Vol. II July, 1925 No. 1
THE
Pan-Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
HAMLIN, IMPORTANT OPUNTIA INSECTS OF THE UNITED STATES .... 1
BARNES AND BENJAMIN, NEW AND RARE LEPIDOPTERA FROM TFIE SOUTHWEST 12
BARNES AND BENJAMIN, NOTES ON ARIZONA PHALAENID^ 16
OSBORN, A NEW ACINOPTERUS FROM SOUTHERN CALIFORNIA 22
CHAMBERLIN, THE CONIFEROUS TREES OF THE UNITED STATES WITH THE
SCOLYTID BEETLES SAID TO ATTACK THEM 23
VAN DUZEE, A NEW MIRID FROM ARIZONA 35
ROCKWOOD, ON NIGHT FLYING AND ATTRACTION TO LIGHT IN ACRIDIID^ . . 36
HUBBELL, A NEW SPECIES OF PRISTOCEUTHOPHILUS 39
SHANNON, NORTH AMERICAN SPHECOMYIA 4?
ECONOMIC NOTES 45
EDITORIAL COMMENT 47
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1925
THE PAN-PACIFIC ENTOMOLOGIST
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The Pan-Pacific Entomologist
Vol. II, No. 1.
July, 1925
IMPORTANT OPUNTIA INSECTS OF THE UNITED
STATES: BIOLOGICAL NOTES ON THE
MOTH-BORERS (PYRALID^).
BY JOHN C. HAMLIN
Fresno, California
In 1920 the project for the biological control of introduced
Opuntias was formally undertaken by the Commonwealth
Prickly-pear Board. This project contemplated the intro-
duction to Australia of the natural enemies of prickly-pear
occurring in the Americas. During the writer’s connection
with this work the collecting and rearing of large numbers
of the more important cactus insects afforded an oppor-
tunity for a general study of their biologies. The necessity
of shipping the material intermittently, as well as attention
to the bulk rearings, prevented undertaking more detailed
biological work.
Because of the economic potentialities of these insects
it is thought desirable to record the data secured. The
results herein contained represent, for the most part, aver-
age conditions observed in breeding stocks and in the field
over several 3 ^ears. The present paper deals only with the
more destructive representatives of that group of Opuntia
insects attacking the joints internally.
Melitaea prodenialis Walker
DESCRIPTIVE
Adult. The adult is a large-bodied, sluggish, gray moth
with a wing expanse of slightly more than one inch. The
fore wings are marked with irregular transverse vittae.
Eggs. The eggs are laid one upon another with great
precision. The egg-sticks are normally attached to the
extreme tips of spines or spinules on the upper part of the
plant, being difficult in the field to distinguish from spines.
Sticks contain from 20 to 60 eggs, averaging 35. The sticks
are usually slightly curved but are often partly coiled. The
2
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
individual eggs are so placed that they resemble tiny,
thickened discs. They are elliptical, one diameter being
1.0 mm. and the other, 0.9 mm. In length the egg measures
0.5 mm.
Larva. Until about one-fourth grown the larva has a
light brown color. Thereafter it assumes a rich, dark blue
hue. When full-grown it measures from 30 to 40 mm. in
length.
Pupa. The dark brown pupa is enclosed in a very tough
silken cocoon, often pure white. The cocoon is somewhat
less than one inch in length.
LIFE HISTORY
In summer the eggs hatch in from nine to fifteen days. The
small larvae construct a light web about an areole, or where
two joints make contact, and beneath this they feed gregari-
ously. The web quickly becomes strengthened by the
addition of excrement and the dried juices of the plant.
When about one-fourth grown they enter the joint and
continue to feed together, emitting pellets through one or
more openings in the sides of the joint. The change of color
described above transpires shortly after the larvae enter the
interior of the plant. Normally, they remain within the
joints until full-grown. In summer the larval stage extends
over from six to eight weeks.
Full-grown larvae emerge from the joints and spin their
cocoons near the affected plant. The cocoons are usually
placed in clumps of three or four, as many as fifteen having
been taken in one mass. Sometimes they are found beneath
fallen, dried joints and near-by rubbish or placed at the
base of an adjacent grass clump. In these instances the
cocoons are a beautiful, lustrous white. The majority, how-
ever, seem to construct their cocoons in the sand just at the
base of the plant, and, in these cases, the cocoons have a
considerable admixture of sand and other foreign matter.
The duration of the pupal condition in summer is approxi-
mately one month.
SEASONAL CYCLE
There are three generations annually in the region about
Miami, Florida. Adults from the overwintering larvae
JULY, 1925 ] HAMLIN OPUNTIA INSECTS
3
appear during late February and early March. The mid-
summer adults issue in June. The fall brood of moths
emerges in October. There are not, however, such definite
generations as would be indicated by the above periods.
The dates mentioned show the periods of maximum
emergence. It is possible to find in the field at almost any
season of the year cocoons and all sizes of larvae.
HABITS
The adults hang motionless from spines during the day.
Mating and oviposition occur at night. This insect is much
more exacting than Olyca junctolineella as regards conditions
requisite for oviposition in cages. Moisture seems to be a
most decisive factor. During periods of low relative
humidity in the natural range and under normally dry
conditions elsewhere, it has been found necessary to sprinkle
the cloth of the cages nightly in order to secure more than
a few scattered, subnormal egg-sticks.
The gregarious habit of M. prodenialis larvae is the
particular feature of this species which makes it so much
more destructive to Opuntia than its western relative,
0. junctolineella. The average number of larvae found to-
gether is about twelve, but as many as twenty-six half-
groAvn larvae have been taken in a single small joint.
FOOD PLANTS
Melitara prodenialis has been found infesting all of the
more common prickly-pears of Florida ; namely, Opuntia
austrina, 0. drummondii, 0. lata, and 0. keyensis. These
Opuntias are comparatively tender and succulent. Rearing
experiments showed that this insect will not survive on
the tougher western Opuntias such as O. Imdheimeri and
O. engelmanni. In Queensland, laboratory results showed
that the larvae did not prosper on the “common pest pear,”
O. inermis, its texture being somewhat similar to that of the
Texas species mentioned. The species of second importance
in Queensland, the “Gayndah pear” or “spiny pest pear”
(O. stricta), being similar texturally to the Florida species,
promises to be quite a suitable host for this insect.
4
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
INJURY TO PLANT
The entire damage due to M. prodenialis is accomplished
by the larvae. They enter the upper joints and, always
feeding gregariously, continue downward from joint to
joint, finally invading even the subterranean portions of the
plant. Infested plants are rapidly hollowed out, and the
remaining tissue, together with a certain amount of larval
excrement, is quickly oviposited in by certain scavenging
diptera (Volucella esuriens Fabr., V. fasciata Macq., and
Neorondania sp.). The larvae of these scavengers are often
responsible for considerable extension and aggravation of
injury to large plants not completely killed by the primary
insect. The effectiveness in Florida of this combination of
insects in destroying prickly-pear is abundantly indicated
in the field by disheveled lots of dried, whitened, paper-like
epidermis on the ground, marking the complete destruction
of cactus clumps.
DISTRIBUTION
Melitara prodenialis has been recorded ^ from New Jersey,
Mississippi, and Florida. In the latter State the writer has
taken it at Miami, Cocoanut Grove, Larkin, West Palm Beach,
Palatka, and Gainesville, and on Biscayne Key.
Thus, while the larval habits render this species the most
destructive Opuntia insect in the United States, its climatic
limitations and the different texture of the western prickly-
pears preclude its prospering in our main cactus area.
NATURAL ENEMIES
Undoubtedly the greatest natural handicap to M.
prodenialis in its habitat obtains through the attack of ants
upon full-grown larvae after emerging from the plant for
pupation in or near the ground.
Brachymeria (Chalcis) pedalis (Cresson) is the only para-
site observed. The parasitized host larva spins its cocoon
normally and completes its transformation. A single adult
parasite then emerges from the pupa. This parasite is not
at all abundant in Florida and constitutes little or no check
upon the moth-borer.
1 The Principal Cactus Insects of the United States, by Hunter,
W. D., Pratt, P. C. & Mitchell, J. D., U. S. D. A., Bur. Ent. Bull.
113 , 1912 .
JULY, 1925]
HAMLIN OPUNTIA INSECTS
5
Melitara sp.
Another type of moth-borer was taken in the same
pasture at Uvalde, Texas, on three occasions during the
summers of 1921 and 1922. Doctor H. G. Dyar, having the
opportunity of viewing only a very small number of adults,
and in the absence of data upon the immature stages, tenta-
tively indicated its identity as Melitara prodenialis Walk.^
Observations on the appearance and habits of the immature
stages suggest that this form is perhaps distinct from that
species.
The three-fourths grown larvae are indistinctly banded,
the blue being very pale, and a faint buff suffusion is notice-
able over the entire larva. In each instance of collection,
from twenty-one to twenty-four larvae were found living
gregariously within the heavy basal joints, excreting
through several perforations. The dry pellets were some-
what webbed and hung from the holes in strings from one
to three inches long.
Pupation takes place within the chamber excavated by
the larvae. The cocoon is very similar to that of M. pro-
denialis, being slightly larger. Pupation occurred during
September. ^
The adults emerged late in October and lived about one
month. The first eggs were laid on November 2. They
are deposited in the same fashion as are those of
M. prodenialis and O. junctolineella, but the sticks average
many more eggs. All egg-sticks contained from sixty to
sixty-five eggs each. The incubation period was approxi-
mately four weeks.
A chalcid parasite which emerged from the cocoons was
determined by Mr. S. A. Rohwer as Pareniaca schwarzi
Cwfd.
Unfortunately no further data were secured to establish
the identity of this moth-borer. One portion of the larvae
which was shipped to the writer in Australia perished, and
the remainder was also lost at Uvalde, Texas, when Mr. E.
2 1 have been informed that additional material of this form has
been submitted to Doctor Dyar and he is describing- it as a new species.
3 The further notes on this moth-borer were made by Mr. E.
Mortensen.
6
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
Mortensen changed headquarters to Miami, Florida, for the
winter.
Olyca (Melitara) junctolineella (Hulst)
DESCRIPTIVE
Adult. The adult is a grayish moth with a wing expanse
of one and one-half inches, being larger and more robust
than Melitara prodenialis.
Eggs. The eggs are somewhat larger than those of
M. prodenialis but are arranged in precisely the same manner.
The egg-sticks average about twenty-five eggs each. Some
sticks have been seen which contained as many as fifty eggs.
Larva. The newly hatched larva is about 7 mm. long and
of a dark color with a suggestion of a deep blue hue. When
from one-half to two-thirds grown they become banded,
blue and white. The full-grown larva measures from 40 to
50 mm. in length and has a diameter about equal to that of
the ordinary lead-pencil.
Pupa. The cocoon and pupa closely resemble those of
M. prodenialis , but are considerably larger.
LIFE HISTORY
The preoviposition period is very brief. Instances were
commonly observed where the moths emerged, mated and
oviposited within twenty-four hours.
The eggs of the spring brood hatch in from five to twelve
days. The small larvae immediately burrow into the joint,
several often entering the same hole. The joint so affected
rapidly blackens and falls to the ground in a few days. The
larvae then crawl again upon the plant and enter new joints,
thus becoming well distributed over the cactus clump.
While the very small larvae live for a brief period in the
same joint, they have separate tunnels and are in no sense
gregarious. After the early stages the larvae are quite
solitary, it being rare to find more than one to a joint.
Occasionally two have been found infesting the same joint.
After adopting the solitary habit the larva remains con-
stantly inside the joint throughout its development. During
summer some ten to twelve weeks are passed in the larval
condition. When full-grown the larva cuts an exit hole and
JULY, 1925] HAMLIN OPUNTIA INSECTS
7
crawls to the ground. The cocoon is spun beneath fallen
cactus joints or other debris near the base of the plant, or
may be placed just below the surface of the soil. The
appearance of the cocoon is affected by its location, those
spun on the joints being a beautiful white, while those in
the ground are mixed with soil. Transformation takes
place shortly after completion of the cocoon, and the pupal
stage extends over about four weeks. The length of all
stages of the fall generation is somewhat longer.
SEASONAL CYCLE
Throughout Southern Texas, 0. junctolineella has two
generations yearly. Thus, during three-fourths of the year
the species exists in the larval stage, both the winter and
the dry, hot summer being passed in that condition. The
periods of emergence and oviposition coincide with the
normally rainy months of the spring and fall. At Uvalde,
Texas, the spring brood of moths issues during May and
early June, the maximum emergence obtaining during late
May or early June. These moths produce the summer
larv^ which pupate from late August through September.
The peak of emergence of the fall moths occurs about the
first of October, and these give rise to the overwintering
larvae. The winter larvae require about five and one-half
months to become full-grown. Pupation extends over the
latter half of March and through April.
The upsetting of the seasonal cycle by transportation
into the opposite seasons of Queensland and the promise
of becoming a three-generation species there has been
recently recorded. ^
HABITS
The adults are inactive by day. The usual position is to
hang from a spine by the second and third pairs of legs,
with the front legs folded close against the head, resembling
the palps. Mating and oviposition occur during the night
although some few pairs have been noted to remain in
copulation during the following forenoon. Many attempts
to observe the manner of oviposition were futile.
4 Seasonal Adaptation of a Northern Hemisphere Insect to the
Southern Hemisphere, by John C. Hamlin, Jl. E'c. Ent., V. 16, No. 5,
pp. 420-424, 1923.
I
8
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
The larvae are quite solitary. Usually only a small num-
ber are found in a cactus clump, despite the large number
and arrangement of the eggs. Cannibalism, if it occurs at
all, is not of sufficient prevalence to account for this condi-
tion. In the rearing of huge numbers of individuals, it was
necessary to place together large numbers of the larvae
from the moment of hatching until pupation occurred, thus
offering an opportunity for the display of cannibalistic
tendencies rarely, if ever, obtaining in nature. Despite such
conditions there were no indications of cannibalism. The
explanation of the condition cited probably lies in the
activities of ants as shown elsewhere in this paper.
The rapidity with which a half-grown larva can enter a
fresh, solid joint is one of the most remarkable feats of the
species. The silicious cuticle is quickly punctured and the
larva disappears in an incredibly short time, considering
the density of the medium. This is accomplished by the
larva passing the pulp immediately through its body.
FOOD PLANTS
During the entire period of the writer’s observations the
species has been found feeding only in prickly-pear. The
favored species are Opuntia Undheimeri and O. engelmanni.
In Queensland the species has prospered, under laboratory
conditions, on both 0 . inermis and 0. stricta.
Prickly-pear plants set up a strong reaction to wounds of
any kind. Numbers of instances have been observed in
rearing cages where the exuding juices have drowned the
larvae which punctured the epidermis. This condition is
much more frequent in cages where, because of crowding,
the larvae are forced to enter new joints many more times
during their existence than in the field. Nevertheless, many
larvae perish in this manner under natural conditions. The
prickly-pears of the ficus-indiccc group show a greater degree
of this type of resistance to larvae of 0 . junctolineella than
do the other Opuntias.
CHARACTER OF INJURY
Newly hatched larvae are responsible for the outright
killing of many joints. The summer brood of larvae, appear-
ing simultaneously with the spring growth of the plant, is
JULY, 1925]
HAMLIN OPUNTIA INSECTS
9
responsible for the destruction of large numbers of the
young fruits.
The most reliable indication of injury is the characteristic
swelling of the joints in which larvae are developing. The
larva hollows out a chamber and packs excrement in the
lower part of the cavity. The tissue within is completely
destroyed and the proliferation of such parts increases the
thickness from two to four-fold. When full-grown the
larva cuts an exit hole through one side of the swelling
and leaves the joint to pupate near the base of the plant.
The larval exit hole permits the several scavenging flies
(V olucella esuriens Fabricius, V. avida Osten Sacken,
V. fas data Macquart, V. pusilla Macquart, Copestyhim mar-
ginatum Say, Hermetia illucens Linn., H. chrysopila Loew, and
H. hunteri Coquillett) to oviposit within the swollen portion.
The maggots find ideal conditions in the excrement of the
former inhabitant and the moisture from the injured plant
tissues. These maggots are, during their development, able
to extend the damage so that further healthy areas of the
joint are invaded. In those instances in which exit holes
were placed high enough to prevent the draining away
of the resulting liquefaction, this secondary injury may be
extended into the next joint below. The net result is the
death of joints so affected.
Occasionally a swollen joint is found which contains no
larva and which is without an exit hole. In view of the
plant resistance already noted, it is believed that such
missing larvse were overcome by the plant reaction after
the cavity had been started. In such joints the swelling
diminishes greatly but the sides do not reunite.
DISTRIBUTION
Olyca junctolineclla occurs throughout the cactus area in
Texas. During the writer’s observations it has been found
most abundant in the Uvalde, Texas, region. It has also
been taken at many points in Mexico as far south as
Cuernavaca, State of Morelos. The writer observed one
larva in an Opuntia growing on the floor of the Grand
Canyon in Arizona which appeared to be of this species.
10
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
NATURAL ENEMIES. PARASITES
A number of parasitic insects appeared in rearing cages
containing field-collected material, but only a portion of
them may definitely be listed as parasites of this moth-
borer.
Apanteles miinoristcB Muesebeck is perhaps the most im-
portant. The parasitized host larva attains full size and
spins its cocoon normally, except perhaps for being of
lighter construction. The host larva is destroyed just
before pupation, and shortly the Olyca cocoon is found to
contain large numbers of the Apanteles cocoons. The
number of adult parasites emerging from one host cocoon
was found to vary between twenty-five and ninety.
On one occasion observations were made upon the
happenings in small fruit jars to which had been added
Olyca larvae, Apanteles adults and a cactus joint. The
parasites would not ovipost in the newly hatched larvae.
Their constant reaction on touching these small larvae dur-
ing their rapid crawling about the jar was merely indicative
of increased excitement and hurried avoidance. When
placed with Olyca larvae about one week old, the parasites
oviposted deeply within the larvae immediately upon finding
them. The ovipositor was often allowed to remain inserted
several seconds while the larva twisted and squirmed. The
larvae were so pierced a great number of times during the
observation. These larvae when subsequently placed upon
Opuntia, entered it and fed normally for some two weeks
when the lot was abandoned preparatory to a change of head-
quarters.
These observations indicate that, in the field, this parasite
oviposits in the larvae at the time they leave the originally
infested, fallen joints and crawl again upon the plant to enter
new joints.
Pareniaca schwarsi Cwfd. and Chalcis pedalis Cress, have
been found as common, though not abundant, parasites of
0. junctolineella.
Second in importance to the Apanteles above mentioned
are Frontina aletice Riley and Phaonia n. sp. These dipterous
parasites affect the host larva similarly to Apanteles
JULY, 1925] HAMLIN OPUNTIA INSECTS
11
mimoristcE, the puparia appearing within the host cocoon
shortly after the latter is spun. The numbers are not
nearly so large, however, usually from four to eight being
present.
PREDATORS
This species is subject to attack by ants, not only as
mature larvae seeking the ground for pupation, but also as
small larvae emerging from originally entered joints which
fell to the ground under the combined attack of the newly
hatched larvae from one or several egg-sticks. It has been
noted above that the falling of joints infested with small
larvae brings about the distribution of the larvae over the
cactus clump. It is believed that the depredations of ants
at this time are responsible for the scarcity of larvae in the
average clump.
ZOOLOGICAL RECORD— PART INSECTA
The ‘Tnsecta” part of the “Zoological Record” (as dis-
tinguished from the complete volume) will in future be pub-
lished by, and only obtainable from, the Imperial Bureau of
Entomology. The price for the part will be 15/-, as heretofore.
It is, however, proposed as an experiment to break up a limited
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follows :
Section A. List of Titles and Subject Index 4/-
Section B. Coleoptera 6/-
Section C. Lepidoptera 6/-
Section D. Hymenoptera and Diptera 4/-
Section E. Hemiptera, Orthoptera and remaining
Orders 4/-
The above division has been instituted for the benefit of those
entomologists who are interested in a portion only of the sys-
tematic part of the work. It is in the nature of an experiment
only and cannot be continued unless it is widely supported.
All orders for the “Insecta” part, or any sections of it, should
be addressed to the Assistant Director, Imperial Bureau of
Entomology, 41, Queen’s Gate, London, S. W. 7. Orders for
the complete volume of the “Zoological Record” should con-
tinue to be sent to the Zoological Society of London, Regent’s
Park, London, N. W. 8.
12
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
NEW AND RARE LEPIDOPTERA FROM
THE SOUTHWEST
BY WM. BARNES AND F. H. BENJAMIN
Decatur, Illinois
Timochares ruptifasciatus Ploetz
1884, Ploetz, Jahr. Nass. Ver. Nat, XXXVII, 27, Antigonus.
1896, Godnian & Salvin, Biol. Centr.-Amer., Lep., Rhop., II, 418,
pi. LXXXVIII, ff. 1-2, Timochares.
1901, Skinner, Ent. News, XII, 171, Timochares.
1905, Dyar, Jour. N. Y. Ent Soc., XIII, 119, Timochares.
1921, Lindsey, Hesp. Am., p. 56, Timochares.
1923, Skinner & Williams, Trans. Am. Ent. Soc., XLVIII, 305, fig.
29 6 genit., Timochares.
Apparently U. S. records of this species are rare. Lindsey
records a specimen from Brownsville, Texas, Barnes collec-
tion, but Skinner and Williams have apparently overlooked
this and record only the Runyon catch from Southern
Arizona or California.
Mr. O. C. Poling recently sent a specimen from the
Baboquivari Mts., Pima County, Arizona, 20 September,
1924. This we submitted to Mr. Shaus for comparison with
his Mexican specimens as the markings are somewhat
darker than those of the Texas specimen before us.
Polia stricta race kappa Barnes & Benjamin, nov.
Head and thorax ferruginous, more or less darkened by black
scales. Abdomen luteous, tinged with rufous. Fore wing variable in
color, the ground color some shade of luteous, suffused with bright
ferruginous red and more or less darkened by black scales, in general
the ferruginous cast dominant; basal maculation indistinct but basal
dash usually present on the freshest specimens. Ordinary lines,
spots, and maculation, as in typical stricta. Hind wing of 6 white,
the veins and outer margin more or less soiled by fuscous. An
obsolescent discal spot. Fringe white at tip, with an obsolescent
interline and yellowish-white base. Hind wing of $ similar but with
more fuscous on it.
Expanse : 6 , 23-28 mm., 2 , 24-28 mm.
Similar to typical stricta but differing from all described
forms by the much paler hind wings which are nearly as
white as those of spiculosa.
JULY, 1925] BARNES AND BENJAMIN LEPIDOPTERA
13
At first we considered the present subspecies to be tenisca
and may have determined it as such for other collections.
“ M amestra” tenisca was described by Smith from a series
of specimens from Bozeman, Montana, and Stockton, Utah,
the type 6 being a Bozeman specimen in the Smith collec-
tion. The recent acquisition of a series of specimens from
both of the type localities of tenisca show that Nevada
material is not quite typical as it varies from forms nearly
as pale as kappa to forms as dark, as tenisca. The specimen
from Reno, Nevada, figured by Barnes & McDunnough
(1913, Contrib. N. H. Lep. N. A., II, (1), 22, pi. X, f. 10)
seems nearly typical of 9 tenisca. While all forms of stricta
seem more or less variable in each locality, and Nevada
material is intermediate, producing some specimens which
can be placed under any of the present subspecific names,
nevertheless our series of the white hind winged form is so
distinct from the variations found in the type localities of
the described subspecies, and in itself is so constant with
its pale secondaries, that we have no hesitation in describing
it.
Type locality: Inyo County, California, mostly from
Round Valley.
Nimiber and sexes of types: Holotype 6 , 15-30 Aug.,
Allotype 9 , 220 Paratypes, both sexes, various dates Aug.,
Sept., Oct.; O. C. Poling collector. Types in Barnes collec-
tion.
Graptolitha tarda Barnes & Benjamin, sp. nov.
Head, collar and thorax dull ash gray mixed with black. Thoracic
crest large, marked by a slim black stripe. Abdomen basally gray,
caudally more ochreous. Fore wing dull ash-gray, the costal region
somewhat paler than the remainder of the wing. Ordinary lines and
spots obsolescent, nearly obsolete; no basal line or dash; t. a. line
marked by a few black scales; median line similar; their course
indeterminate. Orbicular absent; reniform mainly visible as a slight
rufous cast at end of cell. T. p. line so poorly marked its exact
course cannot be seen, but apparently with a tendency to be produced
to points on the veins and to be bent inward in submedian fold where
it shows as a V pointed inward, and is connected to a pointed clavi-
form by a small black bar. S. t. line indicated by a few scattered
black scales. Veins of terminal area slightly darkened. Fringe a
little browner than the wing color and obscurely interlined. Hind
14
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
wing pale red-brown with a decided pinkish cast; usual discal dot and
terminal line present but not conspicuous. Fringe pale, whitish
tinged with pink. Beneath: dull; fore wing gray tinged with pink;
hind wing whitish tinged with grey and pink. Discal dot present on
each wing.
Expanse: 36 mm.
Allied to fagina and itata. Smaller, duller, and darker than
the former, with the markings not so distinct, but although
at once separable, closely allied by the paler costal area,
lack of a basal dash, and the proximity of the claviform to
the t. p. line, leaving room for only a very short black dash
in the submedian fold. Closely allied to itata, with which
it agrees in wing-shape, size, and general color. In true
itata, as represented by the types, there is a tendency for
the transverse maculation to be obscured, but the black
dashes in the reniform, and the median and subterminal
parts of the submedian fold are distinct, as is the subapical
shade. While the present species has the distinct bar in the
submedian fold connecting the claviform to the t. p. line,
the remainder of the longitudinal maculation has been so
reduced that it is invisible except for the faint indication
of a few black scales, while the orbicular is entirely absent
and the reniform mainly indicated by a reddish suffusion.
The legs of the present species lack the strong longitudinal
black lines present, one on each leg, of allied species.
Type locality: Douglas, Arizona.
Number and sexes of types: Holotype 6 , 16-23 June,
unique, in Barnes collection.
Xanthothrix ranunculi form albipuncta Barnes & Benjamin,
nov.
Head, thorax and fore wing black dusted with olivaceous,
the latter with a round white spot near end of cell. Sec-
ondaries blackish. Other characters as in ranunculi which is
a very variable species.
Expanse: 6,11 mm.; $ , 10 mm.
No mention is made in the original description of
ranunculi of any white spotted form, nor do the types in the
Henry Edwards collection show any traces of spots on the
primaries.
JULY, 1925] BARNES AND BENJAMIN LEPIDOPTERA
15
Type locality: Kaweah, California.
Number and sexes of types: Holotype 6 , Allotype $ ,
both 10 April, 1902, in Barnes collection.
Laniifera CYCLADES Druce
1895, Druce, Biol. Centr.-Amer., Lep., Het., II, 220, pi. LXI, ff.
12-13, Facliynoa.
1899, Hampson, Proc. Zool. Soc. Lond., p. 184, Layiiifera.
1903, Dyar, Bull. U. S. N. M., LI I, 380, No. 4328, Laniifera.
We received a single specimen from O. C. Poling, Babo-
quivari Mts., Pima County, Arizona, 1-15 August 1923,
which agreed well with Druce’s figures and with a specimen
from Mexico, ex Kearfott collection. A second specimen
has been received from Dr. Vorhies of the Arizona Experi-
ment Station bred from prickly-pear and labeled Santa Rita
Mts., Arizona, VI-23-24.
This species, described from Mexico, was recorded in
Dyar’s list from Arizona, presumably by Prof. Fernald.
In view of the recent research in insect enemies of the
prickly-pear, this note may be of some importance. The
insect is a member of the Pyralid subfamily Pyraustinge, is
of large size and decided possibility in the control of the
cactus.
The Van Duzee Collection of Hemiptera
At the regular August meeting of the California Academy of
Sciences, Mr. E. P. Van Duzee presented to the Academy his
entire collection of Hemiptera under practically the same con-
ditions under which the Van Dyke Collection of Coleoptera
was accepted by the Academy last year, as announced in this
journal (Vol I, page 13). Under these conditions he retains
general supervision over the collection during his lifetime with
the privilege of temporarily withdrawing for study any of the
specimens he may need for use at home. He also stipulated
that each specimen in this collection is to have attached a
label to read “E. P, Van Duzee Collection,” and that all these
specimens are to be incorporated into the systematic collections
of the Academy and that they are not to be used for exchange
or for exhibition purposes. These conditions secure to him the
use of the material during his lifetime and safeguard it against
being dissipated or lost through carelessness or mismanagement
in the future.
16
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
NOTES ON ARIZONA PHALAENIDZE (Lepid.)
BY WILLIAM BARNES AND F. H. BENJAMIN
Decatur, Illinois
Polia luteimacula* Barnes & Benjamin, n. sp.
Male antennae serrate and fasciculate. Head and thorax black,
gray, and white mixed, more or less tinged with purple and rufous.
Fore wing: ground color gray heavily irrorated and overlaid by
black, with some rufous suffusion, and a purplish tinge; basal line
obsolescent, waved; t. a. line double, its course slightly outwardly
oblique, as outcurved crescents, from costa to median vein, thence
to vein 1, and to inner margin; orbicular and reniform not large,
faintly outlined by black, pale filled, the filling obscured by dusky
central marks and heavy powderings, more or less red tinged, which
shade extends beyond the cell to the t. p. line; claviform moderate to
small, variable, distinctly outlined by black, filled with a rufous shade
which continues to the t. p. line; t. p. line double, black, its outer
line indistinct, angled outward below costa, thence oblique to about
vein 5, thence incurved, produced to a strong outward point on vein 1,
otherwise produced to moderate points on the veins; s. t. a yellowish
wavy line defined on both sides by black and terminating in a moder-
ately large, black defined, spot at tornus; a thin black terminal line;
fringe fuscous-gray, with darker interline and tips, paler basally.
Hind wing: nearly uniformly fuscous, the veins slightly darker;
fringe luteous at base, with fuscous interline and pale tips. Beneath:
with a dark common line; in general fuscous, the hind wing some-
what paler basally; discal spot obsolete on fore wing, obsolescent on
hind wing; fringes as on upper side.
Expanse: 6, 27-32 mm.; 9, 30-31 mm.
Closely allied to rubrifusa Hamp., but the uniformly dark
secondaries easily distinguish it. P. gnata Grt. belongs to
the same group but its type, in the Neumoegen collection,
possesses white hind wings P. agnata Sm. is also allied
* Indicates specimens were submitted to Dr. William Scliaus in order
to ascertain that the species had not been described from south of the
U. S. border.
2 A further discussion of gnata and several other types in the Neu-
moegen collection has been incorporated in a separate paper.
3 We wish to call attention to the fact that male agnata Sm. has serrate
antennae, and if Hampson’s characters are employed, belongs to this group.
Also that segregata Sm. and gussata Sm. are incorrectly placed on our
lists, having serrate male antennae and being closely allied to pUcata Sm. and
negussa Sm. In fact, we suspect that negussa is simply smooth gussata,
and the same may be true between segregata and pUcata. Careful study, based
on abundant material, will be necessary to ascertain the exact relation-
ship of these four names.
JULY, 1925] BARNES AND BENJAMIN PHAL^NID^
17
Locality: White Mts., Arizona.
Types: Holotype 6 ; Allotype $ ; 8 6,2 2, Paratypes.
Notes: From Barnes collection, collector unknown;
types in Barnes collection; Paratype in U. S. N. M.
Nephelistis Hamp
Type Nephalistis congenitalis Hamp.
1905, Hampson, Cat. Lep. Phal. B. M., VI, 250, type designated
congenitalis.
Close to, but probably distinct from, Trichopolia Grt., see
below.
Trichopolia Grt.
Type Trichopolia dentatella Grt.
1883, Grote, Pap. Ill, 76, lists dentatella and questionably ptilodonta. ,
1895, Grote, Abh. Nat. Ver. Bremen, XIV, 81, type designated
dentatella.
1905, Hampson, Cat. Lep. Phal. B. M., V, 361, partim., type
designated dentatella.
1923, Barnes & Benjamin, Bull. B’klyn Ent. Soc., XVIII, 123, type
designated dentatella.
Eupolia Sm.
Type Eupolia licentiosa Sm.
1894, Smith, Trans. Amer. Ent. Soc., XXI, 69; p. 70, licentiosa sole
species and therefore type.
1909, Hampson, Cat. Lep. Phal. B. M., VIII, 544, type designated
licentiosa, but erroneously placed as section I of Namangana.
1923, Barnes & Benjamin, Bull. B’klyn Ent. Soc., XVIII, 123, type
designated licentiosa; licentiosa = dentatella, Eupolia =
Trichopolia.
We describe the following species in Trichopolia although
our three specimens lack a dorsal tuft on the second
abdominal segment. This tuft in dentatella is easily lost, is
only visible on some specimens and as all other characters
seem very similar, we think the tuft has probably been
rubbed off our types of suspicionis.
Structurally the species would appear to fall into Hamp-
son’s Nephelistis. We have already pointed out (1923) that
Hampson drew his description of the genus Trichopolia from
specimens of ursina Sm., and found it necessary to erect the
genus Engelhardtia for type ursina.
18
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
We do not know if the genus Nephelistis will fall into the
synonomy of Trichop olia or not. Certainly it seems very
closely allied, and mainly separated on secondary sexual
characters.
Trichopolia comes very close to Scotogramma, but the front
is not as much rounded out. It is also closely related to
Folia, and some species now placed in Eriopyga. Most of
these hairy-eyed genera need careful study to ascertain
which should be retained and what species placed in them.
Tentatively we place Trichopolia between Admetovis and
Lophoceramica. This placement between Trichoclea and
Chabuata brings Trichopolia into contact with its obvious
exotic affinities, which possess pectinate antennae in the
male sex, such as Hydroeciodes and Nephelistis.
Trichopolia suspicionis* Barnes & Benjamin, n. sp.
Male: Antennae bipectinate, apically serrate and simple, ciliated.
Mid tibiae with tuft of long hair but not conspicuously dilated.
$ antennae simple, the joints slightly marked, ciliated. Androconia
absent in both sexes. Palpi pale, marked with black on sides, second
joint reaching about middle of frons, third joint porrect. Head and
thorax brown irrorated with black, abdomen similar, with dorsal
tuft on basal segment only; and slight lateral tufts.
Fore wing: ground color dull wood-brown, grayed by a scattering
of black scales; transverse lines single, black; basal line from costa
to the end of a short basal black dash, forming a loop; t. a. line
outwardly oblique from costa to inner margin, only slightly excurved;
claviform obsolescent; orbicular pale, moderate, slightly oblique,
faintly outlined by black and with black powdered center; reniform
centrally constricted, edged with an obsolescent series of black
scales, then a pale faint line followed by a strong black edging,
centrally with a blackish outlined inner crescent, the whole mark
heavily powdered by black; t. p. line erect on costa, strongly excurved
around cell, inwardly oblique, only a little incurved, below; s. t. line
obsolete; terminal line thin, broken by veins; fringe pale, checkered
by black.
Hind wing: whitish, subhyaline, with small discal spot and darker
veins, some fuscous suffusion outwardly; a thin terminal fuscous
line; fringe white, obsolescently interlined, and with yellowish base.
Beneath: whitish; the fore wing more or less suffused with fuscous,
especially in cell and on veins, with discal spot and t. p. line
showing through, terminal line and fringe as on upper side; hind
wing with only a scattering of fuscous scales, mostly on veins; ter-
minal line and fringe as on upper side.
Expanse: 26-28 mm.; 2, 26 mm.
JULY, 1925] BARNES AND BENJAMIN PHAL^NIDJE
19
From the only allied species, dentatella, known to us, the present
species differs by the pectinations of the male antennae being but
about half as long, while the general coloration is much paler, the
transverse lines much stronger and blacker.
Locality: Sells Post Office, Indian Oasis, Pima County,
Arizona.
Types: Holotype 6 , /Vllotype $ , 1 6 Paratype, all 1-10
May, 1923.
Notes: O. C. Poling, collector.
Athetis Hbn.
Type Noctua dasychira Hbn.
1822, Hubner. Verz. bek. Schmett., p. 209, lists dasychira, palustris,
caliginosa and cubicularis.
1909, Hampson, Cat. Lep. Phal. B. M. VIII, 299, type designated
furvula, apparently for synonym dasychira.
1911, Warren, in Seitz, Macrolepid., Ill, 207; p. 208, type designated
furvula, apparently for synonym dasychira.
1916, McDunnough, Ent. News, XXVII, 397, replaced by Caradrina.
Atypha Hbn.
Type Noctua pulmonaris Esp.
1822, Hubner, Verz. bek. Schmett., p. 213, lists pulmonaris argillacea
and ochroleuca.
1909, Hampson, Cat. Lep. Phal. B. M., VIII, 299, type designated
pulmonaris.
HapxVlotis Hbn.
Type Noctua furvula Hbn.
1825, Hubner, Verz. bek. Schmett., p. 254, lists furvula, lupula, ravula,
fuscula (poly gramma), atratula, candidula and venustula
(hybnerana).
1909, Hampson, Cat. Lep. Phal. B. M., VIII, 299, type designated
furvuila.
Nebrissa Wlk.
Type Nehrissa himacula Wlk.
1862, Walker, Jour. Linn. Soc., Zook, VI, 194.
1909, Hampson, Cat. Lep. Phal. B. M., VIII, 299, type designated
himacula.
Anorthodes Sm.
Type Anorthodes prima Sm.
1891, Smith, Trans. Amer. Ent. Soc., XVIII, 114, prima sole species
and therefore type.
1909, Hampson, Cat. Lep. Phal. B. M., VIII, 299, type designated
tarda, apparently for synonym prima.
20
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. II, NO. 1
Athetis Hbn. as used by Hampson is apparently correct
in place of Caradrina as used in the Barnes & McDunnough
Check List. Caradrina has type i-intactum by Duponchel,
1829, and will likely replace Lycophotia Hbn. as used by
Hampson and Barnes & McDunnough.
Elaphria Hbn., placed by Hampson as a synonym of
Athetis was first described in the Zutr. Exot. Schmett., I,
pi. XHI, grata sole species and therefore type. This will
have priority over the monotypic genera Monodes Gn. (type
nucicolora Gn.) and Alpesa Wlk. (type villicosta Wlk.)
Athetis indigena* Barnes & Benjamin, n. sp.
Palpi cream-colored, black at the sides. Head and thorax pale
brownish with a violaceous cast. Collar with a broad black band
at its base, else concolorous with the thorax. Abdomen pale brownish
above, yellowish beneath, the genitalia of the male surrounded by
long yellow hair. Fore wing: ground color dull pale wood brown,
with a slight violaceous tinge and a silken lustre, darkened and
grayed by scattered black scales, lines black; basal line geminate,
its outer line indistinct, waved from costa to submedian fold,
obsolescent below; t. a. line geminate, its inner line obsolescent,
appearing as a dot on costa and again as a thin irregular line
from costa to inner margin; orbicular obsolescent, more or less
indicated by a faint ring of black scales; claviform obsolete; reni-
form indistinct, black centered due to intersection by an obscure
median shade, marked by four pale yellowish points; t. p. line faintly
geminate, the inner line stronger, as a dot on costa excurved around
cell, incurved below, produced to points on the veins; s. t. obsolescent,
its course indicated by a faint yellowish shade which is marked in-
wardly by slight dusky shades and one or two black dashes; a thin
terminal line broken by the veins; fringe luteous at base, dark and
obscurely interlined. Hind wing: 6, whitish, subhyaline, the veins
and outer portion darkened by fuscous; $, darker, almost appearing
fuscous; fringe yellowish at base, with gray interline and white tips.
Beneath: whitish, the fore wing largely suffused with fuscous with
slight discal dot and transverse line; hind wing with scattered fuscous
scales mainly along veins and toward the outer margin which is
somewhat darkened, with discal dot and transverse line: fringes, all
wings, as on upper side: sexes similar on under side.
Expanse: 6, 29-32 mm.; 2, 31-33 mm.
Allied to triquetra, with a similar set of black subterminal
marks but lacking the black in the costal-subterminal
region ; at once distinguishable by the entirely different type
JULY, 1925] BARNES AND BENJAMIN PHAL^NID^
21
of reniform and the black at the base of the collar. It is
also considerably heavier in build. Perhaps triquetra and
indigena should be placed in Ariathisa; but we prefer to
retain them with their North American allies.
''Caradrina” derosa Morr., is placed by Hampson as a
synonym of capularis Gn,, but a figure of the type would
indicate that it might be more nearly related to indigena.
We know of nothing like this figure, which does not seem
to match Hampson’s plate CXXXII, figure 12 of capularis;
having strong black markings on the reniform. Not know-
ing how much capularis varies, we hold the status of derosa
in abeyance. Even if distinct from capularis it is probably
exotic.
Localities and number and sexes of types: Holotype 6 ,
Brown’s Canon, Baboquivari Mts., Pima County, Arizona,
elevation approximately 5000 ft., (O. C. Poling), 1-15, June,
1923 ; Allotype $ , id. ; 2 6 Paratypes, id. ; 1 6 1 9 Para-
types, Baboquivari Mts., Pima County, Arizona, elevation
approximately 5000 feet (O. C. Poling), 15-30 June, 1923,
1 3 Paratype, Palmerlee, Arizona, 8-15 May.
Notes: Types in Barnes collection: Parqtype in U. S.
N. M.
CeLIPTERA VALINA Schs.
1901, Schaus, Ann. Mag. N. H., (7), VIII, 86, Mods.
1913, Hampson, Cat. Lep. Phal. B. M., XIII, 135, pi. CCXXV,
f. 5 (type), Celiptera.
Two males, three females, were received from the Babo-
quivari Mts., Arizona (O. C. Poling), dates 15-30 July,
1-15 August, 15-30 August, 1-15 September.
The females agreed with Hampson’s figure. Two males,
one female, were sent to Dr. Schaus for comparison with his
type and series.
The species is rather variable. It also appears somewhat
sexually dimorphic, the males darker marked and lacking
the yellow found in the transverse lines of the females. This
is replaced by rather diffused black.
22
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
A NEW ACINOPTERUS FROM SOUTHERN
CALIFORNIA
BY HERBERT OSBORN
Acinopterus aridellus Osborn, n. sp.
Head scarcely as wide as pronotum, vertex broad, distinctly
angulate, with a faint transverse depression, one-third longer at
middle than next the eye, longer and more pointed in male, front
broad, tapering uniformly from antennal pits, clypeus long, expand-
ing towards the tip, lorse large, nearly touching margin of cheek,
cheek margins sinuate, pronotum a little longer than vertex, elytra
long, apex rather broad, acute at tip; female last ventral segment
twice as long as preceding, hind margin rounded, minutely notched
at tip; male valve concealed or wanting, plates narrow, long, reaching
nearly to tip of pygofer.
Color: dark gray, suffused with pinkish, especially at tip of vertex
and on the elytral veins; ocelli whitish, three obscure whitish lines
on the pronotum, extending on to the scutellum; elytra hyaline, the
veins whitish, or tinged with pink, and margined with fuscous;
beneath yellowish, tinged with pinkish.
Length ; male and female, 4.5-5 mm.
Described from one female (type) two males, (allotype
and paratype) collected by the writer at Palm Canon,
Imperial County, California, March 8, 1924. They occurred
upon a small creeping plant outside the canyon mouth and
amongst barrel cactus, creosote, and sagebrush, in distinctly
arid conditions. It is decidedly smaller, especially more
slender, than acuminatus or angulatus, and the vertex is
longer, especially in the male, where it is quite distinctly
angulate. Type in Author’s collection.
Two Arizona Hemiptera
Ganocapsus filiformis Van Duzee was taken by me in large
numbers on a species of sunflower at Patagonia, Santa Cruz
County, Arizona, about the first of August, 1924. It occurred
on other weeds and grasses in the same locality, but seemed
to be breeding on the sunflower only. With it were taken
numbers of Garganus splendidus Distant, not before recorded
from north of the Mexican border. — E. P. Van Duzee.
JULY, 1925]
CHAMBERLIN SCOLYTID^
23
THE CONIFEROUS TREES OF THE UNITED
STATES WITH THE SCOLYTID (IPID)
BEETLES SAID TO ATTACK THEM
BY W. J. CHAMBERLIN
Oregon State College
Corvallis, Oregon
The following list contains such coniferous trees as are
reported to be the host of one or more of the bark or
ambrosia beetles. This list has been compiled from all
available literature, from field and laboratory records of the
Oregon Experiment Station, from ten years collecting in
the western forests by the author and lastly from material
obtained by exchange.
Although the list is far from complete and some of the
records as given in literature, especially in the older papers,
are probably inaccurate, it is felt that this preliminary list
will prove of value and may stimulate others to check it
carefully and make known any errors, corrections or addi-
tions so that a more complete and accurate list may be
compiled at a future time.
A few of the rarer trees are not listed, since no record of
their being attacked by scolytids, has been noted. Un-
doubtedly all of them have their scolytid enemies and
investigation of such trees should be carried on by those so
situated as to be able to do so.
Mr. Ralph Hopping’s paper, “Coniferous Hosts of the
Ipid^ of the Pacific Coast and Rocky Mountain Region,”
which appeared in the Canadian Entomologist, Vol. 54, pp.
128-134, has been of great value in compiling the present
paper.
The trees are listed alphabetically, according to genera
and species.
1. Abies amabilis. Lovely fir
Cryphalus amabilis Chamb.
Gnathotrichus retusus (Lee.)
sulcatus (Lee.)
Poylgraphus rufipennis (Kby.)
Pseudohylesinus nobilis Sw.
Trypodendron sp.
24
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
2. Abies balsamus. Balsam fir
Cryphalus balsameus Hopk.
Crypturgus balsameus Sw.
atomus Lee.
Dryocoetes confusus Sw.
Gnathotrichus retusus (Lee.)
suleatus (Lee.)
Pityokteines sparsus (Lee.)
Pityophthorus balsameus Blkm.
bassetti Blkm.
Pityophthorus eanadensis Sw.
opaeulus Lee.
patehi Blkm.
puberulus Lee.
^ Platypus wilsoni Sw.
Seolytus pieeae (Sw.)
2 Trypodendron lineatus (Oliv.)
ponderosse Sw.
3 Xyleborus xylographus (Say.)
3. Abies concolor. White fir
See species under A. grandis
4. Abies fraseri. Fraser’s fir
Cryphalus fraseri Hopk. Gnathotriehus retusus (Lee.)
5. Abies grandis. Grand fir
Conophthorus montieolae Hopk.
Cryphalus approximatus Hopk.
grandis Chamb.
pubeseens Hopk.
subeoneentralis Hopk.
Gnathotriehus retusus (Lee.)
suleatus (Lee.)
Pityokteines elegans Sw.
Platypus wilsoni Sw.
Pseudohylesinus grandis Sw.
elongatus Sw.
granulatus (Lee.)
Seolytus kigeri Chamb.
praseeps Leo.
unispinosus Lee.
ventralis Lee.
subseaber Lee.
Trypodendron lineatus Oliv.
Xyleborus xylographus Say.
6. Abies lasiocarpa. Alpine fir
Cryphalus borealis Sw.
eanadensis Chamb.
Dryoeoetes eonfusus Sw.
Gnathotriehus retusus (Lee.)
suleatus (Lee.)
Ips borealis Sw.
Orthotomieus lasioearpi Sw.
Pityokteines elegans Sw.
Pityophthorus pseudotsugse Sw.
Platypus wilsoni Sw.
Trypodendron sp.
7. Abies magnifica. Magnificent fir
Gnathotriehus retusus (Lee.)
suleatus (Lee.)
Leperisinus imperialis Eieh.
Pseudohylesinus granulatus
(Leo.)
Seolytus ventralis Lee.
Trypodendron lineatus Oliv.
Pityokteines elegans Sw.
Platypus wilsoni Sw.
Xyleborus xylographus Say.
1 Not a Scolytiae but so closely related in habits and work that the
family (Platypodidae) is included in this list.
2 This may be T. bivittatus (Kirby).
3 X. xylographus is given as breeding in a great many species of coni-
fers in the older publications. These are probably all errors in determina-
tion and this species probably confines its work to hardwood trees in the
Eastern United States.
JULY, 1925]
CHAMBERLIN SCOLYTID^
25
8. Abies nobilis. Noble fir
Gnathotrichus retusus (Lee.) Pseudohylesinus nobilis Sw.
sulcatus (Lee.) Trypodendron lineatus Oliv.
Hylastes longieollis Sw. Xyleborus xylographus Say.
Platypus wilsoni Sw.
9. Abies venusta. Bristle-cone fir
Gnathotriehus retusus (Lee.) Trypodendron lineatus Oliv.
suleatus (Lee.)
10. Chamcecyparis lawsoniana. Port Orford cedar
Phloeosinus punetatus Lee. Pityokteines minutus Sw.
11. Chamcecyparis nootkatensis. Alaska cedar
Phloeosinus punetatus Lee. Phloeosinus dentatus (Say)
nitidus Sw. Pityokteines minutus Sw.
12. Chamcecyparis thyoides. White cedar
Phloeosinus dentatus (Say) | >
13. Cupressus macrocarpa. Monterey cypress
Phloeosinus eristatus Lee. Phloeosinus eupressi Hopk.
14. Cupressus sargentii. Sargents cypress
Phloeosinus eupressi Hopk. Phloeosinus minutus Sw.
15. Juniperus calif arnica. Desert white cedar
Trypodendron sp.
16. Juniperus communis. Common juniper
Phloeosinus sp. Trypodendron sp.
dentatus (Say)
17. Juniperus flaccida. Juniper
Phloeosinus dentatus (Say)
18. Juniperus occidentalis. Western juniper
Phloeosinus juniperi Sw. Phloeosinus rugosus Sw.
19. Juniperus scopulorum. Rocky Mountain Juniper
Phloeosinus hoppingi Sw. Xyleborus scopulorum Hopk.
scopulorum Sw.
20. Juniperus virginiana. Eastern white cedar
Phloeosinus dentatus (Say) Phloeosinus enixus Blkm.
21. Larix alaskansis. Alaskan larch
Trypodendron lineatus Oliv. Xyleborus xylographus Say.
26
THE PAN-PACIFIC ENTOMOLOGIST [VOL. II, NO. 1
22. Larix americana. American larch
Crypturgus pusillus Gyll.
Dendroctonus simplex Lee.
Dryocoetes americanus Hopk.
autographus Ratz.
Ips pini (Say)
balsameus Lee.
Orthotomieus eselatus (Eieh.)
Polygraphus rufipennis Kby.
Seolytus pieea Sw.
* 23. Larix lyallii. Tamarack
Crypturgus atomus Lee.
Dendroetonus simplex Lee.
pseudotsugas? Hopk.
Dryoeoetes amerieanus Hopk.
Orthotomieus eslatus (Eieh.)
Gnathotriehus materiarius (Fiteh)
Hylurgops pinifex (Fiteh)
Ips pini Say
Phloeosinus serratus Lee.
Pityokteines sparsus (Lee.)
Pityophthorus opaeulus Lee.
Polygraphus rufipennis Kby.
Trypodendron lineatus Oliv.
bivittatum Kby.
Xyleborus xylographus Say
24. Larix occidentalis. Western larch
Crypturgus borealis Sw.
Dendroetonus pseudotsugas
Hopk.
valens Lee.
Ips emarginatus (Lee.)
Orthotomieus vieinus (Lee.)
Pityophthorus intextus Sw.
Polygraphus rufipennis Kby.
Seolytus unispinosus Lee.
Xyleborus xylographus Say
25. Lihocedrus d'seurrens. Incense cedar
Phloeosinus attenuatus Sw.
eupressi Hopk.
Phloeosinus punetatus Lee.
hoppingi Sw.
Phloeosinus vandykei Sw.
fulgens Sw.
Platypus wilsoni Sw.
26. Picea hreweriana. Weeping spruce
Dendroetonus valens Lee. Polygraphus rufipennis Kby.
Gnathotriehus suleatus (Lee.) Xyleborus xylographus Say
27. Picea canadensis. White spruce
Carphoborus earri Sw.
Cryphalus mainensis Blkm.
Crypturgus atomus Lee.
borealis Sw.
Dendroetonus engelmanni Hopk.
johanseni Sw.
pieeaperda Hopk.
terebrans (Oliv.)
valens Lee.
Dryoeoetes amerieanus Hopk.
affaber (Mann.)
earyi Hopk.
pieese Hopk.
septentrionis (Mann.)
Gnathotriehus materiarius
(Fitch)
suleatus (Lee.)
Hylurgops pinifex (Fitch)
* Undoubtedly most of the records for i. lyallu should be referred to
L. occidentalis . Lyallii is a rare alpine species.
JULY, 1925]
CHAMBERLIN SCOLYTID^
27
Ips chagnoni Sw.
engelmanni Sw.
interruptus (Mann.)
Ips pini (Say)
Orthotomicus cselatus (Fitch)
Phthorophlseus picea Sw.
Pityogenes hopkinsi Sw.
Pityophthorus dentifrons Blkm.
Pityophthorus intextus Sw.
nudus Sw.
nitidus Sw.
opaculus Lee.
patchi Blkm.
28. Picea engelmanni
* Dendroctonus borealis Hopk.
engelmanni Hopk.
monticolas Hopk.
murrayanffi Hopk.
ponderosae Hopk.
valens Lee.
Dryocoetes afifaber (Mann.)
americanus Hopk.
septentrionis (Mann.)
Gnathotrichus sulcatus (Lee.)
Hylastes macer Lee.
Hylurgops rugipennis (Mann.)
Ips dubius Sw.
engelmanni Sw.
interpunctus (Eich.)
yohoensis Sw.
Pityophthorus puberulus Lee.
shepardi Blkm.
Polygraphus rufipennis (Kby.)
Scierus annectens Lee.
Scolytus piceae Sw.
Stephanoderes quadridentatus
Hopk.
Tryopdendron bivittatus Kby.
borealis Sw.
cavifrons (Mann.)
Xyleborus xylographus Say
Xylechinus americanus Blkm.
Englemann spruce
Orthotomicus vicinus (Lee.)
Pityogenes carinulatus (Lee.)
Pityokteines minutus Sw.
Pityophthorus atratulus (Lee.)
bassett Blkm.
intextus Sw.
nitidulus (Mann.)
occidentalis Blkm.
pseudotsugae Sw.
Polygraphus rufipennis (Kby.)
Procryphalus striatulus (Mann.)
Scierus annectens Lec.
piceae Sw.
Scolytus unispinosus Lec.
Trypodendron lineatus Oliv.
ponderosae Sw.
29. Picea mariana. Black spruce
Crypturgus atomus Lec.
Dendroctonus rufipennis (Kby.)
frontalis (Oliv.)
rufipennis (Kby.)
piceaperda Hopk.
ponderosae Hopk.
terebrans (Oliv.)
valens Lec.
americanus Hopk.
Dryocoetes granicollis Lec.
Gnathotrichus materiarius
(Fitch)
sulcatus (Lec.)
Hylurgops pinifex (Fitch)
Ips grandicollis (Eich.)
pini (Say)
Orthotomicus caelatus (Eich.)
Pityophthorus cariniceps Lec.
1 hirticeps Lec.
tuberculatus Eich.
Polygraphus rufipennis (Kby.)
Trypodendron bivittatum (Kby.)
lineatus Oliv.
Xyleborus xylographus Say
Xyloterinus politus (Say)
* D. borealis and D. engelmanni are probably the same species.
1 P. fulchellus Eich.
28
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
30. Picea pungens. Blue spruce
Dendroctonus engelmanni Hopk.
ponderosse Hopk.
valens Lee.
Dryocoetes affaber (Mann.)
Gnathotrichus sulcatus (Lee.)
Ips pini (Say)
Polygraphus rufipennis Kby.
Trypodendron lineatus Oliv.
Xyleborus xylographus Say.
31. Picea rubra. Red spruce
Cryphalus mainensis Blkm.
rubentis Hopk.
Crypturgus atomus Lee.
Dendroetonus pieeaperda Hopk.
punetatus Lee.
terebrans Oliv.
valens Lee.
Dryoeoetes amerieana Hopk.
Gnathotriehus materiarius
(Fiteh)
suleatus (Lee.)
Hylurgops pinifex (Fiteh)
Orthotomieus eaelatus (Eieh.)
Pityophthorus biovalis Blkm.
briseoei Blkm.
opaeulus Lee.
puberulus Blkm.
shepardi Blkm.
dentifrons Blkm.
Polygraphus rufipennis Kby.
Trypodendron lineatus Oliv.
Xyleborus xylographus (Say)
Xyleehinus amerieanus Blkm.
32. Picea sifehensis. Sitka spruce
Crypturgus borealis Sw.
Dendroetonus obesus (Mann.)
Dolurgus pumilus (Mann.)
Dryoeoetes affaber (Mann.)
septentrionis (Mann.)
Hylurgops rugipennis (Mann.)
Ips eoneinnus (Mann.)
interruptus (Mann.)
Ips vaneouveri Sw,
Pityophthorus atratulus (Lee.)
nitidulus (Mann.)
punetieollis (Lee.)
Platypus wilsoni Sw.
Polygraphus rufipennis (Kby.)
Pseudohylesinus sitehensis Sw,
Trypodendron cavifrons (Mann.)
33. Pinus alhicaulis. White-bark pine
Dendroetonus montieolae Hopk. Gnathotriehus sulcatus (Lee.)
34. Pinus attenuata. Knobcone pine
Dendroetonus montieolae Hopk. Gnathotrichus asperulus (Lee.)
valens Lee.
35. Pinus balfouriana. Foxtail pine
Dendroctonus montieolae Hopk.
valens Lee.
Gnathotrichus sulcatus Lee.
Hylurgops pinifex (Fitch)
rugipennis (Mann.)
Ips grandicollis Eieh.
vaneouveri Lee.
Xyleborus pubescens Zimni.
xylographus (Say.)
JULY, 1925]
CHAMBERLIN SCOLYTID^
29
36. Pimis caribcea. Slash pine
Crypturgus pusillus Gyll.
Dendroctonus frontalis Zimni.
terebrans Oliv.
Ips calligraphus (Germ.)
grandicollis (Eich.)
Pityogenes comatus (Zimm.)
granulatus Sw.
plagiatus (Lee.)
Pityophthorus publicarius
(Zimm.)
37. Finns clausa. Sand pine
Crypturgus pusillus Gyll. Ips grandicollis (Eich.)
Dendroctonus frontalis Zimm. Pityoborus comatus Zimm.
38. Finns contorta (murr ay ana). Lodgepole pine
Conophthorus contortse Hpk.
Crypturgus pusillus?
Dendroctonus monticolse Hpk.
murrayanae Hopk.
ponderosae Hopk.
valens Lee.
Gnathotrichus retusus (Lee.)
sulcatus (Lee.)
Hylurgops lecontei Sw.
pinifex (Fitch)
rugipennis (Mann.)
subcostulatus (Mann.)
Ips concinnus (Mann.)
emarginatus (Lee.)
grandicollis (Eich.)
guildi Blkman.
interpunctatus (Eich.)
latidens (Lee.)
Ips oregoni (Eich.)
pini (Say)
plastographus (Lee.)
radiatse (Hopk.)
vancouveri Sw.
Orthotomicus ornatus Sw.
Pityogenes carinulatus (Lee.)
fossifrons (Lee.)
knechteli Sw.
Pityophthorus atratulus Lee.
confertus Sw.
murrayanae Blkman.
nitidulus (Mann.)
tuberculatus Eich.
Pseudohylesinus sericeus
(Mann.)
Scierus annectens Lee.
39. Finns coulteri. Coulter pine
Dendroctonus terebrans (Oliv.) Dendroctonus valens Lee.
* brevicomis Lee. Ips calligraphus (Germ.)
40. Finns divaricata. Jack pine
Dendroctonus punctatus Lee.
rufipennis Kby.
Monarthrum fasciatum (Say)
Pityogenes hopkinsi Sw.
Pityophthorus annectens Lee.
Phloesinus pini Sw.
41. Finns echinata. Shortleaf pine
Carphoborus bicristatus Chap.
Crypturgus pusillus Gyll.
Dendroctonus frontalis Zimm.
terebrans (Oliv.)
Gnathotrichus materiarius
(Fitch)
Ips avulsus (Eich.)
grandicollis (Eich.)
* Record by Ralph Hopping.
30
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
Orthotomicus caelatus (Eich.)
Pityogenes plagiatus (Lee.)
Pityophthorus annectens (Lee.)
eomatus (Zimm.)
granulatus Sw.
lautus Eieh.
Pityophthorus nudus Sw.
puliearius (Zimm.)
pullus (Zimm.)
Platypus flavieornis (Fab.)
Stephanoderes quadridentatus
Hopk.
42. Pinus edulus. Pinion
Conophthorus edulis Hopk. Ips eonfusus (Lee.)
43. Pinus flexilis. Limber pine
Dendroetonus montieolse Hopk. Pityophthorus nitidulus (Mann.)
ponderosse Hopk. Polygraphus rufipennis (Kirby)
44. Pinus glabra. Spruce pine
Dendroetonus frontalis Zimm. Ips pini (Say)
terebrans (Oliv.) Pityophthorus anneetens Lee.
45. Pinus jeffreyi. Jeffrey pine
Crypturgus pusillus Gyll.
Dendroetonus jeffreyi Hopk.
valens Lee.
Gnathotriehus retusus (Lee.)
Hylastes maeer Lee.
Hylurgops subeostulatus (Mann.)
Ips eoneinnus (Mann.)
emarginatus (Lee.)
Ips grandieollis (Eieh.)
latidens (Lee.)
oregoni (Eieh.)
radiatae Hopk.
Orthotomieus ornatus Sw.
Pityogenes earninulatus (Lee.)
Pityophthorus eonfinis Lee.
Xyleborus impressus Eieh.
46. Pinus lahmertiana. Sugar pine
Conophthorus lambertianse
Hopk.
Crypturgus pusillus Gyll.
Dendroetonus montieolse Hopk.
valens Lee.
Gnathotriehus retusus (Lee.)
suleatus (Lee.)
Hylurgops rugipennis Mann.
47. Pinus monticola.
Conophthorus montieolse Hopk.
Dendroetonus monticolae Hopk.
valens Lee.
Gnathotriehus retusus (Lee.)
suleatus (Lee.)
Hylastes nigrinus (Mann.)
Hylurgops subeostulatus Mann.
Ips eonfusus (Lee.)
emarginatus (Lee.)
oregoni (Eieh.)
latidens (Lee.)
Pityophthorus confertus Sw.
eonfinis (Lee.)
tubereulatus Eich.
Western white pine
Hylurgops pinifex (Fitch)
porosus (Lee.)
rugipennis (Mann.)
subeostulatus (Mann.)
Ips eonfusus (Lee.)
eoneinnus (Mann.)
JULY, 1925]
CHAMBERLIN SCOLYTID^
31
Ips integer (Eich.)
interruptus (Mann.)
latidens (Lee.)
oregoni (Eich.)
vancouveri Sw.
grandicollis (Eich.)
avulsus (Eich.)
Pityogenes fossifrons (Lee.)
48.
Dendroctonus monticolse Hopk.
49. Pinus muricata.
Pityophthorus carmeli Sw.
Pityophthorus atratulus (Lee.)
confertus Sw.
confinis Lee.
nitidulus (Mann.)
Pityokteines sparsus (Lee.)
elegans Sw.
Platypus wilsoni Sw.
Scolytus monticolas Sw.
Pinus monophylla. Single leaf pine
Ips confusus (Lee.)
California swamp pine or prickle-cone
pine
Polygraphus rufipennis (Kby.)
50. Pinus palustris. Long-leaf pine
Crypturgus alutaceus Sz.
pusillus Gyll.
Dendroctonus frontalis Zimm.
terebrans (Oliv.)
Ips avulsus (Eich.)
grandicollis (Eich.)
Orthotomicus cselatus (Eich.)
Pityogenes plagiatus (Lee.)
Pityophthorus annectens Lee.
comatus (Zimm.)
pulicarius (Zimm.)
Xyleborus confusus Eich.
51. Pinus ponderosa.
Carphoborus simplex Lee.
Conophthorus ponderosse Hopk.
Dendroctonus approximatus Dz.
arizonicus Hopk.
1 barberi Hopk.
brevicomis Lee.
convexifrons Hopk.
jeffreyi Hopk.
monticolae Hopk.
ponderosse Hopk.
terebrans? Oliv.
valens Lee.
Gnathotrichus retusus (Lee.)
sulcatus (Lee.)
Hylastes porosus (Lee.)
Hylurgops lecontei Sw.
rugipennis (Mann.)
subcostulatus (Mann.)
Ips concinnus (Mann.)
confusus (Lee.)
grandicollis (Eich.)
emarginatus (Lee.)
Western yellow pine
Ips integer (Eich.)
interpunctus (Eich.)
knausi Sw.
latidens (Lee.)
oregoni (Eich.)
pini (Say)
radiatse Hopk.
Orthotomicus ornatus (Eich.)
Pityogenes carinulatus (Lee.)
fossifrons (Lee.)
Pityophthorus atratulus (Lee.)
cariniceps Lee.
Pityophthorus confertus Sw.
confinis Lee.
nitidulus (Mann.)
tuberculatus Eich.
Platypus wilsoni Sw.
Trypodendron ponderosae Sw.
Xyleborus xylographus (Say)
scopulorum Hopk.
1 D: barberi Hopk. is probably synonymous with D. brevicomis Lec.
32
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
52. Pinus pungens. Table-mountain pine
Crypturgus pusillus Gyll. Ips grandicollis (Eich.)
Dendroctonus frontalis Zimm. Pityogenes plagiatus (Lee.)
terebrans (Oliv.) Pityophthorus confinis Lee.
Gnathotrichus materiarius (Fiteh) puberulus Lee.
53. Pinus radiata. Monterey pine
Conoplithorus radiatae Hopk.
Carphoborus radiatas Sw.
Dendroetonus valens Lee.
Hylastes nigrinus (Mann.)
Hylurgops rugipennis (Mann.)
Ips plastographus (Lee.)
radiatae Hopk.
Pityophthorus atratulus (Lee.)
earmeli Sw.
Pseudohylesinus serieeus (Mann.)
54. Pinus resinosa. Red pine
Dendroctonus frontalis Zimm.
punctatus Lee.
Ips chagnoni Sw.
perroti Sw.
pini (Say)
Monarthrum fasciatum (Say)
Pityogenes hopkinsi Sw.
55. Pinus rigida.
Dendroctonus frontalis Zimm.
Gnathotrichus materiarius
(Fitch)
Ips callographus (Germ.)
Pityophthorus balsameus Blkm.
canadensis Sw.
confinis Lee.
puberulus Lee.
pulicarius (Zimm.)
Trypodendron scabricollis (Lee.)
Pitch or Pond pine
Ips pini (Say)
grandicollis (Eich.)
Polygraphus rufipennis (Kby.)
Trypodendron bivittatum (Kby.)
56. Pinus sahiniana. Gray or Digger pine
Dendroctonus valens Lee. Ips latidens (Lee.)
Ips grandicollis (Eich.)
57. Pinus strobus.
Conophthorus coniperda Sz.
Crypturgus atomus Lee.
Dendroctonus frontalis Zimm.
punctatus Lee.
rufipennis (Kby.)
terebrans (Oliv.)
valens Lee.
Dryocoetes americanus Hopk.
Gnathotrichus materiarius (Fitch)
retusus Lee.
Hylastes porculus Erich.
Hylurgops pinifex (Fitch)
Eastern white pine
Ips avulsus (Eich.)
calligraphus (Germ.)
chagnoni Sw.
longidens Sw.
pini (Say)
Monarthrum fasciatum (Say)
mali (Fitch)
Orthotomicus caelatus (Eich.)
Pityogenes hopkinsi Sw.
Pityokteines sparsus (Lee.)
Pityophthorus annectens Lee.
biovales Blkm.
JULY, 1925]
CHAMBERLIN SCOLYTID^
33
Pityophthorus briscoei Blkm.
canadensis Sw.
cariniceps Lee.
confinis Lee.
granulatus Sw.
lautus Eieh.
opaculus Lee.
patchi Blkm.
Pityophthorus puberulus Lee.
pullus (Zimm.)
puliearius (Zimm.)
ramiperda Sw.
Polygraphus rufipennis (Kby.)
Trypodendron bivittatum (Kby.)
Xyleborus impressus Eieh.
Xyleehinus amerieanus Blkm.
58. Pinus tceda.
Carphoborus bicristatus Chap.
Crypturgus pusillus Gyll.
Dendroctonus frontalis Zimm.
rufipennis (Kby.)
terebrans (Oliv.)
Gnathotriehus materiarius
(Fitch)
Ips avulsus (Eieh.)
calligraphus (Germ.)
grandicollis (Eieh.)
Orthotomicus caelatus (Eieh.)
Pityogenes plagiatus Lee.
Loblolly pine
Pityokteines sparsus (Lee.)
Pityophthorus annectens Lee.
Pityophthorus bisulcatus Eieh.
granulatus Sw.
lautus Eieh.
puliearius (Zimm.)
pullus (Zimm.)
Platypus flavicornis (Fab.)
Stephanoderes quadridentatus
Hopk.
Trypodendron scabricollis (Lee.)
Xyleborus pecanis Hopk.
59. Pinus torreyana. Torrey pine
Dendroctonus terebrans (Oliv.) Pityophthorus torreyanae Sw.
60. Pinus virginiana. Scrub pine
Dendroctonus frontalis (Zimm.) Trypodendron scabricollis (Lee.)
Gnathotriehus asperulus (Lee.)
61. PseudotsugcB macrocar pa. Big-cone spruce
Dendroctonus pseudotsugae Trypodendron ponderosae Sw.
Hopk. Xyleborus xylographus (Say)
Platypus wilsoni Sw.
62. PseudotsugcB taxifolia. Douglas fir
Cryphalus subconcentralis Hopk.
Dendroctonus pseudotsugae
Hopk.
Dryocoetes pseudotsugae Sw.
Gnathotriehus retusus (Lee.)
sulcatus (Lee.)
Hylastes nigrinus (Mann.)
ruber Sw.
Hylurgops grandicollis Sw.
Ips chamberlini Sw.
Pityokteines elegans Sw.
jasperi Sw.
minutus (Sw.)
Pityophthorus confinis Lee.
nitidulus (Mann.)
pseudotsugae Sw.
Platypus wilsoni Sw.
Pseudohylesinus grandis Sw.
elongatus Sw.
granulatus (Lee.)
34
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
Pseudohylesinus nebulosus (Lee.)
sericeus (Mann.)
Scolytus (E) monticolae (Sw.)
subscaber Lee.
tsugae (Sw.)
unispinosus Lee.
Seolytus ventralis Lee.
Trypodendron bivittatus (Kby.)
eavifrons (Mann.)
ponderosae Sw.
Trypodendron sp.
Xyleborus xylographus (Say)
63. Sequoia gigantea (IVashingtoniana) . Big tree
Gnathotriehus suleatus Lee. Platypus wilsoni Sw.
Phloeosinus punetatus Lee. Trypodendron sp.
rubieundulus Sw.
64. Sequoia sempervirens. Redwood
Gnathotriehus retusus (Lee.) Phloeosinus sequoiae Hopk.
suleatus (Lee.) Platypus wilsoni Sw.
Phloesinus eupressi Hopk. Trypodendron sp.
65. T axodium disticum. Bald cypress
Monarthrum mali (Fiteh) Playtypus eompositus Say
Phloeosinus taxodii Blkm.
66. Thuja occidentalis. White cedar
Gnathotriehus suleatus (Lee.)
67. Thuja plicata.
Gnathotriehus retusus (Lee.)
suleatus (Lee.)
Phloeosinus eupressi Hopk.
punetatus Lee.
sequoiae Hopk.
Phloeosinus eanadensis Sw.
dentatus Say
Western red cedar
Phloeosinus minutus Sw.
Platypus wilsoni Sw.
Trypodendron eavifrons (Mann.)
Xyleborinus n. sp.
68. Tsuga canadensis. Eastern hemlock
Anisandrus obesus (Lee.)
pyri (Peek)
Gnathotriehus suleatus (Lee.)
Ips longidens Sw.
Monarthrum faseiatum (Say)
Monarthrum mali (Fiteh)
Trypodendron bivittatum (Kby.)
Trypodendron sp.
Xyloterinus politus (Say)
69. Tsuga caroliniana. Southern hemlock
Anisandrus obesus (Lee.) Monarthrum faseiatum (Say)
pyri (Peek) Trypodendron bivittatum (Kby.)
Monarthrum mali (Fiteh)
JULY, 1925]
C H A M BERLI N S COL YTID JE
35
70. Tsuga heterophylla. Western hemlock
Gnathotrichus retusus (Lee.) Platypus wilsoni Sw.
sulcatus (Lee.) Pseudohylesinus serieeus (Mann.)
Hylastes nigrinus (Mann.) tsugge Sw.
71. Tsuga mertensiana. Mountain hemlock
Gnathotriehus retusus (Lee.) Seolytus tsugas (Sw.)
suleatus (Lee.)
A New Mirid From Arizona (Hemiptera)
BY E. P. VAN DUZEE
California Academy of Sciences, San Francisco
Mesomiris albescens Van Duzee, n. sp.
Whitish; two spots on pronotal base, scutellar, margins of
clavus and an elongated spot near apex of corium and tarsi
black ; antennae, bases and tips of tibiae reddish. Length, 7 mm.
Female: Head broader than in curtulus; median ineised line eon-
fined to vertex whieh is short; front broad, strongly eoneave, almost
tumid. Rostrum just passing middle of mesonotum. Antennae:
segment I as long as head viewed from above; II as long as from
tip of tylus to apex of seutellum; III two-thirds of II; IV four-
sevenths of III. Pronotum scarcely wider than long; median line
scarcely defined; punctures on collum and seutellum more shallow.
Elytra very obscurely punctured. Legs longer than in curtulus.
Color whitish, tinged with yellow on head and pronotum;
mesonotum and mesosternum croceus, marked with black posteriorly;
a large spot on base of pronotum either side; latero-basal edge of
seutellum, scutellar margin of clavus and an elongate discal spot near
apex of corium (sometimes becoming a faint vitta connecting with
the scutellar margin of clavus), black or blackish; antennae refuscent,
becoming darker at apex and paler on base of segment II and upper
disk of segment I; eyes dark castaneous; legs tinged with yellow,
apex of femora and base and apex of tibiae red; tarsi black as is the
extreme tip of the tibiae and the apex of rostrum.
Type : female. No. 1816, Mus. Calif. Acad. Sci., collected by
E. P. Van Duzee, August 8, 1924, along the Babocomari River
near Huachuca Station, Arizona. Paratype, female, same data.
36
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
ON NIGHT FLYING AND ATTRACTION TO LIGHT
IN ACRIDHDZE AND THE RELATION OF
METEOROLOGICAL CONDITIONS
THERETO
BY L. P. ROCKWOOD
U . S. Bureau of Entomology, Forest Grove, Oregon
Published records of night flying and attraction to light
in the family Acridiidse of the Orthoptera appear to be few
and not readily discoverable. This behavior has been well
established for Dissosteira longipennis Thomas in Colorado ^
and New Mexico ^ and a recent article ^ would indicate that
this species was the only one in the United States known to
fly at night. A search of the literature at hand at this
moment, however, adds several species to the list of those
taken at light at night, and the observation recorded in this
paper adds three more.
Riley, Packard, and Thomas ^ were apparently able to
find but one authentic case of observation of flight of
Acridiidse at night in European literature and mention none
for America. They pointed out, however, that circum-
stantial evidence indicated that migratory flights may be
continued after nightfall. A flight of large numbers of
grasshoppers, thought to have been Schist ocerca americana
Dru., was noted ^ at a lighthouse in Lake Erie off the mouth
of the Detroit River between the hours of 11 and 12 p. m.
on the night of July 12, 1899. Mr, W. R. Walton of the
U. S. Bureau of Entomology has stated in correspondence,
and given permission to quote that, in the vicinity of
Washington, D. C., “we frequently find considerable num-
bers of the Carolina grasshopper, Dissosteira Carolina Linn.,
flying around the electric lights during the warm nights of
the summer.”
1 Circular 36, 13th annual report, State Entomologist of Colorado,
p. 35, June, 1922. Fort Collins, Colo.
2 Bulletin 293, U. S. Department of Agriculture, 1915, Washington, D. C.
i3 Second Report, U. S. Entomological Commission, p. 97, 1880,
Washington, D. C.
4 Bulletin 22, n. s., Div. of Ent., U.S.D.A., p. 106, 1900, Washington, D.C.
JULY, 1925]
ROCKWOOD ACRIDIIDJE
37
Rehn and Hebard * have recorded the capture of
the following species at light at night in New Mexico and
Arizona :
Encoptolophus texensis Braner e, 1 2 , Deming, N. M., July 20, 1917 ;
Eoswell, N. M., Aug., 1902,
Derotmema laticinctum Scudder 6, 1 $ , Alamogordo, N. M., July 12, 1907.
Trimerotropis texana Bruner 1 $ , Alamogordo, IST. M., July 12, 1907.
rubripes Relin 6, 1 2 Alamogordo, N. M., July 12, 1907.
fvinculata Scudder®, Alamogordo, N. M., July 12, 1907.
vinculata Scudder 5^ Ig^ Nogales, Ariz., Aug, 13.
fascicula McNeill 1 2 j Nogales, Ariz., Aug. 13.
strenua McNeill s, 2 specs., Tucson, Ariz., July 26.
Aeoloplus ariBonensis Scudder 7 specs., Yuma, Ariz,, July 27, 28.
Melanoplus herbaceus Bruner ®, 1 2 j Alamogordo, N. M., July 12, 1907.
A study of these records indicates that night flying of
Acridiidse may frequently occur in the Southwestern United
States, while nocturnal activity in this family would seem
to be of rare occurrence in northern latitudes. It is, there-
fore, thought to be worth while to record the following
observation made by the late Prof. A. L. Lovett of the
Oregon Agricultural College and the writer, together with
pertinent meteorological data.
On the night of July 22, 1923, Professor Lovett and the
writer were in the city of Boise, Idaho, after a sweltering
day on the train. At about 9 :30 or 10 p. m., w^e decided
to take a short stroll in the open air for the purpose of
cooling off. Our attention was almost immediately attracted
to grasshoppers which were fluttering around the street
lamps like moths. Several of the grasshoppers were on the
sidewalk where many had been stepped on and mangled
by pedestrians and we each collected a double handful of
the hoppers from in front of show windows and doorways.
These grasshoppers have since been identified by the writer
as follows :
Conozoa ‘ivallula Scudder, 3 2 2
Dissosteira spurcata Sauss., 6 6 6,322
Spharagemon txquale Say, 5 6 5,522
5 Proc. Acad. Nat. Sci. Phila. p. 365, 1908.
6 Proc. Acad. Nat. Sci. Phila., p. Ill, 1909.
38
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
Clinton E. Norquest, Meteorologist of the Weather
Bureau, U. S. Department of Agriculture, Boise, Idaho, has
kindly furnished the following meteorological records for
July 22, 1923 ;
Temperature
Wind direction
^Relative humidity 7
89° at 8 p.m.
W at 7 p.m.
25 per cent at 6 p.m
86° at 9 p.m.
NW at 8 p.m.
80° at 10 p.m.
N at 9 p.m.
77° at 11p.m.
N at 10 p.m.
maximum for day, 98°
SE at 11 p.m.
It is the opinion of the writer that the high night tempera-
ture was the principal factor occasioning the night flight
of grasshoppers in this case, although the relative humidity,
wind direction, and the extreme heat of the preceding day-
light hours were doubtless also important factors. It is a
well known fact that temperature is a prime factor in deter-
mining the activity of grasshoppers. Sviridenko ® and
Parker ^ have recently reported that air temperatures in the
neighborhood of 80° F. occasion maximum activity in the
case of Dociostaurus maroccanus Thnb. and Camnula pellucida
Scudder, respectively. Parker ® has recorded much valuable
data on the relation of meteorological conditions to the
behavior of the latter species. In this paper it is stated that
migratory flights “did not begin (in Montana) until the
air temperature had reached 74° F. and were at their maxi-
mum at from 75° F. to 78° F.” Other data in the same
article indicated that the temperature limiting the inaugura-
tion of short flights in Minnesota is near 75° F. Both
authors agree that air temperatures in the neighborhood of
90° F. tend to inhibit activity in these species.
7 A trace of precipitation was recorded for the 23rd, so that the per-
centage of relative humidity was probably considerably higher at 10 p. m.
8 Sviridenko, P. A. Petrograd, 1924, rev. in Rev. of Applied Ent.,
vol. 12, p. 337, 1924.
9 Bulletin 214, U. of Minn. Agr. Exp. Sta., St. Paul, 1924, p. 32.
JULY, 1925] HUBBELL PRISTOCEUTHOPHILUS
39
A NEW SPECIES OF PRISTOCEUTHOPHILUS FROM
THE OLYMPIC MOUNTAINS, WASHINGTON.
(Orthoptera, Tettigoniidge) ^
BY THEODORE H. HUBBELL
Gainesville, Florida
The expedition to the Olympic Mountains of Washington,
sent out in the summer of 1919 by the Museum of Zoology
of the University of Michigan, brought back a small collec-
tion of the Orthoptera of the region. Only a few species
are represented, as the party left the field before most of
the Orthoptera had become mature. Nevertheless the col-
lection contains several species of the subfamily Rhaphi-
dophorinse which are of unusual interest; one of these
proved to be new, and is described below.
Pristoceuthophilus gaigei Hubbell n. sp.
Related to P. tuberculatus and P. cercalis, but easily dis-
tinguishable from all species of the genus hitherto described
by the distinctive armament of the dorsum of the male
abdomen, the form of the male cerci, and the coloration.
Description of holotype male
Size small for genus. Head with frontal spine well developed,
but shorter and blunter than in celatus and pacificus. Antennae slender,
with approximately 120 segments, more than twice as long as body.
Pronotum slightly longer than exposed portions of meso- and
metanota together. Ventral margins of thoracic terga gently arcuate,
with ventro-cephalic and ventro-caudal angles rather abruptly
rounded. Pronotum and mesonotum smooth; metanotum with a
group of small, blunt-pointed tubercles on each side of dorsum,
covering most of exposed lateral portions of tergum, but not reaching
ventral border.
Terga of abdominal segments I to VIII armed with large, heavy,
blunt-pointed spines, or with rounded or conical tubercles; dorsum
of segment IV elevated into a large median prominence of dis-
tinctive form. Spines and tubercles of each abdominal segment
arranged in two slightly irregular transverse rows along caudal
margins of terga, except on VIII, where cephalic row is absent.
On I to IV the spines (smallest on I) confined to lateral and dorso-
lateral portions of terga; dorso-lateral spines largest (some measure
.7 mm. in length), decreasing in size ventrad. Dorsum of segments
1 Contribution from the Department of Biology, University of Florida.
40
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
I to III unarmed. Tergum of III narrower dorsally than the rest,
crowded forward by enlarged dorsum of IV, which bears, in median
line, a large, tall, smooth protuberance of peculiar shape (see figure)
which is approximately three times as long as dorsal length of
segment III. Segment V armed similarly
to segments I to III, but caudal row of
spines continued across dorsum (the
dorsal spines being, however, much
smaller than those of sides), and cephalic
row represented on dorsum by paired
lateral spines and a row of conical
tubercles. Segment VI with same arrange-
ment as V, but dorsal spines of both rows
larger; spines of cephalic row irregularly
spaced, and only about one-half the height
of those of caudal row. Segment VII with
spines of caudal row reduced to about
half the size of corresponding ones of
VI; cephalic row reduced to an indistinct
line of rounded tubercles. Segment VIII
with cephalic row absent; caudal row re-
duced to a line of rounded tubercles.
On all abdominal segments except I the spines or tubercles of the
caudal tend to be larger than those of the cephalic row; this is most
marked in segments V to VIII, in the last of which no trace of the
cephalic row remains. On segments II and III the armature almost
reaches the ventral borders of the terga; posteriorly the distance to
the ventral margin increases, until on VIII the tubercles are confined
to the dorsum.
Supra-anal plate (tenth tergite) laterally narrow, with gently con-
cave latero-caudal margins, produced mesially in a strongly depressed
triangular lobe. Subgenital plate (hypandrium) subquadrate, nar-
rowing distally; distal margin broadly and shallowly emarginate,
disto-lateral angles rounded. Styles short, robust, bluntly pointed,
with faint indications of segmentation.. Penis fully extruded; ventral
lobe sub-chitinous, densely clothed with fine retrorsoappressed
spinules; longer than broad, with rounded tip (which hides distal
portion of supra-anal plate in type).
Cerci similar in form to those of cercalis and tuberculatus. The
length of cercus to tip of basal unsegniented portion about two-
thirds the length of the pronotum.
Legs moderately stout. Cephalic femora unarmed. Median femora
armed on both margins (left) or on caudal margin only (right) with
a very minute sub-apical spinule. Median tibiae armed dorsally with
2 The artist has correctly shown the main features of the abdominal
specialization, but the figure contains some minor inaccuracies with regard
to proportions and spine arrangement.
PristoceutliopJiilus gaigei
Hubbell.
Dorsal view of abdomen of
male holotype.^
JULY, 1925]
HUBBELL PRISTOCEUTHOPHILUS
41
3-4 (left) and 3-2 (right) small, slender spines on cephalic and
caudal margins respectively. Caudal femora slender, unarmed
dorsally; both ventral carinse armed in distal half with distant,
minute, sharp-pointed spinules, more numerous and closely spaced
on caudal than on cephalic margin, the apical ones larger than the
rest. Hind tibiae straight, with no trace of basal curvature or ventro-
proximal lamination.
Length of antennas approximately 21 mm.; of pronotum 2.6 mm.;
of cephalic femora 3.4 mm.; of caudal femora 7.0 mm.; of caudal
tibi^ 8.3 mm.; of cercus to tip of unsegmented basal portion 1.6 mm.
Coloration®: Antennae honey yellow; joints of distal portion in-
fuscated in proximal half; marked with lighter annuli at irregular
intervals. Head barium yellow, with indistinct markings of Vandyke
brown below eyes and antennal sockets, and on median frontal line.
Frontal spine black, the ocelli on its sides bright sulphur yellow.
Eyes black. Occiput barium yellow, with markings of vandyke
brown between eyes. Thorax barium yellow on dorsum, lighter
and semi-translucent along ventral margin. Two broad brownish
black dorso-lateral stripes border the light-colored median area.
They are most distinct on caudal portion of pronotum and on
mesonotum; on cephalic half of pronotum less densely colored, and
marked with small yellowish maculations. Light median stripe
abruptly narrowed in middle of pronotum by convergence of dark
bands cephalad; from this angulation faint longitudinal streaks of
Vandyke brown extend caudad within margins of median band.
Abdomen with dorsal surface ocher yellow, caudad faintly marked
with longitudinal brownish lines; spines and median protuberance
ochraceous tawny. The dark bands of thorax extend caudad less
distinctly along sides of abdomen. Cerci barium yellow proximally,
shading into reddish brown at distal end of unsegmented portion;
terminal segment of distal portion barium yellow. Caudal femora
honey yellow dorsally, indistinctly suffused with brownish on ventral
surface; outer ventral carina marked with distinct alternating light
and dark patches. Traces of color pattern similar to that of cercalis
visible on pagina. Remaining femora and tibia honej'' yellow, faintly
marked with brownish.
Specimens examined: One (holotype male), Lake Cush-
man, Olympic Mountains, Mason County, Washington,
August 1, 1919 (F. M. Gaige). (In collection of Museum of
Zoology, University of Michigan.) A single immature
female taken June 25, 1919, in the same locality shotvs a
similar type of coloration, and may belong to this species.
3 Where definite color names are used the terminologj* is that of
Ridgway’s Color Standards and Color Nomenclature.
42
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
Prist oceuthophilus gaigei is more bizarre than any species of
the genus hitherto described, and one of the most peculiar of
North American Orthoptera. It is most closely related to
P. tuberculatus and P. cercalis, but far surpasses both of these
species in degree of specialization of the male abdomen. All
three species have the same form of male cercus, a more
specialized type ^ than is found in other members of the
genus. P. tuberculatus approaches closer to gaigei in the
character of the male dorsal abdominal armament than does
cercalis.
The function (if it has one) of the peculiar median
prominence on the dorsum of the abdomen can only be sur-
mised. It may be noted, however, that on the concave
sides of the structure there are two elongate light-colored
areas, Avhich are connected across its dorsum at the nar-
rowest point; these areas are not heavily chitinized, but
seem to be membranous. It is possible that they may con-
tain the openings of scent glands, or perhaps mark the site
of an eversible scent organ.
I have named this species in honor of its collector, my
friend, Mr. F. M. Gaige. His field notes on the type are as
follows : “In rotten logs in clearing on Staircase Trail,
Aug. 1.” Mr. Gaige informs me that these logs were in an
apparently natural clearing in maple-alder woods, with
underbrush of stag-alder, vine-maple and willows, which
bordered the dense conifer forest, the edge of Vv^hich was
about sixty yards distant. The altitude was approximately
1050 feet.
4 These cerci are clearly of a more specialized type than the form
normal in the Rliaphidophorinse, hut E. M. Walker (1922, Ann. Ent. Soc.
Amer., xv, 19) suggests that the segmentation of the apical portion of
the cercus in this group of species is an atavistic character.
JULY, 1925]
SHANNON SPHECOMYIA
43
NORTH AMERICAN SPHECOMYIA
(DIPTERA, SYRPHIDiE)
BY RAYMOND C. SHANNON
Bureau of Entomology
The species of the genus Sphecomyia are remarkable for their
close resemblance to the yellow- jacket wasps, Vespa. The simi-
larity of N. vittata is extraordinary, for even the antennae are
greatly elongated, being quite equal to those of wasps; while
pattoni and dyari, which have small antennae, have black and
enlarged fore tarsi which are probably used to simulate long
antennae by extending them forewards. The species are appar-
ently confined to the North American Pacific Coast region,
except vittata, which appears to be of holarctic distribution.
Table of Species of Sphecomyia Latreille
Al. Second, third, and fourth tergites each with a single yellow
crossband, the posterior half of tergites black; fore legs
mostly yellow; width of third antennal joint equal to
length of first two (Ore., Wn., B. C.) nasica Osborn
A2. Second, third, and fourth tergites each with a pair of yellow
crossbands, sometimes partly fused.
Bl. Fore tibia except base and fore tarsi black; arista much
longer than antenna; mesonotum black.
Cl. Scutellum yellow pilose; third vein moderately
curved downward (Calif.) dyari, n. sp.
C2. Scutellum black pilose; third vein practically straight
(Ore., Ida.) pattoni Williston
B2. Fore legs almost entirely yellow; mesonotum with yel-
low markings.
Cl. Arista as long as antenna; mesonotum with lateral
markings (B. C., Wn., ld 2 i)\o) ....occidentalis Osburn
C2. Arista much shorter than antenna; mesonotum with
a pair of median, interrupted stripes.
Dl. Antenna greatly elongated; arista about one-
third the length of second joint (holarctic)
mttata Wiedemann
D2. Antenna moderately elongate, arista longer than
second antennal joint (Calif. Ore., B. C.)
bremcornis Osten Sacken
Sphecomyia dyari Shannon, n. sp.
Female. Closely related to pattoni, but differs in a number
of features as follows : General appearance noticeably more
robust ; head as broad as high (higher than broad in pattoni ) ;
antennal prominence less projecting; face slightly broader and
44
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
less excavated; antennae more elongate, the third joint distinctly
shorter than the first two, mesonotum more shining ; two yel-
lowish pollinose spots between the humeri ; discal crossvein less
oblique; third vein distinctly curved; yellow abdominal bands
noticeably broader, the interrupted ones with their inner ends
squared. Length, 15.5 mm.; wing, 13 mm.
Type-locality. Gold Lake Camp, Plumas County, California,
July 19, 1916 (H. G. Dyar).
Type. Cat. No. 27,834, U. S. N. M.
Named for Dr. H. G. Dyar, who has added numerous Dip-
tera to the National collection, taken during his countrywide
searches for Culicidae. This species may prove to be a race of
pattoni.
A Useful Bibliography
Parshley, Howard M. Bibliography of the North American
Hemiptera-Heteroptera. Smith College Fiftieth Anniversary Publi-
cation. Northampton, Mass. 252 pages.
Of the working tools of the systematic entomologist perhaps
nothing has more value than a good bibliography. In the book
cited above Dr. H. M. Parshley has given the hemipterist just
such a help and has earned the sincere gratitude that will be
fully accorded him. In addition to listing the works dealing
with the systematic side of the subject he has wisely included
papers on the biology of the group and many treating purely
of the economic aspect of the subject. The names of the
authors are arranged alphabetically, and under each author the
titles are in chronological order, with a letter added, where
necessary, so the Marks system of reference can be used if
desired.
In the publication of my Catalogue of the Hemiptera of
America North of Mexico, the very bulk of the volume made
it advisable for me to omit most references to the biologic and
economic literature included in the Henshaw Bibliography of
Economic Entomology and its continuations, and all bibliog-
raphy except the briefest possible reference to the serials and
books whose abbreviations appeared in the pages of the
catalogue. Dr. Parshley’s Bibliography now supplies this
omission very completely. No entomologist whose labors
touch the Heteroptera at all can afford to be without this
new bibliographical help. — E. P. Van Duzee.
JULY, 1925
ESSIG ECONOMIC NOTES
45
ECONOMIC NOTES
BY E, 0. ESSIG
University of California, Berkeley, California
European Earwig
Although the European earwig, Forficula auricularia (Linn,),
has been known to occur in certain parts of the city of Berkeley
for a number of years, the first real damage to growing plants
came to the attention of the writer during the month of June
of the current year. The injury consisted in the destruction of
flowering buds of carnations in great numbers. The earwigs
ascended the plants at night and ate out the bases of the bud,
in most cases destroying them completely. The presence of
the earwig is being watched with considerable concern, chiefly
because of the damage reported to flowers and horticultural
crops in Washington and Oregon. Inasmuch as the infestation
has been generally limited to areas near the Bay, where little
or no gardening is done, there has been no great opportunity
for serious damage. As the pest spreads, however, to the resi-
dential districts, it is not unlikely that more serious injury will
be reported.
The question of control and extermination has been discussed
with the officials of the State Department of Agriculture. An
agreement was reached a year ago, whereby an inspection of
the Bay district was decided upon, but lack of funds and inspec-
tors caused a delay in this procedure. Extermination is prob-
ably impossible, but it may be favorable to conduct a control
campaign to prevent the spread of the insect throughout the
State. — E. O. Essig.
Beet Army-Worm on Cotton
The beet army-worm, Laphygma exigua (Hubner) [L. Havi-
maculata (Harvey)], occurred in great numbers on several
thousands of acres of cotton in the vicinity of Bakersfield dur-
ing the month of May. Very serious damage was being done
to the young cotton plants, many fields of which were entirely
destroyed.
In addition to the beet army-worn there was also present in
considerable numbers the yellow-striped army-worm., Prodenia
prcefica Grote, which was particularly abundant along ditch
banks and roadways.
46
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
A conference of entomologists resulted in the recommenda-
tion of poison bran mash, which was first broadcasted and later
drilled into the rows with seed drills. This material gave com-
plete control, and the situation was cleared up almost immedi-
ately. Caterpillars were fed and adult moths reared which con-
formed in every way to the adults of the beet army-worm. —
E. O. Essig.
The Currant Moth on Peanuts
Large shipments of peanuts infested with the currant moth,
Ephestia elutella Hiibner, were repeatedly received at the San
Francisco port from China, during the fall of 1924. The out-
breaks were checked by fumigation, but litigation involving hun-
dreds of thousands of dollars grew out of the shipments. This
is another instance of a questionable practice causing a heavy
financial loss to both the shippers and carriers. The trade gener-
ally recognizes the danger of importing peanuts from the Orient
during the late summer, and it is the ignoring of such well-
founded customs, by those willing to take long chances, that
leads to the destruction of good food and the introduction into
this country of dangerous insects. — E. R. deOng.
The Alder Flea Beetle
The alder flea beetle, Haltica bimarginata Say, has been par-
ticularly abundant and destructive in various parts of Northern
California this summer. Near Middletown, California,, it de-
foliated all of the alders along the streams for a distance of ten
miles. — Edwin C. Van Dyke.
The Monterey Cypress Bark-Beetle
The Monterey cypress bark-beetle, Phlaeosinus cupressi
Hopk., has heavily attacked, and practically killed, a number
of Monterey cypresses on the University of California campus
this last summer. Old cypress logs piled in the neighborhood
enabled a large brood to develop which, with the beginning of
spring, attacked the trees, those which were weakened by
drought readily succumbing. This beetle is destructive only
when large broods have been enabled to develop by allowing
dead trees or timber to remain in the neighborhood of the living.
— Edwin C. Van Dyke.
THE PAN-PACIFIC ENTOMOLOGIST
Published quarterly by the Pacific Coast Entomological Society
in co-operation with the California Academy of Sciences
E. P. Van Duzee^ Editor F. E. Blaisdell^ M. D., Treasurer
The annual affiliation meeting of the Pacific Coast Ento-
mological Society was held at Portland, Oregon, June 17, 1925,
in connection with the meetings of the Pacific Division of the
A. A. A. S. In all, more than forty entomologists were in
attendance at this meeting, which proved to be an exceptionally
interesting one. There was a full program of papers, crowding
the morning and afternoon sessions. This meeting was followed
by a two days’ meeting of the Western Division of the Ameri-
can Association of Economic Entomologists, which was equally
well attended, with a full and interesting program.
A number of our entomologists from about the Bay region
have been in the field this summer, and the net results will be
a very considerable addition to our local collections. Dr. E. C.
Van Dyke and E. P. Van Duzee spent the ten days before the
Portland meetings, working along the Oregon coast from Wald-
port to Astoria, and after the meetings Dr. Van Dyke spent
another two weeks working on Mt. Hood, Oregon, and Mt.
Adams, Washington, and vicinity. Mr. C. L. Fox is spending
a month collecting about Pullman, Washington, and Moscow,
Idaho, and other nearby places. Dr. F. E. Blaisdell went by
auto to Seattle, Washington, and Vancouver Island, and Pro-
fessor G. F. Ferris is spending several months cruising along
the Pacific Coast of Mexico and Central America, with a small
party who will make many landings for field work in geology
and entomology. Dr. Ernest H. Nast spent a month col-
lecting neuropteroid insects in the Sierra, and Mr. J. O.
Martin made a short excursion to the Coast about Areata,
California, after coleopterous material. The material taken
by Mr. Van Duzee, Dr. Van Dyke, and much of that taken
by Mr. Fox and others, will enrich the museum of the Cali-
fornia Academy of Sciences.
Mr. A. J. Basinger of Riverside, California, has presented
to the entomological department of the California Academy of
Sciences a collection of over 2200 insects, including a very
valuable and most welcome series of Alaskan Diptera.
48
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 1
Mr. H. H. Keifer, Assistant Curator of Entomology at the
California Academy of Sciences, acting as entomologist on the
Academy’s Revillagigedos expedition, returned on June 12
with a fine lot of insects taken during the two months the expedi-
tion was in the field. This material is now being mounted, and
a fuller report on it will be included in the next number of this
Journal.
Mr. Walter M. Giffard of Honolulu has been a most welcome
visitor at the California Academy of Sciences this summer.
He and Mrs. Gififard spent two months in San Francisco, re-
turning home on June 24. As is his custom when here, Mr.
Giffard spent much time and effort advancing the interests of
the entomological department of the Academy. Mr. F. X. Wil-
liams of Honolulu has also been spending some time in San
Francisco and improving the opportunity to do some work on
the Hymenoptera in the collection of the Academy of Sciences.
Another welcome visitor at the Academy was Professor J. B.
Parker of Washington, D. C., who spent a part of July working
over the Bembicidae in the Academy collection. Our local
entomologists have also been favored with a brief visit by
Mr. George P. Engelhardt of the Brooklyn Museum, who
was making a hasty tour of the Pacific Coast region from
Oregon to Arizona.
The recent death of Mr. Ralph E. Dodge of Exeter, Cali-
fornia, is a real loss to Western entomology. Mr. Dodge
was an enthusiastic collector of the Lepidoptera, his special
interest being in the moths. He had been working in con-
junction with his father, Mr. E. A. Dodge of Santa Cruz,
California, in building up a collection of our California
moths which was rapidly becoming one of the best in the
State. Our Western entomologists will feel that their
branch of science has sustained a serious loss in the death
of young Mr. Dodge, and will unite in extending S 5 ^mpathy
to the bereaved family.
WARD’S NATURAL SCIENCE ESTABLISHMENT
College Avenue, Rochester, New York . .
We offer our best-made g'enuine Schmitt boxes, Exhibition
cases, Cabinets, Insect pins, Riker mounts. Insects and Insect
collections, ‘and all other supplies essential for the pursuit of the
study of entomology. Send for free supply catalogue. No. 41.
Eor Sale
PACIFIC COAST ENTOMOLOGICAL SOCIETY
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Address: Dr. F. E. Blaisdell, 1520 Lake Street,
San Francisco, California
From California Academy of Sciences, Proceeding's, Series 4
Williams, F. X., Butterflies and Hawk-Moths of the Galapagos
Islands. 33 pages, colored plate. 1911. 50 cents.
Hebard, Morgan, Dermaptera and Orthoptera (of the Galapagos
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Ferris, G. F., A Catalogue and Host List of the Anoplura. 84
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Essig, E. O., and Kuwana, S. I., Some Japanese Aphididse. 77
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Cole, F. R. and Lovett, A. L., An Annotated List of the Diptera
of Oregon. 147 Pages, Illust. 1921. $1.
Curran, C. H., Revision of the Pipiza Group of the Syrphidse
from North America. 48 pages, Illust. 1921. 60 cents.
Address: California Academy of Sciences,
San Francisco, California, U. S. A.
\
VoL II
October, 1925
No. 2
THE
Pan-Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
HERMS, ENTOMOLOGICAL OBSERVATIONS ON FANNING AND WASHINGTON
ISLANDS 49
CHAMBERLIN, NOTES ON SOME CENTIPEDS AND MILLIPEDS FROM UTAH . . 55
BARNES AND BENJAMIN, NOTES ON LOPHOCERAMICA ARTEGA BARNES . . 63
BARNES AND BENJAMIN, A NEW PYRALID FROM CALIFORNIA, WITH NOTES
ON DECATURIA PECTINALIS B, & MCD 64
VAN DYKE, NEW SPECIES OF CARABID^ IN THE SUBFAMILY HARPALIN^
CHIEFLY FROM NORTH AMERICA 65
%
BLAISDELL, REVISED CHECK-LIST OF THE SPECIES OF ELEODES INHABITING
AMERICA, NORTH OF MEXICO 77
BARNES AND BENJAMIN, THE HYPERBOREUS GROUP OF HEPIALUS ... 81
CHAMBERLIN, A NEW SPECIES OF LEPIDOSAPHES FROM CHINA .... 85
WACHTER, THE HATCHING OF THE EGGS OF PERIPSOCUS CALIFORNICUS BANKS 87
BLAISDELL, TFIOMAS LINCOLN CASEY 90
ESSIG, ECONOMIC NOTES 92
LARSON, FURTHER NOTES ON HUMAN SUFFERING CAUSED BY MITES ... 93
EDITORIAL 96
San Francisco, California
1925
THE PAN-PACIFIC ENTOMOLOGIST
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ENTOMOLOGIST
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The Pan-Pacific Entomologist
Vol. II, No. 2 October, 1925
ENTOMOLOGICAL OBSERVATIONS ON FANNING
AND WASHINGTON ISLANDS, TOGETHER
WITH GENERAL BIOLOGICAL NOTES
BY W. B, HERMS
University of California, Berkeley
The equatorial coral islands of the mid-Pacific Ocean offer
much of interest to the biologist. They are for the most part
unstudied biologically, being off the main routes of travel and
generally more or less difficult to approach. The rather isolated
group with which we are concerned in this paper consists of
the following small islands : Christmas, nearest the equator.
Fanning, Washington and Palmyra, extending from Lat. 1° 57'
to 5° 49' North and Long. 157° 27' to 162° 11' West. With
the exception of the latter, this group is governmentally a part
of the Gilbert and Ellice Islands (British), and the inhabitants
are mainly Gilbertese.
The group was visited by the writer during the summer of
1924, and the voyage was made on a copra vessel, a small
British motor schooner, under sail practically the entire dis-
tance from San Francisco to Fanning Island. The trip required
twenty-three days. The stay on the islands lasted about three
months, very largely on Fanning, affording an unusual oppor-
tunity for the study of its fauna and flora. On the return trip
the Hawaiian Islands were visited, adding much to the value
of the investigation because of the opportunity to compare cer-
tain material in hand with collections in Honolulu, both from
the Hawaiian Islands and the Society Islands, about midway
between which the islands under consideration are situated, i. e.,
separated from these groups by distances of from about 850 to
1200 miles.
Fanning Island is a typical coral atoll, consisting of a narrow
rim of land hardly three- fourths of a mile wide at the most
and enclosing a very beautiful lagoon, opening into the ocean
at three points, thus forming in reality a circle of three narrow
islands. The entire island is approximately ten and one-half
miles long by five and one-half miles wide, with an external
50
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
circumference of about thirty-one miles. It comprises about
8500 acres, of which over 3200 acres are in coconuts. Wash-
ington Island is about four and one-half miles in length by
one and one-half miles wide, with an external circumference of
about ten miles. This island encloses a shallow fresh-water
lake about two miles long, with an average width of about half
a mile. The highest elevation of either of these islands is not
over twelve feet. Since practically all of our collecting was
done on these two islands, a description of the other islands will
be omitted.
The total annual rainfall for Fanning Island for 1918 was
slightly over 102 inches, for 1923 it was slightly over 66 inches,
and for the first eight months of 1924 it was above 55 inches.
The temperature ranges from about 78° F. at night to about
85° F. at noon, with very little variation throughout the year,
except an occasional day when more than 90° F., or a night
when 70° F. may be reached. The annual mean temperature is
slightly in excess of 81° F. The humidity is seldom below 70.
As might be expected, the flora of these coral islands is very
meager in species, although the vegetation in some places pre-
sents tropical luxuriance, notably on Washington and Palmyra
Islands. The writer and an assistant collected forty-six out of
the fifty-four known species of plants from this group of
islands, only two small collections having been made previously.
Of this total number of species, eleven have been introduced
purposely, among them Cocos nucifera Linn,, Musa sapientum
Linn., Artocarpiis communis Forst., Carica papaya Linn., and
Hibiscus rosa-sinensis Linn. Fourteen species have been acci-
dentally introduced, among them Euphorbia hirta Linn., Eu-
phorbia prostrata Ait., Sida fallax Walp and V ernonia cinerea
Less.
According to Dean E. D. Merrill, who identified all the spe-
cies collected by the writer, about twenty-nine may safely be
considered as native species. Most of the bush, which in some
places forms almost impenetrable thickets, consists of such
native species as the Umbrella {T ournef ortia argentea Einn. f.),
which may reach the height of great trees, Buka {Pisonia
grandis R. Br.), also occasionally occurring as tall trees, and
Nashu (Sccevola kcenigi Krause). The screw pine {Pandanus
tectorius Sol.) is also a conspicuous part of the vegetation in
OCTOBER, 1925]
HERMS FANNING ISLANDS
51
some parts of the islands. The coconut, however, dominates all,
and is never out of sight.
With this rather poor flora as a basis for support, the num-
ber of species of insects dependent directly or indirectl)'' on
plant food is naturally not very great, but the paucity in the
number of species is generally well balanced by the abundance
of individuals in a given species, notably ants. The same thing
may be said relative to the fauna in general. The number of
species of native birds is relatively small, there being only thir-
teen species on these islands (eleven sea birds, and two land
birds) exclusive of migrants. The number of individuals is,
however, very great and at times the sky is literally clouded
with birds, almost wholly fish-eating. The noddy tern {Anous
stolidus) is most numerous, and three species of boobies (Sula
leucogastra, Sula dactylatra and Sula piscator) occur in abun-
dance, while the booby’s arch-enemy, the man-o’-war hawk or
frigate bird (Fregata minor palmerstoni is ever present. A rare
species of paraquet (Vinus kuhlii) , a most exquisite creature
is rapidly growing less in numbers, though still fairly abundant.
The very beautiful tropic bird or bo’s’n bird (two species,
Phcethon lepturus and PhcEthon rubricaiida) is fairly common
in this region.
The ground almost everywhere is full of crab holes, and as
one approaches these, the ludicrous omnivorous land crab (Car-
disoma obesum) sidles away and disappears in its burrow.
Hermit crabs (Ccrnobita rugosa) are exceedingly abundant and
pestiferous, apparently attacking almost anything. The so-called
coconut crab (Birgus latro) , a large and powerful species, is
rapidly becoming extinct, but may still be seen climbing coconut
trees at night, hiding in holes in the ground and cavities of
trees during the day. Two species each of gheckos and skinks,
usually measuring not over five or six inches in length, are
remarkably abundant, and their rather large, white eggs are
frequently encountered in pockets in the rough trunks of the
coconut trees and elsewhere.
Rats (Rpimys alexandrinus) are very numerous and a serious
pest to the coconut industry.
During the practically three months’ stay on these islands
much time was given to the study of coconut pests, particularly
the Tahiti coconut weevil (Diocalandra taitensis Guerin). Rela-
52
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
tively little time remained for much else, however, spare hours
were diligently spent in making collections. Although many
insects were collected and safely brought to Berkeley in pill-
boxes, the total number of species taken is quite disappointingly
small, namely, only about ninety.
These ninety species are distributed somewhat as follows,
with probable corrections after all the material has been care-
fully checked up:
Coleoptera, 31 species, included in the following families:
Staphylinidse (2), Hydrophilidas (1), Cleridae, (1), Nidituli-
dae (1), Cucujidae (2), Dermestidae (1), Cryptophagidae (1),
Ostomidae (1), Coccinellidae (1), Elateridae (3), Odomeridae
(4), Tenebrionidae (3), Scarabaeidae (1), Cerambycidae (1),
Anthribidae (1), Curculionidae (5), Scolytidae (2).
Diptera, 12 species, included in the following families : Culi-
cidas (2), Tipulidae (2), Dolichopodidae (2), Ortalidae (1),
Sarcophagidae (1), Muscidae (1), Calliphoridae (2), Hippo-
boscidae (1).
Hymenoptera, 10 species in the following families : Apidae
(1), Formicidae (7), Evaniidae (1), Chalcididae (1).
Hemiptera, 8 species in the following families : Pentatomidae
(1), Cydnidae (1), Miridae (1), Nabidae (1), Coccidae (4).
Orthoptera, 8 species in the following families : Blattidae (5),
Tettigoniidae (1), Gryllidae (1), Forficulidae (1).
Lepidoptera, 7 species in the following families: Noctuidae
(4), Sphingidae (1), Nymphalidae (2).
Isoptera, 2 species.
Mallophaga, 3 species.
Anoplura, 1 species.
Odonata, 3 species.
Thysanura, 2 species.
BUTTERFLIES
Although one specimen of the milkweed butterfly {Danaus
archippus) , taken on Fanning Island, appears in my collection,
the specimen having been given to my assistant, Mr. Kirby, by
a native, none was actually seen by either of us, and the pres-
ence of this one specimen is rather puzzling. On the other hand,
the beautiful Hypolimnas bolina Linn, occurs rather abundantly,
though restricted in range, in certain localities on Fanning. I
took all three color phases during my stay on this island. So
OCTOBER, 1925] HERMS FANNING ISLANDS
53
far as I know, the sole food plant of the larvae on this island
is Sida fallax Walp, an accidentally introduced weed called
“kaura” by the native Gilbertese.
The natives were greatly impressed by the fact that the writer
reared in his room the beautiful “tebuba” (butterfly) from the
ugly ‘‘worm” which feeds on the “kaura.” Although I do not
kpow how long the larval period is, the pupal period is quite
short, for example, two caterpillars pupated Sunday, June 8
(1924), between 8:45 a. m. and 9 a. m., and emerged early
Monday morning, June 16, giving a pupal period of eight days.
The type locality for this species is the Indo-Malayan region.
It occurs in Madagascar and western China.
MOSQUITOES
Mosquitoes (Aedes) were very abundant and annoying on
Washington Island where they breed primarily in the rain
receptacles, from which the water is taken for drinking and
general domestic purposes. The natives dig shallow water holes,
“temanipa,” in which mosquitoes (Culex) breed in enormous
numbers, particularly on Fanning, where these water holes are
of frequent occurrence.
HEAD LICE
Head lice {Pediculus capitis) are very abundant among the
native Gilbertese. These lice are distinctly brownish black in
color, as may be seen from the specimens taken from natives
on Fanning Island. The lice are considered a delicacy by the
natives. If it were not so disgusting, I might go into consider-
able detail to describe the instances in which I witnessed the
evidently joyous tete-a-tetes in which this form of delousing
was in progress.
TERMITES
Two species of termites were taken on Fanning Island,
namely, Kalotermes immigrans Snyder, a Hawaiian species,
and Cryptotermes hermsi sp. nov. (named by Kirby ^). Only
one nest of the former was found in which these insects were
tunneling in the hard wood of a prostrate log of Tournefortia,
the Umbrella tree. The second species was common and abun-
dant in decayed logs and stumps of coconut and Tournefortia.
Cryptotermes hermsi Kirby is a close relative of the Oriental
1 Kirby, Jr., Harold, 1925. Cryptotermes hermsi sp. nov., a termite
from Fanning- Island. University of California Publications in Zoology,
Vol. 26, No. 23, pp. 437-441. T-welve figures in text.
54
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
termites and is interesting because of its occurrence on Fan-
ning and its absence both from the Hawaiian Islands and Aus-
tralia, with which Fanning Island has maritime contact. This
species is said to differ markedly from Cryptotermes primus
Hill of Australia, and even more decidedly from Cryptotermes
piceatus Snyder of the Hawaiian Islands. It is closely related
to Cryptotermes kotoensis Oshima, an Oriental species.
BLISTERING BEETLES
Shortly after boarding the vessel which was to take me to
Fanning Island, I was told about a beetle which blisters on
contact, and is very common. Not long after my arrival I
learned from experience that this was the case. Myriads of
individuals of two species of Sessinia swarm about the newly
opened male flowers of the coconut, feeding on pollen. These
two species are Sessinia collaris (Shp.) and Sessinia decolor
Fairm., both of which are locally called coconut beetles. Both
of these species were readily attracted by a strong electric light,
which was led from the ship to the land for the purpose of
collecting insects. Sessinia decolor certainly causes severe blis-
tering if, for example, it is slapped when crawling on the
forearm while collecting specimens. A specimen came in con-
tact with my forearm on the evening of May 20 (1924) and
was rather lightly brushed away, but the following day a large
blister appeared, measuring about three-quarters of an inch in
length by one-quarter inch in breadth, considerably elevated
and filled with clear liquid. I opened the blister and applied
iodine. The blister did not cause any appreciable pain.
A more detailed study of the species of insects taken on these
islands is in progress and will be published shortly, together
with observations concerning their geographical distribution.
NOTE ON OPUNTIA INSECTS
Doctor H. G. Dyar has just indicated the identity of “Meli-
tara sp.,” page 5, Vol H, this journal. For this form the name,
holla Zeller, is available, but Doctor Dyar considers it advisable
to treat it as a variety of the Florida prodenialis Walker. The
Florida form is, then, Melitara prodenialis prodenialis Walker,
while the Texan “Melitara sp.” is Melitara prodenialis hollii
Zeller. — J. C. Hamlin.
OCTOBER, 1925]
CHAMBERLIN MYRIOPODA
55
NOTES ON SOME CENTIPEDS AND MILLIPEDS
FROM UTAH
BY RALPH V. CPIAMBERLIN
The centipeds and millipeds noticed in this paper were col-
lected mostly by the writer during the spring of 1924, the
greater portion of them coming from Washington County and
adjacent parts of southv/estern Utah. To these were added
specimens collected in the same region by Mr. V. M. Tanner,
to whom I wish here to express my thanks. In addition to the
material from southern Utah, the paper deals also with some
forms taken by myself farther north, chiefly in Salt Lake and
Cache Counties, Utah.
CHILOPODA
Oabius paiutus Chamberlin, sp. nov.
Brown to yellow, the head and posterior legs tending to have an
orange tinge. Antennas short; composed of twenty articles. Ocelli
five or six, in two series, the single ocellus enlarged. Ventral spines
of first legs, 0, 0, 0, 2, 1-0, 0, 1, 1 (2), 1. Dorsal spines of twelfth legs,
0, 0, 3, 1, 1. Dorsal spines of penult legs, 1, 0, 3, 1, 1; ventral, 0, 1,
2, 3, 2; claws, 2. Dorsal spines of anal legs 1, 0, 3, 1, 0; ventral,
0, 1, 3, 2, 0; claw single. None of the coxae laterally armed. Claw
of gonopods of female tripartite, but the outer lobe almost obliter-
ated, the claw for the most part appearing essentially bipartite, the
lobes short; basal spines, 2 + 2. Anal legs of male unmodified.
Length, up to 7 mm.
Locality. Utah: Parowan Canyon. May, 1924. Many
specimens.
Tidabius zionicus Chamberlin, sp. nov.
Agrees with To. honvillens'is Chamb. and differs from other species
of the genus in having the basal spines of the female gonopods
3 + 3. It differs from honmllensis in having only two claws on the
anal legs instead of three. Articles of antennae up to thirt 3 '', mostly
verj^ short. Ocelli few, in two series. Third article of first four
pairs of legs unarmed ventrally. Ventral spines of anal legs, 0, 1,
3, 2, 0. Dorsal spines of penult legs, 0, 0, 3, 1, 0. Length, up to
8.25 mm.
Locality. Utah: Zion National Park. May, 1924.
Lophobius ioganus Chamberlin, sp. nov.
Dorsum yellowish brown; head more orange anteriorly. Legs yel-
low, the caudal pair of an orange tinge. Ocelli eleven, in three series;
e. g., 1+2, 5, 3. Ventral spines of first legs, 0, 0, 2, 3, 2. Dorsal
spines of twelfth legs, 0, 0, 3, 1, 1. Ventral spines of penult legs,
0, 1, 3, 3, 2; dorsal, 1, 0, 3, 1, 1. Dorsal spines of anal legs 1, 0, 3,
1, 0; ventral, 0, 1, 3, 2, 0; the claw unarmed. Last three pairs of
56 THE PAN-PACIFIC ENTOMOLOGIST [VOL. II, NO. 2
coxae dorsally armed; only the last coxae laterally armed. In the
male the anal legs have the last joint furrowed along the mesal side,
as in the next species; the fourth joint is also similarly modified,
bearing a dorsal lobe at distal end, its mesal surface broadly fur-
rowed, also furrowed along ectal surface above. Length, 12 mm.
Locality. Logan Canyon, Cache County, Utah. One male
and two smaller females.
Nearly related to L. francisccE, but differing decidedly in
color, in being broader behind and thus more attenuated cepha-
lad, in having only the last coxae laterally armed, in having the
ventral spines of anal legs, 0, 1, 3, 2, 0 instead of 0, 1, 3, 2, 1, etc.
Lophobius franciscae Chamberlin, sp. nov.
Allied to L. collium (Chamberlin). It differs from that species
obviously, in being reddish or chestnut instead of yellow or orange.
It differs from that species also, in having the ventral spines of the
anal legs normally 0, 1, 3, 2, 1, instead of 0, 1, 3, 2, 0, though the
latter formula may occasionally hold; also in having only the last
two instead of the last three coxae laterally armed. Antennae nor-
mally with twenty articles, but varying up to twenty-six, the num-
ber tending to differ in such cases on the two sides. Basal spines
of female gonopods, 2 2 or 3 -b 3. In the anal legs of both sexes
the last article is furrowed along the mesal side. In the anal leg
of male the fourth article is conspicuously thickened and is broadly
furrowed or excavated on mesal side, the excavation limited above
by a ridge-like margin somewhat as in collium ; at the distal end,
above a conspicuous process or lobe. Length, 13 mm.
Locality. Cedar City, Utah. May, 1924.
Named for Mrs. Frances (Brown) Chamberlin.
PoKABius UTAHENSis (Chamberlin)
Localities. Utah: Bear Lake. Many females. Also Logan
Canyon. Many specimens, both males and females, but the
females much predominating.
A form abundant under damp leaves in shaded places along
canyon streams in the northern part of the State.
Lithobius forficatus Linne
Locality. Utah : San Pete County, Fairview. Many speci-
mens.
This introduced species has become common about settle-
ments in the more northern part of the State.
Gosibius arizonensis Chamberlin
Localities. Parowan and Cedar Canyons. May. Only males
secured.
The specimens agree closely with Arizona individuals, though
OCTOBER, 1925] CHAMBERLIN MYRIOPODA
57
apparently somewhat smaller. The spines of the anal legs in
one adult male 0, 1, 3, 3, 1 instead of 0, 1, 3, 2, 1,
Ethopolys parowanus Chamberlin, sp. nov.
Related to E. bipunctatus (Wood), occurring in northern Utah and
in Nevada, in having none of the dorsal plates produced. It is a
smaller form with shorter antennje. It differs in having only one
tooth ectad of the prosternal spine, the usual formula being from
1 — 4+4 — 1 to 1 — 6+6 — 1. It is also characterized by having the
claw of the female gonopods entire, or essentially so, instead of
tripartite; basal spines in adults 3, 3, more rarely 2 + 3 or 2 + 2.
Ventral spines of anal legs, 1, 1, 3, 2, 0 or 1, 1, 3, 2, 1, the claw single.
Ventral spines of penult legs, 1, 1, 3, 3, 2; dorsal spines, 1, 0, 3, 1, 1;
claws, three. Length, 21 mm.
Localities. Utah: Parowan; also Cedar City. April and
May. Many specimens.
Bothropolys permundus Chamberlin
Localities. Utah: Mill Creek Canyon, Logan Canyon, Bear
Lake.
Many specimens of this species were taken in each of these
localities. It is one of the most common chilopods of the Wah-
satch Mountains.
Kethops leioceps Chamberlin, sp. nov.
Differs from K. utahensis, the genotype, in wholly lacking both the
paired and the unpaired sulci on cephalic plate. First dorsal plate
similar, in having a cervical sulcus and paired sulci that branch and
together form a W behind the cervical sulcus. Last dorsal plate
wider at caudal than at anterior end, its caudal margin convexly
bowed out in the middle, not emarginate. Last ventral plate strongly
narrowed caudad. Coxopleurae with caudal process short; scattered
short spines. The anal legs in general form as in utahensis; the
joints broad but not furrowed ventrally; femur with a median ven-
tral area free from spines, toward mesal side of ventral surface a
series of four stout spines or teeth. Tibia also with a series of four
ventral teeth. Metatarsus with a ventral series of close-set teeth,
not with an edge excised at middle as in utahensis. Length, 24 mm.
Locality. Utah: Mill Creek Canyon. One specimen.
Theatops posticus (Say)
Locality. Utah: Washington County. May. One specimen.
This species is very common in the Southeastern States, but
has not been known previously from the West.
Otocryptops sexspinosus (Say)
Locality. Utah: Zion National Park. May. One young
specimen.
58
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
There is little doubt that the specimen is correctly referred,
although the margiiiation of the head is obscure. The species
is common, both in the Eastern and Southeastern States and on
the Pacific Coast.
SCOLOPENDRA POLYMORPHA Wood
Localities. Utah : Washington, St. George and Zion National
Park. April and May, 1924.
This species ranges northward in the State as far as Salt
Lake City. The Utah specimens are dwarfed, in comparison
with those found in Arizona and southern California.
Scolopendra utahana Chamberlin, sp. nov.
Head without longitudinal sulci, smooth and shining, obscurely
finely punctate. Antennas short; articles twenty, of which only the
first four are nearly glabrous. First dorsal plate with a strongly
marked cervical sulcus like that of S. polymorpha; without longitu-
dinal sulci. Last dorsal plate short and broad; with a fine median
longitudinal impressed line. Dental plates of prosternum each with
four teeth, of which the three innermost are united at base. Anal
legs short; femur short and thick with about fifteen seriate spines
ventrally, none on mesal surface except the series of five teeth along
dorso-mesal edge; process at dorso-distal corner bearing only two
teeth, which are long. Length, 45 mm.
Locality. Utah: Zion National Park. May.
Resembles S. polymorpha Wood, but readily distinguished,
in having only the first four articles of antennae glabrous instead
of eight or more, in having the distal process of femora of anal
legs with only two spines, and in having the mesal surface of
femora free from spines.
Nyctunguis molinor Chamberlin, sp. nov.
Body yellow, near orange, the middle and posterior segments also
of an orange or red tinge. The cephalic plate longer than wide,
longer than in liber colens, a species it resembles; widest in front of
middle from where more strongly narrowed caudad than cephalad;
anterior margin forming an obtuse angle. Prebasal plate exposed.
All joints of prehensors unarmed; claws when closed failing con-
siderably of attaining the front margin of head. Labrum wide, some-
what as in libercolens, but the median arc deeper and shorter, the
teeth proportionately larger and fewer, t 3 ^pically ten in number. Ven-
tral pores present on anterior plates, forming a small circular area
on each plate. Last ventral plate broadly trapeziform, not strongly
narrowed caudad. The coxal pores two on each side, homogenous,
the anterior one covered by last ventral plate, the posterior one
partly covered; posterior gland much larger than the anterior one.
Pairs of legs, 47 (6), 49 (2). Length, 17 mm.
OCTOBER, 1925]
CHAMBERLIN MYRIOPODA
59
Locality. Utah : At mouth o£ Mill Creek Canyon. One
male and one female. May.
Geophilus shoshoneus Chamberlin, sp. nov.
A species resembling the Eastern G. mordax, but a much smaller
form, easily distinguished in not having the prebasal plate exposed,
the cephalic plate extending well over it. Claws of prehensors, when
closed, attaining anterior margin of head. Prosternum and joints of
prehensors unarmed. Chitinous lines present on prosternum, fine.
Antennae short. Ventral pores of sternites numerous, forming a
transverse band broadest at middle in front of posterior margin.
Last ventral plate wide, the posterior margin a little indented at
middle. Coxal pores four or five on each side close to margin of
plate, with no isolated single pore such as is characteristic of mordax.
Pairs of legs, 55-57. Length, 20 mm.
Locality. Utah : Logan Canyon and on the divide between
this canyon and Bear Lake Valley. Two specimens.
Geophilus rubens Say
Locality. Utah: Washington County. Three specimens.
May.
A species also met with frequently in Arizona. The Utah
specimens agree with those from Arizona, in usually lacking a
geminate black stripe along dorsum.
Linot^nia chionophila (Wood)
? Geophilus acuminatus Leach, Trans. Linn. Soc. Lond., 1814, 11,
p. 386.
Strigamia chionophila Wood, Journ. Phil. Acad. Sci., 1862, 5, p. 50.
LinotcEuia miuropus Chamberlin, Amer. Nat., 1902.
Scolioplanes acuminatus Attenis, Arkiv for Zoologi, 1909, 5, No. 3,
p. 25.
Linotcenia chionophila Chamberlin, Can. Ent., 1911, p. 260.
Localities. Utah: Parowan, one male; Trout Creek, two
specimens ; Bear Lake, one male ; Logan, many specimens.
All specimens examined have thirty-nine or forty-one pairs of
legs, excepting one specimen from Logan Canyon, which seems
also variant in other respects.
Gnathomerium xenoporus (Chamberlin)
Localities. Utah : Parowan, Cedar City, Mill Creek Canyon,
Logan Canyon. Many specimens.
This is evidently the most common geophilid in the canyons
of Utah, where it is often abundant in leaf mold along streams.
It occurs also in Colorado and New Mexico.
60
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
DIPLOPODA
Parajulus canadensis (Newport)
Locality, Utah: Mill Creek Canyon, April. Two males.
This is a common species in eastern Canada and adjacent
parts of the United States, but it has not previously been found
west of the Rocky Mountains.
Parajulus tiganus Chamberlin
Localities. Utah: Mill Creek Canyon, Provo Canyon, Logan
Canyon, and Bear Lake. Many specimens.
This is probably the commonest diplopod in the State. It has
not been found as far south as Cedar City, however, where it
seems to be largely replaced by P. paiutus, a new species here
described.
Parajulus paiutus Chamberlin, sp. nov.
This is a smaller form than P. tiganus. The color is brownish with
a row of black spots along each side, corresponding to the repug-
natorial glands, and a median dorsal longitudinal black line. A gemi-
nate lighter band across dorsum between the lateral spots on each
segment. Last segment darker, blackish. Legs yellow. Antennae
blackish. Distocaudal corner of cardo of mandible in male conspicu-
ously produced, not excavated, the angle narrowly rounded. First
legs thickened as usual, but less so than in tiganus. Gonopods of
male of very different form from those of tiganus, being more like
those of P. nigrans. Both lobes of anterior pair short; posterior lobe
glabrous, narrower than anterior, a little bent forward distally. Pos-
terior gonopods narrowed distally, a little curved caudad distally,
the seminiferous duct opening at the caudally directed tip, much as
in nigrans. Number of segments, 54. Length, male, 27 mm.
Localities. Utah: Parowan, nine specimens, in part not
adult, taken in canyon above the town; Cedar City, four speci-
mens also taken in canyon above town; and Zion National Park,
one female.
Tylobolus utahensis Chamberlin, sp. nov.
A form more slender than usual in the genus, the length being
ten times the thickness. Brown in color, the metazonites darker
brown, with a tendency toward chestnut, the ectal caudal border
paler. Repugnatorial glands showing as blackish spots on the sides,
especially in the posterior region. Legs brown. Collum much nar-
rowed at each end. Coxse of third legs in male with the usual uncate
processes, the immediately following coxae also produced. Median
plate of anterior gonopods of male distally pointed, much exceeded
by the coxal plates, which are contiguous at middle and each of
which at its mesal corner is produced ventrad. The telopodite is
OCTOBER, 1925]
CHAMBERLIN MYRIOPODA
61
somewhat enlarged at distal (mesal) end, but is only a little bent
caudad at the end, not uncate. In the posterior gonopods the ter-
minal hook or blade is smooth; at base of hook in the angle is a
stout angular projection or tooth; the bent portion proximad of
blade finely scabrous. Number of segments, 51-53. Length, 54 mm.;
width, 5.2 mm. Female allotype, 7 mm. thick.
Locality. Utah: Zion National Park. Five adult males,
two adult females, and two young specimens,
Nannole uta (Chamberlin)
Locality, Utah : Little Willow Canyon.
Diploiulus luscus (Meinert)
Locality. Utah: Mill Creek.
Many specimens were taken on cultivated ground below
mouth of the canyon. The species is now not uncommon in
other places near Salt Lake City. A native of Europe, it is
frequently brought to this country in soil about the roots of
plants from Holland and other European countries. It long
ago became established in the Eastern States and has now
extended its range to California. It occurs only in settled or
cultivated districts.
Scytonotus orthodox Chamberlin, sp. nov.
General color brown, in part of a weakly reddish tinge. Legs yel-
low. The second tergite bearing four well-developed series of tuber-
cles; the anterior corners of keels produced along ends of collum,
acute; lateral crenuations weak, with a translucent margin. In ter-
gites of the middle region of body a transverse sulcus distinctly
marked, behind the sulcus with three or in part four rows of well-
developed tubercles, but in front of the sulcus the tubercles are low
and are in part obliterated. Prozonites nearly smooth, the granules
numerous, but very small or obscure. Distinguishable in structure
of the male gonopods; the two blades equal in length, rather slender,
the posterior one bidentate apically on caudal side, the teeth acute;
anterior branch apically acute, with caudal margin straight and an-
terior convex. Gonopods very similar to those of S. amandus; but
in the latter the branches are unequal, the dentate one being obvi-
ously shorter and its teeth closer together.
Localities. Utah : Logan Canyon, five adult males and three
partly grown specimens; Bear Lake, about eighteen specimens,
male and female, with some immature.
The young specimens do not show any reduction of the tuber-
cles in front of the suture.
Spirostrephon utorum Chamberlin, sp. nov.
The color is light yellowish brown without distinct markings. Be-
62
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
tween the median keel and the thickened porigerous keel on each
side of a typical segment there are three major setigerous keels, with
a much lower non-setigerous keel between each two major ones;
the porigerous keel much higher than these more mesal keels and
its margin conspicuously convex in profile instead of nearly straight;
below each porigerous keel two somewhat similarly high setigerous
keels with a low, non-setigerous keel between these and one above
the upper one; below on the side other non-setigerous keels which
become lower in going ventrad. The major keels in profile are seen
to be serrate or dentate. Number of segments, sixty.
Locality. Utah: Green River. One male. R. V. Chamber-
lin, Jr., collection 1920.
Apparently related to S. mutans of Southern California; but
aside from more technical structural differences, it is conspicu-
ously distinct in coloration and in the greater number of seg-
ments to the body.
Tingupa utahensis australis Chamberlin, subsp. nov.
In the absence of males a wholly certain determination of the
position of this form is not possible. It agrees in general structure,
the form and sculpturing of the tergites, etc., with typical utahensis
from the northern part of the State, but it is a larger and more
robust form. Color brown, the caudal border of metazonites paler,
as is the prozonite in the constricting furrow adjacent to metazonite.
Sixth and seventh articles of antennse equal in length, the fifth much
longer and distally slightly thicker than the sixth. The eye-patch is
differently shaped, being trapeziform rather than triangular, with the
dorsal side, rather than that along caudal margin, longest; ocelli in
the holotype in six or seven vertical series instead of in four; e. g.,
beginning with most caudal row, 5, 4, 3, 2, 2, 1; and in four or five
transverse, somewhat curving, series, e. g., beginning at top, 1, 7,
5, 3, 1. Length, about 9 mm.
Locality. Cedar City, Utah. One female taken in the can-
yon. May.
Underwoodia tida Chamberlin, sp. nov.
A much smaller form than hespera, differing superficially also in
being distinctly paler in color on the sides below level of pores than
on dorsum; finely mottled. Legs dusky yellow. Antennae of same
color as legs. Head dark above in frontal region with numerous fine,
pale dots, clypeus yellow. Sixth joint of anteniiEe decidedly thicker
and longer than the seventh. Ocelli in a triangular patch with apex
mesad; thirteen in number, arranged in three or four oblique series,
e. g., 6, 5, 2 or 1, 5, 5, 2. Collum not distinctly margined below.
Length, 6 mm.
Locality. Utah: Logan Canyon.
OCTOBER, 1925]
CHAMBERLIN MYRIOPODA
63
Underwoodia hespera Chamberlin, sp. nov.
The body in general is brown, with the nietazonites darker, light
spots on each side indistinct. Head dark, black, or nearly so. Legs
dusky over yellow or light brown, the antennae blackish. In the
antennae the sixth article is narrower than the fifth and is obviously
and considerably longer than the seventh instead of being equal to
it, as is the rule in species of Cleidogona. Ocelli distinct; eighteen,
in five somewhat irregular longitudinal series, as follows: 1, 2, 4, 5, 6;
the patch triangular in outline, with the apex mesad in position.
Body strongly narrowed anteriorly. Collum acutely narrowed down
each side, margined anteriorly on each side, otherwise smooth. Seg-
ments in general smooth, not striate above, under lens seen to be
evenly marked with areas defined by fine lines, the areas mostly from
quadrate to oblong in form. Length, 8 mm.
Locality. Utah: Mill Creek Canyon. One female.
POLYDESMUS MONILICORNIS Koch
Locality. Mill Creek. One immature specimen, apparently
this form, which is common in the Eastern States and Canada.
NOTES ON LOPHOCERAMICA ARTEGA BARNES
(LEPID., PHALAENIDiE)
BY WILLIAM BARNES AND F. H. BENJAMIN
Decatur, Illinois
Lophoceramica ARTEGA Bai'iies
1907, Barnes, Can. Ent., XXXIX, 64, Tricholita.
1908, Dyar, Proc. Ent. Soc. Wash., X, 32, Lophoceramica.
1912, Barnes & McDunnough, Contrib. N. H. Lep. N. A., I, (4), 21,
pi. IX, f. 20; p. 55, pi. XXVI, f. 11, Lophoceramica.
pallicauda Sm.
1908, Smith, Ann. N. Y. Acad. Sci., XVIII (2), 101, Mamestra.
Lophoceramica artega form eriopygoides nov.
artega Auct. (nec Barnes).
1912, Barnes & McDunnough, Contrib. N. H'. Lep. N. A., I, (4),
21, pi. IX, f. 19, p. 55, pi. XXVI, f. 10, Lophoceramica.
The form of artega lacking, or practically lacking, the white
on the reniform appears to be causing some confusion, so we
designate it as ej'iopygoides. It is a parallel development to
Chabuata endiva form vespera.
Type localities and number and sexes of types: Holotype 3 ,
Paradise, Cochise County, Arizona, 1-7 July; allotype 2, id.,
1-7 June; 5 3 2 2; paratypes, Paradise, Baboquivari Moun-
tains, Palmerlee, and Patagonia, Arizona, June, August, Sep-
tember.
Note — In Barnes collection ; a single female paratype in Cali-
fornia Academy of Sciences, having been submitted for deter-
mination by Mr. E. P. Van Duzee.
64
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
A NEW PYRALID FROM CALIFORNIA, WITH
NOTES ON DECATURIA PECTINALIS
B. & McD. (LEPID., PYRALIDiE,
MACROTHECIN^)
BY WILLIAM BARNES AND F. H. BENJAMIN
Decatur, Illinois
Macrotheca lecerfialis Barnes and Benjamin, sp. nov.
Ground color white; fore wing with a few scattered black scales;
t. a. line black, single, waved from costa to inner margin; orbicular
represented by a few black scales; reniform obsolete; t. p. line single,
black, nearly erect, little waved, a trace of a blackish s. t. suffusion
subapically; a terminal row of dots; fringe white, with a few black
scales; hind wing only slightly darker than fore wing; discal spot
and median shade obsolete; an obscure terminal line; fringe luteous-
white basally, distally white. Expanse, 9-11 mm.
Allied to hilinealis and flexinealis. Lack of reniform will
easily sort it from the latter, while its much paler coloring and
more erect t. p. line will easily distinguish it from the former.
Type locality : Dixieland, Imperial County, California.
Number and sexes of types : Holotype 6 , 15-30 April, 1922 ;
allotype 2, 1-15 May, 1922; 8 6 paratypes, 15-30 April, 1922.
Note — We take pleasure in naming this species in favor of
M. LeCerf, who extended every courtesy to the senior author
during his stay in Paris.
DeCATURIA PECTINALIS B. & McD.
1912, Barnes & McDunnough, Contrib. N. H. Lep. N. A., I (5), 41,
pi. Ill, f. 8, Decaturia.
In the original description of the genus and species the
antennae of the female is described as being simple. A series
of females before us, including the type 2 and four 2
‘‘Cotypes,” proves this to be incorrect. The antennae are decid-
edly strongly lamellate, as that term is applied in Lepidoptera :
viz., dentate.
The type series bore no dates. Specimens have been received
from the Baboquivari Mountains, Pima County, Arizona, April,
May, June, September, and October; and from Clark County,
Nevada, June.
OCTOBER, 1925]
VAN DYKE HARPALIN^
65
NEW SPECIES OF CARABIDPE IN THE SUBFAMILY
HARPALINiE, CHIEFLY FROM WESTERN
NORTH AMERICA
BY EDWIN C. VAN DYKE
University of California, Berkeley, California
Bembidium ccerulescens Van Dyke, new species
Form moderately elongate, gracefully proportioned, slightly con-
vex, shining, nigropiceous with a bluish luster above, especially on
head and prothorax, the antennae, palpi, tibiae, and tarsi brownish.
Head scarcely narrower than the prothorax at apex, alutaceous, e 3 '^es
moderately prominent, antennae slender, about one-half length of
body, frontal grooves single, straight and shallow, mandibles promi-
nent. Prothorax as wide as long, three-fourths as wide as elytra at
base, narrower at base than apex; apex feebly emarginate; sides with
the margins distinctly reflexed, moderately arcuate in front, grad-
ually narrowed and sinuate posteriorly; hind angles slightly obtuse
and not carinate; disc alutaceous with median longitudinal groove
well impressed and abbreviated in front; anterior transverse impres-
sion well defined; area in front of it with a number of longitudinal
rugae; basal impressions broad, deep, indistinctly bistriate, the entire
basal area rugose. Elytra elliptical, three-fifths as broad as long;
humeri distinctly rounded; sides slightly arcuate anteriorly, more
decidedly so posteriorly and but slightly sinuate in front of apex;
disc finely striate, the striae entire and with very vague punctures
anteriorlj'-, the outer ones less deeply impressed, the fifth terminat-
ing in a deep groove; third stria with first dorsal puncture slightly
in front of the middle and second about one-fourth distant from
apex; eighth stria at border of marginal groove, the intervals flat.
Length, 7 mm.; breadth, 3 mm.
Holotype, male (No. 1818, Mus. Calif. Acad. Sci.), in my
collection, kindly presented to me by Mr. Merton C. Lane, who
secured it near the margin of the stream in Niles Canon, Ala-
meda County, California, December 27, 1919. Two paratypes
captured at the same time are in Mr. Lane’s collection.
This very graceful species looks not unlike a small specimen
of Platynus funehris Lee. According to the arrangement of
Roland Hayward,’- it would have to be placed in his Group VH
(planatum) , and close to funerenm Lee., though it is very much
larger, almost as large, in fact, as planatum Lee. It resembles
nigrocoeruleum Hyd. only in color. It is also a more elegantly
formed insect than any other in the group.
1 “On the Species of Bembidium of America North of Mexico,” by
Roland Hayward. Trans. Am. IDnt. Soc., Vol. XXIV (1897), pp. 32-143.
66
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
Bembidium utahensis Van Dyke, new species
Form elongate, depressed, color dark rufous. Head large, but
slightly narrower than the prothorax; eyes small and much flattened,
hardly extending beyond the sides of the head; antennas stout, reach-
ing but two segments beyond the humeri; frontal impressions single,
shallow and lunate; mandibles prominent, porrect; palpi rufocasta-
neous, the terminal joint of the maxillary palpi at least twice as long
as the similar joint in Bembidium laticeps Lee. Prothorax almost a
third broader than long, narrower than elytra at humeri; apex feebly
emarginate; base just perceptibly arcuate; sides moderately arcuate
in front, sinuate posteriorly and parallel for a short distance in front
of base; hind angles right; lateral margin narrow; disc barely con-
vex; median longitudinal groove well impressed, especially pos-
teriorly, and almost completely; anterior and posterior transverse
impressions obsolete; basal impressions deep, with double striae
vaguely defined, most of the basal area somewhat rugose. Elytra
elliptical, somewhat more than a third longer than broad; humeri
rounded but with prominent basal ridge running obliquely inwards,
the margin fine; disc somewhat flattened, all striae present, well
impressed near suture, less defined laterally and apically, and finely
punctate; third stria with two dorsal punctures, the first slightly in
front of the middle, the second somewhat more than a fourth of the
distance from apex; eighth stria in marginal groove. Length, 4.5
mm.; breadth, 2 mm.
Holotype, male (No. 1819, Mus. Calif. Acad. Sci.), in my
collection, secured near Salt Lake City, Utah, by Professor
R. P. Chamberlin, and kindly presented to me by Dr. William
M. Mann. A second specimen was in the collection of Dr.
Mann.
This beetle has its closest relative in Bembidium laticeps Lee.
and belongs with that and Bembidium puritanum Hyd. It dif-
fers from the former by being larger with the prothorax more
elongate and less broad, the elytra also more elongate, with all
of the striae well defined and finely punctured, whereas in the
other the striae are but vaguely defined. It is interesting to find
the third member of this small group near Great Salt Lake,
laticeps being found near the seashore in southern California
and puritanum in Massachusetts.
Trechus ruficollis Van Dyke, new species
Body rather stout and somewhat convex, shining, rufopiceous;
head and prothorax rufous; antennae, palpi, legs and epipleurae cas-
taneous. Head more than three-fourths as wide as prothorax; eyes
moderately prominent; frontal sulci long, deep and semi-circular;
antennae moderately slender, segments definitely broadened out-
OCTOBER, 1925]
VAN DYKE HARPALINyE
67
wardly, extending to second fourth of elytra. Prothorax small, not
quite twice as wide as long; sides moderately arcuate anteriorly,
sinuate posteriorly and straight and parallel near base; hind angles
prominent and right, the lateral margin moderately reflexed; disc
with the median longitudinal groove finely impressed, the anterior
transverse impression vague, the posterior pronounced even at
middle, the basal impressions moderately deep and diffuse. Elytra
oval, somewhat truncate at base, almost twice as wide as prothorax
and two-thirds as wide as long; disc with the three inner striae very
fine, though observable and complete and with slight indications of
a fourth stria; third stria with three foveae, the first one-fifth dis-
tant from base, the second at about the middle, and the third close
to the apex. Length, 3.5 mm.; breadth, 1.75 mm.
Holotype and three paratypes, bearing the label, Lawrence,
Massachusetts, found among a series of unplaced specimens in
the entomological collection of Cornell University. They were
part of the Crew collection. The holotype and one paratype
will be deposited at Cornell University, and the two remaining
paratypes will remain with me.
This species belongs in the series with arisonce Casey, hydro-
picus Horn, ovipennis Mots., and pomonce Fall, agreeing with
all in the general form and degree of smoothness. It, however,
differs from all by having shorter and heavier antennae, the pro^
thorax more decidedly sinuate posteriorly, approached slightly
in this regard by ai'isonce Casey, the hind angles more promi-
nent, and by being bicolored, the rufous prothorax, in all speci-
mens seen, strongly contrasting with the piceous elytra. It is
possible that, in spite of the definite locality label, these speci-
mens may have come from some other locality, for it seems
strange that such a well-marked species should occur in Massa-
chusetts and remain unknown so long. It does not seem to
agree with any of the European species and, as stated, bears
a strong resemblance to certain American species.
Patroboidea Van Dyke, new genus
Body very elongate, parallel and flattened. Head large, elongate,
gense prominent and long, reaching from the posterior margin of
the eyes to the antenna, head constricted posteriorly, with deep and
broad frontal grooves and two supra-orbital seta, one rather incon-
spicuous. Antenna long, reaching to posterior third of body, moder-
ately stout, arising under a feeble frontal ridge; first and second seg-
ments glabrous, the following fairly pilose; first robust, second small
and transverse, third long, as long as the two following united and
about five times as long as the second; segments three to six of
68
THE PAN-PACIFIC ENTOMOLOGIST [VOL. II, NO. 2
about equal length and breadth, seven to ten also of about equal
length and breadth, but somewhat shorter than the preceding, the
eleventh about as long as the third. Eyes poorly developed, pro-
jecting but little beyond sides of head, well removed from the mouth.
Clypeus well developed, transversely depressed in front and with
lateral setae. Labrum short, broadly shallowly eniarginate and pluri-
setose in front. Mandibles prominent, prognathous, feebly arcuate,
acute and curved at tip, with setigerous puncture in scrobe. Maxil-
lae slender, acute at tip, ciliate with stiff hairs on inner side, the
outer lobe biarticulate; palpi rather long, the terminal segment grad-
ually tapering to apex, slightly longer than preceding, the point
subacute; penultimate not hairy. Submentum broad, its basal suture
distinct, deeply emarginate and with broad truncate lobe at middle;
epilobes dentiform; ligula moderately prominent, rounded and bise-
tose at apex; paraglossse broad, rounded at apex and very little
longer than the ligula; palpi of moderate length, the terminal seg-
ment conical, somewhat blunt at apex and slightly longer than
preceding segment.
Prothorax flattened, with side margin narrow and with setigerous
punctures near the front and at hind angles. Body not pedunculate,
scutellum distinct. Elytra flattened, elongated, more or less ellipti-
cal, not margined at base; sides narrowly inflexed; margin pos-
teriorly entire and without internal plica; disc with nine complete
striae, counting the marginal; first and second and the third and
fourth united at the apex; third with three dorsal punctures, the first
one-fourth distant from base, the second at about the middle, and
the third one-fourth distant from apex. Prosternum moderately sep-
arating the coxae and declivous at apex. Mesosternum narrowly
separating the coxae. Metasternum long, the epimeron short, the
posterior coxae contiguous. Anterior tibia of the male somewhat
angulated, the emargination deep and about one-third distant from
apex; tarsi slightly dilated and squamose beneath.
Genotype, Patroboidea rufa Van Dyke, new species.
This genus, without doubt, belongs in the tribe Pogonini, as
defined by Horn,^ and in the first of his groups, the group
Pogoni. It belongs near the genus Patrobus, and in general has
much the appearance of the flatter members of that genus, those
classed as Platidius by Chaudoir, but it is much smaller than
any of the species included within that, and in fact looks much
like the European Thalassophilus longicornis Sturm. With this
last, it agrees as regards general size, shape and color, small
eyes, and heavy and long antennae, but it differs in lacking the
basal striae of the elytra and in having a very small second
2 “On the genera of Carabid^, with special reference to the fauna of
Boreal America,” by George H. Horn, M. D., Trans. Am. Ent. Soc., Vol. IX
(1881-82), p. 135.
OCTOBER, 1925]
VAN DYKE IIARPALIN^
69
segment and very long third segment to the antennae, in this
last regard resembling Pogonus. From Pogonus, it differs pri-
marily in having the genae well developed and elongated, the
constriction of the head, therefore, not just back of the eyes
but far removed, in having the eyes poorly developed and the
palpi, both maxillary and labial, quite long and slender and sub-
acute at apex, in contrast to the truncated palpi of the other.
Patroboidea rufa Van Dyke, new species
Elongate, quite parallel, much flattened, rufous, head somewhat
darkened. Head elongated, somewhat elliptical, smooth, impunctate;
frontal impressions deep, straight, extending back as far as eyes;
mandibles long, porrect, and but little curved except at apex; eyes
small, projecting but little beyond sides of head; antennge rather
stout, long, extending to beyond middle of elytra. Prothorax slightly
broader than long and about as long as head, including mandibles;
base narrower than apex; apex slightly emarginate; base truncate;
sides moderately arcuate in front, sinuate posteriorly, the hind angles
prominent and right-angled; disc flattened, smooth, with median
longitudinal groove deep and complete, anterior transverse impres-
sion vague, the posterior mainly indicated by a series of well marked
punctures, the basal impressions deep and broad posteriorly. Elytra
slightly longer than head and prothorax combined, about one-third
wider than prothorax, base emarginate, the humeri rounded; sides
almost parallel, but slightly arcuate; disc flattened; striae well de-
fined, complete and with but a few indistinct punctures near base;
scutellar strise fine. Body beneath smooth and shining. Length, 5.5
mm.; breadth, 2.25 mm.
Holotype, male (No. 1820, Mus. Calif. Acad. Sci.), in my
collection, taken by myself from beneath a piece of driftwood
on the sandy beach along the Snohomish River near Monroe,
Washington, July 4-14, 1905. A second specimen, which I
have designated as a paratype, was collected along Spious
Creek, British Columbia, May 30, 1921, by Mr. Ralph Hopping,
and is now in his collection.
Pterostichus brunneus Dej.
This species, about which there has long been much con-
fusion, due chiefly to the fact that neither Leconte ^ nor Horn
knew the species, is well defined by both Dejean ^ and Manner-
3 “The Pterostichi of the United States,” by John L. LeConte, Proc.
Acad. Nat. Sci., Phil., 18G1, p. 312. The specimen, sent to Dr. LeConte
by Chaudoir as the type of Pterostichus brunneus Dej., could not have been
that, but probably a specimen of Pterostichus castaneus Dej. Chaudoir himself
later insisted that brunneus and castaneus were quite distinct.
4 “Species General des Coleopteres,” by M. le Compte Dejean.
70
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
heim,® and is easily recognized once we compare typical speci-
mens with the original description. It was described from
Sitka, but is a common species on the islands off the coast of
southeastern Alaska and on Orcus Island in Puget Sound, as
well as on the Selkirk Mountains of British Columbia and the
Cascades of Washington and Oregon, at least as far south as
Mt. Jefferson. In the northern Cascades it lives as does Pteros-
tichus protractus Lee. in the Sierras, in the woods under old
logs. Structurally, it is about midway between castaneus, which
is generally to be found in the same environment with it, and
protractus, and in fact if found in the Sierras would no doubt
be taken for a small specimen of the latter. It is most decidedly
not the same as castaneus, as stated by LeConte,® nor synony-
mous with scutellaris, as given by Horn.® Pterostichus scutel-
laris Lee. is nothing more than a fully pigmented or melanotic
offshoot of amethystinus. This last, a common species of the
dense forests of the Pacific Northwest, extends into California
and along the coast only so far as northern Sonoma County,
where scutellaris takes its place, and continues on to Monterey
County. There seems to be a distinct cleavage between the two
species, and the fact that in ancient times a gulf existed at
about the place where one replaces the other might account for
this.
Pterostichus nigrocaeruleus Van Dyke, new species
Moderately elongate, somewhat flattened, gracefully proportioned,
head and pronotum shining black, elytra violet to bluish black, an-
tennae, legs and under surface piceous to rufopiceous. Head rather
large, almost two-thirds breadth of prothorax, very smooth and shin-
ing, frontal grooves long and well impressed; the mandibles promi-
nent and noticeably hooked at apex; eyes moderately prominent;
antennae slender, extending several segments behind the base of
the prothorax. Prothorax slightly wider than long, anterior angles
prominent, sides broadly arcuate anteriorly, sinuate posteriorly, hind
angles right-angled; lateral margin broad and reflexed; disc slightly
convex, very smooth and shining, median longitudinal groove deep,
the anterior and posterior transverse impressions distinct; basal im-
pressions single, long and deep, the basal area external to these
flattened, not margined, and slightly reflexed in a wing-like manner.
Elytra nearly two-fifths longer than head and prothorax combined,
5 “Beitrag zur Kafer-Pauna der Aleutischen Inseln, der Insel Sitkha
und Neu-Californiens,” by Graf. C. G. Mannerheim, Bui. Moscow, Vol.
XVI (1843), p. 201.
6 “Random Studies in North American Coleoptera,” by George H.
Horn, Trans. Am. Ent. Soc., Vol. XIX (1892), p. 41.
OCTOBER, 1925 ]
VAN DYKE HARPALIN^
71
four-sevenths as broad as long, at least 1 mm. broader than pro-
thorax, elliptical; humeri well rounded and humeral angles not denti-
form; disc slightly convex; striae deep and simple, the intervals
convex, without foveae and very shining; scutellar stria vague, punc-
tiform, at base of first stria. Legs slender, first three segments of
posterior tarsi deeply grooved on outer side, posterior trochanters
over one-third length of femora. Apex of prosternum not evidently
margined. Beneath moderately smooth and shining. Length, 12
mm.; breadth, 4.5 mm.
Holotype, male (No. 1821), allotype, female (No. 1822,
Mus. Calif, Acad. Sci.), and several paratypes, in my collec-
tion, captured by myself along the sides of a mountain stream
back of Port Angeles, Washington, May 26, 1907. Paratypes
will be deposited in the United States National Museum. In
addition, I have used for comparison over fifty more specimens
in my collection, which I collected at the following localities :
The Forks, Clallam County, Washington; Marys Peak, Ben-
ton County, Marshfield, Olney near Astoria, and Waldport,
Oregon; and from near the Van Duzen River, Humboldt
County, California.
This elegant and beautiful species at first glance might be
confused with Pterostichus amethystinus (Mann.), for it is
of about the size of the normal individuals of the latter, is of
somewhat similar color, and is to be found in the same terri-
tory. Structurally, it is widely separated, much more brilliant,
with the head larger, the prothorax much broader, with the
basal impressions single, long and deep, and the posterior angles
much more prominent ; the elytra elliptical, not parallel sided,
the humeri well rounded and without having the humeral angles
at all dentiform ; the intervals distinctly convex throughout, and
the apex of the prosternum not margined behind. Though a
member of the subgenus Hypherpes, to which belong all those
American species without discal elytral foveae, it stands by
itself. In habits, it is only approached by one other member
of the subgenus, crenicollis, for it lives along the gravelly mar-
gins of the small densely shaded streams which come down
from the hills. Pterostichus amethystinus (Mann.) and its
close relative, Pterostichus scutellaris Lee., are to be found only
in the damp woods in or under old logs.
Pterostichus beyeri Van Dyke, new species
Elongate, subparallel, slightly convex, rufopiceous. Head large,
but .5 mm. narrower than prothorax and about as long, frontal
72
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
grooves distinct though not prominent; eyes small and flattened;
gense bulbous, invading much of the ocular area; mandibles long
and but slightly curved, prognathous; antennas moderately stout,
slightly enlarged apically, about reaching to hind angles of pro-
thorax. Prothorax slightly longer than broad, narrowed posteriorly,
apex feebly emarginate; base transverse, slightly more than one-
third of the greatest width of prothorax; sides feebly arcuate and
convergent backwards almost to hind angles where they become
sinuate; hind angles small, bluntly right-angled, lateral margins fine;
disc feebly convex, the median longitudinal groove vague, the pos-
terior lacking; basal impressions single, broadly though shallowly
impressed, the basal margin only evident close to angles. Elytra
slightly longer than head and prothorax combined, barely 5 mm.
broader than prothorax, elongate elliptical; humeri distinct, humeral
angles not dentiform; disc slightly convex, the striae well impressed,
simple and complete, the intervals slightly convex, a bit flattened
apically and without dorsal foveae; scutellar stria distinct, short and
within base of first interval. Legs rather slender, anterior femora
robust; posterior trochanters almost one-half length of femora and
blunt at apex; first segment of hind tarsi vaguely grooved on outer
side. Apex of prosternum not margined. Beneath moderately smooth
and shining. Length, 12.5 mm.; breadth, 3.75 mm.
Holotype, male (No. 1823, Mus. Calif. Acad. Sci.), in my
collection, captured in the Bitter Root Mountains, Montana,
by Mr. Gustav Beyer and by him very kindly presented to me.
One other specimen, in the Beyer collection, has been seen.
This species is somewhat related to Pterostichus caligans
Horn. It is of the same length, but broader, with the prothorax
and elytra proportionally shorter, as compared with the length,
the eyes slightly larger, the humeral angles not dentiform, the
hind femora not so clavate nor the posterior trochanters so long
and apically narrowed, and the posterior tarsi not so definitely
grooved externally. To Pterostichus ro stratus Newm. it is even
closer structurally, differing mainly by being smaller and less
robust, with single basal prothoracic impression instead of
double, and by having the elytral striae simple. This places it
as an intermediate species between the group to which caligans
belongs and rostratus, and shows that the two genera, Cylindro-
charis ^ and LeptoferoniaJ established by Colonel Casey upon
insufficient characters, should be suppressed. In this connec-
tion, I will also state that Hyperphes Chaud. is a well-defined
subgenus of Pterostichus and should include, not only those
7 “Memoirs on the Coleoptera,” VIII (1918), by Thomas L. Casey, Lan-
caster, Penn., pp. 326 and 336.
OCTOBER, 1925]
VAN DYKE HARPALIN^
73
species listed under Pterostichus in the Leng Catalogue, but
also those included under Hammatomerus Chaud., Holciophorus
Lee., Cylindrocharis Casey, and Leptoferonia Casey. The char-
acters which are used to separate these are divergent ones of
minor degree, whereas those which all of the species possess in
common are fundamental.
Pterostichus falli Van Dyke, new species
Elongate, subparallel, slightly convex, rufous. Head large, .75 mm.
narrower than prothorax and almost as long; frontal grooves dis-
tinct; eyes rather small; gen^ bulbous and about as prominent as
eyes; mandibles long and but slightly curved, prognathous; antennae
moderately stout, distinctly enlarged apically, reaching a little be-
yond basal angles of prothorax. Prothorax as long as broad, nar-
rowed posteriorly, apex feebly emarginate, anterior angles evident
but not prominent; base feebly emarginate, one-third the greatest
breadth of prothorax; sides feebl}^ arcuate and convergent pos-
teriorly, sinuate in front of hind angles, which are sharply right-
angled; lateral margin fine; disc somewhat flattened, the median
longitudinal groove clearly defined and almost reaching anterior and
posterior margins; anterior transverse impression evident, the pos-
terior a broad, shallow depression; basal impressions double, the
inner distinct and of moderate length, the outer vague, small and
close to hind angles; basal margin vague. Elytra equal in length to
head and prothorax combined, slightly broader than prothorax;
humeri distinct, the humeral angles vague; sides subparallel, but
feebly arcuate; disc slightly convex, flattened at middle, striae dis-
tinct and complete though finer apically, the intervals quite flat-
tened and without dorsal foveae; scutellar stria fine, over .5 mm. in
length and at base of first stria. Legs rather slender, femora not
greatly developed, posterior trochanters not one-half length of
femora and blunt at apex; first segment of hind tarsi distinctly
grooved on outer side. Apex of prosternum distinctly margined.
Beneath moderately smooth and shining. Length, 11 mm.; breadth,
3.25 mm.
Holotype, male (No. 1824), and allotype, female (No. 1825,
Mus. Calif. Acad. Sci.), in my collection, captured by myself
in the canons on the east side of the hills east of Hollywood,
Los Angeles County, California, January 9 and 4, 1912. Be-
sides these there are seven other specimens in my collection
which I have designated as paratypes and which were collected
at various times in the type locality. It is named in honor of
my good friend, Mr. Henry C. Fall, who has contributed so
much to the advancement of our knowledge of the Coleoptera
of western North America.
74
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
This species belongs with caligans and beyeri. It is, however,
smaller than either, more distinctly flattened, with the prothorax
more narrowed posteriorly, with two evident basal thoracic im-
pressions, and the apex of the prosternum distinctly margined.
Pterostichus termitiformis Van Dyke, new species
Elongate, flattened, rufous, antennae, legs and under surface lighter.
Head very large, as broad as prothorax and distinctly longer; front
flattened: frontal grooves distinct, broadly though shallowly im-
pressed; eyes small, flattened; genae not bulbous; sides of head pos-
terior to eyes slightly arcuate and convergent posteriorly; mandibles
long and but slightly curved, prognathous; antennae long, extending
several segments beyond hind angles of prothorax and but slightly
dilated apically. Prothorax as long as broad, cuneiform; apex evenly
though shallowly arcuate; base moderately arcuate, its width almost
one-half the greatest width of prothorax; sides feebly arcuate and
convergent backwards, becoming slightly sinuate as they approach
the hind angles, the latter right-angled; lateral margin fine; disc
flattened, the median longitudinal groove distinctly defined at center,
anterior transverse impression vague, the posterior shallowly im-
pressed, basal impressions single, rather long and lunate, basal
margin lacking. Elytra slightly longer than head and prothorax
combined, almost 1 mm. broader than prothorax, elongate elliptical;
humeri broadly rounded, without humeral angles; disc feebly convex,
flattened at middle; striae simple and deeply impressed basally, finer
apically, the intervals flattened though approaching convexity near
base, without dorsal foveae; scutellar stria minute, punctiform, at
base of second stria. Legs rather long and slender, femora not
greatly developed, posterior trochanters not one-half length of
femora and blunt at apex, the first and second segments of hind tarsi
distinctly grooved on outer side. Apex of prosternum not margined.
Beneath moderately smooth and shining. Length, 9.5 mm.; breadth,
3 mm.
Holotype, male (No. 1826), allotype, female (No. 1827,
Mus. Calif. Acad. Sci.), and paratype male in my collection,
captured by myself at Marshfield, Oregon, the first on June 11,
1914, the others on June 10, 1914. They were found beneath
deeply imbedded logs in the woods.
This species is definitely allied to caligans, beyeri, and falli.
It differs materially, though, in general appearance, propor-
tionately larger head, and more delicate form generally. The
non-bulbous genae will also serve to distinguish this species
from its associates. The large head and mandibles and small
prothorax give it an appearance not unlike that of a soldier
termite, a resemblance which has suggested the specific name.
OCTOBER, 1925]
VAN DYKE HARPALIN^
75
The last three species described belong to a peculiar group
or assemblage within the subgenus Hyperphes of Pterostichus,
the members of which are characterized in the main by being
somewhat elongate and cylindrical, with small eyes, long for-
ward projecting mandibles, often large femora, especially well-
developed posterior trochanters, and, in the more divergent
forms, with the antennae enlarged apically. These are all char-
acters which are to be found among those Carabidae which live
a burrowing life, and such is the case with those under dis-
cussion, they being the most subterranean in their habits of the
American species of this large genus. The California species
are rarely found except during the winter or early spring
months, being deep down in the ground during the dry sum-
mers, as is the case with many of our earth-dwelling insects.
The greater portion of this group is at present restricted to
the Pacific Coast, but it is, no doubt, a very ancient one which
was at one time widely distributed over the more northern
parts of North America during later Pliocene times. Because
of their specialized habits they could not readily migrate, and
hence were in great part exterminated during the Great Ice
Age, the species which remain to us today being but the relicts
and those which dwelt in territory that suffered but little from
the effects of that age. To this group belong the following spe-
cies : grandiceps Chaud. and rostratus Newm. from the Appa-
lachian region in general ; beyeri n. sp. and elongatus Schffr.,
from the northern Rocky Mountain region; arisonicus Schffr.
from Arizona; and inanis Horn, angustus Dej., longicollis Lee.,
fuchsi Schffr., caligans Horn, falli n. sp., and termitiformis
n. sp., from the Pacific Coast. Cylindrocharis sulcatula Casey
and piceata Casey are absolute synonyms of Pterostichus ros-
tratus Newm., the characters given for their separation being
insufficient. Of the forms described by Colonel Casey under
Leptoferonia, crucialis is but a weak phase of angustus, pumilis
a slight variant of longicollis, fugax a synonym of fuchsi, lar-
valis and humilis practically the same and at most but a sub-
species of fuchsi, the posterior prothoracic angles being slightly
more evident, and inopina a weak variant of ovicollis. The
species of this group, like all species which are to any degree
subterranean, are unstable and plastic and thus inclined to vary
76
THE PAN-PACIFIC ENTOMOLOGIST [VOL. II, NO. 2
greatly, especially if they range over a great extent of terri-
tory, as do many of those mentioned above.
Pterostichus lanei Van Dyke, new species
Of moderate length and breadth, gracefully proportioned, black
and shining, antennas, mouthparts and legs rufopiceous. Head of
normal proportions, two-thirds breadth of prothorax and one-third
longer than broad, convex; frontal grooves distinct but not promi-
nent; mandibles normal; eyes prominent; antennas slender, extend-
ing several segments beyond apex of posterior prothoracic angles.
Prothorax over one-sixth broader than long, distinctly narrowed pos-
teriorly; apex distinctly emarginate, anterior angles moderately
prominent; base but slightly emarginate at middle; sides broadly
arcuate for anterior three-fourths, straight and convergent to pos-
terior angles; hind angles obtuse, rather sharply defined; lateral
margin fine and well reflexed; disc convex, median longitudinal
groove clearly and rather deeply impressed, almost complete; an-
terior and posterior transverse impressions faintly defined; basal
impressions single, deep and linear; a faint indication of a secondary
impression at the sides of the broadened base of the preceding; the
area close to the posterior angles somewhat flattened, thus making
the angles more evident; base not margined. Elytra about one-sixth
longer than head and prothorax combined, a third longer than broad,
elliptical; humeri well rounded; disc convex; striae complete, deeply
and uniformly impressed, the faintest indication only of punctures;
scutellar stria distinct and of moderate length, the intervals convex,
the third with a distinct fovea just anterior to the middle, and a
second midway between that and the apex. Legs slender, femora
moderately developed, posterior trochanters rather large; first three
hind tarsal segments distinctly grooved on outer side. Beneath rather
smooth and shining. Length, 11 mm.; breadth, 4 mm.
Holotype, male (No. 1828, Mus. Calif. Acad. Sci.), in my
collection, captured by Mr. Merton C. Lane at V/awawai,
Washington, and kindly presented to me. Several paratypes
are in Mr. Lane’s collection.
This graceful black species is of about the size of some of
the smaller specimens of Pterostichus vicinus Mann., and it
somewhat suggests that, but is far less robust, with prothorax
more narrowed posteriorly and humeri well rounded. Its struc-
ture would place it near lachrymosus , particularly the posses-
sion of two dorsal punctures and deep linear basal impressions
to the prothorax, but it is not closely related to that nor to
any of the species associated with that.
(To be continued)
OCTOBER, 1925]
BLAISDELL ELEODES
77
REVISED CHECK-LIST OF THE SPECIES OF
ELEODES INHABITING AMERICA, NORTH
OF MEXICO, INCLUDING LOWER CALI-
FORNIA AND ADJACENT ISLANDS
BY F, E. BLAISDELL, SR.
San Francisco, California
The following revised list of the species of Eleodes inhabit-
ing America north of Mexico also includes those from the
islands in the Pacific, adjacent to the mainland and from the
islands in the Gulf of California. This list is intended to replace
the one given by Leng in the Catalogue of the Coleoptera of
America, north of Mexico. References are given for the spe-
cies and phases not given in the Leng Catalogue.
This list includes 106 species, 67 subspecies, and 12 varieties.
In my Monograph of the Eleodiini (Bulletin 63, United
States National Museum) no attempt was made to designate
subspecies. In the present list this has been done, based on
more recent observations. My conception of subspecies and
varieties or races has very recently been stated in the Proc.
Calif. Acad. Sci., Ser. 4, Vol. XIV, No. 16, September, 1925.
For information regarding forms and for synonyms, see my
Monograph and Leng’s Catalogue.
Eleodes coloradensis Blais, was first recorded as humeralis
Lee. forma tuhcrculo-muricata.
Eleodes concinna Blais, was first considered as humeralis
Lee. forma gramdato-muricata.
Eleodes apprima Blais, was first given as laticollis forma
insularis.
Eleodes immunda Blais, was first given as hispilabris forma
elongata.
In all other cases where a form name has been raised to the
rank of a variety or subspecies the name hereby validated has
been marked as “new name” (n. n.).
The diagnoses of two new subspecies are here included so
the names can be used in this list :
Eleodes liorni fenyesi Blaisdell, new subspecies
Form quite similar to horni, but smaller, more slender, with
the integuments more opaque ; pronotal punctuation denser and
more distinct; clypeal surface scarcely at all rugose, smooth
and flat between the punctures.
78
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
Measurements (types). Length, 11 - 12.5 mm.; width,
4 - 5.5 mm.
Type locality. Bishop, Inyo County, California. Col-
lected by Dr. A. Fenyes. Holotype, female, and allotype, male,
in my collection. Horni occurs in Plumas County, California.
Eleodes manni sierra Blaisdell, new subspecies
Form somewhat larger and a little more robust than manni;
humeri more strongly developed and moderately narrowly
rounded; luster duller; pronotal punctuation much finer and
equally as dense; sides of the prothorax more strongly sinuate
before the basal angles, where they are parallel and straight
for a short distance. In the female there are evident striae of
punctures on the elytra, in the male these are not as well
marked.
Measurements (types). Length, 16.5 - 15 mm.; width,
5.5 - 5.6 mm.
Type locality. Kings Canon, Fresno County, California.
Collected on August 4, 1917.
BIBLIOGRAPHY
Blaisdell, 1921. Canadian Entomologist, Vol. LIII (No. 6), pp. 129-
132, June, 1921.
Blaisdell, 1921. Stanford University Publications, University Series.
Biological Sciences, Vol. 1 (No. 3), pp. 217-226, June, 1921.
Blaisdell, 1923. Proceedings California Academy of Sciences, Ser. 4,
Vol. XII (No. 12), pp. 201-288, July, 1923.
Blaisdell, 1925. Proceedings California Academy of Sciences, Ser. 4,
Vol. XIV (No. 14), pp. 321-343, September, 1925.
Blaisdell, 1925. Proceedings California Academy of Sciences, Ser. 4,
Vol. XIV (No. 16), pp. 369-390, September, 1925.
Genus eleodes Esch.
Subgenus Melaneleodes Blais,
debilis Lee.
carbonaria Say
subsp. soror Lee.
vicina Lee.
nitida Casey
ampla Blais,
obsoleta Say
subsp. annectans Blais, n. n.
subsp. porcata Casey,
mazatzalensis Blais., 1925, 379.
knausi Blais.
omissa Lee.
subsp. pygmaea Blais,
subsp. borealis Blais, n. n.
subsp. peninsularis Blais,
parowana Blais., 1925, 374.
subsp. mimica Blais., 1925, 375.
quadricollis Esch.
subsp. lassenica Blais., 1925,
373.
subsp. anthracina Blais,
subsp. lustrans Blais,
cuneaticollis Casey
OCTOBER, 1925]
BLAISDELL ELEODES
79
humeralis Lee.
reducta Blais., 1925, 377.
concinna Blais., 1925, 381.
coloradensis Blais., 1925, 380.
fuscipilosa Blais., 1925, 376.
rileyi Casey
tricostata Say
var. asperata Lee.
pedinoides Lee.
neomexieana Blais,
speeulieollis Blais., 1925, 382.
wenzeli Blais., 1925, 381.
Subgenus Litheleodes Blais,
areuata Casey
extrieata Say
subsp. eonvexieollis Blais., n. n.
subsp. eognata Hald.
subsp. arizonensis Blais,
subsp. utahensis Blais., 1921,
131.
granulata Lee.
subsp. obtusa Lee.
subsp. eorvina Blais., 1921, 224.
papillosa Blais,
leteheri Blais,
vandykei Blais,
var. parvula Blais,
var. modifieata Blais., 1921, 131.
Subgenus Tricheleodes Blais,
hirsuta Lee.
pilosa Horn
barbata Wiekh.
Subgenus Pseudeleodes Blais,
granosa Lee.
Subgenus Promus Lee.
insularis Lined
subsp. terrieola Blais,
subnitens Lee.
goryi Sol.
subsp. seriata Lee.
striolata Lee.
fusiformis Lee.
opaea Say
Subgenus Heteropromus Blais,
veterator Horn
Subgenus Eleodes Esch.
obseura Say
subsp. dispersa Lee.
subsp. suleipennis Mann,
subsp. glabriuseula Blais., 1925,
383.
aeuta Say
suturalis Say
var. texana Lee.
grandieollis Mann.
subsp. valida Boh.
sanmartinensis Blais., 1921, 220.
var. moesta Blais., 1921, 221.
sponsa Lee.
hispilabris Say
subsp. nupta Lee.
subsp. attenuata Blais,
subsp. eonnexa Lee.
/. cowvexa, Leng Cat. 228.
subsp. seulptilis Blais,
subsp. imitabilis Blais,
subsp. immunda Blais., n. n.
/. elongata, Leng Cat. 228.
subsp. eomposita Casey
subpinguis Blais,
graeilis Lee.
subsp. distans Blais,
eaudifera Lee.
longipilosa Horn
dentipes Eseh.
var. perpunetata Blais,
subsp. elongata Blais,
subsp. tularensis Blais., 1925,
386.
subsp. montana Blais., 1925,
385.
subsp. marinas Blais., 1921, 218.
subsp. eonfinis Blais,
subsp. elegans Casey
subsp. prominens Casey
subsp. pertenuis Blais,
subeylindriea Casey
armata Lee.
subsp. impotens Blais,
inepta Blais., 1925, 334.
militaris Horn
subsp. femorata Lee.
80
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
loretensis Blais., 1923, 262.
vanduzeei Blais., 1923, 264.
morbosa Blais., 1925, 335.
acuticauda Lee.
var. punctata Blais.
lati'Collis Lee.
subsp. minor Blais, n. n.
subsp. apprima Blais., 1921, 219.
eschscholtzi Sol.
subsp. lucae Lee.
niutilata Blais., 1921, 222.
inflata Blais., 1923, 261.
adumbrata Blais., 1925, 332.
discincta Blais., 1925, 333.
tenuipes Casey
wickhami Horn
ventricosa Lee.
subsp. falli Blais.
Subgenus Blapylis Horn.
tibialis Blais,
snowi Blais,
lecontei Horn
tenebrosa Horn
var, liana Blais,
inculta Lee.
subsp. affinis Blais,
consobrina Lee.
scabripennis Lee.
blanchardi Blais,
neotomse Blais,
fuchsi Blais,
horni Blais.
subsp. fenyesi, new subsp.
subsp. monticola Blais,
manni Blais.
subsp. sierra, new subsp.
subsp. dilaticollis Blais., 1925,
388.
subsp. variolosa Blais,
parvicollis Escli.
subsp. farralonica Blais, n. n.
subsp. planata Esch.
subsp. squalida Blais,
producta Mann.
subsp. alticola Blais., 1925, 387.
subsp. trita Blais,
subsp. constricta Lee.
behri Grinnell
hoppingi Blais,
clavicornis Esch.
scabrosa Esch.
rotundipennis Lee.
var. versatilis Blais., 1921, 217.
cordata Esch.
subsp. horrida Blais,
subsp. patruelis Blais,
var. adulterina Blais,
var. hybrida Blais,
var. sublaevis Blais, n. n. ,
pimelioides Mann,
nunenmacheri Blais.
subsp. verrucula Blais,
brunnipes Casey
subsp. brevisetosa Blais,
propinqua Blais,
caseyi Blais.
Subgenus Metablapylis Blais.
nigrina Lee.
subsp. perlonga Blais,
subsp. difformis Blais., 1925,
389.
dissimilis Blais.
subsp. nevadensis Blais,
schwarzi Blais.
Subgenus Steneleodes Blais.
gigantea Mann.
subsp. meridionalis Blais,
subsp. gentilis Lee.
subsp. estriata Casey
longicollis Lee.
innocens Lee.
Subgenus Discogenia Lee.
marginata Esch.
scabricula Lee.
subsp. acutangula Blais., 1921,
225.
Genus neobaphion Blais.,
1925, 390.
planipennis Lee.
OCTOBER, 1925] BARNES AND BENJAMIN HEPIALUS
81
THE HYPERBOREUS GROUP OE HEPIALUS
(LEPID., HEPIALIDiE)
BY WILLIAM BARNES AND F. H. BENJAMIN
Decatur, Illinois
Hepialus roseicaput N. & D.
1893, Neumoegen & Dyar, Can. Ent, XXV, 125, Hepialus.
1894, Neumoegen & Dyar, Jour. N. Y, Ent. Soc., II, 169, Hepialus.
1895, Ottolengui, Ent. News, VI, 290, Hepialus.
1896, Dyar, Can. Ent., XXVIII, 176, hyperboreus confusus, Hepialus.
1903, Dyar, Bull. U. S. N. M., LII, 580, No. 6608b, hyperboreus con-
fusus, Hepialus.
1911, McDunnough, Can. Ent., XLIII, 292, (? matthe^'i{\) $), Hepi-
alus.
1917, Barnes & McDunnough, Check List, p. 197, No. 8488, mathe^i,
Hepialus.
I hyperboreus Auct. nec. Moesch.
1911, McDunnough, Can. Ent., XLIII, 290, Hepialus.
j pulcher Auct. nec Grt.
1911, McDunnough, Can. Ent., XLIII, 291, Hepialus.
Described from a single specimen from the Cascade Range
near L3Aton, B. C. The type is a $ ; not a male, as stated in the
original description (1893) and by Ottolengui (1895).
Dyar (1896 and 1903) sinks the name to confusus; McDun-
nough (1911) thought it $ mathewi; and Barnes & Mc-
Dunnough (1917) sink it to mathewi; but apparently lack of
sufficient material in the past was prohibitive to properly sort
the various forms allied to hyperboreus. This lack of material,
coupled with misdeterminations on the part of all authors, has
led to a very general confusion of the names in the group.
Some authors appear to desire to call all American species
synonyms of the European H. ganna. Judging from several
figures, this species is nearest to mcglashani, but we think
entirely distinct.
Both authors have independently examined all types involved
except hyperboreus, which is unknown to us except from the
literature and Moeschler’s figure of type (1862, Wien. Ent.
Monat., VI, pi. I, f. 1), and pulcher, of which we possess
specimens agreeing well with Grote’s figure of type (1864,
Prbc. Ent. Soc. Phila., HI, 522, pi. V, f. 3).^
A resume of the species in key form may assist to sort them.
We are inclined to think that each name so far proposed rep-
resents a distinct species. H. roseicaput is the Mt. Hood spe-
1 Since writing the above we have received a photograph of type pulcher
through the courtesy of Dr. H. Skinner.
82
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
cies, alternately called hyperhoreus and pulcher by McDnn-
nough (1911). Normally, it appears to be sexually dimorphic,
the type agreeing with three out of our five females. Another
female shows silver markings as in the male, and we call this
form mutatus nov. We have a single female, with immaculate
primaries, and we are calling this demutatus nov. Our series
of 12 6 , 5 2 were taken by McDunnough on Mt. Hood, Ore-
gon, 8-15 August. We knov/ confusus only from the type ( $ )
in the Bolter collection from Sitka, Alaska, and a female from
“Metlakatla, B. C., August 21, 1903, Rev. J. F. Keen,” in
the Barnes collection. H. pulcher is known to us from 1 6 ,
Glenwood Springs, Colorado, 16-23 August, and 2 6 “Utah.”
H, mcglashani is represented in the Barnes collection by 2 $ ,
1 2 from unknown localities, 1 6 labeled “Nev.” (which may
mean Sierra Nevada or Truckee, California), and a 3 2 from
Placer County, California. H. mathezvi is represented by a
series of sixteen specimens, both sexes, from Duncans and Vic-
toria, B. C., from Messrs. Blackmore and Hanham. All speci-
mens except one are dated, and the dates range from the 15th
to the 29th of September.
Flepialus novigannus Barnes & Benjamin, n. sp.
A series of 21 6 , 17 2 are in the Barnes collection from
Hymers, Ontario, dates ranging from August 8 to Septem-
ber 15. In general all specimens are similar in size and in
shade of ground color to mcglashani, but the silver markings
are much cleaner cut and entire ; aside from the normal V con-
nected to the wing base, these are restricted to one or two spots
below vein 1, some slight markings in the costal region, and a
tendency toward dots along the outer margin, which, in even
the best marked specimens, are obsolescent. No tendency
toward sexual dimorphism. A dusk-flying species.
Hepialus novigannus mackiei Barnes & Benjamin,
race nov.
Another series of 11 6, 12 2 from Edmonton, Alta. (D.
Mackie) and 16,22 from Calgary, Alta. (Dod?) appear to
represent an insect at least racially distinct. Size and silver
markings as in the preceding. While all of the other species
before us do not vary greatly, except mathewi, the present form,
while possessing uniform maculation in both sexes, varies sur-
prisingly in color, ranging from deep brown to salmon-pink. In
the event the present insect should ultimately prove consubspe-
cific with the Ontario species, we are restricting our types to
OCTOBER, 1925] BARNES AND BENJAMIN HEPIALUS
83
5^,5? which show the salmon-colored primaries. The Ed-
monton specimens range in dates from the 24th of August to
the 11th of September. Only one Calgary specimen is dated,
‘T6-23 Aug.”
Superficial key to the species and forms of the hyperboreus
group of Hepialus.
I. Some silver or whitish markings on fore wings.
A. Conspicuous silver markings other than a broadly open
V connected to wing base;
a. Fore wing with silver markings irregularly split
into spots; hind wing marked by a fuscous band
from apex to near wing base. (Labrador)
hyperboreus
b. Not so; both characters not present in one species.
a. i Silver banding on fore wing, aside from a
basal silver connecting dash, as a broadly
open V not very irregular, although some-
times tending to split into spots, the other
silver markings disconnected; hind wing
with uniformly fuscous disc and discoiicolor-
ous pinkish fringes; usually sexually dimor-
phic; flying 1:30 to 3:30 p. m. in bright
sunshine. (B. C.; Ore.)
6 roseicaput, 2 roseicaput form mutatus
b. i Silver banding on fore wing usually forming
a V, but the V decidedly irregular, often
with some silver tending to form a spur or
spurs on the V, either subapically or near
the base, the V seldom clean-cut except in
some 5 2 ; sexes similar; flying at dusk.
(Calif.) 6 2 mcglashani
c. ^ Silver banding on (left) fore wing tending to
form an oblique E, usually with an addi-
tional apical spur. When the E is broken
into spots these spots are connected by black,
a. 2 Silver markings usually broken into
numerous small spots connected by
black; hind wing with uniformly fus-
cous disc. (Alaska; B. C.).... 2 confusus
b^. Silver markings mainly contiguous;
hind wing with salmon-colored disc,
on which there is an irregular blackish
suffusion. (Colo.; Utah) t pulcher
B. Silver markings restricted to a broadly open V connected
to wing base; except for one or two marks on or near
84
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
costa, the markings below vein 1, and obsolescent spots
at base of fringes;
a. Fore wing usually with a black spot at the end of
the cell, the apical portion of the silver V tending
to become obsolescent or obsolete and usually re-
placed by black; hind wing basally more hairy than
in other species, usually of some shade of luteous,
salmon, or rufous, and usually marked by black.
(Vane. Is.) 6 $ mathe'vji
b. Forewing without black spot at the end of the cell;
the apical portion of the silver V not tending to
become obsolescent or to be replaced by black;
hind wing less hairy than in the preceding, and
usually smoother and less mottled-looking;
a. i Some shade of brown, often more or less
slightly tinged with rufous; flying at dusk,
(Ontario) 6 $ novigannus
b. ^ Color variable; types restricted to salmon-
pink form. (Alberta.)
novigannus race (or form) mackiei
II. No silver or whitish markings.
A. Fore wing marked with more or less of a black V. (B. C.;
Ore.)- $ roseicaput
B. Fore wing unmarked. (Ore.).... 2 roseicaput form demutatus
The Blaisdell Collection of Coleoptera
At the November meeting of the California Academy of
Sciences, Dr. Barton Warren Evermann, Director of the Acad-
emy, announced that Dr. Frank E. Blaisdell, Sr., had presented
to the Academy his entire collection of Coleoptera on the same
terms under which the Van Dyke collection of Coleoptera was
accepted by the Academy. Under these terms Dr. Blaisdell
retains control of the collection during his lifetime, with the
privilege of retaining any portions for study at home as he
may require. When the Van Dyke and Blaisdell collections,
which are comparable in size, are finally incorporated into the
Academy collection, it will constitute a remarkably complete
representation of the coleopterous fauna of the western United
States, as well as an excellent general collection of beetles.
These two collections supplement one another, so there is less
duplication than one might expect. The Blaisdell collection is
very complete in the Tenebrionidse, Melyridse and some other
families, while the Van Dyke collection excels in the Carabidae,
Elateridse, and certain groups of forests insects, such as the
Buprestidae and Cerambycidse. — E. P. Van Duzee.
OCTOBER, 1925] CHAMBERLIN LEPIDOSAPHES
85
A NEW SPECIES OF LEPIDOSAPHES FROM CHINA
(HEMIPTERA, COCCID^^)
BY JOSEPH CONRAD CHAMBERLIN
Citrus Experiment Station, Riverside, California
The species of Lepidosaphes, described below, is very similar
in general appearance to L. heckii, the purple scale, with which
it was originally confused. Morphologically, it is clearly dis-
tinct therefrom, as shown below.
I am indebted to Mr. E. E. Green and Professor G. F. Ferris
for confirming my opinion relative to its status as a new species.
Mr. Green suggests that it is possibly the same as L. euryce
Kuwana, as described in the Proceedings California Academy
of Sciences, Vol. HI, p. 80. That description is inadequate,
however, to permit an exact determination, and consequently
this form is best regarded, for the present at least, as distinct.
Lepidosaphes chinensis Chamberlin, sp. nov.
Type host and locality. On “magnolia” from Canton, China,
Collected by F. Silvestri, 1925.
Habit : Occurring on leaves and stem of host. Scale of
female typical for the genus ; dark reddish-brown, very similar
in general appearance to L. beckii, the purple scale. Scale of
male of the type common to the genus.
Morphological characteristics: Adult female (Figure 1). Length
about 1.0 mm. Derm membranous throughout, except for pygidium
and spur-like lateral projections of the abdominal segments; thorax
distinctly separated from abdomen by a constriction; thorax divided
midway by a distinct transverse crease; margin of abdominal seg-
ments distinctly projecting laterally, bearing gland-spines and
numerous small ducts. There is a distinct transverse row of five
or six large ducts immediately posterior to the posterior spiracles.
On each abdominal spur and on the base of the pygidium there is
a round, heavily chitinized boss or raised disc, five pairs in all.
Pygidium (Figure 2) with two pairs of well-developed lobes; median
pair well spaced, mesally notched, as a whole well rounded; second
pair less prominent than mesal pair, distinctly bilobed. Between the
mesal lobes is a simple pair of gland spines; between the mesal and
second lobe is a pair of gland spines and a large submarginal pore;
beyond the second lobe another pair of gland spines and two large
submarginal pores; then two more spines and another pair of sub-
marginal pores, and lastly two more gland spines and a single
submarginal pore. All submarginal pores with conspicuous chitinous
rims. Dorsal ducts smaller than marginal ducts, fairly numerous and
1 Paper No. 134, University of California, Graduate School of Tropical
Agriculture and Citrus Experiment Station, Riverside, California.
86
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
arranged in two conspicuous bands, as shown in Figure 2. No ven-
tral ducts. Circumgenital pores in five small groups of four to six
pores each. Anal opening anterior to cephalic group of circumgenital
pores.
From L. heckii, its nearest relative, chinensis may be distin-
guished by the fewer circumgenital pores and by the greater
number of marginal chitinized bosses. In heckii, these bosses
are found on the two anterior abdominal segments and the pre-
pygidial segment only, three pairs in all.
The holotype is deposited in the collection of the Department
of Entomology of the University of California Citrus Experi-
ment Station. Paratypes are deposited in the Stanford Uni-
versity collection and in Mr. E. E. Green’s collection. Type
material in the United States National Museum.
aspect of adult female
OCTOBER, 1925] CHAMBERLIN LEPIDOSAPHES
87
Figure 2. Lefidosaphes chinensis sp. nov. Pygidium of adult female.
THE HATCHING OF THE EGGS OF PERIPSOCUS
CALIFORNICUS BANKS
BY SIBYL WACHTER
Peripsociis calif ornicus Banks is a species of the family Pso-
cidas (Corrodentia) . It is commonly found on the leaves and
bark of several species of garden shrubs and trees.
The eggs of this species are laid in masses of nine or ten
on the underside of the leaves, beneath a silken web which is,
as a rule, covered with a loosely woven canopy. Occasionally,
the canopy is absent. Observed individuals have been seen to
oviposit six masses of eggs within ten days’ time, the laying
of each mass covering a period of from twenty-four to forty-
eight hours. The eggs hatch in from eight to ten days from
the time of laying.
The first external indication of life in the eggs, as viewed
under the binocular microscope, is the appearance of a black
structure extending from the apex of the narrower anterior
end of the transparent shell backward about a fifth of the length
of the egg. It becomes very distinct the day before hatching.
The immediate sign of hatching is the movement of this mech-
anism or egg-burster, followed practically simultaneously by
the appearance of a bubble of air at its basal extremity.
Under the microscope the egg-burster appears as an elon-
gated crescent-shaped saw, semi-circular at its basal extremity,
88
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
with the serrate edge extending from this point to its sharply
pointed anterior end. The serrate edge indents one-third of its
width. This mechanism is a part of the amnion, to which it is
attached basally by strands of membrane slightly heavier than
the rest of the amnion.
In this latter structural detail of the egg-burster we are able
to verify a previous statement of Peyerinoff, who described a
frontal chitinous organ consisting of a dentate ridge upon the
amnion of a young psocid. We came to this conclusion from
observing the amnion slip from the embryo as the body worked
itself free, for as it contracted into a wrinkled mass at the
opening of the chorion, the amnion carried with it the tiny saw,
with the aid of which the embryo had attained its freedom.
Furthermore, after dissecting the embryo from the chorion
when it was about to hatch, we were able to slip the embryo
out of the amnion, the egg-burster remaining as a part of the
amnion.
In hatching, consecutive movements of the egg-burster in-
crease gradually the extent of the area through which it works
until it finally cuts through the chorion. Two or three move-
ments are followed by a period of inactivity. In some instances
these periods last five to ten minutes. At corresponding inter-
vals bubbles of air may be seen passing into the oesophagus,
where they increase in size and number as the process advances,
and, passing down through the entire digestive tract, inflate the
body. After a few movements of the egg-burster the basal end
protrudes through the chorion, and at the break the edges
wrinkle back as the opening increases. In the posterior region
of the chorion deflation occurs, as though by the creation of a
vacuum at that point, as the frontal area of the embryo breaks
through and begins to push upward. At this time, which aver-
ages five minutes from the first signs of activity, bubbles of air
enter the alimentary canal in rapid succession, some six or eight
following each swing of the egg-burster. This now moves
through a considerable area. The direction of movement is
back and forth in the direction of the longitudinal axis of
the egg.
Each proximal movement of the anterior extremity of the
burster against the anterior extremity of the head of the em-
bryo causes a very decided indentation at that point. This is
followed by a distal movement of the burster upon which the
OCTOBER, 1925] WACHTER PERIPSOCUS EGGS
89
membrane resumes its previous turgid condition. There is
apparently a direct relation between the movements of the
burster and the accumulation of air in the digestive tract by
means of which the embryo issues from its membranal envelope.
This, then, would indicate an additional use of the burster. The
amnion tears triangularly from the basal extremity of the
burster toward its anterior extremity. The pressure exerted
upon the inflated body by the proximal movement of the burster
would thus seem to play some part in freeing the embryo. Pre-
vious references to the function or air and the use of the burster
in the hatching of the Psocidae have been reviewed by Knab.^
In emerging from the chorion the embryo takes barely a
minute to attain an upright position. The body rotates continu-
ally in line with the position of the egg and soon bends for-
ward, so that the burster is hidden from view. It is covered
with the amnion and presents a smooth transparent appearance
with distinct black eye-spots, which appear soon after emer-
gence. The interior is still filled with air bubbles. As the
anterior end bends forward and down, almost touching the
chorion, the amnion snaps off, splitting just below the eyes,
and is thrown off by pressure from within, a part of it remain-
ing as a wrinkled mass at the split in the chorion and a part
extending backward from the burster in the form of a minute
triangle. The body of the insect now straightens out, the con-
striction between the head and thorax becomes evident, the legs
are raised and begin to move with considerable rapidity toward
and away from the body, and the palps and antennae are gently
waved. The slipping off of the amnion from the time the body
bends over to the time of the freedom of the appendages, and
backward movement of the body into an upright position aver-
ages three minutes. The entire hatching period of the egg varies
from twenty to forty minutes. Its length seems to be deter-
mined largely by the duration of the intervals of inactivity of
the burster and the time needed to accumulate air bubbles in
the first stages of hatching.
After moving the appendages up and down for a few mo-
ments, the insect moves forward to rest lightly on its delicate
legs, and finally walks away from the egg mass. It remains
quite inaxtive for a time except for slight movements of the
abdomen, in which the air bubbles are still visible. These grad-
ually disappear. Five minutes later the psocid walks away
and for several hours remains relatively inactive.
1 Knab, Frederick, The Role of Air in the Ecdysis of Insects. Proc.
Ent. Soc., Wash., Vol. XI, 1909, 68-73.
90
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
THOMAS LINCOLN CASEY
BY FRANK E. BLAISDELL
Colonel Thomas Lincoln Casey, the well-known coleopterist,
retired engineer of the United States Army, died at his resi-
dence in the Stoneleigh Apartments, Connecticut Avenue and
L Streets, Washington, D. C., in the early morning of Feb-
ruary 3, 1925, at the age of 67 years. He had been ailing and
unable to attend to his entomological work for considerable
time. The funeral services were conducted at his residence,
and he was buried on February 6, in the Arlington Cemetery
with full military honors.
Born at West Point, New York, Colonel Casey was the son
of the late General Thomas Lincoln Casey and Mrs. Emma
Weir Casey, and was the brother of Edward Pearce Casey, a
noted architect.
Colonel Casey was a member of the Class of 1877 at Yale
and was graduated from the United States Military Academy
in 1879, and entered the Engineer Corps of the Army. He was
promoted to the grade of first lieutenant in 1881 and of captain
in July, 1888. He was made a major in 1898 and a lieutenant-
colonel in 1906. He became a colonel in September, 1909. He
married Miss Laura Welsh of Philadelphia in 1898.
During the Spanish-American War Colonel Casey was in
charge of the defense of Hampton Roads, Virginia. He had
also been at many stations, in charge of river and harbor
improvements.
He had the distinction of having been an assistant astronomer
under Professor Simon Newcomb on an expedition to Cape
of Good Hope in 1882 to make a study of the planet Venus,
He was a member of the Gree Company Commission to Texas
in 1886 and of the Mississippi River Commission in 1902-1906.
He also served on numerous other special boards, doing vari-
ous kinds of engineering and scientific research. In 1904 he
was in charge of the United States Engineering Exhibit at the
St. Louis Exposition, and between the years 1906 and 1910 he
was also an engineer and secretary of the lighthouse board. He
was retired from active service in the Army in 1912.
Colonel Casey was a member of the New York Academy
of Sciences, the Philadelphia Academy of Natural Sciences,
American Society of Naturalists, the Washington Academy of
OCTOBER, 1925]
BLAISDELL T. L. CASEY
91
Sciences, Astronomical Society of the Pacific, the University
and Century Clubs of New York and the Metropolitan, Army,
Navy, and Chevy Chase Clubs of Washington, D. C.
Personally, I first began to correspond with Colonel Casey
about 1890. My letter file yields a letter dated June 30, 1891.
For several years I submitted material to him for identifica-
tion, both while I was resident at San Diego and for the eight
and a half years that I lived at Mokelumne Hill, Calaveras
County. The results of my collecting can be traced through
his writings. I had the pleasure of visiting him in Washing-
ton in 1909, v\^hile I was in that city indexing my Monograph
on the Eleodiini. I called at his apartment in Stoneleigh Court
and from there we went to his mother’s residence at 1419
K Street, N. W. Washington. It was there that I examined a
number of his types.
Colonel Casey was a most hospitable host. In conversation
he told me that he worked but three hours a day. He explained
how he arranged and studied the specimens and species before
he began to write. He endeavored to be very careful in draw-
ing up his descriptions, and he recorded the cha.racters of the
beetles as he saw and interpreted them, not always as we would
have analyzed them — but, then, how many of us see things as
others see them? Colonel Casey has opened up many lines of
discussion regarding the question of species, subspecies, and
forms. Time will prove whether or not his views are correct.
At present it appears that time will work in his favor, if the
recent studies in S erica and Donacia are any criteria. Critics
had better remain conservative until more work has been done
on the genital characters. Colonel Casey’s decisions have been
freely criticized. He has made many more mistakes than some
other writers because he has done many times more work than
they have. The works of Dr. George Horn yield as many
errors, if his writings on the Tenebrionidse can be taken as a
criterion. Dr. Horn was too conservative and thereby retarded
the science of taxonomy. Colonel Casey advanced taxonomy
too rapidly. In spite of all this we are advancing sanely, and
the best of us can only hope to add our mite to the grand total
of the world’s knowledge.
92
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
ECONOMIC NOTES
EDITED BY E. 0. ESSIG
University of California, Berkeley, California
Leptoglossus clypealis Heid.
The Yucca Elower Bug, Leptoglossus clypealis Heidemann,
was taken in great numbers by the writer, in company with
Dr. E. C. Van Dyke and F. H. Wyiiiore, in the large flowers
of Yucca, in the mountains of San Diego County on April 9,
1925. During the heat of midday the bugs were exceedingly
active, and it was almost impossible to capture them, either with
a net or by hand, because of the formidable sharp-pointed yucca
leaves. Our company, however, secured quite a number of
adults, after having received many painful jabs. — E. O. Essig.
Ash-leaf Bug, Neoborus illitus Van Duzee
This very pretty little yellow and black bug was reported
in great numbers on Oregon Ash at Red Bluff, California,
April 23, 1925. — E. O. Essig.
Miscellaneous Notes
Two species of Sparganothis (Tortricidse) were reared from
walnut husks at Saticoy, California.
The common stored products mite, Tyroglyphus farince
DeGeer, has caused considerable damage to stored walnuts
at San Bruno, California.
The caterpillar of Zale nor da Smith injured blackberry vines
at Saticoy, California, by feeding on the growing tips.
Phthorimcea sp. (probably operculella Zeller) was reared
from larvae found boring in tomato stalks in San Diego County,
Three specimens of an extremely pale form of Carpocapsa
pomonella var. simpsoni Busck were reared at Saticoy, Cali-
fornia. — S. E. Flanders.
Great Abundance of Crickets in the
San Francisco Bay Region
On several nights this last August, during a period of unusual
humidity just preceding a storm, great numbers of the common
field cricket were attracted to the lights in the various cities
about San Francisco Bay. During the day the horde sought
shelter wherever they could find it, and as a result congregated
in the stores in such numbers that many of the merchants
became much worried lest their goods be injured. This is the
most numerous appearance of these insects in this territory of
which I know. — Edwin C. Van Dyke.
OCTOBER, 1925]
LARSON BEAN MITES
93
FURTHER NOTES ON HUMAN SUFFERING
CAUSED BY MITES, PEDICULOIDES
VENTRICOSUS NEWP.
BY A. 0. LARSON
Bureau of Entomology, United States Department
of Agriculture
In an earlier article (Jour. Econ. Ent., Vol. 13, No. 3, p. 322)
the writer described the effects of injuries which he received
from the bites of the mites Pediculoides ventricosus Newp.
while handling weevily beans and cowpeas. As he was hand-
ling these mite-infested seeds daily, it was impossible to deter-
mine how soon the characteristic dermatitis appeared after the
bite had occurred, but he was of the opinion that the burning
sensation began within a very short time. On a number of occa-
sions the writer and co-workers have had minor attacks from
these mites, but, owing to the nature of the work, they have
usually been uncertain as to the time elapsing before discom-
fort became noticeable or before the appearance of reddened
spots. They thought that discomfort came within a few min-
utes, and that the spots always appeared in less than twenty-
four hours. They noticed a difference in susceptibility to the
injury. On one occasion, when three persons were similarly
exposed to the attack of mites, one who had red hair and a
ruddy complexion was injured much worse than either of the
others, while the writer showed very much less injury than
the others.
In describing the effects of applications of these mites to
various parts of the body, Dr. H. E. Ewing' (Can. Ent., Vol.
54, No. 5, p. 108) says: ^‘The appearance of reddened spots,
each with a small papule in the center, was noted the day fol-
lowing application. These spots usually did not develop into
wheals, but did itch considerably.”
Since reading the above statements, the writer and co-
workers, to their sorrow, have had another opportunity to get
first-hand information. On Monday, September 15, a laborer,
James Petersen, began sorting beans in the laboratory on a
specially prepared table. He used as a stopper at the lower
end of the table a small sample bag containing about one and
one-half pounds of beans, to prevent the beans on the table
94
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 2
from rolling off while they were being examined. Some time
during the day he substituted a larger bag filled with weevily
cowpeas. Tuesday morning he said he had an “awful dose of
hives” all over his arms and body. The peculiar thing about
these hives was that none were located on his legs. After an
examination I told him that his hives looked like mite bites.
There were no mites in the beans he was examining, and I
knew of no mites in the laboratory, but if mites had been crawl-
ing onto him from beans stored near by, they would have at-
tacked his legs as well as his body. He was left alone on
Tuesday, as he had been on Monday.
Wednesday morning, when I received word that he was
unable to return to work, I began the work where he had left
off, but as soon as I picked up the bag he had been using at
the lower end of the table, I knew why he was not at work.
This bag was filled with weevily cowpeas, which were badly
infested with mites. He had used the bag, not only to keep
the beans from rolling off the lower end of the table, but he
had used it to wipe the dust off the table after removing each
small lot of beans. The mites from the bag had thus been
spread all over the table and had crawled onto his arms and
body. His legs were not in contact with the table, and were
uninjured.
After removing the bag of mite-infested cowpeas and mak-
ing what I thought was a thorough cleaning of the table and
surroundings, I went to see Mr. Petersen. His body and arms
were a solid red blotch “burning like fire.” He had first noticed
this burning on Monday afternoon. It caused much discom-
fort during that night, but the addition of more mites on Tues-
day made it almost unbearable. The external burning was sup-
plemented by turns of fever followed by a cold sweat, making
sleep impossible. As the writer, on previous occasions, had
obtained relief by bathing in warm soapy water and then apply-
ing talcum powder, this treatment was recommended and im-
mediately gave temporary relief. During the night he obtained
little sleep between the several necessary applications of talcum
powder. He was back at work Thursday morning.
On Wednesday morning I went over the table with a kero-
sene-soaked cloth to remove all mites after I had burned the
mite-infested bag of cowpeas. During the afternoon the writer
OCTOBER, 1925]
LARSON BEAN MITES
95
and a co-laborer, Mr. C. K. Fisher, sorted beans on the table.
In a short time Mr. Fisher was scratching where mites had
bitten his arms and body, and before evening wheals had ap-
peared. The writer, having an equal opportunity during the
afternoon to become infested with mites which had remained
in the cracks of the table, and a better opportunity to become
infested in the forenoon while removing the infested bag of
cowpeas and cleaning the table, first became aware of their
ill-effects when he awoke at 5 a. m. on Thursday morning,
scratching a few burning wheals on his body and arms. Mr.
Fisher had not been in the laboratory since 7 :30' a. m. the pre-
vious Saturday, consequently the bites caused wheals to appear
on his body in less than three hours.
The table was again gone over with kerosene on Thursday
morning and work was resumed, with no new attack of mites.
At the end of a week the marks were plainly visible on the
arms of Mr. Petersen and Mr. Fisher.
In this instance, as in the one cited above, not only was the
writer injured much less than the other men, but a longer time
elapsed before the injuries became visible, and their disappear-
ance was more rapid. This indicates that different individuals
may not react similarly either in the severity of the injury or
the length of time required for the wheals and pustules to
appear. It may suggest a tendency toward an acquired immu-
nity to the effects normally produced by the first attack of the
mites.
Where to Find Zacotus matthewsi Lee.
This brilliant purplish copper-colored carabid ranges from
British Columbia to northern California and in the north as
far east as the Bitter Root Mountains of Montana. It prefers
damp situations, and in the humid lowlands is to be found in
the woods under cover not far from streams or springs. It is
here of a very brilliant copper color and rather hard to find
because of the immense amount of cover. On the high Cas-
cades, however, and the mountains of Idaho as well, as I am
told by Mr. Merton C. Lane, it is to be found far more fre-
quently. Here, close to timber line, where there is but little
litter and the old logs rest upon hard ground, they are readily
observed once the logs are turned over. The more alpine forms
are also less robust, narrower, and less brilliant. — Edwin C.
Van Dyke.
THE PAN-PACIFIC ENTOMOLOGIST
Published quarterly by the Pacific Coast Entomological Society
in co-operation with the California Academy of Sciences
E. P. Van Duzee^ Editor F. E. Blaisdell, M. D., Treasurer
Mr. Ralph Hopping, formerly an active member of the
Pacific Coast Entomological Society, but now Forest Entomolo-
gist for the Province of British Columbia, stationed at Vernon,
B. C., and Dr. J. M. Swain, Dominion Forest Entomologist
at Ottawa, were welcome visitors at the Academy of Sciences
in October. On their return from the South a week later, they
were guests of honor at a special meeting of the Society, at
which they gave an interesting account of the work in forest
entomology being done in the Dominion of Canada.
Mr. Dudley Moulton, Horticultural Commissioner of the
County of San Francisco, and well known as a careful student
of the Thysanoptera, has prepared and is sending free to ento-
mologists interested in collecting Thrips, a small “Thysanop-
tera Collecting Kit,” containing vials, alcohol, a brush and
memorandum sheets, and especially wishes to place them with
entomologists in foreign countries. If you desire one of these
collecting outfits, address Dudley Moulton, 244 California
street, San Francisco, California.
Mr. J. C. Hamlin writes us that an official notice, under date
of September 28, has been received from Mexico City, telling
of the death of Professor Jose Maria Gallegos, in the perform-
ance of his duty in Payo Obispo, Q. R., on September 23, 1925.
Professor Gallegos was a versatile student of nature, his col-
lecting showing a broad range of interest. His chief activities
were, perhaps, centered in the birds, in which field he exhib-
ited skill as a preparator rarely excelled, combined with a tire-
less energy. Much of his recent work had been directed
toward a study of the fauna of the State of Fower California,
and included work on the interesting insect fauna of that penin-
sula. Friends of Professor Gallegos will always remember him
as a gentleman endowed with a most pleasing personality. He
was a distinguished member of the Administration and of the
Society of Biological Sciences of Mexico. Both of these insti-
tutions are most unfortunate in the loss of this worker in the
prime of his life.
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From California Academy of Sciences, Proceedings, Series 4
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Address: California Academy of Sciences,
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Vol. II
Tanuarv, 1926
No. 3
THE
Pan-Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
HAMLIN, IMPORTANT OPUNTIA INSECTS OF THE UNITED STATES ... 97
BARNES AND BENJAMIN, NOTES AND NEW SPECIES 106
BARNES AND BENJAMIN, TWO NEW WESTERN PHAL^NID^ Ill
VAN DYKE, NEW SPECIES OF CARABID^ IN THE SUBFAMILY HARPALINJT;,
CHIEFLY FROM WESTERN NORTH AMERICA 113
FALL, LIST OF THE COLEOPTERA TAKEN IN ALASKA AND ADJACENT PARTS
OF THE YUKON TERRITORY IN 1924 - 127
MASON, TWO SPECIES OF THRIPS FROM CALIFORNIA 155
FLANDERS, NOTES ON PARASITES AT SATICOY, CALIFORNIA 157
KESEL, SEX-LIMITED POLYCHROMATISM IN LASIOPHTICUS PYRASTRI . . . 159
EDITORIAL 160
San Francisco, California
1926
THE PAN-PACIFIC ENTOMOLOGIST
Published quarterly in July, October, January and April by
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ENTOMOLOGIST
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'j
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Published at the California Academy of Sciences, Golden Gate Park, San
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Entered as second-class matter, February 10, 1925, at the postofSce at
San Francisco, California, under Act of August 24, 1912.
The Pan-Pacific Entomologist
Vol. II, No. 3 January, 1926
BIOLOGICAL NOTES ON IMPORTANT OPUNTIA
INSECTS OE THE UNITED STATES ^
BY JOHN C. HAMLIN
Fresno, California
INTRODUCTION
The general data recorded in this paper were secured dur-
ing the writer’s North American work in connection with the
Australian Commonwealth Prickly-pear Board’s project for the
biological control of the prickly-pear pests acclimated in Queens-
land and New South Wales. An earlier paper on this subject
was published in the July, 1925, number of this journal.
A Species Attacking the New Growth
Mimorista flavidissimalis Grote ^
DESCRIPTIVE
Adult. The adult, expanding about one inch, is bright yellow
with darker markings arranged in four irregular transverse
bands on the wings. These markings vary from an incon-
spicuous brown to a distinct blackish-brown color.
Egg. To the unaided eye the egg appears as a tiny, sub-
circular, irridescent disc attached to the flat surface of the joint.
The finely roughened surface is very slightly but regularly
convexed. The disc measures 0.75 mm. in diameter.
Larva. Eull-grown larvae measure about one-half inch in
length, and are of a yellowish-white color.
Pupa. The pupa is about two-fifths of an inch long, of a
polished light-brown color, with the head blackish. It rests
against the cactus joint and is covered by a whitish, convexed
cocoon of thin, tightly woven material. Pupation usually takes
place in wrinkles of dried joints.
LIFE HISTORY
The eggs are deposited singly or in small groups on the newly
formed joints, usually near the upper edges. Often two or
1 For determinations of species mentioned herein the author is in-
debted to the appropriate specialists of the Taxonomic Division, United
States Bureau of Entomology.
2 Order Lepidoptera, Family Pyralidae.
98
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3 ,
more eggs overlap. The extremes of the incubation period were
not determined, but in midsummer it is from three to five days.
Soon after hatching the larvae spin a delicate web, beneath
which they feed, either singly or several beneath one web. They
are not entirely external feeders, for they are frequently found
within the young joints or nopalitos.^ When fully developed, the
larvae spin their characteristic cocoons on the joints which have
succumbed to their attack, or occasionally on nearby living
joints. Duration of the larval and the pupal stages were not
independently determined.
During midsummer a generation is produced in four weeks.
The shortest life cycle was recorded at Uvalde, Texas, in
June, 1921. Newly emerged adults were caged on June 1 with
plants bearing nopalitos. The first moths of the succeeding
generation appeared on June 26, while the peak of emergence
was reached on June 29. From the four females originally
placed in this cage were reared 221 adults of the next genera-
tion. These figures indicate a minimum egg-deposition of fifty-
five per female, but the actual number laid was probably in
excess of the number of individuals successfully reaching the
imaginal condition.
In the lower Rio Grande Valley, where M. Uavidissimalis is
most abundant, the species appears to have about eight genera-
tions annually. The first generation is completed at Browns-
ville, Texas, early in April. The last generation of autumn, all
stages of which show a greatly retarded development, extends
over the mild winter. The pupal condition is undoubtedly the
stage in which the species passes the winter farther north.
HABITS AND FOOD PLANTS
During the day the adults rest on shaded parts of the plant.
At dusk they may be seen flying about among the cactus clumps,
usually alighting on the young joints. Mating and oviposition
have not been observed, but these activities evidently transpire
at night.
This species has never been found attacking any cacti other
than the platyopuntias.^ Moreover, in the field its attack was
observed to be confined to the young, tender joints of these
3 Nopalito is a Spanish term which refers to a definite stage in the
growth of the Opuntia joint, being applied to the tender, newly formed
joint before it has thrown off the rudimentary true leaves.
JANUARY, 1926 ] HAMLIN OPUNTIA INSECTS
99
cacti. Among the Opuntias, however, no restriction to species
was noted.
On one occasion a distinct variation in feeding habit was
observed in a cage poorly supplied with nopalitos. The heavy,
basal joints ceased putting forth new joints, leaving the large
number of larvae with an inadequate supply of dying nopalitos.
When the cage was dismantled all nopalitos were dead, being
very dry and hard. By chance, it was found that considerable
numbers of the larvae had successfully met this condition by
boring into the mature joints at the points of attachment of the
dried nopalitos. Some individuals had penetrated the tough
fiber to a depth of two inches, acting much as other internal
feeders.
INJURY
Owing to the small size and tenderness of the joints against
which Mimorista Paijidissimalis larvae direct their attack, these
parts are very quickly killed. Even when only a few larvae are
present their work weakens the attachment of the nopalito to
the plant and it falls to the ground where, if not completely
killed by the insects, it has not the substance to establish itself.
Occasionally, joints recover from the attacks of the larvae, but
they remain deformed. Such recovery is believed to be due to
the eggs having been laid on small young joints which had
toughened considerably.
The many generations during the growing season and its
attack against the new growth render this species a formidable
enemy of the prickly-pears in the region where it is abundant.
Were its distribution more general, it would rank among the
most important Opuntia insects of the United States.
DISTRIBUTION
Hunter et al.^ noted the species as having a restricted range,
recording it in Texas from Hallettsville and San Antonio,
southward, and in Arizona from Tucson. While the writer has
taken specimens at Uvalde, San Antonio, Victoria, and Browns-
ville, Texas, and at Piedras Negras, Mexico, the species has
been observed to be of importance only in the lower Rio Grande
Valley. This distribution seems certainly to be correlated with
4 The Principal Cactus Insects of the United States, by Hunter, W. D.,
Pratt, P. C., and Mitchell, J. D., U. S. D. A. Bur. Ent. Bull., 113, 1912.
100
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
moisture. Since young joints are required monthly through-
out the summer for oviposition and larval development, those
sections of the cactus country which are very dry during mid-
summer are not suited to continuous propagation of large
numbers of this species.
In this connection it is of interest to note that the so-called
prickly-pear rat {Neotoma micropus Baird), in pruning the
Opuntia plants during feeding, stimulates the production of
new joints which, occurring throughout the summer, constitute
a considerable factor in the propagation of this insect.
NATURAL ENEMIES
The foremost parasite of Mimorista Havidissimalis is a spe-
cies of Casinaria, probably infesta (Cresson). A collection of
100 Mimorista larvae from Brownsville, Texas, in June pro-
duced 20 of these ichneunionid parasites. Another parasite, less
abundant than the former, is Eiphosonia brucsi Cushman.
Cottony Cochineal Insects : Dactylopius spp.^
Two species of wild or cottony cochineal insects are impor-
tant enemies of Opuntia in the United States. These are Dacty-
lopius tomentosus Lamarck and D. confusus Cockerell. As
implied by the common names, these forms are distinguished
from the dye-producing species by the profuse covering of pure
white wax in contrast to the light, powdery covering of the
latter.
Dactylopius tomentosus (Lamarck)
The adult female is hidden by the tomentose wax covering.
These white clumps contain several female insects. The indi-
vidual clumps are of somewhat greater extent and are hardly
so symmetrical as are those of D. confusus Ckll, The insects
themselves are soft-bodied, rounded, and of a dull-reddish color.
The adult males are rarely seen in the field, although they are
produced in abundance. The body of the male is 1 mm. long,
while its length, inclusive of the wings, is nearly 2 mm. From
the posterior end of the abdomen project two slender processes
4 Order Hemiptera, Family Coccidee.
JANUARY, 1926] HAMLIN OPUNTIA INSECTS
101
slightly more than 2 mm, in length. The male is red, with the
wings and abdominal processes pure white.
SEASONAL HISTORY
The mature females produce living young which crawl about
over the plant until a suitable feeding place is found. Here
they insert their probosces and suck the juices of the plant.
After once becoming settled these “crawlers’’ are unable to
move to another location. The males are produced in small
white tubes which appear on end in the waxy covering of the
female. Several generations are produced during the summer.
It seems that a generation is produced in from five to seven
weeks. The crawlers of the last generation in autumn seek shel-
tered places to settle themselves for the winter. It has been
observed that the overwintering brood lives upon the roots or
stems beneath the surface of the ground or next to the ground
on prostrate-growing or fallen joints.
HABITS
The summer broods of crawlers travel upward and cluster
on the topmost spines of the plant upon which they were devel-
oped. Undoubtedly, this habit greatly influences the dissemi-
nation of the species, for such tiny creatures, with the few
strands of the tomentum already formed, would be carried very
considerable distances by the wind.
FOOD PLANTS
Dactylopius tomentosus Lamarck has been found only on
Opuntia. In the field, however, the species occurs as distinct
biological strains or races. Thus, cochineals have been found
feeding upon certain prickly-pears which refused to feed upon
certain other species. The Texas strain has been found to
prefer Opuntia lindheimeri Engelm., while the strain from
Tucson, Arizona, lives upon a white-spined Opuntia, and the
Chico, California, strain prefers prickly-pear different from
either of the former. After due time, and under the stress of
starvation, a very small percentage of a new generation may
be induced to settle upon a species of Opuntia which was pre-
viously refused when its favorite type was also available. While
it is known that these strains can adapt themselves to groups
of Opuntias other than those preferred, the extent of such capa-
bilities has not been determined. Structurally, these strains
102
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
were not distinguishable, and all were determined as belonging
to this species.
INJURY TO PLANT
The etfect of the sucking of these insects is to devitalize the
plant and finally to cause affected joints to dry out completely.
The new growth is effectually checked when these insects are
abundant, for the crawlers settle upon it in preference to the
older joints and quickly cause it to drop from the plant. After
its severance from the plant the insects, being unable to migrate,
continue their sucking until it is completely killed. Thus, the
injury of this cochineal is effected by preventing new growth
and by a gradual killing of the older terminal joints.
DISTRIBUTION
Dactylopius tomentosus Lamarck has been secured by the
writer from Chico, California; Tucson, Arizona; and Uvalde,
Texas. Hunter el alP recorded the species from New Mexico,
Arizona, and Mexico.
Dactylopius confusus (Cockerell)
This insect presents very much the same appearance as the
foregoing species, but the white clumps are somewhat smoother
and appear a little more rounded. Its seasonal history, habits,
and character of injury are very similar to those of its con-
genitor. This species seems to be even more closely restricted
in feeding habits than the preceding species. The writer has
collected it in Florida at Miami, Coconut Grove, Larkin, West
Palm Beach, Palatka, and Gainesville, and at Brownsville,
Texas.
NATURAL ENEMIES OF THE COCHINEALS
Aside from the eleven enemies recorded,® the writer has
found Zophodia quadric olorella (Dyar) ® to be a most impor-
tant predator in material from both Brownsville, Texas, and
Miami, Florida. The larvae of this moth burrow beneath the
waxy covering of the cochineal clumps and devour the soft-
bodied coccids. When fully developed, the predaceous larvae
spin cocoons beneath the cochineal wax, leaving no external
evidence that the waxy material of the former inhabitant now
5 U. S. D. A. Bur. Ent. Bull., 113, 1912.
6 This species is assigned to the genus Poujadia in the Barnes and Mc-
Dunnough catalogue, but Doctor Dyar tells me that it is a Phycetine and
should go in Zophodia.
JANUARY, 1926] HAMLIN OPUNTIA INSECTS
103
covers only the cocoons of a predator. As many as three
cocoons of this moth have been found under a single cochineal
clump, while, occasionally, four or five small larvse have been
found beneath a waxy mass feeding on the cochineals.
A Species Inhibiting the Development of Seeds
Asphondylia OPUNTIA Felt
The mosquito-like adult has never been seen in the field by
the writer. During March the puparia may be seen protrud-
ing from the previous season’s fruits, quite commonly upward
of fifty projecting from one fruit. The midges issue in the
canyon country north of Uvalde, Texas, during the latter half
of March. At this time the flowers of Opuntia have not opened.
In early May a slight yellowing of certain fruits indicates the
presence of larvae, which may be found in the interstices formed
by the rudimentary seeds. They are extremely small and are
barely distinguishable to the naked eye as tiny dots of light
yellow. There are no feeding marks to be seen on the seeds,
and the fruits show a normal degree of turgidity.
Later observations showed no change except a deepening of
the yellow color of affected fruits, and perhaps the production
of one or more small joints or fruits from the infested fruit.
In August the larvae were of the same size and color as when
first observed, and no damage to the seeds was apparent. The
seeds showed just the same state of development observed
in May. In October no changes in the larvae or seeds were
apparent. Likewise, in December no changes could be noted.
During midwinter the larvae evidently have a greatly acceler-
ated development, since they attain a fair size and destroy the
interior of the fruit just before the puparia protrude in March.
There is, then, only one generation annually. From the above
observations it seems that the larvae are greatly retarded in
their development, so that for about nine months they undergo
no perceptible changes. Moreover, it seems that their effect of
transforming the fruit functionally from a reproductive to a
vegetative part is accomplished without entering the seeds or
fleshy interior of the fruit. This inhibitory effect upon the
seeds may be caused by a secretion.
This species is widely distributed. It is recorded ® from
7 Order Diptera, Family Cecidomyiidae.
8 U. S. D. A. Bur. Ent. Bull., 113, 1912.
104
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
many Texas localities and ranges southward to San Luis Potosi,
Mexico, and westward to Los Angeles, California.
An Opuntia Longicorn
Moneilema crassa Le Conte ®
The large, robust, wingless adult is from three-fifths to four-
fifths of an inch in length. The female is shining black through-
out, while, anteriorly, the elytra of the male are sparsely mottled
with whitish.
The eggs are cylindrical with both ends abruptly and regu-
larly convexed, measuring 3.00 to 3.25 mm. in length and 1.50
to 1.75 mm. in width. They are deposited either on cactus or
soil where the plant and the earth make contact.
The larvae enter the plant just below the surface of the soil,
arranging their cells so that the excess moisture is absorbed
by the ground immediately adjacent. In many instances this
wet soil is shaped to form the lower part of the larval cell.
Pupation takes place in the earth just below the scene of the
larval activities. The spheroidal cocoon is slightly more than
one inch in diameter and is composed largely of earth, but may
contain a small amount of the fibers of the plant.
This Opuntia longicorn has two generations annually in the
Uvalde, Texas, region. The winter is passed in the larval stage
and the spring adults appear in greatest abundance during late
April and early May. Second generation adults appear in Sep-
tember. The adults live from two to three months in summer.
Various species of Opuntia and Echinocereus, called “pita-
haya,” are the plants in which this species breeds in Southwest
Texas. The adults feed preferably upon the young, tender
joints or nopalitos, but also quite commonly on the fruits and
older joints.
During the day the adults secrete themselves about the base
of the cactus clump or among the debris which constitutes the
massive covering of the pear-rat’s^® abode. With the approach
of dusk the first few beetles leave their shelter and crawl upon
the plants, where they feed and mate during the night. In the
morning, when the sun’s rays strike them, they descend to
cover. It was, therefore, found that the most effective period
of the day for collecting them is between dawn and about 8 a. m.
9 Order Coleoptera, Family Cerambycidse.
10 Neotoma micro-pus Baird.
JANUARY, 1926] HAMLIN OPUNTIA INSECTS
105
Indeed, with little effort one may capture during this brief
period more beetles than could be collected by several persons
during the day by the laborious method of chopping and mov-
ing the heavy Opuntia plants. On cloudy days the beetles
remain on the upper portions of the plant until much later in
the day.
On account of the semi-subterranean life of the larvae and
the nocturnal habits of the adults, these insects are always a
rather obscure factor in the cactus-insect complex, despite the
importance of their injury. Two reliable indications will serve
to reveal the presence of the insect. First, infested plants are
in a disheveled state, much fallen, and the basal portions are
in a state of decomposition. This condition results from the
larval attack and constitutes the principal injury accomplished
by this species. Second, the feeding marks of the adults on
the upper portions of the plant show the presence of the beetles
in the immediate vicinity, for, being dependent upon crawling
as a means of locomotion, they do not travel far. This gnaw-
ing of the joints is insignificant in comparison to the larval
damage.
The species was collected at many points in the field in the
Uvalde, Texas, region. Its habitat is the dry, rolling country
of Southwest Texas.
NATURAL ENEMIES
Two parasites of M. crassum have come under observation.
The first developed from larvae collected in the La Pryor,
Texas, district during December, 1920. During the following
spring three puparia of this dipterous parasite were found
within one dead larva. This Moneilema larva was destroyed
just prior to the pupation of the other individuals. Unfortu-
nately, these parasites were lost during the interval between
dismantling the Florida station and the belated establishment
of Texas headquarters.
During the autumn of 1922, Mr. E. Mortensen found at
Uvalde, Texas, a minute egg-parasite destroying practically all
of the eggs deposited by large stocks of caged beetles. This
parasite has been described as a new species under the name
of Oencyrtus moneilcmce Gahan.
11 Proc. Ent. Soc. Wash., Vol. 27, 1925.
106
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
NOTES AND NEW SPECIES (LEPID.,
PHALAENID^)
BY WILLIAM BARNES AND F. H. BENJAMIN
Decatur, Illinois
Rhizagrotis epipsilioides Barnes and Benjamin, sp. nov,*
Head and thorax cream-gray. Palpi similar, marked with black.
Fore wing: Ground color cream-gray, dusted with black; basal line
and dash obsolete; t. a. line black, diffuse, broad, indicated by pow-
dery black scales, inwardly oblique from costa, ragged, produced to
an inward point on median vein, thence strongly excurved to inner
margin; orbicular obsolete; reniform a powdery, black, nearly round,
mark; t. p. line black, powdery, produced to points on the veins,
excurved from costa to vein 4, thence incurved to inner margin;
veins marked by black scales; s. t. line diffuse, marked by blackish
scales; its course indeterminate other than approximately parallel
to the outer margin; a row of terminal black dots between the
veins; fringes luteous at base, followed by a fuscous line, then a
whitish line, then another fuscous line, and with pale tips. Hind
wing: Pale, whitish, the veins and outer margin suffused with fus-
cous; fringes luteous at base, a fuscous interline, and white tips.
Beneath: General coloration much as above, somewhat paler; with
an obsolescent blackish common line, on the hind wing marked on
the veins only; a diffuse discal blotch on fore wing, and an obsoles-
cent discal dot on hind wing. Expanse, 41 mm.
The roughened, although not tuberculate Irons, allies the in-
sect to certain Rhizagrotis species, notably R. polingi B. and
Benj., to which it seems allied by habitus. The rather rough
thorax suggests Epipsilia, but the vestiture is composed of bi-
and trifurcate narrow scales mixed with some hair. The present
species, like polingi, is another connecting link between the
genera of the “Euxoa^ and Lycophotia groups. European
workers would probably place it in Lycophotia or '"Agrotis,”
but because of the Irons we prefer to place it in Rhizagrotis .
Superficially, the appearance is very similar to smooth-looking
‘‘Agr Otis’’ aurulenta Sm,, but it may be distinguished by its
black-sided palpi, more roughened Irons, and broader-scaled
thoracic vestiture.
Locality: Vineyard, Utah.
Types: Holotype $, VI-11-18, unique.
Notes: In Barnes collection, Tom Spalding, collector.
* Indicates that the type was submitted to Dr. William Schaus in order
to be sure the species had not been described from the neotropics. Thanks
are due Dr. Schaus for his numerous kindnesses.
JANUARY, 1926] BARNES-BENJAMIN NEW MOTHS
107
“Agrotis” scaramangoides Barnes and Benjamin,
sp. nov.*
Antennae of male serrate and fasciculate. Head and collar rufous,
brown, luteous and gray, mixed, the collar tipped with purplish.
Thorax dull wood-brown. Abdomen similar, paler basally. Fore
wing; Ground color pale brownish cream with black and brown
powderings; costa narrowly edged with brownish-black, else the
costal region of the pale ground color, which is conspicuous only
in the filling of the stigmata and edging of the veins which are
narrowly marked by black; all transverse maculation obsolete; or-
bicular narrow, outwardly oblique from radius and obtusely angled
in cell, the lower part nearly erect; reniform nearly erect, narrow,
only slightly constricted and bent; cell before and between the stig-
mata black-filled; some black filling distad of the reniform; a black
broad streak below cell taking the place of the basal dash and clavi-
form; s. t. line indicated only by a slight darkening between the
veins and by the terminal portion of the wing being darker than
the general ground color; an obsolescent terminal row of dots be-
tween the veins; fringe yellowish at base, interlined with fuscous and
tipped with pale luteous. Hind wing: Nearly uniform brownish fus-
cous, with slightly darker veins, and somewhat paler basally; fringe
yellowish at base, obscurely interlined, and tipped with pale luteous.
Beneath: Fore wing nearly uniformly dull fuscous; hind wing
whitish, with darker veins, and a scattered fuscous suffusion which
considerably darkens the wing distally; fringes as on upper side.
Expanse, 38 mm.
Unallied to any North American insect known to us. In
appearance almost exactly like Warren’s figure of scaramangce,^
but differing by the terminal space being nearly as dark as the
s. t. dashes, and in that the black and pale lined veins 3 and 4
run to the outer margin, while the other veins have a similar
tendency.
We describe in ^'Agrotis” in the Hampsonian sense, pend-
ing a decision on the availability of certain Hubnerian genera
now being considered by the Internal Commission.
Locality: Bald Mountain, Ward, Colorado.
Types Holotype 6 , August, unique.
Notes : In Barnes collection, E. J. Oslar, collector.
POLIA FALSA Grt.
(partim 6 nec $ )
1880, Grote, Can. Ent., XII, 215, Perigea.
1893, Smith, Bull. U. S. N. M., XLIV, 153, Perigea.
1 Prom Turkestan, Mongolia, and Tibet; 1909, Warren, in Seitz, Macro-
lepid., Ill, 50, pi. XI, b.
108
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
1903, Dyar, Bull. U. S. N. M., LII, 111, No. 1120, Perigea.
1906, Hampson, Cat. Lep. Phal. B. M., VI, 388 ( 6 nec 9 ), Polia.
micta Hamp.
1918, Hampson, Nov. Zool., XXV, 117, Miselia.
In 1906 Hampson recognized a mixed type series of falsa
Grt. and restricted the name to the 6 type, renaming the spe-
cies represented by the 9 type Bryomima fallax. In Nov. ZooL,
1918, he attempted to change the name falsa back to Bryomima
(type 2 ) and renamed the 6 type Miselia micta, on the
grounds that the 6 type did not fit the original description in
regard to its hairy eyes.
We scarcely see how this enters the matter. When Hamp-
son first selected one species to hold the nam^ falsa out of a
mixed type series, he might well have selected the species best
fitting the original description. However, he chose the hairy-
eyed species to hold the name falsa, and we do not see how this
fixation can be discarded even by himself.
We might mention that we find nothing in the code which
states that a description must be either adequate or accurate,
and call attention to the fact that a number of authors have
erroneously described hairy-eyed forms as possessing smooth
eyes, and vice versa. In fact, the unique type of Grote’s ''Ma-
mestra'' ferrealis is in Oligia.^
We are retaining falsa for the hairy-eyed species and fallax
for the lashed-eyed species.
Septis ona Sm.
1909, Smith, Jour. N. Y. Ent. Soc., XVII, 58, Luperina.
1910, Hampson, Cat. Lep. Phal. B. M., IX, 499, pi. CXLVII, f. 20,
Parastichtis.
stygia Dyar.
1915, Dyar, Proc. U. S. N. M., XLVII, 377, Trachea.
Professor M. Draudt informed us of the probable synonomy
and asked that we investigate. Accordingly, a specimen of ona,
agreeing with the cotypes and series in the Barnes collection,
was submitted to Dr. Schaus to compare with type stygia in
the United States National Museum. Dr. Schaus states our
specimen “is a perfect match of stygiaB
S. ona Sm., burgessi Morr., and relicina Morr. are very close
allies. The abdomen of each is tufted, so that they seem out
2 See 1923, B. and Benj., Can. Ent., LV, pp. 264-265.
JANUARY, 1926] BARNES-BENJAMIN NEW MOTHS
109
of place in Luperina. We suggest Septis as a better placement,
although we might here mention that we see little to separate
Septis from Trachea.
Luperina enargia Barnes and Benjamin, sp. nov.
Head and thorax luteous, strongly tinged with rufous-purple. Fore
wing: Ground color luteous, tinged and irrorated with rufous-purple;
ordinary lines except the s. t. of rufous-purple, single; basal line
obsolescent; t. a. line in the form of three waves from costa to
inner margin; median shade broad, outwardly oblique from costa
through cell filling in space between the ill-defined, luteous, blotchy
stigmata, angled at base of reniform, thence inwardly oblique; t. p.
line outwardly oblique on costa, thence parallel to costa, thence out-
wardly oblique to a sharp point on vein 6, thence inwardly oblique
and produced to points on the veins, which are strongly marked
by rufous-purple; s. t. line pale, rather as a waved transverse of
ground color bordered by a slight powdering of rufous-purple scales
than as a real line; terminal line thin, more or less interrupted by
the veins; fringe rufous-purple, obscurely mottled and interlined,
paler basally. Hind wing: Black, tinged with rufous-purple, only a
slight trace of luteous on disc, fringe rufous-purple, paler basally.
Beneath, ground color luteous, strongly suffused with rufous-purple,
and powdered by black scales, a common black line across wings,
discal black dot on hind wing only; veins, especially of hind wing,
darkened. Expanse: 6, 37 mm.; 9, 34 mm.
Similar to Luperina venosa Sm. ; somewhat smaller, smoother
appearing ; the fore wing, with the fuscous scales and markings,
replaced by rufous-purple; the hind wing much darker, black
tinged with rufuus-purple, and nearly lacking yellowish on
upper side.
Type locality: Monachee Meadows, Tulare County, Cali-
fornia, 8000 feet.
Number and sexes of type: Holotype 6 and allotype $,
July 8-14.
Platyperigea camina Sm.
1894, Smith, Trans. Am. Ent. Soc., XXI, 60, pi. VI, f. 9, Platyperigea.
1903, Dyar, Bull. U. S. N. M., LI I, 109, No. 1096, Platyperigea.
1909, Hampson, Cat. Lep. Phal. B. M., VIII, 417, pi. CXXXIII, f. 4
(type), ignot., Petilampa.
The typical form from Colorado is much like Hampson’s
figure in markings, but the figure is too large and red. We par-
ticularly call attention to the dentate but evenly curved t. p. line
and the short dash in the cell which takes the place of the
orbicular. Besides a short series from several Colorado local!-
110
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
ties, the form is apparently represented in the Barnes collection
by single specimens from Montana and Wyoming.
Platyperigea camina race alpha Barnes and
Benjamin, nov.
A race, or very likely only a maculation form, with mark-
ings similar to those of typical camina, but with the black dash
which takes the place of the orbicular greatly elongated, dis-
tally nearly making contact with the reniform and continued
mesad to the base of the wing.
Type locality: Deer Creek, Provo Canon, Utah.
Number and sexes of types: Holotype 6, lO-IX-18; one 6
paratype, id., 22-VIII-13.
Notes : Barnes collection, from Tom Spalding.
Platyperigea camina race beta Barnes and
Benjamin, nov.
In general similar to typical camina, but the t. p. line when
visible is much more strongly dentate, its course less evenly
curved, a tooth produced inwardly nearly to the reniform and
another strong inward tooth in the submedian fold, nearly mak-
ing contact with a strong outward tooth on the t. a. line. T. a,
and t. p. lines distally marked with white. In many specimens
it is difficult to see the exact course of the lines, but the present
form is usually easily sorted on its decidedly strigate appear-
ance. Such differences of appearance and markings have nearly
universally been accepted as of specific value. Possibly beta is
a valid species. However, we prefer to call it a subspecies
(race) until there is further evidence of its relationship to
other described forms.
It is represented in the Barnes collection from Jemez Springs,
New Mexico, and Clark County, Nevada, as well as from the
type localities.
Type localities and number and sexes of types : Holotype 6 ,
Paradise, Arizona, 24-31 July; allotype 2, id., 1-7 October;
20 6 , 30 $ paratypes, id., various dates, August, September,
and October ; 1 6,5 2 paratypes, Palmerlee, Arizona, no
dates.
Notes : Types in Barnes collection ; 2 6,2 2 paratypes in
U. S. N. M.
JANUARY, 1926] BARNES-BENJAMIN TWO PHAL^NID^ 111
TWO NEW WESTERN PHAL^NID^
BY WILLIAM BARNES AND F. H. BENJAMIN
Decatur, Illinois
Polia wyatti Barnes and Benjamin, sp. nov.
Head, thorax, and fore wing pale gray, with a few luteous and
black scales; basal line faintly geminate, its mesial part deep black,
erect on costa, produced along median vein, inwardly oblique to
vein 1; a black dash from termination of basal line, in submedian
fold, through about one-third of the t. a. space; t. a. line geminate,
mesial part obsolescent, distal part black, outwardly oblique, waved
from costa to below cell, excurved to base of claviform, strongly
dentate inwardly on vein 1, thence strongly excurved to inner
margin; orbicular large, rounded, white, filled with gray, and out-
lined with black scales; two black spots on costa, one each side of
the orbicular; claviform large, outlined by black, with pale gray fill-
ing, extending to or nearly to the t. p. line; reniform large, irregular,
pale, faintly outlined by black with central pale crescent outlined by
gray, somewhat constricted in middle, distally making contact with
the t. p. line, mesially produced as a point on median vein; t, p.
geminate, its outer line obsolescent, inner line marked by black, pro-
duced to long points on the veins, oblique on costa, excurved to
veins 4 or 5, incurved to claviform, at the distal end of which it
forms a small w-mark, outwardly oblique to vein 1, inwardly oblique
to inner margin; s. t. line obsolescent, marked by some sagitate
black dashes in a course nearly parallel to the outer margin; a ter-
minal row of dots between the veins; fringe pale at base, interlined
and checkered by blackish. Hind wing fuscous, with paler, broad,
obsolescent median shade; with a thin dark terminal line; fringe
luteous with a fuscous interline. Beneath: Fore wing dull gray; hind
wing white, with a few scattered fuscous scales. Expanse, 40-44 mm.
A peculiar species, with a rather rough front, but obviously
not a Scotogramma. The habitus is that of a well-marked,
narrow-winged lustralis, but structurally nearer to insolens.
The eyes are strongly lashed from behind, and possibly a few
lashes from near the base of the antennae, suggestive of Lasi-
onycta. The fore tibiae are unarmed, and the tarsi are rather
heavily spined, but not heavily enough to prohibit placement in
Polia, which term we use in the sense of Hampson, knowing
that the correct name is probably Hadena.
Type locality : Fort Canby, Washington.
Number and sexes of types : Holotype $ , and 1 $ para-
type, VIII-7-12, H. Ramstadt collection.
Notes : Received through Mr. A. K. Wyatt of Chicago, Illi-
112
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
nois, for whom we take pleasure in naming the species. Mr.
Wyatt has kindly allowed us to retain the holotype.
Papaipema dribi Barnes and Benjamin, sp. nov.
Head and thorax purple-rufous; basal antennal segments white;
tegulce white-tipped. Fore wing: Ground color yellow, strongly
tinged with rufous and powdered with dull purplish scales; basal
spot white or purple-gray; subbasal area concolorous with median
area; subbasal line strongly excurved from costa to submedian fold,
bent inwardly at base of cell; t. a. area purple-gray above vein 1;
t. a. line double, angled inward on subcosta, thence outwardly ob-
lique to median vein, incurved in submedian space, strongly excurved
above inner margin; claviform indicated by a minute luteous-white
spot defined by purple-gray, below which is another minute spot
containing two white scales surrounded by purple-gray; orbicular
rounded, white, defined by purple-gray with a rufous distal point;
reniform large, oblique, constricted on inner side at middle, with a
luteous center defined by rufous and surrounded by white spots
defined by purple-gray; medial shade purple-gray, outwardly oblique
from costa, inwardly oblique from cell; t. p. line double, excurved
to about vein 4, incurved in submedian area; s. t. line irregular, its
course not well defined; an apical yellowish patch; terminal line thin,
purple-gray; fringe luteous at base, distally purple-gray, interlined
with luteous and purple-gray. Hind wing luteous-white, suffused
with rufous and purple-gray. Beneath: Luteous-white, tinged with
rufous and purple-gray. Expanse, 38 mm.
Allied to marginidens ; but with somewhat the habitus of
merriccata. The genitalia, examined superficially, are much like
those of marginidens , the clasper and ventral arm of the valve
appearing shorter and heavier.
Mr. Henry Bird has seen the specimen, which we had already
compared with the various Western types in the United States
National Museum, and agreed with us that a new species was
involved. When asked to describe it he declined, but very kindly
advised us to do so.
Type locality : High Rolls, New Mexico.
Number and sexes of types: Holotype 6, August, J. G.
Bonniwell collector.
Notes : The name hirdi having been used for a Papaipema,
v/e take pleasure in using the anogram drihi.
JANUARY, 1926] VAN DYKE HARPALIN^
113
NEW SPECIES OF CARABID^ IN THE SUBFAMILY
HARPALIN.E, CHIEFLY FROM WESTERN
NORTH AMERICA
BY EDWIN C. VAN DYKE
University of California, Berkeley, California
(Continued from page 76)
Pterostichus rufofemoralis Van Dyke, new species
Rather robust, somewhat flattened, black, with the base of first
antennal segment, coxae, trochanters and femora, except the apices,
an orange-red, the head and prothorax smooth and shining, the
elytra subopaque and alutaceous. Head large, considerably over one-
half breadth of prothorax and slightly longer than broad; frontal
grooves prominent; mandibles robust and of normal length; eyes
large; antennae moderately robust, extending slightly beyond the base
of prothorax. Prothorax robust, as broad as long, slightly narrowed
posteriorly, apex shallowly but evenly emarginate, base almost trans-
verse, sides distinctly arcuate throughout, hind angles rounded;
lateral margins moderate, reflexed and with well-marked flattened
fossae within, which broaden basally; disc convex, median longitu-
dinal groove deeply impressed and almost complete, anterior trans-
verse impression well defined, the posterior vague, basal impressions
single, broad and deep posteriorly and coarsely irregularly punctured,
a rounded tubercle to the outer side (in the paratype or second
specimen this is less evident), the base not margined. Elytra three-
fifths longer than head and prothorax combined, five-eighths as
broad as long, elliptical; humeri well rounded, humeral angles absent;
disc convex, striae deeply impressed, complete, uniform throughout,
finely rather closely punctured, intervals convex, slightly crenulate
and vaguely rugose near middle; third with four shallow foveae, the
first about one-fourth distant from base, the second near the middle
and the others closer together and not far from the apex; scutellar
striae rather long and on either side of the suture at base. Legs
fairly robust; first segment of posterior tarsi obscurely grooved at
base on either side; posterior trochanters of moderate size. Apex of
prosternum not margined. Beneath, with the sides of head finely
rugose, the lateral thoracic sclerites and sides of first two abdominal
segments rather coarsely punctured, the median portion of body with
the abdomen posteriorly smooth and shining. Length, 13 mm.;
breadth, 5 mm.
Holotype, female (No. 1853, Mus. Calif. Acad. Sci.), and
paratype female in my collection, captured on Iron Creek, sixty
miles north of Nome, Alaska, August 23, 1913, by J. Aug.
Kusche and by him kindly presented to me.
This interesting and most distinct species should be associated
114
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
with Pterostichus punctatissimus Rand. It has the same general
facies, and is about the size of a small specimen of the latter.
It is in general less robust, its head and prothorax smaller, the
elytra more elliptical, the intervals regular, and the thighs a
clear orange-red instead of black. The males will, no doubt,
be found to have the elytra more shining.
Pterostichus pacificus Van Dyke, new species
Small, robust yet graceful, shining, head and prothorax black,
elytra piceous brown, mouthparts, antennae, legs, prothoracic epi-
pleurae and epipleurae rufopiceous. Head rather large, slightly more
than three-fifths as broad as prothorax; frontal grooves well im-
pressed, non-punctate; mandibles of type usual in subgenus Cryobius;
eyes moderately prominent (less convex than in riparius) ; antennae
rather slender, extending beyond hind angles of prothorax. Pro-
thorax slightly broader than long; apex emarginate; anterior angles
prominent, base slightly emarginate at middle; sides narrowly arcu-
ate from apex to one-fourth distance from base, there sinuate and
straight to hind angles which are right-angled; lateral margin well
defined and markedly reflexed; disc convex, median longitudinal
groove sharply defined and almost complete, anterior transverse im-
pression vague, the posterior distinctly defined; basal impressions
each a single flat-bottomed pit projecting forward along the inner
margin, rugosely punctured at bottom, without distinct carina near
hind angles, base with indistinct margin near angles. Elytra one-
fifth longer than head and prothorax combined, three-fifths as broad
as long; humeri distinct though rounded, the sides moderately arcu-
ate and rounded to sutural angles as usual, the margin fine; disc con-
vex, slightly flattened at middle; striae clearly defined and complete,
more deeply impressed basally and apically, punctuation obscure,
intervals flattened at middle, somewhat convex near base and apex;
third stria with a well-defined fovea one-third distance from base
and another one-fourth distance from apex, scutellar stria rather
long and well impressed. Legs slender, first three segments of hind
tarsi deeply grooved on outer sides. Apex of prosternum not mar-
gined. Beneath, with side pieces of meso and metathorax shallowly
and sparsely punctured, the first and second abdominal segments
also vaguely punctured at sides, elsewhere smooth and shining.
Length, 9 mm.; breadth, 3.25 mm.
Holotype, male (No. 1854, Mus. Calif. Acad. Sci.), a unique
in my collection, captured by myself near Hoquiam, Washing-
ton, May 27, 1914.
This species belongs in the subgenus Cryohius Chaud., and is
related to riparius ® and its associates fatuus and pinguedineus,
8 See my notes concerning' these in “The Coleoptera collected by the
Katmai Expedition,” Nat. Geograph. Soc. Contrib. Tech. Papers, Vol. II,
No. 1 (1924), p. 9.
JANUARY, 1926] VAN DYKE HARPALIN^
115
but it differs from these by having the prothorax more elon-
gate, the anterior prothoracic angles more prominent, the sides
posteriorly parallel for a considerable distance in front of the
hind angles, the outer basal impression merged with the inner
in one deep pit, not separate anteriorly as in those, the lateral
margin of both prothorax and elytra more distinctly defined,
the scutellar striae longer, the punctures of the striae almost
obliterated, and the antennae and legs longer and more delicate.
The shape of the beetle, especially that of the prothorax and
elytra, strongly suggests some of the species of Bembidium like
funereum and mceklini.
Anilloferonia Van Dyke, new genus
Body elongate, moderately flattened and parallel. Head moderate
in size, somewhat constricted behind; eyes wanting; genae slightly
prominent, two supra-orbital setae; frontal grooves well developed;
frontal suture distinct; clypeus prolonged beyond base of mandibles;
mandibles porrect, without setae in scrobes; antennae arising under
a distinct frontal ridge, well developed, the three basal segments
glabrous, outer segments somewhat enlarged; submentum biemargi-
nate and with well-defined tooth at middle; maxillae with laciniae
hooked at tip, the maxillary palpi of moderate length, the last seg-
ment elongate conical, the penultimate about two-thirds length of
preceding and obconical; ligula of moderate size and bisetose at tip;
labial palpi with the last two segments of about equal length, the
penultimate bisetose in front. Prothorax with lateral setae one-third
distant from anterior angles and others within posterior angles.
Body not pedunculate. Elytra narrowly indexed, margin interrupted
posteriorly and with evident internal plica, disc without dorsal punc-
tures. Apterous. Prosternum not prolonged at tip and not margined
at apex. Mesosternum almost vertical in front and rather widely sep-
arating the coxae, the epimera narrow, yet wider internally. Metas-
ternum well developed, the episterna about a third longer than broad,
the epimera almost one-half the width of the former; posterior coxae
contiguous. Legs well developed, femorae not greatly enlarged, pos-
terior trochanters large and somewhat elliptical in shape; anterior
tibiae deeply emarginate within and with inner spurs at summit of
emarginations.
Genotype; Anilloferonia testacea Van Dyke, new species.
This genus, according to the characters, should be placed in
the tribe Pterostichini as constituted by Horn,® and close to the
genus Pterostichus. It differs primarily from that by its having
9 “On the Genera of Carabidse, with special- reference to the fauna of
Boreal America,” by George H. Horn, Trans. Am. Bnt. Soc., Vol. IX
(1881), p. 136.
116
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
no vestige whatever of eyes and by the outer segments of the
maxillary palpi being differently proportioned. Superficially, it
looks like a diminutive specimen of one of the species close to
angustus or like a giant Anillus, particularly as regards the
shape of the head.
Anilloferonia testacea Van Dyke, new species
Testaceous. Head one-third longer than broad, one-third as broad
as prothorax, smooth and shining; antennae extending slightly be-
yond posterior thoracic angles. Prothorax about as wide as long,
apex and base feebly emarginate; sides feebly arcuate for anterior
half, straight and slightly convergent posteriorly; hind angles barely
obtuse, lateral margin fine; disc slightly convex anteriorly, flattened
in region of hind angles; median longitudinal groove distinct, yet
fine and almost complete; anterior transverse impression indicated
only by series of small black spots; posterior impression vague;
basal impressions single, not well defined; basal area obscurely punc-
tate; basal margin only faintly indicated at middle. Elytra almost a
third longer than head and prothorax combined, a third longer than
broad, elongate elliptical; humeri rounded; lateral margin fine; disc
feebly convex, flattened at center; striae nine in number, fine yet
complete, very finely and not closely punctured, the eighth with five
well-marked fovese near humeri and five near apex, the seventh with
one fovea near apex, the intervals flat and without punctures; scu-
tellar striae wanting. Beneath smooth and shining, except side pieces
of meso and metathorax and first two abdominal segments which are
obscurely punctate-rugose. Length, 5.5 mm.; breadth, 2 mm.
Holotype, female (No. 1855, Mus. Calif. Acad. Sci.), a
unique captured by myself in the Yakima Indian Forest Re-
serve on the east side of Mount Adams, Washington, at an
altitude of about 4000 feet, July 1, 1925. It was found beneath
an old log in the forest.
This and two species of Anillus are the only completely eye-
less Carabidae which have been found on the Pacific Coast.
Evarthrus montanus Van Dyke, new species
Elongate, robust, black, head and prothorax somewhat shining,
elytra subopaque, slightly more pronounced in female. Head large,
almost four-fifths breadth of prothorax; frontal grooves well marked;
eyes prominent; mandibles robust and projecting; antennas extend-
ing a couple of segments beyond hind angles of prothorax. Pro-
thorax about 1 mm. broader than long, apex feebly emarginate, base
more distinctly so and about equal in breadth to apex; sides slightly
arcuate and convergent posteriorly until near hind angles where
JANUARY, 1926] VAN DYKE HARPALIN^
117
they become gradually sinuate, then straight to the angles which are
well defined, slightly obtuse and blunt at apex; lateral margin well
defined, rather narrow in front but gradually broader posteriorly;
disc convex, with series of fine transverse rugae near middle; median
longitudinal goove linear and well impressed, almost complete pos-
teriorly and ending abruptly anteriorly at transverse impression;
anterior transverse impression sharply defined and complete, the
posterior not definable; basal impressions double, the inner deep and
broad and each with a faint fovea on inner margin, the outer like-
wise deep but more linear, with a fairly well-defined carina to the
outer side of it. Elytra 2 mm. longer than head and prothorax com-
bined, one-sixth broader than prothorax and over two-thirds longer
than broad; humeri pronounced though rounded; sides moderately
arcuate and distinctly but shallowly sinuate before apex, disc convex
though slightly flattened toward base; striae complete, well impressed
anteriorly, less so posteriorly, finely rather closely and distinctly
punctured anteriorly, the punctures gradually disappearing toward
apex; intervals somewhat flattened, the third with a single fovea
back of middle and close to second stria, the seventh distinctly ele-
vated at base; scutellar striae vaguely defined at base of second
interval. Legs moderately robust, hind tarsi without lateral grooves
externally. Prosternum deeply grooved and not margined at apex.
Beneath moderately smooth and shining. Length, 18 mm.; breadth,
6.5 mm.
Holotype, male (No. 1856), allotype, female (No. 1857, Mus.
Calif. Acad. Sci.), and several paratypes in my collection, cap-
tured by myself in the valley at the base of the Black Moun-
tains, North Carolina, during July, 1902, I have a series of
seventeen specimens. It was not uncommon, and both Mr.
William Beutenmueller, my traveling companion, and I took
them frequently. The females are often much broader and more
robust than the males. A pair of paratypes will be deposited in
the United States National Museum at Washington, D. C.
This is a form that is characteristic of the mountains, I have
always associated it with Evarthrus sigillatus Say, which, though
found on both sides of the Alleghanies, is generally more
northern in distribution. I believe that it is derived from the
same common stock, but it is markedly different in appearance,
even when viewed superficially, for it is considerably larger
and more elongate, the prothorax especially so. The prothorax
is also more flattened basally, the sides distinctly though gradu-
ally sinuate posteriorly, the hind angles definitely angulated,
and the basal impressions less separated. The elytra are more
118
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
elongate, more flattened basally and, as a rule, with the striae
more impressed anteriorly.
Evarthms vagans alabamae Van Dyke, new subspecies
This name I am proposing for a very distinct phase of the
species which is characterized by being of large size, 17 mm.
long by 7 mm. in breadth, resembling in this regard the larger
phase generally known as engelmanni, and differing from this
primarily by having the sides of the prothorax evenly arcuate
to hind angles, not at all sinuate posteriorly, the hind angles as
a result very obtuse, the lateral margin more broadly expanded
posteriorly, the basal impressions more distinctly separated, and
the carinae near the hind angles short and rather poorly defined,
not long and conspicuous as in the other.
Holotype, male (No. 1858), allotype, female (No. 1859, Mus.
Calif. Acad. Sci.), and five paratypes in my collection, all
captured near Mobile, Alabama, by Mr. H. P. Loding. Two
other specimens, one from Shreveport, Louisiana, and the other
from Hart, Louisiana, agree with these.
Evarthrus lodingi Van Dyke, new species
Large, elongate, robust, black, rather shining throughout. Head
two-thirds as broad as prothorax, somewhat elongate; frontal
grooves deeply impressed; frontal suture distinct but fine; eyes
prominent; mandibles robust, elongated. Prothorax over 1 mm.
broader than long, apex feebly emarginate; base slightly narrower
than apex, emarginate, narrowly margined close to angles; sides
rather broadly arcuate and convergent posteriorly until close to base,
where they become abruptly sinuate, then oblique to hind angles,
which are well defined though rounded; lateral margin fine, uniform
and well reflexed; disc moderately convex, smooth; median longitu-
dinal groove fine, distinctly impressed, extending from apex to near
base and with small foveate enlargement near apex and a very large
and deep one near base; transverse impressions not defined; basal
impressions treble, a deep foveate inner one, a deep, long linear
median one, well separated from the first, and an oblique shorter
linear outer one joining basally with the second, a well-defined
Carina at the outer side of the last. Elytra one-fourth longer than
head and prothorax combined, over 1 mm. broader than prothorax
and five-twelfths longer than broad, subelliptical; humeri prominent
but rounded; margin rather broad, sharp and distinctly sinuate near
apex; disc moderately convex, striae complete, very deeply impressed,
finer apically, very finely closely punctured, the punctures gradually
obliterated toward apex; scutellar stria long, deeply impressed and
within base of first interval; intervals convex, the third with fovea
JANUARY, 1926] VAN DYKE HARPALIN^
119
about one-third distant from apex, the seventh conspicuously ele-
vated at base. Legs long yet robust, hind trochanters broad, flat-
tened, and subacute at apex; hind tarsi without lateral grooves exter-
nally. Prosternum shallowly grooved and not margined at apex.
Beneath smooth and shining. Length, 20 mm.; breadth, 7.5 mm.
Holotype, male (No. 1860, Mus. Calif. Acad. Sci.), in my
collection, captured at Monte Sano, Madison County, Alabama,
June 9-11, 1911, by Mr. H. P. Coding and by him very kindly
presented to me. Other specimens are in Mr. Coding’s collec-
tion. The species is named after the captor, a kind and generous
friend who has done many favors for me and has contributed
much toward making known the Coleopterous fauna of his
region.
This very distinct and elegant species should, perhaps, be
placed somewhere near sodalis. It is much like a giant phase
of the latter, with the hind angles of the prothorax of about
the same size but more rounded. Its size, graceful form, three
series of distinct basal thoracic impressions, long deep scutellar
stride, deeply impressed striae and convex intervals should set
it well apart from all known species. According to the Ceng
catalogue and the works of Colonel Casey this species would
fall in the genus Eumolops Casey, but that genus is not founded
upon any character of good generic value, therefore I feel that
it should not be recognized.
Platynus prognathus Van Dyke, new species
Elongate, subparallel, flattened, nigropiceous, margins of prothorax,
elytra, basal segments of antennae, mouthparts, legs, and sides of
abdomen testaceous. Head much elongated, with mandibles longer
than prothorax and almost as broad; frontal impressions large,
irregularly and shallowly impressed; clypeus bifoveate in front; eyes
large and moderately convex; mandibles very long and sickle-shaped,
as long as rest of head; antennae long, terminal segments enlarged,
the last three extending beyond base of prothorax. Prothorax one-
fifth broader than long; apex feebly emarginate; base transverse and
equal in breadth to apex, vaguely margined, sides arcuate for anterior
half, straight and oblique posteriorly, the hind angles very obtuse
and rounded and in advance of the basal margin; lateral margin of
moderate width, slightly reflexed and with broad, shallow fossae
within posteriorly; disc slightly convex; median longitudinal groove
distinctly impressed, the anterior transverse impression short but
10 “Memoirs on the Coleoptera,” by Thomas L. Casey, Vol. VIII (1918),
p. 351.
120
THE PAN-PACIFIC ENTOMOLOGIST [VOL, II, NO. 3
deep, the posterior vague, the surface finely transversely rugose.
Elytra over a third longer than head and prothorax combined, one-
third broader than prothorax, over twice as long as broad, elongate
elliptical; humeri well rounded; sides almost parallel, deeply sinuate
posteriorly, the margin fine; disc quite flattened, striae complete,
deeply impressed, punctures very fine and obscure, intervals flat-
tened, the second, fifth, and seventh somewhat broader; the narrower
subconvex third with three punctures near the second stria, the
anterior about at middle; scutellar stria rather long and well im-
pressed. Legs of moderate length. Beneath rather smooth and
shining.
Holotype, female (No. 1861, Mus. Calif. Acad. Sci.), in my
collection, captured on St. Simon Island, Okefinokee Swamp,
Georgia, April 27 to May 12, 1911, by Professor J. C. Bradley,
and by him very kindly presented to me.
This anomalous species might be placed in the group with
cincticollis, but is widely separated from that, as well as from
all of our other species. Its elongate, parallel form, very long
mandibles and somewhat clubbed antennae give it an appear-
ance all its own. Though this species would be classed as an
Anchomenus, I consider that as but a subgenus at most of
Platynus, in this regard following such able students of the
Carabidae as Ganglbauer and Le Conte and Horn.
Colpodes rufiventris Van Dyke, new species
Moderately elongate, head and prothorax decidedly narrowed, flat-
tened, nigropiceous, elytra somewhat darker, antennae, mouthparts,
legs, undersurface, and margins of elytra rufous with a slight piceous
tinge; head and prothorax shining, elytra subopaque. Head about
1 mm. less in breadth than prothorax and about three-fifths as broad
as long; frontal grooves fine; front flattened at outer side of grooves;
eyes of but moderate size; mandibles long and prominent; antennae
long and delicate, extending almost to middle of elytra; submentum
with simple tooth. Prothorax slightly broader than long, about one-
half the breadth of elytra; apex slightly emarginate; frontal angles
not prominent; base feebly arcuate, vaguely margined; sides evenly
but shallowly arcuate from apex to base, the hind angles broadly
rounded, the margin narrow, broader posteriorly and slightly re-
flexed; disc convex, with vague but fine transverse rugae; longitu-
dinal groove finely but distinctly impressed; anterior transverse im-
pression well, though not sharply, impressed, the basal a shallow
sulcus. Elytra a third longer than head and prothorax combined,
a third longer than broad, elongate ovate, broadest back of middle;
base deeply emarginate, the humeri as a result prominent, subangu-
11 ‘‘Die Kafer von Mitteleuropa,” by Ludwig Ganglbauer, Vol. I, p. 248.
JANUARY, 1926] VAN DYKE HARPALIN^
121
late; lateral margin narrow in front, broader posteriorly; apices
rounded; disc slightly convex, striae finely impressed, without dis-
tinct punctures; scutellar stria very long; intervals flat, the third
with an anterior very vague puncture near third stria, a vague punc-
ture near second stria at middle and an evident puncture near second
stria not far from apex. Legs rather long and delicate; anterior
tibiae shallowly grooved on outer face, middle and posterior deeply
grooved on both outer and inner face posteriorly, with caring be-
tween; first and second segments of front tarsi shallowly grooved
on both sides above; first three segments of middle and hind tarsi
rather deeply grooved on both sides above; fourth segment of all
tarsi narrowly bilobed posteriorly, with a slight tuft of pubescence
beneath. Beneath rather smooth and shining. Length, 10.5 mm.;
breadth, 4 mm.
Holotype, male (No. 1862, Mus. Calif. Acad. Sci.), in my
collection, captured at an altitude of 6000 feet on Mount
Washington, near Nogales, Arizona, July 10, 1919, by Mr.
J. Aug. Kusche, and by him kindly presented to me.
This, the second species of Colpodes to be described from
our fauna, differs from Colpodes longiceps Schfr., also from
Arizona, markedly in color as in other features. It does not
agree either with the descriptions of any of the known Mexi-
can species. Except for the rather wide margin of the pro-
thorax, this species might fall in the genus Glyptolenus Bates,
because of the tibial and tarsal characters, but these are quite
variable within the genus Colpodes, therefore not reliable for
establishing a group of more than subgeneric rank at most.
The genus Colpodes is sometimes placed as but a subgenus of
Platynus, but I think that though the character used for its
separation, the deeply emarginate or bilobed fourth tarsal seg-
ment, is not a very marked one ; it is sufficiently definite, espe-
cially when taken in connection with other characters and
backed by the fact that the genus as a whole is more or less
arboreal and generally restricted to the warmer parts of the
world, whereas almost all of Platynus are strictly terrestrial
and confined to the temperate regions.
Pseudaptinus oviceps. Van Dyke, new species
Elongate, depressed, pubescent, rufotestaceous, antennae, palpi, and
legs testaceous. Head elongate oval, one-fourth longer than broad
and three-fourths as broad as prothorax; finely sparsely punctate,
rather shining; frontal impressions well marked, rather broad; eyes
small, but slightly convex and hardly projecting beyond the even
122
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
curve of the side of the head; mandibles prominent, prognathous;
antennae long and slender, extending several segments beyond the
base of the prothorax. Prothorax a little longer than wide, broadest
near apex; apex very shallowly emarginate; base slightly arcuate,
about one-half breadth of widest portion; sides broadly rounded near
anterior angles, oblique and convergent to posterior third, then sub-
parallel to hind angles which are right-angled but not prominent;
disc moderately convex, somewhat coarsely and sparsely punctured;
median longitudinal groove distinctly impressed; anterior transverse
impression well defined, posterior vague; basal impressions long and
shallow. Elytra oblong elongate, humeri broadly rounded, apex trun-
cate, outer angles rounded, disc flattened; striae finely impressed,
complete; intervals flat, finely and densely punctured, somewhat
rugose toward base. Legs long and delicate. Prosternum sparsely
and coarsely punctured, smooth at sides, meso and metapleurae
coarsely closely punctured, the abdomen very finely punctured.
Length, 6 mm.; breadth, 2 mm.
Holotype, female (No. 1863, Mus. Calif. Acad. Sci.), in my
collection, captured by myself about 1889, under a stone near
the Los Angeles River, California, in what is now Griffith Park.
This species in size and general appearance looks very much
like Pseudaptinus tenuicollis (Lee.), but its ovate and nar-
rower head, small eyes, and prominent mandibles separate it
very decidedly from that as well as from Pseudaptinus lecontei
(Dej,).
Thalpius microcephalus Van Dyke, new species
Elongate, depressed, pubescent, rufotestaceous, antennae, palpi, and
legs testaceous. Head elongate, one-fourth longer than broad and
narrower than prothorax, moderately coarsely, sparsely punctured,
somewhat shining; frontal impressions shallow, poorly defined; eyes
rather small, not projecting laterally beyond sides of head; mandi-
bles of usual size; antennae of moderate length, extending somewhat
beyond base of prothorax, segments four to ten oblong. Prothorax
about as long as wide, broadest one-third distance from apex; apex
slightly emarginate, anterior angles projecting forward somewhat
(much more evident than in horn't) ; base pedunculate as usual; sides
shallowly arcuate anteriorly, oblique and sinuate posteriorly, hind
angles acute; disc slightly convex, flattened medially, coarsely, mod-
erately closely punctured; median longitudinal groove sharply im-
pressed; anterior transverse impression well defined, posterior vaguely
marked; basal impressions of moderate length, shallow. Elytra ob-
long, elongate; humeri broadly rounded; apex truncate, outer angles
rounded; disc flattened; striae shallowly impressed, not distinctly
defined, especially posteriorly; intervals flat, rather coarsely and
irregularly punctured. Legs moderately long. Body beneath coarsely,
JANUARY, 1926] VAN DYKE HARPALIN^
123
rather regularly punctured anteriorly, abdomen more finely punc-
tured. Length 4.75 mm.; breadth, 1.75 mm.
Holotype, a unique female (No. 1864, Mus. Calif. Acad.
Sci.), in my collection, secured by myself from beneath a
stone on the hills in Griffith Park, Los Angeles, California,
January 12, 1912.
This species has the same general body proportions as Thal-
pius horni Chd., but it is much smaller. Its narrow head, less
prominent eyes and poorly defined elytral striae also serve to
distinguish it from that as from all other species. In color, it
resembles rufulus, but it is smaller and has a narrower pro-
thorax in addition to the characters just mentioned.
In this connection I wish to state that in Southeastern Cali-
fornia specimens of Thalpius horni Chd., which are entirely
rufous, are not uncommon.
Harpalus washingtoniensis Van Dyke, new species
Robust, black or slightly brownish. Elytra piceous brown, an-
tennae, mouthparts and legs piceotestaceous, tarsi darker, very shin-
ing, elytra of females duller. Head large, over two-thirds breadth
of prothorax, smooth and shining; frontal suture deeply impressed;
eyes moderately prominent; mandibles well developed; antennae
reaching to hind angles of prothorax. Prothorax almost a third
broader than long; apex and base slightly emarginate, the latter as
broad as apex and distinctly and completely margined; sides moder-
ately and evenly arcuate from anterior angles to posterior fourth,
then sinuate and convergent to hind angles, which are obtuse, almost
right-angled, and sharp at apex; lateral margin fine; disc convex,
median longitudinal groove distinctly impressed; anterior transverse
impression distinct, posterior vague, basal impressions deep; surface
smooth and shining, basal area irregularly punctured, more definitely
so in and about basal impressions. Elytra over one-fifth longer than
head and prothorax combined, two-fifths longer than broad; humeri
rounded; sides slightly arcuate, markedly excised near apex, margin
fine; disc decidedly convex, strise well impressed; scutellar stria
rather long; intervals flattened, somewhat convex basally in males.
Legs robust. Beneath finely, sparsely and rather indistinctly punc-
tured in front, the abdomen smooth and shining, except for sides of
first and base of second segment which are slightly punctured.
Length, 8.5 mm,; breadth, 3.25 mm.
Holotype, male (No. 1865), allotype, female (No. 1866, Mus.
Calif. Acad. Sci.), and numerous paratypes in my collection,
captured by myself at Port Angeles, Washington, May 26,
1907. In my series of thirty-two mounted specimens, I have
124
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
them from the following additional localities : Nanaimo, British
Columbia ; Monroe, Lake Quiniault, Seattle, The Forks, North-
bend, and Orcus Island, Washington ; and Marshfield, Oregon.
A pair of paratypes will be deposited in the entomological
collection of the U. S. National Museum, Washington, D. C.
This species varies considerably in size, ranging in length
from 7 to 10 mm., the females are, as a rule, larger than the
males, and somewhat paler in color, the epipleurae often lighter,
while in a few cases the legs are quite piceous. It belongs in
association with Harpalus spadiceus Dej. and Harpalus carolincB
Schaeffer, differing primarily from the former by having the
prothorax more sinuate posteriorly and the hind angles almost
right and sharply defined, and from the latter by having the
legs yellowish or testaceous, the general body color, especially
the elytra, more brownish, the basal impressions of the pro-
thorax more broadly impressed, the basal area itself definitely
punctate, the elytral margin more acutely excised posteriorly,
and by being as a whole somewhat smaller and less elongate.
Harpalus martini Van Dyke, new species
Robust, elongate, subparallel, black, antennae, tibiae, and tarsi some-
what piceotestaceous, very shining above in the male, the female
quite opaque. Head large, two-thirds breadth of prothorax; frontal
suture very fine, not impressed; eyes moderately prominent; mandi-
bles well developed; antennae reaching to hind angles of prothorax.
Prothorax over a third broader than long; apex distinctly emargi-
nate, the anterior angles slightly prominent; base also distinctly
emarginate, completely margined and broader than apex; sides very
shallowly arcuate, almost subparallel from anterior angles to pos-
terior fourth, thence sinuate and divergent to posterior angles, which
are almost right-angled and slightly rounded at apex; lateral margin
fine; disc convex, median longitudinal groove very fine, anterior
and posterior transverse impressions both vague, basal impressions
broadly and shallowly impressed; posterior angles much flattened;
basal area irregularly punctured, especially about basal impressions.
Elytra one-fifth longer than head and prothorax combined and one-
fifth longer than broad; humeral angles distinctly dentate; sides
slightly arcuate, almost parallel, shallowly excised near apex, margin
fine; disc convex, striae finely impressed; scutellar stria rather long;
intervals flat. Legs robust. Under surface subopaque anteriorly;
abdomen posteriorly smooth and shining in male, subopaque in
female; an oblique area on second and third abdominal segments,
immediately beneath hind femora, finely punctate and with short
hairs arising from each puncture. Length, 10 mm.; breadth, 3.75 mm.
JANUARY, 1926] VAN DYKE HARPALINiE
125
Holotype, male (No. 1867), and allotype, female (No. 1868,
Mus. Calif. Acad. Sci.), collected at Bear Lake, San Bernar«
dino Mountains, California, May 5, 1919, by Mr. J. O. Martin,
and kindly presented to me. Four paratypes are in Mr. Martin’s
collection. This species is named after Mr. Martin, an excellent
collector, in consideration of many favors.
The species is probably related to Harpalus somnulentus Dej.,
but is decidedly darker. Its color and general shape also sug-
gest Harpalus cautus Dej. Its peculiarly shaped prothorax, with
base broader than apex, sides subparallel, sinuate and divergent
posteriorly, and the well-marked humeral dentations of the
elytra should, however, readily serve to separate it from the
above and from all of our other species.
Dicheirus blaisdelli Van Dyke, new species
Robust, very convex; head and prothorax rufo-piceous, elytra
black, legs rufous; clothed above with long erect hair, the sternal
and midventral region as well as the legs also very pilose, the hair
in the main shorter than above. Head very coarsely, moderately
densely and closely punctured, antennas about reaching hind angles
of prothorax. Prothorax distinctly broader than long, 3.5 mm. by
3 mm., emarginate at apex, transverse at base, the latter 2.5 mm.
wide, sides broadly rounded in front, straight and convergent pos-
teriorly, just perceptibly sinuate in front of hind angles, the hind
angles obtuse and distinct, not rounded, the disc convex, coarsely
deeply and rather sparsely punctured. Elytra over one-third longer
than broad, .5 mm. broader than prothorax, sides hardly arcuate,
evenly rounded at apex, posterior sinuation faint; disc very convex,
striae distinctly impressed, impunctate, intervals flat and shining,
sparsely irregularly biseriately" punctured, deeper toward base, very
finely near apex. Prosternum coarsely moderately closely punctured,
propleurae finely sparsely and rather indistinctly punctured, the meso-
and meta-pleurae more distinctly punctured and somewhat rugose,
the abdomen finely and closely punctured at middle in front, very
finely and sparsely at middle behind, with sides impunctate. Legs
robust, spines of anterior tibiae trifid, of posterior slender and acute.
Length, 11 mm.; breadth, 4 mm.
Holotype female (No. 1869, Mus. Calif. Acad. Sci.), and
three paratype females in the collection of Dr. F. E. Blais-
dell, captured by Dr. Blaisdell at Poway, San Diego County,
California.
This robust species in both structure and appearance stands
midway between Dicheirus strenuus (Horn) and Dicheirus an-
gularis Casey. It is separated from the former by being smaller
126
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
and by having the propleurse somewhat punctured and from
the latter by being far more convex and robust, as well as by
having the hind angles of the prothorax more obtuse. From
dilatatus, it can always be separated not only by its robustness,
but by having non-rounded hind angles to prothorax and by
having longer pile. The elytra of blaisdelli simulate very closely
those of a female Plastocerus agassisi Lee. The genus Dichei-
rus Mann, is one of the few which might, with benefit, be kept
separate from the genus Anisodactylus Dej.
Habits- of Trachykele nimbosa Fall
While on Mt. Hood, Oregon, during the latter part of June,
I noticed that the rare buprestid, Trachykele nimbosa Fall,
could often be found ovipositing in the blazes made by the
surveyors in fir trees, Abies, along the right-of-way for a forest
road. These beetles are rather silent in flight, as is the case
with the AcmcBodera, and as a result they alight without attract-
ing much attention. Unfortunately, they also depart with as
great ease, simply vanish before you. This is particularly the
case during the heat of the day. If the weather becomes cooler
while they are in evidence, they may readily be approached and
even picked up with the fingers.
Trachykele opulenta Fall, a brilliant green species, which
breeds normally in the native cypress, Cupressus sar genii Jep-
son, on Cypress Ridge, Marin County, California, has been
found this last year to have come down into the valleys and
attacked the planted Monterey cypress, Cupressus macrocarpa
Hartw. As pointed out by Dr. H. E. Burke, this species only
attacks the dead limbs of living trees, the larvae gradually work-
ing into the main trunk of the tree which they in time honey-
comb. — E. C. Van Dyke.
Habits of Cymatodera decipiens Fall
During the summer of 1925, several Cymatodera dicipiens
Fall emerged from twigs of Pinus ponderosa collected during
the winter of 1923 on Mt. Wilson, Los Angeles County,
California. During the spring and summer of 1924 and 1925,
the only other insects to emerge from these twigs were an undes-
cribed species of Carphoborus which came out in some numbers.
It seems from this that the latter insect, in the larval stage, is
preyed upon by the Cymatodera larvae. — Alonzo C. Davis.
JANUARY, 1926] FALL ALASKAN COLEOPTERA
127
A LIST OF THE COLEOPTERA TAKEN IN ALASKA
AND ADJACENT PARTS OF THE YUKON
TERRITORY IN THE SUMMER OF 1924
BY H. C. FALL
Tyngshoro, Massachusetts
It was the writer’s privilege to be one of a party of three
to make a trip to this wonderland of the North during the
summer of 1924. While the excursion was primarily one of
sight-seeing and general recreation, the writer, from long habit,
kept always an eye open for possible coleopterous windfalls,
and as opportunity offered, a few minutes, a few hours, or
even the better portion of a day in places where a long enough
stop permitted, were devoted to special search for beetles. The
results in the aggregate seem worthy of record, in view of the
general interest taken in collections from this somewhat remote
region.
In 1857 Le Conte remarked that at the close of the activities
of the Russian entomologists, the fauna of the region explored
by them was more completely developed than that of any other
part of the continent. This, of course, is no longer true, even
if we confine ourselves to the coast region in which most of
the early collecting was done; while if the entire territory of
Alaska be considered, it would perhaps be nearer the truth to
say that its fauna is less completely known than that of any
other equal area of our country.
The total number of species of Coleoptera listed from Rus-
sian America at the time of Mannerheim’s last supplement in
1853 was 540. Hamilton’s Alaskan list with supplement in
1894 enumerates 578 species, and the few scattering additions
since that time can hardly bring the total much, if any, above
600 species. It is quite probable that there are now in collec-
tions, notably that of the National Museum, a considerable
number of additional species, but these, even if worked up,
have not yet been recorded.
On the present trip, as nearly as can now be determined,
448 species were taken within, or adjacent to, the Alaskan
boundary, of which 117, exclusive of unidentified Stenus and
Aleocharini, appear to be new to the Alaskan list. Of these,
61 occurred in the coast region and 64 in the Yukon Valley,
128
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
8 species being common to both regions. The fact that nearly
half of these 117 species are from the well-explored coast
region is rather surprising, since it was to be expected that
the great majority of future additions to the Alaskan fauna
would be from the interior valleys and mountains, which, eiito-
mologically speaking, are relatively unworked. The disparity
in this case would indeed have been much greater if the numer-
ous species taken at Dawson, Y. T., where considerable collect-
ing was done, had been included, for it is practically certain
that a very large proportion of the species occurring at this
point must also occur across the international boundary only
fifty miles distant, where the topographical and climatic condi-
tions along the Yukon are virtually the same.
In the interest of economy of space, I shall not attempt here
to speak of collecting conditions and experiences, or, except
for the table of dates below, to give the itinerary of the trip,
further than to say that we left Tyngsboro, Massachusetts, on
June 5, via Montreal and the Canadian Pacific, arriving at
Skagway June 18, after making the incomparable trip up the
“Inside Passage” from Vancouver. On August 2 we boarded
the steamer at Ketchikan for Prince Rupert, the return jour-
ney across the continent being made thence via the Canadian
National Railway to Montreal, arriving at Tyngsboro August 10.
The principal dates on which collecting was done are given
once for all in the subjoined table:
Wrangel, Alaska, June 17. Anchorage, Alaska, July 10-14.
Skagway, Alaska, June 18-20. Seward, Alaska, July 16-20.
Skagway, Alaska, July 27.
White Horse, Y. T., June
21 - 22 .
Dawson, Y. T., June 24-30.
Eagle, Alaska, July 1.
Fort Yukon, Alaska, July 2,
Nenana, Alaska, July 6.
Fairbanks, Alaska, July 7.
Mount McKinley Park, July
8-9.
San Juan, Evans Island,
July 20.
Childs Glacier, Alaska,
July 22.
Port Althorp, Alaska, July 23.
Juneau, Alaska, July 25-26.
Sitka, Alaska, July 28.
Scow Bay, Alaska, July 30.
Kasaan, Alaska, July 31.
Ketchikan, Alaska, August 1.
Throughout the 1400-mile trip down the Yukon and up the
Tanana Rivers daily stops were made for fuel at wood-piles
JANUARY, 1926] FALL ALASKAN COLEOPTERA
129
along the banks, at which nearly always some beetles were
taken.
Species new to the Alaskan list ^are marked with an as-
terisk (*).
List of Species
Family Cicindelid^
Cicindela longilabris Say. White Horse, Y. T. Several seen,
but having no suitable net no effort was made to secure
specimens. No tiger beetles whatever were observed in
Alaskan territory.
Family Carabid^
Trachypachus inermis Mots. Skagway; White Horse.
Cychrus (Pemphus) angusticollis Mann. Seward, Sitka.
Cychrus (Brennus) marginatus Fisch. Juneau; Skagway;
Kasaan ; Ketchikan. Common everywhere near the coast ;
individuals from the last two localities are considerably
larger than those from farther north.
Carabus tcedatus Fab. Skagway.
Elaphrus riparius L. Skagway ; McKinley Park ; Fort Yukon,
June 27 (Kusche).
Elaphrus pallipes Horn. Skagway; Dawson. All typical in
coloration.
Loricera 10-punctata Esch. Scow Bay. A single example.
*Opisthius richardsoni Kby. Skagway; Dawson; Fort Yukon,
June 27 (Kusche). Frequent along the river at Skagway,
varying greatly in color, coppery, green, or nearly black.
There are specimens of this species in my collection from
Dolomi, Alaska, collector unknown to me.
Notiophilus borealis Harris. McKinley Park.
otiophilus simulator Fall. Anchorage. A single example;
also taken by me at Jasper Park, Alberta, on the return
trip.
Nebria metallica Fisch, Skagway; Seward; Child’s Glacier;
Sitka; Juneau.
Nebria sahlbergi Fisch. Seward; Juneau. Two forms refer-
able to this species were taken on the trip ; one of narrow
thorax and somewhat more elongate form at Juneau and
Seward is probably the true sahlbergi. Of this a variety
with violaceous elytra (probably violaceus Mots.) and an-
130
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
other with pale legs occurred. The other form is repre-
sented by a single example taken at Coldstream, Vancouver
Island ; it is a little stouter with broader thorax, agreeing
closely with specimens in my collection from Washington
State and the Rocky Mountains of New Mexico. This
form, in its broader thorax, approaches moesta Lee., which,
I think, can hardly be a varietal form of sahlhergi, although
so recorded in the books.
Nebria bifaria Mann. A single typical specimen with bicolored
legs was taken at McKinley -Park.
Nebria gebleri Dej. Juneau.
Nebria viridis Horn. Dawson; Yukon River (Sheep Creek
below Eagle) ; Tanana River.
Nebria hudsonica Lee. White Horse. Two examples.
Nebria testaceipes Cys. Dawson. One example.
*Dyschirius secretus Fall, n. sp.
Slightly smaller than, but of the general form and aspect of,
integer Lee., near which it must be placed.
Body black, bronzed; legs and antennae black. Head four-fifths
as wide as the prothorax; front feebly transversely impressed; epis-
toma broadly and deeply emarginate. Prothorax subglobose, slightly
wider than long, sides broadly, evenly arcuate. Elytra oblong-oval,
three-fifths longer than wide; striae entire, rather deeply impressed
and strongly punctured before the middle, becoming much finer and
more finely punctured on the declivity; eighth stria finely impressed,
distinctly punctate throughout; base not margined; third interspace
tripunctate, a single apical setigerous puncture. Basal joint of hind
tarsi twice as long as wide and evidently shorter than the two
following combined. Length, 2.5 to 2.75 mm.; width, .8 to .85 mm.
Anchorage, Alaska, seven examples, all taken in damp moss
on low ground, July 13-14, 1924.
In integer, with which the present species is compared, the
size is slightly larger, the legs and antennae are never really
black, but rather dark rufous to rufopiceous, the basal joint of
the hind tarsus is more elongate, at least three times as long as
wide and fully as long as the two following; apex of elytra
with two setigerous punctures. There is a strong probability
that integer is distinct from the European ceneus with which it
has been united, judging from two examples of the latter sent
me by Mr. Edmund Reitter. In nigriceps, with which the
present species might also be compared, the size is larger, the
JANUARY, 1926] FALL ALASKAN COLEOPTERA
131
elytra more finely striate and punctate, the eighth stria obso-
lete, and the elytral apex apparently with two setigerous punc-
tures ; the basal joint of the hind tarsus is also more elongate,
as in integer.
Only two species of Dyschirius have hitherto been recorded
from Alaska, viz., the frigidus and transmarinus of Manner-
heim. The former was described from the Keiiai Peninsula,
and my Anchorage species might well be expected to be the
same. I find, however, that my specimens do not fit the descrip-
tion very well, the size of frigidus (3.5 mm.) being materially
larger, while the el 3 dra are said to be scarcely wider than the
thorax and the elytral striae evanescent shortly behind the
middle. If frigidus really equals ceneus, as it stands in our lists,
it certainly is different from the Anchorage species. Trans-
marinus was described from a single example from Sitka; it is
even larger than frigidus.
Asaphidion yukonense Wickh. White Horse; Dawson.
Bembidion simplex Haywd. McKinley Park.
Bembidion planiusculum Mann. Sitka.
Bembidion complanulum Mann. Dawson; McKinley Park;
Seward; Sitka; Kasaan.
Bembidion funercum Lee. Skagway; Dawson.
Bembidion mceklini Haywd. ? Sitka.
Bembidion quadrifoveolatum Mann. Kasaan.
Bembidion yukonum Fall, n. sp.
Form and size about as in grapi, but rather larger and a little
stouter. Black, distinctly green-bronzed, antennae at base and legs
rufous; surface above strongly shining throughout, without alutace-
ous sculpture except very feebly at elytral apex. Antennas scarcely
attaining the middle of the length, intermediate joints about two and
a half times as long as wide. Head a little wider than the thoracic
apex; eyes moderate. Prothorax subcordiform, one-fifth wider than
long, base slightly narrower than apex; sides strongly evenly rounded
from apex to basal constriction, which is parallel-sided for a distance
about equal to one-fifth the basal width; base angles sharply rec-
tangular, evidently but not very strongly carinate; basal impressions
deep, with a few rather coarse punctures. Elytra one-half wider and
three times as long as the thorax, one-half longer than wide; sides
broadly arcuate, humeri well marked, not angulate; striae feebly im-
pressed near the suture; strial punctures moderate, becoming finer
posteriorly, very fine but traceable near the apex; those of seventh
stria very fine in apical half; intervals flat, the third with two setiger-
132
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
ous punctures adjacent to the third stria at approximately the basal
and apical thirds. Length, 4.75 mm.; width, 1.9 mm.
Described from a single male specimen taken by the writer
at Dawson, Yukon Territory, June 30, 1924.
This species belongs to the ustulatum group, and is appar-
ently nearest insopitum Csy. The latter is unknown to me, but
according to description is a considerably larger species with
more transverse thorax, deeply striate elytra, the seventh stria
less distinctly punctured basally. Grapi is a rather smaller and
narrower species, with much finer elytral sculpture, the legs
typically black, though in two Greenland examples in the
Le Conte collection and in many North American examples so
referred, they are more or less distinctly rufous.
Bembidion bimaculatum Kby. Skagway; Dawson; Nenana.
There occurred at Nenana, together with the typical form,
what I take to be a variety of this species with smaller
thorax, browner elytra with or without apical spot, and finer
striae.
*Bembidion mckinleyi Fall, n. sp.
Black, above with feeble deep-green surface luster, beneath black,
femora piceous; trochanters, tibiae, and tarsi reddish-brown. Upper
surface polished throughout, with faint traces of alutaceous sculp-
ture only at elytral apex. Antennae black, basal joint rufescent, about
half as long as the body, slender, intermediate joints not quite three
times as long as wide. Head three-fourths as wide as thorax, eyes
not at all prominent. Prothorax nearly one-third wider than long,
sides rounded anteriorly, sinuate posteriorly, becoming nearly parallel
in basal fifth or sixth; hind angles rectangular, side margins very
finely reflexed; disc convex, median line well impressed and nearly
entire; foveae only moderately deep, flat or slightly tumid at bottom,
with a few obsolete punctures, hind angles not distinctly carinate.
Elytra fully three and one-half times as long and three-fourths wider
than thorax, three-fifths longer than wide, humeri rather wide, nar-
rowly rounded, sides diverging slightly and nearly straight in basal
two-fifths, greatest width at or very slightly behind the middle; striae
finely impressed, moderately strongly and closely punctured basally,
more feebly so posteriorly, nearly disappearing at apex; seventh stria
represented by a few fine punctures near the base. Length, 5.5 mm.;
width, 2.25 mm.
Described from a single pair taken in McKinley Park, Alaska,
July 8-12, 1924, by my fellow-traveler, Mr. C. A. Richmond.
This species belongs to the ustulatum group, and by its nar-
row thorax is somewhat remindful of scopulinum. It is, how-
JANUARY, 1926] FALL ^ALASKAN COLEOPTERA
133
ever, much larger and without maculation. Careful measure-
ment shows that the prothorax is not quite as wide at apex as
at base. On this basis it would, by Hayward’s table, be asso-
ciated with nevadense and consanguineum, to neither of which
is it at all similar. By Casey’s table it would fall near hum-
holdtense, which is even more remotely related. If the thorax
were regarded as narrower at base than at apex (this relation
is at best a rather elusive one) it would fall between scopulinum
and canadense in Hayward’s table, which is perhaps the most
satisfactory position for it.
Bembidion rupicola Kby. (lucidum Lee.). Skagway; Yukon
and Tanana Rivers. Rupicola has long stood on our lists
as a synonym of ustulatum, but wrongly so. The synonomy
here announced is made certain b)'^ an examination of one
of Kirby’s cotypes kindly sent me by Mr. Arrow.
Bembidion scopulinum Kby. Anchorage; Dawson.
Bembidion picipes Kby. {grapi Auct.). Skagway; Dawson;
Yukon River (Sheep Creek, below Eagle) ; Tanana River;
McKinley Park. This is the species we are accustomed
to call grapi, and probably correctly so, although I have
not been able to compare with European specimens. Except
for darker legs, a Sheep Creek specimen is almost an exact
counterpart of one of the cotypes of Kirby’s picipes, which
I have had the privilege of examining. There can be no
doubt that the other examples taken are the same thing,
and that the species long standing as picipes on our lists,
following Le Conte’s misinterpretation, must henceforth
be known as plagiatum Zimm.
Bembidion dentellum Thumb, (incrematum Lee., arcuatum
Lee.). Dawson.
"^Bembidion coloradense Haywd. Anchorage.
Bembidion nigripes Kby. Skagway; Dawson.
Bembidion intermedium Kby. I did not take this species, but
there are examples in my cabinet collected by Kusche at
Fort Yukon, June 27, 1916. This is the true intermedium
of Kirby, which seems never to have been correctly inter-
preted by our systematists. It is very closely allied to
nigripes (perhaps not distinct), and is similarly brightly
variegated with yellow. The much less ornate intermedium
134
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
of the Hayward monograph is an entirely different thing,
for which Le Conte’s name rapidum must be restored,
unless on investigation some still older name comes to
light.
Bembidion convexulum Haywd. Dawson. Two examples.
Bembidion atrolucens Csy. White Horse.
Bembidion versicolor Lee. Wrangel.
Bembidion dubitans Lee. Dawson.
Bembidion mutatum G. & H. Dawson.
^Bembidion sulcatum Lee. Anchorage. A single example.
^Bembidion concurrens Fall, n. sp.
Body oblong, suboval, rather strongly convex; black, polished
throughout, elytra with a small nubilous external pale spot at apical
fourth, the apex barely visibly pallescent. Antennae black, basal joint
rufescent; median joints twice as long as wide. Head only slightly
narrower than thorax; eyes rather large and prominent. Pro thorax
one-half wider than long, apex evidently wider than base, sides
broadly arcuate in front, posteriorly sinuate and subparallel in about
the basal sixth, the angles sharp and just visibly more than right;
basal fovese deep, not evidently punctured, bordered externally by a
fine acute carina. Elytra not quite one-half longer than wide, five-
eighths wider than thorax, parallel, sides feebly arcuate in basal half;
striae finely impressed internally, scarcely at all so in outer half;
strial punctures moderate, becoming much finer but still quite dis-
tinct on the declivity and laterally in apical half. Legs very nearly
black. Length, 3.25 mm.; width, 1.35 mm.
Alaska. Scow Bay (north of Wrangel) two males ; Skag-
way, one female. The type is a male from Scow Bay.
This species belongs to the as simile group and in form most
nearly resembles anguliferum, though materially smaller and
with elytra much less finely punctate-striate. The species of
this group form a difficult study, and are considerably more
numerous than indicated by Hayward’s very conservative treat-
ment. The present species by a rigorous interpretation of
Casey’s table would fall near invidiosum, in which, as in cautum,
the elytra are not truly black (as they are in the present spe-
cies) and the legs are rufous. The el3d;ra are also relatively
shorter and more coarsely striate than in concurrens. It is pos-
sible that invidiosum is not really distinct from cautum, and
I feel tolerably sure that digressum Csy. is the same as con-
nivens Lee.
Bembidion sp. dub. (assimile group). Dawson; Seward.
JANUARY, 1926] FALL ALASKAN COLEOPTERA
135
Bembidion oblongulum Mann. Seward.
Patrobus septenti'ionis Dej. Skagway.
Patrobus (Platidius) aterrimus Dej. Ketchikan.
Trechus chalybceus Dej. Skagway; Anchorage; Seward;
Kasaan.
Pterostichus crenicollis Lee. Ketchikan.
Pterosfichus amethystinus Dej. Kasaan.
Pterostichus validus Dej. Juneau; Kasaan; Ketchikan.
Pterostichus castaneus Dej. Scow Bay; Kasaan; Ketchikan.
Pterostichus adstrictus Esch. Skagway ; Juneau ; Anchorage ;
Seward; Scow Bay; Kasaan; Dawson. Nearly all the names
under Bothriopterus of the Leng list apply to more or less
trifling variations of this ubiquitous northern species.
Pterostichus (Cryobius) riparius Dej. Skagway; Seward;
Juneau.
Pterostichus ( Cryobius) hudsonicus Lee. Childs Glacier.
Pterostichus ( Cryobius ) brevicornis Kby. McKinley Park ;
Dawson.
Pterostichus ( Cryobius ) sp. near mandibularis , and perhaps not
different. McKinley Park; Tanana River.
Pterostichus ( Cryobius ) sp. McKinley Park ; Eagle ; Dawson.
I am unable to place this, even tentatively.
Pterostichus (Cryobius) sp. Anchorage. A single specimen
allied to the preceding, but with entirely black palpi, an-
tennae, and legs.
Amara ( Cyrtonotus ) eschscholtsi Chd. Dawson.
*Amara (Cyrtonotus) consueta Fall, n. sp.
Oblong, moderately elongate, form nearly as in eschscholtzi. Black,
head and thorax polished; elytra minutely alutaceous, moderately
shining in the male, slightly duller in the female; antennae and legs
ferruginous. Head not quite two-thirds as wide as prothorax. Pro-
thorax not quite one-half wider than long, base very slightly wider
than apex; sides broadly, evenly rounded anteriorly, briefly sinuate
before the nearly rectangular hind angles; disc moderately convex,
anterior transverse impressed line feeble; median line distinct, abbre-
viated at both ends; disc in front completely devoid of punctures,
basal impressions deep, a little oblique, numerously punctate; hind
angles obtusely carinate; base at middle impunctate, side margins
narrowly, finely punctured. Elytra ( 6 ) about one-fifth wider and not
quite three times as long as the thorax and about three-fifths longer
than wide; finely striate, striae finely but distinctly punctate, obso-
letely so near apex. Body beneath black, meso- and metepisterna and
136
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
sides of the metasternum and basal segments of abdomen with a
few fine punctures. Middle and hind thighs bisetose on inner margin;
tarsi with first three joints feebly grooved at sides. Length, 10.5 to
11 mm.; width 3.9 to 4 mm.
Described from six examples taken by the writer at Skagway,
Alaska, June 18-21, 1924.
This species is, by a strict interpretation of Hayward’s table,
to be associated with infausta Lec., and is undoubtedly closely
allied with both that species and eschscholtzi, from both of
which it seems constantly separable by the total lack of anterior
discal punctuation of the thorax. It seems highly probable that
we have too many described species already in this genus, never-
theless I am unable to attach any of the published names to the
present species.
Amara littoralis Esch. Skagway; Ketchikan.
Amara 7' emote striata Dej. White Horse.
Amara mterstitialis Dej. Dawson.
* Amara polita Lec. Eagle.
Calathus ingratus Dej. Skagway; Nenana; Anchorage.
Calathus advena Lec. Childs Glacier.
Platynus stygicus Lec. {\\maurus Mots.). Anchorage.
Platynus perforatus Lec. Dawson.
Platynus bogemanni Gyll. Skagway ; Dawson.
Platynus quadripunctatus DeG. Anchorage.
Platynus n. sp. ? near sordens Kby. Anchorage.
^'Platynus nigriceps Lec. Fairbanks.
Cymindis cribricollis Dej. White Horse.
Harpalus fulvilabris Mann. Eagle.
Harpalus plew'iticus Kby. Dawson.
^Harpalus laticeps Lec. ? Fairbanks. A single non-typical
example, somewhat crushed, picked up from the sidewalk.
"^Harpalus cordifer Notm. Juneau; taken also at Goldstream,
Vancouver Island.
Harpalus basillai'is Kby. White Horse ; Eagle.
*Harpalus pacificus Fall, n. sp.
Body oblong, rather markedly convex, black, strongly shining
throughout, the elytra very finely alutaceous. Antennae and trophi
ferruginous, the former scarcely passing the hind angles of the
thorax, their intermediate joints fully twice as long as wide; head
not quite three-fourths as wide as thorax; eyes moderately promi-
nent. Prothorax slightly more than one-half wider than long; base
JANUARY, 1926] FALL ALASKAN COLEOPTERA
137
a little wider than apex; sides subevenly arcuate, a little more
strongly so in front; basal angles obtuse, their apices somewhat
blunted but not distinctly rounded; basal foveae rather strongly,
closely punctured, the adjoining surface externally sparsely punc-
tured, the punctures nearly absent at middle of base; sides strongly
convexly declivous anteriorly, narrowly and feebly deplanate pos-
teriorly from the middle, the marginal bead translucent posteriorly.
Elytra not quite one-half longer than wide, sides parallel; striae fine,
intervals flat with scattered, very minute punctures. Body beneath
black, the epipleura not at all paler. Legs rather stout; femora and
tips of tibiae black; tibiae except at apex and tarsi rufous. Length,
8.5 mm.; width, 3.5 mm.
Ketchikan, Alaska, a single male (type) ; Scow Bay, one
female ; Skagway, one male doubtfully referred.
This species in its typical form agrees nearly with the de-
scription of fugitans Csy. in many respects, including the color
of the legs, but in that species the form is said to be unusually
depressed, the hind angles of the thorax nearly right, and the
pronotal foveae very shallow and sparsely punctured, none of
which statements are at all applicable to the present species.
The Scow Bay and Skagway specimens differ from the type
in having the legs predominently rufous, the tips of the femora
and tibiae only being lightly infuscated, the sides of the thorax
are also less rounded. In the Scow Bay female the elytra are
dull. All three specimens have the mentum emargination vir-
tually without a tooth, although there is an extremely feeble,
broadly rounded, prominence detectable on close scrutiny.
Harpalus rufimanus Lee. Dawson.
Trichocellus cognatus Gyll. Seward.
Tachycellus nigriniis Dej. Skagway; Juneau; Scow Ba,y ;
Wrangel.
FAMILY DYTISCID.F;
Coelambus borealis Fall, n. sp.
Elongate oval, widest at middle of the length, sides of thorax
nearly continuous with those of the elytra. Head and thorax flavo-
testaceous, the former with a dark spot adjacent to each eye and
the hind margin narrowly blackish, the latter with the anterior mar-
gin narrowly dark, and a small central spot which is diffusely pro-
duced to the hind margin. Elytra with the disc broadly fuscoluteous,
due to the diffusion of the customary markings, which are, however,
somewhat defined when the elytra are viewed very obliquely in the
direction of their length. Body beneath black except the head,
thorax, and epipleura; legs rufotestaceous; tarsi blackish. Head
138
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
alutaceous and sparsely punctulate. Prothorax and elytra polished
throughout, the former finely and sparsely punctured on the disc,
more closely toward all the margins, more strongly so post-medially;
elytral punctures nearly equal in the posterior half where they are
separated by about half their own diameters, in the anterior half
they are obviously, though not very conspicuously, unequal and
somewhat less closely placed. Prothorax one-half wider than the
head, the sides moderately divergent and almost straight throughout.
Elytra twice as wide as the head and one-half longer than wide.
Sides of coxal plates and of the abdomen basally, coarsely closely
punctate, the punctures separated by much less than their own
diameters; surface of coxal plates strongly alutaceous.
Male. Four anterior tarsi strongly dilated, the second joint of the
protarsus visibly wider than the first and as wide as the tibiae at
apex; anterior protarsal claw a little thicker and just visibly shorter
than the posterior and more strongly curved basally. Length, 3.7
mm.; width, 1.85 mm.
Dawson, Yukon Territory. A single male taken by the writer
June 30, 1924.
This species is very closely allied to patruelis, and is possibly
only a variety thereof. It differs from all specimens of a
numerous series of patruelis in my collection by the almost per-
fectly straight sides of the prothorax and by the strongly aluta-
ceous coxal plates, these being polished and without alutaceous
sculpture, except in the dull females, in patruelis. The punctua-
tion of the coxal plates in patruelis is also less dense than in
the present species, the elytral disc is more uniformly punctured,
and the tarsi only rarely show any tendency to infuscation.
Hydroporus planius cuius Fall. Scow Bay.
Hydroporus notahilis Lee. Anchorage. Specimens vary con-
siderably in size and indicate the identity of notabilis and
arcticus.
Hydroporus nigellus Mann. Anchorage ; Seward ; Skagway.
Hydroporus tartaricus Lee. McKinley Park.
Hydroporus melanocephalus Gyll. Dawson; Tanana River;
Fairbanks.
Hydroporus polaris Fall. Dawson. A single male of this rare
species.
"^Hydroporus fuscipennis Schaum, Fairbanks.
Hydroporus striola Gyll. Dawson.
Hydroporus yukonensis Fall, n. sp.
Allied to 12-line atus and semiclarus. Moderately elongate, oval;
upper surface pale testaceous with blackish markings as follows:
JANUARY, 1926] FALL ALASKAN COLEOPTERA
139
Head with a subtriangular spot each side adjacent to the eyes, these
spots confluent with each other and with a posterior marginal line
at the middle of the occiput; prothorax with a small transverse spot
each side of the middle in the posterior transverse impression and
another still smaller at the middle of the sublateral longitudinal im-
pression; elytra each with six narrow discal vitte and two or three
exterior longitudinal dashes, the black vittae a little wider than the
pale stripes between them. Prothorax wider at base, twice as wide
as long, and about one-half wider than the head; sides oblique and
nearly straight from the base almost to the apical angles; side mar-
gins fine but distinct. Elytra about four and one-half times as long,
and one-half wider than the thorax, and a little less than one-half
longer than wide; sides markedly discontinuous with those of the
thorax, subparallel and broadly arcuate in basal three-fifths; feebly
sinuate before the apex in the male, obtusely angulate before an
apical sinus in the female; the sutural angles acute and well defined in
both sexes. Upper surface throughout micro-reticulate and sparsely,
finely punctulate. Body beneath black, similarly micro-reticulate and
sparsely punctulate; margins of hypomera and epipleura testaceous.
Antennae and legs testaceous, the apical parts of the terminal joints
of the palpi and outer joints of the antennae and the basal two-thirds
of the femora infuscate.
In the male the protarsi are a little wider than in the female, with
the second joint evidently wider than the first (scarcely wider than
the first in the female), and the protarsal claws slightly less slender,
though mutually equal and scarcely longer than in the female. The
last ventral segment is not appreciably modified in either sex.
Length (type male), 4.5 mm.; width, 2.3 mm.
Described from a single pair taken by the writer at Dawson,
Yukon Territory, June 24, 1924.
This species is very nearly allied to semiclarus, and may prove
to be only a smaller Northern race of that species. The two
agree in having the male protarsal claws slightly thickened,
but the present species, in addition to the smaller size, differs
in its somewhat smaller thorax with nearly straight converg-
ing sides, the sides being much more rounded in semiclarus ;
12-lineatus differs from both by the protarsal claws of the male
not at all thickened, the entirely pale femora and the more
ample prothorax with sides less strongly discontinuous with
those of the elytra.
'^'Hydroporus alaskanus Fall, n. sp.
Similar to the preceding species in color, markings, general form
and structural features, except as follows: Form a little more elon-
gate, more than twice as long as wide (less than twice as long as
wide in yukonensis). Head three-fourths as wide as the prothorax,
the latter with sides straight and parallel in basal half to three-
140
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
fourths, extremely finely margined; hind angles rectangular; disc
with two anterior transverse black dashes parallel with those in the
posterior depression; elytra with sides even more strongly discon-
tinuous with those of the thorax. In the male the protarsi are as in
yukonensis, except that the tarsal claws are virtually as slender as in
the female; the last ventral is broadly transversely impressed medi-
ally, the apical half flattened or feebly impressed longitudinally at
middle. In the female the apex of the last ventral is produced in
the form of a short broad deflexed process with converging sides
and broadly truncate extremity. Length, 4.5 to 5.3 mm.; width, 1.9
to 2.4 mm. Ratios of length to width in four measured examples are
2.37 (6); 2.2, 2.2, 2.1 ($).
Described from one male and seven females, all taken by
the writer at Skagway, Alaska, June 18-21, 1924. It is quite
distinct from the preceding species by the above characters, and
from all allied forms by the peculiar apical process of the last
ventral in the female.
*Hydroporus recticollis Fall, n. sp.
Closely allied to the preceding two species, but more especially
to alaskanus with which it agrees in the short parallel-sided thorax
with rectangular hind angles, the nearly straight sides extremely
finely margined. The form is more strongly elongate, the thorax
smaller than in either of the preceding two species and not much
wider than the head; elytra from five and one-half to six times the
thoracic length. Black markings of the upper surface heavier than
in the preceding species, the thoracic disc becoming diffusely clouded,
leaving the margins and a narrow central line pale; the black elytral
vittae much wider than the intervening yellow lines and the external
fragmentary vittse more developed. There is a tendency here, as in
all species of the group, for the first and second elytral vittae to
become confluent in a small antemedian spot. The epipleura are
entirely blackish. In the male, as in the preceding species, the pro-
tarsi are perceptibly more dilated than in the female, the second
joint a little wider than either the first or third; the claws mutually
equal, slender and not differing sexually. The last ventral is longi-
tudinally medially impressed in apical half in the male. In the female
the last ventral is rather abruptly flattened or impressed around the
apical margin, which is at middle a little deflexed, but not distinctly
produced. The subapical marginal angulation of the elytra in the
female is a little more prominent, being nearly rectangular in the
present species, obviously obtuse in the two preceding. Length (male
type), 5.2 mm.; width, 2.3 mm. In two females the dimensions are
4.9 and 2.15 and 4.8 by 2.15 mm.
Described from one male and two female examples, taken by
the writer at Seward (Kenai Peninsula), Alaska, July 18, 1924.
JANUARY, 1926] FALL ALASKAN COLEOPTERA
141
The five species now known from our fauna, in this restricted
group having the sides of the elytra of the female subapically
angulate, may be tabulated as follows :
Prothorax wider at base than at middle.
Protarsal claws not differing appreciably in the sexes, femora
entirely pale 12-lineatus
Protarsal claws slightly thickened in the male; femora blackish in
basal half to three-fourths.
Sides of thorax arcuate throughout semiclarus
Sides of thorax nearly straight yukonensis
Prothorax not appreciably wider at base than at middle, the sides
virtually straight and parallel in at least basal half; protarsal
claws of male not at all thickened.
Last ventral of female with a broad deflexed apical
process alaskanus
Last ventral of female without broad apical process; general form
a little more elongate; prothorax smaller and with broad diffuse
discal cloud recticollis
Agabus hypomels Mann. Ketchikan; Scow Bay; San Juan
(Evans Island).
^Agabus inscriptus Cr, Skagway. I have also in my cabinet a
specimen taken at Homer by W. S. Me Alpine of Detroit.
"^Agabus sp. Allied to canadensis ; a single male taken at Fair-
banks. It is possible that the lecontei Cr. of the Hamilton
list may be this.
Agabus infuscatus Aube. Anchorage.
Agabus anthracinus Mann. San Juan (Evans Island).
Agabus nigroceneus Er. Fairbanks.
*Agabus kenaiensis Fall, n. sp.
Obtusely oblong oval, black, antennae and legs brownish, the
thighs darker, especially the posterior. Body only moderately con-
vex, surface rather strongly shining, minutely alutaceous and with a
feebly impressed system of much larger reticulations which are quite
irregular in form. Punctuation apparently lacking, except for the
usual irregular finely setigerous elytral series and similar irregular
series along the front margin and the exterior third of the rear
margin of the prothorax. Closer inspection, however, reveals numer-
ous scattered very minute punctures mostly at the intersections of
the fine reticulating lines. Head with two faint rufous occipital spots.
Prothorax a little less than three times as wide as long, sides feebly
arcuate, nearly continuous with the sides of the elytra. Margin mod-
erate, base just perceptibly sinuate each side of the middle. Elytra
two-thirds as long as wide, slightly wider at middle than at base,
sides subparallel in basal two-thirds. Body beneath alutaceous-
reticulate, with a few fine punctures; ventral surface with some
142
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
widely spaced oblique very fine striae, especially on the second seg-
ment. Prosternal process roof-shaped; hind coxal plates distant by
rather less than half their own length from the middle coxal cavi-
ties; hind tibiae with a few punctures along their inner margin.
Male. Anterior tarsi feebly dilated, somewhat compressed, clothed
beneath with rather long squamules bearing at their tips a few small
inconspicuous palettes; prosternal claws a little elongate, slender,
equal, the posterior with a more pronounced basal sinuation of the
inner margin. Length, 6.25 mm.; width, 3.3 mm.
Described from two completely similar male specimens taken
by the writer at Anchorage, Alaska, July 14, 1924. They
occurred in wet moss.
This species is nearly allied to confertus, differing in its much
smaller size, somewhat narrower form, elytra without a trace
of a sublateral pale spot, last ventral segment of male nearly
destitute of the longitudinal strigae so conspicuous in confertus.
In the present species the posterior protarsal claw shows the
more conspicuous basal sinuation, while in confertus it is the
anterior claw.
*Agabus gelidus Fall, n. sp.
Moderately elongate, oblong oval, barely perceptibly inflated pos-
teriorly, elytra and thorax nearly continuous in outline; black, mod-
erately shining; antennas dark rufous; legs piceo-rufous. Very simi-
lar in most of its characters to confertus, with which it agrees nearly
in size and form; the sculpture above consists as in confertus, of a
system of rather large irregular reticulations within which the sur-
face is more finely reticulate, with sparse, very fine scattered punc-
tures, which may lie either within the meshes or on the reticulating
lines. As in confertus, the metastenial groove for the reception of
the tip of the prosternal process is quite rudimentary. The series
of punctures along the inner margin of the hind tibiae are very few
in number, and the present species differs otherwise from confertus
in the narrower side margins of the prothorax, and notably in the
much narrower wings of the metasternum between the middle coxal
cavities and the hind coxal plates, this distance being in the present
species less than half the distance across the coxal plate, while in
confertus it is obviously greater than half the width of the coxal
plate. The sublateral yellow spot of the elytra is wanting in gelidus,
and the longitudinal strigation of the last ventral in the male is only
feebly developed, there being, however, a broad median impressed
area extending from apex nearly to base of the segment, which seems
to be wholly wanting in confertus. The protarsi of the male are
similarly but noticeably more strongly dilated and compressed in
gelidus, the protarsal claws elongate and sinuate on the inner margin
in about the same degree. Length, 8 mm.; width, 4.25 mm.
JANUARY, 1926] FALL ALASKAN COLEOPTERA
143
Described from a single male specimen taken by the writer
at Fairbanks, Alaska, July 7, 1924.
Agabus tristis Aube. Skagway; Sitka; Wrangel; Ketchikan.
Ilybius quadrimaculatus Lee. Wrangel.
Rhantus divisus Aube. Skagway.
FAMILY HYDROPHILID^
Ochthebius holmbergi Makl. Anchorage.
Helophorus inquinatus Mann. Skagway.
"^Helophorus lacustris Lec. ?, McKinley Park.
"^Helophorus nitidulus Lec. Fairbanks.
Hydrobius fuscipes L. Fairbanks.
^Hydrobius scabrosus Horn. Ketchikan.
"^Paracymus moratus Horn, var. Skagway; Juneau. Mr. Lie-
beck, who has kindly compared specimens with the Utah
types in the Horn collection, finds certain small differences
which seem to be constant and indicate that the Alaskan
form may be distinct.
Laccobius ellipticus Lec. Anchorage.
Cercyon marinus Thoms. Dawson.
Cercyon fulvipennis Mann. Skagway; Anchorage; Juneau.
FAMILY SILPHID^
Necrophorus pustulatus var. melsheimeri Kby. Anchorage.
Silpha lapponica Hbst. Anchorage.
Silpha tritub erculata Kby. Dawson ; White Horse.
Pelatines (P elates Horn ||) latus Mann. Sitka.
Choleva spcnciana Kby.? Dawson. Two females. C. alsiosa
Horn is in my collection from Fort Yukon (Kusche).
Although described from Alaska in 1885, this species does
not appear in the Hamilton list of 1894.
Colon magnicolle Mann. McKinley Park. A single specimen.
Colon pusillum Horn. Dawson. Two examples.
Hydnobius substriatus Lec. Skagway; Dawson.
*Hydnobius latidcns Lec. McKinley Park.
Anisotoma punctatostriata Kby. Dawson.
^Anisotoma assimilis Lec. Anchorage.
*Agathidmm revolvens Lec. McKinley Park; White Horse.
"^Clambus gibbulus Lec. Anchorage.
FAMILY SCYDM^NID.®
Lophoderus bifoj'mis Makl. Sitka; Ketchikan.
144
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
*Euconnus longiceps Fall, n. sp.
Of large size (2 mm.) and rather narrow form, polished, impunc-
tate, except for the minute punctures bearing the elytral hairs; brown,
with the sutural edge nigrescent basally. Pubescence yellowish-
brown, stiff and posteriorly directed on the occiput, similarly rather
densely bristling at sides of prothorax; longer, finer, sparser and
more reclinate on the elytra. Head much longer than wide, scarcely
more than half as wide as the prothorax; sides of the occiput nearly
straight and convergent for about four times the length of the small
anterior eyes, to the strongly constricted neck, the latter about one-
half the width of the head. Antennae fully half the length of the
body, without definitely limited club; second joint subcylindrical,
slightly wider apically, twice as long as wide and nearly equal to
the two following; three to six equal in width to the second and
mutually nearly similar, suboblong, slightly longer than wide; seven
to ten gradually wider and becoming a little transverse; eleven not
quite as long as the two preceding. Prothorax distinctly longer than
wide, sides parallel and arcuate before the posterior lateral constric-
tion, subbasal transverse impression feebly trifoveate at middle and
at hind angles, the latter with short carina. Elytra elongate oval,
widest at middle, about two-thirds longer than wide and two-thirds
wider than the prothorax; basal impression strong, subhumeral and
sutural impressions obsolete. Femora moderately clavate; tibiae wider
medially, tarsi rather short and stout, basal joint subequal to the
two following; apical a little longer than the basal; hind coxae sub-
contiguous. Length, 1.95 mm.; width, .75 mm.
Ketchikan, Alaska, August 1, 1924, a single example sifted
from damp moss.
Although possessing the essential characters of Euconnus, this
species does not fit very well in any of the groups of the genus
established by Casey. The very elongate head should make it
easily recognizable among all known species of the genus. The
sex of the single individual is uncertain, but the slightly broader
basal joint of the protarsus may indicate the male.
*Scydmaenus adjutor Fall, n. sp.
Elongate, suboval, only moderately ventricose; black, antennae and
legs brownish ferruginous; surface polished, head and prothorax
impunctate, elytra with fine sparse punctulation; pubescence fine,
sparse and reclinate or inclined. Head wider than long; eyes rather
prominent, subbasal, the tempora very short and rapidly convergent;
excavation of the vertex broad and shallow, vaguely biimpressed at
bottom. Antennae gradually incrassate, the eighth joint slightly, the
ninth and tenth more distinctly, transverse; joints proportionately
nearly as in pacificus. Prothorax a little longer than wide, sides mod-
erately rounded in apical half, constricted at basal third; surface
coarsely biimpressed laterally within the constriction and with a
JANUARY, 1926] FALL ALASKAN COLEOPTERA
145
short subbasal dorsal transverse impression. Elytra oval, widest near
middle, three-fourths wider than thorax, one-half longer than wide;
humeri obtusely rounded but evident; basal impressions nearly as in
pacificus’, legs as in pacificus. Length (head depressed), 1.2 to 1.3
mm.; width, .55 mm.
Described from two examples taken by the writer at Skag-
way, Alaska, July 27, 1924.
This species is a Brachycepsis, closely allied to suhpunctatus
and pacificus. It is very considerably smaller than either of the
two mentioned, and differs from pacificus also in its black color
(rufotestaceous in pacificus), rather more slender antennas and
slightly less ventricose elytra, which are widest more nearly at
the middle. There is no apparent indication of sex in either of
the two specimens.
Eiifheia (Veraphis) scitula Makl. Scow Bay.
FAMILY ORTHOPERID^
^Orthoperus scutellaris Lee. Dawson; Sheep Creek; Yukon
River (below Eagle). This species was wrongly trans-
ferred to Sphaerius in the Leng List.
*Sacium lugubre Lee. White Horse; Dawson; Fort Yukon.
FAMILY STAPHYLINID^
Olisthcerus megacephalus Zett. McKinley Park ; Tanana River.
Megarthrus sinuatocollis Lac. Dawson.
Anthobium pathos Mann. Seward.
"^Pycnoglypta lurida Gyll. Anchorage.
Omalium foraminosum Makl. Dawson; Sheep Creek, Yukon
River (below Eagle).
Omalium sp. Fairbanks.
Phlceonomus lapponicus Zett. Dawson; Anchorage; Fort
Yukon (Kusche).
*Arpedium brunnescens Sahlbg. Skagway ; McKinley Park.
Acidota quadrata Zett. Skagway; McKinley Park,
Micrcedus laticollis Mann. Sitka.
Micrcedus austinianus Lee. Seward; Ketchikan.
P elecomalium testaceum Mann. Skagway ; Seward.
Geodromicus ovipennis Lee. Dawson. One example. This
specimen agrees well with LeConte’s type and cannot safely
be referred to the European plagiatum (see Leng list), a
specimen of which is before me. This latter has the elytra
distinctly longer, with less diverging sides.
Ancycophorus biimpressus Makl. McKinley Park.
146
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
*Trogophloeus alaskanus Fall, n. sp.
Moderately slender, depressed, sublinear; black, legs black, the
tibiae internally, tarsi, mouth and base of palpi testaceous; antennae
piceous. Upper surface densely, finely punctate and feebly shining;
abdomen, especially toward apex, more sparsely punctate and shin-
ing; pubescence very short and fine. Head quadrate, hind angles
subrectangularly rounded; tempora slightly longer than the eyes,
straight and parallel; vertex broadly roundly impressed Eyes small,
just perceptibly more prominent than the tempora, attaining the
middle of the elytra. Antennas moderately slender, gradually in-
crassate, the outer three joints forming a feebly differentiated club;
joints four to eight about as long as wide, fourth smallest,
eighth just perceptibly transverse. Prothorax equal in width to the
head, a little wider than long, front angles nearly right and scarcely
rounded, sides broadly arcuate and feebly convergent from just be-
hind the apex; hind angles obtuse and rounded as viewed from
above; disc with an oblong impression occupying the apical two-
thirds of the length and the middle one-third of the width. Elytra
distinctly wider than thorax, subequal in length to the head and
thorax combined and much longer than wide. Abdomen slightly
narrower at base than the elytra and slightly widened behind, border
rather thick but shallow; ventral segments four to six impressed at
middle, the fourth very feebly, the sixth broadly and deeply, the apex
of the latter behind the impression briefly carinate, the margin
broadly angulate medially. Legs moderately slender. Length, 1.6
mm.; width, .45 mm.
McKinley Park Station, Alaska, July 9, 1924; a single ex-
ample taken by the writer.
This little species belongs to Group IV of Casey’s Revision,
and by the table would be associated with indigens, from which
it differs in a number of details, notably the longer elytra, dark
legs, and ventral impressions.
*Trogophloeus teres Fall, n. sp.
Closely related to the preceding species, but apparently distinct
by the following differences: Size larger (2.3 mm.), elytra piceo-
testaceous, legs entirely pale; prothorax slightly wider than the head
and relatively more transverse, the impression much larger, extend-
ing almost throughout the length, at the base involving almost the
entire discal width, but becoming narrower and more vague anteri-
orly. Elytra even slightly longer than in alaskanus, abdomen more
distinctly narrowed basally and more strongly punctured, less shin-
ing. Ventral segments not impressed.
A single specimen only, taken at McKinley Park Station on
the same date as the preceding, but whether the two specimens
were taken together I do not recall. There is perhaps a possi-
JANUARY, 1926] FALL ALASKAN COLEOPTERA
147
bility that they are sexes of one species, the difference in color
being due to immaturity. This point can only be settled by
further collecting. So far as I know these are the first species
of the genus to be recorded from Alaska.
Aploderus sp. Female. Ketchikan. This seems much like
princeps, but without the male definite determination is
difficult.
Oxytelus invenustus Csy. ? One female. White Horse. A
male example taken at Edmonton, Alberta, on this trip, and
others in my collection from Banff, Alberta, and Beulah,
New Mexico, appear to possess precisely the male sexual
characters described by Casey from his Maryland type.
^'Oxytelus suspectus Csy. Skagway.
Platystethiis americamis Er. Dawson.
Bledius turgidiis Csy. Yukon River, fifty miles below Selkirk,
Y. T.
Bledius sp. Dawson.
Bledius alhonotatus Makl. ? Skagway.
Stenus bipunctatus Er. Dawson.
^Stenus austini Csy. McKinley Park; Anchorage. I have
also a specimen of this species taken by Wickham at Fort
Wrangel; this is, I suppose, one of the two undetermined
species mentioned by Wickham in his list.
Stenus ptej'obrachys G. and H. Juneau; Sitka; Ketchikan.
* Stenus stygicus Say. Eagle.
^Stenus mammops Csy. Anchorage.
Stenus cariniceps Makl. Juneau; Ketchikan. Ten other as
yet unidentified species of Stenus were taken in Alaska.
I had hoped to be able before this to compare these with
Casey’s types, but Mr. Barber writes me that the latter are
not yet available for study.
*Lathrobium sollicitum Fall, n. sp.
Form nearly parallel; black, antennas piceous, base scarcely paler;
legs rufopiceous, the tibiae paler. Head, thorax and elytra polished,
abdomen dull from the density of the minute punctulation. Antennae
rather slender, not quite reaching the thoracic base, intermediate
joints all feebly obconic and a little longer than wide. Head behind
the clypeus subquadrate, sides parallel, hind angles broadly rounded,
base truncate at middle, punctures numerous but well separated,
median line smoother; eyes at about twice their length from the
148
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
base; gular sutures parallel and widely separated; neck one-third the
width of the head. Prothorax slightly wider than the head, about
one-fourth longer than wide, sides parallel, feebly arcuate, front
angles rather abruptly, hind angles more broadly, rounded; punctua-
tion strong, not very close, median line narrowly but indefinitely
smooth. Elytra equal in length and in maximum width to the
thorax, sides very feebly but perceptibly divergent from base to apex,
punctures similar to those of the thorax but rather sparser. Abdo-
men at base equal in width to the elytra, very slightly widened to
apical third, very finely, densely punctulate and dull, both above and
beneath. Legs moderately stout, the front thighs thickened but not
sinuate beneath. Length, 5.2 to 5.4 mm.; maximum width, .9 mm.
Described from two males taken by the writer at Anchorage,
Alaska, July 13-14, 1924.
In these the sixth dorsal abdominal segment is strongly acumi-
nate at apex, the sixth ventral with a very small apical emargi-
nation, triangular in form with its apex narrowly rounded.
By its narrow neck, this species should be referred to the
group Tartopeus, but does not resemble at all any species
therein defined by Casey, unless it be his lacustris, which is
apparently quite different from the other species of his table,
and is seemingly remindful of the more typical Lathrobii. In
lacustris the elytra are said to be shorter than the thorax, and
the sexual characters of the abdomen are not like those in
sollicitum.
Lathrobium finitimum Lee. Scow Bay.
Lathrobium subseriatum Lee. Kasaan.
^Lathrobium sewardi Fall, n. sp.
Form moderately slender and convex. Head, thorax, and basal
half of elytra brownish piceous or castaneous, apical half of elytra
brighter rufous; abdomen piceous, antennae and legs reddish-brown.
Head subquadrate, sides straight and parallel, hind angles broadly
rounded, base truncate; eyes small, at twice their length from the
base; surface densely punctate at sides and base, less closely so
medially. Antennae attaining base of prothorax, median joints ob-
conical and about one-half longer than wide. Prothorax slightly
narrower than the head, one-third longer than wide, widest at apex,
sides nearly straight and convergent to base; all angles broadly
rounded; punctuation close and confused, median impunctate line
subelevated. Elytra as wide as the head, subequal in length to the
prothorax, and one-sixth or one-seventh longer than wide; sides
parallel, surface coarsely and closely sublinearly punctate. Abdomen
very finely not closely punctate.
Male. Ventral segments two to six with median impressions in-
JANUARY, 1926] FALL ALASKAN COLEOPTERA
149
creasing in size and depth, margin of fifth segment broadly sinuate,
apex of sixth with suboval or elliptical emargination, its limiting
angles sharply subrectangular, the impression anteriorly with a dense
patch of short black spicules, narrowly divided along the middle.
Length, 5.8 to 7 mm.; width, .9 to 1 mm.
Described from two males and four females taken by the
writer at Seward, Alaska, July 18-20, 1924.
This species belongs to Casey’s genus Lobrathium and is very
closely related to tacoma; Csy., differing most notably in color
and the less slender antennae; the secondary sexual characters
are nearly identical.
Baptolinus macrocephalus Nordm. Ketchikan ; Seward ; Kasaan ;
Skagway.
Philonthus septentrionis Fall, n. sp.
Deep black throughout, polished, elytra with slightly green bronze
luster. Head quadrate oval in male, a little more oval in the female;
eyes distant their own length from the base of the head. Antennae
barely as long as the head and thorax, entirely black; joints eight to
ten as wide or slightly wider than long, the preceding joints slightly
more elongate. Prothorax quadrate oval, one-third to two-fifths
wider than the head, evidently narrowed in front; dorsal punctures
fine, three in number, equidistant. Elytra a little wider than the
thorax, nearly square or very slightly transverse; punctuation rather
fine and sparse, not distinctly muricate. Abdomen sparsely a little
more strongly punctate than the elytra, beneath still more coarsely so.
Male. Anterior tarsi moderately strongly dilated; last ventral with
a triangular emargination a little wider than deep, and surrounded
by an acutely angulate gutter. Length, 6 to 7 mm.; width, 1.4 to
1.6 mm.
Described from one male and two female examples collected
by the writer at Dawson, Yukon Territory, June 24, 1924.
This species is of about the size and aspect of umhratilis, and
may be placed near it. In the latter the tempora are shorter,
the thorax less narrowed in front and with coarser serial punc-
tures, the abdomen more finely punctured beneath, and the legs
brownish-red.
Philonthus hasalis Horn, var. Dawson. Three examples differ-
ing from the typical form by having the elytra varied with
fuscous.
Philonthus fraternus Fall, n. sp.
Moderately elongate, deep black, shining, tarsi alone brownish.
150
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
Antennae not quite as long as the head and thorax, outer joints
nearly square. Head subquadrate, three-fifths as wide as the thorax,
the latter as long as wide, distinctly narrowed in front; dorsal punc-
tures four, equidistant. Elytra nearly square, a little wider than the
thorax, not at all aeneous, punctuation rather sparse. Abdomen a
little more finely and equally sparsely punctate.
Male. Front tarsi very broadly dilated, last ventral triangularly
emarginate and with a rather deep longitudinal impression tapering
forward and extending the entire length of the segment; penulti-
mate segment very broadly angulato-sinuate. Length, 6 mm.; width,
1.4 mm.
Dawson, Yukon Territory, June 24, 1924, a single male
specimen taken by the writer.
This species can only be compared with quadricollis and the-
veneti, because of the last ventral of the male being longitudi-
nally impressed throughout its length. In both these latter spe-
cies the sides of the thorax are parallel or very nearly so, while
in the present one they are distinctly convergent. Qiiadricollis
has the legs and antennae brown or paler and theveneti is said
to have the elytra not wider than the thorax.
^Philonthus sp. Ketchikan. One female in the vicinity of
crotchi, which it may be.
^'Philonthus distans Horn. Anchorage.
Philonthus siegwaldi Mann. S kag way ; Juneau ; Port Althorp.
Philonthus nigrihdus Grav. Skagway ; Dawson.
^Philonthus decipiens Horn. Wrangel.
"^Philonthus sordidus Grav. Anchorage.
Creophilus maxillosvis L. Juneau.
Heterothrops sp. Near carhonatus, but probably distinct;
Dawson, one female.
Quedius Icevigatus Gyll. Seward.
Quedius molochinus Grav. Sev/ard; Skagway; Juneau.
Quedius brunneipennis Mann. Anchorage ; McKinley Park.
Quedius hyperhoreus Er. ? Anchorage.
Quedius sublimbatus Makl. Skagway.
Quedius sp. Childs Glacier.
Quedius peregrinus Grav. Juneau.
Quedius sp. near cenescens Makl. Fairbanks.
Tachinus nigricornis Mann. Wrangel.
Tachinus instabilis Makl. Anchorage; Juneau.
JANUARY, 1926] FALL ALASKAN COLEOPTERA
151
Tachinus circtmicinctus Makl, Dawson; Tanana River; Fort
Yukon (Kusche).
^Tachyporus jocosus Say. Anchorage.
Tachyporus chrysomelinus Er. Dawson.
Boletohius cincticollis Say.? Anchorage; McKinley Park.
Mycetoporus humidus Say. Eagle.
Mycetoporus nigrans Makl. McKinley Park, one example
which fits Maeklin’s description very perfectly. His type
was said to be from the interior of the Kenai Peninsula.
"^Gymnusa variegata Kies. Fairbanks, one example, differing
slightly from a European specimen in my collection.
"^Myllcena sp. Scow Bay; Ketchikan. No species of this
genus has hitherto been recorded from Alaska.
Placusa complanata Er. Dawson.
Gyrophcuna sp. Anchorage. This is perhaps geniculata Makl.
*Atheta divisa Mark. Anchorage ; McKinley Park.
Atheta aquatica Thom. ? Dawson.
Atheta graminicola Grav. Skagway ; Fairbanks ; White Horse.
Atheta picipennis Mann. Anchorage.
Acrotona fungi Grav. Skagway ; Anchorage ; Juneau.
'^Gnypeta sp. Fairbanks. First record of the genus from
Alaska.
Gnypeta sp. Dawson.
Aleochara bipustulata L. Anchorage ; Fairbanks ; Dawson.
Aleochara pleuralis Csy. Dawson.
Dasyglossa prospera Er. Tanana River. The greater num-
ber of the Aleocharinae taken are as yet undetermined.
FAMILY PSELAPHID^
Sonoma parviceps Makl. Ketchikan.
Trimium clavicorne Makl. Ketchikan, one pair.
*Richenbachia binodifer Csy. Ketchikan. Scow Bay.
^Rybaxis brendeli Lee. var. Anchorage.
^Tychus cognatus Lee. Seward; Ketchikan.
FAMILY PTILIID^
Ptenidium pullum Makl. Scow Bay.
Acratichis {Trichopteryx || Kby.) sitkeensis Mots. Sitka.
Acratrichis laticollis Makl. Seward.
Acratrichis sp. Anchorage.
152
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
Acratrichis sp. McKinley Park.
Acratrichis three species. Dawson.
FAMILY HISTERID^
Hister (Psilocelis) subopaca Lee. var., or very closely allied
species. Dawson, one example; White Horse (Kusche).
FAMILY LAMPYRID^
*Podabrus comes conspiratus Fall. n. var.
This name is proposed for a form of which a single example
was taken at Skagway. It may or may not be a form of the
Californian comes, the amount of variability of which within
specific limits is not definitely known. The present specimen
differs from all my comes (so labelled) by the slightly more
transverse thorax, which is a little less rounded on the sides
and without visible sinuation at the hind angles, which are not
perceptibly prominent. The thorax also lacks the median eroded
line and has the front margin blackish at middle. It agrees
otherwise in color with the typical comes except that the pos-
terior ventral segments are not margined with yellow, the ter-
minal segment being merely feebly paler at tip. Length, 14 mm.
Skagway, Alaska, June 18-20, 1924; a single female.
*Podabrus tetragonoderus Fall. n. sp.
Moderately elongate, black, clypeus at sides, sides of pro-
thorax and basal three joints of antennae beneath, yellow; elytra
with a grayish aspect due to the rather plentiful ashy pubes-
cence. Antennae longer than half the body; second joint a
little narrower but subequal in length to the third ( 6 ), or
a little shorter than the third ( 2 ) ; third joint subtriangular, a
little longer than wide, distinctly wider than the fourth in the
male, less noticeably so in the female; following joints nearly
parallel-sided, becoming gradually more slender, the fourth a
little more than twice, the outer joints fully three times, as long
as wide. Head ( 6 ) just perceptibly narrower than the thorax;
eyes moderate; the eyes less prominent and the head narrower
in the female; clypeus and front as far back as a line joining
the posterior part of the eyes nearly smooth ; occiput closely,
rugosely punctate but shining. Prothorax quadrate, a little wider
than long, not or barely perceptibly wider near the front angles ;
sides nearly parallel and straight ; a very faint sinuation before
JANUARY, 1926] FALL ALASKAN COLEOPTERA
153
the sharply defined hind angles, front angles rounded, posterior
discal elevations rather strong and situated within the broad
dorsal black stripe, which is dilated behind the middle; sur-
face distinctly but not deeply punctate throughout, more finely
so in the yellow margins and between the discal elevations ;
median line finely eroded in posterior two-thirds. Elytra finely
scabrous punctate. Legs black, front tibiae in the single female
a little dilute in color. Claws ( 6 ) finely cleft, the inner por-
tion approximate to, and nearly as long as, the outer on the
front and middle feet, much shorter but still finely acute on
the hind feet. In the female the inner portion of the claw is
much shorter than the outer on all feet, and the claws might
perhaps better be described as having a basal dilatation with
the free angle sharply acuminate. Length, 7 to 9.5 mm.
Described from three males and one female, taken by the
writer at Skagway, Alaska, June 18-20, 1924.
This species will not enter any of LeConte’s groups as defined
though coming nearest to Group C. There are other species
in my collection with ungual characters similar to the present
one and I hope shortly to publish already prepared descriptions
of these and other new species of the genus.
*Podabrus fissilis Fall, n. sp.
Black, feebly shining; muzzle and two or three basal joints
of antennae beneath rufotestaceous. Antennae slender; second
joint but little more than half the third in the male, a little
shorter than the third in the female. Head conspicuously wider
than the thorax ( 6 ), or subequal in width to it ( 9 ) ; sur-
face alutaceous and opaque, front sparsely punctate. Prothorax
small, nearly as long as wide in the male, front angles obliquely
subtruncate, sides a little convergent thence to the somewhat
prominent hind angles. There is a small rufescent marginal
spot just behind the front angles in the male which is entirely
lacking in the female. Prothorax finely punctate and alutaceous,
sculpture as usual. Elytra very finely scabrous ; tarsal claws
all finely cleft in the male, the inner part shorter on the pos-
terior four feet ; all claws toothed at base in the female. Length,
6 to 7 mm.
Described from a single pair (type male) collected at
Emerald Lake, British Columbia, by Dr. A. Fenyes. This is
154
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
the first nearly entirely black species to be described having
tarsal claws all finely cleft in the male.
Since writing the above description I have taken a single
male of this species at Skagway, Alaska.
Podabrus piniphilus Esch. Skagway; Seward. The Seward
example (male) may be different .
^Podabrus extremus Lee. Skagway.
^Podabrus Iccvicollis Kby. Anchorage.
FAMILY CANTHARID^ (tELEPHORID^)
Cantharis (Tclephorus) mandibularis Kby. (= nigritulus
Lee. ) . Eagle ; Skagway ; McKinley Park ; Dawson ; White
Horse. This is a common species in Alaska and ranges
across the northern part of the continent to New Hamp-
shire. Mandibularis has long been erroneously placed as a
synonym of fraxini in our literature, but the above syn-
onymy is definitely established by comparison with one of
Kirby’s cotypes.
^Silis difficilis Lee. Skagway.
FAMILY MELYRID^
Dasytes hudsonicus Lee. Skagway ; White Horse ; Dawson ;
Eagle; Fort Yukon; commonly beaten from spruce.
FAMILY CLERID^
Thanasimus undulatus Say. Dawson.
Trichodes ornatus Say. Dawson.
FAMILY CORYNETID^
^Necrobia violacea L. Dawson; Fort Yukon (Kusche).
FAMILY MORDELLID^
^Anaspis atra Lee. ? Dawson ; Eagle.
Anaspis sericea Mann. Skagway ; Sitka ; Seward.
FAMILY PYTHID^
Priognatha monilicornis Rand. Skagway.
FAMILY ANTHICID.^
Anthicus mclancholicus Laf. Dawson.
FAMILY ELATERIDAj:
Athous ferruginosus Esch. Skagway; Sitka; Ketchikan. As
Dr. Van Dyke has pointed out in his recently published
Katmai List, this is but a slight variety of rufiventris Esch.,
which is the older name.
{To be continued)
JANUARY, 1926]
MASON NEW THRIPS
155
TWO NEW SPECIES OF THRIPS FROM
CALIFORNIA
BY ARTHUR C. MASON
Bureau of Entomology, U. S. Department
of Agriculture ^
Hoplandrothrips sycamorensis Mason, new species
Female. Length, 2.11 mm. Color uniformly dark brown,
with a very little scattered red hypodermal pigmentation ; tarsi,
fore tibiae, and bases of antennal segments III to VI light
brown.
Head nearly 1.5 times as long as broad, widest immediately be-
hind the eyes and converging slightly posteriorly; frons overhang-
ing about half of the first antennal segment. Cheeks nearly straight
and somewhat roughened with small warts, and having several very
short blunt spines. Postocular bristles short, dilated, and truncate
at tip. Ocelli small, circular and equidistant, placed between the
eyes; anterior one near the end of the vertex and the posterior pair
about opposite the center of the eyes. Eyes small, occupying little
more than one-fourth of the length of the head. Mouth-cone long,
almost reaching the mesothorax and running to a sharp point. An-
tenna eight-segmented and one and one-fourth times the length of
the head. First two segments concolorous with the head. III light
yellow with a little brown shading near the tip, IV to VI yellow at
base, shading into brown, VII and VIII light brown, VII having
a very little yellow at base. Segment I a short truncated cone,
II cup-shaped with a rather pronounced pedicle, III wedge-shaped
with rounded apex, IV and V ovate with narrowed pedicle, IV being
considerably wider than V, VI fusiform, VII truncate ovate, VIII
conical with a blunt tip.
Prothorax little more than half as long as the head and about
twice as wide as long, sides diverging rapidly posteriorly; one rather
prominent but short spine on the anterior margin and a row of five
spines on the posterior margin, the first of them being especially
heavy. All spines dilated and truncate at tip.
Pterothorax slightly wider than the prothorax and narrowed some-
what posteriorly. Wings rather short and colorless, except for a
little yellow shading at the extreme base; three blunt spines at base
of fore wings, five to nine (usually eight) interlocated hairs on pos-
terior margin. Legs rather long, the fore femora being heavy and
thickened; fore tibiae and extreme tips of others light yellow. Fore
tarsi armed with a short, stout tooth near base.
Abdomen large, tapering gradually after the fourth segment and
rounded abruptly from the seventh; provided with numerous blunt
1 Published with the permission of the Secretary of Agriculture.
156
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
colorless bristles, the posterior ones and those at the end of the tube
being nearly as long as the tube. Tube half as long as the head and
tapering gradually.
Measurements: Head, length 0.32 mm., width 0.23 mm.; prothorax,
length 0.18 mm., width 0.37 mm.; pterothorax, width 0.38 mm.; abdo-
men, greatest width 0.408 mm.; tube, length 0.15 mm., width at base
0.073 mm., width at apex 0.043 mm.; antenna, total length 0.41 mm.
Segment 12345678
Length 34 54 67 72 59 49 46 28 microns
Width 43 35 38 39 32 29 25 16 microns
Male slightly smaller and with more red pigmentation; otherwise
similar in appearance. Length, 2.03 mm. As with the females, the
fore tarsi are provided with a stout tooth and in addition they have
a smaller tooth on the inside of the fore tibiae near the base and
also a very small one opposite this on the femur.
The larvae are light yellow to colorless, and live under the
bark of the host. Winter is passed in the egg stage, under the
bark.
Described from five females and one male taken under bark
or on the leaves of sycamore trees at Springville, California.
Type in author’s collection.
This species is close to H . xanthopus Hood, but differs in
many characters, including larger size, darker color, absence
of most of the red pigmentation, shorter postocular and pro-
thoracic bristles, etc.
Zygothrips californicus Mason, new species
Length, 1.35 mm. Color brown to dark brown or fuscous;
legs concolorous with the body; antennal segment III yellow,
IV to VIII light brown.
Head slightly longer than broad, rounded in front and slightly
diverging posteriorly, reaching the widest part about three-fourths
of the way back, then narrowing slightly to the posterior end.
Cheeks smooth. Postocular bristles very small and blunt. Eyes
rather small, occupying about one-fourth of the length of the head
and one-third of its width. Ocelli rather large, the posterior pair
being opposite the anterior third of the eyes and the anterior one
being directed forward. Mouth-cone contracted, reaching about two-
thirds across the pronotum, bluntty rounded. Antenna eight-seg-
mented, one and one-half times the length of the head; bristles short
but sense-cones rather conspicuous. First two segments concolorous
with the head; III light yellow; IV to VI brownish-yellow to light
brown; VII and VIII light brown. Segment I a short truncated
cone; II ovate; III clavate with a broad base; IV ovate, tapering
abruptly to a narrow base; V barrel-shaped; VI cylindrical, with nar-
JANUARY, 1926]
MASON NEW THRIPS
157
rowed base; VII oblong and slightly tapered at tip; VIII conical,
tapering gradually to a rather sharp point.
Prothorax about three-fourths as long as the head and nearly
twice as wide as long, the sides gently diverging and broadly rounded
at the posterior end; prothoracic bristles short and blunt.
Pterothorax slightly wider than the prothorax, sides nearly straight
and converging slightly posteriorly. Wings large and clear, with
long membrane extending to or slightly beyond the end of the tube.
Fore pair somewhat constricted in the middle and with nine to eleven
interlocated hairs on the posterior border. Legs long, femora en-
larged, the fore femora especially being much thickened; fore tarsi
unarmed.
Abdomen rather heavy, width about four-ninths of the length, and
tapering rather abruptly from the fifth segment to the tube. Tube
less than half the length of the head; terminal bristles little more
than half the length of the tube.
Measurements: Head, length 0.229 mm., width 0.209 mm.; pro-
thorax, length 0.152 mm., width 0.276 mm.; pterothorax, width 0.304
mm.; abdomen, greatest width 0.342 mm.; tube, length 0.098 mm.,
width at base 0.057 mm., width at apex 0.033 mm.; antenna, total
length 0.328 mm.
Segment 12345678
Length 25 41 49 56 49 41 41 21 microns
Width 32 29 29 37 29 25 18 10 microns
Described from a single female taken on an orange tree at
Orange Cove, California. Type in author’s collection.
Close to Z. harfi Hood, but differing in the following charac-
ters : Larger size, antennal segments IV to VIII lighter colored,
prothoracic bristles blunt, clear fore wings and with 9 to 11
interlocated hairs, contracted mouth cone, and tube less than
half the length of the head.
NOTES ON PARASITES AT SATICOY, CALIFORNIA,
DURING THE YEAR 1925
BY STANLEY E. FLANDERS
Satie oy, California
On February 28 a braconid of the genus M onogonogastra
was reared from a walnut twig which contained the dead larvae
of Polycaon confertus Lee. During April numbers of these
dusky-winged, red-bodied braconids were observed about the
walnut leaves. One individual was seen December 4.
Through June and July a number of the metallic-blue ichneu-
158
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 3
mon-flies Amhlyteles cceruleus (Cr.), and the yellow and black
A. zehratus (Cr.) were reared from the pupse of Sahulodes
forficaria Guen.
Hovering- about the blossoms of a large eucalyptus tree on
April 2 were many of the syrphid parasites, Homotropus deco-
rat us (Cr.).
During October a black chalcid, Zatropis tortricidis Craw-
ford, was reared from the linia-bean pod-borer, Etiella zincke-
nella Treit. It pupates free in the pod, and is in the pupal stage
seventeen days.
On June 11 the gray, oval larvae of a species of Euplectrus
were observed in a cluster on the dorsum of a caterpillar of
Sahulodes forficaria. Later, several of these parasitized cater-
pillars were found.
From the pupae of the lichen caterpillar, Illice nexa Boisd,,
a number of specimens of the tachinid, Pseudochceta argenti-
frons Coq., were reared in May.
On June 20 three females of Epiurus sp. (possibly Pimpla
albipes Cr. described from Mexico) were reared from Ar chips
argyrospila Walk. Their silken cocoons were spun on walnut
about the remains of the caterpillar.
Six small chalcids of the genus Elachertus were reared in
June from a caterpillar of Ar chips argyrospila.
On April 29 a walnut aphis, Chromaphis juglandicola Kalt.,
was found parasitized by Aphelinus perpallidus Gahan. It is
only occasionally that such a parasitized aphis is found.
While collecting codling moth larvae from burlap bands on
walnut trees during March, a light brovm ichneumon-fly, Exe-
tastes n. sp., was taken from a codling moth cocoon.
Between May 1 and July 21 twenty-four tachinids of the
genus Lixophaga were reared from three hundred overwinter-
ing codling moth larvae. The maggot pupates within twelve
hours after leaving the caterpillar. The length of the pupal
stage was nineteen and twenty days.
The identification of the parasites noted were kindly made
by the specialists, R. A. Cushman, A. B. Gahan, S. A. Rohwer,
J. M. Aldrich, and P. H. Timberlake.
JANUARY, 1926] KESSEL LASIOPHTICUS PYRASTRI
159
SEX-LIMITED POLYCHROMATISM IN
LASIOPHTICUS PYRASTRI (LINN.)
BY E. L. KESSEL
Early last June I collected a large population of the common
syrphid fly Lasiophticus pyrastri (Linn.) on the University
Farm at Davis, California. This series, which was taken by
sweeping so as to be representative of the fauna, numbered 277
individuals. When a classification was made, there were found
to be 235 specimens of the normal bicolored phase, of which
number 98 were females and 137 were males. This is a de-
cidedly unbalanced sex ratio. The remaining 42 members of
the population were of the melanic variety, described by Curtis
as unicolor. It was observed that all of these melanic flies were
females, and it is interesting to note that when their number
is added to the 98 females of the bicolored type there appears
the well-balanced sex ratio for the whole population of 140
females and 137 males.
The above data lead to the conclusion that the melanic form
unicolor appears only in the female sex. Moreover, Curtis, in
his description of this variety, gives the impression that he
had but one specimen, and his colored illustration is definitely
that of a female. Still more evidence was obtained from an
examination of the material in the collection of the California
Academy of Sciences. This series contains thirteen specimens
of unicolor collected in various parts of California, and all
of these are females. The evidence, therefore, appears to be
conclusive.
An intermediate form, although comparatively rare, also
occurs. I took three specimens of this variety in Davis last
June, but these were obtained by selective collecting. They
are characterized by having the yellow markings persisting on
the second abdominal segment, although they are much reduced
in size. The California Academy of Sciences has one such indi-
vidual collected by Dr. E. C. Van Dyke on Mount Ranier,
Washington, in 1920. Two other specimens are cited by Cole
and Lovett in their “Diptera of Oregon.” All six of these in-
termediates are also females. We may, therefore, assume that
this variety, as well as the completely melanic one, is sex-
limited to the female. The species, therefore, appears to be
an example of sex-limited polychromatism, in which there is
only one type of male but three types of females.
THE PAN-PACIFIC ENTOMOLOGIST
Published quarterly by the Pacific Coast Entomological Society
in co-operation with the California Academy of Sciences
E. P. Van Duzee, Editor F. E. Blaisdell, M. D., Treasurer
Through an oversight the legend was omitted from the plate
of Hypolimnas holina L. in the October number of this journal.
A gummed slip to be attached to this plate is enclosed.
On March 23 the Pacific Coast Entomological Society gave
a complimentary dinner to Dr. and Mrs. William Barnes of
Decatur, Illinois, and Dr. and Mrs. R. J. Tillyard of Nelson,
New Zealand. Dr. Barnes gave the society an interesting
account of his work on the North American Lepidoptera and
Dr. Tillyard on his work at the Cawthorn Institute of Scien-
tific Research at Nelson. Dr. Barnes spent several days at the
Academy of Sciences, and his suggestions added much of value
to the Academy collection of Lepidoptera.
The recent death of Mr. Richard H. Stretch at his Seattle
home has removed one of the few remaining landmarks of early
western entomology. About fifty years ago he amassed a very
complete collection of the Bombycine moths of the world. This
he presented to the University of California. So much of this
collection as has escaped the ravages of pests has recently come
into the care of the California Academy of Sciences as a per-
manent loan, where it will be accessible to those interested in
the published work of Mr. Stretch.
Mr. E. Piazza, well known to our Lepidopterists as a suc-
cessful collector of moths, died in New York, March 9, 1926.
while on his way to England to visit his sister. Although an
invalid for years he has been able to do much valuable work
in the collection of moths for Scientific Institutions.
Through the generosity of friends we are able to give our
subscribers sixteen extra pages in this number of the Pan-
Pacific Entomologist. This enables us to include two larger
papers that otherwise it would have been impossible to publish
in full. The April number should follow this with little delay.
The October number was mailed January 16, 1926.
Dr. F. E. Blaisdell writes us that it will be impossible for
him to determine Coleoptera for his correspondents after Sep-
tember 1, 1926, and asks that we call the attention of coleop-
terists to this notice.
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Vol. II
April, 1926
No. 4
THE
Pan-Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
FREEBORN, A NEW CHAOBORID GNAT 161
VAN DUZEE, A NEW SPECIES OF SCENOPINID^ 164 -
MACDOUGALL, SOME NEW SPECIES OF MACROSIPHUM FROM BRITISH
COLUMBIA 165
DAVIS AND SMITH, NOTES ON TPIE GENUS STENOPELMATUS 174
FOX, SOME NEW SPECIES OF SIPHONAPTERA 182
BARNES AND BENJAMIN, ON THE PLACEMENT OF “PSEUDANARTA” FLAVI-
DENS GRT 187
BARNES AND BENJAMIN, ON THE DISTRIBUTION OF OMPHALOCERA OCCI-
DENTALIS B. AND B 188
TANNER, A NEW SPECIES OF PLASTOCERID^ IN EUTHYSANIUS .... 188
FALL, A LIST OF THE COLEOPTERA TAKEN IN ALASKA IN 1924 .... 191
BEAMER, NOTES ON GRIBURIUS MONTEZUMA SUF 209
ESSIG, A BUTTERFLY MIGRATION 211
FOX, A NEW NOMADA 212
DOANE, CONTROLLING MEALYBUGS ON ORNAMENTAL PLANTS 213
/
EDITORIAL 216
San Francisco, California
1926
THE PAN-PACIFIC ENTOMOLOGIST
Published quarterly in July, October, January and April by
the Pacific Coast Entomological Society in co-operation with
the California Academy of Sciences.
Annual subscription $2.00 in advance for the United States
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be sent to the treasurer, Dr. F. E. Blaisdell, 1520 Lake Street,
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Pacific Entomologist.”
Manuscripts for publication and communications regarding
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Duzee, California Academy of Sciences, Golden Gate Park, San
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Subscribers failing to receive their numbers will please notify
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PUBLICATION COMMITTEE, PAN-PACIFIC
ENTOMOLOGIST
E. O. Essig, Chairman
G. F. Ferris R. A. Doane
E. C. Van Dyke Grant Wallace
REGIONAL MEMBERS
W. W. Henderson, Logan, Utah
J. C. Chamberlin, Riverside, California
E. P. Van Duzee, Editor
E. C. Van Dyke^ Associate Editor
F. E. Blaisdell, Sr., Treasurer
Published at the California Academy of Sciences, Golden Gate Park, San
Francisco, California.
SP
J-Class matter, February postoffice at
THE
Pan -Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
VOLUME TWO
July, 1925, to April, 1926
San Francisco, California
1926
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CONTENTS
Barnes, William, and Benjamin, F. H.
New and Rare Lepidoptera from the Southwest... 12
Notes on Arizona Phalasnidse 16
Notes on Lophoceramica artega Barnes 63
A New Pyralid from California with Notes on Deca-
turia pectinalis 64
The Hyperboreus Group of Hepialus 81
Notes and New Species 106
Two New Western Phlaenidae Ill
On the Placement of Pseudanarta flavidens Grote 187
On the Distribution of Omphalocera occidentalis
B. and B 188
Beamer, R. H.
Notes on Griburius montezuma 209
Blaisdell, F. E.
Revised Check-List of the Species of Eleodes Inhabit-
ing America, North of Mexico, Including Lower
California and the Adjacent Islands 77
Thomas Lincoln Casey 90
Chamberlin, J. C.
A New Species of Lepidosaphes from China 85
Chamberlin, R. V.
Notes on Some Centipeds and Millipeds from Utah 55
Chamberlin, W. J.
The Coniferous Trees of the United States with the
Scolytid Beetles Said to Attack Them 23
Davis, A. C., and Smith, L. M.
Notes on the Genus Stenopelmatus with Description
of a New Species 174
Doane, R. W.
Controlling Mealybugs on Ornamental Plants 161
Essig, E. O.
A Butterfly Migration 211
Fall, H. C.
A List of the Coleoptera Taken in Alaska and Adja-
cent Parts of Yukon Territory in the Summer of
1924 ...127, 191
Flanders, S. E.
Notes on Parasites at Saticoy, California, during the
Year 1925 157
Fox, Carroll.
Some New Siphonaptera from California. 182
IV
Fox, C. L.
A New Nomada - 212
Freeborn, S. B.
A New Chaoborid Gnat, Chaborus lacustris 161
Hamlin, J. C.
Important Opuntia Insects of the United States 1, 97
Herms, W. B,
Entomological Observations on Fanning and Washing-
ton Islands, Together with General Biological Notes 49
Hubbell, F. H.
A New Species of Pristoceutophilus from the Olympic
Mountains, Washington 39
Kessell, E. L.
Sex-Limited Polychromatisni in Lasiophticus pyras-
tri L - 159
Larson, A. O.
Further Notes on Human Suffering Caused by Mites.. 93
Macdougall, Alice P.
Some New Species of Macrosiphum from British
Columbia 165
Mason, A, C.
Two New Species of Thrips from California.... 155
Osborn, Herbert.
A New x^cinopterus from Southern California ... 22
Rockwood, L. P.
On Night Flying and Attraction to Light in Acrididae
and the Relation of Meteorological Conditions
Thereto 36
Shannon, R. C.
North American Sphecomyia 43
Tanner, V. M.
A New Species of Plastoceridas in the Genus Euthy-
sanius 188
Van Duzee, E. P.
A New Mirid from California 35
Van Duzee, M. C.
A New Species of Scenopinidae from California 164
Van Dyke, E. C.
New Species of Carabidae in the Subfamily Harpalinse,
Chiefly from Western North America ...65, 113
Wachter, Sibyl.
The Hatching of the Eggs of Peripsocus californicus
Banks
87
The Pan-Pacific Entomologist
Vol. II, No. 4 April, 1926
A NEW CHAOBORID GNAT, CHAOBORUS LACUS-
TRIS SP. NOV. (CHAOBORIDT:; DIPTERA)
BY STANLEY B. FREEBORN
University of California, Berkeley, California
For several years complaints have been received at the
University from the vicinity of Clear Lake, Lake County,
California, concerning the pest of gnats which inundates the
country near the lake throughout the summer months. These
insects are non-biting but are so numerous that they make life
almost unbearable by flying into the eyes and mouth, con-
taminating food and filling store windows inches deep with
their dead bodies if white lights are exposed in the evening.
Nearly all the householders and campers in the vicinity of the
lake have been forced to use red shaded lights for illumination
to prevent being swamped with these insects.
An examination in 1925 proved them to be chaoborids of the
subgenus Sayomyia and they were provisionally determined as
Chaohorus (Sayomyia) astictopus Dyar and Shannon. A care-
ful study of new material proves them to be a distinct species,
the description of which follows :
Chaoborus (Sayomyia) lacustris Freeborn, sp. nov.
Male. Head: Pale brown with vestitiire of straw-colored hairs.
Tori large, occupying nearly whole width of face, pale brown, api-
cally excavated and bearing plumose antennae; antennae translucent
except at whorls, hairs of which are golden yellow. Clypeus flat-
tened, prominent, densely covered with golden hairs, pale brown and
darker at margins. Proboscis and palpi transparent with brown
markings. Eyes dark red.
Thorax: Prothoracic lobes large but not contiguous, anterior half
rich brown, posterior white. Posterior pronotum white with brown
stripe along dorsal margin and slight infuscation on posterior margin.
Mesonotum white shading to cream with rich brown markings bor-
dered by lines of darker brown arranged as follows: two median
stripes tapering posteriorly starting at anterior border and ending at
antescutellar space, two subdorsal stripes tapering posteriorb^ start-
162
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
ing on either side of median stripes at point midway of disc and
ending at scutellum; faint triangular spot with base at scutellum and
apex pointing between two median stripes; pale circular spot indefi-
nitely defined between anterior end of each subdorsal stripe and
anterior margin. Scutellum white at center with brown markings at
each side, vestiture of golden hairs. Postscutellum dark brown with
a median, V-shaped, white cleft at the anterior margin. Pleurae pale
cream with numerous brown markings, most conspicuous of which
is that covering lower three-fourths of the sternopleurite. Prothoracic
sclerites densely haired, others smooth, bare except four upper mese-
pimerals. Halteres entirely white. Legs densely hairy, creamy white
with brown markings; forelegs more heavily pigmented than others;
femora with subapical brown bands and brown spots at hair inser-
tions, tibiae with subbasal and subapical brown bands and brown
spots at hair insertions, basitarsus as long as next two segments,
each segment with apical brown ring growing fainter on the distal
segments. Claws simple. Wing veins clothed with narrow, hair-like
scales, anterior margin clothed with narrow, appressed scales, pos-
terior margin with two lengths of lanceolate scales, one short and
other longer than greatest distance between anal vein and hind
margin. Slight brownish stains on wing membrane at base of R2,
at fork of R2-1-3, at r-m and m-cu. Marginal distance between termi-
nations of R1 and Sc almost twice that between R1 and R2, Fork
of R2 and R3 distinctly distad of fork of MH-2 and M3+4. R2
slightly down-curved at margin. Crossveins r-m and m-cu forming
continuous line. Cu2 becomes evanescent at margin, but appears to
incline forward and run parallel with the margin. Tillyard’s “Cu2”
is fully as prominent as the so-called Cu2 paralleling Cu2 to point
near margin, anal vein prominent, down-curved near margin and
paralleled by prominent fold, another anal parallels hind margin.
Abdomen: First segment translucent white, wider than long;
second to seventh longer than wide, translucent white, lateral mar-
gins of tergite and sternite dark brown, the coloring becoming
progressively narrower as the mid-dorsum and mid-sternum is
approached, giving appearance of lateral triangles; seventh largely
brown; eighth funnel-shaped, one-fourth length of seventh and
rotated 180 degrees.
Terminalia: Gonostyles (side pieces) slender at base expanding to
tip, stained with brown on outer half; a subapical, rounded lobe
bears three and sometimes four stout spines, the inner face from base
to lobe heavily studded with bent hairs not markedly different in
size from spines of lobe; dististyles (claspers) bent, pale at base,
brown at tip, which is very slightly expanded, no appendage (claw).
Paraprocts (tenth sternites) curving around base of gonostyles with
onl}'^ terminal points parallel to the latter, pale at base, deeply pig-
mented at tip which is cleft in two parts so that they resemble
inverted high-heeled cow-puncher’s boots. In balsam mounts two
APRIL, 1926]
FREEBORN CHAOBORUS
163
paraprocts are straightened and take V-shaped position with ter-
minal ends pointing laterally and caudally. Length of male 5 mm.
Female. Similar in coloration to male except that pigmentation
is more pronounced and deeper brown; triangle in antescutellar space
not always distinct; first abdominal tergite with dark pigmentation,
dark markings of venter limited to scattered spots; tarsal rings more
distinct than in male^ last segment all brown; stained areas on wings
more pronounced and numerous, all forks and most marginal termi-
nations of veins with stained area. Palpi with basal segments swollen,
the terminal one as long as combined length of two segments pre-
ceding, slender, annulated. Abdominal segments wider than long.
Claws simple. Length, 4 mm.
Type, male, California Academy of Sciences, No. 1873, Lake-
port, Lake County, California, July 19, 1925.
This species falls near C. punctipennis Say and C. astictopus
D. and S. It differs from both in having very pronounced
markings on the mesonotum and in the form of the paraprocts.
The abdominal markings in the female agree closely with those
of astictopus and the chaetotaxy of the gonostyle is very similar
to that of punctipennis.
LABOPS BURMEISTERI STAL
In the March, 1926, number of the Canadian Entomologist
Dr. Knight requests information regarding the occurrence of
Lahops burmeisteri in New York State. With the consent of
Mr. Evans I retained the specimen of this species which I
determined in 1909 and a reexamination of this specimen shows
it to be the true burmeisteri as determined by Dr. Knight. At
the time I made this determination I had a second specimen
from the Adirondack Mountains of New York, which passed
through my hands for study, so the New York determination
can be depended upon. I have not time now to look up the
Alaskan record and it is possible that may refer to the form
later described as tumidifrons Kngt.
In my material of hirtus Kngt. one specimen from New
Mexico has the tibiae altogether black with the long tibial hairs
and male claspers of hirtus, so the color of the tibiae cannot be
relied upon in separating these forms. Labops hesperius Uhl.
as restricted by Knight is still unknown to me. — E. P. Van
Duzee.
164
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
A NEW SPECIES OF SCENOPINID.E FROM
CALIFORNIA (DIPTERA)
BY M. C. VAN DUZEE
Buffalo, Nezu York
Pseudatrachia albocincta Van Duzee, new species
Male. Length, 6 mm.; of wing, 4 mm. Head, thorax, scutellum
and abdomen black, somewhat granulate^ nearly bare; apical seg-
ment of abdomen, hypopygial appendages and a space above fore
coxse with a few pale hairs; palpi 3 ''ellowish white; vertex and upper
orbits shining black; antennas brown, tip of second joint more yel-
low; a small spot back of humeri and another back of wing yellow,
these spots connected by a narrow white line, abdomen with bronze
reflections above; second, third and fourth segments each with a
conspicuous white elongate spot on the hind margin at the sides,
these spots connected by a narrow white line on the venter and
nearly so on the dorsum.
Coxae, femora, tibiae and tarsi yellow; middle and hind femora
with a brown stripe below, the latter brown at tip above; tips of
tibiae a little darkened; tarsi slightly brownish; hairs on legs minute,
white; knobs of halteres white, petiole brown. Wings hyaline, veins
yellowish. Described from one male.
Type, male, No. 1874, Mus. Calif. Acad. Sci., taken by E. P.
Van Duzee, May 23, 1920, at Bradley, Monterey County,
California.
Table of North American Species
1. Head 3 '^ellow - 2
- Head black 3
2. Mouth parts and scutellum yellow (New Mexico)
griseola Coquillett
- Mouth parts and scutellum black (Arizona) ftaAceps Coquillett
3. Legs yellow..-- - 4
- Legs black or brown 5
4. Second and third abdominal segments white on posterior margin
both above and below (California) albocincta new species
- Abdomen without white margins to the segments (New Mexico;
California) unicolor Coquillett
5. Dorsum of thorax and the femora with long mixed white and
black hair; hind margins of abdominal segments 2-5 white
(Arizona) pilosa Coquillett
- Hair on thorax and femora short; length, 8 mm.; abdomen
rather long and slender (Mexico) longtirio Loew
APRIL, 1926 ]
MACDOUGALL MACEOSIPHUM
165
SOME NEW SPECIES OF MACROSIPHUM FROM
BRITISH COLUMBIA (HOMOPTERA, APHIDID^)
BY ALICE P. MACDOUGALL
University of Toronto
The following species of Macrosiphum are described from
collections made in British Columbia during the summers of
1923-25 inclusive. All measurements are in millimeters and
whenever possible are an average of a number of specimens.
Length of body is measured from the vertex to the tip of cauda.
The width given is that of the widest part of the abdomen.
Owing to the large area covered and the great distance between
various collecting points, it was impossible to take a complete
series in every case. However, these descriptions will, it is
hoped, serve as a basis for further study of some interesting
species from a comparatively new territory and aid in their
identification by workers in these districts.
1. Macrosiphum audeni Macdougall, n. sp.
Fig. I
This species occurs in dense colonies on the upper sides of
the leaves of N ymphaea polysepalum, clustering toward the
petiole. The specimens from which the descriptions were made
were collected by Mr. K. F. Auden in the summer of 1924 at
Merritt, British Columbia, after whom the species is named. It
was taken also by Mr. G. Spencer near Victoria, British
Columbia, in 1925, and by the author the same summer at
Lakelse Lake in the northern part of the province. It is thought
to be comparatively rare as many plants have been worked over
in these and other localities which were all quite uninfested. Its
distribution, however, as indicated by the above localities in
which it was taken, is fairly general for the province.
Apterous viviparous female
General appearance: Color pale green^ head and upper portion of
abdomen slightly darker; antennas light except segment VI and apex
of V, which are dusky; coxae and femora pale; tarsi and tip of tibia
dusky; cauda pale; cornicles pale, except tips which are dusky; eyes
dark red; body long and fairly slender; cornicles not reaching to tips
of cauda, reticulated at tip where dusk}% distinctly wider at base;
166
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
cauda long, irregularly ensiform; 2-4 sensoria on ant. seg. III. Beak
fairly short, not reaching below second pair of coxae. Antennae much
longer than body.
Measurements: Body — length 3.5, width 1.4; ant. segs. — III .8,
IV .7, V .5, VI .19+1.2; corn. — .9; cauda — .4; hind tarsi — .19.
Alaie viviparous female
General appearance: Color as above: antennae dusky except first
two segments and extreme base of third; eyes dark red; legs pale
except distal tips of tibia and femora and all of tarsi, which are
dusky; basal half of cornicles pale, rest dusky; cauda clear. Body
long and fairly slender, except when distended with embryos; where
dusky cornicles are distinctly reticulated, long^ slender; cauda ensi-
form, fairly hairy; basal two-thirds of ant. seg. Ill with row of
12-18 sensoria. Antennse slightly longer than body.
Measurements: Body — length 3.5, width 1.6; ant. segs. — III .75,
IV .72, V .65, VI .2-1.2; corn. — .8; caud — .3; hind tarsi — .17; fore
wing — length 4.2, width 1.45; hind wing — length 2.02, width .69.
All collections were made as recounted above on the follow-
ing dates : Merritt, July 25 ; Victoria, British Columbia, August
8; Lakelse Lake, July 28.
Type slides are deposited in the National Collection, Ottawa.
Paratypes in the United States National Collection, Wash-
ington, and in the author’s collectian.
This species is closely related to Macrosiphum aquilegia
Theo.
2. Macrosiphum bisensoriatum Macdougall, n. sp.
Fig. II
Found in abundance on the under side of leaves of Ribes
lacustre during the summer months. No alternate host is
known ; migrants appear at frequent intervals during the sum-
mer. This species can readily be distinguished from Myzus ribis
Linn, by the larger size and the fact that when disturbed they
are inclined to fall from the leaf.
Apterous viviparous female
General appearance: Color dark green, head slightly lighter than
the rest of the body; antennse pale except I and II, which are con-
colorous with body, distal quarter of V, and all of VI, which are
dusky; flange of cornicle dusky ^ rest clear; cauda abruptly ensiform,
clear; beak reaching almost to third pair of coxse; last segment of
APRIL, 1926]
MACDOUGALL MACROSIPHUM
167
leg shading to brown; tips of tibia and all of tarsi dark brown, rest
pale; head, cauda, ant. seg. Ill, and tibia prominently hairy. Two
small sensoria on proximal half of ant. seg. III. Tip of cornicle dis-
tinctly reticulated, rest imbricated. Body slender, graceful; antennae
much longer than bodjG cornicles reaching to tip of cauda.
*
Measurements: Body — length 3.38, width 1.7; ant. segs. — III 9.5,
IV .8, V .65, VI 16+1.15; cornicles — 1.1; cauda — .42; hind tarsi — 1.5.
Alate viviparous female
General appearance: Thorax very dark, rest as above; insect notice-
ably smaller than above. Coloring of antennal segments as follows:
I and II, concolorous with body, rest of antennae dusky; on dusky
portion of seg. Ill is a row of 9 sensoria; near tip of V, one large
one is found, and the usual number occur at base of spur of VI.
Eyes red; beak quite long, clear, except last segment which is uni-
formly dusky; coxae and basal two-thirds of femora pale, shading to
the light brown of the rest of the femora; tibia light brown except
tip, which is dark; tarsi dark brown; cornicles pale at base, shading
to dusky flange, slightly constricted near tip; this part imbricated;
cauda clear; wing veins distinct, dark brown; stigma rather pale.
Antennae much longer than body.
Measurements: Body — length 2.8, width 1.3; antennal segments —
III .86, IV .7, V .6, VI .17-.9; cornicles— .8; cauda— .32; hind
tarsi — 1.5; fore wing — length 3.7, width 1.3; hind wing — length 2,
width .6.
This species differs from Macrosiphum lactucce Pass in hav-
ing fewer sensoria in both forms and minor differences in color
and size. It was collected at Boundary Bay, British Columbia,
at intervals, through July and August, 1924.
Type slides are in the National Museum, Ottawa. Paratypes
in the National Museum, Washington, and in the author’s
collection.
3. Macrosiphum fuscicornis Macdougall, n. sp.
Fig. Ill
Found in colonies with Aphis oenother(B CEst. on stems and
leaf buds of Bpilohium angustifolium, the dark green color of
this form contrasting with the purple of the smaller one. It has
been taken in different parts of the province during the summer.
Alatse are comparatively rare, only having been found in June
and the latter part of August.
168
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
Apterous viviparous female
General appearance: Dark olive-green body; anteiinze brown
throughout, darkening somewhat in VI, much longer than body;
eyes red; legs uniformly dark except base of femora, which is pale;
cornicles brown except at base, reaching beyond the tip of cauda,
broader at base and slightly constricted near tip; cauda long, irregu-
larly ensiform, light brown.
Measurements: Body — length 3.18, width 1.6; ant. segs. — III 1.03,
IV .94, V .83j VI .2-|-1.12; cornicles 1.4; cauda .5; hind tarsi .18.
Alate viviparous female
General appearance: Color approximately as above; thorax slightly
darker than other parts of the body; aiitennse and cornicles strik-
ingly long; eyes dark red; first two antennal segments dusky, III
dusky except extreme base which is pale, IV getting paler, V and VI
quite light. A more or less regular rov/ of sensoria along edge of
III. Coxae and basal half of femora pale, rest of legs dusky; cor-
nicles dusky, slightly paler immediately below flange; cauda pale;
wing veins normal, stigma pale brown.
Measurements: Body — length 2.6, width .89; ant. segs. — III .81,
IV .8, V .72, VI 15-.8; cornicles — .9; cauda — .32; hind tarsi — 1.7; fore
wing — length 3.7, width 1.38.
This species has been collected at Jerico Beach near Van-
couver, British Columbia, on August 24, 1924, and Merritt,
British Columbia, June 29, 1924. It was observed in other parts
of the province during the same summer. It is most closely
related to Macrosiphum fragariellum Theo., but the cornicles
are tapering and there is no indication of a swelling.
Type slides in Canadian National Collection, Ottawa. Para-
type slides in United States National Museum, Washington,
and in the author’s collection.
4. Macrosiphum nigromaculosum Macdougall, n. sp.
Fig. IV
This very colorful Aphis is found, so far as the writer knows,
only in the semi-arid valleys of the lower Cascades. It is
clustered in masses along the stems of Rosa nutkana, seldom
spreading, to the buds or leaves. Although collected from June
to August, no adult or larval migrants were taken.
Apterous viviparous female
General appearance: Body bright red, fading to pink; cervicum
APRIL, 1926]
MACDOUGALL MACROSIPHUM
169
black, abdomen marked with three more or less regular black bands;
appendages black, body broadly ovate with cornicles projecting out-
ward at right angles giving the insect a most peculiar appearance.
Specimens in balsam appear as follows: head and abdomen dark with
lighter thorax^ antennal segments pale except ends of V and VI,
which are dusky. Eyes very dark red; base of femora pale, rest of
leg light brown, except tips of tibia and femora and all seven tarsi,
which are dark; cornicles uniformly black, very broad at base, taper-
ing gradually to the usual-sized flanged tip, reticulated throughout;
cauda irregularly ensiform, shading from pale at base to dusky at
tip; beak light brown except tip which is very dark, reaching to
second pair of coxae; antennae, tibia and cauda prominently hairy.
No sensoria on antennal segment III, but a very large one near tip
of V and the usual group at the base of spur of VI, scarcely as long
as the body. In cleared specimens the black pigment spots seem to
be scattered rather unevenly in those parts of the body which appear
banded.
Measurements: Body — length 3.7, width 1.9; ant. segs. — III .9,
IV .51, V .51, VI .15-. 74; cornicles — .7; cauda — .38; hind tarsi — .2.
Collections of this species were made in Bootahnie Valley,
June 28, 1924, and in Merritt on July 10 of the same year.
During these months and August, no alatse were observed in
either locality. Readily distinguished from other species occur-
ring on Rosa by color, shape and habits.
Type slides deposited in the Canadian National Collection,
Ottawa. Paratypes in United States National Museum, Wash-
ington, and in author’s collection,
5, Macrosiphum pyrifolias Macdougall, n. sp.
Fig. V
This species is fairly common in the vicinity of Merritt,
British Columbia, on the leaves of Pyrus occidentalis during
the summer. Although the sexes have not been found it is
thought to spend the complete season on this host. Migrants
occur regularly, a few of them being always present in a colony.
Apterous viviparous female
General appearance: Dusky brown in color^ head slightly darker
than the rest of the body; antennal segments V, VI and tips of III,
IV, dark brown, I and II concolorous with body; rest clear; eyes
red; legs pale except tarsi and tips of tibise, which are dusky; corni-
cles long, reaching just below tip of cauda, and pale, except distal
third, which is dusky; cauda clear; body long, of average width;
170
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
embryos show distinctly; dusky portion of cornicles distinctly reticu-
lated. Row of 14-17 sensoria on basal three-quarters of antennal seg-
ment III. Cauda typically ensiform. Beak reaching to third pair of
coxae; last segment dusky.
Measurements: Body — length 3.4, width 1.4; ant. segs. — III .9, IV
.75, V .7, VI .2-. 9; cornicles 1.1; cauda — 14; hind tarsi — 1.6.
Alate viviparous female
General appearance: Much darker than the apterous form; body
narrower and antennae longer; body dark green-brown; cauda, basal
half of femora, proximal part of antennal joint III, all pale; corni-
cles dusky, uniformly colored; wider than in apterae; only the tip
distinctly reticulated, barely reaching to tip of cauda; legs and an-
tennae covered with fairly long, capitate hairs. Antennal joint III
with 18-24 irregularly placed sensoria, more or less clustered in
center.
Measurements: Body — length 3.1, width 1.01; ant. segs. — III 1.1,
IV 8.5, V 7.4, VI .19-1.2; cornicles — 1.1; cauda — .43; hind tarsi — .19;
fore wing — length 4.55, width 1.6; hind wing — length 2.8, width .86.
Collected May 27, 1924, in Merritt, British Columbia, by Mr.
K. F. Auden, and in Tulameen on June 15 of the same year by
the author. It was observed at other times in this locality.
Nearest to Macrosiphum ulmarice Schrank, although it re-
sembles this species only remotely.
Type slides in Canadian National Collection, Ottawa. Para-
types in the United States National Museum, Washington.
6. Macrosiphum subviride Macdougall, n. sp.
Fig. VI
This species is found on the stems and leaves of Aster alpinus
which grows on the eastern slopes of the Cascade Mountains
and in the Rockies. As it does not colonize, it is very difficult
to detect on the pubescent surface of the host. Alatae are found
continuously throughout the summer months. Early in August,
however, the insects entirely disappear; the winter host is un-
known. Transfer experiments were made with Aster occi-
dentalis, but were found to be completely unsuccessful.
Apterous viviparous female
General appearance: Color delicate green, approximating that of
the hairs of the host; antennal segments I and II pale. III and IV
pale with dusky tips, V shading to dusky at tip, VI dusky; legs pale
except tip of tibia and all of tarsi, which are black; cornicle slightly
APRIL, 1926] MACDOUGALL MACROSIPHUM
171
constricted a short way from tip, this narrow part dusky and reticu-
lated; cauda clear, ensiform; eyes red. Antennal segment III with
an irregular row of 8-9 sensoria.
Measurements: Body — length 3.1, width 1.05; ant. segs. — III .89,
IV .68, V .59, VI .19-1.28; cornicles — .8; cauda — .38; hind tarsi — .12.
Alate viviparous female
General appearance: Same general color and shape as above; thor-
acic region slightly greener than the rest of the body; antennae very
dark except segments I, II, and base of III, which are pale and form
a striking contrast; coxae and proximal portion of femora pale, rest
light brown, except tarsi and distal sixth of tibia which are dark as
antennae. Wing veins regular, dark, radius slightly clouded, stigma
dusky. Proximal half of cornicle light, rest dark, reticulated, slightly
constricted near tip; cauda clear, broadly ensiform, hairs fairly
noticeable. 20-25 sensoria on ant. seg. III.
Measurements: Body — length 2.6, width .8; ant. segs. — III .91;
IV .88, V .8, VI .2-1.45; cornicles — .82; cauda — .31; hind tarsi — 1.1;
fore wing — length 4.65, width 1.7; hind wing — length 2.4, width .9.
First collected in Bootahnie Valley, June 27, 192'5, and then
at intervals until August 2 of the same year. This species differs
in minor points from M. circumflexa Buck, M. campanulce
Kalt. and M. solanifolice Ash.
Type slides in Canadian National Collection, Ottawa. Para-
types in United States National Museum, Washington, and in
the author’s collection.
Explanation of Figures
Fig. I. a, cornicle of apterras; b, segment III of antennae of apterae;
c, cornicle of alatae; d, antennal segment III of alatae; e, fore wing;
/, cauda of apterae; cauda of alatae.
Fig. II. a, cornicle of apterae; b, antennal segment III of apterae;
c, cornicle of alatae; d, antennal segment III of alatae; e, cauda of
apter^; /, cauda of alatae; g, fore wing.
Fig. III. a, cornicle of apterae; b, antennal segment III of apterae;
c, cornicle of alatae; d, antennal segment III of alatae; e, cauda of
apterae; /, cauda of alatae; g, fore wing.
Fig. IV. a, cornicle of apterae; b, cauda of apterae.
Fig. V. a, cornicle of apterae; b, antennal segment III of apterae;
c, cornicle of alatae; d, antennal segment III of alatae; e, cauda of
apterae; f, cauda of alatae; g, fore wing.
Fig. VI. a, cornicle of apterae; b, antennal segment III of apterae;
c, cornicle of alatae; d, antennal segment III of alatae; e, cauda of
apterae; /, cauda of alatae; g, fore wing.
172
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
APRIL, 1926] MACDOUGALL MACROSIPHUM
173
Fig, TV
fis, V.
174
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
NOTES ON THE GENUS STENOPELMATUS WITH
DESCRIPTION OF A NEW SPECIES
(ORTHOPTERA)
BY ALONZO C. DAVIS AND LESLIE M. SMITPI
In the course of some anatomical work which the writers
were doing' with two undoubtedly distinct species of Stenopel-
matus, it became necessary to know what species were being
dealt with. One of the species was of large size, with the head,
thorax, and legs usually light testaceous or somewhat fuscus.
This species was dominant along the Berkeley hills and rarely
extended its range into the low lying coastal shelf upon which
the cities of Oakland and Berkeley are situated. In the hills this
species was found only on grassy slopes, overhung by an out-
cropping of rock which had strewn the slope with weathered
fragments. Under these rocks this species constructs its
burrows.
The second species was much smaller, with the head, thorax
and legs usually rufo-testaceous. This species was dominant on
the low, flat belt bordering the San Francisco Bay in the neigh-
borhood of Oakland. It was seldom taken in the hills, where the
large species was fairly abundant. Throughout the residence
section of the city this small species is fairly common. It
burrows in the loose soil in backyard gardens and constructs
tunnels beneath boxes, boards, etc., which are closely pressed
to the ground. When living in the hills, it occurs in the same
type of habitat as does the larger species.
It was difficult to induce the large species to mate in the
laboratory, but when this was done, it was found that the
female pursues the male for several minutes prior to mating.
The eggs of this species measure 3.7 — 3.8 x 2.0 — 2.2 mm. These
insects can stridulate loudly enough to be heard at a distance of
six feet. Stridulation is accomplished by rubbing the inner face
of the hind femora against the opposing side of the abdomen.
The smaller species mated readily in the laboratory ; in this
case the male rapidly pursued the female while the latter in-
variably retreated. The eggs of this species measure 2.9 —
3.1 x 1.8 — 1.95 mm. Stridulating organs are present and the
femora are rubbed against the abdomien as in the large species.
APRIL, 1926] DAVIS AND SMITH STENOPELMATUS
175
but the human ear can but rarely detect the sound, even at close
range.
A rather thorough search of the literature on the genus failed
to net anything which would serve to identify these species.
Hebard's table (N. Y. Ent. Soc., vol. XXIV, pp. 70-86)' is by
far the best, but was inadequate in definitely determining indi-
vidual specimens. The spines and spurs of the hind tibiae, upon
which previous taxonomic work was largely based, are so
variable in size and number as to be almost worthless. Speci-
mens of the large and small species were sent to Mr, Caudell,
who says : “I should say that from the variability of the char-
acters used they all might be either longispina or fiiscns. The
simplest way out would seem to be to call all specimens of this
genus Stenopelmatus fuscus as all our crickets are now Gryllus
assimilis/’
In the collection of the California Academy of Sciences there
are specimens of Y. fuscus, longispina and pictus, about four-
teen specimens in all, which were identified by Hebard, appar-
ently on the basis of tibial spines, the relative length of
metatarsus and inner apical spine of the hind tibia. This latter
character, as has been pointed out, is not dependable and varies
from one extreme (S. fuscus) to the other (S. longispina) in
the same locality in Berkeley. From the above it would seem
that there is as yet no definite character by which these two
species may be separated. It is our opinion, based upon exami-
nation of such material as is available, field study, and a study
of breeding habits, that vS. fuscus and S. longispina are not
distinct species, and that the latter name should be made a
synonym of vS'. fuscus. After studying the literature, we have
arrived at the conclusion that S. fuscus is the larger of the two
species at hand and that the smaller species is as yet un-
described, probably having been overlooked in the past as a
nymph.
The genus having comparatively recently lost its wings, the
thorax is in an inconstant or transitional state, and relative
dimensions are not to be relied upon to any great extent. The
form and shape of the head is also inconstant, since the genus
is subject to magacephaly and other abnormalities. The mouth-
176
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
parts, in the series examined, show no characters of value.
The rugosity and dentition of the mandibles is fairly constant,
except as use reduces the teeth. The ovipositor is of some value,
when used comparatively. The male genitalia consist of a
fleshy bursa copulatrix which seemingly presents no characters
of value, and in any case is too difiicult to preserve to be of
value in identifying cabinet material. The punctuation of the
head shows differences of some value. Color is apparently not
very reliable, varying from very light to dark in the same
locality. In preserved specimens, unless in alcohol, it has been
difficult to use measurements of the abdomen because of the
great shriveling of the parts in drying. By injection and hard-
ening these soft parts may be preserved in more or less their
natural condition, but the size and shape of the abdomen varies
much with the condition of the insect, whether or not full of
food, eggs, parasites, etc. In general, the females are large and
full-bodied. The males are usually somewhat smaller, with the
legs longer and the abdomen smaller in proportion and laterally
compressed.
The characters which we have found to be of most value in
distinguishing between S. fuscus Hald. and S. intermedius
n. sp. are the shape of the anterior plate of the prosternum, the
size and shape of the stridulating plates on the inner surface of
the hind femora, punctuation of the head, basal antennal joints,
and size. A number of attempts were made to interbreed 5.
fuscus with S. intermedius. S. intermedius males were placed
with S. fuscus females and S. fuscus males with S. intermedius
females, but in no case did mating occur. No difficulty was
experienced in breeding each species within itself.
Stenopelmatus fuscus Hald.
Punctures distinct along the frontal (epicranial) suture^ especially
above and below the oblique arms between the eyes. On either side
with two vertical lines of punctures, one just behind the eye and
inclining slightly medially above, the other about halfway between
the eye and the suture; these thin out above, but attain the vertex
and in most cases extend to the anterior margin of the pronotum;
angle at which the two oblique arms meet the frontal suture fairly
obtuse; oblique arms nearly straight, their apparent ending being
near the upper margin of the insertion of the basal antennal joint.
Oblique sutures themselves wide and depressed. At and above the
APRIL, 1926] DAVIS AND SMITH STENOPELMATUS
177
fronto-clypeal suture, with a triangular depressed rugose area, the
sculpture tending to run in oblique lines from the center outward
and downward. Punctuation in some immature specimens sparse,
rarely failing to attain the vertex; lines of punctuation behind the
eyes sometimes nearly lacking. Blades of the ovipositor, in compari-
son with S. intermedins, somewhat longer, less rounded and not so
heavy when viewed from the side, when viewed dorsally, meeting at
a more acute angle. The basal antennal joints, while somewhat vari-
able may prove to be of some value; in general, the third joint is
subequal to the second, both together being equal to the first, the
fourth being one-third as long as the third.
The shape of the prosternum^ while somewhat variable, seems to
be constant enough to be of some value, at least in separating the
forms from any one region. It is divided into a large basal plate,
with wing-like lateral areas separated by postero-lateral notches and
a shallow longitudinal groove, and a smaller anterior plate, separated
from the basal by a deep groove. Basal plate large, flat, truncate
posteriorly; lateral notches quite sharp; grooves between the middle
and lateral plates shallow and ill-defined; lateral plates rather sharp,
narrowing anteriorly; anterior plate smaller, posterior margin regu-
larly arcuate, postero-lateral angles evenly rounded, the whole lateral
and anterior margin forming an even curve. In some specimens the
anterior plate is semicircular, and in a few the anterior margin is
squarely truncate, but in the majority of cases this plate is not more
than twice as wide as long, and in many cases is noticeably less.
Stridulating organ * consisting of very small, raised, chitinous
plates on the inner surface of the hind femora, compactly grouped
on the basal portion of the femur near its upper or anterior margin;
these plates arranged in rows which may be either straight or slightly
wavy, the direction of the rows being at an angle of about 30 degrees
with a line connecting the coxa and base of tibia (vertex of the angle
toward the coxa); the width of the rows varies from 10 to 12 microns,
but this variation cannot be correlated with age. In young or recently
moulted specimens the plates rise to a blunt point on their free mar-
gins; in older specimens wear has reduced the free margin to a
smooth flat curve, or to a straight line. The plates are in contact
with one another laterally.
Maximum
Minimum
Mean
Length of hind tibiae, 2
17.0 mm.
12.0 mm.
15.0 mm.
Length of hind tibiae, 6
19.0
17.0
18.0
Length of notum, 2
9.0
8.0
8.3
Length of notum, 6
9.0
9.0
8.3
Total length, 2
48.0
42.0
45.5
Total length, 6
37.0
35.0
36.0
* Mr. C. D. Duncan (E!nt. News, vol. xxxiv, No. 3, p. 74) figured the
differences in stridulating plates as a sexual difference in S. longispina
Brunner. Since we find both of these types of plates in both males and
females this cannot be a sexual difference. We would place his female
(fig. B) in S. intermedins and his male (fig. A) in s. fuscus. Duncan’s figure B
resembles very closely the stridulating plates of specimens from Pasa-
dena, which superficially resemble 5. fuscus.
178
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
Stenopelmatus intermedius Davis and Smith, n. sp.
Punctuation of head sparse, the lines of punctures barely attain-
ing the vertex; a diffuse finer punctuation over the whole frons in
addition to the coarser punctuation of the sutures and punctate lines.
Angle of junction of the frontal suture somewhat more acute, the
oblique arms curved, their apparent ending being above the insertion
of the basal antennal joints, the sutures themselves very fine and
even. Rugose area above the fronto-clypeal suture somewhat larger
in proportion, the rugosity transverse rather than oblique. Third
basal antennal joint longer than second, both together being sub-
equal to the first, the fourth one-third as long as the third. Ovi-
positor of female shorter, stouter, more rounded, more abruptly
curved upward as seen from the side, the blades meeting at a more
obtuse angle (about 40-45 degrees) as viewed dorsally. Basal plate
of prosternum fairly large, the postero-lateral notches wider and
more rounded; posterior margin squarely truncate in the female,
evenly rounded in the male; lateral plates wider in proportion^ not
so much narrowed anteriorly, the posterior angles directed more
laterally; anterior plate narrow, at least two and one-half times as
wide as long, its posterior margin regularly arcuate, the anterior
usually squarely truncate, sometimes rounded. Stridulating organ
occupying the same location on the hind femur as in S. fuscus ; plates
arranged in rows but the rows not as distinct. It is not possible to
assign a definite line of direction to these rows since the plates are
more or less equally spaced, and seem aligned in several directions.
Width of the row (measured from the free margin of a plate to the
free margin of a plate in the adjacent row) varying from 14 to 20
microns; variation in width of a row not correlated with age; free
margins of the plates smooth curves, far more strongly arcuate,
without a blunt tooth arising at the crest of the curve; not reduced
to a straight line by wear, each plate standing apart from all its
neighbors, with no contact laterally; the distance between plates
varying from 2 to 4 microns. Fine secondary punctuation of the
frons almost lacking in some specimens; in most with a fine sculp-
ture, the result of scratching by earth particles while digging. Length
of the basal antennal joints varying, the third, in some cases, being
subequal to the second and the fourth from one-fourth to one-third
as long as the third. In some specimens (especially males) the pos-
terior margin of the basal plate of the prothorax is rounded, but in
most cases it is squarely truncate, and in some specimens it may be
slightly emarginate. The anterior margin of the anterior plate is
also subject to some variation, but in all cases observed is at least
slightly truncate.
APRIL, 1926] DAVIS AND SMITH STENOPELMATUS 179
Maximum
Minimum
Mean
Length of tibise, 2
7.5 mm.
6.0 mm.
6.9 mm.
Length of tibiae, 6
8.1
7.8
8.0
Length of notum, $
■ 5.0
4.0
4.6
Length of notum, 6
4.5
4.5
4.5
Total length (dry), $
48.0
42.0
45.5
Total length (dry), 6
-.37.0
35.0
36.0
Holotype, 9, (No. 1875), Oakland, California, December,
1925, and allotype, 6 (No. 1876), Oakland, California, Decem-
ber, 1925, in the California Academy of Sciences. Three pairs
of paratypes from the same locality are to be deposited, one
pair in each of the three following museums : United States
National Museum, Philadelphia Academy of Natural Science,
and Entomological Department of the University of California.
These specimens were undoubtedly mature, since they suc-
cessfully copulated before they were killed. This species might
at first be taken for a form intermediate between 6'. fuscus and
S', pictus. According to the characters of Hebard’s table it
would come under S', longispina, the spination of the hind
tibi^ being the same as that given for that species. In size and
general appearance this species more closely resembles S', pictus,
except that the dark lines on the front are lacking.
Stenopelmatus pictus Scud. *
Heavy punctuation of the head practically lacking, the secondary
punctuation extremely fine and sparse. Frontal sutures much as in
intermedins, as is the rugosity above the fronto-clypeal suture. Basal
antennal joints about the same, except that the fourth is only one-
fourth as long as the third. Ovipositor of the female longer, less
stout, the angle formed by the joining of the apices of the blades
more acute. Basal plate of the prosternum somewhat arcuate pos-
teriorly, the lateral plates quite wide, their posterior angles rounded
and not laterally directed. Anterior plate very transverse, about seven
times as wide as long, fairly regularly arcuate posteriorly^ sinuately
truncate in front.
* Since these notes were prepared we have had an opportunity to
examine a specimen of S. pictus from Suisun, California, sent by Dr. Stanley
Freeborn, and a series of eleven specimens taken in lower Panoche
Canyon, Fresno County, California, by Mr. F. W. W''ymore and Dr. E. C.
Van Dyke.
The specimen from Suisun agrees closely with the characters stated
previously. Those from Panoche Canyon agree in all except the shape of
the anterior prosternal plate, which, in this series, varies from transverse
and very short to nearly triangular; the black markings of the head and
thorax, however, are very distinctive. This species on the whole is some-
what larger than S. intermedius . As these specimens were all dead when
received there has been no opportunity for observation of habits and
mating reactions.
180
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
Length of hind tibiae, ?, 6.5 mm.; length of notum, $, 4.0; total
length, 2,21.0.
There are no specimens available from which to make slides
of the stridulating plates, but it is to be expected that these
would resemble those of intermedius quite closely. The only
specimen of .S’, pictus available for examination at present is a
female in the collection of the California Academy of Sciences,
taken by E. P. Van Duzee in San Francisco, April 11, 1919.
This specimen was determined as pictus by Hebard in 1920 but
is marked “atypical.” The presence of the dark lines on the
head serve, however, to distinguish it immediately from
S. intermedius.
Preparation of Soft-Bodied Insects
In mounting such soft-bodied insects as Stenopelmatus (Orth-
optera) for the cabinet, it is found that great difficulty is expe-
rienced in preventing the abdomen from shriveling to such
an extent as to be useless for measurement or other identifica-
tion, Removing the viscera through a ventral longitudinal
suture and filling the abdomen with cotton has been tried, with
indifferent success. I have found that by injecting the speci-
men hypodermically with 95 per cent alcohol and then immers-
ing in alcohol of the same strength for a month or six weeks,
very satisfactory results may be obtained, the abdomen shrivel-
ing only to a very slight extent. Formalin or some other rap-
idly hardening agent might be used with about the same or a
little better success. — Alonzo C. Davis.
After a recent severe outbreak of the oak moth (Phryganidea
calif ornica) at Mill Valley, California, I placed a handful of the
chrysalids in a container. At one average inspection there were
five moths and fifty parasites (Ephialtes hehrensi Cress.) in the
box. Later some small Tachinids and a few Chalcu ovata
emerged. This would indicate a high rate of parasitism. — E. P.
Van Duzee.
APRIL, 1926 ]
DAVIS AND SMITH STENOPELMATUS
181
Fig. 1, Prosternum of Stenopelmatus fuscus; 2, same of intermedins ;
3, anterior prosternal plate of pictus; 4, stridulatory plates of inter-
medins; 5, same of fnscns; 6, frontal sutures of fuscns; same of
intermedins.
182
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
SOME NEW SIPHONAPTERA FROM CALIFORNIA
BY CARROLL FOX
Phalacropsylla cummingi Fox, n. sp.
Figs. I and II
Frons provided with two approximate rows of bristles, an anterior
of three weak bristles and a second row of four stout bristles of about
equal size, the innermost being placed at the edge of the antennal
groove above the vestigial ej'^e. Stiperimposed genal spines two, the
lower longer and broader (fig. 1). Labial palpi reaching almost to the
apex of fore coxae.
Hind coxae with a row of six spineform bristles (spinelets) on its
inner side. Outer side of hind femur with a row of four bristles
toward the apex and two or three toward the base, the inner side
being without bristles. Hind tibia with three rows of bristles in
addition to those placed near the fourth, fifth and sixth dorsal pairs
as in jP. paradisea, giving the appearance of having the bristles on
posterior border of tibia in groups of three. First segment of the
hind tarsus one-third longer than the second (6.4).
Process of clasper (fig. 2^ P) prominent and not divided by a ven-
tral suture; finger extending well above the process, its ventral border
convex and provided with thin bristles extending for nearly its entire
length; below the middle with a submarginal row of two or three
stouter bristles on the inner side (fig. 2, F).
Vertical arm of ninth sternite straighter and wider than in P. para-
disea and not divided at apex; horizontal arm with a row of bristles
on its distal half, the upper three of which are stout, the middle one
much the stoutest (fig. 2, IX st). Length, 1.8 mm.
Los Angeles, California, 1925, one male taken off
Diphodomys agilis.
Type No. 28918, U. S. N. M.
Leptopsylla ebrighti Fox, n. sp.
Figs. Ill and IV
This new species of Leptopsylla is quite distinct from the
other American forms.
Genal spines two; three of the bristles in the fronto-marginal row,
at the most prominent part of the frons, short, heavy and spine-like.
Labial palpi reaching a little beyond the middle of the fore coxze.
Pronotal comb containing about twenty-four spines. Mesonotum
about one-fourth longer than the metanotum in both sexes; the
bristles on the metathoracic epimerum eight in number arranged
APRIL, 1926]
FOX SIPHONAPTERA
183
2; 4: 2: in the male, and 3: 3: 2: in the female. Bristles in the pos-
terior row on abdominal tergites II to VI in the female number
15 to 17, in the male about 12. Abdominal sternites IV to VI, each
bearing a single row of 8 bristles in the female^ 4 or 5 in the male.
Seventh sternite in the female slightly incised ventrally, above the
incision with a broadly rounded lobe and carrying a row of 12
bristles; in the male with a row of from 4 to 6 bristles. Eighth tergite
in the male with 5 lateral bristles, in the female with a submarginal
row of bristles and two lateral rows, the anterior containing 4 small
and the posterior 6 large bristles (fig. 4). Abdominal sternite VIII
of the male without bristles ventrally, produced laterally into a
rounded lobe at the apex of which are 2 fairly long and 1 or 2 shorter
bristles (fig. 3, VIII st).
Style short and thick, wide at the base and at the middle, but
narrow at tip where there is a long bristle, and below this three
smaller bristles. Antepygidial bristles 4 (or 5) on each side in the
female (in the type specimen 4 on one side and 5 on the other), and
3 on each side in the male (in the type specimen 3 on on side and 2
on the other). Hind femur with some bristles basally and apically
but no lateral row; the hind tibia with a row of 8 bristles on the outer
side and 4 or 5 on the inner.
Process of the claspers broad at tip, reaching to about the top of
the finger (fig. 3, P) ; the finger almost straight on its dorsal border,
widest near the tip, the ventral border carrying three large and sev-
eral smaller bristles. The finger may be roughly described as having
the shape of a right-angle triangle with the apex at the pedicle and
the caudo-ventral corner broadly rounded; manubrium slender, acu-
minate and gently curved upward (fig. 3, F). Horizontal arm of the
ninth sternite slightly curved upward, its apex bluntly pointed, its
ventral edge convex and carrying six rather long and some smaller
bristles (fig. 3, IX st). Length: male, 1.8 mm.; female, 2.8 mm.
Los Angeles, California, July, 192*5, a series of males and
females obtained from Neotoma fuscipes.
Type No. 28919, U. S. N. M.
Anomiopsyllus californicus Baker 1904
Figs. V and IX
1904, Baker, Invert. Pacifica, I, p. 39.
1915, Rothschild, Ectoparasites, I, p. 45.
Baker’s description was based on one female from Spilogale
phenax taken at Claremont, Los Angeles County, California.
The writer has a series of both sexes from Neotoma fuscipes
and Spilogale pkenax taken at Los Angeles, California. The
male is reported herein for the first time.
184
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
Rothschild acquired the type specimen from Baker. Un-
fortunately, the receptaculum seminis in the type is almost
destroyed. Rothschild, comparing a series of Anomiopsylhts
niidatus which he had from Paradise, Arizona, with the type of
californicus came to the conclusion that calif ornicvus was the
same as nudatus. These fleas are certainly very much alike but
there are minor differences in the shape of the receptaculum
seminis (fig. 5), and discovery of the male of calif orniais shows
Rothschild to be in error when he pronounced californicus to
be a synonym of nudatus.
In both species there are three small spine-like bristles on the
finger of the claspers. In californicus these are arranged close
together in an oblique row near the apex of the finger (fig. 9,
F). In nudatus they are much more widely separated along the
posterior border of the finger, two of them above and one far
below.
Hoplopsyllus powers! Fox, n. sp.
Figs. VIII and XI
Labial palpi reaching to about two-thirds of the anterior coxse.
Pronotal comb composed of from 14 to 16 spines; the metepimerum
with 9 or 10 bristles in the male (5:4 or 6:4) and 12 or 14 in the
female (6:6 or 6:8). Hind femur carrying a row of about 6 bristles
on its inner side, the hind tibia a row of about 10 on its outer side;
one of the apical bristles of the second hind tarsal segment extends
beyond the middle of the fifth segment. Sternite VII in the female
has a row of from 14 to 15 bristles on the two sides together^ in the
male about 4. Sternite VIII in the male bears laterally a group of
about 7 bristles; in the female there is a lateral row of 5 (or 6), while
apically there is a row of 8 bristles on the outer side and a row of
7 or 8 on the inner side.
Style slightly more than twice as long as broad at base (9:4).
Receptaculum seminis distinctly flattened proximally (fig. 8) ; bristled
process of the claspers (fig. 11, P) about one-third longer than wide
at the middle (10:15), the toothed process five times as long as wide,
exclusive of the tooth (5:25) (fig. 11, P. 2). Length: male, 1,6 mm.;
female, 2.3 mm.
Los Angeles, California, 1925, taken on SyliAlagus bachmani.
Type No. 28920, U. S. N. M.
APRIL, 1926]
FOX SIPHONAPTERA
185
Hoplopsyllus minutus Fox, n. sp.
Figs. VI and X
A small flea. Only the female known.
Labial palpi extending to three-fourths of anterior coxae; maxillary
palpi only a little shorter than the labial palpi. Pronotum bearing a
comb of 13 spines; metepimerum with 7 bristles (3:4). Hind femur
with a row of S bristles on the inner side; hind tibia with a row of
8 or 9 bristles on the outer side; longest apical bristle of the second
hind tarsal segment extending to the tip of the fifth segment.
Abdominal sternites II to VI each with a row of about 4 bristles;
sternite VII carrying a row of 12 or 13 bristles; eighth tergite with
a marginal row of 9 or 10 bristles, a submarginal row of 8 and later-
ally a row of 5 bristles. Style about twice as long as broad at base,
carrying one apical and one subapical bristle (fig. 10). The head of
the receptaculum seminis almost round (fig. 6). Length, 1.3 mm.
San Francisco, California, 1908, taken on Sylvilagus hach-
mani.
Type No. 28922, U. S. N. M.
Explanation of Figures
1. Pkalacropsylla cummingi, frons showing the genal spines.
2. Pkalacropsylla cummingi, male clasping organs.
3. Leptopsylla ebrighti, male clasping organs.
4. Leptopsylla ebrighti, female, abdominal segments VII and VIII.
5. Anomiopsyllus calif ornicus, receptaculum seminis.
6. Hoplopsyllus minutus, receptaculum seminis.
7. Leptopsylla ebrighti, receptaculum seminis.
8. Hoplopsylla po'vjersi, receptaculum seminis.
9. Anomiopsyllus calif ornicus, male, ninth tergite.
10. Hoplopsyllus minutus, female, terminal segments of abdomen.
11. Hoplopsyllus ponjoersi, male, ninth tergite.
NOTE
On October 24, under a band on a large crabapple tree 32
codling moth cocoons were found, 22 of which were parasi-
tized by JEnoplex carpocapsce Cush. In the neighborhood of
this tree 32 of the 417 codling moth larvae collected were
parasitized.
In September a number of the chalcids, Dibrachys bouche-
anus Ratz., were reared from a codling moth larva. Appar-
ently, this parasite functions usually as a secondary, for in
December it was reared from the larvae of JEnoplex carpo-
capS(S Cush.
Ephialtes sanguineipes (Cr.) was reared from a codling moth
larva on August 8. — Stanley E. Flanders,
IIIA-
APRIL, 1926] BARNES-BENJAMIN PSEUDANARTA
187
ON THE PLACEMENT OF “PSEUDANARTA’^
FLAVIDENS GRT. (LEPID., PHALAENID2E)
BY WILLIAM BARNES AND F. H. BENJAMIN
Decatur, Illinois
POLIA ^ FLAVIDENS Grt.
1879, Grote, Bull. Geol. Surv. Terr., V, 205, Hadena (Pseudanarta),
1889, Smith, Ent. Amer., V, 179, Pseudanarta.
1893, Smith, Bull. U. S. N. M., XLIV, 149, Pseudanarta.
1903, Dyar, Bull. U. S. N. M., LII, 124, No. 1309, Pseudanarta.
1906, Hampson, Cat. Lep. Phal. B. M., VI, 195, pi. Cl, f 11, Pseuda-
narta.
1908, Smith, Jour. N. Y. Ent. Soc., XVI, 88, Pseudanarta.
Described from specimens from Colorado, Mead and Henry
Edwards. Beutenmueller, 1892, Bull. Am. Mus. Nat. Hist., IV,
does not list a type in the American Museum of Natural His-
tory. Hampson, 1906, figures the species and lists a female
type in the British Museum. We possess a specimen compared
with this type by Dr. J. McDunnough.
The species has been correctly determined in collections, but
all authors appear to have overlooked the fact that it has hairy
eyes and therefore does not belong in the Cuculliinse, but falls
into the Hadeninse. By Hampson’s keys, 1905, Cat. Lep. Phal.
B. M., V, it would run to the genus Trichestra Hamp., but
appears to be there out of place.
The genera of the Hadeninae need revision, so we tentatively
place the species with its allies secedens Wlk. and hasivirida
B. and McD. The three agree in possessing yellow basal areas
on their secondaries, and lashed eyes, but differ considerably
in characters of palpi and vestiture. It is probable that three
distinct genera will ultimately be erected for these three spe-
cies, but for the present they can rest in
1 Sense Hampson, 1905. We are cognisant that PoUa is a Tentamen
genus with type flavidncta, belonging to the subfamily Cuculliinse; merely
republished by Ochsenheimer, 1816, who credits to Hiibner. Hampson,
1907, Ann. Mag. Nat. Hist., (7), XIX, 252, uses Miselia, with type desig-
nated as conspersa, in place of PoUa. But Miselit! is also a Tentamen genus,
with type oxycaniha, belonging in the subfamily Cuculliinae; merely repub-
lished by Ochsenheimer, 1816, who credits to Hiibner. Like most European
workers, Hampson has refused to accept generic names unless they were
accompanied by a description in words, and not only rejects the Tenta-
men, but also the genera of Ochsenheimer. He, therefore, lists both Folu
and Miselia from Treitschke, 1825. Probably the correct generic name for
the bulk of the species placed by Hampson in PoUa, or Miselia, will be
Hadena Schr.
188
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
ON THE DISTRIBUTION OF OMPHALOCERA
OCCIDENTALIS B. AND BENJ. (LEPID.,
PYRALID^, PYRALIN^)
BY WILLIAM BARNES AND F, H, BENJAMIN
j Decatur, Illinois
Omphalocera occidentalis B. and Benj.
1924, Barnes and Benjamin, Contrib. N. H. Lep. N. A., V, (3), 191,
Omphalocera.
Described from a holotype 6 , Clark County, Nevada, 24-30
June, and 1 6 paratype, High Rolls, New Mexico.
A fine pair have been received from Mr. Tom Spalding, taken
at Eureka, Utah, July 4. The female is very similar to the
male in coloration and maculation.
A NEW SPECIES OF PLASTOCERID.E IN THE
GENUS EUTHYSANIUS (COLEOPTERA)
BY VASCO M. TANNER
Brigham Young University, Provo, Utah
In 1853 Dr. John L. LeConte established the genus Euthy-
sanius and described the species lautus as the type of the genus.
In 1866 he described the species pretiosus, which was the second
of the two species in this genus. These descriptions are brief
and rather inadequate because important morphological charac-
ters are ignored. The genitalia of these species, for example,
have valuable characters which may be utilized in separating
them. Some of the recent workers in coleopterology have failed
to base their descriptions upon definite morphological charac-
ters and as a result their descriptions are almost worthless. It
seems much safer to work with many morphological characters
than to split up groups and make new species on a few variable
characters.
Euthysanius blaisdelli Tanner, sp. nov.
(Figs. 2, 4, 7, 8 and 11)
Material studied : Three males, collected by the writer and
Mr. Angus M. Woodbury in Zion National Park, Utah, August,
1925.
Male. General color a rather light brown. Length, 17-19 mm.
Head broader than long; dorsal side pubescent and punctate; ventral
with a few small setae; maxillary palpi about half as long as the
head, with long setae at the distal end of each article; mandibles
APRIL, 1926]
TANNER EUTHYSANIUS
189
190
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
(fig. 7) rather large with numerous stout setse on the dorsal outer
margin; antennae (fig. 8) pectinate, very pubescent, reddish brown
throughout; pronotum almost square, except for the posterior outer
margins which extend back to the humeri (fig. 4) ; elytra about three
times as long as wide, almost parallel, bearing numerous small pale
setae; legs long and slender, of uniform chestnut brown color; abdo-
men long and slender, the ventral side densely covered with small
setae, segmentation clearly discernible, there being ten tergites and
ten sternites present; aedeagus leaving the body between the ninth
and tenth sternites; genitalia (figs. 2 and 11) much like that of E. pre-
tiosus Lee. (figs. 3 and 12).
In this species the ninth sternite is emarginate; in E. pretiosus
Lee., to which it is most closely allied, it is not emarginate.
The inner surface of the tegmen is very different in these two
species, as is also the basal piece. The genitalia (figs. 1 and 10)
and the antennae (fig. 9) of E. lautus Lee. are noticeably dis-
tinct from those of the other two species.
Type, male, (No. 21) in my collection. Paratype in the col-
lection of Dr. Frank E. Blaisdell, Sr. This species is named in
honor of my good friend Doctor Blaisdell, who has contributed
so much to my collection of Coleoptera.
Literature
LeConte, John L.
1853. Revision of the Elateridse of United States. Trans. Am.
Philos. Soc. (2), X, pp. 405-408.
1866. New Species of North American Coleoptera. Smiths.
Misc. Coll. VI, No. 167, 2d ed., pp. 1-177.
Sharp, D., and Muir, F.
1912. Comp. Anat. of Male Genital Tube in Coleoptera. Trans.
Ent. Soc. Bond., pp. 477-642, pi. 42-78.
Explanation of Figures
Fig. 1. Ventral view of the genitalia of Euthysanius lautus Lee.
Fig. 2. Same of E. blaisdelli sp. nov.
Fig. 3. Same of E. pretiosus Lee.
Fig. 4. Posterior outer margin of pronotum in E. blaisdelli.
Fig. 5. Same in E. pretiosus.
Fig. 6. Same in E. lautus.
Fig. 7. Mandible of E. blaisdelli.
Fig. 8. Last two segments of the antenna in E, blaisdelli.
Fig. 9. Same in E. lautus.
Fig. 10. Dorsal view of the genitalia of E. lautus.
Fig. 11. Same of E. blaisdelli.
Fig. 12. Same of E. pretiosus.
ml. — median lobe. 9ths. — ninth sternite.
t. — tegmen. lOths. — tenth sternite.
bp. — basal-piece. 9tht. — ninth tergite.
lOtht. — tenth tergite.
au. — anus.
APRIL, 1926]
FALL ALASKAN COLEOPTERA
191
A LIST OF THE COLEOPTERA TAKEN IN ALASKA
AND ADJACENT PARTS OF THE YUKON
TERRITORY IN THE SUMMER OF 1924
BY H. C. FALL
Tyngsboro, Massachusetts
(Continued from page 154)
Lepturoides (Campylus) variabilis Esch. Dawson; Eagle;
Fort Yukon; McKinley Park. Very common on grass and
bushes along the Yukon River and frequently seen flying
by day about the streets of Dawson.
Ludius (Corymbites) resplendens Esch. Skagway, a single
specimen beaten from spruce on a mountain trail above the
town.
Ludius (Corymbites) lobatus Esch. Seward, two examples,
one entirely dark, the other with the elytra ferruginous.
Ludius (Corymbites) morulus Lee. White Horse; Dawson.
"^Ludius (Corymbites) nigricoUis Lee. Skagway.
Ludius (Paranomus) costalis Payk. Dawson; Eagle.
"^Hemicrepidius ( Asaphes ) oregonus Lee. Ketchikan ; Wrangel.
Cryptohypnus abbreviatus Say. Dawson.
Cryptohypnus nocturnus Esch. White Horse; Skagway.
Cryptohypnus nocturnus var. lucidulus Mann. Skagway ; Mc-
Kinley Park; Seward.
Hypnoidus tumescens Lee. Skagway; Dawson.
Hypnoidus musculus Esch. Skagway.
*Hypnoidus extricatus Fall, n. sp;
Moderately elongate, subdepressed, black scarcely aenescent,
moderately shining, finely thinly cinereo-pubescent. Head quite
strongly densely punctate, vertex impressed. Antennae entirely
piceous, slender, the apical three joints passing the hind angles
of the thorax. Prothorax about one-fifth wider than long, nar-
rowed at apex ; widest at about the middle where the sides are
strongly arcuate, sinuate before the hind angles which are dis-
tinctly divergent and with a fine carina extending forward one-
third or two-fifths the thoracic length; surface numerously
but not very densely, finely, punctate, the punctures evidently
stronger and closer anteriorly but not in the least asperate;
192
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
median line narrowly smooth. Elytra three times as long as
thorax and at base slightly wider than the latter ; humeri obtuse,
sides nearly straight or feebly arcuate in basal two-thirds,
thence convergent and narrowly rounded at apex; striae fine
and entire, finely punctate, intervals flat, numerously finely
punctate. Body beneath finely sparsely punctate, the punctures
distant by about twice their own diameters as a rule, a little
coarser on the prosternum and toward the sides of the pro-
and meta-sterna, finest on the ventral segments. Legs piceous,
the tarsi and sometimes the coxae more or less pale. Length
(type male), 5 mm.; width, 1.8 mm.
Described from three examples, one each from McKinley
Park (type). Child’s Glacier and Seward, Alaska, all taken by
the writer. The Seward example is stouter than the others
and is doubtless a female. This species does not fit well into
any of the groups indicated by Horn. Because of its depressed
form it would by Horn’s table be likely to be referred to the
striatulus group, but the sculpture is much more in harmony
with the melsheimeri group, where I think it may for the
present be placed in the vicinity of tumescens. The size is a
little larger than in tumescens and the antennas are conspicu-
ously longer than in the latter or any of its near relatives.
Hypnoidns striatulus Lee. Skagway.
^Elater sp. near and perhaps the same as behrensi Horn.
Skagway.
FAMILY MELASID^
Epiphanis cornutus Esch. Seward.
FAMILY BUPRESTID.E
Dicerca tenebrica Kby. (prolongata Lee.). Yukon River
forty miles above Fort Yukon, on spruce wood pile; Ram-
part (Kusche) ; Dawson (Kusche). These specimens are
the same as an example compared by Saunders many years
ago with the types and kindly sent to me by Mr. Arrow
for examination. Kirby’s original specimens, so Mr.
Arrow writes me, are for some reason not now in the
Museum collection. Just what tenebrica is has long been
a matter of doubt with our coleopterists. LeConte asso-
APRIL, 1926]
FALL ALASKAN COLEOPTERA
193
dated it doubtfully with his lacustris (= lugubris $ ).
Crotch in 1873 surmised from the description that it might
be the same as prolongata and his surmise proves to be
correct. It stands now in the Leng List as a synonym of
divaricata. The placing of lugubris as a synonym of tene-
brosa in the Leng List is also an error, the former having
the intermediate tibiae of the male simple, whereas in tener
brosa they are toothed.
Dicerca tenebrosa Kby. Yukon River below Eagle; Dawson.
This also is determined with certainty by comparison wdth
one of Kirby’s original specimens.
Melanophila drummondi Kby. Yukon River, west of Fort
Yukon (Arctic Circle).
M elanophila acuminata DeG. Anchorage.
Chrysobothris trinervia Kby. Yukon River forty miles above
Fort Yukon.
Agrilus poUtus Say. Dawson; Fort Yukon (Kusche).
FAMILY HETEROCERID^
Heterocerus sp. Dawson.
FAMILY HELODIDiE
Cyphon variabilis Thungb. Scow Bay; White Horse. The
White Horse specimens are materially larger than those
from Scow Bay. It is possible that careful investigation
would show that one or both of these are distinct from
variabilis,
FAMILY DERMESTIDiE
'^Byturus unicolor Say. Skagway ; Dav/son.
Trogoderma sinistra Fall, n. sp.
Oblong-oval; similar in form to ornata but rather larger than the
average specimens of that species. Body entirely black without trace
of ornamentation, surface shining, pubescence short, subrecumbent,
blackish throughout. Head about two-fifths as wide as the prothorax,
rather coarsely and densely punctate, eyes feebly sinuate on inner
margin. Antennae (female) dull rufous, the first two joints blackish,
club evidently four-jointed but not very abruptly formed, its length
a little less than that of all the preceding joints. Prothorax about
three-fifths wider than long, sides broadly arcuate and rapidly con-
vergent from base to apex; punctuation finer than on the head, not
194
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
at all asperate, a little closer and coarser near the sides, the punc-
tures as a rule separated by about their own diameters. Elytra
slightly wider than, and about three times as long as, the prothorax,
similarly finely, not densely, punctate. Prosternum and sides of meta-
sternum more densely and coarsely punctate; abdomen finely punc-
tured. Legs blackish, tarsi paler. Length, 3.9 mm.; width, 2.15 mm.
Described from a single female taken by the writer at Daw-
son, Yukon Territory, June 26, 1924.
This is the first entirely black Trogoderma to be described
from our fauna. I have in my collection a pair (62) of
similarly black specimens from Aweme, Manitoba, which are
probably specifically identical, but of this I do not feel entirely
assured. They are considerably smaller (2.8-3.25 mm.) and
seemingly a little narrower, the antennae entirely rufous, the
third joint very small in the male.
Perimegatoma perversa Fall, n. sp.
Narrowly oblong, cylindrical, convex; black, antennae, tibiae and
tarsi ferruginous; pubescence short, stiff, inclined, entirely deep black;
surface feebly shining. Head two-fifths as wide as the thorax, the
latter moderately strongly transverse; sides arcuately narrowed from
the base, side margins narrowly refiexed. Elytra about three and a
half times as long as the thorax, sides broadly sinuate before the
middle. Head and thorax coarsely and densely variolate-punctate,
elytra a little less coarsely and densely so, the punctures subasperate
toward the base, especially at sides. Body beneath similarly densely
rather coarsely punctured, more rugosely so anteriorly. Antennal
club about four-fifths as long as the entire stem, basal two joints
moderately strongly transverse; apical joint about one-half longer
than the two preceding, subtriangular, the lower edge strongly
oblique. Length, 4.6 mm.; width, 1.8 mm.
Dawson, Yukon Territory; a single specimen. The antennal
club is rather of the male type, but I am not sure that the
specimen is a male.
This species, by good rights, should be either the cylindrica
of Kirby or Maiinerheim’s angularis, but in cylindrica the body
is said to be very minutely punctured and covered with decum-
bent cinerous hairs, and angularis is also said to be griseo-
pubescent and with condensed white hairs on the hind angles
of the thorax ; absolute statements which are quite at variance
with the present species.
APRIL, 1926]
FALL ALASKAN COLEOPTERA
195
FAMILY BYRRHID^
Pedilophorus subcamts Lee. White Horse.
Pedilophorus acuminatiis Mann. Sitka.
Simplocaria tessellata Lee. ( ? mefallica Sturm). Yukon River
forty miles below Fort Yukon ; Dawson.
Cytihis alternatiis Say. Skagway; Yukon River below Sel-
kirk; Dawson.
Byrrhus concolor Kirby. Skagway; Yukon River (Sheep
Creek below Eagle) ; Dawson; also on the Yukon below
Selkirk, Yukon Territory, where numerous speeimens were
pieked up on the boat while taking wood. Identifieation is
made after study of an example sent me by Mr. Arrow
who had compared it with Kirby's type.
Byrrhus sp. Certain specimens among those taken below Sel-
kirk are of average larger size and have the transverse
closed figure of the el3rtra defined by pale pubescence better
defined than in the preceding. These are possibly cyclo-
phorus Kby. Mr. Arrow writes me, however, that he is
by no means sure that the concolor and cyclophorus types
represent different species. Of cyclophorus he writes “has
the whole of the lower surface and legs light red, but as
the elytra are not quite black I think that very likely indi-
cates immaturity.” Of concolor (type) he remarks: “Has
very faint traces of two series of small spots placed as in
cyclophorus. It is very slightly smaller and relatively
shorter.”
''^'Byrrhus eximhis Lee. McKinley Park; Dawson ; Yukon River
below Selkirk, Yukon Territory.
Lioligla ( Simplocaria ) nitida Mots. Ketchikan.
FAMILY OSTOMID^
Calitys scaber Thunb. Dawson.
FAMILY NITIDULID^
Meligethes scevus Lee. Dawson.
Nitidula bipunctata L. Dawson. Several specimens having the
form and characteristic elytral spots of this common intro-
duced species, but blacker, legs entirely black or virtually
so, and punctuation rather finer and sparser than in my
examples of the latter.
196
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
Nitidula nigra Schf. Dawson ; McKinley Park. With the pre-
ceding occurred examples without the elytral spots. These
are undoubtedly the nigra of Schseffer. They do not differ
specifically from the spotted form, but whether this is
really a form of bipunctata or a distinct species I am some-
what in doubt.
'^Omosita discoidea Fabr. Skagway.
Epurcea terminalis Mann. Anchorage.
Epurcea truncatella Mann. Yukon River below Eagle ;
Dawson.
FAMILY CUCUJIDAI
Pediacus fuscus Er. White Horse ; Dawson ; Anchorage.
Dendrophagus cygncci Mann. Skagway.
FAMILY CRYPTOPHAGID^
Antherophagus suturalis Makl. Skagway; McKinley Park.
*Cryptophagus difficilis Csy. (LeConte MS). McKinley Park.
Crypto phagus sp. near the preceding. Eagle.
Henoticus serratns Gy 11. Anchorage.
Ccenoscelis ferruginea Sahlbg. Anchorage ; White Horse.
Atomaria (Agathengis) sp. Yukon River, Yukon Territory,
below Selkirk.
''^Atomaria (Agathengis) sp. Fort Yukon; Dawson.
Atomaria (Agathengis) vespertina Makl. Anchorage; White
Horse.
Atomaria (Anchicera) kamtschatika Mots. Yukon River
(Sheep Creek below Eagle); Anchorage; Skagway;
Homer (W. S. McAlpine).
Atomaria (Anchicera) fulvipennis Mann.? Skagway; Homer
(W. S. McAlpine).
'^'Atomaria (Anchicera) sp. Yukon River below Eagle.
^Atomaria (Anchicera) sp. Fort Yukon.
^AtomariO' (Anchicera) sp. McKinley Park.
Atomaria (Anchicera) sp. Dawson.
Atomaria (Anchicera) sp. Dawson. None of these unidenti-
fied species seem referable to any of those of Mannerheim
of Maklin.
FAMILY COLYDIIDAI
'^'Lasconotus borealis Horn. Yukon River below Eagle.
APRIL, 1926] FALL ALASKAN COLEOPTERA
197
FAMILY LATKRIDIID^
'‘^Lathridius breviclavus Fall. Yukon River below Eagle.
Lathridius cinnamopterus Mann. {=montanus Fall). Anchor-
age ; Fort Yukon. Specimens taken at Anchorage are with-
out hesitation referred to this hitherto unrecognized
species. I have little doubt that fulvipennis Mann, is iden-
tical. Renewed study at this time enables me to say that
the species that I described ^ under the name armatulus is
the true costicollis of LeConte, while my montamis as
indicated above seems not to differ from cinnamopterus as
now identified. This leaves the species which I had
previously supposed to be costicollis without a name and
for this the name carinifer is now proposed. Making these
changes in name the characters as given in my table apply,
except that I was in error in saying that the front thighs are
not toothed at base in the male of montamis. This tooth
is very small but detectable in the type on renewed study ;
it is a little better developed in an Alaskan specimen. As
now present in my collection the following localities are
represented: L. cinnamopterus Mann. Anchorage and
Fort Yukon, Alaska, Coeur d’Alene, Idaho; Leavenworth
Valley, Colorado; British Columbia (Caraboo District);
Eureka and Little River, northern California : L. carinifer
new name {costicollis Fall, nec. LeConte), Astoria, Ore-
gon ; British Columbia ; Park City, Utah ; Colorado
Springs, Colorado.
'-^Lathridius costicollis Lee. Anchorage.
Eniemus protensicollis Mann. Seward ; Skagway.
Eniemus consimilis Mann. McKinley Park.
^Corticaria varicolor Fall. Skagway ; White Horse.
'^Corticaria arctophila Fall, n. sp.
Fulvotestaceous, elytra more or less dark, piceotestaceous. Head
two-thirds as wide as the prothorax, sparsely punctate; eyes rather
large, moderately prominent; tempora about one-fourth the length of
the eye as view'ed from above. Antennae reaching the hind margin
of the thorax, joints all longer than wide. Prothorax large^ sub-
cordiform, a little wider than long; sides distinctly serrulate; disk a
little flattened along the median line, a shallow fovea at base; sur-
1 Trans. Am. Ent. Soc., 1899, p. 118.
198
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
face rather sparsely evenly punctate. Elytra about one-fifth wider
than the thorax, oblong-oval, three-fourths or four-fifths longer than
wide, striae feebly impressed, strial punctures very little larger than
those of the interspaces, the hairs springing from the latter more
bristling than those from the strial punctures. Beneath sparsely punc-
tate, the abdomen more finely so. Femora stout, a little more incras-
sate in the male. Prosternum as long before the coxae as the length
of the latter; middle coxae very narrowly separated; metasternum and
first ventral segment subequal in length.
Male. Front tibiae distinctly bent and mucronate at tip; last ven-
tral nearly as long as the two preceding, disk broadly concave, apex
broadly truncate.
Female. Front tibiae straight, unarmed at tip; last ventral as long
as the two preceding, disk flattened, apex broadly rounded.
Length, 2 to 2.3 mm.
McKinley Park Station, Alaska, June 9, 1924, one male
(type), two females. The specimens were beaten from an old
bear skin found at a deserted camp a short distance south of the
station.
This species by its characters would fall between rudis and
varicolor in my table (Trans. Am. Ent. Soc., xxvi, 1899, p.
148). The punctures of the elytral striae and interspaces are
scarcely more unequal than on rudis, but the form and facies
are more nearly that of varicolor. From both the present species
differs by its larger prothorax with sides only very feebly
serrulate, and by its much longer last ventral of the male.
*Corticaria cavicauda Fall, n. sp.
Brownish piceous, shining, pubescence fine, thin and recumbent.
Head three-fourths as wide as the prothorax, finely, sparsely punc-
tate; eyes moderate, tempora one-fourth the length of the eye. An-
tennas scarcely attaining the base of the thorax; joints 3-7 decreas-
ingly longer than wide, 8 and 9 each about as long as wide, 10 a
little transverse. Prothorax moderately transverse, feebly cordate,
widest just before the middle; sides rather coarsely denticulate; sur-
face convex, with a moderately deep posterior fovea; punctuation
rather coarse, the punctures separated bj'^ their own diameter or a
little less. Elytra oblong subovate, scarcely three-fourths longer than
wide and about one-third wider than the thorax; striae scarcely im-
pressed, punctures of the interspaces evidently finer than those of
the striae, all becoming finer toward the apex. Beneath very finely
sparsely punctate, the metasternurn more strongly so. Prosternum
slightly longer before the coxae than the length of the latter; middle
APRIL, 1926 ]
FALL ALASKAN COLEOPTERA
199
cox^ separated by about one-third their own width; legs moderate,
femora not distinctly incrassate.
Male. All the tibiae bent and mucronate at tip, the hind ones only
slightly so; fifth ventral with a large and deep transverse excavation
with acute margins.
Female. Tibiae straight and simple; fifth ventral a little longer
than the preceding, evenly convex, apex broadly arcuato-truncate. .
Length, 1.7 to 1.8 mm.
McKinley Park Station, July 9, 1924. Three examples
(1 6, 2$ $ ) taken with the preceding species from the same
dried bear’s skin. The male is taken as the type.
This species is most nearly allied to prionodera, with which
it agrees nearly in size and form ; it is, however, slightly stouter
and more convex, of darker color, with more approximate
middle coxae and much broader and larger apical ventral fovea,
which in prionodera is not acutely margined.
'^'Corticaria Columbia Fall.? McKinley Park; Fort Yukon
(Kusche).
'''"Corticarin dentigera Lee. Eagle; Dawson; White Horse.
Cor tic aria ferruginea Marsh. Anchorage; Dawson.
FAMILY COCCINELLID.®
Scymnus lacustris Lee. Dawson, one example, not quite typical.
Scymnus nanus Lee. Dawson ; White Horse.
'^Psyllobora 20-maculata Say. Tenana River.
Ncemia episcopalis Kby. Dawson.
Hippodamia 13-punctata L. Anchorage.
Hippodamia parenthesis Say. White Horse.
Coccinella transverso guttata Fabr. Dawson.
''^Coccinella 5 -not at a Kby. Skagway.
Anisocalvia 12-maculata Gebl. Eagle.
FAMILY TENEBRIONID^
P hello psis porcata Lee. Juneau; Skagway.
Sc aphid ema ceneolum Lec. Eagle.
Hypophloeus praetermissus Fall, n. sp.
This name is proposed for a species so closely resembling
parallelus in size, form, color and nearly all other characters as
to be easily confused with it. It differs from parallelus in
having the anterior thoracic angles more rounded and only
200
THE PAN-PACIFIC ENTOMOLOGIST [VOL. II, NO. 4
feebly prominent and the sides of the prothorax more narrowly
margined. In these two particulars it agrees nearly with H.
glaher, which species is, however, materially smaller, narrower
and of more southern habitat.
Several examples of the present species were taken on a
telephone pole adjacent to a spruce wood pile at Dawson,
Yukon Territory, June 25, one of which is taken as the type.
I have what seems to be the same species from Tyngsboro,
Lawrence and Framingham, Massachusetts. There is, of course,
a possibility that this may be an introduced species but I have
no means at hand of verifying this suspicion.
'‘'Upis ceramboides L. Anchorage; Dawson; Fort Yukon
(Kusche).
FAMILY MELANDRYID^
^'Tetratoma concolor Lee. Anchorage; Dawson.
Hallomemis punctulatus Lee. Anchorage ; Dav.'’Son.
FAMILY ANOBIIDiE
'-•'Hadrobregmus linearis Lee. McKinley Park; Yukon River
below Selkirk, Yukon Territory; Fort Yukon (Kusche):
^Microhregma emarginatnm Drift. McKinley Park; Yukon
River thirty miles above Fort Yukon on spruce wood pile.
'^Ccenocara scymnoides Lee. Eagle; Skagway.
FAMILY BOSTRICHID^
Stephana pachys (Dinoderiis) siibstriatus Payk. Yukon River,
Yukon Territory; also below Eagle.
FAMILY CISID.E
Cis biarmatus Mann. Sitka ; Kasaan. -
'^Octotemnus denndatus Csy. Skagrvay.
FAMILY SCARAB^ID^
^^gialia cylindrica Esch. Skagway.
SEgialia lacustris Lee. McKinley Park ; Eagle. As remarked
by Horn this may well be the exarata^ of Mannerheim,
though that author’s statement that the clypeus is emar-
ginate hardly applies to lacustris.
Aphodius aleutus Esch. Entirely black form at Juneau. The
form having the etytra with maculation of the pardalis-
leopardus type was common at Anchorage. I am quite
APRIL, 1926] FALL ALASKAN COLEOPTERA
201
unable to separate the eastern leopardus from this. Horn
places them in different groups, one being assumed to have
the hind tibiae fringed with equal, the other with unequal,
spinules. This is here a very illusory character. Strictly
interpreted the spinules are more or less unequal in both ;
more liberally interpreted they are comparatively equal in
both.
Aphodius pectoralis Lee. 'Eagle ; Dawson ; White Horse.
FAMILY LUCANID^
Platycems depressns Lee. Yukon River below Selkirk, Yukon
Territory.
FAMILY CERAMBYCID^
Asemiim atriim Esch. Juneau; Skagway.
Tetropium cinnamo ptemm Kby. Yukon River below Eagle.
Tetropium alaskanum Fall, n. sp.
Elongate, parallel, black, elytra brownish to fuscous, legs and an-
tennas ferruginous; femora sometimes darker. Head finely, sparsely
punctate with distinct median sulcus. Antennae about three-fifths the
length of the body in the male, shorter in the female, stout, grad-
ually attenuated, second joint three-fourths to four-fifths as long as
the third. Prothorax about one-sixth wider than long, widest and
strongly rounded at or very near the middle, apex distinctly wider
than the base, the former broadly sinuate, the latter truncate; disk
feebly or scarcely impressed along the middle; surface polished and
very finely remotely punctate, becoming more closely so at sides.
Elytra about two and one-half times as long as wide, about one-sixth
wider and nearly three and a half times as long as the prothorax;
sides parallel^ apex obtusely rounded, surface finely, densely punc-
tate and dull, rather densely clothed with short, fine, appressed fulvo-
cinereous pubescence. Body beneath shining, sparsely punctate; last
ventral of male subequal in length to the preceding, its apex broadly
truncate or sinuato-truncate; last ventral of female longer with evenly
rounded apex. Legs short; femora robust. Length, 6.3 to 10.5 mm.;
width, 1.3 to 3 mm.
The type is a male and is one of a series of specimens taken
by the writer on a spruce wood pile at Sheep Creek on the
Yukon River below Eagle, Alaska, July 1, 1924. The same
species was taken at Dawson, Yukon Territory, and there are
other Alaskan examples in my collection from Ruby (Kusche)
and Kowak River (Grinnell). The specimens from the last
named locality are referred to by me (Ent. News, 1900, p. 459)
202
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
as T. cinnamopterum, and were reported to have been “thawed
out of birch wood. Notwithstanding the indicated difference in
habits, the specimens are undoubtedly all of one and the same
species and cannot well be referred to any of our hitherto
described species, though by description evidently closely allied
to parvulum Csy. The latter seems to have a similarly polished
and sparsely punctate prothorax, which, however, is said to be
as wide as the elytra, a condition not obtaining in any one of
twenty examples before me. Parvulum moreover was described
from Indiana and it is improbable that it can be the same as
the Alaskan species.
^Criocephalus agrestis Kby. Fort Yukon and Rampart
(Kusche). The third joint of the hind tarsi is a little
shorter than is described of typical agrestis but I think this
is not of importance,
Leptalia macilenta Mann. Skagwa}^ ; Seward.
Pachyta liturata Kby. Juneau.
^Acmceops proteus Kby, Skagway; Yukon River from Eagle
to below Fort Yukon.
'^Acmceops pratensis Laich. Eagle; Dawson; Fort Yukon
(Kusche).
Leptura vexatrix Mann. Skagway ; Dawson, Fort Yukon
(Kusche). With a good series ranging from New England
to Alaska I am not yet able satisfactorily to separate this
from sexmaculata L., assuming our eastern specimens to
be of that species.
'^'Leptura san guinea Lee. McKinley Park.
Leptura suhargentata Yihy. form similis Kby. White Horse.
"^'Leptura tibialis Lee, Skagway, one female.
Merium proteus Kby. Yukon River below Eagle ; Ruby
(Kusche) ; Dawson.
Phymatodes dimidiatus Kby. Yukon River below Selkirk,
Yukon Territory.
^'Xylotrechiis undulatus Say. Yukon River from Selkirk to
below Fort Yukon.
*Neoclytus muricatulus Kby. Yukon River thirty miles above
Fort Yukon.
M onochamu'S scutellaris Say. Yukon River below Eagle.
APRIL, 1926] FALL ALASKAN COLEOPTERA
203
Acanthocinus obsoletus Oliv. Yukon River below Selkirk,
Yukon Territory.
Pogonocherus peniceilatus Lee. Yukon River below Selkirk,
Yukon Territory.
FAMILY CHRYSOMELID^
*Orsodachna atra Ahr. Eagle; Nenana.
Syneta carinata Mann. Skagway.
Zeugophora atra Fall, new name. White Horse, one example.
This is the black fomi with yellow legs and antenna that
Crotch, Horn and others have called abnormis Lee. The
true abnormis is an entirely yellow form with which kirbyi
Baly and reinecki Grote are synonymous. It is not yet
clear whether these variously colored fonus are really dis-
tinct species, so for the present the names are allowed to
have specific standing; I think, however, there can be no
doubt whatever of the synonomy given above.
Pachybrachys peccans Suffr. Dawson, a single female prob-
ably belongs here.
Adoxus obscurus L. Skagway.
^Adoxus obscurus var. villosus Schrank vitis auct.). Anchor-
age, Yukon River.
^Entomoscelis adonidis Pallas. Dawson; Nenana.
Lina laponica L. Skagway.
Lina sp. White Horse; Big Salmon, Yukon Territory. A
small form with uniformly black-bronzed elytra, the thorax
broadly margined at sides with yellow, the identity of
which is not yet clear to me.
Phytodecta americana Schf. Eagle; Dawson; Yukon River
below Selkirk.
Phytodecta notmani Schf. Eagle; Yukon River below Sel-
kirk. Of two examples taken at Eagle, probably from the
same poplar tree, one is americana and the other notmani.
Also of my two examples taken below Selkirk one is
americana, the other notmani!! Again there are in my
cabinet two examples from Mount Washington summit.
New Hampshire, bearing same date, one americana, the
other notmani. One example, Dawson, is americana. These
determinations are based on Schaeffe’s table in Jour. N. Y.
Ent. Soc., 1924, p. 140.
204
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
Phytodecta vidgatissima L. Dawson ; Yukon River, Yukon
Territory. These specimens seem quite identical with
eastern examples of vulgatissima and not with the variety
inter stitialis as described by Mannerheim.
DisonycJm triangularis .Sa.j. Dawson.
Haltica tombacina Mann. {= evict a Lee.). Skagway; An-
chorage; McKinley Park; Nenana; Dawson, Yukon River,
Yukon Territory. I have no hesitation in announcing the
above synonym. The ‘'‘tombacina Mann.” of Horn’s mono-
graph is quite another thing and as specimens of this in the
LeConte collection and perhaps elsewhere bear the unpub-
lished name cupreolus Lee. I would suggest that this name
be adopted for the species.
^'Crepidodera (Chalcoides) fulz/icornis Fab. Fairbanks. This is
the common and widely distributed species long known as
helixines L. in our lists (see Chittenden in Jour. N. Y.
Ent. Soc., June, 1925).
'^Crepidodera (Hippuriphila) modeeri L. Anchorage; Yukon
River below Eagle.
^'Choetocnema protensa Lee. McKinley Park.
"^'Phyllotreta vittata Fab. Nenana.
FAMILY mylaerid-f; (bruchid.f;)
*Mylabris (Bruchus) prolixus Fall, n. sp.
Narrowly elongate, feebly cuneiform, entirely black, finely sparsely
cinereo-pubescent; head and thorax dull, elytra moderately shining.
Antennae ( 6 ) slender, as long as the entire body, pectinate, second
joint very small, third with an acutely triangular process, shorter
than the joint itself; rami of following joints very slender, becoming
gradually longer, those of joints 6-10 fully twice as long as the joints
themselves. In the female the antennae are stouter, attaining the
apical third of the elytra, serrate from the fourth joint. Head except
the clypeus densely punctate; eyes deeply emarginate, strongly promi-
nent in the male, in which sex they are separated on the front by
about three-fourths the length of the basal joint of the antennae; in
the female a little smaller and distant by about the length of the
basal joint. Prothorax moderately transverse, trapezoidal, sides nearly
straight and strongly convergent throughout; a feeble post-median
sinuation; base with a short truncate median lobe, surface very
densely punctate and dull throughout. Elytra about seven-tenths
(6) or one-half (9) longer than wide; sides gradually and feebly
divergent; striae fine, finely punctate, all abbreviated at apex, the
APRIL, 1926] FALL ALASKAN COLEOPTERA
205
fifth and sixth shortest; intervals flat, not distinctly punctate but
with fine scratches and reticulations. Pygidium a little wider than
long, parabolically rounded at apex, impressed each side at about the
middle, finely but not very distinctly punctate. Body beneath closely
punctate anteriorly, becoming more sparsely and finely so on the
ventral segments. Legs very long and slender, slightly less slender
in the female; the hind tibiae in the latter sex with a very short acute
mucro at tip, which is almost completely obsolete in the male; hind
femora without teeth or denticles; basal joint of hind tarsi conspicu-
ously longer than the remainder in both sexes. Dimensions: male
(type), 2.2 x 1 mm.; female, 2.5 x 1.3 mm.
Described from four examples (1 6, 3$ 9 ) taken by the
writer in the sweepnet near McKinley Park Station, Alaska,
June 9-10, 1924.
This very remarkable species is not only notable as being the
first of its family to have been reported from Alaska, but it is
one of the most conspicuously distinct species in the North
American fauna. By its simple femora and pectinate antennae
it must be referred to Horn’s Group IV, from all the members
of which it differs signally in its narrow form and the long
antennae of the male.
FAMILY CURCULIONID^
*Apion alaskanum Fall, n. sp.
Moderately robust and ventricose; black, antennae, except the club
and one or two joints preceding, front tibiae and all the femora except
at tips, yellow; body above thinly^ finely cinereo-pubescent, scarcely
shining; meso- and meta-sternum at sides densely clothed with ap-
pressed white hairs. Beak ( 9 ) subequal in length to the head and
thorax, rather slender, thickened in basal third, which is distinctly
punctured and dull, finely sparsely punctulate and shining beyond
the antennal insertion. First joint of antenuje as long as the next
two, second reaching the eye. Front canaliculose; eyes prominent.
Prothorax very little wider than long, sides parallel in basal half,
thence oblique and nearly straight to apex; basal margin not at all
expanded, surface densely, rather coarsely, punctate; median line
impressed in basal third. Elytra widest at apical two-fifths, twice as
wide as the thorax; humeri moderate, sides divergent and nearly
straight from base to middle; interspaces flat, nearly twice as wide
as the striae at the middle, sub-biseriately punctate. Beneath rather
coarsely and densely punctate. Length (excluding beak), 2.25 mm.;
width, 1.2 mm.
Described from a single female specimen taken by the writer
on the Yukon River, Alaska, thirty miles above Fort Yukon.
This species belongs to my group IV, and by the table is to
206
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
be placed near segnipes, with which it nearly agrees in the color
of the antennae and legs. The latter species differs in many
respects, notably in its more conical thorax with expanded basal
margin, parallel elytra, much longer beak, smaller, not promi-
nent eyes and punctate rather than channeled front. Only one
species of Apion ( cuprascens Mann, from Sitka) has hitherto
been reported from Alaska.
'^Sitona tibialis Hbst. Skagway.
Lophalophus inquinatus Mann. Skagway.
Lepidophorus lineaticollis Kby. McKinley Park ; Eagle ; Daw-
son; White Horse.
Lepyrus capucinus Schall. or possibly errans Csy. Fort Yukon ;
Nenana; Dawson.
Hyperodes maculicollis Kby. Yukon River below Fort Yukon
(Arctic Circle). This is the vitticollis (vittaticollis) of
Dietz’ paper, but there can be very little doubt that the true
vitticollis is the ulkei of Dietz.
Paraplinthus carinatus Boh. Ketchikan ; Kasaan.
Paraplinthus scrohiculatus Mann. Ketchikan.
Dorytomus longulus Lee. Skagway.
Dorytomus rufulus Mann. Anchorage.
Grypidius equiseti Fabr. Dawson.
N Otaris ccthiops Fabr. Skagway; Anchorage; Yukon River
below Selkirk, Yukon Territory.
Proc torus decipiens Lee. Dawson. Le Conte is in error in say-
ing that the thighs are not toothed in P. armatus and that
the venter is unarmed in the male of decipiens. In the
latter the last ventral is similarly but less strongly modi-
fied, lacking, however, the small tooth at the middle of the
anterior margin of the segment, present in armatus.
*Elleschus borealis Carr. Dawson; Fort Yukon.
*Orchestes parvicollis Lee. Dawson; Fort Yukon.
Acalyptiis carpini Hbst. Dawson.
"^Ceutorhynchus rapee Gyll. Nenana.
Ceutorhynchus decipiens Lee. Dawson.
Ceutorhynchus cyanipennis Germ. Dawson.
Ceutorhynchus pusio Mann. Anchorage ; Dawson.
*Ceutorhynchus fulvotertius Fall, n. sp.
Oval, piceous; apical third of elytra, legs and antennae reddish
APRIL, 1926]
FALL ALASKAN COLEOPTERA
207
brown. Above thinly clothed with slender linear yellowish gray
scale-like hairs, with scattered broader white scales on the elytra
which show some tendency to a serial arrangement; the sides and
median line of the prothorax and an elongate scutellar spot more
densely scaly; beneath clothed with larger oval, more or less pointed
gray scales which are densely placed anteriorly, a little sparser and
more oblong in shape on the abdomen. Head densely punctured;
beak slender, striate and finely punctate; antennal funicle seven-
jointed, formed as in pusio. Prothorax wider than long, much nar-
rowed in front, lateral tubercles small but acute, median line feebly
channeled, apical margin nearly straight between the broad and very
feeble angulations, disk very densely punctate throughout. Elytra
one-fifth longer than wide; humeri moderate, sides more or less evi-
dently convergent from the humeri; intervals rather wide, nearly flat,
subrugose, each with a series of acute tubercles on the declivity.
Legs slender, femora unarmed, claws simple. In the male the middle
and hind femora are unguiculate and the last ventral segment is
deeply transversely foveate; in the female the tibiae are unarmed, the
last ventral similarly but much less strongly impressed. Length
(type), 2 mm,; width, 1.15' mm.
Described from a series of specimens of very uniform size,
taken at Nenana, Alaska, July 6, 1924, and a single specimen
from Fort Yukon, July 2. The type is a male from Nenana.
«
This species greatly resembles pusio Mann., which occurred
plentifully at Anchorage. In the latter the claws have a very
small basal tooth (which may easily be overlooked), the scales
of the elytra are more or less aggregated in small spots, giving
a mottled aspect. Fulvotertius must be referred to the squama-
tus group of Dietz, and by its characters seems to fall next to
squamattis , which, however, it does not much resemble.
Pelenomus gracilipes Dtz. Dawson.
FAMILY SCOLYTID^
P olygraphus rufipennis Kby. Dawson ; McKinley Park.
Dendroctonus borealis Hopk. Dawson; Yukon River below
Eagle (Sheep Creek) ; Anchorage.
Hylurgops rugipennis Mann. Ketchikan. I cannot see any
difference between this and many eastern pinifex.
Hylastes nigrinus Skagway.
Hylastes yukonis Fall, n. sp.
Strikingly similar in size, form and general appearance to H.
nigrinus, from which it seems to differ constantly in its somewhat
less coarse sculpture both above and beneath. In nigrinus the punc-
208
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
tures of the discal striae of the elytra are notably coarse, the intervals
not appreciably wider than the striae. In the present species the strial
punctures are evidently smaller, the intervals between the striae
always distinctly wider than the punctures themselves. The sides of
the prothorax are slightly more rounded than in nigrinus, and the ros-
tral Carina is always sharply defined and entire. In my series of
nigrinus the rostral carina is never so well developed as in yukonis
and not infrequently is nearly or quite obsolete except for the short
distance where it crosses the apical convexity. Length, 4.3 to 5.6
mm.; width, 1.55 to 1.85 mm. Dimensions of type, 1.7 x 4.6 mm.; of
prothorax, 1.5 x 1.3 mm.
Described from six examples taken in flight at White Horse
(June 21, 1924), and Dawson (June 24, 1924), Yukon Terri-
tory. The type is a male from White Horse.
In the male the last ventral is apically concave and furnished
with a brush of yellowish erect hairs as in the allied species.
Notwithstanding the rugose sculpture the integuments of the
upper surface are polished and shining throughout, without
trace of alutaceous sculpture. Nigrinus is a little duller though
not visibly alutaceous ; the nearly allied macer, on the other hand,
has the surface between the punctures distinctly alutaceous.
A pair of the present species was submitted to Dr. J. M.
Swaine, who pronounced them a new species most nearly allied
to nigrinus.
Pityophthorus nitidulus Mann. Seward.
'^Pityophthorus borealis Swn. Fort Yukon; Dawson. My
thanks are due to Dr. J. M. Swaine for determining this
species ; he writes that the specimen sent him is perfectly
typical. It is described from the Coppermine River, Arctic
Canada (Can. Ent., 1924, p. 195).
Pityogenes carinulatus Lee. White Horse.
Ips perturhatus Eich. Yukon River below Eagle ; White
Horse; Dawson. Doctor Swaine has kindly verified my
determination of this species.
Ips interruptus Mann. ? Anchorage.
Orthotomicus vicinus Lee. Yukon River thirty miles above
Fort Yukon; Dawson.
Dryoccefes septentrionis Mann. Fairbanks ; Seward ; Anchor-
age; Yukon and Tenana rivers.
Dryoccefes affaber Mann. Dawson; Yukon River below Sel-
kirk, Yukon Territory.
APRIL, 1926]
BEAMER GRIBURIUS
209
NOTES ON GRIBURIUS MONTEZUMA (SUFFRIAN)
(COLEOPTERA-CHRYSOMELID^)
BY R. H. BEAMER
Department of Entomology, University of Kansas,
Lawrence
The University of Kansas Biological Survey party on July 5,
1923, accompanied by Mr. Warren Knaus of McPherson, Kan-
sas, visited the salt-marshes and surrounding hills in Kiowa
County, Kansas. On the roof of a cave in a rocky bluff a large
series of Griburius montezuma (Suffrian) was taken. The cave
extended back into the cliff perhaps eight feet, and was two feet
in diameter at the smallest place. Quite a large number of sticks
and much debris was piled up at the farther end, upon which
were two young buzzards. The stench which accompanies a
buzzard’s nest was not lacking. The specimens of Griburius
were found mating in large numbers on the roof of this cave.
Some were also observed flying in and out.
A small den with two openings and a stick nest at the back
of it was also a favorite place with the beetles. This second
place was perhaps ten feet from the first, along the same cliff.
As this particular species was considered a good find by Mr.
Knaus, a good deal of interest was taken in the actions of the
individuals and their seeming preference of location. We then
searched all the caves, shady cliffs, crevices, and all other places
in that immediate vicinity, but not a single specimen could we
find. On July 12, 1923, however, in Ellsworth County, Kansas,
perhaps seventy-five miles north and fifty miles east of the
first locality, another buzzard’s nest cave was found and here
four more specimens of this same species were taken. This
called forth a further search of all the caves in the vicinity,
but no more specimens were found.
In the rather hasty examination of the debris about the nests
no larval cases or other indications of the insect’s life history
were found at the time of our first visit. This preference, how-
ever, for a very definite location for mating and the knowledge
that some closely related forms are scavengers in ants’ nests
led us to suspect that this beetle did pass its early stages in the
210
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
refuse of rodent or buzzard nests. It was with a great deal of
interest, therefore, that we made our second visit to these same
caves in Kiowa County, Kansas, on April 12, 1925. We were
rewarded by finding both the living larvae in their cases and
the old cast-skins of previous generations.
The sandstone in which the caves occur is friable and the
walls about the entrance have been honeycombed in the process
of erosion. It was in these small holes and under stones at the
mouth of the caves that the larval cases were found. About
three dozen cases were collected. The beetles emerged in due
time, but the attempt to obtain the life history failed, for the
specimens all died before laying eggs, although two pairs were
observed to mate in captivit^c The evidence is conclusive that
they overwinter as larvae, and is suggestive of but one genera-
tion a year. Thanks are due to Mr. H. C. Fall of Tyiigsboro,
Massachusetts, for identification of the species.
A Luminous Zarhipis (Coleoptera)
On April 3, 1926, an adult male of Zarhipis riversi Horn (?)
was examined with a view to finding out whether or not it pos-
sessed the power of luminosity. The insect was taken into a
dark room and, when excited by shaking and moving about, was
observed to be distinctly luminous. So far as could be deter-
mined in the dark this luminosity involved the posterior and
lateral borders of the first, second and third abdominal segments
dorsally, and the sides, apparently just back of the spiracles,
of the remaining segments. The light was greenish yellow and,
though faint, could be seen at a distance of about two feet. The
insect seems to have at least partial control over this power, as
it was luminous only when disturbed and then shone steadily
for some time. Later in the day the insect failed to show any
light, no matter how much it was agitated.
Another male taken on May 3 refused to become luminous
at first, but after several trials over a period of about four hours
it shone very feebly for a short time. The females of this
genus are distinctly luminous and apparently do not have any
control over the light, as they glow steadily. — A. C. Davis.
APRIL, 1926]
ESSIG BUTTERFLY MIGRATION
211
A BUTTERFLY MIGRATION
BY E. O. ESSIG
A true migration of the thistle butterfly, Vanessa cardui
(Linn.), occurred in California from the middle to the last of
March, 1926. Large numbers were first noticed in southern
California about March 10. They were rapidly traveling north
and entered the southern end of the San Joaquin Valley on
the 12th and 14th. On the 17th they were in the Sacramento
Valley, and a portion of the flock passed through Berkeley on
the 23rd, 24th and 25th. In the latter place all were traveling
due north. In the orchard districts the butterflies were attracted
to the opening blossoms of almonds, apricots, cherries, plums,
prunes and pears, all of which had an unusually heavy bloom
this year. Many of the farmers were greatly annoyed by their
presence. In the interior valleys the butterflies were hovering
over the plants commonly called “woolly breeches” {Amsinckia
spp.), which is a favorite food plant of the caterpillars in the
West.
A full explanation of the periodic migrations of this and a
closely related species, Vanessa carye (Hiibner), has not yet
been made, but they are somehow connected with years having
mild, dry winters. Temperature is undoubtedly an important
factor, and the continued hot weather during the month of
March in California, Arizona, and northern Mexico may have
started the species north.
These migrations are always toward the coast or in a north-
erly direction, and no return migrations have been noted. Many
of the butterflies were captured, and their badly battered wings
gave evidence of a long flight. They move rapidly and often
cover from 100 to 200 miles per day. They rest at night and
the maximum flight occurs from 2 to 3 p. m. on bright, hot days.
On cold days the forward movements cease, so that the limits
of the flights are probably also determined by temperature.
Note by the Editor — Mr. W. S. Wright of the San Diego Museum
of Natural History writes me that this migration was observed
throughout southern California from Yuma to San Diego, beginning
March 13, the general direction of the flight being from southwest
to northeast, and the “peak” of the flight lasting about 40 minutes.
212
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
The butterflies were seen feeding at Encelia flowers. Mr. Wright
noticed that the flight followed two days of strong wind from off the
desert, and suggests that probably the butterflies were blown from
the desert toward the west until they came into contact with the cool
air from the ocean and were then deflected toward the northeast.
Mr. Wright noted that the flight of two years ago was also preceded
by a strong wind from off the desert.
A NEW NOMADA
BY C. L. FOX
San Francisco, California
Nomada (Gnathias) klamathensis, C. L. Fox, n. sp.
Aspect of a black and yellow Heminomada, but with the
bidentate mandibles of a Gnathias. Length, 8 mm.
Male. Robust. Face broad; orbits slightly converging below.
Scape swollen; joint three of flagellum equal to four. Pubescence
on head and thorax white, sparse, longer on sides and underneath.
Punctures on head and thorax dense, coarser on vertex. Mesoscu-
tellum low, feebly and broadly sub-bilobate. Abdomen short and
broad; apical plate narrowed, notched. Wings almost clear, slightly
darkened apically; nervures and stigma light ferruginous; second sub-
marginal cell narrowed above, three-quarters of width below, receiv-
ing first recurrent nervure far beyond the middle; third submarginal
cell much narrowed above; basal nervure well basad of transverso-
medial nervure.
Color black with the following yellow maculations: mandibles
except apices; labrum; clypeus except emarginated medial upper
part; lateral face marks, broad-triangular below, upper portion nar-
row, not quite reaching antennal basal line; scape in front; posterior
dorsal border of prothorax, interrupted medially; conspicuous spot
on lower anterior part of mesopleura; large, nearly coalesced spots,
on lobes of mesoscutellum; tubercles except somewhat anterior!}";
tegulas; anterior and intermediate femora in front and apex behind;
posterior femora with apical front and hind part only; tibiae and
tarsi except black stripe behind on posterior metatarsi; broad bands
on tergites 1-6, that on tergite 1 interrupted medially; broad band
on sternite 2 (possibly on the others if uncontracted) and greater
part of the ultimate sternite. Flagellum light red-brown, joints 1-3
blackened behind; apical margins of tergites and sternites somewhat
dark ferruginous, lighter on posterior ones. Described from one male.
Type, male, No. 1877, Mus. Calif. Acad. Sci. Collected by
myself at Klamath Falls, Oregon, May 12, 1924.
APRIL, 1926]
DOANE MEALYBUGS
213
CONTROLLING MEALYBUGS ON ORNAMENTAL
PLANTS
BY R. W. DOANE
Stanford University
During the past few years the common mealybug, Pseudo-
coccus gahani, has been becoming more and more abundant
and destructive on the ornamental plants on the Stanford
Campus. Efforts were made to control it with the usual spray-
ing materials, such as some of the miscible oils, Blackleaf 40,
and soap solutions, but none of these seemed to do the work
satisfactorily. It seemed almost impossible to get the oils to
penetrate the cottony secretion that often covered the masses
of these insects when they were abundant on any particular
plant, and even when the insects could be hit individually with
the spray, they were not always killed. The same factors
seemed to keep the nicotine from giving a good killing. It
was found that the soap mixtures would dissolve this mealy
covering but that they would only kill a part of the insects.
After a series of experiments in which these different sprays,
used singly and in combination, were tried out, we finally hit
upon a combination which has given us very satisfactory results
here on the campus. I have not had an opportunity to make
any extensive tests to determine the most economical combina-
tion of these materials, but as soon as I found a combination
that would do the work I began using that as I was more
concerned in getting results than in keeping down the cost. It
is quite possible that less nicotine or less soap might be used
with equally good results. The combination that I now use is
as follows :
1 gallon Volk Oil, formerly known as “orange oil.”
lbs. Gold Dust Washing Powder,
pint Nicotine (Blackleaf 40).
50 gallons Water.
It seems that the efficacy of this combination depends upon
the fact that the soap solution will cut the secretions which are
covering the insects and the nicotine will then be able to reach
and destroy the insects, and the oil helps to make the material
penetrating. It has been used successfully on many kinds of
214
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
ornamental plants, particularly Choisya, Oregon Grape and
Pittosporum. In cases of very bad infestations we have some-
times found it necessary to make a second application before
the plants were thoroughly cleaned. One application on a badly
infested lemon tree gave very good results.
Some of the large Araucaria on the campus were very badly
infested with the golden mealybug, Pseiidococciis aurilanaHis.
At a time when many of the young mealybugs were wandering
over the leaves these trees were thoroughly sprayed with this
same material, using a spray-gun with a pressure of 250 to 300
pounds. This greatly improved the appearance of the trees
and killed most of the mealybugs, but another treatment was
necessary some weeks later as it was found that all of the
insects had not been reached at the time of the first spraying.
One of the great advantages of using this spray is that it
removes from the tree the sooty mold that grov/s in the
secretion of the mealybugs. This, of course, makes the trees
much brighter and cleaner.
AN ENTOMOLOGICAL CODE
The Entomological Society of London has sent out a sug-
gested entomological code compiled by the British National
Committee on Entomological Nomenclature. The adoption of
such a code, especially if approved by the International Com-
mittee on Zoological Nomenclature, would be a great help
toward unifying the nomenclature of our science.
Most of the thirty articles in the proposed code will doubt-
less meet with the hearty approval of American entomologists,
but one or two might with advantage be slightly modified.
Article 14, for instance, provides that names of a lower rank
than that of a subspecies shall have no status in nomenclature.
The term “variety” is not even mentioned in this code, but it
has been, and still is, largely used by certain entomologists, and
we are left in doubt as to its standing under the proposed code.
Frequently it is used in the sense of subspecies and in any
case cannot be ignored. For many years such names must be
APRIL, 1926] VAN DUZEE ENTOMOLOGICAL CODE
215
given subspecific rank or our nomenclature would soon be in
a condition of chaos.
-Article 20 provides that the law of priority obtains when any
stage in the life history of an insect is named before the adult,
a provision that opens the road to abuses that might easily prove
intolerable. Article 23 provides that if a genus is established
without the mention of a species the first reviser has the right
to select any species as the type, provided it agrees with the
original definition of the genus. Most American students would
accept this but would credit the genus to such reviser, a proce-
dure not mentioned but probably intended by this code. So long
as we employ genotypes for the final delimitation of our genera,
it is illogical to accept any generic concept that contains no
species that can be used as its type. If we are to accept genera
on their described characters alone, the naming of a genotype
is a superfluity, and that way lies chaos. With such a rule there
would be nothing to prevent any adventurer from naming up
a series of genera by assembling imaginary characters and
awaiting the discovery of some insect that would fit into his
concept. The correct naming of genotypes is probably the most
perplexing problem in nomenclature. There are cases such as
Myodocha Latr. where it is quite impossible to name a geno-
type without violating some rule of our zoological codes. In
such cases we must use common sense and possibly depend
upon later validation by some authoritative committee.
The constant splitting up of genera is a form of amusement
quite popular with a certain class of our entomologists. In
many cases these minute subdivisions, of both genera and spe-
cies, are but minor offshoots in a still plastic complex, that soon
will become extinct in the natural course of evolution, but for
the present they may be treated as subgenera or subspecies, and
thus serve a useful purpose in classification and at the same
time be ignored by the worker in economic entomology. It is
conceivable that to the student of evolution a few hundred
years hence the description of such minute subdivisions may
prove of the greatest value, while to us they are but a weari-
• ness. It is for us to select what we wish to use and leave the
rest to posterity. — E. P. Van Duzee.
THE PAN-PACIFIC ENTOMOLOGIST
Published quarterly by the Pacific Coast Entomological Society
in co-operation with the California Academy of Sciences
E. P. Van Duzee, Editor F. E. Blaisdell, M. D., Treasurer
The present number of the Pan-Pacific Entomologist closes
its second volume. The subscription list has shown a steady
growth during the year, and the third volume will start under
much improved conditions financially. For the completion of
this volume there was still a considerable deficit to be made up.
Members of the Pacific Coast Entomological Society have con-
tributed a part of the necessary funds, but the bulk of the deficit
was covered by a very generous check received from Mr. B.
Preston Clark and an emergency appropriation from the funds
of the Department of Entomology of the California Academy
of Science. Twenty- four extra pages, issued with Vol. II, to
cover two papers of unusual length, were paid for by the
authors and other friends of the journal. The numbers of this
volume have been mailed as follows: No. 1, August 27, 1925;
No. 2, January 16, 1926; No. 3, May 8, 1926; and No. 4,
July 14, 1926.
Among recent visitors at the entomological department of
the California Academy of Science may be mentioned Mr. M. C.
Van Duzee of Buffalo, New York; Dr. Carl J. Drake of the
Iowa Agricultural College; Mr. P. H. Timberlake of the Citrus
Experiment Station at Riverside, California; Mr. Otto Swezey
of the Hawaii Sugar Planters’ Association; Mr. G. M. List of
the Colorado Agricultural College ; and Prof. T. D. A. Cock-
erell of the University of Colorado. Mr, Van Duzee spent
several weeks working over certain families of the Diptera in
the Academy collection that were in need of revision ; Doctor
Drake studied our Tingids, Anthocorids and Gerrids, and Mr.
List our Cimicids.
Dr. and Mrs. E. C. Van Dyke left June 3 for two months’
collecting in the mountains of Colorado. Mr. C. L. Fox has
gone to England for a visit at his former home at Plymouth,
returning in October. He will do some collecting for the
Academy of Science while there.
INDEX TO VOLUME II
217
Acinopterus aridellus Osb., 22.
xA.crididas at light, 36.
.^noplex carpocapsas Curt., 185.
Agabus gelidus Fall, 142.
kenaiensis Fall, 141.
Agrotis scaramangoides B. and
B., 107.
Alaskan Coleoptera, 127, 191.
Amara consueta Fall, 135.
Amblyteles caeruleus Cr., 158.
zebratus Cr., 158.
Anilloferonia V. D., 115.
testacea, V. D., 116.
Anorthodes Sm., 19.
Anomiopsyllus californicus Bak.,
183.
Apanteles mimoristas Msb., 10.
Apelinus perpallidus Gah., 158.
Apion alaskanum Fall, 205.
Asphondylia opuntise Felt, 103.
Athetis Hbn., 19,
indigena B. and B., 20.
Atypha Hbn., 19.
Barnes, William, personals, 160.
Barnes and Benjamin, papers by,
12, 16, 63, 64, 81, 106, 111,
187, 188.
Beamer, R. H., paper by, 209.
Bembidion caerulescens V. D., 65.
concurrens, Fall, 134.
intermedium Kby., 133.
mckinleyi Fall, 132.
picipes Kby., 133.
rupicola Kby., 133.
utahensis V. D., 66.
yukonum Fall, 131.
Basinger’s Alaskan Diptera, 47.
Blaisdell, F. E., papers by, 77, 90.
Blaisdell, personals, 47, 160.
Blaisdell collection of Coleoptera,
84.
Brachymeria pedalis Cress., 4.
Cantharis mandibularis Kby., 154.
Carposapsa simsoni Busk, 92.
Casey, T. L., necrology, 90.
Celiptera valina Schs., 21.
Centipeds of Utah, 55.
Ceutorhynchus fulvotertius Fall,
206.
Chalcis pedalis Cress., 10.
Chamberlin, J. C., paper by, 85.
Chamberlin, R. V., paper by, 55.
Chamberlin, W. J., paper by, 23.
Chaoborus lacustris Frbn., 161.
Clark, B. P., personals, 216.
Cockerell, T. D. A., personals,
216.
Coelambus borealis Fall, 137.
Coleoptera of Alaska, 127, 191.
Colpodes rufiventris V. D., 120.
Corticaria arctophila Fall, 197.
cavicauda Fall, 198.
Crickets, abundance of, 92.
Cryptoternies hermsi Kby., 53.
Cymatodera decipiens Fall, 126.
Dactylopius confusus Ckl., 102.
tomentosus Lam., 100.
Davis, A. C., notes by, 126, 180,
210 .
Davis and Smith, paper by, 174.
Decaturia pectinalis B. and M.,
64.
DeOng, E. R., note by, 46.
Dibrachys boucheanus Rtz., 185.
Dicerca tenebrica Kby., 192.
Dicheirus blaisdelli, V. D., 125.
Diocalandra taitensis Guer., 51,
Doane, R. W., paper by, 213.
Dodge, Ralph E., necrology, 48.
Drake, C. J., personals, 216.
Dyschirius secretus Fall, 130.
Earwig, 45.
Eiphosoma bruesi Csh., 100.
Eleodes, list of, 77.
fenyesi Blaisd., 77.
sierra Blaisd., 78.
Engelhardt, G. P., personals, 48.
Entomological code, 214.
Ephestia elutella Hbn., 46.
Ephialtes behrensi Cress., 180,
sanguineipes Cress., 185.
Essig, E. O., notes by, 45, 92.
218
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
Essig., E. O., paper by, 211.
Ethopolys parowanus Chnib., 57.
Euconnus longiceps Fall, 144.
Eupolia Sm., 17.
Euthysanius blaisdelli Tan., 188.
Evarthrus alabamse V. D., 118.
lodingi V. D., 118.
montanus V. D., 116.
Fall, paper by, 127, 191.
Fanning Islands, 49.
Ferris, G. F., personals, 47.
Flanders, S. E., paper by, 157.
Flanders, notes by, 92, 157, 185.
Forficula auricularia L., 45.
Fox, Carroll, paper by, 182.
Fox, C. L., paper by, 212.
Fox, C. L., personals, 47, 216.
Freeborn, S. R., paper by, 161.
Frontina aletiae Riley, 10.
Gallegos, J. M., necrology, 96.
Ganocapsus filiformis V. D., 22.
Garganus splendidus Dist., 22.
Geophilus shoshonesus Chmb., 59.
Giffard, W. M., personals, 48.
Graptolitha tarda B. and B., 13.
Griburius montezuma Suf., 209.
Haltica biemarginata Sa}'-, 46.
cupreolus Fall, 204.
tombacina Mann., 204.
Hamlin, J. C., note by, 54.
Hamlin, J. C., papers by, 1, 97.
Hapalotis Hbn., 19.
Harpalinffi, 65, 113.
Harpalus martini V. D., 124.
pacificus Fall, 136.
washingtoniensis V. D., 123.
Hepialus, 81.
mackiei B. and B., 82.
novigannus B. and B., 82.
roseicaput N. and D., 81.
Hermes, W. B., paper by, 49.
Homotropus decoratus Cress.,
158.
Hoplandrothrips sycamorensis
Mason, 155.
Hoplopsyllus minutus Fox, 185.
powers! Fox, 184.
Hopping, Ralph, personals, 96.
Hubbell, T. H., paper by, 39.
Hydroporus alaskanus Fall, 139.
recticollis Fall, 140.
yukonensis Fall, 138.
Hylastes yukonis Fall, 207.
Hypnoidus extricatus Fall, 191.
Hypolimnas bolina L., 160.
Hypophloeus praetermissus Fall.
199.
Kalotermes immigrans Snyd., 53.
Keifer, H. H., personals, 48.
Kessel, E. L., paper by, 159.
Kethops leioceps Chmb., 57.
Labops burmeisteri St., 163.
Laniifera cyclades Drc., 15.
Laphygma exigua Hbn., 45.
Larson, A. O., paper by, 93.
Lasiophticus pyrastri L., 159.
Lathridius carinifer Fall, 197.
cinnamopterus Mann., 197.
Lathrobium sewardi Fall, 148.
sollicitum. Fall, 147.
Lepidosaphes chinensis Chmb..
85.
Leptoglossus clypealis Held., 92.
LeptopS 3 dla ebrighti Fox, 182.
List, G. M., personals, 216.
Lophobius franciscffi Chmb., 56.
loganus Chmb., 55.
Lophoceramica artega Brns., 63.
eriopygoides B. and B., 63.
Luperina enargia B. and B., 109.
Macdougall, A. P., paper by, 165.
Macrosiphum audeni Med., 165.
bisensoriatum Med., 166.
fuscicornis Med., 167.
nigromaculosum Med., 168.
pyrifoliae Med., 169.
subviride Med., 170.
Macrotheca lecerfialis B. and B.,
64.
Martin, J. O., personals, 47.
INDEX TO VOLUME II
219
Mason, A. C., paper by, 155.
Mealybug control, 213.
Melanophila consputa Lee., 214.
Melitara prodenialis Wlk., 1.
ballii Zell., 54.
junctolineella Hist., 6.
Mesomiris albescens V. D., 35.
Millipeds of Utah, 55.
Minorista flavidissimalis Grt., 97,
roo.
Moulton, D., personals, 96.
Moneilema crassa Lee., 104.
Mylabris prolixus Fall, 204.
Nast, E. H., personals, 47.
Nebrissa Wlk., 19.
Neobaphion Blaisd., 80.
Neoborus illitus V. D., 92.
Nephelistis Hamp., 17.
Nomada klamathensis Fox, 212.
Nyctunguis molinor Chmb., 58.
Oabius paiutus Chmb., 55.
CEncyrtus moneilemae Ghn., 105.
Olyca see Melitara.
Omphalocera occidentalis B. and
B., 188.
Opuntia insects, 1, 54, 97.
Osborn, H., paper by, 22.
Papaipema dribi B. and B., 112.
Parajulus paiutus Chmb., 60.
Parasite notes, 157.
Pareniaca schwarzi Cwf., 5, 10.
Parker, J. B., personals, 48.
Parshley’s bibliography, 44.
Patroboidea V. D., 67.
rufa V. D., 69.
Pediculoides ventricosus Nwf., 93.
Perimegatoma perversa Fall, 194.
Peripsocus californicus Bnks, 87.
Phalacropsylla cummingi Fox,
182.
Philonthus fraternus Fall, 149.
septentrionis Fall, 149.
Phloeosinus cupressi Hpk., 46.
Phryganidea californica Pack.,
180.
Phytodecta notmani Sch., 202.
Piazza, E., necrology, 160.
Platynus prognathus V, D., 119.
Platyperigea alpha, B. and B.,
110 .
beta B. and B., 110.
canina Sm., 109.
Podabrus conspiratus Fall, 152.
fissilis Fall, 153.
tetragonoderus Fall, 152.
Polia falsa Grt., 107.
flavidens Grt., 187.
kappa B. and B., 12.
luteimacula B. and B., 16.
wyatti B. and B., 111.
Preparation of insects, 180.
Pristoceutophilus gaigei Hbl., 39.
Pseudanarta flavidens Grt., 187.
Pseudatrichia, key to, 164.
albocincta V. D., 164.
Pseudoptinus oviceps V. D., 121.
Pseudochseta argentifrons Coq.,
158.
Pterostichus beyeri V. D., 71.
brunneus Dej., 69.
falli V. D., 73.
lanei V. D., 76.
nigrocceruleus V. D., 70.
pacificus V. D., 114.
rufofemoralis V. D., 113.
termitiformis V. D., 74,
Rhizagrotis epipsilioides B. and
B., 106.
Rockwood, L. P., paper by, 36.
Scolopodendra utahana Chmb.,
58.
Scolytidse, 23.
Scydmaenus adjutor Fall, 144.
Scytonotus orthodox Chmb., 61.
Septis ona Sm., 108.
Sessinia collaris Shp., 54,
decolor Fairm., 54.
Shannon, R. C., paper by, 43.
Siphonaptera, 182,
Sphecomjda, 43.
dyari Shn., 43.
220
THE PAN-PACIFIC ENTOMOLOGIST [vOL. II, NO. 4
Spirostrephon utorum Chmb., 61.
Stenopelmatus fuscus Hald., 176.
intermedius D. and S., 178.
pictus Scud., 179.
Stretch, R. H., necrology, 160.
Swaine, J. M., personals, 96.
Swezey, Otto, personals, 216.
Tanner, V. M., paper by, 188.
Termites of Fanning Island, 53.
Tetropium alaskanum Fall, 201.
Tlialpius microcephalus V. D.,
122 .
Thrips, two new, 155.
Thysanoptera, 96.
Tidabius zionicus Chmb., 55.
Tillyard, R. J., personals, 160.
Timberlake, P. H., personals, 216.
Timochares ruptifasciatus PI., 12.
Tingupha australis Chmb., 62.
Trachykele nimbosa Fall, 126.
opulenta Fall, 126.
Trechus ruficollis V. D., 66.
Trichopolia Grt., 17.
suspicionis B. and B., 18.
Trogoderma sinistra Fall, 193.
Trogophloees alaskanus Fall, 146.
teres Fall, 146.
Tylobolus utahensis Chmb., 60.
Tyroglyphus farinae Deg., 92.
Underwoodia hespera Chmb., 63.
tida Chmb., 62.
Van Duzee, E. P., editorials, 47,
96, 160, 216.
Van Duzee, E. P., notes by, 22,
44, 84, 163, 180, 214.
Van Duzee, E. P., paper by, 35.
Van Duzee, E. P., personals, 47.
Van Duzee collection of Hemip-
tera, 15.
Van Duzee, M. C., paper by, 164.
Van Duzee, M. C., personals, 216.
Van Dyke, E. C., notes by, 46, 92,
95, 126, 214.
Van Dyke, E. C., paper by, 65,
113.
Van Dyke, personals, 47, 216.
Vanessa cardui L., 211.
Wachter, Sibyl, paper by, 87.
Williams, F. X., personals, 48.
Wright, W. S., note, 211.
Xanthothrix albipuncta B. and B.,
14.
Zaccotus matthewsi Lee., 95.
Zale norda Sm., 92.
Zarhipis riversi Horn, 210.
Zeugophora atra Fall, 203.
Zoological record, 11.
Zopliodia quadricolorella Dyar,
102.
ZA^gothrips californicus Mason,
156.
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