Vol. yn July, 1930 No. 1
THE
Pan-Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
WILLIAMS, BEEBEA GUGLIELMI, A CACTUS BORER . . , 1
COCKERELL, NEW SUBGENUS OF ANDRENINE BEES 5
BALL, A NEW SPECIES AND VARIETY OF SCOLOPS 9
MUIR, THREE NEW SPECIES OF AMERICAN CIXIID^ 12
DRAKE, NEW SUGAR-CANE TINGID FROM JAVA 15
BLAISDELL, STUDIES IN THE MELYRID^, VIII, 17
FLANDERS, TRICHOGRAMMA MINUTUM 20
HUNGERFORD, NEW CORIXID^ FROM WESTERN NORTH AMERICA .... 22
KEIFER, CALIFORNIA MICROLEPIDOPTERA, IV 27
VAN DUZEE, M. C., DIPTEROUS GENUS SYMPYCNUS 35
EDITORIAL COMMENT 48
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1930
THE PAN-PACIFIC ENTOMOLOGIST
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ENTOMOLOGIST
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Published at the California Academy of Sciences, Golden Gate Park,
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Entered as second-class matter, February 10, 1925, at the postoIBce at
San Francisco, California, under Act of August 24, 1912.
The Pan-Pacific Entomologist
Vol. VII, No. 1 July, 1930
BEEBEA GUGLIELMI SCHAUS, A PYRALID MOTH
BORER OF THE OPUNTIA CACTUS IN THE
GALAPAGOS ISLANDS
BY FRANCIS X. WILLIAMS
Ha’waiian Sugar Planters Association Experiment Station
Beebea GUGLIELMI Schaus
(Zoologica; New York Zoological Society. V, 2, p. 46, PI. II,
Fig. 22, 1923, male, and notes by William Beebe 1, c. v, 3, p. 54, 1924.)
This rather large, silky, grayish-brown moth that belongs
to the subfamily Pyralinse, was found by Dr. Schaus to repre-
sent a new genus and species and was described by him from a
male specimen taken by William Beebe in April 1923, on
Chatham Island, Galapagos, on the Williams Galapagos Ex-
pedition of the New York Zoological Society.
A small reared series (5 $ $ and 3^3) of this moth,
together with its larva, pupa and empty cocoons,^ were col-
lected by the writer while he was field entomologist to the
California Academy of Science’s Expedition to the Galapagos
Islands in 1905-1906. The few drawings and other data refer-
ring to this insect and which for the most part have lain fallow
for over two decades, may be of some little interest in view of
the importance of certain insects that live upon cacti of the
genus Opuntia, and so they are herewith presented.
Beebea guglielmi is evidently widely distributed over the
more arid lowlands of the Galapagos where its food plant, the
Prickly-pear cactus, is abundant. Beebe’s specimen was taken
on Chatham Island ; specimens were reared by me from mate-
rial collected on Albemarle (Cowley Mountain), Bindloe and
Abingdon islands, and I also collected, or observed, hatched
cocoons of this moth on Daphne Islet and South Seymour
Island, both near Indefatigable Island, and on James and
Duncan islands. The adult moths issued during September and
1 All this material is in the collection of the California Academy of
Sciences.
2
THE PAN-PACIFIC ENTOMOLOGIST [vOL. yil, NO. 1
October, 1906, from infested cactus branches collected a short
time previous. The female of Beehea gugtielmi, which seems
not to have been described, appears to average somewhat larger
in size than the male and its antennae, instead of being plumose
as in that sex, are thread-like with two ranks of fine bristle-like
hairs along their length. The largest female specimen has a
wing expanse of 72 millimeters or nearly three inches. The
type, a male, is in the collection of the United States National
Museum and expands 58 millimeters. Unfortunately the large
larva and pupa collected are not now available and so adequate
descriptions of them cannot be made. The larva, when full
fed, is of a dirty whitish color though pinkish above, and bears
brownish plates. It is solitary in habit, at least when well
grown, and its presence in the cactus is revealed by its drop-
pings at, or below, the entrance to its boring which, often situ-
ated in the middle of a flat branch, is irregularly commodious.
Such a gallery is weakly lined with fine silk and its opening
to the exterior may be extended as a delicate silken tube. When
ready to form a cocoon the larva leaves its tunnel and spins
a rather tough cocoon some 33-35 millimeters long by about
a third as thick, securing it along one side to the cactus plant
and covering it to some extent with available material, such as
cactus spines and lichens, so that the structure is not readily
discernible. Soon it moults to change into a reddish-brown
pupa that may be 26 millimeters long, as is the specimen
figured.
The caterpillars of Beebea were not observed doing much
damage to their food plant and without doubt natural enemies
hold it somewhat in check.
Of the rather large number of insects that attack the prickly
pear in the United States and American tropics and subtropics
a considerable proportion are the caterpillars of moths belong-
ing to the family Pyralidae. Of these are several species of
the genus Melitara (sens, lat.) represented by species the cater-
pillars of which are more or less solitary or that live in colonies,
i. e., gregariously within the plant.
Considerable work has been done on cactus insects; of the
publications referring to this subject, “The Principal Cactus
Insects of the United States,” by Hunter, W. D., Pratt, F. C.,
and Mitchell, J. D., U. S. D. A., Bu. Ent. Bull. 113, 192, is
JULY, 1930] WILLIAMS— CACTUS BORER
3
to show excavation made by the caterpillar. Cowley Mountain,
Albemarle Island.
Figure 3. Branch of Opuntia cactus showing profile view of the
delicate silk-spun tube that extends from the entrance to the burrow
of a large caterpillar. Cowley Mountain, Albemarle Island.
Figure 4, Broadside view of portion of Opuntia cactus branch
to show excavation and the ejected frass of the caterpillar. Cowley
Mountain, Albemarle Island.
Figure 5. Pupa. X 1.35. Galapagos.
4
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. VII, NO. 1
important. Of more recent date is J, C. Hamlin’s paper,
‘‘Important Opuntia Insects of the United States”; “Biological
Notes on the Moth Borers (Pyralidae),” Pan-Pacific Ento-
mologist, II, pp. 1-11, 1925. Of still more recent date is a
paper by A. C. Dodd (“The Biological Control of Prickly Pear
in Australia,” Bull. 34, pp. 1-44, 9 plates; Council for Scien-
tific and Industrial Research, Commonwealth of Australia,
Melbourne, 1927). To quote from Dodd’s paper, p. 24: “The
internal- feeding, boring, or tunneling caterpillars form one of
the largest groups of cactus insects” . . . “Most of these cater-
pillars form a compact group of closely related genera, each
genus containing several species. Those genera are Melitara,
Cactoblastis, Yosemitia, Zophodia, and Ozamia, in North
America, and Cactablastis and Tucumania in South America,”
and p. 31 : “Without doubt, Cactoblastis cactorum is the most
promising of the internal- feeding caterpillars.” This species,
which lives in large colonies, is closely related to Melitara.
A NEW EDITION OF COMSTOCK’S MANUAL^
All students of insects, especially beginners and those
engaged in teaching entomology, will welcome a new edition of
Dr. Comstock’s Manual. Professor Herrick has undertaken
the task of cutting down and bringing up to date the old
Comstock Manual on the plan laid out by Dr. Comstock and
has done this well. So we now have available a text book for
students and amateurs that leaves little to be desired. It is
concise and accurate and covers the subjects for which the work
was designed. For more detailed studies one should consult
Comstock’s Introduction to Entomology or some similar work.
The illustrations and typography are excellent and the book is
not overly large or unwieldy.
Personally, and as a student of the Hemiptera, the present writer
does not approve of separating the Homoptera as an order distinct
from the Heteroptera, as the only character available, the position
of the rostrum, is not a difference in structure, but of position, and
there is no clear line of demarkation between these and the Heter-
optera. If used as separate orders the first should have been desig-
nated as Heteroptera, as the name Hemiptera applies to both groups
and is not available for either separately. — E. P. Van Duzee.
1 A Manual for the Study of Insects, Revised Edition. By John Henry
Comstock, Anna Botsford Comstock and Glenn W. Herrick, Nineteenth
Edition. Comstock Publishing Company, Ithaca, New Tork, 1930, $4.00.
JULY, 1930] COCKERELL NEW SUBGENUS . OF BEES
5
A NEW SUBGENUS OF ANDRENINE BEES
BY T. D. A. COCKERELL
I am greatly indebted to Mr. P, H. Timberlake for per-
mission to study and describe another of his remarkable dis-
coveries. It is an Andrenine bee, which is so peculiar that
when a specimen was submitted to a well-known authority some
years ago, it was declared to be apparently a new genus of
Panurgidae.
Ancylandrena Cockerell, new subgenus of Andrena
Medium-sized bees, with the general characters of Andrena; the
male with abundant long erect hair on head, thorax and abdomen;
face with a pale band along each inner orbit; ocelli in a slight curve;
cheeks rounded, not toothed; antennae ordinary, third joint (which
has short brownish hair on upper side) much longer than fourth,
but not so long as fourth and fifth; the second and third joints are
dull and rugosopunctate, contrasting with the very minutely sculp-
tured, somewhat shining joints beyond; last joint obliquely truncate
at apex; malar space practically obsolete; mandibles ordinary; briefly
bidentate; labrum transverse, with a crescentic sulcus, its lower
margin with long straight orange hairs; maxillary palpi long, six-
jointed, the joints more or less equal, but the fifth much shorter
than fourth or sixth, and the second distinctly shorter than first;
labial palpi ordinary; tongue very short, pointed; anterior wings with
small lanceolate stigma; marginal cell ending obtusely just below
costa; basal nervure almost meeting intercubitus, but a very little
distad of it, three cubital cells, the second large and subquadrate,
receiving the first recurrent nervure near its apex; legs ordinary;
pygidial plate large and pointed; genitalia peculiar, the claspers with
a large tubercle below, the saggittse represented by a long curved
swordlike structure (whence the subgeneric name), extending for-
ward;^ the seventh ventral plate terminates in a crescentic struc-
ture, beset with long hairs; the eighth plate is not notched.
Female with the shining clypeus flattened on disc; malar space
very short but evident; labrum obtusely binodose; antennae with third
joint very long, longer than next two together; facial foveae very
short, not extending down to level of antennae, overlapped by long
hairs; lower part of eyes with scattered hairs; area of metathorax
smooth and shining; abdomen with well-defined hair bands; scopa
of hind legs dense and compact, the hairs nearly all simple.
I do not find any hair on the eyes of the male. There has
been some question about the association of the sexes, but
apparently they belong together, as they are sufficiently alike,
1 Mr. E. Nelson points out to me that the otherwise very different
Andrena cratsegi Rob. has a similar downwardly directed structure.
6 THE PAN^PACIFIC ENTOMOLOGIST: [vOL. VII, NO. 1
fly at the same time, and there is nothing else known in flie
fauna to associate with either.
In Robertson’s system this falls nearest to Opandrena, to
which it seems really to be related. It is an isolated type, and
until or unless more species are found it will be difficult to
say how many of the above characters should be included in
the diagnosis of the subgenus. The relationship seems to be
North American entirely. I find nothing similar in the Asiatic
fauna. ^
Andrena (Ancylandrena) heterodoxa Cockerell
new species
Male (type). Length about 8.5 to 10 mm.; black, with a creamy-
white band along each anterior orbit, rather broad just above mandi-
bles, gradually narrowing, and coming to a point just below level
of antennae'; head broad, facial quadrangle about square; clypeus
and sides of vertex shining; face, front and cheeks covered with very
long white hair; vertex with long black hairs, but also some pale;
disc of mesothorax brilliantly polished, the punctures scattered and
feeble, but the marginal areas are dullish; scutellum moderately
shining; area of metathorax triangular, shining, with a transverse
ridge; thorax with abundant long white hair, mixed with black
dorsally, giving a gray effect; tegulse shining black; wings hyaline,
with very dark nervures and stigma; legs with white hair, creamy
white on inner side of hind basitarsi; abdomen dullish; closely and
rather coarsely punctured, the edges above the apical depressions
rather swollen; second tergite in middle depressed about a third;
depressed portions punctured, their hind margins with extremely
narrow pallid bands; tergites with long erect white hair, faintly
stained with brownish; white marginal hair bands fairly distinct on
third and fourth, and well developed on fifth; apex with white hair;
venter shining.
Female. Length about 11 mm.; facial fovese pale reddish; vertex
and disc of thorax (invading middle of scutellum) with much black
hair; legs with mainly white hair, the scopa of hind legs shining;
anterior tarsi with hair soft gray or dilute black on outer side, pale
orange on inner; middle and hind tarsi with white hair, fulvous at
apex of basitarsi within; spurs light ferruginous; abdomen shining,
first tergite with long white hair, second to fourth vtdth thin short
black hair; but tergites 1 to 6 with broad pure white hair bands,
more or less interrupted on first; caudal fimbria pale gray; second
2 The old World Ancyla has marginal cell ending far from costa; hind
legs of male peculiar.
JULY, 1930] COCKERELL^ — NEW SUBGENUS* OF BEES 7
tergite in middle depressed considerably more than half, the margin
of the depression strongly arcuate, so that it is narrow at the sides;
venter with white hair bands. The third and fourth tergites have
a band of pale hair at the base. The flagellum is distinctly rufescent
beneath.
Type (male) at flowers of Cryptanthe intermedia, River-
side, California, April 25, 1930 (Timberlake). Another male
at Eriogonum fasciculatum, May 20, 1926. Females at Lupinus
paynei, May 11, 1929; also at Lotus scoparius and Hugelia
virgata*
The male will be easily recognized by the peculiar face
markings. The following table separates the female from
others more or less similar in appearance, being of medium
size, and with conspicuous hair bands.
Caudal fimbria dusky chocolate color or gray 1
Caudal fimbria pale 4
1. Wings very dark; abdominal bands fulvous-tinted; basal nerv-
ures going basad of nervules fulvipennis Smith
Wings not dark; basal nervures meeting or (some sapellonis)
falling short of nervules 2
2. Disc of mesothorax with black hair heterodoxa n. sp,
(Compare also peckhami Cockerell.)
Disc of mesothorax without black hair 3
3. Clypeus dullish, with a median ridge sapellonis Ckll.
Clypeus polished without median xi6.Qt.....electrica Casad and Ckll.
4. Flagellum, except basally, bright ferruginous beneath 5
Flagellum dark 6
5. Larger; abdomen highly polished gardinari Ckll.
(Equally large; flagellum obscure red beneath, abdomen
finely punctured.. ellisia Ckll.)
(Stigma small; flagellum obscurely reddish beneath
relatia Vier.)
Smaller; abdomen less polished, but mesothorax posteriorly
and scutellum brilliantly polished and impunctate..
persimilis Graenischer
(Also smallish, flagellum very obscurely reddened, but
mesothorax posteriorly and scutellum polished in
same fashion; clypeus with a median ridge, not
present in persimilis. Two cubital cells gibberisYitx.)
* Since submitting the above material to Dr. Cockerell, another male
was taken May 5 on Cryptanthe, at Riverside, California, and on May 15
Mr. C. M. Dammers and the writer collected seven males at flowers of
Calochortus splendens on the Gavilan Mesa about fifteen miles
south of Riverside. — P. H. Timberlake.
8 THE PAN^PACIFie ENTOMOLOGIST [vOL. VII, NO. 1
-6. Mesothorax with short velvety tomentum coachifostica Vier.
(See also plana Yitr., with chocolate fimbria.)
Mesothorax without such tomentum 7
7. Mesothorax entirely dull 8
(Dull in front; clypeus with a ridge ellisia Ckll.)
Mesothorax more or less shining... 9
8. Clypeus dull (Los Angeles, California; Coquillett)
mustelicolor huardi Vier.
Clypeus highly polished in middle (Texas, Bellfrage)
relativa Vier.
9. Clypeus dull; wings fuliginous 10
(Wings with dusky cloud at apex fracta Casad and Ckll.)
Clypeus shining 11
10. Tegulas reddish black (Stanford University, California, March,
on mustard) scurra Vier.
Tegulas testaceous (Southern California)
davidsoni Vier. and Ckll.
11. Upper part of clypeus flattened, no median ridge; basal
nervure meeting nervulus sieverti Ckll.
Clypeus with a median ridge 12
12. Larger; three cubital cells ellisia Ckll.
Smaller; two cubital cells gibheris Vier.
Lepidomys irrenosa Guenee (Lepidoptera)
In arranging the Noctuidae in the collection of the Cali-
fornia Academy of Sciences I found in the Koebele collection a
little moth bearing the above name. It agrees in every respect
with Guenee’s description, except that it is more ochraceous
than one would expect from that description, but it is strongly
tinted with wine color, especially toward the apex of the wings.
This specimen was taken by Mr. Koebele in Florida. Guenee
says his type, a unique, was from New York and was taken by
Doubleday. Dr. J. B. Smith (1893) says this type bears but
one label, “Doubleday,” with no mention of locality, and
strongly suspects the insect to be tropical. Perhaps on the
strength of this it was omitted from the Dyar and the Barnes
and Mcdonnough lists. So far as I can learn there has been
no recorded capture of a second specimen. It must now be
re-established in our North American lists. — E. P. Van Duzee.
JULY, 1930] BALL NEW SPECIES OF SCOLOPS
9
A NEW SPECIES AND VARIETY OF SCOLOPS
WITH NOTES ON OTHERS (RHYNCHOTA
FULGORID^)
BY E. D. BALL
University of Arizona, Tucson
While working up recently collected material in the genus
S colops, the writer was impressed with the fact that most spe-
cies had definite food plants to which they were apparently
confined, and that in no case was the food plant a grass as
Dozier and others have suggested. The fact that osborni Ball
feeds exclusively on the compass plant of the lower plains
region {Silphium laciniatum L.) has already been recorded.
The writer on checking over his notes found that at least four
other species, one of them being new, were apparently confined
to different members of the Compositse, two to different mem-
bers of the Chenopodiacese and one to an Euphorbiacese. The
two most widely distributed species of the genus, sulcipes Say
and pungens Germ., are found abundantly in weedy and waste
places and are likely to be found to be Compositae feeders also.
Breakey states that the typical habitat of the group is in moist
places near the edge of woodlands and in low spots in the
prairies. This, however, conveys a decidedly erroneous im-
pression of their habits as observed by the writer. They are
lovers of hot, dry and open situations, and are rarely found in
damp or shaded locations.
S. robustus Ball has been found by the writer feeding
exclusively on the perennial ragweed {Ambrosia psilostachya
D. C.) in Colorado, Utah, and Arizona. S- viridis Ball, a pale
green species, is exactly the color of the foliage of the salt-
bush {A triplex canescens James), on which the writer has
always found both nymphs and adults feeding.
S colops graphicus Ball, new. species
Intermediate in size, form, and color markings between
maculosus Ball and uhleri Ball, and resembling stonei Break,
in color and pattern, but much smaller. Dark gray with
irregular white streaks and flecks, strikingly resembling the
dorsal pattern of an eastern quail or meadow lark. Length,
6-7 mm.
10 THE PAN-PACIFIC ENTOMOLOGIST [ VOL. VII, NO. 1
Cephalic process slightly longer than in maculosus, much narrower
and straighter, not as long as in uhleri, but even more slender.
Elytra very broad and short giving a peculiar and characteristic
truncate appearance. Cubitus forking just back of the junction of
the claval veins, and medius forking distinctly beyond the cubitus.
Color dark, smoky, with broad white costal margins, a pair of
narrow wavy stripes along the inner-claval veins, a secondary stripe
arising on cubitus before the fork and fusing posteriorly with the
inner stripe. These two stripes set off a definite dark brown or black
stripe centering on medius with four or five white dots along the
vein and a row of black spots around the apex. Vertex greenish
with a pair of obscure spots; disk of pronotum white with a pair of
dark spots. Scutellum with a broader median wavy stripe and a
pair of widely separated dots. Heavily marked examples lose the
inner pair of ivory stripes.
Holotype, female, allotype, male (in E. D. Ball collection)
and four paratypes taken by the writer on the “match brush"
or rayless goldenrod (Gutierresia calif ornica) at Yarnell
Heights, Arizona, October 8, 1929, and six paratypes taken at
Cline, Arizona, August 2, 1929, on the same plant. Only
nymphs were taken July 14, adults and nymphs later in July
and August, and only adults in October. This is the smallest
species yet described and it appears to be strictly confined to
this very fine-stemmed plant.
S colops uhleri Ball has been taken by the writer many times
on the annual Dondia depressa Wat. growing in alkali areas
in western Colorado and southern Utah.
Scolops uhleri marginatus Ball n. subsp.
Resembling uhleri in general size and form, but usually
larger and lighter colored with a pair of broad smoky or black
stripes just inside the costal ivory stripes. The dark stripes
with only two or three light spots on the veins. About four
black spots at the apex. Cephalic process slightly shorter and
much darker than in typical uhleri. Length, 7-8 mm.
Holotype, female, and allotype, male (in E. D. Ball col-
lection), taken by the writer near Sacaton (labeled Tucson),
Arizona, June 19, 1929, and six paratypes taken at St. George,
Utah, July 24, 1908 (Ball), and Glendale, Nevada, June 9,
JULY, 1930] BALL NEW SPECIES OF SCOLOPS
11
1928 (E. W. Davis). This may prove to be a distinct species
when more material is available for study. At least it is a dis-
tinct subspecies apparently confined to the subtropical South-
west. All specimens taken have been from the perennial shrub
(Dondia torreyana Wat.). Large nymphs and just emerging
adults were taken June 8 by the writer and only a few adults
were found June 23.
S. perdix Uhl. was taken by Mr. Stone and the writer in
abundance June 22, 1927, at Sanford, Florida, on a ridge in a
flat woods area where the narrow-leaved sunflower {Helianthus
angustifolius L.) appeared to be the only possible food plant.
The known distribution and abundance of this species coincides
very well with the distribution of this plant.
S. stonei Break. This very distinct species was found by
Mr. William E. Stone (for whom it was named), Mr. J. A.
Reeves, and the writer, to be confined entirely to the milky-
juiced “queen’s delight” (Stillingia angustifolia Torr.). Small
nymphs were found at Sanford, Florida, in June, large nymphs
and adults in July, and adults only in August and early
September.
Both nymphs and adults of S', snowi Break, were taken by
the writer on goldenrod (Solidago trinervata) at Long Valley,
FlagstaflF, and Williams, Arizona, in August 1929.
Breakey gives California and Idaho under the distribution
of S', hesperius. The writer, who has made long series of col-
lections in both areas, is inclined to believe that those were
erroneous determinations and that hesperius is a plains’ species
extending from Dakota and Montana south to Texas. His
statement that Ball records osborni from Ohio is in error.
S. oshomi Ball was described from Iowa and Kansas. The
writer agrees with him in doubting the distribution records of
6'. grossus from California and New Jersey and would add
Idaho. N. grossus is, like hesperius, only known from the
plains region. Breakey describes S. viridis Ball as “cephalic
process long and slender,” but places it in his key under
“cephalic process short.”
12
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
THREE NEW SPECIES OF AMERICAN CIXIID^
(FULGOROIDEA, HOMOPTERA)
BY F. MUIR
Bennarella Muir, g. n.
Body compressed laterally; tegmen when at rest steeply tecti-
form, apical margins closely pressed together. Sc + R forking about
one-third from base, level with Cu fork, the fork slightly more apical,
M rising from basal cell; three apical Rs, five apical Ms, 1, la, 2, 3, 4;
Cu touching M3 + 4 for a short distance; claval fork about one-
third from base of clavus. The costal margin near base is turned
upward where the fourth and fifth abdominal segments are pro-
duced. Vertex very short, width five or more times the length in
middle; in dorsal view a large part of the base of frons visible, base
of vertex sinuate, slightly emarginate, apex truncate. Length of
frons nearly twice the width, sides slightly arcuate; disk slightly
excavate on basal half where the lateral carinas are longer; median
Carina obsolete, only distinct for a short distance in middle. Clypeus
plainly tricarinate. Labium long, reaching beyond hind coxae.
Second joint of antennae much longer than wide, reaching base of
clypeus. Pronotum short, hind margin roundly excavate. Meso-
notum slightly wider than long, bicarinate, the median carina only
faintly represented on anterior portion. The third, fourth and fifth
abdominal pleura produced, flattened and curved dorsad, the fourth
bears three and the fifth two sense pits, each bearing a bifurcate
hair. Hind tibiae without spines; the second hind tarsus with a row
of spines at apex.
This genus comes next to Bennaria, but the size and nature
of the development of the abdominal pleura are very distinct.
Superficially it looks like one of the Achilixiidse, but an exami-
nation of the male genitalia shows that it is a Cixiidae ; also the
claval vein enters the commissure distinctly before the apex.
Type B. bicoloripennis.
Bennarella bicoloripennis Muir, sp. n.
Male. Length, 2.4 mm.; tegmen, 4.4 mm.
Head and pronotum stramineous, light brown over most of frons
and base of clypeus and pronotum behind eyes; mesonotum brown,
carinae lighter; labium fuscous; legs stramineous, the femora and
tibiae slightly fuscous; abdomen light brown, the produced fourth
and fifth pleura and the genitalia lighter. Basal portion of tegmen
to M fork dark brown, the claval margin and suture yellowish, apical
portion of tegmen yellowish, slightly fuscous in costa and in Sc and
R apical cells; veins mostly the color of the cells, without granules
on macrotrichia. Wings fuscous at base, opaquely white with waxy
JULY, 1930]
MUIR ^THREE NEW CIXIID^
13
secretion over rest, veins fuscous at base and yellowish over apical
portion.
Pygofer comparatively short, opening longer than wide, sides
curved, medio-ventral process angular, small. Anal segment pro-
duced into a point at apex; the styles and aedeagus best understood
from the figure.
Female similar to male in size and color. The ovipositor com-
plete, pygofer longer than broad, a longitudinal depression in which
the ovipositor lies when at rest; the ventral surface of anal segment
concave and fits over the apex of ovipositor when at rest.
Described from four males and five females from Tuma-
tumari, British Guiana (July 19, 1923). Type in H. S. P. A.
Experiment Station collection, Honolulu, No. 1216.
Bennarella fusca Muir, sp. n.
Male : length, 2.4 mm. ; tegmen, 4 mm.
In general structure similar to bicoloripennis, but the lateral
carinae of the frons larger and the antennae slightly shorter.
Head and thorax fuscous brown, the carinae lighter, the lateral
portion of the mesonotum darker than the middle; coxae and femora
fuscous; tibiae lighter. Tegmina fuscous, basally lighter, nodal line
lighter. Wings dark fuscous, veins dark.
Pygofer laterally compressed, sides arcuate, medio-ventral process
subconical in outline. Anal segment large, apex produced beyond
anus, curved ventrad and under, asymmetrical, the right side emargi-
nate. Genital styles small, in lateral view bent at an angle in middle,
apex slightly expanded and rounded. Aedeagus not dissected out.
Described from one male from Belem, Brazil (May 1924).
Type in H. S. P. A. Experiment Station collection, Honolulu,
No. 1217.
The above two species were taken by Dr. F. X. Williams.
Nymphocixia vanduzeei Muir, sp. n.
Male: length, 5.6 mm.; tegmen, 5.7 mm.
This species agrees with the generic description and is closely
related to N. unipunctata V. D. The base of vertex is truncate and
overlaps the middle of pronotum and the middle of the anterior
portion of mesonotum. The hind margin of the eye is nearly trun-
cate and there is only a shallow antennal emargination on the ventral
margin.
Pygofer compressed laterally, sides arcuate, medio-ventral process
fairly large, subangular; anal segment median size, apex pointed;
genital styles small, thin, apical third bent at a right-angle dorsad.
14
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
Stramineous or light brown, the Carinas of head and the genae
lighter. Tegmina stramineous or very light brown, clearer over
membrane and costal cell, veins light; a small dark spot near apex
between M 2 and 3. Wings hyaline with brown veins.
One male from Columbia, Cartagena (C, Gogzo, VIII,
1905 ).
Type in the Zoological Museum, Hamburg.
This is the second species of this interesting genus; the
genotype taken on Espiritu Santo Island, Gulf of California.
1. Bennarella, frons,
2. Bennarella, tegmen.
3. Bennarella, dorsal view of head and nota.
4. Bennarella bicoloripennis Muir, male genitalia.
15
JULY, 1930] DRAKE NEW TINGITID FROM JAVA
A NEW SUGAR-CANE TINGITID FROM JAVA AND
SUMATRA (HEMIPTERA)
BY CARL J. DRAKE
Ames, lonva
Abdastartus sacchari Drake, new species
Very closely allied to A. tyrianus Distant, but differing in the
longer basal segment of antennae, longer antennae, slightly wider
paranota, and more deeply emarginate anterior margin of pronotum.
Head and anterior portion of pronotum very strongly reflexed, not
lying in same plane as rest of body. Head very deeply inserted in
the pronotum, the base of the eyes touching collum, strongly swollen
at the base of the anterior spines, with four very long, slender, sharp
spines; anterior pair reaching beyond the first antennal segment,
parallel or converging apically, their apices testaceous; posterior pair
very long, pale testaceous, directed forward, mostly contiguous with
head, their apices reaching to near the middle of first antennal seg-
ment; antenniferous tubercles large, prominent, testaceous slightly
divergent, their tips very slightly curved inwardly. Antennae long,
slender, pale testaceous, the basal portion of first and distal half
of last segment brown; proportions (male), 7:4:57:32; (female)
7:4:42:23. Bucculae long, broad, projecting anteriorly beyond head,
closed in front. Rostral channel deep, narrow, open behind; rostrum
brown, its apex black, extending between intermediate coxae.
Pronotum closely and deeply pitted, a little more tumid than in
A. tyrianus, the collum very strongly reflexed and its anterior margin
deeply emarginate, its anterior angles much more acute; carinae pale
testaceous, slightly thicker; lateral margins slightly more foliaceous,
pale testaceous. Elytra considerably longer than abdomen, with the
costal area slightly broader and the discoidal area a little longer
than in A. tyrianus) wings a little shorter than elytra, clouded with
fuscous; the entire insect also a little larger and darker in color.
Legs long, slender, pale testaceous. Length, 2.60 mm.; width, ,64 mm.
Holotype, male, and allotype, female, Randoeblatoeng, Java,
July 7, 1923, in my collection. Paratypes, specimens taken
with type, Formosa, and one example from Sumatra. This
is the species erroneously recorded by the writer {Philippine
Journal Science, XXXIV, 1927, p. 307), as Abdastartus tyri-
anus Distant from Java and Formosa. The writer is indebted
to Mr. W. E. China of the British Museum, London, for
sketches and for comparing A. sacchari with Distant’s types
oi A. tyrianus.
Abdastartus tyrianus Distant (Fauna Brit. India, Rhynch.,
Vol. V (Appendix), p. 103, fig. 50), was described from
16
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
several specimens, collected by Mr. Max Lefroy, Pusa, Bengal,
India. Two specimens are at hand from Sumatra. Antennae
shorter and slenderer than in the foregoing species, the apical
portion of terminal segment embrowned. Head less swollen
in front; spines long; anterior pair directed forward, almost
parallel or divergent, reaching to near apex of second segment.
Head and collum very slightly reflexed, the latter moderately
concave in front. Antenniferous tubercles prominent, divergent
at base, slightly curved inwardly at tips.
The genus Abdastartus Distant contains only two species,
A. tyrianus Distant and A. sacchari n. sp. A. muiri Drake
from Java and India belongs to another genus and will be
treated in a subsequent paper.
American Entomology Loses a Distinguished Patron
The death of Dr. William Barnes on May 1, 1930, brought
a feeling of personal loss to every entomologist in this country,
and more, perhaps, to those connected with the California
Academy of Sciences than to many others. For a number of
years Dr. Barnes had taken an active interest in the entomolog-
ical department of the Academy. Most of our species of
Lepidoptera have been determined by him, or under his direc-
tion, and he has presented authentically determined specimens
of several hundred species not otherwise represented in our
collection ; and for all he did for us he expected and wanted no
return other than the satisfaction of assisting in the establish-
ment of an institution that in the future would be an important
element in the scientific work of the western states. Dr.
Barnes’ interest in the Lepidoptera began in childhood and
never slacked through a long and active life. His private col-
lection in this order of insects is undoubtedly the largest and
best studied collection of North American forms in the world.
The 500,000 specimens in his collection represent nearly 8,000
species and include 1915 holotypes and nearly 7,000 other
specimens that are really type material. Dr. Barnes had a
special building constructed for the collection and for many
years employed a trained specialist to care for it, and at all
times was most generous in determining material for other stu-
dents. His death seems an irreparable loss to American Lepid-
opterology. — E. P. Van Duzee.
JULY, 1930] BLAISDELL STUDIES IN MELYRID^ 17
STUDIES IN THE MELYRID^, NUMBER EIGHT
BY FRANK E. BLAISDELL, SR.
Stanford Medical School and California Academy of Sciences
San Francisco, California
The very distinct species of Trichochrous described below
belongs to Casey’s second group of species. It is necessary to
give a detailed description, as the increasing number of new
forms makes it obligatory if the numerous species are to be
recognized by description alone. Whenever possible, collectors
should secure a moderate series of species in the locality where
they are discovered, so as to determine the degree of individual
variation. This is exceedingly essential so as to eliminate
guesswork in the definition of species.
Trichochrous quadrinotatus Blaisdell, new species
Form oblong, slightly oval. Color black, trophi slightly
rufo-piceous, tarsi and distal half of tibiae rufous to piceous.
Surface more or less shining and without metallic luster.
Pubescence not dense nor coarse, moderately short, not com-
pletely recumbent and pale in color; brownish to blackish and some-
what coarser hairs form two more or less quadrate spots on each
elytron, the subbasal macules most so, while the larger at apical
third tend to diffuse both toward apex and anteriorly, connecting
more or less narrowly or more broadly with the subbasal; occasion-
ally the brownish hairs become more diffused and nearly replace the
pale hairs. Hairs of the pronotal disk and head more or less
brownish; on the deflexed sides of elytra, especially toward apex
and on apical declivity, they become coarser and semi-erect; brist-
ling erect setse entirely absent. Lateral pronotal fimbriae moder-
ately short, regular and evenly close-set, pale to blackish in color;
those of the elytra a little longer, quite even and more widely spaced,
directed obliquely postero-laterad. Pubescence of undersurface of
body fine, closely recumbent and more plumbeous in color; that
of sternal side pieces a little coarser and more cinereous.
Head slightly transverse, in width about equal to the pronotal
length and apical width, muzzle short; frons plane, surface very
finely and microscopically reticulate, finely rugulose in clypeal
region and there with a slight glabrous convexity, limited at times
by feeble lateral depressions; punctures small, evenly and rather
widely spaced, separated by a distance equal to two or three times
their diameter. Eyes large, oval, evenly convex and prominent,
rather finely faceted. Antennae less than moderately stout, about
as long as head and pronotum; third joint smallest, intermediate
18
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
joints slightly serrate anteriorly, outer four more rounded, gradually
and evenly incrassate.
Pronotum transverse, widest at basal third or about middle,
about a half wider than long and two and a half times longer than
an eye, moderately and evenly convex from side to side; apex feebly
and broadly arcuate in circular arc, apical angles rounded; sides
moderately arcuate, straighter and more convergent before basal
third, thence more strongly and more or less briefly sinuate before
the small obtuse basal angles, margin finely crenulate; base broadly
and moderately arcuate, slightly sinuate within the angles; disk more
or less microscopically rugulose, more densely and strongly so in
lateral fourth, punctures small in the central area, separated by a
distance equal to one to three times their diameter.
Elytra oblong, about two-sevenths longer than wide, three times
as long as pronotum and about a sixth wider; humeri broadly
rounded, slightly prominent and somewhat tumid; sides parallel,
feebly and very broadly sinuate behind humeri, thence feebly arcuate,
becoming more evenly so to parabolically rounded apex; disk slightly
convex on dorsum, becoming more strongly and declivously so later-
ally, very gradually declivous apically; punctures coarser than on
pronotum and separated by a distance equal to one to three times
their diameter, surface somewhat rugose, smoother and punctures
smaller toward apex.
Undersurface of body very finely and more densely punctate.
Legs moderate in length and slenderness; femora not stout, tarsi
somewhat stout and about equal to their tibia in length, the latter
quite as long as the femur, similar in the sexes.
Male. Pronotum more broadly arcuate at sides, widest about
middle; antennas slightly heavier; elytra rather more oblong and
parallel; fifth ventral abdominal segment truncatosinuate at apex.
Female. Pronotum usually widest at basal third; elytra very
slightly wider posteriorly; fifth ventral broadly but not strongly
arcuate at apex.
Measurements. Length (types), 2. 5-2. 7 mm.; width, .9-1 mm.
Largest female of the series measures length, 3 mm.; width, 1.2 mm.
Holotype, male (No. 2969), and allotype, female (No. 2970,
Museum California Academy of Sciences), and paratypes in
my collection; also in that of Mr. Warwick Benedict (Museum
University of Kansas).
Type locality : Idylwild, Riverside County, California. A
series of twenty-five specimens collected by Mr. Benedict on
May 29, 1923.
In the series at hand the brown hairs of the upper surface
at times by diffusion reduce the paler pubescence to a minority.
JULY, 1930] BLAISDELL STUDIES IN MELYRID^
19
In typical individuals the four dark areas on the elytra are
distinctly quadrate with a minimum amount of diffusion.
Quadrinotatus is to follow vilis Casey in our lists. By the
dark discal elytral macules it differs radically from other mem-
bers of Casey’s second group of species. At first sight it might
be associated with insignis Casey and curticollis Casey on
account of the elytral pattern, but those species have the body
bristling with long erect setae and belong to Casey’s third group
in which there is no regular close-set fimbriae on lateral margins
of pronotum.
Trichochrous punctipennis Lee.
The type locality of this species is Santa Catalina Island
off the coast of Los Angeles County. My own series was col-
lected at Avalon on that island. Mr. Warwick Benedict of the
Entomological Department of the University of Kansas has
recently submitted to me for study a series of fourteen speci-
mens, taken by him at Ensenada, Lower California. They
were collected on July 19, 1924. This is the first record of
the species having been taken on the mainland.
Listrus niveicanthus tincticornis Blais.
The type specimens of this subspecies were taken at Pasa-
dena, California. A considerable series has very recently been
sent me for examination that were collected at Warner’s Hot
Springs, San Diego County, by Mr. Benedict. The differential
characters have been fully discussed by me in my paper pub-
lished as one of the Stanford University publications, Univ.
Series Biol. Sciences, Vol. I, No. 3, June 1921, p. 169.
In Memory of Mr. N. R. Gunn
In the death of Mr. Norman R. Gunn the San Francisco
Butterfly Club has lost one of the most enthusiastic of its
younger members. Mr. Gunn was born at Breckenridge, Min-
nesota, June 27, 1913. He was a student at the Berkeley High
School and had planned to make entomology his life work,
intending to carry on his entomological studies at the Univer-
sity of California. His death at Tacoma, Washington, on July
27, 1930, was due to a heart attack while in bathing. The
heartfelt sympathy of his entomological associates is extended
to the bereaved mother and relatives. — R. F. Sternitzky.
20
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
RACES OF TRICHOGRAMMA MINUTUM
BY STANLEY E. FLANDERS
Citrus Experiment Station, Riverside, California
According to Howard and Fiske, the “pretiosa-like” form
of Trichogramma is made up of two races, one arrhenotokous
and the other thelyotokous.
Marchal found that the thelyotokous race near Paris dif-
fered in color from those obtained in central France, and he
suggested that if observations were made on Trichogramma
from various localities a number of strains would be found
having the status of races or elementary species.
In North America a thelyotokous race of Trichogramma
is yet to be observed. Differences in color, however, have
been observed ranging from “black” to light lemon yellow. The
“black” race studied by Martin is dark fuscous throughout. It
is unique in that the males normally have vestigial wings.
In a paper presented at the Ithaca Congress in 1928 the
writer called attention to the fact that the coloration of the
females varied with the developmental temperatures from light
yellow to the dark color characteristic of the male. The color
of the males is much less affected by temperature.
Peterson first noted that a race existed the females of
which did not become yellow when reared at high tempera-
tures. He also found that the two races did not interbreed and
that the light race appeared to be typically northern, the dark
race southern.
In studying the strains of Trichogramma from various
localities in North America, the writer segregated them accord-
ing to color and life cycles into five races. These maintain
their distinctive characteristics when reared under the same
conditions for generations.
Only the minutum-like form of Trichogramma which is
arrhenotokous and normally has winged males is included in
the classification.
The specimens used were all reared in the eggs of Sitotroga
cerealella at constant temperatures. Hoyer solution was used
1930] FLANDERS— TRICHOGILA.MM A 21
for the mounting medium. The observations on the color of
the specimens were made against a white background.
The length of the life cycle of each race differs from the
others. When reared at a constant temperature of 60 degrees
Fahrenheit the differences are very apparent. The races are
listed below in the order of their lengthening life cycles :
1. Dark short-cycle race (from west coast of Mexico and
southern California) :
A form having dark gray and yellow pigments present at
all temperatures. (Life cycle at 81 degrees Fahrenheit
is seven days.)
2. Yellow race (from Massachusetts, Illinois, and California) :
A clear yellow form having dark gray markings only
when reared at a temperature less than 70 degrees
Fahrenheit.
3. Transition race (from California) :
A yellow form having dark gray markings except when
reared at 90 degrees Fahrenheit or above.
4. Gray race (from Louisiana and Georgia) :
A form like the dark races except that the pigmentation
is less dense. The yellow pigment often is absent in both
male and female when reared at 90 degrees Fahrenheit
so that the adult appears silvery gray with dark gray
markings. (Life cycle at 81 degrees Fahrenheit is eight
days. )
5. Dark long-cycle race (from California and Massachusetts) :
A form like the dark short-cycle race except that head
capsule retains fuscous markings at all temperatures and
the life cycle at 81 degrees Fahrenheit is eight days.
The races with the shorter life cycles appear to have higher
optimum developmental temperatures.
The material for this preliminary study was received
through the kind cooperation of C. H. Alden, C. O. Bare,
William G. Bradley, H. J. Franklin, W. P. Flint, Dr. Philip
Garman, E. A. Maier, Dr. Herbert Spencer, W. J. Schoene,
and R. W. E. Tucker.
22
THE FAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
NEW CORIXID^ FROM WESTERN NORTH
AMERICA (HEMIPTERA)
BY H. B. HUNGERFORD
Department of Entomology, University of Kansas, La<wrence
In reporting some new species of Corixidse from the west-
ern part of North America, I take this occasion to designate
two new groups.
Krizousacorixa Hungerford, genus new
Surface nonrastrate. Eyes small, separated from the margins of
the head by a comparatively wide space. Rostrum small, trans-
versely wrinkled. Fourth segment of antenna less than half the
length of the third. Lateral lobe of prothorax a little longer than
broad, the superior margin a little shorter than the inferior one,
deep set, its base nearly obscured by the side of the head in front
and the inflated margin of the mesothorax behind. Male strigil
right. Legs comparatively short. Anterior femora of the only
known species inflated, with large patch of stridular pegs at base.
Genotype Corixa femorata Guerin, the type of which I
have seen in the Paris Museum. A drawing of the head and
front legs of this species was published by me on Plate I in
the Bulletin of the Brooklyn Entomological Society, Vol. XX,
No. 1, 1925.
This curious genus appears to stand quite alone. The wide
space separating the rear margin of the eye from the rear
margin of the head is in common with Graptocorixa, Cymatia,
and Glaenocorisa. From these it is separated by the relatively
small eyes and by a very different pala in the males.
Graptocorixa Hungerford, genus new
Surface of hemelytra rastrate, transversely marked with undu-
late bands. Eyes prominent, a facial area pubescent, face short,
rostrum small. Space beneath the lateral margin of the eye wide.
Postocular space wide. Fourth segment of antenna less than half
the length of the third. Lateral lobe of prothorax quadrate to rect-
angular with anterior distal angle incurved and posterior (upper)
part usually thickened, mesoxyphus bilobate at tip. Dorsum of
abdomen more or less red. Male strigil right and in size varying
from minute to large. Legs comparatively short and stout. Anterior
pala falcate terminated by a stout claw.
Genotype Corixa abdominolis Say. To this genus also
belong: Corixa serrulata Uhler, Corixa melanogaster Kirk.
JULY, 1930]
HUNGERFORD NEW CORIXID^
23
(= Corixa unguiculata Champ.), Arctocorixa calif ornica
Hungerford, Arctocorixa uhleri Hungerford, and Arctocorixa
gerhardi Hungerford.
This genus has some relation to Cymatia Flor. and Glseno-
corisa Thoms. It agrees with them in having the fourth seg-
ment of the antenna less than half the length of the third and
in having a broad postocular space on the head. Like Cymatia
Flor, it possesses a strong claw on front pala, but does not
have a smooth rostrum as in Cymatia.
The distribution of this group is distinctly southwestern
North America. The records from specimens in the Kansas
University entomological collections are as follows :
Graptocorixa abdominalis Say
Sutton County, Texas; Presidio County, Texas; Valentine,
Texas; Alpine, Texas; Palo Verde, Imperial County, California;
Espirita Santo Island, Gulf of California (Calif. Acad. Sci.) ; Organ
Mountains, New Mexico; Mexico; S. E. Kane County, Utah; Cata-
lina Mountains, Arizona; Hot Springs, Arizona; Warsaw, Arizona;
Superstition Mountains, Arizona; Quartzside, Arizona; Culberson
County, Arizona; Sabino Basin, S. Catalina Mountains, Arizona;
Cochise County, Arizona; Gila County, Arizona (many of these
approach the variety bimaculata Guer.) ; Yavapai County, Arizona;
Santa Cruz, Arizona; Oracle, Arizona (Calif. Acad. Sci.); Yuma
County, Arizona (Calif. Acad. Sci.).
Graptocorixa abdominalis, var. bimaculata Guer.
Mitla, Mexico; Mexico; El Paraiso, Lower California, Mexico;
Colima, Vulcano, Mexico; Guatemala; Costa Rica.
Graptocorixa serrulata Uhler
San Antonio, Mexico; Valentine, Texas; Presidio County, Texas;
Gila County, Arizona; Santa Cruz County, Arizona; Yavapai County,
Arizona; Cochise County, Arizona; Oracle, Arizona (Calif. Acad.
Sci.).
Graptocorixa uhleri Hungerford
Described from California. Some from Monrovia Canyon,
Californig., differ slightly from the types.
Descanso, California (Calif. Acad. Sci.); Santa Cruz Island, Cali-
fornia (Calif. Acad. Sci.).
Graptocorixa californica Hungerford
San Jacinto Mountains, California; “Californie Lansweert”; Pasa-
dena, California; San Diego County, California; Huntington Lake,
California (Calif. Acad. Sci.).
24
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
July, 1930] hungerford — new corixid^
25
Graptocorixa gerhardi (Hungerford)
Santa Cruz County, Arizona; Musquiz Canyon, Fort Davis,
Texas; Brewster County, Texas. (These are slightly different from
the types.)
Graptocorixa melanogaster (Kirkaldy)
Costa Rica, Mexico.
It is interesting to note that this genus appears to have a
wider range than my Neocorixa which has been taken only in
Gila County, Arizona, and Torance County, New Mexico. In
some ways Neocorixa and Graptocorixa have a common resem-
blance. In the one the strigil of the male abdomen is on the
left side, and in the other it is on the right side. I cannot find
any indication of the curious abdominal asymmetry in Grapto-
corixa that is possessed by the females of Neocorixa.
Arctocorixa omani Hungerford, species new
Size. Length, 6.6 mm. Width across head, 2.25 mm.
Color. General facies medium brown. Distinct barring on pro-
notum and basal half of clavus, nine to ten dark bands on pronotum.
The pale bands broader than the dark ones on basal half of clavus,
pattern on remainder of hemelytra reticulate, sometimes confused
with only a suggestion of cross-line effect, except embolium which
is sooty brown to black. Vertex, face and limbs light, venter more
or less dark.
Structural characteristics. Pronotum and clavus rastrate. Head
and pronotum short. Interocular space appearing a trifle narrower
than an eye. Metaxyphus about as long as wide, of the usual tri-
angular shape. Male facial fovea shallow, not attaining the eyes
laterally. Male pala of usual form as shown by Figure 3 on Plate 1,
with twenty-one pegs in the row. Strigil of average size, a little
longer than wide, composed of six striae, the outer one incomplete.
Male genital capsule as shown in Figure 4.
Described from thirty-four specimens from Carson City,
Nevada, August 9, 1929, taken by R. H. Beamer. Holotype,
allotype, and some paratypes in University of Kansas col-
lection. Paratypes also in United States National Museum and
the California Academy of Science.
I have to report this species also from : Phillips Station,
Eldorado County, California, July 24, 1921, 7000 feet eleva-
tion, collected by F. E. Blaisdell (Calif. Acad. Sci.) ; Mono
Lake, California, May 12, 1917, C. L. Fox collector (Calif.
26
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1,
Acad. Sci.) ; Mount Kaiser, California, July 19, 1919, Fresno
County, 10,000 feet., F. C. Clark collector (Calif. Acad. Sci.) ;
Eagle Lake, Lassen County, California, July 31, 1921, J. O.
Martin collector (Calif. Acad. Sci.) ; Vancouver Island, Au-
gust 8, 1898, G. W. Taylor collector (E. P. Van Duzee) ;
Chilliwack, B. C., September 20, 1925.
Arctocorixa blaisdelli Hungerford, species new
Size. Length, 8.4 mm. Width of head, 2,7 mm.
Color. General facies of usual sort. Eight to ten dark bands on
pronotum. Pale bands slightly wider than the dark. Cross banding
of hemelytra entire only on base of clavus, elsewhere broken but
transverse somewhat undulate figures, the pale ones more slender
than the dark. Head and limbs pale, venter more or less dark.
Structural characteristics. Pronotum and clavus rastrate. Vertex
of head slightly higher than the eye. Facial fovea of male marked,
not attaining the eyes laterally. Metaxyphus slender. Front leg of
the male as shown in Figure 1 on Plate. A stridular patch of about
a dozen rows of pegs on base of front femur and thirty to thirty-two
pegs in the palar row. These pegs increase in size from the base.
Strigil of male rectangular and transverse, of fifteen striae, Clasper
of male as shown in Figure 2 on Plate.
Described from the following series : Seven specimens,
Vine Hill, Contra Costa County, California, July 5, 1911, col-
lected by F. E. Blaisdell. Holotype in collection of California
Academy of Science. Other specimens of this species as
follows: San Francisco, California, July 9, 1911, E. C. Van
Dyke, two specimens. Six specimens, Lagunitas, Marin
County, California, June 25, 1924, E. H. Nast, collector. Two
specimens, Berkeley, California, September 2'3, 1915, collected
by E. C. Van Dyke.
Some other records worthy of noting are as follows :
A. utahensis Hungerford, Red Lake, altitude 10,500 feet,
Fresno County, California; taken by E. C. Van Dyke (Calif.
Acad. Sci.). A. wileyi Hungerford, Steen Mountains, Harvey
County, Oregon, June 23, 1922, E. C. Van Dyke (Calif. Acad.
Sci.). Eagle Lake, Lassen County, California, July 31, 1921,
J. O. Martin (Calif. Acad. Sci.). San Diego County, Cali-
fornia, Poway Valley, April 9, 1930, C. and D. Martin. (Some
of these specimens are nearly black.)
JULY, 1930]
KEIFER MICROLEPIDOPTERA
27
CALIFORNIA MICROLEPIDOPTERA IV
BY H. H, KEIFER
Sacramento, California
These are four species of the family Gelechiidse. The first
is a small species which runs to the genus Epithectis in Mey-
rick’s key (Genera Insectorum, fasc. 184), and which does not
fit the description of any recorded species. The white sub-
apical fascia and apical markings are the apparent features of
this species.
Epithectis californica Keifer, new species
Second joint of palpi grayish fuscous, unevenly whitish on inner
side and above, and narrowly white at apex; terminal joint whitish,
extreme base fuscous and entire anterior edge and tip blackish. An-
tennae dark fuscous, somewhat whitish below, especially basally.
Head, thorax, abdomen and legs shining gray. Ground color of
forewing dull white, but entirely overlaid and almost completely
obscured by grayish fuscous. Basal half of wing light grayish fus-
cous with a few indistinct marks. Apical half distinctly darker and
contrastingly marked. Costa at extreme base dark fuscous. Fold
with a dark fuscous blotch just within the basal fourth, and another
at one-third, a white infusion of scales between them and after the
second in the fold; the second mark the larger, probably indicating
the plical stigma. Discal stigmata dark fuscous, small, indistinct,
the first at one-half, the second at the apical third. A short, out-
wardly pointed and oblique blackish mark on costa above first discal
stigma, followed by a white infusion and then the dark fuscous
edging on the inner side of the white fascia. Fascia rather narrow
(angled with apex pointing outward) from costa just beyond two-
thirds, extending obliquely outward to just below central line of
wing at apical fifth, thence abruptly running back to tornus, slightly
interrupted just below angle. Extreme apex with a white dot; evenly
spaced between this and the fascia on the costa are two short white
marks; a conspicuous black dash contiguous to the lower side of the
apical white dot; a white line in cilia around apex; cilia light fuscous.
Hindwing whitish overlaid light fuscous, giving a general gray
appearance. Expanse, to mm.
Type, male, No. 2963, Museum California Academy of
Sciences, taken near Fair Oaks, Sacramento County, California,
by the writer on September 19, 1928. Allotype, female. No.
2964, Museum California Academy of Sciences, with same
data. Forty-one paratypes also have this same locality and
date. These paratypes are in the collection of Miss Annette
28
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
F. Braun, the United States National Museum, the author’s
collection, and the Academy collection.
The male genitalia, as shown by paratypes designated in
the Academy collection and the author’s collection, are chiefly
characterized by spines around the aedceagus and the large ven-
tral plates. These ventral plates touch each other midventrally
and are emarginate apically, ending in a short midventral pro-
jection and a longer finger-like process opposite the shorter
projection, just below each harpe (see Figs, la, lb). The male
genitalia of a specimen of E. attributella Walk, furnished by
Miss Braun have been examined. In this species there is a
suggestion of the ventral plates as in calif ornica, the aedoeagus
is similar to the local species, and the harpes are about as long
though differently shaped ; but the uncus and gnathos are radi-
cally different. Specimens from Oroville and Marin County
which cannot be separated from calif ornica by wing pattern
have genitalia markedly at variance with the Sacramento
County form and therefore are not included at the present.
They may prove to be members of one quite variable species.
The next three species are of the type genus and are asso-
ciated in having the same or closely related food plants. These
plants are of the Ericacese and are Madrone (^Arbutus manziesii
Pursh.) and Manzanita (Arctostaphylos spp.). The moths
differ according to the following synopsis :
Adults
A. Almost entirely reddish or brick red; male genitalia with a
short broad uncus and harpes apparently double
Gelechia panella Busck
B. Grayish black with bluish or purplish reflections and a black
rather elongate triangle on dorsal base of forewings; male
genitalia with uncus slender and produced, harpes simple,
vinculum with large median posterior lobe
Gelechia arbutina n, sp.
C. Grayish black with bluish or purplish reflections and an out-
wardly oblique black bar across central line of wing near
base; male genitalia with uncus produced and slender, harpes
simple, vinculum with two posterior lobes
Gelechia manzanita n. sp.
Larv^
A. Yellow green with three prominent dorsal longitudinal pinkish
purple stripes; found on Manzanita and Madrone
Gelechia panella Busck
JULY, 1930] KEIFER— MICROLEPIDOPTERA 29
B. Yellow green, noticeably irrorated with fine black dots; found
on Madrone only Gelechia arbutina n. sp.
C. Yellow green with a reddish purple subdorsal longitudinal
stripe, or this stripe fuscous (sometimes faint and shortened) ;
found on various species of Manzanita..-G^/^cA:a manzanita n. sp.
Gelechia arbutina Keifer, new species
The scales of this moth are light in color at the extreme
base, almost white, but heavily pigmented dark grayish or
blackish on most of the scale, which is often tipped a lighter
shade. The general effect is deep gray with black spots and a
bluish, coppery or purplish irridescence.
Palpi with second joint dark grayish fuscous, brush large,
longer at base, irrorated whitish, the joint somewhat overlaid
whitish on inner side posteriorly, especially near base ; terminal
joint blackish fuscous, irrorated and marked whitish or ochre-
ous whitish: an annulus just above base, tip white. Head deep
grayish fuscous, face on some specimens lighter; thorax as
head above, the lighter irroration not very contrasting. Ab-
domen sordid whitish, darker toward tip and below. Forewing
the same as the head and thorax, but with more contrasting
irroration, scale tufts, and black marks, and a white subapical
fascia. A small black spot within the costa near base, often
followed by two or three more. A comparatively large deep
black triangle near dorsal base, somewhat elongate longitudi-
nally, its inner and most attenuate point touching dorsal edge
near base, its outer angle ending in the fold and partly on the
edge of a scale tuft by the fold at basal fourth, the costal angle
ending a short distance across the fold and near central line
of wing. A second tuft at basal third on disk; a third, small,
obliquely below and beyond second, on the fold. A conspicuous
moderately narrow whitish fascia, slightly curved or angled,
extends from costa, just after beginning of costal cilia, to
tornus. A large blackish area from the last two scale tufts
to the fascia, the costal edge of this area extending from well
within costa near the second scale tuft to the costa at or just
before the fascia, the dorsal edge somewhat concave but
roughly following the fold to tornus. Apical area beyond the'
white transverse fascia somewhat darker than the usual wing
color, with suggestions of longitudinal stripes, the most notice-
30
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
able directly from the fascia to just below the apex. Cilia light
gray, faintly lined parallel to wing margin. Hind wing light
fuscous, darker apically; cilia light fuscous. Legs dark fuscous
or grayish fuscous, irrorated whitish, and white at apices and
joints. Expanse, 19 to 22 mm.
Type, male. No. 2965, Museum California Academy of
Sciences, reared June 7, 1929, from a larva collected on
Arbutus menziesii Pursh (Madrone) at Mill Valley, Marin
County, California; allotype, female. No. 2966, Museum Cali-
fornia Academy of Sciences, reared June 12, 1927, from a
larva with the same data. A series of eighteen paratypes are
included, part of them reared from Mill Valley larvae and part
from Phoenix Lake (Marin County) larvae. There are also
collected adults, all from Mill Valley, part taken by Mr. E. P.
Van Duzee. These are in the collections of Miss Braun, the
National Museum, the Academy, and the author.
The male and female genitalia as shown by certain desig-
nated paratypes are diagrammed in Figs. 2a, 2b, 2c, 2d. The
main features of the male genitalia are the produced uncus,
the gnathos, the ventrad projections of the tegumen, the large
midventral posterior lobe of the vinculum, and the shape of
the aedoeagus. This is compared under the next species.
The larva is comparatively robust and approximately 17
mm. long when full grown. Body yellowish green; tubercles
indicated as small conspicuous black dots; hairs moderately
long, prominent, black; body generally becoming bright pinkish
before pupation. Thoracic shield more yellowish or ochraceous
than the body, variously shaded with black usually along the
lateral or posterior margins. Anal plate more yellowish than
body; anal fork present. Thoracic legs light yellow, dark fus-
cous on tibia and tarsi. Crochets in complete circle, heavier
outwardly, unevenly triordinal, apparently 55 to 60. Head
light brown or yellow-brown, eyes blackish brown.
This larva is typically found in a roll at the outer edge of
that portion of the leaf it is skeletonizing. This roll is securely
anchored, is open at both ends, and is made larger as the larva
grows. The leaves first attacked are those that had developed
the previous year, as the new growth has not appeared when
this caterpillar begins to feed in the early spring. The habits
JULY, 1930]
KEIFER M I CROLEPIDOPTER A
31
of the mature larvae are not known from field observations,
but they live between new leaves in the laboratory. They are
to be collected in Marin County, at Mill Valley and Phoenix
Lake, during March, April and possibly May; collecting dates
are from March 19 to April 16.
Gelechia manzanitae Keifer, new species
This species is very -similar to the foregoing and does not
need to be described in detail as most of the description would
be a reiteration of that of arbutina. The main difference in
the pattern of the fore wings is on the basal part of the wing
where instead of a black triangle there is an oblique black bar,
from well within the costa at the basal fifth it runs across the
fold and ends on the outer side of a scale tuft well within the
dorsal margin at about the basal fourth. In addition the apical
joint of the palpi is not as white, especially at the tip. Wing
expanse of the various specimens is from 15 mm. to 19.5 mm,
averaging somewhat smaller than arbutina.
Type, male. No. 2967, Museum California Academy of
Sciences, reared June 26, 1927, from a larva collected by the
writer at Phoenix Lake, Marin County, California, feeding in
a new terminal of Arctostaphylos sp. (manzanita). Allotype,
female. No. 2968, Museum California Academy of Sciences,
reared June 30, 1927, from a larva having the same data. In
addition sixty-nine paratypes are included : from Phoenix Lake,
the Marysville Buttes, Sutter County, Calif., and Big Bend
Mountain (above Las Plumas), Butte County, Calif. These
are distributed among the collections of Miss Braun, the
National Museum, the Academy, and the author.
This large series of adults from these widely separated
localities shows considerable variation in size and in larval
coloration, but the color of the adults does not vary, nor do
the genitalia to any extent. Those from Phoenix Lake average
about 18.5 mm. in wing expanse, whereas the other two groups
from the inland localities average 16.5 mm. Adults from
Phoenix Lake larvas emerged from June 15 to 30; Marysville
Butte larvae appeared as adults from May 31 to June 15; the
Big Bend Mountain individuals came out about the same time
as those from Phoenix Lake.
32 THE PAN“PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
The male genitalia of this species (Figs. 3a, 3b, 3c, 3d, 3e)
can immediately be seen to differ from that of arbutina: 1. In
the shape of the uncus and gnathos; 2. In the shorter projec-
tions from the tegumen; 3. In the two ventral lobes of the
vinculum; 4. In the shape of the aedoeagus. The male genitalia
of specimens of Gelechia scabrella Busck received from Dr.
William Barnes have been examined and proves that species
to belong to the same section of the genus as the new species,
the main differences being only in degree. Gelechia walsing-
hami Dietz and pennsylvanica Dietz seem also to belong to this
series, but can immediately be separated by the pattern of the
forewings.
The larvae of manzanitce were collected May 14 to 30 at
Phoenix Lake, April 20 to May 11 at the Marysville Buttes,
and May 23 to 28 on Big Bend Mountain. At least three spe-
cies of Arctostaphylos are attacked in this range. On April 26
the larvae were found skeletonizing the old leaves, as the new
growth had not yet appeared. The mature larvae are found in
spune heads of new growth, eating holes in the leaves.
A Phoenix Lake larva when full grown and not yet ready^
to pupate is from 13 to 14.5 mm. long; body light yellow green,
with a subdorsal reddish or reddish purple longitudinal stripe
from the thoracic shield to the ninth abdominal segment; a few
of the body tubercles slightly fuscous ; hairs moderate in length,
brownish fuscous; thoracic shield brown and blackish laterally,
or black with a narrow median line ; crochets in complete circle,
heavier outwardly, 45 to 47, unevenly triordinal. Head dark
brown to black.
The Marysville Butte larvae are smaller in average and
about the same in color except that the subdorsal stripe is indi-
cated by an almost faint fuscous color. The Big Bend Moun-
tain larvae are, if anything, still smaller, some only 12 mm. long
when full grown; the head and shield is never black and the
subdorsal stripe is only shortly indicated on part of the indi-
viduals by faint fuscous shading.
The pupae of manzanitce and arbutina are very similar; that
of manzanitce being 6.5 to 8 mm., and arbutina 8 to 8.5 mm.
They are dark brown, robust, the usual hair fringe on the
34 THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
seventh segment, and no movable segments. In manzanitcB 'the
antennae, wings and legs end near the posterior margin of
the fifth abdominal segment, while in arbutina they end about
the middle of the segment.
Gelechia panella Busck
This species appears here by virtue of its food plants. An
adult was obtained from Manzanita in Marin County during
1927, but the larva was not noted until specimens were taken
from both Manzanita and Madrone on Big Bend Mountain,
May 23 and 28, 1928.
The length of the full-grown larva of this species is approxi-
mately 16 mm. Head light yellow or yellow-brown ; body
yellow-green with three dorsal longitudinal (a dorsal and a
subdorsal) rather wide pinkish purple stripes. The larva
appears with the new growth in the spring considerably after
the appearance of the two above-described species, and feeds
on both Manzanita and Madrone. The adult is interesting in
that it displays the reddish coloring common to a number of
Manzanita insects, which include members of the Coleoptera
and Hemiptera. The male genitalia would place panella in
another section of the genus than that of arbutina and manza-
nitcB. The distribution in California of this species as shown
by specimens in the Academy collection is : Alma, Santa Cruz
County (J. O. Martin, collector) ; Mill Valley, Marin County
(E. P. Van Duzee and M. C. Van Duzee, collectors) ; Phoenix
Lake, Marin County (reared by the writer) ; and Big Bend
Mountain, Butte County (reared by the writer). The adults
taken on the wing were collected from March to October, prov-
ing the species to overwinter in the adult stage. The reared
specimens have emergence dates from June 22 to June 29.
Explanation of Plate
la, Epithectis calif ornica, lateral view of the male genitalia,
lb, same, composite dorsal and ventral view of the male genitalia,
ventral view on right, 2a, Gelechia arbutina, male genitalia, 2b, same,
side view of uncus, gnathos, and tegumen of male genitalia, 2c, same,
basal part of the female genitalia. 2d, same, signum of female geni-
talia. 3a, Gelechia manzanita, male genitalia. 3b, same, variation in
tip of harpe. 3c, same, side view of uncus, gnathos, and tegumen of
male genitalia. 3d, same, basal part of female genitalia. 3e, same,
signum of female.
JULY, 1930] VAN DUZEE GENUS SYMPYCNUS 35
THE DIPTEROUS GENUS SYMPYCNUS LOEW IN
NORTH AMERICA AND THE WEST INDIES
BY M. C. VAN DUZEE
Buffalo, Ne<iv York
The described species of Sympycnus and its subgenus
Calyxochaetus are distributed over the whole world. Including
those described in this paper, 156 species have been recognized.
These are distributed about as follows : European 12 ; Oriental,
including most of Asia, 32; North America 24 and 12 species
of the subgenus Calyxochaetus ; Central America 10 and 2 spe-
cies of Calyxochaetus; South America 47 and 2 species of
Calyxochaetus; Africa 9 and one species of Calyxochaetus; Aus-
tralia and New Zealand 6 species. In South America, below
the 38 degrees south latitude, nearly half of the known species
of the Dolichopodidae belong to this genus ; this is probably
partly due to ecological conditions in the regions where most
of the collecting has been done.
The genus Sympycnus was established by Dr. Hermann
Loew in 1857, Neue Beitr., v, p. 42, for five species; generic
type designation by Coquillett in 1910, Porphyrops annulipes
Meigen (Proc. U. S. Nat. Mus., Vol. xxxvii, p. 610). The
genus Calyxochaetus was erected by J. Bigot in 1888, Ann.
Soc. Ent., France, Ser. 6, viii. Bulletin p. xxiv, 1888, and Ann.
Soc. Ent., France, Ser. 6, x, p. 282, 1890, with type Sympycnus
nodatus Loew.
Synonyms: Nothosympycnus Wheeler, Proc. Calif. Acad.
Sci., Ser. 3, Zool., Vol. ii, pp. 47, 51, 1899, type Sympycnus
nodatus Loew; Gymnoceromyia Bigot, Ann. Soc. Ent., France,
Ser. 6, X, pp. 276, 293, 1890, type of genus and only species
Gymnoceromyia andicola Bigot.
As Nothosympycnus includes the type species of Calyxo-
chaetus, there can be no question as to its being synonymous;
Gymnoceromyia has been considered a synonym of Psilopus,
but lately I described a species as a Sympycnus (Calyxochaetus)
from Chile, but when the type was returned to the British
Museum Mr. Edwards and Mr. Collin compared it with the
type of Gymnoceromyia andicola Bigot and found them the
same. As it is a true Calyxochaetus, we must place Gymno-
ceromyia also as a synonym of Calyxochaetus. This subgenus
36
THE PAN-PACIFIC ENTOMOLOGIST . [vOL. VII, NO. 1
is separated only by having the first joint of fore tarsi shorter
than fifth, and not or but little longer than wide, but it is
shortened in the male only; the male also has the tip of the
arista enlarged in some species, but as we have a South Ameri-
can species with three enlargements on the arista and the first
joint of the fore tarsi longer than the second, it seems necessary
to make this a subgenus if we can give it even that rank.
The genus Sympycnus is characterized as follows: Face narrow
or somewhat narrowed in the male, wider in female, usually a little
wider above; first antennal joint bare above, third joint usually
small, triangular, pointed at tip with the arista inserted near the
base on upper edge; legs and feet slender, tibiae with rather small
bristles, sometimes almost without bristles; pulvilli often enlarged
in the male; abdomen elongated, cylindrical, somewhat compressed
in the male, tapering to a point in the female; hypopygium of male
more or less embedded, small, its appendages usually small, tri-
angular or rod-like; dorsum of thorax with a more or less distinct
flattened area on posterior slope, before the scutellum; acrostichal
bristles usually in a single row, rarely biserial or even wanting;
abdomen with six segments in the male, sixth small and usually not
visible on the venter; female with five visible segments; wings rather
long and narrow, sometimes much narrowed at base in male, usually
wider in the female with the anal angle more prominent; posterior
cross vein at or a little before the middle of the wing; third and
fourth veins parallel toward their tips, sometimes parallel from the
cross vein, but usually the last section of fourth vein is more or less
bent at or before its middle, fourth vein ending in or a little before
the apex of the wing; sixth vein not reaching the wing margin,
sometimes represented by a slight fold in the wing.
Some species vary considerably from the characters given
above. Several have the third antennal joint large or the arista
inserted near the middle of upper edge; in a few the abdomen
is shorter and formed more as in the Chrysotus and in several
the hypopygial appendages of the male are very long and
narrow, nearly as long as the abdomen.
I am greatly indebted to Dr. J. M. Aldrich for the loan of
material from the United States National Museum, from which
several species new to me are described.
Key to the Males of the North American Species of
Sympycnus, Including Mexico and the West Indies
1. Species with all tarsi plain, except sometimes in color or with
a few longer hairs on last joint 2
JULY, 1930] VAN DUZEE— GENUS SYMPYCNUS
37
2 .
3.
4.
5.
6 .
7.
8 .
9.
10 .
11 .
12 .
13 .
Species with one or more pairs of tarsi modified, or with
unusual hairs or bristles, sometimes only the last joint a
little widened 13
Antennae wholly black or brown, sometimes first joint a little
yellow below 3
Antennae yellow or brownish yellow, at least first joint yellow 8
Fore coxae black with yellow tips (California; Alaska)
cuprinus Wheeler
Fore coxae wholly, or almost wholly yellow 4
Dorsum of thorax with a broad, median, violet stripe (Ontario;
New York) canadensis Van Duzee
Thorax without such a stripe 5
Middle and hind coxae black with yellow tips; first and second
joints of hind tarsi of nearly equal length (Mexico)
varipes Aldrich
All coxae wholly yellow; first joint of hind tarsi shorter than
second 6
Dorsum of thorax with distinct lines; third antennal joint
twice as long as wide (Eastern States to Kansas)
lineatus Loew
Dorsum of thorax without lines; third antennal joint short,
not longer than wide 7
Dorsum of thorax dull; third antennal joint as long as wide;
hypopygial lamellae short and stout; abdomen rather short
(California; Washington) breviventris new species
Dorsum of thorax bright shining green; third antennal joint
short; hypopygial lamellae long and narrow; abdomen
long as usual (St. Vincent, W. I.) thoracicus new species
Basal half of first joint of hind tarsi white, apical half black;
hind tibiae enlarged at tip, which is sharply black (Colo-
rado; New Mexico; Arizona) clavatus Van Duzee
Hind tarsi not black and white; hind tibiae scarcely enlarged
at tip, which is sometimes a little brown 9
Humeri yellow (Afexico) angustipennis Aldrich
Humeri metallic or brown, sometimes a little yellow below 10
Hind tarsi with first joint only a little shorter than second
(California) lamgatus new species
First joint of hind tarsi much shorter than second 11
Face golden yellow (Alaska) aurifacies Van Duzee
Face not yellow, usually white 12
Dorsum of thorax with distinct lines (Eastern States to
Kansas) lineatus Loew
Dorsum of thorax without lines (California)
inaqualh new species
Only one pair of tarsi ornamented ....14
More than one pair of tarsi modified or with unusual hairs or
bristles ^..... 25
38
THE AN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
14. Fore tarsi modified dr with long hairs or bristles, other tarsi
plain IS
— Middle tarsi with long hairs or modified, other tarsi plain 19
— Hind tarsi a little compressed and widened, third joint with a
prominent spine at tip or with long hairs 20
15. Second joint of fore tarsi dilated on underside of basal half or
more; hypopygial lamellae very long filaments, fringed
with long hairs (California) caudatus Van Duzee
— Second joint of fore tarsi not swollen below; hypopygial
lamellae short 16
16. First two antennal joints yellow, third black (California)
lavigatus new species
— Antennae wholly black, or first joint yellowish below 17
17. Fore tarsi gradually, but considerably widened from base to
near tip of second joint, then narrowing again to normal
at tip (Utah) latitarsis new species
— Fore tarsi not widened in the middle, if at all 18
18. Fifth joint of fore tarsi a little widened, their pulvilli very
large (Colorado) pulvillus new species
— Second joint of fore tarsi with a spur near tip, and several
small bristles on other joints, fifth joint not at all widened,
their pulvilli large (California) breviventris new species
19. Last two joints of middle tarsi widened, black (Idaho)
montanus new species
— Second joint of middle tarsi with three long hairs (California)
L tripilus new species
20. First two joints of antennae yellow, third sometimes blackish;
third joint of hind tarsi fringed with long hair (Alaska)
tertianus Loew
— Antennae yellow, tip of third joint brown; second joint of hind
tarsi with one wavy hair, which is nearly as long as the
joint (Indiana) aldrichi new species
— Antennae wholly black or brown 21
21. Third joint of hind tarsi fringed with long hairs, fourth with
several hairs which are shorter (Wyoming; Alberta)
marcidus Wheeler
— Third joint of hind tarsi with a prominent spine at tip 22
22. Hypopygium with elongate, whitish lamellae and two or three
long, black, hair-like filaments 23
— Hypopygial appendages wholly black 24
23. Third joint of hind tarsi shorter than fourth; hypopygial
lamellae very large (Colorado; New Mexico)
calcaratus new species
— Third joint of hind tarsi as long or longer than fourth; hypo-
pygial lamellae very short (Victor, Colorado)
setosus new species
JULY, 1930] . VAN, PUZEE— GENUS SYMPYCNUS
39
24. Fore coxae almost wholly black, with black hair; hypopygium
round, unusually conspicuous (Colorado)
globulicauda new species
— Fore coxae almost wholly yellow with pale yellow hairs; hypo-
pygium small, partly concealed (California)
fasciventris Van Duzee
25. All tarsi modified or with unusual hairs or bristles (California)
pugil Wheeler
— Apical joints of fore tarsi slightly widened; third joint of hind
tarsi with a large thorn at tip; fore coxae, all femora and
tibiae wholly yellow (California) fasciventris Van Duzee
S5mipycnus aldrichi Van Duzee, new species
Male. Length, 2 mm. Face green, quite wide above, eyes touch-
ing below; palpi and proboscis yellow; antennae (Fig. 40) yellow,
third joint brown at tip, triangular, as wide as long, arista appears
yellow in certain lights, brown in others; front dull brownish.
Dorsum of thorax green, dulled with gray pollen; lower part of
thorax yellow; abdomen yellow, apical part and the hypopygium
shining brownish black; hypopygial appendages very small.
Coxae, femora, tibiae and tarsi yellow, with very small hairs and
bristles; apical joints of tarsi and tips of some of the other joints
blackish; hind tarsi (Fig. 41) with long wavy hair at tip of second
joint; joints of fore tarsi as 15-5-5-4-5; of middle ones as 26-12-11-6-6;
joints of posterior pair as 18-13-14-9-8. Calypters, their cilia and the
halteres yellow.
Wings grayish; third and fourth veins nearly straight and parallel
beyond the cross vein, fourth ending in the apex of the wing; last
section of fifth vein as 20, cross vein 11.
Described from one male taken by J. M. Aldrich, July 21,
1915, at La Fayette, Indiana. Type in the United States
National Museum.
Sympycnus angustipennis Aldrich
Biologia, Diptera, Vol. i, 344, 1901. Mexico.
Male. Length, 2.1 mm. Face very narrow below; front purple
in the middle; humeri and posterior margin of pleurae yellow;
abdomen with yellow on the sides near the base; hypopygium with
yellow appendages; coxae, femora and tibiae yellow, tarsi slightly
infuscated; first joint of middle tarsi as long as the remaining four
taken together; pul villi not enlarged; calypters, their cilia and the
halteres yellow.
Sympycnus aurifacies Van Duzee
Ohio Journal of Science, Vol. xxiii, 248, 1923. Alaska.
Male. Length, 1.7 mm. Face narrow, golden yellow; front blue
or violet; first two antennal joints orange yellow, third black, twice
40
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
as long as wide; palpi small, yellowish; dorsum of thorax greenish
brown, with a median green line and a poorly defined purple line
each side of it; scutellum blue or violet, green on the edge; fore
coxae, all femora and tibiae wholly yellow; fore coxae with a few
small yellow hairs; all tarsi black from tip of first joint, anterior
ones one and a half times as long as tibia, first joint nearly as long
as two following taken together; hind basitarsus hardly as long as
the second joint; last section of fifth vein twice as long as cross
vein; calypters, their cilia and the halteres yellow.
Sympycnus breviventris Van Duzee, new species
Male. Length, 2 mm. Face very narrow; face and front grayish
pollinose; palpi black with white pollen; antennae (Fig. 3) black,
seen from below the first joint is yellow, from above it appears
wholly black; occiput green with white pollen; lateral and inferior
orbital cilia white.
Thorax greenish; dorsum covered with brown pollen, without
lines; acrostichal bristles few and very small; pleurae white-pollinose,
its posterior edge more or less yellow. Abdomen green, compressed;
first segment yellow at base, second and third shining yellow on
sides, leaving a median stripe on dorsum and the hind margin green;
hairs of abdomen black; hypopygium (Fig. 5) blackish, very small,
with a pair of yellow lamellae; venter yellow, its last segment black
with a pair of large black bristles.
Coxae yellow, posterior ones sometimes a little blackened; anterior
pair with minute yellow hairs; femora and tibiae yellow; fore tibiae
without bristles, middle and hind ones with a few small bristles;
tarsi yellow, more or less infuscated toward their tips; fore tarsi
(Fig. 4) with a spur at tip of second joint and with several small
bristles on other joints, sometimes these are apparently wanting,
second joint and tip of first appear slightly thickened when viewed
from the side; joints of fore tarsi as 23-9-8-6-6; of middle ones as
30-14-10-7-5; joints of posterior pair as 16-24-14-8-7. Calypters
yellow with black tips and cilia; halteres yellow.
Wings a little grayish; third and fourth veins nearly straight and
parallel beyond the cross vein, fourth ending in the apex of wing;
last section of fifth vein as 39, cross vein as 15; anal angle of wing
quite prominent.
Female. Face wider, color as in the male; abdomen tapering;
joints of hind tarsi as 20-21-13-7-8; palpi yellow; venation as in the
male.
Described from seven males and seven females, holotype,
allotype and seven paratypes taken by the author, April 17,
1915, in San Diego County, California, on the edge of the
JULY, 1930] VAN DUZEE GENUS SYMPYCNUS 41
Colorado Desert; the other five paratypes were taken by J. M.
Aldrich, May 7-9, 1906, at Pacific Grove, California.
Holotype and allotype in author’s collection.
Sympycnus calcaratus Van Duzee, new species
Male. Length, 3-3.5 mm. Face silvery white, narrow below;
front green, dulled with gray pollen; palpi brown or yellowish
brown; antennse (Fig. 23) black, third joint longer than wide,
triangular.
Thorax green, dulled with gray pollen; acrostichal bristles repre-
sented by several small hairs before the suture; pleurae white-polli-
nose; abdomen green, base of segments bronze or coppery, its hairs
mostly black; hypopygium (Fig. 25) black; outer lamellae large,
yellow; inner appendages a pair of small, rounded, black lamellae,
fringed with small hairs.
Coxae and most of fore femora black; tips of coxae, base and
broad tips of fore femora, middle and hind femora, trochanters and
tibiae yellow, upper edge of posterior femora sometimes brown; tarsi
blackened from tip of first joint; fore tibiae without bristles, those
of middle and hind tibiae moderately strong; fore and middle tarsi
plain, with tips of second, third and fourth joints a very little thick-
ened; hind tarsi (Fig. 24) with third and fourth joints a little
thickened, third with a stout, curved spur at tip, which is a little
shorter than the joint; joints of fore tarsi as 23-10-6-5-6; of middle
ones as 37-15-11-8-7; those of hind ones as 25-18-8-11-8. Calypters
mostly brownish with yellow cilia; halteres yellow.
Wings grayish; third vein bent backward a little toward tip; last
section of fourth vein bent near basal third, nearly parallel with third
beyond this bend, its tip distinctly before apex of wing; last section
of fifth vein as 50, cross vein as 15; anal angle prominent but broadly
rounded; sixth vein distinct, not reaching wing margin.
Female. Face wide, silvery white, its sides nearly parallel; palpi
yellow, white-pollinose; third antennal joint as long as wide; color
of front, thorax, abdomen, legs and feet about as in the male; all
tarsi slender, plain; joints of fore tarsi as 29-12-9-6-6; of middle ones
as 40-17-13-8-6; those of posterior pair as 33-23-15-11-8; venation as
in the male.
Described from seven males and two females ; holotype,
allotype and two paratypes were taken by the author June 9,
1915, at Colorado Springs, Colorado; four male paratypes were
taken at Cloudcroft, New Mexico, June 18 and 19, 1902; the
female paratype at the same place on June 21. Type in author’s
collection.
42
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VH, NO. 1
Sympycnus CANADENSIS Van Duzee
Canadian Entomologist, Vol. xlix, 339, 1917. Ontario; New
York.
Male. Length, 2 mm. Face rather wide, silvery white; antennae
black, third joint triangular, as long as wide; dorsum of thorax with
a broad, median, violet stripe, more coppery on the sides and with
large, velvety black spots above root of wings; second and third
abdominal segments mostly yellow; hypopygium (Fig. 76) mostly
concealed and with black lamellae; coxae and legs yellow; upper edge
of hind femora on apical half, whole of hind tibiae and tarsi and last
two joints of fore and middle tarsi black; joints of fore tarsi as
16-7-6-4-5; of middle ones as 28-12-8-7-5; joints of posterior pair as
16-13-9-6-7; calypters, their cilia and the halteres yellow; last section
of fifth vein as 27, cross vein as 9.
Sympycnus caudatus Van Duzee
Canadian Entomologist, Vol. xlix, 338, 1917. California.
Male. Length, 2-3 mm. First two antennal joints yellow, third
black (Fig. 13) scarcely as long as wide; eyes contiguous; front and
thorax bronze brown; abdomen a little yellow at base; hypopygium
brown with very long, yellow outer appendages, which are fringed
with long hairs; fore coxae, all femora, tibiae and tarsi yellow, some-
times the tarsi are slightly infuscated from tip of first joint, but
usually almost wholly pale yellow; fore tarsi as in Figure 14; joints
of fore tarsi as 20-7-6-6-6; of middle ones as 29-17-9-7-5; joints of
posterior pair as 15-21-12-9-7; fore pulvilli enlarged; last section of
fifth vein as 35, cross vein as 11; calypters, their cilia and the halteres
yellow.
Sympycnus clavatus Van Duzee
Entomological News, Vol. xxiv, 271, 1913. New Mexico;
Arizona; Colorado.
Male. Length, 2.5-3 mm. Eyes contiguous; first two antennal
joints yellow (Fig. 7), third small, brown; thorax dark green, humeri
yellowish; abdomen largely yellow, hairs and bristles on first seg-
ment yellow; hypopygium blackish; all coxae, femora, tibiae and fore
(Fig. 8) and middle tarsi yellow, last two or three joints of these
tarsi black; hind tibiae enlarged toward tip with a sharply defined,
wide, black tip; hind tarsi scarcely half as long as tibia with basal
half of first joint white; joints of fore tarsi as 22-6-5-5-7; of middle
ones as 35-16-11-7-7; of posterior pair as 26-16-14-10-8; last section
of fifth vein as 20, cross vein as 15.
43
JULY, 1930] VAN DUZEE GENUS SYMPYCNUS
Sympycnus cuprinus Wheeler
Proc. Calif. Acad. Sci., Ser. 3, Zool., Vol. ii, 50, 1899. Western
States; Alaska.
Male. Length, 2,5-3 mm. Antennae (Fig. 1) black; face and
palpi covered with silvery white pollen; front dull black; thorax
dark green with brown pollen; abdomen green, without yellow on
sides; coxae black with yellow tips, anterior pair with several silvery
white hairs; basal half or more of fore femora and upper edge of
hind femora blackish, remainder of femora yellow; tibiae yellow,
tips of hind ones black; fore and middle tarsi from tip of first joint
and hind ones wholly black; all tarsi plain; fore pulvilli enlarged;
joints of fore tarsi as 19-8-8-7-7; of middle ones as 31-14-8-8-5; of
posterior pair as 23-20-12-7-7; calypters, their cilia and the halteres
yellow, the cilia appearing dark in certain lights; last section of fifth
vein as 32, cross vein as 12,
Sympycnus fasciventris Van Duzee
Canadian Entomologist, Vol, xlix, 337, 1917. California; Ore-
gon; Colorado.
Male. Length, 3 mm. Eyes contiguous; lower orbital cilia black,
short; antennae (Fig. 31) black; thorax brown, nearly opaque, with
gray pollen, bristles inserted in brown dots; abdomen green or cop-
pery with apical margins of segments broadly covered with gray
pollen; sometimes this portion of the segments is more green when
anterior part is coppery; fore coxae yellow with white hair; femora
and tibiae yellow; fore and middle tarsi almost wholly yellow; hind
tarsi (Fig. 32) black from tip of first joint; fore pulvilli enlarged;
joints of fore tarsi as 19-8-5-4-7; of middle ones as 40-15-10-7-6;
joints of posterior tarsi as 25-18-8-10-6; last section of fifth vein
as 41, cross vein as 12; fourth vein ending in apex of wing.
Sympycnus globulicauda Van Duzee, new species
Male. Length, 2.7 mm. Eyes touching below, leaving a small
grayish white triangle above; palpi yellowish brown with white
pollen; front dark green, dulled with gray pollen; antennae (Fig. 28)
black, third joint fully twice as long as wide.
Thorax green; dorsum with brownish pollen and three, broken,
shining brownish stripes along the rows of bristles; acrostichal
bristles few and very small; pleurae more black, white-pollinose;
abdomen green with bronze reflections, its hair black; hypopygium
(Fig. 30) large and globular, its outer lamellae brownish black,
fringed with small hairs.
Coxae black, anterior ones with a few black hairs, which are
longer and more bristle-like at tip; fore femora blackened on basal
44
THE PAN -PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
half, yellow at extreme base and on apical portion; trochanters,
middle and hind femora, all tibiae and base of all tarsi yellow, all
tarsi more or less blackened from tip of first joint; fore tibiae without
bristles; middle tibiae with three rather large ones; hind tibiae with
several smaller bristles; hind tarsi (Fig. 29) with three bristles on
side of first joint, second with two bristles, third joint with several
larger ones on the side and a strong one at tip, fourth joint thickened
when seen from the side; fore and middle tarsi plain; pulvilli of all
tarsi a little enlarged; joints of fore tarsi as 18-9-6-5-5; of middle
ones as 25-11-14-11-10; joints of hind tarsi as 22-17-8-11-6. Calypters
with black cilia; halteres yellow.
Wings grayish; last section of fourth vein bent near basal third,
nearly parallel with third at tip, reaching the wing margin before
the apex of wing; last section of fifth vein as 42, cross vein as 12;
anal angle of wing rather prominent.
Described from one male, taken by the author June 9, 1915,
at Colorado Springs, Colorado. Type in author’s collection.
Sympycnus inaequalis Van Duzee, nev^ species
Male. Length, 2 mm. Head in the type shrunken so the eyes
touch; palpi yellowish, almost yellowish brown; antennas (Fig. 11)
with first two joints yellowish, third black, triangular, considerably
longer than wide.
Thorax blue-green, dorsum with brown pollen; acrostichal bristles
few but long; abdomen blackish green, base yellowish, especially
first segment, its hairs mostly yellowish, first segment with long
yellow bristles on sides; hypopygium (Fig. 12) shining black with
moderately long, pointed lamellae, which are yellow at base and have
long hairs on sides and a hair at tip as long as the lemallae.
Coxae, femora and tibiae yellow, middle, hind and upper surfaces
of hind femora brownish, hind tibiae more brown and slightly en-
larged on apical half, but all these appear more yellow in certain
lights; tarsi brown, fore and middle pairs appear yellow when seen
in the right light; anterior and posterior tarsi plain, first three joints
of middle tarsi plain (last two joints missing in type); joints of fore
tarsi as 36-17-19-8-7; first three joints of middle tarsi as 42-19-13;
of posterior ones as 19-32-13-11-7; pulvilli of fore tarsi not or scarcely
enlarged. Calypters, their cilia and the halteres yellow.
Wings grayish; fourth vein ending in apex of wing; last section
of fifth vein nearly straight, twice as long as cross vein; anal angle
of wing quite prominent.
Female. Five females that probably are of this species were
taken with the male but differ considerably from it: Face wide,
gray-pollinose; antennae wholly black, third joint as long as wide;
hind tibiae yellow, scarcely darker at tip; hind femora black on apical
fourth; first joint of hind tarsi nearly as long as second. They agree
JULY, 1930] VAN DUZEE GENUS SYMPYCNUS
45
with the male in having the last section of fifth vein straight and
twice as long as cross vein; third and fourth veins straight and
parallel, fourth ending in apex of wing; all coxae yellow, and pos-
terior margin of pleurae a little yellowish below. Dorsum of thorax
in females shining brownish with quite distinct lines, in male bluish
without lines.
Described from one male and five females, taken by the
author at Konocti Bay, Clear Lake, Lake County, California,
May 10, 1926. Type in the author’s collection.
Sympycnus laevigatus Van Duzee, new species
Male. Length, 3 mm. Face very narrow, eyes touching in middle
of face (head shrunken) leaving a long, white triangle above and
below; palpi rather large, yellow; front and occiput blackish green,
white-pollinose; antennae (Fig. 9) with first two joints yellow, third
joint blackish, a little yellowish at base, very small, rounded; lower
orbital cilia white.
Thorax green, dulled with white pollen, viewed obliquely the
pollen on the middle of the dorsum is more brownish, forming three
distinct brown lines, the middle one reaching the scutellum; humeri
below and the posterior edge of pleurae yellow; acrostichal bristles
not distinguishable in type; dorsum of abdomen shining black,
second and third segments yellowish brown, lighter in color at base
of second, becoming more black on hind margin of third; hairs of
abdomen black, the long ones on lower edge of first and second
segments yellow; venter yellow; hypopygium shining black, its outer
lamellae short, narrow, yellow, inner appendages black.
Coxae, femora and tibiae yellow; hind femora slightly brown
above on apical half; fore coxae with small white hairs on anterior
surface and yellow bristles at tip; fore and middle tarsi yellow with
last joint black, seen in certain lights some of the other joints appear
infuscated; hind tarsi black from tip of first joint; fore tarsi (Fig. 10)
with long hair and with the joints a little thickened, their pulvilli
enlarged; fore tarsi with the joints as 22-9-7-5-7, pulvilli as 4; joints
of middle tarsi as 36-17-14-8-6; of hind ones as 23-25-16-11-10.
Calypters 3 ^ellow with black tip and yellow cilia; halteres yellow.
Wings grayish, darker in front; third vein a little bent backward
at tip; last section of fourth vein without a bend, nearly parallel
with third, ending in apex of wing; last section of fifth vein as 24,
cross vein as 15; wings narrowed at base.
Female. Face wide, black with gray pollen; palpi brownish;
antennae as in male; no acrostichal bristles; abdomen metallic
coppery black with large yellow spots on sides of second and third
segments and small ones on sides of first; legs and feet colored as
in the male; joints of posterior tarsi as 28-30-20-11-10; wings more
46
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 1
tinged with brown; venation as in the male; last section of fifth vein
as 22, cross vein as 25; wings less narrowed at base.
Described from one pair, taken by the author. May 7 and 8,
1915, at Berkeley, California. Types in the author’s collection.
Sympycnus latitarsis Van Duzee, new species
Male. Length, 2 mm. Face, front, occiput and palpi black; face
very narrow with sides parallel below the middle and a little grayish
brown pollen; antennae (Fig. 15) black, first joint rather long, third
small, somewhat triangular, as long as wide; lower posterior orbits
with a few small, white cilia.
Thorax green; dorsum with a little brown pollen and indistinct,
broken, brown lines along the rows of bristles; acrostichal bristles
few and small; abdomen greenish black, second and third segments
largely yellow on the sides and venter; hairs of abdomen black;
hypopygium large, shining black with short pale hairs, (It is broken
off, but still adheres to the abdomen in the only specimen I have.)
Fore coxae yellowish, almost brown; middle and hind ones black;
femora and tibiae dark yellow, fore femora somewhat blackened at
base; all tarsi yellow at base, blackened from the tip of first joint;
fore tarsi (Figs. 16 and 17) flattened, gradually widened from base
to near the tip of second joint, then becoming smaller, last joint
nearly normal; first joint with a long bristle at tip that is a little
shorter than second joint; second joint with three large bristles that
do not show when viewed from above the broad upper surface;
joints of fore tarsi as 20-11-7-5-6; of middle ones as 27-14-10-6-6;
joints of posterior pair as 18-20-13-9-7, Halteres yellow.
Wings grayish, slightly darker in front; third and last section
of fourth vein nearly straight and parallel, fourth ending in the apex
of the wing; last section of fifth vein as 35, cross vein as 12.
Described from one male, taken by the author June 7, 1915,
at Wendover, Utah. Type in the author’s collection.
Sympycnus lineatus Loew
Neue Beitr., Vol. viii, 67, 1861 ; Mon. N. Am. Diptera, Vol. ii,
189, 1864. Eastern States to Kansas.
Length, 2-3 mm. Antennae (Fig. 2) more or less yellow or yel-
lowish; dorsum of thorax grayish brown with darker lines; abdomen
brownish gray to yellowish brown; hypopygium black with narrow,
straight, rather long, brown appendages projecting downward; coxae,
legs and feet yellowish; joints of fore tarsi as 22-9-6-5-5; of middle
ones as 33-14-11-6-5; of posterior pair as 17-22-15-14-7; third and
fourth veins nearly straight and parallel, fourth ending in apex of
wing; last section of fifth vein as 22, cross vein as 10.
47
JULY, 1930] VAN DUZEE GENUS SYMPYCNUS
Sympycnus marcidus Wheeler
Proc. Calif. Acad. Sci., Ser. 3, Zool., Vol. ii, 48, 1899. Wy-
oming; Alberta.
Male. Length, 2.5-3 mm. Antennas (Fig. 33) small, black, third
joint as long as wide; face very narrow, white; palpi minute, yellow;
thorax dull green; abdomen green with coppery reflections; venter
yellow; hypopygium shining black with a rather long, straight, yel-
lowish appendage extending downward; coxae yellow, middle and
hind ones infuscated on outer surface, fore coxae with white hairs;
femora and tibiae yellow; fore tarsi (Fig. 34) shorter than their
tibiae; fore pulvilli white, enlarged; hind tarsi (Figs. 35, 36) blackened
from tip of first joint; joints of fore tarsi as 13-6-5-4-6; of middle
ones as 38-17-13-8-7; of posterior pair as 27-20-14-8-7; fourth vein
ending in apex of wing; last section of fifth vein as 43, cross vein
as 10; calypters and halteres yellow, cilia of former white; hind tibiae
and tips of hind femora sometimes brown.
Sympycnus montanus Van Duzee, new species
Male. Length, 2.2 mm. Eyes contiguous below (perhaps from
the head being shrunken), leaving a gray triangle above; front and
occiput dull blackish; palpi yellow; antennae black, third joint small,
triangular, arista dorsal.
Dorsum of thorax green with brown pollen; pleurae white-polli-
nose, posterior edge of pleurae and spot at root of wing yellowish;
a scarcely visible depressed area before the scutellum; abdomen
shining black with a reddish yellow tinge on basal half; venter
yellow at base; hairs on dorsum of abdomen black, on sides at base
yellowish; hypopygium black, its small outer lamellae black with
several long hairs at tip; inner appendages more testaceous.
Coxae, femora and tibiae yellow; fore coxae with a few yellow
hairs; fore tarsi yellow at base, blackened from second joint; middle
tarsi (Fig. 42) yellow with last two joints black and widened; hind
tarsi black from tip of first joint; joints of fore tarsi as 17-7-5-4-6;
their pulvilli enlarged; joints of middle tarsi as 28-13-11-5-6; of
posterior pair as 17-17-12-8-7. Calypters, their cilia and the halteres
yellow.
Wings grayish, narrowed a little at base; last section of fourth
vein straight, nearly parallel with third, ending in the apex of the
wing; last section of fifth vein as 23, cross vein as 15.
Described from one male taken by J. M. Aldrich, June 4,
1910, on Mount Moscow, Idaho. Type in the United States
National Museum.
THE PAN-PACIFIC ENTOMOLOGIST
Published quarterly by the Pacific Coast Entomological Society
in cooperation with the California Academy of Sciences
E. P. Van Duzee, Editor S. B. Freeborn, Treasurer
EDITORIAL COMMENT
Several members of the Pacific Coast Entomological So-
ciety have been in the field again during the past season. Mr.
J. O. Martin and Mr. Gorton Linsley went by auto to Browns-
ville, Texas, stopping for a little work in Arizona and New
Mexico. Their work will add several thousand insects of the
southern Texas fauna, which has not been as well represented
as it should be in the collection of the Academy. Dr. E. C.
Van Dyke covered some interesting localities in a circuit
including Salt Lake City, the Grand Teton, Yellowstone and
Glacier National Parks and Steven’s Pass in the Cascade
Mountains in Oregon, altogether adding about three thousand
insects to the Academy collections. Mr. J. E. Cottle made two
or three short trips for Lepidoptera, the most important of
which was to Modoc County, California, and a number of
similar collecting trips within California were made by Mr.
Sternitzky. Mr. Robert Usinger and Robert Wind were mem-
bers of a Boy Scout party under the direction of Dr. Ansel
Hall, that visited the Grand Canyon of the Colorado and sev-
eral interesting localities in the Rocky Mountains. Both were
able to make some valuable additions to their collections. Your
editor did some work in the “Mother Lode Country” in the
Sierra foothills, adding about 1800 insects to the Academy
collection, and Dr. F. E. Blaisdell took some interesting ma-
terial during a short stay in the Yosemite Valley. During last
winter Mr. Theo O. Zschokke and Mr. Richard Blackwelder
collected for a time about Gatun Lake, Panama, Canal Zone.
The Hemiptera came to the Academy collection and made a
most welcome addition, as the representation of the Biologia
fauna already there was comparatively meagre.
Among the welcome visitors at the Entomology Department
of the California Academy of Sciences during the summer holi-
days may be mentioned: S. A. Rohwer, of the Department of
Agriculture, Washington ; R. H. Painter, Agricultural College,
Manhattan, Kansas ; Dwight M. De Long, Ohio State Univer-
sity, Columbus, Ohio, and B. Prashad, director. Zoological
Survey of India, Calcutta, India.
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For Proceedings of the California Academy of Sciences address
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Vol. VII
October, 1930
No. 2
THE
Pan-Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
VAN DUZEE, M. C., THE DIPTEROUS GENUS SYMPYCNUS IN NORTH AMERICA
(concluded) 49
VAN DUZEE, E. P., A NEW BEDUNIA 64
CHAMBERLIN, SOME CHILOPODS IMMIGRANT AT HAWAII 65
MARTIN, TWO NEW COLEOPTEROUS INSECTS FROM ARIZONA 70
FALL, A NEW APHODIUS AND A NEW BUPRESTID FROM CALIFORNIA ... 73
LINSLEY, NEW POGONOCHERUS AND ECYRUS 77
BEQUAERT, ASHMEAD’S GENUS POLISTELLA 91
STERNITZKY, A NEW SUBSPECIES OF PLEBEJUS ICAROIDES 93
HINTON, OBSERVATIONS ON TWO CALIFORNIA BEETLES 94
EDITORIAL COMMENT 96
San Francisco, California
1930
THE PAN-PACIFIC ENTOMOLOGIST
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the California Academy of Sciences.
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PUBLICATION COMMITTEE, PAN-PACIFIC
ENTOMOLOGIST
E. O. Essig, Chairman
G. F. Ferris R. W. Doane
E. C. Van Dyke Grant Wallace
REGIONAL MEMBERS
W. W. Henderson, Logan, Utah
J. C. Chamberlin, Riverside, California
E. P. Van Duzee, Editor
E. C. Van Dyke, Associate Editor
S. B. Freeborn, Treasurer
Published at the California Academy of Sciences, Golden Gate Park,
San Francisco, California.
Entered as second-class matter, February 10, 1925, at the postofRce at
San Francisco, California, under Act of August 24, 1912.
The Pan-Pacific Entomologist
Vol. VII, No. 2
October, 1930
THE DIPTEROUS GENUS SYMPYCNUS LOEW IN
NORTH AMERICA AND THE WEST INDIES
BY M. C. VAN DUZEE
Buffalo, New York
(Continued from page 47)
Sympycnus pugil Wheeler
Proc. Calif. Acad. Sci., Ser. 3, Zool., Vol. ii, 51, 1899. Cali-
fornia; Washington.
Length, 2.5 mm. Face gray-pollinose; front dull green; palpi
small, white-pollinose with a white bristle at tip; antenna black,
third joint small; lower orbital cilia white; thorax, scutellum, and
abdomen dull metallic green, the former with thin gray pollen;
abdomen a little yellow at base, its venter yellow; hypopygium large;
coxae, femora and tibiae yellow; hind femora sometimes dark above;
fore tarsi (Fig. 37) black from tip of first joint; middle tarsi (Fig. 38)
yellow with tip of first and whole of last four joints black; hind
tarsi (Fig. 39) black from tip of first joint; joints of fore tarsi as
24-6-5-5-6; of middle ones as 28-15-12-5-5; joints of posterior pair as
20-20-13-10-7; fore pulvilli much enlarged; wings narrowed at base;
fourth vein ending in apex of wing; last section of fifth vein as 22,
cross-vein as 17; calypters, their cilia and the halteres yellow.
Sympycnus pul villus Van Duzee, new species
Male. Length, 2 mm. Face silvery white, narrowed below;
front brown; occiput green, dulled with white pollen; antennas
(Fig. 18) black, third joint as long as wide, pointed at tip; lateral
and inferior orbital cilia white.
Dorsum of thorax green with only a little gray pollen; pleurae
more blackish, with white pollen, depressed space before the scutel-
lum quite conspicuous; abdomen green with coppery reflections on
the base of the segments beyond second, its hair black; hypopygium
(Fig. 20) small, inconspicuous, black, with very small, black append-
ages.
Coxae black with yellow tips; fore coxae with white hair; hair
on middle and hind ones and bristles on posterior pair yellow; fore
femora greenish black with yellow tip, their lower surface with
several long hairs and two bristles near tip; middle femora wholly
yellow; hind femora yellow with apical third metallic black, the
black extending along upper edge nearly to base; all tibiae yellow,
50
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
posterior ones brown on apical third; bristles of middle and hind
tibiae strong; fore tarsi (Fig. 19) yellow with rather long hair, last
joint black, a little flattened and widened, their pulvilli white, each
a little smaller than the fifth joint; middle tarsi brown almost to base,
hind ones wholly black; joints of fore tarsi as 16-6-6-5-5; fifth joint
as 3 wide and pulvilli as 4 long; joints of middle tarsi as 22-11-10-6-6;
of posterior ones as 19-16-11-7-7. Calypters, their cilia and the
halteres yellow, the former with a black tip.
Wings grayish, a little narrowed toward their root; last section of
fourth vein a little bent before basal third, its tip before the apex
of wing; last section of fifth vein as 34, cross vein as 11.
Described from five males ; two were taken by J. M. Aldrich
at Tennessee Pass, Colorado, holotype July 11, at 10,240 feet
elevation, the paratype July 23, 1917 ; three paratypes were
taken by the author at Mill Valley, Marin County, California,
March 25, 1926. Type in the United States National Museum.
The Tennessee Pass paratype may represent another species;
it differs from the other type in having the fore coxae almost
wholly yellow, fore femora less blackened; the fore tarsi are
missing, but the joints of middle and hind tarsi are in about
the same proportion as in holotype. The Mill Valley paratypes
are in such shape as to conceal the hypopygial appendages, the
abdomen appearing obtusely pointed at tip; in one there is a
slender, black lamella showing, which is fringed with hair.
The drawing of the antena is from a Mill Valley paratype and
the hypopygium from the holotype.
Sympycnus setosus Van Duzee, new species
Male. Length, 3 mm. Face silvery white, narrow below; palpi
brown; front blackish with gray pollen; antennas (Fig. 26) black or
brown, third joint as long as wide.
Thorax blackish with purple reflections, dulled with gray pollen;
acrostichal bristles represented by several very minute hairs; ab-
domen cylindrical, reddish coppery with blue or purple bands at base
of segments, its hairs minute, black; hypopygium colored like the
abdomen, its lamellae small, white with two black filaments or
bristles which are twice as long as the lamellae and extend the length
of the lamellae beyond them.
Coxae black, anterior pair with extreme tip yellow, front surface
with very small, black hairs and several black bristles at tip; tro-
chanters, femora and tibiae yellow; fore tibiae without bristles; middle
ones with two rather small ones; hind tibiae with several very small
bristles; fore and middle tarsi yellow, a little darker at tip; hind
tarsi (Fig. 27) black with first joint yellow at base, third joint thick
OCTOBER, 1930] VAN DUZEE GENUS SYMPYCNUS
51
with a stout bristle at tip and several smaller ones on one side, first
joint with small, erect spines below, these are not much more than
stout hairs and are continued on the next joint, but are still more
hair-like; fore pulvilli a little enlarged; fore and middle tarsi plain;
joints of fore tarsi as 23-10-6-5-5; of middle ones as 38-12-7-5-6;
joints of posterior pair as 25-21-10-9-7, bristle at tip of third joint
as 6. Calypters whitish at base, black at tip with black cilia; halteres
yellow.
Wings grayish. They are crumpled and broken in both type
specimens.
Described from two males, taken by the author June 11,
1915, at Victor, Colorado, at about 10,000 feet elevation. Type
in the author’s collection.
This is something like calcaratus, new species, in the form
and color of the hypopygial lamellae, but the lamellae are very
long in that species and very short in this ; the abdomen in
calcaratus is enlarged below at tip in this cylindrical and slightly
tapering to the tip; the third antennal joint is smaller in this
and hind tarsi much more spiney, especially the third joint.
Sympycnus tertianus Loew
Monog., North American Diptera, Vol. ii, 187, 1864. Alaska.
Male. Length, 2.5 mm. Face grayish; palpi and first two an-
tennal joints yellow; third antennal joint black, rounded; thorax and
abdomen grayish green, rather dull; thorax without lines; posterior
margin of pleurae pale yellow; venter of abdomen wholly yellow;
outer lamellae lancet-shaped, of rather dark color; coxae, femora and
tibiae pale yellow; tarsi blackish from tip of first joint; joints of fore
tarsi of decreasing length in the female, in male the third and fourth
joints are of nearly equal length; joints of middle tarsi of decreasing
length in both sexes; first joint of hind tarsi a little shorter than
second and in the male the third joint is somewhat shorter than
fourth, fourth with longer hairs at tip; wings much narrowed at
base; fourth vein ends a little before the apex of the wing; cross
vein at the middle between root and apex of wing.
Sympycnus thoracicus Van Duzee, new species
Male. Length, 2.5 mm. Eyes contiguous, reducing the face to
a very narrow line of white on upper half; front black with white
pollen, narrow; antennae black, third joint very small, triangular.
Dorsum of thorax and scutellum very bright green, the former
with golden reflections; pleurae brown, largely yellowish; abdomen
cylindrical, black with first and second segments reddish yellow, its
hairs yellow, those on venter as long as thickness of abdomen;
hypopygium (Fig. 6) black, small, its outer lamellae long and narrow,
black; also a pair of small, hook-like inner appendages.
52
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
Coxae, femora, tibiae and tarsi yellow, last joint of middle tarsi
black; all tarsi plain, except that the fore tarsi are very slender and
have long hair on all joints, many of them three times as long as
diameter of the joint, their pulvilli very large; joints of fore tarsi as
25-15-8-6-6; of middle ones as 28-15-12-9-7; joints of posterior ones
as 18-23-14-9-6. Calypters yellow with black tips and cilia; halteres
yellow.
Wings grayish; third and fourth veins parallel and a little arched,
tip of fourth back of apex of wing; last section of fifth vein as 27,
cross vein as 17; anal angle of wing prominent.
Described from one male taken at St. Vincent, West Indies,
at 1500 feet elevation. Type No, 33, Arkansas University
collection.
S 3 mipycnus tripilus Van Duzee, new species
Male. Length, 2-2.5 mm. Face silvery white, narrow below, its
sides parallel on lower two-thirds; palpi small yellow; proboscis
yellowish brown; front black in middle, shining green on sides;
antennae (Fig. 21) black, third joint large, nearly twice as long as
wide, rounded at tip; lower orbits nearly bare, a few black cilia
above.
Thorax metallic brown with green reflections and brown pollen;
acrostichal bristles rather long, in a single row; pleurae white-polli-
nose, its posterior edge yellow; abdomen brown above, venter, most
of second segment and sides of others yellow; hairs on dorsum black,
bristles on sides of first segment yellow; hypopygium greenish black,
its outer appendages narrow, short, yellow.
All coxae yellow; fore coxae with a few small yellow hairs, these
become longer and more bristle-like toward the tip; femora and
tibiae yellow, posterior femora and tibiae more or less brown at tip;
fore tibiae without bristles, middle ones with three, hind tibiae with
several rather small bristles; fore and middle tarsi yellow at base,
brown apically, hind ones wholly brown; middle tarsi (Fig. 22) with
three long, curved hairs on anterior surface of second joint, and a
short, straight one near tip, first joint with several very small spines,
second and third joints with a small spine at tip, third and fourth
joints each with a long hair, fifth with long hairs above, but these
are not longer than in some other species; joints of fore tarsi as
19-13-8-6-5; of middle ones as 32-10-7-6-5; joints of posterior pair
as 15-20-12-8-7. Calypters, their cilia and the halteres yellow.
Wings grayish; last section of fourth vein straight, nearly parallel
with third, ending in the apex of the wing; last section of fifth vein
as 25, cross vein as 9; wing narrowed at base.
Female. Face white, wide; proboscis yellow; base of hind tarsi
yellow; abdomen more brown or yellowish brown; otherwise about
as in the male. Sometimes the antennae are brown or yellowish
brown.
OCTOBER, 1930] VAN DUZEE GENUS SYMPYCNUS
53
Described from eleven males and four females, taken by
the author, April 15, 1915, in San Diego County, California, on
the edge of the Colorado Desert. Type in author’s collection.
Sympycnus varipes Aldrich
Biologia, Diptera, Vol. i, 344, 1901. Mexico.
Male. Length, 2.6 mm. Face and front brown pollinose; an-
tennas black, short; dorsum of thorax opaque gray with a brown,
median line and the bristles inserted in brown dots; scutellum bright
blue; abdomen bronze green with anterior part of each segment
purplish, abdomen without yellow; fore coxas, all femora and tibiae
yellow, tip of hind femora and tibiae and whole of hind tarsi blackish;
fore tarsi yellow with last joint black and enlarged; middle tarsi
yellow with tips of first four joints and whole of last joint blackish;
middle coxas with white hairs in front, hind ones with one blackish
bristle; fore pulvilli enlarged; first and second joints of hind tarsi of
nearly equal length; wings grayish; moderately narrowed at base;
fourth vein ending in apex of wing.
Key to Males of the Species of the Subgenus
Calyxoch^etus
1. Fore tibias considerably incrassated; second joint of fore tarsi
with a long slender bristle or hair at tip 2
— Fore tibias not or but little incrassated 3
2. Arista quite thick, gradually, but considerably enlarged to-
ward tip; Figs. 43-46 (New England States; New York;
Pennsylvania; New Jersey; Ontario) fortunatus Loew
— Arista slender with a conspicuous, lamella at tip; Figs. 47-49
(California) ‘vegetus Wheeler
3. Fourth joint of fore tarsi distinctly longer than third 4
— Fourth joint of fore tarsi shorter than third 5
— Fore tarsi with third and fourth joints of nearly equal length 7
4. Arista plain; fourth joint of fore tarsi with hooked hairs or
bristles above; Figs. 50-52 (Maine; Massachusetts; New
York; Pennsylvania; Indiana) frontalis Loew
— Arista gradually but considerably enlarged on apical fifth;
Figs. 53-55 (Utah) distortus new species
— Arista with a large, nearly round lamella at tip; Figs. 56-58
(Colorado; Alaska) cilifemoratus Van Duzee
5. Fourth joint of fore tarsi about half as long as third (Cali-
fornia cereas Wheeler
— Fourth joint of fore tarsi much more than half as long as
third 6
6. Second joint of middle tarsi with several long hairs on basal
half; Figs. 59-61 (Maine; Ontario) luteipes Van Duzee
54
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
— Tarsi plain, without unusual hairs (Kansas)
abbreviatus Van Duzee
7. Third joint of middle tarsi a little longer than fifth, fourth and
fifth of equal length; Figs. 66-69 (Illinois; Wisconsin;
Missouri; New York; Ontario) nodatus Loew
— Third and fourth joints of middle tarsi each much longer than
fifth; Figs. 62-65 (California) clavicornis new species
— Third and fourth joints of middle tarsi each much shorter
than fifth 8
8. Apical enlargement of arista slight, occupying only apical
third; Figs. 70-72 (Idaho; California) sobrinus Wheeler
— Enlargement of arista quite conspicuous, occupying most of
apical half; Figs. 73-75 (California) hastatus new species
Sympycnus (Calyxoch^tus) abbreviatus Van Duzee
Canadian Entomologist, Vol. xlix, 341, 1917. Kansas.
Male. Length, 3 mm. Face narrow, silvery white; palpi yellowish;
proboscis brown; front blue with thick brown pollen; antennae brown,
third joint large, nearly as wide as long, somewhat square with
rounded corners; dorsum of thorax brown, nearly opaque; abdomen
yellowish at base, black from hind margin of second segment,
bronzed at tip; coxae yellow, anterior ones with minute yellow hairs
and yellow bristles at tip; middle and hind coxae with a black bristle
on outer surface; all femora and tibiae yellow, hind femora and tibiae
darkened at tip, the latter with a rather long, slender bristle near
basal third; fore tarsi yellow, darkened at tip, first joint not longer
than thick, second more than half as long as tibia, third a little
longer than fourth, which is scarcely longer than fifth; hind tarsi
black from tip of first joint, which is shorter than second; wings
grayish, rather narrow at base; veins yellowish brown, costa darker;
fourth vein ending in apex of wing; last section of fifth vein three
times as long as cross vein.
Sympycnus (Calyxoch^tus) cilifemoratus
Van Duzee
Proc. U. S. National Museum, Vol. Ixiii, Art. 21, 12, 1924.
Colorado; Alaska.
Male. Length, 2.5-3 mm. Face narrow below, covered with
white pollen; front blackish or blue with brown pollen; antennas
(Fig. 56) black; thorax bronze brown with brown pollen and blue
reflections; abdomen dark metallic green, bristles on first segment
yellow; hypopygial appendages small, black; all coxae, femora and
fore and middle tibiae yellow, apical half of hind femora more or less
blackish; hind tibiae blackish, sometimes yellowish at base; fore coxae
and fore and middle tibiae with yellow hairs; fore tarsi (Fig. 57) and
middle tarsi (Fig. 58) black from tip of first joint; hind tarsi wholly
black; joints of fore tarsi as 3-29-8-11-7; of middle ones as 34-14-
OCTOBER, 1930] VAN DUZEE GENUS SYMPYCNUS
55
11-10-5; those of posterior pair as 18-23-14-11-8; wings narrowed at
base; third and fourth veins a very little convergent at tips, fourth
ending just before the apex of wing; last section of fifth vein as 31,
cross vein as 10; calypters, their cilia and the halteres yellow.
Sympycnus (Calyxochaetus) clavicornis Van Duzee
new species
Male. Length, 2.5 mm. Face narrow, eyes touching below, black
with white pollen; front blackish; palpi black; antennas (Fig. 62)
brown, first joint long, third as long as first, slightly more than half
as wide as long, tip broadly rounded, arista inserted at tip of third
joint, gradually much widened on apical fourth.
Thorax and abdomen dark brown, shining, dorsum of thorax with
brown pollen, which leaves darker lines; scutellum a little bluish;
posterior edge of pleurae partly yellowish; sides of second abdominal
segment wholly yellow, remaining segments slightly tinged with
green; hypopygium (Fig. 65) brown, with very small, triangular,
yellow lamellas, a pair of large inner lamellae and a long, smooth,
curved, yellow central organ; hairs of abdomen partly yellowish.
Coxae, femora and tibiae yellow; fore coxae with a few white hairs;
middle and hind coxae a little darkened on outer surface; nearly
apical fourth of hind femora brown; hind tibiae more or less
brownish, especially at tip and with several long pale hairs below
near tip; fore tarsi (Fig. 63) brown from tip of second joint; middle
tarsi (Fig. 64) brown at tip, second joint enlarged and with several
long, delicate hairs below; hind tarsi wholly blackish; joints of fore
tarsi as 4-21-9-10-7; of middle ones as 30-11-10-8-5; joints of pos-
terior pair as 20-23-14-11-7. Calypters, their cilia and halteres yellow.
Wings grayish; third and fourth veins nearly straight and paral-
lel, fourth ending distinctly before the apex of wing; last section of
fifth vein 45, cross vein ten-fiftieths of a millimeter long.
Female. Face moderately wide, covered with white pollen; front
blue-black; palpi brown; third antennal, joint nearly round in outline;
lower orbital cilia white; abdomen a very little yellowish on the sides
at base; joints of fore tarsi as 20-15-9-6-6; of middle ones as 33-12-
10-8-7; those of posterior pair as 21-21-14-9-9; wings with third and
fourth veins a little farther apart than in male, fourth ending in apex
■of wing; last section of fifth vein 48, cross vein 14; legs almost
wholly yellow, tarsi a little brownish.
Described from two males and one female, taken by J. M.
Aldrich, July 23, 1911, at Mono Lake, California. Type in the
United States National Museum.
Sympycnus (Calyxochaetus) distortus Van Duzee
new species
Male. Length, 2mm. Antennas (Fig. 53) brown, third joint
rounded, but little longer than wide, arista as long as fore tarsi,
apical fifth gradually but considerably enlarged.
56
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
Thorax and abdomen brown, the former with a little brown
pollen; second segment of abdomen largely yellow; hypopygium
about as in davicornis, rounded above with small, triangular, yel-
lowish outer lamellae, which are not longer than wide, inner append-
ages rounded.
Coxae, femora and tibiae yellow; tips of posterior femora and tibiae
brownish; fore coxae with a few short white hairs; hind femora with
black hairs below, which are short at base, long near tip; fore tarsi
(Fig. 54) yellow with last joint brown, third joint shorter than
fourth, which is curved and has a few long hairs; middle tarsi
(Fig. 55) yellow with last three joints brown, first joint with one
long hair near tip, second with about four long hairs on basal half;
hind tarsi wholly brown; joints of fore tarsi as 3-23-9-13-7; of middle
ones as 30-13-12-9-5; joints of posterior pair as 20-24-14-10-8. Calyp-
ters, their cilia and the halteres yellow.
Wings grayish; third and fourth veins nearly parallel, but bent
back and slightly approaching each other at tip, fourth ending just
in front of apex of wing; cross vein about as 6, last section of fifth
vein as 35.
Described from one male, taken by J. M. Aldrich, July 6,
at Soldier Summit, Utah, at 7454 feet elevation. Type in the
United States National Museum.
Sympycnus (Calyxoch^tus) fortunatus Wheeler
Proc. Calif. Acad. Sci., Ser. 3, Zool., Vol. ii, 52, 1899. North-
eastern States; Ontario.
Male. Length, 3.3 mm. Face very narrow, light gray-pollinose;
front violet; antennae (Fig. 43) dark brown; thorax dull black,
covered with brown pollen, except a median vitta and spots above
the humeri; scutellum violet; abdomen brownish black, base of first,
second and third segments and the venter pale yellow; all coxae and
femora pale yellow; fore coxae with silvery white hairs; posterior
femora brown on apical third; all tibiae pale yellow, tips of hind ones
blackish; fore tibiae and tarsi (Fig. 44), the tarsi black from middle
of second joint; middle tarsi (Figs. 45 and 46) blackish from tip of
first joint; hind tarsi wholly black; length of fore tibiae as 43, joints
of fore tarsi as 5-25-18-7-7; of middle tarsi as 43-11-5-8-7; joints of
posterior pair as 23-29-15-10-7; wings much narrowed at base with
a lobe basally from tip of fifth vein; third and fourth veins nearly
parallel, but slightly arched; last section of fifth vein as 40, cross
vein as 10.
Sympycnus (Calyxoch^tus) frontalis Loetv
Neue Beit., Vol. viii, 67, 1861 ; Monog. N. Am. Diptera, Vol. ii,
188, 1864. Northeastern States.
OCTOBER, 1930] VAN DUZEE GENUS SYMPYCNUS
57
Male. Length, 3 mm. Face very narrow below, covered with
white pollen so as to nearly conceal the blue ground color; front
bright violet; antennas (Fig. 50) black or brown; dorsum of thorax
green or blue with thick grayish brown pollen; abdomen black or
greenish black with more or less yellow, especially on the sides;
posterior margin of pleurae, all coxae, femora, tibiae and tarsi yellow,
tarsi more or less infuscated; fore tarsi (Fig. 51) with its joints as
4-20-9-14-6; middle tarsi (Fig. 52) with its joints as 40-14-4-4-7;
those of posterior tarsi as 19-24-14-11-6; wings with third and fourth
veins nearly straight and parallel, fourth ending in apex of wing;
last section of fifth vein as 32, cross vein as 13; calypters, their cilia
and the halteres yellow.
Sympycnus (Calyxochaetus) hastatus Van Duzee
new species
Male. Length, 3 mm. Face narrow, white; front purple, shining,
except a narrow margin of white pollen along the orbits and above
the antennae; antennae (Fig. 73) black, third joint more brown,
rounded at tip, arista rather short, its first two joints as 22-28, third
joint widened before its middle, but pointed at tip; lower orbital cilia
white.
Thorax green, dull, scutellum blue. Abdomen dull green, second
and third segments yellow, except a median dorsal line, which
widens at the sutures, its hair black; hypopygium black, pointed
below, with a pair of short, pale lamellae, fringed with black hair.
Fore coxae yellow with minute yellow hair; middle and hind
coxae black with yellow tips; femora and tibia yellow, posterior
femora on apical half, especially above, and extreme tips of hind tibia
brown; fore tarsi (Fig. 74) yellow, darker toward tip; middle tarsi
(Fig. 75) yellowish brown; posterior tarsi black from tip of first
joint; joints of fore tarsi as 4-29-14-14-7; of middle ones as 50-12-
4-5-8; joints of posterior pair as 22-27-15-10-5. Calypters yellow with
black tip and pale cilia; halteres yellow.
Wings grayish; third and fourth veins nearly parallel, the latter
ending in apex of wing; cross vein 12, last section of fifth vein 43-
fiftieths of a millimeter long; sixth vein short; wing not much
narrowed at base.
Described from one male taken by the author, April 15,
1915, in San Diego County, California, on the edge of the
Colorado Desert. Type in the author’s collection.
Sympycnus (Calyxochzetus) luteipes Van Duzee
Psyche, Vol. xxx, 63, tar salts, Curran, Canadian Entomologist,
Vol. Ivi, 108, 1924. Maine; New York; Ontario.
Male. Length, 2 mm. Face narrow, white; front dark blue; an-
tennae (Fig. 59) black or brown; dorsum of thorax brown, a little
58
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
dulled with brown pollen; scutellum blue; posterior edge of pleurae
yellow; abdomen with yellow on the sides; all coxae, femora and
fore and middle tibiae yellowish; tips of hind femora and the hind
tibiae more brown or black; fore tarsi (Fig. 60) yellow, blackened
from tip of third joint; middle tarsi (Fig. 61) black from tip of first
joint; hind tarsi wholly black, shorter than their tibia; joints of fore
tarsi as 4-16-10-6-5; of middle ones as 31-10-9-8-5; those of posterior
pair as 14-16-9-8-6; wings brownish gray, narrowed at base; last
section of fifth vein nearly four times as long as cross vein; calypters
and halteres yellow, former with black cilia.
Sympycnus (Calyxoch^tus) nodatus Loetv
Loew’s Cent., Vol. ii. No. 68, Berl. Ent. Zeit., Vol. vi, 215,
1862; Monog. North American Diptera, Vol. ii, 191, 1864.
Arkansas; New York; Illinois; Missouri; Wisconsin;
Ontario.
Male. Length, 2.5-3 mm. Face very narrow, white; front brown
with a little gray pollen; palpi small, yellow; antennae (Fig. 66)
brown, first joint yellow below or wholly yellow; dorsum of thorax
brownish gray; posterior margin of pleurae yellow; scutellum usually
blue; abdomen metallic black, yellow on sides and venter; hypo-
pygial appendages black, outer lamellae yellowish; coxae, femora and
tibiae yellow; fore tarsi (Fig. 67) yellow, apical joints black; middle
tarsi (Figs. 68 and 69) black from tip of first joint; hind tarsi with
last four joints black; joints of fore tarsi as 4-18-10-8-8; of middle
ones as 45-15-7-5-6; of posterior pair as 20-29-15-11-8; wings nar-
rowed at base, hind margin a little sinuous on basal half; last section
of fourth vein straight, ending in apex of wing, nearly parallel with
third, but third bent back a little so as to approach fourth at tip;
last section of fifth vein as 42, cross vein as 13; calypters yellow with
black tip, its cilia yellow, but appearing blackish in certain lights;
halteres yellow.
Sympycnus (Calyxoch^tus) oreas Wheeler
Proc. Calif. Acad. Sci., Ser. 3, Zook, Vol. ii, 55, 1899. Cali-
fornia.
Male. Length, 3 mm. Face very narrow, gray pollinose; pro-
boscis and palpi brown; front green; antennas blackish, third joint
large, oval, evenly rounded, arista basal; long, tapering to a point;
thorax and scutellum green, thickly covered with yellow pollen,
especially in front, leaving a median shining vitta; abdomen dull
blackish, sides and venter partly yellow; fore coxas yellow with white
hairs, middle and hind coxas blackish; femora and tibias yellow, apical
half of hind femora and tips of their tibias blackened; fore tibias dis-
OCTOBER, 1930] VAN DUZEE GENUS SYMPYCNUS
59
tinctly swollen; all tarsi black from tip of first joint; first joint of
fore tarsi very short, second and third much longer and subequal,
fourth and fifth subequal and about half as long as third, third and
fourth joints fringed above with long hair; first joint of hind tarsi
distinctly broader and shorter than second; wings much narrowed
at base, somewhat truncate at apex; last section of fourth vein paral-
lel with third on its apical half, ending in or a little before apex of
wing; last section of fifth vein more than three times as long as
cross vein; calypters yellow with brown border and yellow cilia;
halteres yellow.
Sympycnus (Calyxoch^tus) sobrinus Wheeler
Proc. Calif. Acad. Sci., Ser. 3, Zool., Vol. ii, 54, 1899. Idaho;
California.
Male. Length, 2.5-3 mm. Face rather broad, white; front green;
antennas (Fig. 70) black; thorax greenish, nearly opaque with brown
pollen; scutellum blue; abdomen blackish, sometimes the sides yellow
at base, venter yellowish; hypopygial lamellae very smal, yellowish;
fore coxae yellow with a few yellow hairs and bristles; middle and
hind coxae mostly black; femora and tibiae dark yellow, hind femora
blackish apically; tarsi infuscated from tip of first joint; fore tarsi
(Fig. 71) with their joints as 4-21-13-12-8; middle tarsi (Fig. 72)
with the joints as 45-15-4-6-9; joints of posterior tarsi as 21-26-15-
10-9; wings with anal angle prominent; third and fourth veins nearly
parallel, but bent backward toward their tips; last section of fifth
vein as 44, cross vein as 12; fourth vein ending in the apex of the
wing; calypters, their cilia and the halteres yellow, sometimes the
cilia of the calypters is more brown.
Sympycnus (Calyxochastus) vegetus Wheeler
Proc. Calif. Acad. Sci., Ser. 3, Zool., Vol. ii, 53, 1899. Idaho;
California; Oregon.
Male. Length, 2. 5-3. 3 mm. Face very narrow, white-pollinose;
front violet; antennas (Fig. 47) black; palpi blackish; thorax dull
green or blackish with coppery reflections; scutellum blue; abdomen
bronze-black with yellow on sides and venter; hypopygial lamellae
small, yellowish brown; coxae, femora and fore and middle tibiae
yellow; tips of hind femora and whole of hind tibiae brown; fore
tibiae and tarsi (Fig. 48); middle tarsi (Fig. 49), black from tip of
first joint; hind tarsi wholly black; joints of fore tarsi as 4-22-17-8-7;
of middle tarsi as 39-15-4-5-6; of posterior pair as 25-27-16-11-8;
wings narrowed at base; third vein greatly bent backward at tip,
parallel with fourth, which ends in apex of wing; last section of fifth
vein as 55, cross vein as 12; calypters, their cilia and the halteres
yellow.
60
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 7.
Fig. 8.
Fig. 9.
Fig. 10.
Fig. 11.
Fig. 12.
Fig. 13.
Fig. 14.
Fig. 15.
Fig. 16.
Fig. 17.
Fig. 18.
Fig. 19.
Fig. 20.
Fig. 21.
Explanation of Plate 1
North American species of Sympycnus
cuprinus Wheeler, an-
tenna of male.
lineatus Loew, antenna
of male.
breviventris n. sp., an-
tenna of male.
breviventris, fore tarsus
of male.
breviventris, hypopygium
of male.
thoracicus n. sp., hypo-
pygium of male.
clavatus V. D., antenna
of male.
clavatus, fore tarsus of
male.
Isevigatus n. sp., antenna
of male.
lasvigatus, fore tarsus of
male.
inasqualis n. sp., antenna
of male.
insequalis, hypopygium of
male.
caudatus V. D., antenna
of male.
caudatus, fore tarsus of
male.
latitarsis n. sp., antenna
of male.
latitarsis, fore tarsus of
male (top view).
latitarsis, fore tarsus of
male (side view).
pulvillus n. sp., antenna
of male.
pulvillus, fore tarsus of
male.
pulvillus, hypopygium of
male.
tripilus n. sp., antenna of
male.
Fig. 22. tripilus, mid tarsus of
male.
Fig. 23. calcaratus n. sp., antenna
of male.
Fig. 24. calcaratus, hind tarsus of
male.
Fig. 25. calcaratus, hypopygium
of male.
Fig. 26. setosus n. sp., antenna of
male.
Fig. 27. setosus, hind tarsus of
male.
Fig. 28. globulicauda n. sp., an-
tenna of male.
Fig. 29. globulicauda, hind tarsus
of male.
Fig. 30. globulicauda, hypopy-
gium of male.
Fig. 31. fasciventris V. D., an-
tenna of male.
Fig. 32. fasciventris, hind tarsi of
male.
Fig. 33. mercedus Wheeler, an-
tenna of male.
Fig. 34. mercedus Wheeler, fore
tarsus of male.
Fig. 35. mercedus, hind tarsus of
male (side view).
Fig. 36. mercedus, hind tarsus of
male (top view).
Fig. 37. pugil Wheeler, fore tar-
sus of male.
Fig. 38. pugil, mid tarsus of male.
Fig. 39. pugil, hind tarsus of male.
Fig. 40. aldrichi n, sp., antenna
of male.
Fig. 41. aldrichi, hind tarsus of
male.
Fig. 42. montanus n. sp., mid tar-
sus of male.
OCTOBER, 1930] VAN DUZEE — GENUS SYMPYCNUS
61
M. C. Van Duzee, Plate I
62
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
Explanation of Plate 2
North American species of Sympycnus, subgenus Calyxochaetus
Fig. 43. fortunatus Wheeler, an-
tenna of male.
Fig. 44, fortunatus, fore tibise and
tarsus of male.
Fig. 45. fortunatus, mid tarsus of
male (top view).
Fig. 46. fortunatus, last four
joints of mid tarsus
(bottom view).
Fig. 47. vegetus Wheeler, an-
tenna of male.
Fig. 48. vegetus, fore tarsus of
male.
Fig. 49. vegetus, mid tarsus of
male.
Fig. 50. frontalis Loew, antenna
of male.
Fig. 51. frontalis, fore tarsus of
male.
Fig. 52. frontalis, mid tarsus of
male.
Fig. 53. distortus n. sp., antenna
of male.
Fig. 54. distortus, fore tarsus of
male.
Fig. 55. distortus, mid tarsus of
male.
Fig. 56. cilifemoratus V. D., an-
tenna of male.
Fig. 57. cilifemoratus, fore tarsus
of male.
Fig. 58, cilifemoratus, mid tarsus
of male.
Fig. 59. luteipes V. D., antenna
of male.
Fig. 60. luteipes, fore tarsus of
male.
Fig. 61. luteipes, mid tarsus of
male.
Fig. 62. clavicornis n. sp., antenna
of male.
Fig. 63. clavicornis, fore tarsus
of male.
Fig. 64. clavicornis, mid tarsus of
male.
Fig. 65. clavicornis, hypopygium
male.
Fig. 66. nodatus Loew, antenna
of male.
Fig. 67. nodatus, fore tarsus of
male.
Fig. 68. nodatus, mid tarsus of
male (top view).
Fig. 69. nodatus, mid tarsus of
male (side view).
Fig. 70. sobrinus Wheeler, an-
tenna of male.
Fig. 71. sobrinus, fore tarsus of
male.
Fig. 72. sobrinus, mid tarsus of
male.
Fig. 73. hastatus n. sp., antenna
of male.
Fig. 74. hastatus, fore tarsus of
male.
Fig. 75. hastatus n. sp., mid tar-
sus of male.
Fig. 76. Sympycnus canadensis
V. D., hypopygium of
male.
OCTOBER, 1930] VAN DUZEE GENUS SYMPYCNUS
63
M. C. Van Duzee, Plate II
64
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
A NEW BEDUNIA (HEMIPTERA)
BY E. P. VAN DUZEE
From Professor Roland Thaxter I have recently received
for study three interesting exotic Hemiptera, one of which
proves to be undescribed. The other two are Microvelia albo-
lineolata Bueno, taken at the type locality. Fort De Kock,
Sumatra, and Melanolestes argentinus Berg, from Santa Cata-
lina, Llavallol, Argentina. The latter does not entirely answer to
Berg’s description, so the determination is in some doubt. The
Lygaeid is :
Bedunia adusta Van Duzee, n. sp.
Near segmentata, but apparently distinct. Black, posterior
lobe of pronotum, scutellum and elytra marked with pale casta-
neous ; antennae, rostrum and legs honey- yellow. Length, 6 mm.
Head a little longer than pronotum; tylus produced for nearly
half its length before the cheeks. Vertex finely rugosely punctate,
the lateral areas exterior to the arcuate impressed lines minutely
punctate and sparsely dotted with pale hairs. Antennas, Seg. II a
little the longest; I, III and IV subequal. Rostrum reaching on to
first ventral segment. Pronotum a fifth wider at base than long;
anterior lobe a little longer than posterior, finely punctate, collum
fiat, with one row of punctures, area of callosities large, indefinite,
nearly impunctate; posterior lobe and scutellum coarsely punctate;
clavus with four rows of coarse punctures, corium with two rows
of punctures next the suture, one along the radial vein and a few
scattering punctures toward apex. Anterior femora much incrassate,
with a double row of small teeth on apical half; segment I of hind
tarsi three times as long as II and III together.
Color black, tinged with castaneous on apex of tylus, acetabulas,
'coxas and venter; antennse and legs deep hone3'--yellow or ochrace-
'Ous; antennal segments III and IV slightly darker at base; rostrum
and base of femora paler; posterior lobe of pronotum, collum and
scutellum with a castaneous tinge, the former with a vitta either
side the middle of disk and two basal spots between them, and the
raised arc of the scutellum, paler castaneous; tip of scutellum, basal
half of costa, a spot before its apex and two obscure discal vittas,
the inner short, ochraceous; membrane short, paler at base, attaining
sixth tergite; tip of rostrum black.
Holotype, male, and allotype, female, taken at Wai-Ai,
Solomon Islands, received from Dr. Roland Thaxter under the
number 2959. On the female the pale markings are more
obscure (Holotype, No. 2975, Mus. Calif. Acad. Sci.).
OCTOBER, 1930]
CHAMBERLIN — CHILOPODS
65
ON SOME CHILOPODS IMMIGRANT AT HAWAII
BY RALPH V. CHAMBERLIN
University of Utah, Salt Lake City
The present paper records fifteen species of chilopods taken
chiefly from plants or the soil about them in baggage and cargo
of ships arriving at Hawaii. These were submitted to me for
identification by the Plant Quarantine and Control Adminis-
tration of the United States Department of Agriculture. Most
were taken during the year 1928.
The types of the five species described as new are in the
writer’s collection.
S COLO PENDROM ORPH A
1. Paracryptops weberi Pocock
A single specimen of this unusual species was taken in soil
from Singapore, Straits Settlement, at Honolulu on March 10,
1930.
2. Cr 3 rptops navis Chamberlin, new species
Head overlapped by first dorsal plate; without sulci. Antennae
in holotype consisting of fifteen (right) and seventeen (left) articles.
First dorsal plate without transverse or longitudinal sulci. Pre-
hensors with claws of normal length; anterior margin of prosternum
smooth, slightly convex on each side of the median indentation.
Paired dorsal sulci present from the third dorsal plate, inclusive,
caudad.
Last ventral plate with caudal margin a little convex, the pos-
terior corners well rounded and the sides diverging cephalad.
Last coxae caudally truncate; pores small, moderate in number,
the porose area lying along the ventral plate the end of which it
reaches, but failing to reach the caudal margin of coxae and dorsal
plate by a wide area; caudal margin of coxae setose and one or two
setae just cephalad of margin.
Anterior spiracles rather large, elliptic.
Femur and tibia of anal legs subseriately and subdensely setose
ventrally and mesally, the setae passing into finer, ordinary hairs on
outer surface. Metatarsus with a series of five teeth; first tarsal
joint with a series of four teeth which decrease in size distad.
Length, ad 10 mm.
Locality. One specimen intercepted at Honolulu, Hawaii,
March 10, 1930, in soil from Singapore, Straits Settlement.
66
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
3. Newportia sp.
A specimen of a species of this genus was taken at Hawaii
in 1928 in soil about plants in the cargo of a vessel from
Panama. The anal legs are missing, making specific determi-
nation impossible.
4. Otostigmus sinicolens Chamberlin, new species
Body above dark brownish green, with head and first tergite
abruptly light ochre yellow. Antennae yellowish proximally, darker
distally. Legs greenish, the anal pair with a light ring near distal
end of fourth and fifth joint and with the distal half of the sixth
and all of the seventh joint also pale.
Head smooth, without conspicuous sulci or punctae. Antennae
composed of twenty articles of which only the first two are glabrous,
the third being entirely clothed similarly to the more distal ones.
Paired sulci distinct and complete from the sixth tergite caudad.
Tergites laterad of sulci finely rugose and punctate, the roughening
strong on caudal plates. Between the paired sulci of median and
posterior plates three secondary sulci separating off low longitudinal
ridges, these being less clearly defined in the anterior region.
Laterad of the paired sulci in posterior region several weakly devel-
oped longitudinal ridges also indicated.
Last dorsal plate with a deep, elongate pit on the median line
just in front of the caudal margin.
Prosternum with 5 + 5 teeth of which the mesal one on each
side is partly divided. The basal process of femur with three tuber-
cles or teeth on its mesal apical slope.
Last ventral plate strongly narrowed caudad, the caudal margin
mesally concave. Other ventral plates excepting the most anterior
showing paired sulci over anterior border, but these in no case com-
plete though on some of the posterior sternites continued in weak
longitudinal furrows that may simulate sulci.
Two tarsal spines on about the first eight pairs of legs, on all
others single excepting on the twentieth and twenty-first from which
absent.
Process of anal coxae (pseudopleurae) pronounced, densely porose
as usual excepting in a longitudinal ventral stripe which extends
from caudal end of process nearly to corner of last sternite; process
ending in three spines, with one spine on dorsal side a little proxi-
mad of apical group and two spines on the dorso-lateral margin near
base of process.
Femur of anal legs with a ventral row of four teeth; on ectal
side with three oblique rows, 2, 4, 1, and a dorsal longitudinal series
of four teeth of which one is at the distal end of joint and is a little
longer than the others.
Length, 42 mm.
OCTOBER, 1930] CHAMBERLIN — CHILOPODS
67
Locality. The holotype was taken at Hawaii on specimens
of Eleocharis tuherosa, imported from China in 1928.
5. Otostigmus mians Chamberlin, new species
Dorsum and legs greenish yellow, excepting the last two tergites
which are abruptly lighter, more yellow; the head and antennae a
darker green.
Head smooth. Antennae consisting of nineteen articles of which
the first two are wholly glabrous and the third likewise glabrous
excepting over distal third above.
Prosternum with teeth 2 + 2, each tooth, however, being partly
divided, the ectal one having a denticle on its outer side at base and
the inner one being partly divided at end with inner denticle the
smaller.
Paired sulci beginning on fifth tergite.
Last tergite without median sulcus or pit. All tergites smooth
and shining.
No complete paired sulci on any of the sternites, sulci being indi-
cated only across anterior borders of some plates. Last ventral plate
narrowed caudad, its caudal margin mesally incurved.
Two tarsal spines on the first eight pairs of legs; others, except-
ing the last two pairs, with one tarsal spine.
Pseudopleural processes elongate, bearing two teeth at end and
a third subapical one on outer side and a single dorso-lateral one
toward base. Femur of anal legs with ventral longitudinal row of
four teeth and two mesal rows of three and two teeth; none at distal
end of joint.
Length, 35 mm.
Locality. The holotype taken at Hawaii in 1928 on Dio-
scorea sp. from China. On this specimen one anal leg is in
the initial stage of regeneration. A paratype lacking both anal
legs was taken, also on Dioscorea, in a different lot of speci-
mens during the same year.
6. ScoLOPENDRA SUBSPINIPES Leach
One specimen of this oriental species was taken on Pachy-
rhizus sp. from China in 1928.
Geophilomorpha
7. Mecistocephalus maxillaris Gervais
Several specimens of this species, carried widely by ships
in tropical regions, were taken at Hawaii in 1928 in soil about
plants from Panama.
68
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
Lithobiomorpha
8. Lamyctinus cceculus Brolemann
Several specimens of this now tropicopolitan form were
taken in soil about plants from Tahiti on two dates in 1928.
9. Lamyctes species
A specimen of uncertain species was taken in 1928 in soil
about plants in baggage from Japan.
10. Lamyctes fulvicornis Meinert
Taken at Hawaii April 22, 1929, in soil about plants from
Los Angeles, California.
11. Oabius pylorus Chamberlin
A specimen was taken April 2, 1929, in soil about chives
from San Francisco, California.
12. Lithobius borealis Meinert
Taken November 6, 1929, in packing of orchids from
England.
13. Lithobius species
Specimens too young for identification were taken in 1927
and 1928 in soil about plants from China.
14. Nipponobius sinensis Chamberlin, new species
Head and dorsum yellow, the head lighter in front of the frontal
suture; posterior legs also yellow proximally but distad of third
article lighter, whitish; the other legs light yellow. Antennae con-
colorous with head, the color brighter distally.
Antennae short, distally attenuated, the twenty articles of moder-
ate length.
Ocelli small, few, in a narrowly linear patch, in two series to
be interpreted as 1 4- 2, 2, with the lower series in advance of the
upper, or as 1 -[- 3, 1.
Prosternal teeth small and pale, well separated, 2 + 2.
Coxal pores small, circular, 1, 2, 2, 2.
All posterior coxae wholly unarmed. Ventral spines of first legs
0, 0, 0, 0, 1. Penult legs with ventral spines 0, 1, 2, 1, 0; dorsal
spines 0, 0, 2, 1, 0, the claw double. Anal legs with ventral spines
also 0, 1, 2, 1, 0; dorsal spines 0, 0, 2, 0, 0, the claw single.
The anal legs of male with fifth article a little swollen, with a
shallow longitudinal dorsal sulcus and a dorsal crest at distal end
resembling that in males of Nadabius.
Length, 7 mm.
OCTOBER, 1930]
CHAMBERLIN — CHILOPODS
69
Locality. The types, two adult males, were taken at
Hawaii in 1928 on Lillium species imported from China.
This is the second species to become known in the genus
which was established for a species (N. migrans Chamberlin)
the types of which were taken at Philadelphia, similarly on
lily bulbs, from Saitama, Japan. The present form is sepa-
rated chiefly because of its larger size (7 mm. as against
5 mm.) and the fewer ocelli (1 + 3, 1 as against 1 + 3, 3.)
15. Bothropolys migrans Chamberlin, new species
Lateral margins of head smoothly continuous, not broken at level
of posterior suture. Antennae of moderate length.
Ocelli numerous, typically in four series; e. g., 1 -f- 6, 5, 4, 2.
Prosternal teeth typically 5 4- 5 or 6 -|- 6, small and subequal,
the outer end of dental line extending farther cephalad than the
mesal end. Special ectal spine setiform.
Posterior angles of ninth, eleventh and thirteenth dorsal plates
strongly produced, those of the seventh more weakly but distinctly
produced, and those of the sixth weakly or sometimes not at all
produced. Dorsal plates moderately rugose.
Anal legs with the claw single; ventral spines, 1, 1, 3, 2, 1; dorsal
spines, 1, 0, 3, 1, 0. Penult legs also with claw single; ventral spines,
0, 1, 3, 2, 1; dorsal, 1, 0, 3, 1, 1. Ventral spines of first legs 0, 0, 1,
3, 2, to 0, 0, 1, 3, 1. Last four coxae dorsally armed, the last two
laterally armed; anal coxae ventrally armed and in one specimen
one penult coxa is also ventrally armed with a small spine.
Claw of female gonopods tripartite, the lobes acute; basal spines,
2 + 2 .
Length, 11-14 mm.
Locality. The four type specimens were taken at Hawaii
in 1928 on plants of the genus Dioscorea from China.
United States Bureau of Entomology
The Brookings Institution has published as No. 60 of its
Service Monographs of the United States Government, a very
complete account of the history, activities and organization of
the Bureau of Entomology, by G. A. Weber, a book that will
be appreciated by all whose interests bring them into contact
with the activities of that department of the Government. In
addition to an interesting historical sketch the book includes a
list of the various divisions and field stations with an outline
of the staff of each and the salaries paid. It will prove a useful
reference book for all working entomologists. — E. P. Van
Duzee.
70
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
TWO NEW COLEOPTEROUS INSECTS FROM
ARIZONA
BY J. 0. MARTIN
California Academy of Sciences
Genus Euryptera
The genus Euryptera is distinctly American, ranging from
Rio de Janeiro, Brazil, through Central America, to as far
north in the United States as Massachusetts. Up to the present
there are twenty-six described species, of which four are
recorded from the United States, the greatest development of
the genus being in northern South America and Central
America. A table to nineteen of the species, based on color
alone, is offered by R. P. Belou in Ann. Soc. Ent. Belg., XLI,
p. 339, 1897. An examination of the original descriptions of
all the described species has convinced me that the species
described below is new.
Euryptera cruenta Martin n. sp.
Form elongate, widest just before apex of elytra. Head black
except at apex of clypeus and mouth parts which are testaceous,
variously clouded with black; irregularly punctured and with semi-
erect black hairs except a triangular space at base of clypeus which
is impunctate. Behind the antennal insertion is a deep transverse
depression which is more pronounced in this species, because of the
greater development of the antennal process, than in any of our
other native species. Antennae black, extending to beyond the middle
of elytra, Pronotum campanulate wider at base than long, dull
sanguineous, closely punctate, with red depressed fine hairs. Scu-
tellum black, closely punctate, with depressed black hairs. Elytra
wider than pronotum, just behind the base, and gradually widening
to just before the apices which are separately rounded, with a dis-
tinct obtuse angle at outer side of curve, but lacking any distinct
tooth such as is found at this position in lateralis Oliv. and many of
the more tropical species. Elytra of the same color as the pronotum
and with the same depressed red hairs but with smaller punctures
and with three distinct costas not as pronounced, however, as those
in huachuccB Schffr. Underside of body black, shining, finely punc-
tate, with black hairs which become yellow on the middle and hind
coxae; the first two joints of the abdomen more closely punctate than
the last three. Length, 11 mm.; width just before apex of elytra,
4 mm.
Euryptera cruenta differs from any of our other native spe-
cies in its color, vestiture, and the shape of the elytral apices.
OCTOBER, 1930] MARTIN NEW ARIZONA COLEOPTERA 71
It is nearest to longipennis Bates, but differs from that species
in the color of the thorax, in the lack of any constriction of the
sides of the elytra and in the possession of the angulation on
the outer curve of the elytral apex.
Described from a single specimen taken by myself in Carr
Canyon, Huachuca Mountains, Arizona, while beating oak.
Type, probably a female, No. 2976, in the collection of the
California Academy of Sciences.
Genus Ischnocerus
The genus Ischnocerus Schon, until now represented in
our fauna by a single species {infuscatus Fabr.), is more exten-
sively developed in the Mexican region from where the Biologia
Centrali Americana lists no less than seven species. In Jordan’s
table ^ to these species the one described below runs directly to
infuscatus, but is markedly different from that species.
Ischnocerus angulata Martin, n. sp.
Body piceous black, shining. Head and beak coarsely punctate;
eyes oval and slightly more than one-half wider than long, distant
from base of antenna by more than their major axis; front and beak
to base of antennae covered by chalky white recumbent hairs, a black
spot extending from apex of thorax nearly to the eyes; antennae
ferruginous, except club which is black; male antennae barely reach-
ing to end of body when beak is vertical, those of the female extend-
ing to basal third of elytra. Thorax closely, coarsely, rugosely,
punctate; disk with three longitudinal ridges, one, median, is sepa-
rated from the lateral ones by a broad shallow sulcus; the space
subtended by the summits of the lateral ridges and extending from
base to apex in a shield-shaped design is covered in varying degrees
of thickness by recumbent, chalky-white hairs. The remainder of
the prothorax is covered with ferruginous, black and a few white
hairs which present no regular pattern. Antebasal carina continuous
throughout, extended backward at middle in a broad, nearly ninety
degree, rounded angle which extends more than half the distance
to base of elytra; behind this angular extension is another patch of
white hairs extending to base of thorax. The portion of the pro-
thorax behind the antebasal is rugose and shows no trace of a sub-
basal Carina.
Elytra punctate-striate, the punctures coarse and shallow, shining,
distinctly showing through the vestiture, which is recumbent. Vesti-
ture of the sutural interspaces black with a mixture of white hairs.
The remainder of the elytral vestiture is ferruginous, mixed with
1 Biologia Centrali Americana-, Vol. IV, pt. 6, p. 305, 1906.
72
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
white except for the third, fifth, seventh and ninth interspaces, which
have a larger intermixture of black hairs. The third and fifth inter-
spaces are decorated with irregular tufts of elevated black hairs
roughly in pairs and more numerous behind the middle. In the post-
scutellar region the surface is elevated and has two black tufts.
Pygidium densely covered with chalky-white recumbent hairs.
Underside of body coarsely widely punctate, thinly covered with
white hairs which are more thickly placed laterally.
Legs ferruginous with a piceous band at middle of femora and
behind the middle of tibiae, darker at the joints. Thinly clothed with
white and black hairs. Length, 9 mm.; width, 3 mm.
The above species may at onee be recognized by the large
angulate backward extension of the antebasal carina on the tho-
rax. It differs from infuscatus by its smaller size, shorter male
antennae and more numerous elytral tufts of erect hairs, and by
the more pronounced whiteness of the front and thoracic areas.
Described from two male and four female specimens taken
by myself near Patagonia, Arizona, July 9, 1930, on the dead
branches of a large willow tree; also three females taken by
Mr. E. Gorton Linsley at the same time and under the same
conditions.
Holotype, a male, and allotype, female. Nos. 2977, 2978,
respectively, in the collection of the California Academy of
Sciences.
Paratypes in the above collection and that of Mr. E. Gorton
Linsley.
The Norman R. Gunn Collection of Butterflies
Mrs. Joseph Gunn has presented to the California Academy
of Sciences the collection of butterflies gotten together by her
son, Norman R. Gunn, whose untimely death was noticed in the
last number of this journal. There are 3300 specimens in this
collection, a part of which are in papers and a part are spread
for placement in Riker mounts. This makes a valuable addi-
tion to the Academy collection, including as it does a number
of local races but poorly represented in the Academy series. —
E. P. Van Duzee.
A Correction
Prof. Scullen has called my attention to an error on page 47
of Vol. VI of this journal (July 1929), where the reference
for Odynerus margaretellus Rohwer should have read (Proc.
U. S. Nat’l. Mus., XLIX, 242, 1915).— Editor.
OCTOBER, 1930 ] FALL NEW CALIFORNIA BEETLES
73
A NEW APHODIUS AND A NEW GENUS AND
SPECIES OF BUPRESTID^ FROM
CALIFORNIA
BY H. C. FALL
Tyngsboro, Massachusetts
Aphodius davisi Fall, n. sp.
Small, oblong, moderately convex. Head piceous with paler
margins; prothorax piceous, the lateral margins diffusely dull rufous;
elytra yellow, checkered with subquadriform black spots; femora
yellow, tibise and tarsi brownish; upper surface throughout polished
and strongly shining.
Head anteriorly coarsely punctato-rugulose, closely punctate but
smoother behind the frontal suture, which is perceptibly tritubercu-
late; clypeal emargination distinct, limited each side by sharply
defined subdentiform angulations which are not sinuate externally;
gense moderately prominent, rounded.
Prothorax three-tenths wider than long, only slightly narrower
in front, sides feebly arcuate, hind angles well defined, somewhat
obtuse; base arcuate, just perceptibly sinuate each side of the middle,
marginal line very fine; disk sparsely finely punctate with still
sparser intermixed much coarser punctures which, however, are
somewhat more numerous at sides.
Elytra not quite as wide at base as prothorax, slightly widened
posteriorly; strise finely rather closely punctate; intervals nearly flat,
finely sparsely punctate. The markings consist of three longitudinal
series of alternating irregularly quadrate spots, each spot occupying
the width of two interspaces. The first series is made up of three
spots on intervals 2 and 3, the first spot subbasal; the next series of
three on intervals 4 and 5 with first spot basal; the third series on
intervals 6 and 7 has but two spots, these connected externally by a
line on the eighth interval, which may extend forward to the humeral
umbo and externally nearly to the margin.
Mesosternum not carinate between the coxae; metasternum
smooth and sparsely punctate medially, alutaceous and with coarser
punctures at sides; abdomen alutaceous with a few fine punctures.
Anterior tibiae tridentate, crenulate above, smooth in front; hind
tibiae fringed at apex with unequal spinules; basal joint of hind
tarsus not quite as long as the next three.
Length, 3.2 to 3.7 mm.; width, 1.55 to 1.9 mm.
Irvine Park, Orange County, California.
Described from ten examples sent me by Mr. A. C. Davis
of Santa Ana, California, who with associates took them from
74
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
the nests of wood rats (Neotoma). The type is one of the
smaller examples and is probably a male, although I observe
no positive sexual differences in the series at hand; it bears
date “2-1-30.”
When not darkened by alcohol or otherwise this is a very
pretty little species with its black thorax- and bright yellow
sharply spotted elytra. A slight tendency toward variability in
maculation is observed, which might become more marked in
a longer series.
By Horn’s Monograph davisi runs to serval in Group “I-d.”
It agrees nearly with serval in clypeal outline and sculpture of
the head, and in a general way in coloration; serval, however,
is a larger species, the elytral maculation quite different in
detail, the pronotal punctuation much less disparate, the legs
uniformly reddish brown, the tibial spurs distinctly sexually
modified.
The type and two paratypes have been retained for my own
collection; the remaining paratypes are returned to Mr. Davis,
in whose honor the species is named in recognition of generous
contributions to my cabinet and years of pleasant personal
acquaintance in California.
Hesperorhipis Fall n. gen. (Buprestidse)
Sixty-four years ago (December, 1866), Dr. Leconte
erected the genus Xenorhipis for a unique male specimen of
a small Buprestide having flabellate antennae. It has been stated
by Horn (1882) and later by Kerremans that there is no other
known instance of this type of antennae throughout the entire
family; it will therefore be of more than ordinary interest to
announce that there has come to me recently from California a
unique male of another very small Buprestide having similarly
flabellate antennae. In general makeup this species shows a
close relationship with Xenorhipis, but in several respects is
not strictly congeneric and I would, therefore, suggest for it
the above new generic title. The chief differences are indicated
below in parallel columns and these, combined with certain
characters in the specific description, will constitute a suffi-
ciently distinctive diagnosis of the new genus.
OCTOBER, 1930] FALL NEW CALIFORNIA BEETLES
75
Xenorhipis
Antennal rami beginning at
the second joint.
Last joint of maxillary palpus
scarcely or feebly narrowed api-
cally, the tip broadly truncate.
Elytra of normal length, the
pygidium alone exposed.
Sides of metasternum with a
large hairy depression or exca-
vation.
Posterior coxal plates broad,
scarcely narrowed externally.
Basal joint of hind tarsus but
little shorter than the remaining
joints taken together.
Hesperorhipis n. gen.
Antennal rami beginning at
the third joint.
Last joint of maxillary palpus
ovate conical, the apex pointed
or at most very narrowly trun-
cate.
Elytra shorter, the last five
dorsal abdominal segments ex-
posed.
Sides of metasternum not at
all excavated or depressed.
Posterior coxal plates triangu-
lar, the hind margin strongly
oblique.
Basal joint of hind tarsus
equal in length to the three fol-
lowing joints and constituting
scarcely more than one- third the
tarsal length.
Hesperorhipis albofasciatuis Fall n. sp.
Very small, head, thorax and elytra of nearly equal width; length
to tip of elytra 2.6 mm., to abdominal apex 3.25 mm., width of elytra
at base barely 1 mm.
Color black, head and thorax with deep blue surface luster; elytra
each with a transverse slightly oblique narrow parallel-sided white
fascia just behind the middle, the two forming a very obtuse angle
at the suture. Before the white fascia the elytra are of a brighter
blue in a triangular area involving the entire width at base, the
sides converging to the suture at the white fascia. Behind the fascia
the elytra are black with barely detectable bronze luster. Body
beneath black, faintly bluish; tibiae and tarsi pallescent.
Antennae ( ^ ) dull testaceous, flabellate, almost precisely as in
Horn’s figure for Xenorhipis (Trans. Am. Ent. Soc., 1882, PI. IV),
except that the rami begin at the third joint.
Head almost as wide as the prothorax, front flatly convex, a
vague longitudinal impression terminating above in a small vertical
fovea; clypeus scarcely margined in front, very feebly so over the
antennae; surface densely, coarsely, shallowly reticulate. Eyes verti-
cal, rather large but only moderately prominent.
Prothorax transversely quadrate, slightly more than one-third
wider than long; sides parallel, a little rounded in front and just
visibly sinuate posteriorly, hind angles sharply rectangular; base
bisinuate; side margins fine, becoming subobsolete: at the front
angles; disk broadly flattened medially and with a slight trace of a
postmedian transverse impression; sculpture same as on the head.
76
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
Elytra very little wider than the thorax, seven-tenths longer
than wide, feebly narrowed posteriorly; not very densely but rather
roughly asperately punctate except in the white fascia, which is
straight, perceptibly impressed, extends slightly forward from the
suture to the side margin, and is without visible sculpture.
Body beneath closely punctate; prosternum vaguely impressed
medially, intercoxal process as in Xenorhipts, long, gradually nar-
rowed, apex acute, sides not at all angulate. Ventral segments
throughout coarsely punctate, last segment deeply roundly emargi-
nate. Tarsi as in Xemrhipis, the third and fourth joints with short
membranous lobes; claws nearly simple, a feeble obtuse subbasal
angulation.
The unique representative of this remarkable little species
was beaten from mistletoe at Fort Tejon, California, VIII-3-30,
by Mr. A. C. Davis, and was one of a box full of miscellaneous
things sent me for identification. Mr. Davis writes me that he
made strenuous efforts to obtain further specimens, but with-
out success. The type remains in my collection.
Argyresthias Found in Golden Gate Park,
San Francisco
There are four species of these small golden or silvery
moths found in this locality, and their differences may be
summed up in the following synopsis.
A. Argyresthia pilatella Braun. Adult with 7-9 mm. wing
expanse, dull silvery or grayish, a dark mark on the middle
of the hind margin of the forewings ; larva reddish, mining
needles of Pinus radiata Don. from base; pupa in rough
cocoon spun in contact with a needle or other substance.
B. Argyresthia cupressella Wlshm. Adult with 7-9 mm.
expanse, wings golden, head tuft white ; larva yellow
green, ochreous head, mining in tips of twigs of Cupressus
macrocar pa Hartw. ; pupa in rough cocoon spun in con-
tact with two or more surfaces (among twigs).
C. Argyresthia trifascice Braun. Adult with 6-8 mm. expanse,
wings golden, head tuft brown; larva deep green, black
head, pupa in smooth cocoon spun on one surface only
(Larvae collected from C. macrocar pa twigs).
D. Argyresthia franciscella Busck. Adult with 8-9.5 mm.
expanse, wings dull golden, almost smoky, and crossed by
about three blackish lines, head tuft whitish ; larva sordid
brownish and found within terminal “cell” made of twigs of
C. macrocarpa ; pupa inside cell, no cocoon. — H. H. Keifer.
OCTOBER, 1930] LINSLEY POGONOCHERINI
77
NEW POGONOCHERUS AND ECYRUS (COLEOP-
TERA, CERAMBYCID^) WITH NOTES
CONCERNING OTHERS
BY E. GORTON LINSLEY
Oakland, California
The Pogonocherus and Ecyrus are the most closely related
genera of the Pogonocherini, and are classified by Leconte and
Horn as the subtribe Pogonocheri. As there has been no major
paper on Ecyrus in recent years, and since the last papers on
Pogonocherus were in 1909,^ 1910,^ and 1913,^ it seems advis-
able to bring these genera up to date in regard to recently
acquired knowledge and the addition of new species. Appre-
ciation is expressed to Dr. E. C. Van Dyke of the University
of California for the loan of certain species discussed in this
paper, and for reading the manuscript and offering many help-
ful suggestions. For the two excellent drawings, I am indebted
to Mrs. Frieda Abernathy of Berkeley, California.
Pogonocherus Latr.
The members of this genus are separated from all the other
genera which have heretofore been included in the tribe Pogo-
nocherini, by the long flying hairs of the antennas, legs, and
body, and the presence of lateral tubercles on the prothorax.
The species are nocturnal and are usually restricted to certain
types of food plant. The adults are found, usually in the shade,
on dead and dying twigs. Storm-broken branches and wind-
falls are especially attractive to them. The majority of the spe-
cies are North American, although the genus is well represented
in both Europe and Asia. The recent European writers have
divided Pogonocherus into two groups, those species having
truncate-emarginate elytral apices, and those having rotundate-
truncate elytral apices. This classification can be conveniently
followed in regard to the North American species, and is more
nearly phylogenetically correct than the grouping used by Horn,
Schaeffer, Fall, and Casey. L. M. Planet ^ places the species
1 Schaeffer, C. W. Four New Cerambycidae, Journal New York Ent.
Soc., XVIII, 1909, pp. 99-103.
2 Fall, H. C. New Species of Pogonocherus, Ent. News, XXI, 1910,
pp. 5-9.
3 Casey, T. L. Memoirs of the Coleoptera, IV, 1913; Longicornia,
pp. 345-349.
4 Planet, L. M. Hist. Nat. des Longicornes de France, Paris, 1924,
pp. 280-289.
78
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
with emarginate-truncate apices in Pogonocherus (s. str.) and
those with truncate or rotundate-truncate apices in the sub-
genus Pityphilus Muls. The use of subgenera for these two
groups seems a trifle extreme, but the division is a convenient
one to follow in forming a key, and is the method used in this
paper. Leng, in his Catalogue of the Coleoptera of America
North of Mexico, lists nineteen species of Pogonocherus (using
the genus in its broadest sense), but upon close examination,
there appears to be not more than fourteen known forms,
including the one described in this paper, that are worthy of
specific rank. The following table will separate the species :
Key to the North American Species of Pogonocherus
Elytral apices emarginate-truncate {Pogonocherus s. str.) 1
Elytral apices truncate or rotundate-truncate, but not emarginate
{Pityphilus) 6
1. Elytra with tufted subbasal tubercle; lateral tubercle of pro-
thorax obtuse or blunt at apex 2
Elytra without subbasal tubercle; lateral tubercle of prothorax
acute at apex 3
2. Elytral apices distinctly spiniform, pubescence gray, flying
hairs long. 6-10 mm. Northern California.... (1) crinitus Lee.
Elytral apices not spiniform, pubescence grayish brown, elytra
with an oblique antemedian pale marking. 5.5-9 mm.
California (2) propinquus Fall
3. Elytra with two or more postmedian subsutural tufts of erect
black setee. 6 mm. Southern Arizona.... (3) arizorairuj Schffr.
Elytra without tufts of setae 4
4. Elytra subcostate, antennal scape stout. 5. 5-7. 5 mm. Colo-
rado, New Mexico,, Arizona (4) pictus Fall
Elytra without costae, antennal scape less stout 5
5. Prothorax about as wide as long; outer angle of elytral apices
bluntly dentate; body with numerous suberect long hairs.
4.5-6. 5 mm. Central and eastern North America
(5) mixfus Hald.
Prothorax fully one-half wider than long, outer angle acutely
dentate; suberect hairs short or missing. 4.5-6 mm.
Atlantic Coast (6) salicicola Csy.
6. Lateral tubercles of prothorax obtuse or blunt at apex 7
Lateral tubercles of prothorax acute at apex 14
7. Elytra distinctly tricostate 8
Discal costse of elytra indistinct or evanescent at base; lateral
costae more or less evident
9
OCTOBER, 1930] LINSLEY POGONOCHERINI
79
8. Elytra with a short extra humeral carina between lateral and
median costae; eyes with large callosities above; elytra
two-thirds as wide as long. 8 mm. Southern Texas
(7) vandykei n. sp.
Elytra without extra humeral carina; eyes lacking callosities;
elytra only half as wide as long. 6-11 mm. Southern
California (8) pilatei Van D.
9. Prothorax with small shining tubercle on basal half of median
line; elytra with large postmedian white spot, subsutural
costae with tufts of erect black setae. 4.5-6 mm. Alaska,
Canada, and eastern United States (9) penicillatus Lee.
Prothorax without median tubercle; postmedian elytral white
spot never present 10
10. Elytra with a distinct antemedian yellowish brown V-shaped
area; basal region blackish; lateral pubescence gray, costae
with a few subsutural tufts of short black hair in post-
median area. 6-8.5 mm. Southern Arizona
(10) negundo Schffr.
Elytra lacking yellowish patch; either uniformly gray, gray
and black, or with a distinct postmedian black band 11
11. Elytra with a distinct postmedian black band 12
Elytra more or less uniforml}" gray, black markings when
present either vague, basal, or sutural 13
12. Postmedian black band broad at suture; discal tubercles of
prothorax indistinct. 6-8.5 mm. California, Oregon,
Idaho, and Colorado (11) oregonus Lee.
Postmedian black band narrowed at suture; discal tubercles
of prothorax prominent. 6-9.5 mm. California
(12) calif ornicus Schffr.
13. Body generally clothed with a uniform gray pubescence, some-
times with a vague postmedian black bar, or with a
velvety black V-shaped postscutellar patch extending
backwards as a sutural vitta which generally dilates into
a more or less triangular area in apical region; series of
small black tufts of hair on elytra often present but never
forming four series. 6.8 mm. Central California
(12a) californicus concotor Schffr.
Body clothed with blackish and ashy-gray hairs, the latter
predominating, especially in the antemedian discal elytral
area, the elytra also with four series of small black tufts of
short hairs, the inner series close to the suture. 7-9 mm.
Northern Arizona (13) obscurus Fall
14. Antennal scape long, slender, attaining lateral thoracic tuber-
cle; body clothed with, a dense pale brown pubescence,
elytral apices, sides, and basal area darker brown; median
elytral area gray; flying hairs long. 5-8 mm. Lower
California (14) volitans Lee.
80
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
(1) POGONOCHERUS CRINITUS Lee.
This is one of the largest and most distinct Pogonocherus.
It is more closely related to certain European species, espe-
cially perroudi Muls, than to any North American form. It is
of a general gray color with very long flying hairs on antennae,
legs, and body, and is easily separated from the other members
of the genus by the acute apical spine of the elytra and the
prominently crested subbasal tubercles. It is found throughout
the coastal mountain ranges of California from Monterey
County to Mendocino County, and is recorded in literature
from Vancouver Island, B. C. Host plant, Quercus agrifolia
Nee.
(2) Pogonocherus propinquus Fall
Although some\(rhat resembling penicillatus Lee., this spe-
cies is separated by the emarginate clytral apices and the
obliquely set elytral pale markings. Although described from
southern California, it is more commonly found in the northern
and eastern portions of the state. Its range is continuous from
the mountains of Oregon and northern California, through the
Sierra Nevadas, to the San Bernardino Mountains on the south.
It is normally taken at high altitudes on the western yellow
pine, Pinus ponderosa Laws.
(3) Pogonocherus arizonicus Schffr.
This species is closely allied to P. mixtus Hald. and P. pictus
Fall. It differs from them in having two or more postmedian
tufts of erect black setae on the subsutural costae of the elytra.
Mr. Schaeffer, in his original description of the species, states
that there are three such tufts. A specimen collected by the
writer in the Huachuca Mountains, Arizona (type locality),
has only two tufts of setae, although it agrees with description
in all other respects. Neither mixtus nor pictus have tufts of
setae and are therefore easily separated from arizonicus. The
last differs from penicillatus , the only other species with such
conspicuous tufts, in having emarginate elytral apices and in
entirely lacking subbasal elytral tubercles. Habitat, the moun-
tainous regions of southern Arizona, probably on pine.
(4) Pogonocherus pictus Fall
In this species the elytral apices vary greatly as to trunca-
tion and dentation. Colonel Casey in his description of emargi-
OCTOBER, 1930] LINSLEY POGONOCHERINI
81
natus (later stated by its author to be a synonym of pictus),
speaks of the elytral apices as narrowly and deeply sinuate, and
the sutural tooth as longer than the lateral tooth, which he
describes as sharply marked but not prolonged. Certain speci-
mens before me from Long’s Peak Inn, Colorado (Van Dyke
collection), lack the lateral tooth entirely, while others in the
same series agree with Colonel Casey’s specimen. A specimen
from the Catalina Mountains, Arizona, is at the other extreme,
with a well-defined lateral tooth and a poorly developed sutural
tooth. The general color pattern of pictus is similar to mixtus
Hald from which it may be separated by its larger size, sub-
costate elytra, and stout antennal scape. It is typically a Rocky
Mountain species, ranging from Montana and Colorado, down
into Arizona and New Mexico. P. fastigiatus Csy. is merely
a variation of pictus Fall.
(5) PoGONOCHERUS MIXTUS Hald.
This is one of our smallest species. The prothorax is about
as wide as long, color black to piceous, with large pale elytral
markings. The punctuation of the prothorax and elytra is very
variable. The species is widely spread throughout eastern and
northern North America on both coniferous and broad-leaved
trees. P. parvulus Lee. is merely a small form of mixtus.
P. simplex Lee. was placed by Fall, after a critical examina-
tion of the type, as a synonym of mixtus. This fact seems to
have been overlooked by Casey, although the latter’s paper was
published three years later than Fall’s. Leng, in his Catalogue
of the Coleoptera, followed the latest writer and included
simplex in his list. The species should, however, remain as a
synonym of mixtus Hald.
6. PoGONOCHERUs SALicicoLA Casey
(F. salicola Casey®)
The size, color pattern, and superficial appearance of this
species is very similar to mixtus Hald., although the wide pro-
5 When Casey first described this species in 1913, he had in mind the
fact that it is found on S a 1 i x , but the name given to it, salicola,
unfortunately gives a meaning very different from that intended by its
author. Five years after Casey’s description, Mr. F. J. A. Morris, in the
Canadian Entomologist, Vol. 50, p. 41, pointed out the error in the Latin
stem of the word. Soon after the publication of Mr. Morris’ article,
Casey, in Vol. 8 of his Memoirs of the Coleoptera (p. 415, 1918), changed
the name to salicicola, stating that the original spelling was a mis-
print. Under the rules of nomenclature, salicicola will be valid.
82
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
thorax, the acute apical denticles of the elytra, and the scarcity
of erect black hairs on the body, easily separate it. It is found
throughout the Atlantic States on willow, Salix.
(7) Pogonocherus vandykei Linsley, n. sp.
Robust subcylindrical, piceous, clothed with a rather sparse, short,
gray and brown pubescence, with a few longer, scattered flying hairs,
on antennae, legs and entire upper surface. Head broad, distinctly
quadrangular when viewed from the front, longitudinally sulcate
between the antennae; eyes reniform, lower portion subtriangular,
with large callosities above. Antennae, annulate, longer than body;
scape stout. Prothorax, wider than long, about two-thirds as wide
as elytra at base; lateral and discal tubercles, large, obtuse. Scu-
tellum, white. Elytra about three-fifths as wide as long, tricostate,
with a short extra humeral carina between median and lateral costae;
subsutural costa elevated near base into a large tubercle, promi-
nently crested with long black hairs, and again in postmedian area
into a ridge of small black tufted tubercles; median costa with a
series of similar but smaller tubercles in same region; apices trun-
cate; general color pattern, grayish brown, with a lunate patch of
gray pubescence extending from humerus to suture at middle, and
margined posteriorly with an irregular black line; also with a row
of yellow hairs along inner two costae in antemedian gray area;
apices gray; subbasal tubercles connected with a narrow black line.
Beneath, clothed with a sparse gray pubescence. Underside of tarsi
padded with soft yellow hairs. Length, 8 mm.; breadth, 3 mm.
Type (No. 2979, Mus. Calif. Acad. Sci.), a unique, prob-
ably a female, collected by the writer at Uvalde, Texas, June 13,
1930. I take pleasure in naming this species after Dr. E. C.
Van Dyke, as a slight tribute for many favors.
In this species the elytra are much shorter in proportion to
the total length of the insect, and the subbasal tubercles much
more prominently crested than is the case in any other of the
known forms. The former character gives the species a short
robust appearance that is quite distinctive. The general color
pattern is very similar to that of pilatei or calif ornicus. The
general body proportions, the extrahumeral carina, and the less
rotundate elytral apices, will, however, easily separate it from
either of these two species. It further differs in the large cal-
losities above the eyes, the sparse pubescence, and the presence
of two rows of yellow hairs along inner two costae in the ante-
median gray area of the elytra.
October, 1930] linsley — pogonocherini
83
(8) PoGONOCHERUS PiLATEi Van Dyke
The average size of this species is greater than the average
size of any other known Pogonocherus. It differs from all
other species in the second group except vandykei in having
distinctly tricostate elytra. The characters given above will
separate it from the latter species. It is found in the foothill
regions of southern California on the peculiar shrub Fremonto-
dendron calif ornicum Torr.
(9) Pogonocherus penicillatus Lee.
This is a grayish-brown species with a large white spot on
each elytron and tufts of erect black setae on the discal elytral
costae. There is also a small shining tubercle on the median
line of the prothorax. It is distributed throughout eastern
United States, Canada, and Alaska. After critically examining
Schaeffer’s description of P. alaskanus as well as many typical
specimens of the species, the only character that I can find to
separate it from penicillatus Lee., is the rounded elytral apices.
However, examination of a series of each of these species
shows this character to be inconstant. Certain specimens from
Fort Yukon and Rampart, Alaska, the same general territory
in which typical alaskanus is found, have definitely truncate
elytral apices, which would classify them as penicillatus and
from which they cannot be separated. Other specimens from
eastern United States have rotundate-truncate apices as found
in alaskanus. With extreme and intermediate phases to be
found in both regions, and since there seems to be no other
character with which to separate these forms, I am convinced
that alaskanus Schffr. is merely a variant of P. penicillatus Lee.
P. carinatus Csy. is an intermediate phase between typical
penicillatus and the form alaskanus. It was described from a
specimen taken in Colorado, the western limit of the range of
penicillatus.
Pogonocherus fasciculatus De Geer
This European species is listed from Greenland in our
literature, but since this locality has remained unverified, I have
not included it in the table. In the key it would run to penicil-
latus, which it superficially resembles but from which it can be
separated by the indistinct subbasal tubercles of the elytra and
84
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
the more acute lateral spines of the prothorax. Five to seven
mm. Europe and Greenland ( ?).
(10) POGONOCHERUS NEGUNDO Schffr.
In this species the elytra have an oblique yellowish-brown
band which extends from the lateral margin in the subhumeral
region to the suture at middle. The basal elytral area is much
darker. The species is closely allied to calif ornicus Schffr. from
which it differs in the general color pattern and the less distinct
longitudinal sulcus between the antennae. It is found on box
elder, Acer negimdo Linn., in the Huachuca Mountains of
southern Arizona.
(11) PoGONOCHERUS OREGONUS LeC.
This species is closely related to calif ornicus Schffr. and
differs from it in having the postmedian black band of the
elytra about as broad at the suture as at the lateral edge. In
addition, the discal prothoracic tubercles are much less distinct.
This is an alpine form, most commonly taken in the high moun-
tains from Tulare County, California, to Mount Hood, Oregon.
Specimens in the Van Dyke collection from British Columbia
and Mount Timpanogos, Utah, show that the distribution con-
tinues up the Pacific Coast and back down the western ranges
of the Rocky Mountains. This species is always taken on the
true firs, Abies.
(12) POGONOCHERUS CALIFORNICUS Schffr.
Typical specimens of this species are characterized by hav-
ing a gray antemedian V-shaped area on the elytra, a saddle-
like postmedian black band narrowed at the suture, and the sub-
basal elytral tubercles crowned with tufts of black hair. It is
usually found in the high mountains of northern and eastern
California on the western yellow pine, Pinus ponderosa Laws,
and in Arizona on Pinus edulis Engln. and Pinus jeffreyi
Vasey.®
(12a) POGONOCHERUS CALIFORNICUS CONCOLOR Schffr.
This is a very distinct subspecies of calif ornicus. It is
extremely variable within the subspecies, but is easily separated
from the typical form, which is quite constant as to color. The
6 Webb, J. Lf. Proc. Ent. Coc., Washington, Vol. X, 1908, p. 119.
OCTOBER, 1930 ] LINSLEY POGONOCHERINI
85
form concolor varies from a uniform gray, through a phase
suggestive of typical calif ornicus, but much lighter, to a very
divergent form with velvety black pubescence along the suture.
In addition, typical calif ornicus is a high altitude form taken
on the western yellow pine, whereas concolor is always found
in the dry foothill regions on the Digger pine, Pinus sabiana
Dough
(13) POGONOCHERUS OBSCURUS Fall
This species is very close to calif ornicus Schffr., especially
to its subspecies concolor Schffr. The last differs from Fall’s
description of obscurus in one point only, namely, that obscurus
is stated to have four series of small black tufts of short hairs
on the elytra. In a series of twenty-four specimens of concolor
from Tulare County, California, a number of individuals have
more than one series of such tufts, several have only one series,
and many lack the tufts of hair altogether. In none of the
specimens before me, however, are there four series. Up to
the present time, concolor has only been taken in the Sierra
foothills on Pinus sabiana, but calif ornicus has been taken in
Arizona on Pinus edulis and P. jeffreyi. Fall’s specimens
of obscurus were taken in a region where pine is abundant.
Although I have not seen obscurus, I am inclined to believe
from Fall’s description that when a series of the concolor
variety of calif ornicus is examined with a series of obscurus,
the two may be found to run very near together. Type locality
of obscurus. Bright Angel, Arizona.
(14) PoGONOCHERUS VOLITANS Lec.
The long slender antennal scape of this species easily sepa-
rates it from the other members of the genus, although the color
pattern is somewhat suggestive of negundo Schffr. So distinct
is the antennal scape of this species that its author first placed
it in another genus. All other characters are, however, typi-
cally those of Pogonocherus. Type locality. Cape San Lucas,
Lower California.
Ecyrus Le Conte
The members of this genus are confined to eastern and
southern North America, and differ from Pogonocherus in
having a feebly rounded prothorax without lateral tubercles.
The antennae are clothed along the inner side with flying hairs
86
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
which may or may not be present on legs and body. There
appear to be only two valid species, including the one described
in this paper, that are found within the United States. A third
species from the West Indies and a questionable form from
Mexico bring the total of recognizable species up to four.
These may be separated as follows :
Key to the North American Species of Ecyrus
Prothorax with two large, laterally compressed, discal tubercles;
elytra with two more or less prominently crested subbasal
tubercles 1
Pro thorax without large discal tubercles; subbasal area of elytra
more or less evenly convex 2
1. Discal tubercles of prothorax prolonged backwards into a dis-
tinct horn, discal area with large quadrangular patch of
white pubescence; elytra and upper surface clothed with
long flying hairs, pubescence brown and white. 5-6.5 mm.
Southern Texas (1) cornutus n. sp.
Discal tubercles of prothorax not prolonged backwards into a
horn, discal area clothed with tawny-brown pile; elytra and
upper surface without flying hairs, pubescence blue gray.
13 mm. West Indies (2) hirtipes Gah.
2. Elytra with three black lines of hair pencils; anterior discal
area of prothorax broadly convex, finely tuberculose.
8-11 mm. Mexico (3) penicillatus Bates
Elytra without definite hair pencils; prothorax more or less
evenly convex. 6.5-11 mm. Atlantic Coast and Gulf States
from Canada to southern Mexico (4) dasycerus Say
(1) Ecyrus cornutus Linsley, n. sp.
(Figs. 1 and 2)
Robust, subcylindrical, color varying from brown to piceous,
clothed with a rather dense, short, closely appressed pile, in addi-
tion to numerous suberect long hairs. Head transversely impressed
between the eyes and longitudinally sulcate between the antennae;
pile brown at base of antennae, white on face and mouth parts.
Antennae annulate, slightly longer than the body in female, distinctly
longer in male; scape stout, second segment about as long as wide,
third about five times as long as second, fourth about one and one-
third as long as third, fifth about one-third as long as fourth, sixth
and following segments diminishing in length toward apex. Pro-
thorax two-thirds as wide as elytra at base, about as wide as long,
without lateral spines, sides slightly arcuate; disk convex, with a
laterally compressed tubercle arising from each side, which is pro-
longed backwards into a distinct horn; discal area clothed with a
short dense white pubescence, supplemented laterally with longer
OCTOBER, 1930]
LINSLEY POGONOCHERINI
87
Fig. 1. Ecyrus cornutus Linsley
88
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
brownish hairs. Scutellum white. Elytra twice as long as wide,
basal portion of disk somewhat depressed; third interval elevated
near base into an elongate tubercle and again near middle into an
elongate ridge which gradually diminishes toward apices; also a
secondary ridge extending from humerus in an arcuate manner to
almost unite with median ridge at middle; enclosed within this ridge
is a lunate patch of white pubescence extending from third interval
to sutural margin, and a second smaller patch just posterior to sub-
basal tubercles, lateral slope of ridge with two postmedian black
spots. Elytral intervals somewhat flattened, with a scattering of
minute tufted tubercles throughout; apices truncate-emarginate.
Basal area of elytra generally brown; beneath, sides, and elytral
apices, gray. Underside of third tarsal segment clothed with pale-
yellow hairs, the first and second segments clothed with darker,
brownish hairs. Length, 6 mm.; breadth, 2.5 mm.
Holotype, male (No. 2980, Mus. Calif. Acad. Sci.), and
allotype, female (No. 2981, Mus. Calif. Acad. Sci.), and twelve
paratypes in my own collection were taken by the writer at
Brownsville, Texas, June 24 and 25, 1930. All of the speci-
mens were beaten from dead willow branches and show little
variation except in size (5-6.5 mm.).
This attractive insect, taken in the peculiar subtropical
floral region of Brownsville, might at first glance be mistaken
for a Pogonocherus. The absence of lateral tubercles on the
prothorax, however, place it in Ecyrus. It differs from all the
known species of the latter genus, in the elytral sculpturing,
and in being entirely covered with long flying hairs. In all of
the described forms of Ecyrus these hairs are more or less
evident on the legs and antennae, but in no other case are they
found on the upper surface of the body as in cornutus. In the
latter character, which gives the species the appearance of a
Pogonocherus, it differs from Le Conte’s original description
of Ecyrus, but as it seems to be congeneric in all other respects,
I have placed it in this genus. The species shows a relation-
ship to the West Indian, E. hirtipes Gahan, in the presence of
discal prothoracic tubercles, but the shape of these tubercles,
elytral sculpturing, size, and general color pattern, easily sepa-
rate it. E. cormitus is also very distinct from E. dasycerus
Say, differing in the large discal horns of the prothorax, the
prominent subbasal elytral tubercles, and the long flying hairs
covering the entire surface. The average size in my series of
cornutus is 6 mm.
OCTOBER, 1930]
LINSLEY POGONOCHERINI
89
Fig. 2. Side view of Ecyrus cornutus Linsley
(2) Ecyrus hirtipes Gahan
This species was described from Grenada, West Indies.
The average size, 13 mm., is greater than that of any other
species of the genus. It is of a general bluish gray color, with
flying hairs on antennae, legs, and undersurface. The elytral
costae are more or less distinct throughout, the median one
raised to form a tufted tubercle in subbasal region. The spe-
cies is very distinct and may be further separated by the
characters given in the key.
(3) Ecyrus PENCiLLATUs Bates ^
This species was described by Bates, from Vera Cruz,
Mexico. It is evidently very close to E. dasycerus Say, the
range of which extends well into Mexico. The author states
that the anterior discal area of the prothorax is broadly convex
and finely tuberculose, but this condition is also found in cer-
tain variations of dasycerus. The presence of hair pencils on
the elytra, although sometimes suggested in dasycerus (as in
variety fasciatus Ham.), may be a good character, but until a
series of this species can be critically examined in connection
with a series of dasycerus, the correct standing of this species
cannot be determined. Size, 8-11 mm.
(4) Ecyrus dasycerus Say
This species is commonly found along the Atlantic coast and
in the Gulf states, extending into Mexico. It is very variable,
especially in the southern portion of its range, and many forms
and variations have been named. It ranges in size from 4-11
7 Gahan, C. T. Longicorn Coleop., West India Island, Trans. Ent.
Soc., London, 1895, p. 127.
8 Bates, H. W. Biologia Central! Americana, V, 1880, Longicornia,
p. 137.
90 THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
mm., and in color from cinereous in the more typical forms
to silvery white in the variety fasciatus Ham. The flying hairs,
although moderate on antennae, are very short or missing on
legs and body.
(4a) Ecyrus dasycerus fasciatus Hamilton
I have before me a number of specimens of a light-colored
phase of dasycerus from Florida, collected by Mr. J. N. Knull
(Miami), and Mr. W. S. Blatchley (Royal Palm Park), that
agree with Hamilton’s description of fasciatus. The only char-
acter that I can find in them or the description that will sepa-
rate fasciatus from the typical dasycerus is the whiter pubes-
cence. The Other characters mentioned by Hamilton, such as
the three large tufts of coarse black bristles and the different
type of thoracic punctuation may also be found in a series of
the typical dasycerus. After critically examining a long series
of dasycerus as well as a number of probable fasciatus, in con-
nection with Hamilton’s description, I am convinced that the
latter is nothing more than a color variety of E. dasycerus Say. .
(4b) Ecyrus dasycerus texanus Schffr.
Although the description of this variety is rather indefinite,
the author states that it differs from the typical form in the
presence of two postmedian elytral fasciae. Such fasciae are
vaguely seen in f asciatus as a part of the general color scheme.
A series of specimens taken by the writer at Brownsville, Texas
(type locality of both fasciatus and texanus), have these fasciae
indistinctly formed, but in all other respects they agree with
the typical E. dasycerus Say.
(4c) Ecyrus dasycerus arcuatus Gahan®
An examination of the original description of this species
as well as the illustration accompanying it, shows it to be very
close to E. dasycerus, especially the variety texanus Schffr. As
there seem to be no characters to separate it from dasycerus,
except the large white spaces on the elytra, which, as mentioned
above, are seen in certain variations of the latter species, I have
placed it as a variety. Should it later prove to be the same as
the variety texanus, arcuatus would have priority.
9 Gahan, C. T. Longicornia of Mex. and Cent. Am., Trans. Ent. Soc.,
London, 1892, p. 259.
October, 1930] bequaert — polistella
91
ASHMEAD’S GENUS POLISTELLA
(HYMENOPTERA, VESPID^)
BY J. BEQUAERT
Department of Tropical Medicine y Harvard University
Medical School, Boston, Mass.
In 1904 (Proc. U. S. N. Mus., XXVIII, p. 133), Ashmead
proposed the new generic term, Polistella, for Polistes manil-
lensis H. de Saussure, of the Philippines, giving the following
characters to separate it from Polistes Latreille : “Metathorax
smooth, not transversely striate; abdomen fusiform, subpetio-
late; second cubital cell very small and narrow, along the
cubitus hardly one-third the length of the third, receiving both
recurrent nervures close together; eyes extending to base of
mandibles.” I have long been at a loss to place this genus, since
some of its characters are found in Polistes, while others (such
as the peculiar venation) could hardly be expected in the Polis-
tinae. Ashmead’s figure of his ‘^Polistella manillensis'' (PI. I,
Fig. 1) looks more like one of the Polybiinae, though it is quite
different from any Oriental member of that subfamily known
to me. In the “Revision of the Vespidae of the Belgian Congo”
(1918, Bull. Am. Mus. Nat. Hist., XXXIX, p. 255), I was
unable to reach any definite conclusion regarding the status of
Polistella.
During a recent visit to the United States National Museum,
I had the opportunity to examine the material used by Ashmead
to draw up his generic description. I was greatly surprised to
discover that his insect was not one of the Polistinae, nor
in any way related to Polistes, but one of the Neotropical
Polybiinae.
The material comprises two females (or workers), one well
preserved, the other lacking the abdomen, but both of the same
species. They bear no locality labels, but are marked “Col-
lection Fitch,” and one of them has a label in Fitch’s hand-
writing: “Smaller maribaudes, April 15, ’56.” Each pin in
addition carries a small diamond of red paper. In attempting
to trace the origin of these specimens, I was told by Mr. H. S.
Barber that, in Fitch’s original collection, a red piece of paper
placed on a pin seemed to indicate an insect of Neotropical
origin. This supposition is strengthened by the reference to
92
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 2
“maribaudes,” which is a vernacular name applied in the
Guianas to some of the social wasps. The name is mentioned
by many writers, usually in a corrupted form. In Surinam,
for instance, it becomes “marribu” (Christ, 1791, Naturgesch.
Ins. Bienen, Wespen u. Ameisengeschl., p. 217), “malebonze”
(Fermin, 1769, Description de la Colonie de Surinam, II,
p. 300), “marybons” or “marribons” (M. Houttuyn, 1768,
Linnaeus’ Natuurl. Historic, I, 12, p. 259) ; while in British
Guiana the word “marabunta” is in common use (J. Rodway,
1894, In the Guiana Forest, p. 60).
As a matter of fact, the specimens which Ashmead called
“Polistes manillensis'' are nothing but the common and widely
distributed South American Protopolybia sedida (H. de Saus-
sure). A careful comparison with specimens of the latter fails
to disclose any differences of specific value. Moreover, all the
generic peculiarities of Protopolybia are present, including the
characteristic shape of the postscutellum, the hind margin of
which is angularly produced into the propodeum. It seems
strange that Ashmead should have recognized Polistes nmnil-
lensis in these specimens, since they do not in any way agree
with de Saussure’s description. They are, moreover, much too
small (length, h. -)- th. -)- t. 1 -j- 2 : 4.5 mm. ; spread of wings:
9 mm.) ; for de Saussure’s measurement of length of P. manil-
lensis (8 mm.) includes only the head, thorax and first two
abdominal segments, while he gives 19 mm. as the spread of
the wings.
The question now arises as to what should be done with the
generic name Polistella. At present there is no agreement
among taxonomists as to what is the true genotype when a
generic name has been based upon misidentified specimens.
The majority of American entomologists seem to accept the
printed designation of the genotype as binding, regardless of
the characters given in the generic description and of the iden-
tity of the specimens used by the author. I personally favor
this method, which in the case under consideration will place
Polistella Ashmead (genotype by monotypy : Polistes manillensis
H. de Saussure, 1853) in the subfamily Polistinas. Its exact
status there (either as a synonym or a subgenus of Polistes)
will be decided by a further study of the type of P. mmnillensis.
That species was originally described from a male at the British
OCTOBER, 1930]
BEQUAERT POLISTELLA
93
Museum. It does not seem to have been recognized since
de Saussure, although Meade-Waldo mentioned that Polistes
flavobilineata (Cameron), described as Icaria, was very closely
related to P. manillensis.
If one argues, however, that the identity of Ashmead’s
specimens must settle the matter, Polistella Ashmead (1904)
would take precedence over Proto polybia Ducke (1905, Revue
d’Entomologie, XIV, p. 17), in the subfamily Polybiinse. This
would be a most unfortunate change for which I do not feel
there is any justification, considering that Ashmead not only
failed to recognize the true characters and affinities of his speci-
mens, but made an erroneous specific identification, in addition
to a bad guess as to the country of origin.
Ashmead’s claim that his ^‘Polistella manillensis’' is the
“smallest social wasp known in the world,” loses much of its
flavor in view of the fact that several other species of Proto-
polybia are equally small, while some specimens of Leiponieles
lamellaria (Moebius) may be even smaller.
A NEW SUBSPECIES OF PLEBEJUS ICARI-
OIDES BDV.
BY R. F. STERNITZKY
Plebejus icarioides moroensis Sternitzky, n. subsp.
Differs from Plebejus icarioides var. evius, the closest allied
variety by the difference in the ground color of the superior
surface, which is true blue, lacking the violet hue of evius.
The series of this variety compared with a lengthy series of
evius shows a decided difference in size, being much smaller
than the latter.
Male: The blue ground color fuses gradually into the black mar-
ginal border, which becomes darker toward the apex of the prima-
ries. The secondaries have this dark color present along the costal
margin, gradually fading away as it approaches the anal portion of
the wing. Submarginal spots between the median nervules present.
On some specimens there is a slight indication of a black discal cell
spot. Wing fringes white.
Inferior surface bearing a surprising resemblance to Plebejus
pheres Bdv. A row of six spots between the limbal and discal area
prominent, as is the discal cell spot. These are bordered by white
scales. The sixth spot, closest to the inner margin, is a double eye
94
THE PAN-PACIFIC ENTOMOLOGIST [VOL. VII, NO. 2
spot, and on all specimens at hand this peculiarity is constant. A
submarginal row of six other black spots is also present. These are
small, tending to obliteration toward the apex of the wings, and the
sixth inner marginal spot again tending to twin.
On the secondaries the six spots between the discal and limbal
area are well obliterated by white scales showing only indications
of black centers, except the one closest the costal margin which has
a well-defined black center in all specimens before me. TwO' other
spots situated between the basal and discal area are less constant
and appear as white spots in some specimens, while in others the
black centers are present. The white discal cell spot is large and
irregular. A submarginal row of seven white spots with minute
dark centers is also present. Ground color of wings silver gray.
Female: Ground color, superior surface of wings blackish brown,
heavily powdered with blue scales toward the base. Black discal cell
spot present. On the primaries a distinct submarginal band of light
buff, which gradually fuses intO' the ground color, is always present.
The intensity varies in the specimens before me, and the buff color
has a tendency to fade into the dark brown ground color toward
the apex. On the secondaries the band tends to divide into spots
with dark centers, the one between the median veins one and two
being most prominent. Inferior surface as in males, with the excep-
tion that the ground color takes on a creamy appearance and the
black spots of the primaries are somewhat accentuated.
Wing expanse, 27 mm.
Specimens taken by the author at Moro Beach, San Luis
Obispo County, California, June 27, 1929, on the blue flower-
ing beach lupin.
Holotype, male (No. 2982), allotype, female (No. 2983),
and three male and three female paratypes, deposited in the
California Academy of Sciences; two males and two females
in the Southern California Academy of Sciences, and fifteen
male and eleven females, paratypes, in the author’s collection.
OBSERVATIONS ON TWO CALIFORNIA
BEETLES
BY HOWARD HINTON
Velgus Calif ornicus Horn. — While collecting near Riverton,
El Dorado County, California, during August 1930, I was
attracted to three fire-killed sugar pines which were badly
infested with termites. Beneath the loose bark and at the base
OCTOBER, 1930] HINTON NOTES ON TWO BEETLES
95
of the trees in the damp, rotten wood and termite castings all
stages of this insect were to be found. The pupae were lying
in small hollows which looked as if they had been made out
of the castings. Remains of adults found in the neighborhood
seemed to indicate that at least two broods had developed here.
Diphyllostonm nigricollis (Fall). — One day in the latter
part of June while collecting from the power company’s canal
near Riverton, I noticed many of these Luncanidae floating by.
Believing that they were rare, I settled down to collect and took
about fifty. The next day I collected more and thereafter at
weekly intervals collected others, until the latter part of August.
They were most numerous from nine to eleven o’clock and
increased from day to day until toward the last of July, when
they diminished rapidly in numbers. On one day in July, I
caught 112 specimens. I believe that all of these specimens had
fallen into the water and that, because of the rapid current,
none had escaped.
Note. — These observations by Mr. Hinton, a wide-awake young
Junior College student, are most important. Valgus calif orntcus Horn
has been known for a long time to be more or less associated with
termites. The above observation shows that though somewhat a
general scavenger in rotting pine or fir wood, it has a decided pref-
erence for the areas where the termite castes have collected, in this
regard resembling the European rose chafer, Cetonia aurata L. and
the species of our own genus Cremastochilus which pass their early
existence in the nests of ants.
The genus Diphyllostoma Fall is a most peculiar one. It contains
two species, fimbriata Fall and nigricollis Fall, both first captured by
Ralph Hopping; the first near his old home at Kaweah, the second
in Fresno County. These specimens were taken, if I remember cor-
rectly, while flying or resting on the grass and at dusk. During May
and June, 1929, and again in late May, 1930, numbers of the first
species were taken by my students and myself from the flumes of
the power companies situated just below the entrance to the Sequoia
National Park. We had believed that they were dusk flyers. Mr.
Hinton’s observations seem to indicate that they prefer the heat of
the day. All specimens so far taken have been males. Is the female
flightless like the females of Pleocoma, Anorus, Euthysanuis and
Zarhipisl — Edwin C. Van Dyke.
THE PAN-PACIFIC ENTOMOLOGIST
Published quarterly by the Pacific Coast Entomological Society
in co-operation with the California Academy of Sciences
E. P. Van Duzee, Editor S. B. Freeborn, Treasurer
Editorial Comment
On a preceding page of this number of the Pan-Pacific
Entomologist Dr. Bequaert brings up a point in nomenclature
that it seems to me requires further consideration. He says :
“The majority of American entomologists seem to accept the
printed designation of the genotype as binding, regardless of
the characters given in the generic description and of the iden-
tity of the specimens used by the author.” For some time there
has become more or less apparent a tendency toward a blind
worship of the type and a corresponding disregard of the
description, with the claim that this is necessary for stability
in nomenclature. This would be true if type labels were
infallible, but they are far from that, and in the last analysis
the only dependable element in nomenclature is the description.
Recognizing this as a fact we have but one course to follow.
A genotype must not disagree with the description of the genus,
nor should a species type disagree with the species description.
Errors of omission may be allowed to pass, but the type of a
genus or species must possess the characters named in the
description. A case in point is Chermes of Linnaeus. He
wrote “Pedes saltatorii,” and thirteen of the fourteen species
listed by him are saltatorial. Then, on the basis of elimina-
tion, someone named the fourteenth, a plant louse, as the type
of Chermes, and many of our entomologists now have a genus
of saltatorial plant lice that cannot jump. The same incon-
gruity has occurred many times in the case of species. In the
case of Polistella, mentioned by Dr. Bequaert, some clerk
might some day inadvertently shift Ashmead’s labels to another
insect; then who could tell what Ashmead called ‘'Polistes
manillensis’' and Polistella might find itself in the Ichneumo-
nidae. Stability is impossible unless we follow the description.
A MANUAL OF THE GENERA OF BEETLES OF
AMERICA NORTH OF MEXICO
Keys to the genera and higher groups of Coleoptera, with
a taxonomic list of genera. By J. Chester Bradley, Professor
of Entomology in Cornell University. Limited edition. Litho-
printed. xvi, 360 pp. Buckram, $6. Pressboard, $5.
Daw, Illston & Co., 526d West Clinton Street,
Ithaca, New York
Publisher’s charges for reprints from the Pan-Pacific Ento-
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These prices will be charged for author’s extras ordered
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ENTOMOLOGICAL PAPERS RECENTLY
PUBLISHED IN THE PROCEEDINGS OF THE
CALIFORNIA ACADEMY OF SCIENCES
Fall, Expedition of the California Academy of Sciences to the
Gulf of California in 1921. The Chrysomelidae. 15
cents.
Cole, A Study of the Terminal Abdominal Structures of Male
Diptera. 75 cents.
Van Duzee, M. C., A Contribution to our Knowledge of the
North American Conopidae. 1927. 25 cents.
Van Duzee, M. C., New Mycetophylidae Taken in California
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For Proceedings of the California Academy of Sciences address
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Vol. VII January, 1931 No. 3
THE
Pan-Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
ESSIG, CHARLES FAYETTE MCGLASHAN 97
STANFORD, NOTES ON DIPTERA ATTACKING MAN 99
BARRETT, NEW SPECIES OF APHODIUS AND MALACHIUS FROM CALIFORNIA . 101
HATCH, NOTES ON PH^DON 103
HOPPING, NOTES ON POGONOCHERUS 105
LINSLEY, A CORRECTION 106
GUTHRIE, NOTES ON DEVELOPMENT IN LINA 107
VAN DUZEE, A NEW ISCHNORRHYNCHUS 110
BLAISDELL, A NEW SPECIES OF ZOPHERODES FROM CENTRAL CALIFORNIA . Ill
BARE, A BUENOA OF SOUTHWEST UNITED STATES AND MEXICO .... 115
BALL, SOME NEW LEAFHOPPERS OF THE GENUS ALIGIA 119
MOULTON, A NEW ^OLOTHRIPS FROM NEVADA, WITH NOTES ON OTHER
SPECIES 122
WOODBURY, PENTATOMID.® OF ZION NATIONAL PARK 124
BRUCK, TWO NEW SPECIES OF PHLCEOSINUS 126
USINGER, A NEW SPECIES OF PLATYLYGUS 129
KLYVER, CHERMID®: FROM UTAH, NEVADA, AND ARIZONA 131
EDITORIAL COMMENT 144
San Francisco, California
1931
THE PAN-PACIFIC ENTOMOLOGIST
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PUBLICATION COMMITTEE, PAN-PACIFIC
ENTOMOLOGIST
E. O. Essig, Chairman
G. F. Ferris R. W. Doane
E. C. Van Dyke Grant Wallace
REGIONAL members
Dr. Vasco M. Tanner, Provo, Utah
Mr. Jeane D. Gunder, Pasadena, California
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E. P. Van Duzee, Editor
E. C. Van Dyke, Associate Editor
S. B. Freeborn, Treasurer
Published at the California Academy of Sciences, Golden Gate Park,
San Francisco, California.
Entered as second-class matter, February 10, 1925, at the postofflce at
San FYancisco, California, under Act of August 24, 1912.
The Pan-Pacific Entomologist
Vol VII, No. 3 January, 1931
CHARLES FAYETTE McGLASHAN
BY E. O. ESSIG
With the passing of Charles Fayette McGlashan, the West
has lost one of its most interesting men who link the present
with the old and colorful past. Born near Jamesville, Wis-
consin, August 12, 1847, he crossed the plains in an ox wagon
and arrived in California in September, 1854. Later he
attended the Satoyome Institute at Healdsburg, California,
1862-1865, and finished his education at Williston Seminary,
Easthampton, Massachusetts, 1868-1870. He prepared for law
and the many other occupations which he pursued in after
life as a school teacher, editor, historian, and amateur scientist.
As principal of the Placerville high school in 1871-1872, he
took a class four miles to Negro Hill to look through a tele-
scope owned by a Mr. Oldfield,^ an Englishman and a miner
and an ardent entomological collector. This visit was the be-
ginning of two passionate hobbies which at once seized upon
the budding scientist. The first was the study of astronomy,
which he pursued with such thoroughness that it later won him
a place as director of the Chabot Observatory, Oakland. The
other was entomology. It happened that, on the day of his
visit to the abode of Mr. Oldfield, a large number of moths
emerged in a breeding cage and they so fascinated him that,
after receiving all the instructions possible in the art of collect-
ing and rearing moths and butterflies, he became fired with
entomological enthusiasm which never ceased. After he became
principal of the Truckee schools, July 12, 1872, he devoted
much of his time to collecting, rearing, and distributing Lepid-
optera. His work attracted attention of many entomologists.
Henry Edwards visited him and taught him how to sugar for
moths, and during the twelve years from 1875-1887 McGlashan
sent him the most of the moths reared and collected, so that
many specimens in the Edwards collection, now at the Ameri-
1 Oldfield was an amateur entomologist who donated a collection of
Lepidoptera to the California Academy of Sciences, San Francisco. The
specimens were later destroyed by the earthquake and fire of April, 1906.
98
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
can Museum of Natural History, New York, bear the locality
“Truckee.” During the years 1884-1887 he also collected eggs,
larvae, and pupae of butterflies for W. H. Edwards, who gave
him much information as to securing eggs and rearing the
caterpillars of butterflies. In turn Edwards determined all of
the butterflies in his collection. Albert Koebele collected with
McGlashan at Truckee in 1886. During the early eighties
specimens were collected there by B. Neumogen, Gamble
Geddes, Herman Strecker, W. G. Wright, James Behrens,
J. J. Rivers, and others. As a result of his endeavors Mr.
McGlashan amassed a collection of twenty thousand specimens,
especially rich in butterflies. Many specimens were preserved
in patented, wholly glass cases, each of which is described as
follows :
“The case is made wholly of glass, preferably in the form of a
six-sided box. It is made in sections, each side being a separate
piece — that is to say, the top is one piece of glass, its bottom another
piece of glass, and each of its sides one piece of glass. The meeting
edges of each piece are beveled or chamfered to an angle to suit the
particular case, and these joints are thoroughly cemented together,
so that the sections of the case become practically a single piece,
forming a hollow shell or casing which is permanently and hermeti-
cally sealed. The specimens are pinned to a strip of cork cemented
to glass.” 2
Beginning in 1912 and continuing until the World War in
1915, he taught his daughter, Ximena McGlashan,^ how to
collect and rear butterflies and moths, how to sugar for moths,
and the methods of disposing of the adults. A butterfly farm
was established at Truckee, a correspondence course for ama-
teur entomologists given to hundreds of pupils, and a monthly
magazine ^ issued.
His collections are still maintained at his home in Truckee.
For him Rivers named Lemonias macglashani which proves
to be a form of Euphydryas chalcedona (Dbldy. and Hew.),
and Henry Edwards named Hepialus macglashani, a synonym
of H. pulcher Grote.
McGlashan died at his home at Truckee, January 6, 1931,
and was cremated at Sacramento.
2 Pacific Pmral Press, Vol. 35, p. 378 (April 28, 1888).
3 Now Mrs. J. C. Howard.
4 The Butterfly Farmer, Vol. 1, No. 1 (September, 1913) to Vol. 1,
No. 12 (August, 1914), 208 pages.
January, 1931] essig — c. f. mcglashan
99
To his fellow citizens of Truckee he was perhaps best
known as a competent school teacher, a most successful
criminal lawyer, an editor ® of reputation, a searcher of the
stars, a collector and breeder of butterflies and moths, and a
historian of the hardy pioneering days. To the world at large
his name will be perpetuated as the historian of the famous
Donner tragedy which occurred in the high Sierras near his
home at Truckee in the winter of 1846. His book of 261 pages
entitled “History of the Donner Party,” a tragedy of the
Sierras, was first published by Bancroft, San Francisco, in
1880. A reprint by the same company appeared in 1881 and
subsequent reprints by Crocker, Sacramento, 1907, and Car-
lisle, San Francisco, 1918.
While at Santa Barbara he became interested in a system
of railroad telegraph signals and published a small paper of
eight pages entitled : “McGlashan’s train telegraph. A certain
method of preventing railroad collisions. Telegraph offices
upon cars.” No date is given.
NOTES ON DIPTERA ATTACKING MAN IN
SEVIER GOUNTY, UTAH
J. S. STANFORD, PH.D.
Utah State Agricultural College, Logan, Utah
The following species were collected principally on the
writer while in the valley and canyons collecting birds and
mammals in 1929. The hunter became the hunted, so to speak.
Aedes dorsalis (Meigen). — These mosquitoes began to ap-
pear in Salina in May. By July, when the first crop of hay
was being cut, I saw them in great numbers on the work horses
in fields near the river. They attacked man persistently, too.
Upon asking a farmer where the mosquitoes bred, he replied,
“I don’t know where they breed, but I do know where they
feed.” These mosquitoes seemed to attack most viciously at
dusk.
Aedes campestris D. and K. — This species mingles with
dorsalis throughout the state where collections have been made.
Several specimens were taken in Salina.
5 He edited the Truckee Republican and the Santa Barbara Press.
100
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
Aedes vexans (Meigen). — Late in July while climbing
from the experiment station in the mountains east of Salina
(elevation 7800 feet) to the higher ridges, I encountered fre-
quent small swarms of a dark species which followed persist-
ently, settling principally on my back and legs. They were
more numerous in protected places near trees and bushes where
the breezes were less pronounced. Unfortunately I had left
my cyanide bottle in camp, but I easily caught a score or more
in a vial of alcohol by putting the open vial over them. Be-
cause of this wetting, positive identification was very difficult.
Culex tarsalis Coq. — I collected several females (and two
males) in June on the walls and ceiling of a bedroom at night.
They did not attack us before we slept. They doubtless did
so later in the darkness.
Symphoromyia hirta Johnson. — At the experiment station
in late July we were frequently attacked by these flies darting
to our necks, faces and hands with a low hum, and quickly
beginning to drill. This drilling process was rather painful,
but no swelling resulted and the tingling was of short dura-
tion. These flies lost all caution when boring for blood, and
could be easily picked up or crushed. Hence collecting was an
easy matter.
Silvia quadrivitatta Say.
Chrysops fulvaster O. S. — Moderate numbers of these two
species, the latter more common, attacked me promptly when
walking through the grass and brush by the Sevier River.
They were difficult to capture in hand until well engaged in
biting. Only a few specimens were taken, mainly because of
my reluctance to fight the swarms of mosquitoes longer than
necessary.
Culicoides varipennis Coq. — On hot afternoons in the
valley, especially on the leeward side of hills, these small flies
appeared in swarms and crawled under hat band, down the
neck, etc., biting promptly. Killing apparently did no good,
as others instantly took their places.
The writer is indebted to Dr. O. A. Johannsen, Professor
of Entomology at Cornell University, and Mr. D, M. Rees,
Zoology Department of the University of Utah, for the deter-
mination and verification of the species listed.
January, 1931] barrett — new aphodius and malachius 101
NEW SPECIES OF APHODIUS AND MALACHIUS
FROM CALIFORNIA
(Coleoptera)
BY R. E. BARRETT
Saticoy, California
Aphodius vandykei Barrett, n. sp.
Oblong, convex, black, shining, elytra black with intervals 2 to 5
and part of 6 yellow. Head entirely black, moderately convex,
without trace of tubercles, surface rather finely closely punctate;
clypeus feebly emarginate, the angles broadly rounded, sides arcuate,
genae moderately prominent, not as obtuse as in cruentatus\ antenna
brownish, club piceous. Thorax one and one-half times as wide as
long, slightly narrower in front, sides feebly arcuate, hind angles
distinct but obtuse, base arcuate, basal marginal line entire and more
distinct than in cruentatus', disk convex, punctuation fine and dense
slightly denser and finer at the sides. Scutellum small, black.
Elytra at base not quite as wide as thorax, humeri distinct, with
a small acute tubercle; striae moderately deep, rather finely punc-
tate, the striae red where the intervals are yellow, intervals very
slightly convex, punctuation fine and sparse, the first or sutural
interval black, intervals 2 to 5 and part of 6 yellow except the
apical one-eighth which is usually black or dark brown, the inter-
vals from 6 to the side black. Mesosternum not carinate between
the coxae. Metasternum sparsely evenly punctate. Abdomen indis-
tinctly punctate. Anterior tibiae smooth in front, tridentate exter-
nally, finely crenate above the upper external tooth, piceous, first
tarsal segment as long as the second, piceous. Middle tibiae and
tarsi piceous. Posterior femur rather densely punctate, black, tibiae
with unequal spinules, piceous, the first tarsal segment distinctly
longer than the next three, piceous. Length, 4.5 to 6.0 mm.;
breadth, 2.25 to 3.0 mm.
Holotype, No. 2984, in the Museum of the California
Academy of Sciences. Numerous paratypes in my own col-
lection. All were taken by me from nests of the wood rat,
Neotoma, in the East Fork of Santa Paula Canyon (2500 feet
elevation), Ventura County, California, January 26, 1931.^
A triangular black spot on the elytra is noticeable on many
specimens, the base of the triangle being at the base of the
elytra. This black spot is due to the transparency of the elytra
1 A number of specimens of the same species were taken by me and
my students from the flotsam in a reservoir below the Sequoia National
Park during May, 1929. — Edwin C. Van Dyke.
102
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
and the manner in which the wings are folded. Specimens
preserved in alcohol are noticeably darkened. No sexual differ-
ences are noticed in the series at hand.
By Horn’s Monograph this species runs to terminalis and
cruentatus in Group I-d. It may readily be separated from
terminalis by the small acute tubercle on the humeri, the first
tarsal segment of the posterior legs being longer than the next
three, the yellowish markings on the elytra, and by being in
general larger. From cruentatus it may best be separated by
color, smaller size, lack of tubercles on the head, and shape
of the humeri.
This is the fourth Aphodius from Neotonm nests, neotomce
Fall, davisi Fall, and sparsus Lee. having been previously
described from this habitat.
I take pleasure in naming this species after Dr. E. C.
Van Dyke.
Malachius californicus Barrett, n. sp.
Moderately elongate, parallel or slightly wider posteriorly, from
above metallic blue bordering on green. Head with two frontal
and one small vertical impression, finely punctate, shining metallic
bluish green, antenna eleven-segmented, black; male — segments 1
and 2 cylindrical, 3 triangular twice as long as the second, 4 to 10
pectinate, branch on segment 4 one-half as long as branch on seg-
ment 6, branch on segment 5 equal in length to branch on segment
10, branches on segments 6 to 9 equal in length; female — segments
1 and 2 cylindrical, segments 3 to 10 serrate; undersurface of the
distal half of the first segment and the undersurface of the second
segment yellow in both sexes. Prothorax distinctly wider than long,
glabrous, shining, broadly rounded, uniformly colored. Elytra shin-
ing, scabrous, apex not appendiculate in either sex. Undersurface
of body and legs dark green bordering on black, pubescence fine,
yellow. Length, 3.5 to 4.0 mm.; breadth, 1,5 to 2.0 mm.
Holotype, male, No. 2985, in the Museum of the California
Academy of Sciences. Allotype, female, and one paratype in
my collection. Described from three specimens collected by
myself in Marin County, California, April 18, 1926.
In Horn’s Synopsis of the Malachiidae of the United States
this species runs to the group in which the male antenna is
pectinate and the apex of the elytra are not appendiculate. It
may readily be separated from mixtus and ulkei by its uniform
metallic bluish-green color.
January, 1931] hatch — notes on ph^don
103
NOTES ON PH^DON
BY MELVILLE H. HATCH
Dr. H. C Fall (Pan-Pac. Ent., V, 1929, pp. 145-152) has
pronounced invalid the nine new species of Phcsdon that I
described (1. c., 1928, pp. 44-47, 59-62). Of these Mr. Charles
Schaeffer (Bull. Brook. Ent. Soc., XXIV, 1929, pp. 286-287)
has declared P. carri Hatch valid. For the present, P. van-
couverensis Hatch can likewise be considered distinct. It is
separated from oviformis Lee. by characters of no less moment
than those considered valid by Schaeffer; Fall, moreover, was
unable to report intermediates between it and oviformis.
Since Dr. Fall asserts that he has seen intermediates,
P. punctatus and vandykei Hatch can be regarded as varieties
of prasinella Lee., and dietricM Hatch a variety of viridis
Melsh. (csruginosus Suffr. ; microreticulatus Hatch*). These
differ by clear-cut characters, and, until the matter can be
given more detailed study than Dr. Fall has obviously given
it, the progress of our knowledge is better served by keeping
the several varieties separated. As a third member of the
zdridis group, I announce the var. oklahomensis nov. It is
characterized by the almost complete obsolescence from the
interstriae of both rugosity and micropunctulation and by the
very fine punctation of the disk of the pronotum. The type
is from Chickasha, Oklahoma, in my collection.
The case of P. niger Hatch is difficult. It is most closely
related to americanus Schaeffer (armoracice Hatch and Fall,
nec L.) and it must replace that name if it be eventually proved
to be cospecific with it. It is characterized by jet-black elytra
and convex interstriae, and I have two specimens that fulfill
* Dr. Pall intimates that I was not entitled to rely on the published
descriptions of viridis Melsh. and purpurea Linnell ( p u r pu-
re s c e n s Hatch) but that I was under obligation to consult the types.
I can admit no such obligation. Species exist not by virtue of the char-
acteristics inherent in the type, but by virtue of such of those character-
istics as have been published. When specimens exist that differ from
published descriptions of previously described species in reasonably defi-
nite fashion the obligation is to describe them as new. The occasional
synonomy so produced is much preferable to the announcement of really
new species under old names — a practice that, at best, is frequently
unavoidable.
t Dr. Pall’s proposal to ignore the type of this form is reminiscent
of the reputed habit of the older naturalists of throwing out specimens
that didn’t "fit.” How much of the clarity for which Dr. Pall’s mono-
graphs are so justly famous has been obtained at the price of the sup-
pression of a portion of the data?
104
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
both these requirements. Americanus is characterized by more
or less metallic elytra and flat interstriae. A third form, which I
name planus sp. nov. (Type and four paratypes : King County,
Washn., Evans Creek, May 11, 1929. M. H. Hatch, in my
collection), is characterized by jet-black elytra and flat inter-
striae. So far I have not taken americanus, as above defined,
in western Washington, and it is possible that planus replaces
it in this area. All three of these forms possess a nearly
similar elytral microsculpture, are closely related, and may
eventually be shown to be varieties or subspecies of a single
species.
Two corrections in my second paper on Phcedon (1. c., pp.
59-60) should be noted. The division "A^” of my table should
read “elytral intervals not microreticulate.” Cochlearice L. is
erroneously included under the species without a series of
larger punctures on the second interstria, although the larger
punctures are very faintly indicated in my material which,
moreover, exhibits a definite transverse rugosity of the inter-
striae.
Swarming of Two Species of Diptera
On March 22, 1931, while collecting by the Salinas River
at Templeton, Monterey County, California, I noted an unusual
gathering of Odontomyia pilosa Day. They were swarming on
a cluster of stones projecting a little above the surface of the
water, there being perhaps one or two hundred individuals
altogether. Some were resting on the stones while others flew
back and forth close over them with a rather loud humming
noise, much like that produced by certain Bombyllids when
hovering; perhaps a mating flight.
On the bank near where these Stratomyids were gathered
was a patch of Bceria chrysostoma on the flowers of which
were great numbers of the little Bombyllid, Ploas atratula
Loew. This species is always common on these Baeria flowers,
but I have never seen them in anything like such numbers. —
E. P. Van Duzee.
January, 1931] hopping — pogonocherus
105
NOTES ON POGONOCHERUS
BY RALPH HOPPING
Vernon, B. C.
Referring to Mr. Tinsley’s excellent review of the genus
Pogonocherus in the Pan-Pacific Entomologist for Octo-
ber 1930. The restricted distribution of some of the species
seems to call for comment. A few hosts may also be added
to those given by Mr. Tinsley.
Pogonocherus propinquus Fall. — This species has been bred
by us from not only Pinus ponder osa Taws, but from P. con-
torta Toudon and P. monticola D. Don. at Midday Valley, near
Merritt, B. C., and at Trinity Valley, near Vernon, B. C.
Pogonocherus arisonicus Schffr. — This species also occurs
in California, as well as Arizona. One specimen was taken by
the author near Tahoe, California. It is fairly common for
species of Arizona and New Mexico to be taken as far north
as southern British Columbia.
Pogonocherus pictus Fall. — We have bred this species in
numbers from Pinus ponderosa Taws at Midday Valley, B. C.,
and from Larix Occident alis Nuttal at Trinity Valley, B. C.
Pogonocherus mixtus Hald. — I have examined one speci-
men of mixtus from Salmon Arm, B. C. Other specimens in
my collection are from Teduc and Edmonton, Alberta; Minne-
sota, and the Eastern States.
Pogonocherus salicicola Casey. — Typical specimens of this
species have been bred from Pinus ponderosa Taws at Midday
Valley, B. C.
Pogonocherus penicillatus Tec. — This is our commonest
species and is frequently bred by us from Picea engelmanni
Engel., in our cages at Torna, B. C. It is apparently a Rocky
Mountain species breeding in spruce, extending from Alaska
to British Columbia and the Rocky Mountain States to
Colorado.
106
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
Pogonocherus oregonus Lee. — Three specimens have been
taken in British Columbia at Vernon, Creston, and Midday
Valley.
Pogonocherus crinitus Lee. — ^Although we have not taken
this species it has been seen from Vancouver Island, as stated
by Mr. Linsley. The host Quercues garryana Hook, does not
exist on the mainland of British Columbia.
From the above notes it may be seen that seven, or one-half
the known species, are found in British Columbia, and several
new hosts are listed.
A Correction
Since the publication of an article on Pogonocherus and
Ecyrus in the last number of the Pan-Pacific Entomologist,
two errors in synonomy have come to my attention. The first
of these is in regard to the specific standing of Pogonocherus
parvulus Lee. I accepted Horn’s declaration as set forth in his
revision,^ and later restated by other writers, that P. parvulus
was merely a small form of P. mixtus Hald. I now find,
through the kindness of Dr. H. C. Fall, that P. parvulus is not
only a valid species, but is the form that was later described
by Casey as P. salicicola. Dr. Fall writes, Pogonocherus sali-
cicola Casey is the same as P. parvulus Lee. which is itself
a good species and distinct from P. mixtus Hald. Since the
publication of Casey’s salicicola, I have carefully examined the
two type specimens in the Le Conte collection and find them to
agree in all essentials with Casey’s species.”
Ecyrus fasciatus Ham. should be considered a valid species
and not a variety of E. dasycerus Say. Of this species Dr. Fall
writes, “I have an example of E. fasciatus Ham. which has
been compared with the type, and found to agree perfectly. It
is more robust than E. dasycerus, with a denser pubescence of
a different pattern, the elytra with scattered fine erect hairs,
quite different from the oblique short ones of E. dasycerus, and
with rounded elytral apices.” — E. Gorton Linsley.
1 Horn, G. H.. Trans. Am. Ent. Soc., VII, 1878, p. 42
January, 1931] guthrie — notes on lina
107
NOTES ON EGG HATGHING— LARVAL, PUPAL,
AND ADULT DEVELOPMENT IN LINA
SCRIPTA (FABR.)
(Coleoptera: Chrysomelidae)
BY ESTHER GUTHRIE, M.A.
Woodland, California
Early in June 1930, a twig from a poplar tree was brought
into the Stanford entomology laboratory, with a number of
the pupae of the beetle Lina scripta {Melanoma scripta) hang-
ing to it. On June 14, a number of adults, spotted and black,
emerged from these pupae. A pair of spotted beetles was iso-
lated and placed in a glass jar for the purpose of observation.
The pair was supplied continuously with fresh young poplar
leaves. Room temperature, during the course of observations,
varied from 69 to 70 degrees Fahrenheit.
On June 22, eight days after emergence, the pair mated.
On June 26, twelve days after emergence, fifty-nine eggs were
deposited on the underside of the leaf. The eggs were attached
to the surface by an adhesive secretion in an oblique position,
occasionally one lying over the other. They were oval and of
a deep yellow color. This was the first of several batches of
eggs oviposited by the female.
Hatching began July 2, the period of incubation being thus
six days. Just before hatching, the embryo is clearly dis-
cernible through the smooth, tough shell. A double row of
dorso-lateral black spots, each basal to a single black bristle, is
conspicuous. At regular intervals, peristaltic movements of
the embryo occur, which, as they become more vigorous, cause
the egg shell to split along the side, near the anterior end. As
these movements continue, the opening is widened. When the
split is quite long, the posterior tip of the abdomen of the larva
is drawn forward and under, while at the same time the thorax
and first two abdominal segments are pushed against the split
in the shell. Each time this “pushing” occurs, the bristles
stand erect and the skin becomes turgid.
108
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
Conspicuous on the larva are openings of two rows of
dorso-lateral glands, two pairs on the thorax and seven pairs
on the abdomen. When the larva is disturbed the membrane
of the openings of the glands is everted and a secretion with an
odor similar to Prussic acid exudes. As the larva is pushing
itself through the split in the shell, the skin becomes turgid
and this membrane often everts, discharging the milky secre-
tion. As the body relaxes, the membrane is retracted.
The “pushing” continues until the split is widened suffi-
ciently for the extrusion of the larval body. At the tip of the
abdomen, there is a sticky disk by means of which the larva
attaches itself to the inside of the shell, while the body is
erected perpendicularly from this point and moved about in
various directions. When pigmentation becomes evident on
legs, head, and thorax, the larva curves forward and comes
to rest with only an occasional movement, until pigmentation
is complete.
Pigmentation takes place by degrees. The mandibles,
simple eyes, claws, and bases of the large bristles become pig-
mented in the embryo. In the larva, the sequence is as follows :
thorax, segments of the abdomen, and the tubercles at the
openings of the glands.
There are four evident pouches on the thoracic segments
adjoining the tubercles. These are very large and soon become
yellow and turgid with fluid, while those
remain brownish black.
on the abdomen
The first moult occurred July 5, the first in-star thus last-
ing three days. The larvae, preparing to moult, cease to feed
and move in such a way as to indicate that the body is being
pulled away from the larval skin. At the same time the four
prominent yellow pouches become very light in color. The
larval skin splits through the head, thorax, and first abdominal
segments. Immediately after ecdysis the larva is yellow and
remains inactive until pigmentation sets in and becomes
complete.
Three days later, July 8, the second moult occurred. From
this time until prepupation the larvae fed voraciously. Four
January, 1931] guthrie — notes on lina
109
days later, July 12, the larvae ceased feeding, the four yellow
pouches became white, and the prepupae attached themselves
to the surface by means of the sticky disk at the tip of the
abdomen, with the body suspended. The curved body of the
prepupa then straightened out. Soon a split occurred through
the cuticle of the head, thorax, and first abdominal segments.
The pupa worked its way downward and about two-thirds out
of the larval skin, the last three abdominal segments remain-
ing covered by the thoracic region of the exuvia. Here the
pupa remained suspended, adhering to the larval integument,
and attached to it by a pair of processes on the seventh
abdominal segment. These processes fit into pockets on the
first abdominal segment of the exuvia, at the base of the paired
glands.
The larval glands previously mentioned are of the nature
of osmeteria. These have not been studied, but the pouches
into which they opened were found to be situated in the larval
cuticula and are shed with the exuvia. Here they remain
turgid with the fluid with which they are filled throughout
ecdysis. Upon pressure, exvagination of the membrane in the
throat of the tubercular openings takes place and the fluid flows
out through the openings as in the living insect. Upon release
of pressure, the membrane mechanically invaginates.
The pupa, upon emergence, is orange-colored underneath,
with the rest of the body a light yellow. The sequence of pig-
mentation is as follows : spiracles of the abdomen, dorsal area
of abdominal segments, legs, wing cases, edges of the thorax,
and head. At the end of one hour and thirty minutes the pupa
became glossy black.
Seven days later, July 19, the first adults emerged. On
July 20, all had emerged.
The body of the adult, on emergence, is yellow, with the
head, antennae, legs, and center of the thorax black. The elytra
are grayish white with the spots slightly pigmented. The wings
are extended until thoroughly dry, when they are folded under
the elytra. The lateral margins of the thorax are yellowish
white, remaining so for several days, until the adult is finally
matured and ready to mate.
no
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
A NEW ISCHNORRHYNCHUS
(Hemiptera: Lygaeidas)
BY E. P. VAN DUZEE
Ischnorrhynchus obovatus Van Duzee, n. sp.
Distinct from all our other species by its longer head and
ovate elytra. Length, 4 mm.
Male. Head distinctly more produced than in reseda, sides nearly
rectilinear before the eyes; vertex regularly shallowly punctate; buc-
culas very narrow, scarcely elevated anteriorly. Rostrum attaining
apex of fourth ventral segment; segment I surpassing base of head.
Pronotum strongly punctate; broadly depressed just before the
middle, sides sinuate, anterior margin one-half the posterior; callosi-
ties forming a smooth impressed arc either side; humeri prominent;
scutellum coarsely punctate. Elytra broad, subhyaline, obscurely
punctate on the median area, with a row of fine punctures on the
basal half of the radial vein and next the claval suture; clavus with
marginal and one median row of close punctures; costa ovately arcu-
ate and reflexed, apex distinctly bent at radial vein; membrane
nearly hyaline. Antennal segment I not attaining apex of tylus;
II and IV slightly longer than III.
Color rufo-castaneous; tip of tylus, gula, segments I and IV of
antennae, and base and apex of II and III, black, tip of IV paler.
Pronotum pale posteriorly, arcuate lines and hind margin blackish;
scutellum castaneous. Elytra brownish subhyaline; dot at tip and
a geminate dash on disk blackish. Beneath mostly blackish, varied
with castaneous on the propleura; acetabulae and orifices pale; pale
margin of metapleura strongly produced at outer angle. Legs casta-
neous, knees and apex of tarsi black.
Holotype, male. No. 3003, and allotype, female. No. 3004,
Museum California Academy of Sciences, taken by the author,
April 30, 1922, on Cypress Ridge, Woodacre, Marin County,
California, on Sargent’s Cypress. Paratypes 3 males, 17
females, same data, 7 males and 3 females taken May 11, 1919,
and two females taken by Dr. E. C. Van Dyke at same place,
and three examples taken on Monterey Cypress at Carmel,
March 24, 1919, and March 22, 1918, the latter by Mr. Louis
Slevin. The ovate form and dark color recalls Gastrodes.
January, 1931] blaisdell — zopherodes
111
A NEW SPECIES OF ZOPHERODES FROM
CENTRAL CALIFORNIA
(Coleoptera: Tenebrionidas)
BY FRANK E. BLAISDELL, SR.
Stanford Medical School and California Academy of Sciences,
San Francisco, California
The new species of Zopherodes described below is the first
to be found north of San Francisco Bay, west of Mount Diablo
meridian and about 38.5 degrees north latitude. Pm'vicolUs
Casey occurs as far north as Dalton Creek (Henry Dietrich)
and the South Fork of the Kings River, in Fresno County.
My remarks at the present time concern only Californian spe-
cies. The known species of Zopherodes occurring in our fauna
north of Mexico, have a wide range of distribution in the
southwestern United States, inhabiting Southern and Lower
California, Arizona, Texas, New Mexico, Nevada, and Utah.
Zopherodes sanctaehelenae Blaisdell, n. sp.
Form oblong-oval, about three times as long as width of elytra,
size variable and somewhat stout, prothorax and elytra almost equal
in width, both narrowed basally, the bases equal to width of an
elytron. Luster dull, color deep black as usual.
Head in repose deeply retracted into the prothorax, with eyes
hidden from view, surface punctate.
Pronotum slightly wider than long, widest at middle, apex
broadly and moderately deeply emarginate, angles rather prominent
and well rounded; sides more or less angulate at middle; at apex of
the antennal grooves broadly arcuate, most strongly so anteriorly
where the margin is most prominent, very feebly reflexed and nar-
rowly obtuse, posteriorly rather rapidly convergent to base with
margin more rounded and granulate; base transverse, angles obtuse
and not in the least prominent; disk moderately strongly and evenly
convex, finely punctured, punctures small, scarcely submuricate,
separated by a distance equal to two to four times their diameter,
slightly denser in the vicinity of the angles where they become
somewhat granulate. A feeble, narrow and transverse submarginal
groove is present against the basal bead.
Elytra evenly oval, frequently slightly widest behind the middle
to rarely subcylindrical, much more rapidly convergent anteriorly
than posteriorly, humeri obsolete; sides broadly arcuate, apex
obtusely and rather narrowly rounded, not lobed; apical tubercles
somewhat elongate and well developed, parasutural impressions
strong; disk moderately convex, most strongly so and rounded
112
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
laterally, arcuately declivous apically; surface with vermiculate lines
which vary in distinctness, punctures rather small, very little larger
and slightly stronger than those of the pronotum, similar in distribu-
tion, slightly muricate, most so laterally and apically, with few
granules in the humeral area.
Prosternum densely and coarsely punctato-subtuberculate, like-
wise the meso- and metasternum, with their parapleurse smooth and
quite impunctate. Abdomen smooth and very sparsely punctate,
punctures small; fifth ventral segment deeply impressed at base,
impression bilobed apically. Legs moderate in stoutness and length.
Male. Epistomal area plane and deep velvety black, more finely
punctate; abdomen more finely and sparsely punctate.
Female. Epistomal area more or less strongly impressed at
apex and laterally, dull in luster as elsewhere, punctures stronger
and a little larger. Abdominal punctures a little larger and stronger.
Measurements. Length (types), 16-19 mm.; width, 6-7 mm.
Holotype, male (No. 2986), and allotype, female (No.
2987), and eight paratypes in the Museum of the California
Academy of Sciences. Thirty specimens studied, 13 males and
17 females.
Type locality: Mount St. Helena, Napa County, California.
Collected on July 15, 1930, from under bark of a dead oak and
pine, on the eastern and western slopes of the mountain, by
Messrs. A. T. McClay, T. W. Serins and E. G. Linsley, who
also possess paratypes.
The Academy’s series of ten specimens give the following
comparative measurements which show the degree of vari-
ations in size:
Total
Pronotum
Elytra
Specimen
Length
Length
Width
Length
Width
mm.
mm.
mm.
mm.
mm.
1
13
4.5
5
9.5
6
2
15
5
6
10
6
3
15
5
534
10
64
4
16
5
6
10.5
7
5
16
5
6
10
6
6
18
6
7
12
8
7
18
5.5
6.5
11
7.5
8
19
6
7
11
8
9
19
6
7
12
7
10
20
6.5
7
12
8
In the study of the sculptural characters of the species of
Tenebrionidae it is very essential to have the specimens per-
January, 1931] blaisdell — zopherodes
113
fectly clean, in order not to be deceived as regards luster and
punctation. This can quickly be done with a soft camel’s hair
brush and chloroform. In the present study a number of the
specimens were brighter than others, the application of chloro-
form removed an oily exudate and the specimens were really
dull in luster. Unless this is done no reliance can be put on
the statement luster bright or dull; likewise as regards fine
punctation, the exudate obliterates the punctules.
In the classification of LeConte and Horn no mention has
been made regarding the unusual size of the epistoma, which
constitutes fully four-sixths of the exposed front of the head
when retracted into the prothorax as in repose. The frontal
sutures are directed obliquely backward and inward to just
before the upper part of the eyes to arcuately unite in front
of the. narrow interocular surface. As a result the sides of
the front appear as small lateral and irregularly triangular
areas before the eyes. In only one specimen before me are the
eyes exposed ; these are transverse, finely faceted and separated
above by a narrow interocular surface.
In Zopherodes as defined by Casey the antennal cavities
are well rounded at apex, which point corresponds to the lateral
subangulation of the pronotal margin. In all of the species
the labrum is exposed, short, transverse, and densely punctured.
The species known to occur within the State of California
may be recognized by the following synoptic statement of the
essential differential characters :
Elytra subcylindrical, being abruptly narrowed at base and toward
apex, pronotum strongly convex, generally very feebly im-
pressed along the median line for a short distance at center.... 2
Elytra evenly oval in outline; pronotum less convex, never im-
pressed on the median line 3
2. Pronotal punctures strong but not muricate, uneven in dis-
tribution, denser and coarser at the sides; elytra not wider
than the prothorax, unevenly tuberculate, tubercles flat-
tened and shining, sublineate; prosternum more clearly and
discretely, very coarsely, punctate; abdomen coarsely punc-
tate. Arizona (Nogales, VIII-9-06; Yuma, III-17-12; Pres-
cott; Phoenix; Tempe Desert). Lower California. Cali-
fornia (Laguna, San Diego County; Coachella Valley,
Riverside County, V-9-27; Caliente, IX-lS-89) trhtis Lee.
114
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
3. Form stout; pronotum coarsely, submuricately punctate, only
slightly more closely so toward the sides; elytra evidently
wider than the prothorax, finely, strongly and unevenly
tuberculate, tubercles shining; prosternum irregularly, closely
and muricately tuberculate; abdomen coarsely punctate.
California (Julian, San Diego County; Mount Wilson, Los
Angeles County, July, 1905) induratus Casey
— Form more elongate; pronotum very coarsely and strongly
(male) to moderately (female) muricately punctate, punc-
tures slightly denser toward the sides, which are rounded;
elytra rather finely but very irregularly, closely tubercu-
late, sculpture transversely and very unevenly rugulose;
prosternum very unevenly punctato-tuberculate; abdomen
moderately densely punctate, finer toward apex. California
(San Diego County, Van Dyke collection; Hesperia, IX-
24-13) ventricosus Casey
-- Body moderately stout, convex; pronotum coarsely, unevenly
and not very sparsely punctate, punctures muricate or sub-
tuberculate, not differing much at sides; elytra slightly
wider than pronotum, sculpturing rough in irregular trans-
verse plicae, feebler, sparser, and mixed with some small
and irregular tubercles toward side; prosternum rather
closely punctato-tuberculate; abdomen not densely punc-
tured throughout californicus Casey
— Form less stout than in californicus, more inflated posteriorly;
pronotum rather sparsely and somewhat irregularly, moder-
ately, coarsely punctate, punctures feebly muricate, closer,
rather coarser and scarcely muricate laterally; elytra about
as wide as prothorax, surface with moderate and somewhat
sparse punctures, each, attended by a small and feeble,
slightly transverse tubercle, evenly distributed, not evidently
lineate; prosternum irregularly and closely but not very
strongly punctato-tuberculate; abdomen coarsely, moder-
ately closely punctate, gradually less coarsely posteriorly,
California (Fresno County: Dalton Creek, 4800 feet, VII-
30-20; South Fork Kings River, VII-1-10; Herron Hill,
11-28-92) parmcollis Casey
— Form rather stout. Pronotum rather finely, sparsely punc-
tate, slightly closer laterally, surface smooth, punctures not
muricate; elytra finely, sparsely punctate, punctures feebly
muricate, no tubercles, few granules about humeral area;
abdomen very finely and extremely sparsely punctate; pros-
ternum very irregularly, not densely punctate, centrally a
few tubercles. California (Mount St. Helena, Napa County,
July 15, 1930) sanctcehelena, n. sp.
January, 1931]
BARE A NEW BUENOA
115
A BUENOA OF SOUTHWEST UNITED STATES
AND MEXICO
(Hemiptera)
BY CLARENCE 0. BARE
Sanford, Florida
The University of Kansas Science Bulletin, Vol. XVIII,
page 342, and Plate LIV, No. 5, published in April 1928,
figured the genital claspers and spine of the new species,
Buenoa arisonis Bare, with the note that a full description
would follow. Since that date we have secured and studied
many more specimens. These include some from Arizona,
Southern California, Lower California, and the State of
Colima, Mexico. The following notes give the more complete
description.
Buenoa arizonis Bare
Sise: $ $, length 8.5 to 9 mm.; width 2.4 to 2.6 mm.
2 2 , length 8 to 8.9 mm., width 2.6 to 2.9 mm.
Color: Eyes dark; anterior part of scutellum and meta-
thorax showing through elytra very dark or amber-colored;
hind margin of scutellum yellowish; notocephalon, prothorax,
legs, and underside of body mostly yellowish or straw-colored ;
ventral part of abdomen except outer portions of connexiva
and median carina dark; tips of front and intermediate tarsal
claws black; considerable red coloring about wing attachments.
Structural characteristics : Head with eyes slightly narrower
than pronotum at hind margin. Notocephalon slightly sulcate
in some specimens, narrowest ventrad of vertex, widest between
vertex and synthylipsis vx^hich are subequal in width. Tylus
full and knob-like as seen in profile, a slight depression or
division at the point of the knob in the males. Synthylipsis :
vertex : base of tylus : : 9 : 9 : 12 in ^ ; in 2 as 9 : 11 : 12.
Pronotum with slight knob-like prominence at anterior
angle ; of smooth swollen appearance in 2 ; distinctly tricari-
nate in $ ; hind margin very slightly sinuate; one-fifth wider
than long in ^ ; nearly one-half wider than long in 2 .
Scutellum about one-eleventh shorter than pronotum; rela-
tively somewhat shorter in 2 than in the $ ; anterior portion
smooth and shining black.
116
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
Pronotum : scutellum : claval orifice : elytral suture : mem-
brane : : 6.5 : 5 : 1.2 : 6 : 9 in (J , and : : 5 : 5.5 : 1.2 : 7 : 9 in $ .
Seventh abdominal tergite of S with lateral (sinistral)
spine as in Fig. 7, fairly long and slender.
Gonapophyses of $ as in Figs. 5 and 6.
Pleurite of second abdominal segment in $ with glandular
opening darkly chitinized.
Foreleg : Coxa of $ with usually one large stridular seta and
a setal vortex near base; trochanter angulate; femur with a
small setal comb at distal cephalad angle, on lateral face about
eighteen crib-forming setae ; on mesal face in $ , stridular area
and setae as shown in Figs. 1 and 2. Tibial prong in $ with
about fifty-one setal teeth. Tarsus with setal comb near base
of digitate black-tipped claws.
Middle leg : Distal half of coxa coated with many fine setae ;
on mesal face of femur a distinct row of forty-five to fifty
tooth-like setae, besides the many large crib forming setae; at
distal end of tibia two setal combs of about five setae in each;
also a setal comb at distal end of second tarsal segment opposite
the somewhat falciform claws ; inner face of tarsus in both
segments armed with short setal claws arranged mostly in
groups of three.
Hind leg: Caudal margin of femur armed with two rows
of short dentate setce, about seventy in upper or dorsad row and
sixty in lower or ventrad row; setal comb almost circumscrib-
ing distal end of tibia; setal comb about one-third distance
from proximal end of first tarsal segment and between the two
rows of long swimming setae on mesal face.
Relative lengths of parts of legs vary, but are nearly as
follows :
Front Femora Middle Femora Hind Femora
For $ $ 4.9 7 10.8
For $ $ 4.5 8 11
Femur Tibia Tarsus 1 Tarsus 2
Front leg 4.5 6 2.8 1.7
Middle leg 8 6.6 3 2
Hind leg 11 10.5 4 3.5
Type locality: Superstition Mountains, Arizona (Pinal
County). See Tech. Bull. 7, Univ. of Arizona Agr. Exp. Sta.,
pp. 148-9.
January, 1931]
BARE A NEW BUENOA
117
Fig. 1. Foreleg, left: Low power micro., from holotype, slides Nos.
1 and 2 (x 22).
Fig. 2. Stridular area of front femur: High power micro., from
holotype, slide No. 2 (x 94).
Fig. 3. Tibial prong showing stridular detail: High power micro.,
from holotype, slide No. 2 (x 94).
Fig. 4. Third beak segment with prongs bearing stridular areas:
Low power micro., from paratype No. 3, slide No. 1 (x 22),
Fig. 5. Right genital clasper: High power micro., from holotype,
slide No. 5 (x 94).
Fig. 6. Left genital clasper: High power micro., from holotype,
slide No. 5 (x 94).
Fig. 7. Sinistral spine of seventh abdominal tergite: High power
micro., from holotype, slide No. 4 (x 94).
118 THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
Types: Described from 151 specimens, 52 ^ ^ and 99 9 9 .
Holotype $ , allotype 9, and paratypes (3 ^ ^ and 10 9 9 )
Nos. 1-3, 5-13, and 92, from Superstition Mountains (Pinal
County), Arizona, dated November 7, 1922, and collected by
P. A. Glick. The holotype is made up in five slide mounts of
which Nos. 4 and 5 are those from which the original drawings
in the University of Kansas Science Bulletin, Vol. XVIII,
Plate LIV, No. 5, were made. Some of the paratypes are
made up in slide mounts as follows: No. 1 (2 slides) ; No. 2
(2 slides) ; No. 3 (pinned specimen and 1 slide) ; No. 6
(1 slide); No. 14 (2 slides); No. 92 (2 slides). Paratypes,
39 ^ ^ and 79 9 9 , Nos. 4, 14-17, 30-91, 93-143, from Santa
Cruz, Cochise, and Gila counties, Arizona; and Santa Rita
Mountains, in Pima County, Arizona; dates July 25, 29, and
August 4, 5, 6, 24, 1927; collectors, R. H. Beamer, P. A.
Readio, and L. D. Anderson. Paratypes, 3 ^ ^ and 5 9 9,
Nos. 18-25, Boboquivaria Mountains, Pima County, Arizona;
(1898?) ; collector, F. H. Snow. Paratypes, 3 ^ Nos. 144-
146, Warsaw, Arizona, December 4, 1893, U. S. National
Museum, Accession No. 28058. Paratype, 1 ^ , No. 148, Cata-
lina Mountains, Arizona, “18/4,” Sabino Canyon, Hubbard col-
lection. Paratypes, 1 ^ and 3 9 9, Nos. 26-29, Escondido
Bay, Baja California, June 14, 1921; collector, J. C. Cham-
berlin; ex. Coll. Cal. Acad. Sciences. Paratypes, 1 9 , No. 147,
State of Colima, Mexico, L. Conradt Coll. Paratype, 1 ^ ,
No. 149, Lower California, P. R. Uhler collection. (This
specimen also bears the label, ''Anisops platycnemis Fieber, det.
Uhler.”)
Location of types: In University of Kansas, Snow col-
lection: Holotype, allotype, and paratypes. Nos. 18-25, 30-91,
93-143. In author’s collection: paratypes. Nos. 1-17, 28, and
92. In California Academy of Sciences collection: paratypes.
Nos. 26, 27, and 29. In U. S. National Museum: paratypes.
Nos. 144-149.
Comparative notes: Easily distinguished from similar spe-
cies by distinctive coloration of scutellum, the number of setae
on hind femora, and especially by the knob-like projection of
the tylus. Buenoa antigone Kirk, and Buenoa carinata Cham-
pion are similar, but both have fewer setae on the hind femora
and lack the knob on the tylus.
January, 1931]
BALL NEW ALIGIA
119
SOME NEW LEAFHOPPERS OF THE
GENUS ALIGIA
(Rhynchota: Homoptera)
BY E. D. BALL
IJni'vers'liy of ArizonOj Tucson
The members of this genus are nearly all western and
especially southwestern in distribution. They all possess the
two cross nervures, numerous reticulations and a concave
suture between the ocelli apparently bounding the front. Van
Duzee recently described three species from California, and
this paper adds five forms from Arizona. The North Ameri-
can species may be roughly divided into groups as follows :
A — Elytra, and general color, pale or milky with fuscous or dark
brown nervures.
B — ^Vertex angulate disk nearly flat
inscripta Van D., calif ornica Van D., colei Van D., and
descripta Ball
BB — Vertex rounding disk convex
jucunda Uhl., munda Ball, oculea Ball, and dellana Ball
AA — Elytra, and general color, tawny or golden, the nervures
concolorous or darkening posteriorly.
C — ^Ground color of basal half of elytra pale or subhyaline.
Transverse bands pale or wanting
modesta O. and B., occidentalis Van D., and manitou Ball
CC — Ground color of basal half of elytra tawny to dark testa-
ceous nervures obscure. Bands usually definite, ivory
turbinata Ball, chiricana Ball, and santana Ball
Aligia descripta Ball, n. sp.
Resembling inscripta, but longer, darker with a shorter and
transversely furrowed vertex. Length, 5 mm.
Vertex shorter and narrower than in inscripta with an obtusely
rounding apex and a deep transverse furrow, longer and narrower
than in jucunda which it otherwise resembles. Eront broader above
than in jucunda, definitely convex in both diameters instead of being
concave beneath the produced apex of vertex as in inscripta. Elytra
longer and narrower than in inscripta. Venation similar in pattern,
much more heavily inscribed than in jucunda. The inscriptions
along costa numerous and fairly regular.
Color: Vertex creamy, with a pair of round dots at apex, a trans-
verse band widening in the middle and the median line connecting
this band with a basal triangle fuscous, a pair of brown dashes
against the eyes and a spot on the disk on either side brown. Pro-
notum heavily inscribed with fuscous. Scutellum creamy, a pair of
large brown spots at the base, a median line and a pair of lateral
120
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
dark spots on the apical triangle. Elytra ivory, the veins and numer-
ous vermiculate lines fuscous, two pale transverse bands indicated.
Holotype, female ^ and allotype, male, Glenn Oaks, Ari-
zona, July 19, 1929, and five pairs of paratypes from Glenn
Oaks, Granite Dells and Yarnell Heights, Arizona, taken by
the writer from July to October, 1929,
Aligia manitou dellana Ball, n. var.
Slightly larger and much darker than typical manitou with
concolorous males. Resembling jucunda, but much smaller.
Length, 4.-4. 5 mm.
Vertex slightly longer and more definitely angled than in manitou.
This is accentuated by a dark transverse band back of the ocelli and
two definite dark spots on the apex. The elytra are ivory subhyaline
with dark nervures as in jucunda, without any trace of banding. The
males are exactly the same color as the females, while in manitou
they are an almost uniform golden yellow in sharp contrast.
Holotype, female, Glenn Oaks, October 7. Allotype, male.
Granite Dells, October, 6, and ten paratypes taken with the
types and at Yarnell Heights, October 8. All taken, by the
writer in 1929. This may well be a distinct species in the strik-
ingly different males, but if so, then it is probable that a third
species is still being confused.
Aligia turbinata Ball, n. sp.
Similar to manitou in structure, slightly larger with a slop-
ing vertex. Mahogany with a broad white band, a dark head
and apex of elytra. Length, 4-5 mm.
Vertex sloping, one-third longer on middle than against eye. Head
very obtusely conical. Elytra flaring, opaque with the venation ob-
scure to the anteapical cells, few transverse veinlets in the anteapicals.
Color: Vertex black, shading out posteriorly. Ocelli white, pro-
notum, scutellum, and anterior half of elytra rich, shining reddish
brown. A broad, opaque, waxy, white transverse band across the
second cross nervure. This band narrowly margined with smoky
and a heavy smoky band running obliquely from apex of clavus
across the three inner apical cells. The anteapical cells and part of
the two outer apicals subhyaline with reddish or smoky nervures.
Face smoky, the margins black; below pale, the pygofers reddish.
Holotype, female, and allotype, male, Glenn Oaks, Arizona,
July 29, 1929, and one male and eleven female paratypes taken
with the types and at Yarnell Heights by the writer in July
and August.
1 T 3 qoes in the collection of the author.
January, 1931]
BALL NEW ALIGIA
121
Aligia chiricana Ball, n. sp.
Longer and more slender than manitou which it resembles.
Elytra delicate milky subhyaline, female, or golden subhyaline,
male, with concolorous nervures on basal half. Length, 42-5 mm.
Vertex shorter and slightly more conical than in manitou, shorter
and less conical than in turbinata. Elytra longer and less flaring
than in manitou. Claval nervures without cross nervure, central
anteapical cell broader but less reticulate, usually four short nerv-
ures to costa, while in manitou there are three.
Color: Female, vertex creamy with faint traces of the usual
markings. Pronotum irregularly marked with wavy brown lines.
Scutellum washed with dusty orange, omitting three ivory points at
apex. Elytra delicate milky subhyaline with concolorous nervures.
Three pairs of black points on clavus, each anterior to a nervure
and increasing in size posteriorly. A narrow band across the second
cross nervure in which the nervures are smoky or brown, and the
membrane ivory. Male golden with the smoky nervures and mark-
ings of female, face and below pale creamy.
Holotype, female, allotype, male, and six paratypes, Chiri-
cahua Mountains, Arizona, July 6; also two pairs of paratypes,
Huachuca Mountains, Arizona, June 15, all taken in 1930 by
the writer. This is a strikingly distinct species with a delicate
coloring hard to describe.
Aligia santana Ball, n. sp.
Size and form of chiricana nearly, slightly stouter with a
more rounding vertex. Smaller and narrower than modesta
with a broad white band. Length, 4.5-5 mm.
Vertex broad and uniformly rounding, the margins parallel, in
chiricana and modesta the vertex is slightly produced. Elytra, in
shape and venation resembling chiricana, not as flaring or reticulate,
especially along costa, as in modesta. Color, pale smoky brown with
three pale areas as follows: The vertex, the apex of scutellum and
adjacent margins of the claval areas and a broad and definite ivory
band across the second cross nervure. There is also a triangular
subhyaline area extending in from the first and second apical cells
of the elytra. The vertex and face are creamy with the usual pale
markings. Three pairs of dark triangles and usually a pair of white
dots along the suture. The apical veinlets are fuscous and a smoky
cloud occupies the central apical cell.
Holotype, female, Santa Rita Mountains, Arizona, June 22,
1930, allotype, male, the same locality in July and five female
paratypes taken by the writer from these mountains June to Sep-
tember. This species approaches turbinata in the banded effect,
but is easily separated by the triple banding and the short vertex.
122
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
A NEW ^OLOTHRIPS FROM NEVADA WITH
NOTES ON THREE OTHER SPECIES
FOUND IN CALIFORNIA
BY DUDLEY MOULTON
Aeolothrips aureus, Moulton, n. sp.
Female holotype: Color bright golden yellow, including legs, with
head and thorax orange yellow, extreme tip of abdomen brownish
and hind tibiae brownish in the middle. Antennal segments one to
three yellowish, two slightly darker, three brown at extreme tip,
four brownish yellow shading darker in outer third, five to nine
brown. Fore wings with two irregular, light brown cross bands,
otherwise clear yellow. Hind wings without transparent bands.
Crescents of ocelli reddish orange. Prominent spines at tip of
abdomen dark brown.
Measurements: Total body length 1.7 mm.; head, length .16 mm.;
width .20 mm.; prothorax, length .15 mm., width .26 mm. Segments
of antennae: length (width) I, 23 (36); II, 50 (28); III, 100 (26);
IV, 70 (23); V, 66 (23); VI, 20 (20); VII-IX, 48 microns; total
length, 382 microns.
Head and thorax without conspicuous spines or markings. Ocelli
well developed. Antennal segments seven to nine subequal and
together with six about as long as segment four. Anterior longi-
tudinal vein of fore wing with thirty-two and hind vein with twenty-
five regularly placed extremely small transparent setae. The only
prominent body spines are found at tip of abdomen.
Male allotype: Colored as in the female but with fifth antennal
segment lighter in basal half, also with a dark brown spot in the
middle of eighth abdominal segment and the median portions of
ninth and tenth brownish. The clasping organs on segment nine
dark brown. Prominent terminal spines brownish yellow.
Measurements: Total body length 1.3 mm. Segments of antennae:
length, I, 16; II, 46; III, 86; IV, 56; V, 56; VI, 20; VII-IX, 48
microns; total length, 338 microns.
This species belongs in the fasciatus group of the genus
which has two distinct cross bands on the fore wings and is
most closely related to gloriosus Bagnall, but is readily distin-
guished by its larger size and different coloration. The ninth
and tenth abdominal segments and eighth tergite are entirely
black in gloriosus and the distal portions of antennae are black.
January, 1931] moulton — ^olothrips
123
Type material: Female holotype; male allotype; twenty-
nine female and eight male paratypes taken in the flowers of
a composite, April 8, 1930, by Professor E, O. Essig (M. No.
4178). Types in author’s collection and at the University of
California, Berkeley, California.
Type locality: Las Vegas, Nevada.
Cryptothrips rectangularis Hood
Mr. A. T. McClay of the University of California has
reported finding many black thrips with their red-colored larvee
in the burrows of a Scolytid beetle, Micraces hirtellus, Lee.,
the lead cable borer, in a dead Linden tree on the Berkeley
campus. Collections were made when they were first observed
in November, 1929. The tree was subsequently cut up and
held for observation of the beetles. Mr. Steinweden with Mr.
McClay examined the burrows again in May, 1930, and found
that all of the thrips were gone, giving evidence that they had
only been hibernating in the burrows. The November collec-
tions included two species, Cryptothrips rectangularis Hood
and Doccessissophothrips animus Moulton. C. rectangularis
Hood, was first found under the dead bark of peach trees at
Urbana, Illinois, and later one specimen was taken by Dr. H. E.
Burke under bark at West Yellowstone, Colorado. Mr.
McClay’s observations now extend the habitat of this insect to
California (Moulton, No. 3882).
Docessissophothrips animus Moulton
Four specimens of this insect were collected in the same
burrows with C. rectangularis Hood. Three other specimens
have been known up to the present time, one taken by sweeping
nettles at Mountain View, California, and two from Corvallis,
Oregon (Moulton, No. 3882).
PcECILOTHRIPS ORNATUS Hood
This species was originally taken from under bark of a
red-oak tree in Washington, D. C., and later on hickory trees
at Harrisburg, Penn., and Macedon, New York. Professor
E. O. Essig recently found both adults and larvae of this insect
under bark of a live oak tree on the Berkeley campus, which
extends its habitat to California (Moulton, No. 4236).
124
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
A LIST OF THE PENTATOMID^ OF ZION
NATIONAL PARK, UTAH
BY LOWELL A. WOODBURY
University of Utah
I was enabled to make this study through the courtesy of
Mr. A. M. Woodbury, the Park naturalist, who has very gener-
ously made the Park collection of Hemiptera available to me
for study. Dr. V. M. Tanner of the Brigham Young Uni-
versity has also very kindly loaned me several rare specimens
from this locality.
To Dr. R. V. Chamberlin I am indebted for aid and for
many valuable suggestions in preparing this list.
Mr. W. L. McAtee of the United States Biological Survey
and Mr. Robert L. Usinger of Oakland, California, have very
kindly checked or identified most of the material herein listed.
Mr. E. P. Van Duzee very generously checked the determi-
nation of Brochymena hoppingi Van Duzee.
Unless otherwise noted, Mr. A. M, Woodbury is the col-
lector of all specimens recorded.
"''^Brochymena hoppingi \^an Duzee. One female specimen
collected May 17, 1922, by V. M. Tanner.
^Brochymena sulcata Van Duzee. Quite common. Col-
lected at various times of the year. Hibernates under the dead
bark of cottonwood trees in this locality.
Peribalus ahhreviatus Uhler. Very rare.
Peribalus limbolarius Stab Quite common. In the speci-
mens before me the color varies from a dark form tinged with
black through one somewhat rufous to a very pale form.
Rhytidoloma faceta (Say). This has never been collected
in Zion National Park, but occurs in the same general region
at St. George, Utah, and should be found there.
Chlorochroa sayi Stab This pretty little bug, which is quite
common in this locality, varies a great deal in color pattern,
from a very profuse to an almost obsolete white spotting on the
January, 1931] woodbury — pentatomid^
125
hemelytra. The membrane at times assumes a purple tinge.
I also have a specimen of this from Bryce Canyon, National
Park, Utah, collected by V. M. Tanner.
Carpocoris remotus Horv. One specimen from this locality.
Quite rare.
*Euchistus inflatus Van Duzee. Fairly common.
^Menecles incertus (Say). Very rare. I have one speci-
men from this locality and one from St. George, Utah.
"^'Thyanta perditor (Fabr.). Quite common. The pronotal
band is lacking in all specimens from this locality and the
color of the sides of the prothorax, the humeral spines, and
the tip of the scutellum vary in color from green to deep red.
The general color of the individuals as a whole varies from a
light green through a blue green to a deep green. One speci-
men is a straw yellow.
Thyanta custator (Fabr.). Very common. I have swept
this bug quite commonly from Nightshade {Solanum nigrum)
during July and August. The color pattern varies much the
same as in perditor, but the pronotal band varies from obsolete
to very prominent.
Aero sternum hilaris (Say). I swept one specimen of this
species from the Large-toothed Maple (Acer grandidentatum)
at the Narrows of this Park on April 24, 1930.
Banasa dimidiata (Say). Fairly common.
^Banasa euchlora Stal. Rare. I have also collected this
species at Pipe Springs, Arizona, about ten miles from the
Utah-Arizona boundary, but in the same general region.
Dendrocoris contaminatus (Uhler). One specimen col-
lected May 17, 1922, by V. M. Tanner.
Perillus bioculatus (Fabr.). Occurs at St. George, Utah.
Not collected in Zion, but may be there.
^Podisus maculiventris (Say). The specimens of this spe-
cies from this locality range from a light brown to a very dark
color. In some of the specimens the ventral line of spots on
the abdomen and the spots on the femur are partially or entirely
obsolete.
Zicrona ccerulea Linn. Very rare.
• Species not listed from Utah in Van Duzee’s catalogne.
126
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
TWO NEW SPECIES OF BARK BEETLES OF THE
GENUS PHLCEOSINUS CHAPUIS
(Coleoptera, Scolytidas)
BY C. R. BRUCK
Berkeley, California
While critically studying the bark beetles of western North
America, I have noted that two of the species in the genus
Phloeosinus were new. As the members of this genus are of
considerable economic importance, I feel that these should be
made known as soon as possible.
PhlcEosinus variolatus Bruck, n. sp.
Large, 2.5-4 mm. long, robust, dark reddish brown to black
species. Head with front granulate-punctate, a deep median con-
cavity surrounded by pubescence in the male, the depression shallow
in the female and with sparse pubescence, the carina short, extend-
ing longitudinally beyond the median area of the concavity, with a
short tubercular elevation at the base of the carina; antennal club
twice as long as wide, the sutures moderately oblique; eyes strongly
margined. Prothorax wider than long, densely, moderately coarsely
punctate and somewhat pubescent, sides arcuate, very slightly sinu-
ate anteriorly, base broad. Elytra longer than wide, striae wider
than intervals and grossly, variolately punctured, the intervals shin-
ing, slightly rugose and with a double row of small alternating
punctures, with a hair arising from each puncture; first, third, fifth,
seventh and ninth intervals uniseriately serrate, serrations of first
interval just attaining the crest of the declivity in the male and the
apex of the elytra in the female, face of the declivity in the former
flat with sparse scale-like pubescence, first interval greatly widened,
shining, transversely rugose, second much narrowed, with scale-like
pubescence, third densely serrate with large serrations, fourth much
narrowed with scale-like pubescence and met by the sparsely, moder-
ately serrated fifth, seventh and ninth intervals and the shining and
narrowed sixth and eighth intervals. The striae of the declivity are
much narrower than on the disk and the punctures greatly reduced
in size. Beneath rather densely clothed with long pile.
The female, as indicate above, has the front less pubescent and
less concave than the male and the serrations of the first elytral
interval attain the apex. In addition the first, third, fifth, seventh
and ninth intervals are uniseriately serrate, the first and third with
the serrations numerous and of moderate size and globular, the fifth,
seventh and ninth more sparsely serrate, with the serrations smaller.
January, 1931] bruck — new phlceosinus
127
the even intervals have a few minute tubercles as far as the declivity,
the fifth, sixth, seventh and ninth intervals meet the fourth, and the
flatter portions of the declivity are densely clothed with a scale-like
pubescence.
This large species is one of the most distinct because of
the gross strial punctuation. It would follow cupressi Hopk.
in Swaine’s ^ key.
Holotype No. 2993, allotype No. 2994, Mus. Calif. Acad.
Sci., and a number of designated paratypes, collected on Mount
St. Helena, California, February 29, 1931, by E. F. Wohletz
and myself, from beneath the bark of dead Sargent Cypress,
Cupressus sargenti Jepson. A pair of paratypes will be sent
to Dr. Swaine for the Canadian National Museum, one pair
placed in the collection of Dr. Van Dyke, deposited in the
California Academy of Sciences, one pair in the collection of
Mr. Wohletz and the remainder in my own collection. A very
long series of specimens has been studied, three in the col-
lection of the California Academy of Sciences, taken by Mr.
E. P. Van Duzee, October 31, 1923, others by Mr. Wohletz
and myself from Mount St. Helena, and a long series taken on
Mount St. Helena and on Cedar Ridge, Alameda County,
California.
Phlceosinus aciculatus Bruck, n. sp.
Of moderate size, 2.0-3 mm. in length, slender, black with
elytra generally reddish brown.
Head, male: Front densely punctate-granulate and shallowly
impressed or almost flat; female with front convex, sparsely pubes-
cent and with a few long hairs at the epistomal margin, the frontal
Carina in male barely visible at the middle of the impressed area,
obliterated in the female; eyes strongly margined; antennal club
twice as long as wide, the sutures slightly oblique. Prothorax some-
what wider than long, base broad and bisinuate, sides arcuate, sinu-
ate anteriorly, disk densely, moderately coarsely punctate and some-
what pubescent. Elytra longer than wide, striae deep and well
defined laterally, less defined on the disk; the intervals, on the con-
trary, wide and distinctly aciculate on the disk and less aciculate on
the sides; striae sparsely, finely punctate, the punctures deeper on
the sides than on the disk, first interval but slightly wider than the
1 Canadian Bark Beetles, Pt. II, by J. M. Swaine, Bull. 14, Ent. Br.,
Dept. Agr., Ottawa, Canada, p. 69, 1918.
128
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
discal striae, the others quite wide; pubescence moderate and short;
declivity on the male smooth and shining; interspaces uniformly
wide, first, third, fifth, seventh and ninth uniseriately serrate; the
serrations of the first are few, distant and acute, of the third, smaller,
more numerous, closer and acute, of the fifth, seventh and ninth,
still more reduced in size and number and less acute; declivity in
the female with scale-like pubescence, first interval with a row of
moderately large, widely spaced and acute serrations, also a second-
ary series of very small serrations, more or less limited to the crest
of the declivity, the second interval with fewer, smaller and more
widely spaced acute serrations arranged uniseriately, the third with
a greater number of closely spaced acute serrations, smaller than
those of the first but equal to those of the second interval and uni-
seriately arranged; fifth, sixth, seventh, eighth and ninth with a few,
small, widely spaced and acute serrations arranged uniseriately,
those of the fourth with the aciculations somewhat more prominent
at the crest. Legs and underside of body densely clothed with short
pubescence.
The female, as indicated above, has the front convex, the declivity
clothed with scale-like pubescence, the serrations decidedly smaller,
more numerous; each interspace provided with some serrations, none
being smooth, the first having a few smaller serrations at the side
of the larger ones at the crest of the declivity and, as in the male,
the fifth, sixth, seventh, eighth and ninth meet the fourth.
This moderately sized species is most distinct because of
the densely aciculated disk and the general aciculations of the
sides, the smooth declivity of the male, and the pubescence of
the legs and ventral surface. It would follow juniperi Sw. in
Swaine’s ^ key.
Holotype, No. 2995, and allotype. No. 2996, Mus. Calif.
Acad. Sci., and a number of designated paratypes, collected in
the Chiricahua Mountains, Cochise County, Arizona, August 8,
1916, by V. W. Owen, probably from juniper. A pair of para-
types will be sent to Dr. Swaine for the Canadian National
museum, a pair in my collection and the remainder in the col-
lection of Dr. Van Dyke, deposited in the California Academy
of Sciences. A series of twenty-four specimens have been
studied, all from the collection of Dr. Van Dyke, taken by
V. W, Owen.
2 Canadian Bark Beetles, Pt. II, by J. M. Swaine, Bull. 14, Ent. Br.,
Dept. Agr., Ottawa, Canada, p. 69, 1918.
January, 1931] usinger — new platylygus
129
A NEW SPECIES OF PLATYLYGUS
(Miridas, Hemiptera)
BY R. L. USINGER
Oakland, California
The species herein described was taken by the author while
collecting in the National Parks of the western United States
during the summer of 1930. The utmost cooperation was
received in all of the parks and a keen interest was shown in
the subject of entomology. For the holotype from the col-
lection of the California Academy of Sciences I am indebted
to Mr. E. P. Van Duzee after whom the species is named in
appreciation of the patience and help he has shown me in the
study of Hemiptera. The reader is referred to a paper by
H. H. Knight ^ for a discussion of the species of this genus and
illustrations of the genitalia.
Platylygus vanduzeei Usinger, n. sp.
Differs from the other species of this genus by its lighter
color ; flavo-testaceous, marked with rufous on the head, cal-
losities, and cuneus, and fuscous on the pronotum and elytra.
Scutellum yellow, the base and apex yellowish white. Male
genital clasper and chitinous tip of the oedeagus distinctive of
the species.
Male. Length, 7 mm. Head more nearly verti-
cal than in P. grandis Kngt. ; clypeus 'with tviro
longitudinal fuscous vittae; front strongly convex
as contrasted with grandis where it is somewhat
flattened; anterior portion very broad at base
with the cheeks rectilinear or nearly so, the
cheeks distinctly arcuate in grandis forming an
anterior lobe. Oblique striations tinged with
rufous; longitudinal sulcus obscure or wanting, the area flattened
with the oblique striations arising from it. Pronotum with the
anterior median triangular spot extending between the callosities,
and lateral angles fuscous; a posterior median area extending from
near the lateral angles to the posterior joining of the callosities
fulvous-brown, with a narrow longitudinal median pale line. The
rest of the pronotum, including the callosities, yellow; the callosi-
ties large, contiguous, attaining the anterior margin, polished, marked
with irregular rufous lines. Lateral margins evidently sinuate; recti-
linear or slightly arcuate in grandis. Posterior lobe of the scutellum
with the base and apex yellowish white; median portion flecked with
A, left clasper, dorsal
aspect; B, chitinous tip
of cedeagus.
1 Bulletin of the Brooklyn Entomological Society, Vol. XIII, p. 16, 1918.
130
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
rufous. Hemelytra flavo-testaceous ; clavus infuscated near the apex
of the scutellum; corium with a broad maculate area at middle,
extending toward the apex, fuscous; embolium translucent and
immaculate; cuneus in great part sanguineous. Second segment of
the antennae becoming fulvous apically, third fulvous to fuscous,
fourth entirely fuscous. Rostrum almost attaining the hind margin
of the intermediate coxae; surpassing the middle of the posterior
coxae in grandis. Venter yellow with the sides of the ventral seg-
ments tinged with sanguineous; the genital segment with a reddish
brown, basal triangular mark either side of the middle. Femora
flecked with rufous.
Female uniformly lighter, the black markings of the pronotum
less conspicuous and the oblique markings of the corium confined to
a smaller area. Rostrum almost attaining the hind margin of the
posterior coxae; distinctly surpassing the posterior coxae in grandis.
Holotype^ male. No. 2997, Mus. Calif. Acad. Sciences, col-
lected by C. D. Duncan, June 17, 1921, at the Grand Canyon,
Arizona, and allotype, female. No. 2998, Mus. Calif. Acad.
Sciences, collected by the author in Grand Canyon National
Park, Arizona, South Rim, June 29, 1930. Paratypes, twelve
females taken at Mesa Verde National Park, Colorado, July '6,
1930, by the author.
Butterflies of Los Angeles County
It is a pleasure to record the appearance of Mr. Gunder’s
list of the “Butterflies of Los Angeles County, California.^
This list enumerates 178 forms, representing 123 recognized
species, the remaining 55 being subspecies (here termed races),
transitional, seasonal or sexual forms or hybrids. Mr. Gunder
has worked out the complicated relationship of these various
categories as well as can be done with our present knowledge
of that fauna. A perusal of this list would be of interest to
anyone studying intraspecific variation. In this paper Mr.
Gunder does not recognize aberrations. Probably he has placed
the significant among these in other categories to which he has
assigned them. This list, with that by Mr. W. S. Wright ^ on
the “Butterflies of San Diego County,” gives a very complete
insight into the butterfly fauna of coastal southern California. —
E. P. Van Duzee.
1 Bulletin of the Southern California Academy of Sciences, XXIX,
pp. 39-95, August 1930.
2 Trans. San Diego Soc. of Natural Hist., VI, pp. 1-40.
January, 1931]
KLYVER CHERMID^
131
CHERMID^ FROM UTAH, NEVADA, AND ARIZONA,
INCLUDING THREE NEW SPECIES
(Homoptera)
BY F. D. KLYVER
San Mateo Junior College, San Mateo, California
The specimens on which the following article is based were
collected by Mr. E, W. Davis, United States Bureau of Ento-
mology, Richfield, Utah, and were received through the cour-
tesy of Mr. P. N. Annand, Sugar Beet Insect Investigations,
United States Bureau of Entomology.
Incidental to other work being carried on by Mr. Davis
fifty-five separate collections were made by sweeping twenty-
three species of host plants^ in southern Nevada, southwestern
Utah, and northwestern Arizona between April 20 and June 21,
1930,. The entire collection includes approximately twelve hun-
dred specimens representing four of the six subfamilies recog-
nized by Crawford: namely, the Liviinas, Pauropsyllinae, Trio-
zinae, and Psyllinae. Three of the fourteen species represented
are apparently undescribed and three of them are here undeter-
minable because of insufficient and fragmentary material.
One or more slide mounts of cleared and stained specimens
have been prepared for each species represented. In several
instances where the material was limited my preference has
been to mount the available material on slides for detailed study
with the compound microscope rather than to rely on un-
mounted specimens in making determinations. For most of the
species specimens for making other mounts are available.
The writer has undertaken an extensive study of the Cher-
midai and is interested in receiving specimens from any part
of the worM for identification. With few exceptions the imma-
ture stages of the Chermidae are little known. Material includ-
ing nymphs and a sufficient number of adults to establish the
identity of the immature stages will be especially appreciated.
Types of the new species are in the author’s collection.
s
1 A statement of the writer’s viewpoint relative to the host plants of
the chermids and an evaluation of the several methods of collecting’
commonly employed will be found elsewhere (1930, Klyver, F. D., Notes
on the Chermidss. Part I. Canadian Entomologist, 62:167-168).
132
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
Subfamily Liviin^
Genus Aphalara Forster
Crawford’s opinion that this genus contains a large number
of closely related and probably intergrading species is sub-
stantiated by my observations on numerous specimens included
in the collection. My observations, furthermore, confirm his
statement that the size and color of the body, the presence,
absence or degree of maculation or coloration of the fore wing,
and the size of the genital segments are subject to considerable
variation even within single species. On the other hand, my
specimens definitely indicate that there are other characters,
usually disclosed only in properly cleared, stained, and 'mounted
specimens studied with the compound microscope, which are
more significant and of greater utility in separating the species
than are the admittedly variable characters mentioned above.
These other characters are: (1) the morphology of the fore-
wing and hind wind, that is, the character of the wing mem-
brane, the presence or absence and distribution of setae, punc-
tations and alar radulae,^ and the character of the venation of
the hind wing; (2) the shape or form and detailed structure
of both the male and female genital segments; (3) the com-
parative size of the anal valve or proctiger and the forceps
or claspers of the male; (4) the shape or form of both the
proctiger and the claspers of the male; and (5) the presence
or absence of, the exact form of, and the location or distribu-
tion of the various types of setae, spines, hooks, and chitinized
ridges or folds that characterize the male and female genitalia
in this group. In addition to these characters the ovipositor
of the female is itself frequently distinctive.
Relying chiefly on these latter characters, the numerous
specimens at hand, obviously agreeing very well with the char-
acters of the genus Aphalara, have been assigned to eight
species. One of these, Aphalara pulchella Crawford, is appar-
ently subject to few variations and is easily recognized. An-
2 Ferris, G. P., and Klyver, P. D. Report Upon a Collection of Cher-
midse (Homoptera) from New Zealand. New Zealand Journal of Tech-
nology. In press.
January, 1931]
KLYVER CHERMID^
133
other species, A. suaedcB Crawford, is, on the other hand,
apparently subject to wide and significant variations. It is of
special interest here because of the bearing my observations
have on the group of species centering around A. artemisicB
Forster, Two apparently undescribed species of this genus
which are represented by sufficient material are here described
as new.
Aphalara suaed^ Crawford
Fig. 1, A, B, C, D, H
1914, Aphalara suaedce Crawford. U. S. Nat. Mus., Bull. 85:31; fig.
Specimens. Over 350 adult males and females and three last-stage
nymphs from Dondia nigra (Raf.), Overton, Nevada, May 2, 1930
(311^); nine males and females from the same host. Las Vegas,
Nevada, May 2, 1930 (316); three males from the same host. Las
Vegas, Nevada, May 3, 1930 (322); many males and females from
the same host, Glendale, Nevada, May 2, 1930 (303); three females
from Covillea tridentata (D. C.), St. Thomas, Utah, May 2, 1930
(309); one male from Atriplex hastata L., Logandale, Nevada, May 2,
1930 (304).
Length to tip of folded wing, 1. 6-2.2 mm.; length of body
mounted on slide, 2.5 mm.; length of fore wing, 1.4-1. 8 mm.; width
of fore wing, .6-. 8 mm.; width of head, .5-. 75 mm.
Comparisons of specimens of this long series with many
specimens taken by the writer from Suaeda moquini Greene,
east of Altamont Pass, Alameda County, California, and from
the same host at several localities in the San Joaquin Valley,
California, point to their being identical. The fact that the
three nymphs listed above are also apparently identical with
numerous fifth-stage nymphs taken in the Altamont Pass col-
lections, together with the rest of the life history of the species,
is further and conclusive proof of the identity of the specimens
in the Davis collection.
It is exceedingly interesting, furthermore, that the numer-
ous specimens before me apparently include all of the variations
on which have been based the descriptions of several sup-
posedly distinct species. Certainly the color variations given by
Crawford for Aphalara artemisice Forster, A. angustipennis
Crawford, A. viridis Crawford, A. pinicola Crawford, and
3 Unless otherwise specified the numbers in parentheses are the col-
lection numbers of E. W. Davis.
134
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
A. suaedcB Crawford are found in these specimens. Further-
more, with the exception of those for A. angustipennis, the
measurements derived from these specimens might apply to any
of the species here listed. Except for some variations in size,
and this means gross size, the genitalia are said by Crawford
to be similar for all of these species. This is a further indi-
cation of the possibility that the species are identical, since the
genitalia, most frequently those of the male, are the most reli-
able single character separating the closely related species of
this genus. From careful study of my specimens, both mounted
and unmounted, and from Crawford’s descriptions, I regard
it as entirely probable that A. viridis Crawford, A. pinicola
Crawford, and A. suaedcE Crawford are the same species.
A. angustipennis Crawford may be a distinct but very closely
related species, and A. artemisice Forster is probably a different
but similarly a closely related species.
Aphalara pulchella Crawford
1911, Afhalara pulchella Crawford. Pomona Journ. Ent., 3:480.
Specimens. One female fromDipetalia linifolia (Vahl), Las Vegas,
Nevada, May 3, 1930 (323); and two males and one female from
Salsola pestifer A. Nels., St. George, Utah, May 17, 1930 (360).
These specimens are in general agreement with the descrip-
tion of the species except for the dimensions of the female
genitalia. The measurements- of the two females at hand, both
of which are mounted on slides, are .3 mm. and .35 mm. for
the genital segments, .5 mm. and .6 mm. for the abdomens.
In other words, the genital segment is scarcely one-half as long
as the rest of the abdomen when the measurements are taken
from specimens with the abdomen expanded instead of from
specimens that have dried and hence shrunken.
Aphalara gutierreziae Klyver, n. sp.
Fig. 1, P
Specimens. Forty-four males and females from Gutierrezia saro-
thrce (Pursh), Las Vegas, Nevada, April 20, 1930 (228); thirty-seven
males and females from the same host. Las Vegas, Nevada, May 3,
1930 (328); six females from the same host, Glendale, Nevada,
May 18, 1930 (408); and, one male and one female from Gutierrezia
lucida Greene, St. George, Utah, May 1, 1930 (292).
January, 1931]
KLYVER CHERMID^
135
Types. Holotype, male, FK 224.1.1 $, from Gutierrezia
sarothra (Pursh), Las Vegas, Nevada, May 3, 1930 (328) ;
allotype, female, FK 224.2.1 $ .
Adult. Length to tip of folded wing, 2. 2-2. 5 mm.; length of body
mounted on slide, 1.5-1. 7 mm.; fore wing, 2. 0-2. 2 mm,; width of fore
wing, .75-. 8 mm,; width of head, .5-. 6 mm. General color of all
males at hand (twenty-one males) very constant; head and thorax
black including eyes and antennge, legs and abdomen green, fore
wings very light brown with the veins darker brown, especially
toward the apex. General color of the females slightly more vari-
able; general green color with yellowish or light brown markings to
a general light brown color throughout, wings with membrane light
brown to darker brown with faint brown spots over the apical two-
thirds, veins distinctly brown; tip of abdomen dark brown. Charac-
ters of the genus well developed.
Head slightly wider than prothorax, scarcely deflexed. Antennse
ten-segmented; one-fourth longer than width of head.
Thorax distinctly arched; the dorsum scarcely or very slightly
pubescent, with very fine hair-like setae. Posterior tibia with eight
black teeth at the apex; two rather small claws on the basal tarsus
of the hind legs. Anterior wings hyaline to very slightly opaque,
somewhat fumate in the apical portion and faintly spotted apically,
the veins usually distinctly brown; shape and venation very similar
to that of A. •veaziei Patch. Posterior wing very delicate in texture,
beset throughout with very small punctations, the venation scarcely
discernible, developed as irregular rows of minute punctations.
Abdomen with the plates moderately chitinized. Genitalia of the
male (Fig. 1, P) relatively large; the “body”^ of the proctiger being
about equal to its vertical length (/>), the length of the lobes (/)
slightly greater than p, the lobes wide at the base and subacute at
the apex, being distinctly expanded dorsad in lateral aspect. Female
genitalia very similar to those of A. suaeda Crawford in every
respect except for their slightly smaller size.
Nymph. Unknown.
Aphalara minutistylus Klyver, n. sp.
Fig. 1, G, L, M, N, O, I
Specimens. More than two hundred males and females from
Artemisia filifolia Torr., St. George, Utah, April 21, 1930 (264).
4 The comparative measurements here indicated are taken as shown
in Fig. 1, H and I; “p” is the greatest vertical length of the proctiger;
“b” is the length of the “body” of the proctiger; and “1” is the length of
the lobe of the proctiger measured from the posterior extremity of “b.”
The value of these comparative dimensions is apparent from a con-
sideration of their application to the two proctigers illustrated.
136
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
Types. Holotype, female, FK 227.1.1 $ , from Artemisia
filifolia Torr., St. George, Utah, April 21, 1930 (264) ; allo-
type, male, FK 227.1.1 $ .
Adult. Length to tip of folded wing: male 2.0 mm., female 2.5
mm.; length of body on slide: male 2.5 mm., female 2.6 mm.; length
of fore wing: male 1.5 mm., female 2.0 mm.; width of head: male
.5 mm., female .6 mm. General color of both males and females
green to pale yellowish brown throughout, with white to pale brown
markings on the thorax, apex of female genital segment very dark
brown to black. Antennae light colored with the tip slightly dark-
ened. Fore wings slightly opaque with numerous sharply delimited
dark brown spots, sometimes closely crowded on the apical third of
the wing. Characters of the genus well developed.
Head as wide as thorax; somewhat deflexed. Antennae ten^
segmented, the third segment as long as the first and second com-
bined; antennae slightly but distinctly longer than the width of the
head.
Thorax arched; without pubescence. Posterior tibia with five to
six large black teeth at apex; two claws on the basal tarsus. An-
terior wing (Fig. 1, L) slightly opaque, punctate over the entire
membrane, with many brown spots (Fig. 1, M) distributed irregu-
larly as shown in the figure; venation as shown in illustration. Pos-
terior wing rather large but exceedingly delicate in structure, beset
throughout with numerous very minute points, the venation scarcely
discernible, developed as rows of points that are hardly visible even
with the higher power of the microscope.
Abdomen with the dorsal plates somewhat more heavily chitin-
ized than the ventral. Genitalia of the male (Fig. 1, N, O) relatively
large; b of the proctiger very nearly equal to p, the lobes over one-
fourth longer than p, of uniform width in lateral view and broadly
rounded at the apex; the proctiger having two unusually large
heavily chitinized, and meso-dorsally directed hooks at the base of
the lobes; claspers distinctly longer than p. Genital segment of
female scarcely more than one-half the length of the rest of the
abdomen; the anterior half of the segment twice as thick as the
posterior half, this being easily seen even in unmounted specimens
studied with low magnifications (Fig. 1, G) ; the anterior half of the
dorsal valve bearing a number of large setae as illustrated; the apex
of the dorsal valve being roughened and heavily chitinized, the
ventral valve terminating in a strongly chitinized black apex.
Nymph. Unknown.
January, 1931]
KLYVER CHERMID^
137
This species is evidently very closely similar to A. angusti-
pennis Crawford. It is here described as a new species because
of its smaller size; the different proportions of the fore wing;
and the distinctive character of the male and, especially, the
female genitalia. It differs in several important respects from
A. veaziei Patch. It is clearly different from A. veasiei met-
zaria Crawford on the basis of the male, but more importantly
the female genitalia, which are as distinctive or more so than
those of any species in this difficult group.
Aphalara spp.
Specimens. One female from Sophia pinnata (Walt.) Howell,
Ely, Nevada, May 19, 1930 (2531). One male from Chrysothamnus
speciosus Nutt., seventeen miles northwest of Milford, Utah, May 15,
1930 (2502). One female from Chrysothamnus speciosus Nutt., Little-
field, Arizona, May 3, 1930 (342).
These three specimens are undeterminable without addi-
tional representation because of their somewhat fragmentary
condition. Each represents a species different from the species
in this and other collections of the writer.
Subgenus Anomocera Crawford
1914, Anomocera, Crawford. U. S. Nat. Mus., Bull. 85:36-37.
This interesting group is represented by several specimens
in this collection. The antennae, upon casual observation, cer-
tainly appear to be but eight-segmented. The only perfect
antenna in my material is suggestive of two very short seg-
ments being fused to the eighth segment instead of only one
as indicated by Crawford.
Aphalara (Anomocera) minutissima Crawford
Fig. 1, J, K
1914, Aphalara (Anomocera) minutissima Crawford. U. S. Nat. Mus.,
Bull. 85:37; figs.
Specimens. One female irom Dipetalia linifolia (Vahl), Las Vegas,
Nevada, May 3, 1930 (323); two males and seven females from
Artemisia filifolia Torr., St. George, Utah, April 21, 1930 (264).
Male genitalia with the proctiger of the general type repre-
sented by Aphalara suaedcs Crawford, but less robust, with the
lobes relatively longer and bearing a number of stout, sharply
138
THE PAN-PACIFIC ENTOMOLOGIST [vOL, VII, NO. 3
pointed setae; the claspers, as illustrated (Fig. 1, J, K), nearly
as long as the proctiger. Female genitalia similar to those of
A. suaedco Crawford, but somewhat larger, with the valves of
the ovipositor more than usually extended posteriorly.
Genus Aphalaroida Crawford
1914, Aphalaroida Crawford. U. S. Nat. Mus., Bull. 85:38.
This very interesting genus is represented by a single
Fig. 1. Aphalara suaedce Crawford — A, detail of fore wing; B,
genitalia of male; C, inner face of clasper; D, genitalia of female;
H, proctiger, illustrating method of taking measurements. Psyllia
bre<viata Patch — E, male genitalia; F, inner face of clasper. Aphalara
minutistylus n. sp. — G, genitalia of female; L, fore wing; M, detail of
wing; N, genitalia of male; 0, inner face of clasper; I, proctiger.
Aphalara gutierrezia n. sp. — P, ‘outer and inner aspects of clasper of
male. Aphalara (Anomocera) minutissima Crawford — J, K, outer and
inner aspects of clasper of male.
January, 1931]
KLYVER CHERMIDJE
139
species and but a single, somewhat fragmentary male specimen.
All but the first and second antennal segments are wanting,
the posterior wings, and parts of all the legs except one pos-
terior leg, which is complete, are wanting.
In many respects the specimen at hand fits Crawford’s
description of Aphalaroida pithecolobia Crawford, the generic
type, particularly in the anterior wing form and in the male
genitalia. In other respects it is closely similar to A. inermis
Crawford, especially in wing coloration, which here seems
significant.
Aphalaroida intermedia Klyver, n. sp.
Fig. 2, A, B, C, D, E, F, G, M
Specimen. A single male specimen from Covillea tridentata (D. C.),
St. Thomas, Nevada, May 18, 1930 (405).
Type. Holotype, male, FK 233.1.1 S , from Covillea U'iden-
tata (D. C.), St. Thomas, Nevada, May 18, 1930 (405),
Adult. Length of body on slide, 2.1 mm.; width of head, .5 mm.;
length of fore wing, 1.5 mm.; width of fore wing, .65 mm. General
color pale brown throughout. Wings semiopaque, light brown.
Head (Fig. 2, M) slightly wider than width, of thorax; eyes
prominent; antennae unknown; vertex and gense bearing many large,
blunt-ended setae, the genae bearing several other longer and sharply
pointed setae.
Thorax somewhat arched; the dorsum and the sides bearing
many large setae similar to those found elsewhere on the head, wings,
and ventral plates of the abdomen. Legs with many large, chiefly
sharply pointed setae; apex of posterior tibia (Fig. 2, F) with three
large black teeth mesally and a single large tooth and several large
pointed setae on the opposite side; basal tarsus with two relatively
large black claws. Anterior wing having shape and venation as
shown in illustration (Fig. 2, A); the wing membrane bearing large
setae similar to those found on the head, thorax, and ventral plates
of abdomen, and having an intricate system of markings as illus-
trated (Fig. 2, B), more or less punctate over the entire membrane;
five alar radulae developed extensively, but rather feebly, in the two
marginal cells, between and Ra, Ra and ^ and Mg ^ ^ and Cu^^
(Figs. 2, A, C). Posterior wing unknown.
Abdomen with the plates quite heavily chitinized; the dorsal
plates bearing an irregular row of small pointed setae along the
posterior margin; the ventral plates bearing many large setae of the
type found on the head, thorax, and fore wings, Male genitalia
140
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
(Fig. 2, D, E) moderate in size; the proctiger simple, somewhat
elongate; the claspers about half as long as the proctiger, sub-
rectangular in lateral aspect, the apex notched, the anterior process
short, broadly rounded, the posterior process flattened on top.
Female genitalia unknown.
Nymph. Unknown.
Aside from the difference in general proportions, differing
from A. pithecolobia Crawford, to which it is apparently
closely related, in the pronotum being distinctly arched; in the
anterior wing being conspicuously spotted, with the zving mem-
brane bearing relatively large setae, set biseriately with the veins
but always on the wing membrane at some distance from the
veins (Fig. 2, A, B, C, G). Similar to A. pithecolobia Craw-
ford in the proctiger of the male being considerably longer
than the claspers, the proctiger being relatively stout instead of
very long and slender as in that species.
The general appearance of the anterior wing is apparently
similar to that of A. inermis Crawford. On the other hand,
the specimen at hand differs from that species in the following
particulars : in the fore wing being relatively broader and less
rhomboidal ; in the white markings of the fore wing being
uniform in size and distributed over the entire wing surface;
in the presence of small white areas at the ends of Rg, Mi + 2 .
and M 3 + 4 , and Cui, and a small but very noticeable dark
brown spot at the end of Cu 2 (Fig. 2, A) ; and in the ptero-
stigmal vein (Ri) being more complete, delimiting a longer
pterostigmal area.
Subfamily Pauropsyllinaj:
Genus Heteropsylla Grawford
1914, Heteropsylla Crawford. U. S. Nat. Mus., Bull. 85:44.
This genus is represented by over three hundred adult
specimens of Heteropsylla texana Grawford, the type of the
genus, and by two last-stage nymphs of this species.
Heteropsylla texana Grawford
Fig. 2, N
Specimens. More than three hundred adult males and females
and two last-stage nymphs from Prosopis glandulosa Torr., Overton,
January, 1931]
KLYVER CHERMID^
141
Nevada, May 2, 1930 (315); and, adults only, from Artemisia dracun-
culoides Pursh, St. George, Utah, May 3, 1930 (347); from Sophia
pinnata (Walt.) Howell, Ely, Nevada, May 19, 1930 (2530); from
Pluchea sericea (Nutt.) Coville, Glendale, Nevada, May 3, 1930
(338); from Gorton longipes Jones, Littlefield., Arizona, May 3, 1930
(340) ; from Covillea tridenfata (D. C.), Littlefield, Arizona, May 3,
1930 (344); and from Chrysothamnus speciosus Nutt., Littlefield, Ari-
zona, May 3, 1930 (342).
In all of the more important characters my specimens agree
very closely with Crawford’s description of the species. In
coloration they differ considerably, there being a predominance
of bright green color in the abdomen and frequently over the
entire body in the long series of specimens before me. In
several instances of specimens with the head, thorax, and the
dorsum of the abdomen nearly black the abdomen is dark, yet
distinctly green on the ventral side.
Aside from the coloration the morphology of the fore and
hind wings is noteworthy. The membrane of the anterior wing
is covered with small points, the alar radulae being feebly
developed in the two marginal cells and between Ms + 4 and Cui ;
the veins bearing small setae; and the pterostygma is thickened
and beset with numerous punctations. The posterior wing is
exceedingly delicate in texture, the venation being scarcely dis-
cernible with the compound microscope, consisting of a series
of minute punctations arranged in two rows ; the antero-
proximal margin bearing a number of relatively large, stout,
setae.
The claspers of the male are deeply notched at the apex,
with the posterior process somewhat longer than the anterior,
as described by Crawford, but neither is, strictly speaking,
acute at the apex (Fig. 2, N). The anterior process is roundly
pointed ; the posterior process is flattened at the apex and has a
small antero-mesally directed hook as shown in the illustration.
Nymph. The immature stage in my material will be de-
scribed in the series of short papers by the writer devoted to
the descriptions of the immature stages of generic types and
others of special interest.®
5 1930, Notes on the Chermidae. Part I. Can. Bnt., 62:167-175; pi. 14;
1930, Part II. Ibid. In press.
142
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 3
Subfamily Triozin.<e
Genus Paratrioza Crawford
This genus is represented by one commonly known species.
The information here given extends the description by Craw-
ford in several important details.
Paratrioza cockerelli Sulc.
Fig. 2. H, I, J
1909, Paratrioza cockerelli Sulc. Acta Soc. Ent. Bohemiae, 6:102-109.
1914, Paratrioza cockerelli Sulc. Crawford, U. S. Nat. Mus., Bull.
85:71-72; figs.
Fig-. 2. Aphalaroida intermedia n. sp. — A, fore wing; B, detail of
wing membrane; C, alar radulse; D, genitalia of male; E, inner face
of clasper; F, inner and outer aspects of apex of posterior tibia; G,
seta of wing membrane; M, head. Paratrioza cockerelli Sulc — H, I,
and J, distal ends of fourth, sixth, and eighth antennal segments,
respectively. Trioza lobata Crawford — K, genitalia of male; L, inner
face of clasper. Heteropsylla texana Crawford — N, outer and inner
aspects of clasper of male. Psyllia minuta Crawford — 0, genitalia of
female; P, outer and inner aspects of clasper of male.
January, 1931]
KLYVER CHERMID^
143
Specimens. Numerous males and females; from Covillea triden-
tata (D. C.), Glendale, Nevada, April 20, 1930, May 3, 1930 (242;
332, '330); from Erigonum trichopes Torr., Glendale, Nevada, May 3,
1930 (331); from Chrysothamnus paniculatus (A. Gray), Glendale,
Nevada, May 3, 1930 (333); from Hymenoclea salsola Torr. and Gray,
Las Vegas, Nevada, April 20, 1930 (219); from Norta altissima (L.)
Britton, Milford, Utah, May 20, 1930 (2557); from Chrysothamnus
speciosus Nutt, St. George, Utah, May 1, 1930 (282); from Lepidium
scopularum Jones, Garrison, Nevada, May 15, 1930 (2509); from the
same host, Tonopah, Nevada, May 18, 1930 (2525, 2526); and from
Sophia pimiata (Walt.), Ely, Nevada, May 19, 1930 (2528, 2530, 2531).
Observations with the compound microscope (100 x and
440 x) reveal the very interesting and unusual structure on
antennal segment four which has been mentioned by Crawford.
In addition to the one on this segment, however, similar though
smaller structures are found on segments six and eight as illus-
trated in Fig. 2, H, I, J.
The posterior wing of this species, hitherto undescribed, is
very thin, transparent, and delicately membraneous with the
venation obscure, the veins present as a single row of minute,
rather widely spaced punctations, the membrane covered
throughout with exceedingly fine punctations.
Trioza lobata Crawford
Fig. 2, K, L
1914, Trioza lobata Crawford. U. S. Nat. Mus., Bull. 85:86-87; figs.
Specimen. One adult male from Chrysothamnus speciosus Nutt.,
Panaca, Nevada, May 19, 1930 (2538).
This species is represented by a single imperfect specimen,
of which the antennae are wanting. There can, however, be
no question as to its identity because of the distinctive anterior
wings and other characters.
The alar radulae of the anterior wings are moderately de-
veloped in the two marginal cells and between M and Cuj.
Posterior wings relatively large, very thin and membraneous,
with the venation weakly developed as irregularly set single
and double rows of punctations, and with the general surface
of the membrane beset with minute points arranged in circles
of varying sizes suggesting lacework.
Male genitalia (Fig. 2, K, L) as described by Crawford.
His figures of the genitalia lack definiteness and clearness.
(To be Continued)
THE PAN-PACIFIC ENTOMOLOGIST
Published quarterly by the Pacific Coast Entomological Society
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Editorial Comment
In a review of Dr. Howard’s very interesting and valuable
work, A History of Applied Entomology, recently published
as Volume 84 of the Smithsonian Miscellaneous Collections/
Professor Cockerell has called attention to the disproportionate
emphasis heretofore placed upon the purely economic side of
the entomological activities of the United States Government.
I say “heretofore” advisedly, as there recently have appeared
indications of a more suitable recognition of the importance
of purely systematic work in the management of the National
Museum and of the Bureau of Entomology. Speaking of the
prompt and efficient help he had received from the Imperial
Institute of Entomology at London, in the determination of
certain oriental material. Professor Cockerell says “this work
could not have been done in America, or could only have been
done with extreme difficulty. The United States National
Museum has not yet accepted its responsibility for the develop-
ment of an adequate collection of the insects of the world.”
Again he says : “It is at this point that we sense the weakest
aspect of modern American entomology. There are in Wash-
ington and in the experiment stations and universities many
capable workers. But as a general thing they are intellectually
dissipated by a multiplicity of duties. They may be well paid
and not overworked, but they cannot concentrate on compre-
hensive research.” This is essentially true of every scientific
institution I know in this country. The available material may
be ample, but the staff of specialists is pitifully small and
entirely inadequate to study this material and to make it avail-
able through the publication of monographs or the determi-
nation of species for those to whom determinations are impor-
tant. Professor Cockerell’s review should be read by all those
responsible for shaping the policies of our scientific institu-
tions. — E. P. Van Duzee.
1 Science for February, 1931.
A MANUAL OF THE GENERA OF BEETLES OF
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Vol. VII
No. 4
April, 1931
THE
Pan-Pacific Entomologist
Publbhed by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
CONTENTS
THORPE, MISCELLANEOUS RECORDS OF INSECTS INHABITING THE SALINE
WATERS OF THE CALIFORNIA DESERT REGIONS 145
FALL, A NEW GYRINUS FROM CALIFORNIA, WITH REFERENCES TO OTHER
RECENTLY DESCRIBED SPECIES 154
KLYVER, CHERMID.® FROM UTAH, NEVADA, AND ARIZONA, INCLUDING
THREE NEW SPECIES (CONCLUDED) 157
LAWSON, THE GENUS XEROPHLCEA IN NORTH AMERICA 159
HOOD, SYNONYMS IN THE NORTH AMERICAN THYSANOPTERA .... 170
MOULTON, AN INTERESTING NEW CALIFORNIA THRIPS 173
BAILEY, A THRIPS NEW TO CALIFORNIA 175
FENDER, BUTTERFLIES OF YAMHILL COUNTY, OREGON . 179
BARRETT, A NEW BRACHYTARSUS FROM CALIFORNIA 188
CHAMBERLIN, ON THREE NEW CHILOPODS 189
EDITORIAL COMMENT 192
INDEX TO VOLUME VII 193
San FranciscO) California
1931
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The Pan-Pacific Entomologist
Vol. VII, No. 4
April, 1931
MISCELLANEOUS RECORDS OF INSECTS
INHABITING THE SALINE WATERS
OF THE CALIFORNIAN DESERT
REGIONS
BY W. H. THORPE, PH. D., M. A.
Imperial Institute of Entomology
The insect fauna of brackish waters of the western United
States of America is a subject which has attracted the attention
of entomologists since the observations made by A. S. Packard
in 1869. Since that time much new information has been placed
on record, mostly in the form of brief notes scattered through-
out a variety of natural history journals. Many of these ob-
servations are of little scientific value in that no exact data as
to the composition of the water are given. To say that the
water is “brackish” or “salt” is so vague as to be practically
meaningless. In the case of pelagic forms occurring in large
bodies of water, such as Great Salt Lake, this lack of data is
of no great significance, since the salinity is generally fairly
constant and uniform, but in the smaller lakes which are fed
by intermittent streams or springs, and which are liable to
become very much more concentrated during the dry summer
months, it is important to obtain exact data as to the salinity
of the medium at the actual spot where the insects are collected.
In such lakes one frequently finds very great variation in den-
sity within quite short distances and there may be very marked
differences between the surface and the bottom layers. More-
over, pools which appear closely similar may differ enormously
in salt content. It follows, therefore, that unless great care
is taken the observations may be quite misleading.
The following notes were made, as opportunity offered,
during various expeditions into the desert regions of California
during 1928 and 1929. The observations are unfortunately
very disconnected and incomplete, but as they are accompanied
in most cases by exact data as to the environmental conditions
146
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
the writer feels that they are worth putting on record. Owing
to the inaccessibility of many of the localities, details as to the
life history and the degree of salinity which the various de-
velopmental stages are capable of withstanding are not known
and will probably remain unknown for a long time to come.
Such information, combined with a knowledge of the salinity
variations throughout the year, would, however, be of the
greatest interest for the work of Poisson (1924) and others,
and has shown that a density which is quite harmless to an
adult insect or to a late stage larva may, nevertheless, be suffi-
cient to destroy the eggs or young larvae, perhaps by interfer-
ence with the hatching mechanism. Were such details known
the reasons for the apparently capricious distribution of many
of these halophilus insects would doubtless be more obvious.
It is to be hoped that others will follow up this line of work.
A collector with more time at his disposal would not only be
able to add to our knowledge in the ways outlined above, but
would probably extend considerably the list of species from
an area such as Death Valley. Owing to the great variations
in the physical and chemical conditions a species may be very
abundant in one spot, where conditions are at the optimum, and
yet be nonexistent elsewhere, consequently even in a small area
a great deal of collecting is needed before one’s list is complete.
Death Valley Region. — A brief trip to the Death Valley
region of California (Inyo County) was made during Febru-
ary 23-27, 1929. The south end of the valley was first visited.
Where the bed of the Amargosa River was crossed no water
was seen, but at camp near Old Confidence Mill biting midges
were very troublesome, Culicoides varipennis Coq., and Lepto-
conops kerteszi var. americanus Carter, being taken. Various
species of the genus Culicoides are known from salt and
brackish waters in many parts of the world, and the species
above referred to had no doubt bred in the highly saline tempo-
rary pools which represent the Amargosa River. C. varipennis
is already known as a common fresh-water breeder in Cali-
fornia and New Mexico, and L. kerteszi also breeds in fresh
water in north and central California, as well as at Great Salt
Lake, Utah.
Some collecting was done at Salt Creek and McLean
Springs, on the valley floor, about two miles southwest of
APRIL, 1931] THORPE SALINE WATER INSECTS
147
Stovepipe Wells, on the 2'5th of February. A trickle of water
running over the alkali caked mud at Salt Creek yielded the
following results :
Coleoptera:
DytiscideB. Hydroporus panaminti Fall; Hydroporus spp. (larvse
of two species).
Hydrophilidae. Octhebius rectus Lee.; Enochrui diffusus Lee.
Diptera:
Chironomidae. Orthocladius sp. (larvae and adults).
Two still pools at McLean Springs, “a” and “b,” about
eighteen inches deep and three feet in diameter, were also
examined. The first contained the unidentifiable remains of a
dragon fly larva, and at its margin the salt marsh Carabid,
Tachys vittiger, was obtained. Pool “b” gave further speci-
mens of H. panaminti and some empty caddis cases (Limno-
philidas), and in a very small pool near by 0 . rectus was again
taken.
Further south collections were made in shallow pools con-
nected with a slow-flowing stream on the east side of the valley
floor, about one and a half miles north of Old Harmony Borax
Mill. Here H. panaminti was again found, together with two
species of Hydroporous in the larval stage. Besides 0. rectus
and a larva referable to it, larvae of Ephydra sp. (hians ?) were
present in considerable numbers.
Hurried collections made in a less concentrated pool at
Eagle Borax Well yielded another Hydrophilid, Tropisternus
calif ornicus Lee., and larvae of a species of Procladius.
The salinity of these various waters, in parts per million,
is given below :
Salt Creek
McLean
Springs “a"
McLean
Springs “b”
Pools Near
Old Harmony
Mill
Eagle Borax
Well
Cl
7,228
2,378
2,840
11,863
412
C03
48
0
0
0
0
HC03
50
537
500
850
135
so.
3,272
1,610
1,710
3,766
597
Ca
112
100
92
84
128
Mg
78
53
30
27
61
Na
6,200
2,300
2,737
9,010
345
Borax
60
20
40
300
10
Total
17,048
6,998
7,949
25,900
1,688
Sp. gr.
1.016
1.006
1.007
1.024
1.001
148 THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
Owens Lake. — A visit was paid to Owens Lake, Inyo
County, California, on July 7, 1928. The lake at this time was
very low and on the western side, where the collecting was
done, it was impossible to reach water more than half an inch
in depth, overlying a slimy mud, which was swarming with
Dipterous larvae and pupae, nothing else being found. There
were great numbers of Ephydra hians Say, a species already
recorded from this locality by Aldrich (1912). Pupal cases of
a large Tabanus and many larvae and pupae of Odontomyia
were found. The first named belonged to T. punctifer O. S.,
a common inhabitant of alkaline waters in the West (Webb
and Wells), adults of which were caught in the near vicinity,
and the latter are probably referable to 0. tumida Banks, and
0. arcuata Loew., both of which were obtained. No exact
chemical and physical details of the environment of the latter
two species appear to be on record. Brues (1928) records
T. punctifer as occurring in waters with a specific gravity up
to 1.014.
The composition of Owens Lake is already well known.
According to Clarke (1924), the total salinity varies from
15,900 parts per million (specific gravity 1.015) to 213,700
parts per million (specific gravity 1.153). Considering the time
of the year and the level of the lake, the insects obtained must
have been living under conditions of salinity approaching those
indicated by the higher figures.
The percentages of the chief constituents in the more saline
samples are given below. The data have been calculated in
parts per million, and are given in the second column so that
the figures are comparable with those given in the other tables :
Percentage of Total
Salinity
p. p. m.
Cl
24.82
53,425
C 03
24.55
52,463
S 04
9.93
21,220
Na
38.09
81,398
K
1.62
3,461
Specific Gravity
1.153
Mono Lake. — Owing to its greater depth and large size.
Mono Lake is correspondingly more constant in composition.
The following data are recalculated from Clarke :
APRIL, 1931] THORPE — SALINE WATER INSECTS
149
Salinity
Specific gravity
Cl
CO3
SO,
Na
K
51,170 p. p. m.
1.045
11,943 p. p. m.
11,984
6,580
19,408
. 946
The only aquatic insect definitely known from Mono Lake
appears to be Ephydra hians (Aldrich, 1913, and many other
observers), and this was the only one found by the writer on
a very brief visit in October 1928. Aldrich records Tabanus
opacus Coq., and T. phcsnops O. S., but it appears uncertain
whether these were breeding actually in the lake or in streams
running into it. Ephydra subopaca Loew., is also mentioned
as occurring in seepage near the lake. Many other Diptera
were recorded by Aldrich in the vicinity, but the breeding
habits were not known.
Salt on Sea. — This body of water is of particular interest in
view of the great changes in salinity which must have taken
place since its formation forty years ago. From 1891 till 1905
the lake was small and the salt content presumably high,
although no data for that period appear to be available. From
1905-1907 fresh water from the Colorado River flowed in, with
the result that the lake more than doubled its size. In 1911 the
salinity was found to vary from 7164-7348 parts per million
(MacDougal, 1914). After 1907 the inflow was checked, with
the result that the salinity again began to increase and in 1913
had reached a salinity of 10,026 parts per million.
Hubbard in 1897 recorded four species of Coleoptera at a
point near Salton where some springs, said to contain “1 to 6
per cent of saline matter,” discharged into the sea. An un-
described species of Creniphilus (Paracymus) was described
as present in vast numbers, with Philhydrus dijfusus Lee. ; and
Octhebius rectus Lee., and a new species of the same genus
were found less commonly. Hubbard also records an Ephydra
different from that found in Great Salt Lake.” Aldrich (1912)
quotes a record of E. gracilis from this locality, but the speci-
mens in my possession are all E. subopaca, not gracilis. Hub-
bard, in addition, records several salt marsh Staphylinids and
Carabids.
150 THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 4
I was, unfortunately, not able to reach the exact spot men-
tioned by Hubbard, but two visits to localities on the west
shore of the lake failed to yield anything except immature
Corixid^e, which could not be identified. All the specimens that
I have from this particular region are two Ephydrids collected
by Dr. E. A. Andrews, March 3, 1929, and very kindly handed
over to me. They are Ephydra suhopaca, and a Paralimna sp.,
but the latter can hardly have been breeding in the lake and
there is some doubt about the Ephydra.
In contrast to the lack of insects in the Salton Sea, the
insect fauna of a small tule-surrounded pool at Dos Palmas,
a short way from the eastern shore of the lake, is of interest.
This pool is fed by a luke-warm spring, an analysis of which
is given below. Here, on December 12, 1928, larvse of a spe-
cies of Procladius were present in abundance, and larvae of
Odontomyia sp. were also found. Five species of Coleoptera
were taken as follows :
Dytiscidae. Laccophilus mexicanus Aube; Laccophilus terminalii
Sharp; Rhantus binotatus Harris.
Hydrophilidae. Tropisternus dorsalis Brulle.
A few hundred yards away was a pool of fresh water con-
taining abundant plant growth and mosquito larvae, probably
referable to Anopheles pseudopunctipennis Theob., and Theo-
holdia inornata Will., larvae of Odontomyia sp. were also
found. Here also occurred the Desert Minnow, Cyprinodon
macularcus.
Analysis of springs at Dos Palmas :
Dos Palmas
Canyon Springs
Cl
470
124
C 03
0
0
HC03
115
155
so.
120
736
Ca
46
120
Mg
45
65
Na
270
230
Total Solids
1,020
1,388
A stagnant pool at Canyon Springs, about eight to ten miles
away, contained nothing but mosquito larvae, almost certainly
those of T. inornata, a species already known from saline
waters (Brues, 1928).
APRIL, 1931] THORPE SALINE WATER INSECTS
151
Discussion
Little need be said as to the composition of the waters, the
tables will explain themselves. For the sake of comparison it
may be said that the average specific gravity of sea water is
1.027, that of pure water, of course, being 1.000. It will thus
be seen that many of the samples are much saltier than the sea,
the most concentrated being that from Owens Lake, with a
specific gravity 1.153. The waters are for the most part mark-
edly alkaline, owing chiefly to the presence of large quantities
of dissolved alkaline carbonates, and in some cases borax. It
is, however, fairly well established that differences in p.n. of
the medium have very little direct effect on aquatic insects,
although, of course, the indirect effect by influencing the fauna
and flora as a whole, may be very great. The high proportion
of sulphates should also be noted.
In considering these waters as biological environments the
most important point to know is, of course, the osmotic pres-
sure. This cannot be measured in the field and, in the case of
highly concentrated solutions, can only be estimated with some
difficulty in the laboratory. In the case of sea water and brines
produced by the evaporation of sea water. Fox (1926) has
shown that no serious error is involved if it is assumed that
the salts are completely ionized. Then the osmotic pressures
can be taken as identical with those of solutions of pure NaCl
of the same specific gravity, and for which the freezing point
depressions are accurately known. This procedure is, of course,
much less reliable in the case of waters such as are discussed
in this paper, the salts of which are certainly not completely
ionized, and figures obtained in this way will consequently tend
to be too high; nevertheless, one or two examples, while only
approximate, will serve to give some idea of the order of
magnitude of this factor. Thus the osmotic pressure of Mono
Lake would be about 50 atmospheres, that of the most concen-
trated of the Death Valley pools 28 atmospheres, whilst Owens
Lake might reach as high as 175. The fact that the figures for
Death Valley waters are so much lower than those for Owens
Lake is probably due merely to the difference in the season at
which the observations were made.
152
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 4
The observations here recorded supply further illustrations
of the fact, already noted in many parts of the world (e. g.,
Thorpe, 1927), that the Diptera are not only able to support
life in media of an osmotic pressure that is rapidly fatal to
most other insects, but that they have also a greater power of
adaptation to changes in concentration than have other orders.
As will be seen from these notes the waters of greatest salinity
are occupied mainly by Dipterous larvae. Coleopterous and
Trichopterous larvae occurring only at respectively lower con-
centrations.
Of the Ephydrids, gracilis and hians are well known as
forms confined to saline waters. About E. subopaca there is
some doubt. Aldrich says that in the West it is characteristic
of the less dense waters (specific gravity 1.000 to 1.019), while
Ping (1921), working in the East, states that it cannot go
through its early stages in fresh water. The record from
Salton appears to extend its known range in California by some
five hundred miles to the south.^
Although unidentified Stratiomyidae have frequently been
recorded from saline and alkaline waters (Brues, 1928), the
records from Owens Lake indicate a considerably higher salin-
ity range than usual.
No exact data as to the habitat of any of the Coleoptera
met with, excepting T. dorsalis, appear to have been recorded
previously. T. dorsalis is recorded by Brues (1928) at specific
gravity 1.0008 to 1.0015. With the possible exception of
H. panaminti, the records suggest that all of these insects are
also capable of developing in fresh water. The record of this
species from Dos Palmas extends its known range south by
two hundred miles. Dr. Fall tells me it was previously known
only from the Death Valley region. It is seen to have a wide
salinity range (specific gravity 1.001 to 1.024).
Finally I wish to express my indebtedness to Dr. H. C. Fall
and Dr. O. A. Johannsen for determinations of the Coleoptera
and Diptera respectively, and to the chemical department of
the University of California Citrus Experiment Station at
Riverside, for undertaking water analysis ; to Mr. E. C. Jaeger
of the Biology Department, Riverside Junior College, I am
1 Unless E. m i 1 1 b r se, Jones, a common coast species in south-
ern California, eventually proves to be identical.
APRIL, 1931] THORPE — SALINE WATER INSECTS
153
much indebted, for without the advantage of his experience of
desert travel some of the trips would not have been attempted.
Bibliography
Aldrich, J. M. 1912. “The Biology of Some Western Species of
the Dipterous Genus Ephydra.” J. N. Y. Entom. Soc., 20,
pp. 77-98.
Aldrich, J. M. 1913. “Collecting Notes from the Great Basin and
Adjoining Territory.” Entom. News, 24, pp. 214-221.
Brues, C. T. 1924. “Observations on Animal Life in the Thermal
Waters of Yellowstone Park, with a Consideration of the Ther-
mal Environment.” Proc. Am. Acad. Arts and Sci., 59, pp. 371-
437.
Brues, C. T. 1928. “Studies on the Fauna of Hot Springs in the
Western United States, and the Biology of Thermophilous Ani-
mals.” Proc. Am. Acad. Arts and Sci., 63, pp. 139-228.
Clarke, F. W. 1924. “Composition of River and Lake Waters of
the U. S. A.” Dept, of Interior, U. S. Geo. Survey, Professional
Paper No. 135, Washington, D. C.
Fox, H. Munro. 1926. “Cambridge Expedition to the Suez Canal —
General Part.” Trans. Zool. Soc. London, 22, pp. 1-64.
Hubbard, H. S. 1901. “Salton Lake in the Colorado Desert and
Its Insect Fauna.” Proc. Entom. Soc. Washington, 4, pp. 376-8.
MacDougal, D. T. 1914. “The Salton Sea.” Carnegie Inst., Wash-
ington. Publication No. 193.
Packard, A. S. 1869. “On Insects Inhabiting Salt Water.” Proc.
Essex Inst. Salem, 6, pp. 41-51.
Ping, C. 1921. “The Biology of Ephydra subopaca, Loew.” Cornell
Univ. Agric. Exp. Station Memoir, 49, pp. 561-612.
Poisson, R. 1924. “Contribution a Tetude des Hemipteres aqua-
tiques.” Bull. Biologique, Paris, 58, pp. 49-305.
Thorpe, W. H. 1927. “The Fauna of Brackish Pools of the Sussex
Coast.” Trans. S. E. Union Sci. Societies, 1927, pp. 27-34.
Webb, J. L., and Wells, R. W. 1924. “Horse Flies: Biologies and
Relation to Western Agriculture.” U. S. D. A. Dept. Bull.
No. 1218, p. 36. Washington, D. C., 1924.
154
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
A NEW GYRINUS FROM ALASKA, WITH
REFERENCES TO OTHER RECENTLY
DESCRIBED SPECIES
(Coleoptera)
BY H. C. FALL
Tyngsboro, Massachusetts
Since the appearance of my paper on the “North American
Species of Gyrinus” (Trans. Am. Ent. Soc,, xlvii, 1916), three
additional species have been described as follows: G. hatchi
and G. dubius by Mr. J. B. Wallis (Can. Ent., April, 1926),
and G. gehringi by Mr. K. F. Chamberlain (Bull. Brook. Ent.
Soc., October, 1929).
G. hatchi Wallis. The type was collected by Mr. Melville
H. Hatch at Crooked Lake, Washtenaw County, Michigan. It
measures 4.69 mm. in length, color above black with bluish
reflections, margins bronzed, elytra moderately polished but
distinctly micropunctulate, beneath entirely rufotestaceous. It
is perhaps nearest fratermts, but is smaller and with somewhat
different male genitalia.
G. dubius Wallis. Type from St. Anthony, Newfoundland.
According to Wallis, very close to picipes but more alutaceous
and micropunctulate above with outer strial punctures less evi-
dently larger than those near the suture.
G. gehringi Chamb. Type from Moose Pond, Pittsburg,
in the Connecticut Lake region of New Hampshire. Said to
most nearly resemble lecontei, but is uniformly rufotestaceous
beneath and with quite different genitalia, the middle lobe being
triangular, the sides nearly straight and rapidly converging to
the sharply acuminate tip, which is strongly laterally com-
pressed. Length, 5.5 to 6.2 mm.
G. piceolus Blatch. This species, founded on two female
examples, was suppressed by me as insufficiently distinct from
pemitidus. More recently Mr. Wallis (Can. Ent., February,
1926) has restored the species to specific standing on the basis
of two males, one of which having been sent to Dr. Blatchley
was declared by him to be identical with his type of piceolus.
These males while “resembling pemitidus very closely” are said
to show genitalic differences sufficient to warrant their being
APRIL, 1931]
FALL ^A NEW GYRINUS
155
given specific standing. Assuming the pale hypomera and epi-
pleura to be characteristic, the species runs to pleuralis and
affinis in my table, neither of which it at all resembles.
The following new Gyrinus from Alaska, recently received
from Mr. R. E. Barrett of Saticoy, California, is another inter-
esting addition to our fauna.
Gyrinus instabilis Fall, n. sp.
Nearly related to wallm in form, color and most structural
features, but of average smaller size.
Typical form. Moderately convex, black, two outer elytral inter-
spaces more or less bronzed; disc of the elytra in the male with an
obscure greenish reflection between striae 4 and 7, faint but usually
perceptible by oblique illumination; not definitely visible in the
female; punctures not bronzed. Surface shining, the luster a little
dulled in the female by an evident micropunctulation on a minutely
alutaceous ground. In the male the alutaceous ground sculpture is
scarcely if at all detectable and the micropunctulation is sparser and
feebler, though always present. Striae weakly impressed, the outer
ones a little more noticeably so but not canaliculate; eleventh stria
close to the margin; strial punctures slightly coarser externally.
Body beneath, including the epipleura, metallic black. Front legs
rufoferruginous; middle tibiae, and hind tibiae and tarsi flavotesta-
ceous; middle tarsi except at base, and middle and hind femora more
or less infuscate; claws piceous.
Male genitalia. Color piceous, the middle lobe paler basally;
middle lobe nearly parallel-sided in fully apical half, at apex a little
less than half the width of the lateral lobes; upper surface flattened
for a considerable distance from the tip, which is broadly arcuato-
truncate throughout its width.
Length, 4.7S to S.6 mm.; width, 2.4 to 3.15 mm.
Variations. In a few examples the upper surface is entirely
black without trace of metallic luster. In others the surface is
polished, with the suture narrowly and the punctures bronzed.
In these last the micropunctulation is nearly or quite wanting
in the male but present in the female. The genital armature of
the male is identically the same in these variants as in the
typical form.
Described from a good series of specimens sent me by
Mr. Barrett, who collected them all at Circle, Alaska, June 21,
1928. The type is a male in my own collection.
This species very much resembles wallisi, which, however,
in series is a perceptibly larger species (5.2 to 6.8 mm.). The
156 THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 4
total absence of asneous luster along the margins and in the
punctures is much more frequent in wallisi than in the present
species, and the seneous marginal stripe when present is as
a rule narrower, more obscure and less definitely limited in-
teriorly, but in a few examples it diffuses inwardly over the
greater portion of the elytra. This condition does not obtain
in any example of instahilis, in which the aeneous marginal
stripe is more definitely limited and seems never to cross the
ninth stria. The obscure longitudinal deep greenish streak at
the middle of each elytron in males of the typical form of
instahilis (those without bronzed punctures) is faintly detect-
able in two similar males of wallisi.
In final analysis comparison shows that, while not greatly
unlike, there are constant appreciable differences in the geni-
talia of these two species. In wallisi the middle lobe is slightly
more slender, gradually tapering to the narrowly rounded apex,
the upper surface flattened only at the immediate apex. The
form in instahilis has been stated above.
Two Additions to Our Lists
I recently noticed among the Elaterids of the Snow collec-
tion at the University of Kansas two examples of the genus
Cardiophorus, taken in the Baboquivari Mountains, Arizona,
many years ago by Professor Snow and labelled aptopoides
Cand., a species apparently not previously reported as having
been taken north of Mexico. I submitted the specimens to
W. S. Fisher of the National Museum, who confirmed the
determination.
Among the Endomychidae are two examples of Epipocus
suhcostatus Gorham, also taken in the Baboquivari Mountains,
Arizona, by Professor Snow. The determination was made by
Charles Liebeck and the specimens seem to agree perfectly
with description in the Biologia. — Warwick Benedict, Univer-
sity of Kansas.
APRIL, 1931]
KLYVER CHERMID^
157
CHERMID^ FROM UTAH, NEVADA, AND ARIZONA,
INCLUDING THREE NEW SPECIES
(Homoptera)
BY F. D. KLYVER
San Mateo Junior College, San Mateo, California
(Continued from page 143)
Subfamily Psyllinje
Genus Psyllia
1905, Psyllia Kirkaldy. Wien. Ent. Zeit., 24:268.
Psyllia breviata Patch
Fig. 1, E. F
1912, Psylla breviata Patch. Maine Agr. Exp. Sta., Bull. 202:215.
Specimens. Two males from Chrysothamnus speciosus Nutt., St.
George, Utah, May 1, 1930 (282); one female from Salsola pestifer
A. Nels., St. George, Utah, May 17, 1930 (360); one female from
the same host, Ely, Nevada, May 19, 1930 (2527); one female from
Pluchea sericea (Nutt.), Glendale, Nevada, May 3, 1930 (338); one
female from same host, Overton, Nevada, April 20, 1930 (235); one
female from Bassia hyssopifolia (Pall.), Logandale, Nevada, May 2,
1930 (305); one female from Covillea tridentata (D. C.), St. Thomas,
Nevada, May 18, 1930 (405); one male from the same host, Glendale,
Nevada, May 3, 1930 (330); and one male from same host, St.
George, Utah, May 1, 1930 (286).
The specimens at hand are referred to this species with con-
siderable question. They are in agreement with the general
characters of the species as given by Crawford except for the
genitalia. For instance, the female genital segment in my speci-
mens is approximately one-half the length of the rest of the
abdomen, instead of being equal in length as stated by Craw-
ford. This discrepancy may very well be a direct result of the
method of measuring employed. It is a commonly observed
fact in the long series of my material of this and other species
that the genital segment of the female in unmounted or in dry
mounted specimens will always appear to be proportionately
longer than it actually is, due to the collapsing of the remainder
of the abdomen. There is very little in common, furthermore,
between the male genitalia as illustrated in Fig. 1, E, F, and
the male genitalia of Psylla breviata Patch as drawn by Craw-
ford. This discrepancy may, again, conceivably be assigned to
a difference in interpretation by the technique here employed.
158
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
Observations with the higher magnifications of the micro-
scope disclose the following structures not mentioned by Craw-
ford. The fore wing is beset with rather large points at the
distal end, these points becoming gradually less pronounced
from the distal to the proximal end, while nowhere do they
occur on the veins. Alar radulae are sparsely developed as large
points in the two marginal cells, between Rg and Mi + 2, and
between Mg + 4 and Cu. The posterior wings are relatively
large and thick, the venation being conspicuous as a single row
of points which appear somewhat larger than the similar points
that rather densely beset the entire wing membrane.
Psyllia minuta Crawford
Fig. 2, O, P
1914, Psylla minuta Crawford. U. S, Nat. Mus., Bull. 85:142; figs.
Specimens. One male and six females from unknown host, Rich-
field, Utah, June 21, 1930 (no number); and one female from
Lepidium scopulorum Jones, Garrison, Nevada, May 15, 1930 (2509).
The body and wings agree with the description by Crawford.
The color notes given by him, however, are not applicable to my
specimens. There is very little variation, the general color over the
entire body is light brown. The extreme tip of the antennae and the
apex of the female genital segment are black. The fore wings are
fumate throughout with the apical half sometimes distinctly darker,
the wing membrane is beset with numerous small points that
decrease in size and conspicuousness from the distal to the proximal
end. Four alar radulse are present, one in each of the two marginal
cells, one between Ra and and one between Mg ^ ^ and Cuj^. The
posterior wing is delicately membraneous, punctate throughout, with
the venation developed as faint ridges along which are rows of small
points irregularly arranged.
The claspers of the male terminate at the apex in a distinctly
anteriorly directed claw (Fig. 2, P). The female segment is scarcely
half as long as the rest of the abdomen (Fig. 2, O), the ventral
valve being hardly more than one-third the length of the dorsal
valve, the apex of each bearing a number of very short, stout setae,
as illustrated.
APRIL, 1931]
LAWSON — XEROPHLCEA
159
THE GENUS XEROPHLCEA IN NORTH AMERICA
(Homoptera, Cicadellidae)
BY PAUL B. LAWSON
Lawrence, Kansas ^
The genus Xerophloea, erected by Germar in 1839, is one
of the genera in the subfamily Gyponinse. The members of
the genus may be characterized as follows :
Medium sized or large leafhoppers, with ocelli on flattened
vertex and closer to caudal than cephalic margin, vertex with
margins acute; whole dorsal surface distinctly and coarsely
pitted with pits bearing minute setae; elytra more vertical than
in Gypona. Color of females, except in a few cases, nearly
uniformly green ; males green or varying through yellowish to
brown, with vertex and pronotum frequently marked with dark
brown and veins of elytra frequently appearing mottled. In
both sexes small dark spots occur on margins of veins in caudal
half of elytra and sometimes over other parts of the body.
Female ventral segment long, posterior margin bilobed, with
narrow median slit extending nearly to base. Last ventral seg-
ment of male long, hiding valve; plates long and finger-like,
extending to tip of pygofer.
The uniform coloration of the species and the very great
similarity of the genitalia are undoubtedly responsible for the
fact that hitherto but two species, X. uiridis Fabricius and
X. major Baker, have been recognized in the United States.
Both the external and the internal genitalia are so similar in
the species studied that, although they sometimes show small
characteristic differences, it would be impossible to use them
as diagnostic characters of certain value.
Through the kindness of Mr. E. P. Van Duzee, who has
loaned us the material from the California Academy of Sciences
and his own private collection, and through the collections of
the past several summers in the south by Dr, R. H. Beamer
of the University of Kansas and his survey party, a large
number of specimens of the genus from California to Florida
were made available for study. In addition, many specimens
from northern states were available, so that the present study
1 Contribution from the Department of Entomology, University of
Kansas.
160
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 4
was made from ample material from nearly every part of the
United States.
In spite of the fact that two of the chief characters used
in taxonomic work with leafhoppers, namely color and geni-
talia, are of little or no value in this genus, it was soon found
that the shape of the vertex and the degree of inflation of the
front are very valuable and dependable characters. These,
along with size and geographical distribution, were found to
be sufficient to enable a proper differentiation of the several
species. As a consequence eleven new species are here de-
scribed, a number of which have undoubtedly been previously
included in X. viridis.
Following is an artificial key for the separation of the
species :
Key to Species
1. Front distinctly inflated 2
Front at most but slightly inflated 6
2. Female with entire dorsal surface spotted with brown
brunnea, n. sp.
Females uniformly green 3
3. Vertex of female at most about half length of pronotum 4
Vertex of female distinctly more than half length of pro-
notum 5
4. Front very strongly inflated; vertex shorter infiata n. sp.
Front less strongly inflated; vertex longer robusta n. sp.
5. Larger, more robust species, females over 6 mm
‘vanduzeei n. sp.
Smaller, more slender species, females under 6 mm
californica n. sp.
6. Vertex broadly rounded major Baker
Vertex distinctly angulate 7
7. Males not at all greenish 8
Males definitely greenish or yellowish green 9
8. Males robust, ashy gray, strikingly dotted with dark brown
oraclis n. sp.
Males very slender, cream-colored, strongly marked with
brown spots and lines tenuata n. sp.
9. Larger species, females usually over 7 mm majesta n. sp.
Smaller species, females usually under 7 mm 10
10. Vertex of female nearly as long as pronotum 11
Vertex of female distinctly shorter than pronotum 12
11. Vertex of female longer, sides straighter, apex more obtuse
obtusa n. sp.
Vertex of female slightly shorter, sides more convex, apex
more acute zionis n. sp.
APRIL, 1931]
LAWSON XEROPHLCEA
161
12. Species smaller,' females under 6 mm minor n. sp.
Species larger, females 6-7.5 mm viridis (Fabr.)
Xerophloea zionis Lawson, n. sp.
Figs. 11, 11a
Close to X. viridis but with longer vertex. Length, female,
6.5 to 7 mm. ; male, 6 mm.
Head slightly narrower than prothorax; vertex one-third wider
between the eyes than long, anterior margins distinctly convex, apex
obtusely angulate; front slightly inflated. Pronotum one-fourth
longer than vertex, anterior margin moderately convex.
Color: Females uniformly bright green above except for brownish
eyes and ocelli; below more yellow, with tibiae and tarsi bright
green. Males yellowish green; vertex with median, longitudinal
brown stripe; pronotum with posterior half and middle third of
anterior half brownish.
Holotype, female, Zion National Park, Utah, August 13,
1929, Paul W. Oman. Allotype, male, same data, R. H.
Beamer. Paratypes, a female, same data, Paul W. Oman ; three
females and a male, same data, R. H. Beamer; a female. Pine
View, Utah, July 21, 1922, E. P. Van Duzee.
Last paratype in collection of California Academy of
Sciences ; all other types in Snow entomological collection.
Xerophloea vanduzeei Lawson, n. sp.
Figs. 5, 5a
Close to X. sionis but with narrower head and front more
definitely inflated. Length, female, 6 to 6.5 mm. ; male, 5.5 mm.
Head distinctly narrower than prothorax; vertex one-third wider
between the eyes than long, anterior margins nearly straight, apex
obtusely but strongly angulate; front definitely inflated. Pronotum
one-third longer than vertex, anterior margin strongly convex.
Color: Females uniformly bright green above except for brownish
eyes and ocelli; yellowish green below. Males straw yellow or
greenish yellow; median longitudinal line on vertex, posterior half
and median third of anterior half of pronotum, small spots on veins
and larger ones along claval suture, brown.
Holotype, female, and allotype, male, San Jacinto Moun-
tains, California, July 21, 1929, R. H. Beamer. A large series
of piaratypes, taken largely by E. P. Van Duzee and R. H.
Beamer, are at hand from the following localities in California :
San Jacinto Mountains, Lemon Grove, San Diego County,
Pittsburg, Niles Canyon, San Mateo, Lake Tahoe, Coachella
162
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 4
Valley, Mill Creek Canyon, Cazadero, San Francisco, Marin
County, Cay ton, Poso Creek, Selma, Potholes, Alameda
County.
Twenty-four paratypes in collection of California Academy
of Sciences; all other types in Snow entomological collection.
The writer is glad to name this species in honor of Mr.
E. P. Van Duzee, through whose generous loan of material this
study was made possible.
Xerophloea obtusa Lawson, n. sp.
Figs. 8, 8a
Allied to X. wnduseei but with longer and more obtusely
pointed vertex. Length, female, 6.5 to 7 mm.
Head definitely narrower than pronotum; vertex four-fifths as
long as width between eyes, margin very thin, apex broadly and
obtusely rounded; front nearly straight. Pronotum slightly longer
than vertex, anterior margin moderately convex.
Color: Yellowish green except for pink ocelli and brown eyes;
elytral veins often with white spots. Underside more strongly
yellow.
Holotype, female, San Jacinto Mountains, California, July
21, 1929. Paratypes: Orange County, California, July 14, 1929;
San Jacinto Mountains, California, July 21, 1929; Laguna
Mountains, California, July 6, 1929. All four type specimens
taken by R. H. Beamer.
All types deposited in Snow entomological collection.
Xerophloea minor Lawson, n, sp.
Figs. 6, 6a
A small robust species, allied to viridis, but stouter. Length,
female, 5 to 6 mm. ; male, 5 mm.
Head definitely narrower than pronotum; vertex about two-thirds
as long as width between eyes, anterior margins nearly straight,
apex obtusely angulate; front slightly but definitely inflated. Pro-
notum one-third longer than vertex.
Color: Females uniformly bright green above except for pink
ocelli and brown eyes; underside more yellowish. Males usually
yellowish green with usual brown line on vertex, disc and posterior
half of pronotum and spots on claval suture, brown. Some speci-
mens straw yellow, more strongly marked with brown on vertex,
pronotum, scutellum, and spots on veins of elytra.
Holotype, female, Hidalgo County, Texas, July 31, 1928,
R. H. Beamer. Allotype, male, Hidalgo County, Texas, July
28, 1928, R. H. Beamer. Paratypes, a large series from the
APRIL, 1931]
LAWSON XEROPHLCEA
163
following localities : Cameron, Booie, Hidalgo, Harris, and
Orange counties, Texas; Beaugard and Natchitoche counties,
Louisiana; Biloxi, Agricultural College, Mississippi; Fort
Pierce, Fruitville, Key Largo, Hilliard, Florida; Polk County,
Arkansas.
Four paratypes deposited in collection of California Academy
of Sciences ; all other types in Snow entomological collection.
Xerophloea californica Lawson, n. sp.
Figs. 3, 3a
Allied to X. vanduseei but smaller, with front more strongly
inflated and vertex relatively longer. Length, 5.5 to 6 mm.
Head very slightly narrower than pronotum; vertex about three-
fourths as long as width between eyes, anterior margins slightly
convex to obtusely angulate apex; front broadly and strongly in-
flated. Pronotum about one-third longer than vertex, anterior
margin moderately convex.
Color: Varying from bright to yellowish green, often with small
black dots all over surface; elytral veins sometimes spotted with
brown.
Holotype, female, San Diego County, California, July 4,
1929, P. W. Oman. Paratypes, two same data; two others,
San Jacinto Mountains, California, July 21, 1929, R. H.
B earner.
Types deposited in Snow entomological collection.^
Xerophloea brunnea Lawson, n. sp.
Figs. 4, 4a
Allied to X. californica, but females are brown and vertex
distinctly shorter. Length, female, 5.5 to 6 mm.
Head nearly as wide as pronotum; vertex very wide, about two-
thirds as long as width between eyes, margins convex to obtuse
apex; front strongly inflated. Pronotum about one-third longer than
vertex.
Color: Dirty yellow, spotted all over with tiny brown dots, giving
it a distinctly brownish appearance.
Holotype, female, and a paratype. Orange County, Cali-
fornia, July 14, 1929, R. H. Beamer.
Types deposited in Snow entomological collection.
2 Note by the editor: The following: additional specimens of this spe-
cies were returned by Professor Lawson to the California Academy of
Sciences labeled “Paratypes”: Keen Camp, Riverside County, California,
June 6, 1917 (Van Duzee); Alpine, San Diego County, California, Octo-
ber 4, 1913 (Van Duzee): same place, September 13, 1923 (Van Duzee);
San Diego County, California, April 8, 1925 (Van Dyke), Selma, Fresno
County, California, June 4, 1929 (Van Duzee).
164
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
This is the only species in the genus yet known in which
the females depart entirely from the usual uniform green
coloration and assume the color of some of the darker males.
Xerophloea robusta Lawson, n. sp.
Figs. 2, 2a
A very stout species allied to X. hrunnea but more robust
and with front more strongly inflated. Length, female, 6 to 6.5
mm. ; male, 5.5 mm.
Head distinctly narrower than pronotum; vertex nearly twice as
wide between eyes as long, apex well rounded, very slightly, if at all,
angulate, slightly upturned apically; front very strongly inflated.
Pronotum nearly twice as long as vertex and very wide.
Color: Females uniformly bright green except for pink ocelli and
brown eyes; males straw yellow with brown stripe on middle of
vertex, disc and posterior half of pronotum and spots on suture of
elytra, brown.
Holotype, female, Hidalgo County, Texas, July 28, 1928,
R. H. Beamer. Allotype, male, Culberson County, Texas,
July 12, 1927, L. D. Anderson. Paratypes: Chaves County,
New Mexico; Hidalgo County, Texas; Nogales, Oracle, Pata-
gonia, and Navajo and Yavapai counties, Arizona.
Six paratypes in collection of California Academy of
Sciences ; holotype, allotype, and six paratypes in Snow ento-
mological collection.
Xerophloea inflata Lawson, n. sp.
Figs. 1, la
Closely related to X. robusta, but with shorter vertex and
even more inflated front. Length, female, 6 mm. ; male, 5 mm.
Head distinctly narrower than pronotum; vertex fully twice as
wide between eyes as long, anterior margin nearly rounded; front
very strongly inflated. Pronotum fully twice as long as vertex, very
wide.
Color: As in X. robusta.
Holotype, female. Brooks County, Texas, July 25, 1928,
L. D. Anderson. Allotype, male, Tucson, Arizona, June 21,
1924, A. A. Nichol. Paratypes: Mescal, Tucson, Oracle, Hua-
chuca, and Yavapai County, Arizona, and Palm Springs, Cali-
fornia.
Four paratypes in collection of California Academy of
Sciences ; holotype, allotype, and four paratypes in Snow ento-
mological collection.
APRIL, 1931]
LAWSON XEROPHLCEA
165
Xerophloea majesta Lawson, n. sp.
Figs. 9, 9a
Allied to X. major but not so robust and with angulate
vertex. Length, female, 7 to 8 mm. ; male, 6.5 to 7 mm.
Head distinctly narrower than pronotum; vertex nearly one-third
wider between eyes than long, margins very thin, apex obtusely
but distinctly angulate; front nearly straight. Pronotum distinctly
longer than vertex, quite wide.
Color: Females uniformly bright green except for pink ocelli and
brown eyes. Males straw yellow, usually well marked with brown
on middle line of vertex, brown spots all over pronotum, and elytral
veins alternating with white and brown spots.
Holoiype, female, Douglas County, Kansas, trap light, July
10, 1930, Paul B. Lawson. Allotype, male, same data except
taken on July 20. Paratypes: Douglas and Greenwood coun-
ties, Kansas; Hidalgo, Jim Wells, Palopinto, Bee, Kendall,
Cameron, and Brooks counties, Austin and Galveston, Texas;
Vicksburg, Mississippi; Clemson College, South Carolina.
Five paratypes in collection of California Academy of Sci-
ences ; holotype, allotype, and large series of paratypes in Snow
entomological collection.
Xerophloea oraclis Lawson, n. sp.
Figs. 10, 10a
Allied to X. minor but with more angulate vertex and more
strongly marked with dark brown. Length, male, 5.25 mm.
Head distinctly narrower than pronotum; vertex nearly one-half
wider between eyes than long, anterior margins straight, apex
strongly angulate; front slightly inflated. Pronotum one-half longer
than vertex, wide.
Color: Nearly ashy gray, strikingly marked with dark brown as
follows: On middle third of vertex and scattered spots on lateral
thirds, along anterior third to half of pronotum, and narrow band
along posterior margin, two large spots on scutellum, many small
spots on clavus, and particularly bordering elytral veins. Most of
face brown and entire underside with many small brown dots.
Holotype, male, Oracle, Arizona, July 28, 1924, E. P. Van
Duzee.
Type in collection of California Academy of Sciences.
Xerophloea tenuata Lawson, n. sp.
Figs. 7, 7a
Perhaps closest to X. viridis but much more slender and
vertex much longer. Length, male, 6 mm.
166 THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
Head slightly narrower than pronotum; vertex nearly as wide
between eyes as long, margins nearly straight to distinctly angulate
apex; front straight. Pronotum but slightly longer than vertex.
Color: Background straw yellow to creamy yellow, heavily
marked with brown. Vertex creamy yellow with three more or less
distinct brown longitudinal stripes. Pronotum creamy yellow,
irregularly irrorate with brown. Elytra straw yellow, with large
brown spots on claval suture and veins strongly spotted with brown.
Whole insect appearing strongly brown, above and below.
Holotype, male, San Diego County, California, October 17,
1913, E. P. Van Duzee.
Type in collection of California Academy of Sciences.
This species is by all odds the most slender member of the
genus.
Xerophlcea viridis (Fabricius)
Figs. 12, 12a
Cercopis viridis Fabricius, Ent. Syst., iv, p. 50, 1794.
Xerophlcea grisea Germar, Zeits. f. Ent., i, p. 190, 1839.
Xerophlcea virescens Stal, Of. Vet. Akad. Forh., xi, p. 253, 1854.
Parapholis peltata Uhler, Bui. U. S. Geol. Geog. Surv., iii, p. 461, 1877.
A comparatively slender species. Length, female, 6.5 to 7.5
mm.; male, 5.5 to 6.5 mm.
Head slightly narrower than pronotum; vertex about three-fifths
as long as width between the eyes, margins slightly convex to defi-
nitely angulate apex; front nearly straight. Pronotum at least one-
half longer than vertex.
Color: Females light green, often with minute black dots, especi-
ally along elytral veins. Males yellowish green, usually with median
line on vertex, disc and posterior half of pronotum, brown. In rare
cases males are gray, with additional brown markings on vertex and
pronotum and many brown spots on elytra, so that entire insect
appears distinctly brown.
This species has been recorded from a number of states
scattered over the whole area of the United States. In addi-
tion to those mentioned in Van Duzee’s catalogue, the writer
has before him specimens from Montana, Arizona, and New
Mexico.
In the light of the distribution of the new species here
described, there is some doubt in the writer’s mind as to
whether all the specimens called viridis, having as they do such
a wide range of distribution, are really but one species. More-
over, there seem to be some differences in the vertex and front
of specimens from various parts of the country. However, the
APRIL, 1931]
LAWSON XEROPHLCEA
167
writer has not been able to differentiate clearly enough the
specimens from the several regions, and thinks it best to put
under this name all the forms here included. In distribution,
therefore, this species may be said to cover the United States.
The original description of X. viridis does not set aside this
species from many of the others, and the writer has not seen
the type specimen. However, as now understood, the range of
this species extends to the type locality, the original description
fits it as well as any of the others, and commonly accepted
usage would seem to make it advisable to continue the name for
the forms described above. However, an examination of the
type, if available, may show that this name is applied to the
wrong species. In this case it is more than likely that the name
viridis should apply to the species which the writer has named
minor.
Germar undoubtedly had before him a male of this species
when he described grisea, and Stal was most liable to have had
a specimen of this, the commonest species, before him when he
described virescens. Uhler described peltata from Colorado.
This is undoubtedly a synonym for viridis because so far this
is the only species recorded from that state.
Dr. Myron H. Swenk of the University of Nebraska re-
cently told the writer that he had several records of the swarm-
ing in immense numbers of a species of Xerophloea. He was
kind enough to send the writer a number of specimens taken at
the time of swarming, and these proved to be viridis. It is
questionable whether anyone had ever before heard of any
species of Xerophloea occurring in anything like such tremen-
dous numbers and, therefore, we are, through the courtesy of
Dr. Swenk, quoting reports of a number of occasions when this
species occurred in great swarms :
From Neligh Leader (Nebraska), October 1, 1920:
“The 11th day of September was unique in that the city of Neligh
was visited by a cloud of the little green bugs, and windows other
than mine were clouded with them and houses and stores were
closed to shut them out. One of my friends set a small can under
the electric light at the intersection of Main and Putney streets
which was nearly filled by those which were killed and dropped. All
the stores were closed and it was quite a chore next morning to
sweep up the thousands that were caught when the stores were
closed.”
168
THE PAN-PACIFIC ENTOMOLOGIST
[vOL. VII, NO. 4
Z robitfU
10 mtlk
APRIL, 1931]
LAWSON XEROPHLCEA
169
Mr. H. E. Zimmerman, O’Neill, Nebraska, wrote on Sep-
tember 17, 1920:
“Please find enclosed envelope containing portion of bugs which
swarmed into our streets a few days ago. These pests swarmed in
and formed about two inches thick all over the sidewalks on the
west side of buildings. We are interested, as the farmers here are
inquisitive.”
Mr. John S. Marsh, Guide Rock, Nebraska, wrote on Sep-
tember 21, 1924:
“I am mailing you a sample of bugs that made their appearance
last night. One could have swept up a bushel of them in buildings
and on the sidewalks. What are they?”
It 'is evident from the above, that on occasion at least, this
species must be considered as one of the more important leaf-
hoppers.
Xerophlcea major Baker
Figs. 13, 13a
Xerophlcea major Baker, Psyche, viii, p. 285, 1898.
A large stout species with rounded vertex. Length, 7.5 to
8 mm.
Head slightly narrower than pronotum; vertex two-thirds to
three-fourths as long as width between eyes, anterior margin broadly
rounded; front in profile nearly straight. Pronotum but slightly
longer than vertex.
Color: Female bright green, frequently well spotted with minute
black dots.
This species is unique in its rounded vertex and general
robustness. Van Duzee in his catalogue lists it from New
York, New Jersey, Virginia, and Tennessee. The writer has
before him a specimen from Cherokee County, Kansas.
The Odonata of China
Students of the Odonata will be interested in Dr. James G.
Needham’s Manual of the Dragonflies of China, published in
Peiping as fasc. I of Vol. II of Zoologica Sinica, Ser. A., issued
October 1, 1930. Keys are given to the higher taxonomic units
as well as to the genera and species, and adequate descriptions
are given for all. Full accounts of the preparatory stages are
included when they are known, and notes on distribution. A
total of 89 genera and 266 species are treated in the 344 quarto
pages and 20 plates. The work is entirely in the English lan-
guage. Apparently the Odonata are a dominant group in China.
— E. P. Van Duzee.
170
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII, NO. 4
SYNONYMS IN THE NORTH AMERICAN
THYSANOPTERA
BY J. DOUGLAS HOOD
University of Rochester
It has seemed desirable to clarify the records of this group
of insects by eliminating certain synonyms. Three of these
have been known to the writer for some years.
Genus Leucothrips Reuter
1904, Leucothrips Reuter, Medd. Soc. Fauna Flora Fenn., 30: 107.
[Type, L. nigripennis, n. sp., by monotypy.]
1913, Microthrips Morgan, Proc. U. S. Nat. Mus., 46: 19. [Type,
M. piercei, n. sp., by designation and monotypy.]
Body glabrous, without reticulation or close pubescence. Eyes
prominent, strongly protruding, pilose. Antennae seven-segmented,
the last segment long, slender, about equal in length to sixth. Pro-
thorax with two pairs of moderately long bristles at posterior angles.
Fore wings long, slender, with a single longitudinal vein which
bears a few bristles at base and one or more at apex. Abdominal
bristles short and weak.
The affinities of this genus are with Sericothrips, Scirto-
thrips, and Drepanothrips, but from all of them it may easily
be known by the seven-segmented antennae and glabrous abdo-
men. Reuter described the antennae as eight-segmented; but
many years ago he sent me two specimens from his type series
of nigripennis, mounted on small pieces of cardboard (as was
his habit) ; and these have been compared with material from
Belgium collected and determined by Bagnall. Reuter’s mis-
observation was due to the dried condition of the material and
its minute size, and led to the redescription of the genus by
Morgan in 1913 as Microthrips.
Leucothrips nigripennis Reuter
1904, Leucothrips nigripennis Reuter, Medd. Soc. Fauna Flora Fenn.,
30: 108 [Helsingfors, Finland, on ferns in greenhouse]. 1909,
Bagnall, Ann. Soc. Ent. Belg., 53: 172 [Brussels, Belgium, on
fern in greenhouse]. 1909, Bagnall, Journ. Econ. Biol., 4: 38
[Newcastle-on-Tyne, England, and Glasgow, Scotland, on ferns
and other plants in greenhouses]. 1926, Pettit, 64th Ann. Rep.
Seer. State Board Agr., State of Michigan, p. 219 [East Lansing,
Mich., on fern in greenhouse]. 1926, Priesner, Thys. Eur., pp.
266, 718.1
1 On this latter page Priesner says (translation) : “According to Hood,
in letter, this genus has 7- not 8-segmented antennse. A fuller descrip-
tion must be reserved for some later time, as I am at present not in
possession of actual specimens.”
APRIL, 1931] HOOD NORTH AMERICAN THYSANOPTERA 171
1927, Microthrips leucus Herrick, Ent. News, 38: 278. Fig. 2 [East
Lansing, Michigan, on fern in greenhouse].
This species is represented in the material before me by
cotypic material from Finland, sent to me twenty- two years
ago by Reuter himself, by two specimens from Brussels, Bel-
gium, taken in a greenhouse October 8, 1908, on Duvallia, by
Bagnall, and two specimens from the series which furnished
Mr. Herrick’s types, taken on a fern in a greenhouse at East
Lansing, Michigan, by Donald Ries and Miss E. 1. McDaniel.
One of these specimens is a cotype.
Herrick’s specimens were studied by me in 1925 and 1926,
at which times I told him that the species was Leucothrips
nigripennis.
The insect, as will be seen from the citations given above,
appears to be not rare in greenhouses in both the Old and
New Worlds.
Thrips herricki Bagnall
1926, Thrips herricki Bagnall, Ann. and Mag. Nat. Hist., Ser. 9, 18:
545 [Ithaca, N, Y., June 27, 1924, G. W. Herrick, on Veratrum
wiride'\.
1927, Thrips veratri Priesner, in letters [Ithaca, N. Y., June 27, 1924,
G. W. Herrick, on Veratrum viride],
1927, Thrips veratri Hood, Ent. Amer., 7: 218. PI. XX, fig. 2 (June
14) [Ithaca, N. Y., June 27, 1924 (G. W. Herrick) and July,
1926 (J. D. H.), on Veratrum viride].
1927, Thrips veratri Herrick, Ent. News, 38: 276. Fig. 1 (October 31)
[Ithaca, N. Y., June 27, 1924, G. W. Herrick, on Veratrum
viride ] .
Mr. Herrick discovered this species at Ithaca, New York,
on False Hellebore (Veratrum viride Ait.) and sent specimens
for determination simultaneously to Bagnall, Priesner, and
myself, each of whom, exercising the right of a specialist to
publish the results of his studies, prepared descriptions for the
press. Two of these descriptions actually appeared. Upon
receiving the results of his poll, Mr. Herrick drew up a fourth
description himself, and published it. No confusion has re-
sulted from this wasted effort, however, three of the four
authors, in naming the insect after the plant on which it was
taken, having employed the same specific name !
Wherever its food plant occurs, in New York State at least,
this much-described insect may usually be found in June and
172
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
July, feeding, often in abundance, on the broad leaves, usually
in company with its nymphs.
Treherniella amplipennis (Morgan)
1913, Trichothrips amplipennis Morgan, Proc. U. S. Nat. Mus., 46: 33.
Figs. 64-67. [Quincy, Florida, on Hypericum dolabriforme],
1919, Haploihrips orlando Watson and Osborn, Fla. Buggist, 2: 116.
[Orlando, Florida, taken by sweeping shrubs in “flat woods.’^]
1923, Treherniella orlando, Watson, Fla. Agr. Exp. Sta. Bui., 168: 67,
81. [Designated as type of the new genus Treherniella\,
The combination of generic knd specific name given above
is new. The following material has been studied in arriving at
the above synonymy :
Florida: Quincy, May 15, 1910, on Hypericum dolabriforme, A. C.
Morgan and G. A. Runner; 2 $ $ (2 slides), paratypes of Tricho-
thrips amplipennis.
Gainesville, August 9, 1919, oak in “flat woods”; 1 paratype of
Haploihrips orlando.
Las Palmas, Merritts I., May 4, 1924, sweeping, J. R. Watson;
1 determined by Watson as Treherniella orlando.
Tampa, February, 1924, T. H. Hubbell; 1 $, determined by Wat-
son as Treherniella orlando.
Bradentown, January 20, 1919, sweeping, Alexander Wetmore;
S $ $, 1
Megalothrips picticornis Hood
1927, Megalothrips picticornis Hood, Proc. Biol. Soc. Wash., 40: 204
[Blue Lake, Calif., and Salt Lake City, Utah, on dead willow].
1929, Docessissophothrips animus Moulton, Bull. Brooklyn Ent. Soc.,
24: 242 [Mt. View, Calif., and Corvallis, Ore.] 1931, Pan.-Pac.
Ent., 7: 123 [Berkeley, Calif., in scolytid burrows in linden].
There can be little question as to the correctness of the
above synonymy, though Moulton’s specimens apparently lack
the yellow band which I have described as occurring at the
base of the fourth antennal segment. The species is thoroughly
congeneric with the European M. bonannii Uzel (1895) and
the Eastern North American M. spinosus Hood (1908), both
of which have been fully described and figured. Like them,
picticornis is perhaps most frequently found in hollow stems
and in the burrows of various wood-boring beetles. Whether
it is specifically distinct, or later will be found merely entitled
to subspecific rank as a geographical race, further collecting
no doubt will disclose.
APRIL, 1931] MOULTON — A NEW THRIPS
173
AN INTERESTING NEW CALIFORNIA THRIPS
BY DUDLEY MOULTON
Mr. Richard S. B agnail, in his interesting paper on the
Classification of the Thysanoptera, (Ann. & Mag. Nat. Hist.,
Ser. 10, Vol. 5, p. 574, May, 1930), erects a new Superfamily,
Melanothripoidea, for the Family Melanothripidce, Bagnall.
One of the characters included in this Superfamily is “Apex
of fore tibia armed with a dagger-like scoop, or, if simple,
antennal joint one or two so armed.” Other characters of the
included genera are three- j ointed maxillary and two-jointed
labial palpi, single or horseshoe- shaped sense area on each of
third and fourth antennal segments and the absence of a tarsal
hook.
The genus Dactuliothrips with a maxillary palpus of three
and a labial palpus of two segments cannot be included in the
Superfamily Aeolothripoidea Hood, Family Orothripidcs Bag-
nall, although it has many characters in common with this
group. Also it cannot be included in the Family M elanothripidcs
Bagnall, nor in the Superfamily Melanothripoidea Bagnall,
as now recognized although the predominating characters
would seem to place it in this superfamily. I am therefore
erecting a new family, Dactuliothripidce, and genus and placing
them as follows :
Superfamily Melanothripoidea Bagnall
Family Dactuliothripidae Moulton
Genus Dactuliothrips Moulton
(Daktulios — 3 . ring)
Having the general appearance of an Orothrips or Melanothrips.
Antennae long and slender, with all segments freely movable and
reduced gradually beyond the second. Segments three to nine each
with nine to eleven distinct annulations or rings which are sur-
rounded by whorls of setae. Each of segments three and four with
two circular to oblong sense areas near tip. Maxillary palpus with
three segments and geniculate between second and third, labial pal-
pus with two segments. Fore tibia unarmed, each fore tarsus with
a distinct hook-like appendage. Wings fully developed, broadly
rounded at tip, with ring vein, two longitudinal and four cross veins.
Ovipositor curved upward.
The genus Dactuliothrips may be separated from Orothrips
by the shape of the antennae and the three-segmented maxillary
174
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
palpus and two-segmented labial palpus. It may be separated
from Melanothrips by the presence of two circular sense areas
on segments three and four and the presence of a tarsal hook-
like appendage.
Dactuliothrips spinosus Moulton, n. sp.
Female holotype: Color uniformly brown to light brown with
fore tibiae and tarsi and third antennal segment yellowish. Wings
uniformly light grayish brown, without darkened cross bands. Body
and wing spines dark brown.
Total body length, 1.44 mm.; head, length .176 mm., width .205
mm.; prothorax, length .147 mm., width .235 mm.; mesothorax,
width .32 mm.; abdomen, width .47 mm. Segments of antenna:
length (width) I, 23 (36); II, 40 (30); III, 56 (23); IV. 66 (21);
V, 63 (18); VI, 63 (17); VII, 50 (16); VIII, 46 (13); IX, 53 (10);
total length of antenna 456 microns. Length of spines: interocellar
73 microns, postoculars, outer and inner, 80 microns, median dorsal
on ninth abdominal segment 133 microns, on tenth segment 140
microns.
Head slightly longer than wide, depressed in front at base of
antenna, cheeks slightly arched. Interocellar, postoculars, and other
spines prominent. Eyes produced in front, ocelli fully developed.
Mouth cone short. Antenna slender, 2.6 times longer than head.
Prothorax with numerous conspicuous spines, especially those at
angles and margins. Legs with numerous conspicuous spines.
Fore femora thickened and armed with a series of four short spurs
on distal half of inner anterior margin. Fore tibia without arma-
ture except a pair of stout spines at tip. Wings with regularly
placed spines on costa, sixteen on fore vein and thirteen on hind
vein.
Abdomen broadly ovate with tergites transversely striate and
more or less densely pubescent at the sides. Terminal segments
small.
Male allotype: Smaller than female, body length .97 mm. Body,
legs, and wings lighter in color than in female, but with all body,
leg, and wing spines conspicuously prominent by their dark color.
The single male specimen has an armature of only three spurs on
inner distal half of each fore femora instead of four as in the female.
Type material: Female holotype, male allotype, six female
paratypes taken on Prunus demissa (Choke-cherry), May 19,
1929 (G. R. Struble). All types in author’s collection. (Moul-
ton No. 3832.)
Type locality: Willow Ranch, Siskiyou County, California.
APRIL, 1931] BAILEY NEW CALIFORNIA THRIPS
175
A THRIPS NEW TO CALIFORNIA
BY STANLEY F. BAILEY
University of California
On September 2, 1930, Mr. A. E. Michelbacher collected
some very unusual thrips from the bark of a dying coast live
oak {Quercus agrifolia Nee). The tree was situated on the
southwest corner of the campus of the University of California,
Berkeley, California. Specimens of this very interesting thrips
sent to Mr. Dudley Moulton of San Francisco were determined
as Pcecilothrips ornatus (Hood). He stated that this was the
first record from California of this rather uncommon thrips.
This species was first described by J. D. Hood in 1913 as
Phlosothrips ornatus from thirty-seven females taken in May
and June on the bark of a dying red oak tree at Washington,
D. C.
Hood stated that there were both winged and wingless
forms and that they were practically identical save for the
presence or absence of the wings. No mention was made of the
male and apparently it never has been collected.
There seems to be no further mention of this species until
1927 when Hood transferred it to the genus Pcecilothrips and
wrote “I have for some years suspected that my Phlceothrips
ornatus might be congeneric, perhaps even conspecific, with
Uzel’s Pcecilothrips albopictus, described and still known only
from the unique type taken in Bohemia. Dr. Priesner (in litt.)
has suggested the same thing, but he has not yet been able to
make a direct comparison.’’
At the same time Hood (1927) reported this species from
New York and Pennsylvania. However, of all the collections
made of this thrips in this country up to September, 1930, the
total number of specimens was represented by forty adult
females, one pupa and seven larvae. The colony here recorded
on a dying coast live oak in Berkeley was represented by several
hundred larvae and adults, both winged and wingless. Of 201
adult females collected on September 3, 1930, twenty-three, or
11.4 per cent, were winged. No males or pupae were found.
Large numbers of eggs, discernible as white specks, were to
be found singly and in clusters on the surface and among the
cracks of the bark. The long axis of the egg was in some cases
176 THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
at right angles to the surface of the bark and in other cases
parallel. The clusters of eggs exhibited no order of arrange-
ment.
From careful observations made on this colony during the
fall and winter it appeared that this species, like many of the
suborder Tubulifera, were inhabitants of bark. My observa-
tions seem to be substantiated in that collections of this species
were made in Washington, D. C, from the bark of a dying
red oak, and that the specimens collected in Pennsylvania and
New York were taken under the bark of hickory stumps (two
females were taken flying). It was not definitely ascertained
whether the thrips were feeding only on the exudate of the
dying tree, which was issuing from the deeply cracked bark,
or on fungi, although no fungi were visible under a hand lens.
The adults and larvae were found feeding together in scattered
groups, both on the surface of the bark, and particularly on the
freshly cracked portions of the trunk, from the surface of the
ground to about five feet above. All forms present were very
sluggish in their movements and did not even attempt to seek
shelter under the loose bark when disturbed. Neither egg-
laying, nor emergence of larvae was observed, nor was the
number of larval molts or the rate of egg laying determined.
In association with the thrips were various spiders, many mites,
ants, psocids, and collembolans. From all observations the
mites present were not predacious on the thrips.
During September and October both young and mature
larvae were present, as well as a few winged adults. But as the
season progressed the numbers fell off greatly and by early
December only about ninety large larvae and twenty-five wing-
less adults could be found. No eggs were found after the first
of November. These remaining individuals had concentrated
on the north side of the trunk, which was still exuding a small
amount of sap. The bark on the opposite side of the tree had
dried out and was shrinking away from the trunk. There was
an unusually cold snap during the last of December and during
this period the thrips crawled deep into the crevices of the bark
and under portions that had pulled away from the trunk. All
activity was at a minimum. Throughout the following month
very little activity was evident and the numbers began to drop
APRIL, 1931] BAILEY NEW CALIFORNIA THRIPS
177
off more noticeably until February 1, when only seven wingless
adults could be found and only twenty-seven large larvae. These
surviving individuals were clustered on the north side of the
tree about four inches from the ground on a moist section of
bark partly protected by a clump of dead grass. From this it
would seem that both mature larvae and wingless adults wintered
over and that there was a very great mortality Under such con-
ditions. No other thrips of this species have been found in the
locality. On February 11 the tree was removed, as it was
becoming unsightly and observations on the biology of this
isolated colony were concluded.
Description of the Egg
The egg is quite delicate, oblong, rounded at both ends with the
sides practically parallel. Average length 0.447 mm., average width
0.185 mm. Color snowy white due to a covering of waxy material
over the entire surface. This waxy substance is laid down in reg-
ular hexagonal areas the average width of which is 0.026 mm. On
the anterior end the wax is sometimes present in raised areas, giving
the appearance of tubercles. Previous to hatching the yellowish
tinge of the larva is visible under the wax and, with magnification,
the reddish brown eye-spots are evident, as well as the legs and
antennas which are folded down on the ventral surface of the body.
Upon hatching the anterior end of the chorion is apparently forcibly
ruptured by the increased size of the mature embryo.
Description of the Larva
Newly emerged larva. Total length 0.537 mm. Abdomen ten-
segmented, tapering; length 0.184 mm.; length of tube or tenth seg-
ment 0.066 mm.; ninth segment about one-half as long as tenth;
remainder of segments shorter and about equal. Width of first
abdominal segment 0.123 mm.; width of tenth segment at base
0.034 mm. Head very long in proportion to body, as are the
antennas and legs. Length of head 0.114 mm.; width 0.066 mm.
Width of mesothorax, 0.127 mm. Antenna seven-segmented; length
0.211 mm. Length of antennal segments: I 0.022 mm., II 0.028 mm..
Ill 0.039 mm., IV 0.035 mm., V 0.026 mm., VI 0.026 mm., VII 0.035
mm. First segment short, cylindrical; second, barrel-shaped; third,
fourth, fifth and sixth segments narrower at base; seventh long,
tapering. Color gray; anterior half of second segment very light,
remainder of segments a uniform gray. The eyes are reddish brown
and, in general, the color of the larva is pale yellow with a trace
of crimson on the margins of the abdominal segments. No ocelli.
Spines on head, thorax and abdomen branched at tip.
Full grown larva. The mature larva is very strikingly marked
with crimson and white. There is considerable variation in the size
178
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
and position of the white areas but from a dorsal view the general
appearance is as follows: head white with lateral margins and two
longitudinal stripes in center crimson; prothorax white with two
irregular areas of crimson in the center; mesothorax crimson, often
with the lateral margins white; metathorax white. Abdomen: first
and second segments crimson, often with circular area of white
in center; third segment white; fourth crimson with an irregular
white spot in center; fifth crimson; sixth white, often with anterior-
lateral margins crimson; seventh, eighth and ninth crimson with
large irregular spots of white in center; tenth reddish brown. Legs
yellowish, tinged with crimson. Eyes dark reddish brown. Ocelli
and genitalia not apparent. First three abdominal segments equal
in width, the remainder tapering to the tube or last segment.
Head slightly wider at base than below eyes. Spines on head and at
posterior angles of thoracic and abdominal segments strong,
branched at tip.
Total length 1.550 mm. Length of head 0.150 mm., abdomen 0.875
mm., tenth abdominal segment 0.075 mm., ninth slightly less than
tenth. Width of head at base 0.139 mm., prothorax 0.328 mm., meso-
thorax 0.417 mm., metathorax 0.437 mm., first abdominal segment
0.398 mm., tenth abdominal segment at base 0.034 mm.
Length of antenna 0.288 mm. Length of antennal segments:
I 0.019 mm., II 0.039 mm.. Ill 0.069 mm., IV 0.049 mm., V 0.039
mm., VI 0.029 mm., VII 0.023 mm. Color of antenna grayish brown,
first, second and third segments slightly tinged with crimson, first
and base of second very light gray. First segment short cylindrical,
second barrel-shaped, third sub-conical, fourth, fifth and sixth also
sub-conical but more abruptly so, seventh short, tapering, bluntly
pointed.
Descriptions of egg and larvae made from a large number of
specimens collected by the author on the bark of a dying coast
live oak, Berkeley, California, September 3, 1930.
Bibliography
Hood, J. D., Proc. Biol. Soc. Wash., Vol. XXVI, pages 161-166, 1913.
Entomologica Americana, Vol. VII (new series). No. 4, pages
234-235, 1927.
Moulton, D., Pan-Pacific Entomologist, Vol. VII, page 123, 1931.
APRIL, 1931] FENDER YAMHILL COUNTY BUTTERFLIES 179
BUTTERFLIES OF YAMHILL COUNTY, OREGON
BY KENNETH FENDER
McMinn<ville, Oregon
Yamhill County is in the northwestern portion of the State
of Oregon, just south and west of Portland. It has proved a
very fair collecting area for rhopalocera and I thought a list of
the captures over a recent period would be of interest. Some
ninety names are herein recorded, including two newer names
from recent descriptions.
I wish to thank Mr. Robert Rieder and Mr. Ralph Macy,
two local collectors, for their help. Without their notes this list
would be very incomplete.
I have adopted Mr. J. D. Gunder’s style of cataloguing
from his “List of Butterflies of Los Angeles County, Calif.”
(Bulletin So. Calif. Acad. Sci., XXIX, August, 1930), and
believe his use of classification categories, including the use of
transition form in place of the word aberration, to be very
satisfactory. His list quotes original description references,
illustration notices and food plant data, so this information
would, I believe, be superfluous if repeated here.
As this list probably is incomplete, I will record new cap-
tures as they occur, making note of them in future issues of
the Pan-Pacific Entomologist.
1. Papilio bairdii oregonia Edw., a race (or form?).
1 ^ , no date, Carlton. Only specimen thus far reported from
this county.
2. Papilio zelicaon Luc., a species. 1 $ , 1 $ , May 18,
1930, Baker Creek Canyon; 1 $ , April 27, 192'8, McMinnville
1 ^ , May 6, 1928, McMinnville. Well distributed, but seem-
ingly uncommon.
3. Papilio indra Reak., a species. Although this is an
indra territory, I cannot report its capture from this county, but
undoubtedly it is here, as it has been seen.
4. Papilio rutulus Luc., a species. 5 5,3 9 , April 16,
1928, Baker Creek Canyon; 1 5,1 9, May 23, 1928, Muddy
Valley; 9 5,3 9 , July 24, 1930. Common everywhere in the
county and an easy capture.
180
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
5. Papilio eurymedon Luc., a species. 10 5,3 $ , June
16, 1930, Baker Creek Valley; 3 5, May 9, 1930, Muddy
Valley; 9 5,1 9, June 6, 1929, Baker Creek Valley. A
common Papilio, taken at all elevations.
6. Papilio eurymedon albanus F. & F., a form. 1 5 ,
June 6, 1928, mountains above Baker Creek. A seemingly rare
form of this species.
7. Parnassius clodius claudianus Stich., a race. 50 5 ,
10 9 , June 25, 1930, and 20 5 , 7 9 , June 30, 1930, Baker
Creek Canyon. Very common in this particular canyon where
on favorable days it fairly seems to swarm.
8. Parnassius clodius baldur Edw., a form? 2 5,3 9 ,
August 8, 1928, 1 5 , July 12, 1930, and 1 5 , July 16, 1930,
Baker Creek Canyon. Not very plentiful.
9. Neophasia menapia F. & F., a species. 1 5,1 9 , July
27, 1930, 12 5,2 9 , August 6, 1930, 7 5,4 9 , August 12,
1929, Baker Creek Canyon; 1 5 , July 27, 1929, Pea Vine Ridge,
Females difficult to capture, as they fly high.
10. Ascia protodice Bdv. & Lee., a species. 1 9 , August
23, 1930, McMinnville; 1 9, unknown date, Bellview. Prob-
ably a visitor from eastern Oregon.
11. Ascia protodice vernalis Edw. 1 9, August 27,
1930, Bellview. Also a stray.
12. Ascia napi oleracea Harr., a form? This butterfly
is seen on the western boundary of the county.
13. Ascia napi castoria Reak., a seasonal form? 1 9,
August 27, 1930, Bellview. Seldom taken in this county.
14. Ascia napi margin alis Scud., a seasonal form? 3 5,
April 20, and 20 5 9 s, May 3, 1930, Baker Creek Canyon.
Rather common at proper time.
15. Ascia napi pallida Scud., a seasonal form? 3-5,
2 $ , June 18, 1928, 5 5 , June 30, 1929, 2 5,1 9 , June 6,
1930, Baker Creek Canyon. A fragile butterfly hard to secure
in good condition. Females seem to be scarce.
16. Ascia rap^ L., a species. 12 5,9 9 , May 12, 1927.
McMinnville. Common everywhere as usual.
17. Ascia rap^ yreka Reak., a form. 2 5, 1 9 , July
29, 1930, McMinnville. A darker form of the species, espe-
cially dark on the under side.
/ndex
(J)Afc Minnvilte
® Upper Baker Creek "Valley
^ Louver Baher Creek Valley
® Muddy Valley
® Peavine Ridge
®Macy’s farm
® Amity Hills
®3o/a Hills
® Grand Island
® Coast Range
® Field South of McMinnville
® Yamhill County Golf Course
©Grand Roude Hills
MaPcF COLUCTING GROUNDS in YaMHILLCoUNTY, OREGON-Fender
00
APRIL, 1931] FENDER — YAMHILL COUNTY BUTTERFLIES
182
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
18. Anthocharis SARA REAKiRTi Edw., a racc. 12 S ,
April 10, 1929, 6 S , May 16, 1930, Baker Creek Canyon. A
common butterfly. White females not seen.
19. Anthocharis sara reakirti stella Edw., a female
form. 3 $ , April 16, 1927, Baker Creek Canyon ; 5 $ , April
29, 1929, hills north of Bellview; 4 $, May 13, 1930, Baker
Creek Canyon. Certainly a female form of the species in this
locality.
20. Anthocharis sara julia Edw., a race here ? 1 5 ,
May 16, 1930, Baker Creek Canyon; 1 May 27, 1930, Pea
Vine Ridge. Identification uncertain but probable.
21. Eurymus eurytheme Bdv., a species. 1 $ , October
12, 1929, McMinnville. Not common.
22. Eurymus eurytheme autumnalis Ckll., a seasonal
form (?). 1 $, May, 1927, Bellview; 1 $, October 19, 1928,
McMinnville. Divergent emergence may account for the above
records.
23. Eurymus eurytheme amphidusa Bdv., may be a spe-
cies here. 2 S $, August 6, 1927, south of McMinnville;
1 1 9, September 6, 1927, golf course near McMinnville;
5 B , 2 9 , September 6, 1929, near McMinnville. What I take
to be this butterfly is quite common in the county.
24. Eurymus eurytheme amphidusa alba Stkr., a female
form. 2 9 , August 16, 1928, Muddy Valley; 1 9 , October 12,
1930, McMinnville. White females occur continually.
25. Eurymus occidentalis chrysomelas shast^ B. and
Benj., a female form. 1 9 , date unknown, Bellview. The white
female of chrysomelas and rare.
26. Danaus menippe Hbn., a species. 2 S , June 6, 1927,
McMinnville; 1 ^ , August 7, 1930, Baker Creek Valley. Quite
scarce in this county because of absence of the food plant,
which is milkweed.
27. CcENONYMPHA CALIFORNIA ERYNGii Hy. Edw., a race.
1 5 , July 7, 1928, and 1 July 23, 1930, McMinnville. Has
more yellow on than the species.
28. CcENONYMPHA AMPELOS Edw., a species. 100 S $ ,
1927, 25 ^ 9 , July 19, 1929, McMinnville. A very common
butterfly.
APRIL, 1931] FENDER YAMHILL COUNTY BUTTERFLIES 183
29. Cercyonis ariane Bdv., a species. 40 ^ , June 12,
1930; 20 5 , 60 $ , June 19, 1930, Pratt’s Farm. The only
Satyr in the county thus far.
30. Argynnis leto Behr, a species. 3^,2$, July 9,
1929, 10 S, July 12, 1930, 25 $, 1 $, July 15, 1930, Baker
Creek Valley. Most plentiful Argynnis in this section; but
females are difficult to secure.
31. Argynnis bremnerii Edw., a species, 6^ $ , July 14,
1929, 12 5 $ , July 12, 1930, Baker Creek Valley. Common
in locality noted.
32. Argynnis hydaspe rhodope Edw., a race. 2 $ , 1 $ ,
July 12, 1930, 1 $ , July 16, 1930, Baker Creek Valley. Not
plentiful.
33. Argynnis clio Edw. (?) 1 ^ , June 9, 1928, Muddy
Valley. I hesitate to try to identify this specimen until Mr.
Gunder’s revision of the genus appears.
34. Brenthis epithore Edw., a species. 25 $ ,7 $ , May
17, 1930, 16 $ , 3 2 , June 4, 1930, Baker Creek Valley.
Common from May till late July.
35. Euphydryas chalcedona D. and H., a species. 2 $ ,
May 26, 1930, Baker Creek Valley; 1 2 , May 23, 1928, Lower
Baker Creek. Seldom found in this district.
36. Euphydryas chalcedona colon Edw. ( ?) , a race.
1 $ , May 30, 1929, Baker Creek Valley; 2 $, June 4, 1930,
Lower Baker Creek Valley. More local exploration is needed
to determine these specimens with certainty.
37. Phyciodes campestris Behr., a species, 5 $ , 3 2,
May 16, 1928, Muddy Valley; 3 $, August 6, 1930, Baker
Creek Valley. Common in May and June.
38. Phyciodes mylitta Edw., a species. 12 $ , May 8,
1930, Baker Creek Valley; 1 ^ , 1 2 , July 16, 1930, McMinn-
ville. Plentiful throughout the county.
39. Phyciodes mylitta macyi Fender., a transition form.
1 $ , September 6, 1929, Pea Vine Ridge. (Ent. News, June,
1930). Albifusism.
40. Phyciodes mylitta collinsi Gund., a transition form.
1 3 , July 4, 1929, McMinnville. Immaculism.
41. PoLYGONiA SATYRUS Edw,, a species. 7 S 9 , April 25,
1928, 15 ^ 2, May 9, 1929, McMinnville; 6^,1 2, July 16,
184
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
1930, Baker Creek Valley. Common enough in the valleys at
the forest edge.
42. Polygon lA satyrus chrysoptera Wrgt., a form. 1 S ,
2 $ , undated, McMinnville. A lightly colored form of this
species.
43. PoLYGONiA SATYRUS MARSYAS Edw., a race (?). 2 $ ,
April 25, 1928, 2 1 $ , April 27, 1928, McMinnville. A
scarce Satyr.
44. PoLYGONiA FAUNUS RUSTicus Edw., a race. 15^9,
April 12, 1930, 5 ^9, May 4, 1930, Baker Creek Valley.
Many can be captured in the valleys of the foothills.
45. PoLYGONiA OREAs Edw., a species. 1 $ , April 27, 1930,
Baker Creek Valley. This is the only record thus far.
46. PoLYGONiA OREAS siLENUs Edw., a racc. 3 $ , 1 9 ,
May 20, 1928, 2 ^ , May 29, 1929, Muddy Valley; 1^,29,
April 12, 1930, Baker Creek Valley. Considered a rare butter-
fly and a high flier.
47. Aglais californica Bdv., a species. 1 $ , May 16,
1929, Baker Creek Valley; also 1 $ taken near McMinnville.
Strangely rare in this collecting area.
48. Aglais milberti Godt., a species. 1 $ , June 7, 1928,
McMinnville. Rare here.
49. Aglais milberti subpallida Ckll., a form. 1 $ , July
12, 1929, Yamhill County golf course; 2 5, June 16, 1930,
Baker Creek Valley. This form is evidently commoner than
the species itself. Much intergradation.
50. Aglais antiopa L., a species. 4^,1 9 , April 8, 1930,
1 $ , July 16, 1929, Baker Creek Valley. This species seems
to be with us the year round*
51. Cynthia atalanta L., a species. 2 $ , May 12, 1928,
3 $ (bred), June 14, 1929, McMinnville. Found most every-
where.
52. Cynthia virginiensis Dru., a species. 1 $ , May 24,
1929, McMinnville. Not at all common here.
53. Cynthia cardui L., a species. 1 ^ , May 16, 1928,
1 $ , May 24, 1930, Muddy Valley. Scarcely ever taken in the
county, but may be plentiful at times.
54. Cynthia carye Hbn., a species. 4^9, April 25, 1928,
6 (J 9 , April 27, 1928, McMinnville. Common in comparison
APRIL, 1931] FENDER YAMHILL COUNTY BUTTERFLIES 185
to cardui. This species likes to play along the edges of the
woods with the Grapta species.
55. Cynthia carye muelleri Letch., a transition form.
1 $ , July 23, 1930, Macy’s Farm. This specimen was picked
off the front of an auto.
56. Basilarchia weidemeyerii Edw., a species. 1 $ , un-
dated, foothills south of McMinnville. Probably a visitor in
this region.
57. Basilarchia lorquini Bdv., a species. 5 $ , July 24,
1930, Baker Creek Valley; 2 $, 1 $, May 9, 1928, Muddy
Valley. Our commonest Admiral.
58. Basilarchia lorquini eavesii Edw., a form. 1 $ ,
August 6, 1930, McMinnville. A form in which a row of spots
appear on the upper side of secondaries.
59. Basilarchia lorquini burrisoni Mayn., a race (?).
10 $ , 3 $ , May 24, 1930, Baker Creek Valley. There is evi-
dently a local colony of this up Baker Creek.
60. Heterochroa bredowii californica Butl., a race.
1 July 29, 1928, McMinnville; 2 $, 1$, August 6, 1930,
Baker Creek Valley. Quite rarely found.
61. Atlides HALESus Cram., a species. 1^,1 9 , undated,
Macy’s Farm near McMinnville. Considered a very lucky catch
in this region.
62. Strymon melinus Hbn., a species. In local collec-
tions, evidently from this district.
63. Strymon sylvinus Bdv., a species. 9 $ 2 , May 18,
1930, 6 $ 2 , June 14, 1929, Baker Creek Valley. Taken often
in foothills and higher levels.
64. Incisalia iroides Bdv., a species. 1 $ , July 12, 1930,
1 9 , July 16, 1930, Baker Creek Valley. Last year I was glad
to find this butterfly for the first time.
65. Incisalia eryphon Bdy., a species. 1 3 , July 26, 1930,
Baker Creek Canyon. Probably more records of this species
will be available later.
66. Callophrys dumetorum Bdv., a species. 2 3 , June 8,
1930, 1 3 , June 27, 1930, Valley of Baker Creek. Probably
common enough if searched for.
67. Tharsalea virginiensis Edw., a species. S3, 2 9,
June 18, and 2 3 , I 2 , June 27, 1929, Lower Baker Creek
186
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
Valley. A small local habitat of this species has thus far been
the only colony found in this region.
68. Lyc^ena xanthoides Bdv., a species. 1 ^ , 1 $ , June
18, 1928, 1 $ , June 27, 1928, Macy’s Farm near McMinnville.
The northern limit of this species evidently just reaches this
county.
69. Lyc^na helloides Bdv., a species. 3 ^ , 3 9 , June 8,
1928, 7^,39, June 21, 1929, McMinnville; 2^,49, July 12,
1930, Baker Creek Canyon. This pretty little copper is our
commonest butterfly in the genus.
70. Everes amyntula Bdv., a species. 4 5,2 9 , April 8,
1929, 2 5,29, May 18, 1930, Baker Creek Valley and Canyon.
Fairly plentiful.
71. Plebeius MELISSA Edw., a species. 1 5 , June 7, 1930,
fields south of McMinnville. Probably only a chance record,
though there may be a numerous colony.
72. Plebeius maricopa fenderi Macy, a race. 1 5,2 9 ,
May 25, 1929, taken on an isolated patch of lupine just west of
Macy’s Farm. (Ent. News, January, 1931).
73. Plebeius acmon D. and H,, a species. 6 5,29, May
18, 1929, Redmond’s rock crusher; 10 5,6 9, June 11, 1927,
Grand Island. Common enough most everywhere.
74. Plebeius acmon cottlei Grin., a seasonal form. 1 5 ,
1 9 , April 12, 1928, at same locality as species. Has darker
underside. Is an interesting catch.
75. Philotes battoides oregonensis B. and McD., a race.
1 5 , July 27, 1927, Muddy Valley. Only record thus far.
76. Glaucopsyche xerces Polyphemus Bdv., a race (?).
4 5,3 9 , May 16, 1930, 2 5,1 9 , June 7, 1930, Baker Creek
Valley. The synonymy of Barnes and Benjamin check list is
difficult to understand in this group.
77. Glaucopsyche xerces antiacis Bdv., a form or race
(?). 15,29, May 19, 1928, McMinnville. As in No. 76,
this group is difficult to identify.
78. Lyc^nopsis pseudargiolus B. and L. (form or race
of, ?). 12 5 , 5 9 , May 27, 1930, Baker Creek Valley. Quite
common.
79. Lyc^nopsis pseudargiolus neglecta Edw., a sea-
sonal form. 1 5 , June 26, 1930, Baker Creek Valley. Scarcer
than the species.
APRIL, 1931] FENDER YAMHILL COUNTY BUTTERFLIES 187
80. Epargyreus tityrus Fabr., a species. 3 $ , July 12,
1927, Baker Creek Valley; 10 ^ , 8 9 , July 24, 1930, Baker
Creek Valley. Many to be taken in the foothills.
81. Pyrgus ruralis Bdv., a species. 2 $ , 2 9 , July 7,
1927, 1 5,3 9 , July 12, 1929, Baker Creek Valley. This one
locality is the only spot thus far found here for this butterfly.
82. Pyrgus tessellata occidentalis Skin., a race. 1 5 ,
1 9, July 18, 1928, McMinnville; 1 5, June 11, 1927, Grand
Island. Not numerous, but probably taken throughout this
county.
83. Pyrgus scriptura Bdv., a species. 1 5 , July 16, 1928,
Baker Creek Valley. Rare here as it evidently is most every-
where.
84. Erynnis persius afranius Lint. (?), a race. 2 5,
June 8, 1929, Muddy Valley; 2 5,19, July 24, 1930, Baker
Creek Valley. A relatively common skipper.
85. Erynnis Propertius S. and B., a species. 6 5,3 9 ,
May 12, 1926, Muddy Valley; 3 5,3 9 , May 4, 1928, Baker
Creek Valley; 6 5,4 9, May 28, 1929, Eola Hills. A very
plentiful hill skipper.
86. Pamphilida pal^mon Pall., a species. 2 5 , July 19,
1930, Baker Creek Valley. Uncommon skipper here as it is
elsewhere in the West.
87. PoLiTES SONORA Scud., a species. 1 5 , July 12, 1929,
McMinnville. A scarce capture in these parts.
88. PoLiTES MARDON Edw., a species. 1 5 , June 28, 1929,
McMinnville; 1 9, June 10, 1930, Baker Creek Canyon. A
rare butterfly.
89. PoLiTEs SABULETi Bdv., 2 l species. 20 5,9 9 , August
6, 1929; 45 5 9 , August 21, 1930, McMinnville. This skipper
swarms in quantity in the valleys during August.
90. Atrytone vestris Bdv., a species. 3 5 , July 13, 1928,
2 5,1 9 , July 24, 1929, Baker Creek Valley. Quite common
. in collections.
188
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
A NEW BRACHYTARSUS FROM CALIFORNIA
(Coleoptera — Platystomidse)
BY R. E. BARRETT
Saticoy, California
Brachytarsus franseria Barrett, n. sp.
Oval, robust, less than twice as long as wide, brown, thickly
clothed with short, dark brown and yellow pubescence. Head pice-
ous with sparse yellow pubescence; beak with sides slightly oblique,
tip of mandibles castaneous; eyes emarginate; antennal grooves
reaching the eyes; antennae inserted on the sides of the beak; seg-
ments 1 and 2 robust, the second slightly longer and noticeably
broader; segments 3 to 8 shorter and more slender; segments 9 to 11
forming an oval and very slightly compressed club; segments 1 to 9
castaneous; club piceous. Thorax as wide at base as long; basal
Carina strongly arcuate, reflexed at hind angles when viewed from
above, scarcely extending along the sides. Elytral striae evenly
spaced and parallel, tessellated on alternate intervals. Body beneath
finely pubescent; legs with femur piceous, tibia and tarsus casta-
neous, claws minutely cleft. Length, 2.0 to 2.5 mm.; breadth, 1.0
to 1.3 mm.
Holotype, No. 3005, in the Museum of the California
Academy of Sciences. Numerous paratypes in my own col-
lection. All specimens were taken by me on ragweed, Franseria
bipinnatifida Nutt., growing on the sand hills at Hueneme, Ven-
tura County, California. A series of specimens were first col-
lected July 18, 1926, and another series on April 16, 1931.
Considerable variation is shown in the series at hand. The
April specimens are darker and the elytra lack much of the
tessellated appearance. Four specimens collected in April have
the tibia and tarsus piceous.
This species should be placed near alternatus in our lists,
from which it may be readily separated by the color of the legs.
The numbers of Volume VII of the Pan-Pacific Ento-
mologist were mailed on the following dates : No. 1, on Sep-
tember 26, 1930; No. 2, on December 12, 1930; No. 3, on April
7, 1931; No. 4, on June 2, 1931.
APRIL, 1931] CHAMBERLIN CHILOPODS
189
ON THREE NEW CHILOPODS
BY RALPH V. CHAMBERLIN
University of Utah
The three new Chilopods described below were represented
in collections taken at quarantine by United States inspectors
at Honolulu and New Orleans, as indicated, and sent to the
author for determination. The types of the new forms are in
the author’s collection.
Cryptopid^
Cryptops vector Chamberlin, sp. nov.
General color yellow, with anal legs and especially the antennae
orange.
Cephalic plate smooth, wholly lacking the longitudinal sulci;
overlapping the first dorsal plate. Antennae composed of seventeen
very short articles of which the first five or six bear longer hairs.
First tergite with paired longitudinal sulci which meet anteriorly,
in a depression, a median angulation in the free transverse sulcus.
Second tergite with complete paired sulci. Succeeding tergites
also with paired sulci, but with no indication of keels. Prosternal
margin straight or slightly concave from end to end, without median
emargination. Last ventral plate short, strongly narrowed caudad;
the caudal margin rather wide, scarcely convex, the posterior corners
rounded. Coxopleura caudally truncate; bearing short spinescent
setae caudally and laterally; pores small and few in number, far from
reaching the caudal margin or dorsal plate.
Femur of anal legs bearing numerous spines ventrally, laterally,
and dorsally, with a few ordinary setae above, especially toward
distal end; dorsally with a median longitudinal furrow which is
deeper at caudal end; without teeth. Tibia with ventral and misal
surfaces bearing numerous spines, leaving no clear area, the ectal
and dorsal surfaces bearing ordinary setae; a single tooth near caudal
end beneath; a median longitudinal sulcus at caudal end above.
Metatarsus with a series of four teeth beneath. First tarsal joint
with two teeth beneath. Tarsi of all legs, excepting the last two
pairs, wholly undivided.
Length, about 16 mm.
Locality: One specimen taken at New Orleans, Louisiana,
on banana debris from Honduras, July 1, 1930.
This species seems to have nearest affinities with C. hyalinus
Say and C. detectus Silvestri. From the first it may be readily
separated by the absence of a median keel on tergites, the
presence of a dorsal longitudinal furrow on femur and tibia
190
THE PAN-PACIFIC ENTOMOLOGIST [vOL. VII^ NO. 4
of anal legs, and the presence of a single tooth on tibia beneath.
From detectus it differs in having four ventral teeth on the first
tarsal joint instead of only two, etc.
Lithobiid.(E
Tidabius vector Chamberlin, sp. nov.
Dorsally yellowish brown in color, with head and distal ends of
antennae and lights a brighter, somewhat lemon-yellow, color.
Antennae short, composed of twenty-six, mostly very short, arti-
cles. Eye patch linear, composed of about eight ocelli in two series,
e. g., 1 + 4, 3. Prosternal teeth uniform, 2 -f 2. Coxal pores, small,
round, 3, 3, 3, 3.
Ventral spines of first legs, 0, 0, 0, 0, 1. Penult legs with two
claws; ventral spines, 0, 1, 3, 3, 1; dorsal spines, 0, 0, 3, 1, 0. Anal
legs with two claws; ventral spines, 0, 1, 3, 2, 0; dorsal spines, 0, 0,
2, 0, 0. None of coxae armed.
Posterior legs of male simply inflated, with no special lobes.
Length, approximately 7 mm.
Locality: One male (holotype) taken at Honolulu, Hawaii,
July 17, 1930, in packing about orchids from Mexico.
Among species of the genus having the ventral spines of
the anal legs, 0, 1, 3, 2, 0, and the dorsal spines of the penult
legs, 0, 0, 3, 1, 0, agreeing with nasinttis Chamberlin and suitus
Chamberlin in bearing only two claws on the anal leg. From
these two species it seems to differ in smaller number of an-
tennal articles (26 as against 29-32), somewhat fewer ocelli,
and in having the coxal pores 3, 3, 3, 3 instead of uniformly
2, 3, 3, 3.
Ethopolid^
Bothropolys (Synopolys, subgen. nov.) epelus
Chamberlin, sp. nov.
Dorsum brpwn, the cephalic plate and antennae somewhat chest-
nut; legs yellowish, more brightly pigmented distally.
Antennae of moderate length, composed of the usual two articles.
Ocelli numerous, typically in four series in which the ocelli decrease
in size cephalad and ventrad, e. g., 1 + 6, 6, 4, 3. Prosternal teethS
small, black, uniform, in the holotype 6 + 6. Dorsal plates strongly
and irregularly subrugose. Posterior angles of seventh, ninth,
eleventh, and thirteenth dorsal plates produced, the angles of the
seventh plate less strongly but nevertheless distinctly produced.
Ventral spines of first legs, 0, 0, 2, 3, 2. Ventral spines of the
penult legs, 1, 1, 3, 2, 1; dorsal spines, 1, 0, 3, 1, 1; claw single. Axial
legs many. Last two pairs of coxae armed ventrally, laterally, and
APRIL, 1931]
CHAMBERLIN CHILOPODS
191
dorsally; the two preceding poriferous, coxae not armed ventrally.
Claw of female genital forceps tripartite, the lateral lobes small.
Basal spines slender, 3+3.
Length, near 15 mm.
Locality: One female, the holotype, was taken on the dock
at New Orleans, September 2, 1930.
This species, which may have escaped from a ship at the
dock, has its nearest known relations among the North Ameri-
can species of the genus. With these it agrees in the general
characters of head, prosternum, antennae, etc. It differs from
all of them in having the angles of the seventh tergite produced
while those of the sixth are not, and in having the basal spines
of the female gonopods 3 -j- 3 instead of 2 2. These two
characters are the basis of its subgeneric distinction from
PoropolySj in which neither the sixth nor the seventh dorsal
plates have their angles produced ; and from Bothropolys sens,
str. in which only the ninth, eleventh, and thirteenth dorsal
plates have their angles produced.
A New Pest of the Avocado in California
Caulophilus latinasus (Say) was found by the writer feed-
ingin the seeds of avocado in Orange County (April 12, 1931).
Adults and larvae of all sizes were found in the seeds, which
were completely riddled. This weevil has been known in the
Southern States since 1831, but record of its occurrence in the
West is lacking. It is listed as also occurring in the West
Indies, Mexico, and Guatemala. Other known hosts are chick-
peas and Indian corn. — R. E. Barrett.
Macrosargus cuprarius Linn, in California
On May 24, 1930, I noted many larvae of this Stratiomyid
in dry cow manure near Orangevale, Sacramento County. A
number of the slender, metallic bluish-green adults emerged
during June. Dr. J. M. Aldrich writes concerning this species
that it is of European origin, first reported in this country from
the vicinity of Philadelphia in 1896. It was known in Wis-
consin in 1913 and throughout Canada by 1927. The species
was determined by C. T. Green. — H. H. Keifer, Sacramento.
THE PAN-PACIFIC ENTOMOLOGIST
Published quarterly by the Pacific Coast Entomological Society
in co-operation with the California Academy of Sciences
E. P. Van Duzee, Editor S. B. Freeborn, Treasurer
In the April 3, 1931, number of Science^ Dr. Stiles expresses
much concern regarding the future of the international code
of zoological nomenclature, basing his unfavorable prognosis
upon the action of the Padua (1930) meeting of the Inter-
national Zoological Congress in amending the interndtional
rules in the open meeting of the Congress without such amend-
ment having been acted upon by the International Committee
on Zoological Nomenclature. There can be no question but the
work of the International Committee has been of inestimable
value to zoology through stabilization of its nomenclature, and
it is most earnestly to be hoped that no permanent rift will
rob these efforts of their international character. Probably
all active working entomologists follow these rules rigidly or
endeavor to do so. The so-called ‘'Opinions” are really but per-
sonal interpretations of the rules and there is not yet the same
unanimity in their observance. Some of these “Opinions” were
adopted by the committee without the unanimous agreement
required in the case of the rules and probably will be recon-
sidered later.
Among entomologists at least there is still some divergence
of opinion as to what constitutes the valid naming of a geno-
type. Apparently Fabricius was the first to indicate genotypes,
which he first attempted in 1794 by repeating, in certain genera,
the generic diagnosis in his description of the species he in-
tended should be taken as the type of the genus. Lamarck in
1801 undoubtedly was the first to designate types for each
genus of which he treats, and these type fixations are accepted
by many of our entomologists despite the fact that the Inter-
national Committee ruled in Opinion 79 that this work could
not be considered as fixing genotypes. This and other incon-
sistencies found in these “Opinions” will, without doubt, be
eliminated in time, but in any case they do not affect the validity
and great value of the international rules which should be, and
probably are, accepted by all workers in systematic zoology, in
this country at least. The foundation is satisfactory to all; the
superstructure only needs repairs in certain places. — E. P. Van
Duzee.
193
INDEX TO VOLUME VII
Abdastartus sacchari Drake, 15.
tyrianus Di6t., 15,
Aeolothrips aureus Mltn., 122.
Ancylandrena Ckll., 5,
Andrena, partial key, 7,
heterodoxa Ckll., 6.
Anomocera Cwf., 137.
Aphalara Frst., 132.
gutierrezise Klv., 134.
minutissima Cwf., 137.
minutistylus Klv., 135.
pulchella Cwf., 134.
suaedae Cwf., 133.
Aphalaroida Cwf., 138.
intermedia Klv., 139,
Aphodius davisi Fall, 73.
vandykei Brtt., 101.
Arctocorixa omani Hungf., 25,
blaisdelli Hungf., 26.
Argyresthia cupressella Wlshm.,
76.
franciscella Busck, 76.
. pilatella Braun, 76.
trifasciae Braun, 76.
Aligia, key, 119.
chiricana Ball, 121.
dellana Ball, 120.
descripta Ball, 119.
santana Ball, 121.
turbinata Ball, 120.
Bailey, S. F., paper by, 175.
Ball, E. D., papers by, 9, 119.
Bare, C. O., paper by, 115,
Barnes, William, Necrology, 16.
Barrett, R. E., note by, 191.
Barrett, R. E., paper by, 101, 188.
Bedunia adusta V. D., 64.
Beebea guglielmi Schaus, 1.
Bennarella Muir, 12.
bicoloripennis Muir, 12.
fusca Muir, 13,
Bequaert, J., paper by, 91.
Blaisdell, F. E., papers by, 17,
111 .
Bothropolys epelus Chmbl., 190.
Brachytarsus franceria Brtt.,
188
Bruck, C. R., paper by, 126.
Buenoa arizonis Bare, 115.
Butterflies of Yamhill County,
Oregon, 179.
Cactus Moth, 1.
Calyxochaetus, Monog., 53.
abbreviatus V. D., 54.
cilifemoratus V. D., 54.
clavicornis V. D., 55.
distortus V. D., 55.
fortunatus Whl., 56.
frontalis Lw., 56.
hastatus V. D., 57.
luteipes V, D., 57.
nodatus Lw., 58.
oreas Whl., 58.
sobrinus Whl., 59.
vegetus Whl., 59.
Cardiophorus aptopoides Cand.,
156.
Caulophilus latinasus Say, 191.
Chamberlin, R. V,, papers by, 65,
189.
Chermidse of Utah, etc, 131.
Chilopods, 65, 189.
Cockerell, T. D. A., paper by, 5.
Comstock’s Manual, Review, 2.
Corixidse, new, 22.
Cryptops navis Chmb., 65.
vector Chmbl., 189.
Cryptothrips rectangularis Hood,
123.
Dactuliothripidae Mltn., 173.
Dactuliothrips Mltn., 173.
spinosus Mltn., 174.
Diphyllostoma nigricollis Fall, 95.
Diptera, biting, 99.
Docessissophothrips animus
Mltn., 123.
Drake, C. J., paper by, 15.
Ecyrus, N. Am., 85.
arcuatus Gah., 90.
cornutus Linsl., 86.
dasycerus Say, 89.
194
INDEX TO VOLUME VII
Ecyrus fasciatus Hmlt., 90, 106.
hirtipes Gah., 89.
pencillatus Bates, 89.
texanus Sch., 90.
Epipocus subcostatus Gor., 156.
Epithectis californica Kfr., 27.
Essig, E. O., paper by, 97.
Euryptera cruenta Mrtn., 70.
Fall, H. C., papers by, 73, 154.
Fender, K., paper by, 179.
Flanders, S. E., paper by, 20.
Gelechia arbutina Kfr., 29.
manzanitae Kfr., 31.
panella Busk., 34.
Graptocorixa Hungf., 22.
abdominalis Say, 23.
californica Hungf., 23.
gerhardi Hungf., 25.
melanogaster Kirk., 25.
serrulata Chi., 23
uhleri Hungf., 23
Gunder's List of Butterflies, Re-
view of, 130.
Gunn Collection, 72.
Gunn, N. R., Necrology, 19.
Guthrie, E., paper by, 107.
Gyrinus instabilis Fall, 155.
Hatch, M. H., paper by, 103.
Hesperorhipis Fall, 74.
albofasciatus Fall, 75.
Heteropsylla Cwf, 140.
texana Cwf., 140.
Hinton, H., paper by, 94.
Hood, J. D., paper by, 170.
Hopping, R., paper by, 105.
Hungerford, H. B., paper by, 22.
Ischnocerus angulata Mrtn., 71.
Ischnorrhynchus obovatus, V. D.,
no.
Keifer, H. H., note by, 191.
Keifer, H. H., papers by, 27, 76.
Klyver, F. D., papers by, 131,
157.
Krizousacorixa Hungf., 22.
Lawson, P. B., paper by, 159.
Lepidomys irrenosa Gn., 8.
Leucothrips Rent., 170.
nigripennis Rent., 171.
Lina scripta Fab., 107.
Linsley, E. G., papers by, 77, 106.
Listrus n. tincticornis Blsd., 19.
Macrosargus cuprarius L., 191.
Malachius calif ornicus Brtt., 102.
Martin, J. O., paper by, 70.
McGlashan, C. F., Necrology, 97.
Megalothrips picticornis Hood,
172.
Melyridas, Studies in, 17.
Microlepidoptera, 27, 76.
Moulton, D., papers by, 122, 173.
Muir, F., paper by, 12.
Needham’s Odonata of China,
169.
Nipponobius sinensis Chmb., 68.
Nymphocixia vanduzeei Muir,
13.
Odonata of China, 169.
Odontomyia pilosa Day, 104.
Odynerus margaretellus Rohw.,
72.
Otostigmus sinicolens Chrtib., 66.
mians Chmb., 67.
Paratrioza cockerelli Sulc., 142.
Pentatomidse of Utah, 124.
Phasdon, note on, 103.
oklahomensis Htch., 103.
planus Htch., 104.
Phlaeothrips ornatus Hood, 175.
Phloeosinus aciculatus B ruck, 127.
variolatus Bruck, 126.
Platylygus vanduzeei Usng., 129.
Plebejus i. moroensis Strny., 93.
Ploas atratula Lw., 104.
Poecilothrips ornatus Hoodj 123.
Pogonocherus, N. Am., 77, 105.
arizonicus Sch., 80.
californicus Sch., 84.
concolor Sch., 84.
crinitus Lee., 80.
fasciculatus DeG., 83.
mixtus Hald., 81.
negundo Sch., 84.
obscurus Fall, 85.
parvulus Lee., 106.
penicillatus Lee., 83.
pictus Fall, 80.
THE
Pan -Pacific Entomologist
Published by the
Pacific Coast Entomological Society
in co-operation with
The California Academy of Sciences
VOLUME SEVEN
July, 1930, to April, 1931
San Francisco, California
193 1
11
Contents of Volume Seven
Ball, E. D.
A New Species and Variety of Scolops with Notes on
Others 9
Some New Leafhoppers of the Genus Aligia 119
Bailey, Stanley F.
A Thrips New to California 175
Bare, Clarence O.
A Buenoa of Southwest United States and Mexico 115
Barrett, R. E.
New Species of Aphodius and Malachius from Cali-
fornia 101
A New Brachy tarsus from California 188
Bequaert, J.
Ashmead’s Genus Polistella 91
Blaisdell, Frank E., Sr.
Studies in the Melyridse, Number Eight 17
A New Species of Zopherodes from Central California....! 11
Bruck, C. R.
Two New Species of Bark Beetles of the Genus PhlcEO-
sinus Chapius 126
Chamberlin, Ralph V.
On Some Chilopods Immigrant at Hawaii 65
On Three New Chilopods 189
Cockerell, T. D. A.
A New Subgenus of Andrenine Bees 5
Drake, Carl J.
A New Sugar-Cane Tingitid from Java and Sumatra 15
Essig, E. O.
Charles Fayette McGlashan 97
Fall, H. C.
A New Aphodius and a New Genus and Species of
Buprestidse from California 73
A New Gyrinus from Alaska, with References to Other
Recently Described Species 154
Fender, Kenneth
Butterflies of Yamhill County, Oregon 179
Flanders, Stanley E.
Races of Trichogramma minutum 20
Ill
Guthrie, Esther
Notes on Egg Hatching — Larval, Pupal, and Adult De-
velopment in Lina scripta Fabr 107
Hatch, Melville H.
Notes on Phaedon 103
Hinton, Howard
Observations on Two California Beetles 94
Hood, J. Douglas
Synonyms in the North American Thysanoptera 170
Hopping, Ralph
Notes on Pogonocherus 105
Hungerford, H. B.
New Corixidae from Western North America 22
Keifer, H. H.
California Microlepidoptera IV 27
Klyver, F. D.
Chermidae from Utah, Nevada, and Arizona, Including
Three New Species 131, 157
Lawson, Paul B.
The Genus Xerophloea in North America 159
Linsley, E. Gorton
New Pogonocherus and Ecyrus, with Notes Concerning
Others 77
Martin, J. O.
Two New Coleopterous Insects from Arizona 70
Moulton, Dudley
A New Aeolothrips from Nevada with Notes on Three
Other Species Found in California 122
An Interesting New California Thrips 173
Muir, F.
Three New Species of American Cixiidae 12
Stanford, J. S.
Notes on Diptera Attacking Man in Sevier County,
Utah ^ 99
Sternitzky, R. F.
A New Subspecies of Plebejus icarioides Bdv 93
Thorpe, W. H.
Miscellaneous Records of Insects Inhabiting the Saline
Waters of the California Desert Regions 145
IV
Usinger, R. L.
A New Species of Platylygus 129
Van Duzee, E. P.
A New Bedunia 64
A New Ischnorrhynchus 110
Van Duzee, M. C.
The Dipterous Genus Sympycnus Lowe in North
America and the West Indies .....35, 49
Williams, Francis X
Beebea guglielmi Schaus, A Pyralid Moth Borer of the
Opuntia Cactus in the Galapagos Islands 1
Woodbury, Lowell A.
A List of the Pentatomidae of Zion National Park,
Utah 124
INDEX TO VOLUME VII
195
Pogonocherus pilatei V. D., 83.
salicicola Csy., 81, 106.
vandykei Linsl., 82.
volitans Lee., 85.
Polistella Ashm., 91.
Saline Water Insects, 145.
Scolops graphicus Ball, 9.
u. marginatus Ball, 10.
perdix Uhl., 11.
stonei Break., 11.
Stanford, J. S., paper by, 99.
Sternitzky, R. F., note by, 19.
Sternitzky, R. F., paper by, 93.
Sympyenus, Mong., 35.
aldrichi V. D., 39.
angustipennis Aid., 39.
aurifacies V. D., 39.
breviventris V. D., 40.
calcaratus V. D., 41.
canadensis V. D., 42.
caudatus V. D., 42.
clavatus V. D., 42.
cuprinus Whl., 43.
fasciventris V. D., 43.
globulicauda V. D., 43.
inaequalis V. D., 44.
laevigatus V. D., 45.
latitarsis V. D., 46.
lineatus Lw., 46.
marcidus Whl., 47.
montanus V. D., 47.
pugil Whl., 49.
pulvillus V. D., 49.
setosus V. D., 50.
tertianus Lw., 51.
thoracicus V. D., 51.
tripilus V. D., 52.
varipes Aid., 53.
Synopolys Chmbl., 190.
Thorpe, W. H., paper by, 145.
Thrips herricki Bagn., 171.
Thysanoptera, 122, 170, 173, 175.
Tidabius vector Chmbl., 190.
Treherniella amplipennis Morg.,
172.
Trichochrous 4-notatus Blsd., 17.
punctipennis Lee., 19.
Trichogramma minutum, 20.
Trioza lobata Cwf., 143.
Type vs. description, 96.
Usinger, R. L., paper by, 129.
Valgus californicus Horn, 94.
Van Duzee, E. P., notes by, 8,
16, 69, 104, 130.
Van Duzee, E. P., papers by, 64,
no.
Van Duzee, M. C., paper by, 35.
Van Dyke, E. C., note by, 95.
Williams, F. X., paper by, 1.
Woodbury, L. A., paper by, 124.
Xerophloea Germ., key, 160.
brunnea Laws., 163.
californica Laws., 163.
inflata Laws., 164.
majesta Laws., 165.
major Bak., 169.
minor Laws., 162.
obtusa Laws., 162.
oraclis Laws., 165.
robusta Laws., 164.
tenuata Laws., 165.
vanduzeei Laws., 161.
viridis Fab., 166.
zionis Laws., 161.
Zoological Nomenclature, 192.
Zopherodes, key, 113.
sanctaehelenae Blsd., 111.
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