A Stereo-Atlas of Ostracod Shells
edited by J. Athersuch, D. J. Horne, D. J. Siveter,
and J. E. Whittaker
Volume 16, Part 2; 31st December, 1989
V
Published by the British Micropalaeontological Society, London
ISSN 0952-7451
Editors
Dr J. Athersuch, Stratigraphy Branch, The British Petroleum Co, BP Research Centre, Chertsey Road,
Sunbury-on-Thames, Middlesex TW16 7LN.
Dr D. J. Horne, School of Earth Sciences, Thames Polytechnic, Walburgh House, Bigland Street,
London El 2NG.
Dr David J. Siveter, Department of Geology, The University, Leicester LEI 7RH.
Dr J. E. Whittaker, Department of Palaeontology, British Museum (Natural History), Cromwell Road,
London SW7 5BD.
Editorial Board
Dr J.-P. Colin, Esso Production Research - European, 213 Cours Victor Hugo, 33321 Begles, France.
Dr P. De Deckker, Department of Geology, Australian National University, G.P.O. Box 4, Canberra,
Act 2601, Australia.
Dr D. van Harten, Universiteit van Amsterdam, Geologisch Instituut, Nieuwe Prinsengracht 130,
Amsterdam, The Netherlands.
Dr W. Hansch, Ernst-Moritz-Arndt Universitat, Sektion Geologische Wissenschaften, F.-L.-Jahnstr.
17a, 2200 Greifswald, German Democratic Republic.
Dr R. E. L. Schallreuter, Universitat Hamburg, Geologisch-Palaontologisches Institut, Bundesstrasse
55, D 2000 Hamburg 13, German Federal Republic.
Dr Zhao Yuhong, Nanjing Institute of Geology & Palaeontology, Academia Sinica, Chi-Ming-Ssu,
Nanjing, People’s Republic of China.
Officers of the British Micropalaeontological Society
Chairman Dr A. C. Higgins, BP Research Centre, Chertsey Road, Sunbury-on-Thames, Middlesex
TW16 7LN.
Secretary Dr J. B. Riding, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Treasurer Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History),
Cromwell Road, London SW7 5BD.
Journal Editor Dr M. Keen, Department of Geology, The University of Glasgow G12 8QQ.
Newsletter Editor Dr D. J. Shipp, Robertson Research International, Ty’n-y-Coed, Llanrhos,
Llandudno, Gwynedd LL30 ISA.
Conodont Group Chairman Dr P. M. Smith, Department of Earth Sciences, University of Cambridge,
Downing Street, Cambridge CB2 3EQ.
Conodont Group Secretary Mr A. Swift, Geology Department, University of Nottingham NG7 2RD.
Foraminifera Group Chairman Dr A. A. H. Wonders, B.P. Research Centre, Chertsey Road,
Sunbury-on-Thames, Middlesex TW16 7LN.
Foraminifera Group Secretary Dr F. M. D. Lowry, Department of Geology (Micropalaeontology),
University College, Gower Street, London WC1E 6BT.
Microplankton Group Chairman Dr G. L. Eaton, B.P. Research Centre, Chertsey Road, Sunbury-on-
Thames, Middlesex TW16 7LN.
Microplankton Group Secretary Dr A. J. Powell, B.P. Research Centre, Chertsey Road, Sunbury-on-
Thames, Middlesex TW16 7LN.
Ostracod Group Chairman Dr J. Athersuch, B.P. Research Centre, Chertsey Road, Sunbury-on-
Thames, Middlesex TW16 7LN.
Ostracod Group Secretary Dr N. G. Fuller, Phillips Petroleum Company United Kingdom Limited,
Petroleum Products Division, Phillips Quadrant, 35 Guildford Road, Woking, Surrey GU22 7QT.
Palynology Group Chairman Dr D. J. Batten, Department of Geology, Marischal College, University of
Aberdeen, Aberdeen AB9 IAS.
Palynology Group Secretary Dr J. E. A. Marshall, Department of Geology, The University,
Southampton S09 5NH.
Calcareous Nannofossil Group Chairman Mr M. Jakubowski, Robertson Research International,
Ty'n-y-Coed, Llanrhos, Llandudno, Gwynedd LIAO ISA.
Calcareous Nannofossil Group Secretary Dr J. Crux, B.P. Research Centre, Chertsey Road,
Sunbury-on-Thames, Middlesex TW16 7LN.
Instructions to Authors
Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited.
Format should follow the style set by the papers in this issue. Descriptive matter apart from illustrations
should be cut to a minimum; preferably each plate should be accompanied by one page of text only.
Blanks to aid in mounting figures for plates may be obtained from any one of the Editors or Editorial
Board. Completed papers should be sent to Dr David J. Siveter.
The front cover shows a female left valve (OS 13377) of Bromidella reticulata Harris from the Simpson
Group, middle Ordovician, Oklahoma, USA (see M. Williams & D. J. Siveter, Stereo-Atlas Ostracod
Shells, 16, 1-8, 1989).
Printed in the UK by BPCC BLACKPOOL Ltd., Stanley Road, Blackpool FY1 4QN
A Member of BPCC Ltd
A Stereo-Atlas of Ostracod Shells
edited by J. Athersuch, D. J. Horne, D. J. Siveter
and J. E. Whittaker
Volume 16, 1989
Part 1 (pp.1-77); 31st July, 1989
Part 2 (pp. 78-157); 31st December, 1989
Published by the British Micropalaeontological Society, London
Stereo-Atlas of Ostracod Shells 16, ii
Contents
Contents
1 On Bromidella reticulata (Harris); by M. Williams & D. J. Siveter 1
2 On Lophocypris shulanensis Zhang & Zhao gen. et sp. nov. ; by Zhang Lijun & Zhao Yuhong 9
3 On Dabashanella retroswinga Huo, Shu & Fu ; by Zhao Yuhong & Tong Haowen 13
4 On Progonocythere levigata Bate; by M. I. Wakefield & D. J. Siveter 17
5 On Bythoceratina gobanensis Reyment & Reyment sp. nov. ; by R. A. Reyment & E. R. Reyment 21
6 On Fallaticella schaeferi Schallreuter ; by R. E. L. Schallreuter 25
7 On Columatia variolata (Jones & Holl); by R. F. Lundin & D. J. Siveter 29
8 On Microcheilinella distorta (Geis); by R. F. Lundin 35
9 On Sinessites hispanicus Becker; by G. Becker 39
10 On Kullmannissites kullmanni Becker; by G. Becker 43
11 On Vitissites comtei Becker; by G. Becker 47
12 On Rishona epicypha (Kesling & Kilgore); by G. Becker & F. Adamczak 51
13 On Chinocythere curvispinata Su sp. nov. ; by Su Deying 55
14 On Chinocythere shajingensis Su sp. nov. ; by Su Deying 59
15 On Chinocythere tuberculata Su sp. nov. ; by Su Deying 63
16 On Tuberoloxoconcha tuberosa (Hartmann); by D. J. Horne 67
17 On Tuberoloxoconcha atlantica Horne sp. nov. ; by D. J. Horne 73
18 On Buntonia brunensis Riha; by J. Riha 77
19 On Primitivothlipsurella v-scripta (Jones & Holl); by R. F. Lundin & L. E. Petersen 78
20 On Primitivothlipsurella obtusa Petersen & Lundin sp. nov. ; by L. E. Petersen & R. F. Lundin 86
21 On Balticella deckeri (Harris); by M. Williams & D. J. Siveter 94
22 On Macrypsilon salterianum (Jones); by D. J. Siveter & W. Hansch 100
23 On Berolinella steusloffi (Krause) ; by W. Hansch & D. J. Siveter 106
24 On Aurikirkbya wordensis (Hamilton); by G. Becker & F. Adamczak 112
25 On Nodella hamata Becker; by G. Becker 116
26 On Cytheridea sandbergeri Kammerer sp. nov. ; by T. Kammerer 120
27 On Strandesia weberi (Moniez); by D. Keyser & S. B. Bhatia 128
28 On Abyssobythere guttata Ayress & Whatley gen. et sp. nov. ; by M. Ayress & R. C. Whatley 136
29 On Bryocypris grandipes R0en ; by K. Martens 140
30 On Limnocythere hibernica Athersuch sp. nov. ; by J. Athersuch 148
31 On Echinocythereis spinireticulata Kontrovitz; by M. Kontrovitz & Zhao Yuhong 152
32 Index for Volume 16, 1989 156
Stereo-Atlas of Ostracod Shells 16 ( 19) 78-85 ( 1989)
595.337.23 (113.331) (420 : 161.003.52 + 485 : 161.018.57) : 551.351 + 552.54
Primitivothlipsurella v-scripta (1 of 8)
ON PRIMITIVOTHLIPSURELLA V-SCRIPTA (JONES & HOLL)
by Robert F. Lundin & Lee E. Petersen
(Arizona State University , Tempe, U.S.A. & Anadarko Petroleum Corporation , Houston, USA)
Derivation of name:
Diagnosis:
Remarks:
Genus PRIMITIVOTHLIPSURELLA gen. nov.
Type-species: Thlipsura v-scripta Jones & Holl, 1869
Latin primitivus , early, and Thlipsurella , indicating the genus is ancestral to Thlipsurella Swartz.
1932.
Thlipsuridae with one vertical or subvertical preadductorial sulcus and two oblique posterior,
straight to slightly curved sulci which form a variable but distinctly acute angle with each other and
are bounded posteriorly by a distinct lobe which approximately parallels the posterior border of
the carapace. Adductor muscle attachment marked by large subcircular depression at mid-length
slightly above mid-height on interior surface of valves. Hinge distinctly inclined to longitudinal
axis of valves.
The type -species of Primitivothlipsurella is considered to be a direct descendant of P. obtusa
Petersen & Lundin ( Stereo-Atlas Ostracod Shells, 16, 86-93, 1989). This relationship clearly
suggests that the genus is ancestral to Thlipsurella Swartz.
Primitivothlipsurella is distinguished from Thlipsurella by the arrangement of the sulci,
especially the posterior ones. Nevertheless, the two genera show distinct similarities in the basic
shape and sculpture of the valves. Hingement and other interior structures in the type-species of
Thlipsurella , T. ellipsoclefta Swartz, 1932, are not known. However, Lundin’s ( Okla . Geol. Surv.
Bull., 116, 85-87, pi. 17, fig. 2, 1968) description of T. putea Coryell & Cuskley, 1934, which is
closely related to the type-species, indicates that the hinge and contact margin structures of the
Explanation of Plate 16, 79
Figs. 1, 2, car. (ASU X-109, 921 p.m long): fig. 1, ext. rt. It. ; fig. 2, ext. It. lat. Figs. 3, 4, car. (ASU X-116, 865 /am long): fig. 3, ext.
vent.; fig. 4, ext. rt. lat. Scale A (200/u.m; x 65), figs. 1, 2; scale B (200/i.m; x 69), figs. 3, 4.
Stereo-Atlas of Ostracod Shells 16, 80 Primitivothlipsurella v-scripta (3 of 8)
two genera are similar. However, the place of the adductor muscle attachment in Thlipsurella is
represented exteriorly by an adductorial sulcus. The new genus presented here has no S2, but
rather the typical thlipsurid characteristic of a circular depression on the interior surface at the
place of adductor muscle attachment. Swartz’s ( J . Paleont. 6, pi. 10, fig. 6c, 1932) illustration of a
juvenile of the type-species of Thlipsurella, as well as variations observed in and comparison of P.
v-scripta and P. obtusa, suggest a phylogeny in which the posterior sulci became separated and
more parallel with time.
Primitivothlipsurella is presently known only from P. v-scripta (Jones & Holl) and P. obtusa
Petersen & Lundin, both from the Silurian strata of the Welsh Borderland area of Britain.
Primitivothlipsurella v-scripta (Jones & Holl, 1869)
1869 Thlipsura v-scripta sp. nov. T. R. Jones & H. B. Holl, Ann. Mag. nat. Hist., (4), 3, 214, pi. 15, figs. 3a-c.
1887 Thlipsura v-scripta war. discrete nov. T. R. Jones. Notes on some Silurian ostracoda from Gothland, Stockholm, 6 ( nom . nud.).
1887 Thlipsura v-scripta Jones & Holl; T. R. Jones, Ann. Mag. nat. Hist., (5), 19, 403.
1887 Octonaria octoformis var. informis nov. T. R. Jones, Ann. Mag. nat. Hist., (5), 19. 405, pi. 12, figs. 5a, b.
1888 Thlipsura v-scripta Jones & Holl var. discrete Jones; T. R. Jones, Ann. Mag. nat. Hist., (6) 1, 404, pi. 22, figs. 9a-c, 10.
1919 Thlipsura v-scripta var. discreta Jones; J. E. Hede, Geol. For. Stock. Forh., 41, 139, 147, pi. 6, fig. 1.
1932 Thlipsurella v-scripta (Jones & Holl); F. M. Swartz, J. Paleont., 6, 44, pi. 10, fig. 7.
1956 Thlipsurella discreta (Jones); A. Martinsson, Publ. Palaeontol. Inst. Univ. Uppsala, 14, 33, pi. 5, figs. 43-49.
1965 Thlipsurella discreta (Jones); V. Pokorny, Principles Zool. Micropalaeontol. , 229, fig. 852, Pergamon Press, Oxford.
1966 Thlipsurella discreta (Jones); F. J. Adamczak, Geol. For. Stock. Forh., 88, 466, fig. 5.
1968 Thlipsurella v-scripta (Jones & Holl); V. S. Krandijevsky, Paleont. & Stratigr. of the Lower Palaeozoic of Volyn-Podolia,
Acad. Nauk Ukr. SSR, 70, pi. 11, fig. 11.
1968 Octonaria informis Jones; V. S. Krandijevsky, Paleontol. & Stratigr. of the Lower Palaeozoic of Volyn-Podolia, Acad. Nauk
Ukr. SSR, 74.
1978 “ Thlipsura ” v-scripta Jones & Holl; D. J. Siveter, in: R. H. Bate & E. Robinson (eds.), A Stratigraphical Index of British
Ostracoda, Geol. J. Spec. Issue, 8, 74, pi. 3, figs. 1, 2, tab. 3.
Explanation of Plate 16, 81
Figs. 1, 2, juv. car. (ASU X-117, 771 gm long): fig. 1, ext. rt. lat. ; fig. 2, ext. It. lat. Fig. 3, juv. car., ext. rt. lat. (ASU X-118, 808 gm
long). Scale A (200/xm; X 78), figs. 1, 2; scale B (200/^m; x 74), fig. 3.
Stereo-Atias of Ostracod Shells 16, 79
Primitivothlipsurella v-scripta (2 of 8)
Stereo-Atlas of Ostracod Shells 16, 81
Primitivothlipsurella v-scripta (4 of 8)
Stereo-Atlas of Ostracod Shells 16, 82 Primitivothlipsurella v-scripta (5 of 8)
1981 “ Thlipsura ” v-scripta Jones & Holl; R. J. Aldridge, K. J. Doming & D. J. Siveter, in: J. W. Neale & M. Brasier (Eds.),
Microfossils from Recent & Fossil Shelf Seas , 22, 28, pi. 2.3, fig. 17, Ellis Horwood, Chichester.
1984 “ Thlipsura ” v-scripta Jones & Holl; D. J. Siveter, Spec. Pap. Palaeontol. , 32, 81, text-fig. 3:8.
1987 “ Thlipsurella ” v-scripta (Jones & Holl); L. E. Petersen & R. F. Lundin, J. Micropalaeontol. , 6, 80, pi. 1. fig. 1 (authorship
given on pi. 1, fig. 1 is in error).
Lectotype : Designated herein. British Museum (Nat. Hist.) no. I 2078; juvenile left valve. Specimen
Fig. 1, car. ext. It. lat. (ASU X-lll, 996 pm long): fig. 2, RV, int. lat. (ASU X-112, 996 pm long); fig. 3, LV, int. lat. (ASU X-113.
940 pm long). Scale A (2(XVm; X 62), fig. 1; scale B (200/xm; x 60), fig. 2; scale C (200/um; x 65), fig. 3.
Stereo-Atlas of Ostracod Shells 16, 84 Primitivothlipsurella v-scripta (7 of 8)
Figs. 1, 2, car. (ASU X-114, 1034 pm long): fig. 1, ext. dors.; fig. 2, ext. vent. Fig. 3, car., ext. vent. (ASU X-115, 1015/j.m long). Fig.
4, juv. LV, ext. lat. (lectotype BMNH I 2078, 763 p,m long). Fig. 5, LV, ext. lat. (ASU X-110, 1128yu.m long). Fig. 6, car., ext. rt.
lat. (ASU X-lll, 996 yum long).
Scale A (200 /urn; x 59), figs. 1, 2; scale B (200 yum ; x 60), fig. 3; scale C (200 yum; X 72), fig. 4; scale D (200 yum; x 54), fig. 5; scale E
(200 yum; x62), fig. 6.
Stereo-Atlas of Ostracod Shells 16, 85 Primitivothlipsurella v-scripta (8 of 8)
Stereo-Atlas of Ostracod Shells 16 (20) 86-93 (1989) Primitivothlipsurella obtusa (1 of 8)
595.337.23 (113.331) (420 : 162.003.52) : 551.351 + 552.52
ON PRIMITIVOTHLIPSURELLA OBTUSA PETERSEN & LUNDIN sp. nov.
by Lee E. Petersen & Robert F. Lundin
(Anadarko Petroleum Corp., Houston & Arizona State University, Tempe, USA)
Primitivothlipsurella obtusa sp. nov.
Holotype:
Type locality:
Derivation of name:
Department of Geology, Arizona State University (ASU), USA, no. ASU X-119; carapace.
The north bank of the River Severn opposite Buildwas Abbey, Buildwas, Shropshire, England
(National Grid Reference, SJ 6435 0450); approximately lat. 52°39' N, long. 2°33' W. The sample
is from the upper (but not the uppermost) part of the Buildwas Formation, late early
Sheinwoodian Stage, Wenlock Series, Silurian.
Latin obtusa ; referring to the orientation of the posteroventral sulcus relative to the longitudinal
axis of the valve.
Explanation of Plate 16, 87
Figs. 1-4, car. (holotype ASU X-119, 789 long): fig. 1, ext. rt. lat.; fig. 2, ext. dors.; fig. 3, ext. vent.; fig. 4, ext. It. lat.
Scale (200 gm; x77), figs. 1-4.
Stereo- Atlas of Ostracod Shells 16, 88 Primitivothlipsurella obtusa (3 of 8)
Figured specimens: Department of Geology, Arizona State University (ASU) nos. X-119 (holotype, car.: PI. 16, 87,
figs. 1-4), X-120 (paratype, car.: PI. 16, 89, figs. 1-4), X-122 (paratype, car.: PI. 16, 91, figs.
1-3), X-123 (paratype, car.: PI. 16, 93, fig. 1), X-121 (paratype, juvenile car.: PI. 16. 93,
figs. 2, 3).
X-121 is from the Buildwas Formation, sample approximately lm higher in the section than
the sample yielding the holotype. All of the other figured specimens are from the same sample as
the holotype.
Diagnosis: Primitivothlipsurella species in which the posterior lobe of each valve forms or is very close to the
posterior border of the valve. Posteroventral sulci oriented at distinct angle to the ventral border
of the carapace.
Explanation of Plate 16, 89
Figs. 1-4, car. (ASU X-120, 865 gm long): fig. 1, ext. rt. lat.; fig. 2, ext. dors.; fig. 3, ext. vent.; fig. 4, ext. It. lat.
Scale (200/xm; x71), figs. 1-4.
Primitivothlipsurella obtusa (2 of 8)
Stereo-Atlas of Ostracod Shells 16, 87
Stereo-Atlas of Ostracod Shells 16, 89
Primitivothlipsurella obtusa (4 of 8)
Stereo-Atlas of Ostracod Shells 16, 90 Primitivothlipsurella obtusa (5 of 8)
Remarks: Primitivothlipsurella obtusa is ancestral to the type species Primitivothlipsurella v-scripta (Jones &
Holl, 1869) (see Lundin & Petersen, Stereo-Atlas Ostracod Shells , 16, 78-85, 1989) and is
distinguished from the latter species by: its smaller size; the fact that the posterior lobes form or
are nearly coincident with the posterior border of the carapace; and the fact that normally the
posteroventral sulcus is orientated at a more distinct angle to the ventral border of the carapace
than it is in P. v-scripta. The posterior sulci are fused to form a continuous v-shaped sulcus on
some valves (especially right valves) and this fusion seems to be more common in P. obtusa than in
P. v-scripta. This suggests a trend toward separation and more parallel alignment of the posterior
sulci through time. If this is true, we can speculate that P. obtusa was derived from a species such
as Thlipsuroides walensis (Krandijevsky, 1963).
P. obtusa is a diagnostic species for recognition of lower Sheinwoodian strata in the type
Wenlock Series in the Welsh Borderland.
Distribution: Known from the Buildwas and Coalbrookdale formations in the type Wenlock area. Shropshire, in
the Welsh borderland; in strata ranging from the upper Cyrtograptus centrifugus Biozone through
the lower Monograptus riccartonensis Biozone; lower Sheinwoodian Stage, Wenlock Series,
Silurian.
Acknowledgements : R. F. L. acknowledges support of the College of Liberal Arts and Sciences, Arizona State
University, the National Science Foundation (Grant No. EAR-8200816) and NATO. D. J.
Siveter is thanked for help with fieldwork.
Explanation of Plate 16, 91
Figs. 1-3, car. (ASU X-122, 789 yum long): fig. 1, ext. dors.; fig. 2, ext. rt. lat.; fig. 3, ext. It. lat.
Scale (200 yum; x77), figs. 1-3.
Stereo-Atlas of Ostracod Shells 16, 92 Primitivothlipsurella obtusa (7 of 8)
700 720 740 760 780 800 820
Length, Jim
Text-fig. 1 : Size dispersion diagram for 17 right valves of P. obtusa from the type locality.
Explanation of Plate 16, 93
Fig. 1, car. ext. rt. lat. (ASU X-123, 714yum long): figs. 2, 3, juvenile car. (ASU X-121, 620 yum long): fig. 2, ext. rt. lat.;
fig. 3, ext. It. lat.
Scale A (200 yum; x84), fig. 1; scale B (200 yum; x 97), figs. 2, 3.
Stereo -Atlas of Ostracod Shells 16,91
Primitivothlipsurella obtusa (6 of 8)
Stereo-Atlas of Ostracod Shells 16, 93
Primitivothlipsurella obtusa (8 of 8)
Stereo-Atlas of Ostracod Shells 16 (21) 94-99 (1989) Balticella deckeri (1 of 6)
595.337.2 (113.312) (755 : 162 + 766 : 162.097.34) : 551.351 + 552.54
ON BALTICELLA DECKERI (HARRIS)
by Mark Williams & David J. Siveter
(University of Leicester, England)
Balticella deckeri (Harris, 1931)
1931 Leperditella ? deckeri n. sp., R. W. Harris, in C. E. Decker, Okla. Geol. Surv. Bull., 55. 89, pi. 14, figs. 5a-c.
1934 Leperditella ? deckeri Harris; R. S. Bassler & B. Kellet, Geol. Soc. Am. Spec. Pap., 1, 373.
1957a Balticella deckeri (Harris); R. W. Harris, Okla. Geol. Surv. Bull., 75, 242, pi. 8, figs. 7a-c.
1957b Balticella deckeri subsp. elongata n. subsp., R. W. Harris, Okla. Geol. Surv. Bull., 75, 242, pi. 8, fig. 8.
1962 Balticella deckeri (Harris); J. C. Kraft, Geol. Soc. Am. Mem., 86, 57-58, pi. 13, figs. 16, 17, pi. 14, figs. 1-10, text-figs. lOf-h.
1968 Balticella deckeri elongata Harris; R. E. L. Schallreuter, Wissensch. Zeitskr. Der Ernst Moritz -Arndt- Univ. Greifswald. 17.
Mathemat.-Naturwissensch. Reihe, 1/2, 135.
Holotype: The holotype is in the collections of the Museum of Comparative Zoology, Harvard University,
USA, but without a reference number. The slide containing the holotype refers to the original
figures of Harris (1931, pi. 14, figs. 5a-c). This specimen was refigured by Harris (1957a, pi. 8,
figs. 7a-c). In neither publication did Harris refer his type specimen to published figures, or
mention its museum reference number.
Type locality: From the top of Decker's zone 24 (see Harris, 1957), Bromide Formation; approximately 18m
below the top of the Simpson Group, Ordovician. US Highway 99 (Sec. 11, T. Is, R3E), 3 km S of
Explanation of Plate 16, 95
Fig. 1, car. RV, ext. lat. (MCZ unnumbered holotype, 1.47 mm long); fig. 2, car. RV, ext. lat. (MCZ4636, 1.40mm long); fig. 3, juv.
LV, ext. lat. (OS13427, 1.27mm long); fig. 4, juv. LV, ext. lat. (OS13438, 0.94mm long); fig. 5, juv. LV, ext. lat. (OS13439.
0.72 mm long).
Scale A (250 /xm; x37), fig. 1; scale B (250 /urn; x44), fig. 2; scale C (250/^m; x40), fig. 3; scale D (200 ^m; x45), fig. 4; scale E
(200/u.m; x57), fig. 5.
Stereo-Atlas of Ostracod Shells 16, 96 Balticella decked (5 of 6)
Fittstown, Arbuckle Mountains, Oklahoma, USA; approximately latitude 34°35'N, longitude
96° 41' W.
Museum of Comparative Zoology (MCZ), Flarvard University, USA, unnumbered specimen
(holotype car.: PI. 16, 95, fig. 1), MCZ4636 (car.: PI. 16, 95, fig. 2). British Museum (Nat. Hist.)
nos. OS 13427 (juv. LV: PI. 16, 95, fig. 3), OS13438 (juv. LV: PI. 16, 95, fig. 4), OS13439 (juv. LV:
PI. 16, 95, fig. 5), OS13428 (LV: PI. 16, 97, fig. 1), OS13425 (LV: PI. 16. 97, fig. 2, 3), OS13426
(car.: PI. 16, 97, fig. 4), OS13430 (car.: PI. 16, 97, fig. 5), OS13429 (RV: PI. 16, 97, fig. 6).
MCZ4636 is from Decker's Zone 36, Mountain Lake Member, Bromide Formation, H99
Section, Arbuckle Mountains, Oklahoma. OS13430 is from the Mountain Lake Member, Bromide
Formation, at Rock Crossing, Criner Hills, Oklahoma. All other figured specimens are from the
Edinburg Limestone, middle Ordovician, Virginia.
Bisulcate balticellid; adductorial sulcus deep, preadductorial sulcus well developed. Preadducto-
rial node pronounced, sometimes with a nosterodorsally directed spine. Anterior lobe moderately
well developed. Valve surface smooth. No acroidal spines.
Harris (1957b, 242) considered his new taxon Balticella deckeri elongata to be a seperate
subspecies of Balticella deckeri primarily because of its more elongate carapace and its apparently
straighter and longer ventral overlap. Harris (1957b) also considered B. deckeri elongata to be
restricted to the Tulip Greek Formation of the Simpson Group, and to be ancestral to B. deckeri
which he considered exclusive to the Bromide Formation. Harris' referal of the distribution of B.
deckeri elongata to the Tulip Creek Formation is incorrect, as this was based on the Highway 99
Simpson Group section. Reappraisal of the stratigraphy of this section by Fay & Grafham (Univ.
Kansas Paleontol. Contrib. Monograph 1, 14, 1982) and by us shows that the sequence from which
Harris collected his balticellids at Highway 99 in fact represents the lower member of the Bromide
Formation. Balticellid ostracodes are thus restricted in Oklahoma to the Bromide Formation,
Simpson Group, and do not occur in the Tulip Creek Formation.
Explanation of Plate 16, 97
Fig. 1, LV, ext. lat. (OS13428, 1.83 mm long) ; figs 2, 3, LV (OS13425, 1.6 mm long); fig. 2, detail of stop pegs; fig. 3, int. lat. ; fig. 4,
car. vent. (OS13425, 1.61 mm long); fig. 5, car. dors. (OS13430, 1.81 mm long; fig. 6, RV. int. lat. (OS13429, 1.40mm long).
Scale A (250/u.m; x33), fig. 1; scale B (100/u.m x60), fig. 2; scale C (250 ^m x32), figs. 3, 4; scale D (250 /i.m x32), figs. 5, 6.
Figured specimens:
Diagnosis:
Remarks:
Balticella deckeri (2 of 6)
Stereo-Atlas of Ostracod Shells 16, 97
Stereo-Atlas of Ostracod Shells 16, 95
Balticella deckeri (4 of 6)
Stereo-Atlas of Ostracod Shells 16. 98 Balticella deckeri (5 of 6)
Text-fig. 1. Length-height ratios for right valves of all specimens of B. deckeri recovered from the
Bromide Formation and some conspecific material from the Edinburg Limestone of Virginia (dotted).
Holotvpes: B. deckeri elongata of Harris (HE) and B. deckeri Harris (HD).
Remarks (contd.) Studies of length - height ratios for assemblages of balticellids which we have recovered from
Oklahoma also convince us that B. deckeri and B. deckeri elongata should be treated as a single
taxon (Text-figs. 1, 2). Both holotypes plot close together and well within the range of variation
encountered in the assemblages studied. A single histogram based on all specimens of B. deckeri
recovered from the Bromide Formation, together with additional specimens from the Edinburg
Limestone of Virginia (Text-fig. 1), clearly shows that no major difference in the degree of valve
elongation exists between B. deckeri and B. deckeri elongqta. There is also no difference in the
overlap conditions of the valves in the Balticella specimens that we have studied from Oklahoma.
B. deckeri is very similar to the Swedish type-species Balticella oblonga (Thorslund, Sver.
Geol. Unders. Ser. C., 436, 179, pi. 1, figs. 18-20, 1940), differing only in the more pronounced
anterior lobe, the spine on the preadductorial node, and the lack of fine tuberculate ornament.
The ontogeny of B. deckeri is figured for the first time herein (PI. 16, 95). At least four moult
stages are recognised. The spine on the preadductorial node appears to occur in all juvenile stages,
Stereo- Atlas of Ostracod Shells 16, 99 Balticella deckeri (6 of 6)
while the anterior lobe and the dorsal inflation of the posterior lobal area become more
pronounced during ontogeny. Dimorphism is recognised in other species of Balticella (Schallreu-
ter, 1968 op. cit.) but has not been recognised in our specimens of B. deckeri.
Distribution: B. deckeri occurs in the upper part of the Mountain Lake and Pooleville members of the Bromide
formation (Whiterockian-early Mohawkian), Simpson Group, middle Ordovician, Oklahoma,
USA. Also known from the Edinburg Limestone, middle Ordovician, Virginia, USA (Kraft,
1962, op. cit.).
Text-fig. 2. Length-height ratio for right valves of B. deckeri recovered from two assemblages from the
Bromide Formation. A, from the Pooleville Member; B, from the Mountain Lake Member. Holotypes:
B. deckeri elongata of Harris (HE) and B. deckeri of Harris (HD).
Stereo-Atlas of Ostracod Shells 16 (22) 100-105 (1989) Macrypsilon salterianum (1 of 6)
595.336.11 (113.333) (430.2 : 161.013.53 + 013.54 + 438 : 161.017.51) : 551.351 + 552.54
ON MACRYPSILON SALTERIANUM (JONES)
by David J. Siveter & Wolfgang Hansch
( University of Leicester, England & University of Greifswald, GDR)
Genus MACRYPSILON Martinsson, 1962
Type-species (by original designation): Beyrichia salteriana Jones, 1855
Diagnosis: Amphitoxotidinae with very broad lobes. In tecnomorphs the more or less distinctly developed, narrow
prenodal and adductorial sulci are united below the preadductorial lobe into a sulcus. Velum forms a narrow,
tubulous flange extending between the anterior and the posterior cardinal cormers. Crumina large,
subrounded or more elongated posteroventrally with subcruminal velar edge passing over the crumina but
separated from the postcruminal part of the velum. Lobes reticulate to smooth.
Macrypsilon salterianum (Jones, 1855)
Beyrichia salteriana nov. sp. T. R. Jones, Ann. Mag. nat. Hist., ser. 2, 16. 89, pi. 5, figs 15, 16.
Beyrichia salteriana Jones; E. Boll, Arch. Ver. Freunde Nat. Mecklenburg, 16 (7), 135, pi. 1, fig. 12.
Beyrichia salteriana Jones; A. Krause, Z. Dt. Geol. Ges., 29 (1), 35, pi. 1, fig. 17.
Beyrichia salteriana Jones; F. Roemer, Pal. Abh., 2 (5), 109, fig. 356.
Beyrichia salteriana Jones; G. Reuter, Z. Dt. Geol. Ges., 37 (4), 645, pi. 26, figs. 19a, b (collection is lost).
Beyrichia salteriana Jones; M. Verworn, Ibid. , 39 (1), 31, pi. 3, figs. 8, 10 (collection is lost).
Beyrichia salteriana Jones; K. A. Gronwall, Sver. Geol. Unders., ser. C, no. 170, 19 (4), 18.
Beyrichia salteriana Jones; J. Ch. Moberg & K. A. Gronwall, Lunds Univ. Arsskr., N.F. 5 (1), 7-9, 62.
Explanation of Plate 16, 101
Figs. 1-3, 5: cf RV (I 7099, 1410/xm long): fig. 1, ext. lat. ; fig. 2, ext. vent.; fig. 3, ext. post.; fig. 5, detail of syllobium. Fig. 4,
tecnomorphic RV, ext. lat. (lectotype, I 7100, approx. 910 /xm long). Fig. 6, tecnomorphic LV, ext. lat. (I 7118. 890 pm long).
Figs. 7, 8, Cf RV (SGWG 83/3, 1450/xm long): fig. 7, ext. lat.; fig. 8, ext. vent.
Scale A (200 /xm; x37), figs. 1, 2: scale B (200 pm: X50), fig. 3; scale C (200 /xm; x52), fig. 4; scale D (150 /xm; x70). fig. 5;
scale E (200 /rm; x 50), fig. 6; scale F (300 /xm; x35), figs. 7, 8.
Stereo-Atlas of Ostracod Shells 16, 102 Macrypsilon salterianum (3 of 6)
1957 Neobeyrichia salteriana (Jones); R. V. Kesling & K. J. Rogers, J. Paleont., 31 (5), 1003, tab. 1, pi. 128, figs. 14-18.
1962 Macrypsilon salterianum (Jones); A. Martinsson, Bull. Geol. Inst. Univ. Uppsala, 41, 17, 257, 357, fig. 2D.
1964 Macrypsilon salterianum (Jones); A. Martinsson, Geol. For. Stockh. Forh., 86 (2), 126, 128, 133, 156, 159, fig. 15.
1964 Macrypsilon salterianum (Jones); M. J. Copeland, Bull. Geol. Surv. Can., 117, 5, pi. 1, figs. 4, 5.
1965 Macrypsilon salterianum (Jones); L. Gailite, Izv. Akad. Nauk Latv. SSR, 2 (211), 68.
1966 Macrypsilon salterianum (Jones); D. Kaljo & L. Sarv, Izv. Akad. Nauk Est. SSR, ser. F.- techn. nauk, 2, 279, tab. 1.
1967 Macrypsilon salterianum (Jones); E. Witwicka, Kwart. Geol., 2 (1), 48, pi. 2, figs. 9a-c.
1967 Macrypsilon salterianum (Jones); L. Gailite, in: L. Gailite, M. Rybnikowa & R. Ulste, Stratigrafija, fauna i uslovija
obrazovania silurijskich srednej Pribaltiki, 128, pi. 9, figs. 5a, b, Riga (Zinatne).
1967 Macrypsilon salterianum A. Martinsson, Geol. For. Stockh. Forh., 89 (4), 377.
1968 M. salterianum (Jones); L. Sarv, Ostr. Crasp. Beyr. i Primit. silura Estoni, 28, 98, pi. 9, fig. 1, tabs. 2, 3, Tallinn.
1969 Macrypsilon salterianum (Jones); R. W. L. Shaw, Geol. For. Stockh. Forh., 91 (1), 68, fig. 8.
1970 Macrypsilon salterianum (Jones); A. Pranskevicius, Dokl. Akad. Nauk SSSR. 192 (6), 85.
1970 Macrypsilon salterianum (Jones); L. Sarv, in: D. Kaljo (ed.), Silur Estonii, 158, 169, 299, Tallinn (Valgus).
1971 Macrypsilon salterianum (Jones); L. Sarv, Izv. Akad. Nauk Est. SSR, ser. Chimija-Geol. , 20 (4), 353, 355, tabs. 2, 3.
1971 Macrypsilon salterianum (Jones); R. W. L. Shaw, Palaeontology, 14 (4), 599, pi. 109, figs. 7, 8.
1972 Macrypsilon salterianum (Jones); L. Gailite, Izv. Akad. Nauk Est. SSR, ser. Chimija-Geol., 21 (4), 352.
1972 Macrypsilon salterianum (Jones); A. Pranskevicius, Geol. For. Stockh. Forh., 94 (4), 439, 441.
1972 Macrypsilon salterianum (Jones); A. Pranskevicius, Trudy LitNIGRI, 15, 35, 80, 187, tabs. 4, 5, 7, 11, pi. 10, fig. 2.
1973 Macripsilon [s/c] salterianum (Jones); B. Zbikowska, Acta Geol. Pol., 23 (4), 609, 611, 613-614, 625, tab. 2, pi. 4, figs. 9, 10.
1974 Macrypsilon salterianum (Jones); E. Tomczykowa & E. Witwicka, Bull. Inst. Geol., 276, 59, 61, 69, figs. 2, 3.
1974 Macrypsilon salterianum (Jones); B. Zbikowska, Bull. Akad. Pol. Sci., ser. Sci. de la Terre, 22 (1), 47.
1975 M. salterianum ; A. Pranskevicius, Geol. For. Stockh. Forh., 97 (1), 53-54.
1976 Macrypsilon salterianum (Jones); D. Kaljo & L. Sarv, Izv. Akad. Nauk Est. SSR, ser. Chimija-Geol., 25 (4), 326, 328-329.
1977 M. salterianum: L. Sarv, in: D. Kaljo (ed.), Fazii i fauna Silura Pribaltiki, 161, 164, 169, 173, tab. 1-3, 5, 7, Tallinn.
Explanation of Plate 16, 103
Figs. 1-3, $ car. (SGWG 83/4, 1700 /x m long): fig. 1, ext. lat.; fig. 2, detail of syllobium of RV; fig. 3, ext. vent. Fig. 4, $ LV, detail of
ventral part of crumina (SGWG 83/5, approx. 1540 /xm long). Figs. 5-7: $ LV (SGWG 83/6, 1340 /xm long): fig. 5, ext. post. ; fig.
6, ext. lat. ; fig. 7, ext. vent. Scale A (200 /xm ; X 30), figs. 1,3; scale B (75 /xm ; x 120), fig. 2; scale C (20 /xm ; x 540), fig. 4; scale D
(200 /xm; X 45), fig. 5; scale E (200 /xm; x40), figs. 6, 7.
1855
1862
71877
71885
71885
71887
71897
1909
Stereo-Atlas of Ostracod Shells 16, 103
Macrypsilon salterianum (4 of 6)
Stereo-Atlas of Ostracod Shells 16. 104 Macrypsilon salteriimum (5 of 6)
1977 Macrypsilon salterianum (Jones); M. J. Copeland & J. M. Berdan, Geol. Surv. Canada , Paper 77-1B, pi. 2, 3, figs. 17, 18.
71977 Macrypsilon sp. ; M. J. Copeland & J. M. Berdan. Ibid. , pi. 2, 3, fig. 27.
1977 Macrypsilon salterianum: A. Martinsson, The Silurian- Devonian Boundary, IUGS ser. A, no. 5, 48, 329.
1978 Macrypsilon salterianum: D. J. Siveter. in: R. H. Bate & E. Robinson (eds.). Geol. J. Sp. Iss., 8. 68, 8, pi. 8, figs. 1, 2.
1978 Macrypsilon salterianum (Jones); L. Gailite, in: Stratigrafija fanerozoja Pribaltiki, 13, 16, 18-19, 21, Riga (Zinatne).
1980 Macrypsilon salterianum (Jones); D. J. Siveter. Palaeontogr. Soc. (Monogr.), 133 (556), 54, pi. 10. figs. 8, 13.
1982 Macrypsilon salterianum: L. Sarv. in: Ecostratigraphy of the East Baltic Silurian, 75, Tallinn (Valgus).
1985 Macrypsilon salterianum (Jones); W. Hansch, Lethaia. 18 (4), 375, tab. 1.
1986 M. salterianum (Jones); N. Sidaraviciene. in: D. Kaljo & E. Klaamann (eds.), Teorija opvt ekostratigrafija, 120, 124, Tallinn.
1986 Macrypsilon salterianum (Jones); L. Gailite, Ibid. , 114.
1989 M. salterianum (Jones); D. Siveter. in: C. Holland & M. Bassett (eds.). Global standard for the Silurian, fig. 1681. Nat. Mus.
Wales 9. Cardiff.
Lectotype: British Museum (Nat. Hist.), no. 17100; tecnomorphic RV. Martinsson, 1962, fig. 2D.
[Paratypes: British Museum (Nat. Hist.) 17118, tecnomorphic LV ; Jones, 1855, pi. 5, fig. 16,
17099, cf RV ; Jones. 1855, pi. 5, fig. 15a, b.]
Type locality: Erratic boulder no. 5 of Jones. 1855. near Breslau ( Wrowtaw), Poland; approx, lat. 51°5' N, long.
17°E. Upper Silurian.
Figured specimens: British Museum (Nat. Hist.) nos. 17100 (lectotype, tecnomorphic RV: PI. 16, 101, fig. 4), 17099 (paratype,
Cf RV: PI. 16. 101. figs. 1-3. 5). 17118 (paratype, tecnomorphic LV: PI. 16, 101, fig. 6). All from erratic
boulder no. 5 of Jones. 1855. near Breslau (Wroclaw), Poland. Sektion Geologische Wissenschaften der
E.-M.-Arndt-Universitat Greifswald, German Democratic Republic (GDR), nos. SGWG 83/3 (cf RV: PI.
16. 101. figs. 7, 8). from erratic boulder no. Bey. A20, Zarrenthin b. Jarmen, GDR. approx, lat. 53°56'N,
long. 13°2E E; SGWG 83/4 ( 9 car. : PI. 16. 103. figs. 1-3), from erratic boulder no. Bey. E50, Gager, Isle of
Ruegen. GDR. approx, lat. 54° 17' N, long. 13° 35' E; SGW'G 83/5 (9 LV: PI. 16, 103, fig. 4), from erratic
boulder no. Bey. A32, Zarrenthin b. Jarmen, GDR: SGWG 83/6 (9 LV: PI. 16, 103, figs. 5-7) from erratic-
boulder no. Bey. E12, Gager. Isle of Ruegen, GDR. All specimens Upper Silurian.
Diagnosis: Species of Macrypsilon in which the tecnomorphs have narrow, distinct sulci.
Stereo-Atlas of Ostracod Shells 16, 105 Macrypsilon salterianum (6 of 6)
Remarks: M. salterianum differs from M. parvisulcatum (Sarv 1968) in its more distinctly developed sulci.
The wide stratigraphic range of M. salterianum may be due to confusion between closely related
species (Martinsson 1977, Siveter 1989). M. salterianum shows wide variation between populations
in surface ornament, cruminal shape, the development of the supersulcal tubercle-like feature, the
extent of the depression in the posterodorsal part of the syllobium and the development of the
postcruminal wing-like part of the velum. It is not obvious that the variations are restricted to a
stratigraphic level or geographic province. The occurrence of M. salterianum in Scania (L.
Jeppsson & S. Laufeld 1987, Sver. Geol. Unders., ser. Ca, no. 58, fig. 3) is probable because the
species occurs in the coeval “Red Beyrichienkalk boulders” (Hansch 1985).
Distribution: Upper Ludlow Pfidoli series, Silurian, Canada: Stonehouse Formation, Nova Scotia (Copeland
1964, Copeland & Berdan 1977). Pembroke Formation, Maine, USA? (Siveter 1980).
Great Britain: Kirkby Moor Flags and Scout Hill Flags, Lake District; Upper Whitcliffe and
Downton Castle Sandstone formations, Long Mountain region (Shaw 1969, 1971; Siveter 1980).
Peribaltic area of Poland: Chlapowo borehole; post-Ludlow (Witwicka 1967). Leba 1
borehole, Beyrichienkalk pebbles in the Zechsteinkonglomerat (Martinsson 1964). Leba 2 & 8,
Debki 2 and Piasnica 2 boreholes; post-Ludlow, Neobeyrichia incerta to Nodibeyrichia tuberculata
zones (Zbikowska 1973). Miloszewo, Wejherowo, Karwia, Opalino, Salino, Biatogard and
Leba-IG 1 boreholes; post-Ludlow, Frostiella pliculata to Nodibeyrichia gedanensis zones
(Tomczykowa & Witwicka 1974). Chojnice borehole; post-Ludlow (Zbikowska 1974).
East Baltic area, USSR: Ohesaare 1 & 2 boreholes, Isle of Saaremaa, Estonia; Kaugatuma and
Ohesaare formations (Sarv 1971). Piltene 1, 31 & 32, Stoniskjaj, Kolka 4 & 54 and Pavilosta 51
boreholes, Latvia; Minija and Jura formations (Gailite 1967, 1978). Taurage and Kunkojaj
boreholes, Minija Formation (Pranskevicius 1972); boreholes 87, 94, 96, 98, 108, 110, 112 (Minija
Formation), boreholes 89, 94, 96, 98, 108, 110, 112 (Jura Formation), Arjogalskij profile,
Lithuania (Sidaraviciene 1986). Gusev 5 borehole, Minija Formation (Pranskevicius 1972) and
Dubovskoje borehole, Kaugatuma Formation, Kaliningrad district (Kaljo & Sarv 1976).
Sweden: Klinta Formation and Oved Sandstone sensu Jeppsson & Laufeld (1987), Scania; see
also Gronwall (1897), Moberg & Gronwall (1909), and Martinsson (1967, 375).
Erratic boulders: Beyrichienkalk sensu Martinsson (1963, 1967, 1977); Beyrichienkalk type B,
C, D and “Red Beyrichienkalk" sensu Hansch (1985).
Stereo- Atlas of Ostracod Shells 16 (23) 106-111 (1989) Berolinella steusloffi ( 1 of 6)
595.336.11 (113.333) (430.2 : 161.013.52 + 013.54) : 551.351 + 552.54
ON BEROLINELLA STEUSLOFFI (KRAUSE)
by Wolfgang Hansch & David J. Siveter
(University of Greifswald, German Democratic Republic &
University of Leicester, England)
Genus BEROLINELLA Martinsson, 1962
Type-species (by original designation): Beyrichia steusloffi Krause. 1891
Diagnosis: Amphitoxotidinae having a basal crest along a wide tubulous velum in both sexes. The tubulous.
uninterrupted velar edge and the torus, which forms a sharp basal ridge, both continue across the
crumina (slightly modified after Martinsson 1962, op. cit.).
Remarks: The beyrichiacean Amphitoxotidinae Berolinella, Dibolbina Ulrich & Bassler, 1923, Huntonella
Lundin, 1968, and Tropidotoxotis Siveter. 1980 all have a reasonably complete velar edge cross the
crumina, and are distinguished largely on details of subcruminal, velar and basal crest morphology
(see Siveter, Palaeontogr. Soc. ( Monogr .), 133 (556), 69, 1980). Besides B. steusloffi the genus
currently includes only B. praevia Sarv, 1968, and an undescribed species from Baltic erratic
boulders (Martinsson, 1962).
Explanation of Plate 16, 107
Figs. 1, 2, 9 LV (SGWG 83/1, approx. 1040 /xm long): fig. 1, ext. lat . ; fig. 2, ext. vent. Figs. 3, 4, cf RV (SGWG 83/2, approx.
1200/xm long): fig. 3, ext. vent.; fig. 4, ext. lat.
Scale A (200 /xm; x55), figs. 1, 2; scale B (200 /xm; X65), fig. 3; scale C (200 /xm; x45). fig. 4.
Stereo-Atlas of Ostracod Shells 16, 108 Berolinella steusloffi (3 of 6)
Berolinella steusloffi (Krause, 1891)
1891
71894
? 1895
? 1897
? 1908
non 1909
71916
non 1919
7 1934
7 1954
1956
1957
1962
1963
1964
1966
1967
Beyrichia steusloffi sp. nov. A. Krause, Z. Dt. Geol. Ges., 43 (2), 505, pi. 32, figs. 6, 8, 9; ?7a. b.
B. steusloffi Kr.; A. Steusloff, Ibid. , 46 (4), 786.
Beyrichia steusloffi Krause; J. Ch. Moberg, Sver. Geol. Unders., ser. C, no. 156, 7, 14.
Beyrichia steusloffi Krause; K. A. Gronwall, Ibid. , no. 170, 204, 218, 224, 227, 238.
Beyrichia steusloffi Krause; E. O. Ulrich & R. S. Bassler, Proc. U. S. Nat. Mus., 35 (1646), 286.
Beyrichia steusloffi A. Krause; J. Ch. Moberg & K. A. Gronwall, Lunds Univ. Arsskr. , N.F. 5 (1), 7, 9, 12, 25, 63, 81, 86,
pi. 4, figs. 14, 15.
B. steusloffi Krause; J. Botke, Verb. Geol.-Mijnbouwk. Genootschap v. Nederland en Kolonien , 3, 26.
Beyrichia steusloffi Krause; J. Hede, Geol. For. Stockh. Forh., 41, 135, pi. 5.
Beyrichia steusloffi Krause; R. S. Bassler & B. Kellett, Geol. Soc. America, Spec. Pap. 1, 206.
Beyrichia steusloffi ; G. Henningsmoen, Norsk. Geol. Tidsskr., 34, 29.
Dibolbina steusloffi (Krause); R. V. Kesling, Contr. Mus. Paleont. Univ. Michigan, 13 (2), 56, pi. 4, figs. 1-10, pi. 5, figs.
1-6 (incorrect reconstruction).
Dibolbina steusloffi (Krause); R. V. Kesling & K. J. Rogers, J. Paleont., 31 (4), 1000, pi. 127, figs. 15-21.
Berolinella steusloffi (Krause); A. Martinsson, Bull. Geol. Inst. Univ. Uppsala, 41, 107, 253, figs. 39:9, 129A.
Berolinella steusloffi ; A. Martinsson, Geol. For. Stockh. Forh., 85 (3), 293, 295.
Berolinella steusloffi (Krause); A. Martinsson, Ibid., 86 (2), 128. 135, 156, 159, fig. 15 (log).
Berolinella steusloffi (Krause); D. Kaljo & L. Sarv, Izv. Akad. Nauk Est. SSR, ser. Fisiko-Matem. i techn. nauk, 2,
279, tab. 1.
Berolinella steusloffi; A. Martinsson, Geol. For. Stockh Forh., 89 (4), 377.
Explanation of Plate 16, 109
Fig. 1, 9 RV, ext. lat. (I 6007a, 1210 /am long); fig. 2, 9 LV, ext. ant. (SGWG 83/1). Figs. 3, 4, tecnomorphic RV (I 6007b, 1010 /xm
long): fig. 3, ext. lat.; fig. 4, ext. vent. Figs. 5, 6, cf RV (SGWG 83/2): fig. 5, detail of post, margin; fig. 6, detail of velum.
Scale A (200 /xm ; x 45), fig. 1 ; scale B (200 /xm ; x 60), fig. 2; scale C (200 /xm ; x 55), figs. 3,4; scale D (50 /xm; x 190), fig. 5 ; scale E
(50 /xm; X280), fig. 6.
Stereo- Atlas of Ostracod Shells 16, 107
Berolinella steusloffi (2 of 6)
Berolinella steusloffi (4 of 6)
Stereo-Atlas of Ostracod Shells 16, 109
Stereo-Atlas of Ostracod Shells 16.110 Berolinella steusloffi (5 of 6)
1968 Berolinella steusloffi (Krause): L. Sarv, Ostrakody Craspedobolbinidae, Beyrichiidae i Primitiopsidae Silur Estonii, 27, 95,
tabs. 2. 3, pi. 7, figs. 9, 10, Tallinn (Valgus).
1970 Berolinella steusloffi (Krause); L. Sarv, in: D. Kaljo (ed.), Silur Estonii, 158, 169, 299, Tallinn (Valgus).
1971 Berolinella steusloffi ; L. Sarv, Izv. Akad. Nauk Est. SSR, ser. Chimija-Geol., 20 (4), 353, tab. 3.
1973 Berolinella steusloffi (Krause); B. Zbikowska, Acta Geol. Pol., 23 (4), 613, 625, pi. 4, fig. 8, tab. 2.
71975 Berolinella steusloffi ; A. Pranskevicius, Geol. For. Stockh. Fork.. 97 (1), 53.
1977 Berolinella steusloffi ; L. Sarv, in: D. Kaljo (ed.), Fazii i fauna Silura Pribaltiki, 165, 173, tab. 3, Tallinn (Valgus).
1977 Berolinella steusloffi ; A. Martinsson, The Silurian- Devonian Boundary, IUGS ser. A, no. 5, 48.
1978 Berolinella steusloffi (Krause); D. J. Siveter, in: R. H. Bate & E. Robinson (eds.), A Stratigraphical Index of British
Ostracoda, Geol. J. Spec. Issue, 8, 69.
1985 Berolinella steusloffi (Krause); W. Hansch, Lethaia, 18 (4), 274, tab. 1.
1989 Berolinella steusloffi (Krause); D. J. Siveter. in: C. H. Holland & M. G. Bassett (eds.), A global standard for the Silurian,
263, fig. 168 D, Nat. Mus. Wales Geol. ser. no. 9, Cardiff.
Lectotype:
Type locality:
Figured specimens:
Diagnosis:
Museum fur Naturkunde Berlin, German Democratic Republic (GDR), no. MBO 117; 9 RV.
Krause, 1891, pi. 32. fig. 9. Designated by Sarv, 1968.
[Paratype: MBO 118. tecnomorphic RV.]
Erratic boulder, Beyrichienkalk no. 470 of Krause, 1891. From Muggelheim, Berlin, GDR;
approx, lat. 52°32'N, long. 13°25'E. Pfidoli Series, Silurian.
Sektion Geologische Wissenschaften der E.-M.-Arndt-Universitat Greifswald, GDR, nos.
SGWG83/1 (9 LV: PI. 16, 107, figs. 1,2; PI. 16, 109, fig. 2) ; SGWG 83/2 (cf RV: PI. 16, 107, figs.
3, 4; PI. 16, 109, figs. 5, 6). Both from erratic boulder no. Bey. E7, Gager, Isle of Ruegen, GDR,
approx, lat. 54°17'N, long. 13°45'E. British Museum (Nat. Hist.), nos. I 6007a (9 RV : PI. 16,
109, fig. 1), I 6007b (tecnomorphic RV: PI. 16, 109, figs. 3, 4). Both from erratic boulder no. 600
of Krause, Muggelheim, Berlin, GDR. All specimens are of Pfidoli Series age, Silurian.
Species of Berolinella with similar, well developed prenodal and adductorial sulci, in tecnomorphs
extending from the dorsal margin to the narrow depression above the basal crest of the wide
tubulous velum. Torus and velar edge cross the crumina but not parallel to each other. Velum also
occurs in front of the crumina and is restricted posteroventrally in both dimorphs.
Stereo-Atlas of Ostracod Shells 16. Ill
Berolinella steusloffi (6 of 6)
Remarks: Kesling's (1956) reconstruction of B. steusloffi, showing an entire velum, is incorrect (cf. diagnosis
above). B. praevia differs by having less distinct sulci, parallel torus and velar edge across the
crumina and a more acuminate crumina. The type material of Krause, 1891, figs. 6, 7 is probably
lost; according to his figures, it differs somewhat from the typical B. steusloffi. Berolinella sp. n. of
Martinsson (1962, 253, fig. 129B) may be conspecific with B. praevia. The UB. steusloffi"
specimens of Moberg & Gronwall (1909) do not belong to B. steusloffi-, possibly they represent a
new species, but it is impossible to prepare the material in their slabs to confirm this judgement.
Distribution: B. steusloffi is the youngest Berolinella species and is restricted to the Pfidoli Series, Upper
Silurian.
Peribaltic area of Poland: Leba 1 borehole, Beyrichienkalk pebbles in the Zechsteinkong-
lomerat (Martinsson 1964). Debki 3 borehole, post-Ludlow, Nodibeyrichia tuberculata Zone
(Zbikowska 1973).
East Baltic area, USSR: Ohesaare 2 borehole, Ohesaare-Kliff, Isle of Saaremaa; Ohesaare
Formation (Sarv 1968, 1971). Piltene 32 borehole Latvia; Ohesaare Formation (Sarv 1977).
Erratic boulders: Beyrichienkalk sensu stricto (of Martinsson 1963, 1967, 1977 and
Beyrichienkalk type C sensu Hansch (1985).
Stereo-Atlas of Ostracod Shells 16 (24) 112-115 (1989) Aurikirkbya wordensis ( 1 of 4)
595.336.14 (113.6) (775 : 162.102.30) : 551.351 + 552.54
ON AURIKIRKBYA WORDENSIS (HAMILTON)
by Gerhard Becker & Franciszek Adamczak
( University of Frankfurt, Federal Republic of Germany &
University of Stockholm, Sweden)
Genus AURIKIRKBYA Sohn, 1950
Type-species (by original designation): Kirkbya wordensis Hamilton. 1942
Diagnosis: Kirkbyid genus with two distinct lateral lobes joined by a connecting lobe.
Remarks: The adventral structure (‘velum') is a well developed ridge. The outer list of the contact groove is
terminated dorsally by both an anterior and posterior tooth. The right hinge is provided with a list
and the left hinge with a groove. The right valve is the larger valve and overlaps the smaller left
valve along the free margin. The kirkbyan pit is distinct and situated below the connecting lobe.
Distribution: N America; lower Pennsylvanian to Permian. W Europe; late Upper Devonian (upper
Famennian) and Upper Carboniferous (Westphalian).
Aurikirkbya wordensis (Hamilton, 1942)
1942 Kirkbya wordensis sp. nov. I. B. Hamilton, J. Paleont., 16, 713, 714, pi. 110, fig. 13.
1950 Aurikirkbya wordensis (Hamilton); I. G. Sohn, U.S. Geol. Surv. prof. Pap., 221-C, 36, pi. 7, figs. 1-13.
1954 Aurikirkbya wordensis (Hamilton); I. G. Sohn, U.S. Geol. Surv. prof. Pap., 264-A, 9, pi. 4, figs. 9, 21.
1961 Aurikirkbya wordensis (Hamilton); I. G. Sohn, U.S. Geol. Surv. prof. Pap, 330-A, 141.
1961 Aurikirkbya wordensis (Hamilton); I. G. Sohn, in: R. C. Moore (ed.), Treatise Invert. Paleontol. Pt., Q (3), 164,
text-fig. 95/3.
Holotype: University of Wisconsin, Madison, Wisconsin, USA, no. 22373; an adult LV.
Explanation of Plate 16, 113
Figs. 1, 2, adult RV (USNM 110232a, 1430 /um long): fig. 1, ext. lat. ; fig. 2, ext. vent. obi.
Scale (300 x60), figs. 1, 2.
Stereo-Atlas of Ostracod Shells 16, 114 Aurikirkbya wordensis (3 of 4)
Type locality:
Figured specimens:
Diagnosis:
Remarks:
Distribution :
Scarp N of former Old Word Ranch house NE Marathon, Hess Canyon quadrangle. Glass
Mountains, Brewster County, W Texas, USA (US National Museum locality 703c); lat. 30° 16' N,
long. 103° 10' W. Irregular lenses of bituminous limestone in siliceous shales; uppermost Leonard
or lowermost Word Formation, Middle Permian.
United States National Museum (USNM), Washington DC, USA, nos. USNM 110232a (adult RV:
PI. 16, 113, figs. 1, 2; PI. 16, 115, fig. 2), USNM 110232b (adult LV: PI. 16, 115, fig. 3), USNM
110232c (juv. RV: PI. 16, 115, fig. 1).
All of the figured specimens are topotype material.
Aurikirkbya species with a crenulated contact groove, an indistinct connecting lobe and very
indistinct lobal ridges (‘flanges’) both anteriorly and posteriorly. Posterior (postsulcal) lobe
distinct and subtriangular in dorsal view.
Sohn ( 1954, 9; 1961, 140) described in some N American Aurikirkbya species “the shell wall of the
venter [to be] very thick" and considered this feature to be characteristic for the genus. In A.
wordensis the shell of the connecting lobe also seems to be very thick; however, it is (like the shell
of the venter) not solid but is porous ( = economic construction).
The contact groove of the larger, right valve is crenulated. The crenulation is also visible in the
figure as illustrated by Sohn (1959, pi. 7, fig. 12b). A crenulated contact groove is also known in
Ogmoconchella Gruendel, 1964 (Metacopina). In A. wordensis the corresponding contact list is
most probably smooth. These contact features are interpreted here as associated with water
circulation and filter-feeding (“weir-basket”).
In some specimens (see Plate 16, 115, fig. 3) the internal valve margin seems to be thickened.
This does not demonstrate a calcified inner lamella; most probably it is an artifact product of
silicification.
Texas, USA; uppermost Leonard or lowermost Word Formation, Middle Permian.
Explanation of Plate 16, 115
Fig. 1, juv. RV, int. lat. (USNM 110232c, 890 /urn long); fig. 2, adult RV, int. lat., detail ant. vent. (USNM 110232a, 1430 pun long); fig.
3, adult LV, int. lat. (USNM 110232b, 1480/u.m long).
Scale A (300 /urn; x98), fig.l; scale B (30/u.m; X215), fig. 2; scale C (300/u.m; x53), fig. 3.
Stereo- Atlas of Ostracod Shells 16, 115 Aurikirkbya wordensis (4 of 4)
Stereo-Atlas of Ostracod Shells 16(25) 116-119(1989)
595.336 (113.45) (430.1 : 161 .007.50) : 551 .351 + 552.54
ON NODELLA HAMATA BECKER
by Gerhard Becker
(University of Frankfurt, Federal Republic of Germany
Nodella hamata Becker, 1968
Nodella hamata (1 of 4)
1954 Drepanellina? sp. A. K. Krommelbein, Senckenberg. leth., 34, 256, pi. 1, fig. 6.
1968a Nodella hamata sp. nov. G. Becker, Natur u. Museum, 98. 129, 130, text-figs. 16, 17.
1968b Nodella hamata Becker; G. Becker, Senckenberg. leth., 49, 555-557, text-figs. 1, 2, pi. 1, figs. 1, 2, 6-8.
1985 Nodella hamata Becker; M. Coen, Mem. Inst. Geol. Univ. Louvain, 32, 12, tab. 2, pi. 3, fig. 3.
Holotype:
Type locality:
Figured specimens:
Diagnosis:
Forschungs-Institut Senckenberg, Frankfurt am Main (SMF), Federal Republic of Germany, no.
SMF Xe 5676; an adult heteromorph LV.
Quarry “Steinbreche”, about 1km SW of Refrath village, SW Bergisch-Gladbach. Bergisches
Land, Rheinisches Schiefergebirge, Federal Republic of Germany; lat. 50° 59' N. long. 07° 09' E.
Coral limestones with yellowish marls, Refrath Formation, Frasnian, Upper Devonian.
Forschungs-Institut Senckenberg (SMF), Frankfurt am Main, Federal Republic of Germany, nos. SMF Xe
5676 (adult heteromorphic LV, holotype: PI. 16, 117, fig. 2, PI. 16. 119, fig. 2). SMF Xe 5677 (adult
tecnomorphic car., paratype: PI. 16, 117, fig. 1; PI. 16. 1 19, figs. 1, 3, 5), SMF Xe 5678 (adult heteromorphic
LV, paratype : PI. 16, 117, fig. 3), SMF Xe 5679 (adult heteromorphic RV, paratype: PI. 16, 119, fig. 4). All
topotype material.
Nodella species with a distinct, somewhat elongate presulcal lobe below the dorsal margin and a
high, pointed postsulcal lobe. Ventral-anteroventral bend (“carina") and short posteroventral
spine. Extradomiciliar dimorphism showing a comparatively strong and long marginal hamus in
Explanation of Plate 16, 117
Fig. 1, adult tecnomorphic car., rt. lat. (paratype, SMF Xe 5677, 600 /xm long); fig. 2, adult heteromophic LV, ext. lat. (holotype, SMF
Xe 5676. 550 /xm long); fig. 3, adult heteromorphic LV, ext. lat. (paratype, SMF Xe 5678. 600 /xm long).
Scale (100/xm; x 110), figs. 1-3.
Stereo-Atlas of Ostracod Shells 16, 118 Nodella hamata (3 of 4)
tecnomorphs and a long, sickle shaped anterior flange in heteromorphs. Also proportional
dimorphism, in which the tecnomorphs are slimmer. Lateral surface of the valves finely reticulate.
Remarks: The extradomiciliar dimorphic structures in Nodella hamata are clearly marginal in origin. Both
the dimorphic structures, the hamus in tecnomorphs and the flange in heteromorphs, originate
anteriorly from the otherwise free marginal positioned marginal ridge. The additional, domiciliar
dimorphism is proportional. This combined type of sexual dimorphism was termed “hamal
dimorphism” by Becker (1968a, 129).
What we now term hamal structures were first believed to characterise different subspecies
(Zaspelova, Trudy VNIGRI, 60. 173, 174, 188, 189, pi. 3, figs. 1-5, pi. 8, figs. 1-3, 1952).
Referring to the “modern” taxonomic concepts of Jaanusson (Bull. Geol. Inst. Univ. Uppsala , 37,
197-226, 1957), Becker (1968a, 131) proposed the Suborder Nodellocopina and the Superfamily
Nodellacea for palaeocopids with hamal dimorphism. However, Schallreuter (in: T. Hanai et al.
(eds.). Evolutionary Biology of Ostracoda, Develop. Palaeont. Stratigr., Amsterdam, 11, 1047,
1988) considered that hamal dimorphism was a "special modification” of the antral dimorphism
which characterises hollinids. If this were the case, however, then nodellid (marginal) structures
would be considered homologous with features which are velar and not marginal - which they
clearly, in my opinion, are not. Schallreuter (1988, op. cit.) also compared the nodellid marginal
structures with the perimarginal structures of primitiopsids; however, the latter are typically
primitiopsid features and thus are not homologous or even analogous features. Moreover, (the
nodellid) proportional dimorphism is not at all known in hollinids, and the overall morphology of
nodellids is rather drepanellid-like. Thus, the Superfamily Nodellacea Becker, 1968 is considered
to be a valid taxon, closely related to the Drepanellacea Ulrich & Bassler, 1923.
Distribution: Bergisches Land, Rheinisches Schiefergebirge, Germany; Refrath Formation, Frasnian, Upper
Devonian. Dinant Syncline, Ardennes, Belgium; Fromelennes superieur, supposed late Middle
Devonian.
Explanation of Plate 16, 1 19
Figs. 1, 3, 5, adult tecnomorphic car. (paratype, SMF Xe 5677, 600 pm long): fig. 1, vent.; fig. 3, vent, view of atn. end; fig. 5,
anterovent. obi. Fig. 2, heteromorphic LV, ext. atn. (holotype, SMF Xe 5676, 550 pm long). Fig. 4, heteromorphic RV, ext. vent,
(paratype, SMF Xe 5679, 560 /xm long).
Scale A (100 /xm; x 110), figs. 1, 4; scale B (100 /xm; x205), fig. 2; scale C (30 /xm; x 300), figs. 3, 5.
Nodella hamata (4 of 4)
Stereo-Atlas of Ostracod Shells 16, 119
Stereo-Atlas of Ostracod Shells 16, 117
Nodella hamata (2 of 4)
Stereo-Atlas of Ostracod Shells 16(26) 120-127 (1989)
595.337.14 (118.15) (430.1 : 161.008.49) : 551.35
Cytheridea sandbergeri ( 1 of 8)
ON CYTHERIDEA SANDBERGERI KAMMERER sp. nov.
by Thomas Kammerer
(Geologisches Landesamt Rheinland- Pfalz, Mainz, German Federal Republic &
University College of Wales , Aberystwyth, UK)
Cytheridea sandbergeri sp. nov.
1905 Cytheridea muelleri (v. Munster); E. Lienenklaus, Ber. senckenb. naturf. Ges., 1905, 38 (pars).
1955 Cytheridea pernota sp. nov. H. Oertli & A. J. Key (= Keij), Bull. Verein. schweiz. Petrol. Geol. Ing.. 22 (62), 19 (pars), pi. 1.
figs. 8-13 only ( non pi. 1. figs. 1-7, text-fig. 2).
1956 Cytheridea pernota Oertli & Key; H. J. Oertli, Schweiz, palaeont. Abh., 74. 36, pi. 2, figs. 33-38.
1960 Cytheridea pernota Oertli & Keij; F. Gramann, Marb. Sitzungsber. , 82, 59-88 (passim ), pi. 1, fig. 4.
1962 Cytheridea sp. C 66 [aff. mullerii (Munster 1830)] (sic); H. Malz, Ostracoda, in: F. Doebl & H. Malz, Tertidr des
Rheintal-Grabens. Leitfossilien der Mikropaldontologie , Gebruder Borntraeger. Berlin. 394, pi. 58, figs. 1-2.
Holotype: Forschungsinstitut Senckenberg, Frankfurt, no. SMF Xe 14751; 9 left valve.
[Paratypes: nos. SMF Xe 14752-14764]
Type locality : Borehole no. 27 (KB 2), sample 6015/5922, depth 83.50-83.75 m, Bodenheim. near Mainz,
German Federal Republic (grid ref. R 49 140, H 32 385 - map no. 6015; long. 8° 17' 31" E, lat.
49°55'36"N); restricted marine marl, Schleichsand Formation of Mainz Basin; Rupelian, M.
Oligocene.
Explanation of Plate 16, 121
Fig. 1, $ FV, ext. lat. (holotype, Xe 14751, 865 pm long); fig. 2, cf FV, ext. lat. (paratype, Xe 14752, 866 long); fig. 3, juv. -1 FV.
ext. lat. (paratype, Xe 14753, 658 yum long).
Scale A (100/xm; x75), figs. 1-3.
Stereo-Atlas of Ostracod Shells 16, 122
Derivation of name:
Figured specimens:
Cytheridea sandbergeri (3 of 8)
In honour of C.L.F. Sandberger (1826-1898), in appreciation of his work on the geology of the
Mainz Basin.
Forschungsinstitut Senckenberg nos. Xe 14751 (holotype, 9 LV: PI. 16, 121, fig. 1); Xe 14752
(Cf LV: PI. 16, 121, fig. 2); Xe 14753 (juv. LV: PI. 16,' 121, fig. 3); Xe 14754 (9 RV; PI. 16, 123,
fig. 1); Xe 14755 (cf RV: PI. 16, 123, fig. 2); Xe 14756 (9 car.: PI. 16, 123, fig. 3); Xe 14757 (9
LV: PI. 16, 125, fig. 1); Xe 14758 (cf car.: PI. 16, 125, fig. 2); Xe 14759 (cf LV: PI. 16, 125, fig. 3);
Xe 14760 (9 RV: PI. 16, 127, fig. 1); Xe 14761 (cf RV: PI. 16, 127, fig. 2); Xe 14762 (cf RV: PI.
16, 127, fig. 3); Xe 14763 (9 RV: Text-fig. 1).
All specimens are from the type locality and horizon.
Size:
Table 1. Measurements on 120 valves (holotype and 119 paratypes); N = no of specimens, x =
mean, L= length not including marginal denticles, H = height.
Diagnosis: Right valve in lateral view pear-shaped in females, elongate subtrapezoidal in males, ventral
margin slightly concave in posterior third; left valve subovate with straight or very slightly concave
ventral margin. Anterior margin of both valves high and broadly rounded, forming a nearly
symmetrical semicircle; antero-cardinal angle indistinct. Left valve with 5-7 anterior marginal
Explanation of Plate 16, 123
Fig. 1, 9 RV, ext. lat. (paratype, Xe 14754, 796 yum long); fig. 2, cf RV, ext. lat. (paratype, Xe 14755, 796 gm long); fig. 3, 9 car., ext.
dors, (paratype, Xe 14756, 815 yu.m long).
Scale A (100 yum; x75), figs. 1-3.
Stereo-Atlas of Ostracod Shells 16, 121
Cytheridea sandbergeri (2 of 8)
Cytheridea sandbergeri (4 of 8)
Stereo-Atlas of Ostracod Shells 16, 123
Stereo-Atlas of Ostracod Shells 16, 124 Cytheridea sandbergeri (5 of 8)
denticles, right valve with 7-9 anterior and 4 postero-ventral marginal denticles. Surface
punctate, but smooth along the dorsal margin, especially around the antero-cardinal angle.
Remarks: Sexual dimorphism is pronounced, the males being more elongate in lateral view7 and narrower in
dorsal view. The puncta are coarsest centrally, decreasing in diameter towards the periphery; near
the free margin they are aligned in several parallel rows. Along the anterior margin these rows
form 3 or 4 concentric furrows which in the left valve develop into a mesh-like pattern.
The hinge and internal features are very similar to those of genotype. In C. muelleri the
adductor and mandibular muscle scars are larger than in C. sandbergeri and therefore seem to be
positioned closer to each other.
C. sandbergeri was formerly confused with C. muelleri or C. pernota. The former, from the
Chattian of NW Germany, differs in outline and in the number of anterior marginal denticles of its
right valve. The latter, from the Oligocene of Belgium, the Isle of Wight and NW Germany, has a
distinct antero-cardinal angle and coarser puncta. For a review' and re-illustration of C. muelleri
muelleri (v. Munster), C. m. toenisbergensis Weiss and C. pernota Oertli & Key, see Weiss
( Stereo-Atlas Ostracod Shells , 11 (parts 8-10), 1984). C. sandbergeri represents a major part of the
ostracod fauna within brackish and restricted marine sections of the Schleichsand and
Cyrenenmergel formations of the Mainz Basin (Kammerer, in prep.). During the Rupelian and
Early Chattian it was widespread in the Upper Rhine Graben and neighbouring areas with several
allochronous and allopatric, or parapatric subspecies, or ecotypes, occurring. Similar occurrences
are found in the Swiss Molasse (Oertli & Key, and Oertli. opera cit.) and in the Hessian
Depression (Gramann, op. cit.), at the time attributed to C. pernota.
Explanation of Plate 16, 125
Fig. 1, 9 TV, int. lat. (paratype, Xe 14757, 830 /am long); fig. 2, cf car., ext. dors, (paratype, Xe 14758. 882 /am long); fig. 3, cf LV.
int. lat. (paratype, Xe 14759, 818 /urn long).
Scale A (100 /am; x75), figs. 1-3.
Stereo-Atlas of Ostracod Shells 16, 126 Cytheridea sandbergeri (7 of 8)
Distribution: Oligocene, Rupelian and Early Chattian. Mainz Basin: Schleichsand and Cyrenenmergel,
numerous localities; Rhine Graben: Meletta-Schichten and Cyrenenmergel (Malz, op. cit.);
Hessian Depression: Schleichsand, various localities (Gramann, op. cit.); Swiss Molasse:
Meeressand, Blaue Tone and Cyrenensand, various localities (Oertli, op. cit.).
Text-fig. 1. Internal view of C. sandbergeri from
camera lucida drawing and SEM-micrograph of 9
RV (paratype, Xe 14763, 820 /xm long).
No. of
specimens
• 1
• 2
• 3
• 4
700 750 800 850 900 950 1000
Length (in ^jm)
Text-fig. 2. Length/height plot of 60 left valves of
C. sandbergeri (holotype and paratypes).
Explanation of Plate 16, 127
Fig. 1, 9 RV, int. lat. (paratype, Xe 14760, 818/xm long); fig. 2, cf RV (paratype, Xe 14761), int. muse, sc.; fig. 3, cf RV, int. lat.
(paratype, Xe 14762, 800 /im long).
Scale A (100 /um; x75), figs. 1, 3; scale B (100 /xm; xllO), fig. 2.
Cytheridea sandbergeri (6 of 8)
Stereo-Atlas of Ostracod Shells 16, 125
Stereo-Atlas of Ostracod Shells 16, 127
Cytheridea sandbergeri (8 of 8)
Stereo-Atlas of Ostracod Shells 16 (27) 128-135 (1989) Strandesia weberi (1 of 8)
595.337.12 (119.4+ 119.9) (596 : 161.104.12 + 540 : 161.081.25) : 551.312
ON STRANDESIA WEBERI (MONIEZ)
by Dietmar Keyser & S. B. Bhatia
(University of Hamburg, German Federal Republic & Panjab University, Chandigarh, India)
Strandesia weberi (Moniez, 1892)
1892 Cypris weberi sp. nov. R. Moniez, in: M. Weber, Zoologische Ergebnisse einer Reise in Niederlcindisch Ost-Indien , E. J. Brill,
Leiden. 2, 129-135, pi. 10, figs. 6-11.
1912 Cypris weberi Moniez; G. W. Muller, Tierreich, 31, 233.
1923 Cypris magnifica sp. nov. V. Brehm, Treubia , 3, 222, figs. 1-3.
1932 Eucypris weberi (Moniez); W. Klie, Arch. Hydrobiol. , suppl. 11, 459.
1964 Strandesia spinifera sp. nov. G. Hartmann, bit. Revue ges. Hydrobiol. , Syst. Beih., 3, 141-144, figs. 63a-c, 64a-c.
1979 Strandesia weberi (Moniez); R. Victor & C. H. Fernando, Can. J. Zool, 57, 7, fig. 4.
1980 Strandesia weberi (Moniez); R. Victor et al.. Can J. Zool.. 58, 730.
1983 Strandesia spinifera Hartmann; S. B. Bhatia, in: R. Maddocks, Applications of Ostracoda, University of Houston Geoscience,
442-458, pi. 1, figs. 1-6.
1983 Strandesia weberi (Moniez); N. W. Broodbakker. Bijdr. Dierk... 53. 347, fig. 9H.
Explanation of Plate 16, 129
Fig. 1, LV ext. lat. (ZIM K-34 332, 1095 p,m long, 1278 gm long with spines); fig. 2, RV ext. lat. (ZIM K-34 332, 1145p.m long,
1541 pm long with spines).
Scale A (300 gm; x 73), figs. 1, 2.
Stereo-Atlas of Ostracod Shells 16, 130
Lee to type
Strandesia weberi (3 of 8)
Zoologisch Museum, Amsterdam, no. ZMA, Ostr. 150.710A; designated bv Victor & Fernando
(1979).
Type locality:
Figured specimens:
Diagnosis:
Remarks:
Distribution :
[Paralectotypes: ZMA, Ostr. 150.7 10B]
Celebes, Lumu (approx, lat. 2°30'S, long. 119°00'E). Recent, freshwater.
Zoologisches Museum, Hamburg (ZIM) no. K-27 470 (appendages; Text-figs. 1, 2). From a small
lake near Krakor, Pursat Province, Cambodia (Kampuchea) (approx, lat. 12°30'N, long.
104°00'E); coll. Lindberg.
ZIM no. K-34 332 ($ RV and LV: PI. 16, 129, figs. 1,2; PI. 16. 131, figs. 1,2; PI. 16, 133. figs.
1,2; PI. 16. 135, figs. 1,2). From Holocene marls, Indo-Ganges Plain, at Misa Tal, near Lucknow,
India (approx, lat. 25°N, long. 81°E); coll. Bhatia.
A distinctive Strandesia with a long, hollow posterior spine, about half the length of the shell, in
the RV and with two short, curved anterior spines, one-sixth to one-seventh the length, in the LV.
Surface of valves finely pitted with minute granules on intervening ridges. Shell, in life, is brownish
with blue spots.
For extensive discussion of this and related species, see Victor & Fernando (1979, op. cit.). It
could be confused with S. trispinosa Pinto & Purpur (Publcoes Esp. Esc. Geol. Porto Alegre , 7,
1-53, 1965) from South America and the Caribbean, as noted by Broodbakker ( 1983, op. cit. ), but
there is some difference in dorsal gibbosity between the two, as well as slight differences in the
spines on some of the legs.
So far, only females have been found.
Recent, freshwater: Celebes (Moniez, 1892); Java, Philippines, Malaya, India (Victor &
Fernando, 1979); Cambodia (Hartmann, 1964 and herein). Fossil; Holocene marls from the
Indo-Ganges Plain, near Lucknow (Bhatia, 1983 and herein).
Explanation of Plate 16, 131
Fig. 1, RV int. lat. (ZIM K-34 332); fig. 2, LV int. lat. (ZIM K-34 332).
Scale A (300 gm; X73), figs. 1, 2.
Stereo-Atlas of Ostracod Shells 16, 129
Strandesia weberi (2 of 8)
Stereo- Atlas of Ostracod Shells 16, 131
Strandesia weberi (4 of 8)
Explanation of Plate 16, 133
Fig. 1, LV ext. dors. (ZIM K-34 332); tig. 2, RV ext. vent. (ZIM K-34 332).
Scale A (300 yum; X73), figs. 1, 2.
Stereo-Atlas of Ostracod Shells 16, 134 Strandesia weberi (7 of 8)
Text-fig. 2. Soft-parts of Strandesia weberi (K-27 470). a. Maxilla (P I); b, Thoracopod I (P II); c, Thoracopod II (P III); d, Furca.
Scales: 50 /urn.
Explanation of Plate 16, 135
Fig. 1, LV (ZIM K-34 332) int. muse, sc.; fig. 2, RV (ZIM K-34 332), pore cone with broken bristle.
Scale A (50/rm; x270), fig. 1; scale B (5gm; x 1,850), fig. 2.
Stereo-Atlas of Ostracod Shells 16, 133
Strandesia weberi (6 of 8)
Stereo-Atlas of Ostracod Shells 16, 135
Strandesia weberi (8 of 8)
Stereo-Atlas of Ostracod Shells 16 (28) 136-139 ( 1989)
595.337.14 (119.1) (923 : 163.123.11) : 551.353
Abyssobythere guttata ( 1 of 4)
Derivation of name:
Diagnosis:
Remarks:
ON ABYSSOBYTHERE GUTTATA AYRESS & WHATLEY gen. et sp. nov.
by Michael A. Ayress & Robin C. Whatley
(Geochem Laboratories Ltd., Chester & University College of Wales, Aberystwyth)
Genus ABYSSOBYTHERE gen. nov.
Type-species (here designated): Abyssobythere guttata sp. nov.
Alluding to the occurrence of this bythocytherid genus in the abyss.
Carapace large; subovate to subrhomboidal. Anterior margin broadly rounded, posterior margin
with well developed caudal process at mid-height. Dorsal margin of left valve straight or convex.
Moderately thick-shelled. Surface smooth. Inner lamella broad, vestibulate. Radial pore canals
numerous, narrow and straight.
Abyssobythere is assigned to the Bythocytheridae because of its five adductor muscle scars and
lophodont hinge. It differs from Pseudocythere Sars, 1866 in its thicker shell, more ventral caudal
process and its numerous radial pore canals. Also, in Pseudocythere the right and left valve
outlines are always equal. Abyssobythere differs from V elibythere Schornikov, 1982 in lacking an
alar process; from Rhombobythere Schornikov, 1982 in lacking reticulation or costae; and from
Jonesia Brady, 1866 in its ovate outline and blunt caudal process.
Four other species, as yet undescribed (from the Palaeogene of the SW Pacific. DSDP sites 207
and 209) are assigned to Abyssobythere (see K. Millson, The Palaeobiology of Palaeogene
Ostracoda from Deep Sea Drilling Project Cores in the S W Pacific, unpubl. PhD. thesis, Univ.
Wales, 1, 113-121; 2, pi. 4, figs. 24-29, pi. 5, figs. 1-9). ft. fifth species, as yet undescribed, has
been recovered from the lower Miocene of the Loyaltv Basin, SW Pacific (Harlow pers. comm.
1989).
Explanation of Plate 16, 137
Fig. 1, LV, ext. ant. (OS13389, 920 /xm long); fig. 2, LV, ext. lat . , (holotype, OS13386, 960 /xm long); figs. 3-4, RV (OS13387, 950 /am
long): fig. 3, ext. lat.; fig. 4 ext. ant. vent. obi.
Scale A (200 /xm; x60), fig. 1; scale B (500 /xm; x60); figs. 2-3; scale C (500 /xm; x 60), fig. 4.
Stereo-Atlas of Ostracod Shells 16, 138
Abyssobythere guttata (3 of 4)
Holotype:
Type locality :
Derivation of name:
Figured specimens:
Diagnosis:
Remarks:
Abyssobythere guttata sp. nov.
British Museum (Nat. Hist.) no. OS13386. LV.
[Paratypes British Museum (Nat. Hist.) nos. OS13387-OS13389].
Timor Sea, DSDP Site 262, near axis of Timor Trough, lat. 10° 52. 19' S, long. 123° 50.78' E. Water
depth 2298m. Brown foraminiferal ooze. Zone NN19, Pleistocene.
Latin, alluding to the drop-like outline in lateral view.
British Museum (Nat. Hist.) nos. OS13386 (holotvpe, LV : PI. 16, 137, fig. 2), OS13387 (RV: PI.
16, 137, figs. 3, 4), OS13389 (LV: PI. 16, 137, fig. 1; PI. 16, 139, fig. 2), OS13388 (RV: PI. 16, 139,
figs. 1, 3). All from the type locality and horizon.
Carapace subovate to subrhomboidal in lateral view. Dorsal margin in left valve convex. Right and
left valve outlines virtually equal. Each radial pore canal extends into a box-like chamber distally
and emerges at the base of an external peripheral groove.
A. guttata is most similar to an undescribed species from the lower Miocene of the Loyalty Basin,
SW Pacific, but in that species the dorsal margin of the left valve is straight.
Distribution :
Nannoplankton Zone NN19, Pleistocene of the
Timor Trough (DSDP Site 262, Core 36, Section
6), Zone NN21, Pleistocene of northern flank of
Naturaliste Plateau, eastern Indian Ocean
(DSDP Site 258, Core 1, Section 1), Zone NN19-
NN21, Pleistocene of southeast Wharton Basin,
eastern Indian Ocean (DSDP Site 259, Core 1,
Section 3).
Text-fig. 1. Internal features
observed through transmit-
ted light. LV (OS13389,
920 gm long).
Explanation of Plate 16, 139
Figs. 1, 3, RV (OS13388, 920 /xm long): fig. 1, int. lat.; fig. 3, adductor muscle scar detail. Fig. 2, LV, int. lat. (OS13389, 920 /xm long).
Scale A (500 /xm; x70), figs. 1-2; scale B (100 /xm; x 270), fig. 3.
Abyssobythere guttata (2 of 4)
Stereo-Atlas of Ostracod Shells 16, 137
Stereo-Atlas of Ostracod Shells 16, 139
Abyssobythere guttata (4 of 4)
Stereo-Atlas of Ostracod Shells 16 (29) 140-147 ( 1989) Bryocypris grandipes (1 of 8)
595.337.12 (119.9) (671.1 : 161.011.03) : 551.31/551.312
ON BRYOCYPRIS GRANDIPES R0EN
by Koen Martens
(Koninklijk Belgisch Instituut voor Natuurwetenschappen, Hydrobiologie, Brussels, Belgium)
Genus BRYOCYPRIS R0EN, 1956
Type-species (by original designation): Bryocypris grandipes R0EN, 1956
1956 Bryocypris gen. nov. U. Rpen, Bull. Inst. fr. Afr. noire , 18, ser. A (3), 916.
Diagnosis: Cypridopsine genus with elongated carapace, RV overlapping LV frontally, caudally and ventrally, RV with
well developed frontal and caudal inner lists, LV with caudal, submarginal selvage and weak frontal inner list;
4 large adductor muscle scars present.
Antenna with typical cypridopsine sexual dimorphism in the apical armature: male antenna with claw G3
reduced to a short seta, zx a stout claw and z3 missing. Maxillular palp with distal segment rectangular and
elongated. First thoracopod with penultimate segment divided. Second thoracopod with a pincer, i.e. fourth
segment not individually developed. Hemipenis with inner spermiductus showing the typical cypridopsine
coils in parts c and d. Males, as usually in this group, without a furca (see K. Martens & C. Meisch,
Hydrobiologia, 127, 9-15, 1985); females with a furca of the normal type.
Remarks: Bryocypris appears closely related to both Sarscypridopsis McKenzie, 1977 and Plesiocypridopsis Rome,
1965, yet differs from both genera by a number of morphological peculiarities, the most important ones being
the general outline of the valves, the shape of the furcal ramus in the female and the external anatomy of the
hemipenis.
Explanation of Plate 16. 141
Fig. 1, cf RV, int. lat. (paratype, KM. 512, 534 /um long); fig. 2, cf car. vent, (paratype, OC1477. 534 /am long); fig. 3, cf LV, int. lat.
(paratype, KM. 512, 552 yum long); fig. 4, cf LV, ext. lat. (paratype, OC1476, 552 /urn long).
Scale A (100 /rm; X 1 10)
Stereo- Atlas of Ostracod Shells 16, 142 Bryocypris grandipes (3 of 8)
Bryocypris grandipes Roen, 1956
1956 Bryocypris grandipes sp. nov. U. Roen, Bull. Inst. fr. Afr. noire, 18, ser. A (3), 916-920, figs. 7-19.
Zoologisk Museum (Copenhagen): unnumbered specimens labelled “holotype 2> allotype cf” (not
designated in original publication); the ICZN is unclear with regard to the validity of such types, but since the
designation was suggested by the original author, I propose to accept both holotype and allotype and consider
all other specimens paratypes: cf with soft parts dissected in a sealed slide with glycerine, valves stored dry
(KM. 512); 200 cfcf and 648 22 > the majority in toto in spirit (no number). Also 1 cf and 1 2 dissected and
c. 10 in toto specimens in the KBIN, Brussels (nos. OC1475-1481).
Mosses in caves just above the waterfall of Mpoume. N bank of R. Nyong, Cameroon, Africa (approx, lat.
3° 30' N, long. 11° 05' E). The ‘caves' actually consist of spaces between giant boulders and the thick mats of
mosses lining the roofs and walls of the entrances are never submerged, but kept steadily moist by the fog-like
splash from the fall (J. Birket-Smith, Bull. Inst. fr. Afr. noire, 18, ser. 1(2), 567-582, 1956).
Zoologisk Museum, Copenhagen: KM. 512 (paratype, cf: PI. 16, 141, figs. 1, 3; Text-figs. 1 (B-D, G), 2
(A-D, F), 3 (A-C, E, F) ). KBIN, Brussels, all paratypes: OC1477 (cf ; PI. 16, 141, fig. 2), OC1476 (cf: PL
16, 141, fig. 4; PI. 16, 143, fig. 5; Text-figs. 1(F), 3 (D, G)), OC1475 (2: PL 16, 143, figs. 1, 3); Text-figs.
1 (E), 2(E), 3 (H, I)), 00478(2: PL 16, 143, fig. 2). 00479(2: PL 16. 143, fig. 4), 00480 (2 : Text-fig.
i(A)).
Valves elongated, with posterior margin more widely rounded than anterior one and with numerous marginal
setae. Antennula without Rome-organ. Antenna with natatory setae short, hardly reaching beyond tip of
their segment. Left prehensile palp with terminal segment elongated and distally dilated, proximally
narrower, but not folded as in Plesiocypridopsis', right prehensile palp curved, shorter and narrower. First
thoracopod with penultimate segment with only 1 apical seta and distal segment without lateral seta. Furca in
female with a short, conical ramus, a small lateral seta and an extremely elongated and flagellum-like apical
seta. Genital region in the female with a solid, elongated and curved genital hook. Hemipenis with lateral
shield rounded, bearing a pronounced, subapical thumb-like processus. Inner spermiductus with a
supplementary coil, with the bursa copulatryx large and simple and not surrounded by additional trabeculae.
Explanation of Plate 16, 143
All paratypes. Fig. 1, 2 RV int. lat. (OC1475, 586 /xm long); fig. 2, 2 car. dors. (OC1478, 534 gm long); fig. 3, 2 LV int. lat.
(OC1475, 552 /u.m long); fig. 4, 2 car. dors. (OC1479, 517 gm long); fig. 5, cf RV ext. lat. (OC1476, 552 gm long).
Scale A (100/am; xllO).
Type specimens:
Type locality:
Figured specimens :
Diagnosis:
Stereo-Atlas of Ostracod Shells 16, 141
Bryocypris grandipes (2 of 8)
Stereo-Atlas of Ostracod Shells 16, 143
Bryocypris grandipes (4 of 8)
Stereo-Atlas of Ostracod Shell 16, 144
Bryocypris grandipes (5 of 8)
Text-fig. 1 (see 16, 147 for explanation)
Stereo-Atlas of Ostracod Shell 16. 145
Bryocypris grandipes (6 of 8)
Text-fig. 2 (see 16. 147 for explanation)
Stereo-Atlas of Ostracod Shell 16. 147
Bryocypris grandipes (8 of 8)
Remarks: B. grandipes is a typical cypridopsine species, the most important feature being the presence of a reduced
furca in the female, but also because of the sexual dimorphism in the apical armature of the antenna (i.e. the
reduction of z3) and the structure of the inner spermiductus in the hemipenis. Together with Potamocypris
Brady, 1870, Bryocypris is thus far the only genus where well developed genital hooks are known in the
females. However, this region of the body has not yet been illustrated properly for most species and genera
and the feature may prove to be present in a number of other genera. Bryocypris is only the second
cypridopsine group which has adapted to the terrestrial environment, Callistocypris Schornikov, 1980 ( Zool .
Zh., 59. 1306-1319, 1980) being already described from purely terrestrial environments on the Solomon
Islands. The latter genus (placed in a separate subfamily), however, displays far more morphological
adaptations to such environments. For example, all segments, claws and setae in Callistocypris are short and
stout and even the furca, although clearly cypridopsine, appears more solidly built and has a well developed,
complex furcal attachment. Bryocypris has none of these features; its only adaptations to terrestrial
conditions are the reduction of the natatory setae on the antenna (not unusual in Cypridopsinae), the
disappearance of 2 setae on the first thoracopod and the presence of numerous marginal setae on the valves.
This could indicate that its invasion in such habitats is a fairly recent phenomenon, and that it is unable to live
in truly terrestrial situations (e.g. leaf litter in forests, like for example Callistocypris and Terrestricandona
Danielopol & Betsch, 1980), but is rather restricted to semi-terrestrial environments (mosses in splash zones,
etc.). It is of interest to note that species with a reduced furca can apparantly still (re-)adapt to a crawling
locomotion in difficult circumstances, although in one lineage ( Callistocypris ) this caused a secondary
reinforcement of the furca.
Acknowledgements : Dr T. Wolff (Copenhagen) is acknowledged for his help in providing access to the type material. Mr J. Cillis
and Mrs C. Behen offered technical assistance with the illustrations.
Text-fig. 1. A, 9 paratype (OC1480, 574(um long), B-D, G, cf paratype (KM. 512), E, 9 paratype (OC1475, F, O' paratype (OC1476). A. RV, int.
lat. ; B, hemipenis; C. mandibular palp, showing part of chaetotaxy; D, antenna, detail of natatory setae; E, antennula, chaetotaxy of endopodite not
shown; F, Zenker's organ; G, idem, detail in frontal view. Scale: 156/um for A; 81 /xm for F, G; 33 gm for B-E.
Text-fig. 2. A-D, F, O’ paratype (KM. 512), E, 9 paratype (OC1475). A, rake-like organs; B, mandibular palp (respiratory plate and chaetotaxy of
fourth segment not shown); C, first thoracopod; D, idem, detail; E, furcae and genital region, showing genital hooks; F, mandibular coxa. Scale:
81 pm for C, F; 33 yum for A, B, D, E.
Text-fig. 3. A-C, E, F, O’ paratype (KM. 512), D, G, O paratype (OC1476), H, I, 9 paratype (OC1475). A, maxillula; B, second thoracopod; C,
idem, detail; D, left prehensile palp; E, right prehensile palp; F, left antenna in medial view, detail of apical armature; G, antenna, detail of aesthetasc
Y; H, right antenna in lateral view, detail of apical armature; I, antenna, detail of aesthetasc Y. Scale: 81 (im for B; 33/j.m for A, C-E
Stereo-Atlas of Ostracod Shells 16(30) 148-151 (1989) Limnocythere hibernica (1 of 4)
595.337.14(116.222) (261.2: 162.013.52) : 551.312.4
ON LIMNOCYTHERE HIBERNICA ATHERSUCH sp. nov.
by John Athersuch
(BP Research, Sunbury- on -Thames, England)
Limnocythere hibernica sp. nov.
British Museum (Nat. Hist.) no. OS13432; 9 car.
[Paratypes: British Museum (Nat. Hist.) nos. OS13431, 13433-13437]
The well 26/28-1 in the Porcupine Seabight, offshore SW Ireland (approx, lat. 52°02'N, long.
12°33'W); Middle Jurassic, probably Late Bathonian.
Latin, hibernia = Ireland ; alluding to the location of the type locality in Irish territorial waters.
British Museum (Nat. Hist.) nos. OS13432 (holotype. 9 car. : PI. 16, 149, fig. 2), OS13431 (9 car. :
PI. 16, 149, fig. 1), OS 13433 (cf car.: PI. 16, 149, fig. 3), OS13434 (9 car.: PI. 16, 151, fig. 1),
OS 13435 ( 9 car. : PI. 16, 151, fig- 2), OS13436 (cf car. : PI. 16, 151, fig. 3), OS13437 (LV: Text-fig.
1). All specimens from the type locality; OS13432, 13434 at 2370m (cuttings); OS13436 at 2435 m
(cuttings); OS13431, 13433, 13435, 13437 at 2437.4m (core).
Each adult valve bears a group of variably coalesced protuberances and swollen ridges. Single
subcentral and median dorsal tubercles are separated by two vertical sulci from single anterior and
posterior ridges. The anterior ridge is positioned some way from and subparallel to the anterior
margin; in some specimens it appears to be formed of two coalesced tubercles. The posterior ridge
which runs parallel to the posterior margin has a pronounced C-shape, the dorsal branch being
swollen terminally; the largest of the tubercles lies between the end of the ventral branch and the
subcentral tubercle with which it tends to coalesce in some specimens. Surface ornament of
subrounded reticulation; fossae largest and best developed in posterolateral areas, reducing to
Explanation of Plate 16, 149
Fig. 1, 9 car., ext. h- Ht. (paratype, OS13431, 506 /xm long); fig. 2, 9 car., ext. It. lat. (holotype, OS13432, 552/u.nr long); fig. 3, cf
car., ext. It. lat. (paratype, OS13433, 616/u.m long). Scale A (250/ixm; x 105), figs. 1-3.
Stereo-Atlas of Ostracod Shells 16, 150 Limnocythere hibernica (3 of 4)
foveolae towards margins.
Remarks: There is a considerable amount of variation in the development of the tubercles within a
population of individuals of approximately the same size; this variation is not always the result of
abrasion. In some individuals the anterior and posterior ridges are only weakly developed (PI. 16,
151, fig. 1). A posterodorsal concavity shows that the posterior ridge is, at least in part, hollow.
There is also a small ocular sinus. The adductor muscle scar area is not clearly visible. More
elongate specimens are presumed to be males.
This species is placed in the genus Limnocythere principally because of the remarkable
similarity of its external morphology to many Recent and Neogene species of that genus (e.g. the
living L. porphyretica De Deckker, 1981 ( Zoologica Scr., 10, 41-42, figs. 3, 4).
The valve interior is known only from one damaged specimen (Text-fig. 1); it displays a
lophodont hinge and at least 23 slightly sinuous, unbranched radial pore canals anteriorly. These
are also features consistent with the genus Limnocythere.
Distribution: Known only from wells in the vicinity of the type locality in the Porcupine Seabight. By
comparison with living species, L. hibernica is probably indicative of a non-marine lacustrine
episode. At the type locality this species occurs in an interval with Bisulcocypris spp., Darwinula
sp., conchostracans, gastropods and abundant terrestrial miospores and bisaccate pollen.
Monotypic flood occurrences of this species have been observed in thin beds in core samples.
i
i
i
i
i
Text-fig. 1. Internal view of broken left valve; sex unknown (paratype, OS13437). |_ _2oo^m__|
Explanation of Plate 16, 151
Fig. 1, 9 car., ext. It. lat. (paratype, OS13434, 506 long); fig. 2, 9 car., ext. dors, (paratype, OS13435, 552/u.m long); fig. 3,
Cf car., ext-, dors, (paratype, OS13436, 607 /xm long). Scale A (250/xm; X 105), figs. 1-3.
Holotype:
Type locality:
Derivation of name:
Figured specimens:
Diagnosis:
Stereo-Atlas of Ostracod Shells 16, 149
Limnocythere hibernica (2 of 4)
Limnocythere hibernica (4 of 4)
Stereo- Atlas of Ostracod Shells 16, 151
Stereo- Atlas of Ostracod Shells 16 (31) 152-155 (1989) Echinocythereis spinireticulata (1 of 4)
595.337.14 (119.9) (263 : 162.090.29) : 551.351
ON ECHINOCYTHEREIS SPINIRETICULATA KONTROVITZ
by Mervin Kontrovitz & Zhao Yuhong
(Northeast Louisiana University, Monroe , USA &
Nanjing Institute of Geology & Palaeontology, Academia Sinica, Nanjing, China)
Echinocythereis spinireticulata Kontrovitz, 1971
1971 Echinocythereis spinireticulata sp. nov. M. Kontrovitz, Tulane Stud. Geol. Paleont.. 8, 166-168, pi. 1. figs. 1-3, text-fig. 1.
1975 Echinocythereis spinireticulata Kontrovitz; H. V. Howe & W. A. van den Bold, Bull. Am. Paleont., 65, 307, pi. 2, fig. 4.
Holotype: H. V. Howe Collection (HVH), Louisiana State University, Baton Rouge, USA, no. HVH 8595;
9 left valve.
[Paratypes: nos. HVH 8596-8599]
Type locality: Gulf of Mexico, near the delta of the Mississippi River, approx, lat. 29°00'N, long. 90°00'W;
Holocene, marine, sublittoral.
Figured specimens: Geosciences Department of Northeast Louisiana University (NLUGEO) nos. NLUGEO 1021 (9
RV: PI. 16, 153, fig. 1; PI. 16, 155, fig. 3), NLUGEO 1022 (9 LV: PI. 16, 153, fig. 3; PI. 16, 155,
fig. 1), NLUGEO 1023 (juv. RV: PI. 16. 153, fig. 2), NLUGEO 1024 (juv. LV: PL 16, 155, fig. 2).
From the Gulf of Mexico, near the delta of the Mississippi River; Recent, marine (kindly
provided by Ms J. M. Slack (NLU)).
Explanation of Plate 16. 153
Fig. 1, 9 RV, ext. lat. (NLUGEO 1021, 1110/xm long); fig. 2, juv. RV ext. lat. (NLUGEO 1023, 830 pm long); fig. 3, 9 LV, ext. lat.
(NLUGEO 1022, 1110/xm long).
Scale A (250 /xm; x70), figs. 1-3.
Stereo-Atlas of Ostracod Shells 16, 154 Echinocythereis spinireticulata (3 of 4)
Diagnosis: Surface covered with minute, delicate spines, arranged in rows that form a reticulate pattern.
Ornamentation centered around dorsomedial area of valve. Adults have a row of slender spines
behind the denticulate anterior margin. Posterior margin without denticles. Heavy spine
projecting from posterior ventrolateral convexity of adults. Vertical row of 4 adductor muscle
scars: from dorsal to ventral, the first scar is oval, the second subreniform, the third and fourth
elongate; second and fourth scars nearly touch in front of the third.
Remarks: This species differs from Echinocythereis jacksonensis (Howe & Pyeatt) (in Howe & Chambers,
Geol. Bull. La. , 5, 35-37, pi. 1, figs. 23-24; pi. 6, fig. 31, 1935) in being reticulate over the entire
surface and having a higher posterior, therefore appearing to be shorter. The original description
of E. jacksonensis included two forms, one larger than the other. The larger has anterior
reticulations and only coarse spines from mid-length to the posterior.
Krutak (/. Paleont., 35, 783-784. pi 91, fig. 9, 1961) figured examples of E. jacksonensis on
which “tiny nodes are aligned, tending to form hexagonal, pentameral, or angular patterns". The
specimens he reported also differ from this species in being smooth in the dorsal, posterior, and
ventral areas. Muscle scars and length/height ratios are also significantly different.
E. clarkana (Ulrich & Bassler) (in Case etal.. Systematic Paleontology of the Miocene Deposits
of Maryland, 98, pi. 35, figs. 1 — 10. Miocene Volume. Maryland Geological Survey, Baltimore.
1 904) is distinguished from E. spinireticulata by its coarsely reticulate surface with heavy spines at
the junctures of the ridges, denticulate posterior margin, and larger size.
Distribution: Common in shallow, sublittoral, marine waters of the Gulf of Mexico near the delta of the
Mississippi River; also recovered from the Mississippi Mudlumps (Howe & van den Bold, 1975).
Explanation of Plate 16, 155
Fig. 1, 9 LV, int. lat. (NLUGEO 1022, UlO^m long); fig. 2, juv. LV int. lat. (NLUGEO 1024, 830 /xm long); fig. 3, 9 RV, int. lat.
(NLUGEO 1021, 1110/xm long).
Scale A (250 /xm; x70), figs. 1-3.
Echinocythereis spinireticulata (4 of 4)
Stereo-Atlas of Ostracod Shells 16, 155
Stereo-Atlas of Ostracod Shells 16, 153
Echinocythereis spinireticulata (2 of 4)
Stereo-Atlas of Ostracod Shells 16 (32) 156-157 ( 1989)
Index , Volume 16, 1989(1 of 2)
General Index
Abyssobythere guttata Ayress & Whatley gen. et sp. nov.; 136-139
Adamczak, F. & Becker, G., On Aurikirkbya wordensis (Hamilton); 122-115
Adamczak, F. & Becker. G.. On Rishona epicypha (Kesling & Kilgore); 51-54
Athersuch. J.. On Limhocythere hibernica Athersuch sp. nov.; 148-151
atlantica. Tuber oloxoconcha: 73-76
Aurikirkbya wordensis (Hamilton); 112-113
Ayress. M. A. & Whatley. R. C., On Abyssobythere guttata Ayress & Whatley gen. et sp. nov.; 136-139
Balticella deckeri (Harris); 94-99
Becker. G., On Kullmannissites kullmanni Becker: 43-46
Becker. G.. On Nodella hamata Becker; 116-119
Becker. G., On Sinessites hispanicus Becker: 39-42
Becker. G.. On Vitissites comtei Becker; 47-50
Becker, G. & Adamczak. F.. On Aurikirkbya wordensis (Hamilton); 112-115
Becker. G. & Adamczak. F. . On Rishona epicypha (Kesling & Kilgore); 51-54
Berolinella stcusloffi (Krause); 106-111
Bhatia. S. B. & Keyser. D.. On Strandesia weberi (Moniez); 128-135
Bromidella reticulata Harris; 1-8
brunensis, Buntonia : 77
Bryocypris grandipes Roen ; 140-147
Buntonia brunensis Rfha; 77
Bythoceratina gobanensis Reyment & Reyment sp. nov.; 21-24
Chinocy there curvispinata Su sp. nov.; 55-58
Chinocythere shajingensis Su sp. nov.; 59-62
Chinocythere tuberculata Su sp. nov.; 63-66
Columutia variolata (Jones & Holi); 29-34
comtei, Vitissites ; 47-50
curvispinata, Chinocythere ; 55-58
Cvtheridea sandbergeri Kammerer sp. nov.; 120-127
Dabashanella retroswinga Huo, Shu & Fu; 13-16
deckeri, Balticella'. 94-99
distorta, Microclieilinelhr, 35-38
Echinocythereis spinireticulata Kontrovitz; 152-155
epicypha, Rishona ; 51-54
Fallaticella schaeferi Schallreuter; 25-28
gobanensis, Bythoceratina: 21-24
grandipes, Bryocypris: 140-147
guttata, Abyssobythere: 136-139
hamata, Nodella : 116-119
Hansch, W. & Siveter. D. J.. On Berolinella steusloffi (Krause); 106-111
Hansch, W. & Siveter, D. J.. On Macrypsilon salterianum (Jones); 100-105
hibernica, Limnocythere ; 148-151
hispanicus, Sinessites: 39-42
Horne, D. J., On Tuberoloxoconcha atlantica Horne sp. nov.; 73-76
Horne, D. J.. On Tuberoloxoconcha tuberosa (Hartmann); 67-72
Kammerer, T. On Cytheridea sandbergeri Kammerer sp. nov.; 120-127
Keyser, D. & Bhatia, S. B., On Strandesia weberi (Moniez); 128-135
Kontrovitz, M. & Zhao, Y., On Echinocythereis spinireticulata Kontrovitz; 152-155
kullmanni, Kullmannissites: 43-46
Kullmannissites kullmanni Becker; 43-46
levigata, Progonocy there: 17-20
Limnocythere hibernica Athersuch sp. nov.; 148-151
Lophocypris shulanensis Zhang & Zhao gen. et sp. nov.; 9-12
Lundin, R. F. , On Microcheilinella distorta (Geis); 35-38
Lundin, R. F. & Petersen, L. E., On Primitivothlipsurella obtusa Petersen & Lundin sp. nov.; 86-93
Lundin, R. F. & Petersen, L. E., On Primitivothlipsurella v-scripta (Jones & Holl); 78-85
Lundin, R. F. & Siveter, D. J., On Columatia variolata (Jones & Holl); 29-34
Macrypsilon salterianum (Jones); 100-105
Martens, K., On Bryocypris grandipes Roen ; 140-147
Microcheilinella distorta (Geis); 35-38
Nodella hamata Becker; 116-119
obtusa, Primitivothlipsurella: 86-93
Petersen, L. E. & Lundin, R. F., On Primitivothlipsurella obtusa Petersen & Lundin sp. nov.; 86-93
Petersen, L. E. & Lundin, R. F., On Primitivothlipsurella v-scripta (Jones & Holl); 78-85
Primitivothlipsurella obtusa Petersen & Lundin sp. nov.; 86-93
Primitivothlipsurella v-scripta (Jones & Holl); 78-85
Progonocythere levigata Bate; 17-20
reticulata, Bromidella: 1-8
retroswinga, Dabashanella: 13-16
Reyment, E. R. & Reyment, R. A., On Bythoceratina gobanensis Reyment & Reyment sp. nov.; 21-24
Reyment, R. A. & Reyment, E. R., On Bythoceratina gobanensis Reyment & Reyment sp. nov.; 21-24
Riha, J., On Buntonia brunensis Ri'ha; 77
Rishona epicypha (Kesling & Kilgore); 51-54
salterianum, Macrypsilon: 100—105
sandbergeri, Cytheridea: 120-127
schaeferi, Fallaticella: 25-28
Schallreuter, R. E. L., On Fallaticella schaeferi Schallreuter; 25-28
shajingensis, Chinocythere: 59-62
shulanensis, Lophocypris: 9-12
Sinessites hispanicus Becker; 39-42
Siveter, D. J. & Hansch, W. , On Berolinella steusloffi (Krause); 106-111
Siveter, D. J. & Hansch, W. , On Macrypsilon salterianum (Jones); 100-105
Siveter, D. J. & Lundin, R. F. , On Columatia variolata (Jones & Holl); 29-34
Index, Volume 16, 1986 (2 of 2)
Stereo-Atlas of Ostracod Shells 16, 157
Siveter, D. J. & Wakefield, M. I., On Progonocythere levigata Bate; 17-20
Siveter, D. J. & Williams, M., On Balticella deckeri (Harris); 94-99
Siveter, D. J. & Williams, M., On Bromidella reticulata Harris; 1-8
spinireticulata, Echinocythereis; 152-155
steusloffi, Berolinella ; 106-111
Strandesia weberi (Moniez); 128-135
Su, D., On Chinocythere curvispinata Su sp. nov.; 55-58
Su, D., On Chinocythere shajingensis Su sp. nov.; 59-62
Su, D., On Chinocythere tuberculata Su sp. nov.; 63-66
Tong, H. & Zhao, Y. , On Dabashanella retroswinga Huo, Shu, & Fu; 13-16
tuberculata, Chinocythere ; 63-66
Tuberoloxoconcha atlantica Horne sp. nov.; 73-76
Tuberoloxoconcha tuberosa (Hartmann); 61-12
tuberosa, Tuberoloxoconcha', 67-72
variolata, Columatia ; 29-34
Vitissites comtei Becker; 47-50
v-scripta, Primitivothlipsurella ; 78-85
Wakefield, M. I. & Siveter, D. J. On Progonocythere levigata Bate; 17-20
weberi, Strandesia', 128-135
Whatley, R. C. & Ayress, M. A., On Abyssobythere guttata Ayress & Whatley gen. et sp. nov.; 136-139
Williams, M. & Siveter, D. J., On Balticella deckeri (Harris); 94-99
Williams, M. & Siveter, D. J., On Bromidella reticulata Harris; 1-8
wordensis, Aurikirkbya; 112-115
Zhang, L. & Zhao, Y., On Lophocypris shulanensis Zhang & Zhao gen. et sp. nov.; 9-12
Zhao, Y. & Kontrovitz, M., On Echinocythereis spinireticulata Kontrovitz; 152-155
Zhao, Y. & Tong, H., On Dabashanella retroswinga Huo, Shu & Fu; 13-16
Zhao, Y. & Zhang, L. On Lophocypris shulanensis Zhang & Zhao gen. et sp. nov.; 9-12
Index; Geological Horizon
See 1 (2) 5-22 (1973) for explanation of the Schedules in the Universal Decimal Classification
(261.1)
(261.2)
(263)
(411)
(420)
(430.1)
(430.2)
(437)
(438)
(44)
(460)
Index; Geographical Location
See 1 (2) 5-22 (1973) for explanation of the Schedules in the
North Atlantic Ocean: (485)
Bythoceratina gobanensis; 21-24
North East Atlantic
Limnocythere hibernica; 148-151 (510)
Gulf of Mexico:
Echinocythereis spinireticulata', 152-155
Scotland:
Progonocythere levigata', 17-20
Tuberoloxoconcha atlantica', 73-76
England: (540)
Columatia variolata', 29-34
Primitivothlipsurella obtusa; 86-93 (596)
Primitivothlipsurella v-scripta ; 78-85
Progonocythere levigata', 17—20 (671.1)
German Federal Republic:
Cytheridea sandbergeri; 120-127 (744)
Nodella hamata; 116-119
German Democratic Republic: (755)
Berolinella steusloffi', 106-111
Macrypsilon salterianum', 100-105 (766)
Czechoslovakia:
Buntonia brunensis', 11
Poland: (772)
Macrypsilon salterianum', 100-105
France: (774)
Tuberoloxoconcha tuberosa', 67-72
Spain: (775)
Kullmannissites kullmanni ; 43-46
Sinessites hispanicus; 39-42 (923)
Vitissites comtei', 47-50
Universal Decimal Classification
Sweden:
Fallaticella schaeferi ; 25-28
Primitivothlipsurella v-scripta', 78-85
China:
Chinocythere curvispinata ; 55-58
Chinocythere shajingensis', 59-62
Chinocythere tuberculata', 63-66
Dabashanella retroswinga', 13-16
Lophocypris shulanensis', 9-12
India:
Strandesia weberi', 128-135
Cambodia:
Strandesia weberi', 128-135
Cameroon:
Bryocypris grandipes\ 140-147
Massachusetts:
Tuberoloxoconcha atlantica', 73-76
Virginia:
Balticella deckeri ; 94-99
Oklahoma:
Balticella deckeri ; 94-99
Bromidella reticulata', 1-8
Indiana:
Microcheilinella distorta', 35-38
Michigan:
Rishona epicypha; 51-54
Wisconsin:
Aurikirkbya wordensis ; 112-115
Timor:
Abyssobythere guttata ; 136-139
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Stereo-Atlas of Ostracod Shells: Vol. 16, Part 2
16(19) 78-85
16 (20) 86-93
16(21) 94-99
16 (22) 100-105
16 (23) 106-111
16 (24) 112-115
16 (25) 116-119
16 (26) 120-127
16 (27) 128-135
16 (28) 136-139
16 (29) 140-147
16 (30) 148-151
16 (31) 152-155
CONTENTS
On Primitivothlipsurella v-scripta (Jones & Holl ) ; by R. F. Lundin & L. E.
Petersen
On Primitivothlipsurella obtusa Petersen & Lundin sp. nov.; by L. E.
Petersen & R. F. Lundin
On Balticella decked (Harris); by M. Williams & D. J. Siveter
On Macrypsilon salteriamun (Jones); by D. J. Siveter & W. Hansch
On Berolinella steusloffi (Krause); by W. Hansch & D. J. Siveter
On Aurikirkbya wordensis (Hamilton); by G. Becker & F. Adamczak
On Nodella hamata Becker; by G. Becker
On Cytheridea sandbergeri Kammerer sp. nov.; by T. Kammerer
On Strandesia weberi (Moniez); by D. Keyser & S. B. Bhatia
On Abyssobythere guttata Ayress & Whatley gen. et sp. nov. ; by M. Ayress
& R. C. Whatley
On Bryocypris grandipes R0en; by K. Martens
On Limnocythere hibernica Athersuch sp. nov.; by J. Athersuch
On Echinocythereis spinireticulata Kontrovitz; by M. Kontrovitz & Zhao
Yuhong
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