1. TRICHOPTERA

By Martin E. MosEty

(With Plates I—IIT)

THE TRICHOPTERA taken on the British Museum Expedition to East Africa

form a very representative collection. With the exception of the Polycentro-

pidae, all the Families known on the African continent are present.

It is, perhaps, a little surprising that species of the genus Dipseudopsis,

belonging to the missing Family, were not taken as the genus is very widely

distributed throughout the continent.

The Sericostomatidae are represented by four species of the genus Goerodes.

A considerable number of specimens of most of the species were taken. Of the

Calamoceratidae, only one example was secured, Anisocentropus usambarensis.

The Leptoceridae are well represented, species from no fewer than six genera

being present. The Hydropsychidae appear in the genera Hydropsyche,

Cheumatopsyche and Diplectronella, examples in the first genus having been

taken in great numbers. Two species in the Psychomyidae, of which, it is of

interest to note, one is a Lye, a genus not hitherto recorded in Africa, four in

the Philopotamidae, one in the Rhyacophilidae, and four in the Hydroptilidae,

complete the list. The captures in the last two Families have necessitated the

erection of three new genera.

Records of species taken in the Ruwenzori Range are covered by the dates

I.xii.1934 to 31.1.1935, and, unless otherwise indicated, the types are in the

British Museum collection.

SERICOSTOMATIDAE

THE GENERA Goerodes, Crunoeciella AND Crunobiodes

Dr. Edwards brought home from Uganda long series of two species de-

scribed by Navas as Crunoeciella nudata and Crunoeciella inferior as well’as a

few examples of allied species.

As there is some confusion between Crunoeciella, Crunobiodes and Goerodes,

it is desirable to clear up the position as far as possible here.

In the Ann. Mus. Zool. Sc. U.R.S.S., 1927, Martynov pointed out that

certain species which had been placed in Ulmer’s genus Crunoectella, erected to

take a Madagascan species brumnea, failed to conform to the characters of this

ll, I

iS)

RUWENZORI EXPEDITION

eenus; he therefore erected Crunobiodes in which they might be placed. The

main characters of this new genus are firstly, a fold in the post-costal region of

the anterior wing and secondly a pattern of genitalia showing, with some minor

specific modifications, two pairs of dorsal processes and a branched inferior

appendage, one branch of which is always situated on the upper margin, near

the base, and directed upwards. This conformity of neuration and genitalia

appears to be common to a considerable group of species and is first met in

Ulmer’s Goerodes, a genus erected in 1907 and consequently having priority over

Crunobiodes. A study of numerous African species in this group, all of which

ccnform fairly well with this pattern of genitalia, has shown that there is, in

almost every case, a modification in the position and iength of the post-costal

fold which might be considered by some to be of generic value.

In the Lepidostomatinae, the sub-family of the Sericostomatidae to which

these species belong, neuration seems to be extremely abnormal in individual

species, more particularly in the 3 sex, whilst in the 2 the neuration is usually

regular. It has generally been the practice to accept neuration as giving the

generic characters and to base the specific distinctions on the genitalia, but to

make this custom into a hard and fast rule in the Lepidostomatinae would

appear to bring about a position in which, eventually, every species would

require a separate genus to contain it.

It is here suggested that, in this sub-family, the practice should be reversed

and that where species can be conveniently grouped on a similarity of genitalia,

this character should be considered as pertaining to the genus and that the

species should be separated on neuration. This suggestion is not really revo-

lutionary as it was foreshadowed as far back as 1876 when McLachlan, writing

of another genus, Dinarthrum, in this sub-family, states “there probably exist

a number of species that practically will find a home in this genus but

presenting slight differences zmtey se in the antennae, palpi and neuration, the

latter being of such a character as to defy description.” The italics are mine.

I propose, therefore, to re-define Ulmer’s Goerodes and to transfer to it the

species nudata, inferior, and excelsior, represented in Dr. Edwards’s collection

and also the South African species caffrariae Barnard, all unsuitably placed in

Crunoeciella.

With regard to this Madagascan genus, Dr. Zerny of the Vienna Museum

very kindly sent me the type of C. bvunnea with permission to clear the genitalia

in KOH, and Iam thus enabled to supplement here the good figures of the dried

insect published by Ulmer in his original description. It will be noted that, in

addition to the absence of the post-costal fold, the genitalia, while closely akin,

scarcely conform to the Goerodes pattern. This type of C. byunnea was found

to be in fragments, and to preserve them, these have now ‘been returned to

the Vienna Museum in the form of a balsam preparation.

TRICHOPTERA 3

Crunoeciella Ulmer

Crunoeciella Ulmer, 1905, Ann. Naturh. Mus. Wien, 20: 68, figs.

Spurs, 2, 4, 4. Antennae long, basal joint longer than the head. Wings

covered with thickened hairs; neuration as shown in the figure with no post-

costal fold in the anterior wing. The structure of the genitalia is detailed in

the specific description.

Genotype and only known species, C. brunnea Ulmer, of Madagascar.

Crunceciella brunnea Ulmer (Figs. 1-5)

Crunoeciella byunnea Ulmer, 1905, Ann. Naturh. Mus. Wien, 20: 68, figs.

Figs. 1-5.—Crunoeciella byrunnea Ulmer g: 1. wings, 2. genitalia, dorsal,

3. dorsal plate, 4. genitalia, lateral, 5. inferior appendages, ventral.

Genitalia g. From above, dorsal plate membranous and cleft to form a

pair of finger-like processes with truncate apices, one longer than the other,

4 RUWENZORI EXPEDITION

showing as a deep plate from the side; below it is a pair of long spines, each

bifurcate at the apex, the inner fork long and slender, the outer shorter and

broad. Penis short and down-curved. Inferior appendages from the side, very

long and slender, curving slightly upwards with the inner surface bearing short,

clavate hairs as well as long hairs of the usual type; towards the base of the

appendage is a pair, or perhaps a single, widely-bifurcate, upwardly-directed

process, the outer fork bent over towards its apex which is much dilated at its

lower margin, upper margin produced in a small point; inner fork straight and

directed upward; from beneath, there is a small pointed process at the base

of the inner margin of each appendage.

MapaGascar : Fort Dauphin, Sikova.

Type 3, the only known specimen, now mounted entirely in balsam, in

the collection of the Vienna Museum.

Goerodes Ulmer

Goevodes Ulmer, 1907, Cat. Coll. Selys, fasc. 6 (1): 37-38.

Crunobiodes Martynov, 1927, Ann. Mus. Zool. Acad. Sci. U.R.S.S., 28: 471.

Crunoeciella Auct.

The genus is here based mainly upon characters furnished by the male

genitalia. The species show, in the male, a considerable diversity in the form

of neuration of the anterior wings so that the neuration should be considered as

of specific rather than generic value. There is, however, one character of

neuration which all the males have in common and which may be regarded as

of generic importance, namely the presence, in the anterior wing, of a fold in

the region of the post-costa, but the scale-lined grooves which so often appear

in the Lepidostomatinae, particularly in the Dinarthrum group, are wanting.

In the female, the neuration of the anterior wing is regular.

I would here lke to distinguish between the terms “‘groove”’ and “fold.”

The former is a more or less open furrow, usually but not always situated along

the central region of the anterior wing; the latter, a fold of membrane generally

in the post-costal region (Martynov’s anal groove), possibly welded to the

adjacent membrane and simulating a nervure. The insects are uniformly

brown of varying shades, the males with wings densely clothed with hairs

intermingled, but not invariably, with scales; in the female wings, there are

never either scales or folds; the posterior wings in both sexes have a regular

neuration. .

Basal joint of the antenna unbranched. Maxillary palpi 3 generally two-

jointed, shorter than the labial palpi, terminal joint which is rather obscure,

carrying a tuft of dark and much broadened scales. Spurs 2, 4, 4.

Genitalia 3. All species in the genus conform in a production of the centre

PERE 4 se >

TRICHOPTERA 5

of the dorsal margin of the ninth segment in two pairs of processes, the outer

usually appearing as asymmetric, sinuous and very stout spines, the inner

varying in form ; sometimes these processes are replaced by a plate or plates.

The penis short and curved; inferior appendages always branched, the

branches varying in number and form. They agree in all species in carrying a

more or less erect branch, generally with a dilated apex, arising from the

upper margin of the appendage towards the base, as seen from the side.

This pattern of genitalia, though fairly constant in so far as the South

African species are concerned, is in the Indian and Far Eastern forms inclined

to vary in respect to the processes of the dorsal plate, so that it is sometimes

difficult to fix a dividing line between these paired processes and a structure

representing the pairs welded together.

Genotype here designated, Goevodes cornigera Ulmer, in the De Selys

collection in the Brussels Museum, from Japan.

Goerodes edwardsi sp. n. (Figs. 6-8)

The only conspicuous character by which this species differs from excelstor

Navas is to be found in the wings. In the latter, these are clothed with hairs

Figs. 6-8.—Goerodes edwardsi sp. n. g: 6. wings, 7. inferior

appendage, lateral, 8. ventral.

and numerous small scales; in edwardsi, the vestiture is entirely of hairs, there

are no scales. There are also certain differences in neuration, more particularly

at the distal end of the post-costal fold.

6 RUWENZORI EXPEDITION

In the 3g, there are slight differences in the genitalia, especially in the form

of the inferior appendages. As, however, only one example of edwardsi was

taken, allowance must be made for individual variation.

Length of the anterior wing, § 15 mm.

RUWENzORI: Namwamba Valley, 10,200 ft. (Edwards). 1 3 (type).

Goerodes excelsior (Navas) (Figs. 9-12)

Crunoeciella excelsioy Navas, 1931, Broteria Zool., 27: 132-3, fig. 68.

Through the courtesy of Dr. L. Berland of the Paris Museum, I have been

enabled to examine the type of this species and to give additional figures of the

neuration and genitalia.

Figs. 9-12.—Goerodes excelsiov Navas $: 9. wings, 10. genitalia, dorsal,

11. lateral, 12. inferior appendages, ventral.

Genitalia g. Ninth segment produced at the centre of its dorsal margin in

two pairs of processes the inner, short, triangular, with serrate outer margins,

the outer, asymmetrically sinuous, very strongly chitinized and nearly three

times the length of the inner; penis stout, arching strongly downward with

TRICHOPTERA G7

dilated apex, its apical margin slightly excised, inferior appendages four-

branched, the main branch, from the side, narrowing suddenly to a rather

slender termination with a dilated apex; the second branch arises at the point

where the main branch narrows and is short and rather broad, directed distally

and slightly downward; the third branch, seen from the side, arises from the

upper margin of the main branch, near the base, rod-shaped, without any

dilatation of the apex; fourth branch, from beneath, arises at the base of the

appendage on its inner margin and is short, rather broad, directed distally and

downward. ;

Length of anterior wing, 3 15 mm.

RUWENZORI: Namwamba Valley, 10,200 ft., (Edwards) 1 3; Nyamgasani

Valley, 12-13,000 ft., (Buxton) 2 3.

Type g in Paris Museum from ‘“‘Buamba”’ (presumably Bwamba, West

Ruwenzori).

Goerodes inferior (Navas) (Figs. 13-19)

Crunoeciella inferioy Navas, 1931, Broteria Zool., 27: 135.

Head fuscous, antenna with the basal joint rather larger than the breadth

of the head and oculi together, clothed with dense, fuscous pubescence; palpi

membranous, possibly two-jointed, with a tuft of broad, black scales at the

apex and a minute, membranous, projecting process or wart towards the base;

wings dark fuscous, anterior very broad and rounded, apex considerably

dilated, particularly along the posterior margin; both wings covered with long,

silky black hairs but no scales; anterior with forks Nos. 1 and 2 and perhaps 5;

posterior, fork No. 1 only; discoidal cell of the anterior wing long and rather

narrow, about one and a half times as long as its footstalk; that of the posterior,

short, distal nervure very oblique; fourth apical cell extending not so far

inward as the base of the discoidal cell.

Genitalia 3. Ninth segment produced at the centre of its dorsal margin

in two pairs of processes, the inner, rather more slender than the outer; the

processes terminate in strongly chitinized claws which are asymmetric as to the

‘ direction in which they turn, some up, some down; from the side, the outer

process is sinuous along its lower margin; penis short and slender, arching

downward; inferior appendages three-branched ; main branch very stout at its

base, from the side, slightly dilated at about three-quarters of its length on its

lower margin which is there deeply excised so that there is a triangular, clavate

apex to the branch; from above and beneath, the portion of the appendage

8 RUWENZORI EXPEDITION

beyond the excision forms a narrow, finger-like first branch; the second

branch arises towards the base of the appendage from its upper margin; from

the side, it is moderately long with a slender stem and much dilated, clavate

apex; third branch from beneath; very slender, rather short, arising from the

base of the lower margin; from beneath, the appendages are separated by a

Figs. 13-16.—Goerodes inferioyv Navas: 13. wings g, 14. anterior wing 9, 15.

maxillary palpus g, 16. labial palpus 3,

wide, rounded, horseshoe-shaped excision; margin of the ninth ventral segment

produced at its centre and on each side.

Length of anterior wing, g Io mm., 2 9 mm.

RUWENzORI: Namwamba Valley, 4500—-10,200 ft.; Mobuku Valley, 7800 ft. ;

Bwamba Pass, West Side, 5500-7500 ft. (Edwards). 39 5 Qin all.

Type g in the Paris Museum, from ‘“‘ Monts Rouwenzori”’ (Ch. Alluaud,

1900).

TRICHOPTERA 9

This insect is readily separated from the other species of Goerodes,

described in this paper, by the short, somewhat arched wings, with their

broadened apices and dense clothing of long, silky hairs.

Figs. 17-19.—Goerodes inferioy Navas $: 17. genitalia, dorsal, 18. lateral,

19. inferior appendages, ventral.

In addition to the series secured by Dr. Edwards, a single example has

been subsequently placed in the Museum collection, taken by Miss C. Long-

field on the Mobuku River, Ruwenzori, at the slightly lower altitude of

4000 ft.

Io RUWENZORI EXPEDITION

Goerodes nudata Navas (Figs. 20-25)

Crunoeciella nudata Navas, 1931, Broteria Zool., 27: 134.

Head very dark fuscous, basal joint of the antenna rather longer than the

breadth of the head and oculi together; two very large tufts of hairs arising

Figs, 20-25.—Goerodes nudata Navas: 20. wings g, 21. wings 9, 22. maxillary palpus S;

23. genitalia g, dorsal, 24. lateral, 25. inferior appendages, ventral.

from the head between and behind the bases of the antennae; maxillary palpi

of the ¢ possibly two-jointed, with a tuft of broad, black scales at the apex;

aiken

TRICHOPTERA Il

wings black, anterior 3 with the costal margin doubled over at the base, en-

_ closing a conspicuous tuft of black hairs and with a long fold extending almost

to the margin (the fold is opened out in the figure); forks Nos. 1 and 2 present;

wing covered with numerous black scales; discoidal cell long and narrow;

posterior wing with only fork No. r present, thinly sprinkled with narrow, black

scales; anterior wing 2 with forks Nos. 1, 2, 3 and 5; posterior with forks

Nos. I, 2 and 5.

Genitalia §. Excepting for some slight variation in the proportions of the

various parts, particularly in the branches of the inferior appendages, and a

raised, triangular upper-part to the ninth segment, the genitalia of nudata are

indistinguishable from those of znferior.

Length of anterior wing, ¢ 14 mm., 2 13 mm.

RUWENZORI: Namwamba Valley 4500-10,200 ft. (Edwards). 155 3 @

in all.

Type ¢ in the Paris Musuem from Ruwenzori, Kichouchou, 3000 m.

(Ch. Alluaud, 1909).

CALAMOCERATIDAE

Anisocentrops usambarensis Ulmer

Anisocentropus usambarensis Ulmer, 1910, in Sjéstedt’s Kilimandjaro-Meru Expd.,

1905-6, 13, Neur.: 5-6.

Kenya: Thika, 4500 ft., 23.x.1934 (Edwards). 1 9.

LEPTOCERIDAE

Leptocerus asanus sp. n. (Figs. 26-29)

Head black with cream-coloured hairs between the antennae; these are

black with no apparent annulations; legs very dark fuscous, nearly black ;

wings with blackish pubescence, membrane possibly yellowish and conspicu-

ously yellow in the apical area.

Genitalia g. Apical margin of the ninth dorsal segment produced and

excised at its centre with a lower projecting shelf which is still more excised;

superior appendages moderate in size and conical in shape; upper penis-cover

in the form of a pair of strongly chitinized spines curving slightly upwards as

seen from the side; arising from their upper surfaces at their bases is a pair of

small, transparent and bifurcate branches, the apices of the forks generally

truncate and armed with one or two bristles; there is a pair of penis-sheaths

with dilated, bifurcate apices and a membranous penis; inferior appendages

12 RUWENZORI EXPEDITION

with a very stout basal and branched apical portion; upper branch from the

side, fuscous in colour, short and straight and forming an extension of the

upper apical angle of the basal portion; second branch rather transparent,

Figs. 26-29.—Leptocevus asanus sp. n. $: 26. wings, 27. genitalia, dorsal, with

superior appendages and upper penis-cover, 28. genitalia, lateral, 29. ventral, with

inferior appendages, penis and sheaths.

arising at the base of the first and forming a down-curving hook; the appendages

are concave, inner margin set with strong teeth, outer surface strongly fringed.

Length of the anterior wing, 3 9 mm.

RUWENZORI: Nyamgasani Valley, 12-13,000 ft. (Buxton). 1 3 (type).

Leptocerus sp.

In the absence of a g, I refrain from giving a name to or a description of

this insect as, in the female sex, no very definite characters are available by

which closely related species may be separated.

RUWENZORI: Bwamba Pass, West Side, 5500-7500 ft. (Edwards). 3 9.

TRICHOPTERA 13

Leptocerina talopa sp. n. (Figs. 30-33)

Head black, clothed with cream-coloured hairs; basal joint of the antennae

also clothed with cream-coloured hairs, only one or two of the other joints

remaining; these appear to be clothed dorsally.with white hairs as is usual in

the genus; palpi fuscous; legs ochraceous with white hairs at the bases of the

tarsal joints; wings uniformly dark ochraceous, clothed with short, fuscous

hairs; there is no apparent white streak or white patch at the anal angle of the

anterior wing as occurs in other species in the genus.

Figs. 30-33.—Leptocerina talopa sp. n. g: 30. wings, 31. genitalia,

dorsal, 32. lateral, 33. ventral.

Genitalia g. Apical margin of the ninth dorsal segment produced in a wide

triangle with a slight projection of the under surface at the apex; superior

appendages narrow from the side, only partly visible from above; there is a

projecting angle on the inner margin at the base, only to be seen in a cleared

example; upper penis-cover divided in a pair of strongly chitinized plates or

spines, arching over a membranous penis; in the type, the penis, towards its

apex, seems to have been pushed up between the plates so that it appears above

them; inferior appendages large, two-branched, the outer margin of the lower

branch strongly serrate, the upper branch arising towards its base and directed

upward and distally with a truncate and serrate apical margin; there is a

14 RUWENZORI EXPEDITION

triangular projection at the lower margin about midway; the centre of the

margin of the eighth sternite is slightly produced but scarcely enough to make

a ventral process.

Length of anterior wing, ¢ 9 mm.

W. UcanpDA: Fort Portal district, 5000 ft. (Edwards). 1 3 (type).

The species very closely resembles L. mlanjensis Mos. from Mt. Mlanje,

Nyasaland. There are various slight differences in the form of the genitalia

and also in the neuration which should suffice to separate them.

Triaenodes legona sp. n. (Figs. 34-38)

Head and thorax greyish-fuscous; antennae greyish-ochraceous with no

apparent annulations; wings reddish-fuscous with a few distinct, black marks

on the anterior, in the post-costal region towards the base of the wing and three

Figs. 34-38.—Tyriaenodes legona sp. n. g: 34. wings, 35. dorsal plate, superior

appendages and upper penis-cover, 36. genitalia, lateral, 37. upper branch inferior

appendage, lateral, 38. genitalia, ventral.

or four round the lower apical margin; bordering on these spots, on the basal

side, are patches of golden hairs; pubescence otherwise reddish; legs ochraceous.

Genitalia 3. Apical margin of the ninth dorsal segment produced in a pair

of short, parallel, finger-like processes ; superior appendages very slender, rod-like

TRICHOPTERA £5

and fringed with long hairs; beneath them is an upper penis-cover composed

of two long, slender spines, the apices crossing each other in the example under

description; penis broad, arching downward, apex bifurcate; inferior appen-

dages with two branches, the lower, large, bearing on its inner surface, a mass of

strong teeth; the second branch arises at the base of the lower, from its upper

surface; it has a slender stem and broad, furcate apex; from the side, the lower

fork is directed downward with an acute apex armed with a single bristle;

the upper directed backward with a truncate apex furnished with a few bristles;

there is a small angular projection between the two forks; it is probable that

the shape of the branches will vary slightly in individuals.

Length of anterior wing, ¢ 6 mm.

Kenya: Mt. Elgon, 10,000-11,500 ft., 11.1935 (Edwards). 2 3 (type and

paratype).

Triaenodes wambana sp. n. (Figs. 39-43)

Head ochraceous with cream-coloured hairs; antennae, basal joint long,

ochraceous, other joints ochraceous with dark annulations; legs ochraceous;

Figs. 39-43.—Tyviaenodes wambana sp. n. S$: 39. wings, 40. dorsal pro-

cesses, 41. genitalia, lateral, 42. ventral, 43. inferior appendage from within.

wings ochraceous clothed with ochraceous hairs with here and there small

patches of dark hairs, more particularly along the lower apical border.

Genitalia g. Superior appendages long, finger-like and fringed; beneath

them is a strongly chitinized fork or pair of spines arising from a broad base

16 RUWENZORI EXPEDITION

and directed somewhat downward; penis arching downward with a mem-

branous, two-lobed apex; inferior appendages from beneath, caliper-shaped ;

from the side, the lower or first branch is broad at its base, the apical half with

a concave upper margin so that it tapers gradually to an acute apex; at the base

there arises from the upper margin a distally directed second branch with a

very broad apex, the distal margin of which is sinuously truncate; arising from

just above the base of this branch is a third branch in the form of a strongly

chitinized, sickle-shaped spine curving downward so that its apex is nearly

level with the lower margin of the first branch; this branch has a short spur

about midway; the fourth branch is broad, thickly covered with long spines

on its inner surface; ninth segment from beneath, strongly produced with its

apical margin widely excised.

Length of anterior wing, ¢ 7 mm.

RUWENZORI: Bwamba Pass (west side), 5550-7500 ft. (Edwards). 3 (type).

Triaenodella hastata Ulmer

Triaenodes hastata Ulmer, 1910, in Sjéstedt’s Kilimandjaro-Meru Expd., 1905-6,

13, Neurop: 7-8.

RUWENZORI: Namwamba Valley, 6500 ft., 1 ¢; Fort Portal, 5000 ft.,

(Edwards) 5 9.

The type was described in Ty1aenodes, but the presence of a plate and pencil

of hairs attached to the basal joint of the antenna indicates its position in

Triaenodella.

Oecetis kathia sp. n. (Figs. 44-47)

Head pale reddish-ochraceous with similarly coloured hairs; antennae very

heavily fringed in the basal half, but on one side only; palpi and legs pale

ochraceous; wings cream-coloured with darker shading at the anastomosis;

apex of the anterior wing unusually broad, costa considerably rounded. Spurs:

Ti 2ee2.

Genitalia g. Apical margin of the ninth tergite excised, with the centre of

the excision slightly produced; there is a trifid dorsal plate, all the lobes short,

the middle the shortest, outer lobes stout, central lobe slender; the ninth

segment is further produced at the edges of the excision in a pair of long, down-

curving, narrow plates or spines, each with a small, angular projection at its base

on its outer margin; above these, there lies a pair of minute, transparent superior

appendages, each with the apex armed with a long bristle; in the type, one of

these appendages is aberrantly paired; these processes are so small and in-

conspicuous that I have only been able to make them out under a j}-inch

microscope objective; penis strongly chitinized, arching downwards, with a

TRICHOPTERA 17/

small projection at about midway along its upper margin; inferior appendages

very long and sinuous towards the apices which are incurved; they arise from

stout bases which bear large, rather irregular, upwardly-directed branches on

their upper margins towards the base, as seen from the side.

Length of the anterior wing, ¢ 7 mm.

Kenya: Thika, 4500 ft., 23.x.1934 (Edwards). 4 3 (type and paratypes).

The species closely resembles in its genitalia Potamoryza modesta Barnard.

The feathered antennae and details of neuration suggest that possibly a new

genus should be erected to take it.

Figs. 44—-47.—Oecetis kathia sp.n. 3: 44. wings, 45. portion of an antenna,

46. genitalia, dorsal, 47. lateral.

Oecetis angustipennis sp. n. (Figs. 48-52)

Head and body pale ochraceous; antennae cream-colour with dark

annulations; legs cream-colour; wings very narrow and translucent, anastomosis

dark; Spurs o(?), 2, 2.

Genitalia g. Apical margin of the ninth dorsal segment possibly excised

at its centre where there is a trifid dorsal process, the outer lobes short and

shuttle-shaped, the inner, long and slender, apex rounded, almost truncate

III, Ib

18 RUWENZORI EXPEDITION

and fringed with a few short bristles ; at the base of the outer processes, between

them, lies a pair of short, membranous finger-like appendages, one longer than

the other; beneath the dorsal plate is a pair of superior appendages which, from

the side, are broad at the base, apical parts slender and terminating each in a

single bristle; these are difficult to make out from above and are crossed in the

type; penis large, arching downward, asymmetric; on one side of the apex there

is a broad, downwardly directed process; on the other, a short, strongly fringed

Figs. 48-52.—Oecetis angustipennis sp. n. g: 48. wings, 49. genitalia,

dorsal, 50. lateral, 51. penis from the other side, 52. genitalia, ventral.

lobe; inferior appendages large, three-branched, upper branch slender, arising

at the base and directed upward; central branch produced and curving slightly

outward from above; third branch, from the side, appears as a small spur

projecting from the lower margin of the main branch; from beneath, the third

branches are broad with serrate, rounded margins.

Length of the anterior wing, g 7mm.

RUWENZzORI: Namwamba Valley, 6500 ft. (Edwards). 1 § 1 @ (type and

allotype).

TRICHOPTERA I9

Oecetis portalensis sp: n. (Figs. 53-55)

The single example of the species is in very poor condition and lacks its

antennae and most of its legs. The head and body appear to be ochraceous.

Genitalia 3. The apical margin of the ninth tergite is produced in a trifid

process of which the central lobe is much the longest and curves slightly down-

wards; superior appendages arise from beneath the outer lobes and are very

slender, each armed at the apex with a long bristle; the upper penis-cover

seems to be partly attached to the lateral margin of the ninth segment and is in

the form of a pair of long, down-curving spines, flattened from above, apices

Figs. 53-55.—Oecetis portalensis sp. n. g: 53. wings, 54. genitalia, dorsal, 55. lateral.

somewhat rounded but with the inner apical angles slightly produced; penis

doubtful as to form; lower penis-cover trough-shaped, arching downward,

terminating in an acute tooth; inferior appendages broad at the base, then very

long and slender, curving inwards from above; about midway along the inner

margin of the appendage is a minute angular projection; from the side, there is a

slender, upwardly-directed branch arising from the upper margin near the base;

the margin or the ninth sternite is produced at its centre and truncate with an

excision on each side of the produced part.

Length of anterior wing, 3 5 mm.

UcanpDA: Fort Portal district, 5000 ft. (Edwards). 1 3 (type).

The species bears a close resemblance to O. kathia and to Potamoryza modesta

Barnard, a species found in South Africa. It differs from the latter mainly in

the presence of superior appendages which I cannot see in Dr. Barnard’s species.

20 RUWENZORI EXPEDITION

Oecetis thikanensis sp. n. (Figs. 56-59)

The example before me is in very bad condition so that Iam unable to give

any account of its general appearance.

Anterior wing with all the forks sessile and with various dark spots as indi-

cated in the figure. Apex of the anterior wing rounded, of the posterior, acute.

Genitalia 3. Apical margin of the ninth dorsal segment produced at its centre

in a small triangle; beneath are two large, shuttle-shaped superior appendages ;

Figs. 56-59.—-Oecetis thikanensis sp. n. §: 56. wings, 57. genitalia, dorsal, 58. lateral,

59. penis, lateral.

upper penis-cover broad, apex truncate; penis immensely broad and rounded,

occupying nearly the whole width of the segment; upper margin elevated in a

pair of large, rounded, asymmetric lobes, side by side, the whole arching down-

wards with a broad. lip at the apex; inferior appendages long, slender and

caliper-shaped with a strong angular projection on the upper margin towards

the base.

Length of anterior wing, ¢ 6:5 mm.

Kenya: Thika, 4500 ft., 23.x.1934 (Edwards). 1 g (type):

TRICHOPTERA 21

Setodes intricata sp. n. (Figs. 60-63)

Head black with a clothing of black hairs; antennae, basal joint bulbose

with a few silvery hairs on its sides; remaining joints dark chocolate with white

annulations disappearing towards the apex; legs ochraceous; anterior wings

deep fuscous with two well-indicated patches of white hairs, one large and

round, sometimes divided, extending to the costa near the apex, the other,

short and oblique towards the centre of the wing nearer the base; there are

also a few patches of short golden hairs in various parts of the wings.

Genitalia 3. Apical margin of the eighth dorsal segment straight; at the

Figs. 60—63.—Setodes intricata sp.n. 3: 60. wings, 61. genitalia, dorsal, 62. lateral, 63. ventral.

centre of the apical margin of the ninth tergite there is a small dorsal plate

which has the appearance of a pair of shuttle-shaped, brownish lobes, welded

together and leaving an excised apical margin; at the base of the plate on each

side is a minute, brownish spot, possibly an indentation of the segment; beneath

the plate, the apical margin of the segment is produced in two very long, finger-

shaped processes, apices curving slightly inward with their outer margins

heavily fringed, particularly towards the apices; from the margin of the segment

arise two long, slender spines which in the example under description cross

above the dorsal processes; penis large, elbowed downward, base very stout

and broad both from the side and from above; inferior appendages very large,

apical portions produced from somewhat oval bases which, from beneath,

slightly overlap them; inner margins of the basal part set with strong teeth;

22 RUWENZORI EXPEDITION

there is a slight triangular projection on the upper margin near the base, as

seen from the side.

Length of anterior wing, ¢ 6 mm.

Ucanpa: Fort Portal, Nyakasura, 4.xii.1934 (Edwards). 4 3g (type and

paratypes).

This species is very closely allied to Setodes didymata, described by Dr.

K. H. Barnard from South West Africa. It differs in the shape of the dorsal

plate and also in the form of the processes arising from the ninth dorsal segment.

HYDROPSYCHIDAE

Hydropsyche propinqua Ulmer (Figs. 64-66)

Hydropsyche propinqua Ulmer, 1907, Notes Leyd. Mus., 29: 21, fig. 32-33.

Head ochraceous with ochraceous hairs; oculi black; antennae ochraceous

with dark annulations which extend across the joints to give a suggestion of

diagonal streaks; maxillary palpi with the basal joint short, second twice as

Figs. 64—-66.—Hydropsyche propinqua Ulmer g: 64. genitalia, dorsal, 65. penis,

ventral, 66. genitalia, lateral.

long as the first and not quite as long as the third and fourth together; these are

approximately equal, fifth as long as the second, third and fourth together;

legs ochraceous; wings hyaline with yellowish irrorations.

Genitalia 3. Ninth segment with acute sidepieces and a raised upper part ;

there is a dorsal plate produced in two caliper-like processes separated from

TRICHOPTERA 23

each other by a nearly circular excision; small, angular projections about half-

way along the inner margins, slightly nearer the base; from above, there are

two rounded and fringed warts situated at the sides and projecting slightly

beyond the laterial margins; from the side, the plate is very broad at the base

with a rounded and sinuous upper margin, the caliper-shaped process appearing

as an extended finger with a barely indicated notch on the under side; penis

with a branch arising from its under surface; this branch, about midway,

carries brush-like bunches of hairs on each side; the apex is fringed and bifur-

cate, with the two forks curving upward and slightly outward beyond the apex

of the main stem; at the origin of this lower branch are two short, fringed side-

branches, directed towards the base; the apex of the main stem is much dilated

and bifurcate, the forks diverging, with a narrow excision between them at the

base and with widely excised upper margins and acute outer angles; inferior

appendages two-jointed, basal joint dilated at its apex, terminal joint very

short and, from above, bent over to show an obtuse angle on the outer margin;

apex of the joint truncate.

Length of anterior wing, ¢ 8 mm.

Type 3, with one inferior appendage wanting, is in the Berlin Museum and is

labelled N. Kamerun.

Through the courtesy of Dr. H. Bischoff, I have been able to give the above

description and figures of this species.

The species collected by Dr. Edwards so closely resemble H. propinqua that

it is only in minor points of detail that they are to be separated.

Hydropsyche namwa sp. n. (Figs. 67-71)

Head dark ochraceous; oculi black; antennae ochraceous with slightly

darker annulations; maxillary palpi, basal joint short, second rather long,

longer than the third which is longer than the fourth, fifth equal to the third and

fourth together; wings yellowish with numerous pale yellow irrorations and

blotches.

Genitalia g. Ninth segment: there is a raised upper part, narrowing to a

bilobed apex, lobes small and rounded; side-pieces produced in wide angles;

the segment is produced in a large dorsal plate, the sides of which are widely

separate in the apical part; from the side, the upper margin is dome-like with

the lower apical angle produced in a tailwardly directed finger with a well-

defined notch on its lower margin; there is a rounded and fringed wart on each

side; penis with a branch arising from its under surface; this branch is bifurcate

at its apex, from the side, bending upward beyond the apex of the main stem ;

at the origin of the lower branch are two minute transparent warts or processes ;

the apex of the main stem terminates in a truncate apex, the upper margin

24 RUWENZORI EXPEDITION

produced in two tailwardly and upwardly directed prongs; inferior appendages

two-jointed, from the side, terminal joint very short and broad, abruptly

narrowing to a sub-acute apex.

Length of anterior wing, f I11mm., 914mm.

RuweEnzor1: Namwamba Valley, 4500-10,200 ft. (Edwards). 40 3 @

(including type @).

Figs.. 67—-71.—Hydyropsyche namwa sp. n. g: 67. wings, 68. genitalia,

dorsal, 69. penis, dorsal, 70. ventral, 71. genitalia, lateral.

Hydropsyche bwambana sp. n. (Figs. 72-74)

General appearance of the insect, ochraceous; anterior wings light ochrace-

ous with darker transverse bands.

Genitalia g. Apical margin of the ninth, segment produced at its centre in

a large, excised plate, the excision being deep and rounded; the upper part of

the plate appears as two parallel lobes welded together, each with a distinct

rounded apex; from the side, the plate is directed downwards, apex bifurcate ,

TRICHOPTERA 25

upper fork broader than the lower; there is a rounded wart fringed with short

hairs on each side of the plate; penis with the usual ventral fork common to the

African species in the genus; stem constricted before the apex which is widely

dilated in two curving wings; on the upper surface, at the centre of the excision

Figs. 72-74.—Hydropsyche bwambana sp. n. g:

72. penis, dorsal, 73. ventral, 74. lateral.

formed by the two wings, is a pair of small, distally directed, triangular processes ;

on each side of the base of the ventral fork is a minute, basally directed plate,

fringed with short hairs.

Length of anterior wing, ¢ 10 mm., 2 12 mm.

RUWENzORI: Bwamba Pass (west side), 5500-7500 ft.; Kilembe, 4500 ft.

(Edwards). 24 $ 9 (including type 3).

This species, in its genitalia, closely resembles H. namwa, differing mainly

in details of the penis.

Hydropsyche wamba sp. n. (Figs. 75-79)

Head dark ochraceous, with a pair of still darker warts at the hind margin;

oculi black; antennae pale ochraceous, with darker annulations which extend

across the joints to givea suggestion of diagonal streaks; maxillary palpi, basal

joint short, about half the length of the second which is slightly longer than the

equal third and fourth joints, the second, third and fourth joints are together

slightly longer than the fifth; wings yellowish-fuscous with plentiful yellowish

irrorations after the usual Hydropsyche pattern.

Genitalia 3. Ninth segment: there is a raised upper part, narrowing to a

truncate apex; side-pieces strongly produced, apices rounded; the segment

26 RUWENZORI EXPEDITION

is produced in a very broad, bifurcate dorsal plate, each fork terminating in a

blunt finger as seen from above, the excision between being nearly square

towards the apex, the lower side of the square being extended in a deep V ;

from the side, the plate forms a broad triangle, shaped somewhat like the head of

a bird, the eye being indicated by a hairy wart; penis with a branch arising

from its under surface; this branch is bifurcate at its apex, the forks being long

and slender and with a U-shaped excision between; the apex of the branch is

Figs. 75-79.—Hydyropsyche wamba sp. n. g: 75. genitalia, dorsal,

76. lateral, 77. penis, ventral, 78. dorsal, 79. apex of ventral fork of

the penis.

bent up towards the apex of the main stem; at the origin of the lower branch

are two rather stout, fringed side-branches, hooked sharply backward; the

apex of the main stem from above is widely dilated with the apical margin

slightly concave, a small excision at the centre, the outer angles produced in

short pointed processes; inferior appendages two-jointed, basal joint dilated

at its apex where there is a darkly pigmented area; terminal joint very broad

from the side, short and sinuous from above, apex acute, fused to the lower

branch at its upper basal margin. .

Length of anterior wing, g 12 mm., 9 18 mm.

RUWENZORI: Bwamba Pass (west side), 5500-7500 ft. and Namwamba

Valley, 4500-6500 ft. (Edwards); Nyamgasani Valley, 6400 ft. (Buxton).

37 3d @ in all (including type Q).

TRICHOPTERA 27

Hydropsychodes Ulmer (Fig. 80)

Hydvopsychodes Ulmer, 1905, Ann. Soc. Ent. Belg., 49: 34-35, fig. 22.

Dr. Ulmer (in litt., 29.xi:1935) informs me that in H. albomaculata, the

median cell of the anterior wing is open and he kindly sent me a paratype of the

species which has enabled me to confirm the fact. I am re-figuring the anterior

wing here and to prevent any further confusion, I here designate albomaculata

as the genotype of Hydropsychodes.

Numerous African and Eastern species which have been placed in Hydro-

psychodes, will have to be transferred to other genera; the smaller insects with

Fig. 80.—Hydropsychodes albomaculata Ulmer 6,

anterior wing.

brownish, irrorated anterior wings can probably all be placed in Cheumatopsyche

Wallengren.

It is advisable to deal with the position here as, in Dr. Edwards’s collection

are two female flies, the one taken at Kilembe, 4500 ft., I7.xii.1935, and the

other in Mpanga Forest, Fort Portal, 4000 ft., 24.1.1935, which would, on the

mistaken understanding of the genus, have been placed in Hydropsychodes.

In the absence of the male, I prefer to leave these two insects unnamed.

They resemble in outward appearance, “ Hydvopsychodes”’ sexfasciata, but the

white markings which in that species extend to the membrane itself, are here

indicated by patches of white hairs and the membrane itself is almost unmarked.

It is necessary to transfer the following species, represented in the British

Museum collection, from Hydropsychodes to Cheumatopsyche:

afva Mosely, 1935, Ann. Mag. Nat. Hist., (10) 15: 229-230.

bimaculata Ulmer, 1930, Ann. Mag. Nat. Hist., (10) 6: 491.

chinensis Mart., 1930, Proc. Zool. Soc. Lond., No. 6: 80.

clavalis Mart., 1930, Proc. Zool. Soc. Lond., No. 6: 82.

diminuta Walker, 1852, Cat. Neur. Brit. Mus.: 115.

excisa Ulmer, 1930, Treubia, II: 395.

falcifera Ulmer, 1930, Ann. Mag. Nat. Hist., (10) 6: 482.

lateralis Barnard, 1934, Trans. R. Soc. S. Afr., 21 : 362-364.

lesnet Mosely, 1932, Mem. Est. Mus. Zool. Univ. Coimbra, 1: 64.

maculata Mosely, 1934, Eos, 9: 22-24.

mavmorvata Navas, 1922, Konowia, 1: 38.

obscurvata Ulmer, 1930, Ann. Mag Nat. Hist., (10) 6: 485.

plutonis Banks, 1913, Trans. Amer. Ent. Soc., 39: 239.

sovdida Hagen, 1861, Syn. Neur. N. Amer.: 290.

thomasseti Ulmer, 1931, Deuts. ent. Zeit., 1931: 18-19.

triangulavis Ulmer, 1931, Deuts. ent. Zeit., 1931: 19-20.

tvifida Mosely, 1935, Ann. Mag. Nat. Hist., (10) 15: 228.

zuluensis Barnard, 1934, Trans. R. Soc. S. Afr., 21: 360-362.

28 RUWENZORI EXPEDITION

Cheumatopsyche trifida Mosely

RUWENZORI: Bwamba Pass (west side), 5500-7500 ft. (Edwards). I 31 9.

Type in the British Museum from Katanga, Belgian Congo.

Diplectronella afra Mosely

Diplectronella afva Mosely, 1931, Ann. Mag. Nat. Hist., (10) 8: 202-205.

RUWENzoRI: Kilembe, 4500 ft., 17.xi1.1935 (Edwards). 1 ¢.

The type of the species was also taken in Uganda.

PSY CHOMYIDAE

Lype afra sp. n. (Figs. 81-84)

THERE is nothing in the general appearance of this insect to distinguish it

from the European species of Lyfe.

Figs. 81-84.—Lype afva sp. n. g§: 81. wings, 82. genitalia, dorsal,

83. lateral, 84. ventral.

Genitalia g. Terminal dorsal segment produced in a large plate tapering

to a truncate apex, from above; very deep from the side; on each side is a

large, tapering superior appendage, slightly longer than the plate; penis,

TRICHOPTERA 29

as is usual in the genus, with an upper process which is dilated at its apex with

a broad hump, towards its base, corresponding to the spine in the European

species phaeopa and veducta; inferior appendages two-jointed, basal joint broad

and short, about half the length of the apical joint which terminates in a clavate,

inturned apex; margin of the ninth sternite widely excised.

Length of anterior wing, ¢ 5:5 mm.

RUWENZORI: Namwamba Valley, 6500 ft. (Edwards). 1 3 (type).

Paduniella ankya sp. n. (Figs. 85-90)

Head dark ochraceous; antennae black and stout; palpi, maxillary, six-

jointed; labial, four-jointed; legs, anterior, femora ochraceous, tibia ochraceous

inside, fuscous outside; median, ochraceous excepting the tarsi which are

Figs.85—90.—Paduniella ankya sp.n. g: 85. wings, 86. palpi, 87. genitalia,

dorsal, 88. lateral, 89. internal structure, lateral, 90. genitalia, ventral.

fuscous; posterior, femora ochraceous, tibiae and tarsi fuscous; spurs fuscous,

2, 4,4; wings black; discoidal cell of the anterior wings open.

Genitalia g. Apical margin of the eighth dorsal segment straight; ninth

produced at its centre in a blunt triangle with a truncate apex; from the apical

30 RUWENZORI EXPEDITION

margin of this segment, there proceeds a pair of long, finger-like processes

simulating superior appendages but apparently pertaining to the ninth dorsal

segment; they form, in fact, a widely divided dorsal plate; beneath this plate

and covering a slender, arched penis, is a pair of long, curved spines, forming

an upper penis-cover; the penis itself is accompanied by a single fine sheath or

duct; inferior appendages with two branches, the outer long, slender and

sinuous with a slightly dilated and hood-shaped apex, the lower margin slightly

turned under, the inner arising midway, short, with a truncate apex and pro-

jecting outer angle; on the lower margin, towards the base (the inner margin

from beneath), is a large triangular keel fringed with four or five strong bristles.

In P. africana, the base of the penis and its complicated structures extend

back as far as the fifth segment; in ankya, they extend back only to the seventh.

Length of anterior wing, 3 4:5 mm.

RUWENZORI: Namwamba Valley, Kyanjoki, 6500 ft., 3-4.1.1935 (Edwards).

I g (type).

PHILOPOTAMIDAE

Dolophilus kyanus sp. n. (Figs. 91-94)

Head ochraceous; oculi black, antennae ochraceous with no evident annu-

lations; palpi and legs ochraceous excepting the tibiae and tarsi of the posterior

pair which are light fuscous; wings fuscous.

Genitalia g. The dorsal margin of the ninth segment is produced at its

centre in a triangular process with a very long produced apex; on each side of

it is a long and slender superior appendage; penis long, straight, bearing a fine

spine beneath; inferior appendages two-jointed, basal joint long with the apex

obliquely truncate; the second joint arises from the extreme outer angle and

is densely fringed with long, fine hairs; it has an enormously dilated and

thickened apex the distal surface of which is slightly rounded and _ bears

numerous parallel ridges fringed with short hairs; ninth sternite considerably

produced and excised at the centre of its apical margin to leave two rounded

lobes fringed with long bristles.

Length of anterior wing, ¢ 6 mm.

RuweEnzor1: Namwamba Valley, 6500 ft. (Edwards). 1 3 (type).

Chimarrha Leach

Of the genus Chimarrha there are three species represented in the collection,

one of them unfortunately by only a single 2 which has been left unnamed.

The other two are particularly striking insects, clara having uniformly pale

wings, and zorza, jet black, each wing bearing a large, white, round spot.

TRICHOPTERA 31

Chimarrha sp.

Included in the collection is a female Chimarrha species, entirely black in

colour with a long, slender ovipositor. It is possible that this insect may be

Figs. 91-94.—Dolophilus kyanus sp. n. 3: 91. wings, 92, genitalia,

dorsal, 93. lateral, 94. ventral.

new to science. It is not advisable to describe new species from the female sex

and accordingly, I leave this insect unnamed. It is labelled Bwamba Pass

(west side), 5500-7500 ft. (Edwards).

Chimarrha clara sp. n. (Figs. 95-98)

Head bright yellow; antennae black excepting the basal joint, which is

bright yellow; palpi fuscous; legs yellowish excepting the tibiae and tarsi of the

anterior pair, which are fuscous; abdomen yellowish, inclined to fuscous above ;

wings pale cream with very short, yellow pubescence; both with fuscous or

32 RUWENZORI EXPEDITION

nearly black, narrow fringes; the posterior wing is unusual in having no fork

No. 1; as all the other characters pertain to Chimarrha, the species must be

regarded as exceptional.

Genitalia 3. All the genital parts are yellow; apical margin of the ninth

dorsal segment produced at its centre in a tubular sheath with the apex obliquely

truncate as in zoria; on each side is a small superior appendage with a dilated,

Figs. 95-98.—Chimarrha clava sp. nN. $: 95. wings, 96. genitalia, dorsal,

97. lateral, 98. ventral.

clavate apex; penis membranous, furnished with two long, fine, deep black

spines lying parallel along the penis, the one longer and originating nearer the

base than the other; penis-sheaths wanting; lower penis-cover wanting;

inferior appendages broad at the base, trifurcate; from the side, upper fork

produced and slender, middle fork short, blackened and inturned, somewhat

rectangular, lower fork short and triangular, also slightly inturned, a long,

sinuous process to the ninth ventral segment, apex from the side, clavate, with

its lower surface furnished with a mass of short, black setae; a short, wide

process to the eighth ventral segment.

TRICHOPTERA 33

Length of anterior wing, ¢ and 2 9 mm.

RUWENZoRI: Kilembe, 4500 ft., 17.xii.1935; Bwamba Pass (west side),

7000 ft., 28-29.i.1935 (Edwards). 3 $ 1 (including type 3).

Chimarrha zoria sp. n. (Figs. 99-103)

Head, thorax and abdomen yellowish-orange; antennae black excepting

the basal joint which is yellowish-orange; palpi yellowish-fuscous; legs, all the

Figs. 99-103.—Chimarrha zoriasp.n. g: 99. wings, Ioo. genitalia, dorsal, ror. lateral,

102. penis and sheaths lateral, 103. genitalia, ventral.

femora yellow, tibiae and tarsi of the fore and median legs yellowish-fuscous,

of the hind legs, dark fuscous. Both anterior and posterior wings black, each

with a round white spot towards the centre, the veins at the centre of the spot

of the anterior wing, tinged with yellow.

III, IC

34 RUWENZORI EXPEDITION

Genitalia 3. Margin of the ninth dorsal segment produced at its centre in a

tubular sheath or dorsal process with the apex obliquely truncate and open

above, deeply excised beneath; through this tubular sheath projects a mem-

branous penis furnished with two black spines, one situated dorsally at the base

and the second towards the apex which is dilated and bears a small mem-

branous branch, on which this spine is seated; on each side of the dorsal process

is a small superior appendage with a rounded, dilated apex; there is a long,

slender, blade-like lower penis-cover and a pair of slender, divergent and bifur-

cate sheaths, the upper fork very slender, curving upward, the lower slightly

stouter and longer and with a blackened apex; inferior appendages from

beneath, with broad bases and caliper-like apices; these are directed upward

so that the apices are level with the dorsal process; there is a small, keel-like

process to the ninth ventral segment and a smaller one to the eighth.

Length of anterior wing, j 8 mm.

RUWENzoORI: Namwamba Valley, 6500 ft. (Edwards). 15 ¢ (including

type).

RHYACOPHILIDAE

Afragapetus gen. n.

This genus is closely allied to Pseudagapetus McLachlan and Myspoleo

Barnard. The main points of difference lie in the spurs which are 2, 4, 3, the

upper spur of the hind tibia being abnormally stout and curved in the J,

normal in the 9, the slightly dilated median leg of the 9 and the absence of

ventral processes in both sexes,

Genotype: A/fragapetus ungulatus sp. n.

Afragapetus ungulatus sp. n. (Figs. 104-108)

Head black with a clothing of golden hairs; antennae dark fuscous excepting

the terminal seven joints which are snowy white; legs and palpi yellowish;

wings dark brown, densely clothed with golden hairs.

Genitalia 3. Apical margin of the ninth dorsal segment excised; there is a

pair of membranous processes in the excision with strongly chitinized plates at

their extremities, one arising from the upper margin of the lower penis-cover

and twisting upwards round the penis, the other apparently arising from the

under margin of the terminal dorsal segment; penis retracted, its form obscure;

lower penis-cover very strongly chitinized, in the form of a blackened and slightly

downwardly directed trough or lip; inferior appendages large, oval, each with

TRICHOPTERA 35

a black tooth on its concave inner surface towards the apex and a pair, side by

side, towards the base; no ventral process.

Figs. 104-108.—Afragapetus ungulatus sp. n. g: 104, wings, 105.

abnormal spur of the g posterior leg, 106. genitalia, dorsal, 107. lateral,

108. ventral.

Length of anterior wing, d 4 mm., 9 4:5 mm.

RUWENzORI: Namwamba Valley, 6500—10,200 ft. (Edwards); Namgasani

Valley, 8—-gooo ft. (Buxton). 3 3 2 @ (including type 3).

HYDROPTILIDAE

Afritrichia gen. n.

Antennae broken in the type; ocelli present; spurs I, 2, 4; neuration fairly

complete as indicated in the figure.

The genus has some affinity with the European Stactobia.

Genotype: Afritrichia aurea sp. n.

Afritrichia aurea sp. n. (Figs. 109-112)

Head with a tuft of golden hairs between the deep black oculi; antennae

black; anterior wing covered with dense, golden plubescence without any

36 RUWENZORI EXPEDITION

conspicuous markings though the costal area and the apex of the wing are

inclined to fuscous; legs pale honey-colour.

Genitalia g. Apical margin of the terminal dorsal segment produced in a

heart-shaped, membranous plate beneath and beyond which may be seen a

straight penis and a large, trough-like lower penis-cover, the apex considerably

dilated; on each side of the penis and its cover, from above, is a large, triangular

ee oj

Figs. 109-112.—Afritvichia auvea sp. n. g: 10g. wings, 110. genitalia,

dorsal, 111. lateral, 112. ventral.

process; seen from the side, with a downwardly hooked apex; inferior appen-

dages arise from large, somewhat rectangular bases, the inner, apical angles

being produced in long fingers, slightly diverging towards their apices; from

the lower, basal, outer angle is a short, twisted downturned branch; a long

process to the seventh ventral segment.

Length of the anterior wing, 3 2:5 mm.

RUWENzORI: Namwamba Valley, 6500 ft. (Edwards). 1 3 (type), several 9.

Ugandatrichia gen. n.

Antennae about 43 jointed in 3g, broken in 9; ocelli present; spurs 0, 3, 4;

neuration complete as indicated in the figure.

Genotype: Ugandatrichia minor sp. n.

Ugandatrichia minor sp. n. (Figs. 113-116)

Insect deep black; anterior wings covered with erect, clavate, black hairs;

other characters are detailed in the generic description.

TRICHOPTERA 37

Genitalia g. From above, the ninth dorsal segment is excised to its base,

the sides of the excision sinuous; there is a membranous dorsal plate and a

pair of very strong, black, incurving appendages beneath it; penis straight with

a membranous outer sleeve; lower penis-cover broad, rectangular with a pair

Figs. 113-116.—Ugandatrichia minor sp.n. g: 113. wings, 114. genitalia,

dorsal, 115. lateral, 116. ventral.

of bristles at each apical angle; inferior appendages large, from below, finger-

shaped, nearly parallel, bearing on the upper surface, towards the inner apical

margin, a short, pointed, deep-black spur; a short, keel-like process to the

seventh ventral segment.

Length of the anterior wing, gj 2 mm.

KENYA: Thomson’s Falls, 7500 ft., N. of Nakuru, 6.111.1935 (Edwards).

I 3 (type).

Ugandatrichia nigra sp. n. (Figs. 117-121)

Wings black, densely covered with black, clavate hairs. Neuration as in

U. minor.

Genitalia ¢. Ninth segment very deep; from above, excised widely to its

base, much produced and rounded from the side, deeply excised from beneath;

38 RUWENZORI EXPEDITION

from above, on each inner margin of the excision, at the base, is a large, rounded

bulge of which the distal border is armed with a clump of stout hairs; the basal

part carries another smaller bulge which is covered with minute, pointed setae;

in the excision is a broad, membranous dorsal plate, apical margin excised;

penis of the usual pattern in the Hydroptilidae, consisting of a long, fairly

straight tube through which passes a central duct and with a sheath curled

round the middle with its pointed apex free; beneath the penis is a broad lower

penis-cover, apical margin with a wide excision, the apical angles produced in

Figs. 117-121.—Ugandatrichia nigra sp. n.: 117. $ genitalia, dorsal, 118, lateral, 119. penis,

120. genitalia ventral, 121. 2 genitalia, ventral.

blackened spurs; inferior appendages from beneath, appear as a pair of legs,

widely straddled, with the toes of the foot blackened and directed inwards,

a small wart armed with a few bristles on each heel; a short, keel-like process

to the seventh ventral segment.

Genitalia 9. On the under surface of the ninth ventral segment is a small,

bifurcate process, the forks short, rod-like and divergent; margin of the segment

fringed with widely-separated, long hairs; a short, keel-like process to the

seventh ventral segment.

Length of the anterior wing, 3 4-5 mm., 25 mm.

RUWENZORI: Namwamba Valley, 6500 ft. (Edwards). 1 $ (type). 2 9.

TRICHOPTERA 39

Ugandatrichia acuta sp. n. (Figs. 122-123)

I am departing from my usual practice and am giving a name to this female

insect as I think the characters provided by the shape of the wings (if not

aberrant) and the pattern of the genitalia are sufficient for its future recognition.

The insect is black all over; anterior wings densely covered with upright,

clavate hairs; the apices of both wings are very considerably produced and

pointed; in this respect, they differ from the other two species described in the

122

Figs. 122-123.—Ugandatrichia acuta sp. n. 9: 122. wings, 123. genitalia, ventral.

genus although the neuration conforms excepting in so far as it has been modi-

fied by the produced apices.

On the surface of the eighth ventral segment are two oval patches covered

with minute, scale-lke hairs and on the ninth is an urn-shaped design with

projecting knobs on each side about midway along the outer margins; a small

keel-like process to the seventh ventral segment.

Length of anterior wing, 2 4-5 mm.

KENYA: Chania Falls, 4000 ft., near Nairobi, 3.x.1934 (Edwards). 1 9

(type).

TABLE OF SPECTES

Sericostomatidae

Crunoeciella Ulmer

brunnea Ulmer, 3

Goerodes Ulmer

edwardst sp. n., 5

excelsior Navas, 6

inferior Navas, 7

nudata Navas, 10

Calamoceratidae

Anisocentropus McLachlan

usambarensts Ulmer, II

Leptoceridae

Leptocerus Leach

asanus, II

sp., 12

Leptocerina Mosely

talopa sp. n., 13

Triaenodes McLachlan

legona sp. n., 14

wambana sp. n., 15

Triaenodella Mosely

hastata Ulmer, 16

Oecetis McLachlan

kathia sp. n., 16

angustipenms sp. n., 17

portalensis sp. n., 19

thikanensis sp. n., 20

Setodes Rambur

intricata sp. n., 21

Hydropsychidae

Hydropsyche Pictet

propinqua Ulmer, 22

NamMwA SP. N., 23

bwambana sp. n., 24

wamba sp. n., 25

Hydropsychodes Ulmer

albomaculata Ulmer, 27

Cheumatopsyche Wallengren

afra Mosely, 27

bimaculata Ulmer, 27

chinensis Martynov, 27

clavalis Martynov, 27

diminuta Walker, 27

excisa Ulmer, 27

falcifera Ulmer, 27

lateralis Barnard, 27

lesnet Mosely, 27

maculata Mosely, 27

marmorata Navas, 27

obscurata Ulmer, 27

plutonis Banks, 27

sordida Hagen, 27

thomassett Ulmer, 27

triangularis Ulmer, 27

trifida Mosely, 27, 28

zuluensis Barnard, 27

Diplectronella Ulmer

afra Mosely, 28

Psychomyidae

Lype McLachlan

afra sp. n. 28

Paduniella Ulmer

ankya sp. n., 29

Philopotamidae

Dolophilus McLachlan

kyanus sp. n., 30

Chimarrha Leach

Sp:, 32

claya sp. N., 31

zorla Sp. N., 33

Rhyacophilidae

A fragapetus gen. n., 34

ungulatus sp. N., 34

Hydroptilidae

Afritrichia gen. n., 35

aurea Sp. N., 35

Ugandatrichia gen. n., 36

minor sp. N., 36

nigra sp. N., 37

acuta sp. N., 39

2) PRESENTEB

15 MAR 1939

Plate I

RUWENZORI

Stream in Namwamba Valley near Butama, 6000 feet

Habitat of Chimarryha zoria Mosely

RUWENZORI EXPEDITION—VOL. III

Plate II

Thomson’s Falls, north of Nakuru, Kenya

Habitat of Ugandatrichia minor Mosely

RUWENZORI EXPEDITION—VOL. III PLATE Fi

fae ~ Lan 7 Mi F ‘ ' I > ,

{ y : ; =f ; ’ | 7h wens bie 20 Ee / ;

af ; a 1 F ae / : / ; y F

it h { iy " ff f ‘e

: t : i 4 , : ) i !

’ = ‘ \ j ul W ‘ ;

{ ; 1 . ra : x : Y c i ‘ ' }

i a }

é i ¥ t i 7 é /

i pony | ‘ j te :

’ i ' i S we

i 1 he

‘ y : Le " ae Peary

i 7 in z ‘

: 4 i ' ; ‘ a ny

ig i > i

i ’ i) 3 = ad

: 5 i ra ; ry ) e

+ ae. fe ‘ - . : ¢ . i " i 7

Keay ; ; . ,

- : 1, i PA =

e4 , =

J ‘ 4 /

= > \

oes =

; > ‘ (o> 7

‘ ad Lhe =

= . f 2 yr,

é x \ \

)

j j ‘ ; ' :

i ; 3. = ©

j f 5

eS ' 7 1 fare al

; ; P eee

" * - I 2h : :

7c bs Eyl " a 7

asf eay « , 4 ‘ =

z - { te 2 ' f

~ = ! 1 . + ft

: / ss \ ‘i a P x

; 7 j 2 1

“ i a :

= ‘ z = . id : F ij

“i x \

‘ - 1

~ f 4 1 i & a = ”

: cam ~ sf \ ' 4 1 ct

’ fl 4) = ’

> f 7

= . = = 7 cs

: u > 7 7 -

“ = ( ‘

) = 4

~ \3 * 5 !

= 5 ! : oe Wj 1

. ; ' . ri F Cee ig

Ty aes 2 a

. ise - *

E = 1 4 1 »

rf x t

ae “4

= 7 “), Y ae 7

ts < ae '

: = :

= “fp Z ; a

c i ™ y 5

2 ( ’ : = ok ;

q cay }

’ > Tas J - rf

— s 1 e

i 7 vie =

; 4 ) ‘

: bet ~

' > : ‘

; = * z

Fe = P ie ;

te ri