This is another common species which has a wide range of
distribution. It is found throughout the foothills of southwestern
Oregon' where its larvae infest the Garry oak [Quercus garryana
Dough). It is also taken abundantly on the flowers of the com-
posite Eriophyllum lanatum (Pursh) Forbes, along with acuta,
connexa and labyrinthica.
Acmaeodera labyrinthica Fall
Many specimens of this beautiful species were collected in the
Siskiyou Mountains of southern Oregon, the specimens being
most numerous on the upper Applegate in the vicinity of Copper,
Jackson County, near the Oregon-California line. Two specimens
were dug from dead scar tissue of living Garry oaks {Quercus
garryana Dough). In early June many specimens were taken
feeding on the blossoms of Eriophyllum lanataum (Pursh)
Forbes. This species is considered as distinct by some writers
and as a variety of connexa or acuta by others. However, in any
series it can be easily distinguished by its breadth and bronze
lustre. In the last respect it more closely resembles hepburni,
from which it differs by the shape of the pronotum and the
emargination of the prosternum.
A rather interesting observation was made upon the group
acuta, connexa, hepburni and labyrinthica which would seem
worthy of mention. Numerous specimens of labyrinthica with
lesser numbers of acuta, connexa and hepburni were observed on
108
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX NO. 3
three occasions while they were feeding on the previously men-
tioned yellow composites. During this time numerous pairs were
seen mating, hut at no time did specimens outside the species
succeed in copulating, though occasional attempts were made.
Although the four species are distinctive enough to he separated
on external features, this hit of information lends more proof to
the helief that there are four distinct species.
AcMAEODERA MARIPOSA HoRN
This distinctive species is not abundant, hut a couple of dozen
specimens have been taken. About one-half of them have been dug
from their pupal cells in mountain mahogany [Cercocarpus beta-
loides Hook.) where they work through dead trunks or through
dead patches in living trees. During the spring of 1940, several
specimens were swept from the foliage of antelope brush {Put-
schia tridentata [Pursh.] D. C.) on Roxy Anne Butte, five miles
east of Medford, Jackson County, Oregon. All specimens have
been taken on or near the floor of the Rogue River Valley, none
apparently occurring above 2500 feet elevation.
Acmaeodera plagiaticauda Horn
Because this species is seldom obtained while sweeping it is
considered rare. After learning the habits of the larvae, and col-
lecting accordingly, recently transformed adults are quite easily
obtained by cutting through the dead patches of manzanita
[Arctostaphylos viscida Parry) . It ranges over most of south-
western Oregon, extending as far north as Azalea, Douglas
County.
Acmaeodera purschiae Fisher
This species was first found in the state near Medford,
Jackson County, where two specimens were cut from antelope
brush {Purschia tridentata Pursh.). On June 12, 1940, about
thirty specimens were taken while sweeping the host plants on
Roxy Anne Butte, Jackson County. Previously, five specimens
had been cut from the host plant at the base of Mount Shasta,
near Weed, Siskiyou County, California. The range of this spe-
cies is thus extended northward by several hundred miles and
into a third state.
JULY, 1944]
BEER— OREGON ACMAEODERA
109
AcMAEODERA SINUATA VAR. SEXNOTATA VaN DyKE
Numerous specimens of this beautiful variety have been col-
lected, all having been dug or swept from buckbrush {Ceanothus
cuneatus Nutt. ) . They range over most of southwestern Oregon,
especially through the Siskiyou Mountains. No examples of the
species sinuata Van Dyke have as yet been taken.
AcMAEODERA VANDYKEI FalL
This widely distributed species has seldom been found in its
pupal cell. It is not common but is found throughout the por-
tion of the state where the deer brush {Ceanothus integerrimus
H. & A.) occurs. One specimen was also taken from sticky laurel
{Ceanothus velutinus Dough), which must also be listed as a
host plant. Specimens have been taken in the Wallowa and Blue
Mountains of northeastern Oregon, Hood River County, and
south along the eastern slope of the Cascade Range, and across
into the Rogue River Valley and the adjoining foothills.
AcMAEODERA VARIGATA LeCoNTE
This species has the widest distribution in the state. Speci-
mens have been taken from the following Oregon counties:
Douglas, Grant, Hood River, Jackson, Josephine, Klamath, Lake
and Wheeler. The specimens have been cut from two species of
mountain mahogany {Cerococarpus betuloides Hook, and C.
ledifolius Nutt.) and from service berry {Amalanchier sp.) and
are generally found above 3000 feet in elevation. Two varia-
tions are noticeable, the specimens cut from service berry are
smaller and the specimens taken near Glendale, Douglas County,
are more abundantly maculate.
I am indebted to Mr. James H. Baker of Baker, Oregon, for
permission to include a series of fourteen specimens of A. vari-
egata Lee. from his collection. Nine of these were taken near
Baker, Baker County, the other five were collected at Hiatt Lake,
Jackson County. His series, which is typical, thus extends the
species into the northeastern comer of the state.
110
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XXNO. 3
A NEW DOBSONFLY(MEGALOPTERA)FROM CALIFORNIA
BY EDWIN C. VAN DYKE
California Academy of Sciences, San Francisco, California
In 1908, Banks published a paper on the classification of
the Corydalinae, in which he defined the various genera of the
group, at the same time establishing two new genera at the ex-
pense of older genera. One of these was Neohermes which was
to include the following previously described species: fHicornis^
Banks, the genotype, calif ornicus Walker, angusticollis Hagen,
cinerascens Blanchard, and humeralis Banks,
The genus Neohermes is defined by Banks as having the anal
vein of forewings with two forks, the anal cells subequal in
length, the lower with two veins, the upper with one, and no cross
vein between the first branch of the anal vein and the vein from
anal cell, the costal cells about as long as broad, the antennae as
near to the ocelli as to the eyes, the branches of the median vein
simple, and but three cross veins from radial sector to radius.
Neohermes nigrinus Van Dyke, new species
Of moderate size and entirely black except the anterior part of
head which is somewhat fulvous and a few cross veins on the outer
portion of both wings which are white. Head widest across eyes
which are of moderate size and protuberant, convergent in front
and gradually narrowed behind, the ocelli conspicuous, and with
two elongate callosities centrally located behind; antennae fili-
form, with segments slightly longer than broad and somewhat
moniliform, and almost three-fourths length of forewing. Pro-
thorax longer than broad, with sides parallel and disk opaque
except for sides and medial area behind, which are smooth and
shining. Wings about four times as long as broad, the hind pair
with sides more parallel as usual. The wing venation agrees with
the genus and such typical species as calif ornicus Walker.
Holotjpe (No. 5430, Mus. Calif. Acad. Sci., Ent.) and one
paratype. Holotype, expanse of wings, spread, 5 cm. 2 mm.,
length of body 2 cm. ; paratype, expanse of wings 4 cm. 2 mm.,
length of body 1 cm. 5 mm. Both specimens were collected by
me at Richardson’s Spring, Butte County, California, the
holotype. May 12, 1944, and the paratype May 18, 1944.
The black color, moderate size, a rather small head as com-
pared with californicus, and simple filiform antennae should
readily separate this species from any of the others in the genus.
’ According to Weele, Coll. ZooL, Selys Longchamps, 1910, filicornis Banks as
also angusticollis Hagen, concolor Davis, and minutus Davis are synonyms of cali-
fornicus Walker, therefore the last would be the genotype.
JULY, 1944]
LINSLEY AND USINGER— ARADIDAE
111
FURTHER NOTES ON THE HABITS OF SOME FLAT BUGS
WITH A DESCRIPTION OF THE MALE OF ARADUS
PATIBULUS VAN DUZEE
(Hemiptera, Aradidae)
BY E. G. LINSLEY AND R. L. USINGER
University of California
The following observations on aradids were made between
May 10th and June 15th, 1942, in the Sierra Nevada Mountains
of Madera and Mariposa counties in the vicinity of the south
entrance of Yosemite National Park, California. As in our
previous paper,^ biological observations are by E. G. Linsley,
taxonomic notes and identifications by R. L. Usinger.
Aradid spring dispersal flights were observed at Miami Ranger
Station, Madera County. The flights began on May 19th, which
was the third of three successively warmer days after the end of
the spring snow storms and rains. On May 19th, the flight began
at approximately 5 P.M. P.S.T. and was over by 6:15 P.M. A
sample of 102 specimens was taken. This included 6 species, of
which 51 per cent of the examples were Aradus behrensi Bergr.
On May 20th a second flight occurred. The day was warm but
overcast in the late afternoon. The flight began at approximately
4 P.M., was over by 5:30 when the temperature began to fall.
The same sampling method yielded only 39 examples of two
species, with A. behrensi again predominant. On May 21st, a
third flight occurred, this time on a warm but wholly overcast
day. The flight began at about 4 p.m., lasted until approxi-
mately 6:00. Forty-three examples were taken representing six
species, of which more than 50 per cent were again behrensi.
On this date three species appeared which had not been repre-
sented in previous samples. This was the last flight observed. On
both May 22nd, and 23rd, one example of behrensi was taken
but no further examples were captured in flight between May
23rd and June 15th (termination of observation) .
Associated with the aradids in their dispersal flights were
numerous species of Coleoptera (fig. 1). Although the samples
^ Linsley, E. G. and R. L. Usinger, 1942. Notes on some flat bugs from the
vicinity of Mt. Lassen, California. Pan-Pac. Ent., 18 :83-86.
112
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX NO. 3
obtained are too small to be of significance as far as most of the
individual species are concerned, the totals clearly indicate that
the flights of these insects as a group were correlated with those
of the aradids. Of further significance would appear to be the
fact that the majority of these species are bark inhabitants, as
are aradids, the remainder mostly wood borers. This suggests
that the same factors which initiate aradid dispersal also influ-
ence emergence of other insects which occupy the same habitat.
ARADINAE
, Aradus ampliatus Uhler
This species was not encountered until the third day of dis-
persal flights. This is of interest in view of the fact that in the
Mt. Lassen area adults were taken in flight two days prior to the
flights of other species.
Aradus patibulus Van Duzee
Taken in flight only on May 19th. Adults later found under
bark of western yellow pine at Fish Camp, Mariposa County.
Not previously observed in dispersal flight.
Male. Antennae and pronotum as in the female (Usinger,
Ann. Ent. Soc. Amer., 29:515, plate 1, fig. 4A, 1936) but size
smaller, 7 mm. long, 2^ mm. wide across pronotum, and 3%
mm. wide across the abdomen. Male genital capsule similar to
that of ampliatus Uhler (Usinger, Loc. cit., fig. 4) but narrower
than long, 27 : :29; more narowed basally with shallower, rounded
basal sinuation ; and apical lobes a little more strongly produced,
extending beyond general contour of abdomen. Other characters
as in the female.
Neallotype, male, Miami Ranger Station, Mariposa County,
California, May 20, 1942 (E. G. Linsley) .
Aradus depictus Van Duzee
In flight on May 21st. This species was not encountered in
the Mt. Lassen area.
Aradus fuscomaculatus Stal
In flight only on May 19th. Also found under bark of western
yellow pine at Sugar Pine, Madera County. Not previously ob-
served in dispersal flight.
JULY, 1944]
LINSLEY AND USINGER— ARADIDAE
113
Aradus behrensi Bergroth
Tliis species dominated the dispersal flights on May 19-21,
but was most numerous (52 examples) on May 19th. On May
30th, at Bass Lake, Madera County, 67 adult specimens were
found under a six-inch square of bark of Quercus kelloggii.
Tsbl* I
C0120PTERA ASSOCIATED IB AEADID DISFSBSAL FUOETS - WTAin BABOER StATIOB - HAT 1942
Ifay 18
Hay 19
Bay 20
Bay 21
Bay 22
Iby 28
lAtrlnMOB plotim Eaur. (Btaphyllnldae)
a
12
2
a
0
0
Alonothroa oua ralldna Lao. (Throaoldae)
0
14
1
2
0
0
Throaona sendaz Horn (Throaoldae)
0
6
0
2
0
1
Eronyxa expansa Van D:^e (Oatoaldae)
0
5
0
0
0
0
Calil^a Boahra Thnnb.erg (Oatomidae)
0
2
1
0
0
0
Tenebroldaa slnnatua LeConte (Oatomldae)
0
1
5
2
0
0
IiOblopa oblonga Paraona (Hltldulldae)
0
1
1
4
0
1
Pertbalyora murrayi Horn (Hitidnlidae)
0
3
5
0
0
0
Oliaohroohllua ojrilndriona (LaConte) (Bltldolldae)
0
1
2
0
0
0
Laemophloena ap. (Cnoujldae)
0
6
19
7
0
0
lAsmophloeua ap. (Cnou;]idae)
0
1
6
3
0
0
Cryptophagua ap. (Cryptophagldao)
0
5
2
0
0
0
Vyoetophagua oallfomloas Horn (l(yoetophagidae)
0
11
2
0
1
0
laaoonotna oomplex LaConta (Colydildaa)
0
0
9
0
0
0
Anlonlnm longum LaConte (Colydildaa)
0
0
3
1
0
0
Phynaphora oallfomloa Horn (Ihroetaeldae)
1
8
5
6
0
0
PtlnuB Intomjptna LaConta (Ptlnldae)
0
2
0
2
0
0
Emoblua ponotulatna (LaConte) (Anoblldae)
0
2
10
4
0
0
Stephanopaohya paolflona Casey (Bostrlohidaa)
0
7
6
3
0
0
Spondylls uplfonsla Idannerhalm (Cerasdiyoldae)
2
6
11
3
1
1
Aaemum mokelnmne (Casey) ( Cerambyo Idae )
0
3
1
0
0
0
Atlmla dorsalis LaConta (Ceranbyoidae)
0
4
14
9
0
0
Gylptosoells aequolaa Blaladall (Chrysomalldae)
0
2
11
0
1
0
Eblnomaoar boablfrons LeConte (C roullonldae)
0
1
3
0
0
0
Pieaodes yoaemlte Hopkins (Cnronllonldae)
0
2
6
3
0
1
CosaonuB ponderoaaa Van Dyke (Curoullonldaa)
0
8
11
5
1
0
Cosaonus orenatns Horn (Curoullonldaa)
0
4
7
0
2
0
Dandrootonua valens LeConte (Soolytldae)
2
38
63
16
2
0
Qylurgopa leoontal Snlna (Soolytldae)
0
9
12
2
0
2
^lastea nlgrlnua (Mannerhelm) (Soolytldae)
0
19
34
4
1
2
Hylaatas maoer LeConte (Soolytldae)
14
148
113
78
14
6
Pltyophthorus sp. (Soolytldae)
0
6
2
0
1
0
Ips latldens (LeConte) (Soolytldae)
0
2
4
3
0
0
Gnathotrlohus suloatus (LeConte) (Soolytldae)
0
3
17
2
0
0
TOTAL
19
415
376
160
24
14
where they had apparently congregated for mating. No imma-
ture stages were present.
Aradus apicalis Van Duzee
In flight only on May 21st. Dispersal flights of this species
have not been previously recorded.
114
THE PAN-PACIFIC ENTOMOLOGIST [vOL. XX NO. 3
Aradus debilis Uhler
Small flight on May 19th. This species overwinters as adults
or last instar nymphs. First instar nymphs were found on
Polyporus volvatus fungi on fallen western yellow pine on May
28th, 9 days after the dispersal flight was observed. Adults were
also found in large numbers on the dark upper surface of the
fungus, Pomes penicola, on fallen white fir at Mariposa Big
Trees, June 6th. Because of perfect cryptic coloration they were
very difl&cult to detect and several minutes were required to com-
plete examination of each fungus. The maximum number of
individuals encountered on a single fungus was fourteen.
Aradus blaisdelli Van Duzee
In flight on May 19th. Dispersal flight not previously recorded.
Adults also found under bark of western yellow pine.
Aradus compressus Heidemann
In flight on May 19th, 20th, and 21st. Largest flight on May
19th.
Aradus cinnamqmeus Panzer
Commonly beaten from branches and new foliage of western
yellow pine at Miami Ranger Station and Fish Camp, Mariposa
County, from May 19th to June 10th. A few examples were also
found on Abies concolor at Sugar Pine, Madera County.
MEZIRINAE
Mezira reducta Van Duzee
Found at Miami Ranger Station in colonies of Zootermopsis
in western yellow pine, sugar pine, white fir, and red fir, and in
nests of Xylocopu californica in incense cedar.
Mezira pacific a Usinger
Taken under bark western yellow pine, sugar pine. Digger
pine, white fir, incense cedar, Quercus kelloggii, Alnus rhomhi-
folia, and Populus trichocarpa.
JULY, 1944]
TILDEN AND MANSFIELD— COENONYCHA
115
NOTES ON THREE SPECIES OF THE GENUS COENONYCHA
HORN
( Coleoptera, Scarabaeidae)
BY J. W. TILDEN AND G. S. MANSFIELD
In view of the somewhat limited information available on
species of the genus Coenonycha Horn, it was thought the fol-
lowing notes concerning three recently described species might
prove of interest.
The type series of C. howlesi Cazier was collected about three
miles northeast of Benton, Mono County, California, by J. W.
Tilden and C. W. Bowles. The first specimens taken were at-
tracted to a gas lantern at dusk. The remainder were collected
from the tips of Artemisia tridentata, one person collecting while
the other held the light. Most of them had the forelegs extended,
a habit common to species of this genus. They seemed indifferent
to light that was held on them, though some were attracted to the
light if it remained stationary for a period of time. The vegeta-
tion of this area is predominantly Artemisia tridentata, with an
admixture of other bushy composites. C. howlesi may occur on
other plants, although to date it has not been taken elsewhere.
C. testacea Cazier, described from Cuyama Canyon, Santa
Barbara County, California, was taken in large numbers about
ten miles west of Simmler, San Luis Obispo County, by the
authors on March 20-22, 1940. This species was closely associ-
ated with Eriogonum fasiculatum here as in the type locality. On
each of the three evenings in the Simmler area the first speci-
mens noted came to a gas lantern shortly after sundown. Mat-
ing pairs were observed about dark and commonly later in the
evening.
The insects were taken in numbers from the tips of the plants,
and normally sat with the forelegs extended as in howlesi. They
have well developed flight wings but seem averse to flying. The
number taken at light was very small in proportion to the popu-
lation that was found to exist in the immediate area. The few
that came to light must have flown there, since the collecting
sheet was clear of the ground. Specimens were taken from the
plants until about 12:00 P. M. No doubt they could have been
taken later, although we found the insects less numerous in the
late hours than they were earlier in the evening.
116
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX NO. 3
A third species, C. ampla Cazier, was also found near Simmler,
but on Juniperiis californica, the only tree in the vicinity. The
first was taken by beating about 10:00 P. M., March 21. Exten-
sive beating and inspection yielded several others on the same
night.
On the succeeding night, these were observed carefully. At
first we thought them rare, since so few were found, but one
location was discovered where about 18 were taken. Though
relatively common, they did not approach testacea in numbers,
in this area at least.
Careful approach enabled us to observe these insects without
disturbing them. Each was found at or near the tip of a twig.
This species proved to be negatively phototropic. They would
nearly always drop at the close approach of the light, or would
edge around to the other side of the twig. They were seen to
extend their forelegs as did the other two species. It was noted
also that they dropped if our breath was blown on them. Whether
this was due to objection to the air current or to another factor
we do not know.
At first no feeding was observed. It occurred to us that this
might be due to the light. Such proved to be the case. When
the light was weakened, the beetles became active and fed. When
in moderate to strong light they suspended activity. One of us
obsrved while the other slowly removed the light. The beetle
was seen to crawl deliberately along the limb and begin to feed.
With a slow return of the light, the beetle first ceased to feed,
then stopped all movement, and as the light grew stronger, ex-
tended its legs and dropped to the foliage beneath. It made no
effort to extricate itself from the position in which it landed as
long as the light was present. With the removal of the light,
activity was resumed gradually. Several were observed to behave
in this same manner, and although the quiescent individuals were
conspicuous at all times, they made no effort at concealment.
This habit might easily lead to their dropping before being
taken by beating, and it seems likely that the species is more
common than our collecting results show.
All three were found at the tips of their respective forage
plants with forelegs extended, but in the case of ampla this ap-
peared to be the first stage in a negative response to light, for
with an increase in the intensity of the light they extended the
other pairs of legs as well, and dropped from the trees.
JULY, 1944]
TILDEN— PHOEBIS SENNAE MARCELLINA
117
None of these species was found in daytime, all activities of
mating and feeding being strictly nocturnal, nor were any taken
on any plant other than the one listed for each species.
BREEDING OF PHOEBIS SENNAE MARCELLINA CRAM.
IN SAN JOSE, CALIFORNIA
( Lepidoptera, Pieridae)
In October, 1941, Piazza first noted larvae of this species on
cultivated shrubs of Cassia tomentosa, and later located pupae
and eggs. Reared specimens emerged continuously during the
fall. Piazza and Tilden took a considerable number of all stages
from a shrub of the same species, on December 2, 1941, and
these were reared by Tilden.
From 17 chrysalids removed from the shrub and kept indoors,
2 perfect males, 4 perfect females emerged, while 6 females
emerged with damage from defective chrysalids. Two chrysalids
were parasitized. Three failed to produce adults. Emergence
dates range from December 4 to January 9 (1942). The propor-
tion of females seems unusual, there being 8 females and 4 males,
a ratio of 2:1. Adults reared from specimens collected as larvae
emerged in better condition than those obtained from specimens
that had pupated before being taken, suggesting probable damage
to the chrysalids in removing them from the food plant.
Emerged specimens refused to mate in captivity. Eggs taken
on December 2 did not hatch before the project was abandoned
in June, 1942. All specimens taken as larvae matured and pu-
pated; some still had not emerged in June, indicating a long
pupal stage under some conditions.
Although adults of P. s. marceilina Cram, have been noticed
repeatedly in the San Jose area over a period of several years,
this appears to be the first time that the entire life cycle has been
observed at any locality so far north. Santa Barbara County is
given by Dr. J. A. Comstock as the usual northern limit of the
(breeding?) range.
It is possible that the species may be establishing itself in the
San Jose area, probably due to the cultivation of suitable food
plants, which enable stragglers of this strong-flying butterfly to
form at least temporary colonies north of its normal range. —
J. W. Tilden.
118
THE PAN-PACIFIC ENTOMOLOGIST [vOL. XX NO. 3
A BRIEF NOTE ON THE HABITS OF AMOBIA
(PACHYOPHTHALMUS) FLORIDENSIS
TOWNSEND
(Diptera, Sarcophagidae)
BY E. GORTON LINSLEY
University of California, Berkeley
Amohia {Pachyophthalmus) floridensis Townsend appears to
be a common inhabitant of the nests of Sceliphron servillei
(Sauss.) in California. On numerous occasions the writer has
found pupae of what was apparently this species in the mud
cells of this wasp. In February, 1944, however, in connection
with another study, large numbers of Sceliphron nests from vari-
ous localities in the San Joaquin Valley were examined and a
small percentage from each locality contained pupae of this fly.
Pupae were found in sealed cells under conditions suggesting that
larvae or eggs were deposited during the provisioning period.
The larvae had apparently fed on the Sceliphron larvae as well
as on the stored spiders. Seven adults reared on April 4 from a
cell series taken under a culvert at Chowchilla, Madera County,
on February 15, were sent to H. J. Reinhard, v/ho identified them
as Amohia {Pachyophthalmus) floridensis Townsend. Allen^
summarizes the known habits of Pachyophthalmus as follows:
“The members of the genus, so far as known, are biologically
superimposed on wasps which build mud nests under shelter or in
hollow stems, including members of the families Sphecidae and
Eumenidae. The manner in which the female establishes her
progeny and the subsequent larval development have not been ob-
served. Rau found the puparia of one species lodged in a hollow
stem containing the nests of Odynerus. When associated with the
common “dirt daubers” Pelopaeus and Trypoxylon, which build
their nests indoors, it is not known whether the maggots pupate
within the. mud cells of their hosts or leave them to seek other
shelter for the transitional stage. The members of the genus
Pachyophthalmus are more distinctly aerial in habit than the other
North American genera of Miltogrammini. So far as is known, all
of the activities of adult, larval, and pupal stages take place in a
stratum extending from a foot or more to several feet above the
ground. This is a noteworthy biological specialization in a group
of flies, most of which pass the immature stages underground, and
never rise, as adults, but a few inches above the ground.”
A. {P.) floridensis has been previously recorded from cells of
Sceliphron cementarium (Drury) in Maryland and Mississippi.
^ Allen, H. W., 1926. North American species of two-winged flies belonging to
the tribe Miltogrammini. Proc. U. S. Nat. Mus., 68 (9)1-106, pis. 1-5.
JULY, 1944]
HEMMING— NOMENCLATURE COMMISSION
119
RECENT WORK BY THE INTERNATIONAL COMMISSION
ON ZOOLOGICAL NOMENCLATURE
BY FRANCIS HEMMING
Secretary to the International ComTnisisoyi on
Zoological Nomenclature
The International Commission on Zoological Nomenclature are
now engaged in the publication of decisions taken before the out-
break of war in 1939. These decisions have been embodied in
Opinions 134-183 and Declarations 10-12. Of these, Opinions 134-
155 and the three Declarations have already been published or are
in the press. The remainder are ready for printing and will be
published as soon as funds are available.
These Opinions are of particular interest to entomologists, since
in addition to nine Opinions relating to the interpretation of vari-
ous aspects of the International Code, no less than 38 of these
Opinions are directly concerned with entomological subjects.
The Opinions relating to the interpretation of the Code are:
Opinions 138 (meaning of phrase “definite bibliographic refer-
ence" in Article 25) Opinion 141 (naming of families) ; Opinion
145 (status of names first published in invalid works) ; Opinion
147 (generic names of same origin and meaning as older generic
names) ; Opinion 148 (status relating to names published as emen-
dations of, or substitutes for, older names) ; Opinion 164 (position
as regards types when two or more genera are united) ; Opinion
168 (supplementing Opinion 65 regarding genera based upon erro-
neously determined species) ; Opinion 172 (status of type — desig-
nations of genera in abstracts, etc.) ; and Opinion 183 (interpreta-
tion of Article 8 relating to form in which generic names should
be published) .
Of the Opinions specially concerned with entomological subjects,
six deal with the status of particular works or with the dates of
such works: Opinion 136 (Latreille, 1810) affects all Orders of
insects; Opinion 135 (“Erlangen List," 1801) is of special interest
to hymenopterists ; Opinions 134 (Freyer, Neue Beitrdge) , 138
Hubner, Sarmnl. exot. Schmett., 1807 and Fabricius Mag. Insek-
tenk. (Illiger) 1807) and 150 (Hubner, Verz.hek. Schmett) are
concerned with Lepidoptera and Opinion 152 (Meigen, 1800) with
Diptera. Opinions 140 and 143 deal with certain family names in
insects.
The remaining 30 Opinions deal with particular generic names
in various Orders of insects. These Opinions either fix the types
of these genera or add the names to the Official List of Generic
Names in Zoology, or do both. Three of these Opinions are con-
cerned with Orthoptera; 13 with Hymenoptera and 14 with Lepi-
doptera.
Each Opinion is published separately but Opinions are consecu-
120
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XXNO. 3
lively paged to facilitate the publication of an index on the com-
pletion of the volume concerned.
The International Commission are most anxious to secure that
O'pinio'ns are published as rapidly as possible but they are greatly
hampered by lack of funds. The Commission therefore appeal to
scientific institutions and individual scientific workers for dona-
tions to a special fund to be used for the issue of publications.
Full particulars of this appeal are given in Part 2 of the Com-
mission’s official organ, the Bulletin of Zoological Nomenclature,
published in 1943.
Contributions, however small, will be warmly welcomed and will
be acknowledged in the Bulletin. Bankers’ drafts, checks, and
money orders should be made payable to the International Com-
mission on Zoological Nomenclature and sent to the Commission
at their publications office, 41 Queen’s Gate, London, S.W.7. All
orders for the Commission’s publications should be sent to the
same address. Inquiries relating to the work of the Commission
should be addressed to me at 83 Fellows Road (Garden Flat),
London, N.W,.3.
CLOVER LEAF WEEVIL IN CALIFORNIA
The clover leaf weevil, Hjpera punctata (Fab.), an introduced
species, has spread over much of California and occurs on both
cultivated and uncultivated areas. Although acreage of clover in
this state is very limited, the spring brood is common on Melilo-
tus and other wild clovers. We have records of its presence on
alfalfa in only the following counties; Alameda, Butte, Merced,
Sacramento, San Joaquin, Stanislaus, and Yolo. Its presence can
be expected on alfalfa anywhere it is grown in northern and
central California and in some parts of southern California.
However, the clover leaf weevil has not caused economic damage
to alfalfa in California although it has been reported as an im-
portant pest of clover and alfalfa in other sections of the United
States. On only two occasions have we found it in abundance in
California. Once it was found commonly on Melilotus indica
growing along a roadside on the lower southern slope of Mt.
Diablo, Contra Costa County. More recently (July, 1941) it was
taken in the same area on a very small patch of dry-farmed
young alfalfa. Our climatic conditions in the spring are appar-
ently so favorable for the development of a fungus disease that
destructive numbers are generally checked before serious damage
occurs. Limited acreage of clover may also partially account for
the fact that this insect is not an important pest in California. —
Ray F. Smith and A. E. Michelbacher.
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VOL. XX
October, 1944
No. 4
THE
Pan -Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
CAMRAS, NOTES ON THE NORTH AMERICAN SPECIES OF THE
ZODION FULVIFRONS GROUP 121
DE LONG, FOUR NEW SPECIES OF CLOANTHANUS AND TUMEUS
FROM MEXICO AND BRAZIL 129
MITCHELL, NEW SPECIES AND RECORDS IN MEGACHILE 132
NUNENMACHER, STUDIES AMONG THE COCCINELLIDAE— No. 9 144
POTTS AND SMITH, ABERRANT AGLAIS ANTIOPA 146
VAN DYKE, NEW SPECIES OF NORTH AMERICAN OSTOMIDAE 147
CALDWELL, PINTALIA STAL WITH SPECIAL REFERENCE TO
MEXICO 164
D. B. MACKIE PERPETUAL MEMORIAL LOAN FUND 160
TITLE PAGE, TABLE OF CONTENTS AND INDEX FOR VOLUME XX.
San Francisco, California
1944
THE PAN-PACIFIC ENTOMOLOGIST
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The Pan-Pacific Entomologist
VOL. XX, No. 4 October, 1944
NOTES ON THE NORTH AMERICAN SPECIES OF THE
ZODION FULVIFRONS GROUP
(Diptera, Conopidae)
BY SIDNEY CAMRAS
Chicago, Illinois
In a previous paper (Ent. News, 54:187-191, 1943) the
Oblique-fasciatum Group species has been covered and the
sources of material acknowledged.- As a result of the present
study of the large series of specimens that has been aggregated, a
number of species previously recognized appear to be synonyms.
Some of these may prove to be separable in the future, and the
necessity of depending on descriptions, since most of the types
have not been seen, increases this possibility; but I believe that
the present contribution will aid considerably in a better under-
standing of this difficult group. While the majority of specimens
fail into definable species, individual variation frequently over-
laps two species so that an artificial line of division must be
established. This refers to species occurring in the same region
as well as to intermediates between geographical races.
ZODION FULVIFRONS SaY
Zodion fnlvifrons Say, 1823, Jour. Acad. Nat. Sci. Philadelphia,
3:83; Compl. Writ., 2:74 (Maryland).
Zodion ahdominalis Say, 1823, Jour. Acad. Nat. Sci. Philadelphia,
3:84; Compl. Writ., 2:74 (near Rocky Mountains).
Myopa rubrifrons Desvoidy, 1830, Mem. Sav. etr. Acad. Sci. Paris,
2:247 (Philadelphia).
Zodion flavipennne Bigot, 1887, Ann. Soc. Ent. France (6)7:204
(Mexico) .
Zodion lativentre Graenicher, 1910, Canad. Ent. 42:26 (Wisconsin:
Yellow R., Burnett Cr.).
Zodion sayi Banks, 1916, Ann. Ent. Soc. Amer. 9:194 (Virginia:
Falls Church).
Zodion obscurum Banks, 1916, Ann. Ent. Soc. Amer. 9:194 (Cali-
fornia: Bear Valley, San Bernardino County)
‘I Zodion bilineata Van Duzee, 1927, Proc. Calif. Acad. Sci., 16:586
(Oregon: Wallowa Mountains).
122
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
This species is characterized by the larger size: usually 6-8
mm.; the short, wide, thick female genital plate; and the reddish
abdomen of the male. The female abdomen is usually dark, but
may be reddish on the basal segments. The first posterior cell is
practically always open.
There are two main forms of this species. One has the dorsum
of the thorax covered with bluish pollen and with two dark lines ;
the other has more brownish pollen and five lines on the dorsum
of the thorax {= sayi) . Most individuals however are somewhat
intermediate, and the intergradation is complete. With the de-
crease in thoracic lines and more bluish pollen, the abdomen is
usually more reddish, and the proboscis tends to become longer.
Such individuals are more common in the West. A further ex-
treme are those examples with so much pollen that the dorsal
thoracic lines are indistinct. Obscurum seems to be based on
such specimens.
In the past, it has been considered that the abdomen of the
male of this species may be reddish or entirely dark; but I have
found no contra-indication to referring all specimens with en-
tirely dark abdomens to intermedium, although there is com-
plete intergradation between the two types. There is also com-
plete intergradation between this and a smaller species for which
the name ahitus is used here. Formerly, abdominale was used for
this smaller species, but the name clearly belongs to a species
“rather more than one-fourth of an inch” in length. The speci-
men mentioned by Say under abdominalis as “less than half the
above size” does of course belong to the smaller species. I have
set the dividing line between the two species at 5^2 mm., but the
majority of individuals are well above or below this size.
I have examined the type of lativentre and find it to be a
greasy specimen of the present species. Bilineatum based on an
example with a very long third antennal joint, will, I believe,
prove to be an aberrant individual.
Distribution: Southern Canada, United States, and Mexico.
363 specimens examined.
ZODION INTERMEDIUM BaNKS
Zodion intermedium Banks, 1916, Ann. Ent. Soc. Amer. 9:193
(Pennsylvania: Pocono Lake).
Zodion occidentale Banks, 1916, Ann. Ent. Soc. Amer. 9:194 (Ore-
gon: Montaville).
OCT. 1944]
CAMRAS— ZODION
123
Zodion reclusum Banks, 1916, Ann. Ent. Soc. Amer. 9:194 (Cali-
fornia: Redlands).
Zodion basalis Van Duzee, 1927, Proc. Calif. Acad. Sci., 16:586
(Idaho: Moscow Mountains).
This specieg differs from fulvifrons by the long, narrow, thin
female genital plate, and the entirely dark abdomen of the male.
As in fidvifrons there is similar variation in the thoracic lines
and length of the prosboscis. Basale is based on indistinct thor-
acic lines, and reclusum on a long proboscis. There is a certain
amount of variation in the size and shape of the genital plate,
and intermediates bridge intermedium with occidentale. The
relative length of the genital plate is frequently deceptive, and
when the abdomen is extended may resemble that of perlongum
which is proportionately longer and narrower. The border of
dark striations is narrow in intermedium and perlongum, but
very wide in fulvifrons.
Distribution: Southern Canada and United States. 162 speci-
mens examined.
Zodion perlongum Coquillett
Zodion perlongum Coquillett, 1902, Canad. Ent., 34:199 (Nerv
Mexico: RioRuidosa).
This species is characterized by its large size: 7-9 mm.; very
long, narrow, thin female genital plate; and the long, narrow
abdomen due to the third and fourth abdominal segments which
are longer than wide.
This species is known with certainty only in the female sex.
The males are probably large individuals with reddish abdomens
that cannot be distinguished from fulvifrons.
Distribution: United States and Northern Mexico. 22 speci-
mens examined.
Zodion abitus Adams
Zodion abitus Adams, 1903, Kans. Univ. Sci. Bull., 2:33 (Kansas).
Zodion bicolor Adams, 1903, Kans. Univ. Sci. Bull., 2:35 (Kansas
Douglas County).
This species is characterized by its small size: usually 4%-5
mm., reddish abdomen of the male, and usually closed first
posterior cell.
The abdomen of the male is usually entirely reddish except
for the first segment, with yellow pollen on the sides and distal
124
THE PAN -PACIFIC ENTOMOLOGIST [vOL. XX, NO. 4
segments. Some specimens have dark markings representing in-
tergradation with the entirely dark male abdomen of cinerei-
ventre. Such individuals and those with an open first posterior
cell are more common in the Western portion of the range of the
species. The first posterior cell varies from narrowly open to
petiolate as long as the anterior cross-vein. The palpi are short,
not clavate, and about as long as the width of the proboscis.
The use of the name abitus instead of abdominale is explained
under fulvifrons. Abitus is based on the female, and bicolor on
the male.
Distribution: Southeastern Canada and Eastern United States,
west to Colorado (Boulder, Buena Vista, Ridgway). 73 speci-
mens examined.
ZoDION CINEREIVENTRE VaN DuZEE
Zodion cinereiventris Van Duzee, 1927, Proc. Calif. Acad. Sci.,
16:585 (California: Huntington Lake, Fresno County).
This species is characterized by its small size: 4^2-5 mm., the
dark abdomen of the male, and the open first posterior cell.
This species is similar to abitus in the female and differs only
in that the first posterior cell is always open. It is probable that
this cell may be closed, showing variation similar to abitus, but
in the East such individuals are referable to abitus and on the
West Coast to triste. Females with rather wide first posterior
cells have been seen from Michigan and New York, but may be
considered extremes of abitus. Typical dark abdomened males
have been examined from as far east as Illinois.
Distribution: Western United States east definitely to Illinois.
34 specimens examined.
Zodion triste Bigot
Zodion triste Bigot, 1887, Ann. Soc. Ent. France (6)7:203 (Cali-
fornia). '
This name is used here for a species in which the female
genital plate is long, moderately wide, and thin in antero-
posterior dimension ; and yellowish narrowly edged with the dark
striations. The first posterior cell is closed or narrowly open.
The length is 4^-5 mm., and it is otherwise similar to cinere-
ventre. The males in which the first posterior cell is open cannot
be distinguished from cinereiventre and would be referred here
only when associated with the female.
OCT. 1944]
CAMRAS— ZODION
125
Distribution: California (San Diego). 12, li^ (1st post, cell
closed), 1^ (1st post, cell open).
ZODION NIGRIFRONS KroBER
Zodion nigrifrons Krober, 1915, Arch. Nat., 81 A., h.4, 97 (Cali-
fornia) .
Zodion hirtipes Van Duzee, 1927, Proc. Calif. Acad. Sci., 16:587
(California: Melrose, Alameda County).
This species differs from the other members of the genus by
the characteristic fourth vein, the last segment of which ends
about halfway toward the wing margin. This character has proved
constant in the twelve specimens examined and was mentioned
in the original descriptions of both nigrifrons and hirtipes. The
species in general has long hairs and is very dark, the front and
antennae being usually almost entirely dark, and the thorax so
dark that the darker dorsal lines may be quite indistinct. The
palpi are short, about l]4x width of the proboscis, and not
clavate. Length, 4]4-5 mm.
Distribution: California (Melrose, Alameda County; Berke-
ley; Fish Ranch, Berkeley Hills). 12 specimens examined.
Zodion americanum Wiedemann
Zodion americana Wiedemann, 1830, Auss. Zw. Ins., 2:242 (Monte-
video) .
Zodion nanellum Loew, 1869, Berl. Ent. Zeitschr., Cent. VIII, 75
(District of Columbia).
Zodion pygmaeum Williston, 1885, Trans. Conn. Acad., 6:381 (Cal-
ifornia) .
"I Zodion occidensis Walker, 1849, List. Dipt. Brit. Mus., 3;676
(Ohio).
This species is characterized by its small size: 3-5 mm., con-
stantly open first posterior cell, and relatively sharply defined
black markings of the thorax and abdomen.
This species differs from cinereiventre and the other small
forms in being covered with finer pollen which is bluish or green-
ish; and in having the dark markings usually more extensive,
more opaque, and more sharply defined.
The variation in dark markings is very great and completely
bridges the difference between nanellum and pygmeum. Three
males from New Mexico (Highrolls, Alamogordo) completely
lack the thoracic lines and spots, and have less black on the
abdomen than usual. These may repreesnt another species. Nor-
126
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
mally patterned specimens heve been examined from Highrolls.
Krober has compared the type of americanum with nanellum
and considers them the same species. I havie seen examples of this
species from a number of localities in South and Central America.
Distribution: Southern Canada, United States, Central Amer-
ica, and South America. 226 specimens examined.
ZODION PALP ALE RoBERTSON
Zodion palpalis Robertson, 1901, Canad. Ent., 33:284 (Illinois:
Carlinville) .
This species is characterized by the black, long, clavate palpi;
which are 2V^-3x the width of the proboscis at the level of the
palpi. The first posterior cell usually has a long petiole, but it
may be narrowly open. The labellae of the proboscis are quite
long, about one-fourth the total length of the proboscis. They
are much shorter in the previous species but apparently are long
also in lovetti and parvum. When the labellae are together it
may be difficult to decide their length. The abdomen of the male
has the second and third segments yellow, more reddish on the
distal part of the third; and is relatively long and slender. The
legs are rather slender and entirely dark. Length, 5 mm. ; 1 $
4 mm.
Distribution: Illinois (Carlinville) and North Carolina
(Raleigh). 19 specimens examined, including the type.
Zodion scapulare Adams
Zodion scapularis Adams, 1903, Kans, Univ. Sci. Bull., 2:34 (Ari-
zona) .
This species has not been seen by the writer, but has been in-
corporated into the key. It represents a species between palpale
and parvum, having the abdomen of the former and the palpi
of the latter. Very few specimens are known of this group of
species and additional material may show that some of these
represent individual variation.
Distribution: Arizona.
Zodion parvum Adams
Zodion parvis {parvus in key) Adams, 1903, Kans. Univ. Sci. Bull.,
2:34 (Arizona).
This species is characterized by its small size: 3% mm.; long
petiole of the first posterior cell: 3x anterior cross- vein; short,
OCT. 1944]
CAMRAS— ZODION
127
pale, non-clavate palpi, lx width of proboscis; and dark ab-
domen of the male. The legs are slender and dark, and the label-
lae about one-third the length of the entire proboscis.
Distribution: Arizona and Colorado (no locality). One speci-
men examined.
ZODION LOVETTI VaN DuZEE
Zodion lovetti Van Duzee, 1934, Ann. Ent. Soc. Amer., 27:323
(Colorado: Fort Collins).
This species is characterized by its long (2x width of pro-
boscis), but not clavate palpi; entirely dark abdomen of the
male; and small size: 4 mm. The first posterior cell is closed in
the wing margin to petiolate as long as anterior cross-vein. The
legs are slender and entirely dark.
This species represents another link between pal pale and par-
vum, having the long, though not clavate palpi of the former,
and the dark abdomen of the latter.
Distribution: Colorado (Fort Collins, Tennessee Pass). One
specimen examined.
The following species, each based on a single female, are not
known to the author, and have not been included in the key:
Zodion albifacies Van Duzee, 1927, Proc. Calif. Acad. Sci., 16:588
(Arizona: Ba,boquivari Mountains).
Zodion angusticornis Van Duzee, 1927, Proc. Calif. Acad. Sci.,
16:589 (California: Los Banos).
Zodion bimacula Curran, 1933, Amer. Mus. Nov., 673:7 (Ontario:
Timagami) .
Key to the North American Species of Zodion
1. Thorax marked with black spots pictulum
- Thorax marked with stripes or unmarked 2
2. Thorax dark with two relatively wide light stripes 3
- Thorax light with relatively narrow dark stripes or un-
marked 5
3. Abdomen with golden yellow pollen on distal segments
albonotatum
— Abdomen with oblique white or bluish pollinose markings 4
4. Abdomen dark with bluish pollinose markings cyanescens
— Abdomen mostly rufous with white pollinose markings
ob liquefasciatum
5. Abdomen with golden pollen on distal segments; never with
anterior submedian thoracic lines albonotatum
— Abdominal pollen if yellowish, never golden 6
128
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 4
6 .
7.
8 .
9.
10 .
11 .
12 .
13.
14.
15.
16.
Abdomen mostly rufous with oblique white pollinose marks;
anterior submedian thoracic lines absent oblique fas datum
Abdomen usually dark, pollinose pattern if oblique not dis-
tinctly so; frequently with anterior submedian thoracic lines..?
Over 5% mm. long from base of antennae to tip of abdomen.. 8
5% mm. long or less 12
Females 9
Males 11
Genital plate short, wide, and thick fulvifrons
Genital plate long, narrow and thin 10
Abdominal segments 3 and 4 shorter than wide.. ..intermedium
Abdominal segments 3 and 4 longer than wide perlongum
Abdomen with rufous fulvifrons
(Some of the larger and more rufous individuals are probably
males of perlongum.)
Abdomen with no rufous intermedium
Last segment of fourth vein ends about half way toward the
wing margin nigrifrons
Last segment of fourth vein extends to wing margin 13
Thoracic and abdominal markings relatively sharply defined;
first posterior cell always open americanuon
Thoracic and abdominal markings relatively less sharply de-
fined 14
Palpi long, clavate; 1st posterior cell usually petio\ 2 ite..palpale
Palpi usually short, never clavate 15
Palpi 2x width of proboscis, first posterior cell closed or peti-
olate lovetti
Palpi 1-1% times width of proboscis 16
First posterior cell p etiolate more than 2x anterior cross-vein
17
— First posterior cell petiolate less than 2x anterior cross-vein,
or open 18
17.
Size larger, about 5 mm.; legs more
with yellow on 2nd and 3rd segments.
reddish ;
male abdomen
scamdare
Size about 3% mm.; legs dark;
male
abdomen
dar'k.... parvum
18.
Females
19
Males
21
19.
Genital plate long and thin
triste
Genital plate short and thick...
20
20.
First posterior cell open
....cinereiventre
First posterior cell closed
abitus
21.
Abdomen with red
abitus
Abdomen with no reddish
22
22.
First posterior cell closed
triste
First posterior cell open
cinereiventre, triste
OCT. 1944]
DE LONG— CICADELLIDAE
129
FOUR NEW SPECIES OF CLOANTHANUS AND TUMEUS
FROM MEXICO AND BRAZIL
(Homoptera, Cicadellidae)
BY DWIGHT M . DE LONG
Department Zoology and Entomology, Ohio State University
Cloanthanus dilatus De Long, new species
Resembling nitmdus in general form and appearance but with
distinct genitalia. Length, 4-4.5 mm.
Vertex produced and bluntly angled, one-fourth longer at mid-
dle than basal width between the eyes.
Color: Vertex black with a short longitudinal white dash on
apex and a white dash on margin either side half way between
apex and eye. A short indistinct dash is on base either side of
middle. Pronotum heavily mottled with dark brown and with five
pale longitudinal lines. Scutellum with basal angles dark brown,
the central third paler, the apical angle white margined. Elytra
pale, veins dark brown and with dark brown coloration and pig-
ment lines rather uniformly distributed. Many white areolar spots
in most of cells of fore wing. Face yellow, dusky above.
Genitalia: Female last ventral segment gradually, broadly, pro-
duced, slightly notched at middle. Male plates triangular, about
as long as combined basal width. Style rather short, deeply ex-
cavated at apex to form a finger-like process on inner apical mar-
gin. Aedeagus curved so as to form a long slender basal portion
which is slender and curves dorsally, and apical portion which
curves dorsally and tapers to apex which is broadened by a tri-
angular process on each side at apex.
Holotype male and allotype female collected at Cosomatepec,
Ver. Mexico, November 30, 1924 (M. B. 103). Paratype male
from Popaiaopam, Ver. Mexico, December 12, 1937 (M. B. 397),
all collected by Dr. Dampf.
Cloanthanus liambletoni De Long, new species
Resembling lineafrons in form and general appearance but with
distinct coloration and male genitalia. Length: male, 5 mm.
Vertex strongly produced and bluntly angled more than twice
as long as basal width! between the eyes.
Color: Vertex pronotum and scutellum tawny, the disc of pro-
notum a little darker. Vertex with a pale longitudinal median line
on apical third and, one next either eye. At the end of the apical
median line a longitudinal line arises either side of middle and
130
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
extends to base of vertex. Pronotum with five pale longitudinal
lines. Scutellum with a pale spot in each basal angle and a longi-
tudinal line either side equidistant from basal anglei and each
other. Elytra pale, veins dark brown and with brown pigment in
most of cells. Apical margin broadly brown and with numerous
brown markings resembling veinlets. Face dark brown, margin
pale and with narrow pale arcs.
Genitalia: Male plates short, strongly convexly rounded to near
apex where they are produced to form bluntly pointed apices.
Style deeply, rather broadly concavely excavated at apex to form
a long, outwardly curved finger-like process on inner apical mar-
gin. Aedeagus composed of a dorsal L-shaped process and a pair
of long ventral processes. The ventral processes are gradually
widened on apical half to form broad truncate apices with a long
pointed spur on dorsal apical angle and a short, blunter spur on
ventral apical angle.
Holotype male, collected at Campinas Sag Paulo, Brazil,
April 12, 1936, by E. J. Hambleton.
Cloanthanus fasciatus De Long, new species
A robust, blunt headed species with banded elytra, and distinct
male genitalia, most closely related to phimmen. Length, male,
5.5 mm.
Vertex short, blunt, scarcely angled a little shorter at middle
than basal width between the eyes. Vertex slightly depressed on
median line.
Color: White irr orate with brown. Vertex and pronotum rather
evenly irrorate with brown except for depressed portion just poste-
rior to apex of vertex which is brown. Scutellum irrorate with
brown, basal angles dark brown. Face heavily irrorate with brown.
Margin above appearing to have four black spots on a white band.
A short white transverse band just beneath margin. Elytra white
with irr orations forming two rather definite broad bands. The
anterior band extends across basal half of clavus. The posterior
band extends from apex of clavus to tip of elytra.
Genitalia: Male plates short, broad at base, convexly rounded
to blunt apices. Style short and broad. The finger process on the
inner margin rather heavy and more than half as long as the basal
portion of style, sloping caudally and outwardly. Aedeagus with
a short, thick curved dorsal portion which is bifid on basal half.
This is connected by a slender process to the basal portion which
is rather broad in ventral view and has a pair of rather short
thick divergent spines at apex.
Holotype male collected at PuEBLA, Puebla, Mexico, eleva-
tion 8,500 feet, October 18, 1941, by E. E. Good, J. S. Caldwell,
C. C. Plummer and the author. This blunt headed species is
large, robust and distinct from any others of this genus.
OCT. 1944]
DE LONG— CICADELLIDAE
131
Tumeus irrorellus De Long, new species
Resembling divisus in general form but with distinct coloration
and male genitalia. Length, male 6 mm.
Vertex produced with margin convexly rounded at apex to a
blunt tip, twice as long at middle as basal width between the eyes.
Color: Vertex, pronotum and scutellum brown with pale irrora-
tions. The margin of the vertex at apex is dark brown and the
disc of the pronotum is darker. There are traces of the five pale
longitudinal lines on the pronotum. Elytra pale with dark brown
irrorations and ramose pigment lines,, a few white areolar spots
on all portions of the fore wing. Face dark brown with pale irro-
rations.
Lateral view of male genitalia of species as labelled.
Genitalia: Male plates rather short, triangular, combined width
at base about one-third greater than length. Valve broad and
rounded. Style deeply narrowly notched on outer margin near
apex to form a narrow finger-like process on apical third. Aedea-
gus iri lateral view rather broad, with a basal spur-like process
extending anteriorly. The apical half is bent sharply dorsally and
is inflated, the apex is blunt and rounded.
Holotype male collected at Campinas, Sag Paulo, Brazil,
April 12, 1936, by E. J. Hambleton.
132
THE PAN-PACIFIC ENTOMOLOGIST
[ VOL. XX, NO. 4
NEW SPECIES AND RECORDS IN MEGACHILE
(Hymenoptera, Megachilidae)
BY THEODORE B. MITCHELL^
University of North Carolina, Raleigh
The material described and recorded here was received in
large part from C. D. Michener, P. H. Timberlake and G. E.
Bohart in California and R. H. Beamer in Kansas, with various
scattered records from other sources.
Subgenus Argyropile Mitchell
Megachile (Argyropile) parallela Smith
Megachile parallela Smith, 1853, Cat. Hymen. Brit. Mus., 2:191.
Megachile verhesinae Cockerell, 1908, Ann. Mag. Nat. Hist.
(8)1:264. New synonomy.
In the revision of Nearctic Megachile^ the suggestion was
made that verhesinae should be considered a synonym of paral-
lela, but the observation was unsupported by personal examina-
tion of the types. The specimen here recorded, which is a
typical parallela in all respects, has been compared with the
type of verhesinae by Timberlake and found to be identical.
Arizona: 12 Sonoita, August 9, 1940 (Timberlake, on Ver-
hesina ) .
Megachile (Argyropile) nebraskana Mitchell
Colorado: 12, White Rocks, near Boulder, June 29, 1939
(Timberlake, on Monarda ) .
Megachile (Argyropile) subparallela Mitchell, new species
Female. Size: Length, 8.3 mm.; breadth of abdomen 3 mm.;
anterior wing, 6 mm.
Structure : Face slightly broader above than distance from
clypeal margin to anterior ocellus; eyes very slightly convergent
below; clypeal margin with a small acute triangular projection
separating a pair of small emarginations ; mandibles 4-dentate,
^ Research Contribution No. 20, published with the aid of the State College
Research Fund, Department of Zoology, North Carolina State College of Agricul-
ture and Engineering of the University of North Carolina.
- 1937, Trans. Amer. Ent. Soc., 63 :67.
OCT. 1944]
MITCHELL— MEGACHILE
133
the teeth acute, a conspicuous cutting edge between the third and
fourth, a mere vestige of one between the second and third; basal
segment of flagellum very slightly longer than pedicel, the follow-
ing segments only slightly longer than broad; lateral ocelli very
slightly nearer to edge of vertex than to eyes; vertex nearly
flat and hind margin only slightly incurved; cheeks below subequal
to eyes in width, somewhat narrower above; metatarsi somewhat
shorter and narrower than their tibiae, but relatively robust; ab-
domen cordate in form, the sixth tergum straight in profile, with
suberect hair visible toward base; apical margin of the sixth
sternum thickened and slightly reflexed.
Puncturation : Fine and close on dorsum of thorax, more coarse
on pleura, vertex, clypeus and cheeks below, fine and close on
cheeks above and face above antennae; minute and densely crowded
on basal segments of abdomen, becoming relatively coarse and
sparse on fifth tergum, fine and crowded on the sixth.
Color: Black; wings subhyaline, nervures ferruginous to fus-
cous; tegulae yellowish-ferruginous along outer margin, darker
within; spurs yellowish-ferruginous.
Pubescence : White and I'ather dense on face and clypeus, cheeks
below, pleura, propodeum and base of abdomen; vertex with some
inconspicuous darker hairs interspersed with pale hairs, mesono-
tum, scutellum, and discs of segments 2-5 with short inconspicuous
dark pubescence, the sixth sparsely covered with subappressed
yellowish hairs; segments 2-5 with narrow entire yellowish apical
fasciae; scope creamy- white, that on sixth sternum very sparse
and including a few dark hairs.
Male: Size: Length, 9.7 mm.; breadth of abdomen, 3,4 mm.;
anterior wing, 7 mm.
Structure: Face about equal in breadth to distance from clypeal
margin to median ocellus; eyes convergent below; clypeal margin
beneath beard smooth and shining, slightly incurved; mandibles
3-dentate, the inner tooth robust and triangular, the basal inferior
projection slender; basal segment of flagellum longer than pedicel,
the following segments twice as long as broad, lateral ocelli sub-
equally distant from eyes and edge of vertex; vertex nearly flat,
hind margin nearly straight; cheeks subequal to eyes in width, not
excavated below; front coxal spines short, the tips triangularly
acute; front tarsi long, slender and unmodified; front tibiae and
femora unmodified; mid and hind legs slender, the metatarsi nar-
row and quite short, the mid tarsal segments not at all produced
apically; segments of abdomen quite deeply grooved basally, the
apical margins deeply and abruptly depressed except on the basal
segments medially, the disc of the fifth tergum carinate apically
and slightly overhanging the depressed apical margin; carina of
sixth tergum semi-circular in form, with a deep rounded median
emargination, apical margin of the tergum with robust subtruncate
carinate median teeth which are much nearer the small triangular
lateral angles than to each other; seventh tergum obscure, rounded
134
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
apically; fourth sternum exposed; presternite of fifth sternum
almost linear medially, with a patch of pubescence apically on each
side, the medasternite beset with setae, the tips of which are
broadly dilated and flexed, a fringe of simple setae around basal
margin, the poststernal strip straight and simple; medasternal
areas of sixth sternum contiguous medially at base, closely cov-
ered with short robust setae with strongly dilated and flexed tips,
the poststernal lobe very broadly semi-circular in outline; cardo.of
genital armature quite narrow, stipites entirely bare, slightly di-
lated and flattened apically and slightly exceeded in length by the
simple sagittae, volsellae quite robust, triangularly pointed apically.
Puncturation : Fine and rather sparse on dorsum of thorax, the
surface dull and granular; somewhat more coarse and close on
pleura and propodeum, but obscured by dense pubescence; moder-
ately coarse and rather sparse on vertex medially, becoming fine
and close on cheeks below; rather fine and close on basal segments
of abdomen, becoming more coarse and more widely separated to
the fifth tergum, close and substriate on the sixth above carina.
Color; Black; antennae beneath more brownish; apices of front
and mid tibiae ferruginous; tegulae yellowish-ferruginous on outer
portion; wings subhyaline, nervures piceous to ferruginous; spurs
yellowish.
Pubescence: Pale throughout; long and dense on face, clypeus,
cheeks, pleura, propodeum, and base of abdomen, front tarsi with
a rather thin white fringe; segments 2-4 of abdomen with entire
white apical fasciae, the fifth not fasciate but with some basal
white tomentum, the sixth with dense white tomentum covering
most of disc above.
f
Holotype female Douglas, Arizona, August 16, 1936 (W. W.
Jones), allotype male, topotypical (W. W. Jones), [both Tim-
berlake].
The male of suhparallela differs from that of parallela in the
following respects: The definitely 3-dentate mandibles (either
obscurely or distinctly 4-dentate in parallela) ; the absence of
a median tubercule on the clypeal margin; the median apical
teeth of the sixth tergum are truncate and carinate (usually
slender and acute in parallela but subject to considerable varia-
tion) ; the seventh tergum lacking the slight median indentation
found in parallela’, and the lack of a distinct apical fascia on
the fifth tergum. The range of variation in parallela is consid-
erable, but this form seems to lie outside that range.
The female of subparallela runs to sahinensis in the key re-
cently published.^ However, sahinensis is much larger and the
3 1943, Pan-Pac. Ent., 19 :12.
OCT. 1944]
MITCHELL— MEGACHILE
135
clypeal margin is relatively straight, the median tubercle being
minute and not delimited by the emarginations found in suh-
parallela. Moreover, in sabinensis the scopa on the sixth sternum
is entirely black, this being almost entirely pale in suhparallela,
and the sixth tergum bears conspicuous white tomentum medi-
ally, with darker erect basal hairs and appressed fuscous tomen-
tum apically. In suhparallela the disc of this segment is quite
uniformly covered with rather sparse suberect yellowish pubes-
cence, with neither definitely black nor white hairs in evidence.
Subgenus Delomegachile Viereck
Megachile (Delomegachile) frigida Smith
Before her death. Dr. Grace Sandhouse called my attention to
the fact that I was in error in my use of the name vidua instead
of frigida for this species. Smith proposed both names, vidua
for the female on p. 192 and frigida for the male on p. 193 (Cat.
Hym. Brit. Mus. 1, 1853). As the name vidua has page preced-
ence, it was thought proper to use. HoweVer, Cresson in his
Synopsis of Hymenoptera, p. 303, 1887, made vidua a synonym
of frigida, in spite of the page precedence of vidua, and accord-
ing to article 28 of the International Rules of Zoological No-
menclature, “If the names are of the same date, that selected by
the first reviser shall stand.”
Subgenus Xeromegachile Mitchell
Records of four species, namely deflexa^ manifesta, nevad-
ensis, and wheeleri, are omitted in the following list, since they
add little to the present knowledge of their distribution.
Megachile (Xeromegachile) alata Mitchell
California: 1$, Mt. San Jacinto, Riverside County, May 28,
1939. 1 $ , Argus Mts., Inyo County, June 4, 1939. 1 5 , N. Inyo
Mts., Inyo County, June 7, 1939 (all R. M. Bohart) . 2$, Taquitz
Canyon, Riverside County, April 16, 1938 (R. M. and G, E.
Bohart). 2$, Independence, Inyo County, June 1, 1937 (W. C.
Reeves). 15 Palm Canyon, April 9, 1938 (E. P. Van Duzee) .
Megachile (Xeromegachile) bradleyi Mitchell
Utah: 1$, Orr’s Ranch, September 11, 1935 (G. F. Knowl-
ton) .
136
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 4
Megachile (Xeromegachile) brimleyi Mitchell
Florida: 1^, Cedar Keys, August 10, 1939 (R. H. Beamer).
North Carolina: 3^, 10 miles south of Lillington, July 7,
1938 (Mitchell and Hill). 1$, Carolina Beach, July 25, 1941
(T. B. Mitchell).
Megachile (Xeromegachile) casadae Cockerell
Colorado : 1 $ , 8 miles south of Grand Mesa, July 12, 1938,
scrub oak zone (U. Lanham). 12, Mishawauka, July 11, 1937
(H. T. Peters). Oklahoma: 1$, Hinton, June 5, 1939 (Kaiser-
Nailon) .
Megachile (Xeromegachile) coloradensis Mitchell
Colorado: 12, Buffalo, August (E. S. Tucker).
Megachile (Xeromegachile) dakotensis Mitchell
Iowa: 12 2, County 3, July 10-14, 1936 (Anderson, Barry,
Jacques and Millspaugh). Texas: 12 , Romney, June 30, 1936
(R. H. Beamer).
Megachile (Xeromegachile) fucata Mitchell
California: 1^, Essex, May 29, 1937 (M. A. Cazier). 3^,
Inyo Mts., Inyo County, May 23, 1937 (W. C. Reeves). 1^,
Mazourka Canyon, Inyo Mts., 6,000 feet, June 1, 1937 (Mich-
ener, on Encelia farinosa ) . 1 ^ , Argus Mts., Inyo County, May
22, 1937. 1^, Inyo Mts., June 1, 1937 (E. C. Van Dyke).
Megachile (Xeromegachile) histrata Mitchell
Arizona: 12 , Tucson, April 24, 1937 (W. Benedict). Cali-
fornia: 12 , Mohave Desert, 1932. 12 , N. Inyo Mts., Inyo
County, June 7, 1939 (R. M. Bohart).
The specimen from N. Inyo Mts. bears the same data as a
male of Xeromegachile alata (see page 135) suggesting the prob-
ability that the two are the same. If this proves to be the fact,
histrata will become a synonym of alata since the latter has page
precedence.'*
^ 1934, Trans. Amer. Ent. Soc., 59 :324, 334.
OCT. 1944]
MITCHELL— MEGACHILE
137
Megachile (Xeromegachile) hookeri Cockerell
Colorado: 1$, (no data) (T. Pergande). Utah: 39, Desert,
September 9, 1938 (L. L. Hanson). 3 9, Valley Mts., 7,000 feet,
September.
Megachile (Xeromegachile) impartita Mitchell
California: 19, Westgard Pass, Inyo County, May 27, 1937
(R. M. and G. E. Bohart). 19, Lone Pine, Inyo County, May
23, 1937. 19, Inyo Mts., June 1, 1937 (both E. C. Van Dyke).
Megachile (Xeromegachile) Integra Cresson
Florida: 5$, 3 9, Suwannee Springs, August 2 and 3, 1939
(Beamer, Hardy and Wegenek) . 19, Suwannee Springs, July
29, 1930 (P. W. Oman). 2S, Branford, July 31, 1930. 1^,
Lacoochee, August 18, 1930 (both J. Nottingham). 19, San-
ford, August 8, 1939 (R. H. Beamer).
Megachile (Xeromegachile) legalis Cresson
Megachile legalis Cresson, 1879, Trans. Amer. Ent. Soc., 7 :209.
Megachile {Xeromegachile) couleeana Mitchell, 1938, Pan-Pac.
Ent., 14:171. New Synonomy.
The true identity of the female described under the name
couleeana has become evident by the collection by Timberlake
of a series of 2 males and 6 females at the same time and on the
same flower, in the absence of any other closely related species.
Only 3 specimens of this series have been examined personally,
the record of which, together with others, follows.
Arizona: 15, Flagstaff, June 11, 1909 (F. C. Pratt, on Iris).
California: 15, Independence, Inyo County, June 11, 1937
(W. C. Reeves) Nevada: 15, 29, Kyle Canyon, Charleston Mts.,
5,000 feet, June 3 and 4, 1941 (Timberlake, on Cirsium). Ore-
gon: 19, Culver, July 2, 1935 (R. H. Beamer).
Megachile (Xeromegachile?) maurata Mitchell
For a note concerning the status of this species, see M. {Dero-
tropis) subanograe, page 139.
California: 35, Panamint Mts., 5 miles north of Wildrose
Canyon, 5,000 feet, Inyo County, May 27, 1937 (Michener, on
Sphaeralcea ambigua) . 15, Panamint Mts., May 30, 1937 (E. C.
Van Dyke).
138
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
Megachile (Xeromegachile) mojavensis Mitchell
California: 1$, Lone Pine, Inyo County, May 19, 1937
(W. C. Reeves) . 1 $ , Lone Pine, May 23, 1937 (E. C. Van Dyke) .
Megachile (Xeromegachile) mucorosa Cockerell
Colorado: 2$, (Snow). Kansas: 1^, Clark County, (F. H.
Snow). Texas: liJ, Romero, June 22, 1940 (E. E. Kenaga),
Megachile (Xeromegachile) oslari Mitchell
Colorado: 2$, Durango, 6,300 feet, June 3 and 11, 1936 (B.
Rotger) .
Megachile (Xeromegachile) palmensis Mitchell
California: 2^, Taquitz Canyon, Riverside County, April
16, 1938. 5 ^ , 4 $ , Palm Canyon, Riverside County, April 15,
1938, on Sphderalcea ambigua and Encelia farinosa (both R. M.
gnd G. E. Bohart) .
Megachile (Xeromegachile) soled adensis Cockerell
Arizona: 1^, Picacho Pass, August 7, 1940. 15, Pepper
Sauce Canyon, Santa Catalina Mts., August 16, 1940 (both E. S.
Ross). 15, Sonoita, August 9, 1940 (Timberlake, on Verhesina
enceloides) . Texas: 1 5 , El Paso, August 21, 1908 (F. C. Pratt).
Megachile (Xeromegachile) subnigra Cresson
California: 55, Leavitt Meadow, Walker River, Mono Coun-
ty, June 25, 1937 (E. C. Van Dyke) . 15, Siskiyou County, June
2, 1911. 75, 32, W. Walker River, Mono County, 7,200 feet,
June 25, 1937 (Michener, on Aster and Senecio) . Idaho: 12,
Craters of Moon, June 20, 1938 (E. C. Van Dyke). Nevada:
15, 25 miles northwest of Geriach, Washoe County, May 29,
1939 (Ting, Cazier, Downes and Aitken). Washington: 12,
Clilfdell, July 7, 1935 (E. I. Beamer) . Wyoming: 15, Grand
Teton National Park, July, 1937 (R. M. and G. E. Bohart
Megachile (Xeromegachile) subnigra var. angelica
Mitchell
California: 15, Westgard Pass, Inyo County, June 3, 1937
(N. W. Frazier). 15, Westgard Pass, June 15, 1937. 25, Ma-
zourka Canyon, Inyo Mts., 6,000 feet, June 1, 1937 (both Mich-
OCT. 1944]
MITCHELL— MEGACHIDE
139
ener, on Encelia farinosa) . , Feather River, Butte County,
June 14, 1940 (Cazier and Aitken). 2 c?, Westgard Pass Plateau,
June 3, 1937. 6 $ , Lone Pine, Inyo County, May 23-24, 1937
(both E. C. Van Dyke). 1^, Mt. San Jacinto, Riverside County,
May. 28, 1939 (both R. M. Bohart). Oregon: 1^, Quartz Creek,
20 miles east of Ely, Lake County, 5,500 feet, July 8, 1937. 1 $ ,
Cherry Creek, west side Klamath Lake, 4,175 feet, June 26, 1937
(both Bolinger and Jewett). Idaho: 19, {hlaisdelli) , Craters
of Moon, June 20, 1938 (E. C. Van Dyke).
Subgenus Derotropis Mitchell
Four species of Derotropis are known in both sexes, gravita
( 9 astata) , pascoensis ( 9 gabrielensis) , xerophila and astragali.
Four other species are known only in one sex, the male only of
alamosana and the females only of anograe, suhanograe'^' and
laurita. It now seems probable that alamosana is the male of
anograe, judging from the range, these being the easternmost in
distribution, and from the similarity of the two forms to the
respective sexes of gravita and pascoensis. This relationship,
however, remains to be demonstrated. Some evidence concerning
the identity of the male of subanograe can now be presented, but
that of laurita still has to be discovered.
Megachile (Derotropis) subanograe Mitchell
A series of five females of this species received from C. D.
Michener was accompanied by three males of M. {Xer omega-
chile) maurata Mitchell, the two series bearing identical data as
to locality, date and flower record. This suggests their common
identity, and if proven correct, results in obscuring the distinc-
tions between the males of Xeromegachile and Derotropis. They
are quite similar in size and in general appearance, and to that
degree the association seems a reasonable one.
An additional question concerns the range of variation of the
females of this species. A number of specimens from Arizona,
California, Nevada, and Texas have been received, all bearing
strong resemblances to typical subanograe, but with enough dif-
ferences to indicate the possibility, at least, that they represent
different species. Were these accompanied by males, a more
* The specimen recorded on p. 174, Pan-Pacific Ent. 14, 1938, evidently is a.
female, not a male as indicated.
140
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
positive opinion in the matter would be warranted. Since the
males are not known, two of these variants are described below,
and the test of their validity is left to the future. Records of the
more typical subanograe follow.
California: 1$, Panamint Mts., Inyo County, May 29, 1937
(N. W. Frazier). 5$, 5 miles north of Wildrose Canyon, Pana-
mint Mts., Inyo County, 5,000 feet, May 27, 1937 (C. D. Mich-
ener, on Sphaeralcea amhigua) . Utah: 29, 12 miles east of
Jensen, June 23, 1939 (Timberlake, on Sphaeralcea) . Nevada:
1 9 , Lee Canyon, Mt. Charleston, Clark County, May 24, 1940
(Reeves, Cazier and Ting).
Megachile (Derotropis) melanderi Mitchell, new species
Female. Size: LfCngth, 9 mm.; breadth of abdomen, 3.5 mm.;
anterior wing, 7 mm.
Structure: Length and breadth of face equal; eyes nearly paral-
lel; apical margin of clypeus nearly straight and entire; mid tooth
of mandible about twice as distant from inner angle as from apex
of mandible; second joint of labial palpus not greatly exceeding
the first in length (ratio of about 10-7) ; second segment of flagel-
lum subequal to pedicel, the first considerably longer, middle seg-
ments about as long as broad, apical one longer; lateral ocelli
subequally distant from eyes and edge of vertex; vertex nearly
flat, its posterior margin only slightly curved; cheeks subequal to
eyes in width; mid and hind metatarsi nearly as broad as their
tibiae and about four-fifths as long; sixth tergum straight in pro-
file, with erect hairs evident.
Puncturation : Fine and close on middle of face, supraclypeal
area, upper margin of clypeus, on either side of middle of vertex,
on scutellum except in center and on upper portions of pleura;
more coarse and irregular on clypeus medially, fading out toward
the margin; shallow and obscure on cheeks, becoming sparse be-
low; relatively sparse above ocelli; deep and distinctly separated
on mesonotum, rather widely so in center, as also on scutellum;
more coarse on pleura below; minute and obscure but quite close
on abdomen basally, becoming increasingly sparse to the fifth
tergum, more close, deep and coarse on the sixth.
Color: Black; flagellum more fuscous; tegulae ferruginous with
fuscous blotches; wings subhyaline, somewhat clouded apically,
nervures fuscous to blackish; front and middle tibial spurs yellow-
ish, hind spurs more fuscous.
Pubescence: Entirely white on head, thorax, legs and segments
1 and 2 of abdomen; black and erect on discs of segments 3-5, the
sixth with scattered erect black hairs and subappressed silvery
tomentum; segments 1-5 with dense, entire and conspicuous white
apical fasciae; scopa white, black on fifth and sixth sterna.
OCT. 1944]
MIT CHELL^MEGA CHILE
141
Holotype female: Marfa, Texas, April 28, 1942 (A, L. Mel-
ander) [Timberlake] .
Megachile (Derotropis) yumensis Mitchell, new species
Female. Size: Length, 9.3 mm.; breadth of abdomen, 3.5 mm.;
anterior wing, 6.5 mm.
Structure: Breadth of face about equal to distance from clypeal
margin to median ocellus; eyes slightly convergent below; clypeal
margin broadly impunctate, straight and simple; mandibles 3-
dentate, distance from apex of inner tooth to that of middle tooth
not quite twice that from the middle to the apical tooth, a promi-
nent cutting edg-e between the second and inner teeth; second joint
of labial palpus not greatly exceeding the first in length (ratio of
about 11-8) ; basal segment of flagellum longer than pedicel or the
second segment, the median segments only about as long as broad;
lateral ocelli subequally distant from eyes and edge of vertex;
vertex slightly rounded; its hind margin very slightly incurved;
cheeks subequal to eyes in width; all the metatarsi slightly nar-
rower and considerably shorter than their respective tibiae; abdo-
men cordate in form; sixth tergum straight in profile, with abun-
dant erect hair visible.
Puncturation : Rather fine on head, irregular and rather deep
on clypeus beneath the dense pubescence, close on face, rather close
on vertex medially, more sparse laterally, irregular and shallow
on shining cheeks; slightly more coarse and deep on thorax, well
separated in center of mesonotum and scutellum, close on pleura,
but becoming fine and sparse anteriorly; minute on abdomen, close
and indistinct toward base, increasingly sparse to the fifth tergum,
quite close on the sixth.
Color: Black, antennae deep ferruginous beneath; apices of
front and mid tibiae ferruginous; tegulae piceous; wings hyaline,
slightly clouded apically, the nervures fuscous; front and middle
spurs yellowish, hind spurs more fuscous.
Pubescence: Entirely white on head, thorax and segments 1
and 2 of abdomen, dense on face, clypeus, cheeks below and pleura;
discs of third to fifth terga with intermixed dark and light hairs,
that on the sixth entirely white, rather sparse; segments 1-5 with
broad dense entire white apical fasciae; scopa white, black on sixth
sternum.
Holotype female: Yuma, Arizona, March 30, 1940 (R. C.
Dickson) [Timberlake]. Paratype, 19, topotypical.
In the paratype a slight amount of variation is to be noted,
in that the tegulae and hind spurs are somewhat paler in color,
the short pubescence on the discs of the abdominal terga is almost
entirely pale, and there is a slight amount of black in the scopa
along the hind margin of the fifth sternum.
142
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
Megachile (Derotropis) anograe Cockerell
California: 1$, Santa Paula, June 15, 1927. Colorado: 1$,
Mishawauka, July 11, 1937 (H. T. Peters). Also recorded from
Inspiration Point, Denver, July 2, 1933. Montana: 1$, Jeffer-
son County, July 24, 1924.
Megachile (Derotropis) gravita Mitchell
California: IS, Jamesburg, Hastings Nat. Hist. Reserve,
Santa Lucia Mts., Monterey County, 1,900-2,700 feet, May 10,
1938 (Michener). 1$, Cal. Hot Springs, Tulare County, June 2,
1939 (E. C. Van Dyke). 1^, Cuyamaca Lake, July 6, 1929
(R. H. Reamer). Oregon: IS, Grave Creek, at Reuben Creek,
20 miles north of Grants Pass, June 16, 1937 (Bolinger and
Jewett). 1$, Prospect, June 20, 1924 (C. L. Fox).
Megachile (Derotropis) pascoensis Mitchell
California: 2S, Coalings, May 14, 1938 (M. Cazier). IS,
1$, Antioch, May 8 and 11, 1937. 1$, Mt. Diablo, May 12,
1937 (both R. M. and G. E. Bohart) . 1 $ , Briceburg, Mariposa
County, June 3, 1938 (R. M. Bohart). 2S , 2 9, Jamesburg, Hast-
ings Nat. Hist. Reserve, Santa Lucia Mts., Monterey County,
1,900-2,700 feet. May 21, June 2 and 20, 1938, on Convolvulus
villosus and Trifolium (C. D. Michener). 19, Cal. Hot Springs,
Tulare County, June 2, 1939 (E. C. Van Dyke). 19, San Luis
Obispo, June 17, 1938 (1. McCracken). IS, Antioch sand dunes,
June 4, 1942 (H. A. Scullen). Idaho: 1^, St. Joe River, July
12, 1927 (M. C. Lane).
Megachile (Derotropis) alamosana Mitchell
Oregon: 1^ , Boardman, Morrow County, June 4, 1943 (G. R.
Ferguson) .
Key to Species of Derotropis
FEMALES
1. Scopa entirely black laurita Mitch, (and varieties)
- Scopa pale on segments 1-4 2
2. Mesonotum dull, tessellate, with very fine and sparse punctures
xerophila Ckll.
- Mesonotum more coarsely and closely punctate, the surface be-
tween the punctures shining 3
3. Mesonotum and scutellum uniformly densely and rather finely
punctate gravita Mitch.
OCT. 1944]
MITCHELL— MEGACHILE
143
- Punctures in center of mesonotum separated at least by a punc-
ure width : 4
4. Vertex and scutellum dull, with the punctures fine and very
close, no shining spaces evident pascoensis Mitch.
- Vertex and scutellum more shining, with definite spaces evi-
dent between the punctures 5
5. Scopa on the fifth sternum entirely white, largely dark on the
sixth; distance from middle tooth of mandible to the inner
tooth not quite twice that from the middle tooth toi the apex
yumensis Mitch,
- Scopa entirely black on fifth and sixth sterna 6
6. Pubescence of discs of segments 1-4 of abdomen entirely or
largely pale, with very inconspicuous darker hairs or none;
the sixth with erect black hairs and subappressed inconspicu-
ous silvery tomentum swhanograe Mitch,
- Third and usually the second terga with conspicuous black
pubescence 7
7. Sixth tergum with erect black hairs, otherwise bare, with no
evidence of pale tomentum; distance from middle to inner
teeth of mandible fully three times that from middle tooth to
apex anograe Ckll.
- Sixth tergum with at least some silvery tomentum; distance
from middle to inner teeth of mandible not so extreme 8
8. Sixth tergum uniformly covered with fine suberect silvery
pubescence, with no dark hairs in evidence, the punctures uni-
formly fine and close even at base; hind metatarsi very narrow,
hardly more than half as wide as their tibiae.. ..astragali Mitch.
-""Sixth tergum with conspicuous erect black hairs; hind meta-
tarsi nearly equal to their tibiae in width melandon Mitch.
MALES
1. Front tarsi black, entirely simple, neither dilated nor exca-
vated astragali
- Front tarsi yellowish or ferruginous, dilated and excavated....2
2. Mesosternum with a pair of acute tubercles anterior to the
mid coxae gravita
- Mesosternum not spinose 3
3. Mesonotum dull and tessellate, almost impunctate medially;
seventh tergum simple xerophila
- Mesonotum more shining, distinctly punctate medially; sev-
enth tergum wits a slender apical spine 4
4. Fifth tergum polished, the punctures sparse and minute, its
apical margin not fasciate alamosana
- Fifth tergum more closely and coarsely punctate, and with a
distinct white apical fascia 5
5. ‘Apical margin of clypeus with a broad and deep median emar-
gination pascoensis
- Apical margin of clypeus entire maurata (subanograe?)
144
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 4
STUDIES AMONG THE COCCINELLIDAE— No. 9
(Coleoptera)
BY F. W. NUNENMACHER
Piedmont, California
Zenora tricolor Nunenmacher, new species
Light yellow, black and brick red. Head and pronotum yellow,
with the exception of a faint trace of black in front of the scutellum
where the black of the elytra overlaps. The simplest way to de-
scribe the elytra is to say they are black with yellow margins and
a brick red disc on the suture at about the center. The size of this
disc is from about one-fourth the elytral length to beyond the mid-
dle, and from not quite the center of each elytron. Unequal coarse
punctures. The little remaining pubescence is white. Ventral sur-
face, mouth parts, antennae, legs, sides and segments yellow, breast
a little darker. Length 3 mm., width 2.25 mm.
Holotype female, in the author’s collection, from Para Ama-
zon, no other data.
Crotch, Revision of the Coccinellidae, pp. 277-278, gives the
following description: Z. revistita Muls. Var. E. Elytra metallic
green, the margins from the shoulder to the apex pale ochrous
but the disc pale. I do not think my species belongs to this group.
Th. Dobzhansky, in Beetles of the Genus Hyperaspis Inhabit-
ing the United States, 1941, says he cannot place the following
three of my species: falli, floribunda and walcotti. I would like
to make this clear to the students: falli is a true Hyperaspis’,
floribunda and walcotti belong in Hyperaspidius. And to clear
the note part of his statement, at that time I did not think that
the genus Hyperaspidius would stand. The genus is principally
based on the fova or depression of the epipleura which receives
the knees of the hind pair of legs. I was then working on a lot
of South American species of Hyperaspis and had noted that in
many cases this fova is very shallow, and must be viewed in a
very good, clear light to be seen. I still think it is a weak genus.
The following three species of Hyperaspidius are undescribed,
and I take this opportunity to make them known.
OCT. 1944]
NUNENMACHER— COCCINELLIDAE
145
Hyperaspidius juniperus Nunenmacher, new species
Head straw yellow. Pronotum rufescent, margins and angles
lighter. Elytra about as long as wide with tip slightly truncate,
with a median and a common sutural black vitta. The median
vitta somewhat broader than the light space at either side, starting
at the callus and running parallel to the margin about five-sixths
the length where it curves, becoming a narrow line which joins
the sutural vitta. The sutural vitta at base a little wider than
the scutellum, gradually widening, broadest at about one-third the
leng-th, then narrowing to about two-thirds, from here remaining
about the same width as at base to the tip of the elytra. Very
finely punctured. Ventral surface and legs rufescent. Length 2
mm., width 1.25 mm.
Holotjpe male, in collection of author, from TehacHAPI Pass,
Kern County, California, June 8, 1940, on Juniperus califor-
nicus. Collected by R. G. Dahl who very kindly gave it to me.
Hyperaspidius rossi Nunenmacher, new species
Black and yellow. Head and pronotum deep shiny black, very
finely punctured. Elytra longer than wide, tip slightly truncate;
with a narrow yellow marginal band starting at the side of the
scutellum to about the middle where it is interrupted for a short
space, then it starts again just around the humeral angle along
the margin not quite to two-thirds the length where it is a little
broader; also a small spot at the tip, the same distance from the
sutural margin and the tip of the elytron*, and a small elongate
spot at two-thirds, about its own width from the suture and about
twice as long as wide. Finely punctured, a little stronger than on
pronotum. Ventral surface and legs black, anterior tibiae and
tarsi pitchy. Length 2 mm., width 1.25 mm.
Holotype female, in collection of author, from Oregon, July,
1938, with no other data. This pretty little specimen was kindly
given to me by Dr. E. S. Ross some years ago.
Hyperaspidius shauli Nunenmacher, new species
Yellow and black. Head and pronotum rufescent, margins and
angles lighter; females slightly darker than males. Elytra about
as long as wide, tip not truncate. With a median and a common
sutural black vitta. The median vitta starts at the callus and runs
parallel to the margin to about five-sixths, where it curves and be-
comes a narrow line joining the sutural vitta. In the male this
vitta is only about half as broad as in the female. The sutural
*I think a series of specimens will show this band and also the spot at the tip
united to form a solid vitta.
146
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
vitta starts at the base, just covering the scutellum, and is the
same width to the tip. Very finely punctured. Ventral surface
rufescent, the epipleura yellow. Length 2 mm., width 1.25 mm.
Holotype male, allotype female and four paratypes, all from
Perryton, Texas, August 13, 1937, collected by R. E. Shaul.
Types and two paratypes in the author’s collection, one paratype
female in J. E. Blum’s collection.
This species is narrower, lighter in color, and the vitta are
narrower than in oblongus Casey, as I remember his type.
Coccinella ampla var. rufa Nunenmacher, new variety
Coccinella ampla seems to be very variable even among those
with complete maculations to albino forms. In July, 1937, A. R.
Mead collected Coccinellidae in Mexico for me. Among his ma-
terial I find seven specimens of this species collected in Oaxaca,
Mexico, that are rufescent. The color and pattern of the prono-
tum is fairly constant, with the exception of a yellow border,
which in one or two specimens has a pale reddish color. The
elytra are rufescent with a narrow pale margin. In the darkest
colored specimens there is a trace of the black markings of the
type form. This variety may take the name rufa Nunemacher,
n. var.
Types and five paratypes remain in the author’s collection.
AN ABERRANT SPECIMEN OF AGLAIS ANTIOPA (LINN.)
On September 3, 1944, while collecting in the open forest-
land near the Donner Summit Inn, at an altitude of approxi-
mately 7,000 feet an aberrant specimen of Aglais antiopa was
seen. The specimen was observed closely, although it subse-
quently evaded capture and disappeared. It was undoubtedly of
the type which has been named hygiaea Heydenr., as all other
named aberrations of this species vary in other respects than
the great widening of the yellow border and loss of the sub-
marginal band of purple spots. In this specimen the yellow
border was greatly broadened and diffused inwardly, occupying
at least one-third of the width of the wing. In this respect it
equalled, or perhaps even surpassed the width in the specimen
figured by J. A. Comstock in Butterflies of California, Plate
42, figs. 3 and 5. Only a slight darkening apparently remained
of the submarginal band of spots. — R. W. L. Potts and R. F.
Smith.
OCT. 1944]
VAN DYKE— OSTOMIDAE
147
NEW SPECIES OF NORTH AMERICAN OSTOMIDAE
(Coleoptera)
BY EDWIN C. VAN DYKE
Calif of-nia Academy of. Sciences, San Francisco
In 1915, I published in the Bulletin of the Brooklyn Ento-
mological Society, descriptions of a number of new species of
the genera N emozoma and Corticotomus and gave a short review
of the known species found north of Mexico. In 1916, in the
same journal, I described another N emozoma, the name of which
had to be modified later because it was found to have been previ-
ously used in the same genus. In 1918, in the Journal of the
New York Entomological Society, Schaeffer contributed an arti-
cle in which he described new species in both N emozoma and
Corticotomus while in 1916, Casey described in his Memoirs, a
closely related genus and made certain remarks concerning Corti-
cotomus. Recently, other undescribed species of both N emozoma
and Corticotomus as well as a related species which will have to
have a new genus erected for it, have come into my hands. These
recent additions I will now describe and inasmuch as there is
still considerable confusion concerning previously described spe-
cies and some new information, make revised synopses of the
two genera.
Nemozoma cupressi Van Dyke, new species
Elongate, about five times as long as broad, cylindrical, moder-
ately shining, black with a rufous patch covering a little more than
a fourth of the base of the elytra yet not reaching the lateral
margins, and the three segments of the antennal club also' some-
what rufous. Head about one-eighth longer than broad, deeply
sulcate anteriorly and with an acute angular emargination in front
with the apices of its lateral boundaries slightly everted, with
distinct and well spaced punctures and a linear groove at the bot-
tom of the sulcus and at the base of the head in line with it;
antennae 11 — segmented, reaching the base of head, segments 2-8
small, 3-8 gradually broader and as a whole subequal in length to
the club the first two segments of which are almost three times as
broad as the eighth segment; eyes moderately large but much flat-
tened as seen from above as they hardly project beyond the side
margin of the head. Prothorax equal to the head in length, nar-
rower at base than elytra, gradually broader toward apex where
about equal in breadth to head and elytra, disk convex, with punc-
148
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
tures similar to those of head, side and basal margin fine. Elytra
about twice as long as broad, sparsely, finely and irregularly punc-
tured except near base and suture where arranged in rows, the
general surface also less shining than head and pronotum, and the
apex with the sutural striae well impressed. Beneath rather finely
and sparsely punctured on head and entire thorax and still more
finely punctured on abodmen. Length, 6.5 mm.; breadth, 2 mm.
Holotype (No. 5426, Mus. Calif. Acad. Sci., Ent.) and one
paratype^ the latter 7.5 mm. in length, both beaten from Sargent
cypress at Occidental, Sonoma County, California, August 1,
1939, by Arthur T. McClay. The paratype is in the collection of
Mr. McClay.
This species is in general a bit larger than N. fissiceps (Fall),
our largest species hitherto described and somewhat similar ex-
cept that the latter has the pronotum an orange red and the legs
as well as base of elytra reddish, and elytral punctures more
regularly arranged. N. attenuatum Van Dyke is smaller and
much narrower, more attenuated, has somewhat rufous legs as
well as 10 segmented antennae; N. punctulata Van Dyke is simi-
larly colored, smaller, has the orange patch at base of elytra
more extensive and the elytral punctures more regular; while
N. schwarzi Schaeffer is very much smaller, has shorter elytra
and orange patches near apices of elytra as well as near the base.
Nemozoma fenyesi Van Dyke, new species
Elongate, not quite five times as long as broad, cylindrical, mod-
erately shining, head, pronotum and narrow basal area of elytra
black, the greater portion of elytra a bright red, and the antennal
club, legs and entire under surface more or less rufous. Head
almost as broad as long, deeply sulcate anteriorly as usual but
with the frontal margin feebly trisinuate, almost transverse, the
median portion narrowly and feebly emarginate, not deeply and
angularly cleft as in our Pacific Coast species, with distinct and
well spaced punctures and the median linear groove not apparent
in the sulcus nor at the base of the head; the antennae 11 — seg-
mented, reaching the base of the head, with segments 2-8 small,
as a whole subsqual in length to the club, the club three-segmented
as usual and very distinctly unilaterally dilated; the eyes moder-
ately large but flattened as in the preceding species. Prothorax equal
to the head in length, just perceptibly narrower at base, the sides
straight and parallel, disk convex, with punctures similar to those
of head but finer and denser, side and basal margins fine but distinct.
Elytra more than two and a half timesi as long as broad, striate,
the sutural stria well impressed and complete, the discal distinct
but the lateral lacking, the strial punctures fine and close together
OCT. 1944]
VAN DYKE— OSTOMIDAE
149
medially, less regular laterally. Beneath rather finely and sparsely
punctured. Length, 7 mm.; breadth, 1.75 mm.
Holotype (No. 5427, Mus. Calif. Acad. Sci., Ent.), a unique
from the Fenyes’ collection of the California Academy of Sci-
ences, collected by Dr. Fenyes at Cordoba, Mexico.
This species is very distinct from any of our species, by its
color pattern and physical characters, particularly the modifica-
tions at the front of the head, and the distinct elytral striation,
and from the Guatemalan species, signatum Sharp, by the color
pattern. Dr. Sharp did not clearly describe the peculiarities of
the head in his species but from the illustration given in the
“Biologia,” I am inclined to believe that fenyesi and signatum
have the front of the head quite similar.
Synoptic Table for the Separation of the American
Species of Nemozoma, Found North of Mexico.
1. Antennae with 10 segments Nemozoma Latr.
Very narrow and about six times as long as broad, head broader
than prothorax or elytra, black with basal third of elytra
orange. Carmel, California, on Monterey pine. Cannon Beach,
Ore., on Sitka spruce. 4-4.5 mm attenuatum Van Dyke
- Antennae with 11 segments Subgenus Monesoma Lev. 2
2. Pronotum, base of elytra, antennae and legs orange or rufous,
the remainder black, from 3-4 times as long as broad. Sierra
Nevada Mts. of California and normally on Lihocedrus, a stray
on other conifers. 5-6 mm fissiceps Fall
— Head and pronotum similarly colored 3
3. Head and pronotum piceous or somewhat rufous, an orange
patch at the base of elytra and another near apex of each
elytron, elytra shorter than abdomen, head narrowly grooved
in front, not broadly sulcate. Arizona and Yuma, California.
3.5 mm schwarzi Schaef.
- Head and pronotum black, base of elytra only orange or red-
dish 4
4. Larger species, head broadly sulcate in front, rufous patch at
base of elytra and not reaching side margins, legs piceous or
unicolorous with body beneath, elytral punctures quite irregu-
lar. Occidental, Sonoma County, California, on Sargent’s cyp-
press. 6.5-7. 5 mm cupressi Van Dyke
- Smaller species, head narrowly sulcate in front, orange patch
at base of elytra fully one-third length of elytra and generally
reaching side margin, legs somewhat rufous, elytral punctures
more or less regularly arranged in rows. Br. Col., Or. and
northeastern California, on pines. 4. 5-5. 5 mm
punctulata Van Dyke
150
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
As indicated in the table, attenuatum is the only species in
our fauna which agrees with the European elongatum L,, the type
of the genus Nemozoma of Latreille, in having ten-segmented
antennae. Its geographical position is also significant in this
connection for it is strictly a member of the Vancouveran insect
fauna, the fauna which has in many ways a close relationship
to the Eurasian fauna. All our other species have eleven-seg-
mented antennae and hence fall into Monesoma of Leveille
which is generally considered, and I think correctly so, to be
but a subgenus of Nemozoma. The bulk of the species in the
subgenus are confined to the New World: western North Amer-
ica, Mexico, Central and South America. Our Pacific Coast spe-
cies are somewhat similar in having the head very deeply sulcate
in front and angularly divergent apically, also with a somewhat
similar color pattern. Our Arizona species, schwarzi, is some-
what different in that it has the head narrowly grooved in front,
the elytra short and quadrimaculate. The Mexican species,
fenyesi, which I have described above, differs from all the species
found north of it in not having the apex of the head distinctly
and triangularly notched but by being almost transverse. The
Guatemalan species, signatum, I believe also approaches it in
this regard.
Genus Corticotomus Sharp
The members of this genus superficially resemble those of
Nemozoma and have quite similar habits, also often have similar
color patterns. They differ by having the head flattened above
or even somewhat concave, with the mandibles directed upward,
not downward as in N emozoma, and as discovered by Casey,
have the elytra with a perforate foramen within the humeri. It
is entirely a New World genus, found in both North and South
America. In the Leng Catalogue, the species are correctly placed
but in the Coleopterorum Catalogus, they are placed partly in
N emozoma and partly in Corticotomus. The genus Parafilumis
Casey with the single species, estriata Casey, is very closely re-
lated to Corticotomus and apparently only differing from cali-
fornicus Van Dyke by having the elytra diffusely punctured, not
with the punctures more or less regularly arranged in rows.
Three of the species of Corticotomus are found on the Pacific
Coast, three on the Atlantic Coast, with one subspecies in south-
ern Texas. Parafilumis is from Oregon.
OCT. 1944]
VAN DYKE— OSTOMIDAE
151
Corticotomus apicalis Van Dyke, new species
Small, elongate, about four times as long as broad, moderately con-
vex above, more or less flattened as seen from the side, shining black,
with antennae, palpi, legs and disk of elytra anteriorly testaceous.
Head almost as broad as long, depressed in front and with a feeble
yet distinct triangular excavation, apical margin transverse, man-
dibles prominent and feebly upturned, punctures well impressed
and dense in frontal impression, more sparse at base and some-
what aciculate at sides; antennae 11-segmented, reaching several
segments behind base of head, segments 2-8 small, gradually
broader outwardly, club well developed, distinctly unilaterally
dilated, segments 9-10 triangular and eleventh elliptical; eyes of
fair size and feebly convex and projecting from the side of head
as seen from above. Prothorax somewhat longer than head, rounded
at base and with sides straight and parallel, side and basal mar-
gins fine but distinct, punctures rather evenly placed and well
spaced. Elytra testaceous over three-fourths of area anteriorly
and piceous at apex, the dark area extending forward along suture
and sides, about one^fourth longer than head and prothorax to-
gether, at least twice as long as broad, feebly striate at sides of
disk, more obscurely so toward suture and side margin, punctures
fine and close in striae, more vague where striae, poorly defined,
intervals flat and with a series of fine punctures, generally close
to the strial punctures giving at times the appearance of double
striation, and a perforate foramen within the humeri. Beneath
finely, sparsely punctured in front, abdomen quite smooth. Length,
3.5 mm.; breadth, 1 mm.
Holotype (No. 5428, Mus. Calif. Acad. Sci., Ent.), from Mt.
Wilson, California, August 5, 1924, in the Blaisdell collection
of the California Academy of Sciences, and two paratypes; An-
derson’s Valley, California, March 12, 1914, from Sugar pine
and Mt. St. Helena, California, March 29, 1931, collected by
myself from the burrows of Cryphalus in Douglas fir. The para-
types have the elytral striae less marked than they are in the type.
The specimens mentioned above, I have had set aside for
sometime, considering that they perhaps belonged to a pygmy
race of caviceps. Upon more critically reexamining them, how-
ever, I find characters which set them aside from that species.
It is not only smaller, but proportionately narrower, more shin-
ing, the prothorax longer than broad, about as broad as long in
caviceps, and with finer and less dense punctuation. The elytra
have a different color pattern and are less convex, in fact the
entire insect is more depressed, with a dorsoventral measurement
much less than in the other.
152
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
Synoptic Table for the Separation of the Species of
CORTICOTOMUS FoUND NORTH OF MeXICO
1. Species bicolored or piceous 2
- Species uniformly castaneous or testaceous, not over 4 mm. in
length - 6
2. The prothorax as broad as long 3
3. Larger species, averaging 5 mm. in length, cylindrical, pro-
thorax rather densely punctured with punctures spaced their
own breadth apart, piceous with rufous antennae and legs and
a rufous patch extending obliquely inward and backward from
base of elytra. Rocky Mountains west to Cascades and Sierra
Nevada Mountains. On pines caviceps (Fall)
- Smaller species, 4.5 mm. in length, depressed or somewhat
flattened, prothorax densely, closely punctured, elytra distinctly
striato-punctate, the serial punctures of intervals also quite
evident, piceous with rufous antennae and legs, also sometimes
head, and a rufous area alofng the sides of the elytra. New
Jersey and adjacent Atlantic States. On pine. .depressu^ Schaef.
4. Distinctly cylindrical with head and pronotum densely, closely
punctured, the strial punctures of elytra quit© evident at least
in front 5
- Somewhat depressed or flattened, head and pronotum black,
shining and with punctures as a rule spaced their own width
apart, elytra testaceous over basal two-thirds, apex piceous.
Calif, on Douglas fir and Sugar pine. 3.5 mm...apicalis new spec.
5. Head, prothorax, antennae and legs rufous, basal third or more
of elytra orange, apical area black. Atlantic States especially
the South, on pines. 3-4 mm cylindricus (Lee.)
- Entire insect above piceous or more or less rufous if somewhat
immature. 3-4 mm cylindricus texanus Schaef.
6. Castaneous or rufous above, legs lighter, head and pronotum
finely, sparsely punctured, elytra evidently striato-punctate in
front, obsoletely so behind. Penn, and other Atlantic States.
3.25 mm parallelus (Melsh.)
- Castaneous or testaceous, punctures of head and prothorax
numerous but spaced their own width apart, elytra with vague
strial and minute, serially arranged punctures over greater
part of disk. Northern Calif, on Ponderosa pine. 3-3.5 mm.
calif ornicus Van Dyke
Pseudocotomus Van Dyke, new genus
Similar to Corticotomus Sharp in most particulars such as the
elongated and cylindrical form, 11-segmented antennae with en-
larged, loose, three-segmented club, slightly upturned mandibles,
serial arrangement of elytral punctures, and short legs but dif-
fering in the following regards : very small size, head convex
above, not flattened or concave, the submentum not distinctly sep-
OCT. 1944]
VAN DYKE— OSTOMIDAE
153
arable from the gular in front, trisinuate anteriorly and with the
outer angles extending forward as a tooth or prominent angular
process beneath and beyond the base of the antennae and partly
overlapping the base of the mandibles, the elytra convex, very
markedly alutaceous, without the perforate foramen within the
humeri first observed by Casey as so evident in Corticotomus, and
the serial punctures very distinctly but very shallowly impressed.
Genotype: Pseudocotomus mclayi Van Dyke.
Pseudocotomus mclayi, Van Dyke, new species
Very small, linear and cylindrical, testaceous, legs and antennae
lighter in color, moderately shining, the micro-reticulation very
evident with good magnification. Head as wide as the prothorax
anteriorly and somewhat shorter, with sides parallel behind the
eyes, convex above, clypeus with a feeble emargination at center,
with deep and well spaced punctures dorsally and laterally; an-
tennae about reaching the base of head, with segments 3-8 small
and hardly broader outwardly, together about equal in length to
the club which is three-segmented, prominent and bilaterally di-
lated; eyes moderate in size, granular, and slightly projecting
outward from the sides of the head as seen from above. Prothorax
about one and a half times as long as wide, rounded at base, with
sides straight and feebly diverging to apex where thorax slightly
broader than elytra, the apical margin arcuate, disk finely, sparsely
punctured with a few minute setae laterally, lateral and basal
margins fine. Elytra about two and a half times as long as broad,
with sides straight and parallel, apex rounded, disk with sutural
striae faintly defined toward apex otherwise without striae but
with punctures serially arranged, well spaced, shallowly impressed
yet quite evident against the alutaceous background. Beneath
slightly darker on after-body and with a few distinct punctures
especially at sides of metathorax and with a scattering of setae.
Length, 2 mm.; breadth, about .75 mm,
Holotype (No. 5429, Mus. Calif. Acad. Sci., Ent.) and eleven
paratypes, all reared from the burrows of a Scolytid, Pseudo-
thysanous hartoni Bruck, in Malvastrom, from Westwood Hills,
Los Angeles County, California, by Arthur T. McClay. The
specimens emerged on various dates as October 22, 1936, May 1,
1937 (Holotype) and June, 1937, Half of the specimens are now
in the McClay collection and half in the collection of the Cali-
fornia Academy of Sciences through the kindness of Mr. McClay,
154
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
PINTALIA STAL WITH SPECIAL REFERENCE TO MEXICO
(Homoptera, Cixiidae)
BY JOHN S. CALDWELL
Circleville, Ohio
Specific placement in this genus is very difl&cult because the
older descriptions were based on color and marking which is
somewhat confusing; in this group. StaP described nine species
from Brazil, Lethierry^ one from Venzuela, Uhler^ two from the
West Indies and Fabricius* * one, Fowler^ four from Mexico and
one from Guatemala, Van Duzee® one from the United States,
MuiU ^enty-seven species and one variety from Brazil, British
Guiana, and Ecuador, Osborn® four from Puerto Rico, and Met-
calf® two from Panama. Muir has stated that consohrina and
ustulata Stal are not typical tentatively leaving forty-nine spe-
cies with discrepans provisionally included in Pintalia. Un-
doubtedly intensive collecting will more than double the number
of known species. Unless stated otherwise all types are in the
author’s collection.
Pintalia dorsovittata Van Duzee (figs. 1, la)
If I interpret this species correctly it has been confused with
delicata Fowler. In a series of twenty specimens from Florida,
South Carolina, Tennessee, and Virginia the elytra are not always
unicolorous with the distinct dorsal stripe. The intensity of color
has a wide range and most of the specimens have a more or less
mottled elytra. Aspersa Dozier^® from Mississippi either belongs
to this species or is new.
Pintalia delicata Fowler (figs. 2, 2a)
Delicata is widely distributed throughout Mexico and it is
1862. Bidr. Rio Janeiro-trakt. Hemip., 2:4-6.
“ 1890. Ann. Soc. Ent. France, (6)10:148.
* 1896. Proc. Zool. Soc. London, pp. 64, and 1901. Proc. Ent. Soc. Washing-
ton, 4 :611.
* 1803. Systema Rhyng., p. 64.
® 1904. Biologia Centrali-Americana, 1 :86-88.
“ 1908. Proc. Acad. Nat. Sci. Philadelphia, p. 491.
’ 1934. Trans. Royal Ent. Soc, London, 82:421-441.
9 1936. New York Acad. Sci., (2)14:198-202.
“ 1938. Bull. Mus. Comp. Zool., Harvard College, 82:290-293.
1926. Mississippi Agric. Exp. Sta. Tech. Bull., 14 :68-69.
OCT. 1944]
CALDWELL— PINTALIA
155
possible that it occurs in the United States. Specimens are pres-
ent from the States of Chiapas, Guerrero, Jalisco, Michoacan,
Morelos, Oxaca, Quintana Roo, Sinola, Sinora, and Vera Cruz in
Mexico, and from Guatemala. The male genitalia will serve to
separate this from any known species. The anal segment is. long
and slightly asymmetrical caudad. The lateral margins of the
pygofers are acutely produced dorsad; the medio-ventral process
is long and rounded apically. The aedeagus does not have many
processes. Fowler’s description of the styles is perfect.
Pintalia lineat^ Caldwell, new species (figs. 3, 3a)
Length, 5.8-6 mm. Vertex, pronotum, and mesonotum with mid-
dorsal yellow stripe that is carried onto the elytra to the apex of
the clavus. Elytra transparent, browned basad and caudad with
smoky areas in between; terminations of veins light.
Anal segment of male with short apex. Lateral margins of
pygofers with small projection dorsad; medio-ventral process
large, rounded apically. Styles widely separated in basal half,
meeting at mid-length then strongly divergent.
Holotype male, Vergel, Chiapas, Mexico, June 3, 1935;
paratype, Mapastepec, Chiapas, Mexico, December 7, 1932
(Dampf) .
Pintalia marginata Caldwell, new species (figs. 4,4a)
Length, male 8 mm., female 9 mm. Elytra transparent, macul-
late with fuscous; maculae tending to be concentrated around
wing margins; apical margin fumate. Veins punctate. Hind wings
evenly smoked.
Lateral margins of male pygofers with blunt projection caudad;
medio-ventral process acute, triangular. Styles widely separated
basad forming a diamond-shaped space; apical three-fourths di-
vergent.
Holotype male, and one paratype from Tierra Blanca, Vera
Cruz, Mexico, August 20, 1932, allotype female, Huatusco,
Vera Cruz, Mexico, September, 1932 (Dampf).
Pintalia stigmata Caldwell, new species (figs. 5, 5a)
Length, 6.8 mm. Head, body and legs yellow-brown. Elytra
hyaline, grayish, slightly fumed apically; stigmal spot brown;
cross-veins smoky. Frons almost twice as long as broad, scarcely
narrowed between the eyes. Hind tibiae with one prominent spine
156
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 4
at mid-lengrth and two smaller spines more basad. Veins of
elytra finely but distinctly g'ranulate.
Anal segment of male long, slender, extreme apex with square,
shallow notch. Lateral margins of py gofers with large projection
basad; medio-ventral process small, acute. Styles slender, lunate,
approximate apically, very pubescent.
Holotype male, from Bartica, British Guiana, June 12, 1901
(H. S. Parish), is in the H. Osborn collection at Columbus, Ohio.
Pintalia fumata Caldwell, new species (figs. 6, 6a)
Length, 8.5 mm. Head and thorax amber with lateral compart-
ments of mesonotum darker. Elytra whitish, hyaline; fuscous spot
present either side of brown stigma and at apex of clavus; basal
and claval areas lightly fiimate; indistinct fuscous maculae pres-
ent basad; extreme apex of elytra fuscous. Hind tibiae quadri-
spinose.
Anal segment of male asymmetrical at apex; right side not pro-
duced as much as left. Lateral margins of pygofers greatly pro-
duced caudad. Styles broadest just before base, finely serrate
basad on inner margins.
Holotype, male, “CosTA Rico, Suizatur’lba, (P. Schild)” is
in the H. Osborn collection at Columbus. Ohio.
Pintalia punctata Caldwell, new species (figs. 7, 7a)
Length, 6.7-8 mm. Head and thorax ferruginous. Elytra trans-
parent, dusty; costal margin dark from stigmal spot to apex; en-
tire apex broadly smoky; granulations on veins broadly fuscous;
entire surface very evenly maculate. Entire insect slender.
Anal segment of male extremely asymmetrical with apex
broadly bilobed on right side. Lateral margins of pygofers with
two processes caudad; medio-ventral process very acute. Styles
slightly enlarged apically, straight in ventral aspect.
Holotype male, and allotype, female, from ZooGOCHi, Oaxaca,
Mexico, June 21, 1935, two male paratypes and one female para-
type from Chiltepec, Oaxaca, Mexico, December 10, 1937,
(Dampf). This species should truly be named maculata but the
name has already been used by both Fowler and Osborn. For
maculata Osborn I propose the name osborni.
Pintalia pseudomaculata Caldwell, new species (figs. 8,8a)
Length, 7.5 nun. Head and thorax dark brown. Elytra mark-
ing similar to panctata except that the stigmal spot is brown and
OCT. 1944]
CALDWELL— PINT ALIA
157
the maculae along the veins tend to run together. The elytra are
much broader, especially apically.
Anal segment of male elongate, extremely asymmetrical api-
cally with the right side forming a broad pendulant lobe. Lateral
margins of pygofers with two large projections caudad; medio-
ventral process broad, triangular. Styles gradually enlarged api-
cally, divergent in caudal half.
Holotype male, from Tamazunchale, San Luis Potosi,
Mexico, November 15, 1938, (Caldwell).
Pintalia chromata Caldwell, new species (figs. 10, 10a)
Length, 7-7.5 mm. Head and pronotum yellow. Mesonotum
darker yellow. Elytra clear, yellowish, especially apically. Veins
sparingly punctate with black; a black spot present at union of
claval veins and at apex of clavus. Anal segment of male asym-
metrical at apex with a short and an elongate lobe on right side.
Lateral margins of pygofers with two hooked projections caudad;
medio- ventral projection acute. Styles gradually enlarged apically.
Holotype male, allotype female, and paratypes from Fortin,
Vera Ckuz, Mexico, October 9, 1941 (De Long, Good, Caldwell
and Plummer) .
Pintalia geometra Caldwell, new species (figs. 9,9a)
Length, 7.5-8. 5 mm. Face black. Vertex and center of prono-
tum brown. Center of mesonotum brown, lateral compartments
black. Elytra fuscous, hyaline, with the following markings clear
to white: a spot basad reaching into claval area but not to sutural
margin, sutural margin yellowish punctate with white, a triangle
with base touching union of claval veins and apex of clavus and
the apex reaching to the radius, the stigmal spot and surrounding
area, the submarginal area sometimes broken, and the extreme
apex sometimes clear.
Anal segment of male short asymmetrical, right side broadly
pendulant. Pygofers with short caudal projection; medio-ventral
process short, rounded. Styles slender.
Holotype male, and two paratypes from Vergel, Chiapas,
Mexico, June 3, 1935, and female allotype from Esmeralda,
Chiapas, Mexico, August 18, 1930 (Dampf).
Pintalia contra Caldwell, new speeies (figs. 11, 11a)
Length, 6 mm. Head and thorax dull orange. Elytra fuscous
with light area in humeral angle, a large spot present in costal
158
THE PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
cell, and a broad stripe across the entire surface cephalad to
stigmal spot. Underwing solid fuscous. Hind tibiae with one strong
spine.
Anal segment of male with right side of apex cut away. Lat-
eral margins of pygofers with small caudal projection; medio-
ventral process large, triangular. Styles strongly divergent in
apical half.
Holotype male, and paratype male, from COROICO, Bolivia,
are in the H. Osborn collection at Columbus, Ohio. Paratype in
writer’s collection.
Pint ALIA aspersa Fowler
(Figs. 12, 12a)
This is a short and compact species with the lateral carinae
of the frons wide and flattened. This latter character is similar
to discrepans Muir which Muir only provisionally places in this
genus. Likewise Fowler believed that aspersa may not belong in
Pintalia. However, from the species and specimens before me
this character appears to be gradational with several species ex-
hibiting carinae with varying degrees of flatness making it im-
possible to draw any definite line between the two groups. Most
of the specimens with flattened carinae have the radius usually
four-branched but this is not consistent. Most of the specimens
before me are darker than the illustrations in the Biologia (pi.
9, fig. 24) with the mid-dorsal stripe less pronounced. The size
ranges between 5 and 6.4 mm. with several females as well as
males only 5 mm. The most spines visible on the hind tibiae
are four with three being more common. The lateral margins of
the male pygofers have a rather long, acute projection caudad
and well dorsad; medio-ventral process is very small. Specimens
from Oaxaca, San Luis Potosi, and Vera Cruz in Mexico, and
two from Guatemala.
Pintalia neoaspersa Caldwell, new species (figs. 13, 13a)
Length, 6-6.5 mm. Color and marking identical to aspersa, the
elytra with possibly more solid colors. Upper third of lateral
carinae of frons similar to that species.
Lateral margins of male pygofers truncate, straight; medio-
ventral process small, triangular. Styles sharply excavate on inner
third of basal margins. Apex of anal segment slender. Aedeagus
with coarsely serrated spur on right side of apical portion; peri-
andrium serrate around joint of aedeagus.
OCT. 1944]
CALDWELL— PINTALIA
159
Figs. 1, Pintalia dorsovittata Van D., lateral view of abdominal
apex of male; la, same, ventral view of half of abdominal apex
of male. Figs. 2, 2a, P. delicata Fowler, same views. Figs. 3, 3a,
P. lineata Caldwell. Figs. 4, 4a, P. marginata Caldwell. Figs. 5,
5a, P. stigmata Caldwell. Figs. 6, 6a, P. fumata Caldwell. Figs. 7,
7a, P. punctata Caldwell. Figs. 8, 8a, P. pseudomaculata Caldwell.
Figs. 9, 9a, P. geometra Caldwell. Figs. 10, 10a, P. chromata Cald-
well. Figs. 11, 11a, P. contra Caldwell. Figs. 12, 12a, P. adspersa
Fowler. Figs. 13, 13a, P. neoaspersa Caldwell. Figs. 14, 14a, P.
acarinata Caldwell.
160
PAN-PACIFIC ENTOMOLOGIST
[VOL. XX, NO. 4
Holotype male, and paratype male, Esmeralda, Chiapas,
Mexico, November 18, 1930, male paratypes Santa Isabel,
Chiapas, Mexico, November 17, 1930, allotype female, and five
female paratypes Santa Isabel, Chiapas, November 17, 1930
(Dampf), and one male paratype Jesus Carranza, Vera Cruz,
Mexico, Oct. 14, 1941 (De Long, Good, Caldwell & Plummer).
Pintalia acarinata Caldwell, new species (figs. 14, 14a)
Length, 6 mm. Head and thorax concolorous with lateral com-
partments of mesonotum darker. Elytra clear to clouded with
brown; a definite white spot present at union of claval veins;
another extending from apex of clavus almost to apex of elytra;
area around stigma clear; a definite fuscous stripe originates at
fork of medius and projects toward apex of costal margin, this
stripe is sometimes broadened to include the entire apex of the
elytra in which case the fuscous is broken by white areas. Apical
terminations of veins thickened, white. Carinae of face similar to
aspersa.
Male styles with inner margins deeply excavate in basal fourth,
contiguous in next fourth, divergent in next fourth, and parallel
in apical fourth. Anal segment relatively short.
Holotype male, and two paratypes from B. Esperanza, Guate-
mala, December 15, 1925 (Dampf).
This species and neoaspersa can be separated from aspersa by
their much more slender appearance, greater length, and distinct
male genitalia.
D. B. MACKIE PERPETUAL MEMORIAL LOAN FUND
The California Entomology Club is sponsoring the establish-
ment of a D. B. Mackie Perpetual Memorial Loan Fund for the
benefit of graduate students in Entomology at the University of
California. The immediate goal is $1,000.00 of which more than
$750.00 has already been subscribed. Mr. Mackie’s entomological
work is an especially suitable memorial for such a fund since he
so greatly benefited California Agriculture through his pioneer
work in vacuum fumigation, both with cyanide and methyl bro-
mide. Contributions may be sent to H. H. Keifer, Secretary-
Treasurer, California Entomology Club, State Office Building
No. 1, Sacramento, California.
OCT. 1944]
INDEX TO VOLUME XX*
161
Acarina, 7.
Acmaeodera acuta, 105.
angelica, 106.
bishopiana, 106.
connexa, 106.
fenyesi, 106.
gemina, 107.
hepburni, 107.
labyrinthica, 107.
mariposa, 108.
plagiaticauda, 108.
purschiae, 108.
sinuata sexnotata, 109.
vandykei, 109.
variegata, 109.
Adamson, A. M., Trinidad
termites, 39.
Aelothrips kuwanii, 84.
Alexander, C. P., Tipulidae, 91.
Alfalfa insects, 37.
Aglais antiope, 146.
Altinote alcione bassleri, 32.
a. hermosa, 33.
a. leucothoe, 33.
a. mengeli, 32.
a. moneta, 32.
carmentis, 31.
ozomene alicunde, 34.
stratonice albunea, 34.
Amobia floridensis, 118.
Anaphothrips longipennis, 86.
zeae, 86.
Andrena angustitarsata, 67.
auricoma, 67.
carliniformis, 67.
saccata, 67.
Anoplotermes, 40.
Anthobium bonelli, 103.
californicum, 101.
farrarae, 101.
grayae, 102.
minskae, 104.
orientale, lOl.
Aradidae, 111.
Aradus ampliatus, 112.
apicalis, 113.
blaisdelli, 114.
behrensi, 113.
cinnamoneus, 114.
compressus, 114.
debilis, 114.
depictus, 112.
fuscomaculatus, 112.
patibulus, 112.
Asilidae, 67.
Atimia dorsalis, 30.
Bailey, S. F., Thysanoptera, 81.
Beer, F. M., Acmaeodera, 105.
Bohart, R. M., Stenodynerus, 69
Boletobius cascadensis, 27.
c. hatchi, 28.
intrusus wallisi, 29.
kelleyi, 26.
kremeri, 23.
ruber, 24.
Book review, 100.
Buprestidae, 35, 53, 105.
Buprestis aurulenta, 35, 53.
Caldwell, J. S., Pintalia, 154.
Californiulus vicinus, 80.
Caliulus furcifer, 80.
pcarcei, 80.
Callinicus calcaneus, 67.
Camras, S., Zodion, 121.
Cantharis dentatus, 20.
divisus, 21.
lautus, 21.
Ceononycha ampla, 115.
bowlesi, 115.
testacea, 115.
Chamberlin, R. V., Myriopods,
79.
Chilopoda, 79.
Chirothrips mexicanus, 82.
Cicindellidae, 76.
Cicindela nevadica tubensis, 77.
theatina, 76.
Cicadellidae, 41, 98, 129.
Citellus, host of Thrassis, 14,
18, 19.
Cixidae, 154.
Cladura m^xnabi, 95.
Clavigeridae, 77.
Cleridae, 54.
Cloanthanus dilatus, 129.
fasciatus, 130.
hambletoni, 129.
Clover leaf weevil, 120.
Coccinella ampla var. rufa, 146.
Coccinellidae, 144.
Coleoptera, 12, 20, 22, 23, 30,
35, 39, 53, 54, 61, 76, 101,
105, 115, 120, 144, 147.
Coleoptera, dispersal flights, 113.
Conopidae, 121.
Coproporus ventriculus, 30.
Corticotomus, 132.
apicalis, 151.
californicus, 152.
caviceps, 152.
cylindricus, 152.
c. texanus, 152.
depressus, 152.
parallelus, 152.
Ctenothrips reticulatus, 86.
Culicidae, 19.
Cynomys, host of Thrassis, 19.
Dactylolabis postiana, 94.
* New names in bold face type, synonyms in italics.
162
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. XX, NO. 4
Dasymutilla aureola pacifica, 60
DeLong, D. M., Cloanthanus, 129.
Hebenarus, 41,
Premanus, 98.
Dineutus landaisi latilimbus, 22.
nuirginatits, 22.
mellyi, 22.
or i entails, 22.
Diplopoda, 80.
Dipodomys, host of Thrassis, 15,
16.
Diptera, 19, 67, 91, 118, 121.
Dolichovespula arenaria, 80.
Duncan, C. D., Rosalia funebris,
39.
Elaphrothrips angusticeps, 90.
Ethopolys pusio, 79.
Fender, D. M., Cantharis, 20.
Frankliniella cephalica, 83.
insularis, 82.
Fustiger knausi, 77.
Gonomyia brookmani, 96.
Gosibius benespinosus, 79.
Gynaikothrips uzeli, 83.
Gyrinus orientalis, 22.
Haplothrips fasciculatus, 82.
gowdeyi, 83.
Hatch, M. H., Anthobium, 101.
Book Review, 100.
Hebenarus, 41.
abruptus, 44.
altus, 44.
arcuatus, 48.
cornutus, 47.
curtus, 47.
huachucus, 49.
luteus, 52.
maculosus, 51.
marginellus, 48.
moiiticalus, 50.
pallidus, 42.
ritanus, 42.
r. rubralineus, 43.
rubranotus, 46.
spinosus, 52.
Hemiptera, 78, 111.
Hemming, F., Nomenclature
Commission, 119.
Heterothrips decacornis, 84.
Homoptera, 41, 98, 129, 154.
Hoplandothrips raptor, 88.
Hymenoptera, 39, 60, 69, 80, 132.
Hypera punctata, 120.
Hyperaspidius floribunda, 144.
juniperus, 145.
rossi, 145.
shauli, 145.
walcotti, 144.
Hyperaspi falli, 144.
International Commission on
Zoological Nomenclature, 119.
Isoptera, 39.
Ixodes neotomae Cooley, 7.
Keifer, H. H., Applied taxo-
nomy, 1.
Leech, H. B., Atimia dorsalis, 30
Gyrinidae, 22.
Lepidoptera, 31, 117, 146.
Leptothrips mali, 83.
Limonia (Dicranomyia) illus-
tris, 91.
Linsley, E. G., Amobia flori-
densis, 118.
Aradidae, 111.
Buprestis aurulenta, 35, 53.
Callinicus calcaneus, 67.
Stored products insects, 35.
Liothrips mexicanus, 89.
Lophobius castellopes, 79.
Lygus bradleyi, 78.
convexicollis, 78.
distinguendus, 78.
d. tahoensis, 78.
?nubilatus, 78.
perplexus, 78.
stratus, 78.
usingeri, 78.
Mackie Loan Fund, 160.
Malkin, B., Boletobius, 23.
Mansfield, G. S., Ceononycha,
115.
Mansonia perturbans, 19.
Megachile alamosana, 142, 143.
alata, 135.
anograe, 142, 143.
astragali, 143.
bradleyi, 135.
brimleyi, 136.
casadae, 136.
coloradensis, 136.
dacotensis, 136.
frigida, 135.
fucata, 136.
gravita, 142, 143.
histrata, 136.
hookeri, 137.
impartita, 137.
integra, 137.
laurita, 142.
legalis, 137.
maurata, 137, 143.
melanderi, 140, 143.
mojavensis, 138.
mucorosa, 138.
nebraskana, 132.
oslari, 138.
palmensis, 138.
parallela, 132.
pascoensis, 142, 143.
soledadensis, 138.
sp., 39.
subanograe, 139, 143.
OCT. 1944]
INDEX
163
Megachile (Cont.)
subnigra, 138.
s. angelica, 138.
subparallela, 132.
verbesinae, 132.
xerophila, 142, 143.
yumensis, 141, 143.
Megachilidae, 132.
Megaloptera, 110.
Mezira pacifica, 114.
reducta, 114.
Michelbacher, A. E., Hypera
punctata, 120.
Miridae, 78.
Mischocyttarus f la vi tarsus, 80.
Monesoma, 149.
Mordella lottini, 61.
salomonensis, 61.
subaenea, 61.
Mordellistena abrupta, 65.
maxillaris, 64.
oceanica, 66.
Nasutitermes, 39.
Nemozoma, 131.
attenuatum, 149.
cupressi, 147.
fenyesi, 148.
fissiceps, 149.
punctulata, 149.
schwarzi, 149.
Neohermes nigrinus, 110.
Neotoma, host of Ixodes, 7.
host of Thrassis, 14, 15, 19.
Nomenclature, 119.
Nunenmacher, F. W., Coccinel-
lidae, 144.
Zacotus fredericki, 12.
Onychomys, host of Thrassis,
15, 16, 19.
Orectochilus nigroaeneus, 22.
melli, 22.
Osmia brevis, 67.
cara, 67.
densa pogonigera, 67.
lignaria, 67.
Ostomidae, 147.
Otocryptops sexspinosus, 79.
Oxydiscus (O.) pacificus, 92.
Pacific Coast Entomological
Society, Proceedings, 35.
Peromyscus, host of Thrassis,
14, 16, 19.
Phoebis sennae marcellina, 117.
Pieridae, 117.
Pintalia acarinata, 160.
aspersa, 158.
chromata, 157.
contra, 158.
delicata, 155.
dorsovittata, 154.
Pintalia ( Cont. )
fumata, 156.
geometra, 157.
lineata, 155.
marginata, 155.
neoaspersa, 158.
pseudomaculata, 157.
punctata, 156.
stigmata, 155.
Pokabius pitophilus, 79.
Potts, R. W. L., Acraeinae, 31
Aglais antiope, 146.
Dasymutilla aureola, 60.
Premanus, 98.
hebatus, 98.
Prendergast, B., Megachile, 39.
Prince, F. M., Syl vatic plague,
39.
Thrassis, 13.
Pseudocorticotomus, 152.
mclayi, 153.
Ray, E., Mordellidae, 61.
Rees, D. M., Mansonia, 19.
Rosalia funebris, 39.
Rotger, B., Cicindela, 76.
Sarcophagidae, 118.
Scaphytopuis ritanus, 42.
Scarahaeidae, 115.
Sceliphron servillei, 60, 118.
Siphonaptera, 13.
Smith, R. F., Aglais antiope,
146 .
alfalfa insects, 37.
Dasymutilla aureola, 60.
Hypera punctata, 120.
Mischocyttarus, 80.
Staphylinidae, 23, 101.
Stenodynerus anatropus, 71.
claremontensis, 70, 72.
foxensis, 73.
fundatiformis, 71, 74.
f. gonosceles, 74.
fundatus, 70, 76.
microstictus, 70, 72.
ochrogonius, 74.
papagorum papagorum, 70,
71.
p. tinctifer, 71.
phoenixensis, 70, 72.
taos, 70, 73.
xanthianus, 70, 74.
Stored products insects, 35.
Sylvatic plague, 39.
Tenuaris, 52.
Termites, Trinidad, 39.
Thrassis aridis, 14.
bacchi, 18.
campestris, 15.
pansus, 18.
setosis, 13.
164
THE PAN-PACIFIC ENTOMOLOGIST [vOL, XX, NO. 4
Thysanoptera, 81.
Tilden, J. W., Ceononycha, 115.
Phoebis sennae marcellina,
117.
Tomotaenia laevipes, 79.
Trichodes apivorus, 55.
a. borealis, 55.
basalis, 55, 56.
bibalteatus, 55, 58.
bicinctus, 55, 56.
bimaculatus, 55.
borni, 55, 56.
nexus, 55.
nutalli, 55, 60.
oresterus, 54.
ornatus, 55, 58.
o. obsoletus, 59.
o. tenellus, 55.
peninsularis, 55.
simulator, 55, 58.
Tumeus irrorellus, 131.
Unbidden House Guests, re-
view, 100.
Usinger, R. L., Aradidae, 111.
Lygus, 78.
Van Dyke, E. C., Neobermes,
110 .
Ostomidae, 147.
Type localities, 37.
Vespidae, 69.
Wolcott, A B., Tricbodes, 54.
Zacotus fredericki, 12.
Zapus, bost of Thrassis, 14.
Zenora tricolor, 144.
Zodion abditus, 123, 128.
abdominale, 122.
albonotatum, 127.
americanum, 125, 128.
basalis, 123.
bicolor, 124.
bilinmtwm, 122.
cinereiventre, 124, 128.
cyanescens, 127.
fulvifrons, 121, 128.
hirtipes, 125.
intermedium, 123, 128.
lativentre, 122.
lovetti, 128.
nanellum, 125.
nigrifrons, 125, 128.
obliquefasciatum, 127, 128.
occidentale, 123.
palpale, 126, 128.
parvum, 126, 128.
perlongum, 123, 128.
pictulum, 127.
pygmaeum, 125.
reclusum, 122.
scapulare, 126, 128.
triste, 128.
THE
The Pan-Pacific Entomologis
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
VOLUME TWENTY
1944
EDITORIAL BOARD
E. G. LINSLEY, Editor
E. C. VAN DYKE, Associate Editor
R. W. L. POTTS, Assistant Editor
G. F. FERRIS, E. S. ROSS, R. L. USINGER
R. C. MILLER, Treasurer
1944
G. F. Ferris
E. O. Essig
PUBLICATION COMMITTEE
E. O. Essig, Chairman
1945
M. A. Stewart
F. E. Blaisdell
1946
C. D. Duncan
H. H. Keifer
San Francisco, California
1944
11
CONTENTS FOR VOLUME XX
Alexander, C. P,
Undescribed species of Tipulidae from the western United
States 91
Bailey, S, F.
A review of D. L. Crawford’s species of Thysanoptera.... 81
Beer, F. M.
Notes on the hosts, habits and distribution of Oregon
Acmaeodera 105
Bohart, R. M.
Stenodynerus fundatus and related species in North
America 69
Caldwell, J. S.
Pintalia StM with special reference to Mexico 154
Camras, S.
Notes on the North American species of the Zodion fulvi-
frons group 121
Chamberlin, R. V.
Some records of myriopods collected by W. M. Pearce in
California 79
Cooley, R. A.
Ixodes neotomae, a new species from California 7
De Long, D, W.
A new genus — Hebenarus — and thirteen new species of
leafhoppers closely related to Cloanthanus 41
A new genus — Premanus — and species of Mexican leaf-
hopper related to Omanana. 98
Four new species of Cloanthanus and Tumeus from Mex-
ico and Brazil 129
Fender, D. M.
Notes on Cantharis, II 20
Hatch, M. H.
Book review: Unbidden House Guests 100
Studies on North American Staphylinidae I : Anthobium.. 101
Hemming, F.
Recent work by the International Commission on Zoo-
logical Nomenclature 119
Keifer, H. H.
Applied entomological taxonomy 1
Ill
Leech, H. B.
List of six species of gyrinid water beetles collected by
J. L. Gressitt in China and Japan 22
Spring flight of Atimia dorsalis... 30
Linsley, E. G.
Larval longevity in Buprestis aurulenta Linn 53
Prey of the robber fly Callinicus calcaneus Loew 67
A brief note on the habits of Amobia (Pachyophthalmus)
floridensis Townsend 118
Linsley, E. G. and R. L. Usinger
Further notes on the habits of some flat bugs with a de-
scription of the male of Aradus patibulus Van Duzee 111
Malkin, B.
New species of Boletobius from the northwest with a dis-
tributional note on Coproporus 23
Mitchell, T. B.
New species and records in Megachile 132
Nunenbacher, F. W.
A new species of Zacotus 12
Studies among the Coccinellidae — No. 9 144
Pacific Coast Entomological Society
Proceedings for 1943 35
Potts, R. W. L.
Systematic notes concerning American Acraeinae, II 31
Potts, R. W. L. and R. F. Smith
Hibernation of Dasymutilla aureola pacifica 60
An aberrant specimen of Aglais antiopa (Linn.) 146
Prince, F. M.
Descriptions of three new species of Thrassis Jordan and
the females of T. bacchi (Roths.) and T. pansus
(Jordan) 13
Ray, E.
Notes and descriptions of Mordellidae from the Solomon
Islands 61
Rees, D. M.
A new mosquito record from Utah 19
Rotger, B.
A new species of Cicindela and two new records of Cole-
optera 76
IV
Smith, R. F.
A note on the hibernation of Mischocyttarus flavitarsus
in a nest of Dolichovespula arenaria 80
Smith, R. F. and A. E, Michelbacher
Clover leaf weevil in California 120
Tilden, J. W.
Breeding of Phoebis sennae marcellina Cram, in San
Jose, California 117
Tilden, J. W. and G. S. Mansfield
Notes on three species of the genus Ceononycha Horn 115
Usinger, R. L.
Host plant records of western Lygus 78
Van Dyke, E. C.
A new dobsonfly (Megaloptera) from California 110
New species of North American Ostomidae 147
Wolcott, A. B.
The American species of Trichodes 54
MAILING DATES FOR VOLUME XX
No. 1. January 31, 1944.
No. 2. April 29, 1944.
No. 3. July 31, 1944.
No. 4. October 30, 1944.
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