4. MUSCIDAE: B.—COENOSIINAE

By F. I. van EMDEN

(Imperial Institute of Entomology)

TuHIs second part of the report on the Muscidae of the Ruwenzori Expedition

deals with the subfamily Coenosiinae, of which almost goo specimens of 93

species and subspecies were collected. Together with this material numerous

collections received from East African countries (Kenya, Uganda, Sudan,

Tanganyika) by the Imperial Institute of Entomology, and the collections and

accessions of the British Museum were studied. The Muscidae collected by

Dr. Scott and Mr. J. Omer Cooper in Abyssinia were worked out at the same

time and the new species in this collection have been incorporated in the keys

except for the genus Microcalyptra, of which the Ruwenzori Expedition found

only a single specimen and which, therefore, will be keyed in the report on

Dr. Scott’s expedition to Abyssinia.

Our knowledge of these flies is much less advanced, as far as the Ethiopian

region is concerned, than that of the Muscinae and Stomoxydinae. Striking

testimony to this is the fact that Speiser (1910, Sjéstedts Kilimandjaro—Meru

Exp. 10. Diptera: 165) listed only a single species of this subfamily from the

Kilimanjaro and that Séguy’s recent paper (1938, Miss. scient. Omo 4, Zool.:

277) contains not more than seven species of it. It was, therefore, deemed

advisable and proved even inevitable to refrain from any zoogeographical

conclusions but to present complete keys to the material studied. These keys

are the only means of showing fairly exactly the author’s concept of the

previously known species. This is so much the more necessary as the existing

keys of Stein, Malloch? and Curran? often lead to the same name for species

which are not at all closely related and, moreover, include between them only

64 (Malloch alone 51) species, less than half the number of species accounted

for in the present keys, which cover 158 species and subspecies, just over half

this number being new to science. As the number of described Ethiopian species

(incl. Seychelles and Madagascar), not represented in the various collections

studied, is not very large (36 species, of which 21 could be incorporated in the

key from descriptions), all the described species of this region, as far as possible,

have been included.

1 7913, Ann. Mus. nat. Hungar. 11: 565-567 (Coenosia s. lat.); 1919, Arch. Nat.-gesch.

83 Ar: 88—1or (genera).

2 7922, Ann. Mag. nat. Hist. (9)10: 380-381 (Pygophora), 575-580 (genera of Coeno-

siinae, Caricea); 1924, l.c. (9)13: 411-413 (Atherigona excisa and setifemuy groups).

3 4935, Amer. Mus. Novit. 776: 1-12 (Caricea).

II, 4

g2 RUWENZORI EXPEDITION

Unfortunately my attempts at seeing the types failed owing to the imminence

and outbreak of the war. Except for the types in the Entomological Museum

of the University of Kansas, which through the kindness of Prof. H. B. Hunger-

ford I was able to study at an earlier stage of this paper, and for the types in the

British Museum (Natural History) and in Mr. J. E. Collin’s collection, no types

were therefore checked. My thanks are due to Prof. Hungerford and Mr.

Collin as well as to Mr. G. C. Varley of the University Museum of Zoology,

Cambridge, who informed me that the species described by Stein from the

Cambridge Museum have never been returned and seem to be lost. Dr. C. H.

Curran was kind enough to run down his species, so far as they are in the

American Museum of Natural History, in my key, and I wish to thank him

for the important help afforded in this way. Thanks for information on

several native names of plants are due to Sir Guy A. K. Marshall and Mr.

J. E. Dandy. For the conscientious execution of the drawings I would express

my thanks to Miss D. Fitchew (hypopygial prominences and trifoliate processes

of Atherigona, slightly shaded and unshaded), Miss M. Mackay (hypopygial

prominences and trifoliate processes of Atherigona, shaded), and Miss O. F.

Tassart (remaining figures). Finally I thank Dr. F. W. Edwards and Sir Guy

A. K. Marshall sincerely for their original suggestion that I should undertake

the study of this rich material.

As in the first report of this series, the abdominal segments are counted in

the usual way, ie. the first two connate segments as one, so that the last

entirely free tergite of the female is the fourth. Genera and forms mentioned

in this report but not represented in the material of the Ruwenzori Expedition

are put in square brackets. Species incorporated from literature are marked

with an asterisk, those incorporated on the authority of Dr. Curran or confirmed

by him with a cross.

The abbreviations used are the same as in my first report, with the following

additions:

m-m posterior cross-vein

ph posthumeral seta.

The types of the new forms are in the British Museum (Natural History).

CHARACTERS AND CLASSIFICATION OF THE COENOSIINAE

The Coenosiinae as understood in this paper are defined as Muscidae with

the following characters: Inner margin of lower calyptra more or less strongly

diverging from sides of scutellum, apex rounded or almost triangularly pointed,

not truncate (distinction from Muscinae). Sixth vein (ci, + 2¢a according to

Comstock, cw, + a, according to Hendel) not reaching margin of wing even

as a fold, ventral surface of scutellum not bearing fine, short, erect hairs

MUSCIDAE: B.—COENOSIINAE 93

(distinction from Anthomyiinae).! Parafacialia bare (haired in Lispinae). Ptero-

pleura bare? (with a tuft of hairs in Lispinae, haired in Stomoxydinae and

others). Frons of both sexes of subequal width (sometimes slightly narrower

Fics. 1-4. 1.—Coenosia hetervocnemis sp. n. 2.—Orchisia costata Meig. 3.—

Amicitia lucens sp.n. 4.—Cayvicea argentipes sp.n. Heads in three-quarter view.

in female), more or less broad (at least a seventh the width of the head at the

narrowest point), without cruciate interfrontal or proclinate orbital setae and

1 The character of the sixth vein would also separate the Coenosiinae from the Gastro-

philinae, but there is much evidence of the latter’s belonging to the Acalyptrata (second

antennal joint, position of abdominal spiracles, size of calyptrae).

2 The male of the Patagonian genus Raymondomyia Mall. is unknown. Besides the few

short hairs of the pteropleura the female shows several other characters which are unusual

in the Coenosiinae, and only the discovery of the male will settle the systematic position

of this genus.

94 RUWENZORI EXPEDITION

with only one (Figs. rand 4) or two more or less strong and conspicuous reclinate

frontal setae, of which the anterior one is always longer, if two are present

(Figs. 2-3), the reclinate setae always conspicuous, well separated and never

bent outward (distinction from Phaoniinae and Fanniinae).

To these characters, which are strictly diagnostic, the following important

distinctions may be added, though some of them are only valid for the species

of the Ethiopian region: Mentum rather heavily chitinized but shorter than

the head and only slightly tapering to apex. Never more than one strong

(but sometimes two weak or a moderate and a strong) prst dc and three strong

(sometimes 2—3 weak and two strong) Post dc. stpl usually in an almost equilateral

triangle, generally three present. Lower prostigmatal seta usually curved down-

ward. Apical half of subcosta distinctly curved forward (not straight as in

Fanniinae), seventh vein (3°a according to Comstock, a, according to Hendel)

gently and slightly curved forward (not sigmoid and strongly curved forward

round apex of sixth). Costal spine usually indistinct. Veins always wholly

bare (except, of course, costa).

As far as known the adults are predators on other insects, and this would,

according to Séguy (1937, Gen. Ins. 205: 224) also more or less apply to the

somewhat aberrant genus Atherigona, about which he writes that they are

occasionally zoophagous and eat small insects with soft integument, which they

catch in flying. I have no evidence to the contrary, but Malloch (1929, Ins.

Samoa 6: 157) states that the adults of Athevigona are not predacious. In the

material worked out on the present occasion I saw six species of Coenosiinae

with their prey: Caricea inaequivitta Mall. with the prey Nephrotoma sp., a

Tipulid, C. calopoda Bezzi with Trentepohlia sp., a Tipulid, C. punctigera Stein

with Culex fatigans Wied.?, C. pachypoda Big. with an indeterminable Dolicho-

podid, C. hargreavest Emd. with Sciara sp., and Pygophora acromiata Speis. with

Sciara flavoscutellata End.? The prey has kindly been identified by Dr. F. W.

Edwards. Nematocera play an important part in the records of the prey

of Coenosiinae found in literature,’ and in this respect the group must be

regarded as essentially beneficial. Little is known of the biology of the larvae,

which seem to be largely scavengers or predators, but the species of Atherigona

s. str. are phytophagous (see p. 102). This fact in combination with the two

small distant prst de and more than three (though mainly small) post dc tends

to give that genus a somewhat isolated position, though I am convinced that

it is really closely related to the Coenosiinae.

The subfamily is usually divided into two groups: Coenosiini and Chelisiini,

the latter being characterized by two prst dc. Chelisia itself is, however, hardly

related to the other genera in this aggregate, as it presents cruciate interfrontal

setae, proclinate frontal setae, fine erect hairs on the ventral surface of the

1 See e.g. A. M. Evans, 1930, Trans. Ent. Soc. London 78: 325; B. M. Hobby, 1931,

Proc. Ent. Soc. London 6: 13; A. Cuthbertson, 1937-1939, Trans. Rhodesia Scient. Assoc.

35:3 215- 36; LTO}, 37%_139-

MUSCIDAE: B.—COENOSIINAE 95

scutellum, two subequal, strong frst dc, a distinct though small costal spine,

more richly setose mid and hind tibiae, and a sixth vein which almost reaches

the wing-margin as a fine fold. It should in my opinion be united with the

Anthomyiinae, and Mr. Malloch agrees in a letter with this view. Collin (1927,

Ent. Mo. Mag. 63: 130) also enumerates Chelisia among the genera of the

“Hylemyta-Chortophila-group.”” Practically all’ the Coenosiinae have smaller

setae or hairs in the dc rows between the strong setae, one of them rather close

to and in front of the first dc. This hair may be shorter or longer and is very

variable even within one genus. Though there is a distinct tendency for this

hair in the aggregate around Coenosia and Caricea to become small and

inconspicuous and in the group around Lispocephala to remain quite conspicuous,

its development is very fluctuating in the different species of Pygophora,

Lispocephala and others. The presence of one or two reclinate frontal setae

affords a much better and certainly very natural character for the primary

classification of the Coenosiinae. Malloch even goes so far as to unite the group

with two reclinate frontals with the Phaoniinae (1934, Dipt. Patag. S. Chile

7: 177, 209; personal communications) with which they really seem to link the

Coenosiinae. I do not like this idea, as Pygophora especially is very closely

related to the Coenosia group, and I am, on the other hand, not fully convinced

that the character of a single reclinate seta may not occur in any Phaoniinae.

It may be mentioned that in two specimens seen the second reclinate seta is

absent on one side, but present on the other. It is hardly desirable to name the

two groups of Coenosiinae, and the Chelisiini will have to disappear, as far as

their type-genus is concerned, into the synonymy of the Anthomyiinae, and as

far as their main contents is concerned, into that of the Coenosiinae.

KEY TO THE GENERA OF COENOSIINAE OF THE ETHIOPIAN REGION}

1(14). One pair of reclinate frontal setae (figs. 1, 4). Hind tibiae usually without

a pd and never with two pd setae.

2(13). Hind tibiae without an outstanding a bristle near middle, but usually? with

an av bristle inserted well distad from the ad (Fig. 64, p. 155) or with two

ad or av.

1 The genus Tenwicosta Stein from Madagascar has been omitted from the key, as no

specimen was available. It is characterized by only two strong post dc and the costa ending

at or slightly beyond 74,5. The mid-tibiae with a subbasal p seta, which is very long in

the male (evidently besides the submedian ad and pd setae),

2 The av is absent in both sexes of Anaphalantus (usually) and a few Athevigona and in

males of some Microcalyptva and a few Coenosia, and the a in the males of a few Caricea

(vittata, xenia, hippelates, nigritibia, flavivibrissata?, avistalis, and gilvicova). In these males of

Caricea, which could otherwise, of course, not be distinguished by the key from the males

without av of the genera mentioned, the pyst dc are strong (whereas they are inconspicuous

in Atherigona), the thorax and abdomen are never largely glossy black (distinction from

Microcalyptva and Anaphalantus); if a pd is present on the hind tibiae the upper prostig-

matal seta is much weaker than the propleural (distinction of Cayicea vittata from Coenosia

heterocnemis), and if no pd is present on hind tibiae, either at least the front coxae are

almost entirely yellow (distinction of Caricea flavivibrissata, xenia, aristalis and gilvicoxa

from Coenosia aurvifacies), or the front femora are more or less black (distinction of Caricea

hippelates, nigvitibia and some specimens of inaequivitta from Coenosia aurifacies).

T2(1I).

13(2).

T4(1).

15(22).

16(17).

RUWENZORI EXPEDITION

Pyst dc very small, but little longer than the setulose hairs. Front femora

with but one or two, seldom without any, outstanding pu subapical bristles.

Lower calyptra well developed, strongly projecting beyond upper one.

Atherigona Rond., p. 97

1+3 or 2 dc distinct and much longer than the setulose hairs of the thorax.

Front femora with a complete series of pv bristles, or seldom without

any distinct outstanding bristles, but then the lower calyptra hardly

projecting beyond the upper one.

Hind tibiae with two av and two ad setae near middle. Mid-tibiae with a

strong av seta (in the Ethiopian species). 1+ 3 dc. Two pairs of

scutellar bristles. Outer vertical setae strongly developed. [Schoenomyza]

Hind tibiae with not more than one av (and one or two ad). Mid-tibiae

without an av seta.

1 + 2strongdc; if with 1 + 3 (in Anaphalantus three post pairs present, but

the first of them usually very small), then the front femora without any

outstanding pu setae towards apex. Lower calyptra hardly project-

ing beyond upper one. Thorax with the humeri, pteropleura and two

more or less conspicuous vittae shining black, abdomen black and

shining, not or slightly dusted. Front tibiae with a pv bristle.

Scutellum with but one pair of strong setae. Face and antennae not longer

than frons, the latter rounded in profile, hardly projecting beyond eyes.

Front femora with a complete row of pv bristles, front tibiae of male

not feathered. . c . 5 Microcalyptra Stein, p. 143

Scutellum with two pairs of pees Bees and antennae much longer than

frons, the latter strongly projecting beyond eyes at base of antennae.

Front femora without any pv setae, front tibiae of male feathered,

those of female with a row of short spinose setulae on ventral surface.

[Anaphalantus Loew]

1+ 3 strong dc. Front femora with distinct pu setae up to apex. Thorax

entirely covered by dust though the latter often forms darker vittae.

Scutellum with two pairs of setae.

. Hind tibiae with two ad and one av bristles, mid-tibiae with two p bristles.

Frons strongly projecting beyond eyes in profile. Wings rather broad.

Lower calyptra large, about twice as long as the upper one, strongly

projecting beyond it. Abdomen very slender in male, depressed.

Parafrontalia finely haired on anterior half. . Andersonosia nom. n. p. 143

Hind tibiae with one ad and with or (some males) without an av bristle

(Fig. 64, p. 155). Frons gate projecting beyond eyes. Wings

narrower. Coenosia Meig. p. 145

Hind tibiae without; an av but mn one outstandine a and one ad bristle

near middle (usually close to each other), without or with a pd seta

(Figs. 65-67, p. 155). The a seldom missing, see note (2). Caricea R.-D. p. 154

Two pairs of reclinate orbital setae (Figs. 2, 3). Hind tibiae with one or

two pd (except in Spanochaeta with only a single row of acr), though

these may be small, never with an a and ad seta closely placed.

Scutellum with four strong bristles, the lateral pair as long as or longer than

the first or second post dc. Hind tibiae with two strong or small pd,

two strong ad, and one distinct av. Antennae, palpi, and legs never all

entirely black.

Frontal triangle not passing middle of frons, the latter much longer than

17(16).

18(21).

19(20).

20(19).

21(18).

22(15).

23(26).

24(25).

25(24).

26(23).

This genus, which is no doubt the most difficult one of the Coenosiinae,

MUSCIDAE: B.—COENOSIINAE

wide and narrowed behind. Hind tibiae with two strong pd, front

tibiae with a strong submedian fu bristle, mid-tibiae with two (always?)

pd setae and without an ad seta. Arista long-plumose to middle. 1-+-3

dc, the other setulae in the dc line shorter than the acy, rarely another

smaller dc in front of the prst dc. : ; . Pygophora Schin.,

Frontal triangle reaching anterior margin of frons, the latter not much

longer than wide, usually subparallel (seé Fig. 2). Hind tibiae with

two usually rather small pd setae, front tibiae without a submedian

bristle, mid-tibiae with one pd.

Costa reaching m. 2-+3 dc, as a bristle in front of and rather close to the

prst dc is longer than the prst acr.

Arista with rather long hairs up to tip, bare on ventral side in male?.

Pectiniseta Stein,

Arista with more or less short hairs, or longer hairs only in basal half or

third, the apical half bare! . : :

Costa reaching to just beyond 74,5. 1-+3 dc, the small setulae in the dc

line slightly shorter than the acy. Arista short-haired from base to

apex, the hairs shorter than the width of the third antennal joint.

Lower calyptra not much longer than the upper one. Brevicosta Mall.,

Scutellum with only the apical pair of setae distinct, the lateral pair missing

or less than half as long as the first or second post dc. Arista short-

plumose to almost bare, the hairs not longer than the third antennal

joint wide.

Front tibiae without a submedian pv seta. Frons more or less rounded in

profile, not or slightly protruding at base of antennae. Antennae, palpi,

and legs yellow. Dorsum of thorax not shining, entirely dusted, not

vittate. Arista not geniculate. Well developed prst dc and three strong

post dc present.

One pair of prst dc. Hind tibiae with one small pd and one small and one

long ad (and one small av) setae. Abdomen short. Frontal triangle

reaching anterior margin (Fig. 2). Lower calyptra rather short.

Orchisia Rond,

Two distinct pairs of prst dc, the anterior one much smaller. Hind tibiae

without a pd, and with at most one ad (and one av) setae. acy very

distinct, in one single row. Abdomen very elongate, yellowish trans-

lucent on first and second segments. Frontal triangle hardly reaching

middle of front. Lower calyptra twice the length of the upper one.

Spanochaeta Stein,

Front tibiae with a strong submedian pu seta. Frons protruding in profile

above base of antennae, by the width of the third antennal joint or

more (Fig. 3). Dorsum of thorax with a pair of broad dark vittae.

Frontal triangle not passing middle of frons. Hind tibiae with one or

two pd setae. 5 , : 5 a - . Amicitia gen. n.,

Atherigona Rond.

Pp:

. Lispocephala Pok., p.

Pp.

242

246

247

has

three very well defined groups, the first two of which are recognised in the

1 Lispocephala australis Mall.

to Pectiniseta. The male is unknown.

2 has a long-plumose arista and would run in this key

98 RUWENZORI EXPEDITION

male sex among other characters, by the lack of the long-stalked trifolzate

process of the superior forceps (Fig. 4A) which is invariably present in Atherigona

s. str. In pinned specimens the process is sometimes exposed, and almost

always part of the stalk is visible. After relaxing the specimen for one night

the trifoliate process can usually be freed by bending the genitalia dorsalward

with the point of a pin, while supporting the dorsum of the abdomen with the

head of another pin. All the species which possess this process have modified

3 (4) 4 (5) hp.

ZO tbs WZ,

ie \

o8) GED)

2(3)

Fic. 44.—Abdomen of Atherigona trapezia sp.n., extended; (a) oblique dorsal view,

numbers indicate the tergites, (kp) hypopygial prominence, (#p) trifoliate process,

—(b) oblique ventral view, numbers indicate the tergites and the ventrites referred

to them, the arrow shows the pocket into which ¢p fits, (ft) forceps inferior, (fs)

forceps superior, (p) penis, paramera blackened.

palpi in the male sex, as shown in Fig. 60, and most of them have, in the same

sex, on the fifth abdominal tergite an unpaired knob-shaped, bi-lobed, bifurcate,

or tridentate process, the hypopygial prominence (Fig. 4A), which is directed

beneath and backward. Whereas the species of the setifemur and excisa groups

are neither numerous nor very difficult to separate, those of Atherigona s. str.

MUSCIDAE: B.—COENOSIINAE 99

Ney,

/la,

16.

Fics. 5-17. Hypopygial prominences of Atherigona, posterior view, unless other-

wise stated. 5.—conigeva sp. n. 6.—laevigata Loew, from Doukham, also for

bimaculata Stein. 7.—pedunculata sp. n. 8.—tvuncata sp. 0D. 9.—aurifacies

sp. n., oblique view in posterior and lateral direction. 10.—binubila sp. n., from

Nairobi. 11.—tviangularis sp. n. (a) posterior, (b) ventral view. 12.—gviset-

ventris sp.n. 13.—marginifolia, ventral view. 14.—fevvuginea sp. n., ventral view.

15.—vubyicornis Stein, from Salisbury. 16.—lineata Ad., from Salisbury. 17.—

_ piceiventris sp. n. (in the ventral views the anterior side, i.e. the side facing the

genitalia, is above in the drawing).

Fics. 18-30. Hypopygial prominences of Athevigona, Figs. 18-22 posterior,

23-30 ventral view. 18.—lineata ugandae ssp. n. 19.—indica infuscata ssp. n.,

from Bukalaca. 20.—yvuficoynis Stein, from Mabungo Camp. 21.—tomentigera

sp. n. 22.—1tvidentata Mall. 23.—budongoana sp. n., from Budongo Forest.

24.—astey sp. n. 25.—longifolia sp. n. 26.—perfida Stein, from Butandiga-

Bulambuli. 27.—bedfordi sp. n. 28,—gilvifolia sp. n., from Ibadan. 29.—

steeleae sp. n., from Kilembe. 30.—trapezia sp. n., from Mt. Muhavura.

30,

RUWENZORI EXPEDITION

Fics. 31-39. Trifoliate processes of Atherigona, ventral views, unless otherwise

stated; in Figs. 33, 34, 35, 38, and 39 the lateral lobes are not drawn in natural,

but in a plane position. 31.—hancocki sp. n. 32.—longifolia sp. n., lateral view.

33-—binubila sp. n., from Nairobi, (b) median lobe in lateral view. 34.—bimacu-

lata Stein, from Silhouette, (b) median lobe in lateral view. 35.—haplopyga sp. n.,

(b) lateral view. 36.—lineata Ad., from Salisbury, (b) lateral lobe, interior surface

in plane position. 37.—tomentigera sp. n., (b) lateral lobe, exterior surface in

plane position. 38.—gviseiventris sp. n., (b) median lobe in lateral view. 39.—

lineata ugandae ssp. n.

406. 40a.

Fics. 40-49. Trifoliate processes of Atherigona, ventral views, unless otherwise

stated, lateral lobes not drawn in natural but in a plane position. 40.—aurifacies

sp. n., (b) median lobe in lateral view. 41.—indica infuscata ssp. n., from Buka-

laca. 42.—1triangularis sp. n., (b) front view of tip of median lobe. 43.—tridentata

Mall. 44.—marginifolia sp. n., (6) median lobe in lateral view. 45.—truncata

sp. n., lateral view. 46.—astey sp.n. 47.—piceiventris sp.n. 48.—tvapezia sp. 0..,

from Mt. Muhavura, (b) median lobe in lateral view. 49.—pedunculata sp. n.

MUSCIDAE: B.—COENOSIINAE IOI

are both to a very high degree. I assumed at first that the colours as well as

the shape of the hypopygial prominence presented a considerable amount of

variation. Finding, however, that in similar forms a certain type of hypopygial

prominence was coupled with a certain type of coloration and moreover with

a particular type of trifoliate process, I had toadmit a great number of “‘species’’.

Only a detailed study of material of a greater area will show to what extent

these forms may eventually be regarded as subspecies. As far as my evidence

Fics. 50-59. Trifoliate processes of Atherigona, ventral views, unless otherwise

stated; lateral lobes not drawn in natural but ina plane position. 50.—vruficornis

Stein, from Mabungo Camp, (b) median lobe in lateral view. 51.—vubricornis

Stein, from Salisbury, (b) median lobe in lateral view. 52.—gilvifolia sp. n., from

Ibadan, (b) median lobe in lateral view. 53.—conigeva sp. n. 54.—fervuginea sp.

n. 55.—bedfordi sp.n., from Shendi. 56.—laevigata Loew, from Doukham. 57.—

perfida Stein, from Butandiga-Bulambuli. 58.—steeleae sp. n., from Kilembe.

59.—budongoana sp. n., from Budongo Forest, (b) median lobe in lateral view.

goes, certainly some of the species described in the present paper have a wide

distribution and show no difference in the essential characters in such distant

places as Nigeria and Uganda, or Morocco and Uganda, or Aden and Kenya.

Unfortunately, in the females of this genus, which are evidently more

common than the males, no structural characters have been found either by

previous authors or by myself, as far as Atherigona s. str. is concerned, whereas

structural characters for the distinction of the females of the two other groups are

102 RUWENZORI EXPEDITION

indicated in the key. In order to enable the females of Atherigona s. str. to be

recognised as approximately as possible, colour-characters have largely been

used in the remaining part of the key, as a separate means for the identification

of the males is provided in Figs. 5-30 and 31-59, which represent the hypopygial

prominences and trifoliate processes. This part of the key can only be regarded

as preliminary to a more satisfactory one, based on structural characters of

both sexes.

The biology of the species strongly confirms the validity of the groups founded

on systematic characters. The larvae of the species of the setfemur-group

evidently feed on decaying organic matter, e.g. horse-dung (aberrans) and

“cocoon of Oryctes”’ (pallidipes). In the exctsa-group various kinds of decaying

fruits seem to be the main food, such diverse fruits being recorded on specimens

in the B.M., Imperial Institute of Entomology, and in the literature, as banana,

orange, cotton bolls, maize cobs, tomato and other species of Solanum, Capsicum

minimum, peaches, pumpkin, marrow, melon, breadfruit, pepper, hog-plum

(Spondias) and pawpaw (Carica papaya) for excisa, Tahitian chestnut (ivi =

Inocarpus edulis) for poecilopoda, and Elaeis guineensis for pallidipalpis. The

latter was originally bred from seed-potatoes. A. excisa has further been

obtained from dura-stalks (Andropogon sorghum), Chinese Yam (Dzoscorea

batatas), sugar-cane (Saccharum officinale; “larva a scavenger in Diatraea

tunnels’’), kitchen-refuse, rotten fish, cow-dung, and human faeces. A. integri-

femur was bred from bamboo, but will hardly attack the sound plant. Females

of an undescribed species of this group have been “‘bred as parasites on dura-

stemborer’’, and some of the labels on specimens of A. excisa express the opinion

that the larvae have attacked the boll-worm (Earias insulana Boisd.), Leucinodes

orbonalis Guen., Nacoleia octasema Myr. and other pests. The larvae may, as a

whole, be scavengers and perhaps occasionally attack other insect larvae.

They can, however, hardly be strictly carnivorous, as remarks by Cuthbertson

(1936, Occas. pap. Rhodesian Mus. 5: 56) and Séguy (1937, Gen. Ins. 205: 224)

would suggest. However this may be, the first two groups are either innocuous

or more or less beneficial insects in the larval as well as in the adult stage.

The species of Atherigona s. str., on the other hand, are in the larval stage

strictly phytophagous, as far as known, and often very noxious to cereal crops.

This was first made known by Malloch (1923, Ann. Mag. nat. Hist. (9) 12: 177;

1925, Mem. Dept. Agric. India, Ent. Ser. 8: 111-122) and Ramachandra Rao

(l.c. 123-125). The species 1-3 and 8-15 in Rao’s list belong to Atherigona

s. str. and are pests in various species of Panicum, Setaria, Andropogon, Eriochloa,

Pennisetum, and Paspalum, as well as rice and wheat. As will be seen from the

following descriptions, the larvae of the African species of this group attack

mainly dura (Andropogon sorghum), berseem (Trifolium alexandrinum) and

Rawsonia, the latter being the only non-gramineous plants apparently proved

to be attacked by these flies. Cuthbertson (1936, Occas. pap. Rhodesian Mus.

MUSCIDAE: B.—COENOSIINAE 103

5: 57) mentions Rhodes Grass (Clovis gayana) as being infested by larvae of an

Atherigona.

With regard to the described species, mentioned in Séguy’s work or published

since, but omitted from the following key, nothing can be said about the position

of humeralis Wied., perpulchra Bezzi (pedibus nigris, coxarum apice rufescente),

unicolor Stein (legs entirely black), and subnigripes Karsch (apex of femora

and all the tibiae and tarsi perfectly black). The following suggestions may be

made on the remaining species:

divergens Stein 1913 = ? (cinarina Ség. 1938).

flavicoxa Stein 1913, ? near bimaculata Stein, but legs dark brown, except

for knees and front coxae, which are reddish or yellowish.

immaculata Stein 1910! may from the description of the interfrontals

be supposed to belong to the excisa group and may be related to rather

similar females of an undescribed species from Sudan.

(magnipalpis Stein 1906) = excisa Thoms. 1868 (already stated by Bezzi,

1928, Dipt. Brachyc. Atheric. Fiji: 171).

meripalpis Stein 1913, Ann. Mus. nat. Hungar. 11: 539) = lineata Ad. 1905,

n. syn.

(nigripes Stein 1913, non 1900) = tridentata Mall. 1923, teste Mall. 1923,

p. 192.

parvipuncta Stein 1913, ? near ruficornis Stein 1913.

(pharalis Ség. 1938) = ruficornis Stein 1913, n. syn.

(secrecauda Ség. 1938) = perfida Stein 1913, n. syn.

Dr. Edwards’s material does not contain any representatives of the first

two groups, though four at least (excisa, integrifemur, aberrans and setifemur)

occur in East Africa. All the new species except the first belong to Atherigona

s. str. and present the following characters which are, therefore, not mentioned

in the descriptions: palpi of the male dilated and soft-haired at apex, with

about 3-4 strong, black setulae near base of dorso-exterior surface (Fig. 60),

palpi of the female (Fig. 61) band-shaped, beset with fine, black, stubby setulae

along dorsal edge (palpi not band-shaped in A. haplopyga), front femora with

one (or two) pu preapical setae in both sexes (front femora missing in the

type of tviangularis), front tibiae without a submedian bristle, lateral sub-basal

seta of scutellum of much less than a third the length of the lateral seta, unless

lor 1931? A note on the cover of the second half-volume of vol. 71 of the Denk-

schriften remarks that the papers in it have been obtainable through the book-sellers from

1907. This can obviously not be true for the part on Diptera which was presented for

publication only in November 1909 and can thus hardly have appeared before 1910, the

year which Séguy cites, even if this small part was issued separately at all before 1931.

The Zoological Record reports on the paper in the latter year.

104 RUWENZORI EXPEDITION

otherwise stated. In addition some other characters are common to or extremely

similar in all these species, which nevertheless are mentioned in the descriptions,

and other characters are described though they are expected to be liable to

variation in a larger material. This is done in order to show which characters of

the material studied have been checked in the single species described.

6/.

Fics. 60-62. Left palpus of Athevigona, exterior view. 60.—perfida Stein, 3.

61.—perfida Stein, 9. 62.—evcisa Thoms., ?.

KEY TO THE ETHIOPIAN SPECIES OF ATHERIGONA

1(6). Lower stpl and pu preapical seta of front femora absent. Either the base

of the hind femora with a short comb-like pv row of setulae (short and

stubby in female, longer and more conspicuous in male), or 7-m dis-

tinctly beyond middle of discal cell. Palpi of both sexes band-shaped

and without stronger sub-basal setulae, fuscous. Front femora without

a dorsal emargination near apex. Hypopygium of male without a

prominence immediately behind hind margin of fourth segment and

without a trifoliate process.

2(3). Hind femora with only the normal, short appressed hairs. v-m distinctly be-

yond middle (at about three fifths) of discal cell. Second abdominal

tergite of male with a dense tuft of long, black, flexuous bristles on

each anterior lateral angle, the tergite usually overlapping venter so

far that the tufts appear to be situated on venter; fourth tergite (3) with

a long pale bristle on each side of disc, which extends forward along

venter almost to base of abdomen. Discal cell (¢) rounded at outer lower

angle, m,-+cu, not extending beyond the latter; both normal in female.

Tibiae with the submedian setae minute. : : Labervans Mall.]

3(2). Hind femora with a short row of black pv setulae at base, which are minute

and stubby in female, but at least a third as long as the femoral diameter

in male. yv-m slightly anterior to middle of discal cell. Second ab-

dominal tergite of male without tufts of long flexuous bristles, fourth

tergite of male with only a few short black bristles. Discal cell normal,

ms,+cu, continued to margin.

4(5). Hine trochanters without a short comb of black, spinulose setulae, the

MUSCIDAE: B.—COENOSIINAE 105

femoral comb placed on a slight tubercle. Front and mid-tarsi without

fine outstanding curled av and pv hairs. The normal submedian setae

absent on all the tibiae. Fourth ventrite with two narrow processes

which are pointed at tip, fringed below with short black setulae, and

have each one short stout apical bristle. (gf; 9 unknown).

[*tuberculata Mall. |

5(4). Hind trochanters ($; 9?) with a short comb of black, spinulose setulae,

similar to the femoral comb, the latter not on a tubercle. Front and

mid-tarsi of male with fine outstanding curled av and pv hairs on first

(fore tarsi) or first to third (mid-tarsi) joints, these hairs being most

striking on mid-metatarsus. The submedian setae minute but present

on hind tibiae (1 pd, 1 ad, 1 av), absent or present (1 pd) on mid-tibiae.

Fourth ventrite ($) with two palpiform processes which are rounded at

tip and beset, on inner edge and apex, with unequal, dense, short,

black setulae. : : {[setifemuy Mall. }

6(1). Lower stpl and pu Preapical aa of fone femoral always present. Base of

hind femora without a short pu row of setulae.

7(12). Palpi of male band-shaped, of normal length and without sub-basal setae,

of female long-clavate (Fig. 62). Scutellum, besides the two strong pairs

of setae, with a distinct sub-basal lateral pair which is almost or fully

half as long as the lateral setae. v—m at or beyond middle of discal cell.

Abdomen rather broadly oval, in male without a hypopygial prominence

and a trifoliate process. Front femora of male usually with a shallow

excision before apex of dorsal surface and the apex beyond it beset with

dense, short, black hairs. Interfrontalia strongly dilated anteriorly.

(Acritochaeta Grimsh.)

8(9). Rs very much narrowed at apex, section of costa between 74,5 and m not

more than two-thirds as long as m—m in female (half as long as m—m in

male), the apex of R; with a fuscous spot in male, but only with a

shadow of it in female. Front tarsi of male with fine, outstanding wavy

to curled av and pv hairs, the av ones being twice to thrice as long as

the tarsal diameter and more numerous, especially on apical two or

* three joints. Pollinosity golden with three broad brown vittae on

thorax, ground-colour yellowish-brown to brown. Interfrontalia

orange-red. (Seychelles.) : : .[maculipennis Stein]

9(8). Rs slightly narrowed at apex, pectin of Cece neeqeen vg+5 and m four-

fifths to fully as long as m—m, the apex of Rs hyaline in both sexes.

Front tarsi of male without any hairs that are longer than the tarsal

diameter.

10(11). Seldom up to 4mm. long. Abdomen grey dusted, with well defined, paired

more or less dark brown, roundish spots on the second to fourth seg-

ments. Pleurae, including mesopleura, piceous in ground-colour.

Interfrontalia dull orange-red. y—-m very distinctly beyond middle

of discal cell. Front femora of male with the dorso-subapical excision

present. . ; : [excisa Thoms. |

11(10). 5-6 mm. Abdomen not algae daeeed TaONe well defined spots except

on fourth segment, but these elongate and not very dark. Pleurae

mainly pale. Interfrontalia brownish-black. y—m at or slightly beyond

middle of discal cell. Front femora of male without the dorso-subapical

excision. 0 : : ; : . [integrifemuy sp. n.], p. III

106 RUWENZORI EXPEDITION

12(7). Palpi of male shortened, dilated at apex and with several strong, black

setulae near base of dorso-exterior surface (Fig. 56), of female band-

shaped (Fig. 57), very seldom slightly subclavate. Scutellum with the

lateral sub-basal seta inconspicuous or small, seldom up to one-third

the length of the lateral seta. v—m well before middle of discal cell.

Abdomen more slender, in male always with a trifoliate process and

usually with a hypopygial prominence (see p. 98). Front femora never

with a dorso-subapical excision. Interfrontalia parallel or slightly

dilated anteriorly. (Atherigona s. str.).

13(20). Occiput rather black and glossy, sparsely dusted, up to the occipital setulae

in dorsal half or third. Parafrontalia largely glossy and black. Hypo-

pygial prominence of male absent or simple and knoblike.

14(15). Interfrontalia orange-yellow on front half or third. Third antennal joint

(and face) very long, fully four times as long as wide. Palpi yellow.

Pleurae and humeri entirely ferrugineous, lateral part of mesonotum

usually ferrugineous, dorsum with one or three brown vittae or piceous

with three more strongly shining vittae. Abdomen ferrugineous, the

pattern piceous, consisting in male in a median dark brown spot on

first segment, and a pair of spots on second and third segments, in

female more extended, often most of dorsum entirely glossy black.

Hypopygial prominence simple (Fig. 6), trifoliate process Fig. 56.

laevigata Loew, p. 113

15(14). Interfrontalia black. Third antennal joint of moderate length, up to about

thrice as long as wide, antennae entirely fuscous, except for base of

arista.

16(17). Four propleural setae which are inserted on a strong, knobshaped promi-

nence. Pleurae dark, even in male. Fourth abdominal segment with

a pair of dark spots and a pair of strong marginal setae, hypopygial

prominence absent. Second and third abdominal segments each with

a pair of piceous spots, infuscate and pruinose around and between

them, third segment without a piceous lateral streak. Wings hyaline.

Third antennal joint broad. (Fig. 31) A : . hancocki sp. n., p. 113

17(16). Three propleural setae which are inserted on a much weaker prominence.

Pleurae pale in male, variable in female. Palpi fuscous.

18(19). Larger: 4-5 mm. Fourth abdominal segment (g$) uniformly pale, with a

number of setulose hairs, hypopygial prominence absent. Second and

third abdominal segments entirely piceous on dorsum, except for the

margins. 74;; and m slightly sinuate and infuscate at apex. Third

antennal joint much broader in male. Lateral vitta of thorax broad,

median vitta narrow. (Fig. 35a, b) ‘ : . haplopyga sp. n., p. 114

19(18). Smaller: 3-4 mm. Fourth abdominal segment ($) with a pair of black

spots and a pair of rather strong marginal setae, hypopygial prominence

present (as Fig. 6). Second and third abdominal segments each with

a pair of piceous spots that is usually distinctly separated on median

line, third segment of male in addition with a lateral piceous longitu-

dinal streak. 7,,, and m straight at apex, not infuscate, wings hyaline

in female, with two brownish spots in male : one around h, the second

before apex of y,.. Third antennal joint subequal in both sexes, about

thrice as long as wide. (Fig. 34a, b) ; ; . bimaculata Stein, p. 116

20(13). Exterior part of occiput as densely dusted in upper half as the temples along

27(26).

28(25).

29(30).

30(29).

MUSCIDAE: B.—COENOSIINAE

the eyes, parafrontalia usually mainly dull and dusted. Hypopygial

prominence of male always present and usually truncate, emarginate,

bifurcate, or tridentate, seldom simple, knoblike.

Interfrontalia yellow, at least on anterior half or third. First and second

antennal joints ferruginous, third joint usually fuscous in female,

with the base narrowly ferruginous, variable in male.

Interfrontalia yellow, at most the borders somewhat infuscate near base.

. Palpi yellow (¢ 9). :

. Front legs entirely pale or only the tibiae and tarsi more or less piceous

(vubricornis).

. Third antennal joint of male fuscous except for a narrow basal part. Female

either almost entirely ferruginous to testaceous, or with the femora

infuscate or piceous in at least apical third (and then not falling in this

group).

. Mesonotum pale in front of suture, except for one or three narrow brown

vittae, scutellum wholly or largely pale. Hypopygial prominence

Fig. 14, trifoliate process (Fig. 54) with piceous-black lobes, the lateral

ones with a blunt tooth interiorly, median one clavate in lateral

view. Thorax and abdomen of female ferruginous to testaceous,

except for the vitta on thorax and a V-shaped spot at apex of fourth

abdominal segment. 5 : : [ferruginea sp. n. 3 9],

Mesonotum largely piceous, even in front of suture, where only the humeri

and a lateral stripe to the outer side of the prst seta are pale. Hypopy-

gial prominence as Fig. 24; trifoliate process (Fig. 46) with hyaline

lateral lobes with a brown seam, median lobe gradually and slightly

narrowed from middle to apex in lateral view. (Iemale with the front

femora partly dark.) . é . : ; [aster sp. n. 3],

Third antennal joint of male mainly ferruginous. Mesonotum and usually

the pleurae piceous to black in ground-colour, abdomen testaceous, the

first segment unspotted, some of the others with pairs of small, dark

spots, but without a V-shaped pattern at apex.

Hypopygial prominence as Fig. 5. Trifoliate process (Fig. 53) yellow, with

the apices piceous, the lateral lobes each with a yellow, conical promi-

nence on interior side. Third antennal joint with the dorsal edge and

the apex conspicuously piceous. Wings hyaline. conigeva sp. N.,

Hypopygial prominence larger, notched or emarginate at apex. Trifoliate

process purplish-black, lateral lobes with the interior margin simple.

Third antennal joint in male with the dorsal edge or apex infuscate but

not piceous.

3: apex of wings hyaline. Antennae entirely ferruginous, at most the

extreme apex somewhat infuscate. Hypopygial prominence Fig. 15 ;

trifoliate process Fig. 51. (Q unknown) ° : {vrubricornis Stein}

3g. Apex of wings narrowly infuscate along costa from middle of Ft, to m.

Antennae ferruginous, with the dorsal margin infuscate. Hypopygial

107

p. 116

Ds Itz

p. 118

prominence Fig. 27, trifoliate process Fig. 55 ; [bedfordi sp. n.] p. 121

Front femora piceous or infuscate on at least apical third.

Female with at most the apical half of the front femora piceous, front tibiae

only infuscate. (Male with wholly pale legs.)

The paired, roundish spots of the abdomen brownish and not very con-

spicuous.

IT, 4 (d)

{astey sp. n. 9], p. 117

108

35(34)-

39(38).

40(41).

41(40).

42(21).

43(64).

44(45)-

45(44)-

46(47).

47(46).

48(55)-

RUWENZORI EXPEDITION

Front femora of both sexes piceous in about apical half or more, front tibiae

largely piceous. Hypopygial prominence Fig. 20, trifoliate process

Fig. 50a, b. Third antennal joint ferruginous in male, with the dorsal

edge concolorous (or narrowly infuscate: pharalis Ség.). vuficoynis Stein, p. 121

. Palpi fuscous. (2; g unknown) : : : (sp. indet. Chyulu Hills]

. Interfrontalia yellow on anterior half or third, dark brown or black in at

least basal half. Palpi yellow.

. Third antennal joint of male fuscous (except no doubt for a narrow basal

part). Apical fourth of front femora in male (two-thirds! or more in

female) piceous, front tibiae, except for basal fourth, piceous. Abdomen

in male with a pair of dark spots on second and third segments, in

female testaceous with a pair of small, round spots on third and a more

or less indistinct one on fourth and, sometimes, second segments.

($ not seen.) : : : 3 : : : : [valida Ad.]

Third antennal joint of male ferruginous with the apex or dorsal margin

more or less infuscate. Abdomen of female (that of tomentigeva un-

known) more greyish on dorsum, more conspicuously dusted, with the

spots of the fourth segment more or less oblong.

¢. Front femora and tibiae largely piceous, except for the base (and a

narrow apical ring of the femora). Third antennal joint at least two and

a half times as long as wide. Pleurae blackish. Fourth abdominal seg-

ment with a pair of small round piceous spots. Hypopygial prominence

Fig. 21, trifoliate process (Fig. 37a, b) purplish-black. tomentigeva sp.n., p. 121

3S. Front femora and tibiae entirely pale. Third antennal joint extremely

broad, twice as long as broad. Pleurae ferruginous. Fourth ab-

dominal segment unspotted. Hypopygial prominence Fig. 13, lateral

lobes of the trifoliate process (Fig. 44a, b) largely pale brown, apical

and outer margins with a piceous border ; [marginifolia sp. n.), p. 122

Interfrontalia entirely dark brown to black, sometimes suffusedly dull

reddish on anterior quarter.

Palpi yellow.

Hypopygial prominence of male distinctly tridentate, the lateral teeth being

twice as long as the median one. Front femora and tibiae infuscate at

apex. Humeri probably blackish in ground-colour. Mesonotum

without evident vittae. Second abdominal segment with two large

round blackish spots. 92 unknown. . i é : [*cinarina Ség.]

Hypopygial prominence either knoblike or truncate or more or less bi-lobed

or bifurcate, never tridentate. :

Hypopygial prominence strongly bifurcate (Fig. 19). Humeri black or grey

in ground-colour. Front femora of male with the apical third or half

piceous. Mesonotum without distinct vittae. (Q with the palpi

fuscous, see par. 72) (Fig. 41) 3 : [indica infuscata ssp. n.], p. 123

Hypopygial prominence of male knoblike, truncate or bi-lobed. Humeri

more or less yellow in ground-colour.

Hypopygial prominence of male bi-lobed (Figs. 26-30). Pleurae dark, at

least in female. Fourth abdominal segment of male entirely pale.

1 At most the apical third of the femora somewhat infuscate even in female, dorsum

largely ferrugineous: see remarks on A. laevigata, p. 113.

49(50).

50(49).

51(54).

52(53).

53(52)-

54(51).

55(48).

56(57)-

57(56).

58(59).

59(58).

6o0(61).

MUSCIDAE: B.—COENOSIINAE

Front femora of male with the apical third strongly infuscate. Lateral lobes

of trifoliate process (Fig. 52a, b) ferruginous with whitish base, median

lobe testaceous, with a hyaline blade towards body. Larger species,

4-5 mm. (Fig. 28). : 6 ; : gilvifolia sp. n., p.

Front femora of male entirely yellow, or, ae most, slightly brownish along

dorsum of apical part. Lateral lobes of trifoliate process black, much

broader, median lobe styliform or compressed, but without a hyaline

blade.

Smaller species, 2-5—4:6 mm. Hypopygial prominence (Figs. 26, 30) slightly

bi-lobed in posterior view. [Front tibiae of male usually infuscate at

apex. Second antennal joint usually more or less infuscate or fuscous.

Thorax usually with three narrow brown vittae. Pleurae dark in both

sexes.

First abdominal segment entirely ferruginous or slightly brownish in

middle or with a piceous spot of moderate size, i.e. confined to the

median third or half, spots of the other segments small to moderate.

Basal joints of antennae ferruginous in ground-colour, but infuscated

on disc. Hypopygial prominence Fig. 26, median lobe of trifoliate

process (Fig. 57) clavate and incurved at apex in lateral view, slightly

compressed elsewhere. Slightly smaller. . 4 perfida Stein,

Tirst abdominal segment broadly piceous, the outer margin of this dark

pattern approaches the side-margin of the segment and reaches or sur-

passes the level of the outer margin of the dark pattern on the second

segment, spots of the other segments large. Basal joints of antennae

iuscous. Hypopygial prominence Fig. 30, median lobe of trifoliate

process (Fig. 48a, b) attenuated to apex in lateral view, not incurved,

more strongly compressed. Slightly larger. - tvapezia sp. N., p.

Larger species, 4:2—5°5 mm. Hypopygial prominence (Fig. 29) subcordiform

in posterior view. Front tibiae of male entirely pale. Second antennal

joint ferruginous. Thorax usually without distinct vittae. Pleurae

dark in female, mainly pale in male (Fig. 58) : . steeleaé:sp. Ni., p.

Hypopygial prominence truncate, with the angles somewhat produced, or

knoblike.

Small species, § 3mm. Apical third to half of front femora piceous in male.

Median lobe of trifoliate process (Fig. 32) filiform with the apex by no

means dilated in ventral view. Parafrontalia entirely dusted. Wings

quite hyaline. Pleurae dark. Hypopygial prominence Fig. 25.

longifolia sp. N., p.

Larger species, 3:8-4:8 mm. Front femora of male entirely pale (tviangu-

lavis?). Median lobe of trifoliate process more or less dilated at apex

in ventral view, lateral lobes broad, subtriangular or suboval, at least

in apical half.

Parafrontalia entirely dusted. Wings quite hyaline. Hypopygial prominence

Fig. 23, trifoliate process Fig. 59. Pleurae pale in male, hardly in-

fuscate in female. Thorax usually with three faint, narrow, brown

vittae, which do not reach scutellum. Fourth abdominal segment of

male unspotted. 5 : : : . budongoana sp. N., p.

Parafrontalia largely black and glossy. Eponetl prominence more or

less knoblike.

Virst abdominal segment entirely pale. Thorax without distinct vittae.

109

127

130

131

IIo

68(67).

69(70).

70(69).

71(72).

72(72).

73(64).

74(75)-

RUWENZORI EXPEDITION

Apex of 7,+,; and m not infuscate. Upper prostigmatal minute.

¢$. Pleurae pale. Fourth abdominal segment with a pair of small piceous

spots. Hypopygial prominence Fig. 8, trifoliate process Fig. 45.

[tvuncata sp. nj, p. 133

. First abdominal segment with a conspicuous piceous spot. Thorax with at

least a distinct, brown median vitta. Apex of 74,, and m narrowly

infuscate (in male).

. Upper prostigmatal conspicuous. Pleurae largely pale in male. Fourth

abdominal segment unspotted. Thorax with a pair of faint, linear

sublateral vittae and a not very narrow median vitta which reaches

prsc but not scutellum. Hypopygial prominence Fig. 7, trifoliate

process Fig. 49, median lobe slightly broader and gradually widened

in a hardly appreciable measure in lateral view. pedunculata sp. n., p. 134

. Upper prostigmatal minute. Pleurae dark in male. Fourth abdominal

segment with a pair of piceous spots. Thorax with a single, rather

dark, median vitta which dilates towards scutellum and reaches apex

of the latter, occupying most of the width of its disc. Hypopygial

prominence Fig. 11, trifoliate process Fig. 42a, b. tviangularis sp. n., p. 135

. Palpi fuscous, or mainly fuscous with the apex somewhat ochraceous or

whitish.

. Parafrontalia entirely dusted, at most with small glossy points at the base

of the setae.

. First abdominal segment entirely pale or only with some brownish shadows

in both sexes.

. Hypopygial prominence rather large, knobshaped (Fig. 9). Parafacialia

golden dusted, distinctly wider below than the base of the arista.

Humeri pale. Median lobe of trifoliate process (Fig. 40a, b), in lateral

view, hardly broader at apex than in middle. . aurifaciés sp. N., p. 136

Hypopygial prominence small, bifurcate, the two branches slender and

simple. Parafacialia silver-dusted, as wide below as the base of arista.

Humeri often more or less dark. Median lobe of trifoliate process

filiform in lateral view, more or less finely clavate at apex.

Apical two-fifths of front femora infuscate on outer surface in male, apical

joints of the mid and hind tarsi black in female. Lateral lobes of

trifoliate process (Fig. 36) on a long testaceous stalk which is longer

than the lobe. Hypopygial prominence Fig. 16 (lineata lineata Ad.}

Front femora of male entirely pale, apical joints of the mid and hind tarsi

more or less infuscate in female. Lateral lobes of the trifoliate process

on a short stalk which is hardly more than half as long as the lobe.

Both sexes with fuscous palpi. Abdominal spots of female roundish, small

and ill defined or more or less reduced. (Figs. 18, 39.)

lineata ugandae ssp. 0., p. 137

(Male with yellow palpi, see par. 40.) Abdominal spots of female oblong-

quadrate, rather large, piceous, and sharply defined.

{indica infuscata, sp. n. Qj, p. 123

First abdominal segment with a piceous spot. Thorax with three distinct

brown vittae. Hypopygial prominence simply transverse-knobshaped

or emarginate at apex, with two small and blunt lobes, transverse and

more or less bean-shaped in vertical view.

Lower stpl as far from the posterior as from the anterior one. Second and

MUSCIDAE: B.—COENOSIINAE aa EAL

third abdominal segments testaceous, greyish in places, densely yellowish

dusted and with a brown median vitta and a pair of piceous spots, which

are inverted-triangular on both segments (?) or on second ($). Wings

of male with two smoky suffusions beyond h and apex of sc. Hypopygial

prominence Fig. 10, trifoliate process Fig. 33a, b.

[binubila sp. n.J, p. 138

75(74). Lower stpl slightly closer to the posterior than io the anterior one. Second

and third abdominal segments more largely piceous and dark grey, the

spots elongate-quadrate. Wings unspotted. Hypopygial prominence

conspicuously emarginate, lateral lobes of trifoliate process much

shorter.

76(77). Viceous spots of second abdominal segment fused and occupying the entire

width of the segment, except for narrow side-margins, at most with

some very faint dust in a tongue along median line, abdomen glossy

with distinctly reddish-testaceous ground-colour. Median lobe of

trifoliate process (Fig. 47) very slightly compressed. Smaller: 2°5—-3°8

mm. (Fig. 17) . Q : : : . piceiventris sp. n., p. 139

77(76). Piceous spots of second abdominal eoement separated on median line by a

broad, more or less bi-lobed tongue of very conspicuous grey dust or of

ground-colour, abdomen strongly dusted and the ground-colour usually

more or less greyish, especially on middle and on anterior parts of sides.

Median lobe of trifoliate process very strongly compressed (Fig.38 a, b).

: Larger: 4-5 mm. (Fig. 12). 5 : . grviseiventyis sp. D., p. 140

78(65). Parafrontalia not dusted, but black and glossy, pceee near base of antennae.

79(80). Two or three propleural setae, which are inserted on a slight prominence.

Propleurae and humeri entirely testaceous to ferrugineous. Thorax

densely yellowish-grey dusted, the pale parts golden dusted, one or

three vittae distinct. Hypopygial prominence with three minute

teeth or as Fig. 22, trifoliate process Fig. 43. . tridentata Mall., p. 142

80(79). Four propleural setae, which are inserted on a conspicuous, knobshaped

prominence.! Propleural depression and humeri rufo-piceous to black,

the humeri sometimes partly yellowish. Second and third abdominal

segments almost entirely, front femora entirely piceous.

hancocki sp. n., 2, p. 113

[Atherigona integrifemur sp. n. 9 9}

Length 5:0-5°7 mm.; of wings 4:6-5:2 mm.

$. Head testaceous below and in front, fuscous on occiput and frons,

densely pale grey dusted, except on median dorsal part of occiput, which is

fuscous, and interfrontalia which are dull brownish-black. Interfrontalia rather

strongly widened in front, parafrontalia with 4-5 inclinate setae (the second

or second and third small), two more inclinate small setae on uppermost part

of parafacialia, level with the second antennal joint. Inner vertical very strong,

1 4. nigvithovax Stein from Togo must be very similar to the female of hancocki, but it

may be expected to have only two or three propleurals on a slight prominence. The third

abdominal segment is ferruginous with a pair of black, round spots and an indistinct

median vitta, “the front femora are only infuscate at the apex, etc.

II2 RUWENZORI EXPEDITION

outer vertical, postverticals and ocellars rather long. Antennae fuscous, third

joint thrice as long as wide, arista very short-pubescent but more conspicuously

so than usual in the genus. Palpi fuscous, band-shaped, somewhat excavate

along interior surface, apex densely beset with short, fine, erect, yellow hairs,

base without setae. Jowls slightly narrower at the narrowest point than the

parafrontalia in middle.

Thorax testaceous, the dorsum, excluding the humeri, piceous, the ptero-,

sterno-, hypo- and metapleura and the metaphragma more or less brownish,

the pale parts golden-dusted, the dark ones grey-dusted, mesonotum with

three rather conspicuous, not very narrow vittae, scutellum suffusedly brown.

prst acy in 5-6 rows, the last frst pair not distinctly stronger, the prsc well

developed, 2 + 3—4 dc, only the last two strong; scutellum with the lateral

sub-basal seta almost half as long as the lateral one, the latter and apical setae

subequal. Three propleural setae, one of them minute, the other two rather

strong, upper prostigmatal rather well developed. stf/ in a triangle, the posterior

side of which is considerably shorter and which bears some rather strong setulae,

lower stf/ much weaker than the two others which are subequal.

Abdomen short and robust, brownish-testaceous with some brownish

shadows on posterior segments, third tergite meeting on venter with its margins

raised along median line, the first three segments with some rather strong

marginal setae towards sides. Hypopygial prominence and trifoliate process

absent.

Legs testaceous, front femora infuscate on more than apical half, front

tibiae, except basal third, and hind tarsi, except apical joint, piceous, front

tarsi and basal two joints and bases of the next two joints of mid-tarsi also

piceous, the apices of the basal two joints of mid-tarsi narrowly paler, hind

tibiae slightly infuscate. Front femora with two strong pv preapical setae,

front tibiae unarmed in middle, mid-tibiae with a moderate submedian fd seta

(and on one side an abnormal av), hind tibiae with moderate ad, av and pd setae,

the latter two on the same level. ;

Wings hyaline, veins brown with testaceous base, 74, and m conspicuously

converging though straight near apex; v-m at middle of discal cell. Calyptrae

hyaline with pale-yellow border. Halteres yellowish-white.

Q very similar to male, third antennal joint narrower, more than thrice as

long as wide, palpi elongate-subclavate, without yellow hair but with minute

black setulae, abdomen longer, more depressed, pointed at apex, the hind

margin of the segments and a pair of elongate, oblique spots on the fourth

segment brownish. Front femora piceous except for the basal quarter to sixth.

y—m at or slightly beyond middle.

Kenya: Limuro, vi. 1937, bred ex Bamboo (Van Someren) 3 type 4 2

paratypes (2 2 paratypes returned to Coryndon Memorial Museum).

MUSCIDAE: B.—COENOSIINAE II3

Atherigona laevigata Loew

UGANDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 1 2; typical locality:

Portuc. East ArricA: Inhambane.

In B.M. from AByssIniA. TANGANYIKA: Tanga (J. Ogilvic, Miss A. Mackie);

PortuGc. East AFRICA: Beira (Miss A. Mackie); Kenya (from Coryndon

Mem. Mus.): Naivasha, Ngong and Nairobi, in Coryndon Mem. Mus. from

Rabai and Chyulu Hills; seen from S. RHODEsIA: Salisbury (A. Cuthbertson).

The female specimen from the Seychelles, identified by Stein as scutellaris

Stein (1910, Trans. Linn. Soc. Lond., Ser. 2, Zool., 14: 158), has the lateral

parts of the occiput dusted, it traces near valida in the key. The specimen is

quite similar to ferruginea m., but the interfrontalia are black on posterior two

thirds, and the two parts of the V-shaped spot on the fourth abdominal segment

do not meet at apex. No males are available caught together with females of

this kind, but it is probable that they are aberrant laevigata.

Atherigona hancocki sp. n. ¢ 2

Length 3:5 mm.; of wings 3 mm.

Head: black, jowls and parafacialia yellowish, occiput sparsely dusted and

glossy on dorsal half, parafrontalia not dusted, glossy, except at anterior end,

parafacialia, jowls and temples whitish-grey dusted. Interfrontalia dull black,

parallel, parafrontalia with 5 inclinate frontal setae. Outer vertical shorter than

inner one though very distinct, post-verticals and ocellars well developed.

Antennae fuscous, with the arista rufous at base, third antennal joint very broad,

two and a quarter times as long as wide. Palpi brown at base, yellow on apical

three fifths. Jowls as wide as the parafrontalia in middle.

Thorax: piceous-black, with the humeri and anterior part of the pleurae

ferruginous, greyish dusted, with three narrow brown vittae. acy in four rows

on presutural part of mesonotum. The four propleural setae on a strong,

knob-like prominence. Three stf// in an equilateral triangle, the lower one

weaker, the posterior one stronger than the others.

Abdomen: First segment testaceous, infuscate on centre of dorsum, second

segment piceous on dorsum and rather densely dusted, with three elongate

darker spots, which almost reach hind margin, third and fourth segments each

with a pair of small piceous spots and a pair of moderate hind marginal bristles,

the dorsum of the third segment somewhat infuscate and rather densely dusted.

Hypopygial prominence absent. In this species the “‘trifoliate’’ process (Fig. 31)

has actually five narrow lobes, the median lobe very long and filiform, strongly

curved, thinner before apex and ending in a little knob, which is perfectly bare,

the lateral lobes small, lineiform in lateral view, club-shaped in ventral view,

less than half as long as the median lobe; at their base originates another

II4 RUWENZORI EXPEDITION

deeply bifurcate lobe, the branches of which are filiform near base, then slightly

dilated (in ventral view) and suddenly narrowed and strongly incurved, so that

they almost meet the apical part of the median lobe.

Legs: testaceous, with the front femora, tibiae (except knees), and tarsi

fuscous; mid and hind tarsi fuscous. Mid-tibiae with a pd, hind tibiae with

an ad, av and pd setae, the latter not much beyond the av, all these setae very

small.

Wings hyaline, with testaceous veins. 7-m on second fifth of discal cell,

m and 7,,, slightly converging at apex. Halteres and calyptrae whitish.

9. Occiput densely dusted on most of lateral parts, third antennal joint

thrice as long as wide. Anterior part of the pleurae, except for the prominence

bearing the propleural setae, and humeri piceous to black, seldom the humeri

more or less yellowish, the lateral vittae broader than the median one. Dorsum

of abdominal segments almost entirely piceous with very narrow pale hind»

margins and slightly broader lateral margins of the segments, and a V-shaped

pattern on middle of third and fourth segments, which is formed by denser

grey dust and sometimes even pale ground-colour, and which reaches hind-

margin with its pointed end. Front legs piceous, except for the knees, hind

femora often and sometimes even mid-femora more or less infuscate, hind

tibiae strongly infuscate, mid and hind tarsi piceous.

Ucanpa: Kampala, r8.iv.1926 (G. L. R. Hancock) 3 type; Ruwenzori,

Kilembe, 4500 ft. (F.W.E.) 4 2 paratypes. KENYA: Nairobi, vil.1930: (Van

Someren) 2 9 paratypes; Naivasha, vi.1936 and vii.1937 (H. J. A. Turner)

2 2 paratypes; Ngong v.1936 (Miss M. Steele) 1 2 paratype (3 paratypes

returned to Coryndon Memorial Museum).

Atherigona haplopyga sp. n. ¢ 2

Length 4-2-5 mm.; of wings 3°8-4-7 mm.

3S. Head black, parafacialia, jowls and lower part of the occiput yellowish,

occiput sparsely dusted and glossy on more than dorsal half, parafrontalia not

dusted, glossy, except at anterior end, parafacialia, jowls, temples, and lower

part of occiput whitish-grey dusted. Interfrontalia dull, black, parallel or very

slightly widened anteriorly, parafrontalia with five or four inclinate frontals.

Outer vertical much shorter than the inner one, though very distinct, post-

verticals and ocellars well developed. Antennae fuscous with the arista ferru-

ginous at base, the third antennal joint about twice and two thirds as long as

wide. Palpi brown. Jowls slightly wider than the parafrontalia in middle.

Thorax piceous-black on dorsum, including scutellum, the humeri and

pleurae rufo-testaceous, densely golden dusted, the dorsum grey dusted with

a narrow median and a pair of rather broad sublateral vittae of brown dust, the

latter reaching from the outer row of acy almost to the za and from the neck to

MUSCIDAE: - B.—COENOSIINAE II5

the apex of scutellum, occupying almost the lateral thirds of the latter. prs¢

acy in four to five rows, one pair of distinct frsc, 1+-2 dc, only the last two

pairs strong, scutellum with the lateral setae slightly shorter than the apical

pair. The three propleural setae on a moderate prominence. s¢f/ in an almost

equilateral triangle, the lower one much weaker, the upper one somewhat

stronger than the anterior one.

Abdomen: first segment testaceous, the posterior half of dorsum and a

median vitta, reaching base, piceous; second segment entirely piceous, except

for a narrow, pale-yellow posterior border, which widens and turns ferru-

ginous towards venier, taking up almost half of the length on middle of lateral

surface; third segment piceous, with a deep median longitudinal impression

(normal?) and a rufous spot on either side of hind margin, the first to third

segments with a few moderate marginal setae towards sides; fourth segment

uniformly ferrugineous, with a number of fine, setulose hairs, but without

setae, hypopygial prominence absent. Genitalia translucent testaceous with

large deep-black plates. Trifoliate process (Fig. 35a and b) with the median

lobe black, rectangularly bent, rather short, its basal part triangularly narrowed

from base to apex in ventral view, subtriangularly dilated from base to apex

in lateral view, the apical part (at right angles to the basal part and almost

parallel to the lateral lobes) rod-shaped, shghtly tapering, with two long white

hairs on either side of apex, lateral lobes hyaline yellowish, with a black point

at exterior basal angle and a short black streak just inward of it, longer than

median lobe, and twice as long as the apical part of the latter, rather slender

and gradually tapering, the apex rather narrowly rounded.

Legs testaceous, with the apex of the front femora, the greater apical half

of the front tibiae, and the whole of the front tarsi piceous, the hind tarsi

slightly infuscate. Mid-tibiae with a small fd, hind tibiae with moderate ad,

av and fd setae, the latter not much beyond the av.

Wings hyaline, the veins testaceous, but brownish towards margin, 74,5

extremely narrowly bordered with fuscous and slightly sinuate towards apex;

y—m on second fifth of discal cell. Calyptrae yellowish, slightly smoky, halteres

yellowish.

Q similar to male, but the antennae more than thrice as long as wide, pleurae

sometimes strongly infuscate, abdomen rufo-testaceous, the first segment with

a rather broad piceous median vitta, the second to fourth segments entirely

piceous on dorsum, except for more or less broad lateral margins, and on the

third segment, a narrow posterior margin. In specimens with dark pleurae

the first abdominal segment is mainly piceous on dorsum. Front femora at

apex, front and hind tibiae and tarsi, and mid-tarsi piceous to blackish. In the

female from Sudan the lateral vittae of the mesonotum are absent, whereas

the median vitta is well developed.

Ucanpa: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 3 type 5 Q paratypes;

116 RUWENZORI EXPEDITION

SuDAN: Loti, in banana-clearing near stream, 19.xi.1933 (Miss M. Steele),

Q paratype.

Atherigona bimaculata Stein.

UcanpDA: Ruwenzori: Kilembe, 4500 ft. (F.W.E.) 1 g$ 1 9; Kyarumba,

4500 ft. (D. Rk. Buxton) 1 9; typical localities: SEYCHELLES: Silhouette and

Mahé (both in B.M.).

The discovery of this species on the African continent is rather unexpected.

The specimens are in no way distinguishable from the nine typical specimens

in the B.M., the trifoliate processes of the males being also alike (Fig. 36a, 6),

and striking in their narrow lateral lobes.

[Atherigona ferruginea sp. n. 2 3]

Length 3-5 mm.; of wings 3-2 mm.

2. Head testaceous with the upper half of occiput and the mentum dark

brown, parafrontalia (except for a small spot around the insertion of each seta),

face, parafacialia, jowls, temples and occiput (the latter except for the usual

median dorsal band) densely whitish dusted, the dust almost absent on a line

along the occipital setulae. Interfrontalia dull testaceous, somewhat widened

in front; parafrontalia rather broad, as wide as interfrontalia in front and in

middle, with 4—5 inclinate frontal setae. Outer vertical distinctly weaker than

the inner one, postverticals and ocellars not very strong. Antennae ferru-

ginous, the third joint fuscous from base of arista to apex, four times as long

as wide, last joint of arista infuscate. Palpi yellow. Jowls as wide as the

parafrontalia in middle.

Thorax ferruginous, slightly paler on pleurae, with a narrow brown median

stripe on anterior half, thin white dust between the dc rows, shining on lateral

dorsal part, and with thin golden dust on pleurae, scutellum with a narrow pale

median vitta. acy in 4-5 rows, a small pair of prsc, 2+ 2 dc, only the last two

pairs strong, scutellum with the lateral setae slightly weaker than the apical

pair. Two. propleural setae, the lower one very small. st¢f/ in an equilateral

triangle, which is practically bare of any hairs, the lower seta slightly weaker

and the posterior slightly stronger than the anterior one.

Abdomen yellowish-ferruginous, with a slight brownish shadow on centre

of first segment, some irregular brownish shadows on second, a pair of brownish

spots on third and a dark brown V- or horseshoe-shaped spot on apex of fourth

segment. This spot is open anteriorly, and its lateral parts reach almost to the

basal two fifths of the segment. Dorsum depressed, hairs black and small, the

marginal ones somewhat stronger, especially at sides, but no distinct setae

present.

MUSCIDAE: B.—COENOSIINAE LE,

Legs testaceous with the front tibiae and tarsi infuscate. Mid-tibiae with

a small fd, hind tibiae with moderate ad, av and fd setae, the latter two on the

same level.

Wings hyaline, the veins pale testaceous, 7,,; slightly converging with m

at apex, though the last section of the latter is straight. y-m just beyond basal

third of discal cell. Calyptrae yellowish-hyaline, halteres pale yellowish.

SupAN: Shambat, 30.xii.1930, on berseem (Tvifolium alexandrinum)

(H. W. Bedford) 9 type.

A male with the same data probably belongs to this species. The para-

frontalia and lateral parts of occiput are entirely dusted. Third antennal joint

not quite thrice as long as wide. Palpi yellow. Thorax infuscate from suture

to base of scutellum, with three narrow vittae in front of suture, the lateral

ones not very distinct and abbreviated in front. Lower stp/ very weak. Second

abdominal segment and base of third infuscate (normal?, these segments are

somewhat greasy), each with a pair of round piceous spots, the first three seg-

ments with some moderate marginal setae towards sides, fourth segment

entirely testaceous, without distinct setae though with setulose hairs. Hypo-

pygial prominence (Fig. 14) stalked though rather small, broadly trapezoidal in

outline, resulting from two fused subtriangular knobs. Trifoliate process

(Fig. 54) with the stalk fuscous at base, whitish-hyaline at apex up to the base

of the lobes, the latter piceous-black, median lobe rather short, strongly club-

shaped in ventral view, with the normal minute black setulae at apex, incurved

and somewhat clavate in lateral view, lateral lobes suboval with the exterior

and interior margins rather straight, the base strongly emarginate exteriorly

and with a blunt tooth interiorly.

[Atherigona aster sp. n. ¢ Q]

Length 3°3-4°4 mm.; of wings 3°3-4°0 mm.

3g. Head testaceous, the occiput fuscous except beneath, frontal triangle

and posterior part of parafrontalia fuscous, median dorsal part of occiput

glossy, interfrontalia dull testaceous, sometimes ferruginous near base, rest of

head densely dusted, the pruinosity whitish-grey on the dark and pale-golden

on the pale parts. Interfrontalia slightly broadened anteriorly, parafrontalia

with four inclinate setae, the first of them weak. Inner vertical rather strong,

outer vertical much weaker, postverticals and ocellars well developed. Antennae

ferruginous, third joint fuscous except for a narrow basal part, two and a half

times as long as wide; arista fuscous, its basal two joints lighter brown. Palpi

yellow. Jowls distinctly wider than parafrontalia in middle.

Thorax piceous on dorsum including scutellum, the part to the outer side

of the ph and prst setae and the whole of the pleurae testaceous, golden dusted,

dorsum yellowish-grey dusted, without vittae. fyrst.acy in 4-5 rows, the last

118 RUWENZORI EXPEDITION

prst pair distinctly longer, a pair of small prsc, 2+-4 dc, the two prst ones rather

distinct, but only the last two pairs strong; scutellum with the lateral seta

slightly shorter than the apical one. A strong and a rather small and some-

times a minute propleural seta. stp] in an almost equilateral triangle which

bears 1-2 minute hairs, lower stp/ rather weak, the posterior one slightly longer

than the anterior stl.

Abdomen ferruginous with base and apex testaceous, densely yellowish-

grey dusted on dorsum, first and fourth segments unspotted, second with a pair

of large, oblong, piceous spots, third with a pair of very small, round, dark

brown spots; marginal setae fine though distinct on lateral parts of first to

third segments. Hypopygial prominence (Fig. 24) formed by a blunt central

tubercle and four more sharply defined tubercles in the corners of a quadrate

around it, thus reminding of a simple star. Trifoliate process (Fig. 46) with the

stalk pale brown, the base of the median lobe and most of the lateral lobes

hyaline, and the rest of the median lobe, which is gradually clavate in ventral

view, but curved and band-shaped, and slightly narrowed from middle to apex

in lateral view, brown. The lateral lobes semicircular with the straight margin

interiorly and a strong emargination at basal third of exterior margin, the

latter dark brown from the emargination to the apex and somewhat twisted.

Legs wholly pale testaceous. Mid-tibiae with a small f#d seta, hind tibiae

with small ad, av and pd setae, the latter considerably beyond the av.

Wings hyaline, veins testaceous, 7,,, and m slightly converging at apex,

y—-m slightly before two fifths of discal cell. Calyptrae yellowish-greyish-

hyaline, slightly smoky towards margins. Halteres yellowish-white.

2 similar to male. Pollinosity of head whitish, third antennal joint thrice

as long as wide. The dorsum of the thorax less broadly yellowish on sides,

though the humeri, propleurae and anterior part of mesopleurae are always

testaceous and golden dusted, pleurae elsewhere more or less infuscate. Abdo-

men testaceous with irregular brownish-testaceous shadows, the third and

often second and fourth segments with a pair of rounded, brownish, not very

conspicuous spots.

Kenya: Naivasha, vii.1937 (H. J. A. Turner) $type 2 3 6 2 paratypes.

(t 3 3 2 returned to Coryndon Memorial Museum.)

Though the male characters are very distinctive, the female is extremely

similar to that of rwficornis, and it is very doubtful whether the females of

these two species can always be distinguished. Three female specimens from

the same locality have the characters of vuficornis and have had, therefore,

thus to be identified.

Atherigona conigera sp. n. ¢ 2

Length 3-3°8 mm.; of wings 2-8-3-6 mm.

3. Head fuscous behind, translucent-yellowish above, in front and below,

MUSCIDAE: B.—COENOSIINAE IIg

densely yellowish-grey dusted, the median dorsal part of occiput glossy fuscous,

interfrontalia orange-yellow, parafrontalia, parafacialia, and jowls pale yellow-

dusted. Interfrontalia slightly widened anteriorly, parafrontalia with four

inclinate frontal setae. Outer verticals slightly weaker than the inner verticals,

the latter not very strong, postverticals and ocellars rather weak. Antennae

ferruginous, the third joint not quite thrice as long as wide, its dorsal margin

and apex rather narrowly piceous, arista fuscous. Palpi yellow. Jowls slightly

wider than parafrontalia in middle.

Thorax piceous, humeral callus and propleura yellow, densely yellowish-grey

dusted, without any traces of vittae. rst acy in four rows, the last prst pair

hardly longer, a pair of small prsc, 2+-4 dc, of which only the last two pairs are

well developed; scutellum with the lateral and apical setae equal, moderately

strong. One strong and one fine propleural seta. s¢f/ in a triangle, which bears

a few single minute hairs, and the posterior side of which is shorter, the lower

stpl weak, the posterior one slightly stronger than the anterior one.

Abdomen ferruginous, the second segment with a pair of elongate and the

third with a pair of small, round piceous spots, which on both segments approach

but do not reach the hind margin, fourth segment ferruginous with a pair of

punctiform brown shadows on disc; marginal setae rather fine on segments 1-3,

very fine but distinct on fourth. Hypopygial prominence (Fig. 5) small, simple,

bluntly conical, longitudinally compressed, a deeply impressed area above it.

Trifoliate process (Fig. 53) brownish-yellow, the apices and the apical half of

outer margins of the lateral lobes suffusedly piceous, the apex of the median

lobe very narrowly infuscate, median lobe lineiform with the apex triangular

in ventral view, slightly compressed, narrowly band-shaped with the apex

hardly broader in lateral view, lateral lobes with their stalk broadly winged,

these wings fused near base, lobes oblong, with a broad, shallow emargination

on exterior margin near base and a tooth-shaped process on interior margin

almost in middle.

Legs testaceous, the basal two joints of the front tarsi infuscate, the apex

of the front tibiae very slightly infuscate. Mid-tibiae with a small fd, hind

tibiae with small ad, av and pd setae, the latter considerably beyond the av.

Wings hyaline, veins testaceous, 7,,, and m slightly converging at apex,

y—m at two fifths of the discal cell. Calyptrae whitish-hyaline at base, slightly

smoky in apical half, border yellowish-white, the lower calyptra much pro-

jecting. Halteres yellowish-white.

2 similar to g, pollinosity of head more whitish, thorax with slight traces

of three vittae, two odd prsc acy present, first abdominal segment with a brownish

spot on median line, second and third with an incomplete brown median vitta,

second to fourth with a pair of roundish-triangular, not very large but very

conspicuous, dark brown spots, the apical half of the front femora (except

knees) and front tibiae as well as the front tarsi piceous.

I20 RUWENZORI EXPEDITION

Kenya: Chania Falls, 4000 ft., x.1934 (F.W.E.) 3 type. W. ADEN

PROTECTORATE: Jebel Jihaf, ca. 7100 ft., x.1937 (B.M. Exp. S.W. Arabia,

H. Scott and E. B. Britton), 1 g paratype, 2 2 paratypes. The Aden male is

-almost identical with the type, the spots of the second abdominal segment are,

however, more roundish, the impression above the hypopygial prominence is

strongly transverse (in the type slightly transverse, more subcircular), the

median lobe of the trifoliate process is slightly more clavate in lateral view,

and even the basal joint of the front tarsi is hardly infuscate.

[Atherigona bedfordi sp. n. 3 9]

Length 3-3-4 mm.; of wings 3-3°3 mm.

3g. Head testaceous, occiput and haustellum fuscous, head densely yellowish-

grey dusted, except for the interfrontalia, mouthparts and the dorsal median

part of the occiput. Interfrontalia dull, testaceous, usually somewhat widening

anteriorly, parafrontalia almost as wide or even slightly wider than inter-

frontalia, except behind, with (3—) 4—5 (—6) inclinate and one reclinate

frontal setae. Outer vertical much shorter than the inner one, though very

distinct, postverticals and ocellars well developed. Antennae ferruginous,

with the dorsal edge of the third joint slightly and the apical half of the arista

rather strongly infuscate, third joint thrice as long as wide. Palpi yellow.

Jowls slightly narrower at the narrowest point than the parafrontalia in middle.

Thorax piceous-black, including scutellum, densely whitish-grey dusted,

the humeri and propleurae testaceous, the pleurae sometimes only infuscate.

prst acy in 4-5 rows, the last pair slightly stronger, the prsc pair still stronger,

2+ 3-4 dc, only the last two pairs strong, the others, however, rather distinct ;

scutellum with the lateral seta almost equal to the apical one. Two propleural

setae, the lower one small to moderate. s¢f/ in an equilateral triangle, which is

entirely or practically bare of hairs, lower seta weaker, the posterior one

stronger than the anterior one.

Abdomen testaceous, rather densely greyish dusted, somewhat rufescent on

the dorsum of the second and third segments, which present a pair of round

piceous spots each (the spots on the third segment forming a dark-dusted part

of a larger blackish spot, which is otherwise covered by pale dusting), the first

three segments with some rather strong marginal setae towards sides, the fourth

uniformly testaceous, with fine bristly hairs. Hypopygial prominence (Fig. 27)

stout, truncate and notched at apex, the two apical knobs depressed and

subtriangular. Trifoliate process (Fig. 55) with the base whitish, the lobes

purplish-black, the outer lobes broad, subsinuate at margin, the median lobe

filiform with strongly clavate apex.

Legs entirely testaceous. Mid-tibiae with a small fd, hind tibiae with

moderate ad, av and pd setae, the latter two on the same level.

MUSCIDAE: B.—COENOSIINAE I21I

Wings hyaline, the apex narrowly infuscate from the middle of Rk, to m,

veins testaceous, 7,,; and m very distinctly converging though straight at

apex; v—m between two fifths and one half of the discal cell. Calyptrae whitish,

halteres pale yellowish.

9. Similar to male. Third antennal joint fuscous, except at base, more

than thrice as long as wide. Thorax much paler, the pleurae ferruginous, the

dorsum brownish under the dusting, lateral parts of the mesonotum and the

scutellum yellowish. Anterior pairs of dc very distinct. Abdomen pale testa-

ceous, with a pair of round brown or piceous spots on the dorsum of segments

2-4 or 3 only. No distinct marginal setae present. Front tarsi brownish.

74,5 and m hardly converging, y-m on two-fifths or even slightly basad.

SUDAN: Shendi, 14.xi.1g28, bred on dura (Andropogon sorghum) (A. H.

Husein) § type; paratypes: Shambat, 30.xii.1930 I Q, 5.1.1929 I 9, II.xli.1926

3 6, 15.11.1926 2 5, on berseem (Trifolium alexandrinum) (H. W. Bedford);

Khartoum, 14.ii.1926 on cotton (Gossypium herbacewm) (H. W. Bedford) 1 3,

4.1.1931, bred in laboratory from berseem (H. W. Bedford), 1 8.

Atherigona ruficornis Stein

UGANDA: Kigezi District, Mabungo Camp, 6000 ft., 18.xi.1934 (J. Ford),

2449; Mt. Muhavura, 7000 ft., 29.1x.1934 (F.W.E.), 2 2; Typical localities:

S. RHopeEsIA: Salisbury, and AByssrnrA: Diredaua.

In B.M. from UGANDA: Jinja, 19.vil1.1930, and Katosi, on debris at edge

of lake, 20.ix.1936 (E. G. Gibbins); KeNyA: Kabete, xii.1925, in house (G. B.

Purvis); ZANZIBAR; ABYSSINIA: Maraquo and Doukham. Seen from Rabai

and Naivasha in KENYA.

In the specimens from Mabungo Camp the interfrontalia are suffusedly

blackish along the margins near ocellar triangle; in two females, however, they

are evenly brown on posterior two-thirds, and in these two specimens the

abdomen is distinctly darker with the ground-colour grey on middle of segments

and laterally on anterior parts. In the single female from Doukham (Abyssinia),

the margins of the interfrontalia are very narrowly blackish behind, but the

humeri are grey, hardly paler on dorsum than the mesonotum.

Atherigona tomentigera sp. n. 3

Length 3-1 mm.; of wings 2-9 mm.

3g. Head fuscous behind, testaceous in front and below, densely greyish

dusted, the dust more whitish dorsally, yellowish-grev behind and golden in

front and below, median dorsal part of occiput glossy fuscous, interfrontalia

dark brown with the anterior third yellowish-orange. Interfrontalia very

slightly widened anteriorly, parafrontalia with four inclinate frontal setae.

Outer vertical much weaker than inner vertical, the latter not very strong,

122 RUWENZORI EXPEDITION

postverticals and ocellars rather weak. Antennae ferruginous, third joint two

and a half times as long as wide, its dorsal margin narrowly infuscate near

apex; arista rufous in basal half, fuscous in apical half. Palpi yellow. Jowls

as wide as parafrontalia in middle.

Thorax piceous, humeral callus and propleura yellow, densely yellowish-

grey dusted with three very faint brownish vittae on dorsum. prst acy in four

rows with one or two additional setulae on median line, the last frst pair hardly

longer, a pair of small prsc, of the 2+4-5 dc only the last two are well de-

veloped; scutellum with the lateral pair hardly weaker than the apical pair,

moderately strong. Two rather strong and a minute propleural setae. s¢p/ in

a triangle which bears a few single hairs, and the posterior side of which is

shorter, the lower st// weak, the posterior one distinctly stronger than the

anterior one though not longer.

Abdomen testaceous, the apex ferruginous, densely yellowish-grey dusted

on dorsum, first segment with a suffused piceous spot in middle, second with a

pair of round piceous spots on posterior half, which are connected along front

margin with a narrow brown median vitta by an arcuate brown line, third and

fourth segments each with a pair of round piceous spots (very small on fourth

segment); marginal setae rather fine on all segments. Hypopygial prominence

(Fig. 21) broad, bifurcate, the branches slender and rounded at apex, broadly

separated by a slight tubercle. Trifoliate process (Fig. 37a, b) purplish-black,

median lobe gradually but strongly incurved, compressed and gradually slightly

clavate in lateral view, lineiform with the apex strongly clavate in ventral

view, lateral lobes oblong-oval with the interior margin rather straight, the

interior third of the outer surface covered with minute, dense, white hairs.

Legs yellow, front femora, except for the basal quarter and the knees, front

tibiae, except for base, and front tarsi piceous. Mid-tibiae with a small fd seta,

hind tibiae with small ad, av and fd setae, the latter slightly beyond the ad.

Wings hyaline, veins testaceous, 74,, and m conspicuously converging at

apex, v-m slightly before two-fifths of the discal cell. Calyptrae hyaline, the

apical half of the lower calyptra yellowish-smoky, border yellowish-white.

Halteres pale yellow.

Ucanpa: Kigezi District, Mabungo Camp, 6000 ft., 18.x1.1934 (J. Ford),

3 type.

[Atherigona marginifolia sp. n. 3 Q]

Length 2:8-3°3 mm.; of wings 2°7-2-9 mm.

¢g. Head blackish or fuscous on frons and occiput, except most of para-

frontalia, anterior third of interfrontalia and ventral third of occiput which

are yellowish, as are the ventral and anterior surfaces of head. Interfrontalia

subparallel, very slightly widened anteriorly, parafrontalia with four inclinate

frontal setae. Outer vertical much weaker than the inner one though quite

MUSCIDAE: B.—COENOSIINAE 123

strong, postverticals moderate, ocellars rather strong. Antennae ferruginous,

the third joint extremely broad, twice as long as broad beyond middle, the

apical half of the dorsal edge and the apex fuscous, arista fuscous. Palpi yellow.

Jowls slightly wider than the parafrontalia in middle.

Thorax piceous on dorsum, humeri and pleurae yellowish, yellowish-grey

dusted on dorsum, golden dusted on pleurae, without any vittae. frst acy in

four rows, two additional setulae on median line, the last prst pair distinctly

longer, a distinct pair of prsc,2-+-3dc of which only the last two pairs are strong,

the two prst pairs, however, quite distinct; scutellum with the lateral pair

slightly weaker than the apical pair. Two propleurals, the lower one moderate,

the upper prostigmatal stronger than usual, as long as the lower propleural.

stpl in an almost equilateral triangle (the posterior side slightly shorter), which

bears one or two minute hairs, lower stf/ considerably weaker than the two

others.

Abdomen ferruginous with the base testaceous, rather densely yellowish-

grey dusted, the first segment unspotted, the second and third each with an

oblong piceous spot of moderate size, fourth segment unspotted; marginal

setae rather strong on segments I-—3, fine though distinct on fourth. Hypopygial

prominence (Fig. 13) small, subtruncate; subtrapezoidal, almost quadrate in

vertical view, with the angles slightly protruding. Trifoliate process (Fig. 44,

a, b), pale brown, median lobe darker brown, lateral lobes with the outer and

apical borders piceous; median lobe filiform with the apex somewhat club-

shaped in ventral view, rod-shaped with the apex suddenly dilated into a small

hook in lateral view, lateral lobes bean-shaped, separated on exterior margin

from the basal support by a strong emargination.

Legs wholly testaceous, only the front tarsi infuscate. Mid-tibiae with a

small pd, hind tibiae with moderate ad, av and fd setae, the latter slightly

beyond the av.

Wings hyaline, the veins testaceous, 7,,; and m straight near apex and very

slightly converging, 7m slightly before two-fifths of the discal cell. Calyptrae

yellowish-hyaline, the border pale yellowish. Halteres pale yellow.

2 with the pleurae more or less largely infuscate, the upper prostigmatal

seta small, second abdominal segment with two indefinite, large brownish spots,

third and fourth each with a rather small oval spot, front femora and tibiae

almost entirely piceous, all the tarsi strongly infuscate.

UGANDA: Busana, 22.x.1933 (I. W. Chorley), § type, I g paratype, 3 9

paratypes.

[Atherigona indica infuscata ssp. n. 3 9]

Length 3:2-3°8 mm.; of wings 3°2-3°5 mm.

§. Head fuscous behind, pale yellowish on parafacialia, jowls and lower

part of occiput, densely whitish-grey dusted, except on median dorsal part of

124 RUWENZORI EXPEDITION

occiput and interfrontalia. The latter dark brown, dull, almost parallel,

slightly widened at front end; parafrontalia with 4-6 inclinate frontal setae.

Outer vertical distinctly weaker than inner one, postverticals and ocellars well

developed. Antennae fuscous, third joint about two and a half times as long

as wide, the base of the arista reddish. Palpi yellow, with the base brown.

Jowls approximately as wide as the parafrontalia.

Thorax piceous, humeri grey, propleurae slightly paler, densely yellowish-

grey dusted. prst acy in four rows with one or two odd setulae on the median

line towards suture, the last frst pair slightly stronger, a distinct pair of prsc.

2+3-5 dc, only the last two pairs strong, the two frst pairs rather distinct;

scutellum with the lateral and apical pairs equal and not very long. Three (to

four) propleural setae, two of them quite strong. s¢f/ in an equilateral triangle,

which contains one or two setulae, lower s¢/l much weaker than the two others.

Abdomen testaceous, the first segment with two large pale brown spots and

a pale brown median vitta which extends over the next two segments, the

second with a pair of elongate piceous spots which do not quite reach the hind

margin, the third with a pair of smaller elongate spots, the fourth with a pair

of round, small, brown spots, dorsum of abdomen rather densely yellowish

dusted. The lateral marginal setae rather distinct on the first three segments,

hardly distinct on the fourth. Hypopygial prominence (Fig. 19) broad, strongly

bifurcate, the branches broadly separated, short, truncate and subemarginate

at apex. Trifoliate process (Fig. 41) with the stalk and basal supports of the

lobes fuscous, the lobes piceous, median lobe filiform in basal half, subtriangu-

larly dilated with the apex rounded in apical half (ventral view), strongly

incurved and slightly club-shaped in lateral view, lateral lobes dull, finely

rugose in longitudinal direction, elongate-halfmoon-shaped, the outer margin

broadly emarginate in basal half, the inner margin almost straight, very

slightly emarginate towards apex, the latter narrowly rounded.

Legs testaceous, the apical third or half of the front femora (except knees),

the apical half of the front tibiae, and the front tarsi rather strongly infuscate.

Mid-tibiae with a small pd, hind tibiae with moderate ad, av and pd setae, one

beyond the other.

Wings hyaline, the veins testaceous, 7,,, and m distinctly converging though

not curved at apex, 7v-m just beyond second fifth of discal cell. Calyptrae

hyaline, .pale-yellowish, halteres whitish.

2 similar to male. Third antennal joint fully thrice as long as wide. Palpi

fuscous. The second to fourth abdominal segments each with a pair of elongate

piceous spots, which do not touch the hind margins and are of similar elongate

trapezoidal shape, but ‘decreasing size. The rows of marginal setae rather

distinct. Front femora with the apical two thirds piceous (except for the

knees), front tibiae piceous with the base narrowly pale, front tarsi piceous, the

other tarsi and hind tibiae infuscate.

MUSCIDAE: B.—COENOSIINAE 125

UcanpDA: Bukalaca, xi.1929, larva in leaf-bud of Andropogon sorghum

(H. Hargreaves), 3 type (in B.M.) and 1 g paratype (returned), r 9 paratype

(in B.M.); Morocco: Fez, x.1938, 15.xii.1938, 22.11.1938, from Andropogon

sorghum saccharatum (C. A. Bléton), 2 $3 Q paratypes. The Moroccan specimens

of this species were originally identified as A. soccata Rond., though the

palpi of the female are yellow in this species according to Rondani’s description.

Only later was it recognised that these specimens are conspecific and, in

fact, entirely identical with the Uganda form bred from the same plant.

Though this form would not trace to indica Mall. in Malloch’s key to the

Indian species (1925, Mem. Dept. Agric. India, Ent. Ser. 8: 119) nor zndica to

this species in the present key, it will nevertheless only be a subspecies of it.

In indica the front femora are entirely pale without any trace of an infuscation

in male, and with only the apical half infuscated in female, the front tibiae not

distinctly infuscated at apex in male, infuscated but not piceous in female, the

interfrontalia rufous-brown. The abdominal spots are smaller in zudica, and

the first segment is entirely pale. The hypopygial prominence and the tri-

foliate process are, however, identical in both forms, the median lobe of the

trifoliate process is in both forms filiform in basal half. In Malloch’s figure the

basal half is hidden, and only the dilated apical half can be seen; a study of

Malloch’s types in the B.M. reveals, however, that the process of tudica s. str.

is quite as shown in Fig. 41. The biology of imfuwscata was described under the

name of indica Mall. by Harris (1935, Rept. Dept. Agric. Tanganyika 1934: 84).

Atherigona gilvifolia sp. n. gd 9°

Length 3-8-4-7 mm.; of wings 3°5-4:2 mm.

3. Head fuscous behind, yellowish in front and below, densely grey dusted

with the parafacialia yellowish-grey dusted and the median dorsal part of

occiput glossy fuscous. Interfrontalia dull fuscous, subparallel, parafrontalia

with 4-5 inclinate frontal setae. Outer vertical considerably weaker than

inner vertical though quite strong, postverticals and ocellars rather strong.

Antennae with the first and second joints ferruginous, the third fuscous except

on base which is narrowly ferruginous, somewhat more widely so on. ventral

side, almost two and a half times as long as wide, arista fuscous with the base

narrowly ferruginous. Palpi yellow. Jowls not wider than the parafrontalia

in middle.

Thorax piceous with the humeri and propleurae yellowish, densely grey

dusted, on dorsum more yellowish-grey dusted, no distinct vittae. rst acy in

4-5 rows, the last frst pair slightly stronger, a pair of distinct frsc, 2-4-5 de,

the prst pairs comparatively strong, the last two pairs strong; scutellum with

the lateral sub-basal setae approximately a third as long as the lateral ones,

the apical ones not longer than the latter. Two strong and one small propleural

126 RUWENZORI EXPEDITION

setae. stl in an almost equilateral triangle, which bears a few fine setulae, the

lower stpl much weaker than the two others which are subequal.

Abdomen testaceous, yellowish dusted, slightly infuscate on dorsum of first

and second segments, with a pair of oblong-oval piceous spots on second and

of small round spots on third segment, fourth segment unspotted. Hypopygial

prominence (Fig. 28) stout, short, shortly bi-lobed at apex, the lobes short, oval,

somewhat depressed. Trifoliate process (Fig. 52 a, b) whitish at base (the stalk

however largely black), the lateral lobes ferruginous except for the whitish

base, rather elongate, strongly emarginate at base exteriorly, median lobe

styliform (in lateral view) with a hyaline blade on dorsal side, which is much

broader than the lobe itself, and which connects the recurved, blackish apex

with the stalk, the apex with two or three small black setulae, in ventral view

the median lobe is quite lineiform.

Legs testaceous, the apex of the front femora and the greater apical part of

the front tibiae as well as the basal 3-4 joints of the front tarsi infuscate.

Mid-tibiae with a small fd, hind tibiae with moderate ad, av and pd setae, the

latter slightly beyond the av.

Wings hyaline, the veins testaceous, 7,,; and m straight and very slightly

converging at apex, 7m slightly beyond two-fifths of discal cell. Calyptrae

whitish-hyaline, the border slightly yellowish, halteres pale yellowish.

9 similar to male. Parafacialia with whitish dust. Antennae almost thrice

as long as wide. Palpi fuscous at base, ferruginous in apical two-thirds.

Abdomen testaceous with some brownish shadows on first and especially second

segments, the third and fourth each with a pair of oblong piceous spots, those

of the fourth segment much smaller. Front femora, tibiae and tarsi almost

entirely blackish, hind tarsi strongly, hind tibiae and mid-tarsi slightly infus-

cate. v—m at two-fifths of discal cell.

NIGERIA: Ibadan, 19.x.1935, ¢ type, 14.x.1935 I 2 paratype (Ff. D. Gold-

ing); UGANDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.), 1 3 paratype, 3 9.

Except for their size these latter females might as well belong to steeleae.

Atherigona perfida Stein

n. syn. secrecauda Ség., 1938, Miss. scient. Omo, 4 Zool.: 372.

Ucanpna: Mt. Elgon, between Butandiga and Bulambuli, 8000 ft., 7.viil.

1934 (J. Ford), 19 3, 61 2; Butandiga, 7000 ft., sweeping short grass, 5—6.Viil.

1934 (J. Ford), 6 2; Kigezi District, Mabungo Camp, 6000 ft., 18.xi.1934

(J. Ford), 18 3, 22 2; Mt. Muhavura, 7000 ft., 29.1x.1934 (F.W.E.), 1 3, 2 9;

Ruwenzori, Namwamba Valley, 6500 ft. (F.W.E.), 1 g; Entebbe, 13.xii.1934

(F.W.E.), 1 3. Kenya: Aberdare Range, Mt. Kinangop, goo0 ft., I.xi.1934

(F.W.E); 1g; 9000 ft., cedar forest, 27.x.1934 (F.W.E.), r 3.

Typical locality: AByssin1A: Diredaua; of secrecauda: Mt. Elgon, East-

slope, Elgon saw mill, 2470 m.

MUSCIDAE: B.—COENOSIINAE 127

In B.M. from AByssINIA, TANGANYIKA, UGANDA: Busana, 22.x.33 (T. W.

Chorley); Arua, 1919 (R. E. McConnell); KeNyA: Trans-Nzoia District near

Cherangani Hills, 40 m. E. of Mt. Elgon, 6200 ft., 6.-9.11.1925 (C. R. S. Pitman).

Seen from KENYA: Naivasha, Nairobi, Karura.

Séguy’s description of the abdomen of secrecauda (first abdominal segment

of male not conspicuously spotted, hypopygial prominence “‘biglobuleuse’’,

abdomen of female with reduced spots) proves that his species is identical with

Stein’s perfida and not with the species hereafter described as trapezia. The

darker specimens noted in the key are those caught between Butandiga and

Bulambuli, in which the lateral lobes of the trifoliate process are not dilated

interiorly near base, as is the case in specimens from Mabungo Camp, Abyssinia,

etc. These differences are too slight, however, to cause me to regard this form

as separate. The specimen from Entebbe has the pleurae pale, the hypopygial

prominence longer in ventral view, and the lateral lobes of the trifoliate process

simple interiorly.

Atherigona trapezia sp. n. 3

Length 3:1-4:6 mm.; of wings 2-9-4:3 mm.

§. Head fuscous behind and above, yellowish in front and below, yellowish-

grey dusted on occiput, frontal triangle and parafrontalia, golden dusted on

face, parafacialia, and ventral surface, glossy fuscous on median dorsal part of

occiput. Interfrontalia dull brownish-black, parallel, parafrontalia with 3-4

inclinate frontal setae. Outer vertical rather strong, though conspicuously

weaker than the inner vertical, postverticals and ocellars moderate. Antennae

fuscous, the third joint two and a half times as long as wide, arista fuscous,

rather stout. Palpi yellow. Jowls one and a half times as wide as the para-

frontalia in middle.

Thorax piceous, humeri and propleurae yellowish, yellowish-grey dusted,

with three brown vittae on dorsum, which are not very wide nor very striking,

of equal width, pleurae largely golden dusted. frst acy in 4—5 rows, the last

prst pair distinctly longer, a distinct pair of prsc. 2 + 3 dc, the prst ones quite

distinct though small, only the last two pairs strong; scutellum with the

lateral setae slightly shorter and weaker than the apical pair. One strong,

one moderate, and sometimes a minute propleural seta. s¢f/ in an equilateral

triangle, which bears one or two fine setae, lower stfl considerably weaker,

posterior one much stronger than the anterior sép/.

Abdomen testaceous, first segment with a large, more or less bi-lobed spot

or with two transverse broad spots along hind margin, a narrow hind margin

(more or less dilated on median line) and about a tenth of the width of the

segment on each side remaining testaceous, second segment with a pair of

large, oblong-quadrate spots which converge somewhat behind and occupy

128 RUWENZORI EXPEDITION

the disc of the segment but for a lateral margin as wide as on the first segment,

a narrow hind margin, and a median vitta which is broader in front but is

sometimes strongly infuscate or divided by a brownish median line, third

segment with a pair of oblong spots which do not reach either front or hind

margin, fourth segment unspotted; lateral marginal setae rather strong on

first three segments, fine though distinct on fourth. Hypopygial prominence

(Fig. 30) strongly triangularly dilated in posterior view with the apical angles

strongly projecting sideward, somewhat notched in middle, broadly trapezoidal

in ventral view. Trifoliate process (Fig. 48a, 6) dull black, median lobe filiform,

very gradually and slightly clavate in ventral view, somewhat compressed and

more or less dilated in middle in lateral view, tapering to apex, the latter not

incurved; lateral lobes suboval with the inner margin almost straight, very

slightly sinuate at. base of outer margin.

Legs testaceous, front femora at most with a slight brownish dorsal streak,

front tibiae strongly infuscate except at base, front tarsi piceous, the apex

slightly paler. Mid-tibiae with a pd, hind tibiae with ad, av and pd setae, the

latter beyond the fd or level with it.

Wings hyaline, veins testaceous, 7,,,; and m slightly converging at apex,

y—m just before two fifths of the discal cell. Calyptrae hyaline, the lower one

largely yellowish, border pale yellow. Halteres whitish-yellow.

2 similar to male, but the parafacialia more whitish dusted, the third anten-

nal joint less broad, almost thrice as long as wide, front femora largely piceous,

except for part of the base, mid and hind tarsi infuscate, second abdominal

segment more or less wholly piceous on disc, third with a pattern similar to

that of the second segment of the male, fourth similar to the third of the male,

but the median and lateral parts more or less distinctly infuscate, though

yellowish-grey dusted, often the disc of the abdominal segments wholly piceous.

UGANDA: Kigezi District, Mabungo Camp, 6000 ft., 18.xi.1934 (J. Ford)

3 type, 37 3, 39 2 paratypes; Mt. Muhavura, 7000 ft., 29.ix.1934 (F.W.E.)

23 3, 22 9 paratypes; Mt. Mgahinga, 11000 ft., 22.xi.1934 (F.W.E.), 1 3 para-

type. Ruwenzori, Kyarumba, 4500 ft. (D. R. Buxton) 2 9 paratypes. Mt. Elgon,

Butandiga, 7000 ft., vili.1934 (J. Ford) 1 3, 1 2 paratypes; Bulambuli, 9500 ft.,

vili.1934 (J. Ford) 1 g paratype; Mudangi, 11,000 ft., viii.1934 (J. Ford)

I Q paratype. Kenya: Aberdare Range, Katamayo, 8000 ft., x.1934 (F.W.E.)

I 3, 4 Y paratypes; Mt. Kinangop, 8000 ft., 26.-27.x.1934, I 6, 2 2 paratypes,

8500 ft. 1 § paratype, gooo ft. 1 2 paratype, gooo ft. in cedar forest I J, 3 8

paratypes, 10000 ft. 2 2 paratypes (J. Ford and F.W.E.); Naivasha, vii.1937

(H. J. A. Turner) 2 3 paratypes; Nairobi, iii.1935 (Van Someren) 1 9 paratype;

Karura, vi.1937, caught on Lantana (Van Someren), I 9 paratype. ABYSSINIA:

Djem-Djem Forest, ca. 8000-9000 ft.,. 21.-29.ix.1926 (H. Scott) 2 3, 7 Q para-

types; Wouramboulchi, Serpent Lake, ca. goooft. (J. Omer Cooper) 1 9

paratype.

MUSCIDAE: B.—COENOSIINAE I29

Atherigona steeleae sp. n. ¢ 9

Length 4:2-5°5 mm.; of wings 4—5:2 mm.

3§. Head fuscous behind, yellowish in front and below, densely yellowish

to golden-grey dusted, except on median dorsal part of occiput and on inter-

frontalia. The latter dull blackish-brown, usually narrowly rufescent in front,

parallel to very slightly widened anteriorly; parafrontalia with (3—) 4 (—5)

inclinate frontal setae. Outer vertical considerably weaker than the inner

vertical though quite strong, postverticals and ocellars rather strong. Antennae

ferruginous, third joint infuscate along the dorsal edge and towards apex,

not quite thrice as long as wide, arista brown, ferruginous at base. Palpi

yellow. Jowls almost twice as wide as parafrontalia in middle.

Thorax piceous, humeri and pleurae pale, the latter slightly infuscate

below the wings, densely pale yellowish-grey dusted, with at most slight traces

of three narrow vittae. frst acy in four to five rows, the last prst pair slightly

stronger, a pair of distinct prsc. 2 + 4 dc, the two prst pairs rather conspicuous,

the last two pairs strong; scutellum with the lateral sub-basal seta almost a

third as long as the lateral pair which is almost as long as the apical one. Three

propleural setae, two of them quite strong, upper prostigmatal seta as long as

the second propleural. s¢f/ in an equilateral triangle, which bears, in addition,

a few single, small hairs, the lower st// much weaker and the posterior one

slightly stronger than the anterior stpl.

Abdomen pale ferruginous, with not very dense yellowish dust, the first

segment unspotted, the second and third each with a pair of oblong piceous

spots of about half the length of the segment, which do not reach the hind

margin though they are much closer to it than to the anterior end; the first

three segments with rather strong marginal setae, even the fourth with a

marginal row of quite conspicuous setae. Hypopygial prominence (Fig. 29)

stout, large, strongly bi-lobed, subcordiform, when seen from behind, the two

lobes large, depressed and subtriangular, with the outer and anterior angles

broadly rounded, when seen from below, well defined from each other all

round. Stalk of trifoliate process (Fig. 58) fuscous in basal half, ferruginous

in apical half, as are the supports of the lobes and the median lobe, the latter

filiform, slightly dilated and shortly bifurcate at apex in ventral view, quite

filiform in lateral view, lateral lobes dull piceous-black, with the margins of

the ventral surface glossy, half-moon shaped, slightly emarginate at base of

outer margin, the inner margin extending into an incurved, very blunt tooth.

In the male from Kyarumba the lateral lobes are more slender with the apex

much more narrowly rounded than in the males from Kilembe and Kampala.

Legs entirely testaceous. Mid-tibiae with a small fd, hind tibiae with

moderate ad, av and pd setae, the latter slightly beyond the av or level with it.

Wings hyaline, costal and subcostal cells slightly infuscate, the apex

130 RUWENZORI EXPEDITION

extremely narrowly infuscate along the costa from the middle of Rk to m, veins

brown, 74,; and m slightly converging at apex, m straight, 7,,, almost un-

noticeably sinuate, y-m just beyond two-fifths of the discal cell. Calyptrae

yellowish, the lower one slightly infuscate in apical third. Halteres pale

yellowish.

2 similar to male. Head and pleurae more whitish-grey dusted. Third

antennal joint almost four times as long as wide, fuscous, except at base.

Pleurae much darker though the propleurae are pale. Abdomen testaceous to

light ferruginous at sides and base, the first segment with a large, bi-lobed

brownish shadow on dorsum, the second to fourth each with a pair of piceous

elongate spots, which do not reach the hind margin, and a narrow, brownish,

incomplete median vitta, the latter mainly occupying the basal part of the

segments; marginal setae quite distinct though not very strong. Front legs

piceous, except for base and extreme apex of femora and base of tibiae, hind

tarsi slightly infuscate, rest of legs testaceous. Wings not infuscate. Calyptrae

more greyish-hyaline, not distinctly infuscate.

UcGanbDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 3 type, 3 3, 3 para-

types; Kampala, 30.1.1927 (H. Hargreaves) 1 g paratype, I6.vii.19g26 and

6.v.1932 (G. L. R. Hancock) 2 2 paratypes. SUDAN (paratypes): Loti, 19.xi.1933

(one marked 19.1v.1933), ‘in undergrowth in forest to E.’’, “in forest-clearing

along path”, “‘on path downhill to forest, mostly elephant grass’’ (Miss M.

Steele) 5 9; 2.xi.1933 on path to forest, mostly elephant grass (Miss M.

Steele) 1 9; Shendi, 21.xi.1927, bred from dura (Andropogon sorghum) (J. W.

Cowland) 12. ABYSSINIA: Mt. Chilldlo, forest, ca. gooo ft., 14.x1.1926 (H. Scott)

1:9.

One of the males from Kiiembe measures only 3:5 mm., but through the pale

pleurae, wholly ferruginous second antennal joint, and the strongly bi-lobed

hypopygial prominence it points to this species. The lateral lobes of the

trifoliate process are somewhat more slender, the dilatation on the inner side

more obtuse, and the apical half of the median lobe infuscate.

Atherigona longifolia sp. n. 3

Length 3 mm.; of wings 2°8 mm.

Head fuscous behind, translucent yellowish in front and below, densely

whitish-grey dusted, the median dorsal part of occiput glossy fuscous, inter-

frontalia blackish, narrowly rufescent on front margin (teneral?). Interfrontalia

slightly widened anteriorly, parafrontalia with four inclinate frontal setae.

Outer vertical slightly weaker than inner vertical, postverticals and ocellars

rather weak. Antennae fuscous, the margins of the second joint ferruginous,

third joint less than two and a half times as long as wide; arista fuscous,

MUSCIDAE: B.—COENOSIINAE DeSie

slightly paler at base. Palpi yellow. Jowls not quite as wide as parafrontalia

in middle.

Thorax piceous, including dorsal surface of humeral callus, propleura and

ventral surface of humeral callus yellowish, densely pale yellowish-grey dusted,

dorsum with three narrow and very faint vittae which do not reach scutellum.

prst acy in four to five rows, the last frst pair distinctly longer, a pair of small

prsc. 2 + 3—4 dc, the last two pairs moderately strong, scutellum with the

lateral setae slightly weaker than the apical pair, both moderately strong.

Two rather strong propleural setae. s/f/ in a triangle, the posterior side of

which is shorter, and which bears a few setulae, the lower stf/ weak, the anterior

one slightly weaker than the posterior one, both moderately strong.

Abdomen testaceous, almost entirely infuscate and dusted on dorsum, the

first segment with a deeply bi-lobed large piceous spot, the second and third

with a narrow median vitta and a pair of oblong, slightly converging spots

which do not quite reach hind margin, fourth segment entirely ferruginous,

marginal setae rather distinct at the sides of the first to third segments, one

pair of smaller setae on the fourth segment. Hypopygial prominence (Fig. 25)

small, subtruncate, somewhat knoblike, with the angles slightly projecting,

trapezoidal in vertical view. Trifoliate process (Fig. 32) with the lobes very

long and slender, band-shaped, median lobe blade-shaped, not dilated at apex

in ventral view, elbowed in lateral view, lateral lobes gradually slightly curved,

about twice or thrice as wide and almost twice as long as their basal support.

Legs testaceous, apical two fifths of front femora piceous, except for a narrow

yellow apical ring, front tibiae, except base, and the basal 2~3 tarsal joints

strongly infuscate, mid-coxae blackish, except for the margins which are

ferruginous. Mid-tibiae with a small fd, hind tibiae with moderate ad, av and

pd setae, the latter very little beyond the av.

Wings hyaline, the veins testaceous, 74,; and m straight and slightly

converging at apex, 7m slightly before two-fifths of the discal cell. Calyptrae

greyish-hyaline, the border yellowish-white. Halteres yellowish-white.

UGANDA: Ruwenzori, Kilembe, 4500 ft. (’.W.E.) 3 type.

Atherigona budongoana sp. n. 3 2

Length 3-8-4:8 mm. ; of wings 3°6-4:5 mm.

§. Head fuscous behind, yellowish in front and below, densely whitish-

grey dusted, yellowish-grey on parafacialia, the median dorsal part of occiput

glossy fuscous. Interfrontalia dull blackish-brown, narrowly rufescent in

front, parallel, parafrontalia with 4-5 inclinate and one reclinate frontal

setae. Outer vertical much weaker than inner vertical, though quite distinct,

postverticals and ocellars rather strong. Antennae fuscous, third joint not

132 RUWENZORI EXPEDITION

quite thrice as long as wide, the base of the arista rufous. Palpi yellow. Jowls

wider than the parafrontalia in middle.

Thorax: Mesonotum piceous, usually with three narrow brown vittae, its

lateral part, the humeri, apex of scutellum and pleurae pale testaceous, thorax

densely whitish-yellow dusted. frst acy in 4—5 rows, the last prst pair slightly

stronger, a pair of distinct prsc. 2 +. 3—4 dc, the prst pairs comparatively

strong, the last two pairs strong; scutellum with the lateral seta slightly weaker

than the apical one. Two strong and one minute propleural setae. s¢p/ in an

equilateral triangle, which bears a few fine hairs, the lower stf/l much weaker,

the posterior slightly stronger than the anterior one.

Abdomen testaceous to ferruginous, distinctly yellowish-dusted, the first

segment slightly infuscate at apex, the second and third rather strongly in-

fuscate, each with a pair of oblong piceous spots of approximately half the

length of the segment, which do not reach the hind margin, though they

approach it; fourth segment entirely testaceous, marginal setae rather distinct

on the first three segments, fine on the fourth. Hypopygial prominence

(fig. 23) stout, short, trapezoidal, truncate at apex, with the angles slightly

projecting behind and hardly in front. Trifoliate process (Fig. 59) with the

stalk testaceous and the lobes black, largely dull, median lobe linear, slightly

dilated and shortly bifurcate at apex in ventral view, very strongly com-

pressed, broadly fusiform with the apex incurved in lateral view, almost

as wide at widest point (near base) as lateral lobes, the latter band-shaped,

outer margin obliquely truncate at base, broadly rounded elsewhere, inner

margin concavely rounded.

Legs entirely testaceous or—in the specimen from Nyamgasani Valley—

with the front tarsi fuscous and the apex of the front tibiae, the mid and hind

tarsi slightly infuscate. Mid-tibiae with a small fd, hind tibiae with moderate

ad, av and pd setae, the latter slightly beyond the av.

Wings hyaline, the veins testaceous at base, brown at apex, 74,; and m

straight and very slightly converging in apical part, 7m slightly beyond two-

fifths of the discal cell. Calyptrae pale yellowish, the lower one slightly infuscate

in apical third, halteres white (yellowish-white in specimen from Nyamgasani

Valley):

@ similar to male. Parafacialia whitish dusted. Pleurae largely though

very slightly infuscate, appearing pale, humeri and propleurae entirely pale.

Antennae as in male. Abdomen testaceous with some brownish shadows on

the second segment and a pair of round spots on the third, or more or less

infuscate on the dorsum and with large fuscous spots, a bi-lobed one on the

first, a pair of elongate spots on the second, a pair of narrow, elongate spots

on the third, and a pair of small, round, brown spots on the fourth segment.

Legs testaceous, the apical half or third of the front femora (except for the

extreme apex), the apical three-fifths of the front tibiae, the front and hind

MUSCIDAE: B.—COENOSIINAE 133

tarsi strongly infuscate, mid-tarsi slightly infuscate. y-m at two-fifths of

discal cell.

UGANDA: Budongo Forest, 7.-8.i1.1935 (F.W.E.) 3 type, 23 paratypes,

2 2 paratypes; Ruwenzori, Nyamgasani Valley, 6400 ft. (D. R. Buxton) 1 3;

Busana, 22.x.1933 (I. W. Chorley) 1 9. Kenya: Solai Dist., Sonje Valley,

Laikipia Escarp. 12.ix.1919 (7. J. Anderson) 1 9. The specimen from Busana

is somewhat greasy and its identity not quite certain.

There is a single male from Ruwenzori: Mobuku Valley, 7300 ft. (F.W.E.),

in which the pleurae are mainly dark, the hypopygial prominence somewhat

more outstanding (though very similar in shape), the outer lobes of the trifoliate

process not concavely rounded at interior side but slightly convex (slightly

concave only near basal support), and the median lobe simple at apex, shorter,

not incurved at apex, and more slender in basal half. This is no doubt a local

form of budongoana.

[Atherigona truncata sp. n. 3]

Length 3-4 mm.; of wings 3 mm.

Head fuscous behind and above, pale ferruginous in front and_ below,

densely grey dusted, except on median dorsal part of occiput, parafrontalia

and frontal triangle, which are glossy fuscous. Interfrontalia dull fuscous,

subparallel, parafrontalia with four inclinate setae, the first of which is small.

Outer vertical distinct, though not very strong, inner vertical moderately

strong, postverticals and ocellars well developed. Antennae ferruginous,

third joint slightly infuscate along dorsal edge, very large, almost as long as

the head high, though not quite thrice as long as wide, arista long, light brown

near base, fuscous at apex. Palpi yellow. Jowls as wide as parafrontalia in

middle.

Thorax ferruginous, mesonotum and scutellum piceous, except for lateral

presutural part, grey dusted without distinct vittae, apex of scutellum narrowly

ferruginous. prst acy in 4-5 rows, the last prst pair longer, post acr in six rows,

2 + 3 dc, only the last two pairs rather strong; lateral sub-basal seta of

scutellum almost a third as long as the lateral seta which is strong though

moderately long and equal to the apical one. Two rather strong propleural

setae and a minute one. s¢p/ in an equilateral triangle which bears 1-2 minute

hairs, lower stfl rather weak, anterior moderately strong, posterior one

rather strong.

Abdomen ferruginous with paler base, second segment with a pair of

rather large oblong piceous spots which do not reach front nor hind margin,

third with a pair of small, roundish spots very close to hind margin though

not quite reaching it, fourth with a pair of similar, still smaller spots, lateral

marginal setae well developed on all the four segments. Hypopygial promi-

nence (Fig. 8) rather strongly projecting, knob-shaped with sub-truncate apex.

134 RUWENZORI EXPEDITION

Trifoliate process (Fig. 45) black, median lobe filiform, the apex shortly clavate

with the normal minute hairs, more long-clavate in lateral view, lateral lobes

narrow and hardly dilated in basal half, the apical half semi-circular.

Legs testaceous, the front tibiae slightly infuscate in apical half, front tarsi

strongly infuscate, less so towards apex, hind tarsi slightly infuscate. (Mid-

legs missing in type). Hind tibiae with small ad, av and pd setae, the latter

almost at the same level.

Wings hyaline, veins testaceous, 7,,; and m distinctly converging at apex,

v—m slightly before two-fifths of the discal cell. Calyptrae hyaline with pale

testaceous border. Halteres pale-yellow.

KENYA: Rabat, viii.1930 (Van Someren) 3 type.

Atherigona pedunculata sp. n. ¢

Length 4-2 mm.; of wings 4:4 mm.

Head fuscous on frons and occiput, yellowish in front and below, densely

grey dusted, except on median dorsal part of occiput which is glossy fuscous,

as are the parafrontalia and frontal triangle. Interfrontalia dull black, sub-

parallel, parafrontalia with 4-5 inclinate (one or two of them small) frontal

setae. Outer vertical much weaker than inner one, though quite strong,

postverticals and ocellars rather strong. Antennae ferruginous, the third

joint very long and broad, more than thrice as long as wide, the dorsal margin

and apex infuscate, the latter rather broadly, arista long, longer than the

head is high, fuscous, its basal quarter ferruginous. Palpi yellow. Jowls one

and a half times as wide as the parafrontalia in middle.

Thorax piceous, humeri and pleurae largely yellowish, only partly and

slightly infuscate, yellowish-grey dusted on dorsum, with a pair of very narrow

brown vittae along the dc and a somewhat broader median brown vitta, which

is slightly dilatate to hind end but does not reach the scutellum; pleurae

whitish-grey dusted. rst acr in five rows, the last prst pair distinctly longer,

a distinct pair of prsc. 2 + 4 dc of which only the last two pairs are strong;

scutellum with the lateral sub-basal seta almost a third as long as the lateral

seta, the latter subequal to the apical pair. Three propleural setae of which

two are rather strong, the upper prostigmatal rather strong though conspicuously

weaker than the second propleural. stf/ in an almost equilateral triangle,

which bears a few fine setulae, lower stfl considerably weaker than the two

others.

Abdomen ferruginous, with the base testaceous, the first segment with a

large piceous spot which reaches hind margin and is as broad as the dark pattern

on the second segment, the latter entirely piceous on dorsum except for a

lateral margin of about one eighth of the segment-width on either side, this

stripe densely whitish-grey pollinose, third segment with a pair of oblong

MUSCIDAE: B.—COENOSIINAE 166}

piceous spots which slightly converge in front and are fused except behind,

fourth segment entirely ferruginous, marginal setae rather strong on segments

1-3, finer though very distinct on fourth. Hypopygial prominence (Fig. 7)

knob-shaped, rather large. Trifoliate process (Fig. 4g) black, median lobe fili-

form, very slightly dilated at apex in ventral view, with the apical setae hair-

like, longer and more numerous than usual, slightly compressed, straight and

gradually very slightly dilated towards apex in lateral view, lateral lobes with

the basal part filiform, stalk-like, as long as the apical part which is broadly

axe-shaped.

Legs testaceous, the apex of the front tibiae very slightly and the first to

third joints of the front tarsi strongly infuscate. (Mid-tibiae missing), hind

tibiae with rather strong ad, av and fd setae, the latter hardly beyond the

av, whereas this is inserted rather far from ad.

Wings hyaline, slightly smoky, the veins testaceous, very narrowly infuscate

along 7,,; and m at apex, these veins distinctly converging and slightly sinuate,

the apex of the wing truncate and rectilinear between them, 7m at two-fifths

of the discal cell. Calyptrae hyaline, slightly smoky, the border testaceous.

Halteres yellowish.

UGANDA: Ruwenzori, Namwamba Valley, 6500 ft. (FE. G. Gibbins) 3 type.

Atherigona triangularis sp. n. 3

Length 4 mm.; of wings 3-8 mm.

Head fuscous behind, testaceous in front and below, grey dusted, the median

dorsal part of the occiput glossy fuscous, parafrontalia glossy black, temples,

jowls and face (incl. parafacialia) golden dusted. Interfrontalia dull, almost

velvety blackish-brown, parallel, parafrontalia with 2—3 inclinate frontal setae.

Outer vertical evidently very weak, inner vertical strong (the pores of both

only present in type), postverticals moderate, ocellars rather long. Basal joints

of antennae reddish-brown, third joint pale ferruginous, with the apex broadly

infuscate, less so on ventral side, two thirds as long as head high, thrice as long

as wide, base of the arista testaceous. Palpi yellow. Jowls (at narrowest

point) almost as wide as parafrontalia in middle.

Thorax piceous, with yellowish humeri and propleurae, densely grey dusted,

humeri and sides more golden dusted, mesonotum with a brown median vitta,

which broadens behind, reaches apex of scutellum and occupies all the latter

except for a narrow outer stripe. rst acy in four to five rows, a pair of distinct

prsc, 2 + 4 dc, only the last two pairs rather strong. Two strong and a small

propleural setae. s¢f/ in a triangle which bears a few fine short hairs and the

posterior side of which is slightly shorter, the lower s¢f// much weaker.

Abdomen ferruginous, the first segment with a suffused piceous median

spot, second segment with a pair of large, oblong piceous spots which do not

136 RUWENZORI EXPEDITION

reach the hind margin, third with a pair of small, roundish, slightly oblong

piceous spots towards hind margin, fourth with a pair of small, roundish,

slightly transverse piceous spots; marginal setae quite strong on first three

segments, very distinct though rather fine on fourth segment. Hypopygial

prominence (Fig. 11 a, 6) almost knoblike, moderately large, slightly broadened

to apex which is somewhat convex in posterior view and little wider than long

in vertical view, with the posterior angles slightly marked, the anterior ones

rounded. Trifoliate process (Fig. 42 a, 6) purplish-black, median lobe very

strongly depressed, very broadly triangular in more than apical half, broader

than the lateral lobes, the apex truncate and the apical angles prolonged and

upcurved, the lateral lobes axe-shaped, rather broad, with a short stalk.

Legs broken, except for coxae and trochanters which are entirely pale

ferruginous. ;

Wings hyaline, veins testaceous, brown towards apex, 7,,,; and m very

narrowly and not very conspicuously infuscate at apex, slightly converging,

y—-m slightly beyond two-fifths of the discal cell. Calyptrae greyish-hyaline,

the border yellowish. Halteres yellowish-white.

UGANDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 3 type. The specimen

is poor, all the legs being broken. Unfortunately the abdomen and trifoliate

process were damaged during the dissection of the latter.

Atherigona aurifacies sp. n. ¢

Length 3-8 mm.; of wings 3-6 mm.

Head fuscous on frons and occiput, yellowish in front and below, grey dusted

on occiput, frontal triangle and posterior two thirds of parafrontalia, glossy

fuscous on median dorsal part of occiput, golden dusted on anterior part of

parafrontalia, on face and parafacialia, yellowish-grey dusted below. Inter-

frontalia dull dark brown, slightly and evenly dilated in front, parafrontalia

with four inclinate frontal setae. Outer vertical rather strong, slightly weaker

than inner vertical (postverticals lost) ocellars rather strong. Antennae fuscous,

second joint with reddish margins, the third joint long and broad, two and a

half times as long as wide, arista fuscous. Palpi fuscous. Jowls one and a half

times as wide as parafrontalia in middle.

Thorax piceous, humeri and propleurae pale ferruginous, yellowish-grey

dusted on dorsum (specimen greasy). pyrst acy in five rows, the last frst pair

distinctly longer, a distinct pair of prsc. 2 + 3 dc, the prst ones rather distinct,

the last two strong, scutellum with the lateral and apical setae subequal, strong.

One strong, one moderate and one minute propleural seta, upper prostigmatal

seta moderately long though weaker than the second propleural. s¢p/ in an

almost equilateral triangle, which bears one or two fine setulae, lower stl

much weaker than the two others.

MUSCIDAE: B.—COENOSIINAE 137

Abdomen light ferruginous, the second segment with a pair of small,

brown, suffused spots; marginal setae rather strong on segments I-3, fine on

the fourth. Hypopygial prominence (Fig. 9) simple, knob-shaped, rather

large. Trifoliate process (Fig. 40 a, 6) with the apex of the stalk and the

support of the lobes whitish, the lobes black, median lobe filiform, clavate at

apex in ventral view; lateral lobes suboval with the inner margin very slightly

emarginate, outer margin broadly though very shallowly emarginate near

base, at the inner end of the emargination a keel arises which reaches inner

part of apex, the disc of the lobe slightly hollowed out outward of the keel.

Legs testaceous, front femora somewhat brownish near apex, front tibiae

infuscate, except near base, front tarsi strongly, the others slightly infuscate.

Mid-tibiae with a moderate fd, hind tibiae with moderate ad, av and pd setae,

the latter considerably beyond the av.

Wings hyaline, veins testaceous, 7,,; and m slightly converging at apex,

their apex straight, v-m slightly before two-fifths of discal cell. Calyptrae

yellowish. Halteres pale yellow.

UGANDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 3 type.

Atherigona lineata ugandae ssp. n. ¢ 9°

Characters the same as in /éneata, but slightly smaller, 3-2—3-9 mm., whereas

the female types of lineata, one of which I have seen, are 4 mm., and the speci-

mens of both sexes from Salisbury (typical locality of lineata Ad.) at hand,

collected by Mr. Alex. Cuthbertson, measure 3°8—4:6 mm. Front femora entirely

yellow in male of ssp. wgandae and the front tibiae less strongly infuscate,

whereas most of the apical half, except for a narrow apex, of the posterior

surface of the front femora is piceous in the typical form. The first abdominal

segment of the latter is much broader and the second therefore more narrowed

posteriorly than in ugandae, the branches of the hypopygial prominence

(Fig. 18, viz. 16) aresomewhat longer and more diverging inthe latter, and the basal

support of the lateral lobes of the trifoliate process (Fig. 39, viz. 36) isnot distinctly

stalk-shaped, only half as long as the lobe and mainly black, whereas it is

distinctly longer than the lobe and blackish only along interior margin in the

typical form. The humeri are black and covered with dense grey dust in both

forms. The only character I find for discriminating between the females of

the two forms (besides the slight difference in the average size) is the colour

of the apical one to three joints of the mid and hind tarsi: these are usually

more or less distinctly paler in the females of the ssp. wgandae. The pores of

the frontal setae are sometimes surrounded by a narrow glossy area, but the

parafrontalia are none the less mainly dusted.

UcanpbA: Kigezi District, Mabungo Camp, 6000 ft., xi.1934 (J. Ford)

6 type, 9 2 paratypes; Arua, 1919 (R. E. McConnell) 4 2 paratypes; Busana,

138 RUWENZORI EXPEDITION

22.x.1933 (T.W. Chorley) 9 9, in these specimens the humeri are partly yellowish.

KenyA: Karura, vi.1937, caught on Lantana (Van Someren), 1 2 paratype;

Nairobi, vi.1g28 (Van Someren) 1 2 paratype (in Coryndon Mem. Mus.).

[Atherigona binubila sp. n. 3 9]

Length 3-7-4-7 mm.; of wings 3-3-4°3 mm.

3. Head fuscous behind and above, testaceous in front and beneath,

interfrontalia dull black, median dorsal part of occiput glossy fuscous, rest of

head densely pale grey dusted. Interfrontalia parallel or very slightly dilated

anteriorly, parafrontalia with four inclinate setae, the first of them small.

Inner vertical rather strong, outer vertical much weaker, postverticals and

ocellars rather strong. Antennae with the basal two joints ferruginous, the

second somewhat infuscate, third joint fuscous, less than thrice as long as

wide, arista fuscous with brown base. Palpi fuscous. Jowls distinctly narrower

than the parafrontalia.

Thorax piceous, with the humeri and propleurae testaceous, densely grey

dusted, the pollinosity with some golden reflections on the pleurae, dorsum

with three quite conspicuous though not very broad brown vittae, the sub-

lateral ones slightly narrower. frst acy in 4—5 rows, the last prst pair distinctly

longer, 2 + 3—4 dc, only the last two pairs rather strong, scutellum with the

lateral sub-basal seta fine though almost of a third the length of the lateral

seta, the apical one slightly longer than the latter. Two strong and one minute

propleural. stf/ in an almost equilateral triangle which bears 1-2 small hairs,

lower stpl considerably weaker, posterior stp/ slightly longer than the anterior

one.

Abdomen testaceous, first segment with a brown to piceous median vitta

from base to four-fifths and a transverse band shortly before hind margin

which is incomplete at middle and sides but the anterior inner end of which is

often broadly connected with the median vitta, second segment with a brown

median vitta which is incomplete behind and a pair of rather large, piceous

inverted-triangular spots, third segment with a short trace of a median vitta

at base and a pair of rather small, oblong, piceous spots, fourth segment

without distinct spots. Hypopygial prominence (Fig. 10) rather strongly

projecting, transversely knob-shaped. Trifoliate process (Fig. 33 a, 6) with

the stalk and base testaceous, the lobes piceous-black, median lobe linear at

base, slightly thickened and filiform behind middle, apex slightly bifid in

ventral view, narrowly bandshaped and strongly incurved in lateral view;

lateral lobes boomerang or elongate-banana-shaped with a long emargination

in basal third of outer margin.

Legs pale ferruginous, the front femora somewhat infuscate in apical half,

except for the extreme apex, front tibiae strongly infuscate in apical half,

MUSCIDAE: B.—COENOSIINAE 139

- front tarsi piceous at base and becoming gradually light brown to the last two

joints. Mid-tibiae with a very small fd, hind tibiae with moderate ad, av and

pd setae, the latter slightly beyond the av or level with it.

Wings hyaline with a brown suffusion at the base of the costal cell beyond

h, which extends to the first and second basal cells, and a yellowish-smoky

suffusion between the apices of sc and 7,. m-m slightly suffused with yellow.

74,5 and m distinctly converging at apex, v-m between one-third and two-fifths

of discal cell. Calyptrae whitish, slightly smoky towards margin, the latter

pale yellowish. Halteres whitish.

2 with the third antennal joint more than thrice as long as wide. Shape

of the abdomen rather short-oval, the spots of the third segment inverted-

triangular, fourth segment with a pair of roundish, slightly oblong spots, the

ground-colour of the third and fourth segments and more or less the second

grey, though densely yellowish dusted. Front legs piceous except for the base

of the femora and the knees, hind tibiae and mid-tarsi infuscate, hind tarsi

strongly infuscate. Wings entirely hyaline.

Kenya: Nairobi, ix.1937 (Van Someren) 3 type; vi.1928 I 3 2 2 paratypes;

Karura, vi.1937 caught on Lantana (Van Someren), 1 $22 paratypes; Naivasha,

vii.1937 (H. J. A. Turner) i 8 paratype (paratypes partly returned to Coryndon

Memorial Museum); Ruiru, 19.viil.1932, on coffee (H. C. James) 1 9 paratype.

Atherigona piceiventris sp. n. 3 2

Length 2:8-4:1 mm.; of wings 2-6-3°7 mm.

3$. Head fuscous behind and above, yellowish in front and below, densely

yellowish-grey dusted, the parafacialia golden dusted, median dorsal part of

occiput glossy fuscous. Interfrontalia dull black, slightly dilated in front,

parafrontalia with four inclinate frontal setae. Outer vertical rather short

though quite strong, inner vertical strong, postverticals moderate, ocellars

rather long. Antennae fuscous, third joint two and a half times as long as

wide, arista fuscous. Palpi fuscous. Jowls distinctly wider than the para-

frontalia in middle.

Thorax piceous with the ventral surface of the humeral callus and the

propleura yellowish, densely yellowish-grey dusted, dorsum with three dark

brown vittae. prst acy in 4-5 rows, the last frst pair slightly stronger, a pair

of distinct prsc. 2 + 4 dc, the prst pairs distinct though small, only the last

two pairs well developed; the lateral pair of scutellar setae conspicuously

shorter than the apical pair, though quite strong. Two rather strong propleural

setae. stp/ in an almost equilateral triangle, the posterior side of which is

slightly shorter, and which bears a few single fine setulae, the lower sé// much

weaker than the others which are subequal.

Abdomen testaceous, the dorsum of the first and second segments almost

II, 4 (d)

I40 RUWENZORI EXPEDITION

entirely piceous, sides and hind margin narrowly testaceous, third segment

ferruginous with a pair of oblong piceous spots and the dorsum infuscate -

between them, fourth segment ferruginous, the dark parts rather sparsely

brown dusted with a rather broad median vitta of grey dust, the hair of abdomen

rather long, setae not very strong. Hypopygial prominence (Fig. 17) not very

large, gradually slightly dilated to apex which is emarginate, the two ensuing

slight lobes rounded, in ventral view the prominence is elongate bean-shaped.

Trifoliate process (Fig. 47) black, except for the stalk which is brown with

testaceous apex, median lobe narrow, gradually slightly dilated to apex which

is truncate, in lateral view the median lobe is slightly compressed, somewhat

narrowed at apex, lateral lobes inverted-ovai, broadly emarginate at base of

exterior margin.

Legs testaceous, the apical half of the front femora somewhat infuscate,

front tibiae piceous except at base, front tarsi piceous, the last two or three

joints paler. Mid-tibiae with a small fd, hind tibiae with small ad, av and pd

setae, the last two on the same level.

Wings hyaline, very slightly smoky, the veins brown, 7,,, and m straight

and very slightly converging at apex, 7-m at two-fifths of the discal cell. Calyp-

trae yellowish-smoky, the border testaceous, halteres pale yellowish.

9°. Third antennal joint thrice as long as wide. Propleurae and humeral

callus entirely yellow to entirely piceous. Dorsum of abdomen almost entirely

piceous, even on third and fourth segments, the spots of these two segments

usually on a strongly infuscate greyish-dusted ground, seldom more distinct

on a lighter ground, in one female (Mabungo Camp) all the segments show a

pair of elongate spots and a median vitta on ferruginous ground. Front

femora, except for a narrow basal part and the extreme apex, and front tibiae,

except for the base, and front tarsi piceous, hind tibiae and tarsi and sometimes

mid-tarsi infuscate.

Ucanpa: Mt. Elgon, between Butandiga and Bulambuli, 8000 ft. 7.vili.1934

(J. Ford) 3 type 2 2 paratypes; Kigezi District: Mabungo Camp, 6000 ft.,

resp. 18. and 21.x1.1934 (J. Ford) 1 9 paratype, I 2 with paler abdomen, see

above. KrENyA: Chyulu Hills, 5600 ft., vi.1938 (Coryndon Mus. Exped.)

22 paratypes; Kiambu, 25.vi.1931 (R. H. Le Pelley) 1 2 paratype.

Atherigona griseiventris sp. n. 3 2

Length 4-5 mm.; of wings 4:2—4°5 mm.

3. Head fuscous behind, pale yellowish in front and below, densely yellowish-

grey to golden (on parafacialia) dusted, except on median dorsal part of occiput,

which is glossy fuscous, and on interfrontalia which are dull black, subparallel.

Parafrontalia with (3-) 4-5 (—6) inclinate frontal setae. Outer vertical rather

strong, slightly weaker than the inner vertical, postverticals and ocellars rather

MUSCIDAE: B.—COENOSIINAE I4t

strong. Antennae fuscous, third joint not quite thrice as long as wide, arista

fuscous. Palpi fuscous. Jowls almost twice as wide as the parafrontalia in

middle.

Thorax piceous, densely yellowish-grey dusted, humeri and propleurae

yellowish, the latter golden dusted, dorsum with three narrow brown vittae,

the lateral ones slightly wider than the median vittae. rst acy in 4-5 rows,

the last frst ones not stronger, a pair of distinct prsc. 2 + 3—4 dc, only the

last two pairs strong; scutellum with the lateral setae distinctly shorter than

the apical pair. Two to three propleural setae, two of which are quite strong,

upper prostigmatal bristle rather long though very fine and much shorter than

the second propleural. s¢f/ in an equilateral triangle, which bears a few single

small hairs, the lower s¢// much weaker, the posterior slightly stronger than the

anterior one.

Abdomen ferruginous, dorsum of the first and third segments slightly, of

the second rather strongly infuscate and rather densely whitish-grey dusted,

first segment with a large bi-lobed piceous spot, the two lobes of which almost

reach the hind margin, second segment with a pair of elongate piceous spots

of almost the length of the segment, which do not, however, quite reach the

hind margin, and a faint incomplete median vitta in basal half; third segment

with a pair of small, rounded spots close to hind margin; fourth segment

entirely ferruginous, distinctly dusted; the first three segments with the

marginals distinct though not very strong, the fourth with a few longer hairs

along hind margin. Hypopygial prominence (Fig. 12) stout, not very broad,

subcordiform, broadly emarginate at apex, bean-shaped in vertical view, with

the emargination towards front end. Trifoliate process (Fig. 38 a, b) black,

narrowly pale yellow at furcation, with the lateral lobes short-banana-shaped,

strongly emarginate at base, the central part broader (in lateral view) than

the lateral lobes, broadly claw-shaped, and filiform with a short terminal club

in ventral view.

Mid-legs entirely testaceous (the others missing, except for coxae and

trochanters, which are pale ferruginous), mid-tibiae with a small fd seta.

Wings hyaline, veins brown with more or less testaceous bases; 74,; and

m distinctly converging though straight at apex, y-m on two-fifths of discal

cell. Calyptrae hyaline, the lower one somewhat whitish, halteres pale

yellowish.

2 similar to male. Parafacialia whitish-grey dusted. Third antennal joint

thrice as long as wide. Humeri and propleurae slightly infuscate, the last pair

of prst acy distinctly stronger. Abdomen almost entirely infuscate and rather

densely greyish dusted, first segment with a very large piceous spot, which is

notched behind on median line, second with a pair of slightly oblong spots of

almost the length of the segment and an incomplete faint median vitta, third

with the piceous spots more elongate, pattern otherwise similar; on both these

142 RUWENZORI EXPEDITION

segments the median vitta is connected in front with the spots, and the latter

do not quite reach the hind margin and approach each other slightly behind,

so that they enclose a large bi-lobed median extension of the grey-dusted hind

margin; fourth segment with a short rudiment of the median vitta at base

and a pair of small, oblong spots towards apex; marginal setae distinct only

on the sides of the first to third segments. Front femora, tibiae and tarsi

piceous, except for extreme base of femora and knees; mid and hind tarsi

strongly, hind tibiae slightly infuscate; the latter with moderate ad, av and

pd setae, pd slightly beyond the av. y-m not much beyond basal third of

discal cell.

Kenya: Aberdare Range, Mt. Kinangop, 8000 ft., x.1934 (F.W.E.) 3 type;

gooo ft., xi.1934 (F.W.E.) 9 paratype; Nyeri Track, 10500-11000 ft., x.1934

(J. Ford) 2 Q paratypes; Chyulu Hills, 5200 ft., iv—v.1938, 4 2 (Coryndon

Museum, Nairobi, and B.M.). The specimens from the Chyulu Hills are

slightly different: the lateral vittae of the thorax are not so markedly wider

or not wider at all than the median vitta, and the abdomen is not so much

infuscate and of a more testaceous ground-colour.

Two females (UGANDA: Kigezi District, Mabungo Camp, 6000 ft, 18..xi.1934

(J. Ford)) are quite similar to the Chyulu specimens, but in one of them the

piceous spots are rather large so that only a narrow bifurcate tongue separates

them in the middle. As they are only slightly more than 4 mm. long, these

specimens may perhaps be pale females of piceiventris.

Atherigona tridentata Mall.

UGANDA: Ruwenzori, Namwamba Valley, 6500 ft. (F.W.E.) 1 3; Masindi

District: Budongo Forest, 7.-8.11.1935 (F.W.E.) 1 9.

Typical locality: Kenya: Nakuru. In B.M. from: UGANDA: Segibwa

Falls, 29.vil.1935, attracted by human faeces (£. G. Gibbins); KENYA: Kabete

(T. J. Anderson), Naivasha (H. J. A. Turner); seen from Kiambu, 24.vi.1931

(R. H. LePelley) and Nairobi, vii.1937 (Van Someren).

I have not seen the type nor a specimen that entirely fits the description.

The single male at hand has the hypopygial prominence (Fig. 22) very broad

and subtruncate, in posterior view the lateral angles protrude as small teeth

which are connected by a broad and rounded hump, in ventral view it is

trapezoidal with the anterior and posterior angles protruding as small teeth.

The trifoliate process (Fig. 43) is entirely purplish-black, except for a hyaline,

pale blade on median lobe; this lobe is linear in ventral view, somewhat

compressed and slightly narrowed to apex in lateral view, the hyaline blade

as wide near apex as the lobe, the extreme apex of the latter pale, lateral lobes

semicircular. The female has not been described, and there is no female from

the same locality as the male. Those females which I include in this species

MUSCIDAE: B.—COENOSIINAE 143

exhibit a fair amount of variation, which may, however, be due to geographical

and in part individual variation. Pending the accumulation of a richer material

of this species or group of species it would certainly not seem warrantable to

describe these forms.

Microcalyptra sp. ».

Kenya: Mt. Elgon, heath zone, 10500-11500 ft., 11.1935 (F.W.E.).

A single, evidently male specimen, without hind legs or abdomen, will be

described together with more specimens of this and two other species from

Abyssinia in my report on Dr. Scott’s collection.

Andersonosia nom. n.

Andersonia Mall., 1928, Ann. Mag. Nat. Hist. (10)1: 465 (non. Boul. 1900, non Strickland

19It).

After reading this report, Dr. F. W. Edwards drew my attention to the fact

that Andersonia Mall. is several times preoccupied. I have, therefore, slightly

altered the name given by Malloch.

Andersonosia velutinifrons Mall.

Kenya: Aberdare Range, Mt. Kinangop (west side), 8000 ft., x.1934

(AVE E) 13.

Typical locality: East side of edge of forest, Aberdare Mts., 7300 ft. (type

in B.M.).

The type is a female, and the male has not yet been described. It is similar

to the female, all the generic characters recorded by Malloch are present, and

so are the specific characters of the female. The parafrontalia of both sexes

are finely haired on anterior half. In the female the longest hairs of arista are

about half as long as the greatest width of the third antennal joint, in the

male they measure only a quarter of that width, this being partly due to the

greater width of the antennal joint, partly to real shortness of the hairs. Third

antennal joint longer and wider than in female, practically reaching lower

margin of eyes and mouth-margin. The lower stpl in both sexes is almost

twice as close to the posterior as to the anterior one. Hind femora of male with

2-3 strong av setae near middle besides the preapical. v—m slightly beyond

middle of discal cell in male. Lower calyptra with yellowish border in male,

with yellowish-white border in female. Abdomen parallel-sided, long and

slender, depressed, glossy blackish brown, very sparsely dusted, only the

anterior angles densely grey dusted, a pair of faint paramedian vittae of sparse

greyish dust, close to each other, run over the first three segments. Length

4.5 mm.

144 RUWENZORI EXPEDITION

Andersonosia semifumosa sp. n. 3 9°

Length 5-8 mm.; of wings 6°5 mm.

Head black, densely grey dusted, the median dorsal part of the occiput

brown dusted, interfrontalia velvety black, with strong brown reflections

when seen from in front, parafrontalia grey dusted, with a spot of brown dust

just behind level of lunula, the anterior end of the parafrontalia, the parafacialia

and face silver-white dusted. Frons as wide at vertex as one eye, considerably

widened to base of antennae, as wide in front as long, interfrontalia parallel in

posterior half, rounded-dilated at anterior inclinate seta and constricted at

base of antennae, where the parafrontalia are half as wide, whereas they are

a quarter as wide as the interfrontalia in middle; two strong inclinate frontal

setae. Outer vertical weak, inner vertical very strong, postocellars very long,

ocellars half as long as frons. Antennae fuscous, inserted at dorsal fifth of

eyes, where the frons protrudes in profile by the width of the third antennal

joint, the latter very long, more than thrice as long as wide, almost reaching

mouth-margin, which is level with the lower margins of the eyes, arista short-

haired above, very short-haired beneath, the longest hairs above about twice

as long as the arista wide at its widest point. Palpi black. Jowls not quite as

wide as third antennal segment, parafacialia as wide at narrowest point as the

arista near. base.

Thorax black, densely grey dusted, mesonotum largely glossy black with

slight brown dust, the shoulders and lateral parts of the mesonotum in front

of suture silver-grey dusted. Anterior half of the postsutural part of mesonotum

entirely dark, the dark pattern extending backward in a broad tongue between

the dc; scutellum largely brown dusted, grey dusted at apex. Lower prostig-

matal seta replaced by a few minute hairs. acr hairs practically absent, I +3

strong dc, two strong post za, one strong sa, two strong postalar setae, anterior

notopleural strong, posterior one small. Scutellum with two or three pairs of

hairs and two pairs of subequal strong setae. s¢/ in a triangle, the posterior

side of which is less than half as long as the upper one, anterior stpl weak,

lower stpl very weak, posterior stfl very strong, longer than any other seta.

Abdomen grey dusted on venter and at base of lateral surface, dorsum

almost wholly black and glossy; parallel and slender, somewhat depressed and

arched, the sides with distinct setae and a number of long hairs besides the

short appressed hairs, fourth segment with a transverse discal and marginal

row, each consisting of three pairs of rather strong setae.

Legs piceous, except all the trochanters and tibiae and the basal three-fifths

of the mid and three-fourths of the hind femora which are yellow. Front

femora with a complete row of about six fu and seven fd setae, the most

distal two pairs of fv distinctly stronger than the others, front tibiae with a

rather short and strong submedian pv seta; mid-femora with fine hairs and two

MUSCIDAE: B.—COENOSIINAE 145

preapical (pd and fp) setae, mid-tibiae with two pd setae; hind femora with a

single strong preapical av, about four ad and a fd preapical, hind tibiae with

two ad and a small av, the latter slightly more basad than the second ad seta

which is inserted on second third; there are, besides, one or two longer, appressed

setae distinguishable among the appressed hairs near apex.

Wings long and broad, the apical half brownish, the basal half hyaline, the

border vague, veins brown, 7,,, and m slightly converging at apex, v-m just

_ beyond middle of discal cell, the latter very long, apical section of ms ,cu,

much shorter than m—m. Calyptrae white, with white margin and fringe, the

lower calyptra strongly projecting. Halteres pale yellow.

UGANDA: Ruwenzori, Namwamba Valley, 8300 ft. (.W.E.) 3 type.

Very similar to velutinifrons Mall., but the apical half of the wings is infuscate,

vein 7-m just before middle of the discal vein, border of the calyptrae white,

front femora with a complete row of fv setae (this character at least will

probably distinguish the female of semzfumosa from that of velutinifrons),

hind femora without submedian av setae, even in male, abdomen without

paramedian vittae of greyish dust.

Coenosia Meig.

Whereas in Europe and North America the bulk of the species so far united

in Coenosia are true members of this genus and only a few belong to Caricea,

the latter genus comprises the great majority of the Ethiopian Coenosta and

in fact of the Ethiopian Coenosiinae. Only five species of the large material

worked out have been found to belong to Coenosia, and I am not able to identify

any one of them with a known species. Besides the character of the hind

tibial setae there seems to be a more or less marked difference in the head,

which is considerably longer and more subcubical or trapezoidal in Coenosta.

In all the following species the head is not much higher than long, the arista

is very shortly pubescent or almost bare, the scutellum bears the normal two

pairs of setae, the halteres are yellow, the front tibiae have a pv seta and the

mid-tibiae an ad and a pd bristle.

1(4). Legs entirely or almost entirely yellow. av seta of hind tibiae absent in

male (with very rare individual exceptions). Ocellar bristles moder-

ately developed, moderately diverging.

2(3). Hind tibiae of both sexes without a submedian pd seta. Lower calyptra

normal, more or less broadly rounded at apex, strongly protruding

beyond upper one. Coxae black.! Frons rounded anteriorly in profile,

very slightly protruding. Antennae short, apex falling far short of

level of lower eye-margins . 5 ; : . auyvifacies sp. n., p. 146

1 Front coxae yellowish to brownish, femora black on apical third to one half (whereas

they are entirely yellow in Coenosia aurifacies): see (Xenocoenosia) africa Curr. (1935,

Amer. Mus. Novit. 776: 12). According to information received from Dr. Curran, this

species traces to Coenosia aurifacies but is at once separated by the colour of the legs. As

the arista is said to be ‘“‘rather long pubescent above’’ and the hind tibiae are said to have

a ‘“‘long”’ ad, I should have believed this species to fall within Cavicea rather than Coenosia.

146 RUWENZORI EXPEDITION

3(2). Hund tibiae of both sexes with a small submedian pd seta. Lower calyptra

small, subtriangular, and only slightly projecting beyond the upper one.

Front coxae yellow, the others mainly dark. Frons angular in profile,

protruding by almost the width of the third antennal joint. Antennae

rather long, apex almost reaching level. of lower eye-margins. Face

grey (f. typica) or brown-dusted and apical infuscation of posterior

femora shorter but stronger (ssp. byunneigena, p. 149).

hetevocnemis sp. n., p. 148

4(1). Legs entirely black. av seta of hind tibiae present in both sexes (¢ muicro-

calyptva unknown). Ocellar bristles very strong, strongly diverging

(Fig. 63). Lower calyptra small to very small

5(6). Hind tibiae with a distinct pd seta. Calyptrae very small, the lower one not

projecting. Abdomen with a median vitta and paired spots. Femora

with rather few outstanding setae, the ad row on hind femora consisting

of five to six setae : ‘ : : : microcalyptra sp. D., p. 150

6(5). Hind tibiae without a pd seta. Calyptrae small, though much more con-

spicuous, the lower one slightly projecting. Abdomen entirely dark

dusted on dorsum or with pale dusted margins. Femora with more

numerous outstanding setae, the ad row on hind femora consisting of

seven or more setae.

7(8). Three or four pairs of inclinate frontal bristles. Arista distinctly pubescent,

the longest hairs as long as the thickened base is wide. 2:6—3:5 mm.

3: mid-femora covered with the normal sparse setulae up to apex,

femora and abdomen with the normal setae and setulae forvdi sp. n., p. 151

8(7). Five to seven pairs of inclinate frontal bristles (Fig. 63). Arista almost bare,

the longest hairs shorter than the basal diameter of the arista. 3:3-4

mm. g: mid-femora on apical two-fifths of anterior surface with an

elongate area, devoid of setulae but adorned with dense short brown

microscopic pile; femora (and abdomen) with numerous, long, fine,

erect, curved bristly hairs : é : . tomentigeva sp. N., p. 152

Coenosia aurifacies sp. n. 3 @

Length 3-0-3°5 mm.; of wings 2°8-3-5 mm.

3g. Head black, densely grey dusted, frons and face golden-grey dusted,

interfrontalia black in a strictly dorsal view, golden-grey when seen from in

front. Frons parallel, slightly wider than one eye, fronto-facial stripe very

slightly narrowed not quite to apex of second antennal joint, rather strongly

dilated in an almost straight line from there to lower margin of eyes, where it

is distinctly wider than at vertex; parafrontalia half as wide as interfrontalia,

with three inclinate setae (and one or two minute hairs). Inner vertical strong,

cuter vertical moderate, postverticals and ocellars moderately strong. Antennae

fuscous, the second joint whitish-grey dusted interiorly at apex, third joint

two and a half times as long as wide, reaching ventral sixth of eyes, the latter

not quite reaching level of mouth-margin, apex of antennae thus not much

more distant from base of arista than from mouth-margin; arista sparsely

very short-haired, the longest hairs hardly as long as the basal diameter of the

MUSCIDAE: B.—COENOSIINAE I47

arista. Parafacialia almost half as wide as the third antennal joint, jowls slightly

wider than it.

Thorax black, densely grey-dusted, dorsum with three faint, narrow vittae,

the median one reaching base of scutellum, the others subinterrupted, running

from first to last dc. acy fine and sparse, in two rows, I + 3 strong dc, two

strong 7a; scutellum with a few setulae and the normal two pairs of strong

setae, the lateral pair slightly shorter. Two rather strong propleurals and

prostigmatals, the lower prostigmatal curved downward. stf/ in a triangle,

the anterior side of which is much shorter than the two others, lower stl

considerably shorter, posterior one somewhat longer and stronger than the

anterior stPl.

Abdomen black, densely grey dusted, shorter than thorax, oblong-oval,

strongly compressed at apex; second to fourth segments with a narrow, dark

brown median vitta and second and third each with a pair of round, dark

brown spots; setae of second segment indistinct, third with a transverse row

of three pairs of moderate setae, fourth with a similar row of strong setae and

a marginal row of moderate setae, fifth with two pairs of setae, the interior

pair finer. Hypopygium rather strongly protruding, subglobular and densely

grey dusted in dorsal half, tapering, black and shining towards venter.

Legs entirely pale ferruginous, except for the coxae which are black and

densely grey dusted. Front femora with a pd row of about eight setae, the

four distal ones rather strong, and a pu row of 6-7 moderately fine though

somewhat longer setae, front tibiae with a rather short pv submedian seta;

mid-femora with two stronger a setae near middle and an a, pd and f preapical,

two pv setae are slightly longer than the femoral diameter, mid-tibiae with a _

pd slightly beyond middle and an ad slightly more apicad; hind femora with

seven rather strong though not very long ad, the last of them developed as a

preapical, a strong av on basal third and two in apical half, the other setae of

the av row small, a fd preapical, and a strong fu at second fifth, hind tibiae

with a strong ad in middle, without an av or fd seta.

Wings hyaline, veins testaceous, costa with a small costal spine, 74, and

m parallel, almost inappreciably diverging at extreme apex, vm at three-fifths

of discal cell. Calyptrae whitish-hyaline with pale yellowish border, the lower

one strongly protruding, parallel, broadly rounded at apex. Halteres pale

yellow.

2 similar to male, the pruinosity somewhat more yellowish, abdomen pointed

at apex, more depressed and broader, the second segment with distinct setae

near sides. Front femora with a blackish streak on posterior surface, mid and

hind femora with a not very conspicuous, brownish, incomplete ring on second

third. Hind tibiae with a rather strong though not very long av seta not much

beyond the ad.

KENYA: Aberdare Range, Nyeri Track, 10500 ft., x.1934 (F.W.E.) 3 type;

148 RUWENZORI EXPEDITION

Chyulu Hills, 5200 ft., v.1938 (Coryndon Memorial Museum, Nairobi), 9

paratype.

Coenosia heterocnemis sp. n. 3 2

Length 3:9-5°5 mm.; of wings 4-2-6 mm.

o. Head black, densely grey or bluish-grey dusted, interfrontalia black

with strong brown reflections when seen from in front, parafrontalia grey dusted,

with the interior margin brownish dusted, sometimes entirely brown dusted.

Frons very slightly narrowed to vertex, somewhat broader than one eye in

front, longer than wide; fronto-facial stripe distinctly though slightly narrowed

to beyond base of arista and rather strongly widened to lower margin of eyes,

where it is wider than frons; parafrontalia a quarter to a third (near frontal

triangle) as wide as interfrontalia, with three (and 1-2 minute) inclinate frontal

setae and a few minute hairs on outer anterior part. Inner vertical strong,

outer vertical moderate or weak, postverticals and ocellars rather strong.

Antennae black, not quite reaching lower margin of eyes, the latter not quite

reaching level of mouth-margin, second joint silver-dusted interiorly at apex,

third joint not quite thrice as long as wide; arista short-haired on dorsal surface,

the longest hairs slightly longer than the basal diameter of the arista, the

ventral hairs extremely short and sparse. Palpi black. Parafacialia a third as

wide, jowls slightly wider than third antennal joint.

Thorax black, grey or bluish-grey dusted, dorsum brownish-grey dusted

with five narrow brown vittae, beginning at frst dc, the median one often

obsolescent behind middle. acy fine and small, in two rows, some additional

hairs behind middle, 1-+3 strong dc, two moderate za; scutellum sparsely

setulose with the normal two pairs of strong setae, the lateral pair slightly

shorter. Two rather strong propleurals and prostigmatals, the lower pro-

stigmatal seta curved downward, not surrounded by hairs. s¢f/ in an equilateral

triangle which is almost or entirely bare, the lower and anterior stp/ subequal,

rather strong, the posterior one very strong.

Abdomen oblong-oval, as long as the thorax, distinctly depressed at base

and compressed at apex, densely grey dusted, with a more or less faint brown

median vitta and a pair of usually large, roundish to subquadrate spots on

segments 1-4. Lateral setae distinct on these segments though not very strong

on the third and especially second segments, fifth segment with a pair of discal

and two pairs of marginal setae.

Legs moderately short, pale ferruginous, the front coxae dusted, the mid

and hind coxae usually grey and dusted, the front femora sometimes greyish

and dusted towards base of dorsal surface, the mid and hind femora sometimes

infuscate near apex of dorsal surface, tarsi more or less infuscate and dusted.

Front femora with 4-5 strong fd setae (and I-2 weak ones near base) and

5-6 rather long pv setae which alternate with small ones, front tibiae with a

MUSCIDAE: B.—COENOSIINAE I49

moderate pv seta, mid-femora with 2—3 strong a setae near middle and second

third, 2-3 av near base, four pu near base and two (fd and fp) preapical setae,

mid-tibiae with a moderate fd seta somewhat beyond middle and an ad slightly

more distad; hind femora with about seven strong ad, the last one a preapical,

3-4 strong and several small av, including a strong preapical, one long fu seta

on second fifth, and a fd preapical, hind tibiae with a small fd and a rather

strong ad, usually level with each other and almost in middle.

Wings hyaline, very slightly smoky, veins brown, 7,,, and m very slightly

converging to very slightly diverging at apex, y—-m at three-fifths of discal cell.

Calyptrae hyaline with yellowish border, small, but the lower one distinctly

projecting beyond the upper one, not very narrowly rounded at apex, though

subtriangular. Halteres testaceous.

2 with the third antennal joint two and a half to three times as long as

wide, reaching lower eighth of eyes, jowls as wide as to one and a half times

as wide as the third antennal joint, fourth abdominal segment with a transverse

row of 2-3 pairs of strong discal setae, hind tibiae with a well developed av

seta somewhat beyond the ad.

Kenya: Aberdare Range, Mt. Kinangop, 10,000 ft., x.1934 (F.W.E.) 3 type,

8 § 11 2 paratypes; 8000 ft. (J. Ford) 1 2 paratype; gooo ft., in cedar forest,

27.X.1934 (f.W.E.) 4 § paratypes; 12,000 ft. 1 J 4 2 paratypes; 13,000 ft., on

Senecio brassicaefornis or aberdarica, 28.-30.x.1934 (J. Ford) 3 2 paratypes;

above Nakuru, ca. 9300 ft., on Lobelia aberdarica, 6.11.1935 (F.W.E.) 3 3 39

paratypes; Nyeri Track, 10,500-11,000 ft., 28.x.1934 (F.W.E.) 5 97 2 paratypes.

Mt. Elgon, alpine zone, 12,000-13,000 ft., 11.1935 (f.W.E.) 3 3 3 2 paratypes;

heath zone, 10,500-11,500 ft. 3 9 5 9 paratypes.

The infuscation of the dorsum of the femora varies in the same locality

though it seems to affect a larger proportion of the Elgon specimens. In one

of the males from Mt. Elgon the av seta of the hind tibiae is present (on both

sides). In Abyssinian specimens the lower calyptra is still considerably smaller

(see Emden, 1940, Ann. Mag. nat. Hist.).

Coenosia heterocnemis brunneigena ssp. n.

g. Structural characters as in typical C. heterocnemis, but face, parafacialia

and jowls brown-dusted, the vm vein runs to the middle of the discal cell,

and mid and hind femora with short, but strong apical infuscation; this infus-

cation occupies about the apical sixth and is not confined to the dorsal surface,

but reaches more or less the ventral surface, whereas in the typical form it is

much longer and narrower, and not so conspicuous. The jowls are distinctly

narrower than the third antennal joint.

UGANDA: Ruwenzori, Nyamgasani Valley, 8000-gooo ft. (D. R. Buxton),

3 type.

150 RUWENZORI EXPEDITION

Coenosia microcalyptra sp. n. 2

Length 3-4 mm.; of wings, 3-0 mm.

Head black, brown dusted, face, parafacialia and jowls brownish-grey

dusted, interfrontalia black, with strong brown reflections when seen from in

front. Frons parallel, considerably wider than one eye, very slightly narrowed

anteriorly not quite to apex of second antennal joint, from where the fronto-

facial stripe is evenly widened with almost straight sides to lower margin of

eyes, where it is a quarter wider than at vertex; parafrontalia a third as wide

as interfrontalia, with two large, two moderate and a few minute inclinate

setae and one pair of reclinate (on one side two pairs, the anterior one longer)

frontal setae. Inner vertical strong, outer vertical rather strong, postverticals

rather strong, ocellars very strong and long (slightly longer than the inner

verticals), strongly diverging. Antennae black, second joint grey dusted

interiorly at apex, third less than two and a half times as long as wide, reaching

approximately ventral eighth of eyes, the latter falling far short of level of

mouth-margin, apex of antennae almost as distant from mouth-margin as from

base of arista; the latter very short-haired throughout, the longest hairs of

ventral surface distinctly shorter, of dorsal surface slightly longer than basal

diameter of arista. Parafacialia more than half as wide as the third antennal

joint, jowls almost twice as wide as third antennal joint.

Thorax black, dark brown dusted, the pleurae greyish-brown dusted, dorsum

with three not very conspicuous, darker brown, narrow vittae and traces of

a lateral pair. acy sparse and fine, in two rows, I-+3 strong dc, two moderate 7a;

scutellum with a few single setulae and the normal two strong pairs which are

very long, as are the second postalar and the sa. One moderate and one rather

small propleural, upper prostigmatal rather strong, lower one fine, curved

downward. stf/ in an almost rectangular triangle, the posterior side longer,

the anterior one shorter than the upper one, area devoid of setulae, lower stl

somewhat weaker than the two others which are strong.

Abdomen black, rather depressed, suboval, densely greyish-brown dusted,

with a moderately narrow median vitta on segments 1-4 and a pair of rather

large, subquadrate dark brown spots on second and third segments and a small

one on first segment; lateral setae rather indistinct, except on fourth segment.

Legs wholly black, coxae densely, femora moderately dusted, tibiae and

tarsi slightly dusted. Front femora with a fd and pv row of six setae each,

the basal 2-3 weak, front tibiae with a moderate pu near middle, mid-femora

with four small a and one stronger a preapical seta, one strong av seta almost at

basal third, one long fv at second fifth and two preapicals (pd and p), mid-

tibiae with a rather strong pd and ad almost level with each other at middle;

hind femora with seven ad, the strongest ones towards apex, ending in an ad

preapical, a strong av at a third, another strong one at second third and a

MUSCIDAE: B.—COENOSIINAE I51

strong av preapical, a strong pu on second fifth, and a rather weak fd preapical,

hind tibiae with a rather small submedian fd (on one side with another very

small one basad of it), a strong ad level with the fd, and a fine av somewhat

beyond the other submedian setae. The tibiae with a small additional preapical

seta (besides the d and fd on the front, d on the mid, and d and ad preapicals

on the hind tibiae) to the other side of the d preapical (on the mid-tibiae on

the pd surface).

Wings rather strongly smoky, somewhat more so towards costa, somewhat

less towards hind margin, veins brown, 7,,; and m quite parallel for a con-

siderable distance at apex, 7m slightly beyond middle of discal cell. Calyptrae

very small, the lower one not projecting, even smaller than the upper one,

borders brown with brownish fringe. Halteres pale reddish.

Kenya: Mt. Elgon, alpine zone 12,000—-13,000 ft., 11.1935 (F7.W.E.) 2 type.

Coenosia fordi sp. n. 5 2

Length 2:6-3°5 mm.; of wings 2-7-3°7 mm.

3g. Head black, densely grey dusted, upper part of occiput and frons brown

dusted, interfrontalia black in a strictly dorsal view, brown in anterior view,

face and parafacialia yellowish-grey dusted. Frons subparallel, inappreciably

narrowed at extremities, longer than wide; fronto-facial stripe distinctly nar-

rowed to apex of second antennal joint, rather strongly widened to lower margin

of eyes, where it is wider than frons; parafrontalia a third as wide as interfron-

talia, with 3-4 inclinate frontal setae. Inner vertical strong, outer vertical

and postverticals rather strong, ocellars very long, longer than the inner

verticals, very strongly diverging. Antennae fuscous, almost reaching lower

margin of eyes, the latter not quite reaching level of mouth-margin, apex of

third antennal joint distant from mouth-margin by about width of antennae,

second joint silver-dusted interiorly at apex, third joint not quite thrice as

long as wide, with yellowish-grey reflections on inner surface of basal half;

arista short-haired, the longest hairs on dorsal surface as long as the diameter

of the thickened base. Palpi black. Jowls one and a half times as wide as the

third antennal joint.

Thorax black, dorsum densely dark brown dusted without vittae, pleurae

densely grey dusted. acy sparse but rather strong, in two rows, 1+-3 strong dc,

two strong 7a; scutellum with a few single setulae and the normal two pairs

of strong setae, the lateral pair slightly shorter. Propleural and upper pro-

stigmatal setae moderately strong, lower prostigmatal curved downward, not

surrounded by hairs. s¢f/ in an equilateral triangle which bears a few erect

hairs, lower stf/l moderately strong, the others strong and subequal.

Abdomen depressed, oblong-oval, uniformly dark brown dusted, only the

sides and base more greyish dusted, as is the venter. Lateral setae rather fine

152 RUWENZORI EXPEDITION

though distinct on segments I-3, somewhat stronger on the fourth, fifth segment

with two pairs of setae and some fine hairs, some of which are half as long as

the longer setae.

Legs rather short, wholly black, greyish dusted, only the knees very nar-

rowly yellowish. (Front legs of type missing, but no doubt as in female: front

femora with 6-7 moderately strong pd and fv, tibiae with a moderate pu beyond

middle), mid-femora with three rather weak a bristles on middle half, three

rather long av in basal half, three rather long and a few short pv, the long ones

on middle half, and two preapicals (fp and fd), mid-tibiae with a fine ad and

pd practically on the same level slightly beyond middle. Hind femora with

7-8 rather strong ad, 6-7 av and a pd preapical, hind tibiae with a rather strong

ad seta in middle and a rather fine av seta considerably beyond it.

Wings subhyaline, slightly smoky, strongly iridescent, veins brown, 74,5

very slightly converging to apex, 7-m just beyond second fifth of discal cell.

Calyptrae whitish-hyaline with pale-brown border, small, the lower one sub-

triangular, slightly projecting, rather narrowly rounded at apex. Halteres

yellow.

2 similar to male, the face greyish dusted, abdomen pointed, 7—m slightly

before or at middle of discal cell.

UGANDA: Kigezi District, summit of Mt. Sabinio, 11,000-11,500 ft., xi.1934

(J. Ford) 3 type; Mt. Mgahinga, 11,000 ft., 22.xi.1934 (F.W.E.) 3 2 paratypes;

Mt. Muhavura, 10,000-12,000 ft., xi.1934 (F.W.E.) I 2 paratype (headless).

Coenosia tomentigera sp. n. 5

Length 3°6-4:3 mm.; of wings 3-8—4:4 mm.

3. Head (Fig. 63) black, densely grey dusted, the interfrontalia dull black

in dorsal view, with strong brown reflections in anterior view, the median dorsal

part of occiput and the frons dark brown dusted. Frons subparallel, slightly

wider than one eye, longer than wide, fronto-facial stripe hardly narrowed to

apex of second antennal joint, rather strongly widened with almost straight

sides to lower margin of eyes, where it is a third wider than at frons; para-

frontalia a third as wide as interfrontalia, with 5~7 very fine though rather long

inclinate frontal setae. Inner vertical strong, outer vertical rather strong,

postverticals rather strong, ocellars very long and strong, slightly longer,

stronger and more curved than the inner verticals, very strongly diverging.

Antennae black, second joint grey dusted interiorly at apex, third less than

two and a half times as long as wide, slightly acuminate dorsally at apex,

with brown and whitish reflections, almost reaching level of ventral margin of

eyes, apex almost as far from mouth-margin as from base of arista, the latter

very short-haired throughout, the longest hairs on ventral surface much shorter,

MUSCIDAE: B.—COENOSIINAE 153

those on dorsal surface slightly shorter than basal diameter of arista. Para-

facialia not quite half as wide as the third antennal joint, jowls one and a half

times as wide as it.

Thorax black, dark grey dusted, with three narrow, faint, brown vittae,

pleurae densely grey dusted. acy fine and sparse, though rather long, in two

rows, I-+3 very long dc, two rather long za; scutellum with a few setulae and

the two normal pairs of setae. One moderate propleural seta, two rather strong

63.

Fic. 63.—Head of Coenosia tomentigera sp. n., front view.

prostigmatal setae, the lower one curved downward. s¢f/ long, in an equilateral

triangle, which bears a few long hairs, lower stfl somewhat shorter, posterior

one somewhat longer and stronger than the anterior one.

Abdomen rather depressed, oblong-oval, dark brown dusted on dorsum,

sides, base and venter dark grey dusted, with rather numerous fine and long

hairs on sides and venter, the lateral setae not much stronger, hairs on dorsum

rather short and appressed, fifth segment with a pair of discal and two pairs

of marginal fine setae, and with a few rather long hairs.

Legs wholly black, with rather numerous fine and very long setae, front

femora with about eight pd and fv setae, the latter very long, all the posterior

154 RUWENZORI EXPEDITION

surface clothed with similar long fine setae, among which a # row of five

especially long ones in basal half is striking, front tibiae with a fine, moderately

long p seta just beyond middle; mid-femora with four long a and very long av

setae in basal half, about eight very long and fine fv setae and four preapical

setae on pd and # surfaces, but without ad setae, the apical two-fifths of the

anterior surface without hairs, but covered with dense, short, dark brown

tomentum, mid-tibiae with rather long but very fine submedian fd and ad,

the former just beyond middle, the latter slightly farther on; hind femora

with about ten long and fine ad and av setae, the anterior surface beset with

rather numerous similar setae, hind tibiae with a long ad seta beyond middle

and a shorter av almost at second third.

Wings hyaline, very slightly smoky, veins brown, 7,,; and m parallel or

slightly diverging at apex, v-m at or slightly beyond middle of discal cell, alula

with a fringe of about a dozen very long and fine hairs. Calyptrae whitish-

hyaline with yellowish border. Halteres yellow.

2 similar to male, the margin of the eyes below the antennae more rounded,

all the setae somewhat shorter and stronger, the hairs on the # surface of the

front and the a surface of the hind femora much shorter than in male, very

distinct from the setae, though longer than usual, mid-femora without the

patch of tomentum; abdomen pointed, without the long hairs at sides and

venter, the lateral setae not very strong, somewhat longer on third segment

and rather strong on fourth, on which two lateral discal pairs are conspicuous.

UGANDA: Ruwenzori, Nyamgasani Valley, 12,000—13,000 ft. (D. R. Buxton)

6 type, 2 3 3 Y paratypes.

Caricea R.—-D.

The names Coenosia Meig. 1826 and Caricea R.—D. 1830 are here used in

the same sense as by Stein, Malloch, and others, and the type-fixations of

Rondani are accepted. Huckett in his excellent revision of the North American

Coenosta (1934, Trans. Amer. Ent. Soc. 60:57—119, 133-198) uses Coenosza for

Caricea auct. and Limosia for Coenosia auct., accepting an earlier type-fixation

by Westwood (1840, Introd. Classif. Ins. II, Synops. Gen. p. 143). This change

is, however, not justified in my opinion. For the species with long-haired

arista had in 1830 already been united in a separate genus Caricea by Robineau-

Desvoidy, so that t7gvina F., to which one of the species described by Robineau-

Desvoidy as Caricea had been synonymized as early as 1835 by Macquart

(Hist. nat. Ins. Dipt. 2 : 344) should not have been used in 1840 as genotype

for the old genus Coenosia. Huckett discusses this fact, but remarks correctly

that this recommendation contained in the present rules of nomenclature

would hardly invalidate Westwood’s fixation. On the other hand, it is proved

MUSCIDAE: B.—COENOSIINAE 155

by Huckett and evident from the descriptions that Meigen’s é/gvina is identical

with tigrina F. This implies, however, that Westwood’s genotype of Coenosia

is not the true tigrina F., for in making tigrina F. the genotype of Coenosia,

Westwood mentions expressly that “C. tigrina Meig. belongs to the s.g. Caricea

R.D.” As Caricea was then distinguished from: Coenosia by the long-haired

arista, this means that to Westwood the species of Meigen and Fabricius were

quite different, and belonged respectively to Coenosia and Caricea, i.e. that

tigrina Meig., sensu Westwood was a species with long aristal hairs such as

was later designated as type of Caricea by Rondani (1856), whereas tigrina F.

sensu Westwood must be a species with short hairs of the arista. It is impossible

Fias. 64-67. Left hind tibiae, anterior view. 64.—Coenosia hetevocnemis sp. n.

65.—Caricea transveysalis sp. n. 66.—Caricea calopoda Bezzi. 67.—Caricea

aequivitta sp. n.

to find out which species the misidentified t7gvina F., Westwood really was

(though it must have been one of the other larger species, which belong all to

Coenosia sensu Stein, probably intermedia Fall.), and Westwood’s type-fixation

can, therefore, not be accepted, so that Rondani’s (1866) fixation of genzculata

Fall. is valid as genotype of Coenosia (his former—1856—fixation of sexmaculata

Meig., being inacceptable, as the species was not contained in Meigen’s paper

of 1826).

Robineau-Desvoidy, Meigen, Macquart, Rondani, Meade and others dis-

tinguished Caricea from Coenosia by the longer hairs of the arista, but Stein

(x898, Berlin. Ent. Zschr. 42: 257, note) regarded as Caricea those Coenosia

which were characterized by a laterally compressed head, appreciably shortened

antennae, and strikingly long setae of the tibiae. He mentioned the European

II, 4 (e)

156 RUWENZORI EXPEDITION

species tigrina Mg., nana Zett. (=humuilis Meig.), and cingulipes Zett. (Stein,

non Zett.,—strigipes Stein) as belonging to Caricea, to which list I am for the

time being only able to add atra Meig. Later on Stein considered Caricea a

synonym of Coenosia, and so did most of the other authors. In 1922 (Ann.

Mag. nat. Hist. (9) 10: 576) Malloch separated the African Caricea from Coenosia

by the two very long bristles at middle of hind tibiae, “‘the bases of which are

very close together, one on the antero-dorsal and the other on the anterior or

antero-ventral surface”, whereas in Coenosza the hind tibiae have “one long

strong bristle on antero-dorsal and a weaker bristle on the antero-ventral

surface, the latter nearer apex than the former’’. The large material, which

I have worked out, suggests slight alterations in these definitions, as the bases

of the setae on the anterior surface of the hind tibiae in Caricea may sometimes

be as widely separated as in Coenosia (e.g. in Caricea calopoda Bezzi and

aequivitta m., Figs. 66 and 67, closely related to species in which the setae are

close together), and as in Caricea the seta is always inserted on the anterior

and never on the antero-ventral surface, the decisive character for the dis-

crimination between Caricea and Coenosia thus being the anterior position in

the former genus and antero-ventral in the latter of that seta which is usually

but not always very close to the antero-dorsal seta of the hind tibiae in Carzcea.

I was glad to find at an advanced stage of this paper that Malloch (1921,

Ent. News 32: 107) and Huckett in their studies on the North American

Coenosiinae have come to the same definition, except for some differences in

the rank of the different units and, as far as Huckett is concerned, in the

names used.

On species without an av or a seta and with a fd seta on the hind tibiae

Malloch founded the genera Xenocoenosia and Neodextopsis respectively. I do

not know the American genotypes of these groups, but as far as African species

with the same characters are concerned, the females of those males which lack

the a or av seta show the characters of Coenosia or Caricea quite unequivocally,

and the fd occurs in both genera and is in my opinion not a generic character.

I do not think it right either to erect separate genera on those species with

only two fost de or one pair of scutellar setae, which are in all other regards

typical Caricea.

Caricea is by far the largest genus of Coenosiinae in the Ethiopian region,

and the following key contains, therefore, the groups of species and the isolated

species, whereas keys to the single groups are being presented at the beginning

of each group. The submedian fv seta of the front tibiae is present in all

species, unless otherwise stated.

The following species of “Coenosia”’ described from the Ethiopian region,

which may be expected to belong largely or all to Cavicea, are unknown to me

and could not be incorporated in the key (references given, as far as not con-

tained in Séguy, 1937, Gen. Ins. fasc. 205).

MUSCIDAE: B.—-COENOSIINAE 10S

africa Curr. (Xenocoenosia) N.W. Tanganyika (1935, Amer. Mus. Novit.

776: 12), see note, p. 145.

albipila Stein. S. America or Africa (hardly East African).

(cyclophthalma Thoms. not a Coenosiine, as four post de are present).

gigas Beck. Kenya. calopoda group?, apparently similar to atroapicata.

grauert Curr. Tanganyika. calopoda group? (1935, Amer. Mus. Novit.

776: 8).

inversa Wied. Cape. calopoda group?

laxifrons Curr. Pretoria. = fascigera Stein? (1935, Amer. Mus. Novit.

779: 4).

morops Ség. (Hoplogaster). Kenya. vittata group near vittata? (1938,

Miss. scient. Omo, 4, Zool.: 367).

trichopyga Loew. Mozambique.

Key To THE GROUPS AND ISOLATED SPECIES OF ETHIOPIAN CARICEA

1(8). Hind tibiae with a distinct submedian pd seta (Fig. 65).

2(3). 1-+2dc. Postsutural part of mesonotum with a broad, dark-brown dusted,

transverse band touching suture along front margin. Calyptrae rather

small, lower one narrowly rounded. Hind tibiae with the pd basad of

the anterior pair of bristles and with three preapicals. Legs almost

entirely black. . : : : é s tvansversalis sp. N., p. 160

3(2). 1+3 de.

4(5). Postsutural part of mesonotum with a broad, dark transverse band, touching

suture along front margin. Third antennal joint very long and broad,

almost reaching mouth-margin, arista almost bare. Legs yellow,

dorsal surface of all the femora infuscate, joints 2-5 of the front tarsi

(3) dilated and soled, joints 2-4 with the basal half deep black and the

apical half white, fifth joint almost entirely black with the pulvilli

white. pd seta of hind tibiae well basad of a and ad.

[poecilotaysis sp. n.], p. 161

5(4). Pattern of thorax longitudinal, or absent. Front tarsi not so spotted.

6(7). Scutellum with the lateral pair of setae only, which are very strong; the

apical pair absent. Lower prostigmatal bristle absent. Jowls almost

absent, not wider than the base of the palpi. Front tarsi very long, the

pd of the hind tibiae much beyond the other setae, the a slightly beyond

the ad. Thorax with five narrow brown vittae. Legs yellow with tarsi

and apex of posterior femora piceous. . . scutellavis sp. n., p. 162

7(6). Scutellum with four strong setae, or two strong apical and two weak lateral

bristles. Lower prostigmatal bristle present. ; vittata group, p. 164

8(1). Hind tibiae without a pd seta (Figs. 66, 67).

9(ro). Scutellum with the /ateval pair of setae only, which are very strong, the

apical pair absent. Jowls well developed, as wide as the third antennal

joint. Prostigmatal bristles well developed, surrounded by a few small

hairs. Thorax with five brown vittae. Legs yellow with the tarsi and

the apex of the femora piceous. . 5 : . . bipila sp. n., p. 175

13(16).

14(I5)-

15(14).

16(13).

17(18).

et Ne

18(17).

RUWENZORI EXPEDITION

Scutellum with four strong setae, or the apical pair only strongly developed.

. Legs more or less yellow, at least one pair of femora more or less broadly

yellowish translucent at base, or one pair of tibiae yellowish in middle

or at tip. Halteres yellow.

. The narrowest point of the fronto-facial stripe is almost at or above the base

of the antennae; if it is below the base, then the stripe only slightly

narrower there than at the antennal base, and the eyes distinctly more

widely separated at ventral apex than at dorsal apex (Figs. 66, 67, 74-76).

Antennae inserted between middle and second fifth of eyes (except in

mgahingana, where the antennae and face are rather long), face not

much longer than frons. Jowls often as wide or wider than the third

antennal joint.

All the femora entirely black, except sometimes at apex. Tibiae never

entirely black.

Frons half as wide as one eye, seldom much wider, but then the size not

over 4 mm. (7nanis) or the parafacialia very narrow (tvichocnema). The

parafrontalia and frontal triangle always pale dusted, the interfrontalia

smooth (Fig. 74). : ; E : : 3 humilis group,

Frons almost or fully as wide as one eye. Six or more pairs of inclinate frontal

bristles, of which three pairs are long and decussate. Large species of

5-8 mm. length. Parafacialia never much narrower than the third

antennal joint. The parafrontalia and frontal triangle usually velvety

dark dusted as are the interfrontalia, the latter usually longitudinally

rugose in front. Claws and pulvilli very large, not appreciably shorter,

p. 176

or even longer than the third tarsal joint. . : tigyina group, p. 180

The mid and hind femora at least yellow at base.

Arista with very long hairs, the longest about as long as the third antennal

joint (Fig. 71). Palpi pale yellow, even at apex. Vibrissae black.

Hind femora with two preapical pd not far behind each other. The @

and ad setae of the hind tibiae level with each other. Third antennal

joint fuscous, grey dusted, often somewhat translucent towards apex.

Prostigmatal bristles surrounded by a few small hairs. Thorax with

or without three faint narrow longitudinal vittae, abdomen with a

faint median vitta on segments two to four, and a distinct pair of spots

on segments two and three or two to four. Legs (except mid, hind,

and part of front coxae) entirely yellow, with a small brown point on

the mid and hind femora at articulation with tibiae, and with at least

the last joint of tarsi strongly infuscate; front femora sometimes with

a dark shadow near base along exterior surface. . pachypoda Big.,

Arista at most short-plumose, the longest hairs never much longer than the

third antennal joint wide. Palpi brown with piceous apex, very seldom

yellow, but then the third antennal joint is pale yellow. Hind femora

normally with one preapical pd bristle.

. Small species of 2-5-4 (seldom up to about 5) mm. length. Hind coxae with

one outstanding (and one or two small) hairs on outer hind margin.

pu seta of front tibiae present. . c : stvigipes group,

. Larger species of (4°5—) 5-9 mm. length. Hind coxae with two or three out-

standing (and one or several small) setulae on outer hind margin,

except if no pv on front tibiae and only two scutellar bristles are present.

. Front tibiae without a pu seta. Scutellum with the apical setae only. Hind

p. 186

MUSCIDAE: B.—COENOSIINAE

coxae with one outstanding bristle on outer hind margin, though a small

but rather stout one is close to it. m,-+-cu, strongly thickened and

fuscous near base (3). Legs, including coxae, entirely yellow, but tarsi

strongly infuscate, femora and tarsi very long and slender.

aseta sp. N., Pp.

22(21). Front tibiae with a pv bristle. Scutellum with four setae. Hind coxae with

two outstanding setae and a few smaller ones.

23(24). Tibiae yellow, sometimes more or less infuscate. First post ia, if present,

level with sa. Parafrontalia pale dusted : : calopoda group, p.

24(23). Tibiae entirely black. First post ia before level of sa. Parafrontalia and

interfrontalia equally dark dusted. Mesonotum entirely dark dusted,

except on the sides along humeri and notopleurae.

25(26). Third antennal joint short, about half as long as the face, antennae inserted

at second fifth. Anterior 7a closer to level of sa than of first post dc.

a bristle of hind tibiae absent in male (always?), pu of front tibiae

strong. Face dark golden dusted. : : . nigritibia sp. n., p.

26(25). Third antennal joint long and broad, almost reaching mouth margin,

antennae inserted at dorsal quarter of eyes. Anterior post ia practically

level with first post dc. a bristle of hind tibia distinct, pv of front

tibiae weak in male. [ace silver dusted. 3 mgahingana sp. N., p. 2

27(12). The narrowest point of the fronto-facial stripe is well below the base of the

antennae, the stripe there not much more than half as wide as at

vertex, distinctly narrower there than at the base of the antennae

(Fig. 77). The eyes as distant at dorsal extremity as at ventral ex-

tremity, frons widened to vertex. Antennae inserted at dorsal third of

eyes, which are very large and elongate, face very narrow, much longer

than frons. Parafacialia very narrow, much narrower than the third

antennal joint. Jowls always much narrower than the third antennal

joint, the latter very slender, usually about four times as long as wide

or even longer. 4 é . vebmanni group, Pp.

28(11). Legs entirely black, except pometimes for the coxae and trochanters and

the base of the tibiae.

29(30). Third antennal joint very long and slender, more than four times as long as

wide, pointed at apex of dorsal side, antennae inserted at dorsal third

of eyes. Halteres black. Fronto-facial stripe distinctly narrower

beneath base of antennae than at their base (Fig. 78). Submedian

setae of tibiae extremely long, some of them about three-quarters as

long as the tibiae.t_ Anterior post ia rather long though fine. Anterior

stpl much shorter than the two others. Parafacialia and jowls ex-

tremely narrow. Abdomen of male rather club-shaped and strongly

curved downward. Frons slightly dilated to vertex. moyvio Sp. 0., p.

30(29). Third antennal joint about two to three times as long as wide, inserted at

about middle of eyes, or halteres yellow.

31(32). Frons distinctly dilated from first frontal setae to vertex, often strongly

dilated, uniformly dusted, the parafrontalia mot differentiated by paler

pruinosity, or the frons entirely silver-white dusted (Fig. 79).

niveifvons group, Pp.

32(31). Frons subparallel to vertex or at least subparallel in its middle part, or

1 These setae not more than three-fifths as long as the tibiae, Halteres yellow.

vebmanni group.

159

104

196

See

160 RUWENZORI EXPEDITION

slightly narrowed to vertex, seldom slightly dilated to vertex, the

parafrontalia distinctly differentiated by paler pruinosity though

narrow, frons never entirely white-dusted (Fig. 80). semifumosa group, p. 234

Caricea transversalis sp. n. 3d 2

Length 4:3-4°7 mm.; of wings 3°9-4:2 mm.

g. Head black, greyish pollinose, except on dorsal central part of occiput

which is glossy blackish, interfrontalia which are dull black with strong light-

brown reflections when seen from in front, parafrontalia and face which are

yellowish-grey dusted. Frons subparallel, very slightly narrowed at hind end

of eyes, as wide in front as one eye; fronto-facial stripe with the narrowest

point just beneath the base of antennae, gradually widening to the lower end

of the eyes, where it is hardly wider than in middle of frons; parafrontalia

one-quarter as wide as interfrontalia, with two strong and one or two small

inclinate frontal setae. Inner vertical strong, outer vertical weak, postverticals

and ocellars moderate. Antennae fuscous, the second joint with a small spot

of whitish dust interiorly at apex, third joint more than thrice as long as wide,

not reaching lower margin of eyes, arista short-haired on more than basal half,

the plumosity more than half as wide at base as the third antennal joint.

Palpi fuscous. Parafacialia half, jowls one and a half times as wide as third

antennal joint.

Thorax black, grey dusted, with a brown dusted, dark pattern, consisting

of three spots in front of suture, the lateral ones between dc, ph and prst setae,

and the median one from front end to suture or at least to beyond the frst dc,

and a broad transverse band along posterior side of suture; this band reaches

the infra-alar bullae and is usually connected on median line with the brown-

dusted dorsal surface of scutellum. acy minute and sparse, in two complete

rows, no prsc, I+2 strong dc, one small za (behind level of sa), scutellum

sparsely setulose, with two pairs of strong, equal setae. Two propleurals, the

lower one not much weaker, two prostigmatal setae, the upper one moderate,

the lower one small, directed downward. s¢f/ in a triangle, the anterior (lower)

side of which is distinctly shorter than the posterior and still more than the

upper side, devoid of setulae, the lower and anterior stf/ equal, considerably

shorter than the posterior one.

Abdomen black, grey dusted, with a complete, narrow median vitta and a

pair of roundish spots on segments I-4; these spots occupy the posterior part

of the segments and reach the hind margin of at least the second and third

segments, being at least narrowly connected with the median vitta on one or

several segments; moderately slender, moderately convex on dorsum, slightly

compressed at venter; second segment with one pair of lateral setae, third and

fourth each with one pair of lateral and one pair of dorsal setae towards apex;

fifth segment black with sparse dust, subglobular.

MUSCIDAE: B.—COENOSIINAE 161

Legs black, coxae and femora grey dusted, tibiae reddish-piceous to black,

the base reddish; front femora with approximately eight fine short av setae

in basal half and 6-7 rather long pv, front tibiae with a moderate submedian

pv, mid-femora with an isolated strong submedian av and two preapical (f and

pd) setae; mid-tibiae with rather strong ad and fd setae, the former more

apicad, hind femora with five rather strong ad setae and a fd preapical seta,

about five av setae and one rather long though not very strong pv seta on second

fifth, hind tibiae with one strong fd on second fifth, one ad just before middle

and one a on middle.

Wings hyaline, slightly smoky, veins brown, 7,,; and m slightly converging

owing to the curvation of the former, vm slightly beyond middle of discal

cell. Calyptrae whitish, rather small, but the lower one considerably projecting.

Halteres pale yellow.

® differing from male only by the more depressed and pointed abdomen.

UGaNnDA: Kigezi District, Mt. Sabinio, 8000 ft., xi.1934 (F.W.E.) & type,

I 9 paratype; Kanaba, 7800 ft., xi.1934 (F.W.E.) 2 9 paratypes; Lake Mutanda,

- 6000 ft., xi.1934 (J. Ford) 1 2 paratype. In two of the last-named specimens

the median presutural spot is broader, fused with the lateral spots and not

or only very narrowly connected with the transverse band, the latter not

reaching scutellum.

[Caricea poecilotarsis sp. n. 3]

Length 5 mm.; of wings 4:8 mm.

Head black, whitish-grey pollinose, except on median dorsal part of occiput,

which is glossy blackish, and interfrontalia which are dull, almost velvety

black with light brown reflections when seen from in front, their posterior

corners and outer margin behind middle grey pollinose. Frons slightly though

distinctly narrowed to vertex with straight sides, as wide in front as one eye,

fronto-facial stripe widest at base of antennae, narrowed to middle of face,

where it is as wide as at vertex, and widened again to epistoma; parafrontalia

a third to half as wide as the interfrontalia, with two moderate and two small

inclinate frontal setae. Inner vertical strong, outer vertical weak, postverticals

and ocellars moderate. Antennae fuscous, the second joint grey dusted

interiorly at apex, third joint four times as long as wide, slightly surpassing

lower margin of eyes, arista sparsely haired dorsally in middle half, the longest

hairs hardly as long as the base of arista wide. Parafacialia less than a third

as wide below middle as third antennal joint, jowls as wide as third antennal

joint.

Thorax black, whitish-grey dusted, with a brown dusted dark pattern,

consisting of a pair of oblique, posteriorly diverging spots from neck to just

beyond ph and prst dc, a brown spot around the anterior spiracle, a transverse

band, connecting the bases of the wings along the suture, and another transverse

162 RUWENZORI EXPEDITION

band across the scutellum, leaving the apical third of the latter pale but

encroaching narrowly upon the posterior part of mesonotum. acy minute and

sparse, no prsc, 1+3 strong dc, one moderate za (behind level of sa) ; scutellum

with the normal two pairs of strong setae. Two propleurals, the lower one

much weaker, two prostigmatals, the upper one moderate, the lower one much

weaker, curved downward. s¢f/ in a triangle, the anterior side of which is

shorter than the posterior and still more the upper side, devoid of setulae, lower

and anterior s¢tpl equal, considerably shorter than the posterior one.

Abdomen black, whitish-grey dusted, the second to fourth segments with a

complete, dark, brown-dusted median vitta, the hind margin of the first to

third segments each with a pair of transverse square to semicircular spots,

which are narrowly connected with the median vitta on margin, on the fourth

segment which is very short the apical half is occupied by a dark transverse

band. Second segment with a pair of weak lateral setae, third and fourth each

with a pair of strong lateral and paramedian marginals, fifth segment with a

pair of strong lateral marginals.

Legs testaceous, the dorsal surface of the femora broadly and strongly

infuscate (on the hind femora practically only the ventral part of the base

remains yellowish), hind tibiae largely infuscate except for base, second to fifth

joints of front tarsi dilated and soled, the second and third joints twice as long

as wide, the last two shorter, joints 2—4 with the basal half deep black and the

apical half whitish, fifth jot black, except for a very narrow whitish basal

part; mid and hind tarsi yellow, narrowly blackish at base of second to fourth

joints, apical joint black, pulvilli whitish. Front tibiae with a rather small fv,

mid-femora with two (p and fd) preapicals, mid-tibiae with a rather weak pd

beyond middle and a strong ad at three-fifths, hind femora with five rather

strong ad setae, a pd preapical seta, three av and one long and fine fv seta,

the latter on second fifth, the ventral surface of base with numerous fine, long,

erect hairs, hind tibiae with a strong fd slightly basad of middle and a rather

strong ad and a slightly beyond middle, the a slightly more apicad.

Wings hyaline, slender, veins testaceous, 7,,, and m parallel at apex, 7-m

beyond middle of discal cell. Calyptrae white, the lower one moderately

projecting. Halteres pale yellow.

2 unknown, probably with simple tarsi.

Kenya: Ngong, v.1936 (Miss Steele), § type in B.M. (sent in by Coryndon

Memorial Museum). The thorax of the specimen is badly broken, but all the

more important characters are clearly visible.

Caricea scutellaris sp. n. ¢

Length 6 mm.; of wings 7-2 mm.

Head black, whitish-grey pollinose, the interfrontalia dull black with

MUSCIDAE: B.—COENOSIINAE 163

brownish-white reflections, when seen from in front; frontal triangle yellowish-

dusted anteriorly, very narrow, reaching middle of frons and continuing as a

fine whitish line almost to lunula in anterior view. Frons parallel, narrow,

one-fifth as wide as head, two-fifths as wide as one eye; fronto-facial stripe

narrowest at base of arista, where it is a third as wide as one eye; parafrontalia

a quarter the width of the interfrontalia, with two strong and one small inclinate

frontal setae. Inner vertical very strong, outer vertical weak, postverticals

moderate, ocellars rather weak. Antennae dark fuscous, second joint narrowly

orange at apex, with a small spot of whitish dust interiorly at apex; third joint

almost thrice as long as wide, just reaching lower third of eyes, distance between

its apex and mouth-margin equal to its length; arista bare except near base,

where it is short-haired, the longest hairs being slightly longer than the basal

diameter of arista. Palpi fuscous. Parafacialia linear, jowls not wider than

the base of the palpi.

Thorax blackish, somewhat pale above front coxae, whitish-grey dusted,

the mesonotum brownish-grey dusted with five narrow longitudinal vittae, of

which the median one is complete and extends to beyond middle of scutellum,

whereas the vittae in the dc stripe are incomplete in front and behind and

interrupted at suture and the ones along 7a are only vestigial. acy sparse and

fine, in two irregular rows, no prsc, I + 3 strong dc, two za, the anterior one

very small and level with sa, scutellum almost bare and with only the lateral

pair of setae, which are very long and strong. Two very fine propleurals, lower

prostigmatal absent, upper one fine and not very long, though almost twice as

long as the upper propleural. s¢// in an isosceles triangle, the upper side

slightly longer than the others, a few rather long hairs within the triangle, the

anterior spl weak, the two others very long and strong, especially the posterior

one.

Abdomen slender and parallel in dorsal view, somewhat clavate at apex

in lateral view, fuscous with narrow hind margins and the base and ventral

surface of the first segment testaceous, translucent, hypopygium and fifth

segment ferruginous; setulose hairs rather long, but setae well developed

only at sides of first segment and as a sublateral pair on fourth segment.

Legs, including coxae, yellow, apical quarter to fifth of hind femora and all

the tarsi piceous (mid-tarsi missing). Legs very long and slender, especially

hind femora and front tarsi; front femora with four very long pv setae which

alternate with shorter ones, front tibiae with a long pv seta, mid-femora with

two long av, four long pv and two (pf and fd) preapical setae, mid-tibiae with

a very long and strong ad at two thirds and a somewhat smaller fd beyond

middle, hind femora with four moderate av, three very long pv and one pd

preapical setae, hind tibiae with a long and strong fd beyond middle, a still

stronger ad slightly basad of middle, and a considerably shorter but still very

strong @ seta just before middle.

164 RUWENZORI EXPEDITION

Wings slightly infuscate, slender, veins brown, 7,,; and m hardly appreciably

converging, almost parallel at apex, v-m in middle of discal cell. Calyptrae

hyaline with the border yellowish, of normal size, the lower one widely projecting.

Halteres whitish, rather large.

Ucanpa: Kigezi District, Mt. Sabinio g000-10,000 ft., 25.xi.1934 (F.W.E.)

3 type.

Caricea vittata group

1(16). pd seta of hind tibiae basad of ad or level with it. Thorax with three broad!

brown vittae, which are seldom faint, and which are separated by two

more or less narrow grey vittae along dc. Abdomen largely grey dusted,

usually with a median vitta and pairs of spots.

2(15). The pd seta of the hind tibiae basad of the ad. Both pairs of scutellar bristles

long and strong, seldom the lateral pair much smaller, but then the

median vitta of thorax extended over almost all the dorsal surface of

scutellum.

3(44). Legs more or less yellow. Lower calyptra of at least moderate size, always

distinctly projecting beyond upper one.

4(11). Ocellar bristles well developed, about two-thirds as long as frons. a bristle

of hind tibiae present in male (tarsalis?, fuscifemur?) and female.

Upper prostigmatal bristle not appreciably shorter than propleural

bristle. Arista short-haired, the fringe of the dorsal side broader than

the thickened base of the arista.

5(8). Palpi and front coxae black. The median thoracic vitta not reaching

scutellum. Jowls less than twice as wide as the third antennal joint.

6(7). Legs long and thin, with the setae long and strong in female; some of them

long and thin, the others long and strong in male; tibiae one and a half

times as long as the head wide. Abdomen long and slender. Anterior

ta hardly one-half as long as the anterior sa, not longer than the pos-

terior ia. Frons about three-quarters as wide as one eye. aand ad of

hind tibiae rather far beyond pd. . ‘ . gvacilipes sp. D., p. 165

7(6). Legs short, front tibiae as long as the head wide, mane tibiae slightly longer,

tibiae entirely yellow, tarsi piceous, front femora blackish, or with a

blackish streak on pd surface. Abdomen short and stout. Anterior ia

unusually strong,” stronger than the posterior 7a and more than half as

long as the anterior sa. Frons rather broad, one-sixth narrower than

one eye, slightly narrowed to vertex, with six inclinate frontal setae, the

first and last of which are long, and the second and fourth of which are

moderately long. a seta of hind tibiae slightly beyond ad, ad not much

beyond pd. Jowls one and a half times as wide as the third antennal

joint. Vittae of thorax faint. Abdomen with a very narrow median

vitta and a pair of small indistinct spots on the second and third seg-

ments:>! 9.4): - 5 [tavsalis Walk. ]

8(5). Palpi and front coxae geliow. The medina tonnes vitta extending over

disc of scutellum. Jowls twice as wide as third antennal joint.

! Except in taysalis, where three or five narrow vittae are present.

2 But as in gracilipes in a second female, recently discovered. In it the front femora

have only a blackish streak, and only three inclinate frontal setae and three very small .

hairs are developed.

MUSCIDAE: B.—COENOSIINAE 105

g(r10). Frons parallel, three-quarters as wide as one eye. Femora and tarsi strongly

infuscate, apical half of front tibiae slightly infuscate. Dark spots of

abdomen rather large. The insertions of the three setae of the hind

tibiae can be connected in projection by a straight line. 9. — [ fuscifemur Mall.|

10(9). rons conspicuously narrowed to vertex, slightly wider in front than one

eye. Legs entirely yellow, except for the apical half or two-thirds of

the last tarsal joint, which is blackish, whereas the basal part is pale

yellow. Dark spots and median vitta of abdomen of moderate size.

pd of hind tibiae much basad of a straight line running through the a

and ad setae. ¢. a j . distitaysalis sp. n., p. 167

11(4). Ocellar bristles small, scarcely a third as lone as frons. a bristle of hind

tibiae absent in male. Upper prostigmatal seta weak, about one-half

or two-thirds as long as the propleural seta. Frons distinctly narrowed

to vertex, interfrontalia strongly narrowed to the reclinate seta. Arista

almost bare, the short dorsal pubescence hardly as broad as the thick-

ened base. Lower Sere rather short though well projecting beyond

the upper one. : é [vittata Wied. 12]

12(13). Legs entirely yellow, the apex ot the DOC reriOn femora often narrowly in-

fuscate. Palpi yellow to brown. Pattern of thorax and abdomen light

brown. Face (g) dull golden dusted. : : [vittata vittata Wied.]

13(12). Legs with the tarsi piceous, the tibiae and apex of femora strongly infuscate

to piceous. Palpipiceous. Pattern of thorax and abdomen dark brown.

Face (3) velvety black. 0 : : [vittata chyuluana n. ssp.], p. 169

14(3). Legs entirely piceous-black. Lower calyptra extremely short, shorter than

the upper one and not projecting beyond it. Ocellar bristles very

small, not a third as long as frons. Lateral bristles of scutellum con-

spicuously shorter than apical ones. a seta of hind tibiae present in

both sexes. Upper prostigmatal as long as propleural. Median vitta

of thorax complete to more or less apex of scutellum. [someveni sp. n.], p. 169

15(2). The fd seta of the hind tibiae level with ad, tibiae piceous to black, except

at base. The lateral scutellar bristles much shorter than the apical

ones. The median thoracic vitta not reaching scutellum. Palpi and

front coxae black. Upper prostigmatal bristle long. Lower calyptra

twice as much projecting as the upper one. Front femora black,

posterior femora orange and black. . , . edwardsi sp. n., p. 171

16(1). pd seta of hind tibiae rather far beyond the ad. Thorax with five brown

vittae. Abdomen almost entirely dark brown dusted,! rather shining,

front and hind margins greyish dusted. Lower calyptra projecting by

twice the length of the upper one. Legs very long and slender, all the

femora largely yellow. : : : : VUWENZOVICA SP. D., Pp. 173

Caricea gracilipes sp. n. 5 2

Length 5-8-6:5 mm.; of wings 6.I-7 mm.

g. Head black, fence grey dusted, except on interfrontalia, which are

1 Ground-colour of body reddish-yellow, except on head, mesonotum and scutellum

which are dark; abdomen with brown transverse bands which reach posterior and—except

on first segment—anterior margins in middle. Basal segments of antennae brownish-

yellow, arista very short-plumose: see (Neodexiopsis) colovata Curr. (1938, Amer. Mus.

Novit. 974: 1), which according to information obtained from Dr. Curran traces to Caricea

yuwenzorvica in the above key but differs in colour of abdomen etc.

166 RUWENZORI EXPEDITION

black with brown reflections, and face which is silver-white. Frons practically

parallel, much longer than wide, almost inappreciably narrowed to base of

antennae and to vertex, two thirds as wide in front as one eye, fronto-facial

stripe hardly narrowed to apex of second antennal joint, gradually widened to

lower margin of eyes, where it is one half to two thirds wider than at vertex,

parafrontalia a third to a quarter as wide as interfrontalia in front, less than a

third as wide as interfrontalia in middle, with 3-5 pairs (of which 2-3 are

strong) of inclinate frontals. Inner vertical strong, outer vertical and post-

verticals fine though rather long, ocellars very long and fine. Antennae black,

apex of second joint silver-dusted interiorly, third joint not quite thrice as

long as wide, apex far apart from mouth-margin, but reaching lower fifth of

eyes, the latter reaching level of mouth-margin; arista bare in apical half,

short-plumose in basal half (except where it is thickened), the longest hairs

less than half as long as the width of the third antennal joint. Palpi black.

Parafacialia less than half as wide, jowls as wide as third antennal joint.

Thorax black, grey dusted, with three dark brown dorsal vittae, a rather

broad median vitta which reaches neck, tapers in front, almost reaches the prst

dc laterally, and ends just in front of scutellum, and a pair of sublateral vittae

along the za which are gradually and slightly curved inward in front and behind.

acy fine, in two rows, a few additional hairs behind, 1 + 3 strong dc, two rather

strong za, the anterior one just before level of the sa; scutellum sparsely

setulose, with the normal two pairs of strong setae. Two propleurals and

prostigmatals, the latter surrounded by some fine hairs. s¢f/ in a triangle, the

posterior side of which is much shorter, and which bears rather numerous, long,

fine, erect hairs, anterior stf/ weak, the two others very strong, especially the

upper one.

Abdomen long and slender, somewhat depressed on dorsum but compressed

at apex, black, densely grey dusted, with a narrow, brown median vitta and

sometimes a pair of small brown spots on the third or second and third segments.

In the specimen from Mt. Karangora the brown spots are large and incompletely

fused with the median vitta, and a pair of small spots is also present on the

first and fourth segments; second segment with a pair of lateral setae, third

and fourth segments with two pairs of setae in a transverse discal row, fifth

with 1-2 pairs of setae and some long, erect hairs.

Legs very long and slender, coxae black, grey dusted, trochanters ferru-

ginous, at least at apex, front femora black, except for a narrow yellowish

apex, mid-femora with the basal half, hind femora with the basal two-thirds

ferruginous, the apex black, tibiae ferruginous, infuscate in middle, tarsi

piceous. All the femora with numerous, fine, long, erect hairs, tibiae with dense

and long hair on more than apical half of interior and posterior surfaces, the

submedian fv of the front tibiae very fine and long, the fd of mid-tibiae slightly

beyond middle, very fine and long, the ad almost on second third, moderately

MUSCIDAE: B.—COENOSIINAE 167

long, hind tibiae with the pd somewhat before, the a and ad somewhat beyond

middle, the pd and a-very fine, the ad very strong and long.

Wings somewhat smoky, veins brown, testaceous at base, 74,, and m

slightly converging towards apex, but slightly sinuate, so that they diverge

slightly at extreme apex, 7—m slightly beyond middle of discal cell. Calyptrae

whitish-hyaline with yellowish-white border, the lower one strongly projecting,

largely subparallel with strongly rounded apex. Halteres pale yellow.

2 similar to male, abdomen fusiform, pointed at apex. Femora and tibiae

with the normal short appressed hairs, the setae stronger and much more

conspicuous.

UGANDA: Kigezi District, Mt. Mgahinga, x1.1934 (F.W.E.) 10,000-11,000 ft.

dg type; 8000 ft. 1 § 2 2 paratypes; Mt. Sabinio, xi.1934, 8000 ft. (F.W.E).

I 2 paratype. Ruwenzori, Mt. Karangora, 9900 ft. (F.W.E.) 1 g paratype.

In none of the other specimens are the hairs on the apical half of the tibiae

as long as in the type, where the length of many of them reaches or even

exceeds twice the tibial diameter. In the females the a and ad of the hind

tibiae are inserted in middle, in the male from Mt. Karangora the fd is inserted

practically in middle.

Caricea distitarsalis sp. n. g °

Length 4-7-5 mm.; of wings 5—5°5 mm.

g. Head black, densely grey dusted, the interfrontalia dull black with

strong brownish and pale grey reflections, parafrontalia, parafacialia and face

golden-grey dusted. Frons conspicuously narrowed to vertex, slightly wider

in front than one eye; fronto-facial stripe very slightly narrowed to base of

arista, gradually widened to lower margin of eyes, where it is distinctly wider

than frons at base of antennae; parafrontalia a third as wide as interfrontalia

before middle, with four (the second and fourth stronger) inclinate frontals.

Inner vertical strong, outer one weak, postverticals moderate, ocellars long.

Antennae fuscous, the second joint translucent-reddish at apex, with con-

spicuous golden-grey dust on dorso-interior surface, third joint more than thrice

as long as wide, apex slightly pointed dorsally, not reaching level of lower

margin of eyes, and separated from mouth-margin by two thirds of the length

of the third antennal joint; arista short-plumose, the hairs being longest on

dorsal surface of basal third, where they are a third as long as the width of the

third antennal joint, becoming gradually shorter and disappearing near apex.

Palpi yellow. Parafacialia almost half, jowls almost twice as wide as third

antennal joint.

Thorax black, densely grey dusted, dorsum with three broad, pale brown

vittae, the median one between the dc, along the outer base of which a narrow

vitta remains whitish-grey, narrowing to neck and on scutellum and reaching

168 RUWENZORI EXPEDITION

the apex of the latter, the sublateral ones reaching the za and frst. acr fine

and sparse, almost in one irregular row which ends in a pair of setulae in front

of scutellum, I -+ 3 strong dc, the first post pair somewhat weaker, one 7a;

scutellum sparsely setose, with two pairs of strong setae. Two rather strong

propleurals, two prostigmatals, the upper one slightly stronger than the smaller

propleural, the lower one very small, no small hairs in the vicinity. s¢pl in a

triangle, the anterior side of which is shorter, the triangle almost devoid of small

hairs, lower and anterior stfl subequal, rather strong, posterior stpl very

strong.

Abdomen moderately slender, somewhat depressed on dorsum, somewhat

compressed at apex of venter, black, densely grey dusted, with a brown median

vitta on second and third segments, a pair of conspicuous round spots on the

same segments, and a pair of faint brown spots on the first and fourth segments ;

the second segment with three pairs of moderate setae, the third with three

pairs of strong setae, the fourth with two pairs of strong setae in a transverse

discal row, fifth with a pair on hind margin. Fourth ventrite largely pale, the

margins of its excision densely beset with rather long, erect hairs.

Legs entirely testaceous, except for part of the last joint of each tarsus,

viz. half on front tarsi, two-thirds on mid-tarsi, and four-fifths on hind tarsi,

which are piceous, claws dark, empodia pale.. Front femora with 6—7 rather

strong pv, 5-6 not very strong fd setae, front tibiae with a rather strong but

not long fv, mid-femora with two strong pv near base and two preapicals

(p and fd), mid-tibiae with one ad and one fd, both not very long but quite

strong, the former somewhat longer and on three fifths, the other slightly

beyond middle, hind trochanters (as the apex of the mid and hind coxae) with

numerous, erect, fine hairs, hind femora with four moderate av, the second of

them much stronger, inserted on basal fourth, one strong fv slightly beyond

the strong av, about six rather strong ad and a pd preapical, hind tibiae with a

strong pd somewhat basad of middle and a strong a and ad (the latter a trifle

more basad) somewhat beyond middle.

Wings hyaline, the veins testaceous near base, brown on disc, 74,; and m

very slightly diverging at apex, v-m almost on three-fifths of discal cell.

Calyptrae whitish-hyaline, the border pale yellow, the lower one much longer

than the upper one, strongly projecting, largely subparallel with the apex

semicircularly rounded. Halteres pale yellow.

9 similar to male, but parafrontalia slightly narrower, third antennal joint

thrice as long as wide, longest hairs of arista hardly longer than basal diameter

of arista, only a quarter as long as the third antennal joint is wide, the pleurae,

including humeri, largely ferruginous (normal?), abdomen oblong oval, rather

strongly depressed on dorsum, first segment with a narrow median vitta and

a pair of rather large spots; femora narrowly infuscated on dorsal surface of

apical half, last joint of tarsi piceous, third and fourth more or less infuscate,

MUSCIDAE: B.—COENOSIINAE 169

front femora with 7-10 pd setae, hind trochanters with shorter and sparser

hairs, 7,,; and m not distinctly diverging at extreme apex, v—-m distinctly

beyond three fifths of discal cell, lower calyptra more subtriangular, as the

basal half is broader and not subparallel.

Kenya: N. of Nakuru, ca. 9300 ft., 6.111.1935, on Lobelia aberdarica (’.W.E.)

g type. Ucanpa: Mt. Elgon, 11,000 ft., 10.11.1934 (H. B. Johnston) 2 para-

type, greasy and in rather poor condition.

Very closely related to C. fuscifemur Mall. Most of the characters of

distitarsalis might be regarded as subspecific. The difference in the frons and

in the position of the hind tibial setae, however, would hardly warrant for the

time being uniting the two forms in one species. The ad setae of the mid

and hind femora are less numerous and less conspicuous than in /fuscifemur.

The male of distitarsalis is mature, whereas the female type of fuscifemur is

slightly teneral. The elimination of this influence would tend to increase the

difference in colour between the two species.

[Caricea vittata chyuluana ssp. n. 3]

Length 3-7 mm.; of wings 3-4 mm.

The structural characters are the same as in wittata Wied., which occurs in

Natal, but the coloration is strikingly darker: the face is velvety-black, palpi

piceous-black. The three broad vittae of the thorax, the narrow median vitta

and the paired spots of the abdomen are of the same shape as in vittata, but

dark brown. Front femora with the p and fd surfaces piceous on apical two-

fifths, front tibiae and tarsi strongly infuscate, mid and hind femora piceous

on apical third to quarter, mid and hind tibiae and tarsi very strongly infuscate,

largely piceous, the base of the tibiae and apex of the tarsi slightly paler. There

is a dark spot on each side of the dorsal half of occiput and another one on

and around the anterior prothoracic spiracle, which does not affect the propleural

depression. The wing-veins are brown.

Kenya: Chyulu Hills, 6000 ft., vi.r938 (Coryndon Museum Expedition)

3 type.

[Caricea somereni sp. n. 3 9]

Length 3:3-3°9 mm.; of wings 3°I-3°4 mm.

3. Head black, densely grey dusted, a broad stripe from inner verticals to

neck, a small spot at the base of the reclinate and a larger one, surrounding

the bases of the 2—3 inclinate frontal setae, dark brown dusted, interfrontalia

velvety black. Frons slightly longer than wide in front, distinctly narrowed

to vertex, interfrontalia strongly narrowed to middle, two-thirds as wide

behind as in front, fronto-facial stripe hardly narrowed to middle of face,

I70 RUWENZORI EXPEDITION

slightly widened from there to lower margin of eyes, where it is not or only

slightly wider than on frons; parafrontalia a fifth as wide in front, a third as

wide behind as interfrontalia, distinctly protruding anteriorly in profile, the

face conspicuously receding. Inner vertical rather strong but of moderate

length, outer vertical indistinct, postverticals and ocellars small, the latter

slightly more than a quarter of the length of the frons. Antennae dull black

with greyish reflections, especially on interior surface of second and in apical

part of third joint, the latter thrice as long as wide, approaching though not

quite reaching mouth-margin, which is practically level with the lower margin

of eyes, arista almost bare on ventral surface, longest hairs of dorsal surface

not quite as long as the basal diameter of arista. Parafacialia almost linear

(except near base of antennae), jowls almost as wide as third antennal joint.

Thorax black, densely grey dusted, with three broad, dark brown dusted

vittae, the median one slightly widened from neck to second post dc, reaching

first and second post dc at sides and occupying all the scutellum, except for a

small spot of grey dust between the apical setae, the lateral vittae curved from

level cf humeral seta and the very small p/ to base of wing, usually fused with

the median vitta at the prst dc and the first post dc; pleurae with a dark brown

spot on and around anterior spiracle (but not extending on to propleural depres-

sion). acy sparse and small, the two rows very close together, almost appearing

like one row in places, 1 + 3 strong dc, one small 7a; scutellum with a few

setulose hairs and two pairs of strong setae, the lateral pair distinctly shorter.

Two propleurals and prostigmatals, lower prostigmatal small, curved down-

ward, not surrounded by small hairs. s¢p/ in a triangle, which is bare, and the

anterior side of which is hardly more than half as long as the others, lower and

anterior stl subequal, not very long though quite strong, posterior one long

and strong.

Abdomen elongate-oval, strongly convex, very slightly depressed at base,

somewhat compressed at apex, black, densely grey dusted, with a very con-

spicuous, dark brown dusted median vitta from middle of first to apex of fourth

segment and a pair of rather large, subquadrate spots which broadly reach

hind margin and are sometimes narrowly connected along hind margin with

the median vitta, spots of the fourth segment sometimes fused with median

vitta, in dorsal view the spots reach almost the sides, so that apparently only

a rectangularly bent stripe along median vitta and front margin remains of

the grey dust. Sublateral setae well developed, third segment with a transverse

discal row of two pairs of setae, fourth with a pair of strong paramedian discal

setae and a weaker pair outward and apicad of it, fifth with a pair of long, fine

setae rather far apart and two pairs of smaller setae between them. Lobes of

the fourth ventrite with numerous erect hairs on inner margin, which are not

very long but rather strong.

Legs wholly piceous black, coxae densely grey dusted, femora shining. Front

MUSCIDAE: B.—COENOSIINAE 7X

femora with the pd and pv setae small and fine, four somewhat stronger pd

in apical half, front tibiae with a rather strong but not very long fv seta slightly

beyond middle; mid-femora with 4-5 small ad from base to shortly beyond

middle, the last few of them rather strong (though not long), 3-4 fine and

rather small pv in somewhat more than basal half, and two preapicals (f and

pd), mid-tibiae with a strong ad almost at two-thirds and a much finer pd

not much beyond the middle; hind femora with five rather strong though not

very long ad, a strong pd preapical, 4-5 fine and small av, and about five small

and fine pv, among which a longer one is inserted just beyond basal third, hind

tibiae with a strong pd on middle, a strong ad not much beyond the pd

(approximately by the tibial diameter) and a rather small a at the same

level.

Wings very slightly smoky, veins brown, 7,,, and m slightly converging to

apex, v—m slightly beyond middle of discal cell. Calyptrae whitish, with

whitish border and fringe, extremely short, the lower one still shorter than

the upper one, entirely covered by it. Halteres pale yellow.

Q similar to male, abdomen more depressed, pointed at apex, second to

fourth segments with a narrow dark brown anterior border in addition to the

pattern of the male.

Kenya: Chyulu Hills, 6000 ft., vi. and vii.1938 ¢ type (in B.M.),2 32 9

paratypes (in B.M. and Coryndon Memorial Museum, Nairobi).

Caricea edwardsi sp. n. ¢ @

Length 4:6-5:9 mm.; of wings 4:3-6-2 mm.

6. Head black, densely grey dusted, median dorsal part of occiput and

posterior part of parafrontalia dark brown dusted, interfrontalia dull black

with strong brown reflections when seen from in front. Frons slightly longer

than face, not quite twice as long as wide at vertex, distinctly dilated from

vertex to lunula, slightly wider in front than one eye, fronto-facial stripe sub-

parallel from there to apex of second antennal joint, gradually dilated and

rounded to lower margin of eyes, where it is a third wider than at lunula;

parafrontalia almost a quarter as wide in front, almost a third as wide behind

middle as interfrontalia, with two strong and I~3 (one of them sometimes

rather long) small inclinate frontal setae. Inner vertical strong, outer vertical

and postverticals moderate, ocellars strong, two-thirds as long as frons. Antennae

black, second joint pale dusted interiorly at apex, third joint slightly more

than twice as long as wide, reaching lower sixth of eyes (the latter not quite

reaching level of mouth-margin), falling short of mouth-margin by two-thirds

of the length of the third antennal joint; arista almost bare on apical half,

except for a few short hairs on dorsal surface, short-plumose in basal half, two-

II, 4(f)

172 RUWENZORI EXPEDITION

thirds or three-quarters as wide, including plumosity, as third antennal joint.

Parafacialia approximately a third as wide as, jowls slightly wider than third

antennal joint.

Thorax black, densely grey dusted, dorsum with three broad, brown vittae,

the median one slightly dilated from neck to scutellum, sublateral ones form-

ing a long, inwardly and backwardly curved hook in front, passing through

the prst and ph pores to almost neck and then curved backwards and reaching

the prst dc and suture where they are fused with the median vitta. acr very

sparse and inconspicuous, two rows hardly recognisable; 1 + 3 strong de,

va lacking, scutellum with a few setulose hairs and rather weak lateral and

rather strong apical pairs of setae. Propleural setae strong, prostigmatals

rather weak, the lower one curved downward, very fine though rather long.

stpl in an almost equilateral triangle, the posterior side of which is hardly

shorter, and which bears rather numerous, long, erect hairs, anterior stpl

rather weak, the two others very long and strong, the lower one slightly shorter.

Abdomen rather elongate, slightly narrowed to apex, somewhat depressed

at base, strongly compressed at apex, black, densely grey dusted, first to third

segments with a broad, dark brown band along the hind margin, which reaches

front margin in middle on second and third segments, fourth segment with a

broad median vitta which is more or less extended sideward in middle, the

transverse bands often tending to disintegrate into a median vitta and a pair

of spots. First abdominal segment with the usual vertical fan of setae at sides,

second to fourth segments each with a strong lateral and a moderate sublateral

discal seta, the sublateral seta of the fourth segment strong, this segment

also with 2-3 pairs of finer marginal setae. Hypopygium with fine and rather

short hairs, superior forceps fused, elongate trapezoidal, emarginate at apex,

basal half covered with microscopic pile, apical half bare and glossy, rufescent;

fourth ventrite with rather numerous and long hairs.

Legs with black, densely grey-dusted coxae, front femora black with a

narrow ferruginous apex, sometimes the basal half reddish anteriorly, mid

and hind femora ferruginous with the apical third to sixth black, tibiae and

tarsi piceous, the former with ferruginous basal third or quarter. Front

femora with 5-6 pd, three rather long sub-basal a setae, and five pu which alter-

nate with hairs, front tibiae with a rather fine fu seta beyond middle; mid-

femora with 6-7 av setae from base to second third, 3-4 strong a and four long

pv setae from base to two-thirds, a pv at two-fifths particularly strong, two

strong (p and pd) preapical setae, mid-tibiae with a moderate fd in or beyond

middle, and a similar ad slightly beyond it, hind femora with 6—7 rather strong

ad, about nine, mostly fine av among which two strong ones at a third and two

thirds, pu setae fine and not very distinct, except for a strong one at two-fifths

and a smaller one at three-fifths, one strong pd preapical, hind tibiae with a

strong ad in middle, a smaller a and fd level with it, seldom one of them slightly

MUSCIDAE: B.—COENOSIINAE E73,

more basad or apicad (in one specimen the fd is inserted distinctly basad of

the ad).

Wings slightly smoky, with brown veins which are slightly paler at base,

74,5 and m rather strongly curved to apex, slightly diverging at extreme apex,

y—m distinctly beyond middle of discal cell. Calyptrae hyaline-yellowish with

pale yellow margins, lower one strongly projecting, subparallel, strongly rounded

at apex. Halteres yellow.

2 similar to male, the abdomen pointed with the pattern less conspicuous.

UcanpDA: Kigezi District, Mt. Mgahinga, 10,000-11,000 ft., 22.xi.1934

(F.W.E.) g type, 4d 9 2 paratypes; Mt. Sabinio, 8000 ft., xi.1934 (F.W.E.)

I g 1 Y paratypes; Kanaba, 7800 ft., xi.1934 (P.W.E.) 2 g 2 9 paratypes.

In the specimens from Mt. Sabinio and Kanaba the vittae of the thorax are

not connected in front, the recurved hook being absent, and the setae, especially

the ocellars and outer verticals, are somewhat shorter.

Caricea ruwenzorica sp. n. 3d &

Length 4:8-5:7 mm.; of wings 5:3-6°3 mm.

3. Head black, densely grey dusted, with a dark stripe from vertex to

occipital foramen, interfrontalia black with strong, brown reflections in anterior

view, frontal triangle and parafrontalia brown dusted. Frons not protruding

in profile, more than twice as long as wide, approximately half as wide as one

eye, parallel from vertex to last inclinate frontals, fronto-facial stripe slightly

narrowed from there to base of arista, rather strongly dilated and rounded to

lower margin of eyes, where it is one and a half times as wide as frons; para-

frontalia a third to a quarter as wide as the interfrontalia, with two strong

and 1-3 small inclinate frontal setae. Inner vertical very strong, outer vertical

very distinct though not long, postverticals slightly longer than outer verticals,

ocellars much smaller. Antennae fuscous, second joint with golden-brown

dust interiorly at apex, third joint greyish dusted on apical half, almost thrice

as long as wide, falling short of mouth-margin (which is slightly passed by

the eyes) by the length of the third antennal joint, arista almost bare, sparsely

short-haired in basal half, the longest hairs not twice as long as the basal

diameter of the arista. Parafacialia linear, jowls a third as wide as third

antennal segment.

Thorax black, densely grey dusted, dorsum brown dusted, with five not very

well defined darker brown vittae, the median one reaching neck and continued

over disc of scutellum, the subdorsal and sublateral ones connected in front

at ph. acy very small, fine and sparse, mainly in one irregular row, a few more

setulae present in front of scutellum, 1 + 3 long dc, two moderate za, scutellum

with some fine setulae and two pairs of very strong setae. One fine propleural,

the upper prostigmatal stronger though rather fine, the lower one fine and

174 RUWENZORI EXPEDITION

curved downward, not surrounded by small hairs. s¢f/ in an almost equilateral

triangle, which bears some fine, erect hairs, anterior st// weak, lower one strong,

posterior one very strong.

Abdomen elongate-oval, much longer than the thorax, slightly depressed,

apex compressed, dorsum almost entirely dark brown dusted, with a very

narrow brownish-grey dusted hind margin on first to fourth segments, basal

half of first segment thinly grey dusted. Lateral setae of abdomen well developed,

fifth segment with a pair of fine, long setae among the hairs. Fourth ventrite

with rather dense, fine and short hairs along inner margin of the lobes.

Legs very long and slender, yellow, with the apical third to quarter of the

femora and a long dorsal streak on the anterior femora piceous, posterior tibiae

strongly infuscate to piceous, front tibiae slightly infuscate, base of tibiae

paler, tarsi piceous, first joint of hind tarsi rather strongly compressed. Front

femora with five not very long but quite strong fd setae, a row of fine but

distinct p setae, two of them close to base rather long, 5—6 pv, alternating with

fine setae, three of them from near base to third fifth very long, front tibiae

with a very long and strong fv slightly beyond middle: mid-femora with about

six fine and short av in basal half, 4-5 a setae, a strong one at middle and

another one somewhat beyond middle, three very long fv from a quarter to

three quarters, and two (pf and pd) strong preapicals, mid-tibiae with a strong

pd in middle and a strong and very long ad at three-fifths or two-thirds; hind

femora with 2~4 rather fine ad in basal two-fifths, no ad from there to second

third, and two strong ad in apical third, an av row of six setae which are fine

and short at base and increase gradually in size, three very long pv at a third,

a half and two-thirds, and a strong fd preapical, hind tibiae with a very long

and strong ad at two-fifths, a slightly weaker a practically level with it (very

slightly more apicad) and a strong pd just beyond middle.

Wings rather smoky, veins brown, somewhat paler at base, 74,; and m

very slightly curved to apex, almost parallel, just noticeably diverging; 7-m

slightly beyond middle of discal cell. Calyptrae somewhat smoky, margin

brownish, lower one very long and parallel, rather narrowly rounded at apex,

projecting beyond upper one by more than twice the length of the latter.

Halteres pale yellow.

2 only distinct from male by the stouter, more depressed, more oval, and

pointed abdomen.

UGANDA: Ruwenzori, Namwamba Valley, 6500 ft. (F.W.E.) 3 type, 2 3

3 9 paratypes; Mobuku Valley 7300 ft. (F.W.E.) 3 g paratypes; Bwamba

Pass (west side), 5500-7500 ft. (/.W.E.) 4 $ paratypes; Fort Portal, 4.xii.1934

(F.W.E.) 1g paratype.

In a paler form from Namwamba Valley (1 g) and Mobuku Valley (2 3, 1 9)

the abdominal segments have broader grey transverse bands along the lateral

parts of the front and hind margins, the thoracic vittae are slightly narrower

MUSCIDAE: B.—COENOSIINAE 175

and therefore more detached from each other, the calyptrae are hyaline with

whitish border. In the Namwamba male, which is slightly teneral, the hind

tibiae are, moreover, hardly infuscate at all. In pattern this pale form would

seem to resemble colovata Curr., but the body is black, whereas it is reddish

yellow in colovata. This coloration cannot be due to immaturity, as Curran’s

series consisted of seven specimens collected in the field.

Caricea bipila sp. n. 5

Length 5*5-5:8 mm.; of wings 5:8-6-I mm.

3g. Head black, grey-pollinose, except on interfrontalia which are dull

black with brown reflections, parafrontalia and parafacialia which are yellowish-

grey dusted. Frons almost as wide as one eye, slightly narrowed to vertex,

fronto-facial stripe almost parallel on median third, rather strongly widened

to ventral end of eyes; parafrontalia not quite half as wide in middle as inter-

frontalia, with two strong, and, between them, one or two small to moderate

inclinate frontal setae. Inner vertical strong, outer vertical weak, postverticals

and ocellars long, more than two thirds as long as frons. Antennae fuscous,

the second joint silver-grey dusted at apex on inner dorsal surface, third joint

almost thrice as long as wide, its apex rather far from level of lower eye-margins,

the latter slightly above epistoma, arista short-haired on basal half, the hairs

very sparse on ventral surface, the longest hairs hardly a third as long as the

third antennal joint wide. Parafacialia a third as wide below middle as the

third antennal joint, jowls as wide as third antennal joint.

Thorax black, grey-dusted, the dorsum yellowish-grey dusted with five

vittae, the median one narrow, reaching almost from neck to almost base of

scutellum, subdorsal vitta narrow, reaching from prst to third post dc and more

or less distinctly continued at or to scutellars, lateral vitta broader and shorter,

interrupted at suture. acy fine though distinct, in two rows, no distinct prsc,

1 + 3 strong dc, two ia, the anterior one slightly before level of sa, scutellum

with a few small setulae and a lateral pair of very long and strong setae. Two

rather fine propleurals, the lower one much weaker, two prostigmatals, similar

to the propleurals, surrounded by a few fine though rather long hairs. stf/ in

a triangle which bears some fine and long, erect hairs, and the posterior side

of which is considerably shorter, anterior stfl weak, the two upper stfl very

long and strong.

Abdomen black, parallel and slender in dorsal view, somewhat club-shaped

in lateral view, grey dusted, with a faint and narrow brown median vitta and

a pair of rounded spots on segments 1-4, which are small on 1 and 4, lateral

seta distinct though decreasing in length from first to third segment, much

stronger on fourth segment, where, in addition, a small subdorsal seta is present,

fifth segment with a pair of rather strong, decussate setae.

176 RUWENZORI EXPEDITION

Legs testaceous-yellow, tarsi, apical quarter of posterior femora and a shadow

on dorsal surface of apical quarter of front femora piceous; very slender and

long, especially the front tarsi. Posterior surface of front femora and ventral

surfaces of posterior femora with rather numerous erect, fine, long, soft hairs.

Front tibiae with a fine pu seta, mid-femora with two preapicals (p and pd),

four rather long a setae and three long fv, mid-tibiae with a weak ad and a

moderately strong fd seta, hind femora with six long ad, three very long pu

and a fd preapical, hind tibiae without a fd seta but with about six half-erect

setulose hairs around middle of fd surface, a very slightly basad of ad, both fine.

Wings hyaline, slightly infuscate, veins brown, 7,,; and m parallel at apex,

y-m slightly beyond middle of discal cell. Calyptrae hyaline, border yellow,

small, the lower one, however, rather strongly projecting, subtriangular,

narrowly rounded at apex. Halteres pale yellow.

2 similar to male, the eyes slightly larger and the frons, therefore, slightly

narrower than one eye, ocellars hardly more than half as long as frons. Abdomen

elongate-fusiform, the paired spots larger on first to third segments, indistinct

on fourth. The legs without erect soft hairs, but with the usual appressed hairs

and with the setae much stronger and more conspicuous. Tarsi not so long nor

so slender.

Ucanpa: Kigezi District, Mt. Mgahinga 10,000-11,000 ft., xi.1934 (F.W.E.)

6 type, 2 2 paratypes.

Caricea humilis group

t(10) Frons very slightly projecting beyond eyes in profile, considerably narrower

above antennae than one eye. Eyes reaching level of mouth-margin,

antennae of moderate length, apex conspicuously above level of lower

margin of eyes.

2(5). Frons less than half as wide above antennae as one eye. Pulvilli and claws

rather long. Arista rather long-plumose near base.

3(4). Tarsi pale yellow, except for the last joint, which is infuscate. 3:75-5 mm.

The longest hairs of arista slightly more than half as long as the third

antennal joint. Frons at its narrowest point hardly more than a third

the width of one eye, narrowed from base of antennae to the second

mclinate frontal seta . : : : : [ochvoprocta Speis. |

4(3). Tarsi entirely infuscate. 5:5-6:-5 mm. Arista with very long hairs near

base, bare at apex : : é ; 8 é [*angustifyons Stein]

5(2). rons at least half as wide at narrowest point as one eye (Fig. 74). Pulvilli

and claws short. 3-4 mm.

6(7). Arista long-plumose in basal half, the longest hairs fully as long as the third

antennal joint from arista to apex. Tibiae largely infuscate. 74,5

rather strongly curved downward, the extreme apex, however, slightly

upturned, so that 7,;, and m are slightly diverging at tip.

[setalis sp. n.], p. 177

7(6). Arista pubescent or short-plumose, the longest hairs not longer than the

third antennal joint wide. Tibiae seldom conspicuously infuscate.

MUSCIDAE: B.—COENOSIINAE 177

8(9). Arista short-plumose, about as wide, including plumosity, as the third an-

tennal joint. Frons slightly more than one-half as wide above base of

antennae as one eye (Fig. 74). 744, very gently curved, reaching costa

some distance in front of the wing-tip, very slightly diverging with m

at apex |. : d humilis multimaculata Ad., p. 179

9(8). Arista pubescent in el Hale fie hairs about as long as the base of arista

wide. Frons two-thirds as wide above the base of the antennae as one

eye. 744, rather strongly curved, reaching costa almost in wing-tip,

slightly converging with m at apex : : [curvinervis sp. n.], p. 179

1o(1). Frons projecting beyond eyes by more than width of the third antennal

joint, about three-quarters as wide as one eye, or wider. Eyes not

reaching level of mouth-margin, third antennal joint long, reaching

ventral margin of eyes.1_ Head almost as long as high, eyes less high

than in the species under 1. Abdomen with a brown median vitta and

paired spots. 3-4°5 mm.

11(xz2). Frons almost as wide as one eye, slightly dilated to vertex, with 3—4 inclinate

setae. Hind tibiae without long erect hairs in either sex, though with

the normal setae and an additional preapical (antero-subdorsal between

d and ad on front tibiae). Parafacialia rather wide, not much narrower

than third antennal joint. 3-4mm. . a : [inanis Stein]

12(11). Irons three-quarters as wide as one eye, parallel to vertex, with 4-5 in-

clinate setae. Hind tibiae of male with some long, erect, bristly hairs

on ventral surface. Parafacialia very narrow. Mid-tibiae almost

entirely yellow, the others yellow at base. 445 mm. Madagascar.

[*tvichocnema Stein]

[Caricea setalis sp. n. 9]

Length 3-4 mm.; of wings 2-8 mm.

Head black, densely grey dusted, interfrontalia black, with whitish reflections

when seen from in front, parafrontalia, face and jowls silver-grey dusted. Frons

not quite three-fifths as wide anteriorly as one eye, very slightly dilated to

vertex, almost twice as long as wide in front, fronto-facial stripe gradually

dilated with straight sides from base of antennae to lower margin of eyes,

where it is more than one and a half times as wide as at vertex; parafrontalia

a third as wide as interfrontalia, with three inclinate (the second of them

smaller) frontal setae. Inner vertical strong, outer vertical absent, postverticals

moderate, ocellars rather strong. Antennae fuscous, second joint silver-grey

dusted interiorly at apex, the latter narrowly reddish translucent, third joint

thrice as long as wide, somewhat acuminate dorsally at apex, reaching lower

sixth of eyes but distant from mouth-margin by half the length of the segment,

the eyes practically reaching level of mouth-margin; arista very long-haired

1 Coenosia costata Stein from Durban apparently runs down to this group of species.

It may be distinguished by its frons which is conspicuously broader than one eye, the jowls

which are more than half as broad as the eyes high, the distinct though small costal spine,

and the slightly infuscate cross-veins.

2 This species should run down to the humilis group in the key on pp. 157-160, but it is

repeated in the semifuwmosa group, as two pairs of tibiae are evidently coloured as in that

group and the third is blackish at tip.

178 RUWENZORI-EXPEDITION

at base, the longest hairs at least as long as the third antennal joint from base

of arista to apex, the hairs quickly decreasing in length and quite short at

apex. Palpi pale brown. Parafacialia half as wide, jowls not quite as wide

as third antennal joint.

Thorax black, densely grey-dusted, with a narrow median vitta between

the acr, which is interrupted or subinterrupted behind suture, slightly dilated

towards scutellum and occupies the base and disc of the latter, besides with a

rudimentary vitta or rather a row of small spots at the bases of the dc. acr

small but well developed, in two rows before suture, in four rows behind it,

1 + 3 long and strong dc, posterior 7a small; scutellum setulose, with two pairs

of strong setae, the lateral ones slightly shorter. Two rather strong propleurals

and prostigmatals, the lower prostigmatal curved downward, surrounded by

a few small hairs. stl in an almost equilateral triangle, the posterior side

slightly longer, the area with a few erect hairs, lower stf/ somewhat shorter,

posterior one much longer than the anterior one.

Abdomen black, rather thinly grey dusted, with a dark-brown dusted median

vitta and a pair of rather large rounded spots on each segment; lateral setae

rather strong, third and fourth segments each with a transverse discal row of

two pairs of setae.

Legs with the coxae, trochanters and femora black, tibiae ferruginous,

rather strongly infuscate, tarsi strongly infuscate. Front femora with three

strong d setae towards apex, three rather long fd on basal two-fifths, four short

ones from there to apex, and six rather long and fine fv setae, front tibiae with

a strong and long fv in middle, apex with a strong additional ad preapical seta

(five strong preapicals in all), mid-femora with the av setae fine and short,

three strong a setae from two-fifths to apex, three rather long pu between base

and three-fifths and two preapicals (f and fd), mid-tibiae with a moderate pd

slightly beyond middle and a strong and long ad slightly beyond it ; hind femora

with six strong ad, two long av at one-third and two-thirds, three pv, two of

which—before and behind middle—are rather long, and a fd preapical, hind

tibiae with a very long ad slightly before middle and a long a very little

beyond it.

Wings hyaline, veins pale brown with testaceous base, 7,,, rather strongly

curved downward but slightly simuate before apex, and the latter very slightly

diverging with m, v—-m distinctly beyond middle of discal cell. Calyptrae

whitish with yellowish-white border and fringe, the lower one strongly pro-

jecting and broadly rounded at apex. Halteres pale yellow.

NaTAL: Durban (V. Muir, Sharp Coll.) 9 type; ZANZIBAR, 1.-ii.1925 (H. /.

Snell) 2° paratype (abdomen missing). PortuGc. E. Arrica: Beira, vi.1932

(Miss A. Mackie, J. Ogilvie) 2 2 paratypes. The type is the specimen men-

tioned by Malloch (1922, Ann. Mag. nat. Hist. (9) 10: 582) under tripunctiventris

as “‘probably another species”’.

MUSCIDAE: B.—COENOSIINAE 179

Caricea humilis multimaculata Ad.

UcanvA: Kigezi Dist. Mabungo Camp, 6000 ft., xi.34 (J. Ford) 1 3.

Typical locality: S. Ruopesra: Salisbury. In B.M. from typical locality,

NATAL (Estcourt; Port Shepstone; Willow Grange; Hill Brook), NyASALAND

(Ruo), AByssinIA and SupAN (W. Darfur, Jebel Murra, Killing, 7000 ft.,

7.1V.1932, (Miss M. Steele)).

The African form is certainly not quite identical with the European humilis

Meig., though it was identified thus by Stein, Malloch, and Curran. The frons

is hardly more than half as wide as one eye (Fig. 74), whereas in European

specimens it is about three-fifths to three-quarters as wide as one eye, the

difference in width lying entirely in the interfrontalia. The tibiae are, as a rule,

less infuscate than generally in European specimens, and the plumosity of the

arista is longer than in the latter. The curvature of 7,,; and m, which are

distinctly diverging at apex, is as in humilis s.str.—I have, through the courtesy

of Prof. H. B. Hungerford, seen Adams’ types.

[Caricea curvinervis sp. n. 9]

Length 2.8 mm.; of wings 2-7 mm.

Head black, densely grey dusted, frontal triangle ochraceous dusted, face

silver-grey dusted, interfrontalia dull black, with brownish-white reflections

when seen from in front. Frons projecting in profile by about one half the width

of the third antennal joint, not quite twice as long as wide in front, longer than

face; subparallel, very slightly narrowed from the reclinate seta to vertex,

apparently two-thirds (slightly more than three-quarters if measured) as wide

as one eye, fronto-facial stripe very slightly narrowed to apex of second antennal

joint, moderately dilated and almost straight to lower margin of eyes, where

it is one and a half times as wide as on frons, parafrontalia a third as wide as

interfrontalia, with two moderate and one minute inclinate seta. Inner

verticals rather strong, outer ones indistinct, postverticals and ocellars short

but distinct. Antennae blackish, second joint silver-grey dusted interiorly at

apex, third joint not quite thrice as long as wide, reaching lower sixth of eyes

(the latter practically reaching level of mouth-margin), falling short of mouth-

margin by half the length of the joint; arista bare on more than apical half,

pubescent on basal half, the longest hairs hardly longer than basal diameter of

arista. Palpi fuscous, the base slightly paler. Parafacialia less than half as

wide, jowls not quite as wide as third antennal joint.

Thorax black, rather densely grey dusted, rather shining, dorsum with a

conspicuous, narrow, dark brown median vitta and traces of a pair of linear

vittae along the dc. acy very distinct though small, in two rows, I + 3 de,

two well developed za; scutellum with two pairs of setulose hairs and the

180 RUWENZORI EXPEDITION

normal two pairs of strong setae. Propleural and upper prostigmatal setae

subequal, well developed though not strong, lower prostigmatal small, curved

downward. stf/ in an equilateral triangle which bears some erect hairs,

anterior and lower stfl not very strong, posterior stfl long and strong.

Abdomen oval, pointed at apex, rather depressed, rather densely grey dusted

but nevertheless rather glossy, second and third segments and base of fourth

with a brown median vitta, first to third with a pair of rather large brown,

roundish spots which are essentially confined to posterior half. Lateral setae

of second and third segments rather small, fourth with a transverse row of

two pairs of rather smali setae before apex.

Legs with black, grey dusted coxae, trochanters and femora black, slightly

dusted and rather glossy, tibiae testaceous, hind ones somewhat brownish in

middle, tarsi piceous. Front femora with four fd in apical half, the second of

them stronger, 5-6 small # setae of which the basal two are longer, and 6-7

pv alternating with fine ones, front tibiae with a long fv in middle; mid-femora

with five a setae, the last at two-thirds and rather strong, six fine and short av

in basal half, four long fv, and two strong preapicals (f and fd), mid-tibiae

with a rather strong pd at middle and a strong ad at three-fifths; hind femora

with six rather strong ad and av setae, three rather fine fv, alternating with very

small ones, and a strong fd preapical, hind tibiae with a very long ad in middle

and a strong a slightly basad of it.

Wings hyaline, strongly iridescent, veins brown with testaceous base,

7445 rather strongly curved backward, ending almost in wing-tip, m slightly

converging with it at apex, ym at three-fifths of discal cell. Calyptrae whitish-

hyaline, with yellowish-white borders, lower one slightly dilated from base to

middle, broadly rounded at apex, rather strongly projecting beyond upper

one, not quite twice as long as it. Halteres yellowish-white.

S. RHopeEsiA: Salisbury, 109.111.1939, prey a Chironomid (A. Cuthbertson)

2 type.

Caricea tigrina group

1(2). Parafrontalia pale dusted. Vibrissae just above mouth-margin, not more

distant from it than by the apical width of the palpi. Frons distinctly

narrower than one eye. All the femora distinctly yellow at apex.

Base of m,-+-cu, simple in both sexes. Thorax and first abdominal

segment without pattern. Hind tibiae with an additional preapical seta.

(Mediterranean, etc.) E 5 . ‘ [tigrina F.]

2(1). Parafrontalia largely velvety black, not differentiated from interfrontalia.

Vibrissae distant from mouth-margin by the width of the third an-

tennal joint. Front femora very narrowly, mid and hind femora not

at all yellow at apex. Base of m,-+-cu, thickened and darkened opposite

origin of 7,,, in male. Thorax with a pair of broad longitudinal vittae

along dc to ia, and a narrow median vitta, which may be fused in front of

suture. First abdominal segment usually with a distinct pair of spots.

MUSCIDAE: B.—-COENOSIINAE 181

3(4). Arista short-haired, the longest hairs hardly longer than the base of arista

wide (Fig. 68). Tarsi piceous black. Vittae of thorax fused or narrowly

separated in front. Legs with the usual short setulae, ventral spur of

the hind tibiae about as long as the diameter of the hind femora.

; cubitalis sp. n., p. 181

4(3). Arista short-plumose, the longest hairs almost as long as the third antennal

joint wide (Fig. 69). Tarsal joints 1-4 yellow, 5 piceous. Vittae of thorax

broadly separated throughout. @: First joint of hind tarsi long and

broad, legs rather hairy, hind tibiae and tarsi with dense hairs, which

are longer than the tibial diameter; ventral spur of the hind tibiae

long and curved (Fig. 72), twice as long as the diameter of the hind

femora f 0 : . é c hippelates sp. n., p. 183

Caricea cubitalis sp. n. 3 °

Length 4:7-6-9 mm.; of wings 4:8—7-I mm.

6. Head black, occiput rather thinly, face, parafacialia, jowls and temples

densely whitish-grey dusted, in certain lights dark grey, frons including para-

frontalia wholly brownish-black, with brownish reflections. Frons projecting

in profile by width of third antennal joint, as long as face, almost one half

longer than wide, slightly to considerably wider in front than one eye, slightly

narrowed to vertex, face gradually widened with rounded sides from lunula

to lower margin of eyes, where it is one and a half times as wide as frons;

parafrontalia not very distinct, a third as wide as interfrontalia, with 5~-7

inclinate frontals, the second and last of them strong, the fourth or fifth usually

moderately strong, the others moderate to small. Inner vertical very strong,

outer vertical strong to moderate, postverticals strong, ocellars very strong.

Antennae fuscous, second joint densely silver-grey dusted interiorly at apex,

third thrice as long as wide or slightly longer, apex reaching lower sixth to

eighth of eyes (the latter falling short of mouth-margin by at least width of third

antennal joint), as distant from mouth-margin as from base of arista; the

latter short-haired, the longest hairs as long as or slightly longer than the

basal diameter of arista (Fig. 68). Palpi fuscous. Parafacialia at least as wide

as third antennal joint, jowls at least one and a half times as wide, vibrissae

above mouth-margin by width of the third antennal joint.

Thorax black, densely dark grey dusted, brownish on dorsum, with greenish-

blue reflections on pleurae, mesopleura with a large dark-brown spot from

dorsal margin to middle, dorsum with the following not very striking pattern:

a narrow median vitta and a pair of broad vittae from inward of the de to out-

ward of the za, the vittae often more or less fused in front of suture, reaching

centre of scutellum and lateral scutellar setae respectively. acy rather strong

though short, in two irregular rows, I + 3 very strong dc, posterior za strong,

anterior one strong to absent; scutellum with short setulae and two pairs of

182 RUWENZORI EXPEDITION

strong setae. Upper prostigmatal and propleural rather strong, lower prostig-

matal fine but rather long, curved downward, surrounded by a few small

hairs. stpl in an equilateral triangle, which bears rather numerous and strong,

erect hairs, anterior stjl weak, lower one rather strong, posterior one very

strong.

Abdomen moderately slender, slightly narrowed to apex, somewhat depressed

at base, strongly compressed at apex, black, densely grey dusted, second to

fourth segments without or with a faint, narrow, brown median vitta, first to

fourth each with a pair of dark brown spots; second segment with a pair of

strong lateral setae and weak sublateral setae, third and fourth each with two

Fics. 68-71. Third segment and arista of left antennae of Cavicea, exterior view.

68.—cubitalis sp. n. 69.—hippelates sp. n. 70.—aristalis sp. n. 71.—pachypoda

Big.

pairs of strong discal setae, fourth also with a pair of rather strong lateral

marginals, fifth with a pair of rather long setae and some hairs; fourth ventrite

entirely grey dusted, inner margin of its lobes densely beset with fine, erect

hairs, which are surrounded outwardly by some long, fine, erect hairs.

Legs moderately slender, coxae black, densely grey-dusted, anterior tro-

chanters ferruginous, hind trochanters largely blackish, femora black, greyish

dusted, fore femora very narrowly ferruginous at apex, tibiae pale ferruginous,

very slightly infuscate near middle, rather conspicuously pale grey dusted,

tarsi piceous. Front femora without pd setae or with only 1—2 stronger ones

near middle and a few very small ones beyond them, 1-2 strong a setae near

base and often a row of small ones continuing them, and seven strong fv,

which alternate with fine ones, front tibiae with a long and strong fv at three-

fifths; mid-femora with 6-8 rather strong a setae, 2-3 strong and rather

numerous small a setae, four strong pv, alternating with small (sometimes

MUSCIDAE: B.—COENOSIINAE 183

partly rather strong) ones, and two strong preapicals (pP and fd), mid-tibiae

with a strong fd almost at three-fifths and a strong ad at two-thirds; hind

femora with five strong and sometimes 2—3 weaker ad setae, two strong av at

one-third and beyond two-thirds, and smaller ones of unequal length, four

moderately strong, not very long pv setae and a number of smaller ones, and a

strong pd preapical, hind tibiae with a very strong and long ad somewhat

beyond middle, a shorter though quite strong a seta slightly beyond it, and a

row of fine setulae along more than apical half of pd surface.

Wings hyaline, veins brown, somewhat paler at base, 74, and m distinctly

diverging at apex, vm distinctly beyond middle of discal cell, m,,. and particu-

larly m,-+-cu, strongly thickened and fuscous from the sub-basal transverse

veins to almost level of the apex of 7,. Calyptrae pale yellowish with testaceous

borders, the lower one much longer than the upper one, strongly projecting,

subparallel to beyond middle, broadly rounded at apex. Halteres pale

ferruginous.

? similar to male, the abdomen broader, more oval, pointed at apex, spots

of the fourth segment not very distinct, all the wing-veins normal near base.

Kenya: Aberdare Range, Mt. Kinangop, x.-1.xi.1934 (F.W.E.) 8000 ft.

6 type, I g paratype; gooo ft. 1 2 paratype; 12,000 ft. r g I 2 paratype;

Kinangop, vi.1930 (Turner) I 3 3 2 paratypes. Mt. Elgon, heath zone, 10,500-

11,500 ft., 11.1935 (F.W.E.) 1 g; alpine zone, 12,000-13,000 ft., r g. In the

latter two males, which are otherwise identical, the veins defining the discal

cell are only slightly widened at base. The male from Mt. Kinangop, 12,000 ft.,

has the setae considerably stronger than the other specimens.

Caricea hippelates sp. n. ¢.

Length 5:3 mm.; of wings 5-5 mm.

Head black, grey pollinose, interfrontalia dull black with brown reflections,

dorsal median part of occiput fuscous dusted, lower part of jowls dark brown.

Frons slightly wider than one eye, parallel, longer than wide, slightly shorter

than the face, fronto-facial stripe slightly narrowed to base of arista, con-

siderably widened with almost straight sides from there to lower margin of

eyes; parafrontalia about a fifth as wide as interfrontalia behind base of

antennae, almost half as wide as interfrontalia behind middle, with three strong

and three small pairs of inclinate frontal setae. Inner vertical very strong,

outer vertical rather strong, postverticals strong, ocellars very strong, as long

as frons. Antennae black, second joint silver-grey dusted interiorly, third

joint more than thrice as long as wide, apex separated from mouth-margin by

more than half the length of the joint, reaching lower sixth or seventh of eyes,

the latter not reaching level of mouth-margin; arista rather long-haired

dorsally in more than basal half, the longest hairs slightly longer than the third

184 RUWENZORI- EXPEDITION

antennal joint wide (Fig. 69). Palpi black. Parafacialia two-thirds as wide

as the third antennal joint, jowls with rather numerous, long and strong

peristomal setae, distinctly wider than third antennal joint.

Thorax black, densely grey dusted, the dorsum with a pair of broad, dark

brown vittae almost from anterior end to beyond lateral setae of scutellum and

from just inward of de to just outward of 7a, and a narrow brown median

vitta on presutural part, pleurae with a not very sharply defined dark brown

vitta from propleurae to supraspiracular convexity. acy fine but not very

short, in two rows with a few additional setulae on median line of presutural

72.

Fic. 72.—Caricea hippelates sp. n. Apex of left hind tibia and metatarsus,

anterior view.

Tic. 73.—Left hind tarsus of Cavicea kilembana sp. n., anterior view.

part and outward of the two rows in front of scutellum, 1 + 3 very long and

strong dc, a moderate ia; scutellum with sparse, erect, setulose hairs and two

pairs of strong setae. Two propleurals and prostigmatals, the lower one fine

but rather long, the upper one rather strong and long, prostigmatals surrounded

by some rather long hairs. stf/ in an equilateral triangle which bears rather

numerous, long, erect hairs, anterior stfl rather weak, the others strong, and

an additional strong seta between them.

Abdomen rather short, oblong-oval, rather strongly compressed at apex,

black, densely grey dusted with a narrow brown median vitta on segments

2-4 and a pair of rather large spots on segments 1-4, lateral seta strong, third

segment with a pair of subdorsal setae near hind margin, fourth with a discal

MUSCIDAE: B.—COENOSIINAE 185

transverse row of two pairs of setae, marginal setae indistinct among a row of

long and strong hairs, fifth segment with a pair of setae and a pair of hairs

between them, ventrites long-haired.

Legs very long, but not very slender, coxae black, trochanters pale ferru-

ginous, femora piceous, dusted, tibiae and tarsi ferruginous, front tibiae

somewhat infuscate in middle, last tarsal joint fuscous. Front femora with

numerous long and strong hairs on posterior surface, without a distinct pd

row, with about a dozen fv setae of very different size, about four of them very

long and strong, front tibiae with a long and strong fv seta slightly beyond

middle; mid-femora without a distinct ad row, but with about six moderate

a setae, four long fv in basal half, and two strong preapicals (pf and fd), mid-

tibiae with a long and strong fd slightly beyond middle, but without an ad

(in male at least); hind femora without strong setae, the fd preapical rather

inconspicuous among the coarse hair, hind tibiae without submedian setae but

with a large ad (setiferous?) pore at basal third, ventral (interior) spur very

long and strong, half as long as the tibiae, evenly though not very strongly

curved (Fig. 72), mid and hind tibiae densely haired on ventral surface,

with not very dense, long hairs on ad, d and pd surfaces, tarsi, especially hind

ones, with fine, long, curved hairs among the normal hairs, which themselves

are rather long and strong, the curled hairs on av surface of hind tarsi partly

as long as the femoral diameter.

Wings hardly smoky, strongly iridescent, veins brown, rather strongly

thickened at basal half of discal cell, 74, and m distinctly diverging at apex,

y—-m just beyond middle of discal cell. Calyptrae whitish with pale yellowish

border, the lower one strongly projecting, subparallel with broadly rounded

apex. Halteres orange-yellowish.

2 unknown, probably with the normal two setae on hind tibiae.

UGANDA: Ruwenzori, Bwamba Pass (west side), 5500-7500 ft. (7 W.E.)

3 type.

Caricea pachypoda Big.

Coenosia pachyoida Séguy, 1937, Gen. Ins. 205: 213 (errore).

n. syn. Coenosta longiseta Stein, 1907, Berlin. Ent. Zschr. 51: 78.

UcanDA: Ruwenzori: Kilembe, 4500 ft. (F.W.E.) 5 g¢ 4 9; Kyarumba,

4500 ft. (D. R. Buxton) 1 6 4 9; Mbarara, 15.xi.34 (F.W.E.) I 3.

Typical localities: Assinie; of longiseta: Toco: Bismarckburg. In B.M.

from: UGAnba: Mbale—Kumi Road, 3700 ft., S. of L. Salisbury; Hoima—

Kampala Road, 2.1.12 (R. E. McConnell); Supan: W. Darfur, Jebel Murra,

Killing, 7000 ft., 7.iv.1932, and E. Jebel Murra, Kirima, 5525 ft., 20.v.1932

(Miss M. Steele); TANGANYIKA: Tanga, vi.1932 (J. Ogilvie); ZANZIBAR;

NYASALAND; NATAL; S. NIGERIA; GOLD CoAsT; SIERRA LEONE.

Very reluctantly I sink the excellent and well-known name /Jongiseta as a

186 RUWENZORI EXPEDITION

synonym of pachypoda. The latter was very briefly described in 1891. The

long hairs of the arista (Fig. 71) were not mentioned by Bigot, but the “ palpes

fauves”’ might have suggested the present species, if the rareness of this

character in Caricea had been appreciated. Through the courtesy of Mr. J. E.

Collin I have been able to study the type of pachypoda.

Caricea strigipes group

1(10). Third joint of antennae pale yellow, not very long.

2(3). Vibrissae, anterior inclinate frontal setae, and palpi pale yellow. Scutellum

with only the apical pair of setae. Lower calyptra short, not projecting.

No acr. Body largely reddish-yellow to brownish-yellow. Legs en-

tirely yellow, setae of legs rather short, the hind tibiae of the male

(probably without an a bristle. g 3 mm. : : [*flavivibrissata Stein]

3(2). Vibrissae and inclinate frontal setae black. Scutellum with the usual two

pairs of setae. Body mainly blackish, grey dusted. Lower calyptra long,

strongly projecting beyond upper one.

4(7). Hind ocelli distinctly anterior to level of inner vertical setae. Parafrontalia

distinctly pale dusted, interfrontalia blackish. Three distinct pairs of

inclinate frontals, though the second one is small. Tarsi yellow.

5(6). Frons narrowed from first inclinate seta to vertex, hardly three-quarters as

wide at hind ocelli as at first inclinate seta. Mid and hind femora with

a preapical brown ring, of about a sixth the length of the femora, the

apical sixth yellow. Legs very long and thin. Humeri and pleurae

largely reddish-yellow. Abdomen reddish-yellowish translucent at

base and apex . : : . praeapicalis sp. n., p. 188

6(5). Frons very slightly madened to third molars seta, parallel from there to

vertex, slightly wider at hind ocelli than at first inclinate seta. Mid

and hind femora with a brown apical ring of almost a third the length

of the femora. Legs not very long. Humeri, pleurae, and abdomen

entirely blackish, grey and brown dusted 3 [attenuata Stein], p. 189

7(4). Hind ocelli level with inner vertical setae or slightly posterior. Parafrontalia

indistinct, as the whole frons is uniformly silver dusted in male (prob-

ably uniformly dark dusted in female). Two pairs of long and very

thin inclinate frontals in male. Hind metatarsus (3) twice as wide as

that of the mid-tarsi. (In these silver-fronted males the ocellar bristles

are present; see note, p. 222.)

8(9). All the femora with the apical quarter to third piceous. Hind tarsi piceous.

Dorsum of thorax and abdomen densely grey dusted, the former with

three brown vittae, the latter with a median vitta and a pair of spots on

segments 1-4, which are brown dusted and are more or less fused.

gilvicornis sp. n., p. 189

9(8). Front femora without any apical patch though with a narrow suffused

fuscous stripe on entire dorsal surface, mid and hind femora with a

very narrow apical fuscous ring which extends into a similar dorsal

stripe. All the tarsi yellow. Dorsum of thorax and abdomen shining

black, without conspicuous dusting : 2 nitida sp. N., p. 19

ro(1). Third joint of antennae black, though grey or ital grey dusted.

MUSCIDAE: B.—COENOSIINAE 187

11(12). Front femora black, the mid and hind femora black on the apical fourth or

more, tibiae more or less infuscate. Irons wholly silvery white pollinose

inmale . . i ; é . , , : [tmunvor Curr. |

12(11). Front femora yellow or with a fuscous dorsal longitudinal stripe. Inter-

frontalia dark dusted and parafrontalia pale dusted in both sexes.

13(16). Legs very long and slender, hind tibiae of male without an a seta. Either

the apical third of the posterior femora piceous, or the scutellum with

only the apical pair of setae and the lower calyptra very small. Slender,

somewhat larger species. 3g: 3:0-4:5 mm., abdomen elongate, com-

pressed at apex, third and fourth ventrites densely beset with long,

erect hairs.

14(15). Scutellum with only the apical pair of setae. Lower calyptra small and

narrow, narrowed from base to apex, projecting beyond upper one by

hardly the length of the latter, narrowly rounded at apex. [emora

wholly yellow. Abdomen with a pair of inconspicuous transverse

darker spots along hind margin of segments 1-4, without a median line.

gilvicoxa sp. n., p. 192

15(14). Scutellum with the normal two pairs of setae. Lower calyptra normal,

subparallel in basal half, projecting beyond upper one by more than

twice the length of the latter; rather broadly rounded at apex. Front

femora with a piceous dorsal streak on apical half, posterior femora with

the apical third piceous. Abdomen with a pair of rather faint oblong

spots and a complete median vitta on segments 1-4 : [venta Mall.]

16(13) Legs normal, hind tibiae of male witha quite strong aseta. Posterior femora

with at most a small brown point at the articulation of the tibiae.

Scutellum with the normal two pairs of setae. Lower calyptra normal,

subparallel in middle, rather broadly rounded at apex and projecting be-

yond upper one by approximately twice the length of the latter. Smaller

species, 5 2°5-3'5 mm., 2 3-4 mm. Abdomen of male short, depressed

or slightly compressed at apex, the last ventrites short-haired.

17(18). Arista short-plumose, distinctly wider, including plumosity, than the third

antennal joint, the longest hairs almost as long as the latter is wide.

Mid and hind femora of both sexes, besides setae, with short half-

appressed hairs, which are shorter than the tibial diameter. Hypo-

pygium of male more protruding. Thorax and abdomen (usually) less

dusted and more shining, the former with the median vitta less distinct.

punctigera Stein, p. 193

18(17). Arista pubescent, distinctly narrower, including pubescence, than the third

antennal joint, the longest hairs not more than half as long as the latter

wide. Hypopygium of male not much protruding. Thorax and ab-

domen densely grey dusted and dull.

19(20). Arista conspicuously pubescent, with the longest hairs about one-third to

one-half as long as the third antennal joint wide. Ventral and posterior

surfaces of mid and hind femora of male with numerous fine, erect,

soft hairs, which are almost one-and-a-half times as long as the tibial

diameter . A ° ° ° 5 . : : [stvigtpes Stein]

20(19). Arista very short pubescent, the longest hairs about as long as the basal

diameter of the arista. Femora with short hairs in both sexes besides

setae, the hairs being not quite as long as the tibial diameter.

[exigua Stein], p. 194

Il, 4 (g)

188 RUWENZORI EXPEDITION

Caricea praeapicalis sp. n. 3

Length 4-2 mm.; of wings 4°5 mm.

Head black, densely grey dusted, parafacialia and facial ridges silver-white

dusted, interfrontalia dull black with strong brownish reflections. Frons

almost twice as long as wide in front, shorter than face, strongly narrowed

behind with straight sides, four-fifths as wide in front as one eye, fronto-facial

stripe somewhat narrowed to slightly beyond base of arista, rather strongly

dilated and rounded to lower margin of eyes, where it is one and a half times as

wide as frons at widest point; parafrontalia a sixth as wide in front, a third as

wide behind middle as interfrontalia, with three inclinate frontal setae. Inner

vertical strong, outer vertical weak, postocellars and ocellars minute. First

and second antennal joints piceous, apex of the second joint testaceous-trans-

lucent, pale greyish dusted interiorly, third joint testaceous yellow, more than

thrice as long as wide, apex reaching lower third of eyes but as distant from

mouth-margin as from base of arista; the latter short-plumose, the longest hairs

of dorsal surface as long as the antennal joint wide, the longest hairs of ventral

surface just over half as long. Eyes reaching level of mouth-margin. Palpi

light-brown. Jowls not quite as wide as third antennal joint, with one strong

peristomal seta besides the vibrissae and the adjacent setae.

Thorax testaceous with piceous dorsum, the mesopleura, pteropleura and

sternopleura partly infuscated, dorsum greyish-brown dusted, with a rather

narrow dark-brown median vitta. acy almost absent, 1 + 3 moderate dc, one

za; scutellum almost devoid of setulose hairs, but with the normal two pairs

of strong setae. stfl in an equilateral triangle, which is almost devoid of small

hairs, anterior stpl weak, lower one moderate, posterior one strong.

Abdomen oblong-oval, slightly depressed, first abdominal segment testaceous-

translucent, the bases of the next two segments indistinctly and narrowly

so, sides of the first segment with a large transverse piceous spot along hind

margin, the latter, however, remaining yellow, rest of the abdomen grey,

grey dusted, second and third segments with a narrow median vitta and a pair

of large, transverse, dark-brown patches along hind margin, fourth with a pair

of round, brown spots, fifth segment and hypopygium testaceous, lateral setae

rather strong, fourth segment with a transverse discal row of two pairs of setae,

fifth with a pair of setae in middle.

Legs testaceous, mid and hind femora with a brown preapical ring, front

femora with a slight preapical shadow on anterior surface. Front femora with

three long pv setae, the other setae very small or lacking, front tibiae with a

rather strong pu seta; mid-femora with the av and ad setae very small or lacking,

two moderate pv in basal half, two small ones near two-thirds, and a rather small

p preapical; mid-tibiae with a moderate fd just beyond middle and an ad at

three-fifths, terminal spur long though not strong, slightly curved, somewhat

MUSCIDAE: B.—COENOSIINAE 189

longer than that of the hind tibiae; hind femora with six short ad setae, four

short av, two rather strong and two small fv, the former in the basal third,

the latter in the second third, and a fd preapical, hind tibiae with a strong ad

seta in middle and a rather small a seta at the same level.

Wings slightly smoky, veins brown, 7,,; and m diverging up to apex of

7,3, parallel at apex, y-m beyond middle of discal cell. Calyptrae hyaline with

yellowish-white border, lower calyptra normal, strongly projecting beyond

upper one, subparallel in middle, apex rounded. Halteres testaceous-yellow.

UGANDA: Budongo Forest, 7.-8.11.1935 (F.W.E.) 3 type.

[Caricea attenuata Stein?]

longitaysis Mall., 1922, Ann. Mag. nat. Hist. (9)10: 579, 587; Curr. 1935, Amer. Mus.

Novit. 776: 2, 7 (non Stein).

Typical localities: EGypr: Alexandria and Cairo; of longitarsis Mall. non

Stein. NATAL: Port Natal, Plant leg., in B.M.

The female specimen identified by Malloch as Caricea longiiarsis (Stein) is

in the B.M. Coenosta longitarsis Stein is, however, according to Stein, a synonym

of Spanochaeta dorsalis v. Ros, and there is no reason for querying this synonymy,

so much the less as Stein described both sexes and mentioned, among other

characters, the absence of a seta on the front tibiae, by which character Malloch

distinguishes dorsalis from longitaysis. I consider Malloch’s species to be

attenuata Stein, though the arista is short-plumose in basal half, not quite as

wide as the third antennal joint, whereas it is said to be bare by Stein. The

male of this species is said to have yellow antennae and yellow femora, while

the female should have dark antennae and dark femora. My experience with

the genus Caricea would not make this difference in the colour of the femora

between the sexes of attenuata very probable. On the other hand Stein did not

use the colour of the antennae in discriminating between stvigipes Stein (cim-

gulipes Stein, non Zett.) with dark antennae and attenuata in his key and

remarks of 1913 (Ann. Mus. nat. Hungar. 11: 566, 578). I am satisfied, after

all, that Stein mixed up two or three species in his descriptions of attenuata.

I should think that the females of the original series, described in 1903, belong

to humilis Meig., whereas the males described as attenuata in 1913 belong to

strigtpes or exigua. Stein himself admitted that one would meet with many

specimens, in which the attribution to either of the two species, stvigipes or

attenuata, would remain doubtful. It may then be assumed that the female

specimens of attenuata, mentioned in 1913, were actually humilis multima-

culata Ad.

Caricea gilvicornis sp. n. 3

Length 2-5 mm.; of wings 2-7 mm.

Head black, occiput grey dusted, fronto-facial stripe and jowls uniformly

190 RUWENZORI EXPEDITION

silver-white dusted. Frons parallel, but slightly narrowed anteriorly, fronto-

facial stripe slightly narrowed to middle of second antennal joint, slightly

dilated but almost straight to lower margin of eyes, where it is a quarter wider

than at vertex; parafrontalia hardly defined, with two fine but not short

inclinate frontal setae. Inner vertical rather strong, outer vertical indistinct,

postverticals and ocellars moderately long though fine. The two basal joints

of the antennae fuscous, the second silver-white dusted interiorly at apex, third

joint pale yellow, silver-white dusted, two and a half times as long as wide,

reaching lower fifth of eyes, its apex as distant from mouth-margin as from

base of arista; arista fuscous, very short-haired on basal half, the longest hairs

slightly longer than its basal diameter. Palpi black. Parafacialia slightly

more than half as wide as, jowls slightly narrower than, third antennal joint.

Thorax black, densely grey dusted, with three rather conspicuous brown

vittae on post part, the subdorsal ones extending to prst dc and the median one

just reaching hind end of frst part, scutellum with a not very conspicuous

brown patch on disc. acy fine, in two rows, rather sparse in front of suture;

1 + 3 strong dc, posterior 7a present, rather small; scutellum with two pairs

of fine small hairs and the normal two pairs of strong setae. Two moderately

fine propleurals and prostigmatals, the lower prostigmatal curved downward.

stpl in a triangle which bears a few fine hairs and the posterior side of which is

shorter, anterior s¢fl rather fine, lower one moderately strong, posterior one

strong.

Abdomen suboval, not longer than thorax, depressed at base, compressed at

apex, black, densely grey dusted, each segment with a large, transverse patch

of dark brown dusting, which reaches the hind margin in its entire width and

the front margin in middle, leaving a triangular patch of grey pollinosity on

either side of front margin. Hair appressed and fine, a few stronger, erect hairs

on sides, third ventrite with a pair of long, fine setae at apex.

Legs testaceous, mid and hind coxae dark grey with the apex reddish,

apical third of femora and the hind tarsi piceous, mid-tarsi strongly infuscate,

front tarsi ferruginous. Front femora with two d setae near apex, six rather

small pd (the basal two of them longer) and six moderately long and fine pv

setae, alternating with shorter ones, front tibiae with a long and strong sub-

median pv seta; mid-femora with three moderate ad setae between basal

quarter and two-thirds and two small ones in basal quarter, about six fine and

short av, four moderately long and fine pv setae between base and two-thirds,

and two preapicals (pf and fd), mid-tibiae with a moderate ad and a rather

small pd, the former almost at two-thirds, the latter a trifle more basad; hind

femora with six moderate ad, a long pv in middle, a moderately long one before

and beyond it and some small ones between them, one fd preapical and an

av preapical and two or three other rather small setae between basal and

apical thirds of av surface, hind tibiae with a strong ad and a seta in middle.

MUSCIDAE: B.—COENOSIINAE I9!t

Wings slightly smoky, strongly iridescent, veins brown, with somewhat paler

bases, 7,,, and m slightly converging at apex, y-m just beyond middle of discal

cell. Calyptrae hyaline, borders yellowish-white, the lower one strongly

projecting, broadly rounded at apex. Halteres pale yellow.

UGANDA: Ruwenzori, Namwamba Valley, 6500 ft. (’.W.E.) 3 type.

Caricea nitida sp. n. 3g

Length 2-6 mm.; of wings 2°7 mm.

Head black, occiput glossy black, except for a stripe of whitish-grey dust

between interior verticals and occipital foramen and the temples which are

silver-white dusted, as are the entire fronto-facial stripe and the jowls. Frons

slightly wider in middle than one eye, subparallel, very slightly widened from

first to second frontal setae, longer than wide, fronto-facial stripe slightly

narrowed to apex of second antennal joint, slightly rounded and dilated from

there to lower margin of eyes, where it is about a quarter wider than at vertex;

parafrontalia not defined, with two fine but not short inclinate frontal setae.

Inner vertical strong, outer vertical indistinct, postverticals rather strong,

ocellars rather weak, very close to postverticals (as are the ocelli). First and

second antennal joints fuscous, the second translucent yellowish and densely

silver-dusted interiorly at apex, third pale yellow, silver-white dusted, not

quite thrice as long as wide, reaching lowest sixth of eyes, the latter reaching

level of mouth-margin, apex of third antennal joint falling short of mouth-

margin by almost half the length of the joint, arista fuscous with paler base,

short-haired in basal half of dorsal surface, the longest hairs a third as long as

the diameter of the third antennal joint. Palpi brown. Parafacialia and jowls

half as wide as the third antennal joint.

Thorax black, dorsum, including humeri, glossy, very thinly grey and brown

dusted, pleurae more densely grey dusted. acy very fine and sparse, in two

rows, 1-+3 dc, posterior za not very long, scutellum with two pairs of fine but

rather long hairs and two pairs of setae, which are very long. Two fine pro-

pleurals and prostigmatals, the lower prostigmatal curved downward. stfl

in an equilateral triangle which bears one or two fine but rather long hairs,

anterior and lower stf/ subequal and rather small, posterior one very long and

strong.

Abdomen black, dorsum glossy, thinly brown-dusted at base, devoid of

dust on posterior half of first and on second to fourth segments, fifth tergite

brownish-grey dusted, venter brown-dusted, hair of abdomen rather long,

half-appressed, each segment with a transverse discal row of two pairs of setae

which are not very strong, except for the subdorsal pair. Fourth ventrite

strongly convex, semiglobular, though deeply cleft along median line.

Legs testaceous, mid and hind coxae largely grey, front femora narrowly

192 RUWENZORI EXPEDITION

brownish along dorsal surface, mid-femora narrowly brownish on apical half of

dorsal surface, hind femora with the extreme apex narrowly piceous. Front

femora with 3-4 small fd setae and five moderate pv setae, front tibiae with a

rather long and strong p seta; mid-femora with three small and one moderate

a seta, the latter at three-fifths, av setae hair-like, somewhat longer only in

basal half, four rather long pv setae, alternating with small ones, and two rather

conspicuous preapical setae (p and fd), mid-tibiae with a rather long ad and a

somewhat smaller pd just beyond it, both shghtly beyond middle; hind femora

with five ad setae, the last two stronger, three moderate and several small av,

two rather strong fu and a small one near base, and a fd preapical, hind tibiae

with a strong ad and a seta very close together just basad of middle.

Wings greyish-hyaline, strongly iridescent, veins brown, 7,,; and m slightly

diverging at apex, 7-m distinctly before middle of discal cell. Calyptrae

hyaline-whitish, the borders yellowish-white, both rather small, the lower one,

however, strongly projecting, broadly rounded at apex.

UGANDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 3 type.

Caricea gilvicoxa sp. n. 3g

Length 3-9 mm.; of wings 4:4 mm.

Head black, densely grey dusted, the dorsal part of occiput and the para-

trontalia more yellowish-grey dusted, face and parafacialia pale yellowish-grey

dusted, lower part and jowls whitish dusted, interfrontalia dull black with

brownish and whitish reflections. Frons longer than wide, shorter than face,

rather strongly narrowed to vertex, distinctly wider in front than one eye,

fronto-facial stripe very slightly narrowed to middle of face, slightly dilated

and rounded from there to lower margin of eyes, where it 1s somewhat wider

than frons at widest point; parafrontalia hardly a fifth as wide behind lunula,

hardly a third as wide at reclinate seta as interfrontalia, with three inclinate

frontal setae. Inner vertical strong, outer vertical very weak, hardly longer

than the occipital setulae, postverticals rather weak, ocellars almost half as

long as frons. Antennae black, the second joint pale greyish dusted interiorly

at apex, third joint (broken in type) rather long and slender and reaching about

ventral fifth of eyes (as may be supposed from the length of the shallow im-

pressions on the face). Parafacialia moderately narrow, certainly less than

half as wide as the third antennal joint. Eyes not quite reaching level of

mouth-margin, two-thirds as long as high. Palpi piceous with paler base.

Jowls slightly more than twice as wide as parafacialia.

Thorax black, grey dusted, dorsum greyish-brown dusted with a distinct

brown median vitta, not reaching neck and narrowing behind suture, a pair of

incomplete and not very conspicuous though rather broader lateral vittae along

the prst and za setae and traces of a subdorsal pair of vittae along the dc. acr

MUSCIDAE: B.—-COENOSIINAE 193

small and almost in one irregular row, 1-+-3 moderately strong dc, a long and

strong za; scutellum with a few fine short hairs and the apical pair of setae

which is moderately strong, the lateral pair absent. Two small and fine pro-

pleurals and prostigmatals. s¢// in an equilateral triangle which bears only

one or two fine hairs, anterior stfl very weak, lower one moderate, posterior

one strong.

Abdomen of moderate length, compressed in more than apical half, thinly

grey dusted without a distinct pattern, the hind margins slightly darker except

near middle. Lateral setae of first segment strong, of second and third very

weak and inconspicuous, fourth segment with a marginal row of 2-3 pairs of

rather strong setae, fifth with a pair of distinct setae near middle and some

short hairs, third and fourth ventrites (first and second to a less degree) densely

beset with fine, long, erect hairs, fourth ventrite entirely pruinose, dark grey,

hypopygium largely ferruginous.

Legs very long and slender, wholly testaceous, only the tibiae towards apex

and the tarsi somewhat more ferruginous to brownish. Front femora with a

row of small pd setae, three rather small # setae near base and three rather fine

pv from base to slightly beyond middle, front tibiae with a very fine pv; mid-

femora with about five a setae, the last but one of which is inserted on middle

and is somewhat stronger, four pv, the last of which is inserted on two-thirds

and is rather strong, and a fd and f preapical (setae missing in type), mid-

tibiae with an ad seta at two-thirds and apparently a fd just beyond middle;

hind femora with six rather strong but not very iong ad, 3-4 rather fine av,

a pd preapical, and three moderately long and fine pv setae, alternating with

small ones, hind tibiae (of male) with a single submedian seta just beyond

middle of ad surface.

Wings slender, hyaline, somewhat smoky, veins pale brown, 74,; and m

parallel at apex, y-m just beyond middle of discal cell. Calyptrae yellowish-

hyaline, very small, the lower one slightly projecting, elongate-trapezoidal,

with rather narrowly rounded (though by no means pointed) apex. Halteres

ferruginous-yellow.

UGANDA: Kigezi District, Mt. Sabinio, 8000 ft. xi.1934 (F.W.E.) 3 type.

Caricea punctigera Stein

UGANDA: Ruwenzori, Kyarumba, 4500 ft. (D. R. Buxton) i 3,1 2; Kilembe,

4500 ft. (F.W.E.) 2 9.

Typical locality: UGANDA: Mujenje. In B.M. from Nata: Port Natal and

Pietermaritzburg, the latter specimen with prey.

I think that the present specimens are this species. Malloch identified

as punctigera a form which I believe to be stvigipes. The pubescence of

the arista is always quite distinct against a white background in these forms.

194 RUWENZORI EXPEDITION

In the Seychellean form described as exigua by Stein (typical specimens in

B.M.) it is about as long as the arista is thick at base, in specimens from Kenya

it has the same length or is even somewhat shorter, whereas in specimens from

Port Natal (identified as strigipes by Malloch) and Nyasaland (identified as

punctigera by Malloch) it is distinctly longer. The present specimens, however,

have the hairs of the arista still conspicuously longer. The ventral surface of

the hind femora bears long and fine, dense, erect hairs in the males from Nyasa-

land only in the material at hand, and this form must thus be regarded as

strigipes Stein. It would seem to me that the present form, because of its

longer pubescence of arista, simple mid and hind femora, and more protruding

male hypopygium, must be identified as punctigera Stein. The specimen from

Port Natal presents the same characters, but the pollinosity of thorax and

abdomen is as in stvigipes. The name exigua Stein would then be left for the

form with simple male hind femora and short pubescence of the arista.

[Caricea exigua Stein]

n. syn. Caricea flavipes Ad. 1905, Kansas Univ. Sci. Bull. 3: 206.

n. syn. Coenosia ethelia Curr. 1935, Amer. Mus. Novit. 776: 7.

Typical localities: SEYCHELLES: Mahé; of flavipes: S. RHODESIA: Salis-

bury; of ethelia: TRANSVAAL: Pretoria and S. Arrica: Barberton.—In B.M.

from Mahé and from Kenya: Ruiru, I.vi.1932 2 g, and 13.vii.1932 I 9, on

coffee (Le Pelley).

Stein thought his ex7zgwa might be a synonym of his strigipes (1913, Ann.

Mus. nat. Hungar. 11: 578), but the characters given in my key (p. 187) would

not allow them to be regarded as identical. On the other hand, Curran’s

description of ethelia does not contain any characters which would enable it

to be distinguished from the present species, and the information obtained

from him only reads: traces to strigipes, but the arista is not conspicuously

pubescent, the rays shorter than the thickness of the arista.

C. flavipes Ad. was regarded a synonym of strigipes Stein by Curran (1935,

Amer. Mus. Novit. 776: 10), but Dr. Curran informs me that the paratypes

before him run down to exigua in my key, and not to strigifes. The name

flavipes was preoccupied twice in Coenosia, but not in Caricea, in which Adams

described his species, and there is no reason for supposing that Williston’s

flavipes should be a Caricea, whereas Stein’s species is definitely a Coenosia.

So far the rules do not permit the retention of a secondary homonym though

there is a strong and justified trend for revising the paragraph concerned in this

direction.

Caricea aseta sp. n. J

Length 5-5 mm.; of wings 5°8 mm.

Head black, grey dusted, the dusting denser and pale yellowish-grey on

MUSCIDAE: B.—COENOSIINAE 195

frontal triangle, parafrontalia, parafacialia and jowls; interfrontalia dull

blackish, with brown and whitish reflections when seen from in front. Frons

subparallel, very slightly narrowed to vertex, four-fifths as wide as one eye;

fronto-facial stripe subparallel, very slightly narrowed from base of antennae

to beyond insertion of arista, slightly widened to lower margin of eyes, where

it is about a fifth wider than at vertex, parafrontalia a third as wide as inter-

frontalia, with two rather strong and 1-2 weak inclinate frontal setae. Inner

vertical strong, outer vertical rather weak, postverticals moderate, ocellars

long, more than half as long as frons, occipital setulae very few in number.

Antennae dark fuscous, dust of second and third joints with whitish reflections

in apical half, third joint thrice as long as wide, apex level with lower fifth of

eyes, the latter not reaching level of mouth-margin; arista very short-plumose

on its entire length, except close to base, three-quarters of the width of the

third antennal joint including plumosity. Palpi fuscous at apex, amber on

basal two-thirds. Parafacialia more than half, jowls one-and-a-half times as

wide as the third antennal joint. Vibrissae slightly above mouth-margin.

Thorax blackish, whitish-grey dusted, testaceous on propleurae, ptero-

pleurae, posterior part of sternopleurae, anterior margin of hypopleurae, the

metapleurae and the sclerite behind the hypopleurae (epimeron of metathorax),

dorsum with a not very conspicuous brown pattern: a fusiform median spot

in front of scutellum, a round median spot on disc of scutellum, a narrow,

incomplete, paramedian vitta between acy and dc and closer to the former, and

an indistinct broad brownish area along za, the paramedian vittae distinct

only from prst de to second post dc, but with a narrow vestigial extension to the

neck. acy extremely sparse, almost absent, though in two rows, no prsc, I+3

strong dc, one 7a much behind level of sa and even somewhat behind second

post dc, scutellum apparently bare, with only the apical pair of setae. Two

propleural setae, the anterior one moderately strong, one moderately strong

prostigmatal seta, the lower one replaced by one or two minute hairs. s¢/ in

a triangle, the posterior side of which is somewhat shorter, the anterior stp/

evidently weak.

Abdomen rather slender, translucent ferruginous at base, grey dusted, the

dorsum largely and rather strongly infuscate, except along the margins and a

median line which are rufous, hypopygium ferruginous; lateral setae distinct,

though not very strong, hairs short, fine and sparse.

Legs entirely testaceous with the hind tibiae ferruginous and the front

tarsi and the hind femora. Front femora with two pv and one # seta in basal

fourth, front tibiae without a median seta (mid-legs missing), hind femora with two

av setae in basal third and one on apical third, five rather short ad setae and one

pd preapical seta, hind tibiae with a submedian ad and a, the latter weaker, on

middle and slightly beyond the former.

Wings somewhat infuscate, the base narrowly hyaline, veins brown,

196

RUWENZORI EXPEDITION

ferruginous near base, m3-++-cw, strongly thickened and fuscous near base, 74,5

parallel at apex, 7-m at middle of discal cell. Calyptrae whitish, borders

yellowish, not very large, but the lower one strongly projecting, rather narrowly

rounded at apex. Halteres yellowish.

Kenya: Aberdare Range, Mt. Kinangop, 9000 ft., x.1934 (J. Ford) 3 type.

Caricea calopoda group

1(10). a bristle of the hind tibiae distinctly beyond the ad (Fig. 67) or absent (some

9(8).

males). Parafacialia half or more than half as wide as the third an-

tennal joint (Figs. 75, 76). Frons almost or fully as wide as one eye,

slightly narrowed to vertex. Coxae black or largely black, even usually

the front coxae.

Large species: 6-9 mm. Parafacialia more than half as wide as the third

antennal joint (Fig. 75). The upper and lower prostigmatal bristles

normally surrounded by some small hairs.

y—-m ‘“‘surrounded by a round, dark brown spot’’, m—m ‘bordered with

brown’’. Dark pattern of the thorax largely confluent in middie.

9 mm. ; : : : ; ; [toequaerta

Cross-veins not clouded. Middle-sized species: 6-7-5 mm. Dark pattern of

the thorax not confluent in middle.

Mesonotum with a very broad brown vitta on each side of disc, which is

carried over anterior Jateval angle of scutellum and which reaches or

passes the dc as well as the za laterally, and a fine median brown line

which is neither contiguous to the lateral vittae nor continued over the

disc of scutellum. Front femora entirely or almost entirely black.

Tibiae sometimes more or less infuscate. Arista short-plumose, the

Curr.]

plumosity shorter in apical half : : ; [inaequivitta Mall.}

. Mesonotum with a narrow, sometimes slightly disintegrated vitta along the

dc on each side, a similar vitta along the za, and a well developed median

vitta of about equal width, the latter continued over the disc of the

scutellum, whereas the lateral vittae do not or indistinctly reach the

lateral angles of scutellum. Front femora yellow with a black streak on

dorsal surface which does not reach apex. : [aequivitta sp. n.],

Medium-sized species, 4-6 mm. Parafacialia hardly more than half as wide

as the third antennal joint (Fig. 76).

Arista short-haired, not fully as wide, including the hairs, as the third an-

tennal joint. Usually no small hairs close to the two prostigmatal

bristles. Femora with a conspicuous brown or black preapical spot,

the front femora entirely dark on dorsal surface, except at apex. Thorax

with three or five more or less faint narrow longitudinal vittae, of which

often the median one only is distinct. Abdomen very short with an

incomplete median vitta and paired rounded spots on segments 2 and

3, fourth segment with traces of a similar pattern. cuthbertsoni Curr.,

Arista with hairs, some of which are as long as or longer than the third an-

tennal joint is wide (Fig. 70). The two prostigmatal bristles surrounded

by some small hairs. Femora entirely yellow, without any trace of a

dorsal apical or preapical patch (though with an elongate small brown

patch on av and pv surfaces of apex). A well-developed anterior post ia

present. Pattern much the same as in cuthbertsoni. — aristalis sp. n.,

p. 198

p. 201

10(1).

I1(12).

12(I1).

13(24).

14(17).

15(16).

16(I5).

17(14).

MUSCIDAE: B.—-COENOSIINAE

The a bristle of the hind tibiae basad of or level with the ad. Parafacialia

much less than half as wide as the third antennal joint. Frons much

narrower, at most three-fourths as wide at vertex as one eye.

All the coxae black, grey dusted. Fore stpl long and strong, not weaker than

the lower one. Third antennal joint much longer (almost thrice as long)

than distance from its apex to mouth-margin, pointed at dorso-apical

end, arista, including the plumosity, as wide as the third joint. a

bristle of hind tibiae level with the ad bristle. Thorax indistinctiy

vittate laterally. : : [natalia

At least the front coxae meal, or entirely allore: Fo ore stpl weak, much

smaller than hind and lower stpl. Third antennal joint not much longer

than distance from its apex to mouth-margin.

a bristle of hind tibiae level with or slightly basad of ad. Second abdominal

segment with a distinct lateral and third with a well-developed lateral

and sublateral bristle.

Arista more or less pubescent near base, much narrower, including pubes-

cence, than the third antennal joint. Normally two post 7a, the anterior

one level with sa. Prostigmatal bristles not surrounded by hairs

(seldom one or two minute hairs present in front of the bristles).

Epistoma distinctly though slightly rounded-produced between the

vibrissae. Femora with a distinct dark apical spot. Dark pattern of

thorax and abdomen rather extended though the three patches on the

abdominal segments are never fused; pattern sometimes indistinct.

atvoapicata Mall.,

Dark apical spot of the front femora confined to apical third or two-fifths.

a seta of mid-tibiae long in females and in Tanganyika-males, variable

in Nairobi-males. Dark pattern fainter and less extended, the greyish

dusted paramedian area of abdominal segments 2 and 3 somewhat

broader than the dark median vitta. atvoapicata atvoapicata Mall.,

Dark apical spot of front femora very long, almost reaching base. Mid-

femora with a dark apical streak that reaches middle. a seta of mid-

tibiae very small in male. Dark pattern more conspicuous and more

extended, the greyish-dusted paramedian area of abdominal segments

2 and 3 not broader than the dark median vitta, the dark patches more

triangular and reaching hind margin of segment usually all over their

width. Tibiae sometimes more or less infuscate.

atvoapicata stvigavia Curr., p.

Arista short-plumose, as wide usually, including plumosity, as the third

antennal joint. Usually one post ia. Prostigmatal bristles surrounded

normally by small hairs. If the plumosity of the arista is not as wide

as the third antennal joint (Janipes), these hairs are rather numerous

and long.

. Plumosity of arista confined to basal half, rather short, not as wide as third

antennal segment. Abdomen without a dark pattern, thorax without

a distinct pattern (at least in male).

Femora yellow with the apex rather narrowly blacixish. Ventral surfaces

of abdomen, femora and tibiae with dense, fine, long and erect hairs in

male. 7mm: . 3 lanipes sp. 0.

Front femora black, mid- Game black sit the Base broadly yellowish, hind

femora yellow on a little more than basal half. Ventral surfaces of

197

Mall.]

. 203

198 RUWENZORI EXPEDITION

abdomen, femora and tibiae without abnormal villosity. 5-6 mm.

[tburunga Curr.], p. 204

21(18). Plumosity of arista reaching apex, as wide on basal half as the third antennal

segment. Abdomen and thorax with a distinct dark pattern. Ventral

surfaces of abdomen and legs with the normal bristles and short hairs.

4°5-5°5 mm.

22(23). Femora with a piceous apical spot. Tarsi more or less piceous. Fourth

ventrite of male with a row of long, bristly hairs along its inner margins,

several of which are almost as long as the lateral lobes of the ventrite.

punctipes Thoms., p. 205

23(22). Femora without a distinct apical spot, the front femora without a trace of a

spot, the mid and hind ones with a small point at the articulation of the

tibia. Tarsi yellow. Fourth ventrite of male with one hair only on

each side which is about as long as the lateral lobes of the ventrite.

similis Stein, p. 205

24(13). a bristle of the hind tibiae well basad of ad (Fig. 66) (by only half the tibial

diameter in alba, but then the second and third abdominal segments

without distinct setae).

(28). Femora with at most a small brownish spot at dorsal apex.

7). Tarsi brownish yellow, hardly darker than the tibiae. Lateral bristles of

the scutellum weak or missing. Thorax witha distinct median vitta and

traces of lateral vittae. Abdomen of female very slender, with (not

very striking) hind marginal bands, setae on second and third segments

well developed. Prostigmatal bristles not surrounded by hairs.

longiventris sp. D., Pp. 205

27(26). Tarsi piceous. Lateral bristles of scutellum strong. Thorax and abdomen

without any markings, uniformly silver-grey dusted. Setae on second

and third abdominal segments very weak and inconspicuous. Prostig-

matal bristles surrounded by some small hairs. Wings whitish in basal

half, infuscate in apical half. ¢ é j : alba sp. n., p. 206

28(25). Front femora with a complete broad black grea stripe, mid and hind

femora with a large black apical patch, occupying not much less than a

half or a third of the length of the femora. Prostigmatal bristles sur-

rounded by some small hairs. Scutellum with four strong bristles.

Ventral surfaces of femora, tibiae and abdomen with very conspicuous,

long, dense, and fine hair in male (Fig. 66). Setae on second and third

abdominal segments rather weak, much weaker than on first and fourth

segments . : ‘ ; ; : ; : calopoda Bezzi, p. 208

[Caricea aequivitta sp. n. 9]

Length 6:8-7-5 mm.; of wings 6:8—-7-0 mm.

Head (Fig. 75) black, densely grey dusted, parafrontalia, parafacialia, face,

and jowls yellowish-grey, interfrontalia black, with brown reflections, the

frontal triangle golden-grey. Frons slightly longer from first inclinate seta to

posterior ocelli than wide anteriorly, considerably wider than one eye, shorter

than face, fronto-facial stripe not distinctly narrowed to apex of second

MUSCIDAE: B.—COENOSIINAE I99

antennal joint, widened and hardly rounded from there to the lower margin of

eyes, where it is almost one-half wider than frons; parafrontalia a fifth as wide

behind lunula as interfrontalia, with three strong inclinate setae. Inner

vertical very strong, outer vertical weak, postverticals and ocellars strong.

Antennae fuscous, apex of the second joint reddish translucent at apex, greyish

Fics. 74-76. Heads of Caricea, dorso-frontal view. 74.—humilis multimaculata

Ad. 75.—aequivitta sp. n. 76.—aristalis sp. n.

dusted interiorly at apex, third joint two-and-a-half times as long as wide,

falling short of mouth-margin almost by its own length (lower margin of eyes

falling short of mouth-margin by about width of third antennal joint); arista

short-plumose, hairs of the ventral surface as long as basal diameter of arista,

those of the dorsal surface two-thirds as long near base as third antennal joint

wide. Palpi fuscous with brown base. Parafacialia as wide, jowls twice as

wide as third antennal joint.

200 RUWENZORI EXPEDITION

Thorax black, densely grey dusted, dorsum brownish-grey dusted with five

narrow vittae along acr, de and 7a, median vitta broadly encroaching upon disc

of scutellum. acy rather numerous, small, but distinct, in two rows with single

additional setulae in front of suture and more numerous ones towards scutellum,

I+ 3 strong dc, posterior 7a well developed, anterior one hair-like; scutellum

setulose with two pairs of strong setae, the lateral pair slightly shorter. Pos-

terior propleural and upper prostigmatal setae rather strong, lower ones moderate,

the lower prostigmatal curved downward, surrounded by some small hairs.

stpl in an almost equilateral triangle, which bears some erect hairs, anterior

stpl rather weak, lower one rather strong, posterior one very strong.

Abdomen short-oval, only slightly longer than thorax, convex, pointed,

densely grey dusted, with a narrow brown median vitta and a pair of not very

large roundish brown spots on each segment, second segment with a rather

distinct lateral seta, third segment with a lateral and sublateral pair, fourth

with a transverse row of 2-3 pairs of rather fine setae.

Legs pale ferruginous, coxae black, front coxae with the apex more or less

broadly pale ferruginous, front femora with a rather broad fuscous streak on

dorsal surface from base to two-thirds or three-quarters, mid-femora infuscate

along almost the entire dorsal surface, hind femora with a dark spot on dorsal

surface before apex, the anterior and posterior surfaces with a short preapical

streak near ventral surface, fifth tarsal joint fuscous. Front femora with 2-3

small pd near middle and a rather strong ad preapical, two rather long # setae

near base, which are continued to apex by a row of smallsetae, six long and strong

pv setae which alternate with finer setae, the last (preapical) one conspicuously

stronger though slightly shorter, front tibiae with a strong pv at three-fifths ; mid-

femora with 4—5 rather short but strong a setae from base to two-thirds, about

five short av in basal half, four strong pv in basal three-fifths, and two strong

preapicals (f and pd), mid-tibiae with a very strong fd at three-fifths and a strong

ad at almost three-quarters; hind femora with seven strong ad, 6-7 av, of

which two are long and strong (at about one-third and two-thirds), 2—3 fine but

rather long pu near base, a very long one at a third and a shorter but stronger

“one at two-thirds, and a strong fd preapical, hind tibiae with a very strong

and long ad and a somewhat shorter a seta somewhat beyond it (by up to half

the tibial diameter).

Wings sightly smoky, with brown veins, 7,,; and m parallel at apex, v-m

at middle to distinctly beyond middle of discal cell. Calyptrae hyaline with

yellowish borders and yellowish-white fringe, lower one rather long, subparallel,

rather broadly rounded at apex, strongly projecting beyond upper one. Halteres

ferruginous.

S. RHopeEsta: Vumba Mts., iii.1938 (A. Cuthbertson) 2 type; 11.1938 (A.

Cuthbertson) 1 2 paratype; 1.11.1935 (Major Drysdale) 1 2 paratype (in. coll.

A. Cuthbertson).

MUSCIDAE: B.—COENOSIINAE 201

Caricea cuthbertsoni Curr.

Coenosia cuthbertsoni Curr. 1935, Amer. Mus. Novit. 776: 2, 7.

UGANDA: Ruwenzori, Nyamgasani Valley, 6400 ft. (D. R. Buxton) 1 9.

Typical localities: S. RuoprEstaA: Salisbury; NatraL: Durban—Seen from

Salisbury; in B.M. from Kenya: Ruiru, 19.vili.1932 (H.C. James).

Caricea aristalis sp. n. 3

Length 4:7-5:8 mm.; of wings 4°8-5°5 mm.

3. Head (Fig. 76) black, densely grey dusted, pollinosity of frons and

face more yellowish-grey, interfrontalia black but rather densely yellowish-

grey dusted. Frons longer than wide, almost as wide as one eye, narrowed

to vertex, fronto-facial stripe very slightly narrowed to apex of second antennal

joint, moderately widened with almost straight sides to lower margin of eyes,

where it is one-and-a-half times as wide as frons; parafrontalia not quite a

quarter as wide as the interfrontalia, with 2-3 inclinate setae and 1-2 minute

hairs. Inner vertical very strong, outer one almost indistinct, postverticals

moderate, ocellars strong. Antennae fuscous, second joint ferruginous-trans-

lucent at apex, densely golden-grey dusted interiorly at apex, third joint not

quite thrice as long as wide, reaching lower quarter of eyes (the lower margin

of which almost reaches level of mouth-margin), its apex as distant from mouth-

margin as from base of arista; the latter plumose to apex, the longest hairs as

long as or slightly longer than the diameter of the third antennal joint (Fig. 70).

Palpi fuscous, the base brown. Parafacialia approximately half as wide, jowls

at most as wide as third antennal joint.

Thorax black, densely grey dusted, dorsum yellowish-grey dusted, with

five indistinct, narrow, brownish vittae. acy rather numerous, in two rows with

some odd ones on median line in front of suture and, outward of the two rows,

in front of scutellum, 1-+3 strong dc, two well-developed za; scutellum setulose

with two pairs of strong setae. Posterior propleural and upper prostigmatal

moderately long, lower prostigmatal not very small, curved downward,

prostigmatals surrounded by a few small hairs. stfl in an almost equilateral

triangle, with some fine, erect hairs, anterior stf/ weak, lower one strong,

posterior one very strong.

Abdomen as long as thorax, oval, densely grey dusted, with a brown median

vitta on second to third segments, which reaches neither front nor hind margins,

segments 2—4 each with a pair of transverse, brown spots on disc which are

small and more roundish on fourth segment. Lateral setae rather strong on

the first to fourth segments, the third and fourth besides with a pair of sub-

dorsal setae, fourth, moreover, with a fine lateral marginal seta, fifth with a

pair of long hairs and some short ones, lobes of the fourth ventrite densely

fine-haired along inner margin with long, fine hairs outwardly.

202 RUWENZORI EXPEDITION

Legs wholly testaceous to pale ferruginous, except for the fifth joint of

tarsi which is brown, a small preapical oblong, brown spot on the av surface and

a still smaller one on the pv surface of all the femora, front coxae dark at base,

mid-coxae largely blackish, hind ones largely brown. Front femora with a

small pd preapical and 2-3 very small pd in and beyond middle, 2-3 rather

strong p setae near base, 5-6 rather long fv, front tibiae with a strong and

long pv at three-fifths; mid-femora with a few minute a setae near base and a

small one towards middle, two strong fv in basal half and a small one at three-

fifths, and a single p preapical, mid-tibiae with a moderate fd at three-fifths

and a strong ad at two-thirds; hind femora with a few a setae near base, an

ad and pd preapical (the former more basad), a few small # and fv setae near

base and a rather strong pv towards the basal third, hind tibiae only with a

long and strong ad in middle.

Wings slightly yellowish-smoky, veins iene brown, 74,; and m slightly

diverging at apex, v-m very distinctly beyond middle. Calyptrae yellowish-

hyaline, with the border pale yellow, lower calyptra much longer than the

upper one, strongly projecting, subparallel in middle, broadly rounded at apex.

Halteres testaceous.

9 similar to male, the interfrontalia much less grey-dusted, the abdomen

pointed, the spots of the fourth segment round and very small, the small pu

spot lacking on the front femora, these femora with a complete row of # setae,

the setae on all the femora stronger, the pu of the front femora alternating with

fine, short setae; mid-femora with three long fv setae in basal half, a rather long

one at three-fifths, a few small ones between the others, and the normal two

preapicals (p and fd); hind femora with a complete row of six rather strong

ad, the last of which is the ad preapical, and a strong av at one-third and two-

thirds, hind tibiae with a long and strong ad at middle and a somewhat shorter

a seta distinctly though not very far beyond it.

Ucanpa: Budongo Forest, 7.-8.11.1935 (F.W.E.) 3g type; Kampala,

8.ix.1g18 (C. C. Gowdey) 2 9 paratypes; Mubende, 14.iv.1931 (H. Hargreaves)

I Q paratype (returned to Mr. Hargreaves); Bwamba Forest, Semliki Valley,

2300-2800 ft., 3.-7.xi.1911 (S. A. Neave) 1 9 paratype.

This species would run down to similis Stein in Malloch’s key, though the

words “‘arista short-haired’? would not fit. According to Stein’s description the

arista of similis is “‘breviter pilosa” or “‘bri¢évement villeuse’’. His figure

shows the arista, including pubescence, to be hardly wider at base than the

third antennal joint, whereas in avistalis it is more than twice as wide as the

third joint.

Caricea atroapicata Mall.

UGANDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 2 9.

MUSCIDAE: B.—COENOSIINAE 203

Typical localities: TANGANYIKA: Kondoa Irangi and Ufiomi (both in B.M.).

In B.M. from Kenya: Nairobi, vil.1930 (Van Someren) and Ngong, v.1936

(Miss Steele).

Caricea atroapicata strigaria Curr.

Coenosia strigavia Curr. 1935, Amer. Mus. Novit. 776: 11.

Kenya: Aberdare Range, Thompson’s Falls, 7500 ft., x.1934 (F.W.E.)

2 6,1 9., Mt. Elgon, heath zone, 8500 ft., 11.1935 (F.W.E.) 1 6.

Typical locality: W. TANGANYIKA.—In B.M. from AByssinia. Seen from

Kenya: Chyulu Hills, 5500 ft., iv. and v.1938 (Coryndon Museum E xped.).

Dr. Curran informs me that the type of stvigaria runs down to the present

subspecies in my key, but that the abdominal markings are very poorly defined

and the tibiae are not at all darkened. As in ihe material at hand the tibiae

are only sometimes more or less infuscated, and as the type of pattern of

strigaria is the same as that of my material (see Curran’s description), the

identity is not in doubt. With regard to C. leopoldi Curr. (1929, Rev. Zool.

Bot. afr. 17: 247) from Burungu and Kivu in the Belgian Congo, Dr. Curran

reports that it ““comes out with sévigaria but the disc of the mesonotum is

greyish instead of brown’”’. The differences in coloration are evidently small,

and if the two forms should prove to be synonymous, they would have to take

the name leopoldi Curr.

Caricea lanipes sp. n. 3

Length 6-8 mm.; of wings 7-2 mm.

Head black, densely grey-dusted, interfrontalia dull black with brown

reflections. Frons almost twice as long as wide, parallel, not quite two-thirds

as wide as one eye, almost as long as face, fronto-facial stripe very slightly

narrowed to apex of second antennal joint, rather strongly dilated and very

slightly rounded to lower margin of eyes, where it is two-thirds wider than

above antennae; parafrontalia a third as wide in front, not quite half as wide

in middle as interfrontalia, with four (and a very small) inclinate frontal setae.

Inner vertical strong, outer vertical not distinct, postverticals and ocellars

moderate. Antennae fuscous, the second joint with yellowish-grey dust in-

teriorly, third joint almost thrice as long as wide, reaching lower fifth of eyes,

its apex distant from mouth-margin by almost its length; arista long, smaller

apical half bare, basal half short-plumose, not quite as wide, including plum-

osity, as third antennal joint. Palpi fuscous with paler base. Jowls slightly

wider than third antennal joint.

Thorax black, densely grey-dusted, dorsum, including humeri, yellowish-

grey dusted, without a pattern, except for two short anterior vittae and some

II, 4 (h)

204 RUWENZORI EXPEDITION

irregular small spots of thinner pollinosity. acy fine, in two rows with some

additional hairs behind, 1-++3 long and strong dc, posterior 7a rather strong

(anterior one distinct on one side); scutellum with a few single setulae and

two pairs of strong setae. Posterior propleural rather strong, anterior one

rather fine, prostigmatals fine, the lower one curved downward, surrounded

by rather numerous fine hairs. s¢tf/ in an almost equilateral triangle with

rather numerous long, erect hairs, the posterior side somewhat shorter, anterior

stpl fine, posterior and lower stp/ subequal and very strong.

Abdomen oblong-oval, hardly longer than thorax, rather depressed, uni-

formly and densely grey-dusted, without any pattern, fourth ventrite black

and shining. Lateral setae very strong on segments 2-3, fourth segment with

a transverse discal row of two pairs of strong setae, fifth with two pairs of

setae, venter with rather dense, erect, fine hairs, dorsal hairs appressed on disc,

erect near sides.

Legs pale ferruginous, mid and hind coxae partly grey, front femora with

a dorsal piceous streak over almost apical half, mid and hind femora with the

apical fifth or sixth piceous, tarsi piceous. Front femora with ten rather fine

pd setae and about nine long and fine fu setae, some of which are not very

distinct among the rather long, fine, erect hair that clothes the front femora,

front tibiae with a strong pv seta at two-fifths, v. surface densely covered with

‘fine appressed hairs, # surface with numerous long, fine hairs, ventrai surface

of mid and hind femora and tibiae very densely clothed with fine erect hairs

of moderate length, the hairs somewhat longer but less dense on the anterior

and posterior surfaces of the mid-legs and the anterior surface of the hind

legs, mid-femora with a few ad setae only near base, and five very long and

fine pu setae in basal half, # and fd preapicals long and strong, mid-tibiae with

a rather strong fd seta at middle and a fine ad seta at three-fifths; hind femora

with about eight moderate ad, three long pv in basal half, and two fd near

apex, hind tibiae with a very strong ad and a seta at middle, practically level

with each other.

Wings subhyaline, the apical half slightly infuscate, veins brown with

testaceous base, 7,,; and m distinctly converging to apex, v—-m slightly beyond

middle of discal cell. Calyptrae hyaline with whitish margin, the lower one

strongly projecting, twice as long as the upper one, subparallel (slightly dilated

from base to middle), rather broadly rounded at apex. Halteres pale yellow.

UGANDA: Ruwenzori, Namwamba Valley, 6500 ft. (F.W.E.) 3 type.

[Caricea burunga Curr.]

Coenosia burunga Curr. 1935, Amer. Mus. Novit. 776: 9.

Dr. Curran writes that this species runs in the key p. 196 to danipes (but does

not possess the long pile), if the a is only a little before the ad (it differs on the

MUSCIDAE: B.—-COENOSIINAE 205

two legs in the male but would fall here in the female); if the a is well before

the ad, it would be calopoda but differs at once in the colour of the legs.

Caricea punctipes Thoms.

UGANDA: Ruwenzori, Mobuku Valley, 4500 ft. (F.W.E.) 1 9; KENYA:

Mt. Elgon, Kapretwa, 6500 ft. (F.W.E.), 3 3.

Typical locality: Cape. In B.M. from UGaNnpDaé, 1918 (C. C. Gowdey);

Kampala, 8.ix.1918 and 17.x1.1915 (C. C. Gowdey).

Caricea similis Stein

Kenya: Mt. Elgon, Kapretwa, 6500 ft. (F.W.E:) 1 3.

Typical locality: TANGANYIKA: Kilimanjaro. In B.M. from ABYSSINIA.

Caricea longiventris sp. n. °

Length 5°5 mm.; of wings 5-I-5:2 mm.

Head black, densely grey dusted, interfrontalia black with brown reflections

in anterior view, parafrontalia pale-grey dusted exteriorly, brownish-dusted

interiorly. Frons very slightly narrowed to vertex with almost straight sides,

two-thirds to three-quarters as wide as one eye, fronto-facial stripe widest

above lunula, hardly narrowed to beyond the base of arista, rather strongly

widened and rounded to lower margin of eyes, where it is almost one-and-a-half

times as wide as at vertex; parafrontalia a quarter as wide as interfrontalia,

with two strong and one small inclinate frontal setae. Inner vertical very

strong, outer vertical weak, postverticals rather weak, ocellars small and weak.

Antennae black or fuscous, second joint with pale grey dust interiorly at apex,

third joint two-and-a-half times as long as wide, reaching lower quarter of eyes,

its apex as far from mouth-margin as from base; arista short-plumose through-

out, almost as wide, including plumosity, as third antennal joint. Palpi piceous,

the base somewhat paler. Parafacialia narrow, less than a third as wide, jowls

almost as wide anteriorly as the third antennal joint, vibrissae above mouth-

margin by the anterior width of the jowls.

Thorax black, brownish-grey dusted on dorsum, with five narrow, brown

vittae, the subdorsal ones (along the dc) interrupted, pleurae grey dusted. acy

fine, in two paramedian rows, with a few additional hairs before scutellum, no

prsc, 1+-3 strong dc, anterior post za hair-like, posterior one moderately strong,

scutellum sparsely setulose, apical pair of setae strong, lateral pair weak or

absent. Propleurals and prostigmatals fine, the lower prostigmatal curved

downward. stfl in an equilateral triangle, the area with a few hairs and a

rather strong seta (fourth s¢f/), anterior s¢f/ fine, lower one moderate, posterior

one very long and strong.

206 RUWENZORI EXPEDITION

Abdomen long and slender, fusiform, somewhat depressed and curved down-

ward, not very densely grey dusted, first to third segments with a dark brown

band at hind margin, which occupies the posterior third of these segments and is

sub-interrupted on median line by some faint, greyish pollinosity, first to third

segments with distinct lateral setae, the third besides with a finer pair of sub-

lateral setae, fourth segment with two pairs of fine discal and marginal setae,

forming two transverse rows.

Legs very slender, yellow with a brown subapical shadow on mid and hind

femora, mid and hind coxae blackish exteriorly near base. Front femora with

4-5 short but quite conspicuous d setae before and at middle, a pd row, of which

only 4-5 setae near base are distinct but not very long, a pu row of four very

long setae, alternating with short ones, two of the long setae near middle about

half as long as the femur, front tibiae with a very long pv; mid-femora with a

row of short av setae, two strong a setae near middle, a row of four very long

pv (two of them near middle, about half as long as femur), alternating with

short ones, and with a single strong pd preapical, mid-tibiae with a long and

strong ad at three-fifths and an even longer fd somewhat beyond middle;

hind femora with two strong av at a third and two-thirds, 4—5 ad in basal two-

fifths, one ad at two-thirds and a preapical ad, a preapical pd, and three very

long pv setae, hind tibiae with a strong ad seta at middle and a slightly smaller a

seta basad of the ad almost by the diameter of the tibia.

Wings hyaline, slightly smoky, veins brown, 7,,., and m slightly converging

at apex, the former gently curved, the latter straight, v-m in middle of discal

cell. Calyptrae hyaline with pale yellowish border, the lower one strongly

protruding, subparallel, rather narrow, halteres yellow.

UGANDA: Ruwenzori, Nyamgasani Valley, 8000-9000 ft. (D. R. Buxton) 9

type, I @ paratype.

Caricea alba sp. n. 3

Length 4:7-5°5 mm.; of wings 5:0-5-4 mm.

Head black, densely grey dusted, interfrontalia dull black, with whitish

reflections when seen from in front. Frons rounded in profile, slightly pro-

jecting beyond eyes at base of antennae, almost twice as long as wide, three-

quarters as wide in front as one eye, slightly narrowed to vertex, as long as face,

fronto-facial stripe slightly narrowed to apex of second antennal joint, gradually

widened with weakly rounded sides to lower margin of eyes, where it is one-and-

a-half times as wide as on frons; parafrontalia half as wide as interfrontalia,

slightly more than half as wide at second frontal setae, with two strong and

between them a moderate and a minute, or a minute, a moderate and another

minute frontal seta. Inner vertical strong, outer vertical weak, but rather

distinct, postverticals and ocellars not very strong. Antennae fuscous, second

MUSCIDAE: B.—COENOSIINAE 207

joint greyish dusted interiorly at apex, third joint somewhat grey dusted, two-

and-a-half to three times as long as wide, its apex as distant from mouth-margin

as from base of arista, reaching lower fifth of eyes (the latter not quite reaching

level of mouth-margin); arista short-haired in basal half, the longest hairs a

third to a half as long as the diameter of the third antennal joint. Palpi fuscous

with the base brown. Parafacialia less than half, jowls not quite so fully as

wide as the third antennal joint, a strong seta above the vibrissae.

Thorax black, densely and uniformly pale grey dusted. acy fine, in two

irregular rows, 1-+3 long and strong dc, the three fost dc increasing in size,

posterior 7a well developed; scutellum very finely and minutely setulose, with

two pairs of strong setae, the lateral pair slightly shorter. Posterior propleural

and upper prostigmatal not very strong, lower prostigmatal fine, curved

downward, surrounded by some small hairs. s¢f/in an almost equilateral triangle

which bears rather numerous long, erect hairs, and the posterior side of which

is slightly shorter, anterior stf/ rather weak, lower. and posterior stpl equal,

very long and strong.

Abdomen rather depressed at base, slightly compressed ventrally at apex,

black, densely and uniformly pale grey dusted. Second and third segments

without distinct setae, the fourth with a transverse discal row of 2-3 pairs of

strong setae and a long and fine lateral marginal seta, fifth segment with a pair

of long and fine decussate setae and a pair of smaller ones outward of them,

fourth ventrite densely short-haired along inner margins of lobes, with rather

numerous, long, erect hairs towards outer side.

Legs testaceous, coxae densely whitish dusted, femora with a suffused brown

to piceous patch at apex of dorsal surface which more or less encroaches upon

anterior and posterior surfaces on the posterior femora, tarsi piceous. Front

femora with 6-7 strong fd, about three very long p near base which are followed

by some others which rapidly decrease in length, three long fv in basal half,

alternating with fine ones, and much shorter ones in apical half, front tibiae with

a very long pu seta in middle; mid-femora with 4-5 very long a setae in basal

half, 3-5 long fv in basal three-fifths, a complete row of short and fine # setae,

and two preapicals (f and pd), mid-tibiae with a strong pd on middle and a very

fine but long ad at two-thirds; hind femora with six long ad, the first and last

(preapical) of these shorter, 4-6 fine and rather short av, 2—5 fine # in basal

two-fifths, five long fv in basal two-thirds, the second and last of them shorter,

hind tibiae with a long and fine a at and a very long and somewhat stronger ad

slightly beyond middle (by half the tibial diameter).

Wings whitish in basal half, somewhat smoky in apical half, the difference

very inconspicuous when seen on one wing under higher magnification, but

very evident, if both wings are seen with naked eye behind each other against a

white background, veins brown in apical half and turning gradually pale

testaceous near base, 7,,, and m slightly diverging at apex, v-m slightly beyond

208 RUWENZORI EXPEDITION

(or at) middle of discal cell. Calyptrae whitish-hyaline with whitish border

and fringe, rather large, the lower one strongly projecting, distinctly dilated

from base to middle, apex broadly rounded. Halteres whitish-yellow, the knob

pale reddish at apex.

UcanpDA: Ruwenzori, Mobuku Valley, 7300 ft. (F.W.E.) g type, 1 g para-

type; Kilembe, 4500 ft. (F.W.E.) 3 ¢ paratypes; Kampala, 3.1x.1918, 13.xi.

1917, 14.x1.1917 (C. C. Gowdey) 5 g paratypes; Nairobi, vii.1930 (Van Someren)

I $ paratype (returned to Coryndon Memorial Museum, Nairobi).

Caricea calopoda Bezzi

UGANDA: Ruwenzori, Mobuku Valley, 7300 ft. (F.W.E.) 1 g; Namwamba

Valley, 6500 ft. (F.W.E.) 1 g 1 @. Typical locality: Adi Caié (ERITREA).

Reported from KrENyA: Aberdares (Malloch); UGANDA: Behungi (Curran);

N.W. TANGANYIKA (Curran); BELG. Conco: Kibati (Curran). In B.M. from

KENYA: S. Masai-Reserve, 25.iv.13 (I. J. Anderson), 1 9.

Bezzi as well as Malloch knew females only, when they dealt with this

species. The arrangement of the bristles on the hind tibiae was not described

by Bezzi. Thus some reserve in the use of Bezzi’s name is warranted. The male

described by Curran (1935, Amer. Mus. Novit. 776: 11) isa very pale form; the

two males in the present collection have brown markings, though these are

much less conspicuous than in the female. The outstanding character of the

male is the fine and dense, long, erect hair on the ventral surfaces of the abdomen,

femora and tibiae, the femora being very long-haired all round. The male

form identified as calopoda by Stein (1913, Ann. Mag. nat. Hungar. 11: 578) can

hardly be this species ; 1t may be atroapicata Mall.

Caricea nigritibia sp. n. 3 2

Length 5-3-6-3 mm.; of wings 5:9-7 mm.

3. Head black, densely grey dusted, except on median dorsal part of occiput

which is fuscous, frons which is dark brown dusted (the interfrontalia being dull

black in a strictly dorsal view), face, parafacialia and jowls which are golden

dusted. rons subparallel, very slightly widened to vertex (especially in anterior

half), four-fifths as wide behind as one eye; fronto-facial stripe somewhat

narrowed to base of arista, rather strongly widened to lower margins of eyes,

where it is about a fifth wider than at vertex; parafrontalia one-third as wide

as interfrontalia, with 3(—4) inclinate frontal setae. Inner vertical strong, outer

vertical rather weak, postverticals and ocellars rather strong. Antennae dark

fuscous, apex of second joint golden pruinose on dorso-interior surface, third

joint thrice as long as wide, apex reaching lower fifth of eyes, the latter almost

MUSCIDAE: B.—COENOSIINAE 209

reaching level of mouth-margin; arista very short-pubescent on dorsal surface

near base, the longest hairs shorter than the width of the thickened basal part,

the apical half and ventral surface bare. Palpi black. Parafacialia almost half,

jowls one-and-a-half times as wide as the third antennal joint. Vibrissae slightly

above mouth-margin.

Thorax black, densely grey dusted, the dorsum, including scutellum but

excluding humeri, dark brown dusted. acy sparse and fine but not very short,

in two rows, no prsc, 1+3 strong dc, two strong 7a, the anterior one slightly

before level of sa, scutellum bare except for two or three pairs of setulose hairs

and two pairs of strong setae. Two propleural and two prostigmatal setae,

the dorsal ones strong, the ventral ones moderate, no small hairs round about.

stpl in a triangle, the posterior side of which is somewhat shorter, and the area

of which bears a number of rather long setulose hairs, anterior stpl slightly

weaker, posterior considerably stronger than lower stl.

Abdomen elongate-oval, grey dusted with a large transverse patch of dark

brown pruinosity on each segment, only a narrow front margin (interrupted on

median line) and narrow lateral margins remaining grey, dark patch of fourth

segment subtriangular, its apex reaching hind margin of third segment; second

segment with a pair of distinct lateral setae (and two pairs of smaller setae

rather close to them), third and fourth segments each with three pairs of discal

setae in a transverse row, fifth with a pair of setae and two pairs of setulose hairs.

Legs moderately slender, black, except on basal half of hind and basal third

of mid-femora, which are yellow, base of front tibiae dark ferruginous, front and

mid-trochanters ferruginous. Front femora with six long and fine pu and 5-6

rather short pd, front tibiae with a moderate pv, mid-femora with four strong a

and 3-4 pv in basal half, mid-tibiae with a moderate ad and pd, both somewhat

beyond middle, but the fd very close to it, hind femora with 5-6 moderate ad,

2-3 long av and pv and a pd preapical, hind tibiae with a strong ad seta in middle,

a absent.

Wings hyaline, the veins brown, rather broadly though slightly infuscate,

the infuscation taking up almost all the anterior half of the wing, as only a

harrow apical streak in R, and a narrow long streak in R; remain hyaline.

¥4,5 Slightly curved at apex but parallel, v-m slightly beyond middle of discal

cell. Calyptrae whitish-hyaline, the border yellowish, lower calyptra strongly

projecting. Halteres reddish-yellow.

2 similar to male, the abdomen somewhat broader and more pointed, the

fourth segment only with a median vitta which may be subtriangularly dilated

at apex. In one female the third segment has three separate spots and the

second a median vitta and a pair of very large spots which approach the median _

vitta very closely at some places. The a seta of the hind tibiae well developed,

practically level with the ad.

KENYA: Aberdare Range, Mt. Kinangop, 8000 ft., Cedar Forest, x.1934

210 RUWENZORI EXPEDITION

(F.W.E.) 3 type, 2 2 paratypes; gooo ft., 2 2 paratypes. North of Mt. Kenya,

8300 ft., 18.ii.rg11 (1. J. Anderson), paratype, abdomen missing.

The a bristle and its pore on the hind tibiae of the single male at hand are

lacking, as in the males of a few other species (vittata, some males of mmaequt-

vittata, etc.). The present species, which seems to connect the calopoda and

vebmanni groups, was identified as fumipennis Stein by Malloch (1922, Ann.

Mag. nat. Hist. (9)10: 583). As its hind tibiae, however, are entirely black

(except sometimes for a slightly paler basal zone), whereas they are said to be

“zum grodssten Teil blassgelb”’ by Stein, I refer the species from the Nam-

wamba Valley to vebmanni Speis. (= fumipennis Stein, non Lamb), so much the

more as the antennae are said to be rather slender and long, inserted above the

middle of the eyes. This would apply to the Namwamba species, but not to

the present form.

Caricea mgahingana sp. n. ¢

Length 4-3 mm.; of wings 4-8 mm.

3. Head black, densely grey dusted, median dorsal part of occiput fuscous,

frons dark velvety brown with light reflections, the interfrontalia black in a

strictly dorsal view, face (incl. parafacialia) with silver-white dust. Frons

parallel, slightly wider than one eye, not much longer than wide, much shorter

than face, fronto-facial stripe slightly narrowed to beyond base of arista, widened

to lower margin of eyes, where it is as wide as in middle of frons, parafrontalia

hardly a fifth of the width of the interfrontalia, with 3(-4) inclinate frontal setae.

Inner vertical strong, outer vertical rather weak, postverticals and ocellars

rather strong. Antennae fuscous with slight whitish reflections, third joint long

and broad, more than thrice as long as broad, passing lower margin of eyes

and almost reaching mouth-margin, arista very short-haired on greater middle

half of its dorsal surface, the longest hairs hardly longer than the basal width

of arista. Palpi piceous. Parafacialia very narrow, about a fifth as wide as

third antennal joint, jowls as wide as third antennal joint. Vibrissae level with

mouth-margin.

Thorax black, densely grey dusted, the dorsum, including scutellum but

excluding humeri, dark dusted. acy sparse but rather strong, in two rows, on

stronger prsc, 1-4-3 dc, two strong za, the anterior one much in front of the sa

and slightly behind level of first fost dc, scutellum sparsely setulose, with two

pairs of strong setae. Two propleural and two prostigmatal setae, the upper

ones rather strong, the lower ones rather small, one or two minute hairs close to

the prostigmatals. s¢f/ in an equilateral triangle, which bears one or two long,

erect hairs, the anterior and lower s¢f/ subequal, the posterior one much stronger.

Abdomen elongate oval, rather depressed, grey dusted with a median vitta

and a pair of small spots of dark brown dust on hind margins of second and

MUSCIDAE: B.—COENOSIINAE PPA E

third segments, fourth segment with the median vitta subtriangularly dilated

at apex; second segment with a pair of distinct lateral setae; third and fourth

each with three pairs of setae in a transverse discal row, fifth with two pairs

of setae and a few hairs.

Legs black, with the basal half of hind and basal two fifths of mid-femora

yellow, the tibiae piceous. Front femora with rather short and fine setae, front

tibiae with a fine submedian pv, mid-femora with about four moderate pv

setae in basal half, one beyond middle, and two preapical setae (p and fd),

mid-tibiae with a moderate ad and a small fd, the former much, the latter

slightly beyond middle, hind femora with about six moderate ad, four erect,

fine pu in basal half and one fd preapical, hind tibiae with a strong ad slightly

beyond middle and a weak a just beyond the ad.

Wings hyaline, veins brown, 7,,; gradually but very distinctly converging

at apex, 7-m just beyond middle of discal cell. Calyptrae whitish-hyaline, the

border yellowish-white. Halteres orange-yellow.

UGANDA: Kigezi District, Mt. Mgahinga, 10,000-11,000 ft., xi.1934 (F.W.E.)

3 type.

The species, though placed near nigrotibialis in the key, is perhaps not very

closely related to it. Unfortunately the specimen is somewhat teneral, and the

interfrontalia and jowls are somewhat shrunk.

Caricea rebmanni group

There are perhaps two good species only in the material at hand: tvanslucida

and vebmanni, all the others probably being subspecies of the latter. As I have,

however, seen only one specimen of tvanslucida, I should not like to make the

first bifurcation in the key for it, using the bare arista, short third antennal

joint, and translucent base of abdomen, which characters do not occur in the

other forms. Some of the specimens seen seem to be definitely intermediate

between attenuicornis and kilembana, others between kilembana and ciliventris.

1(10). Front coxae yellow on anterior surface, seldom slightly infuscate; but then

the hind tibiae not paler in middle.

2(7). Tibiae yellow, the mid and hind ones sometimes slightly infuscate. Femora

with not more than apical third piceous (except for dorsal streak which

may pass the apical third).

3(6). Mesonotum entirely dark dusted with the humeral and notopleural calli

grey dusted. Parafrontalia dark dusted like interfrontalia. Fourth

ventrite of male with fine black setulae and an inconspicuous fringe of

short yellow pile along emargination.

4(5). Arista practically bare, third antennal joint half as long as face, blunt at apex.

Hind femora entirely yellow except for a minute piceous point at articu-

lation of tibia. First and base of second abdominal segment somewhat

brownish-yellow translucent, the dark patches on the abdominal

212 RUWENZORI EXPEDITION

tergites not very distinct, the whole dorsum rather shining brown.

tvanslucida sp. N.., p.

5(4). Arista short-plumose except on the apical quarter and the thickened base,

third antennal joint much more than half as long as face, gradually

tapering and pointed in female, very long (four-fifths as long as face)

and less pointed in male (Fig. 77). Hind femora with a brown shadow

or a piceous patch over ae quarter. Abdomen not translucent at

base. : : é attenuicornis Mall., p. 214

6(3). Mesonotum grey anisted, with 3 OF 5 Narrow and partly disintegrated longi-

tudinal vittae (along the acy, dc and za). Parafrontalia distinctly pale

dusted. Fourth ventrite of male with a conspicuous fringe of dense

black setulae along, at least, basal half of emargination.

ciliventris sp. N., p.

7(2). Tibiae piceous, the hind ones sometimes slightly paler in middle. Femora

with at least the apical two-fifths entirely piceous or black.

8(9). Posterior femora with the basal half bright reddish-yellow. The three

basal joints of the hind tarsi strongly compressed, rather stout in side

view, the second and third joint each very distinctly narrower than the

preceding one (Fig. 73). F : : kilembana sp. n., p. 216

9(8). Posterior femora only reddish-yellow Geroicere in basal third or less. The

three basal joints of the hind tarsi moderately compressed, slender, of

almost equal width. . 3 : kinangopana sp. n., p. 218

ro(1).. Anterior surface of the front coxae dlmogt enurely occupied by a piceous,

dark brown dusted streak. Tibiae piceous, the hind ones largely pale

yellow, with base and apex piceous. Femora with a complete blackish

dorsal streak, the front ones practically entirely black, the mid and hind

femora almost entirely black in male with the base somewhat reddish-

yeliow translucent, largely black, with the base more distinctly reddish-

yellow in female. Mesonotum entirely dark dusted, including humeri

and scutellum. Lower calyptra sooty-brown in male. vebmanni Speis. p.

foal

Caricea translucida sp. n. 3

Length 3-3 mm.; of wings 3°7 mm.

Head black, grey dusted on the face and lower part of occiput, dark brown

dusted on the entire upper part of the occiput, frons dull black with whitish

reflections when seen from in front, parafrontalia not pale dusted, only a very

inconspicuous and narrow line of greyish pollinosity ascending along the inner

margin of eyes outward of the frontal setae. Frons subtriangularly dilated

to vertex, with straight sides, twice as wide behind as in front, almost thrice

as long as wide in front, not quite half as wide in front as one eye (measures

estimated); fronto-facial stripe narrowed to middle of third antennai joint,

where it is not much more than a third as wide as one eye, moderately widened

and rounded to lower margin of eyes, where it is as wide as at vertex; para-

frontalia not clearly defined, with two moderate and one minute inclinate

frontal seta. Inner vertical strong, outer one weak though distinct,

MUSCIDAE: .B. COENOSIINAE 213

post-verticals and ocellars moderate. Antennae fuscous, the second joint with

yellowish-grey dust and pale translucent interiorly at apex, third joint thrice

as long as wide, reaching lower third of eyes, its apex as distant from mouth-

margin as from base of arista; arista practically bare, with only a few very

short hairs, the 1-2 longest ones of them as long as the basal diameter of the

arista. Palpi fuscous with paler base. Parafacialia and jowls linear, eyes very

large, slightly passing level of mouth-margin.

Thorax black, densely dark-brown dusted on dorsum, grey dusted on

pleurae, humeri and lateral parts of mesonotum yellowish-grey dusted. acy

fine, in two distinct rows, 1-+-3 very long dc, posterior 7a rather small; scutellum

with a few small hairs on apical half and two pairs of strong setae. The two

propleurals rather small, upper prostigmatal seta rather strong, lower one fine,

curved downward, not surrounded by hairs. s¢fl/ in an almost equilateral

triangle, the posterior side of which is slightly shorter, and which bears some

erect, fine hairs, anterior stp/ rather weak, posterior stp/ very strong, lower one

rather strong.

Abdomen elongate-suboval, slightly longer than thorax, somewhat curved

downward, somewhat compressed at apex, first segment, base of second,

venter of the others, and the hypopygium testaceous translucent, but the

fourth sternite dull black, large and strongly convex, thinly yellowish-grey

dusted; lateral setae distinct, third and fourth segments with a transverse

discal row of four strong setae, apex of the fourth segment with a few long

and strong hairs, fifth with one pair of fine, long setae.

Legs slender, moderately long, testaceous, front and mid-femora with a

brown streak along apical two-thirds of dorsum, hind femora with the extreme

apex narrowly brown, mid-tibiae somewhat infuscated, the basal two joints

of the mid-tarsi piceous, the others whitish-yellow, the basal two joints of the

front and hind tarsi testaceous, the others pale yellow. Front femora with

about six fine and rather short fd setae, five rather long and fine pv setae,

alternating with smaller ones, pv seta of front tibiae more than two-thirds as

long as tibia; mid-femora with 2—3 fine ad setae near base, four long and strong

pv, alternating with small ones, two strong a setae in apical half, and two pre-

apicals (p and fd), mid-tibiae with a strong av and a somewhat smaller fd seta

in middle; hind femora with six ad setae, among them a strong one at two-

thirds, a strong av at two-thirds, a smaller one beyond it, a moderate fv near

base and two very long ones near middle, hind tibiae with a very long ad at

second fifth and a rather long a seta very close to it, the d preapical at three-

quarters.

Wings rather strongly infuscated, veins brown, 7,,; and m very slightly

diverging at apex, y-m beyond middle of discal cell. Calyptrae hyaline, slightly

smoky, the borders testaceous. Halteres ferruginous.

UGANDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 3 type.

214 RUWENZORI1 EXPEDITION

Caricea attenuicornis Mall.

UGANDA: Ruwenzori, Namwamba Valley, 6500 ft. (F.W.E.) 1 3; Bwamba

Pass (West Side), 5500-7500 ft. (F.W.E.) 1 g; Aberdare Range, Thompson’s

Falls, 7500 ft., x.1934 (F.W.E.) I 6.

Fics. 77-80. Heads of Caricea, dorso-frontal view. 77.—attenuicornis Mall. 78.

—movio sp. 0. 79.—niveifrons albisquama ssp. n. 80.—hyalinipennis sp. 0.

Typical locality: Kenya: S.E. edge of Kenya Forest (2 type in B.M.).—In

B.M. from UGANDA: Kampala, I.ix.1g. (C. C. Gowdey) 1 9.

According to Malloch’s key (1922, Ann. Mag. nat. Hist. (9)10: 578) this

species should possess a pd seta on the hind tibiae. This is not mentioned in

the description (l.c., p. 584), nor is it present in the type. The species has thus

been erroneously placed in that section of Malloch’s key. The specimen from

MUSCIDAE: B.—COENOSIINAE 205

Kampala, mentioned by Malloch (1925, Ann. Mag. nat. Hist. (g)16: 363) as

C. rebmanni, belongs to this species. The specimens at hand are certainly con-

specific though not quite identical, the males having the third antennal joint

still more slender and longer and less pointed at apex (Fig. 77), and the apices

of the mid and hind femora blackened, whereas the apices are only infuscate in

the two females. In the specimen from Thompson’s Falls the mid and hind

tibiae are more strongly infuscate.

Caricea ciliventris sp. n. 5 2

Length 3:2-4:2 mm.; of wings 3°5-4°6 mm.

3. Head black, interfrontalia dull black with brown reflections when seen

from in front, parafrontalia, neighbourhood of the ocelli and postverticals

golden-grey dusted, occiput golden-grey dusted above with a fuscous central

vitta which embraces the golden-grey spot around the ocelli like a fuscous

ring, lower part of occiput grey dusted, face whitish-grey dusted. Frons

straight in profile, projecting by almost the width of the antennae, shorter than

face, not much longer than wide at vertex, over three-quarters as wide at

anterior frontal setae as one eye, evenly and strongly dilated to vertex, fronto-

facial stripe rather strongly narrowed to middle or beyond middle of third

antennal joint, where it is apparently half (but actually two-thirds) as wide as

one eye, strongly widened and slightly rounded to lower margins of eyes, which

distinctly pass level of mouth-margin, and where the fronto-facial stripe is

almost twice as wide as at lunula and a sixth wider than at vertex; para-

frontalia not well defined, a third as wide as the interfrontalia at lunula, with

two moderate and one weak inclinate frontal seta. Inner vertical strong, outer

vertical indistinct, postverticals and ocellars strong. Antennae fuscous, second

joint with greyish-brown pollinosity interiorly at apex, third joint four to

almost five times as long as wide, apex somewhat pointed dorsally, reaching

lower quarter of eyes, falling short of mouth-margin by half the length of the

joint; arista short-haired on about basal half, two-thirds as wide, including

plumosity, as third antennal joint. Palpi fuscous or brown. Parafacialia and

jowls almost linear.

Thorax black, pleurae grey dusted, dorsum brownish-grey dusted with five

dark brown vittae along the acy, dc and za, the latter two pairs more or less

disintegrated. acy moderately fine, in two rows and mostly paired, a few addi-

tional ones towards scutellum, 1-+-3 strong dc, two well-developed ta; scutellum

setulose with two pairs of strong setae, the lateral ones slightly shorter. Pos-

terior propleural and upper prostigmatal subequal, not very strong, lower

prostigmatal fine, curved downward, 1-2 small hairs close to it. s¢f/ in an

almost equilateral triangle which bears some fine hairs, anterior and lower

stpl subequal, not very strong, posterior one very long and strong.

216 RUWENZORI EXPEDITION

Abdomen oblong, rather strongly compressed, not very densely grey dusted,

the first to third segments each with a dark brown transverse band along hind

margin, widening towards middle and there more or less reaching front margin,

fourth with a brown median vitta which is triangularly dilated behind; second

to fourth segments each with a transverse discal row of 2-3 pairs of strong

setae, hind margin of fourth segment with some fine, long setae, fifth with a pair

of fine, parallel setae between which a small pair is inserted, fourth ventrite with

a dense fringe of fine, black setulae along at least anterior half of its lateral lobes.

Legs testaceous, coxae brownish exteriorly at base, front temora infuscated

along dorsum and more or less at apex, mid and hind femora infuscated in

apical third, the infuscation extending towards middle or base on dorsal surface,

posterior tibiae more or less, anterior tarsi strongly infuscated, hind ones

piceous. Front femora with 5—7 ad setae in apical three-fifths, five long pv,

alternating with small ones, front tibiae with a very long pv seta at two-fifths,

mid-femora with about five rather fine a setae in basal half, 3-4 long and fine

pv setae from base to two-thirds, and two strong preapicals (f and fd), mid-

tibiae with a moderately strong pd at middle and a strong ad somewhat beyond

middle; hind femora with 7-10 strong ad, the last of them somewhat apart and

level with the rather strong fd preapical, about 5-7 fine though distinct av, a

very long and strong pv at a third, and two smaller ones near base and at

middle, hind tibiae with a long and strong a and an extremely long and strong

ad somewhat before middle, the ad about five-sixths as long as the tibia, the

three basal joints of the hind tarsi somewhat compressed.

Wings slightly smoky, veins dark brown, R, very long and slender, very

slightly dilated to apex, y-m distinctly though slightly before middle of discal

cell. Calyptrae whitish-yellow with pale yellowish border, the lower one much

longer and strongly projecting, broadly rounded at apex. Halteres pale

ferruginous.

92 with the third antennal joint more pointed, the abdomen pointed, the

median vitta of the fourth segment not dilated behind, and the setae of the

legs stronger.

UGANDA: Ruwenzori, Kyarumba, 4500 ft. (D. R. Buxton) g type, 2 3

2 9 paratypes; Kilembe, 4500 ft. (F:W.E.) 2g 2 2 paratypes; Mobuku Valley,

4500 ft. (F.W.E.) 1 2 paratype; Fort Portal, 4.xii.1934 (F.W.E.) 1 3 paratype.

Bugomoba, 24.iv.1927 (H. Hargreaves) 12 paratype (returned to Mr. Har-

greaves).

Caricea kilembana sp. n. 3 2

Length 3-9-4°7 mm.; of wings 3°7-4°6 mm.

3. Head black, interfrontalia black with brown reflections, parafrontalia

dark brown dusted, only the extreme lateral margin narrowly golden dusted,

dorsal part of occiput dark brown dusted, a large, brown-dusted spot round

MUSCIDAE: B.—COENOSIINAE 217

the postverticals defined by a fuscous semicircle, rest of head densely grey

dusted. Frons projecting at base of antennae by the width of the latter, straight

in profile, much shorter than face, two-thirds as wide in front, almost as wide

at vertex as one eye, not much longer than wide at vertex, fronto-facial stripe

strongly narrowed to middle of third antennal joint, strongly widened from

there to lower margins of eyes (which slightly pass level of mouth-margin),

where it is distinctly wider than at vertex; parafrontalia with two strong and

one weak inclinate frontal seta. Inner vertical strong, outer vertical rather

indistinct, postverticals rather strong, ocellars strong. Antennae fuscous,

third joint more than six times as long as wide, reaching lower quarter to fifth

of eyes, falling short of mouth-margin by once or twice its width; arista short-

haired on basal three-quarters, half as wide to almost as wide, including plum-

osity, as third antennal joint. Palpi fuscous. Parafacialia and jowls almost

linear.

Thorax black, pleurae, humeri and lateral parts of the mesonotum densely

grey dusted, mesonotum and scutellum densely dark brown dusted, with a

pair of brownish-grey vittae from neck to suture which are continued along the

front side of the latter. acy fine but rather long, in two rows with a few addi-

tional setulae behind, 1-+-3 very long dc, the first post pair shorter, two well-

developed za; scutellum sparsely and finely setulose with two pairs of strong

setae, the lateral ones slightly shorter. Posterior propleural and upper prostig-

matal moderately long, lower prostigmatal fine, curved downward, surrounded

by a few fine hairs. stpl in a triangle with some long, erect hairs, among which

one near the posterior side forms a fourth stfl, the posterior side considerably

shorter, anterior and lower stfl subequal, not very strong, posterior one very

long and strong.

Abdomen as long as thorax, oblong-oval, slightly depressed at base, distinctly

compressed at apex, somewhat curved downward, black, rather dark grey

dusted, dorsum of the first to third segments with a broad, dark-brown, trans-

verse band on hind margin, which reaches more or less middle of segment at

sides and front margin in middle, fourth segment with a dark inverted-triangular

spot. Second to fourth segments each with a transverse discal row of 2-3 pairs

of rather strong setae, fourth besides with a long marginal lateral seta, fifth

with a pair of long and fine setae and a few fine, erect hairs.

Legs: front coxae pale ferruginous, mid and hind coxae piceous, grey

dusted, trochanters and base of the femora pale ferruginous, apical half and

dorsal surface of the anterior femora piceous, apical two-fifths and most of the

dorsal surface of the hind femora piceous, tibiae and tarsi piceous-black.

Front femora with 6-7 rather strong fd and five long fv setae, the latter alter-

nating with fine ones, front tibiae with a very long and strong fv just beyond

basal third; mid-femora with 6—8 rather strong a@ setae in basal three-fifths,

four long pu setae, alternating with fine ones, and two preapicals (f and fd),

218 RUWENZORI EXPEDITION

mid-tibiae with a moderately strong pd seta at middle and a subequal ad

somewhat beyond it; hind femora with 6-8 ad, about six fine but rather long

av, three long pv, and a fd preapical, hind tibiae with a very strong and long

ad at middle and a moderately long a just before middle. Basal joint of hind

tarsi strongly compressed and dilated, as wide as tibiae, with a rather broad,

bare longitudinal area, the next two joints also compressed and dilated, but to

a less degree (Fig. 73).

Wings strongly smoky and iridescent, veins dark brown, 7,,,; and m slightly

diverging at apex, y-m just beyond middle of discal cell. Calyptrae slightly

smoky with pale testaceous borders, the upper one strikingly short, the lower

one normal, very strongly projecting beyond the upper one, subparallel, broadly

rounded at apex. Halteres testaceous.

Q: abdomen shorter-oval, pointed and not compressed, thorax with five

narrow vittae, the median one complete, the subdorsal ones interrupted at

suture and scutellum, but continued at lateral scutellar setae, and the sub-

lateral ones connected in front with the subdorsal vittae and running over the

prst and sa setae (the presutural part of this pattern is recognisable also in the

male in posterior view, whereas the postsutural part of the pattern can seldom

be traced in that sex).

UcGanDA: Ruwenzori, Kilembe, 4,500 ft. (F.W.E.) 3 type; 73 3 @ para-

types.

Caricea kinangopana sp. n. 3 2

Length 4:3-5°2 mm.; of wings 4°7-5°5 mm.

3g. Head black, densely grey dusted, occiput brown dusted above, frons

dark brown dusted, parafacialia brownish at dorsal end. Head very high, one-

and-a-quarter times as high as long, frons much shorter than face, strongly

dilated to vertex, almost one-half wider at vertex than behind lunula, as long

as wide, fronto-facial stripe rather strongly narrowed to beyond insertion of

arista, considerably widened but hardly rounded to lower margin of eyes, where

it is as wide as at vertex; parafrontalia not distinctly defined from inter-

frontalia, with three inclinate frontal setae. Inner vertical very long, outer

vertical weak, postverticals and ocellars long and rather fine, the former

strongly, the latter moderately diverging. Antennae black, dark brown

dusted, inserted at dorsal third of eyes, long and slender, third joint more than

four times as long as wide, somewhat pointed dorsally at apex, almost reaching

lower sixth of eyes (the latter reaching or slightly passing level of mouth-

margin), falling short of mouth-margin by about twice its width; arista short-

haired, the longest hairs of ventral surface as long as the basal diameter of

arista, the longest hairs of dorsal surface almost half as long as the third

MUSCIDAE: B.—COENOSIINAE 219

antennal joint wide. Palpi piceous. Parafacialia hardly a third as wide as third

antennal joint, jowls much narrower than the latter.

Thorax black, dorsum and scutellum dark brown dusted, scutellum slightly

lighter at sides, humeri and sides of dorsum pale brown, pleurae grey dusted,

mesopleurae with a large brown spot. acy moderately long and numerous, in

two rows, the frsc pair distinctly longer and stronger, 1-+3 strong dc, two

moderate za, the anterior one level with the sa; scutellum finely setulose, with

the two pairs of setae very long and strong. Two strong propleurals, upper

prostigmatal strong, lower one fine, downwards curved, surrounded by a few

fine, small hairs. s¢f/ in an almost equilateral triangle, the posterior side of

which is slightly shorter, and which bears some fine, erect hairs and a stronger

seta between the posterior and lower s¢f/, the anterior and lower stf/ not very

strong, posterior one very long and strong.

Abdomen oblong, rather depressed, densely grey dusted, dorsum with a

broad transverse band on first to third segments which reaches hind margin all

over its width and front margin in middle, a triangle at each anterior angle

remaining grey, fourth segment brown at apex and on median line; second

segment with well-developed lateral and smaller sublateral setae, third and

fourth each with a transverse discal row of two (on the third segment three)

pairs of setae.

Legs almost entirely black, front coxae and all the trochanters ferruginous,

base of the mid and hind femora and of the tibiae more or less ferruginous-

translucent. Front femora with about seven not very long fd and 4-5 very

long pv, alternating with smaller ones, front tibiae with a long and strong pv;

mid-femora with 5—7 strong a setae from base to two-thirds, five fine av in basal

half, three very long pv, alternating with small ones, and two (pf and pd) pre-

apicals, mid-tibiae with a moderate pd slightly beyond middle and an ad of

equal length almost at three-fifths; hind femora with five ad in basal half, a

strong one at two-thirds and a preapical, six av, three long pv, and a pd pre-

apical, hind tibiae with a long and strong ad which is half as long as the tibia,

the a seta slightly shorter and very slightly more basad (by the diameter of

its pore).

Wings rather strongly infuscate, especially along front margin. Veins dark

brown, m slightly curved backward, 7,,; slightly sinuate before apex and

distinctly slightly diverging with m, v-m distinctly beyond middle of discal

cell. Calyptrae hyaline-brown with brown border and fringe, the lower one

much longer, strongly projecting. Halteres pale ferruginous.

2 similar to the male. Abdomen pointed at apex, ad seta of the mid-tibiae

much longer than the pd, two-fifths as long as the tibia. Calyptrae pale brown

with pale brown margin.

Kenya: Aberdare Range, Mt. Kinangop, 8000 ft., cedar forest, x.1934

(F.W.E.) 8 type; gooo ft. 2 2 paratypes.

II, 4 (i)

220 RUWENZORI EXPEDITION

Caricea rebmanni Speis.

Caricea vebmanni Speis., 1924, Beitr. Tierk., Kénigsb. (Widmungsschrift M. Braun):

102; Malloch, 1925, Ann. Mag. nat. Hist. (9)16: 363; Curran, 1935, Amer. Mus.

Novit. 776: Io.

Coenosia funuipennis Stein, 1913, Ann. Mus. nat. Hungar. 11: 575 (non Lamb).

UGANDA: Ruwenzori, Namwamba Valley, 6500 ft. (F.W.E. and E. G.

Gibbins) 18 3, 10 2; Bwamba Pass (West Side), 5500-7500 ft. (F.W-.E.) 1 &.

Typical localities: TANGANYIKA: Kibosho and Kilimanjaro. Reported

from BeLc. Conco: Burunga, by Curran.

Caricea morio sp. n. 3 Q

Length 4:2-5°6 mm.; of wings 4°5-5°8 mm.

3g. Head (Fig. 78) black, dark brown dusted, with some greenish-grey dust

on lower part of occiput. Frons evenly dilated to vertex, in front two-thirds,

at vertex four-fifths as wide as one eye; fronto-facial stripe narrowed to beyond

base of arista, somewhat widened to lower margin of eyes, where it is almost as

wide as at vertex, parafrontalia not defined from interfrontalia, with two strong

and between them one moderate or 2-3 small inclinate frontal setae. Inner

vertical strong, outer vertical weak, postverticals moderate, ocellars strong.

Antennae inserted at dorsal third of eyes, black, microscopic pile of the third

joint white, third joint five times as long as wide, dorsal apex bluntly pointed,

almost reaching level of mouth-margin, eyes fully reaching the latter; arista

practically bare beneath, short-pubescent above beyond the thickened part

for a third of the length of arista, the longest hairs slightly longer than the

thickened part wide. Palpi piceous. Parafacialia linear on lower half, jowls

almost linear. Vibrissae level with mouth-margin.

Thorax black, dark brown dusted, the pleurae, especially sternopleura and

hypopleura, largely greenish-grey dusted. acy rather strong, in two rows, with

a few additional hairs behind, no stronger prsc, 14-3 strong dc, two ta, the

anterior one fine, level with sa, scutellum sparsely setulose, with two pairs of

strong setae. Two propleurals and prostigmatals, the upper ones rather strong,

the lower ones small, no fine hairs around the prostigmatals. s¢f/ in a triangle,

which bears some rather long, erect hairs, and the posterior side of which is

distinctly shorter, the anterior stp/ weak, the other two very strong, especially

the posterior one.

Abdomen slightly depressed, somewhat club-shaped, strongly curved down-

ward, black, dark grey dusted with some parts more brownish, especially a

triangular area or a median vitta on dorsum; the transverse row of two or

three pairs of discal setae distinct on segments 2-4, fifth segment longer than

usual, with two pairs of setae behind each other and some small hairs.

Legs entirely black, front femora with four very long (and some shorter)

pv, front tibiae with the submedian fv and the preapical fd two-thirds to three-

MUSCIDAE: B.—COENOSIINAE 221

quarters as long as the tibia, mid-femora with about nine fine and short av

setae which do not much pass the middle, two strong a setae on and beyond

middle, 2-3 very long pu and two (pf and fd) preapical setae, mid-tibiae with a

long ad and a moderate fd on middle, level with each other, hind femora with

most of the av rather fine, but one strong av about at two-thirds, two very long

and fine pv and one ad and one fd preapical, hind tibiae with the submedian

setae slightly basad of middle, the a@ slightly basad of the ad, the latter three-

quarters as long as the tibiae, d preapical at apical quarter to fifth.

Wings somewhat smoky, strongly iridescent, veins brown, 7,,,; and m

slightly converging towards apex, but inconspicuously sinuate at apex, 7-m

slightly beyond middle of discal cell. Calyptrae whitish with yellowish borders,

the lower one slightly infuscate towards apex, upper one small, lower one

moderate, but strongly projecting, widened to beyond middle and almost semi-

circularly rounded at apex. Halteres black.

® similar to male, the abdomen fusiform, longer and more pointed, not

conspicuously curved downward.

Ucanpba: Kigezi District, Mt. Sabinio, 8000 ft., xi.1934 (F.W.E.) 3 type,

2 g-paratypes; Mt. Mgahinga, 8000 ft., x1.1934 (F.W.E.) 3 3, 9 2 paratypes;

Kanaba, 7300-7800 ft., xi.1934 (F.W.E.) 10 3, 3 2 paratypes.

Very closely related to the vebmanni group in the characters of the head

and chaetotaxy. In Stein’s and Malloch’s keys morio would run to fumisquama

Stein, but the latter is smaller (male 3-3-5 mm.), with the third antennal joint

hardly acuminate and much shorter, the 7,,; and m parallel at apex, and the

abdomen with a very distinct blackish-brown pattern. From brunnezfacies

Curr., with which morio has in common the size and the colour of the legs,

halteres and face, it can be distinguished by the brown dusted and wider frons,

the longer third antennal joint, the infuscate calyptrae, the smaller number

of inclinate frontals, and evidently the curved abdomen.

Caricea niveifrons group

1(6). Pollinosity of frons velvety dark brown to black in both sexes. Arista short-

haired.

2(3). Halteres black. Face velvety black. Longest hairs of the arista shorter

than its basal diameter. Humeri dark brown dusted like the mesonotum.

Lower calyptra distinctly reaching level of first abdominal segment,

calyptrae strikingly white. Epistoma slightly produced, forming a

rounded prominence in dorso-frontal view. Scutellum setulose on disc.

Parafacialia, jowls, and a seta of mid-femora as in aygentipes. Hind

legs of male without silvery pile. 6 : : [velutinifacies sp. n."]

3(2). Halteres yellow. Face silver-grey to silver-white. Longest bairs of the

arista longer than its basal diameter. Humeri grey dusted, mesonotum

dark brown dusted. Lower calyptra not passing hind end of thorax,

calyptrae more hyaline with the border more or less yellowish.

1 A detailed description of this Abyssinian species will be published in Ann. Mag, nat. Hist. to4o.

iS)

11(10).

7(14).

18(19).

RUWENZORI EX <PEDITION

Epistoma slightly produced, forming a rounded prominence in dorso-frontal

view (Fig. 4). Parafacialia about one-half as wide, jowls almost as

wide as the third antennal joint. Scutellum setulose on disc. a setae

of mid-femora of moderate length, not more than a quarter the length

of the femur. Ventral surface of the hind legs with dense silvery pile

inmale. . A : . argentipes sp. N., p. 224

Epistoma truncate in font not produced ene a straight transverse line

in dorso-frontal view. Parafacialia and jowls linear. Scutellum bare,

except for two pairs of strong and one pair of weak setae. The most

distal one of the a setae of the mid-femora of unusual length, one-third

as long as the femur (in male). . : : macochaeta sp. T., Pp. 225

Pollinosity of frons not velvety, silver-white in male, if the halteres are

yellow.

Halteres yellow.

Abdomen immaculate, depressed at base in male. Knees yellow. [*dzluta Stein]

Abdomen with a distinct pattern, subcylindrical and curved, or compressed

in male. Knees not appreciably paler.

. Scutellum setulose. Abdomen with a faint median vitta and paired spots,

setae of the third and fourth segments very strong; abdomen of male

subcylindrical and slightly curved. Thorax without a distinct pattern

or with a more or less complete median vitta. Femora and tibiae

greyish dusted. Third antennal joint pointed, arista short-haired at

base and more than half as wide, including plumosity.

planifrons Stein, p. 227

Scutellum bare except for two pairs both of setae and of setulae. Abdomen

with the first to fourth segments dark brown dusted, except for a grey-

dusted lateral anterior triangle; setae hardly differing from the long

and erect hair of the ventral surface; abdomen of male somewhat com-

pressed. Thorax with five narrow brown vittae. Femora and tibiae

not dusted, black and shining. ~. 6 5 gibbinsi sp. n., p. 22

Knob of halteres entirely dark brown or Noss Scutellum bare, except for

two pairs of setae and two pairs of setulae (one discal sub-apical and one

lateral sub-basal) seldom with 1-2 additional setulae or (byvunnei-

facies) more numerous setulae.

3 3.

Frons brown dusted, moderately widened to vertex. Ocellar bristles present.

Dorsum of thorax dark brown dusted.

. Calyptrae brown. Abdomen somewhat compressed, with a dark brown

median vitta and lateral transverse bands along the hind margins,

which are almost fused with the median vitta. [ fumisquama Stein], p.

. Calyptrae whitish. Seven to nine pairs of frontal setae. Abdomen de-

pressed, entirely dark brown dusted except for a not very conspicuous,

grey dusted, narrow triangle along lateral parts of front margins. prst

and post acy well developed. é : multisetosa sp. D., p.

Frons entirely covered by silver-white prainesity. Ocellars absent.

Third antennal joint pale yellow, silver-pollinose. Frons slightly dilated to

vertex. Thorax and abdomen largely silver-greyish dusted with a

1 The ocellar bristles are absent in all the males with silver dusted frons of this group

which I have seen, whereas the postvertical setae are well developed. According to Curran

the ocellars are present, however, in brunneifacies, which by this character, and the more

numerous frontal setae would appear to be more closely related to multisetosa m.

25(24).

26(27).

27(26).

28(21).

29(30).

30(29).

MUSCIDAE: B.—COENOSIINAE 223

patch of dark brown dust in front of scutellum. pyst and post acy well

developed. Apex of front coxae largely yellowish, with silver-white

pruinosity. Calyptrae whitish. Abdomen depressed. canifrons Stein, p. 230

. Third antennal joint black, though sometimes densely silver-pollinose.

. Calyptrae whitish, with whitish or yellowish border and fringe.

Frons slightly dilated to vertex, much longer on median line than wide at

vertex.

. Face brownish or brownish-yellow pollinose. Seven to eight pairs of frontal

bristles, one or two of them developed as hairs. Dorsum of thorax

silver-grey pollinose, the scutellum and Daas part more cinereous.

Ocellars long and fine. é 3 : [tbrunneifacies Curr.]

Face silver-grey pollinose. Three to four pairs Sof frontal bristles (praeacuta?).

. Third antennal joint and face densely silver-white pollinose. Hind femora

with a complete row of long ad, most of which are almost or fully twice

as long as the greater diameter of the femur, some of the pu almost

two-thirds as long as the hind tibia. Front half of thorax silver-grey,

posterior half and scutellum dark brown pollinose. chaetomevos sp. n., p. 230

Third antennal joint blackish-brown pollinose with at most some slight

greyish reflections. Hind femora with shorter setae, only two to four ad

setae being slightly longer than the greater diameter of the femur, and

with only three rather long pu setae, of which two in the third and fourth

fifths, and none more than half as long as the hind tibia (praeacuta?).

Thorax and abdomen dark brown pollinose, slightly greyish on sides. Ab-

domen somewhat depressed in basal half, somewhat compressed at

apex. Parafacialia very narrow, less than half as wide as the third

antennal joint. pyst and post acy well developed.

praeacuta namwambae ssp. N., p. 232

Thorax and abdomen silver-grey pollinose. Abdomen slightly compressed.

[*praeacuta Stein]

Frons strongly dilated to vertex, almost as wide above antennae as one eye,

hardly longer on median line than wide at vertex (ig. 79). Dorsum

of thorax entirely or largely, dorsum of abdomen entirely silver-grey

dusted.

prst acy absent, post acy very sparse and small. Dorsum of thorax and

abdomen entirely silver-grey dusted. Face brownish.

niveifrons aygentescens ssp. D., p. 234

prst and post acy present, long and fine. Posterior part of mesonotum (about

half of the postsutural part) and the scutellum brown dusted. Face

more or less grey. : : niveifvons albisquama ssp. N., p. 233

. Calyptrae dark brown, with oie border and fringe. Frons strongly

dilated to vertex. Face, thorax, and abdomen entirely dark brown

dusted and dull. Two orthree pairs of prst acy present. niveifrons Stein. p. 232

2 9. Frons dark brown dusted. Ocellars present.

. Calyptrae whitish with the border whitish or yellowish and the fringe

whitish.

Frons moderately widened to vertex, at least a third longer than wide at

vertex.

. Six to nine pairs of frontal bristles, one or two of them hair-like. Face dark

brown with slight greyish reflections.

[tbrunneifacies Curr.] and muitisetosa sp. n., p. 229

1 See note p. 222.

224 : RUWENZORI EXPEDITION

36(35). Three to five pairs of frontal bristles. Face grey pollinose (always?).

37(38). Frons slightly widened from base of antennae to second frontal bristles,

very slightly dilated from there to vertex, at most one-and-a-half

times as wide at vertex as the third antennal joint is long.

canifryons Stein, p. 230

38(37). Frons gradually widened from base of antennae to vertex, broader, almost

or fully twice as wide at vertex as the third antennal joint is long, the

narrowest point of the fronto-facial stripe much below middle of head.

(2 of praeacuta unknown.)

39(40). Frons slightly widened to vertex, one-quarter to one-third wider at vertex

than at first frontal setae. . : : chaectomeros sp. N., p. 230

40(39). F'rons more considerably widened to vertex: one-half wider at vertex (han

at first frontal setae. . : praeacuta namwambae ssp. N., Pp. 232

41(34). Frons strongly widened to vertex, hardly or at least less than a third longer

on median line than wide at vertex.

42(43). prst acy absent. Face dark brown. F niveifvons argentescens ssp. N., Pp. 234

43(42). prst acy present. Face more or less greyish. niveifrons albisquama ssp. N., p. 233

44(33). Calyptrae brown, at least their border.

45(46). Frons strongly widened to vertex, hardly longer than wide at vertex, with

four to five pairs of frontal setae. pyst and post acy well developed.

niveifrons niveifrons Stein, p. 232

46(45). Frons moderately widened to vertex, a third longer than wide, not much

wider at vertex than the third antennal joint is long. Mid-tibiae with

two long setae on posterior surfaces. fumisquama Stein?, p. 228

Caricea argentipes sp. n. 3 2

Length 2:8-4:6 mm.; of wings 3°3-4°3 mm.

3g. Head (Fig. 4) black, occiput grey dusted, frons entirely velvety black,

face wholly silver-white pollinose. Frons slightly projecting at base of an-

tennae, as long as face, distinctly narrower in front than one eye, evenly and

moderately dilated to vertex, fronto-facial stripe not distinctly narrowed from

there to lower margin of eyes, where it is one-and-a-half times as wide as at

vertex, mouth-margin at least as much projecting as frons, forming a short

snout, which is conspicuously rounded anteriorly in dorsal view; parafrontalia

not defined, with 2-3 strong and 3-4 fine and minute setae. Inner vertical very

strong, outer one indistinct, postverticals long, ocellars not very long. Antennae

fuscous, second joint with light brown pollinosity interiorly at apex, third joint

not quite thrice as long as wide, acuminate at dorsal apex, reaching lower fifth

or sixth of eyes, its apex almost as distant from mouth-margin as from base of

arista; the latter short-haired in basal two-fifths, the longest hairs slightly

longer than the basal diameter of the arista. Palpi black. Parafacialia almost

half as wide, jowls almost as wide as third antennal joint.

Thorax black, pleurae grey dusted with bluish-grey reflections, dorsum in-

cluding scutellum dark brown dusted, humeri and notopleurae pale grey

dusted. acy rather long though fine, in two rows, more numerous towards

MUSCIDAE: B.—COENOSIINAE 225

scutellum, 1-+3 very long and strong dc, two rather strong za; scutellum

setulose, with two pairs of long and strong setae. Posterior propleural and

upper prostigmatal moderately strong, lower prostigmatal fine, curved down-

ward, a few small hairs close by. s¢f/ in a triangle which bears some fine, long,

erect hairs, and the anterior and posterior sides of which are considerably shorter

than the upper side, anterior s¢f/ moderately strong, lower stpl strong, posterior

stpl very long and strong.

Abdomen oblong-oval, rather depressed near base, somewhat compressed

at apex, black, densely grey dusted, with a brown median vitta on first to

fourth segments and a pair of roundish spots on second to fourth; these seg-

ments each with a transverse discal row of 2-3 pairs of rather strong setae,

fifth, segment with a pair of rather long, fine setae and rather numerous small

hairs.

Legs wholly black, ventral surface of hind femora silver dusted, of hind

tibiae and tarsi densely beset with fine, minute, appressed silver-white hairs.

Front femora with 6-8 rather strong pd, posterior surface with rather numerous

erect setae, 7-8 rather long pv, alternating with small ones, front tibiae with a

long and strong pv seta slightly beyond middle; mid-femora in basal two-

thirds with five strong a setae which increase in lengtin, four long pv in basal

three-fifths, alternating with fine ones, and two strong preapicals (f and fd),

mid-tibiae with a moderately strong pd at middle and a strong ad almost half-

way between middle and three-fifths; hind femora with 6-7 strong ad, a rather

irregular row of av, among which a strong one at a third and two-thirds, and a

strong preapical, about five fv among which only two—one before and one

behind middle—are long, and a strong fd preapical, hind tibiae with a very

strong pd and a moderately strong @ at middle.

Wings slightly smoky, veins brown, 7,,, and m parallel at apex, vm slightly

before or in middle of discal cell. Calyptrae not very large, the lower one

projecting by about the length of the upper one, hyaline with testaceous border.

Halteres pale ferruginous.

9 not differing from male, except by the formation of the abdomen and the

absence of the silvery pollinosity and hair of the hind legs.

KENYA: Aberdare Range, Nyeri Track, 10,500 ft., x.1934 (F.W.E.) 3 type;

Mt. Kinangop, 10,000 ft., x.1934 (F.W.E.) 2 g 1 Q paratypes; Mt. Elgon,

Alpine Zone, 12,000-13,000 ft., 11.1935 (f.W.E.) 2 3 paratypes (one on Lobelia

elgonensis); NATAL: Ulundi, 5000-6500 ft., ix.1896 (G. A. K. Marshall) 1 °

paratype.

Caricea macrochaeta sp. n. 5

Length 2:8-3°5 mm.; of wings 3:0-3:8 mm.

6. Head black, dark brown dusted, except on lower quarter of occiput,

jowls, face, and parafacialia, which are grey dusted, frons black, dark velvety-

226 RUWENZORI EXPEDITION

brown dusted. Frons rounded in profile, not protruding at lunula, slightly

more than half as wide at lunula as one eye, gradually widened with straight

sides to vertex, where it is two thirds wider than anteriorly, more than twice as

long as wide at lunula, fronto-facial stripe rather strongly narrowed to middle

of third antennal joint, gradually widened and rounded from there to lower

margin of eyes, where it is very slightly wider than at vertex; parafrontalia

not well defined, with two long inclinate frontal setae. Inner vertical strong,

outer vertical almost indistinct, postverticals and ocellars fine. Antennae dull

black with brown reflections, third joint thrice as long as wide, reaching lower

quarter of eyes (the latter slightly passing level of mouth-margin), apex as

distant from mouth-margin as from base of arista; the latter bare on apical

half, short-haired on basal half, the longest hairs slightly longer than the basal

diameter of arista. Palpi black. Parafacialia and jowls linear.

Thorax black, dorsum including scutellum densely dark brown dusted,

humeri and notopleural depression greyish-brown, pleurae grey dusted. acr

fine, sparse, in two rows, I+3 long and strong dc, posterior za well developed;

scutellum with 2~—3 pairs of setulose hairs and the normal two pairs of long and

strong setae. Propleurals and prostigmatals fine, not very long, lower stp/ curved

downward, not surrounded by small hairs. s¢f/ in a triangle which bears some

erect setulose hairs and the posterior side of which is conspicuously shorter,

anterior stpl rather fine, not very long, lower one slightly shorter than upper

one, both very long and strong.

Abdomen black, grey dusted, dorsum dark brown dusted, but the anterior

angles broadly greyish; rather short, not much longer than thorax, somewhat

depressed at base, somewhat compressed at apex, markedly curved downward;

second to fourth segments each with a transverse discal row of two pairs of

setae (the second with two to three finer pairs besides) which increase in length

posteriorly, fourth segment besides with a pair of fine, long marginal setae,

fifth with a pair of fine setae, third and fourth sternites with a pair of long, fine,

erect setae.

Legs wholly black, only slightly dusted, shining. Front femora with 4-5 pd,

some fp setae, and five long pv, partly alternating with fine ones, front tibiae

with a very long (four-fifths as long as tibia) pu seta just beyond two-fifths;

mid-femora with some small a setae near base, a rather long one at middle and a

very long one (a third as long as femur) at three-fifths, four very long fv, and

two preapicals (p and fd), mid-tibiae with a very long a and # seta in middle;

hind femora with five rather strong ad, a row of fine short av setae in basal half

and a very strong one at three-fifths, a moderate pv at one-fifth and two very long

and strong ones at two- and three-fifths, and a pd preapical, hind tibiae with an

extremely long pd (four-fifths as long as tibia) at two-fifths and a somewhat

shorter a seta slightly before it, the d preapical seta very long, at the apical

quarter.

MUSCIDAE: B.—COENOSIINAE 22'7

Wings rather smoky, veins brown, testaceous at base, 7,,; and m parallel at

apex, v-m at middle of discal cell. Calyptrae whitish-hyaline, slightly infuscate

near margin, the border pale testaceous, upper calyptra very small and short,

lower one about thrice its length, strongly projecting, subparallel, rather broadly

rounded at apex. Halteres pale ferruginous.

2 very similar to male, frons half as wide at lunula as one eye, abdomen

pointed at apex, its setae strongest on third segment, the setae of the legs a trifle

shorter in relation to the length of the legs, calyptrae more distinctly smoky

towards margin.

UGANDA: Budongo Forest, 7.-8.i11.1935 (F.W.E.) 3 type; Neagu, Busiro,

5.vili.1929 (H. Hargreaves) 1 9 paratype.

So far as the long a seta of the mid-femora and the parafacialia and jowls,

which do not protrude at all in lateral view, are concerned, this species would

fit Stein’s description of pzlifemur even better than the species thus identified

by Malloch and me, but the additional d seta on the apical third of the front

tibiae in pilifemur, which is mentioned in Stein’s description, and which is

quite exceptional in this genus, proves that Malloch’s identification is correct.

Caricea planifrons Stein

Kenya: Aberdare Range, Mt. Kinangop, 8000-gooo ft., in Cedar Forest,

x.1934 (F.W.E.) 3 3; Katamayo, 8000 ft., x.1934 (F.W.E.) 2 g; Nairobi,

x.1934 (F.W.E.) 1 g. UGanpa: Kampala 12.xii.1934 (F.W.E.) x 2.

Typical locality: TANGANYIKA: Arusha-Ju.

The female is unknown, and though it is remarkable that Dr. Edwards

should have collected seven males without a single female, I do not think that

the latter can erroneously have been referred to another species.

Caricea gibbinsi sp. n. 3

Length 2-9 mm.; of wings 3°2 mm.

Head black, densely grey dusted, frons, face and jowls silver-grey dusted.

Frons subparallel, almost as wide as one eye, slightly shorter than face, fronto-

facial stripe very slightly narrowed from second inclinate frontal to apex of

second antennal joint, rather strongly dilated and slightly rounded to lower

margin of eyes, where it is almost one half wider than at frons; parafrontalia not

defined from interfrontalia, with three inclinate frontal setae. Inner vertical

strong, outer vertical indistinct, postverticals and ocellars moderately long.

Antennae fuscous, second joint silver-white dusted interiorly at apex, third

silver-grey dusted along interior surface of dorsal margin, almost thrice as long

as wide, apex rounded, reaching lower quarter of eyes (the latter not quite

reaching mouth-margin) distant from mouth-margin by not quite the length

ioe)

RUWENZORI EXPEDITION

of the third joint; arista bare in apical three-fifths, short-haired in basal two-

fifths, the longest hairs slightly longer than the basal diameter of arista. Palpi

black. Parafacials more than half as wide as, jowls slightly wider than third

antennal joint.

Thorax black, densely grey dusted, with five narrow brown vittae, which

do not much pass the frst dc in front and broaden and coalesce towards

scutellum, disc of scutellum brown. acy rather strong, in two rows, I+3

strong dc, two strong 7a; scutellum with two pairs (a lateral sub-basal and a

discal sub-apical one) of setulose hairs and two pairs of setae. The two propleural

setae rather fine, upper prostigmatal moderate, lower one fine, curved downward.

stpl in a triangle, the posterior side of which is shorter and which bears several

erect fine hairs, anterior stf/ weak, lower one rather strong, posterior one very

strong.

Abdomen rather short, hardly depressed at base, considerably compressed at

apex, black, densely grey dusted, first to fourth segments each with a transverse

band of dark brown dust along hind margin, which reaches front margin in

middle; hairs appressed on dorsum, erect at sides and venter, the setae not

distinguishable, fifth segment with a pair of rather long, fine setae.

Legs entirely black, coxae dusted, rest glossy. Front femora with four

not very strong pd in apical two-thirds, six pv which decrease in size and

alternate with weaker ones, front tibiae with a long pv which is half as long as

tibia; mid-femora with five a setae in basal two-thirds, six fine, rather short av

in basal half, four pv, alternating with weak ones, and two (fd and /) preapicals,

mid-tibiae with a rather strong ad at three-fifths and a weaker fd slightly

beyond middle; hind femora with six rather strong ad, six fine and short av

in basal two-thirds and a preapical one, 3-4 pv, the last but one strong, at

two-fifths, and a pd preapical, hind tibiae with a strong ad at middle, which is

half as long as tibia, and a slightly shorter a seta at practically the same level.

Wings hyaline, hardly smoky, strongly iridescent, veins brown with slightly

paler base, 74,; and m almost parallel, very slightly converging at apex, 7-m

distinctly beyond middle. Calyptrae hyaline with pale yellow border, the lower

one strongly projecting and broadly rounded at apex. Halteres pale reddish-

brown.

UGANDA: Ruwenzori, Namwamba Valley, 6500 ft. (EF. G. Grbbins) 3 type,

I paratype, (¢?, abdomen lost).

Caricea fumisquama Stein

Ucanpa: Ruwenzori, Namwamba Valley, 10,100 ft. (I. H. E. Jackson),

2 9; Mobuku Valley, 4500 ft. (F.W.E.) 1 9.

Typical locality: AByssinrA: Marako. In B.M. from UGANDA: Segibwa

Falls, attracted by human faeces, 29.11.1935 (E. G. Gibbins).

MUSCIDAE: B.—-COENOSIINAE 229

As no male of this species is available, the identification is somewhat un-

certain; in these females the abdomen is wholly dark brown dusted on dorsum,

whereas the males described by Stein have a black median vitta and transverse

hind marginal bands. The females at hand show an outstanding character

which is unique, as far as I am aware, in Caricea: the mid-tibiae have, instead of

one, two long posterior setae, a pv slightly beyond two-fifths and a fd (or

perhaps rather ~) at two-thirds, whereas the ad (or perhaps rather a) seta is

inserted at middle.

Caricea multisetosa sp. n. 5 2

Length 2-9-3-4 mm.; of wings 2:9-3°6 mm.

¢. Head black, wholly dark brown dusted, only the lower half of the occiput

grey dusted. Frons longer than face, almost two-thirds as wide in front as one

eye, with straight sides, gradually and evenly dilated to vertex, where it is

almost twice as wide as in front, almost a third longer than wide at vertex;

fronto-facial stripe very slightly narrowed to apex of second antennal joint,

gradually widened and moderately rounded to lower margin of eyes, where it is

as wide as at vertex; parafrontalia not defined, with 5~—7 inclinate frontal setae,

the reclinate seta not very distinct from the inclinate ones, often a hair-like

seta between the reclinate and inner vertical setae. Inner vertical strong, outer

one weak though distinct, postverticals moderately strong, ocellars very long.

Antennae fuscous, brown dusted, third joint two-and-a-half times as long as

wide, reaching lower quarter of eyes (the latter just passing level of mouth-

margin), apex as distant from mouth-margin as from base of arista, the latter

bare on almost apical three-fifths, very short-haired in basal part, the longest

hairs hardly longer than the basal diameter of arista. Palpifuscous. Parafacialia

linear, jowls half as wide as third antennal joint.

Thorax black, densely dark brown dusted, pleurae more greyish dusted.

acy sparse, one or two pairs in front of suture, a few more in two irregular rows

on anterior half of postsutural part and some widely scattered ones in posterior

half, 1-+-3 strong dc, two rather strong za; scutellum with two pairs of fine hairs

and two pairs of strong setae. Anterior propleural not much shorter though

much finer than posterior one, the latter moderately strong, lower prostigmatal

rather long, curved downward, upper one moderately strong, with or without

a small adjacent hair. stf/ in an almost equilateral triangle, which bears some

long, erect hairs, one of them forming a fourth sff/ between the posterior and

lower ones, anterior stf/ rather weak, lower one slightly longer, posterior one

very long and strong.

Abdomen oblong-oval, rather depressed, with the apex slightly compressed

on ventral surface, black, dorsum dark brown dusted, with an inconspicuous

greyish triangle in each anterior corner of the segments, ventral surface grey

230 RUWENZORI EXPEDITION

dusted; first segment with a shorter fan of erect hairs behind the normal one,

second and third segments also with some hairs at sides, second to fourth

segments each with a transverse discal row of 2—3 pairs of setae, these rows closer

to hind than to front margins, particularly so on the fourth segment, fifth with

two pairs of fine setae.

Legs wholly black, shining, hardly dusted except on coxae which are brown

dusted. Front femora with 5-6 fd setae, several p rows of more’ than twelve

strong setulose hairs, and seven pv setae, alternating with smaller ones, front

tibiae with a long pv slightly beyond middle; mid-femora with about eight small

a in basal half, three rather long pv setae in basal three-fifths, and two preapicals

(p and pd); hind femora with 7—9 strong ad setae, a not very strong av seta at

two-thirds and an av preapical, a long pu seta at two-fifths, and a fd preapical,

hind tibiae with a long fd slightly beyond two-fifths and a long and fine a seta

close and very slightly basad to it.

Wings strongly infuscate, with the veins dark brown, 7,,, and m almost

parallel at apex, v-m slightly beyond middle of discal cell. Calyptrae whitish

with yellowish-white border, lower one strongly projecting, broadly rounded at

apex. Halteres dark brown.

2 similar to male, the abdomen pointed, the setae somewhat stronger.

UGANDA: Kigezi District, Mt. Sabinio, 7000 ft. (F.W.E.) 3 type, I 2 para-

type; 8000 ft., 1 ¢ paratype; Mt. Mgahinga, 8000 ft., 20.xi.1934 (F.W.E.)

I Q paratype.

Caricea canifrons Stein

UGANDA: Ruwenzori, Fort Portal, 4.xii.1934 (F.W.E.) 1 3; Kilembe,

4500 ft. (F.W.E.) 19. Typical locality: TANGANYIKA: Arusha-Ju.

The female is undescribed. Like all the females in this group it differs from

the male by the brown dusted frons and the presence of well-developed ocellar

setae. The third antennal joint is fuscous, and the thorax and abdomen are

largely dark brown dusted, as in the females of niveifrons albisquama and argen-

tescens. As in the male, the frons is not very narrow in front (approximately as

wide as the third antennal joint is long) and is only very slightly dilated to

vertex, where it is about one-and-a-half times as wide as that joint is long. Two

fine inclinate frontal setae and one small hair.

Caricea chaetomeros sp. n. 3 2

Length 2-8-3°5 mm.; of wings 2:8-3-4 mm.

3g. Head black, grey dusted, fronto-facial stripe densely silver-white dusted.

Irons protruding by almost width of antennae, plane, straight in lateral view,

somewhat concave on disc, only slightly narrower in front than one eye, with

straight sides, a third wider at vertex than in front, much longer than face,

MUSCIDAE: B.—COENOSIINAE 231

fronto-facial stripe slightly narrowed to apex of second antennal joint, rather

strongly dilated with almost straight sides to lower margin of eyes, where it is

at least as wide as at vertex; parafrontalia not differentiated, with two long and

fine inclinate frontalsetae. Inner vertical strong, outer one minute, postverticals

rather fine, ocellars absent (ocelli very small, not prominent). Antennae fuscous,

entirely densely silver-white dusted, the second joint somewhat pale translucent

towards apex, third joint two-and-a-half times as long as wide, reaching lowest

fifth of eyes (the latter reaching level of mouth-margin), apex as distant from

mouth-margin as from base of arista; the latter bare on apical three-fifths,

very short-haired near base, the longest hairs hardly longer than basal diameter

of arista. Palpi black. Parafacialia half as wide, jowls slightly more than half

as wide as third antennal joint.

Thorax black, not very densely pale grey dusted, postsutural part, except

for front margin and anterior half of sides, and scutellum dark brown dusted.

acy sparse, in two rows, at most single setulae outside these rows, I + 3 strong dc,

two rather strong 7a; scutellum with two pairs of setulose hairs (and sometimes

an odd one) and two pairs of strong setae, the lateral ones slightly shorter than

the apical pair. Anterior propleural not much shorter than the posterior one,

lower prostigmatal not much shorter than the upper one, curved downward, one

or two rather long, up-curved hairs close to the upper prostigmatal. Anterior

and lower stp/ more or less indistinct among the numerous, long, strong, erect

hairs, the posterior one long and rather strong, the additional one between the

posterior and lower stf/ long and rather strong.

Abdomen very short, as broad as long, roundish-subtriangular, somewhat

shorter and broader than thorax, strongly depressed, black, thinly blackish-

brown dusted, rather glossy; hairs rather appressed on dorsum, erect at sides,

the setae fine and not very distinct, one stronger marginal seta at sides of

second to fourth segments, hind margin of fourth segment with a row of stronger

hairs, fifth segment with rather numerous, erect, setulose hairs.

Legs entirely black except for the trochanters, which are ferruginous.

Front femora with 6-10 fd setae, 2-3 rather long fp near base, and fine ones up

to apex, surface besides densely fine-haired, and 7-9 very long and fine pu

which alternate with somewhat shorter ones, front tibiae with a very long and

fine pv seta; mid-femora with 7-9 a setae in basal two-thirds, 8—10 fine and rather

long av in basal two-thirds, six long and fine pv, alternating with fine ones, four

very long and rather strong f setae in basal two-thirds, and two strong pre-

apicals (pf and fd), mid-tibiae with a very long and fine fd slightly beyond middle

and a shorter, fine ad almost at three-fifths; hind femora with ro—12 long ad,

about eight long and fine av in basal two-thirds, three very long, and two or

more slightly shorter pv in basal four-fifths, and a pd preapical, hind tibiae

with a very long and rather strong ad somewhat before middle and a finer a

close to it and very slightly basad.

232 RUWENZORI EXPEDITION

Wings strongly infuscate, with the veins dark brown, 7,,; and m parallel

or very slightly converging, v-m at middle of discal cell. Calyptrae whitish with

whitish border, lower one strongly projecting, twice as long as upper one,

broadly rounded at apex. Halteres blackish.

2 uniformly dark brown dusted except for face, parafacialia and apex of

second antennal joint which are silver-white dusted, anterior declivity of thorax

which is greyish dusted with a median extension of the brown dust to neck, and

lower parts of pleurae which are more or less greyish dusted. Frons not concave,

with two strong and 1-2 fine frontal setae, postverticals stronger, ocellars well

developed though not very strong, ocelli well developed. Abdomen pointed

but very little longer than in male, third segment with a distinct though fine

discal lateral, fourth with a discal row of two pairs of rather fine setae. The av

of the mid and hind femora short as usual, the long # of the mid-femora absent,

the pu of the hind femora stronger, only two of them very long, and a smaller

one before and beyond them.

S. RwopeEsiA: Umtali District, Vumba Mts., 11.1938 (Alex. Cuthbertson)

6 type, I @ paratype; iii1.1935, I 2 paratype; UGanpa: Lake Nabugabo,

13.x1.1934 (F.W.E.) 1 g paratype. ;

This species had formerly been identified as praeacuta Stein and was men-

tioned under that name by Cuthbertson (1938, Trans. Rhodesia Sc. Assoc.

36: 119) as preying on Szmulium lepidum De Meill.

Caricea praeacuta namwambae ssp. n.

These specimens run in Stein’s key to praeacuta Stein which was described

from Diredaua. They fit the description well, but the thorax and abdomen are

largely dark brown dusted, only the sides and anterior end grey (but not silver-

grey) dusted. The frons is two-thirds to three-quarters as wide anteriorly as

the eye. The arista is short-haired on basal two-fifths, the longest hairs being a

third as long as the third antennal joint is wide (the arista is said to be bare in

praeacute, but this is also said of nzverfrons, where the arista is similarly haired),

the abdomen is somewhat depressed in basal half, somewhat compressed at

apex in the male.

UGANDA: Ruwenzori, Namwamba Valley, 6500 ft. (F.W.E.) g type, I 3

(without head and abdomen) 2 2 paratypes; Kyarumba, 4500 ft. (D. R. Buxton)

1 9 (teneral); Bwamba Pass (West Side), 5500-7500 ft. (F.W.E.) 1 2 paratype.

Caricea niveifrons Stein

UGANDA: Ruwenzori, Fort Portal, 4.xii.1934 (F.W.E.) 1 $; Kilembe,

4500 ft. (F.W.E.) 1.

Typical locality: TANGANYIKA: Kilimanjaro. Seen from KENYA: Naivasha,

MUSCIDAE: B.—COENOSIINAE 233

vil.1937 (H. J. A. Turner, Coryndon Memorial Museum) and Chyulu Hills,

5600 ft., v._vii.1938 (Exped. Chyulu Hills, Coryndon Memorial Museum).

Caricea niveifrons albisquama ssp. n. 3 2

In Stein’s key this form would point to mveifrons by its broad frons, but to

praecuta by the white calyptrae, whereas in Malloch’s key it would run down to

praeacuta. It is certainly only a form of mzverfrons, which must necessarily be

named owing to the striking difference in the colour of the calyptrae which are

dark brown in niveifrons and white in the present form, and to the fact that the

two forms evidently are mutually exclusive. The face is more or less grey in

both sexes, whereas it is brown in mivezfrons, and the prst acy are well developed

as in that form. The broad frons (Fig. 79) precludes its being united with

praeacuta. Yhe thorax and abdomen are silver-grey dusted with silky gloss in

male, with the posterior half or third of the postsutural part of mesonotum and

the entire scutellum dark brown dusted, whereas the thorax and abdomen are

wholly dark brown dusted in niveifrons. In the female the frons is brown

dusted with the anterior part often more or less greyish, the pleurae, humeri and

anterior declivity of the thorax are also more or less greyish dusted, as are the

sides and anterior angles of the abdominal segments, but this greyish dust is

never very conspicuous.

Kenya: Aberdare Range, Mt. Kinangop, 8000-10,000 ft., x.—-I.xi.1934

(F.W.E. and J. Ford) 3 type, 27 3 28 2 paratypes (8 d 3 Y in Cedar Forest) ;

Nyeri Track, 10,500 ft., 28.x.1934 (F.W.E.) 1 g paratype. UGANDA: Mt. Elgon,

Bulambuli, 9500 ft., viii.1934 (J. Ford) 1 g paratype. Kenya: Mt. Elgon,

Heath Zone, 10,500-11,500 ft., 11.1935 (F.W.E.) 4 3 paratypes; 11,500 ft. on

Artemisia flowers, I 2 paratype.

The males from Mt. Elgon are quite typical, and so is the female mentioned

above. There are, however, two females from UGANDA: Mt. Elgon, be-

tween Butandiga and Bulambuli, 8000 ft., 7.viii.1934 (J. Ford), and KENyA:

Mt. Elgon, Heath Zone, 10,500-11,500 ft., 11.1935 (f.W.E.), in which the face

is brown and the prst acy absent, so one would be inclined to regard them as ssp.

argentescens. On the other hand, two male specimens from Nairobi, x.1934

(F.W.E.) have the face brown and the frst acy absent like ssp. argentescens, but

the scutellum and posterior part of mesonotum brown like ssp. albisguama.

A female from Nairobi, 111.1935 (Van Someren, Coryndon Memorial Museum),

shows the same characters as all these specimens, except of course for the bi-

coloured thorax of the males. Judging from the characters of the males |

regard these specimens as transitions between the two subspecies. This tran-

sition is also found on the Aberdares, though very much rarer than the typical

ssp. albisquama: Mt. Kinangop, 8000 ft. (J. Ford) 1 3 1 9, gooo ft., in Cedar

Forest (F.W.E.) 1 9.

234 RUWENZORI EXPEDITION

Caricea niveifrons argentescens ssp. n. 3 2

Distinct from niveifrons by the white calyptrae and absence of prst acr in

both sexes and by the silver-grey, silky pollinosity of thorax and abdomen

in the male, and distinguished from ssp. albisgquama by the absence of brown

dust on the hind part of the mesonotum and scutellum in the male (though this

is present on the apex of the scutellum in one or two specimens), and by the

dark brown dusted face and the absence of frst acr.

UGANDA: Ruwenzori, Namwamba Valley, 6500 ft. (F.W.E.) ¢ type, 5 3

4° paratypes; Mobuku Valley, 7300 ft. (F.W.E.) 4 g paratypes ; Nkokonjoro,

23.-27.x11.1926 (G. L. R. Hancock) 1 9. Kenya: Londiani, v.1936 (H. J. A.

Turner).

The females from Londiani and Nkokonjoro, as no males from these localities

are available, might equally well belong to the transitional form mentioned

under ssp. albisquama.

Caricea semifumosa group

1(4). Halteres with the knob black or dark brown. Frons subparallel, almost

inappreciably widened to vertex, about half as wide in front as one eye.

Thorax and abdomen conspicuously dusted. 2-4 mm.

2(3). Coxae, trochanters, and antennae entirely black in both sexes. Setae of

femora and tibiae rather short. . ‘ : : [*nodosa Stein]

3(2). Front and mid-legs with the apex of coxae and the trochanters yellow to

whitish in both sexes, the rest of the anterior coxae and the third an-

tennal joint whitish in male. Parafrontalia linear though quite distinct ;

parafacialia linear, jowls hardly as wide as the third antennal joint.

Longest hairs of arista less than one-half as long as the third antennal

joint wide. Scutellum with only two pairs of hairs besides the setae.

albicova Stein, p. 237

4(1). . Halteres with the knob yellow.

5(6). Hind tibiae with three preapical setae (an antero-subdorsal one between the

ad and d preapicals). Fronto-facial stripe narrowed from mouth to

just beyond base of antennae, then subparallel and slightly widening to

vertex, parafacialia at least two-thirds as wide as third antennal joint.

Longest hairs of arista one-and-a-half times as long as the third antennal

joint wide.? Pattern of thorax and abdomen very faint (small paired

spots on abdomen and one or three narrow vittae on thorax). ¢:

Basal third of wings whitish, apical two-thirds brownish; inner half of

each lobe of the fourth ventrite black and shining, somewhat rugose,

outer half opaque bluish-grey dusted. : ‘ [semialba Mall.*]

1“ Coenosia”’ nitidiventris Stein from Madagascar would probably run to this paragraph.

The frons, however, is hardly more than a third as wide as one eye, and thorax and abdomen

are shining black in the species, evidently without any appreciable pruinosity. The halteres

‘“seem to be black’”’ (ex Stein).

2 Arista short-plumose, the longest hairs not quite as long as the third antennal joint

wide. Wings hyaline in both sexes. Tibiae infuscate, but the ground-colour, at least of

the front and mid-tibiae, reddish-yellow: znanis Stein, see humilis group.

3 Two of the typical specimens from Ulundi are males, the other two females. The

female from Willow Grange is not now in the B.M. under semzalba.

MUSCIDAE: B.—-COENOSIINAE

6(5). Hind tibiae with two preapical setae (an ad and a d one).

7(10). Fronto-facial stripe very distinctly narrowed from base of antennae to about

middle of frons, the latter relatively very narrow in middle (Fig. 80).

Parafacialia rather wide, at least half as wide as the third antennal

joint. Arista plumose in basal half, the longest hairs as long as the

third antennal joint wide.

8(9). Last section but one of m four-fifths or at least three-quarters as long as the

last section. ¢: wings whitish in basal half, rather abruptly brown

in apical half, though with vague border; thorax grey dusted without

or with a triangular spot of brown dust in front of and on base of

scutellum; abdomen with a not very conspicuous transverse dark

patch on each segment, somewhat longer than thorax, strongly com-

pressed. 9: wings hyaline, dorsum of thorax with traces of three or five

disintegrated longitudinal vittae, abdomen with a median vitta and a

pair of brown, more or less separated ce at least on some of the

segments. : 5 : . [semifumosa

9(8). Last section but one of m te: fate (3) to about: one-half as long as the last

section, g: wings hyaline throughout, thorax and abdomen, except

the apex of the latter, entirely dark brown dusted on dorsum, darker

brownish-grey on sides than in semifumosa, abdomen distinctly shorter

than thorax. 2 as in semifumosa, the pruinosity slightly darker and

more brownish. : ‘ hyalinipennis sp..n.

10(7). Fronto-facial stripe not meroued fon Base of antennae to middle of HOG

(seldom very slightly narrowed from base of antennae to first inclinate

frontal setae), frons never narrowest in middle.?

11(16). Ocellar bristles short, about one-third as long as the frons from the end of

the lunula to the postvertical setae. Abdomen of male strongly de-

pressed up to apex (nestoy?). Parafacialia narrow, not quite half as

wide as third antennal joint. 1-8—3-2.mm

12(15). Arista not quite as wide, including pubescence, as the third antennal joint.

Front tibiae with a small ad and well developed d and pd preapicals, no

additional d seta present basad of the d, thus with only 2~3 fairly con-

spicuous setae in the apical dorsal part. Abdomen with extended dark

transverse bands,” which occupy almost the whole of the dorsum of the

second and third segments of male, except for a small anterior lateral

triangle of grey dust; in the female these triangles are larger.

13(14). Disc of scutellum setulose. Arista almost as wide, including pubescence, as

the third antennal joint. Frons almost as wide as one eye. Fourth

abdominal segment largely dark dusted on dorsum, similar to the other

segments. Tibiae entirely black or piceous. Fourth ventrite of male

larger, fully as long as the second and third ventrites together, grey

dusted. ‘ : ; : : [nestor

14(13). Disc of scutellum bare, execmt fon eno pairs of setulae and two pairs of

setae. Arista not much more than half as wide, including pubescence,

as the third antennal joint. Frons two-thirds to three-quarters as wide

as one eye. Fourth abdominal segment grey dusted, with a narrow,

235

Stein]

» P» 237

Curr. ]

1 Coenosia fallax Stein from Abyssinia seems to lie near nestor, but is distinguished by

the pattern of the abdomen: a faint median vitta and indistinct paired spots.

2 If the mid-tibiae are almost entirely yellow and the hind tibiae of male beset with

some long, fine, erect hairs on ventral surface, see tvichocnema Stein (humilis group), }

Il, 4 (hk)

236 RUWENZORI EXPEDITION

incomplete, brown median vitta. Tibiae yellowish at base. Fourth

ventrite of male small, slightly longer than the third ventrite, black and

glossy. : : é fascigeva Stein, p. 238

). Arista plumose in basal hale fhe: ieneest nee as long as the third antennal

joint is wide. Front tibiae with the ad preapical seta strong and a

distinct additional d seta on the apical third, thus with four very con-

spicuous setae on the apical dorsal part. Disc of scutellum setulose.

Tibiae entirely black. Abdomen of both sexes largely shining black,

with brownish dust and narrow bands of grey dust along lateral parts

of front margins. Fourth ventrite of male small, black and glossy.

[pilifemur Stein]

16(11). Ocellar bristles strong, about one-half as long as frons, or longer. Abdomen

of male compressed at least at apex. 2-6-5 mm.

17(18). Arista short-pubescent near base, the longest hairs hardly longer than the

basal diameter. Frons slightly dilated to vertex (almost as in the pre-

ceding group, but the parafrontalia narrowly pale dusted). Lower

calyptra in female two-and-a-half, in male five to six times as long as

upper one and (3) more than twice as wide behind middle as the upper

one, reaching third abdominal segment; considerably widened from base

to middle in both sexes. c : . [megalocalyptra’ sp. n.]

18(17). Arista short-plumose, as wide as or vation maclacine pubescence, than third

antennal joint. Frons less dilated, the parafrontalia more broadly pale

dusted. Lower calyptra at most twice as long as upper one, sides sub-

parallel in basal half in both sexes.

19(22). Frons at most three-quarters as wide as one eye. Fourth ventrite of male

entirely dull dusted, or the inner part shining black, in the latter case,

however, the parafacialia narrow. Apex of FR; slightly wider than

middle, its section of the costa about four-fifths as long as that of Rs.

20(21). Hind femora with the normal, single pd preapical. Anterior stpl weaker than

the lower one. Parafacialia much less than half as wide as the third

antennal joint, jowls narrower than third antennal joint. Inner margin

of eyes broadly and evenly rounded in front view. Abdominal spots

not transverse, not fusing with the median vitta. Fourth abdominal

segment of male moderately compressed, not very high, abdomen

oblong in side view, lobes of fourth ventrite black and shining on

interior part. c : : : tvipunctiventris Mall., p. 239

21(20). Hind femora with a second pd soe someuhat basad of the normal pd pre-

apical, Anterior stpl considerably stronger than the lower one, not

much weaker than the posterior one. Parafacialia half as wide as the

third antennal joint or wider. Inner margins of eye practically straight

above and below lunula, thus with a rounded obtuse angle. Abdominal

spots transverse, often fusing with median vitta. g: abdomen strongly

compressed, fourth ventrite entirely grey dusted. hargreavest sp. N., p. 239

22(19). Frons at least as wide as one eye. Parafacialia hardly narrower than the

third antennal joint. Abdomen with three longitudinal spots on seg-

ments 2-4, the median one much narrower. Inner part of fourth

ventrite of male shining black. Apex of R, at most as wide as middle

of this cell, its section of the costa about one-half as long as that of I,.

15(1

tevtviguttata sp. n., p. 241

1 The detailed description of this Abyssinian species will be published in Ann. Mag. nat.

Hist. 1940.

MUSCIDAE: B.—COENOSIINAE 237

Caricea albicoxa Stein

UGANDA: Ruwenzori, Mobuku Valley, 4500. ft. (F.W.E.) 1 g; Kilembe

4500 ft. (F.W.E.) 1 § r 2; Kyarumba, 4500 ft. (D. R. Buxton) 2 9.

Typical locality: TANGANYIKA: Mujenje. Seen from UGANDA: Burunga,

20.v.1926 (G. L. Rk. Hancock).

Caricea hyalinipennis sp. n. ¢ 2

Length 2:7-4:8 mm.; of wings 3°I-4-6 mm.

§. Head (Fig. 80) black, dorsal part of occiput slightly concave, thinly

brown dusted, lower part grey dusted, temples, jowls, face, and parafacialia

densely pale grey dusted, interfrontalia dull black with brown reflections when

seen from in front, parafrontalia ight brown dusted. Frons rounded, hardly

protruding in profile, at most one-half as wide at narrowest point as one eye

(measured, but appearing much narrower), distinctly widening to vertex and

to anterior end, sides somewhat concave, more than twice as long as wide, fronto-

facial stripe evenly widened with straight sides to lower margin of eyes, twice

as wide at base of arista, two-and-a-half times at lower margin of eyes, as frons

at narrowest point; parafrontalia very narrow, a third as wide as interfrontalia,

with about four (3-5) rather small inclinate setae. Inner vertical not very

strong, outer vertical indistinct, postverticals and ocellars small and fine.

Antennae fuscous, second joint interiorly, third entirely brownish dusted, the

latter almost thrice as long as wide, not quite reaching lower margin of eyes

(the latter not quite reaching level of mouth-margin), falling short of mouth-

margin by its width, arista plumose, the longest hairs as long as the third

antennal joint wide, hairs short on apical half. Palpi fuscous. Parafacialia

half as wide, jowls not quite to fully as wide as the third antennal joint.

Thorax black, pleurae grey dusted, dorsum dark brown dusted. acy fine

and small, in two rows before suture, in about four behind suture, 1-+-3 strong

dc, posterior za well developed; scutellum setulose, with two pairs of strong

setae, the lateral pair distinctly shorter. Posterior propleural and upper pro-

stigmatal moderately long and rather fine, lower prostigmatal fine, curved

downward, surrounded by a few small hairs. stf// in an equilateral triangle

which bears a few erect hairs, lower st¢fl fine, anterior one moderately strong,

posterior one very strong.

Abdomen distinctly shorter than thorax, slightly depressed at base, strongly

compressed at apex, dorsum dark brown dusted, somewhat glossy, apex, more

or less narrowly, and venter grey dusted; second segment with a lateral seta,

third and fourth segments each with a transverse row of 2—3 pairs of setae,

fifth with a pair of short and fine setae.

Legs wholly black, the base of the tibiae sometimes narrowly brownish or

238 RUWENZORI EXPEDITION

yellowish. Front femora with 6-7 rather short pd and fv setae, the latter fine

and alternating with smaller ones, setae of the # row very small, front tibiae

with a rather long and fine pv seta between one-half and three-fifths; mid-

femora with 4-6 short a setae in basal half, 3-4 rather short, fine pv in basal

half, and two rather weak preapicals (p and pd), mid-tibiae with a moderate ad

at three-fifths, and a weak fd somewhat beyond middle, hind femora with 6-7

moderately strong ad, a rather fine av at one and at two-thirds, a rather strong

pv at two-fifths, a few small ones near base and beyond it, and a weak pd

preapical, hind tibiae with a rather strong ad slightly beyond middle and a

rather weak a very close to it, at the same level.

Wings hyaline, veins brown at apex, testaceous towards base, 7,,; and m

parallel to slightly diverging at apex, 7-m at two-fifths to two-thirds of discal

cell, the last section but one of m (between 7-m and m-m) two-fifths to slightly

more than half the length of the last section. Calyptrae hyaline, very slightly

smoky, the borders brown, lower one strongly projecting, about twice as long

as upper one, apex broadly rounded. Halteres ferruginous, the knob often

infuscated on disc.

Q. Parafacialia sometimes considerably more than half as wide as third

antennal joint, pollinosity of thorax paler greyish-brown to grey, with five

indistinct or distinct vittae: along the acy, dc and along the prst to inward of

the za, this pattern very variable, the median vitta sometimes abbreviated and

furcate behind suture. Abdomen with a complete or incomplete median vitta

and a pair of rather large, roundish, fuscous spots on first two segments and of

smaller ones on the third and sometimes fourth segments, these spots often

more or less fused with median vitta; abdomen more depressed, not com-

pressed at apex, but pointed. The / row of setae on the front femora often quite

distinct. Border of calyptrae testaceous. Halteres ferruginous, the disc of

the knob much less or not at all infuscated.

UGanpDA: Ruwenzori, Fort Portal, 4.x11.1934 (F.W.E.) 3 type, I ¢ paratype;

Kyarumba, 4500 ft. (D. R. Buxton) 1 2 paratype; Kilembe, 4500 ft. (F.W.E.)

I 2 paratype; Birunga 20.v.1926 (G. L. R. Hancock) 1 3 paratype ; Kampala,

11.x1.1926 (G. L. R. Hancock) 1 9 paratype. KENYA: Chyulu Hills, 3500-5600 ft.,

iv.-v. and vii.1938 (Coryndon Museum Exped.) 2 3 10 9 paratypes (in B.M.

and Coryndon Memorial Mus.). S. RHopEsIA: Salisbury, 14.vi.1937 (Alex.

Cuthbertson) 1 3 4 9 paratypes; I09.iii.1939, 3 2 paratypes (partly returned to

Mr. Cuthbertson).

Caricea fascigera Stein

UcanpDA: Kigezi District, Mt. Muhavura, 7000 ft., 29.ix.1934 (F.W.E.)

2 $ 1 9; Mabungo Camp, 6000 ft., xi.1934 (J. Ford) 5 g 14 9; Mabungo,

6000 ft. (J. Ford) 1 g.3 9. Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 3 2; Mt.

MUSCIDAE: B.—COENOSIINAE 239

Elgon, Butandiga, 7000 ft., 5.vili.1934, sweeping short grass (J. Ford) 1 9;

between Butandiga and Bulambuli, 8000 ft., 7.viii.1934 (J. Ford) 3 9.

Typical locality: TANGANYIKA: Mujenje. In B.M. from S. RHODESIA:

Salisbury, 14.vi.1937 (A. Cuthbertson). Nata: Willow Grange (R. C.

Wroughton). ABYSSINIA.

A female from NATAL: Estcourt, ix.-x.1896 (G. A. K. Marshall) is somewhat

atypical. The arista, stpl, pd preapicals of hind femora, abdominal pattern and

size are as in fascigera, but the ocellar setae and parafacialia as in pygzalis.

Caricea tripunctiventris Mall.

UGANDA: Kalinzu Forest (T. H. E. Jackson) 1 9. S.W. UGANDA: Kigezi

Prov., Lake Mutanda, 6000 ft., xi.34 (J. Ford) i 9.

Typical locality: NyasaALanp: Mt. Mlanje (type g¢ in B.M.). In B.M.

from MASHONALAND: Mt. Chirinda, 3000 ft., 12.vi.11 (C. F. M. Swynnerton) 1 3.

Caricea hargreavesi sp. n. 5

Length 3:1-4:8 mm.; of wings 3°3-4°8 mm.

g. Head black, grey dusted, interfrontalia dull black with shght whitish

and brown reflections when seen from in front, frontal triangle brownish dusted,

parafrontalia, face, jowls and temples densely whitish-grey dusted. Frons

rounded in profile, somewhat protruding at lunula (by at most width of an-

tennae), very slightly (by about a sixth or seventh) dilated with straight sides

from lunula to vertex, two-thirds as wide anteriorly as one eye, more than twice

as long as wide in front; fronto-facial stripe strongly dilated with almost

straight sides from lunula to lower margin of eyes, where it is twice as wide;

parafrontalia well defined, a quarter as wide in front, almost one-half as

wide behind as interfrontalia, with three not very strong inclinate setae (the

second of them weaker). Inner vertical strong, outer vertical absent, post-

verticals not very strong, ocellars strong, at least half as long as frons. Anten-

nae black, second joint interiorly at apex and third joint brownish-grey dusted,

third joint thrice as long as wide, not quite reaching lower margin of eyes (the

latter not quite reaching mouth-margin), falling short of mouth-margin by

width of third antennal joint; arista plumose, the hairs rather long in basal

half, some of them as long as the third antennal joint wide. Palpi fuscous.

Parafacialia one-half as wide or wider, jowls as wide as third antennal joint.

Thorax black, grey dusted, dorsum with a narrow brown median vitta, a

pair of disintegrated vittae along the dc and traces of a somewhat oblique pair

of vittae through the prst and sa pores, scutellum with some faint brownish

spots along base. acy in two rows, rather numerous and not very short, some

additional setulae towards scutellum, 1-+-3 long and strong dc, posterior ta

240 RUWENZORI EXPEDITION

well developed; scutellum setulose with two pairs of strong setae, the lateral

ones conspicuously shorter. Posterior propleural and upper prostigmatal

rather strong, lower prostigmatal rather weak, curved downward, surrounded

by a few fine, erect hairs. stpl in a triangle which bears rather numerous erect

hairs and the posterior side of which is slightly longer than the others, lower

stpl moderately strong, anterior one strong, much longer than the lower one,

the posterior one very strong though only moderately longer and stronger

than the anterior s¢pl.

Abdomen black, grey dusted, first to fourth segments each with a median

vitta and a pair of transverse spots which are often more or less broadly fused

with the median vitta, as long as thorax, oblong-oval, strongly compressed at

apex, second segment with a moderate lateral discal seta, third and fourth

segments each with a transverse discal row of two pairs of strong setae, fourth

besides with a fine lateral marginal seta, fifth with a pair of long and fine setae

and some fine setulose hairs, fourth sternite entirely dark grey dusted.

Legs entirely black and glossy, hardly dusted, except for the coxae which are

dull, grey dusted; the knees, especially the fore knees, sometimes narrowly

yellowish. Front femora with 7-8 rather small fd setae, 3-4 rather long p setae

near base and smaller ones up to apex, and 6-7 long fv, alternating with fine

ones, front tibiae with a long pv seta at almost three-fifths; mid-femora with

4-5 short av up to apex, three long fv in basal half and a smaller one at three-

fifths, and two preapicals (pf and pd), mid-tibiae with a strong ad seta at two-

thirds and a much smaller pd slightly basad of it; hind femora with seven

cather strong ad, a number of small av, two strong ones at one- and two-thirds

and a preapical av, a moderate pv at a quarter, a very strong one at two-fifths,

a moderate one at three-fifths, and two fd near apex, the last of them being the

normal preapical, hind tibiae with a very strong pd at middle and a shorter a

very close to it. >

Wings hyaline, hardly smoky, veins light brown with testaceous bases,

74,5 and m slightly diverging at apex, v-m at slightly beyond middle to three-

fifths of the discal cell. Calyptrae hyaline with pale yellow border, lower one

twice as long as upper one, strongly projecting, sides subparallel with broadly

rounded apex. Halteres pale ferruginous.

similar to male, not differing except by the not compressed, but depressed,

pointed abdomen, the setae of the legs stronger.

UGANDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 3 type, 12 § 14 9 para-

types (one 9 with prey: Sciava sp.); Fort Portal, 4.x11.1934 (P.W.E.) i 3

paratype ; Katosi, 20.1x.1936, on debris at edge of lake (E. G. Gibbins) 1 3g

paratype ; Kampala, 30.1.1927 (H. Hargreaves) I g paratype; Butandiga,

12.1.1930 (H. Hargreaves) 1 3 2 9 paratypes.

A most exceptional character of this species is the double fd preapical seta

on the hind femora.

MUSCIDAE: B. COENOSIINAE 241

Caricea tertriguttata sp. n. ¢ 2

Length 2:6-3-4 mm.; of wings 2-3-3-2mm. |

3g. Head black, densely grey dusted, face, parafacialia, jowls and lower

part of temples silver-white dusted, interfrontalia rather densely grey dusted

with brownish reflections when seen from in front, black only in a strictly dorsal

view. Head similar to that of Coenosia, almost as long as high, frons slightly

rounded in profile, projecting at lunula by width of antennae, subparallel,

slightly wider than one eye, one-half longer than wide, fronto-facial stripe

very slightly narrowed to base of second antennal joint, rather strongly widened

with straight sides from there to lower margin of eyes, where it is a third wider

than at vertex; parafrontalia rather well defined, a quarter as wide in middle

as interfrontalia, with two strong, one moderate and often 1-2 small inclinate

setae. Inner vertical strong, outer one indistinct, postverticals not very

strong, ocellars long and rather strong. Antennae fuscous, second joint silver-

grey dusted interiorly at apex, third joint two-and-a-half times as long as wide,

almost reaching lower margin of eyes (the latter not reaching level of mouth-

margin), falling short of mouth-margin by slightly more than its width; arista

bare on apical third, plumose in basal half, the longest hairs as long as the third

antennal joint is wide. Palpi black. Parafacialia two-thirds to almost as wide,

jowls one-and-a-half times as wide as the third antennal joint.

Thorax black, densely grey dusted, with a rather faint and very narrow dark

brown vitta along the acr, which is triangularly widened before scutellum, and a

disintegrated similar narrow vitta along the dc, scutellum somewhat brownish

dusted on disc. acy small, in two rows which are very close together, a few

additional setulae towards scutellum, 1+3 very strong and long dc, posterior za

distinct though rather small; scutellum sparsely setulose, with two pairs of

strong setae, the lateral ones conspicuously shorter. Propleurals and pro-

stigmatals rather strong, lower prostigmatal curved downward, with 1-2

adjacent minute hairs. s¢p/ in a triangle which bears a few small hairs and the

anterior side of which is considerably shorter, lower s¢f/ moderately strong,

anterior one strong, posterior stp/ very long and strong.

Abdomen black, densely grey dusted, the second to fourth segments each

with an incomplete median vitta and a pair of rather large dark brown spots,

hardly longer than thorax, hardly depressed at base, strongly compressed from

middle to apex, first to fourth segments each with a rather strong lateral discal

seta, third besides with a small and fourth with a strong subdorsal seta, fourth

with two lateral marginals, fifth with a pair of fine, erect setae, fourth ventrite

with numerous, erect, long and partly strong hairs.

Legs black, rather densely grey dusted, tibiae narrowly testaceous trans-

lucent at base; front femora with five rather strong fd setae, three moderately

long p setae near base and small ones up to apex, and six rather long fv,

242, RUWENZORI EXPEDITION

alternating with small ones, front tibiae with a rather strong pv seta; mid-femora

with four a setae in basal two-thirds, the last two of them rather strong, a

distinct a preapical level with the p and fd preapicals, and three long pv in

basal half, mid-tibiae with a strong but not very long pd slightly beyond middle

and a slightly longer ad not much beyond the d; hind femora with five strong

ad setae, some unequal av, among which there are two strong ones at one- and

two-thirds and a rather strong preapical slightly before the level of the pd

preapical, a long pv near base, a strong one at two-fifths and a fine one at three-

fifths, hind tibiae with a strong but not very long ad and a slightly shorter a

close together at middle.

Wings hyaline, hardly smoky, veins brown with testaceous base, 7,,; and

m parallel to slightly diverging at apex, 7—-m slightly beyond middle of discal

cell. Calyptrae whitish with pale testaceous margin, lower one not very large,

about twice as long as the upper one, sides subparallel, rather broadly rounded

at apex. Halteres pale ferruginous.

9 very similar to male, the abdomen somewhat more elongate and depressed,

pointed.

Kenya: Mt. Elgon, heath zone, 10,500-11,500 ft., 11.1935 (F.W.E.) 3 type,

2 $ 2 9 paratypes.

Pygophora Schin.

This genus has, owing to its long frons, short frontal triangle and high

head, more resemblance to typical Coenosiinae than any of the following

genera. Four species have been described from Africa and the Seychelles, but

Coenosia pallipalpis Stein from the Seychelles, the types of which are in the

B.M., is also a Pygophora. In these five species the wings are never spotted

and the legs of the male are entirely pale.

KEY TO THE ETHIOPIAN SPECIES OF PYGOPHORA

1(2). Hind tibiae of male with an apica process on ventral surface. Femora

largely piceous in female. Humeri and_scutellum entirely dark.

Mentum piceous. 2+3 dc. Seychelles. : : é (lobata Stein}

2(1). Hind tibiae of male without an apical process on ventral surface. Legs

entirely pale in both sexes.

3(6). Humeri and scutellum yellow. Mentum yellow.

4(5). Propleural depression yellow. Uppermost frontal seta level with the

ocellars or the anterior ocellus. Mid-femora without a conspicuous

row of dense black setulae on the apical third of the pv surface. Male

with the lateral parts of the fourth ventrite simply angular on dorsal

extremity, with very short and inconspicuous setulae on ventral ex-

tremity. Smaller: 5-6-5 mm. ; : alemella Ség., p. 243

5(4). Propleural depression blackish (though densely wehitighigrey, dusted). Upper-

most frontal seta level with or behind posterior ocelli. Mid-femora with

a conspicuous row of dense black setulae on the apical third of the pu

MUSCIDAE: B.—COENOSIINAE 24

surface; these are increased in number and arranged in a multiple row

in the male. Lateral parts of the fourth ventrite of male pointed and

prolonged on dorsal extremity, and with rather long and conspicuous

setulae on ventral extremity. Larger: 6:5~7 mm. acvomiata Speis., p. 243

6(3). Humeri and scutellum black, grey dusted. Mentum piceous.

7(8). 55-65 mm. Abdomen entirely blackish-grey, not translucent. Third to

fifth tarsal joints very short. Seychelles. 5 : . [pallipalpis Stein}

8(7). 4:5 mm. Abdomen yellowish translucent at base. : [*parvipuncta Stein]

Pygophora alemella Ség.

Séguy, 1938, Miss. scient. Omo 4, Zool.: 368,

UcanpA: Ruwenzori, Nyamgasani Valley, 6400 ft. (D. Rk. Buxton) 1 3

I 9; Namwamba Valley, 6500 ft. (F.W.E.) 2 9; Kilembe, 4500 ft. (F.W.E.)

1 9. Budongo Forest, 7.-8.i1.1935 (F.W.E.) 2 3 3 &.

Typical locality: KENyA: Mt. Elgon, eastern slope, Elgon Saw-mill, 2470 m.

Though Séguy does not compare his species with the known African forms,

it is probable that his description refers to the present species, because the apical

half of the mid-femora is evidently devoid of the setulae characteristic of

acromiata, and the hypopygium is described as yellow without mentioning

another colour for the inferior forceps, which is black in acromzata but yellow

in the present form. Unfortunately the type is not accessible owing to the

war conditions, as M. Séguy was kind enough to inform me.

Pygophora acromiata Speis.

Speiser, 1910, Wiss. Ergebn. Sjéstedts Kilimandjaro—Meru Exp. 2, Diptera: 159 note

(nom. nov. pro Coenosia humeralis Stein 1906, non Wied. 1830).

UcGanpDA: Ruwenzori, Kilembe, 4500 ft. (F.W.£.) 1 § 1 9.

Typical localities: TANGANYIKA: Langenburg (Lake Nyasa), N. CAMEROON,

(? Toco: Bismarckburg). In B.M. from S. NicErta (Abo), N. NicErta (Ibi),

AsHANTI (Obuasi), Port. East Africa: E. of Mt. Mlanje, 3.-7.x.1913 (S. A.

Neave). UGANDA: Bwamba Forest, Semliki Valley, 2300-2800 ft., 3.-7.xi.1g11

(S. A. Neave). Kenya: Embu, Rugendo’s, 4400 ft., 14.vi.1g14 (C. Sz. J. O.

Browne); Ruiru, 6.vii., 13.vil., 19.vili.1935, on coffee (H. C. James); Kiambu,

15.Vii.1932, on coffee (H. C. James). I sawaspecimen from UGANDA: Kampala,

4.x.1934 (H. Hargreaves), mounted with its prey, which has been identified by

Dr. F. W. Edwards as a male of Sczava flavoscutellata End. ?

Though Coenosia humeralis Wied. was moved to Atherigona by Stein in 1902,

humeralis Stein 1906 as a secondary homonym can unfortunately not be used

and must be replaced by Speiser’s nom. nov., as the rules stand at present.

Pectiniseta obscura sp. n. 3

Length 4:0 mm.; of wings 3°3 mm.

Head one and a half times as wide in front view as high, black, dark brown

244 RUWENZORI EXPEDITION

dusted, face, jowls and lower part of occiput reddish-grey translucent, grey

dusted. Frons slightly wider anteriorly than one eye, very slightly narrowed

to vertex, parafrontalia not distinguished by paler dust, with two inclinate and

two reclinate pairs of frontals, the anterior reclinate seta much longer than

the posterior one, fronto-facial stripe very slightly narrowed from base of

antennae almost to vibrissae. Inner verticals very strong, outer verticals weak,

postverticals rather strong, ocellars strong, reaching anterior end of frons.

Antennae inserted at dorsal third of eyes, fuscous, third joint long, almost

reaching mouth-margin (which is distinctly passed by the lower margin of

eyes), covered with fine white, very conspicuous pile, arista bare beneath, long-

pectinate above, the hairs about half as long as the third antennal joint. Palpi

yellow, slightly long clavate, mentum dark-brown, jowls about two-thirds as

wide as third antennal joint.

Thorax piceous-brown, densely dark brown dusted on dorsum, grey dusted

on pleurae. prst acy in three irregular rows, the outer ones longer, post acy

similar but more numerous towards scutellum, 2 ++ 3 dc, the first frst and post

much shorter and finer than the others, the first frst closer to the second prst

than the latter to the suture. Two weak 7a, prealar absent. Scutellum setulose

and with two pairs of strong setae. Upper prostigmatal seta slightly stronger

than upper propleural, lower setae hair-like. st¢p/ in a triangle, the posterior

side of which is somewhat shorter and which bears some fine hairs, posterior

stpl strong, the other two fine though quite long, the lower one somewhat

shorter.

Abdomen subconical, narrowed at base, strongly convex, slightly curved

downward, dorsum uniformly dark brown, brown dusted, base, apex and

ventral surface of abdomen grey dusted, ventral part of first segment testaceous,

translucent, margins of the other segments narrowly yellowish-translucent ;

ventrites reddish-grey with translucent-yellowish margins, dull, greyish dusted.

Lateral setae of first to fourth segments distinct, hairs rather long, half-appressed,

fifth segment with a pair of fine marginal setae and some small hairs. Superior

forceps fused, triangularly pointed, lobes of fourth ventrite deeply emarginate

at apex, so that the apex of their inner margin projects backward as a short

spine, which is shining, as is the apical half of the inner margin.

Legs entirely testaceous (hind legs missing), femora with numerous fine,

erect hairs, front femora with a brown shadow along dorsum and about five

long pv setae, front tibiae without a submedian seta, with three long preapical

setae on the same level (d, pd and #), ventral surface rather densely clothed

with short appressed setulae towards apex, ventral surface of metatarsus densely

beset with similar setulae, whereas the other tarsal joints are soled with a

white pile. Mid-femora with three moderate a setae near middle and a pd

preapical seta, mid-tibiae with a long, fine submedian fd, mid-tarsi not soled.

Wings smoky, the veins brown, 7,,; and m slightly but very distinctly

MUSCIDAE: B.—COENOSIINAE 245

converging at apex, y-m very short, just beyond middle of discal cell, sixth vein

reaching one-third of the distance to margin. Calyptrae brownish-hyaline with

brownish margin and yellowish fringe. Halteres yellow.

UGANDA: Ruwenzori, Bwamba Pass (west side), 5500-7500 ft. (F.W.E.)

3 type.

P. prominens Stein 1910, the only species of Pectiniseta known from the

Ethiopian region, is distinguished from the present species, according to Stein’s

description, by black coxae and femora (except for the extreme apex), whitish-

grey dusted frons, pale cinereous-grey pruinosity of thorax and scutellum, and

the greyish dust of the abdomen, on the second to fourth segments of which a

median vitta and a pair of dark spots are distinct in posterior view. The

sternites are glossy black in prominens, 7,4, and m diverge slightly, the sixth

vein reaches middle, and the calyptrae are white. P. prominens is not in

the B.M.

Lispocephala Pok.

Three species have been listed from the Ethiopian region, and I have the

same number of species before me. Besides, Caricea sexnotata Ad., a type of

which I have seen, proves to be a Lispocephala and a synonym of the form

known as L. miki Strobl. The characters of colour are evidently rather variable

in this genus, and a thorough study of a richer material is wanted. The three

Ethiopian forms at hand may be distinguished by the following key.

1(2). Thorax with a pair of brown vittae along the dc. Paired spots of second to

fourth abdominal segments large and subquadrate, deep black, con-

spicuous even in dorsal view, first abdominal segment not translucent.

Fourth ventrite of male grey and dusted, its ventral half with a vertical

row of four very strong, closely adjacent black setae, which are con-

spicuously stouter than the marginal setae of the dorsum. Superior

forceps spiniform, extremely short, inferior forceps paired, elongate

claw-shaped, about five times as long as the superior forceps. (Frons

distinctly and rather broadly reddish at anterior margin. Front coxae

and basal half of front femora often grey. Third antennal joint light

orange in male, infuscate with light orange base in female.)

squamifera Stein, p. 246

2(1). Thorax without vittae. Paired spots of second to fourth abdominal seg-

ments small, oval, conspicuous and rather black only in posterior view.

Fourth ventrite of male testaceous and shining near apex, with few,

sparse, fine setae and hairs, the strongest among them being considerably

finer than the marginal setae of the dorsum. Superior forceps spiniform,

of moderate length, inferior forceps paired, hardly longer than the

superior forceps. (Frons not or very narrowly reddish at anterior

margin. Front coxae and front femora always yellow, except for a

small greyish patch at the base of the coxae.)

1 See note p. 103; the species was redescribed by Stein in 1910 (Ann. Mus. nat. Hungar.

8: 564) and briefly in 1917 (Arch. Nat. gesch. 83 A 1: 145).

246 RUWENZORI EXPEDITION

3(4). Antennae entirely light orange. First abdominal segment usually (and

second sometimes) translucent yellowish laterally and ventrally.

Inferior forceps rather broad near base, emarginate interiorly near

apex, the latter pointed and curved downward and inward.

miki Strobl, p. 246

4(3). Third antennal joint fuscous, with the base narrowly light orange. Base

of abdomen not translucent yellowish. Inferior forceps parallel and

depressed, simply rounded at apex. j : : : [africana Mall. |

Lispocephala squamifera Stein

Ucanpba: Kigezi District, Mabungo Camp, 6000 ft., 24.x1.1934 (J. Ford)

I 9; Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 1 9.

Typical locality: NatTaL: Durban.

These two females have the front coxae and basal half of the front femora

grey like the typical couple. I have seen two males and three females from

Kenya: Chyulu Hills, 5600 ft., vi. and vii.1938 (Coryndon Museum Exped.),

of which only one female has the typical coloration of the front legs, whereas

the others have the front femora entirely and the front coxae more or less

largely yellow. There is some slight doubt as to the identification of the

species, as Stein describes the ‘‘ Bauchlamellen”’ (ventrites) as bearing at their

apex the black, backward pointing, closely adjacent setae which make them

appear “‘feathered’’. These words are, however, contained in the sentence

describing the hypopygium, and I assume that the lateral lobes of the fourth

ventrite only are meant by “‘Bauchlamellen”’.

Lispocephala miki Strobl

n. syn. Cavicea sexnotata Ad. 1905, Kansas Univ. Sci. Bull. 3: 205.

Kenya: Mt. Elgon, Kapretwa, 6500 ft., ii.1935 (F.W.E.) 2 3; Aberdare

Range, Mt. Kinangop, 8000 and gooo ft., x. and 1.xi.1934 (F.W.E.) 4 35 9.

UcanpDA: Ruwenzori, Mt. Karangora, ggoo ft. (F.W.E.) 1 g; Namwamba

Valley, 6500 ft. and 10200 ft. (F.W.E.) 1 g 2 9.

Typical locality: Datmatia, Zara; reported from EUROPE, CANARY

IsLANDS, ALGERIA, SYRIA and AByssINIA. In B.M. from the latter country.

The type of Caricea sexnotata Ad. (typical locality S. RHODESIA: Salisbury)

has the antennae and front coxae entirely yellow, and the other characters

are also those of L. miki. The Ruwenzori specimens are slightly larger than

the others, but the genitalia do not differ.

Brevicosta africana Mall.

UGANDA: Ruwenzori, Nyamgasani Valley, 6400 ft. (D. R. Buxton) 4 33 9;

MUSCIDAE: B.—COENOSIINAE 247

Namwamba Valley, 6500 ft. (F.W.E.) 2 3 2 9; Kigezi District, Mabungo

Camp, 6000 ft., xi.1934 (J. Ford) 1 9; Masaka, 13.xi.1934 (F.W.E.) 1 9.

Typical locality: S. NiceRrA, Zungeru (in B.M.).

The male is new to science. Its characters are the same as those of the

female. The abdominal segments (Fig. 81) are very much shortened on the

ventral surface, so that the apex of the third ventrite is at about basal third of

abdomen, the fourth extremely long and very deeply cleft, so that it forms two

narrow, long testaceous lobes along the ventral surface of the abdomen, extend-

ing to beyond level of dorsal hind margin of fourth segment, fifth segment much

longer dorsally than fourth, pointed, similar in shape to the female fourth

Fic. 81.—Abdomen of Brevicosta afvicana Mall. 6, lateral view.

segment. The wings of the male are slightly and often not very distinctly

clouded along apex of 7,3, this being very distinctly the case even in one

female (from Masaka). E

Orchisia costata Meig.

UGANDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 2 3; Namwamba Valley,

8300 ft. (F.W.E.) 1 g; Kigezi District, Mabungo Camp, 6000 ft., 18.x1.1934

(J. Ford) i 3.

Typical locality unknown; widely distributed: S. Europe, Cuina, For-

mMosA, Fijt (B.M.), SAmMoa (B.M.). The species has been reported from

MADAGASCAR and many places in AFRICA: S. RHODESIA, NATAL, TANGANYIKA,

UGANDA. In B.M. from AByssINIA, GAMBIA and KENYA.

Spanochaeta dorsalis v. Roser

UGANDA: Ruwenzori, Kilembe, 4500 ft. (F.W.E.) 1 3.

This single African specimen at hand shows some slight differences from

British examples, the hairs of the arista being appreciably longer and the

248 RUWENZORI EXPEDITION

abdomen somewhat shorter (though very similar). More material will be

necessary before any conclusion should be drawn from these deviations.

Amicitia gen. n.

Frons subequal in both sexes, broadly separating the eyes, with two pairs

of reclinate frontal setae, the anterior one of them stronger; frons plane,

strongly protruding in profile at base of antennae, the frontal triangle not

passing middle. Thorax with 2 + 3 dc, of which, however, the first is always

much smaller than the second and sometimes indistinguishable, often the

second prst and first post de are also weak. Scutellum with the apical setae

strong, the lateral pair weak or missing. Two rather strong propleural and

prostigmatal setae, the lower prostigmatal curved downward. Three stpl in

an almost equilateral triangle, or the anterior one absent. Abdomen slender,

parallel. Front tibiae with a fd submedian seta, mid-tibiae with a single sub-

median seta (on pd surface), hind tibiae (Fig. 82) with a moderate and a strong

ad seta, a moderate av seta and one (males) or two (females and some males),

pd setae. Costa without a spine, reaching apex of m, all the other veins bare,

sixth vein not much passing middle of the distance to margin. Lower calyptra

longer than upper one.

Genotype: Amicitia lucens sp. n.

By Séguy’s key the species would be run to the American genus Phyllogaster

Stein, from which they differ, however, by two pairs of reclinate frontal setae,

Phyllogaster having apparently only one pair of reclinate frontal setae (according

to figure in Curran, 1924, Fam. Gen. N. Amer. Diptera, p. 388, Fig. 6). It is,

moreover, evident from Curran’s key (l.c. p. 396-399) that the stfl are placed

as in Mydaeini, not in an almost equilateral triangle. Thus, it does not seem

that Amicitia is closely related with Phyllogaster. In the latter the mid-tibiae

have two pd setae and the ovipositor of the female is adorned with two or four

thorns, protruding behind the fourth segment, which are not present in Amucitia.

The species of Amucitia differ rather strikingly among each other, but they

are, nevertheless, closely related.

The genus is dedicated to friendship, one of the most noble and unselfish

sentiments of mankind, and to all the kind friends, who have given me their

sympathy, encouragement and help in these last seven years after my unprovoked

misfortune.

KEY TO THE SPECIES OF AMICITIA

1(4). 1-+3 strong dc present. Lateral scutellar setae small, less than half as long

as the first two post dc, but distinct. Vittae of thorax brown-pruinose.

Hind tibiae with one or two rather weak pd setae in both sexes, not

densely hairy. Mid-femora with a p and a pd preapical seta.

MUSCIDAE: B.—COENOSIINAE 249

2(3). Front coxae and all the femora yellow, only the hind femora in male with a

slight brown shadow on apex of dorsal surface, in female with the

apical quarter piceous on dorsum. Anterior, paramere oi male strongly

curved, claw-shaped. : c modesta sp. D., p. 249

3(2). Front coxae and femora black except ay a mall apical area, apical half to

third of mid and hind femora black. Anterior paramere very slightly

curved, spine-shaped. . 5 seclusa, sp. N., p. 251

4(1). Only one or two fairly strong post dc present! the each post pair and two prst

pairs very weak. Lateral scutellar setae lacking or hair-like. Vittae

of thorax black and shining. Hind tibiae of male densely beset with

long erect hairs (Fig. 82). Mid-femora with a single p preapical seta.

5(6). Third joint of arista hardly longer than third antennal joint, half as wide,

including plumosity, as third antennal joint. Lateral scutellars absent.

pd setae of hind tibiae (Fig. 82) not distinguishable in male, amongst the

pd hairs, many of which are much longer than the pv hairs. Mid-

metatarsus of moderate length, not longer than the hind metatarsus,

piceous; front metatarsus with short hairs all round. Front and mid-

femora with less than apical half piceous on dorsal surface. Jucens, sp. n., p. 252

6(5). Third joint of arista more than twice as long as third antennal joint, bare on

ventral surface, the longest hairs of dorsal surface not longer than the

diameter of the arista at basal quarter. Lateral scutellars hair-like.

pd setae of hind tibiae distinct, even in male, the longest pd hairs being

much shorter than the pv hairs. Mid-metatarsus distinctly longer than

the hind metatarsus, front metatarsus with a fringe of long erect p hairs.

Front and mid-femora with the entire dorsal surface piceous.

insignis, Sp. D., Pp. 254

Amicitia modesta sp. n. ¢ 9

Length 3°6-5:1 mm.; of wings 4:0-4:8 mm,

3. Head black, densely grey dusted, interfrontalia black with whitish

reflections, face and parafacialia silver-white dusted. Frons slightly wider at

base of antennae than one eye, distinctly narrowed to vertex, protruding in

profile by the width of the third antennal segment; fronto-facial stripe almost

evenly and quite slightly dilated to mouth-margin, face as long as frons.

Parafrontalia well defined, with two pairs of inclinate setae, the posterior one

much weaker. Outer vertical very weak, inner vertical very strong, post-

verticals rather strong, ocellars strong. Antennae fuscous, very long, practically

reaching mouth-margin which is level with the lower margin of the eyes, third

joint more than thrice as long as wide, arista short-plumose, not much more

than half as wide, including plumosity, as the third antennal joint. Palpi

fuscous in apical half, testaceous in basal half. Jowls narrower than third

antennal joint.

Thorax black, densely grey dusted, the pleurae bluish-grey dusted in places,

dorsum with a pair of rather broad vittae of brown dust from the first de to the

lateral setae of scutellum. acy setulae in two rows, with a few additional

250 RUWENZORI EXPEDITION

setulae on posterior half of postsutural part. 2 + 3 dc, the first about half as

long as the others and closer to the second than this one to suture, lateral

setae of scutellum weak, about a third as long as apical pair. stpl in an almost

equilateral triangle, the posterior side slightly shorter, the area of the triangle

usually quite bare (with a few fine erect hairs in the specimen from Mt. Sabinio),

anterior s¢p/ rather weak to moderate, lower one slightly stronger, posterior one

very strong.

Abdomen slender, subcylindrical, very slightly depressed at base, very

slightly compressed at apex, black, densely grey dusted, with a narrow median

vitta on segments I-4 and a pair of conspicuous black, rounded spots on seg-

ments 2-4 (I-4 in specimen from Mt. Sabinio); first to third segments each

with a pair of strong lateral setae (and sometimes a smaller pair more at sides),

fourth with a discal and marginal row, each of two to three pairs of strong

setae, fifth with two pairs of quite conspicuous setae. Hypopygium with

erect, somewhat wavy, bristly hairs, lateral lobes of fourth ventrite long and

broad, the apex prolonged ventrally in form of a rounded, backwardly

directed lobe which bears some strong setae at apex, the inner surface and

the margins of the lateral lobes densely beset with fine, erect, rather short,

black hairs.

Legs testaceous, mid and hind coxae black, the apex of the hind femora

with a brown shadow on fd and d surfaces, tarsi infuscate, the apical joint, at

least of the front tarsi, paler. Front femora with six rather strong pv setae

(which alternate with small ones), basal half with a row of six fine short av

setae, front tibiae with a moderate pv seta, mid-femora with about 6-7 av

setae from base to beyond middle, becoming gradually longer, an a seta just

beyond the last av, an a, pd and p preapical, about seven fv setae and three

very small ones at apex, mid-tibiae with a moderate pd, hind femora with about

seven av and ad setae, the latter mainly confined to the basal two-thirds, the

last ad, a preapical, level with the fd preapical, hind tibiae with one or two

small pd, a small and a strong ad and a rather strong av beyond the second ad

and pd which are inserted almost level with each other.

Wings slightly smoky, veins brown with testaceous base, 74,; and m sub-

parallel at apex (in one specimen from Namwamba Valley slightly diverging

at extreme apex), 7m almost at middle of discal cell, distinctly beyond end of

v,. Calyptrae whitish, with whitish fringe and yellowish-white border, lower

calyptra strongly projecting beyond upper one, more than twice as long, sides

subparallel in middle part. Halteres yellow.

2. Arista as wide, including plumosity, as third antennal joint. Abdomen

fusiform, more depressed. Mid-femora also with a slight brown shadow at

apex of dorsal surface.

UGANDA: Ruwenzori, Namwamba Valley, 8300 ft. (F.W.E.) g type, 1 3

paratype, I 2 paratype. Kigezi District, Mt. Sabinio, 8000 ft., x1.1934 (.W.E.)

MUSCIDAE: B.—COENOSIINAE 251

I ¢ paratype. The three specimens from the Namwamba Valley are at the

upper limit of the size, whereas the Kigezi specimen is at the lower limit.

Amicitia seclusa sp. n. 3

Length 4:0-4-7 mm.; of wings 3:-4—3°8 mm.

Head black, densely grey dusted, interfrontalia black with whitish reflec-

tions, face and parafacialia silver-grey dusted. Frons slightly wider at base of

antennae than one eye, distinctly narrowed to vertex, protruding in profile

by the width of the third antennal joint, fronto-facial stripe almost evenly and

very slightly dilated to mouth-margin, face as long as frons. Parafrontalia

well defined, with two pairs of inclinate setae, the posterior one much weaker.

Outer vertical very weak, inner vertical very strong, postverticals rather

strong, ocellars strong. Antennae fuscous, very long, practically reaching

mouth-margin which is level with the lower margin of eyes, third joint more

than thrice as long as wide, arista short-plumose, not much more than half as

wide, including plumosity, as the third antennal joint. Palpi fuscous. Jowls

narrower than the third antennal joint.

Thorax black, densely grey dusted, dorsum with a pair of rather broad

vittae of brown dust from the first dc to the lateral seta of scutellum. acy

setulae in two rows with a few additional setulae on median line and behind,

2 + 3 dc, the first one about half as long as the others and closer to the second

than this one to suture, lateral seta of scutellum weak, about a third as long

as the apical pair. s¢p/ in an almost equilateral triangle, the posterior side

slightly shorter, the area of the triangle bare or with one or two fine, erect

hairs, anterior stf/ moderate, lower one considerably stronger, posterior one

very strong.

Abdomen slender, subcylindrical, very slightly depressed at base, very slightly

compressed at apex, black, densely grey dusted, with a narrow, faint median

vitta on segments I-3 and a pair of conspicuous, black, rounded, somewhat

oblong spots on segments 2-4 and a less distinct, smaller pair on the first;

first to third segments each with a pair of strong lateral setae (and sometimes

a smaller pair more at sides), fourth with a discal and marginal row, each of

2-3 pairs of strong setae, fifth with two pairs of conspicuous setae, the inner

one weaker. Hypopygium with erect, somewhat wavy, bristly hairs, lateral

lobes of fourth ventrite black and shining, with the dorsal free part hyaline

testaceous, long and broad, the apex prolonged ventrally in form of a rounded,

backwardly directed lobe which bears some strong setae at apex, the inner

surface and the margins of the lateral lobes densely beset with fine, erect,

rather short, black hairs.

Legs testaceous, all the coxae, front femora (except for the extreme apex),

and apical half to third of mid and hind femora black, tarsi somewhat infuscate,

II, 4 (1)

252 RUWENZORI EXPEDITION

less so at apex. Front femora with six rather strong pu setae, which alternate

with small ones, front tibiae with a moderate fv seta, mid-femora with 6-7 av

setae from base to beyond middle, the last two or three somewhat longer, an

a seta just beyond the last av on boundary of pale and dark colour, an a, pd

and p preapical seta, about seven pv setae, about four of them on less than

apical half, and about three very small ones at apex, mid-tibiae with a moderate

pd seta, hind femora with about eight av and ad setae, the last of them.a pre-

apical, the last ad level with the fd preapical, hind tibiae with a small or a small

and a moderate fd, a small and a strong ad and a rather strong av beyond the

second ad and pd which are inserted almost level with each other.

Wings somewhat smoky, veins brown with testaceous base, 7,,; and m

subparallel at apex, very slightly diverging at extreme apex, v—m almost at

middle of discal cell, distinctly beyond apex of 7,. Calyptrae whitish with

yellowish-white fringe and border, lower calyptra strongly projecting beyond

upper one, more than twice as long, subparallel in middle. Halteres reddish-

yellow.

UGANDA: Kigezi District, Mt. Sabinio, 8000 ft., xi.1934 (F.W.E.) 3 type;

Kanaba, 7300-7800 ft., xi.1934 (F.W.E.) 3 $ paratypes.

This form is almost identical with modesta, except for the great difference in

the coloration of the legs and the slight difference in the genitalia described in

the key. One would therefore be inclined to regard these specimens as a sub-

species, but this theory is not supported by the fact that on Mt. Sabinio modesta

was also found.

Amicitia lucens sp. n. 5 2

Length 4:6-5°5 mm.; of wings 4:3-4:8 mm.

3g. Head black, dorsal half of occiput sparsely grey dusted, lower part of

occiput, temples, parafrontalia, parafacialia, facial ridges and jowls densely

grey to silver-grey dusted, face and interfrontalia dull black, with brownish

reflections. Frons as wide anteriorly as one eye, strongly narrowed to vertex

with almost straight sides, plane, projecting in profile beyond eyes by more

than one and a half times the width of the third antennal joint, fronto-facial

stripe very slightly and broadly sinuate between base of antennae and vibrissae,

almost parallel, slightly dilated from above vibrissae to lower margin of eyes.

Parafrontalia well defined, more than two-thirds as wide in front, almost half

as wide in middle as interfrontalia, with two pairs of inclinate frontal setae,

the posterior one much weaker. Outer vertical very weak, inner vertical very

strong, postverticals and ocellars rather weak. Antennae fuscous, very long,

reaching the mouth-margin which is level with the lower margin of eyes, third

joint four times as long as wide, arista elbowed, its third joint hardly longer

than the third antennal joint, very short-haired on most of its length, the

MUSCIDAE: B.—COENOSIINAE 253

longest hairs being as long as the second joint of arista is wide. Palpi fuscous,

slightly paler near base. Jowls narrower than third antennal joint.

Thorax black, pleurae rather densely grey dusted, mesonotum glossy with

a slight median vitta of sparse pollinosity, humeri and sides of mesonotum

rather ‘densely dusted, the ensuing two broad, black vittae very slightly

brownish-dusted, as is the area in front of the scutellum. acy hairs very fine

and inconspicuous, in two rows, with single additional hairs behind. 2-3 dc,

of which only the last two pairs are well developed, the first prst dc sometimes

not distinguishable; scutellum with some fine short hairs, lateral seta not

distinguishable, apical setae long and strong. s¢f/ in a triangle, the posterior

Nii

jE FE <

Rae RE eno

FS =e — 3 << x Saoes

SEEK SSS

82.

Fic. 82.—Left hind tibia of Amicitia lucens sp. n., lateral view.

side of which is somewhat shorter, area with single fine hairs, anterior stp/

weak, lower one rather strong, posterior one very strong.

Abdomen slender, long, subcylindrical, somewhat depressed at base, slightly

compressed at apex, black and shining, sides of base of first segment, a narrow

hind margin, and an inconspicuous, somewhat broader front margin on each

segment grey dusted; first to fourth segments each with a conspicuous lateral

seta rather close to base, fourth besides with a marginal row of four strong

setae, fifth with one pair of long and fine setae. Lateral lobes of fourth ventrite

vertical, translucent testaceous, glossy and bare.

Legs yellow, coxae dusted, mid and hind coxae mainly black in ground-

colour, front femora with a piceous streak on dorsal surface of less than apical

half, mid-femora with a similar streak on approximately apical half, hind femora

on apical quarter. Middle of tibiae more or less infuscate, tarsi piceous. Front

femora with about seven fd and five rather short pv (alternating with very

small ones), the last one stronger, developed as a preapical seta, front tibiae

with a rather long submedian fv; mid-femora with three rather strong (the

middle one very strong) pv and a single p preapical, mid-tibiae with a fine

submedian pd; hind femora and tibiae gently though distinctly curved, hind

femora with 5~7 ad, the last one a strong preapical seta, five a, of which the last

254 RUWENZORI EXPEDITION

one is very strong and is followed by an av preapical seta, and a fine fd preapical

seta, hind tibiae (Fig. 82) with two long ad, about four long av, about ten very

long and erect ventral setae, the longest of which are at least a quarter as long

as the tibia, # surface clothed with very fine, erect, short hairs, pd surface with

two longitudinal rows of very long, erect setae, the less dorsal ones finer,

shorter and curled at apex.

Wings hyaline, with brown veins, very slender, 7,,; and m very slightly

converging, m almost inappreciably upcurved shortly before apex, y-m almost

at middle of discal cell. Calyptrae rather small, the lower one subtriangular

with rounded apex, though distinctly projecting beyond upper one, hyaline

with yellowish border. Halteres testaceous.

9. Antennae somewhat shorter and narrower. Abdomen elongate-fusiform.

Hind tibiae without erect hairs, but with two conspicuous ad, pd and one or

two av setae.

UcanpDA: Kigezi District, Kanaba, 7300-7800 ft., x1.1934 (F.W.E.) 3 type,

3 3 paratypes, 4 2 paratypes.

Amicitia insignis sp. n. 5

Length ca. 5°5 mm.; of wings 5°9 mm.

Head black, grey dusted, interfrontalia dull black in dorsal view, but the

whole frons, including interfrontalia, and face silver-white dusted. Frons as

wide anteriorly as one eye, slightly narrowed to vertex, with almost straight

sides, plane, projecting in profile beyond eyes by more than one and a half

times the width of the third antennal joint, fronto-facial stripe gradually

and extremely slightly dilated to ventral margin of eyes, almost inappreciably

sinuate between base of antennae and vibrissae. Parafrontalia well defined in

dorsal view, two thirds as wide in front, a third as wide in middle as inter-

frontalia, with two rather weak pairs of inclinate setae. Outer vertical very

weak, inner vertical strong, postverticals and ocellars rather weak. Antennae

fuscous, very long, almost reaching the mouth-margin which is level with the

lower margin of eyes, third joint more than four times as long as wide, arista

very long, its third joint more than twice as long as the third antennal joint,

bare on ventral surface, with very sparse and extremely short hair on dorsal

surface, the longest hairs not longer than the diameter of the arista at basal

quarter; second joint of arista longer than the third antennal joint is wide.

Palpi fuscous, the basal half brown. Jowls narrower than the third antennal

joint.

Thorax black, pleurae rather densely grey dusted, mesonotum glossy black

with a moderately broad median vitta of grey dust and the humeri and sides

of mesonotum rather densely grey dusted, the median vitta not very sharply

defined and continued to apex of scutellum. acy hairs very fine, in two rows

MUSCIDAE: B.—COENOSIINAE 255

before suture, a few additional hairs behind. dc very fine and small, only the

last post pair long and moderately strong, the other two post de and the second

prst pair very fine and small, but distinct, anterior frst pair not distinguishable

among the recurved small dc hairs; scutellum with a pair of small lateral sub-

basal hairs, a pair of lateral hairs and long apical setae. Only two stl present,

the anterior one absent, the lower one very long and strong, strictly ventrad of

the posterior one.

Abdomen slender, long, curved downward, rather depressed at base, some-

what compressed at apex, black and shining, sparsely grey dusted, with a

broad though not very conspicuous median vitta on the first three segments,

the fourth segment thinly and evenly dusted; first segment with some very

long lateral setae, second and third without setae, fourth with one discal pair

before middle and one marginal pair, both on lateral part of dorsum, fifth

with two pairs of setae, the more dorsal one smaller; hair of abdomen rather

long, fine and erect, especially on lateral and ventral surfaces.

Legs yellow, extremely long and slender, with very few setae, hind femora

and tibiae slightly though distinctly curved, coxae dusted, mid and hind coxae

mainly black in ground-colour, front and mid-femora with a complete piceous

streak along dorsum, hind femora with a similar streak on apical half of dorsum,

the last two joints of the tarsi blackish, the apex of the first to fourth joints

narrowly yellowish-white, hind metatarsus slightly infuscate, rest of hind tarsi

missing. Front femora with rather fine though distinct setae in pd and p rows,

only with two pv setae in basal half, none in apical half, front tibiae with a

moderate fv seta; mid-femora with three strong ad setae in basal half, one

strong pu near base, and a single # preapical, mid-tibiae with a fine fd seta,

mid-metatarsus extremely long (slightly longer than mesonotum without

scutellum) ; hind femora with a pv basal seta and two ad near apex, without a

preapical seta, hind tibiae with two fd, one ad halfway between them, a small

and a longer av between them (and a longer av just basad of the d preapical),

apical half of ventral surface with about nine very long and strong, erect setae

(of about a quarter of the length of tibiae), / and fd surfaces clothed with fine,

erect setae, which are curled at apex, and the longest (most dorsal) ones of

which are about a sixth as long as the tibia.

Wings slender, hyaline, the veins brown, 74, and m distinctly converging

towards apex, rather conspicuously sinuate immediately before apex, and the

extreme apices even inappreciably diverging (this character will probably be

somewhat variable), y-m at middle of discal cell. Calyptrae hyaline, border

yellowish, lower calyptra strongly projecting, largely subparallel. Halteres

pale yellow.

UGANDA: Kigezi District, Mt. Sabinio, 8000 ft., xi.1934 (*.W.E.) 3 type.

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