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COLOMBO MUSEUM.
GUIDE TO THE COLLECTIONS.
PREFACE.
= SHE first Guide to the Colombo Museum for the use
GN) Of visitors was compiled by Mr. Amyrald Haly in
1886 ; a second, abridged edition of it was printed in
1895.
having the additional advantage of illustrations.
Most of the principal objects will be found mentioned in the
text. One rather important omission may be noted here. Among
the examples of Tamil jewellery referred to on page 17, attention
should have been directed to the “ Thali,” the marriage emblem of
the Tamil race, which is worn round the neck mounted upon a
more or less elaborate necklace.
The collection of rocks and minerals has been entirely re-
arranged and greatly augmented by the Director of the Mineral
Survey, Mr. A, K. Coomaraswamy, B.Sc., who has kindly written
the account of the rocks and minerals of Ceylon for this Guide
(see page 50).
In the compilation of the rest of the Guide I have had the
assistance of the Museum staff, more particularly Mr. Gerard A.
Joseph, Secretary and Librarian ; Mr, H. M. Gunasekara, Assistant
Librarian; and Mr. H. F. Fernando, Taxidermist.
ARTHUR WILLEY,
February 9, 1905. Director, Colombo Museum.
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A GUE TO * THY,
COLLECTIONS IN
THE COLOMBO MUSEUM.
INTRODUCTION.
MIHE collections of objects of antique, local, and general
interest which are exhibited in the Colombo Museum
are intended to illustrate solely the products of human
ingenuity and cultivation, and the forms of nature as manifested
in the Island of Ceylon and its dependencies. With few exceptions,
which are specially noted where they occur, nothing is shown in
the galleries which has not been found in the country or in the
surrounding seas.
Owing to considerations of space on the ground floor, the
various Buddhistical, Ethnographical, and Archeological exhibits
are not arranged in such a strictly systematic manner as could be
desired.
The Zoological collections occupy the upper floor of the Museum,
The Mineral Gallery is situated at the back of the main building.
If reference be made to the ground plan of the Museum it will
be seen that the rooms to the right of the entrance hall are assigned
to the Library and Reading Room. The Library contains an
exhaustive assemblage of books bearing directly or indirectly upon
the religion, agriculture, archeology,and natural history of Ceylon.
There is also a valuable collection of native literature in the
form of Ola Manuscripts, relating in one form or another chiefly to
B 105-04
Cae)
the Buddhist Scriptures. These documents are written in
Sinhalese characters by hand with a pointed iron stylus, upon
properly prepared slips of palm-leaves called “ ola,” and are com-
posed either in Sanskrit, in Pali, or in Elu, the pure Sinhalese
language.
The ola leaves which are employed for the transcription of
the Sinhalese texts are made from the fan-shaped fronds of the
Talipot Palm (Corypha umbraculifera), which grows principally
in the Kandyan Districts; those adapted for Tamil usage by
school children and others are commonly made from the similarly
shaped leaves of the Palmyra Palm (Borassus flabelliformis),
which is especially abundant in the northern parts of the Island.
The Palmyra Olas are narrower, thicker, and less pliable than
the Talipot Olas.
The manuscripts are often consulted by Buddhist priests and
other readers who frequent the Library, and may be inspected, if
desired, on application being made to the Librarian.
CENTRAL HALL.
The first case to meet the eye of the visitor entering the Museum
is that which is placed under the archway before the main stair-
case. It affords a characteristic display of images of Gautama
Buddha, These figures are executed in brass, bronze, wood, and
ivory, and they represent the Founder of Buddhism (who lived
about the fifth century B.C.) in three principal attitudes—sedent,
erect, and recumbent. The head is generally surmounted by a
five-rayed emblem called “sirispota,” which symbolizes the sacred
flame,
On the top of the case there is a large wooden dagaba or relic
case. The ancient dagabas at Anuradhapura and elsewhere are
immense structures supposed to have been erected over various
relics of Buddha and his disciples. Representations of the dagaba
on a small scale, in wood, metal, and ivory, are commonly used as
emblems or as reliquaries, just as the small effigies of Buddhaare
portable copies of the gigantic statues which are scattered about
the country.
The lower portion of this case contains, on the front side, a
selection of Matara Jewellery, dating, at least with regard to the
designs, from the Dutch period (1655-1796 a.D.). The collection
comprises necklaces, brooches, hair ornaments, &c. Most of the
pieces are parures of the so-called Matara diamonds (zircons),
white sapphires, and black tourmalines, in a silver or silver-gilt
setting, manufactured by native jewellers at Matara, Galle, and
Colombo,
findre & Sieich LP
és TALIPOT PALM.
[To face page 2.
Gian)
On the other side of this table case there is a miscellaneous
display of Tamil silver waist-bands, charms, a Mudaliyar’s dress
sword, &e.
Products of the Palmyra’ Palm,—This palm grows in the
low-lying dry parts of the Island. There are extensive native
plantations in the Northern Province, especially in the Jaffna
Peninsula and the outlying islands. It shares with the Cocoanut
Palm and the Date Palm the distinction of providing more
serviceable commodities for the use of man than any other single
species in the vegetable kingdom. As already mentioned, the
leaves are employed in the manufacture of olas; they are also
used for fences, thatching, fans, mats, hats, baskets, water balers,
and umbrellas.
The fruits ripen in the months of August and September, when
they fall to the ground, and are sometimes eaten raw, but more
generally roasted [W. Ferguson]. They vary in qualities of
colour, smell, taste, and shape. From the fleshy part of the fruit
a sweet farinaceous jelly is prepared, called Palmyra Pulp or
“ yunatoo.” The nuts are sown under loose sandy soil, and the
very young subterranean saplings, after being cleaned and dried,
yield the Palmyra Flour.
Palmyra Toddy is prepared from the sap of the flower buds,
which are tapped by the toddy drawers during the months of
November and December, the rainy season of the Northern
Province.
Sugar or “jaggery ” is prepared from sweet toddy,7.e., from the
palm juice which has been prevented from undergoing fermenta-
tion by coating the inside of the toddy receiver with lime or
* ¢hunam,”
On the top of the case there are models of a Jaffna bungalow
with Palmyra Roofing, a shelter for watchers in the paddy fields,
a manger, and a platform for grain.
The Palmyra Palm is dicecious, 7.¢., the male and female flowers
are on different trees. Ina plantation half the trees will be male
and half female. The female tree yields superior timber and a
greater quantity of toddy than the male tree.
The model of the Palmyra Palm and other articles in this case
were presented by Sir W. C. Twynam, K.C.M.G.
Commercial Products.—Until 1880 coffee was the staple export
since the British occupation. During the Dutch administration
the Government held a monopoly of the cultivation of cinnamon,
but this industry is fast disappearing under competition with
other countries, The Cinnamon Gardens of Colombo are noted
for the extreme rarity of the cinnamon shrubs, whole plantations
(fs)
of which have been removed during the past ten years to make
room for building purposes.
From 1880 to 1886 Ceylon passed through a financial crisis in
consequence of the failure of the coffee trees, which were destroyed
by a fungoid disease caused by an organism named Hemileia
vastatrizx, for which no cure could be found. During this time
cinchona and tea planting came into being, and Ceylon is now
chiefly famous throughout the world for the excellence of its tea.
Other products of importance are cacao, cinchona, cardamoms,
and rubber.
More than sixty varieties of rice or “ paddy” are grown in the
Island, all of which, with one exception(the variety called “ el-vi”’),
require more or less continual irrigation.
Another grain of great importance to the poorer natives is that
which is called millet or “kurakkan” (Hleusine coracana). This
isgrown on waste lands called ‘“‘chena,” a corruption of the
Sinhalese word “ hena,” meaning ground prepared for cultivation
at intervals of several years by the cutting and burning of jungle.
The arecanut is the fruit of the Areca Palm, the tall slender
stems of whichafford a pleasing contrast with the unending groves
of cocoanut palms. It is used for chewing with the betel leaf, and
also has some medicinal value.
Native tobacco is extensively cultivated in various parts of the
Island.
On the top of the case are specimens of the gum of the cashew or
caju tree (Anacardium occidentale), a common tree yielding an
edible nut, but not endemic, having, it is thought, been introduced
from Brazil by the Portuguese ; and the resin of the tree called
“hal” in Sinhalese (Vateria acuminata).
Products of the Cocoanut Palm.—The objects exhibited in this
case have on the whole a familiar homely appearance, and bear
eloquent testimony to the world-wide importance of the tree upon
which, in the first instance, the wealth of the Island largely
depends. Almost every part of the tree subserves some useful
purpose, and its general commercial value far exceeds that of the
Palmyra Palm. It begins to bear fruit at about the tenth year,
and a single tree may yield about seventy nuts annually. It is
moncecious, 7.¢., male and female flowers are on the same tree, so
that every tree in a plantation will be fertile.
Most compounds of bungalows in Colombo are planted with
cocoanut palms, which can only be cut down by tenants upon
payment of ten rupees for each tree.
The husk of the fruit yields coir fibre, the shell can be used for
drinking vessels, bowls often handsomely carved, spoons, charcoal,
Caos
&c. The kernel is largely used in cookery, being grated fine by
an instrument called a cocoanut scraper, after which milk can
be expressed from it; when dried in the sun it is known as copra,
from which oil is extracted, the residue being used as cattle food.
The leaves are plaited to form cadjans for thatching roofs, also
baskets; and the trunk yields good timber. The young fruit,
called “kurumba,” furnishes food and ‘drink. The sap of the
unopened flower supplies toddy, arrack, and jaggery.
The dried frond of the palm is twisted into a bundle and used
as a torch. These torches are often employed for purposes of
illumination on festival occasions, being known as “ chulu ” lights,
a corruption of the Sinhalese word “ huluatta.” Torches are also
furnished by the spathes of the tlowers, called “ kolapuwa.” The
midribs of the leaflets are tied into bundles and form excellent
besoms, called “ekel” brooms, a corruption of the Tamil word
“ireku,’ meaning the midrib of a palm leaf.
The preparation of coir fibre is an important industry in the
Western and Southern Provinces. The following account taken
from Dr. Shortt’s Monograph of the Cocoanut Palm applies
equally to the methods in use in Ceylon as to the districts in India,
to which he refers more particularly :—
“The husks, removed from the nuts, are collected and thrown
into pits containing water to soak, and kept there till decompo-
sition sets in.” [Along the railway from Colombo to Galle many
portions of the backwaters and estuaries are fenced in for this
purpose. ] ‘The coir, when taken out of the pit, is beaten with
stout sticks to break up the adhesion and free the fibre from
impurities. Next it is hand-rubbed”’......... and ‘‘ subsequently
rolled into loose pads of about a finger’s thickness preparatory to
being twisted into yarn by the palms of the hands.”
In the bottom sheif is shown the apparatus employed in the
distillation of arrack, and on the top of the case there is a similar
apparatus in native pottery.
Fisheries and Transport,—Many of the models in this case
were made for the Chicago Exhibition of 1893.
On the top shelf are shown models of a bullock cart, a Kandyan
grain store, fish traps, a rattan bridge, and a “chekku” or oil mill
for expressing oil from copraand for the manufacture of gingelly
oil. The “chekku” consists of a huge mortar sunk deeply into the
ground and made of stone inthe Western Province, or of tamarind
wood in the North-Central Province ; in this a heavy pestle
revolves, being worked by a horizontal lever driven round by a
bull or a pairof bulls. A man usually sits on the lever to increase
the weight of the pestle.
Ci}
On the second shelf there are more models of carts and
hackeries, a mud house, and a large native sailing craft called a
‘“‘dhoney.”’
Onthe third shelf there are models of “ kattumarams,”’ a Negombo
canal ‘‘padda” boat, and a boat used in the Pearl Fishery with
representations of the crew and divers. The white man in the
stern holding asuspicious looking bottlein a compromising attitude
is the doctor preparing a dose of medicine.
On the last shelf there are models of outrigger boats, fishing and
passenger boats, a double canoe, and a river raft. There is also a
set of chank shells (Zurbinella pyrum) and several rings cut
from this shell. The chank fishery at Jaffna has been an
important source of revenue. During some years as many as
three millions of these shells have been exported annually to
Caleutta, where they are used for the manufacture of temple
conches and of chank jewellery which is destroyed at funerals.
Other noteworthy exhibits on this shelf are a pearl diver’s coir
basket which has been actually in use, presented by J. Hornell,
Esq., Marine Biologist, and a pearl diver’s sinking stone from
the Pearl Fishery of 1904, presented by the Hon. Mr. E. F. im
Thurn, C.B., C.M.G., then Lieutenant-Governor.
The chank shells and rings were presented by Sir William
Twynam.
Objects from the Maldive Islands.—The Maldive Islands are
an archipelago of coral atolls inhabited by a Mohammedan popu-
lationruled by a Sultan of ancient lineage, who pays annual tribute
to the Ceylon Government. The Maldivians are an artistic
people, the commonest articles in daily use being elegantly shaped,
carved, and lacquered. They make use of Huropean glassware and
earthenware, but protect their dishes and plates and glasses “fin
boxes or cupholders of the most elegant designs and elaborate
carving and colouring. Their boats are also elaborately decorated
when new. On State festivals the capital, Malé, presents a most
gay appearance, the roofs of the houses being covered with richly
coloured cloths, and all the streets profusely decorated with
bunting and curious models of modern steam vessels and little
kiosks furnished with chess tables, the whole being brilliantly
illuminated at night, when the Sultan, amidst a profusion of fire-
works, and preceded by his band, visits the numerous mosques ”
[A. Haly].
The Maldivian sea-going sailing vessels, called buggalows, are
often to be seen in Colombo Harbour, and the view from the end
of the breakwater of one of these boats entering the harbour during
the north-east monsoon is highly picturesque. .
——- = SMES SS ALIA nae
PILLOW AND LACQUERED STAND.
-MAKING
LACE
MALDIVIAN
(Total height, 1 foot 2 inches.)
(To Jue page 6.
RICE MEASURE. PLATE BOX. BOTTLE BOX.
(Diameter, 11 inches.)
MEDICINE BOX FLOWER BOX. INK BOX.
(Height, 8} inches.)
MALDIVIAN LACQUERED BOXES.
To face page 7.)
GE.)
The model of a ship with a mat sail in Case VI. represents, the
kind of boat used for traffic between the numerous islands of the
Maldive group. The models in Case VIII. were presented by the
Sultan of the Maldives, and do not include a copy of the typical
Maldivian buggalow, which is built on characteristic lines unlike
anything shown in these cases. In Case VIII. there are two finely
lacquered drums, spears, and musical instruments. In Case VI.
the chess boards, spinning tops, stands for lace pillows, Nautilus
_ shell spoens, weighing scales, and nautical instruments are
among the more noteworthy objects exhibited.
The lac employed in decorating the fancy boxes, dish covers,
drums, sticks, spears, and stands isimported into the Maldives from
India. The patterns into which it is worked, as well as the designs
followed in wood and stone carving (see below, Maldivian
Tombstones), appear to be exclusively Maldivian.
Many of the objects in Case VI. were presented by H.C. P. Bell,
Hsq., C.C.S., Archeological Commissioner. The rest formed part
of, a collection of Maldivian articles exhibited at the World’s
Columbian Exposition at Chicago in 1893, and were presented
by Sultan Ibrahim Noorudin Iskander, Sultan of the Maldives
from 1882 to 1893.
Masks and Musical Instruments,—Masks are used in plays,
masquerades, and devil-dancing. Their invention is attributed to
the god of curiosities. Those representing various diseases are
said to be employed by devil-dancers to exercise the devils who
occasion the sickness. Their construction appears to be based
upon the principle of eradicating disease from the system by the
homeopathic method of counterfeit presentments.
In spite of their grotesque character and of the fact that they can
be made to order at the present day, these masks possess a profound
interest as affording a clue to the origin of the ancient masks used
in the Greek plays. The Oriental masks of the demons have been
regarded as the prototypes of the Birds of Aristophanes, the
Giants of Pollux, and the frightful forms of Lucian [Upham].
“The mask is the type of the Metempsychosis, the great pivot of
Oriental doctrine, exhibiting to the spectator, scenically, the
changes and forms which in different stages of mundane existence
attach to the vital principle.”.......- ““Had masks originated with
the Greeks, it is fair to conclude that, instead of such frightful
specimens which abound in every museum, they would have given
the human form as they have beautifully embodied it in their
painting and sculpture ; hence the physiognomical character of the
masks may be said to decide their origin and locality to the East”
[Upham].
© 8.5)
The Maha Kéla Sanni Yaka, or Yaksha, represented by the
composite mask in the centre of the case and again over the top of
Case VIII.,* is the great Demon of Fatal Diseases, all of which are
attributed directly to devilish derangements of the three humours,
wind, phlegm, and bile.
The Gopolu, or Gopola Yaka, is the Demon of Cattle, and all
cattle sickness is supposed to proceedfrom him. Heis represented
with horns and tusks and a garment of leaves.
The Gara, or Garra Yaka, is the demon who possesses newly-
built houses, and before a house can be finally occupied a cere-
mony called Gara Yaka Maduwa is generally performed. This
ceremony is presumably equivalent to the European house-
warming.
Stniyama or Huniyama is the art of sorcery, bewitching by
spells and incantations. The word is sometimes anglicized into
Hooniyan, this being the name given to evils inflicted by a man
upon his neighbour or enemy by the agency of charms. In
Hooniyan charms a smallimage of wax or wood is made torepresent
the person whose death or injury is desired. “A few hairs of
his head, some chippings of his finger nails, and a thread or
two from a cloth worn by him, and sometimes a handful of sand
from a place on which he has left his footprint, are required.” The
image is then submitted to aceremony called Jiwama (“ endowing
with life”) performed by a Kattadiya or sorcerer, who recites
mystical wordsover it. Nails made of a composition of five different
metals—gold, silver, copper, tin, and lead—are driven into the
image through the joints, the heart, and the head, and the name
of the victim is marked on the image, which is then buried in the
ground under a stile or at some other spot where the victim is
likely to pass over it. ‘The passing over, or Panna-wana-wa, is
essential to the success of the charm. t .
This Hooniyan charm, or Sunniyan Yaka, as the specimen is
labelled in the case, is of particular interest on account of its world-
wide application in the practice of witchcraft.
On the top of the case, besides more demon masks, there are
large masks called the King and the Queen. These are used in the
native masquerades called K6lama.
The lower portion of the case contains a collection of musical
instruments, The drums are of various kinds, the more charac-
teristic being the flat drum or timbrel called Rabana; the
bobbin-shaped drum called Udikkiya or Udakiya ; Tammettama,
* This Kéla Sanni Yaka was presented by Mr. Justice H. L. Wendt.
{ The account given above of the Hooniyan charm is taken from an article
“On Demonology and Witchcraft in Ceylon,” by Dandris de Silva Gooneratne,
Mndaliyar, in the Journ. Ceylon R, Asiat. Soc., vol. IV., 1865-6, pp. 1-1 17.
GM)
two drums fastened together as kettle drums ; Demala-beré, Tamil
tom-tom ; Yak-beré, demon tom-tom. The Rabana is beaten by
women seated in a group round it on occasions of family rejoicing.
Among the stringed instruments are to be noted the Wenawa or
Vina, the Indian lute, an instrument with a good twang, the
resonator consisting of a cocoanut shell with a skin stretched
across it. The Bandarinha and Viola, presented by H. Holsinger,
Esq., are used by the Mechanics* of Ceylon to accompany their
Lusitanian dances.
There are also a couple of marionettes, employed in a form of
entertainment much in vogue among the Sinhalese. —
Other exhibits in the Central Hallinclude two stands of Kandyan
Spears with lacquered shafts, together with Kandyan blunderbusses
and processional fans.
On the top of Case VIII. there isan interesting relic of the early
conflicts between the British and the Kandyans in the form of a
British drum said to have been captured by the latter.
On a small stand at the foot of the staircase there are some
antique cannon balls, probably of Portuguese origin, which were
unearthed at Medamahanuwara, near Kandy, a place which is
noted for the existence of a cave in which the last King of Kandy
took refuge after his flight from the British, and where he was
captured in 1815.
MEDIA-VAL ROOM.
CASE IX.
Ivory Carvings.—This case contains a varied and valuable
collection of objects made principally of ivory. The specimens
which are worthy of attention include fan handles in ivory and
ebony, combs, panels, dagabas, &c.
The large boxes are carved and shaped after Dutch designs.
The art of making the compressible scent sprinklers is
said to be a secret confined to one family of ivory workers
in the Kegalla District. The little figures of the last King of
Kandy, two of his Ministers or Adigars, and the Chief Priest
are said to be contemporary portraits. Sri Wikrama Raja Sinha
was the last king of the Suluwansa or Lower Dynasty. He
came to the throne of Kandy in 1798 and reigned until 1815,
when he was deposed chiefly on account of his cruelty. The ivory
statuettes of Ehelapolaand his wifearealso of considerable interest.
Ehelapola became First Adigar of the King of Kandy in 1812. He
* The Mechanics of Ceylon are a class of artisans, shoemakers, tailors, blacksmiths,
craftsmen of Portuguese descent, speaking a lingo of their own, Portuguese with
an admixture of Tamil and Sinhalese. Cf. Mr. C. M. Fernando’s article on the
Music of Ceylon in Journ. Ceylon R. Asiat.Soc., vol. XIIT., 1893-1894, pp. 183-189-
Cc 105-04
( 10 )
wasalso Dissave of Sabaragamuwa. Having disobeyed an order to
proceed to Kandy his family was imprisoned by order of the king,
and subsequently his children were beheaded in front of the Maha
Vishnu Dewale at Kandy and his wife was drowned in the tank at
Bogambra, near Kandy. This incident is known as the Khelapola
Tragedy, and constitutes a favourite theme on the modern
Sinhalese stage.
On the lowest shelf of this case there are some more ivory
statuettes of Buddha, ivory flutes, and a richly carved rattle
mounted on a lacquered stick.
There is alsoa handsome ivory Udakiya (without skins) lent by
P. E, Pieris, Esq., C.C.S., and an antique ivory cigar mouthpiece
with receptacle for an extra cigar, presented by Mr. E. R. Goone-
ratne, Gate Mudaliyar.
The quaintly-shaped and lacquered pill boxes and the ola book
covers with the signs of the zodiac deserve notice.
The ivory dagabas are reliquaries or karanduwas, the dome being
screwed upon the base so that it can be removed and a cavity
disclosed in which any small object of veneration or votive offering
can be deposited.
CASE X.
Besides the numerous examples of Kandyan embossed metal
work which are exhibited in this case, the most striking object
is a silver model of the shrine containing the Dalada or Tooth of
Buddha, the reputed original of which is preserved in the Dalada
Maligawa at Kandy. This famous Tooth Relic has played an
important part in the political history of Ceylon. It is esteemed
by Buddhists as the palladium of the country and symbolizes the
inviolability of the Buddhist religion. Itisrelated that the sacred
relic was originally rescued by the sage Khema from the great
teacher’s funeral pyre at Kusinagara and given by him to Brahma-
datta, King of Kalinga, about 2,500 years ago. It was eventually
brought to Ceylon from Southern India by a Brahman Princess
of Kalinga, concealed in the folds of her hair, about the years
310-313 A.D., during the reign of Sri Megahavarna at Anuradha-
pura, where it was wont to be publicly exposed on sacred days
with gorgeous ceremonies. When the relic was first brought
to Ceylon its adventures were recorded in a work called the
Dhatuwansa or Chronicle of the Tooth, written in Elu, the classical
language of the Sinhalese. The tooth is said to represent the left
upper canine or eye-tooth. The legend runs that after all at-
tempts which have been made to destroy the sacred emblem, it
has reappeared resting upon a lotus flower, where it now reposes.
Parakrama Bahu I., surnamed the Great, built a temple for it at
KANDY.
TOOTH.
E OF THE
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[V0 face puge 10.
MODEL OF THK TOOTH RELIC,
To tace paye 11).
( FE)
Pulastipura, the modern Polonnaruwa, between the years 1190 and
1195. About the year 1246 a.p. Vijaya Bahu III. enshrined it at
Dambadeniya, whence some forty years later Bhuvaneka Bahu I.
removed it to Yapahu. Thence it followed the fortunes of the
Suluwansa Dynasty to successive capitals, Kurunegala, Gampola,
and Kotte near Colombo.
In the year 1560 A.D. it is said to have been captured by the
Portuguese and taken to Goa, where it was pounded in a mortar
and consumed in a brazier, but Phcenix-like it rose again from
its ashes and is now at Kandy.*
The vicissitudes of the Tooth Relic are matters of speculation
and controversy, but its political importance as a national
palladium during the dynastic periods seems to be beyond
doubt.
The model here shown was exhibited at Chicago in 1893.
The same shelf contains a handsome display of silverware,
amongst which may be specially noted the large silver dagaba
exhibited at the Colonial and Indian Exhibition of 1886: two
large boxes of beaten silver embossed with deities and. scroll
work ; a large spherical silver box with intricate design in high
relief ; an elegant silver chatty; a silver chalice for sandalwood ;
and a silver scent diffuser of the kind used for sprinkling guests
at wedding ceremonies and for spraying coffins in funeral proces-
sions. There isalsoa finely worked brassdagaba. The leaf-shaped
tassels hanging round the top of the dagabas represent the leaves
of the sacred Bo-tree (Ficus religiosa).
On the next shelf below a large series of brass bowls called
“chembu” is shown. Some of them are further adorned by the
inlaying of alternating strips of beaten silver and copper.
The bottom shelf contains further examples of Kandyan
brasswork, especially betel trays and rice tables, prominent
among them being an antique Kandyan rice table presented by
A. K. Coomaraswamy, Esq., Director of the Mineralogical Survey
of Ceylon. |
The upper shelves of the case contain on one side a set of
embossed silver and brass plates, and a pair of carved silver ola
covers with ola manuscript descriptive of one of the Jatakas or .
Births of Buddha.
On the other side there are some examples of wood carving,
a carved calamander cocoanut scraper from Panadure, between
Colombo and Galle ; sweetmeat moulds or jaggery boards ; game
boards called ‘‘chonka boards,” in which the seeds of the
* Cf. Memoir on the History of the Tooth Relic of Ceylon, by J, Gerson da
Cunha, 1875.
( 2 )
“olinda” (Abrus precatorius) or any other suitable seeds or
shells are placed in two depressions at the ends, and the players
have to make the circuit of the board from pit to pit along the
sides without occupying the same hole at one time. The player
who gets the seeds home first wins.
On the end-wall of the case there are some carved wooden
sweetmeat pats. Hanging from the top of the case down the
middle is a richly embroidered silk cloth said to have been worn
by the wife of Molligoda, the Second Adigar of the last King of
Kandy.
CASES XI. AND XIV.
Embroidered Cloths.—The narrow wall case contains examples
of dress worn by the old aristocracy of the low country.
In the centre is a hat of peculiar shape, somewhat boat-shaped,
called “ Jagalatta Toppiya,” used by Rajapakse, Chief Mudaliyar
of Mahabadde, 1701 A.D.
There is also a Mudaliyar’s dress sword and a sword with hilt
and scabbard of richly carved tortoise-shell dating from the end
of the 18th century, lent by Tudor Rajapakse, Esq.
On the top shelf there are some Dutch swords.
In the case corresponding to this on the opposite side of the
room (Case XIV.) some further examples of woven cloths are
shown, including a handsome old embroidered Kandyan betel
bag, which was formerly carried slung at the side from the
shoulder. There are also some gold embroidered Chetty costumes.
CASE XII.
This case contains an assortment of antique objects in brass
and bronze, among the more interesting of which are three
Sinhalese water clocks; cocoanut oil lamps; elephant bells ;
karanduwas (dagabas); Pattini bangles, hollow armlets and
anklets with a slot along one side and pellets inside, used in
dances on festival occasions such as peraheras, in honour of
Pattini Deviyo, the goddess of chastity ; epaulettes, also worn by
dancers.
The Sinhalese water clock isa clepsydra, consisting of a copper
bowl, of larger and smaller sizes, with a small pinhole in the
bottom and with or without silver datum marks let in at the
sides. The bowl is set floating ina clay water chatty, the water
gradually entering through the pinhole aperture until a datum
level is reached, and eventually the bowl sinks. In the larger
of the clocks shown with graduations the water reaches the
level of the highest datum mark in exactly forty-eight minutes.
The Sinhalese hour or “ peya” consists of twenty-four minutes,
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and the day and night are divided into thirty “peyas” each.
The water clock is called “ pe-tetiya.’”*
On the top of the case there are some interesting examples of old
Sinhalese domestic wood carving, an art which. has almost if not
entirely fallen into disuse. The series includes carved cocoanut
ladles and spoons with more or less ornate wooden handles,
carved wooden spoon racks, and a cocoanut scraper.
CASE XIII.
This case contains a large series of boxes and implements
employed in the services of the arecanut and tobacco. The
arecanut is the principal ingredient in that form of indulgence
known as betel chewing. A fragment of a nut with other spices
is wrapped up in a betel (pepper) leaf and eaten. Then the
finger is dipped in slaked lime called chunam and placed upon
the back of the tongue, or sometimes a spoon or spatula may be
employed for this purpose. The lime is kept in metal boxes,
called chunam boxes, which are elegantly shaped, embossed, and
inlaid. The box is suspended by a chain, at the end of which,
when complete, are carried a small silver or brass earpick, a
toothpick, and a spatula. The very large chunam boxes be-
longed to important personages and were carried by an attendant,
The arecanut is cut into slices by an instrument, resembling
a nut cracker in shape, called an arecanut cutter, The handles
of these cutters afford considerable scope for artistic display,
as will be seen by an examination of the large series here
shown.
Elderly persons with failing powers of mastication are in the
habit of pounding their betel bolus before consuming it. For this
purpose they use a small pestle and mortar called a_ betel
pounder, several of which are exhibited.
Above the chunam boxes there are some antique jewel boxes
made from Dutch designs in brass and copper, embossed and
engraved in various patterns. These are known as Dutch boxes,
other examples of which are to be found in the Ivory Case and
in Case XV.
Below the shelf containing the arecanut cutters there is a
large series of brass tobacco boxes, These have also been made
from Dutch designs, some of them having been actually manu-
factured in Holland, whence they were brought here during
the Dutch Administration.
* Further information on “Sinhalese Measures of Time” is contained in an
article by Mr. Herbert White, C.C.S., in The Orientalist, vol. III., 1888-1889, p. 75 :
and in a paper by Mr. F. H. Modder on Sinhalese Weights and Measures, in Journ.
Ceylon R. Asiat. Soc., vol, XII., 1892, pp. 173-202.
( 4)
The bottom shelf of the case contains some more metal bowls,
trays, and goblets.
The picture on the wall over the case is a temple drawing
representing an incident in the life of Buddha.
CASE XV.
A portion of this case contains a number of “ Dutch boxes,”
many of which, however, have been made in more recent times.
They are made with different kinds of wood—satinwood, cala-
mander, and ebony—and are variously carved and inlaid with
ivory, brass, tortoise-shell, and porcupine quills.
The original native wood carving has largely given way to
the manufacture of these articles and of ebony and cocoanut
elephants. There is also shown here a well-executed carving
of a tortoise in calamander wood, the most valuable wood in
Ceylon.
3esides the boxes there are some examples of carved combs and
hairpins in tortoise-shell and in horn.
On the other side of the case some examples of painted
Kandyan pottery are shown. There are three classes of unglazed
pottery in Ceylon, namely, the plain Village pottery, comprising
the water chatties, cooking bowls, and curry dishes of every-day
use; secondly, the painted pottery of Kandy; and lastly, the
Grotesque pottery of Matara, examples of which are placed upon
the top of the case. This pottery possesses features of ethno-
graphic interest in spite of its grotesqueness ; it is made and sold
chiefly during the time of the Dondra Fair in the summer
months.
CASE XVI.
Kandyan Knives and Swords,—Here are shown numerous
swords and daggers used by the Kandyans during the later Dynastic
Period and still worn on State occasions. Many of them are highly
ornate at the hilt, and the scabbard and base of the blade are often
richly damascened. The handle isfrequently carved out of ivory,
horn, and black coral, and the sheath in some cases is covered with
carved tortoise-shell. At the base of the blade ina few instances
the figure of a lion in brass is let into the steel. ‘This seems to be
of the nature of heraldry. ;
Some of the dagger sheaths contain in addition to the dagger a
receptacle for a stylus for writing upon the ola slip.
On the top shelf there is a set of Kandyan Village jewellery
in the form of numerous brass and glass bangles ; and some old
Dutch swords and powder-horns.
On the top of the case are some antique spear heads.
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TABLE CASE XVII.
A rather heterogeneous assortment of ancient odds and ends is
provisionally placed in this case, gold and silver fragments,
beads, and gems from the ruined citiesof Ceylon. The excavations
which have been carried on for many years under the direction of
the Archeological Commissioner have not led to any sensational
discovery of buried treasure. Such precious relics as have been
unearthed have on the whole been disappointing so far as their
intrinsic value is concerned.
In the reverse half of this case there is an ola horoscope and a
copper sannas or deed conveying a grant of land to a Kandyan
temple by the last King of Kandy. There is also shown an
ebony weighing lever, called Tulappadi in Tamil, still used by
traders in the Vanni and Jaffna, presented by J. P. Lewis, Esq.,
C.C.S.
In the glass box over the case there are two old swords, with
Sinhalese legends dating from the years 1374 and 1416 inscribed
uponthem. The inscriptions relate that the swords were presented
by the Rajas reigning during the years mentioned (1917 in the
Buddhist Era = 1374 in the Christian Hra; 1959 A.B. = 1416 A.D.)
in the town of Jayawardhanapura (the modern Kotte, which lies
in the outskirts of Colombo) to two members of an aristocratic
family upon their appointment as generals. These interesting
swords were presented to the Museum by Mr. C. M. Fernando,
Crown Counsel.
TABLE CASE XVIII.
Containing a valuable collection of coins which have at one time
beencurrentin Ceylon. ‘The coins fall into two classes, namely,
the ancient Sinhalese currency, comprising the coins of the Kings
of Ceylon ; and the foreign coins introduced to this Island by
traders from the days of the Roman Emperors down to the estab-
lishment of British Rule. The devices on the coins of the Sinhalese
Kings represent on the obverse the king standing, holding a lotus
flower in his right hand and a kind of sceptre, sometimes called
the trisul emblem, of questionable significance, in his left hand.
On the reverse the same figure is repeated in a sitting attitude with
the name of the king inscribed to the left of the figure in Nagari-
Sanskrit characters.
It has been a too common practice to forge counterfeits of the
gold coins for the purpose of deceiving collectors.
The most ancient coins represented in the collection are rectan-
gular pieces of silver with or without figures of animals punched
upon them, called Eldlings, which have been found during the
excavation of the ruined cities.
(16 )
Among the rarer Sinhalese dynastic coins may be mentioned
the Lion Coin and the Setu Bull Coin, examples of both of
which are exhibited. ‘To these may be added the very rare gold
Lankeswara coin of Vijaya Bahu, lent by P. E. Pieris, Esq.,
0.C.S.
Among the foreign coins may be noted the Roman and Arabian
coins, Venetian gold sequins, Portuguese silver tangas or tangams,
and gold San Thomé coin, the Dutch dukatoons and silver and
copper stuivers, and challies minted by various States in the Dutch
Confederation. Some of the Dutch copper coins were actually
minted in Ceylon, at Colombo, Galle, and Trincomalee. These
are marked with the letters C, G, and T, respectively.
The establishment of the Dutch United East India Company
(Ostindische Vereenigde Compagnie, indicated on the coins by
the monogram &) on the Island of Ceylon dates from the year
1655 and lasted until 1802, when the Island was formally ceded to
the British (who had occupied it in 1796) by the Treaty of
Amiens.
The copper ingots issued by the Dutch, of the value of 4}
stuivers, are a singular form of money.
The general name applied to the ancient Sinhalese coins is
‘““massa.” ‘They appear to date only from the year 1153 A.D. to
1296 A.D.
The common copper coins of the Dutch of small value were
called challies, a corruption of the Sinhalese word “salliya”
(plural “ salli’’), meaning money orcashin general. The smallest
coin now in use, value half a cent, is still called “tamba-salliya,”
“tamba’’ meaning copper. The proper coins of the Sinhalese
King during the famous captivity of Robert Knox (1659-1679)
were fanams of the size of a spangle.
Another interesting form of money is afforded by the Larins
or Fish-hook money. These are said to have originated at a place
called Lari or Laristan on the Persian Gulf. They were formerly
made in the Maldive Islands, and were also in use in Ceylon in
Knox’s time, anybody being allowed to make them. Portuguese
copper tangams were also current.
Further information on the ancient coins of Ceylon is contained
in the well-known memoir by Professor T. W. Rhys Davids “ On
the Ancient Coins and Measures of Ceylon,” published in 1877 in
the International Numismata Orientalia (London, Triibner &
Co.). .
The ancient beads, coins,and dice discovered at Anuradhapura,
Mihintale, and elsewhere have been described and figured by
Mr. H. C. P. Bell, C.C.S., the Archzxological Commissioner, in his
( iW )
Fourth Progress Report on the excavations at Anuradhapura and.
the North-Central Province (Sessional Papers, 1892).
In the large glass shade over this case are shown numerous
ancient images, mostly sedent figures of Buddha, including eight
thin gold images of Buddha, filled with clay, from Panduwas
Nuwara, and above these a similar gold figure from Tissamaharama
and gold and crystal dagabas from Anuradhapura. At each end
of the cover inside there is a bronze Buddha on a throne backed
by a well executed arch called “makara torana.” These are
called “ Enthroned Buddhas.” There are also figures of Krishna
playing with a ball, and of the goddess Pattini, the latter from
Trincomalee. On the top of the case there is a large sedent
bronze Buddha of unique design in the attitude of teaching,
holding a flower (?) inthe lefthand. This was discovered twelve
miles from Badulla along the new road to Batticaloa, and was
presented by G. F. K. Horsfall, Esq., in 1876. On either side of
this image there are two common village coloured wood-carvings,
one representing a large cobra, the other Buddha seated upon the
folds of a cobra and protected by its expanded hood. The latter
is called a “ Serpent-canopied Buddha.”
TABLE CASE XIX.
A selection of silverware occupies the two halves of this case,
and in the glass shade above there is a set of Tamil silver bangles,
anklets, and toe rings.
Besides some richly damascened Kandyan knives, embossed
silver tobacco boxes, and Dutch relic in the form of a silver plate
presented by a former Dutch Governor of Ceylon to the person
named in the inscription, the principal object in the case is a pair
of handsome silver ola covers with bejewelled button. They
consist of bars of wood painted with dagaba devices on the under
side and overlaid with sheets of beaten silver.
TABLE CASE XX.
Examples of Chetty, Tamil, Sinhalese, and Moorish Jewellery.—
The large gold ornaments are represented here by silver-gilt
replicas. ‘“ Chetty” or “Chitty” is the name applied in India to
all members of the trading castes in the Madras Provinces. The
Colombo Chetties, a caste from Tinnevelly, emigrated to Ceylon
about the middle of the sixteenth century. Their language and
customs are Tamil.
From an ethnographical point of view among the most interesting
objects in this case are the Sinhalese “ nawaratna ” rings set with
the nine principal gems, or as near an approximation as is possible
or can be afforded. The nine gems stand for the nine planets
(including sun and moon), and the ring is worn as a corrective
D 105-04
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for horoscopic purposes ; for example, it isworn bya person born
under an unfavourable star, and its constant usage is supposed to
avert disasters.
STONE GALLERY.
Many remarkable remains of fallen greatness, illustrating the
stupendous grandeur of the ancient religious monuments of
Ceylon during the palmy days of militant Buddhism, are exhibited
in this gallery. The statues, pillars, friezes, and slabs are carved
out of gneiss, the country rock of Ceylon, some of them, however,
consisting almost entirely of crystalline limestone.
Among the more notable pieces are the three principal archxo-
logical treasures of the Colombo Museum mounied in position along
the centre of the room. Facing the south window atthe front end
of the gallery is a perforated carved slab, 4 feet 8 inches high,
2 feet 10 inches wide, and 7 inches thick, known as the Yapahu
window, from Yapahu or Yapahuwa, a village in the North-Western
Province, about twenty miles north of Kurunegala. It consists
of a single block of gneiss cut into the semblance of a frame, which
surrounds a composite hieroglyph consisting of forty-five circles
in five vertical rows joined together in a moniliform pattern, each
circle containing an emblematic figure repeated on both sides of
the stone. The matrix of the slab between the carved portions
was removed by the artist who designed and executed this unique
triumph of stone tracery. In the 13th and 14th centuries there
was a royal palace at Yapahu, and the hall of the palace was
lighted by two of these tracery windows of exquisite workmanship.
We are told by Mr. F. H. Modder that one of these windows “ was
perfect in 1850, but the other had fallen and its fragments were
seattered around. The remaining one would doubtless have soon
shared its fate had not Mr. O’Grady, then Government Agent of
the North-Western Province, removed it to Kurunegala..........
Thence it was transported to Colombo, and now occupies a
prominent place among the archzological exhibits at the Museum.”
The human figures in the lowest circles represent "grotesque
manikins, above these are nautch girls, then animals, some of which
are provided with a trunk and appear to represent the fabulous
“ gaja-sinha” or elephant-lion. The star-shaped radiating emblems
are the “ dharma-chakra ” symbols, the wheel or circle of the laws
and teaching of Buddha. The birds in the top row are the
“hansa” or sacred birds, usually represented by geese, sometimes
by conventional representations of birds.*
* For further remarks quoted from an article by Mr. John Bailey, C.C.S., who
explored the ruins in 1850, see the paper by Mr. F. H. Modder on “ Ancient Cities
and Temples in the Kurunegala District : Yapahuwa.” Journ, Ceylon R. Asiat. Soc.,
vol, XTII., 1893, pp. 97-113.
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The next megalith which claims attention is the colossal figure
of a lion called the Lion of Polonnaruwa. This relic of the past
-is exceptionally valuable and interesting, because there is a
Sinhalese inscription on each side near the base giving the date and
purport of the monument. Upon it was placed the throne of
King Nissanka Malla, a Chakrawarti or Emperor of Kalinga lineage,
who was the Lankeswara or Overlord of Lanka (Ceylon) during the
years 1187-1196 A.D.
The lion formerly stood in the Council or Audience Hall of the
King at Polonnaruwa, whence it was removed to the Colombo
Museum about thirty years ago. The ancient name of the city was
Pulastipura, the modern name is Topawewa, meaning the tank
where the ruined topes or stupas are. It is, however, commonly
known as Polonnaruwa, an Elu term of doubtful derivation
adopted by Sir Emerson Tennent (Ceylon, vol. II., 1847). The
ruins were re-discovered in 1820, and all that remained of the
Audience Hall where the inscriptions were found were ‘48 large
stone pillars with carved capitals supported on a’stone platform,
round the base of which are sculptured a row of lions.” The
great lion-throne “ was lying almost entirely buried at some distance
from the Hall, and was set up with great difficulty ; it had probably
been thrown out of the Hall by the Tamils when they took
Pulastipura, and may formerly have stood between the inscribed
pillars.’”’*
The inscription on the left side of the lion is terminated by the
figure of a fish, a symbol of good omen.
The adventures of the lion during its transport from Polonnaruwa
to Colombo are recounted by Sir William Gregory (Autobiography,
second edit., 1894, p. 343), who was at that time (1872-1877)
Governor of Ceylon: “ Hvery mishap attended the transfer of this
huge stone beast. Its first dray fell to pieces beneath its weight.
On descending from the elevated ground where it stood the two
elephants attached to it pulled over-vigorously, and the dray and
the lion and the elephants flew apart in different directions. It
had then to be drawn over a difficult jungle path a distance of
fifteen miles from the main road; but the elephants had now
learned their business, and these obstacles were surmounted. But
* A facsimile of the inscription on the left of the lion, with translation, is
given by Professor T. W. Rhys Davids in his paper on “ Inscriptions at the
Audience Hall of Parakrama Bahu, Pulastipura, Ceylon,” in the Indian Anti-
quary, vol. II., 1873, pp. 246-249. Pulastipura was the capital of Ceylon from the
end of the eighth to the beginning of the fourteenth century. Previously
Anuradhapura had been the capital for over a thousand years.
Pulastipura enjoyed its period of greatest magnificence during the long reign
of Parakrama Bahu I., surnamed the Great, in the latter half of the twelfth
century, preceding the reign of Nissanka Malla.
( 20)
when it reached the high road the worst of all remained. The
wooden bridges, constructed to sustain a moderate load, were quite
unable to bear the combined weight of the lion and the dray, and
the banks were precipitous and deep. But this, too, was overcome
by digging out asloping passage to the bed of the river and another
on the opposite side. The elephants with their immense strength
and sagacity sustained the strain of letting down the lion, and
easily drew it up again. Much of this took place in the solitary
jungle, but when the inhabited regions were approached, the whole
country turned out in amazement.........
“The procession of elephants, the lion decked with wreaths
and flowers, was a magnificent sight. The tom-tommer from each
village joined the cortege. The headman of the district asked
permission for his little boy to ride the monster into Matale,
whence he was to be conveyed by rail to Colombo. The lion now
stands calmly in the Museum, and few know, or could understand
if told, all the cares it caused and the excitement it created. It
is a most valuable archeological record, and would have been
undoubtedly destroyed ere this had it not been removed.”
The risk of destruction referred to by Sir William Gregory in
the foregoing quotation is demonstrated by the fracture on the left
side of the head, which is said to have been perpetrated by enter-
prising burglars ignorant of the solid nature of dynastic art who
hoped to find treasure hidden within the penetralia of the body.
From the base of the forefoot to the crown of the head the lion
stands six feet.
The third object of distinction is an elaborately carved pillar
which has been recently set up behind the great lion. This is
called the Medagoda Pillar, and formerly stood in the Pattini
Dewale at Medagoda, six miles below Ruanwella, in the Province
of Sabaragamuwa, not far from Yatiyantota. An excellent drawing
of this pillar, accompanied by a lucid description quoted below, is
contained in the “ Report on the Kegalla District of the Province
of Sabaragamuwa,” by H. C.P. Bell, Esq., C.C.S., Archeological
Commissioner (Sessional Papers, 1892, p. 58): “The monolith
must originally have been squared to 1 ft. 2 in., the size it assumes
across the lion’s breast, lotus bosses, and capital fillet. Rising
octagonally from the back of a broad-faced couchant lion of con-
ventional type, with frilled mane and raised tail, the shaft slides
gradually into the rectangular by a semi-expanded calyx mould-
ing. Half way up relief is given by a bordered fillet 2 in. in
breadth, slightly projecting, carved with a single flower pattern
repeated round the pillar. From the fillet depend on each face
twoconcentric ......... pearl-bead strings. A few inches above this
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band stand out from alternate faces full-blown lotus, knops, 5 in.
in circumference, with ornamentation resembling much the Tudor
flower upon the intervening sides. Where the pillar becomes
square there are further loops of pearls, four on each side. A
lower capital of ogee moulding, separated by narrow horizontal
fillets, and finished with ovolos and a rectangular band, is sur-
mounted by a four-faced makara and a lowabacus, From the
centre of the roundlet moulding on all four sides drops the
garlanded chakra symbol.”
Other noteworthy stone carvings in this gallery are the Janitor
Stones from Hanguranketa presented by OC. H.de Soysa, Esq., placed
before and behind the cement base upon which the Yapahu
window now stands ; amystie square stone called yantra-eal, with
twenty-five holes, from Anuradhapura, supposed to be a base stone
sometimes called a yogi stone (cf. H. C. P. Bell, Seventh Report on
Anuradhapura, Sessional Papers, 1896, p. 13) ; the Naga Stone from
Anuradhapura, mounted against the east wall, portraying a large
seven-headed cobra in high relief; friezes from Horana; figures
of Ganesa, the elephant-god, from Horana, presented by Sir C. P.
Layard ; marble statue of Buddha from Tissamaharama, presented
by Sir C. P. Layard; friezes and capitals from Anuradhapura ;
statue of Buddha in spongy gneiss from Ambagamuwa, presented
by Hon. Mr. R. B. Downall (against the west wall) ; cast of the
gigantic statue of Parakrama Bahu the Great at Polonnaruwa; cast
of a remarkable Processional Moonstone from Anuradhapura
(below the south window). The moonstones, perhaps so called in
consequence of their semilunar shape, are employed assteps leading
into the porticos of the temples. They are often of fine design
and execution, and are characteristic of Sinhalese Buddhist
architecture. In the cast exhibited here there is a central lotus
flower surrounded by concentric processions of hansas and other
animals. In front of the stone lion there is another simple
Lotus Moonstone from Hanguranketa, presented in 1878 by
C. H. de Soysa, Esq.
Attention may now be directed to the four wall cases in this
room, three of which contain ethnographical models, and the
fourth a valuable collection of ancient bronzes.
CASE XXI.
Models of a Kandyan Chief or Ratemahatmaya and of a Buddhist
priest with begging bowl ; a temple tapestry hangs at the back of
the case.
CASE XXII.
A Low-country Chief or Mudaliyar and a Sinhalese bride.
Ca cd
CASE XXIII. .
A man and woman of the Veddas, the aboriginal hunting caste or
hill tribe of Ceylon. The bark-cloth bag hanging against the side
of the case is made from the bark of the upas tree, Antiaris
toxicaria, called “riti’” in Sinhalese, “metavil’’in Tamil, belonging
to the same natural order (Urticacezx) as the Bread-fruit and Jak-
fruit trees.
The Veddas used to be an interesting race of forest haunting
nomads, but they are rapidly falling victims to civilization,
exchanging their ancient skill as bowmen and woodmen for a more
sordid if less precarious existence dwindling towards extinction.
They are chiefly to be found in the Province of Uva, but it is
possible to tramp through the Province from top to bottom
without seeing a sign of a Vedda. Occasionally persons are
paraded as Veddas, but when seen away from their natural
environment the effect must be pitiful rather than picturesque.
All the models were executed by a local modeller, Mr. R. G.
Andriesz.
CASE XXIV.
Ancient Bronzes.—On the top shelf a three-branched candela-
brum from Munisseram, presented by Hon. Mr. F. R. Saunders ;
below this a pair of gold-plated bronze curtain frames from Kotte
near Colombo, lent by P. E. Pieris, Esq., C.C.S. ; numerous minia-
ture bronze figures of gods and animals from Dondra Head ; bronze
lamps from Munisseram. On the bottom shelf the central object is
a large bronze Kothali or drinking goblet, with spout fashioned
after the manner of an elephant’s head and trunk, from Ratnapura,
lent by P. E. Pieris, Esq.; also an ancient bronze tripod from Kuru-
negala ; bronze hansas or sacred birds from Munisseram ; a heavy
bronze Contemplation Box with thirty compartments, some of
which contain a few coins and other offerings, presented by the
Royal Asiatic Society.
SOUTH VERANDAH.
Passing through the doorway at the side of the Bronze Case on
to the South Verandah, a Portugese cannon dredged up in the
Colombo Harbour in 1888 is an important relic of the Portuguese
occupation of thecountry. Here isalso exhibited a polished pillar
of Ceylon gneiss from the Mahara quarries employed in the con-
struction of the Colombo Breakwater, presented by John Kyle, Esq.
Returning through the Stone Gallery to the
WEST VERANDAH
a number of inscribed stones will be found, together with two
or three Dutch and Portuguese tombstones. ‘The work of collect-
ing and collating the numerous ancient inscriptions scattered over
“ny ahnd sons or)
‘NOTADO AO SHNIDINORVY AHL * SVACHA
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PETIGAMMANA PILLAR,
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( 23)
the Island was properly organized during the Governorship of
Sir William Gregory, when Dr. P. Goldschmidt was appointed
Archeological Commissioner to the Government of Ceylon in
1874. His reports were published as Sessional Papers from 1875
until his death in 1877. Dr. Goldschmidt was followed by Dr,
Edward Mitller, who compiled a valuable manual on “ Ancient
Inscriptions in Ceylon” (London, 1883), illustrated by a separate
quarto book of plates. Dr. Miiller left Ceylon in 1881, and was
succeeded, after an interval, as Archzological Commissioner by
Mr. H. C. P. Bell, C.C.S., under whose direction the work of
excavation, discovery, and transcription has been continued from
1890 to the present time.
The great slabs placed against the back wall of the verandah are
of interest on account of their antiquity and the characters
employed. The first one, propped up lengthwise on the ground,
is the oldest inscription that has been discovered at Anuradha-
pura, from the Ruanweli Dagaba.* It relates to the restoration
of certain temples during the reign of King Gaja Bahu (113-125
AS Ds Jo
The upright slab next to the Ruanweli Slab is known as the
Tissamaharama Slab, from Tissamaharama near Hambantota in
the Southern Province. It is almost completely preserved, and
according to Dr. Miiller “is the finest specimen we have of an
inscription of the fourth century A.D.”
Adjoining this slab is a narrow flattened stone with an inscrip-
tion on both faces. The inscription is headed on the obverse
side by a symbol of the sun and on the reverse by a crescent
representing the moon, the sun and moon being the usual royal
signs. It is a grant of land to a temple, and concludes (on the
reverse side) with a life-size figure of a crow in sunk relief. This
is the Petigammana Pillar found half buried in a garden within
a few miles of Gampola.t
Many of these inscribed pillars dating from the tenth century
bear, at the top, engravings of the sun and moon as symbols of
royalty [Rhys Davids] or eternity and, at the bottom, the dog and
crow as symbols of instability [Miiller] or meanness; anyone
violating the property of the priesthood renders himself liable te
the penalty of being re-born in the low condition of one of these
animals [Goldschmidt]. The translation of the Petigammana
inscription, according to Mr. Bell, ends with the usual curse:
“* Anyone who disputes this [grant will be born] a crow.”
* Miiller’s Inscriptions, No. 5, p. 27, and Plate 5.
t Miiller’s Inscriptions, No. 67, p. 43, and Plate 67.
{ H.C. P. Bell, Report on the Kegalla District, 1892, p. 79, with plate.
( 24)
In the middle of the outer side of the verandah there is a large
slab, the Dondra Slab,* recording the grant of land to the Temple
of Vishnu at Dondra Head in the fourteenth century. This slab
and the Dondra Pillart at the front outer corner of the verandah
are of particular interest on account of their association with Dondra
Head near Matara in the Southern Province, the most southerly
point of Ceylon. ‘Like Cape Comorin on the Continent of India,”
says Professor Rhys Davids (Indian Antiquary, I., 1872, p. 329),
“Dondra Head has always been a place of pilgrimage, and seems
to have derived its sanctity from its being the extreme southerly
point of land, where the known and firm earth ceases, and man —
looks out upon the ocean—the ever-moving, the impassable, the
infinite.”
‘Opposite to the Dondra Slab is the Mahakalattewa Pillar, from
the bund of a tank of that name six miles from Anuradhapura on
the road to Galkulam. It is remarkable for its perfect preser-
vation, not a single letter missing; the inscription is on all four
sides.t
Occasionally other symbols besides those mentioned above are
engraved upon the pillars, such as a cobra and a priest’s fan. The
latter occurs, for example, on the Kongollewa Pillar§ (placed near
to the Dondra Slab).
The stone slab bearing the Royal Arms of Portugal was found
at Menikkadawara in the Kegalla District by Mr. H. C, P. Bell
(Kegalla Report, 1892, p. 31, and plate).
Leaving now the West Verandah one crosses the Stone Gallery
to the
| NORTH VERANDAH
at the back of the Museum, where more tombstones, capitals,
- inscriptions, &c., will be met with. Here may be noted quaint
Portuguese tombstones||; a couple of Maldivian tombstones
characteristically carved in coralline limestone; a ‘‘ dressed stone”
with a Tamil inscription of the fifteenth century from the Kota-
gama vihare, found by Mr. Bell, who remarks upon the singularity
of discovering a Tamil inscription in the heart of a Sinhalese
district; thisis called the Kotagama Tamil Slab{ ; another stone
* Miiller’s Inscriptions, No. 163, p.71. First translated by Rhys Davids. Journ.
Ceylon R, Asiat. Soc., vol. V., 1870-1871, p. 25. :
+ Miiller’s Inscriptions, No. 159, p. 69. Rhys Davids, oc. cit., 1872, p. 57.
{ Miiller’s Inscriptions, No. 110, p. 55, with plates 110 A-110D.
§ Miiller’s Inscriptions, No. 112, p. 55. Kongollewa lies about two miles north
of Madawachchi in the North-Central Province.
|| A fully illustrated and historical account of these tombstones will be found
in.a paper on “ Portuguese Inscriptions in Ceylon,” by Mr. J. P. Lewis, C.C.S.,
to be published shortly in the Journ. Ceylon R, Asiat. Soc.
q H.C. P. Bell. Report, Kegalla District, 1892, pp. 68 and 85, with figure on
plate facing p. 72.
PZ obid aon OF,
EHUNUGALLA SLAB.
(Photograph by H, C. P. Bell, Esq.)
(Zo face page 24.
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( 3)
slab in a corner of the verandah bearing a short Sinhalese inscrip-
tion with representations of sun (an orb) and the moon (a crescent)
is called the Ehunugalla Slab*; it records a benefaction to a
monastery. Close to this stone is an ancient carved wooden door
from Dewanagala in the Kegaila District. The solid wings of this
door and the left jamb of its frame were found among the lumber
underneath the vihare. There are ten plain panels enclosed by
framework in high relief carved in a foliage scroll. The carving
of the jamb is described as follows: “ Between an outer beading
and inner splayed edge of lotus petals runs a long narrow panel
with gracefully intertwined double scroll of creeper, separating
four figures all different from each other. A space half moulded,
half panelled, in flower design, intervenes between it and the base
panel, in which is placed beside a tree an elephant with head and
right forefoot raised and curled trunk.’
The door, which was presented by F. H. Price, Esq., in 1890, was
reconstructed at the Colombo Museum. The cross beam on the
top with the drooping lotus capitals was brought from the Pinna-
wala Vihare.t
Next tothe wooden door are two carved wooden pillars mounted
as door posts. They belonged toa set of seven balcony pillars
found under the eaves of the porch of the Kumbukgama Vihare.§
Opposite to the wooden door are casts of two Peacock Pillars
from Anuradhapura. At the other end of the verandah there is
a large wooden rice trough or paddy pounder, in which the paddy
which has previously been trodden out of the corn by bullocks is
beatenand husked. Next to this is an old carved rice mortar from
Ratnapura, like those in constant use at the present day, in which
the rice is pounded into flour.
‘BUDDHA SHED.
In the palm-thatched shed in the grounds behind the Museum
further important stone antiquities are exhibited. Chief among
these is the large sedent Buddha found by Mr. Bell in the
jungle near the Nuwarawewa tank at Anuradhapura.| ‘ This
Buddha,” wrote Mr. Bell in 1890, “is admittedly the finest yet
brought to light at Anuradhapura. The wonderful sharpness
* H.C. P. Bell. Report, Kegalla District, 1592, p. 76, with figure on plate
facing p. 72.
+ Id., p. 49.
¢ H.C. P. Bell, op cit., p. 38.
§ Id., p. 22, with figure on plate.
|| H. C. P. Bell. First Report on Archzological Survey of Anuradhapura,
Sessional Papers, 1890, p. 4 [742]. It is known as the Zoluwila Buddha, from
the hamlet of Toluwila. which adjoins the Nuwarawewa buud at the spot where
the statue was found,
B 103-04
( 26 )
and depth of the features, the softness of expression, the symmetry
and repose of the body give the image a tout ensemble which
contrasts markedly with the stolid ‘figure-head’ appearance so
characteristic of these Buddhas in stone.”
In front of the Buddha there is a large moonstone of unique
and admirable design, embossed with wreaths, festoons, and
garlands, a pair of fabulous creatures (makaras) at the sides, and
a pair of two-fold representations of the Svi-patula or sacred
footprints near the base. This is called the Floral Moonstone of
Hanguranketa, and was presented to the Museum in 1894 by
Lady De Soysa.
The representations of the footprint of Buddha, called Syi-
patula, or Sri-pada, of which there are several in this shed carved
in stone, are of considerable interest. The best of them is that
which is known as the Koddaikeni Stone, a double Sri-pada (like
the rest) covered with symbols. The signs on the toes are called
swastika, the radiant emblem in the centre is the dharma-chakra,
in front of which are a couple of flags, at the side a fish-hook
behind a flower vase, a conch shell, a fan, a pair of fishes repre-
senting Pisces, one of the signs of the zodiac ; a complicated cry ptie
emblem occupies the centre of the heel, and on one side of this
opposite to the fishes are the trisul emblems. The exact interpre-
tation of the symbolism of this stone has not yet been attempted.”
Another curious relic is the limestone image representing
aman standing in the jaws of a monster, sent by Mr. C. A. Murray
from Tissamaharama in 1892. The moonstone and carved steps,
with janitors and terminals, which have been arranged at the
entrance to the shed, arrived here from Anuradhapura in the
years 1882 and 1884.t
The stone discs placed round the convex side of the large
Hanguranketa moonstone are described as mural ornaments
from Anuradhapura.
SIGIRIYA FRESCOES.
On the walls flanking the main staircase will be found copies
of the celebrated frescoes discovered in a cave or pocket of the
ancient rock fortress of Sigiriya near Dambulla. Sigiriya, the
lion rock, rises abruptly from the plains of the North-Central
Province to a height of about four hundred feet, with an area of
little more than an acre at the summit. It is said to have been
fortified by the Sinhalese parricide King Kasyapa, who ascended
the throne 475 A.D. and fled to the rock after having immured his
* Of. Memorandum by Messrs. J. P. Lewis and G. M. Fowler, with plate, in
Journ. Ceylon R. Asiat. Soc., vol. XI., 1889-1890, Proceedings, pp. bxii, lxiii.
+ Cf. Administration Reports of the Museum for 1882 (p. 110 D) and 1884(p,18 D).
‘gz abnd aang
‘VALIHIDIS HO MOOU HULYOd LNALIONY
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father King Dhatu Sen, whose capital was Anuradhapura. Kasvapa
made Sigiriya his capital, and took refuge there for eighteen years.
On the western face of the rock chambers have been scooped
out, and in one of these, 160 feet from the ground, protected from
sun and rain, ancient frescoes were painted upon stucco plastered
upon the smooth surface, and still remain in an excellent state
of preservation. The ancient approaches to the summit and to
the chambers having fallen into decay, the rock once more
became nearly inaccessible and, according to local tradition, the
haunt of “yakku” ordemons. It was however tackled by more than
one adventurous climber during the latter half of last century,
and in June, 1889, Mr. Alex. Murray of the Public Works Depart-
ment succeeded in reaching the pocket containing the frescoes
and in making the tracings of them, which he coloured as nearly as
possible like the originals. He hasleftit upon record that the work
of copying took him from sunrise to sunset every day for a week
lying at full length on his back. A Buddhist priest who visited
the chamber gave it as his opinion that the pictures must be the
portraits of some of King Kasyapa’s queens. The portraits are
arranged singly and in couples, the latter representing a maid
offering the sacred lotus on a tray to her mistress.
During the last ten years excavations have been carried on at
Sigiriya under the direction of the Archzological Commissioner,
and fresh copies of the frescoes have been made under his super-
vision in oil colours, but these have not yet been exhi ited in the
Museum.
PALS wl;
ZOOLOGICAL COLLECTIONS.
In the grounds at the back of the Museum a few live animals
indigenous in Ceylon are placed on exhibition in temporary
shelters. The mammals include a leopard, bears, a tiger cat (Felis
viverrina), a jackal, palm or toddy cats, civet cats, porcupines, a
bandicoot rat, mouse deer, hog deer, &c. The birdsare represented
by a pelican ibis presented by Her Excellency Lady Blake, a
pelican, purple herons, Malay bitterns, India koels, Brahminy kites,
scops owl,.and an Alexandrine paroquet. A young rufous-bellied
hawk-eagle has been loaned by His Excellency the Governor
(Sir Henry Blake, G.C.M.G., F.Z.8.). A small tank contains some
monitors or water lizards, called “kabaragoya” in Sinhalese, and
in a small cage there is a chameleon from Chilaw.
On the other side of the block of buildings in which the Mineral
Gallery is situated (see plan) there are two sheds containing
respectively, the skeleton of a sperm whale or cachalot (Physeter
macrocephalus) and of the whalebone whale (Balanoplera indica).
The carcases of whalesare stranded from time to time on the shores
of Ceylon. Some of them seem to have met their death at the
hands of whalers and to have drifted by gale and current to
Ceylon. A whalebone whale was washed ashore in Weligam Bay
in August, 1884,such bones as were recovered being placed on the
front verandah of the Natural History Gallery upstairs; another
carcase drifted ashore at Ambalangoda in September, 1894; the
almost complete skeleton was brought to the Museum, and is the
one now lying in the large cadjan shed; it measured 65 feet in
length. This species of whales has the distinction of being the
largest of all known animals, living or extinct.
A spermaceti whale or cachalot stranded on the south coast of
Mannar in September, 1889; its remains are now exhibited in the
smaller shed, with the exception of the lower jaw, which dropped
into the sea while the work of salvage was proceeding and was
lost. Another carcase arrived at Athuruwila near Bentota in June,
1904, in a high state of decomposition ; the lower jaw, which alone
carries the functional teeth, was missing. In November, 1904,
another decomposed sperm whale minus the lower jaw was
stranded at Mount Lavinia. The teeth of the upper jaw of the
_ sperm whale are vestigial structures imbedded in the gum.
At the foot of the main staircase leading to the upper floor of
the Museum there may be seen a small glass case containing
GROUP OF § 3UR loninyny se :
F SAMBUR (Cervus unicolor) AND WILD BOAR (Sus cristutus)
(Vo face puge 28.
LoAeNae uss © Gite Suk, EE ai |
Be es ee
[62 chnd sanf of
“(srumd snaaypdouiiog) SKAMNOW ANV ‘(supind sijag) GAVdOAT ‘(sivv snita)) AAI GALLOdS
C2)
living insects, leaf insects of the genus Phylliwm, remarkable for
their general resemblance to the leaves of the guava, on which
they are fed. The males are smaller than the females, less
numerous, and carry two long feelers or antenne, held backwards,
nearly as long as the body. The females lay their eggs, which
resemble seeds, freely, and the young hatch out without difficulty-
The Natural History Collection comprises representatives of the
fishes, amphibia, reptiles, birds, mammals, insects, crustacea, and
mollusea occurring within the zoological province of Ceylon.
The centre of the gallery is occupied by a jungle scene
containing sambur deer, commonly known as elk, a wild boar,
a crocodile, peacock, and several smaller birds. Most of the
specimens were obtained from the Hambantota District in the
Southern Province, where all of the species are to be found.
Opposite to this group, over the head of the staircase, is a group
of spotted deer, leopard, and monkeys, the material for which was
also obtained from the same locality. The monkeys on the
tree overhead belong to the common low-country species of the
Maritime Provinces, Semnopithecus priamus.
BIRDS.
The birds of Ceylon comprise large numbers of migratory
species, which visit the Island during the north-east monsoon, but
do not breed here; the chief bird of this kind is the Flamingo.
Then there are numerous resident species, which are indigenous to
Ceylon, but occur also in the Indian Peninsula and elsewhere ; the
chief bird of this kind is the Peafowl. There are no fewer than
forty-nine endemic species, which are peculiar to Ceylon, not being
found beyond the confines of the Island; the chief bird of this kind
is the Ceylon Jungle-fowl. Lastly, there are a few occasional
visitors, which do not come regularly, putting ashore here through
stress of weather ; the chief bird of this kind is the Frigate-bird.
BIRD CASE I.
Turning to the left (east) from the staircase the first bird case,
beginning from the top shelf in front, contains representatives of
the Frogmouths (Podargide); the Trogons (Trogonide); the
Cuckoos, Koels, Malkohas, and Coucals (Cuculide) ; Paroquetsand
Loriquets (Psittacidee); Owls (Strigide). The Ceylonese Frog-
mouth (Batrachostomus moniliger), which also occurs in Travan-
core, is a remarkable bird on account of the oddness of its gape,
curiously resembling the mouth of a frog. Of the cuckoos, the
genus Swrniculus, represented by the species S. /ugubris, the
Drongo Cuckoo, is remarkable for its extraordinary resemblance
to a Drongo or King Crow (Dicrurus). It is said to lay its eggs
( 30 )
in the nests of King Crows, and the latter have been actually
observed in the act of feeding a young Surniculus.* This there-
fore appears to be an example of natural mimicry.
The Red-faced Malkoha (Phenicophaés pyrrhocephalus) is pecu-
liar to Ceylon,t as is also the Ceylonese Coucal (Centropus
chlororhynchus). Both of these species inhabit dense and damp
forests. The Common Coucal or Crow-pheasant (Centropus
sinensis) is known locally as the Jungle Crow. It is a familiar
bird in the low-country jungles and in the vicinity of villages, its
chestnut-coloured wings offering a handsome contrast to the glossy
black body. It is a ground-feeding bird, eating insects, lizards,
and small snakes.
The Indian Koel (Hudynamis honorata) is another familiar
Indo-Ceylonese bird. In the breeding season, from March till
July, its cry of ku-il kw-il, increasing in intensity and ascending
in the scale, is to be heard in almost every grove [Blanford, dc.,
p. 229]. In Ceylon it is known to Europeans as the Brain-fever
bird, on account of the persistency of its cry. It feeds on fruit,
and, like the cuckoos, is parasitic in its nesting habits, laying its
eggs in May and June in the nests of crows, generally the Indian
or Gray Crow (Corvus splendens), less frequently in those of the
Black Crow (Corvus macrorhynchus). The crows bring up the
koels, which at times eject the young crows from the nest after
they have been hatched [Blanford]. The male is black through-
out; the female is spotted with white.
The principal parrot of Ceylon is sometimes called the Alexan-
drine Paroquet (Palwornis eupatria), coloured green, with (in the
male) a broad rose-pink collar round the nape. Thelittle Ceylonese
Loriquet (Loriculus indicus) is peculiar to Ceylon. The parrot
which is commonly used as a cage bird by the natives of Ceylon
is the Rose-ringed Paroquet (Palwornis torquatus).
Of the owls which are shown in the bottom shelf, the Ceylon
Bay Owl (Photodilus assimilis), peculiar to Ceylon, is to be noted
on account of its rarity, being found only in the hills round
Kandy. The Demon bird, or “Ulama” of Ceylon, so-called on
account of its dreadful moaning hoot, is commonly identified with
Huhua nepalensis (= Bubo nipalensis) and also with Syrnium
indrani.
In the other half of this case, commencing at the top, are shown
the Barbets (Capitonide), of which the Yellow-fronted Barbet
(Cyanops flavifrons) and the pretty little Ceylon Barbet (Yantho-
lema rubricapilla) are peculiar to the Island; Indian Rollers
* Blanford, W.T. Birds of India, 1895, vol. ILI. p. 224.
+ The distribution of the different species of birds is indivated in red upon the
small maps plaved below the specimens.
Ge
(Coraciade) ;* Bee-eaters (Meropide) ; Kingfishers (Alcedinide) ;
Hornbillst (Bucerotids), two species, one of which (Lophoceros
gingalensis) is peculiar ; Hoopoes (Upupide) ; Swifts (Cypselide),
one of which, the Edible-nest Swiftlet (Collocalia fuciphaga),
builds the well-known edible nests in caves, small cups made of
grass, moss, and feathers cemented together by inspissated saliva
[Blanford, /.c., p. 177] ; Nightjars (Caprimulgide), which lay their
eggs on the ground without any nest.
BIRD CASE II.
The second case alongside the first contains some of the Doves
and Pigeons (Columbide), including Turtur risorius, the Ring-
dove, and Columba intermedia, the Indian Rock Pigeon, from
which all the breeds of domestic pigeons peculiar to India are
derived. The Galline or true game birds of Ceylon belong to the
Pheasant family (Phasianide). First in order and importance
comes the Peafowl, referred to and exhibited elsewhere ; then
the endemic Ceylon Jungle-fow!l (Gallus lafayetti), characterized
by its peculiar call and by the yellow patch in the centre of the
comb of the male; the endemic Ceylon Spur-fowl (Galloperdix
bicalcarata), which only occurs in the southern half of the Island ;
finally the Quails and Partridges. The Gray Partridge (Francoli-
nus pondicerianus) is common in India, but is only found in the
northern half of Ceylon and in the small islands (Delft, Iranativu,
&e.) off the Jaffna Peninsula. The Painted Partridge (F. pictus),
unrepresented in the Museum collection, is another Indo-
Ceylonese bird localized in Ceylon to the highlands or patanas
between Nuwara Eliya and Badulla.
The Three-toed Quails, represented here by two female Bustard
Quails (Turniz pugnaz), belong to a separate order, Hemipodii,
and family, Turnicide. The females are larger and more highly
coloured than the males, and “the ordinary conduct of the sexes
during the period of incubation is reversed, for the male alone
sits on the eggs and tends the young brood, whilst the females
wander about, uttering a purring call that serves as a challenge,
and fight each other” [Blanford, Faun. Ind. Birds, vol. IV., p. 150}.
Of the Rails, Crakes, Moorhens, and Watercocks (Rallide),
Baillon’s Crake (Porzana pusilla) is to be noted for its rarity in
Ceylon ; the Moorhen (Gallinula chloropus) ranges from Great
* The Indian Roller (Coracias indica) is common at Jaffna, Anuradhapura, &c.,
and is known to Europeans as the Low-country Jay, but it is a Picarian bird
related to the Bee-eaters and Kingfishers, not a Passerine bird of the Crow
family. It is fond of perching on telegraph wires.
jy The Hornbills of the Ethiopian, Oriental, and Papuan regions are the
representatives of the Toucans (Rhamphastide) of South America, and are
sometimes popularly confounded with the latter.
( 32 )
Britain through Europe, Asia,and Africa, and occurs in the tanks
of Ceylon, though rare; finally, the Purple Moorhen (Porphyrio
poliocephalus) is a handsome Indo-Ceylonese bird common in
parts of the low-country.
The bottom shelf in front commences the series of Limicoline
birds, Plovers and Snipes and their allies. The Stone Curlew
(Gdicnemus scolopax) is a north-east migrant to Ceylon, the Great
Stone Plover (Hsacus recurvirostris) is a resident shore bird ; both
belong'to one family, the Gidienemidz. The Crab Plover (Dromas
ardeola), another shore bird, which also breeds in Ceylon, is the
sole type of the family Dromadide.
In the reverse half of this case, commencing from the top, will
be found Courier Plovers and Swallow Plovers, shore birds of the
family Glareolide ; Water Pheasants (Hydrophasianus chirurgus),
Parridz, a common low-country bird: the Turnstone, Lapwings,
Plovers, Oyster-catcher, Stilt,* Avocet,t Curlew, Whimbrel,
Sandpipers,t{ and Stints, all shore birds and waders belonging tothe
family Charadriide; Woodcock (Scolopax rusticula), arare migrant,
and Snipes (Scolopacidz), of which the best known are the
Pintail Snipe (Gallinago stenura), a north-east migrant common
from September to April, and the Painted Snipe (Rostratula
capensis), a resident of the low country, widely distributed in
South Asia and Africa.
Many of the shore birds, like the sea birds, gulls, and terns, have
a very wide distribution. The circum-littoral range of the
Turnstone (Strepsilas interpres) throughout both hemispheres is
remarkable. The Oyster-catcher (Hematopus ostralegus) is con-
fined to the Old World. The range of the Gray Plover (Squatarola
helvetica) is world-wide ; it breeds in the far north and isa winter *
visitor to India, Ceylon, and Burma [Blanford}.
BIRD CASES III. AND IV.
The next couple of bird cases contain the one a group of Acci-
pitrine birds,§ Eagles, Hawks, Falcons, and Kestrels ; the other a
group of Peafowl (Pavo cristatus). This latter is the principal
game bird of Ceylon, and in fact the most stately bird in the
Island. A frequent ornament in parks and gardens in Europe, it
lives here in its native haunts. It feeds and nests upon the
ground, but roosts in the topmost branches of trees, whence it
* Shown in another case (see below).
+ An occasional migrant, not in the Museum collection.
t The Sandpipers are well known locally by the colloquial Sinhalese name
“ siri-biri”; they are also sometimes called Snippets, because they somewhat re-
semble Snipes in their habits.
§ Other Accipitrine birds, Harriers and Falcons, are shown in another case (see
below).
AFOWL.
PE
face page 32.
u
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Oo
wa
surveys the country round and is consequently difficult to
approach. A male in perfect plumage is shown perched upon a
tree ; a dun-coloured female and a partridge-like young are placed
on the floor of the case.
BIRD CASE V.
Passing now towards the other side of the gallery the remaining
bird cases may be inspected, commencing with the case next to
the stairs facing the Sambur and Boar Group. Here are some
of the Passerine birds. The order Passeres comprises about one-
third of the entire avian fauna of Ceylon. The front half of this
case contains Orioles (Oriolidx), represented by the brilliant
yellow-bodied black-headed Oriolus melanocephalus ; Mynas and
Starlings (Sturnide), of which three are peculiar, namely, the
Ceylon Myna, known to ornithologists as the Ceylon Grackle*
(Hulabes ptilogenys), the common Ceylonese Myna (Acridotheres
melanosternus), which is used extensively asa cage bird by the
boutique-keepers and other residents, and the White-headed Myna
or Starling (Sturnornis senex), an inhabitant of the mountain
forests of Ceylon ; Thrushes and Black-birds (Turdidez), of which
the Ceylon Black-bird (Merula kinnis?), a resident of the forests
above 2,500 feet, the Butf-breasted Ceylon Thrush (Oveocincla
imbricata), and the Spotted Thrush (Oreocincla spiloptera) are
peculiar ; Flycatchers (Muscicapidz), of which the Ceylonese Blue
Flycatcher (Stoparola sordida) is endemic, while the Paradise
Flycatcher (Terpsiphone paradisz), locally known as the Ceylon
Bird of Paradise, is an exquisite and highly characteristic Indo-
Ceylonese resident. The Paradise Flycatcher is not uncommon,
though it is rarely seen about Colombo. The male bird undergoes
remarkable changes of plumage, which are illustrated by the
specimens exhibited.
After the autumn moult of the second year the male has the whole head
and crest glossy black [throat brown, breast ashy, belly white], and the whole
upper plumage rich chestnut; the median tail feathers grow toa great length,
and are retained till May or June, when they are cast. After the autumn
moult of the third year the chestnut plumage is again assumed, and also the
long median tail feathers, but the whole lower plumage from the throat down-
wards is pure white, the breast being sharply demarcated from the black
throat. After this moult a gradual transition to the white upper plumage
takes place, the wings and tail being the first parts to be affected, but the
change toa complete white plumage is not affected till the moult of the
fourth autumn. After this moult the male bird is fully adult, and perma-
nently retains the white plumage ; the head, neck, and crest are glossy bluish
black ; the whole body plumage white.t
* This species is placed by Mr. E. W. Oates (Fauna Brit, Ind., Birds, vol. I..
p. 513) in a separate family, the Eulabetide,
t Oates, E. W. Fauna Brit. Ind., Birds, vol. IT., 1890, p. 46.
F 105-04
((, 34 )
Just as the Indo-Ceylonese Peafowl is replaced in Burma by a
distinet species (Pavo muticus) which ranges southwards to Java,
so the Paradise Flycatcher of India and Ceylon is represented in
Burma by an allied species (Terpsiphone affinis).
The robins are closely related to the flycatchers on the one hand
and to the thrushes on the other. The Black Robin (Thamnobia
Julicata) and the Magpie Robin (Copsychus saularis) are, next to
the crows, the sparrows, and the babblers, the commonest birds
in Colombo and throughout the Island; the Long-tailed Robin,
known to ornithologists by its Hindustani name ‘‘ Shama,” is a shy
jungle-bird.
In the reverse half of the case are shown Weaver birds and
Munia Finches (Ploceidz), the Hill Munia (Uroloncha kelaartt)
being peculiar; Sparrows (Fringillide), the House Sparrow
(Passer domesticus) ranging from Great Britain to South Asia;
Swallows (Hirundinidz) ; Wagtails and Pipits (Motacillide) : the
Gray-headed Wagtail (Motacilla borealis) ranges all over Europe,
Asia, and North Africa, and may be seen on the Galle Face Parade
during the north-east season from September to May; Larks
(Alaudide); Sun birds* (Nectariniide) and Flower-peckers
(Diceide), the former with long bills, the latter with short bills,
both families being distinguished from all other Passerine birds
by the serration of both mandibles of the beak ; the Indian Pitta
or Ground Thrush (Pittide), a characteristic north-east migrant.
Finally, the bottom shelf of the case contains the Woodpeckersft
(Picide), a very distinct family of birds well represented in
Ceylon. The type skins of Legge’s Woodpecker (Brachypternus
intermedius) were presented by Sir W. H. Gregory. This variety
is believed to be a hybrid between the Golden-backed Woodpecker
(B. aurantius), and the common Red-backed Woodpecker (B.
erythronotus), which is peculiar to Ceylon. Layard’s Wood-
pecker (Chrysocolaptes stricklandt) is also confined to Ceylon.
BIRD CASE VI.
The case adjoining the one just described, commencing from
the top of the reverse side, contains more Passerine birds, namely,
the Crows, Jays, and Titmice (Corvide), the Ceylonese Jay (Cissa
ornata), coloured chestnut and blue, being endemic, This bird is
sometimes known as the Ceylon Magpie. It is a forest bird of
ps, habits, fesning a good deal on the ground [Oates].
* The Sun Hae are the representatives in the Old World of the eid a birds
of the New World. and like the latter can poise themselves on the wing while
extracting nectar from flowers. They are frequently seen in Colombo feeding
npon the Hibiscus flowers.
+ The Woodpeckers are Picarian birds, not Passerine,
( 35)
The Babblers (Crateropodide) of Ceylon are remarkable for the
large number of endemic species, namely, the Ceylonese Rufous
Babbler (Crateropus rufescens), the Ashy-headed Babbler (C.
cinereifrons), the Ceylonese Scimitar Babbler (Pomatorhinus
melanurus), the Ceylon Yellow-eyed, black-billed Babbler
(Pyctorhis nasalis), the Brown-capped or Quaker Babbler
(Pellorneum fuscicapillum) occurring between 5,000 and 6,000 feet
elevation, the Ceylon Black-fronted Wren Babbler (Rhopocichla
nigrifrons), the Ceylon Arrenga or Whistling Thrush (Arrenga
blight), rare at 4,000 feet, the Ceylon Shortwing or Ant Thrush
(Hlaphrornis pallisert), and the Ceylon White-eye (Zosterops
ceylonensis) occurring above 1,500 feet. |
The Bulbuls also belong to the Crateropodidx, of which they
form a sub-family, Brachypodine. They are shown at the bottom
and continued on the top shelf of the front side of the case. The
Yellow-eared Bulbul (Kelaartia penicillata) is endemic.
Next follow the Drongos or King Crows (Dicruride), so called
because of their remarkable habit of persecuting the crows, which
are double their size. They pursue and chivvy the crows on the
wing with no other apparent object than pure love of mischief ;
they also frequently perch upon the backs of cattle. The species
which is especially tyrannical with regard to the crows is the
Black Drongo (Dicrurus ater). Another exhibited species (D.
leucopygialis) is confined to Ceylon,while the Racket-tailed Drongo
(Dissemurus paradiseus) is distinguished by the great elongation
of the lateral tail feathers with their spatulate tips.
The Warblers (Sylviidee) represent another extensive family of
small birds, the most remarkable of those which occur in Ceylon
being the Indian Tailor bird (Orthotomus sutorius), a nest of
which with the leaf-edges sewn together is exhibited.
Lastly, the Shrikes and Minivets (Laniide) complete the display
in thiscase. The Orange Minivet (Pericrocotus flammeus) bears a
strong resemblance to the oriole in the general colour-pattern of
the plumage.
BIRD CASES VII. AND VIII.
These cases contain groups of birds arranged according to habits
and distribution, but without reference to their position in classi-
fication. One of them consists of a selection of birds from the
Vanni District of the Northern Province, that portion of the
Province which stretches between the settlements of Mullaittivu
and Vavuniya. The other isa group of Waders and Divers, com-
prising Flamingoes, Stilts, Herons, Darter, &c.* The Flamingo
* The Cormorants (exhibited in the adjoining case and also in Bird Vase XII., see
below) of the genus Phalacrocorar are allied to the Darters, and form with the
latter the family Phalacrocoracid, diving and fishing birds,
C865
(Phenicopterus voseus) is a north-east migrant to Ceylon frequent-
ing the salt lakes of the Hambantota and Mullaittivu Districts, Lt
is essentially gregarious, living and breeding in large colonies. The
movements of a flock are remarkably concerted, and at the least
alarm they rise with one consent like a pink cloud across the
horizon. The three specimens exhibited are females in different
stages of plumage.
The Darter (Plotus melanogaster) is sometimes called the Snake
bird, on account of its long serpentine neck, which is persistently
kinked. The method of feeding was described by Mr. W. A.
Forbes as follows* :—‘‘The darters feed entirely under water,
Swimming with its wings half-expanded, though locomotion is
effected entirely by the feet, the bird pursues its prey witha peculiar
darting or jerky action of the head and neck, which may be com-
pared to that ofa man poising a spear or harpoon before throwing
it. Arrived within striking distance the darter suddenly trans-
fixes the fish on the tip of its beak with marvellous dexterity,
and then immediately comes to the surface, where the fish is
shaken off the beak, thrown upwards, and swallowed, usually head
first.” When swimming at the surface the body is submerged, only
the head and neck projecting above the water. The darters are
common about the tanks of Ceylon. When resting ona tree the
wings are held expanded as shown. The stomach of this bird is
provided with a dense hairy plug or sieve guarding the entrance to
the small intestine.
BIRD CASES IX. - XII.
The remaining birds which are placed on exhibition are shown
in the recess facing the Sambur and Boar Group. In Case IX.are
the birds of largest bulk occurring in the Island, gigantesque Storks
and Herons, Spoonbills, and Pelican Ibis or Painted Stork. Case X.
contains Harriers (Falconidz) and Bitterns (Ardeidw, the Heron
family). The Malay Bittern (Gorsachius melanolophus) is a north-
east migrant to Ceylon, arriving towards the end of October and
beginning of November, and fugitive specimens are frequently
captured about this time in Colombo, sometimes landing in the
streets of the Fort and in the ball-room of Queen’s House.
Case XI. contains Gannets or Boobies (Sulide), oceanic birds
sometimes taken here, and Ibis (Ibidide), tank birds. Finally,
Case XII. contains Ducks and Teals (Anatide), Terns and Gulls
(Laride), Frigate birds (Fregatide), Cormorants and Darters
(Phalacrocoracidie), Pelicans (Pelecanide).
* W. A. Forbes, On some points on the Anatomy of the Indian Darter (Plotus
melanogaster) and on the Mechanism of the Neck in the Darters (Plotus), in con-
nection with their habits, P. Zool. Soc., London, 1882, pp. 208-212.
FLAMINGOES AND OTHER WADERS.
[To face page 36,
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( 37)
MAMMALS.
Besides the mammals living in the grounds and those which
have been mentioned above in connection with the groups in the
centre of the gallery, further examples are shown in the western
alcove. Several of the eighty species of mammals recorded from
Ceylon possess insular characteristics ; even such a large creature
as the sambur is said to differ in some respects from its Indian
co-type, but there is nothing like the same degree of endemicity
among the mammals as has been remarked for the birds. As
distinctive Ceylonese mammals may be mentioned the Golden
Paradoxure or Palm Civet (Paradoxurus aureus) and the Ruddy
Mungoose (Herpestes smithi), both of which are called by the same
Sinhalese name “ Hotambuwa.”
There are two principal kinds of monkeys in Ceylon, called res-
pectively in the native language “ Rilawa”’ and “ Wandura,” referred
to by Knox in the anglicized terms “ Rillows” and ‘‘ Wanderows.” .
The former are the Macaques (Macacus pileatus), with cheek
pouches ; the latter are the Langurs, comprising several species of
the genus Semnopithecus, monkeys destitute of cheek pouches.
Troops of “ Rillows” and “ Wanderows” may sometimes be seen on
the same tree, but as arule they keep to themselves.
There are three distinct species of ‘‘Wanderows” in Ceylon. The
commonestis the Madras Langur or Crested Monkey (Semnopithe-
cus priamus), which frequents the low-lying forests of the dry
maritime districts of the North, Kast,and South. In addition to the
crest of hair on the head this monkey is further distinguished by
the fringe of long black hairs of the eyebrows, known as the
supra-orbital fringe.
The Purple-faced Monkey (S. cephalopterus) is without the crest
and fringe ; it inhabits the damp forests of the West at low and
moderate elevations up to about 1,000 feet.
Finally, the Bear Monkey (S. wrsinus), described as endemic,
is the monkey of the mountains, occurring in the country round
Nuwara Eliya. Itis closely related to S. cephalopterus,* of which
it may be a hill variety, and from which it is distinguished by its
longer and denser fur.
The remarkable Prosimian family of the Lemurs, whose head-
quarters are in Madagascar, is represented in Ceylon by a single
species, Loris gracilis, a small tailless, large-eyed, nocturnal,
arboreal creature of retiring habits, sometimes called the Ceylon
Sloth. It is omnivorous, feeding upon young leaves, insects,
* A young live Bear Monkey presented by J. Spearman Armstrong, Esq., from
Kotagala, and a Purple-faced Monkey from Horana, are exhibited in the grounds
at the back of the Museum.
( 38 )
spiders, birds’ eggs, birds, and lizards. It will also eat plantains
and boiled rice, and will drink milk, but is not easy to keep alive
in captivity unless taken young and reared with great care.
The skeleton of an elephant shot by H. W. Varian, Esq., said to
be the largest recorded from Ceylon, is remarkable for the small
size of the tusks, which are reduced to meretushes. The skull of
a large tusker is, however, shown below. Tuskers are rare in
Ceylon, and are believed to be the descendants of imported Indian
elephants. Another elephant skull in section is lying on the floor
next to the skeleton of the sambur. There are also skeletons of
the wild buffalo, wild boar, and the bear. Of the other stuffed
animals, a pair of large leopards, the bear, the otter, and the
pangolin may be noted specially. The Indian Pangolin or Scaly
Ant-eater (Wanis pentadactyla)is one of the most curious mammals
found in Ceylon. It is a nocturnal burrowing animal not often
seen ; its jaws are destitute of teeth (Edentata), and its tongue is
exceedingly long and vermiform, adapted for penetrating into the
burrows of termites or white ants, upon which it feeds. Its
scales are sometimes employed for making imitation tortoise-
shell combs. One such comb, presented by H. J. V. Ekanayake.
Esq., of Balapitiya, is exhibited. The Indian Pangolin is repre-
sented in Burma by an allied species UW. javanica, which ranges
through the Malay Peninsula, Sumatra, Java, Borneo, and Celebes.-
There is also a Chinese Pangolin (MW. aurita) in Nepal, Assam,
Southern China, and Formosa. Several species of the same genus
occur in Africa.
There are three sorts of lying mammals in Ceylon, namely, the
Fruit Bats or Flying Foxes (Pteropodide), the small Insectivorous
Bats (Microchiroptera), and the Flying Squirrel, Pteromys oral
(Rodentia Sciuridze). The Insectivorous Bats comprise the Leaf-
nosed Bats (Rhinolophide); the Vampire Bats (Nycteridz), which
feed upon frogs, rats, and smaller bats, as well as insects; the
Pipistrelles (Vespertilionide) ; and the Sheath-tailed Bats (Embal-
lonuride). The Flying Squirrel has approximately the same
general distribution throughout India, Burma, and Ceylon as the
Flying Fox (Pteropus medius), inhabits the same districts, and is
also frugivorous and nocturnal. It is not however gregarious,
and does not suspend itself head downwards, as do the fruit bats
and other bats, but rests in the ordinary attitudes of arboreal
mammals, :
The aquatic mammalia (Whales, Dolphins, Porpoises, and
Dugongs) are represented by an excellent set of dugongs, male.
female, and young,andaskeleton. The female specimen exhibited
is 10 ft. long ; it wascaptured at Kayts near Jaffna. The Dugong
¥
Te
KRAALIN(
ANT
LEPH
1D)
(10 Face page 38
( 39)
(Halicore dugong) is a gentle creature feeding on seaweeds ; it
allows itself to be handled and killed without resistance. It occurs
off the north-west coast of Ceylon near Jaffna and Mannar, and
ranges from East Africa to Australia. Kelaart, one of the pioneer
naturalists of Ceylon, says that he saw shoals of them on the coast
of Arippu during the Ceylon pear! fisheries of 1835 and 1836, but
they are now scarce. The order Sirenia, to which the dugong
belongs, is represented in tropical America by the manatee. The
tusks of the dugong are the two upper incisors, which, with two
or three molars on each side of both jaws, are the only teeth found
in adults.
On the top of the Dugong Case is the skeleton of a rare cetacean,
Pseudorca crassidens, the Lincolnshire Killer, prepared from a
specimen caught at Moratuwa in 1891, the first recorded from
Indian seas. It is said to feed on cuttle fish, whereas the true
grampus or killer attacks and kills the largest whales. The
Lincolnshire Killer owes its common name to the circumstance
that its skeleton was first discovered in a Lincolnshire fen.
Besides this skeleton there is a stuffed specimen of a dolphin
caught at Negombo in 1883. Its identification is somewhat |
uncertain, but it appears to be closely similar to Elliot’s Dolphin
(Steno perniger).
The smaller mammals of Ceylon (apart from the bats and lemur)
comprise many species of the Rodent order (Squirrels, Rats, Mice,
Hares, and Poreupines). The little squirrel which is a familiar
figure on the trunks and branches of trees in Colombo is called
Sciurus palmarum. The larger tree-squirrel of the low-country
jungles in dry districts is Sciurus macrurus, locally known as the
Rock Squirrel or “* Danduléna.” The largest rat is the Bandicoot or
Pig Rat (Nesocia bandicota), found in all parts of the Island from the
sea-level to Nuwara Eliya. It exceeds a foot in length, exclusive of
the tail. Next to this in size and interest comes the Gerbille or
Antelope Rat (Gerbillus indicus), afield rat. The common House Rat
and the Mouse have been introduced here as to all other parts of
the world. The Musk Rat is more properly called the Musk Shrew
(Crocidura murina) ; it is nota Rodent, but belongs to the family
of Shrews (Soricidz) in the order Insectivora, It is common in
bungalows, outhouses, and compounds in Colombo, and often it
appears in the roads at nightfall. It is pale gray in colour, utters
a characteristic squeak, and hasa long, tapering snout. Finally, the
Black-naped Hare (Lepus nigricollis), the Chevrotain or Mouse
Deer (Tragulus meminna), and the Muntjac or Rib-faced Barking
Deer (Cervulus muntjac), commonly called the Red Deer, must be
mentioned.
( 40 )
REPTILES AND AMPHIBIA.
The collection of reptiles and amphibia is contained under
glass shades over the table cases. The largest reptiles are the Cro-
codiles, Monitors, and the Python. There are two species of cro-
codiles in Ceylon, the Tank Crocodile (Cvocodilus palustris), with
a comparatively short snout, and the River Crocodile (C. porosus),
with a longer and narrower snout. The former is represented by
the skull of a large specimen from the Minneri tank,* the latter
by a young stuffed specimen in the Sambur and Boar Group.
There are also two species of monitors, the large Water Lizard or
“ Kabaragoya ” (Varanus salvator), of which a skeleton is exhibi-
ted in the gallery and some live specimens in the grounds ; and
secondly, the Land Monitor (V. bengalensis), a smaller species,
which lays its eggs in the nests of termites.
Some eighty-one species of snakes have been recorded from
Ceylon, including twenty-six seasnakes (Hydrophide). The
latter are all poisonous, but of the land snakes only six or seven
are poisonous, the most deadly being the Cobra (Naa tripudians),
the Tic Polonga or Russell’s Viper (Vipera russelli), and the
Bungarums or Kraits (Bungarus ceylonicus and B. cowruleus).t
Examples of these are shown over the second table case to the
left of the staircase. The Crotaline or Pit Vipers, so called on
account of the existence of a deep pit of unknown significance on
each side of the snout between the eyes and the nostrils, are not
fatal to man. They are represented in Ceylon by the “ Karawala”’
(Ancistrodon hypnale) and the Green Polonga (Trimeresurus
trigonocephalus).
There is a very common non-poisonous snake which mimics
the dreaded Bungarus ceylonicus in its scheme of coloration,
namely, white transverse bands upon a dark ground colour, and
is sometimes mistaken for it when seen at a distance or when
examined casually. This is Lycodon aulicus, a snake which is
frequently found trespassing in bungalows in Colombo, The
Bungarus occurs chiefly up-country in the country round
Peradeniya, Dimbula, Balangoda, and elsewhere.
A large Python molurus is mounted in the east alcove; some
Hydrophide are shown over the third case; the snakes on the
fifth case include a large Green Polonga and a large Tic Polonga,
somewhat faded ; over the sixth case are the Whip snakes (Dry-
ophis); the “Pol-mal Karawala” (Chrysopelea ornata), which when
fresh shows bright red spots along the back, reputed poisonous,
but in reality harmless and of gentle disposition ; the fresh-water
* Placed on a bench in the east alcove.
+ B. cwruleus has only been recorded from Jaffna,
G
Colour variations of Rhinophis blythii.
UROPELTID,
105-04.
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Head from Tail end. Abnornial
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Rhinophis trevelyanus,
Head-shields of Uvopeltis grandis.
UROPELTID A.
(PABh)
or estuarine snake (Cerberus rhynchops,) which, like another
fresh-water snake, Topidonotus plumbicolor, has a viperine look
but is quite harmless; anda rare fresh-water snake, Gerardia
prevostiana, from the Kelani river, caught twenty yearsago. Over
the seventh case will be found a large specimen of the common
Rat-snake (Zamenis mucosus); another snake which attains a
length of five feet and a wide girth is Dipsas forstenii, of which
only a half-sized example is shown. This snake is represented in
Ceylon by two varieties, the typical variety, brown with angular
oblique black bars, and the red variety, uniformly rich reddish-
chocolate above without black bars, paler roseate flush below; a
fine example obtained from Nambapana in September, 1904, is
shown. It is called locally the Lé-polonga (Blood-polonga), and
is reputed poisonous.
On a bench in the fish-gallery or east alcove there are skeletons
of the python, of asea snake, Distira stokesii, from the Pearl
Banks, presented by Captain Donnan, and of a monitor lizard.
- The remaining smaller snakes and lizards are placed over table
cases near the western end of the gallery. These are chiefly
interesting on account of the relatively large number of endemic
forms, species of lizards of the genera Oeratophora, Lyrvocephalus,
Otocryptis, Cophotis, and Acontias being peculiar to the Island;
examples of these are shown over Table Case XVII. Endemic
species of snakes belonging to the genera Aspidur'aand Rhinophis,
as well as specimens of Lycodon aulicus, to which reference has
been made above, are exhibited over Case XVIII. The snake
Cylindrophis maculatus is also a peculiar species, and is the one
to which the native name “ Depat-naya” (two-headed snake) is
specially applied, though the term is equally applicable to Rhino-
phis and allied snakes. The hinder end of the body of these
snakes is truncated, and bears superficial resemblance to a head.
The colour, yellowish on dark brown, varies considerably,
especially in the case of Rhinophis blythi (see illustrations).
‘The giant tortoise from Aldabra, Testudo elephantina, shown
on the far side of the Peacock Case, does not belong to Ceylon, but
lived here for many years in the grounds of the gilla called
“Uplands” in Mutwal, near Colombo. It was found here at the
time of the British occupation in 1796, and had become thoroughly:
acclimatized.* It died in March, 1894.
A striking display of large marine turtles caught off the coast
of Ceylon is to be seen under the large platform in the eastern or
* The Hog-deer (Cervus porcinus), of which two living specimens are shown in
the grounds, is another example of an acclimatized animal, introduced from
India during the Dutch administration into the Kalutara District, where it now
occurs,
(. 44° )
fish aleove. The edible turtle, Chelone mydas, attaining a length
of four feet, is herbivorous.* The loggerhead turtle, Thalasso-
chelys caretta, is obtained by harpooning and netting by the natives
of Iranativu and elsewhere off the coast ; it is carnivorous, feeding
on crustaceans and molluscs. The leathery turtle, Dermochelys
coriacea, Was presented by C. H.de Soysa, Hsq.; it is not common.
Finally, a specimen of the tortoise-shell turtle, Chelone imbricata,
and a young edible turtle, are shown in small tanks on either side
of the staircase.
Some of the Amphibians of Ceylonare shown over Table Case IV.,
next to the Hagle Group. The large bull-frog is Rana tigrina,
the common toad Bufo melanostictus. The common frog of the
Colombo lake is Rana hexadactyla. The tree-frog,t lvalus
adspersus, peculiar to Ceylon, has been found at Pattipola, 6,200
feet, and is one of the rarest frogs existing. The climbing frogs of
the genus Rhacophorus attach their foam-like nests to the leaves of
shrubs and trees overhanging water, into which the tadpoles drop
when they hatch. The most singular batrachian occurring in the
hills of Ceylon above 2,000 or 3,000 feet is the worm-like, legless
‘ salamander,” { Ichthyophis glutinosus. It burrows in soft mud ;
the female lays eggs of large size, and coils round the clump of
eggs until they are hatched. The larve are aquatic, and are
provided with a respiratory orifice or spivaculwm on each side
of the neck. The development has been worked out by two
Swiss naturalists, Drs, Fritz and Paul Sarasin.
FISHES.
A large series of sharks and bony fishes is shown in the eastern
alcove of the gallery. The largest and rarest is the huge shark
which rests upon the platform along the centre of the room.
This is a specimen of the Basking Shark (Rhinodon typicus), 23
feet long, caught at Moratuwa in 1883. This species has also
been obtained off the Seychelles and the Cape of Good Hope;
it was first recorded from the west coast of Ceylon by Mr.
Amyrald Haly, the former Director of the Colombo Museum.
In spite of its bulk the specimen shown is not full-size, and
it is a harmless shark. It is regarded as one of the gems of
the entire collection.
* Chelonia virgata is synonymous with Chelone mydas {| Boulenger }.
+ The true tree-frogs of the family Hylide are not represented in Ceylon.
{ The tailed batrachians (newts and salamanders) form the order Urodela,
which is unrepresented in Ceylon ; the tailless batrachians (frogs and toads)
form the order Anura; the legless batrachians or the oxcilians belong to the
order Apoda,
NEST OF CLIMBING FROG. (COLOMBO.)
(Rhacophorus maculatus.)
[Vo saute page 44.
vi nah, hy
at Tent e
¢ 6°
Unfortunately the form of the mouth is lost in the mounted
Specimen; when fresh the width of the mouth was 3 feet, but
shrunk to lft. 1lin.in drying. ‘‘ When fresh the lower jaw was
quite straight and flat......... and considerably in advance of
the upper, so that the band of teeth in the lower jaw was quite
uncovered.”
The teeth in both jaws consisted of eleven (in the upper) to
fourteen (in the lower) rows of minute, sharp, recurved denticles,
of equal size, 2 millimeters long.* Another specimen, 18 feet
long, was taken at Negombo in March, 1889, and was presented to
the British Museum by the Government of Ceylon. In the same
year one was caught, 22 feet long, off Madras, andis now exhibited
in the Madras Museum.
At the end of the platform three other stuffed sharks are lying
on their sides; that to the right of the Rhinodon is a fine specimen
of the Hammer-headed Shark (Zygena malleus), characterized by
the shape of the rostrum, which is drawn out sideways into two
hammer-shaped lobes, at the ends of which are the eyes with the
nostrils near to the eyes; that to the left of the Rhinodon is the
Tiger Shark or Tope (Galeocerdo rayneri), distinguished by its
formidable notched teeth, This shark is said to be exceedingly
fierce and very cunning, swelling itself out so as to appear like
a floating mass of animal substance, in order to decoy its prey.
Behind the Rhinodon there lies another shark, 9 feet long, named
Ginglymostoma miilleri, Ginther. Below the front window, at
the back of the Rhinodon, is another interesting shark, A lopecias,
or Alopias vulpes, the Fox or Thresher Shark, which was rescued
by Mr. Haly from the Colombo market in February, 1884.
Against the adjoining north window are two sharks of the saw-
fish family; the larger specimen is an example of Pristis cuspi-
datus, the smaller is Pristis perrotteti. These differ from one
another in the position of the first dorsal fin and in the armature
of the rostrum, a considerable portion of the base of which is
destitute of teeth in P. cuspidatus.
A few more sharks are preserved in spirits in the adjacent
wall case, The skates and rays next invite attention. Over the
wall cases beside the north window are two examples of a large
Sting Ray (Zrygon uarnak), and in the bottom shelf of the left
wall case is a Thorny-backed Ray, said to be common in the Indian
eet called Urogymnus es In a trough on the
* Of. A. Haly. Occurrence of Rhinodon ie Smith, on the West Gonat of
Ceylon, Ann. Nat. Hist, (fifth series), vol. XII., 1883, pp. 48-49.
t G&. Day, F. Faun. Brit. Ind., Fishes, vol. L., p. 33.
t &. Note vy A. Haly in the “ Taueobanian 1886, vol. I., p. 167.
( 46 )
south-east verandah is a young specimen of the “ Two-horned ”
Skate (Dicerobatus eregoodoo), exhibited with the lower side up
displaying the gill-clefts with the gills showing through. In
another trough on the opposite north-east verandah is another
well-preserved Sting Ray of the species T’rygon sephen, and near
to this is a young Beaked Ray (Aétobatis narinari). The skates
and rays feed largely upon crabs and molluscs.
The fairly numerous specimens of Teleostean or bony fishes
which are exhibited in the wall cases are somewhat remarkable
on account of the well-preserved colour markings, which in many
cases have retained their freshness after the lapse of several years,,
in a gum and glycerine mixture adopted by Mr. Haly. Against
the window beside the Thresher Shark is a good example of the
Swordfish (Histiophorus gladius). The large stuffed fish in the
bottom shelf of the adjoining wall case is a Wrasse (Labride).
The parrot wrasses, fishes of brilliant colours, feeding in the
neighbourhood of coral reefs at Galle, for example, with parrot-
like beak consisting of teeth soldered together, belong to the
genus Pseudoscarus of the Wrasse family. ‘The ‘ Red Mullet” of
the Colombo market is Serranus sonnerati, of the Perch family.
The Seirfish, the staple fish food of Colombo, belongs to the
Mackerel family (Scombridz), and is named Cybiwm guttatum.*
The jumping fishes to be seen about the rocks at Mount Lavinia
and Galle are blennies of the genus Salarias; and the mud-
skippers of Negombo are gobies of the genus Periophthalmus.
The principal fresh-water fishes of Ceylon are the Carps (Cypri-
nid), including the Indian game fish called the Mahseer (Barbus
tor), the Ophiocephali, tank-fishes, the Labyrinthici or “ climbing
perches” (Anabas scandens and Polyacanthus signatus), and
the Catfishes (Siluride). One genus of catfishes, Arius, called
“anguluwa” in Sinhalese, occurring at Panadure, Kalutara, and
elsewhere, has the remarkable peculiarity that the males carry the
eggs, 15-20 in number, in their mouths until they are hatched.
INSECTS.
' The Lepidoptera (butterflies and moths) are shown in eight
table cases placed near the Kagle and Peacock Groups.
The Moths or Nocturnal Lepidoptera commence in Table Case II.
with the family Saturniide, the caterpillars of which spin silken
cocoons in which they pupate and from which they emerge in the
adult orimago phase of their life-history. This family comprises
the largest moths found in Ceylon, namely, the Lunar Moth
*Not exhibited. There is an extensive tunny fishery (Zhynnus thunnina) off
Balapitiya during the north-east monsoon, and the fish are daily sent to Colombo.
Salarias Andersoni.
[To face page 46.
ee
(Actias selene), green with a dark bordered whitish circle in the
centre of each wing and with long swallow-tailed hind wings ;
the Atlas Moth (Attacus atlas), deep chocolate with oblique trans-
lucent windows in each wing ; the Tussur Silk Moth (Anthercea
paphia), rich yellow with a transparent round window in each
wing. The Bombycide and Eupterotide complete this side of the
case. The other side contains the Sphingide, which include
among many species the Death’s Head Moths of the genus Aciie-
rontia, the caterpillars of which stridulate, uttering a clicking
sound by rubbing their jaws together ; and the Clear-wing Moths
(Cephonodes hylas), which sometimes become a pest in cultivated
districts. All the caterpillars of this family are provided with a
horn-like appendage on the back near the end of the abdomen.
Case III. opens with the N otodontidzand closes with the Arctiide.
The Syntomide, which follow the Notodontidz, include some
very common species, while the Zygzenide bear the appearance of
butterflies, which they further resemble by their habit of flying
about during the day.
Case IV. continues the Arctiide and commences the extensive
family of the Noctuids, the latter being continued in Case V., which
contains several handsome moths, eg., Phyllodes consobrina with
leaf-shaped forewings, and Ophideres salaminia, which seems to
mimic a Sphingid moth.
Case VI. contains Uraniide, Epiplemidz, Geometride, and the
commencement of the Pyralidz or shining moths. Case VII. con-
tinues the Pyralide and concludes with a few Microlepidoptera
of the family Tineidz. On the other side of this case are shown
some dragon-flies (Neuroptera).
Finally, Cases IX. and X., placed alongside in the central space,
are devoted to the butterflies, which include a fine series of the Leaf
Butterfly, Kallima philarchus.
In addition to the exhibited Lepidoptera the Museum possesses
a large duplicate students’ collection containing many rare species
not shown in the cases. This may be inspected on application.
There are, on the other hand, many moths recorded from Ceylon
which have not yet found their way to the Museum collection.
The other orders of the insects of Ceylon have not been worked
out so thoroughly as the Lepidoptera. They are represented in the
table cases near the west end of the gallery, and are subject to
re-arrangement. Orthoptera (locusts, stick insects, cockroaches,
and mantids) and Coccide (mealy bugs) are shown in Cases XV. and
XVI ; Coleoptera in Case XVII.; Hymenoptera (ants, bees, and
wasps), with a couple of black flies mimicking a eg and a bee
respectively, in Case XVIII.
( 4 9
The Museum is largely indebted for many specimens in the
Insect Department to Mr. E. E. Green, the Government Ento-
mologist,and to Messrs. F. M. Mackwood and O. 8. Wickwar, who
-have also devoted much time to the arrangement of the duplicate
collections of Butterflies, Moths, and Hymenoptera.
CRUSTACEANS AND MOLLUSCS.
Some Crustaceans (crabs, hermit crabs, prawns, barnacles) will
be found among the table cases, chiefly on the tops of the cases.
Marine Shells and Land Shells are contained in the table cases
surrounding the central group. The marine shells of Ceylon are
not remarkable for their exceptional variety and abundance.
Ceylon follows far behind many other localities of the Indo-
Pacific Region in the richness of its Marine Molluscan Fauna, but
a sufficient compensation is afforded by the presence of the
celebrated pearl banks.
On the other hand, the land shells of Ceylon are highly pecu-
liar, and comprise many endemic species. The largest snails of
the Island belong to a genus, Acavus, which is confined to Ceylon.
Mr. Oliver Collett, F.R.M.S., who lived for some years at Amba-
gamuwa, was a great collector and connoisseur of the land-shells of
Ceylon, and published three “Contributions to Ceylon Malacology”
in the Journal of the Ceylon Branch of the Royal Asiatic Society
(vols. XV.-XVI., 1897-1900). On his death, which occurred prema-
turely in 1902, his collection was purchased by the Government
of Ceylon for the Colombo Museum and forms a valuable students’
collection, which can be viewed on application. It contains
several species still unnamed.
PEARL BANKS AND CORAL REEFS.
A number of specimens in Table Cases I. and VIII., some of
which were presented long ago by Captain Donnan, many more
having been added recently by Mr. James Hornell, Marine Biologist
and Inspector of Pearl Banks, illustrate the natural history of the
banks or paars which afford anchorage tothe pearl oyster. Prior
to the year 1903 there had been no pear! fishery for twelve years,
and in 1902 a special mission, consisting of Professor W. A.
Herdman, F.R.S., assisted by Mr. James Hornell, was appointed to
report on the Pearl Oyster Fisheries of the Gulf of Mannar. “The
animal (Margaritifera vulgaris, Schum.=Avicula fucata,Gould)
is not a true oyster, but belongs to the family Aviculiday, and is
therefore more nearly related to the Mussels (Mytilus) than to
the Oysters (Ostvea) of our British seas. One very notable
character of great practical importance, in which it differs from
( 49 )
Ostrea, is that the pearl oyster, like our common mussels, has a
‘byssus’ or bundle of tough threads by which it can attach itself
to rocks or other foreign objects.” *
The collection shows pearl oysters from several paars in various
stages of growth, and other organisms, corals, pennatulids, sponges,
sea urchins, &¢., which also grow on the pearl banks in association
with the pearl oysters. Of special interest are the specimens
prepared in spirits showing pearls 77 situ.
Large specimens of stone corals (Madreporaria), sea shrubs
(Gorgonacea), black corals (Antipatharia), and leathery corals
(Alcyonacea) from Galle and the Maldive Islands are exhibited
in wall cases in the fish aleove and on the south verandah. Those
from the Maldive Islands were presented by Mr. J. Stanley
Gardiner, M.A., who organized an expedition for the investigation
of the coral atolls of the Maldives in the years 1899-1900.t
The dried corals which are exhibited are the basal and axial
skeletons secreted by the soft parts of the living coral polyps.
These form large colonies by a peculiar process of budding and
branching, and deposit the mineral substances which they have
absorbed from the sea water so as to form the wonderful growths
which remain after the living tissues have been removed.
The Madreporaria are the reef-building corals forming extensive
reets at Galle and off Jaffna, and especially at the Maldive Islands,
which are themselves partially elevated coral reefs. The pearl
banks are not coral reefs, but sandbanks, formed of sandstone
and concretions upon which isolated corals grow without forming
reefs,
* Of. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the
Gulf of Mannar, by W. A. Herdman, D.Sc., F.R.S., with supplementary reports
upon the Marine Biology of Ceylon by other naturalists. Published by the Royal
Society, Part I., London, 1903. Professor Herdman and Mr. Hornell arrived in
Ceylon in January, 1902, and the former left in the following April. Since then
the work has been carried on locally by Mr. Hornell.
§ The Fauna and Geography of the Maldive and Laccadive Archipelagoes.
Edited by J. Stanely Gardiner. Vols, I. and IT., 1901-1905 (Cambridge University
Press).
H 105-04
( 50 )
THE ROCKS AND MINERALS OF CEYLON.
By A. K. COOMARASWAMY, B.Sc.,
Director, Mineralogical Survey of Ceylon.
I—ROCKS.
THE rocks of Ceylon are known as Granulites, or, using the
term in its widest sense, as Gneisses; they belong to the same
series as that which Mr. Holland has named in Southern India
the Charnockite Series, The rocks are crystalline and show con-
spicuous mineral banding (foliation), and are very varied in
mineral composition. Rocks of igneous origin form by far the
greater part of the Charnockite Series in Ceylon, although it is
possible that amongst these rocks there are some of sedimentary
origin now highly metamorphosed and incorporated with the
others. The bedded appearance so suggestive of sedimentary
rocks is in this respect deceptive, and is due to the banded
structure produced by flowing movements in the _hetero-
geneous magma during its consolidation. No fossils occur in any
of the crystalline rocks.
The most characteristic types of rock exhibited inelude
‘vranular quartz rock, consisting essentially of quartz, but often
with minute quantities of felspar and garnet; leptynites, composed
of quartz and felspar, and very often containing an abundance of
garnet ; typical charnockite, essentially composed of felspar quartz
and hypersthene; pyroxene granulites, characterized by the
presence of pyroxene (hypersthene or augite or both) with
felspar (usually triclinic) and with or without orthoclase, quartz,
or garnet ; amongst these are rocks with the mineral composition
of norites, diorites, and gabbros ; there are also more basic types
consisting almost entirely of pyroxene, amphibole, and garnet ;
these dark heavy rocks frequently occur as lenticular bands and
inclusions in the more acid types. Coarse-grained pegmatites of
an intrusive character are found in dykes and veins crossing or
parallel to the foliation of the other rocks ; in them the minerals
quartz, orthoclase, mica, and hornblende are most characteristic ;
the quartz and felspar are often intergrown as in graphic granite,
The crystalline limestones are found in wide or narrow bands
interbedded with the other rocks, often separated from them
by a zone of heavy dark green rocks composed essentially
of pyroxene and mica, often with spinel. The limestones
( 1)
themselves vary much in purity and in dolomitic character ; the
minerals most frequently occurring and indeed nearly always
present are forsterite and phlogopite; graphite is frequently
abundant in small flakes; other accessory minerals include
pyrite, blue apatite, spinel, amphibole, chondrodite, scapolite, &c.
Of inclusions in the limestones there are (a) aggregates of the
characteristic accessory minerals ; (0) inclusions of rocks indis-
tinguishable from the pyroxene granulites except for the usual
presence of scapolite and sphene.
Certain rocks composed essentially of pyroxene with scapolite,
amphibole, mica, calcite, and sphene occur sparingly in bands
interbedded with the other rocks; separating limestone from
granulite ; or as inclusions in the limestones.
The Galle Group,—aA group of rocks allied to these but distin-
guished by the presence of wollastonite is found at Galle. These
rocks include types composed of pyroxene, scapolite, sphene,
wollastonite, and graphite, and all transitions from these to rocks
composed of quartz and felspars. There are also remarkably
coarse dykes composed essentially of orthoclase, quartz, and
wollastonite, crossing the foliation; the individuals of wollas-
tonite, moulding the idiomorphic crystals of quartz and ortho-
clase, are sometimes as much as 3 feet in length,
The name Balangoda group is given to a series of granitic rocks,
intrusive in the Charnockite Series or granulites; the granites -
occur in dykes and lenticular masses, the best exposures so far
known being in the Balangoda district. The principal types
include zircon granite, allanite-granite or pegmatite, magnetite
granite, and granite without accessory minerals. But although
not yet discovered in situ (except thorianite, which occurred very
sparingly in a pegmatite vein at Gampola, and ilmenite, which is
common in several types), the following additional minerals are
almost certainly derived from rocks of the Balangoda group:
baddeleyite, beryl, cassiterite, chrysoberyl, fergusonite, geikielite,
ilmenite, rutile, some spinels, thorianite, thorite, topaz, and some
tourmaline.
Rocks which arealteration products of those already mentioned
include bands of chert (opaline chalcedonic rock), which usually
occurs replacing the carbonates of the crystalline limestones,
minerals such as spinel and graphite remaining in the chert
unaltered. Laterite (cabook) is a typical decomposition product
of the granulites, and is of interest on account of its use as a build-
ing material, and on account of its chemical composition, consist~
ing largely of aluminium hydrate. Various ¢lays, including
kaolin (china clay, “ kirimeti,” “ makul”) are derived from the
( 52)
decomposition of the granulites, especially from their felspathie
elements.
Volcanic rocks are very sparingly represented in Ceylon. A
typical dolerite (probably occurring as a dyke) from Kallodai,
Eastern Province, is represented in the collection. A few other
dykes are said to have been met with in Ceylon. No lavas or
tuffs occur, and, with the exception of a few hot springs, no signs
of volcanic activity are found. Denudation has been so long at
work that only the most deep-seated rocks are now exposed at
the surface. Of sedimentary rocks, we have (a) river gravels,
sands, and alluvial clays (used for brick making) deposited
by rivers. In the gravels are found the gems which have been
derived from the crystalline rocks ; many have not yet been found
in situ. (b) Marine deposits, including coralline raised beaches,
blown sand, &e. Fossil shells and corals are abundant in the
raised beaches which are found all round the coast and often some
little way inland, but never very far above the present level of
the sea.
II.—_MINERALS.
Before proceeding to refer in systematic order to the nature and
mode of occurrence of particular minerals it will be useful to
consider briefly their nature and mode of formation. Regarded
from this point of view the minerals of Ceylon fall into three
main groups, which are not, however, separated from each other
by any rigid line of demarcation :—
A.—Minerals composing or associated with the crystalline
rocks,
B.—Vein minerals.
C.—Minerals which are alteration products of other minerals
and rocks.
The first class includes those whose origin is for the most part
directly igneous (7.¢., the majority of Ceylonese minerals), and
which occur as original minerals in the granulites and crystalline
limestones. Many of these have crystallized from an actually
molten or viscous magma, others perhaps from masses of rocks
existing merely in a state akin to fusion, some being developed
as the result of contact interaction between the limestones and
granulites when both possessed a high temperature, whilst others
forming pegmatite veins may even have crystallized from heated
vapours or liquids saturated with mineral matter. When suitable
conditions prevail, these minerals each assume definite and charac-
teristic crystalline forms ; this has, however, rarely been the case
in the granulites themselves, where an irregular “ granulitic”
structure usually prevails; in the crystalline limestones, however.
Ce. grils
Pe ava
PLUMBAGO MINE,
To Face page 58.)
(58 a
the accessory minerals have more often been able to crystallize in
their own forms ; for an example see Fig. 1, an octahedral crystal
of spinel. Such “idiomorphic” erystals are commonly known
in Ceylon as “ devil-cut.”
It is interesting to note that of all the gem minerals so famous
in Ceylon few have yet been found in situ. New records of the
occurrence of these minerals in their parent rock is much desired.
Nearly all the gems of Ceylon, moonstone excepted, are obtained
from the river gravels of the Ratnapura, Rakwana, Galle, and
Morawak Korale Districts; but some are found in superficial
deposits whose situation on hill slopes shows that the contained
minerals must oceur in sity at no very distant spot.
Fig 1.—Octahedral crystal of spinel.
Minerals of the second class include the greater part of the
graphite of Ceylon and perhaps a considerable part of the mica.
Associated with the graphite are often found some of the minerals
characteristic of the granulites, such as quartz and felspar. True
veins of pure quartz are rarely met with in Ceylon. Minerals of
the third class include the deposits of iron ore (limonite) and
manganese ore (psilomelane), which are everywhere common as
decomposition products of the granulites ; and also the minerals
hydrargillite and limonite composing laterite.
A few minerals occur in Ceylon which have not been found
elsewhere; these are thorianite, geikielite, and serendibite. The
mineral baddeleyite was first found in Ceylon, but has since been
met with in Brazil.
Further notes on the more important minerals will be found
below, where a list of all the minerals known to occur in Ceylon
is given.
Graphite, C.—Graphite or plumbago is the most important
mineral product of Ceylon. Its compositionis pure carbon. It is
found in veins and nests in the crystalline rocks, occurring often
i
in a fibrous or flaky form, the flakes being arranged at right angles
to the wall of the vein (see Fig. 2). The veins vary in width from
less thanan eighth of an inch to several feet. Some are found to
follow the foliation planes of the various rocks, others cross them
and ramify in all directions. Much smaller quantities of graphite
occur as flakes in many of the granulites and in the crystalline
linrestones, when it usually forms small tabular six-sided crystals
with well-developed basal cleavage. In these cases the graphite
behaves like the other accessory minerals, and there is no reason
to suppose that it has been subsequently introduced,
Fig. 2.—Vein graphite surrounding a portion of included matrix
(white leptynite).
It is clear, however, that the graphite occurring in veins has been
deposited at atime posterior to the consolidation of the granulites.
The veins are often of the most typical character. Usually they
consist of pure graphite (sometimes there is evidence of more than
one period of deposition ina zoned structure of the vein) ; some-
times the vein shows a central zone of quartz or pyrite with
graphite on either hand, sometimes the graphite is more irregularly
associated with minerals such as felspar quartz and mica and with
fragments of the surrounding rock.
Metamorphism of the surrounding rocks near the veins is found
only on a very small scale ; the rock surfaces in immediate contact
with the veins are not impregnated with scales and flakes of
graphite toa greater depth than half an inch. Nor do. we find that
the quartz and other minerals associated with the graphite veins
Cio a
are filled with disseminated graphite; the latter occurs only in
strings or scales occupying obvious cavities or cracks in the quartz.
It seems that the deposition of graphite has been subsequent to the
formation of the pegmatite veins, though following the same or
similar paths in some cases.
As regards the source of the graphite, we see at once that its
presence cannot be ascribed to the metamorphism of beds of coal
or other carbonaceous deposits, and indeed that the graphite can
have had no direct organic source and is most probably of entirely
inorganic origin. Like other minerals found in veins, it must have
been deposited from vapours or liquids saturated with mineral
(in this case carboniferous) matter; for we cannot suppose that
the graphite was introduced in the form of dykes of molten
carbon.
Sulphur, 8.—Oceurs as a decomposition product of pyrite in
small quantities.
Gold, Au.—Small quantities of native gold are of rare occur-
rence in streams, sands, and alluvial deposits, but there is little
prospect of its ever becoming of commercial importance.
Salt, NaCl.—Obtained by the evaporation of sea waterin natural
and artificial lakes, known as “ lewaya.”’
Cinnabar, Hgs.
Pyrite, FeS,.—Common as an accessory mineral in granulites
and crystalline limestones.
Marcasite, FeS,. ‘
Pyrrhotite, Fe, ,S,.—Occasionally as an accessory mineral in
crystalline limestones and in charnockite.
Ice, H,0.—Frost is sometimes experienced in the hills.
Fig 3.—Corundum.
Corundum, Al,O,,.—Corundumis the most important of the gem
minerals of Ceylon. When red it is known as ruby ; when blue
as Sapphire ; when purple as oriental amethyst ; when yellow as
oriental topaz ; colourless varieties are also found. The peculiar
character of ‘star sapphires ” is‘due to the presence of regularly
distributed minute inclusions arranged along the lines of growth,
producing the appearance known as “ silk ;” a six-rayed star is
seen when the crystalis viewed in the direction of its vertical axis.
Coarser varieties of corundum are used as emery, for which pur-
pose a good deal is obtained in Southern India. Corundum is
found in the river gravels of Ceylon in more or less rounded and
waterworn six-sided crystals, which are either prismatic or doubly
pyramidal in character.
Only two localities are known in Ceylon where corundum
occurs in the parent rock. In one case (Talatu-oya) blue hexa-
gonal crystals occurred in a narrow band of rock composed essen-
tially of orthoclase, microperthite, and oligoclase. In the other
(Haldummulla) violet or purplish hexagonal crystals are found
in a corundum-sillimanite rock (of which a large specimen is
exhibited), but not actually in situ.
In Southern India corundum occurs (a) in felspathic rocks as a
direct product of the magma; (0%) in certain aluminous rocks,
probably as the result of contact metamorphism. Specimens of
these rocks are shown in the collection. In Burma rubies are
found in the erystalline limestones, which in other respects
closely resemble those of Ceylon.
Hematite, Fe,0,-—Less usual than limonite as an iron ore
derived from the decomposition of the granulitie rocks.
Limonite, 2Fe,0,3H,0.—The common ore of iron in Ceylon,
and formerly extensively worked. Almost always found as a
product of the decomposition of the granulites.
Hydrargillite, A1,0,3H,0.—Forms, with limoiite, the greater
part of ordinary laterite (cabook),
Cassiterite, SnO,.
Rutile, TiO,.
Thorianite, ThO,+U0,.— ‘his newly-discovered mineral,
peculiar to Ceylon, is of great commercial importance owing to
the use of thoria in the manufacture of incandescent gas mantles.
It oceurs in very heavy black cubic crystals at Bambarabotuwa,
where over a ton has been obtained. It is valued at £600 sterling
per ton. It is of great scientific interest too, on account of its
chemical composition, one or more new elements being possibly
present ; it contains also a large amount of occluded helium.
Though radio-active, there is no more than a trace of radium
present.
Baddeleyite, ZrO ..
Quartz, Si0,.—Very abundant throughout the Charnockite
Series. The purple variety is amethyst. Drusy groups from
(BF)
cavities in crystalline limestone at Welimada. Doubly terminated
erystals not rare in gem washings.
Fig. 4 shows the ordinary hexagonal prisms of quartz.
Fig. 4.—Quartz hexagonal prisms,
Chert, SiO,.—An opaline chalcedonic rock usually replacing
crystalline limestone.
Chalcedony, SiO,.
Opal, SiO, +H,0.—In some cases a rock consisting entirely of
common opal is found with the less pure cherty varieties.
Psilomelane, MnO+Mn0O,H,O.—Commonly associated with
limonite and hematite in veins and aggregates resulting from the
decomposition of the granulites.
Spinel, (MgFe)OA1,0,;.—Blue, green, and red spinels (especially
the latter, known as Balas rnby) are used as gems, Small well-
formed octahedra (see Fig. 1, page 53) of spinel, usually pink, are
common in the crystalline limestones. Green spinel occasionally
occurs in granulites, and frequently in the heavy dark green
rocks associated with junctions of limestone and granulite. The
gem spinels are obtained from the gravels, but are probably
derived from the crystalline limestones.
Magnetite, FeEOFe,0,.—Common as an accessory mineral in the
granulites ; also in limestones.
Chromite, FeOCr,03.
Chrysoberyl, BEOA],0,.—Includes cat’s-eye and alexandrite.
This important gem stone sometimes occurs in large individuals,
exhibiting characteristic twinnirg. The beautiful chatoyance of
the cat’s-eye is perhaps due to the presence of fine tubular cavities
arranged symmetrically in the crystal. (The much less valuable
I 105-04
(, 58)
“ coast ” or quartz cat’s-eye is of a quite different character, consist-
ing of quartz with included silky fibres of asbestos.) The green
alexandrite appears red by transmitted light and generally also by
candle light, but green by reflected light or daylight.
Calcite, CaCO,.—In the erystalline limestones.
Dolomite (CaMg)CO..—In the crystalline limestone ; sometimes
forming with calcite parallel or ramifying intergrowths.
Forsterite, Mg,Si0,.—This colourless mineral is characteristic
of the crystalline limestones, where it is extremely abundant.
The individuals are usually small. An unusually large crystal
in limestone is exhibited; also some very flattened crystals of a
superficially dark colour. Incipient superficial decomposition
often gives a dark colour to the crystals.
Clinohumite,Mg,(MgF),(Si0,),.—This beautiful yellow mineral
is of rare occurrence in the crystalline limestones.
Pyrozenes :
(1) Hypersthene (FeMg)Si0,.—One of the most characteristic
minerals of the Charnockite Series.
(2) Diopside (CaMg)Si0,.—A colourless to pale green pyroxene
characteristic of the crystalline limestones.
(3) Manganhedenbergite, Ca(FeMn(Si0,),.—Characteristic of
the Galle group,and probably common in similar rocks elsewhere.
(4) Augite, CaMgFe(Si0,). +MgFe(AlFe),(Si0;),.. — Charac-
teristic of many pyroxene granulites.
(5) Wollastonite, CaSiO,.—Characteristic of the rocks of Galle,
in which it occurs disseminated, and also in very large individuals
in coarse pegmatite veins. It has not yet been found elsewhere
in Ceylon.
Amphiboles :
(1) Tremolite, CaMg,(Si0,),.—Includes colourless and pale
amphiboles associated with the crystalline limestones.
(2) Horneblende, Ca(MgFe),(Si0,), &¢.—Includes dark amphi-
boles occurring in the crystalline limestones and the dark green
amphiboles characteristic of many of the more basic varieties or
granulites and of contact zones.
Ilmenite, FeTiO.,.—Of widespread occurrence ; the commonest
ingredient of nd@mbu and black sand. One exceptional and very
large specimen is shown.
Geikielite, MgTiO,.—Not known except in Ceylon, where it was
found in gem refuse.
Titanite (Sphene), CaTiSiO ,— Characteristic of the rocks of the
Galle group; common in junction rocks, &c¢.
Talc (Steatite), H.Mg,(Si0,),.—To be distinguished from mica,
Rare as an accessory mineral in crystalline limestones,
Cae)
_ Serpentine, H,Mg,Si,0,.—Not infrequent as a decomposition
product. of forsterite. ‘:
Apophyllite, (HK),Ca(Si0;)2H,0.
Topaz, (AIF),Si0,.—Commonly found in gem gravels ; yeaa.
colourless, or pink, the latter variety known as king topaz. The
colourless varieties are wrongly knownas watersapphires. The pale
greenish-blue.varietiesarecutas aquamarine. Not met with im s¢tu.
Andalusite, Al(A10)Si0,.
Kyanite, (Al0),Si0.,.—Has only been found in dredgings made
by Professor Herdman off the coast of Ceylon.
Sillimanite, Al,Si0,.—Very rarely in good crystals in gem
gravels. Sillimanite is in some districts a common constituent of
the garnetiferous leptynites, the rocks then resembling the
khondalites of Southern India, of which specimens are exhibited.
Fine coarse sillimanite rocks occur near Haldummulla, the
sillimanite being disposed in sheaf-like and radiating aggregates.
The associated minerals are corundum, garnet, orthoclase-micro-
perthite, ilmenite, and rutile. See also under Corundum.
Kaolinite (China Clay), H,A1,8i,0,.—Common as a decom-
position product of orthoclase.
Felspars :
(1) Orthoclase, KAISi,0, affords the well-known moonstone ; it
occurs in large but well-cleaved crystals in certain acid granulites
associated with crystalline limestones in the Dumbara district,
Central Province. Various pegmatites also, consisting of quartz
and orthoclase, yield moonstone of a poor quality. Large indivi-
duals of idiomorphic orthoclase occur in pegmatite veins at Galle.
An intergrowth of orthoclase with albite is the most usual felspar
of the less basic granulites. The silvery sheen so characteristic of
moonstone is probably the result of the presence of excessively
minute inclusions of kaolin, the products of incipient decom-
position. The bluish-white opalescence of moonstone is best seen
when the crystal is viewed in a direction at right angles to the basal
plane, i.2., when regarding one of the planes of easy cleavage ; the
stone should always be so cut that the flat base of the finished
cabochon gem is parallel to this surface, in order that the opales-
cence may be central and as conspicuous as possible.
(2) Plagioclase (Lime-Soda Felspars).—These are commonly
characteristic of many of the granulites, but are rarely of large size.
A fine blue opalescence has occasionally been observed in the
plagioclase felspars, but none of size suitable for yielding gems have
been found.
(3) Microcline, KAIS,0,.—Rarer than orthoclase in the granu-
lites.
Loe
Micas.—Ceylonese micas are of some importance from a com-
mercial point of view. They include muscovite, biotite, and
phlogopite :—
(1) Muscovite, H,KA1,(Si0,), occurs but sparingly in Ceylon,
and not in crystals large or flawless enough to be of commercial
value.
(2) Biotite, (KH),(MgFe),(AlFe),(Si0,), is chiefly found in
smiil crystals as a microscopic constituent of various rocks
belonging to the Charnockite Series; but a part of vein mica also
belongs to biotite.
(3) PAlogopite, (K,H,Mg,F),;Mg,Al(Si0,), is by far the most
important of Veylonese micas. Minute pale or golden crystals are
almostalways common in the crystalline limestones. Larger mica
crystals occur in veins and bands associated usually with junction
of granulite and crystalline limsstone. ‘hese micas are rarely
colourless, being more usually brown, reddish, bottle-green, or
amber-coloured. The largest crystals found have been two or
three feet in diameter. ‘he veins are usually one or two feet
in width, and are composed of numerous “books” or crystals of
mica which are generally more or less idiomorphic, having a clear-
cut hexagonal outline. Mica is used for the peepholes of stoves,
for lamp chimneys, aud very largely for electrical appliances,
&c.; comminuted mica dust is of use as a non-conducting packing.
Mica has long been used in the Hast for ornamental purposes, also
for medicine. Fine crystals of mica from the Kandy District are
exhibited.
Fig. 5.—Tourmaline,
Tourmaline.—A borosilicate of alumina with magnesia iron and
alkalis. Black varieties associated with quartz are not uncommon.
C361")
The greater part of the foramalli of gemmers belongs properly to
zircon.
Serendibite—A borosilicate of alumina and lime with magnesia
and alkalis. Found only in Ceylon. Insmall crystalsin diopside
rock at the junction oflimestone and granulite. Dumbara district,
Central Province.
Scapolite, Ca,Al,Si,0,;Na,Al,Si,0,,Cl.—Abundant in the
wollastonite-scapolite gneisses of Galle; common in limestone-
granulite junction rocks. Sometimes an accessory mineral in
limestones.
Stilbite, (Na,Ca)Al,Si,0,,6H,0.—In minute crystals; from
Nilhene, near Baddegama, Southern Province.
Zoisite, Ca. A1,(AlOH)(Si0,);.
Allanite, (CiFe),(AlCeFe),(A10H)(SiO,),.—In coarse granitic
dykes near Balangoda.
Beryl, Be, Al,(SiO.;)g.—The pale varieties of emerald known as
aquamarine are abundant in gemgravels. Ceylon specimens with
the true emerald colour have been very rarely met with.
Cordierite (Iolite), (MgFe),Al,Si,0, ,—Inrolled crystals known
as water sapphire. Many stones, however, known as water
sapphire belong to topaz, and are colourless.
Garnets.—Several types of garnet occur in Ceylon. Amongst
these are probably—
Cinnamon stone, Ca,Al1,(Si0,),.—Not known in situ.
Pyrope, Mg,A1,(Si0,)s. These inc!ude the red and pink-
Almandite, Fe, Al,(Si0,)5. | ish-red garnets so Common in and
Spessartite, Mn.Al,(Si0,),. ) characteristic of the granulites.
Fig. 6.—Zircon.
Zircon, ZrSi0,.—A very abundant mineral in Ceylon; occurs
in the granulites as a microscopic constituent; near Balangoda
( 62 )
occurs abundantly in large idiomorphie crystals in a zircon
granite. Common in gem washings, and usually well crystallized.
Colour various : brown, yellowish, green. The colourless varieties
are known as Matara diamonds; the coloured as jargoon and
hyacinth, used as gems.
Thorite, ThSiO,.
Apatite, Ca,(CaF)(PO,),.—A microscopic constituent of many
granulites. Blue apatite is very characteristic of the crystalline
limestones.
Fergusonite, (YErCe)(NbTa)O,.
Anhydrite, CaSO,.
Uraninite (Pitchblende).—Uranate of lead, the chief source of
radium. Not certainly known to occur in Ceylon, as all supposed
specimens may be thorianite.
The above forms a complete list of the minerals at present
known to occur in Ceylon; the discovery of others may -be
expected ; minerals of the samarskite and zschynite groups are
probably present.
For further information as to the rocks and minerals the
following works may be consulted, amongst others :—
Weinschenck, E. Zur Kenntniss der Graphitlagerstatten ; Die
Graphitlagerstitten der Insel Ceylons. Zeit. fur prakt. Geol.:
1900, p. 174.
Griinling, F. Ueber die Mineralvorkommen von Ceylon. - Zeit
fur Kryst, vol. XXXIII., 1900, heft 3.5, pp. 209-239. .
Coomaraswamy, A. K. The Crystalline Limestones of Ceylon.
Quart. Journ. Geol. Soe., vol. LVIII., 1902, pp. 399-422.
Coomaraswamy, A. K. The Point de Galle Group (Ceylon);
Wollastonite-scapolite Gneisses. Quart. Journ. Geol. Soc., vol.
LVIII., 1902, pp. 680-689.
III—ROCKS OF SECONDARY ORIGIN.
Of these, Laterite (cabook) is perhaps the most important. It
commonly occurs overlying quite unaltered granulites, forming
a mantle varying from a few feet to many yards in thickness in
different places, but is of rather local distribution. When dug itis
soft, but as it hardens on exposure it forms a useful building
stone. Typical laterite is a decomposition product of the rocks
beneath, and is especially characteristic of tropical lands. Laterites
are usually found to consist of a fine-grained scaly aggregate of
hydrargillite or similar aluminium hydrate, with also ferrous
( 63 )
hydrate varying in amount according to the nature of the parent
rock. When the latter contained free quartz, it is found in the
laterite in angular grains.
Various Clays, including kaolin (China clay), are also found, and
are also decomposition products of the granulitic rocks. ‘Chey
differ chiefly from laterite in consisting rather of aluminium
silicates than of aluminium hydrate.
Cherts and common-opal rock are sparingly found, often in
association with crystalline limestones, and can sometimes be
shown to have been formed by pseudo-morphous replacement of
the latter ; specimens occur containing the phlogopite, graphite,
and spinel of the original crystalline limestone, as well as others
in which remains of the partially disintegrated carbonates can
still be seen.
Travertine (tufa) is a secondary deposit of carbonate of lime
(apparently sometimes containing magnesium carbonate as well)
removed in solution from the beds of crystalline limestone; but
these deposits are but rarely found in large quantity.
IV.—FOSSILS AND RECENT SEDIMENTARY ROCKS.
Even less is known of these than of the crystalline rocks. A
belt of raised beach deposits is almost everywhere to be traced
around the coast, extending to no great height above sea level.
The material composing the deposit varies from a sandstone
cemented by caleareous material to a rubbly rock composed
entirely of the débris of corals and other calcareous organisms.
A series of recent marine fossils from Palanti-aar is exhibited,
and includes shells in a caleareous breccia, and well-preserved
remains of crabs in nodules of mud. There are also fossil chank
shells from the Kadurawala coast ; the latter are regularly
quarried in the recent deposits of the Jaffna District.
An interesting specimen of a bone and shell breccia from the
floor of a Vedda cave is exhibited ; but little is known, however,
as to the occurrence of really ancient cave deposits; masses of
stalactite are rarely found in caves in the crystalline limestones.
In addition to the recent marine sedimentary deposits, there are
river gravels and alluviums. Thick beds of the former are of
somewhat rare occurrence, but rivers, large or small, are not
infrequently bordered by strips of alluvial deposits, and when
they leave behind the mountain country, and with it their often
torrential character, debouch upon extensive alluvial plains where
fine silty muds are still deposited in times of flood. The fine
muds thus laid down are of great value in the manufacture of
bricks and the coarser kinds of earthenware.
( 64 )
V.—GEMS AND GEMMING.
The gems of Ceylon are of such general interest that it will be
worth while to give a brief account of the manner in which they
are obtained. With the exception of moonstone and some garnets,
none are obtained 7n situ, though all of course are derived from
the crystalline rocks where they originally crystallized, like the
other minerals accompanying them. It is however popularly and
erroneously supposed that they have grown where found, and that
small and flawed gems are merely immature.
The gems (of which a tabulated list is given below) are obtained
from gravels which have been deposited by streams and rivers ;
gemming is now only carried on in the Ratnapura District of the
A GEM PIT.
Province of Sabaragamuwa and the Galle District of the Southern
Province ; buta little is done near Hatton in the Central Province,
and a good many gems were formerly obtained near Nuwara
Eliya and in the Horton Plains. Many districts are now moreor
less exhausted. The process of gemming is briefly as follows :—
A pit is sunk where gem-bearing deposits of gravels are known
to occur; a typical section would show five or six feet of muddy
alluvium, resting on a deposit of gem-bearing gravel not more than
one or two feet thick, and called the i/lam, below which is the
malawa, the decomposed (usually kaolinized) country rock, but
gravels are of course obtained at various depths, from the actual
surface to fifty or sixty feet below. Occasionally two beds of 7//am
CRer 2)
are found, separated by a band of clay. However this may be,
the zllam is removed from the pit and subsequently washed in a
““vemming basket.” This is made of cane, and is of conical form,
about 2 feet wide and 1 foot deep, and has a rim about 23 inches
wide. The washer stands in about 2 feet of water, and holding the
basket in the water gives it a turning movement, depressing the
rim below the water once in every turn, so that the lighter stones
are washed over its edge by the centrifugal movement. Fifteen or
twenty basketfuls are thus washed, and the residue, consisting
only of gems and other heavy minerals,examined. The remaining
material, usually thrown away, is called namobu; it often contains
minerals of scientific interest, and further, it isin this way that
the heavy minerals such as thorianite, containing rare elements,
are obtained.
TABULAR LIST OF GEMS FOUND IN CEYLON.
Corundum.—TIneludes sapphire (blue), ruby (red), star sapphire,
and star ruby. White sapphires have had their original pale
blue or yellow colour discharged by burning. Rubies are almost
always burnt in order to discharge in the same way any trace of
blue colour. Yellow sapphires are “oriental topaz,” and violet
coloured ones oriental amethyst. Pinkish-yellow stones are
called “king topaz.”
Quartz.—Includes rock crystal, amethyst, cairngorm, smoky
quartz, &c.
Spinel.—Green, blue, red. The pink and red varieties are
called balas ruby ; the blue, spinel sapphire.
Chrysoberyl.—Green and yellow ; includes cat’s-eye and alex-
andrite.
Topaz.—Colourless, erroneously called water sapphire; rarely
yellow; pale sea-green, cut as aquamarine.
Othoclase-Feldspar.—Includes moonstone, quarried from the
matrix in the Central Province.
Tourmaline. — Brown and brownish-green and yellow; see
zircon.
Bery/.—Pale sea-green, cut as aquamarine (true aquamarine) ;
the true emerald colour is extremely rare in Ceylon.
Cordierite.—Blue, the true water sapphire ; rarely seen.
Garnet. — Red, pinkish-red, and brownish-yellow (cinnamon
stone).
Zircon.—Green, yellow; the colourless “ Matara diamonds”’ are
got by burning pale zircons and so driving off the colour. Most of
the material called toramalli by gemmers is actually zircon, and
not tourmaline.
K 105-04
( 66 )
VI.—ARRANGEMENT OF THE MINERAL GALLERY.
Of the four wall cases, those facing the entrance are devoted
to rocks ; the two right and left of the door to plumbago, iron
ores, mica, kaolin, &c. Of the seven table cases, the five window cases
contain the systematic collection of Ceylon minerals, beginning
with the elements in the first case on the left-hand, and ending with
zircons, &c., in the fifth case on the right. The two central cases
contain the recent fossils, and collections of a general character
illustrating the geology and mineralogy of Ceylon. Some largeand
interesting specimens are also accommodated on the floor. Geo-
logical photographs and others illustrating the processes of
gemming and iron smelting are hung upon the walls. The collec-
tion has been arranged and largely added to by the staff of the
Mineralogical Survey in 1903 and 1904.
GIANT TORTOISE OF ALDABRA,
(See page 43.)
SPOLIA ZEYLANICA, 67
ON THE PHYTOPHAGOUS AND PARASITIC HYMENOPTERA
COLLECTED BY MR. E. ERNEST GREEN IN CEYLON.
By P. CAMERON.
Plates A and B.
INTRODUCTION.
HE Ichneumonide and other Parasitic Hymenoptera are—
as Dr. Sharp remarks in his volume on Insects (Cambridge
Nat. Hist. Series) —“ One of the most neglected of the great groups
of Insects, though perhaps of greater economic importance to
mankind than any other.” Besides the truly parasitic families,
the group is usually made to inelude the gall-flies—insects which
must be considered as inimical to mankind. It is, however,
with the former families—those that prey upon the vegetarian
insects—that we are chiefly concerned and that form the principal
subject-matter of Mr. Cameron’s paper.
The importance of the Parasitic Hymenoptera will be recog-
nized when it is understood that there is probably not a single
vegetarian insect that does not constitute the host of one or more
species of these parasites. Dr. Sharp states (doc. cit.) that the
destructive “winter moth” (a serious pest of fruit trees) is
known to be attacked by sixty-three distinct species of Hymen-
opterous parasites.
This neglect—by the generality of entomologists—must be
attributed partly to the difficulties of determination and partly
to their somewhat uninteresting exterior, for the family does
not rank among its members many beautiful or remarkable forms.
But the lack of conspicuous external beauty is amply compen-
sated by the interest and complexity of their habits and
development.
In spite of this comparative neglect, nearly 6,000 species of
Ichneumonidz have been described. By far the greater number
of species undergo their early development inside the bodies
of their hosts. The adult female is usually provided with a
prominent slender ovipositor, by means of which the eggs are
inserted into the tissues of the victim. The resulting larve
subsist upon the juices (the lymph or blood), taking up all the
L 8(17)05
68 SPOLIA ZEYLANICA,
nourishment that should go to the building up of the tissues of
the host, until eventually the latter dies of inanition, This
collapse seldom occurs until the contained parasite (or parasites)
is ready to undergo transformation into the pupal state. In the
Ichneumonidse proper the full-grown larva usually spins a
compact cocoon either within the carcass of its victim or by the
side of it.
The early stages of these parasites are not invariably passed
within the body of the host. Three distinct conditions have
been noticed :—
(1) Both egg and larva may be interior.
(2) The egg may be exterior and the larva interior.
(3) Egg and larva may both be completely exterior.
This last condition may be observed in the case of the
parasite of one of our principal tea pests, the “Tea Tortrix”
(Capua coffearia, Niet.) The egg of-this useful little parasite is
attached to the back of the caterpillar, just behind the head.
The young grub fixes itself in the same position and completes
its growth, fully exposed, except for the leafy shelter constructed
by the host. It is noticeable that even those species that attach
their eggs to the surface of the caterpillarsare still provided with
a well-developed piercing ovipositor. With the parasite of the
Tortrix this weapon is employed in piercing the leafy covering
which conceals the victim. It is remarkable, also, that the
periodical moulting of the caterpillar does not dislodge the
parasite.
Besides true insects, spiders are subject to the attacks of
Ichneumon flies.
K. E. G.
FIRST PAPER.
Our knowledge of the Hymenoptera of Ceylon, and more parti-
cularly of the plant-feeding and parasitic species, is very limited.
There is a paper by the Russian Entomologist V. Motsulsky
in the Bull. de la Soc. Imp. des Natur. de Moscow, XXXVI.,
1863, wherein sixty-one parasitic species are described, ineclud-
ing many new genera. The descriptions, however, leave much
to be desired, while it is doubtful if many of the species have
been referred to their proper genera. Motsulsky, for example,
describes two species of Microgaster, but all the species I have
seen, or have been described by recent writers, belong, not to
Microgaster, but to the allied genus Apanteles. Consequently one
is in doubt if the species described by Motsulsky belong to
Microgaster as now limited, or to Apanteles, or even to some new
genus; as is probably the case with the species I have in this
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 69
paper doubtfully referred to Microgaster. The same remark
applies to the species of Microgaster described by Walker. The
systematic position of some of the genera described by the Russian
Hymenopierist is also doubtful in some cases. Thus his genus
Calliopteroma, referred by him to the Jchnewmonide, belongs to
the Encyrtide (Chalcidide) according to that eminent authority
Dr. Wm. H. Ashmead, cf. Proc. U.S. Nat. Mus, XXIII.,152. The
late Mr. Francis Walker described (Ann. Mag. Nat. Hist. 1860,
V1.), a number of species and some new genera, none of which I
have been able to identify, nor has Dr. Ashmead, /.c., been able to
refer them to their proper tribes. I donot know where Walker’s
types are. Apparently they are not in the British Museum,
otherwise Col. C. T. Bingham would have described the aculeates
in his work on the Aculeate Hymenoptera of British India and
Ceylon. In that work he has merely reproduced Walker’s descrip-
tions. Ihave myself (Manchester Memoirs) describeda few species
taken by Mr. G. A. J. Rothney and by Col. Yerbury, and in Proce.
U.S. Nat. Mus. XVIII., Dr. Ashmead has described some species,
mostly reared from Lepidoptera and Coccide, taken or bred by
Mr. Green. And some species have been described by J. O.
Westwood in his Thesaurus Entomologicus Oxoniensis. From
what I have said it is evident that our information regarding the
Hymenoptera of Ceylon is of a very restricted nature—a remark
that applies with even more force to Southern India.
The following species are here described :—
SIRICIDA.
1. Xiphydria striatifrons, n. sp.
BETHYLINA,
Epyris foveatus, n. sp.
3. Rhacoteleia pilosa, n. g. et sp.
> bo
CHALCIDIDA.
4, Spilomegastigmus ruficeps, n. g. et sp.
EVANIID.
Evania interstitialis, n. sp.
Evania peradeniy®, n. sp.
Gasteruption tricoloratum, n. sp.
Gasteruption ceylonicum, n. sp.
ieee iia
: AGA'THIDINA.
9. Agathis kandyensis, n. sp.
10. Agathis oya, n. sp.
11. Agathis ceylonicus, n. sp.
12. Microdus greeni, n. sp.
32.
SPOLIA ZEYLANICA.
CHELONINA.
Chelonus tricoloratus, n. sp.
Phomerotoma hendecasisella, n. sp.
CARDIOCHILINA.
Ernestiella nigromaculata, n. g. et sp.
MICROGASTERIN &.
Microgaster carinicollis, n. sp.
BRACONIN &.
Iphiaulax xanthopsis, n. sp.
Iphiaulax fulvopilosus, n. sp.
Iphiaulax greeni, n. sp.
Iphiaulax ernesti, n. sp.
Iphiaulax kirbyi, n. sp.
Iphiaulax erythroura, n. sp.
Iphiaulax haragamensis, n. sp.
Rhacospathius striolatus, n. g. et sp.
Philomacroplea basimacula, n. g. et sp.
RHOGADINA.
Paraspinaria pilosa, n. g. et sp.
Holcobracon fulvus, n. g. et sp.
Tropobracon luteus, n. g. et sp.
Troporhogas spilonotus, n. g. et sp.
Troporhogas albipes, n. sp.
Troporhogas maculipennis, n. sp.
Troporhogas tricolor, n. sp.
Troporhogas ruficeps, n. sp.
Troporhogas lateralis, n. sp.
Troporhogas trimaculata, n. sp.
CRYPTIN A
Bathycrisis striaticollis, n. g. et sp.
SIRICID A.
1.—NXiphydria striatifrons, sp. nov.
Plate A, fig. 1.
Black, a large irregular mark, broader than long,on the face, two
small irregular oblique marks above the antenna, a line on the
inner eye orbits, with a rounded projecting point in the middle
and dilated above ; on the outer side the line is continued halt
way up the eyes, the upper part projecting obliquely outwards, a
large mark on the top of the temples, irregularly narrowed on the
inner side, the base and lower side of the propleure, the apex, two
small spots on the base of the scutellum,asmallanda large mark
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 7
closely united on the sides of the post-scutellum, a longish mark
on the sides of the firstabdominal segment, asmallone on the third
and fifth, a slightly larger one on the fourth, a large, long, oblique
one on the penultimate, and an oblique mark, transverse behind,
more irregular at the base, and narrowed to a point below on the
last abdominal segment, and a thin curved narrow line on the top
of the mesopleurz, white. Legs dark red, the coxe marked with
white ; the hind tibiz darker, their base and the metatarsus white,
the other joints of the hind tarsi, blackish. The inner side of the
mandibles dark rufous. Wings hyaline, highly iridescent, the
nervures and stigma black, &.
Length 11°5 mm.
Pundalu-oya, February.
Front furrowed in the middle, the furrow widest above; closely,
distinctly striated, the striz becoming strongest below and extend-
ing on to the face. Vertex smooth, shining. Middle lobe of
mesonotum closely rugose ; the apex reticulated; the lateral lobes
irregularly transversely striated. Scutellum coarsely rugose at
the base, the rest smooth and shining. Apex of propleure
sparsely, the mesopleure more closely and strongly punctured
throughout. Transverse marginal nervure received very shortly
beyond the second transverse cubital, which is roundly curved
towards the base of the wings; both the recurrent nervures are
received near the apex of the basal third of the cellule.
The antennz are 15-jointed, have the flagellum thickly covered
with ashort, black, stiff pubescence and taper distinctly and gra-
dually towards the apex.
This species may be known from the two known Indian species
(X. 4-maculata, Cam. & orientalis, West.) by the antenne being
15-jointed, in addition to the other specific differences.
BETHYLIN 43.
2.—Epyris (2) foveatus, sp. nov.
Black, shining, the mandibles,antenne, and legsrufotestaceous ;
wings hyaline, the basal nervures and costa testaceous, the stigma
and stigmal branch darker coloured; the head, thorax, and
ventral surface covered with longish fuscous hair, 6.
Length 4 mm.
Peradeniya, May.
Flagellum of antenne densely covered with long white pubes-
cence, as long as the thorax ; scape about four timesas long as its
thickness at the apex; pedicle longer than wide fully half the
length of the first flagellar joint, which is of the length of the
second ; the last two joints are darker coloured, about three times
12 SPOLIA ZEYLANICA.
longer than thick. Parapsidal furrows and the transverse furrow
at the base of scutellum wide and deep. Post-scutellum with a
deep, semi-circular fovea in the centre at the base. A deep trans-
verse furrow at the base of the metanotum, widened laterally and
bounded at the apex by astout keel. Metanotum witha stout keel
down the middle, stoutly transversely striated ; the apical slope
finely transversely striated in the middle. Pro-and meso-pleure
smooth ; a deep oval fovea near the centre of the mesopleure,
with a smaller one below ; shortly beyond the middle is a deep
depression finely striated in the centre, rounded below, where it
is wider than it is above. Metapleure finely, closely striated.
First abdominal stoutly keeled in the middle at the base and with
a narrower keel on either side; the apex testaceous. Fore legs
and femora slender. Apical tooth of mandibles long, sharply
pointed, the others indistinct. Clypeus projecting, roundly
narrowed at the apex. Basal abscissa of radius straight, obli-
quely sloped ; the apical roundly curved.
This is not a true Hpyris ; but having only asingle 6, I do not
care to found a new genus on it. It has practically only one
long tooth on the mandibles, and in the clypeus projecting at the
apex it differs from the typical species.
KRhacoteleia, gen. nov.
Sub-marginal nervure long, longer than the others united,
marginal nervure very short, half the length of the stigmal, which
has aknob atthe end, the post marginal aboutthree times the length
of stigmal branch. Antenne 13-jointed, the last six forming a
club. Mesonotum and scutellum flat, the parapsidal furrows
distinct, erenulated. Abdomen longish, longer than the head and
thorax united, convex above and below, the sides margined ;
sessile, the basal segment strongly striated, the striz distinct and
clearly separated ; the base of the second segment depressed, stout-
ly striated ; the base of the first clearly margined. Mesopleure
largely and deeply excavated. Scutellum broader than long,
transverse at the base and apex, where there isa crenulated furrow.
The third abdominal segment is longer than the second and fourth,
which are about the same size. Antenne inserted immediately
over the mouth; the scape long. Legs slender, all the tibize one-
spurred, the basal joint of hind tibie as long as the others united,
about three times longer than the second. Metanotum rugosely
punctured, shorter than the scutellum, depressed and transverse
at the apex.
The head is large, rounded in front, transverse behind; the
occiput margined ; temples wide ; ocelli in a triangle. the hinder
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA, 73
almost touching the eyes. Prothorax small. Malar space large.
HKyes gradually narrowed from above downwards. The wings,
when folded, do not reach to the apex of the abdomen.
Allied to Macroteleia and Chromoteleia ; from the former it
may be known by the stigmal vein issuing from near the sub-
marginal, the marginal being very short as it is in Chromoteleia,
but that has two distinct basal cellules; from both it may be
known by the strongly striated basal segments of the abdomen.
3.—Rhacoteleia pilosa, sp. nov.
Plate A, fig. 2.
Black, covered with white pubescence, the antennz except the
apical five or six joints and the legs, except the coxe rufo-
testaceous, wings hyaline, the nervures testaceous, 6 and @.
Length 4 mm.
Peradeniya, August.
Vertex finely, closely aciculated ; the front irregularly rugosely
punctured; malar space closely, obliquely striated. Apex of
mandibles broadly rufous. Hind edge of vertex closely, finely
reticulated ; occiput shining, finely closely, transversely striated.
Mesonotum and seutellum with shallow moderately close, round
punctures, thickly covered with white pubescence ; lateral lobes
of mesonotum clearly separated from the middle one; on the
outer side they are bordered by a crenulated furrow. Apex of
scutellum bordered by a smooth, shining keel. The depression
bordering its sides with stout clearly-separated keels. Pleurz
finely rugose, the mesopleural depression shining, bare, its sides
obseurely striated. The third and following abdominal segments
distinctly closely punctured. Mesosternum strongly closely
punctured. Ventral surface of abdomen closely punctured,
thickly covered with white pubescence.
The pedicle of the antenne is about three times longer than
thick ; the first joint of flagellum is longer than the second and
not quite double the length of the pedicle. The club is clearly
defined ; its last joint conical. The apices of the abdominal
segments are smooth and shining ; the ovipositor is longer than
usual. The 6 appears to have the front more strongly rugosely
reticulated than the &.
CHALCIDIDA.
Spilomegastigmus, gen. nov.
Mandibles bidentate, the teeth blunt. Head and thorax smooth,
the mesonotum only slightly transversely striated. Scutellum
smooth, without a transverse furrow. Antenne long, the scape
74 SPOLIA ZEYLANICA,
slender, its apex reaching to the hinder ocelli. Hypopygium
long, plough-share shaped. Legs long and slender. Apex of
elypeus roundly incised.
This new genus cannot wel] be confounded with any of the
described genera of Megastigmine. It comes nearest to Megas-
tigmus, which may be known from it by the punctured seutellum
with a transverse furrow at the apex, by the distinctly 3-dentate
mandibles and by the shorter antennal scape, the top of which
does not reach to the ocelli. Characteristic is the spotted
abdomen, the red head and the thorax, and the projecting
cultriform hypopygium.
4,—Spilomegastigmus ruficeps, sp. Nov.
Plate A, fig. 3.
Black, the head, antennal scape, prothorax, mesonotum, the
upper part of the mesopleure at the base, and scutellum, red ; the
legs yellow, the femora slightly tinged with rufous; on the sides
of the abdomen are four yellow marks, the second is larger and
rounder than the others, the apical two longer and narrower
than the basal, the wings hyaline, the nervures and stigma black ;
there is a slightly oblique cloud, narrowed below, at the stigma, ?.
Length 6; ovipositor 11 mm.
Kandy, July.
Front excavated in the middle with a keel down the centre, the
sides roundly convex, projecting beyond the eyes; infuscated.
Head much wider than the thorax. Mandibles and sides of
clypeus black. Mesonotum transversely striated, but not closely
orstrongly. Scutellum perfectly smooth. Mesopleurze obscurely
striated. Sheath of ovipositor fringed with long black hair.
EVANIIDA.
5}.—Hvania interstitialis, sp. nov.
Red ; the head and abdomen, except the petiole, black, the four
front legs and hind coxe rufous, the rest of hind legs black.
except the trochanters which are red, mixed with black, the basal
half of metatarsus and calearia which are testaceous and a broad
band on the base of the tibize, which is white; wings hyaline,
the nervures black, the seven or eight basal joints of antennz
testaceous beneath, 6.
Length 5 mm.
Pundalu-oya, October.
Antennal scape as longas the following two joints united ; the
pedicle and first joint of flagellum as long as the second united.
Head thickly covered with white pile; the front and vertex
smooth and shining ; the face, oral region and malar space closely,
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. iB
strongly striated. Apex of clypeus and of cheeks rufo-testaceous.
Mandibles yellow, tinged with testaceous, the teeth black. Hind
ocelli separated from each other by a distinctly greater distance
than they are from the eyes, which very slightly converge above.
Middle lobe of mesonotum with some large scattered punctures ;
the lateral lobes more closely and finely punctured. Scutellum
smooth in the centre, the rest bearing large, round, clearly sepa-
rated punctures. Metanotum closely, reticulated, punctured.
Propleurz smooth, the apex closely striated, the base below with
a few short keels. The dilated lower part of the mesopleure with
shallow,clearly separated round punctures; the edges crenulated.
Sternal process distinctly diverging; the branches stout and
rounded at the apex. Abdominal petiole longer than the rest of
the abdomen, its apical half with scattered punctures, which give
it a rough appearance. On the base of the hind coxe behind is a
smooth shining, raised, elongate space, clearly defined from the
punctured parts and having at the apex a projecting keel, when
pressed together the coxe appear to be united. Tibiz minutely,
sparsely spinose. The long spur of the hind tibiz reaches to
shortly beyond the middle of the metatarsus.
Pronotum transverse in the middle at the base; the sides
broadly rounded, slightly projecting. Cheeks not quite so long
as the antennal scape, parapsidal furrows deep, smooth clearly
defined. Apical abscissa of radius broadly rounded; recurrent
nervure interstitial, as is also the transverse median; the cubitus
is largely bullated at the transverse cubital nervure. There is a
keel on the upper two-thirds of the face, down the centre.
Allied to #. dolichopus, Schl.
6.—Hvania peradeniye, sp. nov.
Length 4 mm. é.
Peradeniya, August.
This species is black with the thorax red as in the preceding
species. The two may be separated thus :—
Recurrent nervure interstitial, oral region rufous at the apex, face
strongly striated and weakly keeled in the middle, hind coxze
rufous above, interstitialis.
Recurrent nervure received beyond the transverse cubital, oral region
entirely black, face weakly striated and strongly keeled in the
centre, hind cox black above, peradeniye.
Scape and pedicle of antennz rufous. Scape of antenne not
much longer than the first joint of flagellum, which is, with the
pedicle, as long as the second. Face closely, weakly striated; a
stout keel on the upper two-thirds; malar space rather strongly,
obliquely striated; thickly covered with white pubescence.
M S(17)05
76 SPOLIA ZEYLANICA,
Front and vertex smooth and shining. Hind ocelli separated
from each other by a slightly greater distance than they are
from the eyes, which converge very slightly above. Shoulders
broadly rounded. Mesonotum with a regular row of punctures
on the inner side of the furrows and a less regular one on the
outer ; the lateral lobes minutely closely punctured in the centre.
Seutellum with a broad smooth band in the middle, the sides
strongly punctured. Hinder division of propleure rugosely
punctured. Mesopleurze with round closely pressed punctures,
except for an oblique smooth band in the middle and a smaller
triangular space on the upper side at the apex. Anterior tibie
and tarsi testaceous, middle trochanters pale testaceous; the
middle coxe and the underside of the hinder pair at the base
rufous ; the base of the hind tibiz and of the hind tarsi white ;
the spurs testaceous; the long spur of the hinder about two-
thirds of the length of the metatarsus; tibiz and tarsi sparsely
spinose. Recurrent nervure received shortly beyond the trans-
verse cubital; lower part of the apical abscissa of the radius
broadly, roundly curved. Abdominal petiole above finely closely
striated; the sides with two stout keels, which become stouter
towards the apex. Hind coxe aboveat the base coarsely punctured,
the middle closely obliquely striated. Apical branches of
metasternal process, straight, obliquely diverging.
Comes close to H. erythrosoma, Sch., also from Ceylon. It may
be known inter alia from our species by its perfectly smooth
abdominal petiole.
7.— Gasleruption tricoloratum, sp. nov.
Black, densely covered with silvery pubescence ; the four front
cox black, the femora rufo-testaceous, the tibiz fuscous, broadly
white at the base, the fore tarsi white, the middle white, infus-
cated at the apex, the hind legs black, except for a narrow white
band near the base of the tibiz. Mandibles testaceous, tinged
with yellow in the middle, the teeth darker coloured ; palpi pale
testaceous, wings clear hyaline, the nervures black: apex of
ovipositor testaceous, ?.
Length 12 mm. ; ovipositor 12 mm.
Peradeniya, May.
Head smooth and shining, covered with a silvery pile. Hind
ocelli separated from each other by the length of the antennal
scape. Thorax thickly covered with silvery pubescence; the
pro- and mesonotum with round, clearly separated shallow
punctures ; opaque, granular. Scutellum with a row of round,
deep fever along the sides. Metanotum with a deep shining
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 77
furrow down the middle; the rest irregularly reticulated. The
third joint of the antenne is about one half the length of the
fourth, and twice the length of the pedicle. Apical half of
mesepleure obscurely reticulated ; on the apex is a deep, shining,
smooth furrow. Second discoidal cellule divided.
8.—Gasleruption ceylonicum, sp. Nov.
Black, the mesopleurz and mesosternum ferruginous ; the legs
black, the fore coxe ferruginous, the fore femora fuscous ; the
base of the fore tibiz and of the tarsi more broadly, white ; the
hind tibie white on the under side at the base. Wings clear
hyaline, the nervures black, the second discoidal cellule divided, ®.
Length 13 mm.; terebra 13 mm.
Pundalu-ova. Febrnary.
Third joint of antenne as long as the scape, more than half the
length of the fourth and not twice the length of the pedicle.
Mandibles rufous. Centre of clypeus covered with pale golden
pubescence. Hind ocelli separated by the length of the third
antennal joint. Thorax thickly covered with silvery pubescence.
Pronotum stoutly keeled down the middle. Middle lobe of
mesonotum closely finely transversely striated ; the apex coarsely,
irregularly reticulated ; the furrows bordering the middle lobe
crenulated ; the furrows bordering the scutellum indistinct.
Metanotum closely transversely reticulated and with a smooth
line down the middle. Shoulders with strong, stout teeth; the
part behind these closely, irregularly reticulated, the middle
depression crenulated. Lower half of mesopleure reticulated.
AGATHIDIN A.
9.—Agathis kandyensis, sp. nov.
Luteous, the antennex, the hind tibiz and the hind tarsi black ;
wings hyaline, the base tinged with yellow, a large cloud at the
base of the stigma, broadest behind and the apex from near the
second cubital cellule dark smoky, the hind wings with a faint
smoky cloud at the apex, 6.
Length 7 mm.
Kandy, May to August.
Head covered with short fuscous hair, lower part of occiput
and the malar space distinctly keeled. Pro- and mesothorax
closely and rather strongly punctured ; the middle lobe raised
in the centre, depressed on either side. Scutellum strongly
and closely punctured ; its apex roundly projecting. Base of
metanotum obliquely depressed, irregularly reticulated ; the
middle strongly irregularly areolated ; the central area irregular;
78 SPOLIA ZEYLANICA.
that next to the spiracular triangular, with a curved keel on the
innerside; the apical slope bordered by are which are widest
below ; the centre with some irreguiar keels. Pleure strongly
and closely punctured ; the crenulated furrow on the mesopleure —
wide, with strong striz ; there isan oblique keel on the centre of
the metapleure.
10.—Agathis oyd, sp. nov.
Length 7°38 mm., ? and 6.
Kandy, July and August. Pundalu-oya, May.
This species is very similar to the preceding ; it may be known
by the apex only of the hind tibie being black; by the middle
lobe of mesonotum being smooth and not raised in the centre, by
the central area on the metanotum being distinctly divided into
a large basal and a smalier apical area; it is joined to the base of
the metanotum by a V-shaped area ; the lateral area is triangular;
on the apical slope is a V-shaped area bordered by a more
irregular V-shaped one. Wings yellowish hyaline, with the
costa, stigma, and nervures yellow on the hyaline parts ; the first
cloud commences at the transverse basal and median nervures, and
is narrowed in front, the apical at the end of the stigma; the
nervures in the hind wings are yellow, with a slight cloud at the
apex. Parapsidal furrows faintly striated.
11.—Agathis ceylonicus, sp. nov.
Luteous, the greater part of the vertex, upper part of occiput,
the third and following segments of abdomen and the hind tibixw
and tarsi, black; the wings yellowish hyaline, the apex witha
fuscous cloud, commencing at the end of the radial cellule ; there
is a small square black mark at the base of the stigma, °.
Length 7 mm.
Kandy, July.
The black antennz thickly covered with short, stiff pubescence ;
they are brownish towards the apex. Face and clypeus thickly
covered with white, the upper part of vertex and occiput with
fuscous pubescence, Thorax smooth and shining, above thickly
covered with white pubescence. Apex of scutellum bifoveate,
the fovexre shallow. On the base of the metanotum are three small
are, the central the smaller, with the sides rounded narrowed
towards the base, the lateral longer, obliquely narrowed towards
the apex ; outside these is a triangular area with the apex on the
inner side; the central area on the apical slope is triangular ; the
two aree outside this are open below ; the outer area large, with
three angles on the outer side.
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 79
12.— Microdus greent, sp. nov.
Ferruginous, the vertex and more or less of mesonotum may
be infuseated or blackish, antennal scape rufous ; the basal half of
‘the flagellum black, the apical testaceous, tinged with yellow :
wings yellowish hyaline to the base of the stigma, beyond that
dark fuscous, the first and second cubital cellules and a curved
cloud below hyaline; the stigma and apical nervures black ; the
hind tarsi infuscated; the basal three segments of abdomen and
the base of the fourth strongly closely longitudinally, striated.?
Length 10mm. terebra 9 mm.
Kandy, July and August.
Head smooth, sparsely pilose; the lower part of the front
deeply bifoveate ; the space separating the two fovee triangular.
Thorax smooth and shining ; the parapsidal furrows deep, smooth.
On the base of the metanotum is a curved furrow forming a
closed area and having a straight keel in the centre, dividing
itintotwo. From its centre two stout keels run to the apex of the
segment forming an elongated area, obliquely narrowed at the
base. Pleurze smooth, the metapleure densely pilose, the lower
edge bordered by a stout waved furrow. Mesosternal furrow
wide, deep, widened at the apex and with some stout transverse
keels.
This species is probably variable as regards the amount of
black on the vertex, thorax, and apex of abdomen. The antennz
are longer than the body.
CHELONINA.
13.—Chelonus tricoloratus, sp. nov.
Black, the antennal scape, and pedicle rufous; the anterior
legs, middle coxe and trochanters, and posterior trochanters rufo-
testaceous, two spots near the base of the abdomen and a broad
band at the base of the hind tibiz pale testaceous. Wings hyaline
to the base of the radius, fuscous beyond it; the nervures and
stigma black. Antenne 23-jointed, &.
Length 4 mm.
Trincomalee Col. Yerbury.
Head and thorax closely finely rugose, covered with a minute
white pubescence. Mesonotum closely reticulated. Scutellum
with some, not very distinct longitudinal strie. Metanotum
irregularly reticulated, more strongly on the apex than on the
base of the basal division; the apical slope shining, smooth,
reticulated above. Base of abdomen longitudinally striated ; the
striz are not numerous and become weaker towards the apex.
Pleure closely reticulated.
80 SPOLIA ZEYLANIGA.
14.—Phanerotoma hendecasisella, sp. nov.
Pallid ferruginous to pale testaceous, the apical segment
ferruginous: antenne infuscated at the apex: wings clear
hyaline, the stigma and nervures fuscous, the former paler at the
base and apex, 6 and &.
‘Length 3 to 4 mm.
Peradeniya July to December.
Bred from beans (Canavalia, sp.) bored by minute pyralid (/)
and from buds of Jasminwm pubescens infested by larve of Hende-
casis duplifascialis, Hmpsn.
Antenne 23-jointed, longer than the body. Head shagreened ;
temples roundly narrowed, occiput deeply roundly ineised; cly-
peus shining, almost smooth, thickly covered with long pale hair;
its sides above with a large, deep fovea. Mesonotum coarsely
shagreened, its apex obscurely striated ; parapsidal furrows only
indicated. The suture at the base of scutellum crenulated.
Scutellum triangular ; its lateral slope stoutly obliquely striated ;
at its apex is a broad shining black transverse stripe. Meta-
notum more coarsely shagreened—almost punctured—than the
mesonotum: there is a not very strong transverse keel across
the middle, above the apical slope; the sides below this project
into a blunt tooth or tubercle. Apex of propleure obscurely
striated ; the lower part, at the apex, depressed, witha few distinct
short keels. Basal two segments of abdomen shagreened ; the
apical at the base finely, closely reticulated—punctured, the
middle and, to a less extent, the apex, finely, closely, longitudi-
nally striated ; the dorsal sutures obscurely crenulated. Second
cubital cellule much narrowed at the apex, the nervures almost
touching there; the basal abscissa of the radius bounding it is
straight and sharply oblique, the abscissa of the cubitus broadly
roundly curved, pale at the base ; the recurrent nervure intersti-
tial; it and the first transverse cubital nervure are pale, but quite
clearly defined. The apex of the hind femora and the hind tibie
ferruginous; the extreme base of the latter anda broad band
above the middle are pallid, almost white. The basal depression .
of the abdomen is bordered by a keel, which is very faint, if not
absent from the middle.
The first and second abscisse of the radius are roundly curved
downwards, the two forming an arc of a circle; the first trans-
verse cubital nervure is straight, oblique to the stigma, then
bends to run along side it to the radius, which it joins close to
its base, quite close to the stigma. The wings are unclouded and
highly iridescent. The second transverse cubital nervure is
very faint and not half the length of the recurrent nervure.
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. Sl
CARDIOCHILINA,
Ernestiella, gen. nov.
Eyes pilose. Malar space large. Front not much excavated,
without keels. Second cubital cellule large, much longer than
wide, wider at the base that at the apex; radius roundly eurved
towards the costa. ‘Transverse median nervure received in the first
cubital cellule, clearly distant from the transverse basal; the
recurrent nervure received in the first cubital cellule. Radial
cellule in hind wings divided. Metanotum areolated. Suturi-
form articulation distinct. Abdomen short: ovipositor short.
Trophi elongate.
The first and second cubital cellules are separated. Parapsidal
furrows deep, the mesonotum distinctly trilobate : the scutellum
raised, not keeled. Areolalarge,7-angled. Theface is notso much
lengthened as in Agathis, the malar space being about one-third
of the length of the eyes. Apex of clypeus broadly rounded: it is
separated from the face by a furrow and is broader than long.
Anal nervure in fore wings interstitial. Antenne about 40-
jointed. Prodiscoidal cellule open at apex ; there is a large cellule
at the base of hind wings. Hind ecalcaria long.
15.—Ernestiella nigromaculata, sp. nov.
Luteous,a large broad mark enclosing the ocelli, three large marks
on the mesonotum, the central the larger, the greater part of the
mesosternum, the antenne and mandibular teeth, black ; wings
hyaline, the apex broadly infuscated, stigma black, yellowish at
the base; the nervures testaceous, the radius pale at the base.
Smooth, shining covered with pale hair. Parapsidal furrows
crenulated at theapex. Scutellar depression with six stout keels,
Behind the large 7-angled area on the metanotum are two small ones
obliquely narrowed behind: on the sides three large irregular ones ;
on the sides of the apical slope a large one, obliquely narrowed
on the outer side, the apex of the middle transverse. Metapleurze
obscurely reticulated. First discoidal cellule at apex half the
width it isat the base. Propleurze deeply excavated, striated below.
Length 6 mm.
Pundalun-oya, March,
MICROGASTERIN A.
16.—Microgasler (?) carinicollis, sp. nov.
Areolet large, rounded in front, transverse below ; cubitus dis-
tinct, radius faint at the base; three cubital cells. Antenne stout,
longer than the body, 19-jointed. Clypeus not separated from the
face, foveate at the sides above. Scutellar depression, with a row
of stout keels; the apex is bordered by a similar row of keels.
82 SPOLIA ZEYLANICA,
Metanotum keeled down the middle ; the apical slope stoutly reti-
culated, almost areolated ; eyes villose ; malar space large. First
abdominal segment with the central area bordered by keels and
with ashorter keelin the centre. Suturiform articulation distinct.
Mesopleural furrow wanting. Tibial spurs short, about one-fourth
of the length of the metatarsus. There is a narrow, but distinct
keel round the centre of the prothorax. Radius and ecubitus in
hind wings indistinct. Shortly below the middle of the mesopleu-
re is a furrow which bifurcates at the apex. On the mesonotum,
in the centre, are three furrows, central being the shorter and it
isalso less distinct. Apices of tarsal joints spinose.
Black : palpiand four front legs testaceous ; the hind femora and
the abdomen, except at the base, red; the basal two-thirds of the
hind tibie and the calcaria white, wings hyaline, the nervures
and stigma black, the latter white at the base. Head and thorax
densely covered with white pubescence, almost opaque. Metano-
tum depressed at the base. The depression at base of post-
scutellum with four stout keels. Post-scutellum depressed at the
base. Frontal depression clearly separated, deep, stoutly keeled
inthe middle. Last joint of tarsi one-half longer than penultimate.
Length nearly 5 mm.
Thisspecies comes closest to Microgaster as now defined, but is
not quite typical.
Bred from larva of Dasychira securis. June.
BRACONINA.
17.—Iphiaulax zanthopsis, sp. nov.
Luteous, the face, oral region, malar space and inner upper eys
orbits, yellow; the antennz very long, black ; hind tarsi fuscous ;
wings yellowish hyaline to the middle of the transverse basal
nervure ; beyond that blackish-fuscous ; the upper part of the first
cubital cellule and a spot below the first transverse cubital nervure
hyaline ; basal half of stigma luteous, ¢.
Length 11; terebra 4 mm,
Elephant Pass, March.
Face smooth, densely covered with long pale hair. Frontal
furrow narrow; vertex thickly covered with long fuscous
hair. Thorax smooth; impunctate, the median segment thickly
covered with long pale hair. Abdomen ovate, broader than the
thorax, short ; the area on the first segment longitudinally rugosely
striolated ;a keel in the centre; basal depression smooth. Basalarea
onthe second segment closely striated, not much narrowed towards
the apex; not limited at the apex; the rest of the segment and the
third closely, strongly, rugosely, punctured ; the lateral depression
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 83
oblique, large, deep, striated ; suturiform articulation, wide, deep,
striated ; the third segment is closely rugosely punctured ; the
furrow on its apex is obscurely striated ; the fourth segment is
punctured, but not closely or strongly; the furrow at its base wide,
deep, striated ; that on itsapex crenulated. The fifth and following
segments smooth ; the apical two are yellowish; on the centre of
the fifth is a deep wide furrow on the apical half ; on the sides, at
the base, is a wide curved striated furrow.
18.—Iphiaulax fulvopilosus, sp. nov.
Luteous, the antenne black; wings yellowish-hyaline, to near
the transverse basal nervure, beyond that dark fulvous; a
hyaline cloud occupying the greater part of the first cubital cellule
and a smaller one along the outer side of the recurrent nervure ;
the base of the stigma fulvous. Pubescence dense, fulvous, ?.
Length 17; terebra 6 mm.
Kandy, June.
Front and vertex sparsely, the face thickly covered with long
fulvous pubescence. Tips of mandibles black. Palpi fulvous,
covered with fulvous hair. Middle of raised part of first abdomi-
nal segment stoutly keeled from near the base; its apex strongly
punctured. Basal area on second segment large, triangular,
obliquely striated, the oblique strize, united by curved short ones,
forming irregular reticulations; the sides of the segment at the
base depressed, without oblique furrows ; its apical part and the
other segments all over closely, longitudinally striated. Suturi-
form articulation narrow, deep, striated, not cleft at the sides ; the
furrows on the following three segments are wider and more dilated
in the middle ; the basal four segments together are as long as the
thorax. Legsstout; the basal four joints of the fore tarsi as long as
the tibiz. Theabdomen thickly covered with fulvous pubescence.
19.—Iphiaulax greeni, sp. nov.
Black, the scape of antenne, head, thorax, and four front legs
red; wings dark fuscons, the nervures and stigma black, ¢.
Length 11; terebra 17 to18 mm.
Peradeniya, August and November.
Face rugosely punctured, a smooth line, gradually widened
towards the apex in the centre; sparsely covered with long fuscous
hair; frontal furrow shallow. First abdominal segment in the
centre with four stout irregular longitudinal keels ; the outer two
unite into one which goes to the apex; the sides of the raised
central part margined ; on the sides of the central keels are irregu-
lar, more or less transverse keels ; the lateral depressions irregularly
transversely striated, The area on the second segment is long,
N 8(17)05
84 SPOLIA ZEYLANICA.
reaching beyond the middle of the segment, becoming gradually
narrowed to a fine point and continued to the apex by a keel ; its
base is irregularly longitudinally striated ; the depression border-
ing it closely strongly transversely striated ; on the outer side is
a long pyriform fovea. Suturiform articulation wide, closely,
strongly striated; the furrow on the apex of the third segment
and on the base of the fourth are striated ; on the apex of the
fourth is a smooth furrow; the third segment is strongly and
closely striated, but not so strongly, as the second; the fourth is
more finely and closely striated. Hind tibie and tarsi densely
covered with black hair. Apex of sheaths of ovipositor white.
Apical abscissa of radius curved, slightly longer than the basal two
united; received in the apex of the first cubital cellule, clearly
distant from the first transverse cubital nervure.
20.—Iphiaulax ernesti, sp. nov.
Head and thorax red; the four front legs yellow, their coxz
tinged with rufous, hind legs fuscous black; wings fuscous,
hyaline, highly iridescent, the nervures and stigma black, ¢.
Length 8; terebra 12 mm.
Peradeniya, October.
Face rugose, thickly covered with long fuscous hair. Temples
obliquely narrowed. Antennal scape rufous, about fourtimes longer
than thick, of equal width throughout. Front deeply furrowed.
Middle area of first segment with two keels, which unite into one
shortly beyond the middle; its sides margined and there is a
curved keel which runs from the central two before they unite ;
the lateral furrows with some scattered transverse keels. The
area on the second segment becomes gradually narrowed to a fine
point shortly beyond the middle; the base finely, distinetly
striated; the depression bordering it has some transverse, not
very distinct, strie ; outside it are four curved longitudinal strie ;
the fovea on the outer side is rounded at the apex and is of equal
width. The third segment is closely, strongly longitudinally
striated ; the suturiform articulation, the furrow at the apex of the
third and at the base of the fourth are striated ; that on the apex
of the fourth is smooth ; there is a J-shaped striated furrow on the
sides of the second segment at the base ; the basal two-thirds of
the fourth segment is finely, longitudinally striated.
A much smaller and more slenderly-built species than J. green;
the fore legs are yellow, not rufous ; the antennal scape is more
slender and does not project at the apex, and the first and second
abdominal segments are finely sparsely striated, not coarsely
striated and reticulated.
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 85
21.—Iphiaulax Kirby, sp. nov.
Luteous ; antennze black ; wings yellowish-hyaline, the apex
with a fuscous border ; the base and apex of stigma black, the
rest of it and the nervures yellow, a small black cloud at the base
of the first cubital cellule, between the radius and the cubitus, ?.
Length 9 mm.; terebra 7 mm.
Kandy.
Sparsely covered with fulvous pubescence. Face irregularly
rugose, covered with pale pubescence. Front and vertex smooth
and shining, bare; the former deeply depressed with a deep,
clearly defined furrow in the centre. First and second abscissa
of radius united equal in length to the third; the cloud at the
stigma may be continued along the basal abscissa of cubitus.
Middle area of first abdominal segment irregularly, longitudinally
striated, intermixed with some transverse striz ; the lateral de-
pression with some transverse keels ; the second segment coarsely
reticulated in the middle ; the basal area triangular, smooth ; the
oblique furrow on the lateral depression crenulated ; the keel at
the basal area short, indistinct. Suturiform articulation stoutly
crenulated, as are also the furrows on the apex of the third and
fourth segments ; they are closely punctured.
22.—Iphiaulax erythroura, sp. nov.
Plate A, fig. 7.
Head, thorax, antennal scape and four front legs rufotestaceous ;
the abdomen black, the fifth and following segments bright
ferruginous, the hypopygium paler, more yellowish at the base ;
wings yellowish-hyaline, paler at the apex; the middle tibize
covered thickly with pale pubescence; the hinder still more
thickly with longer black hair, ?.
Length 12 mm.; terebra 17 mm.
Kandy.
Antennal scape thickly covered with long fulvous hair. Face
coarsely rugosely reticulated. Thorax smooth and shining ; the
apex of median segment black. The central area of basal
segment of abdomen stoutly keeled in the middle; the keel
bordered by stout oblique striz. The second segment is strongly
longitudinally striated ; the basal area small, triangular, smooth,
followed by a stout keel which runs to the smooth apex ; the third
is similarly striated, but with a broader smooth apical area; both
segments are depressed laterally; the suturiform articulation wide,
deep; on the fourth segment, shortly behind the middle, is a trans-
verse furrow which is irregularly striated in the middle. The
basal abscissa of the radius is broadly roundly curved towards
86 SPOLIA ZEYLANICA.
the stigma; transverse median nervure received shortly beyond
the transverse basal. Occiput transverse in the middle, the sides
broadly rounded.
Characteristic of this species is the broadly rounded basal
abscissa—not straight and oblique as usual—of the radius and the
fact of the transverse median nervure being received shortly
beyond the transverse basal. The recurrent nervure is inter-
stitial. The densely haired hind tibiz are also noteworthy.
23.—Iphiaulax haragamensis, sp. nov.
Black; the base and basal half of the sides of the ventral
surface lacteous ; wings fuscous-hyaline, the nervures and stigma
black, ?.
Length 7; terebra 1°5 mm.
Haragam, July.
Head covered with longish hair, fuscous, silvery on the lower
part of the face. Front and vertex smooth ; the former deeply
furrowed. A reddish spot above each antenna. Palpi black,
covered with white pubescence. The pleure and the scutellar
region marked withrufous. The first and basal half of the second
abdominal segment coarsely rugosely punctured ; the following
three strongly, closely, longitudinally striated ; the furrows are
more closely striated ; the last segment white-lead coloured, finely,
closely, transversely striated. Abdomen broad, ovate, as long as
the head and thorax united ; the area on the base of second
segment indistinct, smooth at the base, the apex finely striated.
Rhacospathius, gen. nov.
Plate A, fig. 4.
Transverse median nervure in fore wings received beyond the
transverse basal. Metanotum with three large closed arezx at the
base ; the central keel bifurcates at the apex, forming a triangular
area open at the apex; the rest of the segment and the pleura —
striated. Otherwise as in Spathius.
24.—Rhacospathius striolatus, sp. nov.
Black, the basal two-thirds of the second abdominal segment and
the legs red, the fore legs paler, more yellowish in tint. Wings
hyaline, the stigma and nervures fuscous, ?.
Length 7 ; terebra 5 mm.
Kandy, October.
Mesonotum, except at the base of the central lobe, closely,
strongly, irregularly reticulated. Scutellum minutely aciculated,
the sides with arow of fover, Metanotum irregularly striated and
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 87
reticulated ; the basal aree smooth at the base, the sides furrowed ;
the inner side finely, closely striated along the central keel; the
apical slope irregularly rugosely striated. Propleure strongly
obliquely, the upper part of the mesopleure longitudinally striated;
the lower part, near the base, finely, closely, slightly, obliquely
striated ; the apex rugosely punctured. Basal two-thirds of
mesosternum bordered by a deep furrow. Abdominal petiole
closely, strongly striated. Probrachial nervure in hind wings
interstitial ; the third abscissa of radius longer than the basal two
united. Face, except for a smooth shining line in the centre,
closely transversely striated ; the front closely striated, except at
the sides; the vertex and temples smooth and shining. Legs
sparsely haired ; the hind coxe striated on the outer side,
Philomacroplea, gen. nov.
Apex of sixth abdominal segment broadly, deeply, roundly
incised in the middle. First abdominal segment sessile, excavat-
ed at the base, keeled on the top; it,and the second segment, keeled
down the middle; the sutures of the segments narrow: the
apical segments are curled downwards ; the abdomen short, broad,
not unlike the abdomen of Chelonus. Metanotum smooth and
shining, stoutly keeled down the middle. Parapsidal furrows
distinct, deep, crenulated. Last joint of hind tarsi not much
dilated, as long as the second. Ovipositor short, broad. Antennz
longer than the body, the second joint of flagellum about twice
longer than thick. Malar space large, furrowed down the middle.
Third abscissa of radius about three or four times longer than the
basal two united. Temples obliquelynarrowed. <A broad furrow
behind the ocelli.
Theabdomen is more like what it is in the Chelonine than in the
Braconine. It is allied to Yropidobracon, Baryproctus, and
Plesiobracon, all of which have a stout keel in the centre of the
metanotum ; but, among other differences, the present genus
should be readily separated from them by the keeled base of the
first abdominal segment, by the first and second being keeled
down the centre; by the last segment being roundly broadly
incised, and by the very long third abscissa of the radius.
. 25.—Philomacroplea basimacula, sp. nov.
Head, thorax and the middle of the first and second abdominal
segments broadly rufo-testaceous, the mesosternum and the base
of metapleure above blackish; wings hyaline, the nervures and
stigma fuscous ; the legs pale testaceous, almost white, ? and 6.
Length 3°5 mm.
Bred from Macroplea elisa.
88 SPOLIA ZEYLANICA.
The long black antennze densely covered with a short white
microscopic pile, as are also the thorax and abdomen. The upper
half of pleurz closely, minutely punctured ; the scutellum is less
distinctly punctured, its sides densely pilose. Metanotum very
smooth and shining. Abdomen very closely, uniformly, and
rather strongly punctured ; the apices of the third and following
segments narrowly rufous at the apex. Scutellum triangular,
broad at the base. The second cubital cellule is widened at the
base, the first transverse cubital nervure being very obliquely
sloped. The anal nervure in the fore wing issues from shortly
below the middle of the transverse nervure. The lower half of
the base of the mesopleure is depressed compared with the upper.
RHOGADIN A.
Paraspinaria, gen. nov.
Abdomen with seven segmenis ; the first longer than broad, the
central area large, wide; the second segment without an area;
the sides depressed at the basal half; ovipesitor short, hardly
projecting. Metanotum irregularly reticulated ; the sides at the
apex, with a long stout tooth. Hind wings with an enclosed
probrachial cellule, reaching shortly beyond the middle of the
' basal abscissa of the cubitus ; the nervure broadly rounded at
the apex ; the cubitus and radius complete ; there are no other
apical nervures. Lower part of mesopleure depressed, clearly
separated from the upper; the hind edge of mesosternum has a
stout conical tooth. Radial cellule long reaching to the apex of
the wing; apical abscissa of cubitus longer than the basal two
united ; recurrent nervure received in the first cubital cellule,
Antenne longer than the body; temples obliquely narrowed ;
occiput almost transverse, margined. Palpi long.
The affinities of this genus are clearly with Spinaria. That
genus may be known from it by the abdomen having only
five segments, these being spined and longitudinally striated ; its
anal nervure is interstitial and the prothorax has a spine as a
rule. The antenne in my genus are placed opposite the middle
of the eyes, which are incised. The pronotum is large; its
centre at the base is triangular, the rest is depressed at the sides
and apex. Abdomen as long as the head and thorax united.
26.—Paraspinaria pilosa, sp. nov.
Plate A, fig. 9.
Luteous ; antennal flagellum, apical third of hind tibiwe and
the hind tarsi black; wings hyaline ; the nervures and stigma
yellow ; the costa near the stigma and a small square cloud
below it black, ?.
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 89
Length 9 mm.
Kandy, June.
Shining, thickly covered with white pubescence. Parapsidal
furrows deep, the middle lobe of mesonotum clearly raised.
Base of propleure with three stout keels, the middle one not
reaching, like the others, to the lower edge. Scutellar depression
large, deep, with a stout central keel and a narrower oblique one
on either side. Metanotum with four arez on the centre of the
basal region; the basal large, longer than broad, the apical
smaller, dilated on the outer side; the other arez not clearly
defined. The apical lateral teeth are large, longer than the width
at the base; they become gradually narrowed, with the apex
rounded. First abdominal segment irregularly reticulated, the
sides less strongly than the centre ; the second is also irregularly
reticulated; there is a keel down the centre, reaching to the apex;
the lateral keels are stouter and shorter. Suturiform articulation
stoutly, closely striated. The apical depression of the middle
lobe of mesonotum is reticulated ; there is a distinct furrow on
the apex of the middle lobe. Metasternum bordered by a stout
keel.
Holcobracon, gen. nov.
Lower part of mesopleure with a distinct crenulated furrow.
Median segment closely reticulated, keeled down the middle ;
the sides at the apex below projecting into a blunt tubercle.
Mandibles stout, edentate, the apex transverse in front, oblique,
the basal half slightly roundly incised. Clypeus separated from
the face by a depression, roundly convex, thickly covered with
long hair. Malar space longer than the eyes. Temples broad.
Occiput margined, more distinctly on the sides than above;
roundly incised. Scutellum flat, legs stout; the fore cox
largely project below, the trochanters issuing from the upper
part; fore tibie stoutly toothed. Anal nervure in fore wings
interstitial ; in the hind wings there is a discoidal nervure which
runs from the prediscoidal to the apex, there being thus three
longitudinal nervures in the hind wings; there is a large
closed prebrachial cellule, which becomes gradually wider
towards the apex. Abdomen broad, the basal segments closely
striated ; suturiform articulation deep, roundly curved.
The apical abscissa of radius reaches to the apex of the wings
and is twice the length of the second ; the transverse median
nervure is received distinctly beyond the transverse basal ; the
recurrent in the first cubital cellule.
Therearethree genera known to me which possess the anomalous
character of having a curved nervure issuing from the prebrachial
90 SPOLIA ZEYLANICA.
transverse nervure to the apex of the hind wings. These genera
may be separated as follows :—
(a) Hind coxe with two teeth... Acanthobracon, Cam, (Szep.).
(6) Hind cox toothless. Apex of mandibles stoutly
toothed ; sides of median segment toothed at the sides
above the apex, the scutellum convex... 7Tvichiobracon.
Apex of mandibles not toothed, transverse; sides of
metanotum not toothed above ; the scutellum quite
flat... Holcobracon.
These three genera form a natural tribe, Holcobraconint, allied
to Doryctini, which should be readily known by the hind
‘wings having three longitudinal nervures running to the apex
of the wings; the abdomen sessile, the mesopleure with a
distinct crenulated furrow along the lower border, the metanotum
toothed or taberculated above or below, closely reticulated and
the basal three segments of abdomen longitudinally striated,
with a distinct curved crenulated suturiform articulation and
oblique furrows as in [phiaulax, and stoutly toothed fore tibie.
.
27.—Holcobracon fulvus, sp. nov.
Plate A, fig. 6.
Rufo-fulvous, the antenne black; wings yellowish-hyaline to
near the apex of the transverse basal nervure, with an oblique,
irregular cloud in the first cubital cellule, extending below into
the discoidal ; the stigma luteous; hind wings yellowish-hyaline,
the apical third smoky, ?.
Length 18 mm.
Kandy, November.
Rody and legs thickly covered with long pale pubescence.
Face and oral region closely, finely rugose, thickly covered with
fuscous pubescence, the clypeus with long fuscous hair. Front
and vertex smooth, shining, almost bare. Pronotum punctured,
the punctures large, clearly separated, and each with ajlongish hair.
Mesonotum and scutellum smooth, sparsely haired. The scutellar
depression with five stout keels ; the central the larger. Metano-
tum with a keel down the centre, closely reticulated, the
reticulations shallow, irregular in shape, some hexagonal. The
centre of propleure with some irregular keels. The lower furrow
on the mesonotum closely crenulated ; the oblique furrow below
the tubercles wide, deep, and with some stout keels. The upper
part of the metapleure irregularly, coarsely reticulated ; below the
furrow strongly, irregularly punctured. Legs thickly covered
with long pale pubescence. Basal three segments of abdomen
strongly closely striated ; the suturiform articulation deep,roundly
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 91
curved, crenulated ; there is an oblique furrow on the base of
the second segment; the base of the fourth and fifth segments are
closely striated.
The metapleurz and base of abdomen are yellowish ; the man-
dibles and ocellar region are black.
Tropobracon, gen. nov.
First abscissa of radius slightly longer than the second, both
together as long as the first transverse cubital nervure. Recurrent
nervure widely distant from the apex of the cellule. Second
cubital cellule much narrowed in front, being there less than
half the length it is posteriorly ; apical abscissa four times the
length of the basal two united, reaching to the apex of the wing.
Parapsidal furrows deep, clearly defined, not reaching to the
apex. Basal two jointsof flagellum equalin length. Last joint of
hind tarsi longer thanthe third, as long as the first. First segment
of abdomen short, broader than long, the sides and top of the
apical slope margined.
Abdomen coriaceous, short, broad; the area on the second seg-
ment large, reaching to the apex ; the ovipositor short, not much
longer than half the length of the abdomen.
This genus should be readily known by the recurrent nervure
being received at a distinct distance from the apex of the cubital
cellule, by the deep, clearly defined parapsidal furrows, keeled
basal slope of first abdominal segment, by the short basal two
abscisse of radius, and by thesecond cubital cellule being narrowed
in front and short. Habrobracon, Ashm., seems to be its nearest
ally, but inter alia, that genus should be known by the “basal joint
of hind tarsi being about the length of the third, shorter than the
second,” while in my genus it is double the length of the third
and much longer than the second. The parapsidal furrows are
much deeper and more clearly separated than usual.
28.—Tropobracon luteus, sp. nov.
Luteous, the antenne black, the hind tibie and tarsi infuscated ;
wings hyaline, highly iridescent, the nervures and stigma black ;
the recurrent nervure received shortly beyond the middle of the
cellule, ?.
Length 4 mm.; terebra 1 mm.
Pundalu-oya, March.
Face shining, aciculated, covered with a white pile; a distinct
curved furrow over the clypeus. Malar space long, two-thirds
of the length of the eyes. Front and vertex smooth. Temples
a) 8(17)05
92 SPOLIA ZEYLANICA.
oblique, rounded. Occiput rounded, not transverse. Mesonotum
and scutellum shining ; its middle lobe reaches to the base of the
apical third, is then united to the apex by a stout keel, witha
narrow striated border on either side. Metanotum closely rugose.
First abdominal segment short ; its base rather abruptly sloped, its
sidesand apex margined. The segments are closely, finely punc-
tured ; the area on the second segment large, becoming gradually
narrowed towards the apex; the furrows striated, not uniting at
the apex; there are no lateral furrows at the base. Suturiform
articulation striated : there are no furrows on the other segments.
There is a distinct curved furrow below the middle of the
propleure.
Tropor hogas, gen. nov.
Transverse median nervure received near the base of the cellule,
close to the transverse median ; recurrent nervure received near
the apex of the cellule; second cubital cellule twice longer than
wide, of equal width throughout ; anal nervure not interstitial.
Eyes large, clearly incised on the inner side; malar space small.
Temples short, oblique. Occiput sharply margined, transverse.
Palpi very long, slender, pilose, 4- and 5-jointed- Metanotum
with tworoundly diverging keelsonthe base. A depression on the
lower side of the mesopleure. Basal three segments of abdomen
closely longitudinally striated ; the basal two witha keel down the
centre; suturiform articulation crenulated ; there are crenulated
furrows on the fourth, fifth, and sixth segments ; the base of these
segments is depressed, the apex of the segments being raised and
clearly separated from the base of the following. . Hypopygium
large,cultriform ; ovipositor shortly projecting, the sheaths stout ;
antenne longer than the body, slender, pilose, over 50-jointed,
The abdomen is fully twice the length of the thorax ; the legs long
and slender; the femoranarrowed at the base. The first abscissa
of the radius is not half the length of the second ; the third is the
longest and is curved upwards. The tarsi longer than the
tibie; the metatarsus longer than the two following joints
united.
In Dr. Ashmead’s system (/.c.) this genus could only be con-
founded with Rhogas; that genus may be known from it by the
transverse median nervure being widely distant from the
transverse basal, by the metanotum having a stout keel down the
centre, not two at the base, by the shorter abdomen, with the
segments not sharply separated ; the shorter and stouter legs
with the tarsi not longer than the tibie; and the hypopyginm
is not large and cultriform,
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 93
29.—Troporhogas spilonotus, sp. nov.
Pallid testaceous, the legs paler in tint; the ocellar region,
marks, more or less distinct, on the mesonotum, the metanotum to
the top of the apical slope, the base of the mesopleure and large
marks on the base of the abdominal segments, the penultimate
segment entirely, black. Wings clear hyaline, the nervures pallid
testaceous, the basal half of the stigma fuscous, ¢.
Length 7 to 8 mm.
Peradeniya.
The amount of black on the thorax varies; the hind coxe and
trochanters may be marked with black. The entire body and legs
thickly covered with white pubescence. Firstand second abdomi-
nal segments strongly and closely striated throughout ; the third
is less strongly and closely striated, the striz becoming fainter
towardstheapex. The pleurz may be largely marked with black :
the mesopleural furrow is striated. Parapsidal furrows wide and
deep; the apex of the middle lobe of the mesonotum is depressed
and with a distinct furrow in the centre. Scutellar depression
large, deep ; a stout keel in the centre and a narrower oblique one
onthe sides. Scutellum narrowed towards the apex. Metanotum
finely, irregularly rugose ; the apex with some irregular keels.
30.—Troporhogas albipes, sp. nov.
Rufo-testaceous, the oral region, lower outer orbits, and legs
white ; the four hind coxe brownish-red :; a broad band on the
thickened apex ofthe hind femora (but not reaching to the apex)
black ; scape of antenne rufous, the flagellum yellowish-white.
Wings hyaline, slightly suffused with fuscous ; the apex witha
fuscous narrow cloud round the edges ; the stigma testaceous, the
nervures fuscous, 6.
Length 9mm.
Kandy.
Sides of front stoutly, obliquely striated; face irregularly
wrinkled. Propleure stoutly striated ; the mesopleure wrinkled
and irregularly striated, closely and strongly below ; the meta-
pleure closely, finely, rugosely punctured. Prosternum yellowish,
depressed in the middle and witha longitudinal keel there, which
is bordered by a row of fovex. Parapsidal furrows striated ; the
apex of the middle lobe closely striated and reticulated. Scutellum
convex, roundly narrowed towards the apex, which is rounded ;
a keel runs from the sides of the apex to the wings; the space
inside this is depressed and striated in the middle. Median
segment closely rugosely punctured, the punctures running into
reticulations. Back of abdomen closely rugosely striated ; the
furrows are more strongly and distinctly striated.
94 SPOLIA ZEYLANICA.
31.—Troporhogas maculipennis, sp. nov.
Plate A, fig. 5.
Testaceous, the part between the ocelli, the greater part of the
mesopleure, and the apical two segments of the abdomen blackish ;
the upper part of the thorax infuscated. Wings hyaline; the
apex smoky, before and behind from before the second trans-
verse cubital nervure, the middle from beyond it; there is a
narrow cloud along the anal nervure, which is thick and black,
the apex white and thinner, @.
Length 12 mm.
Kandy, July and August.
Vertex at the sides of the ocelli closely, the front more strongly
obliquely striated. Face thickly covered with long fuscous hair.
Mesonotum with afew scattered punctures, shining, covered with
a short blackish pubescence. Scutellar depression not very deep ;
the central keel not very stout. Metanotum at the sides of the
base closely rugose, the centre and the rest to the middle of the
apical slope irregularly reticulated. Propleurze with some stout
oblique striz ; the mesopleure obscurely punctured, the centre
with an oblique depression; metapleure closely rugosely
punctured. Basal four segments of the abdomen closely longi-
tudinally striated; the fifth closely and strongly, the others
sparsely punctured.
32.—Troporhogas tricolor, sp nov,
Plate A, fig. 8.
Antenne black. Head rufo-testaceous, the vertex behind and
the occiput black; the oral region, mandibles, and palpi pale
yellow. Thorax black, the mesonotum, scutellum, and upper part
of pleurz rufo-testaceous. The first abdominal segment, the base
and the sides, more broadly, especially behind, of the second, the
sides of the third and fourth, the fifth, except for a line on the
sides at the apex, and the apical, entirely yellow, this being also
the case with the ventral surface. Four front legs whitish yellow :
the middle tarsi infuscated ; the hind legsblack. Wings hyaline;
the stigma and nervures black, ?.
Length 7 to 8 mm.
Kandy, June.
Face closely punctured, the middle at the sides of the raised
part striated. The vertex at the sides of the lower ocellus
transversely, the front more coarsely, obliquely striated. Mesono-
tum and scutellum smooth, neither punctured nor striated. The
basal keels on the metanotum are stout and have two transverse
ones between them ; the part on either side, almost smooth, on the
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. oF
outer side distinctly striated ; the apical slope strongly, irregu-
larly reticulated. Base of propleure and of mesopleure strongly
striated ; the metapleurz rugose, the apex striated. First abdomi-
nal segment closely punctured, finely striated on either side of the
keel ; the second strongly, irregularly striated ; the suturiform
articulation strongly, regularly striated; the third and fourth,
except at the apex, closely striated; the other segments closely
punctured, The radius and cubitus from the transverse cubital
nervure are white.
33.—Troporhogas ruficeps, 8p. nov.
Head and thorax rufo-testaceous, the antenne dark fuscous ; the
abdomen pale yellow, the first segment except round the apex
and more narrowly on the sides, a curved mark on the apex of the
second segment,and the greater part of the third, fourth, and fifth
on the back, black ; the four front legs yellow, tinged with rufous ;
the hind coxe, femora, except at the base, and the apex of the tibiz
broadly, black, the rest of them whitish yellow ; the tarsi infus-
cated ; wings hyaline, the stigma and nervures fuscous, ?.°
Length 6°5 mm.
Peradeniya, December.
Face thickly covered with white pubescence ; the sides striated ;
clypeal fovez deep. Vertex behind the ocelli closely, finely
striated. Pro- and mesonotum smooth, not striated, a furrow on
the apex of the latter; scutellum impunctate. The two keels
on the base of metanotum form a A-shaped area, with a thin
transverse keel at the apex. Basal five segments of abdomen
closely longitudinally striated.
34.—Troporhogas lateralis, sp. nov.
Rufo-testaceous, the upper part of the pleurx, the sides of
metanotum broadly, the sides of the first abdominal segment
broadly on the basal, more narrowly on the apical half, a line on
the sides of the second, the line becoming gradually wider from
the base to the apex, the third at the sides and its apex from
shortly behind the middle, black. Legs pale yellow; the apex of
middle femora infuscated, the apical half of posterior black.
Wings hyaline, the nervures and stigma black, .
Length 5 mm,
Peradeniya, August.
Face raised in the centre, striated on either side. Side of front
obscurely striated. Pro- and mesonotum shagreened ; parapsidal
furrows shallow; an indistinct transverse furrow at the apex.
96 SPOLIA ZEYLANICA.
Secutellar depression wide, roundly curved, deep, obscurely
striated. Metanotum closely rugose, the basal keels indistinct, the
sides with some oblique strize. Pro-and mesopleure for the greater
part closely reticulated ; the metapleure finely rugose. The first
and second abdominal segments are closely striated, the third
obscurely so, the other segments smooth ; the suturiform articu-
lation indistinct, only indicated through the apex of the second
being raised. The first abscissa of the radius is half the length of
the second; the second cubital cellule is half the length of the
third,
35.—Troporhogas trimaculata, sp. nov.
Rufo-testaceous, the pleure paler, more yellowish in colour,
the ocellar region and three large marks on the mesonotum black ;
wings hyaline, highly iridescent, the nervures and stigma black:
the costa, base and apex of stigma, and the transverse basal
nervure testaceous, ?,
Length 8 mm.
Kandy, July.
Smooth, shining, impunctate. Scutellar depression large,
shallow, divided by a keel. Scutellum smooth, rufous at the
apex. Post-seutellum rounded, conspicuous, smooth, dark rufous.
Metanotum weakly punctured, a keel down its centre, the keel
indistinct at the base. The basal four segments ofabdomen closely
punctured ; the fifth indistinctly so; there is no keel on the basal
segments.
A NEW GENUS OF CRYPTINA FROM CEYLON.
Bathycrisis, gen. nov.
Abdominal petiole of equal width throughout, the post-petiole
not being dilated ; it is long and slender. Median segment with
two transverse keels ; its spiracles about three times longer than
wide. Transverse median nervure in hind wings broken below the
middle. Disco-cubital nervure broken by a stump of a nervure.
' Areolet of almost equal width throughout. Apex of clypeus
depressed, broadly rounded, above clearly separated from the face,
which is dilated in the centre above, there being thus a depres-
sion between its apex and the clypeus. Flagellum of antenneze
densely pilose, the first joint longer than the second. Front
deeply excavated; a stout keel above each antenna. Eyes dis-
tinctly, roundly incised on the inner side ; the malar space
moderate. Parapsidal furrows deep, wide, reaching to the middle.
There isa wide, deep, curved furrow on the base of the metanotum ;
there is no area there. Thorax strongly rugosely punctured.
The abdomen is more slender than usual.
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 97
I only know the 6 of this genus. It comes near to Ospryncho-
tus, which may be known from it by the rostriform head, with
long malar space, and by the disco-cubital nervure not being
broken by astumpofanervure. In The Entomologist, 1903, 182, I
described an Osprynchotus peronatus from India. I was not then
acquainted with the type of Osprynchotus from the Cape. Now
that I have seen it, I find that my Indian species is a Linnuceras,
Tasch.=Osprynchotus Schmied. non Spin. (Ent. Nachr. XVI., 85).
Linnoceras may be known from the genus here described by the
transverse median nervure in hind wings being broken at, not
below, the middle, by the more elongate clypeus, not séparated
from the face, large projecting labrum, longer, more slender,
mandibles, of which the upper tooth is much the longer. In
Bathycrisis the mandibles are short, thick, with two short stout
teeth of equal length. In Dr. Ashmead’s tables, Bull. U.S. Nat.
Mus. XXIII., 40, Osprynchotus is placed in the division with the
transverse median nervure in hind wings “broken distinctly below
the middle, usually far below the middle,” whereas it is broken
shortly above the middle.
36.—Bathycrisis striaticollis, sp. nov.
Black, the inner orbits, broadly below, narrowly above, a nar-
row line in the centre of the outer, a mark, longer than broad,
transverse at the apex and interrupted in the middle above by a
fovea, a mark in the centre of the clypeus, mandibles, except at
the apex, a small mark on either side of the pronotum, a smaller
one in front of the tegule and tubercles, lemon-yellow; legs, with
the greater part of the femora, the tibiz and the tarsi, except at
the apex, rufo-testaceous; the coxe and femora marked below
with yellow. Wings hyaline, the nervures and stigma black, 6.
Length 9 mm.
Trincomalee, Col. Yerbury.
Head covered with long white hair. Face closely punctured,
the clypeus smooth, the lower part smooth and shining ; the part
immediately below the ocelli strongly transversely striated, and
the striz curved.
The vertex behind the ocelli closely punctured ; the occiput
margined, the temples wide, obliquely narrowed. Pronotum
*strongly irregularly striated. Mesonotum closely punctured; in
the depressed centre the punctuation is closer and runs into
reticulations. The scutellum is less closely punctured and more
shining than the mesonotum. Metanotum closely rugosely
reticulated. Pleurz closely strongly punctured.
YS SPOLIA ZEYLANICA.
ON THE PHYTOPHAGOUS AND PARASITIC HYMENOPTERA
COLLECTED BY MR. E. ERNEST GREEN IN CEYLON.
SECOND PAPER.
By P. CAMERON.
fa ii following species are described in this paper :—
ICHNEUMONIN 2.
Melanichneumon kandyensis, n. sp.
Hoplismenus ceylonicus, n. sp.
Haliphera fulvipes, n. sp.
Aluina erythropus, n. g. et sp.
Deniya pleuralis, n. g. et sp.
Tanyphatnus multimaculatus, n. g. et sp.
Wea Pw Pe
Stictichneumon macariez, n. g. et sp.
CRYPTIN &.
8. Buodias rufipes, n. sp.
9. Melcha cinctipes, n. sp.
10. Melcha annulipes, n. sp.
11. Melcha varibalteata, n. sp.
12. Melcha reticulata, n. sp.
13. Melcha erythropus, n. sp.
14. Melcha maculiceps, n. sp.
15. Friona rufipes, n. sp.
16. Friona bituberculata, n. sp.
17. Fenenias erythropus, n. sp.
18. Skeatia acutilineata, n. sp.
19. Skeatia cyclosiz, n. sp.
20. Skeatia panthone, n. sp.
21. Bathythrix rubriornatus, n. sp.
22. Bathythrix striatus, n. sp.
23. Clitiga excavata, n. g. et sp.
24. Clitiga forticornis, n. sp.
25. Earrana lutea, n. g. et sp.
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 99
OPHIONID 2%.
26. Ophion bicarinatus, n. sp.
27. Pleuroneurophion erythrocerus, n. sp.
28. Eniscospilus spilonotus, n. sp.
29. Eniscospilus melanocarpus, n. sp.
30. Eniscospilus xanthocephalus, n. sp.
31. Eniscospilus unilineatus, n. sp.
32. Eniscospilus dasychire, n. sp.
33. Eniscospilus horsfieldi, n. sp.
34. Spilophion maculipennis, n. g. et sp.
35. Paniscus intermedius, n. sp.
36. Paniscus orientalis, n. sp.
37. Paniscus levis, n. sp.
38. Campoplex greeni, n. sp.
39. Limnerium taprobanicum, n. sp.
40. Nototrachys reticulatus, n. sp.
41. Clatha longipes, n. g. et sp.
42. Trichomma nigricans, n. sp.
TRYPHONIN &.
43. Bassus orientalis, n. sp.
44. Rhorus spinipes, n. sp.
PIMPLIN 2.
45. Epirhyssa ornatipes, n. sp.
46. Theronia maskeliye, n. sp.
47. Erythrotheronia flavolineata, n. g. et sp.
48. Kchthromorpha ornatipes, Cam.
49. Xanthopimpla taprobanica, n. sp.
50. Xanthopimpla kandyensis, n. sp.
51. Xanthopimpla parva, n. sp.
52. Xanthopimpla minuta, n. sp.
53. Philopsyche albobalteata, n. g. et sp.
54. Lissotheronia flavipes, n. g. et sp.
55. Charitopimpla annulipes, n. sp.
56. Lissopimpla rufipes, n. sp.
57. Tanera annulipes, n. g. et sp.
58. Lissonota greeni, n. sp.
ICHNEUMONIN &.
1.—Melanichneumon kandyensis, sp. nov.
Black, the upper inner orbits, narrowly below, more broadly above,
the line extending round the top and narrowly down the upper part
of the outer, a short line at the bottom ; a line on the pronotum, two
P 8(17)05
100 SPOLIA ZEYLANICA.
lines, narrowed in front, on the centre of the mesonotum, the apical
half of the scutellum, the mark dilated laterally at the base, post_
scutellum and the apices of the second to fifth abdominal segments
narrowly, the penultimate more broadly, and the last entirely, yellow ;
the second segments broadly at the base rufous ; antenne with a
broad white band in the middle ; legs red ; the four front cox and
trochanters yellow ; the hind black, red, and yellow. Wings hyaline,
slightly tinged with fulvous, the nervures and stigma black, 9.
Length 15-16 mm.
Kandy, July.
Head smooth, mandibles red, black at apex. On the centre and
on the sides of the clypeus isa reddish mark. Prothorax and meso-
thorax closely punctured ; the scutellum smooth. Median segment
more closely and strongly punctured ; the apical slope thickly covered
with white hair ; the areola open at the base, twice longer than
broad ; the inner side furrowed ; it is of equal width throughout
and has the apex roundly curved inwardly. Disco-cubital and
recurrent nervures broken by a stump of a nervure. Apical half of
post-petiole strongly punctured ; the gastrocceli shallow, yellowish.
Agrees fairly well with Melanichnewmon. Scutellum not quite flat,
depressed at base and apex round, longer than the width at the base.
2.—Hoplismenus ceylonicus, sp. nov.
Black; the face, clypeus, the orbits all round, mandibles, palpi,
a line on the pronotum, tubercles, two lines on the centre of the
mesonotum, the apical half of scutellum, the base of the mark trans-
verse, apex of post-scutellum, and the apices of all the abdominal
segments, pale yellow ; the apex of the first segment has the line
broader than it is on the third and following ; on the apex of the
second the band is twice the width it is on the first and there is a
broader band on its base. Legs red ; the four front coxe and trochan-
ters yellow ; the hind coxe black, the apex red below, above they
are yellow. Wings hyaline, the stigma and nervures black. Antennal
scape and a broad band on the centre of the flagellum white ; the
flagellum brownish beneath, especially on the basal half, 6.
Length 14 mm.
Kandy.
Face and clypeus thickly covered with short, white pubescence
and sparsely punctured ; labrum white, largely projecting. Pro-
mesonotum and scutellum smooth, the latter obliquely sloped at the
baseandapex. Propleurze andmesopleurzesmooth. Base of metano-
tum and areola smooth, the rest strongly punctured ; the posterior
median area irregularly rugose ; the apical slope and metapleure
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 101
thickly covered with white hair, the latter strongly punctured.
Abdomen smooth, except the second segment, which is weakly
punctured ; gastrocoeli smooth, shallow.
This species agrees fairly well with Hoplismenus, having the scutel-
lum as in that genus ; but the areola is not ‘‘ wider than long’’ as in
the diagnosis of Ashmead : it is fully longer than wide, open at the
base, and the posterior median area projects roundly into it. The
lateral teeth are small, but distinct. The clypeus is short, transverse
at the apex ; the labrum projects.
3.—Haliphera fulvipes, sp. nov.
Black; the face, clypeus, mandibles, palpi, upper inner orbits
narrowly, the lower half of the outer broadly—more broadly below
than above—a narrow line on the pronotum, tubercles, a mark on
the tegule, scutellum, a line on the centre of the metanotum,
rounded and narrowed at the top and bottom, one-third of it in the
areola, two-thirds on the posterior median area, a broad line on the
first and a narrower line on the second abdominal segment, lemon-
yellow. Legs fulvous, the coxe black; the fore trochanters
yellow, the middle yellow, black behind, the hinder entirely black.
Wings hyaline, highly iridescent, the nervures and stigma black.
Antennal scape yellow, the flagellum fuscous beneath, 6.
Length 12 mm.
Maskeliya, August.
Face and clypeus distinct, but not very closely punctured, sparsely
covered with white pubescence; front and vertex wrinkled irregularly.
Mesonotum closely, finely, but distinctly punctured ; the scutel-
lum with some scattered punctures. The basal three arex of
metanotum finely, closely, irregularly, transversely striated; the
lateral almost smooth at the base ; the posterior median obscurely
striated at the base, the rest strongly transversely striated ; the
lateral areze with some stout oblique irregular striz. Propleure indis-
tinctly, the mesopleure more strongly and closely and the metapleurze
still more closely and strongly punctured, the latter two more or less
closely striated. Petiole smooth ; the second and third segments
closely, but not strongly punctured; the gastrocceli longish, smooth,
and shining; the apical segments densely covered with short blackish
pubescence ; the apical half of the last yellow.
Aluina, gen. nov.
Scutellum rounded, not flat, its basal three-fourths stoutly keeled.
Areola longer than broad, wider at the base than at the apex ; the
keels roundly curved, the apex slightly rounded inwardly ; its
102 SPOLIA ZEYLANICA.
surface stoutly striated. Abdominal petiole smooth, shining,
impunctate. Ovipositor long, as long as the apical three segments
united. Base of flagellum slender, the joints elongate ; occiput
deeply, widely, roundly incised, sloping,above obliquely from the
ocelli. Disco-cubital nervure angled in the middle, broken there by
a stump of a nervure; the two abscisse straight, oblique, not
rounded.
The head is large ; the eyes large, projecting, distinctly narrowed
below; the malar space moderate. Apex of clypeus transverse ;
labrum hidden. Scutellum longer than broad, rounded at the base
and apex. Face flat, only slightly dilated in the middle, not
separated from the clypeus. Abdomen with the second and third
segments punctured ; longer than the head and thorax united, its
apex acutely pointed.
‘Allied, in some respects, to Cillimus and Hxephanes. It has the
long, projecting ovipositor of the latter, but not its filiform antenne ;
but Exephanes has not the scutellum stoutly keeled ; Cillimus has
the ovipositor short, has the face tumid, not narrowed behind the
eyes; the clypeus is armed at the apex with asmall tooth and the
scutellum short. |
4.—Aluina erythropus, sp. nov.
Plate A, fig. 11.
Black ; the face, clypeus, the orbits all round, the outer broadly
below, a line on the raised centre of the pronotum, scutellar keels,
the sides of the scutellum narrowly, the apex more broadly, post-
scutellum, the apex of the petiolar area, a broad line on the sides of
the apical slope of metanotum, a triangular spot behind the spiracles,
the lower edge of the propleurz, tubercles, a large mark, narrowed in
the middle on the lower part of the mesopleure, a large oblique
mark on the centre of metapleurz, the base of the first abdominal
segment, two spots on its apex, the apex of the second broadly, two
broad lines of almost equal width on the third, two shorter, smaller
ones, obliquely narrowed on the inner side, the greater part of the
sixth and the whole of the apical two, pale yellow. Antenne black,
the under side of the scape and a broad band on the flagellum white.
Legs red, the four front coxee and trochanters yellow. Wings hyaline,
the stigma and nervures black, the costa at the base of the stigma
fuscous, ¥.
Length 11-12 mm.
Pundalu-oya.
Head smooth and shining, the face and centre of clypeus sparsely
weakly punctured ; the front in the centre slightly raised, almost
keeled. Mesonotum closely punctured, the furrows distinct on the
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 103
basal half. Scutellum sparsely punctured. Base of metanotum
almost smooth; the areola with four siout, curved, transverse
keels, the base and apex with a few broken ones, the lateral arez
stoutly obliquely, the posterior median transversely striated; the
base of the spiracular area punctured, the rest closely, obliquely
striated. Apex of propleurz below stoutly striated, above and at
the base punctured ; the base in the middle with some stout, clearly
separated striz, the lower part smooth and shining. Abdomen very
smooth and shining except the second and third segments, which are
closely punctured ; the former stoutly striated at the base.
In the example described, one mandible is black, the other yellow.
Deniya, gen. nov.
Temples almost obsolete ; occiput almost transverse. Eyes
large, projecting. Apex of clypeus rounded, behind not separated
from the face. Antenne dilated beyond the middle. Parapsidal
furrows distinct, crenulated. Apex of mesonotum reticulated.
Scutellum longer than its width at the base, convex, stoutly keeled
to beyond the middle ; the apex with an oblique slope. At the base
of the post-scutellum are two triangular depressions, bordered by
stout keels. Base of metanotum, deeply obliquely depressed at the
base. Areola longer than wide, transverse at the base, the apex
angularly turned inwardly ; the sides near the apex angled. Post
petiole and the following four segments closely punctured ; gastrocceli
large, deep. Areolet 4-angled ; disco-cubital nervure not broken
by a stump of a nervure ; transverse median nervure received
behind tranverse basal. Sheaths of ovipositor broad, projecting.
This genus comes close to Alwina. The two may be separated
thus :—
Temples wide, occiput widely, deeply incised, apex of clypeus
transverse, transverse basal nervure interstitial ; sides of
areola rounded, not angled, parapsidal furrows indistinct,
the apex of mesonotum smooth, post-petiole smooth, Alwina.
Temples very narrow, occiput transverse, apex of clypeus
rounded ; sides of areola not rounded, angled; parapsidal
furrows distinct, apex of mesonotum reticulated, post-
petiole punctured, Deniya.
5.—Deniya pleuralis, sp. nov.
Plate A, fig. 10.
Black ; face, clypeus, mandibles, except at apex, palpi, the inner
and upper orbits, the lower half of outer broadly, a broad line on the
pronotum, tegule, scutellum, the sides of the apical slope of the
metanotum, a line on the lower edge of the propleuree, mesopleuree
104 SPOLIA ZEYLANICA.
from shortly above the middle, mesosternum, tubercles, the apices
of all the abdominal segments—the line on the fourth interrupted—
pale yellow. Legs pale fulvous, the anterior largely tinged with yellow,
the apex of the hind femora, base of hind tibiz, their apex broadly
and the tarsi, black. Wings hyaline, the stigma and nervures
black,?.
Length LO mm.
Peradeniya.
Scape below and a broad ring on the flagellum white. Head
smooth and shining, the face and clypeus sparsely punctured
and haired. Thorax smooth and shining, except the metapleure
and the spiracular area which are closely strongly retjculated,
punctured ; the areola with a stout keel in the centre of the apical
two-thirds ; the metapleure thickly covered with white pubescence.
Post-petiole and the second to fourth abdominal segments closely
punctured.
Tanyphatnus, gen. nov.
6. Areola open behind, more than twice longer than wide, separ-
ated from the lateral arez, the apex almost transverse, the top of the
posterior median area being bluntly rounded. Apex of metano-
tum with a gradually rounded slope. Scutellum not flat, roundly
depressed at the base and apex. Post-petiole in the centre finely,
closely, longitudinally striated ; gastrocceli deep; there are no thy-
ridia. There are eight abdominal segments; cerci long, stout ;
the genital armature much larger than usual; the ventral fold
extends to the apex of the fourth segment. Areolet 5-angled ; the
disco-cubital nervure almost broken by a stump of a nervure. Apex
of clypeus transverse, not separated from the face ; labrum largely
projecting. Upper tooth of mandibles much longer than lower,
projecting twice the length of the latter beyond it; it is sharply
pointed. Spiracles about four times longer than wide.
The temples are moderately broad, rounded ; malar space short ;
the antenne are not much longer than the abdomen, serrate.
Abdominal petiole slender, longish, the post-petiole not clearly
separated. Legs moderately stout. Abdomen slender, more than
twice the length of the head and thorax united; the sides of the
segments spotted with yellow ; the last segments not spotted.
This genus, or sub-genus, has the striated post-petiole of /chneu-
mon sensu str. Comparing it with /. luctatorius, the type given for
Ichneumon s.str., its areolais longer, narrower, and not ofequal width,
it being wider at the base and narrowed roundly in the middle ;
the metanotum has a more gradually rounded slope ; the first
segment of the abdomen is more slender, especially at the apex; the
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 105
scutellum is much more distinctly raised—more convex, and clearly
longer than wide ; the temples are shorter and not so obliquely
narrowed, the malar space smaller, the eyes being longer, and the
upper tooth of the mandibles is longer and more sharply pointed.
6.—Tany phatnus multimaculatus, sp. nov.
Black, the face, clypeus, mandibles, palpi, the inner orbits, the
outer, narrowly above, broadly below, the upper and lower edge of
prothorax, prosternum, the sides of scutellum, narrowly at the base,
more broadly towards the apex, scutellum, a broad band of equal
width on the base of metanotum, extending from the base to the
posterior median area ; the apical slope, except the latter area, the
yellow extending on to the pleure, a mark behind the spiracles, the
lower third of the mesopleure, the apex of the first abdominal seg-
ment and broad marks, closely continuous, on the apices of the other
segments, yellow. Four front legs rufo-yellow, their cox and tro-
chanters yellow; the hind legs rufous, the coxze black, broadly yellow
at the base above, the base of the tibiae broadly yellow. Wings
hyaline, the stigma and nervures black, é.
Length 13 mm.
Pundalu-oya.
Face and clypeus punctured, but not strongly or closely. Front
with some minute punctures. Mesonotum sparsely, irregularly
punctured. Base and areola of metanotum smooth; the rest
closely and strongly punctured, thickly covered with long fuscous
hair. Pleure closely and strongly punctured, the meta irregularly
striated. Basal segment with the post-petiole closely finely striated
in the middle ; the second to fourth more strongly on the basal
half.
The gastroceeli are large, deep, smooth. Transverse median
nervure interstitial. The areola is not separated behind, where
there is no oblique depression ; it is clearly separated from the
lateral arez, is slightly narrowed in the middle, more than twice
longer than wide, and transverse at the apex ; the apex of the seg-
ment has a gradually rounded slope, without teeth. The ventral fold
is on segments two to four.
Stictichneumon, gen. nov.
Clypeus separated from the face by a deep, wide furrow which
unites with the lateral foveze ; its apex not quite transverse. Are-
ola horseshoe-shaped, longer than wide, clearly separated behind;
the petiolar area confluent with lateral. Scutellum roundly convex ;
roundly sloped at the base and apex ; its basal half keeled. Petiole
106 SPOLIA ZEYLANICA.
strongly, closely punctured, as are also the following three segments;
gastroceeli deep, widely separated. Areolet 5-angled ; transverse
median nervure interstitial ; disco-cubital nervure slightly broken.
There is a distinct malar space. The upper tooth of the mandible is
distinctly longer than the lower. Antennz shorter than the body.
Temples obliquely, roundly narrowed. Post-petiole not clearly
separated, becoming gradually wider towards the apex; it is
strongly punctured. There is a distinct depression at the base of
the metanotum. Legs short and stout.
The body is black, spotted with yellow ; it is more closely and
strongly punctured than usual. Temples obliquely narrowed.
This genus should be readily known by the roundly convex scutel-
lum, keeled to the middle, and by the clypeus being separated
from the face by a deep transverse furrow. The middle area of the
petiole is raised, separated from the sides. The apex of the clypeus
is not quite transverse, the middle being slightly dilated
7.—Stictichneumon macarie, sp. nov.
Black, the face, except for a large mark on its lower half, its sides
produced above, the sides of clypeus, inner eye orbits, lower two-
thirds of outer, base of mandibles, palpi, a line round the base of the
prothorax, one on the pronotum, tegulze, two lines on the mesono-
tum, scutellar keels, two small spots on the base of scutellum, a
line on its apex, post-scutellum, two oblique spots on the apical
slope of metanotum, tubercles, a mark on the lower part of the
mesopleurz, contracted in the middle, the apical part shorter and
more oval than the basal, which is narrowed at the apex, an irregular
spot on the apex of the metapleure, transverse marks on the sides
of the basal four abdominal segments, the centre of the penultimate
and the whole of the last, yellow. Legs black, the four front coxze
and trochanters yellow ; their femora and tibie obscure testaceous
infront. Wings clear hyaline, the stigma and nervures black,é.
Length 10 mm.
Maskeliya, July.
Bred from pupa of Macaria, sp.
CRYPTIN&.
8.—Buodias rufipes, sp. nov.
Plate A, fig. 13.
Black. the legs red ; a broad. band on the antennz, the upper eye
inner orbits, an interrupted line on the base of the pronotum, scutel-
lar keels, scutellum, metanotal spines and the apices of the basal
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 107
three abdominal segments and of the sixth more broadly, whitish
yellow. Wings hyaline, the nervures and stigma black, °.
Length 14 mm.; terebra 4 mm.
Kandy, July.
Face and, to a less extent, the base of clypeus irregularly rugose ;
the apex of clypeus smooth and shining. Sides of front coarsely,
obliquely striated. Mesonotum smooth, shortly, densely pilose.
Base of metanotum finely irregularly striated ; the rest stoutly
reticulated—striated ; the spines long, stout. Propleurz stoutly,
closely striated ; the mesopleure irregularly striated at the base,
the rest finely striated—reticulated, but not closely ; the meta-
pleure obliquely striated.
9.—Melcha cinctipes, sp. nov.
Black, face, clypeus, mandibles except at the apex, palpi, the
upper inner orbits, malar space, the upper and lower sides of protho-
rax broadly, metapleurz and apical half of metanotum, the apices
of the basal three segments of the abdomen and the apical pale
yellow ; the scutellum and the keels lemon-yellow. Four front
legs pale yellow ; the hind cox pale yellow, black above, the
trochanters black ; the femora red, the extreme apex black ; the
tibiz black, with a white band near the base, tarsi white, the basal
joint to near the apex, and the apical joint, black. Wings hyaline,
the stigma and nervures black, 6.
Length 7-5 mm.
Peradeniya, August.
Base of metanotum smooth, irregularly punctured to the keel ;
the rest closely reticulated. Propleurze with some stout striz in
the middle behind. Mesopleurz obscurely punctured, with a striated
band in the centre above. Lower part of metapleurz irregularly
punctured—treticulated. The second and following segments of the
abdomen are closely punctured. Tarsi spinose.
Antennal scape fulvous ; flagellum densely covered with short,
thick, black pubescence. There is a yellow line on the centre of the
outer orbits.
10.—Melcha annulipes, sp. nov.
Black, an ovoid mark on the centre of the face, the broad end
above and rounded, clypeus, palpi, mandibles except the teeth, the
upper inner orbits from the antenne, pronotum broadly, tegul,
tubercles, scutellum, metanotum from shortly behind the basal
keel, the top forming two rounded lobes, the yellow extending on to
the metapleurze below the bottom of the lobes and being dilated at
the base below, the apex of the first and second abdominal segments
Q 8(17)05
108 SPOLTA ZEYLANICA.
broadly and the apical four segments entirely, yellow ; the petiole
rufous. Legs rufous, the four front cox and trochanters white ;
the hind tibize fuscous, with a broad ring near the base, the hind
tarsi, except the apical joint, white. Wings hyaline, the stigma
fuscous, the nervures darker, °.
Length 6-7 mm. ; terebra 2mm.
Peradeniya, September.
Antenne broadly ringed with white ; the scape and base of flagel-
lum rufo-testaceous. The part below the ocelli closely, irregularly,
longitudinally striated, keeled in the middle ; the depressed front
with two stout smooth keels. Face finely, irregularly rugose.
Mesonotum finely, closely, rugosely punctured. Scutellum smooth.
Base of metanotum irregularly rugose ; the apex distinctly trans-
versely striated, the strive clearly separated. The centre of propleurz
stoutly striated, the mesopleurze closely, irregularly, obliquely
striated and aciculated ; the metapleure closely, strongly, obliquely
striated. The second and third segments of the abdomen closely,
uniformly punctured.
11.—Melcha varibalteata, sp. nov.
Black, the median segment, apex of mesopleur below, and petiole
rufous ; face, clypeus, mandibles, palpi, lower third of outer orbits,
base of propleurz, pronotum, mesosternum, tegul, tubercles, the
usual plate under the hind wings, the apices of the five abdominal
segments and marks in the centre of the apical two, yellow, the
yellow on the abdominal segments backed with testaceous behind.
Four front legs pale fulvous, the coxz and trochanters white ; the
hind coxe fulvous, their trochanters black ; femora rufous, tibiz and
tarsi fuscous, the tibize broadly rufous at the base. Wings hyaline,
the apex slightly smoky, the stigma fuscous, the nervures darker, 6,
Length 5 mm.
Kandy, July.
Head smooth and shining. Thorax smooth, except the apical
slope and sides of metanotum behind the keel, which are irregularly
rugose. Middle lobe of mesonotum depressed in the centre, and
having a distinct fovea, longer than wide ; the apex at the end of the
parapsidal furrows with a row of fovee. The petiole is black at
the apex, the centre yellowish, as compared with the sides.
12.—Melcha reticulata, sp. nov.
Black, the face, clypeus, mandibles, palpi, the lower fourth of the
outer orbits, malar space, the inner orbits to near the antenne,
a broad line on the apical two-thirds of the pronotum, tegule.
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 109
tubercles, antennal keels, scutellum, the usual mark behind the
wings, the apical slope of the metanotum except for a black mark in
the centre, the apices of the basal four abdominal segments and the
apical entirely, yellow. The four front legs yellow, tinged with
fulvous, the hinder rufo-testaceous, the apex of the coxe, trochanters,
apical half of tibiz, and the tarsi blackish. Under side of antennal
scape testaceous. Wings hyaline, the nervures and stigma black, 6.
Length 7 mm.
Matale, July.—Reared from Psyche subteralbata.
Head shining, impunctate. Mesonotum minutely punctured.
Parapsidal furrows closely, transversely striated ; the part where
they end irregularly punctate—reticulated. The base of metanotum
smooth, the rest strongly, closely reticulated. Propleurz and meso-
pleure smooth; the metapleure irregularly reticulated. The
second and third segments of the abdomen are closely punctured ;
the others smooth, densely pilose.
13.—Melcha erythropus, sp. nov.
Plate B, fig. 2.
Black, the upper inner orbits broadly, more broadly below than
above, a line on the lower part of the inner, the greater part of the
clypeus, labrum, mandibles except the teeth, palpi, a line on the
apical three-fourths of the pronotum, tegule, tubercles, scutellum, a
large crescent-shaped mark behind the wings, the apices of the basal
three abdominal segments and a narrower line on the apex of the
sixth, yellowish-white. Legs rufous, the anterior cox and trochan-
ters white ; the greater part of the hind tarsi blackish. Wings clear
hyaline, the nervures and stigma black. Median segment, apex of
mesosternum and base of metapleure, rufous, ¢.
Length 7 mm.; terebra 2 mm.
Peradeniya, August.
The sixth to twelfth joints of the antenne are ringed with white
except above. Face closely, irregularly, rugosely striated. Clypeus
obscurely punctured. Front with a distinct keel down the centre,
striated on either side, the striz stronger and more irregular above.
' Pronotum and mesonotum smooth and shining ; the propleure in
the middle stoutly striated ; the mesopleurz except behind, closely,
finely reticulated. Metanotum behind the keel smooth and shining :
the rest closely, rugosely reticulated. Metapleurz closely, obliquely
striated, more strongly above than below the furrow. The second
and third segments of the abdomen are finely, closely punctured ;
the others smooth and shining. Tarsi closely spinose. Areolet
square, the apical nervure faint. Base of mesosternum yellow, the
110 SPOLIA ZEYLANICA.
rest black : the second transverse cubital nervure faint. The base
of the first abdominal segment is rufous.
A specimen from Kandy is 12 mm. long, has a stripe on the sides
of the fourth abdominal segment, and the penultimate is entirely
white.
14.—Melcha maculiceps, sp. nov.
Black ; the upper inner orbits, a somewhat pyriform mark on the
centre of the face, the thickened end at the top and rounded, the
clypeus except on the sides and apex, mandibles broadly at the base.
tegule, tubercles, scutellum, the apical slope of the metanotum, the
top of the yellow part narrowed, transverse above, with the sides
slightly oblique, and the apices of the abdominal segments, yellow.
Legs rufous, the fore cox and trochanters yellow, the apical joint
of the tarsi black. Wings hyaline, the nervures and stigma black, ¥.
Length 8 mm. ; terebra 2 mm.
Peradeniya, October. Bred from cocoons of Chrysomelid beetle
feeding on orchid.
The sixth to tenth joints of antennz white below. Face closely
and distinctly punctured. The upper part of the front closely, longi-
tudinally, irregularly striated—reticulated. Mesonotum minutely.
closely punctured ; the apex of the middle lobe closely, strongly.
longitudinally striated, the striated part broad and transverse at
the apex ; separated from the scutellar depression by a smooth
raised part. The scutellar depression is deep and bears stout keels.
Base of metanotum irregularly, closely, rugosely punctured ; the
area with a stout irregular keel in the centre ; the rest is closely
reticulated. Pleurz closely punctured, the propleurz on the lower
half stoutly striated ; the sternum bordering the furrow is on the
basal half, closely, finely, transversely striated. Sides of post-
petiole closely, longitudinally striated.
15.—Friona rufipes, sp. nov.
Plate B, fig. 1.
Black, the face, except for a triangular large black mark in the
centre, clypeus, except for a black line, dilated in the middle, at the
apex, labrum, base of mandibles, palpi, the upper orbits to the end
of the top, the lower half of the outer more broadly, malar space, a
line on the pronotum, tegule, scutellum, a reversed T-shaped line
on the apical, striated part of the metanotum, the cross pieces being
on the apical slope, a semi-circular mark on the lower, apical part of
the mesopleurz, a smaller mark, longish, rounded above, transverse
below, under and in front of it ; a large mark, rounded in front.
gradually widened towards the outer side behind the wings, an
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 111
elongated mark, narrowest behind, on the metapleure and the
apices of the abdominal segments, pale yellow. Legs rufous ; the
four front coxze and trochanters yellow ; the hind tarsi white,
fuscous at the base, the apical joints black, Wings hyaline, the
nervures and stigma black, &.
Kandy, July
Length 13 mm.; terebra 4 mm.
Antenne black, broadly ringed with white. Front with a distinct
central keel ; on the top and bottom it has, on either side, three or
four irregular, curved keels. Face punctured, the centre irregularly,
weakly striated. Thorax smooth and shining. The metanotum
behind the keel and the pleure closely, distinctly striated as usual.
Median segment and hind coxe and trochanters covered with long
black hair. Areolet one half longer than the width at the base.
The hind coxe are marked above at the base with yellow : in the
é they are black, except at the base above, the middle coxe have
a large black mark on the outer side at the apex above. The apex
of the metanotum at the white transverse line is roundly dilated.
16.—Friona bituberculata, sp. nov.
Length 13 mm. 6.
Kandy, July.
In colouration this species resembles closely F. rufipes, but the
pleurz are not marked with yellow, the hind coxe are red, without
black or yellow, and the face wants the black central mark. Other-
wise the two may be separated thus :
Apex of metanotum not raised, near the apex armed with two
stout teeth or tubercles, which are enclosed by the yellow
apical band ; the apical transverse band as long as the basal
central line, this being narrowed towards the base ; pleura,
immaculate, not strongly or closely striated ; the base of the
first and the whole of the apical two segments, white, bituber-
culata, Cam.
Apex of metanotum raised transversely, not tuberculated: the
apical transverse line narrow, the longitudinal one widest at:
the top, pleure marked with yellow, distinctly striated face :
the base of the first abdominal segment not yellow; the
apical two segments only yellow at the apices, rufipes, Cam.
There is a triangular mark on the centre of the metanotum, the
narrow end at the apex ; the striz on it are irregular and run into
reticulations. The striation on the front is not very strong. Face
yellow. Clypeus black, with a yellow mark, transverse, rounded at
the sides in the centre. Legs red ; the hind trochanters black. Areo-
let not very large, almost square (smaller than in rufipes, which
receives the recurrent nervure shortly behind the middle).
1 Ba I. SPOLIA ZEYLANITCA.
17.—Fenenias erythropus, sp. nov.
Plate A, fig. 14.
Black, very shining, the centre of the face and clypeus, the latter
more widely than the former ; the inner orbits narrowly from near
the bottom and round the top, the lower two-thirds of the outer,
base of mandibles and palpi, a narrow line on the apical half of the
pronotum, the base below, a transverse small mark on the apex of
the middle lobe of the mesonotum, scutellar keels, apex of scutellum,
the mark longer than broad ; a line on the centre of the apical slope
of the metanotum, followed downwards, on either side, by a similar
line of the same length and united to a mark on the middle, which is.
longer than broad and of equal width ; tubercles. a pyriform mark
on the lower part of the mesopleure near the base, a smaller, some-
what similar mark on the lower side at the apex, a stripe along the
lower side of the furrow, the usual mark under the hind wings. an
oblique mark near the apex of the metapleurz and narrow lines on
the apices of the abdominal segments (the apical interrupted), pale
yellow. Legsred; the anterior at the base and the hind tarsi yellow-
ish. Wings hyaline, the nervures and stigma black, °.
Length 8 mm.; terebra 4 mm.
Peradeniya, September.
Face and upper part of clypeus distinctly punctured, the former
more closely and rugosely above. Vertex punctured in the centre ;
the front clearly separated from it and with a steep slope. Mesono-
tum distinctly, but*not very closely or coarsely, punctured ; the
furrows deep, wider and striated towards the apex ; they do not
extend near tothe scutellum, which is finely, very sparsely punctured,
its lateral slope is stoutly, obliquely striated at the base, the rest
with some elongated punctures. Base of metano‘um with a smooth,
deep, oblique depression ; in the middle is an indistinct U-shaped
keel ; the centre is irregularly reticulated; the sides much more
finely and regularly reticulated. The apical slope is depressed in
the centre and closely, transversely, irregularly striated—reticulated ;
there is no transverse basal keel; the apical is not very distinctly
defined, especially in the centre ; the teeth are broad and blunt.
Propleure strongly striated except for a punctured space above and
the base below, which is smooth. Mesopleure closely punctured,
coarsely, irregularly, longitudinally striated at the base above, irregu-
larly, closely reticulated in the middle ; the apex with a large smooth
and shining space. First abdominal segment smooth, shining, the
apex with some minute scattered punctures ; the second segment
closely and distinctly punctured, except at the apex ; the basal half
of the third is more closely and finely punctured,
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA, 113
The areolet is minute, closed at the apex ; the transverse median
nervure is received behind the transverse basal; the transverse
median nervure in hind wings broken below the middle. Antenne
long, ringed with white, slender. Metapleural keel reaching close to
the hind coxee. Temples very short, obliquely narrowed. Head not
much wider than the thorax. The basal transverse keel on the meta-
notum is only indicated in the middle.
This species may be separated from the type of the genus thus :—
Metanotal teeth stout ; metanotum with three yellow marks ina
triangle; the mark on the mesonotum oval, longer than
broad, hind tarsi fuscous, albomaculatus, Cam.
Metanotal teeth small; metanotum with the upper part of the
apical slope margined with yellow ; the mark on the meso-
notum transverse, small, broader than long, erythropus, Cam.
In albomaculatus the clypeus is more convex and not so iransverse
attheapex. In Feneniasthe frontis more deeply and widely depress-
ed, the depression reaching closer to the ocelli, than in Skeatia.
18.—Skeatia acutilineata, sp. nov.
Plate A, fig. 12.
Black, face, clypeus, base of mandibles, palpi, the inner eye orbits
to the end of their top, the outer from near the top, the yellow line
becoming gradually wider, malar space, the base of the prothorax
narrowly, the middle of pronotum broadly, tegule, a mark, longer
than broad and of equal width in the centre of the mesonotum,
scutellar keels, scutellum, a triangular mark—the narrow end
above—its base shorter than the sides and laterally united by a
short line to a large mark on the sides of the apical slope, enclosing
the spines and going on to the pleurz, the apices of all the abdominal
segments, tubercles, a large oblique mark on the centre of the meso-
pleure, roundly contracted above and below and narrowed at the
apex, mesosternum, anda mark on the metapleure below the keel,
widest and oblique at the apex, yellow. Antenne broadly ringed
with white in the middle. Legs rufous, the coxe and trochanters
yellow, the hind cox broadly black at the apex above, this part
joined to the base on the lower side by a broad curved line ; the apex
of the hind femora narrowly, the base of the tibiz still more narrowly,
their apex more broadly and the apical joints of their tarsi, black ;
the hind tibiz are infuscated. Wings hyaline, the nervures and
stigma black ; areolet minute, longer than wide, °.
Length 12 mm.; terebra 3-4 mm.
Kandy, July.
Front irregularly rugosely punctured except below the ocelli; the
centre keeled. Face distinctly, but not very closely, punctured:
the clypeus more sparsely punctured above, smooth below. Thorax
114 SPOLIA ZRYLANICA.
closely and strongly punctured ; the metapleure more strongly than
the rest, the metanotum closely reticulated, most strongly and more
irrecularly on the apex. The first abdominal segments sparsely, the
others closely and uniformly punctured.
Characteristic of this species is the acutely narrowed top of the mark
on the metanotum; in the other species it is rounded.
19.—Skeatia cyclosie, sp. nov.
Black, the face, clypeus, mandibles at base, palpi, the upper eye
orbits to the end of the top, the outer from shortly above the middle,
malar space, a broad line on the pronotum, a line on the lower edge
of the propleurz, a small mark, longer than broad, transverse at the
base, rounded at the apex on the centre of the mesonotum, scutellar
keels, scutellum, the sides of the metanotum at the apex broadly,
united above by a rounded line, which has the rounded top broader
than the sides ; tubercles, a large mark on the mesopleure, sharply
contracted in the middle, the basal portion being the larger,
mesosternum and a large mark in the centre of the metapleure, and
the apices of the abdominal segments, yellow. Legs fulvous ; the
coxe and trochanters yellow; the base and a mark on the outer side
of the hind coxz, under side of trochanters, apex of! hind femora,
the base of their tibiz more narrowly and their apex more broadly,
black ; the hind tarsi white. Antenne broadly ringed with white.
Wings clear hyaline, the stigma and nervures black, ¢.
Length 12 mm.; terebra 2-3 mm.
Kandy. Bred from cocoonof Cyclosia panthona, a zygeenid moth.
Front with a distinct keel ; the part next to the keel finely rugose,
outside obscurely striated. Face and top of clypeus punctured,
strongly, but not closely. Mesonotum closely, finely, rugosely
punctured, Scutellum smooth; a small triangular black mark at
the base. Metanotum closely, rugosely reticulated ; the basal area
smooth, wider than long. Propleuree stoutly striated, obliquely
above, below more strongly longitudinally; the mesopleurz and meta-
pleure closely punctured. Base and centre of first abdominal
segment aciculated, the sides of post-petiole closely punctured and
striated ; the other segments closely punctured.
The Malay S. varipes closely resembles this species. They may
be separated as follows :—
The metanotal area small, greatly narrowed at the apex, the top
of the yellow line not dilated, the large line on mesopleure
not contracted in the middle, at apex united to the yellow
on the mesosternum, varipes, Cam.
The metanotal area large, not greatly narrowed at apex, the top
of yellow line greatly dilated ; the large line on mesopleurz
contracted in the middle, not united to the yellow on the
sternum, cyclosi@,
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 115
20.—Skeatia panthonee, sp. nov.
Black ; face, clypeus, mandibles at the base, palpi, inner orbits,
their top, the lower two-thirds of the outer more broadly, malar
space, the edge of the pronotum, the lower edge of the propleure
more broadly, an irregular small mark on the middle of the meso-
notum, scutellum, post-scutellum, scutellar keels, the sides of the
apical slope of the metanotum, united above by a narrow line to a
spot which is longer than broad, rounded above, transverse
below, tubercles, a moderately sized roundish mark on the lower part
of the mesopleurz at the base, a smaller—half the size—somewhat
conical one on the lower side of the apex, a large mark behind the
wings, partly on the top, partly on the pleure, a quadrangular mark
twice longer than broad on the centre of the metapleure, the meso-
pleurze and the apices of the abdominal segments, pale yellow. Legs:
the four anterior yellow, tinged with fulvous, the hinder fulvous.
the cox yellowish, black at the base and with a black large mark
on the apex above, the lower half the larger, and projecting beyond
it at the apex ; the trochanters, apex of femora, the base of tibiz
more narrowly, their apex more broadly, black ;_ the tarsi yellowish.
Wings hyaline, the stigma and nervures black, 6.
Length 11 mm.
Kandy. Bred from Cyclosia panthona.
Face and upper half of clypeus strongly punctured. Front stoutly
irregularly striated. Mesonotum strongly, closely, the scutellum
sparsely punctured. Metanotum behind the keel closely rugosely
punctured, the rest closely strongly reticulated, the reticulations
converging into striz at the apex. Propleure, except at the base,
strongly, closely striated. Mesopleurz closely striated (more closely
and less strongly than the propleurz) obliquely above at the base,
running from the tubercles to the base of the pleure ; the striz on
the lower half are not so oblique and run from the base to the apex.
Base of the metapleurz with some irregular widely separated keels,
the rest closely, strongly, obliquely striated. Petiole obscurely acicu-
lated ; post-petiole smooth and shining ; the second segment thickly
covered with round, clearly separated punctures ; the third similarly
but much less strongly punctured. The area on the base of the
metanotum is wide at the base, not separated from the basal furrow ;
it becomes obliquely narrowed towards the apex.
Although this species has the same host as S. cyclosie it is not,
I believe; its male. The two may be separated thus :
Post-petiole strongly aciculated and punctured ; the mark on the
centre of metanotum broader than long; one long mark cn
the mesopleure, cyclosic,
R : 8(17)05
116 SPOLIA ZEYLANICA.
Post-petiole smooth, shining, impunctate, the mark on the centre
of the metanotum longer than broad ; two widely separated
marks on the mesopleure, panthone.
21.—Bathythrix ? rubriornatus, sp. nov,
Plate B, fig. 4.
Black, the upper part of the propleurze and mesopleure, the meso-
notum, and the scutellum dark red ; the lower part of the propleurse
yellowish; the apex of the first abdominal segment at the sides broadly
testaceous ; the apex of the third and fourth yellow, the latter only
narrowly. Four front legs dark fuscous, the hinder blackish, the
coxe behind, trochanters, and the base of tibiz white. Wings
hyaline, a broad cloud, rounded and narrowed in front behind the
transverse median and the transverse basal, and a much broader one
extending from near the base of the stigma to the end of the radial
cellule, the centre of the apex of the wings, being hyaline ; the apex
of the hind wings smoky, Antenne fuscous, darker at the apex, ¥.
Length 6 mm.; terebra 1 mm.
Peradeniya, April.
In Ashmead’s tables this species would run into the genus Bathy-
thrix, but not having a specimen of that genus for comparison I can-
not say if it be really identical with it or not. In my species the
parapsidal furrows are deep, striated, and broadly rounded behind,
not reaching to the scutellar depression. The areolar is 6-angled,
narrowed behind and much longer than wide ; there is a distinct
petiolar area ; the base of the segment is deeply depressed. The
head (including the clypeus) and thorax are thickly covered with
white long pubescence. Mesonotum aciculated. Scutellum laterally
keeled to near the middle. Median segment closely, finely punctured.
Petiole closely, distinctly, ongitudinally striated ; the second and
third segments are closely punctured. First joint of flagellum as
long as the second.
22.—Bathythrix ? striatus, sp. nov.
Plate B, fig. 5.
Black, the antennal scape, the base of prothorax, tegula, tubercles,
the base of the basai four segments of the abdomen broadly and of
the last narrowly, yellow. Legs rufo-testaceous, the apical four cox
and trochanters and the base of the hind tibiz yellow. Wings
hyaline, a black cloud behind the transverse median nervure anda
large one beginning at the base of the radial cellule, extending to near
its apex and more dilated on the outer side below than on the upper ;
there is a fuscous cloud near the apex of the hind wings, ¥,
Length 7 mm.; terebra 1 mm,
Kandy,
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. D7
Head closely rugose ; the vertex closely striated in the centre,
closely covered with white pubescence; the centre of the face
roundly tuberculate. Mesonotum very finely and closely transverse-
ly striated; the furrows crenulated. Metanotum closely rugosely
reticulated, the basal central area is wide at the base, becomes
gradually narrowed to the apex, where the keels almost unite ; the
areola is open at the apex, being continuous with the posterior
median area ; the lateral basal arez are large and of equal width.
Except at the apex the basal three abdominal and the basal half of
the fourth segments are closely, regularly, finely, longitudinally
striated.
This is a larger and stouter species than B. rubriornatus ; the
abdominal segments are closely longitudinally striated, the metano-
tum less regularly areolated, there being no regularly defined areola,
and the face is more distinctly tuberculated in the centre.
a ;
Clitiga, gen. nov.
6. Antenne as long as the body, 26-jointed, the first joint
of the flagellum longer than the second. Parapsidal furrows indicated
at base only. Scutellum laterally stoutly keeled to near the apex.
Median segment short, smooth, and shining, the apex with a straight,
steep slope ; completely areolated, the areola wider than long,
rounded behind, transverse at the apex ; there are five are on the
apex ; spiracles small, oval; the sides at the apex toothed. Meso-
pleure largely, deeply excavated in the centre. Areolet 5-angled,
the apical nervure faint, but distinct ; disco-cubital nervure rounded,
not angled or broken; the transverse median nervure received
beyond the transverse basal; transverse median nervure in hind
wings broken below the middle. First abdominal segment greatly
dilated at the apex, the base stoutly, angularly projecting ; the base
of second segment depressed, the apex of the depression transverse,
clearly separated. Legs moderately stout. Face flat, not separated
from the clypeus, which is transverse at the apex. Mandibles with
a long apical and a short sub-apical tooth.
2? Antenne stout, dilated towards the apex, longer than the body,
the basal joints of flagellum elongated, The post-petiole is not so
markedly tuberculated on the sides; the apex of the abdomen is
bluntly pointed ; the ovipositor projects and has stout sheaths.
The areola isnot wider than long, as it isin the 6 I have described,
it being nearly as long as wide ; the base of the second abdominal
segment is not tuberculate laterally, but it is depressed at the base.
asin the 4, The abdomen is short, bluntly pointed at the apex,
118 SPOLIA ZEYLANICA.
The difference between the Hemitelini and the Phygadeuonini
appears to be somewhat shadowy, and practically consists in the
absence in the former of the second transverse cubital nervure and
in its presence in the latter. It is clear enough in the present
genus, although bullated in the middle. In Ashmead’s arrangement
of the Phygadeuonini it would come in near Leptodermas, having five
arez on the apex of the metanotum, but it has no keels on the first
abdominal segment. The deeply excavated mesopleura is note-
worthy. The pleural furrow is wide, deep, curved.
23.—Clhitiga excavata, sp. nov.
Plate B, fig. 6.
Black, shining ; the face, except for a line in the centre above,
clypeus, and mandibles except at the apex, palpi, the base of the
prothorax, tegule, tubercles, the lower part of the mesopleural
depression, the mark roundly narrowed above and the apex of the
scutellum, pale yellow. Wings hyaline, the nervures and stigma
black. Front legs fuscous—testaceous ; the cox and trochanters
white ; the middle fuscous, the coxe at the apex and above white ;
the hind legs with the coxe black, the rest fuscous, tinged with
black, ¢.
Length 5mm.
Haputale, ex pupze of Michreseus oblatarius.
January.
Mesonotum clearly, but not closely punctured. The central
excavated part of the mesopleure is smooth, its edges punctured ;
the lower part and the sternum rather strongly punctured. The
upper part of the metapleure strongly, but not closely punctured ;
the lower part smooth and bounded by stout, curved keels. The
posterior median area of the metanotum is surrounded on the inner
side by a row of fovez ; the outer area is stoutly closely striated,
the inner with two or three irregular keels, the central quite smooth ;
the outer is whitish above and there is a white spot on the apex of the
spiracular,
24,—Clitaga forticornis, sp. nov.
Plate B, fig. 7.
Black, the face, except for a black line broad and narrowed below,
a line on the centre of the clypeus broadly dilated at the apex, the
orbits (broadly above) except on the centre of the malar space, basal
half of mandibles, palpi, a broad line on the apical half of the prono-
tum, the lower part of propleure, tegule, tubercles, scutellums, the
sides of the median segment from shortly above the spines, about
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 119
the lower third of the mesopleurz, the mark rounded at the base and
apex, a large oblique mark on the apical half of the metapleure,
united to the lateral mark on the metanotum, the apices of the basal
two abdominal segments broadly, a large mark on the sides of the
third segment, its apical half dilated inwardly, a line, narrowed at
the apex on the sides of the fourth and the apical segments, pale
vellow. Legs rufo-fulvous, the four front coxe and trochanters
yellow, the hind coxe black, tinged with brown at the apex, yellow
for the greater part above, trochanters black below, yellow above,
their tibize and tarsi infuscated at the apex. Wings hyaline, the
nervures and stigma black. Antenne longer than the body, the
middle broadly clear white, ¥.
Length 5-6 mm.
Ex pupe of Michreseus oblatarius.
Haputale, January.
Face punctured, but not closely or strongly ; the clypeus smooth,
with some scattered punctures above. Front and vertex smooth.
Mesonotum, lower half of mesopleure and mesosternum closely
punctured. Base and centre of metanotum smooth, the edges of
the areola and the posterior median areze with a narrow striated
border, the other arez aciculated. Metapleurze more strongly
punctured than the mesopleure. Post-petiole closely rugose and
finely striated ; the second and third segments closely punctured.
Gastrocceli moderately deep, smooth, yellowish testaceous.
Harrana, gen. nov.
Median segment smooth, with one transverse keel behind the
middle ; long, gradually rounded towards the apex ; the spiracles
large, about three times longer than wide. Abdominal petiole long,
slender, not much dilated at the apex, the spiracles placed imme-
diately behind the middle ; the segments smooth ; the ovipositor pro-
jecting. Areolet punctiform as in Mesostenus, open in front ;
transverse median nervure received behind the transverse basal ;
the transverse median nervure in hind wings broken in the middle.
Parapsidal furrows deep. Thorax more than three times longer than
wide. Legs long and slender ; the hind coxe three times longer than
wide; the fore tarsi twice the length of tibiz; claws moderately
long, slender. Head slightly wider than the thorax. Metapleural
keels stout, reaching to the hind coxe. Eyes large, reaching to
the base of the clypeus, there being a clear malar space. Mandi-
bles edentate, gradually narrowed towards the apex.
The type of this genus is a peculiar insect. It has the deep parap-
sidal furrows and the furrow on the lower part of the mesopleurze of
120 SPOLIA ZEYLANIGA.
the Cryptine, and its minute areolet would place it in the Mesostenini,
but its uniform rufo-testaceous colour is very different from anything
found in that group. It might in fact be readily mistaken for a
Paniscus or an Ophionid. It differs from the Cryptine in the spiracles
on the first abdominal segment being placed quite close to, almost at
the middle—further back than in Ophion, but not so far back as in
Paniscus. Inthe position in which the spiracles are placed it agrees
with the Malay genus Cesula. From the resemblance of the
species to Paniscus and Ophion I have no doubt that observation
will show that it is, like the genera mentioned, nocturnal in habits.
The ocelli, too, are large as in these genera.
25.—EHarrana lutea, sp. nov.
Plate B, fig. 3.
Luteous, smooth, and shining, the face, inner orbits to the ocelli
and mandibles, except at the apex, lemon-yellow ; the four front
legs at the base yellowish, the hind tibiz and tarsi fuscous ; wings
clear hyaline, the nervures black; antennz longer than the body,
black, the scape luteous ; it is short, thick, not much longer than
oroad. Pleural sutures crenulated; on the median segment,
between the hind cox, are three stout transverse keels, bordered
on the outer side by a keel which runs down to the base of the hind
cox. Mesopleural furrow deep, obscurely crenulated, °.
Length 10 mm.; terebra 2 mm.
Kandy, October.
OPHIONID&.
26.—Ophion bicarinatus, sp. nov.
Dark rufous, the eye orbits broadly, face, scutellum, and pleura
pallid yellow; antennze pale rufous, wings hyaline, the costa
and stigma rufous, the nervures darker; the recurrent nervure
received twice the length of the transverse cubital nervure behind
the base of the apical abcissa of the radius; the stump of a nervure
as long as the transverse cubital. On the apical slope of the meta-
notum, commencing at the transverse keel, are two stout longitudinal
keels ; the sides of the apical slope are bordered by a stout, waved
keel, which extends to the base of the hind coxe. Face minutely
punctured ; clypeal fovee large, deep. Thorax impunctate, meso-
notum covered with a fuscous pubescence ; the parapsidal furrows
distinct on basal half, &.
Length 21 mm.
Maskeliya, November,
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 12]
The depression at the base of the metanotum is deep, roundly
narrowed towards the apex ; its width at the base longer than the
total length. This species is much larger than any of the recorded
British Indian species. It has keels on the metanotum as in O. areo-
latus, Cam., but, apart from the smaller size of the latter (15 mm.),
areolatus has the basal depression on the metanotum semicircular,
not dilated distinctly at the apex as in the Ceylonese species. 0.
fuscomaculatus, Cam., is also smaller and may readily be known
by the recurrent nervure being received opposite the end of the
basal abcissa of the radius. In one specimen there are indications
of three fuscous lines on the mesonotum.
27.—Pleuroneurophion erythrocerus, sp. nov.
Head pale lemon-yellow, the thorax pale testaceous, the mesono-
tum more rufous in tint; the abdomen, except at the base, rufo-
testaceous, darker towards the apex ; legs testaceous, the anterior
paler. Wings hyaline, the stigma testaceous, the nervures blackish ;
the basal abcissa of the disco-cubital nervure slightly roundly curved
downwards at the apex ; there is hardly an indication of a stump
of a nervure on the disco-cubital nervure; its apical abcissa is
slightly but distinctly longer than the basal abcissa of the cubitus,
which is slightly shorter than the recurrent nervure ; the disco-
cubital is thickened in the middle, with a long bulla beyond. Pro-
thorax and mesothorax closely punctured, the propleure obscurely
striated. Basal depression of metanotum with two stout and some
indistinct keels; the base smooth,the rest with broadly rounded striz
Metapleural keel broad, rounded at base and apex, the middle
slightly turned upwards, ¢.
Except that there can hardly be said to be a stump of a nervure
on the disco-cubital nervure this species fits into Plewroneurophion.
It has the thickened base of the radius of that genus and of Eniscos-
pilus, but it wants the horny points found always in the latter genus.
The transverse median nervure is placed behind the transverse
basal; the median nervure in bind wings is broken far below the
middle. The ocelli are not in a black patch. The antenne are
much more reddish in tint than the body.
Length 14-15 mm.
Peradeniya.
28.—Eniscospilus spilonotus, sp. nov.
Rufo-testaceous, the head, except the ocellar region which is
black, yellow ; the mesonotum with three large black lines, the fifth
and following segments of the abdomen black. Antenne black,
fuscous towards the apex. Wings hyaline, the stigma and nervures
122 SPOLIA ZEYLANICA,
black ; there is one dark horny point, broadly transverse below,
roundly narrowed in front, the base rounded, the apex slightly
rounded inwardly. Transverse cubital nervure two-thirds of the
length of the recurrent, which is more than twice its length from it.
Scutellum yellow, its apical half irregularly striated. Base of
metanotum smooth; the base of the apical part smooth, except for a
few irregular striz ; the centre with roundly curved irregular striz ;
the apex smooth. Propleuree almost smooth, the centre of the
mesopleure finely, closely striated except above; metapleure below
the keel finely, irregularly striated ; the spiracular area coarsely,
irregularly, obliquely striated, much more strongly towards the
apex than at the base. Metasternal keel stout, curved at base and
apex, the base dilated.
‘Brulle (Hymen. IV., 145) describes an O. plicatus from Java
which has three black marks on the mesonotum ; but it is an Ophion,
not an EHniscospilus, it having no horny points in the wings.
29.—Eniscospilus melanocar pus, sp. nov.
Rufous ; the fifth and following segments of the abdomen black ;
the antennz black, broadly rufous at the base ; wings hyaline, the
stigma and nervures black ; the horny point triangular, the sides
rounded, large, and with a long curved tail; there is a small point
beyond the middle of the latter, about twice longer than wide,
rounded behind, transverse at the apex. Transverse cubital nervure
one-third of the length of the apical branch of the disco-cubital and
about one-fourth shorter than the recurrent. Scutellar keels stouter
than usual. Basal depression of metanotum stoutly closely striated,
more closely in the centre than on the sides ; the base smooth ; the
rest closely reticulated, less closely at the base, where there are some
irregular longitudinal keels. Pleure closely, but not strongly
punctured; the metapleure more strongly than the rest; the
propleure striated closely in the centre.
Length 12 mm.
Kandy, August.
30.—Eniscospilus xanthocephalus, sp. nov.
Testaceous, the head and scutellum pallid lemon-yellow; the
ocellar region and the: fifth and following abdominal segments
black ; the occiput and a line in the centre of the face rufous.
Antenne rufo-testaceous ; the basal two-thirds of the first abdominal
segment yellow. Wings clear hyaline, the stigma clear testaceous,
the costa and nervures black ; transverse cubital nervure straight.
oblique, slightly more than one-half the length of the recurrent
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 123
nervure, which is about one-fourth shorter than the apical abcissa of
the disco-cubital nervure ; third discoidal cellule short and wide, in
length not much more than twice its width at the apex. Transverse
basal nervure interstitial. The horny spot is broadly rounded
above, slightly, gradually narrowed to the apex which is transverse ;
the apical spot is much smaller, about three times longer than wide,
oblique at the base, narrowed at the apex, .
Length 11-12 mm.
Peradeniya, June.
Smooth and shining; the upper half of the mesopleure finely
punctured, the lower finely, closely, longitudinally striated. Basal
slope of metanotum with a keel in the centre and one on the sides ;
the basal region smooth, the apical almost smooth in the centre, the
sides transversely striated. Apex.of clypeus broadly rounded.
This species is readily known from the other species here described
by the wide, as compared with its length, third discoidal cellule
31.—Eniscospilus unilineatus, sp. NOV
Pallid testaceous, the head pale lemon-yellow ; the third and
following segments of the abdomen fuscous ; a deep, black, broad
line in the centre of the middle lobe of the mesonotum, the meso-
sternum black; the apex of the first and the second abdominal
segments are more rufous in colour. Legs coloured like the thorax,
the femora deeper in tint. Wings hyaline, the nervures and stigma
black ; the basal horny point is pyriform and is followed by a less
distinct curved line ; beyond it is a small square spot, ?.
Length 17 mm.
Peradeniya, July
Antenne rufous, the scape and the basal 10-11 joints of flagellum
blackish. Head shining, impunctate ; the apex of clypeus broadly
rounded. Mesonotum shining, covered with a pale pubescence.
Basal half of scutellum keeled laterally. Base of metanotum
smooth, irregularly transversely striated laterally, the centre irregu-
larly stoutly reticulated ; the apical slope with some stout, irregular,
roundly curved keels. Propleuree smooth, indistinctly obliquely
striated below. Mesopleure closely, finely punctured. Metapleurz
finely rugose, the upper part with some stout, oblique striz.
32.—Eniscospilus dasychire, sp. nov
Pallid luteous, the abdomen darker coloured, especially towards
the apex ; the head pallid yellow ; antennz rufous. Wings hyaline,
the stigma, costa, and basal abcissa of radius testaceous ; there is
one small pyriform horny point; the transverse cubital nervure
S 8(17)05
124 SPOLIA ZEYLANICA.
slightly roundly curved, as long as the recurrent nervure. Scutel-
lum keeled laterally to the apex. Base of metanotum smooth ; the
basal depression with a stout central and two or three keels on
either side of it ; the base, behind the keel, is irregularly striated ;
the striz in the centre longitudinal, the others irregularly trans-
versely, the rest of the segment with close, rather strong, regularly
roundly curved striz. Propleure closely, obliquely striated. Meso-
pleure closely punctured. Metapleure closely, coarsely, obliquely
striated.
The disco-cubital nervure is roundly curved and thickened in the
centre and with a large bulla beyond the thickened part. Legs
densely pilose ; the pile on the femora longer, paler, softer, and
sparser than on the tibiz or tarsi, ¥.
- Length 21 mm.
Pundalu-oya, January.
Bred from Dasychira horsfield.
33.—Eniscospilus horsfieldi, sp. nov.
Length 18 mm., ?.
This species is very similar to the above described, but is smaller,
more slenderly built, and paler in colour. The two may be separated
thus :—
Length 18 mm.; transverse cubital nervure straight, oblique,
not distinctly rounded ; distinctly shorter than the recurrent
nervure, horsfieldi.
Length 21 mm.; transverse cubital nervure with a distinct,
rounded curve, as long as the recurrent nervure, dasychire.
Base of metanotum shagreened ; the base of the apical part has
one or two longitudinal keels ; the rest with irregular, curved keels
pointing towards the apex ; the rest is closely covered with roundly
curved backwards, strie, which, at the apex, extend on to the
pleure. Propleurz rather strongly obliquely striated in the middle;
the mesopleure in the centre above broadly, and below entirely
striated ; the upper part of the metapleurz coarsely, irregularly
reticulated ; the rest closely, somewhat strongly, obliquely striated.
Stigma, costa, and basal abcissa of radius, rufo-testaceous ; there is
only one small, somewhat pyriform horny point. Legs densely
pilose. The middle lobe of the mesonotum is darker than the latter.
Spilophion, gen. nov.
Transverse median nervure broken shortly, but distinctly, below
the middle. Base of radius thickened. Disco-cubital nervure
roundly, broadly curved, without a stump ; originating before the
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 125
discoidal nervure. Transverse median nervure interstitial. Ab-
domen strongly compressed, more than twice as long as the head
and thorax united. Apex of clypeus transverse ; labrum projecting,
roundly narrowed towards the apex, more than half the length of
the clypeus.
This genus has the thickened base of radius of Pleuroneurophion
and Enicospilus ; the former may be known from it by the not
interstitial transverse basal nervure and by the angled, almost broken
disco-cubital nervure ; the latter by the presence of blister spots
and by the transverse median nervure in hind wings being broken
far below the middle. Characteristic are the projecting labrum and
the spotted wings.
34.—Spilophion maculipennis, sp. nov.
Plate B, fig. 13.
Testaceous, largely marked with yellow; three large marks,
narrowly separated, on the mesonotum, apex of metanotum, a large
mark, obliquely narrowed at the base, on the lower half of the
mesopleure, a large ovoid mark on the metapleure ; the third
abdominal segment above and the greater part fifth and sixth black ;
wings hyaline, the base of radial cellule smoky ; the stigma and the
nervures black ; the former pale at the base; face, oral region, and
orbits lemon-yellow ; antennz rufo-testaceous, ¢.
Length 15-17 mm.
Peradeniya, September.
Basal abcissa of radius thickened at the base. Disco-cubital
nervure roundly curved downwards, the apex broadly, roundly
curved upwards; the cellule bare above, but without horny points.
Head smooth and shining, almost entirely yellow. Prothorax and
mesothorax impunctate. Scutellum laterally keeled to near the apex.
Metanotum behind the keel smooth ; the base depressed, with two
stout keels in the middle ; down the centre of the smooth part is a
fine longitudinal keel. The basal part is smooth laterally ; the
centre with three or four irregular keels ; the central stoutly irregularly
reticulated, its sides obliquely stoutly striated, the strize clearly
separated ; the apical slope has some widely separated irregular
longitudinal striz. Pleurze smooth and shining.
35.—Paniscus intermedius, sp. nov.
Fulvous, the thorax slightly tinged with yellow, the face, oral
region, and orbits pale yellow ; antenne rufous, darker towards the
apex, wings hyaline, the stigma fuscous, the nervures blackish.
Scutellum stoutly keeled, the keels uniting at the apex in a sharply
126 SPOLIA ZEYLANICA.
pointed angle. Areolet oblique, almost appendiculated above ;
the second nervure largely bullated ; the recurrent nervure above
and shortly below the middle widely bullated. Disco-cubital
nervure roundly, broadly curved, not broken by a stump of a nervure.
Transverse median nervure in hind wings broken near the bottom of
the upper fourth. Median segment smooth at the base, the rest
closely transversely striated, the striz running into reticulations at
the apex. Pronotum and mesonotum closely, minutely punctured ;
parapsidal furrows distinctly defined, especially in the middle. Legs
coloured like the body, the anterior paler than the posterior ; tarsal
joints spinose at the apex and more sparsely on the rest, ¢ and 6.
Length 10-11 mm.
Maskeliya; Pundalu-oya.
Owing to their uniform colouration the species of Paniscus are
difficult to define. The present species appears to be sufficiently
distinguished from the known Oriental forms by (1) its smaller size; (2)
by the transverse median nervure in hind wings being broken higher
up, nearer the upper fourth than the upper third as with the others ;
(3) by the keels on the scutellum uniting in a sharp angle, while in the
others they do not unite, but are widely separated. As with the
other species the ocellar region is black. The apex of the clypeus is
more transverse, less rounded than it is in the other species. The
second joint of the hind tarsi is distinctly shorter than the third and
fourth united. The temples are little developed and obliquely,
sharply, roundly narrowed.
36.—Pamniscus orientalis, sp. nov
Rufous, the orbits narrowly pale yellow ; stigma testaceous, the
nervures black. Scutellum more strongly punctured than the mego-
notum, whose furrows are distinct on its basal two-thirds only; the
scutellar keels are stout and do not unite at the apex. Metanotum
smooth at the base only, the rest closely, strongly, transversely
striated. Pleurz closely, distinctly punctured ; the lower part of
the propleurz and metapleur closely, distinctly, obliquely striated.
Transverse median nervure in hind wings broken shortly above the
middle ; disco-cubital nervure not broken by a stump of a nervure.
Ocellar region black. Face strongly and closely, the clypeus sparse-
ly punctured, the apex of the latter bluntly rounded. Wings hya-
line, the stigma testaceous, the nervures black. Tarsal spines long,?.
Length 13 mm.
The pleure are much more strongly punctured than the mesono-
tum, which is almost smooth.
This is a larger species than tntermedius, but still smaller than the
other Indian species ; from intermedius it may be known by its
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 127
deeper rufous colour; by the transverse median nervure in hind
wings being broken lower down, by the scutellar keels not converging
at the apex, by the temples being more largely developed and not so
obliquely narrowed, and by the metapleural keel being much more
broadly developed, especially at the base. Antenne dark rufous,
darker towards the apex.
Pundalu-oya, October.
Paniscus ceylonicus, Cam., is a much larger (nearly 25 mm.) and
stouter species than the two I have described here ; the stigma is
dark fuscous ; the disco-cubital nervure is broken distinctly by a
stump of a nervure; the recurrent nervure forms two rounded curves,
the upper being more broadly rounded, while in those here described
it forms one curve only.
37.—Pamniscus levis, sp. nov.
Pallid yellow, entirely smooth and shining ; the antenne with a
rufous tinge, not darkened towards the apex, wings hyaline, the
stigma pallid yellow, the nervures blackish, ?.
Length 8 mm.
Kandy, July.
This species differs from the other Oriental species in being perfect-
ly smooth, the head and thorax being impunctate and the metano-
tum not striated. It differs also in the transverse median nervure
being received at a greater distance from the transverse basal, and
the eyes below are separated by the same distance as they are at the
top, while in the others, e.g., orientalis and intermedius, the distance
is clearly greater below than it is above. The parapsidal furrows,
too, are much less strongly indicated.
The scutellar keels do not unite at the apex. Apex of clypeus
rounded ; above it is not separated from the face. There is no
malar space, the eyes touching the base of the mandibles. Temples
distinct, obliquely narrowed. Transverse median nervure in hind
wings broken close to the bottom of the upper fourth, the lower part
with a more sharply oblique slope than usual. The antennz are
much more densely pilose than usual, the pile being close, long, and
white. The metasternal keel is wider at the base than at the apex.
It is a more slenderly built species than the others.
38.—Campoplex greent, sp. nov.
Black, the third, fourth, and fifth abdominal segments red ; the
apex of the fore femora, tibize, and tarsi testaceous; the nnddle
femora near the apex and the tibia more broadly at the base,
128 SPOLIA ZEYLANICA.
obscure testaceous ; calcaria dark testaceous. Wings hyaline, the
nervures and stigma black, °.
Length 10 mm.
Hatton ; Pundalu-oya, February.
Head and thorax thickly covered with silvery pubescence. Head
closely punctured. Mandibles uniformly dark rufous including the
teeth, their base thickly covered with white pubescence. Palpi dark
testaceous. Thorax closely, distinctly punctured, the pleurz less
closely than the mesonotum ; the middle and lower part of the pro-
pleure with curved, oblique, clearly separated striz; the apical
central depression on the mesopleurz closely, regularly striated.
On the base of the metanotum are two roundly curved keels, forming
two aree, which are as long as wide ; the centre is depressed and
transversely striated closely above, more widely and broadly below.
Areolet small, oblique, shortly appendiculated.
39.—Limnervum taprobanicum, sp. nov.
Black ; the mandibles and palpi pale testaceous ; the four front
legs dark rufous, the coxe black at the base, trochanters pale yellow ;
the hinder darker red, their coxe and basal joint of trochanters, apex
of tibize, and the tarsi black ; wings hyaline, the nervures fuscous
black ; the stigma paler below, ?.
Length 6 mm.
Maskeliya.
Head and thorax opaque, covered with white pubescence finely
closely punctured. In the centre of the base of the metanotum is a
clearly defined area about three times longer than wide and of equal
width throughout ; from the apex of this two keels run ; they are
at the base roundly curved outwardly, then run obliquely to the
outer side of the apical slope, uniting with the pleural keel above the
middle of the coxe. Propleurz closely, and rather strongly oblique-
ly striated. Areolet shortly appendiculated, 4-angled, its width
at the angles as long as its length ; the recurrent nervure is received
in the middle.
40.—Nototrachys reticulatus, sp. nov.
Black, the four anterior legs and the antennal scape testaceous
tinged with fuscous, wings hyaline, the apex slightly fuscous, the
nervures and stigma black, ?.
Length 7-9 mm.
Peradeniya, January.
Antenne 20-jointed, the last joint as long as the preceding
two united, densely covered with a microscopic pile ; testaceous
towards the apex; the first joint of flagellum distinctly longer
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 129
than the second. Front and vertex shining, the former transversely
finely striated, keeled down the middle. Face and clypeus closely
punctured. Mandibles rufous. Mesonotum closely, irregularly reti-
culated, the sides closely strongly punctured. Scutellum more widely
and irregularly reticulated, its sides stoutly keeled. Metanotum
reticulated, more widely and distinctly on the basal than on the
apical slope ; the base bordered by a keel and with a small area,
longer than broad and of equal width in the centre. Propleuree
smooth, the apex with a stoutly striated border. Mesopleure finely
rugose and irregularly obliquely striated. Metapleuree rugose and
irregularly reticulated. Abdomen smooth and shining; the basal
two segments as long as the head and thorax united; the first
shorter than the second and dilated at the apex.
Transverse basal nervure interstitial ; disco-cubital roundly curved ;
third discoidal cellule at base more than half the width at the apex ;
transverse median nervure in hind wings unbroken. Hind tarsi
longer than tibiz, claws minute, the calcaria of moderate length ;
tarsi minutely spinose, slender. Antenne, slender, as long
as the head, thorax, and first segment of the abdomen. Scutellar
depression large, wide, deep.
Thave, unfortunately, not an example of Nototrachys for com-
parison, but the species I have described agrees fairly well with the
descriptions of that genus.
Clatha, gen. nov.
Hind wings with only two cellules, a large anterior, reaching to
shortly beyond the middle, and with the apex broadly rounded, and
a posterior, not reaching to the middle of the anterior and with its
apex straight and slightly oblique. Disco-cubital nervure intersti-
tial with the discoidal, rounded at the base, the third discoidal cellule
narrowed and pointed at the base ; second discoidal cellule one-third
wider at apex than at base. Hind tibiz as long as the trochanters
and femora united, curved and narrowed at the base; the basal
joint of the trochanters about four times longer than the apical.
Mandibular teeth large, unequal. Hind metatarsus as long as the
other joints united.
The transverse median nervure interstitial. Claws small, simple ;
apex of clypeus rounded. Eyes bare, converging below. Occiput
transverse, margined.
This genus comes close to Agrypon, Foer., as defined by Ashmead
(Bull. U. 8. Nat. Mus., XXTIT., 89), which apparently is identical
with Atrometus, Schmied. (Zeits. fiir Hymen. u. Dipt. 1902, 361) non.
Ashm. The total absence of apical nervures in the hind wings should
130 SPOLIA ZEYLANICA.
readily separate the present genus. In them the anterior nervure
forms one piece, it being in one continuous rounded curve; the
second nervure is joined to it by.a transverse nervure. The hind legs
are very long.
41.—Clatha longipes, sp. nov.
Rufo-testaceous, a line of equal width, enclosing the ocelli, across
the vertex, a line on the centre of the mesonotum extending from the
base to near the apex, a shorter, broader line on the sides, more
irregular in form, not commencing at the base, but reaching to the
apex, the scutellar depression, the base and apex of the scutellum,
the base of the metanotum, a line down its centre and one of similar
size on the sides, a band on the centre of the propleurz, a smaller one
on the mesopleurz, and one on the base of the metapleure, black.
Four front legs yellowish testaceous, the hinder black, their coxe
testaceous, with a black line on the outer side. Antenne filiform,
longer than the body, black, the base testaceous below. Wings
hyaline, the nervures and stigma black, 6 (7?)
Length to apex of second abdominal segment 9 mm. (the other seg-
ments missing).
Kandy, July.
Head smooth and shining ; the lower part of the face, clypeus,
mandibles, the parts behind and before the ocelli, yellow. Meso-
notum and scutellum rugosely punctured ; the metanotum closely
reticulated. Pleurz coarsely, rugosely reticulated, the centre of the
propleur and mesopleure striated. Legs densely shortly pilose ;
the hind femora streaked with testaceous below. The basal two
segments of the abdomen are as long as the thorax ; are thin, of
equal length, and the first is dilated at the apex.
42.—Trichomma nigricans, sp. nov.
Black ; face, clypeus, mandibles, orbits all round, a broad band
narrowed at the base on the pronotum, tegule, scutellum, four front
cox and trochanters, the fore femora and the four front femora
and tibiz anteriorly, the calcaria and the hind coxe broadly at the
base above, yellow. Wings hyaline, the nervures and stigma
black, °.
Length 16 mm.
Kandy, August.
Head covered with white pubescence, smooth ; the front with a
striated band which gets narrower below. Mesonotum opaque,
the furrows and the part at their apex irregularly reticulated ;
the apex in the middle finely, closely, transversely striated. Scutel-
lum smooth. Metanotum coarsely reticulated, as are also the pleure.
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 131
The apex of the propleure from above the middle stoutly striated ;
the mesopleurze more finely, closely striated. Thorax except the
mesonotum thickly covered with silvery pubescence.
Eyes large, pilose, converging below, incised on inner side, reaching
close to the base of the mandibles. Transverse median nervure in
hind wings unbroken. Second recurrent nervure received beyond
the transverse cubitus; the transverse median about the same
distance beyond the transverse basal. Mesonotal furrows distinct.
Second discoidal cellule not much wider at apex than at base ;
the third half the width at the base it is at the apex. Scutellum
laterally keeled. Mesopleure strongly, closely, longitudinally
striated. The second discoidal cellule not much wider at the base
than at the apex ; the third half the width at the base it is at the
apex. Ovipositor half the length of the abdomen. Occiput margin-
ed, not quite transverse. Temples not narrowed. Mandibular
teeth of equal length. Basal joint of hind trochanters more than
twice the length of the apical; the hind femora narrowed at the
base.
TRYPHONIN &.
43.—Bassus orientalis, sp. nov.
Black ; the face, clypeus, mandibles, palpi, the eye orbits to near
the ocelli, a large mark on the sides of the mesonotum reaching to the
tegule, broad in front, becoming gradually narrowed towards the
apex, the upper hinder edge of the propleurz touching the tegule,
tubercles, tegulee, scutellum, post-scutellum, and a small triangular
mark below the hind wings, lemon-yellow. The apices of the basal
three segments of the abdomen narrowly lined with pale yellow ;
the apical two lines widely interrupted in the middle. Four front
legs yellow, their femora tinged with fulvous ; the hind coxe yellow,
tinged with fulvous ; the femora fulvous ; the hind tibiz yellow to
shortly beyond the middle, the rest of them and the tarsi black.
Scape of antennz yellow, the flagellum brownish beneath. Wings
hyaline, the stigma and nervures black, 6...
Length 6 mm.
Peradeniya.—Bred from Syrphid larvz feeding on the tea aphis.
April.
The lateral sutures bounding the clypeus straight, oblique, the top
rounded, the apical half narrowed, the sides roundly curved, the
apex slightly, but distinctly roundly incised ; the face and oral region
covered with short white pubescence. Vertex punctured but not
strongly or closely ; the front less strongly punctured and distinctly
furrowed down the centre. Thorax closely punctured ; the metano-
tum is more closely, more rugosely punctured ; the central apical
L 8(17)05
132 SPOLIA ZEYLANICA,
part is bounded by a rounded keel and is closely distinctly reticulated.
The basal three segments of the abdomen are closely, distinctly,
rugosely punctured, as is also the basal half of the fourth and less
strongly the basal third of the fifth ; the apical half of the fourth is
sparsely punctured ; in both segments the punctured basal part is
bounded by a transverse furrow, there being also a similar furrow on
the third and a less distinct one on the second. The raised central
part of the first segment of the abdomen is depressed in the centre.
The metapleure below the keel are smooth, above it rugosely
punctured.
Allied to the European B. multicolor. The now cosmopolitan, B.
letatorius, Gr., probably is found in Ceylon. I have seen it from
India, and lately from South Africa.
44.—Rhorus spinipes, sp. nov.
Black ; face, apex of clypeus, mandibles, malar space, inner orbits,
a mark on either side of the mesonotum, the mark triangularly
dilated on the inner side at the apex, scutellum, a broad band on
the apex of the metanotum, a roundish mark on the upper side of the
base of mesopleure, a larger quadrangular one on the apex below,
the usual mark under the hind wings, the apex of metapleure, the
mark united to that on the metanotum and the apices of the abdominal
segments, yellow. Four front legs yellow, the hind coxe black,
yellow above, the trochanters for the greater part black, femora
rufous, tibie dark red, blackish at the apex, tarsi black, Wings
hyaline, the stigma fuscous, °.
Length 7 mm. ; terebra 2 mm.
Kandy.
Antenne longer than the body, the scape marked with yellow,
the flagellum brownish below. Smooth and shining ; the metano-
tum obscurely, fincly, transversely striated, the basal half furrowed
down the middle. Abdomen smooth, shining, densely, shortly pilose.
Areolet oblique, triangular, shortly appendiculated. Four hind
tibize and tarsi thickly spinose.
This is probably not a typical Rhorus, but there is no other genus
into which Ican make it enter. I do not know the type of the genus.
In my species the transverse median nervure in hind wings is broken
near the bottom, the transverse median nervure is interstitial, and
the recurrent nervure is interstitial with the second transverse cubital.
PIMPLIN 4%.
45.—E pirhyssa ornatipes, sp: nov.
Pale fulvous yellow ; the middle of front, the ocellar region, the
mark united to the eyes by a black band of equal width throughout,
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 133
the occiput in the centre, its black mark united to the black on the
vertex by a short line ; mesonotum except for a large mark in the
centre, longer than broad, of equal width and transverse at the base
and apex, post scutellum, a narrow line on the base of metanotum, a
large, somewhat triangular mark on the apical half, a line on its
apex, a conical mark on the lower side of the propleure, the lower
side and apex of metapleurz and bands on the base and apex of the
back of the abdominal segments, deep shining black. Wings
hyaline, their apex smoky, the nervures and stigma black. Legs
coloured like the body, the base of the four hinder trochanters, a
broad band on all the femora behind in the centre above, a shorter
line on the hinder pairin front; the base and apex of the hind tibize
broadly and the four hinder tarsi, black, 6.
Length 10 mm.
Hantane, March.
Antenne black. Mandibles and the parts at their base black.
Head and thorax shining, smooth, except for the usual striation on
the mesonotum and the scutellum which is finely transversely
striated. Head and pleure covered with a short, white pubescence.
The black basal and apical bands on the penultimate abdominal
segments are united with a black line down the centre. The apical
joint of the fore tarsi is black.
46.—Theronia maskeliye, sp. nov.
Pallid yellow, the ocellar region, the mesonotum except for two
lines on the basal three-fourths in the centre and the outer side, a
mark in the centre of the scutellum, dilated roundly at the apex,
the base of the metanotum, narrowly in the centre, broadly at the
sides, a large mark roundly narrowed at the apex, incised at the base,
and broad bands on the base of the other segments, black. Antenne
stout, as long as the body, black, the scape yellow below, the base of
flagellum fuscous. The femora broadly marked with black above,
the middle tibize infuscated behind ; the base of the hind tibiz and
a broad band below it, fuscous black ; the four hinder tarsi blackish.
Wings clear hyaline, the nervures and stigma black, ¥ and 6.
Length 10 mm.; terebra 3 mm.
Kandy, August ; Maskeliya, April.
Smooth and shining, covered with a short, white down. There is
no central area on the base of the metanotum, but two lateral ones,
longer than wide, narrowed towards the apex; the outer keel is
straight, oblique, the inner rounded ; from its outer side a keel
runs along the sides of the segment to its apex, there being thus a
134 SPOLIA ZEYLANICA.
large central area extending from the base to the apex of the metano-
tum, the base being narrowed owing to the basal aree. There is a
curved black line on the occiput.
A variable species as regards the size of the black markings. The
scutellum may have no black mark ; the size of the black lines on the
legs varies, that on the hind femora may be cleft, wholly or in part,
the pleurze may be marked with black, and, as is the case with many
Pimplides, there is a considerable variation in size.
Erythrotheronia, gen. nov.
Median segment areolated, not toothed ; the spiracles longer
than wide. Scutellum raised, stoutly keeled laterally to near the
apex. Abdomen smooth, without furrows. Eyes incised, but not
deeply on the inner side; malar space almost absent. Last joint of
antenne shorter than the preceding two united. Areolet oblique,
5-angled ; the transverse median nervure interstitial. Transverse
median nervure in hind wings broken distinctly above the middle.
Temples moderately large, obliquely narrowed. Legs stout; the
claws large, edentate.
The type of this genus looks like a red Xanthopimpla or Theronia.
In Dr. Ashmead’s system it comes near to the New Zealand
genus Allotheronia, from which it differs in the regularly areolated,
not dentate median segment. Theronia and Neotheronia differ,
inter alia, in having pectinated claws.
47.—Erythrotheronia flavolineata, sp. nov.
Plate B, fig. 12.
Ferruginous, smooth, and shining ; the face, clypeus, mandibles,
palpi, lower outer orbits, malar space, lower half of propleurz, two
lines on the mesonotum, inner side of tegule, the edges of mesonotum
at the base, scutellar keels, apex of scutellum, post-scutellum, apical
slope of metanotum, pleurz except above and the apices of the
abdominal segments, yellow. Legs ferruginous, the four anterior
coxe and trochanters yellow, as are also the hinder trochanters.
Wings hyaline, with a slight fuscous-violaceous tinge and highly
iridescent ; the stigma testaceous, the nervures black, 2 and 6.
Length 14 mm.; terebra 3 mm.
Kandy, July.
Antenne black, the scape yellow, the flagellum brownish beneath.
Face sparsely, but distinctly punctured, the sides raised. Thorax
and abdomen smooth, shining, impunctate ; the mesonotum thickly
covered with fuscous pubescence. On the base of the metanotum
are five large arev, a large central, twice longer than wide, with two
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 135
wider than long on either side of it, the basal being the larger, the
keel bordering it being broadly rounded at the apex; the apical
are widest on the outer side; there is only one larger area on the
apical slope, on the sides is a large spiracular area, followed by a
much smaller, somewhat triangular one at the apex.
48.—Echthromor pha ornatipes, Cam.
Plate B, fig. 8.
This species (described from the Khasia Hills, Assam) has been
taken at Peradeniya by Mr. Green.
49.—Xanthopimpla taprobanica, sp. nov.
Pale yellow, a triangular mark enclosing the ocelli, a broad band,
irregularly trilobate—one on each lobe—across the base of the meso-
notum at the tegule, a mark on the tegule continuous with it, a
large transverse mark, roundly curved at the base and extending
on to the scutellar depression, a broad band, narrowed in the centre,
on the base of the metanotum, two large oblique marks on the centre
of the first abdominal segment, dilated in the centre at the base, a
small oblique mark, obliquely narrowed at the base and apex on the
second, alarge broad band on the third, irregularly narrowed laterally
and slightly incised in the middle at the base and apex, two large
broad bands, rounded on the outer side on the fifth, a broad band on
the basal half of the penultimate and two marks on the last, black.
Legs coloured like the body; the middle knees, base of middle
tarsi, the greater part of hinder trochanters, a broad band on the
lower part of the hind femora, apex of femora narrowly above, base
of tibiz, and the tarsi (the middle joints paler in colour), black.
Wings hyaline, their apex with a fuscous cloud, °.
Length 10 mm.; terebra 2 mm
Kandy.
Face rugosely punctured ; the upper part of the clypeus sparsely
punctured. Mesonotum smooth, impunctate ; parapsidal furrows
reaching shortly beyond the middle. Scutellum roundly convex,
the keels not very stout. Areola large, open at the base, transverse
at the apex, longer than broad, the sides angled shortly behind the
middle ; the basal lateral area wider on the outer side, the: sides
roundly curved ; the second large, broad, oblique, of equal width
throughout ; the apical lateral area triangular; the apex on the
outer side, the upper angle longer than the lower. First segment
of abdomen in length clearly longer than its width at the apex,
smooth, except for some punctures in the middle, the second to
136 SPOLIA ZEYLANICA.
fourth segments closely and strongly punctured, the furrows crenu-
lated. Areolet small, oblique, triangular, shortly appendiculated.
The occiput is immaculate, except that the ocellar black spot
slightly projects into it- Pleuree impunctate. Ovipositor as long
as the hind tarsi.
In Professor Krieger’s work on Xanthopimpla this species would
come in near X. splendens. Cf. Bericht d. Naturf. Ges. zu
Leipzig, 1898, 69.
50.—Xanthopimpla kandyensis, sp. nov.
Yellow ; the ocellar region, the mark longer than broad, three
almost continuous marks on the mesonotum at the base, the central
placed more in advance than the others, longer than broad, transverse
at the base, rounded behind, the lateral longer compared with the
width, more irregular, slightly incised in the middle at the base,
rounded and narrowed at the apex ; two irregular, slightly oblique
marks, narrowed on the inner side, on the first segment ; two large
marks, broader than long, with the sides rounded, on the third ;
two somewhat similar ones on the fifth, but broader, more slender
compared with the length, and two similar but larger (the largest of
all) on the seventh and two marks on the base of the metanotum,
broader than long, transverse behind, rounded at the apex, black.
Antenne black, the flagellum brownish, the scape yellow below.
Wings clear hyaline, the nervures and stigma black, .
Length 11 mm. ; terebra 4 mm.
Kandy, July.
Face and clypeus closely, distinctly, but not very strongly punc-
tured. Thorax smooth and shining. The areola on metanotum
broader than long, slightly obliquely narrowed from the apex to the
base ; the lateral arez are of almost the same width, the outer side
obliquely narrowed ; following them is an area which becomes
obliquely narrowed from the base on the inner to the apex on the
outer. Basal two segments of abdomen smooth, the others closely
but not strongly punctured ; the furrows closely striated.
The occiput has no black.
Allied to X. punctata, F.
51.—Xanthopimpla parva, sp. nov.
Pallid yellow, a triangular mark enclosing the ocelli, an irregular
transverse line on the base of mesonotum, with ragged edges, a mark
at the base of the scutellum, two marks, narrowed from the outer to
the inner side, on the base of the metanotum, and marks, broader
than long, on the first, third, and following segments, black. Legs
coloured like the body, two small marks on the hind trochanters, a
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 137
mark on the base of the hind tibiz, and a smaller one on the base of
the hind tarsi, black. Wings clear hyaline, the nervures blackish, é.
Length 6 mm.
Peradeniya, August.
Face closely punctured. Thorax impunctate; the parapsidal
furrows indicated only at the base. There is no central area on the
base of the metanotum, but two semicircular large lateral ones
enclosing the black spots ; there is none on the apex. The basal
two abdominal segments smooth ; the furrows striated ; the other
segments are closely punctured. The apex of the wings slightly
infuscated. Base of mesonotum rounded.
This and the following species are very much smaller than any of
those described hitherto.
52.—Xanthopimpla minuta, sp. nov.
Yellow; a triangular mark enclosing the ocelli, the scutellar de-
pression, two marks on the first abdominal segment, a transverse line
on the third, two closely continuous ones on the fourth, two more
widely separated ones on the fifth, two still more widely separated
marks on the sixth, and a line on the seventh, black. The base of the
hind tibiz marked with black. Wings hyaline, the nervures and
stigma black, &.
Length 5 mm.
°
Kandy, July.
Face minutely punctured. Base of mesonotum transverse, the
parapsidal furrows distinct, the middle lobe clearly separated.
Smooth and shining. There is no area on the base of metanotum :
there is a clearly defined lateral area, broader than long, broad on the
outer, becoming gradually narrowed to a fine point on the inner side;
the sides of the metanotum are keeled. Abdomen closely punctured,
with distinct, striated furrows. The apical half of the first segment
is raised, clearly separated, smooth, of equal width and twice longer
than wide. Antennal scape for the greater part yellow, the flagel-
lum brownish beneath. The antennz are clearly longer than the
body. The ovipositor short.
This species can easily be distinguished from X. parva by the
distinct parapsidal furrows and by the mesonotum being transverse,
not rounded at the base.
Philopsyche, gen. nov.
Kyes very large, reaching to the base of the mandibles, there being
no malar space ; on the inner side they are distinctly incised. Face
keeled down the middle, clearly separated from the clypeus by a
138 SPOLIA ZEYLANICA.
furrow ; the extreme apex of clypeus depressed, not quite transverse.
Last joint of antenne as long as the preceding two united. Scutel-
lum roundly convex. ‘Transverse median nervure in hind wings
broken far below the middle. Areolet triangular, small, appendi-
culated. Transverse median nervure in fore wings placed immediately
behind the transverse basal, almost interstitial. Abdomen and
legs as in Pimpla. Metathoracic spiracles round. Claws slender,
curved, untoothed. The sharp lateral ridges of the mesonotum do
not extend on to the scutellum. The eyes do not converge anteriorly
The last joint of hind tarsi about three times longer than the pre-
ceding.
If the apex of the clypeus is to be considered ‘‘ impressed anteriorly
at apex ” this genus, in Dr. Ashmead’s arrangement, would come in
near [toplectis and Hremochila, with neither of which can it be con-
founded ; if ‘‘ not impressed’’ then it would come nearest to Tromato-
bia, which is easily known from it by the entire eyes. The meta-
pleural keel is distinct, complete. The disco-cubital nervure is
angled beyond the middle, but not broken by the stump of a nervure.
The genus should be readily known by the large, clearly incised eyes,
reaching close to the base of the mandibles, round metathoracic
spiracles, angled disco-cubital nervure, transverse median nervure
in hind wings broken near the bottom, and smooth, white-banded
apex of abdominal segments
53.—Philopsyche albobalteata, sp. nov.
Plate B, fig. 10.
Black ; the antennal scape except above, palpi, hind edge of
pronotum, tegulz, and the apices of the basal six abdominal seg-
ments, white. Four front legs white, the hind coxe and femora
red ; the basal joint of trochanters red, the apical white ; the hind
tibiz at extreme base, and broadly in the middle, white, the rest
black ; the hind tarsi black, the basal joint white to near the apex, 6.
Length 7-8 mm
Bred from Psyche albipes and P. subteralbata.
Matale ; Peradeniya, August.
Head smooth and shining, the face covered with white, the clypeus
with longer white hair ; the former obscurely shagreened. Mesono-
tum and scutellum closely punctured, thickly covered with white
pubescence. Metanotum smooth, shining, and bare at the base and
apex, the middle punctured and thickly covered with long white
hair. Abdomen closely and strongly punctured except on the white
apices of the segments and thickly covered with black pubescence.
Pleurzsmooth, almost bare, except on the metapleure above the keel.
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 139
Lissotheronia, gen. nov.
Abdominal segments broader than long, perfectly smooth, shin-
ing, impunctate; the first segment bituberculate at the base.
Median segment strongly, closely, transversely striated, the spiracles
about three times longer than wide, broader below than above.
Hyes large, incised on the inner side ; malar space small. Clypeus
clearly separated, its apex obliquely depressed. Transverse median
nervure in hind wings broken distinctly above the middle. Trans-
verse median nervure received beyond the transverse basal. Disco-
cubital nervure broadly rounded, unbroken. Claws simple ; tarsi
thickly spinose. Antennz slender, longer than the body, the last
joint nearly as long as the preceding two united.
This genus might be mistaken for a Pimpla, from which, however,
it can be readily known by the perfectly smooth, shining abdomen.
In Dr. Ashmead’s system it would come in near Allotheronia, which
may be known from it by the metathorax having a strong transverse
apical area, with the upper angles dentate or tuberculate, the whole
thorax, too, being “ closely, finely, rugosely punctate ” except for a
spot on the middle of the mesopleure. It has the metanotum closely
transversely striated as in Lissopimpla, having also the smooth
abdomen of that genus ; but otherwise is readily separated from it
by the absence of parapsidal furrows and of a tooth on the hind
femora.
54.—Lissotheronia flavipes, sp. nov.
Black, shining, except the median segment which is closely striated,
the pleure more closely and obliquely than the dorsal surface ; the
legs yellow, except the fore coxe in the middle below, and the
four hind coxe and trochanters, which are black ; wings hyaline,
slightly suffused with yellow. A spot on the scutellum in the centre,
a curved one on the post-scutellum, and one on the base of the
tegule are yellow. The depressed apex of the clypeus is smooth and
shining ; the raised upper part sparsely punctured. Face closely
punctured ; the sides of the lower part roundly convex. Tarsi
spinose, much longer than the tibie. Sternum and under side of
cox thickly covered with fulvous pubescence, .
Length 15 mm. ; terebra 4 mm.
Pundalu-oya, February.
55.—Charitopimpla annulipes, sp. nov.
Plate Le pifea UP
Black; antennal scape except above, tegule, hind edge of prono-
tum, and palpi, yellow; the apex of the abdominal segments narrowly
yellow ; the yellow lines dilated at the sides and tinged with rufous
U 8(17)05
140 SPOLIA ZEYLANICA.
there. Four anterior legs yellow, tinged slightly with rufous ; the
hind coxe and femora rufous, their trochanters yellow ; hind tibiz
black, their base narrowly and the middle broadly white ; hind tarsi
black, the metatarsus to near the apex black. Wings hyaline, the
nervures and stigma black, the areolet small, triangular, with a
long pedicle, it being as long as the branch of the first transverse
cubital nervure, &.
Length 12 mm.; terebra 5 mm.
Pundalu-oya.
Face sparsely pilose and punctured, the middle keeled. Apex
of clypeus brown round the central incision. Mandibular teeth
piceous. Front and vertex smooth and shining. Prothorax smooth,
except along the upper edges. Mesothorax and metathorax closely
punctured, thickly covered with pale pubescence ; on the metanotum
the pubescence is longer and more fulvous in tint. Except at the
apices of the segments the back of the abdomen is closely and strong-
ly punctured ; the apical segments are only slightly punctured.
The antennz are stout, as long as. the abdomen ; the last joint
is as long as the preceding two joints united. The apices of the
tarsal joints are spinose. The metapleurze below the keel are
smooth. The hind coxz below and the extreme apex of the hind
femora are black.
Charitopimpla was described by me in the Journ. Str. Br. Royal
Asiatic Society, 1902, 48.
56.—Lissopimpla rufipes, sp. nov.
Plate B, fig. 14.
Black ; the upper orbits broadly, the outer narrowly above, broad-
ly below, the malar space, face, clypeus, labrum, mandibles, palpi,
a raised line on the basal half of the pronotum, interrupted by the
parapsidal furrows, a mark on the apex of the middle lobe of the
mesonotum, transverse at the base, gradually roundly narrowed to
the apex, scutellar keels to shortly beyond the middle, the apex of
scutellum, the lateral keels, post-scutellum, the three tubercles on the
apex of metanotum, a line on either side of the base of pronotum, the
lower edge of propleurz, tubercles, a small roundish mark on the
base of mesopleure above the middle, the basal half below, a
large mark, roundly narrowed at the apex, obliquely truncated at
the base on the apex of the metapleure, the apex above the hind
cox, two narrow lines on the centre of the first abdominal segment
and about the apical third of the others, yellow. Legs rufous, the
four coxee and trochanters yellowish, the apex of the hind coxe
marked with black, as is also the base. Wings clear hyaline, highly
Se
at
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 141
iridescent, the nervures and stigma black or fuscous black. The
hind tibiz are darker coloured, the hind tarsi fuscous, &.
Length 9-12 mm.; terebra 3-4 mm.
Maskeliya, April; Peradeniya, August.
Antenne longer than the body, slender ; the 8-12 joints of flagel-
lum white. Labrum and clypeus closely and rather strongly punc-
tured. Face depressed, stoutly keeled in the centre, the sides of the
depression roundly curved. Prothorax and mesothorax smooth,
shining ; the pleural sutures closely striated. Metanotum, except in
the centre at the base, closely transversely, the upper part of the
metapleure obliquely, striated. Areolet, as in the typical species of
the genus, oblique, the nervures uniting above. Femoral tooth
small, but distinct.
The occurrence of Lissopimpla in Ceylon is of great interest, the
genus hitherto having been regarded as Australian.
‘Tanera, gen. nov.
Abdominal petiole long and slender, narrowed at the base, as long
as the following two segments united; the segments smooth and
shining, longer than wide ; there is a projecting ovipositor. Median
segment uniformly coriaceous, without a transverse keel; there is
a stout metapleural keel. Legs slender ; the claws not pectinated ;
hind coxz three times longer than wide. Areolet with a long pedicle,
the cubital nervures roundly curved; the recurrent nervure is
received at its apex; transverse median nervure _ received
behind the transverse basal. Clypeus roundly convex, separated
from the face ; its apex broadly rounded. Occiput transverse ; the
ocelli placed close to the edge. Temples very short, almost obsolete
behind the eyes. Antennz nearly as long as the body ; the last
joint longer than the penultimate, but not so long as the preceding
two united.
The eyes are large, converging slightly above ; there is a distinct
malar space ; the ocelli are on the outer edge of the vertex. The
keel on the edge of the metasternum projects at the apex into a
distinct tooth. Metathoracic spiracles small, oval. The abdominal
petiole is longer, more slender and narrower at the base than it is in
Lissonota. If it is to be called “sessile,” in Ashmead’s table (U. 8S.
Nat. Mus. X XIII., 49) it would fit in near 7'revoria, with which it
cannot be confounded ; if “ petiolate’ then it would come in near
Atropha.
57.—T'anera annulipes, sp. nov.
Plate B, fig. 11.
Black, smooth, and shining, except the pleursze and metanotum,
which are closely and strongly punctured ; the latter more coarsely
142 SPOLIA ZEYLANICA.
than the former ; the clypeus, mandibles, palpi, a spot on the malar
space, the inner orbits, more broadly above than below, scutellum,
a line on the centre of the pronotum, tegule, tubercles, the apex
of metapleure ; the basal third of the first abdominal segment,
about the basal third of the second and third, the apex of the third
more narrowly, and the apical segments, yellow. Legs rufous, the
four front cox and trochanters tinged with yellow ; the hind tibize
and tarsi fuscous, tinged with rufous, their base white. Wings
hyaline, the apex of the radial and of the third cubital smoky ; the
stigma fuscous, ?.
Length 8 mm.; terebra 5 mm.
Hatton.
58.—Lissonota greent, sp. nov.
Black ; the clypeus broadly at the apex, more broadly in the centre,
mandibles except the teeth, palpi, underside of the antennal scape,
and the lower edge of the propleure, yellow ; first abdominal seg-
ment, basal third of the second, the base of the third narrowly,
and of the fourth still more narrowly, red ; the apices of the second
and third yellow, tinged with rufous. Legs red ; the anterior coxe
and trochanters and the middle coxe at the base, yellow; the hind
tibize and tarsi black. Wings hyaline, the stigma and nervures dark
fuscous, &.
Length 8 mm.; terebra 5 mm. .
Peradeniya, March.
Closely and regularly punctured. First segment of abdomen
closely, distinctly, longitudinally striated, except in the middle at
the apex ; the second and third segments and the base of the fourth
closely and regularly punctured ; the apical smooth and shining; the
last and the apex of the penultimate white. Areolet not appen-
diculated.
¢
SPOLJA ZEYLANICA.
*
Plate A.
West, Newman ad.nat.lith
HYMENOPTERA.
ree ae
ori
er. rs a
ri ir rs
> “i,
a! 4 a
Te _— i,t ®
oc oe ye a
wi sy tse ee
fa tte Bi i 5 ee
a
ve eat ¥ os
%
7 .
SPOLIA ZEYLANICA. Plate BE
West,Newman ad.nat.lith.
HYMENOPTERA.
PHYTOPHAGOUS AND PARASITIC HYMENOPTERA. 143
EXPLANATION OF THE PLATES
illustrating Mr. P. Cameron's Paper on Phytophagous and Parasitic
Hymenoptera.
PLATE A.
PAGE
1.—Xiphydria striatifrons os Pes 70
2.—Rhacoteleia pilosa see ari 73
3.—Spilomegastigmus ruficeps wt a 74
4,—Rhacospathius striolatus ae ae 86
5.—Troporhogas maculipennis ase it 94
6.—Holeobracon fulvus fee es 90
7.—Iphiaulax erythroura ahs wee 85
8.—Troporhogas tricolor “ee a 94
9.-—Paraspinaria pilosa as “ee 88
10.—Deniya pleuralis... es ssav) SOS
11.—Aluina erythropus ae saat ge OZ
12.—Skeatia acutilineata ss eee 0c
13.—Buodias rufipes... ae Sse OG
14.—Fenenias erythropus She woos vehe
PLATE B.
1.—Friona rufipes Ne bas ane er acl)
2.—Melcha erythropus Se caer) ey
3.—EHarrana lutea “Ae eae see — eh
4.—Bathythrix rubriornatus Pe See
).—Bathythrix striatus a oe Bb
6.—Clitiga excavata ... site Ruled Abe
7.—Clitiga forticornis ... Aas ala a
$.—Hehthromorpha ornatipes ses eG)
9.—Charitopimpla annulipes oes PE BSL,
10.—Philopsyche albobalteata es loo
11.—Tanera annulipes ... cas ppl (Fa
12,—Erythrotheronia flavolineata sve 134
13.—Spilophion maculipennis oe sone lea
14.—Lissopimpla rufipes re oe _ 140
144 SPOLIA ZEYLANICA.
NOTES ON SNAKES COLLECTED AT HAKGALA, CEYLON.
By F. WALL, C.M.Z.S., CAPTAIN, I.M.S.
Fyzabad, United Provinces, India.
HROUGH the kindness of Mr. Nock I have received two
collections of snakes at different times from Hakgala,
Ceylon (5,600 feet).
The number of species included is small (6), and all are
common; nevertheless, collections such as these are of distinct
value with reference to distribution. It will be seen also that
some very interesting information has been gleaned from the
numerous specimens of Aspidura trachyprocta and Ancistrodon
hypnale. have omitted to give measurements, as the specimens
were all preserved in spirit, and under these conditions do not
lend themselves to accurate investigation.
Aspidura trachyprocta.
(48 specimens. )
With reference to this species I notice Gunther says (Rept.
Brit. Ind., p. 203): ‘‘ This species is nearly as common in Ceylon
as A. brachyorrhos.” In the locality of Hakgala A. trachy-
procta is evidently not only vastly more numerous than A. brachy-
orrhos, but by far the commonest snake to be met with at all.
No single specimen of A. brachyorrhos was included.
Food.—Several specimens had their stomachs distended witha
soft putty-like mass, the nature of which I could not determine.
Once I thought I could distinguish a slug, and once agrub. On
four occasions without doubt the ingested material consisted of
earth-worms. I never found traces of any hard-cased insects.
Breeding.—The ovarian follicles are normally about } inch in
length. In one specimen I found 8 follicles (4 in each ovary),
about } inch long, and in another 6 (3 in each ovary) about the
same size, and these appeared to be impregnated. The dates of
both are not known. A third specimen obtained on the 10th of
August, 1904, had 14 follicles (6 in one ovary and 8 in the other)
enlarged to ,2, of an inch. The smallest specimens, which I
believe were hatchlings, measured 4} and 4,% inches. The latter
were found in company with an adalt female on the 10th of
SNAKES COLLECTED AT HAKGALA, CEYLON. 145
August. The adult, however, showed nothing unusual upon
anatomical investigation. The navel involved 2 shields in
both cases. In the 6 10 ventrals intervened between the navel
and the anal shield, in the ? 7.
Sexes.—Of the 48 specimens collected, 21 were males and 27
females. Without actually measuring them, I think there is
no doubt that the females were usually larger. Adult males had
all the scales about the anal region keeled. In the median rows
these were tuberculate, and often bi, tri, or pluridentate, and
in the lowest lateral rows markedly spinose, with the points
directed backwards. The spines were hooked, reminding one
forcibly of rose-thorns, and were attached by a long base to the
anterior three-fifths of each scale.
In young males, or at least in some, the keels were obtusely
tuberculate as in females. The tails of the males were longer, and
the subcaudals varied from 21-26, whereas in females these
shields numbered from 1]-16. The ventrals in males were
however 135-148, against 139-151 in females.
Scale peculiarities.—The scales anteriorly (two head-lengths
behind the head) number 15, mid-body 15, posteriorly (two head-
lengths before vent) 15, keels were absent in all these situations.
The vertebral row was the narrowest of the series, and the
ultimate row very slightly largest. The supracaudals were in odd
rows, as is the rule where the subcaudals are entire. The first
subcaudal was as often divided as entire.
Abnormalities. —One specimen had no preocular. The upper
postocular was confluent with the parietal in one. The labials
were 7, of which the 4th only touched the eye on one side in one
specimen ; 7 with the 5th only touching the eye on the left side
in two specimens.
One very aberrant specimen requires special remark, viz., a
male in which the subcaudals numbered only 16. In addition
the scales anteriorly (two head-lengths behind the head were
‘15 or 14, mid-body 18, posteriorly (two head-lengths before the
vent) 13. Investigation showed that the 2nd and 3rd rows above
the ventrals blended on both sides, one rather before the other,
at the site I select to count the scales anteriorly. The large row
occasioned by this coalescence divided and blended two or three
times before the rows were finally established as 13. One sees
the same vacillation frequently exhibited in snakes whose scales
normally reduce in number, as for instance in T'ropidonotus
piscator, where the scales number 19 in the anterior and mid-
body, and 17 posteriorly. In all other respects this specimen was
normal, and so probably does not deserve specific isolation.
146 SPOLIA ZEYLANICA.
Colour.—Uniform or nearly uniform black, or blackish
dorsally, or varying shades of brown, olive-brown, or olive-green
more or less spotted or speckled blackish. The spots vertebrally
and laterally have a tendency to confluence, and may form lines,
Belly pale yellowish, yellow, ochraceous, or ruddy, sparsely or
profusely spotted, speckled, or marbled with black.
Tropidonotus stolatus.
One female killed on the 12th September, 1904, besides
containing a much digested frog was found to be pregnant.
There were 8 eggs (6 in one ovary and 2 in the other). The
ventrals were 139, subcaudals 66, of which the 32nd, 50th, 51st,
59th, and 60th were entire. As in Malabar specimens the labials
were 8, with the 3rd, 4th, and 5th touching the eye. The nasals
were in contact with the Ist and 2nd labials, a very unusual
character in Indian examples of this species. Usually it is only
the Ist labial that is apposed to the nasal.
Zamenis MUCOSUS.
One specimen about one-third grown, with nothing remarkable
to mention.
Naia tripudians.
I received the head of one of this species, which is uncommon
at the same altitude in India.
Ancistrodon hypnale.
Twenty-seven specimens of this species were collected, so that
it must be a very common snake in this locality.
Food.—It evidently subsists mainly upon the skink, Lygosoma
taprobanense, as no fewer than 11 specimens contained one or
more of these lizards, or in some instances the tails only. I also
found in the stomach of 5 examples small soft-shelled eggs about
half an inch long, probably lacertine, but possibly ophidian.
From one male specimen one such was seen protruding from the
anus, which I extracted and examined. It consisted of the egg
envelope only, compressed, and wrinkled longitudinally, but with
no opening that I could discover, so that it was a mystery to
me how the contents had been absorbed unless by a process
of exosmosis. I floated it in water, teased out the wrinkles,
and examined it most closely, then cut it carefully from pole to
pole and re-examined it from the interior, but could find no sus-
picion of a breach in its continuity. Birds’ eggs are occasionally
passed entire “* per anum,” but are more frequently, I believe,
wholly dissolved by the digestive juices. Perhaps the coriaceous
investment of ova such as this offers a greater resistance to the
SNAKES COLLECTED AT HAKGALA, CEYLON. 147
action of the alimentary secretions than the calcareous envelope of
the former. One specimen contained about two inches of the
tail of a lizard such as a Calotes. Two others contained frogs in a
condition too dissolved to identify, and in one example I found a
snake Aspidura (trachyprocta).
Breeding.—Three specimens were pregnant, but unfortunately
the dates of capture were not noted in any instance. The
prospective mothers measured 1 ft. 2 in., 11} in., and 11} in.
Two contained 6, and one 4 immature eggs.
Sexes.—Of the 27 specimens 10 were males, 13 females, and in 4
the sex was doubtful owing to mutilation. Both sexes appear to
grow to about the same length, and the tails also show little, if any,
disparity. The ventrals and subcaudals were as follows :—
Ventrals 6 123-131 (?); subcaudals 29-37
Ventrals 2 121-138 —; subcaudals 27-36
In 7 at least out of 9 males the tails were bent over ventrally
in a hook-like manner, but in no single female was this seen.
The clasper of the male was bifid on each side as in other
members of the family Viperide, and was surmounted with the
usual array of falciform tentacles. The testicles were not nearly
so elongate as is the rule in snakes, but, as in some other vipers
I know, were oval in form, I could discern no difference in
colouration between the sexes, nor were the keels more pro-
nounced in the scales of males.
Scale peculiarities —One or two things deserve notice. The
nasal shield touches only one labial, viz., the first, and this often
is produced upwards behind the nasal to meet the supraloreal, or
when this is not the case one or more minute intercalary scales
occupy this situation. The lowest preocular is often entire,
often broken up into two. The swbhocular may be entire, in
which case it meets the 3rd and 4th labials, or a small portion is
often detached anteriorly. The 2nd labial variably contributes
to the formation of the loreal pit, often it does not do so, a separate
furrowed scale lying above furnishing a floor to the pit. The
scales are anteriorly (2 head lengths behind the head) 17, midbody
17, and posteriorly (2 head lengths in front of the vent) 17.
Abnormalities.—In one example there were 9 /abials on the left
side. The swblinguals touched 2 infralabials only in 3 specimens.
The 5th swhcaudal was entire in one example, and the 8th, 9th,
and 10th in another.
, Vipera russelli.
Two small specimens were procured, showing that in Ceylon as
in India this species sometimes ascends to a considerable elevation,
though really an inhabitant of the plains.
x 8(17)05
148 SPOLIA ZEYLANICA.
NOTES CHIEFLY ON BIRDS SEEN AT THE PEARL FISHERY CAMP,
MARCH AND APRIL, 1905.
By W. E. Warr.
‘TDIRD-LIFE in the dry littoral zone of the north-western part
of the Island possesses many distinctive features, and as the
birds which for three-quarters of the yearare left in undisturbed
possession of the camp site do not appear to be scared away by the
bustle during the fishery, one constantly saw forms or marked
habits unnoticed elsewhere.
Among the flocks of Brahminy Kites and Gray Crows which
gathered over the offal on the beach, one generally saw two or
three Marsh Harriers (Circus eruginosus). The Common and
Blue-tailed Bee-eaters (Merops viridis and phillipensis) chose the
same unsavoury haunts to feast on the swarms of flies. There
was a foetid pool fed by the drain which carried off the refuse
water from Mr. Dixon’s oyster-washing machine. Here they
might be seen not only hawking for insects on the surface, but
dipping their beaks into the water, probably to fish out the
drowned maggots which were carried down in myriads.
On the short walk from my bungalow just above the beach to
the Kachcheri a few hundred yards iniand I invariably met with
the Rufous Rumped Shrike (Lanius caniceps) flitting among the
mustard trees. ‘This bird, which is I believe almost confined to the
Manaar District, was one of the commonestincamp. Towards the
end of the Fishery while out on an evening stroll with Mr. Ferguson
we saw one attack a large rat-snake.
The bird swooped down into some high grass near the road, and
we watched to see what it had caught. Immediately a six-foot
rat-snake came out into the open, followed by the shrike, which
pecked viciously at its head. The snake made straight for a
hollow tree close by, up which it disappeared, and the bird,
catching sight of us, flew away. It had probably been driving off
the intruder from its nest, but the wonder is that a bird no larger
than a thrush cowed such a big snake.
Between the Governor’s and the Government Agent’s bungalows
lay asmall openspace on which stood two or three gnarled trees.
These were frequently occupied by a few Chestnut-headed Bee-
eaters (I. Swinhoti) and a pair of Hoopoes (Upupa ceylonensis).
BIRDS SEEN AT THE PEARL FISHERY CAMP. 149
Close by was asandy bank, in the side of which a pariah belong-
ing to Mr. Dixon had excavated some burrows wherein she
brought up a litter of puppies. I noticed that another pariah had
dug a similar burrow in the bluff above the Government kottu.
They were doubtless made for the sake of coolness and shade,
which could not be obtained otherwise. I wonder whether the
jackal, or any species of wild dog, is ever in the habit of making
such lairs: whether in fact this was a reversion to the instincts
of the fox or an adaptation to environment. No one that I asked
had noticed pariahs elsewhere make burrows like these.
The neighbourhood of the tani set apart for drinking water
was a favourite roosting-place fora large flock of the Lesser White
Egret. A few Whistling Teal occasionally came there, while the
strip of jungle on the north sheltered several pairs of the Gray
Indian Dove (Turlur risorius). A solitary Grey Heron (A.
cinerea) used often to fish in the shallows at the head of the tank.
To the south of this tank on the opposite side of the cart track
leading to the Manaar road lay several insignificant little tanks
irrigating a small tract of fields which during the fishery lay
fallow. In the small trees and bushes which fringed the water
the Black Drongos (D. atra) had their headquarters, while on
two occasions I saw a pair of the pretty little Yellow-fronted
Woodpecker (Picus mahrattensis) round some Euphorbias which
grew near one of the tiny bunds.
South of the camp, the bare tidal flats of the lagoon at the
river mouth were the happy hunting grounds of innumerable
waders, from the Large Stone Plover tothe Little Stint. Several
species of tern flocked here, sometimes accompanied by a colony
of the beautiful Brown-headed Gull, which usually kept to the
sand-spit on the south side of the bar. On the northern bank, in
the face of the sand cliff just under the Governor’s bungalow, a
large Fish Owl (Ketupa ceylonensis) had his burrow, at the
entrance of which he used to sit of an evening blinking and
ruffing his plumage until it was time for him to fly off on his
rounds. A little further up the river I occasionally met with
the Pied Kingfisher (Ceryle rudis) skimming along under the
bank.
Of other noticeable birds, a species of Sky-lark could be heard
twittering inthe air any morning, and a Gray Partridge (Ortyg-
ornis pondiceriana) might be found on the outskirts of the camp.
A large Fish-eagle came several times to the lagoon ; whether it
was H. leucogaster, or P. ichthyewius I never got close enough
to ascertain. I also heard of the occurrence of a Pariah Kite
(Milvus govinda), but did not see it myself.
150 SPOLIA ZEYLANICA.
In conclusion I would express the hope that some one possess-
ing a good knowledge of ornithology could spare a visit to the
camp when a pearl fishery is in progress. My observations were
made at odd intervals during two very busy months ; my know-
ledge of birds is limited, and if it had not been for Mr. Ferguson,
who knows the Natural History of the district well, and in whose
company I saw practically all the species mentioned, I should
have been, in many cases, uncertain of my identification.
The circumstances are particularly favourable for observation.
Most of the birds have paired before their quiet isinvaded. They
are unwilling to leave the breeding-places they have chosen, and
soon become accustomed to the busy crowd, which passes them
by unheeded. For the species that feed on insects and carrion
there is the attraction of a food supply, the like of which they
can find nowhere else.
An official at the camp, however good his knowledge of the
subject might be, has but little leisure. To the ornithologist an
interesting but out-of-the-way district is rendered easily accessible
by the steamers which during the fishery run from Colombo to
the camp. His visit could not fail to be of interest to himself.
and would, I feel sure, add considerably to the knowledge of the
avi-fauna of Ceylon.
KANDYAN HORN COMBS. 151
KANDYAN HORN COMBS.
By ANANDA K. COOMARASWAMY, B.Sc., AND ETHEL M.
COOMARASWAMY.
With three Plates and two text-figures.
S its title shows, the present paper has no reference to the
merely ornamental fortoise-shell combs now worn by the
low-country Sinhalese (a comparatively modern fashion), nor to
the practical and beautifully carved zvory combs formerly used,
and still often preserved in Kandyan families ; theseivory combs
are pretty well known and are seen in most collections; there is
a fine series of them in the Colombo Museum. Horn combs,
though more generally used, are less familiar, but in some respects
even more interesting than the ivory ones ; for the latter, though
for the most part certainly made in Ceylon, sometimes seem to
show Dravidian influence, and at any rate are often very like
Indian combs of the same type, while the horn combs are more
purely native in form and decoration.
A short account of the making of the ordinary 10-cent horn
comb (Pl. B, fig. 3), used even by the poorest classes,* will now
be given. The horn comb-makers are men of low caste, Rodiyas
in the Bandarawela district for example, and blacksmiths (achari
caste) near Kandy. At Ratnapura and Balangoda the work is
done by Beruwayas and Durayas ; it is never done by goiyas,
who would consider it very degrading. The pictures and descrip-
tion apply specially to the manufacture of combs by Rodiyas
near Bandarawela.
The combs are made of buffalo horn cut transversely, the tops
of the horns being used for tool and knife handles and the like.
The usual tools of the horn comb-maker include a small adze (14
inch cutting edge), a coarse rasp, one or two knives, a marking
awl, a vice, two or three saws, a file, and one or two tools for
incising patterns ; the iron tools are made by local blacksmiths.
A flat piece of horn is sawn off and cut roughly into the
shape of the comb, and then made thinner with adze and rasp
(PI. A, fig. 1), and the shaping then finished with a knife. Next
the guide lines for the inner ends of the teeth and for the patterns
* Those who are better off would now use ordinary European-made combs,
instead of decorated horn or ivory ones of native make.
152 SPOLIA ZEYLANICA.
are scored with the marking awl, which is a small bone tool with a
sharp point inserted near one end. The comb is now set in the
vice, which is held conveniently against a stone by pressure of the
worker’s leg (Pl. A, fig. 2).
The vice (fig. I.) consists of two pieces of wood tied together
near one end, and having a wedge inserted at the other, which,
when it is driven in, causes the short ends to grip very tightly
whatever is placed between them. The vice is usually orna-
mented with a little primitive carving. The teeth are now ent
(P1.. A, fig. 2), first on one side, small teeth with a fine saw, then
on the other, coarse teeth with a thicker saw ; these teeth are cut
very regularly and with astonishing quickness. Each tooth is
next filed back and front to make its edges smooth. Finally the
comb is taken out of the vice and a simple pattern incised with a
chisel-shaped iron tool, as in the upper comb (PI. B, fig. 3), or
Cc
Fig. I.
A, the vice, length 114 inches (Bandarawela District).
B and C, two pattern-marking tools (Pallebedde, Atakalan Korale, Province of
Sabaragamuwa) (reduced ).
with a two or three-pointed tool asin the lower comb (Pl. B, fig. 3).
The pointed tools (B and ©) in fig. I. were used in ornamenting
the comb last referred to, which was made at Pallebedde, Atakalan
Korale, Province of Sabaragamuwa. Last of all the comb is
polished by rubbing with wood ashes and horn shavings.
More elaborately carved and pierced horn combs are sometimes
met with, but their actual manufacture has not been seen by us,
and it is very doubtful if they are any longer made.
The comb shown in fig. [I. is pierced, and deeply carved with
a characteristic Kandyan pattern. Fig. 4, Pl. B, shows a
more elaborate comb, carved and pierced with the nari lata
(woman creeper) design. The nari latd is a well known and
interesting Kandyan pattern, consisting essentially of a female
figure holding a creeper in each hand. In the horn comb shown
the figure seems almost to spring from the tree itself, suggesting
a possible origin of the design in the idea ofa sacred tree with its
KANDYAN HORN COMBS. ] 53
in-dwelling or guardian spirit ; but in other examples the lower
part of the figure is fully developed, and the dress carefully
indicated (as for example on an embroidered betel bag in our
possession), and the pattern is a good deal less suggestive of a
mythological origin ; it seems likely, however, that it may once
have had a symbolical significance of the sort suggested. The
design is a very usual one and is found on all sorts of Kandyan
decorative work.
A still more elaborate comb is seen in fig. 5, Pl. C3; this
unusually large comb is carved and pierced with a hansa putuva
design (geese with intertwined necks), and four heads of makaras
in the corners, and inlaid as well with ivory pegs, square for one
bird and round for the other. This hansa putuva is also a well
known Kandyan design; it may be found for example on such
Orr a>
Se est
Bd) : aN a2,
Ea ain
Fig. II.—Carved and pierced horn comb, Ratnapura District, x 2.
—_
various things as key plates, embroidered betel bags, and painted
pottery. It will be seen that these decorated horn combs are
striking and appropriate in character, and varied and individual
in their ornamentation, and it is a reproach to the modern
Sinhalese that there is now-a-days so little demand for good home-
made wares of this or any kind.
Nots.—Since this paper on Horn Combs was written it has
occurred to us that the nari lata design so frequently seen in
Kandyan decorative art may possibly be derived from the early
representations of the goddess Siri (Lakshmi), seated on a lotus
and holding lotus flowers in either hand. If so, the abundance
of foliage is a late rather than a primitive feature; but its
154 SPOLIA ZEYLANICA.
development from the original simple sprays appears quite
natural considering the fondness for “ creeper” decoration shown
in Kandyan design. On this view the most elaborate types, in
which human faces appear like flowers amongst the foliage (the
main stems of which are always grasped in the two hands of the
seated figure), would be a still later development,
A. K, C.
EH. M. C.
DESCRIPTION OF PLATES.
PLATE A,
Rodiyas making Horn Combs at Ella, near Bandarawela.
Fig. 1.—Using the rasp; a vice, knives, and other tools are seen
in the foreground, ready for use.
Fig. 2.—Cutting the teeth ; an adze is seen on the right.
PLATE B.
Fig. 3.—Above : horn comb made by Rodiyas at Ella; incised
pattern done with a chisel-shaped tool. Below: horn
comb made at Pallebedde (Sab.) ; incised pattern done
with the three-pointed tool shown in the text, fig. I.
x §. Authors’ collection. |
Fig. 4.—Horn comb carved and pierced with nazi laté design.
x 4. Kegalla Kachcheri collection.
PLATE C.
Fig. 5.—Horn comb carved and pierced with hansa putuva design
and four heads of makaras, and inlaid with ivory pegs.
x 8. Authors’ collection.
Fig. 6—Another example of the ndri latd design, from an
embroidered betel bag. x 3}. Authors’ collection. The
embroidery is chain stitch in red and white cotton on
(Kandyan) blue cotton. Compare this with fig. 4, Pl. B.
ZHYLANICA.
SPOLIA
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CARVED AND PIERCED HORN COMB WITH HANSA PUTUVA
NARI LATA DESIGN ON EMBROIDERED BETEL BAG.
Lo be compared with fig. 4, Plate B,
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NOTES. 155
NOTES.
1. On the Loris in captivity.a—With any wild animal in ecapti-
vity the first and most important thing to find out is what
food is most likely to suit its taste. And with a very small
nocturnal beast, such as the Loris, concerning whose habits I
imagine that but little is Known, this is not at first easy. A
list of what I have found Loris eat and thrive on, and of what I
have known him eat and not thrive on, may therefore be of
interest.
First of all, I am convinced that the Loris is by nature purely
insectivorous, and even carnivorous. If not fed for a day or so he
will eat plantains, pumpkins, boiled rice, &c., so will a hungry
leopard eat rice, or a starving man his boots, but not with any
marked gusto.
What a Loris really enjoys is a heavy meal of grasshoppers ;
all varieties seem to be equally appreciated, from the large brown
red-underwinged piéce de résistance to the thin pale green
“salad.” Ihave known a Loris eat 60 mixed grasshoppers at a
sitting. Crickets, moths, ordinary flies, most beetles, and cock-
roaches are all accepted eagerly ; while the rather horrid skill
with which a loris seizes and manipulates a strongly struggling
gecko seems to suggest that this is a not unfamiliar prey.
Worms, brightly-coloured butterflies, certain evil-coloured
beetles, and meat, such as chicken, beef, &c., are rejected, some-
times rather indignantly, and I have known the brown shell-
backed variety of cockroach (I do not know its name) make a
Loris very sick.
About a teaspoonful of water a day seems to be the correct
quantity of liquid, but I fancy the little beast can go fora long
time without any drink at all ; he must often have to do without
it in the hot windy weather from June to September.
After food, perhaps the next most important subject is medicine.
With the Loris I have only experienced two forms of ill-health,
diarrhoea and cramp, and both seem curable by the same remedy,
viz., opium, taken in the form of laudanum; three drops in a
teaspoonful of milk and water is in my experience a safe dose,
and it can be repeated twice in a day if the patient can be got to
take it ; he can be forced to do so of course, but he is rather frail.
we 8(17)05
156 SPOLIA ZEYLANICA.
If moral characteristics be added to physical, the Loris is very
high in the scale ; for his manners are excellent, rather like those
of an old-fashioned child who does not play, but takes his
pleasure in some more dignified manner. For the Loris never
plays; life is for him a very serious matter. Except the Pan-
golin, all other animals I have kept or known, play, all but the
Loris. If one comes to think of it, any Loris who in his natural
haunts attracted attention by frisking about and playing, would
very soon be snapped up by an owl or other enemy, for he is
very weak, and very slow, and his safety, like that of a new boy
in a public school, must depend almost entirely on unostentatious
self-effacement. Hence his good manners.
But besides modesty he has one other means of defence, and a
very curious one indeed it is. For he imitates a cobra. Before
relating the instance in which I saw this done, I must supportmy
testimony by the only reference to this habit that I know of.
Sinhalese of the North-Central Province have several times told
me little stories about the “ Unahapuluwa,” and I have thought
them far-fetched and absurd. And among the stories of its habits
told me by jungle men has been this : “ That the Loris copies the
sound of the cobra as a means of self-defence.”
Now that I have had an opportunity of testing this story and
of finding it true, the other yarns do not seem so absurd. It
happened this way.
I was sitting in the verandah spoiling my eyes by reading by
the last flicker of afterglow, when I heard the regular breathing
sound made by acobra when he inflatesand deflates his body. A
cobra moving ordinarily, or hunting anything much weaker than
himself, does not make this sound, but only when with expanded
hood he sits up to fight.
With the breathing sound came the occasional quick hiss of a
strike. So I got up and took a stick, for I thought that a cobra
might be attacking my Loris, who was not in his cage, but only
tethered to the top of it.
The sound came from my room, where, although it was dusk,
there was plenty of light to kill a snake.
As I went into the room I looked at the cage, which was on the
floor, and on the top of it I saw the outline of a cobra sitting up
with hood expanded, and threatening a cat who crouched about
sixfeetaway. This wasthe Loris, who, with hisarms and shoulders
hunched up, was a sufficiently good imitation of a cobra to take
me in, as he swayed on his long legs, and every now and then let
out a perfect cobra’s hiss. As I have said, it was dusk at the time,
but the Loris is nocturnal, so that his expedient would rarely be
NOTES. 157
required except in the dusk or dark; and the sound was a perfect
imitation. I may mention that I have kept snakes, including a
cobra, and am therefore the less likely to be easily deceived by a
bad imitation.
And, as I said above, the native legend bears me out.
Another native legend concerning the Loris, which obtains
universal acceptation in the North-Central Province, is that he
kills peafowl by creeping slowly up their backs when they are
roosting, and then, twining his long arms round the poor bird’s
neck, clings on until he has bitten to the brain.
The Loris is a charming and most original pet; his fault is
peevishness. But any one who has seen one thrust his long arm
into a bottle full of grasshoppers, pick one out and eat it like a
radish ; who has seen one sit on the table at dinner and catch the
moths and “ poochies ” that fly against the lamp; or has felt one
climb up his face and pounce into his moustache as into an
insect—will agree with me that our not so very distant cousin
is as fascinating a being as any in the forests of Ceylon.
JOHN STILL.
Anuradhapura, February 27, 1905.
2. On the larval habits of the butterfly, Parata alexis, Fabr.
Peradeniya, 27th March, 1905.—It is not often that the larve
of Hesperide (“ Skipper” butterflies) occur in large numbers in |
close proximity to each other. Parata alexis appears to be an
exception in this respect. On the 19th March I observed a fair-
sized tree of Pongamia glabra that had been completely
defoliated by the larve of this butterfly. So much so that they
had been compelled to descend and pupate amongst the foliage
of the shrubs below, for lack of shelter on the tree itself. There
must have been many hundreds of these pupz within a space
of a few square yards. From 50 pupze—collected at haphazard—
there emerged 18 males and 16 females; the remainder failed.
The flies were disclosed on two successive days (March 24 and 25).
On the first day 5 males and 12 females appeared ; on the second
day, 13 males and 4 females, the proportion of the sexes being
almost exactly reversed.
EK. ERNEST GREEN.
3. Curious behaviour of a snake in captivity. Peradeniya,
27th March, 1905.—A small example of Coluber helena which has
been an occupant of my vivarium for several weeks, and has
158 SPOLIA ZEYLANIOA.
hitherto exhibited a gentle and even timid disposition, commenced
to behave in a quite extraordinary manner to-day. On looking
into the cage I found it tightly coiled round atoad many times too
large for its capacity, visibly constricting it, but not attempting
to swallow it. It presently abandoned the toad and then struck
out wildly at anything that approached it, including my hand,
though I have previously repeatedly handled it with impunity.
The excited helena then—without any provocation—seized a
large “‘ Whip-snake” by the middle of the body, around which it
instantly coiled itself in a tight knot, remaining in that position
for half an hour—its head free, but its body tightly constricted—
defying all attempts of the whip-snake (which appeared greatly
inconvenienced) to dislodge it. When offered a small lizard—
as a distraction—it bit at it viciously, but would not retain it. A
large T'ropidonotus stolatus was next assailed in the same manner,
but that snake easily threw it off. In the intervals of these
attacks it moved about the cage with the anterior part of its body
laterally compressed and twisted to one side, thereby displaying
to unusual advantage the white lateral ocelli. The constricting
habit was most marked, and is probably employed when capturing
prey ; but I have not yet detected it in the act of feeding.
K, ERNEST GREEN.
4. On the nesting of the snake, Bungarus ceylonicus.—I have
received specimens of some eggs and young snakes which I have
identified as Bungarus ceylonicus, Giinth. My correspondent
tells me that they were found in a shallow depression, beneath a
wood-pile, in the month of January, 1905. ‘Both parents were
curled up in the hollow (made like a duck’s nest, but not lined
in any way, just scooped and hollowed out of earth), and under
them were lots of eggs and little snakes.”
The eggs have a soft leathery shell and are lightly agglutinated
together. They are cylindrical, with rounded ends, varying in
length from 29 to 35 mm., with a breadth of 17 mm, The young
snakes are brownish black above, with well-defined white annuli;
white beneath. The two specimens in my possession show 24
and 16 annuli respectively, exclusive of the broad fascia across
the head and the white tip to the tail. They measure 230 and
260 mm. in length. Even at this early age the vertebral row of
broad hexagonal scales is well demarked. The parent snakes
were unfortunately destroyed without any measurements being
taken.
KE. ERNEST GREEN.
NOTES. 159
5. Millipede killed by Reduviid bug.—While watching a large
millipede crawling over my lawn I noticed an immature
specimen (nymph) of the Reduviid bug (Physorhynchus linnet,
Stal.) following and manceuvring about it. Suddenly the bug
closed and fastened upon the hinder extremity of the millipede,
plunging its proboscis into the intersegmental tissue on the
ventral surface between the legs. The millipede immediately
writhed about apparently in great pain, but failed to dislodge
its enemy. It tried to crawl away, but rather more than half the
body and limbs were paralyzed. After feeding for about a
minute the bug left its victim, which I then picked up and
confined in a box, meaning to note how soon the paralyzing
effects would pass off. Twelve hours later the millipede, instead
of having recovered, was found to be quite dead. The victim
measures 53 inches in length with a proportionate thickness.
The predatory bug was scarcely } of an inch long. The bite of
many of the Reduviid bugs is—as I have experienced—extremely
painful; but I was astonished to find its action so fatal in this
case. The poison evidently acted directly upon the ventral
nerve cord.
Peradeniya, May, 1905. K. ERNEST GREEN.
6. Remedies adopted against the Paddy Fly.—One of the great-
est enemies of the paddy plant (Oryza sativa) is the so-called rice
sapper or paddy fly (Leptocorisa varicornis), called in the vernacular
goyan-messa.
The common remedy adopted against this pest is smoking the
fields by smother-burning vegetable refuse to windward. The
suffocating effect of the dense smoke that is thus raised is some-
times intensified by adding to the burning mass such substances
as tar, sulphur, tobacco, leaf, and margosa oil (from Azadirachta
indica).
The paddy fly is believed to object strongly to the odour of
resin, and for this reason cultivators often draw a rope saturated
in melted resin across the field.
The flies are sometimes captured by means of “ bird-lime,” in the
following manner. A winnow smeared with the sticky latex of
the jak tree (Artocarpus integrifolia) is fixed to the end of a pole
and drawn over thetips of the plants. Thefliesas they are captured
are collected in a pot strung to the waist of the operator and after-
wards destroyed. The process is carried on in the early morning
or late in the evening.
A method of decoy is also employed thus. An earthenware
160 SPOLIA ZEYLANIOA.
pot riddled with holes is set up in the field with a burning
lamp inside, The flies collect round the heated pot and are
thus destroyed.
But there are often extraordinary “ remedies ” (favoured by the
more superstitious) carried out through the agency of the village
soothsayer. One form of this is where the soothsayer, choosing
his day and hour, enters the field with some tender cocoanut
leaves plaited into a rough mat or “ ola,” and supervises the erection
of a tiny booth or “ mal-messa” on which a floral offering is placed.
In this booth the soothsayer burns some camphor, while he
indulges in a weird incantation. The final act in this ceremony
consists of the stretching of ropes from the mal-messa to various
points in the boundaries of the field.
The following is another form of charming. The soothsayer, as
before choosing his day and hour, proceeds to the field clothed in
spotless white, erects a mal-messa on which he gets some rice
boiled in cocoanut milk (the expressed juice of the kernel of the
cocoanut), and not the liquid contents of the cocoanut as generally
understood, and then with a branch from the inflorescence of
the areca palm proceeds to sprinkle the liquid from the boiling
pot over the fleld, muttering incantations the while.
A variation of this last consists in sprinkling charmed water
from a new clay pot by means of the twig of a lime tree.
In some Catholic villages it is the custom to carry a few paddy
flies tothe nearest church, there to be anathematized, and let loose
again in the field to influence the colony of pests to quit.
For much of the information in this note Iam indebted to Mr.
P. Samaranayaka, late Agricultural Instructor, now of the Veteri-
nary Department.
C. DRIEBERG.
Government Stock Garden,
June 15, 1905.
7. The Lacteal Tract of Loris gracilis—In Flower and
Lydekker’s well-known “ Introduction to the Study of Mammals”
(London, 1891) it is pointed out that in the order Primates “there
are always two mamme in the pectoral region, except in Chi-
romys”’ (l.c., p. 681).
The long-tailed African lemurs of the genus Galago have four
teats, namely, two pectoral and two inguinal (/.c., p. 690).
In the tailless Oriental lemurs belonging to the sub-family
Lorisinz there are said to be two pectoral mamme only (/.c., p. 691).
Chiromys, the Aye-aye of Madagascar, has two inguinal mamme
only (/.c., p. 695).
NOTES. 161
In a much more recent treatise on Mammalia by Dr. Max
Weber (Die Siugetiere, Jena, 1904) the statement that the
Lorisine are distinguished from the Galagine, among other cha-
racters, by the possession of two pectoral teats only, is repeated
on p. 760 of that work,
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Situs mammarum of Loris gracilis, Ceylon.
The villagers who, Iam told, procure the Loris in Ceylon, while
snaring monkeys for purposes of trade, occasionally bring them
alive to Colombo for sale. I have recently procured two females,
each carrying a young Lorisine clasped to her bosom. LEach of
162 SPOLIA ZEYLANIOCA
the mothers had four equal pendulous mamme placed conse-
cutively in two pairs in the posterior pectoral (below the last rib)
and in the anterior abdominal regions, surrounded by a nearly
hairless tract.
Each teat is about a quarter of an inch in length; the anterior
pair is distant about one inch and a quarter from the axillary
region ; the posterior pair is about two and a half inches from the
inguinal region.
The tetramerous arrangement of the teats in the Loris gracilis
of Ceylon is a constant character, and may be observed in young
females as well as in females during lactation. ‘he more primi-
tive mammals of the orders Edentata and Sirenia, represented in
Ceylon by the pangolin and the dugong respectively, have only
one pair of teats in the axillary region, but this fact does not
militate against the idea that the four teats of Loris may bea
primitive feature, at least within the limits of its own order.
Not only the organization but the strictly arboreal habits of the
Loris suggest that it is a creature of remote antiquity. Whether
the character of the lacteal tract affords a further indication of
this may be a matter of opinion, but the numerical data should be
correctly given. The plurality of teats is remarkable also on
account of the fact that the Loris, like bats and monkeys generally,
only produces one young ata birth, which remains clamped to
the parent by its extraordinary prehensile limbs until able to look
after itself.
ARTHUR WILLEY,
Colombo, July, 1905.
SPOLIA ZEYLANICA.
163
HELMINTHES FROM THE COLLECTION OF THE COLOMBO
MUSEUM.
By Dr. O. VON LINSTOW.
( Gottingen.)
With three Plates.
HIS report on Parasitic Worms from Ceylon is a continuation
part IV. (1904).
of my description of Nematodes in Spolia Zeylanica, vol. I.,
There are altogether in the second consignment
which was sent to me fifty-one species, of which thirty belong
to the Nematoda, one to the Acanthocephala, seven to the
Trematoda, and thirteen to the Cestoda; of these, thirty are
described as new, and three new genera of Cestoda have had to be
established,
The following species are dealt with :—
bo no 89 bo LO
Pian be
N
Se Re ee ee ee
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NEMATODA,
Ascaris spiculigera, Rud., from Plotus melanogaster.
Ascaris fissicollis, n.sp., from Haliastur indus.
Ascaris coronata, n.sp., from Ardeola Grayt.
Ascaris brachycheilos, n.sp., from Tropidonotus asperrimus.
Physaloptera brevispiculum, n.sp., from Felis rubiginosa.
Spiroptera secretoria, n.sp., from Plotus melanogaster.
Spiroptera orca, n.sp., from Manis pentadactyla.
Spiroptera sanguinolenta, Rud., from Canis familiaris.
Sptroptera, spec. ? from Cittacincla macroura.
Spiroptera, spec. ? from Pawo cristatus.
Spiroptera, spec. ? from Sciurus palmarum.
Heterakis pusilla, n.sp., from Gallus Lafayettt.
Heterakis granulosa, n.sp., from Gallus gallinaceus.
Strongylus digitatus, n.sp., from Bos taurus.
Kalicephalus willeyi, v. Linst., from Typhlops braminus.
Oxyuris megaloon, n.sp., from Hemidactylus Leschenaultir.
Oxysoma falcatum, n.sp., from Nicoria trijuga.
Dispharagus macrolaimus, n.sp., from Plotus melanogaster.
Sclerostomum equinum, Miiller, from Hquus caballus.
Ankylostomum minimum, n.sp., from Felis rubiginosa.
Ankylostomum trigonocephalum, Rud., from Canis familiaris.
Syngamus trachealis, v. Sieb., from Gallus gallinaceus.
Filaria ? Zschokkei, Meyer, from Manis pentadactyla.
Filaria immitis, Leidy, from Canis familiaris.
8(17)05
ol.
32.
33.
34.
35.
36.
37.
38.
39.
40.
41.
42.
43.
44.
45.
46.
47.
48
49.
50.
51.
SPOLIA ZEYLANICA.
Filaria piscicola, nu.sp., from Marine Fish.
Filaria equina, Abilg., from Equus caballus.
Filaria digitata, n.sp., from Bos indicus.
Filaria tuberosa, v. Linst., from Mabwuia carinata.
Filaria flavescens, Castellani and Willey, from Calotes versicolor.
Trichocephalus discolor, nu.sp., from Bos indicus.
Tetradenos tiara, v. Linstow = Ctenocephalus tiara, v. Linst.
ACANTHOCEPHALA.
Echinorhynchus tener, n.sp., from Spilornis cheela.
TREMATODA.
Lyperosomum squamatum,n.sp.,from Dissura episcopus.
Distomum, spec. ? from Plotus melanogaster.
Distomum hepaticum, L., from Bos indicus.
Paramphistomum calicophoron, Fisch., from Bos indicus.
Paramphistomum gracile, Fisch., from Bos indicus.
Gastrothylax crumenifer, Crepl., from Bos bubalus.
Tristomum megacotyle, n.sp., from Histiophorus.
CESTODA.
Hymenolepis septaria, n.sp., from Upupa ceylonensis.
Hymenolepis clausa, n.sp., from Dendrocygna javanica.
Hymenolepis spinosa, n.sp., from Rostratula capensis.
Dipylidium caninum, L., from Canis familiaris.
Tenia, spec. ? from Haliastur indus.
Diorchis occlusa, n.sp., from Phenicopterus roseus.
Davainea polycalcaria, n.sp., from Corvus macrorhynchus.
Diplochetos volvulus, n.gen. et sp., from Lobipluvia malabarica.
Ophryocotyle ceylonica, n.sp., from Lophoceros gingalensis.
Brochocephalus paradoxus,n.gen. etsp., from A’gialitis mongolica.
Cittotenia bursaria, n.sp., from Lepus nigricollis.
Ichthyotenia cryptobothrium, n.sp., from Chrysopelea ornata.
Aphanobothrium catenatum, n.gen. et sp., from Phenicopterus
TOSeuUs.
I.—NEMATODA.
Ascaris.
The definition given previously (S. Z., part 1V., p. 91) is to be
amended in so far that the intestinal ccecum lies dorsad of the
cesophagus ; the cesophageal gland is produced backwards ventral
to the intestine.
1.—Ascaris spiculigera, Rud.
From the csophagus and stomach of the Darter, Plotus
melanogaster, L.; Wirawila, Southern Province.
2.—Ascaris fissicollis, n.sp.
Pl. I, figs. 1-2.
From intestine of the Brahminy Kite, Haliastur indus, Bodd.;
Nedunkeni, Northern Province.
HELMINTHES : COLOMBO MUSEUM. 165
Three females, 15, 34, and 36 mm. long, 0°79-1'25 mm. wide.
Cuticle annulate; behind the lips the annulation is so deep
that the contours appear fringed. Lips with intermediate lips;
without teeth ; dorsal lip semi-circular, ‘083 mm. x ‘11 mm.”
the two papille lie in front ; intermediate lips obtusely conical,
attenuate from the middle, with an outer groove. Cisophagus §,
acuminate tail ,),.t Eggs thin-shelled, spherical, 065 mm.; the
small vitellus distant from the shell.
3.—Ascaris coronata, n.sp.
J2iL Uy ate ok
From the csophagus and stomach of the Nestling Pond
Heron, Ardeola Grayi, Sykes ; Tissamaharama.
Cuticle narrowly annulate ; lips edentulous with large trian-
gular interlabia ; dorsal lips nearly circular, ‘065 x -078, with two
inner anterior prominences ; papille anterior; cesophagus ¢.
Male, 17 x :79; tail conical, very short, ;4;. On each side of
the caudal extremity 17 pre-anal papille in a row becoming
more closely placed behind ; post-anal papille absent; the cirri
are 2°37 mm. long, straight, the end bent falciform.
Female, 26 x 1:5; tail rounded, anus nearly terminal ; at the
posterior end a small finger-shaped appendix ; the vulva lies in
front of the middle, dividing the body in the ratio 7: 15; eggs
immature.
4.— Ascaris brachychetlos, n.sp.
Pl. IIL., fig. 48.
From intestine of Tvropidonotus asperrimus, Boulenger ;
Colombo,
Cuticle smooth ; lips depressed, dentiferous, with small conical
interlabia; pulp emarginate; papille large; dorsal lip
078 x -177; cesophagus ;},- 1; ; tail in male 545, in female ;3,,
rounded in both sexes carrying a small terminal appendix.
Male, 55 x *83; cirri broad, curved like a sabre, rounded at
the end, 1:19 x :035; at each side of the tail three very small
pre-anal and two post-anal papille; the last of the latter lies
dorsally exactly at the hinder end of the body dorsad of the
styliform appendix.
Female, 104 x 1°58; anus nearly terminal ; vulva near the
end of the anterior third, dividing the length in the ratio 14: 39 ;
eggs spherical, thick-shelled, closely beset with small granules,
086 mm.
* All measurements are in millimetres, and the length always precedes the
breadth.
} These are fractions of the total body length throughout.
166 SPOLIA ZEYLANICA.
d.—Physaloptera brevispiculum, n.sp.
Pl. L, fig. 4.
From stomach of Felis rubiginosa, Geoffr.; Kandy.
Cuticle finely annulate ; head end with a thickened cuticular
collar from which two round lips protrude, each of which carries
three small peaks at the summit; at the tail end there is a simi-
lar preputium-like cuticular ring from which the tail emerges ;
cesophagus z=.
Male, 11:1 x °95, with rounded tail 344 ; at each side of the
cloacal orifice four stalked papille ; behind these four paired
ventral post-anal papilla, unstalked, in successive couples; cirri
very short, feebly curved, ‘79-81.
Female, 11-4 x 1:06; tail conical, 545; eggs thick-shelled,
036 x ‘0381.
P. preputialis, v. Linst., which occurs in Felis catus in Brazil,
has a similar caudal sheath with a body length of 21-30 mm.
6.—Spiroptera secretoria, n.sp.
PIS, feb}
From esophagus and stomach of Plotus melanogaster, L. ;
Wirawila ; in company with Ascaris spiculigera.
Dimensions up to 382 x 1:6, but all specimens immature; both
ends strongly attenuated ; cuticle annulate; some are larve in
process of exuviation. At the head a dorsally placed obtusely
conical papilla; in a circle behind this are six roundish papilla,
and behind these in the submedian lines four truncate papille
with a very small one at the inner side. Alongside the
cesophagus runs a long gland containing a long granular
secretion often projecting through the orifice which lies close
behind the papille. Tail short ‘with small finger-shaped
appendix; lateral lines strongly developed, one-fifth of the
diameter of the body, enclosing a lateral canal.
7.—Spiroptera orca, D.sp.
Pl. L., figs. 6-8.
From stomach of Manis pentadactyla, L.; Horana.
Cuticle annulate; head with two large lips placed dorsal and
ventral, expanded in front; behind these in the submedian lines
four finger-shaped procumbent processes.
The mouth leads into a vestibule, ‘2 mm. long; csophagus
in the male 4, in the female s+; coarse cuticular rings ‘053
apart, fine rings ‘0054 mm. apart.
HELMINTHES : COLOMBO MUSEUM. 167
Male, 25 x °71; tail involute; cirri long; the left cirrus
measured 3:74 mm. in a young specimen of 11°8 mm., ¢.e. nearly
one-third of the body length; the right cirrus is broader and
much shorter, ‘57 mm.; on each side there are four pre-anal
and two post-anal papille, large and round; tail rounded, 5.
Female, 32 x ‘95; tail rounded, 2,; vulva lies behind the
middle dividing the body in the ratio 7: 5; immature eggs
elliptical, 044 x :026; mature, flattened at the poles, ‘047 x :029,
barrel-shaped, surrounded in front and behind by a raised hoop.
8.—Spiroptera sanguinolenta, Rud.
From cesophagus of Canis familiaris, L.; Colombo.
Cf. A. Railliet, Traité de zoologie médicale, Paris, 1895, pp.
536-538, figs. 373-375.
9.—Spiroptera, spec. ?
From intestine of Long-tailed Robin, Cittacincla macrouwra,
Gmel.; Nedunkeni.
One entire and one half specimen spirally wound, not to
be identified.
10.—Spiroptera, spec. ?
From csophagus of Pavo cristatus, L.; Buttuwa.
Three imperfectly preserved fragments.
11.—Spiroptera, spec. ?
From peritoneum of Sciwrus palmarum, 1..; Colombo.
Fragments of a female.
12.—Heterakis pusilla, n.sp.
PL. L, fig. 9.
From rectal coeca of Jungle Fowl, Gallus Lafayetti; Mamadu,
Northern Province.
Head with small roundish lips; cuticle smooth; cesophagus
thickened, club-shaped behind, in the male 1}, in the femaie are
Male, 5 x 0°19; tail »',, finely pointed, adhering to the ventral
side by a granular, opaque cement; cirri very unequal, the
left 0°53, the right 0°15; in front of the cloacal orifice a round
sucker, and behind this, on each side, four large post-anal
papille, of which the last lies at a greater distance from the
third than the three anterior from each other.
Female, 5:13 x 0°24; tail gis. very long and pointed ; vulva
far behind the middle, the pre-genital region to the post-genital
region as 12: 5; eggs, thick-shelled, 0:065 x 0-031.
168 SPOLIA ZEYLANICA.
13.—Heterakis granulosa, n.sp.
Pl. III., fig. 49.
Intestine of domestic fowl, Gallus gallinaceus, L. ; Colombo.
Cuticle annulate ; head with three low semi-circular lips; tail
pointed.
Male, 27 x 0:59; cesophagus |, ; tail Jj); the two straight
rod-shaped cirri, 0°57; sucker long, oval, with a small circular
groove behind, and surrounded by granulations ; on each side
three pre-anal and six post-anal papille, of which the most
anterior is transversely elliptical, the fourth, seventh, and ninth
spherical and marginal ; radial muscles pass to the sucker.
Female, 55 x 0°79; wsophagus ;,; tail ~,; vaginal orifice
immediately in front of the middle dividing the body in
the proportion 16: 17; eggs thick-shelled, 0:078 x 0042.
14.—Strongylus digitatus, n.sp.
Pl. L., fig. 10.
Stomach of Bos indicus ; Colombo.
Cuticle without longitudinal lines, but with very fine annu-
lation; head attenuate, mouth surrounded by four papille ;
cesophagus of male +, of female ,', ; the nerve ring surrounds the
cesophagus at the limit of the first and second quarters and
below it the porus excretorius opens.
Male, 24 x 0°36; cirri very long, 4°54, coalescent throughout
their entire length and thickened fusiform at the end; the
lateral lobes of the bursa are curved claw-like inwardly; each
lobe is supported by six ribs, of which one lies at the inner
margin, two side by side at the hinder border, and three in
a group at the outer margin ; there is no median lobe; the end of
the body is rounded with a pair of finger-shaped, slightly curved
hyaline cuticular lobes.
Female, 29 x 0:47; genital orifice lies far back dividing the
body in the ratio 8: 1; tail pointed, ;j53 eggs 0097 x 0:053.
Eleven species of Strongylus have been found in Bos
taurus, the European domestic ox ; ten of them have short cirri;
only Strongylus filicollis, Rud., has long cirri, but in this species
the cuticle shows 18 longitudinal ridges.
15.—Kalicephalus willeyi, v. Linst.
Rectum of Typhlops braminus, Russell ; Colombo.
Cf. O. v. Linstow, Spol. Zeyl., vol. I., part IV., 1904, pp. 99-100
pl. L, figs. 14-18.
HELMINTHES : COLOMBO MUSEUM. 169
16.—Ozyuris megaloon, n.sp.
Intestine of Gecko, Hemidactylus leschenaultii, Dum. et Bibr.
Mamadu.
Females only in the collection, 6°52 x 0°91; cuticle deeply
annulate; head with three small lips; cesophagus <, narrow,
ending behind in a spheroidal bulb; tail conically pointed, + ;
eggs very large, 0:083 x 0:047.
Oxysoma.
Head with three or more lips ; cesophagus witha spheroidal bulb
behind ; male with two equal cirri and three paired pre-anal
papille ; number of post-anal papille variable; tail in both sexes
finely pointed; Secernentes-Meromyaria ; in reptiles and amphibia.
17.—Oxysoma falcatum, n.sp.
Pl. IIL., figs. 50-51.
Intestine of the Tortoise, Nicoria trijuga, Schweigg. ; Colombo.
Cuticle smooth; head broadly rounded with six stalked
papille ; the stalks are divided giving off an inner branch ;
behind the head are four large papille in the submedian lines ;
cesophagus thin, in the male oe, female me the bulb is
embraced by the cup-shaped commencement of the intestine ;
excretory pore behind the middle of the cesophagus dividing the
latter in the ratio 21 : 16.
Male, 11°6 x 0°55; tail 54, bent hook-like towards the ventral
surface ; cirri equal, falciform, very broad before the middle, 0°44,
pointed at the end; three pre-anal and five post-anal papille
(paired).
Female, 141 x 0°56; tail 54; genital orifice behind the
middle dividing the body as 5 : 3 ; the vagina runs forwards ;
eggs 0°14 x 0-097.
Dispharagus.
Head with two lips; in the region of the so-called neck the
cuticle shows four longitudinal pleats; male with two unequal
cirri ; on each side of the tail four pre-anal papille; it belongs
to the Secernentes-Polymyaria; occurring in the cesophagus,
stomach, and gastric submucosa of birds,
18. —Dispharagus macrolaimus, nusp.
Stomach ofthe Darter, Plotus melanogaster, L. ; Wirawila.
Females only in the collection, 7°3-11°4 x 0°28-0°47 ; cuticle
annulate, with elevated lateral lines; head with two small,
conical, rounded lips; the mouth leads into a long vestibule ;
170 SPOLIA ZBYLANICA.
cesophagus very long, ais <tail zy, terminating in a small finger-
shaped point; the nuchal pleats run 0°80 mm. backwards, rather
beyond the first section of the cesophagus; immediately before
their termination there is on each side a cone-shaped nuchal
papilla; eggs very numerous, with a double shell, 0-031 x 0°011.
Sclerostomum.
Head with buecal orifice set with one or several rows of teeth ;
male with two equal cirri and trilobate costiferous bursa ; female
genital orifice posterior ; Secernentes-Meromyaria ; in mammals
and birds.
19.—Sclerostomum equinum, Miller.
Intestine of horse ; Colombo,
Of. A. Looss, The Sclerostomidz, Records Egyptian Government
School of Medicine, 1901, pp. 76-77, pl. I., figs. 1-5.
Ankylostomum.
Head with chitinous buccal capsule, bent and open towards the
dorsal side ; often with teeth in its fundus; male with trilobate
bursa and two equal cir7i; bursal lobes supported by ribs;
female genital orifice behind the middle; Secernentes-Mero-
myaria; in the intestine of mammals.
20.—Ankylostomum minimum, D.8p.
Pl. L., figs. 11-12.
Stomach of Felis rubiginosa, Geoffr.; Kandy.
Cuticle thick and annulate; buccal capsule dorsally inclined
with four ventral longitudinal “ribs” and a tooth at the bottom.
Male, 2°38 x 0°24; cesophagus }; cirri brown and very short,
0-062 ; lateral lobes of bursa rounded, supported by six ribs, of
which the two anterior lie close together, the third, fourth, and
fifth form a group, and the sixth lies isolated.
Female, 4°46 x 0:20; cesophagus a ; tail 3,3; genital organs
almost confined to hinder half of body ; genital orifice posterior,
dividing the body as 31 : 12; eggs not numerous, 0°088 x 0-053 ;
one branch of the uterus runs forwards, the other backwards.
21.—Ankylostomum trigonocephalum, Rud,
Intestine of dog ; Colombo.
Of. A. Railliet Traité de Zoologie Médicale, Paris, 1895, pp.
470-473, figs. 8327-330.
HELMINTHES : COLOMBO MUSEUM. 171
Syngamus.
Mouth with a chitinous capsule, the wall of which is enlarged
dise-like in front ; male small with a buvsa and two equal cirri ;
female orifice anterior ; male and female in permanent copula ;
Secernentes-Meromyaria; in the trachea, bronchi, and nose of
birds and mammals.
22.—_Syngamus trachealis, v. Sieb.
Trachea of domestic fowl ; Colombo.
Of. A. Railliet, op. cit., pp. 453-455, fig. 312, also Neumann’s
Parasites of Domesticated Animals, p. 607, fig. 318.
23.—Filaria ? Zschokkei, Meyer.
Peritoneum of Manis pentadactyla, L.
Cf. Meyer, Archiv f. Naturgesch. Jahrg.. 61, Berlin, 1896, pp.
56-69, Taf. IV., figs. 1-9. Sexually immature.
24.—Filaria immitis, Leidy.
Pleural cavity of dog; Ragama.
Of. A. Railliet, op. cit., pp. 509-513, figs. 354-356.
25.—Filaria piscicola, n.sp.
From supraorbital region of a marine fish (? Lethrinus, sp.).
One incomplete female, 225 x 0:57; the diameter at the head
is 0°13; the head is rounded, destitute of lips, teeth, and papille ;
the tail is lost from the specimen ; genital orifice quite anterior,
only 0°79 from cephalic extremity ; eggs 0:031 x 0:023 ; it is ovo-
viviparous ; the embryos with acuminate tail measure 0°53 x
0-016.
Filarie are very rare in fishes, and it is therefore to be regretted
that only an imperfect description of this species can be given.
26.—Filaria equina, Abildg.
From eye of horse.
Of. A. Railliet, op. cit., pp. 524-526, figs. 364-366.
27.—Filaria digitata, n.sp.
Pl. IIL., figs. 52-55.
Peritoneum of Bos indicus ; Colombo.
Head with two straight teeth notched at the sumunit, thereafter
four papille in the submedian lines; cuticle finely annulate; a
long vestibule (in the female 0°59) leads into the cesophagus, }
2A 8(17)05
1 ly iy SPOLIA ZEYLANICA.
in the male, } in the female; tail rounded, carrying in the
female a spherical appendix; in the male the tail is coiled in
a close spiral, in the female a loose spiral; in the male 31,5;
in the female ,},; in both sexes in front of the extremity on each
side a finger-shaped appendage, 0-044 long in male, 0-10 in female.
Male, 42°5 x 0°35; the curved cirri 0°16 and 0-065 respectively ;
four pre-anal* and four post-anal* elongated papille ; in front
of the former on each side about 140 small rod-shaped, close-set
papille reaching forwards 1°9 mm. from the tail end.
Female, 56°9 x 0°55; genital orifice anterior dividing the body
as 1:71; eggs 0-039 x 0-026.
28.—Filaria tuberosa, n.sp.
Filaria Mansoni, Castellani and Willey, Sol. Zeyl., vol. I1.,
1904, pp. 79-80, pl. VI., figs. 1-6.
In the peritoneum of Mabwia carinata, Schneid., the Brahminy
Lizard ; Colombo. Larve in the blood.
Only two females in the collection, of which the larger measures
34 x 0°37; tail attenuate, ending with a hemispherical dilatation,
in front of which there is a papilla on each side; anus absent;
head rounded without teeth and papille ; cesophagus very short
#5 ; genital orifice 0°44 from head end; thin-shelled eggs 0-018 x
(013 ; ovoviviparous.
The original name of the species must be altered because
Cobbold named one F. Mansoni, from the orbit of Gallus
gallinaceus in 1880.
29.—Filaria flavescens, Castellani and Willey.
From Calotes versicolor, Dum. et Bibr.; Colombo.
Of. Castellani and Willey, imprim.
Head rounded without teeth and lips; body attenuate and
rounded behind ; anus absent ; esophagus very short, +’, in male,
gx in female. Male, 19 x 0°33; tail, ;}5, with five very small
post-anal papille ; cirri, 0°16, conical with very broad base.
Female, 56 x 0°63; genital orifice 1°97 from head end ; eggs
0-021 x 0:016 ; ovoviviparous.
Trichocephalus.
Body strongly attenuate, cesophagus cellular, hind body
thickened, anus terminal. Male with one spiculum; female
genital orifice at the end of the esophagus at the limit between the
* These papille are always paired unless otherwise given.
HELMINTHES : COLOMBO MUSEUM. 173
thin fore body and thick hind body; eggs barrel-shaped ;
belongs to the Pleuwromyaria, with muscles in place of the lateral
fields; in the cecum of mammals.
30.—Trichocephalus discolor, n.sp.
Pl. L, figs. 13-14.
From Bos indicus, Colombo; said to have occurred in the
stomach.
Cuticle annulate at intervals of 0:0091 mm.; contours, at the
fore body, serrate.
Male white, 45 x 0°14 in front, 0°55 behind; cirrus sheath
spinose ; cirrus long and narrow, of even width except at the
rounded apex, 1°76 x 0°011; hence not to be confounded with
T. affinis, Rud., from Bos taurus.
Female, 50 x 0:13 in front, 0°67 behind ; fore body white, hind
body yellow ; eggs 0°065 x 0-031, with large spherical opercula
at the poles.
Tetradenos tiara, v. Linst.
In Spolia Zeylanica, vol.1., 1904, p. 102, I described a Nematode
from Varanus bengalensis under the name Ctenocephalus tiara.
It happens that Tholenati gave the name Clenocephalus to a
Dipteron in the year 1857, and I have therefore altered the
generic name of the Nematode to Vetradenos.
Cf. Arch. f. Naturg., Berlin, 1904, p. 301.
Il.—ACANTHOCEPHALA.
Echinorhynchus.
Nemathelminthes without intestine; at the head a proboscis
beset with hooks which can be retracted into a proboscis sheath,
alongside which are two lemnisci (fluid reservoirs); male with two
testes, six cement glands (“ prostate”), and a protrusible, bell-
shaped bursa with penis; female with a ligament in which the
placentule arise; the mature eggs are passed through the
sphincter apparatus of the bell-shaped uterus into the vagina ;
the species live in the adult condition in the alimentary canal of
vertebrates.
31.—LHehinorhynchus tener, n.sp.
P). 1:, figs: 15-17.
Intestine of the Serpent Eagle, Spilornis cheela, Daud.; Nedun-
keni, Northern Province.
Body extended very long, delicate and fragile, head feebly
thickened ; rostellum broad and short, thickened in front, 0°43 x
0°39 (in front), the so-called “neck,” 0°28, closely covered with
174 SPOLIA ZEYLANICA.
hooks; rostellum hooks in forty-five transverse rings with twenty
hooks in each ring; hooks of the twelve anterior rings, 0:042,
with a long root, those of the thirty-three posterior rings, 0°021,
thorn-like without root.
Male, 39x 0°52; the group of genital organs extends forwards
9°45 mm. from the thickened tail end.
Female, 72 x 0°79; placentule ovate, 0°17 x -097; eggs with
double shell, the outer shell showing wavy longitudinal iines,
0:053 x 0°023; tail end after copulation carries a layer of cement
substance.
IIl.—TREMATODA.
Lyperosomum.
Distomids with long intestinal rami ; testes behind one another
and behind them lies the ovary; vitellarium on each side laterally
behind the ventral sucker ; genital orifice in front of latter.
32.
Lyperosomum squamatum, n.sp.
Pl. I, fig. 18.
(Hsophagus of the White-necked Stork, Disswra episcopus ;
Gould ; Palatupana.
Length 4°86, breadth 1°78. Oral sucker 0°59; ventral sucker
0:87, its middle point lying at the limit of the second and third
fifths of the body; the cuticle of the ventral surface alone closely
beset with scales; cuticle everywhere very thick; the thickness of
the body is to the width as 1: 2; the cesophagus divides after a
short course into two intestinal diverticula, the epithelium of
which is strongly developed ; they extend to the hinder end; the
genital orifice lies immediately in front of the anterior margin of
the ventral sucker; behind this the two testes follow one behind
the other and, behind the last testis, the small ovary with the
shell-gland ; the vitellaria occur as two tracts behind the ventral
sucker occupying about 4 of the body length; the coils of the
uterus liein the posterior half of the body; the eggs are small and
numerous, narrow and elongate, 0083 x 0:036; they present a
large double black spot which corresponds with the two ocelli
of the embryo; the cirrus sac is small, destitute of a cirrus.
33.—Distomum, spec. ?
Esophagus of the Darter, Plotus melanogaster, Lin.; Wirawila,
Only one opaque specimen which could not be determined
without sections, thus destroying it; moreover a single indivi-
dual would not have sufficed for the investigation. The genus
could therefore not be ascertained since the old genus Distomum
has been broken up into more than 80 genera.
HELMINTHES : COLOMBO MUSEUM. iG
34.—Distomum hepaticum, Lin.
Liver of Bos indicus, [Scinde Cow] ; Colombo.
Of. A. Railliet, Traité de Zoologie médicale, Paris, 1895, pp.
342-356, figs. 219-235.
The generic name Distomwm is here adopted instead of
Fasciola, which was established by Linneus in 1746 for
Distomum hepaticum, Dendrocelum lacteum, and Schistocephalus
solidus, these being taken to be one and the same species.
Fasciola, L., is therefore a scientific impossibility, incapable of
being diagnosed. The definition of Distomum, s. str., 18 as
follows :—
The two intestinal rami richly branched towards the outside ;
ventral sucker lying well forwards with the porus genitalis in
front of it, the uterus and branched ovary ‘behind; farther
back the much-branched testis and the vitellarium at the
margin of the body ; parasitic in the liver of mammals.
Paramphistomum.
A small sucker in front, a large sucker behind; intestinal
rami long; two testes lying obliquely one behind the other,
lobate, without cirrus sac; porus genitalis in front in the
median ventral line with the ductus ejaculatorius and the
uterus opening into it; ovary and shell-gland behind the last
testis; vitellaria lateral close beneath the surface of the body ;
parasitic in the stomach and bile passages of ruminants.
35.—Paramplhistomum calicophorum, Fisch.
Stomach of Bos indicus; Colombo.
Cf. F. Fischoeder, Die Paramphistomiden der Saugethiere,
Zool., Jahrb. Syst. XVII., Jena, 1903, pp. 541-546.
36.—Paramphistomum gracile, Fisch.
Stomach of Ceylon black cattle (Bos indicus).
Cf. Fischoeder, op. cit., pp. 520-524.
Gastrothylax.
Amphistomids with ventral pouch which commences as a
transverse groove shortly behind the mouth; genital pore inside
the ventral pouch, male and female genitalia immediately in
front of the caudal sucker.
37.—Gastrothylax crumenifer, Crepl.
Stomach of Bos bubalus, the country-bred buffalo of Ceylon.
Of. Fischoeder, op. cit., pp. 557-563.
176 SPOLIA ZEYLANICA.
Tristomum.
Body ovate or circular, flat, with two round suckers in front and
a large, disc-shaped posterior sucker with seven rays and small
rods ; cerebral commissure with four ocelli; genital orifices and
vagina in front on the left ventral side; testes numerous; ecto-
parasitic on the gills and surface of the body of marine fishes.
38.—Tvristomum megacotyle, n.sp.
Pl. L., figs. 19-20.
From the surface of the body of a sword fish, Histiophorus, sp.;
Beruwala, February 5, 1904. .
Length 7:5, breadth 6 mm.; ventral cuticle, except on the
suckers and head, thickly beset with circular papille, measuring
0°31 mm.
The two anterior suckers are equal to } of the body length;
the caudal sucker has a diameter equal to 4 the body length, and
shows seven radial ribs (pl. 1, fig. 19) which abut upon a central
roundish area produced backwards; the sucker is bounded at the
periphery by a striated border; in the posterior region of the sucker
there are two straight undivided rods attenuated at the ends, 0°48
mm.long; these rods are always described as hooks in the defini-
tion of the genus 7’ristomum, but they do not deserve this name
since they do not serve for attachment, but, on the contrary, for
releasing the sucker from its adhesion, assuming a vertical posi-
tion by the action of special muscles for this purpose.
A similar, though much larger species, also living upon Histio-
phorus, is Tristomum leve, Verrill=T7. ovale, Goto; the length of
this species amounts to 13 mm. and its breadth 12 mm.; the
diameter of the anterior suckers equals } of the body length; the
rays of the caudal sucker are narrow, its margin is unstriated,
and the rods are expanded and irregularly laciniate at the
roots.
Of. S. Goto, Studies on the Ectoparasitic Trematodes of Japan.
Journ. Coll. Science, Japan, VIII., part 1, Tokyo, 1894, pp.
241-244, .
IV.—CESTODA.
Hymenolepis.
Teniids with armed scolex; proglottids generally broader than
long ; genital orifices marginal and unilateral ; in each segment
three testes ; the mature uterus completely fills the proglottids ;
parasitic in mammals and birds.
HELMINTHES : COLOMBO MUSEUM. ET
39.—Hymenolepis septarva, n.sp.
PIPE het 21:
Intestine of Upupa ceylonensis, Reich.; Weligatta. :
Length 25 ; the proglotiids begin directly behind the scolex and
measure, in front, 0°13 broad x 0°022 long, farther back, 0°79
broad x 0°18 long; towards the end of the chain they become
longer than broad, 0°35 broad x 0:48 long, the last member round-
ed ; the dorsoventral diameter is to the transverse as 2: 3.
The scolex is short, 0°13 long x 0°22 broad, truncate in front ;
suckers 0:089 ; rostellum small and knob-shaped ; hooks absent,
no doubt through casual loss. On each side are two vessels, a
large dorsal and a small ventral; outside these occurs the nerve.
Bundles of longitudinal muscles appearing circular in trans-
verse section course beneath the thick cuticle ; calcareous bodies
absent.
Cirrus-sac with inner side directed obliquely ventrally ; three
small oval testes lie in a row in the middle of the proglottids at
the dorsal side. Vagina dorsad of cirrus sac, expanding to a large
roundish veceptaculum seminis, reaching the middle line and
touching the anterior margin of the segment. Ovary strongly
developed occupying the whole longitudinal extent of the seg-
ments; ventrally a broad transverse branch from which right and
left two broad cornua extend horseshoe-shaped towards the
dorsal side, leaving room for the testes, the vitellarium, and the
shell-gland ; the roundish vitellarium (“ Dotterstock ”’) lies in
the middle line ventrad of the testes; the small shell-gland is
still more ventral in position. The uterus fills the last segments
completely and is subdivided by dorsoventral septa; the eggs
have a triple membrane, the outer 0:073 x 0-064, the inner 0:031
x 0023.
40.—Hymenolepis clausa, n.sp.
Pl. II., figs, 22-23.
Intestine of the Whistling Teal, Dendrocygna javanica, Horst.:
Tissamaharama.
Length 18; proglottids commence at once behind scolex,
measuring in front 0°053 long x 0°35 broad, behind 0°47 x 1°56,
Scolex small, 0°10 x 0°23; suckers 0°10; rostellum hemis-
pherical, carrying eight hooks of 0-057 length. In the paren-
chyme are to be found two layers of longitudinal muscles, outer
small numerous bundles, inner large sparser bundles ; calcareous
bodies not present ; two very large vessels traverse the chain:
outside them the nerves.
Genital pores absent; on one and the same side in each pro-
glottid, + of the cross-diameter distant from the margin, the
178 SPOLIA ZBYLANICA.
cirrus-sac (bursa) and vagina merge into one another directly ;
three testes lie dorsally and posteriorly in each segment, the
central one somewhat backwards; cirrus-sac very large, about
3 of the cross-diameter of the body, containing a very long cirrus,
a uniforraly thin chitinous tube coiled several times ; the orifice
of the cirrus-sac where it passes into the vagina is closely beset
with small equal hooks. The ovary lies unsymmetrically ventrad
of the vagina and cirrus-sac,a racemose body, in the centre of each
follicle a black granular nucleus ; vitellarium ventral, median,
transversely elongate, in front of itthe round shell gland ; no eggs
developed.
I agree with Wolffhiigel in the opinion that the absence of
genital pores is not of systematic importance, and therefore the
species is assigned to Hymenolepis.
4).—Hymenolepis spinosa, n.sp.
Pl. II., figs. 24-25.
Intestine of the Painted Snipe, Rostratula capensis, Lin.;
Vavuniya.
Length 15 mm., anterior proglottids 0°12 broad x 0°044 long;
hindmost proglottids greatly expanded at the hinder margin,
0°62 broad x 0°35 long; all are therefore broader than long.
Scolex thickened towards the fore-body with breadth of 0°22,
the foremost portion of the chain being 0°11 broad.
The rostellum carries ten hooks 0°028 long; the hooks are
slender and have a long root and small hook and lever; genital
orifices marginal and unilateral, approximately at the end of
the first quarter of each member of the chain. The cirri are
remarkably large, 9°14 long, 0:018 broad at the base, closely spi-
nose. The broad cortical layer occupies on each side } of the
dorsoventral diameter; a layer of transverse muscles occurs at
its inner side and inside these numerous small groups of longi-
tudinal muscle-bundles, inside these again eight stronger bundles
of longitudinal muscles; on each side a large ventral and a
smaller dorsal vessel, ectad of these, the nerve.
The large cirrus-sac occupies nearly % of the transverse diam-
eter; dorsad in the middle of the proglottis, three large testes,
one in front beside the cirrus-sac, the two others side by side
farther back. The coiled vagina lies below the cirrus-sac and
expands to form a small receptaculwm seminis which does not
reach to the middle line; the ovary lies in the middle third of
the transverse diameter, behind it the vitellarium, a transverse
strand of about 4 the cross-diameter; the ovate shell-gland
HELMINTHES : COLOMBO MUSEUM. 179
occurs in the middle between the second and third testes. The
eggs have a triple membrane, the outer irregular, 0:047; the
oncosphere is 0:026 long x 0:018 broad.
Dipylidium.
Rostellum with several circlets of rosethorn-shaped hooks ;
genital pores marginal and bilateral; genital organs in each pro-
glottid duplicated ; testes numerous ; parasitic in mammals.
42,—Dipylidium caninum, Lin.
Intestine of Canis familiaris ; Colombo.
Of. A. Railliet, op. cit., pp. 284-290.
43.—Teenia, spec. ?
Intestine of Haliastur indus, Bodd.; Nedunkeni.
Defective fragments without scolex, indeterminable.
Diorchis.
Scolex with simple crown of hooks; in each member two
testes ; genital pores marginal and unilateral ; the mature uterus
fills the proglottids completely ; parasitic in birds.
44,—Diorchis occlusa, n.sp.
Pl. II., figs. 26-27.
Intestine of the Flamingo, Fhenicopterus roseus, Pall.; Weligatta.
Length 75 mm.; the body is thick, ovate in cross section, the
dorsoventral diameter is to the transverse as 7: 9; formation of
proglottids commences at once behind the scolex; anterior
proglottids 0°40 broad x 9°018 long, posterior 0°97 broad x 0°18
long, the breadth always exceeding the length and the contours
serrate.
The scolex, 0°53 broad, is triangular in profile ; the rostellum
is in some cases retracted, in others protracted, appearing short
and broad ; it carries eight hooks, 0:14 mm. long ; the root-branch
is shorter than the hook, in the proportion 25: 29; at the base
there is a small finger-shaped prolongation, and the concavity
thus produced articulates with a roundish body, eight of which
occur in a circle at the summit of the rostellum.
Genital pores absent ; male and female ducts fuse together on the
same side in all segments at a distance of 0°03 from the margin ;
radial bundles of longitudinal muscles occur not far from the
cuticle ; calcareous bodies are present in small quantity ; on each
2B 8(17)05
180 SPOLIA ZEYLANICA.,
side a large ventral and small dorsal vessel, the latter strongly
and regularly sinuate, outside these the usual nerve, round in
section.
A long thin convoluted chitinous cirrus lies in the cirrus-sac,
the duct is finely spinulose, and a seminal vesicle leads into the
cirrus-sac ; two small oval testes, 0°10 x 0:06 lie dorsally.
The wide vagina is ventral to the cirrus-sac; it is finely
spinulose internally and presents a forcipate apparatus ; it leads
into a receptaculum seminis reaching to the middle line; the
fusiform dilatation of the vagina is 0°042 wide. The ovary lies
ventrally in the middle third and consists of separate aggregates ;
the horseshoe-shaped vitellarium lies near the dorsal side and
dorsad of it, the shell-gland.
_Eggs not present.
Davainea.
Scolex with hundreds of very small, generally hammer-shaped
hooks in two circles; suckers generally beset with hooks at the
margin; genital pores unilateral or irregularly alternating ;
numerous testes; eggs in capsules; parasitic in mammals and
birds.
45.—Davainea polycalcaria, n.sp.
Pl. IL., figs. 28-29.
Intestine of Corvus macrorhynchus, Tem. ; Colombo.
Length 65 mm.; in front the proglottids measure 0:03 long x
0°25 broad, in the middle 0°20 x 1°78, behind 0°99 x 1°34; they
become at the end slightly narrower and longer, but always
broader than long; the scolex is button-shaped and very short,
0°31 broad by 0°088 long ; the suckers, 0°10, carry several hundreds
of small hooks at the margin ; the likewise very numerous hooks
of the rostellum are closely packed in circlets, they are pointed
and measure (015. A transverse muscular layer marks off the
broad cortical zone from the medullary layer ; ectad of the trans-
verse muscles are numerous longitudinal muscles which do not
form bundles ; calcareous corpuscles are closely packed through-
out the body ; two large vessels run 3, of cross-diameter distant
from margin and are connected by a transverse ramus communi-
cans in the posterior part of the segment; close beside them
outside is the nerve. The pyriform cirrus-sac occupies +) of the
cross-diameter ; numerous testes occur in the medullary sub-
stance ; they are oval, about 0°039 by 0°026; female organs not
yet developed.
HELMINTHES : COLOMBO MUSEUM. 181
Diplochetos, n.gen.
Genital pores marginal bilateral ; genital glands in only one
group ;.at the scolex a double crown of hooks ; testes numerous ;
three layers of longitudinal muscles; segments much broader
than long; uterus with ventral orifice right and left of the
middle line.
46.—Diplochetos volvulus, n.sp.
Pl. IL., figs. 30-31.
Intestine of Yellow-wattled Lapwing, Sarciophorus malabari-
cus, Bodd.; Weligatta. aes
Length 8°5; anterior segments 0°018 long x 0-21 broad,
middle segments 0°10 x 0:40, posterior 0°37 x 0:70, always
broader than long. The pyriform scolex is 0°35 broad x 0:26
long; the rostellum carries twenty-four hooks in two circles of
twelve, the hooks 0:047 long ; in the parenchyme are three layers
of longitudinal muscle-bundles, which become stronger towards
the interior ; on each side two strongly sinuate vessels, the dorsal
larger ; the nerve runs along much nearer the margin ; calcareous
deposits not present. Genital pores marginal, right and left in
each proglottid; the cirrus-sac occupies } of the cross-diameter
and contains coils of the vas deferens; entad of the cirrus-sac
occur extensive convolutions of the vas deferens; the vagina
lies ventral to the cirrus-sac and expands into an irregularly
shaped receptaculum seminis which reaches about 4 of the cross-
diameter. In each transverse section about ten oval testes are to
be found. The ovary lies in the inner third of the medullary
substance and consists of separate follicles ; the round vitellarium
lies ventrally in the middle; the site of the uterus breaks
through the musculature and parenchyme ventrally about 4
of the cross-diameter from the margin, sometimes to the right,
sometimes to the left without opening to the exterior; I have
not found it in full development. The eggs are 0:016 long
x 0°013 broad. Other Tzeniid genera with biserial genital pores
are the following :— ;
Diploposthe, with simple hook-crown and three testes in each
segment.
Amabilia, with double cirrus-sac and simple vagina opening
ventrally.
Diplophalius, with double male organs and simple female
organs.
Dipylidium, with several rings of hooks on the rostellum and
elongate segments. ee
The large genera from ruminants and rodents do not concern
the question.
182 SPOLIA ZEYLANICA.
Ophryocotyle.
Scolex without rostellum, with five depressions, the margins of
which are beset with numerous small hooks; the margins of the
suckers are also armed with numerous small hooks ; genital pores
marginal, irregularly alternating or bilateral ; parasitic in birds.
47.—Ophryocotyle zeylanica, n.sp.
PL. IL., figs. 32-34.
Intestine of the Ceylonese Hornbill, Lophoceros gingalensis,
Shaw; Nedunkeni, Northern Province.
Length 55; anterior proglottids 0°035 long x 0:097 broad,
posterior 0°12 long x 0°70 broad. Scolex, 0°29 long x 0:19
broad, club-shaped, the so-called neck behind it, strongly
attenuate, 0°079 broad; no rostellum; the suckers occur
laterally behind the middle of the scolex; the inner ring
enclosing the lumen is beset with very numerous hooks ; at the
apex two parallel loops forming five finger-shaped rays whose
margins are beset with very numerous small thorn-like hooks
without root and lever, measuring 0°0053.
The cortical zone is very wide and separated from the
medullary zone by a transverse muscular layer, outside of which
are regularly placed large bundles of longitudinal muscles and
farther outside numerous, irregularly grouped, smaller bundles ;
two large vessels run ventral in the medullary layer forming a
posterior anastomosis in each segment ; the nerve occurs near the
transverse muscles. In the layer between the subcuticular cells
and the outer longitudinal muscles lie caleareous bodies. Genital
pores marginal bilateral at the anterior lateral margin of the seg-
ment ; cirri protruded, 0:19 long and 0°:035 broad at the base, they
are finely spinulose ; cirrus-sac carries outwardly longitudinal
muscles, thereunder circular muscles; it occupies almost } of
the cross-diameter ; numerous testes lie in the medullary substance,
about twelve appearing in a transverse section ; the vas deferens
is coiled and leads into a sigmoid seminal vesicle which has a
narrow lumen and a very thick hyaline wall.
The vagina courses ventral from the cirrus-sac and the
receptaculum seminis reaches almost to the middle of the
segment ; the ovary consists of large isolated cells and occupies a
large place in the medullary substance, not forming a closed
body ; the vitellarium abuts ventrally upon the transverse muscles
and is a roundish body of ,!, the size of the cross-diameter ; the
eggs are 0°036 by 0:029.
The genus Ophryocotyle hitherto comprised three species :—
O. proteus, Friis (1869), from Tringa, Callidris, Charadrius, and
HELMINTHES : COLOMBO MUSEUM. 183
Larus ; O. Lacazei, Villot (1875), from Limosa ; and O. insignis,
Lénnberg (1850), from Hematopus and Mergus.
Friis and Villot describe at the apex of the scolex five distinct
suckers in a row armed with hooks, whereas Lénnberg speaks of
undulating loops ; in O. proteus and O. insignis the genital pores
alternate irregularly ; the similarity in the formation of the
scolex seemed to require that the species here described should
be placed in the genus Ophryocotyle.
Brochocephalus, n.gen.
Scolex with long rostellum with six backwardly directed loops
of hooks; genital pores marginal, regularly alternating ; cirri
large, strongly and closely spinulose ; no veceptaculum seminis ;
three testes in each segment ; segments (proglottids) broader than
long ; eggs two-shelled, the inner shell narrowed at the poles,
48.—Brochocephalus paradoxus, n.sp.
Pl. II., figs. 36 and 38; Pl. IIL., figs. 35 and 37.
Duodenum of the Lesser Sand Plover, Agialitis mongolica,
Pall. ; Weligatta.
Length 85 mm.; without “neck ;” anterior proglottids 0-062
broad x 0-0088 long, posterior 0:75 broad x 0:18 long, always
much shorter than wide; scolex with four suckers and long
protruded knobbed rostellum, 0:10 long x 0-062 broad in front ;
at the apex six backwardly directed tracts of thirteen hooks each,
v.é.. seventy-eight hooks in all; the hooks are slender with very
small hook and lever, measuring 0:029. ;
Two layers of longitudinal muscle-bundles are seen in cross-
section under the cuticle ; on each side two vessels, the larger
forming a posterior anastomosis in each segment; the nerve runs
near the margin. Genital pores marginal and almost regularly
alternating, rarely two follow on the same side ; no calcareous de-
posits. Cirri very large, broad and spinulose, as long as half the
cross-diameter of the proglottid; the large muscular cirrus-sac
is expanded outwardly and oceupies about 4 of the ecross-diameter :
three small ovate testes in the middle line, one in front, the
others symmetrical behind it; behind the testes a transverse
vitellarium occupying } of the cross-diameter:; the Ovary sur-
rounds the testes in the middle third; vagina and cirrus-sac
lie between the two vessels ; the eggs are elliptical and two-
shelled ; outer membrane 0-081 x 0-047, oncosphere spherical ;
the inner thick shell is narrowed at the poles.
The genus Gyrocelia shows a zigzag line, interrupted at eight
points, with forty hooks on the rostellum; four testes in each
segment, and the spherical eggs have two appendices,
184 SPOLIA ZEYLANICA.
Oittoteenia.
Both the genital organs and their ducts are duplicated in each
segment; proglottids broader than long ; scolex unarmed ; uterus
transverse; eggs with pyriform apparatus ; parasitic in rodents.
49.—Cittotenia bursaria, n.sp.
P1.IL., figs. 39-40.
Intestine of Lepus nigricollis, Cuv. ; Nedunkeni.
No complete examples were present in the collection, the
largest fragments measuring 55 mm. in length. The body is
broad, rounded in front ; no ‘‘neck”; anterior segments 0°20 long
x 5 broad, posterior 1:34 long x 7 broad. The scolex is not
protruding ; the four suckers measure 0°14 in diameter ; rostellum
absent. In the parenchyme, two layers of longitudinal
muscles, transverse muscles and many dorsoventral muscles ;
on each side two longitudinal vessels, ventral larger, dorsal
smaller ; outside these, the nerve ; no calcareous bodies. In each
segment there are two groups of genital organs, each occupying
1 of the cross-diameter and having their several ducts The cirri
protrude behind the middle of the margin of the proglottid ; they
measure 0°44 long x 0°035 broad at the base ; the cirrus-sacs are
short and club-shaped; entad of them lies a fusiform bursa
ejaculatoria provided with longitudinal and circular muscles ;
entad of this a fusiform seminal vesicle and still further inwards
a convoluted vas deferens ; the very numerous spherical testes,
0:044 diameter, are distributed through the entire proglottids
without forming two lateral groups. ‘the vagina is broad, with
high endothelium, expanding toa large receplaculum seminis ; il
runs ventrad of the cirrus-sac: entad of the receptaculum and
ventral in position lies the roundish ovary surrounding the
vitellarium.
The uterus traverses the proglottid from right to left and has
roundish protuberances in front and behind ; eggs round, 0-078,
with triple membrane ; the inner membrane forms the so-called
pyriform apparatus ; the spherical oncosphere measures 0:014.
Ichthyotenia.
Secolex unarmed with four suckers, and often a fifth apical
sucker; genital pores marginal, irregularly alternating, testes
numerous; the vagina forms coils at the posterior margin of the
proglottids in the middle, which replace a receptaculum SEMINAS :
parasitic in fishes and reptiles.
HELMINTHES : COLOMBO MUSEUM. 185
50.—Ichthyotenia cryptobothriwm, n.sp.
Pl, IIl., figs. 41-42.
Intestine of Chrysopelea ornata, Russell, a tree-snake; Kuru-
negala.
Length 130 mm.; head end rounded, scolex retracted, the four
suckers being found only in transverse sections surrounded by a
parenchyme-ring with eight vessels ; the suckers extend 0°40 mm.
backwards ; their lumen communicates outwardly by two lacune,
a dorsal and a ventral, which ceases in the midst of the suckers ;
a median plug projects freely.
The anterior proglottids are very short, 0'02 long x 1:00 broad ;
the middle are 0°59 long x 1°69 broad; the posterior 2°48 long
x 0°87 broad, much longer than broad; the last proglottid is
rounded behind.
Caleareous bodies sparsely distributed ; two layers of longitu-
dinal muscle-bundles parallel with the cuticle traverse the
parenchyme, outer thinner numerous bundles, inner thicker and
sparser; at the margin on each side run two vessels, ventral
stronger, dorsal thin and highly sinuous; outside these the stout
nerve.
The genital pores are marginal and irregularly alternating in
the anterior third of the margin of the proglottid. The cirrus is
small, rod-shaped and plain; the cirrus-sac occupies } of the cross-
diameter ; the genital sinus is retracted ; the vas deferens forms
abundant coils reaching to the middle of the cross-diameter ; the
numerous testes lie in a transverse row in the dorsal half of the
medullary substance and measure 0°052 to 0:065. The vagina
runs ventral straight inwards; ventrad of the ovary it forms
numerous coils as equivalent of a receptaculum seminis.
The ovary consists of very large cells and occupies searcely } of
the cross-diameter ; dorsal lies the roundish vitellarium ventral
to which the shell-gland is ‘applied and ventral to this a haust-
orium (Schluckapparat). The spherical eggs have a double
membranous shell, measuring 0:047 to 0:052.
Aphanobothrium, n.gen.
Body broad and thick like a Schistocephalus larva, with short
posteriorly acute-angled proglottids; in the frontal region a
dorsoventral slit leading to five suckers ; destitute of scolex and
hovks; genital organs simple, ducts double, marginal ; the uterus
discharges ventral in the middle line; cortical layer broad ;
cirrus closely beset with hooks; on each side two vessels, the
larger with transverse anastomosis; ovary resolved into coils
which lie dorsoventral and transverse; belongs to the Anoplo-
cephalinz. .
186 SPOLIA ZEYLANICA.
51.—A phanobothrium catenatum, n.sp.
Pl, IL., figs. 43 and 46; Pl. IIL, figs. 44, 45, and 47.
Duodenum of the Flamingo, Phanicopterus roseus, Pall,; Weli-
gatta.
Length up to 135 mm.; breadth 9-10 mm.; body attenuated in
front, truncate behind and very thick; the dorsoventral diameter
isto the cross-diameter as9: 20; anterior end rounded and retracted
in the middle and confined by arching cross lines; scolex not
visible externally ; proglottids sharply delimited, at first 0:20,
then 0:27, behind 1:18long. The middle 2 of the body width are,
dorsally and ventrally, occupied by five longitudinal rows of flat
dells which are also disposed in transverse rows; posteriorly
these deepen into small grooves; the cortical layer is to the
medullary substance dorsoventral as 2: 5; in the tissue occur
oval calcareous bodies measuring on the average 0:018 x 0:012.
Under the cuticle are circular and longitudinal muscles; a strong
transverse muscular layer divides the cortex from the medullary
substance; in the cortex run radial bundles of longitudinal
muscles; on each side two vessels run close together, dorsal a
smaller thick-walled vessel, ventral a larger thin-walled; the
latter forms in each proglottid a strongly sinuous cross-
anastomosis; the longitudinal nerve-trunk runs close outside the
vessels.
The scolex is retractile; in transverse sections 0°05 to 0°07 mm.
from the frontal point are found four suckers of 0:10 diameter,
and a fifth in the middle; these are circular and open outwards
through a dorsoventral slit. The cirrus, 0°39 x 0:079 (0°12 broad
at the base), is regularly beset with hooks like the rostellum of an
Echinorhynchus, arranged in transverse rings of sixteen each ; it
protrudes from a hemispherical protuberance ; the cirrus-sac
occupies } of the cross-diameter; the numerous round testes of
0:26 diameter lie in the medullary substance approximated to the
transverse muscles.
The vagina runs dorsad of the cirrus-sac ; both lie between the ~
vessels. The ovary consists of strands extending from the dorsal
to the ventral side and then bending inwards where they unite in
a short transverse branch; dorsad of this lies the radiate shell-
gland, and dorsad of the latter the small vitellarium consisting of
nodular strands ; the ovarian cells are 0-013 large, those of the
vitellarium 0°0052; only the efferent duct of the uterus was
developed, perforating the transverse muscles ventral in the
middle line; eggs not yet present.
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HELMINTHES : COLOMBO MUSEUM. 18
EXPLANATION OF PLATES
Illustrating Dr. von Linstow’s Paper on Helminthes of Ceylon.
g. vessel ; ”. nerve ; ¢. cirrus-sac ; vd. vas deferens ; /. testis ;
v. vagina ; 7s. receptaculum seminis ; k. ovary ; d. vitellarium ;
u. uterus; s. male seminal vesicle ; sch. shell-gland; schl. haus-
torium.
PLATE I.
Fig. 1.—Ascaris fissicollis. Dorsal lip.
Fig. 2—Same. Accessory lip.
Fig. 3.—Ascaris coronata. Dorsal lip with accessory lips.
Fig. 4.—Physaloptera brevispiculum, 6. Tail from below.
Fig. 5.—Spiroptera secretoria. Head; f.free projecting secretory
filament.
Fig. 6.—Spiroptera orca, 6. ‘Tail from below.
Fig. 7.—Same, . Immature egg.
Fig. 8.—Same. Mature egg.
Fig. 9.—Heterakis pusilla, 6. Tail from below.
Fig. 10.—Strongylus digitatus, 6. Tail from below.
Fig. 11.—Ankylostomum minimum. Head from the right side.
Fig. 12.--Same, 6. Tail from the right side.
Fig. 13.—Trichocephalus discolor, 6. Tail from right side.
Fig. 14.—Same, 2. An egg.
Vig. 15.—Hchinorhynchus tener. Anterior hook of rostellum.
Fig. 16.—Same. Posterior hook.
Fig. 17.—Same. An egg.
Fig. 18.—Lyperosomum squamatum, from below.
Fig. 19.—Tristomum megacotyle, from below.
Fig. 20.—Same. Rod from the caudal sucker.
PLATES TT, AND III,
Fig. 21.—Hymenolepis septaria, Transverse section.
Fig. 22.—Hymenolepis clausa. Transverse section.
Fig. 23.—Same. Hook.
Fig. 24.—Hymenolepis spinosa. ‘Transverse section.
Fig. 25.—Same. Hook.
Fig. 26.—Diorchis occlusa. Transverse section.
Fig. 27.—Same. Hook.
Fig. 28.—Davainea polycalcaria. Transverse section.
Fig. 29.—Same. Hook.
Fig. 30.—Diplochetos volvulus. Transverse section ; on one
side the vas deferens is figured, on the other the vagina.
2¢ 8(17)05
188 SPOLIA ZEYLANICA.
Fig. 31.—Same. Hook.
Fig. 32.—Ophryocotyle zeylanica. Transverse section ; on the
one side the male organs are figured, on the other the female.
Fig. 33.—Same. Scolex from the frontal surface.
Fig. 34.—Same. Hook.
Fig. 35.—Brochocephalus paradoxus. Horizontal section; this
figure is reproduced on Plate III.
Fig. 36.—Same. Rostellum in side view.
Fig. 37._Same. Hook (on PI. III.).
Fig. 38.—Same. An egg.
Fig. 39.—(Pl. I1.). Cittotenia bursaria. Portion of a trans-
verse section ; s. 7. Bursa ejaculatoria ; s. 77. Seminal vesicle.
Fig. 40.—(PI1. II.). Same. An egg.
Fig. 41.—(P1. IIL.) Ichthyotenia cryptobothriwm. Transverse
section through the scolex, 0°30 mm. from the apical point.
Fig. 42.—(P1. II1.).. Same. Transverse section.
Fig. 43.—(P1. II.) Aphanobothrium catenatum. Head end in
flat view.
Fig. 44.—(Pl. III.). Same. Head end in frontal view.
Fig. 45.—(Pl. III.) Same. Transverse section through the
retracted suckers.
Fig. 46.—(PI1. I1.). Same. Surface view of posterior end.
Fig. 47.—(Pl. III.). Same. Portion of a transverse section.
N.B.—The remaining figures are on PI. III.
Fig. 48.—Ascaris brachycheilos. Dorsal lip.
Fig. 49.—Heterakis granulosa, 6. Tail from below.
Fig. 50.—Oxysoma falcatum. Head end.
Fig. 51.—Same, 6. Tail from right side.
Fig. 52.—Filaria digitata. Head end.
Fig. 53.—Same. Tail end of male from right side.
Fig. 54.—Same. Tail points of male from below.
Fig. 55.—Same. Tail points of female from below.
SPOLIA ZEYLANICA. 189
NEW AND INTERESTING LIZARDS IN COLOMBO MUSEUM.
By N. ANNANDALE, D.Sc.
(Deputy Superintendent of the Indian Museum.)
HANKS to Dr. A. Willey I have lately had an opportunity
of examining the Lizards in the collection of the
Colombo Museum, the specimens in which are, without exception,
from Ceylon. Notes on many of them have been published in
Mr. A. Haly’s “ Report on the Collection of Reptilia and Batrachia
in the Colombo Museum” (1891); but several additions have
been made more recently, while a re-examination of some speci-
mens has had interesting results, the most important of which is
the establishment of a new genus for the reception of Nevill’s
Euprepes halianus.* Another new Skink is also described.
GECKONID:.
GYMNODACTYLUS NEBULOSUS, Bedd.
The collection contains a half-grown specimen of this species
from a locality 18 miles north of Kandy (see A. Haly, Adminis-
tration Report, Colombo Museum, 1900). Boulenger records
another Ceylonese example in the British Museum; but the
species, common in some parts of Southern India, must be rare in
Ceylon.
GYMNODACTYLUS FRENATUS, Gthr.
An examination of the two males and two females in the
collection enables me to point out a peculiarity of the adult male
which is possibly assumed at the breeding season and is quite
absent in the female. The ventral surface of the base of the tail
is swollen in the former sex, and there are two large closely
adjacent papille close behind the opening of the penis on each
side,
*H. Nevill. Taprobanian, II., 1887, p.56 ; also Boulenger, Reptiles, Fauna Brit.
India, 1890, p. 213,
190 SPOLIA ZEYLANIGA.
AGAMID 4.
CALOTES OPHIOMACHUS, Merr.
Judging from a young specimen from Kandy in the collection
and from an adult lately captured by myself in Colombo, the
“two groups of spines on each side of the head” noted by
Boulenger may be united into a single series.
SCINCIDAL.
MABUIA BIBRONII (Gray).
Though Boulenger only records this species from the Carnatic
in the “ Fauna,” it is probably not uncommon in some parts of
Ceylon, whence there are several specimens in the Colombo
Museum which I have re-examined (see Haly’s Report on
Reptilia, &c., Colombo, 1891, p. 14, where the species is recorded
from Mullaittivu).
LYGOSOMA MEGALOPS, sp. nov.
Sub-genus Kenewxia, Gray (see G. A. Boulenger, Catalogue of
the Lizards in the British Museum, Second Edition, vol. IIL,
1887, pp. 210 and 214).
Habit lacertiform ; length from snout to fore-limb contained
about 14 times in the length from axilla to groin; limbs well
developed, pentadactyle, overlapping when adpressed; snout
short, obtusely pointed ; eye large; diameter of orbit as great as
length of snout ; distance from orbit to ear-opening much longer
than snout; ear-opening much smaller than eye, circular,
without denticulations.
Rostral much broader than deep, forming a straight suture
with the frontonasal ; no supranasals ; nasal undivided. Frontal
nearly as long as the frontoparietals and the interparietal
together ; interparietal completely separating the parietals ; no
distinct nuchals. Four large, subequal supraoculars ; seveu or
eight superciliaries; six upper and five lower labials. Dorsals
and laterals smooth, ventrals feebly keeled ; body scales subequal,
imbricate, in twenty-four to twenty-six rows round the body ;
anals and caudals not enlarged ; no enlarged scale on the heel ;
middle toe with twelve to fourteen subdigital plates. Colour
almost uniform dark brown. Length of head and body, 2
inches ; length of tail, 23 inches.
Localities.—One specimen from Puttalam ; another from
Kitulgala.
LIZARDS IN THE COLOMBO MUSEUM. 191
THECONYX, gen. nov,
Limbs well developed, pentadactyle ; claws retractile ; other
characters as in Lygosoma. This new genus is intermediate
between Lygosoma and Ristella, from the latter of which it differs
in having five digits on each foot and in the other points which
separate the former genus from the latter, except as regards the
claws.
Fig. 1.—Young Theconyx halianus showing general form and coloration,
from above, X 3.
192 : SPOLIA ZEYLANICA.
THECONYX HALIANUS (Nevill).
As the description of this species drawn up by Haly and
published by Nevill, who gave it its name (Huprepes halianus),
is not very clear, I have based the following diagnosis on three
specimens in the Colombo Museum.
Habit lacertiform, rather stout ; limbs approaching one another
or slightly overlapping when adpressed ; tail cylindrical, of the
same length as the head and body. Tympanum as smal) as
nostril, deeply sunk. No postnasal ; a narrow supranasal which
does not meet its fellow; rostral much broader than deep;
frontal three times as long as broad, much longer than inter-
parietal, from which it is completely separated: four large
supraoculars, seven or eight superciliaries ; no distinct nuchals.
Lower eyelid scaly ; body scales large, imbricate ; dorsals with
three or five indistinct keels; laterals smooth ; twenty -four scale
round the body ; anals slightly enlarged.
Fig. 2.—Left hand from below. Fig. 3—Left foot from below.
x about 4. x about 4.
SEES
Fig. 4.—Claw. xX 12.
Coloration.—Dorsal surface olive (yellow in young) with six or
seven dark transverse bars on the body which are narrower than
the interspaces, and eight or nine on the tail, These are much
more conspicuous in the young than in the adult and equal to
the interspaces. Head variously marked with olive and dark
brown. Ventral surface dirty yellow. Length of head and
body 13 inch : length of tail 1} inch.
Localities.—In addition to Nevill’s types (an adult from the
Western Province and a young specimen from Anuradhapura)
the Colombo Museum possesses a third example (half-grown)
from Horana, collected and presented by G. H. Swayne, Esq., 8th
November, 1901. It is this specimen I have measured, as the
tail is injured in the adult, which is at least twice as large.
SPOLIA ZEYLANIGA. 193
STALKED BARNACLES (CIRRIPEDIA PEDUNCULATA) IN
THE COLOMBO MUSEUM.
By N. ANNANDALE, D.Sc.
(Deputy Superintendent of the Indian Museum.)
MONG the Cirripedes in the Colombo Museum I have been
able to identify the following :—(1) Lepas anserifera, Linn.,
the commonest pedunculate form on floating objects in this part of
the Indian Ocean ; (2) Dichelaspis pellucida, Darwin, a somewhat
scarce species only taken on sea-snakes; and (3) Dichelaspis
equina, Lanchester, which was not described until about three years
ago [Lanchester, P. Zool. Soc., London, 1902 (2), p. 375] but appears
to be common on shallow-water crabs of the east coast of India
as wellas in some parts of Malaya.
The specimens of L. anserifera are attached in dense masses to
pieces of wood and to a bottle ; those of D. pellucida are scattered
on the body of a sea-snake (Hydrus platurus) ; while D. equina
is represented by numerous individuals crowded together on the
posterior walking legs and carapace of a Dorippe dorsipes (Linn.),
and by others scattered on the dorsal and ventral surfaces and
mouth parts of Scylla serrata (Forsk.).
The distribution of Z. anserifera is world-wide. So far as I
am aware, neither species of Dichelaspis has been reported
hitherto from the immediate neighbourhood of Ceylon. D.equina
was described from the east coast of the Malay Peninsula, while
D., pellucida is an Oriental species which probably has a fairly
extensive distribution.
DICHELASPIS TENUIVALVATA, Sp. nov.
Diagnosis.
Capitulum compressed ; carinal edge rounded ; oceludent edge
sinuous, slanting outwards from above; lower edge straight, }
horizontal ; opening large. Five imperfectly calcified plates ;
carina not reiching the upper edge of the capitulum above,
194 SPOLIA ZEYLANICA,
furcate below, the two rami extending along the base of the
capitulum ; tergum large, triangular, almost entirely covering the
upper part of the membrane between the carina and the opening ;
scutum large but very imperfectly developed inferiorly (the
lower border being quite indistinct), in contact with the tergum
along the tergal margin above, feebly separated from the carina
behind, not cleft. Peduncle very short, transversely wrinkled.
Mouth-parts well developed; the mandible very large, with five
teeth ; the labrum feebly bullate ; the maxilla furnished with
stout bristles along its free edge, the inner half of which is
almost straight, while the outer half is deeply concave,
s,
*
we,
-* Scales of Sake
D. tenuivalvata Annand,. x 30. The specimen has been removed from the
body of the snake to which it was attached, some of the scales of the snake still
adhering to the short stalk of the barnacle.
STALKED BARNACLES IN THE COLOMBO MUSEUM. 195
Measurements.
Length of capitulum ... «. OS DTM.
Breadth of capitulum ... .. 4 mm.
Length of peduncle Fe aia, Set Ot TENNIS
This interesting species is represented by several specimens,
growing, side by side with D. eqguina, on the ventral surface of a
sea-snake (Hydrus platurus) from the coast of Ceylon.
All the members of the genus Dichelaspis are notable for the
reduction of the capitular plates which they exhibit. In D.
tenuivalvata, however, the degeneracy is of structure rather than
of form. The five plates represented are all large, but their
calcification is so imperfect that although the mantle is very
transparent, I was unable to see their limits without staining the
specimens. Hven when thus prepared they were by no means
easy to examine critically, and failed entirely to discern the
lower termination of the scuta. On the whole, the affinities
of the new form would seem to lie with Lanchester’s D. occlusa
(P. Z. S., 1902 (2) ) from the east coast of the Malay Peninsula.
From this species it may be distinguished by its extremely short
peduncle and imperfectly differentiated but undivided scuta.
2D 8(17)05
196 SPOLIA ZEYLANICA,
NOTES.
1. Lacteal Tract of Oriental Lorisine.—Dr. Nelson Annandale
Deputy Superintendent of the Indian Museum, writes from
Calcutta under date September 25, 1905 :—
“Tn connection with what you told me regarding the mamme
of Loris gracilis* I have examined the only two specimens we
have in spirit in the Indian Museum of Nycticebus tardigradus,
viz., (1) a male foetus 55 mm. long, and (2) a large female. The
first is quite hairless, and I had no difficulty in detecting two
pairs of mamme as minute tubercles on its skin. The first lies
in a line with the centre of the upper arm when the limb is
pressed down along the side ; the second only a short distance
above the umbilicus, but in the same position relative to the
sides of the body as the first.
* A somewhat prolonged search among the hair of the large
female, which was not in lactation, also resulted in the discovery
of four mamme in the same position. One pair therefore may
be said to be pectoral and one abdominal, though there is not any
very great separation between the two pairs.”
N. ANNANDALE.
2. Curious action of a Toad when confronted by a Snake,—lI
have in my vivarium a “striped ground snake” (Z'vopidonotus
stolatus, L.) which feeds exclusively upon toads and is always
ready for a meal. Under ordinary circumstances it seizes its
prey before the latter is aware of its presence. But, if the toad
catches sight of the snake in time, it can always (temporarily)
avert its doom by assuming a very characteristic attitude that
seems to completely disconcert its would-be captor. The toad
faces its enemy, depresses its head till the snout almost touches
the ground, erects itself on tip-toe, in which position—owing to
the superior length of the hind limbs—the rump is elevated.; at
the same time inflating its abdomen to its utmost extent. If the
snake does not immediately move off (which it usually does),
the toad alternately lowers and raises the hinder part of the
body, a movement which finally routs the enemy. This curious
action is taken by even quite small toadsand cannot be the result
of individual experience. It would appear to be an inherited
instinct.
E. ERNEST GREEN.
Peradeniya, October 26, 1905. ;
* Willey in Spolia Zeylanica, part X., 1905.
NOTES. -- “797
-3. On the constricting habit of Coluber helena.—With reference
to my note (vol. IIL, part X., p. 157) on the constricting habit of
Coluber helena, I have since been able to prove the correctness of
my supposition that this action is normally employed in the
capture of its prey. I have several times observed its capture of
a small skink (Lygosoma, sp.). On one occasion the Coluber had
captured a lizard and was tightly constricting it, the whole body of
the snake being twisted into a complicated knot. It commenced
work upon the tail, which became detached from the body of the
lizard and was promptly swallowed. The snake then apparently
forgot that the largest part of its captive was still enclosed in the
folds of its body, and began looking about for another victim.
Eventually it re-discovered its original capture and commenced
toengulph it, drawing it gradually through the encompassing coils.
In hunting the lizardsit seems to be very inexpert, and repeatedly
failed to effect a capture. As soon as the lizard stopped the snake
lost interest in it and was attracted only by moving objects.
EK. ERNEST GREEN.
4. Scorpion stings.—The sting of the small gray scorpions
(Lsometrus, spp.) is popularly reputed to be more virulent than
that of the large black—or rather, dark blue-green—scorpion
(Palamneus, sp,). If thisis really the case the sting of the latter
must be almost negligible. I have been recently stung on the
palm of my hand by afull-grown male Jsometrus europeus. The
pain was by no means intense—not nearly as severe as that
caused by the sting of a bee-or wasp—and completely passed off
within a very few minutes, without the application of any
remedy, leaving no trace or mark of any kind. It is possible that
the poison spine could not fully penetrate the thick skin, though
the puncture was in the median softer area of the palm. Butl
have observed a case in which a child of three years old was
stung on the finger. The skin of a child at such a tender age
cannot afford any serious obstacle to penetration. In this case
also the pain was very transient. Though productive of
momentary tears, within ten minutes the incident had been quite
forgotten. I have been assured by other persons that no seriously
unpleasant effects have followed the sting of Palamneus.
Are these cases exceptional, or is the popular apprehension of
scorpion stings greatly exaggerated ? It would be interesting to
collect evidence on this question. Will readers of this Journal,
who may have had personal experience in the matter, comeforward
with particulars of their sensations under such circumstances ?
EK. ERNEST GREEN.
198 SPOLIA ZEYLANICA.
5. Minerals new or rare in Ceylon.—Of the following
minerals, some were omitted from the list of Ceylon minerals
given in Spolia Zeylanica, part IX., 1905; the remainder have
been discovered since.
A.—Minerals new to Ceylon.
Galena.—Occurs in small cubes and rolled fragments in nambu
from the Getaheta-oya at Murutangala in Getaheta. Reported
from other localities.
Cassiterite.—Occurs as small black pebbles in nambu at
Niriella and Noragala, and in larger fragments at Kuruwita.
Selenite-—Massive at Kukulawalakanda, Ratmale, Pasdun
Korale, intergrown with pyrite between veins of graphite [James
Parsons | ; also in crystals in fossiliferous marine clays, Matti-aru,
Kuchaveli, north of Tricomalee.
Monazite.-—Occurs in waxy yellow well-worn grains in sands
of the Getaheta-oya and other rivers near Avisawella; but was
first detected by Dr. J. W. Evans in specimens of heavy sands from
the Niriella-ganga sent to the Imperial Institute by the Mineral-
ogical Survey.
Orangite.—It has been pointed out that the “thorite” of
Ceylon really belongs to the variety known as orangite, dis-
tinguished by slightly greater specific gravity and orange colour,
The apparently orthorhombic form of the mineral however
remains unexplained. The crystals are not good enough to give
satisfactory measurements, and too much altered internally for
optical examination.
Platinum.—Occurs very sparingly, associated with gold in
river sands at Dombagomuwa and Karawita, Sabaragamuwa.
[James Parsons].
The following minerals have been reported, but on present
evidence cannot be accepted as found in Ceylon. The sulphides
are quite likely to be found:—Cinnabar, Stibnite, Realgar ;
Epsomite, Olivine [see Bertha Vukits Centralblatt fiir. Min. ete.,
1904, No. 23, p. 715]. Minerals sent by the Mineralogical Survey
to the Imperial Institute were provisionally identified as
Annerodite and Adschynite or Polycrase.
B.—Mineral names omitted in the List.
Molybdenite.—Recorded by Dr. Gygax, andin Gustav Leonhard’s
“ Handworterbuch der topographischen Mineralogie,” 1843. I
re-discovered this mineral this year in the Kegalla District,
where it occurs [at Hettimulla| in a vein of pegmatite cutting the
granulites.
NOTES. 199
Wolframite is also recorded by Leonhard, as well as Saltpetre
and Rose quartz, both of which latter are certainly found in
Ceylon.
“ Oriental Emerald ” ismentioned by Max Bauer in his “ Edels-
einkunde,” which is however totally unreliable as regards Ceylon.
Greenish sapphires do occur, but rarely, and hardly deserve the
name of Oriental Emerald. N.B.—Max Bauer states erroneously
that Beryl does not occur in Ceylon.
Beccarite.—A biaxial variety of Zircon from Ceylon.
Andesine.—(Des cloiseaux, 1884).
Borax.—(Mentioned by Dana).
I am indebted to Mr. L. J. Spencer for several of these refer-
ences.
C.—Some early Records.
Nicolo Conti, who travelled in the East between 1419 and 1444,
speaks of a ‘‘ very noble island, called Zeilan, in which they find
by digging, rubies, saffares, garnets, and those stones which are
called cat’s-eyes.”
Albanasius Nilikin, who travelled between 1468 and 1474,
speaks thus of Ceylon: ‘‘ Ceylon is another not inconsiderable
port of the Indian sea. There, ona hill,isthe tomb of Adam, and
in the vicinity are found precious stones, antimony, fastisses,
agate, cinchai, crystal, sumbada.” A little after he says: “ At
Ceylon you findammone, antimony, fatisses.” Of these, antimony
is not known to occur in Ceylon; cinchai doubtless refers to
spinel (kivinchi). Agate ofa sort does occur sparingly in Ceylon,
but itis more likely that agate of Indian origin is here referred to.
Ludovico de Varthema, who travelled from 1503 to 1508,
speaks of the rubies, garnets, sapphires, jacinths, and topazes of
Ceylon.
These references are extracted from Sir George Birdwood’s
* Report on the Old Records of the India Office,” London, 1891.
A. K. COOMARASWAMY.
6. Recent Marine Clays at Kuchaveli, Ceylon: Ananda K.
Coomaraswamy.
Note on some Post Tertiary Mollusca from Ceylon: R. Bullen
Newton.
Geol. Mag., Dec. v. vol. II., No. XI., November, 1905,
A recent marine clay is exposed near Nachchiarmalai on the
Matti-aru (near Kuchaveli, 22 miles north of Trincomalee) about
200 SPOLIA ZBYLANICA.
14 mile from the coast, at about sea level. The section shows
about 9 feet of alluvium resting on the clay bed which is about
21 feet thick, the lower part being under water. The bed is full
of marine shells, and contains also concretionary nodules with
shells, serpule, and crab remains. All these also occur commonly
washed out and scattered over the sand banks of the Matti-aru.
The common crab, used medicinally, is Macropthalmus Latreiller
(Desmarest), the “ medicine” crab of the Chinese pharma-
copoeia; it has been found in the Post-tertiary clays of Southern
China. A single chelate pincer belonged to Scylla serrata (De
Haan), the great Indian swimming crab. The following shells
together with annelid tubes, Balani, fossil wood, and the crabs
complete the list of organisms collected :—
Telescopium telescopium (Linnzeus).
Potamides fluviatilis (Potiez and Michaud).
Purpura carinifer'a (Lamarck).
Nassa ornata (Kiener).
Melongena pugilina (Bour).
Natria, sp.
Scapharca rhombea (Bour).
Arca, sp.
Placuna placenta (Linneus). This is the ‘* window-pane
oyster,” and it is exceedingly abundant in the bed.
Tapes undulata (Bour).
Tapes textrix (Chemintz).
Diplodonta, cf. oblonga (Hanley).
Tellina, sp.
Dosinia salebrosa (Roemer).
Chione allied ¢mbricata (Sowerby).
Three points in Mr. Bullen Newton’s paper must be
noticed :—
(1) As stated in my paper, nodules occur in the clay bed,
and one crab-nodule was found im sitw; there is
therefore no foundation for his suggestion that the
fossils in the nodules may be younger than those in
the clay.
(2) It does not appear how the identity of littoral fauna
can prove the recent connection of Ceylon with
India.
(3) The particular specimens examined were not presented
as stated,
ANANDA K, COOMARASWAMY.
NOTES, 20]
7. “ Contributions to the Geology of Ceylon. 4. Intrusive
Y y
Pyroxenites, Mica-pyroxenites, and Mica-rocks in the Charnockite
Series or Granulites in Ceylon”: Ananda K.Coomaraswamy
Geol. Mag., Dec. v., vol. II., p. 363, August, 1905.
The following is a summary of the observations recorded in
this paper. A group of pyroxenites in which the minerals
diopside, phlogopite, horneblende, and scapolite predominate,
while sphene, plagioclase, pyrite, apatite, calcite, and spinel are
often accessory, occurs in small sills and dykes intrusive in the
charnockite series or granulites of Ceylon, and in one case
apparently also in a zircon granite of the Balangoda group. The
pyroxenic intrusions have usually a zoned or laterally symmetri-
cal structure analogous to that of mineral veins. The intrusions
rarely exceed 6 feet in thickness, and are usually smaller ; they
are widely distributed in Ceylon. The intrusive material never
shows a chilled edge, but there is a rapid transition from the
pyroxenite to the granulite. The course of the small sills and
dykes of pyroxenic material is usually determined by the
foliation planes and joints of the granulites.
ANANDA K, COOMARASWAMY.
8. Snake lore—Some extraordinary ideas about the cobra are
entertained by the Sinhalese—at least in some parts of the
Island, as will be gathered from the following :—
(1) The cobra drops a segment of its body —beginning of course
at the tail end—after every biteit inflicts, and is ultimately
reduced to a mere head or “ hood,” when it is known as “ Kobo-
nayi,” in which form it is capable of great activity and is able
to perch, bird-like, on the branches of trees.
(2) The cobra grows shorter and thicker as it grows older, and
finally attains to gigantic proportions, “as thick as the stem of a
full-grown arecanut palm.” An ancient reptile of great age and
size is believed to be in occupation of the cave to be found in
Elakande, Horana, where a princess of the Sinhalese dynasty,
who was afflicted with an incurable disease, is said to have lived
in retirement. The trail of this cobra is described as like that
of a log drawn over the surface of the ground.”
(3) When honey bees work for many years in the same spot
they build in the centre of their hive a dome-shaped comb
resembling, in shape, the ash-pumpkin, and hence referred to as
“puhul.” ‘To such hives the cobra finds its way, and coiling
202 SPOLIA ZAEYLANICA.
itself round the “pumpkin” feasts upon bee larve and honey.
The colour of the reptile found in such situations is reddish-
brown, and its sting is innocuous.
C. DRIEBERG.
9. Snakes and fowls—While sitting on the verandah of
Gokarella resthouse, some 12 miles from Kurunegala, I observed
a pullet pursuing a snake 12 to 15 inches in length. At intervals
the latter turned upon its antagonist, and attempted to entwine
itself round the neck of the bird, which, however, snatched it
away without much difficulty with its claws, and proceeded to
peck at its head, when the snake would again try to get away.
In the end, when the snake was about half dead, the fowl
started to swallow it, taking the head first, and after persistent
efforts the reptile, wriggling to the last, disappeared down the
throat of the bird.
To me this was a novel experience, but I subsequently learned
from the resthouse-keeper and others that it was a common
enough occurrence in the countryside, and that village poultry as
a rule attack and make a meal of such snakes as Haldanda,*
Aharakuka,f and Khetuwa.t
C. DRIEBERG.
10. Moths at sea.—During the forenoon of November 18, 1905,
when about sixty miles from the coast of Ceylon, several specimens
of Ophideres fullonica were found on board H.M.S. Sealark and
were brought to me. The noon position of the ship was 6° 115’
N., 79° OL’ E., Colombo Clock Tower bearing N.49 E. 67 miles ; so
that the nearest land was the coast about Barberyn. The wind was
northerly and light (force about 2). Itseems probable that these
specimens came off with the land breeze of the preceding night,
but the distance from land which they had attained appears
noteworthy.
Several specimens of Cephonodes hylas were also flying about
the ship at the same time, but this species is a well-known
migrant,
THOS. BAINBRIGGE FLETCHER,
H.M.S. Sealark.
December 4, 1905.
* Dendrophis pictus. + Tropidonotus stolatus. { Dryophis mycterizans.
SPOLIA ZEYLANICA. 203
TERMITE TRUFFLES. *
By Dr, F. DOFLEIN (Munich).
MONG social insects the Termites [or “ white ants” ] have
been the object of the least amount of biological investiga-
tion. What has become known concerning their life and political
organization is to be found scattered chiefly in systematic works.
So it happened that one of the most interesting discoveries
connected with Termite life was hardly recognized in Science
although the analogous conditions among the ants had awakened
the highest regard.
The faculty possessed by certain Termites of cultivating fungi
in their hillocks has been remarked and more or less clearly
portrayed by a succession of naturalists from the time of
Smeathman, 1781. I will not review the literature here as I shall
do this in a more detailed work. It may be mentioned that
hitherto no one has investigated the phenomenon in all its aspects
and that consequently, in spite of its importance, it has hardly
been touched upon in general biological literature.
Thus I was completely taken by surprise as I opened a Termite
hillock in Ceylon not far from Colombo in order to procure well-
preserved material of Termite brains which I had promised to a
colleague. The nest had the form of a high arched dome,
terminating above in two chimney-like funnels. The height
amounted to 1-13 metre; the funnels had a length of about 20-30
centimetres.
Upon opening the hillock it was at once evident that its thick,
solid clay mass was traversed by a large number of chambers.
Hach of these chambers had approximately the size of a cocoanut.
The walls were smooth and several or many narrow passages
connected each chamber with the neighbouring compartments,
Every chamber was filled with peculiar formations, namely,
brown cakes of a moist, friable substance which in form, size, and
structure strongly resembled middle-sized bath-sponges. One
* Translation of an article by Dr. Doflein entitled “ Die Pilzkulturen der
Termiten,” published in the Verhandlungen der Deutschen Zoologischen
Gesellschaft (xv., Breslau, 1905), Leipzig, 1905, pp. 140-149, two text-figures.
25 10-06
204 SPOLIA ZEYLANIOA.
or several of such cakes were present in each chamber; when
several occurred they were piled one over the other like dishes.
These cakes were occupied by thousands of Termites. The
plan of composition of the “cake” consisted of a multitude of
small cells each of which had its numerous inhabitants ; especially
the passages between the cells seethed with larves and nymphs of
all stages.
The framework of the cake was furthermore beset with rela-
tively numerous white nodules, about the size of a pin’s head,
i-2 mm. diameter, which at first I took to be Termite eggs.
Microscope examination showed me however that they were
structures of quite a different nature.
One could perceive with the unaided eye that the entire “ cake”
was covered, inside and outside, with a fine felt of fungus hyphe
These associated themselves in masses at certain places and as
theirterminal parts branched and expanded ina particular manner
globular aggregates of club-shaped bodies—in other words, the
white nodules, which I will henceforth designate mycelial nodules—
were produced. At present I will not discuss the botanical side
of the question.
The resemblance to the growths observed by Modller in the
fungus gardens of ants of the genus Aéta in Brazil, immediately
occurred to me. As a matter of fact the mycelial nodules serve
the same purpose to the Termites that Moller’s nodules do to the
South American ants. At the same time there are certain bio-
logical peculiarities to be mentioned which are of interest and
have not hitherto been observed.
I was able to prove by different ways that the mycelial nodules
are eaten. In the first place I opened the intestine of numerous
individuals and found them in the crops of all the larve and
nymphs which I examined. Indeed the crops were completely
filled with them, and nothing besides. The cells of the mycelial
nodules were all quite uninjured. The foregoing applies alike to
the larve of workers and soldiers and to the larve and nymphs
of the sexual individuals.
IT also succeeded in aetually feeding the larvee of the workers
and soldiers as well as the larve and nymphs of the sexual
insects with the mycelial nodules. By offering a single nodule
on the point of a needle to animals which had hungered for some
hours or for a day, they accepted the proffered diet. It was
interesting to observe how they first of all investigated it with
their palps, then took it between the mouth-parts, and slowly
turned it round for a long time working it with the points of the
mandibles. It is not easy to observe these operations because the
TERMITE TRUFFLES, 205
insects are readily disturbed by the breath, and it is necessary to
watch them through a fairly strong lens. It is remarkable that a
nodule exactly fills the space between the mouth-parts when
these are fully open. Even the queen accepted a nodule and
consumed it in the same way as the others.
On the contrary, I have never succeeded in inducing an adult
soldier or worker to accept a nodule. Indeed I found their
stomachs to be always filled with vegetable detritus, consist-
ing exclusively of finely divided particles of wood. In spite
of careful examination I could detect no trace of mycelial nodules.
My experiments are not sufficiently numerous to permit a final
opinion as to the mode of feeding of the Termite species investi-
gated by me. Meanwhile it seems reasonable to suppose that
with this species the larvae receive a concentrated and easily
digested food in the form of the mycelial nodules and that these
constitute the permanent food of the sexual forms, while the
larve of workers and soldiers after a certain age obtain other
fodder. Through this consideration the further inference is
suggested that the fodder plays an important part in the differ-
entiation of the “ cells” in the community of Termes obscuriceps,
Wasm.
I may here add some remarks upon the structure of the nest,
the material of the fungus plantations and the bringing of the
latter within the Termite enclosure, which I have investigated
partly in Colombo, partly in Peradeniya and in the Northern
Province.
It has been mentioned above that the dome-shaped nest with
its chimney-like terminals consists of a clayey substance. In its
construction the Termites employ earth, sand, clay from the
neighbourhood, and mix these materials together with their
glutinous saliva. They eject this saliva upon the human skin
when they bite, whereby viscid, brown, tenacious spots arise.
The bite is hardly painful.
For the purpose of observation I placed portions of a nest into
a large glass receptacle and covered the entire mass which com-
prised fragments of the wall, fungus cakes, and thousands of
Termites, with a bell-jarin sucha way to allow access of air. The
whole apparatus was exposed to diffuse daylight. It is well-
known that the majority of Termite species, especially the blind
workers, avoid the light. They immediately commenced in a
methodical manner to construct a roof over the fungus plantations,
using the débris of the nest for this purpose, notwithstanding
the fact that the royal cell with king and queen had been taken
away from them. Within a few hours they had roofed over the
206 SPOLIA ZEYLANICA.
space of a square foot by unceasingly applying small clumps of
masonry mixed with their saliva.
An interesting observation could now be made. Owing to the
fact that evaporation was greatly reduced under the bell-jar, the
newly built roof remained soft and unstable and retained this
consistency for several days until I exposed it to the free air,
when it completely hardened after a few hours.
The mixing of the building material with the saliva of the
Termites not only ensures a very firm construction, but makes
the walls in a high degree resistent towards wetting. The Ter-
mite buildings brought by me to Munich still show this property
with great clearness. When I tried to saturate one of the nests
with lime water in order to strengthen it, it could not be wetted,
whereas this method was constantly employed with advantage to
the nests of Kuropean Hymenoptera, &c. The Termite nest
showed an equal resistance against being wetted with alcoholic
solution of shellac.
From this observation something may be inferred which other-
wise the inspection of the internal disposition of the Termite nest
reveals. For this purpose I will first describe the fungus cakes
in regard to their structure and in their relations to the nest.
Upon opening a Termite nest one finds that the substance of the
‘truffle ”’* is friable and soft. Only when this is the case are the
recesses of the truffle covered with mycelial nodules and the
entire structure populated by numerous Termites with their
larve. Sometimes I came across places in the nests where the
truffles were hard and dry; then they were destitute of nodules
and were not inhabited, except for afew scattered workers in the
cells. Fresh “cakes” dry rather readily when exposed to a current
of air. It is thus very easy to prepare them for transport, and they
have frequently been received into zoological collections, although
their actual nature has not been recognized. They become as
hard as wood, but always remain brittle and very fragile owing to
their delicate composition.
Microscopic examination of their substance shows that the fine
brown scaffolding of which they are composed consists exclusive-
ly of finely chewed wood. Thus the great wood-hunger of the
Termites and the cause of their extraordinary destructiveness
became clear tome. Just as the species of Atéa in South America
occasion great damage to the leaves of living plants in that they
employ their substance for their fungus plantations, so the
* The term truffle is used in the translation as an alternative term for fungus
cake.
TERMITE TRUFFLES. 207
Termites are the destroyers of wood. It appears that only a few
species attack living wood ; most kinds content themselves with
dead wood or such as has been in any way already damaged
by fungi. In the open therefore they are beneficial rather than
injurious, since they act as scavengers of rotten wood. But when
they come into contact with man they become some of the most
injurious insects of the tropics, since woodwork which is adapted
for their purposes forms an tenor constituent of human
dwellings and furniture.
The wood is bitten up quite fine, in the substance of the
fungus cakes and in the intestinal contents of the workers
one finds the finest vessels of the wood isolated. This wood-
brew is discharged through the vent as a small clump which
is evidently mixed with saliva and used in building the fungus
cakes. It is interesting to note that in other cases where Termites
build their dwelling or the main part of it out of dung, the same
fundamental arrangement of walls and passages recurs, as here,
where they build for the purpose of the fungus cultivation.
Ifa fungus cake from a Termite nest be exposed to the light
under a bell-jar to protect it from draught and evaporation, the
fructification of the Termite fungus can be easily induced, a
property which distinguishes it markedly from the Rhozites
forms cultivated by the Attia species. Aftera few days numerous
long club-shaped fructifications grow forth from the thick felt of
hyphe which has meanwhile developed. I will say no more
about the species and form of the fungus in this place as my
studies are not yet completed. I need only point out that other
species of fungi appeared very gradually on the cake, whereas
other objects in the vicinity were subject to a daily coating
of mould. The tendency of the fungus cultures of the Termites
to grow in pure culture appears to be very strong, even when the
cake contains scarcely any Termites. The purity of the culture
cannot therefore be placed to the account of the tireless weeding
by the workers, as is.done by Moller for the fungus plantations
of Atta. At the conclusion I will revert to this point once more.
When a Termite truffle is kept under a bell-jar with moderate
access of air, in very short time the inner surface of the jar is
covered with water drops which after some hours run in small
streams down the glass and form small accumulations of water
in which the Termites can be drowned. The fungus cake
therefore gives off a quantity of water by evaporation. Under
such great moisture the fructification of the fungus takes place
and the formation of mycelial nodules is stopped and the entire
structure tends in high degree to mouldiness.
208 SPOLIA ZEYLANICA.
There is a further point worthy of note in the cultivation
under the bell-jar. After a short time, 1-2 days, one sees the
Termites in large numbers lying upon their backs and realizes
that they are suffocated. Termites taken from the same culture
and kept between two hermetically closing watch-glasses without
pieces of cake, which were kept during the same time, lived and
were perfectly active. It follows from this that the suffocation
of the former was not due to lack of air, especially since access
of air to the main culture was not entirely prevented.
Upon raising the bell-jar it was noticeable that the space was
filled with a gas mixture which was clearly very different in its
composition from the atmospheric air. In breathing it one
experienced a strong oppression, as well as a very remarkable
odour like that of the gas proceeding from fermenting substances.
In any case a large amount of carbonic acid gas had formed, it
not other gases, perhaps as a collateral result of the growth of
the fungus at the expense of the wood.
All those appearances which are observed under the special
conditions of culture outside the Termite nest are naturally
absent inside the hillock. There, through the special construction
of the nest, it is arranged that necessary temperature and
moisture for the development of the fungus remains constant.
The building material is up to a certain point waterproof, so that
neither an excessive evaporation from the fungus cake through the
wall of the chamber, nor the entrance of rain from without, ensues.
What however is most important is that the entire method of
construction of the hillock ensures the elaborate ventilation of
its inner spaces; the Termite hillocks with their chimney flues
are hygienic dwellings.
The chimneys are air shafts which conduct away moisture,
carbonic acid gas, and other injurious gases, while fresh air can
enter through the lower openings. These ventilation flues can
at any time according to the increase of the stock, climatic varia-
tions, &¢., be altered and adapted. Thus is explained the quite
different number of chimneys which arise from a Termite hillock,
and it would be of interest to undertake a comparative study of
the size and number of chimneys, size of hillock, number of
fungus chambers, climate and locality, building material, popula-
tion, andsystematic position of the Termite species under question.
That the draught in these chimneys can, under circumstances,
be considerable, is indicated by the application which travellers
have frequently found forthe Termite nests. Quite recently it has
been repeatedly reported that our troops in South-West Africa
have used the Termite nests as ovens, selecting those forms
TERMITE TRUFFLES. 209
in which chimneys open into a great central dome chamber. In
the case of the form investigated by me in Ceylon I found the
chimneys communicating directly with the fungus chambers,
although the design is generally such that rain cannot directly
reach the fungus-cultures. Whether there is any arrangement
for the disposal of water I was unable to ascertain.
In conclusion, I should like to discuss one point briefly. It is
a remarkable fact that we find the cultivation of fungi practised
by ants and termites: that is to say, by the two orders of social
insects which are farthest removed from one another in point of
affinity. How is such a wonderful convergence to be explained ?
In the history of the social insects there are many phenomena
whose first appearance will always remain inexplicable. More
interesting and important is the question whether we can in any
way conceive and analyse the phenomenon. At first sight the
convergence as between Termites and Ants is quite wonderful
and could lead to the rashest hypotheses, like the convergence
between the ant species Oecophylla smaragdina and Campanotus
senex, both of which use their larve as spinning machines in
weaving their nests. Is their any means of tracing the causal
relations in the origin of the fungus cultures of the Termites and
eventually of investigating them experimentally ?
I think that the biology of Termites gives us points which will
_ support a hypothesis, and I very much regret that the thought
did not occur to me while I was in Ceylon so that I might under-
take the not very difficult testing of its principles. My suspicion
is that the fangus which grows in the fungus cakes is a common
fungus occurring in rotten wood in the neighbourhood, and that
it is conveyed with the chewed wood quite unconsciously by the
insects into the nest. The fact that it grows there almost
in pure culture could be explained by supposing that the wood-
brew is partially sterilized by the action of the saliva or gastric
juice, so that the chewed wood becomes a medium for the
exclusive growth of the particular fungus, all others being
suppressed. Thus the structures automatically become fungus
gardens (although, as in many other cases, originally intended
only as passages and resting places for the larvae, &c.), which are -
then methodically utilized by the Termites. It is not excluded
from possibility that the inhabitants acquire a weeding habit, as
Moller assumes for the species of Atta.
Such an explanation still leaves riddles to be answered in con-
nection with the cultivation of fungi by Termites; but perhaps
it helps to elucidate part of the problem and points to the
solid ground of natural law.
210 SPOLIA ZEYLANICA,
DESCRIPTION OF A NEW SPECIES OF “OPIUS” FROM CEYLON.
By P. CAMERON.
Opius dacusit, sp. nov.
" UTEOUS, the flagellum of antennz and the hinder tibiz and
tarsi black; the wings hyaline, with black stigma and
nervures; @. Length near 5 mm.; terebra fully 5 mm., Pera-
deniya; bred by Mr. E. E. Green from Dipterous maggots
infesting fruit of Cucurbitacee.
Antenne longer than the body, over 60-jointed, towards the
apex minutely haired. Head smooth and shining, the face
covered with white pubescence. Prothorax and mesothorax
smooth and shining, the parapsidal furrows deep; middle lobe
of mesonotum clearly separated, long, gradually narrowed from
the base to the apex which reaches close to the scutellum.
Scutellar depression wide and deep; a stout keel in its centre.
Metanotum areolated ; the areola large, its base narrowed to a
sharp point; the rest slightly, gradually narrowed towards the
apex, which does not reach to the end of the segment; it is
closed and transverse there ; there is a somewhat triangular area
bordering the base of the areola; the other arex are not very
clearly defined. The basal two segments of the abdomen are
closely, regularly, longitudinally striated; the other segments
are smooth and shining ; the first is longer (but not much) than
its width at the apex; the second is wider than long. Pleure
smooth. Mandibles black on the apical half. First abscissa of
radius very short, basal abscissa of radius curved; the recurrent
nervure is received in the second cubital cellute ; the transverse
median nervure is received distinctly beyond the transverse
basal. Malar space as long asthe antennal scape ; the clypeal
fovee large,deep. There is astout keel down the centre of the
basal abdominal segment, the lateral strie being weak compared
with it, The abdomen is slightly shorter than the thorax, broad
and sessile at the base; the apex is bluntly rounded. Meso-
pleural furrow smooth.
SPOLIA ZEYLANICA, 211
NOTES.
1. Historic Trees.—The substance of this Note appeared in the
correspondence columns of the Ceylon Observer on December
29, 1905, and January 6, 1906.
It is probably due to Sir James Emerson Tennent that an idea
prevails that Baldzus preached his first sermon in Ceylon under
the big tamarind tree which still flourishes at Point Pedro.
Tennent says: “Close by the beach there is still standing the
tamarind tree ” commemorated by Baldzus, who preached under
its shade to the first Protestant converts in Ceylon (vol. II.,
p. 535).
It isno doubt due to this passage in Tennent that Mr. J.J. Cotton,
M.C.S., in his “ History of Monumenta] Inscriptions in the Madras
Presidency,” recently published, refers to “ the tree at Point Pedro
under which the celebrated Baldzus preached his first discourse
to the natives.”
But Baldzus, though he certainly commemorates the tree by
recording that “just before the church stands a tall Tamarin
Tree,” which affords “a very agreeable shadow in the heat of the
day,” says nothing about having preached under it himself, but
merely that “the people are often instructed by the Minister to
the number of 3,000” under it (pp. 806-7).
It is certainly not true that he preached his first sermon in
Ceylon at Point Pedro, for that he had done at Mannar on 24th
February, 1658, and it is recorded that in the same year, “in the
church at Telippalai, Baldzeus began the introduction of the
reformed religion into the Jaffna Peninsula.” This being the case,
it seems hardly likely that he preached his first sermon to converts
at Point Pedro; at any rate, there is nothing that I know of to
warrant the statement. Perhaps Mr. Donald Ferguson may beable
to throw alight on the subject. (See “ Baldwus and his Book on
Ceylon,” by Donald Ferguson, p. 6.)
Tennent goes on to state that “ this historical tree ” was in his
day 42 feet in circumference at the base of the trunk. I have had
the tree measured at the base of the trunk; it is now 31 feet.
This is doubtless explained by the fact that until 50 years ago the
2F 10-06
ZAZ SPOLIA ZEYLANICA.
tree was surrounded by a masonry platform, built right up to the
trunk. This platform was three or four feet high, and as many
wide, so that in measuring the tree, Tennent’s informant must
have included the platform.
There is no doubt that the tree still existing is the tree referred
to by Baldweus and Tennent. It is situated about 300 yards from
the seashore, and was evidently at one time fully visible from the
landing place. The whole intervening space has since been built
over with the high stone-walled houses, godowns, and temples that
are characteristic of Point Pedro, but the top of the tree can still
be seen from the Customs, and the tree is a conspicuous landmark
for vessels approaching the harbour. It is, I think, the tallest
tamarind tree I have ever seen. | have ascertained from the oldest
inhabitants that there were never any old tamarind trees of any
size in the neighourhood, but that this tree was always said to be
a very old one, I should think it might easily be 300 years old.
The tamarind is a slow growing tree. Opposite it used to stand,
until they were removed by Mr. Dyke to the present sites, the
Police Magistrate’s house and the Police Court, and near them the
Dutch church. There is no trace of any of these buildings now.
The square open space behind the tree is now occupied by the
market. A good many years ago, but within living memory, the
tree and the adjoining Police Court were frequented by monkeys
(the wandura). These have long since disappeared.
Baldeeus alludes to the grateful shade of the tree. There isa
Tamil proverb that a “ widow’s son should not sleep in the shade
of a tamarind tree.” It is supposed to be more pleasant than that
of any other tree, and, therefore, calculated to make any one, who
lies down in its shade, disinclined to move or to bestir himself as
is necessary in the case of the son of a widow.
Christian Frederick Schwarz, the well-known Missionary of
Tanjore, whose monument by Flaxman is in Schwarz’s Church at
Tanjore, visited Point Pedro on 5th September, 1760, for the ex-
press purpose of seeing this tree (see “ List of Monumental In-
scriptions of Madras, by J.J. Cotton). In any case it deserves the
title of “a historical tree.”
As to the age to which tamarind trees attain, I think there can
be no doubt that like the oak they live for some hundreds of years.
I have seen a tamarind tree at Kachchilamadu in the Mullaittivu
District, which is still known as “Pandara’s tree.” Pandara was
a Vanni chieftain, who enteredinto a league with the Sinhalese
chiefs of Nuwarakalawiya who were hostile to the British, and was
defeated here by Captain Driberg on 31st October, 1803. (See
MOGRISH DAGGER AND SHEATH & }.
To face p. 213.)
NOTES. 2a
“ Vanni Manual,” pp. 19-63.) The tradition is that after the fight
the arms of the defeated Sinhalese (for they were chiefly Sinhalese
of Nuwarakalawiya) were piled under thistree. I had it measured
the other day ; the trunk was 21 feet in girth, 4 feet from the
ground, so that it is 10 feet less than Baldzus’s tree. There isno
doubt that this tree was in existence 102 years ago, and in all
probability it was even then a large tree. Native opinion is that a
tamarind tree may live two or three hundred years. I have seen
very large trees inthe Vanni, and they always indicate the sites of
abandoned villages—villages abandoned perhaps 100 years ago or
more.
The Park, Jaffna, J. P. LEWIS.
2. Lhe Moormen’s Weapon.—I annex a photograph of a curious
kind of dagger with its wooden sheath from the collection of the
late Mr. R. W. levers, C.M.G., C.C.S., which has been acquired
for the Museum. It is known as a samusadu or jamyadu,
for both forms of the word are used, and it appears to be the dis-
tinctive weapon of the Moormen of Ceylon, for a representation
of it is one of their brandmarks for cattle. Thereare two forms of
this brandmark given in the paper on “ Brandmarks on Cattle,”
by the late Mr. James de Alwis, published in the Journal of the
Ceylon Branch of the Royal Asiatic Society for 1874 (vol.
V., p. 60). In one (No. 51) it is represented as of the shape
(fig. 1), which shows that the weapon itself was hardly familiar
to the people who used its shape for a brandmark.
In the other (No. 53), which is given as the brandmark of the
Moormen of Chilaw it has become an ordinary dagger (fig. 2),
but in the Tangalla District, where I also found it used as a
brandmark by the Moormen, the shape was nearer to the original.
I met with three forms. 3
Here (fig. 3) the idea of a weapon also seems to have been lost, and
this particular brandmark was described to me as tambu jadiya (a
copper jar) into which the word samusadu had been corrupted in
the course of time—another proof that the true origin of the mark
had been forgotten. The other shapes were known as ulmadakku
samusadu. (samusadu with an inwards bend) (fig. 4), and
pera-madakku-samusadu (samusadu with an outwards bend)
(fig. 5).
I do not know where Mr. levers came across this specimen.
Possibly specimens of the weapon might be found in India,
214 SPOLIA ZEYLANICA.
where in all probability this particular form of dagger had
its origin, 4
[
3
As
This specimen is rather an elaborate one, the handle and sheath
having on them silver work of artistic design. It is a foot
in length. It is noteworthy that the handle is rather too narrow
for a Kuropean hand to grasp comfortably, as I have found to be
the case also with Ceylon knuckle dusters.
January 1906. J. P. LEWIS.
[The weapon described in the foregoing note has the same form
as the Katar or Indian dagger, which has been described as the
national weapon of the Hindu and “is mentioned by Ibn Batuta,
who lived in the days of Mohamed Toghluk—that is, about
1532 A.D.”
NOTES. 215
Examples of it, beautifully damascened, are figured by T. Hol-
bein Hendley, C.I.E., in his work entitled “ Damascening on
Steel or Iron, as practised in India ;’ London, 1892. ]
3. Scorpion stings.—In connection with my Note on this
subject in Part XI. of this Journal, the following extracts from
letters which I have received from Dr. A. K. Coomaraswamy,
speak for themselves.
KE. #. GREEN.
Extract referred to.
‘You know scorpion and centipede stings and bites are
supposed not to be bad, except on “ Pohoya” days, four timesa
month. The bite of venomous serpents is also worse on those days.
Another point of interest is that one should not kill the animal
that inflicts the wound (snake, scorpion, or centipede) until the
next day, as there is an idea that in some way it can take back the
poison if it remains alive.”
“7 was stung in the foot one evening at Avisawella by the small
species of scorpion and it hurt badly, and I expected to have a bad
time. I rubbedin ammoniaandchunam. The pain went off soon
and in ten minutes there was scarcely any. No trace next day.
Mr. W. D. Holland tells me the scorpion stings are of no account
too. I have told you about bad days for bites. People rely on
mantrams as a cure a good deal. Here is one given by the
Kadadeka Pansala Priest to Mr. Saxton as a sastvi for snake
bite :—
Det HS HS acdsee:
at da da groom Osan:
@dt Ja Seo DS OED
if. e.—On turi turi yéswah
On jaya jaya annatyanéswah
On jaya jayan khapdléswah
“Jf the snake bites you on the left side, you must stroke
yourself with the left hand downwards from top of head right
down the arm, and touch the ground with your righthand ; do this
manytimes. If the snake bites youon the right side, do the same
with the other hand. The above (mantram) is to be learnt by
heart and repeated ”’.
216 SPOLIA ZEYLANICA.
* When I was stung by the scorpion they would not let me kill
it till the next day. Probably it was considered that, as with
snakes, the animal withdraws the poison to itself if not killed.”
A. K. COOMARASWAMY.
4. Flight of white and yellow butterflies.—This well-known
phenomenon takes place twice annually, in the months of
February and November, and is always a source of interest, not
only to entomologists but to all who have eyes to see, and many
are the remarks of wonderment at this curious and pretty sight
which enlivens the monotony of the streets and byways of
Colombo. Iam not going to explain this bi-annual migration, for
I cannot, but these few lines are based upon observations taken
during the flights, more especially the flight of November last,
which commenced on the lst November and lasted practically
till the end of that month, although a few stragglers are still to
be found laying eggs which will give rise to the February flight.
The butterflies are almost exclusively composed of different
species of Catopsilia, and as these show a variety of forms and
phases the whole makes up a very varied collection of yellows
and whites.
T have taken great interest in these flights for several years, but
never before have I seen the numbers so prolific as during last
November in Colombo.
At 2 P.M. on the 21st November, when the flight was at its
highest, I counted during one minute 150, during another minute
170, and during a third minute 148 butterflies passing between
two points 30 feet apart and 12 feet high, making an average
of 156 per minute. At two other places I made similar reckon-
ings, resulting in an all round average of about 150 per minute
traversing a given space. In many different parts of the
Cinnamon Gardens I observed equally large quantities in flight so
that, according to my computation, the number of butterflies
passing in one direction between two points, a mile apart and 12
feet from the ground, would be 26,400 per minute, 1,584,000 per
hour, and 6,336,000 between the hours of 11 A.M. and 3 P.M. on
the one day alone.
They were flying particularly fast and in constant procession ;
rarely one would stop for a few seconds to alight on some
NOTES. 217
conspicuous flower, such as the Hibiscus and Cosmos, as though to
quench its thirst, then away again in greater haste than ever as
if to make up for lost time.
As usual in Colombo, the flights were northwards, but this
does not appear to be the case all over the Island, for Major
Manders* observed that they followed the coast line, starting
somewhere north of Trincomalee and working right round the
south coast to Negombo. Major Manders does not record the
direction of the flights in the more central portions of the Island,
for instance, Nuwara Eliya, Kandy, Diyatalawa, &c. In Nuwara
Eliya and Diyatalawa a westerly direction is taken and this is
probably maintained all over the central portion of the Island ;
it would therefore be interesting to know the direction taken by
flights between the central provinces and the east coast, namely,
whether they strike east and join the circuminsular flight or
strike west passing over Diyatalawa, Horton Plains, Nuwara
Hliya, &c.
Major Manders, in his interesting notes, deals with the direction
of flight of those proceeding from Trincomalee to Negombo, 7.e.,
in the maritime regions only, so that we have still to ascertain
where the butterflies come from which pass over the central
provinces:
The abrupt manner in which the flight starts is very remark-
able. For instance, last November a few straggling. Catopsilias
were to be seen on the first of that month ; on the following day
the flight was in full swing, the numbers gradually increasing
until the 2lst, then diminishing to quite a few on the 3]st.
Before the 2nd November there was no regular flight in one
direction.
In the case of Catopsilia pyranthe I have carefully observed
that the females begin laying their eggs towards the end of the
rush, the few straggling females being then responsible for the
next or February flight. In November of last year oviposition
commenced towards the end of the month and is still going on
up to the time of writing, as an examination of the food plant
(Cassia occidentalis) will show.
The question now arises—When and where are the eggs laid
which produce the prolific November flight? The eggs laid
towards the end of November flight are undoubtedly responsi-
ble for the February flight—the period normally occupied from
egg to imago being from three to four weeks. (In countries
having a winter many insects hibernate, but I have not observed
* Trans. Entom, Soc., London, 1904, No, 26.
218 SPOLIA ZEYLANICA,
such a dormant period in any Ceylon butterflies). If the females
of the February flight lay the eggs for the November flight,
presuming they are to hatch in November, the chances are that
the heavy rains of S.W. monsoon (May to August) would destroy
the majority, leaving few for the November flight, whereas the
latter is the more prolific ; but supposing the feeding grounds of
the larve of the latter are in districts not effected by S.W.
monsoon then it is difficult to understand why they, apparently,
do not emerge and form a flight similar to the other two, some-
where about March, April, or May, but, in Colombo, at any rate,
practically no Catopsilias are to be seen during the seven or eight
months preceding the November flight. I can only arrive at two
explanations :—
(1) The offspring of the February flight must lie in a dormant
state during one or other of their stages, for several months; or
(2) the species must breed throughout the year in some favour-
able locality as regards food-plant and weather. The reason they
do not migrate at regular intervals is most probably due to the
habit of their favourite food plant, which is an annual leguminous
shrub ; it dies off during the first dry months of the year after
shedding its seeds ; these spring up again with the commence-
ment of the rains, and about September-October the plant is plen-
tiful and vigorous, thus forming ample food supply for immense
numbers of the larve. Their numbers increase so rapidly and
to such an extent that the females, prompted by some natural
instinct, migrate, followed by males, and seek pastures new
whereon to deposit their eggs. Thus the bi-annual migration
seems to arise from the abundance of food plant available, which
is greatest shortly before the November flight takes place,
gradually diminishing as the dry weather approaches.
The species Catopsilia chiefly concerned in these flights are
Catopsilia pomona and its variety crocale, and C. pyranthe with
its varieties gnoma, ilea, and chryseis. CU. pomona was most
predominant, but I did not secure a sufficient number during
the flight to ascertain which sex was most abundant. In the
case of OC. pyranthe the males appeared to predominate during
the November flight, for out of 100 specimens secured in one
day, 87 per cent. were males,although towards the end of the flight
the remaining stragglers were almost entirely females. In a
previous capture during a February flight in Colombo (referred
to by Manders) 75 per cent. were females.
OSWIN S. WICKWAR,
Colombo, January 20, 1906.
NOTES. 219
5. Notes by the Way.—In company with two Entomologist
friends I have recently visited the low-country below Koslanda
(in the Province of Uva); making Telulla resthouse our head-
quarters. These notes cover a period extending from November
15 to 24.
On a section of the road between Koslanda and Wellawaya a
few examples of the rare and local butterfly Symphedra nais
were observed and captured. Iam toldthat it always haunts this
particular locality. I have previously taken the species only on
the road to Batticaloa. Manders quotes “the eastern side of the
Island near Trincomalee and the grassy country near Haldum-
mulla” as localities for this insect.
The annual migration of butterflies was in full progress.
Large numbers of them were following the cart road, appearing
to find it a convenient route for travel. They were moving ina
northerly direction. Catopsilia crocale and Appias paulina
were most in evidence. But Hebomoia glaucippe, Ixias ceylonica,
Papilio erithonius, jason, crino, and aristolochic were associated
with them. Huplea asela was also on the move, but not in very
large numbers. It was distinctly a flight of Pieridae, more
particularly of the genus Catopsilia. They settled in dense
array at every damp spot in the road, and rose up in whirling
clouds when disturbed by passing carts or pedestrians. Various
species of Cassia along the route had evidently provided food for
their larve. P. avristolochie appeared to be more exclusive,
having its separate congregating spots. Cvino usually settled
singly. The other butterflies, noticed above, might be seen
mingled in the same group. A very remarkable abnormality of
erithonius was captured by one of my companions and has been
presented by him to the Colombo Museum. The area beyond
the postmedial band on both wings is almost uniformly black.
We collected moths, each night, pitching our powerful
acetylene lamps in different spots—often in the heart of the jun-
gle, and secured a prodigious number of moths and miscellaneous
insects, including such rarities as Azygophleps scalaris, Gunda
apicalis, Stawropus viridescens and S. grisea, Mimeusemia ceylo-
nica, and Afgocera bimacula ; with many smaller species probably
new tu our lists. My attention was drawn, one night, toan insect
that, to outward appearances, was an ordinary “ Daddy-long-legs”
(Tipula). It had captured a small Pyralid moth, which it was
holding with its hind legs and devouring. On closer exami-
nation it proved to be one of the so-called “Scorpion Flies”
(Panorpide) and appears to be a species of Bittacus. To the
best of my belief, this family of insects has not previously been
26 10-06
220 SPOLIA ZEYLANICA.
recognized in Ceylon. A long series of this species was taken in
the moth traps.
In the last number of the Journal of the Bombay Nat. Hist.
Soc. (vol. XVI, No. 4, p. 747) is a note by Capt. Nangle, describ-
ing an attack by flies upon flying Termites in India. A similar
occurrence was noticed one night at Telulla, when a flight of
winged Termites invaded the tent of my moth trap. Imme-
diately following the ‘White Ants” came a number of Muscid
flies which proceeded to pounce upon them. They did not
attempt to carry away their prey, but commenced to feed upon
them there and then, piercing their helpless victims between the
segments of the abdomen. Specimens of this fly have been sent
to Europe for determination.
. Beating for moths, in the daytime, was made difficult by the
presence of enormous numbers of the common Noctuid Amyna
selenampha, which rose in clouds whenever the undergrowth
was disturbed. We saw several trees stripped of their leaves by
the larve of this moth. Fortunately this species did not fly at
night or—at any rate—respected our moth traps.
Our traps were pitched, one night, on the bund of the Telulla
tank. Behind us was an abandoned paddy field. As soon as the
lamps were lighted myriads of frogs joined in a deafening
chorus which soon became so intolerable that we had to stop our
ears with cotton wool.
Reptiles were not much in evidence. But we surprised a large
specimen of Dendrophis pictus swallowing a full-grown lizard
(Calotes versicolor). Younger examples of this snake are said to
feed upon grasshoppers. A fine example of the large rough
tailed snake—Uropeltis grandis—was picked up on the cart road
where it was engaged in extracting grubs from a pat of cow
dung.
EK. ERNEST GREEN.
6. On the Species of Leaf Insects (Phylliine) occurring in
Ceylon.—Mr. W. F. Kirby, in his recently issued “ Synonymic
Catalogue of Orthoptera,’ credits Ceylon with three distinct
species of this subfamily of Phasmide. These are,—Pulchrj_
phyllium agathyrsus, Gray ; Pulchriphyllium crurifolium, Serv.,
and Phyllium athanysus, Westw.
I have recently submitted specimens of our commoner species
to Mr. Kirby, for an authentic determination. There has hitherto
been considerable confusion in the nomenclature ; the commonest
Ceylon species having been variously referred to under the names
2 pane
Fic. 1-—a@ PULCHRIPHYLLIUM CRURIFOLIUM.
xX 2
b PHYLLIUM
ATHANYSUS.
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NOTES. 22)
of seccifolium, scythe, pulchrifolium, and bioculatum. In reply,
Mr. Kirby tells me that this species is apparently crurifolium,
which Gray confused with his previously described biocwlatum.
“Tt has nothing to do with Phyllium siccifolium, which does not
occur in Ceylon.” He also states that ». athanysus can be recog-
nized at once by the hinder edge of the front femora being entire,
and that P. agathyrsus “ seems to differ from P. crurifolium in
having the hinder edge of the front femora more deeply and
irregularly excavated.”
Cc j d
Fig 2. (a) Egg of Pulchriphyllium crurifolium eek
(6) Do. Phyllium athanysus )
Fig 3. (c) Femur of front right limb of P.crurifolium x12
(d) Do. do. Phyllium athanysus
Of these three species, crurifolium is comparatively abundant.
Native boys collect them in considerable numbers, in the neigh-
bourhood of Kandy. They find them by waiting under the trees
that they are known to affect and watching for falling pieces of
green leaves dropped by the feeding insects. This species is very
easily raised from eggs laid by captive females.
I have met with very few examples of athanysus.
Agathyrsus is quite unknown to me, though it is recorded only
from Ceylon. If the “more deeply and irregularly excavated
hinder edge of the front femora” is the only distinction between
this species and crurifolium, then a great number of bred exam-
ples of the latter might be classed as agathyrsus, for very few
222 SPOLIA ZEYLANICA.
individuals reach maturity without having this part of the limb
more or less extensively nibbled away by their comrades in cap-
tivity. Can this accidental erosion have led to the supposed
distinction of the species? The slender individuals in the lower
right hand corner of the figure (4) are the males, which have
functional wings and very small wing-covers, the reverse being
the case with the female insects.
There is a well marked difference in the eggs of crurifolium
and athanysus. ‘That of the former has five winged lateral ridges
and a prominent spiked cap. The egg of athanysus is very much
smaller, has only asmall tubercle on the cap, and the lateral ridges
are low and inconspicuous, merely giving to the eggs a slightly
angular appearance.
In the accompanying photograph, the second and third insects
in the middle row, show signs of having been nibbled by their
companions.
E. ERNEST GREEN.
7. A Harbour Worm and a Boxing Crab.—Ever since the
visit of Professor Haeckel to these shores in 1881 it has been
recognized that Colombo offers few inducements to biologists
who cumber themselves about things of the sea. Nevertheless, if
time and opportunity allowed, a great deal might be gleaned
from the outer;reaches of the Colombo Harbour; and the Northern
Arm of the Breakwater is already attracting a host of creatures from
the surrounding depths. Amongst these there comes occasionally
a remarkably fine Annelid worm belonging to the family Am-
phinomide. Its name is Chloéia flava (Pallas); it is particularly
characteristic of the Indian Ocean and has been known to
naturalists for nearly a century and a half. It attains a length of
nearly five inches or more when fully extended and an inclusive
breadth of about one inch. The number of segments is limited,
not exceeding forty, and this number is only reached in the
fully mature condition. Smaller and therefore younger indi-
viduals have fewer segments, the number of the latter being
roughly correlated with the size; thus examples about half to
three-quarters of an inch in length will have 24-26 segments.
The head region is characterized by the presence of a peculiar
organ called the caruncle, which stretches back over several
of the anterior segments. Behind this region, each segment
Fic. 4 —EGGS, LARVA, NYMPH, AND ADULT (4 aND &) OF “FULCHRIPHYLLIUM CRURIFOLIUM.”
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NOTES. 223
carries along the dorsal side a large purplish violet spot, a pair of
arborescent gills* and, at the sides, two dense tufts (upper and
Chloéia flava.
lower) of long, hair-like, golden yellow bristles or seta, with
which: are associated dorsal and ventral sensory cirri. In
* The gills are coloured a dark sepia with rose-coloured rhachis. Violet
bands may also occur laterally between the bases of the gills and the dorsal
bristles, which appear pale, golden-tipped. The colour is of course snbject to
some variation.
224 SPOLIA ZEYLANICA.
preserved specimens it often happens that most of the sete are
lost ; they are deciduous, those of the dorsal tuft being provided
with recurved barbs by which they remain attached to foreign
bodies when touched or handled. ‘The function of the set is
therefore partly defensive, but during what may be called the
swarming period the tufts of sete act as paddles or swimmerets,
working to and fro in succession like the legs of a centipede,
accompanying the swaying movements of the body when swim-
ming. It is not common to obtain full-sized specimens of this
worm, and the three which the Museum has acquired in recent
years have been kindly sent by Mr. A. D. Prouse from the
Harbour Works, one in March, 1904, another in January, 1905,
and a third in January, 1906.
The small crab which I have called the Boxing Crab was taken
by me last November at Weligama from under a rock close
to the shore at the northern end of the bay. It affords a singular
and by no means widely known example of symbiosis or com-
mensalism between a crab and sea-anemones or Actinians. In
each claw the crab holds a small white sea-anemone in full
activity, tentacles expanded, which it presents with great science
in true boxing attitude to the observer, when alarmed. It is
impossible to exaggerate the singularity of the action. The
ground colour of the crab is whitish with a delicate roseate flush
in the anterior half of the carapace, and a characteristic pattern
formed by blackish lines; there are tufts of bristles behind
the frontal margin and on the surface of the carapace.
I forwarded a sketch of the living crab and its two guests to the
Indian Museum, Calcutta, and was favoured by Major A. Alcock,
F.R.S., with the name of the crab and references to the literature
dealing with it.. It is not mentioned in Dr. J. G. de Man’s |
Crustacea of the Mergui Archipelago (Journ. Linn. Soc., London,
NOTES. 225
XXIJ., 1888), nor in the same author’s work on the Crustacea
of the Malay Archipelago (Decapoden des Indischen Archipels,
1892), nor in Dr. J. R. Henderson’s Contribution to Indian
Carcinology (Trans. Linn. Soc., London, vol. V., part 10, 1893).
It was described from Mauritius under the name Melia tresselata
by H. Milne Edwards in 1834 (Hist. Nat. Crust., Paris, vol. I,
p. 431, pl. 18, f. 8) and figured though without the actinians. Its
correct name is Melia tessellata (Latreille) and a capital figure of
*t is given by L. A. Borradaile in his Report on the Marine
Crustaceans in Stanley Gardiner’s Fauna and Geography of the
Maldive and Laccadive Archipelagoes, vol. I., Cambridge, 1903,
p. 249.
Borradaile places this crab in the subfamily Menippine of the
family Xanthide and points out thatits habit of holding actinians
was noted in 1880 by Richter in Mobius’ Meeresfauna, Mauritius,
but since then the fact has been generally overlooked.
Probably both crab and actinians benefit by the association, the
actinians enjoying an increased mobility and the crab sheltering
and defending itself with the living gloves with which it is
provided ; all sea-anemones, coral polyps, hydroid polyps, and
meduse possess microscopic weapons of offence called urticating
or stinging threads, and no doubt these come into play in this case
when occasion requires it, although I was unable to detect their
presence by the sense of touch. On this subject | may quote from
Mr. Borradaile’s work (.c., p. 250). Hesays: “The object of this
habit is not known, but it is certainly a voluntary act on the part
of the crab, for the actinian is not attached, but held between the
fingers of the Melia, and, if it be taken away, will be again seized.
Usually there is an anemone in each hand, but sometimes one or
both hands are empty. The actinians, which are grasped firmly
round the middle below the tentacles, may be useful, by means of
their stinging cells, either for defence or to ‘fish’ for food with,
or perhaps for both purposes. The chelipeds are slender and
feeble, ill-suited for defence, but at the same time mobile and
well adapted to wield the anemones they carry, and, if the crab be
threatened, it will stretch out its arms towards the aggressor, as
though it would ward him off with the disagreeable obstacles it
thus presents to his attack.”
The crab from Weligama is small, measuring only 10mm. in
width between the lateral angles of the carapace ; the actinians
are retained in position in the preserved state, but the ground-
colour has darkened to a reddish brown.
Finally, it must be mentioned that two other species of Melia
have been recorded from these seas by Major Alcock, namely, M.
226 SPOLIA ZEYLANICA.
cestifer (J. A. S. Bengal, vol. LXVII., pt. 2, 1898, p. 231) and UM.
pugil (ibid.).
I desire to express my thanks to Major Alcock for his kindness
in referring me to the above sources of information concerning
Melia.
A. WILLEY.,
Colombo, January 23, 1906.
8. Stridulation of Gongylus gongylodes.—During a recent
visit (December) to Anuradhapura as I was descending the bund
of Tissa tank behind Isurumuniya, I espied a dark brown
Mantid upon a low green shrub, and upon bending closer found
that it was a male Gongylus. Tts actions upon being alarmed by
my approach arrested my attention. Its colour and general
appearance were those of a brown shrivelled leaf, but it appeared
to be intent upon a foraging expedition with no idea of mimicry
or of any conformity to its surroundings. The under surface of
the flattened prothorax had a dull neutral tint paler in the centre,
but not glistening norin any way attractive. Such mimicry and
protective coloration as carnivorous animals—of which Gongy-
lus is one—frequently exhibit appear to be closely correlated with
their infinite capacity for lying motionless in wait for passing
prey.
As I drew near to the Mantid several times in succession,
on each occasion it dressed itself for the fray, squaring the elbows
of its forelimbs nearly at a right angle to the body as shown in
the upper figure of the second plate illustrating this species
in Spolia Zeylanica, part VIII. (vol. II,), 1905, page 198. The
alert defensive attitude which it assumed was evidently intended
to be terrifying in the sense in which this term has become fami-
liar to students of mimicry, and the terrifying effect was enhanced
by a loud rasping sound which it produced by rubbing the
borders of its hind femora against the rim of the forewings.
The male is distinguished from the female by greater slender-
ness and length of body and wings, by the smaller size of
the foliaceous expansions of prothorax and limbs, and above all
by the feathery antennex, these being hair-like in the female.
The outer free border of each forewing is thickened by a distinet
chitinous rim which is finely serrate. In the region of the hind
femur, on each side, the border of the wing is slightly concave,
allowing free play to the thigh under ordinary circumstances.
The femur itself is smooth, carrying a few minute hairs but
without any rough edge,
NOTES. 227
When the insect is alarmed each hind femur is deliberately
rubbed to and fro against the saw-like edge presented to it along
the concave border of the wings and a very effective stridulat-
ing sound results from the friction. The sound can be approxi-
mately reproduced upon the dead insect by gently passing a
porcupine quill backwards and forwards along the wing-border.
The serrate border of the wing is also present in the female
where the concavity is more pronounced ; and it seems likely that
the sound can also be emitted, on occasion, by the female.
Many pther insects, especially beetles, produce sounds by rasp-
ing. The stridulation of the caterpillar of the Death’s Head Moth
has been alluded to on page 47 of this volume. Many grass-
hoppers emit a loud clicking sound when rising from the ground
in flight, but I have not yet succeeded in ascertaining how this is
produced. The vocal apparatus of male locusts and cicadas be-
longs to a different category of sound-producing organs, and the
object of the call is also different.
What may perhaps lend particular interest to the habit of
stridulation as manifested in Gongylus is the deliberation with
which the action is performed by an inseet which is well known
in other ways for its defensive and offensive tactics.
A. WILLEY.
Colombo, January, 1906.
9. Terrestrial Colubride of Ceylon.—In his admirable syste-
matic work on the Reptiles of British India, Ceylon, and Burma
(1890), Mr. G. A. Boulenger notes the existence of a general desire
felt by those not well acquainted with snakes to know at least how
they may distinguish poisonous from harmless kinds. It is not
a simple matter and there is no way of rendering it simple. The
examination of the teeth is the fundamental test, but they are
often broken and some snakes, the Dipsadinz, possess grooved
fangs and yet are not poisonous. It is in fact necessary to know
the snake before pronouncing upon its character.
In an island like Great Britain where only three species
of snakes occur, one of which—theadder—is poisonous though not
deadly, their identification is not a matter of daily or frequent
necessity. In Ceylon, which covers an area of some 25,331 square
miles, smaller in extent than that of Ireland and yet possesses an
ophidian fauna comprising, exclusive of the sea-snakes, upwards
of fifty species, of which eight are poisonous and three or four
24 10-06 °
228 SPOLIA ZEYLANICA.
fatal, their determination not only stands in constant request.
but may even assume a medico-legal or statistical importance.
In the Vital Statistics issued by the Registrar-General of Ceylon
for the year 1903, the number of deaths attributed to snake-bite
during that year amounted to 217, and the average from 1898 to
1902 is given as 198. In 1904 the number of deaths from snake-
bite was 174, and the average from 1898 to 1903, 201°2, or in
round numbers 200 per annum.
In the Indian Empire, which is roughly about seventy times as
large as Ceylon both in area and population, there is only
a slightly greater proportion of deaths attributed to snake-bite
annually ; namely, about 20,000.
I believe it rarely happens that the offending reptile is caught
and identified, so that statistical tables give little or no informa-
tion concerning the number of victims claimed by different
species of poisonous snakes.
Last September a Malay woman living in Colombo was bitten
by a snake during the night while she was asleep. She must
have unconsciously alarmed the snake in some way, other-
wise it would not have bitten her. A vedarala was called in, but
the woman was beyond help, and died within twelve hours.
The snake had been caught and killed at the time of the accident
and was subsequently sent to the Museum by the Coroner for
identification. It proved to be a dangerous poisonous snake of
the kind called “bungarum ” or,in Northern India, ‘ krait,” in
Sinhalese “ karawala.”
Two months later, in November (1905), another snake was sent
to me from Mullaittivu by Mr. R. A. G, Festing, C.C.S.. with the
information that it had bitten and killed 2 woman two nights
previously. This time the snake was a harmless one and very
common in Ceylon, Lycodon aulicus, often called “ karawala ” or
‘tel-karawala” through confusion with Bungarus, since it
resembles the latter very strikingly and may be said to mimic it.
It was generally believed that the victim in this case had died ot
fright, since it appeared from the evidence at the inquest that she
had been bitten, twelve small punetures being visible on the
right forearm. ,
Bungarus may be recognized at once by the circumstance that
the median dorsal or vertebral scales are much enlarged ;, in
Lycodon the scales are equal. It is, however, partly owing to the
extraordinary parallel series of colour variations in poisonous and
harmless snakes, giving the impression of a natural mimicry, that
so much confusion exists with regard to the vernacular names in
this country. There are also other sources of confusion, The
NOTES. 229
late Mr. William Ferguson, F.L.S., pointed out in 1877,* that the
common Pit-viper, well known here as the “ kunukatuwa,” was
originally figured by Dr. Davey as the “ karawala,” and the latter
erroneous name has adhered to it in English writings ever
since.
There are three or four kinds of so-called ‘“karawala” in
Ceylon, namely, the ‘“‘dunu-karawala,” Bungarus ceylonicus, the
“mal-karawala ” or “ pol-mal-karawala,” Chrysopelea ornata ; and
the “mudu-karawala,” Coluber helena. The expression “ tel-
karawala” is sometimes applied indifferently to varieties of
Bungarus ceylonicus and Lycodon aulicus,t in allusion to the oil-
like gloss of the scales. The term “ mapila” belongs to certain
snakes which have a suspicious look but are not dangerous,
namely, species of the genus Dipsas (cf. Spol. Zeyl., vol. 1., part
III., 1903, pp. 81-84).
The most convincing evidence of mimicry as between harmless
and poisonous snakes is afforded by the specimens of a white-
ringed variety of Bungarus ceylonicus and the white-ringed
species of Lycodon, L. carinatus, which have been kept in the
same jar in the Museum. The similarity between these two
forms was remarked upon in 1877 by Mr. William Ferguson. To
the unaided eye the resemblance is quite deceptive, but with a
lens it will be seen at once that the scales of the Lycodon
carinatus are keeled, besides being equal.
Precisely analogous examples of mimicry are known to occur
among the so-called coral-snakes of Brazil, of which there are four
genera, one (Hlaps) poisonous, a second suspicious, the two others
harmless. The colour is described as a “ rich red with black and
yellow transverse bands,” and the species of these snakes
inhabiting the same districts are so like each other that only
a naturalist could distinguish the harmless from the poisonous
kinds (¢f. Darwin, Descent of Man, Second Edition, p. 353).
The accompanying schedule, which is based primarily upon
Mr. Boulenger’s Monograph, is intended to serve as a guide to the
identification of a section of the snakes of Ceylon. It is complete
so far as it goes, including as it does all the members of the three
subfamilies of Colubride mentioned, namely, the Colubrine,
Dipsadinez, and Elapine, which are known to occur in Ceylon.
The sea-snakes (which are poisonous) and two other small sub-
families of water-snakes (Acrochordine and Homalopsine)
* Wm, Ferguson, Reptile Fauna of Ceylon. Letter on a collection
sent to the Colombo Museum. Colombo, 1877 (printed by the Government
Printer).
+ Lycodon aulicus is also called ** alu-polonga”’ at times.
230 SPOLIA ZEYLANIGCA.
are omitted. The earth-snakes (‘“depat-naya”’), the Python or
rock-snake (“ pimbura”), and the Vipers (‘“ polonga’”’) are not
included within the scope of the schedule.
Two points may be noted in connection with the guiding
characters which I have selected in the index. All the species
mentioned have less than thirty rows of scales round the body
(excluding the ventral shields) and in all cases the general con-
dition of the head-shields is normal and remarkably constant on
the whole. Seen from above there is a rostral shield in front
(usually barely visible in dorsal view) followed by a pair of
internasal shields between the nostrils, then a pair of prafrontals,
a median frontal between two supraoculars and a pair of parietals
behind. ‘he shields which appear in side view of the head are
less constant, the most important feature being the presence or
absence of a loreal shield between the preocular and the nasal,
this being one great distinction between Lycodon and Bun-
garus.
When once a character has been observed all the species which
are shown by the index not to possess this character are excluded,
and thus by continuing the process of exclusion at least the genus
of the specimen can be fixed with certainty, and this is enough for
practical purposes.
A. Side view of head of Lycodon aulicus ; 1. Loreal shield; L*., Ninth
upper labial.
B. Side view of head of Bungarus ceylonicus ; showing absence of loreal
shield; L’. Seventh upper labial.
The species which offers the most generalized characters is No.
XIL., Polyodontophis subpunctatus, a harmless snake which may
NOTHS. 231
be found under logs. I took one recently near Puttalam,* and it
occurs round Colombo and Kalutara,
The regularity with which snake-bite returns appear in Vital
Statistics demonstrates the necessity for a more detailed analysis
than is usually given. Whatis wanted is information concerning
the snakes which are alleged to have inflicted bites on human
subjects. Like as not many of them would prove to be harmless.
In the schedule the numbers prefixed by the letter B refer to
the numbers of the species in Mr. Boulenger’s volume on Reptilia
and Batrachia in the Fauna of British India; S. Z. refers to Spolia
Zeylanica ; Tropidonotus asperrimus, No. XXVI., the Ceylon
representative of 7’. piscator was named by Mr. Boulenger subse-
quently to the publication of the volume (see Ann. Mag. Nat.
Hist. vol. VII., 1891, p. 281).
By way of coincidence it is to be noted that the principal and
proper food of Lycodon aulicus consists of the Brahminy Lizard
or “hikanella ” (Wabuia carinata) whichis swallowed entire by
the snake and may be removed practically uninjured from the
stomach of the latter, if taken at the right time. The size of the
ingested lizard in comparison with the snake is sometimes astonish-
ing. This lizard is believed both here and in South India to
inflict a poisonous bite, and in the Vital Statistics for 1904 (Ceylon)
one death is attributed to injury caused by Brahmin Lizard. In
reality the “ hikanella” is harmless. A similar tradition of folklore
is recorded by Dr. Gadow (Cambridge Nat. Hist. Amphibia and
Reptiles, 1901, p. 506) from the South of Spain and Portugal where
geckos, called “osga,” are abundant and are considered to be
dreadfully poisonous by the country folk. Dr. Gadow points out
that geckos are absolutely harmless; they cannot even inflict
painful bites, but in many countries they are feared as much as
the most poisonous snakes.
There are special difficulties in the way of identifying snakes
which have inflicted bites upon the person. Firstly there is the
difficulty of catching the snake and of making certain that the
one caught is the culprit. Then again in the country districts of
Ceylon, as ail residents in the Island are aware, there is a reluct-
ance to kill the snake, and if killed in some cases, notably in
regard to “‘ mapila ” and “ karawala,” the snake must be burnt in
accordance with immemorial custom. Because the “ mapila” and
* At Karawalagaswewa, on the road from Puttalam to Anuradhapura. This
snake was quoted by Bateson (Materials for the Study of Variation, London,
1894, p. 123) as an example of maximum variation in the number of ventral
shields which correspond with the number of vertebra, as may be easily seen by
observing the ribs pressing against them during locomotion. In this case the
number of ventral shields has been observed to vary from 151 to 240,
232 SPOQLIA ZEYLANICA.
“ karawala ”’ are believed to travel in flocks of seven, and when one
is killed and burnt the smoke is supposed to keep away the other
six. This incineration of snakes is an actval and definite practice
in Ceylon aud the burning of a “ pol-mal-karawala” (Chrysopelea
ornata) under the conditions indicated has been witnessed by my
friend Mr. James Parsons of the Mineralogical Survey, as he has
been good enough to inform me.
Another difficulty in connection with the casual identification
of snakes is more apt to trouble those who know something about.
snakes to begin with than those to whom they are a terra incog-
nita. refer to the occasional capture of snakes of unusual size
belonging to well-known species. A case in point has quite
recently occurred to me as concerning the snake Aspidwra copit.
The specimens of Aspidura which I have seen are slender snakes
of rather small size, a length of two feet being exceptional ; and
the head is long and tapering (compare this Journal, vol. II.,
part VIL. plate facing p. 96).
The other day (January, 1906), a bulky snake was sent to the
Museum from Avisawella by Mr. William Ferguson, Jr., which
severely tested the merits of my schedule. The last genus to
which it might belong seemed to be Aspidura. It had a single
internasal, short frontal, no przocular,* 6 upper labials, 17
smooth scales round the middle of the body, 137 ventral shields
mottled with the dark ground colour, anal entire, 20 unpaired
-eaudals : in short, it wasa female example of Aspidura copii. Its
size however, and therewith its general appearance, was remark-
able ; the length, as nearly as it could be measured in the coiled
condition in which the snake had been preserved and hardened,
amounted to rather more than 25 inches, of which the tail only
occupied two inches; the girth was as much as 33 inches, giving an
appearance of great bulk; the head was widest in the occipital
region, close upon one inch across and greatly flattened, the small
eyes occupying a dorsal position and the snout rapidly tapering in
front of theeyes. In the Fauna of British India, Mr. Boulenger
gives the following dimensions of A. copii: Total length 165
inches, tail 2°8, these figures probably relating to a male.
Upon opening the snake now under consideration, the ovaries
proved to be full of large eggs, three-quarters of an inch long by
about half an inch wide, seven in the left ovary, fourteen in the
right. From what has been said it will be clear that thisis quite
* The preocular shields are shown in the text-figures A and B. Aspidwra
copii and Drummondhayi axe the only Colubrine snakes in Ceylon which have no
preocular,
NOTES.
a noteworthy individual.
233
In Mr. Haly’s Report on the Reptiles,
&c., in the Colombo Museum (1891) A. copii is recorded from Dik-
oya and Balangoda and is said to range between 2,000 and 7,000
SCHEDULE OF THE TERRESTRIAL COLUBRIDA OF CEYLON.
Remarks.
Harmless.
eter Hill-snakes, sometimes
brightly coloured.
feet,
Names.
A.—Colubrine.
Tr. Aspidura brachyorrhos ... B342 ...
II. 7 COpll . B343 ...
IIT. . guentheri . B44 ...
EY. » trachyprocta'... B345 ...
WV: » drummondhayi B.S.Z. ...
VI. MHaplocercusceylonensis B&46 ...
VII. Lycodon striatus Ae sor {ee
VIII. ry aulicus 5. 1B33M aac
IX, . carinatus - DSDG ..
X. Hydrophobus nympha ... B357 ...
XI. gracilis .:. Babs ...
XII. Polyodontophis he a
tatus... anf p04: 255
XIII. Ablabes calamaria ~ B64 =.
XIV. Simotes arnensis ios DOU a5
XV. Oligodon templetonii ... B391 ...
XVI. S sublineatus ... B392 ...
XVII. us subgeriseus ... B394 ..:
XVIII. Zamenis mucosus = BBE Son
XEX.. “3 fasciolatus... B402 ...
XX. Coluber helena SSG) ee
XXI. Dendrophis pictus ~ ALT 2.
2 OIE e bifrenalis... B419
XXIII. caudolineatus B420 ...
XXIV. Tropidonotus ceylo-
nensis... . B430 ...
XXV. stolatus ... B434
XXVI. = asperrimus B1891 ...
XXVII. i plumbicolor B439 ...
XXVIII. Helicops schistosus B440 ...
B.—Dipsadine.
XXIX. Dipsas barnesii . B44 .,..
MX: Hd
BMT li eto:
ceylonensis ...
forstenii
MOK, i,
pulverulentus
XXXIV. Chrysopelea ornata B463 ...
C.—Elapine.
XXXV. Callophis trimacula-
GRAYS ccs teenie...
XXXVI. Bungarus ceylonicus B481 ...
XXX VII. -" coeruleus B482..
XXX VIII. Naia tripudians B485 .
<<a
B446 ...
et BS51 sc.
XXXII. Dryophis mycterizans B461 ...
B462 ...
Striated Wolf-snake, Peradeniya.
Bungalow Wolf- snake, “ Tel-
karawala.”
White-ringed Wolf-snake.
Jaffna, Anuradhapura.
“ Ceylon.”
Collared Sand-snake.
Rare.
Black-banded snake.
Paucidentate Dwarf snakes.
Rat-snake, “‘ Garandiya.”
Jaftna.
Ocellated Land-snake, “ Mudu-
karawala.”’
i ; Tree-snakes, ‘‘ Haldanda.”’
Ceylon Ground-snake.
Striped Ground-snake, “ Ahara-
kukka.”
Amphibious Ground-snake,
‘“ Diya-polonga.”
Viperine Ground-snake.
Freshwater snake.
Non-poisonous.
( Nocturnal Tree-snakes,‘‘Mapila.”’
Green Whip-snake, “ Ehetulla.”
Brown Whip-snake, “ Hena-
. kandaya.”
Florid Tree-snake,
karawala.”
“ Pol-mal-
Poisonous.
Dwarf Bungarum.
Ceylon Bungarum,
karawala.”’
Indian Krait (Jaffna).
Cobra.
* Dunu-
Norr.—Nos. VIII. and XXV. are the sun onERE snakes in Ceylon ; next
in abundance is XX VI., and then follow the Tree-snakes and Whip- snakes,
234 SPOLIA ZEYLANICA.
INDEX.
N.B.—The following common characters may be regarded as occurring
normally since they are presented with great frequency and are therefore
not specially noted in the index ; they are to be presumed unless otherwise
stated :—
(a) Anal Shield divided ; (b) Head
Internasals paired ; (d) Loreal single,
not clearly distinct from neck ; (c)
separated from eye ; (¢) Preocular
single ; (f) Pupil round ; (g) Rostral barely visible from above ; (4) Seales
equal and smooth ; (7) Subeaudals paired.
Anal shield entire i to v., Vi., ix., Xi.,
xx.
Head distinet from neck. x., xi., xviil.,
Kiki, XX. XKLbO KEM. | xy Pto
EVil,, Xxtx, tO RRS, \xxza1;
XXXili,, XXXIV , XXXVili.
Internasal single. i to v., vi., xxviii,
Labials* six. i to v., XXXV.
Labials seven. vi., X., Xi., Xiil., XiV., XV
to xXvii., XXvil., XXXVi., XXXVii.,
XXXViil.
Labials eight. vii., xviii. xix., XxXiv.,
REV MRK) RK, OR RTT
Labials nine. viii., ix., xii., (9-10), xx.,
xxi. (9-10)., xxii., xxiii, (8-9),
XXV1., XXviil., (8-9) xxxiv.,(9-10).
Labials indefinite, xxxi., (8-11).
Loreal absent, i to v., vVi., XXXii.,
SEK, KER, SERVE KV.
XXXVIlii.
Loreal entering the eye, x. xi. xv.
Loreals two, xxii.
Loreals three, xviii.
Maxillary teeth equal, i to v., (20).,
vi. (10)., xii. (40).. xiii. xx., xxi to
xxill.
Maxillary teeth enlarged in front,
vil to ix.
Maxillary teeth enlarged behind, x., |
xi, xiv., xv tO XVli., XVili., Xix.,
Xxiv to XXvil., XXvili.
Maxillary teeth fanged behind, xxix |
tO) SEX, KES eee RI
Maxillary teeth fanged in front,
XXXV., XXXVI., XXXVii., XXXviii.
Colombo, January 23, 1906.
Preocular absent, ii., v.
Preoculars two, xXiv., XXVi., XXX1i,,
XXX1ii,
Preeoculars three. xxix.
Preoculars variable (1 or 2)., X., Xxx.
Pupil horizontal, xxxii., Xxxili.
Pupil vertical, vii to ix., X., Xi., XXix.
to Xxxi,
Rostral} large, xiv., xv to xvii.
Rostral produced., xxxil., XXxXiil.
Seales in 13 rows, x., xxili., xxxv.
Scales in 15 rows, iv., v., Xi., Xili., Xv.,
XVi., Xvii., XXL, Xxii., XXxii.,
XXXlil., XXXVi., XXXVii.
Scales in 17 rows, 1., ii., ili., vi., vii.,
Viil., xil., Xiv. Xvili., XxXxiv.
Scales in 19 rows, 1x., XXiv., XXV.,
XXVi.. XXVIIM_, XXIX.
Scales in 19 or 21 rows, xxx.
Scales in 21 or 28 rows, xix.
Scales in 23 to 27 rows, xx., XXVii.,
XXXVili.
Scales in 25 to 29 rows, xxxi.
Scales keeled, vi., ix., xxiv., Xxv.,
KXVi., XXVii., XXViii.
Scales keeled in posterior part of
body only, ii., iv., v., xviii.t
Subcaudal shields in single row, i to
iv., Vi., ix.
Subocular present, xviii., Xix., XXxii.,
XXXiii,
Ventral shields keeled laterally x., xi.,
EX1., KKil., RXV.
Ventral shields with dark spots at
the sides, xii., XxxXV., XXX., XXXiv.
Ventral shields with dark spots along
sides and middle, xv., xvi.
Vertebral scales enlarged, xxi to
XXlil., XXX., XXXIi., XXXIil., XXXVi.,
XXXVii.
A. WILLEY.
* Upper labials or shields bordering the upper lip.
+ The rostral shield covers the front of the snout.
t In the young rat-snake the keeling of the scales may be absent.
(‘JeqQuaNy Jaye AYQI8q) “VIINVIASS SILOHdOO 371vW
TO i ed cl eth ake
Swe, .
: a tod : - oe reed
NOTES, 235
10. Viviparity of Cophotis ceylanica and Oviparity of Cera-
tophora stoddartii.—Cophotis and Ceiatophora are two genera of
Agamoid lizards containing species which are peculiar to Ceylon.*
Cophotis ceylanica and Ceratophora stoddartii are found in the
districts of Nuwara Eliya (6,200 ft.) and Hakgala (5,580 ft.) ; the
former species is distinguished by the possession of a prehensile
tail, the latter by its rostral appendage (¢f. Spolia Zeyl., part L,
opposite p. 11). Another species belonging to the same family
(Agamidz), also peculiar to Ceylon, is common at Hakgala on way-
side bushes ; this is Calotes nigrilabris.
The other families of lizards which are represented in Ceylon
are the Geckonide, Varanide, Scincide, and Chameleodontide,
Here I am only concerned with the Agamide and Scincide.
It is well known that most of the Agamoid lizards are oviparous,
laying their eggsinsmall holesinthe ground. The occurrence of
viviparity or ovoviviparity, where the eggs undergo their develop-
ment within the oviducts so that the young are born active with
the characters of the parent, is both rare and exceptional among
Agamoids. The only case noted in the Fauna of British India, &e.,
of a viviparous Agamoid is that of the genus Phrynocephalus, not
found in Ceylon (op. cit., Boulenger, Reptiles and Batrachians,
p- 110).
During a recent visit to Hakgala, where I occupied the Botanical
Laboratory by kind permission of Dr. J. C. Willis, at the end of
January and beginning of February, I ascertained that the tree-
dwelling lizard, Cophovis ceylanica, is viviparous, At the proper
season the ripe eggs pass into the oviducts and there undergo their
development without any shell being formed, so that the half-
formed embryo is clearly visible through the thin transparent
wall of the oviduct. One female showed three whitish embryos
lying upon the yolk in the right oviduct and two in the
left, all in the same stage of development; a second had two
advanced embryos in each oviduct, with scales and dark pigment
appearing. :
The size of the eggs, 13 mm. in major diameter, causes the wall
of the oviduct to stretch and to invest each egg closely. After
removal of an egg from the oviduct, the embryo with the yolk is
still surrounded by three membranes, which I interpret as follows
from the examination of flat preparations :—(1) a very thin, trans-
lucent, finely fibrous, non-cellular, vitelline membrane ; (2) an
equally thin and transparent chorionic membrane which is cellular
_ * All species of Ceratophoraareendemic. Cophotis has another species in Suma-
tra (Boulenger),
21 10-06 ©
236 SPOLIA ZEYLANICA.
but non-vascular ; this membrane, which I take to be the epidermal
tract of the outer amnion, consists of a pavement-epithelium in
which the cell-boundaries, vesicular nuclei and nucleoli are clearly
visible without staining ; (3) a thicker, though still transparent,
vascular allantoic membrane traversed by blood-vessels which
give rise to dense capillary networks over the surface of the egg.
From the above description it is evident that, from a reptilian
standpoint, the viviparity of Cophotis is very complete. In this
connection it may be noted that the wall of the oviduct is also
well-supplied with blood-vessels, a large trunk proceeding along
the mesometric line. It seems almost certain that gaseous and
probably also liquid transfusion from parent to embryo takes place
through the thin intervening membranes. It is not always easy
to’ suggest an explanation for the exceptional occurrence of
viviparity in groups of normally oviparous animals. It seems to
depend upon various conditions which may react upon the
organism separately or in combination. Of these conditions the
most obvious are habits and habitat (whether arboreal, deserti-
colous, subterranean), temperature and moisture. There are
Embryo of Cophotis ceylanic« lying in a dilatation of the oviduct. The
dotted portion indicates the yolk. At each side of the figure is the cut end
of the oviduct.
‘several parallel examples which may help to throw light on this
subject. Cophotis is arboreal as shown bionomically by its actual
occurrence as well as structurally by its prehensile tail, but less so
than the Chameleon which has modified hands and feet in addition
to its prehensile tail. Nevertheless most Chameleons lay eggs, an
exception being afforded by Chameleon pumilus, the Dwarf
Chameleon of South Africa which is ovoviviparous. This species
is described by Dr. Gadow (Cambridge Nat. Hist. Amphibia and
Reptiles, 1901, p. 580) as “relatively hardy, being as a native of
South Africa, accustomed tocold nights.” In this respect it offers
an analogy to Oophotis ceylanica, which has almost exactly the
same size as the South African Chameleon, namely, 5-6 inches,
and is accustomed to the cold nights of the upper montane zone
of Ceylon. i)
NOTES. 237
Another parallel example is afforded by the Agamoid genus
Phrynocephalus (referred to above) which inhabits the desert
regions of Central Asia, as compared with the viviparous Iguanoid
genus Fhrynosoma* which inhabits the desert regions of Central
America.
In many or most cases of viviparity among reptiles an egg-shell
is formed round the vitellus, but the egg is not laid, the young
hatching out 7m sztwintheoviducts. In Cophotis there isno trace
of an egg-shell.
With regard to Ceratophora the eggs are laid in small holes in
the ground after the manner of Calotes. JI came acrosssuch a hole
containing four freshly-laid, soft-shelled eggs in the Hakgala
jungle in January and disturbed the female who was apparently
attending to it. Ceratophora stoddartii is usually found clinging
to the trunks of trees or the stems of shrubs and saplings in a
vertical attitude with the rostral appendage directed upwards.
This appears to be its normal resting attitude and it remains
motionless for hours together. Its food consists in large part
of earthworms, to obtain which it descends to the ground. The
female descends to the ground also for the purpose of egg-laying,
the mating taking place on the stem of a young or small tree.
The existence of the rostral appendage is, I think, more or less
closely correlated with the vertical resting attitude of the lizard.
In contrast with the Agamidz most of the Scincoid lizards are
viviparous, but an exception occurs in the species, Mabuia
macularia,f which is oviparous. Some eggs which I found in
the month of December at the base of a kumbuk tree at Karawala-
gaswewa, between Puttalam and Anuradhapura, no doubt
belonged to this species. I placed them in a match-box,and within
an hour or two young skinks hatched out which by an oversight I
omitted toidentify. Thespecies however occurs in Ceylonas well
as in India and Burma. It is closély allied to the well known
Brahminy Lizard which is viviparous (Boulenger op. cit., p. 190).
There is an Australian skink, Tvachysaurus,{ which is strictly
viviparous in so far that like Cophotis no egg-shell is formed,
and the embryos “are ripened in uterus-like dilatations of the
oviducts.”
In conclusion, it may be noted that the geckos (Geckonide) are
’ oviparous lizards; “the only species at present known to be
ovoviviparous are the New Zealand Nawltinus elegans and Hoplo-
dactylus pacificus”’ (Boulenger, op. cit., p. 55).
Colombo, February, 1906. A. WILLEY.
* See Gadow, op. cit., p. 533. Tt See Boulenger, op. cit., 180.
$ See Gadow, op. cit., p. 560.
238 SPOLIA ZEYLANICA.
1]. Subsidence at Kokhilay on the East Coast.—The village of
Kokkutoduvay lies on the narrow strip of land which separates
Kokkilay lagoon from the sea and is some half a mile from
the latter. Across a small “ villu”’ from the village and right on
the sands just above highwater mark, where a very moderate
north-east gale would carry the waves, is a large area containing
a deposit of ancient pottery débris, undoubtedly the site of an old
settlement. This, in conjunction with an existing tradition that
Kokkilay lagoon was once astretch of paddy fields, is very strong
evidence of the land having subsided.
The pottery is old—how old it is impossible to say ; perhaps a
thousand years, perhaps more. But the land hereabouts must
have also risen and that long before the time of the potters. For
the coast is strewn with fossils, sea-shells imbedded in sandstone
which was formed I suppose in a considerable depth of water.
Thus at this little village we have evidence, natural and artificial,
of two alterations, in opposite directions, from the present level.
JOHN STILL.
Anuradhapura, April 19, 1905.
12. Association of a crocodile with a tortoise—EKarly in March,
1905, I was at a village named Kanjuramotai, a few miles south of
Nedunkeni in the Mullaittivu District. Two of the villagers
showed me a few ruins in the jungle, just by the edge of their
paddy fields ; and among them was an old well. This had been
dug out six or eight years ago, and then abandoned, after which
the sides had collapsed inwards leaving a hollow about 10 feet
deep, circular in shape, and 12 or 15 feet in diameter. Its sides
are perpendicular save at the very top where they overhang, held
up hy the matted roots of the undergrowth, so that the place is a
regular trap.
In the well there are two captives, both well known to the
villagers : one a crocodile, about six feet long, the other a very
large mud tortoise (Nicoria trijuga). The villagers aver that the
crocodile gotin during the rainy weather three years ago and that
the tortoise has been there for six months. Neither can get food
(except perhaps frogs), neither can get out; and in the dry
weather they must go without water for months on end.
The turtle has no fear of the crocodile, but when stirred up will
walk up to his snout. The crocodile, which is miserably thin,
resents being stirred up by hissing and snapping his jaws, but
NOTES. 239
makes noattack on the tortoise. The villagers say he is afraid of
being left alone if he eats the tortoise, but perhaps the shell is too
much for him in his enfeebled state.
JOHN STILL.
Anuradhapura, March, 1905.
13. Distribution of flying squirrels in Ceylon.—The squirrel
called Pteromys oral which possesses a dermal parachute by means
of which it can travel through the air for limited distances, and on
this account is always called the flying squirrel, belongs to the
Rodent family Sciuride, which is the squirrel family. This
information is given in a condensed form on p. 38 of this volume.
An illustration showing the animal at rest and in flight is given
on p. 361 of Blanford’s Mammalia (Fauna Brit. Ind.) Another
species called the “Small Travancore Flying Squirrel,” Sciwrop-
terus fuscicapillus, is recorded by Dr. Blanford from the hills of
Ceylon, but no information concerning it is available locally.
The Flying Fox (Pteropus medius) which, as stated on p. 38
(this volume), is a Fruit-eating Bat, occurs in the low-country,
forming ‘‘ camps” during the daytime at such places as Beruwala
(Barberyn), Henaratgoda, and Peradeniya. The flying squirrel
also occurs at Peradeniya, whence several examples have been sent
to the Museum by Mr. E. E. Green. Thisis an example of a local
overlapping in distribution, since the flying squirrel hardly
occurs below the elevation of Peradeniya and the flying fox does
not go much above it. Mr. Edgar Turner has kindly written to
say that he has never seen “flying foxes” in the up-country
jungles, but has often seen the “ flying squirrels” in the jungles
up to 5,000 feet or more, particularly in the Hewaheta jungles by
Rookwood estate.
Ed. (5S. Z:
END OF VOLUME III.
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SPOLIA ZEYLANICA.
ISSUED BY
THE COLOMBO MUSEUM,
CHY LON.
VOLUME Iil.
COLOMBO:
G. J. A. SKEEN, GOVERNMENT PRINTER, CEYLON.
—_—
1906.
ss
7 "ind wi
: n / pia
a 4 Lae AS ane
;
mee 3
i.
; ae oF
ero
CONTENTS OF VOLUME IT.
Part [X.—Marcg, 1905.
Guide to the Antiquities, Minerals, and Natural History Collections
bo
in the Colombo Museum
Part X.—OcrosBeEr, 1905.
Cameron, P.—
On the Phytophagous and Parasitic Hymenoptera collected
by Mr. E. Ernest Green in Ceylon (First Paper)
Cameron, P.—
Idem (Gecond Paper)
Wall, F.—
Notes on Snakes collected at Hakgalla, Ceylon
Wait, W. E.—
Notes chiefly on Birds seen at the Pearl Fishery me
March and April, 1905. Se
Coomaraswamy, A. K. and Ethel M.—
Kandyan Horn Combs
Notes.—
1. On the Zoris in captivity. John Still “ct
2. On the larval habits of the Butterfly, Parata alexis, Fabr.
E. E. Green oe vee
3. Curious behaviour of a eenen in apie. E. E. Green..
4. On the nesting of the Snake Bungarus ceylonicus. H. E.
Green nee .
Millipede killed by a Bedagud Bug. E. E. Green ore
Remedies adopted against the Paddy Fly. C. Drieberg...
7. The Lacteal Tract of Loris gracilis. A. Willey
a
Part XI.—Janvuary, 1906.
Linstow, O. von—
Helminthes from the collection of the Colombo Museum...
Annandale, N.—
New and interesting Lizards in the Colombo Museum
Annandale, N.—
Stalked Barnacles Levene Pedunculata) in the Colombo
Museum a ws
Notes.—
1. Lacteal Tract of Oriental Lorisine. N. Annandale
2. Curious action of a Toad when confronted by a Snake.
j EK. E. Green zs
On the constricting habit of Coluber alain EB. B. Green ...
Scorpion stings. KH. EH. Green : one
Minerals new or rare in Ceylon. A. K. Coomaraswamy
oR
PAGE
144
148
151
155
157
157
158
159
159
160
163
189
193
196
196
197
197
198
( Senn
6. Recent Marine Clays at Kuchaveli,Ceylon. A. K. Cooma-
raswamy out a;
7. Contributions to the Geology of Ceylon. A. K. Coomara-
swamy . <P
8. Snake eres C. Daebeneie
9. Snakes and Fowls. C. Drieberg
10. Moths at sea. T. Bainbrigge Fletcher
Part XIT.—AprIL, 1906.
Doflein, F.—
Termite Truffles
Cameron, P. —
Description of a new species of Opius from Ceylon
Notes.—
Historic Trees. J. P. Lewis
The Moormen’s Weapon. J. P. Lewis
Scorpion stings. A. K.Coomaraswamy ...
Flight of white and yellow Butterflies. O.S, Wickwar
Notes by the Way. .E. E. Green ie
On the Species of Leaf Insects occurring in a Geilo: K. E.
Green ...
A Harbour Worm and a Bosine Crab. A ‘Willey
Stridulation of Gongylus gongylodes. A. Willey
Terrestrial Colubride of Ceylon. A. Willey aa
Viviparity of Cophotis ceylanica. A. Willey cs
Subsidence at Kokkilay on the Hast Coast. John Still
Assoclation of a Crocedile with a Tortoise. John Still
Distribution of Flying Squirrelsin Ceylon. Ed. 8. Z.
238
239
SPOLIA ZEYLANICA.
ISSUED BY
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CEYLON.
Vou. IIJ.—Part IX. © MARCH, 1905.
CONTENTS.
Guide to the Antiquities, Minerals, and Natural History Collections
in the Colombo Museum.
With Plates and Illustrations,
[For Rate of Subscription and other Information see back of Cover. ]
; COLOMBO ;
GEORGE J. A. SKEEN, GOVERNMENT PRINTER, CEYLON.
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SPOLIA ZEYLANICA.
A Quarterly Publication designed to promote a knowledge of
the Natural History of Ceylon (exclusive of Botany) for the
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It will contain Records and Contributions, together with Notes,
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ISSUED BY
CEYLON.
SPOLTA ZEYLANICA,
THEHE COLOMBO MUSEUM,
Vou. III.—PArRT X. OCTOBER, 1905.
CONTENTS.
1. Cameron, P.—
On the Phytophagous and Parasitic Hymenoptera collected
by Mr. E. Ernest Green in Ceylon (First Paper)
2. Cameron, P.—
On the Phytophagousand Parasitic Hymenoptera collected
by Mr. E. Ernest Green in Ceylon (Second Paper) ...
3. Wall, F.—
Notes on Snakes collected at Hakgalla, Ceylon
4, Wait, W. E.—
Notes chiefly on Birds seen at the Pearl Fishery Some
March and April, 1905 are Bs
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Kandyan Horn Combs ... ose
6. Notes.—J. Still, E. B. Green, C. Drieberg, A. Willey
With Plates and Illustrations.
COLOMBO :
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SPOLIA ZEYLANICA.
A Quarterly Publication designed to promote a knowledge of
the Natural History of Ceylon (exclusive of Botany) for the
information of residents in the Island, and also for the advance-
ment of Science.
It will contain Records and Contributions, together with Notes,
Abstracts, and Reviews, relating to the economic and systematic
knowledge of the natural resources (Goology, Anthropology,
Geology) of the Island and of the surronnding seas.
Each Volume will consist of four Parts, the size of which will
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Authors will receive 25 copies of their contributions gratis, or
50 copies if desired.
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SPOLTA ZEYLANICA.
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CONTENTS.
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1. Linstow, Dr. O. von—
Helminthes from the collection of the Colombo Museum 163
2. Annandale, N.—
New and interesting Lizards in the Colombo Museum ... 189
3. Annandale, N.—
Stalked Barnacles (Cirripedia Pedunculata) in the
Colombo Museum .., See eee 193
4. Notes.—N. Annandale, E. E. Green, A. K, Coomaraswamy,
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SPOLIA ZEYLANICA.
A Quarterly Publication designed to promote a knowledge of
the Natural History of Ceylon (exclusive of Botany) for the
information of residents in the Island, and also for the advance-
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Abstracts, and Reviews, relating to the economic and systematic
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Topography, Geology) of the Island and of the surrounding seas.
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from Mr. Bernard Quaritch, 15, Piccadilly, London; and from
Messrs. Wyman & Sons, Ltd., Fetter Lane, London, E.C.
SPOLIA ZEYLANICA.
ISSUED BY
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CEYLON.
Vou. I1].—Part XII. APRIL, 1906.
CONTENTS.
PAGE
Doflein, Dr. F. (Munich)—
Termite Truffics ase Abe ae 203
Cameron, P.—
Description of a new species of “‘ Opius” from Ceylon... 210
Notes.—J. P. Lewis, A. K. Coomaraswamy, E, H. Green,
O. S. Wickwar, A. Willey, J, Still me eee 211
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SPOLIA ZEYLANICA.
A Quarterly Publication designed to promote a knowledge of
the Natural History of Ceylon (exclusive of Botany) for the
information of residents in the Island, and also for the advance-
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It will contain Records and Contributions, together with Notes,
Abstracts, and Reviews, relating to the economic and systematic
knowledge of the natural resources (Zoology, Anthropology,
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