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Baehr, Martin:

Revision ofthe Pseudomorphinae of the Australian region / Martin Baehr. - München : Pfeil

(Spixiana : Supplement; ...)

2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman,

Paussotropus Waterhouse and Cryptocephalomorpha Ritsema :

taxonomy, phylogeny, zoogeography ; (Insecta, Coleoptera, Carabidae)

(Spixiana : Supplement ; 23)

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oPIAIAN

Zeitschrift für Zoologie

Supplement 23

ssamm

Revision of the Pseudomorphinae

of the Australian Region

2. The genera Pseudomorpha Kirby, Adelotopus Hope,

Cainogenion Notman, Paussotropus Waterhouse,

and Cryptocephalomorpha Ritsema.

Taxonomy, phylogeny, zoogeography

(Insecta, Coleoptera, Carabidae)

Martin Baehr

Gedruckt mit Unterstützung der Deutschen Forschungsgemeinschaft

Verlag Dr. Friedrich Pfeil «» München, 01. April 1997 + ISSN 0177-7424 « ISBN 3-931516-20-2

Adelotopus rufozonatus, spec. nov. from near Edith Creek, far Northern Territory.

SPIXIANA | Supplement 23 München, 01. 04. 1997 | ISSN 0341-8391 | ISBN 3-931516-

Revision of the Pseudomorphinae of the Australian Region

2. The genera Pseudomorpha Kirby, Adelotopus Hope,

Cainogenion Notman, Paussotropus Waterhouse,

and Cryptocephalomorpha Ritsema.

Taxonomy, phylogeny, zoogeography

(Insecta, Coleoptera, Carabidae)*

Martin Baehr

Baehr, M. (1997): Revision of the Pseudomorphinae of the Australian Region

2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paus-

sotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny,

zoogeography (Insecta, Coleoptera, Carabidae). - Spixiana Suppl. 23: 1-508

As the second part of a general revision of the Pseudomorphinae of the

Australian, Oriental, and Aethiopian regions the genera Pseudomorpha Kirby,

Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocepha-

lomorpha Ritsema are revised.

The 117 valid species and additional 9 subspecies of Adelotopus are grouped in

26 species-groups each of which confidently represents amonophyletic unit. Keys

are provided for the species-groups of the genus Adelotopus, for the species of the

genera Adelotopus, Cainogenion, and Cryptocephalomorpha, and for the Australian

species of the genus Pseudomorpha. For the latter species a new subgenus Austro-

pseudomorpha, subgen. nov., and for the Argentinian and south Brazilian Pseudo-

morpha laevissima Chaudoir and P. glabra Ogueta a new subgenus Notopseudomor-

pha, subgen. nov. of the genus Pseudomorpha is erected. For Cainogenion ephippiatum

(Newman) a new subgenus Procainogenion, subgen. nov. is erected.

Cainogenion cylindricum (Chaudoir) is transferred to Paussotropus Waterhouse

and Adelotopus insignis Sloane is transferred to Pseudomorpha Kirby, subgenus

Austropseudomorpha, subgen. nov.

Following species names are synonymized (valid name second): Adelotopus

bijugus Darlington = A. rufoguttatus (Blackburn); A. brunneus Castelnau = A. gyrinoides

Hope; A. castaneus Castelnau = A. rubiginosus Newman; A. filiformis Castelnau =

A. nemosomoides Westwood; A. longipennis Macleay = A. elongatulus Macleay;

A. mastersii Macleay = A. dytiscides Newman; A. micans Blackburn = A. paroensis

Castelnau; A. niger Notman = A. paroensis Castelnau; A. occidentalis Castelnau =

A. gyrinoides Hope; Cainogenion bicolor (Castelnau) = C. ephippiatum (Newman);

Paussotropus parallelus Waterhouse = Paussotropus cylindricus (Chaudoir).

Adelotopus rufoguttatus (Blackburn) and Cainogenion subopacum (Macleay) have

been revaluated from synonymy to full species status.

Lectotypes, and eventually paralectotypes, are designated for the following

* Supported by four travel grants and a grant for printing expenses of the Deutsche Forschungsgemeinschaft

(DFG).

species (including those names synonymized in present work): Pseudomorpha

insignis (Sloane), Adelotopus analis Macleay, A. apicalis Macleay, A. bimaculatus

Macleay, A. brevipennis Macleay, A. brunneus Castelnau, A. castaneus Castelnau,

A. celeripes Lea, A. dytiscides Newman, A. elongatulus Macleay, A. fasciatus Castel-

nau, A. filiformis Castelnau, A. fortnumi Hope, A. gyrinoides Hope, A. haemorrhoidalis

Erichson, A. laevis Macleay, A. linearis Macleay, A. longipennis Macleay, A.maculipennis

Macleay, A. mastersii Macleay, A. micans Blackburn, A. nemosomoides Westwood,

A. occidentalis Castelnau, A. paroensis Castelnau, A. politus Castelnau, A. rubiginosus

Newman, A. rufoguttatus (Blackburn), A. tasmani Blackburn, A. vicinus Castelnau,

A. zonatus Castelnau, Cainogenion bicolor (Castelnau), C. creberrimum (Blackburn),

C. ephippiatum (Newman), C. ipsoides (Westwood), C. obscurum (Castelnau), C. sub-

opacum (Macleay), Paussotropus cylindricus (Chaudoir), P. parallelus Waterhouse,

Cryptocephalomorpha collaris (Waterhouse), and C. marginata (Waterhouse).

A neotype is designated for Adelotopus affinis Castelnau.

The following 113 new taxa are described: Pseudomorpha brevis, spec. nov.,

P. insignis pilosa, subspec. nov., P. subangulata, spec. nov.; Adelotopus adelaideae, spec.

nov., A. adustus, spec. noV., A. aequus, Spec. NOV., A. angustatus, spec. NnOV.,

A. aterrimus, spec. nov., A. atrorufus, spec. nov., A. bacillus, spec. nov., A. bamagae,

spec. nov., A. basirufus, spec. nov., A. bimaculatus angustior, subspec. nov., A. brevior,

spec. nov., A. brittoni, spec. nov., A. browni, spec. nov., A. calvus, spec. nov.,

A. caniae, spec. nov., A. ciliatus, spec. nov., A. ciliatus tenuipunctatus, subspec. nov.,

A. clepsydra, spec. nov., A. conicollis, spec. nov., A. convexicollis, spec. nov., A. convexus,

spec. nov., A. coriaceus, spec. nov., A. crassus, spec. nov., A. cribricollis, spec. nov.,

A. crucis, spec. nov., A. cuneatus, spec. nov., A. distinguendus, spec. nov., A. doyeni,

spec. nov., A. dubius, spec. nov., A. dubius glaber, subspec. nov., A. dubius hobartensis,

subspec. nov., A. edithae, spec. nov., A. flavescens, spec. nov., A. flavus, spec. novV.,

A. foliaceus, spec. nov., A. geminus, spec. nov., A. gibbosus, spec. nov., A. gippslandi-

cus, spec. nov., A. grossepunctatus, spec. nov., A. gyrinoides orientalis, subspec. nov.,

A. houstoni, spec. nov., A. howdenorum, spec. nov., A. katherinei, spec. nov., A. kurandae,

spec. nov., A. languidus, spec. nov., A. laticaudatus, spec. nov., A. laticollis, spec. nov.,

A. latior, spec. nov., A. latipalpis, spec. nov., A. lawrencei, spec. nov., A. longiformis,

spec. nov., A. longus, spec. nov., A. longus tropicus, subspec. nov., A. lunatus, spec.

nov, A.luteus, spec. nov., A. macilentus, spec. nov., A. mainae, spec. noV.,

A. marginicollis, spec. nov., A. minor, spec. nov., A. montisatri, spec. nov., A. montorum,

spec. nov., A. multipunctatus, spec. nov., A. murrayanus, spec. nov., A. nigricauda,

spec. nov., A.nitens, spec. nov., A. nitidior, spec. nov., A. obsoletus, spec. nov.,

A. ooldeae, spec. nov., A. ovatus, spec. nov., A. palumae, spec. nov., A. parumpunctatus,

spec. nov., A. penelopeae, spec. nov., A. piceus, spec. nov., A. punctatissimus, spec.

nov., A. puncticollis angustemaculatus, subspec. nov., A. punctulifer, spec. nov.,

A. queenslandicus, spec. nov., A. rufescens, spec. nov., A. rufocaudatus, spec. nov.,

A. rufomarginatus, spec. nov., A. rufozonatus, spec. nov., A. sedlaceki, spec. noV.,

A. semilunatus, spec. nov., A. seminitidus, spec. nov., A. sericeus, spec. nov.,

A. seriepunctatus striatus, subspec. nov., A. similis, spec. nov., A. sinuaticollis, spec.

nov., A. sinuaticollis calliope, subspec. nov., A. sparsepunctatus, spec. nov., A. substriatus,

spec. nov., A. ulrichi, spec. nov., A. unicolor, spec. nov., A. victoriensis, spec. noV.,

A. villosus, spec. nov., A. virgatus, spec. nov., A. yorkensis, spec. nov., A. zborowskii,

spec. nov.; Cainogenion clypeale, spec. nov., C. creberrimum gnaltae, subspec. nov.,

C. depressum, spec. nov., C. glabratum, spec. nov., C. interiore, spec. nov., C. ipsoides

occidentale, subspec. nov., C. parumpilosum, spec. nov., C. rotundicolle, spec. nov.,

C. tropicum, spec. nov.; Cryptocephalomorpha australica, spec. nov., C. genieri, spec.

nov., and C. maior, spec. nov.

Following six species remain doubtful, because the types are lost and the species

belong to species-groups in which species distinction is impossible without com-

parison of the types: Adelotopus aphodioides Westwood, A. cornutus Castelnau,

A. hydrobioides Westwood, A. inquinatus Newman, A. papuanus Gestro, and

A. scolytides Newman.

First instar larvae are described for the first time from the oviducts of many

species of the genera Adelotopus, Cainogenion, and Paussotropus. Their special

characters are included in the phylogenetic analysis.

The possible relationships of the genera of Pseudomorphinae, the subgenera of

the genus Pseudomorpha, the species-groups of the genus Adelotopus, and the species

of the genera Cainogenion and Cryptocephalomorpha are discussed and described in

cladograms based on a reconstructed phylogeny employing the methods proposed

by Hennig. The phylogenetic relations of the species of Adelotopus and of Pseudo-

morpha are discussed. Phylogenetic evidence shows that Sphallomorpha is the

adelphotaxon of all other genera, Pseudomorpha the adelphotaxon of the remaining

genera, the highly specialized genus Cryptocephalomorpha presumably the adelpho-

taxon of the Adelotopus-lineage, in which Adelotopus is the adelphotaxon of Caino-

genion + Paussotropus, whereas Paussotropus is the most evolved genus of this

lineage.

There is strong evidence for an increasing grade of adaptation to myrmecophi-

lous habits in Pseudomorphinae in general, but also within the larger genera.

Whereas the general structure in Sphallomorpha and even still in Pseudomorpha may

be rather adaptative to the subcorticolous life, this is reduced in favour of certain

adaptations to myrmecophily in the other genera with Paussotropus and again

Cryptocephalomorpha being most evolved in this respect. Comparable adaptations to

myrmecophilous habits are also noted in the larvae of Pseudomorpha, Adelotopus,

Cainogenion, and Paussotropus, while larvae of Cryptocephalomorpha are still un-

known.

The distribution of the species is depicted in maps. About 25 distinct distribution

patterns are distinguished and most of them may be regarded as related to faunistic

subregions. Phylogenetic evidence shows that those species and species-groups

that are plesiomorphic in many respects concentrate in the southeastern subregions

of Australia, where also the most plesiotypic species of the genera Sphallomorpha,

Adelotopus, Cainogenion, and Pseudomorpha subgenus Austropseudomorpha occur.

Especially in the northeastern, northern, western, and central subregions a larger

number of species shows a higher degree of apomorphic character states. This

pattern of distribution is exemplified by several dichotomous and trichotomous

vicariant relationships, in which the evolved vicariant(s) almost invariably occur(s)

in the northern or western part of the common range.

Distribution of the apparent basal species of all genera either in southeastern

Australia (Sphallomorpha, Adelotopus, Cainogenion, Pseudomorpha subgenus Austro-

pseudomorpha), or in southern South America (Pseudomorpha subgenus Notopseudo-

morpha), or in South Africa (Cryptocephalomorpha) is evidence of aGondwanaland

origin of the subfamily without knowing the exact place of origin. Evidently most

genera apart from Cryptocephalomorpha and perhaps Pseudomorpha originated in

southeastern Australia or the adjacent part of Gondwanaland. Cryptocephalomorpha,

however, is certainly a recent arrival in the Australian region that immigrated from

the Oriental region, where it probably arrived drifting on terranes of the previous

Sundaland. According to chorological, phylogenetic, and paleogeographic evi-

dence the subfamily and most of its genera must have evolved at least during

Upper Jurassic or even earlier.

The further history of the genera in Australia was comparable to that of

Sphallomorpha, but their taxonomic radiation may have been even more recent than

in the latter genus and it is certainly not yet finished. It occurred mainly in

Pleistocene, synchronous with the late taxonomic diversification of eucalypts

during the same period and generally in closer relation to ants. Several lineages of

Adelotopus and Cainogenion then spread over most of Australia, mainly in an anti-

clockwise, northern and western direction. The repeated change of wet pluvials

and dry interpluvials during Pleistocene, with their spreading of suitable plant

communities (namely Eucalyptus forests or savannahs) and the subsequent isola-

tion of these communities together with their Pseudomorphine hosts accounts for

the high number of endemic and highly evolved species especially in the refugia

of central eastern and northern Queensland, far Northern Territory, northwestern

Australia, and central and southwestern Western Australia.

Dr. M. Baehr, Zoologische Staatssammlung München, Münchhausenstr. 21,

D-81247 München, Germany

SEE

Contents

Introduelon te ee RUHE ONE 12

Maäteriaäl....uuu.us een br re ee ann RI. ee 1092

Apbreviations’of celleetions mentioned initext.........u..n..e seen nennen nase anne near n 2

Methods Wan Lern nee 14

4] Raxonomie prmeipless en ne a EEE 14

4.2. Phylogenetie prineiplesr.........eeneueeeseseseeneeenenenenebeseuetnersesesnsnnranenenersgran sten pensheupärasenaedserer anne 14

4:32 Descnpionssrr ee 15

Ad. Tvpesten. nn. neeneeesneneeneedetenehedeenawsneensesnsnnane enranRnen one rerenancer anne Bess sasnsae ann nen ar a 115

45% Orieinol smatenlale. ae en nn nn en en ker ee Eee 16

4:6. Disunsuishing;characterse. nie zen een mn en en ne 2 er er EEE 16

AU CN EOS LE nen 17

4:8. Mustrationsrer...esmseentesenesrensansenneneetanenneee nesaseener ee nuer ea ear en eonnnn ern as eEneTe ee een 17

4:9. Data of examınedmatenial er. see 17

2310 ADistibublon maps en 18

ARIET MEASULEINENTESSH Rn EN REES UE ER ahserT NETTE SEE 18

4.122, Macnilicationse =.....0...20 Hnsnasensszerstennunrpeletgeeten en nekesedeneuense seen eHenssensnsseharae nenne ee er ee 18

4.133: Abbreviations m... re 18

Way of lite and eolleetiine,methodS....eausseesenscnsusancnesessseensnsssnninssuenseer ne we nssnszer ge en er 19

5412 Wayzol.liie. 2.2... 2er Reef nennen 19

S2ANSImecophilousihabitskr. en. nn. nn an nee 19

53, Colleetingmethods and preservation....uıessenssess nee essarenenanenananstensnnnens ernennen ee 21

| LEE ee renanchere 21

6.1. Theilarvae of the different pseudomorphine genera ..........Jneresronsnssssesssanstkesssnsdrnesnensnser ne 22

Classification... na Ba BER 40

7.1 Subfamuly Pseudomorphinaett Sr... seen eennnneenreeesetnnrae Ve hnekkesgenene een ER ER 40

7A MKeytoithe generarof Pseudomorphinae....eurscsssmrecensrseressne saurer sbebetes en sneasn care ncn ses ee 40

7.2. |GENErA ss asese ss Bssnie tr annaanen anne aaenderngenenen sn gnge dag Ede bBrR Henne HEE TE Le ee er 41

7.2.1, Genus PseudomorphauKitby. 20.2.2 enden ensure ee 41

7.221.1., Subgenus Notopseudomorpha,,subgen..noV. = uenunsiet users ee 42

7.2.1.2... SubgenusAustropseudomorpha;subgen. Nov. en. zen een tee 42

7.2.1.3. Description. of Austropseudomorpha. nen ig en 43

7.2.1.4. Key to the Australian species of genus Pseudomorpha Notman, subgenus Austropseu-

domorpha;'subgen: Nov... 2 ar ee 43

7.2.1:9.. Ihe species‘of Austropseudomorpha, subgen!mov. 2... seen een een ee 43

7.3. Genus’ Adelotopus Flope.. un... ses austneneeeeuenneen nen tan sn near aan sgestnae nee re 51

7.3.1. Description of AdelotopusS..=....nne: 2202020220 enne anna ana ae aea een rer ee 5

7.3.2. Key to the species-groups of the genus Adelotopus Hlope.......::2«sa:4u2usu2ue202022225carexentnessneee 58

7.9.3, Key to, the species of the, genus; Adelotopus Elope...2....2.2..2o N 0saneesaeseeter nern ee 60

7.3:4.. The;species of the genus: Ädelotopus Hope»... 22mm 76

7-A.»GenustEainogenion Notmanı..... 1er nel sSehn es re LEE 328

7,4. 1,Desenption’of’Erinogenion u... 200er ee rede are en ee 329

7.4.2. Key'to the species of the genus Camogenion’Notman..n.e..ereeesnsreeststeaeereensse nenne een 334

7.4.3. The:speeies’of the genus Cainogenion! NOLMan..... essen asenseesuneeseenessonennen re 336

7431» Shbgenus-Procainogenion:subgen NOV. ur. mn ernennen ae re 336

7.4.4.2. Subgenus Cainogenion S.,$tr........0.0.0.. ara iassennhesernnan anne sen ent reahineeen teen 340

SREENUSTRAUSSOLTOPUSEVVAtERNOUSE LA neeeteenrauneneessenenvannanendonenldaneneentendenenkenbe ne 370

DES NONE 370

7.5. 2iheispeeies-of thergenus; PaussotropusWaterhouse..........umeseessscsesuenessenesneneseresnenesnnnuenanenne 370

RO Genus!@ripLocenhalomonpharRitsemar.. eskerenesneneheesenrenenertnrenenerneeauecercennnenesgesesnee einen 374

OD EScHIPLIONONC7Ypioeephalomorpha zerenenenaesneeeasesasaenesonnne ce eeneonraesnennenesrenenenanennen san eeneeneneenn 375

7.6.2. Key to the species of the genus Cryptocephalomorpha Ritsema ............enenene: 977,

726.38 Ihesspeeiesiofithe'gemus Criptocephalomorpha Ritsemans...messaceeesueeeteeteeceeseennae ea 378

7.72 Boubthulspeciesr...n.m essen ven neetarezansnen eat arete een nenne ernst rncntersetheteescnateeter nee 391

728 Speciesserroneously.tecordedirom Australian... en. eeeneeessenee era 392

Reconstiüretedphylogenyrın..... ts eeeenneenennseetnee ee nesennense ee een teen 393

Sl zoystematie,posıLon or.the,ssubfamily Pseudomorphinae......unzeeeseceneneenenenee nee 393

8.2. Phylogenetic relationships of the genera of the subfamily Pseudomorphinae ................. 393

8.3. Phylogenetic relationships of the subgenera of the genus Pseudomorpha Kirby ................ 400

8.3.1. Phylogenetic relationships within the subgenus Austropseudomorpha, subgen. nov....... 403

8.4. Phylogenetic relationships within the genus Adelotopus Hope ..............uuneesennee 403

8.4.1. Phylogenetic relationships within the species-groups of the genus Adelotopus Hope ... 412

8.5. Phylogenetic relationships within the genus Cainogenion Notman .........u.eneeesee: 416

8.6. Phylogenetic relationships within the genus Cryptocephalomorpha Ritsema .............- 422

8.7. Evolutionary trends within the subfamily Pseudomorphinae ....u......u.u..00.0 425

OOLEOTTAD UV Reseller tnereche 428

IR Nitro duch One ee ee 428

I2ADIStEIPUHONPaLIeLnS re ne een neben tosene ee 429

9.2.1. Distribution of genera and of species-groups of Adelotopus: phylogenetic correlations 429

3227 Ecogeoeraphical Patterns of distribution... essen. esnesngencheenensssensorssensgrsenrsteegensennsee 433

9.2.3. Distribution patterns'of species: phylogenetie correlations ...........2..u.0nu.2100004222esnasaanenn 440

9.3. Biogeographic subregions, barriers, and their significance for biogeographic history ..... 443

9.4. Biogeographic history of the subfamily Pseudomorphinae and of the included genera . 447

IA BINIKOAUEHON. ee reain irren reeerere ren eNrerrae en Teeeeeeeeeeeee 447

9.4.2. Paleogeological aspects correlated to the origin of the subfamily Pseudomorphinae ... 447

9.4.3. Time of origin of Pseudomorphinae and of the included genera .............nenene 448

9.4.4. Place of origin of Pseudomorphinae and of the included genera ...............ee 449

92:9. Ecologicalregquirements,of ancestral Rseudomerphinae:.......-. 2.2.2.2 449

9.4.6. Biogeographical history of the genera of Pseudomorphinae..............ueseseseesssesesneeneseenenen 450

ION Genus Sphallomonphan ae. Me eines den äiesehah kecnasreesänsneunen 450

IENGDAGENUSPSEUAOMOTPHIn. Sa. rn N ee nn kenne isie nee 450

IEOSNGENUSAdBlOrOHUSEE He Eee ME 451

9.4.0.4. Genera. Cainogenion and Paussotropusen set snrasnsurlenedebeingesasseneheenaneundahnensnssnsnneneeaese 452

94.09. GenusiErypiocephalomonpha ==. een nee nedenssnnesnenssone 453

9.5. Outview to the future biogeographical history of the Australian Pseudomorphinae ...... 455

OPENIgUSSHONSK..2.n..n.ssranzereencssenrsnenss ss süsin kn farssnnnnasrngsgintecniehunngesneggsrsrtngnnnnurnigtt nern nennt nntnennn anna eat 456

BEN EKNOWIEILEN NE SE RE EN Eee 457

Biieures>habitus, elytrallmiıeroseulpture, Mapscz rn u. nesszensensensegeteseenennsuszssnsgaggestsneseerreene 458

REferENGES ee ee ee alesdasiakiesinisee 500

„Alphabetieal:checklist of genera, subgenera, and species..................2:2=.002222820e00r2020c20gsesseusätsenge 505

. Checklist of the species-groups of the genus Adelotopus with the included species .............. 507

1. Introduction

As the second part of the general revision of the pseudomorphine beetles of the Australian region the

remaining genera Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, Cryptocephalomor-

pha Ritsema, and the Australian species of the genus Pseudomorpha Kirby are revised. For completeness

the (few) South Asian species of Adelotopus and Cryptocephalomorpha and also the single, recently

discovered African species of the latter genus have been included in this revision. Apart from the very

large number of Australian species, the revision covers therefore some species from New Guinea, the

Solomon Islands, Indonesia, the Philippine Islands, Vietnam, Thailand, Malaysia, and South Africa.

Since publication of the first part of this revision no important new opinions about the relation-

ships of the subfamily Pseudomorphinae have been developed, because even the fine study of Arndt

(1993) on the larval characters of Carabidae does not contain new ideas on the relations of the

Pseudomorphinae, but only states their highly isolated status. However, there are new ideas to the

phylogenetic relations within the subfamily Pseudomorphinae (Baehr 1994a) that will be referred to

in the discussion of phylogeny and biogeographical history.

The number of species currently recognized are (Moore at al. 1987 for Australian, and Darlington

1968 for New Guinean species, both except for genus Sphallomorpha, and Baehr 1992a, 1993a, b, 1994b

for genus Sphallomorpha): Sphallomorpha: 143 species and one additional subspecies; Adelotopus: 42

species; Cainogenion: 6 species; Paussotropus: 1 species; Cryptocephalomorpha: 3 species. Of the genus

Pseudomorpha thus far no species from outside of the Americas were known.

2. Material

I have examined altogether 4581 specimens of Adelotopus, 941 specimens of Cainogenion, 36 specimens

of Paussotropus, 76 specimens of Cryptocephalomorpha, 10 specimens of Australian Pseudomorpha, and

additional 35 specimens of American Pseudomorpha from the collections of those institutions and

individuals listed below and from my own collection. Of American Pseudomorpha the following species

have been examined: P. angustata Horn, P. argentina Steinheil, P. arrowi Notman, P. behrensi Horn,

P. castanea Casey, P. champlaini Notman, P. excrucians Kirby, P. gerstaeckeri Chaudoir, P. glabra Ogueta,

P. lacordairei Dejean, P. laevissima Chaudboir, P. pilatei Chaudoir, and P. vindicata Notman.

3. Abbreviations of Collections mentioned in text

AMNH American Museum of Natural History, New York

AMS Australian Museum, Sydney

ANIC Australian National Insect Collection, Canberra

ANSP Academy of Natural Sciences, Philadelphia

BMH B. P. Bishop Museum, Honolulu

BMNH The Natural History Museum, London

CAS California Academy of Sciences, San Francisco

CBM Working collection M. Baehr, München

CBS Collection V. Beysak, Sydney

CMC Collection B. P. Moore, Canberra

CMP-WHC Walford-Huggins Collection, Carnegie Museum, Pittsburg

CSB Collection J. Sedlacek, Brisbane

CUIC Cornell University Insect Collection, Ithaca

DEIB Deutsches Entomologisches Institut, Eberswalde

DPIM Department of Primary Industries, Mareeba

FMNH Field Museum of Natural History, Chicago

FMT Frey Museum, Tutzing, now München

HNMB Hungarian National Museum of Natural History, Budapest

IRSNB Institut Royal des Sciences naturelles, Bruxelles

MCSN Museo Civico di Storia Naturale, Genova

Fig. 1. Habitus of different Pseudomorphinae. Above, from left: Pseudomorpha (Notopseudomorpha) laevissima

Chaudoir, Pseudomorpha (Pseudomorpha) behrensi Hom, Pseudomorpha (Austropseudomorpha) insignis insignis

(Sloane), Sphallomorpha froggatti (Macleay), Sphallomorpha flavicollis (Macleay). - Below, from left: Adelotopus

dytiscides Newman, Adelotopus linearis Macleay, Cainogenion (s. str.) obscurum (Castelnau), Paussotropus cylindricus

(Chaudoir), Cryptocephalomorpha gaverei Ritsema.

MCZ

MMS

MNHB

MNHN

MNTD

Museum of Comparative Zoology, Cambridge/Mass.

Macleay Museum, Sydney

Museum National de l’Histoire Naturelle, Paris

Naturhistorisches Museum, Basel

Naturhistorisches Museum, Wien

Naturhistoriska Riksmuseet, Stockholm

The Canadian Museum of Nature, Ottawa

Museum of Victoria, Melbourne

Nationaal Natuurhistorisch Museum, Leiden

Oxford University Museum, Oxford

Queensland Museum, Brisbane

South Australian Museum, Adelaide

Senckenberg Museum, Frankfurt/M.

Staatliches Museum für Naturkunde, Stuttgart

Staatliches Museum für Tierkunde, Dresden

University of Alberta, Strickland Museum, Edmonton

University of Queensland Insect Collection, Brisbane

United States National Museum, Washington, D.C.

University of Vermont Collection, Burlington

Western Australian Museum, Perth

Z.oologische Staatssammlung, München

Museum für Naturkunde der Humboldt Universität, Berlin

Museum and Art Gallery of the Northern Territory, Darwin

4. Methods

4.1. Taxonomic principles

The taxonomic principles are basically the same as in the first part of this revision. However, in the

genus Adelotopus the differentation of taxa is evidently even more difficult than in the genus

Sphallomorpha, mainly due to more recent evolution and diversification of the genus as a whole and of

many of its taxa, and to reduction or complete loss of certain important characters as for example

chetotaxy. However, there has been considerable taxonomic radiation which led to groups of closely

related taxa. Because of the known difficulties in recognizing subspecies from morphological evidence

alone, those closely related taxa have been described generally as species if they are sympatric, and as

subspecies, when they are evidently allopatric. It must be stressed, however, that the actual distribu-

tion of many taxa is very inadequately known, although a very large part of the specimens existing in

the collections throughout the world have been examined. Hence, some closely related taxa that are

regarded as allopatric subspecies at the present state of knowledge may be actually sympatric, and so

their taxonomic status might have to be revised in future. On the other hand, many old, perhaps

inaccurate or mislabelled locality records that seem to establish sympatric distribution of taxa, are

perhaps wrong. As a consequence, those taxa may be actually allopatric.

In some groups of closely related taxa the distinction of taxa may thus appear rather artificial and

perhaps readers do not agree with some of the taxonomic decisions made herein. lam aware that some

of the taxa distinguished and named by me are disputable and may not be justified when other than

morphological methods are applied. Nevertheless, I generally chose the attitude of a “splitter” when

dealing with species or infraspecific units, but of a “Jumper” at the generic level. I did so mainly for

heuristic reasons, in particular to give users or later revisors the opportunity to trace without

difficulties aberrant specimens or populations that likely would get lost in the material, when they

would be regarded only as “variations”. I feel that in some of these taxa morphological taxonomy is

likely to have reached its limits and should be replaced by or supplemented with other methods.

With regard to genera, however, ladopt the term genus in a rather wide sense, on the same reasons

as explained below (see chapter “Genera”) and in the first part of the revision.

4.2. Phylogenetic principles

For the phylogenetic analysis I used the same methods as in the first part of this revision. They were

proposed by Hennig (1966) and firstly adopted by Brundin (1966), and were for example further

explained by Saether (1990). I did not choose the quantitative phyletic or (from my view) “numerical

cladistic” approach, because I think character analysis must be the prior condition to construction of

a phylogenetic tree and the criterion of parsimony to be used posteriorly to character analysis. I feel

that parsimony may be perhaps not as important when tracing phylogeny as most propagators of that

criterium believe, because evolution of species does not necessarily proceed according to that criterion,

but commonly proceeds along roundabouts and so application of the principle of parsimony may

easily generate fictitious results.

Moreover, I feel, that quantitative phyletic methods easily allude users to overestimate the value

of certain rather weakly polarized or even unpolarized characters that would perhaps not be used

when employing non-computerized cladistic methods. When such characters are introduced into the

data matrix and used for establishing branchings in the cladogram, a sort of numerical analysis will

be likely introduced into cladistics.

I will be even go as far as to say: morphological characters alone almost never can give the full set

of synapomorphies necessary for settling all branchings in a cladogram, and moreover, many

characters (ethological, ecological, physiological, genetical and others) important for tracking phylo-

genetic relatioships are missing when using such method. Therefore, I would like to stress that a

cladogram based on a purely morphological analysis that supports all branchings with synapomor-

phies, should be a priori distrusted, at least in parts.

Although speciation occurs according to the rules of the biological species concept, in entomology

at least, its observation under natural conditions is very rarely possible. Hence characters are needed

as a means for distinction of species. Even when morphological structures are most widely used, other

characters may be of the same value, e.g. physiological, etho-ecological, cytological, or genetical. But

even genetics yield only characters and hence do not solve the problem of applying the biological

species concept to the distinction of species.

Because reconstruction of the phylogeny and history of the fauna or of a given superspecific taxon

is primarily based on the acquisition of adaptations (i.e. characters), analysis of the ancestral or

derivative status of characters is generally the only way to reconstruct phylogeny. As Hennig

demonstrated, only derivative (apomorphic) character states are useful in such phylogenetic classifi-

cation, because they alone define monophyletic groups and can be used to establish sister group

relations.

As a first step towards a phylogenetic classification and analysis of faunal history the attempt is

made to determine the state of the used characters. Character analysis, i. e. decision about plesiomor-

phy and apomorphy of characters, is based either on outgroup comparison using character states

found in related taxa of higher categories as explained by Wiley (1981) and Watrous & Wheeler (1981),

or on group trends as explained by Ross (1974), especially when strictly synapomorphic states were

difficult to perceive because of ample parallelisms or reductions of characters. So it will be noted

especially in the phylogenetic analysis of the species-groups of Adelotopus and of the phylogenetic

relations within these species-groups that commonly evolutionary trends only can be traced, rather

than well founded sister group relations. However, outgroup comparisons beyond the limits of

Pseudomorphinae are difficult due to the highly isolated and yet unsettled position of this subfamily.

In few characters, where outgroup comparisons do not yield good results, the somewhat questionable

principle of “common occurrence” was used which usually supports the plesiomorphic status of a

character. Its value, however, is questioned (Watrous & Wheeler 1981).

In the following text "plesiomorphic” or “apomorphic” is attributed to character states, whereas

“plesiotypic” and “apotypic” is used in combination with taxa and means the degree of distance of a

taxon from the base of the group-specific cladogram expressed in the number of dichotomies from the

base of the cladogram.

4.3. Descriptions

As in the genus Sphallomorpha, most species were described in the last century and by the same authors,

namely W. Macleay (1864, 1871, 1888) and Castelnau (1867, 1868), who described together about half

of the species, and Westwood (1837, 1853), Hope (1834, 1845), Newman (1842, 1856), Erichson (1842),

Gestro (1893), and Blackburn (1893, 1901). Few additional species were described by Chaudoir (1862),

Ritsema (1875, 1909), Waterhouse (1877), Lea (1910), Sloane (1910), Notman (1925), and more recently

by Darlington (1968). Most descriptions are very short and of little or no use for recognition of species,

and even the more recent descriptions do not or barely allow an unequivocal distinction of the species.

As in genus Sphallomorpha, most types with exception of those of the species described by Macleay,

Lea, and Sloane, and few syntypes of Blackburn, are deposited in European or American museums.

This illustrates the difficult situation of the older Australian authors who had no access to most types.

Hence, the large number of misidentified species in the available material is understandable, but also

repeated description of the same species was common, and the rather large number of synonyms

stated by older authors and in this paper is easily understood.

The keys in the general review of Notman (1925) were based mainly on the old descriptions rather

than on well identified material or even on types. Because this author saw but few species, only parts

of the keys are useful, and as far as the species of Adelotopus are concerned, identification of species

by use of Notman’s key is virtually impossible with exception of few peculiar species. So, in Adelotopus

at least, identification of species was generally not possible without comparison of types. That this

opinion is right, was demonstrated by the study of the named museum material which is to an even

greater extent wrongly identified than in the genus Sphallomorpha.

4.4. Types

As in the genus Sphallomorpha, the situation concerning the types in the remaining pseudomorphine

genera is quite favourable, because most types still exist and they are generally in good condition. The

types of few named species only are definitively lost or cannot be found in the institutions where they

should be located (BMNH, MCSN, OUM). However, mostly the loss of types is serious, because these

species belong to very difficult species-groups in which species distinction is generally impossible

without consideration of the male genitalia and of certain external characters like microreticulation

and sculpture of the surface. Because the original descriptions are insufficient with respect to those

characters, these species cannot be distinguished in the existing material, or synonyms designated by

previous authors cannot be verified. Hence they must be treated generally as nomina dubia. In some

species described by Newman and Westwood this is especially disadvantageous, because they were

most commonly determinated by earlier authors in the existing material, and their names have been

frequently cited in the older literature. However, it is virtually impossible to know what they meant.

Through courtesy of the respective curators I was able to examine almost all existing types. During the

study it became evident that examination of male and usually also of female genitalia of types was

indispensible. Hence, most types were dissected and it proved that some type series contain more than

one species. On this and on other reasons, lectotypes were generally designated.

4.5. Origin of material

Although almost 5700 specimens have been examined for this study, rather few species are well

represented in the sample, whereas many are represented by few or even single specimens only. As

in Sphallomorpha, the southeastern part of Australia (eastern South Australia, Victoria, New South

Wales, southern Queensland) is fairly well represented, while material from other parts of Australia

is much rarer and has been almost exclusively collected within the last 30 years. Hence, the faunas of

Western Australia, the whole Northern Territory, interior South Australia, as well as those of western

Queensland and western New South Wales, but also that of northern Queensland, especially the Cape

York Peninsula, are very inadequately known, although the faunas are presumably much richer than

previously believed. Especially Western Australia seems surprisingly rich in species, although at

present records are scattered and most species are rare in the available material. Hence, for a more

balanced knowledge of the fauna, careful collecting work in most parts of Australia would be much

desired, and certainly this would yield additional new species in the remote areas of the north, west,

and centre. The same applies perhaps also for New Guinea and the Oriental Region, the pseudomor-

phine fauna of which is certainly not well known due to lack of specialized and systematic collecting

work.

4.6. Distinguishing Characters

Several species of Adelotopus, Cainogenion, and Cryptocephalomorpha bear more or less conspicuous

colour patterns, in most species, however, pattern alone does not offer good determinating characters,

because it varies to a considerable degree and, on the other hand, is quite similar in certain species.

Hence in many species male genitalia, commonly also female genitalia offer the best or even the

exclusive distinguishing characters. Special attention should be paid to shape of aedeagus, structure

and pattern of the internal sac, shape of parameres, genital ring, and sternite VIII, and in females to

shape and chetotaxy of the stylomeres. Generally, dissection and examination of the carefully cleaned

male genitalia is mandatory. Usually they should be leached for some time in KOH to make visible the

complicated and commonly distinctive inner structures. In some species-groups shape and structure

of aedeagus, parameres, and other genital sclerites offer virtually the only chance for species

distinction.

Commonly, however, other characters are also useful for species differentiation:

Female genitalia, especially the stylomeres, are rather useful, although they are generally very

similar within the different genera. However, some species are well recognized by the structure of

their female genitalia.

Body shape, especially shape of pronotum and elytra, in particular shape of the lateral margin of

both and of the basal angles of the pronotum, commonly provide good characters for species

differentiation, but also for distinction of species-groups.

Of the chetotaxy (see below) only the umbilical pores and setae of the elytra and the ambulatory

setae of the abdomen yield useful differentiating characters, but mainly on the species-group level.

Microsculpture (puncturation and microreticulation) of the upper surface and degree of striation

of the elytra are very useful characters, although they seem rather variable in certain species. For their

use, however, they must be very carefully examined, usually at least at 40x or even 65x magnification

and under good light.

Colour and pattern are useful for species distinction, but they should be used with care and

preferably in combination with other characters.

4.7. Chetotaxy

Contrary to the genus Sphallomorpha, chetotaxy is reduced to a considerable degree in the other

pseudomorphine genera and, as a consequence, does not offer as useful distinctive characters as in

Sphallomorpha. Hence, in the descriptions chetotaxy is not given a special paragraph.

4.8. Illustrations

Because identification of species is still very difficult, as much information as possible has been

included in the illustrations. Apart from several structures of male and female genitalia, line drawings

of the whole beetle, the clypeus and labrum, the terminal palpomeres, and the antennae of all species

were provided. Details of striation, puncturation, and microreticulation of the elytra are also provided

for all species. Frontodorsal and ventral surface of head is illustrated of one example of all genera and

subgenera. In all taxa of the genus Cainogenion the frontal view of head is figured. In the line drawings

of the habitus which have been taken from photographs, pattern is indicated by bold lines that are

more or less tightly dotted according to the degree to which pattern is well delimited. Explanations of

the lateral borders of pronotum and elytra have been finely dash-dotted to give an impression of the

convexity of the surface. Because in most species but few specimens are available, these illustrations

have not been provided as SEM pictures which would have been more predicatory. Only some

structural characters of one paradigmatic species each of all genera - except for Pseudomorpha,

subgenus Austropseudomorpha, of which material was too scarce - are presented as SEM photographs.

However, photographs of 48 species, including at least one species of all species-group of Adelotopus,

are added as illustration of the wide variation of shape and colour.

4.9. Data of examined material

Data of examined material are given in full length including date of collection and name of collector,

but without collecting circumstances which have been mentioned under “habits”. In types, the exact

labelling was used, including all ciphers, notes of determinators and curators, and printed labels.

Original spelling of date of collecting, especially of the month (arabics, roman, abbreviations), has been

used only in types, in the other specimens the month is recorded in roman words. Under “material

examined”, types have been repeated only, when non-typical material is also recorded, otherwise the

reader should look under “types”. Records of examined material have been given in an anti-clockwise

geographical arrangement, beginning with South Australia and ending with Western Australia, or

New Guinea, or Malaysia, respectively. Records not specified to a state and pure records from

“Australia”, are added under “Aus”, records not even mentioning “Australia” are cited under “?”.

States are recorded with their usual abbreviations (see under “abbreviations”). State records or data

like “Australia”, “Nov. Holl.” etc. have not been repeated in non-typical material.

Earlier determinations were recorded with quotation of the determinator. When a specimen was

sent from a collection as being arranged there under a certain species name, though without bearing

a label, this name was quoted with preceding “det.”.

4.10. Distribution maps

Distribution maps are based on label data of examined specimens only. Label data I was not able to

read or to localize and pure state records are not indicated in the maps. Many of the older specimens

bear only state records or no data at all, hence in several species distribution maps show by no means

the real distribution.

4.11. Measurements

Measurements were taken using a stereo microscope with an ocular micrometer. Length has been

measured from apex of labrum to apex of elytra. Lengths, therefore, may slightly differ from those of

other authors. Length of pronotum was measured from middle of apex to base, width of pronotum at

widest part. Width of apex of pronotum was measured between the tips of the apical angles, width of

base at the basal angles or just in front of them in the case they are rounded off. Length/width ratios

are somewhat variable in most species, but generally offer rather good measures of relative shape.

4.12. Magnifications

Especially for examination of the generally fine though taxonomically highly important puncturation

and microreticulation of the surface a stereo microscope with at least 40x, better 65x magnification is

needed, and of course a good lamp of high intensity that can be focussed. For good definition of the

microsculpture a lamp giving natural light is preferable, because fibre-glass optics substantially

change the surface structures.

4.13. Abbreviations

Following abbreviations were used in the text, especially in the keys:

Aus Australia

NG New Guinea

NSW New South Wales

NT Northern Territory

Old Queensland

SA South Australia

Tas Tasmania

Vic Victoria

WA Western Australia

? origin not specified

c central

ce central eastern

cW central western

e eastern

northern

ne northeastern

nw northwestern

southern

se southeastern

sw southwestern

Ww western

l/w ratio length/ width

w/l ratio width/length

> larger or more than

< smaller or less than

40x 40x magnification

65x 65x magnification

5. Way of life and collecting methods

5.1. Way of life

Since writing of the first part of this revision, few new records of the way of life of Pseudomorphinae

have been gathered. However, it has been shown during the ample dissections in the course of

preparing the present paper that at least the genera Adelotopus, Cainogenion, and Paussotropus are

ovoviviparous, and this has been confirmed by the discovery of larvae in the oviducts of 38 species and

one additional subspecies of Adelotopus, 5 species and one additional subspecies of Cainogenion, and

of Paussotropus cylindricus. Due to the small number of specimens available in Cryptocephalomorpha and

in the Australian species of Pseudomorpha subgenus Austropseudomorpha ovovivipary has not yet been

confirmed in these genera or subgenera, though is perhaps present. Recently R. T. Bell (in litt.)

informed me that a female Adelotopus gave birth to larvae when drawn into alcohol. Because Liebherr

& Kavanaugh (1985) described the ovovivipary in American Pseudomorpha, this phenomenon is

definitely only absent from Sphallomorpha which seems in several respects the most plesiomorphic

genus of the whole subfamily. For further discussion of the structure of larvae see below.

During recent collections made in the Cape York Peninsula in the course of a joint programme

carried out by the DPIM, QMB, and ANIC a fairly large number of pseudomorphine beetles were

collected in Malaise traps that were exposed for a longer period (one or two months) at the same

locality. Although it is generally not known, at what time of the day or night the beetles were captured

while on flight, they do perhaps fly more readily than reported from collecting work during daytime.

Apart from endogene stimulantia flying behaviour during daytime may also depend on the

surrounding temperature. This has been several times observed while capturing large numbers of

Sphallomorpha and Adelotopus species in different parts of Australia. Best time for collecting is morning,

when the beetles are still rather sluggish and almost always rely upon running away for flight, whereas

in late afternoon during warm days beetles much more easily try to escape by flying away.

5.2. Myrmecophilous habits

The myrmecophily of certain but still very few pseudomorphine species has been stated mainly by

Moore (1964, 1974), Lenko (1972), and Erwin (1981) who described this behaviour in Sphallomorpha

colymbetoides Westwood (= albopicta Newman), 5. nitiduloides Guerin, Pseudomorpha laevissima Chau-

doir, some other Pseudomorpha species from North America, and an unknown genus from Australia.

According to Lenko (1972) the larvae of Pseudomorpha laevissima Chaudboir live in the nests of

Camponotus rufipes (F.), the larvae of which they eat. Two unnamed Pseudomorpha larvae from the

southwestern USA described by Erwin (1981) were found in nests of Camponotus semitestaceus Emery,

Myrmecocystis testaceus Emery, and Neivamyrmex nigrescens (Cresson).

Contrary to the situation in Sphallomorpha, in Adelotopus, Cainogenion, Paussotropus, and the

Australian Pseudomorpha at least, more records of beetles collected with ants are now available, or

beetles have been mounted together with the presumable host ant on the same card. The following

records are available in the material at hand (Tab. 1). For better comparison tab. 2 gives also the

Australian pseudomorphine species recorded from different ant species:

Tab. 1. List of ant and termite hosts of the genera Sphallomorpha, Adelotopus, Cainogenion, and Pseudomorpha,

subgenus Austropseudomorpha.

Pseudomorphinae ants and termites

Sphallomorpha amabilis (Castelnau) Opistopsis haddani Em. nest

Sphallomorpha flavicollis (Macleay) Opistopsis haddani nest

Opistopsis respirinus nest

Sphallomorpha rhomboidalis Baehr Iridomyrmex purpureus nest

Sphallomorpha ruficollis Baehr Iridomyrmex detectus nest

Adelotopus affinis Castelnau Tapinoma sp.

Adelotopus apicalis Macleay Camponotus dorycus (Sm.)

Adelotopus brevipennis Macleay

Adelotopus celeripes Lea

Adelotopus cuneatus, spec. Nov.

Adelotopus dytiscides Newman

Adelotopus gyrinoides Hope

Adelotopus laevis Macleay

Adelotopus linearis Macleay

Adelotopus punctatus Castelnau

Adelotopus zonatus Castelnau

Cainogenion ephippiatum (Newman)

Cainogenion glabratum, spec. nov.

Cainogenion ipsoides (Westwood)

Cainogenion obscurum (Castelnau)

Cainogenion parumpilosum, spec. Nov.

Cainogenion tropicum, spec. NOV.

Pseudomorpha brevis, spec. NOV.

Cryptocephalomorpha papua Darlington

Iridomyrmex sp.

Camponotus sp.

Hypoclinea (= Diceratoclinea) sp. (nov.?)

Wasmannia sp.

Iridomyrmex sp.

Camponotus nr. nigriceps (Sm.)

Iridomyrmex conifer Forel

Iridomyrmex sp.

Coptotermes acinaciformis

Mastotermes sp.

Technomyrmex sp.

Crematogaster sp.

Iridomyrmex sp.

Formica sp.

“under bark attended by ants”

Crematogaster sp.

“Small ants”

Tab. 2. List of ants and termites and their pseudomorphine guests

Ants

Pseudomorphinae

Camponotus dorycus (Sm.)

Camponotus nr. nigriceps (Sm.)

Camponotus spec.

Crematogaster spec.

Cremategaster spec.

Crematogaster spec.

Formica spec.

Hypoclinea (=Diceratoclinea) spec. (nov?)

Iridomyrmex conifer Forel

Iridomyrmex detectus

Iridomyrmex purpureus

Iridomyrmex spec. 1

Iridomyrmex spec. 2

@S Eu > Ep

Iridomyrmex spec. 3

Opisthopsis haddani

Opisthopsis respirinus

Tapıinoma spec.

Technomyrmex spec.

Wasmannıia spec.

unspecified ants

Adelotopus apicalis Macleay

Adelotopus dytiscides Newman

Adelotopus celeripes Lea

Cainogenion glabratum, spec. nov.

Pseudomorpha brevis, spec. nov.

Cainogenion ephippiatum (Newman)

Cainogenion tropicum, spec. NOV.

Cainogenion parumpilosum, spec. nov.

Cainogenion ipsoides (Westwood)

Adelotopus celeripes Lea

Adelotopus gyrinoides Hope

Sphallomorpha ruficollis Baehr

Sphallomorpha rhomboidalis Baehr

Adelotopus cuneatus, spec. NOV.

Adelotopus brevipennis Macleay

Adelotopus laevis Macleay

Cainogenion ipsoides (Westwood)

Sphallomorpha amabilis (Castelnau)

Sphallomorpha flavicollis (Macleay)

Adelotopus affinis Castelnau

Adelotopus zonatus Castelnau

Adelotopus celeripes Lea

Cainogenion obscurum (Castelnau)

Cryptocephalomorpha papua Darlington

Termites

Coptotermes acinaciformis

Mastotermes spec.

Adelotopus linearis Macleay

Adelotopus punctatus Castelnau

As the lists above demonstrate, thus far rather few ant genera are known as hosts of pseudomorphine

species. Although the records are still fairly unsatisfactory, few perhaps more general trends are worth

noting:

1. Very few pseudomorphine-ant host relations are yet recorded in the genus Sphallomorpha.

2. Ants of the primitive genus Camponotus are only recorded as hosts of three decidedly primitive

Adelotopus species, but also of the primitive Pseudomorpha laevissima and of a further North American

Pseudomorpha species.

3. The ant genus Crematogaster is not yet known as host of any Adelotopus species, but of several

species of Cainogenion and of one Australian Pseudomorpha species.

Very little is known about the way of life of pseudomorphine larvae. Apparently there is only

Moore's (1974, 1983) description of the larval behaviour of two Sphallomorpha species. According to

Moore the proved myrmecophilous species of this genus are commensals the larvae of which are fairly

normal shaped and live like cicindelid larvae in burrows around the nests of ants of the genus

Iridomyrmex that they eat. The larvae of the other genera, however, are inquilines and this is in

conformity with the mode of reproduction that is ovoviviparous (larviparous) in these genera, while

oviparous in Sphallomorpha. Larvae of the highly evolved genus Cryptocephalomorpha are still unknown.

5.3. Collecting methods and preservation

Collecting methods for the genera Adelotopus, Cainogenion, and Paussotropus are basically the same as

described for Sphallomorpha. However, even more attention should be paid to ant nests, since species

of the mentioned genera are likely even more dependent on ants than those of Sphallomorpha (see

chapter above). Little is known in this respect about the Australian species of Pseudomorpha, some of

which have been captured in “pitfall traps”, and species of Cryptocephalomorpha are generally found

only at light, but should be carefully sought for in leaf litter near ants.

As in Sphallomorpha, the specimens should be mounted in such a way that at least the lower surface

of the head is visible. Male and female genitalia should be carefully dissected and cleaned, and the

aedeagus should be leached for some time in KOH. The name of the host tree, as well as the name of

the ant species, when the beetle was found by ants, should be added to the label, because there is still

very little knowledge of interdependence of pseudomorphines with their host trees and ants.

6. Larvae

In a highly specialized, rather unform, myrmecophilous beetle group the phylogeny of which is

difficult to trace, larvae might be particularly suitable for phylogenetic reasoning because they might

better preserve the original differences between the genera than imagines. However, little is known

about the larvae of Pseudomorphinae and not even from all genera larvae have been recorded. Moore

(1964) described the first pseudomorphine larva and took it for a larva of the genus Sphallomorpha.

Subsequently, Moore (1974) observed and reared the true larvae of two Sphallomorpha species and

thought that the formerly described larva might belong to either Adelotopus or Cainogenion. Lenko

(1972) figured the outline of the larva of the South American Pseudomorpha laevissima Chaudoir, and

Erwin (1981) summarized what was then known about pseudomorphine larvae and described and

figured the Sphallomorpha larva as well as two further Pseudomorpha larvae. Liebherr & Kavanaugh

(1985) first reported the ovoviviparous (larviparous) parturation in Pseudomorpha, and Moore (in litt.)

and Bell (in litt.) recently observed the same in one Adelotopus species each. Including the observations

made during the dissections for the present revision, larvae are now known from 2 species of

Sphallomorpha, 3 species of American Pseudomorpha, 38 species of Adelotopus, 5 species of Cainogenion,

and from Paussotropus cylindricus, the single species of this genus. No larvae are so far known from

most species-groups of Sphallomorpha, especially from the more plesiomorphic species-groups and

from those groups formerly included in the genus “Silphomorpha”, from the Australian subgenus

Austropseudomorpha of the genus Pseudomorpha, and from Cryptocephalomorpha.

According to the descriptions and figures of Moore (1964, 1974), Lenko (1972), and Erwin (1981)

and the summary of Arndt (1993) pseudomorphine larvae are fairly apomorphic in: the absence of

lacinia, stemmata, and urogomphi; their cicindelid or physogastric habitus; the small, I-segmented

galea; and the peculiar chetotaxy with fungiform or apically split setae on the head. However, some

of these characters do not apply to all pseudomorphine larvae, and in some species or even genera the

galea is 2-segmented, or the head does not bear fungiform or apically split setae.

The known larvae of Sphallomorpha are certainly most plesiomorphic within the family, because

they are neither physogastric, nor do they apparently possess the characteristic fungiform, or club-

shaped, or apically split setae on the head (see figures in Moore 1974 and Erwin 1981) that are present

in the figured Pseudomorpha larvae (Erwin 1981) and in those larvae of Adelotopus I examined.

Moreover, in the figured Sphallomorpha larva the head is rather large and not lengthened and the legs

are still fairly elongate. Moore (1974) described the habits of the two Sphallomorpha larvae as much alıke

to that of certain tiger beetles, because they wait in holes in the ground around ant nests for passing

ants and seize them from their hollows. It must be stressed, however, that these larvae belong to highly

apomorphic species-groups and species within the genus, and it is by no means certain that larvae of

plesiomorphic species-groups of Sphallomorpha have similar morphology and way of life.

The only apotypic feature I can find in the two described Sphallomorpha larvae is the styliform

antenna. But a similar styliform antenna is also present in the larva of Adelotopus macilentus, spec. nov.,

a fairly evolved species within its genus, whereas the known larvae of the other Adelotopus species

have normal shaped antennae. Hence, the apotypic structure in Sphallomorpha equally could have been

evolved within this genus and it is not necessarily an apomorphic state of the genus as a whole as

Erwin (1981) postulated.

The figured Pseudomorpha larvae (Lenko 1972, Erwin 1981) are highly specialized, because they are

conspicuously physogastric, have very short legs, a small and elongate head with small mouth parts,

lack most of the body setae but bear characteristic fungiform setae on the head. The larva mentioned

by Moore (1964) and again pictured in Erwin (1981) is rather similarly shaped but lacks the

characteristic fungiform setae and may actually belong either to a species of Cainogenion or to the

Australian subgenus Austropseudomorpha of the genus Pseudomorpha, if it is at alla pseudomorphine

larva.

During my study I observed larvae only in the oviducts of females, hence all larvae described or

mentioned herein are Ist instar larvae. It is therefore somewhat questionable, whether they can be

compared with the described larvae which are mainly larvae of later states. The number of larvae

observed in a single female is commonly about 5-6, rarely up to about 15. The larvae are coiled together

and tightly packed in the oviducts and do not show very striking differences of size. Even in large

species like Adelotopus dytiscides the observed larvae are rarely longer than 1.5 mm.

6.1. Descriptions of the larvae of the different pseudomorphine genera

Genus Sphallomorpha. For the two known larvae see Moore (1974) and Erwin (1981).

Genus Pseudomorpha. For the three known larvae of the American species see Lenko (1972) and

Erwin (1981). Larvae of the Australian species are still unknown.

Genus Adelotopus. Larvae of the following species have been observed during this study: dytiscides-

group: A. dytiscides, A. ulrichi, A. apicalis, A. zborowskii;, brevipennis-group: A. brevipennis, A. s. sinuaticollis;

marginicollis-group: A. coriaceus, A. seminitidus; politus-group: A. politus, A. doyeni, A. substriatus,

A. haemorrhoidalis, A. minor, A. b. bimaculatus, A. b. angustior, A. languidus; multipunctatus-group:

A. ovatus; similis-group: A. similis; seriepunctatus-group: A. puncticollis angustemaculatus; rubiginosus-

group: A. rubiginosus, A. distinguendus, A. laticollis, A. cribricollis, A. virgatus, A. queenslandicus; laevis-

group: A. laevis, A. c. ciliatus; gyrinoides-group: A. g. gyrinoides, A. d. dubius, A. montorum, A. victoriensis,

A. murrayanus, A. parumpunctatus, A. lunatus, A. rufoguttatus, A. macilentus; fasciatus-group: A. fasciatus;

paroensis-group: A. paroensis; maculipennis-group: A. cuneatus.

Adelotopus dytiscides (Newman)

Figs:2, 3,5-7, 12, 17,18

Length (anterior margin of nasale to apical margin of segment IX): 1.46 mm; width (across head):

0.28 mm; width (across widest abdominal segment): 0.3 mm.

Colour. White, mandibles rufous.

Body shape. Elongate, rather parallel, not decidedly physogastric.

Fig. 2. Adelotopus dytiscides Newman. 1st instar larva, dorsal view. Total length: 1.45 mm.

Fig. 3. Adelotopus dytiscides Newman. 1st instar larva. Head. a. dorsal view. b. ventral view. Scale: 0.1 mm.

Fig. 4. Adelotopus rubiginosus Newman. 1st instar larva. Head. a. dorsal view. b. ventral view. Scale: 0.1 mm.

Head. Rather large and wide, wider than long, laterally about parallel, neck absent. Frontal

sutures broadly v-shaped, joined posteriorly. Nasale slightly sinuate. Antenna elongate, 4-articulate,

3rd antennomere laterally excised, with hyaline bulb, setae partly club-shaped, see fig. 12. Eyes absent.

Mandibles acute, evenly curved, median edge sharp, in middle with small retinaculum, penicillus

absent, laterobasally with normal-shaped seta. Maxilla large, rather elongate, with 4-articulate palpus,

2-articulate, rather elongate galea, without lacinia, but with elongate seta at this place, setae normal-

shaped, see fig. 7. Labium short and wide, anteriorly convex, with 2 terminal, 4 ventral, and 2 dorsal

setae, palpus 2-articulate, rather elongate.

Legs. Rather elongate, all legs with two slightly unequal claws.

Abdomen. Pygopod without crochets, urogomphi absent.

Chetotaxy. Upper surface of head with rather few club-shaped to feebly fungiform setae (Fig. 17),

lower surface with sparse club-shaped setae only. 3rd and 4th antennomeres with few club-shaped

setae, maxilla and labium with normal-shaped setae only. Thorax laterally with few club-shaped setae,

abdomen dorsally and laterally with few short, slightly fungiform setae (Fig. 18), ventrally with some

normal-shaped setae. Terminal segments of abdomen with several elongate, normal-shaped setae.

Differences in certain structures in the larvae of the genus Adelotopus.

In the following enumeration the larvae of the species-groups of Adelotopus are being briefly

characterized with special emphasis to those characters that differ within the genus.

Figs 5-16. Ist instar larvae of different Pseudomorphinae. Mouth parts. 5-6. Adelotopus dytiscides Newman.

5. Labium, ventral view. 6. Mandible. 7-11. Maxilla. 7. Adelotopus dytiscides Newman. 8. A. rubiginosus Newman.

9. A. seminitidus, spec. nov. 10. Cainogenion (s. str.) ipsoides (Westwood). 11. Paussotropus cylindricus Chaudoir).

12-16. Antenna. 12. Adelotopus dytiscides Newman. 13. A. macilentus, spec. nov. 14. A. cuneatus, spec. noV.

15. Cainogenion (s. str.) ipsoides (Westwood). 16. Paussotropus cylindricus Chaudoir).

dytiscides-group. The known larvae of the other species of the dytiscides-group are fairly similar to

that of A. dytiscides.

brevipennis-group. Basically similar to dytiscides-group. Antenna but moderately elongate. Galea

2-articulate. Labial palpus medium sized. Surface of head with club-shaped setae only. Dorsolateral

surface of abdomen with moderately numerous, short, club-shaped setae, lower surface with some

normal-shaped setae.

marginicollis-group. Basically similar to dytiscides-group. Antenna but moderately elongate. Galea

l-articulate (Fig. 9). Labial palpus medium sized. Surface of head with very few faintly club-shaped

setae. Thorax and abdomen with very few club-shaped setae.

politus-group. Very similar to dytiscides-group. Galea 2-articulate. Surface of head with rather few

club-shaped setae, thorax and abdomen dorsolaterally with few club-shaped setae, ventrally with

normal-shaped setae.

multipunctatus- and similis-groups. Similar to politus-group.

seriepunctatus-group. Basically similar to dytiscides-group. Antenna short, 3rd antennomere wide.

Galea 2-articulate, elongate. Mandible rather short. Labial palpi short. Upper surface of head with

club-shaped to slightly fungiform setae, lower surface of head and dorsolateral surfaces of thorax and

abdomen with rather numerous club-shaped setae, all setae remarkably elongate.

rubiginosus-group. Basically similar to dytiscides-group. Antenna rather short and wide. Galea l-artic-

ulate, small (Fig. 8). Lower surface of head with club-shaped setae, upper surface in middle with very

wide fungiform setae, laterally with club-shaped setae (Fig. 4). Thorax and abdomen with regular

pattern of elongate, club-shaped setae, only terminal segments with some normal-shaped setae.

17 18 19 20 21 22 23

Figs 17-23. Types of setae of different Adelotopus larvae. 17-18. A. dytiscides Newman. 17.Seta on head. 18. Seta

onabdomen. 19. A. macilentus, spec. nov., seta on head. 20. A. paroensis Castelnau, seta on head. 21-22. A. fasciatus

Castelnau. 21. Seta on head. 22. Seta on abdomen. 23. A. cuneatus, spec. nov., seta on head.

laevis-group. Similar to rubiginosus-group.

gyrinoides-group. Basically similar to dytiscides-group. Antenna rather elongate, in A. macilentus with

very elongate apical seta (Fig. 13). Galea 2-articulate, elongate. Upper surface of head with narrowly

fungiform setae (Fig. 19), lower surface of head and dorsolateral surfaces of thorax and abdomen with

few club-shaped setae, in A. macilentus also with several short, moderately fungiform setae, ventral

parts with some normal-shaped setae. Penultimate abdominal segment with several very elongate,

normal-shaped setae.

paroensis-group. Basically similar to dytiscides-group. Antenna rather elongate. Galea 2-articulate,

very elongate. Upper surface of head with moderately wide fungiform setae (Fig. 20), lower surface

and dorsolateral surfaces of thorax and abdomen with rather few club-shaped setae, ventral surface

with normal-shaped setae.

fasciatus-group. Basically rather similar to dytiscides-group. Antenna rather short and wide. Galea

2-articulate, elongate. Labial palpus rather elongate. Surface of head with few rather fungiform setae

(Fig. 21), dorsolateral surfaces of thorax and abdomen with few slightly fungiform setae that are

remarkably stout and apically conspicuously serrate (Fig. 22). Lower surface with fairly numerous,

very elongate normal-shaped setae. Penultimate abdominal segmemt with some extremely elongate

normal-shaped setae.

maculipennis-group. Basically similar to dytiscides-group. Antenna short and wide (Fig. 14). Galea

2-articulate, terminal article elongate. Labium short and wide, palpi short. Mandibles rather short.

Legs short. Surface of head with few tridentate fork-shaped setae (Fig. 23). Dorsolateral surfaces of

thorax and abdomen with few faintly fork-shaped setae, lower surface of abdomen with normal-

shaped setae. Penultimate segment of abdomen in middle with some very elongate fork-shaped setae

and several extremely elongate normal-shaped setae.

This short enumeration demonstrates that the larvae of the different groups of Adelotopus are basically

rather similar, but differ mainly in the length of their appendages, the number of articles of the galea,

and in density and structure of head and body chetotaxy. It should be stressed, however, that

unfortunately larvae are not yet known of some of the probably most apomorphic species-groups, ViZ.

the unicolor-, linearis-, and nemosomoides-groups. Moreover, in some of the large groups (e.g. brevipen-

nis- and multipunctatus-groups) only very few larvae are yet known.

Genus Cainogenion. Larvae of the following species have been observed during this study: C. ipsoides,

C. c. creberrimum, C. c. gnaltae, C. obscurum, C. subopacum, C. tropicum. Unfortunately no larvae were

found in C. ephippiatum, the single species of the plesiomorphic subgenus Procainogenion, although

many females were dissected.

Fig. 24. Cainogenion (s. str.) ipsoides (Westwood). 1st instar larva, dorsal view. Total length: 1.85 mm.

Fig. 25. Cainogenion (s. str.) ipsoides (Westwood). 1st instar larva. Head. a. dorsal view. b. ventral view. Scale:

0.1 mm.

Cainogenion ipsoides ipsoides (Newman)

Figs: 10, 15, 24,25

Length (anterior margin of nasale to apical margin of segment IX): 1.8-2.0 mm; width (across head):

0.16-0.18 mm; width (across widest abdominal segment): 0.35-0.4 mm.

Colour. White, mandibles rufous.

Body shape. Moderately elongate, rather physogastric, but widest about in middle of abdomen.

Head. Small, rather narrow and elongate, longer than wide, laterally almost parallel, neck absent.

Frontal sutures narrowly v-shaped, not joined posteriorly. Nasale slightly convex. Antenna moderate-

Iy elongate, 4-articulate, 3rd antennomere laterally excised, with hyaline bulb, setae see fig. 15. Eyes

absent. Mandibles acute, evenly curved, median edge sharp, in middle with small retinaculum,

penicillus absent, laterobasally with normal-shaped seta. Maxilla large, rather elongate, with 4-artic-

ulate palpus, l-articulate, very small galea, without lacinia, setae see fig. 10. Labium narrow, anteriorly

convex, apparently with 2 terminal, 4 short dorsal, and no ventral setae, palpus 2-articulate, rather

elongate.

Legs. Rather short, all legs with two slightly unequal claws.

Abdomen. Pygopod without crochets, urogomphi absent.

Chetotaxy. Surface of head with very few faintly club-shaped or apically very slightly split setae

(Fig. 25). 3rd and 4th antennomeres with few extremely faintly slit setae, maxilla and labium with

normal-shaped setae only. Thorax and abdomen with few normal-shaped setae only. Terminal

segments of abdomen with few elongate, normal-shaped setae.

The larvae of the other examined species are fairly similar to that of C. ipsoides.

Fig. 26. Paussotropus cylindricus (Chaudoir). 1st instar larva, dorsal view. Total length: 0.9 mm.

Fig. 27. Paussotropus cylindricus (Chaudoir). 1st instar larva. Head. a. dorsal view. b. ventral view. Scale:

0.05 mm.

Genus Paussotropus. A single, not well preserved larva of the unique species P. cylindricus (Chau-

doir) has been observed and is described below:

Paussotropus cylindricus (Chaudoir)

Eigs“11,.16,.26,.27

Length (anterior margin of nasale to apical margin of segment IX): c. 0.9 mm; width (across head):

0.085 mm; width (across widest abdominal segment): c. 0.6 mm.

Colour. White, mandibles rufous.

Body shape. Elongate, markedly physogastric.

Head. Narrow and elongate, clearly longer than wide, laterally about parallel, neck absent. Frontal

sutures narrowly v-shaped, posteriorly not joined. Nasale slightly convex. Antenna very short and

stout, 4-articulate, 3rd antennomere very wide, laterally slightly excised, with hyaline bulb, setae see

fig. 16. Eyes absent. Mandibles acute, evenly curved, median edge sharp, apparently without

retinaculum, penicillus absent, laterobasally with normal-shaped seta. Maxilla large, rather elongate,

apparently with only 3-articulate palpus, the terminal palpomere very elongate, galea absent, without

lacinia, setae see fig. 11. Labium narrow and elongate, anteriorly slightly convex, apparently with 2

terminal setae only, palpus 2-articulate, small and rather short.

Legs. Rather short, all legs with two, slightly unequal claws.

Abdomen. Pygopod without crochets, urogomphi absent.

Chetotaxy. Surface of head with few apically faintly split setae (Fig. 27). 3rd and 4th antennomeres

with few normal-shaped setae, maxilla and labium also with normal-shaped setae only. Thorax and

abdomen only ventrally with few normal-shaped setae. Terminal segments of abdomen with few

elongate, normal-shaped setae.

Genus Cryptocephalomorpha. Of this genus larvae are thus far unknown.

Figs 28-31. Photographs. 28. Pseudomorpha (Notopseudomorpha) laevissima Chaudoir. 29. Pseudomorpha (Austro-

pseudomorpha) insignis insignis (Sloane). 30. Adelotopus dytiscides Newman. 31. A. katherinei, spec. nov. Lengths:

6.3 mm; 6.7 mm; 8.8 mm; 5.8 mm.

Figs 32-35. Photographs. 32. Adelotopus brevipennis Macleay. 33. A. rufozonatus, spec. nov. 34. A. atrorufus, spec.

nov. 35. A. marginicollis, spec. nov. Lengths: 5.6 mm; 7.3 mm; 6.8 mm; 5.55 mm.

Figs 36-39. Photographs. 36. Adelotopus coriaceus, spec. nov. 37. A. exactor Darlington. 38. A. politus Castelnau.

39. A. kurandae, spec. nov. Lengths: 6.4 mm; 7.0 mm; 6.8 mm; 4.95 mm.

Figs 40-43. Photographs. 40. Adelotopus languidus, spec. nov. 41. A. ovatus, spec. nov. 42. A. geminus, spec. NOV.

43. A. nitidior, spec. nov. Lengths: 6.0 mm; 5.6 mm; 6.3 mm; 5.6 mm.

Figs 44-47. Photographs. 44. Adelotopus obsoletus, spec. nov. 45. A. villosus, spec. nov. 46. A. similis, spec. nov.

47. A. tasmani Blackburn. Lengths: 5.2 mm; 6.0 mm; 7.5 mm; 5.5 mm.

Figs 48-51. Photographs. 48. Adelotopus nigricauda, spec. nov. 49. A. seriepunctatus striatus subspec. nov.

50. A. puncticollis angustemaculatus, subspec. nov. 51. A. virgatus, spec. nov. Lengths: 4.85 mm; 5.55 mm; 4.8 mm;

5.5 mm.

65)

Figs 52-55. Photographs. 52. Adelotopus queenslandicus, spec. nov. 53. A. angustatus, spec. nov. 54. A. cıiliatus

tenuipunctatus, subspec. nov. 55. A. unicolor, spec. nov. Lengths: 4.95 mm; 5.6 mm; 5.2 mm; 4.65 mm.

Figs 56-59. Photographs. 56. Adelotopus linearis Macleay. 57. A. bacillus, spec. nov.

59. A. gyrinoides orientalis, subspec. nov. Lengths: 4.8 mm; 5.2 mm; 4.65 mm; 5.5 mm.

dal:

celeripes Lea.

Figs 60-63. Photographs. 60. Adelotopus dubius glaber, subspec. nov. 61. A. rufoguttatus

Castelnau. 63. A. punctulifer, spec. nov. Lengths: 5.6 mm; 4.85 mm; 4.25 mm; 5.55 mm.

(Blackburn). 62. A. affınis

Figs 64-67. Photographs. 64. Adelotopus analis Macleay. 65. A. paroensis Castelnau. 66. A. fasciatus Castelnau.

67. A. longiformis, spec. nov. Lengths: 4.4 mm; 5.35 mm; 4.0 mm; 5.5 mm.

Figs 68-71. Photographs. 68. Adelotopus maculipennis Macleay. 69. Cainogenion (Procainogenion) ephippiatum

(Castelnau). 70. Cainogenion (s. str.) ipsoides ipsoides (Westwood). 71. C. (s. str.) interiore, spec. nov. Lengths:

3.9 mm; 4.5 mm; 7.2 mm; 6.4 mm.

Figs 72-75. Photographs. 72. Cainogenion (s. str.) depressum, spec. NOV. 73. Paussotropus cylindricum (Chaudoir).

74. Cryptocephalomorpha maior, spec. nov. 75. C. papua Darlington. Lengths: 6.6 mm; 5.9 mm; 4.4 mm; 3.0 mm.

7. Classification

7.1. Subfamily Pseudomorphinae

For extensive synonymy and for diagnosis of the subfamily see part 1. of revision.

7.1.1. Key to the genera of Pseudomorphinae

The key to the genera is basically the same as in part 1 of the revision. However, because in the

meantime the genus Pseudomorpha has been discovered in Australia, two new subgenera of Pseudomor-

pha are being described herein, and the genus Cryptocephalomorpha has been found in South Africa, the

key must be slightly altered. For the benefit of the reader it is repeated here:

Eyes dorsal and with a continuous border beneath. Australia, New Guinea, Solomon Islands,

SOUth-EaSt Asa a Re Adelotopus Hope

Eyes dorsal or lateral, when more or less dorsal, without a continuous border beneath ............ 2:

Lateral part of maxilla developed into a large, conspicuous lobe. Beneath eye with triangular

process or conspicuous, large concavity. Tibiae distinctly compressed. Dorsal surface very coarsely

punctate .....uaenesessnssssensenenenensennneneenenssnsenentnnsnsenonontnensnssnsnsosesensnensansnsnsnsssensnsnsnsesesenentnertnenenenenentonenenenene 3.

Lateral part of maxilla inconspicuous. No conspicuous process or concavity beneath eye. Tibia not

distinctly compressed. Surface finely punctate or at most with scattered coarse punctures, then

each puncture bearing an elongate bristle .........usesnesensnensenensenensensenensnnsnsenensenennesenenensenennenne 4.

Beneath eye with triangular process. Tibiae less compressed. Tarsi less thickened. Prosternal

process short. Australia ..........esesesesesenensnensenenenenenenensnensenenenenensnsnsnsnsnenenensnenennen Cainogenion Notman

Beneath eye with large circular concavity. Tibia very compressed. Tarsi thick. Prosternal process

absent Australlan..en.cenere sen eessesssnesnsunesenrsnntenssessrteocs rennen enrhesneseenennene Paussotropus Waterhouse

Head ventrally deflexed. Labrum not visible. Mandibles tiny. Small size, body form very convex,

egg-shaped species. South East Asia, New Guinea, Solomon Islands, northern Australia, South

N eo ber BR ERBocrern ERRoren Feranuikken Cryptocephalomorpha Ritsema

Head prognathous. Labrum visible. Mandibles normal. Body form either cylindrical and elongate,

or rather depressed and wide... eesssssstsneueneenecen enenteereetnneneensen ern

Margins of elytra with fringe of elongate setae. Dorsal surface commonly with scattered, elongate

bristles. Mental tooth elongate, very acute. America, Australia. ............o- Pseudomorpha Kirby 6.

Margins of elytra without fringe of elongate setae. Dorsal surface without elongate bristles, at most

with short pilosity. Mental tooth either widely rounded, or slightly bidentate, or absent. Australia,

NEWEGUINe A RR Sphallomorpha Westwood

Anterior margin of head without incision laterally of clypeus. Southern South America .........

rennen dosade alu ntennenan nenne ses Tre ernennen roter Notopseudomorpha, subgen. nov.

Anterior margin of head with incision laterally of celypeus ........uenssssesessenenessnenennsnsnsnesnsesanaenene ?.:

Body form narrow, cylindrical. Labrum 6-setose. Metepisternum quadrate, wings reduced. Aus-

a ERRer roscacace Austropseudomorpha, subspec. nov.

Body form usually wider, not cylindrical. Labrum 4-setose. Metepisternum elongate, wings well

developed. Ameriea......urnseasescsssssunnanssesnenearsesnsehncnareereensnenreee eat a reer ses sseestcat Pseudomorpha s. str.

7.2. Genera

Notman (1925) characterized the then known pseudomorphine genera and provided a key. In the

meantime, the African genus Hydroporomorpha Westwood has been removed from Pseudomorphinae

to the harpaline subtribe Stenolophina (see Baehr 1992a). Adelopomorpha Heller, included in Pseudo-

morphinae by Csiki (1933), was suspected to belong to Oodinae (Baehr 1992a). In the meantime

examination of the type definitely revealed this status.

The other genera are well keyed out in Notman’s key. Only two species have to be removed from

the genera in which they were originally included, namely Cainogenion ceylindricum (Chaudoir) to genus

Paussotropus, and Adelotopus insignis Sloane to genus Pseudomorpha. Although the large genus Adelo-

topus is taxonomically rather heterogenous, for the present I prefer to include all species in this genus,

rather than to erect several new genera. In my view this would be a procedure not justified in a group

passing through a period of very rapid evolution, as Adelotopus does at present. Hence I prefer to

divide the species in certain species-groups that could be later raised to a nomenclatorial valid status

as subgenera or even genera, when better knowledge of the larvae will perhaps throw more light on

the generic concept.

7.2.1. Genus Pseudomorpha Kirby

Pseudomorpha Kirby, 1825, p. 98; Westwood 1853, p. 395; Lacordaire 1854, p. 151; Notman 1925, p. 12, 30; Csiki

1933, p. 1637; Moore et al. 1987, p. 61.

Heteromorpha Kirby, 1825, p. 109.

Axinophorus Dejean & Boisduval, 1829, p. 60, 174.

Drepanus Dejean, 1831, p. 434.

Type species: Pseudomorpha excrucians Kirby, 1825, by monotypy.

Diagnosis. Genus of Pseudomorphinae, delimited by following characters: Body fairly wide to almost

cylindrical, elytra posteriorly gently convex; head prognathous; eyes situated laterally, without ventral

border, ventral part more or less triangular; clypeus partly or completely fused to frons; labrum

separated from clypeus by a sulcus; supraorbital, clypeal, suborbital, and gular setae present,

preorbital seta absent; antennal grooves deep; lateral plate of maxilla not enlarged; antenna elongate,

basal antennomere simple; mental tooth elongate, triangular; glossa fused with paraglossae to a wide

plate, bisetose but sometimes with additional elongate setae; labial palpi very large, markedly

securiform; ventral surface of head large, not concealed by the mouth parts; prosternal process

straight, rather short, depressed between coxae; number of umbilical pores of elytra variable; femora

moderately or strongly compressed, with deep grooves; tibiae and tarsi not compressed, elongate;

d protarsus biseriately clothed at 1st and 2nd tarsomere, mesotarsus uniseriately clothed at Ist and 2nd

tarsomere or not clothed; d sternum VII not excised; d sternum VIII apically divided, highly

asymmetric; aedeagus with simply folded internal sac; parameres fairly similar, though left paramere

always considerably larger; ? stylomeres 1 and 2 separated, though shape very variable; no distinct

dorsal and ventral ensiform setae present, but nematiform setae present though sometimes very short

and not always arising from a pit.

Larvae. 1st to 3rd instar larvae known of some American but not yet from any of the Australian

species.

Distribution. Southwestern part of North America, Central America including the Caribbean area,

and western part of South America to Argentinia; southern Australia.

Systematic position. Pseudomorpha is probably the adelphotaxon of all other pseudomorphine genera

except for Sphallomorpha.

Note. The Australian species differ rather conspicuously in several respects from the American

species. Hence a new subgenus Austropseudomorpha is proposed to accomodate the Australian species.

It should be stressed, however, that the remaining (American) Pseudomorpha are an assemblage of

species of very different phylogenetic stages including one group of (so far) two species that are

plesiomorphic in many characters and do not much differ in some respects from Australian Sphallo-

morpha and the most plesiotypic Adelotopus. The other American species, however, are more or less

apomorphic in external as well as in genitalic characters and form distinctive species-groups. The

Australian species are more closely related to these evolved species-groups of American Pseudomorpha,

but constitute a highly aberrant offshoot of this group (Baehr 1994a).

Although I am at present not prepared to do a revision of the American Pseudomorpha, for

nomenclatorial correctness the unique group of very plesiomorphic South American species (namely

P. laevissima Chaudoir and P. glabra Ogueta) cannot be longer included in Pseudomorpha proper, when

the Australian species are separated in an own subgenus, but must be (at least) included in a separate

subgenus that is named Notopseudomorpha.

A future revisor of the American Pseudomorpha should pay attention to these different phylogenetic

stages within the genus, and for that reason a revisor should perhaps definitely divide the genus into

separate genera.

7.2.1.1. Subgenus Notopseudomorpha, subgen. nov.

Type species: Pseudomorpha laevissima Chaudoir, 1852 by present designation.

Diagnosis (see figs 1, 28, 76, 97, 284): Subgenus of Pseudomorpha, delimited by following characters:

Body fairly convex; anterior margin of head without excision near clypeus; clypeus partly fused to

frons; labrum 4-setose; supraorbital, preorbital, and gular setae absent; basal antennomere of antenna

simple; glossa bisetose; labial palpi markedly securiform; head without additional long setae;

pronotum without elongate setae along margin and on disk, without large punctures, but with

posterior marginal seta; prosternal process straight, rather short, depressed between coxae; elytra with

elongate setae along margin, without setae and large punctures on surface; umbilical pores of elytra

well discernable, numerous; femora strongly compressed, with deep grooves; d sternum VIII apically

not divided, only slightly asymmetric; aedeagus rather short, with simply folded internal sac, apex of

internal sac with a narrow sclerite; parameres fairly similar, though left paramere considerably larger,

both without setae at apex; ? stylomeres 1 and 2 separated, not very elongate; stylomere 2 of normal

shape, slightly dentiform; 2 elongate nematiform setae present and arising from a pit.

Larvae. The larva of P. laevissima Chaudoir was described and figured by Lenko (1972).

Distribution. South America: southern Brazil, Paraguay, northern Argentinia.

Systematic position. In many respects it is the plesiotypic adelphotaxon of all other species of

Pseudomorpha. So far two species, Pseudomorpha laevissima Chaudoir and P. glabra Ogueta, belong to this

subgenus. They are in particular plesiomorphic in following character states: absence of excision at

anterior border of head; absence of punctures and setae on disk of head, prothorax, and elytra; absence

of fringe of marginal setae on pronotum (apart from regular posterior marginal seta in basal angle));

almost symmetric, apically not as deeply excised d sternum VIII; and comparatively primitive, normal-

shaped ? stylomere 2 that is comparable to that occurring in Sphallomorpha.

7.2.1.2. Subgenus Austropseudomorpha, subgen. nov.

Type species: Adelotopus insignis Sloane, 1910 by present designation.

Diagnosis. Subgenus of Pseudomorpha, delimited by following characters: Body elongate and almost

cylindrical; ventral part of eyes less triangular than in the American species; anterior margin of head

with deep excision near clypeus; clypeus completely fused to frons; labrum 6-setose; one supraorbital

seta only present; all fixed setae of head very elongate; antennal grooves deep and elongate; basal

antennomere of antenna remarkably expanded to a large bulbus and with some pilosity; mental tooth

slightly pointed down; glossa fused with paraglossae to a wide plate, with several additional elongate

setae; pronotum elongate, rather cylindrical, at base as wide as or narrower than at apex; lateral margin

of pronotum with some short, but without very elongate setae; number of umbilical pores of elytra

reduced to 8 or 9 and arranged in two groups; dorsal surfaces of pronotum and elytra without very

elongate setae; elytra fused together, wings absent, metepisternum very short; femora but moderately

compressed; d mesotarsus not clothed beneath; aedeagus narrow and very elongate; ? stylomeres 1

and 2 both very narrow and elongate, stylomere 1 spiniform, with only a single, short seta at apex.

Larvae. Unknown.

Distribution. Australia. So far 3 species and one additional subspecies are included in this subgenus.

Systematic position. This subgenus is probably the adelphotaxon of all American Pseudomorpha

except for those of the subgenus Notopseudomorpha. Austropseudomorpha is thus a rather basally

branching offshoot of Pseudomorpha, but has many highly specialized character states.

7.2.1.3. Description of Austropseudomorpha

Because all species of Austropseudomorpha are very similar and differ only in minor characters, no

detailed description of the subgenus beyond the above diagnosis is given, though the description of

P. (A.) insignis (Sloane) should be consulted.

7.2.1.4. Key to the Australian species of genus Pseudomorpha Notman,

subgenus Austropseudomorpha, subgen. nov.

1. Generally larger, more elongate species, length >6 mm, ratios w/l of pronotum <1.15, 1/w of elytra

>1.55. Lateral margins of pronotum almost parallel, base as wide as apex (Figs 285-287) .......... 23

- Generally smaller, shorter species, length <5.5 mm, ratios w/l of pronotum >1.18, 1/w of elytra

<1.45. Lateral margins of pronotum evenly convex, base clearly narrower than apex (Fig. 288). sw.

WER. nee ee ee RR a Ne BE brevis, spec. NOV.

2. Apical angles of pronotum angulate, protruding, apex in middle but slightly convex. Excision

laterally of clypeus deeper, lateral angle of excision distinctly protruding. Eyes less protruding.

Median antennomeres longer, >2 x as long as wide (Figs 80c, 81c). nw. Vic, s. NSW... 3.

- Apical angles of pronotum obtuse, barely protruding, apex in middle markedly convex. Excision

laterally of clypeus shallower, lateral angle of excision barely protruding. Eyes more protruding.

Median antennomeres shorter, <2 x as long as wide (Fig. 82c). s. WA ...... subangulata, spec. nov.

3. Pronotum and elytra with distinct but fine puncturation between the large punctures. Pilosity of

surface very short, somewhat declined. Aedeagus longer, lower surface not evenly concave (Fig.

BIS)SWAVIcH en. are een lehnte insignis insignis (Sloane)

- Pronotum and elytra without fine puncturation between the large punctures. Pilosity of surface

elongate, erect. Aedeagus shorter, lower surface evenly concave (Fig. 818). 5. NSW...

UUHEHRNEHRER ER EDRSIENSORGNG HAB SAELEEEE SS HMI EET ER RL A nl insignis pilosa, subspec. nov.

7.2.1.5. The species of Austropseudomorpha, subgen. nov.

Pseudomorpha insignis (Sloane, 1910) (new combination)

This species includes two subspecies, one in northwestern Victoria, the other in southern New South

Wales. Due to very scarce material, however, this taxonomic decision is somewhat doubtful and may

be altered in one or another way, when sufficient material is at hand.

Diagnosis. Large, cylindrical species, distinguished from related species by large size, rather parallel

lateral margins of pronotum, angulate, protruding apical angles of pronotum, deep excision in anterior

margin of head with protruding lateral angle, elongate antenna, longer, rather narrow left paramere,

and elongate stylomere 2.

Pseudomorpha insignis insignis (Sloane, 1910)

Figs 1, 29, 77-80, 97, 285, 437, 602

Adelotopus insignis Sloane, 1910, p. 405; Notman 1925, p. 6, 29; Csiki 1933, p. 1635; Moore et al. 1987, p. 51.

Types. Holotype: d, Sea Lake Goudie, 250. 5., Adelotopus insignis Sl. mss. Type; A. insignis Sl. Holotype P 19,

Holotype (ANIC).

Type locality. “Sea Lake”, Victoria.

Diagnosis. Distinguished from P. insignis pilosa, subspec. nov. by larger size, less protruding apical

angles of pronotum, presence of extremely fine additional puncturation of pronotum and elytra, very

short, obliquely decumbent pilosity of surface, and longer, on ventral surface not evenly concave

aedeagus.

Description

Measurements. Length: 6.7-7.1 mm. Ratios. Width/length of pronotum: 0.95-1.06; width prono-

tum/head: 1.29-1.32; length/width of elytra: 1.60-1.63; length elytra/pronotum: 1.74-1.77.

Colour. Reddish-piceous to almost black, in dark specimens margins of pronotum and elytra and

elytral suture faintly reddish translucent. Lower surface reddish to dark piceous, posterior abdominal

sterna at apex slightly lighter. Mouth parts, antenna and legs dark reddish to piceous.

Head (Figs 77, 78, 80a-c, 285). Rather short and wide, frons faintly convex, between eyes with a

shallow transverse impression. Lateral margin of head near eyes convex, strongly bordered, near

clypeus deeply excised, the lateral angle of excision distinctly protruding. Clypeus completely fused

to frons, anterior border very gently convex, slightly swollen, on lateral corner with a very elongate

seta originating from a large and deep groove. Labrum short and wide, separated from clypeus by a

deep furrow, base slightly overlapped by apex of clypeus, labrum directed anteriorly, apex gently

convex, 6-setose. Eye rather large, gently surpassing lateral margin of head, outline rather ovalish,

orbit evenly convex, rather gentle. An elongate supraorbital seta present a considerable distance inside

of eye. Ventral part of eye surpassing lateral margin of head, but not as triangular as in the American

Pseudomorpha. Antennal groove very deep, elongate, medially and laterally sharply bordered. Mental

tooth rather large, triangular, apex slightly obtuse, somewhat pointed down. Wings of mentum large

and wide, apex obtusely rounded. Glossa and paraglossae completely fused to a very wide plate with

gently convex apex that is ventrally strongly keeled. Glossa at apical margin medially with 2 very

elongate setae, laterally on either side with c. 6-8 elongate setae, also dorsal surface with elongate setae.

Terminal palpomere of maxillary palpus elongate, narrow, parallel. Terminal palpomere of labial

palpus very large and wide, markedly securiform. Both palpi sparsely pilose. Lateral plate of maxilla

of normal size. Ventral surface of head elongate. A very elongate gular seta present on either side

shortly behind base of mentum. Antenna elongate, slightly depressed, 7th-8th antennomeres >2.5 x as

long as wide, Ist antennomere remarkably enlarged, bulbose, at lateral tip with the usual elongate seta

and with additional short setae. Microreticulation absent, surface with very scattered coarse punctures

and with an extremely fine, rather dense puncturation within the large punctures that each bears a

very short, erect to slightly declined hair. Surface rather glossy. Laterally and below eye with a group

of 12-15 strong, elongate setae. Gula sparsely and shortly pilose.

Pronotum (Fig. 285). Rather narrow, about quadrate, dorsal surface evenly convex, lateral margins

not explanate. Base about es wide as apex. Apex gently convex, though apical angles distinctly

produced and obtusely angular. Apex finely margined. Sides in middle almost parallel or very gently

convex, strongly margined, lateral channel absent. Basal angles evenly rounded off, base in middle

markedly concave, distinctly margined. Whole surface regularly convex, without median line. Apex

near apical angles within the widened margin with 4-5 strong setae, lateral margin on and below

border with few, short setae. Microreticulation absent, surface with scattered moderately coarse

punctures and with extremely fine, dense puncturation within the large punctures that each bears a

very short erect to slightly declined hair. Surface moderately glossy.

Elytra (Figs 29, 285, 437). Rather elongate, parallel, rather cylindrical. Apex wide, truncature gently

convex, lateral apical angles widely rounded off. Base transverse, shoulders rectangular though

Shorty rounded off. Basal margin attaining half of distance to suture, base in middle with a deep

transverse impression. Basal border wide, marginal channel very narrow, not concealed. Basal border

Fig. 76. Pseudomorpha (Notopseudomorpha) laevissima Chaudoir. Ventral view of head.

Figs 77-79. Pseudomorpha (Austropseudomorpha) insignis insignis (Sloane). 77. Frontal view of head. 78. Ventral

view of head. 79. Base of left antenna.

with two rows of strong, elongate setae, also lateral margin slightly below border with a series of

elongate setae. Series of umbilical pores consisting of 4-5 pores behind shoulder and 3-4 widely spaced

pores in posterior third and at apex. Pores large, markedly umbilicate, easily to see, setae elongate.

Scutellar pore present, large, umbilicate, seta elongate. Striae including sutural stria absent, but

position of striae marked by irregular rows of moderately coarse, widely spaced punctures that each

bears a short, slightly declined hair. Punctures much more coarse than those on pronotum. Microre-

ticulation absent, between large punctures with extremely fine, dense puncturation. Surface rather

glossy. Elytra tighly fused together, wings reduced.

Lower surface. Prosternal process rather short, moderately surpassing procoxae, surface de-

pressed, margined inside of coxae, apex straight, rather depressed, surface shortly pilose. Metepister-

num very short, rectangular, almost half wider than long. Abdominal sterna with a pair of elongate

setae. Terminal sternum in both sexes with 3, rarely unilaterally 2 elongate setae at some distance from

apex. Lower surface glossy, sparsely punctate and setose.

Legs. Moderately elongate, 1st tarsomere of protarsus slightly longer than wide, tibial groove of

profemur moderately deep, symmetric, anterior border straight. Femora comparatively narrow and

elongate, tibiae elongate, not at all widened. Metatibia elongate, c. 8 x as long as wide, Ist tarsomere

of metatarsus c. 2x as long as wide. d protarsus not widened, Ist and 2nd segents biseriately

squamose, mesotarsus not squamose.

d genitalia (Figs S0d-i). Genital ring moderately wide, ovalish, slightly asymmetric, with rather

elongate, incurved apex, with large, asymmetric, deeply excised base. Sternum VII rather narrow,

apically divided, with very deep excision, base slightly bisinuate, basal angles acute, lateral parts

elongate, markedly triangular. Aedeagus very elongate, narrow, depressed, slightly narrowed towards

apex, faintly asymmetric. Basal part long, markedly bent. Lower surface in basal % almost straight,

then more concave. Apex narrow, obtusely acute. Orifice very elongate, internal sac rather simly

folded. Both parameres narrow, elongate, with rounded apex, left paramere considerably larger than

right.

Figs 80a-l. Pseudomorpha (Austropseudomorpha) insignis insignis (Sloane). Details of head and genitalia. a. Lower

surface of terminal palpomeres of maxillary palpus. b. Lower surface of terminal palpomeres of labial palpus.

c. 5th-11th antennomeres. d. d sternum VII. e. d genital ring. f. Lower surface of aedeagus. g. Lateral view of

aedeagus. h. Right paramere. i. Left paramere. k. ? sternum VII. 1. ? stylomeres and lateral plate.

? genitalia (Figs SOk,l). Apex of sternum VIII rather short and wide, markedly triangular, laterally

acute, basal process rather narrow and elongate. Both stylomeres very narrow and elongate, stylomere

1 completely divided, stylomere 2 spine-shaped, at apex with 1 short seta. Lateral plate elongate,

rectangular, with 2-3 short medial-apical setae.

Variation. Little variation noted in relative width of pronotum and elytra only.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. One specimen collected in “drift fence pitfall trap”. Dated specimens

captured in January and April.

Distribution (Fig. 602). Northwestern Victoria.

Material examined (3). Vic: 13, Sea Lake Goudie, 250. 5., Adelotopus insignis Sl. mss. Type; A. insignis Sl.

Holotype P 19 (ANIC); 17, Sea Lake 4.1916 G. Goudie, 789 (SAMA),; 14, 14.4 km SE of Walpeup, 1.1986 35°11S

142°11’E, Site 59, A. L. Yen, Adelotopus sp. A, Silphomorpha sp. 2 (NMV).

Pseudomorpha insignis pilosa, subspec. nov.

Figs 81, 286, 602

Types. Holotype: d, Weddin Mt. 26.1X.23 (ANIC). — Paratypes: 19, Blue Mts. N. S. Wales, Glenbrook (MMS); 19,

Australien, NSW 112, Colo River, 20 km n. Windsor, 7.-8.12.1990, M. Baehr (CBM).

Diagnosis. Distinguished from P. i. insignis (Sloane) by lesser size, slightly more protruding apical

angles of pronotum, absence of extremely fine additional puncturation of pronotum and elytra, rather

elongate, erect pilosity of surface, and shorter, on ventral surface evenly concave aedeagus.

Figs 81f-i. Pseudomorpha (Austropseudomorpha) insignis pilosa, subspec. nov. Details of d genitalia. For legends

see fig. 80.

Figs 82a-c, 1. Pseudomorpha (Austropseudomorpha) subangulata, spec. nov. Details of head and ? genitalia. For

legends see fig. 80.

Description

Measurements. Length: 6.0-6.7 mm. Ratios. Width/length of pronotum: 1.09-1.15; width prono-

tum/head: 1.28-1.30; length/width of elytra: 1.53-1.65; length elytra/pronotum: 1.80-1.81.

Colour. Similar to nominate subspecies.

Head. Rather similar to nominate subspecies, but large punctures coarser, fine puncturation

absent, surface therefore glossier, and pilosity longer and erect.

Pronotum (Fig. 286). Rather similar to nominate subspecies, but slightly wider, apical angles

slightly more produced, pilosity of lateral margin longer and more conspicuous, large punctures of

surface coarser, fine puncturation absent, surface therefore glossier, and pilosity longer and erect.

Elytra (Fig. 286). Rather similar to nominate subspecies, but pilosity of lateral margins longer and

more conspicuous, punctures of striae markedly coarser, fine puncturation of surface absent, therefore

surface glossier, and pilosity longer and erect.

Lower surface. Similar to nominate subspecies.

Legs. Similar to nominate subspecies.

d genitalia (Figs 81f-i). Similar to nominate subspecies, but aedeagus shorter, lower surface evenly

concave, and right paramere with slightly widened, transversely cut apex.

? genitalia. Similar to nominate subspecie.

Variation. Some variation noted in relative shape of pronotum and elytra, otherwise very

homogeneous.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. So far collected in September and December, one paratype collected by me

at light in open eucalypt forest.

Distribution (Fig. 602). Southern New South Wales.

Material examined (3). Only the type series.

Etymology. The name refers to the distinct pilosity of the surface.

Pseudomorpha subangulata, spec. nov.

Figs 82, 287, 602

Types. Holotype: ?, AUSTRALIA: WA, Thomas River, Cape Arid,N. Pk. 1.-5.X.1981 H. & A. Howden (ANIC).

Diagnosis. Moderately large, cylindrical species, distinguished from related species by moderate size,

rather parallel lateral margins of pronotum, barely rectangular, not protruding apical angles of

pronotum, more shallow excisions in anterior margin of head with lateral angle of excision barely

protruding, shorter antenna, and rather elongate stylomere 2.

Description

Measurements. Length: 5.9 mm. Ratios. Width/length of pronotum: 1.05; width pronotum/head:

1.22; length/ width of elytra: 1.61; length elytra/pronotum: 1.75.

Colour. Whole beetle including mouth parts, antenna, and legs reddish-piceous.

Head (Figs 82a-c). Rather short and wide, frons faintly convex, without distinct transverse

impression between eyes. Lateral margin of head near eyes convex, strongly bordered, near clypeus

moderately deeply excised, the lateral angle of excision barely protruding. Clypeus completely fused

to frons, anterior border straight, faintly swollen, on lateral corner with a very elongate seta originating

from a large and deep groove. Labrum short and wide, separated from clypeus by a deep furrow, base

slightly overlapped by apex of clypeus, labrum directed anteriorly, apex almost straight, 6-setose. Eye

rather large, gently surpassing lateral margin of head, outline rather triangular, orbit almost perpen-

dicular, rather large. An elongate supraorbital seta present a considerable distance inside of eye.

Ventral part of eye surpassing lateral margin of head, but not as triangular as in the American

Pseudomorpha. Antennal groove very deep, elongate, medially and laterally sharply bordered. Mental

tooth rather large, triangular, apex narrow but transversely cut, somewhat pointed down. Wings of

mentum large and wide, apex obtusely rounded. Glossa and paraglossae completely fused to a very

wide plate with gently convex apex that is ventrally remarkably strongly keeled. Glossa at apical

margin medially with 2 very elongate setae, laterally on either side with c. 6 elongate setae, also dorsal

surface with elongate setae. Terminal palpomere of maxillary palpus elongate, narrow, parallel.

Terminal palpomere of labial palpus very large and wide, markedly securiform. Both palpi sparsely

pilose. Lateral plate of maxilla of normal size. Ventral surface of head elongate. A very elongate gular

seta present on either side shortly behind base of mentum. Antenna moderately elongate, slightly

depressed, 7th-8th antennomeres <2 x as long as wide, Ist antennomere remarkably enlarged, bulbose,

at lateral tip with the usual elongate seta and with additional short setae. Microreticulation absent,

surface with very scattered, rather coarse punctures and with irregular, finest traces only of an

extremely fine puncturation within the large punctures that each bears a rather short, anteriorly

inclined hair. Surface glossy. Laterally and below eye with a group of c. 10 strong, elongate setae. Gula

anteriorly sparsely and shortly pilose, posteriorly smooth.

Pronotum (Fig. 287). Rather narrow, about quadrate, dorsal surface evenly convex, lateral margins

not explanate. Base about es wide as apex. Apex rather convex, apical angles slightly less than

rectangular, not produced, obtusely angular. Apex finely margined. Sides in middle almost parallel,

strongly margined, lateral channel absent. Basal angles evenly rounded off, base in middle markedly

concave, distinctly margined. Whole surface regularly convex, without median line. Apex near apical

angles within the widened margin with 4-5 strong setae, lateral margin on and below border with very

few, short setae. Microreticulation absent, surface with scattered, rather coarse punctures and with

finest, very superficial traces of an extremely fine puncturation only within the large punctures that

each bears a very short erect to slightly declined hair. Surface rather glossy.

Elytra (Fig. 287). Rather elongate, parallel, rather cylindrical. Apex wide, truncature gently convex,

lateral apical angles widely rounded off. Base transverse, shoulders rectangular though shortly

rounded off. Basal margin attaining half of distance to suture, base in middle with a deep transverse

impression. Basal border wide, marginal channel very narrow, not concealed. Basal border with two

rows of strong, elongate setae, also lateral margin slightly below border with a series of elongate setae.

Series of umbilical pores consisting of 5 pores behind shoulder and 4 widely spaced pores from middle

to posterior angles, but non at apex. Pores large, markedly umbilicate, easily to see, setae elongate.

Scutellar pore present, large, umbilicate, seta elongate. Striae including sutural stria absent, but

position of striae marked by irregular rows of very coarse, widely spaced punctures that each bears

a short, slightly declined hair. Punctures much more coarse than those on pronotum. Microreticulation

almost absent, but faintest, extremely superficial traces still visible under very high magnification.

Surface glossy. Elytra tighly fused together, wings reduced.

Lower surface. Prosternal process rather short, moderately surpassing procoxae, surface de-

pressed, margined inside of coxae, apex straight, rather depressed, surface shortly pilose. Metepister-

num very short, rectangular, almost half wider than long. Abdominal sterna with a pair of elongate

setae. Terminal sternum in $ with 2 elongate setae at some distance from apex. Lower surface glossy,

sparsely punctate and setose.

Legs. Moderately elongate, Ist tarsomere of protarsus slightly longer than wide, tibial groove of

profemur moderately deep, symmetric, anterior border straight. Femora comparatively narrow and

Figs 83. Pseudomorpha (Austropseudomorpha) brevis, spec. nov. Details of head and genitalia. For legends see fig.

80.

elongate, tibiae elongate, not at all widened. Metatibia elongate, c. 8 x as long as wide, Ist tarsomere

of metatarsus c. 2 x as long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Figs 821). Apex of sternum VIII as in P. insignis, rather short and wide, markedly

triangular, laterally acute, basal process rather narrow and elongate. Both stylomeres very narrow and

elongate, stylomere 1 completely divided, stylomere 2 spine-shaped, laterally slightly concave, at apex

with 1 short seta. Lateral plate elongate, rectangular, with 3-4 short medial-apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Holotype collected in October.

Distribution (Fig. 602). Southwestern Australia. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the less angulate apical angles of the prothorax compared with P. insignis.

Pseudomorpha brevis, spec. nov.

Figs 83, 288, 602

Types. Holotype: 2, 1878, Mt. Barker W.A., Lea (SAMA). - Paratypes: 19, 17, Cape Freycinet (34.06 S, 114.59 E)

Western Australia 15-18 Nov 1986 T. F. Houston 639c-18, ex pitfall trap in heath (WAM 94/876-877, CBM).

Diagnosis. Short, cylindrical species, distinguished from related species by usually smaller size,

wider pronotum with rather oblique, distinctly convex lateral margins and barely rectangular, not

protruding apical angles, shorter elytra, more shallow excisions in anterior margin of head with lateral

angle of excision barely protruding, short antenna, short, rounded left paramere, and short sty-

lomere 2.

Description

Measurements. Length: 4.3-5.4 mm. Ratios. Width/length of pronotum: 1.18-1.21; width prono-

tum/head: 1.31-1.35; length/ width of elytra: 1.36-1.40; length elytra/pronotum: 1.65-1.75.

Colour. Reddish-piceous to black, margins of pronotum and elytra and elytral suture sometimes

faintly reddish translucent. Lower surface reddish. Mouth parts, antenna and legs reddish.

Head (Figs 83a-c). Rather short and wide, frons faintly convex, without distinct transverse

impression between eyes. Lateral margin of head near eyes convex, strongly bordered, near clypeus

with rather shallow excision, the lateral angle of excision not protruding. Clypeus completely fused to

frons, anterior border very gently convex, faintly swollen, on lateral corner with a very elongate seta

originating from a large and deep groove. Labrum short and wide, separated from clypeus by a deep

furrow, base slightly overlapped by apex of clypeus, labrum directed anteriorly, apex almost straight,

6-setose. Eye rather large, gently surpassing lateral margin of head, outline rather ovalish, orbit

oblique, gently convex, rather feeble. An elongate supraorbital seta present at considerable distance

medially of eye. Ventral part of eye surpassing lateral margin of head, but not as triangular as in the

American Pseudomorpha. Antennal groove very deep, elongate, medially and laterally sharply bor-

dered. Mental tooth rather large, triangular, apex obtuse, somewhat pointed down. Wings of mentum

large and wide, apex obtusely rounded. Glossa and paraglossae completely fused to a very wide plate

with gently convex apex that is ventrally strongly keeled. Glossa at apical margin medially with 2 very

elongate setae, laterally on either side with c. 6 elongate setae, also dorsal surface with elongate setae.

Terminal palpomere of maxillary palpus elongate, narrow, parallel. Terminal palpomere of labial

palpus very large and wide, markedly securiform. Both palpi sparsely pilose. Lateral plate of maxilla

of normal size. Ventral surface of head elongate. A very elongate gular seta present on either side

shortly behind base of mentum. Antenna comparatively short, slightly depressed, 7th-8th antennomer-

es <1.5 x as long as wide, Ist antennomere remarkably enlarged, bulbose, at lateral tip with the usual

elongate seta and with additional short setae. Microreticulation absent, surface with few, very

scattered, rather coarse punctures that each bears a rather short, erect to anteriorly inclined hair,

without any additional puncturation. Surface glossy. Laterally and below eye with a group of 10-12

strong, elongate setae. Gula anteriorly sparsely and shortly pilose, posteriorly smooth.

Pronotum (Fig. 288). Comparatively wide, dorsal surface evenly convex, lateral margins not

explanate. Base clearly narrower than apex. Apex slightly convex, apical angles slightly less than

rectangular, but faintly produced, obtusely angular. Apex finely margined. Sides evenly convex,

posteriorly obliquely convex, strongly margined, lateral channel absent. Basal angles evenly rounded

off, base in middle markedly concave, distinctly margined. Whole surface regularly convex, without

median line. Apex near apical angles within the widened margin with 4-5 strong setae, lateral margin

below border with several moderately short setae. Microreticulation absent, surface with very

scattered, moderately coarse punctures each of which bears a short, erect to slightly declined hair.

Surface without additional puncturation, glossy.

Elytra (Fig. 288). Rather short, parallel, rather cylindrical. Apex wide, truncature gently convex,

lateral apical angles widely rounded off. Base transverse, shoulders rectangular though shortly

rounded off. Basal margin attaining half of distance to suture, base in middle with a deep transverse

impression. Basal border wide, marginal channel very narrow, not concealed. Basal border with two

rows of strong, elongate setae, also lateral margin slightly below border with a series of elongate setae.

Series of umbilical pores consisting of 5 pores behind shoulder and 4 widely spaced pores from in front

of middle to posterior angles, but none at apex. Pores large, markedly umbilicate, easily to see, setae

elongate. Scutellar pore present, large, umbilicate, seta elongate. Striae including sutural stria absent,

but position of striae marked by irregular rows of very coarse, widely spaced punctures that each bears

a short, slightly declined hair. Punctures much more coarse than those on pronotum. Microreticulation

and additional puncturation absent. Surface highly glossy. Elytra tighly fused together, wings

reduced.

Lower surface. Prosternal process rather short, moderately surpassing procoxae, surface de-

pressed, margined inside of coxae, apex straight, rather depressed, surface shortly pilose. Metepister-

num very short, rectangular, a half wider than long. Abdominal sterna with a pair of elongate setae.

Terminal sternum in the male with 1, in both females with 2-3 elongate setae at some distance from

apex. Lower surface glossy, sparsely punctate and setose.

Legs. Moderately elongate, Ist tarsomere of protarsus about as long as wide, tibial groove of

profemur moderately deep, symmetric, anterior border straight. Femora comparatively narrow and

elongate, tibiae elongate, not at all widened. Metatibia elongate, c. 7 x as long as wide, 1st tarsomere

of metatarsus slightly <2 x as long as wide. d protarsus unknown.

d genitalia (Figs 83d-i). Genital ring rather narrow and parallel, asymmetric, with rather elongate,

incurved apex, with very elongate, asymmetric, deeply excised base. Sternum VII rather narrow,

apically divided, with very deep excision, base very faintly bisinuate, basal angles acute, lateral parts

elongate, markedly triangular. Aedeagus very elongate, narrow, depressed, slightly narrowed in

middle, faintly asymmetric. Basal part long, markedly bent. Lower surface almost straight. Apex

narrow, obtusely acute. Orifice very elongate, internal sac rather simply folded. Right paramere rather

narrow, elongate, with rounded apex, left paramere considerably larger than right, short and wide.

? genitalia (Figs 83k,l). Apex of sternum VII rather narrow and elongate, markedly triangular,

laterally acute, basal process narrow and elongate. Both stylomeres narrow and elongate, though

shorter than in other species, stylomere 1 completely divided, stylomere 2 spine-shaped, at apex with

1 short seta. Lateral plate elongate, rectangular, with 2 short medial-apical setae.

Variation. There is considerable variation of size and colour, otherwise a homogenous species.

Vivipary. Not confirmed in the examined material.

Habits. Little known. Holotype undated, mounted on same card with ants of the genus Crematogaster

Lund, both paratypes collected “ex pitfall trap in heath” in November.

Distribution (Fig. 602). Southwestern corner of Western Australia.

Material examined (3). Only the type series.

Etymology. The name refers to the short body.

7.3. Genus Adelotopus Hope

Adelotopus Hope, 1834, p. 11; Erichson 1842, p. 95; Westwood 1853, p. 403; Lacordaire 1854, p. 153; Sloane 1920,

p. 177; Notman 1925, p. 6, 28; Csiki 1933, p. 1634; Darlington 1968, p. 240; Matthews 1980, p. 10; Moore 1983,

p- 78; Moore et al. 1987, p. 49.

Type species: Adelotopus gyrinoides Hope, 1834, by monotypy.

Diagnosis. Genus of Pseudomorphinae, delimited by following characters: Body fairly wide to very

narrow, even cylindrical; head prognathous but directed downward, rather deeply imbedded in

prothorax; eyes situated laterally, with sharp ventral border; clypeus partly fused to frons; labrum

markedly divided from clypeus, commonly rather overlapped by clypeus; supraorbital, preorbital,

suborbital, mental, and gular setae absent; antennal grooves deep and elongate, ventrally widely

overlapped by large, foliaceous plates; antenna short, antennomeres more or less moniliform; glossa

wide, tongue-like, with several elongate apical setae; at least labial palpi markedly securiform; ventral

surface of head short, partly concealed by the mouth parts; prosternal process of pronthorax

moderately elongate; number of umbilical pores of elytra reduced; femora and tibiae rather com-

pressed, femora with deep grooves; tarsi moderately elongate; male sternum VII not excised; sternum

VIII not divided, asymmetric; aedeagus with more or less complex internal sac; parameres fairly

similar, though left paramere considerably larger; female stylomeres 1 and 2 fused, foliaceous; without

distinct dorsal ensiform seta, ventral ensiform setae, or nematiform setae, but with variable number

of apical or subapical setae on medio-apical surface not situated in a pit.

7.3.1. Description of Adelotopus

Species of Adelotopus exhibit the following character states:

Size and shape. Small to moderately large species (c. 3.5-10.5 mm) of rather wide and depressed

to elongate, highly convex, and cylindrical form.

Colour and pattern (Figs 100-225). Surface either unicolourous black or piceous, or more or less

light reddish, or patterned in various ways: either with apex of elytra more or less widely and

distinctly reddish, or with base of elytra reddish, or with base of elytra narrowly black and rest red,

or with reddish sutural stripe or spot of variable shape, or with both, sutural spot and apex of elytra

Figs 84-87. Adelotopus paroensis Castelnau. 84. Surface of left elytron. 85. Ventral surface of head. 86. Labium

and Galeae. 87. Mouth parts, ventral view.

reddish, or bicolourous with head and pronotum black and elytra reddish, or with more or less

distinctly lighter lateral margins of pronotum and elytra. Ventral surface usually slightly lighter than

dorsal surface, especially abdomen commonly reddish or becoming lighter towards apex. Some

species with the commonly protruding terminal tergite reddish and contrasting to the dark elytra.

Mouth parts, antennae, and legs mostly reddish or reddish-piceous, commonly lighter than rest of

body.

Microsculpture (Figs 438-566). Dorsal surface with marked to very fine microreticulation, or

without any microreticulation, or microreticulation present on head or on head and pronotum only.

Surface commonly also with more or less fine and dense puncturation, this commonly different on

head, pronotum, and elytra. Elytral striae usually indistinct or absent, in few species distinct, though

usually marked only by rows of punctures and/or fine, longitudinal strioles. Elytral intervals slightly

convex in few species. Surface without or with fine pilosity, in few species with conspicuous, elongate,

erect hairs.

Head (Figs 84-91, 98, 99). Short and rather wide, deeply imbedded in the prothorax, somewhat

orthognathous, mandibles directed rather downwards. Surface slightly convex. Eyes large, ventrally

with sharp margin, laterally not protruding, though commonly head laterally slightly projecting, in

some species even gently angulate, then lateral margin of head posteriorly of eyes more or less

Figs 88-89. Adelotopus paroensis Castelnau. 88. Right maxillary and labial palpus, ventral view. 89. Apical part

of labial palpus, terminal view.

Figs 90-91. Adelotopus paroensis Castelnau. %. Right antenna, right anterior tarsus, ventral view. 91. Right

antenna, ventral view.

distinctly oblique and narrowed. Ventro-posteriorly of eyes with a row of short bristles, sometimes

also ventral part of orbits with a punctate and shortly setose area. Clypeus fused to frons, bisetose,

clypeal suture rather indistinct, commonly in middle widely interrupted, clypeus anteriorly slightly

surpassing margin of head, border straight. Labrum rather wide and short, divided from clypeus by

deeply impressed suture, in several species-groups even rather widely overlapped by the clypeus.

Anterior margin of labrum more or less deeply excised, labrum bisetose or quadrisetose. Mandibles

wide, depressed, without scrobe, regularly curved, usually mostly concealed by labrum. Mentum not

divided from submentum by a suture, with distinct, more or less elongate, acute, unidentate tooth.

Gula very short, without distincet gular sutures. Glossa very large, tongue-like, directed ventro-

posteriorly, margin with several (c. 6-16) elongate setae, dorsal surface usually shortly pilose.

Paraglosse not visible, perhaps completely fused to glossa. Lateral lobes of mentum moderately

elongate, wide, commonly apically rounded or obtuse. Labial palpus larger than maxillary palpus,

terminal article always distinctly securiform, sometimes extremely wide. Terminal article of maxillary

palpus rather narrow and parallel to moderately securiform. Terminal palpomeres of both palpi on

upper surface rather densely setose, on lower surface almost asetose, apes with glandulose groove, rim

with a very short row of minute hairs. Galea and lacinia small to very small, elongate, completely

covered by the glossa, not visible from below and so deeply hidden below the glossa that they

presumably do not play an important role in the process of absorption of food. Lacinia slightly curved,

with a fine fringe of setae along inner border. Antenna inserted below eye, in deep antennal groove

between ventral surface of eye and orbit and laminate lateral plate of maxilla. Lateral border of

antennal groove usually angulate or carinate. Antenna short to very short, more or less depressed and

widened in apical half, with preapical antennomeres from almost 3 x as wide as long to about square.

Lateral margin of antenna more or less densely hirsute, middle glabrous. apex of terminal antenno-

mere and latero-apical parts of 7th to 10th antennomeres glandulose. Apart from the clypeal and labral

setae no additional fixed setae on head, though whole lower surface in some species covered by more

or less elongate and dense pilosity.

Microreticulation of head distinct, or more or less superficial, or absent; when present, usually

consisting of isodiametric meshes. Puncturation usually present, though generally rather fine or very

fine, in some species even not visible within the distinct microreticulation. Medially of eyes usually

Figs 92-95. Adelotopus paroensis Castelnau. 92. d median leg, ventral view. 9. Adhesive setae of d anterior

tarsus. 94. Posterior leg of d, ventral view. 95. Apex of d abdomen, ventral view.

with shallow sulcus. In some species surface with more or less dense, though usually very fine

wrinkles or lines, surface withouth or with mostly very short and sparse pilosity, rarely with elongate

erect hairs.

Prothorax (Figs 100-225). Pronotum about as wide as elytra, apical margin more or less deeply

excised and apical angles protruding, though to a different degree. Apical angles acute or obtuse, or

almost rounded off, usually rather oblique. Lateral margins slightly to markedly convex. Basal angles

angulate, or obtuse, or more or less widely rounded off. Basal margin straight, or concave, or slightly

convex. Apex and base more or less distinctly bordered, lateral margins very finely bordered, in some

species only border rather coarse. Shape of pronotum varied from very wide, rather depressed, with

explanate lateral margins, to narrow, elongate, highly convex, almost cylindrical with extremely

narrow lateral margins. Commonly lateral margins rather chanelled. Basal part of pronotum in some

species with shallow, transverse impression across surface. Microreticulation coarse, or feeble, or

absent, puncturation varied, in some species laterally coarser and denser than in middle, surface very

dull to highly glossy. Commonly surface with a faint network of irregular strioles, or microsculpture

rugose. Apical and basal angles without setae. Surface without or with short and sparse pilosity, in few

species with elongate erect hairs. Sternum with a rather short to moderately elongate sternal process

jo]

Qı

Fig. 96. Adelotopus paroensis Castelnau. Left wing. Scale: 1 mm.

of varied shape, commonly this process shortly setose. Sternum bordered along anterior margin of

procoxae.

Elytra (Figs 100-225). Free. More or less elongate and convex, or even cylindrical. Lateral margins

rather parallel, or gently convex, commonly faintly excised in basal third. Base laterally rounded or

oblique, shoulders obtuse or almost rounded off, apex rather wide, slightly concave to gently convex,

lateral apical angles rounded. Lateral margin more or less wide, chanelled, in some species extremely

narrow, not covered to almost completely covered when seen from above. Epipleura on ventral

surface, not visible from above. Basal border complete or reduced to different degree, or reaching only

to lateral third of base. Striae most commonly absent, in some species present and then indicated as

rows of punctures and/or longitudinal strioles, striae generally barely impressed, hence intervals

depressed. Surface microreticulate or not, most commonly rather glossy or very glossy. Puncturation

variable, fine to very coarse, more or less dense, rarely almost invisible within the distinct microretic-

ulation. Scutellar seta generally absent, present only in very few species. Umbilical pores few in

number, rarely more than 7, of which 6 are situated behind shoulders, sometimes with an additional

seta at or shortly behind middle. In several species number of setae even more reduced. Base in front

of shoulders with a row of elongate hairs, sometimes also with some hairs behind shoulders, in some

species anterior half or two thirds of lateral margin covered by rather elongate hairs. Surface without

or with short and sparse pilosity, in few species with elongate, erect hairs.

Wings (Fig. 96). Fully developed.

Ventral surface (Fig. 95). Anterior coxae biperforate and closed. Median coxal cavities conjunct.

Metepisternum more or less elongate, c. 1.5-2.3 x as long as wide at anterior border, in some species

posteriorly obliquely bent and hollowed for reception of metafemur and metatibia. Abdominal sterna

with one or two setae on either side of posterior margin, rarely with a row of setae. Sternum VII

sometimes with a row of longer bristles along apical margin, generally without tactile setae. Lower

surface more or less densely pilose.

Legs (Figs 92-94). Moderately elongate. Femora large, wide, depressed, with deep furrow on

ventral surface to receive most of tibiae. Furrows asymmetric, in profemur on posterior side, in

mesofemur and metafemur on anterior side with large plate that widely overlaps the furrow. Tibiae

moderately elongate, somewhat depressed, tarsi rather short, slightly depressed dorsoventrally. Ist

tarsomere of protarsus not or but feebly longer than wide, Ist tarsomere of metatarsus c. 1.5 to almost

3 x as long as wide. Ventral surface of tarsi asetose apart from fixed setae and of a single pair of setae

on ventral surface of 5th tarsomere. Dorsal surface of tarsi asetose. 1st-3rd tarsomeres of male

protarsus with two rows of adhesive pads, mesotarsus without such pads. Tarsal claws smooth.

Male genitalia (100-225). Sternum VIII not divided, asymmetric, with asymmetric excision of

different form at apex. Genital ring variously shaped, though in most species more or less triangular.

Basal plate usually anteriorly more or less excised. Aedeagus conchiferous, of very different size and

shape, form in some species highly characteristic, within several species-groups, however, basically

similar. Internal sac more or less folded in complex pattern, in some species with characteristic oblique

Fig. 97. Female external genitalia of Pseudomorphinae, dorsal view. Above, from left: Sphallomorpha maculigera

(Macleay), Pseudomorpha (Notopseudomorpha) laevissima (Chaudoir), Pseudomorpha (Austropseudomorpha) insignis

(Sloane), Pseudomorpha (Pseudomorpha) behrensi Horn. — Below, from left: Adelotopus dytiscides Newman, Caino-

genion (s. str.) ipsoides (Westwood), Paussotropus cylindricus (Chaudoir), Cryptocephalomorpha papua Darlington.

Abbreviations: des: dorsal ensiform seta; Ip: lateral plate; ns: nematiform seta(e); ves: ventral ensiform seta(e);

stl: stylomere 1; st2: stylomere 2; VII: sternum VIII.

Figs 98-99. Adelotopus dytiscides Newman. 98. Frontal view of head. 99. Ventral view of head.

fold in apical part. Orifice commonly elongate, situated in middle. Parameres moderately to rather

dissimilar, left paramere considerably larger than right.

Female genitalia (Figs 97, 100-225). Stylomeres 1 and 2 fused to a more or less wide, commonly

triangular plate of varied shape. Ventral ensiform setae, dorsal ensiform setae, and nematiform setae

not distinguished, but a varied number of apical or preapical setae present that do not originate in a

pit. Lateral plate very large, with varied number of elongate apical setae on median apical angle.

Larviparous.

7.3.2. Key to the species-groups of the genus Adelotopus Hope

Because the number of species is now very large and the morphological diversity is high, though

erection of new genera or even subgenera is not advisable at the present stage of knowledge, I decided

to arrange the species into species-groups that confidently are concerned monophyletic units. For

heuristic purposes, the criteria for recognizing of these groups have been strict. As a result, several

groups consist of single species only that do not fit properly into other though very closely related

species-groups. For the benefit of the reader and for use as prerequisite of the later discussion of

phylogenetic relationships species-groups are keyed out in a special key.

1 Fabrum.gtiadrısetoser nun ee ee 2.

—Al.abrumubisetose 122.2... 10.0.2 nenn Re teren ee ER 10.

28 Seutellar,pore and'seta present....uue.atenesnctenseerennen guest ann net nen eek ee >

= Seutellar.pore and'setarabsent.........z2000220eatseneeeneeterererernnenanentaenruentne teen ee ee EB 5.

37 Surtacescoverediwithrelongategereet;haitsc..n.are. see A nee analis-group

— Surface without elonsate, erect halrs.....0 0ER N nenne 4.

4. Lateralimarems/ofielytraswithelongate'setae...n.ine.oseestesreueeesnsannese rue ersnsaseeserenr regen celeripes-group

- Lateral margins of elytra without elongate setae ...................22u20c0200c0200ceneneneen gyrinoides-group (part)

5. Rather wide, depressed species with rather explanate lateral margin of pronotum. Elytra unicol-

ourous, or with reddish apex, or with reddish transverse band. Glossa c. 16-setose. Abdominal

sterna with 1 pair of ambulatory setae. Surface not distinctly pilose. Aedeagus without striking

asymmetrie flange,or spinose.band at apex ..........uucnesoensnssntsasseessnsarsesergerereeree gyrinoides-group (part)

— Less wide and depressed species with narrow lateral margin of pronotum. When elytra unicolour-

ous, glossa only 8-10-setose and surface shortly though distinctly pilose and aedeagus with

striking asymmetric flange at apex. When elytra with reddish apex, either body very narrow and

elongate, cylindrical, or aedeagus with conspicuous spinose band at apex of internal sac. When

elytra with reddish transverse band, body rather narrow and convex and abdominal sterna with

2.pairsiof,ambulatory Setae:..u..2.:02..00.un... 0 ee 6.

6. Colour unicolourous. Surface distinctly, though shortly pilose ...........nee paroensis-group

- Elytra either with reddish apex, or with reddish transverse band or sutural spot... T

7° Elytra with’reddishiapex =.........2..222221120202u00n20000020 2222 aee anna anne see ee RE 8.

— "Elytra withrreddishitransverse band or Sutural'spot. ueuenc-neenrneneneunn see 92

8. Body very narrow and elongate, cylindrical. Aedeagus without oblique, spinose band at apex of

INteRalsacn. inne beenden ehren sen ee ae nemosomoides-group

- Body less narrow and elongate and dorsally somewhat depressed. Aedeagus with oblique, spinose

Band: at apex of Internal’ sae..........u.zsuuneressunsanennenaussennssetende sense punctulifer-group

9. Elytra with reddish transverse band. Body less narrow and elongate, more convex. Glossa c.

MO-SELOSC hie: Berne ee ee ia seele ade fasciatus-group

- Elytra with reddish sutural spot. Body narrower and longer, dorsally somewhat depressed. Glossa

ERO=SCLOS EHE ee nee hen ee maculipennis-group

10. Surface. covered. with conspicuous elongate,;erect-hairs .......eeu. en cesausnenroneuesssarennen: villosus-group

— Surface not covered with conspicuous elongate, erect hairs, at most very shortly pilose .......... ll.

13:

14.

1a)

16.

20.

Pr.

. Basal % of abdomen and of elytra except for scutellum red, apical % contrastingly black

Rorepodyzblack#elytrateompletelyredi nn en enden atrorufus-group

Colour different, when bicolorous, elytra at least partly black

esse El Sr he Boa fe In ee 2 ER NIE nigricauda-group

Colour different, when elytra basally red, then at least part of base black ...............nccce. 13

Base of pronotum and basal third of elytra with transverse sulci ..................... marginicollis-group

Base 04 Ppronotum andeelytra without transverse sulei nn en eeneeneneeenseneeeeeteensnen scan 14.

Lateral margin of pronotum distinctly explanate, apical angles elongate, sharply protruding, basal

angles rectangular or obtuse, in a single species shortly rounded, in latter case colour dull black

mom distinet microretieulation’and short and widespeeies......n.n.n...annc sense 15;

Lateral margin of pronotum not distinctly explanate, but sometimes rather broadly channeled,

apical angles less sharp und protruding, basal angles varied, either glossy black, or yellowish-

jeddish, ot patierned, or.less- short and wide species... era een ntekencunetene 17.

Basal border of elytra complete. Abdominal sterna with at least 1 pair of ambulatory setae ........

EN ARTS OO ER ERRRIRSE URS dytiscides-group

Series of umbilical pores of elytra with postmedian pore. Basal angle of pronotum shortly rounded

OL sp N NEE ER, katherinei-group

Series of umbilical pores of elytra without postmedian pore. Basal angle of pronotum angulate or

OS nee een duntesien et brevipennis-group

Skateralimarein.oßelytra with a row of elongate setae ........2.202.0..2.22ucu000ue0eannsatnennsdsneskae: laevis-group

Baterallmareın ot elytra without a row.of elongatessetäae u... 18.

SBasal borderiof elytratalmostieompleteiorllittle äbbreviated" 2.2... en 19:

Basal border of elytra abbreviated, attaining at most two thirds of base, usually less ............... >

. Basal border of elytra almost complete, reaching very close to suture. Lateral margin of pronotum

narrow. Abdominal sterna with 1 pair of ambulatory setae. Sternum VI without fringe of longer

SEILER ee niert politus-group

Basal border of elytra more abbreviated, at most attaining median % of base. Either wide,

depressed species with rather wide lateral margin of pronotum, with 1 pair of abdominal

ambulatory setae, but with sternum VI with fringe of longer setae, or narrow, almost cylindrical

species with very narrow lateral margin of pronotum, with several pairs of abdominal ambulatory

setae, Puwithout-tringe:of longer 'setae on stemmum VL. .......2urataneusasusenagensnssnsgtacens nase ennnescennanene st 20.

Wide, depressed species with rather wide lateral margin of pronotum. Abdominal sterna with 1

pair of ambulatory setae. Sternum VI with fringe of longer setae at apex ...... exactor-group

Narrow, almost cylindrical species with very narrow lateral margin of pronotum. Abdominal

sterna with several pairs of ambulatory setae. Sternum VI without fringe of longer setae at apex

NET EN BR RN Fe ENTE HERE PER SU HAUEN ER ÜRRREEE TER ERELRE fasmani-group

Series of elytral umbilical pores consisting of 10-14 pores along the whole margin. Abdomen with

2 pP airs'ofambulatorysetaek en een N aller erheerheheseranssehe sent similis-group

Series of elytral umbilical pores consisting of 2-8 subhumeral pores and sometimes a single

postmedian pore only. In latter case abdomen with 1 pair of ambulatory setae only ................ 22:

Sbodyzformmarrowaelongate, almostieylindaicalk rn. Hr ee eeneseeoresiescs rennen ZI

Bodystormivuiderlesszelongate, notieylindrieal. re ne kennen eneenn nn eusen nahe tneeseedesetne 20:

. Completely dull black. Elytron with 4 subhumeral umbilical pores and abdominal sterna with 2-3

Patısgokambrlatoryzsetaene nn erseneanserdenseuetanensanadngetsnaannnnssncsnunasusneskehedesekrnssnnnsfenne Hannes unicolor-group

Elytra with reddish apex, or mostly reddish, or with reddish apical spot. Either abdominal sterna

with 1 pair of ambulatory setae only, or with many setae, in latter case elytron with 6 subhumeral

umbıleal’potes: 2.0... u ee ne eo nn 24.

24. Each elytron with 2 subhumeral pores only. Elytra either with indistinct reddish apex or mostly red

apart mommarrowiblackibase.. un een ne linearis-group

Each elytron with 6 subhumeral pores. Elytra either with distinct reddish apex or with reddish

apical’spok ter... ee ner ee A REN IHRE 23.

. Elytra with reddish apex. Abdominal sterna with many pairs of ambulatory setae. Surface not

Markediyuslossyo. nn an. ee tasmani-group

Elytra with reddish apical spot that is in middle prolonged. Abdominal steran with 1 pair of

ambulatory setae only. Surface markedly'glossy .......neuneneeeenneenenn. ern seriepunctatus-group

. Colour reddish to piceous. Each elytron with 6 subhumeral marginal pores only. Abdominal sterna

with 1 pair of ambulatory setae only. Internal sac at apex with oblique fold .... rubiginosus-group

Colour black, without or with reddish apex. When dark piceous, either elytra with a postmedian

marginal pore, or abdomen with 2 pairs of ambulatory setae and internal sac at apex with an

oblique»conspieuously sSpINose/»bandi....2.0.0000000a LS een raganen neeeeenteere eee 27.

27. Surface not or very superficially microreticulate. Abdomen with 1 pair of ambulatory setae.

Labrum fairly large, less overlapped by the clypeus. Apex of aedeagus regularly rounded. Internal

sactatsapex;with an. oblique fOld ....us1s2s18ca80=4=2222202n20ensrsr0renseteseeneehenenigegtengeneeeeen multipunctatus-group

Surface distinctly microreticulate. Abdomen without ambulatory setae. Labrum very narrow,

deeply overlapped by the clypeus. Apex of aedeagus acute, side near apex sinuate. Internal sac at

apex without ansobliquie Loldin...meetsämenereentenes sack ee obsoletus-group

7.3.3. Key to the species of the genus Adelotopus Hope

Because of the extreme morphological similarity within certain species-groups, some species, especial-

ly females, may be very difficult to identify. Therefore it is advisable always to compare the figures,

and in many cases also to conduct very exact measurements to gain proportion ratios that have been

amply used in the key. For tracking the species in the following descriptive text, the reader should use

the checklist at the end of the paper.

Upper surface with conspicuously elongate, hirsute, erect pilosity.......u.s.....e002.000. 0000er 2:

Upper surface without conspicuously elongate, erect pilosity, at most with very short pilosity 3.

Eiykaunieolourousn> Old... een seen spaeter nenn ee sereleunueher meer villosus, spec. NOV.

Elyera with wide, reddish.apex ee, Old naar en ee analis Macleay

Elytra completely reddish, fore body contrastingly black. Rather large, wide species. sw. NSW

atrorufus, spec. nov.

Colouration different, when fore body black, elytra not completely reddish, though at least with

narrow, black basal band, or with wide, black apex .......:.....uu...0u.u222u0001022002n0000 00008000 ee 4.

Elytra maculate, or fasciate, or with distinct light apex, though in some species only a small dark

basal band left, or basal half red and apical half black (doubtful species under both couplets) 5.

Elytra unicolourous black, or piceous, or reddish, or light yellow ..............esessssesssssssessssensensnesnenen 56.

Basal half to two thirds of elytra including base red, apical part contrastingly black ................. 6.

Colouration different, at least base narrowly black (doubtful species under both couplets) ....... 8.

Apex of elytra distinctly red. Surface densely and coarsely though not rasp-like punctate, not

DIerotreTeulate- ne. NSW ...urnnunssnsnerusntnnnnsnnnsnn san una Me ee basirufus, spec. NOV.

10.

11928

19.

16.

17.

Apex of elytra black. Surface either rather sparsely though rasp-like punctate and with distinct

microreticulation, or almost impunctate and without microreticulation «ee: 72

Body form large, wide, depressed, with wide pronotum and explanate, reddish lateral margins

(Figs 33, 122). Elytra with distinct, rasp-like puncturation and dense microreticulation, dull. Whole

surface extremely shortly, but very densely pilose. n. NT ............e. rufozonatus, spec. NOV.

Body form rather small, narrow, elongate, with rather narrow pronotum bearing rather narrow,

not explanate, black lateral margins (Figs 48, 163). Elytra almost devoid of puncturation and

microreticulation, glossy. Whole surface impilose. Northern Cape York Peninsula, n. Qld ...........

re RE Ren N ERBE IRRRLTEIERN. nigricauda, spec. noV.

Elytra with lieht median sutural spot of varying size ...................uucuscssnensosososcsssrnenensnenscarsenensnensnscnen 2

Elytra with light discal spot, or transverse fascia, or with light apex ........ussesnennenn: 10.

Pronotum reddish, elytral spot small, trapezoidal, in centre of elytra (Figs 68, 414). e. Qld ..........

a EEE maculipennis Macleay

Pronotum black, elytral spot very large, wedge-shaped, occupies most of elytra (Fig. 415). nw. Vic,

SWAINSVWV ER SShallro@ VAR ee ee ee ee eeleerreee cuneatus, spec. NOV.

Elytra with discal spot or transverse fascia in basal half....................oneooereeseesosessneonensnsenenensensee: 11:

Elytra without discal spots or fascia, but with light apex .................sceoscnensenensensonensensüsensesensensanee 2Ü.

Each elytron with light discal spot in basal half that not touch at suture or combine to a common

fascia. Whole surface with remarkably coarse and dense puncturation ......ueeenennnn: 19%

Elytra with light fascia in basal half that is in some species rather hourglass-shaped. Surface with

lessicvanse/and.densepuncturation. ...........uu002u0n00Rnnenseunvecersnersneneheninenchenehssesrssnnsnenstenenzoscsntestenessngnesee 13;

Wide species with wide, laterally markedly explanate pronotum. Surface between punctures

without distinct microreticulation. ne. NSW, e. Old, ce. PNG ....................... rufoguttatus Blackburn

Narrow, elongate species with narrow, laterally barely explanate pronotum. Surface between

punctures with distinct microreticulation. Tas, Vic, ACT, NSW, e. Old ........ macilentus, spec. nov.

. Surface of pronotum and elytra densely and more or less coarsely punctate (at 40x) u. 14.

Surface of pronotum and elytra almost impunctate or extremely finely punctate (at 40x)... 16.

. Smaller, narrower species, length <3.9 mm, with narrower, pronotum and longer elytra, ratio w/l

of pronotum <1.65, ratio width pronotum/head <1.58, ratio 1/w of elytra >1.46. Pronotum more

convex, laterally barely explanate. Puncturation far less coarse. Vic, NSW, se. Old...

see es hehehsssnesesesnnntuessnsctsssnnnuuseenshenssnauuuessnenussssntenensssechansrsnnnaensnusceststnnasessnnsecessnnttes zonatus Castelnau

Larger, wider species, length >4.0 mm, with wider pronotum and elytra, ratio w/l of pronotum

>1.69, ratio width pronotum/head >1.66, ratio 1/w of elytra <1.43. Pronotum more depressed,

laterally fairly explanate. Puncturation much coarser ......unnenensenesenenenenenenesennnenennnneenn 15.

Slightly smaller species, length <4.4. mm. Base of elytra more extensively black. Puncturation of

surface remarkably coarse. NSW, se. Old ...........u.u0s00000nenenenennenonenenenenenenenenensnsnnaesenene affinis Castelnau

Slightly larger species, length >4.5 mm. Base of elytra very narrowly black or even completely red.

Puncturation of surface distinctly less coarse. ne. NSW... basirufus, spec. Nov.

Basal border line of elytra short, barely attaining middle of base. Narrow, elongate species with

highly convex, narrow pronotum, ratio w/l of pronotum <1.50, ratio 1/w of elytra >1.64. Pronotum

with fine though at 40x still perceptible puncturation. e. Vic, s. NSW, ACT ... fasciatus Castelnau

Basal border line of elytra elongate, almost attaining scutellum. Wider, less elongate species with

wider, more depressed pronotum, ratio w/l of pronotum >1.60, ratio 1/w of elytra <1.63.

Puncturation of pronotum at 40x not or barely perceptible ..........enenenn 17.

Smaller, length 4.15 mm. Pronotum narrower at base, ratio base/apex 1.38. Elytra shorter in

relation to pronotum, ratio length of elytra/length of pronotum 2.51. Right paramere remarkably

narrow and parallel (Fig. 146i). Australia, exact distribution unknown .......... clepsydra, spec. nov.

18.

19:

Larger, length >4.6 mm. Pronotum wider at base, ratio base/apex >1.43. Elytra longer in relation

to pronotum, ratio length of elytra/length of pronotum >2.56. Right paramere shorter and more

triangular (Figs 1431, 1441, AS ee ae ee 0 RU ENRERFEEHEERAER 18.

Body form wider, with wider, more triangular pronotum bearing more explanate lateral margins,

ratio w/l of pronotum >1.82, ratio base/apex of pronotum >1.62, with wider elytra, ratio l/Yw of

elytra <1.43. Elytra distinctly microreticulate, also pronotum distinctly microreticulate though very

indistinctly punctate. ne. Old..............eessoseseenensnsnensnsenenenensnsesnssnsnenensnsnsrsrnsnene languidus, spec. nov.

Body form narrower, with narrower, less triangular pronotum and less explanate lateral margins,

ratio w/l of pronotum <1.82, ratio base/apex of pronotum <1.61, with narrower elytra, ratio 1/w

of elytra >1.44. Elytra not microreticulate, pronotum very indistinctly microreticulate though

under 64x fairly distinetly punctate ........eeeesssesesessnsenensnsenensnssnenennenensnnsnensenensnsensnsnsenensonsnsnsensennnenn 19:

Body wider with wider pronotum and shorter elytra, ratio w/l of pronotum >1.74, ratio 1/w of

elytra <1.50. Elytral spot almost always distinctly hourglass-shaped (Fig. 330). Parameres shorter,

especially right paramere perceptibly shorter than left (Figs 143i). ne. Qld south to about Mackay

(intermediate population further south)... bimaculatus bimaculatus Macleay

Body narrower with narrower pronotum and longer elytra, ratio w/l of pronotum <1.70, ratio 1/ w

of elytra >1.55. Elytral spot posteriorly usually straight or even convex (Fig. 331). Parameres longer,

right paramere barely shorter than left (Fig. 144i). se. Qld north to about Gladstone (intermediate

population further north) ...........eesesesoseaenrerescscnenenenenensnsnsssenentnsenr bimaculatus angustior, subspec. nov.

20. Only base of elytra narrowly black, apical % red. Body narrow, very elongate, cylindrical. e. Old,

TINTEN A ee linearis Macleay

Base of elytra more extensively black, usually only apex more or less widely red... 21.

. More than apical half of elytra reddish and reddish area anteriorly remarkably convex and

prolonged along suture (Figs 50, 356, 357) and head and pronotum densely and coarsely punctate.

Apex of aedeagus remarkably acute, dentiform (Figs 168g, 169g). Lateral plate of ? stylomeres very

short (Figs 1631, 169) 2.2.2.2... SR ee enrnehananrtresasnsne robnenen ee 22:

Either only apex of elytra reddish or reddish area anteriorly straight or concave or head and

pronotum not coarsely punctate. Apex of aedeagus never dentiform. Lateral plate of ? stylomeres

usually. less short ...........22..8.cu020000M0uanenectenensnnentesnesnensesaksnsrtnstengensrannensnsansnanssurnasss anne an nad dene nee 23:

. Generally larger, length 4.9-6.0 mm. Reddish elytral spot larger, usually widely touching lateral

border of elytra (Fig. 356). Pronotum laterally straight and remarkably conical, at base less

incurved. Surface less deep black. SA, Vic, sw. NSW; Tas ............ puncticollis puncticollis Notman

Generally smaller, length 4.8-4.9 mm. Reddish elytral spot smaller, not touching lateral border of

elytra (Fig. 357). Pronotum laterally slightly convex, not markedly conical, at base rather incurved.

Surface remarkably deep black. sw. WA ......nen puncticollis angustemaculatus, subspec. nov.

. Surface of elytra highly glossy, without microreticulation and striae well marked by a row of

punctures and intervals at most with a single row of punctures in middle and pronotum at least

laterally very coarsely punctate .......ueeesseseseeesesnsererennnnesnsnnnenenonnensnnnannensnenssosnenssnsssnsnsnnssssnenssssesesnesten 24.

Either surface of elytra microreticulate or striae not well marked or intervals more extensively

punctate or pronotum at most finely punctate .......uuneeenneneneneneneneneneenenenenesnnesnsnsnsesnsnsenesnensenenenone 28.

24. Smaller species, length <5 mm. Puncturation of apical part of pronotum in middle almost as coarse

as laterally. Pronotum and elytra longer and narrower, pronotum laterally rather parallel, ratios

w/lof pronotum <1.20, base/apex of pronotum <1.32, width pronotum/head <1.35, 1/w of elytra

EINE EAN) N I ee montisatri, spec. NOV.

Usually larger species, length generally >5 mm. Puncturation of apical part of pronotum in middle

distinctly finer than laterally. Pronotum and elytra wider, pronotum more widened towards base,

commonly rather conical, ratios w/l of pronotum >1.33, base/apex of pronotum >1.39, width

pronotum/head >1.45, 1/w of elytra <1.70 ...uuneeeesesesesesnesenenenennenenenennenenenenonnsnnnsonasnnnesessenensnnnseneenn 25.

26.

27.

28.

29.

30.

. Elytral intervals impunctate and impilose, though striae punctate. Lateral margin of pronotum

fairly wide. Stylomere narrow and sides parallel, lateral plate rather short (Fig. 1661). ne. Qld ...

RER Rn le RS. BRERE nm er rin BR RE ee en. Een RT Tee en calvus, spec. nov.

Elytral intervals more or less finely punctate and pilose, striae punctate or not. Lateral margin of

pronotum narrower. Stylomere wider, sides either convex (Fig. 1641), or rather parallel, but when

parallel lateral plate remarkably elongate and not parallel (Fig. 165)... 26.

Body slightly longer, narrower, with longer elytra, ratio 1/w of elytra 1.69. Pronotum very convex

with narrow lateral margins. Stylomere rather narrow with parallel sides, at apex straight, lateral

Platesvewy.elongate(Fie.s165) Se Oldienzensesensenescenensaensenennesn sn etsanen atneeaunnns seat CONVEXUS, SPEc. NOV.

Body slightly shorter, wider, with shorter elytra, ratio 1/w of elytra <1.65. Pronotum less convex

with slightly wider lateral margins. Stylomere wide, at apex rounded off, lateral plate rather short

(Eee 1 IRRE EN ee ann. 27.

Slightly larger, length 6.0-6.7 mm. Elytral striae almost impunctate. Pronotum with sparser

puncturation in middle of apical half. Vic, s. NSW ............... seriepunctatus seriepunctatus Notman

Slightly smaller, length 4.95-6.1 mm. Elytral striae punctate. Pronotum with denser puncturation

inmiddleior apieal’haltice Old... eersntaennnannsnennssennne seriepunctatus striatus, subspec. nov.

Body very narrow and elongate, more or less cylindrical (Figs 47, 57, 67, 349, 385, 411, 412, 413)

EEE nern aummns insert a iEheie een ran ee ereeeeedenereeerenilrehetusrhtetrgipetie 29.

Bodyawıdensandıshorter, not.cylindrieal uns... ln ara 30

Surface of pronotum and elytra densely microreticulate, not perceptibly punctate (at 40x), apart

from elytral striae that are sometimes finely punctate; surface remarkably dull ......................... 30.

Surface of pronotum and elytra not densely microreticulate, densely though very finely punctate

(aA) Ssurfaeestather SlOSSyas:....2:.2420226:0220000 0006 Dres ses eee tagen Li ggnante rasante se rnigeeusgerugeree nee De

Body slightly larger (length 5.8 mm), wider and shorter, with wider pronotum and not conspicu-

ously cylindrical body, ratios w/l of pronotum 1.30, width pronotum/head 1.36, 1/w of elytra 1.83.

Reddish apical part of elytra rather distinctly delimited, barely prolonged along suture (Fig. 349).

Basal angle of pronotum rounded off, lateral margin narrow, not channeled. Tas ............o-

ee ER RREER TEL RREHELHER OERERRL EREFERLNEREEE tasmanı Blackburn

Body slightly smaller (length <5.2 mm), narrower and longer, with narrower pronotum and

conspicuously cylindrical body, ratios w/l of pronotum <1.17, width pronotum/head <1.26, 1/w

of elytra >2.04. Reddish apical part of elytra ill delimited, prolonged along suture (Fig. 385). Basal

angle of pronotum rectangular, lateral margin rather wide, distinctly channeled. NSW, n. NT (one

oßithese’reeords may be WIONg]).......u..2.00000&20208200:082000608r8020H00000nanantansnsnnenransasshenreneeee bacıllus, spec. nov.

. Body narrower, more cylindrical, with longer and narrower pronotum, ratios w/l of pronotum

<1.10, base/apex of pronotum <1.18. Puncturation finer, just visible at 40x, microreticulation more

developed. Apex of aedeagus conspicuously hooked (Fig. 222g). e. Vic, s. NSW .......enene:

EHRE NEBEN. SEN ART SR Alina ein EL 3 van a a a ee fianesuvhtene longiformis, spec. nov.

Body slightly wider, less cylindrical, with shorter and wider pronotum, ratios w/l of pronotum

>1.23, base/apex of pronotum >1.24. Puncturation coarser, well visible at 40x, microreticulation

very faint. Apex of aedeagus not hooked (Fig. 221g) or unknown... 2

. Body slightly shorter, with wider pronotum and shorter elytra, ratios w/l of pronotum >1.25, 1/w

of elytra <1.91. Only apical third of elytra reddish (Fig. 411). Pronotum slightly less conical. e. SA,

VIERASTASTN SVEN ee ee ehnenterehter nemosomoides Westwood

Body slightly longer, with narrower pronotum and longer elytra, ratios w/l of pronotum 1.23, 1/w

of elytra 1.95. Almost apical half of elytra reddish (Fig. 413). Pronotum slightly more conical.

PistubutionNunknoWN een nnsnneweuenssansanaesasansnarnannassnnnennennehnsnnehengentenaesnnnnnenn conicollis, spec. nov.

SEIYtronsatubase with,seutellarpore and. setan....ar..r ses nn ae ern

Elytron’atbase. without seutellar.pore and Setat....usasessanssasuensnstsnsnsnsenesseoneneneneneesesessheeneetereennerererernee DO:

103)

40.

41.

. Generally smaller, length 4.3-5.2 mm. Reddish apex of elytra usually not well delimited or even

absent. Pronotum narrower, with wider and more chanelled lateral margins and less rounded

basal angles. Puncturation on elytra dense, at least traces of microreticulation present on elytra.

laswer Vie ACHESENSWERSAOIGE er ee dubius dubius, subspec. nov.

Generally larger, length 4.9-6.0 mm. Reddish apex of elytra well delimited. Pronotum wider, with

narrower and less chanelled lateral margins and more rounded basal angles. Puncturation on

elytra less dense, mieroretieulation absent onielytra....................unoenescsusesnscscnenan eenenereee 35:

. Pronotum generally slightly narrower, ratio w/l of pronotum 1.59-1.75. Puncturation on elytra

denser, surfacesless’@lossy?sW. Tas rsesseosessessensersosteesser een dene dubius hobartensis, subspec. nov.

Pronotum generally slightly wider, ratio w/l of pronotum 1.68-1.86. Puncturation on elytra

Sparsers sümfacesslossiersen Vieh. nn eeneeesneeene een en dubius glaber, subspec. nov.

36. Basal border line of elytra reaching to or close to scutellum (Fig. 292, 323-328)... 37.

Basal border line of elytra abbreviated, usually reaching only halfways to scutellum, rarely slightly

longer (doubtful’speeies under both couplets) ...euesreesessneneneneesesenenenenersrersnseanescenensesesese nern re 43.

. Basal border line of elytra clearly attaining suture behind scutellum. Large and wide, glossy species

with wide pronotum with distinctly concave base, and with apically markedly narrowed elytra.

Elytral striae near apex with rasp-like punctures. ne. Qld, PNG .......ee: apicalis Macleay

Basal border line of elytra not attaining suture behind scutellum. Narrower species with narrower

pronotum with straight base, and with rather parallel elytra. Elytral striae near apex without rasp-

Iike,pUnetmes. 22.2 aseserenatvenstresensengaonsnsgsnesasscatnanener aurneensarhehekah ersehen Besen Free 38.

. Elytra without microreticulation, glossy, striae at least in apical half well perceptible and distinctly

Punetdtesse Old. sansnensetnn seen ne ee et nitens, spec. NOV.

Elytra with very fine, though distinct microreticulation (at 40x), rather dull, striae almost not

perceptible and not or barely punctate .........s...2can2s2o2u02:000s00u2x2002z0n2nen<necanasasarasusäncnsrntnensnsessssernsnmeanaee 39:

. Body short and wide, with wide pronotum, ratios w/l of pronotum 1.84, 1/w of elytra <1.4.

Aedeagus short and wide, rather symmetric (Fig. 136g). Parameres, especially right one, conspic-

uously triagonal with angulate apex (Figs 136i,k). c. Old... rufocaudatus, spec. NOV.

Body longer and less wide, with narrower pronotum, ratios w/l of pronotum <1.75, l/w of elytra

>1.55. Aedeagus, when known, longer and narrower, slightly asymmetric (Figs 137g, 138g, 140g).

Parameres less triagonal with less angulate apex (Figs 137i,k, 138i,k, 1401,k) .......ueseenene: 40.

Reddish apical part of elytra anteriorly straight. Stylomere narrow, apical part parallel (Fig. 327).

NESOld nn ee nenne erregen eerherses sparsepunctatus, spec. NOV.

Reddish apical part of elytra anteriorly concave. Stylomere wide, apical part triangular or convex

(Figs 324, 325, 328). Distribution different ...........ssscnnssesenesessunsnenssssenstagenennngntenenensutnnanenndeshesee er ne 41.

Smaller, length 5.5 mm. Elytra shorter, ratio 1/w 1.55. Pronotum wider in relation to head, ratio

1.73. Stylomere wide, at apex obliquely convex (Fig. 141l). Aedeagus unknown. Distribution

ÜINKENO WEN En en Baade sosen Bone andann een Sees ssssge teens nnd een eg semilunatus, spec. NOV.

Larger, length >6.0 mm. Elytra longer, ratio 1/w >1.60. Stylomere see figs 1371, 1381 ............... 42.

. Larger, length 6.4-8.1 mm. Pronotum slightly wider with wider base, ratios w/l of pronotum 1.62-

1.68, base/apex 1.55-1.62. Aedeagus slightly shorter with shorter parameres (Figs 137g-k). Sty-

lomere triangular (Fig. 1371). Tas, e. Vic, ACT, e. NSW, se. Old .............. haemorrhoidalis Erichson

Smaller, length 6.0-6.3mm. Pronotum slightly narrower with narrower base, ratios w/l of

pronotum 1.56-1.64, base/apex 1.51-1.56. Aedeagus slightly longer with longer parameres (Figs

138g-k). Stylomere convex at apex, not triangular (Fig. 1381). Tas, e. Vic, se. NSW .......ene-

VE RL anne h Mbeneusuesnnehhäbnsuatäubhuehnhsnsesherhunderanennsdhnsnmennännen nach See ee minor, Spec. NOV.

WJllabrum.Z-setose.............scssaogsengesssinsaesengereneenn nenn ent hen ee RER 44.

Labrum 2-sefose..u.......0000esannene ne nee ee ee ee ER 46.

ee S. A

44.

45.

46.

47.

48.

49.

Pronotum and elytra wide, dorsally markedly depressed, ratio w/l of pronotum >1.90. Aedeagus

without oblique spinose band at apex of internal sac (Figs 2058,h, 2098,h) .........eee. 45.

Pronotum and elytra narrower, dorsally less depressed, ratio w/l of pronotum <1.65. Aedeagus

with oblique spinose band at apex of internal sac (Fig. 217g,h) e. NSW, se. Old...

ee oe RO CE B EN NRRR CRILEESERREEE EN AARON punctulifer, spec. nov.

Pronotum usually wider, ratio w/l 1.96-2.08, elytra shorter, ratio 1/w 1.37-1.45. Elytra with 5-7

marginal pores. Surface of elytra less punctate, glossier. ACT, se. NSW ........... lunatus, spec. nov.

Pronotum usually narrower, ratio w/l 1.92-1.98, elytra longer, ratio 1/w 1.47-1.51. Elytra with 4,

rarely unilaterally 5 marginal pores. Surface of elytra more densely punctate, less glossy. e. SA, Vic,

INCH +8 NOW EEE lawrencei, spec. nov.

aroeispeeiesnlenpth 0 9m. nee nun en en rn 47.

Smalllersspeeiesylenath 6A mal. nenne ee 48.

Body rather wide, head with markedly produced antero-lateral angles. Ratios w/l of pronotum

1.72, base/apex of pronotum 1.61, 1/w of elytra 1.52. Pronotum with traces of microreticulation

only. Aedeagus unknown. Solomen Islands eneeeeeeeesnenensnensnonehenesteneneranenesenrasnn penelopeae, spec. NOV.

Body rather narrow, head with barely produced antero-lateral angles. Ratios w/l of pronotum

<1.6, base/apex of pronotum <1.5, 1/w of elytra >1.65. Pronotum with distinct microreticulation.

Aedeagus very elongate, narrowed at apex (Fig. 161g). e. Vic, ACT, NSW; ?SA ....... een

ee RE En I EN EEE similis, spec. nov.

Smallerileneth <simm)/and'ratiow/l of pronotum 17er eensse he nteneaenenee ran an 49.

Larger, length >5.1 mm, when length <5.8 mm then pronotum wide, ratio w/l >1.76......... 52.

Pronotum comparatively wide, ratio w/1>1.68, elytra short, ratio 1/ w c. 1.5. Anterior margin of red

apexiolelytra,slishtly. COnCaVessusssesesennsesctessesassssersstagnnsesnensseenenseeneneezesznrsecnenanne ende ssurtehgnnenaraaanndersget 50.

Pronotum narrower, ratio w/1<1.52, elytra longer, ratio 1/w >1.57. Anterior margin of red apex of

elytra slightly convex. At 40x traces of microreticulation on elytra barely visible .............. I1.

. At 40x microreticulation on pronotum and elytra well visible. Pronotum markedly convex,

microreticulation regular. Aedeagus distinctly narrowed to apex, apex less widely rounded (Fig.

O7 )anesOlde re tee n.nannscaseressenenstriensnenenssaresstssnsssntnsatdsntestesnge Shnereunstnenensnratr gibbosus, spec. NOV.

At 40x microreticulation on pronotum and elytra barely visible. Pronotum far less convex,

microreticulation superficial. Aedeagus little narrowed to apex, apex very widely rounded (Fig.

1558): Northern tip of Cape York Peninsula, ne. Old... yorkensis, spec. NOV.

. Pronotum and elytra dorsally very convex. Elytra longer, ratio 1/w 1.64. ne Old .........ee

A ELTERN IRENENENE EL RE EIS HR RE EERRE DE re R- UERLET DEREN convexicollis, spec. nov.

Pronotum and elytra dorsally perceptibly depressed. Elytra shorter, ratio 1/w 1.57. ne. Old ........

FE NEN LE EU TRRERE ee rer ragahäaerpienbeteednnehebelter een ler euhnber kurandae, spec. nov.

. Smaller species, length <5.9 mm, with wider pronotum, ratio w/1 1.76-1.90. Aedeagus, especially

apex, shorter, apical fold of internal sac less markedly denticulate (Figs 153g,h, 154g,h), parameres

less triangular (Figs'153i,k, 1541,k). ne. Old, PNG ...unienasseesesinnensnrenrsrersesenssenensssinsnsnesrostsestäenene 53:

Larger species, length >5.9 mm, with narrower pronotum, ratio w/l 1.66-1.77. Aedeagus, when

known, longer, especially at apex, apical fold of internal sac markedly denticulate (Figs 150g,h),

parameres remarkably triangular (Figs 150i,k) ............u2222242000000n20nrensronsonseosensnnnenerenseneenssnnenssenstnseenenn O4

. Aedeagus c. 1.3 mm long. Traces of microreticulation on elytra visible, puncturation more distinct,

well visible (at 40x). Series of marginal pores of elytra without postmedian pore. PNG ..........

RE EN an iege ser ren UndsehhrtasentEingrehisenensehrsgre in debitor Darlington

Aedeagus c. 1 mm long (at same size of beetle!). Puncturation on elytra barely visible (at 40x),

microreticulation absent. Series of marginal pores of elytra with postmedian pore. ne. Old .........

I N ee sis sueseos Roslasassoesaseenne nee erndernngangenn nitidior, spec. noV.

54.

jo]

e0.

61.

Elytra parallel for basal two thirds, surface posteriorly more depressed. Anterior margin of red

apex of elytra either slightly convex or sinuate (Figs 338, 339). Aedeagus unknown. Australia ...

OT LESE ER CERCS acereneeouckkürnreonen re Bent rende 55.

Elytra distinctly narrowing from behind shoulders, surface posteriorly rather convex. Anterior

margin of red apex of elytra slightly concave (Fig. 337). Aedeagus see fig. 150g-k. Malaysia, Java

a ee a En ern jacobsoni Ritsema

. Microreticulation on pronotum and elytra fine though well visible (at 40x), surface moderately

glossy. Pronotum narrower, ratio w/l c. 1.65, basal angles obtuse. Anterior margin of red apex of

elytra slightly convex, regular. Lateral plate of stylomere elongate (Fig. 151). ce. Old „nn.

De a een geminus, Spec. NOV.

Microreticulation on pronotum and elytra highly superficial, barely visible (at 40x), surface glossy.

Pronotum wider, ratio w/l c. 1.7, basal angles rounded. Anterior margin of red apex of elytra

sinuate, incised at suture. Lateral plate of stylomere shorter (Fig. 1521). Northern tip of Cape York

Beninsula,men Older en ee en ne laticaudatus, spec. Nov.

. Body narrow and very elongate, eylindxieal. SAWAr....... nn. anna een unicolor, spec. noV.

Body wider and shorter, not cylindrical .......nesesesensensenneesensensensensensensensensentententeneneneetonte 57%

. Elytra at base with scutellar pore and seta .......unsenenesensensennensenennennenentnenenenttente 58.

Elytra without scutellar pore and seta ......unsnssnnnennensensensensensensenennensessensensensensensentententententonten 60.

. Body very wide and short, elytra almost as wide as long, ratios w/l of pronotum >2.3, 1/w ofelytra

<1.2. Basal angles of pronotum widely rounded, lateral margins of elytra with elongate setae

(Fig. 386). s. WA .n.canenneeseneenenensenenensenenensensnsnnensnannensnnntennensnsnsonsnsnsenensnstsensesenserentererrerntenensnsentne celeripes Lea

Body far less wide and more elongate, elytra far longer than wide, ratios w/l of pronotum <1.75,

l/w of elytra >1.45. Basal angles of pronotum more narrowly rounded, lateral margins of elytra

without elongate setae. Distribution different ........ussenensssesssenssenssennsennssunssennernensenssennernnsseneen 09,

. Pronotum generally wider, elytra shorter, ratios w/l of pronotum 1.63-1.74, 1/w of elytra 1.46-1.52.

Apex of elytra usually more or less distinctly reddish. Aedeagus near apex convex, apex slightly

wider (Fig. 203g). Tas, e. Vic, ACT, s. NSW; ?s. SIEHE dubius dubius, subspec. nov.

Pronotum generally narrower, elytra longer, ratios w/l of pronotum 1.56-1.65, 1/w of elytra 1.50-

1.55. Apex of elytra never reddish. Aedeagus near apex slightly impressed, apex slightly narrower

(Fig. 2028). ACT. s. NSW ........nesenessteseessnsssesennnnnensnsnsosennnenenensosnsnnsnnnsnsnsnstsenensnsnsnsenenen vicinus Castelnau

Basal border line of elytra reaching to or close to scutellum or suture „use: 61.

Basal border line of elytra markedly abbreviated, never occupying more than two thirds of base

ER ER RE RE RE Eee Sc erd FhBsrensorrrsnnsinncnncn 75.

Lateral margin of elytra setose also behind shoulders (Fig. 295). Aedeagus with asymmetrically

impressed apex (Fig. 1068), d sternum VII very elongate and with shallow apical excision (Fig.

VOSIESSAVV A er eeateeeanseeeen en ee e e e ee howdenorum, spec. Nov.

Lateral margin of elytra asetose behind shoulders. Aedeagus not with asymmetrically impressed

apex (Figs 100g-105g, 128g-132g, 134g), d sternum VII less elongate and usually with rather deep

apical excision (Figs 100e-105e, 128e-132e, 134e) ......nneneseneneensesenenenenseneneneneenennenenennennenenenetononnt 62.

. Lateral margins of pronotum explanate and basal angles angulate, almost rectangular (Figs 289-

294). Wide species with wide pronotum and aedeagus never with oblique fold in apical part of

internal sac (Figs 1008,h-105g,h)) ........u.eeneenesesnsnsenenenensnenenenennneeeeeneneneneneeeeeeensnnnennenttnns 63.

Lateral margins of pronotum not explanate though sometimes rather broadly channeled and basal

angles not rectangular and angulate. In doubtful cases either less wide species with narrower

pronotum or aedeagus with oblique fold in apical part of internal sac or basal border of elytra

abbreviated, attaining only median % of base ..........neessenesennenenenenenennnennnennnnsnsnsesnnenenenenenenenenenenenn 68.

. Surface markedly dull from dense microreticulation (Fig. 446), elytra almost impunctate, not even

striae distinctly punctate. Aedeagus narrow and elongate, symmetric, lower surface evenly curved

(Fig. 105g), genital ring markedly asymmetric (Fig. 105f). sw. WA... SETICEUS, SPEC. NOV.

64.

66.

67.

68.

69.

”D.

ale

Surface dull or glossy, but elytra always punctate, at least striae posteriorly distinctly punctate.

Aedeagus, when known, less narrow and elongate or slightly asymmetric or lower surface not

evenly curved (Figs 100g-103g), genital ring rather symmetric (Figs 1008-1039)... 64.

Generally smaller, length <6.5 mm, with narrower pronotum and longer elytra, ratios w/l of

pronotum <1.8, base/apex of pronotum <1.7, 1/w of elytra >1.40. d unknown. ne. Old .............

EN En. nebenan sartlenincsenkädee iunncsheste Ten atneenochernn zborowskii, spec. Nov.

Either larger, length >7.5 mm or with wider pronotum and shorter elytra, ratios w/l of pronotum

SO Nbaselapexot Promotum>1.8,1Y wiofelytras 1.39. en nn nenne ea te 69.

. Surface fairly glossy, microreticulation fine, not silky. Puncturation at apex of elytra not rasp-like

(Figs 443, 444). Apex of abdomen conspicuously red. Aedeagus short, slightly asymmetric, lower

surface distinetly: eonyex Eig. 103g). ne Old, PNGrrenneneeeeeeneneznenenentnenensetneneeee apicalis Macleay

Surface more or less dull, microreticulation stronger, rather silky. Puncturation at apex of elytra

rasp-like (Figs 438-442). Apex of abdomen not conspicuously red. Aedeagus longer, less asymmet-

nieWlowersuntace barely.convex (618510051028) 2 een seen eeen een enenenenseeneeseseer 66.

Base of pronotum straight (Fig. 289). Punctures at apex of elytra rather fine, not horseshoe-shaped,

though rasp-like. Aedeagus narrow (Fig. 1008). e. SA, Vic, ACT, NSW. s. Old. sw. WA; Tas .....

EEE EEE Nee een Ren ern ee dytiscides Newman

Base of pronotum distinctly concave (Figs 290, 291). Punctures at apex of elytra remarkably coarse,

somewhat horseshoe-shaped (Figs 440, 442). Aedeagus wider (Figs 101g, 102g) .........e 67.

Body wider, with shorter elytra, ratios w/l of pronotum 1.89-2.05, 1/ w of elytra <1.25. Surface less

silky, punctures on apical half of elytra even coarser. Lower surface of aedeagus straight, left

Paramere acute at apex (Figs 102g,k). ne. Old ..........2.222u2e22:222222202020222020 000000020002 0000_8200 latior, spec. nov.

Body narrower, with longer elytra, ratios w/l of pronotum 1.78-1.86, 1/w of elytra >1.35. Surface

more silky, punctures on apical half of elytra slightly finer. Lower surface of aedeagus slightly

convex, left paramere rounded at apex (Figs 101g,k). ne. Qld ........... nee ulrichi , spec. nov.

Surface of elytra glossy, virtually without microreticulation (at 64x). Rather narrow, elongate

species with elongate elytra, ratios w/l of pronotum <1.72, 1/w of elytra >1.6. In doubtful cases

larger species with length >7.5 mm, or smaller species with length <5.5 mm and then with rather

coarse elytral puncturation ..........eseesesessesessssnsennsonnnnorunsnennsnesnsnssnennsnssensesenssersnssssessssnnsssnnnnssssnssnsnseestenn 69.

Surface of elytra not glossy, at least with traces of microreticulation (at 64x). In doubtful cases

either larger and wider species, ratios w/l of pronotum >1.72,1/w of elytra <1.55, or medium-sized

species, length 6.0-6.8 mm, with fine elytral puncturation .......uesenesessesesenenenennnseesenenenenenennenenn 70.

Larger, length >7.5 mm, with wider pronotum, ratio w/1 >1.65. Aedeagus rather elongate, with

wider apex (Fig. 131g), parameres with rounded apex (Figs 131i,k). ne. NSW .......uenseeeneeenn

EBEN NEBEN SE BEL. 4 EEE nn au EEE el aterrimus, spec. NOV.

Smaller, length <5.5 mm, with narrower pronotum, ratio w/1>1.4. Aedeagus rather short, with

narrower apex (Fig. 135g), parameres triangular with rather acute apex (Figs 135i,k). ce. Old .....

nn enenkuubektadkenesg een nennen tegnehhlnn en oncan re igesnnepensanschtänserhe caniae, spec. NOV.

Body narrow and elongate, ratios w/l of pronotum <1.6, 1/w of elytra 1.68-1.70. Aedeagus

N N substriatus, spec. NOV.

Body wider and shorter, ratios w/l of pronotum >1.7, /w of elytra <1.53 een 71.

Body wide, depressed, with distinctly reddish and rather explanate lateral margins of pronotum

and elytra and pronotum >2x as wide as long, markedly narrowed towards apex, ratio base/apex

1.87. Antero-lateral angles of head distinctly angulate. Aedeagus narrow and elongate, rather

parallel, slightly asymmetric (Fig. 1288,h). PNG ...........enesneeesenenseseneesener exactor Darlington

Body less wide and depressed, usually without distinctly reddish and rather explanate lateral

margins of pronotum or pronotum <2x as wide as long, less narrowed towards apex, ratio base/

apex <1.8. Antero-lateral angles of head not angulate. Aedeagus, when known, wider and shorter,

narrowed towards apex, rather asymmetric (Figs 129g,h, 132g,h) ........... een 728

72:

74.

He).

so.

Body narrower, generally with longer elytra, ratios w/l of pronotum 1.72-1.85, base/apex <1.70,

l/w of elytra 1.39-1.53. When pronotum rather wide, then elytral striae at apex not distinctly rasp-

like punctate(at AOx)Eeu nennen en ne en ken een bern hne ob eraeae san dnaree ne osnene bee ee 73.

Body wider, generally with shorter elytra, ratios w/l of pronotum >1.86, base/apex >1.76, 1/w of

elytra 1.38. When pronotum rather narrow, then elytral striae at apex distinctly rasp-like punctate

(at AO essen ernsten nern ee keesen one rennen resorts ER 74.

. Elytra generally slightly shorter, ratio 1/w 1.38-1.48. Parameres longer, more triangular, with more

acute apex (Figs 129i,k). e. SA, Vic, ACT, NSW, e. Oldy Tas ee politus Castelnau

Elytra generally slightly longer, ratio 1/w 1.47-1.53. Parameres shorter, less triangular, with rather

convexX apex (kies.l321,k).me. NSWise: Oldie een doyeni, spec. noV.

Lateral margins of pronotum and elytra distinctly reddish and somewhat explanate, basal angles

of pronotum rectangular. Striae near apex of elytra distinctly rasp-like punctate (at 40x). Northern

Up. ob@ape York Beninsula, n Older sennssee ne ne ae sedlaceki, spec. noV.

Lateral margins of pronotum and elytra not distinctly reddish and not explanate, basal angles of

pronotum rounded. Striae near apex of elytra not rasp-like punctate (at 40x). ce. NSW...

ee A OHR variolosus Lea.

SEabrum-setosens inne an Ener ernennen 76.

BADLUm2-SEtoseH Ar ee en ee Re EEE II HE RER .86.

. Body rather narrow, dorsally convex, ratios w/l of pronotum <1.4, base/apex of pronotum <1.35,

length elytra/pronotum <2.3. Surface rather sparse, though distinctly pilose. Aedeagus with

conspicous hook-like flange near apex (Fig. 219g). e. SA, Vic, NSW, Old; ?WA .......nnnesensesensnsenenenn

SORT N a a en ee A ee paroensis Castelnau

Body wide, dorsally depressed, ratios w/l of pronotum >1.6, base/apex of pronotum >1.5, length

elytra/pronotum >2.5. Surface impilose. Aedeagus without hook-like flange near apex .......... 77.

Body wide, colour deep black, remarkably glossy, with explanate lateral margins and widely

rounded basal angles of the wide pronotum, ratios w/l of pronotum 1.98-2.05, 1/w of elytra

1.35-1.44. Puncturation of surface remarkably sparse, though punctures fairly coarse, distance

between elytral punctures on the average 4-6x as large as diameter of punctures. ne. NSW, se. Old;

BVG see idee Merebennuss een sn nenne ehem s et agene Hanauer unansngp Ferner ee parumpunctatus, spec. NOV.

Body less wide, colour less deep black, generally less glossy, with narrower pronotum, less

explanate lateral margins and less widely rounded basal angles of ponotum, ratios w/l of

pronotum <1.93, 1/w of elytra >1.42. Puncturation of surface denser, punctures either coarse and

very dense, or much finer, distance between elytral punctures on the average <4x as large as

diameter'of punetures 22... ann. eos ge ereeeesrerhtee S SM Sn oe nu 78.

. Punctuation of surface remarkably coarse and dense, especially on pronotum, here distance

between punctures barely larger than diameter of punctures. ne. NSW, se. Old...

REN RE TEN punctatus Castelnau

Punctuation of surface very fine, usually less dense, especially on pronotum, distance between

punctures about 3-4x as large as diameter of punctures ..............uenesesessensnenenssnsnnnenssnsnsnenennsnsasnensnnne 79%

Body rather narrow and elongate, dorsally less depressed, ratios w/l of pronotum 1.62-1.66, base/

apex of pronotum 1.49-1.51, 1/w of elytra >1.6. Colour piceous with distinct reddish translucent

margins of pronotum and elytra. Aedeagus short (Fig. 210g,h), basal angles of d sternum VI

ObtuseN Fig, 2I0E) ENG nennen nennen et ee ee gippslandicus, spec. Nov.

Body generally wider and shorter, dorsally more depressed, ratios w/l of pronotum 1.65-1.92,

base/apex of pronotum >1.54, 1/w of elytra <1.53. Colour mostly black, rarely blackish-piceous

with reddish translucent margins of pronotum and elytra, in latter species pronotum much wider

and 'basal angles of ö sternum VII widely rounded ...........u.su0e2200e0esesesneenenesesnsneenennsenenerensenensarenne 80.

Surface of pronotum and elytra piceous-black or black, but in latter specimens lateral margins of

pronotum and elytra distinctly reddish translucent .........useeeneseseesesessesennenesnrnsnenennenenassenennssnsnsansenn 81.

un UT

81.

82.

84.

85.

86.

87.

88.

89.

Surface of pronotum and elytra deep black, lateral margins of pronotum and elytra not reddish

as en een 85.

At least traces of microreticulation visible on pronotum and elytra (doubtful specimens under both

OT) Le 82.

Mi erorehieulation absent onNPronotumlandvelytran essen eeeeeenessenenenszeneneecannesensnneneseeranensnsnnen 84.

Surface black, puncturation of elytra distinctly rasp-like (Fig. 540. Stylomere narrow and very

elongate1(19.2011) >Aedeagus;unknonmn: sw. WAR. een nee eneeneeensseneneeszennaneenseenne mainae, Spec. NOV.

Surface usually somewhat piceous, puncturation of elytra not distinctly rasp-like (Figs 538, 539).

Stylemeresshorten. andswider(Eigss1991, 200) Een nn nern open seeeenesnueeeerenereeseesetetaesnseecteeeneeenenen 83.

. Colour more distinctly piceous. Microreticulation rather distinct. Puncturation of elytra less dense,

distance between punctures c. 3x as large as diameter of punctures. Stylomere longer and narrower

(IES199)" er Vie se NOWen. een re. gyrinoides orientalis, subspec. nov.

Colour more blackish-piceous. Microreticulation rather indistinct or almost absent. Puncturation

of elytra denser, distance between punctures c. 2x as large as diameter of punctures. Stylomere

shorter andiwıder (Fig. 200278. WA nn nnesentenespenessseneeeeesaeneenenseanneansersneden gyrinoides gyrinoides Hope

Pronotum narrower, ratio w/l 1.65-1.76. Puncturation of elytra coarser and denser, distance

between punctures c. 2x as large as diameter of punctures. Apex of aedeagus rather acute (Fig.

199g), basal angles of d sternum VII angulate (Fig. 199e). s. WA .......... gyrinoides gyrinoides Hope

Pronotum wider, ratio w/11.76-1.84. Puncturation of elytra finer and less dense, distance between

punctures >3x as large as diameter of punctures. Apex of aedeagus rather wide and convex

(Fig. 207g), basal angles of d sternum VII widely rounded (Figs 207e). e. SA, w. Vic, s. NSW; se. Old

ee N RT murrayanus, Spec. NOV.

Body wider and shorter, with wider pronotum and shorter elytra, ratios w/l of pronotum 1.85-1.93,

base/apex of pronotum 1.60-1.65, 1/w of elytra 1.43-1.46. Apex of aedeagus rather transverse

(Fig. 204g). Stylomere very wide (Fig. 2041). ne. Vic, se. NSW, ACT............ montorum, spec. NOV.

Body narrower and longer, with narrower pronotum and longer elytra, ratios w/l of pronotum

1.75-1.86, base/apex of pronotum 1.55-1.60, 1/w of elytra 1.47-1.50. Apex of aedeagus more convex

(Fig. 206g). Stylomere narrower (Fig. 206). n. Tas, e. Vic, se. NSW .............. victoriensis, Spec. NOV.

Pronotum narrow with comparatively narrow base, rather quadrate, with widely channeled lateral

margins, near base with a transverse impression. Elytra with a transverse impression in basal third

ne N EA BRE Bal a Duke RE HEL KiR en 0 PNRCRRE RE NUR RORLE DEE SUR 87.

Pronotum either wide with rather wide base, or pronotum narrow, but in latter case lateral margins

not widely channeled. Pronotum and elytra without transverse IMpresSiOnS unse: 89.

Colour light reddish. Puncturation of pronotum very fine, of elytra fine, not coriaceous, microre-

ikerullaineyayeilhanogn elogieauninve 0) ee RR marginicollis, spec. NOV.

Colour black. Puncturation of pronotum coarse, of elytra coarse, either microreticulation very

distinct, or puncturation remarkably coriaceous. Distribution different... 88.

Surface of elytra markedly dull from dense microreticulation, puncturation rasp-like, moderately

coriaceous (Fig 465). Pronotum distinctly narrowed to base, elytra shorter, ratio 1/w <1.65.

Stylomere narrow and elongate (Fig. 126). c. NT...........eeessesseneeneenene coriaceus, Spec. NOV.

Surface of elytra rather glossy, microreticulation barely visible, puncturation highly coriaceous

(Fig. 466). Pronotum barely narrowed to base, elytra longer, ratio 1/w <1.76. Stylomere wide and

ShozE (iD 71) EVA anne erenenenstennusseense sten eageransosbesnsnensseönseee seminitidus, spec. NOV.

Lateral margins of pronotum explanate, apical angles acute and remarkably protruding, far

surpassing centre of apex (seen from above). Usually basal angles of pronotum rectangular or

obtuse, when narrowly rounded off, colour dark and series of marginal elytral pores with

postmedian pore and surface dull from distinct mieroreticulation een 0.

RD.

94.

Lateral margins of pronotum not explanate though sometimes rather widely channeled, apical

angles less protruding. Usually basal angles of pronotum widely rounded off. Colour either

reddish or black, in latter case surface more or less glossy from weak or absent microreticulation

EEE RL RE LLL EEE TE EEE 103.

Colour black or very dark piceous, generally without or with indistinct reddish translucent lateral

margins of pronotum and elytra or with very wide and distinct reddish lateral margins, but dark

base and apex, and pronotum at base very wide, ratio base/apex >1.8, with lateral margins near

base markedly parallel, puncturation rather dense and distinct, and microreticulation rather

Supentleial). 22... ne en eu ee er er A SEE Sal

Colour piceous to flavous, piceous specimens usually with reddish translucent lateral borders and

base and apex of pronotum and elytra, in that case base less wide, ratio base/apex <1.65, lateral

margins near base not markedly parallel, puncturation rather sparse and fine, and microreticula-

HONSVERVACHStImehe ee ee ERBE 96.

Lateral margins of pronotum and elytra widely and distinctly reddish. Base of pronotum very

wide, ratio base/apex 1.82, lateral borders near base markedly parallel. Puncturation of surface

distinct, rather dense, microreticulation rather superficial, hence surface comparatively glossy.

Stylomere very narrow and elongate (Fig. 110]). ne. NT... rufomarginatus, spec. NOV.

Lateral margins of pronotum and elytra at most narrowly and indistinctly reddish translucent.

Base of pronotum usually less wide, ratio base/apex <1.8, lateral borders near base not markedly

parallel. Puncturation of surface fine, far less distinct, microreticulation distinct, hence surface dull.

Stylomere.widerrandishorter ...2..2.0n000ernesssrensnnnsnsantateustantneutene turen nen eee sense near ee 92.

2. Body elongate, rather narrow, with narrow pronotum and elongate elytra, ratios w/l of pronotum

1.62, 1/w of elytra 1.59. Elytra with fairly sparse though coarse, distinct, rasp-like puncturation.

Aedeagus rather symmetric with wide, evenly rounded apex (Fig. 121g). Tip of Cape York

Beninsula, n5 Older nr ee nennen neurnasene nn nee a enaeneetee bamagae, spec. noV.

Body shorter, wider, ratios w/l of pronotum >1.71,1/w of elytra <1.47. Elytra with far less distinct,

not rasp-like puncturation. Aedeagus usually less symmetric (Figs 107g, 108g, 111g, 123g).

Distnibution different.....n....22.002202seasseseennnesnssnssnsnsnesncnnanennsenensann Kaum sauna ne eassanse se Fre nner es Sue 93.

. Basal angles of pronotum distinctly rounded off (Figs 31, 296). Basal border line of pronotum

covering two thirds of base. Group of marginal pores of elytra consisting of 6 subhumeral pores

and one postmedian pore. Aedeagus see figs 107g,h. n. NT... katherinei, spec. noV.

Basal angles of pronotum rectangular or at most obtuse (Figs 32, 297, 300, 310). Basal border line

of pronotum covering only half of base. Group of marginal pores of elytra lacking at least the

postmedian pore:-Aedeagus:see figs’ 108g,h, 111g;h, 123&, anna 94.

Puncturation of surface almost invisible (at 40x). Marginal pores of elytra consting of only 3

subhumeral pores. Aedeagus very wide, highly asymmetric, with acute apex (Fig. 108g). n. NT, n.

VEN A Een been bene rer ae ee N ee lener brevipennis Macleay

Puncturation of surface well visible (at 40x). Marginal pores of elytra consting of 6 subhumeral

pores. Aedeagus less wide, less asymmetric, with rounded apex (Figs 111g, 1238) ............. 95.

. Generally larger, length 6.1-7.3 mm, body more depressed, with wider pronotum and shorter

elytra, ratios w/lof pronotum >1.80, base/apex of pronotum >1.72,1/w of elytra <1.4. Basal angles

of pronotum angulate, surface densely and distinctly punctate (at 40x) and rather coriaceous.

Aedeagus see figs 111g,h. Stylomere rather narrow, apex medially convex (Fig. 1111). n. NT ......

BÜNDE an be Er a a De DE ET adelaidae, spec. nov.

Generally smaller, length 6.15 mm, body more convex, with narrower pronotum and longer elytra,

ratios w/l of pronotum 1.77, base/apex of pronotum 1.65, 1/w of elytra 1.47. Basal angles of

pronotum obtuse, surface extremely finely punctate, barely visible at 40x, not coriaceous. Aedea-

gus see figs 123g,h. Stylomere wide, triangular (Fig. 123]). n. NT. ............ee.. edithae, spec. nov.

96.

IM:

98.

100

101.

102.

Large, length 6.5-6.9 mm, body wide, markedly depressed, colour light reddish, with very wide

pronotum with wide base, ratios w/l of pronotum 1.84-1.88, base/apex >1.74. Apical angles of

pronotum remarkably elongate, basal angles almost rectangular, markedly projecting posteriorly,

base laterally distinctly concave (Fig. 113). Aedeagus see figs 112g,h. n. WA ....... ee

Generally smaller, length <6.55 mm, body narrower, less depressed, colour either yellow, or darker

reddish or reddish-piceous, with narrower pronotum and narrower base, ratio w/l of pronotum

usually <1.8, base/apex <1.63, only in a single small, yellow species with obtuse basal angles and

distinctly narrower base ratio w/l of pronotum >1.8. Apical angles of pronotum less elongate,

basal angles rectangular or obtuse, more or less projecting posteriorly, base laterally less distinctly

coneayellEio. 114,298, 302-307). Aedeagus seetiss 1095/1115 &,h.1198,h 2.2... 2... 97.

Small, length 5.5 mm, colour yellow, with wide pronotum though narrow base, because lateral

border much incurved towards base, ratios w/l of pronotum 1.83, base/apex 1.58. Basal angles of

pronotum obtuse (Fig. 114). Elytra short, ratio 1/w 1.3. Aedeagus see figs 115g,h. n. NT...............

oe ee ee N flavus, spec. nov.

Generally larger, length >5.6 mm, colour dark reddish to piceous, with less wide pronotum and

longer elytra, ratios w/l of pronotum <1.8, 1/w of elytra >1.38. Lateral margin of pronotum less

incurved towards base, base rectangular or obtuse, in latter case colour piceous with lighter

marsıns»Aedeagus seertiss'109g8,h, 1165,h-119S hans een eannecnsnsrnscanennantangensanen. 98.

Colour piceous, with rather distinct lighter margins of pronotum and elytra. Western Australia

Seo he EN ee SR NR ER LE Re 92%

Colour reddish, without lighter margins of pronotum and elytra. Eastern Australia ............ 100

Body longer, parallel, with narrower pronotum, ratios w/lof pronotum <1.72, 1/w of elytra >1.62.

Basal angles of pronotum rectangular, produced over base, base laterally sinuate. Aedeagus

longer, with longer apex (Fig. 109g). n. WA ..........uneeessenensnenensnsnnnnsäsnsesnsnssenenenens elongatulus Macleay

Body shorter, wider, with wider pronotum, ratios w/l of pronotum 1.8, 1/w of elytra 1.4. Basal

angles of pronotum obtuse, not produced over base, base straight. Aedeagus shorter, with shorter

APERNEISHIMOE).S. WA unentanneansaesssnsstnnnnnstnesensnenssensustegstsusarsssenssnsnnsnncrasnsnsnenesensätsesuene piceus, spec. Nov.

. Microreticulation of surface much reduced, highly superficial, therefore surface glossy. Basal

angles of pronotum barely prolonged over base, base not or far less concave. Elytra generally

longer, ratio 1/w 1.45-1.55. Apex of aedeagus widely rounded (Figs 117g, 118g) ................. 101.

Microreticulation of surface distinct, therefore surface dull. Basal angles of pronotum distinctly

prolonged over base, base distinctly concave or sinuate. Elytra generally shorter, ratio 1/w

1.38-1.46. Apex of aedeagus acute (Fig. 1198) ........eessesenenserenensnsnensnsnsnenensnsnenenenenssssnanssstsesesennnene 102.

Pronotum slightly narrower with narrower base, elytra generally slightly longer, ratios w/l of

pronotum 1.63-1.69, base/apex 1.48-1.53, 1/w of elytra 1.51-1.55. Base of pronotum almost

straight. Aedeagus slightly longer with less widely rounded apex (Fig. 117g). se. Old...

RL Menacahense esestareenkeacn sachsen denen ende pe eeeeeehennetekerrnrenee longus longus, spec. nov.

Pronotum slightly wider with wider base, elytra generally slightly shorter, ratios w/l of

pronotum 1.66-1.74, base/apex 1.58-1.62, 1/w of elytra 1.45-1.53. Base of pronotum slightly

concave. Aedeagus slightly shorter with widely rounded apex (Fig. 118g). ne. Old...

ee N RER longus tropicus, subspec. nov.

Pronotum slightly wider with wider base, elytra generally slightly shorter, ratios w/l of

pronotum 1.64-1.72, base/apex 1.53-1.60, 1/w of elytra 1.38-1.45. Microreticulation on pronotum

distinct, rather coarse, slightly coriaceous. Puncturation of pronotum and elytra coarser (Fig. 458).

Stylomere shorter and wider (Fig. 1191). e. SA, Vic, e. NSWESEA@Olde nee

BE nen ee un rs lunsertsnee araransessntens er Here sinuaticollis sinuaticollis, spec. nov.

Pronotum slightly narrower with narrower base, elytra slightly longer, ratios w/l of pronotum

1.62, base/apex 1.51, 1/w of elytra 1.46. Microreticulation on pronotum very fine, rather

superficial, not coriaceous. Puncturation of pronotum and elytra fine (Fig. 459). Stylomere longer

andlnarrower(Fle- 120). ce Oldi.....rseeeneesensersnsersesenneneneetannneee sinuaticollis calliope, subspec. nov.

106.

107

110.

IM:

Golounblackito,darkpieeous 2... re ee 104.

Colour reddish to flavous (doubtful species under both couplets) ..........eeeeenen: 108.

. Colour black, puncturation of elytra fine or very fine, barely visible at 40x. Aedeagus, when

known, notmarkedlystriate below............n.e2oenerseseizeranuesersnneeeneneneresnereeen nern ee 105.

Colour piceous, puncturation of elytra coarse, well visible at 40x. Aedeagus markedly striate

below (Fig. 179g). nw. Vic, WA .......u.uuessenseesencnneosnensnenroranensnsurneorsnsnrnenentonenrnesnsnnnen adustus, spec. Nov.

5. Surface with distinct microreticulation (at 64). Aedeagus sinuate near apex, apex acute (Fig. 159g).

Stylomere elongate, apex oblique, apical half parallel (Fig. 1591). ce. and ne. Old .......nee

N EN obsoletus, spec. noV.

Surface without microreticulation (at 64). Aedeagus, when known, not sinuate, apex widely

rounded (Figs 147g, 148g). Stylomere wider, apical half not parallel (Figs 1471-149) ............ 106.

Body wider and shorter, with very wide pronotum, ratios w/l of pronotum 2.03, base/apex of

pronotum 1.68, 1/w of elytra 1.33. Elytra narrowed from shoulders. Aedeagus see figs 148g,h. ne.

ee en ovatus, spec. NOV.

Body narrower and longer, with less wide pronotum, ratios w/l of pronotum <1.85, base/apex

of pronotum <1.57, 1/w of elytra >1.38. Elytra parallel in basal half. Aedeagus see figs 147g,h, or

unknown: Distunbutomditterent. ee 107.

Body wider and shorter, with shorter elytra, ratios base/apex of pronotum >1.50, 1/w of elytra

<1.44, length elytra/pronotum <2.57. Puncturation of elytra dense, fine, though well visible at

64x. Aedeagus see figs 147g,h. Stylomere without short seta(e) on lateral margin (Fig. 147). n. NT,

N A ee multipunctatus, spec. Nov.

Body narrower and longer, with longer elytra, ratios base/apex of pronotum 1.46, 1/w of elytra

1.55, length elytra/pronotum 2.69. Puncturation of elytra extremely fine, barely visible at 64x.

Aedeagus unknown. Stylomere with short seta(e) on lateral margin (149). n. NT...

RR DE ri Ale a a a an He ER ER LE IE ER SEE browni, spec. Nov.

. Lateral margin of elytra with a fringe of long setae at least in basal halt... 109.

Lateral margin of elytra without a fringe of long setae ........nnsnesenenenenensenenensnenensosensnensnsnsnennene 112.

. Aedeagus elongate, lower surface evenly concave, apex acute, highly asymmetric, markedly

turned laterally (Fig. 191g). Stylomere either parallel, with obliquely transverse apex, or at apex

bituberculate (Figs 191]). n. Old, n. NT, n. WA; Vic ..........eneesssssssosossenesssessenssnsenen laevis Macleay

Aedeagus shorter, lower surface almost straight, apex less asymmetric, barely turned laterally

(Figs 192g, 194g). Stylomere wider, either widened towards apex, or with convex apex (Figs 1921,

SAD ee een an aaa Pe tu rar 110.

Elytra shorter, ratio 1/w <1.54. Aedeagus less asymmetrically widened in middle (Fig. 194g), left

paramere with distinctly narrowed apex (Figs 194k). Stylomere rather wide, with convex apex

(Bis. 194) SC WMA on. 0. 22saninshenenenensnrnnusngsennaensenssrherenereesnsarnaesern range ten nassen stern dteeee brevior, spec. Nov.

Elytra longer, ratio 1/w >1.6. Aedeagus more asymmetrically widened in middle (Fig. 192g), left

paramere either with transverse apex, or with distinct hump at apex (Figs 192k, 193k). Stylomere

wide, widened towards apex, apex transverse (Fig. 1921). Old .............eseresesesessseneneneneneseenenen 1lalalg

Pronotum generally less wide with narrower base, puncturation rather coarse. Left paramere

straishtratiapex (Fis, 192K). ee: Old „u.a. 2... nn se ne est ciliatus ciliatus, spec. noV.

Pronotum wider with wider base, puncturation fine. Left paramere straight with distinct hump

at/bottomsob,.apex (Eis 19sk)ane Old. Are re ciliatus tenuipunctatus, subspec. nov.

Headrat!s4&x with.distinet traces of mieroretieulaub on"... een esse seen see see sense Se en IS:

Head at 64x without any traces of microreticulation .....ueeeseseeseseeserssnsesnennsnennenennsnssnsnenesonanssanne 129.

ON et een: Malta N ER 114.

ONCE AU RR A BER BR Re Se RP RU NEE A ER RARL TILL SSR EESL ER EL EEE 1 RE ER as ERBE RE 123%

116.

117.

118.

11119)

120.

DT.

. Aedeagus on ventral and lateral surfaces distinctly striate (Figs 1778-1808) .......................... 115:

Aedeagus on ventral and lateral surfaces not distinctly striate ..............nneneennnnn 118.

. Surface mostly piceous, rarely reddish. Pronotum and elytra remarkably densely and coarsely

punctate. Pronotum wide, with wide base, ratios w/lof pronotum 1.69-1.78, base/apex 1.52-1.58.

Aedeagus moderately asymmetric with acute apex, lower surface almost straight (Figs 179g,h),

parameres rather elongate, not markedly triangular (Figs 179i,k). nw. Vic., WA .......neen

ee NE a Te ae na He ae adustus, spec. nov.

Surface reddish. Pronotum and elytra less densely and coarsely punctate. Pronotum less wide,

with slightly narrower base, ratios w/l of pronotum <1.68, base/apex <1.48. Aedeagus variable

(Figs 177g,h, 178g,h, 180g,h), parameres less elongate, apex variable (Figs 177i,k, 178i,k, 180i,k)

TERROR EEE ERTETTITLELTTENFTER OT TEEN tere er 116.

Aedeagus very asymmetric, more convex towards apex, laterally slightly impressed (Fig. 177g).

Left paramere large, somewhat triangular (Fig. 177k). Puncturation on pronotum rather dense

and fairly coarse, on elytra fine and sparse. Elytra longer at the average, ratio 1/w 1.46-1.58. SA,

VI WEN SVEN ee virgatus

Aedeagus less asymmetric, less convex towards apex, laterally not impressed (Figs 178g, 1808).

Left paramere see figs 178k, 180k. Puncturation either dense and moderately coarse or sparse and

very fine on both, pronotum and elytra. Elytra shorter at the average, ratio 1/w <1.45 ....... 117.

Puncturation of surface sparse and remarkably fine (Fig. 517). Aedeagus slightly narrower,

faintly convex towards apex, lower surface convex (Figs 178g,h). Left paramere convex at apex

(ie) unknown. NE. en es desire brittoni, spec. nov.

Puncturation of surface very dense and fairly coarse (Fig. 519). Aedeagus slightly wider, faintly

faintly concave towards apex, lower surface straight (Figs 180g,h). Left paramere triangular, with

SeutFraApESALIE. ISORK)-2 UNKNOWNENT .... nn er nesnteseagenee, punctatissimus, spec. nov.

Wide, remarkably depressed, yellowish species with wide pronotum, ratio w/l >1.7, and with

widely channeled lateral margins (Fig. 363). Aedeagus symmetric, with shortly rounded apex

115.121) luteus, spec. nov.

Less wide and depressed, reddish species with narrower pronotum, ratio w/l <1.7, with

narrower, less channeled lateral margins. Aedeagus see figs 173g,h, 174g,h, 182g,h ............. 119.

Aedeagus rather elongate, with almost straight lateral border and gently rounded apex (Fig. 173g).

Left paramere rather narrow with acute apex (Fig. 173k). On the average rather large species with

wide pronotum, wide base of pronotum and short elytra, ratios w/l of pronotum 1.58-1.70, base/

apex.1.40-1.s1M l/-w ofelytra 1.47-1.31.2e7 SA, 2w.:Vie5w. NSW, w>Old,. cHNT..2..222.222K.22:,

gerne NEE EU RUHR GEL ENEE HEFTE EEE. SEOELTDER MIT BI HER LLEAR SARA TT TR VER laticollis, spec. nov.

Aedeagus shorter, generally with more acute apex (Figs 174g,h, 182g,h). Left paramere wider

with widely rounded or oblique apex (Figs 174k, 182k). On the average smaller species with less

wide pronotum, narrower base of pronotum and longer elytra, ratios w/l of pronotum 1.47-1.60,

base/apex 1.35 1.41, 1/wof elytra 1.4971.60. Distribution’ditferent ............m2ees00sncarsnsertennneshenn 120.

Apical part of aedeagus slightly concave (Fig. 182g), left paramere at apex convex (Fig. 182k),

ö sternum VII wider, base less convex (Fig. 182e). At the average pronotum narrower with

narrower base, ratios w/1 1.47-1.58, base/apex 1.35-1.39. Puncturation of pronotum usually less

rugose, puncturation of elytra usually sparser. ce. and ne. Old ........................ aequus, spec. NOV.

Apical part of aedeagus straight or slightly convex (Fig. 174g), left paramere at apex oblique

(Fig. 174k), S sternum VII narrower, base distinctly convex (Fig. 174e). At the average pronotum

wider with wider base, ratios w/l 1.52-1.60, base/apex 1.36-1.41. Puncturation of pronotum

usually rugose, puncturation of elytra usually denser. NSW, e. Old .......... cribricollis, spec. nov.

Surface usually piceous, rarely reddish, pronotum and elytra remarkably densely and coarsely

punctate (Fig. 518). Pronotum with wide base, ratio base/apex 1.52-1.58. Stylomere with rather

elongate, parallelapexX (EigI179]). nw. Vie, WA een eetzeeneesansneersnnnnneeserternenee adustus, spec. noV.

Surface reddish or yellowish, pronotum and elytra usally less densely and coarsely punctate.

Pronotum commonly with narrower base, then ratio base/apex <1.51. When base wide, punctu-

ration sparse and rather fine and stylomere polysetose (Fig. 189) ..........unsenseeneenen: 122:

Large, length 6.65 mm, with wide base of pronotum, ratio base/apex 1.6. Puncturation of elytra

rather sparse and fine (Fig. 528). Stylomere narrow, in middle barely sinuate, polysetose

(Fig. 189]). d unknown. n. WA ......eeeneseresnensoneneneososWbentoenensnsnnnsrsensnensnsenrsnnenensasnne crassus, Spec. NOV.

Smaller, length <6.1 mm, with narrower base of pronotum, ratio base/apex <1.52. Puncturation

of elytra variable, though commonly less fine on elytra. Stylomere commonly wider, in middle

more or less distinctly sinuate, rarely with more than 4 setae (Figs 1731-1771, 1821, 1901) ..... 123.

. Maxillary palpus rather narrow, apex of terminal palpomere as long as median border (Figs 174b,

182b). Pronotum rather narrow, with narrow base, ratios w/l 1.47-1.60, base/apex 1.35-1.41. e.

ee eoiciontsc: 124.

Maxillary palpus rather wide, apex of terminal palpomere distinctly wider than median border

(Figs 173b, 175b-177b, 190b). Pronotum rather wide, with wider base, ratios w/1 1.58-1.75, base/

apex 1.36-1.56 .u.usesesssenensesesesenenenensnenscornenenensnsneneonentososonnenenssnensnsnsnsensnenensnsnsnensnanenensnetstnsnenehsnnensnenenenenee 125.

. Atthe average pronotum narrower with narrower base, ratios w/11.47-1.58, base/apex 1.35-1.39.

Puncturation of pronotum usually less rugose, puncturation of elytra usually sparser. ce. and ne.

Ol ZAN en ee aequus, Spec. NOV.

At the average pronotum wider with wider base, ratios w/l 1.52-1.60, base/apex 1.36-1.41.

Puncturation of pronotum usually rugose, puncturation of elytra usually denser. NSW, e. Qld

se se Te RE R e V Hhan rrunt cribricollis, spec. nov.

. Pronotum very wide, with wide, distinctly explanate lateral margins, ratio w/l >1.72. Punctur-

ation of surface fine and rather sparse (Figs 514, 515). Stylomere wide, almost regularly

thomboidal, with convex apex (Figs 1751, 17 )..uuss2rreeranneneseontersestecsanncnunnenenencanensnsteae erden rear 126.

Pronotum narrower, with less explanate lateral margins, ratio w/l <1.7. Puncturation of surface

coarser and denser (Figs 512, 516, 529). Stylomere not rhomboidal (Figs 17231,.172, 190) 5P72

). Base of pronotum narrower, ratio base/apex 1.42. Antenna narrow and elongate, barely widened,

ratio w/l of median antennomeres c. 1.5 (Fig. 176d). Puncturation of surface generally slightly

coarser. Lateral plate of stylomere longer, basal margin not markedly concave (Fig. 1761).

EUNKNOWM. © WAL. nsasassesettanensss ehe hernennnnesnenrrtonteeTetunchennthensegnrenegee rende er houstoni, spec. Nov.

Base of pronotum wider, ratio base/apex >1.52. Antenna short and wide, distinctly widened,

ratio w/l of median antennomeres >2 (Fig. 175d). Puncturation of surface generally finer. Lateral

plate of stylomere shorter, basal margin markedly concave (Fig. 175). ne Qld .......eee-

. Palpi very wide, terminal palpomere of maxillary palpus c. 2x as long as wide (Fig. 190b).

Stylomere'see fig. 1901. S unknown. n. NT ..........zz2.202202020000800200n0nenenenensosannnnenr latipalpis, spec. nov.

Palpi less wide, terminal palpomere of maxillary palpus <1.5 x as long as wide (Figs 173b, 177b).

Stylomere see figs’1731, 1771 ..........22.r.0u08.u0000.0000n A212 de nee sterne sterne Er 128.

. Stylomere narrower, not as markedly narrowed towards apex (Fig. 1771). On the average base of

pronotum narrower, ratio 1.36-1.48. SA, Vic, w. NSW, c. NT. ..................2.0.: virgatus, Spec. NOV.

Stylomere wider, markedly narrowed towards apex (Fig. 173l). On the average base of pronotum

wider; ratio 1.40-1.51..e. SA, w.. Vie w. NSW, w. Old,.e NT... 222 laticollis, spec. nov.

ON a Ih ae oe a DL Pa 130.

ee I EIER EEE frfeninfesscec 137.

. Aedeagus irregularly shaped, bisinuate on both lateral margins, apex somewhat knob-like

(FIo218le)2e2 Ode. sn nesunseranssanseseree anne sense nen ne. queenslandicus, spec. NOV.

Aedeagus not irregularly shaped, not bisinuate on both lateral margins, apex shortly rounded or

acute (Figs 170g, 171g, 172g, 183g, 185g, 1868, 1888) .........202200000002res22n0e00esnsesnesessansasensesnsonsssennnenen 131:

1Sjle

1132.

136.

137.

138.

Aedeagus on lower and lateral surfaces distinctly striate, lower surface straight, apex acute

(Figs 183g,h). Left paramere convex at apex (Fig. 183k). Genital ring slightly asymmetric, lateral

arms not markedly convex (Fig. 183f). Pronotum generally wider, with wide base, ratios w/l of

Pronotumal 551.62, base/apex 147-153: ner Old. ee palumae, spec. nov.

Aedeagus on lower and lateral surfaces not distinctly striate, lower surface varied, apex

commonly convex (Figs 170g,h, 171g,h, 172g,h, 185g,h, 186g,h, 188g,h), though when apex rather

acute, then lower surface convex, then left paramere at apex transverse, arms of genital ring

markedly convex, and pronotum narrower with narrower base, ratios w/1<1.51, base/apex <1.42

Be EEE ELTE ERSTE ET erende snäaseahgsntescessnesngensegee esse en tereneste see serseneeteres ee tet 132.

Aedeagus wide, highly asymmetric, apex rather acute, lower surface slightly convex (Figs 172g,h).

INSVVEROTCH He ee ee en Be Te N ee a Re foliaceus, spec. nov.

Aedeagus usually narrower, less asymmetric, apex rounded, lower surface straight or concave

(Figs 170g,h, 171g,h, 185g,h, 186g,h, 188g,h). When aedeagus rather wide, then far less asymmet-

Heandrapewidely:eonverewrr m... anne eier Fin sheet ne aeg 133;

. Aedeagus wide, in middle markedly widened, lateral margins near apex straight or even slightly

concave, apex widely:convex (Fig, 171g). sw. WA... distinguendus, spec. nov.

Aedeagus unusally less wide, when wide, then lateral margins near apex evenly convex

(Eies1205, E91 EOS TSTH) er ee 134.

N\DeXoraedeasus’widely rounded’off (Fiss 185183 )y nn een 135.

Apex.of dedeagus shortly rounded off (Fiss 1708, 1369)... =... ru. een 136.

. Smaller, length <4.3 mm, body more convex, pronotum narrower with narrower base, ratios w/l

of pronotum <1.5, base/apex <1.41. Aedeagus shorter and wider (Fig. 185g), left paramere at apex

Onverlbie 185R): 5.98, 1. Vie,sWE Old Annan. enter sesaneetetisee senken ehren flavescens

Larger, length 5.5 mm, body more depressed species, pronotum wider with wider base, ratios

w/l of pronotum >1.52, base/apex >1.48. Aedeagus longer and narrower (Fig. 188g), left

Paramere at apex straight (ISIISCHHSEH WAL HEN kennen CFUCIS, SPec. NOV.

Aedeagus narrower, lateral margin near apex faintly concave (Fig. 170g). Genital ring markedly

asymmetric (Fig. 170f). Puncturation of elytra less coarse and dense, on the average 3 punctures

Pro,interval.e 5A, Vic, ACT,.NSW, Old, cs NT, s. WA sa essenessnsnssesnenenesesnes rubiginosus Newman

Aedeagus wider, lateral margin near apex convex (Fig. 186g). Genital ring less asymmetric

(Fig. 186f). Puncturation of elytra coarser and denser, on the average 4 punctures pro interval.

SAH KNOW UVA Apache nen uanleeenssenranheebbaungneekentnafkene stern grossepunctatus, spec. NOV.

Tergum VIII with additional lateral setae. Apical setae of stylomere markedly elongate. Apex of

stylomere wide, rather obliquely transverse, the latero-apical angles rounded (Fig. 1721). NSW.

Ole. eisen ae RER re ER Be NEE foliaceus, spec. nov.

Tergum VII without additional lateral setae. Apical setae of stylomere not markedly elongate.

Sytlomere variable, though when wide and obliquely transverse, the latero-apical angles pro-

nounced.(Biss1701.41711,1811,.1831:1851,1871, 88 Dane ee an 138.

Apex of stylomere wide, obliquely transverse, the latero-apical angle pronounced, lateral plate

elongate (Fig. 1701). Puncturation of pronotum dense and rather coarse. SA, Vic, ACT, NSW, Old,

EN N N N rubiginosus Newman

Apex of stylomere different, when rather obliquely transverse, then either lateral plate shorter

and pronotum wider, or puncturation of pronotum sparser and much finer ........ueeee- 139.

. Apex of stylomere markedly convex on median side (Figs 1811, 1831, 1881). Puncturation of surface

always rather coarse and dense. Pronotum rather wide, ratio w/l usually >1.55, when <1.55, then

surfaceswith'markediy coarse and dense puneturation.........2...0eemseseseeseesscseensenasenensenen 140.

Apex of stylomere straight or oblique (Figs 1711, 1841, 1851, 1871). Puncturation of surface varied.

Pronotum rather narrow, ratio w/l <1.55, when >1.5, then surface with less corse and dense

PLUMGLIT TO N N ea ee ee enter nes inaesseinsuinieierte ne 142.

140. Stylomere narrow, apex acute (Fig. 1881). Larger, length 5.5 mm. s. WA ........... crucIs, Spec. NOV.

- Stylomere wider, apex convex (Figs 1811, 1831). Smaller, length <5.3 mm. Old ....................... 141.

141. Base of pronotum narrower, elytra generally longer, ratios w/l of pronotum 1.41-1.46, 1/w of

elytra SI 1.97 el. ae queenslandicus, spec. nov.

— Base of pronotum wider, elytra generally shorter, ratios w/l of pronotum 1.47-1.53, 1/w of elytra

148-154, ney Oldie ee RE A palumae, spec. nov.

142. Smaller, length <4.9 mm, colour yellowish to light reddish, with shorter elytra, ratio 1/w <1.55.

Elytra with rather coarse to very coarse puncturation (Figs 510, 524). Lateral plate of stylomere

short (Figs,1711,,185]) ...s.u:225:2:5x0 20:0 &000Ju0neos0gent anna nnnaenan una er Ser un aen naan aaer eaee 143.

— Larger, length >5.4 mm, colour reddish, with longer elytra, ratio 1/w >1.64. Elytra generally with

less coarse puncturation (Figs 523, 526). Lateral plate of stylomere elongate (Figs 1841, 1871) ....

EEE TE RER RR 144.

143. Pronotum narrower with narrower base, ratios w/1 1.40-1.51, base/apex 1.38-1.41. Puncturation

of elytra very. coarse (Fig, 524). Stylomere see fig. 1851. s.SA,n. Vie, sw. Old... tee re

anmannateelannehkngentesenessahaneeschreunteninstrnsessnssreetnengen neue EL een flavescens, spec. nov.

- _ Pronotum wider with wider base, ratios w/11.51-1.55, base/apex 1.44-1.48. Puncturation of elytra

less coarse (Fig. 510). Stylomere seeifie. 17112 sw. WA... an distinguendus, spec. nov.

144. Puncturation coarser, on elytra also denser (Fig. 523). Pronotum narrower with narrower base,

ratios w/l 1.47, base/apex 1.43. Stylomere see fig. 1841. d unknown. ce. Old ...............neneene

a NE angustatus, spec. NOV.

— Puncturation finer, on elytra also sparser (Fig. 526). Pronotum wider with wider base, ratios w/l

1.55, )base//apex 1:5: Stylomere see’fig. 1871. 3 unknown iw. SAen. nn ooldeae, spec. nov.

7.3.4. The species of the genus Adelotopus Hope

dytiscides-group

Diagnosis. Large, fairly wide, rather depressed, uniformly black species at most with indistinct

reddish apex of elytra. Labrum bisetose; glossa c. 8-setose; lateral margin of pronotum more or less

widely explanate, basal angles angulate; basal border line of elytra complete or almost so, usually

meeting behind scutellum, always completely visible; scutellar pore absent; lateral margin of elytra

usually without elongate setae behind shoulders, only in one species a short row of setae present;

series of umbilical pores with 6 subhumeral pores and 1 postmedian pore; abdominal sterna with at

least 1 ambulatory seta each side; sternum VI usually with a fringe of longer setae at apical margin;

tibiae, especially metatibia depressed; mesofemur and metafemur wide and depressed, profemur less

wide; internal sac of aedeagus simple, without oblique fold near apex; apex of ? tergum VIII without

setae.

Larvae. 1st instar larvae known of 4 species.

Distribution. 7 species from eastern Australia, southwestern Australia, and Papua New Guinea.

Systematic position. This group is perhaps most generalized in several aspects, e.g. large size, wide

body shape, rather large number of umbilical setae and of abdominal setae, comparatively simple

aedeagus. Most species look externally rather similar to primitive species of Pseudomorpha. Apparently

A. dytiscides is the most primitive species of this group and perhaps altogether of the entire genus.

Figs 100a-l. Adelotopus dytiscides Newman. Details of head and genitalia. a. Labrum. b. Lower surface of

terminal palpomeres of maxillary palpus. c. Lower surface of terminal palpomeres of labial palpus. d.5th-11th

antennomeres. e. d sternum VII. f. d genital ring. g. Lower surface of aedeagus. h. Lateral view of aedeagus.

i. Right paramere. k. Left paramere. 1. ? stylomeres and lateral plate.

Adelotopus dytiscides Newman, 1842

Figs 1, 30, 97-100, 289, 438, 603

Adelotopus dytiscides Newman, 1842, p. 365; Westwood 1853, p. 405, pl. 14, fig. 2; Lacordaire 1854, p. 154; Sloane

1898, p. 514 (dytiscoides); Blackburn 1901, p. 18; Lea 1910, p. 121 (dytiscoides); Notman 1925, p. 7, 10, 28

(dytiscoides); Csiki 1933, p. 1635; Matthews 1980, fig. 78 (dytiscoides); Erwin 1981, p. 64, fig. 28; Moore et al.

1987, p. 50.

Adelotopus fortnumi Hope, 1845, p. 105; Westwood 1853, p. 406; Lacordaire 1854, p. 154; Notman 1925, p. 28; Csiki

1933, p. 1635; Moore et al. 1987, p. 50 (synonymy confirmed by Notman 1925).

Adelotopus mastersii Macleay, 1871, p. 94; Notman 1925, p. 7, 10, 29; Csiki 1933, p. 1636; Moore et al. 1987, p. 51

(new synonymy).

Types. Of dytiscides. Lectotype (by present designation): 9, Type H.T., A. H. Davis, Adelaide, Adelotopus

dytiscoides Type Newman, Ent. Club 44-12 (BMNH).

Of fortnumi. Lectotype (by present designation): d, Type Westwood, Trans. Ent. Soc. 1845. p. 105 Coll. Hope

Oxon., Fortnumi Hope, Type Col: 20 4/4 Adelotopus fortnumi Hope, Hope Dept. Oxford (OUM). - Paralectotypes:

12, A. Fortnumi Hope N. Holland, Type Col: 20 2/4 Adelotopus fortnumi Hope, Hope Dept. Oxford (OUM);

12, Adelotopus Fortnumi Hope, Type Col: 20 3/4 Adelotopus fortnumi Hope, Hope Dept. Oxford (OUM); 19,43 ten,

Adelotopus Fortnumi Hope, Type Col: 20 1/4, Hope Dept. Oxford (OUM).

Of mastersii. Lectotype (by present designation): ?, Gayndah, Syntype, Adelotopus mastersii Macl. Gayndah

Old Syntype mastersii (ANIC-MMS). — Paralectotype: ?, K 12204, Adelotopus mastersii Macl. Gayndah, Holotype

(AMS).

Note. The sample from BMNH includes another specimen of dytiscides bearing a (printed) label

“Type”. It cannot serve as a type, however, because it lacks a locality label.

The synonymy of fortnumi with dytiscides firstly stated by Notman (1925), as well as the new

synonymy of mastersii with dytiscides, is confirmed in present paper by comparison of the types.

Type localities. Of dytiscides: “Adelaide”, South Australia. - Of fortnumi: From description: “Ad-

elaide”, South Australia. - Of mastersii: "Gayndah”, Queensland.

ZEN,

Diagnosis. Large, wide, moderately depressed, uniformly black species with straight base of prono-

tum and rasp-like punctures marking the elytral striae. Distinguished from other species by large size,

not horseshoe-shaped elytral punctures, and elongate, narrow, rather symmetric aedeagus.

Description

Measurements. Length: 7.4-10.2 mm. Ratios. Width/length of pronotum: 1.75-1.88; width base/

apex of pronotum: 1.80-1.95; width pronotum/head: 1.92-2.03; length/ width of elytra: 1.38-1.46; length

elytra/pronotum: 2.48-2.66.

Colour. Black, lateral borders of pronotum and elytra somewhat reddish-piceous translucent.

Lower surface, antennae, mouthparts, and legs dark reddish-piceous, lower surface of head dark

piceous.

Head (Figs 98, 99, 100a-d, 289). Short and wide, rather depressed. Anterior border feebly convex,

lateral border feebly angular. Clypeal suture shallow, though usually complete and distinct, semicir-

cular. Labrum comparatively large, apex slightly concave, bisetose, though sometimes asymmetrically

with an additional seta. Antennal groove laterally barely bordered, latero-posteriorly with convex, not

carinate area. Mental tooth triangular, though wide, at apex rounded off. Wings of mentum wide, near

apex strongly rounded. Glossa wide, tongue-like, apically rounded, ventrally with keel, at border with

c. 8 elongate setae and additional dense pilosity on upper and lower surface and along border.

Terminal palpomere of maxillary palpus narrow, apex feebly widened, barely securiform. Terminal

palpomere of labial palpus large, strongly widened, trhough not wider than long, securiform. Antenna

relatively elongate, 8th-9th antennomeres c. as long as wide. Microreticulation dense and distinct,

puncturation rather dense, surface with a shallow sulcus medially of eyes and usually with some fine

longitudinal strioles on frons, impilose, markedly dull. Ventrolaterally of eyes with a row of short

setae. Suborbital field densely punctate and setose. Both palpi, as well as gula rather densely setose.

Pronotum (Figs 30, 289). Very wide, but moderately convex, base almost 2 x as wide as apex.

Apical angles remarkably produced, acute, attaining at least middle of eyes. Apex deeply, rectangu-

larly excised, slightly convex in excision, not bordered. Sides strongly and evenly curved, widest in

basal third or even at basal angles, barely or not at all bordered, but, especially apically, widely

explanate. Basal angles rectangular or slightly obtuse, at apex slightly rounded off, base almost

straight, not bordered. Microreticulation dense and very distinct, almost isodiametric, puncturation

fine, difficult to detect, surface with several fine, irregular strioles, impilose, dull.

Elytra (Figs 30, 289, 438). Moderately elongate and convex, in basal two fifth rather parallel, then

gently narrowed. Apex slightly oblique, truncature feebly concave, dehiscent at suture. Shoulders

obtuse, basal margin oblique, with a row of many short setae. Basal border line complete, meeting at

suture. Marginal channel rather wide, slightly widened in basal fourth, completely visible from above.

Lateral border asetose or with but 1-2 short setae just behind shoulders. Series of umbilical pores

consisting of 6 pores behind shoulder and 1 pore at or slightly behind middle. Setae short. Striae almost

completely reduced, but sometimes sutural stria just visible as a row of extremelvy fine strioles, or even

traces of the internal striae visible at apex. Microreticulation dense and very distinct, puncturation

dense and rather coarse, punctures becoming fairly rasp-like towards apex. Surface impilose, marked-

ly dull.

Lower surface. Prosternal process rather elongate, straight, moderately depressed, apex rectangu-

lar, shortly setose. Metepisternum elongate, >2 x as long as wide, posteriorly constricted and

hollowed. Abdominal sterna with 4-6 setae each side, sternum VI in both sexes with many (c. 16-30),

fairly elongate setae at and near apical border. Whole lower surface rather densely pilose.

Legs. Elongate, 1st tarsomere of protarsus not wider than long, tibial groove of profemur

comparatively shallow, anterior plate only at apex distinctly overlapping the groove, posteriorly

border of groove not sharp. Metatibia rather narrow, >5 x as long as wide, 1st tarsomere of metatarsus

>2 x as long as wide. d protarsus barely widened.

d genitalia (Figs 100e-k). Genital ring rather asymmetric, laterally convex. Sternum VII apically

deeply excised, basally slightly bisinuate, laterally short. Aedeagus elongate, narrow, depressed,

rather symmetric, lower surface straight. Lateral border wide throughout. Apex widely rounded off.

Internal sac rather simple. Both parameres rather similar, elongate, with obtuse apex.

? genitalia (Fig. 1001). Stylomere triangular, rather wide, with 2-4 subapical setae. Lateral plate

moderately elongate, with 2-3 apical setae.

un _ u _ rn m ZU

Variation. Due to extensive range and large number of specimens available, rather wide variation

noted, especially in shape and relative width of pronotum which may be slightly narrower and more

incurved to base, by relative length of elytra, and density of microreticulation and puncturation.

Otherwise, a rather homogenous species. The types of mastersii are founded on relatively small, convex

specimens with basally more rounded pronotum.

Vivipary. Confirmed by discovery of larvae in the 2 oviducts.

Habits. A common and widespread species, apparently occurring in a variety of more or less open

forests and woodlands, collected by me under bark of different eucalypts. Although few collecting

circumstances are noted, these show a rather different range: repeatedly under bark of unspecified

eucalypts, one “from Red Gum bark at night”, but also “on Melaleuca”, “in woodland”, “on car”, and

some specimens “from ant nest”. One specimen is mounted together with the presumable host ant of

the genus Camponotus Mayr (species near nigriceps Sm.) on the same card. Dated specimens collected

from September to April and in June, but most in the period from October to November.

Distribution (Fig. 603). Southeastern Australia from eastern half of South Australia to central

Queensland, southwestern corner of Western Australia, ? Tasmania. This last record is rather doubtful.

Material examined (658). SA: 17, Type H.T., A. H. Davis, Adelaide, Adelotopus dytiscoides Type Newman, Ent.

Club 44-12 (BMNH); 19, Ent. Club 44.12, A. H. Davis, Adelaide, det. dytiscoides (BMNH); 14, Adelaide, 9.X1.12,

Griffith, Griffith Coll., Id. by A. M. Lea (SAMA); 1? (?), Adelaide 1871, gyrinoides Hope, Nov. Holl. (NHMW);

338,1?, Adelaide, A. H. Elston, A. H. Elston Coll., Adelotopus dytiscoides Newm. (AMS); 13, Adelaide, gyrinoides

Hope, Coll. L. W. Schaufuss, det. dytiscoides (MNHB); 12 (2), 41406 Adelaide Schaub., det. dytiscoides (MNHB);

1?, Adelaide, Bowring 63.47, A. Fortnumi Hope, det. dytiscoides (BMNH); 19, Sharp Coll., Adelotopus ipsoides

Westw., Adelaide, det. dytiscoides (BMNH); 1%, Dytiscoides Newm., gyrinoides Germ., Adelaide, Sharp Coll.

(BMNH); 15, 222, Adelaide Hart, Sharp Coll., det. dytiscoides (BMNH); 1? (2), Adelaide, Sharp Coll., det.

dytiscoides (BMNH); 17, Adelaide, det. dytiscoides (BMNH); 19, Adelaide Griffith "04, E. W. Ferguson Coll.,

Adelotopus dytiscoides Newm. (ANIC); 15,1%, Adelaide, Adelotopus gyrinoides Hope, H. Burnside (?) 239 (SAMA);

222, Adelaide Blackburn (SAMA); 13, Adelaide, dytiscoides W., Id. by A. M. Lea, Lea Dup. (SAMA); 16,

Adelaide, Lea Dup. (SAMA); 19, Adelotopus Adelaide (OUM); 1?, Gyrinoides Hope Adelaide (OUM); 1%,

Adelaide (OUM); 13, A. H. Davis, Adelaide E. Newman, W (OUM); 19, Adelaide, ?, W. (OUM); 1®, Adel.,

Adelotopus gyrinoides Hope (OUM); 62%, Adelaide Coll. Castelnau (MCSN); 1?, Adelaide, Adelotopus fortnumi

Adelaide, Adelotopus dytiscoides Newm. (NHMB); 19, Adelaide (NHMB); 1?, Adelaide, Axinophorus n. sp.,

A. haemorrh. inguinatus Newm. (NHMB); 14, Adelaide, inguinatus Adelaide (NHMB); 1%, Clarendon, Griffith

Coll. Id. by A.M. Lea (SAMA); 14, Mt. Pleasant, 8.X1.86 Tepper, det. dytiscoides (MCZ); 19, Mt. Lofty, J. G. O.

Tepper, A. dytiscoides N. As id. by Blackburn (MCZ); 1?, Mt. Lofty, J. G. ©. Tepper, Ex coll. SA Mus., A. dytiscoides

Newm. As id. by Blackburn (ANIC); 22%, Mt. Lofty Rgs. H. Curnow, E. W. Ferguson Coll. (ANIC); 13, 228,

Mt. Lofty, J. G. ©. Tepper (SAMA, UOIC); 15, Mt. Lofty, J. G. ©. Tepper, Adelotopus dytiscoides Newm., Id. by

R. Harvey (SAMA); 25, 62%, Mt. Lofty Rgs. S. H. Curnow, dytiscoides Newm., Id. by A. M. Lea, Lea Dup.

(SAMA); 15,322, Mt. Lofty Rgs. S. H. Curnow, ac. 23246, dytiscoides Newm. (AMNH); 19, Belair 4.X.85, Tepper

(SAMA); 15, Hahndorf, H. J. Carter Coll., det. dytiscoides (NMV); 18, 522, Yorketown, CNHM 1955, Karl

Brancsik Coll, ex Eduard Knirsch, Adelotopus dytiscoides det. Ball (FMNH, UASM); 1?, Maitland, Yorke Pen.

11.X1.1964, D. Wright (SAMA); 13, Cleve (UOIC); 1%, Pt. Lincoln: Lea (SAMA); 12, Kangaroo Is. J. G. ©. Tepper

(SAMA); 15, 12, Kangaroo I1., F. Wood Jones (SAMA); 22%, Blackburn, ditto S. Australia 797 Quor (SAMA);

15, Parachilna, H. Hale, Flinders Range (SAMA); 15, Williamston, 20.X.88 Tepper, dytiscoides Newm. Id. by A.

M. Lea (UOIC); 1%, A. dytiscoides Newm. v. Müller 77 (SMNS); 1%, v. Müller 77, Müller 1872, Alte Sammlung

(SMNS); 1 (2), dytiscoides Newm. (ANIC); 12, Janson Acq. 1884 (MNHN); 22%, Adelotopus colymbetoides 9°24.

(Bakewell), det. dytiscoides (BMNH); 12, Pascoe Coll., det. dytiscoides (BMNH); 15, det. dytiscoides (BMNH);

19, W. Edwards, det. dytiscoides (MCZ); 18, 12 (2); ?, 10.X.32, Ex Coll. H. V. Southcott (SAMA); 1, Rev. A. F.

Burgess, Lea Dup. (SAMA); 12, C. French’s Coll., det. mastersii (NMV); 1?, C. French’s Coll., Adelotopus

gyrinoides (Hope), det. mastersii (NMV); 256, 1? (NMV). - Vic: 13, Mallee District, C. French’s Collection, det.

mastersii (NMV); 18, 12, Kaniva, 27.X., 28.X., J. Sedlacek (CSB); 18, 8km W. of Kaniva, 21.X.1963, J. Sedlacek

(BMH); 42%, Little Desert, 21.X.46-173 (NMV); 1%, Sea Lake, IV.1914 Goudie, Adelotopus dytiscoides Newm.

(NMV); 15, Grampians X1.92 (NMV); 19, K. Guichard, Hall’s Gap, Adelotopus dytisccoides (BMNH); 19, Birchip,

20, Adelotopus dytiscoides Newm., J. C. Goudie Collection (NMV); 14, 15 km W. Ballarat, 12.X1.1969, E. F. Riek

(ANIC); 19, Bacchus Marsh XII., gyrinoides Hope C. Langenhan, Samml. O. Langenhan (SMTD); 2? ?, Bacchus

Marsh, XI1.1904, Van Dyke Coll., Adelotopus dytiscoides Newm. (CAS); 12, Bacchus Marsh X11.03 (NMV);

13, Geelong, Davey, dytiscoides N., Id. by A. M. Lea, S. Aust. (SAMA); 1%, 1683, Goul, dytiscides Newm. Port

Philipp (MNHB); 1, Pt. Philip, 691 Adelotopus fortnumi Hope (OUM); 37%, Melbourne Coll. Castelnau, Adel.

Dytiscoides New. Melbourne (MCSN); 12, Melbourne Coll. Castelnau, Dytiscoides New. gyrinoides Germ. Melb.

(MCSN); 18,322, Melbourne Coll. Castelnau (MCSN); 12, Melbourne, Deane (UOIC); 18, Adelotopus gyrinoides

Hope Melbourne, det. dytiscoides (MNHB); 27°, Melbourne, det. dytiscoides, fortnumi Hope, Pascoe Coll.

(BMNH); 18, Melbourne, det. dytiscoides (BMNH); 1%, Melbourne Bowring 63-47, det. dytiscoides (BMNH);

12, Melbourne, W. Edwards, det. dytiscoides (MCZ); 12, Melbourne, Adelotopus Hope (HNMB); 12, Melbourne

No. 664, Ejnar Fischer (NHRS); 17, Melbourne F. E. Wilson 14.X11.18, Adelotopus dytiscoides Newm. Id. by T.G.

Sloane, F. E. Wilson Coll. (NMV); 222, Melbourne, Coll. C. Felsche, det. dytiscoides (SMTD); 1?, Melbourne

(NHRS); 1%, From D. Howitt, Adelotopus dytiscides Newm. Melbe. (NMV); 272%, Austral mer. Melb. Melly,

Australie, Ex Musaeo Mniszech, det. dytiscoides Newman (MNHN); 1%, A. ditiscoides Nwm. P. Philip (?), Soc. Ent.

Belg, Coll. PUTZEYS, Adelotopus dytiscoides Newm. det. J. Putzeys (IRSNB); 12, Ringwood, 15.X1.11. 18/22,

H. Pottinger (QMB); 222, F. T. Gully (UOIC); 19, Kerrisdale 1.1.24 (NMV); 32, Vic 142, Big River, 20 km ssw

Jamieson, Lake Eildon SP, 16.X11.1990, M. Baehr (CBM); 19, 12km SE Merrijig, Howqua River 30.X1.1971,

Neboiss, det. dytiscoides (NMV); 32%, Vic 140, 20 km s. Mansfield, 16.XI1.1990, M. Baehr (CBM); 622, Vic 130,

15 km s. Mt. Beauty, 13.X11.1990, M. Baehr (CBM); 359,322, Eltham, 17.1V.23, 17.IX.27 (NMV); 12, Eltham ]J. F.

Dixon (NMV); 14, 1?, Benalla Helms X-X11.89, Adelotopus gyrinoides Hope, Helms Coll. (BMH); 27%, Benalla

Helms, Adelotopus mastersii Macl. (BMH); 1%, Benalla, Adelotopus dytsicoides Newm., Lea Dup. (SAMA);

12, Benalla (SAMA); 1%, Black Rock, J. F. Dixon (NMV); 18, Tarawara 10.X1.1912 (NMV); 18, Ocean Grove

22.X.54 A. N. (NMV); 18, C. E. Clarke Coll., Koetong N. Vic. 19.X1.30 (BMNH); 1%, Cadwega 20.X1.30,

C. E. Clarke Coll. (BMNH); 1%, Clifton 22.X.04, Adelotopus dytiscoides Newm., Griffith Coll. Id. by A. M. Lea

(SAMA); 1%, Melton, X1.40. F. E. W., J. G. Brooks Bequest, 1976 (ANIC); 1%, Rushworth 18.X1.51, F. E. Wilson,

Adelotopus dytiscoides Newm., ex Coll. F. E. Wilson (WAM 87/2169); 17, Rushworth XI1.45, FEW, J. G. Brooks

Bequest, 1976 (ANIC); 32?, 6 ml. SE. of Rutherglen, 3.X1.1950, J. H. Calaby (ANIC); 13, 17, Kandella,

S. Gippsland, French, Coll. Kraatz, det. dytiscoides (DEIB); 17, Sea Lake, X.1915, Goudie (SAMA); 19, Alexandra,

CNHM 1955, Karl Brancsik Coll, ex Eduard Knirsch, dytiscoides Newm. det. G. E. Ball (FMNH); 17, Alexandra,

CNHM 1955, Karl Brancsik Coll, ex Eduard Knirsch (UASM); 22%, Upper Ovens R., 25.X11.59, B. P. Moore

(CMC); 2364, 22%, Victorian Alps Blackburn, Dytiscoides Newm., Adelotopus dytiscoides N. 797 AL (SAMA);

2?2, Victorian Alps Blackburn, 797 AL (SAMA); 16, Vict. Alps No. 664 Ejnar Fischer, Adelotopus dytiscoides

Newm. Id. E. Fischer (NHRS); 1%, Vict. Alps No. 664 Ejnar Fischer (NMV); 14, 2?%, Mitcham, 15.X1.58,

B. P. Moore, Adelotopus nr. dytiscoides Newm., det. B. P. Moore (CMC); 333, 922, 34.355, 132.46E, Robinvale,

25.X.-3.X1.1988, T. Weir, J. Lawrence & M. Harrison (ANIC); 13, Ocean Grove, H. W. Davey (UOIC); 16,

19, Kiata, 31.X.50 F. E. Wilson, F. E. Wilson Collection, det. dytiscoides (NMV); 19, Kiata X1.58 F. E. Wilson,

F. E. Wilson Coll., det. dytiscoides (NMV); 12, Woori Yallock 29.X11.36. F. E. Wilson, F. E. Wilson Coll., det.

dytiscoides (NMV); 12, Noble Park, F. E. Wilson 9.X1.18, Adelotopus dytiscoides Newm. (CAS); 15, Linga, X.1922

F. E. Wilson, Adelotopus dytiscoides Newm., F. E. Wilson Collection (NMV); 14, 19, Janson Acgq. 1884 (MNHN);

229, CNHM 1955, Karl Brancsik Coll, ex Eduard Knirsch, Adelotopus dytiscoides Nemw. det. G. E. Ball 1987

(FMNH, UASM); 684,212, det. dytiscoides (MNHB); 1%, gyrinoides Hp., det. dytiscoides (MNHB); 15,17, Lane,

S. Rolle V. 94911, det. dytiscoides (MNHB); 14, Sharp Coll., Adelotopus dytiscoides Newm. (BMNH); 13, Edwards,

Adelotopus dytiscoides Westw., Sharp Coll. (BMNH); 1%, Edwards, Fry Coll., det. dytiscoides (BMNH); 1%, Fry

Coll., det. dytiscoides (BMNH); 1%, 1937, M. F. C., det. dytiscoides (BMNH); 3?%, M. F. C. 1935, det. dytiscoides

(BMNH); 229, Helms, W. W. Froggatt Coll. Adelotopus dytiscoides Newm. (ANIC); 1%, D. Castelnau 1867,

Adelotopus dytiscoides Newm. teste Putzeys (MCSN); 14, Adelotopus dyticides (sic!) Newm., 8 Howitt Colln.

(NMV); 18, 12, NO. Collection Hy. Edwards, det. dytiscoides (AMNH); 15, 2?%, Edwards Coll., det. dytiscoides

(AMNH); 22%, 2593, det. dytiscoides (AMNH); 15, Museum Leiden ex collection C. J. Louwerens rec. 1979,

Adelotopus dytiscoides Newm. det. Darlington ’48 (NNML); 1%, (not readible), Adelotopus dytiscoides Newm.

(NMV); 18, 42? (NMV). - Tas: 12, 25903, det. dytiscoides (AMNH). - ACT: 19, J. W. Evans, Canberra F.C.T.

6.X1.1919 (ANIC); 18, J. W. Evans, Canberra FCT 6.11.1929 (ANIC); 18, 12, Canberra 1.1960, D. Carne (ANIC);

1? (?); Canberra 16.X.63, J. Sedlacek (CSB); 1?, Canberra, NSW 1926 (ANIC); 22 2, Red Hill, 23.X1.1968 K. Pullen,

Kim Pullen Coll. (ANIC); 14, Black Mtn. 20.X1.66 K. Pullen, Kim Pullen Coll. (ANIC); 1%, Black Mtn. 31.X.67 K.

Pullen, Kim Pullen Coll. (ANIC); 16, Black Mt., X1.65, K. Pullen, Adelotopus mastersii, Kim Pullen Coll. (ANIC). —

NSW: 12, Bombala K 2389 (AMS); 32%, Illawarra, G. E. Bryant, 17.1X.08, G. Bryant Coll, det. dytiscoides (BMNH),;

15, 34.245, 143.14E, 3km NW of Bramah H.S. NW of Balranald, 24.X.1983, D. C. F. Rentz & M. S. Harvey Stop

35 (ANIC); 12, Brocklehurst (32.115, 148.38E) 29.IX.71, S. Misko (ANIC); 234, 2??, Sydney Coll. Castelnau

(MCSN); 19, Sydney K.12228 (AMS); 2543, 37%, S. Australia, Sydney, Taylor, Adelotopus dytsicoides Newm. Bel.

02 (AMS); 13, Sydney Deane (NMV); 12, Sidney, Adelotopus dytiscoides Newm., Coll. L. W. Schaufuss (MNHB);

42 2,Sydney X.02, Helms Coll., Adelotopus gyrinoides Hope (BMH); 17, N. Sydney, G. E. Bryant, 7.X.08, G. Bryant

Coll, Adelotopus Dytiscides Newm. (BMNH); 284, N. Sydney, G. E. Bryant, 3.X.08, 30.X1.08, G. Bryant Coll, det.

dytiscoides (BMNH); 32%, Newport, N. Sydney, G. E. Bryant 23.X.08, G. Bryant Coll., det. dytiscoides (BMNH);

?, North Shore, Taylor, Coll. Hacker, Adelotopus dytiscides Newm. (DEIB); 12, Sydney district, J. J. W. X.1900,

Adelotopus gyrinoides Hope, det. dytiscoides (BMNH); 148, between Orange and Sydney, Hale 5.X.29 (ANIC);

15, Lane Cove, 15.11.1948, H. W. Rudd (AMS); 19, Canterbury IX.01, H. J. Carter Coll., Adelotopus dytiscoides

Newm. (NMV); 18, Mt. Wilson, 23.1X.1972, D. A. Doolan, D. A. Doolan Coll. (AMS); 18, Mt. Wilson X.1930

F. E. Wilson, F. E. Wilson Coll., det. dytiscoides (NMV); 18, Adelotopus diticides (sic!) Goulburn Riv. (OUM)

1?, Bogan River, X1.1935, J. Armstrong (AMS); 14, Newcastle 1.1951, J. Sedlacek (CSB); 13, Up. Hunter Riv.

Valley 20.X1.69, Collr. J d’Apice 5122, Cainogenion ipsoides (Westwood) [Series det. by A. Walford-Huggins]

(CMP-WHC); 12, Caldwell, V. Robb, det. dytiscoides (AMS); 22?, Gunning Gap, 17.X.66, B. P. Moore (CMC);

12, Bathurst 15.1V.1950, Adelotopus dytiscoides (CSB); 12, Bathurst, det. dytiscoides (NMV); 2 (2), Wagga (MMS);

12, Wahroona, H. J. C. 1.15, Adelotopus dytiscoides Newm. Id. by T. G. Sloane, H. E. Andrewes Coll. (BMNH);

18, Wahroona, H. J. Carter (ANIC); 18, 2 mi. S. Mendooran, 4.-5.X.1970, D. K. Mc Alpine and G. A. Holloway

(AMS); 12, Bulga 24.IX.20, WWF (ANIC); 228, Greta 1.X1.51, J. Sedlacek (CSB); 1%, Gunnedah, 5.X1.1982,

J. Doyen coll. (ANIC); 18, 12, Gunnedah 5.11.59 F. E. Wilson, F. E. Wilson Coll., det. dytiscoides (NMV); 1%,

A. mastersii Macl. Tamworth (SAMA); 1?, Boggabri, Tamworth, 8.X1.1932, K. C. Mc Keown (AMS); 14,12, Forest

Reefs, Lea, Field Museum 1970 ex South Australian Museum (FMNH); 14, Forest Reefs, Lea, Adelotopus

dytiscoides N. Victoria (QMB); 19, 17, 5602 For. Reefs, A. mastersii teste Lea, Adelotopus mastersii Macl. (SAMA);

256, Adelotopus mastersii Macl. Forest Reefs, G. C. Champion Coll. (BMNH); 14, Forest Reefs Lea, C. French’s

Coll., det. mastersii (NMV); 12,50 km N. Glen Innes, 20.9.1985, leg. T. Hawkeswood (CBM); 12, 28 km SW of

Tenterfield, 5.X.71, S. Misko (ANIC); 18, 12, 3km NW of Tooraweehnah, 1.X.1971, S. Misko (ANIC);

12, Clarence River Coll. Castelnau (MCSN); 19, Gundagai X.1930 F. E. Wilson, F. E. Wilson Coll., A. dytiscoides

(NMV); 15, Adelong, X.1930, F. E. Wilson, Adelotopus dytiscoides Nw. (MCZ); 12, Quirindi, G. E. Bryant, X1.1908,

Adelotopus dytiscoides Newm. (FMT); 12, Guta, Coll. Dr. Reitter, Adelotopus dytiscoides Newm. Det. B. P. Moore’66

(FMT); 12 (0), Willington 1906 W. W. Froggatt Coll, Adelotopus dytiscoides Nw. (ANIC); 12, Blackheath,

4.X11.1946, C. Oke (NMV); 17, Capertree, XI1.33, Deuquet Coll., det. dytiscoides (CAS); 1%, Ihule (?), X1.30

G. Goudie, J. C. Goudie Collection (NMV); 12, Uah T. G. 5. 8.X.23 (ANIC); 2? 9, Masters, Fry Coll., det. dytiscoides

(BMNH); 22%, Lea, A. Fenyes Coll., dytiscoides Newm. Adelotopus Hope (CAS); 18, Gyrinoides Hope sec.

W. Macleay, Ex Museo H. W. Bates 1892 (MNHN); 15, Adelotopus dytiscoides Newm. Carabidae (CAS); 12, Coll.

Carl Schuchardt, det. dytiscoides (SMF); 15, Coll. v. Schönfeldt, det. gyrinoides (SMF); 12, Coll. v. Schönfeldt,

gyrinoides Hope (SMF); 15, Schaufs. N. Holl., Museum Leiden Adelotopus gyrinoides Hope Det., Gyrinoides Hope

(NNML); 12, Coll. Hacker, det. dytiscoides (DEIB); 12, T. G. S. 24 (ANIC); 1?, 82 Adelotopus dytiscoides Newm.

(OUM); 334,429, Janson Acgq. 1884 (MNHN); 15, Ex Museo L. Fairmaire 1896 (MNHN); 12, Edwards Coll., det.

dytiscoides (AMNH); 1%, 15174, det. mastersii (SMTD); 1? (MNHN); 1? (BMNH). - Qld: 12, Stanthorpe 5.X1.1974,

J. Sedlacek coll. (CSB); 18, Wyberba, 1.1.37, E. Sutton, E. Sutton Coll. (QMB); 1%, Fletcher, E. Sutton, E. Sutton

Coll. (QMB); 17, Brisbane Coll. Castelnau, Brisbane (MCSN); 1%, Brisbane 6.1X.30 (UQIC); 12, Brisbane Illidge

(UOIC); 17, Brisbane: H. Hacker 30.X.21 (QMB); 16, Brisbane H. Hacker 8.1X.15 (QMB); 12, Brisbane. O. W. Tiegs

Adelotopus gyrinoides Hope (QMB); 17, Brisbane, H. Hacker (QMB); 22%, Brisbane (SAMA); 14, Brisbane Mc

Gregor, Ditto Queensland (SAMA); 1%, Brisbane, Ex Musaeo Mniszech, ditiscoides West. (MNHN);

2?%, Archerfield, 19.IX.1964, B. Cantrell (UOIC); 484, 12, Sandgate, Coll. F. Muir IX.1919 (BMH); 1%, Belmont,

16.V1.1964, A. Terauds (UOIC); 17, Nudgee, Brisbane H. Hacker 8.X.23 (QMB); 14, 17, Moreton Bay, Stradbroke

I., IX.1915, J. C. Bridwell Coll., det. dytiscoides (USNM); 15, 2? (?), Moreton Bay, det. dytiscoides (BMNH),;

12, Moreton Bay, Bowring 63-47, det. dytiscoides (BMNH); 13 (2), 1?, K 12228, Ipswich, det. gyrinoides (AMS);

13, Yarraman, 10.X.1979 (CSB); 12, Rivertree 9.X.31, E. Sutton, E. Sutton Coll. (QMB); 12, Milmeran S.Q., IX.50.

SME, J. G. Brooks Bequest, 1976, dytiscoides New. (ANIC); 12, Millmerran SQ, 1.1959, J. Macqueen, Adelotopus

dytiscides Newman [Series det. by A. Walford-Huggins] (CMP-WHC); 17, Qld 19, Burnett R. 10 km n. Eidsvold,

9.X1.1990, M. Baehr (CBM); 1%, Adelotopus mastersii M“L. Jun. Burnett River, 52 Howitt Colln. (NMV);

12, Gayndah, Syntype, Adelotopus mastersii Macl., lectotype! (ANIC-MMS); 1%, K 12204, Adelotopus mastersii

Macl. Gayndah, Holotype, paralectotype! (AMS); 17, X1.1986, Monto, J. Sedlacek Igt. (CSB); 14, 1?, Dawson dist.

(Barnard coll.) (MNHN); 1% (?), Mackenzie River, 29.1.1968, leg. G. Hangay (HNMB); 14 (?), Bowen A. Simson

294 (SAMA); 17, Fry Coll., det. dytiscoides (BMNH); 1%, Janson Acq. 1884 (MNHN); 1%, North N. Holl, Janson

Acq. 1884 (MNHN), 1%, N. Queensland, 1989 25, det. dytiscoides (SMTD); 12, N. Queensland, Adelotopus

gyrinoides, c.151 )IRSNB); 15, 22, det. mastersii (NMV); 12, C. French’s Coll., Adelotopus mastersii (Macleay)

(NMV); 1%, Klages Coll. Exot. Coleopt. C. M. Acc 2275, Adelotopus dytiscoides Newm. (CMP). - WA: 1?, Swan

River Curlois 1869 (OUM); 1?, Newcastle, Lea, Adelotopus dytiscoides Newm. W.Aust.: S. Aust. (SAMA); 19,

WA 49, 33 km nnw. Raventhorpe, 13.X1.1987, M. Baehr (CBM); 1%, Fortnumi Hope, Dytiscides Nw. = Fortnumi

Hope (OUM); 15, 801 263, ipsoides (HNMB). - Aus: 1?, Australia mer. Coll. Kirsch, det. gyrinoides (SMTD);

17, Costa orientale D. Castelnau 1867, Adelotopus ditiscoides (sic !) Newm. t. Putz. (MCSN); 17, A. Deyr. N. Holl.,

Museum Leiden Adelotopus dytiscoides Newm. Det., fortnumi Hope (NNML); 12, 480, Adelotopus ipsoides Wst.

C. Langenhan, Sammlung O. Langenhan (SMTD); 1? Coll. Castelnau (MCSN); 12, A. Fortnumi Hope, Type Col:

20 2/4 (OUM); 15, Type Westwood, Fortnumi Hope, Type Col: 20 4/4 (OUM); 1%, Adelotopus Fortnumi Hope,

Type Col: 20 3/4 (OUM); 1%, 43 ten, Adelotopus Fortnumi Hope, Type Col: 20 1/4 (OUM); 19, Baly (OUM);

12, colymbetoides 14.25 Muir Coll. (OUM); 14, 12, Koebele, Adelotopus dytiscoides Newm. det. H. Notman

(USNM); 18,22%, Koebele, det. dytiscoides (USNM); 18, 222, Koebele, Koebele Coll. det. mastersii, Adelotopus ?

mastersii Macleay (CAS); 1?, CNHM 1955 Karl Brancsik Coll, ex Eduard Knirsch, Adelotopus mastersii Macleay,

Adelotopus dytiscoides Newm. det. G. E. Ball (FMNH); 1%, 1872 leg. Müller, Adelotopus dytiscoides Newm. det.

M. Baehr (CBM); 14, 12, Müller 1872, Alte Sammlung (SMNS); 19, 12, Alte Museums-Sammlung, A. dytiscoides

fortnumi Hope (NHMB); 1%, Dohrn 91, Adelotopus dytiscoides Newm., dytiscoides Cast. (NHMW); 13,222, Collect.

Plason, Adelotopus gyrinoides Hope (NHMW); 17, Ipsoides, Ex Museo L. Fairmaire 1896 (MNHN); 22%, Putzeys,

Coll. E. Witte, det. dytiscoides (SMF); 15, Coll. E. Witte, decipiens ?, det. dytiscoides (SMF); 15, Fundort ?, Coll.

B. Schwarzer, Adelotopus dytiscoides Newm. (SMF); 1?, Fundort ?, Coll. B. Schwarzer, det. gyrinoides (SMF);

13, 19,C. Strobl, Coll. B. Schwarzer, det. gyrinoides (SMF); 1%, Adelotopus dytiscoides Newm. Coll. Schaum (DEIB);

1?, Coll. Kraatz, det. dytiscoides (DEIB); 15, Coll. Kraatz, det. ipsoides (DEIB); 15, 17, det. dytiscoides (MNHB),;

12, Fortnumi Hope, Coll. L. W. Schaufuss, det. dytiscoides (MNHB); 12, Adelotopus gyrinoides Hope, Strobl a 73,

det. dytiscoides (MNHB); 1%, Adelotopus ipsoides Westw. Köchl., det. dytiscoides (MNHB); 1%, Pars, det. dytiscoides

(MNHB); 1?, 1882 I, det. gyrinoides (NHMW); 1%, 1882 I, Adelotopus mastersii, det. gyrinoides (NHMW); 1%, 801

263 (HNMB); 1%, 737, det. dytiscoides (BMNH); 12, Adelotopus gyrinoides Hope, Sharp Coll., det. dytiscoides

(BMNH); 334, 19, 58124, det. dytiscoides (BMNH); 18, Fry Coll., det. dytiscoides (BMNH); 15, 13524, Austr., Fry

Coll., det. dytiscoides (BMNH); 2? %, Bowring 63.47*, det. dytiscoides (BMNH); 277, Blackburn, Sharp Coll., det.

dytiscoides (BMNH); 2? 2, Blackburn, G. C. Champion Coll. (BMNH); 19,22%, Edwards, det. dytiscoides (MCZ);

322, W. Edwards, dytiscides Newm. (MCZ); 17, W. Edwards, Adelotopus dytiscoides (MCZ); 18, 12, Adelotopus

dytiscoides Newm. 554 (MCZ); 1%, Coll. French (ANIC); 15, Simson’s Coll. 294-442 (SAMA); 18, Blackb’s Coll.,

mastersii Macl. (SAMA); 234,2? 2, Coll. Schaum, det. dytiscoides (DEIB); 1%, Coll. Schaum, Adelotopus gyrinoides,

det. dytiscoides (DEIB); 12, fortnumi Hope, Coll. Schaum, det. dytiscoides (DEIB); 17, 13524, 6.660 Muir Coll.

(OUM); 12, 090 Adelotopus Hope (OUM); 19, gyrinoides Hope quest. W. Macl., Ex Museo Van Lansberge

(MNHN); 12, Ex Musaeo E. Steinheil, det. dytiscoides Newman (MNHN); 14, Adelotopus dytiscoides Newm. det.

P. Dupuis, Adelotopus dytiscoides Newm. (IRSNB); 17, Adelotopus gyrinoides Hope det. P. Dupuis, Adelotopus

gyrinoides Hope (IRSNB); 1%, Dohrn, Adelotopus dytiscoides Newm. det..., Adelotopus dytiscoides Newm. Austral.

(IRSNB); 12, Austral. Adelotopus gyrinoides Hope det... (IRSNB); 1%, A. ditiscoides Nwm. N. Holl (2), Soc. Ent. Belg,

Coll. PUTZEYS, Adelotopus dytiscoides Newm. det. J. Putzeys (IRSNB); 13 (OUM). - ?:2?%, Inglewood 11.XI1.20

(NMV); 42%, Oakleigh (NMV); 12, Elmore 11.11.10 (NMV); 12, Whittlean (2), J. A. K. 11X1.08 (NMV);

19, Crayton 26.1V.17 (NMV); 1?, Champion Bay Douboulay (OUM); 12, W. Dist. Kershaw XI.01, Griffith Coll.

12, Oakleigh, A. dytiscoides (NMV); 17, Morcatta, dytiscoides Newman (OUM); 12, Maitland 29.X1.57, Adelotophus

(sic!) sp., det. dytiscoides (AMNH); 2??7, Morcatta (OUM); 14, Adelotopus sp. Taraittah K 12399 (AMS);

12, Petersham 128233 (AMS); 1 (2), E. Plains X1.93 (ANIC); 1%, T...don 16.X.86 (SAMA); 13, Type, Adelotopus

Dytiscides Newman (BMNH); 229, K 12228, det. gyrinoides (AMS); 19, Adelotopus dytiscoides Rev. Zool. 1853,

Adelotopus fortnumi (OUM); 18, 22%, Bridwell Collection, det. dytiscoides (USNM); 12, 1083, det. dytiscoides

(MNHB); 2? 9, 41066, det. dytiscoides (MNHB); 15, Type Adelotopus Dytiscoides Newm. (BMNH); 15, 17, Sharp

Coll., det. dytiscoides (BMNH); 234, 34, Adelotopus gyrinoides Hope (UQIC); 18, A. gyrinoides Hope (ZSM);

222, det. dytiscoides (MNHB); 14, Ad. dytiscoides 20.X1.92 (ANIC); 15, 12, 125, 9, 16 Howitt Colln., det. dytiscoides

(NMV); 1?, Janson Acgq. 1884 (MNHN); 234, 17, Collection E. Rousseau, Adelotopus dytiscoides Newm. det.

E. Rousseau (IRSNB); 13, Collection E. Rousseau, Adelotopus gyrinoides Hope det. E. Rousseau (IRSNB); 15, 61,

Soc. Ent. Belg. Coll. PUTZEYS, Adelotopus dytiscoides Newm. det. J. Putzeys (IRSNB); 12, 797, Silphomorpha

dytiscoides Newm. det..., 797 Silphomorpha dytiscoides Chaud. (IRSNB); 15, Lea Dup. (SAMA); 12, Moc (?) (OUM);

13,12, W (CSB); 1? (UQIC); 1? (ANIC); 1? (SAMA); 14,4? 2 (OUM); 1%, Coll. Schwarzer, det. gyrinoides (SMF);

1?, ac. 23246, mastersii Mcl. (AMNH); 18, 22%, L 540 (NMV); 28, 1322, 1-7, 20-15, 17, 18, Howitt Colln., det.

dytiscoides (NMV); 458,7? , Gunbower X.04 J. A. Kershaw (NMV); 322, 1961 (NMV); 2??, Nat. Mus. Victoria,

From Mr. Duboulay 2.1V.11 (NMV); 1%, C. French’s Coll. (NMV); 434,272, Ex Musaeo Chaudoir, det. dytiscoides

Newman (MNHN); 12, Ex Musaeo Mniszech, det. dytiscoides Newman (MNHN); 14, 1? (NMV).

Adelotopus ulrichi, spec. nov.

Figs 101, 290, 439, 440, 604

Types. Holotype: d, Australien, Qld 27, Rolf Ck., 134km n. Dingo, Fitzroy Dev. Rd., 12.11.1990, M. Baehr

(ANIC). - Paratypes: 834, 4?9, same data (CBM, NMV, ZSM); 18, Brisbane H. Hacker 13.12.11, Adelotopus

dytiscoides Newm. det. Sloane (QMB); 1%, Ipswich S.Q. 2/67 JK, M.54, J. G. Brooks Bequest 1976 (ANIC);

13, Greenbank Qld. 13.X11.62 G. Monteith (UQIC); 13, Maryborough Queensland (SAMA); 17, Maryborough

Queensland E. W. Fischer (SAMA); 1%, Queensland, Rockhampton XI 1986, J. Sedlacek Collector (CSB);

484,19, Australia: Qld Rockhampton 26-27.X1.1967, J. & M. Sedlacek Collectors BISHOP (BMH); 1%, Australia,

Qld, Mackenzie River, 29.1.1968, leg. G. Hangay (HNMB); 19, Edungalba, CQ 9.1.1964, E. E. Adams, 1580,

Adelotopus dytiscides Newman [Series det. by A. Walford-Huggins] (CMP-WHC); 14,N. Holl P. Denison Dämel

(MNHB); 14, Buckley (?) Qld 6.12.26, Australia, G. E. Clarke Collection 1957 (BMNH); 14, N. Queensland,

Mastersii Macl. sec. description, Ex Museo H. W. Bates 1892 (MNHN); 1?, Australie Queensland Salle,

Ex Musaeo Chaudoir (MNHN); 12, N. Holl. Q’land Janson Acq. 1884 (MNHN); 15, Port Darwin, N. Territory

(MMS); 12, Barrimore 16.-1-44, Brock & Sutton, E. Sutton Coll. 1964 (QMB); 148, K 12005 A. politus (AMS).

Figs 101a-l. Adelotopus ulrichi, spec. nov. Details of head and genitalia. For legends see fig. 100.

Diagnosis. Large, moderately wide, moderately depressed, uniformly black species with distinctly

concave base of pronotum and rasp-like and horseshoe-shaped punctures marking the elytral striae.

Distinguished from relative A. latior, spec. nov. by pronotum narrower and more convex, elytra longer

and duller, and elytral striae visible at most in apical half.

Description

Measurements. Length: 7.5-9.1 mm. Ratios. Width/length of pronotum: 1.78-1.86; width base/

apex of pronotum: 1.69-1.85; width pronotum/ head: 1.77-1.93; length/ width of elytra: 1.36-1.43; length

elytra/pronotum: 2.51-2.63.

Colour. Black, lateral borders of pronotum and elytra sometimes slightly reddish-piceous translu-

cent. Lower surface, antennae, mouthparts, and legs dark reddish-piceous, lower surface of head

almost black.

Head (Figs 101a-d, 290). Short and wide, rather depressed. Anterior border feebly convex, lateral

border rather angular. Clypeal suture shallow, in middle usually absent, semicircular. Labrum

comparatively large, apex slightly concave. Antennal groove laterally strongly bordered, latero-

posteriorly with convex, though barely carinate area. Mental tooth triangular, rather wide, at apex

slightly rounded off. Wings of mentum wide, near apex strongly rounded. Glossa wide, tongue-like,

apically rounded, ventrally with feeble keel, at border with c. 8 elongate setae and additional dense

pilosity on upper and lower surface and along border. Terminal palpomere of maxillary palpus

narrow, apex feebly widened, barely securiform. Terminal palpomere of labial palpus large, very

widened, wider than long, markedly securiform. Antenna moderately elongate, Sth-9th antennomeres

slightly wider than long. Microreticulation dense and distinct, puncturation rather dense, though more

or less difficult to detect, surface with a shallow sulcus medially of eyes and usually with some fine

longitudinal strioles on frons, impilose, markedly dull. Ventrolaterally of eyes with a row of short

setae. Suborbital field impunctate and asetose. Both palpi sparsely setose, gula almost smooth.

Pronotum (Fig. 290). Rather wide, moderately convex, base clearly less than 2 x as wide as apex.

Apical angles produced, acute, attaining almost middle of eyes. Apex deeply, rectangularly excised,

slightly convex in excision, not bordered. Sides strongly and evenly curved, widest near basal angles,

barely or not at all bordered, but anteriorly more or less widely explanate. Basal angles rectangular or

slightly obtuse, at apex barely rounded off, base distinctly concave, though in middle feebly produced,

not bordered. Microreticulation dense and distinct, very fine, almost isodiametric, puncturation fine,

difficult to detect, surface with some fine, irregular strioles, impilose, dull.

Elytra (Figs 290, 439, 440). Moderately elongate and convex, in basal two fifth rather parallel, then

gently narrowed. Apex slightly oblique, truncature feebly concave, dehiscent at suture. Shoulders

obtuse, basal margin oblique, with a row of many short setae. Basal border line complete, well

developed, meeting at suture. Marginal channel rather wide, slightly widened in basal fourth,

completely visible from above. Lateral border asetose or with but 1-2 short setae just behind shoulders.

Series of umbilical pores consisting of 6 pores behind shoulder and 1 pore or, rarely, 2 pores at apical

third. Setae short. Striae basally reduced, indicated only by irregular rows of extremely fine strioles,

but sutural stria posterioly slightly impresed, and internal striae visible in apical third as rows of corse,

rasp-like punctures, becoming even larger towards apex. These punctures laterally rather confused.

Microreticulation dense and distinct, basally no puncturation visible. Surface impilose, markedly dull.

Lower surface. Prosternal process rather elongate, straight, moderately depressed, apex rectangu-

lar, shortly setose. Metepisternum elongate, c. 2x as long as wide, posteriorly constricted and

hollowed. Abdominal sterna with 3-6 setae each side, sternum VI in both sexes with many (c. 14-24),

rather short setae at or near apical border. Whole lower surface rather densely punctate and pilose.

Legs. Elongate, Ist tarsomere of protarsus not wider than long, tibial groove of profemur

comparatively shallow, anterior plate only at apex distinctly overlapping the groove, posteriorly

border of groove not sharp. Metatibia rather narrow, >5 x as long as wide, 1st tarsomere of metatarsus

>2 x as long as wide. d protarsus barely widened.

d genitalia (Figs 101e-k). Genital ring rather asymmetric, laterally slightly convex. Sternum VII

apically deeply excised, basally slightly bisinuateor straight, laterally short. Aedeagus moderately

elongate, wide, moderately depressed, rather symmetric, lower surface almost straight. Lateral border

wide throughout. Apex wide, rounded off. Internal sac rather simple. Right paramere elongate, with

obtuse apex. Left paramere much larger, convex, with widely rounded or somewhat obtuse apex.

? genitalia (Fig. 1011). Stylomere 2 triangular, apically rather narrow, with 2-3 subapical setae.

Lateral plate rather elongate, with 2-3 apical setae.

Variation. Only some variation noted in relative shape of pronotum the lateral explanations of

which may be rather narrow, distinctness of puncturation of head and pronotum, and degree of

puncturation of elytra. Otherwise a rather homogeneous species.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Not specified. A series collected by me under bark of River Gum at the border of a dry creek

in open eucalypt woodland. So far collected from November to February only.

Distribution (Fig. 604). Eastern Queensland from Brisbane to about Townsville, perhaps also north-

ern part of Northern Teritory.

Material examined (34). Only the type series.

Etymology. Named in honour of my little son, who helped me collecting mainly by not walking too far into tall

grass and by not getting a sunstroke in extremely hot eastern Queensland.

Adelotopus latior, spec. nov.

Figs 102, 291, 441, 442, 604

Types. Holotype: , 17.585, 144.51E, 43 km SW of Mt. Garnet, Qld. 8 Dec.1985 J. Balderson (ANIC). — Paratypes:

13, Australien, Qld., 3 km w. Atherton 29.12.1981, M. Baehr (CBM); 14, Mareeba Dodd 08 (ANIC); 14, Mareeba

1/49, gyrinoides Hope 1584, J. G. Brooks Bequest 1976 (ANIC); 13, Mareeba NQ 2/53 GB, Adelotopus hydrobioides

Westw. det. B. P. Moore '69, J. G. Brooks Bequest 1976 (ANIC); 15 (defect), Chillagoe, Exp. Dodd, 3.08 (ANIC).

Diagnosis. Large, wide, rather depressed, uniformly black species with distinctly concave base of

pronotum, and rasp-like punctures marking the elytral striae. Distinguished from most closely related

species A. ulrichi, spec. nov. by pronotum wider and more depressed, elytra shorter and less dull, and

elytral striae visible at least from basal third.

Figs 102a-k. Adelotopus latior, spec. nov. Details of head and d genitalia. For legends see fig. 100.

Description

Measurements. Length: 7.0-8.2 mm. Ratios. Width/length of pronotum: 1.89-2.05; width base/

apex of pronotum: 1.83-1.90; width pronotum/ head: 1.90-1.99; length/ width of elytra: 1.23-1.25; length

elytra/pronotum: 2.27-2.55.

Colour. Black, lateral borders of pronotum and elytra faintly reddish-piceous translucent. Lower

surface, antennae, mouthparts, and legs dark reddish-piceous, lower surface of head dark piceous.

Head (Figs 102a-d, 291). Short and wide, rather depressed. Anterior border feebly convex, lateral

border rather angular. Clypeal suture shallow, in middle more or less reduced, semicircular. Labrum

comparatively large, apex slightly concave. Antennal groove laterally strongly bordered, latero-

posteriorly with convex, though barely carinate area. Mental tooth triangular, rather wide, at apex

slightly rounded off. Wings of mentum wide, near apex strongly rounded. Glossa wide, tongue-like,

apically rounded, ventrally with feeble keel, at border with c. 8 elongate setae and additional pilosity

on upper and lower surface and along border. Terminal palpomere of maxillary palpus rather narrow,

apex feebly widened, barely securiform. Terminal palpomere of labial palpus large, very widened,

wider than long, markedly securiform. Antenna moderately elongate, 8th-9th antennomeres distinctly

wider than long. Microreticulation dense and distinct, puncturation rather dense, fairly well seen,

surface with a shallow sulcus medially of eyes and usually with some fine longitudinal strioles on

frons, impilose, dull. Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and

asetose. Both palpi sparsely setose, gula almost smooth.

Pronotum (Fig. 291). Wide, moderately convex, base almost 2 x as wide as apex. Apical angles

produced, acute, attaining almost middle of eyes. Apex deeply, rectangularly excised, slightly convex

in excision, not bordered. Sides strongly and evenly curved, widest near basal angles, barely or not at

all bordered, but anteriorly rather widely explanate. Basal angles rectangular or slightly obtuse, at apex

barely rounded off, base perceptibly concave, sometimes in middle feebly produced, not bordered.

Microreticulation dense and distinct, very fine, almost isodiametric, puncturation minute, though

easily detected under high magnification, surface with some fine, irregular strioles, impilose, moder-

ately dull.

Elytra (Figs 291, 441, 442). Rather short and wide, moderately convex, in basal two fifth rather

parallel, but in middle faintly widened, then gently narrowed. Apex slightly oblique, truncature feebly

concave, dehiscent at suture. Shoulders obtuse, basal margin oblique, with a row of many short setae.

Marginal channel rather wide, slightly widened in basal fourth, completely visible from above. Basal

border complete, well developed, meeting at suture. Lateral border asetose or with but 1-2 short setae

just behind shoulders. Series of umbilical pores consisting of 6 pores behind shoulder and 1 pore or,

je s)

rarely, 2 pores at apical third. Setae short. Striae near base reduced, indicated only by irregular rows

of extremely fine strioles, sutural stria posterioly slightly impressed. All striae visible from at least

basal third as rows of corse, rasp-like punctures, becoming larger towards apex. These punctures

laterally rather confused. Microreticulation dense and very fine, distinct, basally no puncturation

visible. Surface impilose, but moderately dull.

Lower surface. Prosternal process rather elongate, straight, moderately depressed, apex rectangu-

lar, shortly setose. Metepisternum elongate, slightly <2 x as long as wide, posteriorly constricted and

hollowed. Abdominal sterna with 3-6 setae each side, sternum VI with many (c. 14-24) short setae at

or near apical border. Lower surface rather sparsely punctate and pilose.

Legs. Elongate, Ist tarsomere of protarsus not wider than long, tibial groove of profemur

comparatively shallow, anterior plate only at apex distinctly overlapping the groove, posteriorly

border of groove not sharp. Metatibia rather narrow, >5 x as long as wide, 1st tarsomere of metatarsus

>2 x as long as wide. d protarsus barely widened.

d genitalia (Figs 102e-k). Genital ring rather asymmetric, laterally slightly convex. Sternum VII

apically deeply excised, basally slightly bisinuate or straight, laterally short. Aedeagus moderately

elongate, wide, moderately depressed, rather symmetric, lower surface almost straight. Lateral border

wide throughout. Apex wide, rounded off. Internal sac rather simple. Right paramere elongate, with

obtuse apex. Left paramere much larger, convex, with widely rounded apex.

? genitalia. Unknown.

Variation. Little variation noted due to limited material.

Vivipary. Not confirmed due to lack of females.

Habits. Generally not specified. One specimen captured by me under bark of gum-type eucalypt. So

far collected from December to March.

Distribution (Fig. 604). Western slope of Atherton Tableland and western adjacent country.

Material examined (6). Only the type series.

Etymology. The name refers to the shorter and wider shape in comparison to the related A. ulrichi.

Adelotopus apicalis Macleay, 1864

Figs 103, 292, 443, 444, 605, 655

Adelotopus apicalis Macleay, 1864, p.113; Blackburn 1901a, p. 19; 1901b, p. 113; Notman 1925, p. 7, 10, 28; Csiki

1933, p. 1634; Darlington 1968, p. 241; Moore et al. 1987, p. 49.

Types. Lectotype (by present designation): d, Pt. Denison, Syntype, Adelotopus apicalis Macl. Port Denison

(ANIC-MMS). - Paralectotypes: 2??, same data (ANIC-MMS); 1%, Pt. Denison, Syntype (ANIC-MMS).

Type locality. “Pt. Denison”, Queensland.

Diagnosis. Large, wide, rather depressed, black species with reddish elytral apex, bright red apex of

abdomen, and usually perceptibly concave base of pronotum. Further distinguished from relative

species by elytra glossy, elytral striae punctate and visible from apical half, but punctures barely rasp-

like.

Description

Measurements. Length: 6.1-8.5 mm. Ratios. Width/length of pronotum: 1.98-2.11; width base/

apex of pronotum: 1.80-1.88; width pronotum/head: 1.84-1.90; length/ width of elytra: 1.22-1.34; length

elytra/pronotum: 2.47-2.78.

Colour. Glossy black, apex of elytra usually more or less distinctly reddish, though sometimes not

even perceptibly lighter than rest of elytra. Lower surface of head and thorax dark piceous to almost

black, of abdomen reddish-piceous, posteriorly usually red. Mouthparts, antennae, and legs reddish-

piceous or even red.

Head (Figs 103a-d, 292). Short and wide, rather depressed. Anterior border feebly convex, lateral

border feebly obtuse. Clypeal suture shallow, in middle more or less reduced, semicircular. Labrum

comparatively wide, apex gently concave. Antennal groove laterally strongly bordered, latero-

Figs 103a-l. Adelotopus apicalis Macleay. Details of head and genitalia. For legends see fig. 100.

posteriorly with almost depressed area. Mental tooth triangular, short, apex acute. Wings of mentum

wide, near apex strongly rounded. Glossa wide, tongue-like, apically widely rounded, ventrally with

feeble keel, at border with c. 8 elongate setae and additional pilosity on upper and lower surface and

along border. Terminal palpomere of maxillary palpus moderately narrow, apex slightly widened,

weakly securiform. Terminal palpomere of labial palpus large, very widened, wider than long,

markedly securiform. Antenna rather short, 8th-9th antennomeres markedly wider than long. Micro-

reticulation very fine, dense, puncturation minute, difficult to see, surface with a shallow sulcus

medially of eyes, impilose, moderately dull. Ventrolaterally of eyes with a row of short setae.

Suborbital field impunctate and asetose. Both palpi sparsely setose, gula almost impilose.

Pronotum (Fig. 292). Wide, moderately convex, base almost 2 x as wide as apex. Apical angles

produced, rather acute, attaining posterior third of eyes. Apex deeply, rectangularly excised, slightly

convex in excision, not bordered. Sides strongly and evenly curved, widest at or near basal angles,

barely or not at all bordered, usually not much explanate. Basal angles rectangular or slightly obtuse,

at apex barely rounded off, base usually perceptibly concave, sometimes in middle feebly produced,

not bordered. Microreticulation very fine, almost isodiametric, puncturation minute, more or less

easily detected under high magnification, surface with some fine, irregular strioles, impilose, rather

glossy.

Elytra (Figs 292, 443, 444). Rather short and wide, moderately convex, in basal two fifth almost

parallel, then gently narrowed. Apex slightly oblique, apical angles widely rounded, truncature feebly

concave, dehiscent at suture. Shoulders obtuse, basal margin oblique, with a row of many short setae.

Marginal channel narrow, completely visible from above. Basal border complete, well developed,

meeting at suture. Lateral border asetose. Series of umbilical pores consisting of 6 (rarely unilaterally 7)

pores behind shoulder and 1 pore behind middle or at apical third. Setae very short. Striae in basal half

absent, only vaguely indicated by irregular rows of extremely fine strioles, sutural stria posterioly

slightly impressed, suture raised. All striae visible from slightly behind middle as rows of basally fine,

not or but faintly rasp-like punctures, becoming larger towards apex. In basal half intervals also with

scattered, extremely minute punctures. Microreticulation very fine, superficial, much less distinct than

on fore body, consisting of slightly transverse meshes. Surface impilose, glossy.

Lower surface. Prosternal process moderately elongate, wide, straight, rather depressed, apex

widely rounded, shortly setose. Metepisternum moderately elongate, slightly <2 x as long as wide,

apically constricted and hollowed. Abdominal sterna with 1-2 setae each side, sternum Vl in both sexes

moderately punctate and shortly setose, but without more distinct setae at apical border. Lower

surface rather sparsely punctate and pilose.

Legs. Rather elongate, Ist tarsomere of protarsus not wider than long, tibial groove of profemur

moderately deep, anterior plate in anterior half distinctly overlapping the groove, posteriorly border

of groove not sharp. Metatibia rather narrow, c. 5 x as long as wide, 1st tarsomere of metatarsus c. 2 x

as long as wide. d protarsus barely widened.

d genitalia (Figs 103e-k). Genital ring rather asymmetric, laterally slightly convex. Sternum VII

apically deeply excised, basally slightly bisinuate or straight, laterally short. Aedeagus moderately

elongate, wide, moderately depressed, rather symmetric, lower surface gently convex. Lateral border

only in apical half wide. Apex wide, rounded off. Internal sac rather simple. Right paramere

moderately elongate, with obtuse apex. Left paramere much larger, convex, with widely rounded

apex.

? genitalia (Fig. 1031). Stylomere 2 widely triangular, apically rather wide, with 2-3 subapical

setae. Lateral plate rather elongate, with 3-4 apical setae.

Variation. Some variation noted in relative shape of pronotum and elytra, in degree of reddish

colour on apex of elytra, and in degree of elytral striation. There are some specimens, however, that

are barely reddish on elytra and abdomen, and some specimens posessing very wide pronota with

fairly marked lateral sulcus and barely sinuate basal margin that come all from extreme northern Cape

York Peninsula and represent perhaps a geographical variation that is, however, from my view not of

much taxonomical value. There is a single specimen having the head and pronotum extensively

microreticulate and shagreened by many coarse wrinkles. Because it is in other ways similar to other

specimens, I do not believe that it represents more than an abnormal variant.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Little specified. Specimens were collected by me under bark of river gums and other gum-type

eucalypts. The large series from Granite Gorge near Mareeba was found in immediate vicinity of a nest

of Camponotus dorycus (Sm.). Specimens collected by P. Meyer were found “under bark of bloodwood-

type eucalypt” and “under bark of box-type eucalypt”. Dated specimens were captured from

September to February and in May and July, though comparatively few specimens are dated, and,

apart from those recently collected by me, most are from captures made by F. P. Dodd in October 1902.

Distribution (Figs 605, 655). Northeastern Queensland from about Rockhampton to tip of Cape York

Peninsula, apparently also New Guinea. An old record from Tasmania is highly doubtful.

Material examined (89). Tas: 13, 294, Tasmania, A. Simson, Adelotopus sp., Adelotopus hydrobioides Westw. det...

hydrobioides Westw. suivant Simson (IRSNB). — Old: 1?, Rockhampton, Coll. C. Felsche, det. gyrinoides GMID);

17, Dipperu M. E. Q., X1.1971, B. Baldwin (QMB); 12, Port Bowen 75.12, Adelotopus hydroporoides Westw. Id. By

T. G, Sloane (BMNH); 19, Bowen A. Simson 294 4127, Silphomorpha hydrobioides By Simson’s number (SAMA);

12, Bowen, Janson Acg. 1884 (MNHN); 15, 27%, Pt. Denison, Syntype, Adelotopus apicalis Macl. Port Denison

(ANIC-MMS); 1%, Pt. Denison, Syntype (ANIC-MMS); 1%, Adelotopus apicalis M. L. Port Denison, 48, Howitt

es (NMV); 12, P'. Denison, Ex Musaeo Mniszech, apicalis M'Leay, det. haemorrhoidalis Westwood (MNHN);

7, Port Denison Coll. Castelnau, apicalis M. L.j".Port Denison, apicalis ? Mael. (MCSN); 234, Port Denison Coll.

en det. apicalis (MCSN); 272%, Townsville 14.X.02 F. P. Dodd (ANIC); 2 (?), Townsville X.02 F. P. Dodd,

Adelotopus hydroporoides Westw. = apicalis Macl. Id. by T. G. Sloane (ANIC); 15, 1, Townsville X.02 F. P. Dodd,

Griff. 2562, A. apicalis Macl. comp. type 6.VIl.20, Adelotopus hydrobioides West. Id. by T. G. Sloane (SAMA);

236, Townsville, G. Bryant Coll. 1919, det. hydrobioides (BMNH); 2358, Townsville X.02 F. P. Dodd, Ex Coll.

T. Sloane, Adelotopus hydrobioides West. ? Id. by T. G. Sloane, H. E. Andrewes Coll. 1945 (BMNH); 1%, Townsville

14.X.02 F. P. Dodd, G. Bryant Coll. 1919, det. hydrobioides (BMNH); 12, Townsville X.02 F. P. Dodd, det.

hydrobioides (BMNH); 14, Townsville 29.1X.02 F. P. Dodd, C. Bryant Coll. 1919, det. hydroporoides (BMNH);

19, 7286, Townsville 3.X.02 F. P. Dodd (SAMA); 14, Townsville Oct. 02 F. P. Dodd, Adelotopus hydrobioides West.

= apicalis Macl. Id. by T. G. Sloane (SAMA); 18, 12, Townsville (BMNH); 286, Tville 2562, Townsville 2.X1.02

F. P. Dodd, Griffith Coll. Id. by A. M. Lea (SAMA); 19, Hervey Range Rd., 5.3 km W Bohle Rd., W Townsville

29.V. EA P. A. Meyer coll. (CBM); 12, Ayr, N. Q., received from W. du Boulay, REN acl.(NMV);

338, ', Qld 93/67, Einasleigh R. b. Carpentaria Downs, 12.-13.6.1993, M. Baehr (CBM); 1%, Qld 93/64,

ee R. 2km e. Einasleigh, 11.-12.6.1993, M. Baehr (CBM); 1%, Cairns dist. F. P. a Adelotopus

hydrobioides West. = apicalis Macl. Id. by T. G. Sloane (SAMA); 19, Mulgrave Res. 405. Adelotopus hydrobioides

Figs 104a-d, 1. Adelotopus zborowskii, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Westw. Id. by T. G. Sloane Coll. Hacker (DEIB); 19, 17, Kuranda, Griffith Coll. Id. by A. M. Lea, Silphomorpha,

2562 (SAMA); 2?%, Kuranda, Queensland Dodd & Griff. Adelotopus hydrobioides West. Id. by T. G. Sloane

(SAMA); 1?, Mareeba, XI1.52, J. G. Brooks, Adel. gyrinoides Hope (MCZ); 1559, 32%, Qld 93/76, Granite Gorge,

15km wsw. Mareeba, 15.6.1993, M. Baehr (CBM, ZSM); 19, Qld 93/51, Emu Ck., 5km e. Petford, 8.6.1993,

M. Baehr (CBM); 12, 40 Mile Scrub area, SW Ravenshoe, 14.V.1987, P. A. Meyer coll. (CBM); 12, N. of Mareeba

11.58 Darlingtons (MCZ); 18, 18km N. Mareeba, 2.1.1989, H. & A. Howden (Ottawa); 19, 17, Qld 93/4,

Mt. Molloy, 22.5.1993, M. Baehr (CBM); 15, Station Ck. NO, 10 m S. Mt. Carbine, 20.X11.1971, A. &M. Walford-

Huggins, 6128 (CMP-WHC); 1?, Einasleigh River, 33km w. Mt. Surprise, 4.1996, leg. A. Floren (CBM);

222, Toolkoor N. Queensland (NMHB); 1%, Endeavour R., C. French’s Coll., det. A. apicalis (NMV); 17, Somerset,

C. T. Mc Namara (SAMA); 13, Australia Somerset 1.75 L. M. D’Albertis, apicalis Macl. det. Gestro (MCSN);

12, 22/1 Bamaga N.Q. Jan. 1984, ]. H. Sedlacek (CSB); 1?, Cape York, Lea has not, Adelotopus Cape York (QMB);

2? 2, Cape York (MMS); 15, E. Weiske N. Queensland, 15054, det. gyrinoides (SMTD); 18, 17, Simson, Fry Coll.

1905, det. hydrobioides (BMNH). - NG: 1, Redscar Bay, Brit. N. G. (MNHN); 1%, K. Plat. N. N. G. 16.VII.94,

Should be A. hygrobioides from des 5.X11.02 Return (ANIC). - ?: 15, D. Sharp Coll. 1932 (BMNH).

Adelotopus zborowskii, spec. nov.

Figs 104, 293, 445, 605

Types. Holotype: ?, 15.395 144.31E Split Rock QLD 18 Feb-25 Apr 1993 Malaise Trap P. Zborowski (QMB). -

Paratypes: 17, 15.395 144.31E Split Rock QLD 16 Sep-19 Oct 1993 Flight Intercept Trap P. Zborowski & D. Rentz

(DPIM); 1?, Hann R. via Laura, n. Qld Dec. 93-Jan. 94 P. Zborowski Malaise Trap (CBM).

Diagnosis. Medium sized, fairly wide, moderately depressed, black species with more or less

distinctly reddish translucent lateral margins of pronotum and elytra. Further distinguished from

relative species by lesser size, barely marked and only in apical half finely punctate elytral striae, fine

and not silky microreticulation, and absence of elytral punctures.

Description

Measurements. Length: 5.55-6.40 mm. Ratios. Width/length of pronotum: 1.76-1.80; width base/

apex of pronotum: 1.60-1.67; width pronotum /head: 1.66-1.75; length/ width of elytra: 1.40-1.42; length

elytra/pronotum: 2.51-2.54.

Colour. Black, lateral margins of pronotum and elytra more or less distinctly reddish translucent.

Lower surface of head and thorax black, of abdomen reddish-piceous, posteriorly lighter. Mouthparts,

antennae, and femora reddish-piceous, tibiae and tarsi blackish.

Head (Figs 104a-d, 293). Short and wide, fairly convex. Anterior border feebly convex, lateral angle

obtuse, slightly projecting. Clypeal suture shallow, indistinct, widely interrupted in middle, of

completely absent. Labrum comparatively wide, apex gently concave. Antennal groove laterally barely

bordered, latero-posteriorly with convex, slightly carinate area. Mental tooth triangular, short, apex

acute. Wings of mentum wide, near apex strongly rounded. Glossa wide, tongue-like, apically widely

rounded, ventrally with distinct keel, at border with c. 8 elongate setae and additional pilosity on

upper and lower surface and along border. Terminal palpomere of maxillary palpus elongate,

moderately widened, slightly securiform. Terminal palpomere of labial palpus very wide, markedly

securiform. Antenna short, 8th-9th antennomeres >2 x as wide as long. Microreticulation fine, dense,

slightly superficial, puncturation almost invisible, surface with a shallow sulcus medially of eyes,

impilose, moderately glossy. Ventrolaterally of eyes with a row of short setae. Suborbital field

impunctate and asetose. Both palpi rather sparsely setose, gula apparently asetose.

Pronotum (Fig. 293). Moderately wide, moderately convex, base <1.7 x as wide as apex. Apical

angles moderately produced, rather acute, attaining posterior third to half of eyes. Apex deeply,

rectangularly excised, slightly convex in excision, not bordered. Sides evenly curved, widest near basal

angles, very finely, inconspicuously bordered, moderately explanate, but less so than the other species

of this group. Basal angles almost rectangular, at apex barely rounded off, base faintly concave or

almost straight, not bordered. Microreticulation fine and somewhat superficial, almost isodiametric,

puncturation extremely fine, almost invisible, surface with some fine, irregular strioles, impilose,

moderately glossy.

Elytra (Figs 293, 445). Comparatively elongate, moderately convex, in basal three fourth almost

parallel, then gently narrowed. Apex slightly oblique, apical angles widely rounded, truncature feebly

concave. Shoulders obtuse, basal margin oblique, without setae behind shoulders. Marginal channel

narrow, completely visible from above. Basal border almost complete, well developed, ending close to

suture. Lateral border asetose. Series of umbilical pores consisting of 6 pores behind shoulder and

1 pore in middle. Setae very short. Striae in apical half more or less distinctly indicated as rows of fine

punctures, basally absent, sutural stria invisible, suture apically faintly raised. Puncturation almost

invisible. Microreticulation fine, somewhat superficial, isodiametric, slightly coarser than on fore

body. Surface impilose, moderately glossy.

Lower surface. Prosternal process moderately elongate, rather wide, straight, rather depressed,

apex straight, shortly setose. Metepisternum moderately elongate, c. 1.5 x as long as wide, posteriorly

constricted and deeply hollowed. Abdominal sterna with 1 seta each side, sternum VI without longer

setae at apical border. Lower surface rather sparsely punctate and pilose.

Legs. Rather elongate, 1st tarsomere of protarsus not wider than long, tibial groove of profemur

moderately deep, anterior plate only in apical half distinctly overlapping the groove, posteriorly

border of groove fairly sharp. Metatibia rather narrow, c. 5x as long as wide, Ist tarsomere of

metatarsus c. 2x as long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia. (Fig. 1041). Stylomere rather wide, apically slightly narrowed, with 2-3 apical setae.

Lateral plate rather elongate, with 2-3 apical setae.

Variation. Some variation noted in size and relative with of pronotum.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unknown, all specimens collected in “Malaise trap” or “flight intercept trap”. Thus far

collected during the periods of February-April, September-October, and December-January.

Distribution (Fig. 605). Lower Cape York Peninsula, northeastern Queensland.

Material examined (3). Only the type series.

Etymology. Named in honour of the collector.

Adelotopus sericeus, spec nov.

Figs 105, 294, 446, 605

Types. Holotype: d&, WAM Goldfields Survey, R. A. How MRR5 7/1979, Mallee 32°44'30"S, 120°01'30"E,

Mc Dermid Rock, PIT/F, Adelotopus sp. det. T. A. Weir 1981 (ANIC). - Paratype: 15, same data (WAM 94/847)

Diagnosis. Rather large, moderately wide, rather depressed, uniformly black species with dull,

markedly silky surface. Further distinguished from relative species by lesser size, almost invisible and

impunctate elytral striae, almost invisible elytral punctures, rather asymmetric d genital ring, and

narrow and evenly curved aedeagus with wide apex.

Figs 105a-k. Adelotopus sericeus, spec. nov. Details of head and d genitalia. For legends see fig. 100.

Description

Measurements (paratype somewhat damaged, not measurable). Length: c. 6.6-c. 7.0 mm. Ratios.

Width/length of pronotum: c. 1.63; width base/apex of pronotum: 1.70; width pronotum/head: 1.84;

length/width of elytra: c. 1.45; length elytra/pronotum: c. 2.43.

Colour. Dull black. Lower surface of head and thorax black, of abdomen reddish-piceous,

posteriorly lighter. Mouthparts, antennae, and tibiae and tarsi reddish-piceous, femora blackish, at

anterior border piceous.

Head (Figs 105a-d, 294). Short and wide, rather depressed. Anterior border feebly convex, lateral

angle feebly obtuse. Clypeal suture shallow, complete, semicircular. Labrum comparatively wide, apex

gently concave. Antennal groove laterally barely bordered, latero-posteriorly with convex, not carinate

area. Mental tooth triangular, short, apex acute. Wings of mentum wide, near apex strongly rounded.

Glossa wide, tongue-like, apically widely rounded, ventrally with distinct keel, at border with

c. 8elongate setae and additional pilosity on upper and lower surface and along border. Terminal

palpomere of maxillary palpus elongate, feebly widened, not securiform. Terminal palpomere of labial

palpus moderately widened, longer than wide, feebly securiform. Antenna short, 8th-9th antennomer-

es c. 1.5 x as wide as long. Microreticulation fine, dense, puncturation invisible, surface with a shallow

sulcus medially of eyes, impilose, fairly silky. Ventrolaterally of eyes with a row of short setae.

Suborbital field almost impunctate and asetose. Both palpi rather sparsely setose, gula sparsely

punctate and setose.

Pronotum (Fig. 294). Moderately wide, moderately convex, base c. 1.7 x as wide as apex. Apical

angles produced, rather acute, attaining posterior third to half of eyes. Apex deeply, rectangularly

excised, slightly convex in excision, not bordered. Sides strongly and evenly curved, widest near basal

angles, very finely, inconspicuously bordered, moderately explanate. Basal angles almost rectangular,

slightly produced backwards, at apex barely rounded off, base laterally faintly concave, in middle

feebly produced, not bordered. Microreticulation fine though distinct, almost isodiametric, punctur-

ation almost invisible, surface with some fine, irregular strioles, impilose, rather silky.

Elytra (Figs 294, 446). Moderately elongate and wide, moderately convex, in basal three fourth

almost parallel, then gently narrowed. Apex slightly oblique, apical angles widely rounded, truncature

feebly concave. Shoulders obtuse, basal margin oblique, with 2-3 short setae behind shoulders.

Marginal channel narrow, completely visible from above. Basal border complete, well developed,

meeting at suture. Lateral border asetose. Series of umbilical pores consisting of 5 or 6 pores behind

shoulder and 1 pore in middle. Setae very short. Striae in basal two thirds very vaguely indicated as

extremely fine, wavy lines, apically absent, sutural stria invisible, suture apically barely raised.

Puncturation becoming apically denser and more distinct, though generally very difficult to detect

within the strong microreticulation. Microreticulation rather fine, though very distinct, isodiametric,

coarser than on fore body, becoming coarser towards apex. Surface impilose, markedly silky.

Lower surface. Prosternal process moderately elongate, rather wide, straight, rather depressed,

apex straight, shortly setose. Metepisternum elongate, >2 x as long as wide, posteriorly constricted and

hollowed. Abdominal sterna with 1-2 setae each side, sternum VI with c. 12 rather elongate setae at

apical border, and some short setae near border and on surface. Lower surface rather sparsely punctate

and pilose.

Legs. Rather elongate, 1st tarsomere of protarsus not wider than long, tibial groove of profemur

comparatively shallow, anterior plate only at apex distinctly overlapping the groove, posteriorly

border of groove not sharp. Metatibia rather narrow, >5 x as long as wide, 1st tarsomere of metatarsus

>2 x as long as wide. d protarsus barely widened.

d genitalia (Figs 105e-k). Genital ring narrow, rather asymmetric. Sternum VII moderately wide,

apically deeply excised, basally slightly convex. Aedeagus rather elongate, narrow, depressed,

symmetric, strongly and evenly curved. In posterior part of upper surface laterally with sharp edge.

Apex wide, rounded off. Orifice elongate, internal sac rather simple. Both parameres rather narrow

and elongate, though left larger than right.

? genitalia. Unknown.

Variation. Very little variation noted.

Vivipary. Not confirmed due to lack of females.

Habits. Unknown, so far collected in July.

Distribution (Fig. 605). Interior of southwestern Australia. Known only from type locality.

Material examined (2). Only the holotype and one paratype.

Etymology. The name refers to the highly sericeous lustre of the surface.

Adelotopus howdenorum, spec. nov.

Figs 106, 295, 447, 605

Types. Holotype: d, Australia: WA, Mt. Madden, 23-24.1X.1981, H. & A. Howden (ANIC). - Paratype: 17,

Coolgardie hd.(?), 29 Aug 1958, Le Souef (NMV).

Diagnosis. Rather large, wide, rather depressed, uniformly black species with dull, though somewhat

silky surface. Further distinguished from relative species by lesser size, almost invisible and impunc-

tate elytral striae, small and barely rasp-like elytral punctures, wide d genital ring, rather wide, basally

very curved aedeagus with wide, slightly turned apex markedly acute ? stylomere.

Description

Measurements. Length: 6.70-6.75 mm. Ratios. Width/length of pronotum: 1.79-1.81; width base/

apex of pronotum: 1.76; width pronotum/head: 1.85-1.88; length/width of elytra: 1.23-1.27; length

elytra/pronotum: 2.23-2.33.

Colour. Dark piceous to black, lateral borders and apex of abdomen piceous. Lower surface of

head and thorax black, of abdomen dark reddish-piceous, posteriorly lighter. Mouthparts, antennae,

and legs dark reddish-piceous.

Head (Figs 106a-d, 295). Short and wide, rather depressed. Anterior border feebly convex, lateral

angle feebly obtuse. Clypeal suture shallow, in middle more or less reduced, semicircular. Labrum

comparatively wide, apex gently concave. Antennal groove laterally barely bordered, latero-posteri-

orly with convex, not carinate area. Mental tooth triangular, short, apex acute. Wings of mentum wide,

near apex strongly rounded. Glossa wide, tongue-like, apically widely rounded, ventrally with distinct

keel, at border with c. 8 elongate setae and additional pilosity on upper and lower surface and along

border. Terminal palpomere of maxillary palpus distinctly widened, apex securiform. Terminal

palpomere of labial palpus large, extremely widened, much wider than long, markedly securiform.

Antenna rather short, 8th-9th antennomeres wider than long. Microreticulation very fine, dense,

puncturation minute, difficult to see, surface with a shallow sulcus medially of eyes, impilose, slightly

Figs 106a-l. Adelotopus howdenorum, spec. nov. Details of head and genitalia. For legends see fig. 100.

silky. Ventrolaterally of eyes with a row of short setae. Suborbital field finely punctate and densely

setose. Both palpi rather sparsely setose, gula punctate and setose.

Pronotum (Fig. 295). Wide, moderately convex, base c. 1.75 x as wide as apex. Apical angles

produced, rather acute, attaining posterior third to half of eyes. Apex deeply, rectangularly excised,

slightly convex in excision, not bordered. Sides strongly and evenly curved, widest at or near basal

angles, barely or not at all bordered, moderately explanate. Basal angles slightly obtuse, at apex barely

rounded off, base straight, sometimes in middle feebly produced, not bordered. Microreticulation fine

though distinct, almost isodiametric, puncturation minute, more or less easily detected under high

magnification, surface with some fine, irregular strioles, impilose, rather silky.

Elytra (Figs 295, 447). Rather short and wide, moderately convex, in basal two fifth almost parallel,

then gently narrowed. Apex slightly oblique, apical angles widely rounded, truncature feebly concave.

Shoulders obtuse, basal margin oblique, with a row of 6-8 short setae a distance down the lateral

margin, margin there slightly serrate. Marginal channel narrow, completely visible from above. Basal

border complete, well developed, meeting at suture. Lateral border asetose. Series of umbilical pores

consisting of 6 pores behind shoulder and 1 pore (rarely unilaterally 2 pores) behind middle or at

apical third. Setae very short. Striae in basal two thirds vaguely indicated as extremely fine, wavy lines,

apically absent, sutural stria invisible, suture apically raised. Puncturation at base indistinct, apically

becoming gradually denser and more distinct, though still fine. Near apex punctures faintly rasp-like,

rather irregular. Microreticulation fine, though distinct, isodiametric. Surface impilose, somewhat

silky.

Lower surface. Prosternal process moderately elongate, wide, straight, rather depressed, apex

widely rounded, shortly setose. Metepisternum moderately elongate, slightly <2 x as long as wide,

posteriorly constricted and hollowed. Abdominal sterna with 1-2 setae each side, sternum VI in both

sexes moderately punctate and shortly setose, with many (c. 16-24), fairly elongate setae at and near

apical border. Lower surface rather densely punctate and pilose.

Legs. Rather elongate, 1st tarsomere of protarsus not wider than long, tibial groove of profemur

comparatively shallow, anterior plate only at apex distinctly overlapping the groove, posteriorly

border of groove not sharp. Metatibia rather narrow, >5 x as long as wide, Ist tarsomere of metatarsus

>2 x as long as wide. 3 protarsus barely widened.

d genitalia (Figs 106e-k). Genital ring wide, rather asymmetric, with elongate apex. Sternum VII

very wide, apically moderately excised, basally slightly concave, lateral parts elongate. Aedeagus

moderately elongate, fairly wide, moderately depressed, rather symmetric, lower surface near apex

gently convex. Apex very wide, rounded off, slightly turned laterally. Orifice elongate, internal sac

rather simple. Right paramere moderately elongate, apex unknown, broken. Left paramere much

larger, margins straight, apex transversely cut.

7 genitalia (Fig. 1061). Stylomere 2 triangular, apically narrow and acute, with 3 subapical setae.

Lateral plate rather elongate, with 1-2 elongate and 1-2 short apical setae.

Variation. Little noted due to scarce material. The ? specimen is slightly narrower and more

convex.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. So far collected in August and September.

Distribution (Fig. 605). Interior of southwestern Australia.

Material examined (2). Only the type series.

Etymology. Named in honour of the collectors of the holotype.

katherinei-group

Diagnosis. Medium-sized, rather depressed, uniformly black species. Labrum bisetose; glossa

c. 10-setose; lateral margin of pronotum widely explanate, basal angle rounded off; basal border line

of elytra abbreviated, attaining inner quarter of base; scutellar pore absent; lateral margin of elytra

without elongate setae behind shoulders; series of lateral pores with 6 subhumeral pores and

1 postmedian pore; abdominal sterna without ambulatory setae; sternum VI without longer setae at

apical margin; tibiae, especially metatibia depressed; mesofemur and metafemur wide and depressed,

profemur less wide; internal sac of aedeagus complicate, with oblique fold near apex.

Larvae. Unknown.

Distribution. A single species from the northern part of the Northern Territory.

Systematic position. This group is certainly closely related to and is perhaps the adelphotaxon of the

brevipennis-group, though it is slightly more plesiomorphic in the longer basal border of elytra and the

presence of the postmedian marginal pore.

Adelotopus katherinei, spec. nov.

Figs 31, 107, 296, 448, 605

Types. Holotype: d, Australien, NT, Katherine Gorge, 6.-8.11.1984, M. & B. Baehr (MNTD).

Diagnosis. Medium-sized, rather short and wide, convex, dull piceous-black species with explanate,

at base rounded margin of pronotum. Further distinguished from related species by dense and coarse

elytral microreticulation, impilose surface, short and wide apex of prosternal process, and narrow,

elongate parameres with somewhat acute apex.

Description

Measurements. Length: 5.8 mm. Ratios. Width/length of pronotum: 1.85 ; width base/apex of

pronotum: 1.63; width pronotum/head: 1.69; length/width of elytra: 1.21; length elytra/pronotum:

2,30.

Colour. Black, lateral margins of pronotum and elytra reddish-piceous translucent. Lower surface

piceous. Mouth parts and antenna reddish-piceous, legs piceous.

Head (Figs 107a-d). Short and rather wide, moderately depressed. Anterior border gently convex,

lateral angle obtusely rounded, laterally slightly projecting, lateral borders oblique. Clypeal suture

semicircular, in middle narrowly interrupted. Labrum rather narrow, apex deeply concave. Antennal

groove laterally sharply bordered, latero-posteriorly with carinate area. Mental tooth triangular, short,

apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like,

Figs 107a-k. Adelotopus katherinei, spec. nov. Details of head and Ö genitalia. For legends see fig. 100.

apically widely rounded, ventrally with distinct keel, at border with c. 10 elongate setae and additional

pilosity on upper and lower surface and along border. Terminal palpomere of maxillary palpus

moderately wide, slightly securiform. Terminal palpomere of labial palpus very wide, markedly

securiform. Antenna short, Sth-9th antennomeres almost 2x as wide as long. Microreticulation

extremely fine, though distinct, puncturation very fine and fairly dense, surface with a shallow sulcus

medially of eyes, impilose, moderately glossy. Ventrolaterally of eyes with a row of short setae.

Suborbital field at least laterally faintly punctate and setose. Both palpi rather sparsely setose, gula

almost asetose.

Pronotum (Fig. 296). Wide, convex, base wide, apex rather narrow. Apical angles moderately

produced, fairly acute, slightly oblique, attaining posterior third of eyes. Apex moderately deeply

excised, rather convex in excision, not bordered. Sides curved throughout, widest near basal angles.

Margins faintly bordered, fairly explanate. Basal angles shortly rounded off. Base almost straight, not

bordered. Surface near base without transverse impression. Microreticulation rather fine, very distinct,

puncturation very fine, moderately dense, rather difficult to detect, surface with several short,

irregular strioles, impilose, rather dull, faintly coriaceous.

Elytra (Figs 296, 448). Short, rather convex, margins slightly convex throughout. Apex rather wide,

oblique, truncature faintly concave, apical angles rounded off. Shoulders rounded off, basal margin

slightly oblique, without setae behind shoulders. Marginal channel rather wide, completely visible

from above. Basal border incomplete, present in lateral % of elytron, ending gradually. Lateral border

asetose. Series of umbilical pores consisting of 6 pores behind shoulder and an additional pore behind

middle. Setae very short. Striae including sutural stria almost absent, only inner striae in basal half

vaguely indicated by extremely fine longitudinal lines. Microreticulation coarse, isodiametric, very

distinct, puncturation present though difficult to see. Surface with some faint irregular transverse

strioles, impilose, rather dull.

Lower surface. Prosternal process rather short, fairly wide, straight, depressed, apex short and

wide, rounded off, shortly setose. Metepisternum moderately elongate, c. 1.8 x as long as wide,

posteriorly not constricted nor hollowed. Abdominal sterna including sternum VI apparently without

setae at apical border. Lower surface rather densely punctate and pilose.

Legs. Moderately elongate, 1st tarsomere of protarsus about as long as wide, tibial groove of

profemur deep, anterior plate distinctly overlapping the groove for some distance from apex, posterior

border of groove sharp. Femur rather wide. Metatibia moderately elongate, <5 x as long as wide, 1st

tarsomere of metatarsus c. 2 x as long as wide. d protarsus not widened.

So genitalia (Figs 107e-k). Genital ring rather narrow, triangular, slightly asymmetric, arms feebly

convex, with rather narrow, asymmetric base. Sternum VII rather wide, apically straight, with rather

shallow excision, basally almost faintly trianglar and bisinuate, lateral parts triagonal, rather elongate.

Aedeagus short, moderately depressed, in middle rather widened, slightly asymmetric. Lower surface

gently convex. Apex rather wide, widely rounded. Orifice rather short, internal sac fairly complex,

though in single available specimen partly everted. Both parameres rather elongate, with obtusely

rounded apex, left considerably larger than right.

7 genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Collected under bark of gum-type eucalypt in November.

Distribution (Fig. 605). Northern part of Northern Teritory. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the type locality, the Katherine Gorge.

brevipennis-group

Diagnosis. Medium-sized, rather depressed, uniformly black to yellowish, or vividly patterned

species. Labrum bisetose; glossa c. 10-setose; lateral margin of pronotum widely explanate, basal angle

angulate or obtuse; basal border line of elytra abbreviated, rarely longer than half of base; scutellar

pore absent; lateral margin of elytra without elongate setae behind shoulders; series of lateral pores

with 2-6 subhumeral pores only; abdominal sterna usually without ambulatory seta each side, only in

one species with 1-2 setae; sternum VI without longer setae at apical margin; tibiae, especially

metatibia depressed; mesofemur and metafemur wide and depressed, profemur less wide; internal sac

of aedeagus complicate, with oblique fold near apex.

Larvae. Ist instar larvae known of 2 species.

Distribution. 12 species and additional 2 subspecies from whole eastern Australia, the northern part

of Northern Territory, northwestern Australia, and the interior of southwestern Australia. The bulk of

the species, however, concentrates in northern and northwestern Australia.

Systematic position. This group is perhaps generalized in some aspects, e.g. depressed body with

explanate margins of pronotum, but apomorphic in several others, e.g. light colour of some species,

absence of ambulatory setae of the abdomen, complexiy shaped internal sac of aedeagus. It is

presumably the apomorphic adelphotaxon of the katherinei-group.

Adelotopus brevipennis Macleay, 1888

Figs 32, 108, 297, 449, 606

Adelotopus brevipennis Macleay, 1888, p. 459; Sloane 1898, p. 514; Lea 1910, p. 120, 121; Notman 1925, p. 7, 28; Csiki

1933, p. 1634; Moore et al. 1987, p. 49.

Types. Lectotype (by present designation): d, Syntype, Adelotopus brevipennis Macl., Kings Sound (ANIC-

MMS). - Paralectotype: 13, same data, on same card (ANIC-MMS).

Type locality. “Kings Sound”, Western Australia.

Diagnosis. Medium-sized, short and wide, rather depressed, dull black to dark piceous species.

Further distinguished from relative species by low number of umbilical pores, absence of distinct

reddish margins of pronotum and elytra, very wide, markedly triangular d genital ring, and depressed

and extremely wide aedeagus with acute apex.

Figs 108a-l. Adelotopus brevipennis Macleay. Details of head and genitalia. For legends see fig. 100.

Description

Measurements. Length: 5.3-6.8 mm. Ratios. Width/length of pronotum: 1.71-1.83; width base/

apex of pronotum: 1.69-1.80; width pronotum/ head: 1.74-1.81; length/width of elytra: 1.25-1.29; length

elytra/pronotum: 2.27-2.34.

Colour. Dark piceous to dull black, sometimes suture and lateral borders slightly lighter. Lower

surface reddish-piceous. Mouthparts, antennae, and legs dark reddish to reddish-piceous.

Head (Figs 108a-d, 297). Short and wide, rather depressed. Anterior border feebly convex, lateral

angle slightly obtuse, though angles laterally distinctly projecting. Clypeal suture shallow, semicircu-

lar, in middle interrupted. Labrum narrow, apex concave. Antennal groove laterally sharply bordered,

latero-posteriorly with convex, not carinate area. Mental tooth triangular, short, apex acute. Wings of

mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically widely rounded,

ventrally with distinct keel, at border with c. 10 elongate setae and additional pilosity on upper and

lower surface and along border. Terminal palpomere of maxillary palpus moderately elongate, slightly

widened, not markedly securiform. Terminal palpomere of labial palpus very wide, markedly

securiform. Antenna very short, 8th-9th antennomeres almost 2 x as wide as long. Microreticulation

extremely fine, dense, superficial, puncturation very fine, dense, though easily visible under high

magnification, surface with a shallow sulcus medially of eyes, impilose, fairly silky. Ventrolaterally of

eyes with a row of short setae. Suborbital field impunctate and asetose. Both palpi rather sparsely

setose, gula almost asetose.

Pronotum (Fig. 297). Moderately wide, moderately convex, base c. 1.75 x as wide as apex. Apical

angles produced, rather acute, attaining posterior third to half of eyes. Apex deeply, rectangularly

excised, rather convex in excision, not bordered. Sides strongly and evenly curved, widest near basal

angles, very finely, almost unbordered, moderately explanate. Basal angles almost rectangular, shortly

rounded off, base straight to faintly concave, in middle more or less distinctly bordered. Microretic-

ulation extremely fine and dense, rather superficial, almost isodiametric, puncturation fine, though

rather distinct, dense, surface with or without some very fine, irregular strioles near base, impilose,

fairly glossy, though somewhat silky.

Elytra (Figs 297, 449). Rather short and wide, moderately convex, in basal three fourth almost

parallel, then gently narrowed. Apex wide, almost transverse, truncature feebly convex, apical angles

shortly rounded. Shoulders rounded, basal margin oblique, without setae behind shoulders. Marginal

channel moderate, completely visible from above. Basal border incomplete, reaching only to middle

of base, ending abruptly. Lateral border asetose. Series of umbilical pores usually consisting of 2 pores

behind shoulder, sometimes only 1 pore or, more commonly and almost invariably in specimens from

Hamersley Range, 3 pores present (in one specimen unilaterally even 5 pores!). Setae very short. Striae

even in basal half almost invisible or most vaguely indicated as extremely fine, wavy lines, apically

absent, sutural stria invisible. Microreticulation very fine, somewhat superficial, isodiametric to

slightly transverse and arranged in irrgular transverse rows, less distinct than on fore body, becoming

weaker towards apex. Puncturation fine, rather sparse, basally very indistinct, apically becoming

denser and more distinct and faintly rasp-like. Surface impilose, moderately glossy, slightly silky.

Lower surface. Prosternal process rather elongate, rather wide, straight, depressed, apex markedly

widened widely rounded off, shortly setose. Metepisternum rather elongate, slightly <2 x as long as

wide, posteriorly constricted and hollowed. Abdominal sterna including sternum VI without setae at

apical border. Lower surface rather sparsely punctate and pilose.

Legs. Moderately elongate, 1st tarsomere of protarsus distinctly wider than long, tibial groove of

profemur rather deep, anterior plate at apex distinctly overlapping the groove, posteriorly border of

groove sharp. Metatibia rather elongate, c. 5 x as long as wide, Ist tarsomere of metatarsus c. 2x as

long as wide. d protarsus slightly widened.

d genitalia (Figs 108e-k). Genital ring very wide, markedly triangular, with very narrow base.

Sternum VII wide, apically feebly excised or only concave, basally triangularly convex. Aedeagus

rather short, depressed, in middle remarkably widened, highly asymmetric, left border straight of

faintly concave, markedly oblique, right border convex. Lower surface gently convex. Apex short,

acute. Orifice rather short, internal sac fairly complex, with a large, oblique fold near apex. Right

paramere fairly narrow, apex rather acute, left large, upper border strongly convex, apex narrow,

gently rounded or almost acute.

? genitalia (Fig. 1081). Stylomere triangular, rather wide, apex narrow, rounded off, with 3-4

subapical setae. Lateral plate elongate, with 4-7 apical setae.

Variation. Slight variation noted in relative width of pronotum and elytra and in shape of d and

? genitalia. There is a very large specimen from , however, that varies so some degree from the rather

uniform appearance of this species, mainly by large size, wide pronotum, and wide and distinctly

lighter margins of pronotum and elytra. It is but tentatively alluded to this species.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unknown. Specimens collected by me under bark of river eucalypts near the borders

of dry creeks and dry and water-bearing rivers, also at light, aspecimen captured by T. F. Houston “on

trunk of Eucalyptus sp. camaldulensis”, others in “Malaise trap”. Some specimens mounted together

with ants of the genus Iridomyrmex Mayr on same card. So far collected in the period from October to

April.

Distribution (Fig. 606). Northern part of Northern Territory east to border of Arnhem Land, adjacent

tropical Western Australia south to Fortescue River.

Material examined (59). NT: 2??, Humpty Doo 12°35’S 131°05’E, Mt. Mortgage, 3 Dec 1991-17 Jan 92, Wells

& Webber (MNTD); 3? 2, Humpty Doo 12°35’S 131°05’E, Mt. Mortgage, 18 Jan -28 Feb 1992, Wells & Webber

(MNTD); 18, 17, Mary River, 110 km e. Darwin, 1.11.1984, M. & B. Baehr (CBM); 1%, NT 95/52, Mary River,

115 km e. Darwin, 29.-30.8.1995, M. Baehr (CBM); 19, Port Darwin, (MMS); 18, NT 95/48, 30 km n. Edith River

Cr., 28.8.1995, M. Baehr (CBM); 14, Katherine Gorge, 6.-8.11.1984, M. & B. Baehr (CBM); 1%, N. T. Katherine

Gorge NP, 4-6 Dec.1980, M. B. Malipatil (MNTD); 15, 22%, Katherine IV.71 (CSB); 2??, Daly, N.W.A., W. D.

Dodd (SAMA); 13, NT 95/41, 70 km e. Kununurra, 23.8.1995, M. Baehr (CBM). - WA: 238, 1%, Ord River b.

Ivanhoe, 11.-13.11.1984, M. & B. Baehr (CBM); 12, Wyndham, S. Stephens 20.11.01, Adelotopus (SAMA);

235g, Kimberley district, Mjöberg, Adelotopus brevipennis Macl. Id. by T. G. Sloane (NHRS); 19, Ord R. brevipennis,

Adelotopus brevipennis M. (SAMA): 12 >22, 135 km n. Halls Creek, 14.11.1984, M. & B. Baehr (CBM, MCZ,

ZSM); 12, WA 95/33, Elvire River, ! Halls Creek, 20.8.1995, M. Baehr (CBM); 16, Fitzroy Crossing,

18.-20.11.1984, M. & B. Baehr (CBM); 1 A 95/30, Fitzroy Crossing, 18.-19.8.1995, M. Baehr (CBM); 15,

WA 95/31, 110 km se. Fitzroy Crossing, 995, M. Baehr (CBM); 14, 150 km e. Derby, 2km w. Windjana

Gorge, 22.11.1984, M. & B. Baehr (CBM),; berley Lennard River Xing Gibb River Road 17.235, 124.44E 31

9 8

Figs 109a-l. Adelotopus elongatulus Macleay. Details of head and genitalia. For legends see fig. 100.

March 1988 T. F. Houston 679-1 (WAM 94/ 879); 23d, Syntype, Adelotopus brevipennis Macl., Kings Sound

(lectotype!, paralectotype!)(ANIC-MMS); 13, W. W. Froggatt Coll., Adelotopus elongatus Macl. Kings Sound

(ANIC); 15, Derby, Mjöberg, Oct. Adelotopus brevipennis Macl. Id. by T. G. Sloane (NHRS); 13, D. D. Giuliani

6.X11.1968, Derby, Adelotopus sp. E. B. Britton det. 1971 (WAM); 536, 32%, Fortescue R. Hammersley Range:

W.D. Dodd, Adelotopus (SAMA); 2%, Millstream, 3.-5.12.1984, M. & B. Baehr (CBM).

Adelotopus elongatulus Macleay, 1888

Figs 109, 298, 450, 607

Adelotopus elongatulus Macleay, 1888, p. 459; Notman 1925, p. 6, 28; Csiki 1933, p. 1635; Moore et al. 1987, p. 50.

Adelotopus longipennis Macleay, 1888, p. 460; Notman 1925, p. 7, 29; Csiki 1933, p. 1635; Moore et al. 1987, p. 51

(new synonymy).

Types. Of elongatulus. Lectotype (by present designation): d, N. W. Austr., Syntype, Adelotopus elongatulus Macl.

Kings Sound. N. W. A. (ANIC-MMS). — Paralectotype: 15, N. W. Austr., Syntype, Adelotopus elongatulus Macl.

Kings Sound. N. W. A. (ANIC-MMS).

Of longipennis. Lectotype (by present designation): ?, N. W. Austr., Syntype, Adelotopus longipennis Macl.

Kings Sound. N. W. A. (ANIC-MMS). - Paralectotype: 17, N. W. Austr., Syntype, Adelotopus longipennis Macl.

Kings Sound. N. W. A. (ANIC-MMS).

Type localities. Of elongatulus: “Kings Sound”, Western Australia. - Of longipennis: “Kings Sound”,

Western Australia.

Diagnosis. Medium-sized, narrow and elongate, rather depressed, dark reddish-piceous species with

lighter margins and suture. Further distinguished from relative species by moderately wide, on left

side oblique aedeagus with rounded apex, and short and wide parameres.

Description

Measurements. Length: 5.65-6.2 mm. Ratios. Width/length of pronotum: 1.69-1.72; width base/

apex of pronotum: 1.51-1.60; width pronotum/head: 1.55-1.63; length/width of elytra: c. 1.62-1.68;

length elytra/pronotum: 2.59-2.75.

Colour. Dark reddish-piceous, all margins of pronotum, suture and lateral margins of elytra

slightly lighter. Lower surface, mouth parts, and legs reddish-piceous.

Head (Figs 109a-d). Short and wide, rather depressed. Anterior border convex, lateral angle obtuse,

angle laterally slightly projecting. Clypeal suture shallow, semicircular, in middle interrupted. Labrum

narrow, apex concave. Antennal groove laterally sharply bordered, latero-posteriorly with convex, not

carinate area. Mental tooth triangular, short, apex acute. Wings of mentum wide, laterally rounded,

apex rectangular. Glossa wide, tongue-like, apically widely rounded, ventrally with distinct keel, at

border with c. 10 elongate setae and additional pilosity on upper and lower surface and along border.

Terminal palpomere of maxillary palpus rather wide, fairly securiform. Terminal palpomere of labial

palpus very wide, markedly securiform. Antenna very short, 8th-9th antennomeres c. 1.5 x as wide as

long. Microreticulation extremely fine, dense, superficial, puncturation fine, dense, though easily

visible under high magnification, surface with a shallow sulcus medially of eyes, impilose, fairly silky.

Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Both palpi

rather sparsely setose, gula almost asetose.

Pronotum (Fig. 298). Moderately wide, moderately convex, apex rather wide, base >1.5 x as wide

as apex. Apical angles produced, rather acute, attaining posterior third of eyes. Apex deeply,

rectangularly excised, rather convex in excision, not bordered. Sides moderately curved, widest behind

middle, though but slightly curved to base. Sides almost unbordered, moderately explanate. Basal

angles almost rectangular, slightly produced backwards, at apex shortly rounded off. Base laterally

faintly concave, in middle slightly produced, very indistinctly bordered. Microreticulation extremely

fine and dense, rather distinct, almost isodiametric, puncturation moderately fine, distinct, dense,

surface with some very fine, irregular strioles near base, impilose, fairly glossy, though somewhat

silky.

Elytra (Figs 298, 450). Elongate and narrow, moderately convex, parallel, only near apex gently

narrowed. Apex wide, almost transverse, truncature feebly convex, apical angles shortly rounded.

Shoulders obtuse, basal margin slightly oblique, without setae behind shoulders. Marginal channel

rather narrow, though deep, completely visible from above. Basal border incomplete, reaching only to

middle of base, ending abruptly. Lateral border asetose. Series of umbilical pores usually consisting

of 6, sometimes unilaterally 7 pores behind shoulder. Setae short. Striae even in basal half almost

invisible or vaguely indicated as extremely fine, wavy lines, apically absent, sutural stria invisible.

Microreticulation fine, somewhat superficial, isodiametric, less distinct than on fore body. Punctura-

tion very fine, rather sparse, much weaker than on fore body, basally very indistinct, apically more

distinct. Surface impilose, moderately glossy, slightly silky.

Lower surface. Prosternal process rather elongate, narrow, straight, depressed, apex narrow,

straight, slightly rounded off, shortly setose. Metepisternum rather elongate, slightly >2 x as long as

wide, posteriorly constricted but barely hollowed. Abdominal sterna including sternum VI without

setae at apical border. Lower surface rather sparsely punctate and pilose.

Legs. Moderately elongate, Ist tarsomere of protarsus barely wider than long, tibial groove of

profemur rather deep, anterior plate only at apex distinctly overlapping the groove, posterior border

of groove less sharp. Metatibia rather elongate, >5 x as long as wide, 1st tarsomere of metatarsus >2 x

as long as wide. d protarsus slightly widened.

d genitalia (Figs 109a-d). Genital ring moderately wide, slightly assymetric, with narrow base.

Sternum VII rather wide, apically fairly excised, basally triangularly convex. Aedeagus medium-sized,

depressed, in middle moderately widened, asymmetric, left border gently convex, oblique. Lower

surface gentlx convex. Apex rather wide, rounded off. Orifice rather elongate , internal sac fairly

complex, with a short fold near apex. Both parameres short and wide, right with widely rounded apex,

left large, upper border strongly convex, apex narrow, gently rounded.

? genitalia (Fig. 1091). Stylomere triangular, moderately wide, apex wide, rounded off, with 1-2

subapical setae. Lateral plate rather short, with 3 apical setae.

Variation. Only some variation of relative length of elytra noted. The light reddish coloured

syntypes of longipennis apparently immature specimens.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Only old, inaccurately labelled specimens from “Kings Sound” known.

100

Figs 110a-d, 1. Adelotopus rufomarginatus, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Distribution (Fig. 607). Northwestern part of Western Australia. Known only from type locality. The

type locality, however, is so inaccurate, that the species could be distributed throughout the whole of

the Kimberley Division. No recent records available.

Material examined (5). WA: 2593, Adelotopus elongatulus Macl. Kings Sound. (lectotype!, paralectotype!)

(ANIC-MMS); 2?%, Adelotopus longipennis Macl. Kings Sound. (lectotype!, paralectotype!) (ANIC-MMS); 164,

W. W. Froggatt Collection, Adelotopus elongatulus Mel. Kings Sd. (ANIC).

Adelotopus rufomarginatus, spec. nov.

Figs 110, 299, 451, 607

Types. Holotype: ?, Caranbirini W. H., 16.165, 136.05E, 33 km SW of Borroloola, NT, 3 Nov.1975, M. S. Upton

(ANIC).

Diagnosis. Medium-sized, short and wide, rather depressed, glossy black species with wide and

conspicuously reddish translucent borders of pronotum and elytra. Further distinguished from

relative species by low number of umbilical pores and the narrow and elongate stylomere.

Description

Measurements. Length: 7.0 mm. Ratios. Width/length of pronotum: 1.72; width base/apex of

pronotum: 1.82; width pronotum/head: 1.84; length/width of elytra: 1.25; length elytra/pronotum:

2.22:

Colour (Fig. 299). Glossy black, lateral border of pronotum and elytra distinctly reddish translu-

cent. Lower surface reddish-piceous, base and apex of abdomen light reddish-piceous. Mouth parts,

antennae, and legs reddish-piceous, tibiae and tarsi piceous.

Head (Figs 110a-d, 299). Short and very wide, rather depressed. Anterior border almost straight,

lateral angle obtuse, angle laterally distinctly projecting, lateral border concave. Clypeal suture

shallow, semicircular, in middle interrupted. Labrum very narrow, apex concave. Antennal groove

laterally moderately sharply bordered, latero-posteriorly with convex, slightly carinate area. Mental

tooth triangular, very short, apex acute. Wings of mentum wide, laterally rounded, apex rectangular.

Glossa wide, tongue-like, apically widely rounded, ventrally with distinct keel, at border with c. 10

elongate setae and additional pilosity on upper and lower surface and along border. Terminal

palpomere of maxillary palpus moderately wide, slightly securiform. Terminal palpomere of labial

palpus very wide, markedly securiform. Antenna very short, 8th-9th antennomeres almost 2 x as wide

as long. Microreticulation extremely fine, highly superficial, visible only iunder high magnification,

puncturation very fine, moderately dense, surface with a shallow sulcus medially of eyes, impilose,

glossy. Ventrolaterally of eyes with a row of short setae. Suborbital field punctate and finely setose.

Both palpi rather sparsely setose, gula almost asetose.

Pronotum (Fig. 299). Wide, moderately convex, base wide, apex rather narrow. Apical angles

101

produced, acute, attaining posterior half of eyes. Apex deeply, rectangularly excised, rather convex in

excision, faintly bordered. Sides anteriorly strongly curved, posteriorly straight, widest at base. Sides

almost unbordered, markedly explanate and channeled. Basal angles rectangular, at apex shortly

rounded off. Base laterally faintly concave, in middle slightly produced, almost straight, very

indistinctly bordered. Surface near base with shallow transverse impression. Microreticulation ex-

tremely fine, highly superficial, barely visible even under high magnification, isodiametric, punetur-

ation fine, though distinct, dense, surface with some very fine, irregular strioles near base, impilose,

glossy.

Elytra (Figs 299, 451). Short and wide, moderately convex, in basal three fourth almost parallel,

then gently narrowed. Apex wide, almost transverse, truncature feebly convex, apical angles shortly

rounded. Shoulders rounded, basal margin oblique, without setae behind shoulders. Marginal channel

wide and deep, completely visible from above. Basal border incomplete, reaching only to middle of

base, ending abruptly. Lateral border asetose. Series of umbilical pores consisting of 3 widely spaced

pores behind shoulder. Pores and setae very inconspicuous. Striae not visible, sutural stria absent.

Microreticulation extremely fine, highly superficial, barely visible even at high magnification, isodia-

metric, puncturation fine, though rather distinct, fairly dense. Surface impilose, glossy.

Lower surface. Prosternal process rather elongate, narrow, straight, depressed, apex narrow,

straight, slightly rounded off, shortly setose. Metepisternum rather elongate, c. 2 x as long as wide,

posteriorly constricted but barely hollowed. Abdominal sterna including sternum VI without setae at

apical border. Lower surface rather densely punctate and pilose.

Legs. Moderately elongate, 1st tarsomere of protarsus slightly wider than long, tibial groove of

profemur rather deep, anterior plate only at apex distinctly overlapping the groove, posterior border

of groove rather sharp. Metatibia rather narrow, c. 5 x as long as wide, 1st tarsomere of metatarsus

c.2 x as long as wide. S protarsus unknown.

d genitalia. Unknown.

? genitalia (Fig. 1101). Stylomere narrow and elongate, apex narrow, obtuse, with 2-4 subapical

setae. Lateral plate elongate, with 4-5 apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Holotype collected in November.

Distribution (Fig. 607). Northeastern part of Northern Territory. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the wide, rufous margins of pronotum and elytra.

Adelotopus adelaideae, spec. nov.

Figs 111, 300, 452, 607

Types. Holotype: d, Adelaide River 92.-20, Adelotopus sp. ? Id. by T. G. Sloane (BMNH). - Paratypes: 584,322,

Adelaide River, N. W. Australia, J. J. Walker, G. C. Champion Coll. 1927-409 (BMNH, CBM); 19,22, Adelaide

River 92-20 (BMNH).

Diagnosis. Medium-sized, rather elongate, depressed, markedly dull, black species with explanate

margin of pronotum. Further distinguished from relative species by dense and rugose puncturation of

pronotum, moderately wide aedeagus with rounded apex and rather large fold in internal sac, and

large, similarly shaped parameres.

Description

Measurements. Length: 6.1-7.3 mm. Ratios. Width/length of pronotum: 1.80-1.84; width base/

apex of pronotum: 1.72-1.77; width pronotum/ head: 1.80-1.86; length/ width of elytra: 1.36-1.39; length

elytra/pronotum: 2.58-2.68.

Colour. Dull black, borders of pronotum and elytra sometimes indistinctly reddish-piceous

translucent. Lower surface blackish to piceous. Mouth parts and antennae reddish-piceous, legs

piceous.

102

Figs 111a-l. Adelotopus adelaideae, spec. nov. Details of head and genitalia. For legends see fig. 100.

Head (Figs 111a-d). Short and wide, rather depressed. Anterior border gently convex, lateral angle

rounded, angle laterally barely projecting. Clypeal suture almost invisible. Labrum very narrow, apex

concave. Antennal groove laterally moderately sharply bordered, latero-posteriorly with convex, not

carinate area. Mental tooth triangular, short, apex acute. Wings of mentum wide, laterally rounded,

apex rectangular. Glossa wide, tongue-like, apically widely rounded, ventrally with distinct keel, at

border with c. 10 elongate setae and additional pilosity on upper and lower surface and along border.

Terminal palpomere of maxillary palpus moderately wide, barely securiform. Terminal palpomere of

labial palpus rather wide, securiform. Antenna comparatively elongate, Sth-9th antennomeres c. 1.3 x

as wide as long. Microreticulation fine, rather distinct, puncturation dense and moderately coarse,

surface with a shallow sulcus medially of eyes and with some strioles, impilose, dull. Ventrolaterally

of eyes with a row of short setae. Suborbital field almost impunctate and asetose. Both palpi rather

sparsely setose, gula almost asetose.

Pronotum (Fig. 300). Wide, rather depressed, base wide, apex rather narrow. Apical angles

produced, acute, attaining posterior third of eyes. Apex deeply, rectangularly excised, straight in

excision, faintly bordered. Sides anteriorly strongly curved, posteriorly feebly curved, widest slightly

in front of base. Sides almost unbordered, fairly explanate, but not channeled. Basal angles c. 100°, apex

obtuse. Base almost straight, very indistinctly bordered. Surface near base with more or less distinct

transverse impression. Microreticulation fine, rather superficial, isodiametric to somewhat irregular,

puncturation dense and moderately coarse, surface with network of irregular strioles, coriaceous,

impilose, markedly dull.

Elytra (Figs 300, 452). Moderately elongate, depressed, markedly parallel, gently narrowed near

apex. Apex wide, faintly oblique, truncature feebly convex, apical angles shortly rounded. Shoulders

rather distinct, basal margin slightly oblique, without setae behind shoulders. Marginal channel

narrow, completely visible from above. Basal border incomplete, reaching only to middle of base,

ending abruptly. Lateral border asetose. Series of umbilical pores consisting of 6 pores behind

shoulder. Setae very short. Striae including sutural stria absent. Microreticulation fine, though distinct,

isodiametric, puncturation fine, moderately dense, much finer and sparser than on pronotum,

punctures somewhat rasp-like. Surface impilose, markedly dull.

Lower surface. Prosternal process rather elongate, narrow, straight, depressed, apex narrow,

straight, slightly rounded off, shortly setose. Metepisternum moderately elongate, cslightly <2 x as

103

long as wide, posteriorly not constricted nor hollowed. Abdominal sterna with 1-2 setae at apical

border. Sternum VI without elongate setae. Lower surface densely punctate and pilose.

Legs. Rather elongate, 1st tarsomere of protarsus barely wider than long, tibial groove of profemur

moderately deep, anterior plate only at apex distinctly overlapping the groove, posterior border of

groove rather sharp. Femur comparatively narrow. Metatibia narrow and elongate, c. 6 x as long as

wide, Ist tarsomere of metatarsus c. 2.5 x as long as wide. ö protarsus not widened.

d genitalia (Figs 111e-k). Genital ring rather wide, slightly assymetric, with narrow base. Sternum

VII rather wide, apically rather deeply excised, basally almost straight, lateral parts large. Aedeagus

medium-sized, rather depressed, in middle moderately widened, asymmetric, left border gently

convex, oblique. Lower surface gently convex to almost straight. Apex rather wide, rounded off.

Orifice rather short, internal sac fairly complex, with a wide fold near apex. Both parameres large and

rather wide, with widely rounded apex, left considerably largerthan right.

? genitalia (Fig. 1111). Stylomere triangular, moderately wide, apex rather narrow, rounded off,

with 2-3 subapical setae. Lateral plate rather elongate, apparently without apical setae.

Variation. Apart from some differences of size very little variation noted.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Not even the dates of collecting known. This species is actually known only from

old material.

Distribution (Fig. 607). Northern part of Northern Territory. Known only from type locality.

Material examined (12). Only the type series.

Etymology. The name refers to the type locality, the Adelaide River.

Adelotopus rufescens, spec. nov.

Figs 112, 113, 301, 453, 607

Types. Holotype: d, Australia: n WA Kununurra 22.X11.1991-5.1.1992 R. I. Storey (QMB T26064). — Paratypes:

12, same data (DPIM); 12, A. Douglas. leg. 1-12.11.1965, Mt. Hart. W. Aust. (WAM 87/2171).

Diagnosis. Medium-sized, rather wide, depressed, dull, light reddish species with explanate, rectan-

gular margin of pronotum. Further distinguished from relative species by very wide pronotum, rather

narrow, triangular d genital ring, moderately wide aedeagus that is widest close to apex, with rounded

apex and rather small fold in internal sac, and large left paramere with wide apex.

Description

Measurements. Length: 6.5-6.9. Ratios. Width/length of pronotum: 1.84-1.88; width base/apex of

pronotum: 1.74-1.76; width pronotum/head: 1.83-1.86; length/ width of elytra: 1.24-1.33; length elytra/

pronotum: 2.37-2.47.

Colour. Upper and lower surface including mouth parts, antennae, and legs uniformly light

reddish.

Head (Figs 112a-d). Short and wide, rather depressed. Anterior border convex, lateral angle

rounded, angle laterally slightly projecting, lateral borders oblique. Clypeal suture semicircular, in

middle interrupted. Labrum narrow, apex concave. Antennal groove laterally sharply bordered,

latero-posteriorly with carinate area. Mental tooth triangular, short, apex acute. Wings of mentum

wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically widely rounded, ventral-

ly with distinct keel, at border with c. 12 elongate setae and additional pilosity on upper and lower

surface and along border. Terminal palpomere of maxillary palpus rather narrow, barely securiform.

Terminal palpomere of labial palpus rather wide, securiform. Antenna moderately short, 8th-9th

antennomeres slightly <1.5 x as wide as long. Microreticulation fine, rather distinct, puncturation very

fine, rather dense, fairly difficult to see, surface with a shallow sulcus medially of eyes, rather densely

pilose, fairly dull. Ventrolaterally of eyes with a row of short setae. Suborbital field at least laterally

punctate and setose. Both palpi rather sparsely setose, gula almost asetose.

Pronotum (Figs 113, 301). Wide, depressed, base wide, apex rather narrow. Apical angles

produced, acute, attaining posterior half of eyes. Apex deeply, rectangularly excised, faintly convex in

104

Figs 112a-l. Adelotopus rufescens, spec. nov. Details of head and genitalia. For legends see fig. 100.

excision, faintly bordered. Sides anteriorly strongly curved, posteriorly feebly though distinctly

curved, widest slightly in front of base. Sides faintly bordered, explanate, but not channeled. Basal

angles almost rectangular, distinctly produced backwards, apex obtuse. Base laterally concave, in

middle produced, very indistinctly bordered. Surface near base with more or less distinct transverse

impression. Microreticulation fine, though distinct, isodiametric to somewhat irregular, puncturation

dense and moderately fine, surface with faint network of irregular strioles, somewhat coriaceous,

densely pilose, dull.

Elytra (Figs 301, 453). Moderately elongate, depressed, in basal two thirds parallel, then gently

narrowed near apex. Apex rather wide, faintly oblique, truncature feebly convex, apical angles shortly

rounded. Shoulders rather distinct, basal margin slightly oblique, without setae behind shoulders.

Marginal channel rather narrow, completely visible from above. Basal border incomplete, reaching

only to middle of base, ending abruptly. Lateral border asetose. Series of umbilical pores consisting of

6 pores behind shoulder. Setae very short. Striae including sutural stria absent. Microreticulation fine,

though distinct, isodiametric, not coriaceous, puncturation fine, moderately dense, much finer and

sparser than on pronotum, apically punctures somewhat rasp-like. Surface densely pilose, rather dull.

Lower surface. Prosternal process rather elongate, narrow, straight, depressed, apex narrow,

straight, faintly rounded off, shortly setose. Metepisternum moderately elongate, c. 1.8 x as long as

wide, posteriorly not constricted nor hollowed. Abdominal sterna including sternum VI apparently

without setae at apical border. Lower surface densely punctate and pilose.

Legs. Elongate, 1st tarsomere of protarsus slightly longer than wide, tibial groove of profemur

moderately deep, anterior plate only at apex distinctly overlapping the groove, posterior border of

groove sharp. Femur comparatively narrow. Metatibia narrow and elongate, c. 6 x as long as wide, Ist

tarsomere of metatarsus >2.5 x as long as wide. 3 protarsus not widened.

d genitalia (Figs 112e-k). Genital ring rather narrow and elongate, triangular, slightly assymetric,

with narrow base. Sternum VII moderately wide, apically moderately deeply excised, basally feebly

convex, lateral parts small. Aedeagus medium-sized, rather depressed, in apical third moderately

widened, asymmetric. Lower surface gently convex to almost straight. Apex rather wide, rounded off.

Orifice elongate, internal sac fairly complex, with a small fold near apex. Both parameres rather large

and wide, right with rounded apex, left considerably larger than right and with very wide apex.

? genitalia (Fig. 1121). Stylomere triangular, wide, apex rather wide, obliquely rounded off, with

2-3 subapical setae. Lateral plate rather elongate, with 2 apical setae.

105

= =

Figs 113-114. Head and pronotum. 113. Adelotopus rufescens, spec. nov. 114. Adelotopus flavus, spec. nov.

Variation. Apart from some differences in relative width of pronotum and elytra little variation

noted.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. So far collected from end of December to beginning of February.

Distribution (Fig. 607). Kimberley Division, northwestern Australia.

Material examined (3). Only the type series.

Etymology. The name refers to the light colour.

Adelotopus flavus, spec. nov.

Figs 114, 115, 302, 454, 607

Types. Holotype: 3, NT, Mataranka, V-8-83. J. T. Doyen, Adelotopus sp. (not in MCZ) det. J. Liebherr 1987

(CUIC).

Diagnosis. Rather small, fairly wide, depressed, dull, yellowish species with explanate, rounded

margin of pronotum. Further distinguished from relative species by narrow base of pronotum and the

narrow, rather symmetric aedeagus.

Description

Measurements. Length: 5.5 mm. Ratios. Width/length of pronotum: 1.83 ; width base/apex of

pronotum: 1.58; width pronotum/head: 1.74; length/width of elytra: 1.30; length elytra/pronotum:

2.50.

Colour. Head and centre of pronotum light reddish, margins of pronotum, elytra, lower surface,

mouth parts, antennae, and legs yellowish. In apical third of elytra a vague, slightly darker transverse

spot caused by the translucent folded wings.

Head (Figs 115a-d). Short and wide, rather depressed. Anterior border gently convex, lateral angle

rounded, laterally slightly projecting, lateral borders oblique. Clypeal suture semicircular, in middle

interrupted. Labrum narrow, apex concave. Antennal groove laterally sharply bordered, latero-

posteriorly with carinate area. Mental tooth triangular, short, apex acute. Wings of mentum wide,

laterally rounded, apex rectangular. Glossa wide, tongue-like, apically widely rounded, ventrally with

distinct keel, at border with c. 10 elongate setae and additional pilosity on upper and lower surface and

along border. Terminal palpomere of maxillary palpus fairly wide, gently securiform. Terminal

palpomere of labial palpus wide, securiform. Antenna moderately short, Sth-9th antennomeres

slightly <1.5 x as wide as long. Microreticulation fine, though distinct, slightly coriaceous, punctura-

tion rather sparse, mderately fine, fairly difficult to detect within microreticulation, surface with a

106

Figs 115a-k. Adelotopus flavus, spec. nov. Details of head and J genitalia. For legends see fig. 100.

shallow sulcus medially of eyes, rather densely pilose, fairly dull. Ventrolaterally of eyes with a row

of short setae. Suborbital field punctate and setose. Both palpi rather sparsely setose, gula almost

asetose.

Pronotum (Figs 114, 302). Rather wide, moderately depressed, base rather wide, apex fairly

narrow. Apical angles produced, acute, attaining posterior third of eyes. Apex deeply, rectangularly

excised, faintly convex in excision, faintly though distinctly bordered. Sides strongly curved through-

out, widest in basal third. Sides unbordered, explanate, slightly channeled. Basal angles rounded off,

barely produced backwards. Base laterally barely concave, almost straight, faintly though distinctly

bordered. Surface near base with a very shallow transverse impression. Microreticulation fine, though

distinct, isodiametric to somewhat irregular, rather coriaceous, puncturation rather sparse, moderately

fine, difficult to detect within microreticulation, surface with some faint irregular strioles, somewhat

coriaceous, densely pilose, dull.

Elytra (Figs 302, 454). Moderately elongate, depressed, in basal two thirds parallel, then gently

narrowed near apex. Apex rather wide, faintly oblique, truncature feebly convex, apical angles shortly

rounded. Shoulders rather distinct, basal margin slightly oblique, without setae behind shoulders.

Marginal channel rather narrow, completely visible from above. Basal border incomplete, reaching

only to middle of base, ending abruptly. Lateral border asetose. Series of umbilical pores consisting of

5 or 6 pores behind shoulder. Setae very short. Striae including sutural stria absent. Microreticulation

fine, though distinct, isodiametric to slightly transverse, not coriaceous, puncturation fairly coarse,

moderately dense, punctures somewhat rasp-like. Surface densely pilose, rather dull.

Lower surface. Prosternal process rather elongate, narrow, straight, depressed, apex narrow,

straight, faintly rounded off, shortly setose. Metepisternum moderately elongate, c. 1.8 x as long as

wide, posteriorly not constricted nor hollowed. Abdominal sterna including sternum VI apparently

without setae at apical border. Lower surface densely punctate and pilose.

Legs. Elongate, 1st tarsomere of protarsus slightly longer than wide, tibial groove of profemur

moderately deep, anterior plate only at apex distinctly overlapping the groove, posterior border of

groove sharp. Femur moderately wide. Metatibia narrow and elongate, almost 6 x as long as wide, 1st

tarsomere of metatarsus >2.5 x as long as wide. d protarsus not widened.

S genitalia (Figs 115e-k). Genital ring rather narrow and elongate, triangular, slightly asymetric,

with narrow base. Sternum VII moderately wide, apically rather deeply excised, basally bisinuate,

lateral parts small. Aedeagus medium-sized, rather depressed, narrow, rather symmetric. Lower

surface almost straight. Apex rather wide, rounded off. Orifice elongate, internal sac fairly complex,

with a small fold near apex. Both parameres rather narrow and elongate, both with rounded apex, left

considerably larger than right.

? genitalia. Unknown.

107

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Unknown. Holotype collected in May.

Distribution (Fig. 607). Northern Part of Northern Territory. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the yellow colour.

Adelotopus piceus, spec. nov.

Figs 116, 303, 455, 608

Types. Holotype: d, WAM Goldfields Surv. 12/1981 Bungalbin Hill, W. F. Humphreys et al. BHR 2 E.

Salmonophloia woodland, 30°18°S. 119°43’E pit Fall(s) (ANIC).

Diagnosis. Medium-sized, fairly elongate, depressed, dull, reddish-piceous species with darker head

and lighter margins of pronotum and elytra, with explanate, rounded margin of pronotum. Further

distinguished from relative species by obtuse basal angles of pronotum, and fairly wide, somewhat

widened aedeagus.

Description

Measurements. Length: 5.9 mm. Ratios. Width/length of pronotum: 1.80; width base/apex of

pronotum: 1.63; width pronotum/head: 1.71; length/width of elytra: c. 1.40; length elytra/pronotum:

2.60.

Colour. Head piceous, pronotum and elytra reddish-piceous, margin of pronotum and lateral

border of elytra reddish translucent. Lower surface, mouth parts, antennae, and legs dark reddish.

Head (Figs 116a-d). Short and rather wide, rather depressed. Anterior border gently convex,

lateral angle rounded, laterally slightly projecting, lateral borders oblique. Clypeal suture semicircular,

in middle interrupted. Labrum narrow, apex concave. Antennal groove laterally bordered, latero-

posteriorly with carinate area. Mental tooth triangular, short, apex acute. Wings of mentum wide,

laterally rounded, apex rectangular. Glossa wide, tongue-like, apically widely rounded, ventrally with

distinct keel, at border with c. 10 elongate setae and additional pilosity on upper and lower surface and

along border. Terminal palpomere of maxillary palpus fairly wide, gently securiform. Terminal

palpomere of labial palpus very wide, markedly securiform. Antenna moderately short, Sth-9th

antennomeres slightly <1.5 x as wide as long. Microreticulation fine, though distinct, isodiametric,

puncturation rather sparse, very fine, rather difficult to detect, surface with a shallow sulcus medially

of eyes, impilose, moderately dull. Ventrolaterally of eyes with a row of short setae. Suborbital field

almost impunctate and asetose. Both palpi rather sparsely setose, gula almost asetose.

Pronotum (Fig. 303). Rather wide, moderately depressed, base fairly wide, apex rather narrow.

Apical angles rather produced, fairly acute, slightly surpassing posterior boder of eyes. Apex deeply,

rectangularly excised, rather convex in excision, unbordered. Sides strongly curved throughout,

widest in basal third. Margins unbordered, fairly explanate, slightly channeled. Basal angles rounded

off, barely produced backwards. Base slightly concave, laterally faintly bordered. Surface near base

with a very shallow transverse impression. Microreticulation fine, though distinct, isodiametric,

slightly coriaceous, puncturation moderately dense and fine, rather distinct, surface with several faint

irregular strioles, slightly coriaceous, impilose, rather dull.

Elytra (Figs 303, 455). Fairly elongate, depressed, in basal two thirds parallel, then gently

narrowed to apex. Apex rather wide, faintly oblique, truncature faintly concave, apical angles rounded

off. Shoulders rather distinct, basal margin slightly oblique, without setae behind shoulders. Marginal

channel rather narrow, completely visible from above. Basal border incomplete, reaching only to

middle of base, ending abruptly. Lateral border asetose. Series of umbilical pores consisting of 6 pores

behind shoulder. Setae very short. Striae including sutural stria absent. Microreticulation rather fine,

distinct, isodiametric, not coriaceous, more conspicuous than on fore body, puncturation distinct,

moderately dense, apically punctures slightly rasp-like. Surface sparsely pilose, rather dull.

Lower surface. Prosternal process rather elongate, narrow, straight, depressed, apex narrow,

108

Figs 116a-k. Adelotopus piceus, spec. nov. Details of head and d genitalia. For legends see fig. 100.

straight, faintly rounded off, shortly setose. Metepisternum moderately elongate, slightly <2 x as long

as wide, posteriorly not constricted nor hollowed. Abdominal sterna including sternum VI apparently

without setae at apical border. Lower surface rather densely punctate and pilose.

Legs. Fairly elongate, 1st tarsomere of protarsus c. as long as wide, tibial groove of profemur deep,

anterior plate distinctly overlapping the groove for some distance from apex, posterior border of

groove very sharp. Femur moderately wide. Metatibia rather elongate, >5 x as long as wide, Ist

tarsomere of metatarsus >2 x as long as wide. d protarsus not widened.

d genitalia (Figs 116e-k). Genital ring rather narrow, slightly asymetric, with narrow, deeply

excised base. Sternum VII moderately wide, apically deeply excised, basally almost straight, lateral

parts short. Aedeagus medium-sized, rather depressed, in middle moderately widened, slightly

asymmetric. Lower surface gently convex. Apex wide, rounded off. Orifice rather short, internal sac

fairly complex, with a small fold near apex. Both parameres large and wide, with widely rounded apex,

left considerably larger than right, right short.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Holotype collected in December in pit fall trap in Salmonophloia woodland.

Distribution (Fig. 608). Interior of southwestern Australia. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the reddish-piceous color.

Adelotopus longus, spec. nov.

This species occurs in a southern and a northern subspecies that vary by minor differences in shape

of pronotum and elytra, degree of microsculpture, and shape of aedeagus and parameres.

Diagnosis. Medium-sized, elongate, fairly convex, glossy, reddish species with moderately explanate,

rectangular, posteriorly not produced basal angles of pronotum. Further distinguished from relative

species by reduction of microreticulation and convex apex of aedeagus.

109

Adelotopus longus longus, subspec. nov.

Figs 117, 304, 456, 608

Types. Holotype: d, Biggenden, O. X11.1973, H. Frauca (ANIC). - Paratypes: 359,722, same data (ANIC, CBM);

334, 222, Biggenden Old, 5.Jan 1972, H. Frauca (ANIC); 1%, Australia, Nanango Q. S., 6.XIl.86, K. Houston

(CMC); 12, 15 km n. Goomeri, s. Qld., Australien, 23.1.1982, M. Baehr (CBM); 17, 20 km n. Biggenden, s. Qld.,

Australien, 22.1.1982, M. Baehr (CBM); 1%, 122, castaneus Masters, castaneus, Gayndah of Aust. Mus 1.3.04

(ANIC); 338, Aus. Q., 16.X1.86, Gayndah, V.R. Bejsak, legt. (CBS, MMS); 15, Australien, Qld G 38, Raglan Ck.,

10km nw. Mt. Larcom, 21.11.1990, leg. R. Gerstmeier (CBM); 18, Biloela 3.12.46, A. R. Bird (UOIC);

18, Rockhampton, det. castaneus (SAMA); 14, Rock. J. Sedlacek Collector (CSB); 13, 47??, Australia: Qld

Rockhampton 26-27.X1.1967, J. & M. Sedlacek Collectors BISHOP (BMH); 19, Australia: Q. Edungalba 19,

]J. Sedlacek Collector (CSB); 14, Queensland: Roma, 20.1.1978, J. Sedlacek Collector (CSB); 14, Australien, Qld 27,

Rolf Ck., 134km n. Dingo, Fitzroy Dev. Rd., 12.11.1990, M. Baehr (CBM); 1?, Peak Downs, Q., rec. from

W. du Boulay (NMV); 15, 80501, Goedffroy Collection, Adelotopus castaneus (Casteln.) Peak Downs, Queens.

(NMV); 12, 15, Queensland dedit Mac Leay 1877, Adelotopus castaneus Casteln. Queensland D. Macl. 1877, nec

castaneus Cast. det. Gestro (MCSN); 17, Queensland W. M° Leay, castaneus Casteln., Ex Musaeo Chaudoir

(MNHN); 22%, Nov. Holl., 10 518, det. castaneus (NHRS); 222, W. Australia, C. French’s Coll. 5.11.08, Adelotopus

castaneus (Castl.) W. Australia, brunneus Cast. ?, A. castaneus (NMV).

Diagnosis. Distinguished from northern subspecies A. longus tropicus by narrower pronotum and

elytra, convex rather than concave basal border of pronotum, less dense puncturation on pronotum

and elytra, narrower aedeagus, and narrower parameres.

Description

Measurements. Length: 5.6-6.5 mm. Ratios. Width/length of pronotum: 1.63-1.69; width base/

apex of pronotum: 1.48-1.55; width pronotum/ head: 1.57-1.65; length/ width of elytra: 1.51-1.55; length

elytra/pronotum: 2.55-2.69.

Colour. Reddish, head and centre of pronotum slightly darker. Lower surface, mouth parts,

antennae, and legs reddish.

Head (Figs 117a-d). Short and rather wide, moderately depressed. Anterior border gently convex,

lateral angle widely rounded off, laterally slightly projecting, lateral borders oblique. Clypeal suture

semicircular, in middle interrupted. Labrum rather narrow, apex concave. Antennal groove laterally

sharply bordered, latero-posteriorly with carinate area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically

widely rounded, ventrally with distinct keel, at border with c. 10 elongate setae and additional pilosity

on upper and lower surface and along border. Terminal palpomere of maxillary palpus fairly wide,

gently securiform. Terminal palpomere of labial palpus very wide, markedly securiform. Antenna

rather short, 8th-9th antennomeres slightly <2 x as wide as long. Microreticulation extremely fine,

highly superficial, almost invisible, puncturation fine, rather dense, distinct, surface with a shallow

sulcus medially of eyes and some faint strioles, impilose, glossy. Ventrolaterally of eyes with a row of

short setae. Suborbital field finely punctate and setose. Both palpi rather sparsely setose, gula almost

asetose.

Pronotum (Fig. 304). Moderately wide, moderately convex, base fairly wide, apex rather narrow.

Apical angles rather produced, fairly acute, slightly surpassing posterior border of eyes. Apex deeply,

rectangularly excised, rather convex in excision, laterally faintly bordered. Sides moderately curved,

almost straight in front of basal angles, widest in basal third. Margins unbordered, fairly explanate,

slightly channeled. Basal angles almost rectangular, at apex obtuse, barely produced backwards. Base

slightly convex, laterally faintly bordered. Surface near base with or without a very shallow transverse

impression. Microreticulation extremely superficial, almost invisible, puncturation moderately dense

and fine, distinct, surface with several faint irregular strioles, sometimes fairly coriaceous, very

sparsely pilose, moderately to fairly glossy.

Elytra (Figs 304, 456). Elongate, moderately convex, in basal two thirds markedly parallel, then

gently narrowed to apex. Apex rather wide, transverse, truncature convex, apical angles rounded off.

Shoulders rather distinct, basal margin slightly oblique, without setae behind shoulders. Marginal

channel rather narrow, completely visible from above. Basal border incomplete, reaching only to

middle of base, ending abruptly. Lateral border asetose. Series of umbilical pores consisting of 6 pores

behind shoulder. Setae very short. Striae including sutural stria absent. Microreticulation extremely

110

Figs 117a-l. Adelotopus longus longus, spec. nov. Details of head and genitalia. For legends see fig. 100.

superficial, barely visible even under high magnification, puncturation distinct, moderately dense.

Surface sparsely pilose, glossy.

Lower surface. Prosternal process rather elongate, narrow, straight, depressed, apex narrow,

straight, faintly rounded off, shortly setose. Metepisternum rather elongate, c. 2x as long as wide,

posteriorly not constricted nor hollowed. Abdominal sterna including sternum VI apparently without

setae at apical border. Lower surface rather densely punctate and pilose.

Legs. Fairly elongate, Ist tarsomere of protarsus c. as long as wide, tibial groove of profemur deep,

anterior plate distinctly overlapping the groove for some distance from apex, posterior border of

groove sharp. Femur moderately wide. Metatibia rather elongate, >5 x as long as wide, 1st tarsomere

of metatarsus >2 x as long as wide. d protarsus not widened.

d genitalia (Figs 117e-k). Genital ring rather narrow, slightly asymetric, arms evenly convex, with

rather narrow, excised base. Sternum VII moderately wide, apically deeply excised, basally bisinuate,

lateral parts short. Aedeagus medium-sized, rather depressed, in middle slightly widened, slightly

asymmetric. Lower surface faintly bisinuate. Apex wide, rounded off. Orifice rather short, internal sac

fairly complex, with a small fold near apex. Both parameres rather elongate, with widely rounded

apex, left considerably larger than right.

? genitalia (Fig. 1171). Stylomere triangular, moderately wide, apex rather wide, obliquely

rounded off, with 2-3 subapical setae. Lateral plate rather elongate, with 2 apical setae.

Variation. Little variation of relative length of pronotum and elytra and of distinctness of

microsculpture noted.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. Specimens collected by me under bark of different gum-like eucalypts. So

far captured from November to January, and in March.

Distribution (Fig. 608). Southeastern Queensland north to somewhat north of Rockhampton. Two old

specimens from “W. Australia” certainly wrongly labelled.

Material examined (43). Only the type series.

Etymology. The name refers to the elongate shape.

Figs 118b-c, e-k. Adelotopus longus tropicus, subspec. nov. Details of head and d genitalia. For legends see fig.

100.

Adelotopus longus tropicus, subspec. nov.

Figs 118, 305, 457, 608

Types. Holotype: d, Charters Towers Old, 13.X11.1976, B. B. Lowery (ANIC). - Paratypes: 1d, Australia, Old.

17-20 mi W. Atherton, 1.11.1975, H. & A. Howden (ANIC); 1%, 3.7 km E Chillagoe, 13-V-1987, P. A. Meyer coll.,

Under bark of gum-type eucalypt (CBM); 17, Australia, Qld 93/67, Einasleigh R. b. Carpentaria Downs,

12.-13.6.1993, M. Baehr (CBM); 258, Australia, Qld 93/64, Einasleigh R. 2km e. Einasleigh, 11.-12.6.1993,

M. Baehr (CBM); 15, Mareeba: NQ Davies Ck. Rd, 22.Feb.1972, A. & M. Walford-Huggins 5856 (CMP-WHC);

15, M' Molloy Queensland, F. H. Taylor, Adelotopus bimaculatus Macl. (ANIC); 18, Cooktown, N.Q., May 1951,

C. Oke (NMV).

Diagnosis. Distinguished from southern subspecies A. longus longus by wider pronotum and elytra,

concave rather than produced basal border of pronotum, denser puncturation on pronotum and elytra,

wider aedeagus, and wider parameres.

Description

Measurements. Length: 5.6-6.5 mm. Ratios. Width/length of pronotum: 1.66-1.74; width base/

apex of pronotum: 1.58-1.62; width pronotum/ head: 1.69-1.74; length/ width of elytra: 1.45-1.53; length

elytra/pronotum: 2.45-2.54.

Colour. Similar to nominate subspecies.

Head (Figs 118b,c). Similar to nominate subspecies, though palpi slightly narrower, puncturation

even denser, and microreticulation usually even less distinct.

Pronotum (Fig. 305). Generally similar to nominate subspecies, though comparatively wider,

laterally more evenly curved, especially near base, basal angle more produced backwards, and base

perceptible concave. Surface with denser puncturation, microreticulatuion almost invisible.

Elytra (Figs 305, 456). Generally similar to nominate subspecies, though elytra slightly shorter and

wider, less parallel, Microreticulation even less distinct, almost invisible, and puncturation slightly

coarser and denser, hence surface even faintly glossier.

Lower surface. Similar to nominate subspecies.

Legs. Similar to nominate subspecies.

d genitalia (Figs e-k). Largely similar to nominate subspecies, but genital ring wider, aedeagus

slightly wider, and both parameres also wider, at apex more widely rounded off.

? genitalia. Similar to nominate subspecies.

Variation. Generally little variation noted in relative shape of pronotum and elytra and degree of

puncturation of surface, but there is one specimen from Einasleigh River, North Queensland, collected

together with two normal-shaped specimens that has at the same time a wider pronotum, considerably

shorter elytra, a remarkably wide genital ring, an unusually short and wide aedeagus with widely

rounded apex, and narrower and more elongate parameres (Figs ). More material is required to decide,

whether this specimen represents a nomenclatorial valuable taxon, e.g a subspecies or even an

independent species, or is merely an example of unusual strong variation.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. Specimens collected by me under bark of river gums near large river, a

specimen collected by P. Meyer “under bark of gum-type eucalypt”. So far captured in February, May,

June, and December.

Distribution (Fig. 608). Northeastern Queensland from about Townsville to Cooktown.

Material examined (10). The type series and one additional specimen that has been not included in the type

series because of certain remarkable structural differences: 13, Australia, Qld 93/64, Einasleigh R. 2km

e. Einasleigh, 11.-12.6.1993, M. Baehr (CBM).

Etymology. The name refers to the northern distribution of this subspecies.

Adelotopus sinuaticollis, spec. nov.

This species occurs in a southern and a more northern subspecies that vary by minor differences in

shape of pronotum and elytra, degree of microsculpture, and shape of ? stylomeres.

Diagnosis. Medium-sized, rather elongate, fairly convex, more or less dull, rusty reddish species with

moderately explanate, rectangular, distinctly produced margin of pronotum. Further distinguished

from relative species by presence of microreticulation and acute apex of aedeagus.

Adelotopus sinuaticollis sinuaticollis, subspec. nov.

Figs 119, 306, 458, 609

Types. Holotype: d, Sea Lake Goudie, E. W. Ferguson Collection, Adelotopus (ANIC). — Paratypes: 17, Adelaide

Coll. Castelnau, det. aphodioides (MCSN) [marked “s”, mounted with 2 paratypes of s. sinuaticollis, spec. nov. on

same card]; 27? 2, Lea’s, Adelotopus sp. ?, Id. by T. G. Sloane, Sea Lake Goudie (SAMA); 15, 17, Sea Lake Goudie,

Adelotopus aphodioides Westw, J C Goudie Collection (NMV); 13, 22%, Woorinen. V., 5.1.36, F. E. Wilson,

F. E. Wilson Collection, Host Iridomyrmex detectus (NMV); 484, 22%, Morilla N. S. W. (ANIC, CBM); 18, 499,

Australia N. S. W., M. F. L. (BMNH, CBM); 1%, 1 (sex?), castaneus Cast., Gayndah, Adelotopus castaneus Cast.

Queensland 3889 (SAMA); 18, Aust. (OUM); 244, K. 12375, Adelotopus castaneus Casteln. Gayndah (AMS);

15, E. W. Ferguson Collection (ANIC).

Diagnosis. Distinguished from northern subspecies A. sinuaticollis calliope by slightly wider pronotum

and elytra, coarse instead of superficial microreticulation of pronotum, coarser puncturation on

pronotum and elytra, and wider $ stylomere with wider apex.

Description

Measurements. Length: 5.70-6.55 mm. Ratios. Width/length of pronotum: 1.64-1.72; width base/

apex of pronotum: 1.53-1.60; width pronotum/ head: 1.66-1.69; length/ width of elytra: 1.38-1.45; length

elytra/pronotum: 2.36-2.40.

Colour. Upper and lower surface, mouth parts, antenna, and legs rusty red, sometimes head

faintly darker.

Head (Figs 119a-d). Short and rather wide, moderately depressed. Anterior border gently convex,

lateral angle widely rounded off, laterally slightly projecting, lateral borders oblique. Clypeal suture

semicircular, in middle interrupted. Labrum rather narrow, apex concave. Antennal groove laterally

sharply bordered, latero-posteriorly with carinate area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically

widely rounded, ventrally with distinct keel, at border with c. 10 elongate setae and additional pilosity

on upper and lower surface and along border. Terminal palpomere of maxillary palpus wide,

markedly securiform. Terminal palpomere of labial palpus very wide, extremely securiform. Antenna

Figs 119a-l. Adelotopus sinuaticollis sinuaticollis, spec. nov. Details of head and genitalia.

Fig. 1201. Adelotopus sinuaticollis calliope, subspec. nov. Details of ? genitalia. For legends see fig. 100.

moderately short, Sth-9th antennomeres c. 1.5 x as wide as long or faintly wider. Microreticulation

fine, though distinct, puncturation fine, rather dense, more or less easily visible, surface with a shallow

sulcus medially of eyes and some faint strioles, impilose, rather dull. Ventrolaterally of eyes with a row

of short setae. Suborbital field almost impunctate and asetose. Both palpi rather sparsely setose, gula

almost asetose.

Pronotum (Fig. 306). Moderately wide, moderately convex, base fairly wide, apex rather narrow.

Apical angles rather produced, fairly acute, slightly surpassing posterior boder of eyes. Apex deeply,

rectangularly excised, rather convex in excision, laterally faintly bordered. Sides rather curved

throughout, widest in basal third. Margins unbordered, fairly explanate. Basal angles almost rectan-

gular, at apex barely obtuse, distinctly produced backwards. Base laterally distinctly sinuate, in middle

slightly produced, laterally faintly bordered. Surface near base without transverse impression.

Microreticulation fine though very distinct, sometimes even coarse, puncturation moderately dense

and fine, more or less distinct, surface with rather dense network of irregular strioles, usually distinctly

coriaceous, impilose, dull.

Elytra (Figs 306, 458). Rather elongate, moderately convex, in basal two thirds almost parallel, then

gently narrowed to apex. Apex rather wide, transverse, truncature convex, apical angles rounded off.

Shoulders rather distinct, basal margin slightly oblique, without setae behind shoulders. Marginal

channel rather narrow, completely visible from above. Basal border incomplete, reaching only to

middle of base, ending abruptly. Lateral border asetose. Series of umbilical pores consisting of 6

(sometimes unilaterally 7 or even 8) pores behind shoulder. Setae very short. Striae including sutural

stria absent. Microreticulation fine, isodiametric, fairly distinct, though much more superficial than on

fore body, puncturation distinct, moderately dense. Surface impilose, moderately glossy.

Lower surface. Prosternal process rather elongate, narrow, straight, depressed, apex narrow,

114

straight, faintly rounded off, shortly setose. Metepisternum rather elongate, slightly <2 x as long as

wide, posteriorly not constricted nor hollowed. Abdominal sterna including sternum VI apparently

without setae at apical border. Lower surface rather densely punctate and pilose.

Legs. Rather elongate, Ist tarsomere of protarsus slightly longer than wide, tibial groove of

profemur deep, anterior plate distinctly overlapping the groove for some distance from apex, posterior

border of groove sharp. Femur moderately wide. Metatibia elongate, almost 6 x as long as wide, Ist

tarsomere of metatarsus >2 x as long as wide. d protarsus not widened.

d genitalia (Figs 119e-k). Genital ring moderately wide, slightly asymmetric, arms evenly convex,

with rather narrow base. Sternum VII moderately wide, apically deeply excised, basally gently

triangular, lateral parts rather short. Aedeagus medium-sized, rather depressed, in middle widened,

almost symmetric. Lower surface almost straight. Apex narrow, obtuse. Orifice rather short, internal

sac fairly complex, with a small fold near apex. Both parameres large, with fairly rounded apex, left

considerably larger than right. Both parameres with a distinct edge in upper quarter of outer surface.

? genitalia (Fig. 1191). Stylomere triangular, moderately wide, apex wide, rounded off, with 2-3

subapical setae. Lateral plate rather elongate, with 2-3 apical setae.

Variation. Little variation noted in relative shape of pronotum and elytra.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Unknown. This species is exclusively known from old specimens, no specimen is dated and

the locality labels are rather inaccurate.

Distribution (Fig. 609). Southeastern Australia from eastern South Australia through Victoria, eastern

New South Wales, and southeastern Queensland north to Gayndah.

Material examined (26). Only the type series.

Etymology. The name refers to the sinuate basal margin of the pronotum.

Adelotopus sinuaticollis calliope, subspec. nov.

Figs 120, 307, 459, 609

Types. Holotype: 17, Australien, Old 50, Calliope River 27 km se. Mt. Larcom, 20.11.1990, M. Baehr (ANIC).

Diagnosis. Distinguished from southern nominate subspecies A. sinuaticollis sinuaticollis by slightly

narrower pronotum and elytra, superficial instead of coarse microreticulation of pronotum, finer

puncturation on pronotum and elytra, and narrower stylomere with narrower apex.

Description

Measurements. Length: 6.55 mm. Ratios. Width/length of pronotum: 1.62; width base/apex of

pronotum: 1.51; width pronotum/head: 1.63; length/width of elytra: 1.46; length elytra/ pronotum:

2.41.

Colour. Reddish, though head and centre of pronotum perceptibly darker, piceous.

Head. Generally similar to nominate subspecies, though microreticulation and puncturation finer.

Pronotum (Fig. 307). Similar to nominate subspecies in shape, though apical and basal margin

with more distinct borders. Microreticulation fine and rather superficial. Puncturation finer, though

more easily visible due to faint microreticulation and to much more superficial striolation. Surface not

at all coriaceous, but moderately glossy.

Elytra (Figs 307, 459). Similar to nominate subspecies, though microreticulation more superficial

and puncturation perceptibly finer, hence surface fairly glossy. In the single specimen elytra with 6

umbilical pores on both sides.

Lower surface. Similar to nominate subspecies.

Legs. Similar to nominate subspecies, though posterior legs even longer.

d genitalia. Unknown.

? genitalia (Fig. 1201). Stylomere triangular, rather narrow, apex narrow, rounded off, with 2-3

subapical setae. Lateral plate rather elongate, with a longer and a shorter apical seta.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Holotype collected under bark of large river eucalypt, together with several other Adelotopus

species.

Distribution (Fig. 609). Central eastern Queensland near Gladstone. Known only from type locality.

Occurs almost 200 km north of the northernmost known locality of the nominate subspecies.

Material examined (1). Only the holotype.

Etymology. The name refers to the type locality, the Calliope River.

Adelotopus bamagae, spec. nov.

Figs 121, 308, 460, 608

Types. Holotype: d, Bamaga N. Cape York, Q Jan.’58 Darlingtons (MCZ).

Diagnosis. Medium-sized, elongate, fairly depressed, dull, black species with moderately explanate,

obtusely rectangular, faintly produced margin of pronotum. Further distinguished from relative

species by slightly rasp-like elytral punctures and rather densely pilose surface.

Description

Measurements. Length: 6.65 mm. Ratios. Width/length of pronotum: 1.62; width base/apex of

pronotum: 1.59; width pronotum/head: 1.67; length/width of elytra: 1.59; length elytra/pronotum:

2.56.

Colour. Dull black, lateral margin of pronotum feebly reddish translucent. Lower surface reddish-

piceous. Mouth parts, antennae, and legs dark reddish to reddish-piceous.

Head (Figs 121a-d). Short and rather wide, moderately depressed. Anterior border gently convex,

lateral angle widely rounded off, laterally slightly projecting, lateral borders oblique. Clypeal suture

semicircular, in middle interrupted. Labrum rather narrow, apex concave. Antennal groove laterally

sharply bordered, latero-posteriorly with carinate area. Mental tooth triangular, short, apex acute.

Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically widely

rounded, ventrally with distinct keel, at border with c. 10 elongate setae and additional pilosity on

upper and lower surface and along border. Terminal palpomere of maxillary palpus rather wide,

slightly securiform. Terminal palpomere of labial palpus very wide, markedly securiform. Antenna

short, 8th-9th antennomeres c. 2x as wide as long. Microreticulation very fine, fairly distinct,

puncturation very fine, rather dense, though easily visible, surface with a shallow sulcus medially of

eyes and some faint strioles, sparsely pilose, rather dull, somewhat silky. Ventrolaterally of eyes with

a row of short setae. Suborbital field at least laterally faintly punctate and setose. Both palpi rather

sparsely setose, gula almost asetose.

Pronotum (Fig. 308). Moderately wide, moderately convex, base fairly wide, apex rather narrow.

Apical angles rather produced, fairly acute, slightly surpassing posterior border of eyes. Apex deeply,

rectangularly excised, rather convex in excision, laterally faintly bordered. Sides rather curved

throughout, widest in basal third. Margins faintly bordered, fairly explanate. Basal angles c. 100°, at

apex obtuse, feebly produced backwards. Base laterally faintly sinuate, in middle slightly produced,

faintly bordered. Surface near base with a shallow, transverse impression. Microreticulation fine

though rather distinct, puncturation dense, distinct, laterally less dense, though somehwat rasp-like,

surface with several fine, irregular strioles, rather densely pilose, rather dull, somewhat silky.

Elytra (Figs 308, 460). Elongate, moderately convex, on disk rather depressed, in basal two thirds

almost parallel, then gently narrowed to apex. Apex rather wide, transverse, truncature convex, apical

angles rounded off. Shoulders rather distinct, basal margin slightly oblique, without setae behind

shoulders. Marginal channel narrow, completely visible from above. Basal border incomplete, reach-

ing only to middle of base, ending abruptly. Lateral border asetose. Series of umbilical pores consisting

of 6 pores behind shoulder. Setae very short. Striae including sutural stria absent. Microreticulation

fine, isodiametric, fairly distinct, somewhat superficial, puncturation rather fine though very distinct,

rather sparse, punctures distinctly rasp-like. Surface densely pilose, moderately glossy, somewhat

silky.

Lower surface. Prosternal process rather elongate, narrow, straight, behind coxae almost keeled,

apex narrow, straight, faintly rounded off, shortly setose. Metepisternum elongate, c. 2.2 x as long as

116

Figs 121a-k. Adelotopus bamagae, spec. nov. Details of head and genitalia. For legends see fig. 100.

wide, posteriorly not constricted nor hollowed. Abdominal sterna including sternum VI apparently

without setae at apical border. Lower surface rather densely punctate and pilose.

Legs. Rather elongate, 1st tarsomere of protarsus about as long as wide, tibial groove of profemur

deep, anterior plate distinctly overlapping the groove for some distance from apex, posterior border

of groove sharp. Femur moderately wide. Metatibia fairly elongate, c. 5x as long as wide, Ist

tarsomere of metatarsus slightly >2 x as long as wide. d protarsus not widened.

d genitalia (Figs 121e-k). Genital ring fairly wide, rather asymmetric, arms evenly convex, with

rather narrow base. Sternum VII rather wide, apically convex, rather deeply excised, basally bisinuate,

lateral parts rather short. Aedeagus medium-sized, moderately depressed, in middle fairly widened,

slightly asymmetric. Lower surface straight. Apex wide, widely rounded. Orifice rather short, internal

sac fairly complex. Both parameres rather narrow, with obtusely rounded apex, left considerably larger

than right.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Unknown. Holotype collected in January.

Distribution (Fig. 608). North Queensland, extreme tip of Cape York Peninsula. Known only from

type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the type locality.

Adelotopus edithae, spec. nov.

Figs 122, 309, 462, 609

Types. Holotype: d, Australia, NT 95/47,5 km e. Edith River Cr., 27.-28.8.1995, M. Baehr (MNTD). - Paratypes:

238, same data (CBM); 12, 12.405, 132.54E Jabiru NT 5-9 Jul.1977 R. Pengilly, pitfall trap (MNTD); 15, Australia,

NT 95/9, Litchfield NP, 30 km w. Batchelor, 6.8.1995, M. Baehr (CBM).

Diagnosis. Medium-sized, rather short, convex, dull piceous- black species with barely explanate,

obtusely rectangular margin of pronotum. Further distinguished from relative species by extremely

fine, slightly rasp-like elytral puncturation, densely, though extremely shortly pilose surface, laterally

strongly bordered and near apex excised prosternal process, and moderately wide, somewhat shoe-

like aedeagus with rounded apex.

117

i k

Figs 122a-l. Adelotopus edithae, spec. nov. Details of head and genitalia. For legends see fig. 100.

Description

Measurements. Length: 4.80-6.15 mm. Ratios. Width/length of pronotum: 1.76-1.78; width base/

apex of pronotum: 1.57-1.64; width pronotum/head: 1.66-1.72; length/ width of elytra: 1.26-1.35; length

elytra/pronotum: 2.27-2.42.

Colour. Dull piceous-black, lateral margins of pronotum and elytra indistinctly dark reddish

translucent. Lower surface dark piceous. Mouth parts, antenna, and legs dark reddish-piceous.

Head (Figs 122a-d). Short and rather wide, moderately depressed. Anterior border gently convex,

lateral angle widely rounded off, laterally slightly projecting, lateral borders oblique. Clypeal suture

semicircular, in middle widely interrupted. Labrum rather narrow, apex concave. Antennal groove

laterally sharply bordered, latero-posteriorly with carinate area. Mental tooth triangular, rather short,

apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like,

apically widely rounded, ventrally with distinct keel, at border with c. 10 elongate setae and additional

pilosity on upper and lower surface and along border. Terminal palpomere of maxillary palpus

moderately wide, slightly securiform. Terminal palpomere of labial palpus very wide, markedly

securiform. Antenna short, 8th-9th antennomeres almost 2 x as wide as long. Microreticulation very

fine, though distinct, regular, puncturation extremely fine, rather sparse, surface with a shallow sulcus

medially of eyes only, apparently impilose, very smooth, though fairly dull. Ventrolaterally of eyes

with a row of short setae. Suborbital field at least laterally faintly punctate and setose. Both palpi rather

sparsely setose, gula almost asetose.

Pronotum (Fig. 309). Rather wide, convex, base fairly wide, apex rather narrow. Apical angles

moderately produced, fairly acute, slightly surpassing posterior border of eyes. Apex rather deeply,

rectangularly excised, rather convex in excision, very inconspicuously bordered. Sides curved through-

out, widest in basal third. Margins unbordered, barely explanate. Basal angles c. 100°, at apex obtusely

rounded, feebly produced backwards. Base irregularly concave, in middle faintly produced, very

inconspicuously bordered. Surface near base with a very shallow, transverse impression. Microretic-

ulation fine, though distinct, puncturation fine and rather dense, moderately distinct, surface with

several short, irregular strioles, densely though very shortly pilose, rather dull.

Elytra (Figs 309, 462). Rather short, convex throughout, basally almost parallel, then gently

narrowed to apex. Apex rather wide, slightly oblique, truncature apparently faintly concave, apical

118

angles rounded off. Shoulders rounded off, basal margin slightly oblique, without setae behind

shoulders. Marginal channel very narrow, completely visible from above. Basal border incomplete,

reaching only to middle of base, ending abruptly. Lateral border asetose. Series of umbilical pores

consisting of 6 pores behind shoulder. Setae very short. Striae including sutural stria absent.

Microreticulation very fine, isodiametric, slightly superficial, puncturation very fine though distinct,

rather dense, punctures faintly rasp-like, especially towards apex. Surface rather densely pilose,

moderately glossy, silky, pilosity extremely short.

Lower surface. Prosternal process rather elongate, narrow, straight, anteriorly depressed, laterally

strongly bordered, surface behind coxae remarkably concave, apex narrow, straight, rectangular, very

shortly setose. Metepisternum rather elongate, slightly <2 x as long as wide, posteriorly not constrict-

ed nor hollowed. Abdominal sterna including sternum VI apparently without setae at apical border.

Lower surface rather densely punctate and pilose.

Legs. Moderately elongate, Ist tarsomere of protarsus about as long as wide, tibial groove of

profemur deep, anterior plate distinctly overlapping the groove for some distance from apex, posterior

border of groove sharp. Femur rather wide. Metatibia rather elongate, c. 5x as long as wide, Ist

tarsomere of metatarsus slightly >2 x as long as wide. ö protarsus unknown.

d genitalia (Figs 122e-k). Genital ring fairly wide, highly asymmetric, left arm straight, right

convex, with rather narrow, asymmetric base. Sternum VII rather wide, apically rather deeply excised,

basally markedly bisinuate, lateral parts triagonal, rather elongate. Aedeagus medium-sized, moder-

ately depressed, in middle rather vwidened, asymmetric, shoe-like. Lower surface straight. Apex

wide, rounded. Orifice elongate, internal sac fairly complex. Both parameres rather large, triangular,

with narrowly rounded apex, left considerably larger than right.

? genitalia (Fig. 1221). Stylomere short and wide, markedly triagonal, apex obtuse, with 2-3

subapical setae. Lateral plate elongate, with 3 apical setae.

Variation. Some variation noted in size and relative shape of pronotum.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. Specimens collected by me under bark of gum-type eucalypts (E. alba or

E. bigalerita). Female paratype collected in “pitfall trap”. So far captured in July and August.

Distribution (Fig. 609). Eastern and southern margins of Arnhem Land, northern part of Northern

Territory.

Material examined (5). Only the type series.

Etymology. The name refers to the type locality.

Adelotopus rufozonatus, spec. nov.

Figs 33, 123, 310, 461, 608

Types. Holotype: 3, Fly Ck., 37 km S. of Darwin, N. T. 4.11.1979 R. Piper (QMB T26068). - Paratypes: 234, Port

Darwin N. Territory (MMS); 15, 19, Port Darwin end’09 (ANIC); 13, G. F. Hill, Kodpenyay (?) Darwin,

N. T. (SAMA); 13, Australien, NT, Mary River, 110 km e. Darwin, 1.11.1984, M. & B. Baehr (CBM); 334, 428,

Australia, NT 95/48, 30 km n. Edith River Cr., 28.8.1995, M. Baehr (CBM); 13, Australia, NT 95/11, Edith River

Cr., 60 km n. Katherine, 6.-7.8.1995, M. Baehr (CBM); 538, 222, Australia, NT 95/47, 5 km e. Edith River Cr.,

27.-28.8.1995, M. Baehr (CBM, MNTD, QMB, ZSM); 18,222, Katherine N. T. IV-77, J. Sedlacek Collector (CBM,

CSB).

Diagnosis. Medium-sized, moderately elongate, fairly depressed, black species with broad red band

across basal % of elytra, and moderately explanate, obtusely rectangular, faintly produced margin of

pronotum. Further distinguished from relative species by slightly rasp-like elytral punctures and

rather densely pilose surface.

Description

Measurements. Length: 5.65-7.55 mm. Ratios. Width/length of pronotum: 1.62-1.71; width base/

apex of pronotum: 1.41-1.62; width pronotum/head: 1.55-1.77; length/ width of elytra: 1.51-1.68; length

elytra/pronotum: 2.57-2.78.

Colour (Figs 33, 310). Piceous black to black, margins of pronotum and elytra and basal % to % of

119

Figs 123a-l. Adelotopus rufozonatus, spec. nov. Details of head and genitalia. For legends see fig. 100.

elytra light red. Lower surface reddish, below head and pronotum in parts piceous. Mouth parts,

antenna, and legs reddish to reddish-piceous.

Head (Figs 123a-d). Short and rather wide, moderately depressed. Anterior border gently convex,

lateral angle widely rounded off, laterally slightly projecting, lateral borders oblique. Clypeal suture

semicircular, in middle widely interrupted. Labrum rather narrow, apex concave. Antennal groove

laterally sharply bordered, latero-posteriorly with carinate area. Mental tooth triangular, short, apex

acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically

widely rounded, ventrally with distinct keel, at border with c. 10 elongate setae and additional pilosity

on upper and lower surface and along border. Terminal palpomere of maxillary palpus moderately

wide, slightly securiform. Terminal palpomere of labial palpus very wide, markedly securiform.

Antenna moderately short, Sth-9th antennomeres slightly >1.5 x as wide as long. Microreticulation

very fine, rather superficial, sometimes difficult to detect, puncturation double, very fine and fairly

dense, and coarse, the latter becoming dense towards base, surface with a shallow sulcus medially of

eyesand with more or less distinct, sometimes confluent strioles on vertex, densely pilose, moderately

glossy, though posteriorly becoming more coriaceous. Ventrolaterally of eyes with a row of short setae.

Suborbital field at least laterally faintly punctate and setose. Both palpi rather sparsely setose, gula

almost asetose.

Pronotum (Fig. 310). Moderately wide, moderately convex, base fairly wide, apex rather narrow.

Apical angles moderately produced, fairly acute, slightly surpassing posterior border of eyes. Apex

deeply, rectangularly excised, rather convex in excision, fairly distinctly bordered. Sides rather curved

throughout, widest in basal third. Margins unbordered, more or less widely explanate. Basal angles

c. 100°, at apex obtuse or narrowly rounded off, feebly produced backwards. Base laterally faintly

sinuate, in middle slightly produced, perceptibly bordered. Surface near base with a shallow,

transverse impression. Microreticulation very fine, rather superficial, sometimes difficult to detect,

puncturation dense, distinct, anteriorly very dense, posteriorly becoming sparser, double, coarse and

moderately fine, surface with several short, irregular strioles, apically and on disk rather coriaceous,

densely pilose, moderately glossy.

Elytra (Fig 310, 461). Rather elongate, moderately convex, on disk rather depressed, in basal two

thirds almost parallel, then gently narrowed to apex. Apex rather wide, transverse, truncature convex,

120

apical angles rounded off. Shoulders rounded off, basal margin slightly oblique, without setae behind

shoulders. Marginal channel narrow, completely visible from above. Basal border incomplete, reach-

ing only to middle of base, ending abruptly. Lateral border asetose. Series of umbilical pores consisting

of 6 pores behind shoulder. Setae very short. Striae including sutural stria absent. Microreticulation

fine, isodiametric, very superficial, difficult to detect, puncturation rather fine though very distinct,

moderately dense, punctures distinctly rasp-like. Surface densely pilose, moderately glossy, slightly

silky, pilosity fairly elongate.

Lower surface. Prosternal process rather elongate, narrow, straight, apex narrow, straight, faintly

rounded off, shortly setose. Metepisternum rather elongate, slightly <2 x as long as wide, posteriorly

not constricted nor hollowed. Abdominal sterna including sternum VI apparently without setae at

apical border. Lower surface densely punctate and pilose.

Legs. Rather elongate, 1st tarsomere of protarsus about as long as wide, tibial groove of profemur

deep, anterior plate distinctly overlapping the groove for some distance from apex, posterior border

of groove sharp. Femur moderately wide. Metatibia rather elongate, >5 x as long as wide, 1st

tarsomere of metatarsus slightly >2 x as long as wide. d protarsus not widened.

d genitalia (Figs 123e-k). Genital ring fairly wide, asymmetric, arms evenly convex, with rather

narrow, asymmetric base. Sternum VII rather wide, apically rather deeply excised, basally almost

straight, faintly bisinuate, lateral parts triagonal, rather elongate. Aedeagus medium-sized, moderately

depressed, in middle fairly widened, slightly asymmetric. Lower surface straight. Apex wide, widely

rounded. Orifice rather short, internal sac fairly complex. Both parameres rather large, with very wide,

rounded apex, left considerably larger than right.

? genitalia (Fig. 1231). Stylomere rather narrow, barely triagonal, apex obliquely rounded, with 2-3

subapical setae. Lateral plate elongate, with 2-3 apical setae.

Variation. There is considerable variation of size and even more variation of relative width of

pronotum and elytra. One extremely narrow and elongate specimen, collected together with two very

wide specimens from Katherine, demonstrates the wide range of shape in this species. In other respects

all specimens show a high degree of similarity.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. Specimens collected by me under bark of river gum near a tidal river,

others under bark of E. alba or E. bigalerita near a river bed. So far captured in February, April, August,

and November.

Distribution (Fig. 608). Northernmost Northern Territory from Darwin to Katherine.

Material examined (25). Only the type series.

Etymology. The name refers to the broad, red band across the elytra.

atrorufus-group

Diagnosis. Medium-sized, rather depressed, contrastingly black and red species. Labrum bisetose;

glossa c. 10-setose; lateral margin of pronotum only moderately explanate, basal angle shortly rounded

off; basal border line of elytra abbreviated, attaining % of base; scutellar pore absent; lateral margin of

elytra without elongate setae behind shoulders; series of lateral pores with 6 subhumeral pores only;

abdominal sterna without ambulatory setae; sternum VI without longer setae at apical margin; tibiae,

especially metatibia depressed; mesofemur and metafemur wide and depressed, profemur less wide;

d genitalia thus far unknown.

Larvae. Unknown.

Distribution. A single species from southwestern New South Wales.

Systematic position. This group is certainly closely related to the brevipennis-group, though slightly

more apomorphic in the less explanate lateral margins of the elytra and the presence of a contrasting

pattern.

Figs 124a-d, 1. Adelotopus atrorufus, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Adelotopus atrorufus, spec. nov.

Figs 34, 124, 311, 463, 609

Types. Holotype: ?, Australia: Coombah N. S. W. 16.X11.72 B. P. Moore (ANIC).

Diagnosis. Medium sized, rather depressed, markedly bicoloured species, that has the forebody black

with reddish lateral margins of the pronotum, and the elytra uniformly red.

Description

Measurements. Length: 6.3 mm. Ratios. Width/length of pronotum: 1.76; width base/apex of pro-

notum: 1.67; width pronotum/head: 1.82; length/ width of elytra: 1.42; length elytra/pronotum: 2.56.

Colour (Figs 34, 311). Black, lateral margins and apex of pronotum distinctly reddish translucent.

Elytra red. Lower surface of head and prothorax black, rest of lower surface reddish. Mouth parts,

antenna, and legs reddish.

Head (Figs 124a-d). Short and wide, moderately depressed. Anterior border but faintly convex,

lateral angle rounded, laterally not projecting, lateral borders behind eyes not narrowed. Clypeal

suture indistinct, semicircular, in middle widely interrupted. Labrum rather short and wide, apex

slightly concave. Antennal groove laterally sharply bordered, latero-posteriorly with slightly carinate

area. Mental tooth triangular, short, apex acute. Wings of mentum wide, laterally rounded, apex

rectangular. Glossa wide, tongue-like, apically widely rounded, ventrally with distinct keel, at border

with c. 10 elongate setae and additional pilosity on upper and lower surface and along border.

Terminal palpomere of maxillary palpus moderately wide, slightly securiform. Terminal palpomere of

labial palpus very wide, markedly securiform. Antenna elongate, 8th-9th antennomeres c. 1.3 x as

wide as long. Microreticulation fine, distinct, puncturation extremely fine, moderately dense, difficult

to detect, surface with a shallow sulcus medially of eyes, impilose, rather dull. Ventrolaterally of eyes

with a row of extremely short setae. Suborbital field at least laterally faintly punctate and setose. Both

palpi rather sparsely setose, gula impilose.

Pronotum (Fig. 311). Wide, moderately convex, base very wide, strongly narrowed to apex. Apical

angles rather produced, fairly acute, slightly oblique, surpassing posterior third of eyes. Apex

moderately deeply excised, rather convex in excision, very finely and irregularly bordered. Sides

strongly convex throughout, widest in basal third. Margins rather wide, fairly explanate, faintly

bordered. Basal angles shortly rounded off. Base slightly convex, rather coarsely bordered. Surface

near base without transverse impression. Microreticulation very fine, distinct, puncturation extremely

fine, moderately dense, rather difficult to detect, surface with several very fine, irregular strioles,

impilose, rather dull.

Elytra (Figs 311, 463). Short, rather wide, moderately convex, on disk slightly depressed, lateral

margins in basal two thirds parallel, though slightly sinuate behind base, apically evenly narrowed.

Apex moderately wide, transverse, truncature faintly convex, apical angles widely rounded off.

Shoulders rounded off, basal margin almost straight, without setae behind shoulders. Marginal

159)

channel rather wide, almost completely visible from above. Basal border incomplete, present in lateral

% of base, ending gradually. Lateral border asetose. Series of umbilical pores consisting of 6 rather

spaced pores behind shoulder. Setae rather short. Striae including sutural stria absent. Microreticula-

tion fine, slightly superficial, isodiametric, puncturation very fine, moderately sparse, rather difficult

to see. Surface impilose, rather dull.

Lower surface. Prosternal process rather short, fairly narrow, straight, convex, apex short and

narrow, evenly rounded off, shortly setose. Metepisternum rather short, c. 1.5 x as long as wide,

posteriorly not constricted nor hollowed. Abdominal sterna including sternum VI apparently without

setae at apical border. Lower surface rather sparsely punctate and pilose.

Legs. Rather elongate, Ist tarsomere of protarsus slightly longer than wide, tibial groove of

profemur deep, anterior plate distinctly overlapping the groove for basal half, posterior border of

groove sharp. Femur rather wide. Metatibia elongate, >6 x as long as wide, 1st tarsomere of metatarsus

>2 x as long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Figs 1241). Stylomere rather narrow and elongate, strongly widened towards base,

apex evenly rounded, with 3-4 subapical setae. Lateral plate elongate, with 2-3 short apical setae.

Variation. Unknown.

Vivipary. Not confirmed in examined material.

Habits. Unknown. Holotype collected in December.

Distribution (Fig. 609). Southwestern New South Wales.

Material examined (1). Only the holotype.

Etymology. The name refers to the contrasting black and red colouration of pronotum and elytra.

marginicollis-group

Diagnosis. Medium-sized, rather elongate, depressed, yellow or piceous-black species. Labrum

bisetose; glossa c. 16-setose; lateral margin of pronotum rather wide, fairly explanate, basal angle

shortly rounded or rectangular; basal border line of elytra abbreviated, attaining only middle of base;

scutellar pore absent; lateral margin of elytra without elongate setae; series of lateral pores with 6

subhumeral pores and 1 postmedion pore; abdominal sterna with 1 ambulatory seta each side;

sternum VI without a fringe of longer setae at apical margin; tibiae, especially metatibia rather

depressed; all femora wide and depressed; adeagus so far unknown.

Larvae. 1st instar larvae known of 2 species.

Distribution. 3 species in northwestern Queensland, central Northern Territory, and central Western

Australia. Apparently a decided inland group.

Systematic position. The relationships of this group are still rather obscure, because the Ö genitalia

are thus far unknown. However, it is probably the adelphotaxon of the brevipennis + atrorufus + katheri-

nei-groups. Due to many autapomorphies it has a rather isolated systematic position. It is plesiomorphic

in the absence of pattern and the large number of umbilical pores, and perhaps also in the rather

explanate lateral margin of the pronotum. It is apomorphic, however, in the bisetose labrum, high

number of glossal setae, abbreviated basal elytral border, low number of abdominal ambulatory setae,

and presence of more or less distinct transverse impressions on pronotum and elytra.

Adelotopus marginicollis, spec. nov.

Figs 35, 125, 312, 464, 610

Types. Holotype: ?, Aus. Qu. Mt. Isa March, 1986, J. Sedlacek, lg. (QMB T26079).

Diagnosis. Medium-sized, uniformly light reddish species, further distinguished from A. coriaceus,

spec. nov. and A. seminitidus, spec. nov. by the faint, not rasp-like puncturation of the surface and the

lack of a distinct microreticulation and coriaceous surface of the elytra.

Figs 125a-d, 1. Adelotopus marginicollis, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Description

Measurements. Length: 5.55 mm. Ratios. Width/length of pronotum: 1.38; width base/apex of

pronotum: 1.32; width pronotum/head: 1.45; length/width of elytra: 1.72; length elytra/pronotum:

2.41.

Colour. Upper and lower surface including mouth parts, antennae, and legs uniformly light

reddish.

Head (Figs 125a-d). Rather short, wide, rather depressed. Anterior border gently convex, lateral

angle rounded, laterally slightly projecting, lateral borders slightly narrowed behind eyes. Clypeal

suture semicircular, laterally distinct, in middle widely interrupted. Labrum large, moderately

overlapped by the clypeus, apex moderately deeply concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly convex area. Mental tooth triangular, rather short, apex

acute. Wings of mentum moderately wide, laterally rounded, apex obtuse. Glossa wide, tongue-like,

apically convex, ventrally with distinct keel, at border with c. 16 elongate setae. Terminal palpomere

of maxillary palpus wide, distinctly securiform. Terminal palpomere of labial palpus very wide,

markedly securiform. Antenna fairly narrow, Sth-9th antennomeres c. 1.8 x as wide as long. Microre-

ticulation extremely fine, superficial, puncturation also very fine, both difficult to detect. Surface with

very weak sulcus medially of eyes, very shortly pilose, fairly glossy. Ventrolaterally of eyes with a row

of rather long setae. Suborbital field punctate and shortly setose. Gula impilose.

Pronotum (Fig. 312). Rather wide, in middle moderately convex, much wider than long, base

distinctly wider than apex, though widest about in middle. Apical angles rather produced, at apex

obtusely rounded, fairly oblique, almost attaining posterior third of eyes. Apex well excised, convex

in excision, finely bordered. Sides evenly rounded. Margins wide, explanate, finely bordered. Basal

angles wide, obtusely rounded off. Base faintly convex, rather irregularly bordered. Surface near base

with distinct, though not very deep transverse impression. Microreticulation absent, puncturation

very fine, rather dense, surface with some extremely fine irregular wrinkles, very shortly pilose, glossy.

Elytra (Figs 312, 464). Rather elongate, slightly depressed on disk, rather parallel, though faintly

narrowed in basal third and widened in apical half. Disk in basal third with a fairly shallow transverse

impression. Apex wide, transverse, truncature slightly convex, apical angles widely rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

rather narrow throughout, slightly widened in basal third, partly concealed. Basal border incomplete,

reaching about halfways to suture. Lateral border asetose. Series of umbilical pores consisting of 6

rather spaced pores behind shoulder and an additioal seta behind middle. Setae fairly elongate. Striae

including sutural stria absent. Microreticulation absent, puncturation rather fine, fairly dense, slightly

rasp-like, surface shortly and rather densely pilose, glossy.

Lower surface. Prosternal process rather short, rather narrow, lower surface convex, apex NarroW,

compressed, passing over in a very wide angle from ventral surface, sparsely setose. Metepisternum

elongate, c. 1.8 x as long as wide, in posterior third not hollowed. Abdominal sterna with 1 elongate

seta each side. Lower surface moderately sparsely punctate and shortly setose.

Legs. Rather elongate, Ist tarsomere of protarsus about as wide as long, tibial groove of profemur

124

Figs 126a-d, 1. Adelotopus coriaceus, spec. nov. Details of head and % genitalia. For legends see fig. 100.

moderately deep, anterior plate overlapping the groove for apical third, posterior border of groove

sharp. Femur moderately wide. Metatibia narrow and elongate, >6.5 x as long as wide, Ist tarsomere

of metatarsus c. 2.3 x as long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Figs 1251). Stylomere moderately narrow, elongate, apex rather wide, obliquely

convex, with 2 elongate apical setae. Lateral plate elongate, with 1-2 elongate and 0-3 short apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined holotype.

Habits. Unknown. Holotype collected in March.

Distribution (Fig. 610). Northwestern Queensland. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the wide margins of the pronotum.

Adelotopus coriaceus, spec. nov.

Figs 36, 126, 313, 465, 610

Types. Holotype: ?, Yuendumu Central Aust., J. H. Sedlacek (QMB T26076). - Paratypes: 12, Yuendumu Centr.

Aust., J. H. Sedlacek Collector (CBM).

Diagnosis. Moderately large, dull black species, distinguished from the most closely related A. semi-

nitidus, spec. nov. by narrower base of pronotum, shorter elytra, coriaceous surface due to strong

microreticulation, rasp-like puncturation on the elytra, and narrow, elongate stylomere.

Description

Measurements. Length: 6.1-6.4 mm. Ratios. Width/length of pronotum: 1.49-1.52; width base/

apex of pronotum: 1.29-1.34; width pronotum/ head: 1.49-1.52; length/ width of elytra: 1.63-1.65; length

elytra/pronotum: 2.58.

Colour. Dull black, lateral margins of pronotum faintly reddish-piceous translucent. Lower

surface blackish-piceous. Mouth parts and antennae reddish-piceous, legs piceous, tibiae and tarsi

slightly lighter.

Head (Figs 126a-d). Rather short, wide, rather depressed. Anterior border gently convex, lateral

angle rounded, laterally slightly projecting, lateral borders slightly narrowed behind eyes. Clypeal

suture indistinct. Labrum large, moderately overlapped by the clypeus, apex deeply concave.

Antennal groove laterally sharply bordered, latero-posteriorly with slightly convex area. Mental tooth

triangular, rather short, apex acute. Wings of mentum moderately wide, laterally rounded, apex

obtuse. Glossa wide, tongue-like, apically convex, ventrally with distinct keel, at border with

c. 16 elongate setae. Terminal palpomere of maxillary palpus wide, distinctly securiform. Terminal

palpomere of labial palpus very wide, markedly securiform. Antenna fairly narrow and elongate,

rather parallel, 8th-9th antennomeres c. 1.8 x as wide as long. Microreticulation distinct, rather coarse,

puncturation rather coarse, moderately dense, somewhat irregular. Surface with very weak sulcus

medially of eyes and some irregular wrinkles, shortly pilose, dull, rather coriaceous. Ventrolaterally

of eyes with a row of rather long setae. Suborbital field punctate and shortly setose. Gula impilose.

Pronotum (Fig. 313). Rather wide, in middle moderately convex, much wider than long, base

distinctly wider than apex, though widest about in middle. Apical angles rather produced, at apex

obtusely rounded, fairly oblique, almost attaining posterior third of eyes. Apex well excised, slightly

convex in excision, bordered. Sides evenly rounded. Margins wide, explanate, finely bordered. Basal

angles wide, obtusely rounded off. Base faintly convex, rather coarsely bordered. Surface near base

with very deep and wide transverse impression. Microreticulation distinct, puncturation coarse, rather

dense, coriaceous, surface with several irregular wrinkles, pilose, dull, markedly coriaceous.

Elytra (Figs 313,465). Moderately elongate, rather wide, slightly depressed on disk, rather parallel,

though faintly narrowed in basal third and widened in apical half. Disk in basal third with a deep

transverse impression. Apex wide, transverse, truncature slightly convex, apical angles widely

rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders.

Marginal channel rather narrow throughout, slightly widened in basal third, partly concealed. Basal

border incomplete, reaching about halfways to suture. Lateral border asetose. Series of umbilical pores

consisting of 6 rather spaced pores behind shoulder and an additional seta behind middle. Setae fairly

elongate. Striae including sutural stria absent. Microreticulation distinct and rather coarse, punctura-

tion coarse, rather dense, markedly rasp-like, surface shortly setose, markedly dull.

Lower surface. Prosternal process rather short, rather narrow, lower surface convex, apex narrow,

compressed, passing over ina very wide angle from ventral surface, sparsely setose. Metepisternum

very elongate, c. 2.0 x as long as wide, in posterior third not hollowed. Abdominal sterna with 1

elongate seta each side. Lower surface moderately sparsely punctate and shortly setose.

Legs. Rather elongate, Ist tarsomere of protarsus about as wide as long, tibial groove of profemur

moderately deep, anterior plate overlapping the groove for apical third, posterior border of groove

sharp. Femur moderately wide. Metatibia very narrow and elongate, c. 7.5x as long as wide, 1st

tarsomere of metatarsus c. 2.5 x as long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Figs 1261). Stylomere narrow, elongate, apex narrow, acute, medially obliquely

convex, with 2-3 elongate apical setae. Lateral plate elongate, with 3-4 elongate apical setae.

Variation. There is some variation in the relative width of the base of the pronotum and in the

shape of the stylomere.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Unknown.

Distribution (Fig. 610). Central Northern Territory. Known only from type locality.

Material examined (2). Only the holotype and the paratype.

Etymology. The name refers to the markedly coriaceous surface.

Adelotopus seminitidus, spec. nov.

Figs 127, 314, 466, 610

Types. Holotype: ?, Australien, WA 27, Sullivan Creek, 97 km sse. Leinster, 8.11.1987, M. Baehr (WAM).

Diagnosis. Large, rather glossy black species, distinguished from the most closely related A. coriaceus,

spec. nov. by wider base of pronotum, longer elytra, glossy surface due to less distinct microreticula-

tion, irregular, not rasp-like puncturation on the elytra, and wide, rather short stylomere.

Description

Measurements. Length: 7.2 mm. Ratios. Width/length of pronotum: 1.49; width base/apex of pro-

notum: 1.41; width pronotum/head: 1.50; length/width of elytra: 1.76; length elytra/pronotum: 2.80.

126

Figs 127a-d, 1. Adelotopus seminitidus, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Colour. Dull black, lateral margins of pronotum faintly reddish-piceous translucent. Lower

surface blackish-piceous. Mouth parts and antennae reddish-piceous, legs piceous-black, tibiae and

tarsi slightly lighter.

Head (Figs 127a-d). Rather short, wide, rather depressed. Anterior border gently convex, lateral

angle rounded, laterally slightly projecting, lateral borders slightly narrowed behind eyes. Clypeal

suture indistinct. Labrum large, moderately overlapped by the clypeus, apex rather deeply concave.

Antennal groove laterally sharply bordered, latero-posteriorly with slightly convex area. Mental tooth

triangular, rather short, apex acute. Wings of mentum moderately wide, laterally rounded, apex

obtuse. Glossa wide, tongue-like, apically convex, ventrally with distinct keel, at border with c. 16

elongate setae. Terminal palpomere of maxillary palpus wide, distinctly securiform. Terminal pal-

pomere of labial palpus very wide, markedly securiform. Antenna moderately elongate, 8th-9th

antennomeres c. 2x as wide as long. Microreticulation almost absent, only faint traces visible,

puncturation coarse, very dense, Surface with very weak sulcus medially of eyes, shortly and densely

pilose, glossy. Ventrolaterally of eyes with a row of rather long setae. Suborbital field punctate and

shortly setose. Gula impilose.

Pronotum (Fig. 314). Rather wide, in middle moderately convex, much wider than long, base

distinctly wider than apex, widest near base. Apical angles rather produced, at apex obtusely rounded,

fairly oblique, almost attaining posterior third of eyes. Apex well excised, slightly convex in excision,

bordered. Sides anteriorly evenly rounded, in basal half almost straight. Margins wide, explanate,

finely bordered. Basal angles almost rectangular, obtusely rounded off. Base faintly convex, rather

coarsely bordered. Surface near base with very deep and wide transverse impression. Microreticula-

tion almost absent, only faint traces visible, puncturation coarse, very dense, surface densely pilose,

glossy.

Elytra (Figs 314, 466). Elongate, rather wide, slightly depressed on disk, rather parallel, barely

narrowed in basal third and widened in apical half. Disk in basal third with a deep transverse

impression. Apex wide, transverse, truncature slightly convex, apical angles widely rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

rather narrow throughout, slightly widened in basal third, partly concealed. Basal border incomplete,

reaching about halfways to suture. Lateral border asetose. Series of umbilical pores consisting of 6

rather spaced pores behind shoulder and an additional seta behind middle. Setae fairly elongate. Striae

including sutural stria very vaguely indicated by series of punctures set in shallow, irregular furrows.

Microreticulation absent, puncturation rather fine, very difficult to see, because the surface is markedly

coriaceous. Surface shortly setose, glossy, in spite of the highly coriaceous structure.

Lower surface. Prosternal process rather short, rather narrow, lower surface convex, apex narrow,

compressed, passing over in a very wide angle from ventral surface, sparsely setose. Metepisternum

very elongate, c. 2.2 x as long as wide, in posterior third not hollowed. Abdominal sterna with 1

elongate seta each side. Lower surface moderately sparsely punctate and shortly setose.

Legs. Elongate, 1st tarsomere of protarsus distinctly longer than wide, tibial groove of profemur

moderately deep, anterior plate overlapping the groove for apical third, posterior border of groove

127

sharp. Femur moderately wide. Metatibia very narrow and elongate, >8x as long as wide, Ist

tarsomere of metatarsus almost 3 x as long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Fig. 1271). Stylomere wide, tapering to apex, apex rather narrow, shortly rounded,

with 2 elongate apical setae. Lateral plate elongate, with 1-2 short apical setae.

Variation. Unknown.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unknown. Holotype collected by me under bark of river gum in November.

Distribution (Fig. 610). Interior of Western Australia. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the rather glossy surface as compared with the relative A. coriaceus, spec. nov.

exactor-group

Diagnosis. Medium-sized, rather depresed, piceous species with lighter margins. Labrum bisetose;

glossa c. 10-12-setose; lateral margin of pronotum slightly explanate, basal angle obtuse; basal border

line of elytra slightly abbreviated, reaching to median quarter of base, well visible; scutellar pore

absent; lateral margin of elytra moderatel, without elongate setae behind shoulders; series of lateral

pores with 6-7 subhumeral pores and 1 postmedian pore; abdominal sterna with 1 ambulatory seta on

either side; sternum VI with a fringe of slightly longer setae at apical margin; tibiae, especially

metatibia depressed; all femora including profemur wide and depressed; internal sac of aedeagus

complicate, without distinct oblique fold near apex.

Larvae. Unknown.

Distribution. A single rather remarkable species in central Papau New Guinea.

Systematic position. This group is apomorphic in the rather short and wide legs and the complex

internal sac of the aedeagus. It is rather plesiomorphic in the rather depressed body, the fairly wide

pronotal margins, the but slightly abbreviated basal border of elytra, the number of umbilical pores,

the presence of ambulatory setae on the abdomen, and the presence of a fringe of longer setae at the

margin of sternum VI. The systematic position of the group is thus far rather obscure.

Adelotopus exactor Darlington, 1968

Figs 37, 128, 315, 467, 610

Adelotopus exactor Darlington, 1968, p. 240, fig. 153.

Types. Holotype: d, Wareo Finsch Haven New Guinea Rev. L. Wagner, Holotype Adelotopus exactor Darl.

(SAMA). - Paratype: 17, same data, S. A. Museum specimen, exactor Darl., Paratype Adelotopus exactor

Darlington (MC2Z).

Type locality: “Wareo Finsch Haven”, Papua New Guinea.

Diagnosis. Medium-sized, rather wide, moderately convex, piceous species with wide, basally

rounded and slightly explanate, laterally distinctly reddish translucent pronotum. Further distin-

guished from relative species by rather distinct puncturation of head and pronotum, though much

finer puncturation of elytra, narrow, elongate aedeagus with wide, though acute apex, and remarkably

narrow right paramere.

Description

Measurements. Length: 6.7-7.0 mm. Ratios. Width/length of pronotum: c. 2.02-2.06; width base/

apex of pronotum: c. 1.87; width pronotum/head: c. 1.92; length/width of elytra: c. 1.38-1.40; length

elytra/pronotum: c. 2.90-3.10.

128

Figs 128a-l. Adelotopus exactor Darlington. Details of head and genitalia. For legends see fig. 100.

Colour (Figs 37, 315). Dark piceous, margins of pronotum and elytra distinctly reddish translu-

cent. Lower surface, mouth parts, antenna, and legs reddish.

Head (Figs 128a-d). Short and wide, moderately depressed. Anterior border gently convex,

though laterally of clypeus faintly concave, lateral angle angulate, laterally distinctly projecting, lateral

borders oblique. Clypeal suture invisible. Labrum rather large, apex markedly concave. Antennal

groove laterally sharply bordered, latero-posteriorly without carinate area. Mental tooth triangular,

short, apex acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa wide, tongue-like,

apically widely rounded, ventrally with indistinct keel, at border with c. 10-12 elongate setae and

additional pilosity on upper and lower surface and along border. Terminal palpomere of maxillary

palpus rather narrow, barely widened, not securiform. Terminal palpomere of labial palpus wide,

distinctly securiform. Antenna short, Sth-9th antennomeres >2 x as wide as long. Microreticulation

extremely fine, fairly superficial, puncturation moderately fine, distinct, fairly dense, though some-

what irregular, surface with a shallow sulcus medially of eyes, impilose, glossy. Ventrolaterally of eyes

with a row of short setae. Suborbital field impunctate and asetose. Both palpi rather densely setose,

gula apparently asetose.

Pronotum (Fig. 315). Wide, moderately convex, base wide, apex fairly narrow. Apical angles

moderately produced, fairly acute, slightly oblique, attaining posterior third of eyes. Apex moderately

deeply excised, slightly convex in excision, faintly bordered at least laterally. Sides curved throughout,

widest at basal angles. Margins rather wide, somewhat explanate, faintly bordered. Basal angles

shortly rounded off. Base almost straight, not bordered. Surface near base without transverse

impression. Microreticulation extremely fine, superficial, puncturation moderately fine, distinct,

rather dense, though somewhat irregular, surface with some fine, irregular strioles, impilose, fairly

glossy.

Elytra (Figs 315, 467). Moderately elongate, fairly convex, margins faintly convex throughout.

Apex rather wide, slightly oblique, truncature apparently faintly concave, apical angles rounded off.

Shoulders rounded off, basal margin slightly oblique, without setae behind shoulders. Marginal

channel moderately wide, completely visible from above. Basal border slightly abbreviated, absent in

median quarter, ending gradually. Scutellar pore absent. Lateral border asetose. Series of umbilical

pores consisting of 6-7 (in both specimens on left side 7, on right side 6) pores behind shoulder and

1 additional pore behind middle. Setae short. Striae including sutural stria absent. Microreticulation

very fine, rather superficial, though slightly more distinct than on fore body, almost isodiametric,

129

puncturation extremely fine, sparse, becoming slightly more distinct towards apex. Surface impilose,

rather glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex rather

short and wide, straight, shortly setose. Metepisternum short, c. 1.6 x as long as wide, in posterior third

obliquely bent and slightly hollowed. Abdominal sterna apparently with 1 elongate setae on either

side. Sternum VI with a row of slightly longer setae along apical border. Lower surface sparsely

punctate and pilose.

Legs. Medium-sized, 1st tarsomere of protarsus about as long as wide, tibial groove of profemur

very deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia rather short, c. 4.5 x as long as wide, Ist tarsomere of metatarsus c. 1.8 x aslong

as wide. d protarsus not widened.

d genitalia (Figs 128e-k). Genital ring rather narrow, triangular, slightly asymmetric, arms evenly

convex, with rather narrow, asymmetric base. Sternum VII rather wide, apically straight, with deep

excision, basally gently bisinuate, lateral parts short. Aedeagus elongate, moderately depressed,

narrow, parallel, barely asymmetric. Lower surface straight. Apex rather wide, rouded, though the

very tip acute. Orifice rather short, internal sac fairly complex, without a distinct, oblique fold at apex.

Both parameres, especially the right, rather elongate, left considerably larger than right, both with

rounded apex.

? genitalia (Fig. 1281). Stylomere short and rather wide, with wide, markedly rounded apex, with

2-4 subapical setae. Lateral plate rather short, with 4-6 long apical setae.

Variation. Due to scarce material little variation noted. Most measurements and ratios are

somewhat inaccurate, because both specimens are slightly damaged.

Vivipary. Not confirmed in the examined material.

Habits. Unknown.

Distribution (Fig. 610). Northeastern New Guinea. Known only from type locality.

Material examined (2). Only the holotype and the paratype.

politus-group

Diagnosis. Medium-sized to rather large, convex, black or patterned species. Labrum bisetose; glossa

c. 10-12-setose; lateral margin of pronotum narrow, not explanate, basal angle obtuse or shortly

rounded off; basal border line of elytra almost complete, ending immediately near suture, always well

visible; scutellar pore absent; lateral margin of elytra narrow, without elongate setae behind shoulders;

series of lateral pores usually with 6 subhumeral pores and 1 postmedian pore (only in A. substriatus,

spec. nov. the posthumeral pore absent); abdominal sterna with 1, rarely unilaterally 2 ambulatory

setae on either side; sternum VI without longer setae at apical margin; tibiae, especially metatibia

depressed; all femora including profemur wide and depressed; internal sac of aedeagus complicate,

with oblique fold near apex.

Larvae. Ist instar larvae known of 7 species and one additional subspecies.

Distribution. 16 species and one additional subspecies in eastern Australia from eastern South

Australia to northeastern Queensland, and in Tasmania.

Systematic position. This group is apomorphic in the convex body shape, the narrow margins of

pronotum and elytra, the rather short and wide legs, the presence of a pattern in some species, and the

complex internal sac of the aedeagus. It is, however, rather plesiomorphic in the almost complete basal

border of elytra, the number of umbilical pores, and the presence of ambulatory setae on the abdomen.

It may be the more primitive adelphotaxon of the following villosus-linearis-groups.

130

en ET er

Figs 129a-m. Adelotopus politus Castelnau. Details of head and genitalia. For legends see fig. 100. m. Stylomere

of specimen from Adelaide.

Adelotopus politus Castelnau, 1867

Figs 38, 129, 316, 468, 610

Adelotopus politus Castelnau, 1867, p. 31; 1868, p. 117; Lea 1910, p. 121; Notman 1925, p. 29; Csiki 1933, p. 1636;

Moore et al. 1987, p. 52.

Types. Lectotype (by present designation): d, Clarence River Coll. Castelnau, politus Cast. Clarence River,

Syntypus Adelotopus politus Castelnau, 1867 (MCSN) (see note below).

Type locality. “Clarence River”, northern New South Wales (but see note below).

Note. The description gives as type localities” Clarence River” and “Brisbane”. The latter locality,

however, refers to asyntype specimen belonging actually to A. montorum, spec. nov. The reasons, why

the name politus is not applied to this specimen, but to the syntype from Clarence River, are the

following: 1. In the description the surface is described as “of a polished black” which matches better

the specimen chosen here for the lectotype. 2. Already Gestro added two specimens to the type series

that are conspecific to the lectotype chosen by me, which is evidence that Gestro used the name politus

for the same species that I do.

Diagnosis. Medium-sized, rather elongate, very convex, black species with not explanate, at apex

obtusely rounded margin of pronotum. Further distinguished from relative species by extremely fine

microreticulation and puncturation of surface and short, barely asymmetric aedeagus with wide,

obtusely rounded apex.

Description

Measurements. Length: 5.35-7.5 mm. Ratios. Width/length of pronotum: 1.72-1.85; width base/

apex of pronotum: 1.58-1.69; width pronotum/head: 1.67-1.75; length/ width of elytra: 1.38-1.48; length

elytra/pronotum: 2.52-2.69.

Colour. Black, sometimes margins of pronotum and of elytra vaguely translucent. Lower surface

of head and prothorax piceous, of abdomen reddish. Mouth parts, antenna, and legs piceous or

reddish-piceous, cemmonly femora lighter, even reddish.

Head (Figs 129a-d). Short and rather wide, moderately depressed. Anterior border gently convex,

lateral angle obtusely rounded, laterally slightly projecting, lateral borders oblique. Clypeal suture

semicircular, in middle narrowly interrupted. Labrum rather wide, apex feebly concave. Antennal

groove laterally sharply bordered, latero-posteriorly without carinate area. Mental tooth triangular,

short, apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-

like, apically widely rounded, ventrally with indistinct keel, at border with c. 12 elongate setae and

additional pilosity on upper and lower surface and along border. Terminal palpomere of maxillary

palpus rather narrow, barely widened, not securiform. Terminal palpomere of labial palpus wide,

distinctly securiform. Antenna short, 8th-9th antennomeres >2 x as wide as long. Microreticulation

very fine, though distinct, slightly superficial, puncturation extremely fine, sometimes almost invisible

even under high magnification, surface with a shallow sulcus medially of eyes, impilose, rather glossy.

Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Both palpi

rather sparsely setose, gula almost asetose.

Pronotum (Figs 38, 316). Moderately wide, highly convex, base wide, apex fairly narrow. Apical

angles moderately produced, fairly acute, though obtuse at apex, slightly oblique, surpassing posterior

border of eyes. Apex moderately deeply excised, rather convex in excision, not bordered. Sides curved

throughout, widest at or near basal angles. Margins very narrow, not bordered. Basal angles more or

less obtusely rounded off. Base almost straight, faintly bordered. Surface near base without transverse

impression. Microreticulation very fine, rather superficial, puncturation extremely fine, sometimes

almost invisible even under high magnification, surface impilose, rather glossy.

Elytra (Figs 38, 316, 468). Moderately elongate, convex, margins faintly convex throughout. Apex

rather wide, slightly oblique, truncature slightly convex, apical angles rounded off. Shoulders rounded

off, basal margin slightly oblique, without setae behind shoulders. Marginal channel narrow, com-

pletely visible from above. Basal border almost complete, absent only very close to suture, ending

gradually. Scutellar pores absent. Lateral border asetose. Series of umbilical pores consisting of 6

(sometimes unilaterally 7) pores behind shoulder and 1 additional pore behind middle. Setae short.

Striae including sutural stria absent. Microreticulation very fine, highly superficial, isodiametric to

slightly transverse, puncturation extremely fine, rather sparse. Both, microreticulation and punctura-

tion visible only under high magnification. Surface impilose, glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex rather

short and wide, straight, shortly setose. Metepisternum short, c. 1.5 x as long as wide, in posterior third

obliquely bent and deeply hollowed. Abdominal sterna with 1 elongate setae on either side. Sternum

VI without elongate setae at apical border. Lower surface moderately punctate and pilose.

Legs. Medium-sized, 1st tarsomere of protarsus about as long as wide, tibial groove of profemur

very deep, anterior plate widely Sver/apping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia rather short, c. 4 x as long as wide, 1st tarsomere of metatarsus <1.8 x as long

as wide. d protarsus not widened.

d genitalia (Figs 129e-k). Genital ring rather wide, triangular, slightly asymmetric, arms evenly

convex, with rather narrow, asymmetric base. Sternum VII rather wide, apically straight, with deep

excision, basally gently concave, lateral parts rather short. Aedeagus short, moderately depressed, in

middle slightly widened, slightly asymmetric. Lower surface convex. Apex rather wide, obtusely

rounded. Orifice rather short, internal sac fairly complex, with a large, oblique fold at apex. Right

DAtamIesE.S rather elongate, left considerably larger than right, both with obtusely rounded apex.

? genitalia (Figs 1291,m). Stylomere short and wide, with wide, rather transverse apex, with 2-3

Sbapieal setae. Lateral plate moderately elongate, with 2-4 apical setae.

Variation. This species is rather variable with respect to size and shape of pronotum and elytra.

Whereas relative width of pronotum depends to some degree on sex, because the larger females

usually possess a relatively wider and more convex pronotum than the smaller and narrower males,

relative length of elytra is perhaps not connected with sex. There is also some variation in shape of the

basal angles of pronotum which may be more or less evenly rounded, and in microreticulation and

puncturation of the surface.

The two specimens from South Australia differ somewhat in their ? genitalia (Fig. 129m) and may

belong to another taxon, but this is doubtful, as long as no males are available.

Vivipary. Confirmed by discovery of larvae in the 2 oviducts.

Habits. Little known, although the number of specimens is rather numerous. Specimens collected by

me under bark of river gums, single other species captured in “Malaise trap” and in “flight intercept

trap”. So far collected in all months, but more commonly during summer. This is a common and

widespread species, though apparently most common in southeastern Queensland, apparently rare in

northern Queensland.

Distribution (Fig. 610). Southeastern and eastern Australia from eastern South Australia through

eastern Victoria, eastern New South Wales and eastern Queensland up to lower Cape York Peninsula.

A single old record from Tasmania is doubtful. Perhaps the specimens from South Australia belong

to another taxon (see above under “variation”).

Material examined (112). SA: 17, Port Lincoln Blackburn, Sharp Coll. 1906, Adelotopus sp. ? Id. by T. G. Sloane

(BMNH); 1%, Adelaide Blackburn, hydrobioides Westw., Adelotopus hydrobioides W.N.S. W.: SA 17109 (SAMA). —

Vic: 15, 1?, Melbourne Stevens, Ex Musaeo Chaudoir, det. hydrobioides Westw. (MNHN); 22%, Ringwood C.

Oke, Adelotopus hydrobioides Westw. (NMV); 1 (sex ?), Delatite R. 8.11.59 B. P. Moore (CMC). - Tas: 1 (sex?),

scolytides Mw. V. Diem. Prob. this ex. = gyrinoides Hope. It is not Newmans type but det as his sp. in Coll. WMS

t.in B.M. jan. 23 1922, G. J. Arrow (OUM). - NSW: 1?, N. Sydney, 6.VII.1912 Musgrave, Adelotopus sp. L 30436

(AMS); 12, Sydney district J. J. W. XII.1900, hydrobioides= apicalis (ANIC); 15,12, Sydney, R. Mus. Hist. Nat. Belg.

I. G. 12 595 (IRSNB); 13, 17, Megalong Vy, Blue Mts., 22.1.32, 1000 ft. Australia Harvard Exp. Darlington,

Adelotopus hydrobioides Westw. (MCZ); 2? 2, Mt. Victoria D’Albertis 73, politus Cast., Adelotopus politus Cast. det.

R. Gestro (MCSN); 12, Mt. Victoria 1.1931 C. Oke, Adelotopus politus Cast. (NMV); 12, Mt. Wilson 1.1931; C. Oke

(NMV); 222, Hunter R. (MMS); 1%, haemorrhoidalis Goulburn R. (OUM); 14, 4-8 km SW Lake Cathie 11.X1.1982

J. Doyen Coll. (ANIC); 15, Bonville 26.1.78 B. P. Moore (CMC); 1%, Yuragir NP Station Creek 20.X1.1982, J. &

E. Doyen (ANIC); 230, Greta 1951, J. Sedlacek (CSB); 272, Eccleston T. G. S. III.21, A. politus Cast. comp. spec

in H. Coll. 26.X1.21 (ANIC); 1%, Ac. Ck. T. G. S. 25.X1l.10, A. politus Cast. (ANIC); 1?, West Head 25.X1.1978

D. A. Doolan, D. A. Doolan Coll. (AMS); 145, Bostobrick 15.11.1981 D. A. Doolan, D. A. Doolan Coll. (AMS);

22% (2), Gordon 16.X11.1962 D. A. Doolan, D. A. Doolan Coll. (AMS); 15, Clarence River Coll. Castelnau, politus

Cast. Clarence River, lectotype politus! (MCSN); 1%, Clar. R., politus Cast. 50 Howitt Colln. (NMV); 15, A. politus

Tweed R., Adelotopus politus Cast. (SAMA); 1% (2), Rope’s Ck. (MMS); 1%, Coll. Castelnau, N. S. W., hydrobioides

Westw. (MCSN). - Old: 15, Warwick X.1949, E. (2), det. politus (NMV); 82%, Stanthorpe 21.VIll.24 E. Sutton,

E. Sutton Coll. 1964 (QMB); 13, Stanthorpe, 4.V.23 (QMB); 12, Stanthorpe, 111.82 J. Sedlacek (CSB); 15, Qld 69,

15 km ssw. Rathdowney, 26.X1.1990, M. Baehr (CBM); 17, Mt. Maroon, 400-700 m, 13.X11.1981, M. Baehr (CBM);

15, Qld 61, 6 km e. Spicer’s Gap, 22.X1.1990, M. Baehr (CBM); 1?, Tambourine Mt. 10.V.-28, E. Sutton Coll.

(QMB); 15, 12, Brisbane Illidge (UOIC); 17, Birö 1900, Brisbane (HNMB); 1%, Brisbane H. Hacker 8.VIl.11,

Adelotopus politus Cast. Det. Sloane (QMB); 12, Brisbane 1.07 Adelotopus politus Cast., Coll. Hacker, A. politus Lap.

(DEIB); 1?, Brisbane: H. Hacker 23.X.21 (QMB); 15, Brisbane Mt. Gravatt 2.1.52 C. Oke, Adelotopus politus Cast.

compared with paratype (NMV); 1%, Bickdale Brisbane H. Hacker 16.11.25 (QMB); 15, Sandford, 18.11.1973,

J. Collingwood (UQIC); 18, Sandgate, Coll. F. Muir 9.X1.1919 (BMH); 2? 9, Caloundra H. Hacker 28.IX.13 (QMB);

12, 42156, Moreton Bay, politus Cast. (MNHB); 13, Moreton Bay, Brisbane VIL.-VIN.1915, J. C. Bridwell Coll., det.

Scolytides (USNM); 12, E. Sutton TE 18.X11.31 Rivertree, E. Sutton Coll. 1964 (QMB); 17, Deception Bay,

17.1X.1961, R. G. Winiks (UQIC); 1, Dunwich, Stradbroke Is. 27.1V.1963, G. Monteith (UQIC); 15, Moreton Bay,

Stradbroke 1. 20.1X.1915, J. G. Bridwell Coll., A. scolytides det. T. L. Erwin (USNM); 15, Upper Cedar Ck. Via

Samford, Qld. 6.X11.62 G. Monteith (UQIC); 15, Glen Aplin, Old. 6.1.1964 P. Kerridge (UOIC); 17, Kenilworth

State Forest, 5.X11.1966, G. Monteith (UQIC); 15, Hampton S. Q. 1.65 T. M. J. G. Brooks Bequest, 1976 (ANIC);

12, Lawes 10.1V.52 Prarad (UOIC); 1%, Fraser Island 1.-6.V.1967, C. R. Hembrow (UOIC); 19, 1%, Frazer Is.

15.V.81 (CSB); 13, Bundaberg, Coll. F. Muir 11.1919 (BMH); 1%, Bundaberg, Perkins (BMNH); 17, Yeppoon,

11.1.1962, L. A. Powell (UOIC); 222, Qld 20, Cania Gorge, 25 km nw. Monto, 9.-11.X1.1990, M. Baehr (CBM);

12, Kroombit Tops 10.XI. 700 m, J. Sedlacek (CSB); 17, Qld 22, Cariboe Ck., 26 km sse. Biloela, 11.X1.1990,

M. Baehr (CBM); 13, Carnarvon Rge, N. Geary, 5.1.59, det. politus (AMS); 17, Carnarvon R. A. 1IX.54 JGB, M. 188,

J. G. Brooks Bequest, 1976 (ANIV); 17, Maryborough (SAMA); 12, Rockhampton 26.X11.1967 J. & M. Sedlacek

collr. (BMH); 1?, Rockhampton (OUM); 13, Mackay, Adelotopus (ANIC); 15, 5161 Mackay, apicalis Macl.,

Adelotopus apicalis Macl. (SAMA); 15, 131 Mackay (SAMA),; 22%, Mackay, C. French’s Coll. (NMV); 15, 15.395

144.31E Split Rock QLD 24 Nov-13 Dec 1992 Malaise Trap P. Zborowski & W. Dressler (DPIM); 14, 15.395

144.42E Split Rock QLD 29 Jun-24 Aug 1992 Flight Intercept Trap P. Zborowski & ]J. Cardale (DPIM); 15, 1%,

C. York, Soc. Ent. Belg. Coll. PUTZEYS, R.1.Sc.N.B. 1.G. Coll. gen. (IRSNB); 14, Caborlah (?) Q. T.G.S. 24.12.10

(ANIC); 12, Ex Museo H. W. Bates 1892 (MNHN); 12, Thorey 1867, North Holl. bor.(NHMW). — Aus: 12, 58.124,

det. gyrinoides (BMNH); 14, Ex Museo L. Fairmaire 1896 (MNHN). - 2:12, Mt. Tonah 11.32. HJC (ANIC); 18,328,

Palm B. 31.X11.33, M. 169, J. G. Brooks Bequest, 1976 (ANIC); 12, E. W. Ferguson Coll. Adelotopus haemorrhoidalis

Erich. (ANIC); 12, Thorey 1867 (NHMW); 1? (NMV).

Adelotopus variolosus Lea, 1910

Figs 130, 317, 469, 611

Adelotopus variolosus Lea, 1910, p. 121; Notman 1925, p. 6, 29; Csiki 1933, p. 1636; Moore et al. 1987, p. 53.

Types. Holotype: %, variolosus Lea Type Sydney, 2494 Adelotopus variolosus Lea N. S. Wales, Type (SAMA).

Type locality: “Sydney”, New South Wales.

Diagnosis. Medium-sized, rather wide, convex, black species with wide, basally rounded and

laterally distinctly beaded pronotum. Further distinguished from relative species by the extremely fine

microreticulation and puncturation of surface.

Description

Measurements. Length: 7.0-7.3 mm. Ratios. Width/length of pronotum: 1.89-1.95; width base/

apex of pronotum: 1.77-1.80; width pronotum/head: 1.79-1.87; length/width of elytra: c. 1.38; length

elytra/pronotum: 2.62-2.74.

Colour. Dark piceous to black, margins of pronotum and elytra vaguely translucent. Lower

surface of head and pronotum blackish-piceous, of abdomen reddish-piceous to reddish. becoming

lighter towards apex. Antenna and legs reddish-piceous, femora lighter.

Head (Figs 130a,d). Short and rather wide, moderately depressed. Anterior border gently convex,

lateral angle obtusely rounded, laterally slightly projecting, lateral borders oblique. Clypeal suture

semicircular, complete. on either side near base with a distinct pit. Labrum rather large, apex feebly

concave. Antennal groove laterally sharply bordered, latero-posteriorly without carinate area. Mental

tooth triangular, short, apex acute. Wings of mentum wide, laterally rounded, apex obtusely rounded.

Glossa wide, tongue-like, apically widely rounded, ventrally with indistinct keel, at border with

c. 10-12 elongate setae and additional pilosity on upper and lower surface and along border. Both,

maxillary and labial palpi in both examined specimens broken. Antenna short, 8th-9th antennomeres

c.2 x as wide as long. Microreticulation very fine, though distinct, puncturation extremely fine, almost

invisible even under high magnification. Surface with a shallow sulcus medially of eyes and with some

faint, irregular strioles, impilose, rather glossy. Ventrolaterally of eyes with a row of short setae.

Suborbital field impunctate and asetose. Gula apparently asetose.

Pronotum (Fig. 317). Wide, rather convex, base wide, apex rather narrow. Apical angles moderate-

ly produced, fairly acute, though obtuse at apex, slightly oblique, surpassing posterior border of eyes.

Apex moderately deeply excised, rather convex in excision, not bordered. Sides curved throughout,

widest near basal angles. Margins narrow, not explanate, distinctly bordered. Basal angles rounded

off. Base faintly bisinuate or concave, not bordered. Surface near base without transverse impression.

Microreticulation extremely fine, rather superficial, puncturation even finer than on head, rather

sparse, difficult to see even under high magnification, surface with some faint irregular strioles,

impilose, glossy, in holotype with some large drop-like impressions.

Elytra (Figs 317, 469). Moderately elongate, convex, margins faintly convex throughout. Apex

rather wide, slightly oblique, truncature slightly convex, apical angles rounded off. Shoulders rather

distinct, obtusely rounded, basal margin slightly oblique, without setae behind shoulders. Marginal

channel narrow, completely visible from above. Basal border almost complete, absent only very close

to suture, ending gradually. Scutellar pores absent. Lateral border asetose. Series of umbilical pores

consisting of 6-7 pores behind shoulder and 1 additional pore behind middle. Setae short. Striae

including sutural stria absent in basal half, inner striae near apex vaguely indicated by extremely fine

rows of strioles and punctures. Microreticulation extremely fine, highly superficial, isodiametric to

slightly transverse, puncturation minute, rather sparse, difficult to see, becoming more distinct

towards apex. Surface impilose, glossy, in holotype with several large and small drop-like impres-

sions, especially on left elytron.

Figs 130a, d, 1. Adelotopus varıolosus Lea. Details of head and ? genitalia. For legends see fig. 100.

Lower surface. Prosternal process rather short, fairly wide, straight, depressed, apex rather short

and wide, straight, shortly setose. Metepisternum rather short, c. 1.7 x as long as wide, in posterior

third obliquely bent and somewhat hollowed. Abdominal sterna with 1 elongate setae on either side.

Sternum VI without elongate setae at apical border. Lower surface moderately punctate and pilose.

Legs. Medium-sized, 1st tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur very deep, anterior plate widely overlapping the groove for apical %, posterior border of groove

sharp. Femur wide. Metatibia moderately short, c. 4.5 x as long as wide, 1st tarsomere of metatarsus

c. 1.6 x as long as wide. d protarsus unknown. In holotype most legs broken, only femora and

mesotibia of left side left.

d genitalia. Unknown.

? genitalia (Fig. 130). Stylomere moderately wide to rather wide, with wide, transverse to feebly

convex apex, with 3 subapical setae. Lateral plate rather elongate, with 3 long apical setae.

Variation. Little variation noted, besides of the conspicuous drop-like impressions in the holotype.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. One specimen captured in September. Certainly this is a rare or even already

extinct species, because the type locality is “Sydney” and it has been never recollected in recent times.

Distribution (Fig. 611). Central eastern New South Wales. Known only from type locality.

Material examined (2). 1?, variolosus Lea Type Sydney, 2494 Adelotopus variolosus Lea N. S. Wales, Type,

holotype! (SAMA); 1?, Australien, A. L Schrader leg, ded 25.1X.1896, Adelotopus hydrobioides Westw ? Det. Sloane

’07 (ANIC).

Note. The drop-like impressions on pronotum and elytra of the holotype emphasized in the descrip-

tion do not constitute a determinative character of this species, though an abnormal structure that is

also rarely seen in other species.

Adelotopus aterrimus, spec. nov.

Figs 131, 318, 470, 611

Types. Holotype: 4, 3902, Adelotopus 3902 N. S. Wales (SAMA). - Paratype: 17, Australien, NSW 82, 50 km

e. Armidale, 29.11.1990, M. Baehr (CBM).

Diagnosis. Rather large, elongate, moderately convex, deep glossy black species with evenly rounded

basal angles, but not explanate lateral margins of pronotum. Further distinguished by glossy surface,

extremely fine puncturation, and rather symmetric aedeagus with widely rounded apex.

Description

Measurements. Length: 7.55-8.55 mm. Ratios. Width/length of pronotum: 1.67-1.72; width base/

apex of pronotum: 1.56-1.58; width pronotum/ head: 1.67-1.70; length/ width of elytra: 1.60-1.66; length

elytra/pronotum: 2.81-2.85.

[09]

[67 |

Figs 131a-l. Adelotopus aterrimus, spec. nov. Details of head and genitalia. For legends see fig. 100.

Colour. Deep glossy black, only apex of elytra faintly reddish translucent. Lower surface of head

and thorax black to piceous, abodmen piceous to reddish-piceous, becoming lighter towards apex.

Mouth parts and antennae reddish, legs piceous.

Head (Figs 131a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle rounded, laterally barely projecting, lateral borders faintly oblique. Clypeal suture semicircular,

almost complete. Labrum large, apex barely concave. Antennal groove laterally sharply bordered,

latero-posteriorly without carinate area. Mental tooth triangular, short, apex acute. Wings of mentum

wide, laterally rounded, apex obtuse. Glossa wide, tongue-like, apically slightly triangular, ventrally

with indistinct keel, at border with c. 10-12 elongate setae and additional pilosity on upper and lower

surface and along border. Terminal palpomere of maxillary palpus rather narrow, barely widened, not

securiform. Terminal palpomere of labial palpus wide, distinctly securiform. Antenna rather short,

Sth-9th antennomeres slightly <2 x as wide as long. Microreticulation fine, though distinct, punctur-

ation almost invisible within microreticulation, surface with a shallow sulcus medially of eyes,

impilose, rather glossy. Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate

and asetose. Both palpi moderately setose, gula apparently asetose.

Pronotum (Fig. 318). Rather wide, fairly convex, base wide, apex fairly narrow. Apical angles

moderately produced, fairly acute, slightly oblique, attaining posterior third of eyes. Apex moderately

deeply excised, slightly convex in excision, unbordered. Sides curved throughout, widest at basal

angles. Margins rather narrow, not explanate, faintly bordered. Basal angles rather widely rounded off.

Base almost straight to slightly convax, finely bordered. Surface near base without transverse

impression. Microreticulation very fine, rather superficial, puncturation extremely fine, almost invis-

ible even under high magnification, rather sparse, surface impilose, glossy.

Elytra (Figs 318, 470). Moderately elongate, fairly convex, margins in basal % almost parallel. Apex

rather wide, slightly oblique, truncature rather convex, apical angles rounded off. Shoulders obtusely

rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel narrow,

slightly concealed in basal part. Basal border almost complete, absent only very close to suture, ending

gradually. Scutellar pore absent. Lateral border asetose. Series of umbilical pores consisting of 6 pores

behind shoulder and 1 additional pore (or unilaterally 2 pores) behind middle. Setae moderately short.

Striae including sutural stria absent or indicated as very indistinct, widely spaced rows of fine

136

punctures. Microreticulation absent, puncturation extremely fine, dificult to detect even under high

magnification, rather sparse, becoming slightly more distinct towards apex. Surface impilose, remark-

ably glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex rather

short and wide, convex, sparsely setose. Metepisternum moderately elongate, c. 1.8 x as long as wide,

in posterior third neither bent nor hollowed. Abdominal sterna with 1 elongate setae on either side.

Sternum VI without longer setae along apical border. Lower surface very sparsely punctate and pilose.

Legs. Medium-sized, Ist tarsomere of protarsus claerly wider than long, tibial groove of profemur

very deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia rather short, c. 4.5 x as long as wide, Ist tarsomere of metatarsus c. 1.5 x aslong

as wide. d protarsus not widened.

d genitalia (Figs 131e-k). Genital ring medium-sized, triangular, rather asymmetric, arms evenly

convex, with rather narrow, asymmetric base. Sternum VII rather wide, apically gently convex, with

deep excision, basally probably gently concave, lateral parts moderately elongate. Aedeagus medium-

sized, moderately depressed, in middle slightly widened, feebly asymmetric. Lower surface almost

straight. Apex wide, rounded off. Orifice rather short, internal sac fairly complex, with a large, oblique

fold at apex. Both parameres, rather wide, left considerably larger than right, both with widely

rounded apex.

? genitalia (Fig. 1311). Stylomere short and rather wide, triagonal, with wide, rounded apex, with

2-3 subapical setae. Lateral plate elongate, with 3 long apical setae.

Variation. Apart from some differences of relative shape and of degree of puncturation of elytra

little variation noted.

Vivipary. Not confirmed in the examined material.

Habits. Little known. The paratype collected by me in November under bark of ash-like eucalypt in

rather dense eucalypt forest.

Distribution (Fig. 611). New South Wales. Only the paratype is exactly labelled and is from the New

England Tableland in northern New South Wales.

Material examined (2). Only the holotype and the paratype.

Etymology. The name refers to the deep black colouration.

Adelotopus doyeni, spec. nov.

Figs 132, 319, 471, 611

Types. Holotype: d, Australia, NSW 14 km N Deepwater, 25.11.1989, H. & A. Howden, Under logs (NMO). -

Paratypes: 17, Mebbin St. For. NSW, 18 km W of Uki, 23-24 Nov.1982, J. Doyen (ANIC); 17, Qld. 13-15 km N

of Quinalow, 29 Nov.1982, J. Doyen coll. (ANIC); 17, Australien, Old 53, 20 km s. Gin Gin, 21.11.1990, M. Baehr

(CBM); 18, Australia. Qld. Blackdown Tableland nr. Ranger Station 900 m, 15.X11.1988, H. & A. Howden (NMO).

Diagnosis. Medium-sized, moderately elongate, moderately convex, black species with obtuse angles,

not explanate lateral margins of pronotum. Further distinguished by fine microreticulation, fine

puncturation, and short, asymmetric aedeagus with narrowly rounded apex.

Description

Measurements. Length: 5.4-6.9 mm. Ratios. Width/length of pronotum: 1.73-1.85; width base/

apex of pronotum: 1.56-1.67; width pronotum/ head: 1.63-1.75; length/ width of elytra: 1.47-1.53; length

elytra/pronotum: 2.60-2.72.

Colour. Black, margins of pronotum and elytra more or less distinctly reddish translucent. Lower

surface of head and thorax blackish to piceous, of abdomen reddish-piceous to reddish, becoming

lighter towards apex. Mouth parts and antenna piceous to reddish-piceous, legs piceous, femora

usually slightly lighter.

Head (Figs 132a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle rounded, laterally barely projecting, lateral borders faintly oblique. Clypeal suture semicircular,

in middle widely interrupted. Labrum rather large, apex gently concave. Antennal groove laterally

Figs 132a-l. Adelotopus doyeni, spec. nov. Details of head and genitalia. For legends see fig. 100.

sharply bordered, latero-posteriorly without or with gently carinate area. Mental tooth triangular,

short, apex acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa wide, tongue-like,

apically convex, ventrally with indistinct keel, at border with c. 10-12 elongate setae and additional

pilosity on upper and lower surface and along border. Terminal palpomere of maxillary palpus rather

narrow, barely widened, not securiform. Terminal palpomere of labial palpus wide, distinctly

securiform. Antenna short, Sth-9th antennomeres slightly <2 x as wide as long. Microreticulation very

fine, more or lass distinct, puncturation either almost invisible even at high magnification or very fine

though fairly distinct. Surface with a shallow sulcus medially of eyes, impilose, rather glossy.

Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Both palpi

moderately setose, gula apparently asetose.

Pronotum (Fig. 319). Rather wide, fairly convex, base wide, apex fairly narrow. Apical angles

moderately produced, fairly acute, slightly oblique, attaining posterior third of eyes. Apex moderately

deeply excised, slightly convex in excision, laterally faintly bordered. Sides curved throughout, widest

at or near basal angles. Margins rather narrow, not explanate, faintly bordered. Basal angles more or

less obtusely rounded. Base almost straight to slightly convex, finely bordered. Surface near base

without transverse impression. Microreticulation very fine, more or less superficial, puncturation in

males extremely fine, almost invisible even under high magnification and rather sparse, in females

fairly distinct and rather dense, surface impilose, glossy.

Elytra (Figs 319, 471). Moderately elongate, fairly convex, margins in basal half almost parallel or

but faintly convex. Apex rather wide, slightly oblique, truncature slightly convex, apical angles rounded

off. Shoulders obtusely rounded, basal margin slightly oblique, without setae behind shoulders.

Marsginal channel rather narrow, barely concealed in basal part. Basal border almost complete, absent

only very close to suture, ending gradually. Scutellar pore absent. Lateral border asetose. Series of

umbilical pores consisting of 6 pores behind shoulder and I additional pore behind middle. Setae

moderately short. Striae including sutural stria absent or indicated as more or less distinct rows of

moderately fine punctures. Microreticulation fine, somewhat superficial, slightly transverse, punctu-

ration in males extremely fine, difficult to detect even under high magnification, sparse, in females

more distinct, fairly dense, becoming more distinct towards apex. Surface impilose, glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

138

rather short, transversely cut, sparsely setose. Metepisternum moderately elongate, c. 1.8 x as long as

wide, in posterior third distinctly obliquely bent and hollowed. Abdominal sterna with 1 elongate

setae on either side. Sternum VI without longer setae along apical border. Lower surface rather densely

punctate and pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur very deep, anterior plate widely overlapping the groove for apical %, posterior border of groove

sharp. Femur wide. Metatibia rather short, c. 4.5 x as long as wide, Ist tarsomere of metatarsus c. 1.7 x

as long as wide. d protarsus not widened.

d genitalia (Figs 132e-k). Genital ring medium-sized, triangular, rather asymmetric, arms evenly

convex, with rather narrow, asymmetric base. Sternum VII rather wide, apically gently convex, with

deep excision, basally slightly excised but gently convex, lateral parts moderately elongate. Aedeagus

rather short, moderately depressed, in middle rather widened, asymmetric. Lower surface almost

straight. Apex moderately wide, rounded off. Orifice rather short, internal sac fairly complex, with a

large, oblique fold at apex. Right paramere rather narrow, apex obliquely rounded, left large,

considerably larger than right, with widely rounded apex.

? genitalia (Fig. 1321). Stylomere rather narrow, fairly parallel, with wide, obliquely rounded apex,

with 2-3 subapical setae. Lateral plate elongate, with 3-4 long apical setae.

Variation. There is considerable variation of size, relative width of pronotum, and degree of

microreticulation and puncturation of pronotum and elytra. Perhaps the latter differences are partly

sexual, because both males have more superficial microreticulation and very sparse and indistinct

puncturation, whereas in the three females both, microreticulation and puncturation are markedly

more distinct.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Little known. One specimen collected “under logs”, another captured by me under bark of

river gum. One specimen collected at the height of 900 m on the Blackdown Tableland. Records are

from November, December, and February.

Distribution (Fig. 611). Northern New South Wales, southeastern Queensland north to about Rock-

hampton.

Material examined (5). Only the type series.

Etymology. Named in honour of the collector of a part of the type series.

Adelotopus substriatus, spec. nov.

Figs;133, 320, 472,612

Types. Holotype: ?, HT, 3896, A. hydrobioides Westw. Forest-Reefs, Lea’s, Adelotopus hydrobioides We. N. S.

Wales 268, Adelotopus haemorrhoidalis Er. Id. by T. G. Sloane (SAMA). - Paratypes: 2? ?, same data, on same card

(SAMA); 19, Australien, NSW 81,5 km e. Armidale, 29.11.1990, M. Baehr (CBM); 12, Australien, NSW 76, 22 km

n. Glen Innes, 28.11.1990, M. Baehr (CBM).

Diagnosis. Rather elongate and narrow, black species with narrow, at basal angles shortly rounded

pronotum with very narrow margin. Further distinguished by the distinct traces of striae on the whole

elytra.

Description

Measurements. Length: 6.0-6.8 mm. Ratios. Width/length of pronotum: 1.53-1.60; width base/

apex of pronotum: 1.48-1.54; width pronotum/head: 1.52-1.58; length/ width of elytra: 1.68-1.70; length

elytra/pronotum: 2.70-2.78.

Colour. Black. Lower surface of head and thorax dark piceous, of abdomen reddish-piceous to

reddish, beoming lighter towards apex. Mouth parts and antenna reddish to reddish-piceous, legs

piceous, femora slightly lighter.

Head (Figs 133a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle rounded, laterally barely projecting, lateral borders faintly oblique. Clypeal suture semicircular,

not interrupted. Labrum large, apex barely concave. Antennal groove laterally sharply bordered,

latero-posteriorly without carinate area. Mental tooth triangular, short, apex acute. Wings of mentum

Figs 133a-d, I. Adelotopus substriatus, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

wide, laterally rounded, apex obtuse. Glossa wide, tongue-like, apically convex, ventrally with

indistinct keel, at border with c. 10-12 elongate setae and additional pilosity on upper and lower

surface and along border. Terminal palpomere of maxillary palpus rather narrow, barely widened, not

securiform. Terminal palpomere of labial palpus wide, distinctly securiform. Antenna short, 8th-9th

antennomeres slightly <2 x as wide as long. Microreticulation fine, though distinct, puncturation

present, though almost invisible even at high magnification due to distinct microreticulation. Surface

with a shallow sulcus medially of eyes, impilose, moderately glossy. Ventrolaterally of eyes with a row

of short setae. Suborbital field impunctate and asetose. Both palpi moderately setose, gula apparently

asetose.

Pronotum (Fig. 320). Rather narrow, convex, base moderately wide, apex distinctly narrower.

Apical angles moderately produced, moderately airly acute, slightly oblique, attaining posterior third

of eyes. Apex moderately excised, slightly convex in excision, unbordered. Sides gently curved

throughout, widest at basal third. Margins narrow, not explanate, distinctly bordered. Basal angles

shortly, though evenly rounded off. Base almost straight, finely bordered. Surface near base without

transverse impression. Microreticulation very fine, slightly superficial, puncturation extremely fine,

difficult to see even under high magnification, though laterally more distinct, moderately dense,

surface with some extremely fine irregular strioles, impilose, glossy.

Elytra (Figs 320, 472). Rather elongate, convex, margins faintly convex throughout. Apex rather

wide, slightly oblique, truncature slightly convex, apical angles rounded off. Shoulders obtusely

rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel narrow,

partly concealed on whole length. Basal border almost complete, absent only very close to suture,

ending gradually. Lateral border asetose. Series of umbilical pores consisting of 6 pores behind

shoulder only. Setae moderately short. Striae including sutural stria faintly indicated as rows of

extremely fine longitudinal strioles and punctures, sometimes even extremely faintly impressed.

Microreticulation very fine, highly superficial, slightly transverse, puncturation fine, though distinct,

dense, surface impilose, glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, straight, sparsely setose. Metepisternum moderately elongate, c. 1.8 x as long as wide, in

posterior third neither bent nor hollowed. Abdominal sterna with 1 elongate setae on either side.

Sternum VI without longer setae along apical border. Lower surface rather densely punctate and

pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur very deep, anterior plate widely overlapping the groove for apical %, posterior border of groove

sharp. Femur wide. Metatibia moderately short, >4.5 x as long as wide, 1st tarsomere of metatarsus c.

1.3 x as long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Fig. 1331). Stylomere fairly narrow, feebly triangular, with wide, rounded apex, with

2-4 subapical setae. Lateral plate elongate, with 2-3 long apical setae.

Variation. Little variation noted.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

140

Figs 134a-d, 1. Adelotopus sedlaceki, spec. nov. Details of head and % genitalia. For legends see fig. 100.

Habits. Largely unknown. Two specimens collected by me in November under bark of gum-type

eucalypts.

Distribution (Fig. 612). Northeastern New South Wales.

Material examined (5). Only the type series.

Etymology. The name refers to the traces of striation on the elytra.

Adelotopus sedlaceki, spec. nov.

Figs 134, 321, 473, 611

Types. Holotype: ?, Bamaga, N.Q., Dec. 1983, J. H. Sedlacek (QMB T26081).

Diagnosis. Medium-sized, rather wide, moderately convex, dull black species with wide pronotum

with rather wide, slightly explanate margins and wide, rectangular basal angles. Further distinguished

by comparatively depressed build, fine, though very distinct microreticulation, rather dull, silky

surface, and elytral striae at apex marked by fairly coarse, slightly rasp-like punctures.

Description

Measurements. Length: 6.7 mm. Ratios. Width/length of pronotum: 1.86; width base/apex of

pronotum: 1.76; width pronotum/head: 1.81; length/width of elytra: 1.38; length elytra/pronotum:

2.506:

Colour. Rather dull black, margins of pronotum and elytra rather distinctly reddish translucent.

Lower surface of head and prothorax piceous, of rest of thorax and abdomen reddish-piceous to

reddish, becoming lighter towards apex. Mouth parts, antenna and legs reddish-piceous, femora

lighter.

Head (Figs 134a-d). Short and wide, moderately depressed. Anterior border rather convex, lateral

angle obtusely rounded, laterally slightly projecting, lateral borders faintly oblique. Clypeal suture

semicircular, in middle slightly interrupted. Labrum large, apex moderately concave. Antennal groove

laterally sharply bordered, latero-posteriorly without carinate area. Mental tooth triangular, short,

apex acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa wide, tongue-like, apically

convex, ventrally with indistinct keel, at border with c. 10-12 elongate setae and additional pilosity on

upper and lower surface and along border. Terminal palpomere of maxillary palpus slightly widened,

feebly securiform. Terminal palpomere of labial palpus very wide, markedly securiform. Antenna

short, $Sth-9th antennomeres c. 2x as wide as long. Microreticulation fine, though very distinct,

puncturation very fine, though distinct, rather dense. Surface with a shallow sulcus medially of eyes,

impilose, fairly dull. Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and

asetose. Gula apparently asetose.

Pronotum (Fig. 321). Wide, rather convex, base wide, apex narrow. Apical angles rather produced,

fairly acute, somewhat oblique, surpassing posterior third of eyes. Apex rather deeply excised, slightly

141

convex in excision, laterally finely bordered. Sides evenly curved throughout, widest at basal angles.

Margins rather wide, slightly explanate, not bordered. Basal angles almost rectangular, barely obtuse,

faintly produced backwards. Base slightly concave, unbordered. Surface near base without transverse

impression. Microreticulation fine, distinct, puncturation extremely fine, difficult to see even under

high magnification, moderately dense, surface with some faint, irregular strioles, impilose, dull, rather

silky.

Elytra (Figs 321, 473). Moderately elongate, moderately convex, margins faintly convex through-

out. Apex rather narrow, slightly oblique, truncature faintly convex, apical angles rounded off.

Shoulders obtusely rounded, basal margin slightly oblique, without setae behind shoulders. Marginal

channel moderately narrow, on whole length visible from above. Basal border almost complete, absent

only very close to suture, ending gradually. Lateral border asetose. Series of umbilical pores consisting

of 6 pores behind shoulder and 1 additional pore behind middle. Setae moderately elongate. Median

striae in basal half faintly indicated as rows of extremely fine longitudinal strioles, in apical third as

irregular rows of rather large, somewhat rasp-like punctures. Microreticulation fine, though distinct,

about isodiametric, puncturation basally extremely fine, difficult to see, sparse, becoming more

distinct and denser towards apex and margins, surface impilose, dull, fairly silky.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, straight, rather setose. Metepisternum moderately elongate, c. 1.8 x as long as wide, in

posterior third obliquely bent and deeply hollowed. Abdominal sterna with 1 elongate setae on either

side. Sternum VI without longer setae along apical border. Lower surface rather densely punctate and

pilose.

Legs. Medium-sized, 1st tarsomere of protarsus about as wide as long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical half, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, slightly <4.5 x as long as wide, 1st tarsomere of metatarsus

c. 1.8 x as long as wide. d protarsus not widened.

d genitalia. Unknown.

? genitalia (Fig. 1341). Stylomere fairly wide, gently triangular, with wide, rather transverse apex,

with 3 subapical setae. Lateral plate elongate, with 2-3 long apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Holotype collected in December.

Distribution (Fig. 611). Tip of Cape York Peninsula, northernmost Queensland. Known only from

type locality.

Material examined (1). Only the holotype.

Etymology. Named in honour of the collector.

Adelotopus caniae, spec. nov.

Figs 135, 322, 474, 612

Types. Holotype: d, Australien, Qld 21, 2 km se. Cania Gorge, 25 km n. Monto, 11.11.1990, M. Baehr (ANIC).

Diagnosis. Rather small, narrow, moderately convex, glossy black species. Distinguished by rather

narrow pronotum, with moderate margins and obtusely rounded basal angles, parallel, on disk

somewhat depressed elytra with dense and rather coarse puncturation, and short and rather high,

symmetric aedeagus with rather triangular parameres.

Description

Measurements. Length: 5.45 mm. Ratios. Width/length of pronotum: 1.37; width base/apex of

pronotum: 1.16; width pronotum/head: 1.23; length/width of elytra: 1.62; length elytra/pronotum:

2192.

Colour. Dark piceous to blackish, margins of pronotum and elytra and apex of elytra slightly

reddish translucent. Lower urface piceous to reddish-piceous, becoming lighter to apex of abdomen.

Mouth parts light reddish, legs reddish-piceous.

S i

4 k

Figs 135a-c, e-k. Adelotopus caniae, spec. nov. Details of head and Ö genitalia. For legends see fig. 100.

Head (Figs 135a-c). Short and very wide, depressed. Anterior border barely convex, lateral angle

obtusely rounded, laterally slightly projecting, lateral borders faintly oblique. Clypeal suture semicir-

cular, though indistinct and in middle widely interrupted. Labrum large, apex moderately concave.

Antennal groove laterally sharply bordered, latero-posteriorly without carinate area. Mental tooth

triangular, short, apex acute. Wings of mentum wide, laterally rounded, apex rounded. Glossa wide,

tongue-like, apically convex, ventrally with indistinct keel, at border with c. 10-12 elongate setae and

additional pilosity on upper and lower surface and along border. Terminal palpomere of maxillary

palpus slightly widened, feebly securiform. Terminal palpomere of labial palpus very wide, markedly

securiform. Antenna broken. Microreticulation very fine, rather superficial, puncturation very fine,

difficult to see, rather sparse. Surface with a shallow sulcus medially of eyes, impilose, fairly dull.

Ventrolaterally of eyes with a row of short setae. Suborbital field punctate, pilosity extremely fine and

short. Gula apparently asetose.

Pronotum (Fig. 322). Moderately wide, rather convex, base not much wider than apex. Apical

angles rather produced, fairly acute, somewhat oblique, attaining posterior third of eyes. Apex

moderately excised, slightly convex in excision, at least laterally finely bordered. Sides feebly but

evenly curved, widest in basal third. Margins moderately wide, but not explanate, faintly bordered.

Basal angles >100°, obtuse. Base straight, finely and irregularly bordered. Surface near base with faint

transverse impression. Microreticulation fine, very superficial, difficult to detect, puncturation fine,

though rather distinct, moderately dense, surface impilose, glossy.

Elytra (Figs 322, 474). Elongate, rather narrow, moderately convex, on disk slightly depressed,

margins largely parallel. Apex rather wide, slightly oblique, truncature faintly convex, apical angles

rounded off. Shoulders obtusely rounded, basal margin slightly oblique, without setae behind

shoulders. Marginal channel narrow, partly concealed on whole length. Basal border almost complete,

absent only very close to suture, ending gradually. Lateral border asetose. Series of umbilical pores

consisting of 6 pores behind shoulder and 1 additional pore behind middle. Setae rather short. Median

striae in apical half only faintly indicated as irregular rows of rows of rather fine punctures.

Microreticulation absent, puncturation rather fine, dense, becoming more distinct and even denser

towards apex, surface impilose, markedly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, straight, rather setose. Metepisternum moderately elongate, c. 1.3 x as long as wide, in

posterior third obliquely bent and deeply hollowed. Abdominal sterna with 1 elongate setae on either

side. Sternum VI without longer setae along apical border. Lower surface rather densely punctate and

pilose.

Legs. Moderately elongate, protarsus broken, tibial groove of profemur deep, anterior plate

widely overlapping the groove for apical half, posterior border of groove sharp. Femur wide. Metatibia

rather elongate and narrow, c. 5 x as long as wide, Ist tarsomere of metatarsus c. 1.8 x as long as wide.

d protarsus unknown.

d genitalia (Figs 135e-k). Genital ring medium-sized, triangular, rather symmetric, arms evenly

convex, with rather narrow, barely asymmetric base. Sternum VII rather wide, apically gently convex,

with moderately deep excision, basally slightly excised, lateral parts rather short. Aedeagus short,

rather high, in middle rather widened, symmetric. Lower surface almost straight. Apex moderately

wide, shortly rounded off. Orifice rather short, internal sac fairly complex, with a large, oblique fold

at apex. Right paramere rather narrow, triangular, apex obtusely rounded, left large, considerably

larger than right, with slightly triangular, rounded apex.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Holotype collected by me in November under bark of large river gum near creek.

Distribution (Fig. 612). Central eastern Queensland. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the type locality, the Cania Gorge.

Adelotopus rufocaudatus, spec. nov.

Figs 136, 323, 475, 612

Types. Holotype: d, 12 km e. Taroom, z. Qld. Austral. 17.12.1981, M. Baehr (ANIC).

Diagnosis. Medium sized, rather wide, convex, dull piceous-black species with vaguely separated

reddish apex. Further distinguished by rather wide pronotum with fairly wide, though not explanate

margins and shortly rounded basal angles; distinctly microreticulate, dull surface, and small, short,

symmetric aedeagus with markedly rounded apex.

Description

Measurements. Length: 5.85 mm. Ratios. Width/length of pronotum: 1.84; width base/apex of

pronotum: 1.66; width pronotum/head: 1.75; length/width of elytra: 1.39; length elytra/pronotum:

Colour (Fig. 323). Piceous, margins of pronotum and elytra vaguely reddish translucent, apex of

elytra reddish, though not well delimited. Lower surface reddish-piceous, becoming slightlxy lighter

towards apex of abdomen. Mouth parts, antenna, and legs reddish-piceous, though tarsi slightly

darker.

Head (Fig. 136a-d). Short and wide, moderately depressed. Anterior border fairly convex, lateral

angle rounded, laterally barely projecting, lateral borders faintly oblique. Clypeal suture semicircular,

in middle barely interrupted. Labrum large, apex moderately concave. Antennal groove laterally

sharply bordered, latero-posteriorly without carinate area. Mental tooth triangular, short, apex acute.

Wings of mentum wide, laterally rounded, apex obtuse. Glossa wide, tongue-like, apically convex,

ventrally with indistinct keel, at border with c. 10-12 elongate setae and additional pilosity on upper

and lower surface and along border. Terminal palpomere of maxillary palpus slightly widened, feebly

securiform. Terminal palpomere of labial palpus very wide, markedly securiform. Antenna short,

Sth-9th antennomeres c. 2 x as wide as long. Microreticulation fine, though very distinct, puncturation

very fine, difficult to detect within dense microreticulation, fairly dense. Surface with a shallow sulcus

medially of eyes, impilose, fairly dull. Ventrolaterally of eyes with a row of short setae. Suborbital field

impunctate and asetose. Gula apparently asetose.

Pronotum (Fig. 323). Rather wide, convex, base fairly wide, apex narrow. Apical angles rather

produced, at apex obtuse, somewhat oblique, attaining posterior third of eyes. Apex moderately

deeply excised, slightly convex in excision, unbordered. Sides evenly curved throughout, widest near

basal angles. Margins moderate, not explanate, faintly bordered. Basal angles shortly rounded off. Base

almost straight, faintly and irregularly bordered. Surface near base without transverse impression.

Microreticulation fine, though distinct, puncturation extremely fine, difficult to see even under high

144

Figs 136a-k. Adelotopus rufocaudatus, spec. nov. Details of head and J genitalia. For legends see fig. 100.

magnification, rather sparse, surface with some very faint, irregular strioles, impilose, rather dull,

somewhat silky.

Elytra (Figs 323, 475). Medium-sized, moderately convex, margins faintly convex throughout.

Apex rather narrow, slightly oblique, truncature faintly convex, apical angles rounded off. Shoulders

obtusely rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

moderately narrow, on whole length visible from above. Basal border almost complete, absent only

very close to suture, ending gradually. Lateral border asetose. Series of umbilical pores consisting of

6 pores behind shoulder and 1 additional pore behind middle. Setae moderately elongate. Striae in

basal half absent, towards apex extremely faintly indicated as rows of very fine, rather irregular

longitudinal strioles. Microreticulation fine, though distinct, slightly transverse, puncturation very

fine, basally difficult to see, moderately sparse, becoming more distinct and denser towards apex,

surface impilose, dull, fairly silky.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, straight, rather setose. Metepisternum short, slightly <1.5 x as long as wide, in posterior

third markedly obliquely bent and deeply hollowed. Abdominal sterna with 1 elongate setae on either

side. Sternum VI without longer setae along apical border. Lower surface moderately punctate and

pilose.

Legs. Medium-sized, 1st tarsomere of protarsus about as wide as long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, slightly <4.5 x as long as wide, Ist tarsomere of metatarsus

c. 1.6 x as long as wide. d protarsus not widened.

d genitalia (Figs 136e-k). Genital ring medium-sized, regularlxy triangular, rather symmetric,

arms barely convex, with rather narrow, barely asymmetric base. Sternum VII rather wide, apically

almost straight, with deep, slightly irregular excision, basally slightly excised, lateral parts short.

Aedeagus short, rather high, in middle rather widened, symmetric. Lower surface convex. Apex very

wide, widely rounded off. Orifice rather short, internal sac fairly complex, with a large, oblique fold

at apex. Both parameres markedly triangular, apex obtusely rounded, left large, considerably larger

than right.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Holotype collected by me in December at light in open eucalypt forest mixed with cabbage

tree palms at the border of a billabong.

Distribution (Fig. 612). Central Queensland. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the reddish apex of the elytra.

Adelotopus haemorrhoidalis Erichson, 1842

Figs 137, 324, 476, 613

Adelotopus haemorrhoidalis Erichson, 1842, p. 126; Westwood 1853, p. 407, pl. 14, fig. 3; Macleay 1863, p. 113;

Blackburn 1901a, p. 18, 19; Sloane 1920, p. 177; Notman 1925, p. 8, 10, 29; Csiki 1933, p. 1635; Darlington 1968,

p- 241; Moore et al. 1987, p. 51.

? Adelotopus inguinatus Newman, 1842, p. 366; Westwood 1853, p. 407; Lacordaire 1854, p. 154; Notman 1925,

p- 29; Csiki 1933, p. 1635; Moore et al. 1987, p. 51 (doubtful synonymy).

Types. Of haemorrhoidalis. Lectotype (by present designation): ? (2), 1082, Schayer, haemorrhoidalis Er* Tasmania

(MNHB).

Of ingquinatus: Not found in BMNH, probably lost.

Type localities. Of haemorrhoidalis: “Tasmania”. - Of inguinatus: From description: “Port Phillip S. A.”,

Victoria.

Note. Since the type(s) of A. inguinatus are probably lost and because several species compete for being

inguinatus, this synonymy is doubtful. Inquinatus is therefore ranked under “doubtful species”.

Diagnosis. Rather large, elogate, convex, black species with rather wide reddish apex of elytra.

Distinguished from related A. similis, spec. nov. species by wider pronotum with relatively wider base,

wider elytra, medium-sized aedeagus with rounded apex, shorter parameres, and rather wide,

triangular stylomere; from A. minor, spec. nov. by larger size, slightly wider pronotum and elytra,

slightly shorter aedeagus, shorter parameres, and distinctly triangular stylomere.

Description

Measurements. Length: 6.4-8.1 mm. Ratios. Width/length of pronotum: 1.62-1.68; width base/

apex of pronotum: 1.55-1.62; width pronotum/head: 1.60-1.70; length/ width of elytra: 1.60-1.68; length

elytra/pronotum: 2.69-2.82.

Colour (Fig. 324). Black, sometimes margins of pronotum and elytra feebly dark reddish translu-

cent. Elytra with well defined red apex, the anterior border of the spot distinctly concave. Lower

surface of head and thorax black to dark piceous, of abdomen reddish. Mouth parts, antennae, and legs

dark reddish, tibiae and tarsi slightly darker.

Head (Figs 137a-d ). Short and wide, moderately depressed. Anterior border fairly convex, lateral

angle rounded, laterally barely projecting, lateral borders faintly oblique. Clypeal suture semicircular,

in middle slightly interrupted. Labrum large, apex barely concave. Antennal groove laterally sharply

bordered, latero-posteriorly without carinate area. Mental tooth triangular, short, apex acute. Wings

of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically convex,

ventrally with indistinct keel, at border with c. 10-12 elongate setae and additional pilosity on upper

and lower surface and along border. Terminal palpomere of maxillary palpus elongate, barely

widened, not securiform. Terminal palpomere of labial palpus rather wide, securiform. Antenna short,

Sth-9th antennomeres almost 2 x as wide as long. Microreticulation fine, though distinct, puncturation

extremely fine, difficult to detect even under high magnification, rather sparse. Surface with a shallow

sulcus medially of eyes, impilose, moderately glossy. Ventrolaterally of eyes with a row of short setae.

Suborbital field impunctate and asetose. Gula apparently asetose.

Pronotum (Fig. 324). Moderately wide, convex, base fairly wide, apex narrow. Apical angles

moderately produced, at apex obtuse, somewhat oblique, attaining posterior third of eyes. Apex

moderately deeply excised, slightly convex in excision, unbordered. Sides evenly curved throughout,

widest in basal third or fourth. Margins moderate, not explanate, faintly bordered. Basal angles evenly

rounded off. Base almost straight, distinctly bordered. Surface near base without transverse impres-

sion. Microreticulation very fine, distinct, though somewhat superficial, puncturation extremely fine,

though slightly more distinct than on head, rather sparse, surface impilose, rather glossy.

146

Figs 137a-l. Adelotopus haemorrhoidalis Erichson. Details of head and genitalia. For legends see fig. 100.

Elytra (Figs 324, 476). Elongate, convex, margins rather parallel in basal half to %, then gently

narrowed. Apex rather wide, transverse, truncature faintly convex, apical angles rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

moderately narrow, almost completely on visible from above. Basal border almost complete, ending

very closely to suture, becoming very weak there. Lateral border asetose. Series of umbilical pores

consisting of 6 pores behind shoulder and 1 additional pore behind middle, sometimes with a small

intercalary pore. Setae moderately elongate. Striae either completely absent, or faintly indicated as

rows of extremely fine, irregular, longitudinal strioles, or as rows of very shallow impressions.

Microreticulation fine, very superficial, sometimes difficult to detect even under high magnification,

puncturation very fine, rather sparse, becoming more distinct and denser towards apex, surface

impilose, glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin transverse, rather setose. Metepisternum elongate, c. 2 x as long as wide or even

slightly longer, neither bent nor hollowed. Abdominal sterna with 1 elongate setae on either side.

Sternum VI without longer setae along apical border. Lower surface moderately punctate and pilose.

Legs. Medium-sized, 1st tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, c. 4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.6 x as

long as wide. d protarsus not widened.

d genitalia (Figs 137e-k). Genital ring medium-sized, gently triangular, rather symmetric, arms

evenly convex, with rather narrow, fairly excised base. Sternum VII moderately wide, apically convex,

with deep excision, basally regularly excised, lateral parts short. Aedeagus medium-sized, moderately

depressed, in middle slightly widened, slightly asymmetric, apex slighty turned laterally . Lower

surface straight. Apex very wide, widely rounded off. Orifice rather short, internal sac fairly complex,

with a conspicuous, oblique fold at apex. Right paramere rather wide, almost parallel, apex widely

rounded. Left paramere considerably larger than right, triangular, apex obtusely rounded.

? genitalia (Fig. 1371). Stylomere wide, distinctly triangular, with narrow, obtuse apex, with 1-4

subapical setae. Lateral plate elongate, with 2-3 long apical setae.

Variation. Some variation noted in size, relative shape, degree of striation of elytra and punctur-

ation of surface.

Vivipary. Confirmed by discovery of larvae in the $ oviducts.

Habits. Largely unknown. One specimen collected on “E. stellulatum trunks by pyrethrum spray”, a

single specimen collected by me under bark of gum-type eucalpt at about 1200m. So far collected from

November to April and in August, most specimens, hovwever, during the period from December until

February. Occurs sympatrically and even syntopically with A. similis (on Picadilly Circus actually on

the same tree!).

Distribution (Fig. 613). Eastern Victoria, Tasmania, Australian Capital Territory, eastern New South

Wales, southeastern Queensland. The latter record is perhaps close to the New South Wales border.

Material examined (38). Vic: 17, Tennyson Creek, 5 km NW of Buldah 37 14°S 149 07’E, 10.-16.1.1982 ANZSES

Expedition, Adelotopus haemorrhoidalis Er. (NMV); 12, Erskine Falls near Lorne, 25.11.1971 Neboiss (NMV);

13, Melbourne Coll. Castelnau, det. haemorrhoidalis (MCSN); 1%, gyrinoides Hope Melb., Melbourne Coll.

Castelnau, haemorrhoidalis (MCSN); 1%; Warburton F. E. Wilson 1.11.26, F. E. Wilson Collection, det. seriepunctatus

(NMV); 14, Macedon C. Oke, det. haemorrhoidalis (NMV); 12, Buff. Adelotopus haemorrhoidalis Er. Id. by

T. G. Sloane (SAMA). - Tas: 17, Launceston, 91-88, 3939, det. haemorrhoidalis (BMNH); 1%, Launceston C. Oke

1.1941, Adelotopus haemorrhoidalis Er. (NMV); 1%, Arthur Plains S.W.Tas, 6.11.1965 Neboiss (NMV); 12, Hobart,

J. J. Walker, det. haemorrhoidalis (BMNH); 1%, G. H. Hardy, Triabunna 25.X11.1920, Adelotopus hydrobioides W.

Id.by A. M. Lea (QMB); 1? (2), 1082, Schayer, haemorrhoidalis Er, (lectotype!) (MNHB); 1%, 2611 A. Simson,

Adelotopus haemorrhoidalis Erich. Id. by T. G. Sloane (SAMA). - ACT: 15, Kangaroo Ck., Corin Dam Rd, 20.11.1986

K.R. Pullen, Kim Pullen Coll. (ANIC); 18, 6 km E. Corin Dam, Smokes Flat, 4.1V.1980, J. F. Lawrence (CBM);

13, ACT 124, Brindabella Rge, 1200 m, Picadilly Circus, 10.X11.1987, M. Baehr (CBM); 15, 1%, Picadilly Circus,

3650", 9.1.1983, J. Doyen, det. haemorrhoidalis Er. (CUIC); 15, Gininito Bimberi, Brindabella Ra., 28.-29.1.1980,

K. R. Pullen, Kim Pullen Coll. (ANIC); 18, Booroomba Rocks, 6.X1.1988, J&R. Bell (UVB).- NSW: 15, The Creel.

Mt. Kosc. 8000 ft, 15.X1L.31, Harvard Exp. Darlington, Adelotopus haemorrhoidalis Er. (MCZ); 15, 17, Kosciusko,

J. Sedlacek (CAS, CSB); 14, 1?, Tumut R., 1500', X11.53, J. Sedlacek (CBM, CSB); 13, 17, Tumut R., 1450 m 1956,

J. Sedlacek (CBM, CSB); 1% (2), Mul. XT1.96 inguinatus Newm. (ANIC); 1%, Rules Pt. H. J. Carter, 11.24, Adelotopus

inguinitus Erichs. (sic !), Id. by T. G. Sloane (ANIC); 13, 23.VIII.86, Isl. Bend Pondage, V. R. Bejsak Igt. (CBS);

12, Forest Reefs, Lea, Lea’s (SAMA); 14, 17, Goniawa (?) (MMS); 12, Bago Forest, Barlow, 10.111.57, T. G. Campbell

(ANIC). - Qld: 14, Nat. Park, 2.X1.53, F. A. Perkins (UQIC). — Aus: 1?, Adelotopus haemorrhoidalis Erichs. Arch.

1892 126 (BMNH); 1%, W. Edwards, gyrinoides Hope (MCZ).

Doubtful specimens (2). 1 (defect, sex ?), Talbingo T.G.S. ?.X11.22 (ANIC); 1 (defect, sex ?), 951, Victoria,

Adelotopus haemorrhoidalis Erich Tasman. Vic, 29 Howitt Colln., det. gyrinoides (NMV).

Adelotopus minor, spec. nov.

Figs 138, 325, 477, 614

Types. Holotype: d, Healsville, V.C. Oke, det. haemorrhoidalis (NMV). - Paratypes: 17, Mt. Buffalo 8-22-35 Aus.

(BMH); 12, Tasmania, A. Simson, Adelotopus haemorrhoidalis Er. Id. by T. G. Sloane (SAMA); 15, 35.305 150.18E,

Kioloa SF 15 km NE Batemans Bay, NSW Apr.87, M. G. Robinson, ex sticky trap (ANIC); 17,4 N Batemans Bay

21 Oct 1952, E. F. Riek NSW (CBM); 12, Mooney Mooney Ck. Near Gosford N.S.W. 16 Nov 1978, D. K. McAlpine

and B. J. Day (AMS); 12, N. Holl. Janson Acq. 1884 (MNHN); 14, Australie Melly, Ex Museo Chaudoir, det.

haemorrhoidalis (MNHN).

Diagnosis. Medium-sized, elongate, convex, black species with rather wide reddish apex of elytra.

Distinguished from related A. haemorrhoidalis Erichson by lesser size, slightly narrower pronotum with

relatively narrower base, slightly narrower elytra, slightly longer aedeagus, longer parameres, and not

markedly triangular stylomere; from A. similis, spec. nov. by lesser size, slightly wider pronotum and

elytra, medium-sized aedeagus with convex apex, and wider, apically mor evenly rounded stylomere;

from A. nitens, spec. nov. by barely concave labrum, distinct microreticulation on surface, even on

elytra, less distinct pucturation of surface, only faintly indicated elytral striae not marked by distinct

punctures, and less triangular stylomere.

Description

Measurements. Length: 6.0-6.3 mm. Ratios. Width/length of pronotum: 1.56-1.64; width base/

apex of pronotum: 1.51-1.56; width pronotum/ head: 1.55-1.65; length/ width of elytra: 1.62-1.67; length

elytra/pronotum: 2.63-2.70.

Colour (Fig. 325). Black, rarely margins of pronotum and elytra feebly dark reddish translucent.

Elytra with well defined red apex, the anterior border of the spot distinctly concave. Lower surface of

148

Figs 138a-l. Adelotopus minor, spec. nov. Details of head and genitalia. For legends see fig. 100.

head and thorax black to dark piceous, of abdomen light reddish. Mouth parts, antennae, and legs dark

reddish, tibiae and tarsi slightly darker.

Head (Figs 138a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle rounded, laterally barely projecting, lateral borders faintly oblique. Clypeal suture semicircular,

in middle slightly interrupted. Labrum large, apex barely concave. Antennal groove laterally sharply

bordered, latero-posteriorly without carinate area. Mental tooth triangular, short, apex acute. Wings

of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically convex,

ventrally with indistinct keel, at border with c. 10-12 elongate setae and additional pilosity on upper

and lower surface and along border. Terminal palpomere of maxillary palpus elongate, barely

widened, not securiform. Terminal palpomere of labial palpus rather wide, securiform. Antenna rather

short, Sth-9th antennomeres slightly <2 x as wide as long. Microreticulation fine, though distinct,

puncturation very fine, rather difficult to see even under high magnification, rather sparse. Surface

with a shallow sulcus medially of eyes, impilose, moderately glossy. Ventrolaterally of eyes with a row

of short setae. Suborbital field impunctate and asetose. Gula apparently asetose.

Pronotum (Fig. 325). Moderately wide, convex, base fairly wide, apex narrow. Apical angles

moderately produced, at apex obtuse, somewhat oblique, almost attaining posterior third of eyes.

Apex moderately deeply excised, slightily convex in excision, unbordered. Sides evenly curved

throughout, widest in basal third or fourth. Margins narrow to moderate, not explanate, faintly

bordered. Basal angles evenly rounded off. Base almost straight, bordered. Surface near base without

transverse impression. Microreticulation very fine, distinct, only slightly superficial, puncturation

extremely fine, though slightly more distinct than on head, rather sparse, surface impilose, fairly

glossy.

Elytra (Figs 325, 477). Elongate, convex, margins rather parallel in basal half to %, then gently

narrowed. Apex rather wide, transverse, truncature faintly convex, apical angles rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

moderately narrow, almost completely on visible from above. Basal border almost complete, ending

very closely to suture, becoming very weak there. Lateral border asetose. Series of umbilical pores

consisting of 6 pores behind shoulder and 1 additional pore behind middle. Setae moderately elongate.

Striae very faintly indicated as rows of extremely fine, irregular, longitudinal strioles. Microreticula-

tion fine, very superficial, though still visible, puncturation ine and rather sparse, becoming more

distinct and denser towards apex, surface impilose, glossy.

149

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin transverse, rather setose. Metepisternum elongate, c. 2 x as long as wide or even

slightly longer, neither bent nor hollowed. Abdominal sterna with 1 elongate setae on either side.

Sternum VI without longer setae along apical border. Lower surface moderately punctate and pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly wideer than long, tibial groove of

profemur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove

sharp. Femur wide. Metatibia medium-sized, c. 4.5 x as long as wide, 1st tarsomere of metatarsus

c. 1.6 x as long as wide. d protarsus not widened.

d genitalia (Figs 138e-k). Genital ring medium-sized, gently triangular, rather symmetric, arms

evenly convex, with rather narrow, fairly excised base. Sternum VII moderately wide, apically almost

straight, with very deep excision, basally regularly excised, lateral parts short. Aedeagus medium-

sized, moderately depressed, in middle slightly widened, slightly asymmetric, apex slighty turned

laterally. Lower surface gently convex. Apex wide, widely rounded off. Orifice rather short, internal

sac fairly complex, with a conspicuous, oblique, distinctly denticulate fold at apex. Right paramere

narrow, almost parallel, apex widely rounded. Left paramere considerably larger than right, slightly

triangular, apex rounded off.

? genitalia (Fig. 1381). Stylomere wide, but not markedly triangular, with wide, widely rounded

apex, with 2-4 subapical setae. Lateral plate elongate, with 2-3 long apical setae.

Variation. Little variation noted.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unknown. One specimen “ex sticky trap”. So far collected in April, August, and from

October to November. Occurs sympatrically and syntopically with A. similis, spec. nov.

Distribution (Fig. 614). Eastern Victoria, Tasmania, southeastern New South Wales to slightly north

of Sydney.

Material examined (8). Only the type series.

Etymology. The name refers to the smaller size compared with the closely related species A. haemorrhoidalis

Erichson.

Adelotopus nitens, spec. nov.

Figs 139, 326, 478, 614

Types. Holotype: ?, Australia, Qld. Mt. Glorious nr. Brisbane. 635 m. 9.11.X11.1988 H. & A. Howden (NMO).

Diagnosis. Medium-sized, elongate, convex, glossy black species with rather wide reddish apex of

elytra. Distinguished from all related species by markedly glossy surface, at least apically well marked,

punctate elytral striae, and complete absence of microreticulation on elytra. Further distinguished

from most similarly looking A. similis, spec. nov. by distinctly concave labrum, shorter and wide

antennae with Sth and 9th antennomeres >2 x as wide as long, and rather triagonal, apically mor acute

stylomere.

Description

Measurements. Length: 6.35 mm. Ratios. Width/length of pronotum: 1.58; width base/apex of

pronotum: 1.51; width pronotum/head: 1.56; length/width of elytra: 1.61; length elytra/pronotum:

2.62.

Colour (Fig. 326). Black, elytra with well defined red apex, the apical border of the spot distinctly

concave. Lower surface of head and thorax black to dark piceous, of abdomen light reddish. Mouth

parts, antennae, and legs dark reddish, tibiae and tarsi slightly darker.

Head (Figs 139a-d). Short and wide, moderately depressed. Anterior border fairly convex, lateral

angle rounded, laterally barely projecting, lateral borders faintly oblique. Clypeal suture rather

triagonal, not interrupted. Labrum large, apex distinctly concave. Antennal groove laterally sharply

bordered, latero-posteriorly without carinate area. Mental tooth triangular, short, apex acute. Wings

of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically slightly

triangular, ventrally with indistinct keel, at border with c. 16 elongate setae and additional pilosity on

150

Io,

Figs 139a-d, 1. Adelotopus nitens, spec. nov. Details of head and 2 genitalia. For legends see fig. 100.

upper and lower surface and along border. Terminal palpomere of maxillary palpus elongate, barely

widened, not securiform. Terminal palpomere of labial palpus rather wide, securiform. Antenna short,

Sth-9th antennomeres >2 x as wide as long. Microreticulation fine, rather superficial, puncturation

fine, though distinct, rather dense. Surface with a shallow sulcus medially of eyes, impilose, rather

glossy. Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula

apparently asetose.

Pronotum (Fig. 326). Moderately wide, convex, base fairly wide, apex narrow. Apical angles

moderately produced, at apex obtuse, somewhat oblique, almost attaining posterior third of eyes.

Apex moderately deeply excised, slightly convex in excision, indistinctly bordered. Sides evenly

curved throughout, widest in basal third or fourth. Margins narrow, not explanate, faintly bordered.

Basal angles evenly rounded off. Base almost straight, distinctly bordered. Surface near base without

transverse impression. Microreticulation very fine, highly superficial, difficult to see, puncturation

fine, though distinct, rather dense, surface impilose, glossy.

Elytra (Figs 326, 478). Elongate, convex, margins rather parallel in basal half to %, then gently

narrowed. Apex rather wide, transverse, truncature faintly convex, apical angles rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

moderately narrow, almost completely on visible from above. Basal border almost complete, ending

rather abruptly closely to suture, but even there rather distinct. Lateral border asetose. Series of

umbilical pores consisting of 6 pores behind shoulder and 1 additional pore behind middle. Setae

moderately elongate. Striae in apical half rather well marked as rows of fairly coarse, punctures, these

being considerably coarser than the other punctures. Microreticulation absent, puncturation fine and

rather sparse, becoming more distinct and denser towards apex, surface impilose, remarkably glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin transverse, rather setose. Metepisternum rather elongate, slightly <2 x as long as

wide, neither bent nor hollowed. Abdominal sterna with 1 elongate setae on either side. Sternum VI

without longer setae along apical border. Lower surface moderately punctate and pilose.

Legs. Medium-sized, 1st tarsomere of protarsus about as wide as long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, >4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.6 x as

long as wide. 3 protarsus unknown.

d genitalia. Unknown.

? genitalia (Fig. 139]). Stylomere wide, rather triangular, with fairly wide, rounded apex, with 3

subapical setae. Lateral plate elongate, with 2-3 long apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Holotype collected at height of 635 m in December.

Distribution (Fig. 614). Southeastern Queensland. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the highly glossy surface.

Adelotopus sparsepunctatus, spec. nov.

Figs 140, 327, 479, 614

Types. Holotype: ?, in garden, Atherton, Qd. 24.11.1975 J. H. Barrett, Adelotopus gyrinoides Hope det.

B.P. Moore '80 (QMB T26065). - Paratypes: 14, N. Queensland, Kuranda, 1100 ft., May 3-June 20, 1913,

R. E. Turner, 1913-438 (BMNH); 15, Millstream Falls, near Ravenshoe, N Old 9 Jan 1976, Collr. A & M Walford-

Huggins (CMP-WHC); 17, Australia: N. Qld 1 km NE of Tolga Jan. 1989 Storey & De Faveri light trap (DPIM);

12, Australien Old, Mareeba, 19.-25.X1I.1961, leg. H. Demarz (ZSM); 17, Old Rifle Ck., Mt. Molloy, 12-V-1987,

P. A. Meyer coll. (CBM); 1?, Australia, Qld 93/4, Mt. Molloy, 22.5.1993, M. Baehr (CBM); 1?, 15.475, 145.14,

Shiptons Flat Old, 17-19 Oct.1980, T. Weir (ANIC); 13, Australia, Qld 93/37, 5km w. Annan R., 35 km sw.

Cooktown, 3.6.1993, M. Baehr (CBM); 1?, Portland Road N. Old., 1. Jan. 1985 A & M Walford-Huggins,

Adelotopus sp. det. B. P. Moore’86 (CMP-CWH); 14, Austral., Coll. Hacker, det. haemorrhoidalis (DEIB).

Diagnosis. Medium-sized, convex, rather glossy black species with wide, well delimited reddish apex

of elytra, that is anteriorly straight. Distinguished from related species by convex shape, shortly

rounded basal angle of pronotum, elytra evenly narrowed from base, almost complete basal border of

elytra, reduced microreticulation and extremely fine puncturation of elytra, elongate, asymmetric

aedeagus with widely rounded apex, triangular left paramere and narrow stylomere.

Description

Measurements. Length: 5.55-6.3 mm. Ratios. Width/length of pronotum: 1.66-1.75; width base/

apex of pronotum: 1.53-1.58; width pronotum/head: 1.58-1.64; length/ width of elytra: 1.55-1.58; length

elytra/pronotum: 2.53-2.74.

Colour (Fig. 327). Black, elytra with rather wide, well defined red apex, the anterior border of

which is straight. Lower surface of head and thorax piceous, of abdomen reddish. Mouth parts,

antennae, and legs reddish, tibiae and tarsi slightly darker.

Head (Figs 140a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle angulately rounded, laterally slightly projecting, lateral borders distinctly oblique. Clypeal

suture semicircular, in middle slightly interrupted. Labrum large, apex faintly concave. Antennal

groove laterally sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth triangu-

lar, short, apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide,

tongue-like, apically convex, ventrally with indistinct keel, at border with c. 16 elongate setae and

additional pilosity on upper and lower surface and along border. Terminal palpomere of maxillary

palpus slightly widened, though not securiform. Terminal palpomere of labial palpus wide, securi-

form. Antenna short, Sth-9th antennomeres >2 x as wide as long. Microreticulation very fine, though

distinct, puncturation extremely fine, sometims difficult to detect even under high magnification,

rather sparse. Surface with a shallow sulcus medially of eyes, impilose, moderately glossy, slightly

silky. Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula

apparently asetose.

Pronotum (Fig. 327). Rather wide, convex, base moderately wide, apex narrower. Apical angles

rather produced, at apex obtuse, somewhat oblique, almost attaining posterior third of eyes. Apex

moderately deeply excised, slightly convex in excision, unbordered. Sides evenly curved throughout,

widest at base. Margins narrow, not explanate, faintly bordered. Basal angles shrtlly rounded off. Base

almost straight, faintly or irregularly bordered. Surface near base without transverse impression.

Microreticulation very fine, distinct, only slightly superficial, puncturation very fine, rather sparse,

surface impilose, fairly glossy.

Elytra (Figs 327, 479). Rather elongate, highly convex, margins evenly narrowed to apex, faintly

convex throughout. Apex rather narrow, slightly oblique, truncature faintly convex, apical angles

rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders.

Marginal channel rather narrow, partly concealed. Basal border almost complete, ending rather closely

to or immediately at scutellum. Scutellar seta absent. Lateral border asetose. Series of umbilical pores

consisting of 6 closely set pores behind shoulder and 1 additional pore behind middle. Setae

moderately elongate. Striae including sutural stria absent. Microreticulation almost absent, only

faintest traces visible, puncturation extremely fine and rather sparse, rather difficult to detect, surface

impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin depressed, slightly convex, rather setose. Metepisternum moderately elongate,

152

Figs 140a-l. Adelotopus sparsepunctatus, spec. nov. Details of head and genitalia. For legends see fig. 100.

c. 1.7 x as long as wide, in posterior third obliquely bent and rather hollowed. Abdominal sterna with

1 elongate setae on either side. Sternum VI without longer setae along apical border. Lower surface

rather punctate and pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, <4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.6 x as

long as wide. d protarsus not widened.

d genitalia (Figs 140e-k). Genital ring rather wide, fairly symmetric, though one arm more or less

sinuate, the other convex, with rather symmetric, little excised base. Sternum VII rather wide, apically

obliquely convex, with rather deep excision, basally bisinuately excised, lateral parts rather short.

Aedeagus rather elongate, fairly depressed, in middle barely widened, rather asymmetric, apical part

slightly turned laterally. Lower surface distinctly convex. Apex wide, widely rounded off. Orifice

rather elongate, internal sac fairly complex, with a large, oblique fold near apex. Right paramere

narrow, elongate, with shortly rounded apex, left wide, considerably larger than right, tapering, with

widely rounded apex.

? genitalia (Fig. 1401). Stylomere rather narrow and elongate, only slightly triangular, with

obliquely rounded apex, with 2-3 subapical setae. Lateral plate rather elongate, with 2 elongate apical

setae.

Variation. Apart from some differences of size there is some variation of shape and relative width

of pronotum and of distinctness of puncturation of elytra.

Vivipary. Not confirmed in the examined material.

Habits. Little known. Single specimens collected “under bark of gum-type eucalypt”, “by sweeping”,

“in garden”, specimens captured by me under bark of river gum or other gum-type eucalypts. So far

collected during the period from October to January, in March, May, and June.

Distribution (Fig. 614). Northeastern Quensland from Atherton Tableland to south of Cooktown.

Material examined (11). Only the type series.

Etymology. The name refers to the sparse puncturation of surface.

iS >

Figs 141a-c, 1. Adelotopus semilunatus, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Adelotopus semilunatus, spec. nov.

Figs 141, 328, 480

Types. Holotype: ?, Bunawang (?), 9/../40, X (ANIC).

Diagnosis. Medium-sized, slightly depressed, blackish species with wide, semilunar, well delimited,

reddish apex of elytra. Further distinguished from related species by laterally not produced head,

widely rounded basal angles of pronotum, almost complete basal border of elytra, rather conspicuous

mircroreticulation, and short and wide, apically obliquely transverse stylomere.

Description

Measurements. Length: 5.5 mm. Ratios. Width/length of pronotum: 1.65; width base/apex of

pronotum: 1.56; width pronotum/head: 1.73; length/width of elytra: 1.55; length elytra/pronotum:

252%

Colour (Fig. 328). Piceous-black, lateral margins of pronotum slightly reddish translucent, elytra

with rather wide, well defined, semilunar, red apex. Lower surface of head and thorax piceous, of

abdomen reddish. Mouth parts and legs reddish, tibiae and tarsi brown.

Head (Figs 141a-c). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle evenly rounded, laterally not projecting, lateral borders almost parallel. Clypeal suture semicir-

cular, in middle not interrupted. Labrum large, apex faintly concave. Antennal groove laterally sharply

bordered, latero-posteriorly with weakly carinate area. Mental tooth triangular, short, apex acute.

Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically

convex, ventrally with indistinct keel, at border with c. 16 elongate setae and additional pilosity on

upper and lower surface and along border. Terminal palpomere of maxillary palpus slightly widened,

though not securiform. Terminal palpomere of labial palpus wide, securiform. Antenna broken.

Microreticulation rather distinct and fairly coarse, puncturation invisible. Surface with a shallow

sulcus medially of eyes, impilose, comparatively dull. Ventrolaterally of eyes with a row of short setae.

Suborbital field impunctate and asetose. Gula apparently asetose.

Pronotum (Fig. 328). Rather wide, slightly depressed, base moderately wide, apex narrower.

Apical angles rather produced, at apex obtuse, somewhat oblique, surpassing posterior third of eyes.

Apex moderately deeply excised, slightly convex in excision, faintly bordered. Sides evenly curved

throughout, widest at base. Margins moderately wide, slightly explanate, faintly bordered. Basal

angles evenly rounded off. Base slightly convex, distinctly bordered throughout. Surface near base

without transverse impression. Microreticulation distinct, though slightly superficial, rather coarse,

puncturation extremely very fine, almost invisible, surface impilose, fairly glossy.

Elytra (Figs 328, 480). Rather elongate, on disk slightly depressed, margins evenly narrowed to

apex, laterally almost straight. Apex rather wide, slightly oblique, truncature faintly convex, apical

angles rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders.

Marginal channel rather narrow, partly concealed. Basal border almost complete, ending near

scutellum. Lateral border asetose. Series of umbilical pores consisting of 6 closely set pores behind

154

shoulder and 1 additional pore at or behind middle. Setae moderately elongate. Striae including

sutural stria absent, indicated only by traces of elongate lines. Microreticulation superficial, punctu-

ration very fine and sparse, surface impilose, glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin depressed, slightly convex, rather setose. Metepisternum moderately elongate,

c. 1.6 x as long as wide, in posterior third barely bent and hollowed. Abdominal sterna with 2 elongate

setae on either side. Sternum VI without longer setae along apical border. Lower surface rather

punctate and pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, c. 5x as long as wide, metatarsus unknown. d protarsus

unknown.

d genitalia. Unknown.

? genitalia (Fig. 1411). Stylomere rather wide and short, with wide, obliquely rounded apex, with

3-5 subapical setae, of which the median one is by far longest. Lateral plate rather elongate, with 3

elongate apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Unknown.

Distribution. Unknown, because the label is neither easily readible, nor it is known, where the -

doubtful - locality is located. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the semilunar reddish apex of the elytra.

Adelotopus kurandae, spec. nov.

Figs 39, 142, 329, 481, 613

Types. Holotype: S, Kuranda Oct, J. Sedlacek Collr. (QMB T26077).

Diagnosis. Rather small, parallel, convex, black species with wide, slightly triangular, well delimited,

reddish apex of elytra. Further distinguished from related species by laterally but slightly produced

head, narrow pronotum with shortly rounded basal angles, almost complete basal border of elytra,

reduced mircroreticulation, and short, rather symmetric aedeagus with widely rounded apex and

triangular parameres.

Description

Measurements. Length: 4.95 mm. Ratios. Width/length of pronotum: 1.51; width base/apex of

pronotum: 1.46; width pronotum/head: 1.47; length/width of elytra: 1.57; length elytra/pronotum:

2.42.

Colour (Figs 39, 329). Black, lateral margin of pronotum faintly reddish translucent, elytra with

rather wide, well defined red apex, the anterior border of which is slightly triangular. Lower surface

of head and thorax piceous, of abdomen reddish. Clypeus, mouth parts, antennae, and legs dark

reddish, tibiae and tarsi slightly darker.

Head (Figs 142a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle rounded, laterally faintly projecting, lateral borders slightly oblique. Clypeal suture semicircular,

in middle slightly interrupted. Labrum large, apex faintly concave. Antennal groove laterally sharply

bordered, latero-posteriorly with weakly carinate area. Mental tooth triangular, short, apex acute.

Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically

convex, ventrally with indistinct keel, at border with c. 16 elongate setae and additional pilosity on

upper and lower surface and along border. Terminal palpomere of maxillary palpus slightly widened,

though not securiform. Terminal palpomere of labial palpus wide, securiform. Antenna short, Sth-9th

antennomeres >2 x as wide as long. Microreticulation very fine, though distinct, somewhat superficial,

puncturation extremely fine, rather difficult to detect even under high magnification, rather sparse.

Figs 142a-k. Adelotopus kurandae, spec. nov. Details of head and Ö genitalia. For legends see fig. 100.

Surface with a shallow sulcus medially of eyes, impilose, moderately glossy, slightly silky. Ventrola-

terally of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula asetose.

Pronotum (Fig. 329). Rather narrow, highly convex, base rather narrow, slightly narrowed to apex.

Apical angles moderately produced, at apex obtuse, somewhat oblique, attaining posterior third of

eyes. Apex moderately excised, distinctly convex in excision, unbordered. Sides curved throughout,

though posteriorly almost straight, widest in basal third. Margins narrow, not explanate, faintly

bordered. Basal angles shortly, but evenly rounded off. Base almost straight, laterally faintly bordered.

Surface near base without transverse impression. Microreticulation very fine, rather superficial,

puncturation extremely fine and sparse, almost invisible, surface impilose, glossy.

Elytra (Figs 329, 481). Rather elongate, highly convex, margins almost parallel in basal half, faintly

narrowed to apex. Apex rather wide, slightly oblique, truncature convex, apical angles rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

narrow, partly concealed. Basal border almost complete, ending near scutellum. Lateral border

asetose. Series of umbilical pores consisting of 6 closely set pores behind shoulder and 1 additional

pore behind middle. Setae short. Striae including sutural stria absent. Microreticulation almost absent,

only faintest traces visible, puncturation fine and moderately sparse, becoming mor distinct towards

apex and here, arranged into irregular rows, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin depressed, slightly convex, rather setose. Metepisternum moderately elongate,

c. 1.7 x as long as wide, barely bent and hollowed. Abdominal sterna with 1 elongate setae on either

side. Sternum VI without longer setae along apical border. Lower surface rather punctate and pilose.

Legs. Medium-sized, 1st tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, c. 5x as long as wide, 1st tarsomere of metatarsus c. 1.6 x as

long as wide. d protarsus not widened.

3 genitalia (Figs 142e-k). Genital ring moderately wide, fairly symmetric, with rather asymmetric,

deeply excised base. Sternum VII rather wide, apically convex, with rather deep excision, basally

deeply excised, lateral parts rather short. Aedeagus rather short, in middle slightly widened, slightly

asymmetric, apical part slightly turned laterally. Lower surface faintly convex. Apex wide, widely

rounded off. Orifice rather elongate, internal sac fairly complex, with a large, oblique fold near apex.

Both parameres triangular, taperig to apex. Right paramere considerably shorter than left.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Unknown. The holotype collected in October.

156

Distribution (Fig. 613). Atherton Tableland, north Queensland. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the type locality.

Adelotopus bimaculatus Macleay, 1864

Diagnosis. Medium sized, rather elongate, very convex, black species with large, hourglass-shaped

red spot behind base of elytra. Further distinguished from similar A. languidus by narrower shape, less

triangular pronotum with less explanate margins, barely microreticulate, glossy elytra, and less

distinct microreticulation but more distinct puncturation on pronotum.

This species includes two subspecies, the nominate subspecies in northeastern Queensland from

Mackay to Atherton Tableland, and a southern subspecies in southeastern Queensland which is clearly

distinguished by shape, pattern and some minor characters of d and ? genitalia. There is, however,

a broad stripe between the nominate and the southern populations, where intermediate forms occur.

Adelotopus bimaculatus bimaculatus Macleay, 1864

Figs 143, 330, 482, 615

Adelotopus bimaculatus Macleay, 1864, p. 113; 1871, p. 94; Castelnau 1867, p. 33; 1868, p. 119; Gestro 1893, p. 287;

Blackburn 1901b, p. 113; Notman 1925, p. 8, 28; Csiki 1933, p. 1634; Darlington 1968, p. 241, 242; Moore et

al. 1987, p. 49.

Types. Lectotype (by present designation): d, Pt. Denison, Syntype, Adelotopus bimaculatus Macl. Pt. Denison

(ANIC-MMS). — Paralectotype: 1%, same data (ANIC-MMSS).

Type locality: “Pt. Denison”, Queensland.

Diagnosis. Medium-sized, rather elongate, very convex, black species with large, hourglass-shaped

red spot behind base of elytra, and rather wide, laterally convex pronotum with obtusely rounded

basal angles and not explanate margins. Distinguished from relative subspecies A. bimaculatus

angustior, subspec. nov. by wider shape, more evenly rounded lateral margins of pronotum, usually

hourglass-shaped, rarely posteriorly straight or even convex elytral spot, more distinct puncturation

of surface, and shorter parameres, the right paramere being perceptibly shorter than the left.

Description

Measurements. Length: 4.6-6.1 mm. Ratios. Width/length of pronotum: 1.74-1.82; width base/

apex of pronotum: 1.52-1.61; width pronotum/head: 1.59-1.71; length/ width of elytra: 1.44-1.50; length

elytra/pronotum: 2.56-2.64.

Colour (Fig. 330). Black, sometimes the narrow lateral margin of pronotum faintly reddish

translucent. Elytra with a large, more or less hourglass-shaped red spot behind base, usually

occupying the basal half to % of elytra. Apart from few exceptions, this spot does not reach the lateral

margin or even the marginal channel. Sometimes the posterior margin of this spot is almost straight,

the anterior border, however, is always deeply v-shaped. Lower surface of head and prothorax

reddish-piceous, of abdomen reddish. Mouth parts, antenna, and legs reddish.

Head (Figs 143a-d). Short and rather wide, moderately depressed. Anterior border gently convex,

lateral angle rounded, laterally slightly projecting, lateral borders oblique. Clypeal suture semicircular,

in middle widely interrupted or even absent. Labrum rather wide, apex feebly concave. Antennal

groove laterally sharply bordered, latero-posteriorly without carinate area. Mental tooth triangular,

short, apex acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa wide, tongue-like,

apically widely rounded, ventrally with indistinct keel, at border with c. 10-12 elongate setae and

additional pilosity on upper and lower surface and along border. Terminal palpomere of maxillary

palpus rather narrow, barely widened, not securiform. Terminal palpomere of labial palpus wide,

distinctly securiform. Antenna rather short, 8th-9th antennomeres slightly <2 x as wide as long.

Microreticulation very fine, slightly superficial, puncturation very fine, though distinct, moderately

dense, surface with a shallow sulcus medially of eyes, impilose, rather glossy. Ventrolaterally of eyes

with a row of short setae. Suborbital field impunctate and asetose. Both palpi rather sparsely setose,

gula almost asetose.

Pronotum (Fig. 330). Moderately wide, highly convex, base wide, apex fairly narrow. Apical

angles moderately produced, moderately acute, obtuse at apex, slightly oblique, surpassing posterior

border of eyes. Apex moderately excised, rather convex in excision, faintly bordered. Sides curved

throughout, widest usually slightly in front of basal angles. Margins very narrow, apically faintly

bordered. Basal angles c. 100°, obtusely rounded off. Base almost straight, usually not bordered.

Surface near base without transverse impression. Microreticulation very fine, highly superficial,

puncturation fine, though distinct, moderately dense, surface impilose, glossy.

Elytra (Figs 330, 482). Moderately elongate, convex, margins faintly convex throughout. Apex

rather wide, slightly oblique, truncature slightly convex, apical angles rounded off. Shoulders rounded

off, basal margin slightly oblique, without setae behind shoulders. Marginal channel narrow, basally

partly concealed. Basal border almost complete, absent only very close to suture, ending gradually.

Scutellar pores absent. Lateral border asetose. Series of umbilical pores consisting of 6 pores behind

shoulder and 1 additional pore behind middle. Setae rather elongate. Striae including sutural stria

absent, though in apical part inner striae more or less distinctly traceable by irregular rows of slightly

larger punctures. Microreticulation absent, traces only visible near margins and apex, puncturation

fine, rather sparse. Surface impilose, highly glossy.

Lower surface. Prosternal process moderately elongate, fairly wide, straight, apex moderately

elongate and wide, straight, shortly setose. Metepisternum rather short, c. 1.6 x as long as wide, in

posterior third obliquely bent and moderately hollowed. Abdominal sterna with 1 elongate setae on

either side. Sternum VI without elongate setae at apical border. Lower surface moderately punctate

and pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly shorter than wide, tibial groove of

profemur very deep, anterior plate widely overlapping the groove for almost apical %, posterior

border of groove sharp. Femur rather wide. Metatibia rather short, slightly >4 x as long as wide, 1st

tarsomere of metatarsus c. 1.8 x as long as wide. d protarsus not widened.

d genitalia (Figs 143e-k). Genital ring rather wide, triangular, feebly asymmetric, arms evenly

convex, with rather narrow, asymmetric base. Sternum VII rather wide, apically gently convex, with

fairly deep excision, basally gently concave, lateral parts moderately elongate. Aedeagus short,

moderately depressed, in middle rather widened, slightly asymmetric. Lower surface straight, near

apex gently convex. Apex rather wide, evenly rounded. Orifice rather short, internal sac fairly

complex, with a large, oblique, distinctly denticulate fold at apex. Both parameres moderately

elongate, right distinctly shorter than left, apex obtusely rounded. Left large, considerably larger ans

slightly longer than right, with widely rounded apex.

? genitalia (Fig. 143]). Stylomere short and wide, not triangular, with wide, rather oblique, faintly

convex apex, with 2-4 subapical setae. Lateral plate moderately elongate slightly shorter than in

southern subspecies, with 2-5 short apical setae.

Variation. Apart from considerable, not purely sexual variation of size, also shape of the elytral

spot, relative shape of pronotum and elytra, and degree of puncturation of surface varies to some

extent. In addition, the intermediate population in central eastern Queensland mentioned above is

generally slightly narrower and more elongate with less triangular pronotum.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unrecorded. Specimens collected by me under bark of river gum and certain other

gum-type eucalypts, some also at light. Other specimens collected in “gum forest”, “under bark”, and

on “Euc. trunk”, and in “F.1.T.” (flight intercept trap). So far collected from October until March, and

in June. Of the northern, typical population, however, rather few specimens have been recently

captured, although the species should be rather common according to the many records.

Distribution (Fig. 615). Northern Queensland from about Mackay to Atherton Tableland. The most

southerly record is slightly south of Sarina; records from “Cape York” and from Tasmania probably

wrong, a record from Western Australia certainly wrong. The area between Rockhampton and

Clermont in central eastern Queensland is inhabited by a population somewhat intermediate between

the nominate subspecies A. bimaculatus bimaculatus and the southern subspecies A. bimaculatus

angustior, but nevertheless more similar to the northern population.

158

Figs 143a-l. Adelotopus bimaculatus bimaculatus Macleay. Details of head and genitalia. For legends see fig. 100.

Figs 144i-k. Adelotopus bimaculatus angustior, subspec. nov. d parameres. For legends see fig. 100.

Material examined (106). Typical specimens: Tas: 17, 608, Tasmanie, A. Simson, Adelotopus sp., Adelotopus

bimaculatus Macl. det..., bimaculatus W.M.L. suivant Simson (IRSNB); 12, 608, Tasmanie, A. Simson, Adelotopus

bimaculatus Macl. det... (IRSNB). - Old: 234, Qld 36, 15 km sw. Sarina, 17.X1.1990, M. Baehr (CBM); 14, 18,

Bowen, Janson Acq. 1884 (MNHN); 19, Port Bowen 75.22, det. bimaculatus (BMNH); 18, 12, 608 4129, Bowen

A. Simson, Adelotopus bimaculatus Macl. By Simson’s number (SAMA); 14, 12, Pt. Denison, Adelotopus bimacu-

latus Macl. lectotype!, paralectotype! (ANIC-MMS); 27%, Port Denison, Ex Musaeo Mniszech, det. bimaculatus

Macleay j (MNHN); 22%, Port Denison, Coll. Castelnau, bimaculata ML.’ J.”, P. Denison (MCSN); 3? 2, Port

Denison, Coll. Castelnau, bimaculatus M. L. J.” (MCSN); 2? 2,322, Port Denison, Coll. Castelnau, det. bimaculatus

(MCSN); 235, 12, Townsville, det. bimaculatus, G. Bryant Coll. 1919 (BMNH); 1%, Townsville, G. Bryant Coll.

1919, Adelotopus bimaculatus Macl. (FMT); 18, 222, Townsville, 23.X.02, X.02, F. P. Dodd, G. Bryant Coll. 1919,

det. bimaculatus (BMNH); 18, 522, Townsville, 8.X.02, 25.X.02, F. P. Dodd (ANIC); 12, Townsville, 16.X.02,

F. P. Dodd, E. W. Ferguson Coll. (ANIC); 13, 12, Townsville 13.X1.02, F. P. Dodd, ephippiatus Newm. according

to Blackburn Adelotopus bimaculatus M. 1 7137 (SAMA); 18, 12, A. 2559, Townsville, 25.X.02, F. P. Dodd,

Adelotopus bimaculatus Macl. (SAMA); 14, 15 mi N. Townsville, J. Sedlacek Collector (CSB); 1? Charters Towers

196, J. & M. Sedlacek Collectors (BMH); 13, 12, 30 m. SSW. of Ayr, 10.X.50, E. F. Riek (ANIC); 1?, Hidden Vy.

nr. Paluma, 13.1.70, J. G. Brooks BM 88 (CMC); 12, Woodstock NQ. II1.52 E. Sutton A. Jackson, E. Sutton Coll.

(QMB); 12, Woodstock, N. Q., 11.X1.1951, E. Sutton (QMB); 12, 4 ml. Ck. Woodstock, 15.1.1975, J. M. Barrett,

Adelotopus bimaculatus Macl. Det. B. P. Moore '79 (DPIM); 18, Kuranda F. P. Dodd (SAMA); 14, Cairns det.

F. P. Dodd (SAMA); 13, Kuranda Atherton Tab. 11.53, J. G. Brooks, K 48755, bimaculatus Macl. (MCZ); 19, Davies

Ck. Rd. Atherton Tab. XI1.57, Darlingtons, det. bimaculatus (MCZ); 18, 17, Qld 93/48, Walsh R. 20 km ssw.

Mareeba, 7.V1.1993, M. Baehr (CBM); 14, 12, Qld 93/49, Walsh R. 8km e. Dimbulah, 7.-8.6.1993, M. Baehr

(CBM); 15,12, 11 km WSW of Petford, 18.X.-15.X1.1992, P. Scammell F.I.T. (DPIM); 15, Cap York, Soc. Ent. Belg.

Coll. PUTZEYS, R.LSc.N.B. 1.G. Coll. gen. (IRSNB); 15, 43585, Masters, det. bimaculatus (BMNH); 15, bimaculatus

Macl. sec. Masters, Ex Museo H. W. Bates 1892 (MNHN); 15, 12, Simson, det. bimaculatus (BMNH); 15, Thorey

1867, Nov. Holl. bor. (NHMW). - WA: 1%, Australia occid. 1192 (HNMB). - ?: 12, Thorey 1837 (NHMW).

Intermediate specimens: Qld: 17, Rockhampton, Coll. Castelnau, Rockhampton, det. bimaculatus (MCSN);

12, Rockhampton 111.58 Darlingtons, det. bimaculatus (MCZ); 458,922, Rockhampton, 26.-27.X1.1967, J. Sedlacek

Collector (BMH); 15, Rockhampton 26.X11.1967, J. & M. Sedlacek Collectors (BMH); 15, Rockhampton

16.X11.1977, J. Sedlacek Collector (CSB); 12, Rockhampt. Dämel, Coll. Kraatz, Adelotopus bimaculatus Macl. Id.

159

by T. G. Sloane (DEIB); 14, Rockhampton N. Australia Damell Higgins 10/7 (OUM); 14, Rockhampton (OUM);

1?, 32km N Rockhampton, 8.11.1964, J. Sedlacek Collector (BMH); 5??, Marmor T. G. S. X.24 (ANIC);

17, Edungalba CQ, 20.X.45, E. Adams E. Sutton, E. Sutton Coll. 1964 (QMB); 17, Edungalba XI1.1968 Smith leg.

(BMH); 15, 15 km s. Marlborough, 21.1.1982, M. Baehr (CBM); 15, Qld 26, Mackenzie R. 79 km n. Dingo, Fitzroy

Dev. Rd., 11.X1.1990, M. Baehr (CBM); 15, Qld G20, 5km n. Mackenzie R., Fitzroy Dev. Rd., 12.X1.1990, leg.

Gerstmeier (CBM); 27%, Qld 27, Rolf Ck., 134 km n. Dingo, Fitzroy Dev. Rd., 12.X1.1990, M. Baehr (CBM),;

1?, Qld 28, Isaac River, 171 km n. Dingo, Fitzroy Dev. Rd., 12.X1.1990, M. Baehr (CBM); 338, Qld 29, 215 km n.

Dingo, Fitzroy Dev. Rd., 12.X1.1990, M. Baehr (CBM); 15, 12, Qld 39, 25 km s. Funnel Ck., Clermont-Marlb. Rd.,

18.X1.1990, M. Baehr (CBM).

Note. This subspecies looks very similar to A. languidus, spec. nov. that occurs in the same area.

Namely large and wide specimens of A. bimaculatus bimaculatus are superficially very similar. Apart

from the differences in microreticulation and puncturation they can be distinguished also by the

narrower and usually not distinctly reddish margins of the pronotum that are always reddish in

A. languidus.

Adelotopus bimaculatus angustior, subspec. nov.

Figs 144, 331, 483, 615

Types. Holotype: d, Biggenden, X11.1973, H. Frauca (ANIC). — Paratypes: 1?, same data (ANIC); 12, Sydney,

R. Mus. Hist. Nat. Belg. 1.G. 12 595 (IRSNB); 13, Stanthorpe E. Sutton, 12.X.29, E. Sutton Coll. 1964 (QMB);

12, Gatton, 27.V.-31 F. A. Perkins (UOIC); 13, Gatton 26.1.35 (UOIC); 3?2, Mt. Maroon, s. Old. 400-700 m,

13.X11.81, M. Baehr (CBM); 17, Old G48, Tamrookum Ck., s. Beaudesert, 26.X1.1990, leg. Gerstmeier (CBM);

2??,Qld 2, 5km s. Ipswich, 4.X1.1990, M. Baehr (CBM); 18, 22%, Brisbane Illidge (UOIC); 12, Brisbane

J. Sedlacek Igt. (CSB); 14, 12, Bris. 9.X1.19, 22.X1.14 (OMB); 1?, Acacia Ridge, 19.IX.66 W. Freeland (UOIC);

27%, Acacia Ridge, Brisbane, E. C. Dahms, 11.111.65, 3.V1.65 (QMB); 14, Acacia Ridge, 15 mls from Brisbane,

6.1V.1966, J. A. Grant. B. M./CSIRO Expedition B. M. 1973-346 (BMNH); 18, Upper Brookfield 10.11.62,

G. Monteith (UQIC); 12, Greenbank 28.11.1965, T. Weir (UOIC); 222, Nth Pine R. 4.X11.62, G. Monteith (UOIC);

15,17, Redland B. RD 27.V111.33, J. G. Brooks Bequest, 1976 (ANIC); 15, 17, Moretonbay Stevens, Ex Musaeo

Chaudoir, det. bimaculatus Macleay j (MNHN); 17, Qld G44, 30 km n. Gympie, 22.X1.1990, M. Baehr (CBM);

13,37%, Wide Bay, Wide Bay, Coll. Castelnau, det. bimaculatus (MCSN); 1%, K 12234, Adelotopus bimaculatus

M‘L. W. Gayndah (AMS); 1%, Gayndah, IV.58, Darlingtons, det. bimaculatus (MCZ); 17, 16.X1.1986, Gayndah

V.R. Bejsak, Igt. (CBS); 17, Gayndah, K 7744, det. bimaculatus (AMS); 15, Qld 56, Spring Ck., 40 km sse

Biggenden, 21.X1.1990, M. Baehr (CBM); 14, Qld 51, Parker Ck., 20 km s. Miriam Vale, 20.X1.1990, M. Baehr

(CBM); 15, 422, Maryborough E. W. Fischer (SAMA); 1?, Qld 19, Burnett R., 10 km n. Eidsvold, 9.X1.1990,

M. Baehr (CBM); 19, 22, 15.X1.1986, 20 km N. of Eidsvold V. R. Bejsak, Igt. (CBS); 12, 20 km N. Eidsvold

V.R. Bejsak, Igt., Adelotopus spp. det. B. P. Moore’87 (MMS); 15, 15.X1.1986 Monto J. Sedlacek Igt. (CSB);

256,2? ?,Qld 20, Cania Gorge, 25 km nw. Monto, 9.-11.X1.1990, M. Baehr (CBM, ZSM); 17, Old 50, Calliope R.,

27 km se. Mt. Larcom, 20.X1.1990, M. Baehr (CBM); 12, Qld G38, Raglan Ck., 10 km nw. Mt. Larcom, 21.X1.1990,

leg. Gerstmeier (CBM); 27%, Gunaldy QO 4.X11.82, J. Sedlacek Collector (CSB); 17, Australia Mid Queensld.,

D. Sharp Coll. 1932 (BMNH); 15, 90 Adelotopus bimaculatus Macleay Queensland (OUM); 1%, 7287 Queensl.,

A. ephippiatus Newm., bimaculatus Macl. according to Sloane Adelotopus ephippiatus N. 17137 (SAMA),; 1%, 43867,

Thorey, bimaculatus Mac Leay Australia, Adelotopus Hope Westw. (MNHB); 14, Australien, Coll. B. Schwarzer

(SMF C 16275); 15, Aust. (OUM); 15, Adelot. bimaculat., Austral. MLeay, Adelotopus affinis Cast., det. bimaculatus

(MCZ); 18, Mitchelton 28.1.72, J. Sedlacek Collector (CSB); 1?, Bridwell Collection, A. maculipennis det.

T. L. Erwin (USNM).

Diagnosis. Distinguished from nominate subspecies A. bimaculatus bimaculatus by narrower shape,

less rounded lateral margins of pronotum, usually posteriorly convex or at least straight, rarely

hourglass-shaped elytral spot, even finer puncturation of surface, and longer parameres, the right

paramere being as long as the left.

Description

Measurements. Length: 4.8-6.0 mm. Ratios. Width/length of pronotum: 1.61-1.70; width base/

apex of pronotum: 1.43-1.49; width pronotum/ head: 1.50-1.58; length/ width of elytra: 1.55-1.63; length

elytra/pronotum: 2.57-2.69.

Colour (Fig. 331). Black or piceous-black, margins of pronotum commonly slightly reddish

translucent. Elytra with a fairly large, anteriorly v-shaped, posteriorly usually convex or at least

160

straight, but rarely hourglass-shaped, red spot behind base in basal half to % of elytra. Usually this spot

does not reach the lateral margin or even the marginal channel. Lower surface of head and prothorax

reddish-piceous, of abdomen reddish. Mouth parts, antenna, and legs reddish, femora commonly

slightly lighter than tarsi.

Head. Generally similar to nominate subspecies, though antenna usually slightly shorter and

wider, 8th and 9th antennomeres almost 2 x as wide as long, and puncturatiion of surface even finer,

almost invisible even under high magnification.

Pronotum (Fig. 331). In most respects similar to nominate subspecies, but altogether narrower,

laterally less evenly curved and less narrowed to apex. Usually also margin less incurved to basal

angles. Punturation of surface even finer than in nominate subspecies.

Elytra (Fig. 331, 483). In most respects similar to nominate subspecies, though altogether more

elongate and narrower and puncturation in basal half even finer.

Lower surface. Similar to nominate subspecies.

Legs. Similar to nominate subspecies.

d genitalia (Figs 144i-k). Genital ring and aedeagus rather similar to nominate subspecies. Both

parameres longer with longer apex, right almost as long or as long as left, apex almost acute. Left large,

with longer and less evenly rounded apex.

? genitalia. Rather similar to nominate subspecies, but lateral plate usually slightly longer than in

nominate subspecies.

Variation. There is some variation of relative shape, especially of pronotum, of shape of elytral spot,

and of degree of puncturationof surface. Altogether, however, a rather homogeneous subspecies.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unknown. Specimens collected by me under bark of river gum and other gum-type

eucalypts. So far collected from August through May. In southeastern Queensland a rather common

species, but rarely collected in larger numbers.

Distribution (Fig. 615). Southeastern Queensland from border to about Gladstone. An old record from

southern new South Wales (Sydney) is highly doubtful.

Material examined (69). Only the type series.

Etymology. The name refers to the narrower shape of this subspecies compared with the nominate subspecies.

Adelotopus languidus, spec. nov.

Figs 40, 145, 332, 484, 616

Types. Holotype: d,6 km SE of Mareeba, 17.1X.-2.X.1990, S. de Faveri, Malaise trap (QMB T26058). — Paratypes:

2??, same locality and collector, but 2.-16.X.1990 and 16.X.-19.X1.1990 (DPIM); 12, 6km SE of Mareeba,

12.-29.1.1991 S. de Faveri MDPI F.I.T. Site 36 (DPIM); 18, Australia, Old. 15 km E Mareeba 12.11.1989 H. &

A. Howden (NMO); 1%, 4 ml. Ck. Woodstock, Q 15-1-1977, J. H. Barrett, Adelotopus bimaculatus Macleay [Series

det. by A. Walford Huggins] (CMP-WHC); 1%, Delta Downs Stn. NQ. 2.X11.82 Walford-Hugins, Adelotopus

bimaculatus Macleay [Series det. by A. Walford Huggins] (CMP-WHC); 15, Australia, Qld 93/42, Sand Flat Ck.,

35 km s Palmer R., 4.-5.6.1993, M. Baehr (CBM); 1°, Australia, Old 93/43, Sand Flat Ck., 35 km s Palmer R., 1 km

s. Hwy. to Cooktown, 5.6.1993, M. Baehr (CBM); 14, Australia, Qld 93/73, 5 km e. Innot Hot Springs, 14.6.1993,

M. Baehr (CBM); 239,229, Australia: Mt. Garnet N. Q., 8.X1.88, B. P. Moore, det. bimaculatus (CMC); 389,2? 2,

Australia, Qld 93/61, Einasleigh R. 33 km w. Mt. Surprise, 10.-11.6.1993, M. Baehr (CBM); 236, Australia, Old

93/63, 20 km n. Einasleigh, 11.6.1993, M. Baehr (CBM); 1534, 322, Australia, Qld 93/64, Einasleigh R. 2 km

e. Einasleigh, 11.-12.6.1993, M. Baehr (ANIC, BMNH, CBM, MCZ, NMV, SAMA, ZSM); 13, 1%, Australia, Old

93/66, Copperfield R. 14 km n. Kidston, 12.6.1993, M. Baehr (CBM); 2? 2, Mc Kinnons Ck., 9 km ssw. The Lynd

Jet., 13.6.1993, M. Baehr (CBM); 18, Normanton Qld. 10 M, March, 1982, J. Sedlacek Collector (CSB);

13, Australia: Hidden Vy nr. Paluma, N.Q. 13.1.70, J. G. Brooks, under bark BM 88, det. bimaculatus (CMC);

?9, Paluma, N.Qld, 13 Jan 1970 Walford Huggins, Adelotopus bimaculatus Macleay [Series det. by A. Walford

Huggins] (CMP-WHC); 18, Australia, Queensland, Mutchilba, 4.1996, leg. A. Floren (CBM); 235, Australia,

Queensland, Einasleigh River, 33 km w. Mt. Surprise, 4.1996, leg. A. Floren (CBM).

Diagnosis. Medium sized, moderately elongate, rather convex, black species with large, rather

hourglass-shaped red spot behind base of elytra. Further distinguished from similar A. bimaculatus

Macleay by wider shape, more triangular pronotum with more explanate margins, distinctly microre-

ticulate, fairly dull elytra, and distinct microreticulation but much more indistinct puncturation on

pronotum.

Description

Measurements. Length: 5.0-6.25 mm. Ratios. Width/length of pronotum: 1.82-1.86; width base/

apex of pronotum: 1.62-1.66; width pronotum/ head: 1.67-1.71; length/width of elytra: 1.39-1.43; length

elytra/pronotum: 2.56-2.60.

Colour (Figs 40, 332). Black, the lateral margins of pronotum and elytra always distinctly reddish

translucent. Elytra with a rather large, more or less hourglass-shaped red spot behind base, usually

occupying the basal half to % of elytra. This spot does not reach the lateral margin or even the marginal

channel. Sometimes the posterior margin of this spot is almost straight, the anterior border, however,

is always deeply v-shaped. Lower surface of head and prothorax piceous, of abdomen reddish. Mouth

parts, antenna, and legs reddish.

Head (Figs 145a-d). Short and rather wide, moderately depressed. Anterior border gently convex,

lateral angle obtusely rounded, laterally slightly projecting, lateral borders oblique. Clypeal suture

semicircular, in middle widely interrupted. Labrum rather wide, apex feebly concave. Antennal

groove laterally sharply bordered, latero-posteriorly with weak carina. Mental tooth triangular, short,

apex acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa wide, tongue-like, apically

widely rounded, ventrally with indistinct keel, at border with c. 10-12 elongate setae and additional

pilosity on upper and lower surface and along border. Terminal palpomere of maxillary palpus

slightly widened, slightly securiform. Terminal palpomere of labial palpus very wide, markedly

securiform. Antenna rather short, $Sth-9th antennomeres slightly <2 x as wide as long. Microreticula-

tion very fine, distinct, puncturation extremely fine, sometimes difficult to detect even under high

magnification moderately dense, surface with a shallow sulcus medially of eyes, impilose, moderately

glossy to fairly dull, somewhat silky. Ventrolaterally of eyes with a row of short setae. Suborbital field

impunctate and asetose. Both palpi rather sparsely setose, gula almost asetose.

Pronotum (Fig. 332). Rather wide, moderately convex, base wide, apex fairly narrow. Apical

angles moderately produced, moderately acute, obtuse at apex, slightly oblique, surpassing posterior

border of eyes. Apex moderately excised, slightly convex in excision, laterally distinctly, in middle

faintly bordered. Sides curved throughout, widest almost at basal angles. Margins more or less widely

explanate, anteriorly faintly bordered. Basal angles c. 100°, obtuse. Base straight or faintly concave,

faintly bordered or not bordered. Surface near base without transverse impression. Microreticulation

fine and dense, distinct, puncturation very fine, sometimes almost invisible even under high magni-

fication, moderately dense, surface with some very faint strioles, impilose, rather dull, somewaht silky.

Elytra (Figs 332, 484). Moderately elongate, convex, margins faintly convex throughout. Apex

rather wide, slightly oblique, truncature almost straight, apical angles rounded off. Shoulders obtusely

rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel rather

narrow, almost completely visible from above, at most basally slightly concealed. Basal border almost

complete, absent only very close to suture, ending gradually. Scutellar pores absent. Lateral border

asetose. Series of umbilical pores consisting of 6 (rarely unilaterally 5 or 7) pores behind shoulder and

an additional pore behind middle that is rarely absent. Setae rather elongate. Striae including sutural

stria absent, though in apical part inner striae more or less distinctly traceable by irregular rows of

slightly larger, sometimes even rather coarse and somewhat rasp-like punctures. Microreticulation

fine, though distinct, puncturation basally extremely fine, rather sparse, apically denser. Surface

impilose, moderately glossy, slightly silky.

Lower surface. Prosternal process moderately elongate, fairly wide, straight, apex moderately

elongate and wide, straight, shortly setose. Metepisternum rather short, c. 1.6 x as long as wide, in

posterior third obliquely bent and moderately hollowed. Abdominal sterna with 1 elongate setae on

either side. Sternum VI without elongate setae at apical border. Lower surface moderately punctate

and pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly shorter than wide, tibial groove of

profemur very deep, anterior plate widely overlapping the groove for at least apical half, posterior

border of groove sharp. Femur rather wide. Metatibia rather short, slightly >4 x as long as wide, Ist

tarsomere of metatarsus c. 1.8 x as long as wide. d protarsus not widened.

162

Figs 145a-l. Adelotopus languidus, spec. nov. Details of head and genitalia. For legends see fig. 100.

S genitalia (Figs 145e-k). Genital ring rather wide, triangular, feebly asymmetric, arms evenly

convex, with rather narrow, asymmetric base, with elongate apex. Sternum VII wide, apically almost

straight, with fairly deep excision, basally gently concave, lateral parts moderately elongate. Aedeagus

short, moderately depressed, in middle rather widened, slightly asymmetric. Lower surface almost

straight. Apex rather wide, evenly rounded. Orifice rather short, internal sac fairly complex, with a

large, oblique, distinctly denticulate fold at apex. Both parameres moderately elongate, right as long

as left, apex obtusely rounded. Left large, considerably larger than right, with widely rounded apex.

? genitalia (Fig. 1451). Stylomere short and wide, not triangular, with wide, rather oblique, faintly

convex apex, with 2 subapical setae. Lateral plate moderately elongate, with 2 short apical setae.

Variation. Apart from some differences of size, of relative shape of pronotum and elytra, and of

degree of puncturation little variation noted.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unknown. Most specimens collected by me under bark of river gum and other gum-

type eucalypts, one specimen captured “under bark”, some specimens in Malaise trap. So far collected

in October, November, January, March, and June.

Distribution (Fig. 616). Northeastern Queensland from Mt. Spec near Ingham to north of Mt. Molloy

and inland across base of Cape York Peninsula to Normanton.

Material examined (50). Only the type series.

Etymology. The name refers to the rather dull, distinctly microreticulate surface.

163

Adelotopus clepsydra, spec. nov.

Figs 146, 333, 485

Types. Holotype: d, ? (unreadable), Nov. Holl. (NHRS).

Diagnosis. Small, rather elongate, very convex, dark piceous species with large, hourglass-shaped

reddish spot behind base of elytra, and moderately wide, laterally rather parallel pronotum with

obtuse basal angles and somewhat channeled margins. Distinguished from similarly patterned species

by smaller size, narrower shape, rather narow base of pronotum, glossy surface, narrow d genital ring,

rather elongate aedeagus, and narrow and remarkably parallel right paramere.

Description

Measurements. Length: 4.15 mm. Ratios. Width/length of pronotum: 1.65; width base/apex of

pronotum: 1.38; width pronotum/head: 1.48; length/width of elytra: 1.53; length elytra/pronotum:

25.

Colour (Fig. 333). Piceous to blackish, head and pronotum darker, elytra slightly lighter. Lateral

margins of prionotum and elytra distinctly reddish translucent. Elytra with hourglass-shaped reddish

spot behind base and occupying the basal %. Lower surface basally reddish-piceous, posteriorly

reddish. Mouth parts, antenna, and legs reddish.

Head (Figs 146a-d). Short and rather wide, moderately depressed. Anterior border evenly convex,

lateral angle rounded, laterally slightly projecting, lateral borders oblique. Clypeal suture semicircular,

in middle barely interrupted. Labrum rather narrow, apex deeply concave. Antennal groove laterally

sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth triangular, short, apex

acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa wide, tongue-like, apically

widely rounded, ventrally with indistinct keel, at border with c. 10-12 elongate setae and additional

pilosity on upper and lower surface and along border. Terminal palpomere of maxillary palpus rather

short, slightly widened, faintly securiform. Terminal palpomere of labial palpus short and wide,

distinctly securiform. Antenna very short, Sth-9th antennomeres >2.5 x as wide as long. Microreticu-

lation very fine, rather superficial, puncturation extremely fine, almost invisible even under high

magnification, rather sparse, surface with a shallow sulcus medially of eyes, impilose, glossy.

Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Both palpi

rather sparsely setose, gula almost asetose.

Pronotum (Fig. 333). Rather narrow, highly convex, base not much wider than apex. Apical angles

moderately produced, moderately acute, obtuse at apex, rather oblique, surpassing posterior border

of eyes. Apex moderately excised, rather convex in excision, not bordered. Sides feebly curved

throughout, widest in basal third. Margins rather narrow, slightly channelled, anteriorly faintly

bordered. Basal angles >100°, obtuse, not rounded off. Base almost straight, laterally faintly bordered.

Surface near base without transverse impression. Microreticulation very fine, highly superficial,

puncturation extremely fine, almost invisible even under high magnification, rather sparse, surface

impilose, glossy.

Elytra (Figs 333, 485). Moderately elongate, rather narrow, convex, margins almost parallel in

basal %. Apex rather wide, slightly oblique, truncature slightly convex, apical angles rounded off.

Shoulders rounded off, basal margin slightly oblique, without setae behind shoulders. Marginal

channel narrow, basally partly concealed. Basal border almost complete, absent only very close to

suture, ending gradually. Scutellar pore absent. Lateral border asetose. Series of umbilical pores

consisting of 6 pores behind shoulder and 1 additional pore behind middle. Setae rather elongate.

Striae including sutural stria absent, though in apical part inner striae more or less distinctly traceable

by wide-spaced, irregular rows of slightly larger punctures. Microreticulation absent, puncturation

extremely fine, almost invisible even under high magnification, sparse. Surface impilose, highly

glossy.

Lower surface. Prosternal process moderately elongate, fairly wide, straight, apex moderately

elongate, narrow, straight, shortly setose. Metepisternum rather elongate, almost 2 x as long as wide,

in posterior third obliquely bent and moderately hollowed. Abdominal sterna with I elongate setae on

either side. Sternum VI without elongate setae at apical border. Lower surface apparently rather

sparsely punctate and pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly shorter than wide, tibial groove of

164

Figs 146a-k. Adelotopus clepsydra, spec. nov. Details of head and J genitalia. For legends see fig. 100.

profemur very deep, anterior plate widely overlapping the groove for almost apical %, posterior

border of groove sharp. Femur rather wide. Metatibia rather short, c. 4 x as long as wide, metatarsus

absent. d protarsus not widened.

d genitalia (Figs 146e-k). Genital ring rather narrow, triangular, feebly asymmetric, arms faintly

convex, with rather narrow, asymmetric base. Sternum VII rather wide, apically gently convex, with

deep excision, basally bisinuate, lateral parts moderately elongate. Aedeagus fairly elongate, moder-

ately depressed, in middle slightly widened, slightly asymmetric. Lower surface straight, near apex

gently convex. Apex rather wide, evenly rounded. Orifice rather short, internal sac fairly complex,

with a large, oblique fold at apex. Right paramere elongate, narrow, almost parallel, apex obtusely

rounded. Left paramere very large, much larger than right, with widely rounded apex.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Unknown.

Distribution. Unknown. The label is unreadible, the type locality is Australia (“Nov. Holl.”).

Material examined (1). Only the holotype.

Etymology. The name refers to the hourglass-shaped elytral pattern.

multipunctatus-group

Diagnosis. Medium-sized, convex, completely black species or black species with red apex. Labrum

bisetose; glossa c. 10-12-setose; lateral margin of pronotum narrow, not explanate, basal angle obtuse

or shortly rounded off; basal border line of elytra incomplete, usually ending halfways to suture, rarely

slightly longer, sometimes not well visible; scutellar pore absent; lateral margin of elytra narrow,

without elongate setae behind shoulders; series of lateral pores with 6 subhumeral pores and 1

postmedian pore, very rarely (in A. debitor Darlington from New Guinea only) without the postmedian

pore; abdominal sterna with 1 ambulatory seta on either side; sternum VI without longer setae at apical

margin; tibiae, especially metatibia depressed; all femora including profemur wide and depressed;

aedeagus always rather wide, slightly asymmetric, with widely rounded apex; internal sac of aedeagus

complex, with oblique fold near apex; apical margin of ? tergum VIII without elongate setae.

Larvae. Ist instar larva known of 1 species.

165

Distribution. 12 species in eastern Queensland, the northernmost parts of the Northern Territory and

Western Australia, Papua New Guinea, the Solomon Islands (Guadalcanal), and Malaysia and Java.

Systematic position. This group is very similar to the politus-group, it is, however, more apomorphic

in the abbreviated basal border of the elytra. It is perhaps the adelphotaxon of the obsoletus-group, and

it is also closely related to the outstanding villosus-group.

Adelotopus multipunctatus, spec. nov.

Figs 147, 334, 486, 616

Types. Holotype: 3, 14.045, 131.59E, Ferguson R., 31 km SE by S of Pine Creek N. T. 14 Nov.1979, T. Weir

(ANIC). — Paratypes: 19, Australia: N. T. Arnhem Land Maningrida, 5 m, 19.11.1961, Gross, J. L. & M. Gressitt

Collectors (BMH); 12, Australia, NT 95/53, 3km w. Mary River, 30.8.1995, M. Baehr (CBM); 13, Australia,

NT 95/9, Litchfield NP, 30 km w. Batchelor, 6.8.1995, M. Baehr (CBM); 14, Australia, NT 95/51, Litchfield NP,

30 km w. Batchelor, 29.9.1995, M. Baehr (CBM); 334, NT 95/48, 30 km n. Edith River Cr., 28.8.1995, M. Baehr

(CBM); 1%, Katherine N. T. IV.71, J. Sedlacek Collector (CSB); 15, Australia: Northern Terr.: Katherine,

19-20.111.1971, J. &. M. Sedlacek, Colls. BISHOP Museum (BMH); 1%, Australien, NT, Katherine Gorge,

6.-8.11.1984, M. & B. Baehr (CBM); 15, Australia, N. T. Victoria R. Crossing 30 m, X-22-1962, Collectors E. 5. Ross

& D. A. Cavagnaro (CAS); 17, N.T. Victoria R crossing of Victoria Hwy, 11 Nov.1984 M. Malipatil (MNTD);

13, Australia, NT 95/44, Victoria River, 5 km w. Victoria River Cr., 25.8.1995, M. Baehr (CBM); 14, 12, Junction

of Ord & Dunham R. W. A. 19.11.1971. A. Allwood, 11420, 11421 (MNTD); 18, Australia, WA 95/26, Galvans

Gorge, 10 km sw. Mt. Barnett, 13.8.1995, M. Baehr (CBM); 17, W. Aust. Kimberley Lennard River Xing Gibb

River Road 17.295, 124.44E 4 April 1988 T. F. Houston 679-1 (WAM 94/878).

Diagnosis. Rather small to medium-sized, fairly wide, moderately convex, glossy black species.

Further distinguished from related species by the rather wide pronotum with moderate lateral

margins and widely rounded basal angles, incomplete basal border of elytra, complete absence of

microreticulation, fine and rather dense puncturation, highly glossy surface, and short and wide

aedeagus with very wide, rounded apex.

Description

Measurements. Length: 4.7-6.1 mm. Ratios. Width/length of pronotum: 1.74-1.83; width base/

apex of pronotum: 1.51-1.57; width pronotum/head: 1.53-1.59; length/ width of elytra: 1.38-1.44; length

elytra/pronotum: 2.50-2.57.

Colour. Glossy black, sometimes margins of pronotum and elytra reddish translucent. Lower

surface of head and thorax piceous, of abdomen reddish. Mouth parts, antennae, and legs reddish,

tibiae and tarsi slightly darker.

Head (Figs 147a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle rounded, laterally fairly projecting, lateral borders slightly oblique. Clypeal suture semicircular,

in middle interrupted, very inconspucous. Labrum rather large, apex faintly concave. Antennal groove

laterally sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth triangular, short,

apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like,

apically convex, ventrally with indistinct keel, at border with c. 16 elongate setae and additional

pilosity on upper and lower surface and along border. Terminal palpomere of maxillary palpus

slightly widened, though not securiform. Terminal palpomere of labial palpus wide, securiform.

Antenna short, $8th-9th antennomeres >2 x as wide as long. Microreticulation absent, puncturation

fine, fairly dense. Surface with a shallow sulcus medially of eyes, impilose, markedly glossy.

Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula asetose.

Pronotum (Fig. 334). Wide, moderately convex, base wide, markedly narrowed to apex. Apical

angles not much produced, at apex obtuse, somewhat oblique, surpassing posterior margin of eyes.

Apex moderately excised, convex in excision, very faintly bordered. Sides strongly and evenly curved

throughout, widest near base. Margins fairly wide, slightly explanate, faintly bordered. Basal angles

rather widely rounded off. Base slightly convex, faintly bordered. Surface near base without transverse

impression. Microreticulation absent, puncturation very fine, dense, surface impilose, highly glossy.

Elytra (Figs 334, 486). Moderately elongate, moderately convex, regularly narrowed to apex,

faintly convex throughout. Apex rather wide, slightly oblique, truncature faintly convex, apical angles

rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal

166

Figs 147a-l. Adelotopus multipunctatus, spec. nov. Details of head and genitalia. For legends see fig. 100.

channel rather narrow, partly concealed. Basal border incomplete, reaching about to median third

between lateral border and suture. Lateral border asetose. Series of umbilical pores consisting of 6

closely set pores behind shoulder and 1 additional pore behind middle. Setae rather short. Striae

including sutural stria absent. Microreticulation absent, puncturation very fine, rather dense, sometimes

in apical third arranged to irregular rows of slightly larger punctures, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly narow, straight, gently convex, apex slightly

widened, margin depressed, slightly convex, feebly setose. Metepisternum rather short, c. 1.5 x aslong

as wide, in posterior third obliquely bent and deeply hollowed. Abdominal sterna with 1 elongate seta

on either side. Sternum VI without longer setae along apical border. Lower surface rather densely

punctate and pilose.

Legs. Rather short, 1st tarsomere of protarsus distinctly wider than long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical half, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, <4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.3 x as

long as wide. d protarsus not widened.

S genitalia (Figs 147e-k). Genital ring rather narrow, barely asymmetric, with slightly asymmetric,

narrow, barely excised base. Sternum VII rather wide, apically almost straight, with rather deep

excision, basally excised, lateral parts rather short. Aedeagus short, rather convex, in middle slightly

widened, slightly asymmetric. Lower surface convex. Apex very wide, evenly rounded off. Orifice

moderate, internal sac fairly complex, with a large, oblique fold near apex. Both parameres triangular,

with shortly rounded apex, left paramere wide, considerably larger than right.

? genitalia (Fig. 1471). Stylomere rather wide, apex wide, shortly rounded off, with 2-3 elongate

subapical setae. Lateral plate moderately elongate, with 1-3 elongate apical setae.

Variation. Some variation noted in size and relative width of pronotum.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. A specimen collected by me under bark of gum-type eucalypt, another

species “on trunk of Eucalyptus sp. camaldulensis”. So far captured in October, November, and from

February to April.

Distribution (Fig. 616). Northern part of Northern Territory, Kimberley Division, adjacent northwest-

ern Australia.

167

Material examined (18). Only the type series.

Etymology. The name refers to the dense puncturation of the surface.

Adelotopus ovatus, spec. nov.

Figs 41, 148, 335, 487, 616

Types. Holotype: d, 15.395 144.31E Split Rock QLD 18 Aug.-16 Sep. 1993 Malaise Trap P. Zborowski &

5. Shattuck (QMB T26063). — Paratypes: 14, 15.395 144.31E Split Rock QLD Malaise Trap 26 June-16 July 1993

K. Halfpapp & S. De Faveri (DPIM); 15, 15.395 144.31E Split Rock QLD 13 Dec 1992-18 Feb 1993 Malaise Trap

P. Zborowski (CBM); 17, 15.395 144.31E Split Rock QLD 10 Oct-18 Nov 1993 Malaise Trap P. Zborowski &

M. Horak (DPIM); 12, 15.108 145.07E, 3.5 km SW by SMt. Baird QLD, 3-5 May 1981, A. Calder, at light (ANIC).

Diagnosis. Medium-sized, wide, moderately convex, glossy black species. Further distinguished from

related species by the wide pronotum with rather wide lateral margins and widely rounded basal

angles, short and regularly narrowed elytra, incomplete basal border of elytra, almost complete

absence of microreticulation, very fine and sparse puncturation, and glossy surface.

Description

Measurements. Length: 5.6-6.1 mm. Ratios. Width/length of pronotum: 2.03-2.06,; width base/

apex of pronotum: 1.64-1.68; width pronotum/ head: 1.73-1.78; length/ width of elytra: 1.28-1.33; length

elytra/pronotum: 2.60-2.69.

Colour. Black. Lower surface piceous-black. Mouth parts, antennae, and legs reddish-piceous.

Head (Figs 148a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle rounded, laterally fairly projecting, lateral borders distinctly concave and oblique. Clypeal

suture not visible. Labrum rather large, apex concave. Antennal groove laterally sharply bordered,

latero-posteriorly with weakly carinate area. Mental tooth triangular, short, apex acute. Wings of

mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically convex, ventral-

Iy with indistinct keel, at border with c. 16 elongate setae and additional pilosity on upper and lower

surface and along border. Terminal palpomere of maxillary palpus slightly widened, though not

securiform. Terminal palpomere of labial palpus very wide, markedly securiform. Antenna short, 8th-

9th antennomeres >2x as wide as long. Microreticulation very fine and somewaht superficial,

puncturation extremely fine, fairly sparse. Surface with a shallow sulcus medially of eyes, impilose,

rather glossy. Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose.

Gula asetose.

Pronotum (Fig. 335). Very wide, moderately convex, base wide, markedly narrowed to apex.

Apical angles not much produced, at apex obtuse, somewhat oblique, surpassing posterior margin of

eyes. Apex moderately excised, convex in excision, laterally faintly bordered, in middle unbordered.

Sides strongly and evenly curved throughout, widest in posterior third, even slightly narrowed to

base. Margins fairly wide, slightly explanate, faintly bordered. Basal angles rather widely rounded off.

Base straight, distinctly bordered. Surface near base without transverse impression. Microreticulation

almost absent, only finest traces visible, puncturation very fine, sparse, surface impilose, glossy.

Elytra (Fig. 335, 487). Rather short and wide, moderately convex, though depressed on disk,

regularly narrowed to apex, margin faintly convex throughout. Apex rather wide, slightly oblique,

truncature faintly convex, apical angles shortly rounded off. Shoulders rounded, basal margin slightly

oblique, without setae behind shoulders. Marginal channel fairly wide, barely concealed. Basal border

incomplete, reaching about halfways to suture. Lateral border asetose. Series of umbilical pores

consisting of 6 or 7 closely set pores behind shoulder and 1 additional pore behind middle. Setae rather

short. Striae including sutural stria absent, though inner striae in apical half faintly marked by

irregular rows of extrenely fine longitudinal lines. Microreticulation absent, puncturation extremely

fine, sparse, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex markedly

widened, margin depressed, slightly convex, feebly setose. Metepisternum rather short, c. 1.5 x aslong

as wide, in posterior third obliquely bent and deeply hollowed. Abdominal sterna with 1 elongate seta

on either side. Sternum VI without longer setae along apical border. Lower surface rather densely

punctate and pilose.

168

Figs 148a-l. Adelotopus ovatus, spec. nov. Details of head and genitalia. For legends see fig. 100.

Legs. Rather short, Ist tarsomere of protarsus distinctly wider than long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical half, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, slightly >4.5 x as long as wide, 1st tarsomere of metatarsus

c. 1.5 x as long as wide. Ö protarsus unknown.

d genitalia (Figs 148e-k). Genital ring rather narrow, barely asymmetric, with rather asymmetric,

narrow, slightly excised base. Sternum VII rather wide, apically almost straight, with deep excision,

basally faintly excised, lateral parts rather short. Aedeagus short, rather convex, in middle slightly

widened, almost symmetric. Lower surface faintly convex. Apex wide, evenly rounded off. Orifice

elongate, internal sac fairly complex, with a very strongly sclerotized, large fold and an oblique fold

near apex. Both parameres triangular, with obliquely rounded apex, left paramere wide, considerably

larger than right.

? genitalia (Fig. 1481). Stylomere rather short and wide, lateral border straight, apex wide,

obliquely cut off, with 2 elongate subapical setae. Lateral plate moderately elongate, with 2 elongate

apical setae.

Variation. Apart from some differences in size and shape of pronotum and elytra, very little

variation noted.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unknown. So far collected only in malaise traps and at light. Captured from May to

February.

Distribution (Fig. 616). Northeastern Queensland, in a very limited area at the base of the Cape York

Peninsula.

Material examined (5). Only the type series.

Etymology. The name refers to the oviform body shape.

169

Figs 149a-d, 1. Adelotopus browni, spec. nov. Details of head and % genitalia. For legends see fig. 100.

Adelotopus browni, spec. nov.

Figs 149, 336, 488, 617

Types. Holotype: ?, NT, Humpty Doo 12°35’S 131°05’E, Mt. Mortgage, 3 Dec 1991-17 Jan 92, Wells & Webber

(MNTD).

Diagnosis. Medium-sized, fairly narrow, rather convex, glossy black species. Further distinguished

from related species by the rather elongate shape, rather shortly rounded basal angles of pronotum,

incomplete basal border of elytra, complete absence of microreticulation, very fine and inconspicuous

puncturation, highly glossy surface, and the presence of an additional small seta on the lateral margin

of the stylomere.

Description

Measurements. Length: 5.3 mm. Ratios. Width/length of pronotum: 1.73; width base/apex of

pronotum: 1.46; width pronotum/head: 1.50; length/width of elytra: 1.55; length elytra/pronotum:

2.09.

Colour. Glossy black. Lower surface of head and thorax almost black, of abdomen bright reddish.

Mouth parts, antennae, and legs dark reddish, tibiae and tarsi darker.

Head (Figs 149a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle rounded, laterally fairly projecting, lateral borders slightly oblique. Clypeal suture semicircular,

in middle interrupted, very inconspucous. Labrum rather large, apex faintly concave. Antennal groove

laterally sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth triangular, short,

apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like,

apically convex, ventrally with indistinct keel, at border with c. 12 elongate setae and additional

pilosity on upper and lower surface and along border. Terminal palpomere of maxillary palpus

slightly widened, though not securiform. Terminal palpomere of labial palpus wide, securiform.

Antenna short, Sth-9th antennomeres c. 2.25 x as wide as long. Microreticulation absent, puncturation

extremely fine, fairly dense. Surface with a shallow sulcus medially of eyes, impilose, markedly glossy.

Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula asetose.

Pronotum (Fig. 336). Moderately wide, convex, base fairly wide, moderately narrowed to apex.

Apical angles not much produced, at apex obtuse, somewhat oblique, surpassing posterior margin of

eyes. Apex moderately excised, convex in excision, faintly bordered. Sides evenly curved throughout,

widest slightly in front of base. Margins rather narrow, barely explanate, faintly bordered. Basal angles

narrowly rounded off. Base straigth, distinctly bordered. Surface near base without transverse

impression. Microreticulation absent, puncturation extremely fine, moderately dense, surface impi-

lose, highly glossy.

Elytra (Figs 336, 488). Rather elongate, convex, slightly narrowed to apex, laterally faintly convex

170

throughout. Apex rather wide, slightly oblique, truncature faintly convex, apical angles rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

rather narrow, partly concealed. Basal border incomplete, slightly surpassing reaching middle

between lateral border and suture. Lateral border asetose. Series of umbilical pores consisting of 6

closely set pores behind shoulder and 1 additional pore behind middle. Setae rather short. Striae

including sutural stria absent, only indicated near apex by indistinct rows of slightly coarser

punctures. Microreticulation absent, puncturation extremely fine, moderately dense, near apex slightly

coarser, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly narrow, straight, gently convex, apex slightly

widened, margin depressed, slightly convex, feebly setose. Metepisternum c. 1.6 x as long as wide, in

posterior third obliquely bent and deeply hollowed. Abdominal sterna with 1 short seta on either side.

Sternum VI without longer setae along apical border. Lower surface rather densely punctate and

pilose.

Legs. Rather short, 1st tarsomere of protarsus distinctly wider than long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical half, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, c. 4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.4 x as

long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Fig. 149]). Stylomere rather wide, apex wide, shortly rounded off, with 2 elongate

subapical setae and 1 or 2 smaller seta(e) on the lateral margin. Lateral plate rather elongate, with 2

elongate apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. Holotype collected in “Malaise trap” in the period of December-January.

Distribution (Fig. 617). Northern part of Northern Territory. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. Named in honour of the curator who made available the specimen.

Adelotopus jacobsoni Ritsema, 1909

Figs 150, 337, 489, 657

Adelotopus jacobsoni Ritsema, 1909, p. 255; Seidlitz 1912, p. 150 (jacobsonis); Notman 1925, p. 8, 29; Csiki 1933,

p- 1635; Darlington 1968, p. 240.

Types. Holotype: 3, E. Jacobson, Tandjong Priok, Java Nov. 1908, Adelotopus jacobsoni Rits. type!, Adelotopus

jacobsoni Rits. Det. Ritsema, Holotypus (NNML).

Type locality. “Tandjong Priok”, Java.

Diagnosis. Medium-sized, convex, rather glossy black species with wide, well delimited reddish apex

of elytra. Distinguished from related species by convex shape, shortly rounded basal angle of

pronotum, elytra evenly narrowed from base, rediuced microreticulation and extremely fine punctu-

ration of elytra, wide, apically very convex d sternum VII with remarkbly long and acute lateral parts,

and aedeagus with conspicuously denticulate, oblique fold within apex of internal sac, and triangular

parameres.

Description

Measurements. Length: 5.9-6.35 mm. Ratios. Width/length of pronotum: 1.66-1.77; width base/

apex of pronotum: 1.54-1.55; width pronotum/head: 1.55-1.57; length/ width of elytra: 1.51-1.52; length

elytra/pronotum: 2.56-2.70.

Colour (Fig. 337). Black, margins of pronotum and elytra faintly dark reddish translucent. Elytra

with wide, well defined red apex, the anterior border of the spot distinctly concave. Lower surface of

head and thorax black to dark piceous, of abdomen reddish. Mouth parts, antennae, and legs reddish,

tibiae and tarsi slightly darker.

171

Figs 150a-l. Adelotopus jacobsoni Ritsema. Details of head and genitalia. For legends see fig. 100.

Head (Figs 150a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle shortly rounded, laterally slightly projecting, lateral borders distinctly oblique. Clypeal suture

semicircular, in middle rather widely interrupted. Labrum large, apex faintly concave. Antennal

groove laterally sharply bordered, latero-posteriorly without carinate area. Mental tooth triangular,

short, apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-

like, apically convex, ventrally with indistinct keel, at border with c. 12 elongate setae and additional

pilosity on upper and lower surface and along border. Terminal palpomere of maxillary palpus

slightly widened, though not securiform. Terminal palpomere of labial palpus rather wide, securiform.

Antenna short, Sth-9th antennomeres >2 x as wide as long. Microreticulation very fine, though

distinct, puncturation very fine, rather sparse. Surface with a shallow sulcus medially of eyes,

impilose, moderately glossy, slightly silky. Ventrolaterally of eyes with a row of short setae. Suborbital

field impunctate and asetose. Gula apparently asetose.

Pronotum (Fig. 337). Rather wide, convex, base moderately wide, apex narrower. Apical angles

rather produced, at apex obtuse, somewhat oblique, almost attaining posterior third of eyes. Apex

moderately deeply excised, slightly convex in excision, unbordered. Sides evenly curved throughout,

widest at base. Margins narrow, not explanate, faintly bordered. Basal angles shrtlly rounded off. Base

almost straight, faintly or irregularly bordered. Surface near base without transverse impression.

Microreticulation very fine, distinct, only slightly superficial, puncturation very fine, rather sparse,

surface impilose, fairly glossy.

Elytra (Figs 337, 489). Rather elongate, highly convex, margins evenly narrowed to apex, faintly

convex throughout. Apex rather narrow, slightly oblique, truncature faintly convex, apical angles

rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders.

Marginal channel rather narrow, partly concealed. Basal border incomplete, ending about halfways to

suture. Lateral border asetose. Series of umbilical pores consisting of 6 closely set pores behind

shoulder and 1 additional pore behind middle. Setae moderately elongate. Striae including sutural

stria absent. Microreticulation almost absent, only faintest traces visible, puncturation extremely fine

and rather sparse, rather difficult to detect, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin depressed, slightly convex, rather setose. Metepisternum moderately elongate, c.

1.7 x as long as wide, in posterior third obliquely bent and rather hollowed. Abdominal sterna with

172

1 elongate setae on either side. Sternum VI without longer setae along apical border. Lower surface

rather punctate and pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, <4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.6 x as

long as wide. d protarsus not widened.

d genitalia (Figs 150e-k). Genital ring medium-sized, asymmetric, one arm straight, the other

convex, with rather narrow, little excised base. Sternum VII wide, apically evenly convex, with deep

excision, basally regularly excised, lateral parts elongate and very acute. Aedeagus medium-sized,

fairly depressed, in middle slightly widened, slightly asymmetric, apical part slightly turned laterally.

Lower surface gently convex. Apex wide, widely rounded off. Orifice rather elongate, internal sac

fairly complex, with a conspicuously denticulate, oblique fold near apex. Both parameres wide,

triangular with obtuse, shortly rounded apex, left considerably larger than right.

? genitalia (Fig. 1501). Stylomere rather narrow and elongate, only slightly triangular, with

obliquely rounded apex, with 2 subapical setae. Lateral plate rather elongate, with 2 tiny apical setae.

Variation. Apart from some differences in relative width of pronotum little variation noted.

Vivipary. Not confirmed in the examined material.

Habits. Little known. The three specimens from the Castelnau Collection were collected “sous les

ecorces”; the tree species, however, is unknown.

Distribution (Fig. 657). Malaysia, Java.

Material examined (4). Malaysia: 234, 17, Malacca Coll. Castelnau (CBM, MCSN).- Java: 15, E. Jacobson,

Tandjong Priok, Java Nov. 1908, Adelotopus jacobsoni Rits. type!, holotype! (NNML).

Adelotopus geminus, spec. nov.

Figs 42, 151, 338, 490, 617

Types. Holotype: ?, Australien, Qld 44, Cooberrie, 15 km n. Yeppoon, 19.11.1990, M. Baehr (ANIC).

Diagnosis. Medium-sized, convex, rather glossy black species with wide, well delimited reddish apex

of elytra, that is anteriorly straight. Distinguished from most closely related A. laticaudatus, spec. nov.

by narrower pronotum, more distinct microreticulation of pronotum and elytra, shorter, anteriorly

slightly convex reddish apex of elytra, stylomere without additional short setae in middle of lateral

margin, and longer and basally less curved lateral plate of stylomere.

Description

Measurements. Length: 6.3 mm. Ratios. Width/length of pronotum: 1.66; width base/apex of

pronotum: 1.55; width pronotum/head: 1.58; length/width of elytra: 1.58; length elytra/pronotum:

2.67.

Colour (Figs 42, 338). Black, elytra with rather wide, well defined red apex, the anterior border of

which is straight. Lower surface of head and thorax dark piceous, of abdomen reddish. Mouth parts,

antennae, and legs reddish, tibiae and tarsi slightly darker.

Head (Figs 15la-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle angulately rounded, laterally slightly projecting, lateral borders distinctly oblique. Clypeal

suture semicircular, in middle slightly interrupted. Labrum large, apex faintly concave. Antennal

groove laterally sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth triangu-

lar, short, apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide,

tongue-like, apically convex, ventrally with indistinct keel, at border with c. 16 elongate setae and

additional pilosity on upper and lower surface and along border. Terminal palpomere of maxillary

palpus slightly widened, though not securiform. Terminal palpomere of labial palpus wide, securi-

form. Antenna short, 8th-9th antennomeres >2 x as wide as long. Microreticulation very fine, though

distinct, puncturation extremely fine, rather difficult to detect even under high magnification, rather

sparse. Surface with a shallow sulcus medially of eyes, impilose, moderately glossy, slightly silky.

Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula

apparently asetose.

Figs 151a-d, I. Adelotopus geminus, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Pronotum (Fig. 338). Rather wide, convex, base moderately wide, apex narrower. Apical angles

rather produced, at apex obtuse, somewhat oblique, almost attaining posterior third of eyes. Apex

moderately deeply excised, slightly convex in excision, unbordered. Sides evenly curved throughout,

widest at base. Margins narrow, not explanate, faintly bordered. Basal angles obtusely rounded off.

Base almost straight, laterally faintly bordered. Surface near base without transverse impression.

Microreticulation very fine, distinct, slightly superficial, puncturation very fine, rather sparse, surface

impilose, fairly glossy.

Elytra (Figs 338, 490). Rather elongate, highly convex, margins evenly narrowed to apex, faintly

convex throughout. Apex rather narrow, slightly oblique, truncature faintly convex, apical angles

rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders.

Marginal channel rather narrow, partly concealed. Basal border incomplete, ending rather abruptly

about halfways to suture. Lateral border asetose. Series of umbilical pores consisting of 6 closely set

pores behind shoulder and 1 additional pore behind middle. Setae moderately elongate. Striae

including sutural stria absent. Microreticulation very superficial, puncturation extremely fine and

sparse, rather difficult to detect, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin depressed, slightly convex, rather setose. Metepisternum moderately elongate,

c. 1.7 x as long as wide, in posterior third obliquely bent and rather hollowed. Abdominal sterna with

1 elongate setae on either side. Sternum VI without longer setae along apical border. Lower surface

rather punctate and pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, <4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.6 x as

long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Fig. 1511). Stylomere rather wide and short, with wide, rounded apex, with 2 subapical

setae. Lateral plate rather elongate, ventrally straight, with 2 elongate apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. The holotype collected in November under bark of gum-type eucalypt.

Distribution (Fig. 617). Central eastern Queensland. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the remarkable external similarity with A. laticaudatus, spec. nov.

Figs 152a-c, I. Adelotopus laticaudatus, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Adelotopus laticaudatus, spec. nov.

Figs 152,339, 491,617

Types. Holotype: ?, 11.455 142.35E Heathlands QLD 08 Dec 1992-19 Feb 1993 Malaise Trap P. Zborowski (QMB

T26059).

Diagnosis. Medium-sized, convex, rather glossy black species with very wide, well delimited reddish

apex of elytra, that is anteriorly somewhat sinuate and incised at suture. Distinguished from most

closely related A. geminus, spec. nov. by wider pronotum, more superficial microreticulation of

pronotum and elytra, stylomere with additional short setae in middle of lateral margin, and slightly

shorter and basally markedly curved lateral plate of stylomere.

Description

Measurements. Length: 6.3 mm. Ratios. Width/length of pronotum: 1.75; width base/apex of pro-

notum: 1.51; width pronotum/head: 1.61; length/ width of elytra: 1.49; length elytra/pronotum: 2.62.

Colour (Fig. 339). Black, elytra with wide, well defined red apex, the anterior border of which is

somewhat sinuate and prolonged posteriorly along suture. Lower surface of head and thorax dark

piceous, of abdomen reddish. Mouth parts, antennae, and legs piceous, tibiae and tarsi even slightly

darker.

Head (Figs 152a-c). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle angulately rounded, laterally slightly projecting, lateral borders distinctly oblique. Clypeal

suture semicircular, almost complete. Labrum large, apex faintly concave. Antennal groove laterally

sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth triangular, short, apex

acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically

convex, ventrally with indistinct keel, at border with c. 16 elongate setae and additional pilosity on

upper and lower surface and along border. Terminal palpomere of maxillary palpus slightly widened,

though not securiform. Terminal palpomere of labial palpus wide, securiform. Both antennae partly

broken, but even basal antennomeres very short and wide. Microreticulation very fine, somewhat

superficial, puncturation extremely fine, rather difficult to detect even under high magnification,

rather sparse. Surface with a shallow sulcus medially of eyes, impilose, rather glossy, slightly silky.

Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula asetose.

Pronotum (Fig. 339). Rather wide, convex, base rather wide, apex narrower. Apical angles rather

produced, at apex obtuse, somewhat oblique, almost attaining posterior third of eyes. Apex moderate-

ly deeply excised, convex in excision, unbordered. Sides evenly curved throughout, widest near base.

Margins narrow, not explanate, faintly bordered. Basal angles obtusely rounded off. Base almost

straight, in middle slightly produced, laterally faintly bordered. Surface near base without transverse

impression. Microreticulation extremely fine and superficial, difficutl to detect, puncturation very fine,

rather sparse, surface impilose, glossy.

Elytra (Figs 339, 491). Rather elongate, convex, margins parallel in anterior half, then gently

narrowed to apex. Apex rather wide, slightly oblique, truncature faintly convex, apical angles rounded

off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal

channel rather narrow, partly concealed. Basal border incomplete, ending gradually slightly medially

of middle of base. Lateral border asetose. Series of umbilical pores consisting of 6 closely set pores

behind shoulder and 1 pore behind middle. Setae moderately elongate. Striae including sutural stria

absent. Microreticulation extremely superficial, barely perceptible, puncturation extremely fine and

sparse, rather difficult to detect, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, rather depressed, apex wide

and rather short, margin depressed, rather setose. Metepisternum moderately elongate, c. 1.6 x aslong

as wide, in posterior third obliquely bent and rather hollowed. Abdominal sterna with 1 elongate setae

on either side. Sternum VI without longer setae along apical border. Lower surface rather punctate and

pilose.

Legs. Medium-sized, Ist tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, <4.5 x as long as wide, Ist tarsomere of metatarsus c. 1.4 x as

long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Fig. 1521). Stylomere rather wide, with wide, rounded apex, with 2 subapical setae and

an additional, short seta in middle of lateral margin. Lateral plate rather elongate, basally strongly

concave, with 2 elongate apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. Holotype collected during the period from December-February in “Malaise

trap”.

Distribution (Fig. 617). Northern part of Cape York Peninsula, northernmost Queensland. Known

only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the wide reddish apex of the elytra.

Adelotopus debitor Darlington, 1968

Figs 153, 340, 492, 618, 655

Adelotopus debitor Darlington, 1968, p. 241, fig. 154.

Types. Holotype (not seen): sex ?, Ne New Guina, Wau, Morobe Dist., 1200 m, Jan. 16.1961, Sedlacek (BMH). —

Paratype: d, Papua, Kokoda. Pitoki, 450 m, 24.111.1956, J. L. Gressitt Coll. Paratype Adelotopus debitor D. borrowed

by Bishop Mus. (MCZ).

Type localities: “Wau” and “Kokoda-Pitoki”, Papua New Guinea.

Diagnosis. Medium sized, wide, moderately convex, glossy black species with very wide, well

delimited, reddish apex of elytra the border of which is slightly oblique and faintly prolonged

posteriorly along suture. Further distinguished from related species by wide pronotum with evenly

rounded basal angles, characteristically tapering elytra with incomplete basal border, superficial

microreticulation, very fine puncturation, absence of postmedian marginal pore of elytra, and short

and wide, rather asymmetric aedeagus with convex lower surface and a triangular fold in internal sac.

Further distinguished from related A. nitidior, spec. nov. by presence of microreticulation on elytra,

larger aedeagus, and shorter and wider parameres; and from A. yorkensis, spec. nov. by larger size,

wider pronotum, and more asymmetric aedeagus.

Description

Measurements. Length: 5.15-5.85 mm. Ratios. Width/length of pronotum: 1.84-1.89; width base /

apex of pronotum: 1.54-1.58; width pronotum/ head: 1.60-1.65; length/ width of elytra: 1.49-1.50; length

elytra/pronotum: 2.77-2.83.

176

Figs 153a-l. Adelotopus debitor Darlington. Details of head and genitalia. For legends see fig. 100.

Colour (Fig. 340). Black, margins of pronotum and elytra reddish translucent, elytra with rather

wide, well defined red apex, the anterior border of which is slightly oblique and faintly prolonged

backwards along suture. Lower surface of head and thorax dark piceous, of abdomen reddish. Mouth

parts, antennae, and legs dark reddish to piceous, tibiae and tarsi piceous.

Head (Figs 153a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle angulately rounded, laterally slightly projecting, lateral borders distinctly oblique. Clypeal

suture semicircular, in middle slightly interrupted. Labrum rather large, apex faintly concave.

Antennal groove laterally sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth

triangular, short, apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa

wide, tongue-like, apically convex, ventrally with indistinct keel, at border with c. 16 elongate setae

and additional pilosity on upper and lower surface and along border. Terminal palpomere of maxillary

palpus slightly widened, though not securiform. Terminal palpomere of labial palpus rather wide,

securiform. Antenna short, 8th-9th antennomeres >2 x as wide as long. Microreticulation very fine and

superficial, puncturation extremely very fine, rather difficult to detect, fairly sparse. Surface with a

shallow sulcus medially of eyes, impilose, glossy. Ventrolaterally of eyes with a row of short setae.

Suborbital field impunctate and asetose. Gula apparently asetose.

Pronotum (Fig. 340). Wide, rather convex, base wide, markedly narrowed to apex. Apical angles

not much produced, at apex obtuse, somewhat oblique, just surpassing posterior margin of eyes. Apex

moderately excised, very convex in excision, laterally faintly bordered, in middle irregularly bordered.

Sides strongly and evenly curved throughout, widest in basal third or near base. Margins fairly wide,

slightly explanate, faintly bordered. Basal angles shortly though evenly rounded off. Base slightly

convex, distinctly bordered. Surface near base without transverse impression. Microreticulation

extremely fine and superficial, puncturation very fine, fairly sparse, surface impilose, glossy.

Elytra (Figs 340, 492). Moderately elongate, moderately convex, regularly narrowed to apex,

faintly convex throughout. Apex rather wide, slightly oblique, truncature faintly convex, apical angles

rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders.

Marginal channel rather narrow, partly concealed. Basal border incomplete, ending about halfways to

suture. Lateral border asetose. Series of umbilical pores consisting of 6 closely set pores behind

shoulder only. Setae rather short. Striae including sutural stria absent. Microreticulation very super-

ficial, puncturation extremely fine, moderately dense, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

1747,

rather short, margin depressed, slightly convex, feebly setose. Metepisternum moderately elongate,

c. 1.6 x as long as wide, in posterior third obliquely bent and deeply hollowed. Abdominal sterna with

1 elongate seta on either side. Sternum VI without longer setae along apical border. Lower surface

mderately punctate and pilose.

Legs. Rather short, 1st tarsomere of protarsus distinctly wider than long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, <4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.4 x as

long as wide. d protarsus not widened.

d genitalia (Figs 153e-k). Genital ring rather narrow, barely asymmetric, with slightly asymmetric,

fairly excised base. Sternum VII rather wide, apically almost straight, with rather deep excision, basally

faintly excised, lateral parts rather short. Aedeagus shorzt, rather convex, in middle widened, slightly

asymmetric. Lower surface convex. Apex rather wide, evenly rounded off. Orifice elongate, internal

sac fairly complex, with a large, oblique fold near apex. Right paramere short and rather narrow, with

shortly rounded apex, left paramere wide, considerably larger than right, with obliquely rounded

apex.

? genitalia (Fig. 1531). Stylomere rather wide, apex wide, convex, with 2 elongate subapical setae.

Lateral plate rather short, with 2-3 elongate apical setae.

Variation. Some variation noted only in relative width of pronotum and degree of microreticula-

tion of surface.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. One specimen captured in “Malaise Trap”. So far collected during the

period from January to March and from end of August to beginning of September.

Distribution (Figs 618, 655). Papua New Guinea.

Material examined (4). NG: 14, Papua, Kokoda. Pitoki, 450 m, 24.11.1956, J. L. Gressitt Coll. Paratype Adelotopus

debitor D. borrowed by Bishop Mus. (MCZ); 19,J. H. Sedlacek Malaise Trap. Bishop, New Guinea: SE Popondetta,

60 m, 31.VIII.-4.1X.63, Adelotopus cf. debitor D. det. Darlington 69 (BMH); 17, Bulolo, 15.1.-14.11.1979, 800 m,

J. Sedlacek (CBM); 13, N. Guinea NE, Kuper Ra. 1.80 m, 25 km SE Sanau (?), J. H. Sedlacek Coll. Adelotopus debitor

Darlington, det. G. Ball 1989 (BMH).

Adelotopus nitidior, spec. nov.

Figs 43, 154, 341, 493, 618

Types. Holotype: 13, 15.295, 145.16E, Mt. Cook Nat. P. Old, 11-12 Oct 1980, T. Weir (ANIC). — Paratypes:

19, Australia, Qld. 93/27, Lakeland NP, Cabbage Tree Ck., 20 km e. Old Laura, 30.5.1993, M. Baehr (CBM);

14, 15.165, 144.59E, 14 km W by N. Hope Vale Mission Qld, 7-10 May 1981, A. Calder (ANIC); 15, Australia, Old

93/29, Normanby R., 40 km w. Hope Vale, 31.5.1993, M. Baehr (CBM); 15, N. Queensld., Adelotopus haemorrhoi-

dalis ER., coll. Hacker (DEIB); 284, Endeavour R. Queensland (MMS); 14, 12, V. de Poll, Cooktown (ANIC);

1?, 15.105, 145.07E, 3.5 km SW by SMt. Baird Old, 3-5 May 1981, A. Calder, by sweeping (ANIC); 12, Australia,

Qld. 93/42, Sand Flat Ck., 30 km s. Palmer R., 1 km s. Hwy. to Cooktown, 5.6.1993, M. Baehr (CBM); 1%, Nth.

Qld. Mulligan Hwy Station Ck., 28:X1:1970 A. & M. Walford-Huggins 5375 (CMP-WHC); 18, Australia, Old 93/

4, Mt. Molloy, 22.5.1993, M. Baehr (CBM).

Diagnosis. Medium sized, wide, moderately convex, glossy black species with very wide, well

delimited, reddish apex of elytra the border of which is slightly oblique and faintly prolonged

posteriorly along suture. Further distinguished from related species by wide pronotum with evenly

rounded basal angles, characteristically tapering elytra with incomplete basal border, very superficial

or even absent microreticulation, very fine puncturation, and short and wide, rather asymmetric

aedeagus with convex lower surface and a triangular fold in internal sac. Further distinguished from

related A. debitor Darlington by presence of postmedian marginal pore of elytra, absence of microre-

ticulation on elytra, smaller aedeagus, and longer and narrower parameres; and from A. yorkensis, spec.

nov. by larger size, wider pronotum, and more asymmetric aedeagus.

Description

Measurements. Length: 5.25-5.8 mm. Ratios. Width/length of pronotum: 1.76-1.90; width base/

apex of pronotum: 1.50-1.61; width pronotum/ head: 1.54-1.64; length/ width of elytra: 1.46-1.55; length

178

Figs 154a-l. Adelotopus nitidior, spec. nov. Details of head and genitalia. For legends see fig. 100.

elytra/pronotum: 2.7-2.8.

Colour (Figs 43, 341). Black, margins of pronotum and elytra sometimes weakly reddish translu-

cent, elytra with rather wide, well defined red apex, the anterior border of which is slightly oblique

and faintly prolonged backwards along suture. Lower surface of head and thorax dark piceous, of

abdomen reddish-piceous, becoming reddish towards apex. Mouth parts, antennae, and legs dark

reddish to piceous, tibiae and tarsi piceous.

Head (Figs 154a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle angulately rounded, laterally slightly projecting, lateral borders distinctly oblique. Clypeal

suture semicircular, in middle slightly interrupted. Labrum rather large, apex faintly concave.

Antennal groove laterally sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth

triangular, short, apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa

wide, tongue-like, apically convex, ventrally with indistinct keel, at border with c. 16 elongate setae

and additional pilosity on upper and lower surface and along border. Terminal palpomere of maxillary

palpus slightly widened, though not securiform. Terminal palpomere of labial palpus rather wide,

securiform. Antenna short, $th-9th antennomeres >2 x as wide as long. Microreticulation very fine and

superficial, puncturation very fine, rather difficult to detect, fairly sparse. Surface with a shallow

sulcus medially of eyes, impilose, glossy. Ventrolaterally of eyes with a row of short setae. Suborbital

field impunctate and asetose. Gula apparently asetose.

Pronotum (Fig. 341). Wide, rather convex, base wide, markedly narrowed to apex. Apical angles

not much produced, at apex obtuse, somewhat oblique, just surpassing posterior margin of eyes. Apex

moderately excised, very convex in excision, laterally faintly bordered, in middle irregularly bordered.

Sides strongly and evenly curved throughout, widest in basal third or near base. Margins fairly wide,

slightly explanate, faintly bordered. Basal angles shortly though evenly rounded off. Base slightly

convex, distinctly bordered. Surface near base without transverse impression. Microreticulation

extremely fine and highly superficial, puncturation very fine, fairly sparse, surface impilose, glossy.

Elytra (Figs 341, 493). Moderately elongate, moderately convex, regularly narrowed to apex,

faintly convex throughout. Apex rather wide, slightly oblique, truncature faintly convex, apical angles

rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders.

Marginal channel rather narrow, partly concealed. Basal border incomplete, ending about halfways to

suture. Lateral border asetose. Series of umbilical pores consisting of 6 closely set pores behind

179

shoulder and 1 postmedian pore. Setae rather short. Striae including sutural stria absent. Microretic-

ulation virtually absent, puncturation extremely fine, moderately dense, surface impilose, highly

glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin depressed, slightly convex, feebly setose. Metepisternum moderately elongate,

c. 1.6 x as long as wide, in posterior third obliquely bent and deeply hollowed. Abdominal sterna with

1 elongate seta on either side. Sternum VI without longer setae along apical border. Lower surface

mderately punctate and pilose.

Legs. Rather short, Ist tarsomere of protarsus distinctly wider than long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, c. 4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.5 x as

long as wide. ö protarsus not widened.

Jg genitalia (Figs 154e-k). Genital ring rather narrow, barely asymmetric, with slightly asymmetric,

fairly excised base. Sternum VII rather wide, apically almost straight, with rather deep excision, basally

faintly excised, lateral parts rather short. Aedeagus small, short, rather convex, in middle widened,

slightly asymmetric. Lower surface convex. Apex rather wide, evenly rounded off. Orifice elongate,

internal sac fairly complex, with a large, oblique fold near apex. Right paramere rather narrow, with

shortly rounded apex, left paramere moderately wide, considerably larger than right, with obliquely

rounded apex.

? genitalia (Fig. 1541). Stylomere rather wide, apex wide, convex, with 2 elongate subapical setae.

Lateral plate rather short, with 2-3 elongate apical setae.

Variation. Rather variable with respect to relative width and shape of pronotum and elytra.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. Specimens collected by me under bark of river gums and other gum-type

eucalypts. One specimen collected “by sweeping”. So far captured in May, June, October, and

November.

Distribution (Fig. 618). Southeastern part of lower Cape York Peninsula from Mt. Molloy to Lakeland

NP, northeastern Queensland.

Material examined (13). Only the type series.

Etymology. The name refers to the more nitid surface in comparison with the closely related A. debitor

Darlington from New Guinea.

Adelotopus yorkensis, spec. nov.

Figs 155, 342, 494, 618

Types. Holotype: 4, 11.455, 142,35E Heathlands QLD 08 Dec 1992-19 Feb 1993 Malaise Trap P. Zborowski (QMB

T26066).

Diagnosis. Medium sized, wide, moderately convex, glossy black species with very wide, well

delimited, reddish apex of elytra the anterior border of which is concave. Further distinguished from

related species by moderately wide pronotum with evenly rounded basal angles, characteristically

tapering elytra with incomplete basal border, very superficial microreticulation, very fine punctura-

tion, and short and wide, rather symmetric aedeagus with convex lower surface and a triangular fold

in internal sac. Further distinguished from related A. debitor Darlington by smaller size, narrower

pronotum, presence of postmedian marginal pore of elytra, and symmetric aedeagus; and from and

A. nitidior, spec. nov. by smaller size, narrower pronotum, presence of traces of microreticulation on

elytra, and symmetric aedeagus.

Description

Measurements. Length: 4.85 mm. Ratios. Width/length of pronotum: 1.7; width base/apex of

pronotum: 1.49; width pronotum/head: 1.55; length/width of elytra: c. 1.48; length elytra/pronotum:

2.48.

180

Figs 155a-k. Adelotopus yorkensis, spec. nov. Details of head and 3 genitalia. For legends see fig. 100.

Colour (Fig. 342). Black, elytra with rather wide, well defined red apex, the anterior border of

which is slightly concave. Lower surface of head and thorax dark piceous, of abdomen reddish. Mouth

parts, antennae, and legs dark reddish to piceous, tibiae and tarsi piceous.

Head (Figs 155a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle angulately rounded, laterally slightly projecting, lateral borders distinctly oblique. Clypeal

suture semicircular, in middle interrupted. Labrum rather large, apex faintly concave. Antennal

groove laterally sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth triangu-

lar, short, apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide,

tongue-like, apically convex, ventrally with indistinct keel, at border with c. 16 elongate setae and

additional pilosity on upper and lower surface and along border. Terminal palpomere of maxillary

palpus slightly widened, though not securiform. Terminal palpomere of labial palpus rather wide,

securiform. Antenna very short, 8th-9th antennomeres almost 2.5 x as wide as long. Microreticulation

very fine, puncturation fine though distinct, fairly sparse. Surface with a shallow sulcus medially of

eyes, impilose, glossy. Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate

and asetose. Gula asetose.

Pronotum (Fig. 342). Rather wide, rather convex, base wide, markedly narrowed to apex. Apical

angles not much produced, at apex obtuse, somewhat oblique, just surpassing posterior margin of

eyes. Apex moderately excised, very convex in excision, laterally faintly bordered, in middle

irregularly bordered. Sides strongly and evenly curved throughout, widest in basal third. Margins

fairly wide, slightly explanate, faintly bordered. Basal angles shortly though evenly rounded off. Base

slightly convex, distinctly bordered. Surface near base without transverse impression. Microreticula-

tion extremely fine and superficial, puncturation very fine, fairly sparse, surface impilose, glossy.

Elytra (Figs 342, 494). Moderately elongate, moderately convex, regularly narrowed to apex,

faintly convex throughout. Apex rather wide, slightly oblique, truncature faintly convex, apical angles

rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders.

Marginal channel rather narrow, partly concealed. Basal border incomplete, ending about halfways to

suture. Lateral border asetose. Series of umbilical pores consisting of 6 closely set pores behind

shoulder and 1 postmedian pore. Setae rather short. Striae including sutural stria absent. Microretic-

ulation extremely fine and superficial, only traces visible, puncturation very fine, moderately dense,

surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin depressed, slightly convex, feebly setose. Metepisternum moderately elongate,

c. 1.7 x as long as wide, in posterior third obliquely bent and deeply hollowed. Abdominal sterna with

181

l elongate seta on either side. Sternum VI without longer setae along apical border. Lower surface

moderately punctate and pilose.

Legs. Rather short, 1st tarsomere of protarsus distinctly wider than long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, c. 4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.4 x as

long as wide. 3 protarsus not widened.

d genitalia (Figs 155e-k). Genital ring rather narrow, slightly asymmetric, with slightly asymmet-

ric, fairly excised base. Sternum VII wide, apically almost straight, with rather deep excision, basally

faintly excised, lateral parts very elongate. Aedeagus small, short, rather convex, in middle gently

widened, almost symmetric. Lower surface gently convex. Apex rather wide, evenly rounded off.

Orifice elongate, internal sac fairly complex, with a large, oblique fold near apex. Right paramere

rather narrow, with shortly rounded apex, left paramere rather wide, triangular, considerably larger

than right, with shortly rounded apex.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Largely unknown. Holotype collected in Malaise trap during the period from December to

February.

Distribution (Fig. 618). Northernmost part of Cape York Peninsula, northeastern Queensland. Known

only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to occurrence at the tip of the Cape York Peninsula.

Adelotopus convexicollis, spec. nov.

Figs 156, 343, 495, 618

Types. Holotype: 2, Intake Cairns 14.10.34, M. 170., Adelotopus apicalis Macl. dt. B. P. Moore '69, apicalis Macl.

1573, J. G. Brooks Bequest 1976 (ANIC).

Diagnosis. Rather small, parallel, highly convex, black species with rather wide, fairly well delimited,

reddish apex of elytra the anterior border of which is triangular. Further distinguished from related

species by laterally but slightly produced head, narrow pronotum with narrow lateral margins and

obtusely rounded basal angles, incomplete basal border of elytra, reduced microreticulation, moder-

ately dense puncturation, and wide stylomere with widely rounded apex.

Description

Measurements. Length: 4.85 mm. Ratios. Width/length of pronotum: 1.52; width base/apex of

pronotum: 1.48; width pronotum/head: 1.56; length/width of elytra: 1.64; length elytra/pronotum:

2.97:

Colour (Fig. 343). Black, elytra with rather wide, moderately well defined red apex, the anterior

border of which is triangular. Lower surface of head and thorax dark piceous, apical half of abdomen

reddish. Mouth parts (as far as they are present) dark reddish, legs piceous.

Head (Figs 156a-c). Rather short and wide, moderately depressed. Anterior border gently convex,

lateral angle rounded, laterally feebly projecting, lateral borders faintly oblique. Clypeal suture

semicircular, in middle slightly interrupted. Labrum rather large, apex faintly concave. Antennal

groove laterally sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth triangu-

lar, short, apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide,

tongue-like, apically convex, ventrally with indistinct keel, at border with c. 16 elongate setae.

Terminal palpomere of maxillary palpus slightly widened, though not securiform. Terminal pal-

pomere of labial palpus rather wide, securiform. Antenna broken. Microreticulation very fine, slightly

superficial, puncturation extremely fine, rather difficult to detect, fairly sparse. Surface with a shallow

sulcus medially of eyes and some fine wrinkles, impilose, rather glossy, slightly silky. Ventrolaterally

of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula apparently asetose.

182

Figs 156a-c, 1. Adelotopus convexicollis, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Pronotum (Fig. 343). Rather narrow, highly convex, base rather narrow, narrowed to apex. Apical

angles moderately produced, at apex obtuse, somewhat oblique, almost attaining posterior third of

eyes. Apex moderately excised, convex in excision, laterally faintly bordered, in middle unbordered.

Sides evenly curved throughout, widest near base. Margins very narrow, not explanate, faintly

bordered. Basal angles obtusely rounded off. Base almost straight, unbordered. Surface near base

without transverse impression. Microreticulation very fine and superficial, puncturation fine, moder-

ately dense, surface impilose, glossy.

Elytra (Figs 343, 495). Rather elongate, convex, margins basally almost parallel, apically evenly

narrowed to apex. Apex rather wide, slightly oblique, truncature convex, apical angles rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

rather narrow, partly concealed. Basal border incomplete, ending about at the inner third between

lateral border and suture. Lateral border asetose. Series of umbilical pores consisting of 6 closely set

pores behind shoulder and 1 aditional pore behind middle. Setae presumably short. Striae including

sutural stria absent. Microreticulation almost absent, only faintest traces visible, puncturation fine,

moderately dense, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin depressed, slightly convex, slightly setose. Metepisternum moderately elongate,

c. 1.6 x as long as wide, in posterior third obliquely bent and rather deeply hollowed. Abdominal

sterna with 1 elongate seta on either side. Sternum VI without longer setae along apical border. Lower

surface moderately punctate and pilose.

Legs. Rather short, Ist tarsomere of protarsus slightly wider than long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, c.4.5 x as long as wide, Ist tarsomere of metatarsus c. 1.4 x as

long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Fig. 1561). Stylomere rather wide, apex wide, widely rounded off, with 2 elongate

subapical setae. Lateral plate rather short, with 2 elongate apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Holotype collected in October.

Distribution (Fig. 618). Northeastern Queensland. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the highly convex prothorax.

Adelotopus gibbosus, spec. nov.

Figs 157, 344, 496, 619

Types. Holotype: d, Kuranda, N.Q. 21.3.52 C. Oke (NMV).

Diagnosis. Rather small, moderately wide, markedly convex, rather dull black species with wide

reddish apex, the anterior border of which is slightly oblique. Further distinguished from related

species by the very convex pronotum with narrow lateral margins and shortly rounded basal angles,

incomplete basal border of elytra, distinct microreticulation, very fine puncturation, markedly dull

surface, and short and rather wide aedeagus with moderately wide, rounded apex.

Description

Measurements. Length: 4.5 mm. Ratios. Width/length of pronotum: 1.68; width base/apex of

pronotum: 1.52; width pronotum/head: 1.57; length/width of elytra: 1.50; length elytra/pronotum:

Colour (Fig. 344). Black, margins of pronotum very faintly reddish translucent, elytra with rather

wide, well defined red apex, the anterior border of which is slightly oblique. Lower surface of head

and thorax piceous-black, of abdomen reddish reddish. Mouth parts, antennae, and legs reddish

piceous.

Head (Figs 157a-d). Rather short and wide, moderately depressed. Anterior border gently convex,

lateral angle obtusely rounded, laterally slightly projecting, lateral borders but faintly oblique. Clypeal

suture semicircular, though very inconspicuous. Labrum rather wide, apex feebly concave. Antennal

groove laterally sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth triangu-

lar, short, apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide,

tongue-like, apically convex, ventrally with indistinct keel, at border with c. 16 elongate setae.

Terminal palpomere of maxillary palpus slightly widened, not securiform. Terminal palpomere of

labial palpus wide, securiform. Antenna rather short, Sth-9th antennomeres c. 2x as wide as long.

Microreticulation very fine, though distinct, puncturation barely visible. Surface with a shallow sulcus

medially of eyes, impilose, rather dull, slightly silky. Ventrolaterally of eyes with a row of short setae.

Suborbital field impunctate and asetose. Gula apparently asetose.

Pronotum (Fig. 344). Moderately wide, markedly convex, base moderately wide, narrowed to

apex. Apical angles not much produced, at apex obtuse, somewhat oblique, just surpassing posterior

margin of eyes. Apex moderately excised, markedly convex in excision, laterally faintly bordered.

Sides evenly curved throughout, widest near base. Margins rather narrow, not explanate, faintly

bordered. Basal angles rather shortly rounded off. Base feebly concave, unbordered. Surface near base

without transverse impression. Microreticulation very fine, though distinct, puncturation fine, fairly

dense, surface with some faint irregular wrinkles, impilose, rather dull, slightly silky.

Elytra (Figs 344, 496). Moderately elongate, highly convex, regularly narrowed to apex, faintly

convex throughout. Apex fairly wide, rather oblique, truncature slightly convex, apical angles rounded

off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal

channel rather narrow, partly concealed. Basal border incomplete, reaching about halfways to suture.

Lateral border asetose. Series of umbilical pores consisting of 5 or 6 closely set pores behind shoulder

and 1 additional pore behind middle. Setae rather short. Striae including sutural stria absent.

Microreticulation fine, distinct, though slightly superficial, puncturation very fine, sparse, surface

impilose, fairly dull, slightly silky.

Lower surface. Prosternal process moderately short, rather narrow, straight, apex slishtly wid-

ened, margin depressed, slightly convex, rather setose. Metepisternum moderately short, c. 1.6 x as

long as wide, in posterior third faintly obliquely bent and slightly hollowed. Abdominal sterna with

l elongate seta on either side. Sternum VI without longer setae along apical border. Lower surface

rather densely punctate and pilose.

Legs. Rather elongate, Ist tarsomere of protarsus distinctly longer than wide, tibial groove of

profemur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove

sharp. Femur moderately wide. Metatibia moderately elongate, c. 5 x as long as wide, 1st tarsomere

of metatarsus c. 1.8 x as long as wide. d protarsus not widened.

d genitalia (Figs 157e-k). Genital ring fairly wide, roundish, barely asymmetric, with slightly

asymmetric, narrow, barely excised base. Sternum VII rather wide, apically convex, with deep

184

Figs 157a-k. Adelotopus gibbosus, spec. nov. Details of head and d genitalia. For legends see fig. 100.

excision, basally almost straight, lateral parts acute and elongate. Aedeagus rather short, convex, in

middle widened, almost symmetric. Lower surface gently convex. Apex rather narrow, rounded off.

Orifice short, internal sac fairly complex, with a distinct oblique fold near apex. Right paramere

triangular, rather narrow, with shortly rounded apex, left paramere considerably larger than right,

wide, with widely rounded apex.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Unknown. Holotype collected in March.

Distribution (Fig. 619). Northeastern Queensland. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the convex, gibbose shape.

Adelotopus penelopeae, spec nov.

Figs 158, 345, 497, 655

Types. Holotype: ? (defect-abdomen absent), Solomon Is. Guadalcanal, Mt. Austen 14/5,1963, P. Greenslade

6020, Solomon Is. Pres. P. J. M. Greenslade B. M. 1966-477 (BMNH).

Diagnosis. Medium-sized, fairly wide, moderately convex, black species with rather wide, fairly well

delimited, slightly semilunar reddish apex of elytra. Further distinguished from related species by

laterally strongly produced head, rather wide pronotum with moderate lateral margins and obtusely

rounded basal angles, incomplete basal border of elytra, almost absent microreticulation, very fine and

moderately dense puncturation, and highly glossy surface.

Description

Measurements. Length: 7.35 mm. Ratios. Width/length of pronotum: 1.72; width base/apex of

pronotum: 1.61; width pronotum/head: 1.67; length/width of elytra: c. 1.52; length elytra/pronotum:

2.67.

Colour (Fig. 345). Black, margins of pronotum and elytra faintly reddish translucent, elytra with

rather wide, slightly semilunar, well defined red apex. Lower surface of head and thorax dark piceous,

of abdomen presumably reddish. Mouth parts, antenna, and legs dark reddish.

Figs 158a-d. Adelotopus penelopeae, spec. nov. Details of head. For legends see fig. 100.

Head (Figs 158a-d). Very short and wide, moderately depressed. Anterior border almost straight,

lateral angle angulately rounded, laterally markedly projecting, lateral borders distinctly oblique-

concave. Clypeal suture almost invisible. Labrum rather large, apex faintly concave. Antennal groove

laterally sharply bordered, latero-posteriorly with weakly carinate area. Mental tooth triangular, short,

apex acute. Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like,

apically convex, ventrally with indistinct keel, at border with c. 16 elongate setae. Terminal palpomeres

of maxillary palpus slightly widened, though not securiform. Terminal palpomere of labial palpus

wide, securiform. Antenna short, 8th-9th antennomeres c. 2 x as wide as long. Microreticulation very

fine, highly superficial, difficult to detect, puncturation extremely fine, fairly sparse, difficult to detect

even under high magnification. Surface with a shallow sulcus medially of eyes, impilose, highly glossy.

Ventrolaterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula asetose.

Pronotum (Fig. 345). Rather wide, fairly convex, base rather wide, markedly narrowed to apex.

Apical angles moderately produced, at apex obtuse, somewhat oblique, almost attaining posterior

third of eyes. Apex moderately excised, convex in excision, laterally faintly bordered, in middle

unbordered. Sides evenly curved throughout, widest near base. Margins rather narrow, barely

explanate, faintly bordered. Basal angles obtusely rounded off. Base almost straight, unbordered.

Surface near base without transverse impression. Microreticulation extremely fine and superficial,

barely recognizable even under high magnification, puncturation equally extremely fine, rather sparse,

surface with some very fine wrinkles, impilose, highly glossy.

Elytra (Figs 345, 497). Rather wide, moderately elongate, fairly convex, margins faintly, but evenly

narrowed to apex. Apex rather wide, slightly oblique, truncature strraight, apical angles rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

moderately wide, little concealed. Basal border incomplete, ending about halfways to suture. Lateral

border asetose. Series of umbilical pores consisting of 6 closely set pores behind shoulder and

l additional pore behind middle. Setae presumably short. Striae including sutural stria absent.

Microreticulation almost absent, only faintest traces visible, puncturation extremely fine, difficult to

detect, moderately dense, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin depressed, square, slightly setose. Metepisternum moderately elongate, c. 1.6 x as

long as wide, in posterior third markedly obliquely bent and deeply hollowed. Abdominal sterna

unknown. Lower surface moderately punctate and pilose.

Legs. Medium-sized, 1st tarsomere of protarsus about as wide as long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, <5 x as long as wide, 1st tarsomere of metatarsus c. 1.6 x aslong

as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Holotype collected in May.

Distribution (Fig. 655). Guadalcanal, Solomon Islands. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. Named in honour of the collector of the holotype: Mrs. Penelope Greenslade.

y 8y yF F

186

obsoletus-group

Diagnosis. Medium-sized, moderately convex, dull black species. Whole surface with fine, though

distinctmicroreticulation; labrum very narrow, mostly concealed, bisetose; glossa c. 16-setose; lateral

margin of pronotum narrow, not explanate, basal angle rounded off; basal border line of elytra

incomplete, reaching halfways to suture, ending abruptly; scutellar pore absent; lateral margin of

elytra narrow, without elongate setae behind shoulders; series of lateral pores with 6 subhumeral

pores and 1 postmedian pore; abdominal sterna without ambulatory setae; sternum VI without longer

setae at apical margin; tibiae, especially metatibia depressed; all femora including profemur wide and

depressed; aedeagus moderately wide, highly asymmetric, with acute apex, in front of apex deeply

sinuate; internal sac of aedeagus complicate, without oblique fold near apex.

Larva. Unknown.

Distribution. A single species in central eastern to northeastern Queensland.

Systematic position. This group is probably the adelphotaxon of the multipunctatus-group, but is

more apomorphic in the structure of the labrum, the lack of abdominal ambulatory setae, and the

unusual structure of the apex of aedeagus. It is perhaps more plesiomorphic in the distinct microre-

ticulation and the absence of an oblique fold in the apex of the internal sac.

Adelotopus obsoletus, spec. nov.

Figs 44, 159, 346, 498, 619

Types. Holotype: d, Townsville, Qld, Oct. '02, F. P. Dodd (ANIC). - Paratypes: 17, 20 km.n. Biggenden, s. Old.

Australia, 22.1.1982, M. Baehr (CBM); 1%, Australia: Queensland, 32 km N Rockhampton 8.11.1964, J. Sedlacek

Collector BISHOP Museum (BMH); 15, 22%, Townsville Queensland, G. Bryant Coll. 1919-17, det. gyrinoides

(BMNH); 234, 222, Townsville, Old 23.10.02 F. P. Dodd, G. Bryant Coll. 1919, det. gyrinoides (ANIC, BMNH);

23d,N. of Mareeba Feb. ’58, N. O. Darlingtons (MCZ); 15, Australia: n. Old. 15 km NE of Mareeba 7.1-12.11.1985

Storey & Titmarsh, MDPI Intercept Trap Site No. 25 (DPIM); 19, Australia, Qld 93/7, Sand Flat Ck., 35 km

s. Palmer River, 23.5.1993, M. Baehr (CBM).

Diagnosis. Rather small to medium-sized, fairly wide, moderately convex, dull black species. Further

distinguished from related species by the rather wide pronotum with moderate lateral margins and

shortly rounded basal angles, incomplete basal border of elytra, very distinct microreticulation, fine

and rather dense puncturation, markedly dull surface, extremely narrow labrum that is deeply

covered by the clypeus, and short and wide aedeagus with strongly tapering, acute apex that is

conspicuously turned laterally.

Description

Measurements. Length: 4.6-5.6 mm. Ratios. Width/length of pronotum: 1.70-1.75; width base/

apex of pronotum: 1.49-1.56; width pronotum/ head: 1.55-1.60; length/width of elytra: 1.42-1.51; length

elytra/pronotum: 2.43-2.53.

Colour. Piceous-black to black, sometimes margins of pronotum and elytra faintly reddish

translucent. Lower surface piceous, posterior borders of abdominal sterna reddish. Mouth parts,

antennae, and legs piceous, tibiae and tarsi slightly lighter.

Head (Figs 159a-d). Rather short and wide, moderately depressed. Anterior border gently convex,

lateral angle rounded, laterally barely projecting, lateral borders but faintly oblique. Clypeal suture

semicircular, in middle interrupted. Labrum very narrow, deeply overlapped by the clypeus, rather

wide, apex deeply concave. Antennal groove laterally sharply bordered, latero-posteriorly with

sharply carinate area. Mental tooth triangular, short, apex acute. Wings of mentum wide, laterally

rounded, apex rectangular. Glossa wide, tongue-like, apically convex, ventrally with indistinct keel, at

border with c. 16 elongate setae and additional pilosity on upper and lower surface and along border.

Terminal palpomere of maxillary palpus rather widened, fairly securiform. Terminal palpomere of

labial palpus extremely wide, markedly securiform. Antenna moderately short, 8th-9th antennomeres

slightly <2 x as wide as long. Microreticulation fine, though distinct, puncturation rather fine, fairly

dense. Surface with a shallow sulcus medially of eyes, impilose, rather dull, slightly silky. Ventrola-

terally of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula asetose.

187

Figs 159a-l. Adelotopus obsoletus, spec. nov. Details of head and genitalia. For legends see fig. 100.

Pronotum (Fig. 346). Wide, moderately convex, base wide, markedly narrowed to apex. Apical

angles not much produced, at apex obtuse, somewhat oblique, surpassing posterior margin of eyes.

Apex moderately excised, convex in excision, laterally, somtimes also in middle faintly bordered. Sides

strongly and evenly curved throughout, widest near base. Margins fairly wide, slightly explanate,

faintly bordered. Basal angles rather shortly rounded off. Base laterally feebly concave, in middle

slightly convex, faintly bordered. Surface near base without transverse impression. Microreticulation

fine, distinct, puncturation fine, fairly dense, surface impilose, dull, slightly silky.

Elytra (Figs 44, 346, 498). Moderately elongate, slightly depressed on disk, regularly narrowed to

apex, faintly convex throughout. Apex rather wide, slightly oblique, truncature almost straight, apical

angles rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders.

Marsginal channel rather narrow, partly concealed. Basal border incomplete, reaching about halfways

to suture, ending abruptly. Lateral border asetose. Series of umbilical pores consisting of 6 closely set

pores behind shoulder and 1 additional pore behind middle. Setae rather short. Striae including

sutural stria absent. Microreticulation distinct, puncturation rather fine, fairly dense, surface impilose,

dull, slightly silky.

Lower surface. Prosternal process moderately short, narrow, straight, apex narrow, straight,

forming a sharp angle with the lower surface, rather setose. Metepisternum moderately short, c. 1.6 x

as long as wide, in posterior third not obliquely bent nor hollowed. Abdominal sterna without

ambulatory setae. Sternum VI without longer setae along apical border. Lower surface rather densely

punctate and pilose.

Legs. Rather elongate, Ist tarsomere of protarsus distinctly longer than wide, tibial groove of

profemur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove

sharp. Femur moderately wide. Metatibia elongate, almost 6 x as long as wide, 1st tarsomere of

metatarsus c. 2.5 x as long as wide. d protarsus not widened.

d genitalia (Figs 159e-k). Genital ring rather narrow, triangular, barely asymmetric, with slightly

asymmetric, narrow, barely excised base. Sternum VII rather narrow, apically convex, with deep

excision, basally convex, lateral parts short. Aedeagus rather short, depressed, in middle markedly

widened, slightly asymmetric. Lower surface almost straight. Apex narrow, acute, turned to left.

Orifice moderate, internal sac fairly complex, without a distinct oblique fold near apex. Both parameres

rather wide, with shortly rounded apex, left paramere considerably larger than right.

? genitalia (Fig. 1591). Stylomere narrow and elongate, apex obliquely convex, with 2-3 elongate

188

subapical setae. Lateral plate elongate, with 1-2 extremely short apical setae.

Variation. Some variation noted in size and in relative shape of pronotum and elytra.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. Specimens collected by me at light and under bark of river gum, another

specimen captured in “intercept trap”. So far collected in January, February, May, and October.

Distribution (Fig. 619). From eastern central Queensland to base of Cape York Peninsula.

Material examined (14). Only the type series.

Etymology. The name refers to the dull surface.

villosus-group

Diagnosis. Medium-sized, convex, black species. Whole surface with very coarse puncturation and

hirsute, very elongate setae; labrum bisetose; glossa c. 16-setose; lateral margin of pronotum narrow,

not explanate, basal angle widely rounded off; basal border line of elytra incomplete, ending halfways

to suture; scutellar pore absent; lateral margin of elytra narrow, without elongate setae behind

shoulders; series of lateral pores with 7 subhumeral pores and 1 postmedian pore; abdominal sterna

with 1 ambulatory seta on either side; sternum VI without longer setae at apical margin; tibiae,

especially metatibia depressed; all femora including profemur wide and depressed; aedeagus moder-

ately wide, slightly asymmetric, with rounded apex; internal sac of aedeagus complicate, with oblique

fold near apex.

Larva. Unknown.

Distribution. A single species in northeastern Queensland.

Systematic position. This groups is probably the adelphotaxon of the multipunctatus+obsoletus-

groups, particularly in view of the similar structure of the aedeagus. It is outstanding in the hairy

surface, but is more plesiomorphic than both mentioned groups in the larger number and large size

of the umbilical pores of the elytra.

Adelotopus villosus, spec. nov.

Figs 45, 160, 347, 499, 620

Types. Holotype: 4, Mareeba. NQ: 12/50 GB, M. 187., Adelotopus analis Macl. det. B. P. Moore '69, analis Macl.

1571, J. G. Brooks Bequest, 1976 (ANIC). - Paratypes: 19, Australia, Old 93/43, Sand Flat Ck., 1 km s. Hwy to

Cooktown, 5.6.1993, M. Baehr (CBM).

Diagnosis. Medium-sized, rather wide, black species. Distinguished from all other species except for

A. analis Macleay by the extremely coarse puncturation and the remarkably hirsute surface. Distin-

guished from A. analis by absence of pattern, absence of scutellar pore, basally much wider pronotum,

and apically markedly narrowed elytra.

Description

Measurements. Length: 5.65-6.0 mm. Ratios. Width/length of pronotum: 1.94-1.95; width base/

apex of pronotum: 1.57-1.58; width pronotum/ head: 1.67-1.71; length/ width of elytra: 1.43-1.44; length

elytra/pronotum: 2.77-2.81.

Colour. Black. Lower surface piceous-black, becoming reddish-piceous towards apex of abdomen.

Mouth parts, antennae, and legs piceous.

Head (Figs 160a-d). Short and wide, moderately depressed. Anterior border almost straight,

lateral angle shortly rounded, laterally distinctly projecting, lateral borders fairly oblique. Clypeal

suture indistinct, semicircular, in middle interrupted. Labrum rather large, moderately wide, apex

slightly concave. Antennal groove laterally sharply bordered, latero-posteriorly with sharply carinate

area. Mental tooth triangular, short, apex acute. Wings of mentum wide, laterally rounded, apex

rectangular. Glossa wide, tongue-like, apically convex, ventrally with indistinct keel, at border with

189

Figs 160a-l. Adelotopus villosus, spec. nov. Details of head and genitalia. For legends see fig. 100.

c. 16 elongate setae and additional pilosity on upper and lower surface and along border. Terminal

palpomere of maxillary palpus slightly widened, not securiform. Terminal palpomere of labial palpus

wide, securiform. Antenna short, 8th-9th antennomeres c. 2.25 x as wide as long. Microreticulation

absent, puncturation double, a very coarse and rather sparse one, and in the space between the coarse

punctures an extremely fine, moderately dense one. Surface with a shallow sulcus medially of eyes,

glossy, with elongate, erect hairs arising from the coarse punctures. Ventrolaterally of eyes with a row

of rather elongate setae. Suborbital field impunctate and asetose. Gula asetose.

Pronotum (Fig. 347). Rather wide, fairly convex, base wide narrow, strongly narrowed to apex.

Apical angles moderately produced, at apex obtuse, somewhat oblique, almost attaining posterior

third of eyes. Apex moderately excised, slightly convex in excision, faintly bordered. Sides strongly

and evenly curved throughout, widest in basal third. Margins rather narrow, not explanate, faintly

bordered. Basal angles very widely rounded off. Base faintly convex, distinctly bordered. Surface near

base without transverse impression. Microreticulation absent, puncturation double, a very coarse and

rather sparse one, and in the space between the coarse punctures an extremely fine, very sparse

puncturation, difficult to detect. Surface with some more or less distinct, fine wrinkles, highly glossy,

with elongate, erect hairs arising from the coarse punctures. Lateral margin provided with elongate

hairs.

Elytra (Figs 45, 347, 499). Moderately elongate, rather convex, though slightly depressed on disk,

evenly narrowe from base to apex. Apex rather narrow, slightly oblique, truncature slightly convex,

apical angles widely rounded off. Shoulders rounded, basal margin slightly oblique, behind shoulders

and along the whole lateral borders with elongate hairs. Marginal channel extremely narrow, barely

concealed. Basal border incomplete, reaching about halfways to suture. Lateral border finely crenulate.

Series of umbilical pores apparently consisting of 7 closely set pores behind shoulder and an additional

pore behind middle, though difficult to detect within the coarse puncturation. Setae elongate. Striae

including sutural stria absent. Microreticulation absent, puncturation very coarse and rather sparse,

without finer puncturation in the space between the coarse punctures, surface highly glossy.

Lower surface. Prosternal process rather short, moderately wide, straight, apex moderately wide,

convex, very setose. Metepisternum rather short, c. 1.5 x as long as wide, in posterior third obliquely

bent and rather hollowed. Abdominal sterna apparently with 1 elongate seta on either side. Sternum

VI without longer setae along apical border. Lower surface very coarsely, though rather sparsely

punctate, with elongate, erect hairs.

190

Legs. Rather short, Ist tarsomere of protarsus distinctly wider than long, tibial groove of profemur

deep, anterior plate widely overlapping the groove for apical half, posterior border of groove sharp.

Femur wide. Metatibia moderately elongate, slightly >4.5x as long as wide, Ist tarsomere of

metatarsus c. 1.5 x as long as wide. d protarsus not widened.

d genitalia (Figs 160e-k). Genital ring rather narrow, triangular, slightly convex, barely asymmet-

ric, with slightly asymmetric, narrow, barely excised base. Sternum VII rather wide, apically almost

straight, with fairly deep excision, basally bisinuate excised, lateral parts rather short. Aedeagus short,

depressed, in middle slightly widened, barely asymmetric. Lower surface almost straight. Apex wide,

widely reounded off. Orifice moderate, internal sac fairly complex, with a distinct oblique fold near

apex. Both parameres rather narrow, triangular, with rather narrow, shortly rounded apex, left

paramere considerably larger than right.

? genitalia (Fig. 160]). Stylomere wide, apex square, somewhat obliquely rounded, with 1-3

elongate subapical setae. Lateral plate moderately elongate, with 2-3 elongate apical setae.

Variation. Little variation noted.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. The paratype collected by me under bark of river gum at the border of a

dry sandy river. So far captured in December and June.

Distribution (Fig. 620). Base of Cape York Peninsula from Mareeba to about Palmer River, North

Queensland.

Material examined (2). Only the holotype and a paratype.

Etymology. The name refers to the conspicuously hairy surface.

similis-group

Diagnosis. Large, convex, black species with red apex. Labrum bisetose; glossa c. 12-setose; lateral

margin of pronotum narrow, not explanate, basal angle rounded off; basal border line of elytra

incomplete, ending halfways to suture; scutellar pore absent; lateral margin of elytra narrow, without

elongate setae behind shoulders; series of lateral pores with 10-14 pores along the whole margin;

abdominal sterna with 2-3 ambulatory setae on either side; sternum VI without longer setae at apical

margin; tibiae, especially metatibia depressed; all femora including profemur wide and depressed;

aedeagus narrow and elongate, symmetric, with narrowly rounded apex; internal sac of aedeagus

moderately complicate, with narrow oblique fold near apex.

Larva. 1st instar larva of the single species known.

Distribution. A single species in southeastern Australia, from eastern Victoria to New South Wales,

perhaps also in eastern South Australia.

Systematic position. This groups is externally similar to the multipunctatus-group, it belongs, howev-

er, to another lineage und is presumably the plesiomorphic adelphotaxon of the tasmani-linearis-

groups.

Adelotopus similis, spec. nov.

Figs 46, 161, 348, 500, 619

Types. Holotype: 4, 35.305, 150.18E, Kioloa SF, 15 km NE Batemans Bay, NSW Dec.86, M. G. Robinson, Flight

intercept trap (ANIC). - Paratypes: 13, 2??, same data (ANIC); 13, Adelaide (OUM); 1?, Australia, Victoria,

Melbourne, singled 2-10 11.1977, leg. G. Harsenyi (HNMB); 1%, Melbourne Coll. Castelnau, Haemorrhoidalıs

Erichs. (Melbourn.) (MCSN); 14, 37.345 145.53E, Cumberland Ck., 13km ESE of Marysville, 18 Jan.1978.

V. Lawrence & Weir, under bark rotten logs (ANIC); 15, Macedon, Vic. 26.12.32. C. Oke, Adelotopus haemorrhoi-

dalis Er. (NMV); 16, E. Eltham, V C. Oke, det. haemorrhoidalis (NMV); 15, P. Meyer 16.5.66., Woodhouse Cr.,

Nunniong Plt., under bark of Eucalyptus sp. (UASM); 14, P. M. 11.5.66, Woodhouse Ck., Nunniong Plt., under

bark of Eucalyptus sp. (UASM); 19, Lyons Villa Victoria J. E. Dixon, Adelotopus haemorrhoidalis Er. (NMV);

13, 2586 Victoria, haemorrhoidalis Erichs (AMNH); 17, 2586 Victoria, det. haemorrhoidalis (AMNH); 14, Victoria,

191

Ex Museo H. W. Bates 1892 (MNHN); 1%, Adelotopus gyrinoides Hope Victoria (NMV); 17, Thredbo R. HJC. 2.26

(ANIC); 14, The Creel, Mt. Kosc. 8000 ft. Dec 13°31 NSW, Australia Harvard Exp. Darlington, det. haemorrhoidalis

(MCZ); 5388, 32%, Australien, NSW 122 Mt. Kosciusko NP, Sawpit Creek, 1400 m, 11.-12.12.1990, M. Baehr

(CBM); 19, Kosciusko HJC 1-37 (ANIC); 12, NSW 20 km S. of Brindabella, 19 Jan.1985, J. F. Lawrence (ANIC);

18, Australien, ACT 126, Brindabella Range, 10 km nw. Picadilly Circus, 800 m, 10.12.1987, M. Baehr (CBM); 1%,

Australien, ACT 118, Picadilly Circus, 1200 m, 25 km w. Cotter Dam, 9.12.1990, M. Baehr (CBM); 15, 35.225,

148.48E, Picadilly Circus, 1240 m. ACT Mar’84, J. Lawrence, T. Weir, M. L. Johnson. coll., flight intercept

window /trough trap (ANIC); 254, 17, ACT, Brindabella Rge, Picadilly Circus, X-19-82, J. Doyen, Adelotopus

En BT Er. det. J. Liebherr 1987 (CUIC); 17, Wagga, N. 5. W., Oct.1936 C. Oke, Adelotopus haemorrhoidalis

., A. haemorrhoidalis (NMV); 18, N. S. W. Brown Mt. 28.1.1976, J. Sedlacek Collector (CSB); 12, Orange NSW

T G.s . 3.07 (ANIC); 1%, A. hydrobioides Westw. Forest Reefs, Lea, Adelotopus hydrobioides W, N. S. Wales,

Adelotopus hydrobioides Westw. Id. by T. G. Sloane (SAMA),; 15, Australia Blackb’s Coll., Adelotopus Australia

(SAMA); 18, 991, 30 Howitt Colln., det. gyrinoides (NMV); 1%, Collection E. Rousseau, I.R.Sc.N.B. 1.G. Coll. gen.

(IRSNB).

Diagnosis. Rather large, elogate, convex, black species with rather wide reddish apex of elytra.

Distinguished from the similar looking A. haemorrhoidalis Erichson, A. minor, spec. nov., and A. nitens,

spec. nov. by abbreviated basal border of elytra and larger number of umbilical pores.

Description

Measurements. Length: 6.9-7.8 mm. Ratios. Width/length of pronotum: 1.51-1.59; width base/

apex of pronotum: 1.44-1.50; width pronotum/ head: 1.55-1.62; length/ width of elytra: 1.67-1.71; length

elytra/pronotum: 2.70-2.79.

Colour (Figs 46, 348). Black, commonly margins of pronotum and elytra feebly dark reddish

translucent. Elytra with well defined red apex, the anterior border of the spot distinctly concave. Lower

surface of head and thorax black to dark piceous, of abdomen light reddish. Mouth parts, antennae,

and legs dark reddish, tibiae and tarsi slightly darker.

Head (Figs 161a-d). Short and wide, moderately depressed. Anterior border gently convex, lateral

angle rounded, laterally barely projecting, lateral borders faintly oblique. Clypeal suture semicircular,

in middle not or slightly interrupted. Labrum large, apex barely concave. Antennal groove laterally

sharply bordered, latero-posteriorly without carinate area. Mental tooth triangular, short, apex acute.

Wings of mentum wide, laterally rounded, apex rectangular. Glossa wide, tongue-like, apically

convex, ventrally with indistinct keel, at border with c. 12 elongate setae and additional pilosity on

upper and lower surface and along border. Terminal palpomere of maxillary palpus elongate, barely

widened, not securiform. Terminal palpomere of labial palpus rather wide, securiform. Antenna short,

Sth-9th antennomeres c. 2 x as wide as long. Microreticulation rather fine, distinct, puncturation very

fine, rather difficult to see even under high magnification, rather sparse. Surface with a shallow sulcus

medially of eyes, sometimes with some faint, irregular strioles, impilose, moderately glossy. Ventro-

laterally of eyes with a row of short setae. Suborbital field impunctate and asetose. Gula apparently

asetose.

Pronotum (Fig. 348). Moderately wide, convex, base moderately wide, apex narrower. Apical

angles moderately produced, at apex obtuse, somewhat oblique, almost attaining posterior third of

eyes. Apex moderately deeply excised, slightly convex in excision, unbordered. Sides evenly curved

throughout, widest in basal fourth or near base. Margins moderate, not explanate, faintly bordered.

Basal angles evenly rounded off. Base almost straight, bordered. Surface near base without transverse

impression. Microreticulation very fine, distinct, only slightly superficial, puncturation extremely fine,

though slightly more distinct than on head, rather sparse, surface with more or less distinct, faint,

irregular strioles, impilose, fairly glossy.

Elytra (Figs 46, 348, 500). Elongate, convex, margins rather parallel in basal %, then gently

narrowed. Apex rather wide, transverse, truncature faintly convex, apical angles rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

rather narrow, partly concealed. Basal border incomplete, ending about halfways to suture. Lateral

border asetose. Series of umbilical pores consisting of 10-14 pores in a rather spaced and somewhat

irregular row running down almost the whole length of elytra. Setae moderately elongate. Striae very

faintly indicated as rows of extremely fine, irregular, longitudinal strioles. Microreticulation fine, very

superficial, sometimes difficlut to detect, though still visible, puncturation fine and rather sparse,

becoming more distinct and denser towards apex, surface impilose, rather glossy.

192

Figs 161a-l. Adelotopus similis, spec. nov. Details of head and genitalia. For legends see fig. 100.

Lower surface. Prosternal process rather short, fairly wide, straight, gently convex, apex wide and

rather short, margin slightly convex, rather setose. Metepisternum elongate, >2 x as long as wide or

even longer, neither bent nor hollowed. Abdominal sterna with 2-3 elongate setae on either side.

Sternum VI without longer setae along apical border. Lower surface rather punctate and pilose.

Legs. Medium-sized, 1st tarsomere of protarsus slightly wider than long, tibial groove of profe-

mur deep, anterior plate widely overlapping the groove for apical %, posterior border of groove sharp.

Femur wide. Metatibia medium-sized, <4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.6 x as

long as wide. d protarsus not widened.

d genitalia (Figs 161e-k). Genital ring medium-sized, basally rather parallel, symmetric, arms

evenly convex, with rather narrow, little excised base. Sternum VII moderately wide, apically convex,

with deep excision, basally regularly excised, lateral parts short. Aedeagus very narrow and elongate,

depressed, in middle feebly widened, almost symmetric, apical part very elongate, spatulate. Lower

surface gently concave. Apex rather narrow, shortly rounded off. Orifice elongate, internal sac fairly

complex, with a small, oblique fold near apex. Right paramere rather narrow, almost parallel, apex

attenuate. Left paramere considerably larger than right, elongate, apex obtusely rounded.

? genitalia (Fig. 1611). Stylomere rather narrow and elongate, only slightly triangular, with

narrow, shortly rounded apex, with 2 subapical setae. Lateral plate elongate, with 2-3 long apical setae.

Variation. Some variation of size, relative shape of pronotum and elytra, and degree of microre-

ticulation and puncturation noted.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Little known. Some specimens collected under bark of “Eucalyptus sp.”, others in “flight

intercept trap”, specimens captured by me under bark of different eucalypts, even in dense forest,

some in rather high altitude up to about 1400 m. So far collected in October, from December to March,

and in May, most specimens, however, in December and January. Occurs sympatrically with

A. haemorrhoidalis Erichson and A. minor, spec. nov., and even syntopically with A. haemorrhoidalis (on

Picadilly Circus actually on the same tree!).

Distribution (Fig. 619). Eastern Victoria, Australian Capital Territory, New South Wales, ? South

Australia. The latter unique record is doubtful, the more, as it is an old specimen from the Hope

Collection that bears only the label “Adelaide”.

Material examined (42). Only the type series.

Etymology. The name refers to the strong external similarity with the related species A. haemorrhoidalis

Erichson.

tasmani-group

Diagnosis. Medium-sized, elongate, highly convex, black species with red apex of elytra. Labrum

bisetose; glossa c. 12-setose; lateral margin of pronotum narrow, not explanate, basal angle shortly

rounded off; basal border line of elytra incomplete, reaching to median third of base; scutellar pore

absent; lateral margin of elytra very narrow, without elongate setae behind shoulders; series of lateral

pores with 6 subhumeral pores only; abdominal sterna with many ambulatory setae on either side;

sternum VI without longer setae at apical margin; tibiae, especially metatibia depressed; all femora

including profemur wide and depressed; aedeagus so far unknown.

Larva. Unknown.

Distribution. A single species from Tasmania.

Systematic position. The relations of this groups are rather uncertain. Perhaps it belongs to the

lineage that combines the similis-linearis-groups and is probably the adelphotaxon of the nigricauda-

linearis-lineage. It is more apomorphic than the similis-group in the lesser number of umbilical pores

of the elytra and in the narrow, cylindrical shape.

Adelotopus tasmani Blackburn, 1901

Figs 47, 162, 349, 501, 619

Adelotopus tasmani Blackburn, 1901, p. 18; Seidlitz 1903, p. 132 (tasmaniae); Sloane 1920, p. 177; Notman 1925, p. 6,

29; Csiki 1933, p. 1636 (tasmaniae); Moore et al. 1987, p. 53.

Types. Lectotype (by present designation): ?, Type, 1491 T. Tasm. Lakes, Blackburn Coll. 1910-236, Adelotopus

Tasmani, Blackb. (BMNH).

Type locality: “Tasm. Lakes”, central Tasmania.

Diagnosis. Medium-sized, narrow, elongate, convex, black species with fairly wide reddish apex of

elytra. Distinguished from similar looking species by the rather elongate basal border of elytra, rather

large number of umbilical pores, and large number of ambulatory setae of the abdomen.

Description

Measurements. Length: 5.8 mm. Ratios. Width/length of pronotum: 1.30; width base/apex of

pronotum: 1.25; width pronotum/ head: 1.36; length/ width of elytra: c. 1.83; length elytra/pronotum:

2.56:

Colour (Figs 47, 349). Blackish, all margins of pronotum and lateral margin of elytra faintly reddish

translucent, apical fourth of elytra reddish, border of reddish spot prolonged along margins. Lower

surface of head and thorax blackish, abdomen reddish. Mouth parts, antennae, and legs reddish, tibiae

and tarsi barely darker.

Head (Figs 162a-d). Moderately short, rather wide, fairly convex. Anterior border gently convex,

lateral angle rounded, laterally projecting, lateral borders concave behind eyes. Clypeal suture distinct,

semicircular. Labrum rather large and wide, apex slightly concave. Antennal groove laterally sharply

bordered, latero-posteriorly with convex area. Mental tooth triangular, short, apex acute. Wings of

mentum wide, laterally rounded, apex angulate. Glossa wide, tongue-like, apically convex, ventrally

with indistinct keel, at border with c. 12 elongate setae and additional pilosity on upper and lower

surface and along border. Terminal palpomere of maxillary palpus narrow, not securiform. Terminal

palpomere of labial palpus moderately wide, slightly securiform. Antenna very short, Sth-9th

antennomeres almost 3x as wide as long. Microreticulation dense and distinct, puncturation not

visible. Surface with shallow sulcus medially of eyes, impilose, dull, somewhat silky. Ventrolaterally

of eyes with a row of very short and inconspicuous setae. Suborbital field impunctate. Gula impilose.

194

Figs 162a-d, I. Adelotopus tasmani Blackburn. Details of head and ? genitalia. For legends see fig. 100.

Pronotum (Fig. 349). Rather narrow, convex, not much wider than long, base slightly wider than

apex. Apical angles barely produced, at apex rounded, rather oblique, barely attaining posterior border

of eyes. Apex feebly excised, slightly convex in excision, faintly and irregularly bordered. Sides slightly

oblique, almost straight, widest near base. Margins narrow, faintly chanelled, finely bordered. Basal

angles moderately widely rounded off. Base faintly convex, distinctly bordered. Surface near base

without transverse impression. Microreticulation dense and distinct, puncturation not visible, surface

with some very fine, irregular wrinkles, impilose, dull, slioghtly silky.

Elytra (Figs 47, 349, 501). Narrow and elongate, cylindrical, almost parallel, though on disk slightly

depressed. Apex wide, transverse, faintly convex, apical angles widely rounded off. Shoulders

rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel narrow

throughout, mostly concealed. Basal border incomplete, reaching to median third of base. Lateral

border asetose. Series of umbilical pores consisting of 6 closely set pores behind shoulder. Setae fairly

short. Striae including sutural stria absent. Microreticulation dense and distinct, puncturation not

visible, surface impilose, rather dull.

Lower surface. Prosternal process rather elongate, moderately wide, slightly convex, apex wide,

gently convex, passing over in an almost right angle from ventral surface, setose. Metepisternum

elongate, c. 2.2 x as long as wide, in posterior third not hollowed. Abdominal sterna with a row of

several elongate setae along base of each. Sternum VI without longer setae along apical border. Lower

surface sparsely punctate and pilose. Prosternum with some deep, rugose, transverse sulci.

Legs. Rather short, Ist tarsomere of protarsus much wider than long, tibial groove of profemur

deep, anterior plate deeply overlapping the groove for apical half, posterior border of groove sharp.

Femur wide. Metatibia fairly short, c. 4.5 x as long as wide, 1st tarsomere of metatarsus c. 1.3 x aslong

as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Figs 1621). Stylomere wide, though apex acute, triangular, obliquely rounded, lateral

border concave, with 1 elongate seta at apex and 1 sorter subapical seta. Lateral plate elongate, with

2 elongate apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Unknown.

Distribution (Fig. 619). Central Tasmania.

Material examined (1). Only the lectotype.

195

nigricauda-group

Diagnosis. Medium-sized, elongate, though dorsally depressed, black species with basal % of elytra

red. Labrum bisetose; glossa c. 12-setose; lateral margin of pronotum fairly wide, somewhat channeled,

but not explanate, basal angle shortly rounded off; basal border line of elytra incomplete, reaching

halfways to suture; scutellar pore absent; lateral margin of elytra narrow, without elongate setae

behind shoulders; series of lateral pores with 6 subhumeral pores only; abdominal sterna with many

1 ambulatory seta on either side; sternum VI without longer setae at apical margin; tibiae, especially

metatibia depressed; all femora including profemur wide and depressed; aedeagus rather short and

wide, slightly asymmetric, with acute apex; internal sac of aedeagus rather complicate, with distinctly

denticulate area, but without oblique fold near apex.

Larva. Unknown.

Distribution. A single species from northern Cape York Peninsula, northeastern Queensland.

Systematic position. The relations of this groups are rather uncertain. Probably it belongs to the

similis-linearis-lineage and is perhaps the adelphotaxon of the seriepunctatus-linearis-groups. It is

apomorphic in the narrow, dorsally depressed shape, the unusual elytral and abdominal pattern, the

low number of umbilical pores of the elytra, the absence of the microreticuation on the elytra, the acute

apex of the aedeagus, and the presence of distinct denticulation at the apex of the internal sac. It is

perhaps plesiomorphic in the absence of a distinct oblique fold at the apex of the aedeagus.

Adelotopus nigricauda, spec. nov.

Figs 48, 163, 350, 620

Types. Holotype: 4, 12.435 142.42E QLD 7 km S of Batavia Downs 19 Jun-22 Jul 1992 Flight Intercept Trap

P. Zborowski & E. 5. Nielsen (QMB T26062).

Diagnosis. Medium sized, elongate, fairly parallel, rather convex, black species with basal % of elytra,

meso- and metathorax, and base of abdomen red. Further distinguished from similarly coloured

species by absence of any black colour at the base of elytra except for the black scutellum, black rather

than red apex of abdomen, narrow shape, narrow, almost parallel pronotum with base little wider than

apex, and short and wide, at apex rather shortly rounded aedeagus.

Description

Measurements. Length: c. 4.85 mm. Ratios. Width/length of pronotum: 1.42; width base/apex of

pronotum: 1.22; width pronotum/head: 1.35; length/width of elytra: 1.79; length elytra/pronotum:

2.58.

Colour (Figs 48, 350). Black, lateral margin of pronotum narrowly reddish translucent. Basal % of

elytra light reddish, apex contrastingly black. Lower surface of head and prothorax black, meso- and

metathorax and base of abdomen red, 5 apical sternites of abdomen contrastingly black. Mouth parts,

antenna, and legs red.

Head (Figs 163a-d). Short and moderately wide, moderately depressed. Anterior border gently

convex, lateral angle obtusely rounded, laterally barely projecting, lateral borders oblique. Clypeal

suture barely visible. Labrum rather wide, apex feebly concave. Antennal groove laterally sharply

bordered, latero-posteriorly with weak carina. Mental tooth triangular, short, apex acute. Wings of

mentum wide, laterally rounded, apex obtusely rounded. Glossa wide, tongue-like, apically widely

rounded, ventrally with indistinct keel, at border with c. 10-12 elongate setae and additional pilosity

on upper and lower surface and along border. Terminal palpomere of maxillary palpus slightly

widened, slightly securiform. Terminal palpomere of labial palpus very wide, >2 x as wide as long,

markedly securiform. Antenna short, $Sth-9th antennomeres c. 2 x as wide as long. Microreticulation

very fine, distinct, puncturation extremely fine, moderately dense, surface with a shallow sulcus

medially of eyes, impilose, moderately glossy, somewhat silky. Ventrolaterally of eyes with a row of

short setae. Suborbital field impunctate and asetose. Both palpi rather sparsely setose, gula asetose.

Pronotum (Fig. 350). Moderately wide, fairly convex, almost parellel, base not much wider than

apex. Apical angles moderately produced, moderately acute, obtuse at apex, slightly oblique, surpass-

196

Figs 163a-k. Adelotopus nigricauda, spec. nov. Details of head and d genitalia. For legends see fig. 100.

ing posterior border of eyes. Apex moderately excised, slightly convex in excision, laterally distinctly,

in middle faintly bordered. Sides faintly curved throughout, widest about in middle. Margins rather

explanate, anteriorly faintly bordered. Basal angles rounded off. Base almost straight, faintly bordered.

Surface near base with very faint transverse impression. Microreticulation extremely superficial,

almost absent, puncturation very fine, rather sparse, surface impilose, glossy.

Elytra (Figs 38, 350). Elongate, convex, almost parallel. Apex rather wide, slightly oblique,

truncature almost straight, apical angles rounded off. Shoulders obtusely rounded, basal margin

slightly oblique, without setae behind shoulders. Marginal channel rather narrow, inner part con-

cealed. Basal border incomplete, absent in median half, ending gradually. Scutellar pore absent. Lateral

border asetose. Series of umbilical pores consisting of 6 pores behind shoulder. Setae short. Striae

including sutural stria absent. Microreticulation absent, puncturation extremely fine, rather sparse,

surface structure difficult to detect because of weakness of elytra. Surface impilose, glossy.

Lower surface. Prosternal process moderately elongate, fairly wide, straight, apex moderately

elongate, narrow , straight, shortly setose. Metepisternum moderately elongate, c. 1.85 x as long as

wide, almost not bent and hollowed in posterior third. Abdominal sterna with 1 elongate setae on

either side. Sternum VI without elongate setae at apical border. Lower surface moderately punctate

and pilose.

Legs. Medium-sized, 1st tarsomere of protarsus about as long as wide, tibial groove of profemur

very deep, anterior plate widely overlapping the groove for at least apical half, posterior border of

groove sharp. Femur rather wide. Metatibia rather elongate, c. 5 x as long as wide, Ist tarsomere of

metatarsus c. 2x as long as wide. d protarsus not widened.

d genitalia (Figs 163e-k). Genital ring rather narrow, markedly triangular, feebly asymmetric, arms

almost straight, with rather narrow, asymmetric base, with elongate apex. Sternum VII rather wide,

apically almost straight, with fairly deep excision, basally gently concave, lateral parts moderately

elongate. Aedeagus short, wide, rather depressed, in middle strongly widened, slightly asymmetric.

Lower surface slightly convex. Apex rather narrow, obtusely rounded. Orifice rather short, internal sac

fairly complex, with a distinctly denticulate area, but no oblique fold at apex. Both parameres large and

elongate, right almost as long as left, apex obtusely rounded.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Largely unknown. Holotype collected in “flight intercept trap”. So far captured in the period

of June-July.

Distribution (Fig. 620). Northern part of Cape York Peninsula, north Queensland.

197

Material examined (1). Only the holotype.

Etymology. The name refers to the contrastingly black apex of elytra and abdomen.

seriepunctatus-group

Diagnosis. Medium-sized, rather narrow and elongate, almost cylindrical, glossy black species with

reddish apex of elytra. Labrum bisetose; glossa c. 12-setose; lateral margin of pronotum very narrow,

basal angle almost rectangular; basal border line of elytra abbreviated, attaining less than half of base;

scutellar pore absent; lateral margin of elytra without elongate setae; series of umbilical pores with 6-8

subhumeral pores only; abdominal sterna with 1 ambulatory seta each side; sternum VI without a

fringe of longer setae at apical margin; tibiae, especially metatibia rather depressed; all femora wide

and depressed; aedeagus rather short and wide, with rounded or acute apex; internal sac of aedeagus

complicate, without oblique fold at apex.

Larvae. 1st instar larvae known of one species only.

Distribution. 5 species and additional 2 subspecies from South Australia through Victoria, NSW

including the southwestern part to northeastern Queensland, and southwestern Australia.

Systematic position. This group is perhaps rather plesiomorphic in the low number of glossal setae

and the large number of marginal elytral pores. It is highly apomorphic, however, in the narrow,

cylindrical shape, presence of pattern, absence of microreticulation and very glossy surface, abbrevi-

ated basal border of elytra, low number of abdominal ambulatory setae, and the complicate structure

of the internal sac of the aedeagus. Probably it is the in severeal respects highly apomorphic

adelphotaxon of the rubiginosus-laevis-groups.

Adelotopus seriepunctatus Notman, 1925

This species occurs in two subspecies, a southern one in Victoria, and a northern one in Queensland,

though there are so far no records available from the area between, namely New South Wales.

The examined material of this species includes a syntype of A. analis Macleay which, however, is

a completely different species. A specimen from the Chaudboir Collection is labelled lioderma Chaudoir

which is perhaps a manuscript name that, at least from my knowledge, has been never published.

Diagnosis. Medium-sized to fairly large, elongate, convex, black species with reddish apex. Distin-

guished from related species by the glossy surface, rather conical, only fairly punctate pronotum,

narrow, though coarsely bordered lateral margin of pronotum, uniseriate puncturation on the elytral

intervals with each puncture bearing an elongate seta, and the wide, medially very convex, laterally

concave stylomere.

Adelotopus seriepunctatus seriepunctatus Notman, 1925

Figs 164, 351, 502, 620

Adelotopus seriepunctatus Notman, 1925, p. 8, 9, 29; Csiki 1933, p. 1636; Moore et al. 1987, p. 52.

Types. Holotype: 2, 2590 Victoria, Type, Adelotopus seriepunctatus Ntm. det. Ntm (AMNH).

Type locality: “Victoria”.

Diagnosis. Large subspecies, distinguished from A. seriepunctatus striatus, subspec. nov. by larger size,

rather distinctly conical pronotum with less convex lateral margins, less dense puncturation on apical

half of pronotum, and absence of punctures along the elytral striae.

198

Figs 164a-1. Adelotopus seriepunctatus seriepunctatus Notman. Details of head and genitalia. For legends see

fig. 100.

Description

Measurements. Length: 6.0-6.7 mm. Ratios. Width/length of pronotum: 1.33-1.44; width base/

apex of pronotum: 1.43-1.51; width pronotum/head: 1.51-1.59; length/ width of elytra: 1.58-1.64; length

elytra/pronotum: 2.29-2.34.

Colour (Fig. 351). Glossy black, apical fourth or fifth of elytra reddish with fairly ill defined border.

Lower surface of head and thorax blackish, abdomen reddish-piceous or reddish, slightly lighter

towards apex. Mouth parts, antennae, and legs dark reddish, tibiae and tarsi not darker.

Head (Figs 164a-d). Moderately short, rather wide, rather depressed. Anterior border gently

convex, lateral angle rounded, laterally projecting, lateral borders concave behind eyes. Clypeal suture

indistinct, almost invisible. Labrum rather wide and short, moderately overlapped by the clypeus,

apex feebly concave. Antennal groove laterally sharply bordered, latero-posteriorly with slightly

angulate area. Mental tooth triangular, short, apex acute. Wings of mentum wide, laterally rounded,

apex angulate. Glossa wide, tongue-like, apically convex, ventrally with indistinct keel, at border with

c. 12 elongate setae and additional pilosity on upper and lower surface and along border. Terminal

palpomere of maxillary palpus elongate, not securiform. Terminal palpomere of labial palpus

moderately wide, barely securiform. Antenna narrow and rather elongate, parallel, Sth-9th antenno-

meres c. 1.5 x as wide as long. Microreticulation absent, puncturation very fine, moderately dense.

Surface with weak sulcus medially of eyes, impilose, very glossy. Ventrolaterally of eyes with a row

of short setae. Suborbital field impunctate. Gula impilose.

Pronotum (Fig. 351). Large, though rather narrow, highly convex, not much wider than long,

rather conical, base distinctly wider than apex. Apical angles fairly produced, at apex rather acute,

fairly oblique, attaining posterior third of eyes. Apex fairly excised, slightly convex in excision,

distinctly bordered. Sides slightly convex, markedly oblique. Margins narrow, barely chanelled,

coarsely bordered. Basal angles rectangular, very shortly rounded off. Base faintly convex, coarsely

bordered. Surface near base with extremely shallow transverse impression. Microreticulation absent,

puncturation in basal half and in middle of apical part very sparse, rather fine, in lateral part of apical

half coarse and fairly dense, though to a rather different degree, surface with scattered, rather elongate

hairs, highly glossy.

Elytra (Figs 351, 502). Rather narrow and elongate, convex, parallel, sometimes even faintly

widened in apical third. Apex very wide, transverse, truncature convex, in middle distinctly drawn in,

199

apical angles widely rounded off. Shoulders rounded, basal margin slightly oblique, without setae

behind shoulders. Marginal channel anteriorly narrow, becoming extremely narrow apically, partly

concealed. Basal border incomplete, attaining outer third of base. Lateral border asetose. Series of

umbilical pores consisting of 7-8 rather spaced pores behind shoulder, but pores difficult to tell from

puncturation of disk, especially posteriorly. Setae fairly elongate. Scutellum very wide. Striae includ-

ing sutural stria absent, not marked by rows of fine punctures. Microreticulation absent, each interval

with a spaced row of slightly rasp-like or umbilicous punctures, each of which bears a fairly elongate,

erect hair, surface markedly glossy.

Lower surface. Prosternal process rather elongate, moderately wide, convex, apex wide, convex,

passing over inan almost right angle from ventral surface, rather setose. Metepisternum very elongate,

c. 2.2-2.3 x as long as wide, in posterior third not hollowed, but becoming very narrow towards apex.

Abdominal sterna with 1 elongate seta each side. Lower surface sparsely punctate and rather

elongately setose.

Legs. Moderately elongate, 1st tarsomere of protarsus slightly wider than long, tibial groove of

profemur deep, anterior plate deeply overlapping the groove for apical half, posterior border of groove

sharp. Femur wide. Metatibia fairly elongate, c.5 x as long as wide, 1st tarsomere of metatarsus c. 1.8 x

as long as wide.

S genitalia (Figs 164e-k). Genital ring moderately wide, rather triangular, barely asymmetric, with

slightly asymmetric, rather small, fairly excised base. Sternum VII rather wide, apically almost straight,

with moderately deep excision, base excised, basal angles obtuse, lateral parts fairly short. Aedeagus

rather short, moderately depressed, in middle markedly widened, evenly narrowed to apex, faintly

asymmetric. Basal part fairly short, rather bent. Lower surface almost straight. Apex moderately wide,

evenly rounded. Orifice rather elongate, internal sac moderately complex, apparently without a

distinct oblique fold near apex. Both parameres rather elongate, slightly triangular, though convex,

with moderately rounded apex, left paramere considerably larger than right.

? genitalia (Figs 1641). Stylomere wide, median border markedly convex, lateral border concave,

with 3-5 elongate setae at apex. Lateral plate rather elongate, with 2-3 elongate apical setae.

Variation. Only some variation noted in shape of pronotum which may be more or less conical,

and in density of the coarse punctures on the lateral parts of the apical half of the pronotum.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Dated specimens captured in January and August.

Distribution (Fig. 620). Victoria, southern New South Wales.

Material examined (7). Vic 1%, Melbourne, inguinatus Newm. (BMNH); 1%, Killara, C. Oke, Adelotopus

seriepunctatus Notm. (NMV); 1%, Launching Place, 1.1.08, C. Oke, Adelotopus seriepunctatus Notm. (NMV);

12, Greendale, 12.VII1.(9)56, A. N. (CBM); 1?, 2590, Type, Adelotopus seriepunctatus Ntm. det. Ntm, holotype!

(AMNH). - NSW: 18, 1?, Jenolan, 21.VI11.33, ©. H. Swezey Collector (BMH).

Adelotopus seriepunctatus striatus, subspec. nov.

Figs 49, 352, 503, 620

Types. Holotype: d, Townsville, Old, 23.X.02, F. P. Dodd, 21 (ANIC). - Paratypes: 1%, Gayndah, Syntype,

Adelotopus analis Macl. Gayndah (ANIC-MMS); 12, Maryborough Queensland E. W. Fischer (SAMA);

12, Rockhampton, N. Australia (Darnelly) Higgins 1864 (OUM); 15, Qld 29, 215 km n. Dingo, Fitzroy Dev. Road,

12.X1.1990, M. Baehr (CBM); 15, 1%, N. Holl. Q’land, Janson Acg. 1884 (MNHN); 1%, Queensl (OUM);

12, Australie sept. Schmeltz, Ex Musaeo Chaudoir, det. lioderma Chaud. (MNHN); 14, Nov. Holl. (NHRS).

Diagnosis. Smaller subspecies, distinguished from A. s. seriepunctatus Notman by lesser size, not

distinctly conical pronotum with convex lateral margins, denser puncturation on apical half of

pronotum, and presence of punctures along the elytral striae.

Description

Measurements. Length: 4.95-6.1 mm. Ratios. Width/length of pronotum: 1.33-1.43; width base/

apex of pronotum: 1.42-1.51; width pronotum/ head: 1.50-1.58; length/ width of elytra: 1.59-1.63; length

elytra/pronotum: 2.22-2.32.

200

Figs 165a-d, 1. Adelotopus convexus, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Colour (Figs 49, 352). Similar to nominate subspecies.

Head. Similar to nominate subspecies.

Pronotum (Fig. 352). Rather similar to nominate subspecies, though less conical and with more

distinctly convex lateral margins, and basal angles more widely rounded off. Puncturation even in in

basal half and in middle of apical half denser and coarser than in nominate subspecies, usually also

middle of apical half distinctly and rather coarsely punctate. In lateral part of apical half puncturation

also denser.

Elytra (Figs 49, 352, 503). Rather similar to nominate subspecies, though striae indicated by rows

of fine punctures.

Lower surface. Similar to nominate subspecies.

Legs. Similar to nominate subspecies.

d genitalia. Rather similar to nominate subspecies. In the few examined males aedeagus slightly

longer and narrower.

? genitalia. Rather similar to nominate subspecies. Stylomere with 2-4 elongate setae at apex.

Variation. Some variation noted in shape of pronotum, density of pronotal puncturation, and

distinctness of puncturation of elytral striae.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. A specimen collected by me under bark of a bloodwood-like eucalypt,

together with several other pseudomorphine species. Dated specimens captured in October and

November.

Distribution (Fig. 620). Central eastern Queensland north to Townsville.

Material examined (10). Only the type series.

Etymology. The name refers to the distinct traces of elytral striation.

Adelotopus convexus, spec. noV.

Figs 165, 353, 504, 620

Types. Holotype: ?, Lawes, 15.12.54, PQ ASA9 (QMB T26067).

Diagnosis. Medium-sized, elongate, very convex, black species with reddish apex. Distinguished

from related species by the glossy surface, laterally rather convex, only fairly punctate pronotum, very

narrow lateral margin of pronotum, uniseriate puncturation on the elytral intervals with each puncture

bearing an elongate seta, absence of punctures along the elytral striae, the moderately narrow, parallel,

apically almost transverse stylomere, and the elongate lateral plate. Also distinguished from the most

closely related species A. seriepunctatus Notman by narrower pronotum and longer elytra.

201

Description

Measurements. Length: c. 5.65 mm. Ratios. Width/length of pronotum: 1.37; width base/apex of

pronotum: 1.41; width pronotum/head: 1.46; length/width of elytra: 1.69; length elytra/pronotum:

2.40.

Colour (Fig. 353). Glossy black, apical third of elytra reddish with fairly ill defined and very serrate

border. Lower surface of head and thorax blackish, abdomen reddish, slightly lighter towards apex.

Mouth parts, antennae, and legs dark reddish, tibiae and tarsi not darker.

Head (Figs 165a-d). Moderately short, rather wide, rather depressed. Anterior border gently

convex, lateral angle rounded, laterally projecting, lateral borders concave behind eyes. Clypeal suture

indistinct, almost invisible. Labrum rather wide and short, moderately overlapped by the clypeus,

apex feebly concave. Antennal groove laterally sharply bordered, latero-posteriorly with slightly

angulate area. Mental tooth triangular, short, apex acute. Wings of mentum wide, laterally rounded,

apex angulate. Glossa wide, tongue-like, apically convex, ventrally with indistinct keel, at border with

c. 12 elongate setae and additional pilosity on upper and lower surface and along border. Terminal

palpomere of maxillary palpus elongate, not securiform. Terminal palpomere of labial palpus

moderately wide, barely securiform. Antenna narrow and rather elongate, parallel, 8th-9th antenno-

meres c. 1.5 x as wide as long. Microreticulation absent, puncturation extremely fine, sparse. Surface

with weak sulcus medially of eyes, impilose, very glossy. Ventrolaterally of eyes with a row of short

setae. Suborbital field impunctate. Gula impilose.

Pronotum (Fig. 353). Large, though narrow, highly convex, barely wider than long, base not much

wider than apex. Apical angles fairly produced, at apex rather acute, fairly oblique, attaining posterior

third of eyes. Apex fairly excised, slightly convex in excision, unbordered. Sides fairly convex, slightly

oblique. Margins very narrow, not chanelled, rather faintly bordered. Basal angles rectangular, very

shortly rounded off. Base faintly concave, coarsely bordered. Surface near base without transverse

impression. Microreticulation absent, puncturation in basal half and in middle of apical half fairly

sparse, moderately fine, in lateral part of apical half coarse and rather dense, surface with scattered,

rather elongate hairs, highly glossy.

Elytra (Figs 353, 504). Narrow and elongate, highly convex, basally parallel, in apical third evenly

narrowed. Apex rather narrow, transverse, truncature convex, in middle distinctly drawn in, apical

angles widely rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind

shoulders. Marginal channel very narrow except for basal sixth, mostly concealed. Basal border

incomplete, attaining outer third of base. Lateral border asetose. Series of umbilical pores consisting

of 6-7 rather spaced pores behind shoulder, but pores difficult to tell from puncturation of disk,

especially posteriorly. Setae fairly elongate. Scutellum very wide. Striae including sutural stria absent,

not marked by rows of fine punctures. Microreticulation absent, each interval with a spaced row of

slightly rasp-like or umbilicous punctures, each of which bears a fairly elongate, erect hair, surface

markedly glossy.

Lower surface. Prosternal process rather elongate, moderately wide, convex, apex wide, convex,

passing over in an almost right angle from ventral surface, rather setose. Metepisternum very elongate,

c. 2.4 x as long as wide, in posterior third not hollowed, but becoming very narrow towards apex.

Abdominal sterna with 1 elongate seta each side. Lower surface sparsely punctate and rather

elongately setose.

Legs. Moderately elongate, 1st tarsomere of protarsus slightly wider than long, tibial groove of

profemur deep, anterior plate deeply overlapping the groove for apical half, posterior border of groove

sharp. Femur wide. Metatibia fairly elongate, c.5 x as long as wide, 1st tarsomere of metatarsus c. 1.8 x

as long as wide. d protarsus unknown.

ö genitalia. Unknown.

? genitalia (Fig. 1651). Stylomere rather wide, fairly prallel, median border straight, lateral border

slightly concave, apex slightly widened, almost transverse, with 3 elongate setae. Lateral plate very

elongate, with 2 elongate apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Holotype collected in December.

Distribution (Fig. 620). Southeastern Queensland. Known only from type locality.

202

Figs 166a-d, 1. Adelotopus calvus, spec. nov. Details of head and ? genitalia. For legends see fig. 100.

Material examined (1). Only the holotype.

Etymology. The name refers to the convex shape of prothorax and elytra.

Adelotopus calvus, spec. nov.

Figs 166, 354, 505, 621

Types. Holotype: ?, 1 ml East Bakerville Q 13-XI-1974 J. D. Brown, Adelotopus sp. det. B. P. Moore '80 (QMB

T26057).

Diagnosis. Medium-sized, rather elongate, convex, black species with reddish apex. Distinguished

from related species by the glossy surface, laterally rather convex, only fairly punctate pronotum, fairly

wide lateral margin of pronotum, impunctate and impilose elytral intervals, punctate elytral striae,

and the narrow, parallel, stylomere.

Description

Measurements. Length: 5.3 mm. Ratios. Width/length of pronotum: 1.40; width base/apex of

pronotum: 1.39; width pronotum/head: 1.51; length/width of elytra: c. 1.55; length elytra/pronotum:

2.21.

Colour (Fig. 354). Glossy black, lateral margins of pronotum and elytra reddish translucent, apical

third of elytra reddish with fairly ill defined border. Lower surface of head and thorax reddish-

piceous, abdomen reddish, slightly lighter towards apex. Mouth parts, antennae, and legs dark

reddish, tibiae and tarsi slightly darker.

Head (Figs 166a-d). Moderately short, rather wide, rather depressed. Anterior border gently

convex, lateral angle rounded, laterally projecting, lateral borders concave behind eyes. Clypeal suture

very inconspicuous, semicircular. Labrum rather wide and short, moderately overlapped by the

clypeus, apex feebly concave. Antennal groove laterally sharply bordered, latero-posteriorly with

slightly angulate area. Mental tooth triangular, short, apex acute. Wings of mentum wide, laterally

rounded, apex angulate. Glossa wide, tongue-like, apically convex, ventrally with indistinct keel, at

border with c. 12 elongate setae and additional pilosity on upper and lower surface and along border.

Terminal palpomere of maxillary palpus elongate, not securiform. Terminal palpomere of labial

palpus moderately wide, slightly securiform. Antenna narrow and rather elongate, parallel, Sth-9th

antennomeres <1.5 x as wide as long. Microreticulation absent, puncturation extremely fine, sparse.

Surface with weak sulcus medially of eyes, impilose, very glossy. Ventrolaterally of eyes with a row

of short setae. Suborbital field impunctate. Gula impilose.

Pronotum (Fig. 354). Large, fairly wide, convex, distinctly wider than long, base not much wider

than apex. Apical angles fairly produced, at apex rather acute, fairly oblique, almost attaining posterior

third of eyes. Apex fairly excised, convex in excision, almost unbordered. Sides fairly convex, barely

203

oblique. Margins moderately narrow, narrowly chanelled, coarsely bordered. Basal angles rather

shortly rounded off. Base straight, irregularly bordered. Surface near base without transverse

impression. Microreticulation absent, puncturation in basal half and in middle of apical half extremely

fine and superficial, visible only at very high magnification, sparse, in lateral part of apical half very

coarse and rather dense, surface impilose, highly glossy.

Elytra (Figs 354, 505). Rarether narrow and elongate, convex, though slightly depressed on disk,

rather parallel. Apex fairly wide, transverse, truncature slightly convex, apical angles widely rounded

off. Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal

channel fairly narrow throughout, partly concealed. Basal border incomplete, attaining almost mid of

base. Lateral border asetose. Series of umbilical pores consisting of 6 rather spaced pores behind

shoulder, pores difficult to see. Setae fairly elongate. Scutellum very wide. Inner 6 striae including

sutural stria marked by rows of fine punctures, outer striae absent. Microreticulation absent, intervals

impunctate, surface impilose, markedly glossy.

Lower surface. Prosternal process rather elongate, moderately wide, convex, apex wide, convex,

passing over in an acute angle from ventral surface, rather setose. Metepisternum very elongate,

c.2.2 x as long as wide, in posterior third not hollowed, but becoming very narrow towards apex.

Abdominal sterna with 1 elongate seta each side. Lower surface sparsely punctate and rather

elongately setose.

Legs. Moderately elongate, 1st tarsomere of protarsus slightly wider than long, tibial groove of

profemur deep, anterior plate deeply overlapping the groove for apical half, posterior border of groove

sharp. Femur wide. Metatibia fairly elongate, c. 5 x as long as wide, Ist tarsomere of metatarsus c. 1.5 x

as long as wide. d protarsus unknown.

d genitalia. Unknown.

? genitalia (Fig. 1661). Stylomere rather narrow, parallel, both median and lateral borders straight,

apex oblique, with 3 elongate setae. Lateral plate rather short, with 1 elongate apical seta.

Variation. Unknown.

Vivipary. Not confirmed in the examined material.

Habits. Largely Unknown. Holotype collected “under bark Bloodwood” in November.

Distribution (Fig. 621). Atherton Tableland, Northeastern Queensland. Known only from type

locality.

Material examined (1). Only the holotype.

Etymology. The name refers to the absence of punctures and hairs in the intervals.

Adelotopus montisatri, spec. nov.

Figs'167, 355,906, 621

Types. Holotype: 4, N. Slope, Black Mtn. ACT, 4 Sept 1971, K. R. Pullen, Kim Pullen Coll. (ANIC). - Paratypes:

18, Bombala (MMS); 1%, Australia: Black Mountain A. C.T. 13.X1.60, B. P. Moore (CMC); 12, Weston, A.C.T.,

8.1V.1969, E. Britton (ANIC); 12, Australien, ACT 119, Tidbinbilla Rge, 750 m, 6 km w. Cotter Dam, 9.12.1990,

M. Baehr (CBM).

Diagnosis. Rather small to medium-sized, elongate, convex, black species with reddish apex. Distin-

guished from related species by the glossy surface, narrow and elongate, coarsely and rather regularly

punctate pronotum, sparsely uniseriately punctate and setose elytral intervals, punctate elytral striae,

and the wide, at apex obliquely transverse, rather parallel stylomere.

Description

Measurements. Length: 4.25-5.0 mm. Ratios. Width/length of pronotum: 1.13-1.19; width base/

apex of pronotum: 1.25-1.32; width pronotum/head: 1.26-1.33; length/width of elytra: 1.85-1.90; length

elytra/pronotum: 2.32-2.40.

Colour (Fig. 355). Black, apical fourth of elytra reddish with fairly ill defined border. Lower surface

of head and thorax blackish, abdomen reddish, slightly lighter towards apex. Mouth parts, antennae,

and legs reddish, tibiae and tarsi barely darker.

204

h k

Figs 167a-l. Adelotopus montisatri, spec. nov. Details of head and genitalia. For legends see fig. 100.

Head (Figs 167a-d). Rather short, moderately wide, rather depressed. Anterior border gently

convex, lateral angle rounded, laterally projecting, lateral borders concave behind eyes. Clypeal suture

indistinct, almost invisible. Labrum rather wide and short, moderately overlapped by the clypeus,

apex feebly concave. Antennal groove laterally sharply bordered, latero-posteriorly with slightly

angulate area. Mental tooth triangular, short, apex acute. Wings of mentum wide, laterally rounded,

apex angulate. Glossa wide, tongue-like, apically convex, ventrally with indistinct keel, at border with

c. 12 elongate setae and additional pilosity on upper and lower surface and along border. Terminal

palpomere of maxillary elongate, not securiform. Terminal palpomere of labial palpus moderately

wide, barely securiform. Antenna rather narrow and moderately elongate, fairly parallel, Sth-9th

antennomeres c. 1.75x as wide as long. Microreticulation absent, puncturation moderately fine,

moderately dense. Surface with weak sulcus medially of eyes, impilose, glossy. Ventrolaterally of eyes

with a row of short setae. Suborbital field impunctate. Gula impilose.

Pronotum (Fig. 355). Rather narrow, elongate, highly convex, barely wider than long, rather

parallel, base barely wider than apex. Apical angles moderately produced, at apex rather acute, fairly

oblique, surpassing posterior border of eyes. Apex fairly excised, slightly convex in excision, faintly

bordered. Sides barely convex, faintly oblique. Margins very narrow, barely chanelled, coarsely

bordered. Basal angles rectangular, very shortly rounded off. Base faintly convex, coarsely bordered.

Surface near base with extremely shallow transverse impression. Microreticulation absent, punctura-

tion in basal half rather dense, coarse, though becoming much finer towards base, also somewhat finer

in middle, surface apparently impilose, highly glossy.

Elytra (Figs 355, 506). Narrow and elongate, convex, rather parallel. Lateral borders faintly convex.

Apex wide, slightly oblique, truncature convex, apical angles widely rounded off. Shoulders rounded,

basal margin slightly oblique, without setae behind shoulders. Marginal channel very narrow

throughout, mostly concealed. Basal border incomplete, attaining outer third of base. Lateral border

asetose. Series of umbilical pores consisting of 6, rarely 7 rather spaced pores behind shoulder, but

pores small and difficult to detect. Setae fairly elongate. Scutellum very wide. Inner 6 striae including

sutural stria marked by rows of moderately fine to fairly coarse punctures, outer striae absent.

Microreticulation absent, each interval with a irregular, very spaced row of fine, not rasp-like

punctures, each of which bears a rather short, erect hair, surface markedly glossy.

Lower surface. Prosternal process rather elongate, moderately wide, convex, apex wide, convex,

205

passing over in a slightly acute angle from ventral surface, rather setose. Metepisternum very elongate,

c. 2.4 x as long as wide, in posterior third not hollowed, but becoming very narrow towards apex.

Abdominal sterna with 1 elongate seta each side. Lower surface sparsely punctate and rather

elongately setose.

Legs. Moderately elongate, Ist tarsomere of protarsus slightly wider than long, tibial groove of

profemur deep, anterior plate deeply overlapping the groove for apical half, posterior border of groove

sharp. Femur wide. Metatibia fairly elongate, slightly >5 x as long as wide, 1st tarsomere of metatarsus

almost 2 x as long as wide.

d genitalia (Figs 167e-k). Genital ring moderately wide, rather convex, barely asymmetric, with

slightly asymmetric, rather small, fairly excised base. Sternum VII rather wide, apically almost straight,

with moderately deep excision, base excised, basal angles obtuse, lateral parts fairly short. Aedeagus

rather short, moderately depressed, in middle markedly widened, evenly narrowed to apex, faintly

asymmetric. Basal part fairly short, rather bent. Lower surface almost straight to gently convex. Apex

rather narrow, obtusely rounded. Orifice rather elongate, internal sac moderately complex, apparently

without a distinct oblique fold near apex. Both parameres elongate, rather parellel, with widely

rounded apex, left paramere considerably larger than right.

? genitalia (Figs 1671). Stylomere rather wide, fairly parellel, lateral border barely concave, apex

obliquely transverse, with 3 elongate setae at apex. Lateral plate moderately elongate, with 1-2

elongate apical setae.

Variation. Due to limited material only little variation noted in size and degree of puncturation of

surface.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. A specimen collected by me under bark of gum-type eucalypt in 750 m

altitude. Dated specimens captured in April, September, November, and December.

Distribution (Fig. 621). Australian Capital Territory and adjacent southernmost New South Wales.

Material examined (5). Only the type series.

Etymology. The name refers to the type locality of this species, the Black Monutain near Canberra.

Adelotopus puncticollis Notman, 1925

This species occurs in two subspecies, an eastern one in eastern South Australia, Victoria, and

southwestern New South Wales, and a western subspecies in the interior of southwestern Australia.

Diagnosis. Medium-sized, rather elongate, convex, black species with large reddish sutural spot in

apical two thirds of elytra, that leaves only a more or less wide lateral margin black. Distinguished

from related species by the glossy surface, fairly wide, slightly conical, coarsely and rather regularly

punctate pronotum, sparsely, though not uniseriately punctate, impilose elytral intervals, punctate

elytral striae, the acute, dentiform apex of the aedeagus, the wide, at apex more or less obliquely

transverse stylomere, and the short lateral plate.

Adelotopus puncticollis puncticollis Notman, 1925

Figs 168, 356, 507, 621

Adelotopus puncticollis Notman, 1925, p. 8, 9, 29; Csiki 1933, p. 1636; Moore et al. 1987, p. 52.

Types. Holotype: 3, 2588 Victoria, Type, Adelotopus puncticollis Ntm det Ntm. (AMNH).

Type locality: “Victoria”.

Diagnosis. Subspecies distinguished by generally larger size, wider elytral spot, less dark colour of

surface, and usually more conical and less convex lateral margins of pronotum that are less incurved

at base.

206

Figs 168a-l. Adelotopus puncticollis puncticollis Notman. Details of head and genitalia. For legends see fig. 100.

Description

Measurements. Length: 4.9-6.0 mm. Ratios. Width/length of pronotum: 1.24-1.35; width base/

apex of pronotum: 1.42-1.47; width pronotum/ head: 1.51-1.57; length/ width of elytra: 1.65-1.73; length

elytra/pronotum: 2.25-2.38.

Colour (Fig. 356). Piceous-black to black, lateral borders of pronotum and elytra more or less

distinctly reddish translucent. Elytra with large reddish sutural spot in apical two thirds that leaves

anteriorly amore or less wide lateral margin black, but the apical fifth is completely red. Lower surface

of head and thorax blackish, abdomen reddish. Mouth parts, antennae, and legs reddish, tibiae and

tarsi barely darker.

Head (Figs 168a-d). Rather short, moderately wide, rather depressed. Anterior border gently

convex, lateral angle rounded, laterally projecting, lateral borders concave behind eyes. Clypeal suture

more or less distinct, somewhat triangular, sometimes almost invisible. Labrum rather wide and short,

moderately overlapped by the clypeus, apex feebly concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly angulate area. Mental tooth triangular, short, apex acute.

Wings of mentum wide, laterally oblique, apex acutely angulate. Glossa wide, tongue-like, apically

convex, ventrally with indistinct keel, at border with c. 12 elongate setae and additional pilosity on

upper and lower surface and along border. Terminal palpomere of maxillary palpus elongate, not

securiform. Terminal palpomere of labial palpus fairly wide, rather securiform. Antenna rather narrow

and moderately elongate, fairly parallel, Sth-9th antennomeres c. 1.6 x as wide as long. Microreticula-

tion absent, puncturation rather dense, double, coarse punctures mixed with much finer ones. Surface

with weak sulcus medially of eyes, impilose, glossy. Ventrolaterally of eyes with a row of short setae.

Suborbital field impunctate. Gula impilose.

Pronotum (Fig. 356). Moderately narrow, fairly elongate, convex, slightly wider than long, more

or less conical, base distinctly wider than apex, widest near base. Apical angles moderately produced,

at apex rather acute, fairly oblique, surpassing posterior border of eyes. Apex fairly excised, slightly

convex in excision, faintly bordered. Sides more or less distinctly convex, slightly oblique. Margins

very narrow, barely chanelled, coarsely bordered. Basal angles rectangular, very shortly rounded oft.

Base faintly convex, moderately coarsely bordered. Surface near base with extremely shallow

transverse impression. Microreticulation absent, puncturation rather dense, coarse, though becoming

finer and less dense towards base, also somewhat finer in middle, surface impilose, highly glossy.

207

Elytra (Figs 356, 507). Rather narrow and elongate, fairly convex, rather parallel. Lateral borders

faintly convex. Apex wide, transverse, truncature markedly convex, in middle distinctly drawn in,

apical angles widely rounded off. Shoulders rounded, basal margin slightly oblique, without setae

behind shoulders. Marginal channel narrow throughout, partly concealed. Basal border incomplete,

attaining outer third of base. Lateral border asetose. Series of umbilical pores consisting of 6 or 7 rather

spaced pores behind shoulder, but pores small and difficult to detect. Setae fairly elongate. Scutellum

very wide. Striae including sutural stria marked by rows of moderately fine punctures. Microreticu-

lation absent, each interval with fine, sparse, irregular, not rasp-like punctures, surface asetose,

markedly glossy.

Lower surface. Prosternal process rather elongate, moderately wide, convex, apex wide, convex,

passing over in an almost right angle from ventral surface, barely setose. Metepisternum very elongate,

c. 2.2 x as long as wide, in posterior third not hollowed, but becoming very narrow towards apex.

Abdominal sterna with 1 elongate seta each side. Lower surface sparsely punctate and rather shortly

setose.

Legs. rather elongate, Ist tarsomere of protarsus slightly wider than long, tibial groove of

profemur moderately deep, anterior plate overlapping the groove only for apical forth, posterior

border of groove sharp. Femur wide. Metatibia elongate, c, 5.5 x as long as wide, Ist tarsomere of

metatarsus c. 2.2 x as long as wide.

d genitalia (Figs 168e-k). Genital ring rather wide, convex, barely asymmetric, with slightly

asymmetric, rather small, fairly excised base. Sternum VII rather wide, apically obliquely convex, with

rather deep excision, base faintly excised, basal angles rounded, lateral parts fairly short. Aedeagus

rather short, moderately depressed, in middle markedly widened, strongly narrowed to apex, faintly

asymmetric. Basal part fairly short, moderately bent. Lower surface strongly convex. Apex acute,

dentiform. Orifice rather elongate, internal sac complex, apparently without a distinct oblique fold

near apex. Both parameres elongate, rather parallel, with widely rounded apex, left paramere

considerably larger than right.

? genitalia (Figs 1681). Stylomere wide, both median and lateral borders slightly concave, apex

obliquely transverse or gently convex, with 3-4 elongate apical setae. Lateral plate short, with 2-4

elongate apical setae.

Variation. Apart from considerable diffences in size, there is some variation in shape of pronotum,

which may be rather conical or laterally rather convex, and in degree of puncturation on head,

pronotum, and elytra. The red elytral spot is always wide, rather well defined, and reaches the lateral

margin on a wide front.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. Dated specimens captured only in December. Most specimens, however,

undated and very old.

Distribution (Fig. 621). South Australia, Victoria, southwestern New South Wales, ? Tasmania. The

latter record is rather doubtful and refers to a single, barely dated species only labelled “Tasmania”

that was even identified “gaverei” which is a name of an asiatic Cryptocephalomorpha species. The

subspecies seems to live inland or at least in rather dry environments.

Material examined (14). SA: 1%, Kilkerran, filiformis Cast., Adelotopus filiformis Cast. (SAMA); 1%, Rev.

A. P. Burgess, Adelotopus filiformis Cast. (SAMA); 15, Rev. A. P. Burgess (SAMA). — Vic: 17, Inglewood, C. Oke,

Adelotopus puncticollis Notm. (NMV); 1%, Ballarat, C. Oke, Adelotopus puncticollis Notm. ? filiformis Cast., det.

puncticollis (NMV); 12, Sea Lake, Goudie, Adelotopus bicolor Cast. (SAMA); 14, 2588, Type, Adelotopus puncticollis

Ntm det Ntm. (AMNH). - Tas: 12, 1916 26, det. gaverei (SMTD). - NSW: 13, 22%, Gnalta Stn. nr Broken Hill,

9.X11.1964, N. Mc Farland (CBM, SAMA); 17, Murray R. S. Australia A. H. Elston, Adelotopus filiformis Cast. 2097,

A.H. Elston Coll., A. filiformis (AMS); 17, Murray R. S. Australia A. H. Elston, Adelotopus filiformis Cast. Id. by

A.M. Lea, A. H. Elston Coll. (AMS). - Aus: 18, F. Walker 1868 (OUM).

208

Figs 169e-l. Adelotopus puncticollis angustemaculatus, subspec. nov. Details of genitalia. For legends see fig. 100.

Adelotopus puncticollis angustemaculatus, subspec. nov.

Eiss 50) 1169, 357, 308,021

Types. Holotype: d, Norseman, 60 km E. 32.055, 122.35E Western Australia 24 Feb. 1989 M. S. Harvey &

M. E. Blosfelds (WAM 94/882). - Paratypes: 15, Broomehill 33.515, 117.38E Western Australia 6 June 1986

R. P. Mcmillan (WAM 94/867); 14, 12, Australien, WA 53, 5km n. Varley, 14.11.1987, M. Baehr (CBM);

12, Australien, WA 49, 33 km nnw. Raventhorpe, 13.11.1987, M. Baehr (CBM); 1%, 31.X. 90 Mi W Coolgardie,

West. Australia, J. Sedlacek Collector (CSB).

Diagnosis. Subspecies distinguished by generally lesser size, narrower elytral spot, deep black colour

of surface, and not conical and more convex lateral margins of pronotum that are distinctly incurved

at base.

Description

Measurements. Length: 4.8-4.9 mm. Ratios. Width/length of pronotum: 1.31-1.39; width base/

apex of pronotum: 1.42-1.46; width pronotum /head: 1.53-1.57; length/ width of elytra: 1.59-1.70; length

elytra/pronotum: 2.29-2.37.

Colour (Figs 50, 357). Deep glossy black, elytral spot narrower, leaving a wide, black margin.

Otherwise similar to nominate subspecies.

Head. Similar to nominate subspecies, though puncturation generally even denser.

Pronotum (Fig. 357). Rather similar to nominate subspecies, though less conical, laterally more

convex, and distinctly incurved to base, hence widest diameter clearly in front of base.

Elytra (Figs 50, 357, 508). Rather similar to nominate subspecies, though elytra generally slightly

wider and slightly more markedly punctate.

Lower surface. Similar to nominate subspecies

Legs. Similar to nominate subspecies.

& genitalia (Figs 169e-k). Rather similar to nominate subspecies, though in the few specimens

available genital ring even wider, sternum VII with bisinuate base and lateral parts rather elongate.

Aedeagus similar, though slightly shorter.

2 genitalia (Figs 1691). Rather similar to nominate subspecies, though apex obliquely transverse.

Variation. Little variation recognized, especially the three specimens from Varley and Rav-

enthorpe very similar, the single specimen from Coolgardie apparently not fully coloured and less

densely and coarsely punctate.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

209

Habits. Largely unknown. Specimens collected by me under bark of gum-type eucalypts. Holotype

“under bark of Eucalyptus sp.”. Dated specimens captured in February, June, October, and November.

Distribution (Fig. 621). Southwestern Australia.

Material examined (6). Only the type series.

Etymology. The name refers to the narrower elytral spot compared with that of the nominate subspecies.

rubiginosus-group

Diagnosis. Small to medium-sized, rather convex, reddish to piceous species. Labrum bisetose; glossa

c. 10-12-setose; lateral margin of pronotum rather narrow, basal angle widely rounded; basal border

line of elytra abbreviated, attaining at most middle of base; scutellar pore absent; lateral margin of

elytra without fringe of setae; series of umbilical pores with 6 subhumeral pores only; abdominal

sterna usually with 1, rarely without ambulatory seta each side; sternum VI without a fringe of longer

setae at apical margin; tibiae, especially metatibia rather depressed; all femora wide and depressed;

aedeagus rather wide and short, asymmmetric, apex rounded off; internal sac complicate, apex with

a distinct oblique fold; apical margin of ? tergum VIII with 4 median setae, very rarely with additional

lateral setae.

Larvae. 1st instar larvae known of 6 species.

Distribution. 21 species that occur virtually throughout continental Australia, though not in Tasma-

nia and New Guinea.

Systematic position. This group is certainly the adelphotaxon of the laevis-group (see below). Both

groups, however, are perhaps the adelphotaxon of the seriepunctatus-group. Compared with the laevis-

group the rubiginosus-group is more apomorphic in the absence of elongate setae at the margins of the

elytra, but more plesiomorphic in the rounded apex of the aedeagus, and perhaps also in the presence

of a distinct oblique fold in the apex of the internal sac. Both groups are more apomorphic than the

whole lineage from the politus-group to the linearis-group by their uniformly light colour, highly

convex body shape, and markedly abbreviated basal border of elytra.

Note. The many taxa of externally very similar, uniformly reddish but, on the other hand, very

variable beetles can be only distinguished by scrutinized examination of their genitalia. The ?

genitalia, however, are generally more variable, so correct identification of females may be quite

commonly difficult or even impossible. Due to the very limited knowledge of distribution and habits

of all species of this group, decisions between what is a species, a valid subspecies, or merely a local

variation are especially difficult. So it is possible that some taxa named herein on the base of single

specimens bearing aberrant genitalia or otherwise external character states, actually do not merit the

rank given herein.

Adelotopus rubiginosus Newman, 1856

Figs 170, 358, 509, 622

Adelotopus rubiginosus Newman, 1856, p. 128; Notman 1925, p. 7, 10, 29; Csiki 1933, p. 1636; Moore et al. 1987,

P- 22.

Adelotopus castaneus Castelnau, 1867, p. 33; 1868, p. 119; Macleay 1871, p. 95; Notman 1925, p. 7, 10, 28; Csiki 1933,

p. 1634; Moore 1967, p. 321; Moore et al. 1987, p. 50 (new synonymy).

Types. Of rubiginosus: Lectotype (by present designation): d, Type H.T., Type, Adelotopus rubiginosus Newm. Z.

5028 (BMNH).

Of castaneus: Lectotype (by present designation): d, Swan River, Coll. Castenau, Castaneus Cast. Swan Riv.,

Holotypus Adelotopus castaneus Castelnau, 1867 (MCSN).

Type localities. Of rubiginosus: Australia, no locality given.

Of castaneus: “Swan River”, Western Australia.

210

Figs 170a-l. Adelotopus rubiginosus Newman. Details of head and genitalia. For legends see fig. 100.

Diagnosis. Small to medium-sized, completely reddish species, distinguished by but moderately

wide, convex pronotum, absence of microreticulation even on head, dense and rather coarse punctu-

ration of head and pronotum, asymmetric d gential ring, moderately widened, on lower surface not

striate aedeagus with shortly rounded apex, apically rounded left paramere, and wide, apically

obliquely transverse stylomere. Further distinguished from the most closely related species

A. distinguendus, spec. nov. by narrower, more convex pronotum, narrower aedeagus with narrower

apex, wider stylomeres and longer lateral plate; and from A. foliaceus, spec. nov. by even narrower

aedeagus, longer stylomeres, and absence of additional setae on the apical margin of ? tergum VII.

Description

Measurements. Length: 3.7-5.5 mm. Ratios. Width/length of pronotum: 1.40-1.51; width base/

apex of pronotum: 1.37-1.48; width pronotum/head: 1.47-1.61; length/ width of elytra: 1.54-1.65; length

elytra/pronotum: 2.20-2.48.

Colour. Upper and lower surface including mouth parts, antennae, and legs more or less light

reddish, sometimes apex of elytra faintly darker.

Head (Figs 170a-d). Rather short, fairly wide, rather depressed. Anterior border but gently convex,

lateral angle rounded, laterally faintly projecting, lateral borders slightly narrowed behind eyes.

Clypeal suture only at base distinct, in middle widely interrupted. Labrum rather wide and short,

moderately overlapped by the clypeus, apex moderately concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly convex area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa fairly wide, tongue-like, apically

convex, ventrally with distinct keel, at border with c. 10-12 elongate setae. Terminal palpomere of

maxillary palpus elongate, slightly widened, though not securiform. Terminal palpomere of labial

palpus fairly wide, rather securiform. Antenna short and wide, Sth-9th antennomeres c. 1.8 x as wide

as long. Microreticulation absent, puncturation modserately fine, fairly dense. Surface with very weak

sulcus medially of eyes, impilose, highly glossy. Ventrolaterally of eyes with a row of short setae.

Suborbital field punctate and shortly setose. Gula impilose.

Pronotum (Fig. 358). Rather wide, markedly convex, slightly humped, distinctly wider than long,

base distinctly wider than apex, widest near base. Apical angles moderately produced, at apex

obtusely rounded, fairly oblique, attaining posterior border of eyes. Apex fairly excised, markedly

convex in excision, feebly bordered. Sides gently convex, rather oblique. Margins rather wide, rather

chanelled, finely bordered. Basal angles widely rounded off. Base faintly convex, moderately coarsely

bordered. Surface near base with shallow transverse impression. Microreticulation absent, punctura-

tion fairly coarse, dense, surface impilose, highly glossy.

Elytra (Figs 358, 509). Moderately elongate, rather convex, though slightly depressed on disk,

rather parallel, though faintly narrowed in basal third. Lateral borders almost straight. Apex wide,

slightly oblique, truncature barely convex, apical angles widely rounded off. Shoulders rounded, basal

margin slightly oblique, sometimes with few elongate setae behind shoulders. Marginal channel

narrow, in basal third even narrowed, then widened again, mostly concealed. Basal border incomplete,

attaining outer third of base. Lateral margin asetose. Series of umbilical pores consisting of 6 rather

spaced pores behind shoulder. Setae fairly elongate. Striae including sutural stria absent. Microretic-

ulation absent, puncturation rather coarse, fairly dense, surface impilose, markedly glossy.

Lower surface. Prosternal process rather elongate, narrow, convex, apex narow, compressed,

passing over in an almost right angle from ventral surface, shortly setose. Metepisternum elongate,

c. 1.8-1.9 x as long as wide, in posterior third not hollowed. Abdominal sterna with 1 elongate seta each

side. Lower surface rather densely punctate and shortly setose.

Legs. Moderately elongate, Ist tarsomere of protarsus about as long as wide, tibial groove of

profemur moderately deep, anterior plate overlapping the groove for apical third only, posterior

border of groove sharp. Femur wide. Metatibia moderately elongate, c. 5x as long as wide, Ist

tarsomere of metatarsus c. 1.6 x as long as wide.

d genitalia (Figs 170e-k). Genital ring rather wide, convex, fairly highly asymmetric, left arm

convex, right almost straight, with elongate apex, with slightly asymmetric, rather small, barely

excised base. Sternum VII rather wide, apically evenly convex, with moderately deep excision, base

bisinuate, basal angles obtusely rounded, lateral parts fairly elongate. Aedeagus rather short, fairly

depressed, in middle moderately widened, slightly asymmetric, left side straight, right side straight to

faintly convex. Basal part rather long, moderately bent. Lower surface straight to gently convex, not

perceptibly striped. Apex moderately narrow, shortly rounded off, rather symmetric. Orifice rather

elongate, internal sac complex, with a distinct oblique fold near apex. Both parameres large, rather

elongate, square, with widely rounded apex, left paramere considerably larger than right, with an

oblique edge, upper part of lateral side moderately striped.

? genitalia (Fig. 1701). Stylomere wide, apex wide, obliquely transverse, lateral margin concave,

with 2-3 elongate apical setae. Lateral plate elongate, with 3-4 elongate and 2-3 short apical setae.

Apical margin of ? tergum VIII without additional setae.

Variation. There is considerable variation of size, relative width of pronotum, length of elytra, and

puncturation. This variation is somewhat regional, as in Central Australian specimens the punctura-

tion on head and pronotum is generally slightly less coarse and dense, and the lateral margin of the

elytra bears sometimes some hairs also behind shoulders. Because d and ? genitalia generally show

little variation, I refrained from giving the Central Australian specimens an separate taxonomic status.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unknown. A specimen collected by me under bark of gum-type eucalypt, other

specimens collected “under bark”. Dated specimens captured from October to April, though relatively

few specimens dated. This species occurs also far inland.

Distribution (Fig. 622). Eastern Australia from South Australia through Victoria, Australian Capital

Territory, New South Wales to north Queensland, also central Northern Territory and southwestern

Australia, ? Batchian. The latter record from this Molukkan island is certainly wrong.

Material examined (336). SA: 14, 32%, 758, Port Lincoln Blackburn, aphodioides Westw., 7104 Adelotopus

aphodioides Westw. (SAMA); 12, 758, Port Lincoln Blackburn, Sharp Coll., det. rubiginosus (BMNH); 388, 9P 2,

Alde 2/4, Adelaide Griffith, Griffith Collection Id. by A.M. Lea, 254 Adelotopus aphodioides Westw. 2472 (SAMA);

14, 42158, rufescens Chd., Schomb., castaneus Casteln. Adelaide (MNHB); 12, Adelaide, det. castaneus (MNHB);

ld, Adelaide, det. castaneus (MNHB); 12, Adelaide, Adelaide Coll. Castelnau, aphodioides (MCSN); 2?, Adelaide

Coll. Castelnau, det. aphodioides (MCSN); 15, ee Mt. Lofty, J. G. ©. Tepper, F. E. Wilson Collection,

A. aphodioides, Westw. As id. by Blackburn (NMV); 14, 12, Yorketown, CNHM 1955, Karl Brancsik Coll. Ex

Eduard Knirsch, Adelotopus castaneus Casteln., ee castaneus Cast. det. G. E. Ball 1987 (FMNH, UASM);

3,329, Yorketown, CNHM 1955, Karl Brancsik Coll. Ex Eduard Knirsch, Adelotopus castaneus det. Ball (FMNH,

UASM); 18, Yorketown, CNHM 1955, Karl Brancsik Coll. Ex Eduard Knirsch (UASM); 15, Yuma Ck. by Yuma

23.X.1973. G. F. Gross (SAMA); 1d, 12, Wentworth Murray R. 25.X1l.30, C. E. Clarke Collection (BMNH);

12, Wentworth Redcliffs Murray R. 24.-27.X11.30 C.E.C., C. E. Clarke Coll. (BMNH); 17, Murray R. A. H. Elston,

A.H. Elston Collection, det. aphodioides (AMS); 753, Mt. Serle N. Flinders Ra. Hale & Tindale (SAMA); 259,27%,

Everard Rgs., S.A. to Warburton Rgs., W.A. A. Brumby (SAMA); 19, 2 mi. from Odlawirra (Adel. site)

23.X11.1973, G. F. Gross (SAMA); 22%, A. H. Elston, 363 Adelotopus aphodioides Westwood Id. by T. G. Sloane

(AMS); 12, C. French’s Coll., det. aphodioides (NMV); 19, A. Austr. Witte., 19, Soc. Ent. Belg. Coll. PUTZEYS,

R.1.Sc.N.B. 1.G. Coll. gen. (IRSNB); 13, 22, det. aphodioides (ANIC); 238, 17, Adelotopus Aphodioides, Westw.

(SAMA); 19, Nat. Mus. Victoria (NMV); 12, Ex Musaeo H. W. Bates 1892 (MNHN). - Vic: 2722, Lake Hattah

J. E. Dixon, Adelotopus micans Blk®. (NMV); 78,72%, Lake Hattah J. E. Dixon (ANIC, NMV); 277, Lake Hattah,

X1.1924 C. Oke (NMV); 13, 322, Echuca 14.XIl.1966, leg. Bornemissza (HNMB); 17, Birchip III.04, Adelotopus

aphodioides Westw., J. C. Goudie Collection (NMV); 12, Birchip, J. C. Goudie (SAMA); 234, Dimboolah 9.1.87

Tepper (SAMA); 17, Mallee District, Nat. Mus. Victoria - C. French’s Collection (NMV); 18, Melbourne:, Pascoe

Coll., castaneus Cast. (BMNH); 238, 1%, K 12380, Adelotopus aphodioides West. Melbe. (AMS); 17, Melbourne, Ex

Musaeo H. W. Bates 1892 (MNHN); 334, 12, Melbourne Coll. Castelnau. Aphodioides W. (Melbourne) (MCSN);

18, 12, Melbourne Coll. Castelnau, det. aphodioides (MCSN); 17, Melbourne Coll. Castelnau, 80, aphodioides

Westw. Melb. (MCSN); 13, 12, Alberton, Adelotopus aphodioides Ww. Det. B. P. Moore "66 (CMC, FMT); 15,17,

Alberton, det. aphodioides (FMT); 15, Woorinen, 5.1.36 F. E. Wilson, F. E. Wilson Coll., A. castaneus (NMV);

238, Kiata, 31.X11.18 F. E. Wilson, F. E. Wilson Collection, Adelotopus aphodioides Westw. Id. by T. G. Sloane

(NMV); 284, Kiata, 1.1918 R. Oldfield, F. E. Wilson Collection, Adelotopus aphodioides Westw. Id. by T. G. Sloane

(NMV); 18, 12, Kiata, 1.1918 R. Oldfield, T.G.S./61 (ANIC); 222, Kiata, 1.1918 R. Oldfield, 363 Adelotopus

aphodioides Westw., A. H. Elston Collection (AMS); 235, Kiata, I.1918R. Oldfield, (ANIC); 15,1, Wallan, C. Oke,

det. aphodioides (NMV); 238, 17, Kerrisdale 1.1.24, J. E. Dixon (NMV); 638, 327%, Kerrisdale 1.1.24 (NMV);

12, Gippsland 2, det. aphodioides (NMV); 234, Field Mus. (F. Psotas Coll.), Adelotopus castaneus det. Ball (FMNH);

18, P!. West Victoria, det. rubiginosus (BMNH); 15, 17, Nat. Mus. Victoria, C. French’s Collection, Adelotopus

Hydrobioides Westw. (NMV); 12, Field Mus. (F. Psotas Coll.), Adelotopus aphodioides Westwood, Adelotopus

castaneus det. Ball (FMNH); 14, 875, 97, Adelotopus aphodioides Westw., 35, Howitt Colln (NMV); 19, 259, W.

Edwards, det. punctatus (MCZ); 12, 2592, castaneus Cast. (AMNH); 377, 2592, det castaneus (AMNH); 12, 2592,

Ex Musaeo H. W. Bates 1892 (MNHN); 1%, Coll. B. Schwarzer, brunneus Cast. (SMF); 234, 2??, Edwards, Fry

Coll. det. rubiginosus (BMNH); 13, Ex Musaeo H. W. Bates 1892 (MNHN); 18, 1%, Janson Acq. (MNHN). —

ACT: 5922, 5 (sex?), Canberra NSW 1926 in WF (ANIC); 2 (sex?), Canberra N.S.W. 1192 In WF* (ANIC). -

NSW: 14, Culcairn, E. W. Ferguson Collection (ANIC); 16, 4929, Dubbo, 328. Rev. A. S. Barrett, K 60444, det.

aphodioides (AMS); 18, Wentworth Murray Rv. 25.X11.30, C. E. Clarke Collection B. M. 1957-24., A. scolytides

Newm., scolytides (BMNH); 15, 2492 Bathurst (SAMA); 229, Coragabal, F. H. Taylor, Adelotopus sp.,J.C. Goudie

Collection (NMV); 238, 12, Coragabal, F. H. Taylor (ANIC); 2??, Grenfell, E. W. Ferguson Collection,

A. aphodioides Westw. As id. by Blackburn (ANIC); 239, Grenfell, E. W. Ferguson Collection (ANIC); 1%,

Caldwell, Goudie, J. C. Goudie Collection (NMV); 42, Morilla E.W.F. 8.1.08, Not A. aphoides (sic!) by obtuse

angles thorax (ANIC); 938, 1522, 1 (sex?), Morilla 11.11.11 (ANIC); 48, 72%, Morilla (ANIC); 12, 1 (sex?,

fragment), sp 1sp ?, Morilla (ANIC); 1%, Westl. Riverina Wait leg. Lüddemann ded. (DEIB); 17, M.F.L. (BMNH);

12, Clarence River Coll. Castelnau, Clarence Riv., det. aphodioides (MCSN); 17, N. Galle, rufescens Chaud., Ex

Musaeo Mniszech (MNHN). - Qld: 12, Qld 2, 5 km s. Ipswich 4.X1.1990, M. Baehr (CBM); 15, 1?, Wide Bay,

..lotop. 2491 ..eensl. (SAMA); 1?, 10508, Godeffroy Collection, Adelotopus castaneus (Casteln.) Peak Downs

(NMV); 14, 1?, Mus. Godefroy Peak Downs, P.D., 10508, Adelotopus castaneus Casteln., Museum Leiden

Adelotopus castaneus Cast. Det., Castaneus Casteln. (NNML); 339, 12, Port Denison, Port Denison Coll.

Castelnau, det. aphodioides (MCSN),; 15, Mt. Isa 11.1986, J. H. Sedlacek Collector (CSB); 27%, Masters, Fry Coll.

det. rubiginosus (BMNH); 15, Simson, Fry Coll. det. rubiginosus (BMNH). - NT: 1%, Alice Sp. IV.71, J. Sedlacek

Collector (CSB); 234, 12, Alice Springs 20-24.111.1971,J. &M. Sedlacek Colls. Bishop Museum (BMH); 15, Alice

Springs 350-650 m 20-24.X1.1968, N. L. H. Krauss Coll. Bishop Museum (BMH); 17, Comiston Station, near Alice

Springs M. W. Mules (SAMA); 12, 23.375 133.54E 10 km N by E of Alice Springs 6.X1.1979 T. Weir (ANIC);

338, 1622, Ellery Ck. e. Hermannsburg, 1.11.1989 leg. P. A. Meyer (CBM, ZSM); 12, Daly Waters, J. Sedlacek

Collector (CSB); 18, Daly Waters (CSB); 1%, IV.71, J. Sedlacek Collector (CSB). - WA: 14, Swan River, Coll.

Castenau, Castaneus Cast. Swan Riv., Holotypus Adelotopus castaneus Castelnau, 1867 (MCSN); 23, Australia

occid. 1192 (HNMB). - Aus: 14, rufescens Chaud., Riv. Paroo Castelnau, Ex Musaeo Chaudoir (MNHN);

13, Rhynie, Blackb’s Coll., 7138 aus lia (SAMA); 18, 12, Adelotopus hydrobioides Westw. Hay (), 493, G. €

Champion Coll. (BMNH); 15, Nov. Holl. N. West, Fry Coll. det. rubiginosus (BMNH); 18, N. Holl. m. Adelotopus

distinctus Chaud., Sammlung O. Langenhan (SMTD); 235, 1214, det. punctatus (MCZ); 12, Coll. B. Schwarzer,

Adelotopus Aphodioides Wst. (SMF); 18,758, Adelotopus aphodioides Westw. Blackb. 1890 (ANIC); 17, W. Edwards,

aphodioides, det. punctatus (MCZ); 15, H. Edwards, Adelotopus aphodioides, det. punctatus (MCZ); 18, Coll. E. Witte,

rufescens (SMF); 12, Dohrn 91, Adelotopus castaneus Casteln., Castaneus Cast. (NHMW); 12, 18947, Fry Coll. det.

rubiginosus (BMNH); 14, 1214, det. punctatus (MCZ); 18, aphodioides Westw. (ANIC); 12, det. rubiginosus

(BMNH); 168, 12, J. Walker 1860 (OUM); 19, Adelotopus castaneus Cast. det... (IRSNB); 14, 1882 I(NHMW). -

?:1?, Mudgee Shphen (?), 254 ?, H. J. Carter Coll., Adelotopus aphodioides Westw. (NMV); 522, Germanton

27.11.11 J.G.S (?) (ANIC); 234, 27%, E. W. Ferguson Collection, Bachantus (?) Valley, 29.X11.85 Tepper (ANIC);

18, 1 (sex?), Gunning (MMS); 12, Berwick (NMV); 13, Type H.T., Adelotopus rubiginosus Newm. Z. 5028,

lectotype! (BMNH); 19, Coll. L. W. Schaufuss, det. castaneus (MNHB); 15, Adelotopus, Ex Musaeo H. W. Bates

1892 (MNHN); 238, 1%, rufescens Chaud., Ex Musaeo Chaudoir (MNHN); 15, 54 35, det. rubiginosus (BMNH);

12, 157, aphodioides Westw., Adelotopus punctatus Cast. (MCZ); 234, 845, 36, 37, Howitt Colln, det. aphodioides

(NMV); 222,2 (sex?, fragments), A. aphodioides (ANIC); 15, 54 35, det. rubiginosus (BMNHA); 15, 1?, Janson Acg.

(MNHN); 228, Collect. Plason (NHMW); 1%, C. G. Oke Collection No locality (NMV); 13, 17, M 254 (ANIC);

2292,70 (ANIC); 13,12 (ANIC); 135 (NMV). - Indonesia: 14, Batchian, Bowring 63-47*, det. rubiginosus (BMNH).

Adelotopus distinguendus, spec. nov.

Fies 171, 359, 510, 622

Types. Holotype: d, Broomehill W. Australia 10.1.82 R. P. McMillan (WAM 94/865). — Paratypes: 17, same data

(WAM 94/866); 12, R. P. McMillan Date 7.1.52 Culham Inquiline, 168, Adelotopus sp. (WAM 94/849); 17,

R. P. McMillan Date 23.12.51 Culham (CBM, WAM 94/848).

Diagnosis. Small to medium-sized, completely reddish species, distinguished by fairly wide prono-

tum, absence of microreticulation even on head, dense and rather coarse puncturation of head and

pronotum, narrow, rather symmetric d gential ring, rather widened, on lower surface not striate

aedeagus with widely rounded apex, and rather narrow, apically obliquely transverse stylomere.

Further distinguished from the most closely related species A. rubiginosus Newman by wider, less

convex pronotum, wider aedeagus with wider apex, narrower stylomeres and much shorter and wider

lateral plate; and from A. foliaceus, spec. nov. by wider pronotum, slightly narrower aedeagus with

widely rounded apex, longer stylomeres, and absence of additional setae on the apical margin of %

tergum VIN.

Description

Measurements. Length: 3.8-4.9 mm. Ratios. Width/length of pronotum: 1.51-1.55; width base/

apex of pronotum: 1.44-1.48; width pronotum/head: 1.56-1.62; length/width of elytra: c. 1.45-1.55;

length elytra/pronotum: c. 2.35-2.50.

Colour. Upper and lower surface including mouth parts, antennae, and legs more or less light

reddish, sometimes apex of elytra faintly darker.

Head (Figs 171a-d). Rather short, fairly wide, rather depressed. Anterior border but gently convex,

lateral angle rounded, laterally faintly projecting, lateral borders slightly narrowed behind eyes.

Clypeal suture only at base distinct, in middle widely interrupted. Labrum rather wide and short,

moderately overlapped by the clypeus, apex moderately concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly convex area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa fairly wide, tongue-like, apically

convex, ventrally with distinct keel, at border with c. 10-12 elongate setae. Terminal palpomere of

maxillary palpus elongate, barely widened. Terminal palpomere of labial palpus moderately wide,

rather securiform. Antenna rather short and wide, Sth-9th antennomeres c. 1.8 x as wide as long.

Microreticulation absent, puncturation moderately fine, fairly dense. Surface with very weak sulcus

medially of eyes, impilose, highly glossy. Ventrolaterally of eyes with a row of short setae. Suborbital

field punctate and shortly setose. Gula impilose.

Pronotum (Fig. 359). Rather wide, markedly convex, slightly humped, distinctly wider than long,

base distinctly wider than apex, widest near base. Apical angles moderately produced, at apex

obtusely rounded, fairly oblique, attaining posterior border of eyes. Apex fairly excised, markedly

convex in excision, feebly bordered. Sides gently convex, rather oblique. Margins rather wide, rather

chanelled, finely bordered. Basal angles widely rounded off. Base faintly convex, moderately coarsely

bordered. Surface near base with shallow transverse impression. Microreticulation absent, punctura-

tion fairly coarse, rather dense, surface impilose, highly glossy.

Elytra (Figs 359, 510). Rather short and wide, rather convex, though slightly depressed on disk,

rather parallel, though faintly narrowed in basal third. Lateral borders almost straight. Apex wide,

slightly oblique, truncature barely convex, apical angles widely rounded off. Shoulders rounded, basal

margin slightly oblique, sometimes with few elongate setae behind shoulders. Marginal channel

Figs 171a-l. Adelotopus distinguendus, spec. nov. Details of head and genitalia. For legends see fig. 100.

narrow, in basal third even narrowed, then widened again, mostly concealed. Basal border incomplete,

attaining outer third of base. Lateral margin asetose. Series of umbilical pores consisting of 6 rather

spaced pores behind shoulder. Setae fairly elongate. Striae including sutural stria absent. Microretic-

ulation absent, puncturation rather coarse, fairly dense, surface impilose, markedly glossy.

Lower surface. Prosternal process rather elongate, narrow, convex, apex narrow, compressed,

passing over in an almost right angle from ventral surface, shortly setose. Metepisternum fairly

elongate, c. 1.6-1.7 x as long as wide, in posterior third not hollowed. Abdominal sterna with 1 elongate

seta each side. Lower surface rather densely punctate and shortly setose.

Legs. Moderately elongate, Ist tarsomere of protarsus almost as long as wide, tibial groove of

profemur moderately deep, anterior plate overlapping the groove for apical third only, posterior

border of groove sharp. Femur wide. Metatibia moderately elongate, c. 5x as long as wide, Ist

tarsomere of metatarsus c. 1.6 x as long as wide.

d genitalia (Figs 171e-k). Genital ring rather narrow, little convex, barely asymmetric, with

elongate apex, with slightly asymmetric, rather small, barely excised base. Sternum VII rather wide,

apically evenly convex, with moderately deep excision, base faintly bisinuate, basal angles obtusely

rounded, lateral parts fairly elongate. Aedeagus short, depressed, in middle rather widened, fairly

asymmetric, both sides faintly concave. Basal part rather long, moderately bent. Lower surface almost

straight, not perceptibly striped. Apex rather wide, widely rounded off. Orifice elongate, internal sac

complex, with a distinct oblique fold near apex. Both parameres large, rather elongate, square, with

widely rounded apex, left paramere considerably larger than right, upper part of lateral side

moderately striped.

? genitalia (Fig. 1711). Stylomere moderately wide, rather parallel, apex wide, obliquely trans-

verse, with 2-4 elongate apical setae. Lateral plate short and wide, with 3-4 elongate apical setae. Apical

margin of ? tergum VIII without additional setae.

Variation. There is some variation of size and relative shape of pronotum and elytra, and of

puncturation of surface.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unknown. One specimen found as “inquiline”, unfortunately without additional

statement. Thus far captured in December and January.

Distribution (Fig. 622). Southwestern Australia.

Material examined (4). Only the type series.

Etymology. The name refers to the high similarity of this species and A. rubiginosus Newman.

Adelotopus foliaceus, spec. nov.

Figs 172, 360, 511, 622

Types. Holotype: d, Leas, 3891. 254 Adelotopus aphodioides Westw. N. S. Wales (SAMA). — Paratypes: 14, Mullaley,

Jan 57, N.S.W. F. E. Wilson, F. E. Wilson Collection (NMV); 18, Kurting 23.12.23 (NMV); 33, Dalby, O. Mrs.

F. H. Hobler (CBM, SAMA); 738, 32%, Australia: N. Queensld. Richmond 1917/1918 (CBM, NHMW).

Diagnosis. Rather small, completely reddish species, distinguished by rather narrow, convex prono-

tum with rather wide apex and narrow lateral margins, absence of microreticulation even on head,

rather fine puncturation on pronotum though coarse puncturation on elytra, markedly convex d

genital ring with elongate apex, very wide, asymmetric, on lower surface only laterally somewhat

striate aedeagus with rather acute apex, large, apically straight, strongly striate left paramere, and

short stylomeres. Further distinguished from the most closely related species A. rubiginosus Newman

by much wider aedeagus, shorter stylomeres, and presence of additional setae on the apical margin

of ? tergum VII; and from A. distinguendus, spec. nov. by narrower pronotum, even wider aedeagus

with rather acute apex, shorter stylomeres, and presence of additional setae on the apical margin of ?

tergum VIN.

Description

Measurements. Length: 4.3-4.9 mm. Ratios. Width/length of pronotum: 1.44-1.51; width base/

apex of pronotum: 1.37-1.42; width pronotum/head: 1.45-1.51; length/width of elytra: 1.56-1.60; length

elytra/pronotum: 2.35-2.42.

Colour. Upper and lower surface including mouth parts, antennae, and legs light reddish.

Head (Figs 172a-d). Rather short, fairly wide, rather depressed. Anterior border faintly convex,

lateral angle rounded, laterally slightly projecting, lateral borders distinctly narrowed behind eyes.

Clypeal suture only at base distinct, in middle widely interrupted. Labrum rather wide and short,

moderately overlapped by the clypeus, apex moderately concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly convex area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa fairly wide, tongue-like, apically

convex, ventrally with distinct keel, at border with c. 10-12 elongate setae. Terminal palpomere of

maxillary palpus slightly widened, barely securiform. Terminal palpomere of labial palpus wide,

securiform. Antenna moderately elongate, in middle distinctly widened, 8th-9th antennomeres c. 1.8 x

as wide as long. Microreticulation absent, puncturation rather fine, fairly dense. Surface with weak

sulcus medially of eyes, impilose, glossy. Ventrolaterally of eyes with a row of short setae. Suborbital

field punctate and shortly setose. Gula impilose.

Pronotum (Fig. 360). Moderately wide, rather highly convex, base distinctly wider than apex,

widest near base. Apical angles feebly produced, at apex obtuse, fairly oblique, barely ataining

posterior border of eyes. Apex fairly excised, very convex in excision, faintly and irregularly bordered.

Sides moderately convex, fairly oblique. Margins rather narrow, narrowly chanelled, finely bordered.

Basal angles widely rounded off. Base straight, rather coarsely bordered. Surface near base with

shallow transverse impression. Microreticulation absent, puncturation moderately fine, rather dense,

surface impilose, highly glossy.

Elytra (Figs 360, 511). Moderately wide, convex, faintly depressed on disk, parallel. Lateral

borders usually slightly excised in basal third. Apex wide, slightly oblique, truncature barely convex,

apical angles widely rounded off. Shoulders rounded, basal margin slightly oblique, without setae

behind shoulders. Marginal channel rather narrow, partly concealed. Basal border incomplete,

attaining about middle of base. Lateral margin asetose. Series of umbilical pores consisting of 6 rather

spaced pores behind shoulder. Setae fairly elongate. Striae including sutural stria absent. Microretic-

ulation absent, puncturation moderately coarse, rather dense, surface impilose, highly glossy.

Lower surface. Prosternal process rather short, narrow, convex, apex short, narrow, compressed,

216

Figs 172a-l. Adelotopus foliaceus, spec. nov. Details of head and genitalia. For legends see fig. 100.

passing over in an almost right angle from ventral surface, shortly setose. Metepisternum elongate,

c. 1.8 x as long as wide, in posterior third not hollowed. Abdominal sterna with 1 elongate seta each

side. Lower surface rather densely punctate and shortly setose.

Legs. Rather elongate, Ist tarsomere of protarsus as long as wide, tibial groove of profemur rather

shallow, anterior border straight, anterior plate overlapping the groove for about apical half, posterior

border of groove sharp. Femur wide. Metatibia rather elongate, c. 5.5-6 x as long as wide, Ist tarsomere

of metatarsus c. 2.3-2.4 x as long as wide.

ö genitalia (Figs 172e-k). Genital ring rather wide, convex, moderately asymmetric, with rather

elongate apex, with slightly asymmetric, small, slightly excised base. Sternum VII wide, apically

irregularly convex, with rather shallow excision, base gently bisinuate to slightly convex, basal angles

obtusely rounded, lateral parts rather elongate. Aedeagus short, depressed, in middle markedly

widened, asymmetric, left side straight or faintly concave, right convex. Basal part moderately long,

moderately bent. Lower surface convex, on left side slightly striped. Apex narrow, acute to slightly

obtuse, slightly asymmetric. Orifice rather elongate, internal sac complex, with a large oblique fold

near apex. Both parameres rather large, right with widely rounded apex, left considerably larger than

right, with transversely cut apex and strong stripes on upper part.

? genitalia (Fig. 1721). Stylomere short and wide, apex wide, obliquely rounded, with 4 markedly

elongate apical setae. Lateral plate rather elongate, with 2-3 elongate apical setae. Apical margin of ?

tergum VIII with 2-3 additional lateral setae.

Variation. Rather little variation noted.

Vivipary. Not confirmed in the examined material.

Habits. Unknown. Dated specimens collected in December and January.

Distribution (Fig. 622). New South Wales, southern and northern central Queensland, ? northern

Victoria. The latter record refers to a specimen labelled “Kurting” without state record. I think that

means a locality in northern Victoria.

Material examined (16). Only the type series.

Etymology. The name refers to the very wide, foliaceous aedeagus.

Adelotopus laticollis, spec. nov.

Figs‘173, 361,512, 623

Types. Holotype: 3, Longreach Q.: A. M. Lea, 7669 Adelotopus castaneus Cast. Queensland (SAMA) [holotype

mounted with seven paratypes on same card, marked HT.].- Paratypes: 334, 4?9, same data (SAMA),;

12, P. Lincoln, Port Lincoln Coll. Castelnau, det. aphodioides (MCSN); 2? 9, Gawler Range SA, Austral. 12.1972,

M. Baehr (CBM); 12, Adelaide SA Griffith, Griffith Coll. Id. by A. M. Lea, 254 Adelotopus aphodioides Westw.

5.A.2492 (SAMA); 27%, Adelaide Lea, 254, Leas, 3890 Adeletopus aphodioides Westw. S. Australia (SAMA);

?,41408, aphodioides Westwd. Adelaide (NHMB); 1%, Adelaide 5. Australia (SAMA), 1?, Adelaide (OUM);

272%, Adelaide Coll. Castelnau, Adelaide, aphodioides (MCSN) [mounted with a specimen of rubiginosus Newman

on same card, the latter marked “r”]; 14, 12, Adelaide Coll. Castelnau, det. aphodioides (MCSN) [mounted with

a paratype of s. sinuaticollis, spec. nov. on same card, the latter marked “s”]; 14, Adel.“ Tarrier, aphodioides ?, Soc.

Ent. Belg. Coll. PUTZEYS, Adelotopus aphodioides Westw. det. J. Putzeys (IRSNB); 222, Austral. Yorketown,

CNHM Karl Brancsik ex Eduard Knirsch, Pseudomorphini Genus ? sp ? det. D. Shpeley 1987 (FMNH); 14, Nat.

Mus. Victoria S. Australia, Nat. Mus. Victoria C. French’s Coll. 5.11.08, Adelotopus aphodioides Westw. S. Australia

(NMV); 15, The Gorge H. S. Davis 24.10.98 (SAMA); 522, 2589, S. Austr., det. aphodioides (AMNH); 13, K 12349,

Adelotopus hydrobioides Westw. S. Aust. (AMS); 14, 12, Nat. Mus. Victoria -S. Austr. (NMV); 12, S. Austral.,

W (OUM); 12,5. Austr., 2559, Ex Musaeo H. W. Bates 1892 (MNHN); 288,19, 5. Austr. (MMS); 338, aphodioides

Westwood, Melbourne Bakewell, Ex Musaeo Chaudoir (MNHN); 1%, Kiata, V. 31.10.50 F.E.W. ?, det. castaneus

(NMV); 14, Deniliquin 21.1.55 NSW F. E. Wilson, F. E. Wilson Collewction (NMV); 13, Dubbo, N.S.W. 28. Rev.

A. J. Barrett, K 60444, A. castaneus (AMS); 13, 1%, NSW (OUM); 538, 522, Longreach Q.: A. M. Lea, Ditto

Queensland, det. castaneus (SAMA); 17, Longreach Q.: A. M. Lea, ac. 23246, aphodioides Westw. (AMNH);

13, Longreach Q.: A. M. Lea, ac. 23246, Adelotopus castaneus Cast. Id. by A.M. Lea (AMNH); 15, Longreach Q.:

A.M. Lea, ac. 23246, det. aphodioides (AMNH); 13, Longreach Q., A. M. Lea, ac. 23246, det. castaneus (AMNH);

13, Winton Q., Lea, castaneus Cast. Id. by A.M. Lea, F. E. Wilson Collection (NMV); 15, Charters T. 300 m IX.66,

J. H. Sedlacek Collector (CBM); 1%, Australia: N. Queensland 1917/1918, Hughenden N. Queensl. F.M. 11.1.1918

(NHMW); 62%, Alice Springs, C.A. Dec. 1896 (CBM, NMV); 22%, Daly Waters N. T., J. Sedlacek Collector (CSB);

13, A. hydrobioides Westw. Bundogendra (?) (ANIC); 15, Wellington Froggatt 1891, W. W. Froggatt Collection

(ANIC); 12, Austral. mer., Sharp Coll. 1905-313., Adelotopus sp. Voisin du distinctus, det. rubiginosus (BMNH);

12, Australien, Coll. E. Witte, castaneus, Adelotopus Castaneus Cast. Aust. (SMFC 16272); 12, Australia, Adelotopus

distinctus Chaud. K. K., Coll. Kraatz, A. distinctus Chaud. (DEIB); 14, Adelotopus castaneus Cast. Austral.,

Adelotopus aphodioides Westw. (MCZ); 17, Australia, Ex Musaeo E. Steinheil (MNHN); 12, Australie, Dohrn,

Adelotopus castaneus Cast. det..., Adelotopus castaneus Cast. Austral. (IRSNB); 15, rufescens Chaud., Ex Musaeo

Chaudoir (MNHN); 1%, Putzeys, Fundort ?, Coll. E. Witte, Adelotopus? (SMF C 16274); 1?, Coll. B. Schwarzer,

det. castaneus (SMF C 16273).

Note. This species has been repeatedly identified as A. aphodioides Westwood. Because the types of

A. aphodioides are not available and presumably lost, it is impossible to verify this determination at

present (see under doubtful species).

Diagnosis. Medium-sized, completely reddish species, distinguished by wide, moderately convex

pronotum, rather dense, moderately fine puncturation of head and pronotum, symmetric d gential

ring, moderately widened, on lower surface not striate aedeagus with shortly rounded apex, apically

angulate left paramere, and wide, apically strongly narrowed stylomere.

Description

Measurements. Length: 4.7-6.1 mm. Ratios. Width/length of pronotum: 1.58-1.70; width base/

apex of pronotum: 1.40-1.51; width pronotum/ head: 1.54-1.65; length/ width of elytra: 1.47-1.51; length

elytra/pronotum: 2.35-2.52.

Colour. Upper and lower surface including mouth parts, antennae, and legs reddish, forebody

faintly darker.

Head (Figs 173a-d). Rather short, fairly wide, rather depressed. Anterior border gently convex,

lateral angle rounded, laterally faintly projecting, lateral borders slightly narrowed behind eyes.

Clypeal suture only at base distinct, in middle widely interrupted. Labrum rather wide and short,

moderately overlapped by the clypeus, apex moderately concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly convex area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa fairly wide, tongue-like, apically

convex, ventrally with distinct keel, at border with c. 10-12 elongate setae. Terminal palpomere of

maxillary palpus moderately widened, faintly securiform. Terminal palpomere of labial palpus very

218

Figs 173a-l. Adelotopus laticollis, spec. nov. Details of head and genitalia. For legends see fig. 100.

wide, markedly securiform. Antenna rather elongate, 8th-9th antennomeres c. 1.5 x as wide as long.

Microreticulation fine, superficial, puncturation very fine, fairly dense. Surface with weak sulcus

medially of eyes, usually with some irregular, fine wrinkles, impilose, fairly glossy. Ventrolaterally of

eyes with a row of short setae. Suborbital field punctate and shortly setose. Gula impilose.

Pronotum (Fig. 361). Wide, rather convex, base distinctly wider than apex, widest near base.

Apical angles moderately produced, at apex obtusely rounded, fairly oblique, surpassing posterior

border of eyes. Apex fairly excised, markedly convex in excision, feebly bordered. Sides gently convex,

rather oblique. Margins rather wide, rather chanelled, finely bordered. Basal angles widely rounded

off. Base straight or faintly concave, rather irregularly bordered. Surface near base without transverse

impression. Microreticulation much reduced, highly superficial, sometimes only faint traces visible,

puncturation moderately fine to fairly coarse, dense, surface with more or less distinct rugosities,

impilose, highly glossy.

Elytra (Figs 361, 512). Rather wide, moderately convex, slightly depressed on disk, rather parallel,

though usually faintly narrowed in basal third. Lateral borders almost straight. Apex wide, slightly

oblique, truncature barely convex, apical angles widely rounded off. Shoulders rounded, basal margin

slightly oblique, without setae behind shoulders. Marginal channel moderately wide, partly concealed.

Basal border incomplete, attaining outer third of base. Lateral margin asetose. Series of umbilical pores

consisting of 6 rather spaced pores behind shoulder. Setae fairly elongate. Striae including sutural stria

absent. Microreticulation absent, puncturation moderately fine to fairly coarse, fairly dense, surface

impilose, markedly glossy.

Lower surface. Prosternal process rather short, narrow, convex, apex very short, narrow, com-

pressed, passing over in an almost right angle from ventral surface, shortly setose. Metepisternum

elongate, c. 1.8 x as long as wide, in posterior third not hollowed. Abdominal sterna with 1 elongate

seta each side. Lower surface rather densely punctate and shortly setose.

Legs. Elongate, 1st tarsomere of protarsus slightly longer than wide, tibial groove of profemur

moderately deep, anterior plate overlapping the groove for about apical third, posterior border of

groove sharp. Femur wide. Metatibia elongate, c. 6.5 x as long as wide, Ist tarsomere of metatarsus

almost 2.5 x as long as wide.

d genitalia (Figs 173e-k). Genital ring rather wide, convex, barely asymmetric, with rather short

2.19

apex, with slightly asymmetric, rather small, fairly excised base. Sternum VII rather wide, apically

evenly convex, with moderately deep excision, base gently bisinuate, basal angles obtusely rounded,

lateral parts fairly elongate. Aedeagus rather short, fairly depressed, in middle rather widened,

asymmetric, left side convex, right side almost straight. Basal part rather long, moderately bent. Lower

surface straight to gently concave, not perceptibly striped. Apex moderately narrow, evenly rounded

off, rather symmetric. Orifice rather elongate, internal sac complex, with a distinct oblique fold near

apex. Both parameres large, rather elongate, right square, with widely rounded apex, left paramere

considerably larger than right, triangular, apex angulate, shortly rounded.

? genitalia (Fig. 1731). Stylomere wide, strongly narrowed to apex, apex rounded off, both lateral

and median margins concave, with 2-3 elongate apical setae. Lateral plate elongate, with 3-5 elongate

and 2-3 short apical setae.

Variation. Very variable species with respect to size, shape and relative width of pronotum, size

and density of puncturation, rugosity of pronotum, and shape of apex of stylomere.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Virtually unknown. The very few dated specimens collected in January, February, September,

October, and December. Only very few recent captures recorded.

Distribution (Fig. 623). Eastern South Australia, western Victoria, western New South Wales, western

Queensland, central Northern Territory. This is virtually an inland species.

Material examined (77). Apart from the the type series one damaged specimen of unknown sex from

“Australia: N. Queensld. Richmond 1917/1918” (NHMW) is not included in the type series.

Etymology. The name refers to the wide pronotum.

Adelotopus cribricollis, spec. nov.

Figs 174, 362, 513, 624

Types. Holotype: d, Mt. Spec, N.Q. 1/68. G.B., M-114., J. G. Brooks Bequest, 1976 (ANIC) [holotype mounted

with a paratype on same card, marked HT.]. - Paratypes: 15, same data (ANIC); 15, Australia: N.S.W. Trangie

6.X11.1965. ©. W. Richards B.M. 1966-325 (BMNH); 28, Morilla N.S.W. 20.1.27 (ANIC); 238, 222, Brisbane

Nlidge (UOIC); 2355, Toowoomba Brisbane distr. (MNHN); 15, Australia: Q’d Rockhampton 26.X11.1967, J. &

M. Sedlacek Collectors Bishop (BMH); 434,7%%, Australia: Qld Rockhampton, 26-27.X1.1967, J. & M. Sedlacek

Collectors BISHOP (BMH, CBM); 25d, Rock (CSB); 13, Australien, Qld 26 Mackenzie R., 79km n. Dingo,

11.-12.11.1990, M. Baehr (CBM); 13, 1755 4.328, Bowen Queensland A. Simson, Adelotopus castaneus Cast. By

Simson’s number (SAMA); 24d, 12, Port Denison Coll. Castelnau, det. aphodioides (MCSN); 15, 27%, Port

Denison, Port Denison Coll. Castelnau, det. aphodioides (MCSN) Imounted with four specimens of rubiginosus

Newman on same needle, upper card = cribricollis]; 12, rufescens Chaud., P. Denison, aphodioides Westw.,

Ex Musaeo Mniszech (MNHN); 24d, Australia: Old Charters Towers 196, J. &. M. Sedlacek Collectors BISHOP,

22-20.X1.1967 (BMH); 336, Australia: W. Paluma NQ 13.1.70 J. G. Brooks, det. aphodioides (CMC); 18, Archers Ck.

NO 1/55 GB., M. 186, castaneus Cast. 1577. J. G. Brooks Bequest, 1976 (ANIC); 15, Archers Ck. NO 11/54 GB,

M. 280, J. G. Brooks Bequest, 1976 (ANIC); 15, Watten Queensland F. H. Taylor (ANIC); 15, E.Q. (ANIC).

Diagnosis. Small to medium-sized, completely reddish species, distinguished by wide, moderately

convex pronotum with comparatively wide apex, rather dense, usually rather coarse and rugose

puncturation on pronotum, rather symmetric d genital ring, short and moderately widened, on lower

surface not striate aedeagus with shortly rounded apex and a distinct keel on upper surface, apically

obliquely convex, strongly striate left paramere, and wide, apically strongly narrowed stylomere.

Description

Measurements. Length: 4.0-5.5 mm. Ratios. Width/length of pronotum: 1.52-1.60; width base/

apex of pronotum: 1.36-1.41; width pronotum/head: 1.49-1.54; length/ width of elytra: 1.49-1.57; length

elytra/pronotum: 2.39-2.47.

Colour. Upper and lower surface including mouth parts, antennae, and legs reddish, forebody

sometimes faintly darker.

Head (Figs 174a-d). Rather short, fairly wide, rather depressed. Anterior border gently convex,

lateral angle rounded, laterally faintly projecting, lateral borders slightly narrowed behind eyes.

Figs 174a-l. Adelotopus cribricollis, spec. nov. Details of head and genitalia. For legends see fig. 100.

Clypeal suture only at base distinct, in middle widely interrupted. Labrum rather wide and short,

moderately overlapped by the clypeus, apex moderately concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly convex area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa fairly wide, tongue-like, apically

convex, ventrally with distinct keel, at border with c. 10-12 elongate setae. Terminal palpomere of

maxillary palpus moderately widened, slightly securiform. Terminal palpomere of labial palpus very

wide, markedly securiform. Antenna rather narrow and elongate, Sth-9th antennomeres c. 1.5 x as

wide as long. Microreticulation fine, superficial, puncturation fine, fairly dense. Surface with weak

sulcus medially of eyes, impilose, rather glossy. Ventrolaterally of eyes with a row of short setae.

Suborbital field punctate and shortly setose. Gula impilose.

Pronotum (Fig. 362). Rather wide, rather convex, base distinctly wider than apex, widest near

base. Apical angles moderately produced, at apex obtuse, fairly oblique, surpassing posterior border

of eyes. Apex fairly excised, markedly convex in excision, feebly bordered. Sides gently convex,

moderately oblique. Margins rather wide, moderately chanelled, finely bordered. Basal angles widely

rounded off. Base straight or faintly concave, rather irregularly bordered. Surface near base without

or with very shallow transverse impression. Microreticulation much reduced, highly superficial,

sometimes only faint traces visible, puncturation moderately fine to fairly coarse, especially laterally,

dense, surface ususally with rather corase wrinkles, impilose, glossy.

Elytra (Figs 362, 513). Rather wide, moderately convex, slightly depressed on disk, rather parallel.

Lateral borders almost straight. Apex wide, slightly oblique, truncature barely convex, apical angles

widely rounded off. Shoulders rounded, basal margin slightly oblique, without setae behind shoul-

ders. Marginal channel moderately wide, partly concealed. Basal border incomplete, attaining outer

third of base. Lateral margin asetose. Series of umbilical pores consisting of 6, rarely unilaterally 5 or

7 rather spaced pores behind shoulder. Setae fairly elongate. Striae including sutural stria absent.

Microreticulation absent, puncturation moderately fine to fairly coarse, fairly dense, surface impilose,

markedly glossy.

Lower surface. Prosternal process rather short, narrow, convex, apex very short, narrow, com-

pressed, passing over in an almost right angle from ventral surface, shortly setose. Metepisternum

elongate, c. 1.9 x as long as wide, in posterior third not hollowed. Abdominal sterna with 1 elongate

seta each side. Lower surface rather densely punctate and shortly setose.

Legs. Rather elongate, Ist tarsomere of protarsus as long or slightly longer than wide, tibial groove

of profemur moderately deep, anterior plate overlapping the groove for about apical third, posterior

border of groove sharp. Femur wide. Metatibia fairly elongate to elongate, c. 5.5-6.5 x as long as wide,

lst tarsomere of metatarsus c. 2.2-2.5 x as long as wide.

d genitalia (Figs 174e-k). Genital ring rather wide, convex, slightly asymmetric, with rather short

apex, with distincly asymmetric, rather small, fairly excised base. Sternum VII wide, apically evenly

convex, with moderately deep excision, base gently bisinuate, in middle markedly convex, basal angles

obtusely rounded, lateral parts fairly elongate. Aedeagus short, fairly depressed, in middle strongly

widened, slightly asymmetric. Basal part rather long, moderately bent. Lower surface straight, not

perceptibly striped. Upper surface about in middle with a distinct keel. Apex rather narrow, evenly

rounded off, rather symmetric. Orifice rather elongate, internal sac complex, with a distinct oblique

fold near apex. Both parameres large, rather elongate, right slightly triangular, with widely rounded

apex, left paramere considerably larger than right, square, apex conspicuously oblique.

? genitalia (Fig. 1741). Stylomere wide, more or less strongly narrowed to apex, apex rounded off,

both lateral and median margins concave, with 2-3 elongate apical setae. Lateral plate elongate, with

2-6 elongate apical setae.

Variation. There is some variation of size, relative width of pronotum, size and desnsity of

puncturation, rugosity of pronotum, length of legs and shape of stylomere.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Largely unknown. A specimen collected by me under bark of river gum. Dated specimens

captured from November to January.

Distribution (Fig. 624). New South Wales, eastern Queensland.

Material examined (44). Apart from the type series a single, very small specimen is tentatively assigned to this

species, because it has comparatively short legs and slightly differently shaped genital ring and aedeagus:

1, Australien, Coll. E. Witte (SMF).

Etymology. The name refers to the rugose surface of pronotum.

Adelotopus luteus, spec. nov.

Figs 175, 363, 514, 624

Types. Holotype: 4, 15.395 144.31E Split Rock QLD 18 Aug-16 Sep 1993 Flight Intercept Trap P. Zborowski &

S. Shattuck (QMB T26060). — Paratypes: 17, same data (DPIM); 14, same locality, 18 Feb-25 Apr 1993 Malaise

Trap P. Zborowski (DPIM); 1%, same locality, 10 Oct-18 Nov 1993 Malaise Trap P. Zborowski & M. Horak (CBM).

Diagnosis. Rather small, completely light reddish species, distinguished by the very wide, rather

depressed pronotum with markedly wide, channeled lateral margins, very fine puncturation on

pronotum and elytra, rather symmetric d genital ring, short and moderately widened, symmetric, on

lower surface not striate aedeagus with shortly rounded apex, and wide, markedly triangular

stylomere. Further distinguished from most closely A. houstoni, spec. nov. by basally wider pronotum,

distinctly shorter and wider antenna, even finer puncturation of surface, and shorter and basally much

more concave lateral plate of ? stylomere.

Description

Measurements. Length: 4.85-4.95 mm. Ratios. Width/length of pronotum: 1.73-1.75; width base/

apex of pronotum: 1.52-1.56; width pronotum/head: 1.67-1.70; length/width of elytra: c. 1.45; length

elytra/pronotum: 2.45-2.52.

Colour. Upper and lower surface including mouth parts, antennae, and legs yellowish to light

reddish, forebody faintly darker, tibiae and tarsi reddish-piceous.

Head (Figs 175a-d). Rather short, fairly wide, rather depressed. Anterior border gently convex,

lateral angle rounded, laterally faintly projecting, lateral borders slightly narrowed behind eyes.

Clypeal suture only at base distinct, in middle widely interrupted. Labrum rather wide and short,

99)

Figs 175a-l. Adelotopus luteus, spec. nov. Details of head and genitalia. For legends see fig. 100.

moderately overlapped by the clypeus, apex moderately concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly convex area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa fairly wide, tongue-like, apically

convex, ventrally with distinct keel, at border with c. 10-12 elongate setae. Terminal palpomere of

maxillary palpus moderately widened, slightly securiform. Terminal palpomere of labial palpus very

wide, markedly securiform. Antenna short and apically distinctly widened, Sth-9th antennomeres >2 x

as wide as long. Microreticulation very fine, superficial, puncturation extremely fine, difficult to see,

rather sparse. Surface with weak sulcus medially of eyes, impilose, rather glossy. Ventrolaterally of

eyes with a row of short setae. Suborbital field punctate and shortly setose. Gula impilose.

Pronotum (Fig. 363). Very wide, rather depressed, base distinctly wider than apex, widest in apical

third, strongly narrowed to apex. Apical angles moderately produced, at apex obtuse, fairly oblique,

surpassing posterior border of eyes. Apex fairly excised, markedly convex in excision, bordered. Sides

regularly convex, moderately oblique, very wide, markedly chanelled, almost explanate, finely

bordered. Basal angles very widely rounded off. Base faintly convex, distinctly bordered. Surface near

base with very weak transverse impression. Microreticulation almost wanting, sometimes faintest

traces visible, puncturation very fine, sparse, surface impilose, glossy.

Elytra (Figs 363, 514). Rather wide, fairly depressed, rather parallel. Lateral borders almost

straight. Apex wide, slightly oblique, truncature faintly convex, apical angles widely rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

moderately wide, partly concealed. Basal border incomplete, attaining outer third of base. Lateral

margin asetose. Series of umbilical pores consisting of 6, rarely unilaterally 5 rather spaced pores

behind shoulder. Setae short. Striae including sutural stria absent. Microreticulation absent, punctu-

ration very fine, sparse, surface impilose, markedly glossy.

Lower surface. Prosternal process rather short, narrow, convex, apex very short, narrow, com-

pressed, passing over in an almost right angle from ventral surface, shortly setose. Metepisternum

rather elongate, c. 1.7 x as long as wide, in posterior third not hollowed. Abdominal sterna without

elongate setae. Lower surface rather densely punctate and shortly setose.

Legs. Rather elongate, Ist tarsomere of protarsus about as long as wide, tibial groove of profemur

shallow, anterior plate overlapping the groove only in apical fourth, border almost straight, posterior

border of groove sharp. Femur moderately wide. Metatibia fairly elongate, c. 6 x as long as wide, Ist

tarsomere of metatarsus c. 2.2 x as long as wide.

[59]

159)

[09]

d genitalia (Figs 175e-k). Genital ring moderately wide, convex, slightly asymmetric, with short,

obtuse apex, with asymmetric, rather small, little excised base. Sternum VII fairly wide, apically faintly

oblique, with moderately deep excision, base almost straight, basal angles obtusely rounded, lateral

parts rather short. Aedeagus short, fairly depressed, in middle rather widened, slightly asymmetric.

Basal part rather long, moderately bent. Lower surface almost straight, not perceptibly striped. Apex

narrow, obtusely angulate. Orifice elongate, internal sac complex, with a distinct oblique fold near

apex. Both parameres large, rather elongate, with narrowly rounded apex, left paramere considerably

larger than right.

? genitalia (Fig. 1751). Stylomere wide, triangular, apex rounded off, with 3-4 elongate apical setae.

Lateral plate rather short, with 3 rather short apical setae.

Variation. Very little variation noted.

Vivipary. Not confirmed in the examined material.

Habits. Largely unknown. All known specimens collected in “Flight intercept trap” and “Malaise

Trap” during the periods of February-April, August-September, and October-November.

Distribution (Fig. 624). Base of Cape York Peninsula, northern Queensland. Known only from type

locality.

Material examined (4). Only the type series.

Etymology. The name refers to the light colour.

Adelotopus houstoni, spec. nov.

Figs 176, 364, 515, 625

Types. Holotype: 2, 11 km SSE of Banjiwarn HS (27°42°S 121°37’E) W. Aust. 22-28 Feb 1980 T. F. Houston et al.

316-18; Ex pitfall trap Western Australian Museum, Dept. of Biological Survey Site BWRI (WAM).

Diagnosis. Rather small, completely light reddish species, distinguished by the very wide, rather

depressed pronotum with markedly wide, channeled lateral margins, fine puncturation on pronotum

and elytra, and wide, markedly triangular stylomere. Further distinguished from most closely

A. Iuteus, spec. nov. by basally narrower pronotum, distinctly longer and narrower antenna, slightly

coarser puncturation of surface, and longer and basally far less concave lateral plate of ? stylomere.

Description

Measurements. Length: 4.65 mm. Ratios. Width/length of pronotum: 1.75; width base/apex of

pronotum: 1.42; width pronotum/head: 1.54; length/width of elytra: 1.48; length elytra/pronotum:

2.50.

Colour. Upper and lower surface including mouth parts, antennae, and legs yellowish to light

reddish, forebody faintly darker.

Head (Figs 176a-d). Rather short, fairly wide, rather depressed. Anterior border gently convex,

lateral angle rounded, laterally faintly projecting, lateral borders slightly narrowed behind eyes.

Clypeal suture only at base distinct, in middle widely interrupted. Labrum rather wide and short,

moderately overlapped by the clypeus, apex moderately concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly convex area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa fairly wide, tongue-like, apically

convex, ventrally with distinct keel, at border with c. 10-12 elongate setae. Maxillary palpus lost.

Terminal palpomere of labial palpus very wide, markedly securiform. Antenna rather narrow and

elongate, barely widened, 8th-9th antennomeres c. 1,5 x as wide as long. Microreticulation very fine,

superficial, puncturation fine though distinct, moderately dense. Surface with weak sulcus medially

of eyes, impilose, moderately glossy. Ventrolaterally of eyes with a row of short setae. Suborbital field

punctate and shortly setose. Gula impilose.

Pronotum (Fig. 364). Very wide, rather depressed, base distinctly wider than apex, widest in apical

third, moderately narrowed to apex. Apical angles moderately produced, at apex obtuse, fairly

oblique, surpassing posterior border of eyes. Apex fairly excised, markedly convex in excision,

bordered. Sides regularly convex, moderately oblique, wide, chanelled, finely bordered. Basal angles

widely rounded off. Base faintly convex, distinctly bordered. Surface near base with very weak

Figs 176a, c-d, I. Adelotopus houstoni, spec. nov. Details of head and % genitalia. For legends see fig. 100.

transverse impression. Microreticulation absent, puncturation moderately fine, rather sparse, surface

impilose, glossy.

Elytra (Figs 364, 515). Rather wide, fairly depressed, rather parallel. Lateral borders almost

straight. Apex wide, slightly oblique, truncature faintly convex, apical angles widely rounded off.

Shoulders rounded, basal margin slightly oblique, without setae behind shoulders. Marginal channel

moderately wide, partly concealed. Basal border incomplete, attaining outer two fifth of base. Lateral

margin asetose. Series of umbilical pores consisting of 6 rather spaced pores behind shoulder. Setae

short. Striae including sutural stria absent. Microreticulation absent, puncturation moderately fine,

fairly sparse, surface impilose, markedly glossy.

Lower surface. Prosternal process rather short, moderately narrow, convex, apex very short,

narrow, compressed, passing over in an almost right angle from ventral surface, shortly setose.

Metepisternum rather elongate, c. 1.7 x as long as wide, in posterior third not hollowed. Abdominal

sterna without elongate setae. Lower surface rather densely punctate and shortly setose.

Legs. Elongate, Ist tarsomere of protarsus about as long as wide, tibial groove of profemur

shallow, anterior plate barely overlapping the groove, posterior border of groove sharp. Femur

moderately wide. Metatibia elongate, c.7 x as long as wide, 1st tarsomere of metatarsus c. 2.5 x aslong

as wide.

d genitalia. Unknown.

? genitalia (Fig. 1761). Stylomere wide, triangular, apex widely rounded off, with 2 elongate apical

setae. Lateral plate moderately elongate, with 2 elongate apical setae.

Variation. Unknown.

Vivipary. Not confirmed in the examined materal.

Habits. Largely unknown. Holotype collected “in pitfall trap” in February.

Distribution (Fig. 625). Central Western Australia. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. Named in honour of the collector of the holotype.

Adelotopus virgatus, spec. nov.

Eies 91, 177, 365, 516, 625

Types. Holotype: d, Rushworth, V. J. C. le Souef, F. E. Wilson Collection, det. aphodioides (NMV). —- Paratypes:

1?, Adelaide Recu de W. Bates, aphodioides Westw. (MNHN); 16, Adelaide J. G. ©. Tepper (SAMA);

13, Caldwell, N.S.W. 1.31 G. Goudie, J. C. Goudie Collection (NMV); 12, Morilla N.S.W. T.G.S. 31.12.26 (ANIC);

14, Australia: Lake Benanee N.S.W. 14.X11.72 B. P. Moore, Adelotopus gyrinoides Hope det. B. P. Moore 87 (CMC);

18, 322, Australia: Lake Benanee N.S.W. 14.X11.72 B. P. Moore, det. gyrinoides (CBM, CMC); 18, Australia,

middle NSW Round Hill: near Lake Cargillego, No. 67, 11.11.1981 leg. Hangay & Vojnik (HNMB); 14, West.

Riverina Australia Wait leg. Lüddemann ded. (DEIB); 734, 19, Australia: N. T.: Alice Springs 20-24.111.1971,

J. &M. Sedlacek Colls. BISHOP MUSEUM (BMH, CBM); 23, Australia: N. T.: Alice Springs 350-650 m

20-24.111.1968, N. L. H. Krauss Coll. BISHOP MUSEUM (BMH),; 1%, Simson, Nov. Holl. Queensl‘., Fry Coll.

1905.100, det rubiginosus (BMNH); 15, Pascoe Coll. 93-60 (BMNH).

Diagnosis. Small to medium-sized, completely reddish species, distinguished by wide, moderately

convex pronotum with moderately wide apex, rather dense, usually rather coarse puncturation on

pronotum, symmetric d genital ring, short and wide, asymmetric, on lower surface strongly striate

aedeagus with obtuse apex, apically obliquely convex, strongly striate left paramere, and rather

narrow, apically narrowed stylomere.

Description

Measurements. Length: 4.9-5.7 mm. Ratios. Width/length of pronotum: 1.62-1.68; width base/

apex of pronotum: 1.36-1.48; width pronotum/head: 1.47-1.58; length/ width of elytra: 1.46-1.58; length

elytra/pronotum: 2.42-2.58.

Colour. Upper and lower surface including mouth parts, antennae, and legs reddish, forebody

sometimes faintly darker.

Head (Figs 177a-d). Rather short, fairly wide, rather depressed. Anterior border gently convex,

lateral angle rounded, laterally faintly projecting, lateral borders slightly narrowed behind eyes.

Clypeal suture only at base distinct, in middle widely interrupted. Labrum rather wide and short,

moderately overlapped by the clypeus, apex moderately concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly convex area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa fairly wide, tongue-like, apically

convex, ventrally with distinct keel, at border with c. 10-12 elongate setae. Terminal palpomere of

maxillary palpus moderately widened, fairly securiform. Terminal palpomere of labial palpus very

wide, markedly securiform. Antenna rather narrow and elongate, 8th-9th antennomeres c. 1.5 x as

wide as long. Microreticulation fine, superficial, puncturation fine, fairly dense. Surface with weak

sulcus medially of eyes, impilose, rather glossy. Ventrolaterally of eyes with a row of short setae.

Suborbital field punctate and shortly setose. Gula impilose.

Pronotum (Fig. 365). Rather wide, rather convex, base distinctly wider than apex, widest near

base. Apical angles moderately produced, at apex obtuse, fairly oblique, surpassing posterior border

of eyes. Apex fairly excised, markedly convex in excision, feebly bordered. Sides gently convex,

moderately oblique. Margins rather wide, moderately chanelled, finely bordered. Basal angles widely

rounded off. Base straight or faintly concave, rather irregularly bordered. Surface near base with

shallow transverse impression. Microreticulation much reduced, highly superficial, sometimes only

faint traces visible, puncturation fairly coarse, dense, surface rarely somewhat rugose, impilose, glossy.

Elytra (Figs 51, 365, 516). Rather wide, moderately convex, slightly depressed on disk, rather

parallel. Lateral borders usually slightly convex. Apex wide, slightly oblique, truncature barely

convex, apical angles widely rounded off. Shoulders rounded, basal margin slightly oblique, without

setae behind shoulders. Marginal channel moderately wide, partly concealed. Basal border incom-

plete, attaining outer third of base. Lateral margin asetose. Series of umbilical pores consisting of 6,

rarely unilaterally of 5 rather spaced pores behind shoulder. Setae fairly elongate. Striae including

sutural stria absent. Microreticulation absent, puncturation moderately to fairly coarse, fairly dense,

surface impilose, markedly glossy.

Lower surface. Prosternal process rather short, narrow, convex, apex very short, narrow, com-

pressed, passing over in an almost right angle from ventral surface, shortly setose. Metepisternum

elongate, c. 1.9 x as long as wide, in posterior third not hollowed. Abdominal sterna with 1 elongate

seta each side. Lower surface rather densely punctate and shortly setose.

Legs. Rather elongate, 1st tarsomere of protarsus as long or slightly longer than wide, tibial groove

of profemur moderately deep, anterior plate overlapping the groove for about apical third, posterior

border of groove sharp. Femur wide. Metatibia rather elongate, c. 5.5-6 x as long as wide, Ist tarsomere

of metatarsus c. 2.3 x as long as wide.

d genitalia (Figs 177e-k). Genital ring rather wide, rather triangular, barely asymmetric, with

rather short apex, with slightly asymmetric, rather small, fairly excised base. Sternum VII wide,

apically evenly convex, with moderately deep excision, base gently bisinuate, in middle markedly

convex, basal angles obtusely rounded, lateral parts elongate. Aedeagus short, depressed, in middle

strongly widened, markedly asymmetric, left side strongly convex, right almost straight. Basal part

Figs 177a-m. Adelotopus virgatus, spec. nov. Details of head and genitalia. For legends see fig. 100, m. ?

stylomeres and lateral plate of untypical specimen.

rather long, markedly bent. Lower surface straight, markedly striped. Apex narrow, obtuse, asymmet-

ric. Orifice rather elongate, internal sac complex, with a large oblique fold near apex. Both parameres

rather elongate, right rather elongate, with widely rounded apex, left paramere very large, triangular,

apex obliquely rounded.

? genitalia (Figs 1771,m). Stylomere moderately narrow, more or less strongly narrowed to apex,

apex rounded off, both lateral and median margins concave, with 2-4 elongate apical setae. Lateral

plate elongate, with 2-3 elongate apical setae.

Variation. Some variation in shape of pronotum, density and degree of rugosity of pronotal

puncturation, and shape of stylomere noted.

Vivipary. Confirmed by discovery of larvae in the ? oviducts.

Habits. Unknown. Dated specimens captured from December to March.

Distribution (Fig. 625). South Australia, Victoria, western New South Wales, central Northern

Territory.

Material examined (27). In addition to the type series there are some ? specimens doubtfully assigned to this

species, mainly because of differences in the form of their stylomere (Fig. 177m): 1%, Fundort ?, Coll.

B. Schwarzer, det. brunneus (SMF); 1%, Collect. Plason, N. H. (NHMW); 1%, rufescens Chaud., Ex Musaeo

Chaudoir (MNHN).

Etymology. The name refers to the strongly striped lower surface of the aedeagus.

159)

Adelotopus brittoni, spec. nov.

Figs 178, 366, 517, 626

Types. Holotype: d, Wigley Waterhole, 5 miles N. of Alice Springs, NT. 16 Feb 1966 Britton, Upton & Melnnes.

Adelotopus castaneus Cast. E. B. Britton det. 1967 (ANIC).

Diagnosis. Medium-sized, completely reddish species, distinguished by wide, moderately convex

pronotum with moderately wide apex, fine, rather sparse puncturation on pronotum and elytra,

symmetric d genital ring, short and wide, almost symmetric, on lower surface strongly striate

aedeagus with acute apex, and apically convex, strongly striate left paramere.

Description

Measurements. Length: 5.8 mm. Ratios. Width/length of pronotum: 1.65; width base/apex of

pronotum: 1.44; width pronotum/head: 1.65; length/width of elytra: 1.45; length elytra/pronotum:

2.43.

Colour. Upper and lower surface including mouth parts, antennae, and legs reddish.

Head (Figs 178a-d). Rather short, fairly wide, rather depressed. Anterior border gently convex,

lateral angle rounded, laterally faintly projecting, lateral borders slightly narrowed behind eyes.

Clypeal suture only at base distinct, in middle widely interrupted. Labrum rather wide and short,

moderately overlapped by the clypeus, apex moderately concave. Antennal groove laterally sharply

bordered, latero-posteriorly with slightly convex area. Mental tooth triangular, rather short, apex

acute. Wings of mentum wide, laterally rounded, apex obtuse. Glossa fairly wide, tongue-like, apically

convex, ventrally with distinct keel, at border with c. 10-12 elongate setae. Terminal palpomere of

maxillary palpus moderately widened, fairly securiform. Terminal palpomere of labial palpus very

wide, markedly securiform. Antenna rather narrow and elongate, Sth-9th antennomeres <1.5 x as wide

as long. Microreticulation fine, superficial, puncturation very fine, fairly dense. Surface with weak

sulcus medially of eyes, impilose, rather glossy. Ventrolaterally of eyes with a row of short setae.

Suborbital field punctate and shortly setose. Gula impilose.

Pronotum (Fig. 366). Rather wide, rather convex, base distinctly wider than apex, widest near

base. Apical angles moderately produced, at apex acute, fairly oblique, surpassing posterior border of

eyes. Apex fairly excised, markedly convex in excision, feebly bordered. Sides gently convex,

moderately oblique. Margins rather wide, moderately chanelled, finely bordered. Basal angles widely

rounded off. Base straight or faintly concave, rather distinctly bordered. Surface near base with

shallow transverse impression. Microreticulation much reduced, highly superficial, puncturation fine,

rather sparse, surface impilose, highly glossy.

Elytra (Figs 366, 517). Rather wide, moderately convex, slightly depressed on disk, parallel. Lateral

borders even faintly narrowed in basal half. Apex wide, slightly oblique, truncature faintrly convex,

apical angles widely rounded off. Shoulders rounded, basal margin slightly oblique, without setae

behind shoulders. Marginal channel moderately wide, partly concealed. Basal border incomplete,

attaining about middle of base. Lateral margin asetose. Series of umbilical pores consisting of 6 rather

spaced pores behind shoulder. Setae fairly elongate. Striae including sutural stria absent. Microretic-

ulation absent, puncturation rather fine, moderately sparse, surface impilose, very glossy.

Lower surface. Prosternal process rather short, narrow, convex, apex very short, narrow, com-

pressed, passing over in an almost right angle from ventral surface, shortly setose. Metepisternum

elongate, c. 1.9 x as long as wide, in posterior third not hollowed. Abdominal sterna with 1 elongate

seta each side. Lower surface rather densely punctate and shortly setose.

Legs. Elongate, Ist tarsomere of protarsus slightly longer than wide, tibial groove of profemur

moderately deep, anterior plate overlapping the groove for about apical third, posterior border of

groove sharp. Femur wide. Metatibia elongate, c. 6.5 x as long as wide, 1st tarsomere of metatarsus

>2.5 x as long as wide.

ö genitalia (Figs 178e-k). Genital ring rather wide, slightly convex, barely asymmetric, with rather

short apex, with slightly asymmetric, rather small, fairly excised base. Sternum VII very wide, apically

evenly convex, with moderately deep excision, base faintly bisinuate, in middle markedly convex,

basal angles obtusely rounded, lateral parts elongate. Aedeagus short, depressed, in middle rather

widened, slightly asymmetric. Basal part rather long, markedly bent. Lower surface straight to gently

convex, slightly flattened near apex, markedly striped. Lateral border narrow. Apex narrow, acute,

228

Figs 178a-k. Adelotopus brittoni, spec. nov. Details of head and d genitalia. For legends see fig. 100.

slightly asymmetric. Orifice rather elongate, internal sac complex, with a large oblique fold near apex.

Both parameres rather wide and triangular, at apex widely rounded off, left paramere considerably

larger.

? genitalia. Unknown.

Variation. Unknown.

Vivipary. Not confirmed due to lack of females.

Habits. Unknown. Holotype collected in February.

Distribution (Fig. 626). Central Northern Territory. Known only from type locality.

Material examined (1). Only the holotype.

Etymology. Named in honour of the collector and famous coleopteran specialist E. B. Britton.

Adelotopus adustus, spec. nov.

Figs 179, 367, 518, 626

Types. Holotype: d, Champion Bay W Austr. Douboulay (OUM). - Paratypes: 27 9, same data (OUM); 19, Roebuck

Bay, Roebuck Bay (Australia N.W.) Coll. Castelnau, det. scolytides, sp. affinissima all Iıydrobioides ma diff. per gli

elitri punteggiati, n. sp. ?, punctatus Cast. (???), teste Gestro (MCSN); 18, West Australia, Ex Musaeo H. W. Bates

1892 (MNHN); 19, N. W. n. Holl., Janson Acq. 1884 (MNHN); 1%, Du Boulay, Nov. Holl.’Oeei, Fry Coll:

1905.100., det. gyrinoides (BMNH); 15, Ultima, V.G. Goudie (NMV); 1%, Aust. (CBM).

Diagnosis. Medium-sized, piceous species, distinguished by wide, moderately convex pronotum

with moderately wide apex, coarse, dense puncturation on pronotum and elytra, symmetric 3 genital

ring, short and wide, asymmetric, on lower surface strongly striate aedeagus with rather acute apex,

elongate, apically convex, strongly striate left paramere, and more or less triagonal stylomere with

rather narrow apex.

Description

Measurements. Length: 5.9-6.8 mm. Ratios. Width/length of pronotum: 1.69-1.78; width base/

apex of pronotum: 1.52-1.58; width pronotum/head: 1.65-1.70; length/ width of elytra: 1.45-1.55; length

elytra/pronotum: 2.43-2.56.

Colour. Upper surface more or less dark piceous, lower surface including mouth parts, antennae,

and legs reddish-piceous.

Head (Figs 179a-d). Rather short, fairly wide, rather depressed. Anterior border gently convex,

229

Figs 179a-l. Adelotopus adustus, spec. nov. Details of head and genitalia. For legends see fig. 100.